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..\ THE ANNALS
a
Poe
ee
‘
AND
MAGAZINE OF NATURAL HISTORY,
INCLUDING
ZOOLOGY, BOTANY, ann GEOLOGY.
(BEING A CONTINUATION OF THE ‘ANNALS’ COMBINED WITH LOUDON AND
CHARLESWORTI’S ‘ MAGAZINE OF NATURAL HISTORY,’ )
CONDUCTED BY
ALBERT C. L. G. GUNTHER, M.A., M.D., Ph.D., F.B.S.,
WILLIAM CARRUTHERS, Ph.D., F.R.S., F.L.S., F.G.S.,
AND
WILLIAM FRANCIS, F.L.S.
VOL. XX.—SEVENTH SERIES.
—_—~ ~
J a ig ts
/
i >
\ AAZVOS
Ww
wonal Museu
————
LONDON:
PRINTED AND PUBLISHED BY TAYLOR AND FRANCIS,
SOLD BY SIMPKIN, MARSHALL, HAMILTON, KENT, AND CO., LD. ;
BAILLIERE, PARIS: HODGES, FIGGIS, AND CO,, DUBLIN :
AND ASHER, BERLIN.
1907,
As
“Omines res creatz sunt divine sapientiz et potentiz testes, divitie felicitatis
humane :—ex harum usu onitas Creatoris; ex pulchritudine sapientia Domini;
ex ceconomiaé in conservatione, proportione, renovatione, potentia majestatis
elucet. Earum itaque indagatio ab hominibus sibi relictis semper eestimata ;
a veré eruditis et sapientibus semper exculta; malé doctis et barbaris semper
inimica fuit.”—Linnzvs.
“Quel que soit le principe de la vie animale, il ne faut qu’ouvrir les yeux pour
voir qu’elle est le chef-d’ceuvre de la Toute-puissance, et le but auquel se rappor-
tent toutes ses opérations.”—Bruckner, Théorie du Systéme Animal, Leyden,
1767.
Diiiiels & bel ele) see heisyivan powers
Obey our summons; from their deepest dells
The Dryads come, and throw their garlands wild
And odorous branches at our feet; the Nymphs
That press with nimble step the mountain-thyme
And purple heath-flower come not empty-handed,
But scatter round ten thousand forms minute
Of velvet moss or lichen, torn from rock
Or rifted oak or cavern deep: the Naiads too
Quit their loved native stream, from whose smooth face
They crop the lily, and each sedge and rush
That drinks the rippling tide: the frozen poles,
Where peril waits the bold adventurer’s tread,
The burning sands of Borneo and Cayenne,
All, all to us unlock their secret stores
And pay their cheerful tribute.
J. Taytor, Norwich, 1818.
CONTENTS: OF VOL. XX.
[SEVENTH SERIES. }
NUMBER CXV.
I. New African Saturnide. By the Hon. L. W. Roruscuitp,
LE] D eh ee 0c 0 thn yooh NIN Care earta aaa eae yaaa Se errr
II. A Contribution to the Knowledge of the Hymenoptera of the
Oriental Zoological Region. By P. CAMERON .............+.---
Ill. On Three new Mammals from South Africa. By R. C.
WVROUGEBLONG Sopot aes sata ccuercccin ted he ee os ttn Ratio cee hea ohare
IV. Descriptions of new Species of African Spiders and Solifuge.
EK vge Nem Sea EATER STM tener eyare Max «iss sen MPA apoio ohaas ed elas Mel Mist aka otal anon
V. Remarks on Prof. L. von Méhely’s Paper “Zur Losung der
‘ Muralis-Frage.’” By G. A. BouLenesr, F.R.S...............
VI. Description of a new Engystomatid Frog of the Genus
Breviceps from Namaqualand. By G. A. BouLencer, F.R.S.
PETa Coe US) bers carved ch cate Sect era arc nl y Oldocbeievha tea alge d dette conic
VII. Descriptions of a new Toad and a new Amphisbeenid from
Mashonaland. By G. A. BouLencER, F.R.S. (Plate ILI.) ......
VIII. Description of a new Cichlid Fish from Portuguese East
Perce. Dye Cr AL DOULENGER, ES. adcectaclace ss oe sere:
IX. Descriptions of Three new Freshwater Fishes discovered by
Mr. G. L. Batesin South Cameroon. By G. A. BouLENGER, F.R.S.
X. The Atractylis coccinea of T.S. Wright. By E.S. Russet,
WVU Ate peyeags u8 Sa sina en et age AAT PAR ON ae: ah V PA ed are
XI. On the Generic Position of Benson’s Helix hyba and the
Similarity of its Anatomy to that of Khastella vidua, W. T. Blanford.
By Lt.-Colonel H. H. Gopwin-AuvstEn, F.R.S. &. ............
XII. Descriptions and Records of Bees—XV. By T. D. A.
CockKuRELE. University, of Coloradoy . 0 ft dau. ons vce ceo anes
XII. Notes on the Habits and External Characters of the
Solenodon of San Domingo (Solenodon paradoxus), By A. Hyatr
VSR Eee CCE LEbOM NV -)) Sth. s.45 0 5-tcpih ale wi, eee tind ade ocle Ses
XIV. On Three new Mammals from British New Guinea. By
OLDFIELD THOMAS........ Roch bc t €cge both cx PROOTaor PAuOtEL
XV. New and little-known Eastern Moths. By Colonel C.
SIEMPRE NENG dbs yey OC Cararaial sve sh ohsisfeliets Satna Saks eres cata dens s + ¢
New Book :—Books and Portraits illustrating the History of Plant
OJ SEENRET ING Ce BGG n CODOBO URE OMe Obbt Coct noc nonneraort
Pages
5
iv CONTENTS.
NUMBER CXVI.
XVI. A Contribution to the Knowledge of the Hymenoptera of
the Oriental Zoological Region. By P. CAMERON ...........04.
XVII. On a new Leptolepid Fish from the Weald Clay of South-
water, Sussex. By A. SmirH Woopwarp, LL.D., F.R.S. (Plate I.)
XVIII. On new Species of Histeride and Notices of others. By
CRATES; AU <M 153,085 v csctee octets uC CAG EAT RR EOte etek oh eee eRe
XIX. Descriptions of Two new Freshwater Fishes discovered by
Dr. W. J. Ansorge in Mossamedes, Angola. By G, A. BouLENGER,
1 8s] StS eee tele oan einen ota Otitis ane frie Butb ont bas
XX. Description of a new Frog discovered by Dr. W. J. Ansorge
in Mossamedes, Angola. By G. A. BouLencsEr, F.R.S. ........
XXI. On the African Mungooses usually referred to the Herpestes
gracilis Group. By R. C. WroueHtTon
XXII. The Name of the Armenian Wild Sheep. By R.
PG YDWRRER sips kn sia ike 2 aie iat cee is, oe elec ae ae aI Oe TE RRR RI ee
XXIII. Descriptions and Records of Bees—XVI. By T. D. A.
CocKERELL, University of Colorado
©) lo MO! 0ice)Te) alco le) ee! 618 B,eel.@ (ele Lee je8\re
XXIV. Notes on the American Species of Hespertide described
by Elotz.. By FF. D;Gopman, DiC.L.; HORS., &e. 2.0... belle
XXYV. Function and Form with Reference to the Hand and Foot
in Man and Apes. By DunoanC. L. Frrzwiiiams, M.D., Ch.M.,
F.R.C.SS. Ed. & Eng., Demonstrator of Anatomy, King’s College,
London ; Casualty Officer, Hospital for Sick Children, Great Ormond
Sureet. s(blate: Ve). osc crctnncine snc eee eras ei era
XXVI. On Neotropical Mammals of the Genera Callicebus,
Reithrodontomys, Ctenomys, Dasypus, and Marmosa. By OLDFIELD
THOMAS
GAe' 056: G6) ©). 4 -o\.0' (0! 4/6" (0. .0)'6) 61110 10) ,0, 16: n) 6b 0.Feije, 0, 16)'41¢ ce 016) 01016 ce) fa Sib fave) elles! eWatarelie
eee et :0' 19 (07 w. eN elie? 6 e-16 He 0 Geiie helio iene, fe Je! 16,0) 10) 1 (6: (0.10.78! n0 ees el alsa Jee" 8) ew Joga “si ia, et einehelieie: ie
NUMBER CXVII.
XXVIII. Notes from the Gatty Marine Laboratory, St. Andrews.—
No. XXVIII. By Prof. M’Inrosu, M.D., LL.D., F.R.S., &c.
Hila ces VE VIELE) i. oc alere ii ora aaceuten rears Gre eda Pde ono ae
XXVIII. Descriptions of apparently new Species and Subspecies
of Mammals belonging to the Families Lemuride, Cebide, Calli-
trichide, and Cereopithecitde in the Collection of the Natural History
Museum. “By DG. Bunton, DiSe., BORIS EwWen acces ate ce ahs
XXIX. On Mammals from Northern Persia, presented to the
National Museum by Col. A.C. Bailward. By OLpriei.p THomas,
BRAS.
Pie ie oinielvepien tea (d ace ei) 1a) (ero, ce: oie ie) 8j.5) mia ace eile ehtbcie ie ie) fa -9)\87'e) s/h er 8, 1m. v.10: (e116) (ble) s/uemeliaey
Page
81
93
95
155
161
168
169
CONTENTS. Vv
Page
XXX. A Subdivision of the Old Genus Nesokia, with Descriptions
of Three new Members of the Group, and of a Mus from the
Andamans.” (by: OLDRIOLD LH OM ASE cite aici. o)'o vieiejoid @iereleoacs oboe 202
XXXI. On some British Polyzoa. By Canon A. M. Norman,
SAC DOs Gln Dry Boke cru elncn (bate Ik.) Web dc vee < 207
1
XXXII. Three new Spanish Insectivores. By ANGEL CaBRERA, 212
XXXIII. On Four new Pill-Millipedes from the Malay Peninsula
and Siam. By A. 8. Hirsr (British Museum, Nat. Hist.).
GES S.c a Eres 10 ap o Geol gop ORs Gator e Colo ertT. te ore ian an ea 215
XXXIV. Description of an apparently new Lycenid from Mauri-
tiuss) By asm rOn Hh. DRUCES EIZ.S. FONS. fie... < dese e snine 219
XXXV. On an Extinct undescribed Fruit-Bat of the Genus
Pteropus from the Mascarenes. By Gro, E. Mason ............ 220
XXXVI. Brachiopod Nomenclature: Seminula, &c. By 8.58.
WE Tal ORANG be Ce Sich Faaye ey aS craton oda¥ ah si acdtge dhe Gisee ak coheed oe uelehera ob ab cheat gtaracemlc oe
XXXVII. On some Freshwater Amphipods: The Reduction of
the Eye in a new Gammarid from Ireland. By Prof. Dr. Fr.
Wa OViSKeveuy (Gblatest Mul Gg. 2XonIe))) Pttie vercyarc testis fe? st lv oi alexeqster 4 wile ad)
XXXVIII. Observations on the Trematode Parasites of British
Birds. By Witi1AM Nicoxt, M.A., B.Sc., Gatty Marine Laboratory, _
Rear MTT WV SEN way al oeettinyafefcv are. 5)s: orehasetey seeps) sceiereyerore.oei oes Cosanie''ei tis sole a0o¥e 245
XXXIX. Preliminary Report on the Monaxonellida of the
National Antarctic Expedition. By R. KIRKPATRICK .......... 271
XL. Description of an apparently new Species of Monkey of the
Genus Cebus. By D. G. Evuiot, D.Se., F.R.S.E., &e. 2... een 292
XLI. On the Occurrence of <Acanthoglossus in British New
Gauitieasse DY= OLDETET DWE HOMAS ) We /.0e oie'ele © acura Siena ities sea LOD
New Books :—1 Go A-Walking. Through the Woods. With Tllus-
trations by CHARLES Rerp.—Nature in School. No. 2.
Edited by Boris WEINBERG, Priy. Doc. St. Petersburg Univ. . 295
NUMBER CXVIII,
XLII. New South-African Spiders of the Family Drassede in the
Collection of the South-African Museum, By W. F. Purce t,
PhDs Cane Bown. (Plates XL RCV ot os; eietete Sener ayy ones wheraye aces 297
XLIII. Descriptions of Two new African Species of Barbus. By
CrP AG ES OUEAONGIOR OE CEU) (kd acre ahd dcdogeiwledshald = oie tlslatafesovdyop ited se aioks 336
XLIV. Description of a new Pit-Viper from Brazil. By G. A.
EA UUIERIN GIG Rie eek Ua Sepia ave ony atstces ye) ake PARTE Sd ek age og Shen sca ist vores 338
XLY. On new Species of Histertde and Notices of others. By
Gt Lewis, F.L.S.
mii eo. ehel elie ene eve OL Sle. ue, ehelepe re ene) ee) el eseye ie) tie st @ e.wiel.e ane
v1 CONTENTS.
Page
XLVI. Some Remarks on Pteropus mascarinus, Mason. By Knup
JARO ES NAG Sg ac ewe de Sa asse don hyn eae ae Sododencomaecds ds ot. 351
XLVITI. Notes on the Crustacea of the Channel Islands. By
Canon A. M. Norman, M.A., D.C.L., LL.D., F.RS., F.LS.
RE a bese NAN Me Or ONOVMDIIS) © Keema nce wieteiels saje oiels « Graie aig ys etereustesma tates 356
New Books :—100 Photographs from Life. (No. 1.) By DouGias
Eneiisa.—l00 Photographs from Life. (No.2.) By R. B.
Loper.—Les Débuts d’un Savant Naturaliste, le Prince de
lEntomologie, Pierre André Latreille, & Brive, de 1762 a 1796.
Par Lours pk Nussac, Sous-Bibliothécaire du Museum d’His-
toire naturelle—Epm. BorpaGcu. Recherches anatomiques et
biologiques sur l’Autotomie et la Régénération chez divers
Arthropodes.—The Fauna of British India, including Ceylon
and Burma. Published under the authority of the Secretary of
State for India in Council. Edited by Lt.-Col. C.T. Bryauam.
Butterflies. Vol. II. By Lt.-Col. C.T. Binguam...... 372, 373
Proceedings of the Geological Society .................005 374—376
NUMBER CXIX.
XLVIII. Preliminary Notice of new and remarkable Cephalopods
from the South-west Coast of Ireland. By A. L. Massy, Depart-
ment of Agriculture and Technical Instruction, Fisheries Branch,
Dee GS Re ek BA lea date ou Wib,o ots ih fe eae mtetsale She ietene wantnehe tiets 377
XLIX. On the Geographical Races of the Lesser Horseshoe Bat
(Rhinolophus hipposiderus). By KNuD ANDERSEN .............. 384
L. Some new European Insectivora and Carnivora. By Grrrir
roy EN Doty rap hE ES Search ican Cio yen eRCrC en dio aia eehec Gia aon tu 389
LIL Two new Forms of the Spanish Hare. By Gerrit S.
PMSA SEVER coikcve folate etch oleae cialonale ro lors sore ts iels hola eielrer ake leig ss ee tCiet tage 398
LII. Description of Two new Characinid Fishes from South
America; « SBy iC, slam shunGAIN, VL AS 5.0 rie cee et testes torsion te 402
LIII. Note on Raia undulata, Lacep. By C. Tate Reean, M.A. 403
LIV. On a new Hare from the Transvaal. By H. Lyster
AME SON Getveaers ete ole eric aieojtutes. te mine ioon Regan irom our beast p tech pane 404
LV. On anew Dormouse from Asia Minor, with Remarks on the
Subgenus “ Dryomys.” By OLDFIELD THOMAS .............00 406
LVI. Notes onan “ Octopus” with Branching Arms. By EpGar
ARS MITH, LS Oia ZiSes (Plate SXAVAUL:) ote Navs ice top tices oie ee ce ead 407
LVII. Rhynchotal Notes—XLIII. By W.L. Disrant ...... 411
LVIII. Description of a new Genus and Species of Cerambycide
fromeNatal, «By WA DISTANT <. 2 sigs. cau osc cviemee ee pote ewe a ek 423
LIX. Description of a new Species of Cicadide from East Africa.
SV Ve MOIRIDAINI Eat iki te Gn tanh ie «None th whih RAs oie, ae eit cole om ». 425
CONTENTS. Vil
Page
LX. Four new European Squirrels. By Grerrit 8. Minuer.... 426
LXI. New Species of African and TadosWalayan Hesperude. By
(Wolonel C SwinBon,-MeAv.. HRS Ge. is a uSwa 6 ads socked cd 430
LXII. The Significance of the Pattern of the Cubs of Lions (Felis
leo) and of Pumas (Felis concolor). By R. I. Pococr, F.IS.,
¥.Z.S., Superintendent of the Zoloogical Society’s Gardens. (Plates
DG D.C 9. BO) IN 6, SET eee iy OL ORI eSB a C COROOE EC Se IIOM ET EET Tae 436
LXIII. Descriptions and Records of Bees—XVII. By T. D. A.
CockuEMEn.Wmiversity of Colorado. 3 vs :7s.y.ves tie sie d olin som s50 < 445
Echinocrinus versus Archeocidaris, by F. A. Bather, British Museum
(NGM ERISES ptrete rs eed re ttt sc etamiues owen nate Mele ae leads 452
NUMBER Cxx.
LXIV. A Revision of the Medusz belonging to the Family
Laodiceide. By Epwarp T, Browne, University College, London. 457
LXY. On new Species of Histeride and Notices of others. By
Cello aish dial bas bee ep eSiO Ditis Ain Paice eIOIRirhs bio SNe ean mie IPE 480
LXVI. Descriptions of Three new Freshwater Fishes discovered
by Mr. G. L. Bates in South Cameroon. By G. A. BouLENGER,
I GIS odin OA Ow tro OIG Cnn OURO RNR tere Rae Gira RienenD oe 485
LXVII. Descriptions of Three new Fishes from Central Africa.
iBygG. AG DOUMENGRR s Hoku Sis qwerials ta: S40 ssl ciaureste « Gecleee = 487
LXVIII. Description of a new African Snake of the Genus Simo-
cephalus, By G. AS BOUGENGER WE RAS. 55.) 0... ccceecc- sess 489
LXIX. Descriptions of new Species of Land and Freshwater
Shells from Central and South America. By H. B. Preston, F.Z.S. 490
LXX. A new Acanthoglossus from the Island of Salawatti. By
OM PTH DMM ONMAS® oy. ha tere eee sor els rat ais othe oes 498
LXXI. Description of a new Baboon from British East Africa.
Syme) s Coe Mare TOs). OC.5 Ec kuaSats. (OCs aamiere cd ohr-s/ster aro fetes occa 499
LXXII. List of a Collection of small Mammals made by Mr. A. L.
Butler in the Bahr-el-Ghazal. By R.C. Wroucuron .......... 500
LXXIII. Descriptions of Five new Species of Heterocera, By
ERR BERIE ORE CHE MSR OCCk «sic ssloccciis Meme alert d eeeve. serererst. 505
LXXIV. New African Species of the Genus Chrysops, Meigen, in
the British Museum (Natural History). By Ernest E. AuSTEN .. 507
LXXV. Descriptions of Two new Species of Plecotus. By G. E. H.
Eye EL AMCOEETON: ciayetsjersiute aijeoyaNat ve) a ari aye) ews Ce ws 520
LXXVI. Description of a new ‘ ores of Monkey (Cercopithecus
Hamlyn) from the Ituri Forest. By R. I. Pocock, F.L.S., Super-
intendent of the Zoological Society's Gardens ........0+0.0ee ees 52]
Vill
LXXVII. A new Flying-Squirrel from Formosa.
EOMAS | 150%, catia
LXXVIII. Notes on Two Species of African Fresh water Sponges.
By R. Kimkparrick 52
New Books :—Memoirs of the Department of Agriculture in India
(Entomological Series).—Homoptera Andina: Die Zikaden des
Kordillerengebietes von Stidamerika nach Systematik und Ver-
I, Cicadidea.—Précis des Caractéres
génériques des Insectes, disposés dans un ordre naturel par le
Citoyen Latreillecstpsumuewd« a ecapue outs petleeent
breitung, von A, JAcosr.
Pndexiaerre tect
PLATE I,
oT;
III.
IV.
V.
VI.
VU.
VIII.
IX.
x.
XI.
Xi,
XIII.
KIY.
XV.
; Crustacea from the Channel Islands.
XVII:
XVIII.
; Lion and pwna euxbs.
XVI
XIX
XX
CONTENTS.
900 6 00:0 0) 6 O10 Biers ele)6 @ s 6 9 5 dé elefe 6 0 60 0.6 SiGe 0 9.0) sin elena
B,8. 10 GL) 0le.dhese, 0,\9,-6)8, Ciel @ 0 (6.0 0 ple 0. 6 6,6 $169 Ns lo es eletnen ue hs
PLATES IN VOL. XX.
Leptolepis valdensis.
Breviceps macrops.
Bufo anotis.
San Domingo Solenodon,
Hand and foot of Hylobates agilis.
Head of porbeagle shark.
Ommastrephes sagittatus,
Nereis diversicolor.
British Polyzoa.
Pill-Millipedes.
Freshwater Amphipods.
South-African spiders.
Octopus with branching arms.
By Onprirenp
Vol. 20. SEVENTH SERIES. No. 115.
oe Pacey nc enn ae aA A AEA ag Teens sk
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Taytor and Francis, Red Lion Court, Fleet Street, E.C.
Second edition, Vols. 1. & II. Royal 8vo, price 12s, net, each.
CATALOGUE
OF THE
PHYSIOLOGICAL SERIES OF COMPARATIVE ANATOMY
IN THE
MUSEUM OF THE ROYAL COLLEGE OF SURGEONS OF ENGLAND.
Tayzor and Francis, Red Lion Court, Fleet Street, E.C.
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES. ]
Ce ncemeoncennnnOsed per litora spargite muscum,
Naiades, et circitm vitreos considite fontes:
Pollice virgineo teneros hic carpite flores:
Floribus et pictum, dive, replete canistrum.
At yos, o Nymphee Craterides, ite sub undas ;
Ite, recurvato variata corallia trunco
Vellite muscosis e rupibus, et mihi conchas
Ferte, Dez pelagi, et pingui conchylia suceo.”
N. Parthenii Giannettasi, Ecl, 1.
No. 115. JULY 1907.
I.—New African Saturniide.
By the Hon. L. W. Roruscuiyp, Ph.D.
1. Epiphora pelosoma, sp. n.
Body clay-colour ; upperside of thorax hair-brown; meta-
notum, upperside of last abdominal ring, two thin ventral
lateral lines, and a spot on the breast white.
Upperside of wings dark hair-brown, slightly pinkish out-
side the discal band, yellow at the outer margin; eye-spots
as in Antinori, but larger. Fore wing with a white, subbasal,
angle-shaped band and a narrow white discal band, the latter
being nearly straight from costal margin to middle and then
slightly incurved ; apex of wing red. On hind wing a white
subbasal and a white discal band, joining each other in front.
Underside washed with clay-colour, the disk of both wings
densely dusted with white outside the discal band ; the latter
on hind wing continued along costal margin to base,
Length of fore wing 70 mm.
Uluguru Mts., German East Africa; one female in the
Tring Museum.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 1
2 Hon. L. W. Rothschild on
2. Epiphora rectifascia, sp. n
Body blackish brown; antenna, head, and legs clay-
colour; edges of the abdominal segments yellow; metanotum
white.
Upperside of wings dark hair-brown, the distal margin
olive, preceded by yellow spots. Fore wing falcate, a sub-
basal band white, forming an angle of 90° ; eye-spot as in
mythimnia, but longer ; a straight white discal band, sharply
defined ; outside the same and along costal margin white
dusting. Hind wing with a basal and a discal band, both
white and slightly curved; eye-spot large, incurved between
the first and second radials ; disk outside the band pinkish,
as on fore wing, and dusted with white.
Length of fore wing 80 mm.
Yakusu, Stanley Falls, Congo (J. L. Roger) ; one male in
the Tring Museum.
3. Imbrasia ethiops, sp. n.
Nearest to J. rubrocostalis, Kirby (1892).
Upperside of head, thorax, and wings brownish black;
abdomen fulvous. Fore wing with a trace of a grey discal
line ending close to apex in a small costal patch; ocellus
represented by a transparent semicrescent, which is larger
than in rubrocostalis, but very much smaller than in Mitfordi,
Kirby (1892), being thinly edged with fulvous. Costal area
of hind wing rosy red from base to middle and backwards as
fay as anterior edge of cell; ocellus consisting of a deep drab
outer ring, followed internally by a black ring which encircles
an ochraceous iris, bearing a small transparent pupil; no
discal line.
Underside paler than upper, outer margin of fore wing
and the whole hind wing shaded with erey ; on disk of fore
wing near margin a straicht but indistinct blackish band, the
hind wing bearing a likewise blackish shadowy median band
and a discal line, “the latter reaching costal margin close to
angle.
Cameroons ; one male in the Tring Museum.
4, Imbrasia eytherea lucida, subsp. n.
Smaller than South-African cytherea. Resembling yellow
specimens of the same, but the head, collar, metanotum, and
breast purer rosy red, the basal half or two thirds of the
underside washed with rosy red, the brown bands and the
ocelli edged with dark grey-brown, which is usually almost
new African Saturniide. 3
of the same colour as the brown discal line, instead of being
nearly white as in cytherea, the ocelli almost the same size on
fore and hind wing, being much smaller on the hind wing
than in cytherea.
Langenburg, Lake Nyassa, German East Africa; a series
in the Tring Museum.
5. Imbrasia anthina xanthomma, subsp. n.
Ochraceous or red-brown. Fore wing more strongly falcate
than in anthina, the discal band nearer the margin, the yellow
ocellus without transparent centre, or the transparent spot
minute; the grey submarginal area of the hind wing more
extended, being on the underside also less dentate.
Gold Coast and Sierra Leone; several males and one
female in the Tring Museum; a female from T'afo, Gold
Coast, in the British Museum.
6. Imbrasia Gueinzit nyassana, subsp. n.
Smaller than Guecnzii, less uniformly yellow; the sub-
median band of the fore wing more strongly zigzag, the
discal band costally further from the apex, therefore less
oblique ; the ocellus smaller, being subtriangular, a distinct
but shadowy grey submarginal band.
Hind wing more extended rosy red than in Guernzdi, the
base of the abdomen being also red. On the underside the
area between the discal band of the fore wing and the margin
costally wider and posteriorly narrower than in G'uecnz77, and
the grey marginal band prolonged to costal margin. On the
hind wing the distal marginal area posteriorly less widened
than in Guetnziz; the transparent spot of both wings smaller
and less distinctly edged with yellow.
A series in the Tring Museum from Langenburg, Lake
Nyassa, German Hast Africa.
7. Imbrasia Wahlbergi rhodina, subsp. n.
Body and wings varying from tawny ochraceous to reddish
tawny, the ground-colour being much deeper than in Wahi-
bergi ; the costal area of the hind wing above and sometimes
nearly the whole underside more or less distinctly flushed
with rosy red. Ocellus of fore wing smaller than in Wahlberg?,
proximally less strongly rounded than on the outer side, the
pinkish basal patch smaller, the submedian band less sharply
dentate, and the pinkish-grey submarginal band much less
prominent, as is also the case on the hind wing. 4
4 Hon. L. W. Rothschild on
Kigonsera, N.E. of Lake Nyassa, German East Africa ; a
small series in the Tring Museum.
8. Aurivillius triramis, sp. n.
The first subcostal of fore wing from the cell. Antenna
of male much smaller than in A. arata, Westw. (1849), the
branches shorter and therefore stiffer, the apical ventral
branches of the central segments about as long as the next
two segments together, being much shorter than the proximal
branches, the dorsal apical branches also shorter than the
proximal ones of the same segments. In the female the
segments shorter than in arata, the branches, which are
longer, being therefore closer together,
Smaller than West-African specimens of arata. The
pattern almost exactly the same as in yellow individuals of
arata. The tuft at the base of the antenna and the sides of
the breast more yellowish ; the second discal line of fore wing
thinner and more distinctly dentate; the marginal area less
marked with brown.
Dorsal lobe of anal segment of male narrower than in arata,
bearing a dentate crest beneath. Apical lobe of clasper long.
Penis-funnel with a black dentate club on each side.
Sierra Leone, southward to the Kassai River and Angola.
A, arata occurs also in these districts, but is apparently
less frequent.
9. Gonimbrasia belina occidentalis, subsp. n.
Nearest to G. bd. ostris, Druce (1896). Outer margin of
fore wing in male more incurved and in female more straight,
being usually slightly emarginate. The white scaling along
the lines of both wings less dense, this white border to the
discal line of fore wing edged by a fuscous or red line. The
outer margin of hind wing in male more angulate below
centre and the discal line closer to margin; its white border
usually disappearing posteriorly. The ocellus of underside
of fore wing not edged with fuscous. The anal tergite of
male slenderer than in ostris and the processes of the penis-
sheath obtuse.
Gambaga, Volta River, Gold Coast Hinterland (Dr. Bury) ;
a long series in the Tring Museum.
10. Gonimbrasia melanoneura, sp. n.
Nearest to G. acetes, Westw. (1849). Antenna more
shortly pectinated, and only 15 segments bearing side-
branches, about 16 distal segments being without them.
new African Saturniide. 4)
Upperside of wings deeper ochraceous than in acetes, the
distal marginal area narrower, the discal line, especially on
the fore wing, standing nearer the margin; the veins black
from middle of fore wing to outer margin ; the marginal area
of fore wing shaded with fuscous both above and beneath,
except at hind angle.
On underside there is an olivaceous black double spot at
apex of cell on both wings, the inner spot being large and
the outer one small. The clasper (of male) obtuse and the
anal tergite flattened, not compressed as in acetes.
Wassaw district, Gold Coast; one male in the Tring
Museum. Another male, also from the Gold Coast, in the
Leeds Museum ; acetes occurring there too.
11. Lobobunea Morland?, sp. n.
As large as L. alinda, Drury (1780).
Upperside of body and wings maize-yellow, the distal
margin except fringe white in male and fuscous in female.
On tore wing a blackish band just outside apex of cell and in
female a lunate band on the disk nearer the margin than the
cell. Hind wing without bands on upperside in male, there
being two bands in female, one from the ocellus to the abdo-
minal margin, the other about halfway between cell and outer
margin.
On underside the discal lunules in female further away
from the cell than in alinda and the marginal area in both
sexes evenly grey, being without brownish blotches ; the
central line of lunules crossing the wing at tip of cell, being
continuous with the olivaceous black central patch, these
lunules large.
Pemba Island, Hast Africa (4. Morland) ; one pair in the
Tring Museum.
12. Melanocera parva, sp. n.
A little over half the size of MZ. mentppe, Westw. (1849).
Body not washed with white on underside; head and pro-
thorax olivaceous brown, there being no white edge to the
collar.
Upperside of. wings as red as in menippe, hardly at all
speckled with brown along the outer margin; eye-spots as in
menippe, but the white lines absent from both wings.
Underside of fore wing pale red, slightly irrorated with
brown before outer margin; an extremely faint trace of a
discal line. Hind wing less clouded with brown than in
6 Hon. L. W. Rothschild on
menitppe, bearing as in that species on disk a straight white
line bordered outwardly with brown.
Length of fore wing 42 mm.
Clasper bisinuate at apex, the upper angle projecting ;
anal tergite with two acute processes. The segments of the
antenna longer than broad, the branches longer than in
menippe.
Caiala, near Bihé, Angola, 12th October, 1904 (Dr. W. J.
Ansorge), one male; and another male, also from Bihé: both
in the Tring Museum.
PARUSTA, gen. nov.
g. Antenna cylindrical, flattened at apex, the segments
much broader than long, except the last two, each with only
one pair of branches, these long and very slender, being short,
however, on the last three segments ; the apical ventral edge
of the distal segments slightly raised, but not produced into
a process as in the case of Usta and Pseudantherea, Tibiex
normal, without apical thorn-like claws, mid-tibia longer than
the first tarsal segment. Neuration similar to that of Tago-
ropsis; fore wing with four subcostals, the first more distal
than the fourth, the second very short; first radial and first
median at equal distances from the cross-vein ; costal margin
of hind wing nearly straight, the second median much more
proximal than the subcostal.
Nearest to Tugoropsis, differing especially in the structure
of the antenna and in neuration.
13. Parusta wanthops, sp. n.
Cream-buff ; the head ochraceous ; the antenna deep buff
and the abdominal segments brown at the bases and ochra-
ceous at the apical edges. Wings dirty white ; on fore wing
a patch at base, a broad curved antemedian line continued to
base at hind margin, a discal line excurved in centre and there
widened to form a patch, partly encircling the ocellus, further
an ill-defined marginal band and a portion of the veins
brown; the ocellus round, yellow, edged with brownish
black. On hind wing a discal and a submarginal band, both
ill-defined and nearly parallel to outer margin ; ocellus as on
fore wing, but smaller.
Underside nearly as upper; discal band of hind wing
distinct, widened at the ocellus, submarginal band represented
by some ill-defined patches, namely, a costal, a central, and
an anal one, the last two being joined together by a faint
new African Saturniide. iG
line. The eighth ventral ring of abdomen sinuate, the angles
being acuminate.
Length of fore wing 27 mm.
Mpwapwa, German East Africa; one male in the Tring
Museum.
14. Tagoropsis sabulosa, sp. n.
Buff, the head and upperside of the legs being brown-red.
Wings almost uniformly irrorated with brown-red scales,
which give the wings a pinkish-buff appearance. On fore
wing the first and second lines thin, placed asin 7’. flavinata ;
ocellus small, composed of a vitreous dot encircled by a
brown-red line ; the discal line curved gently but distinctly
towards apex of wing, being costally much more distal than
in I’. flavinata; apex of wing produced; no submarginal
spots, but the reddish scaling slightly condensed at the outer
side of the discal line. On hind wing two lines, the first
placed as in 7. flavinata, the second costally excurved, poste-
riorly incurved; the anal angle more prominent than in
7. flavinata, the distal margin being less rounded.
Underside like upper ; the first line of fore wing absent.
The anal tergite black, divided, each lobe compressed.
jlasper excised ventrally before apex, the upper lobe pro-
jecting, the dorsal edge of clasper biemarginate.
Length of fore wing 46 mm.
Moshi, Kilimandjaro ; one male in the Tring Museum.
15. Tagoropsis lupina, sp. n.
Ochraceous buff; palpus and fore coxe vinaceous rufous ;
upperside of legsrosy. Fore wing with three brown-red lines,
the first as in 7. flavinata, broken at median vein, the second
straight across the ocellus, very slightly dentate on the
proximal side; the third line also straight, commencing at
costal margin 6 mm. from the apex, meeting the second line
at inner margin; ocellus composed of a round vitreous spot
edged with brown-red. On hind wing a broad evenly curved
median line and a vestige of an undulate discal line.
Underside washed with brown-red except the posterior area
of the fore wing; the lines as above, but the first line of the
fore wing absent.
Length of fore wing 55 mm.
Lokolele, Congo; one female in the Tring Museum.
8 Hon. L. W. Rothschild on
16. Pseudaphelia apollinaris barotsina, subsp. nv.
Like apollinaris, Boisd. (1847), but the discal line absent
from the hind wing.
Barotseland and west side of Lake Nyassa; several
specimens in the ‘Tring Museum.
17. Cyrtogone Ansorgei, sp. n.
In colour resembling the male of C. nemia, Westw. (1849).
Fore wing shorter and broader, non-falcate, non-dentate,
the outer margin slightly incurved. Hind wing red at base
only; costal margin longer than in nemia, outer margin very
faintly dentate.
Underside more uniformly irrorated than in nemia; fore
wing with trace of a postdiscal line; extreme base of hind
wing red; tarsi partly also red.
Caiala, near Bihé, Angola, lst December, 1904 (Dr. W. J.
Ansorge) ; one male in the Tring Museum.
18. Cyrtogone bilineata, sp. n.
Similar to C. Ansorge?, but approaching nem7a in the shape
of the wings, both wings being more distinctly dentate than
in Ansorgei and the fore wing more falcate.
Upperside of fore wing with two lines as in Ansorgez, but
the discal line more distinct, especially towards costal margin ;
between this line and the yellowish-grey blotch which stands
at the apex of the cell there is a black shadowy patch ex-
tending forward to the costal margin; the outer area more
plainly irrorated with blackish brown, as is also the case on
the hind wing; the costal area of the latter more extended
red.
On underside the disk of the fore wing and the outer area
of the hind wing lighter in colour than in Ansorgei; the fore
wing with a blackish cloud outside the apex of the cell and
the discal line ending in a black spot at the costal margin.
Female similar to the male, but the wings much broader,
the fore wing without a distinct black cloud on the disk, and
the upperside of the hind wing as well as the underside of
both wings, the distal margin of the fore wing excepted,
much paler. ‘There are no white patches, as is the case in
C. nemia.
Kidugala, German Hast Africa; one male and two females
in the Tring Museum ; a pair from Salisbury, Mashonaland
(G. A. K. Marshall), in the British Museum.
Perhaps a geographical representative of Ansorget.
new African Saturniide. 9
19. Holocera suavis, sp. n.
Deeper red-brown than //. smilax, Westw. (1849).
Fore wing broader, less produced at apex, the central band
posteriorly wider, its grey outer border much less curved,
being almost straight from costa to middle; the glass-spot
consists of a half-ring, to which is joined in front an angle-
or inversed comma-shaped spot. Hind wing broader than in
HM, smilax, shorter at abdominal margin, the distal margin
slightly convex ; the eye-spot as in the species of Ludia,
yellow, with a thin, black-edged, central half-ring and a
black outer border; a pinkish-grey submarginal line extends
from close to anal angle forward to the eye-spot, encircling
the same and running back to the abdominal margin, this
last portion being faint ; this line forms the border of a red-
brown band as in Ludia.
On underside the fore wing washed with grey on the disk
outside the vitreous spot from the costal to the abdominal
margin ; some specks at the costal edge and a thin apical
half-ring grey ; outer margin, the cell, and part of costal
margin olivaceous walnut-brown; on hind wing the basal
costal area, a short costal discal band, and the outer margin
the same colour, this costal band continued to hind margin as
a very thin line.
In female the wings much broader than in the male.
Highth abdominal segment (of male) without the apical
spines found in /7. smilax.
Larva yellow; the head, prothorax, a ventral lateral stripe,
the anus, and on each ring a narrow belt bearing setiferous
warts, black, the anterior belts more or less interruped, none
of the belts reaching down to the legs.
Neuelo, Usambara, German Hast Africa ; several specimens
in the Tring Museum ; also in the British Museum from the
same place.
20. Ludia goniata, sp. n.
Similar to Delegorgue?, Boisd. (1847).
Fore wing somewhat longer at inner margin, the apex
rather more produced, the distal margin dentate, the brown
discal area in male wider than in Delegorguet, its outer edge
less curved in front and behind, slightly crenulate. Hind
wing anteriorly longer, the wing therefore larger, distal
margin dentate and below middle distinctly angulate; the
outer edge of the brown discal area also angulate, though
slightly.
10 Mr. P. Cameron on Hymenoptera
On underside the marginal area of fore wing in male nearly
as deep brown as the disk, with a thin grey submarginal
line; in female the whitish submarginal shade of both wings
more restricted than in Delegorgue?, and the brown submar-
ginal line more irregular, being slightly angulate below
middle.
Natal and Grahamstown; several males and females in
the Tring Museum and the British Museum.
21. Ludia eximia, sp. n.
Also closely resembling L. Delegorquet.
Body and wings more blackish brown; basal, costal, and
outer marginal areas shaded with grey, not clay ; submarginal
line on both wings posteriorly a little less close to edge than
in Delegorguei, the margin of fore wing densely shaded with
blackish above and beneath; costal margin of hind wing
longer than in Delegorgued, the wing being less triangular.
Ventral lobe of clasper (of male) short and blunt, the apical
lobe rounded off, not produced into a thorn-like process.
Kampala, Uganda; one male in the Tring Museum.
II.—A Contribution to the Knowledge of the Hymenoptera
of the Oriental Zoological Region. By P. CAMERON.
For the Indian species described in this paper I am indebted
to Col. C. T. Bingham; those from Borneo were sent me by
Mr. John Hewitt, the Curator of the Sarawak Museum. I
have also described a few species taken by Col. Nurse in
Baluchistan and Simla.
Braconidae.
Iphiaulax Preyert, sp. n.
Black ; the head, antennal scape, thorax, and the four ante-
rior legs red, the middle tibie and tarsi darker coloured ; the
wings fuscous, the nervures and stigma black; the head,
metathorax, and legs covered densely with longish blackish
pubescence, the ovipositor densely with stiff black hair;
the first abdominal segment suffused with rufous, the raised
central part almost smooth, the middle with a narrow keel
down the centre; the second segment closely but not very
strongly striated, the keel narrow, extending to the apex,
the base with a small smooth triangular plate; the third is
from the Oriental Zoological Region. It
striated at the base, and there is a triangular striated part on
the sides, the apex extending to the base of the apical fourth ;
there is a keel down the middle of the basal three fourths.
Temples as long as the top of the eyes, rounded behind, the
occiput almost transverse. Face coarsely rugosely punctured.
Antennal scape about three times longer than wide, of equal
width, hardly as long as the second “joint of flagellum, not
hollowed below. Parapsidal furrows shallow. Recurrent
nervure interstitial; the third abscissa of radius as long as
the basal two united.
Length 10, terebra 17 mm.
Kuching ; March (John Hewitt).
In size and form this species resembles J. trichiothecus,
Cam.; the latter may be known by the third and fourth
abdominal segments being striated. J. extraneus, Cam., has
the same abdominal striation, but isa much larger and stouter
species, and may be readily separated by the different form of
the abdominal plate, which is much longer, gradually nar-
rowed to a point near the middle of the segment.
Iphiaulax erlophorus, sp. 0.
Black ; the head (except the sides and top of front) and the
vertex (except the sides behind, and the hinder central part
more narrowly, which are black) dark rufous; the antennal
flagellum black; the prothorax bright red; the depression
bordering the basal plate of second abdominal segment, its
apex broadly in the centre, narrowly on the sides, the centre
of the third and fourth segments broadly at the base and apex,
more narrowly in the middle, and the greater part of the
ventral surface, rufo-testaceous. Median segment and breast
thickly covered with white woolly pubescence, that on the
latter shorter than on the former. The fore legs and the
middle knees broadly rufo-testaceous. Wings hyaline, very
iridescent, slightly tinged with fuscous ; the costa, stigma,
and nervures deep black. 9.
Length 7, terebra 7 mm.
Kuching; October (John Hewitt).
Abdominal segments to near the apex of the fourth finely
closely striated ; the first segment broad at the base, raised,
the sides depressed ; the area on the base of second not very
clearly defined, striated, narrowed gradually towards the apex,
which extends to near the end of the segment; the sur-
rounding parts are much more strongly striated ; there is a
curved furrow on the sides of the third segment and a trans-
verse crenulated one on the apex. Tface “covered with white
12 Mr. P. Cameron on L/ymenoptera
pubescence, smooth; the oral region bordered laterally by
deep wide furrows. There is a distinct malar space about
two thirds as long as the antennal scape. Temples wide,
only slightly narrowed behind. Parapsidal furrows narrow,
deep, clearly defined, reaching from the base to the apex.
The third abscissa of the radius fully as long as the basal two
united ; the recurrent nervure received distinctly before the
transverse cubital.
Iphiaulax cosmiothecus, sp. n.
Black ; the head, palpi, mandibles (except at apex), pro-
thorax, mesothorax, and metathorax (except the sides of
metanotum broadly) red; the fore legs red, darker at the
apex. Wings uniformly fuscous violaceous; the second
abscissa of radius longer than the third; the legs and sheaths
of ovipositor densely covered with stiff black hair; the apex
of the sheaths of the ovipositor with a broad band of white
hair. On the apical three fourths of the raised part of the
first abdominal segment are two stout parallel keels, forming
almost a deep furrow; they are united near the base by a
weak transverse keel ; on the sides are two keels, the outer
(and shorter one) commencing at the base, the other shortly
beyond it; the basal half of the bordering furrows is stoutly
closely striated ; the second, third, and basal half of fourth
segments are closely stoutly striated ; the central keel on
the second extends to the apex, it becoming gradually
narrowed until it is more than twice longer than it is wide at
the base, where it is smooth, the rest being closely longitu-
dinally striated; the sides of the segment are smooth,
shining, triangular; the central area on the outer side bears
two irregular longitudinal keels ; the inner sides bear a few
irregular transverse ones. ‘The first transverse furrow has
the central third stoutly crenulated ; on either side is a large
smooth area, longer than wide, the inner side becoming
gradually roundly curved from the base to the apex ; there
is a somewhat similar but smaller and semicircular area on
the sides of the fourth segment. ‘There is a smooth, raised,
mitre-shaped space in the centre of the face; the sides are
sparsely punctured, the punctures with hairs. The parts
bordering the centre of the front are raised. Metanotum
covered sparsely with long black hair. ‘Temples nearly as
long as the top of the eyes, rounded, not much narrowed
behind. 9?.
Body 13, ovipositor 17 mm.
This species is not unlike J. reticulatus, Cam., which may
from the Oriental Zoological Region. 13
be separated from the present by the first abdominal segment
being flat, not angled, it having also only a single narrow
keel down the centre; the areaon the base of second segment
is an equilateral triangle, small, the fourth seement is striated
to the apex, and the raised central part of the face forms a
large triangle. The central lobe of mesonotum, too, is
flatter, especially at the base.
Kuching, Borneo.
OpuionInz.
Zachresta nigromaculata, sp. n.
Yellow, with the following black spots :—the ocelli, a large
mark in the centre of occiput gradually obliquely widened
below ; a broad line on the basal two thirds of mesonotum in
the centre, slightly dilated and rounded at the apex; a line on
the sides, half in front and half behind the tegule, the two
united at the scutellum by a transverse line ; two semicircular
marks on the base of metanotum ; a mark below the tegule,
narrow and square above, gradually roundly widened below,
where it is transverse ; an oblique mark, rounded at base and
apex, on the mesopleure at the apex close to the sternum; a
broad band on the narrowed base of postpetiole, a broad one
on the middle of second segment, a triangular incision on
its base, the apex transverse; the base of the others broadly,
with a spot on their sides; and a mark on the middle of the
third to fifth ventral segments. Antenne black, the scape
yellow below. Legs coloured like the body, a broad mark on
the base of the hind cox below, the basal joint of the hind
trochanters (except a narrow stripe in the middle above), a
stripe on the outer side of the femora at the base, their base
narrowly, and apex and the tarsi (except the basal half of the
metatarsus), black. Wings hyaline, the nervures and stigma
black ; the areolet minute, with a long pedicle, receiving the
recurrent nervure in the apex; the transverse median nervure
received clearly beyond the transverse basal; the radius and
cubitus in hind wings very faint; the posterior nervures
obsolete. ?.
Length 9, terebra 1 mm.
Sikkim.
Tibiz sparsely, the tarsi more thickly spinose ; the calcaria
long, about three fourths of the length of metatarsus. Head
and thorax closely, somewhat strongly punctured and covered
with white pubescence; the metanotum almost rugosely
punctured ; the apical slope with a broad depression in the
14 Mr. P. Cameron on //ymenoptera
middle. First abdominal segment nearly as long as the
second and third united, the apex distinctly dilated; the
following segments are not compressed, distinctly broad
compared with those of Campoplew.
I refer this species to Zachresta because the spurs, although
longer than usual, are still distinctly shorter than the meta-
tarsus ; in the allied genus Echthronomus, Foer., they are
described as “ very long, scarcely sharter” than it. The
only difference between this species and Campoplea of much
importance lies in the abdomen not being strongly compressed,
the latter genus having also the metanotum longer and more
or less areolated,
Anomalon Bingham, sp. n.
Black ; the face, clypeus, mandibles, malar space, a broad
line on the lower third of the inner orbits, tegulee, and under-
side of antennal scape lemon-yellow; the abdomen (except
the top of the basal two segments) rufo-ferruginous; the four
front legs fulvous, tinged with yellow, their coxe and _ tro-
chanters yellow; the hinder legs rufous, the coxe# and
trochanters black above, the former broadly yellow in the
middle below, the apical third of the tibize black ; the apical
joints of the tarsi tinged with yellow. Wings hyaline, tinged
with fulvous, highly iridescent; the stigma testaceous, the
costa and nervures black. <@.
Length 22 mm.
Sikkim.
Head covered with white hair; the face strongly punc-
tured, the punctures large, clearly separated; the front
stoutly obliquely reticulated; the vertex strongly deeply
punctured. Mesonotum strongly closely punctured, the
punctures running more or less into each other, the apex
stoutly transversely striated. Scutellum roughly reticulated,
depressed broadly in the centre. Metanotum much more
widely reticulated, slightly broadly depressed in the middle,
as are also the metapleure. Propleure stoutly striated,
obliquely above, longitudinally below; the upper part
distinctly punctured. Mesopleure above longitudinally
coarsely reticulated; the apex smooth, bare, below the reticu-
lated part, the rest punctured, the punctures clearly separated,
each having a black hair. Flagellum of antenne rufo-
testaceous, the basal half black above.
Allied to A. tinctipenne, Cam., from the Khasias. The
present species may be known from it by the stoutly reticu-
lated front: in tinctipenne it is striated rather than reticulated,
from the Oriental Zoological Region. V5
the keels, too, being much finer; the yellow mark on the
inner orbits is obliquely narrowed above, not of uniform
thickness, and there is a similar mark above it, which is not
the case with the present species; the puncturation on the
mesonotum is finer and closer and does not run into reticu-
lations and striz at the base; the transverse cubital nervure
is as long as the basal abscissa of the cubitus, while in téne/-
penne it is shorter than it,
Pruprin”.
Rhyssa varilineata, sp. n.
Black, with the following yellow spots:—underside of
scape, face, the orbits broadly (except on top), apex of pro-
notum (its sides more narrowly), scutellar keels, scutellum
(except at apex), median segment (except round the edges),
tubercles, an irregular spot below them, a smaller line under
hind wings, a line down the centre of first abdominal segment
(narrowed in the middle, the apex wider than the base), a
broader line on the apical three fourths of the second (its
apex rounded and dilated), a continuous line near the apex of
the third, an interrupted one on the fourth and fifth, and a
narrower obscure one on the sixth. Legs yellow, the four
anterior cox above, the hinder (except for a broad band
above), the greater part of the femora behind and the poste-
rior in front in the middle, the base of the four front tibia,
the posterior to beyond the middle, and the tarsi, black.
Wings hyaline, iridescent, the stigma testaceous, the nervures
black ; a squarish cloud at the end of the stigma, extending
shortly beyond the areolet. 9.
Length 13, ovipositor 12 mm.
Kuching; September (John Hewitt).
Smooth, except the striated mesonotum and the scutellum,
which is punctured. Pedicle of areolet as long as the straight
basal branch. Clypeus narrowed obliquely, testaceous, its
middle dilated, almost toothed.
This is the smallest Malay species known. It is near to
maculipennis, Smith, which may be separated from it, apart
from its much larger size, by the much larger alar cloud,
which extends beyond the cubitus, and by the metanotum
having three separated yellow spots on the apex, the rest
being black.
Rhyssa claripennis, sp. n.
Black ; the head (except the centre of front), the vertex
16 Mr. P. Cameron on Hymenoptera
and occiput, the mesonotum (except for two lines, rounded
on inner side, in the centre), scutellum (except at apex),
postscutellum, the keels, metanotum, the pleure (except
the propleuree broadly in the centre), the first abdominal
segment broadly in the centre, a narrower stripe down the
middle of the second (widest at the apex, not touching the
base or apex of the segment), and transverse lines before
the apices of the others, the apical two interrupted, and the
ventral surface, pale yellow. Legs pale yellow, the fore
femora behind, the middle above on the apical two thirds, the
posterior entirely above, the base of the tibiee, and the tarsi
black. Wings hyaline, the costa and nervures black, the
stigma fulvous ; pedicle of areolet shorter than lower branches,
the recurrent nervure interstitial. 9.
Length 18, terebra 21 mm.
Kuching ; November (John Hewitt).
Face closely but not coarsely punctured, thickly covered
with short white pubescence. Basal and apical slopes of
mesonotum smooth; the striation strongest in front. Clypeus
depressed, yellow tinged with rufous.
May be known from the described Indo-Malay and
Indian species by the immaculate wings.
Epirhyssa nigrobalteata, Cam.
The markings on this Bornean species vary somewhat,
e.g. the yellow’ mark on the mesonotum may be longer than
wide, with a triangular incision on the apex.
Epirhyssa curvimaculata, sp. n.
Black; the face, clypeus, inner orbits to the ocelli, the outer
narrowly above, broadly below, prothorax (except for a broad
band on the pleure), a pyriform spot on the sides of middle
lobe of mesonotum, a longer one on the inner side of the
Jateral lobes of mesonotum, straight on outer side, rounded
on inner and produced into a sharp point at the apex, scutellar
keels, basal two thirds of scutellum, metanotum (except for a
semicircular mark on apical half), an oblong mark below the
tegule, united to an oblique large mark below it, extending
from the base to the apex, the metapleurse (except at the
base, the black basal spot having the part above the keel
larger than that below it), broad bands (rounded at the base)
on the basal two abdominal segments, the lines on the
following three broad and curved ‘backwards in the middle,
those on the following two broad, curved and widely inter-
rupted in the middle, and the basal ventral segments, pale
from the Oriental Zoological Region. 1%
yellow. Legs pale yellow, the inner side of the four hinder
coxe, the lower side of the posterior, base of four hinder
trochanters, the four hinder femora broadly above, tlie hinder
two also in the middle below, base of tibie and the tarsi, black.
Wings hyaline, a lage fuscous-violaceous cloud at the apex,
commencing shortly beyond the transverse cubital nervure
and extending to the end of the radial cellule. ¢.
Length 15, ovipositor 28 mm.
Kuching ; September (John Hewitt).
Face smooth, finely obscurely striated above ; the stria on
apical central part of mesonotum much finer and closer than
on the rest. Basal half of scutellum closely punctured, as
are also (but more finely) the middle segments of the
abdomen.
In the male the meso- and metapleure are yellow, without
black, as is also the breast, the black on first abdominal
segment is transverse at the apex, the bands are broader, and
the apical segments are reddish brown; the black on the
legs is less extended and the apical cloud in the fore wings
is small, not extending to the cubitus.
This species comes near to EH. (Macrogaster) flavopictus,
Sm., described from Singapore by Smith and from Borneo
by Tosquinet (Mém. de la Soc. ent. de Belg. 1903, p. 258).
It is a much larger species (25 mm. according to ‘losquinet,
15 lines according to Smith). Macrogaster, Brullé, does not
belong to the Pimpline ; Ctenotoma, Cam., is probably the
same. It is purely African, so far as I know.
Epirhyssa cruciata, sp. n.
Black ; the apex of the second and the following abdo-
minal segments reddish brown; the face, clypeus, inner orbits
to near the end of the ocelli, the outer (except for a broad
stripe in the centre of upper third), a large mark (dilated
broadly at the apex below) on propleure, two small oval
spots on the centre of mesonotum at the base, two larger ones
(roundly narrowed on the inner side) at the end of the basal
lobe, a square mark on the sides of the scutellum at the base,
a longer narrower line on the sides of the apex (leaving a
black cruciform mark in the centre), a broad curved band on
the metanotum near the apex uniting with the yellow on
metapleure which occupies almost the whole surface, the
tubercles, the mesopleuree (except round the edges and a large
oblique mark, narrowed above, near the centre below), a large
triangular mark on the sides of mesosternum, a longish line
(dilated at the apex) in the centre of apical three fourths of
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 2
18 Mr. P. Cameron on L/ymeneptera
first abdominal segment, a mark (roundty narrowed behind)
near the apex of the second, a curved line shortly beyond the
middle of the third, two oblique lines in the centre of fourth,
two more widely separated curved ones on the fifth, and more
or less of the ventral surface, yellow. Legs yellow, the four
anterior femora broadly dark rufous above, the hinder pair of
a darkersrufous on the inner and outer side; the four hind
tibia broadly black at the base above, the apex black all
round, and the four hind tarsi black. Wings hyaline, the
apex lightly smoky, the costa and nervures black, the stigma
rufo-testaceous; the recurrent nervure interstitial. @.
Length 18 mm.
Kuching; January (Hewitt).
Face closely distinctly punctured and thickly covered with
white pubescence, the front and vertex smooth. Underside
of antennal scape yellow. Mesonotum striated throughout,
but more weakly towards the apex. Base of scutellum
strongly, the apex more finely and closely striated. Meta-
notum closely punctured, the punctures clearly separated ; at
the end of the central furrow are three or four stout curved
strive.
Epirhyssa lineatiscutis, sp. n.
Length 15mm. ¢.
Kuching ; September (/ewztt).
This species is very similar to L. cruciata, described above ;
the two may be separated thus :
Mesosternum entirely black; mesopleuree black with a
large curved yellow mark below the tubercles and a
smaller, triangular one at the apex below; the mark
on first abdominal segment not much longer than it
is wide at the apex, narrowed at the base, that on
the second triangular, as long as the width at the
apex, the lines on the third and fourth wide, not
Curved, COntINUOUS factor. ak ue citi eet ree eee lineatiscutis.
Mesgosternum black, except for a triangular lateral spot ;
mesopleurs yellow, with an oblique black mark at
the base ; the mark on first abdominal segment long,
narrow, on the second narrow, transverse, the lines
on the third and fourth narrow, not continuous .. eruciata.
The legs are more largely suffused with rufous than in
cruciata, the four hind tibie much darker coloured, the hind
coxee black, yellow above, the line down the centre of
scutellum is narrower than the apical which curves round the
apex, the black band on base of metanotum occupies the
basal fourth and is of equal width, the curved black apical
mark is wider at the apex than it is long, the metapleure
from the Oriental Zoological Region. 19
black round the base, apex, and lower edge ; the four lines on
mesonotum are longish, narrow. Basal three segments black,
except for the yellow marks, the others brownish red. Except
that the black line uniting the eyes with the black on vertex
is wider, the head is coloured as in cruciata. As in that
species, the apex of the wings is narrowly smoky. The
yellow transverse lines on the third, fourth, and fifth seements
are wide, that on the third dilated in the middle at the base.
Xanthopimpla lissonota, sp. n.
Pale yellow, a triangular mark covering the ocelli, a semi-
circular mark, the rounded narrowed end at the apex, on the
centre of middle lobe of mesonotum, a mark widened in the
middle at the base of scutellum, a minute spot on either side
of the base of metanotum in the lateral areze, and two spots
on the seven basal abdominal segments, those on the first
produced into a narrowed line at the base, a mark on the
outer and inner side of the trochanters, and a line on the base
of the hinder tibie, black; the basal joint of the hind tarsi
and the apical two infuscated. Wings hyaline, the apex
shghtly smoky, the nervures and stigma black ; the areolet
small, shortly appendiculated. Flagellum black. @.
Length 7 mm.
Kuching ; November (/ew?#t).
Areola wider than long, six-angled, the apical keel almost
rounded ; the lateral are widened gradually in the middle
almost to a point; the sides slightly angled in the middle,
from which a keel runs towards the apex. Mesonotum
smooth, shining; the furrows short, on basal slope only.
Except the third to seventh abdominal segments the body 1s
impunctate ; the puncturation on the abdomen becomes
gradually weaker. Marks on second abdominal segment
small, irregular, on the third irregularly round, on the others
wider than long, transverse. Face equally roundly convex,
closely punctured, squarish.
Agrees best with Krieger’s section G (cf. Ber. d. naturf.
Gesel. z. Leipzig, 1897-8, p. 92).
Echthrus maculiceps, sp. n.
Black; face (except for a line over the clypeus, dilated at
the sides above), clypeus, labrum, base of mandibles, a line
round the orbits, broadest on the front, palpi, two oblique
marks in the centre of the occiput, base of pronotum, a broad
line, narrowed in front, on the sides, an irregular squarish
) %
A
20 Mr. P. Cameron on Hymenoptera
mark in the centre of the propleuree below, a longish oval
mark on the apex of the middle lobe of mesonotum, tegule,
scutellum (except at apex), a small spot on the sides near the
apex of the yellow part and united to it, scutellar keels, a
band on the base of metanotum, the black bordering its apex,
A-lobate, a large trilobate mark on apex, the central lobe
projecting backwards, smaller than the others, its base incised
in the middle, the sides rounded, a broad large mark, the
base roundly dilated, the apex straight, produced posteriorly
below, this part being also produced backwards as a broad
line to near the apex of the basal third on mesopleure, the
centre of mesosternum broadly, the mark widened towards
the apex and with a black mark in the middle of the apex,
a small mark under the hind wings, a large one almost
covering the metapleure, a broad mark on the apex of the
first abdominal segment, its basal half narrowed, rounded
and incised at the centre, narrower lines, dilated laterally, on
the apices of the second to sixth, the seventh (except for a
large trilobate mark), the apical lobe square, larger than the
lateral, and the ventral surface (except the first and large marks
on the following three segments), pale yellow. Antennze
broadly (sixth to twelfth joints) ringed with white. Legs
yellow, the anterior paler, the hinder femora fulvous ; a spot
on the outer apical side of the four anterior, the posterior
below and broadly at the apex all round, a line on the
trochanters, on the four anterior femora, the apex of the
hind femora narrowly, the base of the hind tibie slightly
more broadly, the apex of the hind tibiz more broadly, and
the apical joint of the tarsi, black. Wings hyaline, the
nervures and stigma black.
Length 19, terebra 5 mm.
Quop, Sarawak ; November (Hewitt).
Closely punctured, densely covered with short white
pubescence ; the mesopleure at the base closely striated ; the
depressed apex of middle lobe finely longitudinally striated.
Base of metanotum behind the keel closely, somewhat
strongly striated, the part between the two keels stoutly
closely striated in the middle, the sides closely reticulated,
the apex much more strongly striated, the striz curved; the
basal keel curved backwards in the middle, the apical stouter,
not curved in the middle. Parapsidal furrows distinct,
narrow, deep. The first abdominal segment broadly smooth
in the centre of narrowed basal part, this smooth line
bifurcating on the widened apex; the punctures on the
segment are stronger and more widely separated than on the
rest of the abdomen. Areolet as long as it is wide at the base,
Srom the Oriental Zoological Region. 21
the apex slightly narrower than the base, through the cubitus
turning slightly upwards from the recurrent nervure. The
transverse basal nervure is received considerably in front of
transverse median.
TRYPHONINE.
Fovaya spinipes, sp. n.
Black, the second, third, and fourth abdominal segments
red, the fourth with the apex narrowly black ; the antenne
with the thirteenth to twentieth joints white ; legs black, the
greater part of the fore femora and all the tibia reddish ;
the anterior tarsi testaceous, the four apical joints black
above ; the basal two joints of the middle tarsi testaceous, the
third whitish, the apical black ; the basal joint of the posterior
tarsi black, testaceous at the base, the second to fourth white,
the fifth black. Wings hyaline, the nervures and stigma
black ; the areolet shortly appendiculated, the basal nervure
straight, oblique, the apical rounded, the recurrent nervure
received near the apex. @.
Length 11 mm.
Simla.
Head and thorax alutaceous, opaque, covered with a pale
down; the lower half of the clypeus shining, somewhat
strongly, but not closely punctured. Apical half of meso-
pleuree smooth and shining above. On the apex of the
metanotum are three areee longer than wide, the central larger
than the lateral and all rounded above; leading from the
base to the central area are two keels which curve inwardly
before the middle, the apical half of the area being wider
than the basal. The anterior tibie sparsely, the four
posterior thickly spinose.
This species may be known from F. annulitarsis, Cam.,
the type of the genus (it is also from Simla) thus :—
Clypeus, lower inner orbits, and a spot on the sides of
mesonotum at the base pale yellow; flagellum
brownish beneath before the white band; second
abdominal segment black, red at the base, the inner
keel of spiracular area distinct ; the four hind femora
HESHACCO USE peters ne metel tel aio tehoekek- kell retere lel ch-Es) felakolial sive
Clypeus, lower inner orbits, and base of mesonotum
entirely black; flagellum black before the white
band; second abdominal segment red, the inner keel
of spiracular area indistinct ; the four hind femora
Cee ate on tet, adsueibeed. cokcdecehate: vce etal cvchatal oo otal «
annulitarsis.
spinipes.
nD
bo
Mr. P. Cameron on Hymenoptera
ss
CRYPTINA.
Silsila striolata, sp. n.
Black, a broad band on the centre of the flagellum, face,
clypeus, mandibles, palpi, a narrow line on the upper inner
orbits, a line on the upper, a broader one on the lower edge
of prothorax, the line on the top, narrowed behind, scutellums,
scutellar keels, a large, broad, triangular mark behind the
hind wings, the apical slope of metanotum, except in the
centre, the mark in the centre continued upwards to near the
transverse keel, this part becoming gradually narrowed,
from its outer edge at the apex a line of equal width runs
along the edge of the pleurze to near the spiracles and below
along the coxe, a broad curved mark on the lower edge of
the mesopleure, the base being narrower and continued
higher up than the apical, and lines of equal width on the
apices of the abdominal segments, pale yellow. Four front
legs pale fulvous, paler at the base; the fulvous tint on the
hind legs is deeper, the lower and outer side of their coxe,
the apical joint of the trochanters, the apex of the femora
narrowly, and the base of the tibiz still more narrowly, black.
Wings hyaline, the nervures and stigma black. @.
Length 12 mm.
Bidi, Borneo; July (/ewiit).
Face, except on the outer edges, closely, roughly punc-
tured, the clypeus smooth. Front stoutly, obliquely striated
below the ocelli, the lower part furrowed down the centre.
The yellow projecting part of the pronotum stoutly striated ;
the upper part of the propleures irregularly, finely striated
and punctured; the middle stoutly striated. Mesonotum
closely, distinctly punctured, finely striated on the edges of
the lobes; the scutellum smooth, its basal depression deep
with four stout keels; the mesopleure stoutly striated,
except in the middle at the apex. On the base of the
metanotum is a band of stout strize, closely pressed, but
clearly separated ; the space between them and the keel is,
in the middle, coarsely, closely shagreened, almost punctured ;
the sides have the punctures clearly separated. Metapleure
closely, rugosely punctured, the punctures running into reti-
culations, especially at the base. ‘The first abdominal segment
smooth and shining, the others opaque, aciculated.
Mesostenvideus cariniscutis, sp. n.
Black ; a line on the inner orbits, a mark, longer than
wide, narrowed above, transverse at the apex, in the centre
from the Oriental Zoological Region. 23
of the face, two large marks, almost united above, in the
centre of the clypeus, the apex of the labrum, an irregular
line on the base of the pronotum, tegule, tubercles, scutel-
Jums, scutellar keels, metanotal spines, and the apices of the
abdominal segments, yellow ; the four front legs yellow, the
femora tinged with fulvous, their tarsi fuscous; the hind
coxe and basal two thirds of the femora fulvous, the tro-
chanters, apex of femora, and almost the apical half of the
tibie, black ; the basal part of the tibize testaceous, the tarsi
white. Antennze broadly ringed with white. Wings hyaline,
the stigma and nervures black, the areolet square; the
transverse median nervure received distinctly behind the
transverse basal. 2.
Length 12, terebra 2 mm.
Sikkim.
Front irregularly transversely striated, the strise weaker
below; a narrow keel down the middle. Face finely,
irregularly reticulated, the clypeus with large scattered
punctures. Mesonotum closely minutely punctured. Scutel-
lum with large lateral keels to near the apex. Base of
metanotum finely, closely, longitudinally striated, the striz
becoming stronger towards the apex; the rest closely,
strongly reticulated ; spines broad, rounded at apex. Abdo-
men finely, closely punctured, the ‘postscutellum smooth.
Mesostenus respondens, Cam., M. brahminus, Cam., JM,
misippus, Cam., JM. clarinervis, Cam., M. reticulatus, Cam.,
M. caligatus, Cam., J. saltator, Cam., M. versatilis, Cam.,
and M. maculiceps, Cam., from the Khasias, belong to
Mesostenoideus.
Friona lineatipes, sp. n.
Black ; the face (except for a short line in the centre below
the antennze and the clypeal fovea), the clypeus (except for a
black triangular mark in the centre of apex), the orbits
broadly (except the upper half of the outer), mandibles (except
the teeth), palpi, a broad line on the pronotum, scutellums, a
large, transverse, triangular mark behind the hind wings, a
broad longitudinal mark on the middle of metanotum, com-
mencing shortly behind the keel and uniting at the apex to a
broader transverse one across the apex, tubercles, a line at
the apex of the pleural furrow, a longish mark, widest below,
in the middle of the metapleuree at “the apex, not quite the
apical half of the postpetiole, a narrower band on the apex
of the second segment, a still narrower one on the apex of
the third, slightly dilated at the middle, broadly laterally,
24 Mr. P. Cameron on Lymenoptera
where it is continued on the sides to the base of the segment,
a similar, but narrower and more irregular line on the fourth,
not dilated in the middle, a narrower one, interrupted in the
middle, on the fifth, and the sides and middle of the apical
two broadly, yellow. Four front legs pale yellow, the
femora tinged with fulvous, the middle tarsi infuscated ;
hind coxe black, with a yellow mark above, extending from
the base to shortly beyond the middle; the trochanters
fulvous, the basal joint black at the base below and entirely
above; the femora fulvous, broadly black above, more
narrowly below; the tibize yellow, tinged with fulvous, the
base narrowly, the apex more broadly black ; the tarsi white
with a narrow black line at the base. Wings hyaline, the
stigma and nervures black. Antenne broadly ringed with
white. @.
Length 13, terebra 5 mm.
Darjiling, 7000 feet; March (C. 7. Bingham).
Frout with a weak, straight, central, and three oblique
keels on either side, the inner oblique one weaker than the
outer two, the outer keel originating from the hind ocelli ;
be low, the front is depressed; the keels are weaker in the
depression. Mesonotum smooth, the apex of its middle lobe
stoutly, metanotum closely, rugosely reticulated ; the space
between the keels stoutly, longitudinally striated; on the
sides the strise run into reticulations; the basal keel is not
very distinct, being confused with the puncturation ; there
is a slight indication of an areola at the base; the second
keel is distinct, broadly roundly curved, dilated laterally.
Basal slope of scutellum with large, clearly separated
punctures, the top is more closely punctured, the punctures
running into each other; it 1s depressed in the middle at
the apex; the apical slope closely, strongly punctured.
Metanotum behind the keel closely, strongly transversely
striated. Pro- and mesopleure stoutly longitudinally
striated; the mesopleurze smooth in the middle behind
and more or less reticulated below ; the metapleuree stoutly
obliquely striated, except for an almost smooth space at the
base. ‘Ihe recurrent nervure is received shortly beyond
the middle. Pubescence on thorax not very dense and pale
in colour.
This species comes near /. frontella, Cam., and the Malay
F, varipes, Cam.
Cratocryptus annulitarsts, sp. 0.
Black ; a semicircular mark on the sides of the face
touching the eyes, a square mark in its centre, touching the
from the Ortental Zoological Region, 25
clypeus, the clypeus, a broad curved line on the inner orbits,
commencing shortly above the antennze and extending above
to the end of the eyes, a longish triangular mark (the
narrowed end above) on the outer orbits, commencing
shortly below the middle and reaching near to the middle of
the malar space, a broad mark on the base of the mandibles,
palpi, a broad line on the middle of pronotum, a line on the
lower edge of propleure, tubercles, tegulze, a line on scutellar
keels, scutellum, a broad, large mark, roundly narrowed
below, on the sides of metanotum on apical slope, and the
apices of the abdominal segments (the lines on the basal
three broad, narrower on the others), pale yellow. The
seventh to thirteenth joints of antennz white, except above.
Legs red, the hinder darker in tint, the tarsi black, the
second and third joints of the posterior white; the four
anterior cox and trochanters yellow ; the hind coxe (except
broadly on the inner side, where they are yellow, as they are
also at the apex below), the apex of femora narrowly, and the
base and apex of tibiw slightly more broadly, black. Wings
hyaline, the stigma and nervures black. ¢.
Length 10, terebra 2 mm.
Sikkim.
Face rugosely punctured, more or less reticulated in the
centre ; the clypeus strongly, sparsely punctured. Front in
the centre, to the apical depression, strongly striated, the
striae intermixing; it is longitudinally striated above,
transverse below. Mesonotum closely, finely punctured,
irregularly transversely striated along the furrows. Scutel-
lar depression wide and deep ; there are four stout keels on
it. Metanotum behind the keel stoutly, longitudinally
striated ; the rest stoutly, closely reticulated. Pleuree closely,
rugosely reticulated; the greater part of the propleure
stoutly striated. ‘The first and the apical segments of the
abdomen smooth; the middle closely punctured. Areolet
wider along the radius than along the transverse cubital
nervures, of almost equal width; the recurrent nervure is
received shortly beyond the middle.
This species has the general coloration (including the
marks on the face) of C. rufipes, Cam., from Deesa, but is
much larger ; they may be separated as follows :—
Base of metanotum punctured, with an areola; the post-
petiole strongly punctured ; meso- and metapleuree
with yellow marks; hind cox red; the hind tarsi
NOGMACWIALS WACHN WHICE! | rere « <ateisio ceeneeeltresie-e o's rufipes.
Base of metanotum striated, without an areola; the post-
petiole smooth ; meso- and metapleure immaculate ;
the hind coxe black and yellow, the hind tarsi
TALC ULCER YEN WEG ae ce ude welch spe sore oars reese annulitarsia,
26 Mr. P. Cameron on [Hymenoptera
Colganta latiscutis, sp. n.
Rufo-fulvous ; the basal six joints of antennze coloured
like the body, the seventh to twelfth pale yellow, the apical
black ; the apical joint of the middle tarsi, the apical two of
the hinder as well as the base of the second and third joints
of the hind pair black. Wings hyaline, highly iridescent,
suffused with brassy tints; the costa and stigma dark testa-
ceous, the nervures black. Median segment closely rugosely
punctured-striated-reticulated ; the two basal keels are irre-
gular; the basal abscissa weaker than the apical, its basal
part straight, the apical longer, weaker, and roundly curved
outwardly. Hye-orbits broadly pale yellow ; the front in the
middle closely rugosely punctured, raised, obliquely narrowed
to a point below. Temples almost obsolete, the eyes above
extending to the extreme edge, below they are abruptly
obliquely narrowed ; the margined edge of occiput is placed
low down below the top of the eyes. Pro- and mesopleure
closely distinctly punctured. Scutellum closely strongly
punctured, thickly covered with longish black hair; the
lateral keels extend to the middle. Mandibles pale yellow,
the apex from shortly behind the teeth black. Centre of face
slightly raised, of equal thickness, not tuberculate above.
Scutellum broad, rounded at the apex, which has a longish,
steep, vertical slope.
Length 17, terebra 3 mm.
Sikkim.
May be known from C. fulvipennis, Cam., by the middle
of the antenne being clear yellow, the antenne being distinctly
three-coloured, by the temples being much less developed
above and more sharply oblique, by the shorter ovipositor,
and by the difference in the metanotal keels: in /ulvipennis
the inner keel is straight, the apical slightly rounded back-
wards in the middle and obliquely sloped; there is a distinct
outer keel inside the spiracles, it having the basal two thirds
roundly curved inwardly, the lower third straight; in the
present species there is no outer keel and the inner is broadly
roundly curved. The difference in the form of the scutellum
is well marked ; in fwlvipennis it is clearly longer than wide
and becomes distinctly narrowed towards the apex, with a
gradual, short, not a vertical slope.
APHADNUS, gen. nov.
Areolet entirely open at apex, the only transverse cubitus
from the Oriental Zoological Region. 27
short, the recurrent nervure widely distant from it, the cubitus
between it and the transverse cubitus sharply obliquely
sloped. Antenne 30-jointed, basal three joints of flagellum
greatly elongated, the basal two of equal length. Transverse
median nervure in hind wings broken below the middle.
Hind ocelli separated from the eyes by a greater distance
.than they are from each other. Eyes very large, the temples
very short, the occiput roundly deeply incised. Metanotum
with three central aree, the petiolar a longish triangle, the
areola longish, 6-angled, obliquely narrowed at the base, the
apex transverse ; the apical area occupying the apical slope ;
there are two large lateral basal areze; spiracles small, not
much longer than wide, broadly rounded before and behind.
First abdominal segment large, longer than it is wide at the
base ; the first, second, third, and base of fourth segments
closely, regularly, longitudinally striated.
The antennee are as long as the body, slender, ringed with
white ; the wings are spotted. Parapsidal furrows distinct,
complete, the metathorax densely covered with long pubes-
cence. Radial cellule wide near the middle ; radius issuing
from middle of stigma, its first abscissa shorter than second.
Eyes higher than the vertex.
This genus cannot be confounded with any of the known
genera of Hemitelline wanting the areolet. The first abdo-
minal segment is shorter and wider than usual; noteworthy
are the striated basal abdominal segments.
Aphadnus rufipes, sp. n.
Black ; tegule, a broad band on the apex of the first
abdominal segment, a stripe in the centre of the apex of the
second, apical half of fourth, an obscure broken narrow line
on fifth, and a narrow complete one on sixth, whitish yellow.
Legs bright red, the four anterior coxe, the base of hind tibie
behind, and the hind spurs white ; the rest of the hind tibiz
and the tarsi black. Wings hyaline, a cloud (rounded and
narrowed in front) before the apex of transverse basal and trans-
verse median nervure, extending slightly beyond the latter, a
large one extending from the base to near the apex of radial
cellule, its lower half projecting backwards along the disco-
cubital nervure, and one in the apical fourth of hind wings,
almost black ; the nervuresand stigma black. Head opaque,
closely punctured, densely covered with white pubescence,
as are also the pleuree and metanotum. Face and clypeus
dilated in the centre. Middle lobe of mesonotum somewhat
28 Mr. P. Cameron on [Hymenoptera
strongly and closely striated, the outer and inner edges of the
lateral more finely striated, their centre opaque, alutaceous.
Base of metanotum irregularly transversely striated, more
finely and closely on the inner side at the base, the apical
striz interlacing, the apical slope more strongly, obliquely,
and transversely striated, the striae more or less interlacing.
Except on the smooth apex the basal segment of abdomen is
closely finely striated, as is also the basal half of the fourth ;
the striation on the second and third is stronger. Abdomen
as wide as the thorax and slightly longer than the head and
thorax united. ?.
Length 7, terebra 2 mm.
Kuching; September (J. Hewitt).
IcCHNEUMONIN2E.
Imeria levifrons, sp. n.
Black ; the entire head smooth, shining; the mesonotum
opaque, closely irregularly striated ; the propleurse almost
smooth ; the mesopleurz smooth at the apex above, the rest
closely longitudinally striated, the striae more or less twisted ;
the metapleuree smooth at the base and below, the rest more
or less striated ; the metanotum (except the enclosed space at
the base) closely reticulated, more or less striated; the
abdomen less shining than the thorax. The following are
pale yellow : face, ely) peus, mandibles, palpi, a spot outside
the lower part of the eyes, the upper inner orbits somewhat
broadly from the antennze to the ocelli; a large broad oblique
mark, obliquely narrowed on the outer side, behind the top of
the eyes; a small irregular mark outside the lower edge of
the eyes. Base of pronotum, a line on the apical half of the
sides above, tubercles, scutellum, apical slope of metanotum,
the mark roundly dilated above in the middle, a large
elongated triangular mark below the hind wings, a line about
the same length on the lower apical half of the apex of
metapleure, its base roundly narrowed, the apex of the first
abdominal segment broadly, a mark on the apex of the second
and base of third, forming united a triangular lateral mark,
narrow lines on the apices of the third to sixth all round,
and the seventh broadly, whitish yellow. Four front lees
whitish yellow, the femora tinged with fulvous, the hinder
yellow, the femora and tibie broadly black, the tarsi fuscous ;
the coxe black, rufous at the base. Wings hyaline, the
nervures and stigma black. @.
from the Oriental Zoological Region. 29
Length 7 mm.
Kuching ; September (John Hewitt).
Antenne longer than the body, the fourth to eighteenth
joints of the flagellum white. Ovipositor-sheath largely pro-
jecting, as long as the apical two joints united. Apex of
scutellum slightly rounded inwardly, the lateral keels of equal
height. Areolet 5-angled, the nervures clearly separated in
front. Apex of abdomen becoming gradually narrowed from
the middle. Transverse keels on metanotum indistinct.
Imeria, Cam., was described from the Khasia Hills, Assam,
in Ann. & Mag. Nat. Hist. xi. (7) 1903, p. 174. Its occur-
rence in Borneo is another proof that Assam belongs to the
Indo-Malay zoological region rather than to the Indian.
It has a considerable resemblance in the form of the head
and abdomen to Aulojoppa, Cam., but that genus has longi-
tudinal as well as transverse keels on the metanotum.
Imeria has only transverse keels.
ALLONOTUS, gen. nov.
Antenne ringed with white, broadly dilated as in the
Joppini. Clypeus clearly separated, raised in the middle
above, the sides above bordered by a furrow. Mesonotuin
with distinct parapsidal furrows on the basal half, the basal
half of the mesosternum bordered by a distinct, if narrow,
furrow. Scutellum keeled laterally to the top of the apex,
which has a longish oblique slope. Metanotum regularly
areolated ; the areola widely distant from the base, longish
triangular, the narrowed end at the base; there is no petiolar
area, the basal lateral arez therefore being confluent ; there
are three aree on the apical slope, the lateral keels of which
are dilated near the top. Abdominal petiole dilated at the
apex, the postpetiole not, however, separated ; there are seven
segments of equal width (except that the second is narrowed
at the base); the ovipositor has a longish sheath, which
projects obliquely over the top of the last segment, which is
bluntly pointed; gastrocceli broad, smooth, broader than the
space separating them. Legs stout, the hind coxe large,
the tarsi spinose. Areolet 5-angled, narrowed in front ;
the discocubital nervure with an indication of a stump; the
transverse median nervure interstitial Head large, the
temples broad, rounded ; the occiput deeply incised.
The thorax is long, about three times longer than wide ;
the eyes are large, parallel; there is a short malar space ;
the transverse median nervure in the hind wings broken near
30 Hymenoptera from the Oriental Zoological Region.
the bottom. The ventral keel extends to the apex of the
fourth segment. Head large, wider than the thorax. Labrum
hidden.
The edentate mandibles refer this genus to the Heresiarchini.
It cannot well be confounded with any of the known genera
in that group. The parapsidal furrows and that on the meso-
notum are quite as distinct as they are in many Cryptine ;
but in other respects the genus belongs to the Ichneumonine.
The antenne are exactly as in the Joppini. Characteristic
is the dilated at the base, clearly separated clypeus. In this
respect the clypeus shows an approach to what it is in
Chiaglas, an Kast-Indian genus belonging to the same group,
but differing considerably otherwise.
Allonotus rufus, sp. n.
Rufous ; the red on the second and following segments of
the abdomen darker, more testaceous in colour; a broad band
at the outer sides of the ocelli (continued as a narrower line
in front of them), the second abdominal segment (except at
the sides and apex), slightly more than the basal halt of the
third and fourth segments, the fifth (except narrowly at the
apex), the sixth (except for a narrow white line at the apex),
and the seventh (except for a wide white band in the middle),
black; the petiole has the bright red colour of the thorax ;
the gastrocceli pale rufous, Legs coloured like the thorax,
except the apices of the four anterior tarsi broadly, the apex
of the hind femora narrowly, and the hind tibize and tarsi,
which are black. Antennze with the scape rufous, the
flagellum black, brownish at the base below, the middle with
a broad white band. Wings hyaline, the stigma fuscous, the
nervures black. ¢@.
Length 10 mm.
Kuching ; December (Hewitt).
Face and clypeus smooth, the former pilose, the latter bare ;
the rest of the head smooth and shining. Except the meso-
notum (which is opaque and slightly shagreened), the thorax
is smooth and shining, as is likewise the abdomen (except
the second and third segments, which are slightly shagreened).
The thorax is three times longer than wide; the scutellum is
not raised above the mesonotum and is longer than wide ;
the metathorax is densely covered with a short pile.
[To be continued. ]
On new Mammals from South Africa. dl
IIl.—On Three new Mammals from South Africa.
By R. C. WRouGHTON.
In a small collection of mammals recently presented to the
Natural History Museum by Mr. C. W. Turner, of Good
Hope, near Aberfeldy, in the north-east of the Orange River
Colony, there are three specimens of an Otomys which differs
from any form hitherto described.
Otomys Turnert, sp. n.
An Otomys of the general form of O. Sloggett?, but slightly
larger and paler.
Fur long and soft; 17-20 mm., against 10-12 mm. in
O. Sloggettt.
General colour above between “raw umber”? and tawny
olive; the individual hairs “ slate-grey,” with a “clay-
coloured” tip (5 mm.), but a certain small proportion of
them, of greater length, with black tips (5-10 mm.).
Under surface “ pinkish buff”; the hairs with slaty bases.
Hands and feet silvery white; hairs on tail, like those on
back, markedly longer than in O. Sloggett’; in the latter the
hairs of its upper surface are coloured black, whereas in
O. Turnert all are dull white except a few on the median line,
above, towards the end.
Skull and dentition in all essential characters the same as
in O. Sloggetti, but upper molar row distinetly larger and
nasals more spatulate (6°5 mm. at broadest, instead of 5 mm.
in O. Sloggetti).
Dimensions of type specimen :—
Head and body 134 mm.; tail 493; hind foot 24; ear 17.
Skull: greatest length 385 ; basilar length 29; zygomatic
breadth 19 ; length of upper molar series 8°5; bullee 7-5.
Hab. Aberfeldy, N.E. Orange River Colony. Alt. 3500/.
Type. Young male. B.M. no. 7. 5. 25. 5. Original
number 27. Collected on 9th July, 1906.
Only one of the three specimens sent has a skull, and
unfortunately it is the youngest. Though young, however,
the type is adult, and the skull dimensions given above may
be accepted as practically normal for the species, but the
body-measurements are small. The largest of the three
specimens measured head and body 160 mm. and tail 59;
this was almost certainly a fully-grown animal.
This new species is undoubtedly very close to O. Sloggett/,
from which, however, it is separable externally by its longer
14
32 On new Mammals from South Africa.
fur, paler general colour, white feet and longer hind feet, and
by the skull having broadened spatulate nasals and a longer
upper molar tooth-row.
In my key to this genus (published in this Magazine in
October last, p. 266) the last section under A. a! might now
stand as follows :—
b°, All four lamin of m, complete ; tail short
and slender,
a‘, Fur long; hind foot 24 mm.; upper molar
series:S:0 mami. s( Oden.) cre pears O, Turnert, sp. n.
b*. Fur short ; hind foot 22mm. ; upper molar
series 7°55 mm. (Deelfontein.) ........ O. Sloggetti, Thos.
In order to compare a specimen of Pedetes contained in
Mr. Turner’s collection, I had occasion to lay out all the
specimens available in the Museum, with the result that it
became at once apparent that there are at least three local
races of the South-African Pedetes cafer separable on colour-
characters. They are distinguishable as follows :—
A. Tail dark above; paler below than above, but not
white ; ears (ctve.) 75 mm. (Cape Colony.) ...... P. cafer.
B. Tail pale above, white below.
a. General colour above “ wood-brown ”; ears longer,
(ere) -formm:) (OMRe Colony.) Vester eh tells e + 5:- P. c. orangia.
b. General colour above redder, nearest to “ vinaceous
cinnamon,” but darker ; ears shorter, (czc.) 70 mm.
(NE Dinas walls) Fea ao.ca, «ate yoxm 2 y-savegercunieiny tele topscahna > P.c. saline.
The following are descriptions of the new subspecies :—
Pedetes cafer orangie, subsp. n.
Size and general characters as in true P. cafer.
General colour above “ wood-brown.” Hairs on the
shoulders and back pale slate-coloured at their bases, then
‘ pinkish buff” with black tips ; a small proportion of wholly
black hairs sparsely scattered among the others. (In all
three forms the slaty bases, black tips, and scattered black
hairs disappear on the lower back, rump, and tail.) Colour
below pure white, the hairs white to their bases. Tail above
coloured like the rump, below pure white (except a discoloured
area near the base) ; tip for about 100 mm. black above and
below.
Skull-characters as in true P. cafer.
Dimensions of type :—
Head and body 400 mm. ; tail 410; hind foot 130; ear 75.
Skull: upper length 89; basilar length 62; zygomatic
breadth 57.
On new African Spiders and Solifuge. 33
Type. Adult female. B.M. no. 7. 5.25.11. Original
number 24. Collected by Mr. C. W. Turner on 27th Feb-
ruary, 1906.
Hab. Aberfeldy Dist., O.R.C.
Other specimens from Ladybrand and Vredefort show that
this form extends all over the Orange River Colony, and even
to Bechuanaland.
Pedetes cafer saline, subsp. n.
Size and general characters as in true P. cafer.
General colour above as in P. cafer, though slightly paler,
much redder than in P. c. orangie; the ground-colour of the
hairs ‘‘ vinaceous cinnamon” rather than “ pinkish buff,”
and the proportion of black greater. The underside quite
as in P. c. orangie.
Skull-characters as in true P. cafer.
Dimensions of type :—
Head and body 398 mm.; tail 464; hind foot 144; ear 70.
Skull: upper length 90; basilar length 63; zygomatic
breadth 57.
Type. Adult female. B.M. no. 6. 4.3.86. Original
number 1345. Collected by Mr. C. H. B. Grant on 30th
December, 1905, and presented to the Museum by Mr. C. D.
Rudd.
Hab. Woodbush, Zoutpansberg Dist., N.W. Transvaal.
T'wo other specimens taken at the same time and place do
not materially differ from the type.
1V.— Descriptions of new Species of African Spiders and
Solifuge. By A. S. Hirst.
Pterinochilus Hindet, sp. n.
3 —Cephalothorax equal in length to patella, tibia, and
tarsus of maxillipalp and considerably shorter than patella
and tibia of fourth leg. Ocular tubercle considerably longer
than broad, the clypeus narrow.
Legs. 4,1, 2, 3. Metatarsus of first leg sinuate, with the
distal end slightly thickened, and leaving but little space
between metatarsus and spur of tibia when closed on tibia ;
tibia of first leg more slender than femur. Patella and tibia
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. a
34 Mr. A. S. Hirst on new
of first leg longer than the corresponding segments of the
fourth.
Palpal organ resembling closely that of P. murinus, the
terminal portion of the style very long and slender (fig. 1).
Fig. 1.
\
\\
t
\ Y
BY \\ \
AN
‘ }
4
AHS a if
\
Palpal organ of Pterinochilus Hindez.
Measurements in mm. Length of cephalothorax 13°5, of
patella, tibia, and tarsus of palp 14°75, of first leg 49, of second
leg 44°5, of third leg 41, of fourth leg 50, of patella and tibia
of first leg 16°5, of patella and tibia of fourth leg 15°25; total
length 27.
Hab. A single male specimen was collected by Mr. 8. L.
Hinde at Fort Hall (4400 feet), British East Africa.
Remarks. The male of P. Hindei differs from that of
P. murinus, to which it is closely allied, in the much smaller
size and in that the cephalothorax equals the patella, tibia,
and tarsus of the maxillipalp, whilst in murdnus the cephalo-
thorax considerably exceeds the patella, tibia, and tarsus of
the palp in length.
Pterinochilus murinus, Pocock.
Pterinochilus murinus, Pocock, Proc. Zool. Soc. 1897, p. 753, pl. xl.
fig. 4; 1898, p. 501.
3 .— Cephalothorax longer than patella, tibia, and tarsus of
maxillipalp, and shorter than patella and tibia of fourth leg.
Legs. Metatarsus of first leg sinuate and dilated distally,
tibia of first leg more slender than femur. Patella and tibia
of first leg exceeding patella and tibia of fourth in length.
Size. Vhis species 1s much larger than P. fHinde?,
African Spiders and Solifuge. 35
Measurements in mm. Length of cephalothorax 19, of
patella, tibia, and tarsus of palp 17°25, of patella and tibia
of first leg 23, of patella and tibia of fourth leg 20; total
length 32.
Hab. Mombasa (Capt. Crawshay).
Pterinochilus meridionalis, sp. n.
&¢ .—Obolour. Cephalothorax greyish black, with lines of
yellowish hairs radiating from the fovea; sternum and ventral
surface of coxee of legs deep black ; abdomen greyish yellow,
the ventral surface marked with a dark patch.
Cephalothorax almost equal in length to patella and tibia
of fourth leg, much shorter than patella and tibia of first leg,
and considerably exceeding patella, tibia, and tarsus of
maxillipalp in length. Ocular tubercle longer than wide,
clypeus of moderate length.
Legs. Metatarsus of first leg moderately stout, curved,
thickened apically, of much less length than tibia, and leaving
but little space between spur and hbase of metatarsus when
closed on tibia.
Lalpal organ with the style fairly long and slender (fig. 2).
Palpal organ of Pterinochilus meridionalis.
Measurements in mm. Length of cephalothorax 14, of
patella, tibia, and tarsus of palp 11°75, of tibia of first leg 11,
of metatarsus of first leg 9°5, of patella and tibia of first leg 17,
of patella and tibia of fourth leg 14°5 ; total length 25.
Hab. A single male specimen was collected at Dowa,
British Central Africa, by Mr. A. R. Andrew during March
1907.
ee
BIS
36 Mr. A. S. Hirst on new
Fleteroscodra crassipes, sp. 1.
9 —Colour. Legs marked with dark spots, which are
distributed much as in H. maculata.
Cephalothoraz longer than wide, of much less length than
patella and tibia of fourth leg, and exceeding patella and
tibia of first leg in length.
Sternum a little longer than broad; posterior sigilla
situated in the posterior third of the sternum and at but little
distance from the lateral margins.
Legs. ‘Tibia and patella of first lez much shorter than the
corresponding segments of the fourth. Fourth leg long-
haired and very stout, the femur being much swollen and the
patella and tibia of considerable thickness. Metatarsus of
fourth leg a little longer than tibia (fig. 3).
Fig. 3.
—S = = - SS \ .
ZA
a
BoB Pf) LD" lft” PR —_
LEP eae Ze =S_ on
Fourth leg of Heteroscodra crassipes from above.
Measurements in mm. Length of cephalothorax 21, of first
leg GO, of second leg 57, of third leg 57, of fourth leg 77, of
patella and tibia of first leg 20, of patella and tibia of fourth
leg 25 ; total length of body 50 ; width of cephalothorax 18°25,
of femur of fourth leg 7°75, of tibia of fourth leg 5:5.
Hab. An adult female from Efulen, Camaroons, and
another from the forest 25 miles inland to the east of Kribi,
Camaroons. These specimens were collected by Mr. G. L.
Bates.
Remarks. This spider differs from H, maculata, the only
other species of the genus, in the much greater thickness of
the fourth pair of legs and in the great dilatation of their
femora.
Cladomelea ornata, sp. n.
?.—Colour pale yellow; tarsi and distal portions of
metatarsi of legs fuscous, the metatarsi of the legs of the
anterior pairs being marked in the middle of their length
with an additional dark patch; patelle and tibie of legs
African Spiders and Solifuge. 37
with light brown spots; cephalothoracie spines apically
darkened.
Cephalothorax. Ocular tubercle moderately elevated as
compared with that of C. longipes, the three spines of the
cephalothorax of fair length and the middle one slightly
curved in an anterior direction (fig. 4).
Fig. 4.
Side view of cephalothorax of Cluwomelea ornata.
Abdomen. Dorsal tubercles of abdomen small, almost
uniform in size, and distributed much as in C. longipes.
The two median tubercles of the second row are replaced,
however, by a single tubercle. Additional tubercles are
present in the posterior median part of the abdomen, a
couple being situated between the row of three tubercles
and the lozenge-shaped group of four tubercles and another
pair placed posteriorly to the lozenge-shaped group.
Legs. Patella and tibia of first leg a little longer than
metatarsus and tarsus, and with tibia, metatarsus, and tarsus
more slender than is the case in C. longipes.
Measurements inmm. Length of first lez 27, of second
leg 19°5, of third leg 10, of fourth leg 18°5, of posterior
cephalothoracic spine 2°75, of ocular tubercle *5, of cephalo-
thorax 4°75, of abdomen 10; total length 14:5; breadth of
cephalothorax 4°75, of abdomen 12.
Egg-cocoon pale yellow in colour, hemispherical, the surface
smooth, the pedicle very short.
Hab. Dry foot-hills, South-east Ruwenzori, altitude
3400 feet; a single female specimen was collected by the
British Expedition to Ruwenzori in May 1906.
Remarks. he more important differences between this
new form and C. long/pes are shown below. The characters
of the latter species are in part taken from a female specimen
from San Salvador, Congo.
Ocular tubercle moderately elevated, the three
spines of the cephalothorax fairly long; tu-
bercles of dorsal surface subequal in size; patella
38 On new African Spiders and Solifuge.
and tibia of first leg together a little longer than
the metatarsus and tarsus .............0005- C. ornata, sp. n.
Ocular tubercle high, the three spines of the cephalo-
thorax long; abdomen dorsally with a pair of
antero-lateral enlarged tubercles ; patella and
tibia of first leg together a little shorter than
the metatarsus and tarsus. ...0 0... 0h = acne C. longipes, Cambr.
Solpuga Fordi, sp. n.
S.—Colour pale yellow; mandible and head-plate pale
brown above, abdomen dorsally pale or darkened.
Head-plate exceeding tibia or metatarsus and tarsus of
maxillipalp by nearly two thirds of its breadth.
Mandible with the end curved. Anterior teeth two in
number and separated from the succeeding large tooth by
two intermediate ones, the first of these being exceedingly
BK nay
Alfeae: oO.
A. Mandible of Solpuga Fordi from the inner side.
B. Distal half of flagellum of 8. Fordi from above.
C. Side view of terminal portion of flagellum of S, Fordi.
minute. Flagellum dorsally provided with a spined projection
on angle, situated at the junction of the dark-coloured anterior
part with the distal and paler part. Inner side of flagellum
furnished with a smooth ridge (sometimes with a few spines
towards the distal end) running from the angular point and
* Zur Lésung der ‘ Muralis-Frage.’” 39
terminating towards the distal end of the flagellum in a spined
projection. Distal part of flagellum dilated and squarely
truncate, Flagellum inserted above the interval between the
two anterior teeth and extending posteriorly to the middle of
the mandible ; basal lamina moderately elevated (fig. 5).
Maxillipalp. Tibia, metatarsus, and tarsus of maxillipalp
ventrally provided with numerous cylinder-bristles, those of
the metatarsus and tarsus unequal in size and more numerous
than those of the tibia.
Measurements in mm. Length of tibia of maxillipalp 13°75,
of metatarsus and tarsus of maxillipalp 12; breadth of
cephalothorax 8. (In another specimen tibia 12, metatarsus
and tarsus 11, cephalothorax 6:5.)
Hab. Two males of this new form were collected by
Captain R. Ford in British East Africa (near Lake Baringo ?).
Remarks. The specimens are dry and somewhat shrunken,
and therefore the total length cannot be given. The colour
also is somewhat faded. ‘The species may be readily distin-
guished by the peculiar form of the flagellum.
V.—Remarks on Prof. L. von Méhely’s Paper “Zur Lisung
der ‘ Muralis-Frage.’”” By G. A. BouLENGER, F.R.S.
PurRsuING his studies towards the solution of the Lacerta
muralis problem, studies which, it is already pertectly clear,
will result in a plethora of ill-defined Jordanian species, the
distinguished Hungarian zoologist Prof. von Méhely has
recently published a short paper * in which he attempts to
prove that the derivation of forms must have taken place in
a direction the reverse of that postulated by Eimer f and since
endorsed by most workers at this difficult group, including
myself, and to which Dr. Gadow’s recent investigation of the
American lizards of the genus Cnemidophorus t has given
further support.
The numerous variations in the pattern of markings in
LL. muralis fall roughly under five heads—striated, spotted,
reticulated, barred, ocellated. It is evident to any one
studying large series that these different types of markings
are insensibly connected by every possible gradation §. ‘lhe
* Ann. Mus. Hung. v. 1907, p. 84, pl. iii.
+ Arch, f. Naturg. 1881, p. 375,
t Proc. Zool. Soc. 1906, i. p. 277.
§ Eimer has pointed out that the pattern on the posterior part of the
body anticipates the evolution of that on the anterior part. This is true
40 Mr. G. A. Boulenger—“ Zur Lisung
question is merely to determine which is to be regarded as
the most primitive. Ontogeny, in these lizards as well as in
their American analogues the Cnemiédophor’, indicates the
direction, as it is the general rule for new types of markings
to be produced in adult males and to be then transmitted to
females and young *, and strong evidence, derived from other
features, would be needed to convince us that, as now held
by Méhely, “striation is not the phyletic initial form, as
believed by Eimer, but the phyletic terminal stage.” His
opinion rests, in the first place, on the assumption + that
L. saxicola and L. chalybdea represent the most primitive
forms of wall-lizards, and he now attempts to strengthen his
position by arguments which seem to me based on a miscon-
ception of the evolution of cranial characters.
In my contribution published in 1905 { I expressed my
full agreement with Himer in regarding the striated type of the
var. campestris as the most primitive among all the wall-lizards,
and I added that “ we are led to regard the var. campestris
as the most ancient form from which the others were derived ;
and this, I think, is also supported by the structural cha-
racters, which differ less from what we may assume to be the
more normal or generalized form of Lacerta before adaptation
to climbing petrophilous habits had been reached.” A form
with massive convex skull, like the var. campestris, would
lead through a number of almost insensible gradations, such
as actually exist, to the much flattened skull which has been
distinguished by Eimer as the platycephalous type in oppo-
sition to the pyramidocephalous. I have never been able to
draw a satisfactory distinction between the two types, and I
do not quite understand how Prof. v. Méhely manages to
group his “ species’’ according to this character. In the list
he gives I notice that L. tiliguerta is regarded by him as
pyramidocephalous. In the paper of mine§ to which he
refers I have described the head of the true “ Tiliguerta”
from Sardinia as “ rather strongly depressed, the occiput quite
flat or even slightly concave”’; it is certainly as a rule more
of the markings on the tail. In primitive striated forms, suchas Z, agilis,
L. taurica, L. campestris, the tail is frequently more or less striated or
“ maculato-striata,’ whilst in extreme reticulated forms, such as LZ. ovy-
cephala, L. sardoa, L. nigriventris, it is more or less distinctly barred.
Reproduced tails, if bearing any markings, are always longitudinally
striped.
* This is clearly shown in the vars. campestris and serpa. We cannot
imagine the reverse.
+ Ann. Mus. Hung. ii. 1904, p. 376.
{ Trans. Zool. Soc. xvii. 1905, p. 388.
§ L.c. p. 409, pl. xxvili. fig. 7.
der ‘ Muralis-Frage.’” 41
platycephalous than that of the typical Z. murals, which
falls in the platycephalous group of Méhely. Besides, the
works of Eimer and of Bedriaga show, in several instances,
that these authors have been unable to correctly appreciate
the character to which, in my opinion, they have attached too
great importance *,
Yet, when we compare extreme forms, such as var. cam-
pestris or fiumana, on the one hand, and var. Bedriage + or
sardoa on the other, the difference in the two types of heads
stands out very strikingly. Weare not much the wiser when
the skulls have been prepared, as the characters pointed out
by Prof. v. Méhely are, for the most part, correlative of the
degree of elongation or depression of the head, which can be
appreciated without injuring the specimens. It must be
borne in mind that skulls of lizards cannot be extracted as we
do in the case of mammals. Preparing the skull means the
partial destruction of the specimen, and in a discussion of
this kind, dealing mainly with individual variations, annectant
examples cannot always be sacrificed. Prof. v. Méhely has
eiven us figures of two extreme types of skulls, but I could
easily lay out a series that would to such an extent bridge over
the differences as to show of how little practical value they
are for the definition of species. A discovery of Prof. v.
Méhely’s is the incomplete ossification of the supraocular
region in the most pronounced platycephalous wall-lizards.
But even here he is obliged to make this restriction—that in
some, in the var. Bedriage, for instance, the fontanelle in the
supraocular bony plates is ‘nicht immer vorhanden”? in adult
males. The character is therefore not of so great importance
after all.
What surprises me most is to find that Prof. v. Méhely
is not at all aware of the individual variations which occur in
the skulls of the forms which he classifies as “rein platy-
cephal” and ‘rein pyramidocephal.” Thus he attaches a
* For instance, in the vars. zzgriventris, serpa, and quadrilineata, some
specimens have been referred by Bedriaga to the pyramidocephalous
group (L. muralis neapolitana, Bedr.) and others of the same race to the
platycephalous (LZ. muralis fusca, Bedr.). Eimer regards the Maltese
lizard as platycephalous and its Filfola derivative as pyramidocephalous,
a distinction which is not borne out by the material at my disposal.
Werner has also fallen into the same pit, in describing examples of the
same form (Z. muralis htteralis, Werner) under the two groups (vars.
lissana and fiumana). I myself do not blush at confessing similar errors,
due, perhaps, more to the nature of things than to any want of ‘ Scharf-
blick” on my part.
+ Prof. v. Méhely calls this lizard Z. reticulata, Bedr. But this name
is otherwise employed in the genus Lacerta (Z. muralis reticulata,
Schreiber, Eimer).
42 Mr. G. A. Boulenger—“ Zur Lisung
great importance to the size and position of the true supra-
orbital bone, and yet a skull of an adult male of the var.
Bedriage which I have prepared for the purpose of verifying
his statements has the supraorbital bone entirely concealed
under the first osteodermal supraocular and the first supra-
ciliary, and this bone is rather smaller than in a male of the
var. téliguerta from Cagliari. It is clear to
me that Prof. v. Méhely is not familiar with
the skull of the true Z. tiliguerta, which, as
stated above, he regards as_ pyramido-
cephalous. The sketch here given of the
bony plates of the supraocular region, care-
fully prepared for me by Mr. E. Degen
from an adult male from Cagliari, shows that
the ossification of that region may in indi-
vidual cases be incomplete; and I should
add that the nasal apertures may be nearly
as large asin L. Bedriage, also that ossifi-
cations are altogether absent from the
temporal region. I have already pointed
out *, and still believe, that the var.
tiliguerta constitutes in a certain sense a
link between the var. Brueggemanni on the one hand, and
the vars. Bedriage and sardoa on the other.
Mr. Degen has also found the supraocular region incom-
pletely ossified in male specimens of the vars. Lilfordi and
Jiumana,.
Prof. v. Méhely thinks a study of the cranial characters
affords a key to the solution of the Z. muralis problem from
the point of view of the phylogeny. He starts from the
assumption that the more feeble development of the osteo-
dermal plates of the head indicates a lower stage of evolution,
and as the ultra-platycephalous forms of wall-lizards belong
more to the reticulate type of pattern, whilst the striated
lizards (with six light streaks) are pyramidocephalous, he
declares ‘so ist es klar, dass die Liingsstreifung nicht die
phyletische Ausganesform kennzeichnet, wie Kimer annahm,
sondern gerade die phyletische Endstufe anzeigt.”
I wish to give here my reasons for differing from this view.
In most groups of lower vertebrates, in which we have some
indication of orthogenetic derivation of forms, we find that a
massive skull degenerates into a more feebly ossified one
so far as the “roof”? is concerned. Among Silurid fishes we
have a beautiful example in the series Clartas—A llabenchelys
Supraocular region
of L, tihguerta.
* Tc. p. 404.
der ‘ Muralis- Frage.’ ” 43
— Clariallabes— Gymnallabes—Channallabes, in which we
witness the gradual disappearance of the plates which roof
over the sides of the skull, concurrently with the eel-like
elongation of the body, the reduction of the caudal fin, and
the reduction and ultimate suppression of the paired fins—a
most suggestive series, the direction of which is unmistakable.
Again, in the Characinid fishes, as pointed out by Sagemehl,
the more primitive types, with large toothed maxillary bone,
have a massive skull, the fontanelles appearing together with
the reduction of the maxillary bone. The same story is told,
in a somewhat different way, by Chelonians (Chelydra—
Staurotypus, Emys—Cistudo, &c.). In Lizards, also, when
we have to deal with an unmistakable orthogenetic series,
ithe drift of which is open to no question, as in Chalcides, for
instance, the more generalized type has a more convex skull,
better protected by osteodermal plates. But there is another
point which is of great importance, and which Prof. v.
Méhely does not appear to have considered. The Lacerte
with massive skulls, from which I would assume the platy-
cephalous lizards to have been derived, have teeth on the
palate (pterygoid bones). These teeth are nearly constantly
present in Lacerta taurica and constantly absent in the forms
of L. muralis with supraocular fontanelles mentioned by
Prof. v. Méhely. Now, the only cases in which I have found
teeth on the palate in L. muralis have been in examples of
the pyramidocephalous vars. campestris and serpa*, a fact
which, in my opinion, goes a long way to support the view
of Eimer as to the general drift of evolution in this group of
lizards.
Much as I value the careful investigation of neglected
points of structure, whether external or osteological, to which
Prof. Méhely is devoting himself, I cannot help regretting
the too frequent appeals he makes to phylogeny in order to
give importance to characters which, from a strictly syste-
matic point of view, must be regarded as trivial and had
better be omitted from specific diagnoses. I have pointed out
on various occasions } that some of the lepidosis characters on
* T have looked for these teeth in a large number of examples of the
typical form without ever succeeding in finding any. Siebenrock (Sitzb.
Akad. Wien, ciii. i. 1894, p. 254) must therefore, in all probability, have
had skulls of some other form before him when he wrote that six or
seven pterygoid teeth are present in Z. muralis. As his specimens are
stated to be from Dalmatia, it is most likely that they belong to one of
those pyramidocephalous forms which were grouped by Bedriaga under
L. muralis neapolitana.
+ Proc. Zool. Soc. 1904, ii. p. 333; Nov. Zool. xii. 1905, p. 75; Trans.
Zool. Soe. xvii. 1905, p. 351.
44 Mr. G. A. Boulenger— Zur Lésung
which he lays stress hopelessly break down when put to the
test of large series from more extensive areas than it is the
custom for faunists to deal with. An interesting example of
the danger of hasty generalizations of this kind has just come
under my notice.
As I mentioned in describing the typical form of Lacerta
muralis*, one of the two specimens (topotypes) from near
Vienna, received from my ever-obliging friend Dr. Werner,
proved to be highly aberrant in several respects. The
parietal shields were abnormally divided by a transverse cleft.
On recently enquiring from Dr. Werner as to whether he had
other examples from the same locality (Véslau, near Baden,
Lower Austria), I was greatly surprised to hear that an
examination of his material had satisfied him that this
division, instead of being anomalous or accidental, is the rule
in Lower Austria. Among his specimens from Médling,
Baden, Véslau, Reichenau, and Miesenbach, not one is without
at least an indication of it, whilst he cannot find such a thing
in any of his other specimens from various parts of Europe.
In order to further confirm this observation, Dr. Werner has
made excursions to Baden and Véslau, whence he sent me
six living examples, all showing a complete or incomplete
cleft across the parietal shields. Therefore this anomaly,
which very seldom occurs in other parts of the very exten-
sive habitat of the wall-lizard, although it is frequent in the
viviparous lizard t, appears to have become fixed in a small
district near Vienna. E. Martint mentions the case of
the inhabitants of a small secluded village in France,
nearly all of whom, at the end of the eighteenth century,
had an extra digit to both hands and feet; gradually,
however, as intercourse with neighbouring communities
became frequent, the deformity was wiped out. Some
years ago, when reporting his interesting discovery of
Lacerta praticola in a valley near Herkulesbad in Transyl-
vania §, Prof. v. Méhely pointed out the frequent presence of
an accessory shield between the interparietal and the occipital,
observed in 48 specimens out of 78; and as it so happened
that the unique specimen on which the species was estab-
lished by Eversmann offered the same anomaly, which occurs
* Trans. Zool. Soc. xvii, 1905, p. 354, pl. xxv. fig. 4.
f+ In which it may be transmitted to the offspring, as shown by a
female from the dunes near Ostend, which produced four young in
captivity (Aug. 4-7), all showing the same anomaly,
{ Histoire des Monstres, 1880. Quoted from Delage, L’ Hérédité, p. 194.
§ Math. Naturw, Ber. Ungarn, xii, 1894, p. 255, and Zool. Anz. 1895,
p. 474.
der ‘ Muralis-Frage.’”’ 45
occasionally in many other species of lizards, he was led to
regard it as “ ein ausgesprocheness Merkmal der Art [Z. prati-
cola], das mit der Zeit wohl eine vollkommene Bestindigkeit
erlangen wird, besonders da drei Schildchen hiufiger bei den
Weibchen, zweie aber bei den Minnchen auftreten und bei
Lacertiden bekanntermaassen die weiblichen Charactere viel
alloemeiner auf die Nachkommenschaft vererbt werden ” *,
Now the case of the Z. praticola from the valley near
Herkulesbad is analogous with that of the LZ. muralis near
Vienna, for in other parts of the habitat of the former species
the intercalated shield is only exceptional, as pointed out by
Kessler and by Bedriaga {, not being found in any of the
fifteen specimens from Caucasia examined by them ; it is
absent in the three specimens from Sukum Kaleh, Caucasia,
and in six out of the nine from the Comana Forest, Roumania f,
preserved in the British Museum. And yet Prof. v. Méhely
included this character of the three azygous shields between
the parietals among the points which, in his opinion, militate
in favour of regarding L. praticola as more nearly related to
LL. muralis (“besonders die auch bei L. muralis hiiufige
(uerteilung des Interparietale”’) than to LZ. vivipara §.
Needless to say, the intercalation of a shield between the
interparietal and the occipital (or division of the interparietal)
occurs occasionally also in L. vivipara. I have before me
specimens presenting such an anomaly from Falmouth,
Brussels, T'alomitza Valley and Brosteni, Roumania, and
Moscow. It should also be mentioned that a few (two to
four) granules may be present between the supraoculars and
the supraciliaries in L. vivipara, whilst, on the other hand,
these granules may be reduced to three or four in Z. praticola.
Great is no doubt the interest attaching to the record of
such individual deviations from the normal condition, great is
also the danger of introducing them in the diagnosis of species.
I seize this opportunity to point out that the genus
Apathya, recently proposed by Méuely || for Lacerta cappa-
docica, Werner, does not seem separable from Latastia,
Bedriaga. Iam indebted to Dr. Werner for a specimen of
Latastia cappadocica, and Dr. J. Roux has shown me another
(labelled as from Mesopotamia). ‘The structure of the digits
* T am not aware of any evidence in support of this statement.
¥ Zool. Anz. 1895, p. 261.
t I am here at variance with Kiritezcu, Bull. Soc. Sc. Bucarest, x.
1901, p. 318, but he does not state how many examples he has examined.
§ A view which has since been abandoned (cf. Ann. Mus. Hung, li,
1904, pp. 875 & 377),
|| Termész. Késlén., Budapest, Ixxxy. 1907, p. 2
-
46 Ona new Engystomatid Frog from Namaqualand.
is similar to that of ZL. longicandata, so is the shape of the
posteriorly narrowed frontal shield, with truncate antero-
lateral angles, and the coloration, as described and figured by
Werner, is highly suggestive of some examples of that species.
The three shields behind the nostril (“ postnasals ” of Werner)
cannot be regarded as a generic character, especially in
view of the well-known variability of these shields in the type
species of the genus Lacerta. Nor can generic importance be
ascribed to the scaling of the lower eyelid if we bear in
mind the modifications which it undergoes within the limits
of the genera Lacerta and Hremias.
VI.—Deseription of a new Engystomatid Frog of the Genus
Breviceps from Namaqualand. By G. A. BouULENGER,
HRS:
[Plate II.]
Breviceps macrops. (P1. II.)
Head comparatively larger and body less gibbose than in
B, gibbosus and allies, eyes much larger, their diameter 63 to
8 times in the length of head and body; interorbital space
narrow, barely half the width of the upper eyelid. Digits
shorter and thicker than in the other species of the genus ;
first and second fingers equal, twice as long as the fourth,
a little shorter than the third, which is not longer than the
eye; toes extremely short, increasing in length from the
rudimentary first to the fourth, which is not longer than the
third finger, the fifth as long as the second; subarticular
tubercles very small, very feebly prominent under the fingers,
\% <F
Lower aspect of hand and foot.
absent under the toes; two large, feebly prominent metatarsal
tubercles, the inner longer than its distance from the extre-
mity of the first toe. Body with small smooth warts above,
limbs and lower parts smooth. Pale sand-colour above, with
e On new Reptilia from Mashonaland. 47
dark brown spots or marblings; a broad dark angular or
curved band from one eyelid to the other across the back of
the head anda narrow straight dark streak across the forehead,
between the anterior borders of the upper eyelids ; upper lip,
limbs, and lower parts white.
mm
IR Royoa MPAGUERR IHC) IMG Apo ooo Dada bo ao oUD 48
Flea diye mere crete ser on ae clarhiamaene 3
WsiGhay Ge INN. oo oonecobeeuveaaauuer ity
Dram SleRO yO eek verte scar tek eoNere idee. 6
Interorbitall wadthy $e... fe swe sts 0 hat 24
Awake Ge mnOldd svoogodeecavananedoc 12
Moreglimnll ieee seer eckoi tna Mole cise cone: 22
Elaine setae cus wre a Bicseecicd? ates song its 10
Mim de lining se tey ee teeta cud cestaiond ccs 30
1 SHG 0) rt oth 0 oa Bn at Ao Sar ne eR 3
I am indebted to my friend Mr. L. Péringuey, Director of
the South African Museum, for three specimens of this most
distinct new species, one of which he has kindly presented to
the British Museum, Mr. Péringuey, who has been able to
keep them alive for some time, writes to me that they spend
the whole day buriedin damp sand. At night the pupil, which
contracts to a narrow horizontal slit, expands over nearly the
whole eye. Like B. gibbosus, this frog, when teased, or
merely touched, inflates its body tremendously and utters
shrill screams—a habit which seems to be frequent among
burrowing batrachians whatever their affinities. Unlike
B. gibbosus, B. macrops does not cover itself with a viscous
secretion when alarmed.
EXPLANATION OF PLATE II.
Breviceps macrops, adult and half-grown,
VII.—Descriptions of a new Toad and a new Amphisbenid
from Mashonaland. By G. A. BouLeNGER, F.R.S.
[Plate III. ]
In November last I had the pleasure of drawing attention in
these ‘ Annals’ to the Chirinda Forest in 8.E. Mashonaland,
where Mr. Guy H. K. Marshall had discovered a new
chameleon of the genus Rhampholeon. I remarked that its
reptile and batrachian fauna, when explored, was likely to
afford further startling additions to South-African herpetology.
48 Mr. G. A. Boulenger on
A small collection, including examples of only two species,
was since made there by Mr. C. F. M. Swynnerton, and
has been presented by him to the British Museum. The
following descriptions show that there was some foundation
for my optimism.
Bufo anotis. (PI. IIL.)
Crown without bony ridges; head as long as broad; snout
pointed, projecting beyond the mouth, with strong canthus ;
loreal region nearly vertical, concave ; interorbital space flat,
a little broader than the upper eyelid; no tympanum, no
eustachian tubes. Fingers’ short, blunt, with strong double
subarticular tubercles, first and second equal; toes short, one-
third webbed, the web covered with coarse granules ; sub-
articular tubercles under the fourth toe double and very
prominent ; two round, subconical, metatarsal tubercles ; no
tarsal fold. The tibio-tarsal articulation reaches the posterior
border of the eye. Upper and lower parts covered with small
smooth warts; parotoid glands very large, moderately promi-
nent but well detined, only a little longer than broad, extending
on the sides to the base of the arm. Upper parts brown and
sides of head and body blackish, or entirely blackish above ;
liinbs, if light-coloured, with blackish cross-bars ; lower parts
bright gainboge-yellow, the breast speckled with black.
Male without vocal sac.
From snout to vent 40 mm.
Three male specimens, in nuptial dress, from small holes
containing water at the foot of large trees.
This remarkable new toad agrees with B. Preusst, Matschie,
and B. taitanus, Peters, in the absence of a tympanum. It
is readily distinguished from the former by the warty integu-
ment and the distinct parotoid glands, from the latter by the
more pointed snout with less oblique lores and the much
larger parotoids extending down the sides.
CHIRINDIA, gen. nov.
Distinguished from Amphisbena by the absence of an
ocular shield, the eye being situated under the posterior part
of a large shield (nasal + labial + prefrontal + ocular) covering
each side of the snout, and by the absence of preanal pores.
Teeth very small.
Chirindia Swynnertont.
A small triangular rostral, a pair of large shields covering
° 7 Ea . . o
the snout, a pair of small frontals in contact with a large
new Reptilia from Mushonaland. 49
shield extending, to the oral border, a pair of postfrontals and
a pair of parietals, both broader than long, and four small
square shields between them and the oral border; four lower
labials, first very large and forming a short suture with its
fellow behind the point of the elongate symphysial. 246
annuli on the body and 24 on the tail; in the middle of the
body an annulus contains 24 segments, 12 above and 12
below the lateral lines; dorsal and lateral segments a little
longer than broad, the two median ventrals about twice as
broad as long. Lateral line moderately distinet. Anal
segments scarcely enlarged. Hind of tail rounded. Flesh-
coloured, tinged with purplish.
Chirindia Swynnertont.
Total length 135 mm.; diameter of body 3; length of
tail 14.
A single specimen of this remarkable Amphisbenid worm-
like lizard was obtained. In the fusion of the ocular with
the large paired shield which covers the whole of the snout, in
addition to the small azygous rostral, Chirindia resembles
Placogaster, Blgy., the unique species of which inhabits
Senegambia ; but it is at once distinguished from that genus
by the paired ventral shields and the absence of preanal
ores. Chirindia is certainly, with Placogaster, the most
remarkable type of Amphisbeenid discovered within the last
twenty-five years.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 4
50 On anew Cichhd Fish.
VIII.—Deseription of a new Cichlid Fish from Portuguese
East Africa. By G. A. Boutencer, F.R.S.
A SMALL series of fishes obtained by Mr. C. M. F. Swynnerton
in the Idunda River, a limestone-stream flowing into the lower
Buzi River, in Portuguese Gazaland (altitude about 100 feet),
and presented by him to the British Museum, consists of
examples of Barbus decipiens, Blgr., Tilapia shirana, Bler.,
T. melanopleura, A. Dum., and a new species of Ttlapia,
here described.
Tilapia Swynnertont.
Depth of body 24 times in total length, length of head 3
times. Snout obtuse, with straight or convex profile, as long
as the eye, which a little exceeds interorbital width and is
contained 31 times in length of head; mouth extending to
below anterior border of eye or a little beyond ; maxillary
exposed at the end; lips rather thick; 3 series of tricuspid
teeth and an outer series of rather large bicuspid teeth (40
to 42 in the upper jaw), the cusps subequal and very obtuse ;
3 or 4 series of scales on the cheek; large scales on the
opercle. Gill-rakers short, 8 on lower part of anterior arch.
Dorsal XV 9-10; spines increasing in length to the last,
which measures nearly half length of head; longest soft rays
% length of head. Anal III 8-9; third spine shorter than last
dorsal. Pectoral barely $ length of head, not extending to
origin of anal. Ventral reaching origin of anal or a little
beyond. Caudalrounded. Scales ctenoid, 20231 lateral
lines oa Dark olive-brown, with ill-defined black cross-
bars ; a black vertical bar below the anterior half of the eye
and a black opercular spot; fins grey; male with two or
three round yellow spots on the anal.
Total length 95 mm.
Described from three specimens.
This species bears a great resemblance to H wlochromis
Moffatt, Casteln. (Chromis philander, M. Weber), but differs
in the dentition and in the larger eye.
IX.—Deseriptions of Three new Freshwater Fishes discovered
by Mr. G. L. Bates in South Cameroon. By G. A.
BouLENGER, F.R.S.
Synodontis Butesi?.
Depth of body 33 to 43 times in the total length, length of
head 3 to 33 times. Head a little longer than broad, rugose
On Three new Freshwater Fishes. ult
above; snout obtusely pointed, as long as postocular part of
head ; eye supero-lateral, its diameter 4 to 5} in length of
head, 12 to 23 times in interorbital width; pramaxillary
teeth forming a short and broad band; movable mandibular
teeth not half diameter of eye, 30 to 40 in number; maxillary
barbel not distinctly margined, extending about to middle of
pectoral spine ; mandibular barbels with ubtuse or tubercular
branches, outer at least twice as long as inner and § to 2
leneth of head. Gill-openings not extending downwards
beyond root of pectoral spine. Occipito-nuchal shield rough
like the occiput, obtusely tectiform, not or’but slightly longer
than broad, the posterior processes rounded. Humeral
process rugose, acutely pointed, extending as far back as
occipito-nuchal process. Skin without villosities. DorsalI 7;
spine nearly as long as head, striated, strongly serrated in
front, with a few very feeble serree behind. Adipose dorsal
rather small, not longer than its distance from the rayed
dorsal. Anal III 8-9. Pectoral strongly serrated on outer
side, still more strongly on inner. Caudal fin deeply notched,
crescentic. Caudal peduncle longer than deep. Brown,
blotched and marbled with darker, with three broad irregular
dark bands, the first below the dorsal fin, the second below
the adipose, the third in front of the caudal fin; fins whitish,
with round black spots.
Total length 90 mm.
Several specimens from the Ja River.
The smallest species of the genus. The first specimens
received were thought by me to be young S. obesus, Blgr., a
species which has not yet been discovered in Cameroon.
Clarias longior.
Depth of body 8 to 9 times in total length, length of head
5 to 54 times. Head extremely depressed, once and } as long
as broad, smooth; occipital process very short, pointed ;
frontal fontanelle knife-shaped, about 3 times as long as
broad ; occipital fontanelle smaller, in advance of occipital
process; eye very small, 24 times in length of snout, 5 or 54
times in interorbital width, which equals width of mouth and
3 length of head; band of preemaxillary teeth 4 times as long
as broad; vomerine teeth granular, forming a curved band
which is as broad as the preemaxillary band; nasal barbel 4
to ? length of head, maxillary once and 4 to once and 3, outer
mandibular once, inner mandibular 3 to 3. Gill-rakers very
few, 12 on anterior arch. Clavicles concealed under the skin.
Dorsal 80, its distance from occipital process 3 to 2 length of
2 Mr. E. S. Russell on
head, its distance from the caudal equal to diameter of eye.
Anal 70-73, nearly reaching the caudal. Pectoral 2 to 2
length of head, the spine smooth and about 2 the length of
the fin. Ventral once and 2 to once and ? as distant from
base of caudal and from end of snout. Caudal 2 length of
head. Dark brown above, whitish beneath.
Total length 225 mm.
One specimen from the Kribi River and one from the Lobi
River.
Eleotris kribensts.
Body cylindrical or a little compressed, its depth 4 to 5
times in total length; length of head 3 to 3} times in total
length. Head broader than deep, naked; snout broad,
rounded, as long as or a little longer than the eye, the diameter
of which is 4 to 4} times in length of lead and equal to or a
little less than interobital width; lower jaw projecting ;
maxillary extending to below anterior third or centre of eye ;
no canine teeth ; no preopercular spine. Dorsals VI, I 8-9,
well separated from each other, longest rays not longer than
head. Anal I 7, opposite to second dorsal. Pectoral 2 to #
length of head, a little longer than ventral. Caudal rounded,
alittle shorter than head. Caudal peduncle once and 4 to once
and 2 as long as deep. Scales smooth on the nape, strongly
ciliated on the body, 52 to 35 in a longitudinal series, 12
between origin of dorsal and anal. Yellowish to brown,
dotted with darker, with or without five or six ill-defined
dark cross-bands; a blackish bar at the root of the caudal
fin; fins brown or blackish and white-edged in males, whitish
with blackish spots in females.
Total length 50 mm.
Numerous specimens from the Kribi River.
X.— The Atractylis coccinea of T. S. Wright.
By E. 8. Russevy, M.A.
Tus hydroid was described by Wright (Ann. Nat. Hist.
ser. 3, viii. (1861) p. 180) in the following words :—
© Atractylis coccinea, n. Sp.
“ Polypidom creeping, widely reticulate. Polyp fusiform,
set at an obtuse angle to its stalk, rich crimson or pink, with
eight alternating tentacles, four long and four short.”
the Atractylis coccinea of T. S. Wright. 53
No gonophores were found. Hincks (¢ Hydroid Zoophytes,’
1868) provisionally transferred the species to the genus
Lerigonimus. Allman does not mention it at all, and I have
not been able to find any subsequent record of it.
In 1905 1 obtained near Millport Marine Biological Station
numerous specimens of what is almost certainly the Atractylis
cvecinea of Wright. They differ from Wright’s specimens in
having twelve equal tentacles, but they all have the hydranth
set at an obtuse angle with the stalk, a very characteristic
point.
The species was not figured by Wright, and his description
was in some respects incomplete. I therefore give here a
detailed description of my specimens and a figure.
The species must be assigned to the genus Wrightia,
Allman (1872). The genus Atractylis, Wright (1859),
contained forms which are now distributed among the genera
Bougainvillia, Perigonimus, and Wrightia. Hincks’s genus
Atractylis is synonymous with Allman’s Wrightia, but the
name Wrightia is to be preferred, since Atractylis ig the
long-established name of a genus of plants. Wrightia, when
constituted by Allman, contained one species, Wrightia
arenosa (Atractyls arenosa, Alder, Suppl. Catalogue, p. 7,
pl. x. figs. 5-7), and the diagnosis of the genus contained
some of the specific characteristics of Wrightia arenosa (Alder),
namely, the funnel-shaped stems, the retractile hydranth, and
the position of the gonophores on the hydrocaulus. I propose
the following definition of the genus, which is in all essentials
the same as Allman’s definition, but leaves out any reference
to the purely specific characters of either of the two species
which the genus contains, arenosa (Alder) and coccinea
(Wright).
WRIGHTIA.
Atractylis (in part.), Wright, Edin, New Phil. Journ. ix. p. 106 (1859).
Atractylis, Hincks (1868).
Wrightia, Allman (1872),
Hydrocaulus erect, unbranched, arising from a creeping
hydrorhiza. Perisare expanding above to form a protective
sheath which clothes the hydranth up to the base of the
tentacles.
Reproduction by fixed sporosacs, which are partially or
wholly invested by a chitinous envelope.
Wrightia coccinea (Wright).
Atractylis coccinea, Wright (1861).
Perigonimus (?) coceineus, Hincks (1868).
54 On the Atractylis coccinea of 7. S. Wright.
Trophosome.—The hydrocaulus consists of a number of
short stems }—} inch in length (slender, somewhat contorted,
unbranched, or with small offshoot bearing a hydranth), which
arise at short intervals from a
creeping and anastomosing
hydrorhiza, which resembles
the stems. The hydranth
makes an obtuse angle with
the stem. It is closely in-
vested up to the roots of the
tentacles by a hydrothecal ex-
pansion of the perisarc, but is
not retractile into it. The
tentacles are 10-12 in number,
short and straight, disposed in
a single verticil round the
bluntly conical hypostome.
The perisare is straw-coloured,
and is wrinkled transversely
where it expands to cover the
hydranth. The colour of the
hydranth is pink, turning to
white at the tip of the hypo-
stome. The tentacles are
translucent white ; the cceno-
sare pink to scarlet.
Gonosome.— The gonophores
aresporosacs, ‘hey are globu-
Jar and arise from the bydro-
rhiza, to which they are at-
tached by a slender pedicel.
They are invested by a
chitinous covering which is
continuous with the perisare.
There is a short blunt spadix,
in the outer layer of which the gametes are matured.
The gonophore resembles in structure that of Garveda
nutans, as figured by Allman (‘Gymnoblastic Hydroids,’ i.
. 44).
The colour of the gonophore is translucent white; the
spadix is brick-red ; the gametes pink.
Wrightia coccinea was taken by Wright at Inch Garvie,
Firth of Forth, on the roots of Laminaria saccharina, and by
the writer at Hunterston Perch, near Fairlie, Firth of Clyde,
in 12 fath., on Tubularia, and in Castle Bay, Little Cumbrae,
in 15-20 fath., also on Zubularta. It is common where it
Wrightia coccinea.
On the Generic Position of Benson’s Helix hyba. 55
does occur. J obtained my specimens in May and June, and
they bore numerous gonophores.
In studying Wrightia and the allied genera one cannot help
noticing their resemblance to Calyptoblasts. The family
Bougainvilliidse, to which they belong, is practically alone
among Gymnoblastea in possessing a single verticil of filiform
tentacles surrounding a conical hypostome. All the Calypto-
blasts have this conical hypostome and single verticil of
filiform tentacles. Further, many of the genera of Bougain-
villiidee have quite a distinct protective cup for the hydranth,
resembling greatly the hydrotheca of the Calyptoblast. Indeed,
were it not for the fact that these genera are classified with
the Gymnoblastea, their protective cups would receive the
name of hydrothece.
These facts point to the conclusion that a close relationship
exists between the family Bougainvilliide and the suborder
Calyptoblastea. The Bougainvilliide, perhaps, form a
transition-stage between the suborders Gymnoblastea and
Calyptoblastea.
XI.—On the Generic Position of Benson’s Helix hyba and
the Similarity of its Anatomy to that of WKhasiella vidua,
W. T. Blanford. By Lt.-Colonel H. H. Gopwin-AUSTEN,
F.R.S. &e.
Ever since the discovery of this species about 1860 it has
been impossible to locate it in any Indian genus without
considerable doubt. It is apparently very rare. I have never
come across it in the field, and | do not think it is to be
found in many collections. I am informed by Mr. S. F,
Harmer, of the University Museum of Zoology, Cambridge,
that two specimens (Benson’s types) are in the MacAndrew
collection. Fortunately I have recently discovered two
specimens among some other species preserved in spirit by
Mr. W. Theobald, marked “Chamba,” a small State in
the N.W. Himalaya, 8.E. of Kashmir Territory. The one
from which the subjoined description was taken was in a
very good state of preservation. All we knew with any
certainty was that it belonged to the Zonitide, Mr. Theobald
having noted the presence of the mucous gland at the extre-
mity of the foot at the time of capture.
Mr. Benson described the shell in the Ann. & Mag. Nat.
Hist. ser. 3, vol. vii. (Feb. 1861), in his usual excellent way,
and goes on to say :—
56 Lt.-Colonel H. H. Godwin-Austen on the
“ This interesting form, discovered in the mountains near
the Bari Do-ab by J. ‘Doy le Smithe, Esq., F.G.S., of
Madhopore, and kindly communicated to me by his broken
the Rev. Fred. Smithe, of Churchdown, approaches more
nearly to the Nilgherry H. Gwerint, Pfr., than to any Hima-
layan species. It is notable for its shorter vaulted spire, sharp
keel, rather open umbilicus, and sculpture. In one imperfect
specimen, with a higher and more rounded hive-shaped spire,
the keel of the penultimate whorl overhangs the anterior part
of the last whorl.”
We now know that 7. Guerin, Pfr., is a Thysanota in a
very distinct family of Land-Mollusca. The overhanging of
the keel of the whorl I note in these shells, and it produces
the appearance of a thread running with the suture. Theobald
records the finding of a large variety of this species at
* Aijas” (“ Ajjas”’ on the map of Kashmir, 2 m.=1 inch),
in the hills east of the Walar Lake, Kashmir, measuring
171 x162x10mm. ‘This shell appears to have been lost.
‘Yhe specimen Benson described measured: diam. major
14mm., minor 18, axis 7; the one I now figure: 12 x 11:5 x
54 respectively
On removing the shell the visceral sac is pale-coloured,
sparsely and finely spotted with black on the line of the
rectum. The foot is short and very narrow, indistinctly
divided, dark grey ; a distinct mucous gland overhung by a
blunt lobe, peripodial grooves, and a well-marked fringed
margin to the foot. ‘There is a small but distinct and serrated
right shell-lobe, which would spread further in life ; the lett
isa marginal band on the edge of the peristome. The genera-
tive organs were of great ‘interest as ey were unfolded.
‘The amatorial organ is of the usual form; the penis retractor
muscle given off from a distinctly coiled czcum at the head
of the main penis-sheath. There is a long epiphallus thence
to the junction of the vas deferens, with a long kale-sac
adjacent in which a spermatophore had been developing.
The sperm atheca is very long, gradually enlarging to the
distal end, and contained hives « spermatophor es; these are of
the type I have described in various species of CEO
Austenia, &c. ‘The flume had no large spines, but ver
minute ones could be detected on the edges of it. This being
a single specimen, which I have mounted in gly cerine-jelly,
I did not like to destroy the sac of the spermatheca to get a
spermatophore out entire. In separating out the lengthened
genitalia of a close-wound shell such as this is, it is not easy
to do so without occasionally something breaking away. The
oviduct is peculiar, very straight up to the albumen-gland,
Generic Position of Benson's Helix hyba. D7
am.or
Fig. 1.—Khasiella hyba, Bs.; shell. x 4:8.
Fig. 2.—Side view of mantle-edge from right side, showing the obscure
right shell-lobe, foot, &e. x 8,
Fig. 3.—Genitalia. x 4:5.
Figs. 3a, 36.—The penis, as seen from two opposite sides, to show the
coiled czecum and position of the retractor muscle attachment, the
kale-sae or flagellum, &e. x 8.
Fig.3¢e—A portion of same organ more enlarged, slightly pressed
between two glass slips and viewed by transmitted light, to show
the close-coiled ceecum.; the dark portion is part of a spermato-
phore. \ x 12:
Fig. 5d.—Terminal end and distal end of an organ not located in the
genitalia and with which it may have no connexion, X 12.
Fig.4.—Jaw. xX 24.
Fig. 5.—Central tooth and three admedian teeth of the radula. x 368.
Fig. 5a.—Kighth to eleventh admedian and sixteenth to eighteenth
lateral teeth.
ep. Epiphallus. | pr. Prostate.
k. Wale-sac or flagellum. | rsl. Right shell-lobe.
vd. Vas deferens, | f. Foot.
ce. Crecum, vs, Visceral sac.
p. Penis. ov. Oviduct.
rmp. Retractor muscle of penis,
am.or. Amatorial organ,
58 = On the Generic Position of Benson's Helix hyba,
so was the line of the prostate; but lying nearly parallel to
this I observed a long narrow ribbon—no doubt a duct—lying
on the surface of the jelly-like oviduct, of a hard nature and
ochraceous in colour, clearly pointed and with a fine retractor
muscle, while floating free among the parts of the genitalia
was a similar-sized duet with a swollen open end, where it
had evidently broken away. ‘Vhis may be an accessory gland
of some kind given off from the free oviduct or base of the
spermatheca. It might very easily have been set down as a
spermatophore, but fortunately in this case the spermatophore
of this species is before me, and narrows its possible function
and connexion with the genitalia considerably. We must
wait for more material to clear up this point.
The second specimen, the shell of which is here figured
(p. 57), I do not like to destroy, as the species would appear
to be so rare.
The jaw is strong and solid, slightly arched into a central
projection.
The radula has the formula
NO elo a lO
20 elie Zee
The teeth are of the usual form in so many genera of the
Zonitidse, the laterals being bicuspid, with the outer cusp
below the inner, becoming very small on the margin.
On comparing these anatomical details with those of other
Indian species, I find there is a remarkable similarity to those
of the genus Ahasiella (Godwin-Austen, Moll. Ind. vol. ii.
p- 129, pl. ¢. figs. 1-5 d@) as seen in the type species vtdua,
W.T. Blanf. ‘There is (1) the same small obscure right
shell-lobe ; (2) same form of foot and mucous gland; (3) the
jaw and radula are precisely alike; (4) the generative organs
differ in no appreciable way, merely that the short free cecum
retractoris penis of vidua becomes a close-wound coil in hyba,
and is thus similar to the same part in ALacrochlamys indica.
It is extremely interesting to find such close resemblance
in the anatomy of two land-molluscs with such very distinct
forms of shell as presented in hyba and indica; differing so
widely, conchologically they would take their place in separate
genera. The shells of vidua and hyba also present at first
sight considerable differences, but the variation becomes less
apparent when hyba is compared with the sharply keeled
species of Ahasiedla, such as climacterica, Bens., and Austeni,
W. T. Blanf.
I think I am right in considering //. hyba by its anatomy to
Descriptions and Records of Bees. 59
belong to the genus Khastella, with these shell-characters :—
Openly umbilicated; sharply keeled ; spire very conoid, with
sides and apex very convex.
Nore, Godalming,
oth June, 1907,
XII.—Descriptions and Records of Bees.— XV.
By T. D. A. CocKERELL, University of Colorado.
Triepeolus Nore, sp. n.
? .— Length about 83 mm.
Black, with the usual markings only slightly yellowish ;
hair on middle of face pure white; legs clear red, spurs of
middle and hind legs black ; no red colour on thorax ; tegulee
apricot-colour; scutellum low, obscurely bilobed, the lateral
teeth black and short. Clypeus shining, with many very
minute punctures and a few large ones; mandibies red except
at base and apex; labrum black, a little reddish at sides ;
first three antennal joints and base of fourth ferruginous;
sides of vertex with large well-separated punctures on a
shining ground ; mesothorax exceedingly densely punctured,
the two median stripes of hair short, not attaining the ante-
rior margin; pleura hairy in fiont and witha large transverse
patch of hair ; lower part of pleura densely punctured, but
at one place a little of the shining surface shows; dark trans-
verse mark on first abdominal segment essentially as in
T. occidentalis, but the posterior band interrupted; segments
2 to 4 with even entire bands, that on 2 with lateral processes
directed forwards so as to make with the band an angle of
about 45°; sides of fifth segment with white tomentum ; last
ventral segment normal.
In nearly all respects exactly like a small edition of
T. occidentalis, but the hair of the abdomen is less yellow,
the pygidial patch is narrower, and the scutellar teeth are
less developed. In size and general appearance it closely
resembles 7’. callopus, Ckll. ‘The colour of the spurs will
readily separate it from 7’. Hopkinst.
Hab. Mesilla Park, New Mexico, at flowers of Spheralcea
lobata, Wooton, May 16 (Ikiss Nora Newberry).
Triepeolus remigatus (Fabr.).
Mesilla, New Mexico, June 30 (Cockerell).
This is the true remigatus, as defined by Cresson and
60 Mr. T. D. A. Cockerell—Desertptions and
Robertson, and is new to New Mexico. The species has a
wide range eastward. Mr. N. Banks sends me specimens
from Glenearlyn, Virginia, July 26, and Falls Chureh,
Virginia, Aug. 7, 2 at “flowers of Eupatorium linearifolium.
Triepeolus agaricifer, sp. 0.
3d .—Length 9 mm. or rather more.
Black, with the markings orange-buft, all the abdominal
bands of the same colour ; legs red, the cox and trochanters
black, anterior femora black above except at apex, middle
femora mainly black above and beneath, hind femora black,
with a broad red stripe above; hind tibiw with a large black
patch behind; spurs red. Middle of mandibles red; labrum
black; hair of face orange; antenne black, the third joint
with a large red mark in front; vertex very coarsely punc-
tured ; mesothorax very coarsely punctured, with two very
distinct stripes which reach the anterior margin, which latter
has no light border; markings of thorax as usual; tegule
and most of tubercles red; scutellum entirely black, strongly
bigibbous, with short but very distinct lateral teeth : lower
part of pleura showing a black (but not entirely nude) area,
which is densely and coarsely punctured. Wings with the
apical margin broadly dark fuliginous; stigma red; nervures
fuscous; second s.m. much narrowed above. Abdomen
broad, with all the bands broad and perfectly entire, except
the basal one on the first segment, which is rather broadly
interrupted ; black median mark on first segment transversely
fusiform, with obliquely truncate sides, much shorter than in
T. helianthi, but not triangular as in J’, lunatus; band on
second segment with a broad low prominence on each side in
front, but no projection forming an angle; apical plate
black, narrow.
Looks at first sight like a small TZ. lunatus, but easily
separated by the characters italicized,
flab, Beulah, New Mexico, August (Cockerell).
The name aga icifer 18 derived from the black mark on
the first abdominal segment, which, when the insect is seen
from the front, looks like an agaric.
Epeolus argyreus, sp. n.
g .—Length about 8} mm.
Black, but largely covered with the usual pubescence,
which on the abdomen is pale cinereous with a yellowish tint,
on the thorax above the same, but on the face and pleura
(both of which are entirely mrad densely covered) brilliant
Records of Bees. 61
silvery white. Eyes pale green. Antenne brown-black, the
third joint partly red. M:sothorax covered with appressed
hair, except a somewhat U-shaped patch posteriorly ; area of
metathorax nude, surrounded by dense hair on all sides;
tegule very hairy, piceous, with reddish margins. Wings
hyaline; in the type the upper half of the second t.-c. 1s
wanting on both sides. Legs black, with silvery hair, that
on inner side of basitarsi golden ; small joints of tarsi ferru-
ginous ; spurs of middle and hind legs black ; first abdominal
segment covered with light hair, except a slenler transverse
band, which is only moderately long and is ill-defined at the
ends; all the light bands entire and broad, that on second
segment greatly enlarging laterally, but the enlargement
forming with the band a very obtuse angle; erect fringe on
fourth and fifth ventral segments fuscous.
Hab. North Yakima, Washington State, Aug. 4, 1903
(Eldred Jenne).
E/peolus bihamatus, sp. n.
g-—Nize, colour, and general appearance agreeing with
E. argyreus, with the same silvery hair covering the pleura,
yieuey not quite so densely, and also on the ‘face, though
failing below, so as to leave the lower part of the very minutely
and densely punctured clypeus visible. The black spurs also
are the same, and the hyaline wings. ‘he following differ-
ences are ree :—Maniibles with more red; margin of
labrum red ; face less narrowed below; third Site joint
longer ; flagellum ferruginous beneath except at base ; meso-
thorax less ‘hairy, leaving a large anchor-shaped black area
very densely punctured, its stem reaching the anterior margin ;
tegulz piceous basally, red in the middie, and with hyaline
margin; area of metathorax almost entirely covered with
hair; marginal cell less slender; second t.-c. complete ;
knees, femora, and tibise red; transverse band on first abdo-
minal segment large and clean-cut ; band on second segment
with a large hook-like process on each side ; erect fringe on
fourth and fifth ventral segments white.
Hab. North Yakima, Washington State, June 26, 1903
(Eldred Jenne).
This species and the last are very distinct by the general
appearance and hairy pleura, together with the black spurs,
from the other American members of the genus.
The following three species of Zrtepeolus superficially
resemble J’ Renata and helianthi, and were confused with
?
62 Mr. T. D. A. Cockerell—Deseriptions and
those species until critically examined. A table is offered to
facilitate their separation :—
Females.
Black mark on first abdominal segment distinctly
triancular, the lateral corners pointed ...... lunatus, Say.
Black mark on first abdominal segment a trans-
VERSO MUAM igus stercgs bese oan acme Ore ee eae toe if
dL eAMUeTIOTetem ora ted pie me see wilersktoen ies eneoke segregatus (Ckll.).
Anterior femora biack, or black with ared stripe. 2.
2. Pleura hairy all over, though more thinly below. Townsend, Ckll.
Lower part of pleura bare.............5-000.- helianthi, Rob.
Males.
All, or nearly all, of the abdominal bands narrowly
ATIFETMUP LCA” qarere er-yaiciae etter aa eines eusat sublunatus, Cll.
The bands beyond the first segment all, or nearly
allientire: ee kercemiate pele eens Thy
1, Larger; anterior femora black; pleura hairy all
OVER ce neers sities Meter eine een Frasere (Ck1l.).
Smaller; anterior femora red and black, or red.. 2.
2#Ploura deity iallvoyer se near tee an ete «ieee where Townsendi, Cll.
Lower part of pleura bare or little hairy ..,... segregatus (Ckll.).
Triepeolus sublunatus, sp. n. (Cressont’, Rob., race ?).
g .—Leneth 11 mm.
With a relatively long and narrow, cylindrical abdomen.
Hair of face distinctly yellowish. Labrum, basal part of
mandibles, and first three joints of antenne red ; the two
stripes on mesothorax very clear and distinct; no band along
anterior border of mesothorax, or patches of hair in the ante-
rior corners; tegule bright ferruginous. Wings dusky ;
lower part of pleura with a bare, very densely punctured
patch. Legs red, the anterior femora black, with the apex
and the lower edge red; middle and hind femora with much
black in front; spurs of middle and hind legs black ; trans-
verse band on first abdominal segment with broad anterior
and posterior projections, the latter dividing the apical light
band ; all the abdominal bands practically of the same colour.
By the colour of the tegulee this species is like 7. occidentalis,
Cresson, but it differs from that by the dark femora and inter-
rupted abdominal bands. In Robertson’s table(Canad. Entom.,
Oct. 1903) it runs to Z. Cressonii, Rob., and agrees with the
diagnosis there given, except as to the pleura. According to
Robertson, Cresson? as found in Illinois is exceeding variable,
and it may be that sublunatus represents a geographical form
Records of Bees. 63
of it*. In our insect the scutellum and tubercles show no
red whatever, and the teeth at the sides of the former are
very little developed. The second abdominal segment has
the band hooked at the sides in one specimen, but the hook-
like projection is wanting in the other, which is certainly
conspecific. The marking of the mesothorax is very different
from that of Hrasere, which I described as a subspecies of
Cressonit.
Hab, Dripping Spring, Organ Mts., New Mexico, Aug. 10
(C. H. T, Townsend). Two males.
Triepeolus segregatus (Ckll.).
3g .—Length about 10 mm.
Compact; labrum black and mandibles with little red;
clypeus very densely punctured; hair of face silver-white.
Antenne black, or scape with a red spot in front, and third
segment and base of fourth red; the two stripes of meso-
thorax very distinct; no pale anterior border; scutellum
entirely black, the lateral teeth pointed but not long; pleura
with a nude patch below, on which are scattered strong
punctures on a shining ground; femora entirely red, or with
suffused dusky patches.
¢ .—NScutellum entirely black ; the abdominal bands beyond
the first segment entire, or that on the second inclined to be
interrupted.
The female ascribed to segregatus in the original descrip-
tion probably represents a distinct species, or at least variety.
I was misled by it into thinking the present insect distinct
from segregatus, but I cannot see that the males are anything
but true segregatus, and the females taken at the same time
and place certainly belong with them. The insect is related
to T. pectoralis, Rob., but the black band on the first abdo-
minal segment is not so long, the tegule are differently
coloured, and there are other differences. The spurs are
brown, not black.
Hab. Dripping Spring, Organ Mts., Aug. 10, 2 ¢,2 2
(C. H. T. Townsend).
Triepeolus Townsend, sp. n.
In size, colour, and markings just like the 7’. segregatus
from the Organ Mts., but differing as follows :—
* However, FE. Cressonii was based primarily on the mercatus of
Cresson (1878), and Cresson’s full description appears to indicate an insect
specifically distinct from swb/wnatus. Virginia is to be taken as the type
locality.
64 Mr. T. D. A. Cockerell—Descriptions and
3 .—The two stripes of mesothorax broader, broadly
reaching anterior margin, and more or less connected with
large patches of pale hair which occupy the anterior lateral
corners ; pleura densely covered with hair ; teeth at sides of
scutellum somewhat longer and appreciably curved inwards ;
spurs of middle and hind legs black; black band on first
abdominal segment longer, rounded instead of obliquely
truncate laterally, and without an anterior projection breaking
the basal pale band; angles formed by lateral processes of
band of second segment less acute; apical plate rounded.
The antenne and labrum are entirely black, and there is no
red about the thorax.
? —Antenne black ; anterior lateral corners of mesothorax
with pale patches ; teeth at sides of scutellum smaller; pleura
hairy, the lower part densely punctured. Wings hardly so
dark; spurs black; apical pale band of first abdominal
segment more broadly interrupted, but basal band not
interrupted.
Hab. Rio Ruidoso, New Mexico, about 6700 ft., at flowers
of Hrigeron macranthus, July 27, @ (C. H. T. Townsend) ;
Rociada, New Mexico, Aug. 8, 2 (Cockere/f).
ISEPEOLUS, gen. nov.
$ .—Similar in appearance and markings to Hpeolus, but
differing as follows :—Third antennal joint very long, longer
than the scape; apical half of galea slender and elongated ;
labial palpi very long (the last two joints minute as usual) ;
axillar teeth present but rounded, blunt; b. n. falling far
short of t.-m. ; submarginal cells subequal, but the third the
largest ; the second large, square, a little shorter below than
first or third, receiving first r.n. near its end; third s.m.
receiving second r. n. near its end; outer side of third s.m.
strongly bulging; marginal cell. with its apex obliquely
truncate, the tip away from the costa, the part of the lower
edge joining the s.m. cells a trifle shorter than the part
beyond; apex of abdomen rounded, without a_ projecting
plate, the upper surface of it covered with fine hair. The
maxillary palpi appear to be quite as in Epeolus. Claws
bifid; a very long pulvillus, as long as the claws, with a
black apical disk.
For years I have had this insect marked ‘ Leiopodus?” ;
but having seen the type of Lezopodus in the British Museum,
it is evident that Zsepeolus is very distinct by the oval abdo-
men, large but not especially peculiar hind coxe, second s.m.
a little narrower below than first, &c.
Records of Bees. 65
. Isepeolus albopictus, sp. n.
3 .—Length about 82 mm.
Black, with markings of white tomentum like those of
Hpeolus; mandibies rather feebly bidentate, dark reddish
except at base; labrum black; face covered with white hair,
but clypeus nearly all bare, dull and roughened with very
minute punctures ; apex of scape, and the following three
joints entirely, red; the remaining part of the flagellum,
which is stout, red beneath; mesothorax dull, with minute
close punctures, anteriorly with two triangles of black hair
surrounded by white, the anterior lateral corners also covered
with white hair; tubercles red ; upper part of pleura covered
with white hair, lower part bare, densely punctured ; scutel-
lum bigibbous, red, with black hair at sides; postscutellum
red, with white hair; metathorax black, the area bare; tegula
reddish. Wings clear, the apical field with dusky stains ;
stigma well developed, red; nervures dark fuscous. Knees,
anterior and middle tibia, apex and base of hind tibiz, and
all the tarsi red ; anterior tarsi long and slender; anterior and
middle tibize marked with two bars of white on the outer side,
hind tibie with a broad bar or patch on apical half; hind
femora swollen ; spurs ferruginous ; first abdominal segment
with white hair at base, a spot at each side, and a pair of
semicircular marks on hind margin; second segment with a
broadly interrupted band on hind margin, its proximal ends
subclavate, its upper lateral margin with a very large bifid
process or extension ; third segment ornamented like second,
except that the band is obsolete in the subdorsal region,
leaving the proximal ends as spots; fourth segment with a
pair of spots; fifth with two transverse hammer-shaped
marks; sixth four-spotted ; apex red; venter dark brown,
with some white hair-spots.
Hub. Carcarana, Argentina; received from Mr. J. C.
Crawford. Collected, I believe, by Professor L. Bruner.
I believe there are other specimens in the U.S. National
Museum.
GRONOCERAS, gen. nov.
Large bees allied to Megachile, with the nesting-habits
(cf. G. combusta) of Chalicodoma: male with flagellum exca-
vated beneath, like a trough ; abdomen with two long spines
at apex; anterior coxee with very long spines ; anterior tarsi
pallid, broadened and flattened ; claws bifid at apex : female
with mandibles 2- to 4-dentate ; scopa red or red and black ;
claws simple. ‘lhe maxillary palpi are bristly.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 5
66 Mr. T. D. A. Cockerell—Deseriptions and
The general build resembles that of Chalicodoma, but the
structure of the apex of the abdomen and of the male antenn
is quite different.
Type, G. Wellmani, sp. n. Also including G. combusta
(Megachile combusta, Sm.), G. guineensis (Apis guineensis,
Fabr.), G. tricolor (Megachile tricolor, Friese), and G. stuppea
(Megachile stuppea, Vachal).
Gronoceras Wellmant, sp. n.
? .—Length 17-19 mm.
Black, including legs and antenna, the hind margins of the
abdominal segments ferruginous ; hair of head, thorax, and
legs black or brown-black, of abdomen bright fox-red, above
and below, but largely black or dark fuscous on the first
segment above; mandibles long, with two apical teeth, the
rest of the cutting-edge toothless ; clypeus normal, strongly
punctured, the punctures well separated in the middle ; meso-
thorax densely punctured and dull; tegule densely punctured,
black, with dark ferruginous margin. Wings subhyaline,
strongly yellowish.
Very nearly agreeing with G. combusta, but the wings are
not nearly so dark.
é .—Lenegth 16-19 mm.
Like the female, except in the sexual characters mentioned
in the generic diagnosis, but the hair of the first abdominal
segment is red like that of the others (in combusta ¢ it is
black); the hair of the face is light yellowish or yellowish
white, with black hairs intermixed ; the cheeks below have
a fringe of long white hairs; the anterior tibia are more or
less pale apically, with a little elevated keel in front; first
basitarsus broad and flat, a sort of pale honey-colour, with a
raised ferruginous keel or line along the outer anterior edge,
short white hair on the outer surface, except basally and more
or less anteriorly, where there are fuscous bristles, a fringe
of dense short black hair on the anterior interior edge, and
a long fringe of ferruginous-tipped hairs behind; second
joint of anterior tarsus dark reddish or yellow, the remaining
joints black, except that the tip of the last is red ; extreme
apex of middle tibia red; bases of claws red; red hind
margins of abdominal segments very broad; venter of abdo-
men mainly red; apex of abdomen with some long dark
hairs, and two long, straight, red-tipped spines, mesad of
each (and springing from the same base) is a short incon-
spicuous spine; a red spine, easily overlooked, on each side
of penultimate segment; anterior coxal spines long and
Records of Bees. 67
straight, reddish above. The second specimen, manifestly
conspecific, has at the apex of the abdomen a pair of short
black spines on one side and a single one on the other ; it is
asymmetrical and evidently abnormal. This male is also
very like G. combusta, but separated by the much paler
wings, red hair on first abdominal segment, form of apical
teeth, &e.
Hab. Portuguese West Africa, long. E. 15° 05', lat. S.
12° 44’, alt. 1360 metres, at flowering mint, Molanthus sp.,
Dec. 1906 (middle of rainy season) ; two of each sex collected
by Dr. F. Creighton Wellman.
Gronoceras benguellensis, sp. n.
2? .—Length about 16} mm.
Black, including legs and antennee ; hair of head, thorax,
and legs brown-black, except that there is a little pallid hair
about the bases of the antenne, and the hair of the thorax
above is dark coffee-colour on the mescthorax, becoming
bright fox-red on hind part of scutellum and upper part of
metathorax ; the hair of the first two abdominal segments
above is bright fox-red, but on the ethers blaek, the hind
margins of the segments narrowly whitish ; ventral scopa red
in the middle and black at the sides, but entirely black on the
last two segments. Wings dusky hyaline, the apex clouded ;
hind spurs red; mandibles 4-dentate, the third tooth trun-
cate, the fourth very small.
¢.—Length about 13 mm.
Hair of face yellowish white, of cheeks below white, of
vertex and occiput brown-black, of thorax and abdomen
brown-black, without red; whitish at sides of abdomen
beneath ; apex of abdomen with a large stiff tuft or brush of
long black hairs ; abdomen above practically bare, the hind
margins of the segments dark reddish; clypeus densely
punctured, with a narrow shining median raised line ; middle
of mandibles with a large tubercle beneath Gia Wellman with
a similar process, but hardly so large); labrum broadly
rounded at apex; anterior tarsi formed in general as in
Wellmani, but the apical joints are red, not black, and the
basitarsus is greyish, with no anterior ferruginous line, but
with a broad cream-coloured stripe down the anterior margin,
and continued on to the second joint; the fringe of hair on
the inner anterior edge is pale reddish instead of black, and
the long posterior fringe extends as far as the penultimate
joint, and is fuscous for its upper half, ferruginous with a
white base for its lower, ¢.e. from the last quarter of the
ry
68 Mr. A. H. Verrill on the
basitarsus on; anterior coxal spines well developed ; spines
at apex of abdomen long and straight, without any inner
pair; the short sublateral spines are black.
Hab. Same locality and date as G. Wellmani; taken by
Dr. Wellman at flowers of Composite, one of each sex.
Megachile caricina, sp. n.
? —Length about 103 mm.
Black, with broad head and broad shovel-shaped abdomen ;
hair of face pale yellowish, of cheeks white, of occiput pale,
but about ocelli short and black; antenne entirely black ;
mandibles 4-dentate; clypeus normal, strongly punctured,
with a smooth, shining, discal area; hair of thorax white at
sides and beneath, above black, with some whitish on meso-
thorax in front and in mesothoracico-scutellar suture ; meso-
thorax and scutellum very densely punctured; tegule black.
Wings strongly dusky. Hair of legs white, pale reddish on
inner side of tarsi; hind basitarsus very broad and flat ;
claws simple; abdomen punctured, not strongly or closely,
and without bands; apical segments above with coarse
black bristles ; scopa bright orange-red, but white basally
and black on last segment.
3 .—Size about the same, as also general appearance.
Face densely covered with yellowish-white hair, a few dark
hairs on each side near upper part of clypeus; black hairs on
vertex ; antennz black ; anterior tarsi quite simple ; anterior
coxe with rather short but stout spines; metathorax and first
abdominal segment with copious white hair; apex with a
strong transverse keel, which is broadly emarginate but not
serrate ; no subapical ventral teeth; claws bifid at end.
This belongs to Megachile, s. str., as defined by Friese and
Robertson. In its general appearance it is much like the
American Jf. mendica, Cresson.
Hab. Same locality and date as Gronoceras Wellmani ;
2 6,1 ¢, taken by Dr. Wellman, the males marked “ on
flowering sedges, side of stream,” the female ‘‘ on sedges.”
XII1.—WNotes on the Habits and External Characters of the
Solenodon of San Domingo (Solenodon paradoxus). By
A. Hyatt VERRILL *.
[Plate IV.]
AurHouGH Solenodon paradoxus of San Domingo and Haiti
was discovered and imperfectly described as early as 1839,
* Reprinted from the Amer. Journ. Sci. for June 1907. From an
advance proof communicated by the Author.
Solenodon of San Domingo. 69
several years before the Cuban species (Solenodon cubanus)
was known to science, it is still practically unknown to recent
zoologists. The published descriptions of this rare and inter-
esting mammal are vague and unsatisfactory. For many years
it has been commonly considered extinct, and when, in Decem-
ber 1906, I undertook a collecting-trip to San Domingo with
the avowed intention of obtaining the Solenodon, prominent
zoologists stated that the quest was hopeless, one of them
saying that I would be as likely to secure specimens of ghosts
as of Solenodon paradocus.
During the five months spent on the island I devoted a
great deal of time hunting for the Solenodon and in inter-
viewing natives from the remote and little-known parts of
the island.
I soon found that the animal was well known to the natives
in certain isolated localities, but that over the greater portion
of the Republic it was absolutely unknown.
This is readily accounted for by the presence of the mon-
goose in most parts of the country, and. it is only a question
of time when this pest will overrun the entire island and the
Solenodon will become actually exterminated.
The natives have several names for the Solenodon, calling
it ‘ Orso,” “ Milqui,” ‘‘ Homigero,” and Juron,” while the
English-speaking negroes from the British West Indies know
it as ‘Ground Hog.” The name- “ Juron” (ferret) is also
applied to the mongoose, and. for some time I was misled by
this confusion of the two animals. In its habits the Solenodon
resembles a hog, rooting in the earth and cultivated grounds,
tearing rotten logs and trees to pieces with its powerful front
claws, and feeding on ants, grubs, insects, vegetables, reptiles,
and fruit, and at times proving destructive to poultry. On
several occasions it has been known to enter the houses in
search of roaches and other vermin, and has been captured in
rat-traps.
It is strictly nocturnal, and spends the day in caves, holes
in the coral-limestone rocks and in hollow trees and logs. It
is a slow, stupid creature. It is unable to run rapidly, but
shambles along with the zigzag sidewise motions of a planti-
grade. It is doubtless owing to this that it obtained the
native name of “ Orso ”’ (bear).
Its long snout and stout front feet, with their curved claws,
and its thick short neck prove impediments to forward progress,
According to the natives, it is incapable of running straight.
They also claim that when pursued it frequently trips itself
and tumbles heels over head. When hunted with dogs, it
thrusts its head into the nearest hole or shelter, and allows
itself to be captured without resistance.
70 Mr. O. Thomas on
The only specimen that I obtained was a female, which
was captured alive and uninjured. A few days after its
capture it gave birth to three naked young. These the
mother promptly devoured, and she died three days later.
This specimen {see Pl. IV.), as preserved in formol, is
14 inches in length, exclusive of the tail, which measures
about 13 inches in length.
The body and head are covered with sparse coarse hair,
which is reddish ferruginous from the eyes to the shoulders
and dusky brown on the rest of the body.
The hair becomes very thin and scattered on the hind-
quarters, which for some distance on the back and sides are
naked, roughly corrugated, and warty, with a sparse, short,
woolly growth between the excrescences.
The legs, snout, and eyelids are naked, and, with the bare
skin of the rump, are pinkish white. The ears are short,
thin, rounded, and are bluish grey with light edges. ‘The
heavy rat-like tail is dark brown and naked. The claws are
horn-colour. The front feet and claws are large, heavy, and
mole-like, and well adapted to digging and tearing asunder
rotten wood &. They are much smaller in proportion than
in the Cuban species, however. The snout is also more
flexible than in S. cubanus, trom which it also differs im
the naked skin of the rump, the colour, size, and other
characters.
XIV.—On Three new Mammals from British New Guinea.
By OLDFIELD ‘THOMAS.
In a further consignment of small mammals presented to the
National Museum by Mr. C. A. W. Monckton, Resident in
Northern British Guinea, there are examples of the two
following new Rodents, one of them forming a new and most
striking genus allied to Hydromys, but even more highly
specialized for an aquatic life.
‘The collection also contains specimens referable to Macro-
glossus australis, Pseudochirus corinne and Forbest, and
Phalanger carmelite, all being valuable acquisitions to the
Museum.
In determining the last-named animal, a new Phalanger
allied to it has been noticed and is now described. It was
obtained by Mr. A. Meek.
Crossomys, gen. nov. (Hydromyine).
A highly specialized aquatic form. Fur thick, soft, and
new Mammals from British New Guinea. 71
very woolly. Har-conches practically aborted, a mere rudi-
ment, 1-2 mm. in length, being all that is left of them.
Whiskers not so thick or long as in the allied forms.
Fingers free, toes rather more broadly webbed than in
Hydromys ; claws, both fore and hind, small, delicate, strongly
curved ; hind feet broad, more twisted than in Hydromys,
those of Parahydromys * being less so; sole-pads broad and
smooth, a large part of the elongate hallucal pad visible in an
upper view of the foot. Tail provided with a strongly
marked swimming-fringe below, formed of hairs about 8 mm.
in length, the fringe bifureating into two lateral ridges on
the proximal inch of the short-haired part of the tail.
Skull with a proportionally short slender face and very
large, smoothly rounded, broad and low brain-case. The
distance from the supraorbital foramina to the occiput is
therefore greater instead of less than that to the tip of
the muzzle. Nasal and interorbital region slightly built, not
broadly swollen as in Parahydromys. Cranial ridges
practically absent. Interparietal sutures almost obsolete in
the type, which is an old specimen. Structure of ante-
orbital and palatal foramina as in Hydromys, the latter not
so far forward as in Parahydromys. Bullee very small, in
correlation with the abortion of the external ear-conches.
Dentition as in Hydromys. Upper incisors narrow, con-
siderably bevelled laterally. Molars small in proportion to
the size of the animal; in structure like those of /Hydromys
except that the laminz are more directly transverse and the
middle lamina of m! is scarcely broadened internally.
Type Crossomys Moncktont.
This beautiful animal forms a most striking new genus,
and Mr. Monckton is to be congratulated on its discovery.
In specialization for an aquatic life it far surpasses Hydromys,
as indicated by its woolly fur, aborted ear-conches, twisted
hind feet, and fringed tail, in which last character it re-
sembles the European water-shrew (Veomys, long known
as Crossopus). Indeed in the accumulation of these characters
it stands at the head of all rodents, for while Fiber has
an even more specialized tail and the same fur and feet, it has
retained its ear-conches. Perhaps the nearest analogue to
* Described as Limnomys, Ann. & Mag. N. H. (7) xvii. p. 325 (1906).
This name being preoccupied (Mearns, 1905) the genus was given (Zool.
Anz. xxx, p. 326, 1906) the clumsy name of Parahydromys by Poche,
to whom a perusal of page 14 line 10 of the Stricklandian Code of
Nomenclature (1863) is to be recommended. My own substituted name
of Drosomys (P. Biol. Soc. Wash. xix. p. 199, 1906) was a few months
later in date.
72 Mr. O. Thomas on
Crossomys, both in structure and habits, 1s Anotomys, from
the torrents of the high Andes, which has altogether lost its
ear-conches, has equally aquatic fur and feet, but in which
the tail, although longer-haired below, has not such a
specialized swimming-fringe.
The structural modifications of the Beaver are of so different
a nature as hardly to come into the comparison.
Crossomys Moncktoni, sp. n.
Size considerably less than in //ydromys. Fur soft and
glossy, the comparatively sparse longer fur of the back about
18 mm. in length, the wool-fur very thick, soft, and close,
about 10-11 mm. in length, resembling that of Fiber.
General colour above grey (grey no. 6), washed on the
back with pale yellowish olivaceous, the longer hairs with
black tips and isabella subterminal rings; the wool-hairs
silvery white or greyish white for seven-eighths their length,
their tips black. Under surface silvery white, the line of
demarcation well-marked, high up on the sides, the hairs
white to their bases or very faintly greyer below the tips.
A narrow line down front side of arms like back, the
remainder white; upper surface of hands pale brown, the
fingers naked. Feet practically uaked, the few minute hairs
glossy white. Tail grey throughout on the short-haired
part, the swimming-fringe white.
Skull and teeth as described above.
Dimensions of the type (measured in the skin) :—
Head and body 205 mm.; tail 220; hind foot (wet) 48.
Skull: greatest length 40°5 ; basilar length 31:5; zygo-
matic breadth 22:2; nasals 11°45; interorbital breadth 5:8 ;
breadth of brain-case 19°7 ; height of brain-case from basion
12; palatilar length 17:3; diastema10°8; palatine foramina
5; length of upper tooth-row 5:3, of m’ 4:2.
Hab. Serigina, Brown River, N.E. British New Guinea.
Altitude “‘ not less than 4500 ft.”
Type. Adult female. B.M. no. 7.5. 22.3. Original number
36. Collected 12th October, 1906, and presented by C. A. W.
Monckton, Esq.
“Tris dark brown. Caught while swimming down a
rapid creek.”—C, A. W. M.
Uromys anak, sp. n.
A very large species with a wholly black tail.
Size largest of the genus. Fur harsh; longer hairs of
back 28-81 mm, inlength, underfur about 20 mm. General
new Mammals from British New Guinea. 73
colour coarsely mixed grey-brown, becoming bistre on the
middle back and rufous or burnt-umber on the rump. When
first appearing the hairs of the fore-back at least are really
grey, that is ringed with black and white, but owing to the
bleaching of the black to brown, and of the white to bufty
white, the general tone soon approaches bistre. Under
surface mixed brown and whitish, without sharp line of
demarcation, most of the hairs brown with whitish tips, but
a certain number along the median area white to their bases.
Muzzle and chin dark brown. Ears short, naked. Arms
grizzled brown, like body, the inner aspect rather lighter ;
upper surface of hands reddish brown. Hind legs dark
rufous, becoming browner on the metatarsals ; toes naked,
except for the few brown hairs at the roots of the claws.
Tail long, of the usual Uromys structure, wholly black,
except for the short basal furry portion, which is deep
reddish.
Skull large and heavy, but otherwise as in the other
members of this group. ‘The molar series markedly longer
than in any other species.
Dimensions of the type (measured on the skin) :—
Head and body 310 mm.; tail 400 ; hind foot (wet) 69 ;
ear (wet) 24.
Skull: palatilar length 38°5; nasals 27 x 8°6 ; interorbital
breadth 10°3; diastema 23°5 ; palatine foramina 7°3 ; upper
molar series (crowns) 14:2.
Hab. Ifogi, Brown River, N.E. British New Guinea.
Altitude ‘* not less than 4000 ft.”
Type. Old male. B.M.no. 7.5.22.2. Original number 27.
Collected 2nd October, 1906, and presented by C. A. W.
Monckton, Esq.
“Tris dark brown. Native name ‘ Felek.” A ground
animal, living in burrows, though occasionally found in the
hollow of a leaning tree.”—C. A. W. M.
Under the names of Uromys validus*, Hapalotis papuanust,
and Mus barbatus {, three members of this group of the genus
have been described from South-eastern New Guinea, though it
is possible that all of these names, or either two of them, may
be synonymous with each other. I have examined the types
of the first and third, and find that their upper molar series
measure 11°3 and 11 mm. respectively, and each of these has
the terminal portion of the tail yellow.
The case of “Hapalotis papuanus” is not so clear, for nothing
* Peters & Doria, Ann. Mus. Genov. xvi. p. 703 (1881).
+ Ramsay, P. Linn. Soc. N.S.W. viii. p. 18 (1883).
{ Milne-Edwards, Bull. Mus. Paris, 1900, p. 167.
74 On new Mammals from British New Guinea.
is said as to the colour of the tail, and the skull and tooth-
measurements are quite inconsistent with each other. How-
ever, both upper and lower tooth-seriesaresaid to be “0°49 in.”
(=12°5 mm.), and if this be taken as correct the size of the
animal wonld be little larger than in U. validus, and con-
siderably smaller than in UW. anak.
Phalanger sericeus, sp. n.
A dark brown species like Ph. carmelite, the fur very long
and silky.
Size and general characters as in Ph. carmelite, to which
the specimen had been hitherto referred. Fur very much
longer than in that species (hairs of back about 38 mm. in
length instead of 27 or 28), exceedingly soft and silky, quite
unlike the rather coarse close fur of the allied species.
Colour essentially as in carmelita, chocolate-brown above and
pure sharply defined white below, but the brown above is
darker and more glossy, resulting from the comparative
silkiness of the hairs. The median dorsal area blacker than
the sides, but no defined stripe perceptible. A patch of paler
brown just above the base of the tail. Mars very small,
thickly clothed internally as well as externally with short
brown hairs. Tail with the proportions of the hairy and
naked portions about as in PA. carmelite, but the proximal
part of the latter is smoother and less shagreened.
Skull and teeth very much as in Ph. carmelite, except that
the secators, both above and below, are less developed, their
apical ridge, which has three or four distinct crenulations in
carmelite, reduced above to an indistinetly bifid point, and
below to an undivided one. Molars rather narrower than in
carmelite. Coronoid process of lower jaw rather higher
and less slanted backwards than in the allied species.
Dimensions of the type (measured in skin) :—
Head and body 455 mm.; tail 810; naked part of tail
above 170 ; hind foot (s. u.) 57.
Skull: basal length 75; greatest breadth 51; greatest
diameter of upper secator 4°38; combined length of three
anterior molariform teeth 16°95.
Hab. Owgarva, Angabunga River (near the Aroa River),
S.E. New Guinea. Altitude 6000’.
Type. Adult male. B.M. no. 5, 11. 28.23. Collected
30th October, 1904, by Mr. A. Meek. One specimen.
This Phalanger was placed on arrival with Ph. carmelite,
but the further material since received from Messrs. Monckton
and Meek shows the Angabunga specimen to represent
quite a distinct form.
On new and little-known Eastern Moths. ras
XV.—New and little-known Eastern Moths.
By Colonel C. Swinuog, M.A., F.L.S., &e.
Family Eupterotide.
Apona khasiana, nov.
& ?. Of a uniform pale ochreous-brown colour, darker and
more ochreous than in A. pallida, Walker, from Sikkim, but
about the same size; the plumes of the antenne about half
the length and brown in colour; the transverse lines and
bands similarly placed, the medial and discal lines single, the
latter curving inwards in its middle; a black dot at the end
of the cell of fore wings; the underside of both wings with
some large red-brown patches towards the outer margin.
Expanse of wings, g 33%, 9 44% inches.
Khasia Hills.
This is probably the insect referred to by Mr. Elwes as
Apona cashmtrensis, Kollar, expanse 120 mm., from the
Khasis, in Journ. Bo, N. He Soc. xi. p. 250 (1897); but
cashmirensis is a small inseet differing from both pallida and
khasiana in the shape of its bands. I have it from Solon,
taken by Bayne Reed.
Family Arctiidae.
Diacrisia procedra, nov.
3. Above and below of a uniform pale ochreous-buff
colour; antenne and palpi black, frons with black sides ;:
fore wings with two black costal spots, one before and the
other beyond the middle; a black spot at the upper end of
the cell, another close to hinder margin below the first spot,
a spot at the base, the discal band of spots below showing
through the wing with two spots on it on the hinder margin
above, and two dots on veins 2 and 3: hind wings w ith a
large spot at the end of the cell and a discal band “composed
of two very large black spots joined together, each divided by
veins 1 and 2, and a small spot divided by vein 5; cilia
concolorous, without markings ; abdomen pinkish, dorsal and
lateral rows of small black spots. Underside marked as
above, except for a larger black spot at the end of the cell of
fore wings, and the discal band clear and prominent, com-
posed of three large spots divided by the veins and connected
with each other by small black dots; pectus bright ochreous,
with black hairs ; legs with black stripes.
76 Colonel C. Swinhoe on new
Expanse of wings 1,5, inch.
Padang, Sumatra; one example.
Superticially resembles D. punctata, Moore, which I have
from Java.
Diacrisia amilada, nov.*
3. Palpi and frons black ; antennz greyish white ; head,
thorax, and fore wings brownish buff, spots and bands on the
latter black; a basal spot, a spot near the hinder margin at
the middle, another above it, and then in continuation five
small spots in a line obliquely inwards to the costa; three
bands of paired linear marks divided by the veins, the first
from the hinder margin a little beyond the middle obliquely
to the costa a little before the apex, the second running
almost parallel, starting a little before the hinder angle, the
third close to the outer margin, the second spot of the row
being absent: hind wings pale pink, a large black spot at
the end of the cell and a discal band composed of two large
black spots in the lower disk and two in the upper; cilia of
both wings ochreous grey, with black spots on the upper
third ; abdomen pale pink, with dorsal and lateral rows of
black spots. Underside: wings paler, hind wings nearly
white, markings much as above, except for a large black
lunular spot at the end of the cell of the fore w ings; fore
legs scarlet, hind legs grey, all with blackish stripes.
“Expanse ‘of wings 1,8, inch.
Padang, Sumatra; one example.
Chionema rhadota, nov.
@. Palpi and antenne dark orange-colour; frons white,
with an orange spot; head and thorax white, ‘bands on each
side of the thorax scarlet : fore wings white, bands scarlet ;
a subbasal, rather thin, sinuous band not reaching the hinder
margin; a broad antemedial band slightly bent inwards in
the middle, with a thick black line on its inner side; a large
black spot at the end of the cell; a postmedial erect band as
broad as the other, with a thick black line on its outer side
which slightly curves inwards above its middle; a marginal
scarlet band of somewhat lesser width: abdomen and hind
wings pinkish ochreous without markings ; cilia of both wings
yellow.
* This I expect is the insect identified by Piepers and Snellen in their
list of Javan Lepidoptera as D. casigneta, Kollar, a Palearctic species
not unlike it in general appearance, but quite a different insect with black
antenne.
~l
S|
and litile-known Eastern Moths.
Expanse of wings 1,%5 inch.
Sitoli, Nias; one example.
There is a female from Nias in the B. M. with C. javanica,
Butler. J have no doubt when the male is known it will be
found to be quite different to the male of the Javan species.
Family Drepanulide.
Drepana albiceris, nov.
@. Whitish buff-colour; palpi black; antennz orange-
brown ; a black band on the upper half of the frons; legs
brown above; thorax and abdomen without markings: wings
sparsely covered with very minute orange-brown atoms and
a few larger black atoms; a transverse brown band composed
of three lines close together from near apex of fore wings,
where there is a smail brown patch with a pale centre, to the
middle of the abdominal margin of the hind wings ; on the
hind wings the band is accompanied by some slight blackish
suffusion, and is obsolete above vein 6, and at the end of cell
touching the inner margin of the band is a rather prominent
black spot ; on both wings there are submarginal black dots,
close to the margin at the apex of fore wings, widening from
the margin hindwards: on the underside the band is
blackish brown, broad and complete, and there is a similar
band on the outer margin.
Expanse of wings 2,*, inches.
Padang, Sumatra; one example.
In shape this species is like Drepana curvaria, Walker,
from Mysol, figured in Cat. Het. Mus. Oxon. i. pl. vii. fig. 1,
but it is larger and the fore wings more falcate.
Family Lymantriide.
Leucoma ecnomoda, nov.
3S. Palpi bright pinkish orange; head dark chestnut-
colour; front of thorax pure white, remainder of thorax,
abdomen, antenna, and wings rather pale pinkish brown:
both wings with a large irregular hyaline patch, evenly curved
on its inner side at one third from the base, running to a
point below the costa, very irregular on its outer side, with
three square lobes projecting outwardly on the fore wings and
one on the hind wings and occupying the best part of the
wings; on the fore wings above and close to the hyaline
patch there is a duplex hyaline spot, almost subapical; on
the hind wings there is one, with another disconnected patch
outside. On the underside the coloration is pinkish white.
78 Colonel C. Swinhoe on new
Eixpanse of wings 1,%) inch.
1 g, Buitenzorg, Java (type).
1 g, Padang, Sumatra.
Allied to Leucoma fenestrata, Hmpsn., from Ceylon, the
hyaline patches much larger and differently shaped.
Euproctis xanthura, nov.
g. Antenne and frons dull ochreous ; palpi brown above,
white beneath and at the tips; body and wings blackish
brown; thorax orange in front; abdomen with an ochreous
tuft: fore wings with the costal line and a patch at the apex
ochreous white, a prominent black spot at the apex, a little
ochreous white on the outer margin below the middle, and
the broad cilia ochreous white: hind wings without markings,
the cilia pure white. On the underside the inner portion of
the fore wings is pale blackish brown, the base and margins
broadly whitish; the hind wings are all white except for
some blackish-brown suffusion on the abdominal third ;
abdomen blackish brown, thorax and legs whitish.
Expanse of wings 1,5 inch.
Padang, Sumatra ; one example.
Belongs to the scintillans group, but is not very closely
allied to any of the forms of that section.
Euproctis neola, nov.
2. Of a uniform blackish brown ; antennz, palpi, thorax
beneath, legs, and abdominal tuft pale whitish ochreous;
thorax in front dull orange: fore wings with a large apical
bright yellow patch, containing two prominent black spots,
one at the apex and the other below it ; another yellow smaller
patch in the shape of a half-circle on the outer border below
the middle; cilia concolorous with the two colours of the
wing: hind wings without markings, the cilia bright yellow,
with a little of this colour entering within the outer margin.
On the underside the fore wing is the same as it is above,
except that the apical spots are absent and the costal line is
yellow ; on the hind wings there is a marginal yellow band
and yellow cilia.
Expanse of wings 1$ inch.
Padang, Sumatra; one example.
Family Chalcosiide.
Soritia zebra.
Laurion zebra, Butler, Ann. & Mag. Nat. Hist. (5) xiv. p. 85 (1884).
Erasmia laja, Pag. Nass. Ver. xxxvili. p. 11, pl. 1. fig. 5 (1885).
Sitoli, Nias.
and little-known Eastern Moths. 79
Butler’s and Pagenstecker’s types were females. I have
received both sexes from Sitoli. ‘he male differs from the
female in having a narrow discal white band on the fore
wings, narrower than in the male of Soritia obliquaria, Fabr.,
but it is broken into three pieces, the centre piece being a
small white spot ; the black marginal band of the hind wings
is narrower, and the black portions of both wings on the
undersides are covered with blue streaks and spots as in the
female.
amily Notodontide.
Stauropus virescens.
Stauropus virescens, Moore, P. Z. S. 1879, p.404; Hmpsn. Journ. Bo.
N. H. Soe. xiii. p. 42 (1900).
Sikkim.
Moore’s type isa male; I have lately received from Sikkim
what I believe to be the female. The fore wings have the
same kind of green above, with four irregular, dark, trans-
verse bands ; the hind wings are pale brownish grey and the
undersides of both wings are grey and quite unmarked as in
the male; the fore wings are rather broad and the expanse
2,3, inches.
Family Hadenide.
Polia illoba.
Agrotis illoba, Butler, Ann. & Mag. Nat. Hist. (5) i. p. 162 (1878).
Polia illoba, Hmpsn. Phal. y. p. 151 (1905).
Graphiphora pacifica, Butler, /. c. p. 165.
Mamestra declinans, Staud. Stett. ent. Zeit. 1888, p. 250.
Khasia Hills; several examples.
Recorded from Siberia, Japan, and China; not previously
recorded from the Indian Region.
Family Focillide.
Iluza eugrapha, nov.
6 ¢. Of a uniform grey colour, very slightly tinged with
pinkish ochreous and minutely irrorated with black atoms ;
antenne blackish brown ; sides of the palpi and legs black:
fore wings with a black dot in the middle of the cell and
another at the end; a broad straight band from apex of fore
wings to near anal angle of hind wings, obsolete from vein 6
to the costa; this band is composed of a thick black outer
line, a thin black inner line, the narrow space between them
erey-pink, and close to it on the inner side on both wings is
80 Bibliographical Notice.
another fine black line for two thirds upwards ; there is also
a black spot just below the middle and close to the outer side
of the band on the fore wings; a thin marginal interrupted
line and black points close to the outer margin on both wings;
a black dot at the end of the cell on the hind wings. Under-
side paler, no markings; the cell-dots and a Le
festooned black line on the outer margin of both wings, most
prominent in the female.
Eixpanse of wings, ¢ 17%, ? 1,5 inch.
3, Gédé, W. Java.
?, Padang, Sumatra.
Allied to Ituza transversa, Moore, from Sikkim and Assam.
There is a male from Bali in the B. M. unnamed, Quadrifid
drawer no. 222.
Family Boarmiide.
Opthalmodes plesia, nov.
3. Moss-green; palpi and frons ochreous white, palpi at
the sides, head, and antenne black: wings somewhat pale
between the bands, making them more or less variegated in
colour, with four transverse, duplex, green, slightly dentated
bands at equal distances apart, commencing from black spots
on the costa of the fore wings and with black spots throughout
on each inner band, the “outer margin of both wings with
equally prominent black: spots; cilia white, a black spot at
the end of each cell. Underside whitish, suffused with dull
ale green; a black spot at the end of each cell and a dull
blackish submarginal band more or less macular.
Expanse of wings 1,’ inch.
Padang, Sumatra ; one example,
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U1. A Contribution te the Knowledge of the Hymenoptera of the
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eenereerreocee eee eee
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Veep ou. (PETC LEV)! wai jaiddelepte aoe ib cela Min(iete gis ait retekenetdie, sisi meets
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IcuvEvmontn# (continued).
Chiaglas nigripes, Cam.
Chiaglas nigripes, Cam. Ann. & Mag. Nat. Hist. (7) ix. p. 152.
Runjit Valley, Sikkim, 1000 feet; April (C. 7. Bingham).
Described from the Khasias.
In Chiaglas the discocubital nervure is broken by a stump
and the transverse median nervure is received shortly beyond
the transverse basal. The species mimics a M/yermo, it
having the same size and colour-markings as, e. g., the
common J. maculitarsis, Cam.
Faliphera latibalteata, sp. n.
Black ; the face, clypeus, base of mandibles, a line on the
inner orbits, above extending to near the hind ocelli, the
lower half of the outer orbits, malar space, prothorax (except
for a line in the centre of the pleure dilated at the base), a
large mark in the centre of the mesonotum (trilobate at the
base, roundly incised at the apex), scutellums, a broad mark
down the middle of the metanotum (half in the upper, half in
the lower part, of equal width) rounded above, transverse
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 6
82 Mr. P. Cameron on Hymenoptera
below, tubercles, a large mark on the lower half of the meso-
pleure (rounded below and extending on to the sternum,
rounded and narrowed above at the apex), an oval spot below
the hind wings, a large mark in the middle of the metapleurae
(transverse at the base, the three other sides broadly rounded),
the postpetiole, almost the apical half of the second segment,
two broad marks (obliquely narrowed towards the base) on
the apical half of the third, two smaller oblique triangular
marks on the sides of the fourth, a narrow line on the apex of
the fifth and sixth, and the seventh from shortly behind the
middle, bright orange-yellow, as are also the legs (except the
fore femora behind, the middle at the base all round and at
the apex, slightly more broadly above, the middle yellow part
being about as long as the black apical) ; the basal and apical
third of the posterior all round, the base narrowly of the four
hind tibie (their apices more broadly), the four anterior tarsi
above, and the hind coxe and trochanters, black. ‘The
underside of the antennal scape yellow, the sixth to fourteenth
joints of the flagellum white. Wings hyaline, the nervures
and stigma black. ¢.
Length 14 mm.
Runjit Valley, Sikkim, 1000 feet; April (C. 7. Bingham).
Head closely punctured, the apex of clypeus and a large
round depression over each antenna smooth, shining. ‘Thorax
closely punctured, the depression in the centre of propleurse
obliquely irregularly striated; the apex of the mesopleure
finely, closely, obliquely striated below. Median segment
closely rugosely punctured, the apex of the posterior median
area transversely striated; the lateral apical areze more
strongly, irregularly, and much less closely striated; the
apex ot the pleure irregularly, stoutly, obliquely reticulated.
Postpetiole smooth, the second and third segments closely
punctured. Gastrocceli longer than wide, somewhat pyri-
form, smooth on the inner side at the base, the outer side
coarsely aciculated. Tarsi closely spinose. ‘he areolet
5-angled, large, the recurrent nervure received shortly beyond
the middle.
For a synopsis of the Indian species of Haliphera see ‘The
Entomologist,’ 1904, p. 307. Inter alta, the present species
may be known by the broad continuous band on the second
abdominal segment.
DIPLOPTERA.
Odynerus Hewittit, sp. n.
Black ; the mandibles (except at extreme apex), clypeus,
from the Oriental Zoological Region. 83
the eye-incision entirely, a line down the centre of the front
(narrowed on the upper half, dilated below the middle, and
the apex still more widely triangularly dilated), the outer
orbits almost entirely, a triangular mark between the ocelli
and the eyes, a large pyriform mark on the sides of the pro-
notum, the two united by a narrow line, two longish lines
down the centre of the mesonotum, a mark on the basal half
of the scutellum (roundly incised in the middle), a large
mark on postscutellum (its base transverse, the apex obliquely
narrowed), a large curved triangular mark covering the sides
of the metanotum, the narrowed part below, two large marks
on the mesopleure at the base (the upper broader and shorter
than the lower, the two divided by the suture), tegule (except
for a fuscous spot), the first abdominal segment at the top of
apical slope, its apex and the apices of the following seven
segments, the lines dilated laterally and that on the seventh
also in the middle, a mark in the centre of the eighth (nar-
rowed towards the apex), the sides of the second ventral
broadly, its apex more narrowly (the black central mark
formed thereby is obliquely narrowed at the apex), and the
apices of the other segments, yellow. Legs yellow, the coxe,
trochanters, and femora black behind. Wings hyaline, the
anteriorsuffused with fuscous; the stigma fuscous, the nervures
black. .
Length 10 mm.
Kuching, Borneo; June (Juhn Hewitt).
Clypeus as broad as long, the apical half roundly narrowed,
the apex transverse. Upper part of head and the thorax,
except the apex of the mesopleure and the base of meta-
pleure (which are bare, smooth, and shining), closely, rather
strongly punctured. Underside of antennal scape yellow, of
the flagellum brown, as is also the hook, which is as long as
the joint. Base of thorax not quite transverse, the sides of
metanotum margined ; the postscutellum is obliquely nar-
rowed. ‘The pubescence all over is short, dense, and pale.
In the Journ. Linn. Soc. 1857, p. 112, Mr. F. Smith
described an Odynerus multipictus from Borneo; in the same
journal, 1859, p. 165, he described another O. mulupictus
trom Aru. In his ‘Catalogue of Malay Hymenoptera’ tlhe
Aru multipictus is duly noted, but not the earlier one.
Saussure (Stett. ent. Zeit. xxui. p. 200) redescribes what he
regarded as the Borneo multipictus, and renamed it guttu-
latus—“ parce que l’auteur |’a aussi donné a une autre espéce
qui pourra le conserver.” It is, however, the Aru mu/tipictus
of 1859 that ought to have been renamed. Bingham (‘ Fauna
of Brit. India,’ Hymen. 1. p. 868) describes and figures the
Ge
84 Mr. P. Cameron on [Tymenoptera
Borneo multipictus, but only the female. Col. Bingham has
given me from Sikkim the male of what is no doubt the
Indian multipictus (1. c. pl. ii. fig. 13). I have unfortunately
only males of these yellow-banded species. Those known
to me may be separated as follows :—
a, The pubescence dense and black, the apex of
the clypeus not transverse; pygidium black ;
the lateral marks on metanotum not much
widened above.
Pronotum and tegule black ; pleuree and scutel-
lum immaculate , second ventral segment
black, with two large yellow spots, the
othersemments black sce, snisice ae ieee « multipictus, Sm.
Pronotum, pleuree, and scutellums marked with
yellow ; the ventral segments (except the
apical two) for the greater part yellow.... septemfasciatus, Sm.
6. The pubescence short and pale; the apex of
clypeus transverse; pygidium with a large
yellow mark; the marks on metanotum much
WIECH ADOVE Moms eel aiseretetoinic! vetlel eee melaiets Hewitt, sp. n.
Odynerus heterospilus, sp. n.
Black; the clypeus, the eye-incisions, a longish triangular
mark over the antenne, the greater part of the mandibles, an
oblique mark opposite the ocelli touching the eyes, the outer
orbits broadly, the base of pronotum, apex of tegule, the base
of scutellum to near the middle, a broad curved line (widened
above) on the sides of metanotum, a large conical mark
(longer than wide) below the tegula, a smaller narrower spot
near the apex of mesopleure below, the first abdominal
segment (except narrowly at the base), the apices of the
second to fifth, the whole of the sixth, the base of the second
ventral, and the apices of the middle ventral narrowly, yellow,
the yellow on the abdomen tinged with rufous ; a transverse
line on the vertex behind the ocelli, the apex of pronotum
from shortly behind the middle, the sides of mesonotum
largely, and a spot below the hind wings surrounding a
yellow mark, blood-red. Underside of antennal scape yellow,
of flagellum brown. Legs yellow, the four anterior coxe,
trochanters and femora behind, and the greater part of the
posterior, black. Wings clear hyaline, the radial cellule
smoky, the stigma testaceous, the nervures black; the first
and second transverse cubital nervures united in front, 9.
Total length 9 mm.
Kuching, Borneo; February (Hewitt).
Belongs to Odynerus (sensu str.). Densely covered with
short white pubescence. First abdominal segment large,
from the Ortental Zoological Legion. 89
cup-shaped, not quite sessile, slightly longer than it is wide
at the apex; the second longer than wide, the apex with a
narrow but distinct depression. Clypeus with the apex trans-
verse, as it is also above. Apex of postscutellum transverse ;
the sides of metanotum bluntly rounded, the apex transverse,
not much depressed in the middle. The base of the thorax
is not quite transverse and with the sides rounded.
A distinct species.
Ancistrocerus megaspilus, sp. 1.
Black ; clypeus, mandibles (except the inner edge), the
eye-incision, the yellow extending beyond it, a longitudinal
line (extending from shortly above the top of the eye-incision
to the antenne, its top narrowed, the bottom dilated), a short
oblique line on the vertex running into the top of the eyes,
the outer orbits broadly, a broad line on the sides of pronotum
(extending from the base to shortly beyond the middle),
tegulz, basal two thirds of scutellum, the apex of the marx
transverse, the sides of the metanotum broadly, the mark
curved and narrowed below, a large almost semicircular mark
below the tegule, a much smaller oblique conical mark on
the apex of mesopleure below, a small transverse mark on
the sides of the first abdominal segment at the top of basal
slope, a line on the apex of the first to fifth segments (that on
the first dilated in the middle, the dilated part with an
incision), and a large conical mark on the sides of the second
segment, yellow. Legs yellow, black behind at the base.
Wings hyaline, the radial and the apex of the costal cellule
fuscous violaceous, the stigma dark fuscous; the recurrent
nervure interstitial, the second cubital cellule much narrowed
in front, but with the nervures apart. ¢.
Total length 10 mm.
Kuching; March (Jewitt).
Apex otf clypeus transverse, moderately wide. Thorax
more than twice longer than wide, the sides at the base
broadly rounded, the apex transverse. Apex of postscutellum
obliquely narrowed towards the centre. First abdominal
segment cup-shaped, not quite sessile at the base; at the
centre of its apex is a distinct fovea, there being similar fovese
on the second and third; the segments are smooth, silky.
The base of the first abdominal segment is hardly separated
from the rest, but is more glabrous and shining.
Sphegide.
Cerceris baluchistanensis, sp. n.
Yellow; asmal! longish spot over each antenna, the vertex
3 g i
86 Mr. P. Cameron on [Tymenoptera
from the hinder ocelli (except for a triangular spot behind
each of the latter), the basal two thirds of the pronotum and
the central apical furrow, three broad lines on mesonotum
and a similar one across the base, the space bordering the
sides of scutellums, a large triangular mark covering the
basal area of metanotum, a broad line covering the central
apical furrow, and the space between the meso- and metapleure,
black ; the flagellum of antennze and the space separating the
abdominal segments rufo-fulvous. Legs yellow, the femora
tinged with fulvous. Wings hyaline, the apex narrowly
smoky, the costa and stigma fulvous, the apical nervures
black. ¢.
Length 12 mm.
Quetta; June (Nurse).
Metanotal area closely but not very strongly transversely
striated. Head, thorax, and, to a less extent, the abdomen
covered with white pubescence, closely strongly punctured,
the metathorax more closely than the rest. Clypeus fringed
with pale golden hair, its apex bluntly rounded. Hyes
slightly converging above. Hinder ocelli separated from each
other by a little greater distance than they are from the eyes.
Apical half of penultimate joint of antenne abruptly narrowed,
the last bluntly rounded at the apex. First abdominal seg-
ment slightly narrowed at the base, longer than it is wide at
the apex. Pygidium strongly punctured throughout, of equal
width, the apex bluntly rounded, keeled; the apex of
epipygium bluntly rounded.
Comes near to C. Waltli’, Spin., from Egypt.
Cerceris quettaensis, sp. n.
Black ; the face, a broad line on the lower half of the inner
eye-orbits, clypeus, mandibles (except the teeth), antennal
scape, two marks (broader than long) obliquely widened
towards the apex on the outer edge of pronotum, tegule,
tubercles, postscutellum, basal half of second abdominal
seoement, the third entirely, the fourth narrowly in the centre,
widely at the sides, the fifth (except for a curved black band
on the base), the sixth entirely, and the greater part of the
ventral segments, bright lemon-yellow, as are also the legs.
Basal four joints of antennal flagellum dark yellow, the apical
brownish below. Wings hyaline, the apex narrowly smoky,
the stigma dark fuscous, the nervures black. @.
Length 11 mm.
Quetta (Nurse).
Metanotal area smooth, shining, the central furrow wide,
from the Oriental Zovlogical Region. 87
opaque, finely rugose. Entire body closely strongly punc-
tured. ‘Third antennal joint as long as the scape, shorter
than the second and fourth united. First abdominal segment
longish oval, about one half longer than wide, as long as the
second. Abdominal segments strongly but not very closely
punctured, the second smooth in the centre at the apex, the
others smooth at the base, the last strongly punctured, de-
pressed at the apex. Pygidium closely strongly punctured,
slightly gradually narrowed towards the apex.
Comes near, apparently, to C. ewgenta, Schlett.
Cerceris basimacula, sp. n.
Black ; a mark on the centre of the clypeus (obliquely
narrowed above and below), a broad line on its apex, a broad
line on the inner orbits from the antenne, two large oblique
marks on the apex of the metanotum (extending on to the
pleuree), the basal third of the abdominal petiole on the sides,
more than the basal third of the second segment, and narrow
lines on the apices of the fourth and fifth segments, yellow ;
the sides of the central and lower parts of the clypeus, pro-
thorax, seutellums, more than the apical third of the second
and third abdominal segments, a line behind the yellow on
the fourth and fifth, and the apical two segments entirely,
dark rufous ; the apex of the first segment of a paler rufous
colour. our front legs pale rufo-testaceous ; the tibiz and
base of tarsi yellow ; the hind legs similarly but much darker
coloured. Wings hyaline, the radial cellule smoky, the apical
cubital cellule of a deeper smoky colour ; the costa and stigma
dark testaceous, the nervures black. Antenne rufo-testa-
ceous, the scape yellow below, the eight apical joints blackish,
paler below. 2.
Length 10 mm.
Sikkim.
Face strongly but not closely punctured ; the clypeus more
finely and closely punctured; there is a short keel in the
centre of the latter, with a fovea on either side. Front and
vertex strongly, somewhat closely punctured. Middle of
pronotum closely punctured, the sides almost smooth ; meso-
notum somewhat strongly but not closely punctured; the
scutellum is more strongly but not so closely punctured as the
latter. Postsecutellum smooth. Metanotum strongly punc-
tured, the punctures clearly separated; the area shining,
smooth. Head and thorax densely covered with white
pubescence. First abdominal segment of equal width, about
four times as long as wide, longer than the second; the
88 Mr. P. Cameron on Hymenoptera
segments strongly punctured ; the pygidium strongly but not
very closely punctured; the sides rounded, narrowed at the
base and apex, the latter slightly rounded inwardly. The
whole abdomen covered with longish white pubescence.
Hypopygium depressed at the apex; the latter has a slight
rounded incision.
Allied to C. lencha, Cam.
Crabro violaceipennis, sp. n.
Black; the antennal scape, a narrow transverse line on the
postscutellum, a spot (broad and rounded at the base, roundly
narrowed towards the apex) on the base of the metanotum in
the centre, a band (rounded at the apex, the base with two
incisions in the middle, the space between projecting beyond
them) on the apex of the first abdominal segment, a broad
irregular band on the basal half of the second, a slightly
narrower band (almost continuous) on the base of the third,
and similar bands (but more clearly interrupted) on the base
of the fourth and fitth, bright lemon-yellow. ‘The legs black ;
the fore tibize in front and above, the middle and posterior
(except below and narrowly at the apex), the middle femora
broadly below, and the basal joint of the tarsi (except at the
apex), lemon-yellow. Wings fuscous violaceous, the nervures
and stigma black. ¢.
Length 13 mm.
Sikkim.
Mesonotum and scutellum closely punctured, the latter
more closely at the base than at the apex, where the punctures
run into striz. Postscutellum smooth at the base, the apex
strongly but not closely punctured. Metanotal area closely
longitudinally striated, the rest of the metanotum more
strongly obliquely striated, the striae curved and interlacing
more or less. Face, clypeus, and cheeks densely covered
with silvery pubescence; the thorax densely covered with
white to silvery pubescence, which is longest on the meta-
notum. Pygidium shining, longish, the basal third with
deep clearly separated punctures; the rest is less strongly
and more sparsely punctured, especially in the middle.
‘There is a deep crenulated furrow down the base of the meso-
pleure. Abdomen sessile, triangular.
Belongs to Bingham’s section B. c. 6'; with none of the
species therein can it be confounded.
Crabro clenopus, sp. Nn.
Black ; a line above on basal half of mandib es, two broad
from the Oriental Zoological Region. 89
lines on pronotum, a large, irregular, oblique mark on the
sides of second abdominal segment, a line (about three times
longer than wide) on the sides of the third, a slightly longer
and narrower one on the sides of the fourth, with two small
lines between, a line on the basal half of the fitth, with two
black lines in the centre at the base, a band on the centre of
the sixth (with two rounded black lines, united at the base,
in the centre), bright lernon-yellow. Anterior legs testaceous,
the femora with the lower half yellow in front ; the tarsi
blackish towards the apex ; the middle trochanters and femora
clear lemon-yellow, with a broad black line in the middle and
a short line on the apex below; the tibia lemon-yellow,
blackish at the base and apex, and with a broader blackish
band in the centre; hind legs black, the metatarsus to near
the middle pale yellow. Wings hyaline, iridescent, the costa
and stigma black. ¢.
Length 9 mm.
Sikkim.
Abdominal petiole fully one fourth longer than it is wide
at the apex ; it becomes gradually widened from the base to
the apex and is closely punctured. Base of metanotum with
a small central area of equal width and a large area on either
side (wider than long), the apical keel less distinct than the
lateral and rounded ; the top of the apical slope is irregularly
widely reticulated, the middle smooth, the apex transversely
striated. Top and apex of mesopleure above and at the apex
smooth, the latter bounded by a rounded curved keel; the
apex below with a narrow band of stout striw. Metapleura:
(except at the apex above) striated closely, ‘The pubescence
on the head, thorax, and sides of abdomen white, on the back
of the abdomen bright golden fulvous. Basal joint of middle
tarsi dilated, straight on inner side, rounded on outer to near
the apex, which is narrowed, below it is fringed with a
comb of stout bristles, the apex projects and ends in two
long spines ; the basal joint of the anterior broad, curved,
slightly narrowed at the base, the following three joints are
broad, the second as long as the following two united ; the
first joint of the posterior longer than the following united,
broad, narrowed at the base. ‘he middle of the mandibles
rufous.
In having the metanotum areolated this species agrees with
C. agycus, Vam., from the Himalayas (‘The Entomologist,’
1904, p. 261).
Psen montanus, sp. 0.
Black, shining, sparsely covered with short white pubes-
90 Mr. P. Cameron on Hymenoptera
cence. Wings clear hyaline, the nervures and stigma black ;
the second cubital cellule much narrowed in front, one fourth
of the length of the third in front; the first recurrent nervure
received very shortly beyond the transverse cubital, the
second at the apex of the basal fourth; metanotum without
keels, except two indistinet longitudinal ones leading down to
the furrow, which is distinct, smooth on upper, crenulated on
lower half; the narrowed part of petiole half the length of
thorax. Body smooth ; the pubescence densest and longest
on the sides. 9.
Length 7 mm.
Simla (Nurse).
May be known from the other black Indian species by the
absence of keels and arew on the metanotum, it being also
smaller than them. P. kashmirensis, Nurse, has a “ remark-
able tubercle” between the antenna, and it agrees otherwise
with the present species closely, but it differs in having a
triangular depression at the base of the metanotum, which
also bears outwardly diverging striz, this not being the case
with the present species.
Psen rufoannulatus, sp. n.
Black, shining, densely covered with longish white pubes-
cence ; the second and third abdominal segments bright red ;
the apical four joints of the tarsi rufo-testaceous ; the wings
clear hyaline, the stigma and nervures black; abdominal
petiole as long as the thorax ; the front with a distinct keel
down the middle. @.
Length 9 mm.
Simla; August (Nurse).
A. stout species.
Metanotal area bare, shining, large, clearly defined, the
apex gradually roundly narrowed to a point shortly above
the middle of the metanotum ; in its centre are two parallel
keels; at a distance from these is a keel; separated from it
at the outer edge are three or four less distinct ones. The
rest of the metanotum is opaque, irregularly striated, the
strize more or less curved; the pubescence is long and dense.
The first recurrent nervure is received near the apex of the
basal third of the cellule, the second very shortly beyond
the second transverse cubital. Hind tibie with about a
dozen white spines. The second abscissa of radius not quite
half the length of third ; the second cubital cellule wide in
front.
from the Oriental Zoological Region. 91
Psen puncticeps, sp. n.
Black ; the knees, tibie, and tarsi whitish testaceous, the
hinder darker tinted, with the tibie fuscous from shortly
beyond the middle ; the second and following segments of the
abdemen piceous, the narrowed part of petiole half the length
of the thorax. Wings clear hyaline, the stigma dark fuscous,
the nervures black. Metanotum without a basa] area, the
sides keeled, the keels with a narrow striated band on the
inner side ; the furrow on the apical slope is wider and deeper
on the basal than on the apical part. Antenne stout,
brownish below, the third joint about one fourth longer than
the next, which is as long as the second. First recurrent
nervure received close to the base of the cellule, the second at
double the distance. Front closely, distinctly, the vertex
less strongly punctured ; there is a narrow keel down the
centre of front, ending in a large stout projection between
the antennze. Parapsidal furrows distinct on basal half of
mesonotum ; the furrow on the base of metanotum is deep,
crenulated, it is separated in the middle by a smooth space
bounded by curved keels. 9.
Length 7 mm.
Simla.
Palarus fortistriolatus, sp. n.
Black, shining, sparsely covered with white pubescence ;
the three lobes of clypeus, lower half of front, basal third of
mandibles, a broad line on the lower three fourths of the outer
eye-orbits, the apex and the sides of pronotum to near the
middle, a line on its base, a large mark in front of the
tegule roundly, irregularly narrowed at the base, tegule,
tubercles, a line on the base of mesopleure, gradually narrowed
below to a sharp point, not reaching to the breast, an irre-
gular triangular spot on the outer lower side of its apex,
scutellums, their lateral keels, and the greater part of the
abdomen, bright lemon-yellow. Legs yellow, suffused with
rufo-fulvous, the four hinder cox and the greater part of
trochanters black; the anterior metatarsus with seven long
spines of equal length (except the basal, which is shorter),
the apical two close together. Wings hyaline, the apex very
slightly suffused with fuscous, the stigma and nervures
fulvous, the apical nervures darker coloured. @?.
Length 13 mm.
Quetta ; June.
The basal slope of the first abdominal segment, the base
92 Hymenoptera from the Oriental Zoological Region.
of the second to sixth narrowly, and the ventral (except for a
line on the apex of the second and irregular marks on the sides
of the others) black; the depressed apices of the first to
fourth brownish ; the line on the second broad, slightly dilated
in the middle, that on the third forming a broad triangle, that
on the fourth narrowly dilated backwards to the apex of
basal third in the middle; there is a deep furrow in the
centre of the apical two thirds of the fifth. The second
ventral segment becomes gradually raised in the centre
towards the apex (forming about one third of the segment),
the apex of the raised part slightly roundly narrowed; the
part beyond this is raised also, is stoutly striated, and has the
apex bluntly rounded and curled up. Pygidium long,
gradually roundly narrowed from the base to the apex,
sparsely weakly punctured. Eyes at the top separated by
not quite the length of the third antennal joint. Hinder
ocelli irregularly oval, depressed in the centre, the depression
behind them longer than wide, the apex deep, the base much
shallower and narrowed; the anterior ocellus roundish, sur-
rounded by a rounded furrow. ‘Third antennal joint as long
as the second and fourth united; front and face densely
covered with silvery pubescence. Pronotum with a steep
oblique slope from the top to the bottom, the top rounded,
clearly separated, widened laterally. Base of mesonotum
closely, the sides and apex sparsely punctured, the scutellums
still more sparsely punctured. Metanotum depressed in the
middle, strongly striated, transversely on base, obliquely on
sides of apex, the centre being smooth; the rest closely,
strongly, rugosely punctured, with an irregularly reticulated
area, widest on outer side, on the inner side at the base.
Pro- and mesopleure shining, sparsely punctured; the meta-
pleure closely irregularly reticulated. The last ventral
segment is flat at the base, gradually narrowed to a fine point,
then prolonged into a keel; the penultimate flat. The front
over the antenne is roundly raised. Middle of first abdo-
minal segment keeled laterally. ‘The striation on the meta-
notum is iregular, being more transverse in one example
than in the other; it is strong in both.
Allied to P. flavipes.
On a new Leptolepid Fish from the Weald Clay. 93
XVIT.—On a new Leptolepid Fish from the Weald Clay of
Southwater, Sussex. By A. SMira Woopwarp, LL.D.,
ERS:
[Plate I.]
TuIn cycloid scales which might have belonged to a Leptolepis-
like fish have already been neticed in the English Wealden
formation, but no complete example of a Wealden member
of the Leptolepide has hitherto been described. A well-
preserved specimen, however, which may be referred to the
family just mentioned, has lately been found in the Weald
Clay at Southwater, near Horsham; and I am indebted to
the Directors of the Southwater Brick and Tile Co., Ltd.
(through Mr. C, H. Aldersmith, A.M.I.C.E.), for the oppor-
tunity of studying this interesting fossil, which has now been
presented to the British Museum.
The new specimen is preserved in counterpart in a slab of
clay, and its best half is shown of one half nat. size in
Pl. I. fig. 1. It is exhibited in direct side-view, only
lacking the anterior part of the head and the hinder half of
the caudal fin. The maximum depth of the trunk is con-
tained somewhat less than three times in the length from the
pectoral arch to the base of the caudal fin, and would probably
equal about one fifth of the total length of the fish. The
same depth is not quite three times as great as the depth of
the caudal pedicle. The fragmentary remains of the head
show that the mandibular suspensorium is inclined forwards,
so that the articulation of the lower jaw must have been
directly beneath the hinder part of the orbit. The hyo-
mandibular bone (dm.) bears a long process for the suspension
of the operculum (op.), which is shown in impression, trape-
zoidal in shape, and about as deep as broad. The preoper-
culum (pop.) has a long upright ascending limb, expanding
below into a triangular plate. The suboperculum (sop.), best
seen in the counterpart not figured, must have been about
four times as broad as deep. Fifteen branchiostegal rays (dr.)
can be counted, the upper seven being expanded and in close
series, the lower eight being narrower bars and more widely
spaced. ‘The opercular apparatus is quite smooth, not
ornamented. The total number of vertebra is about sixty,
half being in the abdominal region. ‘The centra are about
as long as deep in the anterior part of the caudal region, but
are somewhat shorter than deep both in the abdominal and
94 Onanew Leptolepid Fish from the Weald Clay.
in the hinder part of the caudal region. They are well ossified
and their primitive double-cone is strengthened by secondary
bone arranged in fine, close, longitudinal ridges (fig. 2).
The ribs are stout, apparently borne on very short transverse
processes, and clearly extending to the ventral border of the
fish. ‘The fixed neural and hemal arches in the caudal
region are also very stout and gently arched. The hinder
extremity of the vertebral column turns only slightly upwards,
and its heemal arches are somewhat expanded without fusion
into plates. ‘The intermuscular bones are almost completely
obscured by the scales in the fossil, but there are traces of
them above the vertebral column in the abdominal region,
and both above and below this column in the caudal region.
The post-temporal (pt.) is a thick plate, almost triangular i in
shape, and the supraclavicle (sc/.) is a deep and narrow bone.
The clavicle (c/.), as shown in impression, is expanded into a
large smooth plate above the pectoral fin, which is inserted
close to the ventral border. When adpressed to the trunk
this fin extends halfway to the insertion of the pelvic fins;
its rays have a very long and stiff base, but are finely divided
and articulated at the extremity. The pelvic fins (plv.) are
smaller than the pectorals, though similar in character, and
inserted midway between the pectorals and the anal. The
dorsal fin (d.) arises well in front of the middle point between
the occiput and the caudal fin, somewhat in advance of the
insertion of the pelvic fins. It comprises eighteen to twenty
rays, of which the three foremost are closely pressed together,
undivided, and gradually increase in length. The length ot
the fourth or longest ray much exceeds half the depth of the
trunk at its insertion, and, like the following rays, is finely
divided and articulated distally. The anal fin (a.) resembles
the dorsal in character, but is much smaller and comprises
only thirteen or fourteen rays. It is far behind the dorsal,
and its origin is much nearer to that of the caudal than to
the insertion of the pelvic fins. he remains of the caudal
fin-rays (¢.) show that they were comparatively stout. There
are no fulcra on any of the fins, The scales are relatively
large, cycloid, and smooth, occasionally with feeble traces
of a slight radiating pectination at the hinder border, but
usually exhibiting structural lines, including wavy concentric
markings. They are scarcely displaced in ‘the fossil, and are
seen to be deeply overlapping, with the exposed area narrow
and deep. The “lateral line” is scarcely traceable, but
seems to produce a slight depression along some scales in a
series above the vertebral column.
So far as the skeleton is preserved there is nothing in the
On new Species of Histeride. 95
Wealden fossil just described to prevent its reference to an
Hlopine or Clupeoid fish; but as the European Wealden
fish-fauna is essentially of a Jurassic type *, the specimen is
more likely to belong to a member of the Leptolepide, with
which it equally agrees. ‘The skeleton of the trunk resembles
that both of Leptolepis itself and of Athalion ; but the lack
of jaws prevents an exact determination of the genus. The
number of the vertebree exceeds that of all known species of
both genera except dthalion Vidali +, and the relative short-
ness of the hinder caudal, as well as the anterior abdominal
vertebral centra, is a feature peculiar to the new Wealden
fish. The fins, as described, also distinguish this fossil from
all species with which it can be compared. It therefore
represents a new species, which I propose to name Leptolepis
valdensis until the discovery of the head determines its
precise generic position.
EXPLANATION OF PLATE LI.
Fig. 1. Leptolepis valdensis, sp. n.; right side view of type specimen, one
half nat. size-—Weald Clay; Southwater, Sussex. [{ Brit. Mus.
no. P. 10440.] a, anal fin; 6r., branchiostegal rays; ¢., caudal
fin; c/., clavicle ; d., dorsal fin ; Am., hyomandibular ; op., oper-
culum ; plv., pelvic fins; pop., preoperculum ; pt., post-temporal ;
sel., supraclayicle ; sop., suboperculum.
Fig. 2. Ditto; caudal vertebrae of same specimen, nat. size.
XVIII.—On new Species of Histeride and Notices of others.
By G. Lewis, F.L.S.
Tus is the thirty-first paper of this series, which dates from
the year 1884. In the Histeride the absence or otherwise of
prosternal striz is sometimes of great significance, and serves
to distinguish both genera and species. Thus, in Teretrio-
soma the striz are wanting, in Teretrius they are well marked
and very useful as specific characters; and in Paromalus, as
the genus is now defined, the prosternal keel is marginate—
that is, the striae meet at both ends. In Grammostethus,
also, the prosternal striew are of importance, as being con-
stantly similar in a series of fourteen cognate species, but
which possess, however, inter alia, good specific characters.
* A.S. Woodward, “ Note on the Affinities of the English Wealden
Fish Fauna,” Geol. Mag. [4] vol. iii. (1896) pp. 69-71.
t H. E. Sauvage, “ Noticia sobre los Peces de la Caliza litografica de
la Provincia de Lérida,” Mem. R. Acad. Cienc, Barcelona, [3] vol. iv.
no. 35 (1903), p. 18, pl. ii. fig. 2. a
96 Mr. G. Lewis on
But it is necessary to note that in other genera the strie
only help to identify certain species. In Plescus and Placodes
they can only be used as specific characters. Plestus cossy-
phus, Mars., and striatipectus, Lew., are extremely similar,
but the latter alone has a bistriate prosternum, and the same
differences clearly separate edlipticus, Mars., from javanus,
Er. In the genus Placodes, caffer, Hr., has strie, and they
are wanting in ebeninus, Lew., species which otherwise are
superficially very much alike.
It is to be regretted that at the present time there are
genera which are not well defined, notably Lioderma, Apo-
bletes, Phelister, and others, but, as time goes on, with in-
creasing knowledge, some future student of the group may
see a way to point out some salient characters not at present
obvious, and, perhaps, by further subdivision introduce a
more intelligent systematic arrangement,
Marseul’s monograph will, however, remain the initial
work on the family, and the figures of the species, drawn by
him at the time of writing his descriptions, are so charac-
teristic that they lead generally to the certain identification
of the species. But in his time more material was wanted
for an efficient classification, and not much advance has been
made since for the same reason.
List of Genera and Species.
Lioderma pervalidum, Blaisdell. Hister similis.
Hyposolenus, gen. nov. Microlister Sheppardi.
Apobletes poenalis. Margarinotus, Marseul.
Carcinops Voulogeri, Théry.
pumicatus.
Platylister niponensis, Lev. Monoplius Braunsi.
Platysoma germanum. Probolosternus brevistrius.
Cornillus, gen. nov. Exosternus rufulus.
Contipus, Marseul. Reninus Salvini, Lew.
Eugrammicus, gen. nov. puncticollis.
Zabromorphus ordinarius. Eretmotus planifrons.
Coptochilus, Rey. foveisternus.
Heterognathus, Rey. Onthophilus niponensis.
Hister Marseuli.
Lioderma pervalidum, Blaisdell, Lew. Ann. & Mag. Nat.
Hist. xiv. p. 139 (1904).
L. 12-16 mill. (absque mandibulis).
A series of this Californian species has been sent to me,
and I understand that it will be shortly described in America.
It is very distinct from yucatecum, Mars., by its more
elongate and parallel outline, by being slightly more convex,
by the second dorsal stria being complete, by the elytra being
new Species of Histeridex. 97
sparingly punctured on the apical margin, by the propygidium
being wholly punctured on the disk and not at the sides only,
and by the third and fourth abdominal segments being trans-
versely punctured. JZ. yucatecum has sometimes, but rarely,
the second dorsal stria complete, but this variety was not
known to Marseul.
HYPOSOLENUS, gen. nov.
Bedy oval or oblong-oval, somewhat convex ; head retrac-
tile; mandibles dentate; labrum narrow and transverse ;
forehead striate; antenna, scape long and bent; thorax
transverse, lateral stria strong and abruptly terminating after
passing the anterior angle; prosternum bistriate ; meso-
sternum with a deep anterior marginal furrow; pygidium
convex and closely punctate. The other characters agree
with those of Plescus, in which genus the species levigatus,
Mars. (type), and levis, bengalensis, and hamatus, Lew., have
been hitherto included.
Apobletes peenalis, sp. n.
Ovalis, deplanatus, niger, nitidus; fronte excavata, punctulata ;
clypeo transverso, margine late sinuato; pronoto lateribus sub-
rugose punctato, stria marginali antice interrupta; elytris striis
dorsalibus 1-3 integris, 3 sinuata, 4 apicali, ceteris nullis ; pro-
pygidio punctato in medio excepto ; pygidio toto profunde punc-
tato ; mesosterno stria marginali haud interrupta.
L, 34 mill.
The above is similar to A. cavifrons, Lew., in that the
mesosternal stria is complete. A. excavatus and feriatus,
Lew., Schaum? and tener, Mars., belong to the same group,
and the six species all closely resemble one another, but in
the last four the mesosternal stria is interrupted. The type
example of A. excavatus has the third dorsal stria interrupted,
but in a longer series the stria is complete and in several the
pygidium is deeply foveolate on each side at the base.
Hab. Andaman Islands.
Apobletes pumicatus, sp. n.
Oblongo-ovatus, deplanatus, niger, nitidus, antennis pedibusque
piceis; fronte plana, haud striata, impunctata; pronoto stria
marginali tenuiter impressa, brevi in angulo antico ; elytris striis
1-3 integris, 4 apicali, dimidiata; propygidio disperse punctu-
lato; pygidio basi transversim punctato; prosterno lato haud
striato; mesosterno bisinuato, immarginato ; tibiis anticis multi-
denticulatis.
L. 2% mill.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 7
98 Mr. G. Lewis on
Very similar to A. Marseuli, Lew., and pauperatus, Sch.,
which together form a local section of the genus in which
the head is smooth and without striae, the thorax has a very
fine marginal stria close to the edge and confined to the region
of the anterior angles, the pygidia are indistinctly punctured,
and the prosternum broad and not striate. Specifically
pumicatus may be known by its more oblong form and by the
third dorsal stria being complete.
Hab. Cape S. Juan, Biafra, West Africa, In the Museum
of Madrid and my own collection.
Platylister niponensis, Lew.
There is an example of this species in the Museum of
Paris from the island of Kiushiu, and I think it likely that
this is the true locality of the species, not Kioto on the main
island, as I have recorded it. ‘The last place is very far north
for the occurrence in it of a genus which is essentially tropical
or subtropical.
Platysoma germanum, sp. n.
Oblongum, subparallelum, nigrum, nitidum; fronte stria recta,
integra ; pronoto stria marginali antice interrupta; elytris striis
1-3 integris, 4-5 dimidiatis, suturali brevissima ; mesosternum
late emarginatum, stria integra, conspicue impressa; tibiis
anticis 4-dentatis.
L. 33 mill.
Oblong, rather parallel, slightly convex, black and shining ;
the head, frontal stria straight anteriorly and complete, surface
obscurely punctulate; the thorax punctured laterally, stria
continued behind the neck and laterally it is close to edge
(differing in this respect from Alewandri, Mars.) ; the elytra,
striae 1-3 complete, 4—5 dimidiate, sutural very short and
occupying a median position to the fifth stria, the humeral
strice are wanting; the propygidium is coarsely punctured,
but the punctures are not closely set; the pygidium, the
punctures are similar but closer together; the prosternum,
keel rather narrow, without strie; the mesosternum widely
emarginate, with a well-marked marginal stria; the femora
and tibiee are red.
Smaller but somewhat similar to P. Alexandri, Mars. ; the
thoracic stria, however, is close to the edge of the thorax
laterally and is interrupted behind the neck ; the thorax is
punctured at the sides and there is no vestige of a sutural
stria in Marseul’s species. Marseul says of P. Alexandri
new Spectes of Histeride. 99
that there is a short oblique humeral stria, but this oblique
stria, very common throughout this family, is not usually
called a ‘ humeral” stria—humeral striz, as usually under-
stood, are longitudinal.
Hab. Mikindani, E. Africa (Raffray).
CoRNILLUS, gen. nov.
Body oval or oblong-oval; forehead impressed longitu-
dinally, stria biarcuate ; thorax, antennal fossa in the anterior
angle; elytra, strie fine, crenate, or punctiform ; pro-
pygidium finely and evenly punctured, binodulose; pro-
sternum bistriate ; anterior tibize 4-dentate.
This genus has much general similarity to Omalodes ; the
species to be included in it are tuberculipygus, Sch., bullatus,
tuberculatus, and tuberosus, Lew.
Contreus, Marseul, Mon. p. 545 (1853).
I propose to include in this genus C. flezuosus, Sch., as the
type, because it is the commonest species existing in collec-
tions, and digitatus, Mars., with ¢mmarginatus and oblongus,
Lew. For the present I would also include the American
species, abhorrens, Sch., platanus and subquadratus, Mars.
The African species with didymous elytral stria I assign to
the new genus below.
EUGRAMMICUS, gen. nov.
Body oval, more or less convex ; head transverse, man-
dibles equal and strongly dentate, labrum short and transverse,
frontal strie complete; thorax with two lateral strie; an-
tennal fossee are open from below and shallow and somewhat
similar to those of Contipus; elytra, strie are wide and
shallow, with fine crenate edges (Marseul termed them didy-
mous), striz 1-4 complete, 5 and sutural nearly complete,
sutural turning outward before the base; pygidia almost
impunctate; mesosternum feebly sinuous anteriorly ; tibizx
dilated, anterior strongly 3-dentate. Other characters similar
to those of Contipus.
Type, Contipus didymostrius, Mars.
The other known species are sinuosus (convex), prowimus,
and fractistrius, Lew. (less convex) ; at present only unique
examples are in collections, so that it is highly probable
the species have secretive and peculiar habits.
~
100 Mr. G. Lewis on
Zabromorphus ordinarius, sp. N.
Ovatus, parum convexus, niger, nitidus; fronte irregulariter punc-
tata; elytris striis 1-3 dorsalibus integris, 4 antice abbreviata,
5 subobsoleta, suturali fere dimidiata; propygidio pygidioque
dense punctatis.
L. 6 mill.
Oval, rather convex, black and shining; the head irregu-
larly punctured, stria complete ; the thorax smooth, with two
lateral strie, interstice at and near the anterior angle with a
few scratches and punctures ; the elytra, striz, inner humeral
strong and abbreviated anteriorly for about one third, 1-3
also strong with fine crenate edges, 4 somewhat similar but
finer and but a little longer than the inner humeral stria, 5
very short, apical, and traceable only as punctures, sutural
similar to the fourth but apical and not reaching beyond the
middle; the pygidia are densely and very evenly punctured ;
the mesosternum is very feebly sinuous, with a well-marked
marginal stria ; the anterior tibize 3-dentate.
The most peculiar characteristic of this species is the sculp-
ture of the thoracic interstice.
Hab. Fort Salisbury, Matabeleland.
Macrouister, Lewis, Ann. & Mag. Nat. Hist. xiv. p. 145
(1904), = Coptochilus, Rey, L’ Exchange (Lyon), part 14,
p- 4 (1888), type Hister major, L.; Pachylister, Lewis,
l. c.,= Heterognathus, Rey, l. ¢., type Hister inequalis,
Ol.
Rey’s names do not appear in the ‘ Zoological Record’ nor
in “ Scudder” or any of the later lists of genera; Rey writes
of them as ‘* sous-genres,”’ but they are natural genera, and
must be recognized as such. ‘There are some specific names
in Rey’s paper also, but they are synonymic and of no
importance.
As Rey’s lines are very short and have escaped notice for
nearly twenty years, I reproduce them :—
“1? Hister major, Lin., & cause de son labre échancré et
des cétés du prothorax densement ciliés-frangés, mériterait de
constituer au moins un sous-genre (Coptochilus, Rey).”
“T) Hister inequalis, F., en raison de ses mandibules
inégales, pourrait également donner lieu & une coupe sous-
eénérique (Heterognathus, Rey).”
new Species of Histeride. 101
This is all; and I regret reinstating names introduced so
casually, but the rules of nomenclature apparently demand it *.
Hister Marseul, sp. n
Breviter ovatus, convexiusculus, niger, nitidus ; fronte stria integra,
labro trigono ; pronoto stria laterali interna integra basi haud
incurvata, externa ante medium abbreviata; elytris striis 1-3
integris, 4-5 basi multo abbreviatis ; pygidio levissime punctu-
lato; mesosterno margine bistriato ; tibiis anticis 3-dentatis.
L. 87 mill.
Shortly oval, somewhat convex, black and shining; the
head, frontal stria complete, labrum triangular; the thorax,
outer lateral stria confined to the region of the anterior angle,
inner strong and complete, with a rather wide interstice, not
incurved at the base, finely crenate behind the neck ; the
elytra, striz, subhumeral deep but shortened at both ends,
1-3 dorsal complete, but the third is lightly impressed apically,
4-5 faint, punctiform and well shortened anteriorly, sutural
stria discal and bending from the suture at both ends; pygidia
somewhat opaque, microscopically punctured, and there is a
fine raised exterior margin to the pygidium ; the mesosternum
is emarginate, with a stria on each side of the emargination,
and behind it, leaving a rather wide margin, is a complete
stria which turns to the outer edge at the suture and does not
join the metasternal stria; the anterior tibiz are 3-dentate.
This species closely resembles £. adjectus, Mars., but it is
less oval and the thoracic stria is not incurved at the base,
the pygidia are more minutely punctured, and the meso-
sternum has two marginal strie. Hister adjectus, Mars., is
reported from Natal, and the specimens I have determined as
such are from Kamerun and the French Sudan; and I may
not know Marseul’s species except from description.
Hab. Region of the Upper Congo.
The form of the pygidium of the above is similar to
the pygidia of Pachylister cafer, Ex., ceylanus, Mars., pygidi-
alis, Lew., besides [Zister adjectus, Mars., and others; but
this structure does not at present suggest itself to be of generic
importance.
Hister similis, sp. n.
Oyatus, convexus, niger, nitidus ; fronte leviter punctulata, stria
* Since writing the above I find that Rey’s names are preoceupied—
the first in Hemiptera (1843), the second in Pisces (1854) and in Coleo-
ptera (1864).
102 Mr. G. Lewis on
integra antice recta; pronoto striis duabus pone oculos coales-
centibus, interstitiis pone angulos punctatis; elytris striis 1-4 et
humerali integris, 5 basi abbreviata, suturali magis longa, humerali
externa brevissima; propygidio pygidioque dense punctatis ;
mesosterno haud sinuato, stria marginali integra; tiblis anticis
ra :
3-dentatis.
L. 77 mill.
Extremely similar to H. eneus, Lew., but differs in being
black and rather less convex. Also there is a fine punctuation
only within the stria at the anterior angle, not a band of
conspicuous points; the outer humeral stria is median and
very short, scarcely more than an elongate puncture, and the
pygidia are not quite so densely punctate.
Hab. Gambesi River.
Mierolister Sheppardt, sp. n.
Ovalis, parum convexus, niger, nitidus; antennis pedibusque nigris ;
fronte stria transversa in media interrupta, stria anterius arcuata ;
pronoto lateribus distincte punctato, stria marginali integra ;
elytris striis humeralibus nullis, 1-3 dorsalibus integris, validis,
4 et suturali equalibus antice abbreviatis, 5 dimidiata; pro-
pygidio sparse punctato; pygidio fere levi; prosterno bistriato,
striis in medio interruptis; mesosterno late emarginato, stria
integra; tibiis anticis 4-dentatis,
L. 33 mill.
This species is larger than WZ. coronatus, Lew., but extremely
like it. It differs by its more oval form, by the transverse
frontal stria being broken in the middle, by the thorax being
more distinctly punctate, by the elytral striz being deeper
and distinctly crenate, and the interstice between the first and
second striz is markedly wider at the base ; the prosternal
stria are interrupted like those of coronatus, and this may
prove to be a generic character of many species. In coro-
natus the elytral striez are less deep and very feebly crenate ;
in both species there is a transverse crenate stria marking the
suture of the mesosternum, and laterally it is continued along
the metasternum.
It is satisfactory to find a second species which possesses
the characters on which the genus was lately established ; to
these characters may now be added the form of the frontal
striz and the similarly interrupted prosternal strie.
Hab. Beira, East Africa (P. A. Sheppard). Several
examples.
“°
new Species of Uisteride. 103
MarGarinotus, Marseul, Mon. p. 549 (18538).
The species of this genus have open circular antennal fosse,
and this structure suggests its position to be near Grammo-
stethus and Atholus. It cannot remain between Contipus and
fTister, where Marseul placed it.
Carcinops Voulogert, Théry.
Tn the Catalogue of 1905 I placed this species erroneously
in Kissister. Mons. Théry has kindly given me the type
example.
Monoplius Braunst, sp. n.
Orbicularis, convexus, niger, opacus ; fronte plana; elytris seriebus
septem plagarum levium subobsoletis, striis dorsalibus nullis ;
prosterno bistriato; metasterno antice foveolato,
L. 43-5 mill.
Orbicular, convex, black, and opaque; the head, lateral
striz reach a little beyond the eye, surface flat and rugosely
punctured ; the thorax with three lateral small punctures and
one behind the eye (these are seen in other species), surface
densely and strigosely punctured; the elytra, strize, outer
humeral distinct and complete, inner also complete but less
marked and varying in individuals, dorsal obliterated; the
smooth circular spaces, as seen in pinguis, Lew. (pl. xx a.
fig. 8, Ann. & Mag. Nat. Hist. xi. 1893), are obscured by
the dense longitudinal strigosity of the general surface ;
the propygidium is densely and longitudinally strigose; the
pygidium is punctured on the disk and rather bro oadly 1 ugose
along its base; the prosternum is clearly punctured, keel
bistriate, the striee bend towards each other in the middle;
the mesosternum, marginal stria complete; the metasternum
has a fovea close to the suture, and there is a more or less
deep median linear impression which connects the fovea in
the base edge, and sometimes there are two small foves on
the first abdominal segment ; the pilosity on the thighs &e.
when existing is reddish and always sparse. The female has
a very faint longitudinal impression before the apex of the
pygidium, but it is only seen in certain lights.
he above is somewhat similar in size to s¢yillatus, Péring.,
in which, however, the ely ‘tral smooth spaces are much more
distinct.
Hab. Willowmore, Cape Colony. Many examples captured
by Dr. H. Brauns, occurring with Llodotermes viator, Latr.
(Havilandi, Sh.). ; be
104 Mr. G. Lewis on
Probolosternus brevistrius, sp. n.
Ovalis, parum convexus, niger, nitidus; fronte minute punctulata,
antice haud impressa, stria marginali integra ; pronoto levissime
et sparse punetulato ; elytris striis 1-4 integris, 4 basi hamata,
2 Cons brevissima, suturali dimidiata ; propygidio fere levi.
L. 3? mill.
an rather convex, black and shining; the head, marginal
stria complete, not impressed ; the epistoma, surface very
minutely and sparsely punctulate ; the thorax, marginal stria
complete, surface sparsely and finely punctured, with an
antescutellar fovea ; the elytra, interstices of strize smooth,
striz, outer humeral fine, inner humeral and 1-4 dorsal strong
and complete, 4 being markedly hamate at the base, 5 is very
short and dorsal, being just behind the middle of the disk ;
pygidia are almost smooth, but a few fine points are seen in
certain lights; the prosternum narrow and bistriate; the
mesosternum markedly marginate ; the tibize are dilated.
The short discal fifth stria distinguishes this species from
the other four at present known.
Hab, Sierra Leone. A single example in the British
Museum.
Eaosternus rufulus, sp. 0
Ovatus, convexus, nigro-piceus, nitidus; fronte conspicue rugoso
punctata ; pronoto stria marginali integra ; elytris striis tenuiter
impressis, margine late mares ; prosterno ane antice conjunctis ;
mesosterno parum acuto, marginato ; pedibus rufis.
L. 33 mill.
Oval, convex, piceous and shining; the head rugosely
punctured, especially on the anterior part, marginal stria
interrupted in the middle, apparently obliterated by the
rugosities ; the thorax, marginal stria complete, surface
rather closely punctured at the sides, less and more finely so
behind the head, and on the disk nearly smooth ; the elytra
somewhat broadly red at the sides and along ‘the apical
margin, the outer humeral is complete, inner very short and
basal, dorsal 1-4 complete, 6 very short and apical, sutural
complete and hamate at the base ; the propygidium is densely
punctured ; the pygidium has a fine punctuation at the base
and is nearly smooth on the apex; the prosternum, striz
widen out slightly at the base, are parallel along the sides,
and join, forming an arched outline anteriorly ; the meso-
sternum distinctly marginate and somewhat acute in the
middle ; the legs are red and the anterior tibize are 6-7-dentate.
new Species of Histeride. 105
This is the third species of the genus known.
Hab. The Matopo Mountains, Matabeleland (Guy A. K.
Marshall).
Reninus Salvini, Lew., occurs in the nests of Geodoma
cephalotes, and it is probable that all the species of the genus
Reninus and Discocelis ave formicicolous. Hpiglyptus (Lister)
costatus, Mars., also associates with the Gicodoma.
Reninus puncticollis, sp. n.
Breviter ovatus, convexus, niger, nitidus; fronte depressa, stria
angulata; pronoto punctato (basi excepta); elytris striis sub-
humerali et 1 dorsali integris, 1-3 postice, 5 et suturali totis
punctiformibus ; tibiis extus valde dilatatis.
L. 33-34 mill.
Shortly oval, convex, black and shining; the head, frontal
stria strong and angulate before the eyes, depressed before
the epistoma, surface with bowed strize representing incom-
plete parts of shallow punctures and set very irregularly ;
the thorax, marginal stria very fine and apparently very
narrowly interrupted behind the middle of the neck, the inner
stria is close to the marginal stria and traceable anteriorly to
a point behind the eye, anterior angles obtuse and reddish,
surface with coarse punctures, points most dense on the outer
part, and the scutellar region is smooth, the basal edge is
narrowly and microscopically strigose; the elytra, stric, inner
humeral and first dorsal are complete and somewhat carinate
on the basal part, 1-3 are abbreviated on or before the middle,
but are continued apically by shallow punctures arranged
more or less in two rows, the fourth stria is hamate at the
base, 5 and sutural are indicated by shallow punctures, the
fifth is shortened at the middle, the sutural continues to the
base ; the pygidia have a shallow sculpture, consisting appa-
rently of crescents or half punctures which have an imbricate
outline ; the prosternum, lobe somewhat truncate anteriorly
and rather strongly marginate, with a few irregular and
shallow punctures, surface microscopically strigose, keel
bistriate, striz join anteriorly before reaching the suture, the
inner stria is abbreviated behind and the interstices of the
strize are smooth, the anterior area of the keel has a few
irregular punctures somewhat like those of the lobe ; the
mesosternum, marginal stria very fine, sinuous in the middle,
and not very close to the edge, from each basal angle pxo-
ceeds a fine bowed stria, unconnected with the lateral striz,
and both ends of them turn outwards, the sides are bistriate,
106 Mr. G. Lewis on
the stria being also common to the metasternum ; the tibia
are dilated on the outer edge.
This species resembles seminitens, Sch., in the punctuation
of the thorax and in the punctiform sutural stria.
Hab. British Guiana, “ Bartica, Demerara” (2. J. Crew) ;
also Cayenne.
Eretmotus planifrons, sp. n.
Orbicularis, convexus, niger, nitidus ; fronte subopaca, immarginata,
haud striata ; pronoto basi utrinque foveolato ; elytris striis 1-2
brevibus ; prosterno opaco minutissime rugoso, antice indistincte
punctato ; pedibus rufo-piceis.
L. 23 mill.
Orbicular, convex, black and shining; the head is some-
what opaque, being microscopically rugose, very slightly
impressed anteriorly, and without a frontal stria; the thorax,
stria feebly sinuous before the basal angles, anterior angles not
very prominent, less so than those of Lucasz, Mars., and foveo-
late at the base close to the outer margin ; the elytra, strig,
outer humeral dimidiate, inner complete, 1-2 dorsal basal and
very short; the pygidia are finely and somewhat irregularly
punctured, the points being most distinct at the bases of the
segments; the prosternum, surface opaque and microscopi-
cally rugose, with indistinct punctures on the anterior half of
the keel, striae rather fine, widely sinuous laterally and widening
out anteriorly somewhat similarly, but rather more, to those
of corpulentus, Lew. (see figure, Ann. & Mag. Nat. Hist. x.
pl. xix. fig. 1, 1892); the mesosternum, marginal stria is
very fine and the fovea shallow.
Hab. Mount Babor, Algeria (A, Théry). Found with
Aphanogaster testaceopilosa.
Eretmotus fovetsternus, sp. n.
Orbicularis, convexus, niger, nitidus ; femoribus nigris, tibiis obscure
rufis ; fronte stria laterali carinata; elytris striis 1 integra, 2-3
postice evanescentibus ; prosterno striis antice abbreviatis ; meso-
sterno foveis laterali profunde et circulariter excavatis.
L. 2} mill.
Orbicular, convex, black and shining; the femora are
black and the tibise obscurely red; the head, surface opaque,
rugosely punctured, sculpture shallow, stria laterally carinate ;
the thorax margined laterally, punctured chiefly behind the
neck and anterior angles, angles truncate ; the elytra, strie,
1 complete, 2-3 basal and evanescent before the middle, outer
new Species of Histeride. 107
humeral carinate and extends beyond the middle, inner com-
plete but only carinate at the base; the pygidia are evenly
and not densely punctured; the prosternum, strie sinuous
and well-marked, but abbreviated anteriorly before the suture,
keel distinctly punctured and obsoletely rugose ; the meso-
sternum, lateral foveze are deep and circular and the margin
is wider than that of H. Leprieuri, Mars.
The species of Hretmotus are difficult to differentiate, but
the punctuation of the keel, the deep circular fovez in the
mesosternum, and the colour of the legs will distinguish the
above from the other two small Algerian species known, viz.
carinatus, Lew., and Leprieurt, Mars.
Hab. Yakouren, Algeria.
There are sixteen species of Hretmotus now known.
Onthophilus niponensis, sp. n.
Ovatus, niger, subnitidus; fronte leviter concava, clypeo quadrato ;
pronoto 6-costato vix dense punctato; elytris 3-costatis, basi in
medio foveolatis ; pygidio in medio longitudinaliter costato, dense
punctato.
L. 22 mill.
Oval, black, somewhat shining ; the head slightly concave
between the eyes, the clypeus is nearly quadrate, being less
widened posteriorly than that of O. globulosus, Ol. ; the
thorax is rather densely punctured, lateral margins narrowly
and slightly raised, surface 6-costate, outer costa short and
near the base, intermediate one third longer and feebly oblique,
the two before the scutellum are short, parallel to each other,
and clearly separate ; the elytra are 3-costate, the interstices
with five fine carinules, and between the third costa and the
suture the outer carinules are crenate, at the base of the
elytra is a deep fovea between the second and third coste ;
the propygidium is densely punctured, with a median longi-
tudinal carina.
This species closely resembles O. globulosus, F., but it
differs by the clypeus being less wide at the base, by the
thoracic punctures being closer, smaller, and less deep, and
the elytral fovea is conspicuous and occupies the whole
breadth of the interstice. As regards the Japanese species,
it is most similar to ostreatus, Lew., which is, however, much
larger, and the median thoracic coste have appendages
behind the neck.
fab. 'Tokio, Japan. Many specimens in the Museum of
Paris and in my own collection.
108 On Two new Freshwater Fishes.
X1IX.— Descriptions of Two new Freshwater Fishes discovered
by Dr. W. J. Ansorge in Mossamedes, Angola. By G. A.
BouLENnGER, F.R.S.
Labeo Ansorgti.
Body strongly compressed, its depth equal to length of
head and contained 4 times in total length. Head once and
2 as long as broad; snout broad, truncate, feebly projecting,
beset with conical horny tubercles; eye nearly perfectly
lateral, its diameter 4 times in length of head and once and 2
in interorbital width; width of mouth, with lips, about half
that of head; rostral flap not denticulated ; lips bordered
with rounded papille, their inner surface with numerous
transverse plice ; two barbels on each side, anterior 4, poste-
rior 3 diameter of eye. Dorsal IV 10, with concave upper
border, a little nearer root of caudal than end of snout, longest
ray slightly longer than head. Anal IIT 5, nearly reaching
root of caudal. Pectoral nearly as long as head, not reaching
base of ventral, which is below middle of dorsal. Dorsal
deeply forked, with acutely pointed lobes. Caudal peduncle
slightly longer than deep. Scales 36 3, 4 between lateral
line and root of ventral, 16 round caudal peduncle. Silvery,
back brownish.
Total length 65 mm.
A single specimen from shallow swamps at Dongwenna.
A very distinct species, to be placed between L. barbatus,
Bler., and L. capensis, A. Smith.
Paratilapia angusticeps.
Depth of body about 3 times in total length. Head very
strongly compressed, 23 to 3 times in total length; lower
jaw projecting ; preemaxillary processes very long, extending
to between the eyes ; snout pointed, twice as long as the eye
in the adult (not longer than the eye in the young); eye 3}
(young) to 5 times in length of head, its diameter at least
equal to interorbital width ; mouth very oblique, extending
to below anterior border of eye; teeth in 3 or 4 series, outer
largest ; scales on cheek small, in 7 to 9 series; large scales
on the opercle. Gill-rakers short, 11 or 12 on lower part of
anterior arch. Dorsal XV-XVI 14-16; spines slightly
increasing in length to the last, which measures } to 2 length
of head. Anal JI] 11-13; third spine stronger and as long
as or a little shorter than last dorsal. Pectoral 3 to $ length
On a new Frog from Mossamedes. 109
of head, not reaching origin of anal. Caudal rounded,
Caudal peduncle as long as deep. Scales denticulate,
36-39 =; lateral lines 21-24/13-17. Brownish above,
yellowish beneath, with dark brown spots or marblings or
with 7 or 8 indistinct dark bars on the body; a blackish
opercular spot; dorsal, anal, and caudal fins with round
brown or blackish spots.
Total length 160 mm.
Several specimens were obtained by Dr. Ansorge. <A
specimen measuring 215 mm., from the Zambesi, has been
submitted to me by Dr. J. D. F. Gilchrist.
Closely allied to P. robusta, Gthr.
XX.— Description of a new Frog discovered by Dr. W. J.
Ansorge in Mossamedes, Angola. By G. A. BOULENGER,
E.R.S.
Rana eryptotis.
Tongue rather feebly notched behind. Vomerine teeth in
two oblique groups between the choanee. Habit very stout.
Head short, snout rounded ; interorbital space narrower than
the upper eyelid; tympanum completely hidden under the
skin. Fingers short, pointed, first and second equal; toes
short, one-third webbed ; subarticular tubercles of toes very
prominent, conical; inner metatarsal tubercle very large,
very prominent, shovel-shaped, sharp-edged, at least as long
as the inner toe ; asmall, round, outer metatarsal tubercle ; a
round tarsal tubercle, just below the tibio-tarsal articulation.
The tarso-metatarsal articulation reaches the eye. Skin
smooth. Pale greyish or brownish above, with dark-edged
grey or brown spots disposed with greater or less symmetry ;
an interrupted dark cross-bar between the eyes ; back some-
times with white or pink dots ; a light vertebral streak often
present ; hind limbs with dark transverse spots, not forming
complete cross-bars ; lower parts white.
From snout to vent 31 mm.
Numerous specimens were obtained by Dr. Ansorge at
Catequero, Ponang Kuma (Dongwenna), and in the Kafitu
Swamps.
The hidden tympanum and the tarsal tubercle well distin-
guish this small frog from &. Delalandii and other allied
species from Africa.
110 Mr. R. C. Wroughton on
XXI.—On the African Mungooses usually referred to the
Herpestes gracilis Group. By R. C. WRouGuHTON.
THE following notes refer to the section of the Herpestine
containing the forms of the small mungoose with a dark
tail-tip (usually black, rarely brown). ‘The members of the
section are found all over Africa, varying somewhat in size
and colour, as was to be expected.
The following is a list of the names already given to
members of this section, so far as I have been able to
ascertain :—
1835. Herpestes sanguineus, Riippell, N. Wirb. Abyss. p. 27.
1835. Herpestes gracilis, Riippell, N. Wirb. Abyss. p. 29.
1835. Herpestes mutgigella, Riippell, N. Wirb. Abyss. p. 29.
1836. Ichneumon ratlamucht, Smith, App. Rep. Exp.C. Afr. p. 42.
1836. Ichneumon Cauu, Smith, App. Rep. Exp. C. Afr. p. 42.
1836. Cynictis melanurus, Martin, P. Z. 8. p. 36.
1838. Herpestes badius, Smith, Ill. Afr. Zool. ii. pl. iv.
1839. Ichneumia nigricaudatus, Geoffroy, Mag. Zool. p. 18.
1847. Herpestes Galiniert, Guérin & Ferret, Galinier, Voy. Abyss.
Atlas, Zool. pl. i.
1848. Herpestes ochraceus, Gray, P. Z. S. p. 138.
. 1849. Herpestes punctulatus, Gray, P. Z.8. p. 11.
. 1850. Herpestes Lefebvret, Desmurs & Prévost, Lefebvre, Voy.
Abyss. Atlas, Zool. pl. i.
13. 1852. Herpestes ornatus, Peters, Reis. Moss. p. 117.
OIE SU CO OIE
coed geet peed
=
14, 1855. Herpestes ochromelas, Pucheran, Rev. Zool. vii. p. 393.
15. 1861. Herpestes iodoprymnus, Heuglin, Nov. Act. Ac, Leop. xxix.
p. 23.
16. 1864. Herpestes Granti, Gray, P. Z.S. p. 561.
17. 1864. Calogale venatica, Gray, P. Z. 8. p. 564.
18. 1877. Herpestes ruficauda, Heuglin, Reis. N.O.-Afr. p. 41.
19. 1877. Herpestes mutscheltschela, Heuglin, Reis. N.O.-Afr. p. 43.
20. 1894. Herpestes Newmanni, Matschie, SB. Ges. naturf. Fr. Berl.
Ope 2
21. 1904. Terpesles ochraceus perfulvidus, Thomas, Ann. & Mag. Nat.
Hist. p. 96.
22, 1904. Herpestes ochraceus fulvidior, Thomas, Ann. & Mag. Nat.
Hist. p. 97.
23. 1905. Herpestes Bocageit, Thomas & Wroughton, Ann. & Mage.
Nat. Hist. xvi. p. 170.
Practically all these forms were described as distinct
species, some even as belonging to separate genera. In
1882, in his paper on the African Herpestine (P. Z.8. p. 59),
Mr. Thomas brought all these forms together as varieties of
one species, i.e. gracilis, Riipp. In 1898 Mr. de Winton,
with increased material to guide him, pointed out that
Mr. Thomas’s variety “d” must rank as a distinct species on
account of differences in skull-characters. At present the
African Mungooses. aL
material is at least five times that which Mr. de Winton
examined. It divides easily into two groups—a smaller, with
a hind foot (cire.) 50 mm. and a skull-length (cire.) 57 mm.,
and a larger, with a hind foot (circ.) 60 mm. and a skull-
length (cire.)62mm. The group of smaller animals subdivides
into (1) Mr. Thomas’s variety “d,” that is Mungos ochra-
ceus, Gray, with its races; and (2) into a group of which the
oldest representative is Cynictis melanurus, Mart. Similarly
the larger forms are separable into two groups, viz. a northern,
in which the crown and nape are coloured like the back, and
a southern, characterized by a tawny suffusion on the head
and neck. ‘This colour-pattern is so striking and constant
that (combined with the fact that a band across Africa from
east to west, in which only a small form is found, separates
the habitats of the northern and southern groups) I deem
myself justified in ranking this form as a distinct species (or,
more correctly, group of species).
Key.
I. Size smaller ; hind foot (cire.) 50 mm,
A. Brain-case swollen anteriorly, postorbital
constriction sudden,
a. Hairs of body annulated. (Abyssinia.) (1) ochraceus, Gray.
6. Hairs not annulated in dorsal region,
which, with tail, is darker than back,
(Somali pacts ta dele alaeiecde se) 2) Gn eeesazor, Los
ce. Hairs of body not annulated. (Somali.) (8) 0. perfulvidus, Thos,
B. Brain-case narrowing gradually forward
to postorbital constriction.
a, Skull larger ; condylo-basal length 60-
62 mm.; zygomatic breadth 33-34.
a‘, General colour “burnt umber”;
face and crown like body; length
c-m' 23mm. (Gold Coast.) .... (4) melanurus, Mart.
b'. General colour “burnt umber”;
crown black; length c-m! 21'7 mm.
(Zanzibar Island.) ate de stares: (5) m, Last, subsp. n.
6, Skull smaller; condylo-basal length 58-
60 mm. ; zygomatic breadth 29.
a‘. General colour yellow ; length e—m' [ Wr.
20mm ATP OLA) s ceclocre soar (6) m. Bocaget, Thos, &
6‘, General colour drab; length c-m!
AZlmni (Cape: Verd:)\..\5 «crass as (7) m. canus, subsp. n.
ce’, General colour red-umber: length
c-m' 20°77 mm. (Zomba.) ...... (8) m. zombe, subsp. n.
II. Size larger ; hind foot (cire.) 60 mm,
A. No tawny suffusion on head and neck.
a. Tail-tip chestnut. (Kordofan.)...... (9) sanguineus, Riipp.
6. Tail-tip black.
a‘, General colour pinkish drab ; hind
foot 583 mm, (Abyssinia.) ,..... (10) s. gracilis, Riipp.
112 Mr. R. C. Wroughton on
&'. General colour brown; hind foot
623mm, W(Abyssinia.) 6a sila (11) s. mutgigella, Riipp.
c!, General colour “ seal-brown ”; hind
foot 58 mm. (Ruwenzori.)...... (12) s. proteus, Thos.
d', General colour “ oliye-buff”; hind
foot 58 mm, (Fort Hall, B.E.A.) (13) s. wee, subsp. n.
B. Tawny suffusion on head, neck, and
shoulders.
a, Hair not or only obscurely annulated.
a‘. Tail-tip brown. (Ugogo.) ...... (14) Granti, Gray.
6. Tail-tip black. (N.W. Transvaal.) (15) vatlamuchi, Sm.
6. Hair annulated. (S. Africa.) ...... (16) Cauui, Sin.
1. Mungos ochraceus, Gray.
1848. Herpestes ochraceus, Gray, P. Z. 8. p. 138.
Gray’s type is in the collection.
The following are 21g MO-SUENE dimensions :—
Head and body 250 mm.; tail 240; hind foot 48.
Skull: condylo-basal length 58; basilar length 53;
greatest breadth 31; palate breadth across p* 20; length
c—m' 19°6.
The skull is recognizable from that of MW. gracilis &e. by
the inflation anteriorly of the brain-case, recalling somewhat
the shape of the quite young skull in both these species.
Gray describes the colour as “ Pale brownish (yellow, very
minutely mixed or punctated with a darker tint.’
Hab. Type locality “ Abyssinia.” (Type, B.M. no.
44. 7. 30. 44.)
5.5.11.3. Near Berbera (Mr. Drake Brockman).
98.6. 9.3. Jefa Muder, Somaliland (Messrs. Hawker &
Cheetham).
59.7.9.10. Coast of Africa (Verreauz).
2. Mungos ochraceus fulvidior, Thos.
1904. Herpestes ochraceus fulvidior, Thomas, Ann. & Mag. Nat. Hist.
xiv. p. 97
Thomas describes this form as grizzled ochraceous, darker
than in the typical form, the median dorsal area unannulated,
strong tawny ochraceous, in continuity with the tawny of
the tail.
Dimensions :—
Head and body 250 mm.; tail 240; hind foot 51; ear 26.
Skull: condylo-basal length 58; basilar length 53 ;
greatest breadth 31; palate breadth across pt 19; length
c-m! 20°5.
African Mungooses. 113
Hab. Type locality Mandeira, Somali. (Type, B.M.
NOVO. 8.95 7 2 rs Atkenson.)
98.3.9.3. N. Haud, Somali (C. V. A. Peel).
3. Mungos ochraceus perfulvidus, Thos.
1904. Herpestes ochraceus perfulvidus, Thomas, Ann. & Mag. Nat. Hist.
xiv. p» 96:
Thomas describes this species as unannulated, “ uniform
bright ochraceous on head, body, and outer side of limbs.”’
Dimensions of the type (measured in the flesh) :—
Head and body 275 mm.; tail 220; hind foot 51; ear 25.
Skull: condylo-basal length 51:5; basilar length 53°5;
greatest breadth 34; palate breadth across p* 19; length
c—m' 20.
The above measurements of body and tail seem to me to
be open to doubt. There is nothing to show that this form
differs at all strikingly in size from the two preceding ones.
Hab, Type locality Wardair, Somali. (Type, B.M. no.
4.5.9.9: Capt. Dunn.)
7.4,4,1-2. Ber, near Burao, Somali (Ir. Drake Brock-
man).
4. Mungos melanurus, Mart.
1836. Cynictis melanurus, Martin, P. Z. 8, p. 56.
Martin’s description is very meagre, but, besides the actual
type, there are several specimens from the same and_ neigh-
bouring localities. The following is a short general descrip-
tion of this form :—General colour “ burnt sienna.” Hairs
of back short (10-12 mm.), ‘‘orange-rufous,” ringed and tipped
with black, forming a grizzling which tends to produce trans-
verse alternate dark and light bands, this tendency extending
much further forward than in the Abyssinian forms, visible
even on the neck behind the ears; face the same colour as
the back, but much more finely grizzled; tail coloured as
back, with the usual black tip.
Dimensions :—
Head and body 300 mm.; tail 280; hind foot 53.
Skull: condylo-basal length (cire.) 63; basilar length
(cire.) 58; zygomatic breadth 34; palate breadth across
p* 21; length c—m'* 23.
Hab. Type locality Sierra Leone. (Type, B.M. no.
55. 12. 24. 229.)
72.2.22.2. Ashanti.
76.10.28.9. Gold Coast ( Col. Strachan).
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 8
114 Mr. R. C. Wroughton on
5. Mungos melanurus Last?, subsp. n.
General colour near “ Mars brown,” with the usual black
tail-tip. Hairs of back 10-12 mm. long, black, with a lower
pale buff and a subterminal tawny ring. Face and crown
almost black, very finely grizzled with tawny; tail coloured
like the back, individual hairs 20 mm. long.
Dimensions :—
Head and body (circ.) 270 mm. ; tail (cire.) 250; hind
foot 50; ear 25,
Skull: condylo-basal length 62; basilar length 56; zygo-
matic breadth (?); palate breadth across p* 21°5; length
c—m' 21°7.
Hab. Type locality Zanzibar Island. (Type, B.M. no.
6.6.9. 11.)
6.6.5.8-16. Zanzibar Island (J. T. Last).
6.6.5. 27-29 (skulls only). Zanzibar Island (J. 7. Last).
The black mask is very noticeable; in a series of nine
specimens it 1s absent in only one, in which the face-hairs
are markedly longer, more coarsely annulate, and a red-
brown is substituted for the usual pale buff.
In the general colour there is a strong likeness between
M. m. Lasti and M. melanurus of the West Coast, but the
black face suffices to distinguish the former. Even the excep-
tional individual mentioned above has a quite different look
to the minutely but distinctly grizzled mask of IL. melanurus.
6. Mungos melanurus Bocaget, Thos. & Wr.
Herpestes gracilis punctulatus, Bocage.
1905. Herpestes Bocager, Thomas & Wroughton, Ann. & Mag. Nat.
Hist. xvi. p. 170.
The general colour is a bright ochraceous, strikingly
different from the red-brown and drab of the preceding more
northern forms.
Dimensions :—
Head and body 265 mm.; tail 235; hind foot 52; ear 25.
Skull: condylo-basal length (probably circ.) 58; zygo-
matic breadth 29; palate breadth across p* 19; length
c—m* 20.
Hab. Caconda, Angola. (Type, B.M. no. 5. 5. 9. 13.)
7. Mungos melanurus canus, subsp. n.
General colour drab. Hairs of the back 15 mm. long,
black, with two cream-buff rings, one of which is subterminal,
African Mungooses. 115
leaving only a very short black tip. Face coloured like
back, but the grizzling finer ; tail coloured like back.
Dimensions :—
Head and body (circ.) 260 mm.; tail (cire.) 250; hind
foot 52.
Skull: condylo-basal length (cire.) 58; basilar length
(cire.) 53; zygomatic breadth 29; palate breadth across
p’ 19; length c—m’ 21°5.
Type (young adult), B.M. no. 72. 12. 12. 5.
Hab. Cape Verd.
8. Mungos melanurus zombe, subsp. n.
General colour “ raw umber,” with black tail-tip.
Hairs of back 10-12 mm. long, dirty white at base, followed
by four rings, black and cream-buff alternately, with a black
tip, but in avery large proportion of the hairs the subterminal
pale ring only gradually darkens to black at the extreme
tip. Face darker than back, almost black, very finely grizzled
with white ; tail and feet coloured like back.
Dimensions :—
Head and body (cire.) 260 mm. ; tail (cire.) 220; hind
foot 48; ear 24.
Skull: condylo-basal length (cire.) 60; basilar length 55;
zygomatic breadth 29; palate breadth across p* 19; length
c—m’ 20°7.
Hab. Type locality Zomba, Nyasa. (Type, B.M. no.
GUGLOL EGE)
93.5. 2.2-3. Zomba, Nyasa (Sir H. H. Johnston).
97.10.1.67. Zomba, Nyasa (Sir H. H. Johnston).
9. Mungos sanguineus, Riipp.
1835, Herpestes sanguineus, Riippell, N. Wirb. Abyss. p. 27,
This form has hitherto been accepted as a species distinct
from M. gracilis. The Natural History Museum has no speci-
men from Kordofan, the type locality of I. sanguineus, but a
specimen from the hills near Suakim can, I believe, be nothing
but a form intermediate between M. gracilis and sanguineus.
It is very pale in general colour and the tail-tip is half
chocolate-brown and half black. There is a cotype of M.
gracilis in the collection, and comparing these two specimens
with Riippell’s plate of WM. sanguineus and gracilis, the Suakim
individual resembles the figure of J/. sanguineus quite as much
as (if not rather more than) the cotype agrees with that of
M. gracilis, In the dimensions recorded by oor there is
116 Mr. R. C. Wroughton oa
practically no difference between those of these two forms.
I have no skull of either typical JM. sanguzneus or even of
the Suakim form which I believe to be closely allied to it.
Riippell gives figures of skulls of M. sanguineus and gracilis,
and at first sight the emargination of the lambdoid crest
(which he particularly notices in the letterpress) and the almost
complete absence of a postorbital constriction in the former
seem to point to a fundamental difference in skull-shape
between the two forms. An examination of the long series
of skulls in the Natural History Museum shows, however,
that in this group both these characters merely indi-
cate immaturity. I think I am justified in concluding that
M. sanguineus and gracilis cannot be specifically separated ;
and as JV. sanguineus is the earlier name that form must be
accepted as the typical one of this group, which extends
through the length and breadth of Africa, with no variation
other than of size and colour.
I follow Riippell in describing M/. sanguineus as “ reddish
isabella colour, grizzled with chestnut and with a rust-
red tail-tip.” Riippell’s dimensions are based on a quite
young individual, and I offer the following (based on adult
M. gracilis and the Suakim specimen) as probably those of a
normal specimen :—
Head and body 300 mm.; tail 325; hind foot 58.
Hab. Type locality Kordofan.
(?) 6. 10. 2.9. Erkowit, Suakim (Af. A. ZL. Butler).
10. Mungos sanguineus gracilis, Riipp.
1835, Herpestes gracilis, Riippell, N. Wirb. Abyss. p. 29.
1847. Herpestes Galinieri, Guérin & Ferret, Galinier, Voy. Abyss. Atlas,
Zool. pl. i.
1850. Herpestes Lefebvrei, Desmurs & Prévost, Lefebvre Voy. Abyss.
Atlas, Zool. pl. i.*
1855. Herpestes ochromelas, Pucheran, Rey. Zool. vii. p. 393.
1861. Herpestes todoprymnus, Heuglin, Nov. Act. Ac. Leop. xxix.
p. 63.
1861. Herpestes adailensis, Heuglin, Peterm. Geog. Mitth. p. 17,
Riippell describes the colour of his type as “ cinereo
flavicans”” and “‘ gelb grau,” but I think the term “ pinkish
drab” used in my key gives a much better idea both of a
cotype in the collection and of the animal represented in
Riippell’s plate. Riippell’s description is evidently based on
a young individual and the cotype mentioned above is also
* This is the name given in the index to the Atlas, but at the foot of
the plate itself is printed “ Herpestes gracilis.”
African Mungooses. 117
young. The proportionally long tail is specially mentioned
by Riippell.
Dimensions :—
Head and body 300 mm.; tail 340; hind foot 58.
Skull: condylo-basal length 62 ; basilar length 37; zygo-
matic breadth 82; palate breadth across p* 20; length c—m'
PAS
Fab. Type locality near Massowa, Abyssinia. (Cotype,
B.M. no. 163 a.)
69.10. 24.11. Soaroo, Abyssinia (W. 7. Blanford).
69. 10. 24. 18. Adigrat, Tigre, Abyssinia (W. IT.
Blanjford).
69. 2.2.6. Abyssinia. Purchased.
Oli Tali “lh. Zuar, Abyssmia (Zephiro, HH. iN.
McMillan).
Hlerpestes Galiniert is from the figure plainly a very
young M. gracilis. Mr. Thomas has quite recently made a
special examination of all the specimens in the Paris Museum
and assures me that the animal figured by Desmurs and
Prévost as H. gracilis, on which both the names Lefeburet
and ochromelas were based, is a pale example of MW. s. gracilis,
in spite of the resemblance that the figure bears to
M. ochraceus.
11. Mungos sanguineus mutgigella, Riipp.
1835. Herpestes mutgigella, Riippell, N. Wirb. Abyss. p. 29.
1889. Ichneumia nigricaudatus, Geoffroy, Mag. Zool. p. 18.
1877. Herpestes mutscheltschela, Heuglin, Reis. N.O.-Afr. p. 43.
Riippell defines MW. mutgigella as “* blackish umber-brown.”
There is considerable individual variation in the extent to
which the body-hairs are annulated. In a cotype from near
Masowa all annulation is completely absent, while in a
specimen from Hrythrea almost all the hairs are annulated,
though very coarsely. In all cases, however, it would seem
that annulation of the hairs is absent on the tail. ‘he form
is rather larger than typical JM. sanguineus and has a rather
markedly short tail.
Dimensions as follows :—
Head and body 350 mm.; tail 320; hind foot 62.
Skull: condylo-basal length 65; basilar length 60;
zygomatic breadth 34 ; palate breadth across p* 21; length
c—m 23.
Hab. Type locality Dembea and Simen Dists., Abyssinia.
(Cotype, B.M. no. 164 a.)
46.6. 15.37. Abyssinia.
118 Mr. R. C. Wroughton on
69. 10. 24. 12. Adigrat, Tigre, Abyssinia (W. T.
Blanford).
3.12.5. 2-3. Chadi Saati, Erythreea (Schrader).
12. Mungos sanguineus proteus, Thos.
1907. Mungos gracilis protets, Thos. Ann, & Mag. Nat. Hist. xix.
p. 119.
A very variable form, much subject to melanism. Mr.
Thomas describes the colour as ranging from “ grizzled
tawny ochraceous”’ to ‘ blackish bistre.”
The recorded dimensions are :—
Head and body 306 mm.; tail 260; hind foot 959;
ear 25.
Skull: condylo-basal length 61 ; basilar length 56 ; zygo-
matic breadth 30°5; palate breadth across p* 19°5; length
c—m! 21°7.
Hab. Type locality Ruwenzori. (Type, BoM. aio;
6. 12. 4. 35.)
6. 12. 4. 31-36. Ruwenzori.
13. Mungos sanguineus ibe, subsp. n.
Size about as in M. s. mutgigella; general colour ‘‘ olive-
buff’ Hairs of back 15-20 mm. long, black, with subbasal
and subterminal rings bright buff, each 2-3 mm. wide;
grizzling of back, as in all the preceding forms, tending to
assume transverse dark stripes on hinder back and flank ;
head and feet more finely, tail more coarsely grizzled ; hairs
of latter, 25-30 mm. long, each with three or four buff rings.
Skull distinctly longer than in any of the preceding
forms, the extra length almost wholly behind postorbital
constriction.
Dimensions of the type specimen :—
Head and body 330 mm.; tail 300; hind foot 60; ear 26.
Skull: condylo-basal length 67; basilar length 62;
zygomatic breadth 35; palate breadth across p* 22; length
c—m! 23°5.
Hab. Kikuyu, British East Africa. (Type, adult 9 , B.M.
no. 4.12.6.7. Collected by 8. L. Hinde, Esq., at Fort Hall,
British East Africa.)
Like the other forms it seems that J. s. cbew has a tendency
to melanism, though much less so than in JZ. s. mutgigella or
s. proteus. Of the two specimens taken at the same place
by Mr. Crawshay, one is quite normal; in the other the
width of the buff rings is much reduced everywhere, while on
the face and along the median dorsal line they are absent.
African Mungooses. 119
92.12. 3.6. Machakos (Ff. J. Jackson),
0.3.27.11. Lé, Galaland (Lord Delamere).
0. 5.3.1-2. Roromo, Kikuyu (2. Crawshay).
0.6. 21.8. British East Africa (Lord Delamere).
2.7.6.13. Fort Hall, B.E.A. (8S. Z. Hinde).
3.4.4.1. Wort Hall; BEA. (S. L. Hinde).
Sells 2. Hort Hall, B.E.A. (S. L. Hinde).
4.2.5. 3-4. Fort Hall, B.K.A. (S. ZL. Hinde).
4.11.5.11-12. Fort Hall, B.H.A. (2. Metnertzhagen).
14. Mungos Grantt, na
1864, Herpestes Granti, Gray, P. Z. 8. p. 5
i se ‘pestes Neumanni, Matschie, SB. ‘Ges. naturf, Fr. Berl,
p. 121.
Matschie’s Md. Newmanni is from Ugogo, and is therefore
a topotype of Grant, Gray, and a comparison of Gray’s type
with Prof. Matschie’s description of JZ, Newmanni shows that
without doubt they are the same form.
General colour ochraceous, all the hairs on the median line
from nose to tail tipped with tawny ; tip of tail chestnut.
Dimensions :—
Head and body (cire.) 320 mm.; tail (circ.) 270; hind
foot 58; ear 25.
Skull: condylo-basal length 63; basilar length 58 ; zygo-
matic breadth 30; palate breadth across p* 21°5; length
c—m} 22.
Hab. Type locality Mgunda Mkali. (B.M. no. 63.7.7. 18,
Capt. Speke.)
15. Mungos ratlamucht, Sm.
1836. Ichneumon ratlamuchi, Smith, App. Rep. Exp. C. Afr. p. 42.
1838. Herpestes badius, Smith, Il. Zool. S. Afr. pl. iv.
«‘ Above chestnut, deepest on the head, neck, and tail; the
hair of the back ringed with dull yellow, that of first half of
tail with brown and yellow ; tip of tail deep black.” This
is Smith’s description of JZ. ratlamucht. In his description of
M. badius he substitutes “ bright bay ” for ‘* chestnut,” and
mentions the black annulations of the hairs of the head and
neck which cause the deepening of colour noted in his
first description. Both descriptions apply to the type
specimen in the Museum Collection.
Dimensions :—
Head and body 300 mm.; tail 290 ; hind foot 58 ; ear 25.
Skull : condylo-basal length Gai: basilar length 58 ; ZY Zo=
120 On African Mungooses.
matic breadth 33; palate breadth across p* 20°5; length
c—m! 21°5.
Hab. Type locality (of IZ. ratlamucht) ‘ between Latakoo
and the Tropic” ; (of MZ. badius) “ between Old Latakoo and
Karichane, 120 miles eastward.” (Type, B.M. no. 46.6.1.14.)
16. Mungos Cauui, Sm.
1886. Ichneumon Cauui, Smith, App. Rep. Exp. C. Afr. p. 42.
1849, Herpestes punctulatus, Gray, P. Z. 8. p. 11.
1852. Herpestes ornatus, Peters, Reise Moss. p. 117.
1864. Calogale venatica, Gray, P. Z. S. p. 563.
I have been able to examine about 40 specimens from
localities spread over 10° of longitude and 15° of latitude,
and I have to confess that I cannot differentiate any of them
as even racially distinct from the rest. It is true that a
series from Matabeleland is on the average paler than the
rest, as a series from Mashonaland and the Zambesi basin is
somewhat redder, but single individuals could be picked from
other localities which are quite as pale or as red. It is to be
regretted that Smith’s name Cauuz is the oldest, it was given
to a young specimen from the neighbourhood of the Kalahari
Desert, a quite extreme example of the pale desert type.
Peters’s M. ornatus is also based ona quite young specimen *
of a darker redder type, while JZ. punctulatus, Gray, un-
doubtedly represents the normal form,
The general colour is hard to describe, it is some shade of
drab tinged with greenish, having in all casesa greater or less
suffusion of reddish on the loins and rump.
Dimensions :—
Head and body 300 mm.; tail 275; hind foot 58; ear 25.
Skull: condylo-basal length 65 ; basilar length 60; zygo-
matic breadth 33-35 ; palate breadth across p* 20-23 ; length
c-m' 22.
Hab. Type locality Currichaine (N.W. Transvaal).
45.4.4. 2. Durban, Natal. (Type of 1. punctulatus, Gray.)
5.3.8.13. Illovo, Natal (udd Collection).
4.12.3. 36-38. Umvalosi, Zululand (Rudd Collection).
4,.5.1.33-36. Htshowe, Zululand (Rudd Collection).
4.9.1. 32-34. Zuurbronn, EH. Transvaal (Rudd Collection).
* Peters’s figure of the skull of M. ornatus shows all the sutures
which are only visible in youth. A specimen in the collection from the
Zambesi not far from Tette, the type locality of M. ornatus, answers both
in dimensions and colouring to Peters’s description ; itis quite young: an
older specimen from the same locality is identical with other S. African
individuals in dimensions and colour-pattern.
On the Armenian Wild Sheep. 121
98.4.4.13. Krugersdorp, West Transvaal (17. P.
Thomasset).
46. 6.2.23. Mohopaui Berg, Bechuanaland.
97.1.4.1-4. Matabeleland (/’. C. Selous).
99. 2.23.1. Ngamiland (Capt. Lugard).
93.11.21.1. De Kaap (Dr. P. Rendall).
6. 4. 8. 31-36. Woodbush, N.W. ‘Transvaal (Rudd
Collection).
5.12.9.19-20. Klein Letaba, N. Transvaal (Rudd
Collection).
6.8.2.49. Legogot, N.E. Transvaal (Rudd Collection).
6.4.7.1. Sabi River, N.K. Transvaal (J. S. Hamilton).
(sey 25, 4, Salisbury, Mashonaland (Guy Marshall).
97. 1. 4.5. Salisbury, Mashonaland (Ff. C. Selous).
99. 8.3.4, Mashonaland (Boyd Alezander).
Gaile a 21. N. Rhodesia (S. A. Neave).
1.6.26.2. Pasa (Col. Manning).
9
7.10.1.66. LL. Nyasa (Sir H. Johnston).
XXII.—The Name of the Armenian Wild Sheep.
By R. LypeKKer.
CONSIDERABLE diversity of usage prevails among naturalists
with regard to the scientific name of the Armenian wild sheep,
some writers adopting Ovis Gmelini, Blyth (1840), while
others prefer Ovis orientalis, When the latter name is used
the authority is generally given as G. I’. Gmelin, ‘ Reise,’
vol. 11, p. 486 (1784); but the animal is there referred to
merely as “das orientalische Schafe,’” which is, of course,
not a technical name. On the other hand, in Brandt and
Ratzeburg’s ‘Getreue Darstellung und Beschreibung der
Thiere, etc.,’ Berlin, 1829, p. 54, pl. ix. fig. 1, we find the
wild sheep of the “ Ceraunian Mountains ”’ of Persia described
and figured as Ovis musimon, var. ortentalis; and this name,
moditied to O. orientalis, consequently stands. It may be
added that the work in question is really composed of extracts
from ‘ Medizinische Zoologie,’ a serial of which the first
volume, containing the name in question, appears to have
been published in 1827.
As to the “ Ceraunian Mountains” of Persia, I take it
that this must be an application of the name in a sense
analogous to that in which the term “ Alps” is often used,
seeing that the Ceraunian or Acroceraunian Mountains are
122 Mr. T. D. A. Cockerell— Descriptions and
in Albania. It probably refers to the south side of the
Elburz range.
To the sheep of this locality I have given (1905) the name
Ovis Gmelini Erskinet, but this will now have to give way
to O. orientalis typica, while the Armenian race will have to
be called O. orientalis Gmelint, Blyth’s specimens having
come from Erzerum.
X XIII.—Descriptions and Records of Bees —XVI.
By Tl’. D. A. CocKERELL, University of Colorado.
Osmia Bennette, sp. n.
$ .—Length about 9 mm.
Very brilliant, shining, Auwgochlora-green, with coppery
and golden tints on the face and mesothorax, and a certain
amount of golden lustre on the abdomen; antenne entirely
black, not moniliform ; clypeus with long white hair; hair of
front, vertex, and thorax above yellowish white or pale
yellowish, without any admixture of dark hairs except a very
few about the ocelli; tegule brilliant green. Wings clear,
the apical margin a little dusky. Legs green, with mostly
pale hair, but some dark, e. g. the middle tibia has dark hairs
intermixed, and the middle basitarsus has much black hair.
Dorsal hair of abdomen partly light and partly black, the
black prevailing posteriorly ; apex of sixth segment very
feebly notched ; seventh bidentate; venter with strong blue
or purple tints.
From O. gaudiosa, Ckll., this is easily known by its larger
size and the conspicuous black hair of the hinder part of the
abdomen. ‘The size and green tegule suggest aftinity with
O. bruneri, Ckll., but I do not think it can be the male of
that insect, the colour of the pubescence being so entirely
different. ‘The quite different colour of the tegument is not
so important, as that may differ sexually in Osmia, e. g. in
QO. versicolor, Latr. From QO. bella, Cress., it is readily
known by the absence of dark hair on the thorax above.
‘The width of the abdomen is 3 mm., thus much broader than
O. fulgida, Cress.
Hab. Campus of University of Colorado, Boulder, Colorado,
at flowers of Zarawacum taraxacum, May 8, 1907 (Mrs. C.
Bennett).
Records of Bees. 123
Osmia Edne, sp. n.
? .— Length about 7 mm.
Very brilliant golden-green, a little reddish on second
abdominal segment; hair of head and thorax above long, pale
ochraceous, with no dark hairs intermixed; on clypeus the
hair is yellowish white; flagellum ferruginous beneath, not
moniliform ; mesothorax densely rugoso-punctate ; tegule
brilliant green. Wings dusky hyaline. Legs green, with
yellowish-white hair; hind femora almost black behind.
Abdomen subglobose, with the hair enterely pale yellowish or
yellowish white ; sixth segment with a minute feeble notch ;
seventh strongly notched rather than bidentate.
Hab. Campus of University of Colorado, Boulder, Colorado,
at flowers of Zarawacum taraxacum, May 9, 1907 (Aliss Edna
Baker).
The following table separates this species from some other
bright green males :—
Posterior part of abdomen dorsally with some or
much: black, hair srsize larger oa. + 6 0s )3 0003s UG
Posterior part of abdomen dorsally with wholly light
Haine Size smaller racecar ae wists cae casts) «shee 2.
1, Hair of thorax above with some dark hairs inter-
BUCO US a eePerk vapeur atttts tales oletel sites ae sls adnate « bella, Cresson.
Hair of thorax above without dark hairs intermixed. Sennette, Ckli.
2. Size smaller; pubescence white; abdomen dullish,
MO SHINING SOLMENM see ealel clelele cis zsoe ee ws mals gaudiosa, Ckll.
Size larger; pubescence very yellow; abdomen
brilliant; sbimine eoldeni ii. Gesclecten sce sees Edne, Cll,
Osmia leonis, sp. n.
? .—Length about 113 mm.
Head and thorax partly dark green, partly black ; abdomen
shining dark blue; legs black ; ventral scopa black ; cheeks
normal, Head ordinary, with large strong punctures ; man-
dibles tridentate, the apical tooth long and falciform, the
second triangular and sharp, the third a long undulating
ridge; anterior part of clypeus purple-black, this colour
extending also up the middle; edge of clypeus straight, but
at the middle next to the edge there is @ transversely oval
shining pit of quite large size; supraclypeal region with a
shining green mark like an inverted V; antenne entirely
black, comparatively short. Hair of clypeus, lower sides of
face, and cheeks black; of front black and pale ochreous
mixed ; of vertex mostly black, but of occiput pale ochreous,
extending forward over ocelli; hair of pleura black, but of
124 Mr. T. D. A. Cockerell—Descriptions and
thorax above light ochreous, with black hair sparsely inter-
mixed on scutellum and hind part of mesothorax ; tegule
black. Wings clear in the middle, but with the hind margin
broadly and very distinctly infuscated ; a very dark streak
in upper part of marginal cell. Hair of legs black, dark
reddish on inner side of tarsi. Abdomen short and convex,
shining dark blue, with slight crimson or purple tints on
second and third segments: hair of first segment pale except
at extreme sides ; of second black at extreme base, at sides,
and apex, but long and pale on disk; remaining segments
with black hair.
In my table of Boulder County Osmia (Univ. of Colo.
Studies, ined.) this runs to O. nigrifrons and O. gaillardia,
resembling the latter in superficial appearance (especially the
shining abdomen), but differing in the colour of the pubes-
cence, the clypeal pit, &c. The second r. n. joins the second
s.m. nearer its end in gadllardie than in leonits. In gatl-
lardie the little brushes of hair from beneath the clypeus are
yellow; in deonds they are pure black. From nigrifrons the
new species is more easily separated; among other things the
punctures of the face are finer and smaller in negrifrons than
in feonts. "There is evident affinity with O. juata, Cresson,
but the admixture of black hair on the thorax above and the
light hair on second abdominal segment, as well as the clypeal
pit, should suffice to distinguish deonts from that insect.
Hab. Campus of University of Colorado, Boulder, Colorado,
at flowers of Taraxacum tararacum, May 8 (Edna Baker).
On the hair of the abdomen were some mites of the genus
Trichotarsus, which Mr. N. Banks says are 7. osmie, Duf., or
more probably a new species allied to it.
Osmia integrella, sp. n. (possibly wntversitatis var.)
3g .—Length 10 or 11 mm.
Steel-blue, with the head and thorax above, the first abdo-
minal segment, and a strong tinge or suffusion on the apical
part of the other segments green; hair Jong, the pale hair
white, not ochreous, no admixture of black hair on head and
thorax above, except that some black hair on upper part of
cheeks has a few outlying hairs on extreme sides of vertex;
cheeks with hair white, but some black behind and above.
Antenne black, flagellum not moniliform; clypeus normal;
pleura with white hair except on posterior margin, starting
with a patch just beneath wings, where it is black; extreme
sides of metathorax with black hair; tegule green in front.
Wings little dusky. Legs black, not at all metallic, their
Records of Bees. 125
hair largely black, but some pale on middle and hind femora,
and long and mainly pale on anterior femora and _ tibie
behind. First two abdominal segments with long white hair,
some black at extreme sides of first segment, and short black
hair along extreme base of second ; third with mixed black
and pale; the other segments nearly all black; in certain
lights there is a strong suggestion of pale hair-bands; sixth
segment quite entire; seventh with a pair of comparatively
short teeth, wide apart; first ventral segment entire, third
deeply emarginate, the emargination ciliate with reddish.
The second and third joints of middle tarst have a strongly
swollen or inflated appearance. ‘Third antennal joint shorter
than fourth. Spurs normal.
This must be closely allied to O. integra, Cress., which I
know only from the description ; but it is smaller, the hair
not (or barely) ochreous-tinted above, the legs apparently
with more pale hair ; the abdomen with more pale hair; and
the fringe on middle of third ventral segment can hardly be
said to be long and golden. Cresson also makes no allusion
to any peculiarity of the middle tarsi of dntegra, though in
the species described just before he describes and figures
tarsal structures. It does not fit into any of the Robertsonian
subgenera.
In my table of Boulder County species O. integrella runs
nearest to O. viridior, or, perhaps, to cyaneonitens. The
three are readily separated thus :—
Larger ; hair on anterior tibiee behind dense, rather short,
and wholly black, contrasting with the long white
hair.on' their femora behind)... oi 2s eee asks viridior.
Smaller; hair on anterior tibiz behind long and mainly
WALLS Herp 8 cedars AN Ah ae es tats etehiafale Meu MAGA Lah: integrella.
Size about as in integrella, but very different by the dark
purple-blue abdomen, without conspicuous light hair,
much larger head and broader face, notched sixth
SIITETE OOF anne anicoino dip. cocnt: Opec coneces bienn cyaneonitens,
The face of integrella is rather unusually narrow, the eyes
converging below, and is densely covered with white hair.
O. universitatis, Ckll., has the same type of coloration and
the same sort of middle tarsi as integrelia; but it is smaller
than the latter and has not the conspicuous black hair on the -
sides of the thorax posteriorly or on the cheeks. In wniversi-
tatis the middle femora and tibia have the hair behind
brilliant white, with some black intermixed ; in dntegredla
this hair is black, with a few glittering white hairs. The two
are obviously of the same immediate group, but apparently
not varieties of a single species.
126 Mr. T. D. A. Cockerell—Descriptions and
The stipites of O. ¢ntegrella are divided at the end into a
brush of black hairs and a divergent long, linear, corneous
process.
Hab, Campus of University of Colorado, Boulder, Colorado,
at flowers of Taraxacum taravacum, May 8 (Edna Baker).
Osmia Ramaley?, sp. n.
3 .—Length about 73 mm.
Head olive-green, the front almost golden; thorax above
yellowish green, at sides bluish green; the metathorax blue,
contrasting with the green scutellum and_ postscutellum.
Abdomen a sort of Prussian green or greenish blue. Legs
greenish blue, but the anterior legs black in front; tegule
green. Wings clear, with only a very faint duskiness, the
hind margin no darker than the rest. Hair of head and
thorax long and white (not ochreous above), very dense on
face ; no dark hairs anywhere; hair of legs white, fulvous on
inner side of tarsi; middle tarsi rather long, slender, the
joints normal. Antenne black, the flagellum not moniliform ;
third joint shorter than fourth. Abdomen rather coarsely
sculptured; hind margins of segments concolorous with the
rest ; sixth segment with a broad, very shallow emargination ;
seventh strongly bidentate; first ventral truncate. Hair on
abdomen above white, without any black.
In the table of Boulder County species this runs to
O. proxima, Cresson, which is, however, much smaller and
otherwise different. There is a good deal of resemblance to
O. atriventris, Cress., but the green tegulz and the structure
of the sixth abdominal segment are distinctive. O. Wheeler
has the legs much less metallic, the tegule reddish in the
middle, and, especially, the teeth of the seventh abdominal
seoment very much larger and triangular.
Hab. Campus of University of Colorado, Boulder, Colorado,
at flowers of Taravacum taraxacum, May 10 (Hdna Baker).
Named after Professor Francis Ramaley, of the University
of Colorado, in recognition of his work on the natural history
of Colorado.
Since the above was written, on May 21, Mrs. C. Bennett
has taken at Boulder 2 2? and 1 ¢ of O. Ramaley?, all at
flowers of Astragalus goniatus, Nuttall. ‘The male is exactly
like the type, except that the first ventral segment is very
distinctly emarginate. This character is considered by
Robertson to be of generic value, but after minutely com-
paring every part of the two bees I am sure that they are of
the same species. ‘The female has an orange scopa and is
Records of Bees. 127
almost exactly like O. hypochrysea Rohwert, Ckll., except
that the clypeus is quite normal (not quadridentate) and the
disk of the mesothorax is very much more densely punctured.
The legs are blue.
This certainly belongs to the subgenus Xanthosmia, but it
is not Robertson’s X. cordata, being much smaller, with the
third antennal joint in the male nearly as long as the fourth.
O. tridis from New Mexico, which also has the first ventral
seoment (fg) emarginate, is easily separated from O. Ramaleyi
by its larger size and the character of the hair on the abdomen.
Osmia olivacea, Ckll.
This was described from a male, but Mrs. C. Bennett has
taken a female at Boulder, May 21, 1907, at flowers of
Astragalus goniatus, It is a remarkably fine insect, coloured
like the male, but nearly 12 mm. long, very robust, the
ventral scopa black ; coarse black hair on the clypeus, but
pale ochreous on sides of face, and these colours mixed on
front; hair of thorax above as in male; hair of pleura pale
ochreous, rather scanty; clypeus normal; mandibles _ tri-
dentate, but the inner tooth broad and notched; legs black,
not metallic. The abdomen is a very beautiful deep oliva-
ceous green, with some slight crimson stains.
Also at the Astragalus goniatus, on the same day,
Mrs. Bennett took two females of Osmia nigrifrons, Cresson,
variety, and one of O. coloradella, Ckll.
Celioxys Portere, Ckll.
Mr. N. Banks sends me a 9 of this New Mexico species,
which he took at Falls Church, Virginia, July 21—a most
unexpected extension of range.
Melissodes Boltonie, Robertson.
Falls Church, Virginia, August and September (V. Banks).
Very like MW. perplexa, but smaller. Differs from MV, dllata
by the smaller average size, band on middle of second abdo-
minal segment entire or almost, and tuft on end of hind
femora pale.
Melissodes manipularis, Smith.
Falls Church, Virginia, 2 ¢, one from flowers of Eupa-
torium, Sept. 4 (V. Banks).
Much like JM. trinodis, but differs by the black hair on
128 Mr. T. D. A. Cockerell—Deseriptions and
middle of thorax above. Smith does not mention this hair,
but I have examined his type.
Melissodes nivea, Robertson.
Falls Church, Virginia, 4 ¢, Sept. 4 and 8 (NV. Banks).
Andrena nigre, Robertson.
Boulder, Colorado (Rohwer).
New to Colorado. At Boulder, on May 22, 1907,
Mr. Rohwer took females of three species of Andrena, all
having red abdomens, at the flowers of Salix luteosericea,
Rydberg. These, upon examination, prove to be A. nigra,
Robertson, A. Marie, Robertson, and A. erythrogastra
(Ashmead). The known range of nigre is extended about
800 miles westward.
Andrena saccharina, sp. n., Cockerell and Rohwer.
3 .—Length about 8 mm.
Black, with greyish-white hair, not nearly dense enough on
thorax above to hide thesurface. Head rather large, quadrate,
facial quadrangle much broader than long; front with coarse
vertical striz ; cheeks broad, with the rounded posterior angle
a little above level of middle of eye. Antenne long, third
joint longer than fourth, but a trifle shorter than 4+5;
flagellum very obscurely brownish beneath, the middle joints
longer than broad. Mandibles strongly grooved, bidentate,
the apex reddish ; process of labrum broadly rounded, not at
all emarginate ; malar space large and shining ; clypeus much
produced, mainly light yellow, with strong but very sparse
punctures ; the yellow is invaded by black above and below,
so that its lower margin is convex (the apical margin of
clypeus being black) and its upper part is notched above and
deeply on each side; mesothorax and scutellum dull, minutely
tessellate, with sparse feeble punctures ; area of metathorax
merely roughened, scarcely defined; tegula dark in front,
shining brown behind. Wings yellowish, iridescent, stigma
(of normal size) and nervures ferruginous; second s.m.
narrow, receiving the r.n. near its middle. Legs black, with
light hair. Abdomen with asericeous surface, not punctured,
the hind margins of the segments obscurely reddish, and with
very thin, not conspicuous, bands of white hair.
On account of the produced clypeus and large malar space
this is related to A. leptanthi, V. & C., but it is a very distinct
Records of Bees. 129
species. It was given to me by Mr. 8. A. Rohwer, who had
already studied it and determined that it was new.
Hab. Sugar Loaf Mountain, Boulder County, Colorado,
8500 ft., May 18, 1907, at flowers of Arctostaphylos uva-uasi.
Collected by Miss Edna Baker,
Perdita quadrangularis, sp. n.
3 .—Lenegth 4 mm.
In my tables of Perdita runs to P. spheralcee, of which it
looks like a small edition. From P. erigeronis it is easily
known by the pallid nervures and the coloration of the
abdomen. The face-markings are of a chrome-yellow
instead of lemon-yellow, but otherwise agree, as do all the
other markings of the head, with spheralcee. The face
below the antenne is ail yellow, the lateral marks being large
and quadrangular, ending on the orbital margin at an angle
of about 45°. Antenne chrome-yellow, the first four or five
joints black-spotted above. Thorax, legs, and wings essen-
tially as in spheralcee, but margin of stigma yellowish.
Abdomen with the venter yellow, as in spheralcee, but on
the dorsum the dark colour is reduced, the general effect
being that of equally broad light and dark bands, with the
apical segments a sort of yellowish ferruginous. The thorax
has much white hair.
Hab. Alamogordo, New Mexico, May 15, about eighty-five
specimens (H. L. Viereck).
P. spheralcee does not appear on the wing until long after
midsummer.
Perdita Vierecki, sp. n.
The female runs in the tables of Perdita to male P. tarda,
the male to P. exclamans. Both sexes were taken in large
numbers; otherwise one would suspect the insect to be a
variety of P. exclamans, to which it is very closely allied.
? .—Length about 43 mm.
Differing from eaclamans as follows :—Size smaller ;
abdomen above piceous, with the yellow markings reduced
to a variable series of spots occupying the middle of the
segments; thus the first segment may have a pair of dots or
no light marks at all; the second may have a large quad-
rangular pale yellow patch ora yellow band on the middle
third; the third segment, and also the fourth, may be more
or less banded, the band in no case approaching the lateral
margin; or the fourth and fifth may each have a couple of
spots ; thus the abdominal markings are extremely variable,
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 9
130 Mr, T, D, A, Cockerell—Descriptions and
but in all cases the colour is very different from that of
exclamans, which has broad yellow bands, mostly reaching
the lateral margins.
3 .—LEssentially as in exclamans, but the head-like ex-
tension of the yellow above the antenne in middle line is
much smaller, and the yellow patch on the pleura does not
send a band to the middle coxe.
Hab. Alamogordo, New Mexico, April 26 to May 15,
about 110 specimens (ZH. L. Viereck).
Although this series is manifestly distinct from P. eacla-
mans, some of the specimens show more or less evident
transitional characters, suggesting that the insect should
perhaps take only subspecific rank, This is one of those
cases, of which we now know several in Perdita, which
promise to yield facts of extreme interest to the evolutionist
when carefully studied in the field.
Perdita phacelie, Ckll.
Alamogordo, New Mexico, May 13 and 15 and June 6,
33 specimens (HH. L. Viereck).
I have compared the Alamogordo specimens with cotypes
of phacelie and cannot see any difference. The latter,
however, were taken early in September. Are we to suppose
that this species winters over in the adult state, or is it
double-brooded? The small size and dull mesothorax readily
separate this from P. @nedfrons.
Perdita pectidis, Ckll.
This also has been known as an autumn species, flying in
September. Mr. Viereck took at Alamogordo, April 24 and
May 3, a series of about 65 specimens, which exhibit much
variation, but do not seem to be separable from P. pectidis.
In the female the clypeus may be three-spotted or may
lack the middle spot, and even the lateral spots may be very
small. The banding of the abdomen varies from a couple of
dots only to well-developed bands. In general, however, the
insect agrees excellently with pectedis, and I do not think it
possible to regard it as distinct.
Perdita chamesarache, Ckll.
Highrolls, New Mexico, June 11, 1902 (H. L. Viereck).
This species was found flying at Albuquerque and Santa I'é
in August.
Records of Bees. Poe
Perdita Rehni, sp. n.
? .—Length about 43 mm.
In the table of New Mexico Perdita runs to P. pallidior,
but is easily distinguished from that species by the smaller
size, much darker abdomen, colour of antenne, &c. Head
and thorax yellowish green, the mesothorax dullish (not
brilliantly polished, as it is in many species) ; head small ;
front shining green; clypeus and supraclypeal area purplish
black ; the only pale face-marks are the small L-shaped pale
yellowish lateral marks, which send a very fine broken line
up the orbital margin to a little above level of antenne,
Mandibles whitish ; cheeks dark, with white hair. Antenne
dark above, pale yellowish beneath, the dark more prominent
than the light; border of prothorax and tubercles light
yellow; pleura all dark. Anterior legs entirely light yellow,
or the femora may have a brown patch, and the tibiz a line,
behind; middle femora all light yellow, but the tibize mainly
brown on outer side and the tarsi somewhat darkened ; hind
femora with the apical half above dark brown and their tibize
and tarsi brown; nervures colourless, but stigma margined
with brown; marginal cell with the substigmatal portion
much the longest ; third discoidal distinct. Abdomen above
dark brown, the first segment with a transverse light yellow
discal mark ; segments 2 to 4 each witha basal light yellow
band, but the bands on 3 and 4 may be nearly concealed by
the retraction of the sezments, giving the appearance of a
one-banded abdomen; fifth segment, apex, and ventral surface
reddish yellow. Sometimes the fifth segment has a very
broad yellow band on a dark ground, or it may be all dark.
Hab. Alamogordo, New Mexico, June 9, 1902 (HZ. ZL.
Viereck).
Named after Mr. Rehn, the well-known orthopterist and
mammalogist, who was Mr. Viereck’s companion on the 1902
expedition.
Prosopis Cressoni, n. n.
Prosopis pygmea, Cresson, Proc. Boston Soc. Nat. Hist. xii. (1869)
p- 272 (Lllinois).—Not P. pyymea, Schenck, 1853,
Nomada flavoguttata Alfkent, n. n.
Nomada pygmea, Schenck, Berlin. ent. Zeits. xviii. (1874) p, 342.—
Not WV. pygmea, Cresson, 1863,
g*
132 Dr. F. D. Godman on American
Megachile Grantiana, n. n.
Megachile punctatissima, W. F. Kirby, Bull. Liverp. Mus. iii, (1900)
p. 20.—Not of Spinola, 1806.
Hab. Sokotra (Ogilvie- Grant and Forbes).
I informed Mr. Kirby of the preoccupation of the name
proposed by him; but as he is no longer working on Hy-
menoptera, he asks me to rectify the matter.
Megachile paucipunctulata, W. F. Kirby.
Megachile paucipunctulata, W. F, Kirby, Bull. Liverp. Mus. iii. (1900)
p- 21.—Sokotra.
N. syn.: Megachile sokotrana, Friese, Zeits. f. Hym. und Dipt. 1903,
p- 287.—Sokotra (Simony).
Megachile subsericans, n. n.
Megachile vicina, Moraw. Here Soc. Ent. Ross. xxviii. (1894) p. 87.—
Turkestan.—Not M. vicina, Mocsary, 1879.
Megachile mixtula, n. n.
Megachile mixta, Radoszkowski, Bull. Soc. Nat. Moscou, xlvii. (1874)
p. 188.—Caucasus.—Not M, mixta, Costa, 1863.
Boulder, Colorado, U.8.A.,
June 5, 1907.
XXIV.—WNotes on the American Species of Hesperiide
gourd by Plétz. By F. D. Gopman, D.C.L., F.R.S.,
ec
J HAVE recently had an opportunity of examining the large
number of coloured drawings made by the late Carl Plitz,
and am at last enabled to identify most of the Mexican and
Central-American species described in his various writings.
These drawings not only illustrate the numerous Hesperiids
named by him, but nearly all those tabulated or described by
Herrich-Schiiffer and other continental authors, and they are
therefore of the greatest interest to all students of Rhopalo-
cera. The following notes on his figures of the American
species, generally, give the necessary corrections to the
synonymy, &c.* ‘The numerals in parentheses after each
name indicate the number of Pl6tz’s drawings, and the locality
& Drawings nos. 1328-1833 are unfortunately missing, and one
Central-American species, Netrocoryne coronus, Plotz, still remains
unidentified by me.
Hesperiide described by P itz.
133
is added either from the drawing or the published description.
In a few cases the locality has not been recorded.
Coloured
copies of the figures of all the unidentified American species
have been made and presented by me to the Natural History
Museum at South Kensington.
Goniurus pilatus, Plétz (2), Brazil
and Guiana.
9 procne, Plotz (3), Brazil.
zagorus, Plotz (4), Alla-
era.
zalanthus, Plotz (5), Alla-
ora.
» élongatus, Plotz (8), Brazil.
» nicasius, Plétz (9), Brazil.
Eudamus flammiula, H.-S. (18),
Loe. ?
Goniurus procerus, Plotz (21), Peru.
AS elius, Plétz (22), Para.
retractus, Plotz (24), La
Guayra.
» larius, Plotz (26), Cuba.
Pf velinus, Plotz (27), Bahia.
» géalbula, Plotz (80), Brazil.
» Keferstemt, Plotz (32),
Caracas.
» protecides, Plotz (55),
N. Am.
rtvion, Selotz «(36)5) Rae
Janeiro.
5, herophilus, Plotz (43), Rio
Janeiro.
» leucodesma, Plitz (47),
Para.
» Aypozonius, Plétz (53), La
Guayra.
Eudamus fulminans, H.-S, (63),
Loc. ?
. zopyrus, Plétz (64), Su-
rinam,
= Eudamus semplicius, Stoll.
= Eudamus simplicius, Stoll.
Probably a small form of Zuda-
mus eurycles, Latr.
= Eudamus eurycles, Latr.
Very near Eudamus undulatus,
Hew., but with the outer dark
band only present on the
secondaries beneath.
= Eudamus undulatus, Hew.
= Eudamus indora, But]. Herrich-
Schaffer’s name has priority.
Belongs to Hudamus. Not in
the G. & S. coll.
Belongs to Eudamus. Not in the
G. & S. coll.
The St. Vincent and Grenada in-
sect recorded by G.& 8. under
the name Eudamus santiago,
Luc., belongs to this species.
=Eudamus santiago, Lue. (=cory-
don, Butl.).
Probably a var. of Eudamus do-
rantes, Stoll.
Belongs to Eudamus. From Co-
lombia, Guiana, and Brazil in
the G. & §. coll.
= Eudamus dorantes, Stoll.
= Eudamus proteus, L., var. with
the hyaline spots on the pri-
maries very small. Specimens
from the Lesser Antilles in the
G. & S. coll.
= Eudamus catillus, Cr.
= Eudamus virescens, Mab.
= Eudamus cholus, Plotz. Descrip-
tion unpublished ?
= Goniurus ceelus, Cr,
= Thymele mephitis, Hew. Herrich-
Schaffer’s name has priority.
Probably belongs to Thymele. The
markings are very similar to
those of Eudamus miltas, G.
& 5.
134
Eudamus erycina, Plotz
(66),
Brazil.
4 aulus, Plotz (67), Brazil.
Telegonus granadensis, Méschl. (70),
Venezuela.
Eudamus briccius, Plétz (72), §
Am.
» passalus, H.-S. (74),
Loc. ?
55 orphne, Plotz (79), Rio
Janeiro.
5 orpheus, Plotz (80), Para.
» justus, Plotz (81), S. Am.
Telegonus bahianus, H.-S. (89),
Loe. ?
” pseudochalybe, H.-S.
(87), Loe. ?
Eudamus Hopfferi, Plotz (88), 8
Am.
Telegonus cretellus, H.-S.
(89
Loe. P )
Eudamus xagua, Lue. (90), Cuba.
49 blasius, Plétz (93), Cuba.
5 latimargo, H.-S. (94),
Loe. P
5 albicuspis, H.-S. (95),
Loc. P
r Gundlach, Plotz (99),
Porto Rico.
% clavicornis, H.-S. (100),
Loe. ?
» panthius, H.-S. (109),
Loe. ?
x rochus, Plotz (116),
Brazil.
i. scheba, Plotz (117), S.
Am.
7 chersts, H.-S. (118), Loc. ?
3 phhus, Plotz(124), Brazil.
Dr. F. D. Godman on American
Unknown tome. ? Genus Thymele.
d =Thymele enotrus, Cr. 9 =T.
Sulviluna, Mab.
= Thymele aulestes, Cr., var.
= Thymele aulestes, Cr., var.
A Goniurus, near talus, Cr. Not
in G. & 8. coll.
A Telegonus, very near T. eudemus,
Mab., from Central America.
AT. Rymele, very near 7’, enotrus,
Cr.
ASSL, hymele, dg, near 7. pervivar,
Hiibn.
A Chrysoplectrum, probably a var.
of otriades, Hew. From San
Paulo in the G. & S. coll.
A close ally of the variable 7. cre-
teus, Or.
= Telegonus creteus, Cr,, var.
Plotz’s figures appear to represent
two “species : the lower one
agrees with 7. jaira, Butl.,
the upper one being very
like 7. chiriquensis, ” Stand.
Herrich-Schafier’s name has
priority.
The identity of this species with
Goniloba malefida, H.-5., was
noted by Plotz.
= Telegonus elorus, Hew.
= Telegonus grullus, Mab. Herrich-
Schiiffer’s name has priority.
A Thymele(?), near phalecus,
G.&S.
A form of Proterdes idas, Cr., very
near Angasi, G. & S., but
without the subapical spots.
= Lpargyreus exadeus, Cr.
= Epargyreus asander, Hew., var.
=Epargyreus enispe, Hew., var.
with the subapical ” spots
wanting.
Very near LEpargyreus asander,
Hew., but with a straicht
Ww hite band on the secondaries
beneath.
= Zichydrus evelinda, Butl. Her-
rich - Schiffer’s. name has
priority.
Bone Cogia(?), near cajeta,
Hesperiidae described by Plotz.
Eudamus cajeta, H.-S. (128), Loe. ?
9 valeriana (valerius), Plotz
(150), Mexico.
3 nicomedes, Plotz (153),
Brazil.
» galapus, Plotz (184),
Mexico.
FS casica, H.-S. (135), Loe. ?
Proteides zethos, Plotz (137), Para.
a antiope, Plitz (138), Para.
as nicola, Plotz (140), Para.
Telemiades vulpecula, Plotz (145),
S. Am.
Telegonus epicalus, Hiibn. (arctu-
rus, H.-S.) (147),
Loe. ?
ceramina, H.-S. (148),
Loe. ?
)
Netrocoryne seneca, Plotz (160),
Brazil.
Eudamus porcius, Feld. (161), Rio
Negro.
Netrocoryne damias, Plotz (162),
S. Am.
Telegonus gaurus, Plotz (165), Rio
Negro.
fulvius, Plotz (166), Ca-
meta,
”
os probus, Moschl. (168),
Surinam.
- cenosa (H.-S.), Méschl.
(169), Venezuela.
a5 tychios, Plotz (179),
Bahia.
lucea, Plétz(183),S. Am.
135
Plotz’s figure does not quite agree
with the insect identified as
Cogia cajeta, H.-S., in the
‘ Biologia.’
Belongs to Cogia, near cajeta, H.-S.
Belongs to Telemiades, near me-
gallus, Mab.
A Rhabdoides, near casica, H.-S.
Her-
has
=Lhabdoides epigena, Butl.
rich-Schatler’s name
priority.
= Telemiades amphion, Hiibn., var.
= Telemiades amphion, Hibn., var.
Belongs to Cabares(?), near potrillo,
uc.
The type of the genus Physalea,
Mab. Males from Colombia
and Venezuela in G. & §. coll.
= Telemiades phasias, Hew. Hib-
ner’s name _ has __ priority.
Plotz’s figure was taken from
a Brazilian specimen.
Plotz’s figures represent the two
sexes of a Telemiades, from
Surinam, very near 7’. littera,
Mab. The supposed type of
T. ceramina, in the G. & 8.
coll., isa Q of Lerema accius,
Smith and Abbot, as stated in
the ‘ Biologia.’
Possibly belongs to Ardaris. Not
in the G. & S. coll.
= and 9, Dyscophus doriscus,
Hew., and Netrocoryne cecu-
tiens, H.-S. Felder’s name
has priority.
= Bungalotis ramusis, Cr., 3.
= Nascus euribates, Cr., 3, var.
= Telegonus erythras, Mab. There
is a specimen of it from Villa
Nova, Amazons, in the G. &58.
coll. Mabille refers the species
to Dyscophellus. Plotz’s name
has priority.
Genus? NotinG. & S$. coll.
= Bungalotis phaselis, Hew., ¢ @.
Plotz’s figures seem to represent
vars. of Bungalotis salatis,
Cro
Probably a var. of Nascus cephisus,
Hew., 2. A specimen of it
136
Pellicia macarius, H.-S. (191),
Venezuela.
» albangula, H.-S. (192),
Guatemala.
» ephora, H.-S. (193), 8. and
Centr. Am.
» licisca, Plotz (196), Nica-
ragua.
dimidiata, H.-S. (cormna,
Plétz) (199), Mexico.
» theon, Plétz (200), S. Am.
zamia, Plotz (201), 5. Am.
» tyana, Plotz (202),
Paulo.
» crispus, H.-S. (204), Ve-
nezuela,
Arteurotia demetrius, Plotz (205),
Brazil.
A‘thilla toxeus, Plotz (212), Mexico.
» nocera, Plétz (218), Co-
lombia.
primus, Plétz (214), Brazil.
melas, Plotz (216), Rio
Janeiro.
”
”
Cogia punctilia, Plotz (222), San-
tarem.
Cecropterus longipennis, Plotz (227),
S. Am.
55 koluthos, Plotz (228),
Colombia.
ss orontes, Plotz (229), La
Guayra.
+ lunulus, Plotz (281),
. Am.
bocus, Plotz (283), Para.
Lychnuchus ee) hiarbas, Cram.
(242), S. Am.
clearchus, Plotz (247),
S. Am.
9)
Dr. F. D. Godman on American
from Santarem in the G. & S
coll.
Pl6tz’s figures (¢ 2) do not agree
with the insect identitied under
this name in the ‘ Biologia,’
but belong to P. castolus, Hew.
The specimen (2 ) figured by Plotz
is stated to be from “ Rio,” not
Guatemala, The species can-
not be satisfactorily identified.
tiphys, Godm. __ Herrich-
Schaffer’s name has priority.
=P. thyestis, Godm., g. Plotz’s
naine has priority.
didia, Méschl. Herrich-
Schaffer’s name has priority.
Achlyodes nivonicus, Plotz, is
doubtless the female of the
same species.
Apparently not represented in the
G. & S. coll.
Apparently not represented in the
Wie
=P,
. coll,
Probably belongs to Mycteris, near
cer ‘ulead, Mab. Specimen
figured isa ¢.
Probably belongs to Mycteris, near
cambyses, Hew. Specimen
figured isa g.
Unknown to me.
=Murgaria albociliata, Mab., but
has darker cilia.
There is a worn specimen of this
species in the G, & 8. coll.
Also in G. & S. coll.
Very like 2. echina, Hew., but
with the secondaries almost
black beneath.
A close ally of C. calchas, H.-S.
=C. aunus, F., Gs
=C. zentus, Moschl., ¢. Occurs
also in Venezuela.
=, aunus, E., 9).
= C. neis, Hiibn., 9, var.
=C. ttylus, Hiibn., var.
Plotz figures a Lychnuchotdes
(P ozeus, Hew.) under this
name. It is possible that
Cramer’s species has been cor-
rectly identified by him.
This figure represents the insect
identified as Ancistrocampta
Hesperiides described by Plétz.
Sophista plinius, Plétz (249), 58. Am.
Hesperia paria, Plota (259), Chi-
riqul.
y ma, Plotz (261), Chiriqui.
» wmsignis, Plitz (262), La
Guayra.
» wmfuscata, Plotz (265),
Brazil.
Pamphila lurida, U.-S. (266),
Loc. P
Hesperia rubida, Plotz (268), S.
Am.
Cobalus derasa, H.-S,* (269), Loe. ?
Goniloba sandarac, H.-S. (271),
Cuba.
Hesperia depuncta, Plotz (278),
Rio Janeiro.
Cobalus tertianus, H.-S.
(274),
Loe. ?
Hesperia crispinus, Plotz (277),
Mexico.
5 rivera, Plotz (278), Rio
Janeiro.
5 bias, Plotz (279), 8S.
m.
137
hiarbas, Cram., in the British
Museum. Pl6tz’s figure (246)
of L. celsus, F., appears to
represent the same species.
Very near S. aristoteles, D. & H.
Fhere is a specimen of it from
Novo Friburgo in the G. & S.
coll.
=Lutychide achelous, Plotz, 3.
Specimen figured is from La
Guayra, not Chiriqui.
= Methionopsis modesta, Godm.
Plotz’s name ina has priority.
= Mnasitheus simplicissima, H.-S.
This is not the species identified as
Papias infuscata, Plotz, in the
‘ Biologia,’ for which Mabille’s
name wteger can be used.
HT. infuscata, Plotz, has a
brand formed of two narrow
elongate streaks on the pri-
maries in the ¢. It comes
very near Metiscus atheus,
Godm.,
Probably belongs to Mnasitheus.
Specimen figured is from
Brazil.
Probably belongs to Papias.
Probably the 2 of a species of
Papeas.
= Hesperia palea, Hew., Herrich-
Schaffer’s name having pri-
ority. Belongs to genus As-
bolis, Mab.
= Cobalus cinnamomea, H.-S., the
latter name having priority.
Belongs to a genus near Dion.
Numerous specimensin G, &S.
coll.
Belongs to Cobalus. Pamphila
warra, Moschl., is the ¢, and
P. zola, Moschl., the 2, of the
same species,
Looks very like Mnasicles geta,
Godm. The drawing of the
underside is, perhaps, too
highly coloured.
Probably a Mnasicles,
Very like Mnaseas bicolor, Mab.
* Corresp.-Blatt Regensb. 1870, p. 159.
138
Tesperva gabinus, Plétz (280), Rio
Janeiro.
», Herminieri, Latr. (281),
Carolina.
» perloides, Plotz
Brazil.
perla, Plotz (283), Rio
Janeiro.
» ewcellata,
Brazil.
Pamphila subcostulata, H.-S. (286),
Brazil.
(282),
Plitz (284),
Hesperia leucopogon, Plotz (287),
La Guayra.
antistia, Plotz (290), Rio
” ?
Janeiro,
lycenoides, Plotz (291),
Rio Janeiro,
pruinosa, Plotz (293), 8.
Am.
Plitz (294),
abdon,
Brazil.
acrea, Plétz (296), Co-
lombia.
aon, Plotz (800), Co-
lombia.
Jinbriata, Plotz (801),
Mexico.
ochrope, (311),
Para.
Plotz
Erycides apicalis, H.-S. (818), Loe. ?
Hesperia peratha, Plotz (815),
Bahia.
» _ nealces, Plotz (816), Rio
Janeiro.
alda, Plotz (321), Brazil.
”
Dr. F. D. Godman on American
Genus? Two @Q from Santa
Catherina in the G. & S.
coll.
Plétz figures a Venezuelan speci-
men as this species.
Very like the Mexican Mastor
anubis, Godm.
Not represented in the G. & S. coll.
Apparently not represented in the
G.& 8. coll.
Probably belongs to Papias. A ¢
from Iquitos in the G. & S.
coll. may belong to it.
Apparently not represented in the
G. & 8. coll.
A Cymenes and probably a form of
the species identified as C.
stlius, Latr., in the ‘ Biologia,’
Plotz’s figure of the latter
represents a larger insect, very
like an unnamed specimen
from Chiriqui in the G. & S.
coll.
A Cymenes, and perhaps a var. of
malitiosa, H.-S.
Belongs near the genus Dion.
There are two specimens of it
from Chapada in the G. & S.
coll.
Belongs to the genus Zenida, Mab.
There are three specimens of
it in the G. & 8. coll.
Unknown to me.
=Lignyostola lacydus, _ Druce.
Plotz’s description does not
seem to have been published.
Unknown to me.
=Pyrrhopygopsis telmela, Hew.,
var. with a small hyaline spot
on the primaries exterior to
the transverse fascia. Plétz’s
figure (310) of P. telmela is
very like P. cleanthes, Latr.
= Pyrrhopygopsis cleanthes, Latr.,
var. with the spots on the pri-
maries very small.
Probably a TZhracides, near luda,
Hew.
= Thracides luda, Hew., and T.
hundurensis, Mab.
A very close ally of T’hemesion cer-
tema, Hew. There is a 9 of
it from Santa Catherina in the
G. & S. coll.
Hesperiidae described by Plotz.
H.-S. (822),
Cobalus quadrata,
Loe, ?
Hesperia trimaculata, Plotz (826),
Brazil.
» elisa, Plotz (832), Brazil.
5 adjuncta, Plotz (340),
Colombia.
» Besckei, Plotz (348),
Novo Friburgo.
» xanthotrix, Plétz (852),
Rio Janeiro.
» brinoides, Moschl. (860),
Surinam.
» nanneta, Plotz (870), Rio
Janeiro,
catochra (catochia), Plotz
(3874), Mahrida.
Goniloba matthiolus, H.-S. (375),
Loe. ?
Hesperia pelora, Plotz (884),
Brazil.
valentina, Plitz (889),
Surinam.
¥ replana, Plotz (890),
Brazil.
. socles, Plétz (892), 8.
Am.
. emacareus, Plotz (393),
Venezuela,
Goniloba macareus, H.-S. (395),
Loc. ?
Hesperia eas, Plotz (400), Chiriqui.
» eabenta, Plotz (401), 8.
Am.
139
Belongs to genus Tisizs. There is
a ¢o of it from Minas Geraes
in the G. & S. coll. exactly
agreeing with Plotz’s figure.
Probably a Cobalus near gabina,
Godm. There is a ¢ of it
from Minas Geraes in the
G. & S. coll.
=H. crotona, Hew. Genus near
Perichares.
= Talides sergestus, Cr.. g, dark
var.
= Mcontades cydia, Hew., var., 3,
with a broader white band on
the secondaries beneath.
Probably a var. of Paraides or-
chamus, Cr., ¢. The Phocides
canthothrix of Mabille, from
Bolivia, is perhaps a different
species, Plétz’s description
has not apparently been pub-
lished.
=H. noseda, Hew. Genus?
Probably belongs to Thraeides,
near c@sena, Hew.
Incorrectly quoted amongst the
unidentitied Mexican species
in the ‘ Biologia.’ No doubt
an eastern form,
= Thracides salius, Cram., var.
Plotz’s figure is too green on
the upperside.
Probably not the 9 of Cobalopsis
edda, Mab., as suggested in the
‘ Biologia.’
= Carystoides basochesi, Latr., var.
with the three hyaline spots on
the primaries reduced in size
and the patch on the second-
aries broken up into small
spots.
=Carystoides basochesi, Latr., var.
with the hyaline patch on the
secondaries reduced to two
small spots.
= Vacerra litana, Hew., var.
The insect identified as Thespicus
macareus, H.-8., in the ‘ Bio-
logia.’
= Thespieus othna, Butl.
= Vacerra htana, Hew., var.
Probably a var. of Vacerra litana,
Hew., with the white markings
more extended beneath,
140
Hesperia diores, Plotz (407), 8. Am.
elana, Plotz (409), Brazil.
» stlanion, Plotz (410),
Bahia.
5 mitella, Plotz (411),
Brazil.
Jebus, Plotz (412), Brazil.
melaleuca, Plotz (413),
Rio Janeiro.
Cobalus dissoluta, Plotz (416),
Loe. ?
Hesperia ozeta, Plo6tz (417), Bahia.
ciniea, Plotz (418), Para.
”
”
7
Goniloba caniola, H.-S. (419), Loe. ?
» conformis, H.-S. (422),
Loe. ?
» luctuosa, H.-S. (426),
Loe. ?
Hesperia chlorus, Plotz (428), Suri-
nam.
» xsa, Plotz (430), Rio
Janeiro.
hypodesma, Plétz (481),
Para, Rio Janeiro.
cuneata, Plétz (482), 8
Am.
hesiodes, Plotz (434),
“Cap Rico.”
elvira, Plotz (436), S.
Am.
lyrcea, Plotz (442), Brazil.
eucherus, Plotz (445), Su-
rinam,
», monacha, Plotz (451),
Blumenau.
peninsularis, Plotz (452),
Para
senen, Platz (459), Rio
Janeiro.
Dr. F. D. Godman on American
Very like Carystus Claudianus,
Latr., but with the seeondaries
similarly marked above and
beneath.
Comes near the genus Pseudo-
sarhia, Berg. There are three
males of it from Chapada in
the G. & S. coll.
Very like the 2 of Carystus hy-
laspes, Cram., the type of Ma-
bille’s genus Synale. The
single specimen (Q) in the
G. & §. coll. has the head
green above.
A very close ally of H. elana,
Piitz, and belonging to the
same genus. Occurs at the
same locality, Chapada, from
which place there is a pair in
the G. & 8, coll.
= Zenis minos, Latr.
= Zenis minos, Latr., var.
= Zenis calvina, Hew.
= Zenis ozota, Butl., var.
Probably a Tirynthia, near conflua,
= Vacerra canente, Butl. Herrich-
Schaffer’s name has priority.
Probably a var. of Prenes nero, F.
Belongs to Prenes, near evadnes,
Cram., and pauper, Mab.
= Prenes evadnes, Cram., 3.
Rr obably: an Oxvynthes, near corusca,
H.-S.
A close ally of Metron chrysogaster,
Butl.
= Metron chrysogaster, Butl., and
goza, Hew
= Cobalus fidicula, Hew. The lo-
cality quoted is a misprint for
Costa Rica.
Probably a Vettius, near laur a
Hew.
= Carystus marcus, F.
= Carystus fantasos, Cram.
Belongs to Vettius, near triangu-
laris, Hiibn. Plotz’s figure is
taken from a Para specimen.
Probably a Vettéus, near Lafres-
nayt, Latr.
= Paracarystus hypargyra, H.-S,
var.
Hesperiidee described by Plitz.
Hesperia rexia, Plétz (466), Brazil.
; aquilina, Plotz (473),
Loe. ?
Cobalus catocala, H.-S. (485), Loe. ?
Hesperia distigma, Plotz (488),
Loe. ?
” sabina, Plotz (490),
Novo Friburgo.
hersilia, Plota (492), Rio
Janeiro,
Cobalus bistrigula, H.-S.
Loe. ? 2
Hesperia cyrus, Plotz (498), Rio
Janeiro.
9 parvipuncta, Moschl.
(500), Surinam.
Goniloba hemeterius, H.-S. (501),
Loe. ?
(497),
Carystus erebina, Moschl. (502),
Colombia.
Cobalus sameda, H.-S. (503), Loe. ?
Hesperia degener, Plotz (504),
Loe. ?
_ lydora, Plotz (505), Ve-
nezuela,
Cobalus grossula, H.-S. (507),
Loe. ?
rH deleta, H.-S. (508),
Loe. Pp
Hesperia duroca, Plétz (509), Rio
Janeiro.
Goniloba aphilos, H.-S. (511), Loe, ?
141
Genus? Probably a var. of Hes-
perta almoda, Hew., with the
lighter markings of the under-
side wanting. Carystus meta-
nira, Mab., is no doubt synony-
mous.
Perhaps an Artines, near epitus,
Hiibn.
Plotz’s drawing of this species
shows that it is not the 2 of
Megistias tsus, Godm.
Very like Hutychide cingulicornis,
H.-S., and probably Ameri-
can.
Unknown to me. Genus? The
¢ has a brand formed of three
short streaks,
= Cobalus virbius, Cram.
=Rhinthon alus, Mab. Herrich-
Schaffer’s name has priority.
Near Hesperia obeda, Butl. Genus?
=Pamphila Méschl.
Genus P
There is a pair of this species from
Cuba in the G. & 8. coll.; the
3, however, wants the two
small hyaline spots on the
primaries shown by Plotz.
Genus ?
Plotz figures the ¢ of a different
species from Brazil under this
name. C, erebina, Moschl.,=
Hesperia cynea, Hew.,as stated
in the ‘Biologia,’ under the
genus Rhinthon.
This may be the 9 of Plét2’s C.
erebina, which cannot be in-
cluded in the genus Rhinthon.
Herrich-Schiffer’s name has
to be used in any case,
Belongs near Gonus. A ¢ from
Bolivia in G. & 8. coll. is very
like it.
Belongs near Gonus. Two males
from Venezuela in the G.& 8.
coll.
Very like Oligoria maculata, Edw.
Unknown tome. ? American,
= Oligoria maculata, Edw.
angularis,
Perhaps the 2 of H. cyrus, Plotz.
=Carystus obeda, Butl., ¢. Ge-
nus? Herrich-Schatter’s name
has priority,
142
Cobalus elegantula, H.-S. (518),
Loe. ?
Hesperia quadrangula, Plotz (514),
Brazil.
Cobalus subcordata, H.-S. (515),
Loe. ?
Hesperia olympia, Plotz (516),
Brazil.
Ps norus, Plétz (518), New
Orleans.
Pamphila theogenis, Capr.* (519),
Surinam.
Cobalus dama, H.-S. (522), Loe. P
Hesperia cornelius, Latr. (525),
Cuba.
Cobalus tripunctus, H.-S. (526),
Cuba.
Hesperia gura, Plétz (528), Chi-
riqui.
Goniloba complana, H.-S. (529),
Loc. ?
corope, H.-S. (530),
Loe. ?
Hesperia eteocla, Plotz (581), Rio
Janeiro,
Carystus orope (Plétz), Capronn.
(533), Botafogo, Brazil.
Cohalus nerotdes, H.-S. (554), Loc. ?
”
Hesperia dalima, Plotz (688),
Brazil.
dispersa, Plotz (541),
Cuba.
» philerope, Plotz (542),
razil.
i; credula, Plotz (5851),
Brazil.
Goniloba cubana, H.-S. (556), Cuba.
Hesperia osca, Plétz (557), Caracas.
sabea, Plotz (560), Brazil.
”?
ulrica, Plétz (561), Rio
” i
Janeiro.
Dr. F. D. Godman on American
Genus? Several specimens of it
from Brazil in the G. & S.
coll.
A close ally of C. elegantula, H.-S.
Genus? Also contained in
the G. & S. coll.
Genus? Occurs in Guiana and
Brazil.
= Cobalus subcordata, H.-S., var.
= Oligoria maculata, Edw., var.
Not identified in the G. & S. coll.
Not identified in the G. & §. coll.
@ looks like Prenes vada,
Mab.
Plotz’s figure represents Prenes
ocola, Edw., and his deter-
mination is, parhaps, correct.
Near Megistias. Occurs also in
S. America. Pamphila ancus,
Méschl., is the same species,
= Eutychide midia, Hew., and Hes-
peria favetta, Plotz.
= Lutychide midia, Hew. Plitz’s
drawing is taken from a Nica-
raguan specimen. Herrich-
Schiatfer’s name has priority.
= 9, Damas clavus, Er.
Probably a Codalus, allied to ga-
bina, Godm.
Unknown to me,
Not identified in the G. & S. coll.
? American.
Probably belongs to Vacerra.
There is a ¢ very like it in
the G. & S. coll.
=Lerodea eufala, Edw. Descrip-
tion unpublished ?
Not identified in the G. & S. coll.
Not identified in the G. & S. coll.
Very like Hesperia quadrangula,
Plotz, 2. Genus?
Not identified in the G. & S. coll.
A ¢ from Colombia in the G. & S.
coll. very like this, but with
small hyaline spots.
Not identified in the G. & S. coll.
* Figured by Capronnier in Ann, Soc. Ent. Belg. xvii. t. i. fig. 8.
Hesperiidee described by Plotz.
Fesperia xanthosticta, Plotz (562),
Para.
Goniloba complanula, H.-S. (564),
Loe. ?
Hesperia yema, Plotz (565), West
Indies.
is verticalis, Plotz (567),
Brazil.
o commodus, Plotz (568),
Brazil.
ef dyma, . Plétz (569),
Brazil.
99 dedecora, Plotz (571),
La Guayra.
Goniloba exoteria, H.-S. (572),
Loe. P
Hesperia Grote, Plotz
United States.
. lochius, Plotz (576), La
Guayra.
A dido, Plotz (577), Vene-
zuela,
F phocylides, Plotz (578),
La Guayra.
p edata, Plotz (580), La
Guayra.
= tyrteus, Plétz (581), La
Guayra.
5) judas, Plitz (584), Novo
Friburgo, Brazil.
(573),
5 wimico, Plétz (585),
Florida.
53 pudorina, Plotz (587),
Brazil.
4 lupulina, Plotz (588),
Loc. ?
3 lunata, (589),
Loe. ?
stlene, Plétz (590), Loe. ?
oA corticea, Plotz (591), La
Guayra.
Plotz
nt vetulina, Plotz (597),
Loc. ?
143
Not identified in the G. & S. coll.
=Turesis Lucasi, F. Plotz’s
drawing is taken from a Chi-
riqui specimen,
=Turesis Lucast, F. Carystus
hebon, Mab., and Pamphila
stlacea, Moschl., are probably
varieties of the same specier.
Description unpublished ?
Genus? A female from Rio
Janeiro in the G. & BS. coll.
may belong to this species,
but it has only one outer spot
on the primaries instead of
two.
Numerous specimens (¢ ?) in the
G. & 8. coll. Genus ?
Genus? A pair in the G. & S.
coll.
Not identified in the G. & S. coll.
= Amblyscirtes nanno, Edw. Her-
rich-Schaffer’s name has
priority.
= Atrytonopsis hianna, Scudd., 3,
var.
= Lerema bipunctata, Mab. Plitz’s
name has priority.
Probably Lerema accius, 8. & A.,
Waits, o:
Probably a small Q of Lerema
aceius, S. & A.
P=Megistias sus, Godm, 9.
Plétz’s name has priority.
= Megistias telata, H.-S., var.
Unknown to me.
Pnear genus
Atrytonopsis.
=Prenes panoquin, Seudd. De-
scription unpublished ?
Belongs near Catia. Numerous
specimens (¢ 2 )intheG. &S8.
coll.
= Atrytone inimica, Butl. & Druce.
=Cymenes berus, Mab. Plitz’s
name has priority.
= Cymenes berus, Mab., var.
= Megistias epberus, Mab. Hes-
peria noctis and lysias, Plotz,
are doubtless varieties of the
same species. Plétz’s name
corticea has priority over epi-
berus, Mab.
= Euphyes verna, Edw., ¢.
144
Hesperia lidia, Plitz (607), Loc.?
» eulogius, Plotz (608),
Mexico.
» vesuria, Plotz (612), Ja-
maica.
» gemma, Plitz (618),
Loe. ?
» myrona, Pléitz (614),
Venezuela.
» floridensis, Plitz (616),
Florida.
*) PELTED Plotz (617),
Loe. ?
» lysias, Plotz (623), Chi-
riqul.
» mutius, Plotz (625),
Georgia.
53 Huebneri, Plotz (626),
West Indies.
Pamphila magdatia, H.-S. (681),
Cuba.
Hesperia magica, Plitz (682), Cuba.
Plitz (633),
ammonia,
Loe. P
radians, Lue. (684), Cuba.
’”
clara, Plétz (635), Cali-
fornia.
Hy combinata, Plotz (636),
Colombia.
55 unna, Plotz (637), Phila-
delphia.
enuna, Plétz (639), Chile.
ancora, Plitz (641), Loe. ?
ignorans, Plotz (647),
Loe. ?
dares, Plotz (648), Loe. ?
Dr. F. D. Godman on American
Probably a var. of Limochores
manataaqua, Seudd,
= Atrytone mellona,GGodm. Plotz’s
name has priority.
A Catia, near Druryi, Latr.
in G. & 8. coll.
= Pamphila ravola, 8. & G., oS,
from Dominica, as already
noted by Weymer and Mabille.
Belongs near the genus Catia.
Plotz’s name has priority.
= Pamphila kenava, Butl., d. Be-
longs near Molo.
=Limochores palatka, Edw., 3.
Also
Probably a var. of the N.-American
Erynnis ottoe, Edw., 3.
=Megistias epiberus, Mab., and
Hesperia corticea, _— Plotz.
Plétz’s names antedate that of
Mabille.
= Atrytone arogos, Boisd., var., 3.
= Atrytone vitellius, F. Descrip-
tion unpublished ?
This Cuban insect has stood for
many years under the name
Thymelicus nanus, H.-8., in
the G. & 8. coll, This latter
species, according to Plotz’s
figure of it, isan Ancyloaypha.
There is a specimen of 7. na-
nus in Mr. Druce’s collection.
= Choranthus radians, Luc., vayr.,
dQ.
Perhaps a dark form of the 2 of
Choranthus radians, Lue.
This figure, if really belonging to
H, radians, Q, represents a
dark, almost immaculate form.
Apparently a pale var. of Thyme-
licus brettus, Boisd., 3.
=Thymelicus viber, Hiibn. rm Vein
Description unpublished ?
= Thymelicus brettus, Boisd., 3.
Blanch.,
=Hylephila fasciolata,
unpub-
@. Description
lished ?
= Hylephila phyleus, Drury, 3 9.
Probably belongs to Pol.tes. There
isa 9 very like it, from Me-
rida, Venezuela, in the G. & 8.
coll.
Belongs to Thymelhcus, There are
several specimens (d¢ 2) of
this species, from Brazil and
Bolivia, in the G. & 8. coll.
Hesperiidee described by Plétz.
Hesperia lina, Plitz (649), Bogota.
5) reticulata (650), la
Guayra and Chiriqui.
zacheus, Plitz (652), Su-
rinam.
”?
Cobalus vitellina, H.-S. (653), Loe. ?
Hesperia erratica, Plitz (656),
Guatemala.
subreticulata,Plitz (658),
Loe. ?
ob
francisca, Plitz (666),
California and Mexico.
Hf amphissa, VPlotz (667),
Venezuela.
grynea, Plotz (670),
Chile.
5 augustus, Plotz (676),
Brazil.
6 angulina, Plotaz (677),
Brazil.
Pamphila fasciata, Moschl. (678),
Surinam.
Hesperia ulphila, Plotz (679),
Mexico,
Pamphila lamida, Moschl. (680),
Colombia.
Hesperia statius, Plétz (686), Vene-
zuela.
Pamphila antiqua,
Cuba.
Hesperia serina,
Mexico.
H.-S. (687),
Plitz (697),
145
Probably belongs to Polites. There
isa @ of it from Colombia in
the G. & S. coll.
=Famphila_suffenas and meton,
Mab, Plotz’s name has pri-
ority. The species is referred
to Phlebodes by Mabille. The
four specimens figured by Plitz
seem to represent more than
one species.
= Atrytone melane, Edw., var., 3,
with the secondaries immacu-
late above.
=Atrytone melane, Edw. Plitz’s
figures (¢ 2) were made from
Mexican specimens. The two
names were published in the
same year, 1869.
= Altrytone zabulon, Boisd., ¢.
Plotz’s figures (655) of A.
zabulon, from Buffalo, represent
A, hobomok, Harr.
=Cherephon rhesus, Edw., 9.
Plotz’s drawing was made
from a Mexican specimen,
= Ochlodes agricola, Boisd., 3 9.
The Mexican habitat requires
confirmation.
= Atalopedes campestris, Boisd.,
do.
=Hylephila fulva, Blanch., 2.
Probably belongs to <Atrytone.
There is a similar unnamed
form from Brazil in the G. & S.
coll.
= Hesperia oropa, Hew., 5. Be-
longs near Metron.
= Hesperia oropa, Hew. Q@.
Wrongly identified in the
‘ Biologia’ as the 2 of Metron
chrysogaster, Butl. Méschler’s
name has priority.
An Aftrytone, near monticola,
Gedm., with larger spots on
the primaries and the under-
side of the secondaries differ-
ently marked. ‘The figures
seem to represent two species.
Probably the 2 of a species of
Airytone. Unknown to me.
=Serdis venexuele, Westw., and
S. fulgens, Mab., 5 ?.
= Phemiadesutha, Hew. Hervich-
Schafter’s name has priority.
An Eastern species of Padraona,
not Mexican.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 10
146
Hesperia tropica, Plotz
Mexico.
(698),
im evilis, Plitz (706), Cali-
fornia.
os letis, Plétz (710), Rio
Janeiro.
Apaustus argynnis, Plotz (758),
Brazil.
; odilia, Plitz (784),
Brazil.
Fj acroleuca, Plétz (748),
Brazil.
5 interpunctata, Plotz
(753), Bahia and Para.
Papilio saturnus, F. (755), 8. Am.
Apaustus levina, Plotz (760), Rio
Janeiro.
» jflavocostata, Plotz (761),
Rio Janeiro.
» ferrago, Plétz (762),
Loe. ?
. eudesmia, Ploitz (763),
Mexico.
Ke vicinus, Plotz (764),
Loe. P
5 imerius, Plotz (765),
Brazil.
> Prittwits, Plotz (771),
Mexico.
* euphrasia, Plitz (778),
Mexico.
Goniloba singularis, H.-S. (776),
Cuba.
Thymelicus isidorus, Plotz (780),
Mexico.
nanus, H.-S. (781),
Cuba.
Butleria mesoxantha, Plétz (815),
Venezuela.
» vantholeuca, Plitz (816),
Venezuela.
» lagilla, Plotz (820), Loc. ?
» apertus, Plotz (824),
Loe. ?
Dr. F. D. Godman on American
An Eastern species of Padraona.
Java? is pencilled on the
drawing.
Probably the @ of a species of
Vinius, near arignote, Godm.
Locality doubtless incorrect.
= Vinius nicomedes, Mab., 3. Both
names were published in 1883.
Unknown to me. Genus?
Near Phlebodes unia, Butl., from
Haiti. Genus?
Near genus Artines. NotinG.&8.
coll.
= Callimormus vetula, Mab.
This figure, taken from a Para
specimen, seems to represent
Vorates decora, H.-S.
Genus? Unknown to me.
Genus? An allied form from
Guiana in the G. & S. coll.
= Padraona epictetus, F., var.
= Padraona epictetus, F., var.
Genus? An allied form from Co-
lombia in the G. & 8. coll.
Genus? From the “ Kaden ” col-
lection in the G. & 8. coll.
= Adopeoides simplex, Godm. (nec
Feld.). Felder’s species, to
judge from Plotz’s figure (775),
appears to be a species of
Zariaspes unknown to me.
= Ancyloxypha arene, Edw., @,
dark var. A. leporina, Plotz,
doubtless belongs to the same
species.
= Copeodes aurantiaca, Hew., 3.
Plétz’s figure is taken from a
Mexican specimen. Herrich-
Schiiffer’s name has priority.
= Oarisma poweshetk, Park., var.
An Ancyloxypha, near arene, Edw.
= B. cypselus, Feld., var., a species
included by Mabille in his
genus Dalla,
= B. dimidiata, Feld.,var., referred
by Mabille to his genus
Dalla.
Very near B. caicus, Hew.
Near the Chilian B. bisserguttata,
Philippi.
Hesperiidee described by Plétz.
Butleria pruna, Plotz (828), San
Domingo,
Carterocephalus flavimargo, Plotz
(829), Chile.
Cyclopides gyrans, Plotz (843),
Mexico.
3 vitus, Plotz (848),
Chile.
Leucochitonea ligania, Plotz (862),
S. Am.
* eulalia, Plotz (865),
Venezuela.
Pyrgus aconyta(H.-S.), Plotz (871),
Georgia.
Syrichthus valdivianus,
(875), Chile. Rees
Pyrgus aqutrix, Plots (882), Bu-
enos Aires.
yy wnsolacna, \Plotz (887),
Mexico.
3) albescens, Plotz’ (889),
Mexico.
» varus, Plotz (900), Mexico.
Sey cungus, ue lowe (QO),
Centr. America.
Carcharodus radiatus, Plotz (907),
Texas.
Eiphyriades variegata, Plotz (911),
Rio Janeiro.
Pythonides dilucida, Moschl. (916),
Surinam.
* ceruleus, Plotz (930),
Brazil.
Hesperia tertulanus, IF. (935),
Loe, P
Pythonides alaricus, Plotz (988),
Bahia.
Syrichthus leucodesma, Hr. (939),
Guiana.
Pythondes servatius, Plotz (944),
Para.
fs prudens, Plotz (954),
Surinam.
147
There is an allied form from Rie
Janeiro, with a similarly
coloured underside, in the
G. &S. coll.
= Argopteron aurerpenne, Blanch.,
OF
= Dalla (Butler va) anomala, Mab.
Plotz’s name has priority.
Probably belongs to Batleria, near
tr ipunctata, } Mab.
= Heliopetes alana, Reak., var.
=Heliopetes nivella, Mab. Plotz
figures (864) a very similar
form, almost oy white
above, from Cayenne, as &.
maimon, fF. Deésc Cancer un-
published ?
Very like Helopetes domicella, Kv.,
but greener. Description un-
published P
A. Flesperia, very near H. trisig-
nita, Mab. and HH. notata,
Blanch.
?= Hesperia montwaga, Reak.
Plotz’s figures were taken
from Mexican specimens.
? = Hesperia notata, Blanch.
?= Hesperva montivaga, Reak.
2 = Iesperia montivaga, Reak,
?= Hesperia notata, Blanch.
= Celotes nessus, Edw.
Belongs to Dieus, near lacena,
Hew.
Unknown to me.
=Paches lmea, Hew.
Butl.).
Plotz figures a 2 specimen, from
Minas Geraes, of Leucochitonea
lancea, Hew., as this species.
Genus?
A Paramimus, near scurra, Hiibn.
Not represented in the G, & S.
coll.
= Paramimus scurra, Hiibn., var.,
without the red spot.
An Atarnes, near Sallei, Feld.
(iabesa,
Probably belongs to Achlyodes.
From Cayenne and Chapada
inG. & 8. coll.
10*
148
Achlyodes nivonicus, Plitz (956),
Mexico.
thiena, Plétz (957),
Loe. ?
a serapton, Plotz (959),
Novo Friburgo.
Hi plumbago, Plotz (960),
Loe. ?
x protius, Plotz (961),
Brazil.
‘5 gorgona, Plotz (963),
Gnatemala.
9% basiguita, Ploitz (964),
S. Am.
7 noctula, Plotz (965),
Para,
Hesperia bigutta, Prittw. (977),
Loe. ?
Achlyodes blanda, Plotz (980),
Loe. P
5) servius, Plotz (981),
Brazil.
% erisichthon, Plotz (982),
Loe. ?
% cnidus, Plotz (983),
Loe.?
“ balma (H.-S.), Plotz
(984), Brazil.
Antigonus sericus, Plotz (990),
Chiriqui.
5 hborws (H.-8.), Plotz
(991), Bahia.
% bypuncta, Plotz (995),
Mexico.
aura, Plotz (998),
Brazil.
is badia, Plotz (999),
Chanchamayo.
£ robigus, Plotz (1006),
S. Am.
Achlyodes auxo, Méschl. (1008),
Colombia.
Antigonus patens, Plotz (1009), Rio
Janeiro.
% tortricinus, Ploéta (1010),
Panama and Vene-
zuela.
Dr. F. D. Godman on American
Apparently = Pellicia dinuidiata,
H-S., Figured from a
damaged specimen.
An Achlyodes, near Fredericus,
Hitbn. From British Guiana
and Para in the G..& S. coll.
Not identified in the G. & S. coll.
= Achlyodes chalybs, Mab. Plotz’s
name has priority. From
Para and Ega in the G. & 8.
coll,
= Achlyodes pulverea, Mab. From
Rio Janeiro in the G. & S.
coll,
= Chiomara gesta, H.-S., var.
= Chiomara punctum, Mab.
= Chiomara nuthrax, Moschl.
= Cnimara gesta, H.-S., var.
= Chiismara sssta, H.-S., var.
= Pythonides hyacinthinus, Mab.
A Pythonides, near phila, G. & S.
A Pythonides, unknown to me.
= Pythonides zera, Butl.
tion unpublished P
= Systasea corrosa, Mab.
Descrip-
A Systasea, near corrosa, Mab.
From Chapada in G. & S. coll.
Description unpublished ?
= Theagenes lactifera, Butl., 2, var.
Plotz’s figures (994) of T,
noctud, Feld., § 9, agree with
T. lactifera, Buti.
= Diphoridas dichrous, Mab. This
species may be the Hesperia
palpalis of Latreille. |
= Kbrietas infanda, Butl.
=Echelatus luctuosus, G. & S.
Plotz’s name has priority.
= Camptopleura theramenes, Mab. °
Unknown to me. Perhaps a
species of Gorgophone.
3 = Hbrietas anacreon, Staud. ;
2 =. infanda, Butl., or ~
EE, tsus, Mab.
Hesperiidee described by Plétz.
Antigonus elaudia (H.-S.), Plotz
(1012), La Guayra.
trisertata (H.-S.), Plotz
?
(1013), Venezuela.
of eremita, Plotz (1014),
S. Am,
3 simplicior, Plétz (1015),
Brazil.
re alburnea, Ploétz (1016),
Para.
. cajus, Plétz (1023), Peru.
a adamas, Plétz (1051),
Brazil.
Thanaos zarucco, Lue. (1035),
Cuba.
Antigonus andere Plotz (1041),
uba.
= heteropterus, Plotz
(1044), Brazil.
Jamaicensis, Plotz
(1045), Jamaica,
Nisoniades flavipalpis, Plotz (1051),
Copiapo.
eusebtus, Plotz (1055),
Centr. America.
Achlyodes trochilus, Hoptt. (1055),
”
Peru.
Hesperia chlorocephala, _ Laty.
(1056), Brazil. — i
Nisoniades auricapilla (Hopff.),
Plotz (1057), Para,
norica, Plotz (1059),
Brazil and Cayenne.
eta, Plitz (1060), Chi-
riqui and Brazil.
aterea, Ploitz (1062),
Rio Janeiro.
Tagiades teniatus, Plota (1068),
Oaxaca.
doria, Plotz
Mexico.
”
(1075),
”
149
An Ebrietas, near ecliptica, Butl.
Description unpublished ?
An Lbrietas, probably a var. of
E. elaudia.
? Belongs to Ebrietas.
to me.
Belongs to Echelatus. Typeisa od.
Unknown
An Echelatus, and probably the @
of E. simplicior. There is a
specimen of it from Paraguay
in the G. & §. coll.
Belongs to Eudamidas. There are
three specimens of it from
Peru in the G. & S. coll.
3 =Sostrata leucorrhoa, G. & S.;
2 =S. scintillans, Mab.
= Thanaos martialis, Seudd. Lu-
cas’s name has priority,
Belongs to Thanaos. Also from
Cuba in the G. & 8. coll.
Belongs to Thanaos. There is a
long series of this species from
Chapada in the G. & §. coll.
Belongs to the genus Melanthes,
Mab. Very near the variable
M., zephodes, Hiibn. (figs. 1037,
1038 of Piétz), from Cuba and
the Bahamas. The ¢ only
of the Jamaican insect is
figured by Plotz.
Not identified in the G. & 8. coll.
Genus Staphylus? Figure
taken froma ¢.
= Staphylus giselus, Mab.
Belongs to Gorgopas. This insect
isidentitied as Hesperia chloro-
cephala, Latr., in the G, & 8.
coll.
Plotz’s figure of this species repre-
sents a Staphylus.
Genus? Not Staphylus auro-
capilla, Staud. Description
unpublished ?
Genus? Not identified
G. as. coll:
A Staphylus, 3. ?=mazans, Reak.
The figure is taken from a
3razilian specimen,
= Systasea incisa, Mab., var.
in the
= Systasea pulverulenta, Veld., var.
A. Timochreon, and probably a var.
of satyrus, Feld.
150
Tagiades monophthalma, Plotz
(1077), Brazil.
% diophthalma, Plotz
(1078), Loe. ?
» jacobus, Plotz (1082),
Rio Janeiro.
5 morvus, Plotz (1088),
Brazil.
s} hiera, Plotz (41103),
Loe. ?
Phareas ferrugineus (1121), Bahia.
» cervinus, Plotz (1122),
Loe. ? ur
» epimethea — (epiminthea),
Plétz (1123), Brazil.
» Anne, Plitz (1135), Para.
Entheus concinna, Plétz (1142),
Para.
Erycides imbreus, Plotaz (1198),
Centr. America.
Eudamus batabano, Wue. (man-
cinus, H.-S.) (1200), Cuba.
Erycides erebus, Plitz (1201),
Bahia.
. Herrichi, H.-S. (1204),
Loc. P
59 lincea, H.-S. (1209),
Loe. ?
Pyrrhopyge Marti, Plotz (1237),
Brazil.
Myscelus epigona, H.-S, (1248),
Loe. ?
Pyrrhopyge _ deniiculata, H.-S.
(1289), Loe. ?
Goniurus Platowti, Plotz (1817),
Toe.
brevicauda, Plétz (1819),
Chiriqui.
dominicus, Plotz (1821),
Loe.?
gideon, Plotz
Loe. ?
(1824),
Dr. F. D. Godman on American
Belongs to Cyclosemia. From
Parana in the G. & 8. coll.
A Cyclosemia, near earina, Hew.
Belongs to Cyclosemia.
Belongs to Cyclosemia. <A speci-
men very like it from Entre
Rios in the G. & S. coll.
= Celenorrhinus eligius, Cram.
Probably a Lignyostola, near de-
specta, Butl. Unknown to me.
=Eudamus pausias, Uew., var.
Genus ?
Belongs to Lignyostole.
= Entheus lemna, Butl., 2.
Unknown to me.
A Phocides, near alemon, Cram.
Unknown to me.
Belongs to Phocides.
Probably a Tarsoctenus. Unknown
to me.
A Tarsoctenus, near plutia, Hew.
Very like Naseus phocus, Cram., 3.
The drawing is marked = £.
grandimacula, Mab., a Bra-
ailian insect. Herrich-Schaf-
fer’s name has priority.
Probably belongs to Sarbia,
This species was wrongly identified
in the ‘ Biologia’ as synony-
mous with 2habdotdes epigena,
Butl., and the reference to it
in that work must be erased.
M. orbius, Mab., is very nearly
related to M. epigona, H.-S.
Ploétz’s figures are taken from
Venezuelan specimens,
A Mysoria, near pelota, Plotz.
Plétz’s drawing was made
from a Rio Negro specimen.
Probably a var. of Eudamus esme-
yaldus, Butl., with the dark
bands on the underside of the
secondaries coalescent.
= Thymele eniopeus,G.& 8. Plotz’s
name has priority.
= Eudamus albimargo, Mab.
= Goniurus celus, Cram.
Hesperiidee described by Plétz.
ELudames tellus, Pistz (1327), Bu-
enos Aires.
7 tynatius, Plotz (1328),
oc. P
+) albociliata, Plétz (1329),
Colombia.
Felemiades dianina, Plotz (1331),
S. Am.
Netrocoryne coronus, Plétz (1332),
Chiriqui,
Feleyonus corentinus, Plotz (1333),
Surinam.
5 ophiuchus, Pléta (1334),
Surinam.
3 mutius, Plétz (1835), Co-
lombia.
pertica, Plétz (1336), S.
Am.
Arteurotia meris, Plétz (1839), Co-
lombia.
epipola, Plitz (1340),
Cayenne.
Aithilla Weymeri, Plotz (1342),
Loe. ?
Hesperia caura, Plotz (1853), Suri-
nam.
» Ldlicht, Plotz (1354),
Loe. ?
» salma, Plotz (1555), Pa-
nama.
» eda, Plitz (13857), Blu-
menau.
» eamposa, Plotz (1360),
Brazil.
» gquispica, Plotz (1861),
Peru.
angulis, Plétz (1567), Pa-
nama.
» luscinia, Plotz (1869),
Blumenau.
» gtla, Plotz (1870), Ari-
zona.
licta, Plétaz (1372), Loe. P
» emeritta, Plitz (1878),
Brazil.
diana, Plotz (1383), San
Paulo.
hilda, Plotz (1384), Blu-
menau.
A Naseus, near cephise, H.-S.
Drawing missing.
) 7
7 »
” ”?
” 97
= Bungalotis sebrus, Feld., 3.
= Bungalotis salatis, Cram., 2 , var.
Near Lychnuchus clearchus, Plotz,
but with three subapical spots
and an oblique band on the
primaries white.
Probably a Pellicia.
= Sproniades clinias, Mab. Genus ?
=Telegonus chiriquensis, Staud.,
and 7. meretriz, Hew. Ploitz’s
drawing is taken from a Chi-
riqui specimen.
May belong to Papvas.
Genus? Typeis Q.
Possibly = Mnasilus penicillatus,
Godm.
Genus? Not unlike Perimeles
remus, F,
Belongs to Pyrrhopygopsis. Not
represented in the G. & 5S.
coll.
A Pyrrhopygopsis, near socrates,
Mén., but with yellow cilia
and a black head.
= Damas clavus, Er., g. Proteides
cervus, Moschl., is the 2 of the
same species.
Belongs to Perichares, near agrippa,
Godm., ¢.
= Cogia hippalus, Edw., 3.
= Xenades orchamus, Cram., ¢,
var.
Not identified in the G. & S. coll.
152
Hesperia uruba,
=
Dr. F. D. Godman on American
Plotz (1385),
Brazil.
zygia, Plotz (1388), Loc. ?
yva, Plétz (1889), Loe. ?
angellus, Plotz
(1393),
Chiriqui.
velleius, Plétz (1394), Su-
rinam.
corisana, Plotz (1895),
Surinam.
zeppa, Plotz (1397), Suri-
nam.
éthra, Plétz (1899), Suri-
nam.
mulla, Plotz (1401), Suri-
nam.
Savetta, Plotz (1408), Chi-
riqui.
conta, Plitz (1410), Minas
Geraes.,
autumna, Plotz (1411),
Centr. America.
cabella, Plotz
Puerto Cabello.
subviridis, Plotz (1426),
San Paulo.
noctis, Plétz (1481), Chi-
riqui.
(1419),
gereon, Plotz (1482),
Para.
aurinia, Plotz (1436), Ja-
maica.
Zenckii, Plétz (1487),
Mexico.
monica, Plotz (1489),
Blumenau.
piso, Plétz (1440), Pa-
nama.
zela, Plitz (1441), Monte
Video,
hyana, Plotz
(1448),
Chile.
Not identified in the G. & S. coll.
? American. Unknown to me.
Very like Cobalets argus, Moschl.
The specimen figured is from
Blumenau, Brazil.
Perhaps the 2 of Halotus saxula
(Mab.), Godm., with whiter
spots. Plétzs name has
priority.
=Rhinthon bistrigula, H.-S. (alus,
Godm.).
Unknown to me.
= Hutychide complana, H.-S. (mi-
dia, Hew., gura, Plotz). Plotz’s
figure of upperside too green,
as in many other cases.
Perhaps= Pamphila ancus, Moschl.
= Cobalopsis edda, Mab. ¢o. H.
pelora, Plotz, is apparently a
large 2 of the same species,
and this is a still older name,
both antedating that of Ma-
bille,
= Rhinthon chiriquensis, Mab., 3.
Not identified in the G. & S. coll.
= Megistias epiberus, Mab., dark
var. See note under Hesperia
lysias, Plotz (fig. 623).
Not identified in the G. & S. coll.
Probably belongs to Limochores or
Serdis. Not contained in the
G. & 8. coll.
= Thymelicus vibex, Hiibn., 3, vars.
H, combinata, Plotz (686), is
the same species.
An Aftrytone, near eulogius, Plotz.
There isa ¢ of it, from Novo
Friburgo, in the G. & S. coll.
= Atrytone rolla, Mab., ¢.
Probably belongs near <Atrytone.
Not identified in the G.& 8.
coll.
= Hylephila fulva, Blanch., 3.
Hesperiidz described by Plétz.
Hesperia morganta, Plotz (1444),
Ss. Am.
» catilina, Plotz (1445),
Blumenau.
» genoa, Plitz (1446), Ne-
vada.
» axtus, Plotz (1448), Colo-
rado,
» Ubrita, Plotz (1453), Pa-
nama.
» hyboma, Plotz
Minas Geraes.
(1455),
Pamphila humeralis, Mab. (1456),
Para.
Apaustus vopiscella, Plétz (1471),
Minas Geraes.
» facths, Plotz (1474),
Surinam.
; scheria, Plotz (1475),
Para.
a alsimo, Plotz (1479), Su-
rinam.
5 polita, Plotz (1481),
Loe. ?
» fabulinus, Plotz (1482),
Surinam.
» bebarus, Plétz (1488), Co-
lombia.
» servilius, Moschl. (1487),
Surinam.
» sulla, Méschl. (1488),
Colombia.
Heteropterus Jelskyi, Ersch. (1490),
Peru.
Apaustus krexvos, Plotz (1491),
Para.
a matuta, Plotz (1498),
Loe. ?
Thymelicus macra, Plotz (1500), N.
Am,
3 tucumanus, Plétz(1503),
Cordova.
Cyclopides facetus, Plitz (1580),
Loe. ?
Pythonides tnanca, Plitz (1535),
Loe. ?
153
=Thymelicus brettus, Boisd., 3,
var. The specimen figured by
Plotz is from North America.
Genus? H. sulfurina, Mab., is
probably the same species; and
if this is the case, Mabille’s
name has priority.
? = Polites sabulett, Boisd., var.
Belongs to Cherephon, and pro-
bably =rhesus, Edw., d 2.
= Augiades hecale,Godm, Plotz’s
name has priority.
Perhaps belongs to Padraona.
Underside of secondaries
marked very much as in Phe-
nuades propertius, F,
Near Molo. Not in G. & S. coll.
Not identified in the G. & S. coll.
Probably Mnasitheus simplicissima,
By-85:9).
Genus? A specimen () from the
Amazons in the G. & 8S. coll.
Probably = Callimormus gracilis,
Feld.
A Callimormus,
Scudd.
Probably a Callimormus, near vetula,
Tab.
Genus? A @ from the Lower
Amazons in the G. & §. coll.
very near it.
Genus? Not identified in the
G. & 8. coll.
Not identified in the G. & S. coll.
near juventus,
Unknown to me.
Not unlike A. imerius, Plitz (765).
Perhaps an ally of Hesperia reticu-
lata, Plotz. Genus?
= Coveodes singularis, H.-S. (auan-
traca, Hew., procris, Edw.),
var.
Belongs to Ancylorypha, and pro-
bably =arene, Edw., 9.
A Butleria, near flavomaculata,
Blanch. Specimen figured is
from Chile, and there is one
from Mendoza in the G. & 8.
coll.
= Heliopetes petrus, Hiibn., and
not H. arsalte, L., as queried
in the ‘ Biologia.’
154
Leucochitonea pampina, Plotz
(1536), Buenos Aires.
Pyrgus Willi, Plotz (1587), Minas
Geraes.
» erisia, H.-S. (1550), Cuba.
Pythonides dilucida, Moschl., (1552),
Surinam. °
Achlyodes onorbo, Moschl, (1555),
Surinam.
Pythonides (Papilio) certalis, Cram.
(1555), S. Am.
Pythonides portulana, Plétz (1557),
Ss. Am
Achlyodes fatinitza, Plotz (1558),
Colombia.
®: ancholis, Plotz (1559),
Colombia.
Antigonus ceclydes, Plotz (1560),
Colombia.
5 tolimus, Plotz (1561),
Colombia.
5 obliqua, Plotz (1562),
Loe. ?
ss gorilla, Plotz (1567),
Panama.
Nisoniades tucumanus, Plotz (1568),
Cordcva.
astur, Plow (1569), Su-
” Z
rluam.
oH bibiana, Plotz (1571),
Colombia.
os cecus, Plotz (1572),
Loe. ?
Tagiades chacona, Plitz (1578),
Panama.
monartus, Plétz (1579),
Panama.
vincula, Plotz
Panama.
(1580),
#)
On American Hesperiide described by Piotz.
Belongs to Heliopetes. From
Corumba, Chapada, and Para-
guay in the G. & 8. coll.
= Helwopetes domicella, Ex.
Belongs to Hesperia. Plotz’s figure
is taken from a Porto Rico
specimen.
= Chiomara asychis, Cram., var.
A Chiomara, near asychis, Cram.
Plotz figures aspecimen of P. assecla,
Mab., from Brazil, as this
species. There is a specimen
of the latter from Chontales
in the G.& S. coll., hitherto
overlooked.
=P, festivus, Er.
Not identified in the G. & S. coll.
= Hesperia tetra, Mab. Referred
by Mabille to his genus Bolla.
= Aithilla echina, Hew., var.
Probably belongs to G‘orgophone,
near varius, Mab., ¢.
Belongs to Anastrus, There is a
specimen of it from Santarem
in the G. & 8. coll.
Closely related to Bolla giselus,
Mab., but smaller and with the
two small hyaline spots on
the primaries well separated.
Specimen figured isa ¢.
Belongs to Staphylus or Bolla.
has a costal fold.
A species with peculiarly formed
antenne (if correctly drawn)
and a costal fold. Unknown
to me.
Belongs to Anisochoria, near sub-
limbata, Mab.
A Cyclosemia,
Stoll.
An Ebrietas, near ecliptica, Butl.,
with bluish grey on the outer
part of the secondaries beneath.
Figure not good.
= Celenorrhinus chiriquensis, Mab.
Ploétz’s name has priority.
Probably a large Q of Staphylus
mazans, Reak. There is a
very similar unnamed specimen
from Caché, Costa Rica, in the
G. & BS. coll.
near herennius,
On the Hand and Foot in Man and Apes. 155
eee serenus, Plotz (1687), =Entheus ewmelus, Cram., °.
oc.
Pyrrhopyge porus, Plitz (1595) =Mimoniades pityusa, Hew.
Colombia.
cn parima, Plotz (1597), Belongs to Yanguna. Not repre-
Surinam. sented in the G. & S. coll.
5 leucoloma, Ersch. Very near P. sergius, Hopff., but
(1599), Peru. with the white patch on the
underside of the secondaries
® extending further inward.
XXV.—Function and Form with Reference to the Hand and
Foot in Man and Apes. By Duncan C. L. Frrzwi.uiams,
M.D., Ch.M., F.R.C.SS. Ed. & Eng., Demonstrator of
Anatomy, King’s College, London; Casualty Officer,
Hospital for Sick Children, Great Ormond Street.
[Plate V.]
TurouGH the kindness of Professor Cunningham I was
enabled to carry out a systematic anatomical study of a gibbon
(Hylobates agilis) in his possession. In this paper I wish to
deal only with the description of the hands and feet of the
animal, and incidentally to draw attention to the differences
existing in the hands and feet of man.
The orang-utangs, chimpanzees, gorillas, and gibbons are
the four great tribes which form the anthropoid family. The
larger members of the family resemble man in stature and
outward form more nearly than the gibbon, but on closer
investigation the gibbon presents certain characteristics which
have led many eminent anatomists to place this ape next to
man in the scale of animal life. With the single exception
of man, the gibbon can assume the erect posture more
completely than any other animal.
The Hand.
The hand of the gibbon (PI. V. fig. 1, A) is remarkable for
the great length it possesses in comparison with its width,
Measured from the crease in front of the wrist to the tip
of the longest tinger the length amounts to 15} cm., but the
breadth of the widest part, just above the root of the little
finger, does not exceed 31 cm. The length is due to the
development both of the metacarpals and phalanges.
The thenar and hypothenar eminences are small; indeed,
so feeble is the development of the latter that it can scarcely
156 Dr. D. C. L. Fitzwilliams on the
be said to exist. The thumb is short in comparison with the
rest of the hand; it extends slightly beyond the level of the
metacarpo-phalangeal joint of the second digit, but not as far
as the web between the second and third. Inman the thumb
reaches to the level of the proximal interphalangeal articu-
lation of the index, while the interphalangeal joint of the
thumb is on a level with the metacarpo-phalangeal joint of
the second digit.
The web of the gibbon’s thumb is very small, not reaching
halfway down the first metacarpal bone. This gives rise to
an appearance as if the palm and the thumb sprang separately
from the region of the wrist. In marked contrast to this, the
webs of the fingers are much more extensive than those of
man, and reach nearly halfway down the elongated proximal
phalanges, thereby lengthening the palm at the expense of
the fingers. This extensive webbing prevents separation of
the fingers to any great extent. As in man, the third digit
is the longest. ‘The fingers are placed parallel to one another,
but show slight ability to oppose the thumb.
The positions of the deep creases of the skin are as follows :—
1. In front of the wrist there is a deep transverse crease
due to the flexion of this joint.
2. From just below the midpoint of the wrist-crease another
starts, and runs downwards and outwards into the deep cleft
which intervenes between the thenar eminence and the rest of
the palm. This crease and cleft are caused by the adduction
of the thumb. In man the great development of the thumb-
muscles has filled out the cleft and increased the web, while
the upper limit of the crease curves outwards round the thenar
eminence well below the creases of the wrist.
3. Starting from the same point as the preceding, a crease
passes downwards and inwards to the inner margin of the
palm, just above its centre. ‘The causation of this line is not
very evident, as the muscles of the hypothenar eminence are
smal]. Of such a crease little or no trace is to be found in
the human hand; but in the foot of the ape a very similar
marking is found in front of the outer part of the heel, the
part which corresponds to the hypothenar eminence of the
hand (see Pl. V. tig. 1, 3B):
4. In common with the two preceding yet another crease
starts and runs longitudinally down the palm to the root of
the third digit.
5. Parallel and to the outer side of the last-mentioned
crease is a shorter crease which ends at the cleft between the
second and third digits. These creases result probably from
the contraction of the contrahentes and palmar interosseous
Hand and Foot in Man and Apes. eat
muscles, which causes the digits to converge on the central
one. Both these creases are usually found in the hand of
man, but are fainter, more irregular, and more obliquely
placed. I believe that cheiromancy attributes great brain-
power to the individual possessing one of these lines in a
well-marked degree, an assumption highly complimentary to
the ape.
6. About a third of the way down the palm is a crease
passing transversely across.
7. Halfway down the palm a crease passes across in a
curved manner with the concavity towards the wrist.
8. A curved crease, parallel to the last mentioned, is
situated about two thirds of the way down the palm. ‘These
last two creases evidently correspond to the two deeply
marked creases which run obliquely across the human palin.
All three transverse creases are due to the folding of the
palmar tissues during flexion of the fingers. The obliquity
of the lines in the human hand results from the great amount
of opposability which exists between the human thumb and
the inner four digits. The extra line found in the palm of
the ape is probably accounted for, partly by the increased
length of the palm, and partly by the constantly flexed
position of the hand.
9. Deep creases are present at the roots of the fingers on a
level with the webs. These creases run transversely in the
case of the second and third digits and obliquely in the case
of the fourth and fifth. ‘This same arrangement, though to a
less marked degree, is present in the hand of man; the creases
of the two outer fingers are transverse, while those of the two
inner have an oblique tendency.
The long slender fingers of this ape (Pl. V. fig. 2) are
remarkably flat on their palmar aspect. They all exhibit, to
a greater or less degree, a longitudinal crease which runs
down the centre of the palmar surface. This crease is best
marked on the middle digit. It will be seen later that the finer
skin-lines of the fingers converge on these central creases.
Owing to the proximal interphalangeal joint of the middle
digit being marked in front by two widely separated trans-
verse lines, this digit appears to possess four instead of three
phalanges. The thenar eminence is marked by a series of
oblique lines running downwards and inwards. On adducting
the human thumb similar lines may appear.
The terminal phalanges are bulbous and projecting. The
free part of the nail stands well away from the back of the
phalanx ; in man the nail hes quite close to the back of the
ungual phalanx. As the nail of the thumb is the shortest,
158 Dr. D. C. L. Fitzwilliams on the
broadest, and least arched, it most closely resembles the
human nail. As one approaches the little finger the nail of
each digit becomes successively longer, narrower, more arched,
and therefore more claw-like, ina manner analogous to that
noted by Wiedersheim in the human hand. This is due to
the greater and more varied use to which the nails of the
outer digits are put; that of the little finger, being the least
used, retains in a greater degree than the others its primitive
claw-like character.
The fine lines on the palmar skin are much coarser than
those on the human hand, and in many cases pass inter-
ruptedly across the deep creases of the palm.
The coarseness of the finer lines is probably a response to
the requirements of function: the animal uses its hands as
hooks by which to suspend the weight of the body from the
branches, and the friction produced by the finer lines gives
firmness to the grip. If the skin of the palm were devoid of
lines and perfectly smooth a firm grip, with no slipping,
would be difficult to obtain. The delicacy of the lines of the
human hand is due to the fact that the hand is no longer an
organ merely of grasp, but used for many varied and highly
specialized movements.
In the gibbon the lines are arranged in a definite manner
and one well calculated to obviate the risk of the hand
slipping from a branch. In the lower part of the palm the
lines show a general tendency to converge towards the central
digit. In the fingers the lines are arranged in achevron-like
manner, converging from the margins downwards towards
the longitudinal crease which runs along the centre of each
digit. This arrangement is best seen on the proximal and
middle phalanges of the third and fourth digits, where the
weight of the body chiefly rests. The lines of the proximal
and middle phalanges of the second and fifth digits, like the
lines of the palm, have a tendency to converge towards the
central digit. This tendency survives in the human hand
only on those parts which are used solely for grip and not for
the execution of any of the finer movements. It is well seen
on the palm near to, and on the proximal phalanges of, the
index and little fingers, whilst elsewhere the lines run in an
irregular manner, forming whorls and triangles which differ
in individual hands. ‘This wonderful difference in the
patterns met with on the thumb is used by the police in the
identification of criminals.
Oblique lines are found to be more efficient mechanical
agents for the prevention of slipping than transverse. For
Hand and Foot in Man and Apes. 159
this reason the driving-wheels of all heavy traction-engines
are provided with sloping or chevron lines.
Professor Goodsir [1] many years ago pointed out that the
hand of man was the only perfect hand, and that while the
ape’s hand was fitted to grasp a cylinder such as the branch
of a tree, it was much less able than the human hand to grasp
a sphere. Hepburn [2] has compared the hands of the four
anthropoids with that of man, and shown how the develop-
ment in the latter of the muscles of thenar and hypothenar
eminences has modified the position of the creases. In the
gibbon the fingers are capable of flexion and adduction to
the middle line, but have little tendency to oppose the thumb ;
the transverse and longitudinal creases are therefore met
with. In man opposition of the thumb to the fingers is one
of the most prominent characteristics of the hand, and the
creases in consequence are oblique.
The Foot.
The foot (Pl. V. fig. 1, B), from the point of the heel to the
end of the middle toe, measures 14 em. in length, of which two
thirds belong to the sole and one third to the toes. The width
of the sole, exclusive of the great toe, is 2°6 cm. The hallux
reaches to the level of the proximal interphalangeal joint of
the second toe. Unlike the foot of man, the hallux is not
bound up parallel to the other digits, but becomes free just
beyond the mid-point of the sole. Behind this point it is
marked off from the rest of the foot by a deep crease. The
hallux is a much more powerful digit than the pollex. All
the toes have a tendency to point outwards, being set at an
angle to the plane of the long axis of the sole. In conse-
quence of this angle the four outer digits, on flexion, oppose
the great toe. The web of the four outer toes reaches about
halfway down the proximal phalanges and prevents any great
separation of these digits.
The following are the chief skin-creases found in the sole :—
1. On looking at the sole it is seen to be divided into two
unequal parts by a longitudinal crease which starts on the
inner side of the heel and, passing forwards, gradually deepens
into a cleft which divides the great toe from the rest of the
sole. The smaller of these portions carries the hallux, while
the larger bears the four outer toes. This dividing crease
and clett are caused by the opposability of the hallux.
2. From the preceding crease near its commencement,
another smaller crease passes outwards and forwards in front
160 On the Hand and Foot in Man and Apes.
of the heel, and corresponds to a similar crease in the hand
of the ape. In the human foot neither of these creases is
seen.
3. A well-marked crease runs transversely across the sole,
even on to the hallux just behind the point where that digit
becomes free from the rest of the foot.
A, A still deeper crease runs transversely across the sole
just behind the metacarpo-phalangeal articulations. This
marks the place at which the long axes of the sole and digits
meet at an obtuse angle.
5. At the root of the four other toes there is a deep double
crease which slopes from the margins of the foot forward to
the middle digit. Creases 3, 4, and 5 are caused by the
folding of the tissues on flexion of the toes into the sole.
Hepburn [2] figures a crease in the foot of the gibbon
running longitudinally, and ascribes its presence to the con-
tractions of the adductor (contrahentes) layer of muscles.
In my specimen this crease was absent, and it is interesting
to note that the adductor muscles of the four outer toes were
also wanting.
The creases and the finer lines on the toes correspond to,
but are not so well marked as, those found on the fingers.
The finer lines on the sole start from the heel and pass in
wide curves to the margin of the foot: those on the outer
side to the outer margin; those in the centre forward with a
slight general inclination inward; and those on the inner
side forward and inward to the cleft between the sole and
great toe. This same arrangement is reproduced on a
smaller scale on the skin of the ball of the great toe.
The description of the finger-nails applies equally to the
toe-nails. The ungual phalanges of the toes were, if any-
thing, a trifle more ‘bulbous than those of the fingers.
From my account it is clear that there is much less resem-
blance between a man’s and a gibbon’s foot than there is
between their hands. In fact, of the hand and foot of the
gibbon, it is the latter w hich more closely resembles the
human hand. ‘The central digit of the ape’s foot and hand
corresponds to the central digit of the human hand, namely
the third, while the central digit of the human foot is the
second. The marked difference between the hand and foot of
man is due to specialization. Man uses his hand, said
Goodsir, ‘as an instrument for acting on matter, in the terms
of his human faculty of thinking in space’; while his foot
is an organ merely of support and progression. Commonly
all other functions are precluded, for the foot is enclosed in a
boot.» But among peoples, such as the natives of India,
On Neotropical Mammals. 161
where custom does not confine the feet in coverings, the
human foot retains a certain amount of its prehensile power.
In the ape the foot is far superior to the hand as a prehensile
organ, for not only is the hallux better developed than the
pollex, but it has much greater power to oppose the other
digits. Grasping and progression in animals of arboreal
habits are to some extent synonymous. In the human foot
the hallux is so bound up with the rest of the foot that it is
practically incapable of any independent action.
References.
[1] ‘Anatomical Memoirs of John Goodsir.’ Vol. I. Edited by W.
Turner,
{2] ‘Journal of Anatomy and Physiology,’ 1893.
EXPLANATION OF PLATE YV.
Fig. 1. A, hand, B, foot of Hylobates agilis, to show the arrangement of
the creases. Owing to the hook-like position of the hand, the
finger-tips are out of focus.
Fg. 2. Skin from the palmar aspect of the fingers, to show the cheyron-
like arrangement of the fine lines,
XXVI.—On Neotropical Mammals of the Genera Callicebus,
Reithrodontomys, Ctenomys, Dasypus, and Marmosa. By
OLDFIELD THOMAS.
Callicebus pallescens, sp. n.
Allied to C. donacophilus, d’Orb., with which it shares the
greyish-white hands, feet, and tail, but the head and body are
almost of the same pale colour, so that the whole animal is
one of the palest and most uniformly coloured species of the
group.
Size very small. Tur thick and soft; the longer hairs of
back about 60, the shorter 35 mm. in length. General
colour of body pale greyish, suffused with pinkish buff; the
long hairs indistinctly ringed with whitish and black, the
underfur pinkish buff for its terminal half, its basal half dark
brown. Under surface and inner side of limbs rufous, rather
paler than in C. donacophilus. Head rather yellower than
back, owing to the hairs being tipped with yellow, but the
difference is not conspicuous. Muzzle and lips whitish.
Hands and feet greyish white. Tail also greyish white, but
the hairs inconspicuously ringed with blackish.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. bi
162 Mr. O. Thomas on
Skull chiefly remarkable for its small size as compared
with that of any other species.
Dimensions of the type (aes sie on skin) :—
Head and body 365 mm.; tail 390; hind foot 84.
Skull : Bade length 58°5 ; basal length 44:5; zygo-
matie breadth 39; mastoid breadth 34° 3% front of canine to
back of m? 18:2.
Hab. Chaco of Paraguay; type from 30 miles N. of
Concepcion.
Type. Male. B.M. no. 94. 3.6.1. Collected October
1893 by Dr. J. Bohls. Two specimens.
In the absence of fresh specimens of C. donacophilus these
skins from Paraguay were provisionally referred to that
species. Now, however, a series of skins from Bolivia,
collected by Mr. Steinbach, and nearly topotypical of
d’Orbigny’s species, prove conclusively that the present form
needs description. Its differential characters are given above.
Grison* furax, sp. n.
One of the smaller species of the group, of a strongly
yellowish colour.
General colour comparatively yellow or buffy, the facial
streaks and the ends of the dorsal hairs buffy or buffy ochra-
ceous, the lateral streak behind the ear more deeply ochra-
ceous. Underfur of back brown, darkening terminally.
Under surface and limbs wholly black, the belly without any
light tipping. ’ail-hairs brown basally, broadly tipped with
buffy ochraceous ; parted on the upper surface in such a way
as to show a narrow, median, ochraceous-buffy line along the
centre, composed entirely of short woolly hairs.
Skull with the carnassials and molars of medium propor-
tions, the lower carnassial without a supplementary internal
cusp.
Dimensions of the type tule fully adult) :—
Head and body 317 mm.; tail 110; hind foot 54; ear 22.
Skull: basal léneth 60; oreatest breadth 37 3 length of
p* on outer edge 7°5; greatest diameter of m’ 6° aa
Hab. Southern Brazil. Type from San Francisco dos
Campos, 8. Minas Geraes. Altitude 1580 m.
Type. Immature male. B.M. no. 1. 6. 6. 25. Original
number 622. Collected 29th March, 1901, by Alphonse
Robert.
This is the common Grison or “ Furao” of Southern
Brazil, generally known by the name of “ Galictis vittata” ;
* Oken, 1816. Better known as Galictis.
Neotropical Mammals, 163
but it seems to be without a name properly applicable to it.
The Viverra vittata of Schreber was based on a Surinam
animal, probably one of the group with a supplementary cusp
on the lower carnassial, and certainly not the present form, to
which its name has been so commonly applied. Thunberg’s
Ursus brasiliensis is, again, clearly the larger type of this
group, and would appear to be the same as Nehring’s
Galictis crassidens.
The Chilian G. cuja, Mol. (syn. G. quiqut, Mol.), agrees
with G. furaz in the essential characters of size and tooth-
structure, but in colour is much greyer, the buffy parts of
G. furax being replaced by white or whitish.
Grison furax luteolus, subsp. n.
Similar to true furaz, but paler throughout.
Size and general characters as in G. furax, but the facial
bands, the tips of the dorsal and caudal hairs, and the pale
short-haired line along the top of the tail are all much paler
in colour, being approximately ‘ cream-buff” instead of
ochraceous buff. Underfur dull whitish butt, giving a con-
spicuously paler tone to the whole animal. Hairs of belly
tipped with pale buffy, the throat, chest, inguinal region, and
limbs being, as usual, black. Extreme tip of tail with a
siaall tuft of white hairs. Four pairs of mammze, all inguinal.
Dimensions of the type (measured in flesh) :—
Head and body 380 mm.; tail 170; hind foot 60; ear 22.
Skull: basal length 66°5; greatest breadth 41°3; mastoid
breadth 37°5; palatal length 33; length of p* on outer
side 7'1 ; greatest diameter of m’ 5°2.
Hab. Chulumani, Bolivia, 67° W., 16° S. Alt. 1800 mn.
Type. Old female. B.M. no. 1. 6. 7. 27. Original
number 1305. Collected 31st December, 1900, by Perry O.
Simons ; presented by Oldfield Thomas.
‘This form may be readily distinguished from the Brazilian
animal by its more creamy-buff colour, and especially by its
light underfur.
Reithrodontomys modestus, sp. n.
A small species of dark colour, with a short and uniformly
dark tail.
Size about asin F. saturatus. General colour dark, nearest
to Ridgway’s bistre, the sides more drabby, the dorsal area
rather blacker. Under surface soiled greyish (grey no. 6),
not sharply defined, the bases of the hairs blackish slate; a
large patch between the fore limbs drab, like the lower flanks.
Ears short, uniformly blackish. Upper surface of hands
164 Mr. O. Thomas on
white, of feet dull white with a darker area along the outer
side of the metatarsus, but this area is not sharply contrasted
as is sometimes the case. ‘Tail little longer than the head
and body, well-haired, uniformly blackish above and below,
the proximal inch below only inconspicuously lighter.
Skull thin and papery, without strong ridges. Palatal
foramina reaching to the level of the second lamina of m!.
Bulle comparatively small.
Dimensions of the type (measured in the flesh) :—
Head and body 59 mm.; tail 70; hind foot (s. u.) 16°5 ;
ear 12°5.
Skull: greatest length 21; basilar length 15; length ot
nasals 7°8; interorbital breadth 3°5; breadth of brain-
ease 10°5; palatal foramina 4°6; length of upper molar
series 3°3.
Hab. Jinotega, Nicaragua. Altitude 4650’.
Type. Adult male. Original number 29. Collected 20th
January, 1906, by Mr. M. G. Palmer.
This Retthrodontomys is distinguishable by its small size,
short ears, dull coloration, and short unicolor tail.
Mr. W. H. Osgood has been kind enough to compare this
mouse with the specimens in the U.S. Department of Agri-
culture collection, and he tells me that it does not appear to
resemble very closely any of the species there. ‘It is,
perhaps, nearest to Ff. australis of Costa Rica, but is darker
on the upper parts, slightly smaller, and the tail is shorter
and more nearly unicolor.”
Ctenomys Stetnbacht, sp. n.
A fairly large species, of a uniform coppery-grey colour.
Size rather above the average in the genus. Fur straight,
fine and glossy; hairs of back about 138 mm. in length.
General colour a peculiar dark drabby grey-brown or coppery,
quite unlike that of any known Ctenomys, but very similar
to that of certain Geomyide, e. g. Zygogeomys trichopus,
Merriam. ‘This colour is quite uniform over the whole of
the head, upper surface, and sides. Under surface creamy
white, the hairs dull slaty for their basal two thirds; line of
demarcation on sides fairly sharply defined. Whiskers white.
Chin and a band across the lower neck in front of the arms
brown, separated by a broad whitish patch running across
the interramial region and narrowing on the sides to a point
about half an inch below each ear. Arms and legs light-
coloured, except a narrow band running down the front of
the forearms; hands and feet almost naked above, pale
brown, the lateral fringes whitish. ‘Tail very thinly clothed,
its sparse hairs dull white.
Neotropical Mammals. 165
Dimensions (taken on the skin) :—
Head and body 245 mm.; tail 86; hind foot, s. u. 41,
c. u. 405.
Hab. Campo of Province Sara, near Santa Cruz de la
Sierra, Bolivia.
Type. Adult male. Collected by Mr. J. Steinbach.
This striking species differs so widely in its coloration
from every known Ctenomys that even in the absence of the
skull I have no hesitation in describing it as new. Its colour
above is extraordinarily like that of Zygogeomys trichopus
and some other Geomyide, which have a similar coppery-
brown tone, while all the hitherto known Ctenomys are ot
some fawn or buffy tint.
A baby specimen of Cé. Steinbachi, only 100 mm. in
length, is of precisely the same colour as the adult.
The Local Forms of Dasypus sexcinctus, Linn.
While the extreme forms of the Dasypus seacinctus group
are, as shown in a former paper *, so widely different in size
that it seems impossible for them to belong to the same
species, further material from intermediate localities tends to
fill up the gap between them, and I am now disposed to
regard them as forming but a single species with several
geographical races.
These races may be briefly distinguished as follows :—
1. Size smallest; greatest skull-length 95 mm.,
cephalic shield in an average specimen
87x65. Colour brown. About 31-33
scales in movable bands. Back thinly
haired, the pelvic shield practically naked. [ Linn.
(Bara? ) Bears eR COME A cremedenated oicheFtesk cos eceser D, sexcinctus sexcinctus,
. Size medium ; greatest skull- length of adult
114 mm., cephalic shield about 10377.
Colour paler, more yellow. Hairs as in
eexcinctiso. (Bahia: i .\5 <)sc cae aes .... D. 5s. setosus, Wied.
. Size largest; greatest skull-length attaining
126 mm., cephalic shield 12588. Colour
brown. Hairs as in sexcinctus. (Matto
Grosso, Paraguay, and South Brazil.) .... D. s. gilvipes, Ul.
4, Size rather less than in gilvipes; cephalic
shield 11489 mm. Colour pale horny.
Movable bands with 386 scales. Back
scantily haired. Skull short and broad ;
molars broad and rounded, (Bolivia.).... D. s. bolivia, subsp. u.
5. Size as in bolivie ; cephalic shield 11678
mm. Colour sandy. 36 scales in movable
bands. Back well haired, the pelvic shield
with many white hairs. Nasals rather [subsp. n.
peculiar in shape. (Tucuman.).......... D, s. tucumanus,
bo
co
* P. Z. 5, 1903, ii. p. 242.
166 Mr. O. Thomas on
The new forms may be more fully described as follows :—
Dasypus sexcinctus bolivia, subsp. n.
Size rather less than in gilvipes. Colour pale; hairs of
carapace white, of soft parts brown, nowhere really black.
Hairs not more numerous than in gilvipes, the scales of the
movable bands each with a couple of white bristles about
2-4 cm. long at their posterior end; pelvic shield almost
naked, its few bristles rarely more than 1 em. in length.
Scales more numerous than in the Eastern forms, the median
movable bands consisting of 36 scales. Frontal shield very
broad in proportion to its length.
Skull broad and stout, with broadly and abruptly expanded
zygomata. Frontal region but little convex. Nasals nearly
parallel-sided, not of the peculiar shape found in tucumanus.
Palate broader. Molars more broadly rounded, the fifth
maxillary tooth 6°4 x 4°7 mm.
Dimensions of the type (measured on the flattened skin) :—
Head and body 500 mm. ; tail 250.
Frontal shield 114 x 89.
Skull: condylo-nasal length 119; basal length 100°5 ;
zygomatic breadth 75; nasals, length 43°3, breadth ante-
riorly 18, mesially 18°3, posteriorly 20; palatal length 68 ;
breadth of palate between fifth maxillary teeth 16:4,
Hab. Near Santa Cruz de la Sierra, Bolivia.
Type. Old male. Original number 139. Collected
17th May, 1906, by Mr. J. Steinbach ; presented by Oldfield
‘Thomas. One specimen.
Lhe pale colour and more numerous scales ally this form
to the next subspecies, while it resembles D. s. gilvipes in
its scantily haired pelvic shield. Its very broad and rather
flattened skull is peculiar to itself,
Dasypus sexcinctus tucumanus, subsp. n.
Size rather less than in the large Paraguayan D. s. gilvipes.
Colour paler, the carapace itself sandy, its hairs white.
These latter are fairly numerous both on the movable bands
and on the pelvic shield, and attain from 3-5 cm. in length
along the sides. Scattered hairs of underparts, arms, and
legs mostly black, with a few whitish ones intermingled.
Scales of carapace rather smaller than in the Eastern forms,
there being 33-86 scales on each of the movable bands, the
numbers on the shoulder and pelvic shields proportionally
increased.
Nasals narrow in front and behind, angularly broad in the
Neotropical Mammals. 167
middle, the middle part projecting outwards on each side in
an angle at the premaxillo-maxillary suture. Posterior
narial opening narrow, the notch sharply V-shaped. ‘Teeth
narrow throughout, the fifth maxillary tooth 6:2 x 4-2 mm.
Dimensions of the type (as taken by collector in the
flesh) :—
Head and body 410 mm. ; tail 230.
Cephalic shield 116 x 78.
Skull: back of frontals to end of nasals 85; greatest
breadth 64; nasals, length 40, breadth, anteriorly 11, in
middle 19, posteriorly 12°5; palatal length 64; breadth of
palate between fifth maxillary teeth 13:3.
Hab. Tapia, Tucuman. Alt. 700 m.
Type. Adult male. B.M. no. 3. 6.6.16. Original num-
ber 1910. Collected 23rd September, 1902, by L. Dinelli;
presented by Oldfield Thomas. ‘T'wo specimens examined.
Marmosa chloe, sp. n.
A small dark-coloured species, with creamy chest and
inguinal region.
Size about as in MV, fuscata, Thos., and M. Klagest, All.
Fur soft and fine; hairs of back about 6 mm. in length.
General colour above sepia along the dorsal area, shading off
into bistre on the sides. Under surface, on throat, chest, and
inguinal region, pale pinkish or creamy buff, the hairs this
colour to their bases ; but on the belly the slaty-based hairs
encroach on each side, so as to narrow the creamy part to a
mere median line. Sides of face with a large ill-defined
black patch surrounding the eyes, the paler area between
them less marked than in K/ages¢. Lower cheeks cream-buff.
Ears naked, dark grey. Hands pale brown, feet whitish.
Tail with its short-haired part about half an inch in length,
the remainder naked, uniformly pale grey.
Skull shaped very much as in M. Klagesi; similarly
broad, with expanded zygomata and fairly well defined supra-
orbital ledges. Nasals expanded posteriorly.
Dimensions of the type (measured in the flesh) :—
Head and body 143 mm. ; tail 170; hind foot 21; ear 19.
Skull: greatest length 33°7; basal length 30; greatest
breadth 18; nasals 14x 4; interorbital breadth 5-2; breadth
of brain-case 13°5; palate, length 18:2, breadth outside
m® 10°35; combined length of three anterior molariform
teeth 5°6.
Hab, Demerara River, 29 miles above Georgetown, British
Guiana.
168 Bibliographical Notice.-
Type. Male. Original number 13. Collected 6th De-
cember, 1906, by Mr. 8S. B. Warren.
This species is very much darker throughout than
M. Klagesi, to which it appears to be most nearly allied.
The similarly dark Jf. fuscata has a wholly grey-mixed
belly and a much more elongate skull, without trace of
supraorbital beads.
BIBLIOGRAPHICAL NOTICE.
Catalogue of the Madreporarian Corals in the British Museum
(Natural History). Vol. VI. By H. M. Bernarp, M.A.
London: Printed by Order of the Trustees of the British Museum,
1906.
In this volume Mr. Bernard completes the description of the genus
Porites and gives also a descriptive list of the genus Gontopora
supplementary to that given in vol. iv.
Besides the Corals there are probably few groups in the animal
kingdom which present such formidable difficulties to the taxonomist
and morphologist alike—difficulties which at the present moment
appear to defy solution. Mr. Bernard, who has devoted to this
group many years of patient study, has, however, certainly laid the
foundations of a more exact knowledge of coral-forming animals,
and has at the same time brought to light some important evidence
with regard to the effect of isolation and the influence of environ-
ment on living organisms, more especially with regard to sessile
forms.
The Author has, however, so it seems to us, become entangled in
the toils of that seductive question, What is a species? Dissatisfied
with our present definition, and unable to supply any better, he
has endeavoured to compromise. We venture to think, however,
that his suggestion will not be favourably received.
Recognizing a number of local forms in the several genera which
he has so far examined, he proposes to regard these not as so many
geographical races to be distinguished by trinomial specific names,
but as indeterminate incipient species, which are to be distinguished
by numerals. So that we get such names as Goniopora Queens-
landie quintadecima, Goniopora Australie occidentalis septima,
Porites incerte sedis quartadecima !!
However, in spite of this positive drawback, Mr. Bernard has
contrived to bring together a mass of most valuable information,
which will prove of great value not merely to the students of corals,
but also to those who are interested in the study of animal life in
general.
The plates which illustrate the volume are of great beauty.
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XVI. A Contribution to the Knowledge of the eee of the
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XX. Description of a new Frog discovered by Dr. W. J. Ansorge
Be sas eee ve 109
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CockrruLi, University of Colorado............6+. RMA SUE gt 122
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by Plotz:: By #.D.:-Gopman, DiC DPS... Wer es est as 132
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Direct. (Plate Vi). os. cee ey ae uae Pn te Re Ra Re .. 155
XXVI. On Neotropical Mammals of the Genera Callicebus,
Reithrodontomys, Cienomys, Dasypus, and Marmosa. By O.pFreip
Sg CNA pe a i a AT ete uy Sighs airs hese eee 161
BIBLIOGRAPHICAL NOTICE.
Catalogue of the Madreporarian Corals in the British Museum (Natural
History)... Vol. Vi. By H. M. Bexwanp, MCAS Se. ©. 2ocene 168
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XXVII.—Notes from the Gatty Marine Laboratory, St.
Andrews.—No. XXVIII. By Prof. M‘Intosu, M.D.,
LL.D., F.R.S., &e.
[Plates VI.-VIII.]}
. On a Porbeagle Shark with a large Suboral Aperture.
. On the Young of the Ling.
. On a large Example of Ommastrephes sagittatus, d’Orb.
. On Genetyllis citrina, a new Phyllodocid.
. On the Reproduction of Nereis diversicolor, O. F. Miiller.
Ov > CO bD
1. On a Porbeagle Shark with a large Suboral Aperture.
The number of porbeagle sharks caught in the gill-nets of
the fishermen in St. Andrews Bay has been a subject of
remark in former notes, and as time advances they do not
appear to be diminishing. A female 9 feet long and in a very
healthy condition entangled itself in the gill-nets for cod last
November, and as the fishermen stated that it had two mouths,
it was brought to the Laboratory. Inspection showed that
all the internal organs were in a normal state. At a distance
of 5 inches behind the tip of the mandible is a large aperture
(PI. VI. fig. 1,a), with a smoothly cicatrized margin, measuring
in the preserved specimen 4 inches in transverse diameter,
and blocked inferiorly by the basihyal, covered with cicatrized
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 12
170 Prof. M‘Intosh’s Notes from the
skin and freely movable with the branchial apparatus. The
truncated tip of the basihyal is marked by various hard
wrinkles, and it projects from the ventral surface a consider-
able distance in profile (Pl. VI. fig. 3). When the hyoidean
apparatus is pushed forward the blunt tip has a tendency to
close the aperture.
Viewed internally (PI. VI. fig. 2) the basihyal is depressed,
much of it passing into the aperture in the floor of the mouth
and projecting ventrally. The ceratobyals are nearly normal
or only very slightly lowered. In the fresh specimen the
movements of the parts were quite free, as in an ordinary
example, and apparently prehension and deglutition were
unimpaired.
So far as observation goes, it would seem that the fish had at
one time been captured by a powerful shark-hook which had
struck it from below and pierced the basihyal and the soft parts
near it. In its struggles it probably wrenched the tip of the
basihyal, which, with the soft parts, gave way under the powerful
strain, so that about 2 inches of the basihyal with the tip of
the tongue were lost and a large gap externally was caused.
Whether the hook remained a short or a long time in the wound
would depend on the soundnessof the tackle. Though the whole
hyoidean apparatus must have received a considerable wrench,
yet the healing and contraction of the wound and the cicatri-
zation of the coverings of the depressed basihyal have been
so complete that comparatively little inconvenience has
resulted. The large ventral aperture would take in water,
whilst the plug of the basihyal would prevent the loss of
food. Moreover, swallowing would not be interfered with.
The slender gill-nets prove more deadly to such a powerful
fish than a hook, for they yield on every side, and, though
torn, soon envelop fins and tail and impede branchial respira-
tion. On the other hand, it is comparatively rare for such a
form to envelop itself with the lines, from which this shark
is an adept at picking off the fishes after they are hooked.
As indicated elsewhere, the porbeagle shark has never been
known to attack the human subject, yet the great muscular
power and weight of an example measuring 9 feet and the
length and sharpness of its teeth undoubtedly fit it for any
predatory function. If by chance it acquired such a habit,
and was as common in St. Andrews Bay in the warmer
months as it is in the cold, those who swim in the inshore
waters would have to face a new danger.
Gatty Marine Laboratory, St. Andrews. 1 7ae
2. On the Young of the Ling (Molva molva, L.).
J. Schmidt *, as the result of the recent unequalled inter-
national opportunities for obtaining the early stages of the
food-fishes, gives an account of the pelagic postlarval stages
of the ling. Yet he has only procured a single example
5 mm. longer than that described and figured in the
* Researches ’ t, and which really adds little to our knowledge,
since the pigment forming the commencement of the longitu-
dinal band characteristic of the later stage of 34 inches is
already indicated in the memoir cited. The 33-inch stage
has not been procured by the Danish author, his figure having
been taken from the ‘ Researches.’ He seems to be in doubt
as to the passage of this longitudinally striped form into the
transversely barred one found at the rocks near the pier at
St. Andrews, since this coloration is diagnostic of the post-
larval stages of the blue ling (Molva byrkelange, Walb.), a
deep-sea form. Now only one species of ling frequents the
eastern shores of Scotland, the eggs and young of which
were long ago described at St. Andrews. Moreover, there is
little doubt that the longitudinally striped stage of 3} inches
by-and-by changes into a transversely barred one with
blotches along the sides. Moreover, at 13} inches the
coloration remains very much as at the 9-inch stage described
in the ‘ Researches.’ The golden colour of the pectorals and
the pallor of the ventrals are the same, the median division
of the tip of the latter, however, being larger and broader,
The barbel bas additional black pigment. Perhaps the
pigment-bars on the tail are less bold, though the margin is
still white. In general outline the chief change, in com-
parison with the 9-inch stage, is the elongation of the snout
in front of the eyes, the spaces between the eyes and the
nostril and between the latter and the tip of the snout having
increased. This example was procured in May, and is
probably about six months older than the 9-inch stage; and
as the barred condition is assumed in all probability in its
second year, the rate of growth given in the ‘ British Food-
Fishes’ { may ke too rapid.
Schmidt’s hesitation in believing that a longitudinally
striped young ling of 34 inches subsequently becomes trans-
formed into a fish with bold transverse bars is natural.
* Meddel. fra Komm. for Hayunder, Serie Fiskeri, Bd. ii. no. 8
(1906).
+ Trans. R. 8. Edin. vol. xxxv. p. 830, pl. xvii. fig. 4.
{ M‘Intosh & Masterman, p. 283,
172 Prof. M‘Intosh’s Notes from the
There is, however, no reason to doubt it, (1) because no
smaller form with transverse bars is known at St. Andrews ;
(2) because all the examples (and they are many) above this
size are transversely barred, the blotches remaining even to
the size (133 in.) already indicated; (3) because no other
species of ling has hitherto been found in the neighbouring
waters.
The above writer, in criticizing the figure of the early ling
in the ‘Researches ’*, had forgotten to look at the text. The
outline of these delicate young forms is considerably altered
by strong alcohol, and whether a slip had occurred in
Prof. Prince’s figure or in the work of the lithographer it is
now difficult to determine. At any rate, it is clearly stated
in the text that “the notochord passes almost in a straight
line backward to the tip of the tail, and the caudal fin is
continuous with the unbroken marginal fin dorsally and
ventrally. The great development of the ventral or per-
manent rays, however, slightly pushes the tip with the
embryonic radial striations upward. he hypurals, two of
which are very distinct, are developing inferiorly, and the
epiurals dorsally, but they have only slightly affected the
direction of the notochord. The early development of the
upper caudal rays in this form is of interest, as it is in
marked contrast with such forms as the Pleuronectide, in
which the inferior fin-rays alone appear.”’ A re-examination
of the specimen in the University Museum bears out the
accuracy of this description, and shows that the slightly
tapered tip of the notochord projects backward nearly in the
centre of the caudal rays, which are now taking the place of
the embryonic fin-rays. ‘he hypural and epiural elements
are clearly visible. There is nothing in the preparation to
cause any confusion with the condition in a Pleuronectid,
from which the pelvic fins alone would at once distinguish
the young ling.
3. Ona large Example of Ommastrephes sagittatus, d’ Ord.
Information was given by Mr. Andrew Brown, of Queen’s
Gardens, St. Andrews, early in January of the stranding of
a large cuttlefish at the East Rocks, near the Rock and
Spindle. It had, unfortunately, been much destroyed by
birds before it was seen—the tentacles, funnel, all the viscera,
beaks, and even a large part of the mnscles of the mantle
having been removed. A glance at the specimen showed that
* Trans. R. S, Edin, vol, xxxy.
Gatty Marine Laboratory, St. Andrews. 173
it was of unusual size, no similar form having been met with
for forty years, when a somewhat smaller example (mantle
20 inches and tentacles 3 feet) was procured on the beach.
The visits of the swarms of smaller squids which prove so
troublesome to the liners are as uncertain, one having been
noted in 1885 and the last in 1891 *. On the last-mentioned
occasion a vast number took possession of the ground between
the Bell Rock and St. Andrews, entered the latter, and even
filled the salmon stake-nets, though most escaped through
the meshes when the nets were hauled. Very large squids in
British waters appear to be rare, one slightly larger than the
present example having been described by Dr. Goodrich f,
and two are mentioned by him as in the British Museum.
Foreign specimens of great size are well known, especially on
the American coasts {. A gigantic one was seen in 1876,
when a Scotch herring-boat, containing amongst others the
attendant at the Laboratory (AW: Brown), leaving Howth
Point, was summoned by the cries of the occupants of a
diver’s boat. ‘They found the diver, who had been surveying
a sunken tug off St. John’s Point, on the ladder, to which
he had been hauled (as no signal had been made for some
time), with his arms pinioned by a huge cuttlefish. ‘The men
rapidly cut the diver free and allowed the cuttlefish to
escape. ‘Their impression was that the body of the cuttlefish
was about a foot in diameter and the arms about 7 or 8 teet
in length §.
In the present mangled example now in the University
Museum the length of the mantle from the tip of the tail to
the collar is 25 inches. ‘The caudal fin is 11 inches from the
apex to the centre, 11¢ inches along the outer slopes, and
fully 10 inches across the base on each side, though the latter
is probably underestimated, as the central portion had been
lacerated by the gulls. ‘The total breadth of the base of the
caudal is thus more than 20 inches. The peniecles are
absent, but the eight arms have an average length of 13} inches,
and the breadth of the winged or keeled arm is fully 22 inches.
The pen, characterized by its great strength and stiffness, 1s
in fragments, but when laid caretully ‘together it measures
23 inches, though in all probability in its complete condition
* Fourth Ann. Rep. Scotch Fishery Board (1885), p. 204, and Tenth
Report (1891), part ii. p. 299,
+ Journ. Mar. Biol. Assoc. vol. ii. n.s. p. 314 (1893).
{ An interesting account of one of these is given by Prof. D’Arcy
Thompson, Proc. Zool. Soc., Dec. 18, 1900, p. 992.
§ Part of an arm was secured for G lasgow Museum, but Prof. Graham
Kerr, who kindly made a search lately, could find no trace of it.
174 Prof. M‘Intosh’s Notes from the
it approaches 25 inches. The size of the terminal pocket, in
contrast with that of a specimen witha body 15 inches long,
is great, viz. 17 mm. in depth at the proximal edge, and from
that to the flexible tip 20 mm. It is marked by the usual
ridges.
The description of the arms by Dr. Goodrich would do
fairly for the present specimen, the latero-ventral frill of the
third right arm (PI. VII. fig. 1) being conspicuous, and at
its distal third, the widest portion, is at least an inch and
a half in breadth. As mentioned by Dr. Goodrich, it is
supported by thick ridges. Hach arm has two rows of
suckers (Pl. VII. fig. 2), which differ from those described
and figured by Goodrich, but agree with those of the common
Ommastrephes sagittatus, and this is confirmed by Dr. Hoyle,
who kindly examined both the example and its suckers, and
to whom I am indebted for much information. ‘The largest
suckers on the arms measure 15 mm. over all, and the aperture
with the teeth 10 mm. én stu. The proximal half of the rim
(as it lies on the arm) has a series of small teeth, whilst the
distal half has larger recurved teeth, the whole surrounded
by the mobile muscular investment. The central cushion is
attached to the tough pedicle. When removed from its
investment (Pl. VIJ. fig. 3) the horny rim is distinctly
oblique, the edge trending from the large teeth forming a
Jong hoof-like process which apparently gives a larger surface
for its attachment. ‘The arrangement of the teeth on the
rim is more or less uniform (Pl. VII. fig. 3), viz. a median
very large tooth, often more or less blunted by use, and two
or three somewhat smaller teeth on each side, the points of
these being very sharp. Occasionally a smaller tooth occurs
between the lateral teeth of one side. ‘The teeth spring from
a thickened horny rim and are sheathed externally in a hard,
glistening, pale yellow substance, which appears to perform
the part of enamel. So far as could be seen, no fold
supporting horny plates existed round the outer margin. As
the tentacles were absent nothing can be said about their
suckers, but they must have been of large size, and probably
with a more or less even series of teeth, as in the smaller
examples.
In a specimen which has a mantle about a foot in length,
and which has a diameter at its widest part of 3 inches, the
arms range from 7-7} inches in spirit, the third right arm
having a keel, which, however, is but slightly developed in
contrast with the old example. ‘The tentacles are 10-11 inches
in length and bear larger suckers than on the arms. ‘The
length of the caudal fin along its outer edge is 4} inches.
Gatty Marine Laboratory, St. Andrews. io
Such forms a useful contrast with the large specimen, which
seems to have successfully eluded the agencies which, as a
rule, render examples of this size rare on our shores, though,
erhaps, abundant in the deeper waters.
4, On Genetyllis citrina, a new Phyllodocid.
This form was procured on a stone—coated with corallines,
a bight yellow sponge, and several ascidians—brought up
by a fisherman’s hook in the Minch in 1865.
The head is rounded ovate, with two black eyes of con-
siderable size, The four tentacles had been removed, but they
probably resemble those of G.dutea. The tentacular cirri are
similarly arranged, viz., two shorter anteriorly and two
Jonger posteriorly. Body about 3 inches in length, much
tapered anteriorly. Posteriorly it terminates in two caudal
cirri. ‘The dorsal surface is convex, the ventral marked by
two elevated ridges on each side of a slightly depressed central
area. ‘The entire animal is of a most brilliant chrome-
yellow—deepest on the middle third, which here and there
showed blackish-brown patches on the lamelle (cirri). It
tinged the water with a yellowish mucus, and also dyed the
spirit in which it was immersed of the same hue.
The dorsal region of the foot has a massive, short, bluntly
conical process devoid of spine or bristles, and bearing the
unequally cordate lamella (cirrus), which is marked ‘by a
series of lines and reticulations trom a central rib. The
imbricate lamellae are borne more or less horizontally along
the sides of the dorsum, leaving the central region bare.
The semicircular gap at the base of the lamella fits the
rounded extremity of the division to which it is attached.
The short setigerous region is bifid at the tip and supported
by a black spine, and a group of bristles shorter than in
G. lutea, the translucent shafts being slightly bent, and with
a dilated distal end which has a few spikes on each side.
The terminal process is finely tapered and shorter than in
G. lutea. The edge shows no distinct serrations, though the
adherence of particles would indicate them, Attached to the
ventral and posterior part of the region is an_ irregularly
reniform ventral cirrus, vertical in position. The inferior
border is rounded, but the superior is truncated, with a
tendency to a point at the upper and outer angle. The
cirri—both superior and interior—vary litle in shape
throughout the body.
The stone was coated with the yellowish sponge, but its
connection with habits of the Genetyllés is unknown. The
176 Prof. M‘Intosh’s Notes from the
annelid crawled actively amongst the ascidians and other
growths to escape capture.
This form clearly approaches Genetyllis lutea, Malmgren,
but the setigerous region supported by a black spine and
carrying shorter bristles with shorter terminal processes than
in that species point to separation.
5. On the Reproduction of Nereis diversicolor, O. F. Miiller.
Various statements have been made about the reproduction
of this species. Thus Max Schultze * refers to the occur-
rence of the young in the body-cavity, having apparently
considered Koch’s statements in regard to Marphysa sanguinea
as true.
Cuvier and Grube, again, thought Arenicola was herma-
phrodite, and Rathke placed Ampjztrite in the same category,
though each might be more or less male or female. An
interesting résumé of the views of the period is given by
Frey and Leuckart f. A. Krohn, again, describes from Nice
a viviparous Syllis (S. wvipara) allied to S. Armandt, only
the new form has simple tips to the terminal pieces of the
bristles. The enclosed young form has 23 segments and is
like the adult.
In Balfour’s ‘Embryology’ { it is stated that “a few
forms (e. g., Hunice sanguinea, Syllis vivipara, and Nereis
diversicolor) are viviparous.” Considerable reliance in more
recent times has been placed on the observations of Mendthal
on the supposed hemaphroditism of Nereis diversicolor. This
author seems to have been attracted to the subject by the
previous work of Schréder§, who found embryos in the
morula-stage. Mendthal described the species from the bay
of Pillau as carrying both eggs and testes in June, and he
was the more inclined to believe in its hermaphroditism since
Schréder met with only 3 males out of 48 examples. He
places and figures the testes in the dorsal region as pear- or
flask-shaped masses to the exterior of the dorsal longitudinal
muscles. The supposed testes, therefore, occupy the same
position as the outer limb of Dr. Goodrich’s dorsal organ, and
it may be that the cilia were a source of misapprehension.
On the other hand, he describes the ova as developing at the
bases of the feet, the figure representing them in the position
of the segmental organ.
* ¢Entwickelung Arenicola piscatorum &c.,’ p. 214 (Halle, 1856).
+ Beitr. z. Kenntniss wirb. Thiere, p. 82 (1847).
ft Volar ole:
§ This author’s paper has not yet been obtainable.
Gatty Marine Laboratory, St. Andrews. Lay
De St. Joseph, however, could not corroborate the fore-
going observations. Gravier, on the other hand, considers
that hermophroditism and viviparity may exceptionally exist
in this form, and in the ‘Cambridge Natural History ’ it is
stated that viviparity is characteristic of it.
The annelid is very common on many parts of the British
shores, burrowing in clay, sand, mud, peat, and similar media,
and at St. Andrews it abounds at the upper end of the
harbour amidst muddy clay on the sides of the Kinness Burn
which enters there. During last winter especially careful
observation of the species was carried out to test the condition
as regards reproduction ; yet though both hermaphroditism
and viviparity have been discountenanced, it has not been
possible to find the worm either discharging from its tunnel
in the mud its eggs and sperms or becoming pelagic at
maturity and thus dispersing the sexual elements. So far
as can be observed, the former, perhaps, seems to be the more
likely, though no certainty exists on this head.
In the middle of October the majority of the examples—
both large and small—were females with fairly developed
ova, which were almost visible to the naked eye and easily
under a lens. In these the vascularity of the feet had slightly
increased, but no change in the lobes of the feet or in the
bristles had occurred. Amongst the ova in the ccelomic space
were numerous pale granular cells, apparently moditied
perivisceral corpuscles. Very few males were obtained at
this time, and these for the most part were undeveloped.
In November comparatively few ova were attached to the
ventral plexuses, most being free in the perivisceral space.
On the other hand, the vascular plexuses in the region of the
ciliated organ were laden with dense groups of rather coarsely
granular cells—sometimes in lobular masses, and it was con-
sidered that these pale cells were associated with the growth
and maturation of the ova in the ccelomic space. ‘The ova
had considerably increased in size towards the end of the
month.
In transverse section* the body of the annelid at this
time differed little from the type except in the presence of
ova, which appear most abundantly, in the sections, at the
bases of the teet and extending into their lobes. The number
in the coelom was not large, many, in all probability, having
fallen out. ‘The dorsal and ventral longitudinal muscles
showed little change, the pennate fold of the latter being well
* T am indebted to Dr. Tosh for valued aid in making these sections
and in other respects.
178 Prof. M‘Intosh’s Notes from the
developed. The oblique muscles pass at each side slightly
below the nerve-cord to be attached to the basement-mem-
brane; and at least three neural canals are present, the larger
lateral having a coagulable fluid internally, and each runs
along the outer border of the nerve-trunk between the pairs
of ganglia. ‘The median or dorsal canal, again, appears to
have a separate strand in the interganglionic region, and
when it reaches the ganglia it splits into two trunks in certain
sections. The muscles of the bases of the feet and of the
bristles are also strong. The wall of the alimentary canal
appears to be of normal thickness.
The annelids remained very much in the same condition
during December, the great majority of those examined being
females, so much so that it was at first considered probable
that, as authors had stated, a complex sexual condition
existed (e. g. alternate development of the reproductive
elements); but as a few males were still met with and no
trace of an intermediate stage occurred, such complexity
could not be proved. This month the only difference in the
sections of the females was the increase in the size of the ova,
the abundance of the coelomic corpuscles (PI. VILI. fig. 1),
and the distention of the body-cavity and the bases of the feet,
so that the muscles of the wall were stretched. Thevascularity
of the outer surface of the gut also appeared to be increased.
The segmental organs showed no feature of note. Inthe males
the perivisceral cavity and the bases of the feet contained
dense masses of translucent granular cells, the large nuclei of
which stained deeply with eosine.
The great increase of the large granularcells (PI. VILL. fig. 3)
in the ceelomic cavity in January was a feature of moment,
especially in those females in which the ova were small.
The enormous masses of these cells distended the bodies of
the females and they probably increased by division, each
being filled with spherules. In glancing at the living
annelids a pale, or greyish, green hue characterized the
posterior region of the body in the females; whilst the
anterior segments had their vascularity increased, the dorsal
vessel of the foot and its branches especially were distinct.
When the posterior region was punctured, the masses of ova
had a pale greenish colour as in the previous months, and
were similarly unfertilized. As many have discharged ova,
it would appear that fertilization is external as in allied
forms.
The sexes are not always distinguished by colour, both
males and females being greenish or dull yellowish, though
the males are often paler. No change in eyes, feet, or
Gatty Marine Laboratory, St. Andrews. 179
bristles is apparent. In many an opaque, dull whitish con-
dition occurs in the anterior feet, which are filled with the
large granular perivisceral corpuscles. Moreover, almost all
have a touch of white at the base of the dorsal cirrus—from
one end of the body to the other, a similar touch occurring
at the tip of the ceratophore of each tentacular cirrus.
Mendthal* figures two examples, one of the general
brownish-orange hue of the ordinary type, and the other of
this colour in the anterior fourth, whilst the rest is of a grass-
green tint. He considers that the latter (green) colour is
due to a diet of green alge, a condition which is not in
accordance with the observations at St. Andrews, where the
greenish hue appeared to be the result of the presence of
masses of more or less ripe ova. There is no reason, how-
ever, why adiet of Ulva or other green alga should not tint
the digestive canal green.
In the male the sperm-cells (PJ. VIII. fig. 2) filled the
various cavities even to the bases of the dorsal cirri through-
out January, but though the body-wall was frequently
distended, no degeneration of the muscular bands could be
observed. In the females the size and abundance of the
ova in particular examples varied, but, as a rule, the granular
ova with a slightly stained nucleus and a more deeply stained
nucleolus were more or less advanced towards maturity.
Occasionally an example with few and small ova appeared.
In some of these the muscular walls of the body were con-
tracted, and very few ova, and these of small size, occurred
in the perivisceral space. More frequently they were found
in the feet. Masses of ovigerous tissue were attached to the
vessels near the bases of the latter; and cellular masses (it
may be parts of the dorsal organ) with boldly stained nuclei
passed upward at the bases of the feet within, as well as
without, the lateral lobe of the dorsal longitudinal muscles,
and similar cells passed into all the spaces of the feet—in
specimens having a considerable number of large ova. ‘These
loose coelomic masses probably represented the cells alluded
to in the living forms.
In February the increase in size and the division of the
sperm-cells formed the most noteworthy feature, but no free
sperms were seen. In the other sex the ova in many had
considerably increased in size, and from the appearance
of the annelids a large number of ova must have been
discharged.
* Untersuch. u. d. Mollusken u. Anneliden des frischen Hatfts, p. 9,
Taf. figs. 1 & 2 (Konigsberg, 1889).
180 Prof. M‘Intosh’s Notes from the
Signs of rapid division of the sperm-cells characterized
many males in the middle of March, and towards the end
of the month the perivisceral cavity formed a vast reservoir
for the male elements. The body-walls were stretched,
the muscular layers diminished, and the alimentary canal
collapsed. The tubes of the segmental organ had also en-
larged, so that sperms could readily find issue by the
segmental papilla. Though the nuclei of the walls of the
organ and its tubes were everywhere distinct, no trace of
sperms was at any time found within it, so that they probably
escape by rupture of the body-wall, as in allied forms.
The ova in the various examples throughout March
appeared to attain full size and maturity, viz. about "1524 mm.,
and they were probably shed by rupture of the body-wall,
the vast numbers set free, even from a limited area, affording
an indication of the almost illimitable resources of nature in
the sea. Whether the sexes discharge their elements i sctu
or by a terminal pelagic stage could not be ascertained.
None were captured in the tow-nets dragged, by day and by
night, up stream at ebb-tide, and no indication of any change
in the structure of the feet, bristles, or eyes, and no shrivelling
of the posterior region of the body as in the very interesting
Ceratocephalus Osawat of Akira Izuka*, common in the
estuary of the Sumida River in Tokyo, occurred. If one
may hazard a conjecture, it is probable that the sperms and
ova are discharged on the sites inhabited by the annelids, and
the larve by-and-by carried seawards. ‘There is no doubt,
however, that, without change in the condition of the feet
and bristles, the species is an apt swimmer, progressing
through the water swiftly in graceful screw-coils.
Towards the end of May (29th) and for some time pre-
viously signs of degeneration appeared in certain ova, as if
they were in process of absorption. ‘These ova were smaller,
minutely granular, and with larger oil-globules. The larger
ova had lost the germinal vesicle and spot, and transmitted
light more readily than formerly. Nothing was seen to
suggest the view that certain ova were undergoing develop-
ment, for, when kept in vessels both of sea-water and fresh
water, they were rapidly disintegrated.
Some females at this date have shed all their ova, and are
of a pale brownish-yellow colour, occasionally with a minute
dusting of yellow grains along the dorsum.
Having failed to secure the early larve by any of the
methods alluded to, masses of the clayey mud with the adults
* Journ. Coll. Sc. Univ. Tokyo, vol. xvii. art. ii, (1902).
Gatty Marine Laboratory, St. Andrews. SI
én situ were brought from the banks of the Kinness Burn
near the harbour towards the end of May, and a strict scrutiny
made of the tubes and the mud lining them. Numerous
postlarval forms were thus obtained, but no trace of trocho-
phores or other early stages. Whether these stages had been
passed in such an environment before the examination, or
whether the later larve to be subsequently described had
settled in the mud of the tubes after a pelagic stage is yet
undetermined,
The youngest stage observed has three bristled segments
(Pl. VIII. figs. 4 & 5), a head with two short palpi, and two
short frontal tentacles. The eyes are imperfectly differentiated,
consisting of an irregular group of black pigment-granules
(fig. 5). A tentacular cirrus occurs on each side opposite
the eyes. The three feet are nearly alike, each with a bristle-
tuft and a small dorsal cirrus, or the first is rudimentary
(fig. 4). A minute caudal cirrus is at each side of the posterior
end. Between the last foot and the pygidium is a projection,
indicating a segment. The proboscis has a pair of jaws each
with three long teeth, including the anterior fang (Pl. VIII.
fig. 6). The opaque part of the gut extends over the last
two bristled segments. ‘he minute bristles already present
the typical structure of camerated shaft and homogomph
articulation of the end of the shaft, and are in two bundles in
each foot.
No younger form has yet been found amongst the mud or
the adults, so that it is probable that they settle down at this
stage. Moreover, no pelagic larve appeared in the vessels;
yet as the postlarval forms with three bristled segments
occurred in the mud of the tubes of the adult, it would appear
that all do not wander.
When the postlarval form has four bristled segments
the head has two eyes on each side, placed close together
and posteriorly, a pair of short frontal tentacles, short
palpi which present no distal articulation, a pair of tentacular
cirri, and two short anal cirri. Behind the head, which
bears the tentacular cirri, is a region with only bristles on
each side; a foot with a large bristle-tuft and a minute
dorsal cirrus follows, and then a second foot of similar
structure. A rudimentary foot comes next, with a bristle-
tuft on one side. A rudimentary ventral cirrus occurs on
each foot. A pair of minute jaws having two teeth behind
the anterior fang is found in the proboscis. Vast swarms of
Infusoria (like monads) frequent the moist and odoriferous
mud in which the young Nereids occur.
The postlarval form of the 28th May (Pl. VIII. fig. 7)
182 Prof. M‘Intosh’s Notes from the
has a head with two comparatively large frontal tentacles
and two stumpy palpi with very short terminal segments
that only occasionally are visible, two pairs of tentacular
cirri, the last with a spine in the dorsal, five pairs of distinctly
bristled feet, with more evident dorsal and ventral cirri and
slightly brownish spines, besides two rudimentary ones
behind and a large pygidium with two caudal cirri. The
cirrus of the first bristled segment is elongating, but the
bristles are shorter than in the succeeding foot. The mouth
is a wide aperture, followed by the somewhat ovoid proboscis,
which is armed with two translucent jaws having three
prominent teeth which are proportionately longer than in the
adult. ‘The proboscis is narrowed posteriorly and joins a
large opaque, yellowish, glandular region, broad and truncated
in front and diminishing posteriorly and terminating in the
pale rectal part of the gut and its dorsal anus. ‘The opaque
glandular region stretches from the space between the second
and third feet to the last bristled foot, and shows the groups
of oily granules so characteristic of the glands on the walls of
the alimentary canal. A pair of slightly opaque ovoid
glandular bodies, the segmental organs, lie behind the first
complete foot, just in front of the opaque region of the gut,
and active ciliary action is occasionally noticed in them.
In the postlarval form with six bristled feet the eyes are
better defined and the palpi, frontal tentacles, and tentacular
cirri are larger. The dorsal cirrus of the first bristled seg-
ment is the longest in the animal, but there isno spine. The
spines are darker, the posterior end remains bifid, and three
teeth occur behind the anterior fang in each jaw.
When seven bristled feet are present the caudal cirri are
considerably longer, the last foot (seventh) having a few
short bristles, a short dorsal cirrus, two short spines, and a
minute rudiment of a ventral cirrus. Behind is an indication
of the eighth foot as a lateral projection, with a minute
papilla representing the dorsal cirrus. Internally is the tip
of a minute spine, but no bristles. The ventral of the first
tentacular cirrus (opposite the eyes) is a short subulate
process; that of the next segment is indistinct, though, with
the exception of the caudal cirri, the dorsal is the longest of
the series. The bristles of this segment are always short and
in contrast with the next segment, which shows the segmental
organs immediately behind. Opacities in the corresponding
parts of the following feet indicate the early development of
these organs throughout. ‘Traces of a fourth tooth behind
the great anterior fang of the maxille are visible. This
description might also apply to a postlarval form with eight
Gatiy Marine Laboratory, St. Andrews. 183
bristled feet (Pl. VIII. fig. 8), the latest stage observed
towards the end of May.
If the young Nereids of the foregoing stages are left in the
vessels beside the adults the larger examples rapidly disappear.
They are probably devoured by the adults, only the minute
stages escaping capture, and in all likelihood they, too, would
by-and-by be found out and captured ; yet, as in the fishes,
these checks have little influence on the permanent abundance
of the species.
On the 17th July the young forms are found on the same
sites, but considerable progress has been made in develop-
ment. ‘The head in outline nearly resembles that of the
adult and is marked by whitish pigment in front of the eyes
in the smaller, brownish in the older, which are definitely
formed on each side, the anterior pair being somewhat larger
and furnished with lenses. The frontal tentacles and the
palpi are well formed and show numerous fine palpocils, the
mobility of the distal segment of the palpi being noteworthy.
The two pairs of teutacular cirri are now much longer, the
dorsal stretching outward like fine hairs as the animal pushes
its snout forward. They retain the proportions to each other
of the adult, though they have not yet reached full develop-
ment in any case. The ventral of each is considerably
shorter than the dorsal, but projects clearly on each side, the
first rather longer and thicker at the base than the frontal
tentacles. The body is now elongated, having from sixteen
to twenty bristled feet, and is marked along the centre by a
yellowish or brownish streak, often slightly moniliform or
zigzag from the intestinal contents, and a whitish dot is
observed in some at the bases of the feet. The massive and
somewhat ovoid proboscis has six teeth behind the great
anterior fang, and the narrow part of the canal behind it is
usually firmly contracted (and empty). The first part of the
gut behind the post-proboscidian narrow region is the largest,
and for some distance it is not marked by lateral constrictions,
as in the succeeding portion. Most of the feet anteriorly
present a condition approaching that of the adult, though the
dorsal cirrus is shorter. Only the last two feet are devoid of
bristles externally. The caudal cirri are now considerably
longer. The circulation of the red blood in the dorsal and
ventral trunks is now evident. The segmental organs
extend backwards to the posterior feet, though not quite to
the tail.
So far as observed at St. Andrews, therefore, there is no
foundation for the statement that the Scotch representatives
are hermaphrodite, and still less that they are viviparous, as
184 Notes from the Gatty Marine Laboratory.
mentioned by Max Schultze, by the ‘ Cambridge Natural
History,’ and by Gravier. Max Schultze * gives a circum-
stantial account of finding female examples in April at
Greifswald with ciliated eggs and pear-shaped larvee in the
body-cavity, and he figures two of the latter, hundreds of
which occurred in the cavities at the bases of the feet. His
figures, however, give rise to doubt as to the nature of the
ciliated forms, which have the narrow end of the pear
anteriorly with the two eyes at some distance behind it.
In one, indeed, the mouth is indicated behind the eyes as a
small radiated disk. The posterior end of the larva is broad
and rounded. In the light of the condition at St. Andrews
doubt arises as to correctness of the interpretation, more
especially in view of the structure of the larva; but it is
right to give a margin for possible variations in regard to
internal fertilization under certain circumstances. Yet it
seems unlikely.
The foregoing interpretation as to the mode of reproduction
would seem to be most in accordance with observation, since
on the 28th May masses of clayey mud brought from the
habitat of the species and with the annelids cm stu showed
many free ova amongst the mud coating the walls of the
tube, and, further, of various postlarval examples.
EXPLANATION OF THE PLATES.
PLATE VI.T
Fig. 1. Ventral view of the head of a porbeagle shark, 9 feet long,
showing the rounded tip of the hyoidean apparatus (a) pro-
jecting from its smoothly finished aperture.
Fig. 2. Aspect of the same region from the interior of the mouth.
ig. 8. Lateral view of the preparation, indicating the projection of the
hyoidean boss.
N.B.—The first figure is drawn to a larger scale than figs. 2 and 3,
which are nearly equally reduced.
PuaTeE VII.
Fig. 1. Third right arm of the large Ommastrephes sagittatus, d’Orb.,
showing the keel to the left, the web with its supporting rays,
and the suckers. A portion of the web has been removed.
Slightly reduced.
* ‘Entwickelung von Arenicola piscatorum nebst Bemerk. iiber Ent-
wickel. and. Kiemenw. 1856, p. 214, Taf. ix. figs. 11 & 12.
+ I am indebted to the Carnegie Trust for these figures, those of
Pl. VIL., and four of Pl, VIII.
On apparently new Mammals. 185
Fig. 2. Inner face of the same arm with four suckers. Slightly enlarged.
Fig. 3. Horny rim of a sucker, showing the arrangement of the sharp
teeth, usually on the distal margin of the rim. Enlarged.
Puate VIII.
Fig. 1. Masses of perivisceral corpuscles near the bases of the feet of
Nereis diversicolor, O. F. M., 7th December, 1906, x Zeiss
oc. 2, obj. F.
Fig. 2. Male elements. Similarly magnified.
Fig. 3. Large corpuscles floating freely in the ecelomic space, 21st J anuary,
1907. x Zeiss oc. 2, obj. F.
Fig. 4. Postlarval form of 28th May, 1907, with three bristle-tufts (one
rudimentary).
Fig. 5. Another of same date with three bristle-bundles.
Fig. 6. Mandible of the foregoing. x Zeiss oc. 2, obj. D.
Fig. 7. Postlarval form of 28th May with tive bristle-bundles. Magnified,
Fig. 8. Postlarval form of 28th May with eight bristle-bundles.
Maguified.
XXVIII.—Descriptions of apparently new Species and Sub-
‘ species of Mammals belonging to the Families Lemuride,
Cebide, Callitrichidee, and Cercopithecidee in the Collection
of the Natural History Museum. By D.G. Evutot, D.Sc.,
F.R.S.E., &e.
Havine for some time been engaged in the study of the
Primates, it was found necessary to investigate the material
contained in the various great museums in the Old World,
and, beginning with the vast collection of the Primates con-
tained in the Natural History Museum, London, my friend
Mr. Oldfield Thomas, Curator of Mammalogy in that institu-
tion, not only most kindly gave me every facility for pursuing
my studies, but also requested that I would describe any
specimen that I found in the collection that 1 considered
might be new. In various genera the Museum is very rich
both in number of species and examples, and the advantage
one possessed in working with such splendid material is
exemplified in the comparatively large number of new forms
contained in this paper. My thanks, therefore, are especially
due to Mr. Thomas for the opportunity, not only of examining
the great collection under his care, but of making known to
mammalogists the various forms that seemed worthy of
special recognition.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 13
186 Dr. D. G. Elliot on apparently new
Family Lemurida.
Genus GALAGO.
Galago zuluensis, sp. n-
Type Iceality. Zululand, East Africa.
Gen. char. Similar to G. Hindet, but browner and with
much larger ears ; tail darker and shorter. Skull one third
larger than that of G. Hindet, teeth larger.
Colour. Head and upperparts broccoli-brown and grey
mixed, darkest on the head; outer side of limbs wood-brown ;
dorsal line washed with Mars brown; underparts and inner
side of limbs yellowish white; hands and feet greyish brown ;
tail above pale Mars brown, beneath paler ; ears black.
Measurements. otal length about 570 mm.; tail 320;
ears 31 (skin). Skull: occipito-nasal length 70; hensel 57 ;
zygomatic width 463; intertemporal width 19; palatal
length 28; breadth of brain-case 33; length of nasals 19;
feneth of upper molar series 29; length of mandible 46 ;
length of lower molar series 23.
B.M. no. of type 94. 6. 29. 1.
This animal is of about the same size in head and body-
length as G. Hinde7, but has a considerably shorter and
much darker tail. When the two are placed side by side
G. Hindei appears like a grey animal in comparison. The
ears of the present form are about one third larger both in
length and width. In their dried and shrunken state they
measure 31 mm. in length and 27 mm. wide, while those of
G. Hindet are 24 mm. by 22 mm. respectively. There is
such a vast difference in the size of the skulls and teeth that
they hardly admit of comparison. In colour G. zuluensis is
much like G@. crassicaudatus, but has a much smaller skull, it
being midway between G. crassicaudatus and G. Hindet,
with all the differences such a disparity of size would create.
Galago Hindei, sp. n.
Type locality. Kitui, Athi River, British East Africa,
Altitude 3500 feet.
Gen. char. Size large, colour pale; ear small ; tail very
long. Smaller in total length than either G. crasstcaudatus
or G. Garnetti.
Colour. Head and upperparts pale wood-brown, washed
on head and dorsal region with darker brown ; arms and
hands like head; outer side of legs isabella-colour ; feet dark
Species and Subspecies of Mammals. 187
brown; chin vinaceous cinnamon ; rest of underparts white ;
tail above pale wood-brown, beneath whitish.
Measurements. Total length about 575 mm.; tail 370;
hind foot 86; ear 39. Skull: occipito-nasal length 63;
hensel 49; zygomatic width 42; intertemporal width 20;
palatal length 23; breadth of brain-case 29; length of
nasals 19; length of upper molar series 19; length of man-
dible 41; length of lower molar series 20.
B.M. no. of type 1. 5. 6. 2.
The skull of this form is considerably smaller than that of
either G. crassicaudatus or G. Garnettt. In colour it differs
in being paler and in the very long, pale, almost white tail.
Two specimens are in the Museum collection varying slightly
in colour, the paratype having unfortunately lost half its tail.
Galago gabonensis Batesi, subsp. n.
Type locality. Como River, Gaboon, West Africa.
Gen. char. Similar to G. gabonensis, but much darker
above and has a black tail and light grey feet, and from
G. Allen? it is distinguished by its black tail and grey feet
and legs below the knee.
Colour. Forehead, base of ears, cheeks, stripe between eyes,
and nose light grey ; top of head and hind-neck and upper-
parts dark mummy-brown ; outer side of arms dark tawny; a
patch of tawny on thigh, the upper portion darker than the
lower; rest of legs brownish grey, becoming clear grey on
feet ; throat and front of neck yellowish, rest of lower parts
whitish; hands greyish mummy-brown; tail seal-brown,
sprinkled with grey on basal half. Ears large, blackish.
Measurements. Total length 470 mm.; tail 250. Skull:
occipito-nasal length 48; hensel 38; zygomatic width 32;
intertemporal width 18; palatal length 19; width of brain-
case 24; length of nasals 138; length of upper molar series 16 ;
length of mandible 31; length of lower molar series 14.
B.M. no. of type 96. 10. 9. 4.
While this race hasa general resemblance to both G. Alleni
and G. gabonensis, it can readily be distinguished from both:
by its grey legs and feet from G. Allend, and from G'. gabon-
ensis by its grey feet, darker upperparts, and black tail.
The two forms come together on the Como River, but there
are no intermediate specimens.
Galago braccatus, sp. n.
Type locality. Mount Kilimanjaro, Kast Africa.
Gen. char. Similar to G@. gallarum, but darker grey above,
%
188 Dr. D. G. Elliot on apparently new
and the bright buff of the limbs ends abruptly on meeting
the grey colour, and does not grade into it as in the allied
species.
Colour. Head and neck buff, the hairs tipped with black,
giving to these parts a grizzled appearance; rest of upper-
parts iron-grey ; orbital ring black; stripe between eyes,
nose, upper lip, neck, and chin grey ; outer side of arms and
legs buff; hands and feet yellowish grey; inner side of
thighs and underparts yellowish white; tail dark Prout’s
brown, hairs tipped with white; ears large, naked, black.
Measurements. Total length about 480 mm.; tail 300
(skin). Skull: occipito-nasal length 45; hensel 32; zygo-
matic width 29; intertemporal width 19; palatal length 15;
width of brain-case 24 ; length of nasals 12; length of upper
molar series 13; length of mandible 26; length of lower
molar series 13.
B.M. no. of type 2. 11. 5. 1.
This rather handsome species was obtained by Mr. A. B.
Percival on Mount Kilimanjaro, East Africa. While allied
to G. gallarum, Thomas, it is easily distinguished from that
species by its dark grey colour and the abruptness with which
the buff and grey come together on the legs, As is to be
expected of an animal dwelling at a high altitude, the fur is
thick and long. ‘There are no perceptible differences in the
skulls.
Galago nyase, sp. n.
Type locality. Mountains south of Lake Nyasa, Central
Africa.
Gen. char. Fur woolly; tail bushy; skull, though much
broken, exhibits great differences from that of G. sennar-
ensis, ranging from the White Nile south to Ankola west of
the Victoria Nyanza. The rostrum is long and more slender,
the nasals long and narrow ; the palate is long and narrow,
and not so wide posteriorly as in the skulls of G. sennar-
ensis ; the anterior line of the orbit is in front of m' instead
of in front of pm’, as seen in the other; there is only a slight
rise of the frontal above the rostrum, thus causing the superior
outline of the skull to be flatter.
Colour. General hue above broccoli-brown ; outer side of
arms broccoli-brown; legs cream-buff; chest cream-buff ;
underparts and inner side of limbs yellowish white.
Measurements. Total length about 355 mm. ; tail 185 (skin).
Skull: from frontal suture to end of nasals 27; length of
nasals, median line, 10; width of rostrum at canines 7 ;
length of palate 15; width between last molars 7; length of
Species and Subspecies of Mammals. 189
upper molar series 13; length of mandible 24; length of
lower molar series 13.
The type and an example in alcohol from Zomba, Nyasa-
land, are the only representatives of this species in the
collection.
B.M. no. of type 64. 12. 10. 15.
While perhaps this species resembles G. gallarum in colour
more than any other, the skull, in its long and narrow rostrum
and low crown, is very different. The type was procured by
Dr. Kirk, when he was accompanying Dr. Livingstone, the
famous African explorer.
Subgenus HemiGaLaGo,
Galago Thomast, sp. n.
Type locality. Fort Benin, Semliki River, Central Africa.
Gen. char. Larger than G. Demidofi; colour quite
different. Skull much larger; differently shaped brain-
case, much broader in occipital region and higher over roots
of zygomata ; teeth much larger.
Colour. Head and upperparts drab, washed with Mars
brown on head and dorsal region ; stripe between eyes and
nose yellowish white; outer side of limbs drab; underparts
and inner side of limbs buff; tail Mars brown.
Measurements. Total length 347 mm.; tail 210; hind
foot 58; ear 28. Skull: occipito-nasal length 40; hensel 28 ;
zygomatic width 25 ; intertemporal width 16; palatal
length 14; breadth of brain-case 21; length of nasals 12 ;
length of upper molar series 11; length of mandible 22;
leneth of lower molar series 11.
B.M. no. of type 6. 12. 4. 58.
This, the fourth member of the subgenus Hemigalago,
differs from all the others in colour and dimensions, being the
largest of all, and cannot well be compared with or mistaken
forany of them. The skull shows many and great differences
from those of the other species. The type was procured on
the boundary-line of Uganda and the Congo Free State, and
a second and somewhat darker specimen at Dumo, Uganda.
Whether it penetrates farther into the Congo Region or is
confined to Uganda is unknown.
Family Cebidz.
Genus AOTUS.
Aotus boliviensis, sp. n.
Type locality. Province of Sara, Central Bolivia.
Gen. char. Similar in colour to A. Azare, but cranial
190 Dr. D. G. Elliot on apparently new
characters quite different. Orbits wider and higher ; orbital
portion of frontal bulging outward, forming a decided curve
from the frontal to the nasals; nasals longer and wider ;
brain-case longer and narrower posteriorly ; extreme width
across orbits much greater ; superior outline of skull much
less curved; angle of occipital region much less; teeth
larger, particularly the last upper molar; space from poste-
rior edge of foramen magnum to interparietal much greater ;
basioccipital between bull much wider.
Colour. Similar to A, Azare, but more tinged with red on
upperparts ; a white spot near each eye extending back upon
the head; a broad black line on middle of forehead from
nose to between ears and a narrow black line from corner of
eye on each side of the head bordering the white spot ; upper-
parts mixed iron-grey and russet, becoming more brownish
on lower back; cheeks and chin white; sides yellowish
brown; inner side of limbs and underparts pale orange-
ochraceous ; hands and feet dark greyish brown ; tail mixed
ochraceous rufous and black on basal half, remainder black.
The hairs of tail are all ochraceous rufous at base, and this
shows more or less throughout the entire length.
Measurements» Total length 720 mm.; tail 400; hind
foot 100; ear 35. Skull: total length 64; occipito-nasal
length 61:5; hensel 44; zygomatic width 40:5; inter-
temporal width 33; extreme width of orbits 45; height of
orbits 21; length of nasals 12; width of brain-case 35 ;
distance from foramen magnum to interparietal 113 width of
basioccipital between bulle anteriorly 3-5; length of upper
molar series 14; length of mandible 41; length of lower
molar series 16.
B.M. no. ot type 7. 8. 2. 9.
While similar in colour to A. Azare, the present species
differs greatly in its cranial characters. The skull is much
larger and the brain-case considerably longer, while the
orbits are enormous, very large even for these big-eyed
animals. Two specimens (a male and female) were obtained
by Mr. J. Steinbach in the Province of Sara, Central
Bolivia.
Genus SAIMIRI.
Saimiri macrodon, sp. n.
Type locality. Copataza River, Ecuador.
Gen. char. Similar to S. sciurea, but hands and feet much
darker. Skull has a much higher and narrower brain-case,
much wider palate, larger teeth, with the external line of the
Species and Subspecies of Mammals. 191
upper tooth-row much more curved; zygomatic arch wider
and intertemporal width greater ; bulla narrower and longer.
Geogr. distr. Upper waters of the Amazon in Ecuador, and
Peru.
Colour. General colour like 8S. sczurea, with the back
darker, that of the type being tawny and black on the dorsal
region; golden yellow and black on the flanks ; arms above
elbows dark grey washed with yellow; legs paler; underparts
yellowish white ; the forearms, hands, and feet tawny ; head
and tail like S. sciurea.
Measurements. Size similar to S. sevurea. Skull: occipito-
nasal length 64°5 mm. ; zygomatic width 43; intertemporal
width 32; length of nasals 11; width of brain-case 36 ;
height of brain-case above zygomata 35; palatal length 19;
length of upper molar series 16; width of palate between
canines 12; width of palate between last molars 13.
B.M. no. of type 80. 5. 6. 15.
While the general colour of this animal resembles that of
S. sciurea from the east coast of South America, it is at once
noticeable by its much darker forearms, hands, and feet.
The greatest differences, however, between the two forms are
exhibited in the skulls and in the large teeth of the present
species. The brain-case has quite a different shape, being
long and narrow, with an elevated forehead sloping rapidly
downward to the occiput, which is narrow and rounded. The
palate is wider throughout its length; the teeth much larger,
the canines longer and stouter. Several examples were
obtained in the type locality and others from the Jurua River,
a tributary of the Amazon, and from Marcapata, Peru.
Genus CALLICEBUS.
Callicebus usto-fuscus, sp. n.
Type locality. “ Brazil.”
Gen. char. Allied to C. cupreus, but much darker in
colour; teeth much larger; palate longer and narrower ;
brain-case wider; space between pterygoid processes and
bulle and the width of basioccipital throughout its length
greater. Practically the skull is larger in every way and
more massive. Mandible longer and heavier, and the depth
of the ramus greater.
Colour. General hue above burnt-umber, the hairs being
slaty-grey at base, then annulated with two bands of slate-
and two of clay-colour, and a dark tip. Face naked, black ;
top of head a mixed dark ochraceous rufous and black, the
192 Dr. D. G. Elliot on apparently new
black predominating on the forehead; the rump is redder
than the back and is a burnt-sienna on the outer side of the
limbs; hands and feet claret-brown ; sides of head, throat,
inner side of limbs, and underparts maroon; basal third of
tail black, the hairs being chestnut with broad black tips,
rest of tail mixed black and yellowish grey or very pale clay-
colour, the underside of tail being almost altogether clay-
colour; ears black.
Measurements. Size about the same as C. cupreus. Skull:
occipital region has been cut away; intertemporal width
32 mm.; zygomatic width 41; palatal length 21; width
between last molars 12; breadth of brain-case 35; length of
nasals 9; leneth of upper molar series 14; length of m’ 5;
length of mandible 42; extreme height of mandible 35;
length of lower molar series 17°5.
BM. no; of typed. 7.351.
This species is nearest C. cupreus, but is altogether different
in colour and darker in all its hues. The skulls also are not
at all in accord, the differences mentioned being very con-
spicuous when they are compared. The unique example has
no history beyond the statement that it came from Brazil.
Callicebus subrufus, sp. n.
Type locality. Pachitea, Ucayali River, Peru. Altitude
400 to 500 feet.
Gen. char. Allied to C. leucometupa, but colour entirely
different.
Colour. Face black; a narrow black bar on forehead
above eyes, succeeded by a broader one of white; rest of
head on top, neck, and entire upperparts bright russet,
becoming darker and more reddish on the rump, the hairs
being slate at base, then alternately ringed with slate and
bright russet, or on the rump with slate and dark russet or
reddish ; arms to elbows and thighs to knees grey, the hairs
being seal-brown at base, then russet, and tips grey, this
colour overlying the rest ; hands and feet and rest of limbs
bright chestnut-red ; sides of face, whiskers, inner side of
limbs, throat, chest, and middle of abdomen bright chestnut-
red ; fingers and toes yellowish grey ; tail, basal third black,
with chestnut hairs mixed with black at the root, remainder
light grey above, whitish beneath; hair on ears white.
Measurements. Size about equal to that of C. leucometopa.
Skull: occipito-nasal length 51 mm.; hensel 40; zygomatic
width 35; intertemporal width 29; palatal length 18;
breadth of brain-case 33; nasals broken; length of upper
Species and Subspecies of Mammals. 193
molar series 16; length of mandible 35; length of lower
molar series 16.
BM. no. of type 42.7, (22:
While allied to C. leucometopa, the great difference in
colour the present species exhibits makes it easily recog-
nizable. In its yellowish-grey fingers and toes it shows a
leaning towards C. ornatus, but in other respects it has no
resemblance to that species.
Genus LAGOTHRIX.
Lagothriv lugens, sp. n.
Type locality. Mountains 2° 20’ north of Tolima, Colombia.
Altitude 5000 to 7000 feet. .
Gen. char. Body stout, heavy, as in L. lagotricha, but
colour very different; fur thick, woolly; limbs moderately
long ; tail very long and very broad at base.
Colour.— Male. Head, arms, and body dark purplish
brown, almost black; legs and tail blackish brown washed
with grey, the hairs being blackish brown at base, then black
and tipped with grey or yellowish; breast reddish chestnut,
rest of underparts black.
Measurements. Size same as L. lagotricha. Skull: total
length 112 mm.; occiput broken ; occipito-nasal length 105 ;
zygomatic width 743; intertemporal width 45; palatal
length 84; breadth of brain-case 57 ; length of brain-case
from end of nasals 79; length of nasals 13 ; width of nasals
anteriorly 13; length of upper molar series 24; length of
mandible 74 ; length of lower molar series 30°5.
B.M. no. of type 90. 2. 22. 2.
‘'wo specimens are in the collection which differ so
markedly from all other members of the genus that it is
impossible to assign them to any described species. The fur
is soft and very thick, particularly so at the base of the tail.
One is dark purplish brown or blackish on the upperparts to
rump, and blackish to grey on legs and tail; the other is
darker. The skulls resemble, as may be expected, those of
LL lagotricha and L. infumatus in general, but the nasals have
a depression in the middle and the anterior portion stands at
a right angle to the posterior and are very broad anteriorly ;
the brain-case is shorter than in the skulls of the other species
and the narial opening is of a different shape, like a heart
but not so pointed, more rounded on the lower side.
194 Dr. D. G. Elliot on apparently new
Family Cer copithecide.
Genus PAPIo.
Papio strepitus, sp. n.
Type locality. Fort Johnston, Nyasaland, 8.K. Africa.
Gen. char. Size large, exceeding the dimensions of
P. pruinosus; hair very long, loose; face partly naked ;
brain-case about two thirds the length of the facial region ;
nasals only slightly raised above rostrum, wide anteriorly ;
pit in side of lower jaw long and deep ; palate narrow and of
nearly equal width throughout its entire length; tooth-rows
straight.
Colour. Forehead mixed pale yellow and black; crown
and nape dull tawny ochraceous, centre of crown darker, the
hairs ringed with dull tawny ochraceous and black, those on
side Prout’s brown at base, rest tawny ochraceous. The
appearance of the crown and nape is more reddish than
yellow, with a dark central portion ; sides of head below ears
buff; lower part of neck to middle of back purplish drab
and ochraceous buff; base of hairs purplish drab, the rest
ringed with black and ochraceous buff, some hairs tawny
ochraceous tipped with black. The purplish drab of the
base of hairs dominates the other colours and gives a kind of
dark patch to this part of the back. On the shoulder is a
patch of hairs, buff at their roots, graduating to cream-buff at
their tips, not ringed; lower part of back paler than the
upper, more yellow showing, and over all the upperparts are
numerous long hairs with whitish tips; flanks ochraceous
buff; upper part of arms, entire legs, and feet ochraceous
buff; forearms and hands mixed ochraceous buff and black,
the latter being the colour of the base of the hairs showing
through ; underparts yellowish grey ; tail at base like back,
mixed black and ochraceous, tip ochraceous buff; cheeks and
sides of nose and the lips covered with short yellowish hairs ;
upper eyelids flesh-colour, space beneath the eyes and the
nose black.
Measurements. Head and body 915 mm. ; tail C09 (skin).
Skull: total length 195; oceipito-nasal length 160; hen-
sel 139; intertemporal width 58; zygomatic width 115;
palatal length 86; bieadth of brain-case 79; length of
nasals 72; anterior width of nasals 14; length of upper
molar series 45; length of mandible 115; length of lower
molar series 60.
BA no. ot type 97. 10. 1.9:
Species and Subspecies of Mammals. 195
The general appearance of this species is that of a yellowish
animal, with a brownish back and a reddish head and limbs,
and underparts whitish grey. ‘This is the effect the various
colours of the hairs produce when glancedat. In coloration it
belongs to the light-hued baboons, of which section P. babuin
may be considered a representative. Looked at in certain
lights the hairs have a greenish-yellow tint, but when care-
fully examined the colours are as given in the description
and unlike any of the other species. ‘I'wo specimens were
obtained by Sir H. H. Johnston in Nyasaland, the type at
Fort Johnston and the other at Zomba on Lake Nyasa.
The species bears no resemblance whatever to P. prudnosus,
Thomas, also procured at Fort Johnston, either in colour or
in the characters of the skull.
Genus COLOBUS.
Colobus tephrosceles, sp. n.
Type locality. Ruahara River, Toro, altitude 4000 feet,
Centra] Africa.
Gen. char. Similar to C. rufomitratus, but differs in not
having any black on the head between tufts, in the pale
coloured arms and legs, in the grey-brown basal portion of
the tail, in the absence of black stripe between ears and
shoulders, and in the feet and hands being brownish black
instead of dark olive-brown. Upright tufts on sides of crown
above ears.
Colour. A narrow black band on forehead extending
backwards to ears; top of head and nape dark rusty brown ;
upright tufts dark rusty brown, mixed with some brownish-
black hairs tipped with yellow, these showing chiefly on the
outer side of the tufts; sides of head between ears and eyes
blackish grey, the hairs hiding the ears; side of lip purplish
grey, this extending over the lower jaw; hairs on upper part
of back long, covering the shoulders, brownish black, grading
into dark Prout’s brown on sides and rump; outer side of
arms pale greyish brown; outer side of legs pale brownish
grey, lighter than the arms; underparts and inner side of
limbs greyish white; hands brownish black; feet Vandyke
brown ; tail, basal third greyish brown, remainder blackish
brown, grading into black at tip.
Measurements. No skull to type specimen, Another skull
without skin, procured by Sir H. H. Johnston in Toro, has
total length 116 mm.; occipito-nasal length 95; hensel 84;
zygomatic width 78; intertemporal width 44; palatal
196 Mr. O. Thomas on
length 45; length of nasals 16; length of upper molar
series 29; length of mandible 81; length of lower molar
series 35.
B.M. no. of type 1. 8. 9. 129.
Three examples of this remarkable monkey were procured
by Sir H. H. Johnston (one adult and two young) on the
east side of Mount Ruwenzori at an altitude of 4000 feet.
It is evidently closely allied to C. rufomitratus from the
coast, but exhibits quite sufficient differences in colour and
markings to entitle it to a distinctive rank. As the species
was not seen by the members of the Ruwenzori Expedition,
lately returned to England, it must be considered as rare
even in its own district.
XXIX.—On Mammals from Northern Persia, presented to the
National Museum by Col. A. C. Bailward. By OLDFIELD
‘THomas, F.R.S.
In 1905 Col. A. C. Bailward made a shooting-trip across
Persia, taking with him Mr. R. B. Woosnam to pay special
attention to the collecting of natural history specimens, and
he then obtained the series of mammals of which [ gave an
account during the succeeding winter *.
During the present year Col. Bailward has again been to
Persia, taking Mr. Woosnam with him, but this time to the
northern part of the country, between ‘Teheran and the
Caspian, and there Mr. Woosnam has collected the specimens
enumerated below. Few in number as they are, for the trip
was quite a short one, they include no less than five new
forms, whiie all of them are most valuable accessions to the
Museum, which had previously possessed almost nothing from
that region. We thus have reason to be most grateful to
Col. Bailward for taking advantage of his trip to increase our
National Ccllections in a region as yet so poorly represented
in the Museum; and the marked success of the present
expedition should encourage him and others to repeat the
experiment.
1. Pipistrellus Kuhl, Natt.
g. 120. Teheran, Persia. 4600’.
A pale form, probably representing P. lepidus, Bly.
* PZ. 8, 1908, ii, p. 519.
Mammals from Northern Persia. 197
2. Pipistrellus pipistrellus, Schr.
gd. 118. South coast of Caspian. 150’.
3. Pterygistes noctula, Schr.
gd. 134, 135. Resht, S. coast of Caspian.
4, Myotis myosotis, Bechst.
6. 132; 9.131. Elburz Mts., near Resht. 400’.
5. Miniopterus Schretbersi pallidus, subsp. n.
?. 113. South coast of Caspian. Alt. —25 m. 25th
March, 1907, B.Momnonwd. Te las Type:
Similar to typical J/. Schreibersi in all essential respects,
but the general colour is paler, approximating to Ridgway’s
** wood-brown,”’ while Schrezbersi is rather darker than
‘ broccoli-brown.” Below, the colour, smoky grey in
Schreibersi, is more or less suffused with buffy, especially in
the inguinal region and along the proximal edge of the
interfemoral.
Dimensions of the type (the starred measurements taken
in the flesh) :—
Forearm 46°5 mm.
*Head and body 55; *tail 57; *ear 12.
Skull: condylo-basal length 15; basisinual length 12 ;
mastoid breadth 9.
All the European Miniopteri in the Museum collection are
quite similar in colour and equally different from this pale
Caspian example.
6. Crocidura russula caspica, subsp. n.
&. 106. South coast of Caspian Sea. Alt. —25 m.
16th March, 1907. B.M. no. 7. 7.14.8. Type.
Like C. russula monacha, Vhos.t, but larger, with shorter
tail and darker belly.
Size markedly larger than in monacha. General colour
above of exactly the same smoky brown as in that animal,
but below it is even darker, being, in fact, of a brown very
little lighter than the upper side. Tail of normal vwssula
proportions, therefore much shorter than in monacha,
Skull large, stouter and heavier than in true russula, there-
fore much larger than in monacha, whose skull little exceeds
+ From the neighbourhood of Trebizond. Ann, & Mag. Nat. Hist. (7)
xvii. p. 417 (1906).
198 Mr. O. Thomas on
that of C. mimula, Mill. Brain-case of the narrow high
type characteristic of C. russula.
Dimensions of the type (measured in flesh) :—
Head and body 77 mm.; tail 38*; hind foot 14; ear 9.
Skull: condylo-basal length 21; basal length 19; greatest
breadth 9°6; height of brain-case from basion 5:4; length of
upper tooth-series 9-4.
Fab. and type as above.
This Eastern representative of the common C. russula is
readily distinguishable from any of the described forms of
that species by its dark colour, and especially its very dark
under surface, which is scarcely lighter than the upper side.
7. Crocidura leucodon persica, sp. n.
gf. 127; 2.128. Elburz Mts., near Demavend. Alt.
6500’. 2nd May, 1907. B.M. no. 7. 7.14.9. Type.
‘Trapped near a stream in oak-forest.’”’—R, B. W.
Quite like true German C. leucodon in size, proportions,
and skull, but the fur is shorter (hairs of back about 3°5 mm.)
and the colour of the adult is much paler, that of the type
being somewhat paler than Ridgway’s “drab.” The distri-
bution and sharp definition of the upper and lower colours
quite as in lewcodon.
Skull with the characteristic flattened shape of the brain-
case found in true C. leucodon.
Dimensions of the type (measured in the flesh) :—
Head and body 72 mm.; tail 35 ; hind foot 12; ear 8.
Skull: condylo-basal length 19; basal length 17; greatest
breadth 9°2; height of brain-case from basion 4°6 ; length
of upper tooth-series 9.
Hab. and type as above.
This pretty little shrew is readily distinguishable from the
European C. leucodon by its paler and more drabby colour.
From C. Guldenstaedti, Pall., it is at once separable by its
much shorter tail, as that species, with a trunk-measurement
of 72 mm., has a tail 47 mm. in length.
Thanks to the kindness of Dr. Camerano and Dr. Festa, of
the 'urin Museum, I have been allowed the loan of the
typical specimens of de Filippi’s Crocidura fumigatat, and
find that that animal is also a longer-tailed form, the spirit-
specimen measuring 60 mm, for the head and body and
42 mm. for the tail. All traces of the original colour have
* Measured on skin. Mr. Woosnam writes 46, which is probably a
lapsus calami for 36.
+ Viaggio in Persia, p. 343 (1865).
Mammals from Northern Persia. 199
unfortunately long vanished from the specimens, but the
proportions and skull readily show that it is a different shrew
from that now described.
The Trebizond form which I described as C. leucodon
lastus * differs still more from normal Kuropean C. leucodon
both in size and length of tail, and [ am now disposed to
think it should be recognized as an independent species,
whose name would therefore be Crocidura lasia.
8. Mus rattus, L.
?. 102, 108, 109. South coast of Caspian. 0,
9. Mus musculus, L.
6. 100, 101, 104, 133. South coast of Caspian. 0.
6. 125,126. Elburz Mts., near Demavend. 6500!.
10. Micromys sylvaticus, L.
6 103, 100,100) Pit oe tA id PG. | South: coast
of Caspian. 0!.
6. 129,180; 9.124. Elburz Mts., near Demavend.
6500!
11. Nesokia Bailwardi, sp. n.
a. ?. 351. Bunder-i-gaz, 8. shore of Caspian. Alt. —25m.
25th March, 1907. B.M. no. 7. 7. 14. 30. Type.
A member of the N. Huttoni group.
External characters as usual, the general colour of the type
wood-brown above, rather paler below.
Skull larger and heavier than that of N. Huttoni. Superior
outline strongly bowed. Nasals short, not broadened in
front, evenly but slightly narrowing backwards to the more
or less rounded posterior end. Supraorbital ridges very
thick and heavy at the postorbital point, abruptly dying away
halfway across the parietals. Interparietal fairly large.
Bulle as in WV. Huttont. Molars rather small in proportion
to the size of the skull.
Dimensions of the type (measured on the spirit-specimen
before skinning) :—
Head and body 167 mm.; tail 123; hind foot 33; ear 24.
Skull: condylo-basal length 42; basilar length 37; zygo-
matic breadth 27; nasals, length 12-2, breadth anteriorly 3-9,
* Ann. & Mag. Nat. Hist. (7) xvii. p. 416 (1906),
200 Mr. O. Thomas on
mesially 3°7; interorbital breadth 6:1; tip to tip of post-
orbital processes 10°7 ; interparietal 4 x 8; palatilar length 23;
diastema 14; palatal foramina 5; length of upper molar
series (crowns) 6°5, (alveoli) 8:5.
Fab, and type as above. Thespecimen not old, the molars
being little worn down and the basilar suture not quite
closed.
This Nesokia is probably the form referred to N. Huttont
by Radde and Walter * in their paper on the mammals of
Transcaspia, while they described as new a species (“ N.
Boettgert”) which appears to me to be really assignable to
the true N. /Tutton?, as also, I think, is probably the case
with Nehring’s “ NV. Huttont var. Sutunint” T from Merv.
The majority of the measurements of the skull given by
Radde and Walter for N. Boettgert might actually have been
taken from the typical skull of N. Huttont now in the British
Museum (no. 79.11. 21. 499), while a topotype of N. H. Satu-
nini is also quite similar.
NV. Bailwardi mainly differs from N. Huttoni by its larger
size, more bowed skull, and the heavier supraorbital ridges,
these characters being the more noteworthy as the typical
skull is distinctly younger than that of the Kandahar species.
A close ally is described in the succeeding paper.
12. Microtus terrestris persicus, de Fil.
g. 110. S. coast of Caspian Sea. Alt. 0.
‘Trapped on banks of a stream.”—Z. B. W.
As with the type of Crocidura fumigata, | have been most
kindly allowed the loan of de Filippi’s examples of his
« Arvicola amphibius var. persicus”’ by the authorities of
the ‘Turin Museum.
Jo that form two voles from Van, obtained in 1896 by
Major W.H. Williams, were referred by Barrett-Hamilton f,
and the interest in examining the type was to see if it
presented the peculiar rounded character of the molars, to be
described further on, which was present in these Van
examples.
De Filippi’s specimens consist of a stuffed specimen and
an imperfect skull ; but as it was possible that the two indi-
viduals might differ in the character of their teeth, I have
been permitted to extract the skull of the stuffed specimen,
which must be regarded as the main type, since the external
* Zool. Jahrb. Syst. iv. p. 1036 (1889),
+ SB. Ges. nat. Fr. Berl. 1899, p. 108,
}{ Ann, & Mag. Nat. Hist, (7) ii. p. 224 (1899),
Mammals from Northern Persia. 201
characters were alone used by de Filippi to distinguish his
variety.
The skull so extracted proves fortunately to be perfect,
and is therefore a great improvement on the broken skull on
which he based his statement that the osteological characters
of the Persian vole were absolutely the same as those of
Arvicola amphibius. But the statement itself needs practi-
eally no modification, for this typical skull has no special
peculiarities, and in particular has the normal angular molars
found in European water-voles in general. It therefore
precisely agrees with Mr. Woosnam’s specimen no. 110 from
the low-lying shores of the Caspian Sea, a region through
which de Filippi also passed; and I am disposed to. believe,
therefore, that though that author first saw animals of this
group at Sultanieh, which is on the plateau south of the
Elburz, the actual specimen he brought home and described
was from the lower and more northern region. Moreover, in
speaking of them at Sultanieh, he says they were abundant
then and onwards to the end of his time in Persia, a state-
ment that covers the Caspian as well as the Elburz region.
Actual Sultanieh specimens may hereafter upset this con-
clusion ; but in any case I feel compelled to distinguish the
round-toothed forms as a different subspecies, to which all
the specimens as yet certainly known to come from the
plateau belong.
13. Microtus terrestris armenius, subsp. n.
3. 123. Elburz Mts., near Demavend. Alt. 9000!.
[g. 16,17. Van, Armenia. Alt. 5000’. Presented by
Major W. H. Williams, R.A. ]
** Shot in a small stream.”—R. B. W.
“ Dug out of hole by stream.” —W, A. W.
External characters quite as in IZ, t. persicus, except that
the general tone is slightly greyer and the ears are rather
shorter.
Skull essentially as in that animal, though the brain-case
is, perhaps, a little longer in proportion to its breadth and
the nasals incline to be more expanded anteriorly.
Molars with quite the same pattern as in the lowland
form, but all the enamel-bound spaces, instead of being
sharply angular externally and internally, are rounded, often
almost circular, so that there are no real angles, external or
internal, either above or below, the teeth having therefore a
very peculiar and characteristic appearance, quite unlike that
found in normal water-voles,
Ann. & Mag. N. Hist. Serv. 7. Vol. xx. 14
202 Mr. O. Thomas on
Dimensions of the type (taken in flesh) :—
Head and body 195 mm.; tail 126; hind foot 29; ear 16.
Skull: condylo-basal length 41°5; basilar length 37:2 ;
greatest breadth 25; nasals 11:2X5°6; interorbital con-
striction 4°8; palatal foramina 7; length of upper molar
series (crowns) 9°6, (alveoli) 10:1.
External dimensions of Mr. Woosnam’s specimen (mea-
sured in flesh) :—
Head and body 174 mm.; tail 136; hind foot 33; ear 16.
Hab. Armenia and N.W. Persia, on plateau. Type from
Van, 5000!.
Type. Adult male. B.M. no. 97. 6.4.10. Original
number 17. Collected 23rd December, 1896, and presented
by Major W. H. Williams, R.A.
I can find no evidence that the peculiar rounded character
of the molar spaces, as contrasted with their normal angular
condition in the type of persicus and the Caspian Sea
specimen no. 110, is due either to sex, age, or individual
variation, and it therefore seems advisable to give a special
name to the specimens that show it in spite of their resem-
blance to the Jowland examples in other respects.
14. Microtus sp.
gS. 121. Elburz Mts., near Demavend. 4000!.
A small species of the JZ. arvalis group.
15. Cervus elaphus, L.
Young ?. 122. Elburz Mts., nea Demavend. 5000/.
XXX.—A Subdivision of the Old Genus Nesokia, with
Descriptions of Three new Members of the Group, and of a
Mus from the Andamans. By OLDFIELD THOMAS.
THe genus Nesokia contains three such very distinct and
natural groups that in accordance with modern ideas they
should be recognized as distinct genera. ‘Their respective
characters have already been described by Anderson, Blanford,
and myself, but the most tangible may be briefly recapitulated
as follows :—
new Mammals from the Andamans. 203
I. Nesoxra, Gray, Ann. & Mag. N. H. x. p. 264 (1842).
Type. Arvicola indica *, Gray & Hardwicke. (Nesokia
Hardwicket auctorum.)
Skull short and broad. Palatal foramina short. Molars
laminate, least M/us-like.
Mammex 2—2=8.
II. Gunomys f, gen. nov.
Type. Arvicola bengalensis, Gray & Hardwicke. (Nesokia
bengalensis a.uct.)
Skull broad. Palatal foramina long.
Mamme irregular, 14-18 in number.
III. Banpicota, Gray, Ann. & Mag. N. H. (4) xii. p, 418
(1873).
Type. ‘ Bandicota gigantea,” 1. e. B. bandicota, Bechst.
Skull comparatively long and narrow. Palatal foramina
long. Molars most Mus-like.
Mamme 3—3=12.
These three genera are specialized in the order given,
Nesokia being the most extreme and the farthest from
Mus, both in skull, tooth-structure, and external characters,
and Bandicota the nearest, while Gunomys is intermediate
between the other two.
Nesokia suilla, sp. n.
Nesokia Bacheri, Nehring, Anderson & de Winton, Zool. Egypt,
Mamm. p. 286, pl. L. (1902).
Closely allied to NW. Bailwardit. Distinguished from
N. Bacheri§ by smaller size and smaller bullee.
Colour and other external characters as in WV. Badlwardi.
Skull, as compared with that of N. Bailwardi, averaging
about the same size, but with various differences. in detail.
Zygomata more boldly expanded and forming a strongly
convex shoulder opposite their anterior root, those of N. Bail-
wardi evenly broadening outwards to their posterior part.
* Bechstein’s Mus indicus being now removed, as a Bandicota, from
the genus Nesokia, the specific term given by Gray and Hardwicke again
becomes tenable for the animal usually known as Nesokia Hardwickei.
The same specimen (B.M. no, 99 a) is the type of both names.
{ yodvos, fruitful, prolific.
{ Supra, p. 199.
§ Dr. Nehring (SB. Ges. nat. Fr. Berl. 1901, p. 219) states that the type
of Wagner's “* Meriones myosuros,” which is a Nesokia, is of the same
size as NV. Bachert. It would therefore be larger than either N. Batlwardi
or N, suilla.
Was
204 Mr. O. Thomas on
Nasals decidedly longer, broadened and projected forward
anteriorly, then abruptly narrowed posteriorly in their middle
third, and running backwards to a point, their edges, behind
the anterior third, very faintly concave. Supraorbital ridges,
even in the oldest specimens, not so thick as in N, Bailwardi.
Interparietal smaller. Bulla as in N. Batlward?, conspicu-
ously smaller than in N. Bacheri. Molars larger than in
N. Bailwardi, the crown measurement of an immature
specimen 7°6 mm., as against 6°5 in that animal (9-0 in
NV. Bacher).
Dimensions of the type (measured in skin) :—
Head and body 185 mm. ; tail 113; hind foot 32; ear 16.
Skull: condylo-basal length 44:5; basilar length 39;
zygomatic breadth 27; nasals, length 15, anterior breadth
5, middle breadth 3°6; interorbital breadth 6°1; inter-
parietal 3°2 x 6; palatilar length 24:5 ; diastema 15-5; palatal
foramina 6°2; length of upper molar series (crowns) 7:8,
(alveoli) 8°8.
Hab, Eastern Egypt. Type from Shaluf, Suez.
Type. Old male. B.M. no. 4. 8.2.29. Original number
59. From the collection of the late Dr. John Anderson ;
presented by Mrs. Anderson.
Several examples of the Egyptian Nesokia, all agreeing in
the above-mentioned characters, have now been received
by the Museum. Besides Dr. Anderson’s original series
there are three, purchased alive in Cairo by the Hon. N. C.
Rothschild, and one, said to be from the Fayoum, presented
by Capt. 8. 8. Flower.
Gunomys varius, S}). Nn.
The Malay representative of G'. bengalensvs.
Size decidedly larger than in G. bengalensis. Fur very
coarse and harsh, though not to be called spiny; freely
mixed with longer piles attaining 83-4 cm. in length. General
colour above coarsely mixed black and cream-buff, without
the brown tints of G. bengalensis. Individually the ordinary
hairs are slaty at base, black at tip, with a broad cream-buff
subterminal band ; the longer piles black, a few of them
white. Under surface dull grizzled greyish, the slaty-grey
bases of the hairs more conspicuous than their duli whitish
tips. Head and dorsal line particularly heavily pencilled
with black. Ears brown. Hands and feet brown on the
middle of the metapodials, whitish laterally and on the digits.
Tail well clothed with coarse hairs 2-3 mm. in length; dark
brown above, rather paler below.
new Mammals from the Andamans. 205
Skull similar in general characters to that of G. bengalensis,
but larger and heavier throughout.
Dimensions of the type (measured in the flesh) :—
Head and body 266 mm.; tail 197; hind foot 40; ear 19,
Skull: condylo-basal length 48; basilar length 43;
greatest breadth 26°7 ; nasals 16 x 5:2; interorbital breadth
6°5; frontal height * 15°5; palatilar length 25°5 ; diastema
163; palatal foramina 10X2°6; length of upper molar
series (crowns) 7°3, (alveoli) 8°5.
Hab. Georgetown, Pinang, Malay Peninsula.
Type. Adult male. B.M. no. 98.8.3.3. Collected 8th
April, 1898, and presented by Capt. 8. S. Flower.
This is the “ Mus set’fer, Horsfield,” of Dr. Cantor’s “ List
of Malayan Mammals” ft, a specimen collected by him in
Pinang having been received with the Indian Museum
collections in 1879.
G. varius is distinguishable from G. bengalensis by its
coarse fur, variegated colour, and comparatively large size.
Gunomys varillus, sp. n.
Like G. varius, but very much smaller.
Size very small, scarcely equalling the smallest S. Indian
species of the genus. Fur coarse; longer piles almost con-
fined to the posterior back. General colour above about as
in G. varius, or rather more buffy, but the mixture is finer,
not so coarsely variegated. Under surface dull greyish, the
hairs slaty at base, dull creamy terminally. Hands and feet
brown. ‘Tail more finely scaled than in G. varus, uniformly
brown.
Skull very much smaller in all dimensions than that of
G. varius, but essentially similar in form. Nasals short and
narrow. Supraorbital ridges less heavily developed. Anterior
zygomatic plate well projected forwards, much more so than
in the equally small $8. Indian G. kok. Palatal foramina of
equal breadth for their anterior two-thirds, narrowed in their
posterior third.
Dimensions of the type (measured in skin) :—
Head and body 184 mm. ; tail 137; hind foot 36°5; ear 16.
Skull: condylo-basal length 40°5; basilar length 35 ;
greatest breadth 23; nasals 13x4°3; interorbital breadth
5°7; frontal height 12°5; palatilar length 21; diastema 13 ;
palatal foramina 8xX2°4; length of upper molar series
(crowns) 7, (alveoli) 8.
* From the supraorbital ridge to the alyeolus between m? and m’,
+ J.A.S. B. xv. p. 254 (1846).
206 On new Mammals from the Andamans.
Hab. Georgetown, Pinang.
Type. Adult male. B.M. no. 98.8.3.5. Collected and
presented by Capt. S. 8. Flower. Another specimen obtained
by Dr. Cantor.
Both Dr. Cantor in 1845 and Capt. Flower half a century
later obtained in the little island of Pinang examples repre-
senting two speeies of this genus, a large anda small. The
former marked both his specimens as “ ALus setifer,” evidently
taking them for the same species, but there cannot be the
slightest doubt that they are quite distinct animals, their
difference in size being far too great to be due to individual
variation.
Mus Rogerst, sp. n.
A spinous-haired species with 1—3=8 mamme.
Size of M/us norvegicus. Fur coarse, profusely mixed with
spines, which on the back are about 16 mm. in length by 0°4
mm. in breadth. General colour coarsely grizzled ochraceous
brown, the bases of both hairs and spines pale grey, the tips
of the spines black and of the ordinary hairs ochraceous.
The few long bristle-hairs are wholly black. Sides greyer.
Under surface not sharply defined, pale buffy greyish, the
hairs pale slaty at base, dull cream-buff terminally. Hars
finely haired, dark grey. Limbs dark grey externally, light
grey like belly along their inner aspect. Hands and feet
white above, the metapodials slightly darker; fifth hind toe,
without claw, reaching to the end of the first phalanx of the
fourth. ‘Tail rather shorter than head and body, almost
naked; rings of scales about 10 to the centimetre; dark
brown above, whitish flesh-colour below. Mamme 1—3=8.
Skull strongly built, with well-marked supraorbital beads,
which are continued across the parietals to the corners of the
interparietals. Muzzle rather narrow, parallel-sided. Palatal
foramina not reaching back to the level of the molars.
Mesopterygoid fossa broadly rounded in front, its anterior
limb slightly anterior to the front end of the parapterygoid
fosse on each side of it. Bulle of medium size. Molars
small in proportion to the general size.
Dimensions of the type (measured on the spirit-specimen
before skinning) :—
Head and body 195 mm.; tail 188; hind foot (s. u.) 41;
ear 28.
Skull: greatest length 48°5; basilar length 40; zygomatic
breadth 22°5; nasals 18x5; interorbital breadth 7:2;
greatest separation of parietal ridges 16; palatilar length
On some British Polyzoa. 207
23°3; diastema 14-2; palatal foramina 9-2; length of upper
molar series 7°8.
Hab. W. Coast of South Andaman Island, north of Iké
Bay.
Type. Adult female. B.M. no. 6. 4.13.2. Collected
February 1904, and presented by C. G. Rogers, Esq.
In spite of the number of rats recently described by
Mr. G. 8. Miller * from the Andaman group, this fine species
does not appear to have been previously obtained. Its very
unusual mammary formula, 1—3=8, is alone shared, in the
whole of the Muride, by Mus bagobus, Mearns, from the
Philippines, and Mus pulliventer, Miller, from the Nicobars,
of which latter it may be the Andaman representative, but
from which it differs by its markedly larger size and distinctly
bicolor tail.
XXXI.— On some British Polyzoa. By Canon A. M.
Norman, M.A., D.C.L., LL.D., F.R.S., F.L.S.
[Plate [X.]
*
Micropora impressa (Moll). (Pl, TX. figs. 1-3.)
1803. Eschara impressa, Moll, Eschara, p. 51, pl. ii. fig. 9.
ae Eschara andegavensis, Michelin, Icon. Zoophyt. p. 329 (nec auct.
ur.). Le
1848, nen gracilis, Reuss, Foss. Polyp. des Wiener Tertiirbeckens,
p- 98, pl. xi. fig. 12 (mee Von Miinster).
me Pee Pes calpensis, Busk, Brit. Mus, Cat. p. 60, pl. civ.
oS. 0
1867. Membranipora bifoveolata, Heller, Bryozoen des adriatischen
Meeres, p. 19, pl. ii. fig. 1
1871. Membranipora calpensis, Manzoni, “Supp. alla Fauna Bryoz.
Medit.,” Sitz. k. Akad. d. Wissensch. vol. lxiil. p. 3, pl. i. figs. 2, 3.
1879. Micropora impressa, Waters, Ann. & Mag. Nat. Hist. ser. 5,
vol, iii. p. 123.
I have recently found among material put by for further
examination a little box which contained three small pieces of
the above Polyzoon, and labelled Guernsey. I cannot recall
to mind whether I procured these specimens myself at
Guernsey in 1865 or whether they were given to me.
Other specimens are in my collection trom Naples, where
I found it in 1887 to be abundant, and from the Adriatic,
given to me by my late friend Professor Heller under his
name * Membranipora bifoveolata.”
* Pr, U.S. Nat. Mus. xxiv. p. 758 (synopsis of species) (1902).
208 Canon A. M. Norman on
Genus TEREBRIPORA, d’Orbigny.
This interesting genus, the exact position of which cannot
be determined until the animal shall have been examined,
but which is presumed to be a burrowing Polyzoon, was
instituted by d’Orbigny in 1841* to include two species,
Terebripora ramosa and T. irregularis, which he had found
in shells of Calyptrwa, Crepidula, and Pecten off the South-
American coast.
In 1865 Paul Fischer published an excellent paper on the
family}, inwhich he enumerates all the species both recent and
fossil presumed to be referable to his ‘ Famille des Térébri-
porides.” In this paper eight recent and fourteen fossil species
are recorded. ‘I'wo of the recent species had been found in
European seas—one, Terebripora Orbigniana, Fischer, bur-
rowing in shells of Ostrea edulis at Arcachon, and in Conus
mediterraneus and Triton nodifer in the Mediterranean ; the
other, Spathipora sertum, Fischer, found at La Rochelle,
Arcachon, and the Mediterranean in shells of Lutraria ellip-
tica, Cardium norvegicum, Pectunculus glycimeris, and Triton
nodifer, in 1880 M. J. Jullien} added another recent
species, 7’. Fischeri, which was found in a shell of Buccinum
from Cape Verd Islands. *
Terebripora ditrupe, sp.n. (Pl. LX. figs. 4-7.)
Terebripora has a mode of growth analogous to that of
Flippothoa divaricata, but instead of running over the surface
of shells &c. as in the latter species, the whole polyzoary is
buried in its substance, except that the orifices of the zocecia
open through the surface. The thread-like connecting fibres
or stolons in all species hitherto described appear to be quite
simple, but in 7’. ditrupe they consist of lines interrupted on
one side by small lateral projecting processes (fig. 5). The
zocecia are not in the same plane as the connecting fibre, but
at right angles to it, in such a manner that they are also
perpendicular to the surface (fig. 6). Owing to this position
of the zocecia their lower portion is too deeply seated to be
seen with the microscope; the oral opening has a somewhat
* d’Orbigny (A.), ‘ Voyage dans Amérique méridionale,’ yol. vi. p. 28,
xs
+ Fischer (P.), “ Etude sur les Bryozoaires perforant de la Famille des
Térébriporides,” Nouv. Arch. du Museum, vol. ii. pp. 293-318, pl. xi.
¢ Jullien (J.), “ Desc. nouv. Espéce de Bryozoaire perforant du genre
Terebripora, dOrbigny,” Bull. Soc. Zool. de France, 1880, pp. 1-4 and
woodcut (separate copy).
some British Polyzoa. 209
irregular margin, but would appear to have a lip-like
projection (fig. 7).
The calcareous shells of the Annelidan genus Ditrupa are
dredged in extraordinary profusion on some parts of the
“ Haat” off Shetland. I had saved a large box full of
specimens of these, selected on account of encrusting growths
on them, mostly of Polyzoa. On examining the contents of
this box, which contained at least two thousand Ditrupe, I
discovered in one specimen the Yerebripora which I have
here described.
Schizoporella Alder (Busk).
This species is subject to some variation in its mode of
growth and in the absence or presence of lateral avicularia.
It has been several times described. The original illustra-
tions of Busk are very good. The following will give the
synonymy of the species :—
Var. a.—Chain-like growth ; without avicularia.
1856, Alystdota Alderi, Busk, Quart. Journ. Micr. Sci. vol. iv. p. 311,
pl. ix. figs. 6, 7.
1869. Alysidota Alderi, Norman, “Last Report Dredging Shetland,”
Brit. Assoc, Rep. for 1868, p. 306.
Var. b.—Chain-like growth; with avicularia. (Chiefly
arctic, rare Shetland.)
1867. Mollia vulgaris, forma ansata (partim), Smitt, “ Kritisk Forteck.
&e.,” Csfvers. K, Vet.-Akad. Forhand. p. 15, pl. xxv. fig. 81,
1880. Schizoporella Alderi, Hincks, Hist. Brit. Marine Polyzoa, p, 243,
pl. xxxvi. figs. 9, 9a,
Var. c.—Clustered growth; without avicularia. (Shetland,
&e.)
1860, Lepralia Barleet, Busk, Quart. Journ. Micr. Sci. vol. vii. p. 143,
pl. xxvi. figs. 1, 2.
1867. Mollia vulgaris, forma ansata (partim), Smitt, 2. c. figs. 79, 82.
1880. Schizoporella Alderi, Hincks, /. c. fig. 10.
Var. d.— Clustered growth ; with avicularia, (Arctic.)
1867. Mollia vulgaris, var. ansata (partim), Smitt, J. ¢. fig. 80.
1900. Schizoporella Elmwoodie, Waters, ‘ Bryozoa from Franz Josef
Land,” Journ. Linn. Soc., Zool. vol. xxviii. p. 66, pl. ix. figs. 1 & 18.
1905. Schizoporella Stormi, Nordgaard, Hydrog. and Biol. Invest.,
Invert. Norweg. Fiords, p. 166, pl. v. figs. 1, 2.
1906, Schizoporella Elmwoodie, Kluge, Ergiinz. u. die ‘Olga’ Exped.
gesamm,. Bryozoen, p. 40.
1906. Schizoporella Stormi, Nordgaard, Bryozoa from Second ‘Fram’
Exped. 1898-1902, p. 17, pl. i. figs, 10, 11.
210 Canon A. M. Norman on
The following examples are in my collection :—
Var. a. Aldert—Shetland (types Barlee); Shetland and
Bergen Fiord (A. MW. N.).
Var. b—Shetland (with Barlee’s types).
Var. c. Barlee’.—Shetland (types Barlec); Shetland and
Bergen Fiord (A. M. NV.) ; ‘ Porcupine,’ 1869; Spits-
bergen (Smitt, as “‘ Mollia vulgaris, forma ansata”’).
Var. d. Elwoodie.—Grey Hook, Spitsbergen, 90 fath. (Smee,
as Mollia vulgaris, var. ansata); Hammerfest (Nord-
gaard, cotypes of Schizcporella Storm).
Escharina Dutertret (Audouin). (PL IX. figs. 8-12.)
Mastigophora Dutertret, Hincks, Hist. Brit. Marine Polyzoa, p. 279,
pl. xxxvii. figs. 1, 2.
Hincks describes two forms of this species, to which I
would call a little more attention. The Shetland form, which
is abundant there in 80-170 fathoms (figs. 8, 9), has the
zocecia tumid, the surface somewhat roughly granulated,
with a tendency in the granulations to arrange themselves in
radiating lines ; six mouth-spines are developed; the vibra-
cula are of moderate length ; the incision of the lip has the
angles generally rounded off. In one of the ‘ Porcupine’
dredgings of 1869 the Shetland form just noticed occurred
in abundance, but with these were also two specimens which
had a very different aspect. The zocecia were much longer
(fig. 10 as compared with fig. 8, both drawn with the same
magnifying-power) and much more flattened ; the vibracula
were very long, but there were no mouth-spines, while the
incision of the lip had the angles sharply defined. In the
Antrim variety described by Hincks the zocecia are not so
large, but they are nearer this second variety, since they are
similarly flattened and have the oral opening of similar form
(see fig. 12). I have an allied form from Madeira, which is
probably that which has been more than once recorded as
L. Dutertret. It is smaller, much more tumid, and the
vibracula arise from elevated shoulders. It may be a distinct
species.
Phylactella pygmcaa (Norman).
1869. Celleporella pygmea, Norman, “ Last Report Dredging Shetland,”
Brit. Assoc. Report for 1868, p. 308.
1880. Cedlleporella pygmea, Hincks, Hist. Brit. Marine Polyzoa, p. 415.
As has been already recorded by Hincks, this minute
some British Polyzoa. 211
Polyzoon has been dredged by me in 80-170 fathoms off
Shetland and also in deep water in Bergen Fiord. The late
Mr. C. Peach also sent me a specimen taken by him off
Wick, and it was also procured by the ‘ Porcupine’ Expe-
dition of 1869.
As the species has not been figured, it is illustrated here
in the annexed woodcut. I have nothing to add to the
description which has already been given. I have pro-
visionally placed the species in the genus Phylactella, as,
perhaps, it comes nearer to P. collar’s than to any other
form; but it has little in common with the type of that
genus, P. labrosa. The small size of the zocecia may be
judged by fig. 2, which represents the outline of a zoarium
which will contain forty to fifty zocecia.
1. Phylactella pygmea (Norman). 2. Size of a zoarium.
Cellepora surcularis (Packard).
1856. Cellepora cervicornis, Busk, Ann. & Mag. Nat. Hist. ser. 2,
vol. xviii. p. 82; and 1858. Mon. Fossil Polyzoa of the Crag, p. 57;
and 1880. Journ. Linn. Soe., Zool. vol. xvi. p. 258, pl. xiii. figs, 6-8
(nee Cellepora cervicornis, Johnston).
1863. Celleporaria surcularis, Packard, “ List of Animals dredged near
Caribou Island,” Canadian Naturalist, vol. viii. p. 410; and 1867.
“Obs. Glacial Phenomena of Labrador and Maine,” Mem. Boston
Soc. Nat, Hist. vol. i. p. 274.
1867. Cellepora incrassata, Smitt, “ Krit. Forteck. &c.,” Gifvers. K.
Vet.-Akad. Forhand. p. 83, pl. xxviii. figs. 212-216 (nec Ceéllepora
incrassata, Lamikk.).
1886. Cellepora cervicornis, Lorenz, Bryozvén yon Jan Mayen, p. 13,
fie. 12.
A young specimen of this species. encrusting stone and
Palo Senior A. Cabrera on
embracing an upright growing Serpula was dredged by the
‘Porcupine’ in 1869. The station was not preserved, but a
box contained a large number of stones the numerous species
encrusting which were, with the one exception of this
Cellepora, identical with the deep-sea fauna of Polyzoa with
which I am so familiar in the Shetland seas; and there can
be little doubt but that the species was taken within the
British area. The species along with it were Amphiblestrum
trifolium, Escharina Dutertrec, Ramphonotus minax, Mega-
pora ringens, Anarthropora monodon ; ‘* Schizoporella” ansata,
Alderi, and sinuosa; Porella bella; Escharella abyssicola,
laqueata, and microstoma ; Hemicyclopora polita, &c.
EXPLANATION OF PLATE IX.
Fig. 1. Micropora impressa, Moll: a living zocecium,
Fig. Ditto: a dead zocecium.
Ditto: operculum.
. Shell of Ditrupa arietina (Miller).
Segment of this shell magnified, to show the perforations of
Terebripora ditrupe.
Terebripora ditrupe, sp. n.: upper portion of a zocecium.
. Ditto: oral aperture of zocecium.
. Escharina Dutertrei, Audouin, the deep-water Shetland form.
Fig. 9. Ditto, its operculum.
Fg. 10. Ditto: variety taken in company with the last.
Fig. 11. Ditto: ditto, its opercula.
Fig. 12. Ditto: oral opening of a specimen from the Antrim coast.
A
SQ
OMND OP OOr
XXXII.—Three new Spanish Insectivores.
By ANGEL CABRERA.
AMONG a number of Spanish small mammals lately arrived
for my private collection there are a few apparently new
forms of Insectivores that I now propose to describe. Some
of them are also represented in the Natural Science Museum
of Madrid.
Talpa ceca occidentalis, subsp. n.
Characters. A small form of J’. ceca, with a flatter, but
not lower, skull, and very hairy tail and feet. Width of
fore foot considerably greater than its length without nails.
Colour. Brownish black, the hairs being dark silvery grey
with deep brown tips. Middle of under surface without the
Jast colour, the general hue becoming dark silver-grey.
Hairs of the tail very long, black; those of the feet very
new Spanish Insectivores. 213
dark brown. The fur is very glossy, showing silvery re-
flections in certain lights; if wet, it exhibits a brilliant
metallic lustre, green on the upper surface, dark purple on
the belly.
Skull. Similar to that of typical ceca, but the brain-case,
although rather high (more than 9 mm.), is flatter on the
upper surface, so that, viewed from behind, its greatest
breadth appears above the middle horizontal line of the skull,
as in 7’. c. levantis.
Measurements (type in flesh). Head and body 102 mm. ;
tail 24; fore foot, breadth 17°6, length (s.u.) 15°5; hind foot
(s.u.) 15:5.
Skull: greatest length 31°5; basal length 22:5; zygo-
matic breadth 11; breadth of brain-case 15:2 ; palatal length
14 ; upper tooth-row 13:5.
Habitat. Guadarrama Mountains, Central Spain. Alt.
1200-1300 m.
Type. Adult male, from La Granja (Segovia), collected
by Sr. M. de la Escalera, September 1906. No. 122,
collection of A. Cabrera.
Remarks. By its smaller size this mole is easily distin-
guishable from the Italian and Asiatic forms, in which
the head and body length exceeds 120 mm. It therefore
appears to need a subspecific name.
Crocidura russula pulchra, subsp. n.
Characters. A small, long-tailed shrew, like C. mimula in
size, but with a longer tail and the typical skull of C. russula.
Colour. Upperparts pale sepia, with a very slight reddish
tinge, and showing bright silvery reflections on the back.
Ventral surface ashy white. The hairs are everywhere dark
slate at the base, and this colour appears externally on the
underparts. ‘Tail sepia above, dirty white below.
Skull. The skull and teeth are identical in form with those
of OC. russula russula, but a little smaller,
Measurements (type in flesh). Head and body 71 mm. ;
tail 41-5; hind foot (s.u.) 12; ear 8.
Skull: greatest length, exclusive of incisors, 18°9; breadth
of brain-case 9°1 ; greatest antorbital breadth 6; interorbital
breadth 4:2; upper tooth-row 8°4.
Habitat. Kastern Spain, Valencia. A specimen from
Minorea (Balearic Islands) in the Madrid Museum belongs
probably to the same form.
Type. Adult male, from Valencia, collected by Sr. José
M. Benedito, January 1907. No. 117, collection of A.
Cabrera,
214 On new Spanish Insectivores.
Remarks. I think it best to treat this form as a subspecies
until the true relationship of the different shrews of the
russula group is made out. It is noteworthy that almost all
the southern forms of this group (cypria, monacha, caudata,
pulchra) have a remarkably long tail.
Neomys anomalus, sp. n.
Characters. Smaller than typical NV. fodiens; tail rounded,
its lower surface without a keel of hairs.
Colour. Upperparts glossy brownish black, the hairs
being dark iron-grey with reddish-black ends. Underparts
white, slightly washed with yellowish under the neck ; the
white sharply separated from the dark colour on the sides.
Hands and feet white, the latter with a blackish patch running
from the heel along the posterior half of the external border.
The long hairs fringing the foot white. Tail bicolor, brown-
ish black above, white below; the hair on its lower surface
long enough to mask the scales, but not to form a fringe as
in N. fodiens; it is only a little elongated about the end,
hardly forming an inconspicuous terminal tuft.
After a long immersion in alcohol the colour of the dorsal
surface of the body becomes a dark reddish chestnut.
Skull. Compared with N. fodiens, the brain-case is higher
and less rounded, its anterior part being not convex, but
forming a smooth slope. The occiput is also flatter in its
upper part. The teeth show no peculiarities.
Measurements (type, after a short immersion in alcohol).
Head and body 73 mm.; tail 60; hind foot (s.u.) 17:5;
ear 8.
Skull: greatest length, exclusive of incisors, 20°5; breadth
of brain-case 10; greatest antorbital breadth 6°2 ; interorbital
breadth 4; upper tooth-row 9°6.
Habitat. Central Spain. 1 have seen specimens from
Salamanca and Madrid provinces.
Type. Adult male, from San Martin de la Vega (province
of Madrid, on the Jarama River), collected in December 1892.
No. 1140, Museum of Natural Science of Madrid.
Remarks. 'This Neomys is not alone in the lack of a hairy
keel under the tail. ‘The same peculiarity has been found by
Mr. Charles Mottaz in another new form from the Vaud
Alps, Switzerland. Mr, Mottaz has kindly sent me a speci-
men (skin and skul!) and an unpublished description of his
animal, and from comparison it results that both the Swiss
and the Spanish forms, although similar in the tail-structure,
are very different in other points. In the same season the
Mr. A. S. Hirst on new Pill-Millipedes. 205
hair of the Swiss form is shorter and greyer than that of
N.anomalus. The brain-case of the skull in the former species
is rounded and somewhat globular, while in the Spanish
animal it is flat in the anterior part and about the occiput, the
entire outline being not evenly convex, but nearly angular.
Owing to the absence of hair-fringe on the under surface of
the tail, NV. anomalus has hitherto been confounded by Spanish
naturalists with Sore araneus, a species that I have never
seen in the Peninsula. Under that name the specimens in
the Madrid Museum were exhibited.
XXXIII.—On Four new Pill-Millipedes from the Malay
Peninsula and Siam. By A. 5S. Hirst (British Museum,
Nat. Hist.).
[Plate X.]
THE four forms which I describe below as new seem to be
somewhat closely allied to one another. Their copulatory
feet present much resemblance and the walking-legs in all
four species are furnished with three spines above the claw.
Three of them come from the Malay Peninsula, and the
remaining one from Siam. Z. anthracina, Pocock, from the
Malay Peninsula, Z. ¢mpunctata, Pocock, from Penang, and
Z. semilevis, Pocock, from South ‘lenasserim, are also
members of this species-group. ‘The legs of these last species
were described by Mr. Pocock * as being provided with two
spines above the claw; in reality, however, they are provided
with three.
Zephronia rugulosa, sp. n.
Colour (faded, in spirit). Head, nuchal plate, and the first
tergite dark brown or black : tergites dark brown, the ante-
rior borders yellowish brown and ornamented with several
small dark spots +; the last tergite with irregular dark spots.
Head, Anterior region of the head marked with tairly
numerous punctures, the posterior part sparsely punctured.
The anterior border with a single tooth.
Nuchal plate with sparse and fine punctures.
First tergite convex anteriorly, the usual angle being almost
* Ann. Mus. Civ. Genova, ser. 2, vol. x. no, 30, p.5 (1890); Ann. &
Mag. Nat. Hist. ser. 6, vol. xvi. p. 413 (1895).
+ These spots are probably due to bad preservation.
216 Mr. A. S. Hirst on new Pill-Millipedes
rounded off and effaced ; marginal excavated area in the form
of a narrow groove expanding a little anteriorly and con-
fined to the lateral part of the tergite. The two lateral
depressed areas punctured and connected with one another by
a narrow transverse strip, which is finely punctured but not
hollowed out.
Tergites. Punctures almost absent; the anterior margins
roughened and granular ; last tergite minutely punctured in
its anterior half.
Legs provided with three spines above the claw.
Copulatory feet. Anterior forceps with the immovable
finger short, much curved, and flattened, the hollowed-out
inner surface bearing a membranous tooth; movable finger
straight and composed of two segments; the proximal
segment with the upper inner corner projecting and almost
rectangular; distal segment with a spine on the inner side.
Posterior forceps with the immovable finger stout and
bearing a forked membranous tooth on the inner edge;
movable finger stout, curved, and composed of two segments
(figs. 6-8).
Measurements inmm. Total length 53°5, middle breadth 29:5.
Hab. Pahang, Malay Peninsula. Three specimens col-
lected by Mr. Craddock. (Brit. Mus. Coll.)
The following form is apparently a local variety of the
above species :—
Colour (in spirit). Head and nuchal plate black; first
tergite black, with the exception of the middle lateral area,
which is yellowish brown: tergites with the posterior two
thirds dark, the anterior border yellow; last tergite black,
with a narrow yellow border posteriorly.
Nuchal plate smooth, the punctures minute and very
sparse.
First tergite (fig. 1).
Tergites with the anterior margins but little roughened.
Legs with 3-4 spines above the claw.
Copulatory feet. Proximal segment of movable finger of
anterior forceps with the angular projection sharper, less
rectangular, and more conical than is the case in the form
from Pahang (fig. 9).
Measurements in mm. Total length 48°5, middle breadth 25.
Hab. Kelantan (Mr. J. D. F. Gemlette). (Brit. Mus.
Coll.)
Zephronia Ridleyt, sp. n.
Colour (in spirit). Head and nuchal plate black: tergites
from the Malay Peninsula and Siam. 21
‘ y
dark green, the anterior margins pale yellow ; first tergite
with the anterior and posterior margins black and the middle
part yellow.
Head. Anterior part of head furnished with many hairs,
the posterior part with a few scattered hairs.
Nuchal plate with a few fine hairs.
First tergite smooth and shining anteriorly, posteriorly
with many minute punctures. Lateral depressed area in the
form of a very narrow groove, which becomes enlarged in its
inner anterior portion ; the depressed areas of the two sides
are not continuous (fig. 2).
Tergites. Anterior third of tergites finely roughened and
very minutely and densely punctured. The rest of the
surface without granules, the middle part being marked with
fine and numerous punctures, the posterior part more sparsely
punctured. Last tergite minutely punctured.
Legs provided with three spines above the claw.
Copulatory feet. Anterior pair with the immovable finger
of the forceps short, flattened, strongly curved, and bearing
a membranous conical tooth on the inner side; movable
finger straight, of much greater length than the immovable
finger and composed of two segments, the distal one being of
moderate size, the proximal one with the upper inner angle
large and projecting. Posterior pair with the immovable
finger bearing a bifid membranous tooth on the inner edge ;
movable finger bisegmented, a little curved, and fairly
stout (figs. 10 & 11).
Measurements in mm. Total length 34, breadth 19.
Hab. Selangore (in a cave). A single specimen was
collected by Mr. H. N. Ridley during the year 1897. (Brit.
Mus. Coll.)
Zephronia Flowert, sp. n.
Colour deep brown.
Head with punctures bearing hairs anteriorly, posteriorly
with a few scattered hairs.
Nuchal plate smooth, the anterior and posterior margins
with minute punctures, however.
First tergite provided with a smooth, transverse, almost
keel-like elevation, which is situated posteriorly to the ante-
rior groove or depression, and forms its boundary, the ridge
itself being limited posteriorly by a narrow lineal groove
(sometimes indistinct) terminating laterally in diverging
lines similar to those present in Z. nigriceps, Pocock (fig. 3).
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 15
218 Mr, A. 8. Hirst on new Pill-Millipedes.
Tergites smooth, devoid of granules, and ornamented with
a fine scale-like sculpturing, punctures absent.
Legs provided with three spines above the claw (fig. 13).
Vulva. Distal sclerite reduced to a band or strip, which
increases in length (antero-posterior) towards the inner angle
fig. 12
oe rements in mm. Total length 41, middle breadth 19.
Hab. Singapore. Two specimens collected by Capt. 8. 8.
Flower. (Brit. Mus. Coll.)
Zephronia stamensis, sp. n.
Colour (in spirit). Dark green, the tergites usually orna-
mented in the middle part of their anterior half with two clear
yellow patches and with a transverse stripe occupying the
middle part of the posterior border. In many specimens,
however, the anterior and posterior markings are fused
together, leaving a dark green spot (often triangular in shape)
in the middle of the tergite.
Head. Anterior part of the head with many small hairs,
the posterior part sparsely punctured; the anterior margin
with a single tooth.
Nuchal plate unpunctured or sparsely punctured.
first tergite with the depressed area rather narrow in its
lateral portion and connected by a still narrower and shallow
strip with the area of the other side; lateral portions of
depressed area and the connecting strip covered with dense
hair. A smooth and unpunctured transverse strip of the
surface is situated posteriorly to the depressed area, the rest
of the tergite being hairy (fig. 4).
Tergites densely covered with minute hairs (in worn
specimens densely and minutely punctured).
Legs provided with three spines above the claw.
Copulatory feet. Immovable finger of anterior forceps
flattened, much curved, and bearing a membranous process
on the inner side; the movable finger fairly stout and very
indistinctly bisegmented, the serrations of the inner side of
the lower segment ending ina slight projection. Immovable
finger of posterior forceps provided with two membranous
teeth; the movable finger curved, of moderate stoutness, and
composed of two segments (figs. 4 & 15).
Vulva with the distal piece or cap increasing in size
towards the inner side, where, however, it is nearly twice as
broad as long; the outer corner narrowed and extending a
little down the outer side of the proximal segment. Lower
segment with the opening long and V-shaped, the point of
On an apparently new Lycenid from Mauritius. 219
the V (which is continued some distance as a groove) being
directed towards the lower inner corner (fig. 16).
Measurements in mm. Length 26:5, middle breadth 12°5.
Hab. Kosichang and Chantaboon, Siam. <A large number
of specimens collected by Capt. 8. 8. Flower. (Brit. Mus.
Coll.)
EXPLANATION OF PLATE X.
Fig. 1. Zephronia rugulosa, var. First tergite.
Fig. 2. Zephronia Ridley?, sp. n. First tergite.
Fug. 3. Zephronia Floweri, sp.n. First tergite.
tig. 4. Zephronia siamensis, sp.n. First tergite.
Fig. 5. Zephronia impunctata, Pocock. First tergite.
Figs. 6, 7. Zephronia rugulosa, sp.n, Anterior copulatory forceps.
Fig. 8. Ditto. Posterior copulatory forceps.
Fig. 9. Zephronia rugulosa, var, Anterior copulatory forceps.
Figs. 10, 11. Zephronia Ridleyi. Anterior copulatory forceps.
Fig. 12. Zephronia Klowert. Vulva.
Fig. 13. Ditto. Walking-leg.
Fig. 14. Zephronia sitamensis. Anterior copulatory forceps.
fig. 15, Ditto. Posterior copulatory forceps.
Fig. 16. Ditto. Vulva.
XXXIV.—Description of an apparently new Lycenid from
Mauritius. By Hamitton H. Druce, F.Z.S., F.E.S.
Lieut.-Cot. N. MAnpers, whilst residing in Mauritius,
collected a series of a Lyceenid which I am unable to determine
and which he has asked me to describe. It is not included in
Boisduval’s work on the Lepidoptera of the island and is not
mentioned by Dr. Trimenin his list. At first sight I thought
it would be well placed in Dr. Butler’s genus Cyclyrius, but
the coloration of both sexes is so different from the typical
species of that genus (C. Webbianus, Brullé)—it is also
without the chequered cilia—that it seems to me better placed
amongst the tailless group of Nacaduba, some species of which
it much resembles on the upperside.
Nacaduba Manders?, sp. n.
3 .—Upperside uniform dull violaceous blue; costal and
outer margins very narrowly greyish brown. Cilia paler.
Underside: ground-colour pale brownish grey with irregular,
darker, sordid-white edged spots and markings. Fore wing:
an elongate spot crossing the middle of the cell, followed by
another much the same closing its end. Beyond this an
t5*
220 Mr. G. E. Mason on an
ultramedian band of ovular spots commencing just below the
costa and reaching to the submedian nervure, that portion of
it which is opposite the cell being placed further outwards
towards the apex. A dark anteciliary line followed by a row
of minute dark lunules inwardly bordered with sordid white.
Hind wing: several irregular spots near the base and along
the anal margin, and beyond these an irregular much broken
band of spots. commencie on the costa, becoming divided on
the disc, and angled to the anal margin. A dark anteciliary
line and shades as in fore wing. There are three deep
black marginal spots (the centre spot being the largest)
supporting “metallic-blue scales and ringed with pale orange
situated in the three anal nervular interspaces. Cilia of both
wings grey.
Palpi black above, clothed with black and white hairs
below. Thorax and abdomen below more or less clothed
with whitish hairs.
When examined with a glass the whole surface of both
wings below appears to be suffused with sordid white scales.
? .—Upperside: fore wing bright blue, with the costa,
apex, and outer margin blackish brown. Hind wing blackish
brown, slightly paler along the costal edge; the basal
third bright blue. Three dark marginal spots "in the anal
interspaces crowned with bright blue. Cilia of both wings
brown, paler towards apex of hind wing.
Underside as @, but ground-colour darker and spots more
prominent.
Expanse, ¢ 2, 1} inch.
Hab, Mauritius.
Types, ¢ ¢, coll. Manders. Co-types in B.M.
Also in coll. Dr uce, presented by Lieut.-Col. Manders.
The species is without tails.
Lieut.-Col. Manders writes that it flies all the year round,
except in the coldest weather, there being a succession of
broods. He did not find it in the Island of Réunion.
XXXV.—On an Extinct undescribed Fruit- Bat of the Genus
Pteropus from the Mascarenes. By Gro. E. Mason.
A cursory survey of the small island known as La Ronde,
laying about 15 miles north-east of Mauritius, was made a
year ago Py a South African syndicate interested in the
guano trade, and on the face of a deep-wooded gorge situated
m the central part of the island numerous fissures were
met with containing a superficial deposit of red earth, from
Extinct Frut-Bat from the Mascarenes. 221
which a member of the survey collected and transmitted to
me the specimens forming the subject of this communication.
The material consists of a skull, the right and left femur, and
portion of the humerus, all of which can most certainly be
attributed to the same individual, and they clearly indicate a
very characteristic and hitherto unrecorde:l species of frugi-
vorous bat of the genus Pteropus, which has, however,
now ceased to exist on the island, neither does it occur on
any other of the Mascarene Islands. Associated with the
remains were also found the bones of tortoises, birds, and
introduced animals, the presence of these latter, which are
referable to the goat (Capra hircus) and rabbit (Lepus
cuniculus), being of particular interest and affording reliable
data by which we may, with every degree of certainty,
assign the age of the deposit to a period succeeding the
advent of man in the island. Contemporary with Dédus,
Pezophaps, and other members of the then existing fauna of
the Masearenes, this bat must have long survived those
forms, lingering on until a comparatively recent period, the
state of preservation and general condition of the remains under
discussion strongly bearing out this hypothesis. In seeking
for a possible cause for its extinction it is perhaps, in this
case, more reasonable to assign the chief weight to those
changes in the conditions affecting a due and plentiful supply
of the soft fruits and berries so essential for the sustenance
of these voracious creatures, brought about by a series of dry
or tempestuous seasons unduly prolonged, as the means
of subsistence within the very restricted area of the island,
even under most favoured influences, must have been very
limited, rather than to the direct agency of man, which we
know has been instrumental in bringing about the varied
physical transitions and concomitant changes in the endemic
animal life of the whole Mascarene group of islands as knowa
to us at the present day.
For this interesting species I propose the name of
Pieropus mascarinus, sp. n.
Skull.—With moderately long but heavy muzzle, flattened
and scarcely concave frontal region. Sagittal crest weakly
developed, almost obsolete.
Measurements.—A few of the measurements can only be
given approximately owing to the damaged condition of the
specimen :—
Upper length (approximate) 49 mm. ; condylo-basal length
(approximate) 46 ; basilar length (approximate) 44; median
palate length 28 ; breadth between outer sides of canines 9°8,
inside canines 4°3 ; outside p. 4 13°38, inside p. 4 8; outside
222, Onan Extinct Fruit-Bat from the Mascarenes.
m.213, inside m.2 9°2; zygomatic breadth (approximate) 27 ;
least interorbital breadth 7:8; the dimensions of the struc-
ture behind postorbital process cannot be taken; greatest
breadth of brain-case 15 ; occipital depth 11; mandible 40°6 ;
maxillary tooth-row exclusive of incisors (alveoli) 20-1;
mandibular tooth-row exclusive of incisors (alveoli) 22°3.
Teeth—With the longitudinal grooving characteristic of
the genus Pteropus, very strongly developed, somewhat
large and heavy for the size of the animal; canines long and
sharp, both above and below, with unusually acute and
prominent postero-internal basal ledges, those of the maxillary
in particular ; first upper premolars very minute, but would
have been still persistent in the type and standing in the
tooth-row.
Measurements.—The sizes of the upper incisors and second
upper molars are omitted, as these teetl are wanting in the
specimen. Vertical length of upper canine 7°5 mm.; hori-
zontal length of upper canine 3°5 ; horizontal length of p. 3
$°8, width 2°3; horizontal length of p.4 3°8, width 3;
horizontal length of m.1 4°1, width 2-1. Lower teeth—com-
bined breadth of incisors —?; height of canine (from basal
ledge behind) 5; horizontal length of anterior premolar 2 ;
of p.3 3°8, width 2°1; horizontal length of p. 4 3°5; of m.1 4°3,
width 2; horizontal length of penultimate molar 3, width
1:8; horizontal length of m. 3 1°5, width 1:4.
Locality.—Round Island, North-east Mauritius.
As compared with the now existing Pteropt of the
Mascarenes, this species occupies a place intermediate between
Pteropus vampyrus and Pt. rodricensis, in size only, the
dentition being typical of Pteropus, whereas the two above
species fall into the subgenus Speetrum. The only other
fruit-bat occurring in the Mascarenes is Pt. rubricollis, and
this has been placed by Matschie in his subgenus Serico-
nycteris. A great analogy, in fact, exists between the dentition
ot Pt. mascarinus and those species of Pteropus (tonganus,
Gouldit, and conspicil/atus) inhabiting Australia and the
islands of the Pacitic Ocean.
The limb-bones call for no special description, the most
perfect specimen being the right femur, which measures
40 mm.
Since the above was written a few additional bones referable
to this new species, and representing two younger individuals,
have been received. ‘Their fragmentary condition has,
however, failed to add any additional particulars to the above
description of this interesting and lost species.
On Brachiopod Nomenclature. 223
XXXVI.—Brachiopod Nomenclature: Seminula, &c.
By 8. 8. Buckman, F.G.S.
In this Magazine (vol. xviii. 1906, p. 324) I put forward
certain views regarding the genus Seminula. In the same
Magazine (vol. xix. 1907, p. 194) Dr. Vaughan contested
my conclusions. I have also been favoured with certain
verbal criticisms concerning them.
The gist of the verbal criticisms may be given first. They
are to this effect—that Terebratula pentaédra, Phillips, ought
not to be taken as the type of Seminula: that M‘Coy, in
using a trivial name as a generic term, indicated exactly the
type of his genus: that, therefore, 7’. semznula is the type of
Seminula: that M‘Coy himself subsequently confirmed this,
as Dr. Vaughan points out. To which I may add that, if
the type was considered doubtful before, then M‘Coy becomes
the first one to select a type to his genus; and that therefore
subsequent authors are barred from selecting outside his
limits,
The difficulty in this case is that M‘Coy himself, when
he made this selection, confused as Seminula seminula speci-
mens of Dielasma; but we have it on Davidson’s authority
(Carb. Mon. p. 16) that the original of Seminula pisum, as
M‘Coy called Phillips’s Zerebratula seminula, is a Rhyncho-
nella [Camarophoria]. As that is what M‘Coy originally
had in his hand in naming his species and genus, then if the
views prevail that M‘Coy’s selection of a trivial name for
a generic is a better indication of his type than his giving
a figure, the type of Seminula, M‘Coy, will be 7. pisum=
Ter. seminula. The result will be the same as in my previous
paper—that Semenula is a genus akin to Camarephoria.
Now as to Dr. Vaughan’s observations on Ter. pentaédra.
He says that the type of this species is in the British
Museum; but I had come to the conclusion that this was
not the type. This alleged type Dr. Vaughan says is con-
specific with S. ambiguus: I tind so many differences that I
cannot regard it as congeneric. ‘lhe most important point
is the contour of the beak-region. In S. ambiguus the dorsal
umbo is not prominent, and on each side of it the two valves
join flush: it has a thorough Terebratuloid contour. In
the “ 7. pentaédra” the beak-region has what may be called
a spiriferoid contour: the umbo is very prominent and the
two valves join to make a flange each side of it, features
which are seen in Spiriferids. ‘These same features I tind
in the specimens accompanying the alleged 7. pentaédra :
224 Mr. 8. 8S. Buckman on
eight of these Dr. Vaughan admits are fringed Athyrids—
and I agree. I claim, however, that the alleged 7. pentaédra
is also a fringed Athyrid in imperfect preservation ; and
Dr. Vaughan’s statement (p. 196) that the remaining speci-
men in the series, which he says “‘ approaches closely to the
[alleged] type,” does exhibit glabristriation supports this
view. My contention is that these two specimens supplement
one another; that they belong to a series of globose fringed
Athyrids not yet generically distinguished ; that they are
allied, as the characters of their beak-regions show, to the
glabristria- Roysii forms; and that they are generically
separable from S. ambiguus by their beak-region characters,
J have examined many specimens of S. ambiguus, some of
which Dr. Vaughan kindly sent me; and the terebratuloid
contour of the beak-region is very distinctive.
Composita.
The terebratuloid appearance of S. ambiguus struck
Sowerby (Min. Conch. iv. p. 105), and the combination of
Terebratula and Spirifer characters in the shell caused him
to give a hint about constructing a new genus for it. Brown
took the hint, and emphasized the composite character in
his name. Dr, Vaughan says (p. 197) that Brown’s figures
represent Spirifer glaber: he gives as reasons the large size,
the shape, and other characters. Brown’s figures, however,
are exactly the same size and shape as the larger of
the syntypes figured in Sowerby’s plate: in fact, Brown’s
figures are obviously made out of the details given by the
four figures of Sowerby—the size and shape are taken from
the larger figures, and the characters of the smaller figures
have been enlarged to fit. Brown’s fig. 4 (Foss. Conch.
pl. liv.*) is obviously based on a tracing of the middle figure
of Sowerby’s plate: then the valve has been depicted from
the outside—the details, even to a bit of coil seen through a
break, being taken from the N.H. fig. of Sowerby’s plate
(Min. Conch. iv. pl. 376).
It is hardly necessary to pursue any further the idea that
Brown figured S. glaber in this case; but in his pl. li. it
may be seen how differently he did represent it.
Type Specimens.
Scepticism with regard to the identity of alleged type
specimens is necessary, as I have shown before+. A case
ii point now concerns a Carboniferous species. In the
t+ Ann, & Mag. Nat. Hist. (7) vol. xiv. p. 392 (1904).
Brachiopod Nomenclature. 225
Sowerby Collection (British Museum, Natural History) under
No. 43464 are four specimens—one in one box, three in
another. The one alone is said to be the figured specimen
of Anomites crumena, Martin (Petrif. Derb. pl. xxxvi. fig. 4) ;
but Martin’s figure is coloured light ochre, while this is a
blackish-grey fossil of much smaller size and with less
marked costee. In the pedicle-valve of this blackish speci-
men I cannot find any mesial septum. It has the appearance
of a Lower Lias Rhynchonella, and it is possibly the example
mentioned by Sowerby as from Pickeridge (Min. Conch. i.
. 190).
: Of the three specimens in a box, one is claimed as the
original of the example of J’. crumena figured by Sowerby
in fig. 3 of pl. 83. This and another specimen in the box
may both have supplied details of what is perhaps a com-
posite figure—what Schuchert calls a synthetograph *. But
these three specimens are not from Mountain Limestone as
claimed: they are from Middle Lias Marlstone and are the
well-known Rhynchonella northamptonensis, Walker. David-
son’s Ool. & Lias. Brach., Suppl. pl. xxix. fig. 8, represents
them exactly.
The T.-globata series.
The Inferior-Oolite and Fuller’s- Earth species, which
hitherto have been designated by the above term, form a
remarkable group ; but their identification with Terebratula
globata is erroneous. It is necessary to revise.
Terebratula globata, J. de C. Sowerby.
1823. Min. Conch. pl. 436, fig. 1.
An examination of the types of the species shows that the
identification usually made, on the lines of the specimen
figured as 1’, globata by Davidson in Ool. & Lias. Brach.,
Suppl. pl. xvi. 3, is quite incorrect. Sowerby’s species is
a very globose, almost uniplicate, barely biplicate shell, not
at all well depicted by Davidson, Ool. Brach. pl. xiii. 2, 3.
Sowerby’s species is the shell which the late J. F. Walker
has for years distinguished and distributed by the MS. name
of a village near Frome: that will be a guide to its identifi-
cation in many cases.
I suspect that Dav. Suppl. pl. xvii. 5 is really 7. globata
and not 7. bullata. These two species are remarkably
alike: they are isochronous homeeomorphs—members of two
* “Catalozue of Type Specimens,” Bull. U.S. Nat. Mus. vol. liii.
p- 15 (1905),
226 On Brachiopod Nomenclature.
families. 1. globata has much the appearance of my J’. with-
ingtonensis *, but is much more tumid. It has the same
peculiarly truncate beak.
This identification of T. globata leads to the following
change of name :—
Terebratula nunneyensis, nom. nov.
1878. T. globata, Dav. (non Sow.) Ool. & Lias, Brach., Suppl.
pl. xvii. 3.
_ Much more plicate, but much less tumid than 7. globata.
Common in the Fuller’s Earth.
Various Cotteswold Inferior Oolite Terebratule were
identified by Davidson with 7. globata ; but of late years it
has generally been recognized that they themselves require
to be separated as well as parted from 7. globata=T. nun-
neyensis. They and TZ. nunneyensis belong to the same
group; but the true 7. globata belongs to quite a different
series—that of T. spherotdalis.
Terebratula cotteswoldensis, nom. nov.
1878. T. globata, var., Davidson (non Sow.), Suppl. pl. xvi. 1.
Like 7. intermedia, Sow., but more plicate and much more
tumid. Common in Clypeus-grit of the Cotteswolds.
Terebratula cheltensis, nom. nov.
1878. 7. globata, Davidson (non Sow.), Suppl. pl. xvii. 2 (type); 1851,
pl. xiii. fig. 7.
Oppel (Juraf. p. 497) notes how Davidson’s pl. xiii. 7
differs from his 7. Fleischert. ‘The other figure cannot repre-
sent one of Oppel’s types, for he does not mention Cheltenham
in his list of localities, and 7’. Fleischer’ belongs to the
Cornbrash.
Common in the Clypeus-grit of the Cotteswolds.
T. birdlipensis, Walker, of which Dav. Suppl. pl. xvii. 18,
may be taken as type, and 7. tuméda, Dav., mentioned in
Suppl. p. 149 as 7. globata var. tumida, are two more forms
of what used to be called the globata-series. Presumably the
specimens depicted in Ool. Brach. pl. xii. figs. 5, 6, are
what Davidson intended as 7’. tumida: Leckhampton and
Cheltenham are really terms for the same locality.
* Proc. Cotteswold Club, xiii. p. 246 (1901).
On some Freshwater Amphipods. 227
XXX VIT.—On some Freshwater Amphipods : The Reduction
of the Eye in a new Gammarid from Ireland. By Prof. Dr.
Fr. VEJDOVSKY *.
[Plates XI. & XII.]
THE paper by W. F. de Vismes Kane (23) on the Amphipods
of the Irish lake, Lough Mask, is worthy of attention in
many ways. It records that the author collected there about
130 specimens of Niphargus Kochianus, among which there
were three ‘* which had well-ileveloped pigmentation and a
fourth which showed a cloudy shading in the optic region.”
As I had already stated (22) that in “ N. Kochianus” rudi-
ments of eyes without pigment are present, I was obliged to
devote special attention to the work of de Vismes Kane,
according to which pigment was still actually to be found in
certain individuals.
Mr. de Vismes Kane, with the greatest kindness, placed at
my disposal, for purposes of investigation, a portion of his
collection of the above-mentioned Gammarids from Lough
Mask, including examples of the wholly blind Niphargus and
also the four individuals with the eye-flecks which had been
preserved in formol. This valuable material, for which I
here wish to thank Mr. de Vismes Kane, proved to be of
great importance for several reasons, viz. :—
(1) Because it furnishes evidence as to the extent of the
first stage in the reduction of the compound eye ;
(2) Because by reason of this degeneration it is possible to
explain the significance of the ‘ pigment-veil” [Pigment-
schleier] already known in Crangonyx subterraneus ; and
(3) Because, lastly, it offers the possibility of deciding
the relationship of the freshwater shrimp [Flohkrebs] found
in the wells of Munich, usually alluded to as Niphargus
“ Kochianus,” with the similarly designated species from
England and Ireland.
I. On BATHYONYX, gen. nov.
In his paper de Vismes Kane refers all the Gammarids
found in Lough Mask to a single species, viz., Niphargus
Kochianus, Bate. I have been able to confirm this identifi-
cation with the exception of those four individuals with eye-
* Translated by D. J. Scourfield from the ‘Sitzungsberichte der
koniglichen bohmischen Gesellschaft der Wissenschaften,’ No, 28.
Prag, 1908.
228 Prof. Dr. Fr. Vejdovsky on
pigment, which in no way possess the characters of the genus
Niphargus, but are representatives of a new genus inter-
mediate between Crangonya and Gammarus, which I
designate Bathyonyz. ‘To the species I give the name of
B. de Vismest, gen. nov., sp. n., in honour of the discoverer.
Generic and specific characters :—Bathyonyx with both
patrs of antenne very short (the flagellum of the first usually
6-jointed, that of the second 4-jointed), with 2-jointed secondary
branch on the first antenna, Eyes consisting of crystalline
cones diffusely distributed in the pigment. Gnathopoda with
weakly developed hands ; the hands of the anterior gnathopods
broader than those of the second pair. Telson split for two-
thirds of tts length, consequently bilobed. Third pair of uro-
pods with an outer 2-jointed, and a short inner \-jointed
branch, scantily furnished with sete, as in Crangonyx. Seg-
mental capsular epidermal sensory organs fusiform.
Bathyonyx de Vismest, gen. et sp. nn.
The four specimens examined were all of equal length,
namely 3 mm., and of similar form. Being preserved in
formol they still showed traces of the original pale orange-
red coloration which was especially noticed by de Vismes
Kane in the living condition.
The antenne are remarkably short, attaining about one-
fourth of the body-length ; there is no well-marked distinction
between the joints of the peduncle and the flagellum, for the
three basal joints pass gradually into those of the flagellum.
In this way the total number of joints of the first antennze
amounts to nine in two specimens and eight in the third.
(The fourth example was cut into serial sections.) The
third joint carries the 2-jointed secondary branch, of the
same form as in Niphargus for example, 7. e. with an
elongated and greatly swollen basal joint, and a short and
slender distal joint (Pl. XI. fig. 2,2). The antennal sete
are weak, short, and scanty, from 1-4 on each joint as in
Niphargus. The sensory sete, which I have described from
the antennee of Crangonyx as sensory brushes [Sinnespinsel],
are also present in Bathyonyx, but they are very feebly
developed, so that they can be easily overlooked.
These remarkable sense-hairs are characteristic not only of
these genera, but also of Viéphargus and Gammarus (especially
those of the first joint of the first antennee), occurring, in fact,
in the last-named genus in the greatest number.
Thus in G@. fluviatilis from Herzegovina (collected by
Dr. Thon) I invariably found seven sensory brushes on the
some Freshwater Amphipods. 229
distal outer edge of the first joint in the space between the
large ordinary sete. In addition, there are in the middle of
the same joint two further sensory brushes, accompanied by
a thicker ordinary seta, exactly as in Crangony«.
In Bathyonyx also there are always two sensory brushes
in the middle of the first joint and four on the distal edge.
Their structure is the same as in Crangonyx, Niphargus,
and Gammarus. Hach sensory brush consists of a basal
cup or urn sunk into the fibrous inner cuticle. (This fibrous
inner cuticle is covered with a thin homogeneous outer
cuticle, Pl. XI. fig. 14, c).
From the cup there arises the stem of the seta, which, at its
distal end, is furnished with two lateral rows of fine hairs
(Pl. XI. fig. 14). I have designated these sense-hairs in
Crangonyx as “ quaking-hairs ” [Zitterhaare], because in the
living state they are found to be subject to periodical
vibratory movements, which points to some definite sensory
action. ‘he quaking movement of these hairs of Amphipods
is certainly worthy of remark, especially as it resembles
ciliary action, although produced in a different manner from
the latter. The quaking of the sense-hairs reminds one
rather of the flame-cells [Zitterorgane] of the Rotatoria, or
of the tufted hairs [Btischelhaare] which I have described in
Bothrioplana bohemica *.,
By the examination in profile of the transparent first
antenna it is easy to demonstrate that the sensory brushes
are supplied with nerves from the antennal ganglion. Long
and fine processes from the ganglion-cells are connected with
special club-shaped cells, the short process from each of which
is spread out like a dish at the base of the cup from which the
seta arises (Pl. XI. fig. 14, ne). The structure of the ganglia in
the antenne of the Gammarids is, according to what I have
so far been able to demonstrate in Gammarus pulex, &c.,
very characteristic and deserves special attention from com-
parative nerve histologists.
On the lower antenne the sensory brushes only occur
singly ; thus I have observed two on the second joint and
one each on the third and last joints of the same torm and
size as on the first antennee.
I only found the hyaline clubs, so-called olfactory clubs,
on one specimen (PI. XI. fig. 3); they are very short, not
attaining the length of the joints, and therefore very difficult
to observe.
* “Zur vergleich, Anatomie der Turbellarien,” Zeit. f. wiss. Zool.
Bd. Ix. (1905).
230 Prof. Dr. Fr. Vejdovsky on
Sensory brushes are present on the telson * as well as on
the antenne, as has already been shown in many Amphipods.
Their position upon the surface of the telson appears to be
characteristic of the species of Niphargus and Gammarus.
In Bathyonyx the telson is split for two-thirds of its length,
thus being bilobed posteriorly, the lobes gradually narrowing
and each being furnished at the point with a long and a short
simple seta (Pl. XI. fig. 13).
In addition there is always present here a short sensory
brush. Somewhat anterior to this group of hairs and towards
the exterior edge there are always two longer sensory brushes
of the same form and size as on the antenne. The inner-
vation of all the sensory brushes on the telson evidently
proceeds from the lateral nerves of the last ventral nerve-
cord ganglion, as I have been able to demonstrate, for
example, on the telson of the New Zealand species Cran-
gonyx compactus, Chilton.
Quite different in structure from the sensory brushes are
the segmental sensory capsules, which, among different
Gammarids, exhibit a form typical for the genus. By means
of these sense-organs the genera Niphargus, Crangonyx, and
Gammarus can be determined with perfect safety. But they
are somewhat difficult to find, especially when only a few
occur on the segment, as they are characterized by their
extremely minute dimensions. It is only on this account
that I have not made an earlier reference to their occurrence
in Orangonyx. I refer to the capsules as segmental sense-
organs because they are distributed partly on the body-
segments and partly on the epimera. The form, number,
and distribution of the capsules can be seen without much
difficulty in preparations of the second epimera, where they
appear regularly disposed in a semicircle on the margin, and,
in consequence of the sharply contoured cuticular walls, they
stand out as plainly in the living animal as in preparations.
The sense-hairs, too, which are connected with the capsules,
make the recognition of the latter easier.
Among the species of the genera already mentioned I have
made out the following forms of sensory capsules :—
In Niphargus they are mostly flask-shaped, gradually
narrowing towards the exterior, where they pass into the little
sense-hair. ‘The sense-hairs are mostly plumose, or like a
paint-brush, rarely simple and pointed.
I have examined the sensory capsules of various species of
* Chilton has also observed them on the penultimate joint of the
fourth pereiopods in Gammarus fragilis.
some Freshwater Amphipods. 231
Niphargus as regards shape and have been able to determine
the following forms :—
Niphargus puteanus from Prague (Pl. XII. figs. 23 & 24).
Sensory capsules 12 w long and 4 w broad, with usually a
short sense-hair ending in a tuft. Less frequently the sense-
hairs are simple, not branched.
Niphargus elegans from Modena (PI. XII. fig. 22). Sensory
capsules of the same form as in N. puteanus from Prague,
but the sense-hairs have forked ends with a long tuft on one
side,
Niphargus tatrensis (Switzerland, fig. 26), more strongly
swollen, with a simple sense-hair ending in a thread.
A Niphargus from Lille has very small sensory capsules
with sense-hairs of the paint-brush type (fig. 27).
In Niphargus Caspary, from Munich, the sensory capsules
are very pale, thin-walled, and short. They give rise toa
simple short sense-hair (fig. 28).
Niphargus Kochianus (Lough Mask in Ireland) is provided
with nearly spherical sensory capsules narrowed towards the
exterior and tufted sense-hairs (fig. 29).
In Crangonyx subterraneus, great numbers of which I have
been able to examine recently from the point of view of the
sense-organs now in question, the capsules are almost
cylindrical, longer than broad, the sense-hairs long, not
plumose ; sensory capsules 4—5 w long, sense-hairs 20 uw long
(fig. 30).
In Crangonyx compactus, from New Zealand, the sensory
capsules are very slender, terminating with a simple hair
(fig. 25).
In all the species of Gammarus the capsules are, in the
main, spindle-shaped, slender, pointed exteriorly, not infre-
quently constricted in the middle, the sense-hairs short, not
plumose. The length of the capsules varies, according to the
species, from 16-22 w (figs. 20 & 21). The same details of
the form of the capsules apply, in general, to Bathyonyz also,
but the structures are much smaller and difficult to find
(figs. 17, 18, spo, spo’).
'The internal structure of the sensory capsules differs from
that of the sensory brushes, but is the same in all genera, as
can be demonstrated with the greatest precision on the larger
forms occurring in the Gammarids. ‘The proximal portion
of the cuticular wall of the capsule turns inwards and forms a
hollow axial stalk, to which the sensory seta is attached. In
some species of Gammarus and in Crangonyz the distal end
of the stalk, 2. e. where the sensory seta arises, is thickened
into a little head. It is not difficult to make out that the
932 Prot. Drs Er. Vejdovsky on
capsules are innervated from the cutaneous nerves, for a pale
but sharply contoured nerve-thread approaches the base of each
capsule and then clearly passes along the stalk to be joined
to the sensory seta. ‘The nerve-fibre always originates from
one ganglion-cell, as is accurately shown by Hamann.
The sensory capsules are found on all the segments. On
the cephalic segment of Bathyonyz, close behind the diffuse
eye, there are 4-5 capsules arranged in a curved line
(fig. 17, spo), and not far behind there is a second row of
3-4 capsules (fig. 17, spo’). On the following segments
the number may be less, but they occur more or less
regularly also on the hinder segments.
In Crangonyx I always found, in the hinder region of the
body, four capsules on each segment, two anterior and two
posterior, while on the anterior segments they were rarer and
Jess regularly distributed, although one must take into account
the circumstance that the structures can easily escape
observation by reason of their minuteness and their usually
small refractive power.
The attention of earlier investigators was very often given
to the sensory capsules. Thus La Valette observed them on
the back and therefore referred to them as “ capilliin corporis
dorso siti.”” Humbert recognized them as sense-organs and
designated them accurately as ‘‘ capsules sensitives.” As
Hamann justly remarks, Leydig combatted the correctness of
this expression, “for it is not a question of capsules but of
modified cuticular canals. But from his description it is
evident that he classed these organs with the similarly formed
hair-structures of the antenne and only examined the latter.”
Vom Rath and more recently Hamann have correctly described
and figured the sensory capsules (referred to by Hamann
as sensory clubs). Della Valle calls the organs “ peli,” but
his figures are reproduced on a small scale, so that the
structure of the capsules, nerve-endings, and sensory sete do
not clearly appear.
The most remarkable organs of Bathyonya are undoubtedly
the EYES, and for this reason I propose to describe them in
detail. Not one of the four above-mentioned specimens has
normal compound eyes such as are found in Gammarus, but
visual organs with scattered components on both sides of the
head. Unfortunately, owing to the fixing agent, the pig-
ment was not perfectly preserved, so that it is “impossible to
state exactly the number of the pigment-cells.
One example was cut into a complete set of serial sections,
principally with the object of showing more precisely the
relation of the nerves and optic ganglia to the external visual
some Freshwater Amphipods. 233
apparatus. Unfortunately the preservation in formalin had
so much damaged the internal nervous structure and the
brain that nothing definite can be said in this connection. It
is only possible to refer to the superficial appearance of the
eye, and in addition at most to the position of the dioptric
elements under the hypodermis as observed in preparations of
the entire animals under moderate and high-power magni-
fication. But even in this way interesting results are
obtained.
In general the eye of Bathyonyx is not a compact organ,
such as we are accustomed to picture the compound eye of
Arthropods, but forms a large irregularly defined fleck of
unequal size with a variable number of crystalline cones in
each of the available specimens. I figure such eyes in
figs. 16-18.
In fig. 16, which is drawn under a low magnification,
there are twenty-four cones. They occur in a dark (brown
to brownish black) fine-grained pigment which appears to be
most thickly developed in the centre of the eye.
Here was evidently situated the original eye, as, for example,
in Gammarus. In this central pigment most of the crystal-
line cones are collected, without, however, forming a compact
organ.
They are as irregularly distributed in the pigment, without
any definite position, as the peripheral crystalline cones,
which mostly lie under the ordinary hypodermis without any
pigment. All the crystalline cones are commonly composed
of two segments or simply constricted, the hemispheres being
sometimes equal and sometimes unequal in size. Now and
again at the periphery small wholly isolated cones are also
found. The crystalline cones consist of a strongly refractive
homogeneous brownish substance, and therefore stand out
clearly in the preparations.
In this condition the individual cones appeared in all four
specimens. Another eye, as seen under strong magnification
(as with Hom. Imm. Apochr. Olj. 2 mm., Oc. 4) and showing
interesting details in the structure of the cones, is reproduced
ime Te
Here the scattered components of the eye are still more
striking ; in the centre of the eye-fleck one finds large, simple,
rarely double, hexagonal or irregularly branched pigment-
cells in the midst of whose granular substance a small rounded
nucleus can be detected (p). The whole system formed
by the cones and the central pigment-cells retains in the
main a reniform shape, as is also typical of the compound
eyes of Gammarus. In Bathyonyzx, however, there exists no
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 16
234 Prof. Dr. Fr. Vejdovsky on
connection between the individual cones; they are isolated and
mostly situated at the periphery of the pigment-cells. The
total number of the cones in the specimen represented amounts
to only fourteen, and the variable form and size of the com-
ponents can be well seen in the drawing.
The number, form, and size of the crystalline cones vary,
however, not only in different individuals, but also in the
right and left eyes of one and the same animal. The eye
just described belongs to the left side of the head (fig. 17).
In fig. 18 is shown the eye from the right side of the same
animal. In the centre of the pigment-cells there is only one
crystalline cone; the others are situated at the periphery of
the pigment-cells, irregularly scattered, and two even lying
not far from the frontal margin of the head. In this case
there are only eleven crystalline cones of the most diverse
size and form.
The structure of the crystalline cones here reproduced is
certainly very peculiar and difficult to correlate with the
details of an ordinary compound eye of Gammarus. In each
cone there is a dark, finely granular, sharply defined matrix,
at whose margin, when in a favourable position, a little body
ean be seen which I am inclined to consider as a nucleus.
In this matrix an irregular number of smaller rounded bodies
are present. In the simplest case there is a large central
sphere and with it one or more little spheres. There can,
however, be such an increase of both kinds that the matrix
appears to be filled with vacuoles. These inner vacuolar
spherical bodies are filled with a homogeneous slightly re-
fractive substance which is only stained by carmine in a very
feeble and diffuse way.
In the above-mentioned serial sections into which one
specimen was cut, I have unfortunately found little explaining
the connection of the appearance in side view of the crystal-
line cones, as described, with the nervous organization. In
fig. 19 one such section is reproduced. Nothing is here to
be seen of the nerves. The cuticle of the head (c) stands
off a long way from the eye. The hypodermis (/p) forms a
thin layer which contains only the regularly placed nuclei.
Underneath there are three groups of large pigment-masses
without nuclei, between which occur rounded vacuolar bodies.
These are the rudiments of the crystalline cones, and they
are not nearly so refractive in these sections of 5 mw in thick-
ness as in the preparations showing the animals from the
side. A direct connection between the outer larger pigment-
masses and the deeper layer of smaller pigment-cells (r¢)
does not exist in the serial sections.
some Freshwater Amphipods. 235
Between the two layers there is a hollow space, whether
artifically produced or not I am unable to decide. But
according to the known structure of the eye in Gammarus
we should be dealing, in the lower pigmented layer of cells
arranged almost like epithelium, with retinula-cells, to which
also the outer pigment-masses belong, they being certainly
only separated by the unsatisfactory method of fixing. From
this it would seem that in Bathyony« the retinula apparatus
was still retained, although the dioptric elements were
gradually destroyed. For the crystalline cones described
above must be considered as being in process of degeneration :
firstly, because they do not form a single compact eye, but
are only loosely scattered under the hypodermis, and, secondly,
because the actual substance of the normal crystalline cone is
gradually degenerating.
Special investigation of the mouth-organs of Bathyonya
shows that they resemble in form those of Crangonyx, except
that they are much more feebly developed, thus corresponding
to the general organization of Bathyonyx. he drawings of
the mandibles and their palps (fig. 4), of the maxille of the
first (figs. 5 & 6) and second pairs (tig. 7), and, lastly, of the
maxillipedes (fig. 8) entirely support this conception. Tie
palpi of the maxillipedes are almost as long as the gnathopods,
which are also very weakly developed, especially the hands,
which, under low magnification, can scarcely be distinguished
from the almost equally broad or even broader proximal joints
(carpopodites). The form of the hands of the gnathopods of
both pairs reminds one of Crangonyz, but in general in this
case, as in Crangonyx, there is some variability. In order
to show this I give in figs. 9-11 camera drawings of the
hands of three specimens. Figures 10 and 11 are shown
under the same and fig. 9 under a slightly higher magni-
fication. From a comparison of the three it is apparent that
the only character common to all is that the hands of the
first pair are obviously broader and shorter than those of the
second pair. It can also be seen from the drawings that the
oblique outer edge of both hands is finely toothed and that the
armature of strong sete which is characteristic of the hands
of Gammarus, Crangonyz, and in part Niphurgus (as I have
specially remarked in connection with Crangony2#) is entirely
absent in Bathyonyx. In other respects a slight variability
in the form of the hands is to be observed in all three cases,
which is also true of Crangonyx. A comparison of the form
of the gnathopod hands of the last-named genus, as I have
figured them in my paper (21, figs. 12-14), with those figured
>A
(ay
236 Prof. Dr. Fr. Vejdovsky on
by Chilton (7, fig. 4, gn, gn”) might easily lead to the view
that the Crangonyse found in Bohemia in a certain measure
represented a different species to that found in English wells.
Indeed Stebbing, on the ground of my representation and his
own experience, has considered that the Cr ‘angonyx found at
Radotin near Prague belongs to a distinct genus and species
“ Hucrangonyx Vejdovskyt. 4 Only after comparison of the
original specimens from Bohemia and England, which we
made in Prague with Chilton, was it demonstrated, beyond
all doubt, that we were dealing with one and the same species,
in which the form of the gnathopod hands in different
individuals is subject to a noticeable variability.
I have been able recently to convince myself that this is
actually the case by an examination of numerous specimens
of Cranyonyx which were collected in the course of last year
in great quantities in wells at Podbaba, near Prague, by my
energetic pupil Herr Niessner.
I have not specially investigated the other feet, as, in the
main, they resemble in form the corresponding extremities of
Crangonys. The little double hooks on the thighs of the
first and second pleopods also differ only in a small degree
from the same structures in other Gammarids, especially
Gammarus (fig. 15). Further, the last pair of uropods
correspond with those of Crangonya; they are rather short,
the basal joint almost as high as broad; the outer branch
consists of a long proximal and a short thin distal joint
(fig. 12).
The proximal joint bears on its side only one or two sete,
and differs essentially in this respect from the corresponding
joint of the Niphargd and Gammarids, which is armed with
tufts of strong sete.
The inner branch is 1-jointed and reaches more than half
the length of the outer branch. Moreover, it is comparatively
longer than in Niphargus and shorter than in Gammarus. 1
must, however, remark that I have only examined these
wropods i in two specimens.
From the point of view of general biology the discovery of
Bathyonyx is very important. So far as its systematic
position 1s concerned, the genus comes between Gammarus
and Crangonyz, as it agrees with the former in its capsular
segmental sense-organs and with the latter in the rest of its
organization. Only the absence of the so-called secondary
gills, which are so characteristic of Crangonye (and Boruta),
and the two-lobed telson separate Bathyonyx trom the genera
some Hreshwater Amphipods. 2370
mentioned and bring it in some respects near to Gammarus
and in others to Niphargus.
According to de Vismes Kane, Bathyonyx was obtained
from a depth of from 130 to 150 feet in Lough Mask. From
the general habitus of its body and most of the details of its
organization it may be regarded as a degenerate form, a
conception which is strongly supported by the sense- orgs ans,
especially the eyes. All the above-mentioned cuticular
sense-organs, as the sensory brushes and capsules, are present
in all species of Gammarus living in ordinary tresh waters,
but in much greater number and development than in Bathy-
onyx, where they are subject in both directions to a reduction
corresponding to that of the eyes, and cannot therefore be
considered as compensatory organs. The ancestors of Bathy-
onyx evidently possessed the same sense-or gans as, for
example, the common Gammarus, and in the same number
and development, but they degenerated in the course of time
in the depths of Lough Mask. There is no reason for sup-
posing that these crustaceans reached the lake in the water
from the springs ; their organization strengthens us rather
in the opinion that the progenitors are to be sought perhaps
only in a species of Crangonyx or Gammarus which graduall
adapted themselves to life at the bottom of Lough Mask.
Although species of Crangonyx possessing eyes are known,
among which especially C. recurvus, according to Grube,
lives in Lake Vrana, on the island of Cherso, unfortunately
its organization has not been hitherto carefully studied *,
Irom what has been said it appears that we must attach the
greatest importance to the eyes.
The gradual adaptation to life in the darkness of deep
lakes and subterranean waters generally produces the result
that the organs of sight are gradually, not suddenly, reduced,
until at length the animal appears quite eyeless, and trans.
mits its blindness to the following generations. So far as the
freshwater Amphipoda are concerned, we now know a series
of cases in which we must conclude that eyes were formerly
present. Reference has often been made to the observation
of Leydig that certain individuals of Niphargus puteanus were
provided with eye-pigment just in that part of the head where
the eyes are situated in such a form as the common
Gammarus.
* From an interesting epaten by M. Grochowski (11) T learn that he
and Professor B. Dybowski found in 1895 a large number of C. recurvus
in the lake mentioned. A special account of this species therefore may
be expected,
238 Prof. Dr. Fr. Vejdovsky on
Although in the course of thirty years I have investi-
gated from this point of view hundreds of examples of the
species mentioned, I have not been able to find anything of
the pigment in question in a single individual, and I believe
therefore that we are driven to the conclusion that Leydig
only had Crangonyx before him. In this genus I have in-
variably found, not only in examples from Radotin, near
Prague, but also recently among great numbers of individuals
from the wells of Podbaba, near Prague, pigment-flecks on
both sides of the head, consisting of large branched cells such
as I have described in an earlier paper.
Very interesting is the further statement of Moniez (14),
from which it appears that he observed a ‘Gammarus fluvia-
tiis”’ in the drinking-water of Hmmerin, near Lille, whose
eyes appeared in the form of dark flecks,
They were not so compact as in the normal freshwater
shrimps, but appeared to consist of single ommatidia sur-
rounded by black pigment, and the crystalline cones were not
so round as in the typical species. Moniez therefore described
the form he had observed as G. fluviatilis, var. d’Hmmerin.
According to the description we should have here a case of
rudimentary eyes exactly corresponding to what we have
specially described in Bathyonyx, and it is to be hoped that
the Emmerin variety may be subjected to an examination as
to its other characters in order to see whether it may not
represent a distinct species closely allied to ,Bathyonyx
de Vismest.
Not less important is the communication of R. Schneider
(19) about the Gammarus pulex living in the underground
waters of Claustal, in which the author found the eyes to be
in a peculiar condition. They are of irregular form, without
detinite outline, with a little blackish pigment which is con-
fined to the centre of the eye, disappearing towards the
periphery. The crystalline cones are separated from one
another, and the eye is therefore diffusely formed as in our
Bathyonyx. Ik. Schneider designates the form as G. pulex,
var. subterraneus *.
According to Garbini, the G. fluviatilis observed by him
near Verona exhibits the same arrangement with regard to
the eyes as Schneider’s G’. pulex, and he calls the form
G. fluviatilis, var. monophthalmus. ‘The same author mentions
also NV. eleyans, var. imperfectus, with small brownish pigment-
* (The original description by Schneider is given by Prof. Vejdovsky
in a long footnote, but it has not been thought necessary to reproduce it
in this translation.— 7Zraxslator’s note. |
some Freshwater Amphipods. 23
flecks, like those I have referred to in connexion with the
specimens from Modena.
The large pigment-flecks of Crangonyx subterraneus, which
I have described as a pigment-veil [‘ Pigmentschleier els
suggest by their position the former presence of eyes; L
have not, however, referred to them as rudiments of eyes so
long as ‘there was no definite proof that the pigment was
directly connected on the one hand with the crystalline cones
and on the other with the optic ganglia.
We know now, however, that on the one side there are
species of Crangonyx with normal eyes, and ou the other that
the eyes of C. compactus from New Zealand are, according
to Chil ton, only represented by two or three little lenses ””
without pigment. If we turn to C. subterraneus, we must
regard the pigment-veil as a rudiment of an eye in which the
crystalline cones have completely disappeared and only the
pigment-cells remain. We find, therefore, that the genus
Crangonyx is characterized by visual organs in all possible
stages of reduction; and we might expect to find similar
series of degenerating eyes in other genera. For Gammarus
I have already mentioned the observations of Moniez and
Schneider ; a completely eyeless species G. fragilis, has
been described from New Zealand by Chilton, and I myself
know a large species from Herzegovina the two examples of
which in my possession lack all trace of eyes.
The same series may be made out in Niphargus. It is
true that species with normal eyes are not known, but
N. elegans, which is characterized by possessing only the
eye-pigment, permits of the assumption that there are species
with eyes, and there exists a whole series of completely blind
species, as N. Kochianus, Caspary, puteanus, &c. (The work
of Viré has not been accessible to me.)
Now it is possible that there are eyeless forms which oceur
at the same time and in the same place with those possessing
reduced and normal eyes. At least “ Gammarus pulew,”
from Wiadenschwyl in Switzerland, deserves a renewed
investigation, as, according to the statement of Asper, it is
represented at a depth of 40 m. by individuals with and
without eyes. This case of variable development of the
visual organs in different individuals of the same species is
contirmed among other species of animals by Packard, and
Forel particularly notes that in rare and exceptional instances
the blind Asellus Foreliz, Blanc, still bears vestiges of eyes ;
and in this connexion it may be remembered that years ago
I found in a well in Prague both eyed and eyeless forms of
the rhabdoccel Gyrator notops, Duges. ‘The eyeless form
240 Prof. Dr. Fr. Vejdovsky on
has been described by L. v. Graff as a distinct species—
Gyrator coccus.
The foregoing cases may support the opinion expressed by
Packard, and more recently by Chilton, that it is possible
that different species, and even individuals from different
localities, may exhibit different stages in the reduction of the
eyes.
The discovery of Bathyony# offers now excellent evidence
as to the probable first stage in the degeneration of the eyes.
The ancestors of this genus were certainly closely related
to a Gammarus with normal eyes, as may be seen from
the whole organization and in particular from the capsular
segmental cuticular sense-organs. The visual organs also
represent the well-known kidney-shaped eyes of a Gammarus-
like ancestor. The descendants, adapting themselves to live
in the dark depths of the lake *, gradually lost the power
of seeing with the compound eye, as individual ommatidia
remained functionless and consequently the crystalline cones
became subject to degeneration. Only the pigment-cells
remained and the ommatidia which still retained their
functional power appeared then as scattered components of an
eye on each side of the head. It is in this stage that
Bathyonyx now presents itself to us.
But the crystalline cones still present have also undergone
degeneration, as is shown by the little homogeneous spheres
within the cones. The hypothesis of the progressive re-
duction of the eyes can, I believe, be supported by the
following facts :—
(1) From the variable number of the crystalline cones in
one and the same animal on the right and left sides of the
head.
(2) From the extremely variable size of the individual
crystalline cones, which in one and the same eye present
instances of diminution in size so as to finally appear simply
as little refractive particles.
(3) From the form of the crystalline cones themselves,
The homogeneous spheres within the fine-grained matrix
must, I think, be regarded as evidence of degeneration. In
this way the crystalline cones lose their dioptric property.
The degeneration, however, does not occur in any particular
order, but may affect widely separated ommatidia, while the
intermediate cones may remain intact.
In consequence of the foregoing, the originally compact
* From the small number of specimens obtained it may be supposed
that the species lives perhaps in the mud and only occasionally swims
about freely in the water.
some Freshwater Amphipods. JA1
eye breaks up into a variable number of scattered elements
which occupy a considerable space without any definite
arrangement.
From what has been said it appears that we must consider
the structure of the eyes in Bathyonys# as representing the
first stage of degeneration, and therefore this genus as the
first which could arise from the Gammarus-like ancestor with
normaleyes. ‘heancestry of the blind species of Gammarus
and of Bathyonyx may be represented as follows :—
1. Species of Gammarus with eyes.
Gammarus pulex, fluviatilis, &e.
So
2. The forms with re- 4. Species of Gamma-
duced eyes mentioned by rus without eyes (G. fra-
Schneider and Moniez gilis, Chilton),
(G. pulex vay. subterra-
neus,Schn., G, fluviatilis 3. Bathyonyx, Vejd.
var. d’Emmerin, Mon.), (B. de Vismesi, Vejd.).
An exactly similar series can be made out in Crangonya,
and adopting the same method as above the following result
1s obtained :—
1. Species of Crangonyx with eyes.
Crangonyx gracilis, recurvus, &e.
J a
2. C. compactus 3. C. subterra- _ 4. Blind species 5. Boruta
Chilt. (With 2 news, Bate. With inNorthAmerica, Wrzes, (ZB. ‘tene-
to 8 erystalline pigment-veil. brarum).
cones.)
For Niphargus also a corresponding series of species can
in all probability, be drawn up in spite of the fact that no
species bearing eyes are yet known. That such must have
existed, however, is proved by the species with rudimentary
visual organs.
The series in this case may be shown in the following
manner :—
1. Niphargus (hypothetical, species with eyes),
_ 2. N, elegans, Garb, (With 3, Eyeless species.
little pigment-flecks.) (2) With optic ganglia.
N. Kochianus, Bate.
N. Caspary, Pratz.
(6) Without optic nerves.
N, puteanus, &e.
242 Prof. Dr. Fr. Vejdovsky on
Literature.
(1) Asprr, G. “Beitriige zur Kenntniss der Tiefseefauna der
Schweizerseen,”’ Zool. Anz. ii. 1880, pp. 180-184, 200-207.
(2) Barr, C.Spence. Nat. Hist. Review, 1859, vi. Proc. Soc. p. 165.
Quoted by de Vismes Kane.
(3) Bare, C. Spencr, and Westwoop, J. O. “A History of the
British Sessile-eyed Crustacea,” 2 vols. London, 1863-1868.
(4) Barr, C. Spence. ‘Catalogue of the Specimens of Amphipodous
Crustacea in the Collection of the British Museum.’ London, 1862.
(5) Caspary, Ros. ‘“ Gammarus puteanus,” Verhandl. nat. Ver.
preuss. Rhein]. u. Westphal. Bonn, 1849, pp. 89-48.
(6) Cuizton, Cuas. “ The Subterranean Crustacea of New Zealand :
with some general Remarks on the Fauna of Caves and Wells.”
Trans. Linn. Soc. London. Second Series. Zoology, vol. vi.
1894, pp. 163-284, pls. xvi.-xxiil.
“The Subterranean Amphipoda of the British Islands,”
Journ. Linnean Society, Zoology, vol. xxviii. November 1900,
pp. 140-160, pls. 16-18.
(8) Detra VaLLtR. “ Gammarini del Golfo di Napoli,” Fauna und
Flora, Monograph xx. 1893.
(9) Fornt, F. A. “La Faune profonde des Lacs suisses,” Mém. de
la Soc. Helvétique des Sciences nat. vol. xxix. 1885.
(10) Garpini, ADR. “ Gammarus ciechi in aque superficiali basse,”
Acead. di Agricoltura Arti e Comercio di Verona, vol. Ixx. ser. iil.
8 pp.
(11) Cai M. “O rodzaju studniczka (Niphargus). Odbitka
z ezasopisma ‘ Kosmos.’” S 45 rycinamicynkograticznemi. Lwow,
1904, 75 S.
(12) Hostus, Aue. “ Ueber die Gammarus-Arten d. Gegend von
Bonn,” Arch, Nat. 1850, pp. 282-248.
(13) Lrypie, F. “ Amphipoden und Isopoden, Anat. u. zool. Bemer-
kungen,” Z. f. w. Z. xxx. Suppl. pp. 235-274, Taf. ix.—xii. 1878.
(14) Monrez, R. ‘ Faune des Eaux souterraines du Département du
Nord de la France,’ t. i. 1888-89, 68 pp. Lille, 1889.
(15) Packarp, A.S. “The Cave Fauna of North America, with Re-
marks on the Anatomy of the Brain and Origin of the Blind
Species,” Proc. Acad. Nat. Sci. Phil. vol. iv. pp. 8-156, pls. i.-
XXVil.
(16) Parker, G.H. ‘The Compound Eyes in Crustaceans,” Bull.
Mus. Comp. Zool. Harvard College, vol. xxi. no. 2, 3 pls. Cam-
bridge, 1891.
(17) Prarz, Ep. “ Ueber einige im Grundwasser lebende Thiere,”
Beitr. z. Kenntn. der unterird. Crustaceen,” Dissert. inaug. Peters-
burg, 1866, 64 pp., 4 Taf.
(18) Rougemont, PH. pE. ‘Etude de la faune des eaux privées de
lumiére.” 4°, avec 5 planch. Paris, 1876.
(19) ScunemeEr, Ros. “ Der unterirdische Gammarus von Clausthal
(Gammarus pulex var. subterraneus),” Sitzber. konigl. preuss,
Akad. Wiss. Berlin, 1885.
(20) Sreppine, T. R. R. ‘Report on the Amphipoda collected by
H.M.S. ‘Challenger’ during the Years 1873-1876.’ Zool. vol. xxix.
London, L888.
(21) Vespovsky, F, “ Ueber einige Siisswasseramphipoden. I. Uebe?
Crangonyx, Sp. Bate,” Sitzber. konigl. bohm. Gesellsch. Wiss:
Prag, 1896, x.
(7)
some Freshwater Amphipods. 243
(22) Vespovsny, F. “Ueber einige Siisswasseramphipoden, IL. Zur
Frage der Augenrudimente von Niphargus,” ibid. 1900.
(23) VismEs Kann, W. F. pg. ‘ Niphargus Kochianus, Bate, in an
Trish Lake, and NV. subterraneus, Leach, in Kent,” Ann. & Mag.
Nat. Hist. ser. 7, vol. xiv. 1904, pp. 274-2883, pl. viii.
(24) Wrzesnrowsxtr, Avaust. “O trzech kielzich podziemnych. De
tribus Crustaceis Amphipod. subterraneis,” Pam, fyziogr. viii.
1888.
“Ueber drei unterird. Gammariden,” Z. f. w. Z. B. 50,
pp. 600-725, 1890, Taf. xxvili-xxxii.
(25)
EXPLANATION OF THE PLATES.
General significance of the letters.
e. Cuticle. n. Secondary branch of the
dl, Ordinary sete. first antennee.
hk. Uyaline spheres in the crystal- p. Pigment-cells.
line cones. r. Retinula-cells.
hp. Hypodermis. sp. Sensory brushes.
kr, Crystalline cones. spo, spo’. Segmental cuticular
sense-organs,
PLATE XI.
Fig. 1. Bathyonyx de Vismesi under low magnification. The appendages
of the left side only are shown.
Fig. 2. Portion of the third joint of the upper (first) antennz, with the
accessory branch (7).
3. Terminal portion of the antenna, with two hyaline clubs [so-
called olfactory clubs] and a sensory brush.
Fig. 4. Mandible, left side, from below.
5, Maxilla of first pair; the inner lobe is not visible in this position,
6. a, inner, b, outer teeth of the middle lobe of maxilla of first
alr.
: Marl of the second pair.
. Maxillipede of the left side.
Figs. 9-11. Gnathopods of the first (a) and second (6) pairs of three indi-
viduals under nearly equal magnification, in order to show the
moderate variability of their form.
Fig. 12. Uropod of the third pair.
Fig. 13. Telson from above, in order to show the position of the sensory
brushes.
Fig. 14. Outer (¢) and fibrous (e’) cuticle, in which the urn of a sensory
brush is embedded. ne, nerve-ending.
Fig. 15. Double hooks of the first and second pleopods.
&
on
Prarn Xl:
Fig. 16. Distribution of the crystalline cones in the pigment and on its
periphery under low magnification. Bathyonyz.
Fig. 17, Head from the left side, with basal joints of the first and second
antennz, the diffuse eye, and the segmental cuticular sense-
organs. Highly magnified.
Fig. 18. ead of the same animal seen from the right side, in order to
show the variable number of crystalline cones.
244 On some Freshwater Amphipods.
Fig. 19. Section through the diffuse eye of a specimen preserved in formol-
The cuticle (c) is widely separated from the hypodermis (Ap).
rt, retinula.
Figs. 20-30. Segmental sensory capsules of different representatives of
the genera Gammarus, Niphargus, and Crangonyx.
Fig. 20. Gammarus sp., from Herzegovina (22 » long).
Fig. 21a, b. Gammarus from Lautenthal (16 p).
Fig. 22. Niphargus elegans from Modena.
Figs. 23, 24. Niphargus puteanus from Prague (12 p).
“1g. 20. Crangonyx compactus.
Fig. 26. Niphargus tatrensis (Switzerland).
Fig. 27. Niphargus from Lille.
Fig. 28. Niphargus Caspary from Munich.
1g. 29. Niphargus Kochianus from Lough Mask ; 4, in optical section.
Fig. 30. Crangonyx subterraneus from Podbaba, near Prague.
[The foregoing paper by Prof. Vejdovsky, of Prague, has
been translated by my friend Mr. D. J. Scourfield, and would
have appeared in print last year, but that it seemed desirable
if possible to supplement the information by the result of
further researches. Unfortunately three days’ dredging
undertaken last summer, and again this year, failed in
securing any additional specimens of Bathyonyx de Vismesi,
although 251 examples of Niphargus Kochianus, Bate, were
taken in Lough Mask. The bright orange colour which
characterizes them all, as well as Bathyonyx, from the lake,
finds a parallel in the observation of M. Chevreux referring
to Niphargus Plateaut, var. robustus, captured in an open
basin formed by the source of the Robine at the foot of the
mountains of Gardiole, the adult specimens of which he
described to be of a salmon-red, paler in the younger indi-
viduals. The probability that these animals might have
been originally introduced from the underground streams that
percolate the limestone strata about Lough Mask suggested
an investigation of these latter. Accordingly, Lord Ardilaun’s
permission having been given, a research was conducted into
the subterranean waters in his extensive grounds near Cong.
This resulted in the discovery of thirteen N. Kochianus in
one cave and a single specimen in another. All fourteen
were of the usual translucent hue which I am accustomed to
see in examples of Niphargus taken from wells, but, with
the exception of their colour, were in every respect identical
with those of the lake. Hence it is now scarcely open to
doubt that the blind species of Niphargus recorded as occurring
in Lough Mask, the Lake of Geneva, the Lake of Zirknitz in
Carniola, and some Swiss lakes, have been derived from their
congeners inhabiting subterranean waters, and that the orange
or salmon-red colour of those in Lough Mask and those cited
On the Trematode Parasites of British Birds. 245
by M. Chevreux is an attribute of the changed environment.
I do not know whether a similar colour characterizes those of
other open waters. As to the origin of Bathyonyx, however,
we have so far no indication of its derivation trom subter-
ranean ancestors; and it appears probable, according to
Prof. Vejdovsky, whose judgment is based upon its general
characteristics and the details of its organization, that its
progenitors lived in the open waters of Lough Mask and that
its present characters are the result of its existence in the
lowest depths. My latest researches have also strengthened
the probability of his suggestion that a habit of burrowing in
mud may have enabled Lathyonyx to escape capture by the
dredge, which skims the surface of the lake-floor. Never-
theless a considerable amount of mud was subjected to exami-
nation on each oceasion, which produced nothing but some
N. Kochianus.
The latter portion of Prof. Vejdovsky’s paper on the
synonymy of
Gammarus Caspary, Pratz,
Gammarus Kochianus, de Rougemont,
Niphargus Kochianus, Vejdovsky,
Niphargus Caspary, Wrzesniowski,
Niphargus Kochianus, Chilton,
has been omitted.— Wm. F. pe VisMES KANE. ]
XXXVIII.—Observations on the Trematode Parasites of
British Birds. By Wi.utam Nicoun, M.A., B.Sc., Gatty
Marine Laboratory, St. Andrews.
Towarps the end of last spring I had an opportunity of
examining several of the commoner species of shore-birds.
These were, unfortunately, in most cases not in the best
condition for helminthological investigation, as I seldom
received them till at least a day after they were killed, when
the parasites were almost always dead. The immediate
application of weak formalin, however, preserved their
anatomical details rather well, although it rendered them
unfit for histological work. For this reason important
features in some cases are only superficially described, a
matter which I hope to improve shortly with the help of
better material. These obscrvations are thus to be regarded,
to a certain extent at least, as merely preliminary.
246 Mr. W. Nicoll on the
The species obtained appeared on a first examination to
be all assignable to well-known genera, e. g. Psilostomum,
Echinostomum, Gymnophallus, Tocotrema, and in particular
Spelotrema. The frequent occurrence of members of the
last-named genus was not surprising, as I had previously
found a Spelotrema species in great abundance in Larus
argentatus. A great variety of sizes occurred, but in this
respect there were no obvious lines of demarcation to
indicate whether one or more species were under considera-
tion. Closer examination proved that not only were there
several different species, but that two distinct genera were
present. They occurred, as a rule, side by side in the same
hosts, occupying the same situations in the intestine and
being equally numerous. I have been able to differentiate
three distinct species of Spelotrema, as well as three species
of the other genus, which is new and for which I propose
the name Maritrema. In addition a new species of Gymno-
phallus and one of Tocotrema have to be recorded.
Altogether 16 species of birds were examined. Of these,
three, Urinator lumme, Alle alle, and Uria troile, yielded no
Trematode parasites. Of the other species 91 per cent. of
the examples were more or less abundantly infected. Of
the total number of all species 68 per cent. were infected.
These figures are very high compared with those of Mtihling*,
Hausmann +, and Wolffhiigel , but this is perhaps to be
accounted for by the fact that I do not include any purely
terrestrial birds (in the sense that they do not frequent
the sea), which are apparently much less subject to infection
than the shore-birds. Other possible explanations are that
this locality is more favourable for infection, or that my
examination has been more exhaustive than that of the
above-mentioned observers.
The absence of Trematode parasites in Alle alle and Uria
troile recalls a similar condition § in Blennius pholis amongst
fishes. Both these birds are, so far as | know, purely
aquatic and feed principally on crustaceans and fish. ‘The
shore-birds, on the other hand, live to a large extent on
mollusks, as well as fish, the diet being thus in great part
different. In the same way the diet of Blennius pholis,
which subsists on small mollusks ||, differs from that of other
* Archiv f. Naturg. lxiv. p. 58.
+ Centralbl. f. Bakter. xxvi. p. 452.
t Beitrag zur Kenntnis der Vogelhelminthen (Inaug.-Diss.), 1900.
§ Ann. & Mag. Nat. Hist. (9) xix. p. 68.
|| Blennius pholis is also fond of Balanus, but the sessile condition
of the latter renders it different from other Crustacea.
Trematode Parasites of British Birds. 247
fish, which feed largely on crustaceans, annelids, and each
other. From these and other considerations it would appear
that, in the littoral zone, at least, where both have a common
feeding-ground, birds derive their Distomid parasites from
mollusks *, while fish obtain theirs from crustaceans,
annelids, and each other. This seems to afford an explana-
tion of the absence of Distomids in the three above-cited
cases.
For ease of reference I subjoin a list of hosts with
their respective parasites :—
Alea torda, Linn. Razorbill.
Tocotrema lingua (Crepl.). Intestine.
? Spelotrema simile, Jdgersk. (juven.). Intestine.
Larus argentatus, Brinn. Herring-Gull.
Gymnophallus deliciosus (Olsson). Gall-bladder.
Maritrema lepidum, sp. n. Intestine.
Parorchis acanthus, Necoll. Bursa Fabricii, rectum,
and intestine.
Spelotrema excellens, sp. n. Intestine and ceeca.
(=Sp. simile, Jégersk., Nicoll.)
Tocotrema lingua (Crepl.). Intestine.
Monostomum sp. (juven.). Intestine.
Larus canus, Linn. Common Gull.
Parorchis acanthus, Nicoll. Rectum.
Larus ridibundus, Linn, Black-headed Gull.
Maritrema gratiosum, sp. n. Intestine.
Phalacrocorax graculus (Meyer). Shag.
? Cryptocotyle concavum (Crepl.). Intestine, ceca, and
rectum,
Ovdemia nigra (Linn.), Scoter.
Gymnophallus dapsilis, sp. n. Bursa Fabricii.
Psilostomum brevicolle (Crepi.). Intestine.
Spelotrema pygmzum (Levins.). Intestine.
Oidemia fusca (Linn.). Velvet Scoter.
Gymunophallus dapsilis, sp. n, Bursa Fabricii (and
P ceca).
Psilostomum brevicolle (Crepl.). Intestine, ceca, and
rectum,
Spelotrema pygmeeum (Levins.). Intestine, czca, and
rectum,
Monostomum sp. Ceca.
Pelidna (Tringa) alpina (Linn.). Dunlin.
Maritrema gratiosum, sp. 0. Intestine.
Spelotrema feriatum, sp. n. Intestine, ceca, and
rectum.
Spelotrema claviforme (Brds.). Intestine, ceca, and
rectum,
Totanus calidris, Bechst. Redshank.
Maritrema humile, sp. n. Intestine.
* An important exception to this is the case of some Spelotrema species,
which are obtained, partly at least, from Cancer pagurus and Carcinus
monas.
248 Mr. W. Nicoll on the
Spelotrema feriatum, sp. n. Intestine, ceca, and
rectum,
Tocotrema jejunum, sp. n. Intestine.
Monostomum petasatum, Deslongch. Ceca.
Numenius arquata, Lath. Curlew.
? Echinostomum secundum, Nicold. Intestine, ceca, and
rectum.
Vanellus vanellus (Linn.). Lapwing.
Spelotrema feriatum, sp. n. (juven.). Intestine, ceca, and
rectum.
Atgialitis hiaticula (Linn.). Ring-Plover.
Maritrema gratiosum, sp. n. Intestine and duo-
denum.
Spelotrema feriatum, sp. n. Rectum, ceca, and in-
testine.
Spelotrema claviforme (Brds.). Intestine, rectum, and
ceca.
Hematopus ostralegus, Linn, Oyster-catcher.
Maritrema gratiosum, sp. n. (juven.). Intestine.
Spelotrema feriatum, sp. n. (juven.). Intestine.
Psilostomum brevicolle (Crepl.). Intestine and czeca.
Genus Srrtorrema, Jiigerskiold.
Spelotrema excellens, sp. 0.
=Sp. simile, Jagersk., Nicoll, Ann. & Mag. Nat. Hist. (7) xvii.
pp. 522-524,
From other members of the genus this species is dis-
tinguished by its comparatively large size. In a former
note I commented on the great difference in size between
my specimens and those of Sp. simile, obtained by Jager-
skidld from the same host. Beyond this I could find no
difference of sufficient importance to warraut specific dis-
tinction, and the occurrence of the two forms in the same
host was a reason for regarding them, if not identical, at
least merely as two varieties of the same species. It not
uncommonly happens, however, that two or even more
species of the same genus inhabit the same host.
Jagerskidld found a similar difficulty in distinguishing
between his form and Sp. pygmeun (Levins.). The two
forms are nearly of the same size and practically the only
differences between them consist in the reversed ratio of the
suckers, which, however, are very nearly equal, and in
the size of the “‘ genital body.” It is to these features that
I, also, am obliged to turn, in order to obtain marks to
differentiate my specimens.
Since first obtaining this form I have met with it several
times again, always in Larus argentatus, never in any other
host. Renewed observations and measurements confirmed
Trematode Parasites of British Birds. 249
those previously noted, except in one respect, the relative
sizes of the suckers. This is by no means an easy matter to
decide, for, the suckers being very nearly equal, slight
contraction on the part of either is sufficient to cause it to
appear smaller. In addition, owing to the method of
preservation, it frequently happens that one or both suckers
are deformed, z. e., elongated in one direction or another,
From previous measurements it was concluded that the
ventral sucker was, if anything, slightly larger than the oral
sucker, thus agreeing with Jagerskidld’s observations. A
further extended series of measurements has caused me to
alter that opinion. In a considerable number of cases I
have found the suckers almost exactly equal, and in all
other instances, except one, the oral sucker exceeded the
ventral to a greater or Jess extent. This is perhaps the most
essential difference between Spelotrema eacellens and Sp.
simile, Jiigersk., apart from the diversity in size. Other
possible features of distinction, such as the size of the
genital body, the length of the ductus ejaculatorius, and
the situation and shape of other organs will be referred
to later.
In most respects my previous description is fairly accurate,
but several features, the importance of which I did not then
realize, were overlooked. In describing the shape, exception
was taken to Jigerskidld’s term “ biscuit-formig.” This
term is occasionally used by continental observers; it is not
very accurate, although it may be apt enough as understood
by them. I preferred the term “ club-shaped,” but that was
in reference more to the outline than to the actual body-
form, in the same sense, I believe, as Brandes used the
specific name claviforme. I can imagine no satisfactory
general term to describe this shape, which is almost peculiar
to the genus. If the outline were made to revolve on its
long axis a club-shape would be evolved; in reality,
however, there is a very considerable amount of dorso-
ventral flattening, particularly in the anterior part of the
body, which is extended, and the sides of which display a
tendency to curl up ventrally. The presence of the large
number of ova causes a bulging in the posterior part of the
body, which, moreover, has an elliptical or almost circular
outline. In describing Sp. pygymeum (Levins.), Odhner *
uses the term “ keulen-formig” and adds that in contracted
conditions the breadth increases gradually from before
backwards. This would involve a somewhat triangular
* Fauna Arctica, iv. (2) pp. 315-316.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 17
250 Mr. W. Nicoll on the
outline, much as Levinsen found it. Such a form is rarely,
if ever, met with in Sp. excellens or Sp. simile. In con-
traction the anterior part of the body becomes broader,
almost as broad as the posterior part, but it is always
separated from the latter by a distinct constriction about the
level of the intestinal bifurcation.
In my second series of measurements, specimens were
found varying in length from ‘71 to 1°39 mm., the average
being °91 mm. ‘The maximum breadth of the anterior part
of the body is °23-—"37 mm.; at the constriction ‘21-28 mm. ;
posteriorly °37—49 mm. The average of these is °29, 24,
‘41 mm., being nearly in the proportion 5:4: 7, and a large
number of examples were found to vary very slightly from
these average figures. Both suckers are small and feebly
developed: the oral sucker measures ‘068-086 mm. in
diameter, the ventral sucker *062—081 mm.; average
‘076 mm. and ‘071 mm. respectively, a ratio of nearly
15:14. The maximum limit ‘095 mm. previously quoted
for the ventral sucker is apparently abnormal. The ventral
sucker is situated at a distance of rather more than 4 of the
body-length from the posterior end, and if it be regarded as
marking the boundary between the anterior and posterior
parts of the body, the ratio between the two is approxi-
mately 5:3.
The pharynx measures ‘037-062 x 026-044 mm., and
the prepharynx is about ? as long. The cesophagus is
narrow and may be any length from -20 mm. to °*45 mm.
The diverticula are about 2 of that length ; they are dilated
and extend back as far as the level of the centre of the
ventral sucker. In a few cases they stop short of, or exceed,
this limit, but the variation in this respect is slight.
Odhner poimted out a distinction between Spelotrema
pygmeum and Sp. simile, which escaped Jagerskidld’s notice.
In the latter species the base of the genital body is much
larger proportionately than in the former. In Sp. excellens
it is still larger in comparison with the size of the ventral
sucker. It measures ‘050--065 mm. in diameter, 7. e., about
4 the diameter of the sucker. It is situated on the left side
of, and close beside, the ventral sucker, which it occasionally
overlaps, and it is on a level with the centre of the sucker
or, as often as not, a little in front of or behind this. The
vesicula seminalis hes in the middle line or a little to the
right, immediately in front of the ventral sucker, sometimes
slightly overlapping it dorsally. Its outline is elliptical, the
long axis being directed obliquely forwards ; size ‘09-11 x
‘07-08 mm. ‘The ductus ejaculatorius, issuing from its
Trematode Parasites of British Birds. yaa) b
anterior end, is not convoluted to the same extent as in
Sp. simile : its course is, in fact, almost direct, albeit slightly
sinuate, towards the genital body. This difference may
be of specific importance. The testes are shaped and
situated exactly as in Sp. pygmeum*, not as represented
in my figure. The outline is oval with the long axis directly
transverse. They are usually enveloped to a large extent by
the uterus. The right testis is just behind the ovary, while
on the left the anterior border of the testis is almost con-
tiguous with the genital sucker. They measure -14—-16 x
‘08-11 mm. The ovary is on the right of the ventral
sucker and is contiguous with it, the right intestinal diver-
ticulum, and sometimes the vesicula seminalis. Its shape is
somewhat variable, being pear-shaped, oval, or more irregular,
but in all cases the long axis is directed obliquely down-
wards to the middle line of the body; size -10-"11 x :07—
‘(OS mm. The uterus starts from the posterior end of the
ovary, and passing down the middle of the body forms a
series of convolutions first on the right side and then on the
left. The convolutions are so voluminous that no regular
arrangement is apparent. The uterus fills practically the
whole of the posterior part of the body not occupied by
the testes and yolk-glands. Its anterior limit is the level of
the anterior border of the ventral sucker. The ova are very
numerous, colourless on the right, bright yellow on the left.
This would appear to be a further point of distinction
between this species and Sp. simile, for, according to Odhner,
the ova in the latter are all colourless, a feature serving to
distinguish it from Sp. pygmeum, in which the condition is
the same as in Sp. excellens. The ova measure ‘023-025 x
‘010-013 mm. The yolk-glands are difficult to make out,
being obscured by the ova. They lie behind and ventral to
the testes, which they overlap to some extent. They appear
to consist of a number of lobes on each side, but the arrange-
ment and the exact number of the lobes I am unable to deter-
mine with certainty. Comparison with Sp. pygmeum and
the next species to be described would point to their number
being about eight on each side. The yolk-ducts are usually
visible, uniting just behind the ventral sucker, the common
duct passing towards the ovary.
Speloitrema feriatum, sp. n.
This species occurred very numerously in Pelidna alpina,
Totanus calidris, and Afgialitis hiaticula, less frequently and
* Odhner’s figure, loc. cit.
L7*
252 Mr. W. Nicoll on the
only in immature condition in Hematopus ostralegus and
Vanellus vanellus. Jn every case the habitat is towards the
lower end of the intestine, the ceca, and the rectum. Occa-
sionally a few were found as far forward as the duodenum.
It is readily distinguished from the other species of the
genus by several well-marked features. The body does not
display ithe familiar club-like outline, it has more that of a
pointed oval, the posterior end being rounded, There is no
marked constriction. The examples from Vanellus are
bluntly oval, presenting the appearance of a species of the
genus Gymnophallus, from which they are certainly hard to
distinguish in their immature condition *.
In point of size it is midway between Sp. simile and
Sp. excellens, the observed limits in length being ‘66-91 mm. ;
average ‘77 mm. In a specimen of average length the
maximum breadth is °36 mm., and is found at the level of
the ventral sucker or a little behind it. The oral sucker is
of large size, measuring ‘09 mm., and is considerably greater
than the ventral sucker, which has a diameter of ‘(075 mm.
The ratio is 6:5. The ventral sucker is almost exactly a third
of the body-length from the posterior end. ‘The pharynx
has an average length of ‘065 mm., and the prepharynx 1s
about as oes but it varies with the state of extension.
The csophagus measures ‘12 mm. in length and the diver-
ticula ‘28 mm. Thus the latter are nearly 24 times as long
as the esophagus, in marked contrast to the condition in
Sp. excellens. Moreover, they extend as far back as the
posterior border of the ventral sucker, and they are not
dilated.
The genital body, situated close to the left side of the
ventral sucker and rather behind the level of its centre, is
comparatively small, measuring only ‘040 mm. in diameter
at the base. The vesicula seminalis is a large ovoid, some-
times almost globular body, lying well to the right side of
* From later observation I am inclined to suspect that more than one
species is under consideration here. What must be regarded as the
typical specimens occur particularly in Pelidna and Aqialitis, and they
present some features which are not met with in the other members of
the genus. These features I have not touched on above, as I at first
considered them to be abnormal; they are as follows:—The body has a
distinct brownish colour, due to the presence of some pigment. The
excretory system is mapped out with surprising distinctness in a series of
intensely black lines, the configuration of which is much as in Jiger-
skidld’s figure (Centralbl. f. Balter. xxvii. p. 794, fig. 2), but, in addition,
numerous fine anastomosing tubules are also visible. Finally, there is a
roundish structure, enclosing apparently two small ovoid bodies, situated
between the genital body and the left intestinal diverticulum, Of the
function of this structure I have at present no knowledge.
Trematode Parasites of British Birds. 253
the middle line. It is thus not entirely in front of the
ventral sucker, with which it is contiguous. Its long axis
is obliquely transverse and the ductus ejaculatorius issues
from its left end and passes round in front of the ventral
sucker in almost direct course to the genital body. The
testes are situated as in the preceding species. The ovary is
a regularly ovoid body, with its long axis transverse and
slightly depressed towards the middle line. It is on the
level of the posterior border of the ventral sucker and is
thus further back than in Sp. eacellens. From it the uterus
passes down the middle of the body towards the left side;
it then runs round the posterior edge of the body to the
right side, where it forms a few convolutions. Pursuing a
transverse zigzag course it returns to the left side, almost
reaching the outer edge of the body, whence it bends back
again and passes forward to the genital sucker. Its topo-
graphy is thus entirely different from that in Sp. excellens.
In addition the ova are almost colourless or of a very dull
hue. They measure ‘020-025 x:010-"012 mm. The uterus
is of very limited extent and does not extend in front of the
testes. The yolk-glands can thus be easily made out. They
occupy the usual position ventral to and a little behind the
testes. In an immature specimen their lobed structure can
be distinctly seen. In Sp. simile Jigerskidld figures six
lobes on each side. Odhner corrects this to eight for both
Sp. simile and Sp. pygmeum. In Sp. feriatum, however,
there seem to be seven lobes on the left side and eight on
the right. Should this be the normal condition it would
be analogous to that in some of the Hemiuride, in which
there are three lobes or tubes on the left and four on the
right. The arrangement of the lobes is characteristic and
apparently fairly constant. In each group there is a central
lobe of roughly quadrilateral outline; the sides of this form,
as it were, facets into which the other lobes fit. The latter
are all of quadrilateral or wedge-shaped outline, broadest at
their distal ends, with rounded corners. The anterior face
of the central lobe and each of the lateral faces accommo-
dates a single lobe, while behind there are three lobes in
the left vitellarium and four in the right. I have observed
the odd number of lobes on the left side in all the specimens
examined, but it is possible that an additional lobe may
be obscured. From Odhner’s figure there appears to be
no such regular formation of the lobes in Sp. pygmeum.
Jaigerskidld * represents a regular arrangement, but there is
no central lobe.
* Centralbl. f. Bakter. xxvii. p. 733, fig. 1.
254 Mr. W. Nicoll on the
Spelotrema claviforme (Brandes).
Bee claviforme, Brandes, Archiv Naturg. liv. p. 247, pl. xvii.
o:
In Pelidna alpina and Afgialitis hiaticula there occurred a
very large number of examples of a specieswhich I am obliged
to identify as Distomum claviforme, Brds., despite its author’s
unsatisfactory description. Brandes’s note on the species 1s
meagre and his figure is erroneous in its most important
feature. It has hitherto not been redescribed. It has been
alluded to, however, by Lithe* and Jagerskidld +, both of
whom were much of opinion that the species must be regarded
as unidentifiable, except by the author himself. In a foot-
note Jiigerskiéld advances the supposition that it might be
included in the genus Levinsenia (=Spelotrema) and gives
an interpretation of Brandes’s figure in this light. I
previously found occasion to agree with this interpretation
and my present discovery shows how correct it is. Ad-
mitting its correctness, however, there remain three or four
errors in Brandes’s figure ; the oral sucker is smaller than the
ventral, the posterior “testes (=the genital body) is much too
large, and the ova are proportionately far too small. The
whole drawing is of a sketchy nature {, so that too much
weight need not be attached to these discrepancies.
The majority of my specimens exhibit the typical club-
like outline in a marked degree. The anterior and posterior
parts of the body are always separated by a constriction
which occurs about the level of the intestinal bifurcation.
The cuticular scales extend exceptionally far back (to the
level of the testes at any rate). The length of the body
never exceeds ‘40 mm. and may be as small as ‘23 mm.
Immature specimens were never observed. The maximum
breadth (posteriorly) is on an average ‘17 mm., while an-
teriorly the breadth is usually about ‘18 mm. As in
Sp. pygmeum, the oral sucker is greater than the ventral,
but the difference is here more pronounced. ‘They have
respectively an average diameter of ‘038 mm. and ‘031 mm.
The ventral sucker is at a distance of slightly less than a
third of the body-length from the posterior one The
pharynx measures ‘020-025 x ‘010-013 mm.; the pre-
pharynx is about the same length as the ahaeane but it
may be considerably longer or shorter. The cesophagus has
* Zool. Anzeig. xxil. p. 537,
+ Centralbl. Bakter. xxvii. p. 759.
{ As witness the representation of the uterus as a continuous tube
having neither beginning nor end,
Trematode Parasites of British Birds. 255
an average length of +12 mm., but it varies very much with
the state of extension or contraction ; the diverticula are
almost constantly ‘08 mm. long. The latter are dilated,
very wide apart, and do not nearly reach the level of the
ventral sucker. In Sp. pygmeum they extend as far back
as the posterior border of the sucker.
The genital body is situated close to the left side of the
ventral sucker, on a level with its anterior half. It is of
small size, the basal diameter being only ‘013-014 mm.
The vesicula seminalis is transversely oval and lies sym-
metrically in the intestinal bifurcation, a considerable
distance in front of the ventral sucker. The ductus ejacula-
torius is slightly sinuate. The ovary is also correspondingly
far forward. It is an irregularly rounded body and is not
contiguous with the ventral sucker, the vesicula seminalis, or
the right intestinal diverticulum, as is the case in Sp. pyge
meum, where it is of much greater size. The uterus
occupies most of the posterior part of the body and extends
forward on each side to the end of the intestinal diverticula.
The ova are fairly numerous and of comparatively large size,
*020—024 x ‘011-014 mm. ‘They are colourless on the
right, bright yellow on the left. The uterus completely
envelops the testes and yolk-glands, so that these organs can
hardly be seen. This is probably responsible for Brandes’s
mistake. The testes have much the same situation, but are
not nearly so large as those in Sp. pygymeum. The yolk-
glands are behind and ventral to the testes, but division into
lobes, if such, indeed, be their condition, is not distinctly
marked.
From the foregoing it is apparent that in point of size,
and possibly on the whole, Sp. claviforme approaches most
nearly to Sp. pygmeum. Several differences have already
been indicated, but they may be briefly recapitulated as
follows: cuticular scales are more extensive ; suckers are more
unequal and the ventral sucker is further back ; surrounding
structures are thus relatively further forward; intestinal
diverticula shorter compared with cesophagus ; genital organs
all smaller; uterus more voluminous, enveloping the testes *.
* A conclusive proof, which I previously neglected to adduce, that
Sp. claviforme is not, as Brandes thought, the adult of the cercaria
described by M‘Intosh from Careinus manas (Quart. J. Micr. Sci.
1865, p. 201) is the fact that this cercaria measures ‘6 x ‘2 mm. (approx.).
It must therefore be the larva of some larger species. A much more
likely cercaria is that described by Miss Lebour (Northumberland Sea
Fish. Rept. 1905, p. 6) from Lzttorina rudis, if it is not an antecedent
stage of the above. It is much smaller in size (‘25 mm.), but it has
rather long intestinal diverticula.
256 Mr. W. Nicoll on the
In addition to the above-mentioned species I have found
species of Spelotrema in Alca torda, Oidemia fusca, and
Oidemia nigra, but their identity is doubtful. From <Alca
only one immature example was taken; it apparently agrees
most closely with Sp. simile. Its length is *53 mm.,
maximum breadth ‘29 mm.; outline club-like; oral sucker
053 mm., ventral sucker ‘050x ‘056 mm.; undivided in-
testine *22 mm., diverticula -17 mm., reaching centre of
ventral sucker ; genital body on level of posterior half of
sucker, not very large. Ovary on same level as sucker,
pear-shaped ; testes transversely oval; yolk-glands almost
clear of the testes, lobed but not well developed.
The species from the scoters is in all probability Sp.
pygmeum, which Jameson ™* has already found in Britain in
the black scoter. I found it in great nnmbers throughout
the intestine of Oidemia fusca, not so numerously m 0.
nigra. The specimens were unfortunately destroyed before
they were thoroughly examined, but from notes made at the
time of collection some idea of their nature is to be gained,
Length up to 50 mm., breadth °24 mm.; oral sucker
‘044 mm., ventral sucker slightly less; pharynx *025 x °022
mm. ; ova °023-"025 x ‘014 mm. These figures agree more
nearly with Sp. pygmeum than with Sp. claviforme, which
can be the only two species in question.
Genus Tocotrema, Looss.
Three species have already been ascribed to this genus,
viz., T. lingua (Crepl.), T. concavum (Crepl.), and T. Muehlingi
(Jagersk.) (= Dist. lingua, Crepl., Mihling). They do not,
however, appear to form a very homogeneous group. Toco-
trema lingua and T. concavum, though agreeing in the
possession of a genital sucker which includes the degenerate
ventral sucker, differ in several important features, which
appear to demand a generic separation. Although it is true
that both possess a genital sucker, the structure of this
organ is not at all similar in both. The peculiar “ kegel-
formiges Koérper ” described by Jiigerskidld + in the sucker
of Tocotrema lingua is not even hinted at by Mihling f{ in
his description of Distomum concavum. It may be concluded
that such a structure is not present in the latter species, for
its presence could not fail to have been noted by Miihling.
My specimens of an almost identical form show no appear-
ance on external examination of such a body, but as I
* Proc. Zool. Soe, 1902, i. p. 158.
+ Bergens Mus. : Aarbog, 1898, 11. p. 10,
{ Arch. f. Naturg. Ixiv. pp. 80-83.
Trematode Purasites of British Birds. 257
possess no sectional preparations this evidence is not con-
clusive. ‘The ventral sucker in Tucotrema concavum appears
to be in a more degenerate condition than in 7. lingua.
Moreover, the voluminous vesicula seminalis of the latter is
very much reduced in the former. A less important differ-
ence is the symmetrical arrangement of the testes in Toco-
trema concavum in contrast to the oblique arrangement in
T. lingua.
The occurrence here of a new species agreeing very closely
with Zocotrema lingua accentuates the difference between
the latter and Distomum concavum. Moreover, considering
the large number of hosts from which Distomum concavum
has been recorded, it is quite possible that two or more
closely similar species may have been confused. The form
which I have obtained from Phalacrocorax graculus is almost
identical with Distomum concavum, but it differs from
Mihling’s description in one or two important features,
which may cause it to be regarded as distinct. Distomum
Muehlingi shows considerable resemblance to Tocotrema lingua,
and if it should prove to have a genital sucker, as opposed to
Miihling’s description, it would be classed in the same group.
For these reasons [am inclined to regard Distomum concavum
as the type of a genus distinct from, but closely related to,
Tocotrema. To this genus Lithe’s name Cry yptocotyle,
hitherto regarded as a synonym of Tocotrema, might con-
veniently be applied. Liihe took Distomum concavum as
type, while the genus Tocotrema is founded on Distomumn
lingua as type.
Tocotrema jejunum, sp. n.
This species occurred fairly frequently in Totanus calidris.
It is of somewhat smaller size than 7. lingua, the length
varying from °87-1'77 mm., but the normal length 1s
probably not beyond 1:3 mm. It displays, however, an
extraordinary amount of extensibility. Almost every ex-
ample was well extended, and in a large number the extension
was so great that they had almost abe appearance of threads,
the body being often bent and twisted in a grotesque fashion,
As might be “expected, the maximum breadth occurring in
the posterior part of the body is not at all constant, varying
from 1 to } of the body-length. The normal figure is
probably about 1, being Sarai in examples of 1:0 mm. to
i-2 mm. long. The shape i is not lke that cf 7. lingua, the
anterior part of the body being more tapering and the
posterior part more rounded. Ina normal specimen (1:0-
1:2 mm.) the oral sucker measures ‘(045 mm. The ventral
258 Mr. W. Nicoll on the
sucker, as in T. lingua, is small and is situated within the
anterior part of the genital sucker. The latter has a trans-
verse diameter of 055 mm. It is placed about the nriddle
of the body, slightly nearer the posterior end. The pharynx
has an almost constant size of ‘038 x:018 mm.; the pre-
pharynx is somewhat shorter. The cesophagus varies from
‘06 mm. in a contracted specimen to ‘14 mm. in an extended
one. The average may be taken as ‘10 mm., or nearly j!; of
the body- length. It is therefore longer than roo lingua.
The intestinal diverticula are much the same as in the latter.
The excretory system is also similar. ‘The cuticle is entirely
covered with scales, with the exception of a small part at the
posterior extremity.
The testes have the same situation as in Tocetrema lingua.
They are irregularly oval, with indentations on their posterior
border, and they are so placed that their long axes are
almost at right angles to each other. The yolk-glands have
also a similar position, but they are not so extensive. As
far forward as the genital sucker their situation is purely
lateral, but at this level they bend in on each side and form
a complete arch in front of the ventral sucker. The anterior
lateral prolongations found in 7. lingua are not present here.
The ovary is a small, roughly oval, lobed body, lying a short
distance in front of the testes on the right side of the middle
line. A receptaculum seminis is present in the middle
line between the ovary and testes. It is of variable size,
being sometimes smaller and sometimes much larger than
the ovary. The uterus lies between the testes and the
genital sucker and laterally does not extend much beyond
the intestinal diverticula. It has the appearance which is
characteristic of Tocotrema lingua, owing to its being
arranged in three fairly regular transverse convolutions.
The ova are numerous and of a light brown colour. They
have a distinctive shape, pointed at one end, much broader
at the other. They differ in this respect from those of
T. lingua, which are more nearly elliptical. The most im-
portant feature distinguishing the two species is the size of
the ova. Jiigerskidld gives the size in 7. lingua as -048 x
‘022 mm., which I am able to confirm m; in 7. jejunum the
size is only ‘031-036 x ‘016-019 mm., the average being
°033 x ‘017 mm.
In this species the vesicula seminalis is even more volu-
minous than in 7: lingua. It extends back as far as the
anterior testis and is highly convoluted. At its anterior end
it becomes distinctly constricted and passes into the pars
prostatica. This is a large pear-shaped body, enclosed by a
Trematode Parasites of British Birds. 259
distinct membrane. It is situated in the long axis of the
body ; broad posteriorly, tapering gradually forwards.
There is some resemblance between this species and
Distomum Muehlingi. They are about the same size and
their ova are equal, but apart from the questionable presence
of a genital sucker in D. Muehling?, it displays the following
differences :—The outline of the body is constricted in the
middle ; the cesophagus is much longer; the yolk-glands do
not extend in front of the ventral sucker, but are more
voluminous posteriorly, uniting between the testes, which
are some distance apart; the ovary is rounded; the vesicula
seminalis is not voluminous; and the ova are few.
Tocotrema lingua (Crepl.).
J have already frequently referred to this species and to
Jigerskidld’s excellent note on it*. It is a very common
parasite of the herring-gull, its favourite habitat being the
anterior end of the intestine, particularly the duodenum.
In addition a new host, Alca torda, must be mentioned.
Only one bird of this species was examined, and it yielded
but a single example of the parasite, so that its occurrence
may be no more than casual. It is a small specimen about
1 mm. long, with few ova, but it is undoubtedly identical with
the specimens from the herring-gull.
The numerous specimens from Larus argentatus show a
remarkable uniformity in size. The length varies from
1-5 mm. to 18 mm., while the maximum breadth is rather
less than 4 of the length, and occurs usually in the anterior
part of the body, giving the outline the familiar tongue-
shape. Occasionally the anterior part is more tapering.
The cuticular scales cover only the anterior part of the body,
but they extend somewhat further back than Jiagerskidld
represents them. ‘The oral sucker has a diameter of -075—
085 mm., and the ventral sucker appears to be nearly of the
same size, although it is difficult to measure the latter owing
to the absence of a limiting membrane. The pharynx
LESTE "06 x °045 mm., and the short prepharynx is about
tas long. The comes is extremely short (04—-06 mm.),
being not more than 5), of the body-length. Jagerskidld’s
statement that it never exceeds } of the body- leneth seems
to be an oversight, for it never in any circumstances
approaches near that length, and in his figure, which is quite
correct, itis not more than 4). It is thus much shorter than
that of T. jejunum or Distomum Muehlingi.
* Bergens Mus. Aarbog, 1898, no. ii,
260 Mr. W. Nicoll on the
Jigerskiéld’s description of the structure of the genital
sucker is accurate enough, but his representation in fig. 1
does not give a true idea of its external appearance. Three
main groups of muscle-fibres are seen: (1) those forming
the genital cup itself, consisting of radial and circular fibres ;
(2) those forming the ventral sucker, consisting almost
entirely of radial fibres; (8) those associated with the
common aperture. These are situated almost entirely in
front of the aperture, very few fibres passing backwards.
Two well-marked circular bands corresponding to Jager-
skidld’s HQM, VQM, fig. 2, pass almost three-quarters
round the aperture. In addition from the sides and anterior
border of the aperture there pass out numerous radial fibres.
Of these the lateral fibres become lost in the surrounding
tissue, but the anterior fibres pass over and round the ventral
sucker and seem to have a certain degree of connection with
it. The ventral sucker is almost separated from the genital
sucker by a constriction partly surrounded by a sort of
sphincter. In extended states of the animal the passage
between the ventral sucker and the genital cup appears as a
narrow tube.
The vesicula seminalis is more voluminous than Jager-
skidld has it. 1t does not reach as far back as the ovary.
At its anterior end there is a distinct constriction where it
passes into the pars prostatica. The latter forms an almost
semicircular (U-shaped) loop directed backwards. The
ascending limb narrows gradually into the ductus ejacu-
latorius, which passes straight forward to the genital sucker,
The ova measure 047-049 x *022-"025 mm.
Genus Cryprocoryie (Liihe).
Liihe gives no definition of this genus, so I offer the
following :—
Body flattened, expanded ; outline oval, slightly pointed
anteriorly, rounded posteriorly, margin usually irregularly
crenated. Cuticle entirely covered with small scales. Ventral
sucker small, situated within the large genital sucker, and
only slightly differentiated from it. Cavity of genital sucker
simple, with no “ plug-shaped body.” Vesicula seminalis
not very voluminous ; pars prostatica? Testes side by side,
symmetrical near hind end of body; posterior border more
or less indented. Receptaculum seminis well developed ;
Laurer’s canal? Ovary irregular, lobed, on right or left side
of middle line. Uterus between testes and genital sucker
forming two or more transverse folds which stretch between
Trematode Parasites of British Birds. 261
the intestinal diverticula (but not beyond them). Ova not
very numerous, light brown, thick-shelled, measuring about
‘036 x:019 mm. Yolk-glands, alimentary system, and ex-
cretory system as in Tocotrema.
Looss’s subfamily Coenogoniminze appears too hetero-
geneous to constitute a natural subfamily. It seems to have
much more the comprehensiveness of a true family, which
might be designated Ccoenogonimide (or Cotylogonimide,
if, as Braun holds, Lihe’s name Cotylogonimus has priority).
Moreover, Looss’s definition of his subfamily does not cover
the forms which he includes within it. For instance, the
following terms do not apply to the genera Tocotrema,
Cryptocotyle, and Scaphanocephalus :—“< Body distinctly
divided into narrow anterior part and plumper posterior
part; genital aperture in front of, or sideways from, the
ventral aac, long axes of testes transverse to long axis
of body ; ovary globular ; windings of uterus extend laterally
to near ae edges of the body ; ova ‘02-03 mm. long.”
That such a family as the Ccenogonimide exists there can
be little doubt, but it is not easy to define its exact limits.
Much depends on which features are considered of greatest
importance. The genus <Ascocotyle appears to indicate a
relationship with the Brachycceliinz, and it is quite possible
that the latter, along with the genera Spelotrema, Levinseniella,
and Gymnophallus, might be included in the same family.
The genera Yocotrema, Cryptocotyle, and Scaphanocephalus,
Jager sk. , represent the furthest stage which the approximation
of “the genital aperture with the ventral sucker has reached.
They form the nucleus of a subfamily, for which the name
Tocotremine, n. subfam., is appropriate,
Cryptocotyle concavum (Crepl.) ?
This form occurred very numerously in Phalacrocorax
graculus; over a hundred examples were taken from one
bird. The favourite habitat is the anterior part of the
intestine and the duodenum. Distomum concavum has been
recorded by several observers from a large number of hosts,
Miihling’s description of the species is the most exhaustive,
and he was the first to discover the true nature of the genital
sucker, but my specimens do not entirely coincide with his
description.
A prepharynx is not entirely absent, but is short, being
only 4 of the length of the pharynx, 'The cesophagus is also
slightly shorter than the pharynx, and thus much shorter
262 Mr. W. Nicoll on the
than Mihling has it. The peculiar inward bend of the
intestinal diverticula as they approach the genital sucker is
a constant feature, as is also their inward inclination at their
termination. The yolk-glands are, as a rule, somewhat more
voluminous than Miihling represents them, and occasionally
form a continuous arch in front of the genital sucker. The
yolk-duets pass along the anterior border of the testes, and
unite in a receptacle of variable size a little to the left of
the middle line. The receptacle is capable of comparatively
enormous dilatation. The fairly large, oval receptaculum
seminis is situated immediately in front of the testes in the
middle line. The testes are much as Mihling represents
them and measure ‘18-26 x:11-"16 mm. With regard to
the situation and shape of the ovary, my observations disagree
entirely with those of Miihling. In my specimens it occurs
on the right side of the body, directly in front of the right
testis. It has the form of a scalene triangle, the longest
side facing forwards ; the shortest side hes along the recep-
taculum seminis, while the third side is parallel to and almost
contiguous with the anterior border of the right testis. The
latter sides are regularly lobed, there being four or five lobes
on the side next the testis and two or three on that next the
receptaculum seminis. The lobes are not of great depth.
The anterior face is more even. The uterus has a character-
istic appearance. It makes first a short turn to the left,
then stretches across the body to the right side, whence it
bends back again to the left side, and turning forward
proceeds to the genital sucker. Its shape is tlierefore G-like,
with two short turns at the ends of the limbs. It never
extends over the intestinal diverticula. In Miihling’s figure
the uterus traverses the body four times. ‘The ova are not
very numerous, light brown in colour, with a thick shell.
They are more pointed at one pole than appears from
Miuhling’s representation. Their size is ‘034-037 x :018-
"019 mm.
Putting aside the possibility of error on Mibling’s part,
which does not seem likely, only two alternatives remain :
we have here to deal with a case of ‘ situs inversus”’ of the
ovary, which is not uncommon in certain species, or my
specimens must represent a distinct variety or species. To
elucidate the matter I examined more than fifty examples
and succeeded in finding one with the ovary on the left side,
the bend of the uterus directed towards the same side and
the yolk-receptacle on the right. Even then it did not
resemble Mihling’s figure, for the ovary retained its charac-
teristic shape, the yolk-receptacle was on the opposite side
Trematode Parasites of British Birds. 263
of the body, and the uterus traversed the body only twice.
The possibility of amphitypy and slight error on Mihling’s
part causes me to hesitate in separating my specimens
specifically from Distomum concavum. I therefore include
them under that species for the present, in the hope that
further investigation will prove them identical or distinct.
My specimens varied in length from ‘68 mm. to 1:06 mm.,
and in breadth ‘40-66 mm., the average size being
°81x'54mm. The maximum breadth is therefore 2 of the
length. The oral sucker has an average diameter of ‘O60 mm.,
and the genital sucker measures *15x‘ll mm. The size of
the ovary is about °13 x ‘07 mm.
Genus GyMNOPHALLUS, Odhner.
Gymnophallus dapsilis, sp. n.
Occurred frequently in the Bursa Fabricii of Oidemia fusca
and Oidemia nigra. It resembles in its habitat and in other
respects Gymn. bursicola, Odhn., from Somateria mollissima.
It differs from all other species of the genus in having the
yolk-glands in front of the ventral sucker, and this is the
principal difference between it and Gymn. bursicola. Mature
specimens varied in length from *84 mm. to 1:13 mm., the
average being very approximately 1 mm., and, in fact, most
of the examples varied but little from that size. One speci-
men *75 mm. long contained ova, but it was an exception.
Immature forms were found as small as ‘68 mm. The
maximum breadth at about the middle of the body was
‘45-80 mm., average ‘63 mm., or nearly 2 of the length.
The general outline is a regular oval with rounded ends.
The cuticle is entirely covered with sharp scales, ‘006 mm.
long; it gives the body a firm consistency and when ruptured
by pressure the fracture is sharp.
The oral sucker is circular, with a diameter of -16—-21 mm.,
i. e., about + of the body-length. The ventral sucker is
smaller, ‘14-17 mm., and lies almost exactly } of the body-
length from the posterior end. In immature specimens the
suckers tend to be more nearly equal. The pharynx is con-
tinuous with the mouth and is almost circular in outline,
with a diameter of ‘068-087 mm. ‘The cesophagus is about
the same length as the pharynx. The diverticula are short
and wide and extend very slightly beyond the middle of the
body. The excretory vesicle resembles that of Gymn. bursi-
cola, but the bifurcation is nearer the ventral sucker. The
limbs end near the edge of the body at the level of the
pharynx,
264 Mr. W. Nicoll on the
The testes are small longitudinally oval bodies, situated
to the outer side of the excretory tubes and on a level with
the posterior border of the ventral sucker or slightly further
forward. They are symmetrical about the middle line and
measure *11x‘08 mm. ‘The ovary is a short distance in
front of the right testis, on the level of the anterior border
of the ventral sucker. It is round and slightly larger than
the testes. The position of the yolk-glands is almost in-
variably in front of and dorsal to the ventral sucker.
Occasionally they are a little further back, but in only one
instance were they found beyond the centre of the sucker.
They are symmetrical about the middle line and consist of a
number of small loosely-knit follicles.
The uterus is confined to the space between the two
suckers. In one case out of sixty a few ova were observed
behind the ventral sucker on the left side, and this was the
case in which the yolk-glands were so far back, thus it must
be regarded as abnormal. The ova are light yellow and
fairly numerous. They present one of the most peculiar
features of the animal, and it is surprising that no mention
has been made of a similar peculiarity in connection with
the other species of the genus. It consists in the presence
in the same specimen of ova of very different sizes, the
largest being as much as eight times the size (volume) of
the smallest. This, moreover, is not a rare occurrence, but
is to be met with in almost every other specimen. In several
instances hardly any two ova were of the same size, and
almost every length between a minimum of ‘012 mm. and a
maximum of ‘03] mm. was observed. The smallest ova dis-
played a fairly constant size of 012 x :009 mm., while what
may be assumed to be the normal ovum (2. e. such as was
found in an example possessing ova of nearly uniform size)
measured ‘025-031 x ‘017-019 mm. This corresponds fairly
well with the size of the ovain G. bursicola. No explanation
of this apparent vagary suggests itself at present ; the pheno-
menon is unique, to my knowledge. It can hardly be the
case that the ova when first formed are of small size and
grow in the course of their passage through the uterus, for
the small ova are found throughout the latter even near its
termination; in addition the shell is of a bright yellow
colour, which is not usually the case in newly-formed ova.
Many large colourless ova are also sometimes to be seen in
the vicinity of the ovary. The condition is much more
probably pathological, but its occurrence in so many speci-
mens seems opposed to this view.
The vesicula seminalis is a large ovoid body on the left of
Trematode Parasites of British Birds. 265
aud slightly overlapping the ventral sucker. It is on the
same level as the ovary. The pars prostatica is also of con-
siderable size ; it lies directly in front of the ventral sucker,
obscured to some extent by the yolk-glands.
From the foregoing it is apparent that this species, while
bearing a close resemblance to G. bursicola, Odhner, has at
least half a dozen constant features of difference. This
species has some claim to be considered as the adult of the
pearl-forming Trematode of Mytilus edulis. Odhner* has a
note on this subject in which he criticizes Jameson’s +
observations. He shows that the specimens which the latter
obtained from Oidemia nigra and identified with Distomum
somaterie, Levins., belong, at least in part, more probably to
Gymn. bursicola. He also corrects Jameson’s obvious error
that such small specimens as he found could be the adults of
the large cercariz in the mussel. He considers Gyman. bursi-
cola as the most probable adult of these cercariz, and there
is no reason why this should not be the case. The claims,
however, of this new species appear equally strong. Its
dimensions allow for a fair increase in size in attaining
maturity, while the immature specimens obtained in Oidemia
fusca are only a little larger than the cercarie in Mytilus.
The difficulty { of proving the identity of the sporocyst stage
in Cardium edule and Tapes pullastra with the cercarive in
Mytilus still remains, and it is quite possible that they may
be distinct. There must be at least six or seven distinct
cercariz of this type to correspond with the number of species
of the genus Gymnophallus.
Genus Marirrema, gen. nov.
Body flattened, leaf-like, more or less elongated oval,
sometimes tongue-shaped. Anterior end usually somewhat
more pointed than posterior end. Cuticle provided with
minute scales over a considerable extent. Suckers small
and nearly equal; the ventral sucker is situated about the
middle of the body. The intestinal diverticula may be short
and sac-like or more elongated and narrow, but in no case
do they extend beyond the testes; the bifurcation takes
place nearer the ventral than the oral sucker. ‘The excretory
system (in Maritrema gratiosum) consists of a small vesicle
from which two narrow tubules run forward; in front of
the intestinal bifurcation they begin to widen and bend in
* Fauna Arctica, iv. (2) p. 312.
+ Proc. Zool. Soc. 1902, i. pp. 151-160.
{ Vide Ann. & Mag. Nat. Hist. (7) xvii. p. 151.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 18
266 Mr. W. Nicoll on the
towards the middle line. Just in front of the pharynx they
unite in the middle line, forming a wide commissure. From
this a short wide branch runs forward on each side. Ovary
situated immediately behind the ventral sucker, median or
somewhat to the right ; it has a characteristic trilobate out-
line, the base facing outwards and the apex median. Uterus
very voluminous, occupying the greater part of the body
behind ovary, but not extending in front of the latter. Ova
numerous, oval, colourless on left side of body, yellow on
right. Size 016-022 x 008-012 mm. Yolk-glands of
limited extent, confined to a fringe along the edge of the
posterior part of the body. At the level ‘of the testes this
fringe bends inwards, passing along the anterior border of
testes to unite in the middle line, so that almost a complete
circle isformed. Receptaculum seminis absent (?). Laurer’s
canal (?). Testes symmetrical about the middle line, a little
behind the ovary ; outline oval, long axis usually transverse,
sometimes decidedly oblique. Vesiculaseminalis large, oval
on the right side of and close to the ventral sucker, behind
which it extends some distance. It is enclosed in a well-
marked cirrus-pouch. This has a somewhat pear-shaped
outline, narrowing in front of the vesicula seminalis. The
narrow part contains a short pars prostatica and passes round
in front of the ventral sucker. The genital aperture is on
the left of the latter, a little behind the level of its anterior
border. The vagina passes up on the left side of the
sucker.
Habitat, the intestine of birds. Type, Maritrema gratio-
sum, sp. n., from Pelidna alpina. Other species, M. lepidum,
sp. n., and M. humile, sp. n.
Maritrema gratiosum, sp. n.
This occurred frequently in large numbers in Pelidna
alpina and Atgialitis hiaticula. A fem immature specimens
from Larus ridibundus and Hematopus ostralegus are pro-
bably best referred to this species, although they do not
wholly agree with the mature examples. The habitat is
most usually the anterior part of the intestine.
The body is flattened and somewhat elongated, with a
tendency to assume a tongue-like outline. There is usually
a slight constriction near the level of the ventral sucker.
The length varies from *45 mm. to 1:10 mm., the maximum
breadth from *24: mm. to*44 mm., or 3-4 of the length. As
in Spelotrema species, considerable difficulty was experienced
in determining the relative sizes of the suckers. Of the
Trematode Pagasites of British Birds. 267
specimens measured about 60 per cent. showed the oral
sucker greater than the ventral, but taking the average of
the figures obtained it was found that both approximated
050 mm. The limits in each case are (043-062 mm. The
suckers are therefore very nearly equal, the oral sucker
being usually a little the larger. In the immature specimens
from Hematopus and Larus ridibundus the oral sucker is
invariably distinctly greater. The oral sucker is subterminal
and the ventral sucker is almost midway between the anterior
and posterior ends of the body. The ‘prepharynx i is usually
of considerable length, -062 mm. on an average, being nearly
twice as long as the pharynx, which measures °033 x 16 mm.
In the Hematopus and Larus specimens the prepharynx was
usually much shorter. The cesophagus measures ‘06-16 mm.,
being 4-1 of the body-length. The diverticula are more
than twice as long, *16—"37 mm., and this forms an important
point of difference between this species and the other two
species of the genus. They always extend as far back as the
anterior border of the testes.
The ovary is situated immediately behind the ventral
sucker, median or a little to the right. The characteristic
trilobate outline is well marked. The uterus is so voluminous
that its convolutions cannot he followed. It first fills the
left side, where the ova are colourless, then passes to the
night, where they are yellow in colour; the condition is
hus the reverse of that in Syelotrema. The numerous ova
yeasure ‘020-022 x:010—012 mm. The yolk-glands have
the situation and extent already noted in the definition of
te genus, and this is constant for all three species. The
testes are not overlapped by the uterus to any great extent,
so that they are probably almost as thick as the body itself.
Their outline is elongated oval, and their long axes are
directed more or less obliquely upwards towards the middle
line. In the specimens from Larus ridibundus the long axes
are directly transverse, while in those from Hematopus the
testes are almost globular. The shape and position of the
cirrus-pouch have been noted in the definition of the genus.
The vesicula seminalis within the pouch had a perfectly oval
outline and is proportionately smaller in this species than in
the other two. It does not extend much further back than
the middle of the ovary. ‘The terminal portion of the vagina
has a sinuate course.
Maritrema lepidum, sp. n
This occurred only in the intestine of Larus argentatus.
It is about the same size as the previous species, but differs
18*
268 Mr. W. Nicoll on the
in the following respects :—The suckers are proportionately
larger and the oral sucker is distinctly greater than the
ventral, their diameters being respectively ‘068 mm. and
‘059 mm. The ventral sucker is situated in front of the
middle of the body. The pharynx is of the same size as that
in M. gratiosum, but the prepharynx is shorter. ‘The ceso-
phagus is about the same length in both; the diverticula are
here distinctly shorter, not extending beyond the level of
the ventral sucker. They are straighter and wider, and are
only 14 times the length of the cesophagus.
The ovary is the same as before, but always shghtly to the
right and somewhat smaller. The condition of the uterus is
also similar; the ova are quite as numerous but smaller,
measuring only ‘018—019 x 009-010 mm. The testes are
further forward, the right testis being almost contiguous
with the ovary and the vesicula seminalis. Their long axes
are always, so far as I have observed, directly transverse,
and they are not overlapped by the uterus to any great
extent. The vesicula seminalis is much larger, extending
back as far as the anterior border of the right testis. The
course of the vagina is distinctly Z-shaped.
Maritrema humile, sp. n.
This was found in large numbers in the intestine of
Totanus calidris. tis a very small species, measuring only
-28—-40 mm. in length, with a maximum breadth of *12 -
‘16mm. The cuticle is almost entirely covered with scales.
Unlike the other two species, it has the oral sucker less than
the ventral, the diameters being respectively 025-031 mm.
and :030—034 mm. The ventral sucker is very nearly in
the middle of the body. The pharynx measures ‘019 x ‘010
mm., and the prepharynx is about the same length. The
cesophagus is ‘04-05 mm. long, or nearly % of the body-
length, while the diverticula are almost twice as long. The
latter are dilated, wide apart, and only reach the level of the
anterior border of the ventral sucker.
The ovary, uterus, and yolk-glands are the same as in the
other two species. The ova are still smaller than those of
M. lepidum, being only ‘016-018 x -008-011 mm. The
testes are almost entirely enveloped by the uterus and their
position is not obvious. They are symmetrical, transversely
oval, and further back than in M. lepidus. The vesicula
seminalis is a comparatively large structure, and the cirrus-
pouch has exceptionally thick walls. It measures *093 x ‘032
mm., and occasionally overlaps the ventral sucker to a con-
siderable extent.
Trematode Parasites of British Birds. 269
In connection with the above described three species it is
interesting to read Jagerskidld’s note, in which he says :—
“ During my search for Levinsenia brachysoma I have found
that there are quite a number of small Distomids, having
the habitat of a Levinsenia, but so different in their copulatory
organs that we are compelled to assign them to other genera.
Thus, for example, in the intestine of Charadrius hiaticula
a small Trematode was found in great numbers, which on
superficial examination might have been mistaken for a
Levinsenia sp., but on closer inspection displayed a cirrus
and a cirrus-pouch” *. I have no doubt that he had before
him members of the genus Maritrema, and in particular
M. gratiosum and M. humile.
Equally interesting is the occurrence in Paludestrina
stagnalis of two cercariz, C. oocysta and C. pirum, first
discovered and described by Miss Lebour f. These show
very close resemblance to species of the genus Maritrema.
C’. oocysta shows well-marked testes, ovary, and cirrus-pouch,
and has thus reached a very advanced stage of development
for a cercaria. In many respects it agrees closely with
Maritrema humile, but the fact that the oral sucker is greater
than the ventral seems opposed to its identification as the
larval form of that species. Cercaria pirum has not attained
such a degree of development, there being no evidence of
genital organs, but the configuration of the alimentary and
xeretory systems leaves little doubt that it is the larva of
pme Maritrema species. The anterior commissure of the
xeretory system differs only from my description of a similar
structure in M. gratiosum in being behind the pharynx in-
stead of in front. The suckers are practically equal, which
agrees with the condition in M. gratiosum, but the intestinal
diverticula are shorter. It is thus impossible at this stage
to identify the cercarize with certainty, but there can be no
doubt that they belong to this or some closely allied genus.
nn Genus Psttostomum, Looss.
Psilostomum brevicolle (Crepl.).
I have found this species not only in the intestine and
cee of Hematopus ostralegus, but frequently also in the
intestine, ceca, and rectum of Oidemia fusca, and in the
intestine of O. nigra. Psilostomum oxyurum (Crepl.) has
* Centralbl. f. Bakter. xxvii. p. 739.
+ Trans. Nat. Hist. Soc. Northumberland &c., new series, vol. 1. pt. 3,
pp. 445-6.
270 On the Trematode Parasites of British Birds.
already been recorded from Oidemia nigra, but this is, I
believe, the first time that P. brevicolle has been met with in the
scoters. The most recent note on the species is by Braun™*,
in the course of which he criticises Mueller’s observations +.
My specimens are small, 1:74-3:41 mm. long with a breadth
of -85-54 mm. They are somewhat cylindrical anteriorly,
more flattened posteriorly, agreeing in this respect with
Braun’s specimens. The posterior end is usually rather
blunt. With regard to the length of the neck (distance
between centres of suckers, as I take it) my specimens are
apparently much more extended than those of Mueller or
Braun, for I find the proportion to be nearly constantly 4 of
the body-length, certainly never less than +, and in young
examples much more. The constriction behind the ventral
sucker is not a mere narrowing in the breadth, but is usually
accompanied by distinct transverse annulation of the cuticle.
The breadth here is only about half that at the widest part
of the body. The oral sucker is slightly larger than the
ventral. ‘The latter is, as Braun says, usually deeper,
although in extension of the anterior part of the body the
oral sucker may be quite as deep. ‘There is practically no
prepharynx. The pharynx is almost globular, measuring
14-21 x 12-22 mm.; it is thus somewhat larger than
Braun has it. He is correct in saying that the cesophagus
is extremely short, the bifurcation taking place almost im-
mediately behind the pharynx.
The testes are contiguous or very nearly so, and th
posterior is most often but not invariably the larger. Wit:
regard to the ova, there seem to be great variations in size.
although when the large size of the ova is taken into con-
sideration the variation is not excessive. The observations
of Mueller and Braun show an apparent discrepancy ; tlie
former gives the dimensions as *12—13 x°l mm., the latter
104x°08 mm. This difference must result from the fact
that Mueller had only measured the larger ova and Bra im
the smaller. With the view to explain this difficul I
measured the ova in upwards of 30 examples, and find), the
limits in length to be *100—124 mm., and in breadth *(32-
‘078 mm. ‘These limits do not represent isolated examples,
for all sizes between them were observed. The average
figures are ‘1118 x -0685 mm., but this does not imply that
such a size is commoner than others. From this it is obvious
that Mueller’s figures, which are at best a rough approxima-
* Zool. Jahrb. Syst. xvi. pp. 12-13, pl. i. fig. 9
+ Arch. Naturg. Ixii. p. 19, pl. iii. fig. 2.
Monaxonellida of the National Antarctic Eupedition. 271
tion, are too high, and that Braun probably was contented
with measuring a very few ova. Both find the breadth too
great. These observations, I may add, were conducted
almost entirely on specimens from Hematopus.
XXXIX.—Preliminary Report on the Monaxonellida of the
National Antarctic Kxpedition. By R. Kirkpatrick.
THE Monaxonellida brought home by the ‘ Discovery’ in-
clude 43 species, of which 24are new. Most of the specimens
came from the neighbourhood of the Winter Quarters (lat.
Ue 29S. long 160) We Be)
Polymastia tnvaginata, sp. n.
Sponge hemispherical, free or attached, covered with a
thick pile of pointed spicules ; with one large oscular papilla
usually completely invaginated, so that the summit of the
oscule is on a level with, or below, the general surface,
Under surface with a fleshy basal pad.
Colour in spirit pale yellow above, and often grey and semi-
transparent on the under surface in free specimens. Con-
sistence dense and firm.
Skeleton.—Choanosomal, formed of fibres curving upwards
from the base to the periphery, penetrating the cortex, and
forming the thick surface pile ; with stellate groups of small
tyles between the fibres.
Cortical skeleton formed of a dense layer of vertical tyles
of various lengths embedded in a tough fibrous layer trom
*d to 1°25 mm. thick.
Basal skeleton consisting of spicules transversely arranged,
and crossing each other in an irregular manner.
Spicules.—Large, smooth, slightly curved styles, or occa-
sionally strongyles, 2240 x 40 pw.
Cortical tyles with small spheroidal head, short neck,
fusiform straight shaft, varying in length from 140 to 350 p,
and in thickness from 12 to 19 w. A few very slender styles
scattered in the choanosome, 70x 6 mw, with head and neck
making an angle with the shaft. Some medium-sized cortical
tyles in the oscular papilla have long, oval heads. ‘Tyles of
the stellate clusters slender, with the head making an angle
with the shaft, 200 x 15 wu.
Localities. Winter Quarters, 10-30 fath.; off Mt. Erebus,
500 tath.
Dee, Mr. R. Kirkpatrick on the Monawvonellida
Spherotylus antarcticus, sp. n.
Sponge dome-shaped or spheroidal, attached or free.
Surface beset with a dense short pile of cortical microtyles ;
with several usually elongated papillee with or without a
large terminal orifice. Dermal pores distributed over the
cortex, each pore opening into a single tubular canal in the
cortex ; the mouth or pore of the pore-canal is guarded with
a ring of radiating cortical tyles. Flagellated chambers
diplodal.
Skeleton formed mainly of radiating fibres composed of
styles, with diverging brushes of spherostyles near the sur-
face. Cortex with a surface-layer of densely packed tufts of
small vertical tyles, and a subcortical layer of tangential
styles and tyles.
Spicules.—Spherostyles 8 mm. in length by 30 p» in
diameter in the middle, and 14 w in the region below the
distal knob; distal knob 28 mw in diameter, hemispherical,
with granular surface and with a few square teeth or serra-
tions on the edge.
Styles straight, fusiform, blunt-pointed, 2°8 mm. in length,
41 w in diameter in the middle, 23 mw in diameter at the
rounded end.
Cortical tyles curved, 146 uw long, head 3°25 wu in diameter ;
neck slender, 2°75 w thick, with broad oar-blade-like shaft,
but circular in section, 7 w thick.
Styles of lower cortical tangential layer, also in choano-
some, 900 x 20 w. ‘Lyles of the same layer nearly straight,
270 w long, with head 7 w in diameter and relatively thick
neck 6°8 w in diameter.
Slender, curved tyles, 460 x 10 w, scattered in choanosome.
Young specimens are oval, with one long closed papilla ;
the bundles of divergent exotyles are more or less separate
and distinct, and the distal knobs retained and not broken
off.
Locality. Winter Quarters, 10-30 fath.
SIGMAXINYSSA, gen. nov.
Cup-shaped Axinellide with longitudinal skeletal fibres
joined by transverse ones on the inner aspect, and with tufts
given off at right angles to these on the outer aspect. Mega-
scleres, oxeas ; microscleres, sigmata and toxa.
Sigmaazinyssa phakellioides, sp. n.
Sponge sessile, cornucopia~ or cup-shaped. Inner surface
of the National Antarctic Expedition. 273
smooth, outer surface coarsely pilose. Consistence rather
hard, but flexible. Colour in spirit greyish drab, Inner
surface with numerous small oscules, each about 1 mm. in
diameter ; outer surface pilose, with dermal membrane per-
forated by round pores 95 mw in diameter.
Skeleton.—On inner surface formed of close-set longitu-
dinal lines joined by cross-bars, and giving off tufts of fibres,
which proceed outwards at right angles to the outer sur-
face, pushing up the dermal membrane, but barely projecting
beyond it.
Spicules.—Oxeas, 835°5 x 42°25 yw, curved at centre, sharp-
pointed. Sigmata, 81:25 w long, 35°75 w broad, and 3:25 yw
thick, often with an angular bend at centre of shaft. ‘Toxa,
130 w long and 3°25 yp thick at centre, with smooth surface.
This species bears in its outward aspect a very close re-
semblance to cup-shaped species of Phakellia ; also the skeletal
arrangement is like that of Phakellia ; the oxeas, sigmata,
and toxa are those of a typical Gellius. The Axinellid genus
Sigmaxinella, Dendy, which has microscleres in the form of
sigmata, has styles for megascleres.
Locality. Coulman I., 100 fath.
Fymedesmia exiqua, sp. n.
The sponge forms a thin translucent greyish-white crust,
about 5 mm. in diameter, on a stone. ‘The surface is smooth,
and the substance of a fleshy consistence.
Skeleton.—The choanogsome contains scattered short acantho-
styles, and the dermal membrane tangential tyles isolated
or in bundles of a few.
Spicules—Megascleres : acanthostyles, 94x19 jy, short,
thick, with spines pointing backwards slightly. Ketosomal
tyles, 157 x 3°5 yu, straight, smooth, with oval heads, 5 w long
and 4°5 w broad.
Microscleres: pluridentate isancore spatuliferce at each end,
with five foliate teeth, 5 ~ in length, sometimes with three
or four; shaft deeply curved, 2°5 » thick, sometimes with
central alate expansions.
Sigmata, 9°6 yw long, 5°6 w broad, *5 wu thick, scattered sepa-
rately in the choanosome.
This new species resembles in several respects H. zetlandica,
Bowerbank, but the ancore of the latter have only three
teeth, the sigmata are much longer (51 «) and in sheaves ;
also the ectosomal tyles are much larger, viz. 328 x 3°25 pw,
and the spines of the acanthostyles more verticillate. Myme-
desmia irritans, Thiele, from Juan Fernandez, has nearly the
974 Mr. R. Kirkpatrick on the Monaxonellida
same spicular elements, but of different dimensions, and has
the labis among its microscleres.
Locality. Off Balleney I., 254 fath.
Hymerrhaphia rufa, sp. n.
The sponge forms a thin mud-coloured brown crust on a
branched Polyzoon. ‘The surtace is smooth, and no pores or
oscules are visible. The consistence is rather tough.
The skeleton of the choanosome is formed of longer and
shorter acanthostyles dressed vertically, that of the ectosome
being formed of tangentially arranged anisotornotes, either
isolated or in bundles.
Spicules.—Megascleres : larger acanthostyles, 312 x 25 m,
swollen at the head, spined all over, with larger curved spines
at the head. Smaller acanthostyles, 131 18°75 wu. Aniso-
tornotes of ectosome, 34412 mw, straight, fusiform, attenu-
ating gradually at one end but abruptly at the other.
Microscleres : isaneore, 28°5 w long, with three or four
teeth at each end; rarely the teeth are not developed, the
ends being in the form of hemispherical cups.
Dredged near Winter Quarters, No. 10 hole, 130 fath.
Ophlituspongia nidificata, sp. n.
Sponge massive, of an inverted pyramidal shape, sessile,
narrow and contracted at the point of attachment. Surface
uniformly coarsely spinous. Circular oscules (‘7 mm. in
diameter) on the upper surface, at the bases of the spines ;
(pores closed). Subdermal cavities flat and shallow. F'lagel-
lated chambers diplodal. Colour dirty grey, the tips of the
spines being yellowish. Consistence hard and tough.
Skeleton consisting of dense branching axes of styles
cemented with spongin and echinated by smooth styles
passing at right angles from the axis to the surface. Hcto~
somal spicules in form of slender straight styles. Consider-
able tracts of dermal membrane were devoid of these spicules.
Spicules—Megascleres: large, straight, smooth styles, on
an average about 1000 x 50 w. Also smooth curved kind,
625 pw long. LEctosomal styles straight, smooth, tapering
gradually to a point, 406 x 9 p.
Microscleres : toxa smooth, 638 yw long, 6°25 w thick at the
centre. ‘These spicules occur in nests or groups of five to
ten.
This new species comes well within the genus Ophlita-
spongia, Bk., as emended by Dendy. O. seriata, Bk., O. sub-
hispida, Carter, and OU. membranacea, Thiele, all have toxa,
of the National Antarctic Expedition. 275
but the first and third species are encrusting, and the second
has long slender branches,
Dredged off Coulman I., 100 fath.
LISSOMYXILLA, Hanitsch.
This genus was established by Hanitsch* to include
Ectyonine Sponges with fibres having a core of smooth
styles echinated by acanthostyles with special ectosomal
megascleres, and with or without microscleres, Unfortunately
the species he selected as type of the genus (Tethea spi-
nosa, Bowerbank) in no way fell in with the definition, since,
as Topsent + points out, this species has neither echinating
spicules nor special ectosomal spicules, and Topsent refers
Lissomyzilla to the limbo of useless names. Among the
‘ Discovery ’ sponges, however, is a specimen which fits in
cls Hanitsch’s definition of ’ Lissomyzeilla, which runs :—
“Skeleton fibres of the choanosome formed of smooth mon-
actinals echinated by spined styles. Megascleres of the
ectosome smooth diactinals or monactinals. Microscleres
(isochelee &c.) may be present.” Accordingly I propose to
revive the name.
Lissomyxilla Hanitschi, sp. n.
There are two much-worn species of this sponge of a dark,
dirty-grey colour, and a third young small specimen, whitish
in colour, growing on a specimen of Hornera. ‘he largest
specimen is 4 cm. high and 5 cm, in diameter at the base ; :
the dermal membrane is worn away, exposing several open-
ings of exhalant canals, 4 mm. in diameter. The dermal
membrane of the smallest specimen is transparent, smooth,
and raised up at one place into a small conical oscule, with
radial tangential spicules in its walls.
The skeleton of the choanosome is formed of branching
fibres on an average about 150 mw thick, echinated by spined
styles in an obscurely verticillate manner, the whole skeleton,
as seen in sections, having a somewhat confused appearance.
The ectosomal spicules partly proceed obliquely from the
main fibres to the dermal membrane, and partly lie tangenti-
ally in that membrane.
Spicules.—Megascleres : styles, 500 x 19 w, smooth, curved
near the head, sharp-pointed. Kchinating acanthostyles,
* Trans. Liverpool Biol. Soe. vol. viii. p. 194 (1894).
+ Archiv, Zool. Exp. et Gén. (3) tome viii. p. 265 (1900),
276 Mr. R. Kirkpatrick on the Monazonellida
219x18:75 jw (without including spines), larger spines
on the head 9 w Jong. Ectosomal amphityles, 356 x 11 4,
straight, very slightly fusiform, subtylote, and with a small
mucro at each end. Microscleres none.
Myzilla victoriana, Dendy (Halichondria pustulosa, Carter) ,
would come under this genus, although, at the same time, it
is in possession of isochele, and the heads of the styli of the
main fibres occasionally have a slight indication of spination.
Localities. Coulman I., 100 fath.; east end of Barrier,
100 fath.
Iophon spatulatus, sp. n.
Sponge slender, cylindrical, branched. Colour pale brown
in spirit. Oscules oval, about 1 x*6 mm. in diameter, slightly
raised ; surrounded by a radiating zone of tyles. Pores in
sieve-like areas 2-3 mm. in length and about 2 mm. in
breadth, on a level with the general surface.
Flagellated chambers 26 x 23 p.
Skeleton.— Dermal: a closely packed layer of dermal tyles.
Choanosomal: a loose network of multispicular fibres
3-6 spicules thick forms the core of the cylinder. From
the central core are given off at right angles fibres one or two
spicules thick which support the dermal membrane.
Sprcules.—Megascleres: smooth styles, 462 x 25 w, with a
mucro at the basal end. Ectosomal subtyles 225 x 12°5 p,
fusiform, with the swelled ends smooth laterally, and with
the extremities only slightly convex, in fact almost truncate,
and covered with spines.
Microscleres: palmate anisochelz 18°7 yw long, 6°2 w broad
(front view).
Bipocilla rare, 13°6 yw long, 1:15 w thick, shaft deeply
curved, slightly twisted, and with ends almost similar, spatu-
late, with crenulated edges ; occasionally with five relatively
large denticles in place of the finer crenulation. A second
kind of bipocilla have a longer, less curved shaft, with scoop-
like rather than spathulate ends, each scoop having four or
tive denticles, One example has three sharp prongs at one
end and the spathulate prolongation at the other.
The species is represented by numerous small, for the most
part fragmentary, slender cylindrical pieces.
Since writing the above description accounts of two new
species of Jophon from the Antarctic have been published by
Topsent, viz. I. unicornis and I. pluricornis. ‘The two new
species described in the present Report and Topsent’s two
species all possess the curious modified acanthostyles.
I. unicornis, 'Vopsent, has ne bipocilla, and Z. pluricornis has
of the National Antarctic Expedition. Dita
bipocilla apparently of the typical form; the present two
species both have spathulate bipocilla with crenulated ends ;
and, further, there are differences, viz. in the shape of the
body, the arrangement of the skeleton, and the character of
the ectosomal spicules, which lead me to regard the
‘ Discovery’ specimens as belonging to distinct species.
Localities. Winter Quarters, 25-30 fath.; Coulman L.,
100 fath.
Tophon flabello-digitatus, sp. n.
Sponge forming a large palmato-digitate or digitate growth
in one plane; branches compressed, usually with oscules
along one edge. Surface finely verruculate ; with elongated
pore-areas supported by fan-like wisps of ectosomal spicules.
Colour dark brown; consistence soft, the sponge being
readily broken.
Skeleton typically formed of a network of spiculo-fibre, the
primary lines of which proceed upwards and outwards from
the inner surface of oscular tubes to the outer surface of the
sponge, where their ends form the verrucee ; the secondary
fibres join the primary at right angles, forming rectangular
meshes about 1°25mm. square. The thickness of the primary
fibres is about *7 mm., that of the secondary about ‘5 mm.
The ectosomal skeleton consists of fan-like bundles and
wisps of spicules, isolated or proceeding upwards and spreading
out from the terminal main fibres.
Spicules.—Megascleres: smooth styles, 590 x 25 yu, curved,
with a mucro at the head end.
Ectosomal spicules, 344 x 12°5 w, subamphitylote, fusiform,
with a marginal ring of vertical spines at both ends and a
terminal central spike at one extremity.
Microscleres: palmate anisochele of two sizes, a large
kind 35 w long and 10°7 w broad, with thick shaft, with
triangular palmate tooth, 17°5 w long, at the large end, not
quite as long nor as broad as the ale; lower margin of alw
convex. Lower central tooth with a curved upper edge
produced into a spine.
A small kind, 17:5 uw long, 6°2 w broad; upper palmate
tooth triangular, rounded above, as broad and as long as the
alee ; lower border of alee concave ; lower tooth with simple
rounded upper edge.
Bipocilla varying in length from 5°5 to 11 yw, according to
the convexity of the shaft, which is usually deeply curved ;
both ends spathulate, nearly similar, and with crenulate
edges, or with 5-7 teeth.
278 Mr. R. Kirkpatrick on the Monaxonellida
The fine specimen (no. 184) which constitutes the type of
the new species is 24 em. wide and 25:6 cm. high.
Locality. Winter Quarters, 28-130 fath.
Mywilla decepta, sp. n.
There are two very small specimens of this species: one is
in the form of an extremely thin incrustation on a piece of
rock ; the surface is pilose, owing to the projection of verti-
cally dressed acanthostyles, each surrounded by tufts of
ectosomal spicules. The other, which incrusts the branch of
a Polyzoon, is thicker, and the surface here is partly smooth,
partly provided with minute sharp-pointed conules supported
by acanthostyles. The colour of both specimens is reddish
brown.
The skeleton in the very thin incrustation at first sight
resembles that of a Hymerrhaphia; each vertical acanthostyle
is isolated and with its head on the base and its pointed end
projecting. In the thicker specimen it is possible to make
out primary and secondary lines of skeletal fibres.
The ectosomal spicules are partly arranged in paniculate
tufts, partly lying tangentially in the dermal membrane.
Spicules.—Megascleres: choanosomal acanthostyles, 468 x
23°5 mw, curved, spined at the head only, with subtornote
points. Ectosomal strongyles, 238 x 4°6 w, straight, smooth,
cylindrical, usually with a pointed mucro at one end.
Microscleres: arcuate isochele, 19°5 w long, 5°6 w broad,
palmate tooth 8 w long, ovoid, with rounded distal edge; with
thick curved shaft; with tongue-shaped ale about 8 w long.
Isancore unguiferze, 15°3 yu long, with slender, curved,
sometimes wavy shaft, with three sharp claw-like teeth at
each end.
Chelate bipocilla 8 long, with deeply curved shaft with
spathulate ends each with three triangular denticles; these
spicules are fairly common and not accidental.
There are also several isochelz arcuate in which the ale
and denticle are replaced at one end by a spoon-like lamella.
The presence in Myailla of chelate bipocilla similar in
many respects to those found in the new species Jophon spatu-
latus and I. flabello-digitatus is exceptional. In other respects
the spiculation is that of a typical J/yxilla. The isochele
arcuate, though only half the length, resemble in shape
those of Myailla nobilis, R. & D., from off the Rio de la
Plata, and J/. digitata, R. & D., from the Cape of Good
Hope. ‘The new species, again, closely resembles Mywilla
dophonoides, Swartszewsky (Mém. Soc. Nat. Kieff, xx. p. 340,
of the National Antarctic Expedition. 279
pl. xi. fig. 7, and pl. xv. fig. 27), from the White Sea; but,
m addition to other differences, the latter species has no
asochele arcuate.
Localities. Winter Quarters, 125 fath.; off Balleney L.,
254 fath.
Tedania variolosa, sp, n.
Sponge in form of a mass of thick flabellate or digitate
fronds arising from a common base; with circular sphinctrate
oscules, each about 1 cm, in diameter, situated at the summits
or along the upper edges of the branches, the canals into
which they lead extending nearly to the base of the branches.
General surface of the sponge covered with circular pore-
areas each about 4 mm. in diameter, the oval or circular pores
being about 90 » in diameter, and the strands of the poral
reticulum about 30 yw in breadth. Colour in spirit pale brown.
Consistence soft and fleshy, being easily torn.
Flagellated chambers, 42x35 yw, oval, aphodal, with
aphodus (in a measured example) 13 wu long.
Skeleton.—Choanosomal skeleton formed of loosely agglo-
merated compound, longitudinal, or main bundles about
1 mm. in diameter, curving out to the surface as they pass
upwards ; the separate fibres of the main bundles about 80 wu
thick. ‘The main bundles joined at right angles by secondary
fibres 1-3 spicules thick. Spongin not perceptible. Kcto-
somal skeleton formed of circles of strongyles, the spicules
isolated or in fan-like wisps, arranged partly vertically,
partly tangentially, round the pore-areas ; the vertical spicules
usually isolated and the tangential ones in wisps. On drying
the sponge the edges of the pore-areas stand up sharply, the
areas themselves sinking in, giving a pock-marked aspect to
the surface.
Spicules.—Megaseleres: choanosomal styles, 402 x13 yp,
curved at about one fourth of the length from the round end,
smooth, but occasionally with a few spines about the head.
Ectosomal strongyles, 261x6°5 mw, smooth, occasionally
slightly swollen at each end.
Microscleres none.
The single specimen is in the form of a squarish mass of
thick fleshy flabello-palmate or digitate lobes; the height is
18 cm. and the breadth 13 cm.
The arrangement of the pores in circular areas each
sutrounded by a zone of ectosomal spicules is not common in
Tedania ; it occurs in the second new species described below,
and something of the kind is found in Tedania tenuicapittata,
Ridley, from the Straits of Magellan. In the present species
250 Mr, R. Kirkpatrick on the Monaxonellida
this feature is so well marked as to give the surface a pock-
marked appearance.
The rhaphides, usually so characteristic of Tedania, have
entirely disappeared.
Locality. Winter Quarters, 10 fath.
Tedania coulmani, sp. n.
The single specimen isin the form of a finger-like fragment
5°5 cm. long and 1:7 cm. in its greatest thickness. he
colour is dirty grey and the consistence soft. The surface
shows the same circular pore-sieve areas as in 1’. variolosa.
Along one side of the sponge the surface has been torn away,
exposing an exhalant canal running along the length of the.
specimen, but apparently the terminal oscule has been torn
away.
Skeleton.—Rings of spicules, partly vertical, partly tan-
gential, isolated or in tufts, surround the pore-areas.
The choanosomal skeleton is formed of primary longitudinal
fibres about 120 w thick, joined by secondary fibres one
spicule in length and two to three in thickness, joining the
former at right angles.
Spicules.—Megascleres : the choanosomal acanthostyles,
475 x 18 w, curved, smooth, or with sparse spines, usually on
the upper and lower thirds of the length.
Dermal ectosoma] tornotes, 319 x 12°5 #, smooth, straight,
fusiform, larger at one end than the other. Under a high
power each end shows a rounded shoulder prolonged into a
mucronate spine.
Microscleres absent.
The present species resembles 7’. vardolosa in having the
circular pore-areas, and in the absence of rhaphides, but
differs widely in the character of the dermal tornotes. Both
species differ from all other species of Tedania in having no
microscleres. The nearest species to the present one are
Tedania tenuicapitata, Ridley, from the Straits of Magellan,
and Trachytedania spinata, Kidley, trom the same locality ; :
both of these have rhaphides, and neither has the circular
pore-areas, though in 7’. tenwicapitata there is a tendency to
a radial arrangement of bundles of dermal spicules. The
spination of the acanthostyles recalls a similar character in
Trachytedania spinata.
Locality. Coulman I., 100 fath.
Mycale acerata, sp. n.
Sponge large, massive, with numerous small rounded
of the National Antarctic Expedition. 281
mammille; surface finely reticulate and finely hispid.
Colour creamy white in spirit. Consistence soft, the tissues
being easily torn. The flesh reddish (but soon decolorized),
and showing the glistening white strands of the skeleton.
Oscules in form of wide, thin-walled, cylindrical chimneys
with rather jagged upper edges, about 1 cm. in height and
1-2 cm. in diameter.
Skeleton.—Ectosomal: a network of triangular meshes
formed by bundles of oxeas, the strands being about *35 mm.
thick and the meshes about °5 mm. across. Main skeleton
formed of long thick anastomosing fibres, which attenuate
gradually from 1°5 mm. in thickness and break up a little
below the surface into panicles of much finer fibres, which
support the dermal membrane and penetrate the strands and
nodes of the dermal reticulum, giving rise to a finely hispid
condition of the surface. Parallel groups of oxeas scattered
in the choanosome.
Spicules—Megascleres: oxeas, 850X16 25 yp, slightly
curved, rather abruptly pointed at one end and more tapering
at the other. These oxeas form the fibres and also are
gathered into bundles, one spicule in length, of parallel oxeas,
_ scattered in the choanosome,
Microscleres: large anisochele palmate, 105 x 50 yw, sepa-
rate or in rosettes, usually with an angular bend in the
shaft; with a triangular upper tooth 60 w& long, about the
same length as the upper ale, which latter are very wide.
With the lower tooth oblong, 12°8 w high, with a slightly
convex edge; in one of the specimens this edge is produced
into a denticle.
A smaller kind of anisochele palmate, 47 « long and 17 pw
broad, at the upper end, with a long oval tooth 20 w long
extending below the ale.
Trichodragmata, 62x12 w, the trichites being very fine,
sharply pointed oxeas.
There are three fine specimens of this species, the largest
forming a thick flabellate body 17 cm. high, 11 em. broad,
and 7 em. thick.
The mammillz are on an average about *75 em. in height,
and 1 cm. in diameter at the base. ‘The new species bears a
very close resemblance to Mycale magellanica, Ridley, which
likewise has a mammillated, finely reticulate surface and
glistening skeletal fibres, but here the surface is smooth and
not hispid, and the megascleres are styles, or subtyles, such
as are normally found in the genus Mycale. The microscleres
also are different in the two species.
A second species of Mycale with oxeate megascleres 1
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 19
282 Mr. R. Kirkpatrick on the Monaxonellida
Mycaleintermedia (O. Sch.), from Kast Greenland, noticed by
Thiele. ‘he Arctic specimen consisted only of a fragment ;
but the spicules, which are all considerably smaller than in
the Antarctic species, have the following dimensions :—Oxeas
450 p long, 10-12 pw thick; large anisocheles 50-60 yw long ;
small anisocheles 18 » long.
Locality. Winter Quarters, 25-178 fath.
Desmacidon meandrina, sp. n.
The material consists of three subcylindrical fragments
tapering at the distal end.
The consistence is hard and dense. The colour in spirit
is dirty brownish grey.
The surface is fairly uniformly level, and presents flattened
papillze or meandrine ridges, slightly roughened at the top by
projecting oxeas (best seen on side view with a lens).
The dermal membrane roofs over the grooves and spaces
between the papilla and ridges. The pores are mostly
circular and about 95 w in diameter.
The small circular oscules, numerous and scattered, are
about 1 mm. in diameter. The skeleton is formed of a thick,
main axis, consisting of rather loose longitudinal strands ;
from this are given off at right angles cylindrical or lamellar
bundles of loose strands, which proceed to the surface and
form the papille and ridges.
Spicules.—Megascleres: oxeas, 579 X 39 w, curved (usually)
or bent at the centre, with sharp, pointedends. Microscleres :
isancore unguifere, 26 w long and 15:8 w broad; shaft
strongly curved and 38°52 w thick; with usually five teeth or
claws, about 5°28 long, at each end, viz. a central, single,
and two lateral bifurcated teeth.
The largest of the three pieces of this sponge is 6°2 cm.
long and 16 mm. in diameter. The fragments appear to be
broken off from some branched specimen, and I shall refer to
them as branches. They are subcylindrical, being slightly
compressed in one plane. The chamber system is aphodal, the
flagellated chambers (44 x 29 w) being pyriform. ‘There is a
considerable amount of variation in the teeth of the isancore,
the number varying from three to six, the most usual number
being five.
In some respects the new species resembles Desmacidon (?)
ramosa (R. & D.), obtained by the ‘ Challenger’ from the
Cape of Good Hope and Marion Island. In both species
there is a central axis of longitudinal fibres, whence fibres
proceed to the surface at right angles. In the ‘ Challenger’
of the National Antarctic Expedition. 283
species the radiate bundles branch in a fan-like manner,
finally forming an almost uniform surface layer of vertical
oxeas. The microscleres in D. (?) ramosa are isochele
arcuate,
Locality. Coulman I., 100 fath.
Desmacidon spinigera, sp. n.
Sponge digitiform or knob-like ; surface coarsely spinous.
Consistence rather hard. Colour pale red. Several small
oscules about 2 mm. in diameter.
Dermal membrane spread like a delicate net between the
spines, and at some distance from the floors of the subdermal
spaces. Flagellated chambers oval, 46 x 32 yp.
Skeleton formed of coarse, longitudinal main strands,
about 180 thick, radiating out to the surface, with loose
scattered spicules between, united in horizontal bundles only
beneath the surface.
Spicules.—Megascleres: oxeas, 731 x 26, curved at centre,
mostly subtornote, though some attenuate gradually, with
sharp pointed ends.
Microscleres of one kind, viz. isochele palmate, 24°64 w
long and 5:28 w broad, on side view; pointed at each end,
with straight axis; with palmate teeth 8°8 w long, and with
narrow ale 8°8 w long.
Four specimens were obtained. The type specimen from
No. 10 hole, 130 fath., is digitate, 7°5 cm. in length and 2 cm.
in diameter. The spines are *2-"5 cm. long, those at the
lower end pointing obliquely upwards, but above becoming
vertical to the long axis.
This species bears much resemblance to Desmacidon setifer,
Topsent, obtained by the ‘ Belgica’ from the Atlantic. The
isochele are o£ much the same character, but those of
D, setifer are very much larger, viz. 75-100 w by 18-20 pw.
Further, the consistence of the latter is soft, the colour
yellowish in spirit, and the surface hispidation much finer.
Localities. Winter Quarters, 20-130 fath.; Coulman L.,
100 fath.
Joyeuxia Belli *, sp. n.
Sponge attached, ovoid, with a thick rind enclosing a soft
pulp; with short conical osular, and long trumpet-shaped
poral papille. Surface finely pilose. Colour of surface
yellow, of the rind whitish, and of the pith deep yellow.
* Named in honour of Emeritus Professor F. J. Bell, of the Zoological
Department of the Natural History Museum, and editor of the “ Reports
on the Natural History Collections ” brought home by the ‘ Discovery’
from the Antarctic.
ge
284 Mr. R. Kirkpatrick on the Monawxonellida
Flagellated chambers 23 x 20 4; diplodal.
Skeleton.—Cortical skeleton formed of layers of strongyles
crossing each other at right angles. The walls of the oscular
and poral papillae supported by a layer of longitudinal
strongyles. The surface of the sponge hirsute, with a fine
pile of strongyles standing out at right angles or obliquely.
Choanosome without spicules.
Spicules.—Slightly flexuous smooth strongyles 850 long,
10 in diameter at the ends, and 13m in diameter at the
centre.
There is one adult specimen 5 cm. long, 3°5 em. broad,
and 3 cm. thick, with a deep groove on the under aspect, by
which it was probably attached to a worm-tube or stem of a
hydroid. There is also a small conical specimen, 6 mm. high,
attached to a piece of rock.
I was at first disposed to regard this remarkable species as
a member of a new genus, partly on account of its very thick
rind, which is in places over a millimetre in thickness, and
partly because of the highly specialized poral papille ; but
apart from these characters, the new form evidently shows the
closest affinities to Joyeuxia. The three hitherto described
species all have a rind enclosing a soft pulp, the latter being
without or almost without a skeleton; then, too, the pulp is
highly coloured. Joyeuxia tubulosa, Vopsent, and J. ascidoides
(Fristedt) have fistule, which, however, appear to be oscular.
Two of the species, J. viridis and J. tubulosa, have strongyles ;
J. ascidioides has tyles and also chele. Accordingly Topsent
places the genus near Desmacidon.
The poral papille attain a height of 1 to 1:2 cm.; they
are expanded at the end.
The inconspicuous oscular papillee are only about 4 mm.
high and are tightly contracted.
Locality. Winter Quarters, 10-20 fath.
CERCIDOCHELA *, gen. nov.
Mycaline with peculiar shuttle-shaped chele or canono-
chele }, with the single tooth from each end fused, and with
a semicircular vertical lamella extending inwards from the
shaft and from the dental bridge, so as to nearly meet.
Cercidochela Lankesteri ft, sp. n.
Sponge elongated, slender, fusiform. Colour white ; con-
sistence soft. Surface smooth to the naked eye, but finely
* xepkis, ldos, a shuttle. + xavay, dvos, a shuttle.
+t Named in honour of Professor Sir E. Ray Lankester, K.C.B., F.R.S.
of the National Antarctic Expedition. 285
hispid under a lens. With several small scattered oscules,
about 1 mm. in diameter, level with the surface. F'lagellated
chambers aphodal, oval, 31 x 21 pw.
Skeleton formed of long longitudinal lines of spicule-fibres
about 100 thick, not forming a definite central axis,
radiating out in plumose manner to the surface ; with a few
isolated spicules arranged in a scalariform manner at right
angles to main fibres. Spongin not perceptible.
Spicu/es.—Megascleres: oxeas, 452x19°5 yw, curved at
centre, attenuating gradually to sharp points slightly planed
away on inneraspect. Microscleres : canonochelez, somewhat
shuttle-shaped, 45°5 w long by 22°75 broad, with the two
teeth fused to form a bridge, and with a semicircular lamella
passing upwards from the shaft and downwards from the
dental bridge, both lamellz being nearly on the same plane
and nearly meeting, the lamelle sometimes with basal
tubercles. Developmental forms in shape of thin, oval,
linear bodies, the oval at first not being complete.
The unique specimen representing the new genus and
species is 12°5 em. long and 1 em. in breadth at the centre.
The body attenuates to fine ends, and apparently has not
been attached to anything.
The remarkable canonochele recall to mind the spheran-
core of Melonanchora, but the latter spicules have three pairs
of fused teeth.
The canonochele are scattered about in the choanosome in
considerable numbers. ‘The shape may be compared with
an oval basin with the bottom cut out, and with two semi-
circular lids or lamella passing horizontally from the upper
edge of the basin, so as to nearly meet; further, it is
necessary to imagine such a basin turned up on its side,
The earliest developmental forms have an _ elongated
C-shape; then the open G@ becomes a closed oval; by this
time the falx at each end is perceptible, and the beginnings
of the lamelle appear. A further change leads to a marked
asymimetry, the thin oval ring becoming a broad band by
widening in a direcilon away from the edges whence the
lamella arise.
Locality. Winter Quarters, 130 fath.
HOPLAKITHARA *, gen. nov.
Mycalinz possessing exotyles with large spherical spined
heads and with fimbriated placocheles.
* onda (pl. of émAos), armour; x:Odpa, a guitar,
286 Mr. R. Kirkpatrick on the Monawonellida
Lloplakithara Dendyi*.
Sponge in form of a small cushion, attached by a narrow
base. Surface smooth to the naked eye. Colour pale brown
in spirit. Consistence, hard externally, soft within. Flagel-
lated chambers 32°5 w in diameter, spheroidal, eurypylous.
Skeleton with protective armour formed by gigantic
spheroidal heads of exotyles, the exotyles being arranged as
radiating bundles in form of inverted cones, with the apices
a little below the cortex; with scattered strongyles.
Spicules.—Megascleres: exotyles, with the heads a little
inclined to the long axis of the spicule, the proximal end (in
the interior of the sponge) rounded, the distal end swollen
into large spherical heads, with short cylindrical spines
covering the distal three-fourths of the head. ‘Total length
358 yw, the shaft enlarging in diameter from 6°5 w at the
proximal end to 16 mw just below the head. Head 55 in
diameter; cylindrical denticles 1°76 to 3°52 in height,
with finely denticulate edge, and with cup-like depression at
the summit.
Strongyles straight, fusiform, smooth, 467°5 wu long, 9°75 pw
in diameter at centre, 6°5 w in diameter at ends.
Microscleres : placocheles, fimbriated, 84°5 w long, 29°25 wu
broad ; length of tooth 87°75 p.
Sigmata very small, slender, C-shaped, 8°8 wu long, 5°28 wu
broad, ‘9 w thick.
The minute spheroidal or cushion-shaped specimen was
2°2 mm. in height and 3 mm. in horizontal diameter; it was
growing on the side of an Alcyonarian creeping over a
branched Cellepora. No pores or oscules were discernible.
The under surface, which was narrowed to the point of
attachment, was paler in colour than the upper.
The new genus is closely related to the Mycaline genera
Rhaphidotheca and Guitarra, to the former by its exotyles,
and to the latter by its fimbriated placocheles.
The distal knobs of the exotyles of &. Marshall-Hallit,
Kent, 49 in diameter, are smooth and spherical, and those
of LK. rhopalophora, Schmidt (R. affinis, Carter), are 104 pu
long and 80m broad, and club-shaped. Lundbeck regards
these two species as probably identical, and certainly the
differences are slight.
Locality. Winter Quarters, 130 fath.
Gellius fimbriatus, sp. n.
Sponge in form of a thick triangular cake, or conico-
cylindrical. ‘Texture soft, easily broken. Colour in spirit
pale buff.
* Named in honour of Professor Dr. A. Dendy.
of the National Antarctic Expedition. 287
Surface level, or almost imperceptibly hispid ; showing
through the dermal membrane a somewhat areolated pattern,
each areola being formed by the end of a main fibre giving
off fimbriated twigs which roof over the subdermal spaces
between the main fibres. A few circular oscules about
5 mm. in diameter, and with slightly raised rims, occur.
Skeleton formed of longitudinal lines of flat, loose, band-
like main fibres, with an irregular and obscure reticulation of
single spicules between. ‘he main fibres spread out in a
paniculate manner a little below the dermal membrane.
Spicules.— Megascleres : oxeas 537 x 16°25 y, slightly bent
or curved at centre, attenuating gradually to sharp points.
Sigmata varying in size, the largest being +40 m long,
17°6 w broad, and 1°76 thick, with one or more angular
bends in the curve, usually one end of the shaft with an
angular bend, the other end curved.
The specimen selected as the type of this species has a
flattened angular pad-like body, 10 em. long, 4°5 em. broad,
and 2°5 cm. thick; it apparently lay free on the bottom.
An “areola” with its central node (the end of the main
fibre) and lateral branchlets occupies on an average an area
of 6x4 mm.
‘The upper surface alone shows the areolated appearance
below the dermal membrane, the under surface being nearly
opaque.
‘The triangular cushion-like shape of the type calls to
mind Gellius flayellifer (Ik. & D.), but there are no flagellate
sigmata in the new species. G. rudis (Topsent) has a much
firmer and denser structure, the oxeas are shorter, thicker,
and with tornote ends, and the sigmata are more slender and
with uniform curve.
The ends of the main fibres are arranged in linear series.
‘The dermal membrane is separated about 3 mm. from the
floors of the large subdermal spaces, and the band-like
supporting pillars are about 2 mm. broad. A vertical section
gives the appearance of a miniature ‘‘hall of a thousand
columns.” ‘lhe dermal membrane on the under surface of
the sponge contains scattered oxeas arranged tangentially.
Localities. Winter Quarters, 12-20 fath. ; Coulman L.,
100 fath.
Gellius pilosus, sp. n.
Sponge in form of an erect flattened triangular or elongate
lamella divided or digitate at the upper edge. Consistence
soft, fleshy, compressible.
Colour in spirit dirty white or very pale yellow.
288 Mr. R. Kirkpatrick on the Monawonellida
Surface finely conulose and pilose, owing to the projection
of the ends of the main skeleton-fibres about 1 min., the
conules being about ‘6 mm. apart from each other.
A few oscules, about 1 mm. in diameter, on a level with
the surface.
Skeleton formed of slender main axial fibres on an average
about 2-5 spicules thick, curving out to the surface, where
they form the pile, and of secondary fibres, usually one, but
sometimes two or three spicules thick, at right angles to the
main ones, with which they form oblong scalariform meshes.
Spongin well developed at the nodes of ‘the network.
Spicules. — Megascleres: oxeas, 537 x 22°75 mw, sharp-
pointed, subtornote, a few being distinctly tornote.
Microscleres: sigmata, very “abundant both in choanosome
and ectosome, C-shaped, 39 x 16°25 w in length and_ breadth
and 1°5 w thick.
There are two specimens. One of them is in the form of
a triangular lamella dividing into two subterete branches, the
total length being 10°5 cm., the breadth 3°5 em., and the
thickness 1 em. ‘The second specimen is 11 cm. long and
3 cm. broad, with little more than a notch at the upper edge,
indicating a division into branches.
‘The new species bears some resemblance to Gellius flagel-
lifer, R. & D., but differs from it in the absence of the
peculiar flagellate sigmata. Further, G. flagellifer has an
even surface, and a dermal skeleton network of spiculo-fibre ;
but in the new species the secondary fibres, usually not more
than one spicule thick, are often not present at or just below
the surtace.
Locality. Winter Quarters, 25-30 fath.
Gellius cucurbitiformis, sp. n.
Sponge small, free, bulbous, with fistular prolongations.
Surface smooth, showing under a lens a fine white reticulum,
Colour in spirit pale brown. Consistence rather soft.
Skeleton.— With a distinct dermal layer of irregularly
arranged tangential oxeas. Choanosomal skeleton a reti-
culum (with square or triangular meshes) of spiculo-fibre,
the strands 2-3 spicules thick, with a faint indication of main
fibres radiating to the surface.
Spicules.— Megascleres: oxeas, 342 x 9°75 yw, slightly
curved, subtornote.
Microscleres: sigmata, varying in size, the smallest being
about 20 w long, C-shaped, and with uniform curve, and the
largest 39 w long, 19°8 yw broad, and 1:2 p thick.
There are two small specimens, both of which were found
in a tangled mass of débris surrounding a worm-tube. The
of the National Antaretic Expedition. 289
larger, the type specimen, consists of a basal bulbous portion,
13 mm. long, 7 mm. broad, and 8 mm. high, from one side
of which arises a rather thick-walled fistula 13 mm. high and
5°5 mm. in diameter; at the opposite side is a broken circular
area, from which, in all probability, a second fistula arose ;
lastly, between these two is a small raised knob with a rounded
orifice on one side of it. The narrow oscular canal is not
central, but along one side of the thick-walled complete fistula.
The second specimen is tubular, with a slightly enlarged
solid base, whence arises a fistula; the total length is 2 em.,
and diameter’5cm. No pores are discernible; the subdermal
spaces are about *2 mm. in depth.
The eurypylous flagellated chambers are 23 w in diameter.
Cellules sphéruleuses, 8-9 w in diameter, are common.
There is no bast-like subdermal layer as in Oceanapia
mollis, Dendy, and the spicules of the latter are smaller, the
oxeas being 200 x 8 w, and the sigmata only 16 w. Lundbeck
describes two species of Gredlius with fistulae, and with a well-
developed dermal bark, viz. G. duridus and G. microtoxa,
but both these species have toxa in addition to sigmata.
Locality. Winter Quarters, 25-30 fath.
Oceanapia tantula, sp. n.
The sponge consists of five small fragments of tubes, the
longest of which is 8 mm. in length, by 4 mm. in diameter:
three of the pieces are hollow, thin-walled, and tubular; the
other two are solid. One of the solid pieces seems to belong
to the top of a fistula.
The colour is transparent white.
Skeleton.—The dermal layer is composed of a chitinous-
looking membrane with strongyles lying tangentially, usually
in one layer and densely packed, but sometimes more or less
scattered.
The white strands of the loose subdermal reticulum are
visible through the surface. They are longitudinal, and only
anastomose occasionally. The strands are less than *l mm.
in diameter. ‘They vary in composition ; in some parts being
composed of strongyles smaller than those of the dermal
layer, in other parts of smooth trichodragmata, or again of
strongyles, amphityles, and trichodragmata. ‘The pale trans-
parent choanosomal tissues are crowded with small spined
rhaphides.
Spicules.—Megascleres : strongyles, 437 x 19 y, slightly
fusiform, curved once or sometimes twice. Occasionally one
end is pointed, the spicule becoming a style.
Amphityles, 395 x 7:25 w, slightly fusiform; heads 13 pz
long, 9°75 yw broad.
290 Mr. R. Kirkpatrick on the Monaxonellida
Microscleres: long,smooth rhaphides, separate or in bundles,
forming part of the “subdermal reticulum, 650 x 2°5 pw.
Short, scattered, spined rhaphides, usually stylote, 162
long and about 2°5 p broad.
H. V. Wilson describes a species of Oceanapia, viz.
O. bacillifera, with strongyles, but it has sigmata.
Oceanapia (Phiewodictyon) singaporensis (Carter) has
strongyles in the dermal layer, but oxeas as well as strongyles
in the skeleton-fibres, and there are no microscleres.
The species of the Gelliine genus Rhaphisia have oxeas,
trichodragmata, and, in one species, toxa; but there are no
fistule, and there isno subdermal reticulum of spicular fibres.
Locality. Winter Quarters, 130 fath.
Petrosia fistulata, sp. n.
Sponge tubular. Surface smooth, showing the round
openings of the inhalant canals about -4 mm. in diameter
and close together.
Inner surface of the tube of the sponge finely or rarely
coarsely pilose, and showing the round openings of the ex-
halant canals about 1 mm. in diameter. Colour in spirit
pale yellow. Texture firm, but slightly compressible. Hury-
pylous flagellated chambers spheroidal, 24°5 w in diameter.
Skeleton formed of main fibres proceeding from the inner
to the outer surface, joined by secondary fibres one spicule
thick, so as to form obscurely quadrangular or hexagonal
tubes about ‘5 mm. in diameter; ends of spicules cemented
with spongin.
Spicules.—Oxeas, 492 x 24°4 wz, bent usually, or curved at
centre, subtornote.
There are four specimens, the two larger being uniformly
cylindrical and the smaller ventricose. ‘The largest is 6 cm,
long, the diameter being 2°1 cm. and the thickness of the
wall 5 mm.
The dermal membrane roofing over the inhalant orifices is
usually supported there by two or three single spicules
radiating to the centre. The pores are 95 w in diameter.
Small embryos about ‘76 mm. in diameter occur. The
new species comes nearest to the species from Kerguelen,
which Carter identified as Thalystas sub-triangularis, Duch.
& Mich., but which Ridley and Dendy regarded as synony-
mous with Petrosia similis (Ridley & Dendy).
The spicules of the Antarctic species are very much larger
than those of Carter’s, and partly in consequence of this the
skeletal network of the latter is much denser from a closer
approximation of the fibres.
of the National Antarctic Expedition. 291
Localities. Winter Quarters, 25-30 fath.; McMurdo Bay,
96-120 fath.
Reniera Scotti, sp. n.*
Sponge consisting of one or more fistulae. Texture very
soft and easily lacerated. Colour in spirit varying from
yellow to pale reddish. Outer surface varying from being
finely hispid to having large conules and meandrine ridges.
Inner surface of fistulae very finely hispid in the spaces
between the numerous orifices of exhalant canals. Flagellated
chambers large, hemispherical, 60 x 40 pw.
Skeleton formed of parallel longitudinal lines of main
fibres, about 2-6 spicules thick, curving outwards from the
inner to the outer surface, where they pass into the conules
and ridges ; secondary fibres at right angles to the main
ones, one or two spicules thick. The spicules are not closely
united, and spongin is only present in very small amounts.
Spicules.—Oxeas, 343 x 14°6 yw, curved or bent at centre,
subtornote.
There are six specimens and fragments. The outward
appearance varies greatly according to age and size. In one
small specimen the surface is finely hispid, in larger ones
conulose, and in very large ones conulated and with high
meandrine ridges. The largest specimen (No. 118) is in
form of a wide thick-walled tube, 12 cm. high and 6 cm. in
diameter, and with walls 1:5 cm. thick, but attenuating
towards the rim of the tube. This specimen is incomplete
below. The orifice is circular, and within the rim is a
diaphragm contracted to a white line.
The surface is covered with large conules and meandrine
ridges rising to a height of nearly 1 cm.
The dermal membrane, in the spaces between the conules
and ridge, shows as a fine lace-like reticulum, with circular
pores 133 w in diameter, and beneath it the orifices (1-1'5 mm.
in diameter) of the inhalant canals are visible. The ex-
halant orifices on the inner wall of the tube are much larger
than the inhalant ; they vary from 1 to 6 or 7 mm., their
edges are smooth and rounded.
The species closely resembles F. spznosella, Thiele, from
Punta Arenas. In Thiele’s species the body is tubular, with
conulated surface, and the texture is very soft; but the
skeletal framework is irregular, and the oxeas, though similar
in form, are much shorter, smaller, and more slender, being
only 150-170 pw long and 7-8 yp thick.
Localities. Winter Quarters, 5-100 fath.; off E. end of Ice
Barrier, 100 fath.
* This fine species is named after Captain Scott, R.N., the leader of
the Expedition.
292 On an apparently new Species of Moukey.
XL.— Description of an apparently new Species of Monkey of
the Genus Cebus. By D. G. Exuior, D.Sc., F.R.S.E., &e.
Cebus apiculatus, sp. n.
Type locality. La Union, Lower Orinoco. Type in
British Museum.
Colour.— Male. Face flesh-colour. Black spot on middle
of crown extending in a narrow line on to the forehead ; rest
of head greyish brown, becoming greyish white on forehead
and sides in front of ears; scape reddish brown; upper half
of back blackish brown; lower half of thighs and root of
tail blackish brown, all the hairs tipped with russet, giving
this part a reddish appearance ; arms to wrist on outer and
inner sides pale yellow, the hairs being blackish brown at
base and tipped with pale yellow, which becomes the dominant
colour; wrists and hands blackish brown; hairs towards
shoulders are pale yellow to the roots, the blackish-brown
Lases appearing at the elbows ; legs on outer side pale yellow,
becoming reddish below the knees, the hairs being blackish
brown tipped with pale yellow to the knees and then tipped
with golden; throat yellowish white; chest yellowish brown ;
rest of underparts dark brown in the centre of body, the
hairs pale yellow at base ; hands blackish brown, feet black ;
tail above like thighs until near tip, the hairs being black
tipped with pale yellow, tip blackish brown, beneath blackish
brown the entire length.
Measurements. Total length about 918 mm. ; tail 459 (skin).
Skull: total length 91; occipito-nasal length 81; hensel 59 ;
zygomatic width 59; intertemporal width 42; palatal
length 32; breadth of brain-case 52; length of nasals 19;
length of upper molar series 21; length of mandible 55;
length of lower molar series 24.
Type. Adult male. B.M. no. 5. 5. 24. 1. Collected
25th Oct., 1903, by Mr. Klages.
A female from the Guayapo, Lower Orinoco, has much
longer hair in front and on sides of head, standing out in the
shape of a semicrest; the upper arms are darker, being a
yellowish brown; the back 1s not so conspicuously tipped
with yellow and is more red at the rump, as are also the
legs; the black of the crown is broader and covers all the
back of the head; the tail is like that of the male; all the
underparts are blackish brown, only the roots of the hairs on
the chest being yellowish white.
Occurrence of Acanthoglossus in British New Guinea. 293
Another female from the same locality is much redder
above, the hairs tipped with yellow on the sides and with
ferruginous on dorsal region and on thighs; middle of head
from a point on the forehead to occiput blackish brown; back
of neck reddish brown like dorsal region; long hairs on
forehead and sides of head pale brown; arms reddish brown,
only a little of the pale yellow so conspicuous on the male
appearing near the shoulder; forearms, legs, hands, feet,
underparts, and tail like in the other female.
These three specimens resemble those of no species with
which I am acquainted; the style of coloration is alike in
all of them, although the tints vary somewhat from a blackish
to a reddish brown. All three are strongly speckled on the
lower back, legs, and tail by the light tips of the hairs, but
only the male has the very light arms. The hair on the head
of the male is short and compressed, but the hair on the head
of the females is long, loose, and in the form of a crest
standing upright on the forehead and away from the sides of
the head. The female last described has the back of the
head reddish brown like the neck, while the other has the
crown and nape alike, of a blackish-brown colour. In this
respect, however, they merely follow the habit of many
species of the genus Cebus, individuals varying greatly
among themselves, even from the same locality, in the pattern
exhibited on the crown of the head, as well as in the distri-
bution and variety of tints on various parts of the body.
I am indebted to the courtesy of Mr. Oldfield Thomas,
Curator of Mammalogy in the British Museum, for the
opportunity of describing the above specimens.
XLI.— On the Occurrence of Acanthoglossus in British
New Guinea. By OLDFIELD ‘THOMAS.
THE Long-nosed Echidna, Acanthoglossus* (otherwise
Zeglossus, still better known as Proechidna), has hitherto
only been known from the north-western part of New Guinea,
whence have come all the examples in the different Huropean
museums. Of these Mr. Rothschild possesses, in the ‘Tring
* IT do not admit that the name <Acanthoglossus is preoccupied and
rendered invalid by the existence of the earlier Acanthoglossa.
294 Occurrence of Acanthoglossus in British New Guinea.
Museum, uo less than ten skins, divided by him into three
subspecies.
It has therefore been with great interest that 1 have
examined a fine example of the genus obtained in British
New Guinea by Capt. F. R. Barton, and presented by him
to the National Museum. It was brought in to him at
Port. Moresby by natives, who had obtained it on Mount
Victoria, in the Albert Edward range, at an altitude of about
8000 feet.
A comparison of this specimen with those in the British
and ‘Tring Museums shows, as might be expected, a certain
number of differences, of very much the same character as
those that distinguish the three Dutch New Guinea sub-
species from each other, as described by Mr. Rothschild.
I therefore propose to form for it a fourth subspecies, and
to name it, in honour of its donor,
Acanthoglossus Bruijnit Bartont, subsp. n.
Fur long and thick, entirely hiding the spines over the
whole of the dorsal area, these being visible only on the
nape, sides of neck, flanks, back of rump, and caudal region.
Limbs densely hairy. No spines on belly. General colour
black (not brown) throughout, the head not lighter than the
body, and the under surface practically as dark as the upper.
The only parts not black are the hands and feet, which are
brown, grizzled with whitish. ‘The bases of the hairs of the
head are also rather lighter than the tips. Spines thin, short
(rarely attaining 80 mm. in length), white, very different to
the long, thick, “blackish spines of A. B. nigroaculeatus.
Skull with rather shorter and more curved beak than that
of the specimen measured in the ‘Catalogue of Marsupials:
Monotremes’; condyloid vacuities present ; palatal foramina
unusually long.
Dimensions of the type :—
Length of head and body 560 mm.
Skull: basal length 175; greatest breadth 59; palate
length 161; anterior palatine foramina 49.
Hab. Mount Victoria, British New Guinea. Alt. 8000’.
Type. Old female. B.M. no. 7. 7.17.5. Collected and
presented by Capt. F. R. Barton.
Or
Bibliographical Notices. 29
BIBLIOGRAPHICAL NOTICES.
I Go A-Walking. Through the Woods. With Illustrations by
Caartes Rerp, London: T. N. Foulis, 1907. 2s. 6d.
Tis book, which bears the above barbarous title, justifies its exist-
ence only through its illustrations, which are really beautiful. The
text, which is merely a compilation of scraps from various second-
rate writers, is puerile. Wie.
Nature in School.—No. 2. Edited by Borts Werysere, Priv. Doc.
St. Petersburg Univ. (Published by I. D. Sitin, Petrovka, Moscow.)
THE appearance of this monthly journal is an interesting sign of the
great activity of late years among the Russians with regard to
scientific pursuits. We are aware that capable and industrious
observers have travelled in regions scarcely known by name, and
undergone innumerable difficulties and hardships in order to enrich
the annals of exploration and research. Since the days of the great
Prjevalsky, for example, colleagues and followers have re-trodden
his paths, opened up new ground, and published their records. But
while some have worked in the desert and on the seas those at home
have not been idle, as the numerous university and society
transactions and proceedings abundantly testify. The journal
before us is devoted to the encouragement of the study of physics,
chemistry, and natural history in middle and elementary schools.
In the natural history section, Mr. N. M. Knipoviteh, a well-known
authority on marine conditions, continues his examination of the
Caspian Sea basin. Referring to plankton, a subject hitherto little
studied, his definition is worth reproducing :—
What is this plankton? A rain of corpses, incessantly falling from above
in proportion to the death-rate among the animal and vegetable organisms
inhabiting in abundance the upper strata of our basin. This rain of corpses
does not present any peculiarity exclusively belonging to the Caspian Sea. In
any considerable extent of salt and fresh water on the terrestrial sphere the
same phenomenon occurs.
Among the Caspian plankton have been found Chiridotea entomon
and Cardium edule, which belong to northern seas and lakes. Next
Mr. Knipovitch discusses the herring-fisheries of the Volga and
Caspian, and urges that care should be taken by fishing communities
to avoid exhaustion and waste of the Volga sources of the important
herring industry. Passing to geology, the author traces the changes
which have taken place since the Sarmatian Sea extended from
the foot of the Alps to Tian-shan, and concludes: “ the Caspian Sea
may be considered an example of a basin with anomalous physico-
geographical and biological conditions.” Mr. V. Poloytsoy deals with
296 Bibliographical Notices.
country excursions. To junior students, names like nightingale,
squirrel, hare, &c. are only familiar from the written symbols in
school books, and the results of inquiry among Russian and German
children are set out in this table:—
German. Russian,
Children who have not seen: per cent. per cent.
Lark rising with song ......seesseeeeee- 25 50-60
Squirrels in a WOOd ......secseseeceeeees 71 58-66
Fumi NANG hace cose sae cei sieceeserieete 3 50-54
INTOGE Tan SAconpaceapsnoocuoncodacdbsocacoddes 22 30-59
Nests on a tree ........... sepseesesenesiet 15 3-11
Snailicrawlin Can vescsntesetiadecneateteses 2 40-59
Not heard :
Nightingale’s Song .........cesseseeeeseess 56 40-44
OPERONS Ae Socancsccnaddancanoo5ons00000 11 20-52
Not visited :
IAW GaihyslOXOSUi sescsneecisesectecrineesse secs 44 10-il
One of acerous leaves ..5..0......s0cc-cn. 37 12-17
Tn other words, about half the children are as if blind or deaf to
rural sights and sounds. This leads the writer to exclaim at the
expenses during a decade of school life and overwork of brains in
order to learn word combinations. This can only be remedied by
teachers, not merely by programmes. Especial interest attaches to
the article by Mr. Sukatshev on the forest as an association, as an
object for winter study. The mutual effects of trees on their growth
side by side are contrasted with the development of solitary
specimens. ‘There is a struggle for life among crowded trees, and
the weakest goes to the wall. There are distinctions between
simple and complex associations. This article is suggestive of
original research and aspects often overlooked. The results of
winter zoological explorations round the St. Petersburg Government
are given by Mr. 8. A. Petrov. The winter habits and colouring of
animals and birds are described.
In the department of chemistry, Mr. S. Sozonov continues his
examination of a government scheme for chemical instruction in
Realschulen. “We writes: *‘ Dogmatism in an elementary course of
chemistry, as of physics, should be reduced to the minimum possible.
To eliminate it altogether is unfortunately out of the question at
present.” Mr. V. Verkhovsky continues his description of heating
apparatus for chemical experiments in middle schools, and the study
of physics in village and urban schools is treated by Mr. V.
Lermantoy. ‘Russian students,” observes the latter, ‘“ are
generally lacking in attention: they understand more easily a
serious but short deduction, than a simple but ‘ many-storied’
reasoned conclusion extending over pages.’’ ‘There is a favourable
notice of a new German solder-paste, “tinol,’ made at a Bonn
factory.
The remainder of space is occupied by reviews of works on in-
organic chemistry and elementary physics.
Francis P, Marcnanr.
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CONTENTS OF NUMBER 117.—Seventh Series.
XXVII. Notes from the Gatty Marine Laboratory, St. Andrews.—
No. XXVIIL By Prof. M‘Intosa, M.D., LL.D., F. R. 8., &e.
(Plates Vi—-ViIth) ae
XXVIII. Descriptions of apparently new Species and Re is
of Mammals belonging to the Families Lemuride, Cebide, Calli-
trichide, and Cercopithecide in the Collection of the Natural History
Museum. By D. G. Extior, D.Sc., F.R.S.E., &.
XXIX. On Mammals from Northern Persia, presented to the
National Museum by Col. A. C. Bailward. By OLDEIELD THomas,
EP CBR A eae Bae aren ee AP ata pe aay SACL Say sige cs ee
XXX. A Subdivision of the Old Genus Nesokia, with Descriptions
of Three new Members of the Group, and of a Mus from the
Andamans, < By OLDFraLD THOMAS 37 f Pee ice ee ae in
XXXI. On some British Polyzoa. By Canon A. M. Normay,
MA. D.C. LED PRS. RS Binte Ty nee cee ae
XXXII. Three new Spanish Insectivores. By Ancun Carrera .
XXXITI. On Four new Pill-Millipedes from the Malay Peninsula
and Siam. SBy-.A. §. Hirsr (British Museum, Nat. Hist.).
(Plate X.) ;
XXXIV. Description of an apparently new Lycenid from Mauri-
dius. By Hamitcon H.-Deuce, PAS SACU. eo ae cee ea wee
XXXY. On an Extinct undescribed Fruit-Bat of the Genus °
Pteropus from the Mascarenes. By Guo. E. Mason.............:
XXXVI. Brachiopod Nomenclature: Seminula, &, By 8. 8.
Buckman, F.G.S8.
XXXVITI. On some Freshwater Amphipods: The Reduction of
the Eye in a new Gammarid from Ireland. By Prof. Dr. Fr.
NBIDOVSEY, << (Plates 6 PW KL os Se ec Fe 8 eee eee
XXXVIII. Observations on the Trematode Parasites of British
Birds. By Wiitram Nicotr, M.A., B.Sc., Gatty Marine Laboratory,
St. Andrews...... ‘
eeceetececsere eee eee ee ew eres ete ee tee ees eesesn
eeervreceete eet teeeeeesteeeseeveeececeeer ees eneee
_ XXXIX. Preliminary Report on the Monaxonellida of the
National Antarctic Expedition. By R, Krrxparrick ............
XL. Description of an apparently new Species of Monkey of the
Genus Cebus. By D. G. Extrot, D.Sc., F.RS.E., &.........0...
XLI. On the Occurrence of Acanthoglossus in British New Guinea.
By Orprmrp THOwAs 25 es oS. ee BS RG Gres Aen RS,
{ Plate IX. will be issued with the October Number.
BIBLIOGRAPHICAL NOTICES.
I Go A-Walking. Through the Woods. With Illustrations by
Cuantes Rerp
ee SP ee Se Se et
Nature in School.—No. 2. Edited by Boris Wetnzure, Priy. Doc.
St. Petersburg Univ.
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185
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. 212
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292
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XLIL.—New South-African Spiders of the Family Drassidze
in the Collection of the South-African Museum. By W.F.
Purce.t, Ph.D., Cape Town.
[Plates XIII.-XV. ]}
THE spiders of the family Drasside are particularly
numerous both in species and individuals throughout all the
non-tropical parts of South Africa. Only very few South-
African species have, however, as yet been described, and
the entire collection in the South-African Museum, with the
exception of one or two forms, is still new to science. The
present paper contains the descriptions of fifty-three new
species, being that portion of the Collection pertaining to the
genera Platyoides, Cambr., Theuma, E. Sim., Drassodes,
Westr., Scotopheus, HE. Sim., and Melanophora, C. Koch,
as defined in Simon’s Hist. Nat. Araign. 2nd ed., together
with two new genera for certain forms which did not seem to
fall into any of the genera recognized by Simon. In the
present uncertain state of the classification of the Drasside
all of these genera are to be considered as more or less of a
temporary nature. All localities are in Cape Colony, except
when otherwise stated.
Genus PLAtTyoIpESs, Cambr.
1. Platyoides bidentatus, sp. n.
(Plate XIII. figs. 1 & 2.)
Specimens.—(a) 3 99 and 1 ¢ from Kentani and
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 20
298 Dr. W. F. Pureell on
District, collected by the Rev. Dr. F. C. Kolbe and Miss
Alice Pegler; May.
2 2 (types).— Colour of carapace and chelicera mahogany-
brown; sternum and three basal segments of legs yellow,
the sternum with a deep blackish-red border all round, the
remaining leg-segments (especially the patellee) and the apex
of each femur more or less infuscated ; abdomen blackened
above, with or without a longitudinal row of more yellowish
areas on each side of the median line, the underside paler,
yellowish, whitish, or more or less infuscated.
Carapace longer than broad, its median length equal to
that of the tibia and ? or slightly more of the metatarsus of
first leg, its margins with slender spines. Anterior median
eyes about a diameter apart, their distance from the anterior
laterals about 12 times their distance from one another.
Posterior median eyes obliquely oval, about two long diameters
apart and about three from the posterior laterals.
Chelicera with spiniform sete above, the inner margin
with a dense stripe of slender setee borne on small granules
and with 2 rather strong and conspicuous teeth in the
stripe.
Legs with spines and spiniform sete on the basal segments
above and with long black sete on the other segments ; tarsi
and metatarsi of two anterior pairs with scopula below, that
on the first metatarsus extending over the distal half at least,
but that on the second confined to the apex of the segment; ,
tibiz not scopulate.
Vulva as in Pl, XIII. fig. 1; the T-shaped depression
with the longitudinal portion finely pointed posteriorly and
the transverse portion slightly recurving at each end.
d.—Very much smaller, being less than half as long as
the 9.
Carapace yellowish, its length as in the @? ; the eyes
closer together, the anterior medians at least as close to the
laterals as to one another, being an eye’s diameter or less
from them; the posterior median eyes about a diameter from
one another and 14 diameters from the posterior laterals.
Chelicera toothed as in the 9.
Pedipalps short; the femur cylindrical, longer than the
patella and tibia together, with mesial spine above; the tibia
shorter than the patella, with small apical process on upper
outer edge; the tarsus strongly convex, ovate, obtusely
lobate at base on outer side; palpal organ as in PI. XIII.
fig. 2.
Length of trunk (carapace and abdomen) 2? ? 113-12,
g 42, of carapace 2 43, ¢ 2; width of carapace ? 44,
& 14mm.
new South-African Spiders. 299
(b) 1 2 from East London (J. Wood).
(c)2 99 from Port St. Johns, Pondoland (Guy
Shortridge).
Somewhat resembling P. Abrahami, O. P. Cambr., and
P. laterigradus, Poc., in the form of the vulva, but
possessing two strong teeth on the chelicera.
2. Platyoides quinquedentatus, sp. n.
é (Plate XIII. fig. 3.)
Specimens.—2 9 § and 1 young from Swellendam (/7/.
A. Fry, 1900).
? 9.—Colour of carapace and chelicera dark mahogany-
brown; legs similarly coloured or lighter and more yellowish,
the anterior pairs more reddish than the posterior ones, all
partially infuscated, the femora not paler than the distal
segment ; abdomen ‘blackened, with a paler, more or less
yellowish, longitudinal dorsal stripe on each side above, the
ventral surface pale yellowish; sternum reddish yellow.
Carapace as long as the tibia and 4—3 of the metatarsus of
first leg, the margin with spiniform sete and slender spines.
Anterior median eyes almost a diameter apart and almost two
diameters from the laterals, their distance from the latter
being double their distance from one another; posterior
median eyes almost transversely oval, about 24 diameters
apart and almost 3 diameters from the laterals.
Chelicera with 5 very strong teeth in the distal half of
the inner margin, besides the row of setee ; upperside with
spiniform sete.
Legs with spiniform sete on basal segments, the distal
segments with slender sete; two anterior pairs of tarsi and
metatarsi scopulate along whole length below, the first tibia
also with tiny apical scopula.
Vulva of two black pear-shaped depressions, appearing (in
spirits) as in Pl. XIII. fig. 3.
Length of trunk in 9 ? 11-123, of carapace 44-5; width
of carapace 44-44 mm.
Genus THEUMA, H. Sim.
1. Theuma capensis, sp. n.
Specimens.—(a) 10 ¢ 2, 6 ¢ g, and 1 young from the
drier slopes of the mountains of the Cape Peninsula, especially
the slopes above Camps Bay. ¢¢ from September to
December.
2 ?.—Colour yellow, the head, chelicera, tarsi, and meta-
tarsi more rufescent; the abdomen pale yellowish, with
yellowish or pale fuscous hairs and darker sete.
20*
300 Dr. W. F. Purcell on
Anterior row of eyes scarcely procurved, the medians
distinctly smaller than the laterals; posterior row of eyes
slightly recurved, the eyes equal, the medians slightly
oblique, about a diameter (rarely distinctly less than a
diameter) apart and 14 diameters from the laterals, the latter
distinctly a little nearer to the anterior laterals than to the
posterior median eyes.
Chelicera with 4-5 superior and only 2 (rarely 3) inferior
teeth.
Legs.—Metatarsus I with 1-2 inner and 2 outer inferior
spines, narrowly scopulate on each side, but not between the
rows of spines below. ‘Tibia I with 4-5 inner and 3-6 outer
spines below, not distinctly scopulate or at most with a few
scopulate hairs along the rows of spines.
Vulva-plate longer than wide, the triangular fovea wide
behind, narrowed in front.
Inferior spinners relatively short, the length of the basal
segment taken below a little less than the width of the ocular
area.
3 3 .—Legs longer; metatarsus I with 2 inner and 3
outer spines below; tibia I with 4 pairs of inferior spines
and sometimes with an inner lateral spine as well.
Inferior spinners longer, the length of the basal segments
exceeding the width of the ocular area.
Tibia of pedipalps short and thick, its outer process longer
than the segment and suddenly deflexed at apex. Tarsus
with a short curved process near the middle of the outer
edge.
Length of trunk (carapace and abdomen), ? 2? 5-7, ¢ o
44-5 mm.
A variety of this form with four minute teeth instead of
two on the inferior inner margin of the chelicera is represented
by the following specimens :—
(b) 1 g from Ceres (W. F. P.).
(c) 1 @ from near Tulbagh Road Station (W. F. P.).
(d) 1 2 from Caledon.
2. Theuma fusca, sp. n.
Specimens.—(a) 6 2 ? from the Montagu Baths, a number
of young from Kogmans Kloof, and 2 ¢¢ from Aston,
Robertson Div., all found by my wife and myself.
? ¢ .—Colour ochraceous, head and chelicera rufescent ;
abdomen pale yellowish below, densely clothed above with
infuseated hairs.
Anterior row of eyes slightly procurved, the laterals very
new South-African Spiders. 301
distinctly larger than the medians; posterior row more
strongly recurved than in 7. capensis, sp. n., the medians
smaller than the anterior laterals, rotund, about a diameter
(rarely less) apart and less than two diameters from the
laterals, the latter eyes perhaps a trifle larger than the
medians and about as far from them as from the anterior
laterals.
Chelicera with 3 larger and 1 small superior and 3-4
inferior teeth.
Metatarsus I with 1-2 inner and 2 (rarely 1) outer spines ;
tibia I with 3 pairs of inferior spines (including an apical
pair) and sometimes with another internal spine as well.
Vulva-plate with the triangular fovea wider behind, its
anterior end opening into a small circular fovea.
Inferior spinners long, the length of the basal segment
much exceeding the width of the ocular area (except in one
specimen where the spinners were perhaps damaged in youth).
$ $.—Legs much longer than in the 9 2 and more
numerously spined. Metatarsus I with 4 strong spines.
Tibia I with 0-1 outer and 1-3 inner spines in addition to
the 3 inferior pairs. Posterior median eyes sometimes less
than a diameter apart. Hairs on abdomen rubbed off.
Pedipalps as in 7’, capensis, sp. n.
Length of trunk, 9 2 82-11, ¢ g 63-72 mm.
(b) 2 $¢ and 1 young from Bergvliet, Cape Peninsula
Ca Es).
3. Theuma Schreinert, sp. n.
Spectmens.—2 2 2 and 5 @g from Hanover (S. C.
Cronwright Schreiner, September to November 1901, and
G. Shortridge, 1903).
Differing from 7. fusca, sp. n., as follows :—
9 2 .—Hairs on abdomen paler, brownish.
Posterior row of eyes straighter, only very slightly re-
curved; the eyes subequal, the lateral eyes distinctly a little
nearer to the anterior laterals than to the posterior median
eyes.
: Chelicera with 4—5 superior teeth, the proximal one small.
Tibia 1 with 3-4 pairs of non-apical and 0-2 apical spines
below, and frequently with 1-2 outer and 1-2 inner spines on
the sides as well.
¢ $.—Metatarsus I with 5-7 spines. Tibia I with 5-6
pairs of spines below, besides 2 outer and 2-4 inner spines on.
the sides.
Length of trunk, 9 9 7-83, 3 g 63-8 mm.
302 Dr. W. F. Purcell on
Another ¢ from the same place and apparently belonging
to the same species is much larger, its carapace alone
measuring 43 mm. in length.
4. Theuma maculata, sp. n.
Specimens.—2 2 3 (one not quite mature), 1 g, and 1
juv. from Beaufort West (W. F. P., September 1896).
2 2 (types).—Colour yellow, the abdomen pale yellow
below, spotted with black on the sides; the upper surface
with a broad irregular band of black on each side and a
narrow median black stripe, connected with the lateral bands
by a series of oblique black stripes.
Anterior row of eyes very distinctly procurved, the laterals
very distinctly larger than the small medians; posterior row
distinctly recurved, the median eyes almost rotundate and
rather large, considerably larger than the anterior medians,
and nearly as large as the anterior laterals, their distance
apart distinctly less than a diameter, the lateral eyes equi-
distant from the medians and anterior laterals.
Chelicera with 3 large and 1 small superior and 3-4
inferior teeth.
Spines of legs, vulva, and spinners much as in the 7’ fusca,
sp. Nn.
: ¢ .—Like that of 7. fusca, apart from the ocular cha-
racters, except that the tibia of the pedipalps is much more
elongate, being distinctly longer along the inner side than
thick, its length equalling that of the outer process.
Length of 2 7, & 74 mm.
d. Theuma cedr?, sp. n.
1 ? from Boschkloof Waterfall, Cedar Mountains, Clan-
william Division (i. Pattison, November 1897).
Colour yellow ; the head, chelicera, and extremities of the
legs rufescent ; abdomen pale yellow, the upperside clothed
with dark hairs, which form spots posteriorly (the hairs
rubbed off in the middle, but probably arranged as in 7. macu-
lata, sp. n.).
Eyes and spinners much as in T. maculata.
Metatarsus I with 4 strong spines. Zvb¢a I with 3 pairs of
inferior spines (including an apical pair) and 2 internal spines
in addition.
Chelicera with 3 superior and 3 inferior teeth, the latter
stronger than usual.
Vulva-plate with the fovea elongate and narrow behind.
Length 9 mm.
oe
new South-African Spiders. 30
~
6. Theuma mutica, sp. n.
1 @ found at Kogmans Kloof, Robertson Division, in
August 1900, by my wife.
Colour.—Carapace more or less infuscated, darkest at the
lateral margins, paler and yellowish along the middle of ante-
rior half and along posterior border. Chelicera and sternum
ochraceous. Legs yellowish, faintly infuscated in parts.
Abdomen pale yellowish below, deeply infuscated above and
at the sides.
Anterior row of eyes slightly procurved, the laterals much
larger than the medians. Posterior row of eyes straight, the
medians oval, oblique, very close together, separated by less
than half a length, but at least an eye’s length from the
laterals; these latter scarcely larger than the medians and
close to the anterior laterals, being less than a diameter from
them.
Chelicera with 2 larger proximal and 3 smaller distal teeth
in the superior row, and only 1 minute tooth in the inferior
row.
Legs with numerous fine small spines scattered irregularly
on the under surface, the first leg without any strong spines
below; second leg with a strong mesial spine on tibia; third
and especially fourth leg strongly spined on underside as well
as on upper.
Vulva-plate broader than long, consisting of a pair of oval
reddish disks separated by a narrow triangular fovea.
Inferior spinners long.
Length 45 mm.
A very distinct species.
7. Theuma parva, sp. n.
1 g from Eierfontein, 8-9 miles west of Hanover (S. C.
Cronwright Schreiner, January 1902).
Colour very pale yellowish.
Anterior row of eyes slightly procurved, the medians a little
smaller than and touching the laterals, but a little separated
from each other. Posterior row of eyes straight (scarcely
recurved at all), the subrotund median eyes comparatively
large, being larger than the laterals and at least as large as
the anterior laterals, their distance from one another less
than an eye’s diameter and slightly greater than their distance
from the posterior laterals, the latter eyes about as far from
the anterior laterals as from the posterior medians.
Chelicera with 4 superior and 3 smaller inferior teeth.
304 Dr. W. F. Purcell on
Legs.—Metatarsus I with 3 long spines at base. Tibia I
with 3 pairs of inferior spines (including an apical pair).
Pedipalps very like those of T. capensis, sp. n., except that
the bulb is relatively much larger than in any of the foregoing
species.
rics spinners shortish, their length only just exceeding
the width of the ocular area.
Length of carapace and abdomen 3? mm.
A very distinct species.
Table of the Species of Theuma * described above.
a. Posterior row of eyes straight, the medians
being a little nearer to the laterals than to
one another. (Hanover Div.) .......... 3 T. parva, sp. n.
&. Posterior row of eyes more or less recurved,
the medians always nearer to each other
than to the lateral eyes.
a*, Posterior row of eyes only slightly re-
curved, the laterals being distinctly a
little nearer to the anterior laterals than
to the posterior median eyes.
a’. Females.
a‘, Inferior spinners shortish, the length
of the basal segment taken below
being less than the width of the
pore row of eyes. (Cape, Tul-
pagh, and Caledon Divs.) ........ © T. capensis, sp. 1.
b*. Inferior spinners long.
a’, Anterior pair of legs strongly
spined below. (Hanover.)...... Q T. Schreineri, sp. 0.
b°. Anterior pair of legs with no strong
spines below. (Robertson Div.) Q T. mutica, sp. n.
6°. Males.
a°. Tibia I with 4 pairs of spines below.
(Cape and Ceres Divs.) ........ .. & T. capensis, sp. n.
b°. Tibia I with 3 pairs of spines below.
(Hanover) 7 ie kiscnccemeweeer do T. Schreineri, sp. n.
&*. Posterior row of eyes more strongly re-
curved, the laterals being about as far
from the anterior laterals as from the
posterior median eyes.
a. Q with the posterior median eyes
smaller, about a diameter apart. 6
with the outer process of the tibia of
pedipalps much longer than the short
tibia. (Robertson and Cape Divs.).. ¢ 2 T. fusca, sp. n.
* Two other South-African species, T. zylina and T. aprica, E. Sim.,
have been described (Hist. Nat. Araign. 2nd ed. p. 351), but no locality
is given in either case.
new South-African Spiders. 305
b", 9 with the posterior median eyes
largish, less than a diameter apart.
¢ with the outer process of the tibia
of pedipalps about as long as the
rather elongate tibia.
a*. Vulva of 2 with the fovea widened
behind. (Beaufort West.) ...... 3 Q T. maculata, sp. vn.
b°, Vulva with the fovea narrow behind.
(Clanwilliamy Diy. )\ie.r. ccrtie sce csi © T. cedri, sp. n.
Genus DrassopEs, Westr.
1. Drassodes lophognathus, sp. n.
(Pl. XIII. figs. 4-6.)
Specimens.—(a) 52 2?,4¢ 3, and7 young from the north-
western slopes of the Devil’s Peak, Cape Town, under stones
(WaE-P.):
2 2 (types). Colour.—Carapace pale ochraceous, the head
often darker. brown; chelicera ochraceous to brown; legs
pale yellow, the femora very pale, the anterior pairs of legs
often brownish yellow distally ; sternum pale ochraceous to
brownish yellow ; abdomen pale yellowish, the upper surface
and the posterior part of the sides marked with numerous
short black bars and round spots, the anterior half also with
a broad, wedge-shaped, median black band, the under surface
and the sides pale yellow or more or less infuscated.
Carapace broad in front, the width of the head across the
second line of eyes being at least 3 of the greatest width of
the carapace.
Eyes.—Anterior row considerably procurved, the eyes sub-
equal; the posterior row lightly procurved, the median eyes
elongate oval or triquetrous, subcontiguous or as much as
half a long diameter apart, and almost or quite a long
diameter from the smaller lateral eyes. Median eye-area
longer than wide and parallel-sided.
Chelicera with the two inferior teeth of moderate size, the
distal one being usually a trifle smaller; middle superior
tooth very strong, the other two superior teeth small.
Legs.—Metatarsus I with Q-2 inferior basal spines.
Tibia I unspined, IV without dorsal spines. ‘Tarsi I-III
and metatarsi I-II distinctly scopulate at the sides below ;
tarsus IV and metatarsus III with narrow strip of scopular
hairs, the fourth metatarsus with a few external scopular
hairs in some specimens at least.
Vulva-plate (Pl. XIII. fig. 6) very large, consisting of a
brown, horseshoe-shaped curved rim enclosing a large deep
306 Dr. W. F. Purcell on
cavity, which is divided longitudinally by a broad, pallid,
wedge-shaped keel, dilated anteriorly to form a T; hind part
of the rim with a pair of dark admedian spots.
& g.—Anterior part of head and the chelicera and other
mouth-parts often reddish black, the abdominal spots also
generally more sharply marked.
Coa of pedipalps strongly depressed, transversely ridged
in the middle, the depression bordered on the outer side as
well as on the inner by a well-marked keel, which is absent
on the outer side in the female.
Chelicera with the inner margin sinuated near the middle
and lobate at the base, the superior distal edge of the inner
margin with the two proximal teeth fused to form a low keel
on which the point of the proximal tooth is usually not dis-
tinguishable; this margin also much more oblique distally
than in the @.
Pedipalps short; the tibia together with its strong superior
process longer than the patella, this process broadly and
obliquely truncated at apex and slightly sinuated on upper
inner margin near apex, the lower edge acute at apex; tarsus
large, as long as the patella and tibia together (inclusive of
the tibial process), somewhat polygonal in outline (Pl. XIII.
fig. 4), the outer edge being slightly sinuated, the inner edge
with a few fine spiniform sete. Palpal organ very large,
asnin 2), OU fies 5:
Length of trunk, 2 2 53-8, ¢ ¢ 5-6 mm.
(b) Other specimens from the Cape Peninsula (Table
Mountain, Camps Bay, Cape Flats, Simonstown). In one
of these (a 2) the chelicera have three inferior and four
superior teeth.
(c) 1 Q from Stellenbosch (L. Péringuey).
(d) 5 2 Q from the Pass at Avontuur near Stormsvlei,
Swellendam Division, collected by my wife and myself.
(e) 1 g and 1 @ from St. Helena Bay, Malmesbury
Division (J. L. C. Goold).
(£) 1 g from Onder Berg Vlei, Clanwilliam Div. (C. Z.
Leipoldt).
(g) 1 2 from the Hex River Valley, Worcester Div.
(Ff. Treleaven).
(h) 3 2 2 trom Touws River, Worcester Div. (W. F. P.).
(i) 8 g@ dg ancd7 Q 2 from Matjestontein, Worcester Div.,
and
(j) 1 g and 4 2? 9 from Laingsburg, Pr. Albert Div.,
coliected by Mr. R. M. Lightfoot, my wite, and myself.
(k) 1 ? from Beaufort West (W. F. P.).
new South-African Spiders. 307
(1) 3 2 2 from Willowmore (Dr. H. Brauns).
The T-shaped median ridge of the vulva is normally white
in colour, but occasionally it is more or less lightly browned ;
the posterior part, too, 1s sometimes broad behind and
parallel-sided, instead of being wedge-shaped. The carapace
of the 2 does not exceed 2# mm. in length in the largest
specimen, that of the largest g being slightly less. ‘lhe
anterior lateral eyes are their own diameter or less from the
anterior margin of the carapace. The spots on the abdomen
are occasionally absent. In some ¢ ¢, notably those from
Matjesfontein, the two proximal teeth of the superior margin
of the chelicera are less completely fused and separately
distinguishable.
The species is closely allied to D. morosus (O. P. Cambr.)
from Palestine, the figure of whose palp, given by Cambridge
(P. Z. 8. 1872, pl. xv. fig. 9), almost exactly resembles the
palp of D. lophognathus, sp.n. In D.morosus, however, the
eyes of the posterior row are said to be equidistant from each
other. D. emissus (O. P. Cambr.), from Palestine, has a
very similar vulva (P. Z. 8. 1872, pl. xv. fig. 17).
2. Drassodes tessellatus, sp. n.
(Pl. XUL. figs. 7 & 8.)
Specimens.—(a) 17 g g and 45 2 9, besides a pair in
copulé ; also6 2 ¢ with variety of the vulva; all collected
at Hanover by Mr. 8. C. Cronwright Schreiner in 1901.
Very closely allied to D. lophognathus, sp. n., but larger,
and differing principally in the form of the proximal tooth on
the inner side of the large sigmoid process of the palpal
organ, this tooth (which is hidden under the tarsus) forming
a broad truncated lobe and much broader than the distal
tooth (Pl. XIII. fig. 8).
Colour.—Carapace often rufescent, the head and chelicera
red, the legs yellowish or reddish yellow, the abdomen
speckled as in lophognathus.
Posterior median eyes slightly more than an eye’s diameter
from the laterals, and the anterior lateral eyes generally
slightly more than a diameter from the anterior margin of
the carapace.
Chelicera of § 8 with the two proximal teeth on superior
inner margin fused to a keel or more or less separate.
Vulva of 2 2 normally resembling that of lophognathus,
the median T-shaped ridge being generally whitish ; often,
however, this ridge is browned or reddened, and in the
308 Dr. W. F. Purcell on
6 ¢ ¢ mentioned above it is dark brown or black, the trans-
verse cavity in front of it then often with a brown margin,
as an IE) exo ties 7;
Length of trunk in 2 reaching 103 mm.; length of cara-
pace in 2 up to4 mm.
(b) 1 g and1 ? from Steynsburg Division, Cape Colony
(G. G. Ponder).
3. Drassodes solitarius, sp.n. (PI. XIII. fig. 9.)
1 ¢ from Hanover (S. C. Cronwright Schreiner).
Very like J. tessellatus, sp. n., but still larger and with
somewhat different vulva,
Colour.—Carapace rufescent, the head and chelicera deeper
red, legs ochraceous; abdomen pale yellowish, the dorsal
spots only faintly marked, the anterior half with a median
pale dorsal stripe bordered on each side by a darker stripe.
Anterior median eyes slighty nearer together than in ¢essed-
latus, their distance apart being scarcely twice their distance
from the lateral eyes; posterior median eyes distinctly more
than a diameter from the posterior laterals.
Chelicera with all the teeth rather strong, the middle
superior tooth very strong.
Vulva asin Pl. XIII. fig. 9, the horseshoe rim being rela-
tively broader and more transverse and with its anterior ends
more strongly converging than in the preceding forms.
Length of trunk 10, of carapace 45 mm.
4, Drassodes caffrerianus, sp.n. (PI. XIII. fig. 10.)
1 g from the Keneha Bridge, about 22 miles west of
Maclear, Pondoland (A. S. Weisbecker, August 1903).
Colour pale yellow, with black reticulation and hairs; legs
pale yellow, many of the segments faintly infuscated ; abdo-
men black above, the underside yellowish but infuscated ;
sternum pale yellowish, with black edges. (The specimen,
when caught, had evidently but recently moulted, and
maturer specimens would probably be much darker.)
Closely allied to D. lophognathus, sp. n., but differing
principally as follows :—
Chelicera only very feebly sinuated on inner edge, the
large tooth of the superior inner margin not keel-like, but
erect and conical, and accompanied by a tiny second tooth on
its proximal side.
new South-African Spiders. 309
Pedipalps closely resembling those of lophognathus, but
with different dentition on the medial distal process (Pl. XIII.
fig. 10).
Length of trunk 42 mm.
5. Drassodes calceatus, sp.n. (PI. XIII. fig. 11.)
2 2 2 from Matjesfontein.
Very like D. lophognathus, sp. n., but with the vulva
different and shaped as in Pl. XIII. fig. 11.
Length of trunk 5-5$ mm.
The abdomen is much more thickly spotted on the ventral
surface than is the case in any of the female specimens of
lephognathus from the same locality.
6. Drassodes Gooldi, sp.n. (Pl. XIII. fig. 12.)
2 2 2 from Stompneus, St. Helena Bay, Malmesbury
Div. (J. E. C. Goold).
Closely allied to D. calceatus, sp.n., but larger and without
the infuseate spots on the abdomen.
Vulva as in Pl. XIII. fig. 12; the anterior margin of the
anterior cavity brown, and therefore much more distinct than
in calceatus, and also more strongly emarginate in the middle.
Length of trunk 8, of carapace 34 mm.
7. Drassodes lyratus, sp.n. (Pl. XIII. fig. 13.)
1 ? from Matjesfontein, Worcester Div.
Very like 1). lophognathus, sp. n., but with the vulva
different, as in Pl. XIII. fig. 13.
The abdomen has two fine longitudinal stripes below
behind the vulva, but there are no dark spots on the dorsal
surface.
Length of trunk 5} mm.
8. Drassodes helene, sp. n.
(Pl. XIII. figs. 14 & 14a.)
1 ¢ from Stompneus, St. Helena Bay, Malmesbury Div.
(J. E. C. Goold).
Allied to D. lophognathus, sp.n., with similar maxille, &e.,
but of larger size and with somewhat different pedipalps.
Carapace rufescent, the head and chelicera blackish red ;
abdomen numerously spotted above, the under surface
without spots.
310 Dr. W. F. Purcell on
Anterior legs unspined.
Posterior median eyes large, angular, subcontiguous, and
about their own diameter from the laterals.
Pedipalps with the tarsus narrower and symmetrically oval
in outline when seen from above, its length almost equalling
that of the patella and tibia together (inclusive of the tibial
process) ; surface of the tarsus with slender scattered spines
and spiniform setee, the inner edge with some stronger curved
spines, the outer edge convex, not sinuated. Palpal organ
also smaller, appearing as in P]. XIII. fig. 14, when seen from
below; the medial distal process of the bulb with simple apex
and provided near the middle on the upper surface with a
single erect tooth (only seen on lifting the tarsus, as in
Pl. XIII. fig. 14a).
Chelicera with strongly oblique inner distal margin, the
upper margin with a strong keel-like tooth remote from the
apex, the medial sinus short and deep.
Length of trunk 73, of carapace 3? mm.
This may, perhaps, be the ¢ of D. calceatus, sp. n., from
the same locality, but in the latter the infuscated markings
on the abdomen are obsolete, while they are strongly marked
in D, helene.
9. Drassodes Dreget, sp.n. (Pl. XIII. fig. 15.)
2 2 2 from Port Elizabeth (J. LZ. Drége, July 1899).
Colour of carapace yellow or rufescent, the head and cheli-
cera reddish; legs yellowish or reddish yellow; abdomen
pallid, with numerous black spots in rows, the under surface
with or without spots ; sternum reddish yellow, with darker
edges.
Closely resembling D. lophognathus, sp. n., but with
different vulva, the transverse cavity and fold in front of the
horseshoe-shaped ridge found in all the females of Drassodes
described in the preceding pages being absent. Appearance
of vulva in spirits as in P]. XIII. fig. 15.
Chelicera with the two inferior teeth rather small, slightly
unequal.
Length of trunk 6 mm.
10. Drassodes ereptor, sp. 0.
(Pl. XUE: tess iio ae a7.)
Specimens.—(a) 5 ¢ 2 from the Hot Baths at Montagu
(W. F. P., November 1902).
? 9. Colowr.—Carapace and sternum mahogany-brown,
new South-African Spiders. S11
the chelicera slightly darker ; legs yellowish brown, the two
anterior pairs with the three distal segments darker brown ;
abdomen more or less infuscated, the ventral surface generally
aler.
: Carapace broad; width of the head across the second line
of eyes a little more than half but less than two thirds of the
greatest width of the carapace. Anterior row of eyes pro-
curved, the eyes subequai, the posterior row lightly procurved,
the medians obliquely elongate-oval, subcontiguous, and
about an eye’s diameter or slightly more from the smaller
lateral eyes; median eye-area long, slightly wider in front
than behind.
Chelicera with 2 strong, equal (rarely slightly unequal)
inferior teeth (occasionally with a third minute tooth in
addition); the superior teeth 3 in number, the middle one
very strong, the proximal one small.
Legs robust; the anterior pair unspined below, the tarsus
rather short, its length being only about 2 of that of the
metatarsus ; tarsi and metatarsi of two anterior pairs densely
and broadly scopulate to the base below ; posterior tarsi more
feebly scopulate, but the metatarsi with only a few scopular
hairs; fourth tibia with a dorsal spine near base.
Sternum broad.
Vulva as in Pl. XIII. fig. 16.
Length of trunk 8-104, length and width of carapace in
largest @ 44 and 34 mm.
(b) 1 ¢ from Clanwilliam (C. L. Letpoldt, October 1897).
(c) 1 ¢ from Bosch Kloof in the Cedarberg Range,
Clanwilliam Div. (C. L. Letpoldt, December 1897).
(d) 1 ¢ from Salt River Flats, near Cape Town (W. F. P.,
April 1896).
3 .—Closeiy resembling the ? ¢, with similar chelicera,
legs, eyes, &e.
Mawille normal, without keel on outer edge of lower
surface.
Pedipalps.—Tibia cylindrical, strongly spined at apex and
provided with a small external diverging spur, which is
tipped with a sharp, curved, black tooth, its length (exclusive
ot the spur) subequal to that of the patella; tarsus narrow-
ovate, acuminate, small, about as long as the patella and half
the tibia (exclusive of the spur), its inner side with several
spines ; palpal organ with a long slender spine commencing
near base and extending along under surface to apex (Pl. XIII.
ji DE)
Length of trunk 6? mm.
The abdomen of the ¢ has a small scutellum in front.
312 Dr. W. F. Purcell on
Genus Scoropuzus, E. Sim.
Scotopheus relegatus, sp. n.
(Pl. XIII. figs. 18 & 18a.)
Specimens.—(a) 1 g from Cape Town (EF. A. Morris),
Colour.—Legs and carapace pale ochraceous ; the head and
chelicera testaceous yellow; abdomen yellowish, darker
posteriorly, with large scutum in front.
Carapace narrowed in front, the width of the head across
posterior line of eyes being more than 4 but considerably less
than 2 of the greatest width of the carapace; head much
depressed. Anterior row of eyes procurved, the medians
largest, their lower margins, seen from in front, in a line with
the centres of the laterals, the length of the clypeus equal to
that of a lateral eye, the median eyes only narrowly separated
from the laterals. Posterior row of eyes lightly procurved,
the median eyes slightly oval, almost rotund, small, being
only slightly larger than the lateral eyes, their distance apart
a little less than an eye’s diameter and their distance from
the laterals equal to nearly 14 diameters.
Chelicera much attenuated at apex, the upper margin with
one large and a couple of small teeth and the lower margin
with one small tooth.
Maville rather long, dilated externally at apex, the outer
margin being emarginate.
Sternum rather narrow, strongly attenuated in front and
produced.
Legs stoutish and rather short, the tarsi and anterior meta-
tarsi scopulate to the base, the posterior metatarsi scopulate
at apex only; anterior metatarsus with a pair of basal spines ;
first tibia with 8 inner and 2 outer spines below and some
distal scopular hairs on inner side.
Pedipalps.—Patella slightly longer than the tibia (exclu-
sive of the process), the two segments together almost equal
in length to the small tarsus; lateral process of tibia small,
terminating in a sharp incurved claw; palpal organ with
black spiral-shaped distal spine and a red hook-like process
on outer side next to the spine (Pl. XIII. figs. 18 & 18a).
Length of trunk 8} mm.
(b) 1 g from Robben Island in Table Bay (A. Tucker),
measuring only 5 mm. in length.
There are also a number of females in the Collection appa-
rently belonging to several species.
new South-African Spiders. 313
Genus DIAPHRACTUS*, nov.
Carapace longish ovate, depressed, broad in front, with
thoracic stria. Anterior row of eyes almost straight, the eyes
a little separated from one another, the medians largest.
Posterior row of eyes considerably wider and slightly pro-
curved, the medians subrotund, small. Lateral eyes on each
side a little nearer together than the anterior and posterior
median eyes. Chelicera strong, somewhat attenuated at apex,
the oblique superior margin with three small teeth remote
from one another ; no inferior teeth present. Labium elon-
gate, narrow, and parallel-sided, only ‘slightly attenuated,
emarginate at apex, and reaching almost up to the inner
angles of the maxille, the lateral margins narrowly keeled,
the surface depressed between the keels. Maxille broad,
strongly depressed, slightly dilated externally at apex and
emarginate behind the dilation, the base attenuated, the inner
margin straight, the outer margin strongly convex in posterior
two-thirds, the posterior three-fourths of maxilla bordered
along inner, posterior, and outer margins by a strong con-
tinuous keel. Sternum long-ovate, strongly attenuated in
front and produced. Legs robust, short, the posterior pairs
numerously, the anterior pairs more sparsely spined.
Apparently closely allied to Scotopheus, EK. Sim., but
resembling a Clubiona in appearance.
Type: D. Letpoldti, sp. n.
Diaphractus Leipoldti, sp.n. (Pl. XIII. fig. 19.)
1 ? from Rondegat, near Clanwilliam (C. LZ. Ledpoldt).
Colour.—Carapace testaceous yellow, the head in front and
the chelicera reddish ; legs pale yellow, the distal seoments
of the anterior pairs reddish; sternum and mouth-parts
reddish testaceous below ; abdomen pale yellowish, with
brownish hairs.
Carapace.—W idth of head across posterior line of eyes at
least 2 of the greatest width of the carapace. Clypeus slightly
less than a lateral eye in length ; posterior eyes of equal size,
small, the medians about a diameter and a half apart and
quite two diameters from the lateral eyes.
Legs.—All the tarsi and the anterior metatarsi scopulate
to the base; metatarsus I with 1-2 basal spines; tibia I
stout, with tiny scopula on each side at apex, and with 2 short
outer and 2-3 short inner spines below.
* Hedged round, alluding to the maxille.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 21
314 Dr. W. F. Purcell on
Vulva (Pl. XI1II. fig. 19) with broad median keel, a cavity
(bordered externally by a curved ridge) on each side of the
keel, and a large receptaculum partly on outer side of and
partly below each cavity.
Length of trunk 11} mm.
Genus XEROPH AUS, nov.
Carapace moderately convex, attenuated in front, with
thoracic stria. Anterior row of eyes strongly procurved, the
median eyes large. Posterior row of eyes wider, moderately
or strongly procurved, the median eyes generally large,
obliquely oval or angular and rather close together. Lateral
eyes of each side much closer together than the anterior and
posterior median eyes. Chelicera hardly or not attenuated
at apex, with three (sometimes two) superior teeth and one
inferior tooth, the latter absent in some species. Sternum as
in Drassodes, rather broad in front, or, at any rate, not
strongly attenuated nor produced. Legs generally rather
short and robust, the anterior pairs spined below on the meta-
tarsi and tibie ; tarsi and anterior metatarsi scopulate to the
base, the posterior metatarsi and often also the anterior tibia
scopulate distally. Abdomen of male scutate above at base.
‘Type: X. capensis, sp. n.
This genus is evidently closely allied to Scotopheus, which
differs from it, according to Simon’s diagnosis, principally in
having the sternum strongly attenuated in front. In Scoto-
pheus, also, both rows of eyes are apparently straighter, the
chelicera more strongly attenuated at apex and the head more
depressed.
The species fall into two sections :—
§ 1. Speeies in which the median groove of the vulva of the female
terminates anteriorly in a small transverse pocket. (Spp. 1-9.)
1. Xeropheus capensis, sp. n.
(Pl. XIII. figs. 20 & 20a.)
Specimens.—(a) 11 6 g and 7 2 @ from the northern
base of Devil’s Peak, Cape Peninsula ; also 2 2 2 from the
northern base of ‘lable Mountain.
Characters of a 3 specimen (type) :—
Carapace narrowed in front, its width across the posterior
row of eyes more than } but less than 3% of the greatest
width. Anterior row of eyes strongly procurved, the median
eyes close to the laterals and larger, much further from one
another, their lower margins on a level with (or slightly
new South-African Spiders. 315
higher than) the centres of the latter; clypeus very slightly
or scarcely longer than an anterior lateral eye’s diameter ;
posterior row of eyes strongly procurved, a line joining the
hind margins of the laterals passing slightly in front of the
centres of the medians; the median eyes ovate, larger than
the laterals and distant about their own length from the
latter and nearly half their own length from one another;
median eye-area longer than wide, broadest in front; lateral
eyes on each side subequal and between 3—? of a diameter
apart ; width of the posterior row of eyes slightly more than
‘ half the width of the head at the same place, the posterior
row also about half a lateral eye’s diameter wider on each
side than the anterior row.
Chelicera with 1 weak inferior and 3 superior teeth, only
the middle one being large.
Legs robust; all the tarsi and the two anterior pairs of
metatarsi scopulate to the base, the posterior scopule divided
by a broad band of sete; two posterior pairs of metatarsi
with a distal scopular band on the side; tibia I with an inner
row of 2 spines below; metatarsus I with a pair of basal
spines.
Pedipalps.—Tibia(Pl. XIII. fig. 20) measured along upper
inner edge subequal to the patella in length, its outer side
produced at apex into a long, stout, curved process, which is
closely appressed to the tarsus along its whole length with the
exception of the apex; this process very broad at base, then
suddenly narrowed, the narrow portion long, straight, slightly
constricted above and below near apex and ending in a black
curved claw. Tarsus large, broadly ovate, acuminate, its
length equal to that of the tibia (inclusive of the process), its
inner margin with wide shallow sinus. Palpal organ very
large and compact, occupying the whole width of the under-
side of the tarsus excepting at the apex, its lower surface
divided longitudinally by a dark ridge and provided at the
apex with two short processes.
9 ?.—Legs more densely scopulate, the first tibia with a
short, internal, distal scopula (almost obsolete in the ¢).
Vulva consisting of a dark, convex, hairy, nearly rotund
plate (slightly longer than wide), divided longitudinally by a
deep groove, which is bordered on each side by a low, smooth,
black ridge, the ridges united anteriorly in front by a trans-
verse bridge spanning the anterior end of the groove and
ending separately behind in a rounded black convexity; the
edge of the bridge (which is visible only when dry, as in
PI. XIII. fig. 20 a) is situated on a level with the anterior
edge of the rotund plate, and beneath it the groove is con-
21*
316 Dr. W. F. Purcell on
tinued anteriorly for a very short distance, forming a tiny
brown pocket, which is truncated or rounded at the anterior
end and distinctly visible in spirits through the integument.
Length of trunk, 2 ? 83-113, ¢ ¢ 6-85 mm.
In the maturer specimens of both sexes the carapace is
testaceous yellow and thickly clothed with silky yellowish
hairs and some scattered fine black bristles ; the chelicera are
testaceous and the legs yellowish, with the distal segments
testaceous; the abdomen is densely clothed with silky, light
or dark, somewhat bronzy brown hairs.
The first tibia has 2-3 spines in the inferior row, and the °
posterior median eyes are often a little more than an eye’s
length distant from the laterals, especially in the ? ¢.
(b) 2¢ 4 andd @ @ from Stellenbosch (Dr. Rk. Broom).
(c) 2 6S from near Tulbagh Road Station (W. F. P.).
(d) 2 ¢ 2 from Clanwilliam (2. WM. Lightfoot and C. L.
Letpoldt).
(e) 1 2 from Boschkloof Waterfall, Cedarbergen, Clan-
william Div. (2. Pattison).
2. Xeropheus delphinurus, sp. n.
(Pl xa tie. 21)
Specimens.—(a) 6 ¢ g from the Cape Peninsula (Cape
Flats, Bergvliet, Silver Mine Stream, and Hout Bay:
Wake sP.3).
Characters of a & specimen from Bergvliet :—
Closely agreeing with the type of X. capensis, sp. n., and
also with similar palpal organ, but differing in having the
lateral process of the tibia of the pedipalps distinctly curved
just before the apical claw, and with a short but rather deep
sinus on the underside here, asin Pl. XIII. fig. 21,
The posterior median eyes are also a little closer together
and the tibia of the first leg has 2-3 inferior spines in the
inner row.
The other males are similar, but in one of them the first
tibia has an extra spine on the inner surface besides the three
in the inferior row.
Length of trunk 63-73 mm.
(b) 1 g from the Hottentots Holland Mountains, Caledon
Div. (near Gordons Bay: R. I. Lightfoot).
3. Neropheus interrogator, sp. n.
(Pl. XPV. tig.22.)
3 6 from the Cape Peninsula (Bergvliet and Signal
Hille Wyre 2)
new South-African Spiders. a17
Characters of a & specimen from Bergvliet :—
Closely agreeing with the type of X. capensis and with
similar palpal organ, but differmg in having the lateral
process of the tibia of the pedipalp strongly curved near apex,
much more strongly so than in X. delphinurus, sp. n., and
with a much larger inferior sinus (Pl. XIV. fig. 22).
In the other ¢ ¢ the first tibia has three inferior spines,
and in one case the first metatarsus has two pairs of basal
spines.
The anterior row of eyes are also slightly less procurved
in this species than in the two preceding species, the lower
edges of the medians being slightly below the level of the
centres of the laterals.
Length of trunk 8 mm.
In the Museum are also 7 ? 2 from the Cape Peninsula
belonging either to this or the foregoing species or to both,
and differing from the @ 2 of X. capensis in that the median
eroove of the vulva is continued anteriorly for some distance
beyond the rotund dark plate. It is noteworthy that none of
these females, nor the males of X. delphinurus and tnterro-
gator, have been found on the northern slopes of Devil’s Peak
and 'Vable Mountain, which are, on the other hand, the only
localities in the Peninsula where the males and females of
X. capensis have been met with.
4. Xeropheus flavescens, sp. un. (PI. XIV. fig. 23.)
1 ¢ from Rondegat, near Clanwilliam (C. L. Leipoldt).
Colour pale yellow, the chelicera and anterior part of
carapace reddish yellow ; the abdomen pallid, with dark hairs
and yellow scutellum.
Carapace narrower than usual; anterior median eyes large,
touching the much smaller lateral eyes, whose centres are, if
anything, a trifle above the level of the lower margins of the
medians, the clypeus subequal to a lateral eye in length ;
posterior median eyes a little larger than the laterals, about 4
of a long diameter apart and about a diameter distant from
the lateral eyes.
Legs longish, the first tibia with an inner row of two spines.
Chelicera with 1 strong and 2 feeble superior teeth, the
inferior tooth feeble or obsolete.
Pedipalps.—Patella decidedly longer than the tibia, the
two segments together shorter than the narrow tarsus ; lateral
process of tibia long and narrow, bent somewhat downwards
from the base, but otherwise straight, reaching nearly to end
of tarsal organ and minutely inturned at the pointed apex,
318 Dr. W. F. Purcell on
very similar to that of X. crustosus, sp. n., but straighter ;
tarsal organ with the greater part of its underside white and
membranous, with a long, outcurved, very fine filament
arising from a small dark mesial sclerite, the inner basal
angle produced into a short horn (Pl. XIV. fig. 23).
Length 8} mm.
5. Neropheus spiralifer, sp. n.
(Pl. XIV. figs. 24 & 25.)
Specimens.—2 $ 3 and 5 ? 3 from Hanover, anda ¢
and ? from Hierfontein, 8-9 miles west of Hanover (S. C.
Cronwright Schreiner).
3 3 (types).—Colour like that of X. capensis, sp. n.
Carapace shaped as in X. capensis. Anterior row of eyes
very strongly procurved, the medians large, very close to the
laterals, and with their inferior margins a little above the line
joining the centres of the latter; clypeus equal to or very
slightly longer than a lateral eye; posterior row of eyes only
slightly wider than the anterior row, strongly procurved, a
line joining the hind margins of the laterals cutting the
medians in front of their centres, the median eyes very large,
ovate, separated by not more than } of their long diameter
from one another and by about a diameter from the laterals ;
anterior and posterior lateral eyes slightly less than a poste-
rior eye’s diameter apart.
Chelicera and legs as in capensis, but with fewer spines,
the first metatarsus having 1-2 basal spines and the first
tibia only 1 inferior spine.
Fedipalps.—Tibia, viewed from above, turbinate, much
broader distally than long, its length, measured along inner
upper edge, a little less than that of the patella; its distal
part strongly produced laterally on outer side, the process
very thick and strong, truncated at apex and bearing on its
anterior side a slenderer acuminate process, which is directed
forwards almost at right angles to the other and ends in a
claw curving downwards ; tarsus large, strongly acuminate
and incurved distally, its length very much greater than that
of the patella and tibia together (inclusive of the process),
the distal portion projecting for more than 4 of the whole
length beyond the cavity containing the palpal organ ; palpal
organ complicated, bearing two long spines, viz. a white one
arising from the middle of the outer edge, thence running in
a slight curve forwards, and ending just before reaching the
apex of the tarsus, and an extremely long red spine, which
starts at anterior end, and, after describing a spiral curve and
new South-African Spiders. 319
a half, runs backwards to the base of the tarsus and then
curves forwards again, running along the outer edge of the
tarsus alongside of the white spine, and ending together with
the latter. In spirits the underside appears as in Pl. XIV.
fig. 24.
? ¢.—Tibia and metatarsus of first leg often unspined
below, the tibia with conspicuous internal scopula.
Vulva appearing im spirits as in Pl. XIV. fig. 25, and
consisting of four dark convexities, joined in pairs on each
side by a curved deep black ridge and separated by a large
longitudinal groove which terminates in front in a small
pocket.
Length of trunk, ¢ ¢ 73-82, 2 @ 7-104 mm.
6. Xeropheus aridus, sp.n. (Pl. XIV. fig. 26.)
1 2 from Tsabis in Bushmanland, 20 miles north-east of
Concordia, Namaqualand Div. (J. H. C. Krapohl).
Closely allied to X. spiralfer, sp. n., but larger.
Carapace similar, but clothed with pale pubescence ; ante-
rior row of eyes strongly procurved, the clypeus exceeding
the lateral eyes in length ; posterior row of eyes also strongly
procurved, a line joining the posterior margins of the laterals
only just cutting the larger median eyes, the latter eyes
about + of a long diameter apart, and, if anything, slightly
more than a diameter from the lateral eyes. i
Legs.—Tibia I scopulate on both sides distally, with one
inner apical spine.
Chelicera with 3 superior but no inferior teeth.
Vulva (Pl. XIV. fig. 26) with two deep angular lateral
cavities, the anterior pocket situated more posteriorly than in
X. sptralifer.
Length of trunk 12 mm.
7. Xeropheus lunulifer, sp. n.
(Pl. XIV. figs. 27 & 28.)
Spectmens.—9 3 3 and13 ? ? from Signal Hill and the
northern and western slopes of Table Mountain and Devil’s
Peak in the Cape Peninsula (/. Treleaven, W. F. P.).
3 3 (types). Colour of dark specimens.—Carapace maho-
gany-red, with yellow pubescence ; chelicera dark red; legs
pale testaceous yellow; abdomen with dark brown or nearly
black hairs above, the underside paler.
Carapace.— W idth of head across the posterior line of eyes
almost or quite 2 of the greatest width of the carapace.
3
Anterior median eyes larger than the laterals and separated
320 Dr. W. F. Purcell on
from them, their lower margins, if anything, a trifle below
the level of the centres of the latter, the clypeus considerably
longer than a lateral eye’s diameter; posterior row of eyes
considerably wider than the anterior row and moderately
procurved, a line joining the hind margins of the laterals
passing slightly in front of the centres of the medians; the
latter oval, a little larger than the laterals, their distance
apart about 3 a long diameter (more or less), their distance
from the laterals slightly exceeding a long diameter ; the
lateral eyes on each side about a diameter or slightly less
apart.
Chelicera with 3 well-developed teeth on the superior
margin, the middle one being very large, the inferior margin
with a strong tooth.
Legs with the scopule and spines as in X. capensis, sp. n.,
but the first tibia with only 1-2 inferior spines and occa-
sionally with an internal spine in addition, but no basal spine.
Pedipalps longish ; the tibia cylindrical, longish, its length
(exclusive of the process) a little exceeding that of the
patella, its upper surface with a strong mesial spine, its lateral
process slender, acuminate, straight or slightly upturned
distally when viewed from the side, but slightly curved when
viewed from below and simply pointed; tarsus small and
narrow, acuminate, the apex obtuse, palpal organ narrower
than the tarsus, appearing in spirits as in Pl. XIV. fig. 27,
the apical spine very slender, out-turned.
2 ¢ Byes more spaced out than is generally the case in
the male, the posterior medians often separated by a space
only slightly less than a long diameter and generally about
11-1} diameters distant from a lateral eye, the posterior
medians also often scarcely larger than the laterals.
First tidza with the distal scopula distinct and occasionaliy
with three inferior spines below in the distal half, the basal
spine always absent.
Vulva appearing in spirits as in Pl. XIV. fig. 28; the
posterior part with a pair of blackish somewhat reniform
convexities, each with an intensely black cavity on the medial
side, and separated by a long median keel, which is strongly
convex from behind to in front and sharply marked off on
each side; in front of the anterior end of the keel is a small
pecket opening posteriorly and visible through the intecu-
ment as a brown half-moon.
Length, ¢ 3 93-114, 2 2 84-143 mm.
new South-African Spiders. 321
8. NXeropheus spoliator, sp. n.
(Pl. XIV. figs. 29 & 30.)
Specimens.—2 gg and 2 ¢ 2 from Hanover (S. C.
Cronwright Schreiner).
Allied to X. lunulifer, sp.n., the darker specimens similarly
coloured.
3 S (types).—Carapace and eyes much as in the ¢ of
lunulifer, except that the clypeus is shorter and may be
subequal to the length of a lateral eye.
Chelicera with 3 (sometimes 4) superior teeth, of which
the proximal one is minute; the inferior margin without
teeth.
Legs with the scopule much as in X. capensis, sp. n.;
metatarsus I with a pair of basal spines; tibia I with 3 (in
one leg with 5) pairs of inferior spines.
Pedipalps shortish ; tibia (exclusive of the process) sub-
equal in length to the short patella, produced at apex on
outer side into a stout, dark red, simple process, which hardly
equals the rest of the joint in length; tarsus very like that of
X. lunulifer ; palpal organ simple, narrower than the tarsus,
the apical spine short and stoutish, directed inwards and
upwards (P]. XIV. fig. 29).
2 ¢ .—Legs. ‘Tibia I with small scopula on outer as well
as inner side below and with a single inferior row of 3 spines.
Eyes more spaced than in the ¢, the posterior medians at
least + of a long diameter apart and nearly or quite 2
diameters from the laterals, which may be distant a diameter
or more from the anterior laterals.
Vulva appearing in spirits as in Pl. XIV. fig. 30; the two
oblique cavities (lighter in the figure) are separated by a
broad keel, which is grooved along the median line and
diverges posteriorly ; in front of the keel is another cavity
terminated anteriorly by the brown pocket.
Length of trunk, ¢ ¢ 9-11, 2? 9 15 mm.
9, Xeropheus Lightfooti, sp. n.
(Pl. XIV. figs. 31 & 32.)
Specimens.—1 g andl ¢ from Triangle, Worcester Div.
(Rk. M. Lightfoot, May 1898).
Closely allied to X. spoliator, sp.n., and similarly coloured.
& .—Carapace narrower in front, the width of head across
the posterior line of eyes less than 2 of the greatest width of
the carapace. Anterior median eyes close to the much
O22 Dr. W. F. Purcell on
smaller laterals, their lower margins on a level with the
centres of the latter, the clypeus longer than an eye’s
diameter. Posterior row of eyes strongly procurved, the
median eyes distinctly larger than the laterals and nearly 3 a
long diameter apart, their distance from the laterals also
decidedly exceeding their own length.
Chelicera with only 2 distinct teeth in upper margin, that
corresponding to the proximal tooth of the other species
being quite obsolete, the lower margin without teeth.
Legs scopulated much as in X. capensis, sp. n.; meta-
tarsus I with two basal spines ; tibia I with an inferior. row
of 3 spines.
Pedipalps longer than in spoliator, the tibia attenuated at
base, its dorsal side a little longer than the patella and angu-
larly produced at apex, the angle ending in a short slightly
out-turned spur ; outer distal edge of tibia obliquely truncated
and provided with a small, blunt, subcylindrical process,
which curves slightly upwards and inwards at apex; tarsus
rather broadly ovate, acuminate at apex; palpal organ very
like that of spoliator but larger, the apical spine hook-like,
strongly curving inwards and upwards (Pl. XIV. fig. 31).
2? —Chelicera sometimes with a minute proximal third
tooth.
Tibia I with small internal scopula and an interior row of
2-3 inferior spines.
Vulva-plate appearing in spirits as in Pl. XIV. fig. 32, as
broad as long, allied in form to that of spoliator, but smaller,
the pair of cavities (light in the figure) separated by a
broader keel, which is shallowly grooved longitudinally, its
edges diverging anteriorly as well as posteriorly, each cavity
with a large dark-coloured convexity behind it, the median
groove of the keel deepened anteriorly but not posteriorly and
not reaching hind end of the vulva-plate.
Length of trunk, ¢ 2, 9 mm.
§ 2, Species in which the longitudinal groove of the vulva of the female is
provided anteriorly with a flexible tongue-like appendage instead of a
pocket. (Spp. 10-16.)
10. Xeropheus communis, sp. n.
(Pl. XIV. figs. 33, 33a, & 34.)
Specimens.—(a) 1 g and 3 ? 2 from the Willowmore
District (H. H. L. Schwarz).
3 (type).—Very like X. capensis in form, colour, and in
the characters of the eyes, legs, and chelicera.
new South-African Spiders. 323
Posterior median eyes about a third of a long diameter
apart, the clypeus slightly less than a lateral eye’s diameter.
Legs.—Tibia I with 1 outer and 1-2 inner spines below.
Fedipalps.—Patella very slightly longer than the tibia
(exclusive of the process), the two segments together equalling
the tarsus in‘ length; tibia (Pl. XIV. fig. 33 a) oval, cylin-
drical, provided at its outer distal margin with a strong
spiniform process, which is only slightly shorter than the
tibia itself, the apex of the process pointed and minutely
incurved but not upcurved; palpal organ as in Pl. XIV.
fig. 33, the apex with a short, fine, curved spine, the apex of
the black sclerite at the inner basal angle produced upwards
into a minute process.
? ¢.—Tibia I scopulate on both sides distally, but espe-
cially on the inner side, the under surface with an inner row
of 2 spines.
Vulva (Pl. XIV. fig. 34) with a large median groove, con-
taining in its anterior part a large tongue-shaped appendage,
the convexity on each side of the posterior half of the groove
bordered externally by a curved, black, comma-shaped mark
and furrow, the anterior part of the groove flanked on each
side by a small depression, the apex of the appendage
marked with a small brown spot.
Length of trunk, ¢ 8, 2 2? 83-113 mm.
(b) 1 g from the Baths near Montagu (W. F. P.).
(ec) 1 g andl ¢ from Touws River Station, Worcester
Diva (We.
(d) 1 g from Laingsburg, Prince Albert Div. (W. F. P.).
(e) 2 ¢ g from Prince Albert Village (W. F. P.).
(f) 2 ¢ 3 and5d 2 ¢ from Hanover and neighbourhood
(Vlagkop, Eierfontein: S. C. Oronwright Schreiner).
(g) 1 $ from Dunbrody (Rev. J. A. O'Neil).
(h) 1 g and3 2 @? from the Kentani District, Transkei
(Rev. F. C, Kolbe, Miss A. Pegler, and H. P. Abernethy).
(i) 1 $ from Rhode, Mount Ayliff Distr., Pondoland
(A. Weisbecker).
Also many other specimens, apparently females of this
species, from various parts of Cape Celony.
This is the commonest and most widely distributed species
in Cape Colony. In the ¢ ¢ the first tibia has 1 outer and
usually 2 (rarely 1 or 4) inner spines below, including an
apical pair, but in the ? there is no outer apical spine below.
The length of the clypeus sometimes equals the length of an
anterior lateral eye, and the posterior median eyes are some-
times as much as half an eye’s diameter apart. The apical
324 Dr. W. F. Purcell on
spine of the palpal organ is frequently much straighter than
in the type.
11. Xeropheus aurariarum, sp. n.
(Pl. XIV. figs. 35 & 36.)
Specimens.—4 3 S (types) and 3 ¢ 9 from Witwaters-
rand, Transvaal (Miss A. Pegler); also several ? 9 from
Johannesburg (H. A. Fry) and Krugersdorp, Transvaal.
Closely resembling X. communis in general characters,
except :—
Length of clypeus exceeding that of an anterior lateral eye,
the lower margins of the median eyes also slightly above the
level of the centres of the laterals.
Tibia of first leg with 1 outer and 1 inner apical spine
(rarely with 2 inner spines below) in the g, but with only
1 inferior spine in the ¢.
Pedipalps of g short, the patella a little longer than the
tibia (exclusive of the process), the two segments together
distinctly shorter than the tarsus; lateral process of tibia
resembling that of X. communis in shape, but subequal to
the rest of the joint in length; palpal organ as in Pl. XIV.
fig. 35.
Walve of ? as in Pl. XIV. fig. 36, the receptaculum
seminis on each side large, embracing the comma-shaped
black grooves.
Length of trunk, ¢ g 7-83, 2 2 103-123 mm.
12. Xeropheus exiguus, sp. n.
(Pl. XIV. figs. 37 & 37a.)
1 gS from Laingsburg, Prince Albert Div. (2. A. Light-
foot, August 1903).
Closely resembling X. communis, sp. n., but smaller,
Pedipalps.—Tibia (exclusive of the process) subequal to
the patella in length, produced on its outer side into a stout
acuminate process, which is slightly upturned at apex and is
as long as the rest of the joint (PI. XIV. fig. 37 a) ; tarsus
ovate, acuminate, longer than the patella and tibia (exclusive
of the process); palpal organ very like that of communis
(PL XV gig. 37).
Length of trunk 63 mm.
13. Xeropheus rostratus, sp. n.
(Pl. XIV. figs. 38, 38a, & 39.)
Specimens.—1 fg (type) and 1 ? from Graaff Reinet
(J. Paynter, September 1902).
new South-African Spiders. 325
Closely resembling X. communis, sp. n., in characters of
carapace, eyes, spine-armature, and scopulee of legs, Ke.
Length of elypeus subequal to the length of an anterior
lateral eye.
Legs rather longish, the first tibia in the ¢ with a few
inner scopular hairs.
Chelicera with a strong inferior tooth.
Pedipalps of $ longer than in X. communis, the patella
as long as the tibia (exclusive of the process), but the two
segments together longer than the tarsus; lateral process of
tibia short and stout, the apex suddenly narrowed and claw-
like, strongly curved inwards and a little downwards
(Pl. XIV. fig. 38) ; palpal organ larger and more extensively
chitinized than in communis, the apical spine very short
(EE XIV fig. 38 a).
Vulva of 2 nearly but not quite like that of communis,
differing mainly in the form of the receptaculum seminis
(ERUXIVE fies 39).
Length of trunk, ¢ 103, 2 12 mm.
14. Xeropheus crustosus, sp.n. (Pl. XV. fig. 40.)
1 from East London (J. Wood).
Allied to X. communis, sp. n., but larger.
Carapace mahogany-red, the head narrow ; anterior median
eyes with their lower margins situated slightly below the
level of the centres of the lateral eyes, the clypeus a little
longer than a lateral eye; posterior row of eyes much wider
than the anterior row, the median eyes equal to the laterals
in size and at least ? of a long diameter apart, distant from
the laterals about 1} times a long diameter.
Chelicera with weak lower tooth, but strong upper teeth.
Legs very robust, the first tibia with 0-1 outer and 2 inner
spines below.
Pedipalps.—Tibia (viewed from above) attenuated at base,
its length (exclusive of the process) slightly less than that of
the patella, the lateral process resembling that of X. com-
munis in shape, but very long, much longer than the rest of
the joint, and reaching as far as the distal end of the palpal
organ, its distal part slightly curving upwards, the apex
pointed and minutely incurved; palpal organ very like that
of X. communis, the apical spine short (Pl. XV. fig. 40).
Length of trunk 11 mm.
326 Dr. W. F. Purcell on
15. Xeropheus appendiculatus, sp. n.
(Pl. XV. figs. 41 & 42.)
Specimens.—(a) 3 $ § and 1 ? from Hanover (C. S.
Cronwright Schreiner).
3 3 (types).—Colour of carapace light testaceous yellow,
the legs slightly paler; the chelicera blackish red; the
abdomen clothed with brown hairs.
Carapace.—Width of head across posterior median eyes
almost or quite # of the greatest width of the carapace.
Anterior row of eyes strongly procurved, the lower margins
of the median eyes slightly above the level of the centres of
the laterals, the clypeus long, almost twice as long as a
lateral eye; posterior row of eyes much wider than the
anterior row, moderately procurved, a line joining the hind
margins of the laterals cutting the medians a little in front
of their centres ; the posterior median eyes oval, much larger
than the laterals, distant from the latter about 1} times a
long diameter or slightly more, but only } of a diameter from
one another ; lateral eyes on each side separated by a little
more than a posterior eye’s diameter.
Chelicera with the inferior tooth rather small.
Legs longish, the first tibia conspicuously scopulated on
each side distally, the posterior metatarsus strongly scopulate
on each side; metatarsus I with 2 basal spines; tibia I with
an inferior row of 2-3 spines.
Pedipalps.—Tibia (exclusive of the process) short, equal
to the patella in length, its shape turbinate when seen from
above, its lateral process very long, as long as the patella and
tibia together, closely appressed to the tarsus except at the
apex, slender, pointed, appearing perfectly straight when seen
from the side (except at base, where it is slightly bent and
suddenly and strongly thickened); tarsus long, ovate, acumi-
nate; palpal organ rather complex, provided in the middle at
apex with a short, curved, black spine or spur and a long,
extremely fine filament, and on the inner side at the base
with a stout, curved, obtuse process (Pl. XV. fig. 41).
2 yes slightly more spaced, the clypeus 1} times the
length of an anterior lateral eye, the median eyes a little
removed from the laterals, the posterior median eyes distant
at least 14 times a long diameter from the posterior laterals.
Legs.—Metatarsus | with 1-2 basal spines.
Vulva (Pl. XV. fig. 42) with a couple of large lateral
depressions, which are bordered on the inner side by a pair of
sharp black edges converging posteriorly, each depression
containing a large brown convexity posteriorly ; anterior
new South-African Spiders. 327
part of vulva-plate with a median, flexible, tongue-like
appendage lying in a groove.
Length of trunk, ¢ ¢ 9-92, 9 12 mm.
(b) 1 g and 1 2? from Zululand (W. Anderson). The
& does not apparently differ from the type, but in the ? the
appendage of the vulva reaches further back relatively to the
lateral cavities.
16. Xeropheus Patricki, sp.n. (Pl. XV. fig. 43.)
1 ¢ from the Punegwe, 50 miles east of Umtali, in Portu-
guese Hast Africa (DV). L. Patrick).
Colour, torm of carapace, and the ocular area resembling
that of X. communis, except that the clypeus is distinctly
longer than an anterior lateral eye; the posterior median
eyes oval, large, much larger than the laterals, and 4 of a
long diameter apart.
Chelicera with the inferior tooth very small.
Legs \ongish, slender; metatarsus I with 0-1 basal spine ;
tibia I sparsely scopulate on both sides distally, the underside
with an inner row of 3-4 spines.
Pedipalps.—Tibia (exclusive of the process) slightly
longer than the patella, its lateral process resembling that of
X. communis in shape, but straighter, almost as long as the
rest of the joint, its apex pointed and minutely incurved, not
quite reaching to the middle of the tarsus; palpal organ
somewhat like that of X. appendiculatus, with similar internal
process, but with only a short apical spine, which is directed
outwards and downwards (Pl. XV. fig. 43).
Length of trunk 83 mm.
Genus MELANOPHORA, C. Koch.
1. Melanophora fuliginea. (Pl. XV. figs. 44-46.)
Specimens.—(a) 7 2 2 from Signal Hill, Cape Town, and
several ¢ ¢ from the same locality. Females of this species
have not been found in other parts of the Peninsula.
9 2 (types).—Colour black, the legs brownish distally
and frequently also at base, the first pair of femora with
large pale yellowish area on both inner and outer surfaces ;
pulmonary opercula pale yellowish or brownish, the post-
genital portion of the abdomen often more or less pale
yellowish.
Anterior row of eyes strongly procurved, the lateral eyes
considerably larger than the medians, the length of the clypeus
328 Dr. W. F. Purcell on
a little exceeding that of a lateral eye; posterior row of eyes
almost straight (scarcely recurved), a little wider than the
anterior row, the median eyes flattish and appearing (in
spirits), if anything, slightly smaller than the convex lateral
eyes, their distance apart also slightly less than or subequal
to their distance from the laterals.
Chelicera normally with 1 large and 3 small superior and
2 small but distinct inferior teeth on the inner margin.
Legs.—Anterior tarsi and metatarsi scopulate below, the
first metatarsus with 2 inferior basal spines, the tibia unspined.
Vulva as in Pl. XV. fig. 44.
3 S. Pedipalps.—Patella a little longer than the tibia
(exclusive of the lateral process); tibia with simple straight
lateral process, which is slightly longer than the rest of the
segment; tarsus oval, acuminate, slightly shorter than the
patella and tibia, together with the lateral process; tarsal
organ as in Pl. XV. fig. 45, the distal spine curved, strong,
with a smaller spine on its side at some distance from the
apex (fig. 46), the apex itself not projecting out of the cavity
of the tarsus.
Length of trunk, 2 ?, 62-9 mm.
(b) 2 9 2 and 1 ¢ from Slanghoek, Worcester Div.
CW ELE.)
2. Melanophora invida, sp.n. (Pl. XV. fig. 47.)
Specimens.—(a) 9 2 ¢ from various parts of the Cape
Peninsula (Cape Flats, Camps Bay, Hout Bay, Kalk Bay
Mountain) and 1 @? from Maitland Flats, Cape Div.*
Vulva as in Pl. XV. fig. 47, being very wide and practically
filling the space between the lung-opercula.
Other characters as in J. fuliginea, sp. n., but the inferior
teeth of the chelicera very minute and sometimes apparently
obsolete.
Length of trunk, 2? ?, 6-74 mm.
(b) 1 2 from Stellenbosch (Dr. &. Broom).
(c) 1 2 from near Bethlehem, Stellenbosch Div. (G.
French).
(d) 1 2 from Ceres (W. F. P.).
(e) 2 2 2 from Rabiesberg, Worcester Div. (W. F. P.).
(f) 1 2 from Touws River, Worcester Div. (W. F. P.).
* Tam unable to identify the males of this and the following Penin-
sular forms. Some of these males are apparently scarcely distinguishable
from those of M. fuliginea, sp. n.
/
new South-African Spiders. 329
3. Melanophora Lightfoot’, sp.n. (PI. XV. fig. 48.)
2 2 2 from Ceres (R. M. Lightfoot).
Vulva as in Pl. XV. fig. 48.
Closely allied to MZ. envida, sp. n. Chelicera with 2-3
distinct inferior teeth.
Length of trunk 63-84 mm.
4, Melanophora Simoni, sp.n. (PI. XV. fig. 49.)
Spcimens.— (a) 12 2 2 from various parts of the Cape
Peninsula (Cape Flats, Camps Bay, Hout Bay, Kalk Bay
and ‘l’able Mountains), including a specimen from Signal
Hill.
Vulva as in Pl. XV. fig. 49.
Closely resembling M. fuliginea, sp. n., but smaller and
often less black. Posterior median eyes more convex and
more sharply defined.
Length of trunk, 9 9, 54-64 mm.
(b) 1 2 from Grahamstown (ev. J. A. O'Neil).
5. Melanophora redunca, sp.n. (Pl. XV. fig. 50.)
Specimens.—(a) 5 9 2 from the Cape Peninsula (Cape
Flats, Kalk Bay Mountain).
Vulva as in PI]. XV. fig. 50.
Other characters as in WM. Simont, sp.n. Chelicera with
2-3 minute inferior teeth.
Length of trunk, 9 ¢, 53-63 mm.
(b) 1 ¢ from Brandvlei, Worcester Div. (W. F. P.).
(c) 1 2 from Touws River, Worcester Div. (W. F. P.).
(a) 1 @ from Avontuur, near Stormsvlei, Swellendam
Div. (W. F. P).
6. Melanophora montana, sp.n. (Pl. XV. fig. 51.)
Specimens.— (a) 3 2 2? from the Cape Peninsula (Table
Mountain, &c.).
Vulva as in Pl, XV. fig. 51.
Other characters as in J/, fuliginea, sp. n. Chelicera with
2~3 inferior teeth.
Length of trunk 6?-83 mm.
(b) 1 2 from Port Elizabeth (J. L. Drége).
7. Melanophora Broom, sp.n. (Pl. XV. fig. 52.)
Specimens.—(a) 1 2 from Stellenbosch (Dr. R. Broom).
Vulva as in Pl, XV. fig. 52.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 22
330 Dr. W. F. Purcell on
Other characters as in M/. Simoni, sp. n.
Length of trunk 5 mm.
(b) 1 2 from Bergvliet, Cape Peninsula (W. F. P.).
(c) 2 99 from St. Helena Bay, Malmesbury Div.
(J. H. C. Goold).
(d) 2 2 2 from Porterville Road (formerly Piquetberg
Road) Station, Tulbagh Div. (R. MW. Lightfoot).
(e) 1 9 from Tulbagh Road (W. F. P.).
8. Melanophora Gooldi, sp.n. (PI. XV. fig. 53.)
3 2 ¢ from Stompneus and Steenbergs Cove, St. Helena
Bay, Malmesbury Div. (J. 7. C. Goold).
Vulva as in Pl. XV. fig. 53.
Other characters as in I, fuliginea, sp. n.
Length of trunk 63-8} mm.
9. Melanophora Cronwrighti, sp. n.
(Pl. XV. figs. 54 & 55.)
Specimens.—(a) 19 9 9 and 4 ¢ ¢ from Hanover, and
1 9 and2 ¢ 8 from Vlagkop, 5-6 miles north of Hanover,
all collected by Mr. 8. C. Cronwright Schreiner.
Very closely allied to 1. Goold’, sp. n., but the 2 2 with
somewhat different vulva (Pl. XV. fig. 54).
Chelicera with 1-2 inferior and 3-4 superior teeth on inner
margin.
Pedipalps of 3 .—Patella slightly longer than tibia (exclu-
sive of the lateral process) ; lateral process of tibia longer
than the rest of the joint ; tarsus as long as the patella and
tibia together with the lateral process; tarsal organ very like
that of Md. fuliginea, sp. n., but with the distal spine
different (Pl. XV. fig. 55).
Length of trunk, 9 2 5-83, ¢ 3 42-6} mm.
(b) 3 2 2 from Worcester (2. AL. Lightfoot).
10. Melanophora caldaria, sp. n.
(Pl. XV. figs. 56 & 57.)
Specimens.—2 9 ¢ and 1 g from the Hot Baths at
Montagu, collected by my wife and myself.
2 2 (types).— Colour as in the preceding forms.
Eyes.—Anterior row strongly procurved, the lateral eyes
considerably larger than the medians; posterior row straight,
scarcely or not at all wider than the anterior row, the eyes
equidistant, the laterals larger than the medians.
new South-African Spiders. 331
Chelicera with 4 superior but apparently only 1 inferior
tooth on inner margin.
Two anterior pairs of legs without any spines below.
Vulva as in Pl. XV. fig. 56.
3. Pedipalps.—Tibia short, its entire length (inclusive of
the lateral process) being only about equal to that of the
patella, the process forming about } the whole length, the
tarsus considerably longer than these two segments together ;
palpal organ with a long, fine, curved, black spine on outer
side distally (Pl. XV. fig. 57).
Length of trunk, 9 9 54-52, ¢ 43 mm.
11. Melanophora zonognathus, sp. n.
(Pl. XV. figs. 58 & 59.)
Specimens.—4 3 9 and 4 ¢ g from Hierfontein, 8-9 miles
west of Hanover (S. C. Cronwright Schreiner).
2 9 .—Abdomen black; carapace and legs black to dark
brown; underside and tarsi paler; the anterior pair of
femora dark, without the large pale areas on outer and inner
surfaces,
Eyes.—Anterior row procurved, the lateral eyes large, much
larger than the small medians; posterior row wider, straight,
the eyes equidistant, the laterals larger than the medians.
Labium and mawille longish, the latter deeply emarginate
on outer side.
Chelicera with 1 large and several smaller superior and
2-3 well-developed inferior teeth on inner margin.
Legs.—Metatarsus I unspined, Il with a pair of basal
spines below.
Vulva as in Pl. XV. fig. 58.
3 g. Pedipalps.—Tibia (including lateral process) as
long as the patella, the process small, shorter than the rest
of the joint; tarsus longer than the two preceding joints
together ; palpal organ with long slender black spine arising
distally on outer side and curving backwards and then
forwards on lateral side (Pl. XV. fig.
@., 59).
Length of trunk, ¢ 2? 53-64, dd 43-
a]
64 mm.
12. Melanophora humilis, sp. n.
(Pl. XV. figs. 60 & 61.)
Specimens.—1 @ (type) and 1 g from Ceres (W. F& P.,
October 1897).
Infuscated, the underside and the tarsi paler, the anterior
pair of femora with large pale yellowish area on each side,
332 Dr. W. F. Purcell on
Eyes as in M. conognathus, sp. 0.
Chelicera with 2 inferior teeth.
Legs.—Anterior metatarsus with a pair of inferior spines.
Vulva of 2 asin Pl. XV. fig. 60.
Pedipalps of &.—Tibia (exclusive of the process) slightly
shorter than the patella, but inclusive of the process longer,
the process shorter than the rest of the segment; tarsus
shorter than the tibia ‘and patella together; palpal organ as
in Pl. XY. fig. 61.
Length, 2? ¢, 24 mm.
13. Melanophora O'Neili, sp.n. (Pl. XV. fig. 62.)
Specimens.—2 ¢ 2 from Dunbrody, Uitenhage Div.
(Rev. J. A. O'Neil).
Colour.—Carapace and chelicera mahogany-brown to
blackish red, the legs only slightly paler than the carapace.
Abdomen infuscated or black, pallid below.
JZyes.—Anuterior row procurved, the lateral eyes decidedly
larger than the medians; posterior row almost straight (a
trifle procurved), the medians smaller than the laterals and
slightly nearer to the latter than to one another.
Labium and maxille as in zonognathus, sp. n.
Chelicera with 3 large superior but no inferior teeth.
Legs.—Metatarsi I and Il unspined.
Vulva as in Pl. XV. fig. 62.
Length of trunk 54-7 mm.
14. Melanophora corrugata, sp. n.
(Pl. XV. figs. 63 & 64.)
Specimens.—(a) 16 9 2? and 2 g¢ from Hanover and
1 o and 3 @ @ from Vlagkop and Hiertontein, near Hanover
(S. C. Cronwright Schreiner).
2 2. Colour.—Carapace and sternum light testaceous, the
legs paler except the patella and tibia of I and the tibia of IT,
which are generally darker or more or less infuscated, the
fourth tibia and metatarsus also often darkened. Abdomen
more or less infuscated above, pale below, the scutellum of the
& testaceous.
Eyes rather large and conspicuous, the anterior row
strongly procurved, the laterals largest; the posterior row
wider than the anterior row and very slightly procurved, the
medians oblique, very close together, a little larger than the
laterals and much nearer to one another than to the latter ;
clypeus slightly shorter than the Jength of an anterior lateral
eye.
new South-African Spiders. 333
Mazille broad at base, obtuse at apex, emarginate exter-
nally.
Chelicera with about 3 minute teeth on superior inner
margin, but none on lower margin.
Legs stout, especially the femur, patella, and tibia of first
pair; tibia I ‘and LI unspined below ; ; metatarsus I unspined,
II with 1 (rarely 0) outer and 2 inner, stout, short spines.
Vulva as in Pl. XV. fig. 63, with a large, transversely
striated, anterior area.
3 go. Legs.—Tibie I and Il with 3-4 outer and 2-4 inner
splues ane II with 1-3 upper spines as well; metatarsi [
and II with 2 inferior pairs of spines, IIL with an inner
superior spine as well.
Pedipalps.—Tibia together with its large process much
longer than the patella; palpal organ as in Pl. XV. fig. 64,
with a long slender black spine, which arises anteriorly and
coils round the upper and outer side and ends on the lower
surface.
ee of trunk, 2? ? 48-8, ¢ ¢ 55 mm.
(b) 1 ? from era in British Bechuanaland (Gia Ei
Beare).
Very distinct from any of the species described above.
15. Melanophora arida, sp.n. (Pl. XV. fig. 65.)
Specimens —1 @¢ and 1 juv. from Laingsburg, Prince
Albert Div., collected by my wife and myself.
Very closely allied to the foregoing, but with the abdomen
pallid above, the posterior row of eyes not wider than the
anterior row, the posterior lateral eyes being nearer the
medians (but further from them than the latter are from one
another), the chelicera with a minute inferior tooth and 4—5
minute superior teeth on inner margin; tibia IL with or
without an inferior spine, and the vulva as in Pl. XV.
fig. 65.
Length of trunk in 2 55 mm.
16. Melanophora acanthognathus, sp. n.
(Pl. XV. fig. 66.)
Specimens.—7 ¢ 2? from Cape Town and the Cape
Peninsula.
Colour testaceous, the legs sometimes somewhat darkened,
the sternum bordered with black ; the abdomen blackened,
its ventral surface pallid, or, at least, paler.
Anterior row of eyes strongly procurved, the laterals larger
than the medians, the clypeus longer than a lateral eye ;
334 Dr. W. F. Purcell on
posterior row straight (scarcely procurved), slightly or
scarcely at all wider than the anterior row; the eyes large,
the medians triangular, larger than the laterals and slightly
nearer to one another than to the laterals.
Chelicera with some superior but no inferior teeth on inner
margin; the anterior surface densely covered with numerous
longish spines.
Mazille obtuse, slightly emarginate externally, the labium
shortish.
Legs.—Tibix I and II with 0-1 inferior spine; metatarsi
T and II with two long bands of slenderer spines and generally
also a basal pair of stouter spines below ; tarsi I and II also
furnished with two rows of small spines below and scopulate
on each side.
Vulva as in Pl. XV. fig. 66.
Length 7-83 mm.
EXPLANATION OF THE PLATES.
PLATE XIII.
Fig. 1. Platyoides bidentatus, sp.n., 9. Vulva.
Fig. 2. Ditto, ¢. Tibia, tarsus, and palpal organ of left pedipalp from
below.
Fig. 3. Platyoides quinquedentatus, sp.n., 2. Vulva.
Fig. 4. Drassodes lophognathus, sp. u., 5. Tibia and tarsus of right
pedipalp from above.
Fig. 5. Ditto, g. Right palpal organ removed from the tarsus and
seen obliquely from the medial side and below.
Fig. 6. Ditto, 9. Vulva.
Fig. 7. Drassodes tessellatus, sp.n., 9. Vulva.
Fig. 8. Ditto, ¢. Apex of process of right palpal organ in same
position as in fig. 5.
Fig. 9. Drassodes solitarius, sp.n., 2. Vulva.
tg. 10. Drassodes caffrerianus, sp. u.. 3. Distal part of right palpal
organ in same position as in fig. 5.
Fig. 11. Drassodes calceatus, sp.n., 9. Vulva.
Fig. 12: Drassodes Gooldi, sp.n., 2. Vulva.
Fig. 13. Drassodes lyratus, sp.n. Q. Vulva.
Fig. 14. Drassodes helene, sp. u., 6. Right pedipalp from below.
14a, Distal process of palpal organ in same position as in
fig. 5, to show the dorsal tooth.
Fig. 15. Drassodes Dreget, sp. n., Q. Vulva.
Fig. 16. Drassodes ereptor, sp.u.. Q. Wulva.
Fig. 17. Ditto, g. Right pedipalp from below.
Fig. 18. Scotopheus relegatus, sp.n., ¢. Tibia of right pedipalp from
outer side. 18a. Tarsus and palpal organ of right pedipalp
from below.
Fig. 19. Diaphractus Letpoldti, sp. n.. 9. Vulva.
Fig. 20. Xeropheus capensis, sp.n., §. Right pedipalp from outer side.
20a. 2, vulva.
Fig. 21. Xeropheus dephinurus, sp.n., $. Tibia of right pedipalp from
outer side.
Fig.
Fig.
Fig. 2
Fig. 2
Fig. 2
Fig. :
Fig. 2
Fig. *
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fug.
Fig.
. 42.
. Xeropheus Patricki, sp. n.. g. Tarsus and palpal organ of
new South-African Spiders. 335°
' Prate XIV.
. Xeropheus interrogator, sp. n., o. Tibia of right pedipalp
from outer side.
. Xeropheus flavescens, sp... 3. Right pedipalp from below.
. Xeropheus spiralifer, sp. n., 6. Right pedipalp from below.
. Ditto, 9. Vulva.
. Xeropheus aridus, sp.u., 9. Vulva.
. Xeropheus lunulifer, sp. n., ¢. Right pedipalp from below.
. Ditto, 9. Vulva.
. Xeropheus spoliator, sp. n., 6. Right pedipalp from outer side.
. Ditto, 2. Vulva.
. Xeropheus Lightfooti, sp. u.. do. Right pedipalp from outer
side.
. Ditto, 2. Vulva. :
. Xeropheus communis,sp.n., o. Tarsus and palpal organ of right
pedipalp from below. 33a. Tibia of right pedipalp from outer
side.
. Ditto, 9. Vulva.
5. Xeropheus aurariarum, sp.n.. 3. Tarsus and palpal organ of
right pedipalp from below.
. Ditto, 2. Vulva.
. Xeropheus exiguus, sp.n.. &. Tarsus and palpal organ of right
pedipalp from below. 37a. Tibia of right pedipalp from
outer side.
. Xeropheus rostratus, sp. n., d. Tibia of right pedipalp from
outer side. 88a. Tarsus and palpal organ of right pedipalp
from below.
. Ditto, 2. Vulva.
PLATE XV.
. Aeropheus crustosus, sp.n., ¢. Tarsus and palpal organ of right
pedipalp from below.
. Xeropheus appendiculatus, sp. nu. S. Right pedipalp from
below.
Ditto, 2. Vulva.
right pedipalp from below.
. Melanophora fuliginea, sp. n., Q. Vulva.
. Ditto, ¢. Tarsus and palpal organ of right pedipalp from
below.
. Ditto, ¢. Distal part of right palpal organ from outer side
with the tarsus removed.
. Melanophora invida, sp. n., 9. Vulva.
. Melanophora Lightfooti, sp.n., 9. Vulva.
. Melanophora Simoni, sp.n. Q. Vulva.
. Melanophora redunca, sp.n., 9. Vulva.
. Melanophora montana, sp.n., 2. Vulva.
. Melanophora Broom, sp.n., Q. Vulva.
. Melanophora Gooldi, sp.u., 2. Vulva.
. Melanophor'a Cronwrighti, sp. n., 9. Vulva.
. Ditto, g. Right palpal organ from outer side, removed from
the tarsus,
. Melanophora caldaria, sp.n., 2. Vulva.
. Ditto, ¢. Right palpal organ from outer side, removed from
the tarsus.
336 Mr. G. A. Boulenger on new
Fig. 58. Melanophora zonognathus, sp. n., 2. Vulva.
Fig. 59. Ditto, g. Tarsus and palpal organ of right pedipalp from
below.
Fig. 60. Melanophora humilis, sp.n., 9. Vulva. }
Fig. 61. Ditto, g. Tarsus and palpal organ of right pedipalp from
below.
Fig. 62, Melanophora O'Neili, sp.u., 2. Vulva.
Fig. 63. Melanophora corrugata, sp.n., Q. Vulva.
Fig. 64. Ditto, 6. Right pedipalp from below.
Fig. 65. Melanophora arida, sp.n., Q. Vulva.
Fig. 66. Melanophora acanthognathus, sp.n., Q. Vulva.
XLIU.—Descriptions of Two new African Species of Barbus.
By G. A. BouULENGER, F.R.S.
THE paradoxical section of Barbels without barbels (Systomus,
McClelland, Bleeker), long believed to be restricted to South-
eastern Asia, has, within the present century, received several
additions from Africa. ‘I'wo further species are here described,
bringing the number of African species up to eight, which
may be distinguished by means of the following key :—
A. Lateral line complete.
Sc. 29-32 77; depth of body 3} to 33 times in
total length ; caudal peduncle wwice as long [1906 (L. Victoria).
as deep is005 ROG GoW SORTA AE Se ooo ar B. Magdalene, Bier.,
Sc. 28 ae depth of body 3 times in total length ;
caudal peduncle once and 3} as long as [1902 (Conzo).
GEGD eet Mhax hcick sisices hoaistrome ale teres ian B. Brazze, Pellegr.,
Se. 25 3 depth of body 23 times in total [(S. Cameroon).
length ; caudal peduncle as long as deep.. B. aspilus, sp. n.
Se. 22-25 a; depth of body 22 to 53 times
in total length ; caudal peduncle not or but [ (Nile),
slightly longer than deep .............. B. anema, Blgr., 1903
Sc. 23 ai depth of body 3} times in total
length; caudal peduncle once and 3 as long [(Congo P),
SIL CCP era ererevettvele aie rersrem siclegs cle eh eee B, trispilomimus, sp. n.
B. Lateral line absent or reduced to a few tubules; depth of body
equal or nearly equal to length of head, 3 to 53 times in total
length.
Sc. 22-25 ae origin of dorsal midway be- [1908 (Nile).
tween end of snout and root of caudal.... B. stigmatopygus, Blgr.,
Sc. 20-23 Et origin of dorsal nearer end of [(S. Cameroon),
Anoutthan caudal: voc. cee een B. Je, Bigy., 1903
4
Se. 19-20 a ; origin of dorsal midway between [ (Nile).
end of snout and root of caudal.......... B. pumilus, Blgr., 1901
African Species of Barbus. 337
Barbus aspilus.
Depth of body 24 times in total length, length of head 4
times. Snout rounded, a little shorter than the eye, the
diameter of which is 3 times in length of head and equals
interorbital width ; mouth small, subinferior, with moderately
developed lips; no barbels. Dorsal III 8; last simple ray
not ossified, once and 4 as long as head; the border of the
fin slightly emarginate, its origi nearer end of snout than
root of caudal. Anal III 5. Pectoral a little shorter than
head, reaching ventral; latter a little behind vertical of
origin of dorsal. Caudal forked. Caudal peduncle nearly as
deep as long. Scales 25 es 24 between lateral line and
ventral, 12 round caudal peduncle. Yellowish brown above,
the scales dark brown at the base, silvery below; no
markings.
Total length 125 mm.
A single specimen from the Ja River, 8. Cameroon (Coll.
G. L. Bates).
Barbus trispilomimus.
Depth of body 3} times in total length, length of head 3
times. Snout rounded, projecting beyond the mouth, shorter
than the eye, the diameter of which is 2? times in length of
head and equals interorbital width; mouth small, with thin
lips; no barbels. Dorsal JI 8, last simple ray not ossified,
as long as head, the border of the fin not emarginate, its
origin midway between end of snout and root of caudal.
Anal IlI 5. Pectoral shorter than head, not reaching
ventral; latter below middle of dorsal. Caudal forked.
Caudal peduncle once and a half as long as deep. Scales
23 a 2% between lateral line and ventral, 8 round caudal
peduncle. Pale brownish above, silvery below ; three large
round black spots on each side of the body, on the lateral
line, the first in front of the vertical of the dorsal, the second
behind the vertical of the dorsal, the third in front of the
caudal; two small black spots at the base of the dorsal.
Total length 35 mm.
The locality of this fish, which strikingly resembles
B. trispilus, Blkr., in form and markings, is unknown. The
single specimen was found in an unlabelled jar containing
examples of Clariallabes melas, Blgr., and Channallabes apus,
Gthr. It therefore probably comes from the Congo.
338 On a new Pit-Viper from Brazil.
XLIV.—Deseription of a new Pit-Viper from Brazil.
By G. A. BouLenGer, F.R.S.
Lachesis ttapetininge.
Snout obtusely pointed, with strong, slightly raised canthus.
Rostral a little deeper than broad; nasal divided; upper
head-scales small, imbricate, strongly keeled; supraocular
large, separated from its fellow by 7 to 9 longitudinal series
of scales; internasals large and in contact with each other ;
a large canthal; two or three postoculars and a subocular,
which is separated from the labials by one series of scales ;
loreal pit separated from the labials; temporal scales keeled ;
8 upper labials, third and fourth largest. Scales strongly
keeled, in 25 rows. Ventrals 150-152; anal entire; sub-
caudals 28-29. Pale reddish brown above, with large
transversely oval or quadrangular dark light-edged spots
disposed in two alternating series, some of the spots of the
two sides meeting on the middle line of the back, others, on
the sides, broken up into two or more; sides of snout
whitish ; a broad oblique dark band below the eye; a trans-
versely oval dark spot on the upper surface of the snout, and
a wavy dark band, continuous or interrupted, on each side of
the top of the head from the interocular region to the nape ;
belly white, much speckled or blotched with brown, the white
of the lower parts sometimes sharply defined from the brown
of the upper, on the lower row of scales.
Total length 400 mm. ; tail 55.
Two specimens, females, from Jtapetininga, a city in the
State of Sio Paulo. The British Museum is indebted for
these specimens to Dr. Vital Brasil, Director of the Serum-
therapic Institute of S. Paulo, who recognized them as
belonging to a distinct species, remarkable for its small size,
the above-described specimens being the largest obtained so
far. The name Lachesis ttapetininge was suggested to me
by Dr. Vital Brasil.
This Lachests itapetininge is nearly related to L. Neuwiedit,
Wagl., from which it is easily distinguished by its stouter
form, as expressed by the lower number of ventral and caudal
shields.
Among other specimens of Lachesis from the State of
Sio Paulo I was pleased to examine one of the “ Jararacucu,”
a remarkable variety of LZ. lanceolatus growing to a length of
over 2 metres, the pattern of coloration of which has been
figured by Jan (Icon. Ophid. xlvii. pl. ii. fig. 3). It agrees
entirely in structure with the true “ Fer-de-Lance,” with
which it is completely connected by intermediate patterns of
coloration.
On new Species of Uisteridee. 339
XLV.— On new Species of Histeridee and Notices of others.
By G. Lewis, F.L.S.
MARSEUL, in his Monograph, collated descriptions of species
other than his own for the sake of easy reference, and I have
from time to time followed his example. In this, the
thirty-second paper of the series, 1 have introduced a few
more.
List of Species.
Teretriosoma argentinum.
Eutidium peruanum.
Teretrius montanus, Horn.
levatus, Horn.
Sternaulax caledonixw, Fauv.
Apobletes angolensis, Lev,
Migneauxi, Mars.,
=foliaceus, Payk.
Platysoma connexum, Fauv.
Simeani, Muls. § God.
Silinus, gen. nov.
Zabromorphus deflexus.
Hister subsulcatus, Mars.,
=hottentotta, Hr.
Hister falsus, Sols.
Bruchi.
foveicollis.
Grammostethus socius.
Atholus atricolor.
Kpitoxus felix.
Microlomalus filum, Reiét.
Pachycrerus montanus, Lew.
nanus.
Paratropus aptistrius.
Homalopygus latisternus,
Murexus, gen. nov.
Saprinus sparsutus, Sols.
lateristrias, Sols,
turanus, Sods.
Teretriosoma argentinum, sp. n.
Cylindricum, subelongatum, obscure seneum, undique sat dense
punctulatum ; antennis pedibusque rufo-brunneis ; elytris trans-
versim basi impressis; mesosterno stria marginali prave integra ;
tibiis anticis 7-denticulatis.
L. 27 mill.
Cylindrical, somewhat elongate, faintly coppery, clearly
but not very closely punctured above; the head sometimes
has a small fovea on its vertex which may be sexual; the
thorax, the lateral stria continues behind the head; the
elytra are impressed transversely behind the anterior angles ;
the pygidium is slightly convex and with the propygidium is
clearly punctulate like the upper surface; the sterna are
microscopically strigose, with punctures evenly and not
closely set; the prosternal keel is arched at the base and
anteriorly the marginal rim is very narrow; the mesosternum
is obtusely produced anteriorly and the marginal stria is
apparently complete but irregular, owing to the interposition
of punctures; the metasternum has a well-marked lateral
stria which turns outwards from the anterior edge; the legs
340 Mr. G. Lewis on
and antenne are reddish brown, anterior tibia with seven
denticulations.
This species is more cylindrical and elongate than any
other known, There are now nearly forty species of this
genus described, and others exist in collections, and they are
very difficult to characterize intelligently.
flab. Argentina, Province of Buenos Aires (£7. Richter).
Eutidium peruanum, sp. n.
Oblongo-ovatum, convexiusculum, nigrum, nitidum; fronte plana
haud striata; pronoto impunctato; elytris striis 1-2 brevibus,
ceteris nullis; pygidio grosse punctato, apice transversim levi.
L. 6? mill. (absque mandibulis).
Oblong-oval, slightly convex, black and shining; the
forehead is not striate, but in certain lights transverse
impressions may be seen which apparently correspond to
striz; the thorax is laterally impunctate; the elytra, stria
1-2 are basal and very short, the outer one longest; the
propygidium is sparsely punctate on its lateral edges and
rather widely biimpressed apically ; the pygidium is coarsely
punctured, with the posterior margin smooth; the prosternal
keel is narrowed before the coxe.
The large size, the more oblong form, and the coarse
punctuation of the pygidium distinguish this species from the
other four known.
Hab. Peru.
Teretrius | Teretriosoma?] montanus, Horn, Tr. Amer. Ent.
Soe. vill. p. 143 (1880).
“ Black, shining. Thorax a little wider than long, rather
finely punctate, the punctures denser in front. Elytra not
more densely punctured than the posterior portion of the
thorax. Prosternum sparsely punctate, the tip distinctly
grooved. Mesosternum sparsely and finely punctate, without
marginal line. Anterior tibize rather strongly bidentate at
middle, the posterior bispinose near the tip.
“ Length *10 inch (2°5 mm.).
“ Hab. Colorado.
“Of the same form as americanus [Teretriosoma], but
larger and with different dentate tibiz.”
Teretrius levatus, Horn, Proc. Calif. Ac. iv. p. 365 (1894).
“Cylindrical, piceous black, shining; legs brown. Head
finely, not closely punctate. ‘Thorax sparsely punctate, fine
new Species of Uisteride. 341
at apex and sides, coarser near base. Elytra more coarsely
punctate, the punctures finer near sides than on apex; a
smooth umbonal space; a short very oblique stria at base.
Pygidium sparsely punctate. Mesosternum distinctly mar-
gined in front, sparsely and coarsely punctured. Anterior
tibiee 4-denticulate; middle tibize 4-spinose ; hind tibie bi-
spinose near apex, with a smaller spine at middle.
“* Length *10 inch (2°5 mm.).
« Resembles obliquulus, but much smaller [the measure-
ment given is ‘10 inch for both species], and with one less
spine on the middle and hind tibia, the spines being at the
same time more slender.
“ Hab, San José del Cabo, Lower California.”
Sternaulax caledonic, Fauv. Rev. d’Ent. x. p. 164 (1891).
Fauvel in his description of this species points out the
differences between it and zealandica, Mars. :—
Voisin du zealandica, Mars., mais distinct surtout par
Ja disposition du stries; celle de la téte, presque complete en
avant; celle du corselet plus forte et entiére jusqu’au milieu ;
la 2° dorsale des élytres trés nette, bien que raccourcie ; le
propygidium plus finement ponctué,- entouré dun rebord
cariniforme.””
The anterior femora of Marseul’s species are also more
markedly punctate.
L. 9 mill.
fab, New Caledonia.
Apobletes angolensis, Lew. Ent. Mag. xvi. p. 76 (1879).
Oblongo-ovalis, complanatus, nigro-piceus, nitidus; fronte plana,
stria fere recta; pronoto stria marginali antice late interrupta,
interna tenuiter impressa, obliqua, margine distanti; elytris
striis 1-3 integris, ceteris nullis ; propygidio transversim punc-
tato; pygidio grosse punctato, margine postice elevato; prosterno
haud striato; mesosterno stria integra, utrinque valde ‘impressa.
L, 5-6 mill.
Hab. Angola, Congo River, and Togoland.
Apobletes Migneauxi, Mars. (1860),=A. foliaceus, Payk.
(1811), syn. n.
The species I referred to in a note (Ann. & Mag. Nat.
Hist. vu. p. 241, 1901) with a complete mesosternal stria is
undescribed at present, as I do not know its habitat.
342 Mr. G. Lewis on
Platysoma connerum, Fauv. Rev. d’ Ent. x. p. 166 (1891).
« Forma plana, striis elytrorum structuraque pygidii ut in atisterno,
Mars., conformatis, sed pronoti lateribus minus distincte punctu-
latis, stria laterali marginali conjuncta, hac margine apicali haud
interrupta, prosterno minus lato, mesosterno toto marginato,
antice arcuatim profundius sinuato, sine linea inter hoc et meta-
sternum bene perspicua sat facile distinguendum.”
L. 4 mill.
Hab. New Caledonia.
I copy Fauvel’s description of the above, as I apparently
erroneously placed P. connexwum as a synonym of P. Mont-
rouzieri, Mars., in the Catalogue of 1905.
Platysoma Simeani, Mulsant & Godart, Ann. Soc. Linn.
Lyon, xxi. p. 420 (1874).
“Oblong, noir, brillant; antennes et pattes brundtres ;
strie frontale entiére ; prothorax quadrangulaire ; élytres a
trois stries marginales entiéres ; les trois suivantes raccourcies ;
pygidium couvert de gros points ocelles ; tous les tarses
[tibiee] tridentés.”
Mulsant also says that the prosternum is bordered with a
stria, but this refers to the anterior lobe, not to the keel.
The anterior tibiz are 3-dentate, but the others are 4-spinose.
The head is finely punctulate anteriorly and somewhat
coarsely punctured behind, the frontal stria is strong and
widely transverse in front, not quite straight, and continues
well-marked over the eye; the lateral stria is very similar to
that of Confuci?, Mars., but it continues a little round the
posterior angle and behind the neck it widens a little away
from the edge and is feebly and irregularly crenate, surface
lightly punctulate, especially laterally ; the pygidia are
closely ocellately punctured; the mesosternum is rather
widely sinuous anteriorly, but the marginal stria is bisinuous
and fine and not very close to the edge, and it is broken at
the suture, not joining the metasternal lateral stria. The
elytra, as Mulsant says, have the 1-3 dorsal complete, 4-5
and sutural not quite dimidiate, the fourth and fifth (the
shortest) are rather shorter than the sutural. The outer
humeral stria is fine and complete, and continues along the
apical margin to a point opposite the sutural stria.
The form of the humeral stria will serve to identify this
species from others known.
Hab. Beirut, Syria, “ sous l’écorce d’un olivier.”
new Species of Histeride. 343
SILINUS, gen. nov.
Body elongate, somewhat cylindrical, but a little depressed ;
the head impressed, frontal stria complete, mandibles more or
less canaliculate at the base, antennal fossa circular and open ;
thoracic marginal stria complete and partly or wholly con-
tinued along the base; elytral strie 1-4 complete, 5 dimidiate,
sutural wanting or very faintly marked; propygidium punc-
tate; pygidium smooth at the apex or marginate ; prosternum
sometimes striate; mesosternum emarginate, stria complete ;
all the tibize dilated.
The species known have hitherto been placed in Platylister,
and are pinnigera (type), palmipes, and ewtrarius, Lew., and
robustus, Sch. The frontal stria in the last is very deep and
the thoracic stria is simply “complete,” not, in the usual
sense of this term, passing along the base. In other respects
extrarius and robustus are, judging by the descriptions, very
similar.
Zabromorphus deflezus, sp. n.
Ovalis, convexus, niger, nitidus; pronoto striis 2 lateralibus in-
tegris ; elytris striis dorsalibus 1-2 integris, 3 dimidiata, ceteris
nullis ; propygidio pygidioque dense punctatis ; mesosterno
truncato, marginato.
L. 8-83 mill.
Oval, convex, black and shining, surface very finely
punctulate; the head, frontal stria complete and_ nearly
straight anteriorly ; the thorax, inner lateral stria complete
and continues behind the head and is markedly deflected near
the eye, outer lateral ceases after passing the anterior angle,
behind the angle the outer interspace has some iregular
marks which seem to be the rudiments of a third lateral stria,
these marks vary in individuals; the elytra, inner humeral
apical dimidiate, somewhat faint and broken at intervals, 1-2
dorsal complete, 3 basal but reaching the middle and it is
somewhat irregular and lightly impressed, the other dorsal
strie are wanting, but there are impressions and punctures
along the apical margin which apparently represent ap-
pendages; the pygidia are densely punctured; the meso-
sternum is truncate and the marginal stria complete ; the
anterior tibize are 3-dentate and the anterior femora clearly
punctured.
The peculiar dorsal striation will serve to distinguish this
species.
Hab. Isansu, German East Africa.
344 Mr. G. Lewis on
Ilister subsulcatus, Mars. (1854),= 7H. hottentotta, Er.
(1834), syn. n.
Lister turanus, Sols. Reise Fedsch. Turkest.,
Col. ii. p. 223 (1876).
*‘ Subquadrato-ovatus, convexus, niger, nitidus, antennis piceis ;
fronte biimpressa, stria integra, antice recta; pronoto transverso,
antrorsum parum angustato, utrinque breviter ciliato et bistriato,
striis integris; elytris striis dorsalibus 1-3 integris, 4 ee
abbreviata, haud appendiculata ; margine inflexo impresso
bistriato, levi; propygidio basi disperse, minus subtiliter punc-
tato, apice levi, pygidio sat crebre subtilius punctato; prosterno
apice rotundato; mesosterno lato, sat profunde sinuato, stria
integra; tibiis anticis valide 3-dentatis, dente apicali bilobo,
posticis biseriatim 5—6-denticulato-spinosis.
6c Ip. D2 ”
ese
TIab. In valle Sarafschan, specimen unicum.”
The above is Solsky’s diagnosis, and there is more in the
Russian language descriptive of the species.
Hister falsus, Sols. Reise Fedsch. Turkest.,
Col. ii. p. 229 (1876).
“ Oblongus, subquadrato-ovatus, convexus, niger, nitidus, supra
obsolete, subtiliter punctulatus, antennis rufo-brunneis, basi
nigricantibus. Stria frontali antice recta, medio subsinuata ;
mandibulis supra plana, subtiliter rugulosis. Pronoto antrorsum
parum angustato, stria laterali interna integra, externa ante
medium abbreviata. Elytris thorace longioribus, stria sub-
humerali nulla, 1-3 dorsalibus integris, levibus, 4-6 abbreviatis,
apicalibus, 5 fere nulla, 6 dimidiata; fossa marginali levi bi-
striata. Propygidio pygidioque minus crebre punctatis, illic
apice levigato. Mesosterno apice emarginato, stria integra.
Tibiis anticis quadridentatis, dente apicali bilobo, posticis 7—8-
spinoso-denticulatis.
“ Var. striis 4 et 5 dorsalibus omino deletis.
* Long. 6, lat. 44 mill.”
“Ad Maracandam.” ‘Turkestan.
Fister Bruchi, sp. n.
Oyatus, convexus, niger, nitidus; fronte haud impressa, stria antice
recta; pronoto stria interna integra, e externa brevi; elytris striis
1-3 integris, 4-5 apicalibus, suturali medium versus abbreviata ;
pygidio tenuissime punctulate ; mesosterno marginato, antice
sinuato; tibiis anticis 5-dentatis.
neu Species of Uisteride, 345
Oval, convex, black and shining; the head very finely
punctulate, stria complete and straight anteriorly, forehead
not impressed; the thorax, inner lateral stria complete, outer
short and almost confined to the angle; the elytra, stria,
outer humeral short and just before the middle, inner well-
marked apical and slightly overlaps the outer stria, 1-3
complete, 4—5 very short and apical, sutural is much short-
ened anteriorly ; the pygidia are very finely punctulate (like
those of cavifrons, Mars.) ; the mesesternum is slightly
sinuous and marginate anteriorly; the anterior tibiz are
5-dentate.
This species belongs to a section of the genus peculiarly
American.
Hab. Argentina, Province Salta and Tucuman (C. Bruch
and G. A. Baer).
Fister foveicollis, sp. n.
Ovyalis, convexiusculus, niger, nitidus; fronte plana, stria semi-
circulari; pronoto in medio bifoveolato; elytris striis 1-4
dorsalibus integris, 5 et suturali abbreviatis; propygidio bifoveo-
lato pygidioque punctulatis.
L. 53-6 mill.
Oval, slightly oblong, somewhat convex, black and shining ;
the forehead plane, not impressed, stria complete and semi-
circular, surface very finely punctulate; the thorax also very
finely punctulate, with two discal fovee separated by about
the width of the head, inner lateral stria complete, outer
dimidiate or sometimes longer; the elytra, striz, inner
humeral shortened one third at the base, dorsal 1-4 complete,
5 nearly dimidiate, sutural one third longer; the propygidium
and pygidium very finely punctulate, the first has two shallow
foverw near the outer edge; the mesosternum emarginate,
stria complete ; the anterior tibiae 5—6-dentate.
This species is rather more oblong than H. punctifer, Payk.,
but it is otherwise very similar. ‘The thoracic fovew are
very peculiar, and had I only a single example I should
have considered them incidental.
Hab. Brazil, Minas, Serra de Diamantina (EZ. Gounelle) ;
three examples.
Grammostethus socius, sp. n.
Ovalis, convexiusculus, niger, nitidus; fronte minute punctulata,
stria arcuata, integra; elytris striis 1-4 integris, 5 et suturali
dimidiatis ; prosterno striis posticis abbreviatis.
L. 4 mill.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 25
346 Mr. G. Lewis on
Oval, somewhat convex, black and shining; the forehead
minutely punctulate, stria complete and rather widely arched ;
the thorax very finely punctulate, inner stria complete and
minutely crenate behind the head, outer stria wanting ; the
elytra, outer humeral stria basal, curved and dimidiate, inner
stria wanting, dorsal 1-4 complete, 5 and sutural nearly equal,
but the sutural just passes the middle; the pygidia are clearly
not closely punctured ; the prosternum, the strize are short
and do not pass between the coxe; the basal lobe is im-
pressed; the mesosternum is truncate, stria complete and
close to the edge ; the anterior tibize are 5—6-denticulate.
The fourth dorsal stria in this species is complete, but this
does not seem to be an important character in the genus, as
in some not otherwise dissimilar species (e. 9. fractistrius,
Lew. Ann. & Mag. Nat. Hist. xvii. p. 342, 1906) the fourth
stria is sometimes complete on one side and not on the other.
Hab. Kashmir, North India.
Atholus atricolor, sp. n.
Ovalis, convexiusculus, niger, nitidus ; fronte stria integra, antice
leviter biimpressa; pronoto stria externa abbreviata, interna
integra; elytris striis 1-4 validis, integris, 5 apicali, suturali
basi paululum abbreviata; propygidio aliquantum grosse punc-
tato ; tibiis anticis 4-dentatis.
L, 33-4 mill.
Oval, somewhat convex, black and shining; the head,
mandibles canaliculate, stria complete, strong at the sides,
feebly sinuous anteriorly, with two circular impressions
behind it, surface punctulate ; the thorax, outer lateral stria
short, passing from the anterior angle nearly halfway along
the side, inner stria rather strong and complete, crenate behind
the neck; the elytra, striz, inner humeral strong, apical, and
reaching just beyond the middle, dorsal 1-4 also strong and
complete, 5 apical and not quite reaching the middle, sutural
shortened a little at the base and turning slightly from the
suture at both ends; the propygidium is coarsely, evenly,
aud not densely punctate, on the pygidium the punctures are
smaller; the prosternum is narrow between the coxe; the
mesosternum, marginal stria complete and close to the anterior
edge, slightly broken at the suiure, not quite joining the
metasternal stria; the anterior tibie are 4-dentate.
Hab. Mashonaland (Guy A. K. Marshall).
A stercoraceous species which sometimes flies to light.
new Species of Histeride. 347
Epitoxus felix, sp. ns
Snborbicularis, convexus, niger, nitidus; fronte plana tenuissime
punctulata, stria semicirculari integra; pronoto lateribus punc-
tato, ante scutellum transversim rugose punctato; elytris striis
1-2 subintegris, 3 integris sinuatis, 4 fere dimidiata, 5 obsoleta,
i suturali integris basi arcuata ; tibiis anticis multispinosis.
3 mill.
Nearly orbicular in outline, convex, black and shining ;
the legs and antenne dark brown; the head, surface with an
extremely fine punctuation, stria semicircular ; >; the) thorax,
marginal stria complete and finely crenulate behind the head,
disk almost smooth, outer area rather coarsely but not densely
punctured, the scutellar impression is transverse and narrow
and somewhat rugosely punctured; the elytra, striz, outer
humeral apical and nearly dimidiate, 1 dorsal a little short-
ened apically, 2 a little shortened at the base, 3 complete,
slightly sinuous, 5 indicated only by two or three apical
punctures, sutural complete but very fine before the base,
seen only in certain lights to join an appendage to the fourth
stria; the pygidia are clearly, not densely, punctured, and
the points vary in size and are fewer near the apices; the
prosternum is sparingly and evenly punctured, stria well-
marked laterally, but very fine along the base and not quite
joining anteriorly ; the mesosternum is impunctate, marginal
and transverse striz well-marked and crenulate ; the anterior
tibiee are 9-10-spinose.
The epipieurz of the species in this genus are bistriate.
Hab. Usambara, Hast Africa.
Microlomalus (Paromalus) filum, Reitt. Deutsche ent. Zeitschr.
XXVill. p. 256 (1884).
** Prosternum antrorsum angustum, obtuse carinatum, striis laterali-
bus non impressum, postice rotundatim lobatum, subdeplanatum.
Metasternum valde elongatum, in medio maris longitudinaliter,
minus profunde sulcatum. ‘Tibiis leviter curvatis, angustis,
intermediis subtiliter tridentatis, anticis dilatatis (magis curvatis),
extus parce denticulatis, apice extus oblique truncatis,’
Hab. Bulgaria.
Pachycrerus montanus, Lewis (1907).
Oblongus, subcylindricus, viridi-cyaneus, nitidus ; fronte impressa,
stria integra; pronoto stria marginali antice late interrupta ;
elytris striis 1-4 integris, 5 obsoleta, suturali dimidiata; pro-
pygidio pygidioque fortius vix dense Punchavis 5 prosterno
348 Mr. G. Lewis on
bistriato, striis rectis haud conjunctis; mesosterno antice im-
marginato ; tibiis anticis 6-dentatis.
L. 53-6 mill.
Oblong, somewhat cylindrical, greenish blue and shining ;
the head slightly impressed anteriorly, marginal stria com-
plete, surface microscopically punctulate ; the thorax, mar-
ginal stria is widely interrupted behind the head, the disk
before the scutellum is almost smooth, but behind the head
and laterally it is conspicuously punctate ; the elytra, strie,
outer humeral fine bent and reaching the middle from the apex,
1-4 complete, 5 short and apical or wanting, sutural apical
well-marked and reaching just beyond the middle; the
pygidia are closely and somewhat coarsely punctate; the
prosternum bistriate, striz straight and not joining at either
end and do not quite reach the base; the mesosternum is
somewhat acute and immarginate anteriorly ; the anterior
tibiae: 6-dentate.
In form and colour this species somewhat resembles
P. Rafray’, Lew., but it is slightly greener and being broader
is a little less cylindrical.
Hab. Kilimanjaro (Sjéstedt). Two examples.
Pachycrerus nanus, sp. n.
Oblongus, parum convexus, subniger, nitidus, pedibus rufo-brunneis ;
fronte stria utrinque angulata, antice late interrupta; propygidio
parum grosse punctato; mesosterno stria transversa arcuata,
stria marginali late interrupta.
L, 24 mill.
Oblong, rather convex, nearly black and shining, legs and
antennee reddish brown; the head very feebly impressed
anteriorly, surface with punctures of varying sizes not closely
set, stria angulate near the eye, oblique anteriorly and
abruptly shortened ; the thorax, marginal stria fine and not
passing behind the neck, punctuation not close and consisting
of somewhat large points and fine ones intermixed; the
elytra, outer humeral stria fine and complete, inner short and
nearly median, 1-4 dorsal complete, 5 wanting, sutural a
little shortened at both ends, apices very sparingly punctured ;
the propygidium is somewhat coarsely, but not closely
punctured ; the pygidium, punctures much smaller and fewer ;
the prosternum is bistriate, strize sinuous and not joining at
either end, keel moderately broad; the mesosternum is
bisinuous anteriorly, not pointed, and the marginal stria is
widely interrupted, the transverse stria is detached and in
the form of an arc, with a few irregular notches along the
new Species of Histeride. 349
edges, it is anterior to the suture; the anterior tibia are 5-6-
denticulate.
The small stature and the singular frontal and mesosternal
striz distinguish this species from the others known.
Hab. 'Togo-land.
Paratropus aptistrius, sp. n.
Ovatus, brunneus, tenuissime punctulatus, subnitidus ; fronte stria
marginali integra, utrinque carinata ; pronoto anguste marginato ;
elytris striis 1-4 et suturali integris, 4 arcu basali suturali
coéunte, 5 ultra medium abbreviata ; prosterno angusto, bistriato.
L. 33 mill.
Oval, brown, somewhat shining, surface microscopically
punctulate; the head convex on vertex, frontal stria com-
plete, angulate before the eyes and carinate especially laterally
and by the angles; the thorax is narrowly marginate; the
elytra, striz, humeral strong and complete, 1-4 and sutural
finer and also complete, 4 and sutural join at the base, 5 is
shortened before the middle ; the pygidia are feebly punctulate;
the prosternum, the keel is narrow and bistriate, strie parallel
until just behind the anterior lobe, where they turn outwards ;
the mesosternum is sparingly punctulate, bistriate anteriorly,
with a erenate straight stria over the suture.
This species is extremely similar to ovddes, Mars., but the
fourth and sutural strize are complete and join anteriorly and
the surface is not ‘ levis nitidus,” owing to a fine though
sparing punctuation and a roughened sculpture seen under
the microscope.
flab. Portuguese Guinea.
Homalopygus latisternus, sp. n.
Oblongus, depressus, piceus, nitidus ; fronte tenuiter punctulata ;
pronoto ad angulos tristriato ; elytris striis dorsalibus abbreviatis ;
pygidio basi excavato; prosterno lato, impunctato; mesosterno
profunde inciso.
L. 24 mill.
Oblong, rather depressed, piceous, shining; the head,
surface uneven, free, finely punctulate and densely micro-
scopically strigose; the thorax, marginal stria fine and
interrupted behind the neck, the anterior angles are some-
what impressed, at the basal angle there is a fine short stria
and close to it are two longer strize also very fine and bent
and pointing towards the head; the elytra, striz very fine,
inner humeral complete, 1 dorsal indistinct, 2—8 fine, basal,
but well-marked, 4 a little longer and joined to the sutural stria,
350 On new Species of Visteridee.
which is complete and onthe dorsal disk geminate ; the pro-
pygidium is finely not closely punctured; the pygidium is
triangularly excavated at the base ; the prosternum, the keel
is rather wide (twice the width of that of remex, Lew.), and
the strie are feebly sinuous between the cox and are widely
separate at both ends; the mesosternum is very deeply
incised on either side of the projection, which is somewhat
large and obtuse, the marginal stria is fine and complete, but
does not follow the anterior edge closely.
In outline the above is similar to H. remea, Lew., but the
aeaceea dorsal strize are very different, and the excavated
pygidium and the width of the prosternal keel also distinguish
it from remee.
Hab. Jatahy, Province of Goyaz, Brazil.
MUREXUS, gen. nov.
Body oblong and with the legs densely muricate, forehead
convex ; thorax explanate laterally and anteriorly cut out in
semicircular outline; elytra costate; prosternum roughly
carinate, anterior lobe tuberculate ; mesosternum bisinuous ;
legs elongate, tibiz angulate on the outer edge. Type
Terapus muricatus, Lew. Ann. & Mag. Nat. Hist. vi. p. 286,
pl. 10. fig. 2 (1900).
The above characters are selected from those drawn for a
single species; at present only Terapus Mniszecht, Mars., and
bicarinatus, Lew., are properly referable to Terapus. T. Mar-
seult, Westw., more nearly resembles a species of aye but it
has no mesosternal tubercles : Westwood’s fioure, ‘ Thesaurus
Ent. Oxon.’ p. 67, tab. 67. f. 9 (1874), is of course excellent.
Saprinus sparsutus, Sols. Reise Fedsch. Turkest.,
Col. 1. p. 238 (1876).
Saprinus brunnensis, Fleicher, Wien. ent. Zeit. ii. p. 179 (1888).
“ Saprino tenuistrio, Mars., affinis, subquadrato-ovatus breviusculus,
nitidus, niger, elytris apice picescentibus, pedibus antennisque
piceis, his clava ferruginea. Fronte crebre punctulata, stria nulla.
Thorace medio subtilissime disperse punctato, lateribus fortius et
erebre ruguloso-punctato, longitudinaliter impresso, impressione
paulo ante angulum posteriorem fovea oblonga, margini approxi-
mata, terminata, foveolis postocularibus distinctis. Elytris
undique punctulatis, punctis circa scutellum subtilioribus, magis
dispersis, postice fortioribus, sat crebris; stria suturali integra,
basi cum 4 dorsali conjuncta, striis dorsalibus in medio abbreviatis ;
stria humerali postice abbreviata, cum subhumerali interna haud
On Pteropus mascarinus, iLuson. 351
eonjuncta, subhumerali externa breyi, basali. Pygidio sat crebre
punctato. Prosterno striis convergentibus, stria mesosternali
integra. Tibiis anticis 5-6-denticulatis..
“Long. 42, lat. 3 mill.
** In valle Sarafschan.” Turkestan.
Saprinus lateristrius, Sols. Reise Fedsch. Turkest.,
Col. ii. p. 240 (1876).
* Oblongo-ovatus, convexus, piceo-niger, nitidus, antennis pedibus-
que piceis, his rufescentibus. Capite subtiliter, minus dense
punctulato, fronte antice transversim impressa et utrinque sub-
carinata, stria frontali medio interrupta. Thorace transverso,
basi utrinque subsinuato, versus angulos anticos arcuatim
angustato, conyvexo, disco subtilissime, minus dense, versus
marginem anteriorem et latera paulo fortius et crebrius punctuata,
summa basi punctis majoribus immixtis; stria marginali
integra, interne striola, antice et postice abbreviata, margini
laterali parallela, adjecta. Elytris lateribus parum rotundatis,
apice solum versus angulum internum subtiliter, disperse punctu-
latis ; striis dorsalibus paulo pone medium abbreviatis, prima
ceteris breviore, 4-ta antice cum suturali confluente, subhumerali
interna abbreviata, cum humerali conjuncta, externa basi notata.
Pygidio crebre, profunde, minus subtiliter punctato. Prosterno
striis lateralibus antrorsum sensim approximatis. Tibiis anticis
4—5-denticulatis, dentibus tribus ultimis validis.
“Long. 3, lat. 2 mill.
“Ad urbem Taschkent.” Turkestan..
XLVI.—Some Remarks on Pteropus masearinus, Mason.
By Knup ANDERSEN.
In the September number of this Journal (pp. 220-222)
Mr. Geo. E. Mason gives a description of a supposed extinct
species of fruit-bat, Péeropus mascarinus, from the small
island La Ronde, about 15 miles north-east of Mauritius.
The species is based on a single skull found in a fissure, in a
superficial deposit of red earth, associated with bones of
introduced animals, such as goat and rabbit.
A perusal of the description and measurements of the skull
and teeth of Pt. mascarinus has satisfied me that the species
must be exceedingly like the now living Pé. rodricensis,
Dobson. ‘ 1 ;
The skull of Pt. mascarinus is described as having a
352 Mr. K. Andersen on
“ moderately long but heavy muzzle, flattened and scarcely
concave frontal region; sagittal crest weakly developed,
almost obsolete.” In Pt. rodricensis the muzzle is moderately
long but heavy ; the sagittal crest in young adults undeveloped
or almost obsolete, in aged individuals well developed but
low; the frontal region (between postorbital processes) in
young adults flattened and scarcely concave, in aged specimens
distinctly concave. The type of Pt. mascarinus is no doubt
a young or young adult specimen.
The canines in Pt. mascarinus are “long and sharp, both
above and below, with unusually acute and prominent postero-
internal basal ledges, those of the maxillary in particular”’ ;
the same is the case in Pt. rodricensis. p* in Pt. mascarinus
is “ very minute, but would have been still persistent in the
type and standing in the tooth-row ”; p® in Pt. rodricensis is
minute and deciduous; the presence of the alveoli of this
tooth in the type of Pt. mascarinus is additional evidence
that it is a young or young adult individual.
It must be admitted that the above characters of the skull
and teeth in Pt. mascarinus are rather vague; they are, in
fact, common to a large number of species of Pteropus, and
consequently do not give any conclusive evidence of the close
affinity of Pt. mascarinus to Pt. rodricensis, far less of its
possible identity with the latter species. But when we
turn from the description to the measurements of the skull
and teeth we are on much safer ground; nearly all the
measurements of Pt. mascarinus show the most striking
similarity to those of Pt. rodricensis. In the table below I
compare the principal measurements of the skull and teeth of
Pt. mascarinus, as given by Mr. Mason, with those of three
skulls of adult specimens of Pt. rodricensis (one of them the
type of the species) in the collection of the British Museum.
It will be seen that the measurements of the two species are
practically identical, with the following two exceptions, both
of which are, most probably, apparent only, not real :—
(1) The approximate basilar iength of the skull of Pt. masca-
rinus is stated to be 44 mm., the approximate condylo-basal
length 46 mm.; in Pt. rodricensis the measurements are
respectively 46 and 51:‘7 mm.; the basilar length is practi-
cally the same in the two skulls (taking into consideration that
the measurement of ma:scarinus is approximate only, and the
type probably a young adult), whereas the condylo-basal
length would seem to be widely different; but it must be
said that, provided the basilar length in mascarinus is
approximately 44 mm., the condylo-basal length cannot
Pteropus mascarinus, Jfason. 303
possibly be 46 mm., but must be considerably greater; the
latter measurement cannot have been taken by Mr. Mason in
the usual way (from condylion to gnathion), or, if it has, it
is wrong. (2) All the measurements of the teeth in the
supposed two species are exceedingly alike, with one excep-
tion; in Pt. rodricensis m, is a trifle (0°1-0°2 mm.) shorter
(antero-posterior extent) than Py in Pt. mascarinus it would
seem to be no less than 0°8 mm. longer than py; that m,, in
a species of Pteropus, is considerably longer than py, looks
sufficiently strange to arouse doubt as to the correctness of the
statement : but still more strange the character becomes from
another point of view; in all species of Pteropus the width
of m, is (very nearly) between two thirds and three fourths
the length of the tooth, in Pé. mascarinus the width would
be less than half the length of the tooth, and at the same
time all other teeth, above and below, would be practically
precisely similar to those of Pt. rodricensis! I have not
much hesitation in saying that the width of m, is probably
correctly given by Mr. Mason, but there must be some
misprint or mistake in his measurement of its length.—lf,
now, I eliminate the two points discussed above, viz. the
condylo-basal length and the length of m; in Pt. mascarinus
(and I think I am justitied in doing so), there remains a
skull which, so far as Mr. Mason’s description goes, agrees
exactly with that of Pt. rodricensis.
In discussing the probable affinities of Pt. mascarinus,
Mr. Mason comes to the conclusion that it “‘ occupies a place
intermediate between Pieropus ey [i.e. Pt. Edwardsi]
and Pt. redricenstis, in size only, the dentition being typical
of Pteropus, whereas the two above species fall into the
subgenus Spectrum”; and ‘a great analogy, in fact, exists
between the dentition of Pt. mascarinus and those species of
Pteropus (tonganus, Gouldii, and conspicillatus) inhabiting
Australia and the islands of the Pacific Ocean.” A glance
at the subjoined table of measurements is suflicient to show
that Pt. mascarinus is not intermediate in size between
Pt. Edwardsi and Pt. rodricensis, but similar to this latter
species. Whether mascarinus is a distinct species or not
cannot be decided with any degree of certainty, so long as
its external characters are unknown ; closely related species
of Pteropus often agree in cranial and dental, and differ in
external characters, and the distance between Mauritius and
Rodriguez is great enough to make it poss¢b/e that mascartnus
is distinguishable from rodricensis. But so much is beyond
all doubt, that in order to find “a great analogy ” to Pt. mas-
354 Mr. K. Andersen on
carinus we need not go so far as Australia and Polynesia,
but only the much shorter way from Mauritius to Rodri-
guez; itis with Pt. rodricensis, and this species only, that
Mr. Mason ought to have compared the “ Round Island” skull
before he regarded it as representing a new species,
Mr. Mason calls Pt. mascarinus an extinct species. On
what evidence? As already mentioned, the skull was found,
superficially in a fissure, together with bones of introduced
animals; the “state of preservation and general condition of
the remains”? show, Mr. Mason admits, that they are of
quite recent origin. What, then, are the proofs that the skull
is of an extinct species, rather than of an individual that died
last year? Is the bat fauna of “ Round Island” so well
known that we can be sure that P#. mascarinus does not exist
there at this moment? In Mr. Mason’s own words, only a
“cursory”? survey of the island was made by the party,
“interested in the guano trade,” who visited the island in
1906, and a member of which obtained the skull. Supposing,
for the sake of argument only, that, for some reason or other,
Ft. mascarinus has ceased to exist on the island, where are
the proofs that it does not live in Mauritius, which is situated
only 15 miles from “ Round Island”? It is perfectly in-
credible that Pt. mascarinus should have been confined to the
small “ Round Island,” which is not only situated in the
closest proximity to the main island, Mauritius, but even
connected with this latter by several small islands! The
“ possible cause for its [unproved] extinction” is supposed
by Mr. Mason to be scarcity of food, “brought about by a
series of dry or tempestuous seasons unduly prolonged.”
Is this mere speculation, or has Mr. Mason some proof that
a series of dry ‘‘or’’ tempestuous seasons have occurred in
La Ronde? Is it likely that all the individuals of a species
of fruit-bat died for want of food when they had only to fly
15 miles to find abundance of food?—'The plain facts are
these: a person finds a bat skull lying superficially in a
fissure ; the skull is that of a species hitherto not recorded
from the place where found or its nearest neighbourhood ; the
safest conclusion would seem to be: this person has come
across the (actual or deserted) hiding-place of some fruit-
bats, and this skull is unquestionable evidence that this very
species, though hitherto not known from that place, must
live there or in the neighbourhood; Mr. Mason’s conclusion
is: I am unable to refer this skull to any species recorded in
literature from that place; consequently it is an extinct
species.
Pteropus mascarinus, Jason. 355
The above remarks may be briefly epitomized as follows :—
Pt. mascarinus is very closely related to, perhaps identical
with, Pt. rodricensts; Mr. Mason was unaware of this fact,
and therefore failed to point out any character by which
it can be distinguished from Pt. rodricensis ; and he has not
given a particle of evidence that Pt. mascarinus is an ewtinct
species.
Measurements of Pteropus mascarinus * and rodricensis.
Pt. mascarinus. Ft. rodricensis.
Type Type.
(after Mason). ¢ ad. 3 ad. © ad.
76.3.11.1. | 76.3.15.14. | 76.3.15.15.
mm. mm. mm. mm.
Skuil: upper length ...... (49) noi 52:2
», condylo-basal length . (46) Bete 51:7 Siciale
3) basilarlengthy jasc. (44) ees 46 ries
» zygomatic width.... (27) 30°8 ate 28°2
» across m*—m? ext. 3 12°8 13°3 3
» across p*—p* ext. 2). . 13:8 13°6 14 13:1
» interorbital width 7:3 TO ‘is 88
Mani dtlal Oia. Sp ste tis) tears oe 40°6 412 41:2 39-2
Upper teeth, c-m? ........ 2071 19:8 19 185
Lower teeth, e-m, ........ 22°3 22 22 20°7
(AG (Sy Ys 8 A ae A, A 38 37 38 37
TPRNVLCLG LEY syrstersteaveralaeealastinites 2°3 2°7 2°6 2-6
Popa a aan Recline incest 6 3:3 3°38 39 o7
spf. WCDI ca cpanel Spououepaists Payal 3 2:8 2°8 2°77
ml length .ccererais testers ne tar 41 4:1 4°] 3:9
gps 9 WCHL cr vaptarectetens: «teks 2°1 21 25 2:2
im, lengthy. as sis GuisioDCaor Ae 2 2 2
5a, WLGUE ss a lsrereevctatnralers oat IEF 16 167)
Png LETC bait rater cs siaxsiei ciel «toe 2 2 2 18
Mabe WWE votees raat tete x se, oF oe 18 138 16
appl CH vratwste ccarateveter fares 38 41 4-1 39
Pee SWACLLLG oe ratisucy aveieiseessrsies sre 21 2:2 2:2 2:2
Piey LON lee rer charesnersterssoctshe ots 35 39 4:1 38
WAG thw rota ckaroctesitsta st. ae 2°3 2:5 2:3
sd std Nee ORCL CI OSE 4:5 3:7 39 37
pA aire tn eeticns Che eRe 2 2:1 2:2 Pal |
Mew LOT SUN U state! ala jel cet aiostevens oe 3 2°8 2:9 3
jy hy Md Uimararetocetey ene en ctere(cscres 1:8 2 Z 2
Meas LEDGES wercjeforers cotcies fom 15 15 a7 1-4
yy. CWE areas Boe BOOED OE 14 1:2 13 12
* The measurements in parentheses of P¢, mascarinus are approximate.
356 Canon A. M. Norman on the
XLVIT.—Wotes on the Crustacea of the Channel Islands. By
Canon A. M. Norman, M.A., D.C.L., OL.D., F.RB.S.,
BLES:
[Plates XVI. & XVII.]
TuE following publications relate to those Crustacea which
are already recorded from the Channel Islands :-—
1. ‘The Channel Islands.’ By D.T. Ansrep and R. G, Larnam.
1862, pp. 231-235. In this work the list of Cirripedia was
supplied by Mr. Collings, that of the Stalk-eyed Crustacea by
Mr. Lukis and Myr. Le Lievre, that of the Isopodaand Amphipoda
by Mr. Le Lievre, and that of the Entomostraca by Mr. Collings.
2. ‘Recherches sur la Faune Marine des Iles Anglo-Normandes.’ By
Dr. René Ka@uier. 1885.
3. WALKER (A. O.) and Hornet (J.). “ Report on the Schizopoda,
Cumacea, Isopoda, and Amphipoda of the Channel Islands,”
Journ. Marine Zoology and Microscopy, vol. ii. 1896, pp. 49-55.
. THompson (Isaac C.). “Report on the Plankton Copepoda of the
Channel Isles,” Journ. Marine Zoology and Microscopy, Dee. 1897.
I have not seen this paper, but, according to Mr. Sinel, it contains
thirty-one species.
. SineL (JosEPH). “A Contribution to our Knowledge of the
ane of the Channel Islands,” Trans. Guernsey Soc. of Nat.
Sci. 1906.
rc
cn
M. Chevreux collected at Jersey, and sent a list of species
to Messrs. Walker and Hornell for use in their Catalogue.
THE STALK-EYED CRUSTACEA (EXCEPT SCHIZOPODA).
The following is a full list of the Stalk-eyed Crustacea
which have been recorded. I have found almost all the
species myself, but they are here given on the authority of
(A.) Ansted, (.) Koehler, (S.) Sinel. Where no letter
follows a name all three writers include the species; where
an initial letter or letters follow the name it must be under-
stood that the writer or writers thus indicated record the
species.
After the list I add some notes on a few species.
Many of the Stalk-eyed Crustacea which are found in
the Channel Islands have been figured by Mr. J. Sinel
in his recently published ‘An Outline of the Natural
History of our Shores,? 1906 (Swan, Sonnenschein, & Co.).
This is a very useful work for any person commencing the
study of marine zoology.
Crustacea cf the Channel Islands.
Ebalia tuberosa, Pennant.
— tumefacta, Montagu.
Cranchii, Leach.
Corystes cassivelaunus, Montagu.
Atelecyclus septemdentatus, Mon-
tagu. A., S.
Thia polita, Leach.
Polybius Henslowii, Leach. A., 8.
Portumnus latipes, Pennant. K.,
Portunus puber, Linné.
corrugatus, Pennant.
depurator, Linné. K., 8.
marmoreus, Leach. K., S.
holsatus, Fabricius.
pusillus, Leach.
arcuatus, Leach.
Bathynectes longipes, Risso. SS,
Carcinus meoenas, Linné.
Perimela denticulata, Montagu.
Pilumnus hirtellus, Zrnné.
Xantho floridus, Montagu.
hydrophilus, Herbst.
Cancer pagurus, Linné.
Gonoplax angulatus, Pennant. S.
Nautilograpsus minutus, Linné. A,
Pinnotheres pisum, Pennant.
Eurynome aspera, Pennant,
Mamaia squinado, Herbst.
Pisa tetraodon, Pennant.
biaculeata, Montagu.
Hyas coarctatus, Leach.
araneus, Linné, K.
Inachus dorsettensis, Pennant.
dorynchus, Leach.
leptochirus, Leach. K., S.
Acheeus Cranchii, Leach. K., S.
Macropodia rostrata, Linné.
longirostris, Fabricius.
— «xg eyptia, H, M.-Edwards.
KS
Dromia vulgaris, H, M.-Edwards.
Anapagurus levis, W. Thompson,
307
Anapagurus Hyndmanni, W.
Thompson. K., 8.
Pagurus bernhardus, Linné.
Prideauxii, Leach.
— cuanensis, W. Thompson.
I EGET,
Galathea strigosa, Linné.
squamifera, Montagu.
—-nexa, Embleton K.?, 8.
intermedia, Ldljeborg. S.
Porcellana platycheles, Pennant.
longicornis, Pennant.
Callianassa subterranea, Montagu.
Upogebia stellata, Montagu.
Axius stirynchus, Leach.
Arctus ursus, Dana.
Palinurus vulgaris, Latreille.
Homarus gammarus, Linné.
Crangon vulgaris, Linné.
Pontophilus s spinosus, Leach. 8.
fHgeon(?) trispinosus, fazlstone.
fasciatus, Risso. K., 8.
sculptus, Bell. A., 8.
Nika edulis, isso. K., S.
Lysmata seticaudata, Risso. S.
Alpheus macrocheles, Hailstone.
ES, 3.
Athanas nitescens, Montagu.
Hippolyte varians, Leach. K., S.
, var. fascigera, Gosse. S.
Prideauxiana, Leach. S.,
Spirontocaris Cranchii, Leach. K.,
Pandalus annulicornis, Leach. K.,
S,
Pandalina brevirostris, Rathke. S.
Anchistia scripta, Heller. 8S,
Paleemon serratus, Pennant.
squilla, Leach. KS:
Paleemonetes varians, Techs ALS:
Squilla Desmarestii, Paso.
I dredged several specimens of
this species near St. Peter’s Port Harbour, Guernsey, last
summer (1906), and 4. Hyndmannt and P. cuanensis in the
same neighbourhood,
Galathea dispersa, Bate.
intermedia, Lilljeborg.
Both these species occur off Guernsey, but I have not seen G. neva
from the Channel Isles.
It is possible that the species which has
been recorded as G. newa was really G. dispersa.
358 Canon A. M. Norman on the
Lysmata seticaudata, Risso.
1816, Melicerta seticaudata, Risso, Hist. nat. Crustacés de Nice, p. 110,
pl. ii. fig. 1 ' :
1826. Lysmata seticaudata, Risso, Hist. nat. de l’Europe mérid. vol. v.
0. 62.
1828. Lysmata seticaudata, Roux, Crust. de la Médit. pl. xxvii.
1837. Lysmata seticaudata, H. Milne-Edwards, Hist. nat. des Crust.
vol. ii. p. 386, pl. xxv. fig. 10; Atlas Rég. Anim. de Cuvier, pl. liv.
fio. 3.
1863. Lysmata seticaudata, Heller, Crust. des siidlichen Europa, p. 234,
pl. vill. fig. 1. :
1906. Lysmata seticaudata, Sinel (J.), An Outline of the Natural
History of our Shores, p. 134.
Antennules with three flagella. Rostrum with six teeth above,
of which two or three are on the carapace and two below. First
feet didactyle, tolerably strong. Second feet of both sides greatly
lengthened and having the carpus divided into very numerous
jointlets. Telson with two pairs of dorsal spines. Outer uropod
two-jointed. The colouring is peculiarly characteristic, being
brilliant red, with four pale or white lines extending longitudinally
along the entire length of the animal.
About the year 1889 Mr. J. Sinel procured a specimen of this
beautiful Mediterranean prawn in a lobster-pot at La Rocque,
Jersey. He wrote to ask me what it was, introducing into his
letter a pencil sketch. That sketch undoubtedly represented
Lysmata seticaudata, a most interesting addition to the fauna.
Unfortunately the present existence of this specimen is unknown.
Mr. Sinel tells me that he parted with it to some collector, but he
cannot recall to mind who was the recipient. Perhaps this note
may bring to light the possessor of this unique British specimen.
Spirontocaris pusiola, Kroyer. I haye a Jersey specimen sent to
me by Mr. Sinel many years ago.
Anchistia scripta, Risso.
1816. Alpheus scriptus, Risso, Hist. nat. de l'Europe mérid. vol. v.
pave:
(?) Periclemenes elegans, Costa, Fauna del Regno di Napoli, pl. vi.
figs. 1-6.
1849. Palemon biunguiculatus, Lucas, Anim. Artic, de Algérie, p. 45,
pl. iv. fig. 4.
1861. Dennisia sagittifera, Norman, Ann. & Mag. Nat. Hist. ser. 3,
vol. vill. p. 8 (separate copy), pl. xiii. figs. 8-13.
1863. Anchistia scripta, Heller, Crust. des siidlichen Europa, p, 256,
pl. vii. figs. 18, 19.
I dredged the specimen described by me in 1861 in St. Catherine’s
Bay, Jersey, among Zostera in 1859. Mr. Sinel took a second
example in St. Clement’s Bay, Jersey, in 1890, and Mr. Frank
Wright captured a third near St. Peter’s Port, Guernsey, in 1905.
Crustacea of the Channel Islands. 359
SCHIZOPODA,
In the lists which follow, initials affixed to a locality
make known the naturalist on whose authority it is inserted.
‘Thus :—
H, VWornell (James).
N. Norman (A. M.).
S, Sinel (Joseph).
S. §& H. Sinell and Hornell.
W. Walker (A. O.).
W. & H. Walker and Hornell.
In cases where no initial follows the locality or localities
it will be understood that they are on my own authority.
Nyctiphanes Couchii, Bell. Jersey (Sinel, fide Kahler).
Siriella jaltensis, Czerniavsky. Guernsey and Jersey.
Clausii, G. O. Sars. Guernsey.
armata, H. M.-Edwards. Jersey.
Gastrosaccus sanctus, P. J. van Beneden. Jersey.
spinifer, Goés. Jersey (Museum at Bootle, Lancashire, fide W.).
Anchialina agilis, G. O. Sars. Off Alderney, tow-net (Dr. Gough
in Mus. Nor.) ; Guernsey (W.).
For the genus Anchialina, see Norman and Scott, ‘Crustacea of
Devon and Cornwall,’ 1906, p, 24.
Heteromysis formosa, 8. I, Smith. Guernsey.
Mysidopsis gibbosa, G. O. Sars. Guernsey (A. O. Walker, in Titt.,
Sept. 4, 1892).
Leptomysis mediterranea, G. O. Sars. Guernsey and Jersey.
Mesopodopsis Slabberi, P. J. van Beneden, Jersey (Musewin at
Bootle, Lancashire, fide W.).
Macromysis fleauosa, Miller. Abundant.
Mr. Sinel, whom I had the pleasure of seeing at Jersey last
year (1906), told me that species of Mystdea were caught at
Jersey and made into a paste called ‘ Chervé,” used in
fishing. He subsequently, in answer to my enquiries, sent
me the following account of this preparation :—
“ At present the fishers of ‘Chervé’ have to obtain a
licence trom the chief fishery inspector to enable them to
carry on this very old practice—a very absurd law. ‘hey
take it both day and night during the spring tides in the
large gravelly pools, among the Zosrera, in the southern bays.
“Tt is taken in large hand-nets, with usually a cross-bar
360 Canon A. M. Norman on the
to the handle, so that the fisher’s breast bears on it [see
figure]. The nets are from 3 feet to 4 feet 6 inches across
the wooden (ash) blade which forms the base. The best
nets are of horsehair cloth, as used for sieves, but ordinary
linen cheese-cloth is mostly used. At about 18 inches from
the pocket of the net a cane ring with netting of about three
quarters of an inch mesh is inserted across it, so as to prevent
the entrance of larger objects (crabs, shrimps, &c.) among
the Mysids.
“The men usually wade waist-deep, pushing the net before
them for ten minutes or so, when they untie the pocket end
and empty their catch into their basket.
“When the ‘Chervé’ is brought home it is mixed with
a sufficient quantity of coarse salt to preserve it, and then it
is pressed into pans. It forms a pink-coloured compost.
This is sold to mullet-anglers at about four shillings a gallon,
though many anglers obtain a licence and take their own
supply.
“The favourite places for mullet are not far from the
shore, and high water is the best time for the sport. The
angler takes a spoonful of ‘ Chervé’ from his tin, uses it as a
ground-bait, and waits awhile. If no mullet turn up in ten
or twenty minutes, he tries another spot or goes home; but
almost invariably a few are soon attracted, to be followed by
a crowd. The ‘Chervé’ tints the water pinkish for some
yards around, and has a strong and not pleasant smell, like
that of boiled prawn or lobster gone bad.
Crustacea of the Channel Islands. 361
““T am constantly consulted respecting some substitute for
‘Chervé’: crabs and shrimps pounded in a mortar and
salted; ray-fish boiled with sugar; bread soaked in shrimp
soup ; cabbage boiled in fat, &c., &e.; but ‘Chervé’ still
holds its own. I have often been astonished when sitting on
a jetty fishing for Labrus &e., and not a mullet in sight for
hours; then down comes a ‘ mulleter,’ who throws in a spoon-
ful of ‘Chervé,’ and in a minute one or two dark forms come
into the field, then more and more, and their silvery sides
flash as they turn about and suck up the ‘ Chervé.’ ”’
Mr. Sinel kindly sent me a bottle of ‘‘ Chervé”’ as taken
from the net. Small pieces of seaweed were mixed with the
animals. Both in numbers and in bulk Macromysis flexuosa
predominated, but Schistomysts spiritus was present in large
numbers, together with afew Siriella armata. No doubt the
admixture of species would vary in different spots and in
different seasons. That the Mysidea were used as bait was
quite a new fact to me, as it probably will be to other
carcinologists.
Macromysis neglecta, G. O. Sars. Guernsey and Jersey.
mermis, Rathke. Guernsey.
Schistomysis spiritus, Norman. Guernsey and Jersey.
ornata, G. O. Sars. Off St. Sampson’s Harbour, Guernsey.
Helleri, G. O. Sars. Guernsey and Jersey.
Neomysis vulgaris, J. V. Thompson. Arnold’s Pond, Guernsey, 1865.
SyMPoDA (=CUMACEA).
Iphinoe trispinosa, Goodsir. Off St. Sampson’s, Guernsey (N.);
Jersey (W. & #.).
serrata, Norman. (Guernsey.
Cumopsis Goodsiri, P. J. van Beneden. Abundant from washing
sand dug at low-water spring tides in Belgrave Bay, Guernsey.
Eudorella truncatula, Bate. Dredged Guernsey, 1565.
Diastylis levis, Norman. Dredged near St. Martin’s Point,
Guernsey.
Pseudocuma similis, G. O. Sars. Shallow water, sand, Guernsey.
longicornis, Bate. Jersey (W. & H.).
Cumella pygmea, G. O. Sars. Guernsey.
Nannastacus unguiculatus, Bate. Guernsey (W.); Jersey (W.).
ISOPODA.
Apseudes Latreillii, Milne-Edwards. Very abundant off Guernsey
(A. M. N.); Jersey (W. & H.). Bate and Westwood state
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 24
362 Canon A. M. Norman on the
that I had taken A. talpa off Guernsey, but the species I sent
them was A. Latreillit.
Apseudes talpa, Leach. Jersey (Kehler).
Tanais Cavolinii, Milne-Edwards. Guernsey and Jersey (S. & H.).
Leptochelia Savignii, Kroyer. Guernsey (V.); Herm (Kehler) ;
Jersey (S. & H.).
Tanaissus Lilljeborgii, Stebbing. In sand, low water, Belgrave
Bay, Guernsey.
Paratanais Batei, G. O. Sars. Jersey (S. & H.); Sark (Kehler), as
Paratanais foreipatus.
Anthura gracilis, Montagu. Jersey (S. § H.).
Paranthura nigropunctata, Lucas. Guernsey, 1865 (N.); Jersey
GS. g7.).
Gnathia maxillaris, Montagu. Guernsey (N.); Jersey (S. § H.).
Anilocra physodes, Linné (=A. mediterranea, Leach). Herm, 1865
(NV.); Jersey (S. f& H.); Guernsey (7. Wright).
Aga rosacea, Risso, = Aga bicarinata, Leach, and Bate and West-
wood. A specimen of this species was brought to me by a
trawler last year at Guernsey. In 1894 (Ann. & Mag. Nat.
Hist. ser. 7, vol. xiv. p. 433), and again last year in Norman
and Scott (‘The Crustacea of Devon and Cornwall,’ p. 38), I
questioned whether ga rosacea could with certainty be
regarded as a British species; but I had overlooked the fact
that Sinel and Hornell had recorded the occurrence (in Walker
& Hornell’s paper) of two specimens taken at Jersey on the
fish Squatina angelus.
Cirolana Cranchvi, Leach. Off St. Sampson’s Harbour, Guernsey
(V.); Jersey (Kehler).
borealis, Lilljeborg. Guernsey, 1865, rare.
Conilera cylindracea, Montagu. Guernsey and Jersey.
Eurydice spinigera, H. J. Hansen. Jersey (Sinel, in Mus. Nor.).
Limnoria lignorum, Rathke. Jersey (S. & #.).
Spheroma serratum, Fabricius. Herm (N.); Jersey (S. & #.).
Hookeri, Leach. Guernsey, Cobo Bay &c., 1865.
Cymodice truncata, Montagu. Guernsey and Jersey.
Nesa bidentata, Leach. Guernsey, Herm, Jersey.
Campecopea hirsuta, Montagu. Guernsey (NV.); Jersey (W. & H.),
Idotea balthica, Pallas. Common.
neglecta, G.O. Sars. Guernsey, Jersey.
emarginata, Fabricius. Jersey, tow-net (Kehler).
—— lmearis, Linné. Guernsey, very large (N.); Jersey (S. ¢ H.).
salinarum, Dollfus (=? viridis, Slabber). Arnold’s Pond,
Guernsey, 1865.
Zenobiana prismatica, Risso. Jersey (Sinel, in Mus. Nor.).
Crustacea of the Channel Islands. 363
Stenosoma acuminatum, Leach. <A single specimen, St. Aubin’s,
Jersey (Kehler).
laneiferum, Leach. Jersey. In Walker & Hornell’s list, but
no habitat is given.
Astacilla longicornis, Sowerby. Guernsey, 1865.
Asellus aquaticus, Linné. Guernsey.
Janira maculosa, Leach, Jersey (S. § H.).
Jera Nordmanni, Rathke. Jersey (S. & H.); Sark (Kehler).
marina, Fabricius. Guernsey, Jersey.
Jeropsis brevicornis, Keehler. The types, Gouliot Caves, Sark
(Kehler, who kindly gave me a specimen).
Munna Kréyeri, Goodsir. Jersey (S. & H.).
limicola, G. O. Sars. Dredged, Guernsey.
Ingia oceanica, Linné. Guernsey and Jersey.
Oniscus asellus, Linné. Abundant.
Porcellio scaber, Latreille. Common.
Platyarthrus Hoffmanseggvi, Brandt. St. Sampson’s, Guernsey.
Armadillidium vulgare, Latreille. Abundant.
Bopyrus squillarum, Latreille. Guernsey and Jersey.
Bopyrina Giardi, Bonnier. Many on Hippolyte varians, Jersey,
Bopyroides hippolytes, Kroyer. “ Parasitic on Hippolyte” (S. & #.).
Pleurocrypta galathee, Hesse. In Galathea squamifera, Guernsey
(N.); Jersey (Kehler).
strigosa, Giard & Bonnier. In Galathea strigosa.
intermedia, Giard & Bonnier. In Galathea intermedia,
Guernsey.
Athelges paguri, Rathke. On Pagurus bernhardus, Guernsey, 1865
(V.); Jersey (Kehler).
Gyge ee Cornalia & Pancer. In Upogebia stellata, Jersey,
1859.
Tone thoracica, Montagu. In Callianassa subterranea, Jersey
(S. § H.).
Liriopsis pygmea, Rathke. Guernsey, 1865.
AMPHIPODA.
Hyperia galba, Montagu (S. & #.).
Talitrus locusta, Pallas. Common.
Orchestia gammarellus, Pallas. Guernsey (N.); Jersey (Kehler).
mediterranea, Costa. Jersey (Chevreux, W.).
Hyale Nilssoni, Rathke. Jersey (Chevreux, W.).
—— Lubbockiana, Bate. Jersey (Kehler, S.).
24%
bol
364 Canon A. M. Norman on the
Lysianassa plumosa, Boeck,=L. Coste, B. & W. (nec Milne-Edw.).
Guernsey and Jersey.
ceratina, Walker, =? longicornis, Lucas. Guernsey and Jersey.
Acidostoma obesum, Bate. Jersey (S.).
Socarnes erythrophthalmus, D. Robertson. Guernsey.
Perierella Audouiniana, Bate. Jersey (Chevreua, S.).
Tryphosites longipes, Bate. Guernsey (N.); Jersey (S.).
Hippomedon denticulatus, Bate. Jersey (S.).
Haplonyx cicada, Fabricius, = A. Holboll, B.& W. Jersey (S. § H.).
Callisoma Hopei, Costa,=C. crenata, Bate. Jersey (S.).
Orchomene humilis, A. Costa,=O. Batei, G. O. Sars. Jersey.
Tryphosella Sarsi, Bonnier. Jersey (W. & H.).
Nannonyx Goesi. Grand Azette, Jersey, spring tide (W. § Z.).
Menigrates obtusifrons, Boeck. Jersey (S. dg: #.).
Lepidepecreum longicorne, Bate. Guernsey (NV., W.); Jersey
(Sig -.):
Bathyporeia Guilliamsonia, Bate, = B. norvegica, G. O. Sars.
Guernsey (V.); Jersey (S. & H.).
—— pelagica, Bate. In sand, low water, Belgrave Bay, Guernsey
(W.), Jersey (S. § H.).
Robertsoni, Bate. Jersey (S.).
Haustorius arenarius, Slabber. Jersey (S.).
Urothoe marinus, Bate. Jersey (Chevreux, S.).
—— brevicornis, Bate. Guernsey (W.); Jersey (S. & #.).
elegans, Bate. Jersey (S. f§ H.).
pulchella, Costa. Jersey (Chevreux).
Metaphowus simplex, Bate. Guernsey (N.).
Fultoni, T. Scott. Jersey (Chevreux); off Noirmont Point
(S.5d7 7):
Harpinia neglecta, G. O. Sars, =Phowus plumosus, Bate. Jersey (S.).
pectinata, G. O. Sars. Dredged off Belgrave Bay, Guernsey,
several specimens. It was not a species which I could have
expected to find in the Channel Islands.
Ampelisca typica, Bate. Guernsey (1V.); Jersey (S. §- H.).
tenuicornis, Lilljeborg. Guernsey (V.); Jersey (Chevreux).
assimilis, Boeck. Off St. Martin’s Point, Guernsey.
—— spinipes, Boeck. Guernsey.
brevicornis, Costa,= A. Belliana, B. & W. Guernsey (N.);
Jersey (S. & H.).
—— gibba, G. O. Sars. Off St. Peter’s Port, Guernsey, in 7 fath.,
1892 (W.).
Amphilochus manudens, Bate. Jersey (S. § H.).
Crustacea of the Channel Islands. 365
Amphilochus neapolitanus, Della Valle, = A. melanops, Walker.
Bordeaux Harbour, Guernsey; and Jersey.
Peltocova Marioni, Catta, = Cyproidea damnoniensis, Stebbing.
Jersey.
Stenothoe monoculoides, Montagu. Guernsey and Jersey.
marina, Bate. Jersey (Kehler).
Metopa rubrovittata, G. O. Sars. Bordeaux Harbour, Guernsey.
borealis, G. O. Sars. Off St. Martin’s Point, 25 fath. (W.).
Metopa sarniensis, sp.n. (Pl. XVII. figs. 1-5.)
A very small species. The antennules and antennz are
subequal in length, and the peduncles of both are quite
smooth and devoid of sete. The first joint of peduncle of
the antennules is stout and equals in length the two following
joints; the flagellum is 8-jointed, the first of these joints
equals in length the last two joints of the peduncle. The
first gnathopod (fig. 1) is remarkable on account of the very
short carpus, which is somewhat triangular; the propodos
equals nearly three times the length of the carpus, it gradually
tapers for the attachment of the gently curved finger; the
propodos has one seta on the side near the end and a bundle
of three below the attachment of the finger; the finger (fig. 2)
has its distal portion minutely denticulated or serrated ; two
minute sete are on its side above the serrations, and there
are no others.
The second gnathopod (fig. 3) is subchelate and resembles
that of many members of the genus in its general form. The
hand widens gradually and slightly from its base, and distally
is abruptly truncated, so that the palm is transverse (fig. 4);
the nail is strong and about equals the transverse palm in
length; it has on its back four setae, and on the palm where
the tip of the finger meets it there is a group of slender spines
and a few sete.
The last perzeopods (fig. 5) have the basos outspread behind
into a thin lamina, which is devoid of sete or spines; the
meros is expanded behind and projects down to near the
extremity of the carpus.
The first gnathopods are, perhaps, more valuable in this.
genus for specific characters than the second, though the
latter are so much the larger. In this species the first
gnathopod is quite different from that of any other species
of the genus known to me; both the very short carpus and
the serrated tip of the finger are quite exceptional.
A single specimen was dredged, July 1906, not far from
the harbour of St. Peter’s Port, Guernsey.
366 Canon A. M. Norman on the
Leucothoe spinicarpa, Abildgaard. Guernsey (W.); Jersey (S.d:1.).
LTilijeborgii, Boeck. Guernsey.
—— “ furina, Savigny.” Jersey (Snel). Perhaps the same as the
last.
Pereionotus testudo, Montagu. Jersey (S.).
Laphystius sturionis, Kroyer,=Darwinia compressa, Bate. Jersey
(S21 #.).
Colomastix pusilla, Grube. Jersey (Chevreua) ; Sark (Kehler).
Odius carinatus, Bate. Jersey (S.).
Epimeria cornigera, Fabricius, = Acanthonotus Owent, B. & W.
Jersey (S.).
Iphimedia obesa, H. Rathke. Guernsey (W.); Jersey (S.).
—— minuta, G. O. Sars. Guernsey.
Eblane, Bate. ‘‘ Not common, Vermarette, Herm” (S.).
Perhaps the preceding species, which the author did not know.
Lilljeborgia pallida, Bate. Jersey (S.).
picta, Norman. The type specimens were taken by me off
Guernsey in 1865.
Monoculodes carinatus, Bate. Off St. Sampson’s Harbour, Guernsey
(N.); Jersey (W. & H.).
Perioculodes longimanus, Bate & Westwood. Guernsey and Jersey.
Pontocrates arenarius, Bate. ‘St. Clement’s Bay, Jersey, not
common” (S.). Perhaps not this, but the next species.
norvegicus, Boeck. Guernsey (NV.); Jersey (Chevreux).
Synchelidium haplocheles, Grube. Guernsey (NV.); Jersey (W.g H.).
Calliopius leviusculus, Kroyer,=C. Rathkei, Zaddach. Guernsey
(N.); Jersey (W. & H., S.).
Apherusa cirrus, Bate,=A. borealis, G. O. Sars. Guernsey and
Jersey.
ovalipes, Norman & Scott.
1906. Apherusa ovalipes, Norman & Scott, Crustacea of Devon and
Cornwall, p. 75, pl. viii. figs. 1-8.
Fermain Bay, Guernsey.
bispinosa, Bate. Guernsey (V.); Jersey (W. & Z., 8.).
—— Jurinei, H. M.-Edwards. Tide-marks, Belgrave Bay,
Guernsey (V.); Jersey (W. & #.).
Sympleustes latipes, M. Sars. Jersey (S.).
Nototropis Swammerdamii, H. M.-Edwards. Jersey (N.).
vedlomensis, Bate. Fermain Bay, Guernse SR ;
(W. & H., 8). : Recut
Dexamine spinosa, Montagu. Guernsey and Jersey.
thea, Boeck. Guernsey and Jersey.
Triteta gebbosa, Bate. Jersey (W. & H., S.).
Crustacea of the Channel Islands. 367
Guernea coalita, Norman. Bordeaux Harbour, Guernsey; and
Jersey.
Eusirus longipes, Boeck. Shore near St. Helier’s, Jersey (W. S H.).
Amathilla Homeri, Fabricius. Jersey (S. § Z.).
Isea Montagu, H. M.-Edwards. On Mamaia squinado, Guernsey
(N.); Jersey (S. & #.).
Niphargus fontanus, Bate. Sinel records this from four covered-in
wells in Jersey.
subterraneus, Leach, =N. aquilex, B.& W. Mr. Frank Wright,
of the Library, St. Peter’s Port, has recently sent me a specimen
of this species taken in a well at Guernsey.
Cheirocratus Sundevalli, H. Rathke. Fermain Bay, Guernsey (JV.) ;
Jersey (Chevreua).
assimilis, Lilljeborg. Near St. Martin’s Point, Guernsey,
Gammarella brevicaudata, H. M.-Edwards. Guernsey (1V.); Jersey
GWE Gc S.)s
Gammarus locusta, Linné. Common.
pulex, De Geer. Common.
— marinus, Leach. Guernsey and Jersey (W. & H., S.).
campylops, Leach. Jersey (Chevreux, W. & H., S.)
Echinogammarus Berillont (Catta). (Pl. XVI. figs. 1, 2.)
1878. Gammarus Berilloni, Catta, ‘‘ Note sur le Gammarus Berillont,
n. sp.,” Bull. Soc. Borda Dax. Lier trimestre, p. 1, pl. i. figs. 1, 2.
1896. Gammarus Berilloni, Chevreux, Bull. Soc. Zool. France, vol. xxi.
p. 24, and woodcuts.
1906. Echinogammarus Berillont, Stebbing, Das Tierreich, Amphipoda,
I. Gammaridea, p. 481,
I am indebted to M. Chevreux for one of the examples of this
interesting species which he procured in Jersey. It is at once
distinguished from our previously known British allies by the dense
clothing of spines and setz on the metasome. It was found by
Catta at Mondarran (Basses-Pyrénées), and has subsequently been
met with again, but only in the same district, namely at St. Jean
de Lux and near Biarritz, and I have received from Don Ignacio
Bolivar, of the Madrid Museum, specimens which came from San
Sebastian. It is remarkable that this freshwater species should
have been met with at such a place as Jersey.
Melita palmata, Montagu. Guernsey (NV.); Jersey (W. & ZZ, S.).
obtusata, Montagu. Guernsey (V.); Jersey (W. & H., S.).
gladiosa, Bate. Guernsey (NV.) ; Jersey (S.).
Mera grossimana, Montagu. Guernsey, Herm, Jersey.
othonis, H. M.-Edwards. Guernsey.
— Batei, Norman. I dredged the type specimens off St. Martin’s.
Point, Guernsey, in 1865.
Ceradocus semiserratus, Bate. Guernsey, off St. Martin’s Point.
368 Canon A. M. Norman on the
Megaluropus agilis, Norman. Guernsey, in tow-net.
Microdeutopus danmoniensis, Bate. Guernsey and Jersey (W.).
gryllotalpa, Costa. (Pl. XVI. fig. 3; Pl. XVII. figs. 6, 7.)
1893. Microdeutopus gryllotalpa, Della Valle, Faun. und Flor. des
Golfes von Neapel, Gammarini, p. 44, pl. i. fig. 12, pl. xi. figs. 25-43.
1894. Microdeutopus gryllotalpa, G. O. Sars, Crust. Norway, Amphi-
poda, p. 543, pl. excil. fig. 2.
1906. Microdeutopus gryllotalpa, Stebbing, Das Tierreich, Amphipoda,
I. Gammaridea, p. 590.
Found at Jersey by M. Chevreux and Mr, Sinel.
I have given figures of the one-jointed accessory flagellum of the
antennules (Pl. XVII. fig. 6), which is shorter than the first joint of
the flagellum, and of the first gnathopod of the male, for comparison
with the same parts in the following species. This gnathopod has
2-4 toothed processes on the hinder margin of the carpus, which
inerease in size distally. Della Valle gives only two such processes.
I figure (Pl. XVII. fig. 7) a gnathopod with three such processes
from Plymouth (as figured by Sars and by Bruzelius, Autonoe
grandimana), and (Pl. XVI. fig. 3) another carpus from Valencia,
Spain, with four processes (as figured by Costa).
stationis, Della Valle. (Pl. XVI. fig. 4; Pl. XVII. figs. 8-11.)
1888. Microdeutopus gryllotalpa, Nebeski, Beit. zur kennt. der Amphip.
der Adria, p. 45, pl. iv. fig. 41.
1893. Microdeutopus stationis, Della Valle, Faun. u. Flor. des Golfes
von Neapel, Gammarini, p. 415, pl. v. fig. 9, pl. x. figs. 31-41.
1906. Microdeutopus stationis, Stebbing, Das Tierreich, Amphipoda,
J. Gammaridea, p. 590.
Sowinsky has referred a Microdeutopus found in the Black Sea to
this species ; but I cannot reconcile the fig. 6 in his paper of 1891
with the first gnathopod (male) of this species. Nebeski gives an
interesting series of figures illustrative of the gradual change in
form of the gnathopod from the young to the adult.
I last year dredged many specimens of this addition to our fauna
off St. Peter’s Port, Guernsey.
I have figured some of the most salient features which distinguish
this species from M. gryllotalpa. The secondary appendage of the
antennules is 3—4-jointed and is equal in length to the first three
joints of the flagellum (Pl. XVII. fig. 8). The first gnathopod in the
male has not quite so massive a carpus as in that species (Pl. XVII.
figs. 9 & 10), The carpus is armed with three teeth at the extre-
mity (not passing down the hind margin); these teeth are not in
the same straight line, the innermost and bluntest tooth bends away
from the large central tooth, so that it would appear that the finger
closes down between them. Pl. XVII. fig. 11 represents the distal
portion of the first gnathopod of the female, and Pl. XVI. fig. 4 the
second gnathopod of the male.
Stimpsonella chelifera, Bate. Guernsey (V.); Jersey (S. § #7.).
Crustacca of the Channel Islands. 369
Coremapus versiculatus, Bate.
1906. Coremapus versiculatus, Norman & Scott, Crustacea of Devon
and Cornwall, p. 85, pl. ix. figs. 8-10.
Guernsey, common (V.); Jersey (Chevreus).
Lembos Websteri, Bate. Sark (Kehler).
Aora typica, Kréyer,=A. gracilis, Bate.
Stebbing has synonymized these two names, and I think that he
is right. I have taken Aora occasionally off Guernsey.
Leptocheirus pectinatus, Norman.
1906. Leptocheirus pectinatus, Norman & Scott, Crustacea of Devon
and Cornwall, p. 87, pl. ix. figs. 1-3.
Off St. Sampson’s, Guernsey (V.); Jersey (Chevreuz).
Leptocheirus guttatus, Grube.
1887. Ptilocheirus tricristatus, Chevreux, “ Crust. Amphip. de la céte
ouest de Bretagne,” Bull. Soc. Zool. France, vol. xii. p. 23, pl. v.
figs. 3, 4, and woodcut at p. 6.
1906. Leptocheirus guttatus, Norman & Scott, Crustacea of Devon and
Cornwall, p. 88, pl. ix. figs. 4-7.
Off St. Sampson’s, Guernsey.
Gammaropsis maculata, Johnston, Jersey (N., S. § H.).
Microprotopus maculatus, Norman. Guernsey (NV.); Jersey
(Chevreux).
—— longimanus, Chevreux. Herm and Guernsey.
Photis longicaudata, Bate. Guernsey (V.); Jersey, off La Rocque
(Si GL):
Podoceropsis Sophie, Boeck. Jersey (S.).
Megamphopus cornutus, Norman. Guernsey (W.); Jersey (W. & H.).
Amphithoe rubricata, Montagu. Guernsey and Jersey.
Pleonexes gammaroides, Bate. Guernsey and Jersey.
Sunamphithoe pelagica, H. M.-Edwards,=S. conformata, Bate.
Guernsey (V.); Jersey (Chevreux).
Bruzeliella falcata, Montagu. Jersey.
ocia, Bate. One male, Jersey (W. g§ H.).
Jassa pelagica, Leach,=Podocerus capillatus, B. & W. Among
Styelopsis, Jersey (Kehler).
Erichthonius abditus, Templeton. Guernsey (V.); Jersey (S. f Z.).
Hunter, Bate. Guernsey.
difformis, H. M.-Edwards. Jersey (S.).
Cerapus crassicornis, Bate. Jersey (S.).
Corophium volutator, Pallas,=C. longicorne, Latreille. Guernsey
(N.); Jersey (S. & H.).
crassicorne, Bate. Jersey (S.).
Bonelli, H. M.-Edwards. Guernsey (.).
Canon A. M. Norman on the
370
Unciola crenatipalmata, Bate. Guernsey, in Fermain Bay in 1906,
and also taken in 1865.
Chelura terebrans, Philippi. Jersey (W. & H., S.).
Dulichia porrecta, Bate. Between the Castle and St. Martin’s Point,
Guernsey (W.); Jersey (W. & H., S.).
Platophium Darwini, Bate. Guernsey (NV.); Jersey, tide-marks ( W.).
Phtisica marina, Slabber. Guernsey and Jersey.
Pseudoprotella phasma, Montagu. Jersey (Awhler).
Guernsey (NV.); Jersey (S.).
Caprella tuberculata, Guérin. Guernsey.
linearis, Linné. Guernsey (1V.); Jersey (W.).
acanthifera, Leach. Common, Jersey (S. d H.).
Guernsey (J.).
Jersey (W. & H.).
Periambus typicus, Kroyer.
, var. levissima, Mayer.
acutifrons, Latreille.
OSTRACODA.
The following list contains the species known to me from
the Channel Isles. The freshwater species have not been
sought for, except the very fine Cypris bispinosa, which was
sent to me many yearsago by the late Dr. Lukis of Guernsey.
The species which have the letter 6 after them are inserted
on the authority of Dr.G. 8. Brady; the rest have been
found by myself.
Cypris bispinosa, Lucas.
Paracypris polita, G. O. Sars.
Pontocypris trigonella, G. O. Sars.
Erythrocypris mytiloides, Norman.
Argyllcecia cylindrica, G. O. Sars.
Bairdia acanthigera, G. S. Brady.
inflata, Norman. B.
Cythere lutea, O. F. Miller.
cyamos, Norman.
— confusa, brady & Norman.
pellucida, Baard.
porcellanea, G. S. Brady.
tenera, G. S. Brady.
badia, Norman.
crispata, G. S. Brady.
Robertsoni, G. S. Brady.
convexa, Buird.
marginata, Norman.
— albomaculata, Baird.
— villosa, G. O. Sars.
Jeffreysi, GS. Brady. B.
cuneiformis, G. S. Brady. B.
Cythere finmarchica, G. O. Sars.
tuberculata, G. O. Sars. B.
runcinata, Baird.
antiquata, Bazrd,
Cytheridea elongata, G'. S. Brady.
torosa, 7’. &. Jones.
Eucythere declivis, Norman.
Loxoconcha impressa, Baird.
viridis, O. F. Miiller.
— multifora, Norman.
— tamarindus, 7. &. Jones.
guttata, Norman.
Xestoleberis aurantia, Baird.
labiata, Brady §- Robertson.
depressa, G'. O. Sars.
Cytherura nigrescens, Baird,
cornuta, G. S. Brady.
sella, Gi. O. Sars.
acuticostata, G. O. Sars.
angulata, G. S. Brady.
producta, G. S. Brady.
fulva, Brady § Robertson.
Crustacea of the Channel Islands. 371
Cytherura striata, G. O. Sars. Paradoxostoma obliquum, G. O.
—— cellulosa, Norman. Sars.
Bythocythere constricta, G. O. pulchellum, G, O. Sars.
Sars. B. fasciatum, Brady & Norman.
Pseudocythere caudata, G. O. Sars. | —— Normani, G. S. Brady.
Sclerochilus contortus, Norman. hibernicum, G. 8S. Brady.
Cytherideis subulata, G. S. Brady. orchadense, Brady § Robertson.
Macherina amygdaloides, G. S. flexuosum, G’. S. Brady.
Brady. Philomedes interpuncta, Baird.
Paradoxostoma variabile, Baird. Asterope Mariz, Baird.
— ensiforme, G. S. Brady. teres, Norman.
——- abbreviatum, G. O. Sars.
CoPEPODA.
In the commencement of this paper it is mentioned that my
late friend Mr. Isaac C. Thompson recorded thirty-one species
of Copepoda from Channel Island plankton. On the Harpac-
ticoida and fish-parasites nothing has been published, and the
few species in my own collection are not worth recording. It
is probable that many minute species new to our fauna will
hereafter be found in what may be expected to prove a
very rich district. It is to be hoped that some naturalist
with good eyes and a steady hand will before long work at
these minims. But very much remains to be done in other
orders, especially among the Sympoda and Ostracoda.
EXPLANATION OF THE PLATES.
PuatE XVI.
Fig. 1. Gammarus Berillont, Catta.
Fg. 2. Ditto. Last uropod.
Fig. 3, Microdeutopus gryllotalpa, Costa. First gnathopod (3), from
Valencia, Spain.
Fig. 4. Microdeutopus stationis, Della Valle. Second gnathopod (¢).
Puate XVII.
Fig. 1. Metopa sarniensis, sp.n. First gnathopod.
Fig. 2. Ditto, The distal joints of the same, more magnified.
Fig. 3. Ditto. Second gnathopod.,
. Ditto. The last perzeopod.
3, Microdeutopus gryllotalpa, Costa. Seconfary flagellum of
antennule.
1
2
3
Fig. 4, Ditto. The distal joints of the same, more magnified,
5
6
7. Ditto. Distal portion of first gnathopod (¢), from Plymouth.
Fig. 8. Microdeutopus stationis, Della Valle. Secondary flagellum of
first antennule.
Fig. 9, Ditto. First gnathopod (¢).
Fig. 10, Ditto. Ditto, to show variation,
Fig. 11. Ditto. First gnathopod (@).
372 Bibliographical Notices.
BIBLIOGRAPHICAL NOTICES.
100 Photographs from Life. (No.1.) By Dovetas Enetisn.
London: Bousfield & Co., 1907.
THis is a Sa wonderful shilling’s worth, containing most charming
pictures of shrew-mice, dormice, house- and field-mice, and the
meadow-mouse.
In some thirty pages of text the Author briefly enumerates the
salient points of each species and the more interesting facts with
regard to the life-history of each. Accurate and pleasantly written,
this addition to the volume will be most welcome.
100 Photographs from Life. (No. 2.) By R. B. Lover.
London: Bousfield & Co., 1907.
Mr. R. B. Lopez has earned for himself a considerable reputation
as a bird-photographer, and this is thoroughly well sustained in this
little booklet—a companion volume to that on the Rodents.
Though all the pictures herein reproduced have appeared else-
where, they will find a no less hearty welcome among bird-lovers
on this account.
Mr. Lodge’s notes on the habits are extremely interesting, and
not less so because he has included therein many details concerning
the making of his pictures. Wik: 72.
Les Débuts Cun Savant Naturaliste, le Prince de VEntomologie, Pierre
André Latreille, a Brive, de 1762 a4 1796. Par Lovis pE Nussac,
Sous-Bibliothécaire du Museum d’Histoire naturelle. 8vo.
Paris, 1907. Pp. 264.
LarreIte is not the least distinguished among the numerous French
clerics who have done so much good work in entomology for more
than a century, and we may also mention Engramelle, Lacordaire,
David, and Joannis. At a time when entomology was in its infancy
in Britain, Latreille laid the foundations of the science deeply and
firmly in the sixteen volumes of bis ‘ Histoire naturelle, générale
et particulicre des Crustacés et des Insectes, published between
1802 and 1805.
But the interesting work before us deals only with the earlier
part of his life, before the publication of his principal works; for,
though he had previously published several entomological papers,
his ‘ Précis,’ the first of the long series of books which made his
name famous, did not appear till 1796. He was born at Brive in
1762, and died in Paris in 1833.
We do not propose to speak of Latreille’s career further, except
to relate again one of the most interesting episodes of his early
life. At the time of the French Revolution, Latreille, who had
Bibliographical Notices. 373
risen to the dignity of an Abbé, was thrown into prison and con-
veyed to Bordeaux for deportation to Cayenne, with other offenders
guilty of belonging to the priesthood, During the visit of a surgeon
to the prison the latter saw Latreille pick up an insect and look at
it carefully. “Is it rare?” he asked. “Yes,” said Latreille.
**Give it me,” said the surgeon, “ for I have a friend who is inter-
ested in insects.” ‘Take it, and ask him to give me its name,”
said Latreille. But the surgeon brought back the reply that he
could not discover the name, and that it was probably a new
species ; and then Latreille gave him a message to his friend Bory
de St. Vincent, a man of considerable influence, who was able to
exert it to release him from his imprisonment, and thereby saved
his life, for the vessel in which he was to have sailed was lost with
all on board, except some of the crew, who escaped in a boat. The
insect, the capture of which led to such important results, was
Necrolia ruficollis, Fabricius, a small beetle belonging to the
Malacodermata.
Epa. Borpaer. Recherches anatomiques et biologiques sur 0 Autotomie
et la Régénération chez divers Arthropodes. (Reprinted from
‘ Bulletin scientifique de la France et de la Belgique,’ vol. xxxix.
1905, pp. 807-454, pl. vi. & 20 text-figs.)
A TREATISE on the power possessed by various insects (especially
Phasmide) of reproducing lost limbs, either when a limb has been
cast by the insect itself to facilitate its escape from an enemy, or
when a limb has been injured, or amputated for experimental
purposes. ‘The author is the Director of the Museum of Natural
History in the island of Réunion, and has paid special attention to
the question in the case of two large Phasmids (Monandroptera
inuncans, Serv., and Rhaphiderus scabrosus, Serv.) which inhabit
that island. The brochure will be very interesting to entomologists
who occupy themselves with insect physiology.
The Fauna of British India, including Ceylon and Burma. Pub-
lished under the authority of the Secretary of State for India in
Council. Edited by Lt.-Col. C. T. Brinenam. — Butterflies.
Vol. II. By Lt.-Col. C.T. Bryenam. London: Taylor & Francis,
1907. 8vo. Pp. vili, 480; pls. xi.—xix.
Tue second volume of this important work includes the Papilionide,
the Pieride, and the following five subfamilies of Lycaenidee :—Gery-
dine, Lycwenine, Curetine, Liphyrine, and Poritiine. Two more
subfamilies of Lycznide (Arhopaline and Thecline) remain over
till the next volume.
It will thus be seen that the volume includes a considerable
portion of the most conspicuous and interesting butterflies. In the
Papilionidee six genera are included—Armandia, Leptocircus, Teino-
374 Geological Society.
palpus, Papilio, Parnassius, and Hypermnestra; the 76 species of
Papilio not being broken into small genera, as was attempted by
F. Moore and others, but arranged in 26 groups, according to the
scheme of Felder, as expanded by the Hon. W. Rothschild, even
Ornithoptera being treated simply as a group. In the Pieride and
Lycenide the genera are much more numerous, and seldom include
many species.
Comparatively few European species extend to India; but it is
worthy of note that of the twelve Indian species of Pieris seven are
European, including our three common Whites and Pieris daplidice.
Our two Clouded Yellows are also among the nine Indian species,
the only other European Colias found in India being an Alpine
form of the Polar Colias nastes. Our Clouded Yellows stand as
C. hyale, Linn., and C. croceus, Fourcroy.
The section on Lyczenide includes much that will be read with
great interest respecting the relations between these butterflies and
ants.
The letterpress is written with great care, and it would be
difficult to overestimate the value of this book either to collectors in
the field or to home students of butterflies. The illustrations are
all by Horace Knight, and include 10 plates, excellently reproduced
in colour-photography, and 104 (often multiple) numbered figures
in the text. Wa ake
PROCEEDINGS OF LEARNED SOCIETIES.
GEOLOGICAL SOCIETY.
February 6th, 1907.—J. E. Marr, Se.D., F.R.S.,
Vice-President, in the Chair.
The following communication was read :—
‘Note on the Cervical Vertebra of Zeuglodon from the Barton
Clay of Barton Cliff (Hampshire).’ By Charles William Andrews,
B.A., D.Sc., F.R.S., F.G.S.
The Author gives a brief description of a cervical vertebra from
the Barton Clay of Barton Cliff. It is referred provisionally to
Zeuglodon Wanklyni, a species described in 1876 by Prof. H. G.
Seeley. The skull on which this description was founded is totally
lost, so that this vertebra is the only bone of a Zeuglodon from the
Barton Clay, and, with the possible exception of a vertebra from
the Brockenhurst Beds (which is the type of Balenoptera Judd),
the only one found in the British Isles that now exists.
Geological Society. 375
May 1st, 1907.—Sir Archibald Geikie, D.C.L., Sc.D., Sec.R.S.,
President, in the Chair.
The following communication was read :—
‘On the Xerophytie Character of Coal-Plants, and a Suggested
Origin of Coal-Beds.’ By the Rev. Prof. George Henslow, M.A.,
FLS., F:G:S:
Of the Paleozoic flora, the Hquisetites, now represented by the
sole genus Hquisetum, are decidedly hygrophytic, if not hydrophytic.
The Ferns, which appear to have much resemblance to certain
modern types, especially the Marattiew, seem to have lived under
more or less similar conditions to the present; that is, varying from
the hygrophytic habit of Hymenophyllum to the xerophytic Ceterach.
The Cycadofilices, Cordaites, etc. are decidedly xerophytic; and
the same is the character of the Lycopodiales, represented now by
Lycopodium and NSelaginella, and of Psilotum, Salisburia, and
others. In fact, the characteristic feature of the great coal-forests
was xerophytic, and the vegetation appears to be of an upland
type. Illustrations are given from recent and Carboniferous plants,
to show the characters of leaf, root, and stem which separate these
classes of plants. The position of coal-seams is accounted for by
the action of earth-movements in late Carboniferous times: these
threw the forest-bearing surface into shallow waves and troughs,
which became gradually accentuated, the latter being gradually
filled with sediment, upon which, during intervals of rest, new
forest-growth took place.
June 5th, 1907.—Sir Archibald Geikie, D.C.L., Se.D., Sec.R.S.,
President, in the Chair.
The following communications were read :—
1. «A Marine Fauna in the Basement-Beds of the Bristol Coal-
field” By Herbert Bolton, F.R.S.E., F.G.S.
Tsolated and rare fossils have been occasionally discovered in the
Bristol Coalfield, but the list of genera and species hitherto recorded
is ashort one. During the progress of an exploration-branch at
the Ashton-Vale Colliery, fossiliferous shales were traversed in tho
lowest Coal-Measures resting upon the Millstone Grit. A section
of the Coal-Measures in this part of the Coalfield is given, the Gays
Seam being the lowest workable seam, and the chief fossiliferous
shale is localized at a depth of 84 feet below it. The most striking
feature of the fossils is their dwarfed condition, especially among
the fish-remains, Fossils found in the spoil-banks of other pits
indicate that other marine horizons occur in the Coalfield. The
thickness of the Millstone Grit appears to be about 980 feet. The
paleontological description embodies a tabular list of fossils from
the marine horizon, which shows a close correspondence with the
list drawn up from the marine beds associated with the Gin Coal of
376 Geological Society.
North Staffordshire ; but it does not appear to be desirable to con-
clude that the horizons are identical, until further evidence of faunal
development has been obtained from the Bristol area. The brachio-
pod-fauna contains forms identical with, or closely approximating
ey species occurring in the Cyathaxonia- and Dibunophyllum-
ones,
The paleontological description contains an account of the different
species collected, including notes on Productus, Choncetes, Derbya,
and Orthothetes, contributed by Dr. A. Vaughan, and new species of
Chonetes, Raphistoma, and Lowonema.
2. ‘Brachiopod Morphology: Cincta, Eudesia, and the Develop-
ment of Ribs.” By $8. 8. Buckman, F.G.S.
The test-ornament of brachiopods is found in three main phases :
smooth, ribbed, and spinous; and of these three, a costate
species is more advanced than a smooth one, and less advanced
than a spinose one. There are catagenetic developments, also,
in reverse order; but these may, for the present, be neglected.
Dall has stated that Cincta (Terebratula-numismalis Group) is a
synonym of Hudesia (T7.-cardium Group); but, although it would be
quite possible for costate forms to be developed from the smooth
Cincta, yet they would not be costate forms of the Hudesia-type:
in the Cincta-type the coste developed would be of the kind
which produced opposite carination of both valves, while in
Eudesia the costa on the two valves are alter nate, the carine
of one valve being opposed by sulci in the other. This fundamental
difference not only prevents the inclusion of Hudesia and Cincta in
one genus, but shows that they belong to entirely different series.
The first phase of development dealt with may be called the
lenticular stage, which might develop in either of the two
directions indicated. The next phase would be the Cincta-stage,
in which the front margin is rounded in youth, truncate in
adolescence, incipiently excavate and bilobate in the adult, as the
growth-lines of the specimens show. The Cincta-stage may develop
in two directions—out of broad forms the quadrifid stage, out of
narrow forms the cornute stage. The next development may be
called the quadricarinate or trigonellid stage; and the fourth
stage, the multicarinate or pectunculus-stage. In Hudesia
there is a highly-developed multicarinate stage, but the carine are
alternate, not opposite. In degree of ribbing it is higher than
Oincta, and in a way even higher than the pectunculus-stage ; but
both the ribbing and the loop forbid connexion with Cincta. The
preceding stage is exemplified by Jsmenia pectunculoides. A prior
stage may be seen in Megerlia Munieri; and, as an example of
the incipient uniplicate stage, Terebratula Whitakert may be given.
Certain emendations in nomenclature appear to be necessitated as
a result of this communication ; new names are given, and their
application defined. A Table is added to show the successive stages
of development along the two lines.
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Page
XLIL. New South-African Spiders of the Family Drasside in the
‘Collection of the South-African Museum. By W. F. Purcett,
PhD; Cope. Town: (Plates XDA.) onset. en eae ees 297
XLII. Descriptions of Two new African Species of Barbus. By
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Pe i ea 356
BIBLIOGRAPHICAL NOTICES.
100 Photographs from Life. (No. 1.) By Dovetas Enetiso...... 372
100 Photozraphs from Life. (No. 2.) By R. B. Lopez .......... ib.
Les Débuts d’un Savant Naturaliste, le Prince de I’Entomologie,
Pierre André Latreille, 4 Brive, de 1762 41796. Par Lovis pz
Nvssac, Sous-Bibliothécaire du Museum d’Histoire naturelle .. id.
Epu. Borpacr. Recherches anatomiques et biologijues sur
YAutotomie et la Régénération chez divers Arthropodes ...... 3793
The Fauna of British India, including Ceylon and Burma. Published
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XLVIII.—Preliminary Notice of new and remarkable
Cephalopods from the South-west Coast of Ireland. By
A. L. Massy, Department of Agriculture and Technical
Instruction, Fisheries Branch, Dublin.
THE species mentioned below were all met with at 50 to 70
miles off the south-west coast of Ireland, and form part of a
collection made by the Fisheries Cruiser ‘ Helga’ during the
years 1901 to 1907. A detailed report of the whole collection,
with illustrations of the new species, is in preparation and
will appear in the “ Scientific Investigations” of the Fisheries
Branch of the Department,
Family Polypodida.
Genus Potypus, J. G. Schneider.
Polypus profundicola, sp. n.
Head slightly narrower than body. Eyes large and promi-
nent. A few slightly marked tubercles usually present round
each eye. Skin otherwise smooth in all parts. Arms some-
what slender, largest at the base, tapering gradually to very
fine points. Order 1, 2, 3, 4. Their length is on an average
six times longer than the body. Ventral arms the slightest,
the others not differing much in bulk. Suckers sessile with
circular aperture, and radial grooves extending almost to the
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 25
378 Mr. A. L. Massy on Cephalopods
margin. Suckers relatively small and placed rather far apart,
especially in the distal half of umbrella region. The space
here between twosuckers frequently twice the diameter of either
sucker. Suckers of extremities of arms perfect, not reduced to
papille, and extending to within less than half a millimetre
from tip of arm. Web much developed, extending nearly one
quarter the lengthof arms. Least development between ventral
arms. Web continued on arms in the form of large lateral
crests. Radula formed of 7 series of teeth: 1 median, 4 lateral,
and 2marginal. Dentalformula3 2 2 1 2 2 3. Median row
consisting of large acute teeth without lateral denticles. Inner
lateral teeth much the smallest, broad at the base, terminating
in short conical points. Outer lateral teeth broader and more
elongated. Marginal teeth elongated and claw-like. Siphon
moderately large. Organ of siphon consisting of a pair of
heart-shaped folds, of which the thickened rims are con-
tinuous and distinctly projecting all round. Mantle bursiform.
Pallial aperture not as large as head. Body soft. Lateral
adductor muscles in form of flat bands placed rather close
together, the anterior much the smallest, the posterior reaching
their greatest width in the portion attached to the body.
The two cartilaginous pieces in the dorsal mantle are long
and rod-like; they are widest in the anterior portion. The
ligula copulatoria * of the hectocotylized arm has from eight
to nine laminee copulatoriz in the concave interior, in the
median portion of which there is a flat-topped longitudinal
ridge. Calamus brachialis moderately short. Sperm-canal
smooth, shallow, and like a white band on surface of arm.
General colour vinous, due to an immense number of small
light and dark purple and dark red chromatophores distributed
on a white ground. Polypus profundicola appears to be very
nearly allied to Octopus ergasticus, Fischer, particularly in
the form of the hectocotylized arm. ‘The ligula copulatoria
is, however, only crossed by 6 or 7 lamin in O. ergasticus.
The hectocotylus is also larger in proportion in P. profundicola,
and the arms are longer, and with smaller and more numerous
suckers. ‘lhe web is also more developed, forming large
crests on the arms, and the siphon projects much less. The
specimens examined consist of 7 males and 7 females,
obtained in the trawl at 385-720 fath. The principal dimen-
sions of a large male and female are given below.
* The terms used in describing the different parts of the modified arm
are those suggested by P. and J. Fischer in “Diagnoses d’espéces
nouvelles de Mollusques Céphalopodes recueillis dans le cours de
VExpédition scientifique du Talisman” (1883), Journ. de Conch. 1892,
xxx. p. 297.
from the South-west Coast of Ireland. 379
Dimensions. <&.
mm.
Total length ...... Ho BDO 010 LO HO OIOR HIG Ck 582
End of body to mantle-margin ............ 103
ra x OVOM ie ers pa erence sdaiiyeh oh since Hal
Breadth’ of body) eens. cen. Mere aero ones 70
A h@ad®. O13 Pea as hate oe honk 62
Hye toledgerofsumbroliaier rss ah eacls oe ae 102
Diameter of largest sucker .........+000s 4
Length of arms * :-—
Ist on right 471 mm. Ist on left 460
2nd sand oa LaOlonn 2nd =, ~—s mutilated.
srd » 264 ,, (hect.). 3rd ,, ditto.
4th a olOn Ath 01 327
Approximate number of suckers, Ist arm on right, 254.
Dimensions. .
mm
Potaltlens th 7.0:5 «shee ase ncetaer eee . 457
Kind of body to mantle-margin ....,..... 32
ne ss OYCN Po cekachetneres aire se oaks id
Breadth of bodygny sae. 9cedacdis .ecctee se 50
Hye to. edge of umbrellay cic. acoso 6 105
Diameter of largest sucker, nearly ........ 4
Length of arms :—
Ist on right 877 mm. Ist on left 379
nde Ht she 35 ONT s 2nd ite eon
Stdery et TOO Sth) gn ele
Galies err, Pasion 4th , 290
Approximate number of suckers, 1st arm on right, 218.
Polypus Norman, sp. n.
Head rather broader than body. Eyes large and prominent,
circumference of aperture in the type 11 mm. A number of
deep wrinkles extend across the surface between eyes,
originating from dorsal side of eye-aperture. Surface other-
wise smooth.in all parts. Arms 4-sided and keeled, some-
what slender, largest at the base, tapering gradually to very
fine points, and arranged in the order 1, 2, 3, 4. Arms about
four times the length of body, the two dorsal pairs more bulky
than the rest. Suckers sessile, large, and with the arrange-
ment in the centre of arm noticed by M. Joubin in O. levis f.
Each sucker with circular aperture and numerous radial
* The arms are measured from the mouth.
+ “ Entre les ventouses, sur la face orale des bras, on voit une créte
peu saillante mais bien nette qui passe d’une ventouse de droite a celle
immédiatement supérieure 4 gauche, puis de 1a revient 4 droite et ainsi
de suite. L’ensemble de cette disposition a l’aspect d’un lacet reliant
alternativement deux rangées de boutons.” Campagnes Scientifiques du
Prince de Monaco. Fascicule xvii. (1900): Céphalopodes provenant des
campagnes de la Princesse Alice (1891-1897), par Louis Joubin.
20%
380 Mr. A. L. Massy on Cephalopods
grooves, the first two usually placed singly, remainder
arranged in pairs, the seventh or eighth pair (about centre of
web in type) usually the largest, gradually diminishing towards
tip, where they are very small but not reduced to papille. In
the type the suckers extend to within about 2° mm. of the
extremity of each arm. Web much developed, extending about
one quarter the length of the arms. Maximum development
between the dorsal pairs, least development between the
ventral arms. Web not continued on arms. Buccal mem-
brane and mandibles apparently very small, almost concealed
in the type by the basal suckers. Siphon moderately large,
extending rather more than one third of the distance between
pallial aperture and web between ventral arms. Mantle
bursiform and somewhat pointed posteriorly, its aperture
exactly the width of basal portion of head. Body firm. The
ealamus brachialis of the hectocotylized arm short and but
little projecting; ligula copulatoria narrow and pointed, the
deeply concave interior looking like a longitudinal groove,
and crossed by nine slightly defined Jamine copulatoriz.
Sperm-canal extending to umbrella-margin, quite shallow
and like a narrow gradually widening white band, smooth
except for a few faint grooves at a little distance from the
calamus brachialis. The end of hectocotylized arm when
viewed in profile much resembles that of O. polyzenia*, except
that the point of the calamus brachialis is not quite so acute.
Colour much like that of P. profundicola, but of a warmer
tone. Dorsal surface of body, head, and web the darkest ;
arms, inner and under surface much lighter, with patches of
bluish grey on web between each arm on ventral side. The
only specimen of this graceful-looking form was taken in the
sprat-net on trawl at 707-710 fath.
Dimensions.
mm.
Motal length cir tne cine cites ier soba 206
End of body to mantle-margin ............ 29
+ ae CONLTOLOINEC YC (Macnee pileers 42
Breadthiotsbody cs. cet. eie seit sek . 30
- heads. ch pumue cre nee noe eet 32
Centre of eye to lateral edge of umbrella .. 50
Length of calamus brachialis ............
* ligula copulatoria epoca e 3
Breadth of ligula copulatoria ..........-. 2
Diameter of largest sucker.........+..0+0+ 4
Siphon projects above mantle-margin ...... 15
Diameter HetweenOy.es ag iis «ios eveleyelveiisteinte 26
* Report on the Zoological Collections made in the Indo-Pacific Ocean
during the Voyage of H.M.S. ‘Alert,’ 1881-2. Mollusca (by E. A. Smith),
pl. AV. eee
from the South-west Coast of Ireland. 381
Length of arms :—
Ist on right 157 mm. 1st on left 158
PANTY | yee Aires 2nd, loz
ord gays Sor ya(hectje vard 4) S147
Ath =,,, | 1s3e"5, 4th ,,, 183
Approximate number of suckers, 1st arm on right, 144.
Family Gonatide.
Genus Gonatus, Gray, 1849.
Gonatus Fabrictt, Lichtenstein.
Nine specimens taken in the trawl at depths varying from
465-666 fath. All were examined without success for any
trace of a hectocotylus. The largest specimen measured
275 mm. in length, and the smallest 118mm. Not met with
before in British-and-Ivish waters.
Family Enoploteuthide.
Genus OcroPpoDOTEUTHIS, Riippell, 1884.
Octopodoteuthis sicula, Riippell, 1884.
A specimen taken in the trawl at 550-570 fath. This
individual agrees closely with Dr. Jatta’s description. The
tips of all the arms being mutilated, the sex cannot be
determined without injury to the specimen. ‘his species
appears never to have been met with outside the Mediterranean
before, or of such a size. It measures 117 mm. across the
fins; from end of body to mantle-margin 107 mm.; from
end of mantle-margin to bifurcation of first pair of arms
oo mm.
The siphon projects 17 mm. above the mantle-margin.
No trace exists of the roots of the tentacles, which in this
species are moulted or re-absorbed in youth.
Family Histioteuthide.
Genus HistioreuTuis, Orbigny.
FHlistioteuthis bonelliana, Férussac.
Two specimens. One with a total length of 31 mm. was
taken in the triangle net at 70-80 fath. from the surface over
soundings of about 800 fath. The other measures 129 mm.
and was taken in the sprat-net on trawl at 775-795 fath.
The smaller specimen possesses no trace of web between the
arms, or of the black elongate swellings present in the larger
specimen at the tips of all exeept the ventral arms.
Not previously recorded from British-and-Irish waters.
382 Mr. A. L. Massy on Cephalopods
Family Cranchiide.
Genus HELICOCRANCHIA, gen. nov.
Body elongated, chalice-shaped, tapering gradually to a
rounded point. Mantle-substance tough, smooth, pale, with
many small chromatophores. Fins considerable, oval, pedun-
culate, attached to end of dorsal surface of body. Eyes
sessile, large, in the form of a low cone. Arms rather long,
with keel and lateral membrane moderately developed.
Tentacles long and expanded into a club. Siphon extremely
large.
felicocranchia Pfeffert, sp. n.
Body broadest in the middle, tapering gradually, and
rounded posteriorly. Breadth of body rather less than half
the length of mantle. Surface smooth ; colour creamy white ;
dorsal surface closely freckled with dull red oblong chromato-
phores arranged irregularly ; ventral surface and sides with
eight transverse rows of chromatophores, as well as a number
of spots arranged in no particular order. Fins narrowly
pedunculate, broadly pyriform in outline, somewhat fleshy,
rather more than one fifth the length of the mantle, attached
to dorsal surface close to (in type at 1 mm. from) posterior
end of body. Mantle-margin depressed in middle line
dorsally and there joined to siphon. LHyes sessile, large;
their visual parts in the form of a low cone, of which the
posterior face is occupied by a large bluntly conical process..
‘Towards the posterior end of each eye is an oblong, soft,
white papilla, possibly the olfactory papilla. Buccal mem-
brane mutilated in the type, but apparently seven-angled.
Siphon extremely large, extending about two thirds of the
length of the ventral arms. The arms are about one third
as long as the mantle. They are unequal, the apparent order
of length being 3, 2, 1, 4 *, slender and tapering, with
transparent keel moderately developed on distal two thirds
of all, but least developed on dorsal arms. Lateral mem-
brane moderate, extending entire length of arms. Margin
usually straight. The arms appear to be quite free. The
suckers are stalked and arranged in two rows until the distal
third of arm is reached, when they suddenly become very
minute, crowded, and arranged irregularly. These minute
suckers are stalked and have a circular aperture; towards
the extreme tip they appear to be imperfectly formed. The
large suckers have a circular horny ring and about four rows
* Tips of ventral arms absent.
Strom the South-west Coast of Ireland. 383
of papille. The latter when they cross the edge look like
teeth. The tips of the ventral arms being absent in the type,
it is not possible to say if the small suckers are present on
these arms also. The suckers on all the arms are placed
furthest apart on the proximal portion, gradually becoming
placed closer together and reaching their maximum size just
before the commencement of the distal third, where they are
abruptly succeeded by the tiny suckers. The large suckers
of the dorsal arms are smaller in proportion than those of the
other arms. The tentacles are long, slender, and round.
When bent back they extend rather more than three quarters
of the length of the mantle. The stem is thickest at its base,
whence it narrows gradually but considerably, again expanding
into a club furnished with moderate swimming-crest and
lateral membrane. ‘The suckers of the club are in four rows,
of which the two median are perhaps slightly the largest.
About 60 suckers are present on each club, all with circular
horny ring, with about tour rows of papillae. About 16 pairs
of minute suckers occur on the inner surface of the stem,
placed close together near the club, becoming gradually
more distant. Apparently none are modified into fixing-pads.
There are none on the proximal portion of the stem, which
is quite smooth in the type for the last 10.mm. Organ of
siphon consisting of two quadrangular folds on sides and a
median dorsal organ; the latter is composed of a thin
plate occupied by an anterior and two lateral tubercles.
From the anterior tubercle on pressure a needle-like process
appeared, in the type extending a length of 1 mm. Similar
processes claw-like in shape are situated in each of the
lateral tubercles. The specimen was taken in the trawl at
350 fath.
Dimensions.
mm
Motalilon pth 3 patos sclenteteaecs teovders Sie od cones 80
End of body to mantle-margin dorsally .,.. 39
as oF EOP: Of CY Oxcre « ccoreret Pessis yer ¢ 4]
Breadtaol Dod yrs. veda vees ad cee « 18
A, head between eyes ............ 3
. jp) ACLOSS! CV. CSharelejoieia eiete © tale 10
Wong th, Ok fine: 2! s< «ein clasieencr tas anaes Oe aa ii
Breage, Of He 6. eenc sc acne a eras ae 7
7 (diameter) of peduncle of fin...... 2
Length of arms :—
Ist on right 11 mm. Ist on left 11
2nd; 5; LST iss 2nd , 13
SrOuaen 3 14 ,, SLOee al Le
4th ,, es 4th ,, 8
Length) of tentacle’? iovisatrvivv co viesetels . OF
384 Mr. K. Andersen on the Geographical Races
The arms were measured from the mouth, but the arm-
measurements must be considered only approximate, as, with
the exception of the first and third arms on the left, the
extreme tips are missing.
XLIX.—On the Geographical Races of the Lesser Horseshoe
Bat (Rhinolophus hipposiderus). By KNup ANDERSEN.
In a paper published two years ago in the ‘ Proceedings of
the Zoological Society of London’ (1905, 11. pp. 189-144) I
pointed out the existence of three geographical races of the
Lesser Horseshoe Bat, viz. a small southern form (2A. h.
minimus), distributed, broadly speaking, over the Mediter-
ranean Subregion, south-eastwards to Sennaar and Keren; a
large northern form (Rh. h. hipposiderus), ranging, broadly
speaking, from the extreme N.W. Himalayas (Gilgit),
through N.W. Persia and Armenia, over the whole of Central
Europe N. of the Balkans and the Alps; thirdly, a form
(Zh. h. minutus) apparently confined to England, Wales, and
Ireland. The southern differs from the northern form, I
wrote, in being in every respect smaller—in some respects,
as it seems, absolutely smaller, in others at least on an
average; I found the length of the forearm to be the most
convenient means for a ready discrimination: in minimus
34°7-38 mm., in hipposiderus 39-41°7. As to the characters
of the English form, they have no bearing on the subject-
matter of this paper, and are therefore left entirely out of
consideration here. My conclusions were based on an exam-
ination of ninety-five adult specimens, mostly from the collec-
tion of the British Museum, and obtained in localities dotted
over practically the whole area occupied by the species.
Since I wrote that paper several other specimens, partly
material added to the British Museum collection, partly
examples sent for inspection and identification by corre-
spondents here and abroad, have passed through my hands.
Every specimen has agreed precisely with the characters
pointed out by me, with the exception of three, all of which
are from a place in which I had already predicted* that both
forms or intermediate individuals would most probably be
found.
* “T have some reason to believe that in certain border districts (e. g.
S.W. Switzerland ....) the two forms occur together, perhaps side by
side, but intermediate examples 1 have never seen. They will probably
be found.” (Proc. Zool. Soc. 1905, ii, p. 141.)
of the Lesser Horseshoe Bat. 385
In the ‘Mémoires de la Société zoologique de France’
(1907, pp. 21-22) M. Charles Mottaz, Geneva, records the
results of an examination of 169 specimens of the Lesser
Horseshoe (presumably obtained at or near the place where he
lives or somewhere else in Switzerland, though this isnot stated
in his paper) as follows :— Jusqu’ici le résultat de cette étude
nous laisse perplexe et ne saurait nous convaincre de la validité
de la sous-espéce minimus. En effet, sur 169 sujets actuelle-
ment entre nos mains, mesurés et scrupuleusement étudiés,
nous avons df constater que: 1°. Ceux qui répondaient aux
caractéres du minimus ou qui pouvait étre taxés d’intermé-
diaires étaient tous, jeunes [s/c] et vieux,des g. 2°. Ceux qui
furent classés hipposideros étaient tous, jeunes [sze] et vieux,
des 2. Queconclure? Y a-t-il coincidence fortuite, ou plutét
la différence signalée serait-elle seulement une question de
sexe?.... Pour]’instant nous ne saurions enregistrer défini-
tivement le Rh. hipposideros minimus dans notre faune suisse
ni admettre la validité de cette sous-espéce tout au moins
pour ce qui concerne les représentants de notre région.” —In
afew words: M. Mottaz has come to the conclusion that what
Ihave taken to be a difference between a southern and northern
form of this bat is, in reality, at least in Switzerland, a sexual
difference only.
Although it must be granted, I think, that in committin
a mistake of such kind as suggested by M. Mottaz I should
have made myself guilty of an almost unthinkable carelessness
(a quality of which, I hope, my zoological papers do not in
other respects bear too abundant witness), and at the same
time been singularly unfortunate in having had before me
from the whole area inhabited by minimus males only, and
from the whole area of hippostderus females only ; although
M. Mottaz, in dealing with a question of s¢ze in a series of
individuals, does not give any measurements at all to enable
the reader to control the correctness of his results ; although
M. Mottaz, in discussing a question for the decision of which
it is of the highest importance to know the exact places in
which he obtained the individuals examined, does not give a
single locality (they were “ recueilli.... partout,” is the
only information given, so that it cannot even be seen with
certainty whether they were obtained “ partout”’ in Switzer-
land or “partout”’ in Europe generally) ; although, there-
fore, his two “ conclusions,” as given in his paper, are merely
postulates not supported by a single fact or figure ; although
the insertion of the word “ jeunes ” in both of his conclusions
admits of one explanation only, viz. that M. Mottaz lacks
the necessary training in dealing with questions of this
586 Mr. K. Andersen on the Geographical Races
kind (for, inasmuch as the difference between minimus
and hipposiderus is a well-marked but small difference of
size, nobody would, of course, be able to tell whether a young
individual, obtained in a region where we have all reason to
believe that both forms occur, is referable to the one rather
than to the other form),—I shall now, in the interests of the
subject, challenge his conclusions and prove, by means of
indisputable figures and facts, that M. Mottaz’s result, strange
as it is, is totally wrong as well. Strictly speaking I cannot,
of course, prove anything about the very specimens examined
by M. Mottaz, and which I have not seen, but this is also not
necessary ; if only I succeed in proving, on the basis of the
large British Museum series, the validity of the characters of
minimus and hipposiderus, and their perfect independence of
the sex of the individuals, then M. Mottaz will get nobody to
believe that what in southern as compared with northern
specimens of the Lesser Horseshoe is a well-marked racial
difference, is in Switzerland transformed into a_ sexual
difference.
Subjoined I give the locality, sex, and length of forearm of
all the full-grown examples of minimus and hipposiderus
which I have had the opportunity of examining up to this
moment. Be it noted that the measurement of the forearm
has been taken from the most backward projecting point of
the forearm to the front curve of the carpus (wing folded).
All the measurements are of perfectly adult specimens, with
one exception, mentioned below.
Rh. h. minimus.
Erythrea: Keren.—One male, young adult, type of Rh. minimus,
Heuglin: 863 mm. (The not quite consolidated condition of the
finger-joints of this individual proves it to be slightly immature ;
it is, however, in all probability full-grown; I give the measure-
ment here only because this example is the type specimen.)
Sennaar.—One female: 36°5.
Cyprus.—Five males: 34:7, 35°8, 36:0, 86°8, 87°2; one female: 37:7,
Smyrna.—One female: 37°5.
Crete.—One male: 35:5.
Malta.—F our males: 36:0, 86:2, 86°83, 36°90; two females: 36:0, 36:0;
two unsexed: 36:0, 37:0,
Sicily: Ficuzza.—One male: 385°7; four females: 36:2, 36:2, 36:8,
36'8
Middle Italy: Ostia.—Two unsexed : 35°7, 36°8.
Sardinia: near Siliqua.x—One male: 36°7; one female: 37°8.
Corsica.—Three males: 37°8, 37:8, 38:0; one female: 37-7.
Balearics.—Four males: 36:2, 86°6, 36:8, 36:9; three females: 36:8,
37°6, 387°6.
Spain: Seville—One male: 37:7.
Portugal: Cintra.—One male: 36:2.
of the Lesser Horseshoe Bat. 387
Morocco: Tangiers.—One female : 37:7.
S.W. Switzerland: St. Cergues.—One male: 37:7.
N. Switzerland: near Baar.—One male: 38:0.
Intermediate individuals (nearest to mznimus).
S.W. Switzerland : Geneva.—Two males: 38:2, 38°7. (In one of these
specimens (no. 6. 2. 6,3; forearm 38°7) the right elbow is damaged,
and I do not feel quite satisfied that the same is not the case
with the left elbow.— A third Geneva specimen in the collection,
also a male, is slightly immature, but probably full-grown ; its
forearm measures 38°2.)
Rh, h. hipposiderus.
Gilgit.—One male: 39°8.
N.W. Persia: Urmi.—One male: 89:8.
Armenia: Van.—One male: 39:2; one female: 39:3.
S. Caucasus.—Two males: 39:0, 389°8; one female: 38:7.
[Cyprus.—One female : 39-6 *,]
N. Bulgaria: Rustshuk.—One male: 39:0,
Roumauia.—Five males: 39:0, 39°5, 59:8, 40:0, 41:0; eight females:
39°5, 39°8, 40:0, 40:0, 40:0, 41:0, 41:0, 41:2. ;
Transsilvania: Kronstadt.—Two females: 40:0, 41-0.
8. Carpathians.—One male: 39°83.
Hungary: Offner Mts.—One female: 41:7.
Bavaria: Burgheim.—Two males: 39-0, 40:0; one female: 40-0.
Schlangenbad.—One male: 40:0; one female: 40:0.
N. Switzerland: Thurgau.u—One male: 40:2; three females: 40-0,
40°6, 41:7,
S.W. Switzerland: St. Maurice.—One unsexed: 41:3.
The above details nay be summed up in tabular form as
follows :—
Specimens | Min, | Max. |Average.
mm. mm, mi.
TRETITUS LAMLESI cies « aici hare « 24 347 38:0 30°7
| kipposiderus, males ......+: ir MG 39°0 41:0 39°7
minimus, females .......... | 15 560 378 37°0
| hipposiderus, females ...... 19 38'7 41°7 40°83
| minimus, all examplest .... 43 84:7 38:0 | 86-7
_ hipposiderus, all examplest.. 36 38'7 41-7 40:0
% On this specimen, see Proc. Zool. Soc. 1905, ii. p. 142, footnote f,
+ Including also the few unsexed specimens enumerated above.
388 On the Lesser Horseshoe Bat.
These figures need nocomment. It is a well-known fact
that in many bats females average a trifle larger than males,
but so small is the average difference in size between
the sexes of this species (0°3 mm. in minzmus in favour of
the females, 0°6 mm. in hipposiderus in favour of the same
sex) that it is scarcely detectable except on careful measuring
of a tolerably large series.
As already emphasized in my paper two years ago, there
are certain border districts in which the two races meet and,
to a certain extent, merge into one another. As yet we know
exceedingly little about the exact limits of this transitional
zone, but the detailed lists of specimens examined and
measured, as given above, may, perhaps, throw a little light
on the question. It will be noted that of the five full-grown
specimens | have seen from 8.W. Switzerland (St. Maurice,
St. Cergues, and Geneva), one is indistinguishable from
minimus, one decidedly a hipposiderus, whereas three (all
from Geneva) are rather intermediate but nearest to minimus ;
of the five full-grown specimens from N. Switzerland (Baar,
Thurgau), one is indistinguishable from minimus though
having the maximum size of this form, whereas four are
decidedly hippostderus ; the series is much too small to allow
of any safe conclusions, but, so far as it goes, it shows
that in S.W. Switzerland both forms occur as well as
intermediate examples, in N. Switzerland hipposiderus, as
might be expected, is the dominant form (four of five
examples), whereas a small percentage (one of five) is refer-
able to minimus. Although, as already said, derived from a
very small series of specimens, this result will probably prove
to be approximately correct, and it is so far from being
unexpected that it is, on the contrary, entirely in accordance
with what I could predict without hesitation two years ago.
—'l'o this I can now add one fact more: not only does
minimus go a certain distance northward into the area of
hipposiderus, becoming probably rarer the farther north, but
I know on excellent authority * that Aipposiderus goes a
* Dr, Senna, Florence, writes (translation from letter, Dec. 19, 1905) :—
“ You have pointed out that Aipposiderus ranges (so far as our continent
is concerned) over Central Europe N. of the Balkans and the Alps,
minimus over the Mediterranean Subregion, This seems to be perfectly
true, generally speaking; I find, for instance, that nine specimens from
Cyprus are decidedly menimus, several examples from 8. Italy (Calabria,
Sicily) are, without exception, munimus; but tm middle and northern
Italy we begin to meet with hipposiderus, about 15 per cent. of the indi-
viduals belonging to this form, as against 85 per cent. of minimus ; still
farther north, as you say, we find Acpposiderus..... I hope I shall
eet so much spare time that I can work out the range of these forms in
Italy on the basis of the collections preserved in all, or most, of the
Italian Museums.”
On new European Insectivora and Carnivora. 389
certain distance southward into the area in which ménimus is
the dominant form. But all this is precisely what we must
expect, in view of the fact that these races do not occupy
isolated (insular) localities, but the central and southern part
of one continuous land area.—Very likely there is also in
W. Asia a certain region in which both forms or intermediate
examples occur (see the female from 8. Caucasus, which I
have referred to hippostderus, but which has the minimum
size of this race).
From France and the whole of the Balkan Peninsula 8. of
Rustshuk we completely lack information; it would be parti-
cularly interesting to know whether French specimens are
hipposiderus or minimus, or, possibly, identical with the
British form, minutus.
I should not have taken the trouble to give the proofs—
once more, and in a much more detailed form—of the exist-
ence in continental Kurope of two well-marked races of
the Lesser Horseshoe Bat were it not for the following
reason :—It is a matter of course that on the basis of the
collection in one single Museum—be such collection even so
rich as that of the British Museum—it is impossible to give
more than a rough sketch of the range of these two races of
Th. hipposiderus; the working out of the details must be
left to the local naturalists interested in the subject. But
the stimulus to do such useful work is naturally taken away,
or greatly weakened, when a writer, claiming to base his
conclusion on a careful examination of an unusually large
series of specimens, declares that he cannot see that the
supposed racial difference is anything but a difference
between male and female of the same species. To show that
this opinion is entirely wrong is the object of these lines.
Naturalists may safely take it as an established fact that
these two races do exist; what we want to know now is,
(1) the exact area occupied exclusively by the one or the other
form, and (2) the area where both of them occur together.
This latter is the transitional zone between the regions
inhabited by the two races.
L.—Some new European Insectivora and Carnivora.
By Gerrit 8. MILuer.
In the course of some studies of the European mammal
fauna, undertaken at the invitation of Mr. Oldfield Thomas,
I have found the following hitherto unnamed Insectivora and
Carnivora.
390 Mr. G. S. Miller on new
Crocidura russula cintre, subsp. n.
Type.—Adult male (skin and skull), B.M. no. 98. 2.2.11.
Collected at Cintra, near Lisbon, Portugal, January 26, 1896,
by Oldfield Thomas. Original number 47.
Diagnosis.—Smaller than true Crocidura russula (hind
foot 11:4—12°2 mm. instead of 11:7-14; condylo-basal length
of skull 17:8-19:2 instead of 18-20°4); colour dark and
rich, in striking contrast with the pallid tints of the Spanish
C. r. pulchra, Cabrera, and with a strong coppery lustre rarely
indicated in the typical race.
Colour.—Type: upperparts between the mars-brown and
russet of Ridgway, very faintly darker along middle of back,
the hairs everywhere with metallic coppery and silvery
reflections. Underparts and feet pale wood-brown. ‘Tail a
dull indefinite broccoli-brown, darker above than below.
Skull and teeth— Except for their smaller size the skuil
and teeth resemble those of true russula.
Measurements.—T ype: head and body 64 mm. ; tail 33;
hind foot 11:4; ear 8°6: skull, condylo-basal length 18 ;
width of brain-case 9; depth of brain-case 4°8 ; mandible
(including incisors) 11:8 ; upper tooth-row 8°6.
Specimens examtned.—Kleven, all from the type locality.
Remarks.—In its small size the Cintra shrew agrees with
the Spanish race, but the colour is conspicuously darker.
The skins show no specially noteworthy variations, though
in a few individuals the fur is less glossy than usual. ‘Taken
as a whole the series is about as dark as in French and
Belgian russula; but the peculiar coppery lustre is highly
characteristic of the Portuguese form.
Crocidura cyrnensis, sp. n.
Type-—Adult male (in alcohol). B.M. no, 6.3.14. 1.
Bastia, Corsica. Collected and presented by E. R. Southwell,
Ksq.
en onic: than the Sicilian Crocidura caudata
(hind foot 12-12°4 mm. instead of 14 mm.) ; tail relatively
about as long as in cawdata, but not unusually thickened (its
ratio to head and body about 70, its diameter at middle 2 mm.
instead of 3 mm.).
Colour.—Upperparts broccoli-brown slightly washed with
sepia, the slate-grey bases of the hairs showing through at
surface and producing a general effect nearly the drab of
Ridgway. Most of the hairs with faint silvery reflections in
certain lights. Underparts and feet light smoke-grey.
European Insectivora and Carnivora. 391
Skull and teeth.—No perfect skull has been seen, but there
are apparently no special cranial peculiarities. Teeth
essentially as in C. russula.
Measurements.—Type: head and body 67 mm. ; tail 48 ;
hind foot 124; ear 10; skull, from front of incisor to back
of glenoid fossa 11:6; mandible (including incisor) 11°6;
upper tooth-row 8.
Specimens examined.—T wo, the type, and a skin from La
Foce de Vizzavora, presented by Col. J. W. Yerbury.
Crocidura balearica, sp. n.
1901. Crocidura russulu, Thomas, Proc. Zool. Soc. London, i. p. 59.
Type.—Adult female (skin and skull), B.M. no. 0. 7. 1. 42.
Collected at San Cristobal, Minorca, Balearic Islands,
April 7, 1900, by Oldfield Thomas and R. I. Pocock.
Original number 263.
Diagnosis.—Similar to Crocidura cyrnensis, but teeth much
smaller.
Colour.—Type: above hair-brown tinged with sepia, the
hairs nearly without metallic reflections ; below dull smoke-
grey. Feet and tail a dull, indefinite brownish, the tail
scarcely lighter below.
Skull and teeth.—Except for the smaller teeth there appear
to be no cranial or dental peculiarities to distinguish the
species from C. cyrnensis.
Measurements.—Type: head and body 42 mm.; tail 45;
hind foot 12°5; ear 9; distance from front of upper incisor
to back of second molar 7-4.
Specimens examined.—Three, all from the type locality.
Remarks.—This species, C. cyrnensis, C. caudata, and C. cy-
prie (Bate) form a group, of which no continental member
is yet known, characterized, as compared with C. russula,
C. leucodon, and C. mimula, by the noticeable elongation of
the tail.
Vulpes ichnuse, sp. n.
. Type.—Adult male (skin and skull). B.M. no. 88. 12.1.2.
Collected at Sarrabus, Sardinia, February 26, 1885, by
G. B. Travers, and presented by the Marquis G. Doria.
Diagnosis.—Size less than in any of the known continental
members of the Vulpes vulpes group; both hind foot and
condylo-basal length of skull less than 130 mm, in adult male,
ear from crown only 60-70 mm., colour rather dark.
Colour.—F ace and head dark rufous, becoming lighter and
more dull on base of ears and on neck, and fading to
392 Mr. G. S. Miller on new
ochraceous-rufous on shoulders and back. Sides of neck,
outer surface of upper arm, and region just behind axille a
tawny buff. Underfur of back drab-grey at base, tawny
clay-colour at tip. Longer hairs of head, sides, and back
(behind shoulders) much speckled by the presence of a buffy
white subterminal area (about 5 mm. long) on each hair;
extreme tips reddish. Feet and legs ochraceous-rufous,
slightly clouded with blackish and a little speckled with buffy
white. Tail like back at base, the ochraceous-rufous gradually
fading through a buffy grey to the whitish buff tip, the
longer hairs everywhere except at tip with 30-40 mm. black
terminal area. Underparts in front of fore legs buffy white
tinged with hair-brown, the latter becoming nearly clear on
middle of throat. Rest of underparts a mixture of hair-
brown and dull tawny, the latter predcminating laterally.
Skull and teeth—Except for their smaller size the skull and
teeth are essentially as in Vulpes vulpes.
Measurements.—Type: hind foot 123 mm.; ear from
crown 70: skull, condylo-basal length 129; zygomatic
breadth 78; mastoid breadth 47; rostral breadth over
canines 22; depth of brain-case 41; frontal depth at last
molar 33; rostral depth behind canine 17; mandible 100°4 ;
maxillary tooth-row 59; mandibular tooth-row 65°4.
Specimens examined.—T wo, both from Sardinia.
Remarks.—While the Sardinian fox is readily distin-
guisbable from its continental allies by its small size, it
closely agrees in this respect with the small fox of Crete.
It retains, however, the usual dark, bright coloration of the
ordinary European animals, while in the Cretan fox the
rufous parts are faded to ochraceous buff.
Vulpes indutus, sp. n.
1904. Vulpes vulpes, Bate, Proc, Zool. Soc. London, 1908, ii, p. 345
(April 1, 1904).
Type.—Adult (skin only). B.M. no. 3, 12.4. 25. Cape
Pyla, Cyprus (Miss D. M. A. Bate).
Diagnosis.—Similar to the small Vulpes tchnuse of
Sardinia, but general colour paler (face ochraceous-buff
instead of dark rufous, sides dull yellowish buff instead of
tawny buff) and legs grizzled blackish in strong contrast
with colour of sides.
Colour.—F ace and head ochraceous-buff, becoming lighter
and more buffy on base of ears and fading to dull yellowish
buff on neck and body. Underfur of back a slaty grey at
base, becoming nearly russet at tips of hairs, this darker
European Insectivora and Carnivora. 393
colour appearing at surface from between ears to base of tail
and over thighs and shoulders. Longer hairs of head, sides,
and back with broad subterminal light area, buffy on neck
and back, dull whitish on flanks and sides, the extreme tips
blackish. Legs a grizzle of blackish and whitish in strong
contrast with colour of sides, the inner surface washed with
dull ochraceous. Feet blackish, the hind foot much suffused
with dull ochraceous. ‘Tail buffy clay-colour, the tip nearly
white, most of the hairs, except at tip, black terminally, and
those of underside with a whitish subterminal area. Under-
parts to fore legs buffy white, clouded by the dark underfur
and becoming buff on inter-ramia and dusky on tips. Behind
fore legs the clouded whitish extends as a narrow median
area between the clear dull buff of lateral portions of under-
arts.
; Skull and teeth—The skull is conspicuously smaller than
that of Vu/pes vulpes, in this respect agreeing with that of
V. ichnuse. It resembles the latter also in its greater relative
breadth as compared with skulls of mainland foxes. The
teeth show no special peculiarities.
Measurements.—Type: ear from crown 62 mm. (in two
other specimens 65 and68). Skull of adult (probably male),
from type locality, No. 3. 12. 4. 26: condylo-basal length 123;
zygomatic breadth 73:4; mastoid breadth 42°6; rostral
breadth over canines 22°6; depth of brain-case 388; depth
behind last molar 31°6 ; rostral depth behind canine 16:4;
mandible 97 ; maxillary tooth-row exclusive of incisors 55:4 ;
mandibular tooth-row exclusive of incisors 63°4.
Vulpes vulpes stlaceus, subsp. n.
Type.—Adult male (skin and skull), collected near Silos,
Province of Burgos, Spain, January 1907, by the Rev. Father
Saturio Gonzalez. Original number 1.
Diagnosis.—Like Vulpes vulpes vulpes of Central Europe,
but with the reddish tints mostly replaced by buffy and
greyish.
Colour.—Head and face tawny-ochraceous, becoming a
yellowish ochraceous-buff behind ears and brightening to a
dull rufous about eye and between eye and base of whiskers ;
upper surface of muzzle ochraceous-buff ; forehead from level
of front of eyes, and cheeks behind eyes, strongly suffused with
creamy white, this becoming less evident between ears and
disappearing entirely on ochraceous-buff area behind ear ;
hairs of outer margin and inner surface of ear pale cream-
buff, those of inner margin light buff; back of ear with the
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 26
394 Mr. G. §. Miller on new
usual blackish area; back a coarse mixture of black, creamy
white, and russet, the black predominating along middle of
neck and between shoulders, the russet along median line of
back; on sides the black nearly disappears, and the russet
changes to a light ochraceous-buff which becomes nearly clear
(that is, scarcely overlaid with creamy white) around base of
fore leg and on shoulder nearly to median line of back ; legs
a tawny ochraceous-buff noticeably darker than that of ‘sides,
the hairs on outer surface rather conspicuously black-tipped ;
feet somewhat yellower than legs, the upper surface strongly
washed with black; tail a light buffy grey, tinged with a
tawny ochraceous-buff like that of legs above, “whitish at
extremity, the longer hairs everywhere (except at tip) with
terminal 20-30 mm. black, the dark clouding thus produced
most noticeable on lower surface ; underparts dull whitish,
everywhere clouded with slaty black, this most conspicuous
on throat and along middle of chest.
Skull and teeth.—Kssentially as in the fox of Central
Europe.
Measurements.—Type: head and body 750 mm.; tail-ver-
tebre 370; hind foot 150: skull, condylo-basal iene th 143 ;
zygomatic breadth 78; mastoid breadth 48° 8; rostral breadth
over canines 24 ; depth of brain-case 41; depth behind last
molar 35°4 ; rostral depth behind eanine 17° 6; mandible 99 ;
maxillary tooth-row, exclusive of incisors, 65; mandibular
tooth-row, exclusive of incisors, 73.
Specimens examined.—Nine, from the following localities
in Spain: Province of Vitoria, Arrechavaleta, 1; Province
of Burgos, Silos, 1; Palacios,1; vicinity of Burgos, 1;
Galicia, Forreo del Allo, 1; Province af Seville, vicinity of
Seville, 3; Province of Alicante, Elche, 1
Meles arcalus, sp. n.
1899. Meles meles mediterraneus, Barrett- eis Ann. & Mace. Nat.
Hist. (7) iv. p. 131 (November 1899) (part.).
1906. Meles meles mediterraneus, Bate, P. TAS: 1905, ii. p. 318
(April 5, 1906).
Type.—Y oung female (skin and skull). B.M. no, 5.12.2.17.
Collected on the Lassethe Plain, Crete, by Miss D. M. A.
Bate. Original number 25.
Diagnosis.—Smaller and paler than in the ordinary
European badgers (upper length of skull in adult less than
110 mm.; maxillary tooth-row, exclusive of incisors, about
35 mm.) ; audital bulle not flattened, their form as in the
Caucasian Meles minor (Satunin).
European Insectivora and Carnivora. 395
Colour.—The colour so closely resembles that of the
common badger as to need no detailed description. It is,
however, somewhat paler than in true Me/es meles, agreeing
in this respect with the Spanish M. m. mediterraneus.
Skull.—In. general form the skull appears to agree with that
of Meles meles (no perfect adult skulls examined), though it
is readily distinguishable by its smaller size (upper length
in adult 107 instead of 124-137 mm.) and by the form of the
audital bullae. These are strongly inflated, the highest region
near middle of bulla proper (exclusive of meatal tube) and so
broadly rounded as to show no longitudinal ridge, the region
between highest portion and meatus not noticeably concave.
Their form resembles that in the much larger Meles minor
(Satunin) as figured in the original description * and as
represented by specimens collected by Mr. A. Robert in the
neighbourhood of Trebizond.
Teeth—The teeth are smaller than those of Meles meles,
and the smaller cusps tend to be better developed, but other-
wise they show no special peculiarities except that the
postero-external border of the upper molar appears to be more
strongly concave than usual in JZ. meles. ‘he material at
hand is, however, not sufficient to show whether this character
is constant.
_ Measurements—Type: hind foot 80 mm.; ear from
crown 17 (both from dry skin): skull, condylo-basal length
100 ; upper length 105 (107) +; distance from anterior rim
of orbit to gnathion 386 (38°6); zygomatic breadth 55°6 ;
mastoid breadth 48°6 ; depth of brain-case at front of basi-
occipital 36; mandible 68; maxillary tooth-row, exclusive
of incisors, 36 (36°8) ; mandibular tooth-row, exclusive of
incisors, 42°6; mandibular molars (crowns) 2074.
Specimens examined.—Three, all from Crete.
Putorius erminea ricine, subsp. n.
1904. Putorius erminea, Barrett-Hamilton, Annals of Scottish
Natural History, p. 203 (October 1904).
Type.—Adult male (skin and skull). B.M. no. 7. 10.19. 1.
Collected at Islay House, Island of Islay, Scotland,
February 6, 1896, by P. Mackenzie. Presented by Hugh
Morrison, Hsq.
Diagnosis.—Size less than in Putortus erminea stabilis of
Southern England (hind foot in adult females 35 to 40 mm.
* Mitteilungen des Kaukasischen Museums, ii. (1905), pl. i. 1906.
+ Measurements in parentheses are those of an imperfect adult skull
without exact locality.
26*
396 Mr. G. S. Miller on new
instead of 38 to 44 mm.) ; skull with zygomatic arches very
wide-spreading, the ratio of zygomatic breadth to condylo-
basal length ranging from 58 to 60 instead of from 53
to 57.
Measurements.—Type: head and body 254 mm.; tail
105; hind foot 43: skull, condylo-basal length 47; zygo-
matic breadth 24:2; mastoid breadth 23:2 ; occipital depth to
lip of foramen magnum 13; frontal depth behind tooth-row
13°2; mandible 26°38; maxillary tooth-row, exclusive of
incisors, 12 ; mandibular tooth-row, exclusive of incisors, 15.
Specimens examined.—Seven from Islay and six from Jura,
the latter kindly placed at my disposal by Mr. Harmer, of
the Cambridge Museum.
Felis grampia, sp. n.
Type.—Young adult male (skin and skull). B.M. no.
4.1. 25.3. Invermoriston District, Inverness, Scotland,
January 16,1904. Presented by A. H. Cocks, Esq. Original
number 60.
Diagnosis.—Similar to Felis silvestris, Schreber, but darker
(the general effect of back and sides broccoli-brown instead
of smoke-grey) and with black markings more extensive and
better defined.
Colour.—Underfur of back and sides a light ochraceous-
buff, the basal half of the hairs mouse-grey. Light annu-
lations of longer hairs very nearly the cream-buff of
Ridgway. Black tips to longer hairs more noticeable than
in Felis silvestris, and general effect of ground-colour dis-
tinctly browner and with no trace of the frosted appearance
often very noticeable in the continental animal. Upperside
of feet and inner surface of hind legs ochraceous-buff,
becoming duller and somewhat drab-tinged on underside of
body. Pectoral and intercrural white areas well defined and
strongly contrasted with surrounding colour. Black spotting
on middle of chest conspicuous. Soles and palms blackish.
Dark markings on tail, legs, and upperparts similar to those
of Felis silvestris in arrangement, but more definite in outline,
particularly the transverse stripes on outer side of fore legs
and those on posterior half of body, the latter nearly always
appearing as definite stripes.
Skull and teeth.—Asg in Felis silvestris.
Measurements.—T ype: head and body 534 mm. ; tail 338 ;
hind foot 127: skull, condylo-basal length 87; zygomatic
breadth 66; mastoid breadth 43°8; postorbital constriction
34°6; interorbital constriction 18°6; breadth of rostrum over
European Insectivora and Carnivora. 397
canines 25; depth of brain-case 37; frontal depth behind
tooth-row 31:4 ; rostral depth behind canine 16; mandible
63; maxillary tooth-row, exclusive of incisors, 29°6 ; mandi-
bular tooth-row, exclusive of incisors, 32°4.
Specimens examined.—KHleven, all from Scotland.
Felis tartessia, sp. n.
Type.—Adult male (skin and skull). B.M. no. 7. 6. 4. 1.
Coto Dofiana, near Jerez de la Frontera, Spain. Collected
and presented by B. F. Buck, Esq.
DPiagnosis—Larger and darker than Felis silvestris and
with conspicuously larger teeth.
Colour.—The colour is noticeably darker than in Felis
silvestris, scarcely or not distinguishable from that of
F. grampius. Underfur more slaty at base than in the two
related species, about the grey No. 6 of Ridgway, its terminal
portion a dull cream-buff. Pale annulations of longer hairs
nearly as light as in J. silvestris. Inner surface of hind legs
a light ochraceous-buff ; rest of underparts as in /. grampius.
Dark markings well defined, their arrangement and extent as
in the British Wild-cat.
Skull.—In fully adult males the skull is very large,
apparently exceeding that of any of the other members of the
group. ‘The form is not peculiar.
Teeth—While in form the teeth show no peculiarities their
size immediately distinguishes them from those of other
members of the group, including the domestic cat. This is
particularly noticeable in the premolars both above and below.
In the three skulls examined (two males and one female) the
length of the posterior two upper premolars together is
19°8 mm., while in fifteen skulls of / stlvestris and /. grampius
it ranges from 16°6 to 18:8. The combined length of the
three lower cheek-teeth in the three Spanish cats ranges from
23°4 to 23°6, while in the fifteen northern specimens the
extremes are 18°8 and 21°2.
Measurements.—Vhe specimens were not measured in the
flesh, but the hind foot and tail both appear to be longer than
in the northern forms. Skull of type: condylo-basal length
93+ mm.; zygomatic breadth 764; mastoid breadth 47:2 ;
postorbital constriction 33; interorbital constriction 21°6 ;
breadth of rostrum over canines 29; frontal depth behind tooth-
row 333 rostral depth behind canine 20°4; mandible 69 ;
maxillary tooth-row, exclusive of incisors, 33; mandibular
tooth-row, exclusive of incisors, 3d°4.
Specimens examined.—Three, the type and two others
(probably from the type locality).
os
(Lo)
oQ
Mr. G. S. Miller on
Lynx pardella, nom. nov.
1824, Felis pardina, Temminck, Monogr. de Mamm. i, p, 116. Not
Lynx pardina, Oken, 1816,
Type.—Adult female (skin and skull). B.M. no. 4, 12. 12. 2.
Coto Dojiana, near Jerez de la Frontera, Spain. Collected
and presented by Abel Chapman, Esq.
In applying the well-known name Lynx pardina to the
Spanish lynx the fact has been overlooked that Temminck
took his specific name from Oken, or at least that he supposed
his animal to be the same as that of the earlier author.
Oken’s Lynx pardina was a striped cat from “Turkey and
Barbary ” *, so that, whatever the true identity of the species,
it cannot have been the spotted lynx of Spain.
The material in the British Museum shows that two colour-
patterns oceur among Spanish lynxes, in one of which,
apparently the more usual, the spots on the back and sides
are small, mostly about 10 mm. or less in diameter, the rows
indistinet, but containing evidently more than 25. spots
between shoulder and base of tail; while in the other the
spots are larger and more distinct, many of them 20 mm. in
diameter, the rows containing only about a dozen spots
between shoulder and base of tail. From the skins at hand
it is impossible to determine the status of these two forms,
though the similarity of their skulls indicates that they are
merely colour-phases of a single species. ‘To avoid any
possible ambiguity I have designated a type specimen for
this new name, and have selected for this purpose a skin
showing the better-known, small-spotted type of coloration,
‘Temminck’s animal came from the neighbourhood of Lisbon,
Portugal, but to which of the two phases it belonged the
description gives no clue.
LL—Two new Forms of the Spanish Hare.
By Gerrit 8. MILLER.
THE series of fifteen specimens of the Spanish Hare in the
British Museum shows that this strikingly characterized
species } is represented by three readily distinguishable forms,
which may be briefly defined as follows :—
* “In der Titrkei und Barbarei....rothbraun, Bauch falb, Gurgel
weiss, ttberall voll schwarzer Streifen oben, Flecken unten, auf Ohren
solche Querstreifen.” (Oken, Lehrbuch der Zoologie, iii. Th. ii, Abth.,
» 1051.) :
: + Fora full discussion of the status of the Spanish Hare, see de Winton,
Ann, & Mag, Nat. Hist. ser. 7, i. p. 163 (February 1898). The name
new Forms of the Spanish Hare.. 399
Lepus granatensis granatensis, Rosenhauer.
Type locality.—Vicinity of Granada, Spain.
Geographical distribution—The greater part of Spain,
extending at least from the Province of Burgos in the north
to the entire Mediterranean coast; Balearic Islands. Exact
limits of range not known.
Diagnosis.—Ear long, its height from crown in dried
specimens 105-115 mm.; general coloration pallid, the
eround-colour of back nearly the cream-colour of Ridgway
and much in excess of black.
Colour.—Underfur (17 mm.) bluish grey at base, the hairs
becoming buffy terminally, their extreme tips frequently
darker, though never definitely blackish; longer hairs
(30 mm.) greyish at base, then black to tip, each with a
conspicuous, sharply defined, subterminal annulation (4 mm.)
of pale cream-buff; clear area along sides (ill defined) be-
tween cinnamon-rufous and ochraceous-rufous, becoming
somewhat less bright on inguinal patches and darker and
more intense on outer surface of thighs; outer surface of fore
leg a duller shade of the same colour, the inner surface white
to wrist, where the white crosses to outer side and forms a
conspicuous patch on metacarpals. Collar pale wood-brown,
strongly washed with pale, dull, ochraceous-buff. Chin
slightly clouded with dark brown.
Skull and teeth.—In the typical form of Lepus granatensis
the skull is of maximum size for the species, the occipito-
nasal length in full-grown individuals ranging about from
895 to 90 mm.
Measurements.—Adult from Granada: hind foot 115 mm. ;
ear from crown 120. Adult from Seville (type of Lilfordc) :
hind foot 116; ear from crown 120. Adult male from Silos,
Province of Burgos (no. 7384 G. 8S. M.): head and body 520;
tail-vertebree 87; hind foot 115; ear from crown 120 *.
Skull of adult male from Seville (type of Lilfordi) and of
adult male from Silos, Province of Burgos (no. 7384 G.8. M.),
those of latter in parentheses: occipito-nasal length 88°6 (88) ;
condylo-basal length 77°6 (78) ; zygomatic breadth 42°8 (42) ;
interorbital breadth 19 (18°8) ; postorbital breadth 12°4 (13-4) ;
breailth of brain-case 31°8 (29°6); nasal (diagonal) 37°4 (39°6) ;
greatest breadth of both nasals together 20 (18°8); diastema
Lepus Lilfordi applied to the animal by Mr. de Winton has been shown
by Hilzheimer (Zool. Anzeiger, xxx. p. 512, August 14, 1904) to be ante-
dated by the Lepus granatensis of Rosenhauer (¢ Die Thiere Andalusiens,’
p- 3, 1856).
* In the flesh the hind foot and ear of this specimen measured 118 and
140 mm. respectively.
409 Mr. G. S. Miller on
24:4 (25°6); mandible 67 (65°6); maxillary tooth-row
(alveoli) 16°2 (15); mandibular tooth-row (alveoli) 17°6 (15°8).
pectmens examined.—Iwelve, from the following lo-
ealities:—Silos, Province of Burgos, 2; Castrillo de la
Reina, Province of Burgos, 2; Seville, 4; Las Marismas,
Province of Seville, 1; Granada, ie Selva, Majorea,
Balearic Islands, 1; no exact locality, 1.
ftemarks.—The typical form of the Spanish hare is at once
recognizable by its long ears and pallid coloration. The
specimen from Majorca appears to belong to this subspecies,
though its hind foot and ear are both rather shorter than
usual (108 mm.). It is in a bleached abraded pelage that
evidently gives no idea of the animal’s normal appearance,
though its colour is matched by. that of a skin in similar
condition from Seville.
Lepus granatensis gallecius, subsp. n.
Type.—Adult male (skinand skull). B.M. no. 94.2. 16.1.
Collected at La Coruiia, Province of Corufia, Spain, January
28, 1894, by Dr. V. L. Seoane.
Geogr aphicat distribution.—At present this form is known
from the extreme north-west corner of Spain only.
Diagnosis.—Similar to Lepus granatensis granatensis, but
colour throughout dark and rich, the ground-colour of back
nearly the ochraceous-buff of Ridgway, and scarcely in excess
of black ; white markings on fore leg not so extensive as in
the typical form.
Colour.—Underfur with buffy portion distinctly brighter
than in Lepus granatensis granatensis, and extreme tips of the
hairs becoming definitely black. Subterminal annulations of
longer hairs light ochreous-buff. Clear area along sides dull
cinnamon-rufous, the inguinal patches similar, but outer
surface of thigh and of front leg distinetly darker and
brighter. Inner surface of front leg with white either ex-
tending to wrist or not beyond elbow, the usual white patch
on metacarpals present in either case, though slightly less
extensive than in LZ. granatensis granatensis. Collar a dark
wood-brown washed with yellowish clay-colour. Head and
ears noticeably darker than in the typical form, the eye-ring
thrown into rather strong relief. A conspicuous, dull, cinna-
mon-rufous patch at base of whiskers. Chin heavily clouded
with dusky, in strong contrast with surrounding parts.
Skull and teeth—As in Lepus granatensts granatensis.
Measurements—Type: hind foot 107 mm.; ear from
crown 105. A second specimen (no. 82. 12. 8.1), Vigo,
new Forms of the Spanish [Tare 401
Province of Pontevedra, measures: hind foot 110; ear from
crown 105.
Specimens examined. —Two, both from North-western
Spain.
Lepus granatensis iturissius, subsp. n.
Type.—Adult (skin and skull). B.M. no. 97. 6. 15. 1.
Collected in the Basses Pyrénées by G. Greig *. Presented
by J. E. Harting.
Geographical distribution.—Basses Pyrénées. Limits of
range unknown.
Diagnosis.—Colour essentially as in Lepus granatensis
granatensis; ear short, its length from crown only 95 mm. ;
skull small and slender, its occipito-nasal length about 82 mm.
instead of 85-90 mm.
Colour. —TVhe colour so exactly agrees with that of Lepus
granatensis granatensis as to require no special description.
Skull and teeth—The skull differs trom that of Lepus
granatensis granatensis in its distinctly smaller size, in the
even less convex dorsal profile, and the more slender rostrum.
The rostral depth at front of tooth-row is 17 mm., while in
LL. granatensis granatensis it ranges from 19 to 21 mm.
Other cranial characters as well as teeth as in the typical
form.
Measurements.—Hind foot 110 mm.; ear from crown 95,
Skull : occipito-nasal length 82°4 ; condylo-basal length 72°6;
zygomatic breadth 39-4; interorbital breadth 16 ; postorbital
breadth 13; breadth of brain-case 29°8; nasal (diagonal) 36:4 ;
greatest breadth of both nasals together 18°8; diastema 22:6 ;
mandible 61 ; maxillary tooth-row (alveoli) 14°8; mandibular
tooth-row (alveoli) 15-8.
Specimen examined.— The type.
Remarks.—Vhis strikingly marked form, which may prove
to be specifically distinct trom Lepus granatensis, is imme-
diately recognizable by its short ear and small slender skull.
Its range is probably confined to the region of the Basses
Pyrénées ; at least, the Central Pyrenean hare, as described
by Hilzheimer f, is a very different animal, a large member
of the Lepus europeus group.
* Probably bought in market at Biarritz (see note by Mr. Greig in
‘Field, no, 2801, p. 155, January 30, 1897).
+ Lepus europeus pyrenacus, Hilzheimer, Zool, Anzeiger, xxx. p- 612
(August 14, 1906). Type locality: “ Bagnéres.”
402 On new Characinid Fishes from South America.
LII.—Deseription of Two new Characinid Fishes from |
South America. By C. Tare Recan, M.A. i
MIMAGONIATES, gen. nov.
Body moderately elongate, compressed ; abdomen keeled,
but not strongly compressed toanedge. Mouth small; teeth
tricuspid, in a single series; no maxillary teeth; palate
toothless. Nostrils close together. Gill-membranes not
united, free from the isthmus. Scales cyeloid, of moderate
size; lateral line incomplete. Dorsal fin short, posterior in
position ; adipose fin present ; anal fin elongate.
Intermediate between Chirodon, Girard, and Leptagoniates,
Bler.
Mimagoniates Barberi, sp. n.
Depth of body 3 to 32 in the length, length of head 4 to
2, Snout much shorter than eye, the diameter of which is
24 to 23 in the length of head and a little less than the inter-
orbital width. Cleft of mouth nearly vertical; maxillary
not extending to below the eye. 42 to 45 scales in a longi-
tudinal series ; lateral line on 4 to 8 scales only. Dorsal 10;
origin equidistant from gill-opening and base of caudal, above
the anterior part of the anal. Anal 34-38; origin equidis-
tant from anterior part of eye and base of caudal; anterior
rays the longest, about 2 the length of head; free edge
straight or slightly concave. Pectoral extending to or a
little beyond the base of ventral. Caudal forked. A lateral
band (blackish in preserved specimens) from the lower part
of eye to the lower lobe of caudal. An oblique dark stripe
on the dorsal; anal with a dark margin.
Hab. Arroyo Yacé, Estacién Caballero, Paraguay.
Several specimens, the largest 40 mm. in total length,
collected by Dr. A. Barbero.
CTENOCHARAX *, gen. nov.
Body oblong, compressed; abdomen rounded. Mouth
small ; teeth slender, subconical, in a single series ; maxillary
toothed ; palate toothless. Nostrils close together ; gill-
membranes not united, free from the isthmus. Scales cycloid,
rather large ; lateral line incomplete. Dorsal fin short, nearly
entirely in advance of the rather short anal; no adipose fin.
Related to Aphyocharax, Gthr.
* The name Ctenocharax is given on account of the comb-like appear-
ance of the single series of teeth in the jaws.
On Raia undulata, Lacep. 403
Ctenocharax bogotensis, sp. n.
Depth of body 22 in the length, length of head 33. Snout
as long as eye, the diameter of which is 44 in the length of
head and 13 in the interorbital width. Cleft of mouth
oblique, the maxillary extending to below the anterior } of
eye. 33 scales in a longitudinal series; lateral line on 5 or
6 scales only. Dorsal 11, origin equidistant from eye and
base of caudal. Anal 14, origin below the last ray of the
dorsal; longest rays 4 the length of head; free edge
straight. Pectoral 2 the length of head, not reaching the
ventrals. Caudal notched, with rounded lobes. Coloration
uniformly olivaceous (in spirit).
Hab. Bogota.
a specimen, 63 mm. in total length, purchased in
1868.
LITI.—WNote on Raia undulata, Lacep.
By C. Tate Reean, M.A.
In the British Museum is a specimen of Raia undulata,
Lacep., from the coast of Cornwall, presented by the late
Mr. Harcourt Powell in 1880. As this species is not usually
regarded as belonging to the British fauna, I have tried to
find further evidence of its occurrence on the Cornish coast,
which is furnished by Couch, who, as has been pointed out
by Moreau, certainly had an example of Rata undulata, which
he regarded as a variety of R. microcellata, Montagu.
Raia undulata is described in detail by Moreau (Poiss. de
France, i. p. 434, 1881), who states that it is found on all
the coasts of France (Mediterranean, Atlantic, and Channel).
It is not a little curious that the original describer, Lacepéde
(Hist. Nat. Poiss. iv. p. 675, 1802), wrote that it occurred
‘entre les rivages si fréquentés de la France et de
PAngleterre.”
Of other British species Raia clavata, L., appears to be
nearest to R. undulata, but the latter should generally be
easily recognized by its system of coloration, which has been
thus deseribed by Couch (Fish. Brit. Islands, 1. p. 108, 1862) :
“ Another example differed considerably from the former
fi. e. Rata microcellata] in the nature and distribution of its
colours, which were still more beautiful. The ground-colour
was a brilliant yellow, marked with numerous gyrations,
which were lyre-shaped, each gyration being formed of a
404 Mr. H. L. Jameson on a
dark line, margined on each side by a series of pale yellow
spots, like beads.”
According to Moreau, the pale spots bordering the dark
undulated stripes may frequently be absent, or in large speci-
mens they may be greyish white in colour and the stripes may
be less distinctly marked than in the young.
LIV.—On a new Hare from the Transvaal.
By H. Lyster JAMESON.
In June of this year I noticed a large grey Pronolagus on
the Observatory Kopje adjoining the town of Johannesburg.
After some difficulty I succeeded in trapping this specimen,
a full-grown and pregnant female, which proved to belong to
a hitherto undescribed form. Pending a fuller study of the
genus Pronolagus I shall describe this hare as a High-Veld
race of Pronolagus Ruddi (Thomas and Schwann, Abstract
Po ZS. no, 18) p. 23, April 25, 1905; and ‘PZ. 1905,
vol. i. p. 272).
The Witwatersrand Pronolagus is specially interesting
because it extends considerably the range of the P. Ruddi
group westwards, hitherto known only from Zululand and
the Kastern Transvaal.
This form differs trom the type species (so far as I can
gather by comparing it with Thomas’s description, for I have
not had an opportunity of comparing it with the type) in its
generally grey colour, in the dark tail, black for its distal
third, in the absence of slaty bases to the fur, and in having
black soles to its feet.
Pronolagus Ruddi randensis *, subsp. n.
Size as in P. Ruddi (much larger than in P. crassi-
caudatus). Coat harsh, as in P. Ruddi. Ground-colour
buff, heavily pencilled with black, giving the impression of
rather dark grey when seen at a distance. Colour a little
lighter on rump. The long stiff hairs are black, with a
subterminal buff zone, as in P. Ruddi, and are about 18 mm.
long. Among these, especially along the back, are a few
very long hairs (40 mm. or more), which are white at the
base and black distally, sometimes with a subterminal white
* Subspecific name from Rand, popular abbreviation for Witwaters-
rand, the name of the range of hills on which Johannesburg is situated.
new Hare from the Transvaal, 405
ring. These hairs give the middle part of the back a parti-
cularly dark grey appearance. Wool-hairs buff, without the
slaty-grey bases characteristic of P. Ruddi.
Under surface buff, passing into grey on the flanks, so that
there is no sharp line between back and belly. This grada-
tion is due to the amount of black in the long coarse hairs
becoming less towards the ventral surface, until, in the mid-
ventral region, these hairs are entirely buff-coloured.
Head bluish grey, with a slight rufous tinge on the fore-
head. Nape-patch small and inconspicuous, rather more
rufous than back. Throat and chest as back. Ears grey, as
in P, Ruddi, but without the black edge. Limbs buffy
rufous, not any lighter towards digits. Soles black.
Tail very large and bushy, about 140 mm. long, including
the hairs ; dark reddish brown, pencilled with black, at base ;
distal third of tail black.
Skull as in P, Rudd?, but slightly larger and rather
narrower in the nasal region. The palatal foramina are not
quite so much narrowed posteriorly, their inwardly directed
edges, so conspicuous in P. Ruddi, being reduced to mere
narrowed ridges of bone, so that the walls of the nasal
chamber are exposed, as in P, crassicaudatus. The notch in
the incisors is more distinct than in Thomas’s figure of the
skull of P. Ruddi (P. Z. 8S. 1905, i. pl. xvi. fig. 4). The
upper molars have the uncrenulated anterior enamel-wall of
the posterior lamina extending considerably less than halfway
across the tooth, thus presenting conditions intermediate
between P. Ruddi and P. crassicaudatus. ‘The anterior lower
premolars also show an intermediate condition, their anterior
walls having a single notch on the left side and a double one
on the right.
Dimensions of the type (measured in the flesh) :—
Head and body 480 mm.; tail 110; hind foot 100;
ear 97; ear-opening 87.
Skull dimensions: greatest length 94; basilar length 74;
zygomatic breadth 41; nasals 47 x 20; interorbital breadth 18,
intertemporal breadth 15; palatal foramina 29 x 8°5; palatal
bridge 10; diastema 32°53 greatest breadth of nasal region,
at level of anterior upper premolar, 23. For measurements
of P. Ruddi see ‘Thomas and Schwann (loc. cit.) ; for dimen-
sions of two races of P. crassicaudatus see Thomas, Ann, &
Mag. Nat. Hist. (7) vol. x. pp. 244-246 (1902).
Hab. Witwatersrand Range, Transvaal.
Type. Female. No, 108 in my collection. Observatory,
Johannesburg, 5900 ft., 25th June, 1907.
In Aabits this hare resembles P. crassicaudatus and P, Ruddi,
406 On a new Dormouse from Asia Minor.
It frequents the stony kopjes of the Witwatersrand, spending
the day in holes under rocks and boulders, and coming out to
feed at dusk. I have only once found it lying out in the
daytime.
Like other Pronolag?, it frequently returns to the same
spot to defecate ; hence its presence in a locality can often
be detected by the piles of droppings on the hill-sides. Itisa
very retiring species, and seems little known among local
sportsmen, although it occurs right up to the outskirts of
Johannesburg.
Its short ears, heavy build, and general pose at once recall
the European rabbit. It is very fleet on foot; in fact, the
“red hare’’ (a collective name for the members of the genus
Pronolagus) is regarded by sportsmen as the fastest of the
South-African hares.
The uterus of the type specimen contained a single large
leveret.
University College, Johannesburg,
September 1907.
LV.—On a new Dormouse from Asia Minor, with Remarks
on the Subgenus “ Dryomys.” By OLDFIELD 'THOMAS.
In 1906* I formeda special subgenus to contain the dormouse
previously known as Hliomys nitedula, Pallas (syn. E. dryas,
Schr.), and gave it the name of Dryomys; but I now find
that this name is preoccupied {, and would propose to replace
it by Dyromys, an anagram of the same word.
Since I formed the subgenus there has been discovered the
large but nearly related Central-Asian species to which I
applied the specific name angelus, and Mr. Gerrit Miller has
drawn my attention to additional points of distinction between
* P. Z. 8. 1906, ii. p. 348.
+ “ Dryomys parvulus, Tschudi, Fauna Peruana, p. 179, lam. xiii.
fig. 1.” Philippi, An. Mus. Chile, Murideos de Chile, p. 20 (1900).
Although this is, no doubt, merely an erroneous rendering of Dry-
momys, yet, as it occurs with a specific name attached and a reference
both to a description and figure, it seems to be technically too valid as a
name to be used again for another animal.
Another of my generic names, Neotomys, was used as a misprint for
Jectomys by Wallace some years before I published it, but there the
misprinted term was without any mention of a species or reference to a
description, and consequently, viewed simply by itself, was a mere nomen
nudum, which was not the case with Philippi’s Dryomys.
On an “ Octopus ” with Branching Arms. 407
this group and Hliomys ; I am therefore, at his instance,
prepared to give it full generic rank.
The new form may be called
Dyromys nitedula phrygius, subsp. n.
Size about as in subsp. prctus, therefore rather larger than
in European nitedula. General colour above a light buffy
yellowish, markedly lighter and more yellowish than in either
typical netedula of Southern Russia, Winged of Greece, or the
Persian pictus. Tail lined grey, with a faint tinge of buffy.
Bulle larger than in the European forms.
Dimensions of the type (measured in flesh) :—
Head and body 100 mm.; tail 85; hind foot 21; ear 14.
Skull: greatest length 26°7; basilar length 20-4; greatest
breadth 15; nasals 8; length of upper molar series 3°6.
Hab. Murad Dagh, Ushak Province, Asia Minor (Mount
Dindymus, Phrygia, of classical maps). Altitude 7500’.
Type. Adult male. B.M. no. 5. 10. 6.1. Original
number 37. Collected and presented by W. Griffiths
Blackler, Esq. ‘Two specimens.
This pretty dormouse may be distinguished from the other
forms of D. nitedula by its brighter and yellower colour.
The Museum has recently received from Mr. Herbert Sykes a
specimen nearly topotypical of B'anford’s Myoxus pictus from
Central Persia, which might have been the same as the Asia
Minor animal, but proves to be a comparatively greyer and
less yellowish form. Specimens quite similar to the latter
have also been obtained in the Kurrum Valley, N.W. Frontier
of India, by Mr. C. H. T. Whitehead, thus forming a
considerable extension of the eastward range of this type of
dormouse.
LVI.— Notes on an “ Octopus” with Branching Arms.
By Epear A. Smita, I.8.0., F.Z.8.
[Plate XVI]
A COLLECTION sent home this year by Mr. R. Gordon Smith
from Japan contained a very remarkable specimen of an
Octopus with branching arms. It has been presented by him
to the British Museum.
Furcation in the arms of Cephalopods appears to be of
408 Mr. E. A. Smith on an
rare occurrence, judging from the few records of such abnor-
malities. C. Parona* has described and figured bifurcation
in an arm of ‘ Eledone moschata,” an additional arm in
E. Aldrovandi, and a bifureate arm in “ Octopus vulgaris.”
These are the only records I have been able to discover of
such irregularities of growth.
The present specimen presents a most remarkable instance
of arm-branching, the fureation not being confined to one
arm, but five out of the eight arms branch more than once.
Unfortunately the specimen, although sent home in formalin,
had evidently been dried previously, and all the viscera and
part of the head had been cut away, leaving only the body-sac,
the dorsal skin of the head, and the arms.
The latter are connected at the base by a rather broad
membrane, broadest between the ventral pair and the lower
laterals, rather narrower between the two laterals, and quite
narrow between the upper laterals and the dorsal arms and
between the latter also. The membrane between the two
ventrals has been removed, so it is impossible to say anything
definite regarding its extent. I think, however, we may
regard it as fairly certain that it was as wide at this part as
any where.
Arms.—The proportional length of the arms in such an
abnormal specimen is not of much importance. ‘They are
rather equal in thickness, excepting the dorsal pair, which
are a trifle more slender than the rest.
The right dorsal is the only arm that does not exhibit
furcation.
The left dorsal has two branchings, the first at about
6 inches from the mouth and the second an inch higher up on
the other side of the arm. ‘This, again, trifurcates at the
end.
The left upper lateral is normal for about 18 inches from
the oral centre, then bifurcates, the right branch again
bifurcating at a distance of 2 inches further up.
The right upper lateral is regular for 13 inches and then
bifurcates, the branch being a mere stump half an inch in
length and looking as if the rest of it bad been bitten off.
An inch and a half further up the arm a second fureation
occurs, and 24 inches further the arm trifurcates, one of the
branches, like that lower down, being only a stump an inch
long and without any suckers. One of the longer branches
trifurcates at the tip.
* Boll. Mus. Zool. Anat. Comp. Genova, 1900, no, 96,
?
“Octopus” with Branching Arms. 409
The left lower lateral arm is normal for 20 inches from the
mouth, then bifurcates.
The right lower lateral is regular for about 11 inches, then
bifurcates, the smaller branch again forking at a distance of
about 3 inches.
The left ventral is normal for 12 inches, then trifurcates,
one of the three branches being single to the end, the other
two, about 3 inches further up, again bifureating, with two of
the small branches again bifurcating ; so that this arm has,
in fact, nine branches altogether.
The right ventral is simple for about 15 inches, then
divides, one branch being single to the end, the other bifur-
cating 2 inches further up.
Altogether, then, it is seen that this remarkable animal
exhibits thirty-three branches to the eight normal arms,
mostly towards their ends. They are, with the exception of
the two stumps on the right upper lateral arm, provided with
suckers alternating in two rows, those on some of the most
slender branches being, in their dried and shrivelled con-
dition, rather obscure. The largest suckers are on the lateral
arms and are about 13 mm. in diameter.
The skin of the back, head, and the back of the arms,
excepting the ventral pair, are of a dirty purplish colour,
closely spotted with a darker tint and quite smooth. The
web between the arms is also spotted, excepting the portion
between the ventral pair and between the ventrals and lower
laterals. The spotting does not extend far up the arms,
which exteriorly are of a dark dirty purplish colour. The
eyes are about 1? inches apart, with a small cirrus behind each
aud a still smaller one in front of them. Behind each eye,
at a distance of about 15 mm., is a round pore-like circle
14 mm. in diameter. The ventral side of the body is pale,
excepting towards the sides, where the darker colour of the
back conimences.
Dimensions.
in.
Leneth of body from posterior end to between the eyes... 7}
Length of arms, from the mouth to the end of the longest
branch of each :—
Dorsalipatn sss: aso sc4 as) ea sveig acieeeiel Patetaics) shal oat 223
ight upper laterally... <icscractee cate acl teeta nee 25
Left o sine Gk Hake etna ed tae el 26
VEO WE Ge cioborbondoodceuCbocoDoue 20
Lett ” fh Doooonconoon oo so nono 0 oo 25
5 VONULall secs cites Rear SenVarohRedeatnerd as) star) aie) « 18
ii ee sone nie re hice bee c or cy tec Coron 19
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 27
410 On an “ Octopus” with Branching Arms.
The species appears to be the Polypus cephea (Gray) *
described from a single animal in the British Museum.
Although no special mention is made by Gray of the small
cirri in front of the eyes, they really exist in his specimen,
and the round pore-like circle behind each eye is also trace-
able. Gray did not give the colour of his species, but it is
spotted just in the same way as the specimen now described.
Upon the cause of the peculiar abnormality of this animal
I cannot venture an opinion. That it is an aberration from a
normal type is, I think, very evident, for were it not so we
should expect to find some regularity in the forking of the
arms. Such, however, is not the case, as neither the dorsal
pair, the laterals, or the ventrals exhibit any similarity.
With regard to the use of the generic name Polypus of
Schneider, Dr. Hoyle t+ has shown that it must take the
place of the commonly accepted Octopus, but his supposition
that it had been overlooked by modern writers is not quite
the case, for Jeffreys {, in describing Octopus, observes that
it is “the genus Polypus of Schneider, but Leach seems to
be the only modern naturalist who has adopted that name.”
This paragraph seems to have escaped Dr. Hoyle’s notice,
for he remarks :—“ Jeffreys’ attribution of the genus Loligo
to Schneider might seem to indicate that he was acquainted
with it [Schneider’s work], but on that hypothesis it is
difficult to explain why he quotes Sepiola as of Leach and is
quite silent as to the prior names for Octopus and Eledone.”
In justice to Jeffreys I may mention that he does not quote
Leach as the author of Sepiola, but Rondelet. I may also
refer to the fact that Owen § was acquainted with Schneider’s
work, and gave an outline of his classification, and, had the
“law of priority ” been observed at that time, doubtless he
would have employed the name Polypus. Finally, I would
point out that Philippi, in his ‘ Handbuch der Conchyliologie,’
p- 93, published in 1853, has employed the genus Polypus,
Schneider, sinking Octopus as a synonym.
* Cat. Cephalopoda Antepedia Brit. Mus. p. 15 (1849).
+ Mem. Manchester Phil. Soc. vol. xlv. no. 9, pp. 1-7.
{ Brit. Conch. vol. v. p. 148.
§ Trans. Zool, Soc. 1838, vol. ii. p. 125.
Mr. W. L. Distant on Homoptera. 411
LVII.— Rhynchotal Notes—XLIII. By W. L. Distant.
WITH one exception all the Cicadide described in this paper
are from the continent of Australia. The materials from which
these species were described are (1) from the Northern
Territories of South Australia collected by Mr. W. Stalker
and sent to the British Museum by Sir William Ingram and
the Hon. John Forrest, and also from another locality in the
same territory collected and sent to the Museum by Mr. H.J.
Hillier; (2) a collection made in Queensland by Mr. F. P.
Dodd; and (3) a number of specimens sent to me by
Mr. W. W. Froggatt for identification, with the intimation
that they were not included in the Monograph of Australian
Cicadidz written by Messrs. Goding and Froggatt. For
some other specimens I have to thank Mr. H. Ashton, of
Sydney.
Subfam. Creaprv2z.
Division THOPHARIA.
Thopha colorata, sp. n.
fg. Head and pronotum reddish ochraceous; apical and
basal margins of front and a broad transverse fascia on vertex
between the eyes black; pronotum with the margins a little
paler than disk, small obscure black spots at the anterior and
posterior margins of the central longitudinal impression;
mesonotum black, two obconical spots at anterior margin
only indicated by their pale margins, on each side of which are
two large discal, angulated, reddish-ochraceous spots, the
interior angles of which form the inner margins of the
obconical spots ; lateral margins of pronotum and mesonotum
and basal cruciform elevation reddish ochraceous, the latter
with a central longitudinal black spot ; abdomen above
black, the tympana reddish ochraceous, the apical segment
(excluding centre) cretaceous white; head beneath black,
anterior margins between face and eyes ochraceous, lateral
margins of face narrowly and obscurely ochraceous ; sternum
greyishly pilose; legs reddish ochraceous, anterior cox
streaked with black, intermediate and posterior coxz more or
less cretaceously pilose; opercula pale ochraceous ; abdomen
beneath piceous, with the posterior segmental margins
testaceous, the anal plate and segment reddish ochraceous,
basal and lateral margins of abdomen, and a large spot at
base of tympana, cretaceously white; eyes sessile, the breadth
27%
412 Mr. W. L. Distant on Homoptera.
between their outer margins equalling the length between the
anterior margin of the pronotum and the base of the cruci-
form elevation; tegmina and wings subhyaline, talc-like ;
tegmina with the base and about basal two thirds of venation
ochraceous or reddish ochraceous, apical venation brownish
ochraceous, costal and postcostal areas and base of posterior
claval margin black ; wings with the base, venation, and
basal halves of margins to anal area ochraceous or reddish
ochraceous, at apical : margins the veins are darker in hue;
face very elobose, with a central impressed longitudinal line,
on each side of which is a longitudinal series of obscure
nodules, lateral areas transversely striate ; rostrum reaching
the posterior cox.
Long., excl. tegm., ¢ 35 mm.; exp. tegm. 98 mm.
Hab. N. T., 8. Australia (H. J. Hillier, Brit. Mus.).
Division CICADARTIA.
Macrotristria Goding?, sp. n.
?. Vertex ochraceous, anterior area of front and a fasciate
line connecting it with base castaneous, irregular markings
surrounding the ocelli, a curved fasciate line at inner margins
of eyes, and the central posterior margin, black ; pronotum
ochraceous, the posterior and lateral margins a little paler in
hue, two central, discal, contiguous lines (united posteriorly
and ampliated at anterior mar gin), the furrows, and an oblique
and a curved spot on each side of the central lines, black
mesonotum ochraceous, with four obconical spots, the two
central ones darker and smaller and meeting two curved
angulated spots, which emerge from the ochraceous cruciform
elevation, black, extreme lateral margins also black; abdomen
above black, the margins of the abdominal segments on apical
half more or less ochraceous; body beneath black; face
castaneous, space between face and eyes ochraceous, inner
margins of eyes, posteriorly connected with a short oblique
fascia, black; coxze and legs ochraceous, longitudinal streaks
to coxee and femora, and the tibiz more or less, castaneous or
piceous, anterior and intermediate tarsi black; tegmina
hyaline, tale-like, the venation piceous, extreme base, costal
area, upper part of costal cell, and the claval veins ochraceous,
posteostal area, and basal half of posterior claval margin,
black; wings hyaline, tale-like, extreme base and most of
veins on basal half ochraceous. or stramineous, venation on
apical half and the veins to anal area piceous or black ; eyes
= . 3
robustly sessile; face broadly globose, finely transversely
Mr. W. L. Distant on Homoptera. 413
striate, medially longitudinally interrupted ; rostrum reaching
the intermediate coxze ; body beneath ereyishly pilose and
the abdominal segments narrowly margined with ochraceous.
Long., excl. tegm., 2, 36 mm.; exp. tegm. 118 mm.
Hab. Queensland (fF. P. Dodd, Brit. Mus.).
Macrotristria Hilliert, sp. n.
Vertex black, the lateral margins, two transverse basal
lines near inner margins of eyes, and a central basal point
behind ocelli pale ochraceous ; pronotum black, the lateral
areas gradually broadening from disk piceous brown, a central
longitudinal line, the posterior margin (narrowly) and which
is extended upwardly near each ‘posterior angle, and the
anterior angle of lateral margins pale ochraceous ; mesonotum
black, two central, obconical, dark castaneous spots, which
are narrowly margined with pale ochraceous, the lateral
margins greyishly pilose, the cruciform elevation ochraceous ;
lateral margins of the metasternum ochraceous; abdomen
above black, the abdominal margins (narrowly transversely
and broadly. laterally) and the apical segment caleareously
tomentose, the white lateral margins spotted with black ;
body beneath more or less ereyishly tomentose; a small
central spot on basal margin of face, a large transverse spot
between face and eyes, posterior margins to the sternal and
abdominal segments, opercula, coxe, and apices of the femora,
ochraceous ; rostrum black, ochraceous at base; tegmina and
wings pale hyaline, tale-like, tegmina with the venation
ochraceous at base, piceous on apical two thirds, costal and
postcostal areas black margined with ochraceous ; wings with
the veins mostly ochraceous, a few black on basal area; head
above subangularly produced, the front prominent; face
globosely compressed, strongly transversely ridged, with a
central longitudinal carinate line; rostrum reaching the
posterior cox; opercula in the male transverse, well sepa-
rated internally, moderately broadly convex posteriorly, not
extending beyond the basal segment.
Long., excl. tegm., 6 9, 27-32 mm.; exp. tegm. 91-
102 mm.
Hab. N.'T., 8. Australia; Hermansburg (H. J. Hillier,
Brit. Mus.).
Cicada Knowlest, sp. n.
Vertex from base to the anterior margins of the eyes black,
its lateral margins, ocelli, and front ochraceous, base of front
and a transverse spot at the apex of each lateral margin
414 Mr. W. L. Distant on Homoptera.
black ; face ochraceous, with a central discal spot, the cheeks,
and clypeus black, a large irregular spot at inner margin of
each eye and a longitudinal line to clypeus ochraceous ; pro-
notum ochraceous, the lateral and posterior margins, anterior
margin (centrally interrupted), and the furrows black ; meso-
notum ochraceous, with two large anterior, central, obconical
spots, on each side of which is a smaller obconical spot, two
spots in front of the basal cruciform elevation, and the poste-
rior lateral margins black ; abdomen above ochraceous, with
a basal, central, greyish spot, the margins of the segment, two
oblique, longitudinal, central fasciz on the apical and anal
segments, and a sublateral series of spots, black; body
beneath black ; legs, transverse abdominal fascie, and lateral
marginal spots ochraceous; coxe, apical spots to anterior and
intermediate femora beneath, and tarsal claws, black ; tegmina
hyaline, talc-like, the venation ochraceous, much darker on
apical area, base of costal area and the postcostal area black,
basal area tinged with pale sanguineous; wings subhyaline,
the venation ochraceous, basal area tinged with pale san-
guineous; vertex with three central longitudinal furrows ;
face centrally smooth, laterally transversely striate, pronotum
with the lateral furrows behind the eyes broadly foveately
excavate ; opercula in male short, transverse, rounded poste-
riorly, scarcely extending over base of abdomen, not meeting
inwardly, black, with their posterior and lateral margins
narrowly and obscurely ochraceous; rostrum just passing the
posterior coxee.
Long., excl. tegm., 6, 33 mm.; exp. tegm. 101 mm.
Hab. Fiji Islands (C. Knowles, Brit. Mus.).
Subfam. Gamaninz.
Division CICADATRARIA.
Tamasa tristigma.
Cicada tristigma, Germ. in Silb. Rev. Ent. ii. p. 69 (1834).
Tettigia tristigma, Stal, Ann. Soc. Ent. Fr. 1861, p. 617.
Tibicen kurande, God. & Frogg. Proc. Linn. Soc. N. S. Wales, 1904,
p. 605.
Tibicen Doddi, God. & Frogg. loc. eit. p. 602.
A species very variable in size; the British Museum now
possesses a fair series of specimens collected by Mr. Dodd in
Queensland, and I am indebted to the kindness of Mr. Froggatt
for cotypes of his species.
Mr. W. L. Distant on Homoptera. 415
Subfam. Vrerervrwz.
Division TAPHURARIA.
Abricta Stalkert, sp. n.
3d. Vertex black; front testaceous ; ocelli testaceous, and
in some specimens a distinct small, central, ochraceous spot at
base of vertex ; pronotum ochraceous, the margins, a broad,
central, longitudinal fascia (which is dilated anteriorly and
posteriorly), and the furrows pale castaneous, edge of basal
margin black ; mesonotum pale castaneous, two obscure
obconical spots at anterior margin, which are only denoted by
their paler margins, lateral margins, and also the lateral
margins of the metanotum, greyishly pilose ; abdomen above
pale castaneous, the posterior segmental margins narrowly
obscurely virescent and the anterior margins broadly dark
castaneous ; body beneath and legs brownish ochraceous, the
area between face and eyes black; coxee, base of rostrum,
and opercula in male pale ochraceous ; tegmina hyaline, the
venation ochraceous, extreme base, costal and postcostal areas
ochraceous, the latter centrally streaked with greyish, upper
apical area, and broad margins to the transverse veins at
bases of second, third, and fourth apical areas, fuscous brown,
posterior claval margin from about one third from base
fuscous brown ; wings hyaline, the venation, extreme base,
and anal area pale ochraceous, at outer posterior angle of
anal area a somewhat large fuscous-brown spot ; face reddish
castaneous, with a central, longitudinal, linear incision, the
lateral areas broadly transversely striate; clypeus with a
piceous spot on each side; rostrum reaching the posterior
cox, its apical area black; opercula short, somewhat
obliquely directed inwardly, widely separated internally, their
posterior margins subtruncate and not passing base of
abdomen.
Long., excl. tegm., ¢ 9, 18-18} mm.; exp. tegm. 50-
55 mm.
Hab. N. T., S. Australia ; Alexandria (W. Stalker, Brit.
Mus.) ; W. Australia; Nicol Bay (Coll. Dist.).
Abricta Frencht, sp. n.
3. Body above black; front, vertex, and pronotum with
a broken, central, longitudinal, testaceous fascia, in some
specimens scarcely visible; lateral margins of vertex, poste-
rior and lateral margins of pronotum, two discal inwardly
angulated longitudinal fasciz and lateral margins to meso-
416 Mr. W. L. Distant on Homoptera.
notum, margins of metanotum, and abdominal segmental
margins, testaceous ; body beneath black ; face, lateral areas
.
of sternum, opercula, and abdominal margins testaceous; legs
ochraceous, spots to coxe and trochanters, longitudinal streaks
or spots to femora, and apices of tibiz and tarsi black ;
tegmina bronzy-brown subhyaline, veins mostly ochraceous
before and piceous beyond middle, transverse veins at bases
of first, second, and third apical areas broadly margined with
black, costal margin black, costal area sanguineous ; wings
subhyaline, with the venation ochraceons; body robust ;
pronotum strongly furrowed; opercula short, transverse,
well separated internally, posterior margins moderately con-
vex, not passing the basal abdominal segment; rostrum
reaching the intermediate cox ; tegmina short and broad.
Long., excl. tegm., g ?, 16-18 mm.; exp. tegm. 34-
38 mm.
Hab. Victoria; Woori Yallock (C. French, Jun.).
Allied to the Tasmanian A. aurata, Walk., from which it
differs by the more outwardly rounded opercula, infuscate
tegmina, and globose abdomen, not depressed on lateral areas
as in Walker’s species.
Burbunga venosa, sp. n.
Vertex and front castaneous, lateral margins of vertex
ochraceous, ocelli red; pronotum ochraceous, the furrows
and sometimes a central longitudinal fascia brownish; meso-
notum greyish ochraceous, with four obconical castaneous
spots, the two central ones shorter ; abdomen above ochraceous,
the anterior margins of the last four segments castaneous, a
castaneous spot on each side of the anterior margin of the
apical segment; body beneath and legs ochraceous, face
testaceous, abdomen beneath shaded with brownish ; tegmina
and wings hyaline, tale-like ; tegmina with the venation as
far as the apical areas stramineous, veins and transverse veins
of the apical areas fuscous, costal margin fuscous, costal area
stramineous ; wings with the veins pale stramineous, the outer
marginal and tips of the apical veins palely fuscous; head
with the front prominent, in breadth considerably narrower
than lateral margins of vertex ; vertex with three longitudinal
impressions ; face prominent, globose, strongly transversely
striate, centrally longitudinally interrupted ; opercula in male
somewhat obliquely directed inwardly, their posterior margins
truncate, well separated internally ; rostrum only just passing
the intermediate coxe. 5
Long., excl. deems, ¢ 6.7015 jmm.; exp. tegm. 49-
50 mm.
Hab. N. T., 5. Australia (Hl. 4. Hillier, Brit. Mus.)
Mr. W. L. Distant on Homoptera. 417
Burbunga albofasciata, sp. n.
?. Body above pale ochraceous; pronotum with four
silvery-white, sericeous, longitudinal fasciz, the two central
shorter and not extending beyond the anterior margin of the
basal cruciform elevation, the two outermost longer and at
lateral margins; abdomen above with three similar discal
fascive ; body beneath and legs pale ochraceous, lateral
margins of the sternum and abdomen sericeously white ;
tegmina and wings hyaline, talc-like; tegmina with the costal
area and veins stramineous Sythe veins defining the posterior
margin of the radial area, the veins defining the second ulnar
area, and almost the whole venation defining the apical areas,
fuscous ; wings with the venation stramineous, the outer and
upper venation more or less fuscous; head with front promi-
nent, in breadth considerably narrower than lateral margins
of OIE vertex with three longitudinal incisions; pro-
oD
notum with two central discal ridges enclosing a flattened
5 5
somewhat darker fascia; face strongly produced, laterally
compressed, strongly transversely striate, narrowly longitu-
dinally interrupted at middle ; rostrum reaching the posterior
COX.
Long., excl. tegm,, 2, 15 mm.; exp. tegm. 46 mm.
Hab. N. T., 8. Australia; Alexandria (W. Stalker, Brit.
Mus.).
Division CHLOROCYSTARIA.
Beturia varicolor, sp. n.
Virescent ; posterior margins of the abdominal segments
above a little darker ; disk of abdomen beneath, tibia (ex-
cluding bases), and the tarsi sanguineous ; tegmina and
wings ‘hy aline, the venation greenish or ochraceous, tegmina
with the costal area more or less ochraceous and with eight
apical areas ; wings with six apical areas.
Var. a.—Head, pro- and mesonota pale testaceous; abdo-
men above testaceous, abdomen beneath and costal area of
tegmina sanguineous.
“Var. b.—Body above and costal area of tegmina san-
guineous ; body beneath and legs ochraceous ; tibiz, tarsi,
and diseal posterior marginal abdominal segments san-
euineous.
¢. ‘Tympana entirely exposed ; opercula elongate, oblique,
not reaching base of abdomen, and only inwardly covering
the cavities ; rostrum reaching the posterior cox, its apex
piceous; face somewhat elongate, lateral areas oblique,
418 Mr. W. L. Distant on Homoptera.
strongly transversely striate, centrally finely longitudinally
sulcate ; abdomen moderately inflated.
Long., excl. tegm., ¢ 19-20, ? 18 mm. ; exp. tegm., ¢
47-49, ¢ 54 mm.
Hab. Queensland (F. P. Dodd, Brit. Mus.).
Beturia modesta, sp. n.
3S ?. Virescent or ochraceous, probably virescent in fresh
or living specimens ; ocelli red ; tibize and tarsi more or less
testaceous ; tegmina and wings hyaline, the venation virescent
or ochraceous.
3. Uniformly ochraceous in typical specimen; front
anteriorly transversely subangulate, and centrally sulcate
between the ocelli; pronotum with a central, flattened,
longitudinal, concolorous, fasciate impression, which neither
reaches the anterior nor posterior margin, the lateral posterior
angles somewhat broadly posteriorly produced ; face with the
lateral areas oblique, strongly transversely striate, narrowly
longitudinally suleate; rostrum reaching the intermediate
cox; opercula very small, oblique, apically subangulate,
scarcely extending halfway across the cavities, and leaving
the posterior and interior areas widely exposed.
?. Uniformly virescent in typical specimen.
Long., excl. tegm., g 15, 2 12 mm.; exp. tegm., ¢ 40,
2 38 mm.
Hab. Queensland (F. P. Dodd, Brit. Mus.).
Allied to B. varicolor, Dist., but a smaller species ; the
opercula in the male much shorter and smaller and the cavities
much more exposed.
Mardalana suffusa, sp. n.
Vertex, pro- and mesonota virescent ; anterior margin of
front, a transverse spot to vertex near anterior margins of
eyes, surrounding area to each ocellus, two central longitu-
dinal fascize to pronotum and the furrows and lateral margins,
four obconical spots to mesonotum, the lateral longest, ex-
tending whole length of mesonotum, the central short and
meeting two slightly curved fasciz which start from two spots
in front of cruciform elevation, purplish brown ; head beneath,
sternum, and legs virescent, posterior area of the face, coxa,
and femora more or less spotted with purplish brown; teg-
mina and wings hyaline, the venation dull virescent or
brownish ochraceous, tegmina with the costal area virescent
streaked with testaceous, extreme bases of both tegmina and
wings ochraceous.
Mr. W. L. Distant on Homoptera. 419
3. Abdomen somewhat strongly inflated, pale greenish
ochraceous, the margins of the abdominal segments testaceous
brown.
?. Abdomen normal, not inflated, dull virescent, a broad
obscure, dark, central, longitudinal fascia above, and a
narrower and more distinct, longitudinal, central, purplish-
brown fascia beneath.
3 2. Rostrum reaching the posterior coxe, its apex
purplish brown; face strongly transversely striate, centrally,
narrowly, longitudinally suleate ; tegmina with eleven, wings
with five apical areas.
Long., excl. tegm., ¢ 25, ? 20 mm.; exp. tegm., 3 60,
2? 62 mm.
Hab. Queensland (fF, P. Dodd, Brit. Mus.).
Division MELAMPSALTARIA.
Melampsalta Froggatti, sp. n.
? . Head, pronotum, and mesonotum black, shortly greyishly
pilose; vertex with the ocelli, a short basal longitudinal sulca-
tion between them, and a spot near middle of lateral margins
purplish red, anterior margin of front more or less reddish ;
pronotum with a central longitudinal fascia not reaching
base, the furrows and extreme posterior margin, testaceous ;
mesonotum with two faint obconical spots margined with
purplish at middle of anterior margin, the lateral margins and
cruciform elevation also purplish red ; abdomen above some-
what purplish red; head beneath black, face and clypeus
purplish red, face with the basal margin and two central
longitudinal fasciz united posteriorly black; body beneath
and legs purplish red; apex of rostrum black; disk of
abdomen beneath piceous ; tegmina and wings hyaline, talc-
like, with their bases purplish red, the tegmina with a
slightly pale ochraceous tint, costal area and the venation
ochraceous, base of costal area purplish red; face globose,
centrally suleate for half its length, its lateral areas strongly
transversely striate, rostrum reaching the intermediate coxe.
Long., excl. tegm., 9, 13 mm.; exp. tegm. 40 mm.
Hab. Queensland (#. P. Dodd, Brit. Mus.); Cairns
(Colls. Froggatt and Dist.).
Melampsalta hermansburgensis, sp. n.
Body virescent, inclining here and there to ochraceous ;
legs ochraceous (in fresh and living specimens probably
uniformly virescent) ; tegmina and wings hyaline, talc-like,
420 Mr. W. L. Distant on Homoptera.
bases of both narrowly ochraceous, venation and the costal
area of tegmina ochraceous.
9. Body very robust; front somewhat subangularly
prominent ; ocelli purplish red; pronotum with two central
longitudinal lines, which broaden outwardly a little ante-
riorly and very widely so posteriorly ; between these lines on
anterior disk the surface is deepened or sulcate ; face globose,
strongly, centrally, longitudinally sulcate, lateral areas
robustly transversely striate ; rostrum slightly passing the
intermediate coxe ; tegmina with the bases of the upper vein
to lower ulnar area and the lower vein to radial area fused
for less than the length of basal cell.
Long., excl. tegm., 9, 17-18 mm.; exp. tegm. 50 mm.
Hab. N.'l., 8S. Australia; Hermansburg (H. J. Hiller,
Brit. Mus.).
Melampsalta kewelensis, sp. n.
@. Vertex and front black; a central longitudinal fascia
to front, lateral margins of vertex, and a basal spot between
the ocelli pale ochraceous; pronotum brownish ochraceous,
mottled with piceous, a broad, central, hourglass-shaped,
black fascia, containing a central, elongate, pale ochraceous
spot, anterior, posterior, and lateral margins pale ochra-
ceous, sublateral and subbasal margins black; mesonotum
black, with two central longitudinal fascize, the anterior
halves of which are narrow, the posterior halves widened,
notched, and continued to basal cruciform elevation, and a
large oblong spot on each lateral margin, pale ochraceous ;
cruciform elevation pale ochraceous ; abdomen above black,
the segmental margins and the apical segment pale ochra-
ceous, the latter with three longitudinal black fasciz ; head
beneath black ; a basal spot and margins to face, a trans-
verse basal line between face and eyes, and apex of clypeus,
pale ochraceous ; body beneath and legs pale ochraceous ;
cox and legs spotted or streaked with black; tegmina
hyaline, talc-like, the venation dull ochraceous, postcostal
margin, upper margin of basal cell, and base of posterior
claval margin black ; wings hyaline, the venation ochraceous ;
rostrum reaching the posterior cox ; tegmina with the upper
vein to lower ulnar area and the lower vein of radial area
fused for about the length of basal cell.
Long., excl. tegm., 2, 17 mm.; exp, tegm. 40 mm.
Hab. Victoria; Kewel (fill).
Allied to JZ. Waterhousei, Dist.
‘Mr. W. L. Distant on Homoptera. 421
Melampsalia murrayensis, sp. n.
&. Vertex, front, pronotum, and mesonotum black; a
small central elongate spot between and a similar one in front
of ocelli, a central longitudinal line on anterior half of pro-
notum, and two linear discal spots to mesonotum, dull ochra-
ceous; lateral posterior margins of pro- and mesonota, and
base of cruciform elevation, pale ochraceous ; abdomen orange-
yellow, above with two transverse basal fascize and three
longitudinal segmental series of four spots each, one series
central, the other two lateral; head beneath and sternum
black ; legs ochraceous, femora and coxe strongly streaked
with black ; rostrum reaching the posterior coxe; opercula
small, rounded, not extending inwardly beyond the posterior
coxee, not backwardly beyond the base of abdomen ; tegmina
with the bases of lower vein to radial area and upper vein to
lower ulnar area fused for less than the length of basal cell.
Long., excl. tegm., ¢, 11 mm.; exp. tegm. 30 mm.
Hab. Victoria; N.W. Murray River (C. French, Jun.).
Melampsalta sulcata, sp. n.
g. Vertex and front black, greyishly pilose; basal lateral
margins of vertex and a spot on lateral margins of vertex
sanguineous ; a longitudinal sulcation pouveen the ocelli
testaceous ; pronotum testaceous, centrally longitudinally
sulcate, submarginally narrowly piceous; mesonotum testa-
ceous, with four obconical black spots—two small, central,
and not extending far from anterior margin, and a larger one on
each lateral area reaching the anterior angles of the cruciform
elevation ; abdomen above testaceous, the segmental margins
sanguineous, a series of obscure fuscous segmental spots on
each lateral area; body beneath and legs sanguineous, face
with two central, broad, longitudinal, ‘black fascia, united
anteriorly ; the cheeks and area between face and eyes black,
thickly palely pilose; rostrum reaching the intermediate
coxe, face medially longitudinally suleate ; opercula narrow,
roundly obliquely directed inwardly, but widely separated ;
tegmina and wings hyaline, tale-like, their bases narrowly
sanguineous, the ‘venation dull greyish ; tegmina with the
bases of the lower vein to radial area and the upper vein to
lower ulnar area fused for a little less than the length of basal
cell.
Long., excl. tegm., ¢, 13 mm. ; exp. tegm. 37 mm.
Hab. Cape York.
422 Mr. W. L. Distant on Homoptera.
Melampsalia arenaria, sp. n.
3. Vertex, front, and pronotum black, thickly somewhat
longly pilose ; lateral margins of vertex, a central longitu-
dinal fascia to vertex and pronotum, on the latter widened
posteriorly, and the lateral and posterior margins of pronotum
pale ochraceous, on each side of the central fascia the whole
pronotal lateral areas are more or less testaceous, with the
furrows blackish ; mesonotum black, with two central, longi-
tudinal, angulated, pale ochraceous fascize, which are narrow
at anterior margin and angularly widened before the anterior
angles of the cruciform elevation, which is ochraceous, with
its anterior angles black ; abdomen above black, the seg-
mental margins obscure brownish ochraceous ; head beneath,
sternum, and legs ochraceous ; abdomen beneath dull obscure
ochraceous ; lateral margins and apex of face and apices of
tarsal claws black ; tegmina and wings hyaline, the venation
and costal area of tegmina ochraceous ; face with the lateral
areas oblique and strongly transversely striate ; rostrum
reaching the intermediate coxe, its apex black ; metasternum
black; opercula short, broadly transverse, posteriorly sub-
tr uncate, not meeting but not very widely separated internally ;
the bases of the lower vein to radial area and the upper vein
to lower ulnar area shortly fused and only for about one third
the length of basal cell.
Long., excl. tegm., ¢, 15-17 mm.; exp. tegm. 31-
37 mm.
Hab. Sydney (H. Ashton, on the sands).
Pauropsalta endeavourensis, sp. n.
g. Vertex and front black, a spot at apex of front and
the lateral margins of vertex ’ pale ochraceous ; ocelli red ;
pronotum reddish testaceous, with a central, longitudinal, pale
ochraceous fascia margined with black, margins pale ochra-
ceous, the furrows, a curved line on each lateral area, and a
spot on lateral margin black; mesonotum black, two discal
longitudinal fascize, which are inwardly angulated and dilated
and almost meet at middle, testaceous, ochraceous near ante-
rior margin ; cruciform elevation and margins of metanotum
pale ochraceous ; abdomen above ochraceous, the segmental
margins and a series of central and lateral spots more or less
black and greyishly pilose; head beneath black, margins of
face ochraceous ; sternum, legs, and abdomen beneath ochra-
ceous ; spots to coxe and sternum, femora (excluding apices),
margins of metasternum, and a spot on each side of apical
On a new Genus and Species of Cerambycidee. 423
segment piceous or black; tegmina and wings hyaline, the
venation mostly fuscous, costal area to tegmina and veins to
anal area of wings pale ochraceous ; opercula short and poste-
riorly rounded; rostrum reaching the intermediate coxe ;
bases of the lower vein to radial area and upper vein to lower
ulnar area fused for more than the length of basal cell.
Long., excl. tegm., ¢ 12, 9 14 mm.; exp. tegm., ¢ 32,
? 35 mm.
Hab. Queensland ; Endeavour River.
Mr. Froggatt kindly sent me a male and a female specimen
of this species.
Urabunana festiva, sp. n.
3. Vertex of head and front black ; a spot on each lateral
margin of front, lateral margins of vertex, and a spot before
each eye pale ochraceous; pronotum pale ochraceous, with a
broad, central, longitudinal, shining black fascia, which
inwardly contains a short pale discal streak, posterior margin
narrowly inwardly black; mesonotum paler or more stra-
mineous than the pronotum, with a broad, central, shining
black, longitudinal fascia extending through the cruciform
basal elevation, and a long obconical spot of the same colour
on each lateral area; abdomen above stramineous, first and
second segments (excluding lateral margins) and disks of the
other segments (gradually decreasing in width towards apex)
shining black; body beneath and legs very pale ochraceous ;
central area of face, anterior and posterior lateral margins of
face, anterior tibie and tarsi, and the rostrum black ; tegmina
and wings hyaline, tale-like, the venation stramineous or
pale virescent, postcostal margin and the outer margins to
the apical areas black ; opercula small and transverse; abdo-
men globose beneath ; wings with four apical areas (in a male
specimen now before me only three apical areas, which is
clearly a malformation).
Long., excl. tegm., ¢ 2, 13 mm.; exp. tegm. 32 mm.
Hab, Victoria; Kewel (Hill).
LVIII.—Description of a new Genus and Species of
Cerambycide from Natal. By W. L. Disranrt.
Subfam. Creraupycin 2.
GAHANIA, gen. nov.
g. Head with front short, slightly concave from side to
424 Ona new Genus and Species of Cerambycide.
side, marked off from clypeus by a distinct arcuate groove ;
eyes rather coarsely granulate, emarginate, the lower lobes
large ; antenne about three fourths the length of body, stout,
first joint slightly curved, gradually thickened to apex, third
shorter than fourth or following, these angulate in front of
apex; pronotum transverse, armed with a short conical
tubercle at middle of each side, and three obtuse tubercles on
disk ; mesonotum without stridulatory area; elytra feebly
convex, semitransparent, polished, obtusely rounded at apex,
with a short spine on each side at suture; anterior coxe
strongly transverse, but having their acetabula closed behind
by extension inwards of epimera to meet the prosternum,
acetabula of middie coxe extending to epimera ; posterior
legs much longer than anterior or intermediate legs, their
femora reaching to apex of elytra, first joint of posterior tarsi
almost as long as the following two united ; first abdominal
segment almost as long as the following three together.
Allied to Megacelus, Lac., but with larger and more
coarsely granulated eyes; elytra longer and not dehiscent at
the apex ; femora less thickened and the anterior coxal cavities
closed behind.
I have dedicated this genus to my friend C. J. Gahan,
the well-known authority on the Longicornia and an ever
helpful adviser to other workers.
Gahania Simmonds?, sp. n.
&. Head, antenne, thorax, body beneath, and legs pale
brownish ochraceous; eyes black; elytra pale shining
ochraceous, the basal area and sutural margins pale brownish
ochraceous, the subcostal areas behind middle distinctly paler ;
head finely somewhat sparingly punctate, coarsely rugulose
on disk between eyes, centrally, finely, longitudinally suicate
on posterior disk and between the antennal bases, front longly
pilose ; pronotum coarsely punctate and granulose, strongly
transversely depressed behind anterior margin, a much
narrower and finer transverse impression before basal margin,
tubercles as in generic description, of the three discal the
central is smaller and situate a little behind the others, the
conical lateral tubercles have their apices black; elytra
polished, semitransparent, sparingly finely punctate, the
punctures more distinct on the darker basal area ; the first
three joints of the antennee have outwardly a few long hairs,
and the lateral margins of the pronotum are longly sparingly
pilose; body beneath and legs somewhat thickly pilose.
Long., ¢, 28 mm.
On a new Species of Cicadide. 425
Hab. Natal; Durban (H. W. Simmonds, Coll. Dist.).
A male specimen of this species was taken by Mr. Simmonds,
of New Zealand, when on a recent visit to Natal. That
specimen is now in my collection, having been presented to
me by its discoverer. Mr. Bell Marley also tells me he took
another specimen at about the same time and locality. I
hope to eventually figure the species in my ‘Insecta
‘Transvaaliensia.’
LIX.—Description of a new Species of Cicadide from
Last Africa. By W. L. Distant.
Pycna Liliotti, sp. n.
¢. Head, pronotum, and mesonotum pale virescent,
thickly greyishly pilose; some spots to front, area of the
ocelli, and inner margins of the eyes piceous ; pronotum with
two central longitudinal fasciz, united into a broad fascia on
posterior half, and from the middle of which there proceeds on
each side an oblique line to eyes, the furrows, and a broad
oblique spot on each side of the posterior margin near the
outer angles, piceous ; mesonotum with two small obconical
spots on anterior margin, on each side an oblique line
reaching middle, where it is inwardly recurved, a transverse
linear spot near each anterior angle of the cruciform elevation
and the lateral areas broadly (enclosing a large pale spot at
base and apex) piceous ; abdomen above piceous, the seg-
mental margins broadly and a central longitudinal fascia
virescent ; body beneath and legs pale ochraceous; head
beneath (including face), large spots to femora, bases and
apices of tibiz and tarsi, apex of rostrum, basal abdominal
segment, and the segmental margins piceous or black; basal
margin of face, a basal transverse spot between face and eyes,
and the clypeus pale virescent ; tegmina with nearly basal
half opaque, greyish, with the veins virescent, two spots on
costal membrane, two in apical half of radial area, and a
transverse fascia at end of pale opaque area _piceous,
remaining area of tegmina hyaline, the veins testaceous, with
piceous spots, more particularly in transverse marginal and
submarginal apical series; wings bright pale ochraceous,
opaque, and outwardly margined with piceous for more
than half their length, the apical area hyaline; rostrum
reaching the second abdominal segment; opercula short,
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 28
426 Mr. G. S. Miller on
transverse, not meeting inwardly, pronotal lateral margins
only moderately angularly peceaes
Long., excl. tegm., ¢, 22 mm.; exp. tegm. 68 mm.
Hab. Brit. East Africa ; Nairobi (Cae: Elliott, Brit. Mus.).
Allied to P. hecuba, Dist., from which it differs by the
very much less produced pronotal margins, the opercula not
meeting internally (in P. hecuba they overlap), the opaque
colouring of the wings is more extensive and extends to the
anal area, tegmina paler, &e.
LX.—Vour new European Squirrels.
By Gerrit 8. MILier.
AMONG the 275 skins of European squirrels in the British
Museum are representatives of the following four hitherto
unnamed forms :—
Scturus vulgaris rutilans, subsp. n.
1899. Seiurus vulgaris rufus, Barrett-Hamilton, Proc. Zool, Soc. Lond.
p- 5 (part.). Not of Kerr, 1792.
© 1906. Seiurus vulgaris rufus, Trouessart, Bull. Mus. d’Hist. Nat. Paris,
xli. p. 360 (part.). Not of Kerr, 1792.
Type.— Adult male (skin and skull). B.M. no. 95. 4. 18. 7
Collected at Rudolstadt, Schwarzburg, Germany, December
24, 1894. (Lilford Collection.)
Diagnosis. —Colour much brighter than in Scvurus vulgaris
vulgaris, the body clear rutous * in summer, rufous tinged with
light smoke-grey along sides in winter ; tail at alll seasons clear
rufous, usually somewhat darker than body. Brown phase
(occasional but much less frequent than red phase) : body a
grizzled hair-brown, suffused with mummy-brown over back ;
tail slaty black.
Measurements—Type: head and body 223 mm.;_ tail-
vertebre 175; hind foot 62; ear from meatus 27. Skull:
condylo-basal length 46°6 ; mastoid breadth 25; postorbital
constriction 18; interorbital constriction 17; rostral breadth
at front of nasals 8°6; nasal 15°4; diastema 12:2; man-
dible 33; maxillary tooth-row (alveoli) 9°6; mandibular
tooth-row (alveoli) 9°6
Specimens examined. — Fifty-five from the following
* The exact shade in the type between the cinnamon-rufous and
orange-rufous of Ridgway, but somewhat lighter than either.
Four new European Squirrels. 427
localities :— Germany, Marxheim, Bavaria, 5; Blumenthal,
Hannover, 2; Marburg, Hessen-Nassau, 2; LIlsenbyrg,
Saxony, 7; Magdeburg, Saxony, 1; Wernegerode, Saxony, 1;
Strass, near Burgheim, Schwaben, 8; Niesky, Silesia, 1 ;
Rudolstadt, Thiiringen, 4; Ummerstadt, Thiiringen, 12 ;
Neustadt, Wied, 4: Austria-Hungary, Haida, Bohemia, 1 ;
Karlsbad, Bohemia, 1; Hatszeg, Hungary, 1: Roumania,
Bustenari, 5.
Remarks.—This is the bright-coloured squirrel of Central
Kurope, ranging trom Germany eastward through Austria-
Hungary to Roumania. In Switzerland it apparently inter-
grades with S. vulgaris ttalieus, in which the brown phase is
dominant, but the exact relationship between these two forms
is not yet clearly understood, and for this reason I have
omitted the Swiss specimens from the list of material examined.
To this animal and the next the name rufus has been applied
by Barrett-Hamilton and Trouessart, but Kerr’s Securus vul-
garts rufus* is strietly a synonym of S. vulgaris vulgaris,
and therefore cannot be used in this sense.
Sciurus vulgaris russus, subsp. n.
1899. Seiurus vulgaris rufus, Barrett-Hamilton, Proc. Zool. Soc. Lond.
p- 5 (part.). Not of Kerr, 1792.
1906. Securus vulgaris rufus, Trouessart, Bull. Mus. d’Hist. Nat. Paris,
xii. p. 360 (part.). Not of Kerr, 1792.
Type.—Adult male (skin and skull). B.M. no. 97. 11. 6. 2.
Dinan, Cétes-du-Nord, France, October 29, 1897. Collected
and presented by W. Jennings Bramley, Esq.
Diagnosis.—Similar to Sciurus vulgaris rutilans, but with
colour in red phase darker and less bright, the body cinnamon-~
rufous strongly tinged with chestnut in summer, hazel + much
suffused with dull light smoke-grey along sides in winter;
tail at all seasons a light bright chestnut (Ridgway). Brown
phase infrequent, similar to that of rwtzlans.
Measurements.—Vype: head and body 202 mm. ; tail-
vertebre 166; hind foot 54; ear from meatus 30. Skull:
condylo-basal length 46; zygomatic breadth 29; mastoid
breadth 24°6 ; postorbital constriction 16°8 ; interorbital con-
striction 15:4; rostral breadth at front of nasals 7°4; nasal
15:2; diastema 12:4; mandible 31°4; maxillary tooth-row
(alveoli) 9; mandibular tooth-row (alveoli) 8°6.
* © Animal Kingdom,’ p. 255 (1792).
+ In the type the exact shade of the red is slightly ae than hazel,
but the general effect produced by the light and dark annulations is
intermediate between hazel and chestnut.
28%
498 Mr. G. S. Miller on
Specimens examined.— Twenty-two, from the following
localities :—Holland, Oosterbeek, 2; Graveland, 4: France,
Dinan, Cédtes-du-Nord, 4; Duclair, Seine-Inférieure, 2;
Seine-Inférieure, no exact locality, 3; Manonville, Meurthe-
et-Moselle, 2; Meurthe-et-Moselle, no exact locality, 5.
Remarks.—While the exact limits of the range of this
squirrel are still unknown, it is probable that the animal
occupies the coast-region from Denmark southward at least
to Brittany. In the Landes of South-western France it is
apparently replaced by the North-Spanish squirrel or a closely
related form. Westward it extends across France to the
Valley of the Moselle.
Tn colour, as well as in range, Sciurus vulgaris russus and
S. v. rutilans correspond with the two races of red-backed
voles, Evotomys hercynicus rubidus and EH. h. hercynicus,
occurring in the same regions.
Sciurus vulgaris numantius, subsp. n.
1905. Sfetwrus] sp., Cabrera, Bol. Real Soc. Espan. Hist. Nat. iv.
pp. 224, 231 (April 1905).
1905. Sfetwrus | rufus, Cabrera, Bol. Real Soc. Espati. Hist. Nat. iv.
p. 225 (April 1905).
Type.—Adult female (skin and skull) collected by Gerrit
S. Miller, at Pinares de Quintanar de la Sierra, Province of
Burgos, Spain, October 28, 1906. Original number 7418.
Diagnosis.—Size greater than in the Central-Kuropean
forms, but less than that of Sccurus dnfuscatus, Cabrera.
Colour of upperparts in light phase not as dark asin S, infus-
catus, and tail never with white median area on lower
surface.
Colour—Type: head, back, sides, and outer surface of
legs a uniform indistinctly grizzled brown, intermediate be-
tween the broccoli-brown and wood-brown of Ridgway; a
faint russet tinge along middle of back. Muzzle and fore
part of face between ochraceous-buff and clay-colour. Cheeks
drab. Lar-tufts blackish brown. Sides of neck pale dull
wood-brown. Inner surface of legs and ill-defined stripe
along sides of belly dull light hazel. Feet like inner side of
legs, but paler. Tail a very dark rufous, approaching the
chestnut of Ridgway, especially near base, but rather more
red ; median portion of tail below lighter, the hairs buffy grey
through basal half, each with two drab annulations. Under-
parts buffy white, the chin and interramial region light ecru-
drab.
In the dark phase the tail is clear bluish black, very faintly
Four new European Squirrels. 429
grizzled along median region below, and the back is much
darkened by a blackish suffusion.
Colour variation shows itself chiefly in the greater or less
tendency to assume the dark phase. Occasionally the hazel
of sides brightens nearly to a dull rufous and spreads to
lateral portion of dorsal area, the portion bordering white of
ventral surface at the same time becoming nearly buff. Har-
tufts either blackish or reddish.
Skull and teeth —TVhe skull and teeth are intermediate in
size between those of the ordinary Central-Huropean forms
(russus and rutilans) and the large S. infuscatus. In form
they show no special peculiarities.
Measurements —Vype: head and body 237 mm.;_tail-
vertebrae 230; hind foot 66; ear from meatus 34. Skuil:
condylo-basal length 50; zygomatic breadth 333; mastoid
breadth 26; postorbital constriction 17-4; interorbital con-
striction 18°8; rostral breadth at front of nasals 8:2;
nasals 16; diastema 13; mandible 34; maxillary tooth-
row 9°8; mandibular tooth-row 10.
Specimens examined. — Nineteen, from the following
localities :—France, Solférino, Landes, 1; St. Jean de Luz,
Basses Pyrénées, 1: Spain, Sierra de Dubros, Asturias, 2 ;
Arrechavaleta, Vitoria, 2; Panticosa, Huesca, 6; Pinares de
Quintanar de Ja Sierra, Burgos, 7.
Remarks.—This squirrel is evidently the Sccuwrus sp. ? of
Mr. Cabrera’s review of the Spanish members of the genus,
as its characters agree with those assigned to this problematic
form. It is also without much doubt the rufus of the same
paper, though on this point the evidence is not so clear.
Whatever the squirrel of the Lower Ebro Valley may be, it
is very unlikely to prove identical with either russus or
rutiluns, the two forms confused under the name rufus at the
time when Mr. Cabrera wrote.
Sciurus vulgaris lileus, subsp. n.
1906. Sciwrus vulgaris italicus, Trouessart, Bull. Mus. d’Hist. Nat,
Paris, xil. p. 364 (part.).
Type-—Young adult female (skin and skull). B.M.
no. 7.9.8.1. Agoriani, north side of Lyakura (Parnassus) Mts.,
Greece. Received from Wilhelm Schliiter.
Diagnosis.—Colour in brown phase peculiar in the notice-
able contrast of the very dark almost blackish posterior half
of back with hair-brown shoulders and neck. Red phase not
known, and probably rare or absent.
Colour.—T ype: general colour above a grizzled hair-
460 Colonel C. Swinhoe oz
brown, paler and more grey on cheeks and across muzzle,
‘much darkened with blackish on posterior half of back and
on outer surface of hind legs. Inner surface of hind legs and
line along sides of body bordering white of underparts tawny-
ochraceous, becoming lighter and duller anteriorly and con-
tinued along sides of neck and spreading over outer surface
of fore legs. Feet dull tawny-ochraceous suffused with
blackish. ‘Tail blackish suffused with tawny-ochraceous
beneath surface. Underparts creamy white; chin and inter-
yamial region light drabby~grey. :
Skull and teeth.—The skull and teeth show no special
peculiarities.
Measurementsi—Type: hind foot 60 mm.; ear from
meatus 29. Skull: zygomatic breadth 31°4; postorbital
constriction 17°8 ; interorbital constriction 17; rostral breadth
at front of nasals 7-4; nasal 14; diastema 12°8; man-
dible 32°4; maxillary tooth-row 9°8; mandibular tooth-
row 9°8.
Specimens excamined.—Three, all from the type locality.
LXI.—New Species of African and Indo-Malayan
Hesperiide. By Colonel C. SwinHog, M.A., F.L.S., &e.
Family Hesperiidz.
Casyapa kallima, nov.
3 @. Dark olive-brown ; frons, palpi, body below, and
legs bright orange-ochreous ; antenne with ochreous and
brown bands: fore wings with a very broad and prominent
orange-ochreous band from the outer margin above the hinder
angle, which it does not reach, to the middle of the costa,
running inwards for some distance along the costal vein, just
below its middle the band is slightly contracted ; cilia of both
wings dark brown ; no other markings above or below.
Expanse of wings, g 23%, @ 25 inches.
Milne Bay, New Guinea (types in B. M.).
Allied to C. callixenus, Hew., from Dorey.
Tagiades louisa, nov.
¢. Blackish brown; palpi white beneath ; frons with a
white spot on each side: fore wings with two large hyaline
spots at the end of the cell, one outside its lower angle and
new African and Indo-Malayan Hesperiide. 431
another close beneath it, all more or less triangular ; a sub-
apical row of six small spots in the usual recurved line: hind
wings with about one half of the lower portion white, the
white running up the abdominal margin to the base ; two
very large black spots in the middle of the dise touching the
inner side of the outer curve of the brown portion of the
wing; no marginal marks or spots. Underside with two
additional hyaline spots on the fore wings near the hinder
angle; hind wings with a somewhat narrow black costal
border, the two discal spots much smaller, and one minute
black mark on the outer border below the middle ; legs and
body white. *
Expanse of wings 2 inches.
Rossel Island (type in B. M.).
Sarangesa haplopa, nov.
$. Dark blackish brown with a slight red tinge; palpi
whitish below : wings above with spots and bands much as
in subalbicans, Bethune-Baker, but the submarginal band of
the fore wings is curved evenly with the outer margin ; a
hyaline spot on the upper part of the cell at three fourths and
another immediately above it ; a small round spot below the
lower end of the cell and a larger triangular spot immediately
below it, also three small subapical spots close together, the
centre spot a little inwards. Underside very different to
subalbicans, being uniformly blackish brown, the spots on
the fore wings as above ; a blackish discal band, a pale band
just beyond it, even with the outer margin on both wings;
internal spots as above; a dark macular submarginal band.
Expanse of wings 1% inch.
E. Ruwenzori, 7000’ (G. Legge) (type in B. M.).
Celenorhinus chinensis, nov.
¢g. Antenne brown beneath, white above, with thin
brown bands ; club white, with brownish tips ; head, body,
and wings uniform blackish brown, with an olive tint ; palpi
white beneath ; abdomen with thin ochreous bands on the
last four segments and an ochreous tip: fore wings with two
large, nearly square, semihyaline white spots or patches, the
first nearly filling up the end of the cell and extending right
across it, the other immediately below it of the same size and
shape, the two forming a band which is very slightly out-
wardly oblique; a small round spot outside the centre of this
band, another below its outer lower edge, and a still smaller
spot below this and more inwards ; subapical dots and large
432 Colonel C. Swinhoe on
orange-ochreous spots on the hind wings as in C. pyrrha,
de Nicéville; cilia of fore wings white, of hind wings
ochreous.
Expanse of wings 2,/5 inches.
Omei-shan, W. China (Crowley Bequest) ; two examples
(type 1 in B. M. ie
The central white semihyaline band is more erect than in
any other species of the leucocera group.
Celenorhinus Plétzi, nov.
3. Dark blackish brown with an ochreous tinge; upper
half of the antennee and club ochreous on the underside: fore
wings with a very broad orange-ochreous discal band, not
quite touching either margin, slightly narrower towards costa,
its inner margin a little before the middle of the wing and
nearly straight, i its outer margin somewhat roundly curved
and uneven in its upper half; an orange-ochreous subapical
spot: hind wings with a large orange-ochreous apical patch.
Underside slightly paler ; fore wings as above; hind wings
without markings ; palpi and face orange- -ochreous ; ; abdomen
and legs below smeared with ochreous.
Expanse of wings 1,55 inch.
Bipindi, Cameroons ; 3 one example.
Allied to C. atratus, Mab., but quite distinct.
Abaratha siamtca, nov.
@. Pale pinkish brown, almost as pale as A. saraya,
Doherty ; palpi with some ‘white hairs beneath : fore wings
with a semihyaline white spot in the cell at one fourth, with
a blackish spot on its outer side and a mark below it, this
being the limit of the basal brown space ; following this is a
pale pinkish band of three large square spots joined together,
the centre one outermost ; a broken lunular spot near end of
cell, a round spot below the end, a minute white spot above
this, another below it, both beyond it; a little larger spot,
below again and inwards, not far from the centre of the
hinder margin, the usual three subapical spots: hind wings
with four brownish bands of conjoined spots, with pale
pinkish spaces between. Underside paler and more pinkish,
the hyaline spots as above, the bands more prominent and
more macular, many of the spots being disconnected ; legs
whitish.
Expanse of wings 1,85 inch.
Shan States, Siam (type in B. M.).
new African and Indo- Malayan Hesperiide. 433
Arnetta Binghami, nov.
&. Dark blackish brown ; antennee white at the tips and
with white crook ; palpi brown beneath, not white as is usual
in this genus: fore wings narrower than usual; two white
semihyaline spots at the end of the cell, a small lunular semi-
hyaline mark below the end and a little beyond it, a small
white spot a little beyond the upper end of the lunule, two
white subapical dots: hind wings without markings. Under-
side nearly as dark as above; markings of the fore wings
similar, except that the two subapical dots are absent.
Expanse of wings 1} inch.
Tavoy Valley, Burmah (Bingham) (type in B. M.).
Salween Valley (Bingham).
Superficially looks like a Parnara, but has the venation
and antenne of Arnetta.
Telicota ternatensis, nov.
g. Orange-ochreous, as in palmarum, Moore, but darker
and brighter coloured ; antennz ochreous brown, with black
bands and white apiculus; palpi pale ochreous, and a band of
that colour between the antenne; thorax and abdomen
brown, the former with long ochreous hairs on each side:
fore wings with a thin black streak on the subcostal vein and
all the veins black, another on the internal vein, a dull brown
band from the base to the middle under vein 2 ; a black oblique
band containing the sexual mark quite straight to the end of
the cell, then outwards below vein 6; the outer marginal
band much asin palmarum : hind wings blackish brown, with
a large ochreous spot above the middle of the centre, and a
broad ochreous patch in the disk, as in palmarum. Under-
side: fore wings with the black discal band extended hind-
wards, filling the basal space below the cell, and extended in
a paler form along the hinder margin to the angle; the outer
band is pale and has a black macular band limiting its inner
margin; the hind wings are ochreous irrorated with black
atoms except the spot and patch, the former has an outer
black line and the latter a black macular border.
Expanse of wings 1,%5 inch.
Ternate (Wallace) (type in B. M.).
Morotai (Bernstein).
This species is not unlike palmarum, Moore, but that insect
belongs to Mabille’s genus Corone, the species of which,
though resembling Yelicota, have no sex-mark,
434 Colonel C. Swinhoe on
Caltoris laraca, nov.
6. Of the usual olive-brown colour; palpi below and
pectus whitish: fore wings with a white diaphanous spot at
upper end of cell, another obliquely below it before the lower
end, a large square spot touching the lower end of the cell
and a smaller equally square spot obliquely above it, its inner
lower corner nearly touching the upper outer corner of the
lower spot; a dot in a line beyond these two spots, and above
this are the three usual subapical dots in the usual curve; a
small spot in the middle of the internal vein; hind wing
with two spots close together in the middle of the disk ; the
centre part of the fore wing is darker than the remainder of
the wing, and the hind wing is broadly margined with darker
brown along the costa. Underside much paler, with an
ochreous suffusion ; spots as above, the lower internal part of
the fore wing suffused with black.
Expanse of wings 1,8) inch.
Woodlark Island (eek) (type in B. M.).
Notocrypta aluensts, nov.
3 @. Uniform dark black; palpi white beneath: fore
wings with an outwardly curved, curiously shaped medial
band composed of four spots from the middle of the costa to
near the hinder margin before the angle; first spot quite
small and nearly touching the costa, the second large, nearly
round, the third cylindrical, its outer two thirds extending
beyond the round spot, all three touching each other ; the
front spot small, excavated on its inner side like the letter C
below the outer end of the third spot and slightly separated
from it: hind wings unmarked; cilia black. Underside
blackish brown, the band not macular; male with two white
subapical dots, first near the costa, the others well separated
from it and nearer the outer margin ; female with a series of
five subapicai dots, curving outwards and then downwards.
Eixpanse of wings, ¢ 1%, ¢ 1,8, inch.
Alu Islands.
Somewhat resembling wokana, Plots, from Aru and Keé
Islands.
Ismene lusca, nov.
?. Palpi, face, and pectus scarlet-orange, last joint of
palpi brown; antennee black; head, body, and wings olive-
brown ; abdomen with scarlet-orange bands on the last two
segments ; anal tuft similarly coloured: fore wings without
new African and Indo-Malayan Hesperide. 435
any markings whatever: hind wings with the abdominal
margin from vein 2 scarlet-orange. Underside: thorax and
legs with orange hairs ; abdomen scarlet-orange with black
bands on each side; wings paler than above, tinted with
scarlet-orange ; the lower part of fore wings from vein 2
whitish grey; the hind wings with the interspaces streaked
with scarlet-orange, becoming more intense towards the
abdominal margin, where the wing is for about one third
entirely of that colour.
EXxxpanse of wings 2-4, inches.
Maros, 8. Celebes (type in B. M.).
flasora minsona, nov.
dg. Palpi ochreous grey; eyes with a white ring round
them ; last joint of palpi, antenne, head, body, and wings of
a uniform olive-brown with a slight ochreous tint; frons and
head tinted with blue-green; abdomen with dark brown
segmental bands: wings without any markings ; cilia pale
greyish ochreous. Underside much paler: fore wings with
the cell-space and a short transverse band beyond the cell
pale and ochreous tinged; hinder margin also pale: hind
wings darker, no blue reflections ; anal lobe, which is re-
stricted and blunt, with a very large blackish patch; an
attenuated white band from the costa near apex to the abdo-
minal margin above, the blackish patch, which it touches,
the band broken above the patch; body greenish grey ;
legs ochreous grey.
kixpanse of wings 2,}, inches.
Borneo; two examples (Crowley Bequest) (typein B. M.).
Hasora wortha, nov.
3. Palpi ochreous grey, whitish at the sides ; eyes ringed
with white; antenne brown, tinged with red in parts and
whitish on the underside below the club; head, body, and
wings olive-brown, tinged with ochreous, especially towards
the base of fore wings and on the lower and abdominal
portions of the hind wings; no markings ; cilia ochreous
erey. Underside paler; a broad darker shade on the fore
wings below the cell and a discal shade: hind wings darker ;
a broad pale shade across the disk and on the abdominal
margin ; anal lobe as much restricted as in anura, de Nicé. ;
alarge blackish patch on and inside of it, on the upperside of
which is a small ochreous spot ; body brown; legs ochreous.
Expanse of wings 2y'p inches.
Java (Crowley Bequest) (type in B. M.).
436 Mr. R. I. Pocock on the Pattern of the
Hasora hobroa, nov.
6. Palpi, head, and thorax green ; abdomen pale brown,
its basal half covered with dull green hairs ; antenna black:
wings olive-brown, paling towards base with an ochreous
tinge; a green patch on the costa of fore wings one third
from the base; a slight greenish-grey suffusion below costa
at the base; the hind wings with the basal and abdominal
areas covered with greenish-grey hairs; cilia brown, with
pale tips ; wings without any other markings. Underside:
wings paler and suffused with ochreous; fore wings with the
apical and outer marginal spaces darkest ; hind wings with a
broad pale discal shade, the wing dark on both sides of it and
merging into blackish in and above the anal lobe, which is
much restricted ; cilia with a white short basal line above the
anal line; pectus and body grey tinged with green; legs
darker green.
Expanse of wings 2;% inches.
Celebes (type in B. M.).
The fore wings are shorter than usual and the hind wings
are very deep and round, and it has no stigma.
flasora meala, nov.
3. Palpi below and aring round the eyes ochreous grey ;
head, body, and wings olive-brown, as in chromus, Cram. ;
cilia brown; wings without markings: “wings below paler and
with a gloss on them; fore wings with the costa broadly
smeared with blue-green above the cell, a short dark trans-
verse shade beyond the end: hind wings with a thin, straight,
transverse white band or thick line from the costa a little before
the apex to the abdominal margin one fifth from the anal
angle, the whole space inside this line blue-green; anal lobe
damaged on both wings, but is evidently much restricted and
has a blackish patch ; face, pectus, and entire body brown;
legs ochreous grey.
Expanse of wings 275 inches.
Celebes (Wallace) (type in B. M.).
A good and distinct species.
LXII.— The Significance of the Pattern of the Cubs of
Lions (Felis leo) and of Pumas (Felis concolor). By
R. I. Pocock, F.L.S., F.Z.S., Superintendent of the
Zoological Society’s Gardens.
{Plates XIX. & XX.]
As a very general, perhaps invariable, rule, members of the
cat tribe (Felis) that are spotted or striped when adult are
Cubs of Lions and of Pumas. 437
similarly spotted or striped when young—that is to say, the
pattern undergoes no very marked change with growth, apart
from gaining or losing in distinctness. Even when it is
evanescent in the adult, such indications of it as are pre-
served coincide with the more clearly defined pattern of the
cub or kitten. It is therefore permissible to conclude that
those species in which the adult is self-coloured and the cub
variegated were marked as their cubs are marked and in no
other way. Hence the pattern of the cub must be treated
as a specific character, and may be regarded as affording a
most useful clue to affinity. Possibly, indeed, the true
relationships of some of the self-coloured species of Felis
will never be certainly ascertained until the pattern of the
foetal or newly born young is known.
It is in the truth of the above-stated propositions that lie
the chief terest and importance of the pattern exhibited by
the cubs of lions and pumas.
It is well known that the newly born cubs of those species
show a definite pattern of dark marks upon a tawny or pale
brown ground-colour; but although the presence of this
pattern has been cited repeatedly as evidence of the descent
of the species concerned from striped or spotted ancestors,
I am not aware that it has been used previously as a guide
in determining their affinities with other existing forms of
the genus Felis.
Examination of a series of skins of lion cubs shows that
the pattern, which has been described sometimes as “ spots,”
sometimes as “ stripes,” varies considerably in intensity with
individuals. The meaning of this variation is unknown to
me, since all the skins I have seen belong to specimens born
in menageries from parents of unknown geographical origin,
Sportsmen and collectors, unaware of the interest of the
question, have never apparently brought skins of wild-born
cubs from different localities. There are therefore no data
from which an opinion can be formed as to the local con-
stancy of the coloration of the cubs and of the value of the
variation, if any, in taxonomy.
One of the best-marked examples I have seen was born at
the Clifton Zoological Gardens in the spring of 1904 and
is preserved in the Bristol Museum. An account of it was
published * by Mr. Herbert Bolton, F.R.S.E., F.Z.S., the
curator; and I am indebted to him for kindly lending
me this and one other specimen for examination and
description +.
* Proc. Bristol Society, (2) x. pp. 248-249 (1904).
+ I have also seen other specimens in the Bristol Museum and in the
Museum of the Zoological Gardens at Clifton, where they were bred,
438 Mr. R. I. Pocock on the Pattern of the
The ground-colour is a sandy or in parts a golden yellow
fading to white on the lips, the chin, the mterramal area,
the chest, the posterior part of the belly, and the inner side of
the limbs. The underside of the tail is also whitish in the
middle line ; but the throat and the median part of the belly
are washed with yellow. ‘There is a conspicuous and rather
large whitish patch over the inner half of the eye. The
back of the ears is jet-black with a narrow edging of white.
The pattern, which consists for the most part of spots, is so
abundant and diffused that the interspaces look like pale
stripes on a dark ground. The spots are rosette-spots like
those of an ounce, an Indian leopard, or a jaguar—that is to
say, they consist of a black or dusky brown more or less
broken up rim surrounding an area which, though much
lighter than the rim, is decidedly darker than the intervening
spaces. Everywhere on the body and on the upper portion
of the limbs the spots are markedly wider than the inter-
spaces. On the upper surface of the head and along the
spine the spots are more heavily pigmented than elsewhere.
except on the hind leg between the knee and the hock and
on the distal end of the tail, where they are as black as on
the back. On the head the spots run into six rather con-
fused and broken up longitudinal stripes, two admedians
mostly blended together, which pass backwards from above the
eyes on to the nape of the neck, and two laterals on each side,
which converge inwards over the occiput and fuse with the
admedians on the fore part of the nape. The external of
these rises just above the ear on each side; the internal
rises much further forwards above the inner angle of the eye.
As in the chitah (Cynatlurus jubatus), there is a patch of
pigment extending downwards from the inner angle of the
eye to the white of the upper lip. There is also a patch of
black pigment above the outer half of the eye, and the area
of the cheek behind and below the eye is clouded with black.
On the sides of the neck and shoulders the pattern is obscure,
but where visible the spots show indications of transverse or
vertical arrangement. This transverse arrangement is very
clearly expressed upon the body, especially upon its thoracie
portion, where the spots are most manifestly transversely
or vertically elongated and not subcircular as in leopards
(F. pardus) and jaguars (F. onea), nor longitudinally elongate
Their coloration agrees substantially with that of the examples described
above. None, however, are quite so heavily pigmented, though in some
the tigrine nature of the pattern is equally strongly, if not more strongly,
in evidence.
Cubs of Lions and of Pumas. 439
as in some ounces (F. uncia). They are, moreover, set end
to end, one above the other, forming in some cases inter-
rupted double wavy stripes from two to three inches long,
extending from the median spinal line on to the belly. They
might perhaps be described as transverse chains of spots
comparable to the longitudinal chains of spots seen in
ocelots (F. pardalis). The duplication of the stripes is formed
by the fusion of the anterior rims of the superimposed
rosettes and of the posterior rims of the rosettes, the two
resulting streaks being separated by a paler area representing
the fused central portions of the rosettes. In some cases
these rosette-stripes extend uninterruptedly across the spine,
meeting those of the opposite side at an obtuse angle and
beimg divided from the preceding and succeeding rosette-
stripes by the also uninterrupted narrow intervening strip
of pale ground-colour.
On the right side of the body the fusion of the rosettes
into lines is less pronounced than on the left side. On the
spinal area where the rosettes are more heavily pigmented,
the pale central area of each is less clearly shown than on
the sides. On the sacral region and on the root of the tail
the rosettes show a distinct arrangement in four longitudinal
stripes, such as may be frequently seen in leopards, jaguars,
and ounces. On the upper part of the thighs the arrange-
ment of the rosettes is irregular, but on the lower and
posterior parts a longitudinal arrangement with an upward
angulation, such as is shown in a more emphatic manner in
tigers, is noticeable. Above and below the hocks the spots
are more solid and they extend on to the inner surface of
the limb above the hock and halfway down the front of the
leg between the hock and the paw. The paw is white and
spotless. The fore legs are more tawny than the hind legs
and are rather faintly rosetted from the elbow to the paw,
but on the inner side below the elbow very decided indications
of the brachial stripes so constant in cats are retained. The
tail is spotted from the root to the tip, the spots, which are
more strongly pronounced terminally, showing distinct
transverse arrangement (Pl. XIX.).
The second cub, which is larger and probably older than
the one just described, has the same style of pattern, but the
pattern is everywhere much fainter, the backs of the ears
being the only parts as heavily pigmented as in the other
example. In both specimens the hair on the neck is directed
backwards as in young tigers and leopards, and not forwards
on the sides of the neck aud forming a median cervical crest
as is the case, at all events usually, im adult leopards and
440 Mr. R. I. Pocock on the Pattern of the
tigers and also in adult lions. There are distinct signs of
the whorl of hair on the shoulder *.
It seems to me that the pattern of lion cubs affords very
strong support to Dr. Bonavia’s view respecting the origin
of the stripes of the tiger from the fusion of rosette-spots,
such as are seen in Asiatic leopards, into subvertical
or obliquely transverse lines. In tigers the stripes are
seldom quite vertical, except upon the upper part of the
shouldersand hind-quarters. Onthesides of the body beneath
the lumbar region they are oblique with a decided dorso-
ventral inclination backwards. Moreover, they seldom form
continuous streaks. Quite commonly each is broken up
into three constituents, a dorsal, a medio-lateral, and a
ventral, which frequently overlap at their juxtaposed extre-
mities. The medio-laterals are often suppressed on the
thoracic area behind the shoulder, as may be seen in two
specimens from Nepal now living in the Zoological Society’s
Gardens and in a “Siberian” specimen mounted in the British
Museum. It is not unusual to see one or more of the
above-mentioned constituent stripes continued by a row of
faint spots; or there may be rows of such small spots on
the interspaces between the stripes. Quite commonly, too,
one or more of the constituent stripes may be doubled in the
form of a long loop. More rarely where there is a greater
degree of fusion between the constituents a continuous
double stripe results ; and these double stripes may, I think,
be truthfully compared with what may be called the rosette-
stripes of lion cubs, the anterior and posterior dark rims of
the rosette-stripes in the lion corresponding respeotively to
the anterior and posterior moieties of the double or loop-
stripe in the tiger. This, I understand, is substantially
Dr. Bonavia’s interpretation of the origin of the pattern in
the tiger. He did not, however, cite the pattern of the
lion cubs in support of his hypothesis, but depended upon
that of leopards or jaguars, which supply less cogent evidence
in its favour, because in these species the rosettes do not fuse
into stripes as they do in lion cubs f.
* On account of the erroneous belief held by some people that young
lions are born with their eyes open, it may be added that the eyes in these
two specimens, as in all others I have seen, are closed, as is the case,
so far as I know, in all species of Felis.
+ Although I have attempted to show that ‘the pattern of lion cubs
bears out Dr. Bonavia’s views of the origin of stripes of tigers from
yosette-spots such as are seen in jaguars, I do not agree with that author
in believing that the pattern in Felide was originally of that type. It
must be admitted, I think, that HKimer was right in holding that the
pattern in these animals consisted primarily of longitudinal suipes.
Cubs of Lions and of Pumas. 44]
Another point in which the pattern of these lion cubs
differs from the pattern of leopards, jaguars, and ounces, and
approaches that of tigers is the following. In the three
spotted species if the spots upon the spine show definite
arrangement that arrangement is decidedly longitudinally
linear. It is always so, I believe, more or less upon the
sacral region, but less decidedly so on the lumbar and
thoracic areas. In tigers, on the contrary, the upper extre-
mities of the stripes almost invariably turn forwards on the
spine, and meeting from opposite sides in the middle line at
an angle form a series of A-shaped figures, a corresponding
shape being naturally assumed by the intervening spaces.
In the lion cub above described a similar pattern is quite
clearly indicated in certain places, the spine being crossed
from right to left by broad irregular A-shaped blotchy
stripes separated by narrower interspaces of a golden-tawny
hue. In tigers, of course, the interspaces are wider than
the stripes, the converse being true of the lion. This cireum-
stance is quite in keeping with the theory that the narrow
stripes of the tiger resulted from the antero-posterior com-
pression of the broader rosette-stripes such as are seen in
young lions.
On the tail and lower half of the thigh, as stated above,
the disposition of the spots in the lion cub also foreshadows
the stripes of these regions seen in tigers, but not seen, or
not nearly so clearly seen, in leopards, jaguars, and ounces.
Another truly tigrine feature is the presence of a white
patch over each eye. This is a feature in which the lion cub
differs not only from adult leopards, jaguars, and ounces, but
also from adult lions.
If there is any truth in the above-stated assumption as to
the origin of the tiger’s pattern and also in the claim that
the pattern of the lion cub is nearly intermediate in character
between that of the leopard and that of the tiger—and I can
see no strong argument against either,—it appears to me to
be necessary to conclude that these three species of Felis are
nearly related forms, a conclusion by no means obvious when
the coloration of the adults alone is considered. It was
largely no doubt owing to the differences in the coloration of
the adults that each of these species has been referred to a
distinct subgenus of the genus Felis.
I have elsewhere * pointed out that the prevalent belief in
near affinity between leopards and jaguars, so forcibly
suggested by their patterns, is confirmed by the resemblance
* P. Z. S. 1907, p. 677 (Oct.).
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 29
442 Mr. R. J. Pocock on the Pattern of the
between the roars of the two species; and as an additional
argument in favour of the view that lons and tigers are
related, it may be urged that the roar of a tiger is sufficiently
like that of a lion to be easily mistaken by those who have
never noted the differences between the two sounds. The
differences, of course, are obvious, but the similarity is also
unmistakable. The roar of the tiger, in fact, is much more
like the roar of the lion than it is like the roar of any other
species of Felis that I have heard.
It is, in my opinion, quite evident that too much import-
ance has been attached by earlier authors to absence of
pattern in adult examples of some species of Felis, The lion
and the puma, for example, are suggestively juxtaposed both
in Jardine’s monograph of this eroup, published in 1834, and
in that of Dr. Elhot, published in 1883; and Trouessart, even
as lately as 1904, kept the puma and the hon in the same
subgenus. But if appeal be made to the primary pattern
of these two species, as shown by the cubs, and not to the
secondary coloration of the adult, which is probably of
comparatively recent origin, very little support will be found
for the view that the two are nearly related forms.
I have only had the opportunity of examining the skins of
two newly born puma cubs, one in the collection of the
Zoological Society of London, the other in the Museum of
the Zoological Gardens at Clifton. Although the pattern of
the two is in the main identical, they differ in certain
respects so markedly from each other that it is probable that
one or both of the parents of the one were specifically or
subspecifically distinct from one or both of the parents of the
other *.
In the Zoological Society’s specimen (Pl. XX.) the ground-
colour is a brownish fawn, fading to white on the underside and
on the inner side of the limbs. There is white above the eyes,
on the upper lip, lower lip, and chin, the cheek below the post-
ocular stripe being a dirty white. The sides and top of the
muzzle are dark brown, and both the front and hind legs
from the elbow and hock to the tips of the toes are also
dark brown and without spots. On the side of the head a
black stripe extends backwards from the corner of the eye
beneath the ear, where it expands into a large dark patch.
Above the inner corner of each eye a black stripe runs back-
wards on to the summit of the head, and between these
are two narrower stripes. On the head these four stripes
apparently become zigzagged and more or less broken up.
* The puma (L" concolor) of the older authors has been divided into a
number of species and subspecies of late years. It would be extremely
interesting to know what the cubs of all these forms are like.
Cubs of Lions and of Pumas. 443
Unfortunately the hair on the posterior part of the head and
the anterior part of the nape is rubbed off, so it is impossible
to say for certain that the stripes extended right over this
region. That they probably did so, however, is attested by
the presence of four cervical stripes upon the posterior area of
the nape. The admedians of these, though narrow and faint,
can be traced on to the middle line between the shoulders.
The externals, on the other hand, are broad and very distinct,
the one on the right side being continued by a well-marked
suprascapular stripe, a stripe which is persistent in so many
of the smaller species of the genus Felis. On the sides of
the neck outside the external cervical stripe there are some
darker and fainter elongated spots or abbreviated stripes
running downwards towards the throat and chest. On the
shoulders below the suprascapular stripe there is also a large
transversely elongated spot, and below this some smaller
spots which become lost in the fuscous tint of the fore leg.
On the lumbar and sacral regions of the body there are three
very definite rows of large spots showing a decided tendency
to coalesce into longitudinal stripes. On the sides of the
body below the external of these (the dorso-lateral stripe)
there are about three rows of large spots of irregular shape
and sometimes more or less fused, especially on the abdominal
region, where they tend to run into short zigzag abbreviated
stripes. The thighs are spotted like the sides of the body.
On the thoracic region, both dorsally and laterally, the spots
are less symmetrically arranged than upon the abdominal
region, and the posterior part of the thoracic region is
marked dorsally with a pair of very large abbreviated stripes,
representing two or three fused spots, and inclining obliquely
backwards from a point close to the median dorsal line. The
greater part of the tail is lost; but the one inch of its basal
portion which persists exhibits a large dorsal spot.
The spots are all solid and of nearly uniform intensity.
The example in the Museum of the Clifton Zoological
Society differs from the one above described principally in
the complete absence of spots and stripes from the top of the
head and the nape of the neck, in the tawny hue of the legs,
and in the noticeably fainter tint of the spots on the sides of
the body as compared with those of the dorsal area. On
the head there is a short stripe above the inner corner of
the eye, a narrow stripe descending backwards from the
outer corner of the eye beneath the ear, and a brown patch
above the corner of the mouth. The back of the ear is jet-
black. Extending along the dorsal area from the shoulder
to the root of the tail there are three rows of solid spots,
those of the median or spinal row forming a more definite
Pa ie
444 On the Cubs of Lions and of Pumas.
line than those of the lateral row, which are about nine in
number on each side. Below the latter the spots form
roughly about three rows, but their arrangement is not
obviously either vertical or longitudinal. On the shoulders
the spots tend to run into abbreviated transverse bars; at
least, on to the root of the tail the median spinal stripe
extends. It expands along the tail into triangular blotches
constituting transverse bars, about eight in number; the tip
of the tail is black. The legs are practically without spots.
Broadly speaking, the pattern of the two specimens of
puma described above agrees with that of the examples
depicted in pl. 11. of Elliot’s ‘ Monograph of the Felidee.’
In the large size and small number of the spots, in their
solidity and definite arrangement in three rather widely
separated lines along the back, or, at least, along its lumbo-
sacral area, in the nature of the markings on the tail, on the
shoulders, and on the nape of the neck, when they persist
there, the pattern of the puma is quite different from that of
lions, leopards, jaguars, and ounces. On the evidence
supplied by the pattern, the puma cannot, I think, be regarded
as nearly related to any one of those species. Nor do I
know of any special point, apart from size, in which the
puma resembles the three spotted species just mentioned,
while practically the only likeness he presents to the lon is
the adult coloration, which must be set aside as a valueless
criterion of relationship.
It is a very difficult matter to decide to which group of
species of the genus Felis the puma is really related. The
pattern is not like that of any existing form; but in the
characters above enumerated, in which it differs from the
pattern of leopards and lions, it approaches the pattern of
several of the smaller species of the genus, species in which
the pattern is, im my opinion, of a more primitive type than
it is in the giants of the family.
I can find nothing in the structure of the skulls opposed
to the view here put forward, that the puma cannot be
associated with the group comprising tigers, lions, jaguars,
leopards, and probably ounces, nor anything in disaccord
with the suggestion that its nearest allies must be sought
amongst some of the smaller species. Rather the contrary.
And I do not think the resemblance between pumas and
“domestic cats” in the ossification of the hyoidean suspen-
sorium and in the expression of friendly feeling by
“purring”? should be altogether lost sight of in future
discussions of the subject *.
_ * In lions, tigers, and leopards, according to Mivart, the suspensorium
is ligamentous. ‘These species do not “ purr.”
Y
Descriptions and Records of Bees. 445
Summary.
1. The patterns of the cubs of lions and pumas are specific
characters. These species, usually described as
uniformly coloured, were formerly marked as their
cubs are marked and in no other way.
2. The pattern of lion cubs is intermediate between the
spotted pattern of leopards or jaguars and the striped
pattern of tigers.
3. From this it may be inferred that leopards (including
jaguars), lions, and tigers are nearly related one to
another,
4. On the assumption that spots preceded transverse stripes
in evolution, it may also be inferred that the stripes
of tigers originated from the fusion of rosettes into
transverse chains, as Dr. Bonavia maintained.
5. The pattern of puma cubs affords no support to the
belief that pumas are nearly allied either to leopards
or lions.
6. Rather, in my opinion, does the pattern of puma cubs
suggest that pumas may be regarded as large self-
coloured representatives of one of the groups of
smaller species of Felis, in the same way that lions
may be regarded as large and otherwise modified
representatives of a group exemplified by leopards.
EXPLANATION OF THE PLATES.
Puatre XIX.
Copy of a photograph of the dorsal view of a mounted lion cub in the
collection of the Bristol Museum, showing the formation of
transverse stripes from rosettes and attesting the relationship
between lions and leopards on the one hand, and lions and tigers
on the other.
PuaTE XX.
Drawing of the flat skin of a newly born puma cub in the collection of
the Zoological Society of London. The unshaded area on the
fore part of the neck shows where the hair has been rubbed
away. Since the tail was absent from this skin, the drawing of
that organ was copied from the example in the Museum of the
Clifton Zoological Gardens.
LXIII.—Descriptions and Records of Bees—X VII.
By I’. D. A. CockERELL, University of Colorado.
Osmia fulgida, Cresson, 1864.
THis species was described from the female. Mr. S. A.
Rohwer collected five males at Florissant, Colorado, June 15
to July 6, 1907; one was at flowers of Hrigeron. The male
is about 9 mm. long, very bright green, the abdomen shining ;
446 Mr. T. D. A. Cockerell—Descriptions and
form very slender, with the abdomen long and parallel-sided;
pubescence white throughout ; scape green; flagellum long
and slender, ferruginous, darkened above ; apical dorsal seg-
ment concave above, its apex broadly snout-like, not at all
notched ; a small spine on each side of the penultimate
segment.
Var. a.—Hind margins of abdominal segments narrowly
brilliant purple; vertex suffusedly crimson and golden. One
specimen, July 6.
By the narrow form and the structure of the abdomen,
this species is allied to Monumetha, but the antennx are
normal for Osmia.
Osmia pentstemonis, Ckll., 1906.
Var. a. 9.—Light hair of thorax strongly yellowish.
Florissant, Colorado, at flowers of Besseya plantaginea, June 1,
1907 (S. A. Rohwer).
Osmia physarie, sp. n.
g.—Length about 85 mm.
Blue-green, shining; head and thorax with copious long
hair, which is faintly yellowish dorsally, but otherwise white,
with no dark hairs intermixed ; flagellum dark reddish and
very strongly crenulate beneath ; legs black, not at all metallic,
their hair white, last tarsal joint terruginous; form of tarsi
normal ; tegule piceous with a large green spot. Wings
hyaline, a little dusky, but not at all reddish ; b. n. meeting
t.-m. Abdomen subglobose, with abundant dull white hair,
which tends to form marginal bands; sia@th dorsal segment
perfectly entire ; seventh bidentate ; first ventral entire.
In the Florissant table of Osmia (Bull. Am. Mus. Nat.
Hist. 1906) this runs to O. faceta, but that is very different
by the black hair on the abdomen. In the Boulder County
table (Univ. of Colo. Studies, 1907) it runs to O. prowima,
but that is a smaller and otherwise different species. The
antenne suggest O. ch/orops, but the flagellum is shorter, and
the hind tarsi are different; the wings also are differently
coloured.
Hab. Florissant, Colorado, at flowers of Physaria, June 1,
1907 (S. A. Rohwer).
Osmia seneciophila, sp. n.
3 .——-Length 9 mm.
Head and thorax very dark dull blue, a little greenish on
scutellum ; abdomen shining indigo-blue, closely punctured ;
hair of head and thorax white ; hair of cheeks white, except
just behind the eyes, where it is long and black; extreme
sides of vertex with a few black hairs, and even a very few
Records of Bees. 447
on front; flagellum long, dark, submoniliform; legs black,
not metallic, their hair partly black and partly pale; tarsi
normal, but the last joint red and rather elongate. Wings
hyaline, a little stained along the veins. Abdomen subglobose,
hind margins of segments very narrowly reddish ; third and
following segments with short black hair; sixth with a small
notch ; seventh bidentate ; first ventral normal ; third ventral
with a semicircle of long pale orange hairs in the median emar-
gination; fourth broadly elevated in the middle, the long
margin of the elevation ciliate with black hairs.
In the Florissant table runs to O. faceta, and it agrees
with certain Rocky Mountain males which I have called
jfaceta; but the real faceta is a steel-blue insect allied to
O. chalyba. In the Boulder County table it runs to
O. aprilina, which is, however, a smaller species, with the
hair a different colour and the flagellum not submoniliform.
Disregarding the few black hairs at sides of vertex, it runs
to Wheeleri and cyaneonitens. O. Wheelerd is easily separated
from it by the absence of the peculiar characters of the third
and fourth ventral segments, as well as the narrower form
and different colour of the abdomen; O. cyaneonitens is
considerably larger, and also lacks the semicircle of orange
hairs on the under side of the abdomen.
Hab. ¥lorissant, Colorado (type-locality), at flowers of
Senecio tridenticulata, June 26 (S. A. Rohwer). Top of
Las Vegas Range, New Mexico, 11,000 ft., June 28
( Cockerel/).
Osmia amala, sp. n.*
¢ .—-Length about 9 mm.
Head and thorax very densely punctured, but glittering,
dark blue, giving way to green on clypeus and mesothorax ;
abdomen broad-ovate, brilliant purple-blue, the hind margins
of the segments narrowly rufous ; legs black, without metallic
tints; hair of head and thorax dull white, with no dark
intermixed on thorax above; hair of cheeks black, and a few
black hairs on extreme sides of vertex; hair on pleura white,
but just under the wings, and posteriorly on sides of meta-
thorax, it is black; clypeus normal; antenne wholly dark,
flagellum very long, not at all crenulate or moniliform.
Wings hyaline; b. n. meeting t.-m. Hair of legs partly pale
and partly dark; hind femora with scattered black hairs ;
hair on inner side of hind basitarsus very dark purplish
fuscous. Sixth dorsal segment of abdomen entire, not at all
produced; seventh bidentate, with a brush of black hair
beneath each tooth; first ventral not emarginate; second
* Amala, a Malayan word for blue.
448 Mr. T. D. A. Cockerell—Desecriptions and
ventral and sides of third with fringes of long black hair ;
emargination of third with short whitish hair.
In the Florissant table runs to O. faceta, but differs by the
non-moniliform antenns and other characters. In the
Boulder County table runs to O. viridior, but is much
smaller and of a different colour. Disregarding the few
black hairs at sides of vertex, it runs to O. Wheeler?, which
it closely resembles ; but it is easily known from Wheelerz
by the black hair at sides of metathorax, structure of apex of
abdomen, &c. The first two small joints of the middle tarsi
are swollen, as in O. untversitatis and integrella: wniversttatis
has the hair on the sides of the metathorax light and the
abdomen greenish; ¢ntegredla is considerably larger than
amala and the abdomen is of a very different colour. In
size and the colour of the abdomen QO. amala resembles
O. coloradella.
Hab. Florissant, Colorado, June 30, 1907 (S. A. Rohwer).
Osmia mertensi@, sp. n.
3d .—Length about 8 mm.
Headand thorax green, glittering; abdomen deep blue-green,
very shiny, the hind margins of the segments concolorous ;
legs strongly tinged with green; hair of head and thorax
long and white; no dark hairs on thorax, but a few long
dark hairs on anterior part of cheeks ; hair of legs pale, with
some black intermixed ; hair of lind femora partly pale and
partly dark ; clypeus normal, covered with a dense brush of
hair; flagellum only moderately long, not at all moniliform,
ferruginous beneath ; wings strongly dusky; tarsi normal.
Abdomen with short black hair beyond second segment;
sixth notched ; seventh bidentate ; venter normal.
In the Florissant table runs to O. Wheeleri, of which it
may possibly be a variety, but the antenne are differently
coloured, and the sixth abdominal segment is only feebly
notched, and does not bulge at the sides as it does in
Wheeleri. Inthe Boulder County table it runs to O. aprilina,
but that has the pubescence, antennae, &c. quite different.
The vertex of mertensiw not only shows some black hairs at
the sides, but there are black hairs about the ocelli.
Hab. Florissant, Colorado, at flowers of Mertensia lanceo-
lata, June 19, 1907 (S. A. Rohwer).
Osmia enena, sp. n.*
$ Length about 9 mm.
Dark blue. Superficially similar to O. mertensie, but
differing thus: antenne considerably longer, the flagellum
* Enena, a Malay word meaning small,
Records of Bees. A49
faintly brownish beneath, but quite dark ; legs not metallic ;
abdomen narrower and less shining ; mar sinal cell longer and
narrower apically. The head is rather large ; there are dark
hairs about tle ocelli ; the thorax is wholly without dark
hair, There is a dark stain at the apex of the marginal cell.
Notch in sixth abdominal segment strong.
In the Florissant table it runs to O. faceta, but differs in
the antenne, &e. Compared with O. Wheeler?, the abdomen
is duller and the sixth segment differently shaped. The
clypeus is practically black; in Wheelert it is blue-green.
In the Boulder County table, O. enenaruns to aprilina, from
which it differs in the form of the abdomen, the colour of the
pubescence, Ke.
Hab. Florissant, Colorado, June 23, 1907 (S.A. Rohwer).
Osmia vallcola, sp. n.
6 .—Leneth about 8 mm.
Superficially similar to O. mertenste, but smaller, with the
flagellum wholly dark, the apical tooth of mandibles longer,
the head and thorax greenish blue; the vertex, front, and
thorax above with scattered very long black hairs; upper
part of pleura with the hair black; marginal cell longer ;
second submarginal cell very long. ‘The legs are strongly
metallic ; the abdomen is very shiny, and of the same tint
as that of mertensie; sixth segment rather feebly notched ;
venter and tarsi normal.
In the Florissant table runs to 3, and runs out because of
hair of pleura half black and half light. In the Boulder
County table it runs to O. propinqua, but the flagellum is
not moniliform, and there are many other differences.
(O. propinqua does oceur at Florissant, both sexes having
been taken by Mr. Rohwer from flowers of Salix brachy-
carpa, June 11, 1907.)
Hab. Florissant, Colorado, at flowers of Lzbes vallicola,
June 11, 1907 (S. a Rohwer).
Osmia nigrifrons, Cresson, 1878.
Three females at flowers of Senecio tridenticulatus, F loris-
sant, Colo., June 14, 1907 (S. A. Rohwer).
Osmia Wheelert, Ckll., 1906.
One male at flowers of Castilleta integra, Florissant, Colo.,
June 23, 1907 (S. A. Rohwer).
Osmia florissanticola, Ckll., 1906.
Two females at flowers of Aragallus Lamberti (sens. lat.),
Florissant, Colo., June 1907 (7. D. A. Cockerell).
450 Mr. T. D. A. Cockerell— Descriptions and
Osmia subtrevoris, Ckll., 1906.
Florissant, Colo., June 16, 1907 (S. A. Rohwer).
Sphecodes (Macheris) Rohwer?, sp. n
? .—Leneth a little over 5 mm.
Head and thorax black, shining, with white hair, which is
not at all infuscated dorsally; head transverse, front very
densely punctured, flagellum thick, testaceous beneath ;
mandibles simple, ferruginous except at extreme base; disk
of mesothorax with strong punctures, about as close though
perhaps not quite so large as in S. Cressonii (the thorax itself
is smaller than in Cressonit, and not quite the same tint,
appearing a sort of blue-black by contrast) ; area of meta-
thorax semilunar, well defined, with strong radiating ridges ;
tegule rufo-testaceous; wings dusky reddish; tarsi dark
brown, with glittering hairs. Abdomen shining, of a yellower
red than that of S. eusé¢ctus, the red including only the first
three segments (but these without black), the fourth and
following black; second and third segments punctured
basally.
Close to S. stygius, Rob., from Illinois, but separable by
the characters italicised.
Hab. Florissant, Colorado, at flowers of Salix brachycarpa,
July 7, 1907 (S. A. Rohwer). Mr. Rohwer also took Pro-
feraner rhois, Ckll., and Sphecodes Sophie, Ckll., at the flowers
of the same species of Saltz, at Florissant, the first June 2,
the second June 6.
While on the subject of Sphecodes, I take the opportunity
to offer a table of the larger species more or less like
S. dichrous. Many of these are very much alike superficially,
and are not easy to recognize without a table. ‘I'he table is
based on females.
When the abdomen is looked at from the
side, a strong constriction is seen dorsally
at the base of the second segment ;
punctures of mesothorax strong and
dense; area with strong longitudinal
TIAGOS! Gsaiutaancen ts ame hele see eae pecosensis, Ckll.
When the abdomen is looked at from the
side, the constriction is seen to be feeble
OLa@sent fice ee Men Pere eeire ili:
]. First abdominal segment with a blackish
discal spot ; disk of mesothorax shining,
with the punctures widely separated
First abdominal segment without a black-
Tel GUISCHI WO Fs Ras ao Ng 0b 0.04 do Soo 3.
2. Area of metathorax with widely separated
longitudinal ridges and a few cross ones ;
abdomen distinctly but not at all closely
bo
Records of Bees. 451
unctured, first segment very sparsel
te)
UU bere nanclos oododcbebecc Monee dichrous, Smith.
Area subcancellate with irregular wrinkles
(male abdomen largely red) .......... hesperellus, Ckll,
3. First abdominal segment with the apical
half very distinctly and rather closely
punctured ; area irregularly wrinkled ;
mesothorax rather more densely punc-
tured than in dichrous ; abdomen broad,
the apex hardly at all blackish........ arvensis, Patt., Rob.
First abdominal segment with very sparse
(sometimes hardly any) punctures .... 4.
. Second abdominal segment, beyond the
base, with very fine, relatively close,
ro
TES lar PUNChUNES. 37, wy sncpscenoieta eae 5.
Second abdominal segment, beyond the
base, with very sparse punctures ...... 6,
5. Abdomen broad, chestnut colour; wings
very dark and very yellow............ arcvensiformis, Cll.
Abdomen narrower, paler, the last seg-
ment blackened ; wings hyaline, the
apical half smoky, but not yellowish ;
aspect ofS. columbier.<cas incisors: lautus, Lovell & Ckll.
6. Apex of abdomen broadly blackish...... 7.
Apex scarcely or not blackish ........ yo ch
7. Larger; mesothorax very shiny; wings
iyalineees sae wets Nicaea een olympieus, Cll.
Smaller; mesothorax duller; wings with
a strong reddish suffusion ............ obscurans, Lovell & Ckll.
8. Face with a little white hair, but princi-
pally coarse black bristles ; abdomen long
and pale; mesothorax densely punctured,
the posterior middle sparsely ........ columbie, Ckll.
Face with hoary white hair ............ 9.
9, Mesothorax smaller, very shiny, less
densely punctured, the median sulcus
evident tege okie a ee eeu ee ee arroyanus, Ckll,
Mesothorax larger, duller, more densely
punctured, the median sulcus not or
hardly evident ...... SI no ectee persimilis, Lovell & Ckll.
The three species of Lovell and Cockerell are from Maine,
and are described in a paper which has been sent to ‘ Psyche’
for publication.
Falictus scrophularie, Ckll., 1906.
Mr. S. A. Rohwer took 17 females at Florissant in June
1907 ; of these, 15 were at flowers of Salix brachycarpa, one
at Ribes vallicola, and one at Taraxacum tararacum. The
thorax varies from blue-green to brassy green.
Andrena Portere, Ckll., 1900.
Florissant, 7 females, June 16, 1907 (S. A. Rohwer). Five
were at Ribes longiflorum*, two at R. pumilum.
* The mountain form of R. longiflorum is R. leiobotrys, Koehne, as I
learn from Dr. N. L. Britton. I understand that it will be regarded
as distinct in a forthcoming part of ‘ North American Flora.’
452 Miscellaneous.
MISCELLANEOUS.
Echinocrinus versus Archeocidaris.
By F. A. Barner, British Museum (Nat. Hist.).
Amoné many preliminary studies for a memoir entitled ‘ Triassic
Echinoderms of Bakony ” *, now passing through the press, I had
to make up my mind on this long-vexed question. My friend
Professor R. T. Jackson had to do the same for a forthcoming
memoir on Paleeozoic Echinoids, and on his asking my opinion I
sent him the MS. of the present note. Had he not urged its
publication, so as to clear the ground, I should have preferred
to let someone else play this ungrateful réle of nomenclatural
chiffonier.
The name Echinocrinus was proposed by Agassiz (1841, “ Obs.
sur les progres récents de l’hist. nat. des Echinodermes,” Monogr.
d’ Kchinodermes, ii. p. 15) for Cidaris Urii Fleming, and Cidarites
Nerei, Protei et priscus of Mimster, and some unpublished species.
The genus was thought by him to be a crinoid precursor of
Kchinoidea, possessing “ la forme sphéroidale des oursins avec des
ambulacres étroits et de longs piquans épineux comme certains
Cidaris,” and “ circonscrit dans les terrains de transition et dans les
terrains houillers.” On p. 20 Agassiz mentioned a new species of
Echinocrinus sent to him by Austin.
According to modern rules and customs it is clear that the name
Echinocrinus, unless preoccupied, which has never been maintained 7,
is perfectly valid, and that one of the four species mentioned by
name must be taken as genotype.
T. & T. Austin (1842, Ann. Nat. Hist. x. p. 111, Oct.) accepted
the name “Hchinocrinus, and mentioned under it HL. pomum, E. spi-
nosus, EH. anceps, and £. cidariformis? The last three were their
own MS. species, while the first, ascribed to Agassiz, doubtless
referred to the specimen above mentioned as sent by Austin to
Agassiz, and was also stillin MS. Consequently, except as showing
that the name was accepted, this paper has no bearing on the inter-
pretation of the genus.
The species #. spinosus and E. anceps were described by the
Austins in March 1843 (Ann. Nat. Hist. xi. p. 207), but not in
such a way as to permit of their recognition, while they did not
state their horizon. The latter was compared with LZ. pomum,
which, however, remained undescribed. Here occurs the first
objection to the generic name: “It appears to us,” they wrote,
“that the name of our genus Sycocrinites and that of the Echino-
crinus of Professor Agassiz require amendment, as their terminations
imply affinities which do not exist.” The meaning of this is not
very clear, since by the title of their paper they still retained them
under Crinoidea, and since by the words “ Column unknown ” they
* ‘Resultate der wissenschaftl. Erforschung des Balatonsees,’ i. Bd.
1 Th., Pal. Anh.
+ Echinoencrinus, H. v. Meyer, 1826, is unfortunately similar, but it is
far from identical.
Miscellaneous. 453
implied that both genera were at any rate Pelmatozoa. Unpublished
drawings by the Austins of some of their species of Sycocrinites
have convinced me that those species were Crinoids after all. The
criticism, therefore, was not at that time well founded.
In 1844 the name Zehinocrinus was still maintained by no less a
person than M‘Coy (‘Synops. Carb. Foss. Ireland’ *, p. 173), who
then gave a systematic diagnosis, recognizing the genus as an
Kehinoid, and distinguishing it from Palechinus on the one hand
and Cidaris on the other. Under this name he proceeded to
describe 4. Munsterianus? (Kon.), E. glabrispina (Phill.), E. tri-
serialis sp. n., H. Urit (Flem., incl. Cidaris benburbiensis Portlock),
and H. vetustus (Phill.). Now, if by chance any objection were
raised to Hehinocrinus Agassiz, on the ground of insufficient or
misleading description, and if the Austins’ use of the name were
ignored on similar grounds, the objector would still be compelled to
accept the name as here confirmed by M‘Coy; and since all the
species described by M‘Coy were based on radioles only, with the
exception of H. Uri, that species would naturally be selected as
genotype, just as it would in the case of Agassiz’ name tT. So far
the situation is unchanged. M‘Coy, however, took the unfortunate
step of mentioning that he “had long + ago distinguished this
species in” his “* MSS. under the name of Archeocidaris.” Ob-
viously this casual remark could not give any sanction to the name
Archeocidaris, nor was it intended to do so. On the contrary, it
is a question whether this mention did not put the name out of
court at once and for ever as a mere homonym of Echinocrinus §.
Once these fossils were clearly understood to be Echinoids and
unrelated to Crinoids the obvious appropriateness of M‘Coy’s MS.
name led to its adoption by some authors. Thus, in 1845, Murchi-
son, Verneuil, and Keyserling (‘ Geol. Russia,’ 1. pp. xiv & 396), in
assigning Cudaris rossicus Buch, to the newly established genus,
asserted their preference for the name Arch@ocidaris. Nowhere in
the volume, however, is this species called anything but Crdaris (or
Cidarites) rossicus. It would, therefore, scarcely be possible to
regard the name Archeocidaris as given any standing by those
authors; nor, if occasion arose, could it be restricted on these
grounds to any genus that might be established with C. rossicus as
* T am aware that, strictly speaking, this work was first published in
1862 by Messrs. Williams and Norgate ; but ntany copies were privately
distributed by Sir Richard Griffith towards the close of 1844 and
subsequently. Desor, however, had not yet seen it in 1857.
+ If anyone were to insist on the first species mentioned being taken
as the genotype, he would select £. Uri if he accepted Agassiz, but
E. glabrispina if he accepted only M‘Coy. 2. Munsterianus being
marked with a ? could not be selected.
{ None the less in 1842 he called this species merely Cidaris Urit.
See R. [ J.] Griffith, ‘ Notice respecting the Fossils of the Mountain Lime-
stone of Ireland &c.,’ p. 12: printed Dublin, 1842.
§ In the legend to the lithographed plates the name <Archeocidaris,
which had been printed, was erased, and the name Echinocrinus inserted
by hand.
A454 Miscellaneous.
genotype. Admittedly the name remains a pure homonym of
Lchinocrwmus.
That the view just expressed was the one at that time adopted by a
man of sane judgment is proved by Bronn’s “ Index Palzontologicus”
(‘ Nomenclator,’ p. 443; 1848), where Echinocrinus is definitely
accepted with Archeocidaris asa synonym. To this genus Bronn
refers all the species hitherto mentioned, except MS. names and
except L. triserialis M‘Coy, while he adds Cidaris deucalionis
Eichw. as a possible synonym of #. rossicus.
In November 1846, however, Desor, in Agassiz & Desor (* Cat.
raisonnée des Kch.,” Ann. Sci. Nat. (3) vi. p. 340) founded the
genus Paleocidaris for the Cidaris Neret, Protei, and prisca of
Miinster, previously placed by Agassiz in Echinocrinus. That the
intention was simply to supplant the name Hchinocrinus by one
more appropriate to the echinoid nature of the fossils, appears
from a note in Desor’s nos (p. 154), as well as from the
omission of Hchinocrinus from the ‘Catalogue.’ Since, however, no
such statement was made at the time, it might be possible to assign
to Echinocrinus the genotype Cidaris Uri, and to Paleocidaris the
genotype Cidaris Nere?, and ultimately, should subdivision of genera
proceed on these lines, to use both names. Down to this present,
however, no writer has doubted that C. Meret and C. Ur are con-
generic, and Paleocidaris must therefore be regarded as either a
homonym or a synonym of Hehinocrinus.
The name Paleocidaris was no doubt proposed in ignorance of
M‘ Coy’s sname Archeocidaris, and M‘Coy (1849, Ann. chat Hist.
(2) ili. p. 252) rightly pointed out that, if Hchinocrinus were to
Ce place on any grounds to a later name, then Archeocidaris was
prior to Paleocidaris; and this view was accepted by Desor (1857,
‘Synopsis, p. 154). M‘Coy’s reasons for reversing his own previous
action were thus expressed :—(1) ‘‘ Agassiz neither indicated the
affinities nor gave any descriptive notice of the genus Echinocrinus,
while I have done both for my Archeocidarts.” (2) ‘ Several of
the continental geologists have not followed my example in rejecting
my own name, but prefer Archwocidaris.” (38) Agassiz and Desor,
as above mentioned, have given up Agassiz’ own name. Of these
reasons the only one that could have any validity is the first ; but
the statement is not entirely correct, since Agassiz did indicate
what, in his opinion, were the affinities of Pchinoer tue. and he did
give a descriptive notice two lines long. The incorrectness of his
opinion and the insufficiency of his notice were fully compensated
for by his mention of four well-known species. No contemporary
worker failed to understand precisely what Agassiz meant by
Echinocrinus. That the name accords ill with present knowledge
may be admitted, but it is no more misleading than M‘Coy’s own
name Codaster, proposed in the very same paper for a fossil
“‘ obviously allied to Pentremites,” or than Agelacrinus for an Edrio-
asteroid, or Phrissocystis for an Echinoid. As for the ‘several
continental geologists” of M/‘Coy’s paragraph (2), I have been
unable to discover them; but it may be added that d’Orbigny
Miscellaneous. 455
(1850, ‘ Prodrome,’i. p. 154) adopted Echinocrinus with Palwocidaris
as a synonym, and left Archeocidaris out of consideration, presumably
as a mere MS. name.
Thus far, then, the result of our examination is to reinstate
Echinocrinus Agassiz, with Archeocidaris as a pure homonym and
Paleocidarts as a synonym. The name Archeocidaride will also
have to go. There remain for consideration various attempts at
subdividing the original genus Hehinocrinus, or the relegation of
certain species to other genera.
Desor (1857, ‘Synopsis, p. 155) distinguished a new genus
Eocidaris from Archeocidaris (i, e. Echinocrinus) on the ground
that the primary tubercles of the interambulacrals were devoid of a
halo (‘second anneau”), and he included in this genus, inter alia,
KE. Vernewlana (King), EF. rossica (Buch), and #. Munsteriana
(Kon.), all species which had at one time been referred to Echino-
erinus or Archeocidaris.
To the interpretation of Hocidaris I shall ask permission to recur
in a subsequent note, and confine myself here to pointing out that
the result of Desor’s action at any rate was to restrict Echinocrinus
to species with primary interambulacral tubercles perforate, non-
crenelate, scrobiculate, and, above all, annulate (7. e. with a “ basal
terrace ”’).
The next step appears to have been that taken by Meek and
Worthen (1869, Proce. Acad. Nat. Sci. Philadelphia), who, after
describing Hocidaris ? squamosa (p. 79), showed its distinction from
Archeocidaris and Kocidaris, and tentatively proposed the name
Lepidocidaris (p. 81). This resembles Hocidaris in the absence of
a basal terrace from the interambulacral primary tubercles, but is
presumed to differ from it in the presence of eight columns of inter-
ambulacrals at the ambitus and demiplates alternating with primary
ambulacrals. The structure of these parts in Hocidaris is still
unknown ; but the genus Lepidocidaris has been generally accepted
(see Jackson, 1896, p. 220; and Tornquist, 1897, pp. 51=773),.
In 1883 Pomel (‘ Class. Meth.’ p. 113) erected the genus Cidaro-
tropus, with genotype Archeocidaris Worthenit Hall. Pomel drew
no distinction between the interambulacral plates, but based the
genus on the two series of regular primary ambulacrals, each with
two pores in the middle of it. Our knowledge of these structures
in the original species of Hchinocrinus is not enough to warrant a
distinction on these grounds.
Tornquist, however (1897 *, pp. 52=774), has divided the species
into two main groups. That including the genotype £, Urii,
together with HL. Nerei, EH. Werveket, and E. Grueneri, possesses
interambulacrals with a clear basal terrace from which thick wedge-
shaped ridges radiate to the margin of the plate. The other group,
which includes £. rossicus, E. Trautscholdi, E. Wortheni, E. biangu-
latus, E. megastylus, and E. Norwoodi, has interambulacrals with a
basal terrace well developed on some, but sometimes entirely absent
* “Tas fossilfiihrende Untercarbon &c.,” Abh. geol. Karte Elsass, y,
456 Miscellaneous.
from others, with smooth scrobicule around which are numerous
small scrobicular tubercles in one or more circles; these tubercles
are usually especially numerous on the adoral and adapical margins,
particularly the latter, where they form many close-serried rows.
E. Munsterianus is, according to Tornquist, the representative of
yet a third group, the characters of which he does not give. Does
he know more of it than the radioles ?
Tornquist has given no generic or subgeneric names to the groups
defined by him, but Lambert (1900*, p. 42), while retaining for
the first group the name Eehinocrinus (in accordance with the views
above expressed), has suggested the resuscitation of Archwocidaris
for the second group. This, of course, is quite inadmissible, as
already explained (p. 453). If a name be required, ‘“ Cidaro-
tropus”’ is ready to hand, with genotype C. Worthenr. But it is
really too early to make this division. Unfortunately so many
species have been based on radioles or on isolated interambulacral
plates that they cannot be referred with any security to either of
these groups. Perhaps we may be allowed to retain the word
“ Archeocidaris ” as a convenient appellation for such doubtful cases,
much as the word “ Ammonites ” still has its uses.
The following statement summarizes the conclusions with regard
to the principal generic names mentioned in this study :—
Ecurinocrinus Agassiz, 1841.
Genotype Cidarts Uri Fleming.
Syn. Archeocidaris, M‘Coy, MS., 1844.
Paleocidaris, Desor, 1846.
Archeocidaris, M‘Coy, 1849.
Crparotrropus Pomel, 1883.
Genotype Archwocidaris Worthent Hall.
Syn. Eocidaris Desor (pars), 1857.
Archeocidaris Lambert (non M‘Coy), 1900.
Lepirpocrparis Meek & Worthen, 1869, em. Jackson, 1896.
Genotype Hocidaris? squamosa Meek & Worthen, 1869.
Of these genera, Echinocrinus and Lepidocidaris are well estab-
lished, but Cidarotropus rests, for the present at any rate, on an
insecure basis, and, if not accepted, must with its synonyms be added
to the list of synonyms of Hchinocrinus.
The choice of a name to replace Archeocidaride is therefore
limited to two. Echinocrinide, the natural successor, does not
suggest the systematic position of the Family so well as Lermo-
crparip#, and I therefore propose to use the latter.
* “Etude sur quelques Echin. de 1’Infra-Lias &c.,” Bull. Soc. Sci.
Yonne, liii. lev semestre, pt. 2.
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LXIV.—A Revision of the Medusce belonging to the Family
Laodiceide. By Epwarp T. Browne, University
College, London.
THIS is an old family-name to which I have given a new
definition. Although the character selected is a conspicuous
one, it has not hitherto taken an important position in classi-
fication, but has been chiefly used as a character for distin-
guishing certain genera. This character, now selected for
the family, is the presence of cordyli, commonly called
sensory clubs, on the margin of the umbrella.
Some of the genera which possess cordyli were placed by
Heckel amongst the Thaumantide, of which the Laodiceidze
formed a subfamily, and was distinguished from the other
subfamilies not by the presence of cordyli, but by the number
of radial canals. The other genera, on account of their
having branched radial canals, were placed in the Cannotide,
a family which Maas (1904) has recently revised.
In the family Laodiceidea I have placed the following
genera :—Laodice, Staurophora, Ptychogena, Staurodiscus,
Toxorchis, and Melicertissa. 'The characters of these genera
have been revised, but the revision has not led to a trans-
ference of species. The species of all the genera have been
subjected to an impartial examination, which has resulted in
a reduction of their number. For the purpose of making
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 30
458 Mr. E. T. Browne on the Meduse
this revision as complete as possible I have briefly given the
characters of new species of Stawrophora and Ptychogena.
A cordylus is an organ of a well-marked character, and
when once recognized it is not likely to be mistaken for
anything else on the margin of the umbrella. It is quite
distinct from marginal bulbs and tubercles or sprouting
tentacles and cirri. Its shape varies slightly in different
genera, but it always has a clear translucent appearance,
without any coloration, and is free from nematocysts. It is
also without otoliths and such concretions as are generally
found in sense-organs. Its function, however, has not yet
been definitely found out, but it is generally regarded as a
sensory organ. The first adequate description of a cordylus
was given by Brooks (1895), to whom the sensory theory is
due.
Hartlaub’s positive statement (1897) that the cordyli of
Staurophora develop into tentacles led me to carefully examine
early and intermediate stages of Laodice. If cordyli are the
forerunners of tentacles one would naturally expect to see
them in the earliest stage or in the very early stages; but
they do not make their appearance until the Medusa has at
least trebled the original number of its tentacles.
After searching the margins of the umbrella of several
dozen young Laodice, I did find two specimens which showed
cordyli being converted into tentacles. They showed, how-
ever, an exception to the normal course of development of a
tentacle which needs an explanation.
When there is ample room between two tentacles one finds
a cordylus, a cirrus, and a tentacular bud in a single row and
isolated from one another. The tentacular bud increases in
size until it becomes a bulb, from which sprouts out the
tentacle. Under this condition there is not the slightest
indication of a cordylus becoming converted into a tentacle.
The stalk of the cordylus arises direct from the margin of the
umbrella and does not touch the tentacular bulb.
In most young specimens the interval between two tentacles
has frequently the appearance of being overcrowded, owing
to the marginal appendages developing faster than the margin
of the umbrella. The tentacular buds arise alongside of, or
even underneath, the stalk of a cordylus, so that a cordylus is
often seen on the side, or on the top, of a tentacular bulb.
One specimen was seen with a number of buds and bulbs with
tentacles developing ; each bulb had a cirrus on its outer side
and a cordylus on its inner side. It was evident that the
tentacular bud had forced its way up between the cirrus and
belonging to the Family Laodiceide. 459
the cordylus. On the development of the bud into a bulb the
cirrus and cordylus were carried up on to the side of the
bulb. It is rare to find a cirrus and cordylus on the side of
a basal bulb of a large tentacle, so that these organs either
change their position or disappear. They cannot develop into
a tentacle, because the tentacle is already formed.
In the two specimens showing the cordyli being converted
into tentacles it was fairly evident that the tentacular bud
made its appearance right underneath the already fully deve-
loped cordylus. There were a sufficient number of bulbs with
cordyli to trace out the various stages of growth. One bulb
showed very distinctly the conical apex of the sprouting
tentacle beneath the translucent stalk of a cordylus, and later
stages showed the translucent cells of the cordyli becoming
opaque as the tentacles advanced in size. The cordylus in
the process of conversion becomes very large, and finally
loses its characteristic shape. It seems to me that the cells
of the cordylus are converted into tentacular cells, and as
soon as that process is completed the rounded end of the
cordylus becomes pointed and indistinguishable from an
ordinary half-grown tentacle.
As the conversion of cordyli into tentacles was only seen in
two young stages, it is probably due to the cordyli being in
the way of rapidly growing tentacles, and consequently they
were absorbed.
A time comes when tentacular growth stops and the bulbs
remain in an arrested state of development. This, I think,
accounts for some adult specimens having their cordyli upon
small bulbs and also upon bulbs with ocelli.
Family Laodiceide, L. Agassiz, 1862.
Character of the Family.—Leptomeduse with cordyli,
commonly called sensory clubs, on the margin of the
umbrella.
Genus Laopice, Lesson, 1843.
Generic character.—Laodiceide with four radial canals ;
with a central stomach and mouth; with ocelli on the basal
bulbs of the tentacles.
This is the best-known genus of the family. Although I
have excluded several species which were formerly placed in
the genus and reduced others to synonyms, still I am not
quite satisfied with the result, owing to the difficulty of
. 30*
460 Mr. E. T. Browne on the Medusee
finding suitable characters for the determination of the
species. As the means of distinguishing the species the
following characters are used :—
a. The presence or absence of cirri.
b. The presence or absence of a spur at the base of the
tentacles.
c. The number of cordyli between the tentacles.
d. The shape of the gonads.
Laodice undulata (Forbes & Goodsir), 1851.
Thaumantias undulata, Forbes & Goodsir, 1851, p. 313, pl. x. fig. 7.
Thaumantias confluens, Forbes & Goodsir, 1851, p. 314, pl. x. fig. 8.
Thaumantias mediterranea, Gegenbaur, 1856, p. 237, Taf. viii. figs, 1-
Cosmetira punctata, Heeckel, 1864, p. 334.
Laodice calcarata, Browne, 1898, p. 823, pl. xlix. fig. 4.
Laodice cruciata, Maas, 1904, p. 18.
5.
Laodice calearata, A. Agassiz, 1862.
Laodicea calcarata, A. Agassiz, 1862, p. 350.
Lafoea calearata, A. Agassiz, 1865, p. 122, figs. 184-194,
Laodice calearata, Heckel, 1879, p. 134.
Laodice calearata, Brooks, 1895, p. 287, pl. xvii.
Laodice ulothrix (Heckel), 1877.
Cosmetira ulothrix, Heeckel, 1877.
Laodice ulothrix, Heckel, 1879, p. 183, Taf. viii. figs. 5-7.
Laodice ulothrix, Mayer, 1900, p. 49; Mayer, 1904, p. 14, pl.iv. fig. 30.
In 1851 Forbes and Goodsir described as new species
Thaumantias undulata and Thaumantias confluens, which
they found on the west coast of Scotland. I consider 7’. con-
jfluens to be an earlier stage of J. undulata. It is quite
evident from the description and figure that 7. undulata
belongs to the genus Laodice. The specimens were seen
alive, and in their description the authors state that each
tentacle “springs from a bulbous base, bearing a small but
distinct black ocellus. Between each pair of tentacula is a
minute, transparent, mobile, pedunculated tubercle. [The
figure shows these tubercles, which have the appearance of
roughly drawn cordyli.} Down the four gastrovascular
canals, very nearly from their divergence to the margin
of the umbrella, run the four linear genital glands, tinged
with rose-colour. They are very peculiarly formed, each
hanging from the surface of the subumbrella in the shape of
a pair of undulated membranous curtains, strikingly reminding
us of the appearance presented by Staurophora, but differing
in their nature ; for, in the animal we are describing, they are
belonging to the Family Laodiceide. 461
assuredly distinct from the stomach-lobes. The stomach is
rather large and quadrangularly campanulate, rose-coloured,
and slightly fimbriated at the margins.” ‘he umbrella is
hemispherical, about 14 inches (38 mm.) in diameter. The
tentacles are about 160 in number. ‘The authors do not
mention or figure cirri, which should have been present in the
specimens.
The Laodice which I have found on the British coasts I
called Laodice calcarata, and used the name before I had
seen the paper by Forbes and Goodsir. Otherwise I should
have no doubt ealled the British form Laodice undulata.
The fact that Forbes and Goodsir mention the presence of
one ‘“‘ pedunculated tubercle’ between every two tentacles,
by which they evidently mean a cordylus, shows, I think,
clearly that they had found a Laodice, and, so far as I know,
there is only one species of Laodice on the British coasts.
Forbes and Goodsir say nothing whatever about cirri,
which they would have seen if the living specimens had
been caretully examined. In preserved specimens cirri are
sometimes scarce and also the cordyli, as these organs
are rather fragile. In the second species, Thaumantias
confluens, the figure of the margin of the umbrella does show
two or three projections between the tentacles. ‘They may
possibly represent the bases of broken off cirri and a cordylus.
The authors state that this species has also pedunculated
tubercles.
To Gegenbaur the credit must be given for the first
adequate description with good figures of a Laodice when he
described Thaumantias mediterranea, 1856.
It is futile to consider Aledusa cruciata of Forskal, 1775, as
a Laodice, because the essential character of the family is not
mentioned or figured. Hzeckel, moreover, has caused utter
confusion by placing several species clearly belonging to
other genera as synonyms of Laodice cruciata. ‘lhe law of
priority is carried too far when it is extended to species
which have never been either described or figured, so as to
indicate the character of the family or genus.
In the above list of references there are six distinct specific
names; three of them may be safely regarded as synonyms,
I have made several endeavours to find a single character or
combinations of characters whereby the remaining three
species—L. undulata, L. calcarata, and L. ulothria—could be
distinguished from each other and readily recognized. When
the descriptions and figures of these species have been
analyzed one finds that new figures, with more detailed
descriptions based upon more specimens, are needed. It is
462 Mr. E. T. Browne on the Medusa
solely for that reason that I have refrained from joining all
the above-mentioned species under one name.
The normal number of cordyli between every two tentacles
in the British form is one, and one is also found in the Medi-
terranean form. Jaodice ulothriz, according to Heckel’s
figure, has two cordyli, but Mayer has described specimens
with one cordylus between every two tentacles. Laodice
calcarata, according to Agassiz, has one or two cordyli
between the tentacles, but Brooks mentions specimens with
only one. It is evident that there is one cordylus between
every two tentacles and that some specimens may have one
or two; but there is no evidence that any of the North-
Atlantic species have always two cordyli between every two
tentacles. ‘The same is the case with the cirri, either one or
two between every two tentacles. Allowances must be made
for development and also for breakage in preserved specimens.
Brooks records a variety of Laodice without cirri from
the Bahamas, but Mayer records specimens with cirri from
the same region.
The ocelli are certainly very variable in number. In some
specimens every tentacle is provided with an ocellus, whereas
in others comparatively only a few tentacles have ocelli.
Gegenbaur figures an ocellus at the base of the cordylus in
L. mediterranea, and Brooks also mentions a variety with
ocelli in the same position from the Bahamas. The ocellus
belongs really to a tentacular bulb in an arrested state of
development, upon which the cordylus is situated. I have
found that the British form of Laodice has a very variable
number of irregularly distributed ocelli, so that they are of
little use for a specific character.
The length of the gonads along the radial canals is useless
for a specific character, as the length depends upon growth.
There is certainly a difference in colour, but colour unfor-
tunately usually disappears after preservation, and, moreover,
the descriptions do not always state whether the colour
described is that of the living medusa or of a specimen in
alcohol or some other fluid. I haveseen large living specimens
of the British form which were quite colourless, and other
specimens from the same locality with pink gonads. ‘There
is, however, a tendency for the European forms to have
pinkish gonads and the American forms to have dark yellowish
to brown gonads, Mayer describes L. ulothriz from the
Bahamas as being dull pink, brownish, or greenish white, so
that it appears to me that colour is of little use as a specific
character.
On bringing together the characters of the three species
belonging to the Family Laodiceide. 463
found in the North Atlantic and Mediterranean, one finds
that between every two tentacles there are always one
cordylus (rarely two) and either one or two cirri. The ocelli
are variable in number and not quite constant in position,
and their colour is variable—dark brown, dark violet, or
black. The colour of gonads and other organs is also vari-
able—dark yellow, brown, pink, or pale violet. ‘The tentacles
are numerous and have, when fully developed, an endodermal
basal spur, which is variable in length and shape. The
gonads, when mature, form undulating bands upon the radial
canals.
Distribution. North Atlantic; Europe, British coasts
(L. undulata, Forbes and Goodsir ; L. calcarata, Browne).
Mediterranean (L. mediterranea, Gegenbaur ; L, cruciata,
Maas).
Canary Is. (ZL. ulothrix, Heckel).
Bahamas (L. ulothriz, Mayer).
North Atlantic ; American coast (Z. calcarata, Agassiz,
Brooks, Hargitt).
Tortugas, off Florida (LZ. ulothriz, Mayer).
The Hydroids belonging to Laodice calcarata and
Laodice undulata.
A. Agassiz (1865, p. 124) gives a brief description with a
figure of the hydroid which he believed to belong to Laodice
calcarata. It is necessary, however, to criticize this con-
nexion of the hydroid with the medusa, as it is a matter of
some importance.
Agassiz discovered a small hydroid which he considered
to belong to the genus Lafoea, hence the name Lajfoea calcarata.
The hydroid is a small creeping form and was found just
below low-tide mark in Buzzard’s Bay, Naushon. ‘The
hydranths are arranged “ in a quincunx manner on both sides
of a long slender creeping stolon, which does not branch.”
The figure shows that the perisarc is tube-like, and there is
no evidence of an operculum. ‘The hydroid has a few very
large gonothecew, inside of which develop meduse. The
medusa on liberation has ‘‘ two long tentacles, two slightly
developed ones, and four more hardly perceptible in the
middle of the space between the chymiferous tubes (radial
canals). The basal bulbs of the two tentacles and the other
six tentacular buds each have one dark pigment-spot. This
medusa on liberation from its hydroid has only two long
tentacles, no cirri, and no cordyli.
The next stage mentioned is much older than the earliest
464 Mr. E. T. Browne on the Meduse
stage. As there is no evidence to show that Agassiz reared
the meduse in an aquarium, I presume that this later stage
was taken out of the sea. It has sixteen tentacles and a
cirrus between every two tentacles. The basal bulbs of all
the tentacles are provided with ocelli; but there are no
cordyli. It seems to me that the similarity between the
above two stages is the presence of ocelli, and it is well to
remember that there are other genera besides Laodice with
dark ocelli on the basal bulbs.
Metschnikoff (1886, p. 83, Taf. iv. figs. 17-31, Taf. v.
fig. 1) has given an excellent description of the development
of the ova of Laodice cruciata and splendid figures of the
hydroids which he reared from the ova. His work is entirely
embryological, and no description of the medusa is given. I[
presume he means Laodice cruciata according to Heckel, and
that his species was really Laodice mediterranea of Gegen-
baur ; for this is the only species of Laodice among Heckel’s
many synonyms. The hydroid which Metschnikoff reared is
similar to Cuspidella humilis, Hincks.
Hincks (1568) described three species of Cuspidella—
C. humilis, C. costata, and C. grandis. The descriptions are
based upon the shape of the hydrothece. It is evident to me
that the shape of the hydrotheca of Cuspidella and its allies
is not sufficient to base specific characters upon, and that the
structure of the hydranth must be taken into consideration,
and also the gonosome. It is quite likely that Cuspidella
costata is only another form of C. humilis. For the purpose
in view it is sufficient to know that Metschnikoff reared
from the ovum of Laodice a hydroid belonging to the genus
Cuspidella, which is distinguished generically from Agassiz’s
Lafoea by the presence of an operculum on the top of the
hydrotheca.
During June 1906 I received a letter from Miss M. Delap,
of Valencia Island, stating that she had kept a colony of
Cuspidella costata under observation and had seen the colony
liberate meduse. Later on I received drawings of the
hydroid and its medusa and also specimens. The hydro-
theca is like Hincks’s figure and has a few transverse rings
and an operculum. ‘The gonotheca is somewhat similar to
the hydrotheca, but is about twice the length and is without
transverse rings. The figure drawn by Miss Delap shows
two medusa-buds inside the gonotheca and a medusa just
escaping through the operculum. The medusa on liberation
has two opposite perradial tentacles and two opposite per-
radial tentacular bulbs. On each side of the two tentacles
there is a cirrus, adradial in position, and cirrus-buds occupy
belonging to the Family Laodiceidee. 465
the other four adradial positions. The umbrella is nearly as
broad as high, about 1 mm., and has a few nematocysts
scattered over the exumbrella. The four perradial tentacular
bulbs have black ocelli on their inner side. The medusa on
liberation is without cordyli.
Metschnikoff figures two specimens of the young hydroid,
one with a short hydrotheca and the other with a long hydro-
theca; both are without transverse rings, which are generally
considered to be lines of growth. Miss Delap’s figure is
similar to Metschnikoff’s hydroid with a short hydrotheca,
but shows the transverse rings.
The comparison of Cuspidella costata with Lafoea calcarata
of Agassiz shows that the two hydroids are not of the same
genus. The hydrotheca of Lafoea calcarata is without an
operculum, and its gonotheca is also without an operculum
and is quite different in shape. Agassiz’s hydroid is not a
Cuspidella, and it is not a true Lafoea, because it liberates
medusee. Lafoea has a peculiar gonosome, which until
recently was regarded as a distinct hydroid, generically known
as Coppinia. Moreover, the meduse liberated from these
two hydroids are not similar. Agassiz’s medusa has two
tentacles and six tentacular buds, all with ocelli, and no cirri.
Miss Delap’s medusa has two tentacles and two tentacular
buds, all with ocelli, and four cirri, Hither Agassiz’s hydroid
does not belong to Laodice calcarata, or if it does, then the
adult medusze found on the American coast and on the British
coast should show specific differences, sufficiently conspicuous
to distinguish one from the other.
I have in my collection of British medusze some young
stages of Laodice taken in tow-nets at Valencia in 1897 and
at the Scilly Isles in 1899 and 1903. ‘The earliest stage,
about 1°25 mm. in diameter, has four perradial tentacles, each
with a black ocellus on the basal bulb, four interradial, eight
adradial, and a few scattered buds or bulbs, all without
ocelli. Between every two bulbs there is generally a cirrus ;
but there is not the slightest trace of a cordylus. As develop-
ment proceeds tentacles sprout out from the bulbs, more buds
or bulbs appear, and more cirri come into existence. It is
not until the umbrella is 3-4 mm. in diameter that cordyli
are clearly recognizable. (Many of the early stages were
examined alive.)
Laodice is the only medusa on the British coasts with
black ocelli on the inner side of the basal bulbs and with
cirri, so that these early stages, without cordyli, are not likely
to belong to another genus. ‘T'he presence of black ocelli and
cirri in the medusa liberated from Cuspidella costata indicates
466 Mr. E. T. Browne on the Meduse
a Laodice, and there is no reason for supposing that after a
little further development it would not become similar to the
earliest stage in my series of young Laodice.
The absence of cirri in the medusa liberated from Lafoea
calcarata seems to indicate that it 1s not a Laodice. The
later stage, with cirri, described by Agassiz, has the characters
of a Laodice, and agrees with one of my early stages before
the cordyli begin to develop ; but, as I have already pointed
out, there is no evidence that this particular specimen was
reared from the hydroid.
Laodice indica, Browne, 1905, p. 136, pl. i. fig. 5, pl. iv.
figs. (—E1:
This species is very much like Laodice undulata, but the
tentacles are without a basal spur. Cirri present. One
cordylus between every two tentacles.
Distribution. Indian Ocean, Ceylon.
Laodice marama, Agassiz and Mayer, 1899, p. 162, pl. ii.
figs. 7-8.
This species closely resembles Laodice indica, but can be
distinguished from it by the presence of usually two or three
cordyli between every two tentacles. Curri present. The
tentacles have long tapering basal bulbs and are without a
spur. ‘The size of the umbrella and the general appearance
of the gonads suggests the description having been based
upon a young immature stage.
The presence of cirri distinguishes this species from
LL. pulchra.
Distribution. Pacific Ocean, Fiji.
Laodice pulchra, Browne, 1902, p. 280.
In this species there are generally three to four cordyli
between every two tentacles, and they are situated upon small
bulbs. Cirri absent. The tentacles are without a basal spur.
Gonads arranged in a series of short folds along both sides of
very large radial canals.
Distribution. South Atlantic, Falkland Islands.
Laodice Maasti, nov. nom.
Laodice fijiana, vay. indica, Maas, 1905, p. 25, Taf. 11. figs. 14-15, Tat. v.
figs. 32-35.
It was not without some hesitation that I decided upon
giving a new name to the Laodice described by Maas in the
belonging to the Family Laodiceide. 467
report on the ‘Siboga’ medusz. Maas believes his specimens
to be either identical with, or closely related to, Laodice
Jyjiana of Agassiz and Mayer. If not identical, he suggests
that they should be regarded as a variety under the name of
indica. In describing the specimens Maas took the oppor-
tunity to criticize the genus Laodice and its allies. His is an
excellent criticism, and after I had independently investi-
gated the literature on the species I was pleased to find myself
in agreement with him.
Laodice Maasii is twice to three times the size of L. fijiana,
with more than twice the number of tentacles, and with many
more cordyli. The gonads extend much further along the
radial canals. Both species are provided with ocelli on about
two thirds of the basal bulbs of the tentacles. Taking the
above characters alone there is no reason for not imagining
the smaller LZ. jyzana growing to a larger size and possessing
more tentacles, more cordyli, and longer gonads. Then it
would resemble L. Maasii. ‘lhe medusa figured by Agassiz
and Mayer does not look at all like Maas’s medusa. ‘The
gonads of L. fijiana are adjacent to the stomach and on
conspicuous diverticula of the radial canals, whereas in Maas’s
medusa the diverticula are not visible in the figures, though
the author states that there are outgrowths along the radial
canals. It is a question of degree between a slight outgrowth
and a conspicuous one, ‘There is, however, one character by
which the two species can be distinguished. Maas figures
and describes the tentacles with basal spurs, which are not
present in L. fyiana.
Distribution. Kast Indian Archipelago.
Laodice fjiana, Agassiz and Mayer, 1899, p. 163, pl. iii.
figs. 9-10.
This species has a very few cordyli; only about eight are
present, though the tentacles number about seventy. Cirri
absent. ‘l’entacles without a basal spur. Vhe gonads are
upon short lateral diverticula of the radial canals. The
scarcity of cordyli and the presence of conspicuous diverticula
on the radial canals carrying the gonads appear to be the
principal characters of this species.
Distribution. Pacific Ocean, Fiji.
The following species are excluded from the genus
Laodice :—
Laodice cruciata, Heeckel, 1879.
I think it would be a distinct advantage if this specific
468 Mr. E. T. Browne on the Meduse
name were regarded as obsolete. In the first place, it is
impossible to identify the original Medusa cruciata of Forskal,
as its description and figures are too indefinite. In the second
place, Heckel has produced great confusion by putting under
the name of Laodice cruciata several species which clearly
belong to other genera. I have criticized in detail Heckel’s
synonyms in the Proc. Zool. Soc. (1896, p. 482), and it is
not necessary to do so again. There is only one genuine
Laodice amongst the lot, namely Thaumantias mediterranea,
Gegenbaur.
Cosmetira salinarum, du Plessis, 1879, p. 39, pl. xi.
Laodice salinarum, Heeckel, 1880, p. 636.
This species was found by du Plessis in brackish-water
ditches in a salt-marsh near Cette. Du Plessis says that “it
is curious that it is a miniature copy of a much larger species,
Cosmetira punctata, which occurs in the sea near Cette.”
Cosmetira punctata is a synonym of Laodice mediterranea.
The description given by du Plessis is rather vague, and the
photograph, which is the only figure, is too fuzzy to show any
details. From the description | rather think that the medusa
is more likely to be an Olindias or one of the Olindiade. It
was found suspended by the long tentacles from the lower
surfaces of masses of alge. ‘This points to the tentacles
having adhesive disks. ‘Lhe tentacles are provided with rings
of nematocysts, and between the tentacles at regular intervals
are some little reddish sacs, which have a pigment-spot and
some crystalline concretions. The sensory clubs of the
Laodiceide are without otoliths or crystalline concretions.
There is no clear evidence that this medusa belongs to the
Laodiceide, and it should be searched for again and properly
described. Maas (1905) has also expressed an opinion to the
same effect.
Laodice cellularia, A. Agassiz, 1862, p.350; id. 1865, p. 127,
figs. 195-196.
Thaumantias cellularia, Heckel, 1879, p. 129; Murbach and Shearer,
1803, p. 172, pl. xvii. fig. 2.
Agassiz, in his original description of this species, was
doubtful whether it belonged to the genus Laodice, for the
examination of the tentacles could not be made sufficiently
accurate to determine this point. Murbach and Shearer have
again found this medusa. 'I'hey definitely state that specimens
preserved in formalin do not show ocelli or cirri. As nothing
belonging to the Family Laodiceide. 469
is said about sensory clubs, it may be presumed that these
organs are also absent, and therefore the medusa is not a
Laodice. Whether it is a Thaumantias or not depends upon
the result of a revision of the Thaumantide and Hucopidee.
Thaumantias cellularia inhabits Puget Sound and the straits
between Vancouver Island and British Columbia.
Laodice Chapmani, Giinther, 1903, p. 425, pl. ix. figs. 1-3.
Maas has already expressed an opinion that this species is
not a Laodice. It certainly does not look like one, and the
absence of cordyli excludes it from the Laodiceide. The
description is based upon a single specimen found in the
North Atlantic.
Laodice neptuna, Mayer, 1900, p. 48, pl. xx. figs. 50-52.
This medusa was found at the Tortugas, off the coast of
Florida. It has been well described and figured by Mayer,
who does not mention the existence of cordyli; consequently
I exclude it from the Laodiceidee. In general appearance
this medusa does not look like a Laodice, but more like a
medusa belonging to another family at an intermediate stage
in development.
Genus Srauropuora, Brandt, 1835.
Staurophora, Heckel, 1879.
Staurostoma, Heckel, 1879.
Generic character.— Laodiceide with four radial canals ;
with a narrow cross-shaped stomach and mouth extending
across the subumbrella; with ocelli on the basal bulbs of the
tentacles.
Although Brandt established the genus Staurophora, it was
Louis Agassiz who, in his description of Stawrophora laciniata,
first gave an accurate account of a Staurophora, and clearly
demonstrated the existence of a mouth and stomach. One
of his figures shows distinctly a cordylus, though no mention
is made of this organ in the description. Agassiz was per-
fectly right in associating his species with Brandt’s genus
Staurophora,.
Heckel has certainly misinterpreted Brandt’s figures of
Staurophora in considering the lobes of the stomach to be
blind lateral branches of the radial canals ; hence his placing
Staurophora in the family Cannotidee. 'Lhis error led to his
introducing a new genus, Staurostoma, for Agassiz’s species,
which was placed amongst the Thaumantide. Hartlaub and
470 Mr. E. T. Browne on the Meduse
Maas have also expressed their disapproval of Heckel’s
Staurostoma.
The most interesting character of Staurophora is the posi-
tion of the stomach, mouth, and the gonads. How they
obtained their present position will be more readily understood
after considering the position of these organs in Laodice. It
appears to me that Stawrophora is descended from a Laodice-
like medusa.
In Laodice pulchra the radial canals are extremely large
and the gonads are situated upon them. The gonads are
arranged in a series of short folds forming a row on each
side of the canals, close to the subumbrella. They extend
along the whole length of the enlarged canals right up to
the central stomach, where they very nearly meet the gonads
belonging to the adjacent canals. In my original description
of Laodice pulchra the enlarged portions of the radial canals
were regarded as lobes of the stomach, and not as radial
canals. I considered the very short canals between the lobes
and the circular canal to be the true radial canals. This
species certainly has the appearance of possessing a very
large four-rayed stomach with gonads extending along the
lobes and a large central mouth with the margin in folds.
If one were to slit open along the middle the enlarged
portions of each radial canal of Laodice pulchra, and imagine
the cut margins to be the margins of a mouth, then the
position of the mouth, stomach, and gonads would be similar
to those of Staurophora.
I think the mouth of Stauwrophora has arisen by the out-
growth of a central mouth along the enlarged portions of the
radial canals of a Laodice-like medusa, and consequently
those portions of the radial canals have been converted into
a four-rayed stomach. The gonads have not changed their
position, but in Stawrophora they have lengthened slightly
and meet in the centre of the cross.
The earliest stages of Staurophora laciniata are very similar
to those of a young Laodice. hey have a small central
stomach and mouth and four radial canals. A. Agassiz has
traced the development of the mouth of S. lactndata, and his
figures clearly show how the mouth grows out to form a
perradial cross.
There is no disputing the fact that in Staurophora the
gonads are upon the walls of the stomach and occupy the
position of the gonads of an Anthomedusa. There is, how-
ever, very good evidence that Laodice undulata comes from
a calyptoblastic hydroid belonging to the genus Cuspidella,
and there are also Laodiceide with gonads on the radial
belonging to the Family Laodiceide. 471
canals far away from the stomach; so that the position of the
gonads of Staurophora does not justify its removal to the
Anthomeduse nor allow one to look upon the enlarged radial
canals of Laodice as lobes of the stomach. Hartlaub (1897)
has suggested that Staurophora is related to the Tiaride,
because the early stages bear a resemblance to Tvara.
Staurophora Mertensii, Brandt, 1835; id. 1838, p. 400,
Taf. xxiv.—xxv.; Heckel, 1879, p. 149.
This is the type species of the genus, and as it has not
been taken since the days of Brandt, a fresh description to
meet modern requirements and detailed drawings are much
needed.
Brandt’s figures show that the cross-shaped stomach an]
mouth, which extend right across the subumbrella, have a
large number of short lateral lobes. These lobes form the
characteristic feature of the species, as they are arranged ina
definite manner, either alternating or in pairs, and have a
definite shape.
Distribution. North Pacific; Norfolk Sound and off the
Aleutian Islands.
Staurophora arctica (Heckel), 1879.
Staurostoma arctica, Heckel, 1879, p. 131; Levinsen, 1892, p. 145;
Auriyillius, 1896, p. 194; Linko, 1900, p. 4, Taf. ii. figs. 22-25;
1904, p. 218; 1907, p. 151.
This species, according to Heeckel, has the gastro-genital
cross extending completely across the subumbrella, but the
mouth extends for only half that distance, so that the distal
half of each ray is a closed tube.
Linko (1900) states that all the tentacles (over 400 in
number) are equal in size and similar, and that on their inner
side, close to the velum, there is blackish ocellus. He figures
a long cordylus between every two tentacles and also a sensory
vesicle above the velum, one opposite every tentacle. So far
as I know, a sensory vesicle has not been yet found in any
other species of the Laodiceidee, It is a small vesicle, witha
single otolith, embedded in the ectoderm, and situated just at
the juncture of the velum with the subumbrella.
Distribution. Arctic Ocean ; Spitzbergen (Heckel). Ba-
rents Sea, Kolafjord and Ekaterinen Haven in Lapland
(Linko). West coast of Greenland (Levinsen).
472 Mr. E. T. Browne on the Meduscee
Staurophora laciniata, L. Agassiz, 1849.
Staurophora laciniata, L. Agassiz, 1849, p. 300, pl. vii.; A. Agassiz,
1865, p. 136, figs. 215-219; Wagner, 1885, p. 80, Taf. iv.; Fewkes,
1886, p. 958; Fewkes, 1888, p. 233; Hartlaub, 1897, p. 484, Taf. xvi.c,
Taf. xxii.; Broch, 1905, p. 7.
Staurostoma laciniata, Heeckel, 1879, p. 180; Hargitt, 1905, p. 43.
Staurophora Ketthii, Peach, 1867, p. 358, pl. i.
Thaumantias melanops, M‘Intosh, 1890, p. 40, pl. viii.; Hartlaub, 1904,
p. 103.
This species has the mouth extending along the whole
length of the stomach, and the tentacles form alternating
series of large and small ones, but the difference in size is
very slight. Both this species and S. arctica require further
examination, and careful drawings should be made of the
organs on the margin of the umbrella, especially of the
tentacles. It is not yet definitely proved that the latter is a
distinct species.
Peach states that his specimens agreed in every detail
with L. Agassiz’s description of S. laciniata, except that the
four rays of the stomach meet to form a perfect cross, whereas
Agassiz figures an imperfect cross. This slight difference is
not a specific character, though Peach attached great import-
ance to it.
M‘Intosh described under the name of Thaumantias
melanops an abnormal Hydromedusa without stomach or
mouth. Hartlaub (1904) has examined M‘Intosh’s specimen,
and states that it is a typical S. daciniata.
Hartlaub (1897) records the capture of a large specimen
at Heligoland, but he is a little uncertain whether it belongs
to S. arctica or S. laciniata. He also obtained some very
early stages, about 2 mm. in diameter, and kept them alive
for several weeks in an aquarium. ‘They fed on copepods
and grew at a great rate. It was whilst watching the
development of these young stages that Hartlaub saw cordyli
develop direct into tentacles. The figures of these young
stages do not show cordyli, but only tentacular buds.
Distribution. Arctic Ocean ; White Sea (Wagner). North
Atlantic; America, Boston Harbour (L. Agassiz); Nahant
(A. Agassiz) ; Woods Holl (Hargitt). Liat. 38° N., long. 68°
W.(Lewkes). Bay of Fundy ; Grand Manan Is. and Frye’s Is.
(Fewkes, 1888). Europe; Norway (Broch). Scotland, east
coast (Peach, M*Intosh). Heligoland (Haurtlaub).
Staurophora falklandica, sp. 0.
This new species was taken by the Scottish Antarctic
belonging to the Family Laodiceide. 473
Expedition (8.8. ‘Scotia’) in Stanley Harbour, Falkland
Islands. A description of it, with figures, will be given in
the Report on the Meduse collected by the expedition, to be
published in the ‘Transactions of the Royal Society of
Edinburgh.’
It is very much like S. daciniata, but has a series of long
tentacles and a series of very short rudimentary tentacles.
Genus PrycHoaena, A. Agassiz, 1865.
Generic character.—Laodiceide with four radial canals ;
with a central stomach and mouth; with the basal bulbs of
the tentacles without ocelli.
Ptychogena lactea, A. Agassiz, 1865.
Ptychogena lactea, A. Agassiz, 1865, p. 157, figs. 220-224; Heeckel,
1879, p. 147; Hargitt, 1905, p. 45.
Ptychogena pinnulata, Heckel, 1879, p. 148; Heeckel, 1882, p. 7, pl. ii. ;
Gronberg, 1898, p. 465; Levinsen, 1892, p. 145.
Ptychogena pinnulata, var. intermedia, Linko, 1904, p. 217.
This species was first discovered by A. Agassiz, who found
it abundant for a few days in Massachusetts Bay, about 1864,
and it has not again been recorded for the North-American
coast. This single record rather indicates that this medusa is
not a native of that region, but has probably drifted down
south from the Arctic regions.
According to Heckel P. pinnulata differs from P. lactea in
the shape of the gonads. ‘Those of the latter have fewer
lateral diverticula, but some of the longer ones are slightly
branched.
Linko has found in Barents Sea a form which he considers
to be a variety intermediate between P. lactea and P. pinnu-
lata. ‘This variety has gonads shaped like P. lactea, but with
about as many diverticula as P. pinnulata. It agrees in
colour with P. dactea. I do not think that the differences in
the shape and size of the gonads are, taken by themselves,
sufficient for a specific character, and Linko’s variety shows
a connexion between the two species. The differences in shape
and size are more likely due to the development of the gonads.
This species is probably a scarce Arctic medusa which
drifts south into the North Atlantic. There is no trustworthy
evidence that it is a deep-sea form.
Distribution. Arctic Ocean: Greenland (Grénberg). Barents
Sea (Linke). North Atlantic: America, Massachusetts Bay
and Nahant (A. Agassiz). Iceland (Levinsen). Between
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 31
474 Mr. E. T. Browne on the Meduse
Treland and Iceland, lat. 59° 7’ N., long. 13° 32! W., lat.
42° 8’ N., long. 63° 39’ W. (Heckel).
Ptychogena antaretica, sp. n.
This new species was taken off Cape Adare, Victoria Land,
by the ‘Southern Cross’ Expedition. There is only one
specimen, which unfortunately has a large hole through the
top of the umbrella. ‘lhe stomach and mouth are completely
gone and only the distal halves of the four gonads remain.
The margin of the umbrella is in perfect condition, The
basal bulbs of the tentacles are laterally compressed, and
there are no ocelli. They belong to the same type of bulb as
that figured by Agassiz and Heckel for P. dactea, and are not
like the basal bulbs of Stawrophora or Laodice.
The gonads are large, with broad lateral folds. In the
region of the gonads the radial canals show a wavy margin
corresponding to the principal folds of the gonads, but
the canals have no lateral diverticula like P. lactea. The
shape of the gonads is intermediate between P. lactea and
P. longigona.
The new species can easily be distinguished from P. lactea
by the absence of diverticula on the radial canals and by the
colour of the tentacles, which are red. It is not so easy to
distinguish it from P. longigona, because the organs on the
margin of the umbrella of P. longigona have not been
described in detail or figured. The gonads of the P. ant-
arctica have much broader lateral folds and do not extend so
far along the radial canals.
A description with figures of P, antarctica will appear in
the Report on the Meduse collected by the ‘ Discovery’ and
‘ Southern Cross’ Expeditions.
Ptychogena longigona, Maas, 1893, p. 64, Taf. vi. figs. 7-9.
Maas, in the description of this species, states that it has
* Randkolben,” by which J presume he means cordyli, As
he has omitted to figure the margin of the umbrella, it is
necessary to rely upon the brief description. The gonads are
very long, extending the whole length of the radial canals,
and are arranged in a series of lateral folds or lobes, but the
radial canals have no lateral diverticula as in P. lactea.
Distribution. North Atlantic, off the north-west coast of
Scotland,
belonging to the Family Laodiceide. 475
Genus Srauropiscus, Heckel, 1879.
Generic character.—Laodiceida with four main radial
canals, each with lateral branches.
The two genera Staurodiscus and Voworchis are distin-
guished from the other genera of the Laodiceide by the
presence of branched radial canals. It was on account of the
branching of the canals that Heckel placed these genera in
the Cannotide, The Cannotide, as a distinct family, has
now ceased to exist; its destruction was due to a revision of
its genera by Maas (1904). Staurodiscus and Toxorchis
were transferred by Maas to the Berenicide, to which he
gave an emended definition.
Berenice is the type genus of the Berenicidee. One species
(B. rosea) is without marginal bulbs, but the other (B. Hux-
feyz) has bulbs. I am uncertain whether these bulbs are
cordyli or only tentacular bulbs ; if they should turn out to
be cordyli, then the species should be transferred to the genus
Staurodiscus.
Staurodiscus tetrastaurus, Heckel, 1879.
Staurodiscus tetrastaurus, Heckel, 1878, p. 145, Taf. ix. figs. 1-3;
Mayer, 1900, p. 46, pls. xviii—xix. figs. 47-49; Maas, 1904, p. 440.
Staurodiscus heterosceles, Heckel, 1879, p. 146.
In this species each of the radial canals has a pair of lateral
branches which do not join the circular canal. The gonads
develop upon the blind branches and also upon the portion of
the main canal between the branches and the circular canal.
There are eight to sixteen tentacles and two or three cordyli
between every two tentacles. Cirri absent. A black ocellus
at the base of all the tentacles and cordyli. Mayer describes
young stages as well as adult, and states that the ocelli are
endodermal.
Distribution. North Atlantic: Canary Is. (Heckel).
Tortugas (Mayer).
Staurodiscus nigricans, Agassiz and Mayer, 1899, p. 164,
pl. iv. figs. 11-12.
This species has radial canals with a pair of lateral branches,
which do join the circular canal. The gonads are upon the
branches and the portion of the main canal between the
branches and the circular canal. ‘Twelve tentacles present
and six or seven cordyli between every two tentacles. Curri
and ocelli absent.
Distribution. Pacific Ocean; Fiji (Agassiz and Mayer).
aL
476 Mr. E. T. Browne on the Medusce
Genus Toxorcuis, Heckel, 1879.
Generic character.—Laodiceide with six main radial canals,
each one widely forked or with lateral branches.
Toxorchis arcuatus, Heckel, 1879, p. 157, Taf. ix. figs. 6-8.
This species has only been seen by Heckel. His figures
show that each radial canal is widely forked, with gonads in
the fork of the canals. The margin of the umbrella is
provided with twenty-four tentacles, and between every two
tentacles there are a cirrus and a cordylus. The basal bulbs
of tentacles have ocelli.
Distribution. North Atlantic; Canary ls. (Teckel).
It is probable that the genus Cladocanna, Heeckel (1879,
p- 160), will ultimately become a synonym of Yowxorchis.
There are two species, C. thalassina (Péron, 1809), which
has not been well described, and C. polycladia, which
Heckel has described but not figured. The latter species
has six radial canals with several lateral branches, each of
which is again dichotomously divided. The tentacles are
very numerous and between them are cirri and marginal clubs.
If the marginal clubs turn out to be cordyli, then I would
suggest that the species be placed in the genus Toworchis.
Maas (1904) considers C. polycladia to be identical with
C. thalassina, and doubtfully refers it to the genus Toworchis.
Genus MELIcERTISSA, Heckel, 1879.
Generic character.—Laodiceide with eight radial canals,
without lateral branches.
This genus Heeckel placed in the Thaumantide, in the
subfamily Melicertide, containing genera with eight canals.
This subfamily will probably disappear on the completion of
the revision of the Thaumantide.
Melicertissa clavigera, Heckel, 1879, p. 135, Taf. vii.
figs. 8-12.
This is another species which has only been seen by
Heckel. It has only eight tentacles, and between every two
tentacles there are three cordyli but no cirri. The basal
bulbs of the tentacles and cordyli have ocelli.
Distribution. North Atlantic : Canary Is. (ecke?).
belonging to the Family Laodiceide. 477
Melicertissa malayica (Maas), 1905.
Melicertidium malayicum, Maas, 1905, p. 28, Taf. v. figs. 29-31.
This species is readily recognized by the large number of
tentacles (about one hundred and fifty). There is roughly
one cordylus to every two tentacles, but not between every
pair of tentacles; the total number of cordyli is about half
that of the tentacles. The cirri are rather scarce. About
one fourth of the tentacles are provided with ocelli. The
gonads are on the proximal half of the radial canals.
Distribution. Kast Indian Archipelage (Maas).
Maas placed this species in the genus Melicertidium as it
agreed with Heeckel’s definition of the genus, which happened
to be an erroneous one. I have recently emended the genus
Melicertidium. ‘The species belonging to it have eight radial
canals and numerous tentacles, but they are without cirri or
marginal bulbs of any kind.
The following genera and species have for the present been
excluded from the Laodiceidee :—
Octonema eucope, Heckel, 1879, p. 127.
The genus Octonema was established by Heckel for a
single species found at Honolulu, Sandwich Is. According
to Heckel’s classification, the genus is distinguished from
Laodice by the presence of only eight tentacles. The Species
has a large number of marginal bulbs, knobs, and cirri.
Unfortunately there is no figure of this medusa, so that the
exact meaning of ‘f Randkolben” remains doubttul. Heckel
also states that a black ocellus is situated on the outer side of
the basal bulb of the tentacles. ‘The Laodiceide usually
have the ocelli on the inner side of the basal bulbs, and there
is no trustworthy evidence to show that any species of the
family has ocelli on the outer side. It is quite probable that
Octonema eucope belongs to another family.
Octonema gelatinosa, Mayer, 1900, p. 8, pl. vi. figs. 20-21.
The description of this species is based upon a single
specimen taken in Charleston Harbour, U.S.A. It has the
appearance of a young stage, with only four tentacles and
twenty marginal bulbs. Hach bulb has an ocellus, which,
according to the description, is situated in the endoderm.
There are eight marginal clubs, and a figure shows that their
478 Mr. E. T. Browne on the Medusee
distal end is provided with cells which look like nematocysts.
Cordyli proper are without nematocysts, so that these bulbs
are more likely to be tentacular bulbs.
Octorhopalon fertilis, von Lendenfeld, 1884, pp. 919, pl. xlii.
figs. 14-15.
This is a little medusa, 2 mm. in diameter, having eight
tentacles and eight marginal clubs, and was found by Len-
denfeld at Port Jackson, Australia. The description is rather
concise and the figures have been badly reproduced, so that
they do not help out the short description. ‘The author states
that cirri are absent, but nothing is said about ocelli, though
in the figure there are indications of an ocellus on the inner
side of the basal bulbs of the tentacles.
The gonads are very large, extending along the whole
length of the radial canals and also round the base of the
stomach. They are folded transversely.
Before one can classify this medusa among the Laodiceidee
there are two points which require further elucidation. Is it
a young stage with gonads just appearing or a fully grown
adult? Are the clubs true cordyli? ‘The clubs in one figure
have the appearance of cordyli lying across the velum ; in
the second figure they project outwards and have the
appearance of auditory clubs, which should contain an
otocyst.
List of Books and Memoirs quoted in the Text.
Agassiz, A. 1865. “ North-American Acalephe.”’ Catalogue Mus.
Comp. Zool. Harvard, No. ii.
Agassiz, A., and Mayer, A.G. 1889. “Acalephs from the Fiji
Islands.” Bull. Mus. Comp. Zool. Harvard, vol. xxxii. pp. 157-
189, 17 pls.
Aaassiz, L. 1849. ‘ Acalephe of North America.” Mem. Amer.
Acad, Arts & Sci. vol. iv. pp. 221-316, 8 pls.
—. 1860-1862. ‘Contributions to the Natural History of the
United States of America, yol.iv. Boston.
Auvriviuuius, C. W.8. 1896. “Das Plankton der Baffins Bay und
Davis Strait.” Festskriff for Lilljeborg, pp. 181-212. Upsala.
Branpt, J. F. 1838. “ Ausfiihrliche Beschreibung der von C. H.
Mertens auf seiner Weltumsegelung beobachteten Schirmquallen.”’
Mém. Acad. Imp. St. Pétersb. sér. 6, tom. ii. (Sci. nat.) pp. 237-
411, 31 pls.
Brocu, H. 1905. ‘Zur Medusenfauna von Norwegen.” Bergens
Mus. Aarbog, No. xi. 8 pp.
Brooxrs, W.K. 1895. “ The Sensory Clubs or Cordyli of Lacdice.”
Journ. Morphology, vol. x. pp. 287-804, pl. XVii.
Browne, E.T. 1898. ‘On British Meduse.” Proc. Zool. Soc. 1897,
pp. 816-836, pls, xlviii.-xlix.
belonging to the Family Laodiceide. 479
Browne, E. T. 1902. “A Preliminary Report on Hydromedusee
from the Falkland Islands.” Ann. & Mag. Nat. Hist. (7) vol. ix.
pp. 272-284.
—. 1905. ‘“Meduse,” in Herdman’s ‘ Pearl Fisheries, Ceylon,’
part iv. pp. 181-166, 4 pls. Royal Soc. London.
Frewxes, J, W. 1886. “ Report on the Meduse collected by the
‘ Albatross’ in the Region of the Gulf Stream in 1883-1884, Ann.
Rep. Comm. Fisheries tor 1884, pp. 927-977, 10 pls. Washington.
——. 1888. “On certain Meduse from New England.” Bull. Mus.
Comp. Zool. vol. xiii. pp. 209-240, 6 pls.
Forsrs, E., and Goopstr, J. 1851. “On some remarkable Marine
Invertebrates new to British Seas.” ‘Trans, Roy. Soc. Edinb.
vol. xx. pp. 307-315, pls. x.—xi.
GEGENBAUR, C. 1856. “Versuch eines Systemes der Medusen.”
Zeitschr. wiss. Zool. Bd. viii. pp. 202-273, Taf. vii.—x.
GronBere, G. 1898. “Die Hydroid-Medusen des Arktischen
Gebiets.” Zool. Jahrb, Abt. Syst. Bd. xi. pp. 451-468.
GintHerR, R. T. 1903, “ Report on the Celenterata from the
Intermediate Waters of the North Atlantic.” Ann. & Mag. Nat.
Hist. (7) vol. xi. pp. 420-480, pls. ix.—x.
Hacxer, EK. 1864. “ Beschreibung neuer craspedoter Medusen aus
den Golfe von Nizza.” Jena. Zeitschr. Naturw. Bd.i. pp. 825-842.
—. 1879-1880. ‘Das System der Medusen.’ Jena.
—. 1882. ‘Report on the Deep-sea Meduse dredged by H.M.S.
‘Challenger.’ ” Report 1V.— Die Tiefsee Medusen der Challenger-
Reise.’ Jena, 188].
Hareirr, C. W. 1905. “The Meduse of Woods Holl Region.”
Bull. Bureau Fisheries, 1904, vol. xxiv. pp. 21-79, 7 pls.
Hartiaus, C. 1897. “Die Hydromedusen Helgolands.” Wiss.
Meeres deutsch. Meere, Bd. ii. pp. 449-536, 10 pis.
1904. “ Bericht iiber eine zoologische Studienreise nach Frank-
reich, Gross-britannien und Norwegen, ausgefthit im Fruhjahre
1902.” Wiss. Meeresunter, Bd. v. Abt. Helgoland, pp. 97-106.
Hincxs, T. 1868. ‘A History of the British Hydroid Zoophytes.’
Ray Soc. London.
LENDENFELD, R. von. 1884. “ Addenda to the Australian Hydro-
medusee.” Proc. Linn. Soc. New South Wales, vol. ix. pp. 908-
924, pls. xl.-xliii.
LEVINSEN, G. M. R. 1892. “ Meduser, Ctenophorer og Hydroider
fra Gronlands.” Vidensk. Meddel. Nat. Foren. pp. 145-212,
tab, v.—vill.
Linxo, A. 1900. “Ueber den Bau der Augen bei den Hydro-
medusen.” Mém. Acad. Imp. Sci. St, Pétersb. sér. 8, tom. x.
pp. 1-28, Taf. 1-11.
——. 1904. “ Zoologische Studien im Barents-Meere.” Zool. Anz.
Bd. xxviii. pp. 210-220.
—. 1907. “Untersuchungen iiber das Plankton des Barents
Meeres.” Wiss. practische Murman-Expedition, 245 pp., 21 figs,
(Russian. )
Maas, O. 1893, ‘Die craspedoten Medusen. Ergebnisse der
Plankton-Expedition.’ ‘ National.” Hd. ii. K. c., 107 pp., 8 Tafn.
—. 1904. “ Bemerkungen zum System der Medusen. Revision
der Cannotiden, Heckel.” Sitz. math.-phys. Klasse kgl. bayer,
Akad. Wiss, Munchen, Bd. xxxiv. pp. 421-445,
—. 1904. “Méduses provenant des Campagnes des Yachts
(‘ Hirondelle’ et ‘ Princesse-Alice’), 1886-1903.” Résult. Camp.
Scient. Monaco, fasc. xxviii., 71 pp., 6 pls.
480 Mr. G. Lewis on
Maas, O. 1905. ‘Die craspedoten Medusen der Siboga-Expedition,’
Monograph x., 84 pp., 14 Tafn.
M‘Intosu, W.C. 1890. ‘Notes from St. Andrews Marine Labora-
tory: Abnormal Hydromeduse, &c.” Ann. & Mag. Nat. Hist.
(6) vol. v. pp. 40-48, pl. viii.
Mayen, A.G. 1900. “ Descriptions of new and little-known Medusee
from the Western Atlantic.” Bull. Mus. Comp. Zool. vol. xxxvii.
pp. 1-9, 6 pls.
—. 1900. “Some Meduse from the Tortugas, Florida.” Bull.
Comp. Mus. Zool. vol. xxxvii. pp. 11-82, 44 pls.
——. 1904. ‘“Meduse of the Bahamas.” Mem. Nat. Sci. Mus.
Brooklyn Instit. vol. i., 33 pp., 7 pls.
MeETscHNIkOFF, EH. 1886, ritnsbryaloeiaels Studien an Medusen.’
Wien.
Murpacu, L.,and SHEARER,C. 1903. ‘On Meduse from the Coast
of British Columbia and Alaska.” Proc. Zool. Soe. ii. pp. 164-
192, pls. xvli.—xxil.
Pracu, C. W. 1867. ‘On new British Naked-eyed Medusz.”
Journ. Roy. Instit. Cornwall, vol. ii. pp. 8355-860, 2 pls.
Puiessis, G. pu. 1879. “ Etude sur la Cosmetira salinarum, nouvelle
méduse paludicole des environs de Cette.” Bull. Soc. Vaud. sér. 2,
tom. xvi. pp. 39-45, pl. ii. Transl. in Ann. & Mag. Nat. Hist. (5)
vol. iii. pp. 885-389.
Waaner, R. 1885. ‘Die Wirbellosen des Weissen Meeres.’ Leipzig.
LXV.—On new Species of Histeridze and Notices of others.
By G. Lewis, F.L.S.
Tus paper, the thirty-third of the series, is a short appendix
to the last (Ann. & Mag. Nat. Hist. xx. p. 839, 1907), and
furnishes at the close of the year sundry descriptions which
render the preceding papers, as a whole, more complete.
List of Species.
Teretrius placitus, Horn. Hister monitor.
Diplogrammicus, gen. nov. Carcinops lauta, Zemm.
intermedius. Echinodes decipiens, Horn.
Zabromcerphus rugicollis. Saprinus ovillum, Sols.
Teretrius placitus, Horn, Tr. Amer. Ent. Soe. viil. p. 143
(1880).
‘Black, shining, elytra red. Thorax nearly square, very
narrowly margined, surface moderately densely and equally
punctured. Elytra a little more coarsely but less densely
punctured than the thorax, and without trace of strie, colour
red. Propygidium and pygidium moderately densely punc-
tured, the latter more finely. Prosternum truncate in front.
new Species of Histeride. 481
Mesosternum with a fine entire marginal line, surface coarsely
and moderately densely punctured. Legs rufous. Anterior
tibie with four or five minute teeth, the middle with two
larger teeth ; posterior finely bispinose, the upper spine, which
is near the middle, very small.
“ L. °08 inch ; 2 mm.
‘“‘ A more robust species than americanus and less coarsely
punctured, and differing especially in the dentation of the
tibia. ‘he colour-character is of secondary importance, and
while the three specimens before me are alike, it is possible
that others will occur entirely black.
“ Collected by Mr. H. K. Morrison in Western Nevada.”’
All the specimens I have seen have red elytra.
DIPLOGRAMMICUS, gen. nov.
Body oblong, somewhat convex, punctulate above; elytra
with three dorsal strize, third sometimes broken, depressed in
the sutural area; propygidium convex, not impressed nor
uneven; prosternum bistriate; mesosternum marginate
anteriorly. The other characters correspond with those of
Omatlodes, of which genus omega, Kirb., is the type.
The species to be included in Diplogrammicus are ebeninus
(Omalodes), Er., type, Marseuli, Sch., and tntermedius, Lew.
Diplogrammicus intermedius, sp. n.
Oblongus, parum parallelus ; fronte impressa stria retrorsum angu-
lata; pronoto lateribus et anterioribus dense punctato ; elytris
striis 1-3 dorsalibus integris; propygidio pygidioque dense
punctatis ; prosterno in medio bistriato.
L. 9 mill.
Oblong, somewhat parallel laterally, black and shining ;
the head impressed longitudinally, stria complete, turning
backwards in the middle, surface rather densely punctured ;
the thorax densely punctured laterally, less distinctly behind
the head, and the points gradually lessen to the disk, scutellar
region smooth, the lateral stria is not well-marked owing to
the encroachment of the punctuation and it ceases at the
posterior angle. The lateral strie of Marseuli, Sch., is well-
marked and turns inward at the base and passes the angle.
The elytra are finely not closely punctulate (more distinctly
so than in Marseulr), the striz 1-3 are complete, but the
third is somewhat irregular in its course, the others are
wanting ; the propygidium is coarsely punctured and some-
what gibbous along the sides; the pygidium is similarly
482 Mr, G. Lewis on
punctured ; the prosternum is bistriate before the coxe ; the
mesosternum (not quite so shortened as that of Marseulv) is
punctulate, with marginal stria complete.
The description above is drawn from a specimen from the
Chevrolat collection which Marseul named ebeninus, Er., but
it is clearly not so. Marseul gives an excellent figure of
ebeninus in his monograph (pl. xv. fig. 22), showing the
dorsal striz and punctuation very clearly, and the sternal stria
also are separately shown. Lrichson’s species is a native of
Argentina.
Hab. Chili (ex coll. Chevrolat).
Zabromorphus rugicollis.
Oblongo-ovalis, convexus, niger, nitidus; fronte rugcso-punctata,
stria carinata, antice recta ; pronoto striis 2 lateralibus integris,
utrinque grosse et dense strigoso-punctato; elytris striis sub-
humerali 1-6 dorsalibus omnibus integris, 57 cum suturali con-
nexa; propygidio pygidioque dense et grosse punctatis; tibiis
anticis 3-dentatis.
L. 83 mill.
Oblong-oval, convex, black and shining ; the head coarsely
and rugosely punctured, frontal stria carinate and straight
anteriorly, mandibles concave, also rugose; the thorax,
marginal stria fine and complete, inner and outer lateral strize
also complete, the former continuing behind the head, surface
behind the middle of the neck punctured not very closely but
laterally and broadly behind the angles, the surface is strongly
and longitudinally strigose, the strigosities continuing but
gradually narrowing to the base, behind the middle of the
disk and along the base there is an irregular space almost
smooth ; the elytra, strie deeply impressed, subhumeral and
dorsal all complete, the fifth and sutural joining at the base ;
the propygidium and pygidium are densely and coarsely pune-
tate; the prosternum, keel narrow, anterior lobe bistriate ;
mesosternum feebly and widely emarginate, stria complete ;
the anterior tibiz are 3-dentate.
This species differs from pachysoma, Ancey, and longicollis,
Mars., by the conspicuous strigose sculpture of the thorax,
and from the first by having only two lateral thoracic striz.
I have seen an example of rugicollis in which the sutural and
fifth are not joined, and I have an example of pachysoma in
which these striz almost join. The dorsal striz resemble
those of apis, Mars., a species from the Cape which I have
not seen.
Hab. Benguella.
new Spectes of Histeride. 483
Hister monitor, sp. n.
Breviter ovalis, convexiusculus, niger, nitidus ; fronte stria integra ;
pronoto stria interna integra, basi haud incurvata, externa ante
medium abbreviata; elytris striis 1-2 integris, 3-5 obsoletis,
suturali vix abbreviata ; propygidio pygidioque minute punctatis,
hoe anguste marginato ; mesosterno stria arcuata integra ; tibiis
anticis 3-dentatis.
L. 8-103 mill.
Shortly oval, rather convex, black and shining ; the head
flat and impunctate above, stria well-marked, complete, and
nearly straight anteriorly; the thorax, inner lateral stria
strong and complete, not incurved at the base, interstice rather
wide, outer stria short, not passing the angle anteriorly, but
extending a little behind ; the elytra, striz, inner humeral
apical and just passing the middle, 1-2 dorsal complete, but
the second is somewhat feeble at the base, 3-5 are traceable
only in certain lights and are indicated by fine points, the
sutural is fine but distinct, abbreviated anteriorly but only
indistinctly shortened behind; the propygidium and the
pygidium are very minutely and not closely punctured, the
latter is narrowly margined on the outer edge; the meso-
sternum is emarginate, with a stria on both sides of the
emargination, the usual marginal stria is complete and well-
marked and is arched in front and does not follow the contour
of the segment. HH. gorilla, Sch., has the mesosternal stria
sinuous anteriorly and no short lateral striae. HH. adjectus,
Mars., has the lateral stricze, but they are not so conspicuous.
The anterior tibiz are strongly 3-dentate.
The dorsal striz of this species are very peculiar, but the
three species mentioned have some characters in common.
Hab. British Uganda (Grauer). In the Deutsches Ento-
mologisches National Museum and my own collection. Three
examples.
Also found by Messrs. Legge and Wollaston at Old Camp,
KE. Ruwenzori, 6000-7000 feet, 1906.
Carcinops lauta, Zimm. Tr. Amer. Ent. Soc. p. 253 (1869).
“‘ Oval, convex, very shining, piceous ; antenne and legs
ferruginous ; head and thorax finely and sparsely punctured,
punctures of different sizes ; marginal stria of prothorax very
fine, scarcely obvious in front ; elytra with the sutural stria
feeble, composed only of separate punctures and abbreviated
in front, five dorsal strix entire, slightly curved inwards and
distinctly punctured ; the inner lateral stria is well developed
and finer than the dorsal strie; pygidium and propygidium
484. On new Species of Histeridee.
smooth; mesosternum deeply emarginate in front and
surrcunded by a fine stria; front tibise 2-toothed.
“LL. 2 line.
“ Hab. North Carolina.”
Horn, in his ‘Synopsis,’ 1873, p. 308, says the pygidia
are punctulate.
Echinodes decipiens, Horn, Tr. Amer. Ent. Soc. x. p. 286
(1883).
“Broadly oval, piceous, brown, moderately shining.
Head smooth, obtusely carinate at middle, deeply impressed
on each side. Thorax shining, surface with sparsely placed,
slightly muricate punctures, each bearing a short erect hair.
Elytra with a marginal and three vaguely impressed entire
strie at the side, each with a single row of coarse and closely
placed punctures bearing a short erect hair; between these
strize and the suture are four series of punctures rather irregu-
larly placed, bearing setze, the sutural row the most distinct.
Propygidium and pygidium shining, sparsely punctate. Body
beneath shining, smooth, Prosternum bicarinate, the carine
divergent posteriorly. LL. *06 inch; 1°5 millim.
“The species resembles KH. setiger, Lec., and is but a little
longer. It differs in the elytral sculpture. The present
species has the outer striae more impressed, the inner rows of
punctures single, while in setger the outer striz are less
impressed and the inner rows composed of two or more series
of punctures.
“ Collected by H. K. Morrison in Arizona.”
Saprinus ovillum, Sols. Reis. Fedsch. Turkestan, ii. p. 240
(1876).
“ Oblongo-ovatus, convexus, piceo-niger, supra nigro-zneus, nitidus,
antennis pedibusque piceis. Fronte crebre punctata, antice obso-
lete transversim carinata, stria nulla. Pronoto transverso, basi
leviter rotundato, vix bisinuato, lateribus rotundato, antrorsum
angustato, convexo, disco basin versus subtiliter et minus crebre,
ceterum crebre, minus subtiliter punctato; stria marginali integra.
Elytris apice ad suturam subtiliter disperse punctatis, stria
suturali integra, antice cum quarta dorsali arcuatim conjuncta,
dorsalibus paulo pone medium abbreviatis, 1 et 2 subbrevioribus,
subhumerali interna cum humerali conjuncta, postice abbreviata,
externa nulla. Pygidio crebre punctato. Prosterno striis
parallelis, apice confluentibus. ‘Tibiis anticis 5-dentatis, dentibus
tribus ultimis fortibus.
“L. 26 mill”
Hab. “In Valle Sarafschan.”’
On Three new Freshwater Fishes. 485
LXVI.— Descriptions of Three new Freshwater Fishes disco-
vered by Mr. G. L. Bates in South Cameroon. By G. A.
BouLENGER, F.R.S.
Nannocharax ocellicauda.
Depth of body 44 to 5 times in total length, length of head
4 times. Head twice as long as broad, deeper than broad ;
snout rounded, feebly projecting, a little shorter than eye,
which equals length of postorbital part of head and inter-
orbital width. Gill-rakers short and few. Dorsal III 9-10,
originating above root of ventrals, equally distant from eye
and from root of caudal; longest ray about ? length of head.
Anal IIL 9. Pectoral much shorter than head, not reaching
root of ventral. Lobes of caudal acutely pointes Caudal
peduncle once and 4 aslong as deep. Scales 50 © 3 O between
lateral line series and root of caudal ; lateral line incomplete,
reduced to a few tubercles. Yellowish in spirit, with a large
black, light-edged ocellus at the root of the caudal fin ;
vertical fins greyish, dorsal black in front.
Total length 35 mm.
Two specimens from the Ja River.
This fish brings up to nine the number of species of Nanno-
charaz. I have prepared the following synopsis for their
distinction :—
1. Dorsal originating above ventrals, with 9 to 11 branched rays.
A. Lateral line complete.
1. Less than 50 scales in lateral line, 43 in transverse series above
lateral line ; length of head 33 to 4 times in total length.
Head not deeper than broad; depth of
body 53 to 53 times in total length ;
pectoral as long as head, extending [ (Gold Coast, Gaboon, Congo).
beyond root of ventral ; Sq. 42-46 = 5 =, 1. N. fusciatus, Gthr., 1867
Head deeper than broad; depth of bods
43 to 5 times in total length ; pec-
toral as long as head, extending: Py
yond root of ventral ; Sq. 37-38 3 .. 2, N. brevis, Blgr., 1902 (Congo).
Head deeper than broad ; depth of oe
3% to 4; times in total length ; pec-
toral much shorter than head, not [(Ogowe).
reaching root of ventral; Sq. 388-40 *. 3. M. parvus, Pellegr., 1907
63
2. 47 scales or more in lateral line, 53 in transverse series above
lateral line ; length of head 4 to 5 times in total length,
Head deeper than broad ; depth of body
486 On Three new Freshwater Fishes.
5 to 53 times in total length; pec-
toral as long as head or slightly
shorter, reaching a little beyond (Cameroon),
origin of ventral ; Sq. 47-55 a .. 4, N. intermedius, Blgr., 1908
3!
Head not deeper than broad; depth of
body 5 to 6 times in total length ;
pectoral shorter than head; Sq. { (Nile).
BOEBD eitae tyre in aos cane 5, N. niloticus, Joann., 1835
Head not deeper than broad; depth of
body 63 to 73 times in total length ;
pectoral shorter than head; Sq,
52-57
13
[(Congo).
Hod ten adn stadia aac 6. N. elongatus, Blgr., 1900
B. Lateral line incomplete, reduced to a few tubercles; Sq. 50 Gk
ga
length of head 4 times in total length. ;
7. N. ocellicauda, Blgr., 1907
[ (Cameroon).
II. Dorsal originating well in advance of ventrals, with 11 or 12
branched rays ; head deeper than broad, 4 to 43 times in total
length; depth of body 44 to 5 times in total length; pectoral
shorter than head.
Si 6 EE EE reatlics Ripeap et aac 8, N.tenia, Blgr., 1902 (Congo)
eBay) TA Bea ODM Ay bora AG AO Oo cet 9. N. dimidiatus, Pellegr., 1904
: [(Casamanza).
Barbus callipterus.
Depth of body 3 to 34 times in the total length, length of
head 34 to 32 times. Snout rounded, not longer than eye,
the diameter of which is 22 to 3 times in length of head and
equals interorbital width ; mouth terminal ; lips moderately
developed, interrupted on the chin; barbels two on each side,
subequal, 2 to # diameter of eye. Dorsal III 8, last simple
ray flexible, not enlarged, nearly as long as head ; free edge
of fin slightly concave ; its distance from occiput less than its
distance from caudal fin. Anal III 5, longest ray 3 to 2
length of head. Pectoral a little shorter than head, not
reaching ventral; latter below anterior half of dorsal. Caudal
peduncle about once and 4 as long as deep. Scales 25-26 eo
2 between lateral line and ventral, 8 or 10 round caudal
peduncle. Yellow, brownish on the back, the dorsal and
lateral scales dark brown at the base ; dorsal fin orange in
the basal half, white in the distal half, with a large deep
black spot between the last simple ray and the third branched
ray ; other fins white, the caudal orange at the base.
Total length 73 mm.
Four specimens from Akok, Kribi River.
On new Fishes from Central Africa. 487
Mastacembelus longicauda.
Depth of body 24 times in total length, length of head
(without rostral appendage) 11 times. Vent much nearer end
of snout than base of caudal fin. Length of head 4 times in
its distance from vent and a little greater than its distance
from first dorsal spine. Snout 4 times as long as eye, ending
in a trifid appendage, which is not quite twice as long as eye ;
cleft of mouth extending to below anterior border of eye; no
preeorbital spine; two strong preopercular spines. Vertical
fins united with the very short, broadly rounded caudal.
Dorsal XX VII 150; last spine once and } diameter of eye.
Anal IL 150*. Pectoral one fourth length of head. 14 series
of scales between origin of soft dorsal and lateral line. Brown
above, yellow beneath ; a dorsal series of small dark brown
spots ; sides with lighter spots, some of which are ocellar ;
end of tail with large dark brown spots.
Total length 305 mm.
A single specimen from Akok, Kribi River.
Allied to M/. Greshoffi, Blgr., from the Congo, but form
more elongate, caudal tin more rounded, and no preorbital
spine.
LXVII.— Descriptions of Three new Fishes from Central
Africa. By G. A. BOULENGER, F’.R.S.
Petersius Woosnamt.
Depth of body equal to length of head, 4 times in total
length. Head longer than deep, with convex upper profile ;
lower jaw projecting shghtly beyond upper; diameter of eye
equal to length of snout, 8 times in length of head; maxillary
not extending to below anterior border of eye; 14 teeth (°) in
the upper jaw, 8 in the lower; outer premaxillary teeth
tricuspid, inner multicuspid and inserted directly behind the
outer. Gill-rakers short, 12 on lower part of anterior arch.
Dorsal ILI 8, originating above base of ventrals and at equal
distance from end of snout and from root of caudal; longest
ray about $ length of head. Adipose fin very small. Anal
Ilf 17-18. Pectoral shorter than head, not reaching ventral.
* The numbers of soft rays are approximative, the vertical fins of the
unique specimen being rather stiff, rendering the counting of the rays
very difficult.
488 On new Fishes from Central Africa.
Caudal peduncle as longas deep. Scales 29-30 a) 2 between
lateral line and ventral. No markings, except a silvery
lateral band, which is black-edged above.
Total length 70 mm.
Two specimens from the Aruwimi, in the Congo Forest,
form part of the collection made by Mr. R. B. Woosnam and
presented to the British Museum by the Subscribers to the
Ruwenzori Expedition Fund.
In its dentition this new species stands nearest to
P. occidentalis, Gthr., in which the body is much shorter and
the lateral line incomplete.
Barbus Johnston.
Depth of body equal to ldhgth of head, 4 times in total
length. Snout rounded, a little longer than the eye, the
diameter of which is 44 times in length of head ; interorbital
width 24 times in length of head; mouth inferior, its width
3 times in length of head; lower jaw with a blunt edge,
covered by a thin horny sheath ; lips feebly developed, lower
confined to the sides; barbels two on each side, posterior
slightly longer than the anterior and as long as the eye.
Dorsal IV 9, last simple ray flexible, not enlarged, as long
as head ; free edge of fin deeply concave ; its distance from
centre of eye equals its distance from caudal fin. Anal III 5,
longest ray § length of head. Pectoral slightly shorter than
head, not reaching ventral; latter below middle of dorsal.
Caudal peduncle once and # as Jong as deep. Scales 38 4
3 between lateral line and ventral, 14 round caudal peduncle.
Silvery, darker on the back.
Total length 180 mm.
A single specimen from British Central Africa, between
Kondowe and Karonga, was presented by Sir Harry Johnston
in 1897.
B. rhodesianus, Blgr., which has a similar mouth (ap-
proaching the Capoéia type), is distinguished by its larger
scales (80-32 >) and its shorter barbels.
Amphilius Hargeri.
Depth of body 5} times in total length, length of head 32
times. Head slightly longer than broad; eyes small, in the
middle of the length of the head, 3 diameters apart; inter-
orbital width 2 length of snout; latter broadly rounded,
projecting but slightly beyond lower jaw; posterior nostril
On a new African Snake. 489
midway between eye and end of snout; premaxillary teeth
forming a short crescentic band; maxillary barbel once and
1 length of head; outer mandibular barbel # length of head,
inner $. Dorsal 16, well in front of vertical of ventrals,
but nearer latter than base of pectorals, equally distant
from end of snout and from middle of adipose fin ; first
branched ray 2 length of head. Adipose fin low, 5 times
as long as deep, twice as long as rayed dorsal. Anal II 7, a
little nearer root of caudal than base of ventral. Pectoral
longer than ventral, a little shorter than head. Caudal feebly
emarginate. Caudal peduncle as long as deep. Dark olive
above, whitish beneath; five round whitish spots on the
back—one in front of the dorsal fin, one on each side of the
last rays of the dorsal, one in front of the adipose fin, and one
on the posterior extremity of the latter.
Total length 60 mm.
A single specimen from Mlanji, British Central Africa,
presented by Mr. R. L. Harger.
Closely allied to A. platychir, Gthr., which occurs also in
British Central Africa, but body shorter and barbels longer.
LXVIU.—Description of a new African Snake of the
Genus Simocephalus. By G. A. BouLencer, F.R.S.
Stmocephalus Butler.
Eye moderately large. Rostral once and a half as broad as
deep, the portion visible from above measuring one third its
distance from the frontal ; internasals once and a half ag
broad as long, half the length of the prefrontals, which are
nearly as long as the frontal; frontal slightly longer than
broad, as long as its distance from the end of the snout,
shorter than the parietals ; loreal deeper than long ; one pra-
and three postoculars ; the lower of the latter may be regarded
as a subocular; temporals 1 +2, the first narrowly separating,
in front, the parietal from the fifth labial; seven upper
labials, third and fourth entering the eye; five lower labials
in contact with the anterior chin-shields, which are longer
than the posterior. Scales in 15 rows, strongly keeled, with
secondary tubercular keels. Ventrals 232; anal entire;
subcaudals 58. Black above, each scale with a whitish basal
spot, white beneath ; ventrals edged with black on the sides;
lower surface of tail greyish.
Total length 400 mm.; tail 55.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx.
©>
bo
490 . Mr. H. B. Preston on new Shells
A single female specimen from between Wau and Chak
Jhak, in the Bahr-el-Ghazal Province, presented to the
British Museum by Mr. A. L. Butler.
This snake appears to be most nearly related to S. Chanleri,
described by Dr. Stejneger from a specimen in bad condition
obtained at the mouth of the Tana River, British East Africa.
But the proportions of the head-shields and the coloration
are too different to justify me in referring the Bahr-el-Ghazal
specimen to that species.
LXIX.—Descriptions of new Species of Land and Fresh-
water Shells from Central and South America. By H. B.
PRESTON, F.Z.S.
Polita peruviana, sp.n. (Fig. 1.)
Shell somewhat depressed, thin, pale brownish horn-
colour; whorls 44, sculptured with transverse, arcuate striz ;
sutures deeply impressed ; umbilicus wide and deep; peri-
stome simple ; aperture lunate.
Alt. 3°25, diam. maj. 7°25 mm.
Aperture: alt. 2, diam. 1°5 mm.
Hab. Chanchamayo, Peru.
Closely allied to P. insignis, D’Orb. *, which occurred with
it; P. peruviana is, however, not so depressed and of a darker
colour, the umbilicus is also rather narrower and the sutures
though well detined do not present the almost channelled
appearance which is the case with P. insignis.
Stephanoda bogotensis, sp.n. (Fig. 2.)
Shell orbicular, somewhat depressed, dark brown, showing
traces of having been covered with a hairy periostracum ;
whorls 54, sculptured with fine, arcuate, transverse, and very
minute spiral striz ; umbilicus very wide and deep ; aperture
lunate.
Alt. 4, diam. maj. 8 mm.
Aperture: alt. 2°5, diam. 1 mm.
Hab. Bogota, United States of Colombia.
Pleurodonte (Labyrinthus) Da Costiana, sp.n. (Fig. 3.)
Shell thin, subdepressed, carinate, pale brownish horn-
colour; whorls 43, obliquely striate with lines of growth and
* Voy. Amér. Mérid. t. 26. ff. 14-17.
from Central and South America. 491
minutely granular ; sutures impressed; umbilicus broad and
deep ; peristome continuous, reflexed; aperture irregularly
auriform, a single broad tooth appearing on the parietal wall
and two teeth on basal margin, the inner strong and simple,
the outer not so strong and bifid.
Alt. 13, diam. maj. 29°75 mm.
Aperture: alt. (including peristome) 10, diam. 14:5 mm.
Hab. Chanchamayo, Peru.
Allied to L. Garbet, Hidalgo, but much larger, more de-
pressed, less solid, and more finely granular than that species ;
it shows no traces of spiral striz and, moreover, has a wider
umbilicus.
Bulimulus (Drymeus) carandaitiensis, sp.n. (Fig. 4.)
Shell narrowly perforate, fusiform, white, painted with
longitudinal bands of brown varying in shade from pale
reddish brown to dark chestnut ; whorls 94, convex, marked
with fine longitudinal lines of growth ; sutures well impressed,
subcrenulate; aperture inversely auriform ; peristome simple ;
columella descending obliquely, reflexed above, thus partly
concealing the umbilicus.
Alt. 35, diam. maj. 14 mm.
Aperture: alt. 14, diam. 6 mm.
Hab. Carandaiti, province of Cordillera, Bolivia, 1000
metres.
Bulimulus (Drymeus) chacoensis, sp.n. (Fig. 5.)
Shell narrowly perforate, subulately fusiform, cream-
coloured, painted with irregular longitudinal bands varying
in breadth and in shade from pale brownish horn-colour to
dark chestnut ; whorls 9, flattish, smooth ; sutures impressed ;
aperture elongately oval; peristome simple, acute ; columella
descending somewhat obliquely and reflexed over the narrow
umbilicus.
Alt. 30, diam. maj. 9°53 mm.
Aperture: alt. 10°75, diam. 3 mm.
Hab. To the north of the Rio Pilecomayo, Chaco, Bolivia,
600 metres.
Bulimulus (Drymeus) nigroumbilicatus, sp.n. (Fig. 6.)
Shell subperforate, fusiform, yellowish white, painted with
longitudinal, arcuate, deep brown and_ blackish bands,
umbilical area deep blackish brown; whorls 8, rather flat,
marked with lines of growth and fine, wavy, spiral strie, the
ane
OL
492 Mr. H. B. Preston on new Shells
Fig. 1.
Fig. 3.
Fig. 5. Fig. 6.
Bulimulus (Drymeus) Bulimulus (D.) Bulimulus (D.)
carandaitiensis. chacoensis. nigroumbilicatus.
Fig. 8. Fig. 9.
LP
Bulimulus (D.) Dulimulus (D.) Bulimulus (D.)
Morenot. comis. solidus.
from Central and South America. 493
‘ig. 10. Ries Te
Bulimulus (D.) Bulimulus (D.) Claustlia (Nenia)
ventricosus. vicinus, chanchamayoensis.
Fig. 15.
=a Limnea Selli.
Clausilia ( Nenia) Slosarskit, Stenogyra
var. Rosenberg? colombiana,
Fig. 16. Fig. 17.
costaricensis.
Fig. 18.
Physa balteata. Physa cornea.
Planorbis
meriddaensis.
494 Mr. H. B. Preston on new Shells
latter especially noticeable on the lower half of the body-
whorl; sutures impressed; aperture receding towards the
Lase, narrowly, inversely auriform ; columella expanded over
the very narrow umbilical perforation ; peristome simple.
Alt. 26°5, diam. maj. 9°5 mm.
Aperture: alt. 10°25, diam. 4 mm.
Hab. Yo the north of the Rio Pileomayo, Chaco, Bolivia,
600 metres.
Bulimulus (Drymeus) Morenoi, sp. n. (Fig. 7.)
Shell ovate conic, narrowly perforate, pale yellowish white ;
whorls 64, convex, marked with lines of growth and showing
traces of very fine spiral strive; sutures impressed and
irregularly crenulate with lines of growth; aperture acumi-
nate, oval; peristome simple; columella descending some-
what obliquely and reflexed over the umbilical region.
Alt. 23°5, diam. maj. 11 mm.
Aperture: alt. 10°5, diam. maj. 5 mm.
Hab. Argentina.
Bulimulus (Drymeus) comis, sp.n. (Fig. 8.)
Shell narrowly fusiform, smooth, milk-white, painted on
the lower portion of the whorls with transverse bands of
purplish and greyish black; whorls 6, marked with very fine
lines of growth; sutures impressed; umbilicus narrow;
aperture inversely auriform; peristome thin, expanded
especially below ; columella twisted into a fold, and diffused
into a pale purple callus which reaches the lip above ;
interior of shell pale purple.
Alt. 28, diam. maj. 12 mm.
Aperture: alt. 11°5, diam. 5°5 mm.
Hab. Bogota, United States of Colombia.
Bulimulus (Drymeus) solidus, sp.n. (Fig. 9.)
Shell fusiform, rather solid, very narrowly perforate,
smooth, polished, cream-coloured, painted with two inter-
rupted spiral bands of greyish black and five wavy transverse
Jines and streaks of greyish purple; whorls 54, slightly
convex; sutures impressed ; aperture ovate, somewhat
oblique; peristome white, rather thick, slightly reflexed ;
columella descending obliquely and reflexed over the very
narrow umbilicus, white below, purple above and diffused
into a purple callus which joins the upper margin of the lip ;
interior of shell purple.
‘
from Central and South America. 495
Alt. 32°5, diam. maj. 15 mm.
Aperture: alt. 14, diam. 6 mm.
Hab. Bogota, United States of Colombia.
Bulimulus (Drymeus) ventricosus, sp.n. (Fig. 10.)
Shell fusiform, rather tumid, smooth, pale flesh-colour,
painted with transverse streaks of reddish and purplish brown
and with two interrupted bands of brownish black, which
increase to three on the last whorl; whorls 54, flattish ;
sutures impressed ; aperture inversely auriform ; peristome
thin, white, reflexed; columella slightly arched, reflexed
outwards over the narrow perforation and diffused into a
purple callus which reaches the lip above.
Alt. 34°75, diam. maj. 16°5 mm.
Aperture: alt. 17, diam. 7 mm.
Hiab. Bogota, United States of Colombia.
Bulimulus (Drymeus) vicinus, sp.n. (Fig. 11.)
Shell fusiform, narrowly perforate, smooth, yellowish fawn-
colour, with occasional transverse streaks of a darker shade ;
whorls 54, marked with somewhat coarse growth-lines ;
sutures impressed, faintly crenulate with the lines of growth ;
aperture ovate, much contracted below; peristome white,
slightly reflexed ; columella somewhat oblique and spreading
into a purple callus which reaches the lip above; interior
of shell mauve.
Alt. 33, diam. maj. 15 mm.
Aperture: alt. 13°5, diam. 6 mm,
fab. Bogota, United States of Colombia.
Clausilia (Nenia) chanchamayoensis, sp.n. (Fig. 12.)
Shell fusiform, rather thin, light brown, streaked and
mottled with greyish white; whorls 7, finely and closely
transversely striate; sutures well impressed; aperture
ovate, oblique ; peristome expanded, reflexed above; lamella
superior strong and reaching to the outer edge of the peristome;
Jamella inferior not so strong and reaching only to the inner
edge of the peristome.
Alt. 13, diam. maj. 3 mm.
Aperture (including peristome) : alt. 3°75, diam. 2°75 mm.
Hab. Chanchamayo, Peru.
Clausilia (Nenia) Slosarskit, Lub., var, Rosenberg’, var. n.
(Fig 13)
More coarsely transversely striate and much less closely
spirally striate than is the typical form,
496 Mr. H. B. Preston on new Shells
Hab, Pozuzo, Peru, 800 metres.
Among the fifteen specimens examined there appears to be
a good deal of difference in the diameter of the shell, and I
was at first inclined to think that there might be two distinct
species represented; but as they all agree in every other
respect, I am satisfied that they all belong to the same form.
Stenogyra colombiana, sp.n. (Fig. 14.)
Shell subperforate, fusiform, vitreous, pale straw-colour ;
apex obtuse ; whorls 64, sculptured with very fine transverse
coste ; sutures impressed, subcrenulate, and discoloured with
a narrow band of reddish brown, which appears to be painted
on the interior of the shell; columella descending perpendi-
cularly, slightly excavated and joined to the lip above by a
very thin callus,
Alt. 28°5, diam. maj. 12 mm.
Hab. Bogota, United States of Colombia.
Limnea Selli, sp.n. (Fig. 15.)
Shell ovate, thin, semipellucid, light brownish horn-colour ;
whorls 4, sculptured with very minute punctate spiral striz
or scratches and fine transverse lines of growth; sutures
impressed ; aperture ovate, rather dilated below; peristome
simple; columella descending obliquely, reflexed outward,
thus partly concealing the narrow perforation and spreading
above into a thick callus which reaches the upper margin of
the peristome.
Alt. 12, diam. maj. 7 mm.
Aperture : alt. 7, diam. 3°5 mm.
Hab. Bogota, United States of Colombia.
Planorbis costaricensis, sp.n. (Fig. 16.)
Shell pale brown, corneous, depressed, concave both above
and below; whorls 4—5, sculptured with coarse oblique lines
of growth, the body-whorl inflated and extending upwards
and outwards towards the aperture ; aperture obliquely lunate;
peristome acute, bent slightly outwards, a callus uniting the
points of contact with the whorl.
Alt. maj. 9°5, diam. maj. 26 mm.
Aperture : alt. 9, diam. 6 mm.
Hab, Catalina, Province of Guanacaste, Costa Rica.
This shell is in many respects allied to P. guadeloupensis,
Sow., but is much more inflated than is the case with that
species, and there are no traces of a peripheral keel; more-
from Central and South America. 497
over the upward extension of the last whorl near the aperture,
so noticeable in the present species, is totally lacking in
P. quadeloupensis.
Planorbis Boucardiana, sp.n. (Fig. 17.)
Shell depressed, subcarinate, whitish grey; whorls 4,
sculptured with fine oblique striz ; spire concave ; umbilicus
wide, very concave, especially in the centre; peristome
simple, very oblique, broadly lunate.
Alt. 2, diam. maj. 7 mm.
Aperture : alt. 2, diam. 1°25 mm.
Hab. Mexico.
Planorbis meridaensis, sp.n. (Fig. 18.)
Shell depressed, suborbicular, dull brown; whorls 35,
marked with oblique arcuate lines of growth; right side
deeply excavated, especially in the middle ; left side some-
what concave; peristome acute, a callus ‘on the parietal
whorl joining the two margins of the peristome; aperture
obliquely lunate.
Alt. 3, diam. maj. 8 mm.
Aperture: alt. 3, diam. 1°5 mm.
Hab. Merida, Venezuela.
Physa balteata, sp.n. (Fig. 19.)
Shell fusiform, pale yellowish horn-colour, transversely
marked with oblique, buff-coloured bands; whorls 6, the last
somewhat inflated and sculptured with very fine transverse
strie; sutures rather deeply impressed; edge of peristome
acute, a thickening appearing just within the shell ; columella
straight, white, descending somewhat obliquely, a very thin
callus joining it with the lip above; aperture elongately,
inversely auriform.
Alt. 12°5, diam. maj. 6°75 mm.
Aperture : alt. 8, diam. 3°5 mm,
Hab. Oaxaca, Mexico.
Physa cornea, sp.n. (Fig. 20.)
Shell fusiform, thin, polished, transversely striated with
irregular and somewhat distant lines of growth and spirally
sculptured with very fine wavy strie; whorls 5}; sutures
impressed ; peristome acute; columella twisted, descending
498 On a new Acanthoglossus.
somewhat abruptly, a light callus joining it with the lip
above ; aperture elongately, inversely auriform.
Alt. 11°5, diam. maj. 6 mm.
Aperture: alt. 7:5, diam. 3 mm.
Hab. Merida, Venezuela.
Allied to P. venzuelensis, Mart. ; it is, however, separable
from that species by its more twisted columella and its less
opaque texture, it is also more conspicuously transversely
striate than is the case with that species. The fine spiral
striza mentioned above are only visible by the aid of a strong
lens.
LXX.—A new Acanthoglossus from the Island of Salawattt.
By OLbFIELD THOMAS.
THE National Museum owes to the generosity of Mr. Walter
Goodfellow a fine Long-nosed Echidna which he obtained in
the island of Salawatti, and kept alive, with another specimen,
for some months.
The genus has not hitherto been recorded out of New
Guinea itself, and there mainly or entirely at high altitudes,
for which the thick coat present in A. Brutjnii admirably
suits it. But the island of Salawatti being throughout com-
paratively low, it is not surprising that the Acanthoglossus
occurring there should be different in the development of its
coat from its New Guinea ally.
I would propose to call the new form
Acanthoglossus Goodfellowt, sp. n.
Coat much more spinous and less hairy than in any of the
forms of A. Bruynit. Spines extending on the under surface
nearly to the middle line of the belly, and though absent in
the groin and between the fore limbs they reappear on the
chest and throat to the middle line, though small and far
apart. In A. Bruynii the under surface is without spines
throughout.
Spines of upper surface averaging about an inch in length
and 2°5 mm. in thickness, the longer ones attaining 30 mm.
with a thickness of 3 mm.; the small spines of the chest and
belly about 12x 1mm. _ In colour all over the body they are
white, shading basally into grey. Fur short and scanty, the
skin and bases of the spines not hidden; the hairs only about
half an inch in length ; uniformly black throughout. On the
On a new Baboon from British East Africa. 499
chest and groin the hairs are coarse and bristly, blackish on
the former, yellowish grey on the latter. Hands and feet
thinly covered with short coarse bristles. Claws 3—3, as
usual *,
Skull without marked special characters, its dimensions as
follows :—
Greatest breadth 58 mm.; palate length 160; length of
rostrum 119; palatal foramina 37°5.
flab. Island of Salawatti, N.W. of New Guinea.
Type. Old female. B.M. no. 7. 9. 5. 1. Collected
October 1906, and presented by Walter Goodfellow, Esq.
The predominance of the spines and the almost entire
suppression of the woolly coat will readily distinguish the
Salawatti form from its thickly clothed New Guinea ally.
LXXI1.— Description of a new Baboon from British East
Africa. By D. G. Exutor, D.Sc., F.R.S.E., &c.
Family Cercopithecida.
Genus Papio.
Papio furax, sp. n.
Type locality. Baringo, North-west of Mt. Kenia, Hast
Africa.
Gen. char. Resembles P. doguera from Abyssinia, but
darker and cranial characters very different. The rostrum is
shorter and much broader and flatter, and the nasals do not
rise above the piane of the rostrum, but are flat, not rounded ;
the palate is flatter and the distance between the last molar
and the palatal arch is much greater; the intertemporal
width is much less, as is also the width of the brain-case; the
pit on the side of the jaw is broader, shorter, not so deep,
and the zygomatic width much less.
Colour. General colour seal-brown, the hairs banded with
buff, becoming ochraceous buff on the rump; the black tips
of the hairs so arranged over the ochraceous buff as to form
bands of black, though not distinctly defined ; limbs black
and cream-colour, the hairs having bands of those colours
with black tips; chest black and cream-colour; abdomen
* When describing A. B. Bartont, I omitted to mention that the type
had no less than five claws on both fore and hind feet; but considerable
variation in this respect has already been recorded,
500 Mr. R. C. Wroughton on
black, the hairs banded with ochraceous buff; hands mixed
black and ochraceous buff; feet black, with the base of the
hairs buff; tail mixed black and whitish yellow.
Measurements. Size about the same as P. doguera. Skull:
total length 196 mm.; occipito-nasal length 166; hensel 141;
intertemporal width 56; zygomatic width 122; palatal
length 93; breadth of brain-case 82; length of nasals 77 ;
length of upper molar series 48; length of mandible 145 ;
length of lower molar series 65.
Type. Adult male in British Museum. No. 1. 8. 9. 20.
Presented by Sir H. H. Johnston.
‘'wo specimens were procured by Sir H. H. Johnston, one
at Baringoand one at Nandi, north of Mt. Kenia. Compared
with Abyssinian examples of P. doguera the present species
is a much darker animal and the bands on the hairs are paler
and more yellow, those of P. doguera being more ochraceous,
with less of the black showing. The general tone of the
Baringo specimen, while seal-brown, has a greyish tinge,
while that of Abyssinian examples is more ochraceous. ‘The
cranial characters, however, will easily distinguish the present
form from its ally.
I am indebted to my friend Mr. Oldfield Thomas for the
privilege of describing the species.
LXXII.—List of a Collection of small Mammals made by
Mr, A. L. Builer in the Bahr-el-Ghazal. By R. C.
WROUGHTON.
THE following is a list of the small mammals collected by
Mr. Butler between Renk, on the White Nile, and Dem
Zubeir, in the Bahr-al-Ghazal. ‘The collection, though small,
was obtained under trying conditions. It is of special
interest, not so much on account of the new forms it contains,
as that it was made in aregion whence little has been brought
since Heuglin’s collection nearly fifty years ago.
1. Epomophorus anurus, Heug.
6-003 2). 51. Dad Majok. 225i. .07,
‘Lopotypes of Heuglin’s species.
2. Lavia frons affinis, And. & Wrought.
Ow 1595 2. Renlee WihitesNule yin ieOle
Mammals from the Bahr-el- Ghazal. 501
3. Cherophon sp.
O32.) GamateZan nO One
4, Crocidura sericea, Sund.
od. 41,45; 9. 40,44. Chak Chak. 13. ili. 07.
5. Crocidura sp.
6. 33. Between Chak Chak and Dem Zubeir. 7. ili. 07.
6. Funisciurus multicolor, Riipp.
@- Li Katha (Kor 'Gitty)sseo le One
6 43. Chak: Chak, 13.07
7. Xerus erythropus leucombrinus, Riipp.
6. 22; 2. 20: Chak Chak -22/245 i, 07.
8. Tatera Emini, Thos.
O20,-0;, 2-245 10e(juv.)}e. Chak Chaks 9 19.15 07,
9. Tatera macropus, Heug.
6. 40> 9. 15523 Guv.). Chak Chak. 1, 07.
10. Yatera robusta, Wagn.
3. 30; 2. 31. Halfway between Chak Chak and Dem
Zubeirs de tl. Ots
11. Mus sp. (multimammate).
- 9. Moyen. 211. 07.
o LZ, -Kathas eal 07:
. 26,42; 9. 138,14. Chak Chak. * 13/16. iii. 07.
Oy +0 +0
12. Mus alghazal, sp. n.
3. 24,27. Chak Chak. 24/25. ii. 07.
d. 28,36. Between Chak Chak and Dem Zubeir.
2 Quite Olt:
A rat rather smaller than W. Hindei, Thos.
Fur soft and silky, about 10 mm. long on the back.
General colour above dull tawny (near “ russet”’), the hairs
slate-grey for basal two thirds, then “ ochraceous buff,” a
considerable proportion of somewhat longer wholly black
502 Mr. R. C. Wroughton on
hairs scattered among the rest ; underside white, basal fourth
of hairs slaty. Head and face like back, but a bright buff
patch on cheek below eye. Upper lip and chin white like
belly. Hands and feet pure white. ‘Tail sparsely clad ;
rings 8=10 mm.
Skull markedly smaller in all details than that of 1. Hindet,
except the teeth, which are about the same size.
Dimensions of the type :—
Head and body 147 mm.; tail 156; hind foot 28; ear 16.
Skull: greatest length 36; basilar length 29; greatest
breadth 18; nasals 14; pnrcrorbitall breadth OG iiemeenss
breadth 14°5 ; diastema 9°5 ; length of upper molar series 6 ;
bullee 7.
Hab. Chak Chak, Bahr-el-Ghazal.
Type. Adult male. Collector’s number 24. Taken
25th February, 1907.
Four specimens (unfortunately all males) examined.
13. Mus Blainet, sp. n.
Qe gos | SAMs aoe ON.
O19. ‘ChakiChak:: 227 170%
g. 39. Between Chak Chak and Dem Zubeir. 9. 111. 07.
g. 47. Between Chak Chak and the Pongo. 19. 111. 07.
A small soft-furred mouse with a rather short tail.
Size about as in MM. Allent and M. Dennie, but with a
quite short tail.
Fur soft and silky, about 10 mm. long on the back.
General colour above “ isabella,” the hairs slate-colour
from their bases for three fourths their length, terminal
fourth pale orange-buff with black tips; underside white,
the basal half of hairs slate-colour. Face below the eyes,
sides of neck, and flanks “ orange-buff” with a mixture of
“vinaceous buff”; chin and upper lip white; hands and
feet white; tail sparsely haired, darker above than below,
with about 2 rings=1 mm,
Skull and teeth of the usual type in Afus.
Dimensions of the type :—
Head and body 100 mm.; tail 103; hind foot 22; ear 20.
Skull: greatest length 28 « basilar length 22; greatest
breadth 13° 5; nasals 10; interorbital breadth 4:3: brain-
case breadth 11°3; disseny) 78; length of upper molar
series 4°5 ; bulls 5.
Hab. Chak Chak, Bahr-el-Ghazal.
Type. Young female. Collector's number 19. Taken
22nd February, 1907.
Mammals from the Bahr-el- Ghazal. 503
Five specimens examined.
I have named this species after Mr. Gilbert Blaine, who
was Mr. Butler’s companion on the tour during which the
collection was made.
14. Mus Butleri, sp. n.
9. 37. Between Chak Chak and Dem Zubeir. 8. ili. 07.
A mouse rather smaller than the last, with a longer tail
and pure white beliy.
Size rather smaller than M. Blatne.
Fur soft and silky ; 6-8 mm. long on the back.
General colour above ‘ clay-colour,” the hairs slate-
colour from their bases for two thirds their length, terminal
third a bright fawn, with short black tips; underside pure
white, the hairs white to their bases ; face below the eyes and
sides of the neck and body fawn; line between upper and
belly colours sharply defined; feet white; tail sparsely
haired, darker above than below, about 7-8 rings=5 mm.
Skull and teeth of the type normal in the genus Mus.
Dimensions of the type :—
Head and body 90 mm. ; tail 118; hind foot 20; ear 16.
Skull: greatest length 27; basilar length 21; greatest
breadth 13; nasals 10; interorbital breadth 4; brain-
case breadth 12 ; diastema 7 ; length of upper molar series 4°5 ;
bulle 5.
Hab. Between Chak Chak and Dem Zubeir, Bahr-el-
Ghazal.
Type. Young female. Collector’s number 37. Taken
8th March, 1907.
Though the size and proportions of the skull are so similar
in the two species, the present form is separable at once from
the last by its proportionally much longer tail and snow-white
belly. ‘he species most closely resembling this one is
perhaps MJ. Dennie, Vhos., from Ruwenzori; but in that
animal the tail is proportionally still longer and the slaty
bases of the hairs of its underside distinguish it markedly
from M, Butler.
15. Arvicanthis zebra, Heugl.
§. 48,49. Dad Majok. 30. iii. 07.
'‘l’opoty pes of Heuglin’s species.
5OL On Mammals from the Bahr-el- Ghazal.
16. Arvicanthis sp.
3. 34, 35, 38. Between Chak Chak and Dem Zubeir.
7/9. iii. 07.
17. Thamnomys Macmillani, sp. n.
Gals 20. “Chek Chak, 92401007,
A small Thamnomys of the T. arborarius group.
Size as in 7. arborarius.
Fur soft and silky, about 10 mm. long on back.
Jeneral colour above a dark “ olive-buff,” with a good
deal of ochraceous colouring on the median line, especially
bright on the rump, basal three fourths of hairs dark slate ;
underside pure white, the hairs white to their bases. Feet
pale buff. Tail dark, pale below for basal half; 17 rings=
10 im. ; sparsely clothed with dark hairs; hairs short for
basal half of tail, then lengthening to 3-4 mm. on terminal
fourth.
Skull and teeth as in 7. arborarius, but markedly smaller
in all details judging from Peters’s description and figure.
Dimensions of type :—
Head and body 102 mm.; tail 160; hind foot 22; ear 15.
Skull: greatest length 28; basilar length 21; greatest
breadth 13°5 ; nasals 9°5; interorbital breadth 4°3; brain-case
breadth 12; diastema 6°7 ; length of upper molar series 4°3 ;
bullee 5.
Hab. Wouida, N. of Lake Rudolf. Alt. 6200!.
Type. Adult female. B.M. no. 6.11. 1.38. Taken by
Mr. Zaphiro on the 30th June, 1905, and presented to the
British Museum by Mr. W. N. McMillan.
Two specimens sent by Mr. Butler from Chak Chak are
quite like the type in all essential characteristics (their skulls
unfortunately are imperfect). The colour-pattern is also
similar, but the grey-drab shoulder-patches are even more
clearly marked in Mr. Butler’s specimens.
18. Thryonomys Swinderenianus, Temm.
Skull only. Chak Chak. 1. ii. 07.
19. Gazella albonotata, Roths.
No specimen of this striking species had hitherto been
obtained by the National Museum.
On new Species of Lleterocera. 505
LX XIII.—Descriptions of Five new Species of [leterocera.
By HerBert Druce, F.L.S. &e.
Fam. Castniide.
Castnia angusta, sp. n.
Male.—Head, collar, tegule, thorax, and abdomen dark
brown; legs blackish brown. Primaries dark brown, bluish at
the base; the veins near the outer margin irrorated with pale
blue scales ; a pale yellow spot at the end of the cell, beyond
which a narrow yellow line crosses the wing from the costal
to the inner margin near the anal angle; the fringe yellow :
secondaries dark brown, shot with blue at the base, crossed
beyond the middle by a band of narrow, crescent-shaped,
yellow spots, which extend from near the apex to the anal
angle; asubmarginal row of six rather large round yellow
spots, starting from the apex; the fringes of both wings
yellow. Underside considerably paler brown: primaries,
the yellow band much wider than above, a submarginal row
of eight yellow spots extends from the apex to the anal angle ;
secondaries crossed by two rows cf yellow spots, the first
below the middle, the second submarginal, on the outer
margin the spots are edged with reddish brown.
Expanse 64 inches.
Hab. Ecuador, Los Lanos (Mus. Druce).
Castnia dolopia, sp. n.
Female.—Head, collar, tegule, thorax, and base of the abdo-
men dark brown; abdomen black ; antennze black, the tips pale
brown. Primaries dark brown glossed with green; a large
greyish-brown spot at the end of the cell, beyond which the
wing is crossed trom near the apex by a series of dull greyish-
brown spots, those nearest the apex very indistinct, the spots
are edged with black ; below the spots on the inner margin are
two lunular-shaped black marks : secondaries black, the base
shot with bluish green ; a row of brownish-white spots crosses
the.wing from near the anal angle almost to the apex, the
fringe brownish white. ‘The underside of both wings pale
brown, with all the spots much more distinct and all edged
with black.
Expanse 7? inches.
Hab. Ecuador, Los Lanos (Mus. Druce).
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 33
506 On new Species of [eterocera.
Fam. Thymaride.
Pedoptila thaletes, sp. n.
Female.—Head, collar, tegule, thorax, and abdomen
reddish brown ; abdomen banded with black ; the antenne,
underside of the abdomen, and the legs black; the anus
clothed with greyish hairs. Primaries, the basal half reddish
orange, the outer half black, the veins black ; secondaries
reddish orange as far as the lobe, which is black almost to the
end of the wing; the tip of the tail white.
Expanse 12 inch.
Hab. Cameroons, Ja River (Mus. Druce).
Fam. Syntomide.
Listiva falerina, sp. n.
Male.—Head, antenne, collar, tegule, thorax, and base of
abdomen black; head and collar spotted with metallic blue ;
abdomen metallic blue; legs black. Primaries black, the
basal half of the wing red streaked with black; a broad
oblique yellow band beyond the cell, a metallic-blue spot at
the base, and a blue streak at the end of the cell : secondaries
red, edged with black, widest at the apex and the middle of
the outer margin.
Expanse 22 inches,
Hab. Peru, Marcapata (Mus. Druce).
Fam. Arctiidae.
Automolis marcapata, sp. n.
Female.—Head, antenne, collar, tegule, thorax, and abdo-
men reddish yellow; legs reddish. Primariesreddish, irrorated
near the base with yellow scales ; a large semihyaline yellow
blotch at the end of the cell extending to the costal margin ;
the veins and the edges of the large blotch dark red; the
fringe brownish red: secondaries deep pink, the fringe
yellowish.
Expanse 2 inches.
Hab. Peru, Marcapata (Mus. Druce).
Allied to Automolis sanguinolenta, Cram., but very
distinct.
On new African Species of the Genus Chrysops. 507
LXXIV.— New African Species of the Genus Chrysops,
Meigen, in the British Museum (Natural History). By
Ernest E. Austen.
Tue latest list of African representatives of this genus *
includes the names of eighteen species. As stated in the
“* Notes ” at the end of the present paper, however, two of these
are synonyms. Six new species are described in the
following pages, bringing the number of known African
forms of these handsome and bloodthirsty flies up to twenty-
two. The Museum collection contains examples of at least
two additional species, of which descriptions will be published
in a subsequent paper. The types (or co-types) of all the
new species described below are in the British Museum
(Natural History).
Genus Curysops, Meigen.
Chrysops funebris, sp. n.
?.—Lengtht (5 specimens) 9 to 104 mm.; width of
head 24 to 23 mm.; width of front at vertex ? mm. ; length
of wing 9 to 10 mm. ’
Uniformly black, though the usual tufts of golden pile are
present below humeral calli and on mesopleure ; dorsum of
thorax (denuded) thinly covered with greyish dust ; abdomen
(except a dull black area on the middle of the first segment,
below the scutellum) shining, clothed above with minute,
sparse, whitish hairs; wing-markings uniformly dark brown
or black, distal margin of transverse band straight cr nearly
so; first and second joints of middle and hind tarsi yellowish
white, tips of these joints usually brownish.
Head.—¥ rontal callus large, nearly semicircular in out-
line, almost touching the eye on each side, and separated
from the black ocellar region by a narrow band of greyish
pollen; a small patch of golden pollen between bases of
antennze (perhaps produced downwards in specimens in per=
fect condition) ; face considerably produced downwards and
forwards, very protuberant, separated from the shining black
jowls by a strip of golden pollen running down from the
front margin of the eye; under surface of head thinly clothed
* Cf. Bezzi, Bull. Soc. Ent. Ital., Anno xxxvii. 1905, p. 234.
+ The length in all cases is measured from the front of the face to the
tip of the abdomen, and is exclusive of the antenne.
By
508 Mr. E. E. Austen on new
with rather long yellowish hair ; palpi hazel-brown*, very
slender ; antenne slender, first joint not at all incrassated
and only slightly longer than the second ; second and third
joints dark brown, first joint lighter (brownish tawny) except
at distal extremity. Thorax with a tuft of golden pile
on the metapleure, besides the tufts already mentioned.
Abdomen: the short whitish hair clothing the dorsum some-
what longer and more conspicuous on the sides of the first
and second segments than elsewhere; under side of second
segment also clothed with short whitish hairs ; remainder of
venter inconspicuously clothed with blackish hair. Wings
with extreme base, costal cells, basal fourth of first basal
and basal sixth of second basal cell, marginal cell from a
point above the origin of the third vein to the end, and a
continuation (apical blotch) which fills out the apical portion
of the first submarginal cell, and the upper angle of the
second submarginal, shading off below, dark brown; the
proximal margin of the transverse band runs from the origin
of the third longitudinal vein to the posterior upper angle of
the fifth posterior cell, and thence (sometimes obliquely for-
wards) to the sixth vein, on which it ends; the distal
margin of the transverse band runs from a point below the
second longitudinal vein, and a little beyond the level of the
end of the first longitudinal, to the hind margin of the wing
at the end of the vein separating the third and fourth
posterior cells. Proximally, the transverse band thus fills
out the extreme tips of the first and second basal cells, and
the end of the anal cell; distally it includes the base of the
second posterior cell, and diagonally bisects the third
posterior cell. The whole of the discal cell is included, but
there is sometimes a tiny hyaline fleck at its extreme base.
The apical portion of the wing below the apical blotch is
entirely infuscated with a brownish suffusion, and the
infuscated area is separated from the transverse band by a
milky streak, which ends on the vein separating the second
and third posterior cells. Halteres dark brown. Legs :
coxee dark brown ; femora reddish brown, tips of hind pair
_ darker; tibiz, except extreme tips, front tarsi, and last two
joints of middle and hind tarsi dark brown ; third joints of
middle and hind tarsi brownish; front and middle tibiz in-
crassated, hind tibize also somewhat incrassated.
Uganda: type and three other specimens from north-east
side of Lake Albert Edward, 1906 (Dr. A. D. P. Hodges) ;
* For names of colours see Ridgway, ‘A Nomenclature of Colors
for Naturalists’ (Boston: Little, Brown, and Company, 1886),
African Species of the Genus Chrysops. 509
a fifth specimen from the shore of Lake Victoria, in Buddu,
1903 (Sleeping Sickness Commission of the Royal Society,
per Colonel David Bruce, C.B., R.A.M.C., F.R.S.).
Of the African species already described, Chrysops funebris
appears, so far as may be judged from the description, most
nearly to resemble C. confluens, Lw., from the Cape of Good
Hope, from which, however, it may at once be distinguished
by the slender antennz, single ‘frontal callus, and very
different wing-markings.
Chrysops silacea, sp. u.
? .—Length (17 specimens) 8 to 10 mm.; width of head
2% to 3 mm.; width of front 1 mm.; length of wing 7} to
91 mm.
Head yellow ; thorax black or blackish brown, with yellow
longitudinal stripes; scutellum ochre-yellow, median portion
of extreme base black ; abdomen ochraceous, upper side marked
on basal half with two longitudinal black stripes, not reaching
hind margin of third segment, and sometimes not extending
beyond second ; distal portion of wing from base of third vein
wnfuscated, transverse band inconspicuous, reduced to a small
dark blotch extending from base of third vein to posterior
upper angle of fifth posterior cell; legs ochraceous, front tarsi
and tips of middle and hind tarsi dark brown.
Head rather broad, dark brown on vertex, elsewhere
clothed with ochre-yellow pollen, except on frontal callus, a
mark above base of each antenna, the facial tubercles, and
a small dark brown spot on the jowls below each eye; frontal
callus of moderate size, three fourths as wide as the front
and well separated from the eye on each side, not highly
polished, edged with brown at the sides or sometimes all
round the upper curved margin; facial tubercles highly
polished, elliptical in shape, occupying lower two thirds of
face and extending nearly to margin of buccal cavity,
separated from each other by a narrow stripe of yellow
pollen ; often only the upper ‘portion of the pollinose stripe
is present, when the tubercles coalesce below and assume
the appearance of a single elongate cordate tubercle, which
extends to the lower margin of the face; front, except on
callus, clothed with short brown hair, densest on vertex ;
sides of face and lower surface of head clothed with pale
golden hair; palpi lanceolate, widest part of moderate
breadth, thinly clothed with minute yellowish hairs ;
antenne 31 mm. in length, dark brown, basal portion of
slender first joint tawny ochraceous. Thorav clothed with
510 Mr. E. E. Austen on new
golden-yellow pile, which is thicker and longer on the sides ;
the dorsum exhibits three well-defined blackish-brown
stripes, the median stripe being separated from the others
by a pair of dull ochraceous stripes, which meet together in
front and taper off posteriorly, where each is continuous
with a broad, lateral, light yellow stripe; the median
blackish-brown stripe tapers off in front and extends to the
hind margin; the adlateral blackish-brown stripes, which
are broader behind the transverse suture, are rounded off in
front and behind and do not reach the hind margin; pleurz
marked with a broad, curving, yellow pollinose stripe, thickly
clothed with golden-yellow pile, commencing below the
humeral callosity, encircling the upper part of the meso-
pleura, and extending to the metapleura; sternopleure
yellowish pollinose, golden yellow behind, where they are
clothed with similarly coloured hair; area between front
cox also clothed with golden-yellow pollen and_ hair.
Abdomen clothed above and below with minute orange-
ochraceous hairs ; the black stripes, which commence on
hinder half of first segment, vary greatly in width in
different specimens, and are often somewhat irregular in
outline ; they are sometimes interrupted on hind margin of
first segment, and may terminate before reaching hind
margin of second, to reappear as basal spots on the third
segment ; lateral margin of second, and sometimes of third
and fourth segments as well, narrowly edged with dark
brown or black ; venter unicolorous, a small median black
streak occasionally present on the second and third segments.
Wings: costal cells ochre-yellow; stigma ochraceous ;
whole of apical portion, from a point on costa above fork of
third vein to tip of anal cell, suffused with brown, including
the whole of the fifth posterior cell, but not the bases of the
second, third, and fourth posterior cells; there is thus a
paler, buff-coloured area between the apical blotch and the
remains of the transverse band; the latter includes the tips
of the first and second basal cells and the basal third of the
discal cell ; lower portion of second basal cell usually with a
slight yellowish tinge. Halteres dark brown. Legs: cox
dark brown (front pair paler), clothed with golden-yellow
hair; femora and tibize clothed with minute orange-ochra-
ceous hairs; front tibiz somewhat incrassated, their distal
portion brownish.
Congo Free State; Southern and Northern Nigeria.
Type and four other specimens from the Congo Free State
(Kimwenza, 28. in. 1904; Yakusu, 15. 1x. 1904 ; Kimwosa,
African Species of the Genus Chrysops. 511
Leopoldville, 1904; N’Kussu, Cataract Region, 25. xi. 1904:
Drs. Dutton, Todd, & Christy) ; other specimens from Old
Calabar, Southern Nigeria (the late Miss M. H. Kingsley) ;
Ohumbele, Southern Nigeria, 6. viii. 1900 (Dr. H. E. Annete) ;
Odut and Uwet, May 1906 (G. C. Dudgeon); Sapele,
Northern Nigeria, 1906 (received from Dr. G. F. Darker) ;
Akwatcha, Bassa Province, Northern Nigeria, July 1906,
Jauuary and February 1907 (Dr. G. J. Pirie). According
to Dr. Pirie C. silacea is numerous at Akwatcha in the dry
season ; and at Sapele, in Northern Nigeria, it is stated by
Dr. Darker to be troublesome to human beings.
This species must not be confused with Chrysops dimidiata,
v. d. Wulp, the wing-markings of which are identical with
those of C. silacea, while the buff-yellow abdomen is also
adorned with two black longitudinal stripes. C. dimidiata
may be distinguished by the considerably larger and more
prominent frontal tubercle, whichis only narrowly separated
from the eye on each side ; by the paler (buff-yellow) ground-
colour of the abdomen ; and by the fact that the abdominal
stripes are much broader and reach the fourth segment, when
they either become merged in the dark brown colour of the
terminal segments, as stated by van der Wulp, or meet to-
gether on the sixth segment soastoformaloop. In C. silacea
the black stripes are sometimes so attenuated or interrupted
as to be easily overlooked, and in the case of one specimen
from Odut, Southern Nigeria, little more of them remains
than a couple of small dark flecks on the hind margin
of the first segment, and one or two indistinct markings
on the second.
The range of C. dimidiata, which was described (‘ Notes
from the Leyden Museum,’ vii. 1885, p. 80) from Chimfimo,
Portuguese West Africa, evidently overlaps that of C. silacea,
since the Museum possesses a series of specimens of the
former species from Ologbo, Southern Nigeria, May 1906
(G. C. Dudgeon).
Eye-markings *.—In two specimens from Sapele, pre-
served in spirit, the occipital border is fairly broad, and in
contact with the occipital margin and with the upper frontal
spot, which is also connected with the frontal margin. The
occipital border is also in contact with the shaft above and
the arrow-head below, which are continuous with each other,
* For explanation of terms used see V. A. E. Daecke’s interesting
paper, “ On the Eye-Coloration of the Genus Chrysops” (‘ Entomological
News,’ vol. xvii. 1906, pp. 89-42, pl. 1.).
512 Mr. E. E. Austen on new
forming a zigzag line. The middle and lower frontal spots
are somewhat narrow and in contact with the frontal
margin *,
Chrysops Wellmanii, sp. n.
? .— Length (5 specimens) 7 to 72 mm.; width of head
24 to 24 mm.; width of front at vertex 1mm.; length
of wing 7 to 74 mm.
Head wide, flattened from front to rear ; face whitish grey ;
frontal callus, vertex on each side, and antenne shining black ;
Jjirst joint of antenne elliptical in outline when seen from
above, strongly incrassate, twice the length of the second joint
and, at the widest part, one and a half times as broad ; thorax
uniformly plumbeous above and below, without markings ;
abdomen varying from ochraceous-buff to tawny, brownish at
the tip; wings hyaline, extreme base, costal border, and
sharply defined transverse band, tapering to hind margin,
blackish brown; legs, except cinereous coxe, black.
Head.—F¥rontal callus shallow from above downwards, but
very wide, extending from eye toeye; lower margin straight,
sharply defined, separated from the bases of the antennz by
a narrow transverse band of the whitish-grey pollen which
covers the face; frontal callus somewhat triangular in out-
line, the upper margin being produced in the centre so as to
include the anterior ocellus ; above the callus is a narrow
band of grey pollen, in the middle line sending an offshoot
to the occiput, and leaving on each side of the vertex a
somewhat quadrate, shining, black area; occipital region
grey ; face and jowis entirely whitish-grey pollinose, without
any shining tubercles, but oral margin blackish in front;
face and underside of head clothed with long whitish hair ;
pal cinereous, elliptical in outline, clothed with whitish
hair; first jomt of antennze clothed with blackish hair
(third joint wanting). Thorax clothed with long whitish
hair on sides and beneath. Post-alar calli with a reddish
tinge. Scutellum blackish, hind margin fringed with whitish
hair. Abdomen: a somewhat quadrate dark brown patch on
the middle of the first segment beneath the scutellum,
shading off to the hind margin; extreme lateral margins of
* Chrysops silacea is the species referred to by Newstead (‘ Annals of
Tropical Medicine and Parasitology, vol. i. no. 1 (February, 1907), p. 43,
pl. iv. fig. 1) as “ Chrysops dimidiatus, v.d. Wulp.” I am responsible
for this identification, which I formerly believed to be correct, and it
was not until a series of the true C. dimidiata, v. d. Wulp, was received
from Mr, Dudgeon, as mentioned above, that the error became evident.
African Species of the Genus Chrysops. 513
second and lateral margins of third and following segments
dark brown; on ventral side fifth and following segments
usually dark brown, hinder portion of fourth segment often
more or less brownish ; abdomen clothed above with short
yellowish or whitish hair, beneath with short whitish hair,
sides fringed with long whitish hair. Wings: bases as well
as extreme tips of first and second basal cells blackish brown
apical blotch narrow, extending from the upper end of the
distal margin of the transverse band to a little way beyond
the termination of the upper branch of the third longitudinal
vein ; distal margin of transverse band somewhat irregular,
commencing on the costaa little beyond the level of the fork
of the third vein, and terminating at the tip of the anal cell ;
in the fourth and fifth posterior cells it is somewhat indis-
tinct, and less sharply defined than above; proximal margin
of the transverse band commencing at the base of the third
and running at right angles to the costa almost straight
down to the sixth vein, sometimes with a slight backward
projection at the upper inner angle of the fifth posterior
cell ; veins in hyaline area on basal side of transverse band
pale (cream-colour) ; veins beyond transverse band also partly
pale. Halteres brown, Legs: front tibiz alone incrassated,
indistinctly reddish at the base.
Angola: type and four other specimens from the Chiyaka
district, March 1906 (Dr. F. Creighton Wellman).
The species was met with in thick bush, where antelope
abound, and Dr. Wellman states that it “is very agile and
goes for one’s eyes.” When forwarding these specimens
and examples of a new species of Tabanus, which will shortly
be described, the donor wrote :—‘‘'These are probably the
first insects ever collected in Chiyaka district.” I am glad
to be able to associate with this extremely pretty species the
name of its discoverer, who, although professionally engaged
in another department of science, owing to his energy and
enthusiasm as a collector of Diptera has, during the last
three years, enriched the National Collection with examples
of many interesting new species from Portuguese West
Africa.
By its remarkable coloration C. Wellmanii is easily distin-
guishable from any of its congeners known to me from any
part of the world,
Chrysops Brucei, sp. n.
¢ .—Length (8 specimens) 8 to 93 mom. ; width of head 24
to 3 mm. ; width of front at vertex 3 3 mm. ; length of wing
74 to 9 mm.
514 Mr. E. E. Austen on new
Thick-set, medium-sized, dusky species ; dorsum of thorax
dark brown, with grey longitudinal stripes ; abdomen smoke-
grey or drab-grey, with blackish markings on upper side ; wings
with a brownish tinge, and extreme base, costal border, an
oblique transverse band tapering to a point and not reaching
hind margin, and an apical blotch, intensified round ends of
second longitudinal and upper branch of third longitudinal
veins, dark brown or blackish brown ; legs ochraceous, extreme
tips of all femora and ends of tarsi dark brown.
Head yellowish pollinose, front of same colour or greyer ;
ocellar region usually dark brown, greyer in better-preserved
examples; frontal callus dull black, small, separated from
eye on each side by about one sixth of total width of front
at that point, transverse diameter of callus about two and a
quarter times its height, upper border slightly concave in
middle; a transverse furrow beneath antennz, below which
the central portion of the face is somewhat protuberant ; on
each side of this protuberance a shining brown triangular
area descending to margin of buccal cavity ; a small blackish
triangular fleck on the jowls beneath each eye (not always
distinctly visible) ; front, sides of face, and under side of head
clothed with fine yellowish hair, longer beneath ; palpi of
moderate size, brownish or reddish brown, thinly clothed
with fine yellowish hair ; antenne small, first joint slightly
swollen, not elongate, length of second joint about three
fourths of that of first; first and second jomts blackish
grey above, dull tawny below; third joint blackish brown,
dull ferruginous at extreme base; first and second joints
clothed with blackish hair. Thorax: dorsum with three
dark brown and four much narrower grey longitudinal
stripes, also an extremely narrow, short, dark stripe above
base of wing, running from lower end of transverse suture
almost to postalar callus ; median dark brown stripe about
twice the width of the other two, rounded off about midway
between transverse suture and posterior margin, and thence
to hind margin in well-preserved specimens somewhat
less distinct; paired dark brown stripes interrupted at
transverse suture, and not extending so far forwards as
median stripe ; admedian grey stripes usually with a some-
what yellowish or brownish tinge posteriorly, lateral grey
stripes paler, lavender-grey behind transverse suture ; pleurz
mouse-grey, clothed with yellowish or pale golden hair, of
which there is a tuft below humeral callus, on hind margin
of mesopleura, and on metapleura. Scutellum greyish
brown at base, with broad dull ferruginous hind border.
Abdomen: first segment with a black median blotch beneath
African Species of the Genus Chrysops. 515
the scutellum extending to the hind margin, and on each
side curving outwards so as to form a black hind border to
the segment reaching to the lateral margins; a small
yellowish-grey triangle on the posterior margin, in the
middle line, partially divides the blotch into two halves ;
second and third segments with a pair of large black or
blackish-brown blotches, roughly quadrate in shape, with the
posterior inner angles rounded off, and the outer border (at
least in the case of the blotches on the second segment)
emarginate ; on the second segment the emarginate outer
borders of the blotches run out to the basal angles, thus
forming, with the corresponding markings on the first
segment, an incomplete transverse band ; in the case of the
second segment the hinder edge of the blotches is close to
the hind margin, in that of the third the drab-grey or buff-
coloured hind border is usually a little deeper; fourth
segment with a similar pair of blackish blotches (most
distinct when the insect is viewed obliquely from behind),
more or less rounded off posteriorly, but not emarginate on
the outer side; fifth, sixth, and seventh segments dark
brown, hind borders of fifth and sixth drab-grey, buff, or
ochraceous-buff ; upper surface of abdomen, especially poste-
riorly, clothed with minute golden hairs, lateral margins
of fifth and following segments fringed with longer brownish
hair ; blotches on third segment covered with minute black
hairs; venter mouse-grey, clothed with fine golden hair,
longer than on dorsum and more erect; hind borders of
segments yellowish. Wings: brown area at base including
basal fourth to basal sixth of first and second basal cells, the
tips of which are included in the transverse band; second
costal cell lighter towards the tip; stigma large and well-
marked, dark brown; distal margin of transverse band
beginning on costal border halfway or two thirds of the
distance between end of stigma and level of fork of third
vein, and then running obliquely backwards with two promi-
nences, one on third vein before it bifurecates, the other in
base of second posterior cell; transverse band terminates in
anal cell and does not reach sixth longitudinal vein; discal
cell entirely included in transverse band, but its central
portion as well as base of first submarginal cell usually
lighter; in the apical blotch the intensifications of colour
round the ends of the second longitudinal vein and upper branch
of the third very conspicuous and characteristic; against a
dark background the distal border of the transverse band is
seen to have a lighter edging. Halteresdark brown. Legs:
cox mouse-grey; last three joints of all tarsi dark brown ;
516 Mr. E. E. Austen on new
extreme tips of first and second joints of middle and hind
tarsi brown; front tibize moderately incrassated, middle and
hind tibize not incrassated.
Uganda: type and two other specimens from Kyadondo,
1903 (Colonel David Bruce, C.B., R.A.M.C., F.R.S.); other
specimens from Singo, 1903 (Col. Bruce), Busoga, March
1906 (Dr. A. D. P. Hodges), and the Nile between Wadelai
and Nimule, 1906 (the late Dr. W. A. Densham). With
reference to the specimen obtained by him, Dr. Densham
wrote :—“ Only specimen seen ; taken at an opening in the
swampy river edge.”’ I have pleasure in naming this species
in honour of the distinguished investigator, whose epoch-
making discovery of the cause of nagana (tsetse-fly disease of
animals) laid the foundation of all subsequent work upon
the «tiology and treatment of sleeping sickness and the
various forms of animal trypanosomiasis.
Chrysops Brucei is not closely allied to any other African
representative of its genus hitherto described. In its abdo-
minal markings it presents a certain, though not by any
means a close, resemblance to the following species ; but the
wing-markings (flecks at tips of second and upper branch of
third vein) are alone sufficient to distinguish it from all other
African species of Chrysops at present known.
Chrysops inconspicua, sp. 0.
? .—Length (3 specimens) 5 to 74 mm.; width of head 2
to 24 mm.; width of front at vertex # mm. ; length of wing
5 to 65 mm.
Small species, with rather broadly oval abdomen, the dorsum
of which is marked with conspicuous paired black spots or
transverse blotches, meeting in the middle line in front and
separated by triangular interspaces ; wings hyaline, with base,
costal border to end of second longitudinal vein, and an oblique
transverse band not extending beyond anal cell blackish brown ;
anal cell infuscated, a clear space in first and second basal
cells near their distal extremities ; legs entirely black.
Head.—Front olive-grey, jowls and sides of face chrome-
yellow pollinose; frontal callus very large, shining black,
shaped like an equilateral triangle, the apex including the
anterior ocellus; vertex with a pair of small, transversely
elongate, shining black flecks, one on each side of median
line and including one of the other two ocelli; median
portion of lower part of face with a single large, somewhat
cordate, shining black tubercle, extending to margin of buccal
cavity ; no black fleck on jowls; front, sides of face, and
African Species of the Genus Chrysops. 517
under surface of head thinly clothed with rather long and
fine yellowish hair, longer below ; palpi blackish, slender or
of medium thickness, clothed with minute yellowish hairs
antenneé not elongated, first joint more or less inieradeatedl
second joint two thirds of length of first; first and second
joints dark mouse-grey, somewhat shining, clothed with
rough-looking yellowish-brown hair ; last jomt dull brownish
black. Thorax: dorsum with a pair of well-defined, broad,
shining, blackish-brown longitudinal stripes, rounded off at
each end, but not inter rupted on transverse suture ; these
stripes separated by a broad median mouse-grey stripe, darker
from front margin to a little beyond transverse suture, the
darker portion with a hght grey edging ; pleurz yellowish
poliinose (lower portion ‘of mesopleura blackish), with thick
tufts of cadmium-yellow hair; dorsum clothed with fine,
erect, pale yellow hair; scutelium dark mouse-grey, clothed
with long, thin, yellowish hair. Addomen: ground-colour
of dorsum olive-grey, sides of first and second segments
sometimes ochraceous ; first segment with a blackish-brown
median blotch, extending well beyond scutellum on each
side; paired blotches on the four following segments roughly
triangular in shape, rounded off behind, and not extending
to hind margins, but sometimes transversely elongate, being
produced so as to include basal angles of seements ; sixth
and seventh segments mouse-grey ; dorsum (except on black
blotches, which are covered with minute, appressed, blackish
hairs) clothed with short chrome-yellow hair, most noticeable
on hind margins of segments; sides of segments, from first
to fifth inclusive, fringed with bright cadmium-yellow hair ;
venter yellowish grey, clothed with cadmium-yellow hair,
basal portion of some of the segments sometimes with dark
brown transverse bands. Wings: basal brown area in first
basal cell includes the proximal two thirds ; in second basal
cell it may be confined to proximal third, or, if extending as
far as in first basal cell, may be more or less obliterated
posteriorly ; proximal margin of transverse band running
perfectly straight from base of third vein to inner basal angle
of fifth posterior cell, thus filling out the tips of the first and
second basal cells with colour ; ‘distal margin of transverse
band commencing at end of second longitudinal vein and
running obliquely backwards so as to include basal half of
fifth posterior cell; distal margin of transverse band with
three indentations, the first in first submarginal cell, above
fork of third vein, the second in first posterior cell, and the
third on or near vein separating fourth and fifth posterior
cells ; the transverse band dies away in the anal cell; trans-
518 Mr. E. E. Austen on new
verse band sometimes with lighter areas, e. g. in discal cell
and at base of fourth and fifth posterior cells ; portion of
fifth vein bounding second basal cell below more or less
suffused with brown; no apical blotch. MHalteres dark
brown. Legs: none of the tibiz incrassated.
Angola (Ochilonda, Bihé): three specimens, March 7th-
9th, 1905 (Dr. F. Creighton Wellmann). The donor’s field-
note on this species is as follows :—‘ Taken March 7th, 8th,
and 9th in the edges of bush bordering on small wet plains.
Natives call it the ‘antelope fly, and state that it bites
principally the duyker-bok (Cephalophus Grimmii).”
The slight resemblance between this species and the fore-
going as regards abdominal markings has already been noted ;
the wing-markings, however, at once distinguish it, and it
cannot possibly be confused with any other African Chrysops
as yet described. Since neither of the three specimens
available is in faultless condition, no special type has been
selected ; but the foregoing description has been drawn up
from all three, which are therefore co-types.
Chrysops nigriflava, sp. 0.
? Length (3 specimens) 74 to 8? mm.; width of head
22 to 24 mm. ; width of front at vertex 3 mm.; length of
wing 8 to 8} mm.
Cadmium-yellow, with conspicuous black longitudinal stripes;
dorsum of thorax with three black stripes, middle one extending
on to scutellum ; dorsum of abdomen with two very broad
black stripes, enclosing a narrower median stripe of ground-
colour ; base, costal border, and distal half of wings, from
base of third vein, blackish brown, an ill-defined lighter area
along posterior margin, commencing below upper branch of third
vein and terminating in fifth posterior cell; anal cell (in all
three specimens available) closed before reaching margin of
wing ; legs black.
Head cadmium-yellow pollinose; frontal callus shining
black, rather narrow in the antero-posterior direction, and
on each side ending on a level with outer margin of base of
antenna, upper edge somewhat curved, ends rounded off,
lower edge straighter; vertex marked with a dark brown
nearly circular spot, which includes the ocelli; median tumid
area of lower part of face with shining dark brown triangle
on each side, apices of triangle reaching margin of buccal
cavity ; no dark spot on jowls; sides of face, jowls, and
under side of head clothed with cadmium-yellow hair; palpi
large, lanceolate in shape when viewed from outer side,
African Species of the Genus Chrysops. 519
blackish brown and clothed with brownish hair; antenne
(third joint missing) black, not slender, first joint slightly
incrassate and also somewhat elongate, length of second jomt
two thirds of that of first joint ; hair clothing first and second
joints black. Thoraz: black stripes on dorsum at least
twice as broad as the pair of yellow stripes separating them,
which are somewhat paler than elsewhere ; a dark brown
stripe on side of thorax ona level with base of wing, extending
from front margin to postalar callus; beneath this stripe in
well-preserved specimens there is a thick fringe of cadmium-
yellow hair, and the lateral yellow stripe external to the
outer dorsal black stripe bears similar hair; lower part of
mesopleura and pectus blackish brown. Scufel/um yellow on
sides and posteriorly ; end of median thoracic stripe forming
a dark shield-shaped median spot, not extending to hind
margin; yellow area on scutellum and admedian yellow thoracic
stripes apparently pollinose, the ground-colour being black.
Abdomen: black dorsal stripes starting from a common
base on front margin of first segment beneath scutellum,
dividing just before reaching hind margin of this segment,
and extending without interruption to the seventh segment ;
they approach one another on the sixth segment and almost
or quite come into contact on the seventh ; on the third and
following segments the stripes widen out so as to reach the
lateral margins ; venter, except a scarcely visible black or
blackish median area at extreme base on first segment,
entirely cadmium-yellow without markings, clothed with
short, appressed, similarly coloured hair; median dorsal
stripe and sides of first and second segments clothed with
similar hair. Wings: transverse band extending to hind
margin (though lighter posteriorly), and so broad that wing
may be described as blackish brown with exception of a lar ge,
semihy aline, triangular area at the base ; the upper margin of
this light area, w hich is somewhat blurred, starts at the base
of the anal cell and runs obliquely forward into the marginal
cell a little before the base of the third longitudinal vein ;
the anterior side of the light area is formed by the proximal
margin of the transverse hand, which runs at right angles to
the costa, from the base of the third vein straight down into
the anal cell ; the alula, axillary cell, and apical portion of
anal cell are all infuscated ; lighter area in distal half of
wing widest in first submarginal cell (7. e. below what in the
case of other species would be the apical blotch), and thence
progressively diminishing in width. Halteres dark brown.
Legs entirely black and clothed with short black hair, none
of the tibize incrassated.
520 On new Species of Plecotus.
German East Africa: type and two other specimens from
Neguelo, Usambara (purchased from Hermann Rolle).
The striking markings and coloration of this species are
sufficient to distinguish it without difficulty from any other
African Chrysops at present known.
SYNONYMICAL AND OTHER NOTES.
Chrysops tarsalis, Walk., = C. longicornis, Macq., as
suggested by Gerstaecker (Baron Carl Claus von der Decken’s
‘Reisen in Ost-Afrika,’ Bd. u. Abth. 3, 1873, p. 384).
Miss Ricardo (Ann. & Mag. Nat. Hist. ser. 7, vol. ix. 1902,
p. 368) is mistaken in regarding the two species as distinct.
Chrysops trimaculatus, Bigot, = C. longicornis, Macq.
Through the courtesy of Mr. G. H. Verrall, in whose collec-
tion it now is, I have been enabled to examine Bigot’s type,
which is a ¢, not a 9, as stated by the author and by
Miss Ricardo (loc. cit. p. 871).
Chrysops fuscus, Ricardo (loc. cit. p. 368).—The type and
remainder of the original series of this species are males, not
females, as stated by the author.
LXXV.—Deseriptions of Two new Species of Plecotus.
By G. E. H. Barretr-HAmMILTON.
Amoncst the bats in the British Museum of Natural History
I find two undescribed species of the ‘‘ Long-ear.”” They
may be characterized as follows :—
2 ° 09 fp
Plecotus tenerife, sp. n.
This bat resembles P. aurttus of Britain, but has much
larger wings. The colour is apparently darker, but cannot
be taken as reliable from the old dried skin upon which this
description is based: it is, above, near Ridgway’s “ hair-
brown,” the hair-tips lighter; below, dirty yellowish white,
the dark basal portions of the hairs not evident.
The type has no skull.
The dimensions (in millimetres) of the type are as
follows :—
Head and body 52; tail 45; ear from the notch 35;
tragus 14; thumb without claw 6; longest digit 73; basal
joint of fifth digit 35 ; basal joint of second digit 37; fore-
arm 44; tibia 18°5 ; hind foot (without claw) 8.
Hab. Teneriffe.
On a new Species of Monkey. 521
The type,'a dried skin, is No. 87. 4.18.1 of the British
Museum collection, and was taken at Orotava, Teneriffe, by
Sei. R. Gomez on the 3rd of April, 1887.
The large wings of this bat mark it as distinct from any
other known Plecotus.
Plecotus puck, sp. n.
This bat resembles P. auritus of Britain and is of similar
size, but has a quite distinct skull.
The colour of the basal portions of the hairs is everywhere
dusky: the tips are, above between Ridgway’s “ isabella
colour ” and “ broccoli-brown,” below whitish ; the upperside
has a grizzled appearance.
The skull, as compared with Central European examples
(? austriacus of Geoffroy), is smaller, but about equal in size
to those from England. The facial region and palate are,
however, narrower and the backward extension of the latter
less pronounced ; the auditory bulle are larger.
The dimensions of the type (in millimetres) are as
follows :—
Head and body 40 *; tail 50*; ear 41*; ear from the
notch 32; tragus 16; thumb without claw 8; longest digit
60; basal joint of fifth digit 32 ; basal joint of second digit
34°5 ; forearm 38; tibia 17; hind foot without claw 8.
The type is a skin, No. 5.11.19. 1 of the British Museum
collection. Itis a male taken at Murree, N. India, altitude
7500 feet, by Capt. E. T. F. Birrell, R.A.M.C., on the 20th
of August, 1905.
This bat approaches more closely to European Plecot?
than to others in the Museum from Tor, Sinai, and from
Ladak. ‘he latter are in both cases larger, in which
character they agree with two from Hokkaido, Japan. They
are, perhaps, referable to Hodgson’s P. homochrous.
LXXVI.—Description of a new Species of Monkey (Cerco-
pithecus Hamlyni) from the Ituri Forest. By R. I.
Pocock, F.L.S., Superintendent of the Zoological Society’s
Gardens.
Cercopithecus Hamlyni, sp. n.
Skin of the face a purplish-brown hue, with a narrow
whitish flesh-coloured line extending from the brow between
the eyes and nostrils on to the upper lip. Ears the same
colour as the face, scantily clothed with inconspicuous
* Collector’s measurements from label.
Ann. & Mag. N. Hist. Ser. 7. Vol. xx. 34
522 On a new Flying-Squirrel from Formosa.
speckled hairs, Head with a rather inconspicuous short and
narrow pale brow-band; the rest of the head, the cheeks,
and the dorsal and lateral surfaces of the body uniformly
dark-coloured, the hairs being finely speckled with brownish
yellow. No white on the throat or chest, the underside
bemg for the most part of an olive-grey or brownish-grey
hue. ‘The arms and legs speckled to the wrists and ankles,
but darker than the body, especially upon the upper arm and
thigh, the speckling being less apparent than on the body ;
hands and feet for the most part black above. ‘Tail the same
colour above as below, speckled and tinted like the body in
the basal two thirds of its length, with its distal third black.
The whiskers are full and directed obliquely backwards
and downwards, and the coat is fairly thick and long.
Loc. The Ituri Forest.
A single living specimen belonging to the Hon. Walter
Rothschild and proeured from Mr. J. D. Hamlyn, after
whom the species is named.
This is a very distinct species. By the analytical key to
the groups of species of the genus Cercopithecus published
in my recent monograph (P. Z. S. 1907, p. 681) of these
monkeys it falls under heading ‘‘7’””—that is to say, with
the species of the Leucampyx- and Albogularis-groups ; but
it has not the black fore limbs and white throat of either
section, and further differs from both in the uniformity in
tint and speckling between the head and body, in lacking the
black of the head and nape characteristic of the Leucampye-
eroup and the brighter red or yellow hue of the dorsal area of
the body of the Albogularis-group. Finally, the peculiar
white line down the centre of the face is quite unique.
LXXVIL—A new Flying-Squirrel from Formosa.
By OLDFIELD ‘THOMAS.
Petaurista lena, sp. n.
A beautiful rufous species with a pure white head and
under surface.
General colour above a brilliant chestnut-rufous, much as
in P. grandis, but less dulled by black. This colour extends
over the upperside of the neck from the occiput, the back, and
the whole of the upperside of the parachute down to the
elbows and ankles. Under surface and inner side of limbs
pure white throughout or slightly tinged with yellowish.
Head also pure white above and below, bounded abruptly
behind by the rich rufous of the nape; a few rufous hairs
forming a narrow rim round the eyes. Hars long, oval,
On Two African Freshwater Sponges. 528
mostly naked, the hairs on their bases behind rufous, passing
into a darker postauricular spot, which may be more or less
mixed with blackish rufous. Hands white, forearms and
edges of the patagial cartilage mixed rufous and white, as are
also the edges of the parachute, the proportions of the rufous
and white hairs varying in the two specimens. Hind feet
dark rufous proximally, more or less grizzled with white
terminally. Tail mixed black and rufous, the tip with a
black or black-and-white tuft.
Skull, as compared with that of P. grandis, readily distin-
guishable by its much longer and narrower nasals, which are
far less expanded anteriorly; the nasal opening is conse-
quently much narrower, while of about the same height.
Dimensions of the type (measured in skin) :—
Head and body 350 mm.*; tail 440; hind foot (wet) 76 ;
ear (wet) 40.
Skull: upper length to hinder edge of parietal 68; greatest
breadth 47°5; nasals 23°5 x 12°5 ; interorbital breadth 17:2 ;
palatilar length 32; length of upper tooth-row exclusive of
p 16.
Hab. Tapposha, Central Formosa.
Type. Adult female. Collected 18th February, 1907, by
native hunters for Mr. Alan Owston. Original number 60.
Two specimens.
- This striking flying-squirrel is one of the most beautiful
species ever described, owing to the wonderful contrast
between the rich red body and the pure white head and
underparts. The two specimens are practically the same
throughout, and the skull-difference from P. grandis, also a
native of Formosa, shows that P. dena is not a mere colour-
phase of that animal.
LXXVIII.—WNotes on Two Species of African Freshwater
Sponges. By Kh. KIRKPATRICK.
Mr. J. Stuart THomson has sent to the Natural History
Museum several small specimens of freshwater sponges which
he collected from a pond at Valkenberg Vlei, near Cape
Town. The specimens, which are in the form of crusts on
the stems of rushes, belong, in my opinion, to a new variety
of Ephydatia fluviatilis, Linn, This almost cosmopolitan
species has been found in Europe, Asia, and America, but, I
believe, is now recorded for the first time from Africa.
A specimen of a second species, viz. of Spongilla cerebellata,
Bowerbank, from a pond near Cairo, has been presented to
the Museum by Dr. Innes Bey, through Mr. C. Boulenger.
* Probably shrunk ; the other specimen is 480 mm. in length,
524 On Two African Freshwater Sponges.
S. cerebellata has been recorded from Central India. If
Dr. Annandale is right in regarding this species as a form
of S. lacustris, then the two commonest Huropean species,
E. fluviatilis and S. lacustris, have to be added to the list of
African freshwater sponges, of which twenty-one species are
now known.
A description of the new variety of EF. fluviatil’s is given
below.
wo
Fig. 1.—£phydatia fluviatilis, var. capensis, on reed, nat. size.
Fig. 2—Gemmule.
Fig. 3.—Oxea.
Figs. 4, 5.—Smooth and spined tornotes.
Figs. 6, 7.—Amphidisks.
Fig. 8 —End view of same.
Fig. 9.—Oxea of #. fluviatilis from Thames, London.
Figs. 10, 11.—Amphidisk from Thames specimen, side and end view.
Ephydatia fluviatilis, Linn., var. capensis, nov.
Sponge encrusting ; colour, alive or in alcohol, of a medium
brown; surface in part smooth and uniform, in part pitted
and finely hirsute. Oscules scattered, level with surface,
inconspicuous, about 1 to 15 mm. in diameter. Flagellated
chambers small, oval, 20x 16 yw.
Gemmules large, oval, °75 mm. in length by ‘54 mm. in
breadth, with funnel-shaped tube opening flush with the
Bibliographical Notices. 525
surface ; with thick (10 w) chitinous inner shell and a single
layer of amphidisks embedded in a vesicular layer.
Spicules : slender oxeas (fig. 3) 288 x 8 yw, curved, attenu-
ating gradually, smooth or finely and sparsely spined; tornote
oxeas (figs. 4, 5) 240 x 12 yw, smooth or finely spined.
Gemmule spicules: amphidisks (figs. 6-8) 35°6 p long,
shaft 4°3 mw thick, smooth or with one or several spines ;
diameter of deeply incised disks 24°34; with a knob at
centre of disk.
Numerous developmental forms scattered in the tissues of
the sponge.
Hab. Valkenberg Vlei, near Cape Town (J. Stuart
Thomson).
There are seven specimens of this sponge, all encrusting,
the largest being 5 cm. long and *5 cm. thick. The chief
point of interest lies in a comparison with the typical Euro-
pean form. In the latter the subtornote oxeas (fig. 9) are
considerably longer and thicker, viz. 320 x 20 w; the amphi-
disks (figs. 10, 11) are much shorter, viz. 18°5 w, with disks
21-4 in diameter and shafts 4°5 thick ; lastly, the gemmules
are smaller, on an average about °36x°31 mm. These
differences led me at first to regard the Cape specimens as
representatives of a new species, but I ultimately came to
regard them as belonging to a new strongly marked variety
of Ephydatia fluviatilis.
BIBLIOGRAPHICAL NOTICES.
Memoirs of the Department of Agriculture in India (Entomological
Series). 4to. Agricultural Research Institute, Pusa. Printed
by Thacker, Spink, & Co., Calcutta.
A usEFuL periodical, recently commenced under the editorship of
the energetic Imperial Entomologist, Prof. H. Maxwell-Letroy,
F.E.S., F.Z.S. The following parts have already apppeared :—
Vol. I. No. I. The Bombay Locust.
II. The more important Insects injurious to Indian
Agriculture.
Both by Prof. Maxwell-Lefroy.
III. The Indian Surface Caterpillars of the Genus
Agrotis. By Prof. Maxwell-Lefroy and his
Assistant, C. C. Ghosh, B.A.
IV. Individual and Seasonal Variations in Helopeltis
theivora, Waterhouse, with Description of a new
Species of Helopeltis. By Harold H. Mann,
D.Sc., Scientific Officer, Indian Tea Association.
The following is announced as in the press :—
V. The Coccide attacking the Tea-plant in India and
Ceylon. By E. EK. Green, F.E.S., and Harold
H. Mann, D.Sc.
526 Bibliographical Notices.
Homoptera Andina: Die Zikaden des Kordillerengebictes von Sud-
amerika nach Systematik und Verbreitung, von A. Jaconi. —
I. Cicadide. (Abhandlungen und Berichte des Konigl. Zoologischen
und Anthropologisch-Ethnographischen Museums zu Dresden.)
Band xi. (5). 1907. 4to. Pp. 28:
Tur present work offers us a synopsis of the Cicadidee of all the
Pacific States of South America, from the southern limits of the
fauna included in the ‘ Fauna Boreali-Americana’ to Chile in the
south, and to Venezuela and the Upper Amazon districts on the
east.
Seventy-two species are enumerated in the present work, of
which no less than forty-four are figured (occasionally with additional
details) on the large double plate accompanying the paper. Hight
species are represented in colour, the rest being plain, and seven
species are described as new. New species, or those which have
been insufficiently described, are noticed at length, but the others
often very briefly. On p. 5a useful diagram of neuration &c. is
given according to the system of Stal. Limited faunistic works
like the present, especially when well illustrated, are of great value
in extending our knowledge of the details of a subject so vast as
Entomology. Wabi
Précis des Caractéres génériques des Insectes, disposes dans un ordre
naturel par le Citoyen Latreille. A Paris, chez Prévot, Libraire,
Quai des Augustins, et & Brive, chez F. Bourdeaux, Imprimeur
Libraire. A Brive, de l’Imprimerie des F. Bourdeaux, au 5 de
la R. [1796]. Pp. xiv, 208. Imprimé 4 200 Exemplaires pour
A. Hermann. mbccccvil.
We alluded recently to a work published on the early life of the
great French entomologist Latreille, who stood in a somewhat
similar relationship to Cuvier as that occupied by Fabricius towards
Linné. Latreille was born in 1762, and died in 1833. After his
almost miraculous escape from death during the early days of the
Revolution he devoted himself to entomology with great ardour,
74 entries (from 1792 onwards) figuring in Hagen’s ‘ bibliotheca
Entomologica’ under his name. The work before us is a reprint of
one of the rarest of entomological books, of which only a few copies
are known to be in existence; and it is also the earliest of the
important series of books on systematic entomology which made
Latreille’s name famous. It includes the characters in French of
all the genera of Insects, Arachnida, Crustacea, and Myriopoda,
divided into 14 Orders. No species are mentioned, and no types
are assigned even to new genera; but this omission was fully
rectified by the publication (1802-1805) of the largest and most
valuable of Latreille’s works, his great ‘ Histoire naturelle, générale
et particuliére, des Crustacés et des Insectes,’ in 14 volumes. We
are glad that the numerous entomological libraries which do not
possess the ‘ Précis’ will now be able to place at least the reprint on
their shelves.
Or
INDEX to
~T
VOLE XX
ABARATHA, new species of, 432.
Abricta, new species of, 415.
Acanthoglossus, new species of, 293,
498.
Allonotus,
eenus, 29,
Amphilius, new species of, 488.
Amphipods, on some freshwater,
227.
Ancistrocerus, new species of, 85.
Andersen, K., on Pteropus masca-
rinus, 351; on the geographical
races of Rhinolophus hipposiderus,
384.
Andrena, new species of, 128.
Andrews, Dr. C. W., on the cervical
vertebree of Zeuglodon from the
Barton clay, 374.
Anomalon, new species of, 14.
Aotus, new species of, 189,
Arachnida, new, 53, 297.
Archeeocidaris, on the generic name,
452.
Arnetta, new species of, 433.
Apes, on the hand and foot in man
and, 155.
Aphadnus, characters of the new
genus, 26,
Apobletes, new species of, 97.
Apona, new species of, 75.
Atholus, new species of, 346.
Atractylis coccimea, remarks on,
52.
Aurivillius, new species of, 4.
Austen, E. E., on new African
species of Chrysops, 507.
Automolis, new species of, 506,
Beeturia, new species of, 417.
characters of the new
Barbus, new species of, 336, 486,
488.
Barrett-Hamilton, G. E. H., on two
new species of Plecotus, 520,
Bather, F. A., Echinocrinus versus
Archeocidaris, 452.
Bathyonyx, characters of the new
genus, 227,
Batrachians, new, 46, 47, 109.
Birds, on the trematode parasites of
British, 245,
Bolton, H., on a marine fauna in the
Bristol coal-field, 575.
Books, new :—Books and Portraits
illustrating the History of Plant
Classification, 80; Bernard’s Cata-
logue of the Madreporarian Corals
in the British Museum, 168 ;
Reid’s I Go A-Walking, 295 ;
Nature in School, 295; English’s
100 Photographs from Life, 372 ;
Lodge’s 100 Photographs from
Life, 372; de Nussac’s Les Débuts
dun Savant Naturaliste, Pierre
André Latreille, 372; Bordage’s
Recherches anatomiques et biolo-
giques sur l’Autotomie et la Ré-
eénération chez divers Arthro-
podes, 373; The Fauna of British
India, Butterflies, vol. ii., 373;
Memoirs of the Department of
Agriculture in India, 525; Jacobi’s
Homoptera Andina, 526; Précis
des Caractéres génériques des
Insectes, 526.
Boulenger, G. A., remarks on Prof.
L. von Méhely’s paper “ Zur
Lisung der Muralis-Frage, 39;
528
on a new engystomatid frog, 46;
on a new toad and a new amphis-
beenid from Mashonaland, 47; on
a new cichlid fish, 50; on new
freshwater fishes from S. Came-
roon, 50; on new freshwater fishes
from Angola, 108; on a new frog
from Angola, 109; on new African
species of Barbus, 336; on a new
pit-viper from Brazil, 338 ; on new
freshwater fishes from S. Came-
roon, 485; on new fishes from
Central Africa, 487; on a new
African snake, 489.
Brachiopod nomenclature, notes on,
223; morphology, 376.
Breviceps, new species of, 46.
Browne, E. T., on the medusze be-
longing to the family Laodiceide,
457.
Buckman, S.8., notes on brachiopod
nomenclature, 223: on brachiopod
morphology, 376.
Bufo, new species of, 48.
Bulimulus, new species of, 491.
Burbunga, new species of, 416.
Cabrera, A., on three new Spanish
insectivores, 212.
Callicebus, new species of, 161,
191.
Caltoris, new species of, 434.
Cameron, P., on the hymenoptera
of the oriental zoological region,
OSs
Castnia, new species of, 505.
Casyapa, new species of, 430.
Cebus, new species of, 292.
Celzenorhinus, new species of, 431.
Cerceris, new species of, 85.
Cercidochela, characters of the new
genus, 284.
Cercopithecus,
521.
Chionzema, new species of, 76.
Chirindia, characters of the new
genus, 48,
Chrysops, new species of, 507.
Cicada, new species of, 418.
Cladomelea, new species of, 36.
Clarias, new species of, 51.
Clausilia, new species of, 495.
Cockerell, T. D. A., descriptions and
records of bees, 59, 122, 445.
Coleoptera, new, 95, 339, 423,
481.
Colganta, new species of, 26.
new species of,
INDEX.
Colopus, new species of, 195.
Copepods from the S.W. coast of
Ireland, on, 377.
Cornillus, characters of the new
genus, 99.
Crabro, new species of, 88.
Crangonyx, on the sensory capsules
in species of, 231.
Cratocryptus, new species of, 24.
Crocidura, new species of, 197, 215,
390.
Crossomys, characters of the new
genus, 70.
Crustacea, new, 227, 365, 377; on
the, of the Channel] Islands, 556.
Cryptocotyle concavum, remarks on,
261.
Ctenocharax, characters of the new
genus, 402.
Ctenomys, new species of, 164.
Cyrtogone, new species of, 8.
Dasypus sexcinctus, on the local
forms of, 165.
Desmacidon, new species of, 282.
Diacrisia, new species of, 75.
Diaphractus, characters of the new
genus, 315.
Diplogrammicus, characters of the
new genus, 481.
Diptera, new, 507.
Distant, W. L., rhynchotal notes,
411; ona new genus and species
of Cerambycidee, 423; on a new
species of Cicadide from E. Africa,
425.
Drasside, new 8. African, 297.
Drassodes, new species of, 305,
Drepana, new species of, 77.
Druce, H., on new heterocera, 505.
Druce, H. H., on a new lyczenid from
Mauritius, 219.
Dryomys, new subspecies of, 407.
, definition of the new generic
name, 406.
Kchinocrinus, on the generic name,
452,
Echthrus, new species of, 19.
Eleotris, new species of, 52.
Elliot, Dr. G., on new species and
subspecies of mammals, 185; on a
new species of Cebus, 292; on a
new baboon from British E. Africa,
499,
Epeolus, new species of, 60.
Ephydatia fluviatilis, new variety of,
524,
INDEX.
Epiphora, new species of, 1.
Epirhyssa, new species of, 16.
Kpitoxus, new species of, 347.
Eretmotus, new species of, 106.
Escharina Dutertrei, remarks on,
210.
Eugrammicus, characters of the new
genus, 99.
Euproctis, new species of, 78.
Eutidium, new species of, 540,
Exosternus, new species of, 104.
Felis, new species of, 396.
leo and F. concolor, on the
significance of the pattern of the
cubs of, 436.
Fishes, new, 50, 93, 108, 336, 402,
485, 487.
Fitzwilliams, Dr. D. C. L., on fune-
tion and form with reference to
the hand and foot in man and
apes, 155.
Fovaya, new species of, 21.
Friona, new species of, 25.
Gahania, characters of the new
genus, 425,
Galago, new species of, 186.
Gammarid, on the reduction of the
eye in an Irish, 227.
Gellius, new species of, 286.
Genetyllis, new species of, 175.
Geological Society, proceedings of
the, 374.
Godman, Dr. F. D., on the American
species of Hesperiidze described by
Plotz, 152.
Godwin-Austen, Lt.-Col. H. H., on
Helix hyba, 55.
Gonimbrasia, new species of, 4,
Grammostethus, new species of,
345.
Grison, new species of, 162,
Gronoceras, characters of the new
genus, 65,
Gunomys, characters of the new
genus, 203. ;
Gymnophallus, new species of, 263.
Haliphera, new species of, 81.
Hasora, new species of, 435,
Helicocranchia, characters of the
new genus, 382,
Helix hyba, observations on, 55.
Henslow, Prof. G., on the xerophytic
character of coal-plants, 375.
Herpestes gracilis, on the African
mungooses of the group of, 110.
Hesperiide, new, 430.
529
Hesperiidz, on the American, de-
scribed by Plotz, 132.
Heteroscodra, new species of, 36.
Hirst, A. 8., on new African spiders
and Solifuge, 33; on new pill-
millipedes from the Malay Penin-
sula and Siam, 215,
Tlister, new species of, 101, 544,
485.
Histeridee, new, 95, 339, 480.
Histizea, new species of, 506.
Holocera, new species of, 9.
Tlomalopygus, new species of, 349.
Hoplakithara, characters of the new
genus, 285.
Hylobates agilis, on the hand and
foot of, 155.
Hymedesmia, new species of, 273.
Hymenoptera, new, 10, 59, 81, 122,
445,
Tymerrhaphia, new species of, 274.
Hyposolenus, characters of the new
genus, 97.
Tluza, new species of, 79.
Imbrasia, new species of, 2.
Imeria, new species of, 28.
Iophon, new species of, 276.
Iphiaulax, new species of, 10.
Isepeolus, characters of the new
genus, 64.
Ismene, new species of, 434.
Jameson, H. L., on a new hare from
the Transvaal, 404.
Joyeuxia, new species of, 283.
Kirkpatrick, R., on the Monaxo-
nellida of the National Antarctic
Expedition, 271; on two African
freshwater sponges, 525,
Labeo, new species of, 108.
Lacerta muralis, remarks on, 39.
Lachesis, new species of, 338,
Lagothrix, new species of, 193.
Laodice Maasii, definition of the new
specific name, 466,
Laodiceide, revision of the, 457.
Lepidoptera, new, 1, 75, 219, 430,
505,
Leptolepis, new species of, 93,
Lepus granatensis, new forms of,
398,
Leucoma, new species of, 78.
Lewis, G., on new Histeride, 95,
339, 480.
Limnea, new species of, 496,
Lissomyxilla, new species of, 275,
Lobobunvea, new species of, 5.
Ann. & Mag. N. Hist. Ser. 7. Vol. xs 35
2
5
Ludia, new species of, 9.
Lydekker, R., on the name of the
Armenian wild sheep, 121.
Lynx pardella, definition of the new
specific name, 398.
M‘Intosh, Prof., ona porbeagle shark
with a large suboral aperture,
169; on the young of the iing,
171; ona large example of Omma-
strephes sagittatus, 172 ; on Gene-
tyllis citrina, 175; on the repro-
duction of Nereis diversicolor, 176.
Macrotristria, new species of, 412.
Mammals, new, 31, 70, 113, 161,
185, 196, 202, 212, 221, 292, 293,
390, 398, 404, 406, 426, 498, 499,
500, 520, 521, 522.
Man and apes, on the hand and foot
in, 155.
Mardalana, new species of, 418.
Maritrema, characters of the new
genus, 265,
Marmosa, new species of, 167.
Mason, G. E., on an extinct fruit-bat
of the genus Pteropus, 220.
Massy, A. L., on copepods from the
S.W. coast of Ireland, 377.
Mastacembelus, new species of,
487.
Meduse belonging to the family
Laodiceidee, revision of the, 457.
Megachile, new species of, 68.
Melampsalta, new species of, 419.
Melanocera, new species of, 5.
Melanophora, new species of, 327,
Meles, new species of, 394.
Mesostenoideus, new species of, 22.
Metopa, new species of, 365.
Microlister, new species of, 102.
Microtus, new subspecies of, 201,
Miller, G. S., on new European in-
sectivora and carniyora, 389; on
two new forms of the Spanish hare,
398; on new European squirrels,
426, .
Mimagoniates, characters of the new
genus, 402.
Miniopterus,
197,
Mollusca, new, 490.
Molva molva, on the young of,
Wal
Monaxonellida of the National Ant-
arctic Expedition, on the, 271.
Monoplius, new species of, 103.
Mungos, new subspecies of, 113.
new subspecies of,
30 INDEX.
Murexus, characters of the
genus, 350,
Mus, new species of, 206, 501.
Mycale, new species of, 280.
Myxilla, new species of, 278.
Nacaduba, new species of, 219.
Nannocharax, new species of, 485,
Neomys, new species of, 214.
Nereis diversicolor, on the reproduc-
tion of, 176.
Nesokia, new species of, 199, 205;
subdivision of the old genus,
202.
Nicoll, W., on the trematode para-
sites of British birds, 245.
Niphargus, on the sensory capsules
In species of, 280.
Norman, Canon A. M., on some
British polyzoa, 207 ; on the crus-
tacea of the Channel Islands, 356.
Notocrypta, new species of, 454.
Oceanapia, new species of, 289.
Octopus with branching arms, note
on an, 407.
Odynerus, new species of, 82.
Ommastrephes sagittatus, note on a
large example of, 172.
Onthophilus, new species of, 107,
Ophlitaspongia, new species of, 274.
Ophthalmodes, new species of, 80.
Osmia, new species of, 122, 446.
Otomys, new species of, 31.
Ovis orientalis, note on, 121.
Pachycréerus, new species of, 548.
Palarus, new species of, 91.
Papio, new species of, 194, 499.
Paratilapia, new species of, 108.
Paratropus, new species of, 349.
Parusta, characters of the new
genus, 6.
Pauropsalta, new species of, 422.
Pedetes, new species of, 32.
Pedoptila, new species of, 506.
Perdita, new species of, 129.
Petaurista, new species of, 522.
Petersius, new species of, 487.
Petrosia, new species of, 290.
Phalanger, new species of, 74.
Phylactella pygmea, remarks on,
210.
Phyllodocid, on a new, 175.
Physa, new species of, 497.
Pill-millipedes, new, 215.
Planorbis, new species of, 496.
Platyoides, new species of, 297,
Platysoma, new species of, 98.
new
INDEX.
Plecotus, new species of, 520.
Pleurodonte, new species of, 490.
Pocock, R. I., on the significance of
the pattern of the cubs of lions
and of pumas, 436; on a new
species of monkey, 521.
Polita, new species of, 490,
Polymastia, new species of, 271.
Polypus, new species of, 377.
cephea, note on a specimen of,
407.
Polyzoa, notes on British, 207.
Porbeagle shark with a large suboral
aperture, note on a, 169.
Preston, H. B., on new land and
freshwater shells from Central and
South America, 490.
Probolosternus, new species of, 104.
Pronolagus, new subspecies of, 404.
Psen, new species of, 89.
Psilostomum brevicolle, remarks on,
269.
Pterinochilus, new species of, 33.
Pteropus, on an extinct fruit-bat of
the genus, 220, 351.
Purcell, Dr. W. F., onnewS. African
Drasside, 297.
Putorius, new subspecies of, 395.
Pycna, new species of, 425.
Raia undulata, note on, 403.
Rana, new species of, 109,
Regan, C. T., on two new characinid
fishes from 8. America, 402; on
Raia undulata, 405.
Reithrodontomys, new species of,
163,
Reniera, new species of, 291.
teninus, new species of, 105.
Reptiles, new, 48, 338, 489.
Rhinolophus hipposiderus, on the
geographical races of, 384.
Rhynchota, new, 411, 425.
Rhyssa, new species of, 15.
Rothschild, Hon. L. W., on new
African Saturnide, 1.
Russell, E. S., on the Atractylis
coccinea of T. 5S. Wright, 52.
Saimiri, new species of, 190.
Sarangesa, new species of, 431.
Saturniide, new African, 1.
Schizoporella Alderi, remarks on,
209.
Sciurus, new subspecies of, 426.
Scotopheeus, new species of, 312.
Seminula, remarks on the genus,
223.
ddl
Shark, on a porbeagle, with a large
suboral aperture, 169.
Sheep, on the name of the Armenian
wild, 121.
Sigmaxinyssa, characters of the new
genus, 272.
Silinus, characters
genus, 345,
Silsila, new species of, 22.
Simocephalus, new species of, 489.
Smith, EK. A., on an octopus with
branching arms, 407.
Solenodon paradoxus, on the habits
and external characters of, 68.
Solpuga, new species of, 39.
Spelotrema, new species of, 248.
Spheerotylus, new species of, 272.
Sphecodes, new species of, 450.
Spiders, new African, 33,
Sponges, notes on two African fresh-
water, 523.
Spongilla cerebellata, note on, 525.
Staurophora, new species of, 472.
Stenogyra, new species of, 496,
Stephanoda, new species of, 490.
Swinhoe, Col. C., on new and little-
known Eastern moths, 75 ; on new
African and Indo-Malayan Hespe-
ride, 430.
Synodontis, new species of, 50.
Tagiades, new species of, 430.
Tagoropsis, new species of, 7.
Talpa, new subspecies of, 212.
Tedania, new species of, 279.
Telicota, new species of, 433.
Terebratula, notes on species of,
225.
Terebripora, new species of, 208.
Teretriosoma, new species of, 339,
Thamnomys, new species of, 504.
Theuma, new species of, 299.
Thomas, O., on new mammals from
3ritish New Guinea, 70; on neo-
tropical mammals of the genera
Callicebus &e., 161; on mammals
from Northern Persia, 196; on the
genus Nesokia and on a new
species of Mus, 202; on the occur-
rence of Acanthoglossus in British
New Guinea, 293; on a new dor-
mouse from Asia Minor, 406; on
anew Acanthoglossus from Sala-
watti, 498; on a new flying-
squirrel from Formosa, 522.
Thopha, new species of, 411,
Tilapia, new species of, 50.
of the new
532
Tocotrema, new species of, 257.
Trematode parasites of British birds,
on the, 245,
‘Triepeolus, new species of, 59.
Urabunana, new species of, 423.
Uromys, new species of, 72.
Vejdovsky, Prof. F., on the reduc-
tion of the eye in a new gammarid
from Ireland, 227.
Verrill, A. H., on the habits and ex-
ternal characters of the solenodon
of San Domingo, 68.
Vulpes, new species of, 391.
Woodward, Dr, A. Smith, on a new
INDEX.
leptolepid fish from the Weald
Clay of Southwater, 95.
Wroughton, R. C., on new mammals
from 8. Africa, 381; on the mun-
gooses of the Herpestes gracilis
group, 110; on mammals from the
Bahr-el-Ghazal, 500.
Xanthopimpla, new species of, 19.
Xerophzus, characters of the new
genus, 314,
Zabromorphus, new species of, 100,
343, 482.
Zachresta, new species of, 13.
Zephronia, new species of, 215.
END OF THE TWENTIETH VOLUME.
PRINTED BY TAYLOR AND FRANCIS,
RED LION COURT, FLEET STREET.
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CONTENTS OF NUMBER 120.—Seventh Series.
Page
LXIY. A Revision of the Meduse belonging to the Family Laodi- —
ceide. By Enwarp T. Browns, University College, London
LXY. On new Species of Histeride and Notices of others. By
Gimmie OS. 5. oon ws se ee oe ee ee ee 480
LXVI. Descriptions of Three new fosheaie Fishes discovered
by Mr.G. L. Batesin South Cameroon. By G. A. Bounenerr, F.R.S. 485
LXVII. Descriptions of Three new Fishes from Central Africa.
By G. A. Bovtznesr, F.R.S. Wess e tect eee ees fan ham See eee sae 487
LXVIII. Description of a\new African Snake of the Genus
Simocephalus. By G. A. Boubeyerr, F.RS. ......-... ee ee eee 489
LXIX. Descriptions of new\ Species of Land and Freshwater
Shells from Central and South An ica. By H. B. Preston, F.Z.8. 490
LXX. A new Acanthoglossus ‘Hom the Island of Salawatti.
By OCOFIELD THOMAS o> <r a ek agile 6 2 pos ooo oeinre wie ae 498
LXXI. Description of a new Baboon from British East Africa.
BeOeG. Ersr0r, D.5¢., FBS Ay aes ees og cs ee es bint oe 499
LXXIL. List of a Collection of small Mammals made by Mr. A. L.
Butler in the Bahr-el-Ghazal. By R.C. Wroveuton ............ 500
LXXIII. Descriptions of Five new Sarg of Heterocera. By
Hersert Druce, F.L.S. &e.
LXXIV. New African Species of the Genus Chrysops, Meigen, in
the British Museum (Natural History). By Ervesr E. AUSTEN , . 507
LXXYV. Descriptions of Two new Species of Plecotus. By G. E,
Hi, Bamerre Baer os oo sw ses sa sae ee a 520
LXXVI. Description of a new Species of Monkey (Cercopithecus
Hamlyni) from the Ituri Forest. By R. I. Pocock, F.L.S., Super-
intendent of the Zoological Society’s Gardens ...... gets Sones awe 521
LXXVII. A new Flying-Squirrel from Formosa. By Oxpriexp
PROWES oo as eee ee as ee ee ae ee es re . 522
LXXVIII. Notes on Two Species of African Freshwater Sponges.
Ry RM Marmeravei0e 2.74.5 a eas ae a space ie OE + -523:A :
BIBLIOGRAPHICAL NOTICES.
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