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BULLETIN DU MUSÉUM NATIONAL D’HISTOIRE NATURELLE

3 e série, n° 211, janvier-février 1974, Zoologie 141

Malagasy Endomychidae :

a Collection from the Andringitra Massif

and Anosyennes chains

(Insecta Coleoptera)

by H. F. Strohecker *

Résumé. — Description de deux genres nouveaux : Malagaricophilus (espèce type : M.

spilotus sp. n.) et Bryodryas (espèce type : B. lunidorsum sp. n.). Les autres espèces nouvelles sont

Stenotarsus subglaber, S. nudus, S. microsetosus, S. cordipennis, S. apterus, S. acrophilus, Anagari-

cophilus stillifer, A. nitidus, A. rugosus, A. longus, A. stigmosus. Les espèces Ilaploscelis plagii-

collis (Fairmaire), H. dejectus Strohecker et Cymones atroclavatus (Fairmaire) sont représentées

dans la collection étudiée. Les holotypes et allotypes des nouvelles espèces sont conservés au Muséum

national d’Histoire naturelle (Entomologie), Paris. Sur les 14 espèces récoltées au-dessus de 1 500 m,

7 ont les ailes métathoraciques réduites ou absentes. Les descriptions d ’Anagaricophilus octospi-

lotus, du massif de l’Ankaratra, d’A. vulneratus et de Geoendomychus contusus, de Nandihizana

préfecture de Maroantsetra), ont été ajoutées.

Abstract. — Description of two new généra : Malagaricophilus (type species : M. spilotus

sp. n.) and Bryodryas (type species : B. lunidorsum sp. n.). Other new species are Stenotarsus

subglaber, S. nudus, S. microsetosus, S. cordipennis, S. apterus, S. acrophilus, Anagaricophilus

stillifer, A. nitidus, A. rugosus, A. longus, A. stigmatus. In the studied collection, Haploscelis

plagiicollis (Fairmaire), II. dejectus Strohecker and Cymones atroclavatus (Fairmaire) are repre-

sented. Holotypes and allotypes are kept in National Muséum of Natural History (Entomology),

Paris. Of 14 species taken above 1 500 m, 7 hâve inner wings reduced or absent. Descriptions

added of Anagaricophilus octospilotus, from Ankaratra massif, A. vulneratus and Geoendomychus

contusus, from Nandihizana (Maroantsetra Préfecture).

The Endomychidae taken in the course of the Recherche Coopérative sur Programme

n° 22b 1 , established by the Centre National de la Recherche Scientifique, were sent to me

for study by M. A. Descarpentries of the Entomological Laboratory of the National

Muséum of Natural History in Paris.

This family of beetles is eminently tropical and, while few généra hâve been found

to be represented in Madagascar, numerous species hâve been described, especially of the

généra Haploscelis and Stenotarsus. The collection made in the Andringitra massif (Center

Madagascar) and the Anosyennes chains (East Madagascar) includes, besides material of

these two généra, examples of Mycetaeids, up to now little known in Madagascar.

* University of Miami, Department of Biology, Coral Gables, Florida 33124, USA.

1. 1971, Bulletin de la Société d’Écologie, 2 (2-3) : 189-266 ; 1973, Bulletin du Muséum national

d’Histoire naturelle, 3 e série, n° 118, janv.-févr., Ecologie générale 1 : 1-40.

211, l

302

H. F. STROHECKER

The most notable feature of this assemblage of specimens is the high proportion of

forms with aborted metathoracic wings and highly convex elytra. The apterous condition

is rare in the Endomychidae but I hâve noted it in the case of a few Asiatic species ; in

these cases also the elytra are unusually convex. Of 14 Malagasy species taken above

1 500 meters 7 are flightless. What adaptation may be conferred by this loss of wings

and elytral élévation is not self-evident ; perhaps it increases air space and enables the

insect to survive periods of inundation during heavy rainfall. One may recall the expé¬

rience of Heberdey (as quoted by Wigglesworth, 1939) that removal of the wings of

Dytiscids allowed the beetles to prolong their period of submergence.

Appended to this report are descriptions of three new species based on specimens

collected in other areas of Madagascar.

In the figures, except no. 17, pronotum and elytron hâve been drawn separately,

in each case with latéral margins approximately perpendicular to line of sight.

Holotypes and allotypes of new species described are in the Muséum national d’His-

toire naturelle, Paris. One or more paratypes, when available, are in author’s collection.

Subfamily Eumorphinae

Genus Haploscelis Blanchard

Twenty-eight species of this endemic and essentially tropical genus hâve been described.

Haploscelis plagiicollis (Fairmaire)

Hybopterus plagiicollis Fairmaire, 1883, Le Naturaliste, 2 : 365, type male : Paris Muséum.

The few specimens of this distinctive species which I hâve seen up to now hâve corne

from the Andringitra massif. In the collection now at hand are 18 specimens. The inner

wings are absent and the elytra more gibbous than is usual in the genus but its basic struc¬

ture is similar to the other species.

Andringitra Est : Anjavidilava, 1 850-1 950 m, 18-XII/15-I-1971, FDHM2 ; Maro-

sitry, Est riv. Antsifotra, 2 000 m, 2/5-XII-1970, FDSM ; Andringitra Centre : plateau

Andohariana, 2 000-2 100 m, 9-XI/10-XII-1970, FDHM2.

Haploscelis dejectus Strohecker

Haploscelis dejectus Strohecker, 1972, Cah. ORSTOM , sér. Biol., 16 : 126, type male : Paris Muséum.

The unique type was among specimens which became detached from card-mounts

and its provenance could not be determined.

MALAGASY ENDOMYCHIDAE

303

Andringitra Sud : Andrianony, cirque Manjarivolo, 1 800 m, 26-X/3-XI-1970, FDHM2,

a single male.

Chaînes anosyennes : Massif nord, 1 050 m, moyenne Ranomandry, 11/30-XI-1971,

FDHMA, a female which seems to be of this species.

Genus Cymones Gorham

Cymones atroclavatus (Fairmaire)

Dioedes atroclavatus Fairmaire, 1883, Le Naturaliste, 2 : 365, type female : Paris Muséum.

Cymones cowani Gorham, 1886, Proc. Zool. Soc. London, 1886 : 158, type male : unknown to me.

Encymon armipes Fairmaire, 1899, Ann. Soc. ent. France, 68 : 469, type male : Paris Muséum.

As recognized by me the species has wide distribution in tropical Madagascar. Some

of the différences in color may represent racial divergence. Inner wings normal. Fifteen

specimens with data

Chaînes anosyennes : Massif nord, 1 050 m, moyenne Ranomandry, 11/30-XI-1971,

FDHMA.

Subfamily Stenotarsinae

Genus Stenotarsus Perty

Twenty-one Malagasy species of this large circumtropical genus hâve been described

and additional species are recorded here. Some of these show unusual features but do not

seem to transgress the accepted définition of Stenotarsus.

Stenotarsus subglaber sp. n.

(Fig. 1, 2)

Elliptic in outline, moderately convex for genus. Under surface, legs, antennal stalk,

head and pronotum red, elytra, including epipleura, black with umbo, base narrowly and

apex broadlv red. Length 6,3-7,5 mm.

Although at 30 X fine short hairs can be seen on upper surface the general aspect of

the insect is glabrous. Antenna of male 2,8 mm, articles 3-8 longer than wide, club large,

its last article almost square. Pronotum with médian length 3/5 basal width, raised side

margins low, fiat, rather narrow (.2 mm) and of equal width throughout, dise finely and

sparsely punctured, latéral sulci broad, shallow, transverse sulcus évident only near latéral

sulci. Elytra broader at base than pronotum, umbo prominent, dise finely and sparsely

punctured. The front border of the red apical area runs obliquely forward from suture

to side margin. The first sternite of male is slightly concave at middle and there are two

foveae behind each metacoxa. Inner wings normal.

304

H. F. STROHECKER

Holotype male : Andringitra Est : Anjavidilava, 1 850-1 950 m, 18-XII/15-I-1971,

FDHM2.

Allotype female : same data as holotype.

Paratypes : 1 male, 4 females with same data as holotype.

Stenotarsus nudus sp.

(Fig. 3, 4)

In general appearance much like a miniature of S. subglaber but with raised pronotal

margins proportionately wider and antennal stalk articles bead-like, article 9 as wide distad

as long, narrower than 10, 11 very large, longer than wide, its apex feebly rounded. Upper

surface very finely punctured and apparently glabrous but fine short setae can be seen

at 60 X , especially in région of umbo. Length 3,3 mm, max. width 2,2 mm.

Holotype male : Andringitra Sud : Andrianony, cirque Manjarivolc, 1 650 m, 26-X/3-

XI-1970, FDHM1.

I'ig. 1-10. — Gen. Stenotarsus Perty. — 1, S. subglaber sp. n., edeage, left side ; 2, id., dorsal face. — 3,

S. nudus sp. n., edeage, left side ; 4, id., dorsal face. — 5, S. microsetosus sp. n., edeage, left side ; 6,

id., dorsal face ; 7, id., antennal club. — 8, S. cordipennis sp. n., edeage, left side ; 9, id., dorsal face ;

10, id., antennal club.

Allotype female : same data as holotype.

Paratypes : 1 female with same data as holotype, 1 male, 10 females, Andringitra Est :

Anjavidilava, 1 850-1 950 m, 18-XII/15-1-1971, FDHM2. The females are larger (4-4,5 mm)

than the males and elytral coloration varies front similarity to males to elytra mostly red

with large comrnon black area but this is probably due to âge différences. Inner wings

normal.

MALAGASY ENDOMYCHIDAE

305

Stenotarsus microsetosus sp. n.

(Fig. 5-7)

Of elliptic outline, about equally rounded in front and behind, normally convex for

genus. Inner wings normal. Under surface, including elytral epipleura, dark red, last

four sternites paler, metasternum of male tumid, coarsely and densely punctured at middle.

Sternite 1 convex at middle, very finely punctured ; two foveoles behind metacoxa. Anten-

nae stout, stalk articles bead-like, club large, articles 9-10 transverse, 11 almost square,

truncate. Pronotum red, raised side margins low, wide and fiat, latéral sulci continued

as fine grooves to front margin, dise finely and sparsely punctured with sparse but conspi-

cuous pubescence. Elytra rather coarsely punctured but with interspaces wider than

punctures, pubescence sparse and short. Elytra black with base and apical fifth red.

Length 3,8 mm, max. width 2,5 mm.

Holotype male : Chaînes anosyennes : Massif nord, moyenne Ranomandry, 1050 m,

11/30-XI-1971, FDHMA.

Stenotarsus cordipennis sp. n.

(Fig. 8-10)

Under surface, legs, head, antennal stalk, pronotum, scutellum, elytral base, umbo,

side margins and apex red ; elytra mostly black, densely punctured and with fine recumbent

pubescence. Length 3,5 mm, max. width 2,2 mm.

Antennal articles 5-7 slightly longer than wide, 10 transverse, 11 cycloid. Pronotum with

basal width slightly more than twice médian length, narrowed in front to half basal width,

raised margins rather narrow (. 1 mm), latéral sulci linear and deep, reaching to mid-length

of pronotum, transverse sulcus shallow, indistinct, dise and sides conspicuously punctured

and pubescent. Elytra 1,3 times as long as their combined width, a little wider at base

than pronotum, widening slightly in anterior third then gradually narrowed to apices which

are feeblv attenuate and separately rounded. Metasternum of male densely, coarsely

punctured, sternite 1 densely, coarsely punctured at middle and with two foveoles behind

each metacoxa. Inner wings normal.

Holotype male : Chaînes anosyennes : Massif nord, haute Ranomandry, 1 900 m,

17/26-XI-1971, FDFIM.

Allotype female : same data as holotype. The elytra of this specimen are ferruginous

but this, I feel certain, is a teneral condition. In ail other features, including punctation

of metasternum and sternite 1, it is very similar to holotype.

Paratypes : 3 teneral ? females with same data as holotype.

306

H. F. STROHECKER

Stenotarsus apterus sp. n.

(Fig. 11-13)

Oval in outline, strongly convex, under surface, including elytral epipleura, legs,

antennal stalk and pronotum red. Elvtra shilling black with sparse recumbent tawny

pubescence ; umbo dark red. Length 3 mm, max. width 2,1 mm.

Antennae stout, articles 4-5 short-cylindric, the other stalk articles bead-like, 9 evenly,

widened distad and as long as wide, 10 transverse, 11 elliptic, slightly longer than wide,

1,5 times as wide as 9. Pronotum with raised margins very narrow (for genus) and fiat,

latéral sulci extending forward to mid-length and ending basad in a deep fovca. Elytra

gibbous, overhanging latéral margins, tapering from anterior third and somewhat cordi-

form in outline, punctures conspicuous but well separated. Tuner wings aborted. Meta-

sternum convex, densely and coarsely punctured. Stérilité 1 sparsely punctured and with

foveoles behind metacoxa.

Holotype male : Chaînes anosyennes : forêt au pied Nord du Piton Andohahelo, 1 880 m,

V-1972, FDHM.

Stenotarsus acrophilus sp. n.

(Fig. 14-16)

Oval in outline, highly convex, under surface and legs reddish black, antennal stalk,

head and pronotum dark red, elytra, including epipleura, shining black with very fine,

short, sparse pubescence. Length 3,4 mm, max. width 2,2 mm.

Antennae with articles 2-7 short-cylindric, 9 svmmetrically widened to apex, 11 ovoid

with apex subtruncate. Pronotum shining, sparsely punctured, side margins modcrately

wide, low and fiat, latéral sulci linear and deep, transverse sulcus very close to hind margin.

Elytra gibbous, overhanging side margins, sparsely punctured. Inner wings absent.

Flolotype female : Chaînes anosyennes : Massif nord, haute Ranomandry, 1 900 m,

17/26-XI-1971, FDHM.

Paratypes : 3 females with same data as holotype.

Very similar to S. apterus and I hâve considered it as possibly the female of that spe-

cies. It difTers, however, in several features : the prosternai process is much broader than

in S. apterus, the mesosternum more declivent in front and the antennae slenderer with

longer stalk articles. The under surface is pitchy in S. acrophilus, clear red in 5. apterus.

Subfamily Mycetaeinae

Generic définitions are still difïicult in this group. A number of type species are known

from little material, some from unique specimens.

MALAGASY ENDOMICHYDAE

307

Fig. 11-16. — Gen. Slenolarsus Perty. —- 11, 5. apterus sp. n., edeage, left side ; 12, id., dorsal face ;

13, id., prosternum (same scale as fig. 16). — 14, S. acropliilus sp. n., habitus ; 15, id., antenna ; 16,

id., prosternum (same scale as fig. 13).

Genus Anagaricophilus Arrow

Devised by Arrow for A. pulchellus based on a single specimen from the Seychelles.

J. Vinson added a second species, A. concinnus, from Mauritius. A feature of A. concinnus

not noted by Vinson is a small dimple at the elytral apex of some specimens, probably

males.

Although Arrow described the tarsus of A. pulchellus as having four articles and it

was so illustrated by me from drawings received from the British Muséum (Nat. Hist.),

the late Dr. Vinson and I examined the unique type in 1960 and agreed that the small

penultimate tarsomere of Arrow’ s diagnosis could not be discerned.

The new species here placed in the genus hâve the prosternai process completely mar-

gined, palps large, last article of maxillary palp oval, of labial palp subcycloid ; three tarso-

meres ; latéral sulci of pronotum with external carina coursing far forward.

Anagaricophilus stillifer sp. n.

(Fig. 17)

Reddish black below with elytral epipleura red ; shining black above, each elytron

with three orange markings. Length 2,2-2,4 mm.

Ten antennomeres, 1-2 stout, 3 slender, 4-7 bead-like, 8-10 forming broad loose club

winch is as long as stalk ; stalk red, club black. Pronotum with sides parallel in basal half,

rounded to blunt front angles, narrowly margined, discal punctures and setae fine and

sparse, latéral sulci deep, triangular and connected by a transverse sulcus close to hind

margin. Elytra strongly convex in both axes, parasutural stria deep but evanescent near

308

H. F. STROHECKER

triaugular scutellum, punctures deep, rather coarse and dense, setae hardly évident. Of

the orange markings two are rounded and occupy an oblique line before mid-length, the

third is a transverse bar on apical slope.

In male each elytron has a rounded prominence before apex and distomesad of this

eminence the suturai margin has a dense row of erect setae. Elytral apex of female simple.

Inner wings fully developed.

Holotype male : Andringitra Sud : Andrianony, cirque Manjarivolo, 1 650 m, 26-X/3-

XI- 1970, FDHM1.

Allotype female : Andringitra Centre, plateau Andohariana, 2 000-2 100 m, 9-XI/10-

XII- 1970, HFAM1.

Paratype : male with data of allotype (these two specimens somewhat, teneral). Three

females, Maroantsetra pref., Nandihizina, 10-XII-1938 (J. Vadon) appear also to be of

this species.

Fig. 17-19. — Gen. Anagaricophilus Arrow. —• 17, A. stillijer

sp. n., habitus, (J. — 18, A. vulneratus sp. n., apex of elytron of <J.

— 19, A. nitidus sp. n., apex of elytron of <J.

Anagaricophilus nitidus sp. n.

(Fig. 19)

Of elliptic outline, highly convex, under surface black, antennal stalk dark red, upper

surface shining black, fînely and sparsely punctured, with a minute seta (60x) in each

puncture, legs black, tarsi reddish. Length 1,4 mm, max. width 1,25 mm.

Ten antennomeres, 1-2 long and stout, 3 narrow, short-cylindric, 4-7 bead-like, 8-10

forming a large, loose club which is as long as 1-7 combined. Pronotum nearly twice as

wide as its médian length, sides narrowly margined, nearly parallel up to the curvature

to front angles, latéral sulci triangular and deep, basal sulcus close to hind margin. Elvtra

almost 3x as long as pronotum, their sides continuously rounded, parasutural stria eva-

nescent toward scutellum. Wings fully developed Legs rather long, slender, tarsi

slender, third tarsomere equal to fîrst two combined.

MALAGASY ENDOMYCHIDAE

309

In male elytral tip is arcuately impressed, the impressions in caudal view hâve the form

of a conventionalized heart. Elytral tips of female simple.

Holotype male : Andringitra Est : Anjavidilava, 2 000 m, 18-X1I/15-I-1971, FDSMP.

Allotype female : Anjavidilava, 1 850-1 950 m, FDHM2.

Paratypes : male, Andringitra Centre : plateau Andohariana, 2 000-2 100 m, 9-XI/10-

XII-1970, HFAM1 ; female with same data as allotype, female, Andringitra Sud : Andria-

nony, cirque Manjarivolo, 1 650 m, 26-X/3-XI-1970, dans mousses sur arbre, FDHM1 ;

Chaînes anosyennes : Massif nord, 1 050 m, moyenne Ranomandry, 11/30-XI-1971, FDHMA.

Anagaricophilus rugosus sp. n.

(Fig. 20)

Outline short-elliptic, strongly convex, black below and above, tibiae, tarsi and anten-

nal stalk pale and each elytron with three or four yellow marks, sparselv pubescent. Length

1,6 mm, max. width 1,15 mm.

Antennomeres ten, 1-2 stout but longer than wide, 3 slender, 5-7 bead-like, 8-10 for-

ming large club which is 2/3 as long as stalk. Pronotum coarsely and densely punctured,

sides about parallel basad, raised margin rather broad in front but narrowed to hind angles,

latéral sulci broadly triangular, connected by deep transverse sulcus close to hind margin.

Elytra strongly convex in both axes, side margins rather wide, parasutural stria feeble

in front but deeper caudad, surface densely and coarsely punctured. Each elytron bas

a round yellow spot on mid-line before middle and a similar spot meso-caudad of this.

Behind the outer spot is a long-triangular spot and behind the inner spot is a J-shaped

(left elytron) yellow mark, this tending to coalesce with the triangular spot to form a U.

Elytral tips of female are feebly produced. Inner wings absent.

Holotype male : Andringitra Est : Marositry, Est riv. Antsifotra, 2 000 m, 2/5-XII-

1970, FDSM.

Allotype female : data same as for holotype.

Paratypes : 4 females with same data as holotype and 2 females from Anjavidilava,

1850-1 950 m, 18-XII/15-1-1971, FDI1M2.

Anagaricophilus longus sp. n.

(Fig. 21, 22)

Form long-elliptic, moderately convex, reddish black below, black above, each ely¬

tron with three large yellow markings, coarsely punctured, sparsely pubescent. Length

2 mm, max. width 1,2 mm.

Antennomeres ten of which 7-10 form a club subequal in length to stalk. Pronotum

2,3 times as wide as its médian length, narrowly margined at sides, latéral sulci triangular

and shallow, transverse sulcus narrow, close to hind margin. Elytra about 3,5 times as

310

H. F. STROHECKER

Fig. 20-22. — Gen. Anagaricophilus Arrow. —• 20, A. rugosus sp. n., habitus. —

21, A. longus sp. n., habitus ; 22, id., antenna.

long as pronotum, sides subparallel from base to apical third, evenly rounded to apex,

surface coarsely and densely punctured. Each elytron has a broad yellow vitta extending

from base across umbo to near mid-length, a large semicircular spot touching the parasu-

tural stria and a large triangular yellow mark covering most of caudal slope. Inner wings

normal.

Holotype female : Andringitra Centre : plateau Andohariana, 2 000-2 100 m, 9-XI/10-

XII-1970, HFAM1.

Anagaricophilus stigmatus sp. n.

(Fig. 23)

Lower surface and legs black, elytral epipleura dark red. Upper surface highly con-

vex, sparsely pubescent, black, each elytron with four yellow spots. Antennal stalk and

tarsi pale. Length 1,75 mm, max. width 1,25 mm.

Nine antennomeres, 1-2 stout but longer than wide, 3 slender, cylindric, 4-6 bead-like,

7-9 forming a stout club which is slightly shorter than 1-6 combined. Pronotum 2X as

wide as its médian length, strongly convex in transverse axis, side margins wide in front,

narrowed to the obtuse hind angles, latéral sulci broadly triangular and very shallow basad,

connected by a transverse sulcus, dise distinctly but rather finely punctured. Elytra

highly convex in both axes, narrowly margined, densely and rather coarsely punctured,

parasutural stria feeble on basal half, distinct caudad. Each elytron has a round yellow

spot near suture and mid-length and, slightly anterior to this an angulate spot on latéral

slope. Behind these is a pair of spots similarly placed. Inner wings absent.

Holotype female : Andringitra Est : Anjavidilava, 2 000 m, 18-XII-15-I-1971, FDSMP.

MALAGASY ENDOMYCHIDAE

311

Fig. 23-24. — 23, Anagaricophilus stigmatus sp. n., habitus. —

24, Malagaricopliilus spilotus gen. and sp. n., habitus.

Genus Malagaricophilus nov.

Tarsi trimerous, unlobed, mandible minutely trifid at apex, last article of maxillary

palp subconical, of labial palp strongly transverse. Prosternum briefly prolonged behind

coxae, its apex spatulate, margined. Mesocoxae narrowly separated, their cavities closed

by sterna. Elytra without parasutural stria. Pronotum with base strongly undulate,

i.e. lobed at middle.

Type species : Malagaricophilus spilotus sp. n.

Malagaricophilus spilotus sp. n.

(Fig. 24)

Long-oval in form, moderately convex, black below and above, coarsely pubescent,

each elvtron with two round vellow spots near mid-line. Length 2 mm, max. width 1,4 mm.

Antennal stalk yellow, its fîrst two articles stout, 3 narrow and cylindric, 4-7 bead-

like, 8-10 forming a large black club which is slightly shorter than the stalk. Head eoarselv,

shallowly punctured. Pronotum with sides continuouslv rounded, marginal bead narrowed

behind, latéral sulci broad, triangular, deep, transverse sulcus very close to hind margin,

surface coarsely and rather closely punctured, pubescence grav. Elytra 3x or more as

long as pronotum, subparallel, densely and coarsely punctured. Each elvtron bas two

transversely oval orange marks near mid-line, the fîrst a little before mid-length, the second

on caudal slope. Inner wings strap-like, no longer than elytra.

Holotype female : Andringitra Est : Anjavidilava, 2 000 m, 18-XII/15-I-1971, FDSMP.

312

H. F. STROHECKER

Genus Bryodryas nov.

Highly convex, subhemispherical. Tarsi trimerous, stout, claws appendiculate.

Antennomeres nine, of which the last three form a stout club. Last article of maxillary

palp conoid and minutely truncate, of labial palp very large and broadly truncate. Man-

dible bifid at tip. Prosternai process prolonged beliind coxae, its apex slightly narrowed,

rounded. Mesocoxae rather narrowly separated, their cavities closed by the sterna. Scu-

tellum transversely triangular. Upper surface coarsely pitted, parasutural stria of elytra

very wide and deep. The generic naine is féminine.

Type species : Bryodryas lunidorsum sp. n.

Bryodryas lunidorsum sp. n.

(Fig. 25-29)

Black below and above, legs, elytral epipleura and antennae paler. Length 1,3 mm,

max. width 1,15 mm.

Antennomeres 1-2 oval, stout, 3 much narrower than 2 but hardly longer than wide,

3-(i bead-like, club about as long as stalk, the last article cycloid. Pronotum twice as

wide as médian length with transverse impressed line near front margin, sides continuously

curved, marginal bead wide near front angles, narrowed behind, surface with crater-like

punctures, latéral sulci deep, triangular, transverse sulcus deep and close to hind margin.

Elytra very convex in both axes with dense crater-like punctures, parasutural stria wide

and deep, suturai area smooth. At 60X some minute setae can be seen but the insect

appears glabrous. Inner wings absent.

Fig. 25-32. — 25, Bryodryas lunidorsum gen. and sp. n., habitus ; 26, 27, id., labial palp ; 28, id., maxil-

lary palp ; 29, id., antenna. — 30, Geoendomychus contusus sp. n., habitus ; 31, id., prosternum ; 32, id.,

antenna.

MALAGASY ENDOMYCHIDAE

313

Holotype (sex ?) : Andringitra Sud : Andrianony, cirque Manjarivolo, 1 650 m, 26-X/3-

XI-1970, FD1IM1, mousses sur rocher.

Paratype : female with same data as holotype. This specimen fell apart when relaxed

and lias been mounted in Euparal.

Anagaricophilus vulneratus sp. n.

(Fig. 18)

Under surface black, legs and antennal stalk yellow, above shinning black, eaeh elv-

tron with two transverse yellow areas. Length 1,5 mm.

Nine antennomeres, 1-2 stout, 3 short-cylindric, 4-6 bead-like, 7-9 forming a broad

club which is as long as stalk. Pronotum strongly convex, distinctly punctured, narrowly

margined at sides, latéral sulci triangular, shallow but with long external carina, transverse

sulcus deep, close to hind margin. Elytra coarsely and rather densely punctured, each

with a transversely quadrate orange bar on basal half and a similar luit smaller pre-apical

bar. Inner wings normally developed.

In male each elytron is deeplv impressed at apex and within the impressed area is a

recurved hook, evidently formed of agglutinated setae.

Holotype male : Madagascar : Maroantsetra pref., Nandihizina, 12-X-1938 (J. Vadon ).

Anagaricophilus octospilotus sp. n.

(Fig. 33-35)

Dark red beneath, legs and antennal stalk reddish. Antennal club black. Upper

surface deep black, shilling, each elytron with four yellow spots. Length 1,6 mm, max.

width 1,1 mm.

Tarsi trimerous, unlobed. Prosternai process prolonged behind coxae, its apex rmm-

ded, margined. Antennomeres nine, 1-2 stout but longer than wide, 3 slender, 4 quadrate,

5-6 bead-like, 7-9 forming large club which is 3/4 as long as stalk. hast article of maxil-

lary palp long-oval, of labial palp subquadrate, broadly truncate. Pronotum twice as

wide as long, narrowly margined, strongly convex, finely punctured, setae minute and sparse.

latéral sulci triangular, rather deep, their external carina reaching to middle of dise, trans¬

verse sulcus close to base. Elytra rather coarsely and densely punctured, the interspaces

about equal to punctures. Setae minute, sparse. Wings normal.

Holotype female : Massif de l’Ankaratra, Mont Tsiafajavona, 2 640 m, 7/12-1-1972,

VR.

Paratypes : two females with same data as holotype. In one of the paratypes the

caudal spot is narrowly connected to a small latéral spot at the same latitude ; in the other

paratype there is a dark interval between these spots. It seems évident to me that these

two conditions represent stages in the development of pattern shown by the holotype.

314

H. F. STROHECKER

Fig. 33-35. — Gen. Anagaricophilus Arrow. — 33, A . octospilotus sp.

n., habitus ; 34, id., antenna ; 35, id., female genitalia (ventral).

Genus Geoendomychus Lea

I ara not altogether happy with generic placement of the new species described below.

Referral to Geoendomychus agréés fairly well with Arrow’s use of the name but I hâve

seen but one specimen of the generotype, G. puhescens Lea front Lord Howe Isl., and hâve

now no specimen of it for comparison. As the mycetaeids become represented by more

material in collections it will probably lie useful to deline additional généra.

Geoendomychus contusus sp. n.

(Fig. 30-32)

Lower surface, legs and antennal stalk reddish, black above, each elytron with a large

red marking covering most of caudal half. Length 1,6 mm, max. width 1,2 mm.

Antennomeres ten, 1-2 stout but longer than wide, 3 cylindric, 4-7 transverse, 8-10

fornting a very large club which is longer than 1-7 combined. Pronotum convex, sides

narrowly margined, parallel basad, much rounded to front angles, dise finely and sparsely

punctured, latéral sulci short, triangular and shallow, transverse sulcus close to hind mar-

gin. Elytra but little wider at base than pronotum, roundly widened to mid-length, thence

continuously rounded to apex, latéral margins, in dorsal view, invisible except near base,

surface fairly densely and coarsely punctured (punctures about equal to interspaces), pubes¬

cence not dense, moderately long, semi-erect. Last article of labial palp transverse. Pro¬

sternai process contracted toward apex which is rather narrowly rounded and unmargined.

In the male the elytral apex is dimpled, the suturai margin within the dimple elevated

and with a recurved bristle. Latéral to the dimple is a small conical eminence bearing

a large seta directed caudad. The cône is well removed from the margin but the elytron

MALAGASY ENDOMYCHIDAE

315

lias been posed to bring it into sharp view. The seta is apparently compound ; on the

left elytron it appears double, although single on the right (which has been used in the

figure).

Ilolotype male : Maroantsetra pref., Nandihizina, 10-XII-1938 (J. Vadon).

Manuscrit déposé le 27 février 1973.

Bull. Mus. nain. Hist. nat., Paris, 3 e sér., n° 211, janv.-févr. 1974,

Zoologie 141 : 301-315.

Achevé d’imprimer le 30 septembre 1974.

IMPRIMERIE NATIONALE

4 564 001 5

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Les références bibliographiques apparaîtront selon les modèles suivants :

Bauchot, M.-L., J. Daget, J.-C. Hureau et Th. Monod, 1970. — Le problème des

« auteurs secondaires » en taxionomie. Bull. Mus. Hist. nat., Paris, 2 e sér., 42 (2) : 301-304.

Tinbergen, N., 1952. — The study of instinct. Oxford, Clarendon Press, 228 p.

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