European Journal of Taxonomy 141: 1-138

http://dx.doi.org/10.5852/ejt.2015.141

This work is licensed under a Creative Commons Attribution 3.0 License.

ISSN 2118-9773

www. europeanj ournaloftaxonomy. eu

2015 • Miles C.

Monograph

urn:lsid:zoobank.org:pub:2838F4A0-3FF9-4985-ACED-84CCBF9C59A3

The Earwig Collection (Dermaptera) of the Manchester Museum, UK,

with a complete type catalogue

Claire MILES

Department of Entomology, The Manchester Museum, University of Manchester, Oxford Road,

Manchester, M13 9PF, UK. Email: [email protected]

urn:lsid:zoobank.org:author:527E8319-57C4-4C8D-8896-46FA09795B2E

Table of Contents

Abstract. 1

Introduction . 2

Description of the Dermaptera collection . 2

Development of the Dermaptera collection . 3

Walter Douglas Hincks (1906-1961) . 8

Alan Brindle (1915-2001). 13

The Entomological Archive . 15

Notable earwig specimens in the MMEJE collection . 16

Giant Earwig, Labidura herculeana (Fabricius, 1798) . 16

Parasitic earwigs . 17

Type catalogue of the Dermaptera collection at MMUE . 20

Other material . 123

Acknowledgements . 125

References. 125

Appendix I: Correspondence in the Hincks archive relating to Dermaptera . 134

Appendix II: Correspondence in the Brindle archive relating to Dermaptera . 137

Abstract. This paper presents a description of the worldwide Dermaptera collection of the Manchester

Museum, UK. In total, the collection contains over 16,700 specimens and represents 835 species and

subspecies (approximately 44% of the Dermaptera world fauna). The composition of the collection is

described and a brief account is given of its history and those instrumental in its development, notably

W.D. Hincks (1906-1961) and A. Brindle (1915-2001). Correspondence held at the museum relating

to their work on Dermaptera is listed and examples relating to Labidura herculeana (Fabricius, 1798)

and some of the parasitic earwigs in the collection are explored. Finally, a complete type catalogue of

the Manchester Museum’s Dermaptera is presented; it is arranged alphabetically by species name as

originally described, with photographs of all holotypes and their corresponding data labels. The collection

contains 40 holotypes and 694 secondary types (allotypes, syntypes or paratypes), representing 283

species and subspecies, an estimated 15% of the Dermaptera world fauna.

Key words. Dermaptera, type catalogue, Manchester Museum, W.D. Hincks, A. Brindle.

Miles C. 2015. The Earwig Collection (Dermaptera) of the Manchester Museum, UK, with a complete type

catalogue. European Journal of Taxonomy 141: 1-138. http://dx.doi.org/10.5852/eit.2015.141

European Journal of Taxonomy 141: 1-138 (2015)

Introduction

The collections of arthropods at the Manchester Museum (MMUE) are among the top three in the UK,

containing more than three million specimens, including about two and a half million insects (Logunov

& Merriman 2012; Logunov 2010a). The worldwide Dermaptera collection is one of the particular

strengths of the Entomology Department and it is probably among the most comprehensive in the world

(Logunov 2010b), containing almost a half of the species described worldwide (see Table 1).

During the review of the collection, an updated count was made of all its constituent parts (Table 1).

The total of over 16,700 specimens is approximately 5,000 more than previously thought (Logunov &

Merriman 2012). In the process, accession numbers have been added to all individual dried, mounted

primary and secondary type specimens and to all vials of spirit specimens.

The aims of this paper are (1) to briefly describe the composition of the Museum’s Dermaptera collection

and its history and (2) to provide a complete catalogue of its type material. The Manchester collections

are searchable online at http://www.museum.manchester.ac.uk : however, the Dermaptera collection is

not yet completely databased on the Museum’s collection management system to species level and this

significant resource is currently very little used. It is hoped that the publication of this paper with a

complete type catalogue will encourage and facilitate the use of this important, but underpublicized and

underused collection.

Description of the Dermaptera collection

The collection comprises over 16,700 specimens organised in five divisions:

1. Approximately 10,800 worldwide specimens (accession no. L3233) are dry mounted, housed in three

Watkins and Doncaster wooden cabinets, each with 20 glass-lidded drawers (Tig. 1). The specimens

are pinned or mounted on card and arranged taxonomically. Table 1 gives the numerical composition

of the collection and Tig. 2 gives the percentage of World Dermaptera represented at the Manchester

Museum, by family. Three drawers (approx. 1300 specimens) contain undetermined material, partially

sorted according to the zoogeographical regions. Table 2 shows the geographical distribution of the

undetermined material.

2. The British earwig collection is kept separate from the foreign insects and is housed in one drawer of

a Hill unit (Tig. 3). The four native species are represented, as well as Euborellia annulipes (Lucas,

1847) and Marava arachidis (Yersin, 1860), with 250 specimens in total (Table 3). In addition, there

is one specimen of Labidura riparia (Pallas, 1773) from Prance in this collection.

3. There are around 2000 undetermined specimens of papered material. 1600 are Indian Dermaptera

(accession no. P3254) collected between 1953 and 1972 (Pig. 4). They were supplied by the well-

known Indian entomologist and natural history dealer P. Susai Nathan (1891-1976) of Kurumbagaram

(the ‘P’ is the initial of his father’s name, Prabala Nathan; M.A.N. Jayapaul pers. comm., 2015).

hollowing his retirement in 1969, his daughter-in-law Theresa Rajabi Susai Nathan took over the

dealership (see comments in the text under ‘Alan Brindle’). 185 specimens (accession no. L3241)

are from South America, collected by Czeslaw M. Biezanko (1895-1986), a Polish entomologist and

recognized authority on South American butterflies. A further 200 specimens are from northern India

collected between 1953 and 1956. The collector is identified only as ‘P.S’.

4. The spirit collection contains approximately 3500 Dermaptera specimens stored in 70% industrial

methylated spirit in glass tubes with polythene caps. The tubes are stored upright in plastic boxes

lined with foam plastic. While completing this project, the opportunity was taken to re-curate the

spirit collection by topping up where necessary, restoring specimens which had completely dried

MILES C., Dermaptera of the Manchester Museum

Table 1 . Composition of the Manchester Museum’s Dermaptera collection.

No. of species and subspecies

MMUE’s

MMUE collection representation No. of specimens

Family

World

Fauna 1

Dry

Spirit

Combined

Dry/Spirit 2

of World

Fauna (%)

Dry

Spirit

Total

Anisolabididae

393

170

43.3

1426

532

1958

Apachyidae

53.3

Arixeniidae

40.0

Chelisochidae

33.7

463

464

Diplatyidae

144

29.9

136

Forficulidae

470

185

39.4

3077

820

3897

Hemimeridae

54.5

Karschiellidae

91.7

Labiduridae

47.7

1044

209

1253

Pygidicranidae

185

49.7

734

735

Spongiphoridae

510

250

49.0

2825

1386

4211

Undetermined

—

3326 3

544

3870

Total

1916

834

835

43.6

13,221

3528

16,749

1 Hopkins et al. (2014) (Extant species).

2 Species present in both dry and spirit collections counted only once.

3 Includes both mounted and papered material.

out, and adding accession numbers to all vials. There are two species in the spirit collection not

represented in the dry collection. One is Hemimerus hanseni Sharp, 1895. The other is a species for

which the author was unable to find reference to a published description, Pericomus nigra, identified

by Brindle in 1980. The specimens of this species were part of around 5000 collected by Dr. Henk

Wolda in Panama (see below in the type catalogue). Wolda was an entomologist and staff member at

the Smithsonian Tropical Research Institute. There is no reference to this species in Brindle’s notes

from his analysis of the material (MMUE, Brindle archive, items 129, 138), from which he did,

however, describe Pericomus ater as a new species.

5. 108 specimens of earwigs imported in foodstuffs or plant material form part of a reference collection

(accession number F3231) initiated by W.D. Hincks (see section ‘Walter Douglas Hincks’ below).

Five species in 4 genera are represented and include 89 specimens from East Africa, 14 from West

Africa and 2 from Saudi Arabia, almost all found in imported bark or logs.

In summary, 835 species and subspecies are represented in the entire collection, approximately 44% of

the Dermaptera world fauna. The number of species is comparable to those held at the British Natural

History Museum (BMNH; now Natural History Museum), London (855 species). The number of

specimens at MMUE is 75% that of BMNH, which holds 450 primary and 1200 secondary Dermaptera

types (Natural History Museum 2015b). A complete type catalogue of Dermaptera at MMUE is provided

below.

Development of the Dermaptera collection

Two curators of the Manchester Museum Entomology Department, Walter Douglas Hincks (1906-1961)

and Alan Brindle (1915-2001), were chiefly responsible for developing the extent and geographical scope

of the collection as it is today, both being the world authorities on the order in their time. Hincks was

European Journal of Taxonomy 141: 1-138 (2015)

Table 2. Number of undetermined Dermaptera specimens in the Manchester Museum by geographical

region.

Region

No. of specimens

Dried, mounted

Dried, papered

Spirit

Total

Oriental

464

1622

2086

Afrotropical

288

290

Australasia

191

Neotropical

300

185

470

955

Palearctic

202

253

Nearctic

No data

1317

2009

544

3870

Assistant Keeper of Entomology at Manchester Museum from 1947 to 1961 (the title changed to Keeper

in the 1950s) and Brindle was the Keeper of Entomology from 1961 to 1982.

The Museum was founded by the Manchester Society for the Promotion of Natural History in 1821. A

history of the development of the museum is provided by Alberti (2009). An outline of the establishment

of the Entomology Department is given by Johnson (1996) and Logunov (2010a), who pinpoint the

department’s beginnings to the promotion of John R. Hardy (1844-1921) to Senior Assistant Keeper

and Curator of Entomology in January 1908. Prior to this, the insect collections inherited from the

Fig. 1. The author and two of the three cabinets (drawers 21-60) housing the dry, mounted specimens in

the worldwide Dermaptera collection, the Manchester Museum.

MILES C., Dermaptera of the Manchester Museum

Table 3. Number of Dermaptera specimens in the British Collection of the Manchester Museum, by

species.

Family

Genus

Species

No. of specimens

Anisolabididae

Euborellia

anmilipes

Forhculidae

Forficula

auricularia

Forhculidae

Forficula

lesnei

Forhculidae

Apterygida

media

Labiduridae

Labidura

riparia 1

Spongiphoridae

Labia

minor

Spongiphoridae

Marava

arachidis

Undetennined

—

Total

251

1 Collected in France.

Manchester Natural History Society were in such a poor state that it was considered necessary to start

them again from scratch (Report 1890-94).

In 1951 Hinclcs wrote:

“... it has been stated of various orders or groups of insects that they are ‘neglected’. To none is the

epithet more truly applicable than to the order Dermaptera or Earwigs, the serious students of which, at

any one time, could be counted on the fingers of one hancT (Hincks 1951a: 1).

100.00 -

90.00 -

80.00 -

70.00 -

60.00 -

Family

Fig. 2. A graph showing % of World Dermaptera represented at the Manchester Museum, by family.

European Journal of Taxonomy 141: 1-138 (2015)

It is not surprising then that from the first entry in the Entomology Department’s Accession Register

(Volume ‘F’) in 1889 by Hardy until the 1950s, with one exception, there is no specific mention of

Dermaptera acquisitions in the Museum’s Annual Reports or in the Accession Register. The exception is

three specimens of the parasitic Hemimerus talpoides Walker, 1871 from the Gold Coast, West Africa,

donated by Prof Hickson in 1919 (accession no. FI809; see Parasitic Earwigs below). That is not to

say earwigs were not acquired. In the early part of the century it is likely they were included in the

unspecified ‘Insects’ and possibly ‘Orthoptera’ received from around the world, and certainly from the

time of Hincks in the late 40s onwards, Dermaptera started to flood into the Department. The acquisitions

were not, however, entered into the Accession Register as a matter of routine.

Much research would be necessary to identify the sometimes tortuous routes by which many of the

early specimens came into the collection; as described below, many were acquired through exchange,

particularly by Hincks and Brindle. The earliest dated specimen in the Dermaptera collection is an

Apachyus chartaceus (Haan, 1842) dated 1878 (Fig. 5). It was collected during a Dutch scientific

expedition to Sumatra 1877-1879 under the supervision of the President of the Royal Dutch Geographical

Society, Prof. P.J. Veth (Wentholt & Wilschut 2003). Dermaptera material from the expedition was

sent to Auguste de Bormans of Belgium for identification (Veth 1892), considered ‘the only and final

Fig. 3. A drawer of the British Dermaptera collection, the Manchester Museum.

MILES C., Dermaptera of the Manchester Museum

authority’ on Dermaptera at the time (Burr 1901: 85). De Bormans’ collection was acquired by Malcolm

Burr (see below) and later sold to BMNH in the early 1900s (letter from Burr to Hincks, MMUE, Hincks

archive, item 54; Natural History Museum 2015a).

Other specimens dating from the end of the 19th century were collected by Leonardo Fea (1852-1903)

in Burma (now Myanmar), and by Hans Friihstorfer (1866-1922) in Java in 1893 and Lombok, Lesser

Sunda Islands in 1896. Fea was an Italian zoologist who worked at the Genoa Natural History Museum

and made several foreign trips to collect specimens, exploring extensively in Burma in the late 1880s

(Gorman 1892). Friihstorfer was described as “...a purely entomological and systematic genius, the

like of which we have never met before and are not likely to meet again ’ (Martin 1922: 97). In Lombok

he met and worked with William Doherty (1857-1901), “ Prince of collectors ” (Martin 1922: 99), who

collected MMUE’s specimen of Cordax armatus (de Haan, 1842) in Macassar, Indonesia in 1896.

In 1919, Harry Britten (1870-1954) was appointed as Hardy’s successor as Assistant Keeper at

Manchester Museum. According to Hincks, Britten was “the greatest British entomologist since the

days of Curtis and Stephens ” (Hincks 1954a: 225). He particularly added to the extent of almost all

orders of British insects in the collection and improved their arrangement (Report 1923-24). He and

Hincks each provided 50-60 British earwig specimens and between them collected almost 50% of

the specimens in the British collection. Britten’s specimens are almost entirely from Lancashire and

Cheshire, collected between 1919 and 1948, overlapping Hincks’ which date from 1932 to 1951. The

earliest dated specimens are Forficula auricularia Linnaeus, 1758 collected in 1892 in the Fame Islands,

the label only identifying the collector as ‘Bateson’. The most recent specimen (also F. auricularia ) was

collected in 2013 at Manchester University by the current Curator of Arthropods, Dmitri Logunov.

There is little information about the contribution to the Dermaptera collection of Geoffrey Kerri eh (1909—

2003), who followed Britten as Assistant Keeper in 1937. He was seconded away from the museum to

other duties a year after the start of the war and only returned half-time in 1943. His particular interest

was parasitoid Hymenoptera (Noyes 2003). The Annual Report of 1940—41 notes that he began to build

up “... a collection of immature forms in fluid preservative ... ”, although it is not specified whether this

included any earwigs (Report 1940-41: 7).

Fig. 4. Papered material from India collected by PS. Nathan and T.R.S. Nathan between 1953 and 1972.

Arrows indicate Brindle’s identifications (see co mm ents in the text under ‘Alan Brindle’).

European Journal of Taxonomy 141: 1-138 (2015)

Walter Douglas Hincks (1906-1961)

The development of the Dermaptera collection really took off when Hincks (Fig. 6) was appointed

Assistant Keeper in August 1947. When he took over, “ with the aid of his assistant Stanley Shaw, and

later Alan Brindle, the rearrangement and combination of these collections was commenced with the

object of making the Museum s Entomology Department the finest reference and study centre in the

North ’ (Kloet 1961: 183). Hincks developed his interest in entomology as a member of the Leeds

Naturalists’ Club while studying to qualify as a pharmacist (Thimann 1961). He went on to work for a

large firm of manufacturing chemists. Kloet (1961) describes how Hincks developed his expertise first as

Ex coll,

de Bormans.

f JP

Fig. 5. The collection’s earliest dated specimen, Apachyus chartaceus (Haan, 1842), dated 1878, the

Manchester Museum. Scale bar = 1 cm.

MILES C., Dermaptera of the Manchester Museum

a world authority on Passalidae, with his friend John Dibb, and then extended his studies to Orthoptera,

Chironomidae, Diptera, Cassidinae and parasitic Hymenoptera. At the same time he built up a large

private collection and during his life played a most important role in the organisations devoted to

the promotion of Entomological knowledge ” (Kloet 1961: 182). He is well known for the comprehensive

24-volume Check List of British Insects compiled in collaboration with Kloet, published in 1945

(Kitchen 1962). This publication formed the structure for the future layout of the British collections at

the Manchester Museum (Johnson 1996).

Fig. 6. Walter Douglas Hincks (1906-1961), Keeper of Entomology at the Manchester Museum (1947-

1961), photographed in the late 1950s.

European Journal of Taxonomy 141: 1-138 (2015)

Hincks began to focus on the study of Dermaptera in the early 1930s and was encouraged and supported

by Dr. Malcolm Burr (1878-1954), a pioneer in the field who was originally a specialist in European

Orthoptera but became an accepted authority on Dermaptera (Hincks 1954d). Burr trained as a geologist,

but was also a journalist, author and in his later years, an English professor at Istanbul University

(Harvey et al. 1996). The two men kept up a sustained correspondence. 34 letters from the period

January 1933 to October 1951 are held in the Museum’s entomological archive. In January 1933, Burr

wrote, “/ am so delighted that some one [sic], esp. an Englishman, has come forward to carry on the

work [on Dermaptera] where I gave up ” (MMUE, Hincks archive, item 52) and writing to Hincks from

Istanbul in 1950, Burr wrote, “I should like to see you chuck the beetles and go all out on earwigs ”

(MMUE, Hincks archive, item 42). Hincks did make earwigs one of his special areas of study, becoming

a major contributor to the development of the taxonomy of the order. Kloet said of him :

he realised that here lay the greatest scope for research and a task that would test to the utmost his

skill as a systematist and taxonomist and exercise his talent for the investigation of nomenclature and

synonymy. It is rare, indeed, to find a man capable of all these approaches, yet Hincks was undoubtedly

master of them alF (Kloet 1961: 182).

One of the reasons Hincks gave for the lack of progress with Dermaptera taxonomy was that, until Burr,

no-one had enough material to develop a sound classification of the order (Hincks 1951a). In February

1933, Burr exhorted Hincks, “... the chief thing is to get a representative set of as many genera as poss.

You will find my coll, in the B.M. the completest in existence stilF (MMUE, Hincks archive, item 54).

This was backed up by James Rehn (1881-1965), Acting Curator at the Academy of Natural Sciences of

Philadelphia, who wrote on 3 Mar. 1933:

“It may be possible for me in the not distant future to send material of a few species of earwigs which

are included in our duplicate series, but my experience has been much the same as Dr. Burr S, in that

earwigs rarely come to the student in sufficient series to more than supply his working representation ”

(MMUE, Hincks archive, item 56).

Hincks therefore worked assiduously to build up a comprehensive earwig collection and achieved this

by several means. To widen the geographic range of the collection, he requested or encouraged others to

request collectors to provide specimens from particular areas, and at the same time arranged the purchase

of specimens directly from collectors. Correspondence in the Hincks archive, retained at the Manchester

Museum, evidences his network of contacts with those at the forefront in the study of Dermaptera such

as Friedrich Zacher (1884-1961), J.A. Rehn, Alfredo Borelli (1858-1943), Carlo Menozzi (1892-1943)

(Gilbert 1977) and Grigory Bey-Bienlco (1903-1971) (Rentz 2014; MMUE, Hincks archive, items 48,

72, 56, 58, 85, 90, 387). Appendix I provides a list of Hincks’ correspondence specifically relating to

Dermaptera. In 1938 he advertised in the publication Insektenborse to buy Dermaptera material. This

resulted in the purchase of 370 earwigs from Java (supplied by H. Lucht and M.E. Walsh) and 270-80

from Paraguay (supplied by F.H. Schade) (MMUE, Hincks archive, items 45-7, 104-6, 124).

From all this material, he determined and described new species and this taxonomic work led to

scholarly publications. His philosophy was that the department “... should carry out taxonomic and

biogeographical research at the international level, based upon the collections of overseas museums

as well as its own ” (Johnson 1996: 205). In his first busy year as Assistant Keeper, the Annual Report

records that in addition to all his other curatorial duties, “A monograph of the Earwigs of the world has

been initiated and materials accumulated for the purpose .” (Report 1947—48: 11).

Part I of his Systematic Monograph of the Dermaptera of the World based on material at BMNH was

published in 1955 and the second (and last) of the intended six volumes was published in 1959. Fig. 7

shows an example of Hincks’ original monograph illustrations and Fig. 8 shows a page from his

MILES C., Dermaptera of the Manchester Museum

‘Typomap of Africa’. His contribution to the London Royal Entomological Society’s Handbooks for the

identification of British insects on Dermaptera and Orthoptera was published in 1956. By the time of his

death, he had published more than three hundred papers and notes in international, national and regional

journals (Johnson 1996), including more than 70 on Dermaptera.

Hincks’ expertise led to increased requests for identification of material from Public Health Departments,

amateur naturalists and other inquirers and, particularly, to specialised identification work for various

British and overseas museums and institutions. The accepted custom of retaining part of the material

for the Museum led to a rapid increase in the size and scope of the collection (Reports 1953-58). Some

enquiries arose from the accidental importation and discovery of insects in timber and foodstuffs, and

were numerous enough that Hincks saw the need to create a reference collection of such insects. This

Typomap of Africa

IMO

STv

SW OK TR r<

CC NL

Figs 7-8. Two items from Hincks’ archive retained at the Manchester Museum. 7. An illustration of

Pyragraparaguayensis Borelli, 1904 (<$) by Hincks for his Dermaptera monograph (Hincks 1959: 195)

(item 597). 8. A page from Hincks’ ‘Typomap of Africa’ (item 15).

European Journal of Taxonomy 141: 1-138 (2015)

developed from its inception in 1951 (Report 1950-51) to contain 463 identified insects in 7 orders in

2010 (Logunov 2010a); however, virtually all the Dermaptera material in this collection was collected

between 1948 and 1955 and arranged by Hincks (See ‘Description of the Dermaptera collection’ above).

In addition, material was collected on the many field trips, surveys and courses directed by or attended

by Hincks, Shaw, and later, Brindle, both in Britain and abroad, although there is a comment in the

Annual Report for 1956-57 (p. 10) that collecting and other field activities have been purposely

curtailed this year because of the vast number of unidentified specimens which have accumulated in the

department ”. Fig. 9 shows Hincks and Brindle during one of the Insect Surveys at Freshfield, Lancashire.

Six specimens in the British earwig collection resulted from these surveys. Shaw left the department

in May 1956 and Alan Brindle, “ an entomologist of considerable experience” , joined Hincks as his

assistant in 1958 (Report 1958-9: 8).

By the time of his death in 1961, Hincks had become a world authority on Dermaptera, Passalidae and

Chrysomelidae and improved the Manchester collection such that it was “... now the most important

collection in the country after the British Museum and the Hope Department, Oxford, its types and

figured specimens running into many thousands ” (Report 1960-61: 1). Hincks’ personal collection

of World Dermaptera was purchased by the Museum in 1961 (accession no. F2440) along with his

Passalidae and Chrysomelidae collections. The Dermaptera collection was second only to the British

Museum collections and at that time included some 200 ‘ types and paratypes\ i.e., 27% of the type

material held now. It was housed in twenty cabinet drawers “ already arranged ’, the collections being

“... beautifully documented and of great importance for study ” (Report 1961-2: 14, 8, 2).

Fig. 9. Freshfield Insect Survey, June 1959. Left to right: W.D. Hincks, M.W.R. de Vere Graham,

A. Brindle and C.H. Wallace Pugh (MMLJE archive).

MILES C., Dermaptera of the Manchester Museum

Alan Brindle (1915-2001)

Following Hincks’ death, Alan Brindle (Fig. 10) was promoted to Keeper of Entomology, a post he held

until his retirement in 1982. He had started work in a I an cas h ir e woollen mill and at the same t im e

attended night school to learn German, French and Russian. During the war he worked in intelligence

in Pakistan and Bangladesh and after the war taught secondary biology. He developed an interest in

Fepidoptera in his teens, collecting around Pendle Hill, and also took the opportunity to collect snake

skins, butterflies and other insects during his war postings (Johnson 2003). He was elected Fellow of

the Royal Entomological Society in 1941 and awarded an MSc by the University of Manchester for his

work on caddisfly biology in 1968 (Report 2000-01).

Fig. 10. Alan Brindle (1915-2001), Keeper of Entomology at the Manchester Museum (1961-1982),

photographed in the mid 1980s.

European Journal of Taxonomy 141: 1-138 (2015)

According to Johnson (1996), Brindle subscribed to Hincks’ philosophy that entomologists should

become taxonomic specialists in an order or family, within the international scientific community. After

Hincks’ death, Brindle took up and continued Hincks’ work on Dermaptera, publishing his first paper

in 1964 with G. Friese on Albanian earwigs. Prior to this he had already published over 60 papers, the

majority of them on Trichoptera and the larvae of British Tipulinae and British Diptera. Dermaptera took

over and between then and 1994 he published descriptions of a staggering “357 new taxa (321 species,

11 subspecies, 15 genera and 4 subfamilies) in 126papers ” (Johnson C. 2003: 57). His great work, ‘The

Dermaptera of Africa’, was published in two parts in 1973 and 1978 by the Musee Royal de l’Afrique

Centrale, Tervuren, Belgium.

Large numbers of Dermaptera continued to be received for identification from museums and scientific

institutions around the world. Each year hundreds of specimens were identified by Brindle and returned

and, according to custom, it was allowed that some specimens be retained at MMUE. Most of these

were not recorded in the accession register but this sort of work was a regular feature in the Annual

Reports, a typical one from 1965-66 recounting that of the earwigs received from various institutions,

most were new to the collection and “... a good proportion are species new to science ” (Report 1965-66:

14). In 1973 the workload was so heavy that Brindle had to return some Cassidinae unidentified to Dr

Richard zur Strassen of the Research Institute Senckenberg, Frankfurt as “ lam working on Dermaptera

and do not see any chance of working on the Cassidinae for some years to come ” (MMUE, Brindle

archive, item 174). As another example, Brindle was asked to examine approximately 5000 specimens

from Panama, from a collection organised by the Smithsonian Tropical Research Institute. From this

material he described 14 new species (Brindle 1981, 1982a, 1988). His notes state that 802 specimens

of 31 species from this material were retained at MMUE (MMUE, Brindle archive, item 138). The only

earwig in the collection mounted with wings open was part of this consignment (Fig. 11).

Like Hincks, Brindle also sought out specimens from specific areas. Correspondence between Brindle,

PS. Nathan and his daughter-in-law T.R.S. Nathan shows that material from South India was requested

and received from before November 1964 to at least September 1972 (MMUE, Brindle Archive, items

PANAMA, Fortuna

Wet forest,

atdightD 50 m

5 - M. i

Dr.Henk Wolda,S?PI

Fig. 11. The only specimen at MMUE mounted with wings unfurled, Purex remotus (Burr, 1899) from

Panama, the Manchester Museum. Scale bar = 1 cm.

MILES C., Dermaptera of the Manchester Museum

215-234). There are several hundreds of Nathan contributions mounted, determined and incorporated

into the collection, with 1600 remaining as papered material. Brindle’s handwritten notes on some

packets identify genus and in some cases species, the majority of these packets containing mixed species

or genera. Both Hincks and Brindle named species for Nathan - Diplatys nathani Hincks, 1960 and

Gonolabidura nathani Brindle, 1965. Hincks acknowledged that Nathan’s “... expert field work has

provided many Museums with important insect materials from India ” (Hincks 1960a: 15) and Brindle

that his “... untiring efforts in collecting specimens have considerably increased our knowledge of the

Dermaptera of India” (Brindle 1965: 594).

During 1979-80, Dr Seiroku Sakai (1924-2004) visited the Manchester collection on his travels around

the world gathering original descriptions of new Dermaptera species and photographing existing type

specimens for his encyclopaedic ‘Dermapterorum Catalogus: A Basic Survey for Integrated Taxonomy

of the Dermaptera of the World’ (Masaki 2006). He met with Brindle and with Prof. E.J. Popham

of Manchester University to discuss classification (Report 1979-80: 15). They kept in contact and

the Brindle Archive holds Brindle’s working copy of Sakai’s ‘A new proposed classification of the

Dermaptera with special reference to the checklist of the Dermaptera of the world’ (Sakai 1982), with

further corrections and editions, as well as reprints of Sakai’s papers which he sent to Brindle.

Johnson (1996) estimated that during Brindle’s period as Keeper, the Dermaptera collection doubled in

size to around 10,000 specimens. The great numbers of specimens coming in from such identification

requests led to the re-curation of the Dermaptera in three cabinets with sixty drawers (Fig. 1). The first

part of the newly reorganised collection was arranged according to Hincks’ ‘Systematic Monograph

of the World Dermaptera’ and the rest of the collection was arranged according to “... revisions of

certain groups of the earwigs by the Keeper ” (Report 1967-68: 10). The storage of the Dermaptera spirit

collection today is virtually unchanged since Brindle re-curated it during 1972-3 (Report 1972-3).

Brindle retired in 1982, a much respected scientist who had overseen significant development of the

Museum’s entomological collections and gained a “... high reputation in the fields of classification

of world Dermaptera (Earwigs) and taxonomy of Diptera (fly) larvae ”. He was remembered as a “...

kindly, modest and thoughtful man, who wore his knowledge lightly and was always ready to help and

encourage others ” (Report 1981-82: 17).

Since his time, only around 30 specimens have been added to the collection, the most recent being three

specimens collected by Dmitri Logunov in Peru in 2014.

The Entomological Archive

In addition to a library of specialist books and reprints, the Manchester Museum’s Entomology

department holds thousands of documents relating to the Keepers and associated collectors, organised

into 34 collections. The following extracts summarise material relating to Brindle and Hincks (after

Logunov 2010a: 40):

Brindle, Alan 599 [items]. In five boxes. Correspondence with H.G. Briggs, I.M. Robson, H. Robertson,

T. Cekalovic, T.R.S. Nathan, G. Kerrich, and others (1933-84, 165 items); ‘Insect records’ (10 volumes,

1930-82); species records of aquatic insects in the museum collection, plus locality list; reprint collection

(346 items); index records; typed and handwritten lists of insect species; Lepidoptera records; various

keys; address books; visitor book; Diptera larvae catalogue; etc.

Hincks, Walter D. 611 [items]. In five boxes. Original figures for Dermaptera monograph (383 cards);

published papers by Hincks (94 items); reprints Hymenoptera Parasitica J. Lancs. & Chesh. Ent. Soc.,

133 & 134 (2009 & 2010) 41 and others; address and field notebooks (26 items); notes on Dermaptera

European Journal of Taxonomy 141: 1-138 (2015)

and manuscripts on Passalidae; diaries (1948—49, 1956); correspondence with F. Spaeth, J.T. Salmon,

M. Burr, W. Junk, and others (453 items).

Hincks ’ and Brindle’s correspondence relating particularly to the Dermaptera collection is listed in

Appendices I and II. Due to the particular interest of these two curators, the entomology department has

a comprehensive library of their articles and reprints relating to Dermaptera as well as those of other

authors.

Notable earwig specimens in the MMUE collection

Giant Earwig, Labidura herculeana (Fabricius, 1798)

The collection contains two specimens (one male, one female) of the iconic Giant Earwig Labidura

herculeana (Fabricius, 1798) (accession nos. F3233.1871 (male) and F3233.1872 (female)) (Fig. 12).

The species is notable for being the largest known earwig, with one specimen in the Tervuren collections

measuring 78 mm long and some forceps remains suggesting even larger specimens existed (Brindle

1970c). The MMUE specimens were amongst 40 collected during expeditions in 1965-66 and 1967 by

the Musee Royal de l’Afrique Centrale (RMAC), Tervuren, Belgium. The species is also sadly notable

for being recently assessed extinct. Endemic to the island of St. Helena in the South Atlantic Ocean,

several unsuccessful searches have been made for them since and the species was assessed extinct in

August 2014, the last living specimen having been observed in May 1967 (Pryce & White 2014).

Downgraded to a subspecies of L. riparia by Kirby in 1904, in 1970 Brindle reinstated L. herculeana to

a full species (Brindle 1970c) after examining the 40 RMAC specimens. At the time he suggested that

“ ...the fact that this species is only known from this small area of St. Helena, and is not known from

anywhere else in the world, must give rise to some concern about its future status' ’ (Brindle 1970c: 215).

Pryce & White (2014) record that its decline may be due to the removal, for construction, of surface

stones under which the earwigs would burrow, as well as increased predation by rodents and possibly

invertebrates, including spiders and the centipede Scolopendra morsitans Linnaeus, 1758.

Brindle (1970c) also synonymised L. loveridgei Zeuner, 1962 with L. herculeana (Fabricius, 1798).

L. loveridgei had been described from forceps remains and named for the zoologist Arthur Loveridge

(1891-1980) who had moved to live on the island in 1957. Hincks had seen the forceps and also assigned

3 k Helena f Horst Pni^r

PJ*^rr\ . ls-oe> tr

Helen*,

H’Ji , TVrg u fCn

N. Iglfop T ^ n- _

lEj

dct. A.Brindle I96fe

, Ter „ .

CL)

det. A.Brindle I960

Fig. 12. Male and female specimens of Labidura herculeana (Fabricius, 1798), with corresponding data

labels; MMUE, accession numbers F3233.1871 and F3233.1872. Scale bar = 1 cm.

MILES C., Dermaptera of the Manchester Museum

them to L. loveridgei (Brindle 1970c), but was keen to see more material, according to his letter to

Loveridge in June 1960 (MMUE, Hincks archive, item 360, Fig. 13). L. herculeana forceps were found

with bird bones by Philip Ashmole and Douglas Dorward at Prosperous Bay in 1959. Other remains

found by Philip and Myrtle Ashmole in 1995 suggest that some Giant Earwigs lived in seabird colonies

in rocky places before the seabirds were eradicated by predators ( Ashmole & Ashmole 2004).

In 1960, Loveridge wrote to Hincks outlining his intention to search for further L. herculeana remains

at the same spot where the forceps had been found, and wrote to him again six months later, recounting

the unsuccessful results of his expedition (Fig. 14). Loveridge had worked at Manchester Museum in

the time of J.R. Hardy. He said that if he had known Hincks was at MMUE, he would have asked for

the forceps to be deposited there .. where I know the collections are well-cared for ”, rather than at the

British Museum.

Parasitic earwigs

The collection also contains examples of parasitic earwig species. Two of the five species of the family

Arixeniidae are present. There are 15 specimens of Arixenia esan Jordan, 1909 (11 in spirit, 4 dried and

mounted) (Fig. 15) and 8 specimens of Xeniaria jacobsoni (Burr, 1912) (3 in spirit, 5 dried and mounted)

(Fig. 16). Karl Jordan (1861-1959), who described the family Arixeniidae and also A. esau in 1909,

sent 6 specimens of Arixenia (now Xeniaria) jacobsoni to Hincks, stating, “ ...the sender and collector

are now in Paradise, where all good entomologists go ” (MMUE, Brindle archive, item 55) (Fig. 17).

Jordan’s letter was sent from Walter Rothschild’s natural history museum at the small market town of

Tring in Hertfordshire, England. This museum, now part of the Natural History Museum, London, is still

one of the largest private natural history collections ever assembled (Natural History Museum 2015c).

June anrd, i'jgd.

A- T*v* rldfri„ End - „

c/o« Plan LovArlUfTAp

■Milford Trail** R

CIBVEDAM, BonVffPt.

ft#*I* Pr r Lava-rid#* ¥

Thaol you very such ind**d far y#uy blind and iatairejitl-'ng-

I hnd rot Intended at tail* stnru t-Q publish anythlnr

regard in* the axtraordlikflry r*Iff-rarer p* i4ltfa y#U rilMrnVwrwd

In 34 .Hs is ram. Tl*w*var abould you hm nbl* to diacorar any nora

Tirtti trirl on your raturn how# r it night ba pasglbl* to pr-npuisro

a p-npor on th* subject. The ns tier la of 4ft* Interact

■ fid I regard your discovery of oofmldeirritil# Iwpwrtane#, but

hesltat* to speculate without further ■tidnnde.

E yrvm non4 inters*4*ri to l arm that you started your career

in tha Mane be * tor Viia^ur in 4-be days of Roy Tlrrrty* You wauld find

B.any cbnnges h#r* if you. ware Able to visit US again, and p.j

own Pf|#rt»tnt is ¥*ry such mintitt du* 4* n y *vn activities

during tha lfi*t doe rile end. w# now hP¥# conalderelila collections in

nest orders.

1 have spoken to Ur ,-Bl ci v* r an the f phone shout the n1111pad*a

and of aouran he la twit. Ibt#r*st*d „ hut unfortunately Sirs problaaai

just as wa *11 bur* at th# sonant. Bannr ha will write you and

lat you kft-frv tile ■^ssc ■: slor« the Rfitter quits aces..

¥14 ft kind regards and best wish as +

Your#. (tinegrtljr,

*'7

VARNEYS

[5LAHD <IF ST, HELENA

SOUTH ATLANTIC

1.111.i960.

S. D. Hinoka, Esq, ,

Tbs linn 3 l 2 s ter Huaauai,

TIB Uilnnlty,

MnnokeitaE 13, England.

tail -2 i Singka:

TkiH is to &aknotfJ,edgd reoeipi of yeur let’® 1 of

33.Tfl.6-Q on tae gukjBOt of aaiwlg-foraspa fEois St*

Hslaua.

Blnae raturnlng iiexa In August I kftve teen

ooastwttly awaiting la opportunity to gt% m boat to take

me round TO Prospermia Bay Bklon lisa on ta« ena-t aid*

of tkB 1 Bland, rusxa lending 1« difficult 0*1 fig to tie

fravfiling aoUtllBasfc trade Wind*.

Ca HoTfomho? 36 tk I klred a motor launok find tierae

ial andaxa - It ia an lour aid twenty minutea run fror.

Jamestown jatfcy - and 3j:oat 5 kOUE8 aakore la tke

of flkioa 3 Tara -lewfcsd to digging by 3 ox 4 of ui. 3Te

got a lot of bird balok 'But no aartfig foreSpa sniok 1

ifaa moEB aJlietOUB to gfit, I kad fhadS a SlBtek of tiiam

and 5 ho*Bd tk« picture anil aiplaifiod about taaf. to tils

otfiar two wko tasa asoavatisg.

By way of oonB-olati&n I si® eonding you a doson

earwig a wilii I pereoniill7 found ia*idB tha outer kualf*

of Cans GooB9banie« *kl5k tusy antes by niOaliag a kola

and tlen aftMk tae fruit, km S jigiit in calling tkem

by tha nsma glTSft in ^aliiaa, 137&, St. Halena*? , vl*i

^ Torflnula fluTlpen FabElclUB ..... 13, ffitk tdBE la a_

L'utnyja.-.\r : aa aanotftshaleniae Walter 1, found dead in uouaaj

Jo noaach^ae out .!ax Fatrlolue >. 4+1, taken on two dates,

Looelity labels are pxorldad for all 19 inaaota; please

feal fyee to giro a*ay any that are surpiua to your

7*l]UiraMDta,

)v t .■ j--.m.'..-.. .i.. ..^ V-.-- 1 .■. rVr.-_■,_^ YcuTB flinoarau/j

V.**. :■ c* l| J. V Ip,

1 . ! • Jtr, 't ^ i _

Fig. 13. Letter from Hincks to Arthur Love¬

ridge, 23 Jun. 1960 (MMUE, Hincks archive, item

360).

Fig. 14. Letter from Arthur Loveridge to Hincks

relating his unsuccessful search for the Giant

Earwig (MMUE, Hincks archive, item 317).

European Journal of Taxonomy 141: 1-138 (2015)

Figs 15-16. Parasitic earwigs from the collection of the Manchester Museum. 15. Arixenia esau Jordan,

1909. 16. A. jacobsoni Burr, 1912 (now Xeniaria jacobsoni ), syntype. Scale bars = 1 cm.

Jordan had been curator of insects there since 1893. At the date of the letter in 1947, Jordan was 86

years old, having continued as an ‘unofficial worker’ at Tring since 1939 (Johnson K.R. 2003: 314). All

5 specimens of A jacobsoni now at MMUE are labelled ‘Ex. Mus. Tring via Dr. K. Jordan’. Three are

labelled ‘Cotype’ with Burr’s determination labels and dated 1911. Another, collected in 1919 by a Dr

Hacker, was found ‘on bat’ according to the specimen label.

if?

BRITISH MUSEUM (NATURAL HISTORY)

THE ZOOLOGICAL MUSEUM

TRJNG p Janusry 1^7.

HERTFORDSHIRE

i'ckph'mrr 7'riag ^ 5

Dear Mr. HIpks,

I sin a ending you herewith 6 A.ri^enis

jacobsonljfrm the Malay Peninsula*the exact

locality wb s not given with the specimens;

Federated Malay States will have to do* The

sender and the collector are both in Parsdtae

where all good entomologists go,

Youra sincerely

>/ r* ft f

- A

Fig. 17. Tetter from Karl Jordan to Hincks enclosed with 6 ‘ Arixenia jacobsonf specimens (MMUE,

Brindle archive, item 55).

MILES C., Dermaptera of the Manchester Museum

The Arixenia esau specimens come from Gua Payau (Deer Cave), Sarawak, the specimen label stating

that they were found on a mound of bat guano. The wingless Arixeniidae occur in Indonesia, the

Philippines and the Malay peninsula, and are associated with bats of the genus Cheiromeles, feeding

on the bats’ skin and gland secretions (Kocarelc et al. 2013). They occur in large numbers on guano in

caves (Brindle & Oromi 1994). A. esau is not often found actually on flying bats, although it has been

suggested this is how the earwigs transfer to new roosts (Marshall 1977).

The collection also contains 5 of the 11 species of the Hemimeridae (36 specimens, 15 in spirit), which

are ectoparasites of Giant Rats. Hemimerus talpoides Walker, 1871 (Fig. 18) were the earliest recorded

foreign Dermaptera in the Accession Register, in 1919. The family shows adaptations to a parasitic

life with specialized grooves on the legs for close attachment to the host, loss of wings and eyes, and

straight, narrow cerci (Kocarek et al. 2013). In a study by Mogbo & Akunne (2014) in Awka, Nigeria,

Hemimerus talpoides was found to be the most abundant parasite of Cricetomys gambianus rats, with up

to 9 on those examined. These rats are economically important in the area as a source of protein and are

sold for hide. Their keen sense of smell has been used to detect land mines in Tanzania and to diagnose

tuberculosis (Mogbo & Akunne 2014), so just how injurious the parasites are to their hosts may well be

a topic for further research.

_ . :9V^tr

w ,— w w — ~ ^ , " r ~— —

_ Ifc - I

-j- ( g

liuvJ

: i s o<i

I'aApGcles

Fig. 18. The parasitic earwig Hemimerus talpoides Walker, 1871, the Manchester Museum. Scale bar =

1 cm.

European Journal of Taxonomy 141: 1-138 (2015)

Table 4. Number of Dermaptera species and subspecies represented by type specimens at the Manchester

Museum, by family (nomenclature follows Steinmann 1989c, complemented by latest updates from

Engel & Haas 2007).

Number of species and subspecies

% World

Fauna

No. of

specimens

Family

Holotype

only

Holotype +

paratype(s)

Paratype(s)

only

Syntype(s)

only

Total

Anisolabididae

17.8

157

Arixeniidae

20.0

Chelisochidae

5.3

Diplatyidae

12.5

Forficulidae

10.6

129

Karschiellidae

16.7

Labiduridae

7.9

Pygidicranidae

16.8

Spongiphoridae

100

19.6

312

Total

235

283

15.0

734

Type catalogue of the Dermaptera collection at MMUE

At the Manchester Museum, type material is kept within the main collection. In total there are 733

holotypes, allotypes, syntypes and paratypes, representing 283 species (Table 4). This includes 40

holotypes, of which 32 species were described by Brindle and 8 by Hincks.

The following catalogue provides a complete list of the holotypes, syntypes, allotypes and paratypes in

the collection, arranged alphabetically by species name as originally described, in the following format:

Species name, Author (Original genus)

Literature citation for the original description.

Type material

Number, gender and type of specimens, accession number/s, mount.

Type locality

Transcription of holotype locality label.

Locality label/s

Transcription of locality label/s of other type specimens.

Taxonomy

Current nomenclature following Steinmann (1989c), unless otherwise stated, complemented with latest

changes by Engel & Haas (2007) and reference to the checklist of Hopkins et al. (2014).

Remarks

Including nomenclatural changes and any supplementary information from original descriptions and

Brindle’s Type Index Card system.

Also included in a separate section below is material labelled as type material, which is assumed to

represent unpublished ‘manuscript names’ only.

The following notation is used: (!) - new line; (//) - separate specimen label; [ ] - editorial text.

MILES C., Dermaptera of the Manchester Museum

Abbreviations:

BMNH = British Museum (Natural History), UK

MMUE = The Manchester Museum, UK

STRI = Smithsonian Tropical Research Institute

acutangula Hincks, 1955 (Dacnodes)

Dacnodes acutangula Hincks, 1955c: 807, figs 1-2.

Type material

1 5, paratype, F3233.1318, dried, mounted on pin.

Locality label

Musee du Congo / Elizabethville / XII - 1932 / Mile Kerkvoorde.

Taxonomy

Pygidicranidae, Pygidicraninae, Dacnodes acutangula Hincks, 1955.

adjacens Hincks, 1955 (Diplatys)

Diplatys adjacens Hincks, 1955a: 110, fig. 143.

Type material

1 S, paratype, F3233.1291, dried, mounted on card.

Locality label

W. Almora Divn / Kumaon U.P / July 1917, HGC.

Taxonomy

Diplatyidae, Diplatyinae, Diplatys ( Syndiplatys ) adjacens Hincks, 1955.

Remarks

1. Transferred by Steinmann (1986a: 175).

2. The original description makes no reference to any paratype at MMUE.

3. The specimen carries Hincks’ determination label, stating ‘Paratype’.

4. The collector H.G. Champion is named in the description.

5. The locality Kumaun in the description, and Kumaon on the label, are variant spellings for a

mountainous region in Uttar Pradesh, India.

6. Index card entry: UP = United Provinces, now Uttar Pradesh, India.

affinis Hincks, 1947 (Diplatys)

Diplatys affinis Hincks, 1947b: 12, fig. 6.

Type material

1 5, paratype, F3233.1292, dried, mounted on card.

European Journal of Taxonomy 141: 1-138 (2015)

Locality label

Burma ca 200 m. / Washaung, 20 km / East of Myitkyina / 14.VII.34, Malaise.

Taxonomy

Diplatyidae, Diplatyinae, Diplatys (Hypodiplatys) bormansi Burr, 1910.

Remarks

Diplatys affinis synonymised with Diplatys bormansi Burr, 1910 by Hincks (1955a: 124). Transferred

by Steinmann (1986a: 175).

africana Brindle, 1968 {Labia)

(Fig. 50)

Labia africana Brindle, 1968e: 24, figs 13-15.

Type material

1 S, holotype, F3233.1590 (Fig. 50), dried, mounted on card.

Type locality

Nigeria: / Banchi Prov. / Gadan 25.6.57 / P. Blasdale.

Taxonomy

Spongiphoridae, Geracinae, Pseudovostox africanus (Brindle, 1968).

Remarks

Transferred by Brindle (1970g: 60). Hopkins et al. (2014) refer to this species as ‘ africana Steinmann

(1989c: 424) states Brindle’s use of P. africana in 1970g is a misprint for P. africanus. (See also below

under P. africanus meridionalis and P. africanus sudanensis.)

africanus Brindle, 1973 ( Cordax )

Cordax africanus Brindle, 1973f: 306, figs 417—420.

Type material

1 S, paratype, F3233.1681, dried, mounted on card.

Locality label

S. Africa. / R. E. Turner. / Brit. Mus. / 1924-54. // Port St. John, / Pondoland. / Dec. 1923.

Taxonomy

Forficulidae, Opisthocosmiinae, Syntonus africanus (Brindle, 1973).

Remark

Transferred by Steinmann (1993: 250).

africanus Brindle, 1978 ( Osteulcus )

Osteulcus africanus , Brindle, 1978: 194, fig. 349.

MILES C., Dermaptera of the Manchester Museum

Type material

1 §, paratype, F3233.1745, dried, mounted on card.

Locality label

Cameroun / Yaounde / Vadon! / IX.30.

Taxonomy

Forficulidae, Ancistrogastrinae, Osteulcus africanus Brindle, 1978.

africanus meridionalis Brindle, 1970 ( Pseudovostox )

Pseudovostox africanus meridionalis Brindle, 1970e: 265.

Type material

1 5, paratype, F3233.1591, dried, mounted on card.

Locality label

Coll. Mus. Congo / Tanganika: Musosa, 980m. / (a la lumiere) XI-1953 / H. Bomans.

Taxonomy

Spongiphoridae, Geracinae, Pseudovostox africanus meridionalis Brindle, 1970.

Remarks

1. Hopkins et al. (2014) refer to this species as ‘ africana ’. Brindle named the subspecies P. africanus

meridionalis in the original description and subsequently (Brindle 1973f: 136). Steinmann (1989c:

424) has P. africanus meridionalis.

2. Additional information on the index card: Tanzania, Musosa.

africanus sudanensis Brindle, 1970 ( Pseudovostox )

Fig. 53

Pseudovostox africanus sudanensis Brindle, 1970e: 266.

Type material

1 (?, holotype, F3233.1592 (Fig. 53), dried, mounted on card.

Type locality

Sudan: Yirol / 21.V.54 / E. T. M. Reid.

Taxonomy

Spongiphoridae, Geracinae, Pseudovostox africanus sudanensis Brindle, 1970.

Remarks

Hopkins et al. (2014) refer to this species as 'africana’. Brindle named the subspecies P. africanus

sudanensis in the original description and subsequently (1973f: 138). Steinmann (1989c: 424) has P.

africanus sudanensis.

European Journal of Taxonomy 141: 1-138 (2015)

albocincta Brindle, 1966 (Parablandex)

Parablandex albocincta Brindle, 1966g: 232, figs 21-24.

Type material

1 S, paratype, F3233.1339, dried, mounted on card.

Locality label

Madagascar / e Mus / Geneva.

Taxonomy

Pygidicranidae, Blandicinae, Parablandex albocincta Brindle, 1966.

Remark

The original description makes no reference to specimens at MMUE.

apicalis Hincks, 1951 ( Nesogaster )

Nesogaster apicalis Hincks, 1951b: 568, figs 10-11.

Type material

1 5, allotype, F3233.502; 2 33, paratypes, F3233.500-501; 1 §, paratype, F3233.503; dried, mounted

on card.

Locality labels

F3233.500: New Hebrides: / Malekula, / Ounua. / Feb. 1929 / Miss L. E. Cheesman. / B. M. 1929-280.

F3233.501: New Hebrides: / Santo. / viii-ix. 1929 / L. E. Cheesman. /B. M. 1929-537.

F3233.502: New Hebrides: /N. E. Malekula/ vii-1929 / L. E. Cheesman. /B. M. 1929-514.

F3233.503: New Hebrides: / Malekula, / Ounua. / Mar. & Apl. 1929. / Miss L. E. Cheesman. / B. M.

1929-343.

Taxonomy

Spongiphoridae, Nesogastrinae, Nesogaster aculeatus (de Bormans, 1900).

Remarks

1. Nesogaster apicalis synonymised with N. aculeatus (de Bormans, 1900) by Steinmann (1989b: 23).

2. Locality Ounua given as ‘Ounna’ in the original description (possible mistranscription).

aptera Brindle, 1978 ( Geracodes )

Geracodes aptera Brindle, 1978: 67, figs 81-83.

Type material

1 5, paratype, F3233.1444, dried, mounted on card.

Locality label

IFAN / Nimba (Guinee) / Lamotte et Roy / VII - XII 51.

MILES C., Dermaptera of the Manchester Museum

Taxonomy

Anisolabididae, Isolabidinae, Geracodes aptera Brindle, 1978.

araguensis Brindle, 1974 ( Geracides )

Geracides araguensis Brindle, 1974b: 120, figs 11-12, 14,21.

Type material

1 8, paratype, F3233.1531, dried, mounted on card.

Locality label

Venezuela-Aragua/Rancho Grande / 1100m. 14.VIII.73. // Col. J. L. Garcia.

Taxonomy

Spongiphoridae, Cosmogeracinae, Cosmogerax araguensis (Brindle, 1974).

Remarks

Taxonomy follows Steinmann 1989c. Transferred by Brindle (1982a: 36) as Cosmogerax araguensis.

Steinmann (1989b: 100) has C. araguense. Hopkins et al. (2014) has C. araguensis.

arcuatus Brindle, 1968 ( Auchenomus )

Auchenomus arcuatus Brindle, 1968h: 543, figs 5-6, 9.

Type material

1 8, paratype, F3233.1499, dried, mounted on card, without head.

Locality label

Ripang / N. Luzon.

Taxonomy

Spongiphoridae, Sparattinae, Auchenomini, Auchenomus arcuatus Brindle, 1968.

Remark

Index card entry: Philippine Islands.

argentinus Hincks, 1959 (Esphalmenus)

Fig. 36

Esphalmenus argentinus Hincks, 1959: 203, figs 202, 209.

Type material

1 8, holotype, F3233.1341 (Fig. 36); 1 ?, allotype, F3233.1342; 4 88, paratypes, F3233.1343,

F3233.1346-1348; 8 ??, paratypes, F3233.1344-1345, F3233.1349 (2 specimens), F3233.1350

(2 specimens), F3233.1351 (2 specimens); 2 paratypes, gender not specified, F3233.1352; all dried,

mounted on card.

Type locality

Argentine: / Laguna Cazmiril / Tucuman 4500m. / 10. XII. 1947. / Coll. Sacriste.

European Journal of Taxonomy 141: 1-138 (2015)

Locality labels

F3233.1342-1347, F3233.1349-1352: Argentina / Tucuman: Laguna / Cazmiril 4,500m. / 10. XII.1947

(Sacriste).

F3233.1348: Chile - Tarapaca / Arica / Tahapaca. 4800 / 22.2.48. // Coll. / Kuschel.

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus argentinus Hincks, 1959.

ash mo lei Brindle, 1981 ( Capralabis )

Capralabis ashmolei Brindle, 1981: 77, figs 1-2, 4, 7.

Type material

3 88, paratypes, F3233.1433-1435; 2 $ paratypes, F3233.1436-1437; all dried, mounted on card. In

spirit: 1 c?, paratype, F3448.195; 2 5?, paratypes, F3448.196; 1 5, paratype, F3448.197.

Locality labels

F3233.1433-1437: Ecuador/Los Tayos / Main cave / 12.7.1976 /N. P. Ashmole.

F3448.195: 25 Jul 76 / Los Tayos 815 78°12’W 3°06’S // Main cave. Bottom / of 2’ [illegible]. Rotten

wood / Threshold.

F3448.197: same data except number, 811.

F3448.196: 20 Jul 76 / 930 78°12’W 3°06’S // Bottom of 2’ [illegible] shaft. / Under stones in / stream.

Sta. 1.

Taxonomy

Anisolabididae, Anisolabidinae, Capralabis ashmolei Brindle, 1981.

Remarks

The original description refers to two male and four female paratypes at MMUE, rather than three males

and two females; and in spirit, 1 §, same data except 25 Jul. 1976, and 2 § §, in spirit, same data except

2 Jul. 1976. This last date may be a mistranscription of 20 for 2. There is no reference to a male paratype

in spirit.

ater Brindle, 1988 (Pericomus)

Fig. 52

Pericomus ater Brindle, 1988: 48, figs 1-2.

Type material

1 8, holotype, F3233.1571 (Fig. 52); 9 88, paratypes, F3233.1572-1580; 9 $ $, paratypes, F3233.1581-

1589; all dried, mounted on card. In spirit: 7 c??, paratypes, F3448.37.

Type locality

Panama, Fortuna / Wet forest, / at light, 1050m / 28.6.1979 / Dr. Henk Wolda, STRI.

Locality labels

As for type, except for dates: F3233.1576, 28 Jul. 1977; F3233.1586, 1 Sep. 1977; F3233.1587,

F3233.1589, 24 Sep. 1977; F3233.1580, 13 Oct. 1977; F3233.1582, 15 Nov. 1977; F3233.1584, 23 Jun.

1978; F3233.1583, 12 Jul. 1978; F3233.1573, 20 Jul. 1978; F3233.1574, 2 Aug. 1978; F3233.1578, 12

MILES C., Dermaptera of the Manchester Museum

Aug. 1978; F3233.1579,4 Sep. 1978; F3233.1585, 5 Sep. 1978; F3233.1572,20 Sep. 1978; F3233.1575,

6 May 1979; F3233.1577, F3233.1588, 15 Jun. 1979; F3233.1581, 25 Jun. 1979; F3448.37, 1977-1978.

Taxonomy

Spongiphoridae, Pericominae, Pericomus ater Brindle, 1988.

Remarks

1. The original description refers to 10 male and 10 female paratypes.

2. Dates ofF3233.1576 (28 Jul. 1977) and F3233.1581 (25 Jun. 1979) do not match the date range in the

description, 1 Aug. 1977-15 Jun. 1979.

auchenomoides Hincks, 1954 ( Chaetospanici )

Chaetospania auchenomoides Hincks, 1954e: 15, figs 5-7.

Type material

1 S, paratype, F3233.573; 1 §, paratype, F3233.574; dried, mounted on card.

Locality labels

F3233.573-574: C. Sumba / Fokojengo / 22.9.49 / Biihler & Suiter.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania auchenomoides Hincks, 1954.

Remark

Index card entry: Fesser Sunda Islands.

basidentatus Brindle, 1984 (Esphalmenus)

Esphalmenus basidentatus Brindle, 1984: 287, figs 5, 13, 25.

Type material

1 <S, paratype, F3233.1353; 1 §, paratype, F3233.1354; dried, mounted on card.

Locality labels

F3233.1353-1354: Argentina, Chubut / SO. Esquel / Fa Hoya, 800-1350 m / 24.11.1979 Mision

Cientifica Danesa.

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus basidentatus Brindle, 1984.

basilewskyi Brindle, 1966 ( Diplatys )

Diplatys basilewskyi Brindle, 1966e: 30, figs 1, 4.

Type material

1 [c??], paratype, F3233.1293, dried, mounted on card, without forceps.

European Journal of Taxonomy 141: 1-138 (2015)

Locality label

Coll. Mus. Congo / Tanganyika Terr.: Bunduki / Uluguru Mts., moy. Mgeta / 1300 m. 30-IV/2-V-1957.

Taxonomy

Diplatyidae, Diplatyinae, Haplodiplatys basilewskyi (Brindle, 1966).

Remarks

1. Transferred by Steinmann (1986a: 171).

2. Gender is given as $ on the index card and in the original description, but unspecified on the specimen

labels.

beroni Brindle, 1982 ( Diplatys )

Diplatys beroni Brindle, 1982b: 165, figs 2-3.

Type material

1 S, paratype, F3233.1313, dried, mounted on card.

Locality label

Nigeria / Jos Plateau / State. 5.X. 1976 / P. Beron leg.

Taxonomy

Diplatyidae, Diplatyinae, Diplatys beroni Brindle, 1982.

bhutanensis Brindle, 1975 ( Diplatys )

Diplatys bhutanensis Brindle, 1975c: 12, figs 7-8.

Type material

1 3, paratype, F3233.1294; 1 §, paratype, F3233.1295; dried, mounted on card.

Locality labels

F3233.1294-1295: Chimakothi / 1900/2300 22/5 / Nat.-Hist. Museum / Basel - Bhutan / Expedition

1972.

Taxonomy

Diplatyidae, Diplatyinae, Haplodiplatys bhowmiki (Srivastava & Saha, 1975).

Remarks

Diplatys bhutanensis synonymised with Diplatys bhowmiki (Srivastava & Saha, 1975) by Srivastava

(1982: 62). Transferred by Steinmann (1986a: 171).

bhutanensis Brindle, 1975 {Forficula)

Forficula bhutanensis Brindle, 1975c: 43, fig. 45.

Type material

4 S3, paratypes, F3233.1717-1720; 2 paratypes, F3233.1715-1716; dried, mounted on card.

MILES C., Dermaptera of the Manchester Museum

Locality labels

F3233.1715, F3233.1719: Gogona, 3100 m / 10-12.6 // Nat.-Hist Museum / Basel-Bhutan/Expedition

1972.

F3233.1716-1718, F3233.1720: Kotoka-Gogona / 2600-3400 m 10/6 // Nat.-Hist Museum / Basel-

Bhutan / Expedition 1972.

Taxonomy

Forficulidae, Forficulinae, Forficula bhutanensis Brindle, 1975.

Remark

The original description states that three male paratypes from the first locality are at MMUE, not one

male and one female; and two male and two female paratypes from the second locality, not one female

and three males.

bicarinata Hincks, 1947 ( Guanchia )

Guanchia bicarinata Hincks, 1947b: 28, fig. 14.

Type material

1 (?, paratype, F3233.1734; 1 paratype, F3233.1735; dried, mounted on card.

Locality labels

F3233.1734: N. E. Burma / Kambaiti 7000 m / 12/5 1934 R. Malaise.

F3233.1735: As above except date 13 May 1934.

Taxonomy

Forficulidae, Forficulinae, Guanchia bicarinata Hincks, 1947.

bicolor Brindle, 1973 ( Archidux )

Fig. 32

Archidux bicolor Brindle, 1973f: 296, fig. 402.

Type material

1 c?, holotype, F3233.1697 (Fig. 32); 1 §, allotype, F3233.1698; 1 §, paratype, F3233.1699; dried,

mounted on card.

Type locality

Njombe / 6000-6500 ft. / Tanganyika / 20.2.1951 W. R

Locality labels

As type except dates: F3233.1698, 22 Dec. 1951; F3233.1699, 18 Nov. 1951.

Taxonomy

Forficulidae, Skendylinae, Cosmiella bicolor (Brindle, 1973).

Remarks

1. Transferred by Steinmann (1989c: 669).

European Journal of Taxonomy 141: 1-138 (2015)

2. The collection date for the allotype, F3233.1698, is given in the original description as 31.XII. 1957.

From inspection of the label it is probable that 1957 is a mistranscription of 1951.

3. The original description also refers to two paratypes, one male and one female, with the same data,

dates 29 Oct. 1951 and 30 Nov. 1951, respectively.

bicolor Brindle, 1975 ( Brachylabis )

Brachylabis bicolor Brindle, 1975b: 684, fig. 3.

Type material

1 5, paratype, F3233.1466, dried, mounted on card.

Locality label

Tanzanie: Mts Uluguru / Chenzema, alt. 1700m. / 2-22/VII/71.

Taxonomy

Anisolabididae, Isolabidinae, Isolabis bicolor (Brindle, 1975).

Remarks

1. Transferred by Brindle (1978: 46).

2. The original description gives the date as Jun. 1971, not Jul. 1971 as on the specimen label.

bidentata Brindle, 1971 (Strongylopsalis)

Strongylopsalis bidentata Brindle, 1971b: 565, figs 90-92.

Type material

1 S, paratype, F3233.1616, forceps broken at tips; 1 §, paratype, F3233.1617; dried, mounted on card.

Locality labels

F3233.1616-1617: Colombia: D. E. Bogota/El. 8,600 feet / 13-X-1964/ Susan Bates.

Taxonomy

Spongiphoridae, Strongylopsalinae, Strongylopsalis bidentata Brindle, 1971.

Remark

The original description gives the date as 23 Oct. 1964. Both labels are clearly amended to ‘13’ over

typed figures.

bifurcatus Brindle, 1970 (Cordax)

Cordax bifurcatus Brindle, 1970d: 23, figs 13-16.

Type material

1 5, paratype, F3233.1682, dried, mounted on card.

Locality label

Belinga / Gabon. / 23.3.1963 / H. Coiffait.

MILES C., Dermaptera of the Manchester Museum

Taxonomy

Forficulidae, Opisthocosmiinae, Co relax bifurcatus Brindle, 1970.

bilobatus Brindle, 1973 ( Archidux )

Archidwc bilobatus Brindle, 1973f: 299, figs 407—410.

Type material

1 S, paratype, F3233.1700; 1 5, paratype, F3233.1701; dried, mounted on pin.

Locality labels

F3233.1700-1701: 5 to 7 miles into /KeniaForest. /Near FuchiR. / 9-10.2.1911.

Taxonomy

Forficulidae, Skendylinae, Cosmiella bilobata (Brindle, 1973).

Remarks

1. Transferred by Steinmann (1989c: 670).

2. Index card entry: Kenya.

binodosa Brindle, 1967 ( Gonolabina )

Fig. 24

Gonolabina binodosa Brindle, 1967c: 9, figs 3, 5-6, 9, 10.

Type material

1 holotype, F3233.1451 (Fig. 24); 1 §, allotype, F3233.1452; 1 §, paratype, F3233.1453; dried,

mounted on card.

Type locality

Zarate, Peru / Rio Rimac / 9.7.1953 / H. W. Koepcke.

Locality labels

As type locality.

Taxonomy

Anisolabididae, Gonolabininae, Gonolabina binodosa Brindle, 1967.

bintumanensis Brindle, 1971 (Anisolabis)

Anisolabis bintumanensis Brindle, 1971c: 272, figs 10-13.

Type material

1 <S, paratype, F3233.1368; 1 §, paratype, F3233.1369; dried, mounted on card.

Locality labels

F3233.1368: Savane 520 m / piste / [Keionaduga?] / 29.V. 1963 Mission ENS-IFAN / aux Monts Foma

/ Sierra Feone.

European Journal of Taxonomy 141: 1-138 (2015)

F3233.1369: Bintumane/ 1870 m/20.V. 1963. //prairie acyperacees/20.V. 1963. //MissionENS-IFAN

/ aux Monts Loma / Sierra Leone.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis bintumanensis Brindle, 1971.

Remark

The data label for F3233.1368 does not match any in the original description.

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F3233.1298

Figs 19-27. Holotypes (c?) of Anisolabididae and Diplatyidae in the collection of the Manchester

Museum. 19. Gelotolabis hincksi Brindle, 1964 (now Gonolabis hincksi). 20. G. umbrosa Brindle, 1964

(now Gonolabis umbrosa). 21. Gelotolabis tanganyikae Brindle, 1964 (now Gonolabis tanganyikae).

22. Brachylabis meridionalis Brindle, 1978 (now Antisolabis meridionalis). 23. Gonolabina trinodosa

Brindle, 1967. 24. G. binodosa Brindle, 1967. 25. Idolopsalis koepekei Brindle, 1966. 26. Anisolabis

rougemonti Brindle, 1978. 27. Diplatys hainanensis Hincks, 1940 (now Paradiplatys saivazae). Scale

bars = 1 cm.

MILES C., Dermaptera of the Manchester Museum

bituberculata Brindle, 1966 ( Anechura )

Anechura bituberculata Brindle, 1966e: 43, figs 20-22.

Type material

2 SS, paratypes, F3233.1721-1722; dried, mounted on card.

Locality labels

F3233.1721: Recolte dans / prairie seche / avec buissons. // I.R.S.A.C.-Mus. Congo / Kivu: Terr.

Mwenga, sud Fac / Fungwe, 2750 m. Ill-1953 / N. Feleup - G Marlier.

F3233.1722: I.R.S.A.C.-Mus. Congo / Kivu : Terr. Uvira, Ht Fuvubu, 2750 m. (humus) / N. Feleup

V-1954.

Taxonomy

Forficulidae, Forficulinae, Guanchia bituberculata (Brindle, 1966).

Remarks

1. Transferred by Steinmann (1993: 560).

2. In the original description, the locality of the specimen matching F3233.1721 is given as Fac Fongwe,

not Fac Fungwe and date as 1958, not 1953. For F3233.1722, locality given as Ht Fambo in original

description, not Ht Fuvubu as on specimen label.

boliviana Brindle, 1971 ( Strongylopsalis )

Strongylopsalis boliviana Brindle, 197If: 11, fig. 11.

Type material

1 S, paratype, F3233.1618, dried, mounted on card.

Locality label

Quiaca // N. Holmgren.

Taxonomy

Spongiphoridae, Strongylopsalinae, Strongylopsalis excavata Brindle, 1973.

Remarks

1. S. excavata is a nomen novum provided by Brindle following the transfer of Carcinophora boliviana

Bormans, 1903 to Strongylopsalis (Brindle 1973e: 14).

2. Index card entry: Bolivia.

bolivianus Brindle, 1971 {Vostox)

Fig. 40

Vostox bolivianus Brindle, 1971b: 541, fig. 46.

Type material

1 <S, holotype, F3233.1547 (Fig. 40), dried, mounted on card.

European Journal of Taxonomy 141: 1-138 (2015)

Type locality

Bolivia 1100 m / Cochabamba / Alta Palmar / 3-1961 / leg. F. Walz.

Taxonomy

Spongiphoridae, Spongiphorinae, Vostox bolivianus Brindle, 1971.

browni Hincks, 1954 {Labia)

Labia browni Hincks, 1954c: 159, figs 1-2.

Type material

1 3, paratype, F3233.1540, dried, mounted on card.

Locality label

Seychelles / Praslin: / Grande Anse / 26.V.52 / E.S. Brown.

Taxonomy

Spongiphoridae, Labiinae, Spirolabia browni (Hincks, 1954).

Remark

Transferred by Steinmann (1987: 180).

brunneus Brindle, 1966 ( Emboros )

Emboros brunneus Brindle, 1966g: 255, figs 63, 67.

Type material

1 3, paratype, F3233.1683; 1 §, paratype, F3233.1684; dried, mounted on card.

Locality labels

F3233.1683-1684: Perinet // Institut / Scientifique / Madagascar.

Taxonomy

Forficulidae, Skendylinae, Afrocosmia brunnea (Brindle, 1966).

Remark

Transferred by Steinmann (1975: 217).

burri Hincks, 1955 ( Cranopygia )

Cranopygia burri Hincks, 1955c: 812, fig. 4.

Type material

1 3, paratype, F3233.1316, dried, mounted on card.

Locality label

S. India / Madura Dis. / Shembaganur. / Astruc.

MILES C., Dermaptera of the Manchester Museum

Taxonomy

Pygidicranidae, Pygidicraninae, Cranopygia burri Hincks, 1955.

burri Hincks, 1959 ( Echinosoma)

Echinosoma burri , Hincks, 1959: 158, figs 151-153.

Type material

1 c?, paratype, F3233.1320; 2 paratypes, F3233.1321-1322; dried, mounted on card.

Locality labels

F3233.1320-1321: Sumatra / Montagnes pres de / Pajakombo, mai 1905 // Coll. Gadeau de Kerville.

F3233.1322: H. Fucht, Java / K. O Blawan / 2.3.39 / 900/1500 Mr. / Idjen Plateau.

Taxonomy

Pygidicranidae, Echinosomatinae, Echinosoma burri Hincks, 1959.

Remark

The original description refers to only one specimen at MMUE, corresponding to F3233.1322.

burri Borelli, 1909 ( Pseudisolabis )

Pseudisolabis burri, Borelli, 1909: 1.

Type material

1 S, syntype, F3233.1483, dried, mounted on card.

Locality label

Kaschmir.

Taxonomy

Spongiphoridae, Isolaboidinae, Isolaboides burri (Borelli, 1909).

Remarks

1. Transferred by Hincks (1958b: 132).

2. Additional information in the original description: “... raccolti lungo la strada che conduce da

Sirinagur a Gilghit, N. E. del Kashmir; altitudine 2500 metri circa, maggio 1908.”

camposi Borelli, 1907 ( Gonolabis )

Gonolabis camposi Borelli, 1907a: 1, fig.

Type material

1 S, syntype, F3233.1355, dried, mounted on pin.

Locality label

Chiquiboquio / Ecuador / Ex Coll. Borelli.

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus camposi (Borelli, 1907).

Remarks

Transferred by Burr (1909: 252).

canaca Brindle, 1976 ( Anisolabis )

Anisolabis canaca Brindle, 1976b: 70, fig. 3.

Type material

1 <S, paratype, F3233.214; 1 §, paratype, F3233.215; dried, mounted on card.

Locality labels

F3233.214-215: New Caledonia: /P. D. Montague.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis canaca Brindle, 1976.

carinata Hincks, 1959 ( Cranopygia )

Cranopygia carinata Hincks, 1959: 65, fig. 50.

Type material

1 S, paratype, F3233.39; 1 §, paratype, F3233.40; dried, mounted on pin.

Locality labels

F3233.39M0: Maffin Bay / Dutch New Guinea / IV-IX.44 / E. S. Ross Coll.

Taxonomy

Pygidicranidae, Pygidicraninae, Cranopygia carinata Hincks, 1959.

carinata Brindle, 1978 ( Isolabis )

Isolabis carinata Brindle, 1978: 40, figs 33-34.

Type material

1 c?, paratype, F3233.1467; 1 $, paratype, F3233.1468; dried, mounted on card.

Locality labels

F3233.1467-1468: Recolte dans Fhumus // I.R.S.A.C. / S. Rhodesia: Mt Selinda, / foret de Chirinda,

1300 m. / VII. 1960 N. Leleup.

Taxonomy

Anisolabididae, Brachylabidinae, Brachylabis carinata (Brindle, 1978).

Remark

Transferred by Steinmann (1989a: 303).

MILES C., Dermaptera of the Manchester Museum

carinatus Brindle, 1975 ( Spongovostox )

Spongovostox carinatus Brindle, 1975b: 691, figs 10-11.

Type material

1 5, paratype, F3233.1602, dried, mounted on card.

Locality label

Tanzanie: Mts Uluguru / Morogoro Campus Fac / Agric., piege lum. U.V. / alt. 600 m. V-VI/71. // Coll.

Mus. Tervuren / Mission Mts Uluguru / F. Berger, N. Feleup / J. Debecker V/VIII/71.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox carinatus Brindle, 1975.

carinatus Brindle, 1977 {Vostox)

Vostox carinatus Brindle, 1977b: 122, figs 27-28.

Type material

2 <2 <5, paratypes, F3233.1632-1633; 1 §, paratype, F3233.1634; dried, mounted on card.

Locality labels

F3233.1632-1633: Venezuela, Aragua / Rancho Grande / La Trilla. Periquito / 11-1-1975.

F3233.1634: Venezuela, Aragua / Rancho Grande / 1100m. 11-1-74.

Taxonomy

Spongiphoridae, Spongiphorinae, Vostox carinatus Brindle, 1977.

Remark

The original description states that one male and one female paratype at MMUE are from the La Trilla

locality, not two males as in the collection.

cavernicola Brindle, 1979 ( Mixocosmia )

Mixocosmia cavernicola Brindle, 1979: 153, fig. 6.

Type material

1 2, paratype, F3233.1778; 1 5, paratype, F3233.1779; dried, mounted on card.

Locality labels

F3233.1778-1779: Cuevas de la Puente / 3000 m. 20 km. S. / San Francisco, 40 km/E. San Luis Potosi

/ S. L. P. Mexico. 16-V-1972. // W. M. Elliott / R. Ralph / M. McEachem.

Taxonomy

Forficulidae, Skendylinae, Mixocosmia cavernicola Brindle, 1979.

celisi Brindle, 1966 ( Vandex )

Vandex celisi Brindle, 1966c: 46, figs 3—4, 10.

European Journal of Taxonomy 141: 1-138 (2015)

Type material

1 <S, paratype, F3233.1622; 1 §, paratype, F3233.1623; dried, mounted on card.

Locality labels

F3233.1622-1623: Dans bois mort. // Coll. Mus. Congo / Uganda: E. Ruwenzori, / Kalengire, 2000 m.

/ R. P. M. J. Celis 14-1-54.

Taxonomy

Spongiphoridae, Vandicinae, Vandex celisi Brindle, 1966.

ceylonicus Brindle, 1977 ( Nannisolabis )

Nannisolabis ceylonicus Brindle, 1977a: 456, figs 1-3.

Type material

3 <$<$, paratypes, F3233.1485-1487; 1 §, paratype, F3233.1488; dried, mounted on card.

Locality labels

F3233.1485-1488: Ceylon, Central / Kandy: env. 600 m. / 19.1.1970 / C. Beaucheti I. Lobl.

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis ceylonica (Brindle, 1977).

Remarks

Transferred by Steinmann (1989a: 368). Steinmann (1989a) uses both ‘ Antisolabis ceylonicus’ (p.

342) and ‘ Antisolabis ceylonica’ (p. 368). Steinmann (1989a: 368, 1989c: 331) gives Brindle’s original

name for the species as Nannisolabis ceylonica, not Nannisolabis ceylonicus as in Brindle’s original

description (1977a: 456).

cliopardi Hincks, 1947 {Forficula)

Forficula chopardi Hincks, 1947a: 201, fig. 1.

Type material

1 5, paratype, F3233.1723, dried, mounted on card.

Locality label

Museum Paris / Mt. / Bintumani / 1800 m. // Museum Paris / Sierra Leone / P. Jaeger.

Taxonomy

Forficulidae, Forficulinae, Forficula chopardi Hincks, 1947.

Remarks

Steinmann (1993: 596), in the entry for F. chopardi , cites ‘1947 Anechura chopardi Hincks’ as the

original described taxon. This should read ‘ 1947 Forficula chopardi Hincks’. Hincks described Anechura

chopardi in 1958 (Hincks 1958a: 185). Brindle (1973f: 270) transferred Anechura chopardi to Forficula

with the new name F. borellii (F. chopardi Hin cks, 1947 being the senior homonym). In Ste inm a nn

MILES C., Dermaptera of the Manchester Museum

1989c the two species, F. borellii Brindle, 1973 (p. 815) and F chopardi Hincks, 1947 (p. 817) are

clearly distinguished.

clavijoi Brindle, 1974 (Eugerax)

Eugerax clavijoi Brindle, 1974b: 119, fig. 20.

Type material

1 5, paratype F3233.1532, dried, mounted on card.

Locality label

Venezuela-Aragua/Rancho Grande / 1100m. 17.V.73. // J. Salcedo / J. Clavijo.

Taxonomy

Spongiphoridae, Geracinae, Eugerax clavijoi Brindle, 1974.

coalescens Borelli, 1927 ( Chelisoclies )

Chelisoches coalescens Borelli, 1927: 75, figs 6-9.

Type material

3 $ 5, syntypes, F3233.1676, dried, mounted on card.

Locality label

Fort de Kock / (Sumatra) 920 M. / 1925 / leg. E. Jacobson. // $ // Turin Mus. / ex spt.

Taxonomy

Chelisochidae, Chelisochinae, Proreus coalescens (Borelli, 1927).

Remark

Transferred by Hebard (1929: 338).

coiffaiti Brindle, 1970 {Brachylabis)

Brachylabis coiffaiti Brindle, 1970d: 14, figs 1-2, 5.

Type material

1 S, paratype, F3233.1469, dried, mounted on card.

Locality label

Belinga / 46 / H. Coiffait // Mission Biologique / Au Gabon / P P Grasse Directeur.

Taxonomy

Anisolabididae, Isolabidinae, Isolabis coiffaiti (Brindle, 1970).

Remark

Transferred by Brindle (1978: 52).

European Journal of Taxonomy 141: 1-138 (2015)

collinsi Brindle, 1980 ( Brcichylabis )

Brachylabis collinsi Brindle, 1980: 172, figs 7, 9.

Type material

1 S, paratype, F3233.104, dried, mounted on card.

Locality label

Sarawak / Gunong Mulu / National Park / 1978 N. M. Collins / Loc. D.

Taxonomy

Anisolabididae, Brachylabidinae, Brachylabis collinsi Brindle, 1980.

Remark

Additional information in the original description: ‘Dipterocarp Forest 220 m February-March 1978’.

compressa Borelli, 1907 (Anisolubis)

Anisolabis compressa Borelli, 1907b: 3.

Type material

1 8, syntype, F3233.1440, dried, mounted on card.

Locality label

1906. Bimbia/Viag. Ruwenzori, S.A.R.D. Abruzzi.

Taxonomy

Anisolabididae, Anisolabidinae, Euborellia compressa (Borelli, 1907).

Remarks

1. Transferred by Burr (1915: 544).

2. Index card entry: Uganda.

confusus Hincks, 1955 ( Diplatys )

Diplatys confusus Hincks, 1955a: 103, figs 129-130.

Type material

2 88, paratypes, F3233.1296-1297, dried, mounted on card.

Locality labels

F3233.1296: S. India. / Madura Dis. / Shembaganur. / Astruc.

F3233.1297: S. India / Nilgiri Hills / Moyar Camp / 2,600’ 10.4.54 / PS. Nathan.

Taxonomy

Diplatyidae, Diplatyinae, Diplatys (Syndiplatys ) confusus Hincks, 1955.

Remark

Transferred by Steinmann (1986a: 175).

MILES C., Dermaptera of the Manchester Museum

convolutum Hincks, 1959 ( Echinosoma )

Echinosoma convolutum Hincks, 1959: 152, figs 140-141.

Type material

1 S, paratype, F3233.1325, dried, mounted on card.

Locality label

Teinzo / Birmania / Fea, Maggio 1886.

Taxonomy

Pygidicranidae, Echinosomatinae, Echinosoma convolutum Hincks, 1959.

cornutus Brindle, 1973 {Spongovostox)

Spongovostox cornutus Brindle, 1973f: 207, figs 271, 277-278.

Type material

2 S3, paratypes, F3233.1603-1604; 1 §, paratype, F3233.1605; dried, mounted on card.

Locality labels

F3233.1603: Nigeria 10-IX 1966 /Univ. of Ife / Coll. J. L. Libby.

F3233.1604-1605: Dundo, Angola / 18-IX-1971 / A. De B. Machado. // From old termitory / of

Macrotermes bellicosus.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox cornutus Brindle, 1973.

cristata Hincks, 1954 ( Apolabis )

Apolabis cristata Hincks, 1954b: 104, figs 2-5.

Type material

3 33, paratypes, F3233.1370-1372; 2 §§, paratypes, F3233.1373-1374; dried, mounted on card.

Locality labels

F3233.1370-1374: Mt To (1600 m) / Camp 1. / Museum Paris /Nimba (Guinee) / M. Lamotte II.VI.42.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis cristata (Hincks, 1954).

Remarks

1. Transferred by Brindle (1978: 70).

2. In the original description, the date is given as Feb.-Apr. 1942, not Feb. Jun. 1942 as on the specimen

labels.

cristata Brindle, 1976 ( Nesogaster )

Nesogaster cristata Brindle, 1976b: 79, figs 24-25.

European Journal of Taxonomy 141: 1-138 (2015)

Type material

3 paratypes, F3233.524-526; 2 $ §, paratypes, F3233.527-528; dried, mounted on card.

Locality labels

F3233.524: New Caledonia / Mt. Koghi / 27.1.1963. // C. Yoshimoto / & N. Krauss / Light Trap.

F3233.525: New Caledonia / Plateau de / Dogny. // 30. III. 1968 // J. L. Gressitt / Collector / Bishop

Museum.

F3233.526-527: New Caledonia / Plaine des Lacs / area, 6.XI. 1958. // C.R. Joyce / Collector.

F3233.528: New Caledonia / Foret de Thi to /Hamma, 700-800 m / 16.VII. 1958. //Malkin & J. Rageau

/ Collectors.

Taxonomy

Spongiphoridae, Nesogastrinae, Nesogaster cristatus Brindle, 1976.

Remarks

1. Both Steinmann (1989c: 404) and Hopkins et al. (2014) refer to this species as cristatus. Brindle gives

cristata in the original description and on the 1973 determination labels.

2. In the original description, the data for F3233.524 only matches that for one of the specimens described

under ‘Other material examined’ as being at MMUE, a larva.

cunicula Brindle, 1978 ( Anisolabis )

Anisolabis cunicula Brindle, 1978: 113, figs 113-115,239.

Type material

1 S, paratype, F3233.1375, dried, mounted on card.

Locality label

Nigeria Oluduce / S. E. State l.III. 1971 / Col. J. T. Medler.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis cunicula Brindle, 1978.

curta Brindle, 1975 ( Tagalina )

Fig. 38

Tagalina curta Brindle, 1975a: 151, fig. 5.

Type material

1 S, holotype, F3233.60 (Fig. 38); 1 $, allotype, F3233.61; dried, mounted on pin.

Type locality

T.P.N.G. / 17.VI.56 / Tari / Dr. Peters.

Locality label

F3233.61: T.P.N.G. / 17.VI.56/Tari/Dr. Peters.

Taxonomy

Pygidicranidae, Pygidicraninae, Tagalina curta Brindle, 1975.

MILES C., Dermaptera of the Manchester Museum

Remarks

1. Additional information in the original description: locality New Guinea.

2. The collection date is given in the description as 17 Jul. 1956, not 17 Jun. 1956.

decellei Brindle, 1968 {Labia)

Labia decellei Brindle, 1968d: 18, figs 8-10.

Type material

1 c?, paratype, F3233.1606; 1 §, paratype, F3233.1607; dried, mounted on card.

Locality labels

F3233.1606: Coll. Mus. Tervuren / Cote dTvoire : Bingerville / XI. 1962 / J. Decelle.

F3233.1607: As above except date: Jun. 1961.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox decellei (Brindle, 1968).

Remarks

1. Transferred by Brindle (1973f: 195).

2. The original description does not refer to any female paratype examined other than the allotype, the

collection date for which is Jul. 1961 (not Jun. 1961 as for $ paratype F3233.1607).

delicatula Brindle, 1970 {Chaetospania)

Chaetospania delicatula Brindle, 1970d: 19, figs 8-9.

Type material

2 <$<$, paratypes, F3233.1506-1507; 1 5, paratype, F3233.1508; dried, mounted on card.

Locality labels

F3233.1506: Belinga / 22.6.1963 / H. Coiffait // Mission Biologique / Au Gabon / PP Grasse Directeur.

F3233.1507-1508: Same data except date, 22 Mar. 1963.

Taxonomy

Spongiphoridae, Labiinae, Chaetolabia delicatula (Brindle, 1970).

Remark

Transferred by Brindle (1973f: 164).

delicatula Brindle, 1970 ( Gressittolabis )

Gressittolabis delicatula Brindle, 1970f: 687, figs 63-66.

Type material

1 $, paratype, F3233.929; 2 $ paratypes, F3233.930-931; dried, mounted on card.

European Journal of Taxonomy 141: 1-138 (2015)

Locality labels

F3233.929-930: Solomon Is. / Guadacanal [sic] / Mt. Austen / Sept/Oct. 1965.

F3233.931: Bougainville, / Simba Mission / 2-VII-1956 / E. J. Ford.

Taxonomy

Chelisochidae, Chelisochinae, Gressittolabis delicatula Brindle, 1970.

Remarks

E Steinmann (1989c) spells the genus ‘ Gressitolabis ’ and Hopkins et al. (2014) refer to Gressittolabis

as a synonym of Gressitolabis. Brindle named the genus Gressittolabis for Dr J.L. Gressitt in the

original description.

2. The original description gives the date for paratype F3233.931 as 2 Aug. 1965, not 2 Jul. 1956 as on

the specimen label.

dentatus Hincks, 1959 (Esphalmenus)

Fig. 37

Esphalmenus dentatus Hincks, 1959: 205, fig. 210.

Type material

1 3, holotype, F3233.1356 (Fig. 37); dried, mounted on card.

Type locality

Chile / Ex. S. / Kuschel.

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus dentatus Hincks, 1959.

dentatus Brindle, 1988 (Vostox)

Fig. 42

Vostox dentatus Brindle, 1988: 50, figs 7-8.

Type material

1 (f, holotype, F3233.1637 (Fig. 42); 12 33, paratypes, F3233.1638-1649; 13 §§, paratypes,

F3233.1650-1662; dried, mounted on card. In spirit: 18 33, 12 §§, paratypes, F3448.46.

Type locality

Panama, Fortuna / Wet forest / at light, 1050m / 1.9.1976 / Dr. Henk Wolda, STRI.

Locality labels

As fortype, except dates: F3233.1651,28 Sep. 1976; F3233.1643, F3233.1648,11 Oct. 1976; F3233.1646,

F3233.1650, 17 Nov. 1976; F3233.1638, F3233.1652, F3233.1659, 21 Nov. 1976; F3233.1647, 23 May

1977; F3233.1661, 5 Aug. 1977; F3233.1660, 16 Aug. 1977; F3233.1644,25 Spe. 1977; F3233.1658,27

Sep. 1977; F3233.1642, 3 Jun. 1978; F3233.1655, 15 Jul. 1978; F3233.1649, 12 Sep. 1978; F3233.1662,

21 Sep. 1978; F3233.1640-1641, 25 Sep. 1978; F3233.1639, F3233.1656, 11 Dec. 1978; F3233.1657,

13 Nov. 1978; F3233.1653, 3 May 1979; F3233.1645, F3233.1654, 1 Jun. 1979.

MILES C., Dermaptera of the Manchester Museum

F3448.46: Panama : Chiriqui Province / Fortuna. 1050 m. at light / 1977-1978. Dr. Henk Wolda /

Smithsonian Tropical Research Institute / Vostox dentatus Brindle / Paratypes 18 (?, 12 $ / det. A.

Brindle 1980.

Taxonomy

Spongiphoridae, Spongiphorinae, Vostox dentatus Brindle, 1988.

Remark

The original description refers to 12 female paratypes at MMUE, not 13. Dates for all the MMUE

specimens fall within the range given in the description (28.ix. 1976—1.vi. 1979) and the locality data

match.

denticulatum Hincks, 1959 (Echinosoma)

Echinosoma denticulatum Hincks, 1959: 150, figs 136-137.

Type material

1 c?, paratype, F3233.1323; 1 §, paratype, F3233.1324: dried, mounted on card.

Locality labels

F3233.1323-1324: Sumatra / Mont Ophir / Coll. Gadeau / de Kerville.

Taxonomy

Pygidicranidae, Echinosomatinae, Echinosoma denticulatum Hincks, 1959.

Remark

The original description has the spelling ‘Mt. Opphir. (Ophir (also known as Talakmau or Talamau) is

a volcano in West Sumatra, Indonesia.)

diagonalis Brindle, 1982 ( Cosmogerax )

Cosmogerax diagonalis Brindle, 1982a: 37, figs 3-6, 12.

Type material

2 $ J, paratypes, F3233.1674-1675, dried, mounted on card.

Locality labels

F3233.1674-1675: Venezuela, Aragua / El Limon. 450 m. / 10-XII-1975.

Taxonomy

Spongiphoridae, Cosmogeracinae, Cosmogerax diagonalis Brindle, 1982.

Remarks

1. Brindle (1982a: 38) refers to only 1 §, paratype at MMUE.

2. Taxonomy follows Steinmann 1989c. Steinmann (1989b: 100) has Cosmogerax diagonale. Hopkins

et al. have C. diagonalis.

European Journal of Taxonomy 141: 1-138 (2015)

discalis Brindle, 1989 (Cardnop flora)

Carcinophora discalis BrindlQ, 1989: 14, fig. 17.

Type material

1 5, paratype, F3233.1392, dried, mounted on card.

Locality label

Cerro Jele, Prov. Pana. / ma. enraiz de He- / lecho. Rep. Panama. / 12 Sept. 1985 / Col. Grupo Botanica.

Taxonomy

Anisolabididae, Anisolabidinae, Carcinophora discalis Brindle, 1989.

Remark

Not listed in Steinmann (1989c) or Hopkins et al. (2014).

discedens Hincks, 1959 (Bormansia)

Bormansia discedens Yimeks, 1959: 170, fig. 166.

Type material

3 S3, paratypes, F3233.1334, F3233.1335 (2 specimens on one pin); 2 §§, paratypes, F3233.1336;

dried, mounted on card.

Locality labels

F3233.1334-1336: Mozambique / Luabo 1.1957 / Miss P. J. Usher.

Taxonomy

Karschiellidae, Bormansia discedens Hincks, 1959.

Remarks

1. Listed in Steinmann (1989c: 115) and Hopkins et al. (2014) as discendens. Named ‘ discedens ’ in the

original description and on Hincks’ specimen determination labels.

2. The original description gives the locality as Luaba (possible misspelling of Luabo, Mozambique).

distendens Brindle, 1975 {Forficula)

Forficula distendens Brindle, 1975c: 47, figs 54-55.

Type material

1 5, paratype, F3233.1733, dried, mounted on card.

Locality label

20 km S / Thimphu / 2,300 m 18/5 // Nat.-Hist Museum / Basel - Bhutan / Expedition 1972.

Taxonomy

Forficulidae, Forficulinae, Guanchia distendens (Brindle, 1975).

MILES C., Dermaptera of the Manchester Museum

Remarks

1. Transferred by Steinmann (1993: 585).

2. The original description gives the collection date as 31 May 1972, not 18 May 1972.

distincta Brindle, 1975 ( Chaetospania )

Chaetospania distincta Brindle, 1975b: 688, fig. 8.

Type material

1 S, paratype, F3233.1509, dried, mounted on card.

Locality label

Tanzanie: Mts Uluguru / Kiroka, for. heliophile / alt. 725m 27-31/V/71.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania distincta Brindle, 1975.

diversus Hincks, 1953 ( Emboros )

Emboros diversus Hincks, 1953: 378, figs 21-22.

Type material

1 S, paratype, F3233.1685; 1 §, paratype, F3233.1686; dried, mounted on card.

Locality labels

F3233.1685: Madagascar: / Mt. Tsaratanana / 1800 m. Foret de / mousses. X.49 / R. Paulian.

F3233.1686: Same data except altitude 1500 m.

Taxonomy

Forficulidae, Skendylinae, Paracosmiella diversa (Hincks, 1953).

Remark

Transferred by Steinmann (1993: 167).

doesburgi Brindle, 1982 ( Cosmogerax )

Cosmogerax doesburgi Brindle, 1982a: 38, fig. 2.

Type material

1 5, paratype, F3233.1673; dried, mounted on card.

Locality label

Museum Leiden / Suriname Para Distr. / Onverdacht, on light / 7. VII. 1962, 19-22h. / PH. V. Doesburg jr.

Taxonomy

Spongiphoridae, Cosmogeracinae, Cosmogerax doesburgi Brindle, 1982.

European Journal of Taxonomy 141: 1-138 (2015)

dominicana Brindle, 1971 ( Parasparatta )

Parasparatta dominicana Brindle, 1971e: 14, figs 13-14.

Type material

2 SS, paratypes, F3233.1522-1523, dried, mounted on card.

Locality labels

F3233.1522: Dominica / En Haut Jean / XII-7-1964 / P. J. Spangler.

F3233.1523: Dominica / Pont Casse. 1.7 / mi. e. 12III65 / W. W. Wirth. // 65W66ex / decaying heart /

Euterpe palm.

Taxonomy

Spongiphoridae, Sparattinae, Sparattini, Sparatta dominicana (Brindle, 1971).

Remarks

1. Transferred by Steinmann (1989b: 160).

2. Index card entry: West Indies.

3. Brindle (197le: 16) states ‘All the types are in the United States National Museum, except for one

paratype male (Pont Casse) and one paratype female (En Haut Jean), in the Manchester Museum’;

however, on the determination label for specimen F3233.1522, 6 $ ’ has been overwritten with ‘c?’ so

that both are males.

dorsalis Brindle, 1969 ( Acanthocordax )

Fig. 30

Acanthocordax dorsalis Brindle, 1969e: 276, figs 1-2.

Type material

1 S, holotype, F3233.1279 (Fig. 30), dried, mounted on card;

Type locality

New Guinea / Aiyura / 6000 ft. / III. 1945 / B. A. O’Connor // Imp. Inst. Ent. / Coll. No. 10635.

Taxonomy

Forficulidae, Opisthocosmiinae, Acanthocordax dorsalis Brindle, 1969.

ecarinatus Brindle, 1973 (Esphalmenus)

Esphalmenus ecarinatus Brindle, 1973f: 105, figs 133, 135-136.

Type material

1 paratype, F3233.1357; 1 5, paratype, F3233.1358; dried, mounted on card.

Locality labels

F3233.1357-1358: S. Afr. Cape Prov. / Drakensbergen 8 miles / ENE Rhodes / 10.III.51 No. 223.

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus ecarinatus Brindle, 1973.

MILES C., Dermaptera of the Manchester Museum

edentata Brindle, 1988 ( Sarcinatrix )

Fig. 33

Sarcinatrix edentata Brindle, 1988: 55, figs 3-5.

Type material

1 (?, holotype, F3233.1747 (Fig. 33); 1 $, paratype, F3233.1748; dried, mounted on card.

Type locality

Panama, Fortuna / Wet forest, / at light, 1050m / 12 Sept. 1977 / Dr. Henk Wolda, STRI.

Locality label

F3233.1748: As type locality except date, 28 Nov. 1978.

Taxonomy

Forficulidae, Ancistrogastrinae, Sarcinatrix edentata Brindle, 1988.

elongata Brindle, 1978 (Anisolabis)

Anisolabis elongataBrmdlQ, 1978: 103, figs 205-207, 278.

drt W. p.Hl ntks

det. A. Brindle

F 3233.1743

N. fevw ■

, CM-erVo ZM-OQm.

,■&. V \ St.

Vi .\fi cy v a(A.tk,

i Kcd

r ferine*-

del,. A. Brindle 1961

noIlJyiK.

F3233.1793

**■

HAINAN.

! We 4.0 A if-mv

.,■ 111111(111 *■***■■ VI** + T

G, Ros igg....

F3233.1797

NJOMBE

6000-6^00 ft.

TANGANYIKA

\V.P,

Afe-b<<4iix e?

det. A. Brindle 197 v>

F3233.1697

"j?!* J Jfll ISA Iv,

/? gi

^ P eg 7^6-

f" '■ 'y-vwt ^

B A .O'Con nor *

IMF. INST, ENT.

coll.no. /<? is S'

det. A.Brindle 196^

hesl*^,^ f

PANAMA, Fortuna

Wet forest*

Dr. BeiAtCVtilfia/SiRI

det. A. Brindle 197JT

hole* Fy'pc-

F3233.1747

Figs 28-33. Holotypes (c?) of Forficulidae in the collection of the Manchester Museum. 28. Metresura

rugosa Brindle, 1975. 29. Timomenus pieli Hincks, 1940. 30. Acanthocordax dorsalis Brindle, 1969

(F3233.1279). 31. Skalistesperuviana Brindle, 1970.32. Archidnx bicolor Brindle, 1973 (now Cosmiella

bicolor). 33. Sarcinatrix edentata Brindle, 1988. Scale bars = 1 cm.

European Journal of Taxonomy 141: 1-138 (2015)

Type material

1 S, paratype, F3233.1376, dried, mounted on card. 1 §, paratype, F3233.1377, dried, mounted on pin.

Locality labels

F3233.1376: Br. Cameroons / Sappo. nr. Buea/Mar. 1951; S. Tita.

F3233.1377: Br. Cameroons / Sasse-Sappo-Buea / Mar. 1952; S. Tita.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis elongata Brindle, 1978.

emcirginata Brindle, 1977 ( Marava )

Marava emarginataAdvmdlQ, 1977b: 124, figs 30-32.

Type material

1 S, paratype, F3233.1548, dried, mounted on card.

Locality label

Venezuela - Aragua / Rancho Grande / 1100 m 20-V-75.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava emarginata Brindle, 1977.

Remark

The original description gives the date of the MMUE specimen as 29 Jan. 1967, not 20 May 1975 as on

the specimen label. This date does not match any in the description.

erythrocephala Brindle, 1964 ( Gelotolabis )

Gelotolabis erythrocephala Brindle, 1964: 753, figs 7, 14.

Type material

1 S, paratype, F3233.1378; dried, mounted on pin.

Locality label

Coll. Mus. Congo / Tanganyika Terr. : Kili- / manjaro. Vers. Sud. 4200- / 4600 m. 18/20-VII-1957.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis felix felix Burr, 1907.

Remark

Gelotolabis erythrocephala was synonymised with Anisolabis felix felix Burr, 1907 by Brindle (1978:

127).

excavatus Hincks, 1954 (Spongovostox)

Spongovostox excavatus Hincks, 1954b: 119, figs 21-22.

MILES C., Dermaptera of the Manchester Museum

Type material

1 S, paratype, F3233.1608, dried, mounted on card.

Locality label

IFAN / Nimba (Guinee) / Famotte et Roy / VII XII 51.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox excavatus Hincks, 1954.

Remark

Index card entry: Guinea, Nimba Mts.

felix elgonensis Brindle, 1978 {An is old bis)

Anisolabis felix elgonensis Brindle, 1978: 128.

Type material

1 S, paratype, F3233.1379, dried, mounted on card.

Locality label

Mount Elgon / in rotten tree / 16.1.1955 / leg. M. W. & A. T.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis felix elgonensis Brindle, 1978.

flaviceps Brindle, 1975 ( Metresura )

Metresura flaviceps Brindle, 1975d: 85.

Type material

1 S, paratype, F3233.1739, dried, mounted on pin.

Locality label

Costa Rica. / Aug. 1935. / EQ. A32140.

Taxonomy

Forficulidae, Neolobophorinae, Metresura flaviceps Brindle, 1975.

flavicornis Hincks, 1947 ( Cosmicula )

Cosmicula flavicornis Hincks, 1947b: 40, figs 22-23.

Type material

1 S, paratype, F3233.1712, dried, mounted on card.

Locality label

N. E. Burma / Kambaiti, 1800 m / 11/6.1934 Malaise.

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Forficulidae, Skendylinae, Cosmiellaflavicornis ( Hincks, 1947).

Remark

Transferred by Steinmann (1989c: 671).

flavipes Brindle, 1975 ( Metresura )

Metresura flavipes Brindle, 1975d: 85.

Type material

1 S, paratype, F3233.1740, dried, mounted on pin.

Locality label

San Pedro de / Montes de Oca / C. R. 9-4-33 / C. H. Ballou.

Taxonomy

Forficulidae, Neolobophorinae, Metresura flavipes Brindle, 1975.

Remarks

1. Not listed in Hopkins et al. (2014).

2. The locality is given as Costa Rica in the original description.

flavocincta Brindle, 1982 {Spongovostox)

Spongovostoxflavocincta Brindle, 1982a: 40, figs 16-17.

Type material

2 $ J, paratypes, F3233.1614-1615, dried, mounted on card.

Locality labels

F3233.1614: Panama, Fortuna / Wet forest, / at light, 1050m / 17 Sept. 1977 / Dr. Henk Wolda, STRI.

F3233.1615: Same data except date 2 Aug. 1977.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox flavocinctus Brindle, 1982.

Remark

Changed to ‘ flavocinctus ’ by Steinmann (1989b: 347).

flavohumeralis Brindle, 1988 ( Marava )

Fig. 45

Marava flavohumeralis Brindle, 1988: 52, fig. 16.

Type material

1 3, holotype, F3233.1799 (Fig. 45); 1 ?, allotype, F3233.1800; 6 S3, paratypes, F3233.1801-1806;

6 $ 5, paratypes, F3233.1807-1812; dried, mounted on card.

MILES C., Dermaptera of the Manchester Museum

Type locality

Panama, Fortuna / Wet forest / at light, 1050m / 24.6.1978 / Dr. Henk Wolda, STRI.

Locality labels

As type locality except dates: F3233.1806, 11 Oct. 1976; F3233.1812, 11 May 1977; F3233.1810, 8 Jul.

1977; F3233.1805, 21 Aug. 1977; F3233.1808, F3233.1809, 24 Sep. 1977; F3233.1800-1802, 17 May

1978; F3233.1804, 10 Jun. 1978; F3233.1811, 9 Jul. 1978; F3233.1807, 25 Sep. 1978; F3233.1803, 14

Oct. 1978.

Taxonomy

Spongiphoridae, Spongiphorinae, Maravaflavohumeralis Brindle, 1988.

Remark

The number of paratypes does not match that given in the original description, which states “Paratypes

(all same locality): 8 8 ?, ll.x.1976 - 21.xi.1978 (MM)”.

formosanum Hincks, 1959 ( Echinosoma )

Echinosoma formosanum Hincks, 1959: 151, figs 138-139.

Type material

1 S, paratype, F3233.1326: dried, mounted on card.

Locality labels

Tainan, / Formosa IV. // Echinosoma / formosanum / Hincks / Paratype 3.

Taxonomy

Pygidicranidae, Echinosomatinae, Echinosoma formosanum Hincks, 1959.

fulvescens Hincks, 1953 ( Chaetospanici )

Chaetospaniafulvescens Hincks, 1953: 373, figs 15-16.

Type material

2 $ J, paratypes, F3233.1510—1511, dried, mounted on card.

Locality labels

F3233.1510-1511: Madagascar CE / Perinet (parapluie) / XI. 1946 J. Millot.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospaniafulvescens Hincks, 1953.

fusca Brindle, 1966 ( Brachylabis )

Brachylabis fusca Brindle, 1966e: 39, fig. 13.

Type material

1 paratype, F3233.1470; 1 §, paratype, F3233.1471; dried, mounted on card.

European Journal of Taxonomy 141: 1-138 (2015)

Locality labels

F3233.1470: Foret transition // Col. Mus. Congo / Tanganyika Terr.: Uluguru / Mts., vail Ululu-Ndogo,

/ 1500 m. 8-V-1957.

F3233.1471: Foret de montagne // Col. Mus. Congo / Tanganyika Terr.: Uluguru / Mts., sommet du

Kidunda/ 1800-1950 m. 3-V-1957.

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis fusca (Brindle, 1966).

Remarks

1. Transferred by Steinmann (1989a: 344).

2. In the original description only one set of locality data is given, as for F3233.1470.

fuscata clavata Brindle, 1972 ( Chaetospania )

Chaetospaniafuscataclavata Brindle, 1972c: 148, figs 13e-13f, 13j.

Type material

2 <$<$, paratypes, F3233.602-603; 1 §, paratype, F3233.604: dried, mounted on card.

Locality labels

F3233.602-604: Pilgo River / Guam / 26 May 1945 // G. E. Bohart / J. C. Gressit.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania fuscata clavata Brindle, 1972.

fuscata fuscata Brindle, 1972 ( Chaetospania )

Chaetospania fuscata fuscata Brindle, 1972c: 146, figs 13c-13d, 13k.

Type material

1 S, paratype, F3233.599; 1 §, paratype, F3233.600; dried, mounted on card.

Locality labels

F3233.599-600: Kusaie. 580m / Mt. Matante / 11 Feb. 1953 / J. F. G. Clarke.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania fuscata fusca ta Brindle, 1972.

Remark

Index card entry: Indonesia.

fuscula Brindle, 1970 ( Brachylabis )

Brachylabis fuscula Brindle, 1970d: 15, figs 3—4, 6.

Type material

1 3, paratype, F3233.1472, dried, mounted on card.

MILES C., Dermaptera of the Manchester Museum

Locality label

Belinga / 109T / H. Coiffait // Mission Biologique / Au Gabon / P.P. Grasse Directeur.

Taxonomy

Anisolabididae, Brachylabidinae, Brachylabis fuscula Brindle, 1970.

gariazzoi var. tanganyikae Hincks, 1957 ( Forcipula )

Forcipula gariazzoi var. tanganyikae Hincks, 1957a: 42.

Type material

1 <S, paratype, F3233.1492; 1 paratype, F3233.1493; dried, mounted on pin.

Locality labels

F3233.1492: Baie de Kasoje / Riv. Lubulungu / 15.XII.46 / Miss. Tanganika 18.

F3233.1493: Same data except date 15 Dec. 1946.

Taxonomy

Labiduridae, Labidurinae, Forcipula tanganyikae Hincks, 1957.

Remarks

1. Transferred by Steinmann (1980: 235).

2. The original description has ‘ gariazzoi not ‘ gariazzi ’ as in Borelli’s original description of Forcipula

gariazzi (Borelli, 1900: 1).

3. Index card entry: Tanzania.

gnathonica Brindle, 1970 ( Chaetospania )

Chaetospania gnathonica Brindle, 1970f: 673, figs 28-29.

Type material

1 S, paratype, F3233.597, dried, mounted on card.

Locality label

Forest litter / Solomon Is. / Kolombangara / Nr. Kuzi / 3-9.ix. 1968 / PN. Fawrence / Roy. Soc. Exp. /

B. M. 166-1.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania gnathonica Brindle, 1970.

Remark

In the original description, the paratype at MMUE is stated to be a female. Brindle’s determination label

states S-

gracilis Brindle, 1966 (Allostethus)

Allostethus gracilis Brindle, 1966h: 442, fig. 6.

European Journal of Taxonomy 141: 1-138 (2015)

Type material

1 S, paratype, F3233.298, dried, mounted on card.

Locality label

Momungan / Mindanao.

Taxonomy

Labiduridae, Allostethinae, Allostethus gracilis Brindle, 1966.

Remarks

1. The original description gives the locality as Philippine Islands.

2. Steinmann (1989a: 412) has the taxon as Allostethus gracile.

gracilis Brindle, 1988 ( Marava )

Fig. 47

Marava gracilis Brindle, 1988: 52, figs 20-2F

Type material

1 (?, holotype, F3233.1825 (Fig. 47); 1 <$, paratype, F3233.1826; 5 paratypes, gender unspecified,

F3233.1827-1831; dried, mounted on card.

Type locality

Panama, Barro / Colorado Is., S-M / light trap, ground / level 8.11.1978 / Dr. Henk Wolda, STRI.

Locality labels

As for type except date: F3233.1827, 4 Nov. 1978; F3233.1828, 7Nov. 1978; F3233.1829, 9Nov. 1978;

F3233.1830, 13 Nov. 1978; F3233.1826, 20 Nov. 1978; F3233.1831, 4 Dec. 1978.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava gracilis Brindle, 1988.

Remarks

1. Not listed in Hopkins et al. (2014).

2. The original description refers to three male and four female paratypes at MMUE (i.e., one more than

listed here) with the date range, 15 Jul. 1978M Dec. 1978.

grandicollis Hincks, 1954 ( Antisolabis )

Antisolabis grandicollis Hincks, 1954b: 117, figs 19-20.

Type material

1 5, paratype, F3233.1473, dried, mounted on card.

Locality label

IFAN / N im ba (Guinee) / Lamotte et Roy / VII. XII. 51.

Taxonomy

Anisolabididae, Isolabidinae, Isolabis grandicollis (Hincks, 1954).

MILES C., Dermaptera of the Manchester Museum

Remarks

1. Transferred by Brindle (1978: 46).

2. Index card entry: Nimba Mts.

greensladei Brindle, 1970 ( Brachylabis )

Brachylabis greensladei Brindle, 1970f: 659, figs 17-18.

Type material

1 S, paratype, F3233.112; 1 §, paratype, F3233.105; dried, mounted on card.

Locality labels

F3233.112, F3233.105: Solomon Is. / Guadacanal [sic] / Mt. Austen / 24.VIII.1965 / P. Greenslade.

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis greensladei (Brindle, 1970).

Remarks

1. Transferred by Steinmann (1989a: 359).

2. The original description states that only 1 male paratype was passed to MMUE. F3233.112 bears

Brindle’s 1970 determination label stating ‘Paratype’.

3. The locality, spelled ‘Guadacanal’ on the locality label, is given as ‘Guadalcanal’ in the description.

griveaudi Brindle, 1966 ( Marava)

Marava griveaudi Bundle, 1966g: 248, figs 47^18, 55.

Type material

1 S, paratype, F3233.1549; 1 $, paratype, F3233.1550; dried, mounted on card.

Locality labels

F3233.1549: Sakarana / Lambomakandro / AR.

F3233.1550: Sakarana / Lambomakandro / P. Griveaud.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava griveaudi Brindle, 1966.

guatemalensis Brindle, 1973 ( Geracides )

Geracides guatemalensis Brindle, 1973a: 154, figs 1—4.

Type material

1 S, paratype, F3233.1530, dried, mounted on card.

Locality label

Guatemala / 39 4347 / Primavera logs / N. York / Mar. 29 1939.

Taxonomy

Spongiphoridae, Cosmogeracinae, Cosmogercoc guatemalensis (Brindle, 1973).

European Journal of Taxonomy 141: 1-138 (2015)

Remarks

Taxonomy follows Steinmann 1989c. Transferred by Brindle (1982a: 37) as Cosmogerax guatemalensis.

Steinmann (1989b: 104) has C. guatemalense . Hopkins et al. (2014) have C. guatemalensis.

gurneyi Brindle, 1970 (Setocordax)

Setocordax gurneyi Brindle, 1970a: 99, fig. 1.

Type material

1 S, paratype, F3233.1754, dried, mounted on pin.

Locality label

Colombia / with orchids / at Miami, Fla / Aug. 7, 1963 / 63 18995.

Taxonomy

Forficulidae, Neolobophorinae, Setocordax gurneyi Brindle, 1970.

hainanensis Hincks, 1940 ( Diplatys )

Fig. 27

Diplatys hainanensis Hincks, 1940: 30, figs 1-3.

Type material

1 S, holotype F3233.1298 (Fig. 27), dried, mounted on card.

Type locality

Hainan, Sama/ 18.VII.36 / G. Ros. leg.

Taxonomy

Diplatyidae, Diplatyinae, Paradiplatys salvazae (Burr, 1917).

Remarks

Hincks (1955a: 94) synonymised Diplatys hainanensis Hincks, 1940 with D. salvazae Burr, 1917.

Transferred to Lobodiplatys {Paradiplatys) salvazae (Burr, 1917) by Steinmann (1986a: 178). The

combination Paradiplatys (.Paradiplatys ) salvazae (Burr) was reinstated by Engel & Haas (2007: 20).

halli Hincks, 1949 ( Nesogaster)

Nesogaster halli Hincks, 1949b: 202, figs 1-3.

Type material

1 S , paratype, F3233.541; 1 §, paratype, F3233.542; dried, mounted on card.

Locality labels

F3233.541: In house. / Titirangi / March 1944 / M. W. Carter.

F3233.542: Sieved. /Mt. Albert. / Auckland. / 24th Sep. 1941. /D. Spiller. Coll.

Taxonomy

Spongiphoridae, Nesogastrinae, Nesogaster halli Hincks, 1949.

MILES C., Dermaptera of the Manchester Museum

Remarks

1. Index card entry: New Zealand.

2. Data in the original description which matches that for specimen F3233.542 has the date 29 Sep.

1941, not 24 Sep. 1941.

hamata Brindle, 1978 (Anisolabis)

Anisolabis hamata Brindle, 1978: 132, figs 201-204, 272.

Type material

1 5, paratype, F3233.1380, dried, mounted on card.

Locality label

Dinsha, Bale / Ethiopia / 1971 / G. De Rougemont.

Taxonomy

Anisolabididae, Anisolabidinae, Flexiolabis hamata (Brindle, 1978).

Remark

Transferred by Steinmann (1989a: 36).

hamatus Brindle, 1972 ( Diplatys )

Diplatys hamatus Brindle, 1972a: 281, figs 1-2.

Type material

1 S, paratype, F3233.1301, dried, mounted on card.

Locality label

Costa Rica / Turrialba / VII-15-19-65 / R J. Spangler.

Taxonomy

Diplatyidae, Diplatyinae, Haplodiplatys hamatus (Brindle, 1972).

Remark

Transferred by Steinmann (1974: 201).

hebardi Hincks, 1957 ( Echinosoma )

Echinosoma hebardi Hincks, 1957b: 55, figs 17-18.

Type material

1 S, paratype, F3233.1327, dried, mounted on card.

Locality label

Tanganyika / Southern Prov. / Nachingwea / 3.XI.53 - III.54 / V. F. Eastop.

Taxonomy

Pygidicranidae, Echinosomatinae, Echinosoma hebardi Hincks, 1957.

European Journal of Taxonomy 141: 1-138 (2015)

heliconia Brindle, 1985 {Labia)

Labia heliconia Brindle, 1985: 79, figs 10, 15, 22.

Type material

1 S, paratype, F3233.1545; 1 §, paratype, F3233.1546; dried, mounted on card.

Locality labels

F3233.1545: Ecuador: Prov. Napo / Lago Agrio. in / Heliconia 26.9.1975 / Andrea Langley.

F3233.1546: Same data except date 23 Aug. 1975.

Taxonomy

Spongiphoridae, Labiinae, Paralabella heliconia (Brindle, 1985).

Remark

Transferred by Steinmann (1989c: 583).

hincksi Ramamurthi, 1960 ( Auchenomus )

Auchenomus hincksi Ramamurthi, 1960: 35, figs 2-3.

Type material

1 S, 1 5, paratypes, F3233.1500, dried, mounted on card.

Locality label

India: Madras / B. N. Ramamurthi / ( f gen. Removed)

Taxonomy

Spongiphoridae, Sparattinae, Auchenomini, Auchenomus hincksi Ramamurthi, 1960.

hincksi Brindle, 1964 ( Gelotolabis )

Fig. 19

Gelotolabis hincksi Brindle, 1964: 753, fig. 10.

Type material

1 S, holotype, F3233.1381 (Fig. 19); 1 5, allotype, F3233.1382; dried, mounted on pin.

Type locality

South Kulal, / Aug. 1944 / J. Adamson.

Locality label

F3233.1382: As type locality.

Taxonomy

Anisolabididae, Anisolabidinae, Gonolabis hincksi (Brindle, 1964).

Remark

Transferred by Steinmann (1978: 196).

MILES C., Dermaptera of the Manchester Museum

hincksi Brindle, 1970 ( Pseudovostox )

Fig. 54

Pseudovostox hincksi Brindle, 1970e: 259, figs 1-2, 10.

Type material

1 3, holotype, F3233.1593 (Fig. 54), dried, mounted on card.

Type locality

V. Falls Rd. 148 / S. Rhodesia m. // XI. 1952 / Nat. Museum / S. Rhodesia.

Taxonomy

Spongiphoridae, Geracinae, Pseudovostox hincksi Brindle, 1970.

hincksi Brindle, 1966 ( Vandex )

Vandex hincksi Brindle, 1966c: 44, figs 5-6, 9.

Type material

3 33, paratypes, F3233.1624-1626; 2 5?, paratypes, F3233.1627-1628; dried, mounted on card.

Locality labels

F3233.1624: Coll. Mus. Congo /Ruanda : Rutovu, for. du /Rugege, 2350 m. /R Basilewsky 20/23-1-53.

F3233.1625-1627: Recolte dans / l’humus / (foret Hagenia). // I.R.S.A.C.-Mus. Congo / Kivu : T.

Kabare, contr. / S. E. du Kahuzi, / 13-VIII-1951 /N. Feleup.

F3233.1628: Recolte dans / l’humus. // I.R.S.A.C.- Mus. Congo / Kivu : Terr. Fubero / 2.200 m. 27-XI-

1951 /N. Feleup / (for. mont. avec bamb.).

Taxonomy

Spongiphoridae, Vandicinae, Vandex hincksi Brindle, 1966.

Remarks

1. Index card entry: Africa, Zaire, Ruanda, Burundi borders, Kivu Province.

2. With regard to the locality of F3233.1624, Brindle (1966c: 46) states ‘This species was incorrectly

recorded by the late Dr. W.D. Hinclcs in 1955 as V schubotzi, on a series from the Rugege Forest,

Ruanda. Only a pair from this series is now available, but these clearly belong to the present species.’

humicola Brindle, 1966 ( Brachylabis )

Brachylabis humicola Brindle, 1966e: 37, figs 11, 19.

Type material

1 3, paratype, F3233.1366; 1 5, paratype, F3233.1367; dried, mounted on card.

Locality labels

F3233.1366-1367: I.R.S.A.C.-Mus. Congo / Kivu: Terr. Kabare, S. E. / Kahuzi, 2080/2200 m. / N.

Feleup VIII-1951.

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Anisolabididae, Isolabidinae, Africolabis humicola (Brindle, 1966).

Remark

Transferred by Brindle (1978: 25).

inca Brindle, 1970 ( Neolobophora )

Neolobophora incadBvmdle, 1970a: 102, fig. 11.

Type material

1 <S, paratype, F3233.1752; 1 paratype, F3233.1753; dried, mounted on card.

Locality labels

F3233.1752: Torentoy Canyon / Base Machu. Picchu / 2000-2200m. Peru / VI—VII. 1964. Malkin.

F3233.1753: Machu-Picchu (sobre / ruinas, 2600-2800m) / Peru - 1-2.VII. 1964 / B. Malkin.

Taxonomy

Forficulidae, Neolobophorinae, Neolobophora inca Brindle, 1970.

incisa Brindle, 1970 (Adiathella)

Adiathella incisa Brindle, 1970f: 690, figs 59-60.

Type material

1 S, paratype, F3233.924; 1 §, paratype, F3233.925; dried, mounted on card.

Locality labels

F3233.924: Guadacanal [sic] / Papamanaisiu / 5700’ / 8-XI-1965 / R. S. Exp.

F3233.925: Forest litter / Solomon Is. / Guadalcanal / Popamanisiu / c 7000’. // 6 - 8. XI. 1965 / P. N.

Lawrence / Roy. Soc. Exped. / Brit. Mus. 1966-1.

Taxonomy

Chelisochidae, Chelisochinae, Adiathella incisa Brindle, 1970.

Remarks

1. It is unclear whether F3233.925 is the allotype referred to in the original description, which only refers

to one male paratype at MMUE. The allotype locality data in the description matches F3233.925.

2. The locality ‘Guadacanal’ on the label for F3233.925 is given as ‘Guadalcanal’ in the description.

incisiodes Brindle, 1978 {An iso la bis)

Anisolabis incisiodes Brindle, 1978: 94, figs 136-138, 247.

Type material

1 S, paratype, F3233.1383; 2 5?, paratypes, F3233.1384-1385; dried, mounted on pin.

MILES C., Dermaptera of the Manchester Museum

Locality labels

F3233.1383: Museum Paris / Congo Beige / Eala III. 1935 / J. Ghesquiere.

F3233.1384: Museum Paris / Congo Beige / Eala 1.1935 / J. Ghesquiere.

F3233.1385: Musee du Congo / Eala / - VIII -1935 / J. Ghesquiere.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis incisiodes Brindle, 1978.

Remark

The spelling 'incisiodes’ is used consistently in the original description. Steinmann (1989c: 217) and

Hopkins et al. (2014) refer to this species as incisoides.

indica Brindle, 1966 {Forcipula)

Forcipula indica Brindle, 1966d: 265, figs 26, 36.

Type material

1 S, paratype, F3233.1494, dried, mounted on pin.

Locality label

India: U.P / Song River / Dehra Dun. // Under stone / in water / 17.6.1945.

Taxonomy

Labiduridae, Labidurinae, Forcipula indica Brindle, 1966.

Remark

The date given in the original description is 1948, not 1945 as on the specimen label.

insulana Brindle, 1972 ( Gonolabis )

Gonolabis insulana Brindle, 1972c: 119, fig. 5a-e.

Type material

2 38, paratypes, F3233.188, F3233.190; 1 §, paratype, F3233.189; dried, mounted on pin.

Locality labels

F3233.188: S. / Mapld. Yap Group / Jul-Au 50 / R J Goss. Pac. S. Bd.

F3233.189: Pac. S. Bd. Yap Group / Jul-Au 50 / R J Goss. Gagil Dist.

F3233.190: N. Yap Is. / Micronesia / July-Aug. 1950 / R. J. Goss

Taxonomy

Anisolabididae, Anisolabidinae, Gonolabis insulana Brindle, 1972.

insularis Brindle, 1976 ( Auchenomus )

Auchenomus insularis Brindle, 1976c: 228, figs 27-28.

Type material

1 5, paratype, F3233.561, dried, mounted on card.

European Journal of Taxonomy 141: 1-138 (2015)

Locality label

New Hebrides / Espiritu Santo I. / (SW): Above / Namatasopa, 400m. / Aug. 30, 1957.

Taxonomy

Spongiphoridae, Sparattinae, Auchenomini, Auchenomus insular is Brindle, 1976.

intermedius Brindle, 1966 ( Archidux )

Archidux intermedius Brindle, 1966c: 54, figs 13, 17, 21, 24.

Type material

1 <S, paratype, F3233.1702; 1 paratype, F3233.1703; dried, mounted on card.

Locality labels

F3233.1702: I.R.S.A.C.-Mus.-Congo / Kivu: Mont Muhi, ruiss. / Isale, 3025 m. (humus) / N. Leleup

VII-1955.

F3233.1703: Foret transition//I.R.S.A.C.-Mus. Congo/Kivu : Riv. Nyalcagera, terr. /Kabare 1600/1700

m. / N. Feleup XI-1955.

Taxonomy

Forficulidae, Skendylinae, Cosmiella intermedia (Brindle, 1966).

Remarks

1. Transferred by Steinmann (1989c: 672).

2. Index card entry: Rwanda, Fake Kivu.

3. In the original description the specimen with data which matches F3233.1702 is a female paratype,

not a male.

jacobsoni Burr, 1912 ( Arixenia )

Fig. 16

Arixenia jacobsoni Burr, 1912: 105, fig.

Type material

2 syntypes, gender not specified, F3233.1866-67 (Fig. 16); 1 syntype, F3233.1868.

Locality labels

F3233.1866: E. Jacobson / Babakan. (Ban- / joemas) / Java / Mrt 1911/ Gouwa Fawa (grot) // Cotype //

3550 // A. jacobsoni Burr / det. Burr // Ex Mus. Tring / via Dr. K. Jordan.

F3233.1867: As F3233.1866 except number 3551.

F3233.1868: As F3233.1866 except number 3547 and det. label: Arixenia jacobsoni / Burr S / det. Burr.

Taxonomy

Arixeniidae, Xeniaria jacobsoni (Burr, 1912).

Remarks

1. Transferred by Maa (Nakata & Maa 1974: 319).

MILES C., Dermaptera of the Manchester Museum

2. In the original description, the author refers to the ‘nomotype’ being in his collection and to ‘numerous

specimens: Edw. Jacobson.’

3. In the original description, the locality is given as Banjoumas Residency, Gouwa Lawa cave near the

sea-shore at Babakan, Java.

jacobsoni Borelli, 1927 ( Timomenus )

Timomenus jacobsoni Borelli, 1927: 78, fig. 10.

Type material

1 8, 1 5, syntypes, F3233.1869; 1 8, syntype, F3233.1870; dried, mounted on card.

Locality labels

F3233.1869: Fort de Kock / (Sumatra) 920. M. / 1925 / leg. E. Jacobson.

F3233.1870: Anei Kloof / (Sumatra’s West / Kust) 500 M 1925 / leg. E. Jacobson.

Taxonomy

Forficulidae, Opisthocosmiinae, Timomenus jacobsoni Borelli, 1927.

Remarks

Brindle (1968f: 246) considered T. jacobsoni to be a lighter form of T. bicuspis (Stal, 1860); however,

Steinmann (1982b) presents T. jacobsoni as a species in its own right. In the original description, Borelli

referred to 4 specimens examined: 1 8 with data as F3233.1870, and 2 88 and 1 $ with data as

F3233.1869 except the date 1924. He did not designate a holotype and it appears therefore that all

four specimens should be considered syntypes. According to Steinmann (1982b: 365), 1 8 was held

at Museo ed Istituto di Zoologia Sistematoca, Torino, which Steinmann appears to have designated the

holotype (‘without penis’, although the male genitalia are figured in Steinmann’s description).

javana Hincks, 1955 ( Cranopygia )

Cranopygia javana Hincks, 1955c: 810, fig. 3.

Type material

9 88 , paratypes, F3233.8-11, F3233.17-19, F3233.23-24; 11 $ $, paratypes, F3233.12-16, F3233.20-

22, F3233.25-27; dried, mounted on pin.

Locality labels

F3233.18-21: E. Java / Soember / Brantas / Mt. Ardjuno / 6000’ 2.37 / M. E. Walsh.

F3233.15, F3233.17, F3233.22-24, F3233.27: Java: / Soember / Brantas /11.37 M. E. Walsh.

F3233.8-10, F3233.13-14, F3233.16, F3233.25: Kangean I. / (E. of Madura) /Ardjasa/M. E. Walsh.

F3233.11-12, F3233.26: Kangean I. / (E. of Madura) / Ardjasa 50’ / 5.36 M. E. Walsh.

Taxonomy

Pygidicranidae, Pygidicraninae, Cranopygia javana Hincks, 1955.

javanicus Hincks, 1951 ( Nesogaster )

Fig. 58

Nesogaster javanicus Hincks, 1951b: 570, figs 12-14.

European Journal of Taxonomy 141: 1-138 (2015)

Type material

1 S, holotype, F3233.545 (Fig. 58); 1 §, allotype, F3233.546; 1 S, paratype, F3233.547; dried, mounted

on card.

Type locality

East Java: / Soember Brantas / Feb. 1937 / M. E. Walsh.

Locality labels

F3233.546-547: As for type locality.

Taxonomy

Spongiphoridae, Nesogastrinae, Nesogaster aculeatus (de Bormans, 1900).

Remark

Nesogaster javanicus Hincks, 1951 synonymised with A. aculeatus (de Bormans, 1900) by Steinmann

(1989b: 23).

kambaitiensis Hincks, 1947 (Forficula)

Forficula kambaitiensis Hincks, 1947b: 29, figs 15-16.

Type material

1 (?, paratype, F3233.1729; 1 §, paratype, F3233.1730; dried, mounted on card.

Locality labels

F3233.1729: N. E. Burma / Kambaiti, 7000 ft / 19/4 R. Malaise.

F3233.1730: N. E. Burma / Kambaiti, 2000 m / 6/4. 1934 Malaise.

Taxonomy

Forficulidae, Forficulinae, Forficula kambaitiensis Hincks, 1947.

kelangi Brindle, 1969 ( Brachylabis )

Brachylabis kelangiJBrmdlQ, 1969b: 199, figs 1-3.

Type material

1 S, paratype, F3233.1423, dried, mounted on card.

Locality label

Kapar Mangrove / Forest Reserve / Kelang, West / Malaysia. 7.2.1969 / A. Sase Kumar.

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis kelangi (Brindle, 1969).

Remarks

1. Transferred by Steinmann (1989a: 370).

2. Index card entry: Port Swettenham.

MILES C., Dermaptera of the Manchester Museum

kermadecensis Giles, 1973 {Labia)

Labia kermadecensis Giles, 1973: 306, figs 1-4.

Type material

1 S, paratype, F3233.670, dried, mounted on card.

Locality label

Bells Flat, Raoul Isl, / Kermadec Isis. / 3 January 1967, J. C. Watt leg. / litter sample 67/57.

Taxonomy

Spongiphoridae, Fabiinae, Spirolabia kermadecensis (Giles, 1973).

Remarks

1. Transferred by Ste inm a nn (1987: 181).

2. Additional locality data in the original description: 6 ... litter under Metrosideros kermadecensis ,

Myoporum laetum and Rhopalostylus, dry litter on ‘Oneraki black sandy silt’, sieved, sample U ...’

(Giles 1973: 308).

kivuensis Brindle, 1973 ( Cliaetospania )

Chaetospania kivuensis BrindlQ, 1973f: 155, figs 191-194.

Type material

1 S, paratype, F3233.1512, dried, mounted on card.

Locality label

Recolte sous / ecorce d’arbre mort // I.R.S.A.C. - Mus. Congo / Kivu: Terr. Mwenga, / S.O. Itombwe,

Luiko / 2050 m (for. mont.) 1-1952 / N. Leleup.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania kivuensis Brindle, 1973.

Remark

Index card entry: Africa, Zaire, Kivu.

kjellanderi Brindle, 1967 ( Diplatys )

Diplatys kjellanderi Brindle, 1967b: 148, figs 3, 6, 8.

Type material

1 S, paratype, F3233.1302, dried, mounted on card.

Locality label

Bulbalan, Luzon.

Taxonomy

Diplatyidae, Diplatyinae, Circodiplatys kjellanderi (Brindle, 1967).

European Journal of Taxonomy 141: 1-138 (2015)

Remarks

Transferred by Steinmann (1986b: 109). Steinmann (1989c: 31) and Hopkins et al. (2014) have the

spelling ‘ kjellenderi Brindle dedicated this species to Dr. E. Kjellander (Brindle 1967b: 150).

koepckei Brindle, 1966 ( Idolopsalis)

Fig. 25

Idolopsalis koepckei Brindle, 1966a: 145, figs 3, 7, 14.

Type material

1 S, holotype, F3233.1456 (Fig. 25); 1 §, allotype, F3233.1457; 3 $$, paratypes, F3233.1458-1460; 3

5$, paratypes, F3233.1461-1463; dried, mounted on card.

Type locality

Hda. Taulis Peru / (ca. 6° 50’ S, 79° 10’ W) / 27 Dezember 1952 / leg.: Koepcke.

Locality labels

F3233.1457: As type locality.

F3233.1458: N. Peru, w. slopes /Andes, Hacienda / Taulis ca 6’ 50” / 79’ 10” S. Ca 2700 m. / berg-

urwald. / 27.12.52. Koepcke.

F3233.1459-1462: Same data except 1700m. / montane forest / rotten wood / leaves on ground / H W

Koepcke 1.52. // *Kp. 403j.

F3233.1463: As type locality except mid. Jan. 1952.

Taxonomy

Anisolabididae, Idolopsalidinae, Idolopsalis koepckei Brindle, 1966.

Remark

The original description gives the date for the holotype and the specimen matching F3233.1458 as 24

Dec. 1952, not 27 Dec. 1952.

koepckei Brindle, 1968 {Strongylopsalis)

Fig. 55

Strongylopsalis koepckei Brindle, 1968c: 279, figs 1, 4.

Type material

1 S, holotype, F3233.1619 (Fig. 55); 1 §, allotype, F3233.1620; dried, mounted on card.

Type locality

Peru / Bei Kampanillayo / (bei Tarma) 6 August 1953 / leg. Koepcke.

Locality label

F3233.1620: bei Campanillay / Peru (bei Tarna) / 6.viii. 1953 / H. W. Koepcke.

Taxonomy

Spongiphoridae, Strongylopsalinae, Strongylopsalis koepckei Brindle, 1968.

MILES C., Dermaptera of the Manchester Museum

Remark

The original description gives the locality as Campanillayo (bei Tama). The index card has Bei

Campanillayo. (Tarma is a city in Peru.)

kuscheli Hincks, 1959 ( Esphalmenus )

Fig. 35

Esphalmenus kuscheli Hincks, 1959: 201, fig. 204.

Type material

1 <S, holotype, F3233.1359 (Fig. 35); 1 §, allotype, F3233.1360; dried, mounted on card.

Type locality

Chile. Bio-Bio / Faguna Faja / Fos Barros - 2100 m / 18.1.48. // Coll. / Kuschel // Esphalmenus /

Kuscheli / Hincks / Holotype S / det. W. D. Hincks.

Locality label

F3233.1360: Chile. Bio-Bio / Faguna Faja / Fos Barros - 1500 m / 23.1.48. // Esphalmenus / Kuscheli /

Hincks / Allotype $ / det. W. D. Hincks.

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus kuscheli Hincks, 1959.

labioides Hincks, 1952 ( Geracodes )

Geracodes labioides Hincks, 1952b: 23, figs 7-9.

Type material

1 (?, paratype, F3233.1445; 5 §§, paratypes, F3233.1446-1450; dried, mounted on card.

Locality labels

F3233.1445-1448: Sierra Leone / Njala / 15.ix - 8.xii 1930 / E. Hargreaves / At light.

F3233.1449: Sierra Leone / Njala / 30.X.32;

F3233.1450: Same data except date 21 Nov. 1923.

Taxonomy

Anisolabididae, Isolabidinae, Geracodes labioides Hincks, 1952.

laevigata Brindle, 1978 (Anisolabis)

Anisolabis laevigata Brindle, 1978: 100, figs 142, 145, 251.

Type material

1 5, paratype, F3233.1386, dried, mounted on card.

Locality label

Nimba (Guinee) / Lamotte, Amiet, / Vanderplaetsen / XII 56 - V 57.

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Anisolabididae, Anisolabidinae, Gonolabis laevigata (Brindle, 1978).

Remark

Transferred by Steinman (1989a: 197).

laevis Brindle, 1975 (Archidux)

Archidux laevis Brindle, 1975b: 694, fig. 4.

Type material

2 SS, paratypes, F3233.1704-1705, dried, mounted on card.

Locality labels

F3233.1704-1705: Tanzanie: Mts Uluguru / Chenzema, alt. 1700m. /2-22/VII/71 // Coll. Mus. Tervuren

/ Mission Mts Uluguru / L. Berger, N. Leleup / J. Debecker V/VIII/71.

Taxonomy

Forficulidae, Skendylinae, Cosmiella laevis (Brindle, 1975).

Remarks

1. Transferred by Steinmann (1989c: 672).

2. The original description refers to only one male paratype at MMUE.

34 F3233.1361

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35 F3233.1359

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F3233.1341

fUndii,Col rotators OHf*

Vilptrtl 3500 ft. IMI1

tfM 1038

BrUdb

det. A.Brindle 196*-

F3233.1491

Figs 34-39. Holotypes (c?) of Pygidicranidae and Labiduridae in the collection of the Manchester

Museum. 34. Esphalmenus mucronatus Hincks, 1959. 35. E. kuscheli Hincks, 1959. 36. E. argentinus

Hincks, 1959. 37. E. dentatus Hincks, 1959. 38. Tagalina curta Brindle, 1975. 39. Gonolabidura nathani

Brindle, 1965. Scale bars = 1 cm.

MILES C., Dermaptera of the Manchester Museum

laminata Brindle, 1973 (Strongylopsalis)

Strongylopsalis laminata Brindle, 1973e: 11, figs 1, 6.

Type material

1 S, paratype, F3233.1621, dried, mounted on card.

Locality label

Suri / Bolivia / W. M. Ma nn // Mulford / Bio. Exp. / 1921-22.

Taxonomy

Spongiphoridae, Strongylopsalinae, Strongylopsalis laminata Bundle, 1973.

Remark

Index card entry: Bolivia, Cajuata.

lamottei Brindle, 1971 (An is old bis)

Anisolabis lamottei Brindle, 1971c: 270, figs 5-7, 9.

Type material

3 88, paratypes, F3233.1387-1389; 2 paratypes F3233.1390-1391; dried, mounted on card.

Locality labels

F3233.1387-1388: Savane / Sommet 1550 m / 25.V. 1963. // Mission ENS-IFAN / Aux Monts Loma /

Sierra Leone.

F3233.1389: Savane / 2 metres ou vegetation. Piste de / Bandankoro / 29.V. 1963 // Mission ENS-IFAN

/ Aux Monts Loma / Sierra Leone.

F3233.1390: Savane / 1580 m / 18-V-1963. // Mission ENS-IFAN / Aux Monts Loma / Sierra Leone.

F3233.1391: Prairie a / cyperacees / 20. V. 1963. // Mission ENS-IFAN / Aux Monts Loma / Sierra Leone.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis lamottei Brindle, 1971.

Remark

For the specimen most probably corresponding to F3233.1390, the original description gives the

altidtude as 1630 m, not 1580 m.

lamottei Hincks , 1954 (Diplatys)

Diplatys lamottei Hincks, 1954b: 115, figs 17-18.

Type material

1 5, paratype, F3233.1303, dried, mounted on card.

Locality label

IFAN / Nimba (Guinee) / Lamotte et Roy / VII - XII 51.

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Diplatyidae, Diplatyinae, Paradiplatys lamottei (Hincks, 1954).

Remarks

Transferred to Lobodiplatys lamottei by Steinmann (1974: 199) and to the subgenus Paradiplatys by

Steinmann (1989c: 82). The combination Paradiplatys (Paradiplatys) lamottei was reinstated by Engel

& Haas (2007: 19).

lateralis Brindle, 1989 ( Carcinophora )

Carcinophora lateralis Brindle, 1989: 14, fig. 11.

Type material

3 33, paratypes, F3233.1393, F3233.1839-1840; 4 paratypes, F3233.1841-1844; dried, mounted

on card.

Locality labels

F3233.1393: Panama, Fortuna / Wet forest / at light, 1050 m. / 11 Oct. 1976 / Dr. Henk Wolda, STRI.

Same data except date: F3233.1840, 1 Oct. 1976; F3233.1843, 7 Oct. 1976; F3233.1844, 15 Oct. 1976;

F3233.1841, 24 May 1977; F3233.1839, 6 Sep. 1978; F3233.1842, 24 Sep. 1978.

Taxonomy

Anisolabididae, Anisolabidinae, Carcinophora lateralis Brindle, 1989.

Remarks

1. Not listed in Steinmann (1989c) or Hopkins et al. (2014).

2. The date range of the specimens does not match exactly the date range in the original description:

27.iii.1976-2.viii. 1977.

leleupi Brindle, 1968 ( Anophthalmolabis )

Anophthalmolabis leleupi Brindle, 1968b: 172, figs 1-2.

Type material

1 3, paratype, F3233.101; 1 §, paratype, F3233.100; dried, mounted on card.

Locality labels

F3233.100-101: Galapagos Isles / Santa Cruz / 27.XII. 1964 /N. Leleup.

Taxonomy

Anisolabididae, Anophthalmolabidinae, Anophthalmolabis leleupi Brindle, 1968.

lenkoi Brindle, 1968 ( Brachylabis )

Brachylabis lenkoi Brindle, 1968a: 32, figs 1-2.

Type material

2 33, paratypes, F3233.1424-1425, dried, mounted on card.

MILES C., Dermaptera of the Manchester Museum

Locality labels

F3233.1424-1425: Barueri / SP, Brazil / 20.VIII. 1967 / K. Lenko col.

Taxonomy

Anisolabididae, Brachylabidinae, Br achy lab is lenko i Brindle, 1968.

Remark

Index card entry: Sao Paolo.

lingua Brindle, 1970 (Adiathella)

Adiathella lingua Brindle, 1970f: 689, fig. 57.

Type material

1 S, paratype, F3233.926, dried, mounted on card.

Locality label

J. L. Gressit / Collector // Solomon Is. / Guadalcanal: Suta / 500-1200 m. / IV-27-1956.

Taxonomy

Chelisochidae, Chelisochinae, Adiathella lingua Brindle, 1970.

linsleyi Brindle, 1970 (Skalistes)

Skalistes linsleyi Brindle, 1970b: 226, fig. 4.

Type material

3 $$, paratypes, F3233.1780-1782; 2 §§, paratypes, F3233.1783-1784; dried, mounted on pin.

Locality labels

F3233.1780-1784: Michoacan / Mex. V-13-38 // Pres, by E. G. Finsley

Taxonomy

Forficulidae, Forficulinae, Skalistes linsleyi Brindle, 1970.

longicollis Hincks, 1957 ( Nesolabia )

Nesolabia longicollis Hincks, 1957c: 54, figs 9-11.

Type material

1 <S, paratype, F3233.1832; 1 paratype, F3233.1833; dried, mounted on card.

Locality labels

F3233.1832-1833: La Reunion / Plaine des Cafres / Piton Manuel / 27 I 1955 // Institut / Scientifique

/ Madagascar.

Taxonomy

Spongiphoridae, Geracinae, Nesolabia longicollis Hincks, 1957.

European Journal of Taxonomy 141: 1-138 (2015)

Remark

The original description makes no reference to a female paratype with data matching that for F3233.1833.

lucasi Brindle, 1974 {Skalistes)

Skalistes lucasi Brindle, 1974a: 5, figs 1—4.

Type material

4 88, paratypes, F3233.1785-1788; 4 §§, paratypes, F3233.1789-1792; dried, mounted on pin.

Locality labels

F3233.1785, F3233.1789: Mexico: Chiapas-Oaxaca / border, 21 km W. Rizo / de Oro along ridge SE

Cerro Baul, 1615 m. / 8-IX-1972, K. E. Lucas / within Tillandisa sp.

F3233.1791: Same data except ‘within Tillandsia guatamalensis / specimen #: B3-\

F3233.1786: Same data except ‘within Vriesia sp’.

F3233.1792 : Same data except ‘within epiphytic Vriesia sp. / specimen #: B3-\

F3233.1787-1788, F3233.1790: Same data as F233.1792 except date 8 Jan. 1973.

Taxonomy

Forficulidae, Forficulinae, Skalistes lucasi Brindle, 1974.

lucens Brindle, 1975 (Forficula)

Forficula lucens Brindle, 1975c: 48, figs 56-57.

Type material

1 8, paratype, F3233.1731; 1 §, paratype, F3233.1732; dried, mounted on card.

Locality labels

F3233.1731-1732: Tongsa 2150 m / 24/6 // Nat.-Hist. Museum / Basel - Bhutan / Expedition 1972.

Taxonomy

Forficulidae, Forficulinae, Guanchia lucens (Brindle, 1975).

Remark

Transferred by Steinmann (1993: 584).

lundbladi Hincks, 1938 ( Perirrhytus )

Perirrhytus lundbladi Hincks, 1938: 5, figs 5-6.

Type material

3 paratypes, gender unspecified, F3233.1845-1847; dried, mounted on card.

Locality labels

F3233.1845: Madeira: / Caramujo. / 12.8.35 / “Erica - [illegible]” / Dr. O. Lundblad. / Stockholm Mus.

F3233.1846-1847: Madeira: /Feiteiras / 15.8.35 / Dr. O. Lundblad.

MILES C., Dermaptera of the Manchester Museum

Taxonomy

Forficulidae, Anechurinae, Perirrhytus lundbladi Hincks, 1938.

Remarks

Index card entry: 2 S3, 1 ?• This agrees with the original description, which states that 15 paratypes

were examined, that all paratypes with the same locality information as F3233.1845 were males and that

the only two described from Feiteiras (as F3233.1846-1847) were 1 S and 1 §.

machu-picchuensis Brindle, 1971 ( Marava )

Marava machu-picchuensis Brindle, 197Id: 250, figs 1, 5, 7.

Type material

1 S, paratype, F3233.1551, dried, mounted on card.

Locality label

Torentoy Canyon (Ba- / se Machu-Picchu) / 2000-2000m - Peru / VI-VII.964 B. Malkin.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava machupicchuensis Brindle, 1971.

magnus Brindle, 1988 {Vostox)

Fig. 41

Vostox magnus Brindle, 1988: 50, figs 9-10.

Type material

1 (J,holotype,F3233.1663 (Fig. 41); 4 paratypes,F3233.1664-1667; 5 $$,paratypes,F3233.1668-

1672; dried, mounted on card.

Type locality

Panama, 1300M / Alto Fino, near / Boguete, Chiriqui / 25/31.VIII. 1976 / Dr. Henk Wolda, STRI.

Locality labels

F3233.1664: Panama, 1300M / Alto Lino, near / Boguete, Chiriqui / 12-VI-1978 / Dr. Henk Wolda,

STRI.

Same data except date: F3233.1665, 6 Sep. 1977; F3233.1668, 3-4 Jun. 1978.

F3233.1666: Panama, Fortuna / Wet forest / at light, 1050m / 23 Nov. 1976 / Dr. Henk Wolda, STRI.

Same data except date: F3233.1667, 29 Nov. 1976; F3233.1671, 12 Apr. 1978; F3233.1670, 15 May

1978; F3233.1672, 14 Aug. 1978; F3233.1669, 25 Aug. 1978.

Taxonomy

Spongiphoridae, Spongiphorinae, Vostox magnus Brindle, 1988.

malaisei Hincks, 1947 ( Chaetospania )

Chaetospania malaisei Hincks, 1947b: 19, figs 9-11.

European Journal of Taxonomy 141: 1-138 (2015)

Type material

1 S, paratype, F3233.1513, dried, mounted on card.

Locality label

N. E. Burma / Kambaiti 7000 ft / 30/5 1934 / R. Malaise.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania malaisei Hincks, 1947.

malgacha Brindle, 1966 (Euborellia)

Euborellia malgacha Brindle, 1966g: 236, fig. 28.

Type material

1 S, paratype, F3233.1441, dried, mounted on card.

Locality label

Sahafanjana / Manambato / (Anove). Institut / Scientifique / Madagascar.

Taxonomy

Anisolabididae, Anisolabidinae, Euborellia malgacha Brindle, 1966.

malgacha Brindle, 1966 {Labia)

Labia malgacha Brindle, 1966g: 245, figs 33-35.

Type material

2 $ paratypes, F3233.1541-1542, dried, mounted on card.

Locality labels

F3233.1541-1542: Ampitameloka / 840 m / sud Moramanga / VIII/56 R Griv. // Institut Scientifique /

Madagascar.

Taxonomy

Spongiphoridae, Labiinae, Paralabella malgacha (Brindle, 1966).

Remark

Transferred by St.e inm a. nn (1989b: 486).

mameti Hincks, 1950 {Labia)

Labia mameti Hincks, 1950b: 354, figs 1-3.

Type material

1 S, paratype, F3233.1594, dried, mounted on card.

Locality label

Mauritius / Le Pouce Mt. / 2. XI. 1948 / R. Mamet.26.

MILES C., Dermaptera of the Manchester Museum

Taxonomy

Spongiphoridae, Geracinae, Pseudovostox mameti (Hincks, 1950).

Remarks

1. Transferred by Brindle (1973d: 142).

2. The original description has the collection date as 1949, not 1948.

martensi Brindle, 1987 ( Aborolabis )

Aborolabis martensiBvmdlQ, 1987: 336, figs 12, 17.

Type material

1 3, paratype, F3233.1837; 1 §, paratype, F3233.1838; dried, mounted on card.

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dat.A.Brindle 1990

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F3233.1814

Figs 40-48. Holotypes (c?) of Spongiphoridae from the collection of the Manchester Museum.

40. Vostox bolivianus Brindle, 1971. 41. V magnus Brindle, 1988. 42. V dentatus Brindle, 1988.

43. Maravapygidiata Brindle, 1988. 44. M. pallida Brindle, 1988. 45. M. flavohumeralis Brindle, 1988.

46. M. nigrocincta Brindle, 1988 (no determination label). 47. M. gracilis Brindle, 1988. 48. M. parvula

Brindle, 1988. Scale bars = 1 cm.

European Journal of Taxonomy 141: 1-138 (2015)

Locality labels

F3233.1837-1838: Nepal, Gorkha/Dist. Buri Gandaki / 29 July 1983 / 700-800 m. 219.

Taxonomy

Anisolabididae, Anisolabidinae, Aborolabis martensi Brindle, 1987.

Remark

The original description gives the locality as between Suteo and Labubesi, Shorea.

martensi Brindle, 1974 ( Allodahlia )

Allodahlia martensi ''Brindle, 1974c: 154, figs 13-16, 18.

Type material

1 S, paratype, F3233.1692; 2 paratypes, F3233.1693-1694; dried, mounted on card.

Locality labels

F3233.1692: Nepal. Thalcsang / above Tulcche / 3100-3400m. / 6/7.VII. 1973 / J. Martens.

F3233.1693, same data except: 3150-3200m. / 2/4.VII. 1973.

F3233.1694: Nepal 2600m. / Gosainkund / 23.IV. 1973 / J. Martens.

Taxonomy

Forficulidae, Allodahliinae, Allodahlia martensi Brindle, 1974.

Remarks

The original description states, ‘1 male, 1 female paratypes (32) and 1 male, 1 female paratypes (35)

have been retained in the Manchester Museum.’ [32 and 35 refer to data sets.] F3233.1692 matches data

set 32 in the description except the date 5-10 Aug. 1970. The female with this data appears to be missing

from MMUE. F3233.1693 matches data set 34 in the description exactly but not 32 or 35. The data for

F3233.1694 does not match any in the description, except the collector.

masai Hincks, 1949 (Afrolabia)

Afrolabia masai Hincks, 1949a: 76, figs 1-3.

Type material

1 <$, paratype, F3233.1609; 1 §, paratype, F3233.1610; dried, mounted on card.

Locality labels

F3233.1609-1610: Van Someren / Meru, 7/43.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox masai (Hincks, 1949).

Remarks

1. Transferred by Brindle (1973f: 216).

2. Index card entry: Africa, Tanzania, Mt. Meru.

MILES C., Dermaptera of the Manchester Museum

meridionalis Brindle, 1978 (Brachylabis)

Fig. 22

Brachylabis meridionalis Bundle, 1978: 29, figs 11-13.

Type material

1 $, holotype, F3233.1426 (Fig. 22), dried, mounted on card.

Type locality

S. Africa / Storms R. / 21.9.51 / J F Darbyshire.

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis meridionalis (Brindle, 1978).

Remark

Transferred by Steinmann (1989a: 343).

meridionalis Brindle, 1973 ( Chaetospania )

Fig. 57

Chaetospania meridionalis Brindle, 1973f: 151, fig. 187.

Type material

1 $, holotype, F3233.1514 (Fig. 57); 1 §, allotype, F3233.1515; 1 $ paratype, F3233.1516; dried,

mounted on card.

Type locality

Vumba Mountains / S. Rhodesia / Under bark on dead / tree 21.3.1959 / C. N. Smithers.

Locality labels

F3233.1515—1516: As type locality.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania meridionalis Brindle, 1973.

Remark

The original description refers to two male paratypes, not one.

milloti Hincks, 1953 {Antisolabis)

Antisolabis milloti Hincks, 1953: 365, figs 4-5.

Type material

1 S, paratype, F3233.1484, dried, mounted on card.

Locality label

17-10-51 / (A.R.) // Institut / Scientifique / Madagascar.

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Anisolabididae, Brachylabidinae, Metisolabis milloti (Hincks, 1953).

Remark

Transferred by Brindle (1966g: 234).

minima Brindle, 1966 ( Brachylabis )

Brachylabis minima Brindle, 1966e: 40, figs 6, 15.

Type material

1 <S, paratype, F3233.1474; 1 paratype, F3233.1475; dried, mounted on card.

Locality labels

F3233.1474: Foret transition // Coll. Mus. Congo / Tanganyika Terr.: Uluguru / Mts., Ululu-Ndogo, /

1500 m. 8-V-1957.

F3233.1475: Foret transition// Coll. Mus. Congo / Tanganyika Terr.: Uluguru/Mts., sommet duKidunda

/ 1800- 1950 m. 3-V-1957.

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis minima (Brindle, 1966).

Remark

Transferred by Steinmann (1989a: 348).

minimus Brindle, 1966 ( Archidux )

Archidux minimus Brindle, 1966c: 56, figs 14-16, 20, 23.

Type material

1 S, paratype, F3233.1706; 1 §, paratype, F3233.1707; dried, mounted on card.

Locality labels

F3233.1706-1707: Recolte dans / l’humus // I.R.S.A.C.-Mus. Congo / Kivu : Terr. Lubero / 2.200 m.

27-XI-1951 /N. Leleup / (for. mont. avec bamb.)

Taxonomy

Forficulidae, Skendylinae, Cosmiella minima (Brindle, 1966).

Remarks

1. Transferred by Steinmann (1989c: 673).

2. Index card entry: Africa, Zaire.

minuscula Brindle, 1977 ( Marava )

Marava minuscula Brindle, 1977b: 125, figs 23-24.

Type material

1 S, paratype, F3233.1552; 1 paratype, F3233.1553; dried, mounted on card.

MILES C., Dermaptera of the Manchester Museum

Locality labels

F3233.1552-1553: Venezuela - Bolivar / El Hormiguero Me- / seta de Nuria. / 500m. 13-17.XII.74.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava furia Steinmann, 1989.

Remark

New name by Steinmann (1989c: 502): ‘ Marava furia Steinmann (new name for Marava minuscula

Brindle, 1977 as a junior homonym of Forficula minusculaLatrQiUe, 1817).’

minutissima Brindle, 1972 (Anisolabis)

Anisolabis minutissima Brindle, 1972c: 128, figs 6a-c, 7b.

Type material

1 S, paratype, F3233.1394, dried, mounted on card.

Locality label

Garakayo I. / Pelew Islands / VIII: 9 : 45. // Coll. & pres, by / Henry S. Dybas.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis minutissima Brindle, 1972.

Remarks

1. Index card entry: Pelew Islands = Palau.

2. The original description refers only to the holotype and allotype. There is no reference to paratypes.

mucronatus Hincks, 1959 ( Esphalmenus )

Fig. 34

Esphalmenus mucronatus Hincks, 1959: 202, fig. 208.

Type material

1 <S, holotype, F3233.1361 (Fig. 34), dried, mounted on card.

Type locality

Bolivia / Estacion Condor / 4790 m / 11.1.49. Coll. / Kuschel.

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus mucronatus Hincks, 1959.

nathani Brindle, 1965 ( Gonolabidura )

Fig. 39

Gonolabidura nathani Brindle, 1965: 594, fig. 21.

Type material

1 S, holotype, F3233.1491 (Fig. 39), dried, mounted on card.

European Journal of Taxonomy 141: 1-138 (2015)

Type locality

S. India, Coimbatore Dstr. / Valparal 3500 ft. II—III / coll. P. S. Nathan 1938.

Taxonomy

Labiduridae, Allostethinae, Gonolabidura nathani Brindle, 1965.

nepalensis Brindle, 1974 ( Anisolabis )

Anisolabis nepalensis Brindle, 1974c: 148, figs 4-5.

Type material

1 c?, paratype, F3233.1395; 1 §, paratype, F3233.1396; dried, mounted on card.

Locality labels

F3233.1395: Nepal /Myangdi -Khola/Muri 2100 m. /IV. 1970/ J. Martens. F3233.1396: Nepal: Sitha

- Gorapani / 2300 - 2700m / J. Martens leg. ll.Jul.73.

Taxonomy

Anisolabididae, Anisolabidinae, Aborolabis nepalensis (Brindle, 1974).

Remarks

1. Transferred by Steinmann (1979: 215).

2. The original description gives the altitude for F3233.1395 as 2300 m, not 2100 m. The locality of

F3233.1396 is given in the description as between Sika (not Sitha as on the label) and Gorapani.

nevermanni Brindle, 1966 (Vara)

Vara nevermanni Brindle, 1966f: 138, figs 1—4.

Type material

1 S, paratype, F3233.1749; 2 §§, paratypes, F3233.1750-1751; dried, mounted on card.

Locality labels

F3233.1749: Costa Rica / Farm Hamburg / am Reventazon / 3.XII. 1932. // Urwald welkes Laub / F.

Neverm a rm leg. / Eing. Nr. 104, 1932.

F3233.1750: Costa Rica / Farm Hamburg / am Reventazon / 29.3.31. // F. Nevermann leg. / Eing.

Nr. 164, 1931.

F3233.1751: Costa Rica / Turrialba / 29.VIII.32 // An Carica papaya / F. Nevermann leg. / Eing. Nr. 42

1934.

Taxonomy

Forficulidae, Ancistrogastrinae, Sarcinatrix anomalia (Rehn, 1903).

Remarks

1. Vara nevermanni Brindle, 1966 synonymised with Sarcinatrix anomalia (Rehn, 1903) by Brindle

(1967a: 189).

2. The original description gives the date for the specimen which matches F3233.1751 as 29 Jul. 1932,

not 29 Aug. 1932.

MILES C., Dermaptera of the Manchester Museum

3. In the original description, no reference to a specimen matching the date of F3233.1750 (i.e., 29 Mar.

1931) was made.

nigerrima Brindle, 1966 ( Forficula )

Forficula nigerrima Brindle, 1966b: 148, figs 1-3.

Type material

1 S, paratype, F3233.1724, dried, mounted on card.

Locality label

Nyasaland / Mt Mlange / 8.8.1956 / AWR McCrae. // Ridge 7500’ / A m ESE of / Little Rus Camp /

Little Rus Plat. // Under stones / on peaty / grass mat.

Taxonomy

Forficulidae, Forficulinae, Afroforficula nigerrima (Brindle, 1966).

Remarks

1. Transferred by Ste inm a nn (1993: 544).

2. Index card entry: Africa, Malawi.

3. Mt Mlange (locality label) is a variant spelling of Mt. Mlanje or Mt. Mulanje, Malawi.

nigrescens Brindle, 1987 ( Aborolabis )

Aborolabis nigrescens Brindle, 1987: 337, figs 9, 11.

Type material

1 <S, paratype, F3233.1835; 1 §, paratype, F3233.1836; dried, mounted on card.

Locality labels

F3233.1835-1836: Nepal, Nuwakot / District, Trisuli / 21/22 July 1983 / 600-650 m 907.

Taxonomy

Anisolabididae, Anisolabidinae, Aborolabis nigrescens Brindle, 1987.

nigrescens Brindle, 1977 ( Pyragra )

Pyragra nigrescens Brindle, 1977b: 113, figs 6, 8.

Type material

1 5, paratype, F3233.1340, dried, mounted on pin.

Locality label

Venezuela, Bolivar / El Bochinche Res. / Forestal Imataca / 200m. 6-13.XII.74.

Taxonomy

Pygidicranidae, Pyragrinae, Pyragra nigrescens Brindle, 1977.

European Journal of Taxonomy 141: 1-138 (2015)

nigrinus Brindle, 1984 (Esphalmenus)

Esphalmenus nigrinus Brindle, 1984: 286, fig. 24.

Type material

1 5, paratype, F3233.1362, dried, mounted on card.

Locality label

Ecuador / N. of Azogues / 3000 m.

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus nigrinus Brindle, 1984.

Remark

The original description refers to one male paratype at MMUE. The MMUE specimen is labelled §.

nigrita Brindle, 1971 ( Idolopsalis )

Idolopsalis nigrita Brindle, 197Id: 255, fig. 9.

Type material

1 S, paratype, F3233.1464, dried, mounted on card. 1 §, paratype, F3233.1465, dried, mounted on pin.

Locality labels

F3233.1464: Bromelia / 10m // Est. Biol. Boraceia /Salesopolis, SP / 26.XI. 1968 / E. X. Rabello col.

F3233.1465: Est. Biol. Boraceia/Salesopolis, SP / 18.VI. 1966 / C. Costa col.

Taxonomy

Anisolabididae, Idolopsalidinae, Idolopsalis nigrita Brindle, 1971.

Remark

The original description gives the locality San Paulo.

nigritum Brindle, 1974 {Eugerax)

Eugerax nigritum Brindle, 1974b: 118.

Type material

1 S, paratype, F3233.1533, dried, mounted on card.

Locality label

Venezuela - Aragua / Rancho Grande / 1100m. 12.V.73. // J. A. Clavijo / G. Equis.

Taxonomy

Spongiphoridae, Geracinae, Eugerax nigritum Brindle, 1974.

MILES C., Dermaptera of the Manchester Museum

nigrocinctci Brindle, 1988 {Marava)

Fig. 46

Marava nigrocincta Brindle, 1988: 52, figs 14, 19.

Type material

1 holotype, F3233.1813 (Fig. 46); 10 $$, paratypes, F3233.1850-1859; 4 paratypes,

F3233.1860-1863; dried, mounted on card.

Type locality

Panama, Fortuna / Wet forest / at light, 1050m / 10.5.1977 / Dr. Henk Wolda, STRI.

Locality labels

As type locality except dates: F3233.1862, 13 Nov. 1976; F3233.1860, 28 Jul. 1977; F3233.1861, 15

Aug. 1977; F3233.1863, 11 Sep. 1977; F3233.1850-1854, 24 Sep. 1977, F3233.1855-1858, 25 Sep.

1977; F3233.1859, 24 Oct. 1977.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava nigrocincta Brindle, 1988.

Remark

The original description gives the collection date of the holotype as 21 Jul. 1977, not 10 May 1977 as

on the locality label.

nilgiriensis Hincks, 1955 ( Diplatys )

Diplatys nilgiriensis Hincks, 1955a: 111, fig. 144.

Type material

1 S, paratype, F3233.1304, dried, mounted on card.

Locality label

S. India / Nilgiri Hills / Cherangode / 3,500’ X.50 / PS. Nathan.

Taxonomy

Diplatyidae, Diplatyinae, Diplatys (Syndiplatys) nilgiriensis Hincks, 1955.

Remarks

1. Transferred by Steinmann (1986a: 175).

2. The original description gives the collection date as Nov. 1950, not Oct. 1950 as on the locality label.

nimbaensis Hincks, 1954 (Anisolabis)

Anisolabis nimbaensis Hincks, 1954b: 103, fig. 1.

Type material

1 paratype, F3233.1397; 1 5, paratype, F3233.1398; dried, mounted on card.

European Journal of Taxonomy 141: 1-138 (2015)

Locality labels

F3233.1397-1398: Mt To (1600 m) / Camp 1. Museum Paris /Nimba (Guinee) / M. Lamotte II. VI. 42.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis nimbaensis Hincks, 1954.

Remarks

1. Index card entry: Africa, Guinea, Nimba Mts.

2. The original description refers to 4 paratypes in the Hincks collection.

nodulosa Brindle, 1973 {Labia)

Labia nodulosa Brindle, 1973c: 395, figs 1-2.

Type material

1 S, paratype, F3233.1543, dried, mounted on card.

Locality label

Costa Rica / Coronado / 15.11.1931 / F. Nevermann.

Taxonomy

Spongiphoridae, Labiinae, Paralabella conspicua (Borelli, 1906).

Remark

Synonymised with L. conspicua (Borelli, 1906) by Brindle (1985: 74). L. conspicua transferred to

Paralabella by Steinmann (1989b: 473).

notonoma Hincks, 1952 {Antisolabis)

Antisolabis notonoma Hincks, 1952a: 214, fig. 1.

Type material

1 S, paratype, F3233.1489, dried, mounted on card.

Locality label

Mt. Keira / New South Wales / 4 : 12 : 1949 / C. E. Chadwick // Under rotting log.

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis notonoma Hincks, 1952.

ocellatus Brindle, 1971 ( Vostox)

Vostox ocellatus Brindle, 1971b: 539, figs 63-65.

Type material

1 S, paratype, F3233.1635, dried, mounted on card.

Locality label

Honduras / 3913999 / Primavera / logs. n. y. / Aug. 19, 1939.

MILES C., Dermaptera of the Manchester Museum

Taxonomy

Spongiphoridae, Spongiphorinae, Vostox ocellatus Brindle, 1971.

ochracea Brindle, 1966 ( Chaetospania )

Chaetospania ochracea Brindle, 1966g: 241, figs 45^46.

Type material

1 <$, paratype, F3233.1517, dried, mounted on card.

Locality label

Manambato / (Anove) // Institut Scientifique / Madagascar.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania ochracea Brindle, 1966.

ochroptera Brindle, 1972 ( Allodahlia )

Allodahlia ochroptera Brindle, 1972b: 27, fig. 9.

Type material

1 5, paratype, F3233.1695, dried, mounted on card.

Locality label

Bhamo / Birmania / Fea IX 1886.

Taxonomy

Forficulidae, Allodahliinae, Allodahlia ochroptera Brindle, 1972.

Remarks

1. Bhamo is a city in northern Myanmar.

2. The original description gives the collection date as Oct. 1886, not Sep. 1886 as on the locality label.

pallida Brindle, 1988 ( Marava )

Fig. 44

Marava pallida Brindle, 1988: 51, figs 13, 18.

Type material

1 c?,holotype,F3233.1558 (Fig. 44);4 c?c?,paratypes,F3233.1559-1562; 8 $$,paratypes,F3233.1563-

1570; dried, mounted on card.

Type locality

Panama, Barro / Colorado Is., S-M / light trap, ground / level / 30.9.1977 / Dr. Henk Wolda, STRI.

Locality labels

As type locality except dates: F3233.1562,22 Nov. 1977; F3233.1560,1 May 1978; F3233.1559,17Jun.

1978; F3233.1561, 8 Sep. 1978; F3233.1563, 27 Sep. 1978; F3233.1568, 13 Oct. 1978; F3233.1567,

European Journal of Taxonomy 141: 1-138 (2015)

21 Oct. 1978; F3233.1565, 26 Oct. 1978; F3233.1564, 22 Nov. 1978; F3233.1569, 13 Dec. 1978.

F3233.1566: Panama, Barro / Colorado Is., S-M / light trap, ground / level / WK 20 D / Dr. Henk Wolda,

STRI.

F3233.1570: Panama, Barro / Colorado Is., S-M / light trap, tree / canopy / 30.9.1977 / Dr. Henk Wolda,

STRI.

Taxonomy

Spongiphoridae, Spongiphorinae, Maravapallida Brindle, 1988.

pallidicolle Hincks, 1959 (Echinosoma)

Echinosomapallidicolle Hincks, 1959: 154, figs 143-144.

Type material

1 <S, paratype, F3233.461; 2 $ §, paratypes, F3233.462M63; dried, mounted on card.

Locality labels

F3233.461: Java / Coll. Gadeau / de Kerville.

F3233.462: Java / Mount Tengger / Coll. Gadeau / de Kerville.

F3233.463: East Java: / Soember Brantas / Feb. 1937. / M. E. Walsh.

Taxonomy

Pygidicranidae, Echinosomatinae, Echinosoma pallidicolle Hincks, 1959.

pallidus Brindle, 1968 ( Auchenomus )

Auchenomuspallidus Brindle, 1968h: 543, figs 4, 7-8, 10.

Type material

1 S, paratype, F3233.1501, dried, mounted on card.

Locality label

Virao / S. O. Luzon.

Taxonomy

Spongiphoridae, Sparattinae, Auchenomini, Auchenomus pallidus Brindle, 1968.

Remark

Index card entry: Philippine Islands, Luzon.

pandani Hincks, 1960 (Auchenomus)

Auchenomus pandani Hincks, 1960b: 157, figs 2-3.

Type material

1 S, paratype, F3233.554, dried, mounted on card.

Locality label

New Guinea / Yalu : Lae / i. 1958 / R. W. Paine. // On leaves of / Pandanus / (No. 4 sp.).

MILES C., Dermaptera of the Manchester Museum

Taxonomy

Spongiphoridae, Sparattinae, Auchenomini, Auchenomuspandani Hincks, 1960.

pandani Hincks, 1959 ( Tagalina )

Tagalinapandani Hincks, 1959: 114, fig. 99.

Type material

1 <$, paratype, F3233.58, dried, mounted on pin.

Locality label

Melambi R. Lae N. G, / Mirilunga Vill. 4500 / 29.12.56 on Pandanus / Coll. J H Ardley (New Guinea)

//C. I. F. Coll./No. 15242.

Taxonomy

Pygidicranidae, Pygidicraninae, Tagalina pandani Hincks, 1959.

parvula Brindle, 1988 ( Marava)

Fig. 48

Marava parvula Brindle, 1988: 54, fig. 17.

Type material

1 holotype, F3233.1814 (Fig. 48); 1 paratype, F3233.1815; 5 §§, paratypes, F3233.1816-1820;

1 paratype, gender unspecified, F3233.1821; dried, mounted on card.

Type locality

Panama, 1300m / Alto Lino, near /Boguete, Chiriqui / l.X. 1977 / Dr. Henk Wolda, STRI.

Locality labels

As type locality except dates: F3233.1815, 3 May 1977; F3233.1820, 7 May 1977; F3233.1816, 9 Oct.

1977; F3233.1821, 14 Oct. 1977; F3233.1817, 4 May 1978; F3233.1818, F3233.1819, 23 Jun. 1978.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava parvula Brindle, 1988.

Remark

The original description refers to 7 female paratypes at MMUE.

pauliani Hincks, 1948 ( Chaetospania )

Chaetospaniapauliani Hincks, 1948: 319, figs 2, 6.

Type material

1 S, paratype, F3233.1518, dried, mounted on card.

Locality label

Museum Paris / Cote dTvoire / Reserve du Banco / R. Paulian & C. Delamare.

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospaniapauliani Hincks, 1948.

peruviana Brindle, 1970 ( Skalistes )

Fig. 31

Skalistes peruviana Brindle, 1970b: 223, figs 5, 13.

Type material

1 3, holotype, F3233.1793 (Fig. 31); 1 ?, allotype, F3233.1794; 3 S3, paratypes, F3233.1795-1796 [2

specimens]; dried, mounted on card.

Type locality

N. Peru : Huambos / nr. Cutervo 2400m / 13.VI. 1956. / W. Weyrauch.

Locality labels

F3233.1794: Porculla-Peru /Pass (Weg von Olmos /nach Jaen) IF April / leg.: Koepcke. 1953.

F3233.1795-1796: As type locality.

Taxonomy

Forficulidae, Forficulinae, Skalistes peruviana Brindle, 1970.

philippinense Hincks, 1959 (Echi nosoma)

Echinosoma philippinense Hincks, 1959: 156, figs 148-150.

Type material

2 S3, paratypes, F3233.1328-1329; 2 5?, paratypes, F3233.1330-1331; dried, mounted on card.

Locality labels

F3233.1328-133F Philippine Is. / Mindoro, San / Jose X.1945 / E. S. Ross. // Ex Calif. / Ac. Scie.

Taxonomy

Pygidicranidae, Echinosomatinae, Echinosomaphilippinense Hincks, 1959.

picea Borelli, 1907 ( Gonolabis )

Gonolabis picea Borelli, 1907c: 1.

Type material

1 3, syntype, F3233.1399, dried, mounted on card.

Locality label

1906 Butiti / Viag. Ruwenzori / S. A. R. D. Abruzzi. ex coll. / Borelli.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis duplicata Steinmann, 1989;

MILES C., Dermaptera of the Manchester Museum

Remarks

1. Steinmann (1989a: 94) proposed a new name for Gonolabis picea Borelli, 1907, a junior homonym

of Gonolabis picea Shiraki, 1905.

2. Index card entry: Uganda.

pieli Hincks, 1940 ( Timomenus )

Fig. 29

Timomenus pieli Hincks, 1940: 38.

Type material

1 S, holotype, F3233.1797 (Fig. 29); 1 paratype, F3233.1798; dried, mounted on pin.

Type locality

Hainan / Nodoa 25.III.36 / G. Ros leg. // 43.

Locality label

F3233.1798: Hainan Is. S. China / Tsi-pin-tsuen Fam- / ks-heung, Fai-mo-ling / (Mt. Range), Ktung-

shan / District, May 17-26 / 1935. F. K. To.

Taxonomy

Forficulidae, Opisthocosmiinae, Timomenus pieli Hincks, 1940.

ponapensis Brindle, 1972 ( Chaetospania )

Chaetospania ponapensis Brindle, 1972c: 149, fig. 13g—i.

Type material

1 (?, paratype, F3233.607; 2 $ paratypes, F3233.608-609; dried, mounted on card.

Locality labels

F3233.607: Nanpil, Nett Dist. / Ponape I. / 25 Feb. 1948.

F3233.609: Same data except date 28 Feb. 1948.

F3233.608: Mt. Tama- / tamansakir / Ponape Id. / 1400’ / June - Sept. 50 / PA. Adams.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospaniaponapensis Brindle, 1972.

Remark

Index card entry: Micronesia, Ponape.

pophami Brindle, 1969 ( Vandex )

Fig. 51

Vandex pophami Brindle, 1969d: 190, fig. 1.

Type material

1 c?, holotype, F3233.1629 (Fig. 51), dried, mounted on card.

European Journal of Taxonomy 141: 1-138 (2015)

Type locality

Mufindi / Tanganyika / 1938.

Taxonomy

Spongiphoridae, Vandicinae, Vandexpophami Brindle, 1969.

porrectella Brindle, 1978 (Anisolabis)

Anisolabisporrectella Brindle, 1978: 86, figs 99-101, 116, 238.

Type material

1 <S, paratype, F3233.1400; 1 $, paratype, F3233.1401; dried, mounted on card.

Locality labels

F3233.1400-1401: Recolte dans Fhumus. I.R.S.A.C. - Mus. Congo / Kivu: T. Kalehe, Bitale / N. O.

Kahusi, 1600 m. / (for. transit.) 29-VI-1951 /N. Leleup.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis porrectella Brindle, 1978.

Remark

The original description gives the locality as Kahele Territory, not Kalehe as on the locality label. Bitale

is in Kalehe Territory, South Kivu Province, Democratic Republic of Congo.

prolongata Hincks, 1947 (Sphingolabis)

Sphingolabisprolongata Hincks, 1947b: 21, figs 12-13.

Type material

1 S, paratype, F3233.1599, dried, mounted on card.

Locality label

N. E. Burma / Kambaiti 7,000 ft. / 12/4 R. Malaise.

Taxonomy

Spongiphoridae, Labiinae, Sphingolabis prolongata Hincks, 1947.

Remarks

1. Hopkins et al. (2014) has ‘ prolonga

2. The original description gives the full date as 12 Apr. 1934.

pulchellus Brindle, 1977 ( Purex )

Purexpulchellus Brindle, 1977b: 118, figs 9-11.

Type material

1 $, paratype, F3233.1596; 1 $, paratype, F3233.1597; dried, mounted on card.

MILES C., Dermaptera of the Manchester Museum

Locality labels

F3233.1596: Venezuela - Aragua / Rancho Grande / 1100m. 20.V.75. // En la / Luz. // J. Salcedo / J. A.

Clavijo.

F3233.1597: Same data except date 2 Apr. 1975 and collector J. Salcedo.

Taxonomy

Spongiphoridae, Spongiphorinae, Purexpulchellus Brindle, 1977.

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Figs 49-58. Holotypes (S) of Spongiphoridae in the collection of the Manchester Museum. 49. Aucheno-

mus variabilis guadalcanalensis Brindle, 1970. 50. Labia africana Brindle, 1968 (now Pseudovostox

africanus). 51. Vandex pophami Brindle, 1969. 52. Pericomus ater Brindle, 1988. 53. Pseudovostox

africanus sudanensis Brindle, 1970. 54. P. hincksi Brindle, 1970. 55. Strongylopsalis koepekei

Brindle, 1968. 56. Labia rotundifrons Hinclcs, 1954 (now Paralabella rotundifrons). 57. Chaetospania

meridionalis Brindle, 1973. 58. Nesogaster javanicus Hincks, 1951 (now N. aculeatus). Scale bars = 1

cm.

European Journal of Taxonomy 141: 1-138 (2015)

pusilla Brindle, 1978 ( Bormansia )

Bormansia pusilla Brindle, 1978: 189, fig. 347.

Type material

1 S, paratype, F3233.1337, dried, mounted on card.

Locality label

Ostafrilca: T. T. / Msala (Rufiji) / 27.IV. 1952 / H. Knipper leg.

Taxonomy

Karschiellidae, Bormansia pusilla Brindle, 1978.

Remark

Index card entry: Tanzania.

pygidiata Brindle, 1988 ( Marava )

Fig. 43

Marava pygidiata Brindle, 1988: 54, fig. 23.

Type material

1 S, holotype, F3233.1822 (Fig. 43); 2 §§, paratypes, F3233.1823-1824; dried, mounted on card. In

spirit: 2 paratypes [gender not specified], F3448.31.

Type locality

Panama, Fortuna / Wet forest / at light, 1050m / 15.12.1977 / Dr. Henk Wolda, STRI.

Locality labels

As type locality except date: F3233.1823, 24 Sep. 1977; F3233.1824, 25 Sep. 1977; F3448.31, 1977-

1978.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava pygidiata Brindle, 1988.

Remark

The original description (Brindle 1988) refers to one male and 12 female paratypes at MMUE with a

date range of 12 Aug. 1977-3 Jun. 1979.

pygidiatus Brindle, 1975 ( Vandex )

Vandexpygidiatus Brindle, 1975b: 687, figs 5-7.

Type material

1 <S, paratype, F3233.1630; 1 §, paratype, F3233.1631; dried, mounted on card.

Locality labels

F3233.1630: Tanzanie : Mts Uluguru / Morning Side, Toelo / for., alt. 1450 m / 21—29/VI/71.

F3233.1631: Tanzanie : Mts Uluguru / Chenzema, alt. 1700m. / 2-22/VII/71.

MILES C., Dermaptera of the Manchester Museum

Taxonomy

Spongiphoridae, Vandicinae, Vandex pygidiatus Brin die, 1975.

pygmaea Brindle, 1970 ( Metresura )

Metresura pygmaea Brindle, 1970a: 105, fig. 9.

Type material

1 S, paratype, F3233.1741; 1 paratype, F3233.1742; dried, mounted on card.

Locality labels

F3233.1741: Fa Venta-D.F. /Mexico / 17.XII. 1961 /PE. Pereira Col.

F3233.1742: Desierto Eos Feones / 2900 m. D.F. Mexico / II - 1962 / Olguin E Pereira.

Taxonomy

Forficulidae, Neolobophorinae, Metresura pygmaea Brindle, 1970.

quadricollis Hincks, 1954 ( Apolabis )

Apolabis quadricollis YLincks, 1954b: 106, fig. 6.

Type material

1 S, paratype, F3233.1402, dried, mounted on card.

Locality label

Museum Paris / Nimba (Guinee) / II. VI. 42. Pierre Richaud (900m).

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis quadricollis (Hincks, 1954).

Remarks

1. Transferred by Brindle (1978: 114).

2. Index card entry: Guinea, Nimba Mts.

quadrimaculata Brindle, 1971 ( Sarcinatrix )

Sarcinatrix quadrimaculata Brindle, 197lg: 169, figs 12-16.

Type material

1 S, paratype, F3233.1746, dried, mounted on card.

Locality label

Est. Biol. Boraceia / Salesopolis - S. P. / Brasil - 13 - 18.IV.961 / Reichardt col.

Taxonomy

Forficulidae, Ancistrogastrinae, Sarcinatrix quadrimaculata Brindle, 1971.

Remark

Index card entry: Brazil, Sao Paulo, Esta£ao Biologica de Boraceia, Salesopolis.

European Journal of Taxonomy 141: 1-138 (2015)

rehni Hincks, 1951 ( Nesogaster)

Nesogaster rehni Hincks, 1951b: 573, fig. 17.

Type material

2 SS, paratypes, F3233.537, dried, pinned, double mounted on the same pin.

Locality label

Queensland / Kuranda / Dodd.

Taxonomy

Spongiphoridae, Nesogastrinae, Nesogaster rehni Hincks, 1951.

reichardti Brindle, 1971 ( Mecomera )

Mecomera reichardti Brindle, 197lg: 166, figs 1-7.

Type material

1 c?, paratype, F3233.1520; 1 $, paratype, F3233.1521; dried, mounted on card.

Locality labels

F3233.1520-1521: St. Catharina / Blumenau / [illegible] / 20.339 /11.19.

Taxonomy

Spongiphoridae, Sparattinae, Sparattini, Mecomera reichardti Brindle, 1971.

Remark

Index card entry: Blumenau, Santa Catarina, Brazil.

rhodesiensis Brindle, 1973 ( Afrocosmia )

Afrocosmia rhodesiensis Brindle, 1973f: 288, figs 389-391.

Type material

1 S, paratype, F3233.1679; 1 5, paratype, F3233.1680; dried, mounted on card.

Locality labels

F3233.1679: Recolte dans / Fhumus // I.R.S.A.C.-Mus. R.A.C. / S. Rhodesia : Mt Selinda, / foret de

Chirinda, 1300 m. /VII. 1960 N. Leleup.

F3233.1680: Vase d’une / tete de source / marecageuse // I.R.S.A.C.-Mus. R.A.C. / S. Rhodesia : Mt

Selinda, / foret de Chirinda, 1300 m. / VII. 1960 N. Leleup.

Taxonomy

Forficulidae, Skendylinae, Afrocosmia rhodesiensis Brindle, 1973.

rileyi Hincks, 1955 ( Diplatys )

Diplatys rileyi Hincks, 1955a: 74, figs 75-78.

MILES C., Dermaptera of the Manchester Museum

Type material

1 §, paratype, F3233.1305, dried, mounted on card.

Locality label

Sikkim: Singhik/ 3,500 ft. 23.iv. 1924. /Maj. R. W. G. Hingston.

Taxonomy

Diplatyidae, Diplatyinae, Haplodiplatys rileyi (Hincks, 1955).

Remark

Transferred by Steinmann (1974: 203).

rostrata Brindle, 1978 ( Anisolabis )

Anisolabis rostrata Brindle, 1978: 87, figs 107-109, 237.

Type material

1 3, paratype, F3233.1403, dried, mounted on card.

Locality label

Biot. No. 95 / Humus / en foret. Mus. Roy. Afr. Centr. / Tshuapa : Terr. Ikela, / riv. Lukendu, VIII-1959

/ B. 95 N. Leleup.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis rostrata Brindle, 1978.

rostratus Brindle, 1984 {Esphalmenus)

Esphalmenus rostratus Brindle, 1984: 288, fig. 27.

Type material

1 c?, paratype, F3233.1363; 1 5, paratype, F3233.1364; dried, mounted on card.

Locality labels

F3233.1363-1364: Argentina / Tierra del Fuego / Estancia Viamonte / Auricosta 23.1.1979 / Mision

Cientifica Danesa.

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus rostratus Brindle, 1984.

rotundicollis Hincks, 1952 ( Diplatys )

Diplatys rotundicollis Hincks, 1952b: 19, figs 1-6.

Type material

2 S3, paratypes, F3233.1307, F3233.1849; 2 paratypes, F3233.1308-1309; dried, mounted on

card.

European Journal of Taxonomy 141: 1-138 (2015)

Locality labels

F3233.1307: Sierra Leone / Njala // 7.III.32. / Hargreaves.

F3233.1308: Njala / Sierra Leone / E. Hargreaves. / Date 24-V-26.

As F3233.1307 except dates: F3233.1309, 2 Mar. 1932; F3233.1849, 11 May 1932.

Taxonomy

Diplatyidae, Diplatyinae, Paradiplatys rotundicollis (Hincks, 1952).

Remarks

Transferred to Lobodiplatys ( Heterodiplatys ) by Steinmann (1989c: 79). The combination Paradiplatys

(Heterodiplatys) (Hincks) was reinstated by Engel & Haas (2007: 19).

rotundifrons Hincks, 1954 {Labia)

Fig. 56

Labia rotundifrons Hincks, 1954e: 20, figs 14-15.

Type material

1 S, holotype, F3233.1544 (Fig. 56), dried, mounted on card.

Type locality

Ceylon: / Newara Eliya / in a garden / 6300 ft 19.8.36 / Walford.

Taxonomy

Spongiphoridae, Labiinae, Paralabella rotundifrons (Hincks, 1954).

Remarks

1. Transferred by Ste inm a nn (1989b: 496).

2. The original description gives the locality as Nuwara. Newara Eliya (locality label) is a city in Central

Province, Sri Lanka.

rougemonti Brindle, 1978 ( Anisolabis )

Fig. 26

Anisolabis rougemonti Brindle, 1978: 135, figs 225-227, 270.

Type material

1 S, holotype, F3233.1404 (Fig. 26); 1 §, allotype, F3233.1405; dried, mounted on card.

Type locality

Ethiopia / Arossi Province / Sodere, VII-1971 / G. de Rougemont.

Locality label

F3233.1405: As type locality.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis rougemonti Brindle, 1978.

MILES C., Dermaptera of the Manchester Museum

Remark

The original description gives the locality as Sodare, not Sodere as on the specimen label. Arussi

Province corresponds to the present day Arsi Zone of Ethiopia.

rouxioides Brindle, 1976 ( Brachylabis )

Brachylabis rouxioides Brindle, 1976b: 74, figs 9, 15, 17.

Type material

1 3, paratype, F3233.119, dried, mounted on pin.

Locality label

New Caledonia. /P.D. Montague. / 1918-87. Loc. MtMou477/Date 10.3.14 Sex/Coll. P.D. Montague.

/ New Caledonia Exped.

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis rouxioides (Brindle, 1976).

Remarks

Transferred by Steinmann (1989a: 361) ns Antisolabis rouxiodes. There are inconsistencies in the spelling

of rouxiodes/rowcioides in both Steinmann 1989a and Steinmann 1989c, each publication containing

both spellings. Hopkins et al. (2014) has ‘ rouxiodes ’ and gives ‘ rouxioides ’ as a synonym, but with a

query. Brindle’s original spelling was ‘ rouxioides \

royi Brindle, 1971 ( Anisolabis )

Anisolabis royi Brindle, 1971c: 268, figs 1—4, 8.

Type material

2 33, paratypes, F3233.1406-1407; 3 §§, paratypes, F3233.1408-1410; dried, mounted on card.

Locality labels

F3233.1406-1410: Foret Camp / 1070 m / 26. V. 1963. Mission ENS-IFAN / aux Monts Loma / Sierra

Leone.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis royi Brindle, 1971.

Remark

The original description refers to 12 paratypes at MMUE.

rubescens Brindle, 1973 {Spongovostox)

Spongovostox rubescens Brindle, 1973f: 222, figs 296-297, 300, 306.

Type material

1 5, paratype, F3233.1611, dried, mounted on card.

European Journal of Taxonomy 141: 1-138 (2015)

Locality label

Bitale Kivu 1954 / R. Pierlot / RP. 48a. // Coll. R. Mayne / Com. Et. Bois Congo R. 2558.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox rubescens Brindle, 1973.

rudebecki Brindle, 1969 ( Pseudovostox )

Pseudovostox rudebecki Brindle, 1969c: 147, figs 1-5, 9.

Type material

1 8, paratype, F3233.1595, dried, mounted on card.

Locality label

At light in / the evening // Afr. Angola / Vila Artur de Paiva / 5-8.X. 1956 / coll. G. Rudebeck.

Taxonomy

Spongiphoridae, Geracinae, Pseudovostox rudebecki Brindle, 1969.

ruficollis Hincks, 1957 ( Brachylabis )

Brachylabis ruficollis Hincks, 1957b: 69, figs 29-30.

Type material

2 88, paratypes, F3233.1427-1428; 2 5?, paratypes, F3233.1429-1430; dried, mounted on card.

Locality labels

F3233.1427: Salisbury S. A. / JAN. 1954/R. M. Arnold.

F3233.1428: South Africa / Grahamstown/ 28.9 49 / (R. Bigalke).

F3233.1429: Grahamstown/ July 1951 / coll: V. B Brigg.

F3233.1430: Cape Province / Grahamstown / 8.1951 /CD Quickelbirg.

Taxonomy

Anisolabididae, Brachylabidinae, Ctenisolabis ruficollis (Hincks, 1957).

Remark

Transferred by Steinmann (1989a: 292).

rugosa Brindle, 1966 ( Brachylabis )

Brachylabis rugosa Brindle, 1966e: 38, fig. 12.

Type material

1 8, paratype, F3233.1476; 1 §, paratype, F3233.1477; dried, mounted on card.

Locality labels

F3233.1476-1477: Vest. Foret / ombrophile / dans l’humus // Coll. Mus. Congo / Tanganyika Terr.;

Bundu- / ki, Uluguru Mts., 1300 m. / 2-V-1957 // Mission Zoolog. I.R.S.A.C. / en afrique orientale / (P.

Basilewsky et / N. Feleup).

100

MILES C., Dermaptera of the Manchester Museum

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis rugosa (Brindle, 1966).

Remark

Transferred by Steinmann (1989a: 347).

rugosa Brindle, 1975 ( Metresura)

Fig. 28

Metresura rugosa Brindle, 1975d: 84.

Type material

1 <S, holotype, F3233.1743 (Fig. 28); 1 $, allotype, F3233.1744; dried, mounted on card.

Type locality

Salvador, Isalco / volcano, forest / 1200 m., foot of / Cerro Verde (Mala / Cara) Hac. Buena / Vista

26.7.1951.

Locality label

F3233.1744: As type locality.

Taxonomy

Forficulidae, Neolobophorinae, Metresura rugosa Brindle, 1975.

rugosula Hincks, 1957 ( Forficula )

Forficula rugosula Hincks, 1957b: 89, fig. 37.

Type material

1 c?, paratype, F3233.1725, dried, mounted on card.

Locality label

R. Howell / Grahamstown / Aug 52.

Taxonomy

Forficulidae, Forficulinae, Guanchia rugosula (Hincks, 1957).

Remark

Transferred by Steinmann (1993: 557).

rugosus Brindle, 1971 ( Anisolabis )

Anisolabis rugosus Brindle, 1971c: 274, figs 16-19.

Type material

1 S, paratype, F3233.1411, dried, mounted on card.

Locality label

Camp a 800 m / foret primaire / 17. V. 1963. Mission ENS-IFAN / aux Monts Foma / Sierra Feone.

101

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis rugosa Brindle, 1971.

Remark

Brindle (1971c: 266) refers to two paratypes at MMUE.

rugosus Brindle, 1975 (Archidux)

Archidux rugosus Brindle, 1975b: 693.

Type material

1 <S, paratype, F3233.1708; 1 §, paratype, F3233.1709; dried, mounted on card.

Locality labels

F3233.1708-1709: Tanzanie : Mts Uluguru / Kinola, for. transition / alt. 1500-1750 m. / 6—13/VI/71 //

Coll. Mus. Tervuren / Mission Mts Uluguru / L. Berger, N. Leleup / J. Debecker V/VIII/71.

Taxonomy

Forficulidae, Skendylinae, Cosmiella rugosa (Brindle, 1975).

Remark

Transferred by Steinmann (1989c: 674).

saegeri Brindle, 1968 ( Nala )

Nala saegeri Brindle, 1968e: 21, figs 11-12.

Type material

2 $$, paratypes, F3233.1495, F3233.1497; 2 paratypes, F3233.1496, F3233.1498; dried, mounted

on card.

Locality labels

F3233.1495, F3233.1497: Congo beige, P. N. G. / Miss. H. De Saeger /K. 17, 18-iv-1950 / Rec. H. De

Saeger. 432.

F3233.1496: Ssame data except date and number, 10 May 1950 and 504.

F3233.1498: Same data as F3233.1496 except number, 506.

Taxonomy

Labiduridae, Nalinae, Nala saegeri Brindle, 1968.

salcedoi Brindle, 1974 ( Eugerax )

Eugerax salcedoi Brindle, 1974b: 117, figs 9, 13.

Type material

1 c?, paratype, F3233.1534; 1 paratype, F3233.1535; dried, mounted on card.

102

MILES C., Dermaptera of the Manchester Museum

Locality labels

F3233.1534:Venezuela-Aragua/ Rancho Grande / 1100m. 15-22.VII.74// Col. JonH. Soeder.

F3233.1535: Same data except date and collectors: 19.VII.73 // J. Salcedo / J. A. Clavijo.

Taxonomy

Spongiphoridae, Geracinae, Eugerax salcedoi Brindle, 1974.

Salvadorensis Brindle, 1971 {Spongiphora)

Spongiphora salvadorensis Brindle, 1971b: 531, figs 15, 19.

Type material

1 (?, paratype, F3233.1600; 1 §, paratype, F3233.1601; dried, mounted on card.

Locality labels

F3233.1600-1601: El Salvador: Mt. / San Salvador / 4000’ - 6400’ / VII-8-1963. // D. Q. Cavagnaro /

& M. E. Irwin / collectors.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongiphora salvadorensis Brindle, 1971.

Remark

The original description gives the collection date as 1968 (possible misreading of 1963).

salvazae Burr, 1917 ( Diplatys )

Diplatys salvazae Burr, 1917: 57.

Type material

1 c?, paratype, F3233.1299; 1 5, paratype, F3233.1300; dried, mounted on card.

Locality labels

F3233.1299-1300: Cambodia / Pnom Penh / iv 13 Vitalis. // Burr Coll. B. M. 1915 - 120.

Taxonomy

Diplatyidae, Diplatyinae, Paradiplatys salvazae (Burr, 1917).

Remarks

Transferred to Lobodiplatys ( Paradiplatys ) salvazae (Burr, 1917) by Steinmann (1986a: 178). The

combination Paradiplatys (.Paradiplatys ) salvazae (Burr) was reinstated by Engel & Haas (2007: 20).

schwarzi var. nigrinus Brindle, 1988 {Spongovostox)

Spongovostox schwarzi var. nigrinus Brindle, 1988: 46, 51, fig. 12.

Type material

1 paratype [gender unspecified], F3233.1866; dried, mounted on card.

103

European Journal of Taxonomy 141: 1-138 (2015)

Locality label

Panama, Barro / Colorado Is., S-M / light trap, ground / level 5.5.1978 / Dr Henk Wolda, STRI.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox schwarzi (Caudell, 1907) var. nigrinus Brindle, 1988.

scotti Brindle, 1978 ( Anisolabis )

Anisolabis scotti Brindle, 1978: 131, figs 195-196, 260.

Type material

1 S, paratype, F3233.1412, dried, mounted on card, without head.

Locality label

By sifting humus under trees. Abyssinia: / Mt. Chillalo, / forest, / circa 9,000ft. / 13.xi. 1926. / Dr. H.

Scott.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis scotti Brindle, 1978.

setulosum Hincks, 1959 ( Echinosoma )

Echinosoma setulosum Hincks, 1959: 155, figs 145-147.

Type material

1 S, paratype, F3233.1332; 1 §, paratype, F3233.1333; dried, mounted on card.

Locality labels

F3233.1332: Tonkin / Than-Moi / Juni-Juli / H. Fruhstorfer / Ex Coll. Gadeau / de Kerville.

F3233.1333: Annam / Phuc-Son / Nov. Dez. / H. Fruhstorfer / Coll. Gadeau / de Kerville.

Taxonomy

Pygidicranidae, Echinosomatinae, Echinosoma setulosum Hincks, 1959.

seychellensis Brindle, 1976 ( Brachylabis )

Brachylabis seychellensis Brindle, 1976a: 439, figs 1-3.

Type material

1 S, paratype, F3233.1431; 1 $, paratype, F3233.1432; dried, mounted on card.

Locality labels

F3233.1431-1432: lies Seychelles - Mahe / Centre: Morne Blanc / foret endemique, / 667 m 8/25.

VIII. 1972.

Taxonomy

Anisolabididae, Antisolabidinae, Antisolabis seychellensis (Brindle, 1976).

104

MILES C., Dermaptera of the Manchester Museum

Remark

Transferred by Steinmann (1989a: 350).

simiensis Brindle, 1978 (Anisolabis)

Anisolabis simiensis Brindle, 1978: 134, figs 231-234.

Type material

1 8, paratype, F3233.1413, dried, mounted on card.

Locality label

Ethiopia, c. 3000m, / Simien Mountains / between Debark and / Adesghie. 22.X. 1973 / G. de Rougemont.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis simiensis Brindle, 1978.

Remark

The original description gives the locality as ‘near Derasghie’.

simplex Brindle, 1966 (Archidux)

Archidux simplex Brindle, 1966e: 47, figs 23-25.

Type material

2 88, paratypes, F3233.1710-1711, dried, mounted on card.

Locality labels

F3233.1710-1711: Foret / transition, / dans l’humus // Coll. Mus. Congo / Tanganyika Terr. : Bundu- /

ki, Uluguru Mts., 1500 m. / gorge Mungula 1/6-V-1957. // Mission Zoolog. I.R.S.A.C. / en Afrique

orientale / (P. Basilewsky et / N. Feleup).

Taxonomy

Forficulidae, Skendylinae, Paracosmiella simplex (Brindle, 1966).

Remark

Transferred by Steinmann (1993: 166).

sogai Brindle, 1966 ( Emboros )

Emboros sogai Brindle, 1966g: 252, figs 58, 64.

Type material

1 5, paratype, F3233.1687, dried, mounted on card.

Locality label

Madagascar-Est / dct. Andapa / Anjanaharibe 1600m / IV-60 R Soga.

105

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Forficulidae, Skendylinae, Brachycosmiella sogai (Brindle, 1966).

Remark

Transferred by Steinmann (1993: 165).

solomonensis Brindle, 1970 (Epilandex)

Epilandex solomonensis Brindle, 1970f: 654, figs 9-13.

Type material

5 S3, paratypes, F3233.127-131; 4 5?, paratypes, F3233.132-135; dried, mounted on card.

Locality labels

F3233.127-128, F3233.130-135: Solomon Is. Guadacanal [sic] /Mt. Austen/ Jan. 1966 /P. Greenslade.

F3233.129: Same data except date Sep.-Oct. 1965.

Taxonomy

Anisolabididae, Anisolabididae, Epilandex solomonensis Brindle, 1970.

spicata Brindle, 1972 ( Chaetolabia )

Chaetolabia spicata Brindle, 1972c: 153, fig. 14g-h.

Type material

1 S, paratype, F3233.618, dried, mounted on card.

Locality label

Palau: Peleliu / N. E. Coast / 28 Jan 1948 / H.S. Dybas.

Taxonomy

Spongiphoridae, Labiinae, Chaetolabia spicata Brindle, 1972.

spiculifera Brindle, 1977 (Parasparatta)

Parasparatta spiculifera Bnnd\Q, 1977b: 116, fig. 20.

Type material

1 S, paratype, F3233.1524, dried, mounted on card.

Locality label

Venezuela Aragua / Rancho Grande / 1100 m 11.1.75.

Taxonomy

Spongiphoridae, Sparattinae, Sparattini, Sparatta spiculifera (Brindle, 1977).

Remark

Transferred by Steinmann (1989b: 155).

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MILES C., Dermaptera of the Manchester Museum

spinosci Brindle, 1966 ( Allodahlia )

Allodahlia spinosa Brindle, 1966h: 444, figs 7-9, 11-12.

Type material

1 S, paratype, F3233.1696, dried, mounted on card.

Locality label

Pt Bango / Mindanao.

Taxonomy

Forficulidae, Allodahliinae, Allodahlia ancylura (Dohrn, 1865).

Remark

Allodahlia spinosa Brindle, 1966 was synonymised with Allodahlia ancylura (Dohrn, 1865) by Brindle

(1972b: 29).

srivastavai Brindle, 1981 ( Capralabia )

Capralabia srivastavai Brindle, 1981: 79.

Type material

1 S, paratype, F3233.1438, dried, mounted on card.

Locality label

Panama, Barro / Colorado Is., S-M / light trap, ground / level / 14.11.1978 / Dr. Henk Wolda, STRI.

Taxonomy

Anisolabididae, Anisolabidinae, Capralabis srivastavai Brindle, 1981.

Remark

The genus is named Capralabia in the original description.

steinmanni Brindle, 1981 ( Capralabis )

Capralabis steinmanni Brindle, 1981: 80.

Type material

1 5, paratype, F3233.1439, dried, mounted on card.

Locality label

Venezuela Ara- / gua, El Fimon. / 450 m 13-IX-1978.

Taxonomy

Anisolabididae, Anisolabidinae, Capralabis steinmanni Brindle, 1981.

Remark

The original description refers to a second female paratype at MMUE.

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European Journal of Taxonomy 141: 1-138 (2015)

stemmleri Brindle, 1975 ( Diplatys )

Diplatys stemmleri Brindle, 1975c: 10, figs 4-6.

Type material

1 S, paratype, F3233.1310, dried, mounted on card.

Locality label

21 km O Wangdi / Phodr. 1700 - 2000 / Nat.-Hist. Museum / Basel - Bhutan / Expedition 1972.

Taxonomy

Diplatyidae, Diplatyinae, Haplodiplatys stemmleri (Brindle, 1975).

Remarks

1. Transferred by Ste inm a nn (1982a: 211).

2. The original description gives the locality as Wangdiphodrang and the date as 15 Jun. 1972.

straeleni Hincks, 1955 ( Apolabis )

Apolabis straeleni Hincks, 1955b: 7, figs 1-2.

Type material

1 <S, paratype, F3233.1414; 1 paratype, F3233.1315; dried, mounted on card.

Locality labels

F3233.1414: Congo beige : P.N.U. / Kaswabilenga (r. Dr. Lufira) / (700 m.) 6-9-X-1947 / Mis. G. F. De

Witte. 805a.

F3233.1415: Same data except date, 18-23 Sep. 1947, and number, 769a.

Taxonomy

Anisolabididae, Anisolabidinae, Anisolabis straeleni (Hincks, 1955).

Remarks

1. Transferred by Brindle (1978: 96).

2. Index card entry: Africa, Zaire, Kaswebilenga, Upemba Nat. Park.

submetallica Brindle, 1966 ( Brachylabis )

Brachylabis submetallica Brindle, 1966e: 40, figs 8, 14.

Type material

1 S , paratype, F3233.1478; 1 $, paratype, F3233.1479; dried, mounted on card.

Locality labels

F3233.1478-1479: Recolte dans / racines / de roseaux // I.R.S.A.C. Mus. Congo / Kivu : Terr. Uvira /

Rives L. Tanganika/ 14-11-1952 /N. Leleup.

Taxonomy

Anisolabididae, Brachylabidinae, Metisolabis submetallica (Brindle, 1966).

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MILES C., Dermaptera of the Manchester Museum

Remark

Transferred by Steinmann (1989a: 282).

sumatrana Brindle, 1965 ( Allostethella )

Allostethella sumatrana Brindle, 1965: 591, figs 3, 10, 20.

Type material

1 5, paratype, F3233.1490, dried, mounted on card.

Locality label

Sumatra’s OK/ Sibolangit.

Taxonomy

Labiduridae, Allostethinae, Allostethella sumatrana Brindle, 1965.

sumatrana Borelli, 1927 ( Chaetospania )

Chaetospania sumatrana Borelli, 1927: 71, figs 1-3.

Type material

3 $$, 3 55, syntypes, F3233.1519, dried, mounted on card.

Locality label

Gunung Singgalang / Sumatra’s Westkust / M. 192 / leg. E. Jacobson.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania sumatrana Borelli, 1927.

surinamensis Brindle, 1968 ( Larex )

Larex surinamensis Brindle, 1968g: 34, figs 28-29.

Type material

1 <$, paratype, F3233.1554, dried, mounted on card.

Locality label

Mus. Leiden/D. C. Geyskes. /Waremapau/ l-VIII-1939.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava surinamensis (Brindle, 1968).

Remarks

1. Transferred by Brindle (1971b: 562).

2. Index card entry: Surinam.

3. The locality is given as Waremapan in the original description, not Waremapau (locality label).

(Waremapan Kreek, Sipaliwini, is in the southeastern corner of Suriname.)

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European Journal of Taxonomy 141: 1-138 (2015)

sutteri Hincks, 1954 {Labia)

Labia sutteri Hincks, 1954e: 17, figs 8-11.

Type material

1 S, paratype, F3233.668; 1 §, paratype, F3233.667; dried, mounted on card.

Locality labels

F3233.667-668: O. Sumba / Mau Marru / 19.7.1949 / Buhler & Sutter.

Taxonomy

Spongiphoridae, Labiinae, Paralabella sutteri (Hincks, 1954).

Remark

Transferred by Steinmann (1989b: 488).

tanganyikae Brindle, 1964 ( Gelotolabis )

Fig. 21

Gelotolabis tanganyikae Brindle, 1964: 754, figs 1-2, 15.

Type material

1 <S, holotype, F3233.1416 (Fig. 21); 3 §§, paratypes, F3233.1317-1319; dried, mounted on card.

Type locality

Njombe / 6000-6500 ft. / Tanganyika / I.II. 1957 W. P.

Locality labels

As type locality except date: F3233.1317—1318, 26 Feb. 1957; F3233.1319, 21 Aug. 1952, Dr W. Peters.

Taxonomy

Anisolabididae, Anisolabidinae, Gonolabis tanganyikae (Brindle, 1964).

Remarks

1. Transferred by Steinmann (1978: 193).

2. The date of the holotype does not match that given for the material examined by Brindle in the original

description, i.e., ‘26.ii.57 (holotype, allotype)’, although the rest of the locality data matches. There is

no reference to female paratypes examined except the allotype. The female specimens F3233.1317-

1319 have ‘Paratype’ on Brindle’s determination labels.

taurinus Brindle, 1982 ( Diplatys )

Diplatys taurinus Brindle, 1982b: 163, figs 4-5.

Type material

1 5, paratype, F3233.1314, dried, mounted on card.

Locality label

Tanzania / E. Usambara Mts. / Amani 1000m / 20.VII. 1980.

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MILES C., Dermaptera of the Manchester Museum

Taxonomy

Diplatyidae, Diplatyinae, Diplatys taurinus Brindle, 1982.

tenebrosus Brindle, 1968 ( Hamaxas )

Hamaxas tenebrosus Brindle, 1968h: 546, figs 15, 23.

Type material

1 <$, paratype, F3233.1677; 1 paratype (gender unspecified), F3233.1678; dried, mounted on card.

Locality labels

F3233.1677-1678: Dapa/ Siargao.

Taxonomy

Chelisochidae, Chelisochinae, Hamaxas tenebrosus Brindle, 1968.

Remark

The original description refers only to the holotype, allotype and one male paratype, and states that

holotype and allotype are in the Riksmuseum, Stockholm.

tibetanus Hincks, 1955 ( Diplatys )

Diplatys tibetanus Hincks, 1955a: 76, figs 81-83.

Type material

1 S, paratype, F3233.1306; 1 §, paratype, F3233.1311; dried, mounted on pin.

Locality labels

F3233.1306, F3233.1311: Tibet: / Rongshar Valley. / 9,500ft. 25.vi.1924. /Maj. R. W. G. Hingston.

Taxonomy

Diplatyidae, Diplatyinae, Haplodiplatys tibetanus (Hincks, 1955).

Remarks

1. Transferred by Ste inmann (1974: 204).

2. The original description has the date 24 Jun. 1924, not 25 Jun. 1924. Both specimens have the word

‘Paratype’ on Hincks’ determination labels.

tibialis Hincks, 1953 ( Chaetospania )

Chaetospania tibialis Hincks, 1953: 371, figs 10-12.

Type material

1 S, paratype, F3233.1525, dried, mounted on card.

Locality label

Madagascar C. / Perinet 8.46 / Prof. J. Millot.

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European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania tibialis Hincks, 1953.

tigrina Brindle, 1967 ( Echinosoma )

Echinosoma tigrina Bundle, 1967b: 155, figs 18-21.

Type material

1 S, paratype, F3233.87; 1 §, paratype, F3233.88; dried, mounted on pin.

Locality labels

F3233.87-88: Long / Navang / O. Borneo / Mjoberg.

Taxonomy

Pygidicranidae, Echinosomatinae, Echinosoma tigrina Brindle, 1967.

townesi Brindle, 1979 ( Marava )

Marava townesi Brindle, 1979: 152, fig. 2.

Type material

1 S, paratype, F3233.1555, dried, mounted on card.

Locality label

Guatemala / 37 16 56 / Primavera logs / under bark / N.Y. Jan. 18, 1937.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava townesi Brindle, 1979.

transversa Brindle, 1978 ( Isolabis )

Isolabis transversa Brindle, 1978: 41, figs 35-36.

Type material

1 S, paratype, F3233.1480, dried, mounted on card.

Locality label

Hot for. marec. / avec Pandanus // I.R.S.A.C.-Mus. Congo / Kwango: terr. De Feshi, / rive dr. Kwenge

IV-1959 /B. 78 Mme J. Leleup.

Taxonomy

Anisolabididae, Isolabidinae, Isolabis transversa Brindle, 1978.

Remark

The original description gives the date as Mar. 1959, not Apr. 1959 (locality label).

triangulata Hincks, 1950 (Forficula)

Forficula triangulata Hincks, 1950a: 180, figs 3-4.

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MILES C., Dermaptera of the Manchester Museum

Type material

2 SS, paratypes, F3233.1726-1727; 1 §, paratype, F3233.1728; dried, mounted on card.

Locality labels

F3233.1726-1728: Kilimanjaro T.T. / 14,500 ft. / 27.X.48 Dr. G. Salt. // Under & beside stones on

saddle.

Taxonomy

Forficulidae, Forficulinae, Guanchia triangulata (Hincks, 1950).

Remark

Transferred by Steinmann (1993: 561).

triangulatus Brindle, 1987 ( Haplodiplatys )

Haplodiplatys triangulatus Brindle, 1987: 335, figs 1-3.

Type material

1 S, paratype, F3233.1315, dried, mounted on card.

Locality label

E. Nepal: / Aran Valley / VI-15-1954 / 7,000 ft. F. Swan.

Taxonomy

Diplatyidae, Diplatyinae, Haplodiplatys triangulatus Brindle, 1987.

trinodosa Brindle, 1967 ( Gonolabina )

Fig. 23

Gonolabina trinodosa Brindle, 1967c: 9, figs 2, 4.

Type material

1 c?, holotype, F3233.1454 (Fig. 23), dried, mounted on card. 1 §, allotype, F3233.1455, dried, mounted

on pin.

Type locality

Tragadero / (cerca Jauja) / 3500 m / 3.IV.49 / Coll. Blancas

Locality label

F3233.1455: Acolla (cerca / Jauja) 3460 m / VIII.49 / Coll: F. Blancas.

Taxonomy

Anisolabididae, Gonolabininae, Gonolabina trinodosa ^Brindle, 1967.

Remark

The original description has the altitude 3600 m for the allotype, not 3460 m.

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European Journal of Taxonomy 141: 1-138 (2015)

trinodosci Brindle, 1977 ( Pygidicrana )

Pygidicrana trinodosa Brindle, 1977b: 111, figs 1-3.

Type material

1 S, paratype, F3233.1319, dried, mounted on card.

Locality label

Curimagua / Venezuela, Falcon / 16.VIII. 1975 / R. E. Dietz. / Leg.

Taxonomy

Pygidicranidae, Pygidicraninae, Pygidicrana trinodosa Brindle, 1977.

Remark

The original description has the date 16 Dec. 1975, not 16 Aug. 1975.

trituberculatus Brindle, 1971 ( Purex )

Purex trituberculatus Brindle, 1971a: 171, fig. 24.

Type material

1 S, paratype, F3233.1598, dried, mounted on pin.

Locality label

Venezuela: Ar. / Rancho Grande / 1100m 16-23.X.66.

Taxonomy

Spongiphoridae, Spongiphorinae, Purex sinuatus Brindle, 1971.

Remark

Purex trituberculatus Brindle, 1971 was synonym ised with P. sinuatus Brindle, 1971 by Brindle (1977b:

119).

trituberculatus Brindle, 1973 ( Spongovostox )

Spongovostox trituberculatus Brindle, 1973f: 186, figs 249, 256.

Type material

1 S, paratype, F3233.1612, dried, mounted on card.

Locality label

Coll. Mus. Congo / Kivu : Sanghe, PI. De / Ruzizi (A la lumiere) / XII-1951 / H. Bomans.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox trituberculatus Brindle, 1973.

truncata Brindle, 1978 (Euborellia)

Euborellia truncata Brindle, 1978: 175, figs 291, 322.

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MILES C., Dermaptera of the Manchester Museum

Type material

1 <$, paratype, F3233.1442; 1 §, paratype, F3233.1443; dried, mounted on pin.

Locality labels

F3233.1442-1443: Uganda. / Entebbe. / v - vi 1912 / Mrs. Gowdey.

Taxonomy

Anisolabididae, Anisolabidinae, Euborellia truncata Brindle, 1978.

tuberculata Hincks, 1958 ( Isolaboides )

Isolaboides tuberculata Hincks, 1958b: 133, figs 1-2.

Type material

1 S, paratype, F3233.1484, dried, mounted on card.

Locality label

Liban: Les Cedres / Coiffait. 28.IX. / Museum Paris.

Taxonomy

Spongiphoridae, Isolaboidinae, Isolaboides tuberculata Hincks, 1958.

Remark

Index card entry: Lebanon.

tuberosa Brindle, 1970 (Sphingolabis)

Sphingolabis tuberosa Brindle, 1970f: 678, figs 47—49.

Type material

1 S, paratype, F3233.695, dried, mounted on card.

Locality label

Solomon Is. / Guadacanal [sic] / Popamanasiu / 25. X. 1965 / P. Greenslade.

Taxonomy

Spongiphoridae, Labiinae, Sphingolabis tuberosa Brindle, 1970.

umbrosa Brindle, 1964 ( Gelotolabis )

Fig. 20

Gelotolabis Brindle, 1964: 752, figs 5, 13.

Type material

1 (?, holotype, F3233.1420 (Fig. 20); 1 $, paratype, F3233.1421; dried, mounted on pin.

Type locality

Tanganyika / Nachingwea / Southern Prov. / IX. 53 - III. 54 / V. F. Eastop.

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European Journal of Taxonomy 141: 1-138 (2015)

Locality label

F3233.1421: As type locality.

Taxonomy

Anisolabididae, Anisolabidinae, Gonolabis umbrosa (Brindle, 1964).

Remark

Transferred by Steinmann (1978: 195).

uncinata Brindle, 1978 (Anisolabis)

Anisolabis uncinata Brindle, 1978: 88, figs 110-112, 236.

Type material

1 S, paratype, F3233.1422, dried, mounted on card.

Locality label

Biot. No 15 / marais boises / avec Pandanus. I.R.S.A.C. - Mus. Congo / Kivu : Mwenga / 1250 m. 24-

IV-1958/B.15 N. Leleup.

Taxonomy

Anisolabididae, Anisolabidinae, Flexiolabis uncinata (Brindle, 1978).

Remark

Transferred by Steinmann (1989a: 35).

unicolor Brindle, 1975 ( Brachylabis )

Brachylabis unicolor Brindle, 1975b: 684, figs 1-2.

Type material

1 c?, paratype, F3233.1481; 1 5, paratype, F3233.1482; dried, mounted on card.

Locality labels

F3233.1481: Tanzanie: Mts Uluguru / Kimboza for. heliophile / alt. 600m 24-30/VII/71 // Coll. Mus.

Tervuran / Mission Mts Uluguru / L. Berger, N. Leleup / J. Debecker V/VIII/71.

F3233.1482: Tanzanie: Mts Uluguru / Chenzema, alt. 1700m. / 2-22/VII/71 // Coll. Mus. Tervuran /

Mission Mts Uluguru / L. Berger, N. Leleup / J. Debecker V/VIII/71.

Taxonomy

Anisolabididae, Isolabidinae, Isolabis unicolor (Brindle, 1975).

Remarks

1. Transferred by Brindle (1978: 44).

2. The original description states that all paratypes have the locality Kimboza, as for F3233.1481. The

locality data on the label of the female (F3233.1482) corresponds exactly to the allotype data in the

description.

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MILES C., Dermaptera of the Manchester Museum

unicolor Brindle, 1966 ( Emboros )

Emboros unicolor Brindle, 1966g: 255, figs 62, 65, 69.

Type material

4 $ J, paratypes, F3233.1688-1691, dried, mounted on card.

Locality labels

F3233.1688-1691: Madagascar-Est / dct. Sambava /R. N. XII / Marojejy-Ouest 1140m / XI-59 R Soga.

Taxonomy

Forficulidae, Skendylinae, Cosmiella unicolor (Brindle, 1966).

Remark

Transferred by Steinmann (1989c: 675).

vansomereni Brindle, 1969 ( Chaetospania )

Chaetospania vansomereni Brindle, 1969a: 98, figs 1-2.

Type material

2 S3, paratypes, F3233.1526-1527; 2 paratypes, F3233.1528-1529; dried, mounted on card.

Locality labels

F3233.1526-1529: Van Someren / Meru, 7/43.

Taxonomy

Spongiphoridae, Sparattinae, Chaetospaniini, Chaetospania vansomereni Brindle, 1969.

Remark

Index card entry: Tanzania.

variabilis Brindle, 1973 ( Mixocosmia )

Mixocosmia variabilis Brindle, 1973b: 50, figs 1-3.

Type material

12 33 , paratypes, F3233.1758-1769; 8 §§, paratypes, F3233.1770-1777; dried, mounted on card

except: F3233.1759, 1762-1763, 1766, 1769-1770, 1772-1773; dried, mounted on pin.

Locality labels

F3233.1762: Venezuela - Aragua / Rancho Grande / 1100m. 10-VIII-69 // J. A. Clavijo / col.

Same data except date: F3233.1765,11 Aug. 1969; F3233.1759,20 Aug. 1969; F3233.1758,F3233.1773,

27 Aug. 1969.

Same data except date and collector: F3233.1770, 30-VII-69 // J. Salcedo / J. A. Clavijo; F3233.1766,

5-IX-69// J. Salcedo/J. A. Clavijo; F3233.1768, 16.V.70//F. Fernandez Y. / J. A. Clavijo; F3233.1772,

14-VIII-49 // F. Fernandez Y / - P Fenjues; F3233.1763, 20.VIII.69 // F. Fernandez Y. / C. J. Rosales

/ Cols; F3233.1771, 7.XI.55 // F. Fernandez Y. / C. J. Rosales / Cols; F3233.1761: [9?].VI.55 // F.

Fernandez Y. / C. J. Rosales / Cols; F3233.1760, 4-IV-51 / C. J. Rosales; F3233.1767, F3233.1769,

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European Journal of Taxonomy 141: 1-138 (2015)

F3233.1774-1777, 17.XI.1968 /H. Reichardt col; F3233.1764, as F3233.1768 except 21.V.71 // En la

/ Luz.

Taxonomy

Forficulidae, Skendylinae, Mixocosmia variabilis Brindle, 1973.

Remarks

1. Steinmann (1989c) and Hopkins et al. (2014) have Brindle’s original name as Kleter variabilis and

so have the current name as Mixocosmia variabilis (Brindle, 1973); however, Brindle’s original

description names the new species Mixocosmia variabilis , not Kleter.

2. The number of paratypes does not match that stated in the original description, i.e., of 36 male and

62 female paratypes with the same data as the holotype (dates ranging from 30 Apr. 1969 to 26 May

1971) mentioned in the description, 11 male and 14 female paratypes were at MMUE, not 12 males

and 8 females.

variabilis egoloensis Brindle, 1970 ( Auchenomus )

Auchenomus variabilis egoloensis Brindle, 1970f: 667, figs 23-24.

Type material

1 S, paratype, F3233.559; 1 §, paratype, F3233.560; dried, mounted on card.

Locality labels

F3233.559-560: Solomon Is. / New Georgia Group / nr. Egolo 1-25 m. / 16-VII-1959.

Taxonomy

Spongiphoridae, Sparattinae, Auchenomini, Auchenomus variabilis egoloensis Brindle, 1970.

variabilis guadalcanalensis Brindle, 1970 (Auchenomus)

Fig. 49

Auchenomus variabilis guadalcanalensis Brindle, 1970f: 668, figs 25-26.

Type material

1 c?, holotype, F3233.557 (Fig. 49); 1 $, allotype, F3233.558; dried, mounted on card.

Type locality

Solomon Is. / Guadalcanal / Rua Vata / 21:xi: 1954 / E S Brown.

Locality label

F3233.558: As type locality.

Taxonomy

Spongiphoridae, Sparattinae, Auchenomini, Auchenomus variabilis guadalcanalensis Brindle, 1970.

variegatus Hincks, 1957 ( Blandex )

Blandex variegatus Hincks, 1957b: 45, figs 7-10.

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MILES C., Dermaptera of the Manchester Museum

Type material

1 S, paratype F3233.1338, dried, mounted on card.

Locality label

S. Afr. Cape Prov. / Cape Point / Nature Reserv / 24.X.50 No. 15. Swedish South Africa / Expedition /

1950-1951 / Brinck-Rudebeck.

Taxonomy

Pygidicranidae, Blandicinae, Blandex variegatus Hincks, 1957.

venezuelica Brindle, 1977 ( Marava )

Marava venezuelica Brindle, 1977b: 123, figs 25-26.

Type material

1 (?, paratype, F3233.1556; 1 $, paratype, F3233.1557; dried, mounted on card.

Locality labels

F3233.1556: Venezuela Aragua / Rancho Grande / 1100 m. 16.VIII.74.

F3233.1557: Same data except date 4 Aug. 1974.

Taxonomy

Spongiphoridae, Spongiphorinae, Marava venezuelica Brindle, 1977.

Remark

The original description gives the date for the male paratype (F3233.1556) as 2 Aug. 1974, not 16 Aug.

1974.

venezuelica Brindle, 1982 ( Yepezia )

Yepezia venezuelica Brindle, 1982a: 38, figs 7—11.

Type material

1 S, paratype, F3233.1636, dried, mounted on card, without head.

Locality label

Guigiie / Venezuela - Cara- / bobo m. / 23.11.1968. // J. & B. Bechyne / leg.

Taxonomy

Spongiphoridae, Geracinae, Yepezia venezuelica Brindle, 1982.

venezuelicum Brindle, 1974 (Barygerax)

Barygerax venezuelicum Brindle, 1974b: 114, figs 5-8.

Type material

2 S3, paratypes, F3233.1502-1503; 2 paratypes, F3233.1504-1505; dried, mounted on card.

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Locality labels

F3233.1502: Venezuela - Aragua / Rancho Grande / 1100 m. 25-VII-69. // J. Salcedo / col. // Venezuela

Inst. / Zool. Agricola - / Fac. Agronomia / Univ. Central.

Same data except date: F3233.1503, 30 Jul. 1969; F3233.1504, 10 Aug. 1969; F3233.1505, 25 Mar.

1971.

Taxonomy

Spongiphoridae, Geracinae, Barygerax venezuelicum Brindle, 1974.

Remarks

F The original description has the date 30 Aug. 1969 for the specimen corresponding to F3233.1503,

not 30 Jul. 1969.

2. There are two male and two female paratypes at MMUE, not three males and one female as in the

description.

venezuelicus Brindle, 1973 (Kleter)

Kleter venezuelicus Brindle, 1973b: 49, figs 5-6.

Type material

1 <$, paratype, F3233.1736; 2 5?, paratypes, F3233.1737-1738; dried, mounted on card.

Locality labels

F3233.1736: Venezuela - Ara- / gua- El Limon. / 450m. 22-1-70. // En Pseudo tallos de camburusados /

como trampa. // col. P. Bolivar.

F3233.1737: Venezuela - Ara- / gua-Maracay / 450m. 19-1-70. //En tramp / de cambur. // col. P. Bolivar.

F3233.1738: Venezuela - Cara- / bobo - Mariara / 30-1-1970. // Trampa / de Luz.

Taxonomy

Forficulidae, Skendylinae, Hypocosmiella venezuelica (Brindle, 1973).

Remarks

Transferred by Steinmann (1993: 184). Not listed in Hopkins et al. (2014).

vittipennis Hincks, 1955 ( Cranopygia )

Cranopygia vittipennis Hincks , 1955c: 820, figs 13-14.

Type material

1 c?, paratype, F3233.1317, dried, mounted on card.

Locality label

S. India: S. Coorg / Ammatti 3100 ft. / 3.XI.52 under bark / PS. Nathan.

Taxonomy

Pygidicranidae, Pygidicraninae, Acrania vittipennis (Hincks, 1955).

120

MILES C., Dermaptera of the Manchester Museum

Remarks

Cranopygia vittipennis Hincks was transferred to Epicranopygia by Steinmann (1986b: 271). The

species was transferred to Acrania Burr by Engel & Haas (2007: 19).

weidneri Brindle, 1966 ( Esphalmenus )

Esphalmenus weidneri Brindle, 1966f: 128, figs 9, 11-12.

Type material

1 5, paratype, F3233.1365, dried, mounted on card.

Locality label

Probably near / Santiago / Chile / Dr. G. Rahm.

Taxonomy

Pygidicranidae, Esphalmeninae, Esphalmenus weidneri Brindle, 1966.

wittei Hincks, 1955 (Diaperasticus)

Diaperasticus wittei Hincks, 1955b: 12, figs 4-5.

Type material

1 c?, paratype, F3233.1713; 1 §, paratype, F3233.1714; dried, mounted on card.

Locality labels

F3233.1713: Congo beige: P. N. U. / Lusinga (1760 m.) / 18-vii 1947 / Mis. G. F. De Witte. 604a.

F3233.1714: Congo beige : P. N. U. / Kalule N. r. g. Face Mujinga- / Kalenge (1050m.) 28-II-3-III-49

/Mis. G. F. De Witte. 2399a.

Taxonomy

Forficulidae, Diaperasticinae, Diaperasticus wittei Hincks, 1955.

wittmeri Brindle, 1975 (Irdex)

Irdex wittmeri Brindle, 1975c: 28, figs 24-25.

Type material

2 <$<$, paratypes, F3233.1536-1537; 2 $ paratypes, F3233.1538-1539; dried, mounted on card.

Locality labels

F3233.1536: Wangdi Phodrang / 1300m Darju La. // Nat.-Hist. Museum / Basel - Bhutan / Expedition

1972.

F3233.1537-1538: Km 87 von / Phuntsholing 22/5 / 1680m. Nat.-Hist. Museum / Basel - Bhutan /

Expedition 1972.

F3233.1539: 21 km O Wangdi / Phodr. 1700-2000. Nat.-Hist. Museum / Basel - Bhutan / Expedition

1972.

121

European Journal of Taxonomy 141: 1-138 (2015)

Taxonomy

Spongiphoridae, Spongiphorinae, Irdex wittmeri Brindle, 1975.

woldai Brindle, 1988 ( Mixocosmia )

Mixocosmia woldai Brindle, 1988: 55, figs 25-27.

Type material

1 S, paratype, F3233.1757; 2 5?, paratypes, F3233.1755-1756; dried, mounted on card.

Locality labels

F3233.1755-1756: Panama, Fortuna / Wet forest / at light, 1050m / 29.6.1979 / Dr. Henk Wolda, STRI.

F3233.1757: Same data except date 25 Sep. 1978.

Taxonomy

Forficulidae, Skendylinae, Mixocosmia woldai Brindle, 1988.

wuermlii Brindle, 1975 ( Spongovostox )

Spongovostox wuermlii Brindle, 1975c: 33, figs 33-34.

Type material

1 S, paratype, F3233.1613, dried, mounted on card.

Locality label

km 87 von 1680 m / Phuntsholing 22/5. // Basel-Bhutan / Expedition 1972.

Taxonomy

Spongiphoridae, Spongiphorinae, Spongovostox wuermlii Brindle, 1975.

yepezi Brindle, 1974 ( Cylindrogaster )

Cylindrogaster yepezi Brindle, 1974b: 108, figs 1-2.

Type material

1 S, paratype, F3233.1312, dried, mounted on pin.

Locality label

Venezuela - Aragua / Rancho Grande / 1100m 27.III.67 / C. J. Rosales J. Salcedo.

Taxonomy

Pygidicranidae, Cylindrogastrinae, Cylindrogaster yepezi Brindle, 1974.

122

MILES C., Dermaptera of the Manchester Museum

Other Material: Specimens carrying a type label , but for which no description has

been found

These taxa are not listed in Steinmann (1989c) or Hopkins et al. (2014) and the present author has been

unable to locate any published descriptions. Thus, these species names seem to represent unpublished

‘manuscript names’ only.

flavolinecita Brindle ( Tagalina ) MS

Figs 62-63

Type material

1 S, paratype, F3233.1864 (Fig. 62), dried, mounted on pin.

Locality label

Museum Feiden / Nieuw Guinea Exp K.N.A.G. 1939 / Araboebirak ll-x-1939.

Determination label

Manchester Museum Paratype // Tagalina $ / flavolineata Brindle / det. A. Brindle 1980 / paratype.

Taxonomy

Pygidicranidae, Pygidicranidae, Tagalina.

Remarks

Found recently in a box separate from the collection. A note written by A. Brindle pinned next to the

specimen reads “ Tagalina flavolineata n. sp. (paratype for Manchester) (holotype in Feiden). Not yet

described (paper not written)” (Fig. 63).

The holotype and allotype have been confirmed to be at the Naturalis Biodiversity Center, Feiden (F.

Willemse, pers. comm.), labelled Tagalina flavolineata and bearing Brindle’s determination label.

rossi Hincks ( Carcinophora ) MS

Fig. 61

Type material

1 S, paratype, F3233.1848 (Fig. 61), dried, mounted on pin.

Locality label

PERU: 43 mi. E. / Tingo Maria / 1200 m. XI. 18.54. // E. I. Sehlinger / & E. S. Ross / collectors.

Determination label

Carcinophora / rossi Hincks / TYPE S / det. W. D. Hincks // MS name.

Taxonomy

Anisolabididae, Anisolabidinae, Carcinophora.

spatulus Brindle ( Nesogaster)

Fig.59

Type material

1 5, paratype, F3233.548 (Fig. 59), dried, mounted on card

Locality label

Fiji, Vanua Fevu / Nandrau / 9.10.1977 / G. Kuschel.

123

European Journal of Taxonomy 141: 1-138 (2015)

Determination label

Nesogaster §, / spatulus Brindle / det. A. Brindle 1983 / paratype.

Taxonomy

Spongiphoridae, Nesogastrinae, Nesogaster.

Remarks

This species is fully described in a typewritten manuscript in the Brindle archive, item 65 (Brindle &

Maddison: date not known [unpublished?]), as W. spatulus sp. n. NZAC Endemic’ [NZAC = New

Zealand Arthropod Collection], The label data matches the described material exactly.

subcarinata Brindle ( Metalabis ) MS

Fig. 60

Type material

1 c?, holotype, F3233.1834 (Fig. 60), dried, mounted on card.

Type locality

Brokopondo / Surinam / 20.IX. 1963.

Taxonomy

Anisolabididae, Anisolabidinae, Metalabis.

T O', Uruv

4. k>

c, ff

4 hal u S £wv*cik_

{ det. A. Brindle

pofCcJypc-

F3233.548

po odo

" $\jv \ no_m

3a f*.

t Cl. lCt hi f>

do b£<uA ooJTa,

* BrJ .

del, A, firing l¥t7

F3233.1834

PEBU:43

TioffO

120!) m,XM3-S4

H, I. Seh linger

collector*

d <X iTC r Wm£>

tv pie ><

det. W.D Hmcki

Mnsrmm Trfft’en

Hlfiuw Guinea, Kjcp

k , ,x. m g . im*

Aral) oebi vak

-tfoi/b | (pMta ^6*

det* At Brindle 19®&

F3233.1864

-«e

k AlccVLit nctTcX ft WO,

6 pe?Tc.\ HcAtKtf-fer;)

£ K

ts It-

Ki^T y^r

jT p o e$£ir n o r i^,n fte. n ^

Figs 59-63. Other material, presumed manuscript names, in the collection of the Manchester Museum.

59. Nesogaster spatulus Brindle (det. 1983). 60. Metalabis subcarinata Brindle (det. 1987). 61. Carcino-

phora rossi Hincks (no date). 62. Tagalina flavolineata Brindle (det. 1980). 63. Brindle’s handwritten

label for the Tagalina flavolineata specimen (Fig. 62). Scale bars = 1 cm.

124

MILES C., Dermaptera of the Manchester Museum

Acknowledgements

I wish to thank Dmitri Logunov (Manchester Museum, UK) for the opportunity to work on this collection,

his advice, proofreading, and processing the holotype photographs; Philip Rispin (Manchester Museum,

UK) for the use of his equipment, time and skill in photographing the specimens; Leonid Anisyutkin

(Zoological Institute RAS, St. Petersburg, Russia) for advice on the taxonomy of the type material;

Mary Angela Nellie Jayapaul for information relating to the life of PS. Nathan; Luc Willemse (Naturalis

Biodiversity Center, Leiden, Netherlands) for information relating to Brindle MS species names; and

the two anonymous referees and desk editor Danny Eibye-Jacobsen (EJT) for their detailed and helpful

co mm ents

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zoologique beige aux lies Galapagos et en Ecuador 1: 171-176.

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1. Pericominae, Strongylopsalinae, Sparattinae. Journal of Natural History 2: 273-303. http://dx.doi.

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Decelle, 1961-1964). Annales duMusee royal de VAfrique Centrale, Zoologie (8) 165: 11-20.

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Brindle A. 1968f. Notes on the genus Timomenus Burr (Dermaptera, Forficulidae). Entomologist’s

Monthly Magazine 104: 243-249.

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and other Guyanas 10 (36): 1-60.

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Labiidae). Proceedings of the Royal Entomological Society of London B 38 (7-8): 95-100. http://dx.doi.

org/10,1111/i. 1365-3113.1969.tb00237.x

126

MILES C., Dermaptera of the Manchester Museum

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Entomologist 102: 197-200.

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Entomological. 145-150.

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190-193.

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Monthly Magazine 105: 273-277.

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Forficulidae). Entomologist’s Monthly Magazine 106: 97-105.

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Entomologist 103: 217-228.

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de Zoologie et de Botanique Africaines 81 (3—4): 255-270.

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Brindle A. 1970g. The Hemimerina and Dermaptera of Angola. Publicagoes Culturais da Companhia

de Diamantes de Angola 34: 39-74.

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Regions. IT Geracinae and Labiinae. Journal of Natural History 5: 155-182. http://dx.doi.

org/10.1080/0022293 7100770101

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Frangais d’Afrique Noire 86: 265-281.

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MILES C., Dermaptera of the Manchester Museum

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Brindle A. 1982b. Three new species of African Diplatys Serville (Dermaptera, Diplatyidae).

Entomologist s Monthly Magazine 118:163-166.

Brindle A. 1984. The Esphalmeninae (Dermaptera: Pygidicranidae): a group of Andean and Southern

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tb00053.x

Brindle A. 1985. Synonymical notes on the American species of Labia Leach (Dermaptera, Labiidae)

with a key to species and description of a new species. Entomologists Monthly Magazine 121: 73-80.

Brindle A. 1987. New Dermaptera records from Nepal, with descriptions of new species and a review of

the Himalayan Fauna (Insecta). Courier Forschungsinstitut Senckenberg 93: 333-351.

Brindle A. 1988. Dermaptera at light in Panama. Entomologists Monthly Magazine 124: 45-56.

Brindle A. 1989. Carcinophora americana (Beauvois) and related species (Derm., Anisolabididae).

Entomologist s Monthly Magazine 125: 13-16.

Brindle A. & Maddison P. [Date not known] The Dermaptera of Fiji, with records from other SW Pacific

Islands [unpublished?]. Brindle Archive item 65, Manchester Museum.

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Manuscript received: 29 April 2015

Manuscript accepted: 13 July 2015

Published on: 1 October 2015

Topic editor: Koen Martens

Desk editor: Danny Eibye-Jacobsen

Printed versions of all papers are also deposited in the libraries of the institutes that are members of

the EJT consortium: Museum National d’Histoire Naturelle, Paris, France; Botanic Garden Meise,

Belgium; Royal Museum for Central Africa, Tervuren, Belgium; Natural History Museum, Fondon,

United Kingdom; Royal Belgian Institute of Natural Sciences, Brussels, Belgium; Natural History

Museum of Denmark, Copenhagen, De nm ark

133

European Journal of Taxonomy 141: 1-138 (2015)

Appendix I: Correspondence in the Hincks Archive relating to

Dermaptera, the Manchester Museum

Ordered alphabetically by correspondent, then date.

Item No.

From

Date

No. of pages

387

G. Bey-Bienko

24 Jan. 1937

postcard

G. Bey-Bienko

17 Mar. 1937

Alfredo Borelli, Torino

16 Feb. 1933

M. Burr, Maidenhead

24 Jan. 1933

M. Burr, Maidenhead

02 Feb. 1933

M. Burr, Maidenhead

09 Feb. 1933

M. Burr, Maidenhead

22 Feb. 1933

M. Burr, Maidenhead

21 Jan. 1934

M. Burr, Maidenhead

24 Apr. 1934

M. Burr, Maidenhead

23 Oct. 1934

M. Burr, Maidenhead

25 Nov. 1934

M. Burr, Windsor

23 Mar. 1935

M. Burr

30 Mar. 1935

M. Burr, Windsor

13 May 1935

M. Burr, Windsor

02 Oct. 1935

M. Burr, Windsor

07 Oct. 1935

M. Burr, Windsor

31 Oct. 1935

M. Burr, enclosing letter from O. Piel, Shanghai

10 Nov. 1935

2,1

M. Burr, Windsor

12 Nov. 1935

M. Burr, Maidenhead

30 Jan. 1936

M. Burr, Windsor

14 Feb. 1936

M. Burr, Windsor

07 Mar. 1936

M. Burr, Windsor

26 Sep. 1936

M. Burr, Windsor

17 Dec. 1936

M. Burr, Windsor

06 Jan. 1937

M. Burr, Windsor

11 Mar. 1937

1+3 photos

M. Burr, Worthing

13 Jun. 1938

103

M. Burr, Worthing

14 Jul. 1938

109

M. Burr, Worthing

28 Aug. 1938

110

M. Burr, Worthing

15 Nov. 1938

120

M. Burr to N.D. Riley, Natural History Museum

15 Nov. 1938

111

M. Burr, Worthing

17 Nov. 1938

M. Burr, Windsor

04 May 1939

M. Burr, Istanbul

19 Dec. 1948

M. Burr, Istanbul

10 Sep. 1950

M. Burr, Istanbul,

W.D. Hincks

08 Oct. 1951,

12 Dec. 1951

1,1

134

MILES C., Dermaptera of the Manchester Museum

Appendix I, continued

Item No.

From

Date

No. of pages

389

Unknown person [M. Burr?]

No date

postcard

390

Fritz Carpentier

15 Sep. 1936

postcard

276

Lucien Chopard, Museum National d’Histoire Naturelle

Entomologie, Paris

10 Feb. 1960

275

W.D. Hincks to L. Chopard

07 Mar. 1960

274

Lucien Chopard, Museum National d‘Histoire Naturelle

Entomologie, Paris

15 Mar. 1960

271

Lucien Chopard, Museum National d‘Histoire Naturelle

Entomologie, Paris

30 Nov. 1960

272

W.D. Hincks to L. Chopard

05 Dec. 1960

268

Lucien Chopard, Museum National d‘Histoire Naturelle

Entomologie, Paris

16 Jan. 1961

269

W.D. Hincks to L. Chopard

25 Jan. 1961

Ashley B. Gurney, Divn. of Insect Identification, US Dept of

Agriculture, Washington

03 May 1950

Ashley B. Gurney, Divn. of Insect Identification, US Dept of

Agriculture, Washington

27 Mar. 1951

125

Ashley B. Gurney, Divn. of Insect Identification, US Dept of

Agriculture, Washington

06 Sep. 1960

127

[Probably Ashley B. Gurney, Divn. of Insect Identification,

US Dept of Agriculture, Washington]

06 Sep. 1960

128

[Probably W.D. Hincks to Ashley B. Gurney]

29 Sep. 1960

Kenneth J. Hayward, Director, Instituto de Entomologia,

Tucuman, Argentina

29 Nov. 1949

247

W.D. Hincks to Colin Johnson

31 Aug. 1960

239

W.D. Hincks to PV. Joshi

20 Oct. 1960

237

P. V. Joshi, Dept, of Zoology, N. Wadia College, Poona,

India

31 Oct. 1960

283

Dr. A.P Kapur, Officer in Charge, Entomology Seen.,

Zoological Survey of India, Calcutta

26 Nov. 1959

280

Dr. A.P. Kapur, Officer in Charge, Entomology Seen.,

Zoological Survey of India, Calcutta

21 Mar. 1960

281

W.D. Hincks to Dr. A.P. Kapur, Officer in Charge,

Entomology Seen., Zoological Survey of India, Calcutta

24 Mar. 1960

279

W.D. Hincks to Dr. A.P. Kapur, Officer in Charge,

Entomology Seen., Zoological Survey of India, Calcutta

28 Mar. 1960

278

Dr. A.P. Kapur, Officer in Charge, Entomology Seen.,

Zoological Survey of India, Calcutta

15 Apr. 1960

277

W.D. Hincks to Dr. A.P. Kapur, Officer in Charge,

Entomology Seen., Zoological Survey of India, Calcutta

19 Jul. 1960

310

Dr. H.W. Koepcke, Peru

17 Nov. 1959

309

W.D. Hincks to Dr. H.W. Koepcke, Peru

23 Nov. 1959

359

A. Loveridge, M/V Durban Castle , in the Thames

15 Jun. 1960

360

W.D. Hincks to A. Loveridge

23 Jun. 1960

135

European Journal of Taxonomy 141: 1-138 (2015)

Appendix I, continued

Item No.

From

Date

No. of pages

317

A. Loveridge

01 Dec. 1960

318

W.D. Hinclcs to A. Loveridge

05 Jan. 1961

H.K. Long, Shell Petroleum Company of Colombia, Bogota

26 Mar. 1939

H.K. Long, Shell Petroleum Company of Colombia, Bogota

04 Jul. 1939

106

H. Lucht, Bondowoso, Java

17 Jun. 1938

H. Lucht, Bondowoso, Java

17 Aug. 1938

H. Lucht, Bondowoso, Java

24 May 1939

Raymond Mamet, Mauritius

09 Nov. 1949

T. McCarthy, Chief Entomologist N.S.W. Dept, of

Agriculture

03 Apr.

C. Menozzi

25 Jun. 1936

postcard

E. Morales Agacino, Madrid

11 Jan. 1936

E. Morales Agacino, Madrid

27 Jan. 1936

postcard

Edward P Mumford, Dept of Entomology, Oxford

24 Sep. 1937

Edward P Mumford, Dept of Entomology, Oxford

24 Sep. 1937

102

Ferd. Nevermann, San Jose, Costa Rica

11 Apr. 1938

R.PO. Piel, Director, Musee Heude, Shanghai

28 Feb. 1937

R.PO. Piel, Director, Musee Heude, Shanghai

03 Jan. 1936

R.PO. Piel, Director, Musee Heude, Shanghai

30 Nov. 1936

240

David R. Ragge

12 Oct. 1960

James A. Rehn, Acting Curator, Academy of Natural

Sciences of Philadelphia

03 Mar. 1933

James A. Rehn, Acting Curator, Academy of Natural

Sciences of Philadelphia

04 Apr. 1933

Isabel Sanabria

18 May 1959

124

F.H. Schade, Villarrica, Paraguay

25 Feb. 1939

104

F.H. Schade, Villarrica, Paraguay

19 Jun. 1938

Dr H. Schouteden, Director Musee du Congo Beige,

Tervueren, Belgium

02 Feb. 1935

Dr H. Schouteden, Director Musee du Congo Beige,

Tervueren, Belgium.

22 Feb. 1935

V. Van Straelen, President, Institut des Parcs nationaux du

Congo Beige, Brussels

22 Jul. 1937

105

M.E. Walsh, Soekaboemi, Java

17 Jun. 1938

M.E. Walsh, Soekaboemi, Java

16 Aug. 1938

Dr. Friedrich Zacher, Berlin

20 Nov. 1934

Unknown person, Musee du Congo Beige, Tervuren,

Belgium

20 Jan. 1935

386

Unknown person

22 Mar. 1938

postcard

375

Unknown person [Letterhead ‘F. H. Walz, Buenos Aires’]

22 Jun. 1961

136

MILES C., Dermaptera of the Manchester Museum

Appendix II: Correspondence in the Brindle Archive relating to

Dermaptera, the Manchester Museum

Ordered alphabetically by correspondent, then date.

Item No.

From

Date

No. of pages

147

Annette Aiello regarding publication of ‘Insects of Panama

and Mesoamerica: Selected Studies’

12 May 1992

148

A. Brindle to A. Aiello

27 May 1992

M. Beier

15 Nov. 1963

postcard

242

Dr. T. Cekalovic (in Spanish) regarding supply of Dermaptera

from Chile, Easter Islands, Juan Fernandez Island, Magellan

region and Tierra del Fuego

08 Nov. 1974

243

Handwritten translation of 242

247

Dr. T. Cekalovic, sending 1 box Dermaptera and Isoptera, 2

boxes Lepidoptera

20 Feb. 1975

248

Handwritten translation of 243

249

Dr. T. Cekalovic

03 Apr. 1975

259

A. Brindle to Dr. T. Cekalovic, with certificate of posting for

return of insects

15 Oct. 1975

Karl Jordan to W.D. Hincks

30 Jan. 1947

154

Dr. H.W. Koepcke, Peru to W.D. Hincks regarding progress

on identification

12 Mar. 1964

155

Dr. H.W. Koepcke’s reply to 154

22 Jul. 1964

157

Dr. H.W. Koepcke regarding Brindle’s naming of Idolopsalis

koepcke i

10 Nov. 1965

158

A. Brindle to Dr. H.W. Koepcke enclosing paper reprints

relating to Dermaptera sent for identification plus list of

Peruvian Dermaptera

05 Jul. 1967

218

PS. Nathan, sending 10 boxes of‘desiderata’

21 Jul. 1965

219

A. Brindle’s to PS. Nathan, reply to 218

08 Sep. 1965

220

PS. Nathan, enquiring what families Brindle would like

24 Nov. 1965

221

PS. Nathan

22 Aug. 1966

223

A. Brindle to PS. Nathan

08 Jul. 1967

222

PS. Nathan

23 Nov. 1967

224

PS. Nathan, sending 3 boxes of Dermapera and beetles

13 Feb. 1968

225

Attached to 226. Notification of retirement of PS. Nathan,

handing over of business to his daughter-in-law

226

See 225. Mrs T.R.S. Nathan, sending 1 box of Dermaptera

and receipt of reprint of earwig named for PS. Nathan

02 Jul. 1969

227

Mrs T.R.S. Nathan

15 Apr. 1970

228

A. Brindle to Mrs T.R.S. Nathan, requesting more

Dermaptera, Ptiliidae, Neuroptera or Orthoptera

22 Apr. 1970

229

Mrs T.R.S. Nathan

31 Mar. 1971

230

A. Brindle to Mrs T.R.S. Nathan, requesting further

Dermaptera, Neuroptera, Orthoptera, Hemiptera and

Hymenoptera

14 Apr. 1971

137

European Journal of Taxonomy 141: 1-138 (2015)

Appendix II, continued

Item No.

From

Date

No. of pages

231

Mrs T.R.S. Nathan, sending 3 boxes Dermapera and

Orthoptera

12 Jul. 1971

232

Mrs T.R.S. Nathan

27 Apr. 1972

233

Mrs T.RS. Nathan, sending 4 double size boxes of

Dermaptera, Orthoptera, Hemiptera and Hymenoptera

13 Jul. 1972

234

A. Brindle to Mrs T.R.S. Nathan

18 Sep. 1972

R Oromi with some additions and amendments to Brindle’s

30 Oct. 1990,

08 Nov. 1990

article on cavemicole Dermaptera (No. 63) and Brindle’s

Edward [Popham?]

17 Aug. 1981

235

A. Brindle to S.K. Sircar, Assam, regarding supply of

Dermaptera and Ptiliidae

29 Nov. 1967

236

S.K. Sircar, Assam, reply to 235

09 Dec. 1967

237

A. Brindle to S.K. Sircar

05 Jan. 1968

238

S.K. Sircar, sending 700 earwigs

27 Feb. 1968

239

A. Brindle to S.K. Sircar

04 Mar. 1968

174

A. Brindle to Dr R. zur Strassen regarding return of

Cassidinae unidentified

24 Jan. 1973

175

Dr. R. zur Strassen’s reply to 174

05 Feb. 1973

272

A. Brindle to Taiwan Novelty Company, enquiring about

supply of Dermaptera from Formosa

29 Nov. 1967

273

Au Chun-Cheng, Director, Taiwan Novelty Company, reply to

272

07 Dec. 1967

262

J.S. Taylor, Natal, South Africa

16 Feb. 1966

263

J.S. Taylor, Natal, South Africa

15 Mar. 1966

264

J.S. Taylor, Natal, South Africa, sending vials of insects

including forficulids and staphylinids

04 Apr. 1966

274

A. Brindle to H. Winkler, Vienna, requesting prices for

Coleoptera, Dermaptera and other small orders

21 Nov. 1968

138