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j
JOURNAL
OF THE
ASIATIC SOCIETY OF BENGAL.
VOL. LXV.
PART II. (NATURAL HIsTory, &c.).
(Nos. I & II.—1898.)
EDITED BY THE
NaturaL jiistTo RY pECRETARY,
VS
“Tt will flourish, if naturalists, chemists, antiquaries, philologers, and men of
science in different parts of Asza, will commit their observations to writing, and send
them to the Asiatic Society at Calcutta. It will languish, if such communications shall
be long intermitted ; and it will die away, if they shall entirely cease.”” SIR WM. JONES.
CALCUTTA:
PRINTED AT THE BAPTIST MISSION PRUSS,
AND PUBLISHED BY THE
ASIATIC SOCIETY, 57, PARK SIREET.
1898.
ane
sh i) Am) a
a 4
LIST OF CONTRIBUTORS.
Aucock, ALFRED ;— Materials for a Carcinological Fauna of India,
No. 8. The Brachyura OS eae Part t.* The ane
POOPED 60 s000.000008
Buapouri, C. B., and Based .) re — : Comeatienen ee iis
Chemical Laboratory, Presidency Saileae: Calcutta. On Double
Thiosulphates of Copper and Sodium..
Finn, F.;—Note on the Long-Snouted Whe: ‘caps: (Dryophis
mycterizans)...
;— Note on the Seaannat ahaa of Spinasdeyd an ee ais
of the Purple Honeysucker (Arachnechthra asiatica) and of
an analogous American bird (Coereba cyanea)... ....seceecee severe
Kine, Grorce ;— Materials for a Flora of the Malayan Peninsula..
——-—————- ; and Pratin, D. ;— Descriptions of some new aah
from the North-Hastern Frontiers of India ..
Nic&vitte, LioneL DE ;— On a small collection of Batherflies meas
Buru in the Moluccas ......... ee
——-————-; and Gn, erate “ae ea List
of the Butterflies of the Ké Isles. (Plate is )
Riptey, H. N.;—New species of Hntada from Singapore bong
ate fls' Fe nw nolan pe Sinha dhmnaydaey aneeve.sne sees swbard civjere ave
Page.
67
234
251
305
he + iy! :
. >! i
CRAG E SE CES OR 8 a aoe a Bg + ae a
— $ y
he Ge sa ‘ 5 Rint 2 ade
Dates of Issue. Part II, 1898.
No. I,—Containing pp. 1-250, was issued on 5th August, 1898.
aa {I].—Containing pp. 251—321, was issued on 15th September,
1898.
to
a ‘ ,
Bi boa re
hs cab iat tre
h
California Academy of Sciences
Presented byASiatic Society of —
Bengal.
a ee oe a
AOURNAL
OF THE
ASIATIC SOCIETY OF BENGAL.
== § E>
Vol, LXVII. Part II.—NATURAL SCIENCE.
No. I.—1898.
Materials for a Flora of the Mulayan Peninsula—By Groner Kina,
K.C.1.E., M.B., LL.D., F.R.S., Superintendent of the Royal Botanic
Garden, Calcutta.
No. 10.
I had hoped in the present contribution to have completed, for
these Materials, my account of the whole of the remaining Natural
Orders of Calyciflore. This hope has, however, been frustrated by sick-
ness. I have decided therefore to offer now to the Society the account
of the five Orders which I have been able to elaborate ; trusting, at
some time in the near future, to deal with the remaining Orders of
the Class. Following the sequence adopted by Sir Joseph Hooker in
his Flora of British India, those treated of in the present paper
come to be numbered as below; Nos. 48 Lythracexw, 49 Onagraceex, 50
Samydacee, 52 Oucurbitacex, and 56 Araliacex. And those which re-
main to be described would be Nos. 46 Myrtacex, 47 Melastomacex, 51
Passifloracee, 53 Begoniacew, 54 Ficoidex, 55 Umbelliferee, and 57
Oornacex. After finishing the Calycifloree, I hope, in collaboration with
my friend and successor, Dr. D. Prain, to describe the families which
are embraced in the gamopetalous and apetalous groups.
Order XLVIII. LYTHRACEHAL.
Trees, shrubs or herbs ; branches often quadrangular. Leaves entire,
opposite, sometimes alternate or whorled; stipules 0. Injlorescence
various, often in cymes or panicles, lowers hermaphrodite, regular,
ft |
2 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
rarely oblique, unisexual in Cyrpteronia. Calyx-tube free, persistent ;
lobes 3-6, valvate, some accessory often added. Petals as many as
the calyx-teeth, rarely 0, inserted near the mouth of the calyx-tube.
Stamens definite or numerous, inserted on the calyx-tube. Ovary free
in the bottom of the calyx-tube (rarely inferior), 2-6-celled, style long ;
stigma capitate, rarely 2-lobed; ovules numerous, placentas axile
(rarely parietal). Fruit coriaceous or membranous, free or more or
less adnate to the base of the calyx, 2~6-celled or (by absorption of
the partitions) l-celled, dehiscent or indehiscent. Seeds numerous,
various in shape, angular, sometimes winged; albumen none; embryo
straight, (cotyledons convolute in Sonneratia and Punica.) DistRIB.
Species about 275 in tropical regions and especially in those of the
New World; a few in temperate zones.
Tribe I. AMMANNIZ. Herbs, mostly sub-aquatic, with small
or minute flowers ; the calyx membranous aye . 1, AMMANNIA.
Tribe II, Lyturea. ‘Trees or shrubs with moder ie or large-
sized flowers (minute in Crypteronia), large often wrinkled
petals, and coriaceous or herbaceous calyx.
Stamens not more than 12.
Calyx 6-toothed ; petals 6; stamens 12; capsule circum-
scissile, 1-celled ; seeds cuneate-obovate, angled .. 2 .PEMPHIS;
Calyx 4- or 5-toothed ; petals 0; flowers numerous, minute,
racemose, sub-unisexual; stamens 4 or 5; capsule 2-celled
and 2-valved; seeds minute, narrowly winged on one
side Si as sie ae ... & CRYPTERONIA.
Stamens indefinite.
Seeds free, not imbedded in pulp.
Stamens in 2 or more rows; capsule 3-6-celled; seeds
large, winged laterally ate Je .. 4 LAGERSTREMIA.
Stamens in a single row; capsule 4-8-celled; seeds
minute, narrowly winged at the upper margin an DUABANGA.
Seeds imbedded in pulp, angular; berry 10-15-celled .. 6. SoONNERATIA.
<
1. AmmanniA, Linn.
Annual glabrous herbs growing in damp places; branches often
quadrangular. . Leaves opposite and alternate, sometimes whorled, entire ;
stipules 0. Flowers small, axillary, solitary and subsessile, or in small
trichotomous cymes ; bracteoles usually 2. Calyx campanulate or tubu-
lar-campanulate, 3-0-toothed, often with minute interposed teeth or
folds. Petals 3-5 or 0, small, inserted between the calyx-teeth. Stamens
2-8, inserted on the calyx-tube. Ovary enclosed in the calyx-tube, 1-5-
celled, the septa very thin and often absorbed; style filiform or short,
stigma capitate; ovules numerous, placentas axile. Capsule membran-
ous, globose or elongated-ellipsoid, enclosed in the calyx, 2-3-valved,
1898.] G. King-—Materials for a Flora of the Malayan Peninsula. 3
irregularly breaking up, or circumscissile. Seeds many, small, smooth,
round on the back and with a raphe on the inner face, ellipsoid or nearly
hemispheric; placenta ultimately free central by the absorption of the
dissepiments covered by the seeds. Distris. Species 30; in the tropi-
cal or warm temperate zones of the whole world.
Flowers sessile, calyx-tube elongate-campanulate, capsule
ellipsoid, seeds narrowly oblong, falcate oes .. 1. A. peploides.
Flowers pedicelled, calyx-tube depressed-hemispheric, cap-
sule depressed-globose, seeds sub-hemispheric... vasi-| De, A. baccifera.
1. AMMANIA PEPLOIDES, Spreng. Syst. I, 444. Flowers in short
axillary branches, sessile, solitary in the axils of reduced leaves ; bracts
in pairs, filiform, shorter than the tube of the calyx. Calyx-tube elon-
gate-campanulate, almost smooth, its mouth with 4 acutely triangular
teeth. Petals absent, or 4 and minute. Capsule 2-valved, ellipsoid ;
seeds narrowly oblong, sub-falcate, pink, angular, the hilum obscure.
Leaves opposite, their midribs prominent; those of the flower-bearing
branches linear-oblong, bearing a flower in the axil of each; those
of the main stem elliptic or obovate, narrowed to the base and almost
petiolate. Stems decumbent, often rooting, sometimes erect. Boiss.
Flor. Orient. II, 742; Kurz in Journ. As. Soc, 1877, pt. II, 84; Clarke in
Hook. fil. Flor. Br. Ind. I, 566. A. nana, Roxb. Flor. Ind. I, 427, (not
of Wallich). A. repens, Rottl., DC. Prodr. III, 80. Ameletia indica,
DC. in Mem. Soc. Hist. Nat. Genev. III, 11 (1825) 2, and 82,t. 3f. A.;
Prodr. III, 76; Wall. Cat. 2093; W.& A. Prodr. 303; Blume Mus.
Bot. II, 135, t. 47; Dalz. & Gibs. Bomb. Flor. 96; Wight Ic. t. 257.
A. elongata, Blume Mus. Bot. II, 185, A. acutidens, Miq. Flor. Ind. Bat.
J, Pt. 1,617. <A. polystachya, Wall. Cat. 2094. A. latifolia, Wall. Cat,
2096, (partly Peplis indica,) Willd. Sp. Pl. II, 244.
Soutp ANDAMAN; near the settlements of Port Blair and Port
Mowat; doubtless introduced as a weed of cultivation. Dzistris. India,
China, Persia; in rice and other fields.
2, AMMANNIA BACCIFERA, Linn. Sp. Pl. 120. Flowers in very con-
densed axillary racemes or clusters shorter than the leaves; bracts
filiform, shorter than the flower-pedicels. Calyx-tube widely campanu-
late, short, ridged ; the teeth 4, broadly triangular, acute. Petals none
or minute. Capsule depressed-globose, imperfectly circumscissile above
the middle. Seeds sub-hemispheric, black, excavated on the plane face.
Leaves opposite, rather distant, linear-oblong, sub-acute or obtuse,
narrowed at the base, smaller upwards, 2-5 in. long. Stem erect,
glabrous, 8-24 in. long. Blume Mus. Bot. Lugd. Bat. II, 133; Dalz. &
Gibs. Bomb. Flor. 97; Kurz in Journ. As. Soc. 1877, Pt. II, 85 ; Clarke
in Hook. fil. Flor. Br. Ind. II, 569. A, vestcatoria, Roxb. Hort. Beng. 11
4 G. King — Materials for a Flora of the Malayan Peninsula. [No. 1,
Flor. Ind. I, 426; ed. Wall. I, 447; DC. Prodr. III, 78; W. & A. Prodr.
305; Wall. Cat. 2098, (partly). A. indica, Lamk. Ill. I, 311, No. 1555 ;
DC. lic. 77; W. & A. Prodr. 305; Wall. Cat. 2099; Blume 1. c. t. 46.
A. debilis, Ait. Hort. Kew, ed. 1, I, 163. A. verticillata, Boiss.
Flor. Orient. II, 743, (not of Link). Hapalocarpum vesicatorium and
H. indicum, Mig. Flor. Ind. Bat. I, Pt. J, 618. Cryptotheca apetala,
Blume Bijd. 1128; DC. 1. ec. 76.
Perak ; King’s Collector 303; Curtis 3195. S. Andaman, in similar
situations with the last.
2. Pempuis, Forst.
A maritime shrub or tree, 25-35 ft. high. Leaves opposite, oblong
to oblong-lanceolate, entire, very thick, fleshy. Flowers small, axillary,
solitary, peduncles 2-bracteate at their base. Calya-tube campanulate,
12-co -ribbed ; teeth 6, short, with 6 shorter accessory teeth. Petals 6,
inserted at the mouth of the calyx-tube and nearly as long, obovate,
wrinkled, white or rose. Stamens 12, inserted in two series towards
the middle of the calyx-tube. Ovary free at the bottom of the calyx-
tube, 3-celled at the base; style long, stigma capitate; ovules many,
ascending ; placentas 3, sub-basal. Capsule coriaceous, obovoid or nearly
globose, included in the calyx-tube or exserted nearly half its length,
somewhat irregularly circumscissile, ultimately 1-celled. Seeds very
many, long cuneate-obovoid, angular, smooth, standing out in all direc-
tious from the apparently free central placenta.
Pempis acipuLA, Forst. Gen. t. 34. Young parts more or less
clothed with grey silky hairs, the young branches 4-angled. Leaves
subsessile or very shortly petioled, sub-acute or obtuse, *5—-1°5 in. long.
Flowers white. Capsule ‘4 in. long and ‘2 in. in diam. DC. Prodr. III,
89; Wall. Cat. 2108; W.and A. Prodr. 307; Griff. Notul. IV, 510;
Blume Mus. Bot. II, t. 43; Miq. Flor. Ind. Bat. I, pt. I, 619; Bedd.
Flor. Sylv. Anal. Gen..t. XIV, fig. 5; Kurz For. Flor. I, 518.
PRAingustifolia, Roxb. Hort. Beng. 91; Flor. Ind. II, 465. P. setosa,
Lour. Flor. Filip. ed.1, 410. Maclellandia Griffithiana, Wight Ic. t. 1996.
Lythrum Pemphis, Linn. f. Suppl. 249; Lamk. Ill. II, 408, fig. 2.
Melanium fruticosum, Spreng. Syst. IT, 445.
On the beech in Sincapore and probably in all the provinces.
Awnpaman and Great Coco Isnanps; Prain. Disrris. Burma, Ceylon,
S. of British India,
3. CRYPTERONIA, Blume.
Trees. Leaves opposite, petioled, entire, ovate or lanceolate.
Racemes elongate, in branched panicles. Flowers minute, white or green,
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 5
with short linear bracts at the base of the pedicels, polygamo-dicecious.
Calyzx-tube short, saucer-shaped, or longer and subhemispheric; teeth
5 (rarely 4), valvate, persistent. Petals0. Stamens as many as the
calyx-teeth, inserted between them near the mouth of the calyx-
tube. Ovary free, 2-celled, with numerous horizontal or ascending
ovules attached to the axile placentas ; style long, filiform; stigma capi-
tate, obscurely 2-lobed. Capsule surrounded at the base by the calyx,
globose, pubescent, crowned by the persistent style, 2-celled, dehiscing
so as to divide the style, fruit-pedicel deflexed. Seeds many, elongate-
ellipsoid, narrowly winged on one side. Disrris, Species 5, extending
from the Khasia Hills to the Philippine Islands.
Leaves membranous, usually narrowed to the base, nerves
5 or 6 pairs, calyx less than ‘1 in. in diam. Je .. IL. C. paniculata,
Leaves coriaceous, rounded or cordate at the base, nerves
7 or 8 pairs, calyx more than ‘lin.indiam. ... we oe 6 C, Griffith.
1. CRYPTERONIA PANICULATA, Blume Bijdr. 1151. A tree 20-40
feet high ; young branches glabrous or sometimes puberulous. Leaves
membranous, oblong to oblong-lanceolate or more or less broadly
elliptic, bluntly acuminate or blunt, narrowed to the base, entire,
glabrous on both surfaces or slightly pubescent on the Jower; main
nerves 5 or 6 pairs, rather faint, curved ; length 3-6 in., breadth 1°25-
2°5 in. Flowers on short pedicels, very numerous, in long cylindric
pubescent or glabrous racemes longer than the leaves, the racemes
often panicled, Calya less than ‘1 in. in diam., its teeth triangular or
triangular-lanceolate, acute. Stamens in the hermaphrodite flowers of
the same length as the calyx-teeth, longer in the male flowers. Capsules
globose-conic, puberulous or minutely velvety. Kurz in Journ. As. Soc,
Beng. 187, Pt. 11, 86; For. Flora Burma, I, 519. C. pubescens, Blume
Mus. Bot. Lugd, Bat. IT, 123; Clarke in Hook. fil. Flor. Br, Ind. II,
574; Griff. Notul. IV, 404; Ic. Pl. Asiat. t. 564, fig, II. OC. glabra,
Blume Mus. Bot. II, 123; Clarke in Hook. fil, Flor. Br. Ind. IT, 574.
Henslovia pubescens, Wall. Cat. 4904; Pl. As. Rar. III, 14 t. 221; Mig.
Flor. Ind. Bat. I, Pt. I, 716; Planch, in Hook. Lond. Journ. Bot. IV,
477, t. XVI. B, Henslovia Hookeri, Wall. Cat. 8566. H, affinis, Planch,
Lond, Journ. Bot. IV, 477 (in part). H. leptostachys, Planch. Lond.
Journ. Bot. IV, 478. H. glabra, Wall. Cat. 4093; Pl. As. Rar. III, 14;
Planch. in Hook. Lond. Journ, Bot. IV, 478 ; Miq. Flor. Ind. Bat, I,
Pe, 716.
Penanc; Porter. Matacca; Maingay (Kew Distrib. 650/2).
Perak; King’s Collector No. 5205. AnpaMAaNn IsLANDS; very common ;
King’s Collector. Distris. Burma, Chittagong, Khasia.
I can find nothing better to distinguish the species which have been named
6 G. King— Materials for a Flora of the Malayan Peninsula. \ [No. 1,
C. glabra and C. pubescens from each other than the presence on the latter of a
small amount of hair, neither can I find any tangible character to separate either
from C. paniculata, Blume. To the latter species, as the oldest, I therefore reduce
both,
2. Oryprmronta Grirrirnit, Clarke in Hook. fil. Flor. Br. Ind. II,
574. A tree 40-60 feet high; young branches glabrous. Leaves
coriaceous, broadly elliptic, acute, very slightly narrowed to the rounded
or cordate ‘base, entire, glabrous on both surfaces; main nerves 7 or 8
pairs, distinct on the lower surface, curved, ascending; length 4-8 in.,
breadth 2°25-3°5 in. Racemes much longer than the leaves, rusty-
pubescent, sometimes panicled; flowers numerous but not crowded,
shortly pedicellate. Calyx rather more than ‘1 in. in diam., its teeth
triangular. Stamens exserted, Capsule shorter than the calyx-teeth,
velvety, less than *l in. in diam., crowned by the long stout pubescent
style. Hensloview sp. Griff. Notul. 406; Ic. Pl. Asiat. t. 564, fig. 1.
Maracoa; Griffith 2518. Maingay (Kew Distrib.) 651. Derry
1201. Penang; Curtis 1739. Prrax; Scortechint 221, Wray 2589,
2638. King’s Collector 3473, 4152, 8592.
4, LAGERSTR@MIA, Linn.
Trees or shrubs. Leaves opposite, distichous (or the uppermost
alternate), entire, oblong or ovate. Panicles axillary and terminal,
the ultimate branchlets usually cymose, sometimes dense; peduncles
2-bracteate at their apex; pedicels 2-bracteolate. Flowers often large,
Calyx-tube funnel-shaped, smooth, grooved, angular or sub-alate ; lobes
5-sometimes 7-9, ovate, subacute, valvate. Petals 6, sometimes 7-9
(or 0), inserted at the summit of the calyx-tube, clawed, wrinkled ;
margin crisped, erose, or fimbriate. Stamens very many, inserted in
several rows near the bottom of the calyx-tube ; filaments long, exserted.
Ovary sessile in the bottom of the calyx, 3-6-celled; style long,
bent, stigma capitate; ovules numerous, ascending, placentas axile.
Capsule more or less adnate to the calyx, ellipsoid, coriaceous, smooth,
3-6-celled, 3-6-valved. Seeds many (rarely few), elongate, flat, erect,
winged from their summit. Disrris. Species 18, in South-east Asia
extending to Australia, :
Inflorescence and exterior of calyx stellately ferrugineous-
tomentose ... a ae sh 1. LZ. floribunda.
Inflorescence and outside of calyx minutely cinereous or
whitish-velvety (the calyx sometimes rusty in L. Flos-
Regine).
Leaves glaucous beneath; calyx 10-ribbed, the mouth
with only 5 teeth ... R me a .. 2, L. hypoleuca.
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 7
Leaves not glaucous beneath.
Calyx 8- or 9-ribbed, the teeth as many as the ribs and
alternate with them; flowers 1°5 in. in diam.; leaves
2—3°25 in. long ae oa
Calyx 12-14-ribbed; the teeth half as many as the
ribs, the ribs opposite the teeth broader ; flowers 2-3
in. in diam.; leaves 3°5-8 in. long te wv. 4 OD. Flos-Regine.
3. L, ovalifolia.
1.° LAGERSTR@MIA FLORIBUNDA, Jack in Mal. Misc. I, 38. A tree
15-30 feet -high. Leuves ovate-oblong or elliptic-oblong, sub-acute, the
base rounded, sub-sessile; main nerves 8-12 pairs, sub-horizontal or
curving upwards; both surfaces minutely reticulate when dry, the upper
glabrous and shining, the lower with deciduous stellate pale brown
pubescence, or glabrous; length 5-7 in., breadth 2-2°75 in. Punicle
much longer than the leaves, terminal, erect; the branches long, race-
moid, the ultimate branchlets cymose, ascending, everywhere (as also the
calyces,) covered with more or less deciduous wooly rusty pubescence.
Flowers 1°5 in, in diam.,on short pedicels. Calyx turbinate in bud,
boldly 12-ridged, each alternate ridge passing into one of the 6
triangular calyx-teeth and often forming a mucro at its apex. Petuls
sub-orbicular, with wavy edges, rose-coloured changing to whitish.
Stamens unequal, the outer rows the longest. Capsule *5 in. long,
elliptic, minutely cinereous-tomentose, half enveloped in the calyx,
style persistent. DC. Prodr. III, 93; Wall. Cat, 2115; Mig. Flor. Ind.
Bat, I, pt. I, 623 (not Blume Mus. Bot. I, t. 41) ; Griff. Notul. LV, 509;
Kurz For. Flor. I, 522; Clarke in Hook. fil. Flor, Br. Ind. II, 577.
Kepau; Curtis 2602. Penang; King. Trane; King’s Collector
1407, Matacca; Maingay (Kew Distrib.) 653/2. Distrris. Burma,
Siam, China. ~
2, LaGgerstra@mia HypoLeuca, Kurz in Journ. As. Soc. Beng. for 1872
Pt. II, p. 30. A tree 60-70 feet high; all parts except the inflorescence
glabrous. Leaves thickly membranous, oblong-lanceolate to elliptic
or oblong-elliptic, shortly acuminate, the base rounded; main nerves
8-12 pairs, spreading, not prominent; both surfaces rather distinctly
reticulate when dry, the upper shining, the lower glaucous; length
5-8 in., breadth 2-3 in., petiole *3 in. long. © Panicles minutely cinereous-
velvety, longer than the leaves, terminal, few-branched, the branches
with rather short cymose sub-horizontal branchlets. Flowers about
1°25 or 1°5 in. in diam., on jointed whitish unequal pedicels. Calyx
turbinate in bud, minutely whitish-velvety, boldly 10-ribbed, the
alternate ribs excurrent into the 5 triangular acute mucronate lobes
of the mouth. Petals lilac, oblong, wavy, ‘5 in. long. Capsule woody,
oblong, mucronate, about ‘65 in. long. For. Flor. Burm., I, 528 ;
Clarke in Hook. fil. Flor. Br. Ind. II, 577.
8 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
AnpaMAN Istanps; Kurz, Prain, King’s Collectors, Gruar Coco
IsuAND; Prain.
3. LAGERstR@MiIA oOvaLirouia, Teysm. et Binn. in Nat. Tijdsch.
Ned. Ind. II. (1840) 306. A tree 50 or even 100 feet high. Leaves
oblong, ovate or oblong-obovate, acute, slightly narrowed to the base ;
main nerves 4 or 5 pairs, curved, ascending, slightly prominent on the
lower surface when dry; upper surface greenish with minute black
dots when dry, glabrescent except sometimes the minutely pubescent
nerves; the lower brown when dry, glabrescent; length 2-325 in.,
breadth 1°25-2 in., petiole ‘25 in. Panicle terminal, puberulous, 3-8
in. long, few-flowered ; the branches few, sub-horizontal, cymose, puberu-
lous below ; the pedicels minutely velvety, cinereous. Flowers 1'5 in.
in diam., on jointed velvety pedicels, Calyx turbinate, minutely
cinereous-velvety, with 8 or 9 bold winged ridges not passing into the
calyx-teeth. Calyz-teeth 8 or 9, acutely triangular, the edges thickened
and reflexed. Petals (with claw) °75 in. long, orbicular, clawed, purple,
their edges slightly undulate. Capsule elliptic-ovoid, blunt, minutely
velvety, ‘75 in. long, and °6 in. in diam. Kriudk. Arch. ITI, 440; Pl.
Nov. Hort. Bog. (ed. Vriese) 20. Blume Mus. Bot. Lugd. Bat. II, 127;
Mig. Flor. Ind. Bat. I, Pt. I, 624; Koorders and Valeton, Bijdr. I, 193.
LL. celebica, B31, 1. ¢c. 127. DL. hexaptera, Miq. Flor. Ind. Bat. I, Pt. I, 623;
Clarke in Hook. fil. Flor. Br. Ind. IT, 577.
Matacca; Maingay (Kew Distrib. 653), Prrax; King’s Collectors
8701, 10025 & 10532. Panane; Ridley 2640.
I can find no character to separate L. hexaptera, Miq. from the older species of
Teysmann and Binnindik. Miquel’s name is moreover an unhappy one, as the
calyx-teeth are usually 9 and not 6.
4, Lacerstramra Fros-Recing, Retz (1789) Obs. V, 25, <A tree
30-60 feet high, all parts except the inflorescence glabrous. Leaves
oblong to elliptic-oblong, acute, narrowed (rarely obtuse) at the base,
shortly petiolate; main nerves 10-13 pairs, curving upwards, slightly
prominent beneath when dry; both surfaces minutely reticulate,
glabrous, the upper shining, the lower dull and of a dark brown colour
when dry, length 3°5-8 in., breadth 1°75-3 in., petiole ‘25-4 in.
Panicle terminal, longer than the leaves, its ultimate branchlets cymose.
Flowers from 2-8 in. in diam., on rather thick greyish unequal pedicels.
Calyx turbinate, with 12-14 prominent stout ridges; those opposite the
calyx-teeth broader, the mouth with 6-7 acute triangular spreading
thick coriaceous teeth thickened at the edges. Petals sub-orbicular,
clawed, corrugated and with undulate edges. Stamens all equal in
length. Capsule oblong to sub-globose, minutely apiculate, ‘8-1°25 in.
‘long, and ‘6-75 in. indiam, Kurz in For. Flora Burm, I, 524; Clarke in
1898.] G. King—Materials for a Flora of the Malayan Peninsula. 9
Hook. fil. Flor. Br. Ind. II, 577. DL. Munchhausia, Lamk. Ency. IIT, 375 ;
Ill. t. 473 fig. 2. DL. Reginee, Roxb. Pl. Corom., I, 46, t. 65; Hort. Beng.
88; Hook. fil. Flor. Br. Ind. IJ, 505; Blume Bijdr. 1127; DC. Prodr.
III, 93; W. & A. Prodr. Flor. Penins. Ind, 308; Blume Mus. Bot,
Lugd. Bat. II, 126; Mig. Flor. Ind. Bat. I, Pt. I, 623 and Suppl. 328.
Li. speciosa, Pers. (1807) Ench. II, 72 (not of DC.); Koehne in Engl.
Bot. Jahrb. IV, 28; Koorders and Valeton, Bijdr. I, 190, (excl. from
all where reduced the syn. L. macrocarpa, Wall.).
Matacca, Singapore. Perak. Disrris. Java, British India.
I have adopted Retz’s name (published in 1789) for this plant, as it is pretty
nearly certain what Retz’s plant was. Koehne, Koorders and Valeton and others
however adopt Persoon’s name of L. speciosa on the ground that, although it dates
from only 1807, it preserves the specific name of Linnzeus (Munchhausenia speciosa
1770). But this procedure is rendered inadmissable when Linnezus’s description of
that plant is consulted, for he describes M. speciosa as a shrub, whereas this plant is a
large tree; moreover the rest of his description would apply to various other species
of Lagerstremia; the identity of M. speciosa, L. is thus quite uncertain. L. macro-
carpa of Wall. Cat. 2114, isa tree of about the size of L. Flos-Regine and resem-
bles it in most respects, but differs (1) in having leaves of larger size (5-12 in.
long) more or less broadly elliptic, never oblong or elliptic-oblong, the apex often
sub-acute and the base broad or narrowed into a petiole twice as long as that of
L. Flos-Regine ; (2) in the calyx being very slightly, if at all, furrowed and never
ribbed, and (3) in having a larger more globose capsule (1-1°35 in. long and nearly
as much in diam.). This tree has been in cultivation in the Bot. Garden, Calcutta
side by side with L. Flos-Reginz for many years. It flowers earlier than the latter,
has pink (not lilac) petals, and much larger capsules. In my opinion it is a good |
species and should not be merged in L. Flos-Reginz. Itis found only in Burma
and Chittagong. Kurz, who was familiar with it in its wild state in Burma, con-
sidered it distinct and kept it as a species in his Forest Flora of British Burma.
5. Duapancoa, Ham.
Large glabrous trees with pendent quadrangular branches. Leaves
opposite, distichous, large, short-petioled, elongated, acute, entire, cordate
or rounded atthe base. Panicles large, terminal, with opposite branches ;
flowers large. Calyx-tube wide, adnate to the base of the ovary; lobes
4-7, thick, valvate in the bud. Petals 4~—7, clawed, obovate, crisped
and undulate, white. Stamens very many, inserted on a perigynous
ring. Ovary conical, 4-8-celled; style bent, long; stigma capitate,
4-8-lobed ; ovules very many, ascending, placentas covering nearly
the whole interior surface of the ovarian cells. Capsule sub-globose,
surrounded at the base by the thick spreading calyx, coriaceous, perfectly
or imperfectly 4-8-celled, 4—8-valved. Seeds very numerous, minute,
ellipsoid, testa produced at both ends in two tails much exceeding the
length of the nucleus. Disrxis. Species 2; Hastern Himalaya, Assam
and Malaya, |
J. u. 2
10 G. King— Materials for a Flora of the Malayan Peninsula. (No.1,
DUABANGA SONNERATIOIDES, Ham. in Trans. Linn. Soc, XVII, 178.
A tree 60-100 feet high. Leaves thinly coriaceous, oblong to ovate-
oblong, 7-12 in. long and 2°5-4 in. broad, glabrous, glaucous beneath.
Flowers 2—2°5 in, across, on thick-jointed pedicels tapering to the base ;
panicle short, few-flowered, drooping. Petals 4-7, about 1 in. long.
Capsule ovoid-globose, ]-1°5 in. in diam. Hook. fil. Hl. Him. Pl. t. 11;
Kurz For. Flor. Burm. I, 525; Clarke in Hook. fil. Flor. Br. Ind. II, 579.
D. grandiflora, Walp. Rep. II, 114. Lagerstremia grandiflora, Roxb.
Hort. Beng. 38; Flor. Ind. II, 503; DC. Prodr. III, 93; Wall. Cat.
2111; Blume Mus. Bot. I, 109. Leptospartion grandiflorwm, Griff. Ic.
Pl. Asiat, t. 591.
Pprak; Scortechint, King’s Collector 5912. AnpdAMans AND NICOBARS,
King’s Collectors, Distrrp. Burma, Assam, Khasia, Eastern Himalaya,
6. Sonneratia, Linn. f.
Ever-green trees, growing near the sea, glabrous. Leaves opposite,
petioled, coriaceous, entire. Flowers without bracts, large, three to-
gether at the summits of the branches, or axillary and solitary. Calyx
thick, coriaceous ; lobes 4-8, lanceolate, valvate. Petals 0, or as many
as the calyx-lobes and linear-oblong, Stamens numerous, inserted in a
circular band on the calyx-tube. Ovary nearly free, or adnate at the
base to the calyx-tube, many-celled ; style long, stigma capitate; ovules
numerous, ascending, placentas axile. Berry subglobose, supported by
the persistent calyx, 10-15-celled. Seeds very many, small, curved,
angular, embedded in pulp; cotyledons convolute. Dustin. Species
4—5 ; on the tropical sea-shores of the Hastern hemisphere.
With petals i oes #58 ees .. Ll. §. acida.
Without petals :—
Calyx in bud ovoid, obtuse ... ese ae eo. &. Si@erta,
var. Griffith.
Calyx in bud narrowly ellipsoid, tapering to both ends oe «62. | Oe aLUae
1. Sonneratia acipa, Linn. fil, Suppl. 252. A small tree 10-35
feet high, the branchlets jointed and 4-angled, Leaves oblong to oblong-
obovate or obovate, tapering into a broad short petiole, blunt and
sometimes retuse ; length 2~3 in., breadth 1°35 to 2 in. in the obovate
forms. Slower-buds solitary, ellipsoid, the calyx-tube not angled; the
calyx when fully developed 1 in. long, its lobes 6-8, the lobes triangular,
reflexed, Petals linear, slightly broader towards tle apex than at the
base. Style exserted, sometimes 3 in, long, stigma capitate, Capsule
depressed-globose, sometimes as much as 2 in. in diam,, the apex some-
what concave, the walls thick, Roxb, Hort. Beng. 38; Flor. Ind. II,
506; Roth Nov, Sp. 233; DC, Prodr. III, 231; Wall. Cat. 3641;
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 11
W.& A, Prodr. 327 ; Wight Ic. t. 340; Griff. Notul. IV, 652; Blume
Mus. Bot. I, 336; Miq. Flor. Ind. Bat. I, Pt. I, 496; Dalz. & Gibs. Bomb.
Flor, 98; Brand. For. Flor. 242; Kurz For. Fl. Burm. I, 526; Clarke
in Hook. fil. Flor. Br. Ind, II, 580; Koorders and Valeton, Bijdr. I, 198.
Rhizophora caseolaris, Linn. Sp. Pl. 635. Aubletia caseolaris, Geertn,
Fruct. I, 479, t. 78.
PenanG; Curtis 1108. Perak; Scortechini, Wray 2494. ANDAMANS;
Kurz, Prain, King’s Collectors, Distr1p. the coasts of Burma, the
Deltas of the British Indian Rivers and of those of the Malayan Islands.
Var. Griffithii, Leaves obovate, petals none. 8S. Griffithii, Kurz
Peeu Report, App. B, 54; For. Flora Burma, I, 526 ; Clarke in Flor. Br.
Ind. II, 580. 8S. alba, Griff. (not of Smith) Notul. IV, 652. &. neglecta,
Bl. Mus. Bot. Lugd. Bat, I, 338; Miq. Flor. Ind. Bat. I, Pt. I, 498.
Perak; Scortechini 967. Distris. Burma.
The fruit of this variety is unknown; but the scanty material, so far as it goes,
appears to show that the plant is merely an apetalous form of §. acida with leaves
more obovate than is usual in that species. I have therefore reduced it to a form
of the latter. Blume described other three species of Sonneratia with petals,
viz., 8. obovata, S. evenia, and 8S. lanceolata separating them chiefly by characters
taken from the shapes of the leaves. But in this genus the form of the leaf is very
variable, and I doubt whether these three species are more than forms of S. acida.
§. Pagapat, Blanco, and S. ovalis, Korth. are probably also forms of it.
2. SoONNERATIA ALBA, Smith in Rees Cyclop. XX XIII, No. 2. A small
tree 10-15 feet high ; young branches rather terete. Leaves obovate or
obovate-reniform, decurrent on the short petiole, blunt or retuse, 2—4 in.
long and nearly as broad, petiole *125-25 in. Flower-buds narrowly
ellipsoid, tapering to each end, very slightly ridged; the fully developed
calyx sharply angled ; its lobes 6-8, oblong-lanceolate, acute. Petals none.
Flowers about the size of those of §. acida, usually 2 or 3 together.
Capsule broadly obconic, ribbed, 1 in. or more in diam. at the apex.
DC. Prodr. III, 231; Blume Mus. Bot. Lugd. Bat. I, 338; Mig. Flor. Ind.
Bat. I, Pt. I, 497; Kurz For. Flora Burma I, 526; Clarke in Flor.
Br. Ind. II, 580; Koorders and Valeton, Bijdr. I, 200. 8S. Mossambi-
censis, Klotsch in Peters Reis. Mossamb, Bot. t, 12. S. acida, Benth.
(not of Linn. fil.) Flor. Austral. III, 301; Hiern in Oliv. Flor. Trop.
Afric. II, 483; Wall. Cat. 3641 B.
Singapore; Wallich, Distris. Java, Moluccas.
Order XLIX. ONAGRACEA.
Herbs, rarely undershrubs, sometimes aquatic. Leaves opposite or
alternate, entire or toothed, undivided (in Trapa the submerged leaves
pinnatipartite), exstipulate. Flowers hermaphrodite, mostly axillary
and solitary, or spiked or racemed towards the ends of the branches,
12) G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
sub-irregular. Calya-tube wholly adnate to the ovary (half-adnate in
Traupa), linib with 2-5 valvate lobes. Petals epigynous, alternate with
the calyx-lobes, rarely 0. Stamens as many or twice as many as the
petals, inserted with them. Ovary inferior (half-inferior in Trapa),
1—6-celled, most often 4-celled; style 1, cylindric or subulate, stigma
capitate or nearly 2-lobed or 4-fid; ovules one or many in each eell,
pendulous or half-ascending, placentas axile. Fruit various, dehiscent
or indehiscent, membranous capsular or bony, 1- or several-celled, 1 or
o-seeded. Seeds without albumen, or nearly so, Dusrris. Species 300,
spread throughout the world, most abundant in the North Temperate
Zone.
Stamens twice as numerons as the calyx-lobes... oo. |. JUSSIMA.
Stamens equal in number to the calyx-lobes_ .,. .. 2 LUDWIGIA.
1. Jussima, Linn.
Herbaceous or suffruticose, sub-aquatic. Leaves simple, alternate,
usually entire. lowers white or yellow, solitary, axillary; pedicel
usually bibracteate at the apex. Culyax-tube narrow, only slightly
produced above the ovary ; its teeth 4—6, acute, persistent. Petals 4-6,
epigynous. Stamens also epigynous, twice as numerous as the petals.
Ovary inferior, 4-5-celled ; style simple, usually very short, the stigma
4-or 5-lobed ; ovules numerous, axile, in several vertical rows at the
inner angle of each cell. Capsule narrow, cylindric or angled, 4- or 5-
celled, 8-10-ribbed, dehiscing septicidally. Seeds very numerous, with-
out coma. Disrris. Species 30, tropical, chiefly American.
JUSSIZA SUFFRUTICOSA, Linn, Sp. Pl, 555. Hrect, branching, 1- 4 feet
high. Leaves lanceolate, ovate-lanceolate or almost linear, acute or
acuminate, narrowed to the base, villous, pubescent or arib- pinkie
2-3 in. long, and ‘25-75 in. broad, sessile or very shortly petioled.
Flowers ‘5-75 in. in diam., on very short pedicels, the bracts small or
foliaceous. Petals wholly yellow. Capsule linear, cylindric, 1-2 in.
long, membranous, not woody, 8-ribbed, deciduously villous or pubes-
cent. Seeds sub-hemispheric; the testa with a prominent raphe, shining,
brown, not corky. DC. Prodr. 117, 58; Wall. Cat. 6334; Mig. Flor. Ind.
Bat. I, Pt. I, 628; Kurzin Journ. As, Soc. 1877, Pt. 11, 90; Clarke in Flor.
Br. Ind, Il, 587. J. ewaltala, Roxb. Hort. Beng, 33; Flor. Ind. I], 401.
J, villosa, Lamk. Dict. III, 331; DC. Prodr. III], 57; Wall. Cat. 6333;
W. & A. Prodr. 336; Gibs. & Dalz. Bomb. Flor. 98. J. fruticosa, DC.
lee. J. scabra, Willd.; DC, lic. J. Burmanni, and octophila, DC. 1. c.
J. longipes, Griff. Notal. IV, 689. J. decumbens, Wall. Cat. 6322.
J.angustifolia, Lamk. Dict. III, 331 and IIL, t. 280, fig. 3; DC. Prodr. III,
55; Mig. Flor. Ind. Bat. I, Pt. I, 627, Epilobium fruticoswm, Lour, Flor.
Dopha china 226. Rheede Hort. Mal. I, t, 50.
1898.| G. King-~ Materials for a Flora of the Malayan Peninsula.) 13
SmyncaporeE; Penanc; Perak; ANDAMAN IstanDs, and probably in
all the other provinces; Distris. British India, Ceylon,
A widely distributed plant to which many names have been given. Itis
readily distinguished from J. repens, Linn. (the only other species common to the
tropics of both worlds) by its narrower leaves, membranous capsule and erect habit..
2. Lupwiaia, Linn.
Herbs, Leaves alternate, undivided, sub-entire. Flowers usually
axillary, solitary, sessile or nearly so, peduncle 2-bracteate at its apex.
Culyx-tube scarcely produced above the ovary, linear in the Indian
species; teeth 3-5, acute, persistent. Petals 3-5 (or 0), epigynous,
yellow. Stamens equal in number to the calyx-segments, epigynous.
Ovary inferior, 4-5-celled ; style simple, stigma capitate; ovules very
many, attached in 2 or more vertical rows to the inner angle of each
cell. Capsule linear or oblong (in the Indian species), 4—5-celled, open-
ing by terminal pores or breaking up irregularly along the sides. Seeds
numerous, obovoid, smooth, raphe obscure or prominent but not large,
without coma. Dustris, Species 20, mostly in North America; extend-
ing from the cool temperate zone to the pete chiefly funapienie
marshes.
Capsules inflated, seeds in several rows re wv. IL. DL. parvifiora.
Capsules filiform, not inflated; seeds in a pag row in
each cell
=! pat a, 2. L. prostrata.
1. Lupwieia parvirtora, Roxb. Hort. Beng. 11; Flor. Ind. I, 419.
An erect glabrous herb 8-24 in. high. Leaves taicalbtoite, linear-lanceo-
late or linear-oblong, narrowed to each end, 1-3 in long, and ‘25-°75 in.
broad. Flowers on short pedicels, usually 4-fid. Petals small. Capsule
inflated, obsoletely 4-angled, smooth, crowned for sometime by the
persistent calyx-teeth, ‘35-5 in. long. Seeds in many rows in each cell.
DC. Prodr. III, 59; Wight Ill. t. 101; W. & A. Prodr. 336; Dalz. &
Gibs. Bomb. Flor. 99; Benth. Flor. Austral. III, 307; Boiss. Flor.
Orient. IJ, 752; Kurz in Journ. As. Soc. 1877, Pt. II, 91; Clarke in
Hook. fil. Flor. Br. Ind. IT, 588, lL. lythroides, Blume Bijd. 1134;
DC. 1.c. D. jussizoides, Wall. Cat. 6335 (not of Linn. and others).
Perak; Scortechint 442; Wray 2720. ODistris. British India
and Ceylon. |
2. Lupwicta prostrata, Roxb. Hort. Beng. 11; Flor. Ind. I, 420.
Stem prostrate, or decumbent at the base and then erect, 8-24 in. long,
glabrous. Leaves lanceolate or linear-lanceolate, acute or acuminate,
tapering to the base, 2—4 in. long and °35—75 in. broad. Flowers some-
times more than one in an axil, sessile, 4-fid. Petals lanceolate, longer
than the calyx. Capsule 4-angled, thin, filiform, not at all inflated,
14 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
smooth, crowned by the linear calyx-teeth ; length *5-1 in. diam. ‘02 in.
Seeds oval, ina single row in each cell, DC. Prodr. IIT, 59; Wight Ic. t.
762; Kurz in Journ. As. Soc. 1877, Pt. IJ, 91. Clarke in Flor. Br. Ind,
II, 588. L, diffusa, Ham. in Trans. Linn. Soc. XIV, 301; Wall. Cat.
6336; DC. 1. c DL. fruticulosa, Blume Bijd. 1133; DC.1.¢. Nemato-
pyxrs prostrata, pusilla, and fruticulosa, Miq. Flor. Ind, Bat. I, Pt. I, 630.
Perak; Curtis 3183. Sincapore; King’s Collector 58. ANDAMANS;
King’s Collector, Distris. Burma, Cachar, Sylhet, Assam, and the base
of the Hastern Himalaya.
A species closely resembling L. parviflora, but readily distinguished by having
very slender capsules, through the walls of which the seeds may be distinguished
lying in single row in each cell; whereas in L. parviflora the seeds are in several
rows in each cell and are undistinguishable on the outside of the thick walls of the
short inflated capsules.
Order L, SAMYDACEA.
Trees or shrubs. Leaves alternate, often distichous, petioled, rarely
subsessile, simple, entire or slightly crenate or serrate, often closely
punctulate beneath; stipules small, deciduous. Flowers regular, small,
axillary, shortly pedicelled, densely fascicled or in long simple or
panicled racemes. Calyx coriaceous, persistent; tube short, free, or
longer and adnate to the ovary; limb 3-7-fid. Petals as many as the
calyx-lobes (or 0), perigynous, imbricated. Stamens definite or indefi-
nite, often with staminodes between or united in a tube with them.
Ovary superior or half-superior, 1-celled ; style 1, capitate or 3-fid at the
apex, or styles 2-5; ovules many or several, placentas 2-5 (usually 3),
parietal. Fruit loculicidally 2-5- (usually 3-) valved, valves carrying
the seeds on their mesial line. Seeds several (usually few, sometimes
many), oblong or angular, albuminous, usually drilled. Disrris.
Species 180, scattered through the tropical regions of the globe, rarely
also in the subtropical.
Flowers without petals :—
Flowers in axillary glomeruli, rarely solitary ... .. 1. CASEARIA,
Flowers in long slender racemes or panicles... .. 2 OSMELIA.
Flowers with petals; flowers in axillary or terminal racemes
or panicles ose Bo ee nas .. & HOMALIUM,
1. Casearia, Jacq.
Shrubs or small trees. Leaves alternate, distichous, petioled, un-
divided, entire or slightly serrate, often minutely punctate; stipules
small, lateral, caducous. Flowers small, greenish-yellow, clustered in
the axils (in the Indian species) ; pedicels short, jointed above their
bases, surrounded by small scales. Calyx inferior, deeply 4~5-lobed ;
1898.] G. King—NMaterials for a Flora of the Malayan Peninsula. 15
lobes imbricate, obtuse, persistent. Petals 0. Stamens double the number
of the calyx-lobes or thereabout, united in a tube with staminodes
alternating with the free portion of the filaments; staminal tube hypo-
gynous, sometimes very short so that the filaments are nearly or quite
free. Ovary free, ovoid, 1-celled, style simple, stigma capitate or 3-fid;
ovules many, parietal. wit succulent, globose or ovoid, ellipsoid
(when dry somewhat 3-angular or 6-ribbed), 3-rarely 2-valved. Seeds
many, angular or obovoid, with a fleshy usually coloured aril ; embryo
straight. Distris. Species 140, in the warmer parts of the whole globe,
most plentiful in America.
Leaves pubescent on the lower surface, glabrescent on the
upper :—
Leaves narrowed or rounded, but never cordate or trun-
cate, at the base ; glomeruli few-flowered .., .. Ll. CG. Lobbiana.
Leaves cordate or truncate at the base; glomeruli many-
flowered ... Ae me ae Palate a, grewiefolia.
Leaves glabrous on both surfaces :—
Fruit not exceeding 1 in. in length and usually much
shorter :—
Leaves broadly elliptic, shortly acuminate, thinly mem-
branous ; calyx-teeth 4 ee os .. 3& @. albicans,
Leaves elliptic-oblong, thinly coriaceous or membran-
ous, main nerves 10-12 pairs; stamens 10 .. 4. GC. andamanica.
Leaves elliptic-oblong to elliptic-lanceolate, main nerves
6-8 pairs; stamens 8 ve ae .. 5. C. esculenta.
Fruit large, fleshy, 1°5-2 in. long; leaves narrowly
oblong; coriaceous.
Flowers in glomeruli :—
Leaves tapering slightly to base and apex; fruit
apiculate Kise “ae - 6 GC. Kunstleri.
Leaves acute at the apex, subs acute at the Haines
fruit not apiculate a wv T OC. Clarke.
Flowers solitary or in fascicles of 2, asibtone .» 8. OC. macrocarpa,
1, CaszartA Lopstana, Turez. in Bull. Soc. Nat. Mose. (1858),
XXXI, Pt. I, 463, A slender tree 15-20 feet high; young branches
slender, densely tawny-pubescent. Leaves oblung or oblong-lanceolate,
shortly acuminate, the base rounded or narrowed; upper surface
glabrescent, the midrib and nerves pubescent ; lower surface sparsely
pubescent, densely so on the midrib and nerves; main nerves 8-10
pairs, curving upwards, length 2-5 in., breadth 75-225 in.; petioles
‘2-3 in., tomentose, slender. Flowers few together in the leaf-axils, on
very short pedicels, glabrescent. Calyz-teeth triangular, acute, reflexed.
Staminodes villous, rather shorter than the stamens. Fruit almost
sessile and nearly globose, orange-yellow when ripe, ‘5 in. long and °4
in, in diam. Clarke in Hook. fil. Flor. Br. Ind. II, 594,
16 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
Srvcaporr; Lobb. Porax; King’s Collector 2377, 2631, 10731;
Wray ; 829, 2594; Scortechini (without number), Distris. Tenasserim.
2, CASEARIA GREWI#FOLIA, Vent. Choix. 48. A small tree; young
branches 4-angled, densely rusty-tomentose. Leaves oblong, acute, not
at all or very little narrowed to the cordate sub-truncate base, the
edges entire or minutely crenulate; upper surface shining, almost
glabrous, the lower surface pellucid-punctate when fresh, tomentose
on the midrib and 10-14 pairs of little-curved ascending main nerves,
otherwise sparsely pubescent; length 4°25-6 in., breadth 1°25-2°35 in.,
petiole ‘2 in. Glomeruli many-flowered, chiefly from the axils of fallen
leaves; the flower-pedicels slender, ‘25 in. long, minutely pubescent,
‘ articulated at the base. Flower-bud °15 in. long; sepals minutely
adpressed-pubescent, adnate at the base to the short staminal column.
Stamens 8, the glabrous filaments alternating with the 8 pubescent
staminodes. Ovary pytamidal, hairy at its junction with the style.
Fruit compressed-ellipsoid, nearly 1 in. long, glabrous, boldly ridged
when dry. DC. Prodr. Il,51; Mig. ‘Flor, Ind, Bat. 1, Pi. e706:
Clarke in Flor. Br. Ind. IT, 594. C. variabilis, Blume Mus. Bot. Lugd.
Bat. I, 252. C. subcuneata, Miq. Flor. Ind. Bat. I, Pt. I, 706. C. cinerea,
Turez. in Bull. Soc. Nat. Mosc. (1858), XX XI, Pt, I, 462.
' Manacca; Maingay (Kew Distrib.) 659, 661. Purax; Scortechini ;
2003. Disrris. Java, Bali, and other Islands of the Malay Archipelago.
8. Castaria ALBICANS, Wall. Cat..7197. A shrub 3 or 4 feet high ;
young branches slender, very pale, almost white when dry, glabrous.
Leaves thinly membranous, pale brown when dry and much pellucid-
punctate, more or less broadly elliptic, shortly acuminate; the base
rounded or very slightly narrowed, the edges entire, both surfaces gla-
brous and finely reticulate when dry; main nerves 8 or 9 pairs, curved,
ascending; length 4-7°5 in., breadth 2°25-4 in., petiole *25 in. Glome-
ruli small, few-flowered, axillary ; flower-buds obovoid, almost sessile,
surrounded by numerons acute bracteoles. Calyx-teeth 4, broadly tri-
angular, glabrous. Fruit broadly ovoid, compressed, acute, subglabrous,
1 in. long and ‘65 in. in its broad diameter. ‘‘ Samydee,” Wall. Cat.
7432. | 3
Penang; Wallich. Porax; King’s Collector 3634.
The above description is drawn up from the two sheets of Wall. Cat. above
quoted and of King’s Collector 3634. One of Wallich’s specimens 9197 has fruit
in a pocket detached from the twig, the others have no fruit. The plant here
named C. albicans is not, however, that described under the same name by Mr.
C. B. Clarke in the Flora of British India. Mr. Clarke’s plant is 660 of Maingay’s
Herbarium, and is referred by me (along with 660/2) to C. macrocarpa, Clarke.
4. CASEARIA ANDAMANICA, new species, <A tree 20-40 feet high ;
young branches pale brown, glabrous. Leaves oblong-elliptic, acute,
1898.] G. King— Materials for a Flora of the Malayan Peninsula, 17
slightly narrowed and somewhat oblique at the very base ; both surfaces
glabrous, minutely reticulate when dry; main nerves 10-12 pairs,
spreading, curving upwards, slightly prominent beneath ; length 6-9 in.,
breadth 2°75-3°5 in., petiole *5-"75 in. Glomeruli axillary, large and
many-flowered ; buds sub-globular, glabrous ; pedicels about °25 in. long
glabrous, the bracteoles minute. Calyz-segments ovate, glabrous;
staminal tube wide, nearly glabrous, adherent below to the sepals;
filaments 10, as long as the tube and as the alternating villous stami-
nodes, Ovary elongate-pyramidal, 3-angled, stigma capitate; fruit un-
known.
Anpaman IsLanps ; King’s Collectors.
5. CASEARIA ESCULENTA, Roxb, Flor. Ind. TI, 422. A shrub or
small tree as high as 20 or 30 feet; young branches pale, striate when
dry, glabrous. Leaves thinly coriaceous, elliptic-oblong to elliptic-
lanceolate, acute at the apex and acute or obliquely rounded at the
base, the edges entire; both surfaces reticulate ; main nerves 6-8 pairs,
spreading, not prominent; length 3-7 in., breadth 1°5-2°25 in,, petiole
25 in. Glomeruli axillary, many-flowered ; buds and pedicels glabrous,
the latter ‘2-25 in. long; bracteoles very short, glabrous. Calyx-teeth
4, broadly ovate, concave. Scamens 8, alternating with the staminodes.
Ripe fruit ellipsoid to globular-ovate, glabrous, dehiscing by 2 or 5
valves, length *75 in, or more. Clarke in Flor. Br. Ind, II, 592.
O. lxvigata, Dalz, in Hook. Journ. Bot. IV, 107; Dalz. & Gibs. Bomb.
Flor. 11. ©, Championit and C. Zeylanica, Thwaites, Enum. Pl. Ceylon,
19. OC. varians, Thwaites Enum. 19 (in part).
Sincapore; Lobb, Griffith, Maingay (Kew Distrib.) 657.
Perak; Ridley 5218; Scortechini 804; King’s Collector 4699, 7001,
A widely distributed species presenting a considerable amount of variation.
Perak specimens have larger leaves than those from British India; but they appear
to have smaller fruits; for Roxburgh describes the fruit of the plant, as it grows
in the Northern Circars, as being as large as a nutmeg.
6, Cassaria Kunstiert, King n. spec. A tree 30-80 feet high ;
young branches pale, glabrous. Leaves coriaceous, narrowly oblong,
tapering slightly to each end, the edges entire; both surfaces glab-
rous and minutely reticulate; the upper shining, the lower rather
dull; main nerves 7 or 8 pairs, ascending and only slightly curved,
prominent on the lower surface; length 5-7 in., breadth 1-5-2 in.:;
petiole -25 in, stout. Glomerult mostly in the axils of fallen leaves,
many-flowered; pedicels stout, glabrous ‘35-45 in. long; buds ‘15 in,
long, blunt. Calyx 5-cleft, the segments broadly ovate, obtuse, concave,
minutely pubescent, Stamens 10, broad, sub-acute, the filaments broad,
pointed, their edges pubescent. Staminodes elliptic, their apices acute and
J.u. 3
18 G. King— Materials for a Flora of the Malayan Peninsula. [No. J,
pubescent. Ovary broadly ovate, 5-celled; the stigma capitate-discoid,
almost sessile. Fruit bright yellow, broadly ovoid or obovoid, apicu-
late, tapering also to the base, glabrous, 1°5-2 in. long and *75-1°35 in.
diam., pulpy and smooth when ripe, boldly 3-angled when dry.
Perak ; Wray 3752; King’s Collector 3694, 6936, 7118.
This resembles both C. macrocarpa and C. Clarkei in several respects, but it has
larger and more numerous flowers, and the fruit is broader, in proportion to its
length, than is the case in either of these species. The leaves are less shining
and reticulate than those of C. macrocarpa, but they closely resemble those of
C. Clarke.
7. CasbaRIACLARKEI, King. A tree; young branches thick, glabrous,
striate. Leaves coriaceous, narrowly oblong, acute at both base and
apex, both surfaces glabrous, the upper shining ; main nerves 5-7 pairs,
slightly curved, ascending, prominent beneath; length 5-10 in., breadth
1°5-2°75 in., petiole 25-4. in. long. Glomeruli very condensed, axillary,
many-flowered ; pedicels ‘25 in. long, glabrous like the buds. Ripe
fruit ovoid, bright yellow, 2 in. long and 1:25 in. in diam. C. albicans,
Clarke in Flor. Br. Ind. II, 593 (not of Wallich).
Manracca; Maingay (Kew Distrib.) 660. Sryearore; Ridley 6354. :
This species resembles C. macrocarpa Clarke, but has smaller fruit. Its leaves
are moreover larger, less shining and less reticulate, and the glomeruli are much
more numerously flowered.
8. CasfaRIA macrocarPa, Clarke in Hook. fil. Flor. Br. Ind. IT,
593. A small tree; young branches reddish, glabrous. Leaves narrowly
lanceolate, acute or acuminate, the base narrowed, the edges entire;
both surfaces glabrous, shining, minute, reticulate when dry; main nerves
about 5 or 6 pairs, curved, ascending, length 4-6 in., breadth 1 to 1°5 in.,
petiole *3-4in. long, Glomeruli axillary, one-or few-flowered ; pedicels
about *15 in, long, glabrous, the buds glabrous outside, reddish ; calyx-
teeth 5, oblong, blunt, minutely velvety inside. Fruit obovoid or
ellipsoid, compressed, 1°5-1°75 in. long by ‘65 in. broad; seeds obovoid,
compressed.
Pananc; Maingay 660/2; Curtis 229 and 960,
2. Osmetia, Thwaites.
Trees. Leaves alternate, petioled, ovate or oblong-lanceolate, entire
or obscurely serrate, epunctate; stipules minute, deciduous. lowers
small, very nearly sessile, in long simple or panicled racemes. Calyx
inferior, divided nearly to the base; lobes 4 or 5, rounded, imbricate.
Petals 0, Stamens 8 or 10, half alternating with as many 2-lobed hairy
scales and half inserted in the notches of those scales. Ovary superior,
l-celled ; styles 3, short, with capitellate or bifid stigmas; ovules few ;
1898.} G. King— Materials for a Flora of the Malayan Peninsula. 19
placentas 3, parietal. Capsule subglobose, 3-valved. Seeds few,
subglobose, with a red fleshy aril. Duisrris. Species 6; Malaya, Philip-
pines, Ceylon.
Osmexta Marneayi, King n. spec.. A diceceous tree 20-60 feet high ;
young branches slender, minutely tawny-tomentose. Leaves membran-
ous, oblong or elliptic-oblong to elliptic, shortly acuminate, the base
rounded or slightly narrowed, the edges entire or very obscurely
crenate; the upper surface glabrous, the lower covered with minute
yellowish pubescence especially on the nerves and midrib; main nerves
6-10 pairs, curved, spreading, interarching within the edge; length
4—6 in., breadth 1°75 to 3 in., petiole ‘6-75 in. Male panicles slender,
terminal, several times as long as the leaves; flowers ‘1 in. in diam.
or less, on short pedicels bracteate at the base, in slightly distant small
glomeruli. Sepals 4, membranous, imbricate, rotund, pubescent, concave.
Petals 0. Stamens 8 in two rows, one row with longer filaments
alternating with the row opposite the broad villous glands. Panicles
of female flowers axillary, shorter than the leaves, slightly longer in
fruit. Calyx as in the male, but the segments smaller. Stamens sub-
equal, shorter than the calyx, the filaments very short, glands and
ovary densely hairy; the latter sub-globular, tomentose, crowned by 3
short distant bifid glabrous stigmas, l-celled ; ovules 3, erect. Capsule
‘5-65 in. long, 3-ridged, dehiscing by 3 valves.
MataccaA; Maingay (Kew Distrib.) 1448. Perak; Scortechini
158, 191, 623. King’s Collector 741, 1240, 2339, 4259, 4096, 5667, 7660,
7045, 10017, 10981; Wray 3665. Pauanc; Ridley 2654, Srncarore ;
King, Ridley 3804, 1904, .
This appears to be a very common tree in Perak; for there is large suite of
specimens of it in the Calcutta Herbarium numbering about 150 sheets, The
various gatherings vary somewhat as to the amount of pubescence and number of
nerves on the leaves, as also in the length of the panicles; but I cannot make
out more than one species. Maingay’s specimen (Kew Distrib.) 1439 looks as if
it might be different. There is only a single sheet of it at Calcutta, and no flower
remains on its panicles. Beccari’s Sumatra plant 928 may possibly belong to still
another species. All the species have the facies of Antidesma; the capsular fruit
when present however at once distinguishes then from that genus.
3. Homatium, Jacq.
Shrubs or trees. Leaves alternate, crenate or subentire, petioled or
sessile, rarely punctulate. Flowers hairy, small, in slender axillary and
sub-terminal simple or panicled racemes; bract at the base of the pedicel
often prominent but caducous. Calya-tube funnel-shaped or cylindric,
adnate to the base of the ovary ; lobes 5-10, narrow, persistent, Petals
o-10, inserted in the throat of the calyx, linear-oblong, persistent. Disc
20 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1
tomentose. Stamens solitary or in fascicles of 2-7, opposite the petals,
alternating with large glands. Ovary half-superior, 1-celled ; styles 2-5,
filiform, stigmas capitellate ; ovules many or several ; placentas parietal,
extending only down the upper free portion of the ovary. Capsule half-
superior, coriaceous, 2—5-valved at the apex. Seeds few, angular or
oblong. Duistris.—Species 50, scattered over the hot regions of nearly
the whole globe.
Flowers never more than ‘35 in. in diam. :—
Stamens 1 opposite each petal; leaves glabrous So ke
Stamens 2 in front of each petal; calyx-tube funnel—
+ shaped :—
Leaves glabrous, glaucous beneath; flowers 6-
merous eee vee oop 2 HH. Kunstierz.
Leaves glabrous except on He midrib beneath, not
glaucous; flowers 6- or 7-merous sie 3.
Leaves sparsely pubescent on both surfaces, the
midribs tomentose ; flowers 10-merous we A
Stamens 4 in front of each petal; flowers 6-merous,
calyx-tube cylindric, expanding very slightly at the
mouth see ove aa «. OO. H, frutescens,
Flowers ‘6—75 in. in diam. :—~
Stamens 4 in front of each petal; calyx-teeth 5,
much smaller than the petals : 6.
Stamens 7-9 opposite or sub-opposite to cai ssoulare
calyx-teeth 7-9, larger than the petals and accrescent 7.
H. longifoliwm,
H, propinquum.
H. Griffithianwm.
H. undulatum.
H. grandiflorum.
1, Homanium LonGirotiuM, Benth. in Journ. Linn. Soc. IV, 35
A tree 30-60 feet high; young branches slender,
lenticellate, almost
glabrous.
Leaves coriaceous, oblong or oblong-lanceolate, acute or
shortly and bluntly acuminate, the base narrowed; the edges entire,
sometimes slightly undulate; both surfaces quite glabrous, the lower
darkest when dry; main nerves 7-9 pairs, spreading, curved, faint;
length 3-4°5, in., breadth 1'5-1°75 in.; petiole ‘3 in., stout. Racemes
slender, axillary, solitary, rarely branched, 4-7 in. long, covered with
minute white tomentum. Flowers‘15 in, across, almost sessile, in
fascicles of 3 or 4, 6-fid. Calyw-tube narrow, its segments lanceolate like
the petals but broader Stamens one opposite to each petal alternating
with yellow-glands. Fruit unknown. Clarke in Flor. Br. Ind. II,
596. Blackwellia macrostachya, Turcz. in Bull. Soc. Imp. Mose. 1863
(Vol. XXXVI), 610.
Penang; Phillips, Curtis 201. Maracca; Maingay (Kew Distrib.)
665; Derry 994, Prax; King’s Collector 4444, 7855, 10230, 10763;
Scortechini 487, 2036. SeLancor; Scortechini 1910.
2. Homatium KounstTLERri, n. spec. King. A tree 30-40 feet high ;
young branches glabrous, rather slender, smooth, glaucous. Leaves
1898.] G. King — Materials for a Flora of the Malayan Peninsula, 21
coriaceous, oblong-elliptic, shortly acuminate, narrowed or rounded at
the base, sometimes oblique ; the edges remotely crenate, revolute when
dry ; both surfaces glabrous, the lower glaucous; main nerves 8 or 9
pairs, faint, curving upwards; length 4°5—7 in., breadth 2°25-2°75 in. ;
petiole -2—3 in., thick. Rucemes solitary, axillary, slightly Jonger than
the leayes, bearing rather distant glomeruli of 3.or 4 flowers each, the
rachis densely and minutely tomentose. Flowers *3 in. in diam., the
pedicels about ‘1 in. long. Calya-tube short, widely funnel-shaped ;
teeth 6, oblanceolate, obtuse, spreading, minutely tomentose externally.
Petals 6, broadly lanceolate, sub-acute, equal in length to the calyx-
teeth, villous on the inner surface. Stamens 2 in front of each petal;
the filaments glabrous rising from a small bulb, Ovary hairy. Styles
5, short, sub-erect.
Perak ; King’s Collector 4286, 7109.
3. Homatium Propinguum, C. B, Clarke in Flor. Br. Ind. II, 597.
A tree 60-80 or even 120, feet in height; young branches pale brown
when dry, puberulous. eaves coriaceous, elliptic to elliptic-obovate,
obtuse, or shortly acuminate, more or less narrowed and sometimes
slightly oblique at the base, the edges undulate- or undulate-crenate ;
both surfaces glabrous, the midrib alone sometimes puberulous beneath ;
main nerves 9-1] pairs, curved, spreading, prominent beneath when dry ;
length 4-7 in., breadth 2°25-3°5 in.; petiole ‘25-35 in. Racemes 4-12
in, long, clustered and sometimes panicled at the apices of the branches,
covered with soft pale tomentum., lowers *2in, in diam., 6 or 7-fid, in
close clusters and on pedicels *15 in. long. Calyx-tube funnel-shaped ;
its segments longer than the petals, subspathulate. Stamens twice as
many as the petals, all bearing anthers. Fruit unknown, H. longifolium,
(in part) Benth. in Journ. Linn. Soc. IV, 35. Blackwellia propinqua,
Wall. Cat. 4898. B. spiralis, Wall. Cat. 4897A.
PenanG; Porter, Curtis 1592. Purak; King’s Collector 3748, 3935,
4883, 7936. Matuacca; Grifith; Maingay (Kew Distrib.) 664.
4, Homantom Grirriratanum, Kurz in Journ. As. Soc. Bengl, XL,
Pt, Il, (for 1877), 57. A tree 30-40 feet high, the young parts softly
tawny-pubescent. eaves membranous, obovate-oblong, shortly and
bluntly apiculate, slightly narrowed to the rounded or minutely sub-
cordate base, the edges subentire to coarsely crenate; both surfaces
sparsely pubescent, tomentose or densely pubescent on the midrib
and 7-9 pairs of spreading little curved not prominent main nerves;
length 2°75-5 in., breadth 1°65-2'5 in.; petiole ‘2-"4 in., pubescent,
Racemes solitary, one and a half times as long as the leaves, softly
tawny-tomentose; the glomeruli few-flowered, not crowded together.
Flowers °35 in. in diam., densely villous in all parts. Oalya-tube conical
22 G. King— Materials for a Flora of the Malayan Peninsula. [No, 1,
expanding into a wide mouth ; the teeth 10, linear. Petals oblanceolate
or spathulate, broader and longer than the sepals. Stamens 2 in front of
each petal, glabrous. Ovary short, crowned by 5 short slightly spread-
ing styles, glabrous towards the apex. Fruit unknown. Kurz For,
Flora Burma, II, 531; Clarke in Flor. Br. Ind. II, 597. H. feetidum,
Benth. in Journ. Linn. Soc. IV, 37 (in part). Blackwellia dasyantha,
Turez, Bull. Soc. Imp. Mosc. Vol. XXXVI (J863), 610. Blackwellia
spec., Griff. Notule IV, 584.
KepaH; Curtis 2506. Trane; King’s Collector 1393. Disrrtre.
Tenasserim.
Kurz describes the calyx-lobes and petals as 6 each, and in that he is quite
wrong; for dissection of Griffith’s specimen (which is the type of the Species) shows
10 of each. Although the structure of the flowers is the same in the gatherings
of this plant from Burma, Kedah and Trang, there is considerable difference as to
the amount of hair on the leaves and also as to theiredges. The Burmese specimens
are rather obscurely crenate except when very young; the Kedah plant has its adult
leaves boldly crenate and minutely pubescent on th@lower surface and glabrescent
on the upper (except the midrib and main nerves) ; while the Trang specimens are
glabrescent on both surfaces, with the exception of the nerves and midrib. The
flowers of the Trang plant are moreover slightly larger than those either from
Kedah or Tenasserim.
5. HomatiuM FRUTESCENS, King. A tree 20-30 feet high ; young
branches slender, pale when dry, glabrous, striate. Leaves thinly
coriaceous, elliptic-oblong to elliptic, shortly acuminate, tapering slightly
to the base; main nerves 7 or8 pairs, curving upwards, prominent beneath;
both surfaces glabrous, minutely reticulate, the lower pale-brown and
the upper olivaceous when dry; length 3-5 in, breadth 1:75-2-25
(rarely 2°75 in.), petiole ‘lin. acemes axillary, sometimes with 2 or 3
branches, about as long as the leaves or sometimes longer, the rachis
puberulous ; the flowers numerous but not crowded, in pairs or solitary,
about ‘25in. long, and ‘] in. or*l5 diam. at the mouth; the pedicel
under ‘lin. in length. Calyx narrowly cylindric-conic, boldly ridged;
teeth 6, lanceolate, acute, erect. Petals larger than the petals, oblong-
lanceolate, pubescent on both surfaces but especially on the upper.
Stamens 4 opposite each petal and slightly exceeding it in length,
glabrous. Ovary hairy, elongate; styles 5, as long as the stamens,
cylindric, erect, Fruit unknown. Blackwellia longiflora, Mig. Flor.
Ind. Bat. I, Pt. I, 715. B. caryophyllacea, Zoll. et Moritz. Syst.
Verz. 33; Miq. Flor. Ind. Bat. 1, c. 715; Homalium caryophyllacewm,
Benth. in Journ. Linn. Soc. IV, 38. Cordylanthus frutescens, Blume
Mus. Bot. Ludg, Bat. LH, 27, fig. II. Homalium cordylanthus, Benth.
in Journ, Linn. Soc. IV, 38.
Setancor ; Jiidley 1902. Jonore; Ridley 4050, 4182. Pura;
King’s Collector, 776, 777, 784, 5246, 7040, 10471, 10237, 10102, 10996.
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 23
The flower of this is excellently figured by Blume in his Mus. Bot. as quoted
above. Flowers however of a slightly different shape are found in some specimens.
In these the calyx-tube is funnel-shaped and less cylindric, and the petals are more
spreading. These are associated usually-with larger leaves, more broadly elliptic
in shape; and at one time I was of opinion that they might belong to a different
species. But the examination of nearly a hundred specimens of the two forms as
collected in Perak has convinced that they are not specifically separable. Blume
describes the plant as a shrub, but in Perak it is always a small tree. Miquel’s
Homalium obovale from Sumatra (Flor. Ind. Bat. Suppl. 334) comes very near
this, and perhaps is not distinct.
6. HoMALIUM UNDULATUM, n. spec. King. A tree 40-60 feet high ;
young branches slender, pale-brown, minutely lenticellate. Leaves
thinly coriaceous, elliptic, shortly and bluntly acuminate, slightly
narrowed and sometimes unequal at the base, the edges undulate-
crenate ; both surfaces glabrous and minutely reticulate when dry, the
upper shining, the lower dull; length 3-4 in., breadth 1°75-2 in., petiole
‘2-25 in. Inflorescence consisting of a terminal panicle twice as long
as the leaves and of afew solitary racemes from the upper leaf-axils,
many-flowered, the ‘rachises covered by minute rather sparse pale
tomentum. lowers ‘6 or*7 in. in diam., lax; their pedicels slender,
tomentose, ‘4. in. long. Calya-tube widely funnel-shaped, very slightly
ridged, minutely tomentose; teeth 5, lanceolate, tomentose on both
surfaces like the tube. Petals 5, much longer than the sepals, obovoid,
blunt, clawed at the base, reticulate, tomentose. Stamens shorter than
the sepals and much shorter than the petals, 4 in front of each petal,
glabrous. Ovary hairy, crowned by 3 narrowly conical spreading hairy
styles, Fruit unknown.
Purak ; [King’s Collector 7064, 8184.
7. HoMALIUM GRANDIFLORUM, Benth. in Journ. Linn. Soc. IV, 37,
A tree 30-40 feet high ; young branches rather stout, glabrous. Leaves
coriaceous, elliptic to oblong, acute or shortly acuminate, narrowed near
the petiole, the edges revolute and obscurely crenulate; both surfaces
minutely reticulate when dry, and the upper very shining, the lower
somewhat duller; main nerves 9-12 pairs, ascending, only slightly
curved; length 3°5-6°5 in., breadth 1°75-3°25 im.; petiole ‘2-'3 in.,
stout. Injlorescence consisting of terminal few-branched panicles and
of solitary axillary racemes with tomentose rachises. Flowers numerous
but not crowded, solitary, uot in glomerul, °75 in. in diam. ; the pedicels
‘2 in. long, tomentose. Calywx-tube short, hemispheric ; teeth 7-9, spathu-
late-oblong, spreading, enlarged often flowering. Petals lanceolate,
shorter than the calyx-teeth; the glands alternating with the petals,
large. Stamens 7-12 opposite each petal; the filaments subulate, sparsely
pilose. Ovary free or nearly so, 6-7-ridged, tomentose, conical; styles
24 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
6 or 7, short, glabrous, erect. Fruit unknown. Clarke in Flor. Br. Ind.
II, 598. Pierrea dictyoneura, Hancein Trimen’s Journ. Bot. for 1877, 339-
Matacca; Griffith. Perak; Scortechini, Sincaporn; Ridley 6527.
Gulf of Siam; Hance. Distris. Tenasserim.
An examination of flowers of his Pierrea dictyoneuwra, sent to me by the late
Dr. Hance, proves that that plant is a species of Homaliwm with larger flowers and
more stamens than usual.
Order LIT. CUCURBITACE A,
Climbing herbs or shrubs; tendrils solitary, lateral, spiral, simple
or divided. Leaves alternate, petioled, frequently cordate, simple, lobed
or pedately divided. Flowers moncecious or dicecious, yellow or white,
racemed and solitary, less commonly panicled. Calyx-tube wholly
adnate to the ovary; limb rotate, campanulate, or tubular; lobes 5
(rarely 3), imbricate. Petals 5, inserted on the calyx-limb, united in a
tube, or nearly or quite free, sometimes fimbriated at the margin, valvate
or involute in the bud. Stamens inserted at the mouth or about the
middle or at the base of the calyx-tube, usually 3 (sometimes 5 or 2),
anthers free or united into a tube, one usually 1-celled and the other
two 2-celled, cells straight or flexuose or conduplicate, the connective
sometimes crested or produced. Ovary inferior, usually 3-carpellary ;
style 1 with 3 stigmas, more rarely styles 2-3-4; placentas usually 3,
vertical, in double lines, the edges of the carpellary leaves being often
turned in so far that the ovary (even before fertilization) is spuriously
3-celled; ovules usually many, horizontal, rarely pendulous, sometimes
few and pendulous from near the top of the ovary. Fruit generally
berried or fleshy, indehiscent or dehiscing by valves or by a circum-
scissile lid, often 1-celled, the seeds being often packed in pulp or fibre.
Seeds usually many, often compressed, horizontal, pendulous, fre-
quently corrugated or sub-spinose on the margins, albumen 0, Dzsrris.
Species 600, in the warmer parts of the whole globe especially in the
tropics.
Tribe I, CucuMERINEM. Ovules horizontal ; female flowers
solitary, never panicled ; stamens 3, rarely 2 or 5, free
or variously connate ; cells of anthers straight, curved or
flexnous; ovary bearing 3 (rarely 2 or 5) placentas :—
Anther-cells conduplicate or sigmoid; corolla rotate or
campanulate, divided to the base :—
Petals cirrhiferous or fimbriate :—
Seeds only 6, perfect, very large Vr Hopesonia.
Seeds numerous, testa not fibrous oa .. 2. TRICHOSANTHES.
Petals entire :—
Calyx-tube of male flower elongate; the anthers in-
cluded in the tube, cohering, sessile or subsessile,
tendrils usually simple des vee .. 3 GYMNOPETALUM,
—_
1898.] G, King— Materials for a Flora of the Maluyan Peninsula. 29
Calyx-tube of male flower short; anthers usually
exserted or sub-exserted, free or slightly coherent :—
Stamens inserted near the mouth of the calyx-
tube ; the anthers cohering slightly :—
Male flowers in racemes or clusters; fruit more
or less fibrous, dehiscing by a _ transverse
lid near the apex; tendrils 2—5-fid .. 4. Lurra.
Male and female flowers solitary; fruit not
fibrous, very fleshy, indehiscent ; tendrils 2-fid 5. BrNINcASA.
Stamens inserted below the mouth of the calyx-
tube; anthers more or less cohering; male
flowers in racemes, usually bracteate; tendrils
simple We 6. Momorpica.
Anther-cells straight, the abhdetis sass the fruit
on a slender pedicel F .. 7. MELOTHRIA.
Tribe II. GyNostEMMER. Ovules Baccarat ; nee ne
in panicles; stamens 3-5, anther-cells straight, filaments
connate near the base ; ovary 3-celled and with 3 placentas,
fruit indehiscent es: ... 8. GYNOSTEMMA.
Tribe III. Zanonrm. Ovules Bedi canis flowers in
_panicles or racemes; stamens 5, free, anthers straight ;
ovary with 3 placentas; fruit cylindric or clavate, dry,
1-celled, dehiscence circumscissile ; seeds winged :—
Calyx-lobes 3, seeds winged all round ana «» 9. ZANONIA.
Calyx-lobes 5, seeds winged at the apex only ... LO. ALSOMITRA.
1. Hopesonra, H. f. & T.
A large climber. Leaves coriaceous, 3-5-lobed, long-petioled ;
tendrils 2-3-fid. lowers large, dicecious ; males in long racemes ; bracts
oblong, entire, deciduous; females solitary. Male-calyx long-tubular,
with a shortly campanulate mouth and 5 short teeth; petals 5, connate
at the base, very long-fimbriate; stamens 3, filaments very short ;
anthers exserted, connate, linear, one l-celled, two 2-celled, cells condu-
plicate. Female calyx and corolla as in the male; ovary globose,
l-celled; style long; stigmas 3, oblong, bifid, exserted; placentas 3,
parietal, near the base of the ovary, 2-ovulate oneach side. Prutt.
large, depressed-globular, 12-grooved, flesh hard; perfect seeds usually
six, each having a smaller, commonly barren one, attached to its side,
flat-ellipsoid, with sunk veins.
Hopesonia HereRocLITA, Hook. fil. and Thoms. in Proc. Linn. Soc.
II, 257. Stem very long, often reaching 80 or 100 feet. Leaves palmate,
somewhat cordate at the base; the lobes entire, rarely slightly denti-
culate, acute; both surfaces glabrous, minutely reticulate when dry,
6-10 in, long and as broad, the petioles 2 or 3 in. long. Male racemes
about as long as the leaves; bracts solitary, oblong, acute, *o in. long,
Calyx rusty-tomentose externally, the tube 3 or 4 in. long, with a gland
J. U1. 4
26 G. King—Materials for a Flora of the Malayan Peninsula. [No. 1,
on each tooth of the mouth. Petals sub-obcordate, retuse, about 2 in.
long, covered with brown hairs and 3-nerved externally ; the inner
surface white, the margins fringed with very long spiral villous
yellow threads. Filaments clavate, anthers forming an inverted cone;
pistil 0. Female flowers often on a separate plant; their peduncles 1 or
2 in. long, occasionally in a raceme much shorter than those of the
males; stamens 0. Fruit turbinate, 4-10in. in diam., and less from base
to apex, covered with short reddish-brown tomentum. Seeds 2-3 in.
long, and 1°5-2°25 in. broad. Hook, fil. Ill. Himal. Pl. tt, 1, 2, 3;
Flor. des Serres, t. 1262, 3; Clarke in Hook, fil. Flor. Br, Ind. II, 606.
Hodgsonia macrocarpa, Cogn. in DC. Mon, Phan. III, 349. Trichosan-
thes macrocarpa, Bl. Bijdr. 935; Ser. in DC. Prodr. II, 315; Miq.
* Flor. Ind, Bat. I, Pt. II, 676. J, hexasperma, Bl. Bide: 19555" De:
Prodr. III, 315; Hassk. Pl. Jav. Rar. 192; Mig. Fl. Ind. Bat. 1. c, 678.
T. heteroclita, Roxb. Hort. Beng. 70; Fl. Ind. III, 705; Wall. Cat. 6684.
T. grandiflora, Wall. Cat. 6685 (not of Blume).
Perak; Scortechini, Wray, King’s Collector. Matacca; Maingay.
Prnanc; Wallich. Distris. Sumatra, Java, Borneo, Burma, Assam,
and the base of the Hastern Himalaya.
The earliest name of this species was Roxburgh’s (7. heteroclita), published in
the Hortus Bengalensis in 1814. Blume’s name TJ. macrocarpa dates from 1826,
2. TricHosantHes, Linn.
Scandent herbs. Leaves entire or 3-9-lobed, denticulate ; tendrils
usually 2—5-fid. lowers dicecious or occasionally moneecious, white ; male
peduncles usually in axillary pairs, one 1-flowered caducous, the other
racemose ; bracts large or small or 0; female flower solitary. Male-
calyx long-tubular; teeth 5, lanceolate, entire serrate or laciniate.
Corolla 5-fid nearly to the base, lobes long-fimbriate; stamens 3;
anthers almost included, connate (free in 7’. dioica), long-linear, one
l-celled, two 2-celled, cells conduplicate, Female calyx and corolla
as in the male. Ovary inferior, at the base of the calyx-tube,
1-celled; style filiform, 3-or 6-fid at the apex; placentas 3, parietal ; ovules
very many, horizontal, half-pendulous. J’rwit lanceolate or globose,
smooth, acute or obtuse at the apex. Seeds many, horizontal, packed
in pulp, compressed, ellipsoid, sometimes angular on the margin.
Disrris. Species 38, in South-Hast Asia, extending through Malaya to
North Austraiia, also through China to Japan.
Leaves, although often deeply lobed, always simple :—
Male inflorescence ebracteate ... a wees oL.) DP. cucumerina.
Male inflorescence bracteate :—
Bracts linear-oblong or narrowly rhomboid, quite
entire . eve ai oe we 27, Hearn
1898.] G. King— Materials for a Flora of the Maluyan Peninsula. 27
Bracts small ovate-lanceolate, caudate-acuminate,
obscurely dentate, leaves quite smooth and glab-
rous on both surfaces ex . 38. TT. tricuspidata.
Bracts very large, ovate to sub- oeincalne, ieente
incised or simply serrate; calyx-teeth lanceolate;
fruit usually globular, rarely ovate... - 4 TT. palmata.
Bracts rather large, oblong-ovate, laciniate; calyx-
teeth spreading, entire; fruit always ellipsoid ... 5. TT, Wallichiana.
Leaves usually trifoliolate, but sometimes simple on the.
same or different plants; leaflets membranous, unequal,
much acuminate, the edges undulate-dentate: bracts of
male inflorescence small, oblong, their edges with a few
long distant teeth bu ow 6. PF. Wawrei.
Leaves always trifoliolate ; tail ote coriaceous, hah bet and
bluntly apiculate or subacute, the edges quite entire ;
bracts of male inflorescence deltoid, their edges pectinate 7. T. celebtca.
1. TRiIcHOSANTHES cUCUMERINA, Linn. Sp, Pl. Ed.1,1008. Stems
slender, angled, puberulous. Leaves membranous, orbicular-reniform to
broadly ovate in general outline; the edges remotely denticulate, often
more or less deeply 3-5 or 7-lobed; the lobes broad with acute but not
acuminate apices and often sinuate margins, the base deeply cordate,
the sinus often sub-rectangular ; upper surface sparsely pubescent or
sub-glabrous, the lower deciduously pubescent at first, ultimately sub-
scabrid; length 2-4 in. and breadth about the same; petiole 1-2 in.,
pubescent ; tendrils slender, sulcate, puberulous, 2—3-fid. Male peduncles
rather longer than the leaves, pubescent, bearing a few flowers towards
the apex, ebracteate. Flowers ‘5-1 in. in diam. at the mouth; the
tube 2 or 3 times longer, cylindric, a solitary male flower sometimes
from the same axil as the peduncle. Female flower on a peduncle ‘5 in.
long, fruit 1-3 in. long, narrowly ovoid, the apex conical, red when ripe;
seeds half ellipsoid, compressed, corrugated. Lour. Flor. Cochchin,
588; Ser.in DC. Prodr. IIT, 315; Roxb. Hort. Beng. 70; Roxb. Flor.
Ind. III, 720; Wall. Cat. 6690 A, B, C, D, F; Blume Bijd. 933;
Dalz, & Gibs. Bomb. Flor. 102; Mig. Flor. Ind. Bat. I, Pt. I, 676;
Naud. in Ann. Sc. Nat. Ser. 4, XVIII, 191; Kurzin Journ. As. Soe.
1877, Pt. II, 98; W. & A. Prodr. 350; Miq. Flor. Ind. Bat. I, Pt.
I, 676; Thwaites Enum. Pl. Ceyl. 126; Benth. Flor. Austral. III, 314;
Clarke in Hook, fil. Flor. Br, Ind. IT, 609; Cogn. in DC. Mon. Phan.
III, 358, T. laciniosa, Klein in Herb. Rottler. 1’. pilosa, Wall. Cat,
6691. Bryonia wmbellata, Wall. Cat. 6700 D. Cucumis Missionis, Wall.
Cat. 6728.
Perak; King’s Collector 5622, Disrris. Malayan Archipelago;
British India.
The Perak plant has larger flowers than any Indian specimen, and I refer it to
T, cucumerina with some hesitation,
28 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
2. 2? TricnosantHrs Hearni, F. Mull. in Benth. Flor. Austral. III,
315. Stem slender, glabrous, sulcate. Leaves membranous, broadly
ovate, the apex acute or acuminate, the base deeply and widely cordate ;
the edges rather remotely denticulate, sometimes remotely sinuate-lobed ;
the upper surface glabrous and shining, the lower densely and minutely
pubescent, the nerves, subglabrous and stout; length 4-8 in., breadth
3°5-6 in., petiole 1-1°75 in, Male peduncles often in pairs, axillary,
longer than the leaves, slender, puberulous, angled ; floriferous in the
upper half; bracts linear-oblong, entire, about °25 in. long; several of
the lower ones narrowly rhomboid, all pubescent, flowers under 1] in. in
diam. at the mouth; calyx-tube cylindric, the mouth narrowly infundi-
buliform, puberulous; the lobes of the mouth narrow, acute. Female
flower and fruit unknown.
Anpamans; Kurz, King’s Collector.
The late Mr. Kurz collected a single specimen of this on Rutland island (in the
Andaman group) many years ago, and he referred it in the Calcutta Herbarium to
Trichosanthes reniformis, Miq. He also suggested for it the name Trichosanthes
herpetospermum,—-a name which he never published. In 1890 several specimens of
the plant were collected near Port Mowat, on the Sonth Andaman. Specimens of
this second gathering, as well as of Kurz’s original gathering, were sent by me to Mr.
C. B. Clarke who found that they closely resemble, and are probably identical with,
T. Hearni, an Australian species named by the late Baron von Miiller, and of which
an imperfect description was published by Mr. Bentham in his Flora of Australia.
And to this species I now doubtfully refer both the Andaman gatherings.
3. TRICHOSANTHES TRICUSPIDATA, Lour, Flor. Cochinchin. IJ, 723.
Stem stout, sulcate, smooth, glabrous. Leaves thinly coriaceous, ovate-
sub-triangular, with from 38-5 stout triangular broad, acute or acumi-
nate, spreading lobes, the base broadly cordate, the edges remctely and
minutely denticulate or subentire, glabrous on both surfaces; length
and breadth 3-5 in.; petiole slender, 1-1‘5 in.; tendrils 3-fid. Male
peduncles floriferous for half their length, longer than the leaves, stout,
glabrous or puberulous, 10-20 flowered; bracts ‘thick, rigid, ovate-
lanceolate, caudate-acuminate, obscurely dentate or entire; calyx-tube
tapering to the base, shortly but densely tomentose, the lobes caudate-
lanceolate with a few irregular distant teeth; corolla 1°5 in. in diam.
Female flower from the same axil as the male peduncle, the tube 1°5 in.
long, the corolla °75 in. in diam. ; fruit ovoid when young, subglobular
and 2 in. in diam. when ripe, smooth, yellow with numerous small black
pustules, the pulp thick; the seeds in the very centre, broadly ovate,
compressed, *5 in. long and ‘4 in. broad. Blume Bijdr. 935; Ser. in
DC. Prodr. ITI, 315; Roem, Syn., fasc. II, 95; Mig, Flor. Ind. Bat. I,
Pt. I, 676; Cogniaux in DC. Mon. Phan. IIT, 374.
_ Penane; Curtis 1947. Perak; Scortechini 376; King’s Collector
2202, 5111; Wray 4029.
1898.| G. King— Materials for a Flora'of the Malayan Peninsula. 29
This resembles T. palmata, Roxb. and T. Wallichiana, Wight; but the lobes of
the leaves are shallower and their surfaces more glabrous and not at all scabrid,
while the bracts of the male inflorescence are much narrower and very acuminate ;
and their edges, instead of being laciniate, are entire or at most obscurely dentate.
4, TRICHOSANTHES PALMATA, Roxb. Hort. Beng. (1814) p. 70.
Stem stout, angled and sulcate, glabrous, Leaves thickly membranous,
broadly ovate or orbicular in general outline, deeply cut down to
about the middle into 3-7 lobes; the lobes acute or acuminate, their
edges entire or denticulate, the base deeply but usually narrowly
cordate; upper surface more or less scabrous or scabrid, glabrous, glan-
dular near the apex of the petiole, the lower glabrous or somewhat
pubescent, length and breadth 2-6 in. ; petiole rather slender, 1 to 3 in.
long; tendrils 2- or 3-fid. Male racemes sometimes in pairs, longer than
the leaves, few-flowered; bracts large, ovate to sub-orbicular, deeply
incised or simply serrate, glabrous or with viscid hairs; calyx-tube 1°5
in. long, tomentose or glabrous, the teeth lanceolate, the edges deeply
serrate or laciniate. Female flower solitary, on a peduncle less than
1 in. long ; fruit ovoid, pointed when young, globular when ripe, 1*5-2
in. in diam. T. palmata, Roxb. Flor. Ind. III, 704; Wall. Cat. 6688
(excl. C, F); W. & A. Prodr. 350 ; Wight Ill. t. 104, 105; Dalz. & Gibs.
Bomb. Flor. 103. 1. laciniosa, Wall. Cat. 6689 A,B. T. aspera, Heyne
in Herb. Rottler. 1. tricuspis, Mig. Flor. Ind. Bat. I, Pt. I, 679. T.
cordata, Wall, Cat. 6686 (excl. A and B). TT. anguina, Wall. Cat. 6687
(F partly) ; Voigt Hort. Bot. Sub. 58. TT. bracteata, Kurz in Journ.
As. Soc. Beng. 1877, Pt. II, 99; Cogn. in DC. Mon. Phan. III, 375,
T. pubera, Blume Bijdr. 936; Ser. in DC. Prodr. III, 315; Roem. Syn,
fase. II, 95; Mig. Flor. Ind. Bat. I, Pt. I, 675. Cucurbita Melopepo,
Wall. Cat. 6725. Involucraria Wallichit, Seringe in DC. Prodr. III,
318. Bryonia palmata, Wall. Cat. 6711 F.
Perak; Wray 2181, 2371, 2478, 3049 ; King’s Collector 1848, 4983,
10579. Anpamans ; King’s Collector.
The fruit when ripe is usually globular ; but there are specimens in the Calcutta
Herbarium which have oval fruit like T. Wallichiana, Wight, and at the same time
the scabrid leaves and laciniate calyx lobes which are supposed to be characteristic
of this species.
5. TrichosanrHes Watiicuiana, Wight in Ann. and Mag. Nat.
Hist. VIII, 70. Stem robust, angled and sulcate, glabrous. Leaves
membranous, sub-orbicular in general outline, divided half-way down or
more into 3-5 oblong or triangular acute lobes ; the lobes sparsely denti-
culate, the lower on each side sometimes lobulate, the base deeply
and widely cordate; both surfaces glabrous and usually smooth, the
upper sometimes slightly scabrid and with a few glands near the apex
of the petiole; length and breadth 3-7 inches, tendrils 3-fid. Male
30 G. King— Materials for a Flora of the Malayan Peninsula. {[No. 1,
peduncles at first shorter than the leaves, elongating with age; bracts
oblong-ovate, their edges laciniate; calyx-tube glabrous, the teeth
spreading, entire; petals obovate, laciniate. Female flower solitary ;
fruit ellipsoid, obtuse, smooth, 2-4 in. long. Cogn. in DC. Mon. Phan,
III, 368. Involucraria Wallichiana, Ser. in Mem. Hist. Nat. Geneve III,
Pt. I, 25 t. 5; in DC: Prodr. III, 318; Roem. Syn. fasc. 2, Speees,
Trichosanthes multiloba, Clarke in Hook. fil. Flor. Br. Ind. IT, 607 (not of
Mig.). TT. grandibracteata, Kurz in Journ. As. Soc. Beng. XLVI, 98, 99.
PaHaneG ; Ridley 244. Sincarore; Ridley 296, 446, 4762. Prnane ;
Curtis 2004. Perak; Scortechint 508 and 600. .
A species scarcely separable from T. pubera, Bl., of which in my opinion it
would better be treated as a variety.
6. TricHosantHes Wawr2zt, Cogn. in DC. Mon. Phan. ITI, 384,
Stem slender, sub-glabrous, sulcate. Leaves membranous, trifoliolate
or simple; leaflets of the trifoliolate form unequal, the middle one
oblanceolate, the lateral ones angularly auriculate or lobulate at the
base on the outer side, the edges of all subentire or sparsely denticulate
and the apices much acuminate, all with slender petiolules -25-—3
in, long; the lower surface reticulate when dry, glabrous, the upper
sparsely and minutely pustulate and glabrous; length of the leaflets
2°5-5 in., breadth °75-1'5 in., petiole 1:°25-2 in.; the simple form
triangular-oblong, tapering gradually to the apex, the base with 2 short
sub-horizontal lobules, length 2°5-5 in.; tendrils short, simple or bifid.
Male peduncles shorter than the leaves, very stout, sulcate, glabrous,
many-flowered ; bracts small, oblong, their edges with a few long teeth,
the flowers extending for half their length; calyx °5 in. long, narrowly
campanulate, suddenly contracted into the tube, the teeth erecto-
patent or recurved, lanceolate, acuminate, entire. Female flower with
cylindric tube slightly dilated at the apex, the teeth as in the male;
petals white, yellowish at the base, oblong, 3-nerved, the fimbriz long,
the external surface minutely papillose ; ovary, glabrous, ovoid-oblong ;
fruit subglobular when young, oblong when ripe, smooth, red with
orange or white stripes ; seeds compressed, oblong-ovoid, ‘6 in. long.
Perak; King’s Collector 2203, 4519, 4668, 5380, 5405, 10176.
Wray 2382; Scortechint. (SincApoRE; Wawra 241 in Herb. Vindob.
fide Cogniaux.)
Cogniaux describes this as having its leaves always trifoliolate. But in some
of the Perak specimens both trifoliolate and simple leaves as above described are
to be found; in a few only simple leaves, and in the majority only trifoliolate.
7. TricHosANTHES CELEBICA, Cogn. in DC. Mon. Phan. III, 385,
Stem slender, glabrous, sulcate, sometimes hairy at the nodes. Leaves
coriaceous, trifoliolate; the leaflets unequal in size, the middle one
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 31
the largest, ovate-oblong, shortly acuminate or sub-acute; the base
slightly narrowed, oblique, not cordate, the edges quite entire; both
surfaces glabrous, minutely reticulate when dry, the upper minutely
punctate ; length of leaflets 3-4'5 in., breadth 1°5-2°5 in. ; petiole 1-1°5
in. long, stout; petiolules *15 in. ; tendrils short, bifid. Male peduncle
shorter than or equal to the leaves, stout, deeply sulcate, adpressed-
pubescent, floriferous for half its length; bracts deltoid, pectinate,
tawny glandular-tomentose. Calya-tube narrowly infundibuliform,
minutely pubescent; the teeth lanceolate, acuminate, entire, erecto-
patent. Female flower unknown. Fruit ovate, smooth, 8 in. long
and 3 or 4 in. in diam. with a leathery rind, bright red with 10 yellow
stripes when ripe; seeds obliquely oblong, compressed, smooth, ‘6 in.
long and ‘4 in. broad.
Perak; Wray 2460; Scortechini; King’s Collector 4033. Srnea-
PORE; Ridley 2051, 4592, 6783; Hullett 247. Disrris. Celebes, Beccari.
3. GYMNoPETALUM, Arn.
Twining herbs, tendrils usually simple or 2-fid. Leaves petioled,
5-angular, nearly entire or deeply lobed, Flowers white (or yellow ?),
somewhat large; occasionally moncecious; male peduncles in fully
developed plants 2 from each axil, the earlier 1-flowered, the later
longer with racemes, either often suppressed; bracts on the racemes
persistent, large, lanceolate, incised or small ; females 1-flowered, usually
in separate axils. Male calyx-tube long, contracted near the mouth,
limb of 5 lanceolate segments ; petals 5, not fimbriate on the margin;
stamens 3; anthers included, connate, elongate, 1-2-celled, cells
conduplicate ; rudiments of the ovary 1 or 3, small, linear. Female
calyx and corolla as in the male; ovary oblong; style long, stigmas 3,
short linear; ovules horizontal, many ; placentas 3, long, vertical. Fruit
ovate-oblong, acute at both ends. Seeds many or few, ellipsoid, com-
pressed, margined, nearly smooth. Dusrris. Species 6; in India, China
and Malaya. |
Leaves not lobed ... cag = we Ll. G. integrifolium.
Leaves lobed :—
Leaves 3-5-lobed half way down, reniform to trian-
gular in general outline; lobes triangular acute, not
lobulate ie ada des we 2. G. Cochinchinensis.
Leaves deeply 5-lobed, the lobes lobulate-sinuate,
blunt, their general outline orbicular ... we 3 G. quinqueloculare.
1. GYMNOPETALUM INTEGRIFOLIUM, Kurz in Journ. As. Soc. Beng.
XL, 58. Creeping, only a few feet long; stem scabrid, tendrils simple
or bifid. Leaves reniform, obtuse; the margin undulate or denticulate,
not lobed; upper surface very scabrid, the lower softly tomentose,
32 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
length 1°75-2 in., breadth *8-2°25 in., petiole ‘45-65 in. Flowers mone-
cious, all solitary, axillary and bracteate, the male peduncle 1°5 in. long,
the female only ‘25 in. Calyz-tube in both sexes elongate, densely
covered with long brown hairs; the teeth 5, lanceolate; corolla white,
about 1:35 in. in diam., its lobes obovate, entire, pubescent, veined.
Fruit about ‘75 im. in diam., globular, smooth, orange-red. Kurz in
Flora for 1871, p. 295; Clarke in Hook. fil. Flor. Br. Ind. II, 612.
Cucumis integri folius, Roxb. Flor. Ind. III, 724; Wall. Cat. 6730.
Trichosanthes officinalis, Wall. Cat, 6694. T. integrifolia, Kurz in Journ.
As. Soc. Beng. XLVI, Pt. II, 99; Cogn. in DC. Mon. Phan. ITI, 386.
Perak; Wray 2167; Ridley 3107.
Kurz named this Gymnopetalium integrifolium in the Journal of the Asiatic
Society of Bengal which was issued in March 1871. His publication of it in Flora
dates only from October of the same year.
2. GymnopeTaLUM COocHINCHINENSIS, Kurz in Journ. As. Soc. Beng.
XLVI, Pt. II, 57. Stems slender, angled, slightly scabrid-hairy. Leaves
reniform to triangular in outline, 5-angled or 3-5-lobed half way
down; the lobes triangular, acute, the edges crenate-dentate and
thickened, the base deeply and widely cordate, both surfaces more or
less scabrid, length 2 to 4 in., breadth 2 to 3°5 in.; petiole scabrid-
pubescent, 1-1°5 in. long; tendrils simple or bifid. Male peduncle
longer than the leaves, the flowers racemose, or sometimes solitary ;
bracts large, incised-serrate, *5-"75 in. long; calyx-tube sub-cylindric,
villous, the mouth closed by deflexed hairs, the teeth erecto-patent ;
petals ovate-oblong, ‘5 in. long, entire or sub-crenate. Peduncle of
female flower shorter than the leaf-petiole, sparsely puberulous ; fruct
about 2 in. long. and *75 in. in diam., somewhat scabrid, 10-ribbed,
orange-red, the beak long; seeds about ‘25 in. long. Kurz in Flora
for 1871, p. 295; Clarke in Hook. fil. Flor. Br. Ind. II, 611; Cogn.
in DC. Mon. Phan. III, 391. Bryonia cochinchinensis, Lour. Flor.
Cochinch. 595; DC, Prodr. III, 305. Momordica tubiflora, Roxb.
Flor. Ind. III, 711, (not of Wallich). Zripodanthera cochinchinensis.
Roem. Synops. II, 48. Scotanthus tubiflorus, Naud. in Ann. Sc. Nat. Ser.
4, XVI. 172, t. 3. Trichosanthes cucumerina, Wall. Cat. 6690 EH.
T.? Fatoa, Ham. in Wall. Cat. 6695. Bryonia grandis, Wall. Cat. 6700
K. L. Trichosanthes costata, Bl. Bijdr. 933; Ser. in DC. Prodr. III, 314.
Panane; Ridley 2446. Kepan ; Curtis 2592, Perak; King’s Col-
lector 10563. Distris. British India, Malayan Archipelago, China.
3. GYMNOPETALUM QUINQUELOBUM, Miq. in Flor. Ind. Bat. I, Pt. I,
681. A slender annual; stem striate, with short pubescence or
glabrous. Leaves orbicular in general outline, deeply 5-lobed ; the lobes
sinuately 2- or 3-lobulate, or oblong and subentire; both surfaces
sparsely covered with thick whitish hairs with bulbous bases; length
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 33
1°5-2°25 in., breadth 1°75-3 in.; petiole *75-1 in., sparsely pubescent.
Male flowers usually solitary, rarely in racemes slightly longer than
the leaves, shortly pilose; the bracts oblong, serrate, ‘5 in. long. Calyzx-
_ teeth erect, entire or dentate; petals oblong, acute, puberulous outside.
Peduncle of female flower *5-1 in. long. Fruit brilliant scarlet when
ripe, oblong-fusiform, acutely ribbed, shortly pubescent, 1°5-2 in.
long; seeds blackish, rugulose, obscurely marginate, narrowed to the
base, *2 in. long and about ‘1 in. broad. Clarke in Hook. fil. Flor. Brit.
Ind. II, 611; Cogn. in DC. Mon. Phan. III, 392. Scotanthus Porteanus,
Naud. in Ann. Sc, Nat. Ser. V, Vol. 5, 25. Gymnopetalum hetero-
phyllum, Kurz in Trim. Journ. Bot. for 1875, p. 326.
ANDAMAN AND Nicopar Istanps; Kurz.
4, Lourra, Cav,
Climbers, large or small, pubescent or nearly glabrous; tendrils
2-5-fid. Leaves cordate, usually 5-angular or 5-lobed ; petiole without
glands at its apex. lowers yellow or white, moncecious, males and
females often from the same axil; females solitary or panicled, malés
on long or short racemes or clustered, Mate; calyx-tube turbinate,
lobes 5, triangular or lanceolate; petals 5, obovate; stamens 3, rarely
5, filaments 3, free or connate; anthers exserted, free, one l-celled, the
others 2-celled; cells sigmoid, often on the margin of the broad con-
nective. FEMALE; calyx-tube shortly produced above the ovary ; lobes
and corolla as inthe male; ovary oblong, style cylindric, stigma 3-lobed ;
ovules very many, horizontal, Frat large or small, oblong (not
spherical), smooth or angular or spinous, ultimately fibrous, not succu-
lent, 3-celled, usually circumscissile near the apex. Seeds many, oblong,
compressed, Distrip, Species 6, in the warmer regions of the Old
World and one in America.
Lurra Aleyprraca, Miller Gard. Dict. ed. VIII, ex Hook. fil. in
Oliv. Flor. Trop. Afr. I1, 530. Stem stout, many yards in length; the
young branches glabrous, angled and very deeply sulcate; tendrils
2-3-fid. Leaves large, reniform or reniform-orbicular in general outline,
palmately 5-lobed ; the lobes acute, lobulate and denticulate; both sur-
faces scabrous or scaberulous, punctate, glabrous except the pubescent
nerves on the lower surface ; length 25-6 in., breadth 3-9 in., petiole 2-2°5
in., pubescent, eglandular. Male peduncle 6 in. long; the flowers 4-12,
crowded near the summit, 1'5-2 in. in diam., their pedicels short, each
with a small ovate viscid bract, or ebracteate; petals 5, yellow with
green veins; stamens 5. Female flower solitary on a peduncle 1-3 in,
long, usually from the same axil as the male inflorescence ; fruit 5-12
in, long, sub-cyliudric, with numerous bold ridges; seeds usually black,
J. ut. 5
34 G. Kinge— Materials for a Flora of the Malayan Peninsula. [No. I,
narrowly winged, smooth or slightly tubercled. DC. Prodr. III, 3038 ;
Clarke in Hook, fil, Flor, Brit, Ind. I, 614, Cucumis aegyptiacus, Vesl.
in Alp. Pl. Aegypt. p. 199, t. 58, 59. Momordica Luffa, Linn. Spec. ed. 1,
1009. LL. pentandra, Roxb. Flor. Ind. IJ], 71/2; W.& A. Prodr 343;
Wall. Cat. 6751; Wight Ic. t. 499. DZ. racemosu, Roxb: 1. ec. 715.
I, clavata, Roxb. Hort. Beng. 104; Flor. Ind. Il], 714. LZ. acutangula,
W. & A. 1. ¢., (not of Roxb.). LZ. cylindrica, Roem. Synops. II, 63;
Naud. in Ann. Se. Nat. Ser. 4, XII, p..119; Kurz in Journ. As. Soc.
1877, Pt. II, 100; Cogn. in DC. Mon. Phan. III, 456. JL. Petola and
L. Cattu-picinna, Seringe in DC. 1.c. ‘ZL. Parvala, Wall. Cat. 6758.
LL. Gosa, hederacea and Satpatia, Wall. Cat. 6753, 6755, 6757. Bryon
chetrophylla, Wall. Cat. 6715 A.
Pprak ; King’s Collector 1020. Duistris. British India and in the
Tropics generally ; often cultivated.
The synonymy of this species occupies more than a page in Cogniaux’s excellent
Monograph of the Cucurbitacex in De Candolle’s Suites aw Prodromus, Vol. III. I have
followed Messrs. Cogniaux and C. B. Clarke in reducing here Roxburgh’s three species
I, pentandra, L. racemosa and L. clavata, but I do so with considerable hesitation,
Neither in flower nor leaf do Roxburgh’s figures of his D. clavata and L. pentandra
much resemble each other, whatever relation either of them may bear to M. Aegyp-
tiaca, Miller. The material of the Indian species in the Calcutta Herbarium is very
unsatisfactory, and I do not think the last word on them will be said until they have
been carefully cultivated side by side, and studied as they grow.
5. Bernitncasa, Savi.
A large climber, softly hairy, tendrils 2- or 3-fid, rarely simple:
Leaves cordate, reniform-orbicular, more or less 5-lobed ; petiole without
glands. lowers large, yellow, moncecious, all solitary, without bracts.
Matt; calyx-tube campanulate; lobes 5, leaf-like, serrate; petals 95;
nearly separate, obovate; stamens 3, inserted near the mouth of the
tube; anthers exserted, free, one l-celled, two 2-celled, cells sigmoid.
Frmae ; calyx and corollaas inthe male; ovary oblong, densely hairy ;
style thick, with 3 flexuose stigmas; ovules numerous, horizontal ;
placentas 3. Fruct large, fleshy, oblong, pubescent, indehiscent. Seeds
many, oblong, compressed, margined.
Bunincasa HisPipaA, Cogn. in DC. Mon. Phan. III, 513. Annual.
Leaves on long petioles, reniform-rotund; with 5-9 small lobes, all
toothed, 4-6 in, long and about the same in breadth ; petioles cylindric,
longer than the leaves, tendrils usually 3-fid. Male flowers axillary,
solitary, peduncled, yellow, 1°5 in. in diam.; teeth of the calyx ob-
tusely pinnatifid or undulate. Female flowers like the male, axillary,
solitary, peduncled ; calyx as in the male. Fruit sub-cylindric, obtuse
at the ends, smooth, hairy when young but glabrous and with a whitish
‘1898.}: G. King+~Materials for a Flora of the Maliytin Peninsula. 35
bloom when ripe, 12-18 in. long and from 8-10 in. in diam. Seeds white
with tumid margins, *5 in. long and ‘15 in. broad... Cucurbita hispida,
Thunb. Flor. Jap. (1784), p. 322; Bl. Bijdr. 931; Wall. Cat. 6723.
CO. Pepo, Lour. Flor. Coch.-Chin. p. 593; Roxb. Flor. Ind. III, 718 (not
of Linn.). Benincasa cerifera, Savi in Bibl. Ital. TX, 159; DC, Prodr.
Ill, 303; W. et Arn. Prodr. 344; Mig. Flor. Ind. Bat. I, Pt. I; 665;
Clarke in Hook. fil. Flor. Br. Ind. IT, 616. Cucurbita villosa, Bl. Bijdr.
931; DC. Prodr. IfT, 317. C. farinosa, Bl. Bijdr. 931. Cucurbita alba,
Roxb. in E. Ind. Comp. Mus., tab. 457 (ex W, et Arn.). Gymnopetalum
septemlobum, Miq. Flor, Ind. Bat. I, Pt. I, 679. . Lagenaria dasystemon,
Franch. et Sav. Enum. Pl. Jap. I, 173:
Nicopar AND ANDAMAN IsLANDS; cultivated, King’s Collectors. Dts-
Tris.—Malayan Archipelago, Australasia, China, Philippines, British
India; cultivated.
6. Momorpica, Linn.
Climbing by simple tendrils. Leaves cordate, petioled, undivided
in the Indian species. Flowers yellow or white, moncecious or dicecious,
females solitary, peduncled; males solitary or racemed, bracteate or not.
Mate; calyx-tube short, campanulate with 2-3 basal oblong incurved
scales (ex Hook. f.) lobes 5; corolla 5-partite nearly to the base; stamens
3; filaments short; anthers at length free, one l-celled, one or two
2-celled, cells conduplicate or horse-shoe-shaped.. Famatn; calyx and
corolla as in the male; ovary oblong; style long, stigmas 3; ovules
very many; placentas 3, horizontal. Fruit oblong or spherical, rough or
smooth, indehiscent or 3-valved, many- or few-seeded. Seeds obovate or
complanate, smooth, corrugate or sculptured. Drsrris.—Species 25,
chiefly in the warmer parts of Africa, several in Tropical Asia, a few
in Tropical America. ;
Leaves entire :—
Male pedicels ebracteate .., .. 1. M. Clarkeana.
Male pedicels with a bract close to the ee .. 2, M. subangulata.
Leaves 38-lobed, the lobules entire - .. & M. Cochinchinensis.
Leaves 5-7-lobed, the lobules sinuate-dentate or lobe
late .. yas wa} ee, .. 4 MM. Charantia,
hi i hdeaativl CLARKEANA, n. spec. King. Stem slender, 4-angled,
glabrous, 20-30 feet long. Leaves thinly membranous, broadly ovate,
never lobed, acute, the base deeply cordate or emarginate, both surfaces
quite glabrous ; length 3-5 in., breadth 3-4 in. ; petiole 1°5-2 in., slender,
eglandular. Male flowers ‘75 in. in diam., on filiform pedicels several
form a leaf-axil, or in few-flowered lax racemes; calyx-lobes puberulous,
broad, obtuse, their margins membranous and glabrous; corolla deeply
divided into 5 broad blunt lobes, puberulous.. Female flower unknown.
36 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
Fruit vermillion when ripe, sub-globular, apiculate when young,
glabrous; without ridges or papillae, 2-2°5 in. in diam.; seeds as in
M. Cochinchinensts.
Pprak ; Scortechini 1605 ; King’s Collector 8340; Wray 3273.
This species is allied to M. Cochinchinensis and has seeds exactly alike those of
that species. But the leaves of this are of thinner texture than those of M. Cochin-
chinensis, and they are not lobed ; moreover the petiole in this is eglandular, and the
fruit is quite free from ridges or papilla of any kind. I have named it in honour
of my friend Mr. C.B. Clarke who believes with me that it is a hitherto undescribed
species.
2. Momorvica supancuLata, Blume Bijdr. 928. Stem slender,
glabrous, angled, sulcate, several feet lony; tendrils single. Leaves
broadly ovate, deeply cordate at the base, the apex acute, the edges
with remote cartilaginous teeth; upper surface glabrous, the lower
with sparse small adpressed hairs ; length 1°75-2:25 in., breadth 1:25-
165 in.; petiole 1 in, long, glabrous, eglandular. Male peduncle
‘5-1 in. long, pubescent; the bract close to the flower broader than
long, its apex obtuse, its base cordate, minutely pubescent, the veins
prominent; flower nearly 1 in. in diam.; calyx deep purple; its lobes
oblong, obtuse, glandular, nearly ‘2 in. long; corolla yellow, partite
to the base, the segments oblong. Female flower unknown; fruit
ellipsoid, 2-3 in. long, obscurely ribbed, the ribs broken into joints.
Ser. in DC. Prodr. III, 316; Roem. Syn. fase. 2, p, 58; Mig. in Flor.
Ind. Bat. I, Pt, I, 664; Kurz in Journ. As. Soc. Beng. XLVI, Pt. II,
102; Cogn. in DC. Mon. Phan. III, 443.
Perak ; Scortechini 399. Distris. Java, Brit. India.
3. Momorpica Cocuin-Cuinensis, Spreng. Syst. Veg. III, 14.
A powerful climber ascending tall trees; stem angled, glabrous. Leaves
in general outline sub-orbicular or broadly ovate, the base cordate or
emarginate, usually 3-lobed to the middle or below it (sometimes 5-
lobed), the margins with sparse umbilicate glands, both surfaces
glabrous, length 4-7 in., breadth nearly the same; petiole 2-3 in. long,
glandular at the middle and upper part; tendrils long, stout, simple.
Male peduncle 2-6 in. long with an orbicular-reniform cucullate bract
at its apex embracing the flower-bud ; calyx-segments dark, coriaceous,
hairy ; corolla 1°75-3 in. across, white tinged with yellowish, some of
the petals with black spots at the base, others with yellow glands.
Female peduncle 1-2 in. long, (longer in fruit), with a small bract
about the middle. Frwt ovate, pointed at the apex, fleshy, bright red
and covered with conical points but not ribbed; 4-5 in. long; seeds
numerous, blackish, ovate, compressed, sculptured, the margins undulate-
sub-lobulate, Kurz in Journ. As. Soc. Beng. XLVI, Pt. 2, 102; Clarke
in Hook. fil. Flor. Br. Ind. II, 618; Cogn. in DC. Mon. Phan, III, 444.
1898.] G. King— Materials for a Flora of the Malayan Peninsula 37
M. mixta, Roxb. Hort. Beng. 70; Flor. Ind. III, 709; Wight & Arn.
Prodr. 349; Roem. Syn., fasc. 2, 59; Hook. in Bot. Mag. t. 5145; Mig.
Flor. Ind. Bat. 1, Pt. 1, 664; Naud. in Ann, Sc. Nat. Ser. 4, XII, 182.
M. dioica, Wall. Cat. (not of Roxb.) 6750, Ato F. Muricia Cochin-
chinensis, Lour. Flor. Coch.-Chin. 733; Ser. in DC, Prodr. III, 318.
Perak; Scortechini, King’s Collector, Wray; a common plant.
Disrris.—British India, Malaya, Philippines.
4, Momorpica CuarantraA, Linn. Sp. Pl. ed. I, p. 1009. Stem
slender, branching, striate, pubescent, sometimes tomentose towards
the apex, 3-6 feet long. Leaves reniform-orbicular in general outline,
]-3 in. in diam., deeply divided into 5-7 lobes ; the lobes sinuate-dentate
or lobulate and mucronate, constricted at the base, glabrous or slightly
pubescent; petiole 1-3 in.; the tendrils simple. Male peduncle slender,
with a reniform or orbicular bract at or below the middle. Male flower
°35-"75 in. in diam.; the calyx-lobes ovate, acute; corolla somewhat
irregular, yellow, its segments obtuse or emarginate, anther cells much
bent. Female peduncle 2-4 in. long, slender, bracteate near the base ;
ovary fusiform, muricate. Fruit 1-3 in. long, ovoid, tapering to both
ends, many-ribbed and bearing numerous triangular tubercles. Seeds
compressed, sometimes almost 3-toothed, the margins corrugated, the
sides sculptured. Lour. Flor. Cochchin. IT, p. 598; Bot. Mag. t. 2455;
Ser. in DC. Prodr. III, 311; Roxb. Flor. Ind, III, 707; Wight and
Arn. Prodr. 348 ; Torr. and Gray Flor. N. Amer. I, 543; Wight Ic.
tab. 504; Wight Ill. t. 105 t2s; Miq. Flor. Ind. Bat. I, Pt. I, 663; Cogn.
in Mart. Flor. Bras., fase. 78, p. 14; Clarke in Hook. fil. Flor. Br. Ind.
II, 616. Cogn. in DC, Mon. Phan. III, 436. M. muricata, Willd. Spec.
IV, 602; Roxb. Flor. Ind. III, 707; W. & A. Prodr. 348; Mig. Flor,
Ind. Bat. I, Pt. 1, 663. M. humilis, Wall. Cat. 6747. M. anthelmintica,
Schum. et Thou. Flor. Guin. 423. M. Rowxburghiana, Don Gen. Syst.
Gard. III, 35. M. macropetala, Mart. in Hook. Journ. Bot. V, 504.
Perak; Scortechini. Disrris. Malayan Archipelago, British India,
China, Tropical Africa and America.
7. Meorueta, Linn.
Climbing herbs ; tendrils simple or 2-fid. Leaves petioled, deltoid,
truncate or hastate, entire or deeply 3-lobed, little hairy, often punctate.
Flowers small, white, usually moncecious, males and females often from
the same axil ; male pedicels long, clustered (rarely sub-solitary) in the
axils, or clustered on long racemes resembling branches without leaves ;
female long-pedicelled. Male; calyx-tube short, teeth 5, small; corolla
5-partite ; stamens 3, inserted in the middle of the calyx-tube; anthers
free, one l-celled, two 2-celled ; cells free, straight, simple, more or less
38 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
lateral; connective prolonged, undivided, glabrous. Female calyx and
corolla as in the male; ovary oblong, style long ; stigmas 3, subglobose ;
ovules many, ts placentas 3, retLod. Fruit indehiscent,
globose, acute or fusiform, subrostrate. Seeds many, small, oblong,
much compressed, obscurely margined, smooth or very nearly so. Dis-
TRIB. Species about 55, all tropical.
Fruit not beaked :—
Fruit globular, glabrous me “ w. «€0L.. MM. affinis.
Fruit oblong, glabrous a aes ons te AE anager
Fruit beaked :—
Fruit fusiform, beaked, velvety ... bs . 3 M. marginata.
1. Mbtoruria aFFinis, n. spec. King. ath geaes the stem slender,
glabrous, 4-angled and deeply sulcate, not rooting ns the nodes; the
tendrils rather stout, bifid. Leaves ovate-cordate to triangular, Stee,
3-5-lobed, the edges denticulate ; upper surface densely scabrid-hispid,
the hairs white; the lower paler and with hairs of the same colour
but sparser and more slender ; length 1°35-4'25 in., breadth 1°5-3 26 in.,
petiole ‘5-1 in. Male and female flowers often from the same axil ;
the males in many-flowered umbels on pedicels twice as long as the
leaf-petioles, pubescent below, glabrous above ; pedicels about 20, slender,
unequal, dilated at the apex, glabrous, *25—35 in. long. Flowers ‘1 in.
in diam., globose, pubescent, with 5 slender short diverging subulate
teeth below the mouth. Petals not seen; anthers 3, straight, the conne-
tive not produced. Pedicel of the frwt shorter and stouter than the
peduncle of the male umbel, glabrous. Fruit globose, glabrous, thin-
walled, many-seeded, red when ripe, ‘4-5 in. in diam.; seeds obovoid,
pitted, somewhat compressed, pale.
Perak ; Scortechinit 495; Wray 860, 1404; Curtis ; King’s Collector
‘1069, 2539. Borneo; Bancermassine ; Motley 167.
Mr. C. B. Clarke, who was so good as to examine my specimens of this species
and to compare them for me at Kew, assures me that the Perak specimens agree
exactly with Motley’s 167. Mr. Clarke considers the species as closely allied to
M. marginata, Cogn. from which it differs by its globular glabrous fruit.
2. Metoruria inpica, Lour. Flor. Coch. China, 35. Stem slender,
filiform, glabrous, 3-6 feet long, often rooting at the nodes, the tendrils
simple. Leaves triangular-cordate, acute, entire or 3-lobed (sometimes
deeply); the nerves somewhat hairy, otherwise glabrous, the lobes
irregularly denticulate or lobulate; length 1°25-2 in. and breadth the
same, petiole ‘6-lin. Male pedicels solitary or in racemes of 2 or 3 on
peduncles 1 in. long; calyx-tube broadly campanulate, its teeth subu-
late, spreading ; corolla white, puberulons, its segments ovate-oblong.
Stamens. with thick obconic filaments, glabrous or sparsely villose;
anthers ovate-oblong, ciliate, the connective much produced. Pedunecle
1898.] G. King — Materials for a Flora of the Malayan Peninsula. 39
of female flower solitary, longer that the leaf-petiole. Fruit oblong,
glabrous, white, ‘5-75 in. long. Seeds ovate, attenuate at the base.
Ser. in DC. Prodr. III, 313; Naud. in Ann. Sc. Nat., Ser. 4, XVI, 169
t.2; Hance Suppl, Hongkong Flora, 104; Kurz in Journ. As. Soc.
Beng. XLVI, Pt. II, 105; Clarke in Hook. fil. Flor, Br. Ind. IT, 626.
Bryonia geminata, Blume Bijd. 924; Ser. in DC. Prodr. III, 305;
Roem. Syn., fasc. II, 35; Miq. Flor. Ind. Bat, I, Pt. I, 659. B. tenella,
Roxb. Flor. Ind. II], 725. Aechmandra indica, Arn. in Hook. Journ.
Bot. ITI, 274; Wight in Ann. and Mag. Nat. Hist. VIII, 267; Mig.
Flor. Ind. Bat. I, Pt. I, 658. |
SeLancore; King’s Collector.360. Disrris. British India, Cochin-
China, Philippines, Hongkong.
3. Metoraria MARGINATA, Cogn. in DC. Mon. Phan, IL]. 593.
Stem creeping, rooting at the nodes, angular, sulcate, glabrous; the
tendrils slender, simple, puberulous. Leaves ovate-cordate or more or
less 3-lobed, acute; the edges minutely and distantly denticulate ;
the upper surface coarsely and distinctly and the lower minutely and
more closely strigose, the nerves on both pubescent ; length 1-1-25 in.,
breadth 1-1°75 in. ; petiole *75-1°5 in., tomentose. Male flowers umbellate
on a filiform few-flowered peduncle shorter than the petiole, glabrous ;
the pedicels erect, ‘2-3 in. long; the calyx broadly campanulate, pubes-
cent, its teeth subulate. Petals villose, yellow; anthers glabrous, in-
appendiculate. Female flower solitary on a peduncle ‘5 in. long. Frutt
narrowly cylindric, beaked, tapering to the base, velvety, ‘75 in. long,
its peduncle filiform; seeds 6-8, foveolate. Bryonia marginata, Blume
Bijdr. 924; Ser.in DC. Prodr. III, 305; Roem. Syn., fase. II, p. 36;
Miq. Flor. Ind. Bat. I, Pt. IJ, 660. B. epigzeea, Blume Bijdr. 924; Ser. in
DO. Prodr. II, 306, Aechmandra Blumeana, Roem. Syn. fase. IT, p. 33;
Mig. Flor. Ind. Bat. I, Pt. 11, 657. Melothria Rumphiana, Scheff. Ann.
Jard. Bot. Buitenz. I, 25. Cerasiocarpum ? Maingayi, Clarke in Hook,
fil. Flor. Br. Ind. III, 629. .
Maxacca ; Maingay (Kew Distrib.) 1268. Prrax; King’s Collector
874. Disrris. Java and Sumatra. |
Var. heterophylla, Cogn. in DC, Mon. Phan. III, 594. Leaves quite
entire, ovate-cordate or oblong-sub-hastate, scabrous above. almost smooth
beneath but hairy on the nerves. Bryonia heterophylla, Blume Bijdr.
925; Wall. Cat. 6704; Roem. Syn., fase. IT, 35. B. Blumei, Ser. in DC.
Prodr. III, 305; Miq. Flor. Ind. Bat. I, Pt. I, 659. Cerasiocarpum ?
penangense, Clarke in Hook. fil. Flor. Br. Ind. III, 629,
Prnana; Wallich ; Curtis 1928. Disrrip. Java.
NOTE,
There are in the Calcutta Herbarium specimens of several species of Melothria
40 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
which Mr. C. B. Clarke, (who elaborated the family of Cucurbitacee for Hooker’s
Flora of British India, and who kindly compared my Malayan material in the Kew
Herbarium) considers as probably new. But, as these want either flowers or fruit,
I am obliged to leave them undescribed. The chief amongst these are Perak, Wray
2228 and 3416; and Andamans, King’s Collector 2200.
8. Gynostemma, Blume.
Climbing herbs, tendrils simple. Leaves pedate ; leaflets 3-5, ovate-
lanceolate, serrate, membranous. lowers small, dicecious, in axillary
diffuse panicles, greenish. Male flower ; calyx short, with 5 small lobes ;
corolla rotate, 5-partite, with lanceolate segments; stamens 5, filaments
connate below; anthers 2-celled, the cells straight and elongate.
Female flower, calyx and corolla asin the male; ovary rotund, 2-3-celled;
styles 2 or 3, united below, bifid at the apex; ovules 2 in each cell,
pendulous, Jrwit globose, umbonate, indehiscent, as large as a pea,
1-3-seeded. Seeds wingless, verrucose, sub-muricate. Species 4; all
Indo-Malayan. }
GyNosTeMMA PEDATA, Blume Bijdr. 23. Slender, 10-20 feet long;
young branches and leaves puberulous or glabrous, rarely pubescent.
Leaves membranous, trifoliolate or pedate, the petioles 1-1°5 in. long;
leaflets 3-7, ovate-oblong, lanceolate or oblanceolate, uneqnal, the
middle the longest, their apices acute or sub-acute, the bases narrowed
and sometimes oblique, the edges crenulate or crenate-dentate; length
°75-2'5 in., breadth ‘4-1 in., the petiolules ‘1-2 iu. Panicles longer
than the leaves, 3-6 in. or even a foot long, slender, sparsely branched,
more or less coarsely pubescent. Calyz-segments triangular, acute;
segments of corolla 1-nerved, ciliate-dentate, Fruit ‘15 im. in diam.
Seeds trigonous. Roem. Syn., fase, II, p. 110; Miq. Flor. Ind. Bat. I,
Pt. I, 683; Clarke in Hook. fil. Flor. Br. Ind. II, 683; Cogn. in DC.
Mon. Phan. III, 913. G. lawa, Cogn. Mon. Phan. IIT, 914; Zanonia
lawa, Wall, Cat. 3727; Pl. As. Rar. II, 29; Arn. in Hook. Journ.
Bot. III, 272 (in note). Zanonta cissoides, Wall. Cat. 3726; Pl. As. Rar.
II, 28. LZ. Wightiana, Arn. Pugill. 38; Nov. Act. Acad. Nat. Cur. 18, Pt.
I, 356; Roem. Syn., fase. II,117. Alsomitra laxa, Roem. Syn., fase. IT,
118. Pestolozzia lava, Thw. Enum. Pl. Zeyl. 124. P. pedata, Zoll. et
Moritz. Syst. Verz. p. 31. Alsomitra cissoides, Roem. Syn., fase. II, 118.
Enkylia trigyna, Griff. Pl. Cantor. 27; Miq. Prol. Flor. Jap., pp. 15 and
142. HE. digyna, Griff. Pl. Cantor. 27. Zanonia pedata, Miq. Flor.
Ind. Bat. I, Pt. I, 6838. Gynostemma cissoides, Franch. et Sav. Enum.
Pl. Jap. I, 176. Vitis atroviridis, Wall. Cat. 6040; Vitis trichophora,
Wall. Cat. 9032.
Perak; King’s Collector 2306. Disrris. Malayan Archipelago,
British India and Tonkin.
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 41
This plant varies as to the number of its leaflets in its leaves and as to pubes-
cence. The pedate forms with 5-7-puberulous leaflets have been by some authors
(among whom is M. Cogniaux, the latest monographer of the Cucurbitacex) con-
sidered as belonging to a different species from the plants with trifoliolate glabrous
shining leaves; and the latter has been named G. lava. After carefully examining
about a hundred specimens collected in different parts of British India and
Malaya, I find so many that unite to some extent the characters relied upon as
distinctive that I have been driven, with all respect to M. Cogniaux, to adopt Mr.
Clarke’s view that there is but a single species.
9, Zanonta, Linn.
Climbing herbs; tendrils simple. Leaves long-petioled, simple,
ovate or oblong. lowers small, dicecious, in large compound pendulous
racemes, males pedicelled, females subsessile. Male; sepals 3, oblong
or orbicular, concave; corolla rotate, 5-partite, the segments subacute ;
stamens 5, free, inserted on a fleshy disc, filaments very short; anthers
1-celled, transversely oblong. Female; calyx and corolla as in the male ;
ovary sub-clavate, at first 3-celled ; styles 3, spreading, 2-fid at the apex ;
ovules in each cell 2 or many, pendulous, attached in 2 series to a fleshy
parietal placenta. Capsule large, elongate-cylindric, clavate, 3-valved
at the truncate apex. Seeds large, oblong, pendulous, compressed, sur-
rounded byalarge membranous wing. Disrris. Species3; British India,
Malaya.
Leaves ovate-oblong ; fruit 2°5 in. long eft ww L 2. indica.
Leaves ovate-rotund ; fruit 5-8 in. long ca oo. 2. Z. Clarkei.
1. Zanonta 1npicA, Linn. Spec. Pl. ed. IT, 1457. Slender, climbing
to the extent of 30-50 feet, glabrous. Leaves coriaceous, ovate-oblong,
acute; the base 3-nerved, rounded or somewhat emarginate; main
nerves 6-8 pairs, curved, spreading, prominent beneath; length 3°5-6
in., breadth 2-3'5 in., petiole ‘65-8 in. Fruit cylindric-campanulate,
glabrous, the apex truncate, 2°5 in. long; seeds flat, thin, 1‘5-1°75 in.
long ; the nucleus oval, only ‘6 in. long, the rest being wing. Blume
Bijdr. 937; Ser. in DC. Prodr, III, 298; Roem. Syn. fasc. II, 117;
Wight and Arn. Prodr. 340; Wight Ill. t. 103; Maiq. Flor. Ind. Bat. I,
Pt. I, 682 ; Thwaites’ Enum. Pl. Zey. 124, 442 ; Clarke in Hook, fil. Flor.
Br. Ind. II, 633; Trimen Flora Ceylon II, 261.
Perak ; King’s Collector 7198, 7362.
2, ZanontA CLARKEI, n. spec. King. Slender, 60-80 feet long,
glabrous, the young branches deeply grooved, Leaves coriaceous, ovate-
rotund, acute ; the base broad, emarginate or slightly cordate, with 5-7
nerves radiating from the apex of the petiole (the lateral one on eack
side small); length 2°5-3°5 in., breadth 2°5-3 in., petiole ‘75 in., tendrils
bifid. Flowers unknown. Fruit ovoid-cylindric, smooth, 5-8 in. long,
a... 6
42 G. King— Materials for a Flora of the Malayan Peninsula. [No. J,
and 3 in. in diam. Seeds flattened, about *1 in. thick, ovate, with 6
unequal deep narrow marginal lobes (those at the ends being the
longest), the seed proper or nucleus papillose, lin. long and °75 in,
broad, surrounded on all sides by a thin membranous wing nearly J in.
wide.
Perak; King’s Collector 7230.
This fine species is known only from a single gathering of six specimens, none
of which bears a flower. My friend Mr. C. B. Clarke, F.R.S., who was so kind
as to compare for me at Kew the whole of my Malayan Cucurbits, notes on this as
follows: ‘‘ This is either a new Zanonia near Z. indica, Linn., ora new Alsomitra
near A. Capricornica, F. Mull.—I think a Zanonia, whether the sepals turn out to be
5 or not. The 2-fid cirrhi may do for either genus. The spinose-margined seeds are
not like those of Zanonia; but the ovoid, or sub-globose fruit is like nothing out
of section Macrozanonia, Cogn. I call it Zanonia, n. spec.”
10. Atsomirra, Bth. & Hk. f.
Large climbers; tendrils simple or 2-fid. Leaves with 3 oblong
entire leaflets, Flowers small, dicecious, white, in compound panicles
with filiform branches. Male; calyx rotate, 5-partite, segments oblong,
acute; corolla rotate, 5-partite, segments obtuse ; stamens 5, filaments
short, near together at the base; anthers small, oblong, straight, 1-
celled. Female; calyx and corolla as in the male; ovary elongate-
clavate, l-celled ; styles 3-4, conical, with semi-lunate stigmas; ovules
very many, pendulous; placentas 3, thick, vertical, parietal. Capsule
large, elongate-clavate, truncate and 3-valved at the apex. Seeds very
many, compressed, vertical, in six rows, much corrugated, incised or
horned on the margin with a terminal membranous wing longer than
the seed. Drsrris. Species 9; British India, Malaya, North Aus-
tralia, S. America.
ALSOMITRA CLAVIGERA, Roem. Syn. fase. IJ, p. 118. <A slender
glabrous climber. Leaflets fleshy, the middle the largest, oblong or
oblong-lanceolate, the apex obtuse, the base narrowed, the edges entire,
subsessile, eglandular, length 1'5-3 in., breadth °65-1l in, Panicles
slender, twice as long as the leaves, longer in fruit, Capsule smooth,
1:25-1'5 im. long and ‘4 in. broad. Seeds dark, cinereous, shortly
muricate. Cogniaux in DC. Mon. Phan. III, 927; Hook. fil. in Bot.
Mag. t. 6017; Clarke in Hook. fil. Flor. Br, Ind. II, 634. Zanonia
sarcophylla, Wall. Cat. 3724; Pl. As. Rar. IT, 28, t. 133.
Kapa ; Curtis 2504,
Ordered LVI ARALIACEA,
Trees or shrubs, very rarely herbs, often scandent, sometimes
prickly. Leaves alternate, the uppermost rarely sub-opposite, long-
1898.] G. King—Waterials for a Flora of the Malayan Peninsula. 43
petioled, large, simple or compound; stipules adnate to the petiole,
sometimes inconspicuous or 0. Flowers regular, small, often polygamous,
in umbels racemes or umbellate panicles; bracts and bracteoles small
or conspicuous; pedicels continuous with the base of the calyx or
jointed. Calya-tube adnate to the ovary; limb truncate, obsolete or
with small teeth. Petals 5, rarely 6-7 or many, valvate or sub-imbricate,
expanding or calyptrate. Stamens as many as and alternate with the
petals (very many in Tupidanthus), inserted round an epigynous disc.
Ovary inferior, 2-celled, or cells as many as the stamens, or 1-celled ;
styles as many as the cells, distinct or united; ovules solitary and
pendulous in each cell. Fruit coriaceous or drupaceous, usually small,
one or more cells sometimes suppressed. Seed pendulous, albumen
uniform or ruminate; embryo minute, radicle next the hilum. Distrrs.
Species 400, chiefly tropical and subtropical, a few in the cool temperate
zones,
Petals imbricate (slightly); pedicels of flowers
jointed :—
Styles 2-5, free; leaves compound ... 1. ARALIA.
Styles 3 or 4, free; leaves simple, entire, lobed or
pinnatifid ; ovary 1-celled sae .. 2, ARADIDIUM..
Petals valvate ; stamens not exceeding 12 :—
Albumen uniform, ovary 4-12-celled, pedicels continu-
ous with the flower, leaves simple or digitately com-
pound :—
Fruit boldly ridged, never more than ‘3 in. long;
leaves usually compound .. “i .. 93 HePpTAPLEURUM.
Fruit succulent, aS alee ‘5 in. in pall leaves
simple, large, rotund-reniform, lobed . «ee 4 'PREVESIA.
Fruit succulent, ‘3 in. in diam., obscurely ridged ;
leaves simple, entire (palmately lobed in young
shoots ) ore ae eos eo 5 DENDROPANAX,
Albumen ruminate :—
Ovary 1-celled :—
Ovary 1l-ovulate, leaves compound ... w. 6. ARTHROPHYLLUM.
Ovary with 2 ovules; fruit 2-celled, 2-seeded,
leaves simple “oe a -. % WARDENFA.
Ovary 2-celled, pedicels continuous :— ;
Style distinct ; leaves pinnately decompound ... 8, HETEROPANAX.
Styles. combined; leaves digitate, palmate or
angled ove ove ... 9 BRASSAIOPsIS.
Ovary 4- or 5-celled, pedicels joinind Fe .. 10. HeETEROPSIS.
Petals valvate ; stamens 30-50 es .» Ll. Tupranruus,
1, Aratia, Linn.
Herbs, shrubs or small trees, glabrous, hairy or prickly. Leaves
alternate or whoried, digitate, pinnate or compound-pinnate ; leaflets
44 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
serrate or nearly entire; bracts and stipules not prominent. Umbels
solitary or in racemes or panicles, rarely in compound umbels; pedicels
usually jointed close under the flower. Flowers often polygamo-mone-
cions. Calyx with its margin truncate or 5-toothed. Petals 5, ovate,
imbricate in bud. Stamens 5. Ovary 2-5-celled; styles 2-5, free or
shortly connate at base. Fruit 4-5-celled, 4-5-angular, or subglobose,
2-3-celled. Albwmen uniform. Dustris. Species about 50; extending
from India and Malaya to Japan and North America.
Leaflets entire or minutely serrulate, densely ferrugineous-
tomentose on the lower surface oem oe «. I, A. Thomson:
Leaflets coarsely and somewhat unequally serrate, the lower
surface with scattered flexuose white hairs ove oo. 2 A. armata.
Leaflets obscurely undulate-serrate, quite glabrous on both
surfaces, the lower sub-glaucous _... Sr iteee ww Os A, ferow.
1. ArattA THomsont, Seem. Rev. Hed. 91. A large prickly shrub,
all parts more or less softly pubescent or tomentose. Stem slender,
the prickles remote. Leaves large, 2-3-pinnate ; leaflets thickly membra-
nous, 5-9 in each ultimate pinnule, ovate-lanceolate, acuminate, the
base rounded and somewhat oblique; the petiolules of all except the
terminal very short (usually less than ‘1 in. long), that of the terminal
leaflet °5 in. or upwards, the edges entire or minutely serrulate; upper
surface sparsely strigose, the nerves and midrib pubescent ; the lower
softly tomentose ; length 2°5-3°5 in., breadth 1-1°75 in. Panicles 15-20
in. long, with short sub-horizontal branches; the ultimate branches
umbellate ; the bracts narrowly lanceolate, ‘25 in long. Fruit *15 in.
long, somewhat wrinkled, glabrous, the ridges broad. Clarke in Hook.
fil, Flor. Br. Ind. TI, 723.
Pawane; Ridley 2450. Prnana; King’s Collector 1574; Curtis
462. Prax; King’s Collector 7807, 8704, 10045.
2. ARALIA ARMATA, Seem. Rev. Hed. 91. A prickly shrub; stem
slender, almost glabrous except the puberulous young parts. Leaves
very large, 3-pinnate, the rachises puberulous ; leaflets thinly mem-
branous, 9-11 in each ultimate pinnule, ovate-lanceolate, acuminate,
the edges coarsely and somewhat nnequally serrate ; the base rounded,
slightly oblique or sub-cordate ; sessile except the terminal one; petiolule
of the terminal leaflet under 1 inch ; length of leaflets 3-5 in., breadth
15-2 in.; upper surface with a few sparse adpressed hairs, otherwise
glabrous ; lower with scattered white flexuose hairs especially on the
nerves and midrib. Panicle 15-18 in. long, pubescent in its ultimate
3-fid branches, the lower part glabrous, ultimate branchlets umbellate ;
the pedicels slender ‘3-4 in. long, tomentose. Fruit nearly ‘2 in. long,
glabrous; the ridges broad, flat. Kurz For. Flora Burma, I, 536;
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 45
Clarke in Hook. fil. Flor. Br. Ind. II, 723. Panazx armatum, Wall.
Cat. 4933; G. Don. Gen. Syst. TII, 386.
Kepau ; Ourtis 2526. Distris. British India; in Burma, the Khasia
Hills and on the lower slopes of the Hastern Himalaya.
3. ARALIA FerOX, Miq. Flor. Ind. Bat. I, Pt. I, 750. A lax spread-
ing glabrous shrub, often scandent to 20 or 30 feet ; the stems, branches,
rachises of the leaves and inflorescence bearing numerous short recurved
spines. Leaves 2-3-pinnate, the pinne 3 or 4 pairs; leaflets sub-
coriaceous, 3-5 in a pinna, the pairs opposite, ovate, acute, the bases
rounded, the edges obscurely undulate-serrate ; both surfaces glabrous,
the upper shining when dry, the lower sub-glaucous ; length 1-2 in.,
breadth *75-1 in. ; petiolules unequal, the lateral '15-"3 in., the terminal
5-65 in. long. Panicle terminal, 8-15 in. long and 6 in. across,
with numerous slightly compressed horizontal branches, themselves
branching and ultimately ending in numerous peduncled umbels of
10-15 long-pedicelled oblong flowers ‘1 in. long; the pedicels slender,
3-5 in. long. Calyx-tube campanulate, 10-ridged, the limb with 5
small triangular acute teeth. Petals ovate. Fruit ovate-globose,
boldly 5-ridged, rather more than ‘] in. long.
Perak ; Scortechint 142, 501; King’s Collector 1037, 4434, 5089,
8438, 10568 ; Wray 2155. |
2. ARatipium, Miq.
Leaves large, simple, usually deeply lobed or pinnatifid, glabrous .
Flowers male or hermaphrodite, in large compound panicles, minute.
Calyx-teeth triangular, spreading, the tube campanulate. Petals 5,
imbricate. Stamens 5. Ovary usually 3-celled, two of the cells soon
aborting. Styles distinct, subulate. Pruzt obliquely ovoid, drupaceous,
1 inch or more in length, 1-seeded ; the seed solitary, rugose, pendulous,
vertically sulcate ; albumen very copious, coarsely ruminate, penetrated
by outgrowing folds from the funicle; embryo small. Distrip; two
species, both Malayan.
This is a remarkable genus of doubtful position. The large solitary seed, with
a much developed funicle forming an expansion at the base of the coarsely
ruminated albumen and sending processes into the latter, and the unisexual habit
make it doubtful whether it should not be placed in Cornaceez (to which Seemann
referred it), rather than in Araliacex.
ARALIDIUM PINNATIFIDUM, Miq. Flor. Ind. Bat. I, 763,t.13. A
small tree without prickles, glabrous except the inflorescence. Leaves
thinly coriaceous, irregularly lobed or coarsely pinnatifid, rarely entire
and narrowly elliptic ; length of the lobed or pinnatifid forms 10-18 in.,
breadth 7-10 in.; length of the entire leaves 4-10 in., breadth 2-4 in.;
petiole stout, 15-5 in. long. Pamnicles many-branched, ferruginous-
46 G. King— Materials for a Flora of the Malayan Peninsula. (No. I,
puberulous, shorter than or as long as the leaves. Petals pubescent.
Fruit narrowly ellipsoid, pointed at each end, glabrous, the pericarp:
thin, length 1°35 in., diam °6 in., damson-colored when ripe. Miq. Flor,.
Ind. Bat. Suppl. 340; Clarke in Hook. fil. Flor. Br. Ind. II, 726;
Hemsley in Hook. Ic, Plantar. t. 1549.
Panane ; Ridley 2439. Snrancor; Curtis 2340. Stncarore; Ridley ;:
Hullett, 485, 888; King. Panana; Ridley 2662. Matacca; Griffith:
(Kew Distrib.) 2702; Maingay 676. Prrax ; Scortechini ; Wray ; King’s
Collector (many numbers), a very common plant. Duistris. Sumatra,.
Forbes 2207,
The structure of the seed is well explained by Mr. Hemsley in his description
and figure in Hooker’s Icones Plantarum.
3. Heprapniurum, Gertn.
Large shrubs or trees, glabrous or tomentose, without prickles.
Leaves digitate, rarely compound-digitate or 1-foliolate; leaflets coriaceous,.
entire or remotely toothed or lobed, never closely serrate nor ciliate,
those near the panicle usually entire; stipules often connate within the
petiole and prominent. Umbels (rarely capitate) panicled or in com-
pound racemes; bracts woolly, deciduous or persistent; pedicels not
jointed under the flower; bracteoles few or 0, or rarely densely tufted.
Calyx-margin toothed or truncate. Petals 5-6 or many, valvate. Stamens
as many as the petals. Ovary cells as many as the petals, disc small
or large; styles small, separate or combined in a short comical narrow
cylindric column. Fruit subglobose, 5-6-angled. Seeds compressed,
albumen uniform. Disrris. Species 55, in the tropics of the Old World.
Leaves simple ove BA so o. Ll. H. avene,
Leaves all trifoliolate :—
Panicle longer than the leaves, slender; its branches
long, spreading, laxly-flowered ... Af - 2. H. luridwm,
Panicle shorter than the leaves; its meaneies adit:
densely-flowered ... a wae oo & H. triste,
Leaves all digitately 5-9-foliolate :—
Panicle simple or 2-branched, sub-racemose, only about
3 in. long; the lateral umbels subsessile, lax, few-
flowered an Re sy .. 4. H. sub-racemoswm.
Panicle divided from the base or near it into several
erect narrow branches 6-12 inches long; the lateral
branches very short and bearing dense heads of sub-
sessile flowers :—
Ovary 8-celled, leaves glaucous and with loose wooly
hairs beneath ay Bae aw .. & H. Cephalotes.
Ovary 5-celled :—
Both surfaces of the leaves quite glabrous .. 6. H. Scortechinu.
Lower surface of leaves tomentose we . % HH. tomentosum..
1898.] G. King— Materials for a Flora of the Malayan Peninsula.
Panicle with few narrow erect branches bearing shortly
peduncled umbels of pedicellate flowers :—
Ovary 12-celled; leaflets 5, membranous, 7-12 in.
long a waa ae oni
Ovary 9-celled; leaflets 7-9, very coriaceous, 3-5
in. long ove
Ovary 5-celled :—
Main nerves of leaflets only 2 or 3 pairs, the basal
prominent; reticulatious rather wide, not promi-
nent; panicles very narrow, covered, up to the
bases of the pedicels, with minute stellate hair
Main nerves of leaflets 5 or 6 pairs, basal nerves
prominent ; panicles glabrous {rarely hairy) narrow
in flower, spreading in fruit
Panicle with spreading branches :—
Branches of the panicle umbellate :—
Ovary 5-celled; reticulations of leaves wide, dis-
tinct eee ove ade vee
‘Ovary 6-celled :—
Common petiole 1-2°5 in. long; leaflets lanceo-
late or oblanceolate, caudate-acuminate, much
narrowed to the base, 1°5-3'5 in. long; flower-
buds globular, *1 in. in diam. ;
Common petiole 2-4 in. long; leaflets aie
lanceolate, shortly caudate-acuminate, the bases
rounded, 2-4 in. long; flower-buds oblong, about
‘2 in. long ; fruit 25 in. long
fruit *1 in. long ..
Common petiole 10-24 in. long; leaflets oblong.
elliptic, shortly and abruptly acuminate, 4-12
long ; flower-buds sub-globular, ‘1 in. in Pern
fruit elliptic, ‘25-3 in. long
Common petiole 5-10 in, long; ieaflats abliig:
elliptic or oblanceolate-oblong, acute, 5-10 in.
long; flower-buds globular, *15 in. in diam. ;
fruit ovoid, *25 in. long ... 78 ;
Common petiole 2-2°25 in. long; leaflets lanceo-
late, acute, narrowed to the base, 1°5—2°5 in. long ;
flower-buds oblong, °*2 in. long, fruit ‘3 in, long...
Branches of the panicle racemose
Lower leaves large, digitately decompound :—
Leaflets entire, narrowly oblong, acuminate, 2-3°5 in.
long, panicle 8 in. long see
Leaflets usually entire, 2°5-7 in. long, Bich 10-15 in.
long ; fruit oblong a
Leaves bipinnate, leaflets donmiolke toothed ;
‘globular eee vee
“elie sub-
aye (t
fa |e
12.
aes
. £6,
. 16.
Ui
» 18.
ae 8
3 20.
pels
A7
H. Singalangense.
H., lati-foliolatum.
H. subulatum,
H, venulosum,.
H, ellipticum.
H. scandens.
H, affine.
H, Hullettis.
H. Ridleyi.
H. nervosum.
H. Wray.
H. biternatum.
H. heterophyllum.
A. Curtisi.
1. HeEprapLevRUM AVENE, Seem. ta Hed. 43. Scandent ; young
branches with striate rugulose glabrous bark, brown when dry. Leaves
48 G King—Materials for a Flora of the Malayan Peninsula. [No. 1,
simple, coriaceous, narrowly oblong, acute, narrowed to the base, the
edges slightly recurved ; both surfaces glabrous, dull when dry; main
nerves about 12 pairs, faint, sub-horizontal ; length 3°5-5°25 in., breadth
‘9-1-2 in.; petiole *75-1:25, thickened near the apex. Panicles single or
2 or 3, terminal, puberulous, half as long as the leaves, erect ; the few
branches rather close together, about °25 or *3 in. long, each bearing
3-5 pedicellate flowers ; the buds ‘15 in., ovoid, their pedicels ‘2 in. long ;
bracts lanceolate, deciduous. Calyx-rim narrow; petals, stamens and
stigmas 6. Jrutt ellipsoid, as large as a pea, sulcate. Sciadophyllum
avene, Herb. Korthals.
SinGAPORE; Ridley 5840, 6337. Distris. Sumatra.
At once distinguished in the genus by its oblong simple leaves.
2. HEPraPLEURUM LURIDUM, new species. An epiphytic shrub 2-3
feet long ; branches with rugulose glabrous bark. Leaves trifoliolate,
with a common petiole 1-1'75 in. long, glabrous, fleshy when fresh, ver-
tically rugose when dry; leaflets very coriaceous, narrowly elliptic-ob-
long, slightly oblique, tapering gradually to each end; the edges entire,
much recurved when dry; both surfaces quite glabrous, the nerves
and veins very indistinct even when dry ; length 2°5-4 in., breadth 5-75
in.; petiolules unequal, thick, the lateral ‘15-25 in. long, the terminal
about ‘4 in. Panicle terminal, longer than the leaves; its branches few,
slender, long, spreading, glabrous; the ultimate branchlets 1-1°5 in.
long, bearing umbels of 3-8 broadly ovoid glabrous flowers ‘1 in. long;
their pedicels at first only about *1 in. long, but two or three times
as long in fruit. Calyzx-tube shortly campanulate ; its mouth truncate,
entire. Petals completely united into a calyptra. Stamens 6, erect,
the filaments short, Styles united into a short conical column; the
stigmas 6, small, occupying the corners of its truncate apex.
Perak ; Scortechint 1191; King’s Collector 8304.
A very distinct small species easily recognised by its narrow very acuminate
avenous leaflets borne on fleshy petioles, and by its slender spreading long-branched
few-flowered panicles.
3. HEPTAPLEURUM TRISTE, new species. A tree ; young branches as
thick as a swan’s-quill, glabrous, pale when dry. Leaves 3-foliolate,
the common petiole 15-2 in. long; leaflets thickly coriaceous, glabrous,
broadly elliptic, blunt or shortly apiculate, the base rounded ; the edges
entire, boldly recurved when dry ; main nerves 10 to 12 pairs, close to-
gether, indistinct on both surfaces; length 2°25-3°25 in., breadth 15-2
in.; petiolules unequal, the lateral pair ‘5 in. long, the middle one ‘8 in.
Panicle terminal, shorter than the leaves, glabrous, divided from the
base into 2 or 3 spreading branches, ebracteate; the branchlets about
‘Din. long (longer in fruit), each ending in an umbel of 10-20 ovate
1898.] G. King — Materials for a Flora of the Mulayan Peninsiila. 49
flowers, ‘15 in. long, their pedicels ‘1 in. long. Calyaz-tube campanu-
late ; the limb truncate, narrow, Petals 5, broadly elliptic, blunt, re-
flexed. Fruit narrowly oblong, pointed, deeply 5-ridged, glabrous,
erowned by the short conical style-column, ‘2 in. long, glabrous.
Perak ; on Ulu Batang Padong, at an elevation of about 4900
feet ; Wray 1509.
A species near H. ellipticum but readily distinguished from that in the Herba-
rium by its dull broad coriaceous leaflets, more widely campanulate calyx-tube,
and larger fruit.
_ 4. HepraPpLEURUM SUB-RACEMOSUM, new species. A shrub 2-3 feet
high; young branches with thick corky glabrous bark, pale when dry.
Leaves trifoliolate or sometimes 5-folioiate, the common petiole 1°5-2°5
in. long, glabrous. Leaflets coriaceous, narrowly elliptic-lanceolate,
caudate-acuminate, narrowed at the base, the edges entire and slightly
revolute ; both surfaces glabrous, distinctly and finely reticulate wheu
dry; length 2°25-4 in., breadth °75-1-] in.; petiolules unequal, 3-4 in.
long, that of the terminal leaflet *8-1'25in. long, thickened at the apex.
Panicle solitary, terminal, very narrow, sub-racemose, 2-branched, only
about 3 inches long and about °65 in. across; the branchlets only *1—2
in. long, each ending in an umbel of 3-5 oblong, green flowers °15 in.
long; rim of calyx very narrow. Fruit oblong, *2 m, long, glabrous,
with 5 blunt ridges, pale green tipped with bluish, 5-celled.
‘Perak ; King’s Collector 8283.
Readily recognised by its small caudate-acuminate narrow leaflets, much
and prominently reticulated when dry; and by its short narrow racemoid panicles.
5. HerrarLnevrum Crrpuarotes, Clarke in: Flor. Br. Ind. [I, 731.
‘A large tree. Leaves digitate, their petioles rather slender, 1:5-3°5 in.
long, glaucous; leaflets about 7, oblong-elliptic, the apex abruptly and
shortly acuminate, the base rounded, the edges entire and slightly
recurved when dry; upper surface shining, glabrous; the lower dull ,
glaucous, and bearing when young some quickly deciduous loose wooly
hairs. Panicle minutely tomentose, terminal, consisting of numerous
sub-erect branches 6-12 inches Jong,*bearing shortly peéduncled globose
ebracteolate dense capitula. Fruit oblong, ribbed, 8-celled, covered
with stellate white tomentum and crowned by the broad cluster of
Short styles ; disc large, spongy. H. capitatum, Seem. Rey. Hed. 15
(in part). Vonage ae
Maracea; Griffith (Kew Distrib.) 2700. Sincaporr; Iidley 3973,
6409. Prnana; Onrtis 837. Perak; Wiay 1542; Scortechini 391.
6. Heprartevrum Scorrechtntl, new species, An epiphytic shrub,
the young branches stout. Leaves digitate; the common petiole terete,
glabrous, stout, 2°5-3'5 feet long ; leaflets 8-10, very coriaceous, oblong
oN. 7 ith )
50 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
or oblong-elliptic, the apex blunt or shortly apiculate, the base rounded,
the edges entire and recurved when dry; both surfaces quite glabrous,
not reticulate; main nerves 6-9 pairs, slightly prominent on the lower
surface; length 9-13 in., breadth 4-6 in.; petiolules unequal, 2°5-5 in.
long. Panicle terminal, dividing into several narrow raceme-like
branches, 10-20 inches long, covered with pale scurfy tomentum ; the
branchlets 40 or 50 in number and °35 to ‘75 in. long, each with a
broadly ovate convolute wooly bract at its base, and at its apex a dense
globular umbel of sub-globular flowers ‘1 in. in diam. and borne on
pedicels 05 in. long. Calya truncate, tapering to the base. Petals 6 or 7,
narrowly lanceolate. Stigmas free, ovary 5-celled. Fruit unknown.
Perak; Scortechini 2008.
Collected only by the late Father Scortechini, whose specimens unfortunately
are scanty and rather fragmentary. A very distinct species, in general appearance
resembling H. Cephototes, Clarke and H, rigidwm, Seem. The leaflets of the
latter have however twice as many main lateral nerves, although their length
is not greater.
7. HepraPLEURUM TOMENTOSUM, Hassk. in Cat. Hort. Bot. Bogor.
(1844), p. 165, A half-scandent shrub, 6-8 in. high ; the young branches
stout, deciduously stellate-pubescent, the older glabrous. Leaves digitate ;
the common petiole 9-12 in. long, densely covered with rusty stellate
tomentum, the intra-petiolar stipules about ‘75 in. long. Leaflets 5-7,
coriaceous, oblong-elliptic, caudate-acuminate, slightly narrowed to the
base, the edges entire and slightly revolute; the upper surface boldly
bullate and finely reticulate, quite glabrous, pale olivaceous when dry ;
the lower pale brown, more or less closely covered with pale brown
stellate hairs; length 8-10 in., breadth 2°75-3°5 in.; petiolules unequal,
15-2°5 in. long, tomentose. Panicles usually two together, terminal,
4-5 in. long, racemose ; the lateral branches about 12, short, ‘2-4 1n. long,
each bearing asub-globular umbel of 8-10, ovoid, blunt, glabrous flowers
‘15 in. long; their pedicels ‘1 in. long, pubescent. Calyx-tube funnel-
shaped, the rim narrow. Petals 5, white, glabrous, calyptrate. Stamens
longer than the petals. Styles 5, conjoined, papilliform, their apices
free. Fruit *2 in. long, narrowly oblong, 5-ridged, 5-celled. Sciadophyllum
tomentosum, Blume Bijdr. 877; DC. Prod. IV, 260. Paratropia
tomentosa, Miq. in Bonplandia for 1856, p. 188; Flor. Ind. Bat. I, Pt. I,
753; Aun. Mus. Lugd. Bat. I, 23.
Perak; Scortechini, Wray 2202, 3152; King’s Collector 2066, 2216,
2569, 7253, 8733. SnLanagor; Curtis 2341. Distris. Sumatra; Forbes,
2611.
I have seen no authentically-named specimen of this from the Leiden Herba-
rium, but the Perak plant agrees so completely with Miquel’s description that I
have no doubt of the correctness of my identification.
1898.] _G. King— Materials for a Flora of the Malayan Peninsula. 5%
8. HeprapLevrum SInGaLANGenseE, Seem. Rev. Hed. 42. Scandent,
glabrous. Leaves digitate ; common petiole 12-15 in. long, stout ; leaflets
5, membranous, elliptic, shortly acuminate, the base slightly narrowed,
the edges with a few irregular remote teeth or entire; both sur-
faces glabrous, faintly reticulate; main nerves 7 or 8 pairs, curved,
Spreading ; length 7-12 in., breadth 3'25-5°25 in., petiolules 1*25-2°5 in.,
thickened at the base. Panicle 12-18 in. long, covered with deciduous
rusty steilate pubescence, very narrow, (about 2 in. wide) ; the branches
numerous, about *5 in. long, each ending in an umbel of 5-10 oblong
obovoid pedicelled flowers *25 in. long, their pedicels -2—3 in. long.
Calyx-tube shortly campanulate, minutely scaly ; its limbnarrow, truncate,
undulate. Petals 9, fleshy, narrow, slightly unequal, connate by their
edges. Stamens 9. Stigmas united into a notched fleshy ring, ovary
12-celled. Fruit unknown. Paratropia Singalense, Mig. in Ann. Mus.
Imgd. Bat. I, 23. Agalma redivivum, Seem. Rev, Hed. 25.
Perak ; Scortechini 390; Curtis 3170.
The specimens which I have seen are few. In foliage they agree with the
specimen in Kew named Agalma redivivum, which however Seemann describes as
having 7-8-merous flowers. In other respects they agree with Seemann’s description
of that plant. They also agree with Miquel’s full description of his Paratropia
Singalangense, except as to the length of the petiole which Miquel gives as only 4-6
Inches, a measurement so much out of proportion with those he gives for the
petiolules, (1-2°5 in.) and leaflets, that I cannot help suspecting some clerical error.
9. HePTAPLEURUM LATIFOLIOLATUM, new species. A bush with stout
branches. Leaves digitate; the common petiole stout, glabrous, some-
what compressed, 7-11] in. long; leaflets thickly coriaceous, ovate-
rotund, shortly acuminate or blunt, the base rounded ; the edges entire,
slightly recurved when dry; both surfaces glabrous, the upper shining,
the lower dull and sub-glaucous; main lateral nerves 7 or 8 pairs,
spreading, curving upwards, prominent on the lower surface, depressed
on the upper when dry; length 3-5 in., breadth 1°75-3'5 in.; petiolules
*8-1°5 in., rather stout. Panicles terminal, in pairs, 5-8 in, long,
glabrous; the branches short, horizontal, each bearing at its apex an
umbel of 10-15 pedicelled flowers; pedicels °3 in. long. Fruit broadly
ovoid, deeply 9-ridged, 9-celled, glabrous, crowned by the 9 very short
free styles, and ‘15 in. in diam., red when ripe.
Perak; on Gunong Babu at an elevation of about 5000 feet ;
Wray 3927.
10. HeprarLevrum susuntatum, Seem. Rev. Hed. 42. Scandent,
glabrous. Leaves digitate; the common petiole 2-6 in. long, slender,
glabrous, terete; leaflets coriaceous, oblong-elliptic or elliptic, shortly
acuminate, tapering to the base; the edges entire, revolute when dry ;
both surfaces glabrous and reticulate but not prominently so, the upper
52. G. King — Materials for a Flora of the Malayan Peninsula. [No. 1,
shining, the lower dull when dry ; midrib prominent on the lower surface
as also the 4 oblique, basal, and 2-3 pairs of sub-horizontal main nerves ;
length 2°5-7 in., breadth 1—3°5 in; petiolules unequal, *2—2 in., swollen
near the apex. Panicle terminal, stellately puberulous, shorter than the-
leaves while in flower, longer in fruit, bearing many very narrow
racemoid branches each with an acuminate lanceolate bract 1 in. long’
at its base; branchlets very short, from °15 in. when in flower to ‘5 in.
when in fruit, stellately pnberulous, each bearing an umbel of 12-20:
flowers; buds depressed, globular, glabrous, ‘05 in. in diam. ; their pedicels:
‘2in. long, slender Iruit rather broadly ovoid, bluntly 5-ridged, 5-.
celled, glabrous, ‘15 in. long. Clarke in Flor. Br. Ind. IT, 730. Paratropia
subuiata, Mig. in Ann. Mus. Lugd. Bat. I, 22. .
Maracca; Griffith (Kew Distrib.) 2690; Maincay; 681, (Kew~
Distrib.) Derry. Pananc; Ridley 1632, 5818. Perak; Scortechini;
Wray 872, 3090, .3639; King’s Collector 773, 1102, 2343. Dustrip.
Sumatra.
This closely resembles H: venulosum, Seem., and is not always readily distin-
guishable from that species. The best marks of this appear to me to be the promi- :
nence of the basal nerves and the fewness of the other main nerves (only 2 or 3
pairs); and the longer narrower panicles which, up to the pedicels of the:
flowers, are covered with minute stellate hairs. The majority of the specimens of’
H. venulosum, Seem. have glabrous panicles, but those of var. macrophylla are
hairy, and the leaves have many more lateral nerves.
11. Heprrapteurum ventLosum, Seem. Rev. Hed. 44. Scandent;
young branches rather slender, glabrous. Leaves digitate; common’
petiole slender, terete, glabrous, 4-6 in. long; leaflets thinly coriaceous,
oblanceolate-elliptic to oblong-lanceolate, shortly acuminate, narrowed
to the base, the edges entire; both surfaces shining, glabrous, conspicu-
ously and minutely reticulate; length 3-7 in., breadth 1:25-2-75 in.;
petiolules unequal, slender, varying from *5-1°5 in. long in the same leaf.
Panicle varying in length bnt usually shorter than the leaves, terminal,
glabrous or occasionally slightly pubescent, (stellate-pubescent in var.
macrophylla), dividing into several narrow branches bearing short
branchlets each terminating in an umbel of 10-15 depressed-clobular
flowers ‘] in. in diam.; their pedicels unequal, slender, *15--3 in. im
length. I’ruit ovoid, bluntly 5-ridged, 5-celled, glabrous, ‘15 in. long,
yellow when ripe. Clarke in Hook. fil. Flor. Br. Ind. Il, 729; Brand.
For. Flor. 294; Kurz For. Flor. I, 538. Paratropia venulosa, W. & A:
Prodr. 377; Wight Ill. t. 118. Hedera veuosa, Wall. Cat. 4923. H.
terebinthacea, Wall. Cat. 4920, (partly,). Aralia digitata, Roxb. Hort.
Beng. 22; Flor. Ind. II, 107.
ANDAMAN Istanps ; King’s Collectors. Matacca; Derry.
This species, so common from the base of the Eastern Himalaya southwards
1898.] G. King—Materials for a Flora of the Malayan Peninsula. 53:
through Assam to Burma, is replaced in the Malayan Peninsula by the closely
allied species H. ellipticwm, which differs from this as noted under that species.,
Mr. Clarke, in Flor. Br. India, however, reduces H. ellipticum to H. venulosum, a:
course in favour of which there is a good deal to be said.
12. HepraPLevruM .ELLIpTICcUM, Seem, Rev. Hed. 43. Scandent;.
young branches with pale brown glabrous bark. Leaves digitate; the
common petiole 4-6 in. long, glabrous ; leaflets 5- 7, coriaceous, more-
or less broadly elliptic, sometimes elliptic- rotund, apiculate, subapicu-.
late or obtuse, the base rounded or sub-cuneate; the edges entire,
revolute when dry ; both surfaces glabrous and rather dull when dry ;.
main nerves 4 or 5 pairs, the reticulations wide, inconspicuous ; length
2-5-7 in,, breadth 1°75-4 in. ; petiolules uequal, *75-1'5 in., that of the
middle leaflet 2 in. Panicle about as long as the leaves, terminal,
glabrous, lax, open, the branches long, spreading and bearing few-:
flowered umbels on long slender peduncles ; flowers globular-ovoid, ‘1 in,
Jong or less, on slender pedicels ‘15-2 in. long. Fruit oblong, yellowish,
with 5 ridges and 5 cells, Paratropia elliptica, Maiq. in Bonplandia
1856, p. 1388; Flor. Ind. Bat. J, Pt. I, p.756; in Ann. Mus. Luged.
Bat. I, 20; Sciadophyllum ellipticum, Blume Bijdr. 878; DC. Prodr.
IV, 260.
Sincapore; Ridley 5839, 6399: Matnacca; Derry 1187, 1215.
Penane ; Ourtis 972. Perax; Scortechini; Wray 2020, 2136; King’s
Collector 2541, 4733, 10375, 10534. ANnpaman ann Nicopar Istanps;
King’s Collector.
This Raeauthios H. venulosum, Seem ; but the Eatne Rem on the leaves of this
are wider and less distinct than in that; and the panicles of this have spreading,
quite glabrous, lax branches.
13. HEPrAPLEURUM SCANDENS, Seem. Rev. Hed. 43. A slender
creeper 3-5 in. long, the stem pale and corky. Leaves small, digitate ;
common petiole 1-2°5 in. long, slender ; leaflets 3-5, thinly coriaceons,
lanceolate, caudate-acuminate, tapering much to the base, the edges
entire and somewhat recurved ; both surfaces glabrous, the upper smooth
and. ee the lower dull and reticulate; length 1°5-3°5 in., breadth
“5-8 in.; petiolules subequal, ‘2-25 in, long.. Panicles longer than the
leaves, aie ace. 4-8 in. long; the branches spreading horizontally,
simple, each ending in an‘umbel of flowers on a slender pedicel nearly
‘75 in. long; buds globular,*1 in. in diam. JL’ruit elliptic, boldly 5-
ridged, 5-celled, ‘1 in. long. Paratropia scandens, Miq. in Bonplandia
1856, p. 138; Flor. Ind. Bat. I, Pt. I. 757. P. brachybotrya, Mig. Flor.
Ind. Bat. I, Pt. I, 755. Sciadophyllum scandens, Blume Bijdr. 878.
Perak; Wray 1844, 2401, 2880; Curtis 2687; Scortechini 218,
1352 ; King’s Collector 4304. Distris. Java, Sumatra.
A very slender glabrous species, at once distinguished by its small lanceolate,
caudate-acuminate, digitate leaflets.
54 G. King— Materials for a Flora of the Malayan Peninsula. (No. 1,
14. HeprarpLEURUM AFFINE, new species. A shrub 6-8 feet high,
semi-scandent ; young branches as thick as a goose-quill, shining, gla-
brous; common petiole 2-4 in. long, slender, the stipule bout °65 in., both
glabrous ; leaflets 5 or 6, thinly coriaceous, ovate-lanceolate to elliptic,
acuminate, the base sometimes narrowed but always rounded, the edges
entire ; length 2°25-4 in., breadth 1:25-1°75 in.; petiolules somewhat
unequal, °75-1°5 in. long. Panicle terminal, when young furfuraceously
puberulous towards the base, ultimately quite glabrous everywhere,
25-5 in. long and almost as broad ; the main branches 3 or 4, spreading
and bearing, in pairs or whorls of 3, six to twelve ultimate ebracteate
branches ‘5-75 in. long, each terminating in an umbel of 7-10 pedicel-
late broadly ovate flowers nearly ‘2 in. long, the pedicels ‘2 in. long.
Fruit broadly ovate, apiculate, boldly 6-ridged, 6-celled, glabrous, nearly
°25 in. long.
Perak; at elevations of from 3000 to 5000 feet. Scortechini 333,
486 ; King’s Collector 3827; Wray 4121.
This in many respects resembles H. ellipticum, but has larger flowers and its
fruit is on shorter pedicels.
15. Heprarptevrum Hunter, new species. A small tree, 10-15
feet high ; branches stout, rugulose, deciduously pubescent, Leaves large,
digitate; the common petiole terete, glabrous, 10-24 in. long; leaflets
7-11, coriaceous, oblong or oblong-elliptic, occasionally somewhat broader
in the upper than in the lower half, shortly and abruptly acuminate,
slightly narrowed to the rounded base; both surfaces glabrous, not
reticulate, the midrib prominent ; main nerves 5-8 pairs, distant, slightly
curved and ascending, prominent on the lower surface when dry,,
obsolete on the upper; length 4-12 in., breadth 1°5-3°25 in.; petiolules
1:25-3°5 in., slender, glabrous. Panicles 9-12 inches long, terminal,
several together, long and narrow, with short horizontal slender branches
from ‘5 to 1 in. long, bearing terminal umbels of 8-12 small 6-merous
sub-globular flowers ‘1 in. in diam. Fruit elliptic, 6-ridged, 6-celled,
crowned by the 6 short distinct styles, glabrous, pedicels °25—-3 in. long.
Sincapore; Ridley 447, 4591, 6012; Wray 2323; King’s Collector
3048. Jonore; King and Hullett. |
A species resembling H. dvaricata, Miq.; but having leaflets with fewer nerves
and no reticulations, much longer panicles and narrower fruit. It is allied also to
H. longifolium, Seem., but the leaflets of that species have greatly more numerous.
‘main nerves, and the panicles are densely clothed with broad scale-like hairs and
have longer lateral branchlets.
16. HuprapLevrum Rrpury1, new species. Scandent; young
ranches stout, glabrous. Leaves digitate; common petiole terete,
glabrous, 5-10 in. long ; leaflets 5, very coriaceous, oblong, oblong-elliptic
or oblanceolate-oblong, acute, slightly narrowed at the base; the edges
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 55
entire, very slightly revolute when dry; both surfaces glabrous, the
reticulations faint when dry ; main nerves very slightly prominent on
the lower surface, about 8 pairs, spreading, the intermediate nerves
almost as conspicuous ; length 4—6°5 in., breadth *2-2°75 in.; petiolules
unequal, °75-2°25 in. Panicle terminal, glabrous, 5 or 6 in. long, with
several spreading branches; the branchlets few, about ‘5 in. long, each
bearing an umbel of 10-20 globular flowers ‘15 in. in diam. Calyz-
tube short, widely campanulate, the limb narrowed and truncate.
Petals 5, elliptic, glabrous. Fruit ovoid, somewhat succulent, smooth,
faintly 6-ridged, ‘25 in. long, 6-celled.
Sincapore ; Ridley 6336 and perhaps also 1890a.
17. HepraPLeURUM NERVOSUM, new species. A small shrub;
branches with glabrous bark pale brown when dry. Leaves digitate;
the common petiole terete, 2-2°25 in. long ; leaflets 6, very coriaceous,
lanceolate, acute, the base narrowed; the edges entire, much recurved
when dry; both surfaces glabrous, the upper shining, the lower dull;
main nerves 7-10 pairs, straight, sub-horizontal, very prominent on the
lower surface and deeply impressed on tle upper when dry, length
1-5-2°5 in., breadth 5-9 in.; petiolules unequal, the middle two about
‘75 in. long, the others about half as long. Panicle terminal, from
1-5-2 in. long, rusty-puberulous at first, afterwards glabrous, branches
about 2, spreading, with short bracteoles at the base and above it, each
ending in an umbel of 8-10 oblong pedicelled glabrous flowers ‘2 in.
long, their pedicels *1-15 in. long. Calya-tube cylindric-campanulate,
the limb truncate and entire. Petals narrowly triangular. Fruit
rotund-ovoid, boldly 6-ridged, 6-celled, glabrous, °3 in. long.
PERAK, on Gunong Chabong; Scortechini.
A very distinct small species with rather large flowers and fruit for the genus,
and prominently-nerved very coriaceous leaflets.
18. HeptapLeurum WRrayl, new species. A small tree; young
branches as thick as a swan’s quill, furfuraceous. Leaves digitate;
common petiole 6-9 in. long, slender, glabrous; leatlets 7-9, thinly
coriaceous, elliptic, abruptly shortly and sharply acuminate, the base
rounded, the edges with shallow distant sharp serrations ; upper surface
glabrous, the lower glaucous and with scattered minute stellate hairy
scales; main nerves 7 or 9 pairs, prominent beneath, length 3-5 in.,
breadth 1°75-2°25 in.; petiolules unequal, 1°5-2°5 in. long. Panicle
terminal, longer than the leaves, furfuraceous stellate-pubescent, bearing
a few rather distant, horizontal or deflexed many-flowered racemes,
Flowers 15 in. in diam., their pedicels *2 in. long. Oulya-tube funnel-
shaped, its mouth with 5 short triangular spreading teeth. Petals 5,
elliptic-oblong, glabrous, reflexed. Fruit globular, prominently 5-ridged,
56 GG. King—Materials for a Flora of the Malayan Peninsula’ [No. 1,
‘crowned by’ the long confuent column of styles, elabrous, 5- celled, “15
in. in diam.
Prrak; on Gunong Brumber Paha at an siaiatise of about 7000
feet, Wray 1585.
A very distinct species, at once distinguishable by its racemose panicles, and
leaflets glaucous on the lower surface and with serrate edges.
19. - H&PTAPLEURUM BITERNATUM, Clarke in Hook. fil. Flor. Br. Ind.
II, 735. A shrub several feet high; the young shoots. and the under
surfaces of the leaves deciduously stellate-pubescent. Leaves digitately
decompound or twice pinnate, with ternate leaflets at each node of the
rachis; leaflets coriaceous, narrowly-oblong, acuminate, entire, the
pase slightly narrowed and rounded ; length 2-3°5 in., breadth *5-75 in.,
petiolules *1 in. long or less, that of the terminal leaflet -4 in.; both
surfaces minutely reticuiate and shining, the upper glabrous, the lower
with deciduous stellate pubescence. Panicle 8 in. long, but only abont
15 in. across ; the branches little- divided, stellate-hairy ; bracts decidu-
ous, pedicels 2 in. long.
Matacca; Maingay (Kew Distrib.) 684.
Known only by Maingay’s fragmentary specimens.
20. HepraAPLEURUM HETEROPHYLLUM, Seem. Rev. Hed. 40. A ‘bush
‘or small tree 8-12 feet high. Lower leaves large, ternately decompound,
24 in. across ; common petiole 12—24 in. long; the upper leaves smaller
and only twice digitate ; the leaflets in all 3 to 5 on each petiolule, thinly
coriaceous, variable in shape, oblong-lanceolate ‘to elliptic or broadly
ovate, shortly acuminate, narrowed or rounded at the base; the edges
entire, rarely with 1 or-2 teeth near the apex ; both sernels minutely
reticulate and glabrous, the lower minutely dotted; length 2-5-7 or
even 9 in., breadth 1-2°25 in.; petiolules of the Tones leaflets *1="25 j ins,
that of fig terminal twice as fos Panicles with deciduous pale stellate
‘pubescence, solitary or several together, 10-15 in. long, and only 1°5-2
in. across; the branches horizontal, slender, each ending in an umbel
of flowers on slender pedicels, the flowers bearing fertile pistils smaller
than those with fertile stamens. Irwit narrowly oblong, boldly 5-ribbed,
glabrous, 5-celled, nearly *25 in. long, claret-coloured when ripe.
‘Clarke in Hook. fil. Flor. Br. Ind. II, 731. Hedera heterophylla, Wall.
Cat. 4919; G. Don. Gen. Syst. HT, 394. Paratropia heteropha ylla, Presl
‘Epimel. Bot. 250; Miq. Flor. Ind. Bat. 1, Pt. J, 761.
Penance; Wallich, Curtis 241, 2301 and possibly: 1950. PORAK
Scortechini 145, 664; King’s Collector 718, 2688, 8640, 8769.
21. Kitsaolin ie i aoe Currtsti, new species. - A large shrub. Lower
leaves bipinnate, the upper trifoliolate ; common petioles of both about
8 in. long; leaflets thinly coriaceous, oblong-elliptic, sometimes slightly
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 57
obovate, the apex shortly acuminate, the edges entire in the lower half
but with a few unequal scanty coarse sharp teeth in the upper half; the
base slightly narrowed, sometimes oblique; both surfaces quite glabrous,
shining and finely reticulate when dry; length 3-6 in., breadth 1°5-2°5
in.; petiolules unequal, the lateral -1 in. long or absent, the terminal
35-8 in. Panicle terminal, shorter than the leaf-petioles, with several
rusty stellate-tomentose bracts °75 in. long at its base, 2-branched; the
branches narrow, sparsely covered with scurfy pubescence; the lateral
branchlets about *75 in. long, slender, each bearing at its apex a crowded
umbel of from 10-20 oblong flowers on pedicels *15-—3 in. long. Fruit
oblong, boldly 5-ribbed, crowned by the conical disc bearing 5 small
rounded stigmas at its corners, 5-celled.
Penane; at Pulo Boetong, 1950.
I have seen only two specimens (and they are both of the same gathering) of
this very distinct species.
4. Trevesta, Vis.
Shrubs or small trees, prickly or unarmed, glabrous or stellate-
hairy. Leaves palmifid or palmisect; petioles often united by a wing
at their base; stipules united within the petiole, or obsolete. Flowers
polygamous, large for the Order; umbels panicled ; pedicels not jointed
under the flower; bracts small or 0. Calyz-margin entire or toothed.
Petals 8-12, valvate, somewhat thick, often cohering as a cap in the
fertile flowers. Stamens equal in number to the petals. Ovary with as
many cells as the petals; styles connate into a short column. Fruit
ovoid, large for the Order. Seeds compressed; albumen uniform.
Distris. Species about 10; natives of Hastern India, Malaya and
Polynesia.
TREVESIA PALMATA, Vis. in Mem. Acad. Torino, Ser. 2, 1V, 262, with
fig. A-small single-stemmed tree 10-25 feet high; young shoots ferru-
ginous-pubescent and very prickly. Leaves coriaceous, large (12-24 in.
in diam.), rotund in general outline, deeply palmatifid; or, in young
shoots, palmatisect, widely cordate at the base, the lobes acuminate, their
edges serrate or sometimes lobulate; glabrous when adult or with a few
small rufous stellate hairs on the lower surface; the lobules contracted
in the middle to a pseudo-petiolule (in var. cheirantha); petiole often
prickly, 6 to 20in. long. Panicles 12-30 in. long, the branches spreading,
when young clothed with reddish-brown tomentum ; bracts oblong, 1 in,
long, usually deciduous; pedicels 1-15 in. long. Ilower-buds ‘12 in.
in diam. Fruit ovoid-rotund, the ribs not prominent, crowned by
the stout style, fleshy, ‘5 in.in diam. Seem. Rev. Hed. 77; Kurz For,
Flora Burma, I, 539; Clarke in Flor. Br. Ind. II, 732; Boerlage in Ann.
5. i, 8
58 G. King— Materials for a Flora of the Malayan Peninsula. [No.1],
Jard. Bot. Buitenzorg VI, 108. Gastonia palmata, Roxb. Hort. Beng.
33; Flor. Ind. II, 407; Lindl. in Bot. Reg. t. 894. Giilibertia palmata,
DC. Prodr. IV, 256. Hedera ferruginea and H. palmata, Wall. Cat. 4909
and 4910 (partly). Brassaiopsis confluens, Seem. Rev. Hed. 18 (as to the
leaves). Aralia dubia, Spreng. Syst. Veg. IV, 2, p. 125.
Perak ; Scortechint ; King’s Collector 4436, 6715.
Var. cheirantha, Clarke in Flor. Br. Ind. II, 732; lamine of the
lobes cut away in the middle so as to expose the midrib and form a ~
pseudo-petiolule. Hedera ? cheirantha, Jack in Wall. Cat. 4925; Wall.
Cat. 4910 in part.
Perak ; Wray 2822; King’s Collector 2308 ; Scortechini 344.
5. Denpropanax, Decne and Planch.
Unarmed glabrous trees or shrubs. Leaves simple, entire, (palmate-
ly 3-5-lobed on young shoots). Umbels solitary or in small panicles ;
bracts small or none; pedicels not jointed under the flower. Limb
of the calyx entire or 5-toothed. Petals 5, free, valvate, rather thick.
Stamens 5. Styles united into a column at the base, free at the apex.
Fruit globose or ellipsoid, succulent, distinctly or obscurely 5-ribbed.
Seeds compressed ; albumen uniform. Dusrrir. about 12 species mostly
tropical American ; one Japanese ; one Indo-Chinese.
DenpDROPANAX MAINGAYI, new species. A shrub; young branches with
corky bark, pale-brown when dry, all parts except the umbels glabrous.
Leaves alternate or sub-opposite, thinly coriaceous, oblong-ovate, oblong
or lanceolate, acute ; the base rounded, sometimes slightly narrowed; the
edges entire and slightly recurved when dry; both surfaces glabrous,
dull, the midrib prominent on the lower and sending off near its base two
boid curving nerves running at: some distance from the margin to the
apex and, above the origin of these, 7-8 pairs of faint horizontal nerves ;
length 2-3°25 in., breadth 1-1-75 in., petioles varying from °25-1°5 in,
in length. Umbel simple, terminal, its pedicel °35—5 in. long; flowers
8-12, oblong, pedicelled, -2 in. Jong, their pedicels °25--4 in. long.
Calyx cylindric-campanulate, puberulous, its mouth with 5 sharp
triangular teeth. Petals broadly lanceolate, acute, quite free. Fruit
globular, succulent, glabrons, °3 in. in diam. JD. parviflorwm, Clarke in
Hook. fil. Flor. Br. Ind. (not of Bentham).
Maxacca; Maingay (Kew Distrib.) 682; Grifith 2685-1. Purax ;
Scortechini. 308.
This Dendropanax, found in Malacca and Perak, does not agree with specimens
of D. parviflorum, Benth., collected in Hongkong. It appears to me to be a distinct
species hitherto un-named,
1898.) G. King— Materials for a Flora of the Malayan Peninsula. 59
6. ARTHROPHYLLUM, Blume.
Shrubs or small trees, unarmed. Leaves glabrous or sub-glabrou
the lower large and compoundly pinnate, the uppermost opposite and
simple, the intermediate 3-foliolate; leaflets easily separable from the
rachis; stipules forming a ligule within the petiole. Inflorescence a
terminal compound umbel, the terminal umbellules peduncled; bracts
very small; pedicel not jointed under the flower. Calyx-teeth 5, small.
Petals 5, valvate. Stamens 5. Ovary 1-celled, 1-ovuled; style short,
simple. Fruit (in the Indian species) ovoid, not angular. Seed sub-
globose; albumen ruminate. Disrris. Species 3, Malayan.
The l-celled ovary is anomalous in this Order, and this genus was excluded
from Araliacie by Seemann.
Lower leaves pinnately decompound ye we L. A. diversifolium.
Lower leaves simply pinnate, or at most bipinnate .. 2. A. pinnatum.
1, ARTHROPHYLLUM DiversIFoLIoM, Blume Bijdr. 879. A shrub or
small tree, all parts except the umbels glabrous. Lower leaves large,
pinnately decompound, with pairs of opposite leaflets at the main
divisions, the upper leaves smaller and simply pinnate, and the upper-
most of all trifoliolate or simple ; leaflets coriaceous, oblong or elliptic,
acute, narrowed at the base, entire; length 1°5-3 in., breadth 1°25-2°25
in.; petiolules ‘25-45 in., slender. Inflorescence a compound umbel ;
the peduncles of the ultimate umbels unequal, covered with warm brown
deciduous stellate tomentum ; lengthening in fruit to *5-1°5 in.; pedicels
*25 in. long; fruit ovoid-globose, not ridged, crowned by the conical
disc, glabrous. Clarke in Hook. fil. Flor. Br. Ind. II, 734; Mig. Flor.
Ind. Bat. I, Pt. I, 767. A. javanicum, Blume Bijdr. 879; DC. Prodr.
IV, 266; Kurz For. Flor. I, 540. A. ellipticum, Blume and DC. 1. c.
A. Blumeanum, Zoll. & Mor. Verz. 41; Mig. l. c. 1, 768. A. ovali-
folium, Jungh. & De Vriese in Miq. 1. c. t. 14. Panaz polycarpum, Wall.
Cat. 4930. P. Jackianum, Wall. Cat. 4931. Hedera Jackiana, G. Don
Gen. Syst. III, 394. H.? ovata, Wall. Cat. 4911. Hupteron, sp. nov.
Kurz Andam. Rep. Suppl. B 9.
Manacea ; Griffith (Kew Distrib. 2675). Sincarporr; Anderson 48,
185; Hullett 351, 393; Ridley, 5838. Prnana; Curtis 781. Prrak
Wray 2012, 3063; King’s Collector and Scortechini, many numbers,
ANDAMAN Istanps; Kurz.
2. ARTHROPHYLLUM PINNATUM, Clarke in Hook. fil. Flor. Br. Ind.
If, 734. A bush; all parts glabrous. Leaves pinnate, rarely bipinnate
or simple; the pinnate ones 12-18 in. long and with 5-17 leaflets ;
leaflets varying in size, coriaceous, lanceolate, caudate-acuminate, taper-
ing to the base; the edges entire, glabrous; length 1:25-4 in., breadth
‘4-1 in., petiolule absent or only ‘1 in. in length. Umbels with few
60 G. King— Materials for a Flora of the Malayan Peninsula. [No.1,
umbellules, their peduncles 1°5-2 in. long; pedicels glabrous or with
rusty stellate deciduous pubescence; fruit sub-globose, shining, °15 in.
in diam. Panax pinnatum, Lamk. Dict. I, 715; DC. Prodr. IV, 254;
Wall. Cat. 9057. P. secunda, Schultz Syst. VI, 215. Nothopanaz ?
pinnatum, Miq. Flor. Ind. Bat. I, Pt. I, 766.
PenanG; Wallich, Maingay (Kew Distrib.) 679. Matacca; Main-
gay 677; Griffith (Kew Distrib.) 2676; Ridley 3224. Perak; Wray
330, 1475; Scortechini 352.
7. WaArDENIA, new genus.
A miniature tree with prickly stem, otherwise unarmed. Leaves
coriaceous, simple, on long terete petioles expanded at the base into
a short sheath with 2 minute stipules on its inner surface. Inflores-
cence a terminal shortly-branched compound umbel. Flowers herma-
phrodite. Calyx-twbe narrowly campanulate, its limb with 5, small,
spreading teeth. Petals 5, calyptrate, their edges slightly infolded,
vaivate below, slightly imbricate near the apex; the midribs prominent
on the inner surface. Stamens 5, alternate with the petals; the fila-
ments short, straight; the anthers versatile; the cells linear, quite
separate from each other, each united by. its middle to the tip of
the filament. Disc large, fleshy, convex, covering the whole of the
apex of the ovary, slightly 5-lobed. Styles united to form a short
thick column without any distinct stigmatic enlargement; ovary l-
celled, with 2 parallel pendulous ovules. Frwit 2-celled, by the form-
ation of a dissepiment not present in the ovary, 2-seeded ; seeds com-
pressed. A single species.
This genus is allied to Arthrophyllum ; but its ovaries, although one-celled, have
two pendulous ovules. The fruit, however, is two-celled, by the subsequent form-
ation of a dissepiment, and is 2-seeded. The leaves moreover are all simple. The
seeds of the few specimens which I have seen are quite young and the nature of
the albumen cannot be made out. I have named the genus in honour of my friend
Brigade-Surgeon Lt.-Colonel C. J. H. Warden, a distinguished pharm and
one of the authors of the Pharmacographia Indica.
WARDENIA SIMPLEX, King. A shrub 6-8 in. high, deciduously
rufous-pubescent towards the apex, prickly near the base. Leaves
simple, elliptic, tapering gradually to the shortly acuminate apex, not
narrowed to the slightly cordate base; both surfaces bearing minute
scattered rusty stellate hairs; length 8-15 in., breadth 3°5-7 in.,
petiole 5-10 in. Flower buds ‘1 in. in diam., conical; pedicels *6—9
in. long, slender, rusty-pubescent, the umbels 10-20-flowered. Calyx
slightly rusty-pubescent. Petals glabrous. Fruit elliptic-globose, sub-
glabrous, *2 in, long, crowned by the calyx and by the slender conic
stylar column.
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 62
Perak; near Ulu Kerling, King’s Collector in flower during March
only once collected.
8. HeteEropanax, Seem.
A small unarmed tree. Leaves glabrous or nearly so, very large,
pinnately decompound, stipules not prominent. Panicles large, the
branches bearing umbels, stellate-hairy ; bracts of umbels small, ovate,
obtuse, persistent; pedicels not jointed to the flowers. Flowers polyga-
mous, the female flowers most numerous in the terminal umbel. Calyz-
limb minutely toothed. Petals 5, valvate. Stamens 5, filaments filiform,
anthers ovate. Disk nearly flat; ovary 2-celled; styles 2, slender, free
from the base, spreading, the stigmas sub-terminal. Fruit laterally
compressed, coriaceous, almost didymous, 2-seeded. Seeds compressed,
albumen ruminate. Species 1 or 2; Indo-Chinese.
HETEROPANAX FRAGRANS, Seem. Rev. Hed. 73. A tree 40-60 feet
high ; all parts glabrous. Leaves large, the lower often 3 feet across,
pinnately decompound, tke pinnae with a pair of opposite leaflets at
their forks; leaflets ovate or ovate-oblong, acute or acuminate; the
base slightly oblique, not narrowed; variable in size (2°5-5 in. long,
and 15-25 in. broad) ; petiolules of lateral leaflets ‘1-2 in. long, that
of the terminal one ‘8-1 in. Panicles terminal, longer than the leaves ;
flowers small, whitish-tomentose, in small condensed umbels, their
pedicels under ‘25 in. long; fruit compressed, sub-reniform, sub-
glabrous, slightly glaucous, ‘35 in. across. Brandis For. Flora 249
Kurz For. Flora Burma, J, 541; Clarke in Hook. fil. Flor. Br. Ind. II,
734. Panax fragrans, Roxb. Hort. Beng. 21; Flor. Ind. II, 76; Wall.
Cat. 4929; DC. Prodr. IV, 254 (excl. syn. of Don).
Anpaman Istanps; King’s Collector. Disvris. Brit. India, Java,
China.
9. Brassatopsis, Decne. & Planch.
Large shrubs or trees, glabrous or tomentose, armed or not. Leaves
digitate or palmate or angled; stipules connate within the petiole, not
prominent. Umbels in large compound panicles, young parts at least
stellately tomentose ; bracts not large, often persistent; pedicels rising
from a dense cluster of persistent bracteoles, not jointed under the
often polygamous flowers. Calyx 5-toothed. Petuls 5, valvate. Stamens
5. Ovary 2-celled; styles 2, united, long or short. Fruit broadly
globose or turbinate, 2- or (by abortion) 1-seeded. Seed not com-
pressed; albumen ruminated. Drsrris. Species 11 ; Northern Brit.
India to Java.
Brassalopsis pALMATA, Kurz in Journ. As. Soc. Beng. XXXIX
62 G. King— Materials for a Flora of the Malayan Peninsula. [No. 1,
(1870) Pt. Il, 77. A small sparingly prickly tree with simple stem
slightly branched near the top; young shoots covered with tawny or
rusty scurfy tomentum. eaves crowded at the ends of the stem and
branches, large, 9-15 in. across, coriaceous, rotund in general outline,
cordate at the base, palmately lobed about half or more than half way
down; the lobes 5-9, oblong, or sometimes sinuate towards the base, acumi-
nate, serrate; upper surface glabrous, lower sub-glabrous; the petiole
10-20 in. long, without prickles, scurfy-tomeutose when young, ultimately
glabrous. Inflorescence rusty-tomentose, terminal, panicled, the ultimate
branches bearing many-flowered bracteolate umbels; flowers ‘15 in,
across, their pedicels ‘5-7 in. long; rim of calyx narrow, irregularly
toothed. Fruit broadly elliptic or turbinate, terete, ‘3 in. long, as large
as a pea, crowned by the slender column of connate styles; cocci 1 or 2,
with chartaceous pericarp, l-seeded. Kurz For. Flor. Burma J, 537;
Clarke in Hook. fil. Flor. Br. Ind. II, 735. Panax pulmatum, Roxb.
Hort. Beng. 21; Flor. Ind. Il, 74. Hedera polycantha, Wall. Pl. As. Rar.
iI, t. 190; Cat. 4907 B.
This species is closely allied to B. Hainla, Seem., from which it
differs chiefly in having leaves with deeper narrower more serrate
lobes, and also in having rusty instead of pale tomentum on the
young shoots and inflorescence. Thetwo are in my opinion rather too
closely allied to be kept distinct as species. In his distribution, Wallich
issued both under the name Hedera polycantha and the number 4907. |
Purak ; Scortechinit 17, 1146; Ridley 3018; King’s Collector 2598.
Distris. Brit. India, along the base of the Himalaya; Assam and
Burma,
Var. andamanica, lobes of leaves obovate-oblong, the edges almost
entire; inflorescence a narrow panicle nearly as long as the leaves.
Araliopsis andamanica, Kurz in Andaman Report, App. B, 9.
ANDAMAN Istanns ; Kurz, King’s Collectors.
10. Heprropsis, C. B. Clarke.
A glabrous unarmed tree. Leaves 1-3-foliolate; leaflets lanceolate,
denticulate or nearly entire; base of petiole much dilated; stipules
inconspicuous. Umbels panicled; bracts and bracteoles deciduous .
pedicels jointed close under the flowers. Calyx margined, somewhat
prominently 5-toothed. Petals 5, valvate. Stamens 5. Ovary 5-celled ;
styles connate. Fruit berried, large, sub-globose, crowned by the stout
persistent style. Seeds 5-4; albumen ruminated.
Heperopsis Matneayi, Clarke in Hook. fil. Flor. Br. India, II, 739.
Leaflets of the compound leaves membranous, ovate-lanceolate, acumi-
nate, narrowed at the base, nerves faint, length 3-5 in., breadth 1°5-2
1898.] G. King— Materials for a Flora of the Malayan Peninsula. 63
in., petiolules ‘1-2 in. Simple leaves as long as 8'5 in. and about 4 in.
broad; petiole 1-2 inches. Inflorescence a panicle of umbels with
puberulous peduncles 1 in. or more long. Calya-tube sub-globular,
puberulous. Petals ovate-lanceolate, spreading, ‘15 in. long, Fruit (un-
ripe) more than ‘5 in. long, including the conical disc and persistent
style.
Matacca ; Maingay (Kew Distrib.) 683.
This plant has not been collected since Maingay’s time, and it is known only
from his fragmentary specimens.
11., Touprpantavs,. H, f. &.T.
A large glabrous shrub, at first erect but afterwards a lofty
climber. Leaves digitate ; leaflets glabrous, leathery, entire; stipules
connate within the petiole. Inflorescence a compound umbel or small
panicle ; pedicels thick, not jointed under the flowers. Calyx-margin
obsolete. Petals closely connate, falling offina cap. Stamens very many,
in two or several series. Ovarian cells and stigmas very numerous;
the latter sessile, radiating, crowded but not connate. Fruit globose,
depressed, succulent.
T'UPIDANTHUS CALYPTRATUS, Hook. fil. and Thoms. in Bot. Mag. t.
4908. Leaflets 7-9, oblong or oblong-obovate, acute or blunt, 4-7 in.
long, and 1:75-3'5 in. broad, the petiolules 1-2 in.; the common
petiole 6-15 in. Inflorescence umbellate, 3-4-branched; the branches
stout, short and with large coriaceous bracts at their bases; the
ultimate umbels with 3-7 pedicellate flowers nearly 1 in. across ; calyx-
tube glabrous, thickly coriaceous. Stamens 50-70, crowded. Fruit
sub-globose, succulent, 1°25-1'5 in. in diam. when ripe. Seem, Rev.
Hed, 6; Clarke in Hook. fil, Flor.Br. Ind. IT, 740.
Perak; on Gunong Ulu Sungei, elevat. 4500 feet; Wray 1594.
Distrts. Burma; Khasia Hills and probably Java.
Mr. Wray’s specimens were collected at an elevation much higher than this
species ever ascends to in British India. They have smaller leaves with blunt
leaflets, but are otherwise indistinguishable from the British Indian plant.
64 ¥. Finn— Plumage of Honeysucker, [ No},
Note on the Seasonal change of Plumage in the males of the Purple
Honeysucker ( Arachnechthra asiatica) and of an analogous American
bird (Coereba cyanea).—By F, Finn, B.A., F.Z.8., Deputy Super-
intendent, Indian Museum.
[Received and Read January 4th, 1898.]
Dr. Jerdon in his “ Birds of India” (Vol I, p.870) and Captain
Shelley, in his Monograph of the Crnnyridae, agree in assigning to the
male of our common Purple Honeysucker (Arachnecthra astatica) besides
its characteristic dress, a plumage much resembling that of the female,
but marked with a broad purple streak down the ventral surface.
Dr H. Gadow, however, in the British Museum Catalogue volume
(IX, p. 58), dealing with these birds, ignores this change of plumage;
and Mr. Oates, in his “ Birds of British Burmah” (Vol. I, p. 322),
states that the change does not take place in that country, “for fuall-
plumaged males may be obtained all the year round.” He believes also
that the young males of this species are clothed in female plumage all
through their first winter, and thinks that the abundance of such has
probably given rise to the belief in a change of plumage.
With all due deference to the opinion of so excellent an ornithologist
as Mr. Oates, however, I venture to suggest that he is wrong, and that
the authors previously cited are right, with respect to this change of
plumage, at any rate in Indian examples.
In the first place, the presence of full-plumaged birds all the year
round is of very little weight in disproving this change. Marked in-
dividual variations occur in the period of change of plumage by birds
which possess more than one dress, and specimens of such species may
be found in more or less full-plumage and undress at the same date,
as I have myself seen in Ducks and Dabchicks.
This consideration disposes, I think, of Mr. Oates’ first argument,
but I have better evidence to bring forward.
About the middle of July last year (1897) in view of my approaching
visit to England on leave, I procured a number of Honeysuckers in
the hope of being able to take some alive to the London Zoological
Gardens, where such birds have never previously been exhibited. All
the birds I kept, with one exception, were Arachnecthra zeylonica, but
I had, and brought home safely, one male specimen of the species I am
now considering.
This bird, when I got it, was in heavy moult, and mostly purple
in colour, but to my great surprise (I had taken it fora young male
assuming full-plumage) it gradually lost this hue, and by the time I
1898. | F. Finn — Plumage of Honeysucker. 65
started for England, in the first week in August, it was in the non-
breeding dress, brown above and yellow below, with the median purple
streak, but still retaining the orange axillary tufts.
This specimen, unfortunately, only survived its advent at the
Zoological Gardens for about a fortnight* and I do not know whether
it was preserved ; if it was, it was probably put in spirit, as the moult
had never been properly completed, and so the plumage was in bad
order. It had, however, lived long enough to show that the change
above referred to does really take place; for that captivity could have
so affected the bird as to change the colour of the actually growing
feathers, I am not inclined to admit, and I therefore conclude that the
accounts which give this bird a change of plumage are quite correct.
While on this subject, it seemed to me that I might draw the atten-
tion of ornithologists to a similar change, apparently hitherto unrecord-
ed, in a bird which, though not believed to be allied to our Sunbirds,
and inhabiting the New World, nevertheless in form and habits presents
at least an analogical resemblance to these. I allude to the Yellow-
winged Blue Sugar-bird (Coereba cyanea) of which several specimens
have been exhibited in the London Zoological Society’s Gardens.
During my previous acquaintance with the species there, I had
been struck by the change of plumage that the male appeared to under-
go, and when in England last September, [ found the Society’s single
Specimen, a male which had been acquired as long ago as 1890, actually
passing into the full violet plumage from the undress stage, which had
been olive-green above, and yellowish below, much resembling the
coloration of the female. The tail was black, and the wings yellow and
black, and the legs pink-red, as in the male in full plumage, In fact,
* T ascribe my small measure of success with living Sunbirds to the fact that
I fed them too much on “‘slops”” —~sweetened milk or milk-sop. In addition to some
such food given at first it would, I think, be well to supply crumbled yolk of hard-
boiled egg mixed with powdered sugar, and to-keep them as much on this as possi-
ble, with fruit also. None of the Arachnecthra zeylonica I had survived the voyage
but one, and this died in the train en route from Plymouth to London. I saw this
bird bullying the A. asiatica one occasion at least, and I had previously noticed that
the latter bird appeared somewhat to fear its companions. When all were together
in a big cage in Calcutta it kept almost entirely to one twig in the branch put in this
cage, and was in general less active in its movements than A. zeylonica, though it
seemed less sensitive to cold on the voyage. None of the male A. zeylonica, some
of which were monlting, showed any sign of changing their bright plumage for
a duller one, as suggested by Captain Shelley in his account of the species in the
Monograph above quoted. Neither did they molest each other, while I remember
having had to separate two male specimens of A. asiatica which I had previously
kept, because one was getting so badly bullied by the other,
FF.
on Ir. 9
66 F. Finn— Long-Snouted Whip-Snake. [No. 1,
the bird presented much the same appearance as a skin (21280 in
Register, exhibited on this occasion) in the Museum collection, except
that there were many more violet feathers visible.
The keepers I consulted bore me out as to the regular occurrence
of the change of plumage in the male of this species; and one was of
the opinion that the quills and tail changed also; but this I do not
recollect seeing myself. Unfortunately this bird also soon after died,
and was not preserved.
The existence of this change of colouration in the male of a Coereba
is interesting as tending to confirm the views of those naturalists (Dr.
Sclater and Messrs. Baird, Brewer and Ridgway), who place the
Coerebidae in close connection with the Tanagers, in which group the
male of Pyranga rubra exhibits a similar seasonal alteration of plumage.
Note on the Long-Snouted Whip-Snake (Dryophis mycterizans).—By F.
Finn, B.A., F.Z.8., Deputy Superintendent, Indian Museum.
[ Received and Read, January 5th, 1898.]
A common belief in India accredits the Whip-Snake with the
propensity for deliberately striking at the eye. As this trait is not
alluded to by either Dr. Giinther or Dr. Boulenger in their accounts
of the Indian Reptilia, 1 venture here to bring forward an instance
which shows that the notion above noticed is really correct.
On December Ist, 1897, a bird-catcher, with whom I had pre-
viously had dealings, brought to my quarters two specimens of the
Long-Snouted Whip-Snake (Dryophis mycterizans) for sale. Knowing
them to be harmless, and the vendor having no fear of them, I took
both in my hands and went to show them to a friend who was in an
adjoiing room; the larger one* having meanwhile struck at my hand,
without breaking the skin. As I was exhibiting the snakes, 1 was
rather unpleasantly surprised by finding this large specimen suddenly
dart at my eye, and inflict a bite on it, which, as I had instinctively
closed the threatened organ, only resulted in some small punctures on
the eyelids, which were just sufficient to draw a little blood. The posi-
tion of these, two on the upper, and one on the lower eyelid, sufficiently
shows, I think, the deliberateness of the reptile’s aim. Of course I
suffered no inconvenience from the bite, although on rubbing my eye
* This stuffed skin of this specimen was exhibited, together with a sketch of
the bitten eye made by the Museum artist, to show the position of the tooth-marks.
i,
1898. ] A. Alcock — Careinological Fauna of India. 67
a few hours afterwards, I removed a tooth rather over 35 inch long
from the puncture in the lower eyelid. This, however, after being
examined under the microscope by Dr. Alcock and myself, proved not
to be a grooved one, so that this little experience throws no light on the
possible effects of Dryophis fangs on the human subject; I think,
however, that it may be fairly allowed, in connection with the belief above
mentioned, to upset the reputation for gentleness which Dr. Boulenger
awards to the species.* I may say that I was not holding the snake
roughly or maltreating it in any way, and that when confined afterwards
in a glass case it repeatedly struck at anyone who came near, seeming
to aim particularly at the face, though it soon recognized, apparently,
the futility of attacking glass.
This intelligence in attack was again shown subsequently, when,
having transferred the snake to a large cage of wire gauze, I endeavour-
ed to make it attack a Gecko. This it would not do even when the
lizard was thrown absolutely in its face, darting open-mouthed at me
instead. It similarly refused to bite a handkerchief with which I
teased it, though I have succeeded in getting Dendrophis pictus (a
black Andaman variety) to do this.
Materials for a Carcinological Fauna of India. No.3. The Brachyura
Cyclometopa. PartI. The Family Xanthide.—By A. Aucock, M.B.,
C.M.Z.S., Superintendent of the Indian Museum.
[Received 20th March. Read 6th April, 1898. ]
The family Xanthidsx, as here defined, includes the Cancridx (with-
out Cancer and Pirimela) and the Hriphiide (without Oethra) of Dana’s
system.
It is a family which, as most authors have remarked, it is almost
impossible to divide into groups that shall be at once natural and
sharply defined, owing to the numerous intergradations of form that
exist.
The Indian species of this family, so far as I have been able to
discover, number 153, of which all but the following 14 are represented
in the Indian Museum :—
Carpilodes venosus Hdw., Carpilodes margaritatus A. M. Hdw.,
Lachnopodus rodgerst Stimpson, Lophactzxa fissa Henderson, Lophozozymus
* IT hope this will not be taken as captious criticism of Dr. Boulenger’s work,
for which I entertain the sincerest admiration, especially since I know that gentle-
man to be in the habit of studying reptiles in life when opportunity offers.
F, F.
68 A. Alcock — Carcinological Fauna of India. [No. 1,
cristatus A. M. Edw., Hypocoelus rugosus Henderson, Cycloranthus lineatus
A. M. Edw., Halimede thurstont Henderson, Cymo tuberculatus Ortmann,
Pilumnus labyrinthicus Miers, Actwmnus verrucosus Henderson, Actumnus
nudus A. M. Edw., Heteropanope eucratoides Stimpson, Hurycarcinus
maculatus A. M. Kdw.
The new species described in this paper have almost all been
obtained by the ‘‘ Investigator” and will be figured in the Illustrations
of the Zoology of the Investigator for the year 1899, the original drawings
for which are now in course of preparation,
Tribe CYCLOMETOPA.
Cyclométopes Milne Edwards, Hist. Nat. Crust. I. 264, 363 (part.)
Cancroidea, Dana, U. S. Expl. Exp. Crust. pt. I. p. 142 (part.)
Cyclométopes, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XIV. 1860, p. 183.
Cyclometopa or Cancroidea, Miers, Challenger Brachyura, p. 106 (part.)
Cancroidea Portuninea and Cancroidea Cyclometopa (part.) Ortmann, Zool.
Jahrb., Syst., VII. 1893-94, pp. 65 and 411.
Carapace, almost without exception, broader than long, the antero-
lateral borders generally arched, sometimes very strongly so, the postero-
lateral borders generally convergent, sometimes very strongly so: the
front broadish or broad, horizontal or obliquely deflexed, not rostrate.
Buccal cavern square-cut, commonly broader than long: the palp
of the external maxillipeds articulating at or near the antero-internal
angle of the merus.
Epistome transverse, short fore and aft.
The antennules generally fold nearly transversely.
The abdomen of the male occupies all the space between the last
pair of legs.
Branchie nine pairs; their efferent channels opening on either side
of the palate.
The genital ducts of the male open at the bases of the last pair of
legs.
The Cyclométopes of Milne Edwards includes the genus Mthra
which, following Miers, has been relegated to the Oxyrhyncha in this
series of papers, and excludes the Telphusidx, which by all subsequent
writers have been regarded as true Cyclometopes.
The Cancroidea of Dana includes the genus Acanthocyclus. My
only knowledge of this genus is derived from drawings and descriptions,
which do not as yet satisfy me that Acanthocyclus is more nearly related
to the Cyclometopes than to other groups.
The Cyclometopa of Miers includes not only Acanthocyclus, but,
1898. | A. Aleock— Carcinological Fauna of India. 69
following Claus, the Corystoidea. Now undoubtedly several of the
forms included under the Corystoidea have very close relations with
Cancer and Pirimela; and if Cancer and Pirimela are regarded as typical
Catametopes then such (Corystoid) forms as Atelecyclus and a
tartum may also be classed as Cyclometopes.
In this preliminary paper I prefer not to take Cancer as an ideal
Cyclometope, and to leave the Corystoidea for future consideration.
The Cyclometopa of Ortmann includes the family Parthenopide,
which in this series of papers has, in accordance with the views of
other authors, been considered with the Oxyrhyncha; and also the
Corystoid genera Atelecyclus and Hypopeltarium, the Cancrine affinities
of which have been admitted. I cannot, however, think that the
removal of the Parthenopide from their long approved position, as
Oxyrhynchs showing a connexion between that type and the Cancrine
type, serves any useful purpose.
For the purposes of this paper the Cyclometopa are divided into
the following families :—
I. Cancridx, in which the fold of fe antennules is (oneal
or obliquely longitudinal, and the anterior boundary of the buccal
cavern is somewhat indefinite, being more or less overlapped by the
external maxillipeds.
Of this family, of which Cancer and Pirimela are types, no repre-
sentative is known in the Indian Seas.
II. Xanthidx, in which the fold of the antennules is transverse
or obliquely transverse, and the anterior boundary of the buccal
cavern is raised and sharply defined, so that the external maxillipeds
commonly shut close against it unless they fall short of it.
Ill. Portunide, in which the fifth pair of legs is peculiarly modified
for swimming and usually has the propodite and dactylus singularly
broad thin and paddle-like.
IV. Telphuside, in which the form is Grapsoid, the branchial
regions being much dilated. The members of this family inhabit fresh
water and, sometimes, damp jungle.
The present paper refers to the family Xanthidex.
Family XANTHID A.
Canceriens arqués et quadrilatéres Milne Edwards, Hist. Nat. Crust. I. 369.
Cancridez (exc. Cancrine et ?Polydectine) and Hriphide (exc. Oethrine) Dana,
U.S. Expl. Exp., Crust., pt. I. pp. 147, 228.
Canceriens (exc. Oethra, Cancérides et Pirimélides) A. Milne Edwards, Nouv.
Archiv. du Mus. I. 1863, pp. 177-182.
Cancridz (exc. Cancer), Miers, Challenger Brachyura, p. 106.
Xanthini (exc. Thiidx), Ortmann, Zool. Jahrb. Syst. VII. 1893-94, p, 412.
70 A. Alcock — Carcinological Fauna of India. [No. ],
Carapace transversely oval or transversely hexagonal or subquad-
rate or (rarely) subcircular, but almost always broader than long.
Front broadish or very broad, never in the form of a rostrum. The
fold of the antennules is transverse or obliquely transverse. Antennary
flagella short or slender. Anterior margin of buccal cavern very well
defined, not overlapped by the external maxillipeds. Legs gressorial.
The Xanthide may be divided, according to the character of the
palate emphasized by Dana, into two sections, as follows :—
I. Hyperomerista, in which the efferent branchial channel on either
side is defined by a ridge on either side of the palate,—the ridge extend-
ing right up to the anterior border of the buccal cavern.
II. Hyperolissa, in which ridges defining the efferent branchial
channels are either altogether absent or are present on the posterior
part of the palate only. :
I do not think that these sections, depending on a single variable
character, should be considered as families, or even as subfamilies.
The section Hyperolissa, which corresponds to Dana’s family
Cancride, minus Cancer and Pirimela and Polydectus, is here subdivided
into 3 subfamilies, hereafter characterized, namely, Xanthine, Acting,
and Chlorodine.
The section Hyperomerista, which corresponds to Dana’s family
Eriphiide, minus Oethra, is here subdivided into 4 subfamilies, hereafter
characterized, namely, Menippine, Oziine, Pilumnine and Hriphiine.
The genus Platypilumnus, Wood-Mason MS., Alcock, Ann. Mag.
Nat. Hist. May 1894, p. 401, and Illustrations of the Zoology of the
Investigator, Crustacea pl. xiv. fig. 6, probably belongs to this family
and to the section Hyperolissa, and is probably related most nearly to
Galene ; but as I have only a single female specimen to go by its exact
position must remain undecided.
The following artificial key is meant to serve for the discrimination
of the Indian genera of this family :—
A. Alcock — Carcinological Fauna of India. 71
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A. Alcock — Carcinological Fauna of India.
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1898. ] A. Alcock — Carcinological Fauna of India. Ge
Section I. Hyperolissa.
Xanthids in which the efferent branchial channels are not defined
by a complete ridge on either side of the palate.
Subfamily I. XANTHINA.
Carapace usually much broader than long, usually transversely oval,
sometimes transversely hexagonal. The front is contained from 35 to 53
times in the greatest breadth of the carapace.
Alliance TI. CarprtnioipA. Carapace broad, transversely oval, the
antero-lateral border either entire, or divided into a few broad, shallow,
rounded lobes. Legs sub-cylindrical. Abdomen of the male with the
3rd and 4th, or usually the 3rd, 4th and 5th segments fused together.
Alliance IT. Zozymorpa. Carapace broad, transversely oval, the
antero-lateral border in the form of a sharp crest which may be either
thin and entire (fissured only) or cut into 4 large teeth, Legs with at
least the upper border of the merus carpus and propodus sharply
eristiform. Abdomen of the male with the 8rd, 4th and Sth somites
fused.
Alliance III. EHvuxantnota. Carapace broad, tranversely oval,
very profusely areolated in high relief; the antero-lateral borders are
continued below the orbits to the outer angle of the buccal cavern.
The basal antennal joint has its outer angle prolonged and impacted in
the orbital hiatus, and the antennary flagellum, which is hardly visible
without a lens, arises within the orbit. The abdomen of the male has
the 3rd, 4th and 5th somites fused.
Alliance IV. Xanruorpa. Front almost always prominent, square-
cut (notched or fissured in the middle line) and sublaminar, and almost
always separated from either supra-orbital margin by a deepish notch.
Carapace broad (except Medzxus and Httsodes), usually transversely oval,
but sometimes more hexagonal; the antero-lateral border usually cut
_ into sharp teeth. Male abdomen with segments 3-6 fused.
Alliance V. MHatimepompa. Front prominent and square-cut.
Carapace pentagonal, moderately broad. Abdomen of the male with
all 7 segments distinct, the last segment being more than twice as long
as any of the others.
Alliance VI. Gatenormpa. Carapace broad, pentagonal approach-
ing the quadrilateral, the antero-lateral border hardly longer than the
postero-lateral. The basal antennal joint does not nearly reach the
front. The abdomen of the male has all 7 segments distinct. The sole
type, Galene, is so singular that it might be separated as a distinct
subfamily.
78 A. Alcock — Carcinological Fauna of India. [No. 1,
Subfamily IJ]. ACTAEIN A.
Carapace usually much broader than long and usually very pro-
fusely and profoundly lobulated; the antero-lateral border is either
divided into 4 blunt lobes, or crenated. The front is about a third the
greatest breadth of the carapace, sometimes a little more, sometimes a
little less, and is divided into two rather prominent usually round-
pointed lobes.
Subfamily II. CHLORODINA.
Carapace hexagonal or transversely oval, or subcircular (Cymo) or
approaching the subcireular (Cyclodius). Front from a third to half
the greatest breadth of the carapace—much broader than in the preced-
ing subfamilies.
Alliance I. XantHopnoipa. Carapace transversely oval, front a
third or little less than a third the greatest breadth of the carapace,
fingers not hollowed at tip.
Alliance II. Cutoropioipa. Carapace transversely oval, front
nearer half than a third the greatest breadth of the carapace, fingers
hollowed at tip.
Alliance III. Cymorpa. Carapace subcircular, flat; front about
half the greatest breadth of the carapace: chelipeds remarkably un-
equal.
Subfamily I, XANTHIN A.
Alliance I. Carpilioida.
Carpilius. Liomera. Liagore.
Carpilodes. Lioxanutho.
Lachnopodus.
Carpitius, Leach, Desmarest, A. M. Edw.
Carpilius, Leach, Desmarest Consid. Gen. Crust. p. 104 (footnote).
Carpilius, Riippell, 24 Krabben roth. Meer. p. 13 (part).
Carpilius, Milne Edwards, Hist. Nat. Crust. I, 380.
Carpilius, De Haan, Faun. Japon. Crust. p. 16.
Carpilius, Dana, U. S. Expl. Exp. Crust. I. p. 159.
Carpilius, A. Mitne Epwarps. Ann, Scr. Nar. Zoot. (iv.) XVIII. 1862, p. 46,
and Novy. ArcuHiv. Dv Mos. I. 1865, p. 212, and Miss. Sci. Mex., Crust. p. 238.
Carpilius, Miers, Challenger Brachyura, p. 110.
Carapace broad, very convex in both directions, smooth (except for
some coarse pitting inside the frontal and antero-lateral border), with no
indication of regions; its antero-lateral borders strongly-arched, thick,
entire, smoothly-moulded ; its postero-lateral borders strongly-conver-
1898. | A, Alcock — Carcinological Fauna of India. 79
gent, straight, with a prominent tubercle at the angle of junction with
the antero-lateral.
Front moderately broad, (less than a third the greatest width of
carapace ) deflexed, 3-lobed, the middle lobe prominent and _ bilobulate,
the edges of all thickened.
Orbital margins entire, the upper margin thickened and forming
a well-marked blunt tooth at its junction with the antero-lateral margin.
Eyes on short thick stalks.
Antennules folding obliquely, almost transversely : inter-antennu-
lary septum broad.
Basal joint of antenne long, flat, running up into an oblique cleft
between the margin of the front and the infra-orbital plate; the
antennary flagellum very small, less than half the diameter of the orbit
and lodged in the said cleft.
Merus of the external maxillipeds with its anterior border very
oblique. 3
Chelipeds massive, smooth, unequal in both sexes; the fingers
bluntly pointed, those of the larger cheliped with a single pair of
‘molariform teeth, those of the smaller cheliped with a blunt cutting-
edge.
Legs smooth, sub-cylindrical.
Abdomen of male six-jointed—the 5rd and 4th somites fused with
obliteration of sutures, the 5th somite also immovably adherent to the
Ath. Large crabs.
Key to the Indian species of Carpilius.
1. Carapace with definitely disposed large red blots we OC. maculatus.
2. Carapace irregularly marbled with red oa «. OC. conveaus.
1. Carpilius maculatus, ( Linn.)
Cancer ruber, Rumph, Amboinsche Rariteitkamer, p. 18, pl. x. fig. 1.
Cancer saxatile, Seba, Thesaurus, III. 47, pl. xix. fig. 12.
Cancer maculatus, Linn. Syst. Nat. (xii.) p. 1042: Fabricius, Ent. Syst. II. 447,
and Suppl. p. 338: Herbst, Krabben, I. ii. 185, pl. vi. fig. 41, and I. ii. 263, pl. xxi.
fig. 118, and III. iv. 8, pl. 1x. fig. 2: Desmarest, Consid. Gen. Crust. p. 14,
Carpilius maculatus, Milne Edwards, Hist. Nat. Crust. I. 382, and in Cuvier
Régne Animal, Crust. pl. xi. fig. 2: De Haan, Faun. Japon., Crust. p., 7 (name only):
Dana, U. S. Expl. Exp., Crust. pt. I. p. 160: Stimpson, Proc. Ac. Nat. Sci. Philad.
1858, p. 32: Alph. Milne Edwards, in Maillard’s Pile Réunion, Annexe F, p. 3, and
Nouv. Archiv. du Mus. 1. 1865, p. 214 and 1X. 1878, p. 175: Heller, Reise Novara,
Crust. p.9: Hess, Archiv. fur Naturges. XXXI. i. 1865, pp. 183 and 171: Hoffmann,
in Pollen and Van Dam, Faun. Madagasc., Crust. p. 3; Richters in Mobius Meeresf.
Maurit. p. 145: F. Muller, Verh. Ges. Basel. VIII. 1886, p. 473: Miers, Challenger
Brachyura, p. 111: de Man, Archiv. f. Naturges. LIII. 1887, i. p. 231, and Zool.
80 A. Alcock — Carcinological Fauna of India. [ No. 1,
Jahrbuch., Syst. VIII. 1895, p. 496: Cano, Boll. Soc. Nat. Napol. III. 1889, p. 189:
J. R. Henderson, Trans. Linn. Soc. Zool. (2) V. 1898, p. 353: Ortmann, Zool.
Jahrbuch., Syst. VII. 1894, p. 469.
Front obliquely deflexed, the median lobe very decidedly bilobulate
and separated from the lateral lobes on either side by a deep notch.
Carapace with not less than eleven large roundish dark-red blots
(which seem never to competely fade even in very old Museum speci-
mens) disposed as follows :—two on either side immediately behind the
eye, the smaller and anterior one of these involving the orbital margin ;
three in a transverse curve across the middle of the carapace; four in
another transverse line just in front of the posterior margin.
Hight specimens, from the Andamans, Nicobars, and Palk Straits.
2. Carzilius convexus, (Forskal) Riippell.
Cancer convezus, Forskal, Descr. Anim. p. 88.
Cancer adspersus, Herbst, Krabben, I. ii. 264, pl. xxi. fig. 1.
Cancer marmarinus, Herbst, Krabben, III, iv. 7, pl. lx, fig. 1.
Carpilius convexus, Ruppell, 24 Krabben roth. Meer. p. 13, pl. iii, fig. 2
and pl. vi. fig. 6: Milne Edwards, Hist. Nat. Crust. I. 382, pl. xvi. figs. 9,
10: DeHaan, Faun. Japon. Crust. p.17 (name only): Dana, U. 8. Expl. Exp. Crust.
pt. I. p. 159, pl. vii. fig. 5: Stimpson, Proc. Acad. Nat. Sci. Philad. 1858, p. 32:
Heller, SB. Ak. Wien XLITII. 1861, p. 319: Alph. Milne Edwards in Maillard’s Vile
Réunion Annexe F, p. 3, and Nouv. Archiv. du Mus. I. 1865, p. 215, and IX. 1878, p.
176: Hilgendorf in v. d. Decken’s Reisen in Ost-Afrika III.i. p. 73: Hoffmann in
Pollen and Van Dam, Faun. Madagasc., Crust. p. 3: Miers, P. Z. S. 1877, p. 133,
and Ann. Mag. Nat. Hist. (5) II. 1878, p. 407: Richters in Mébius Meeresfauna
Manrit. p. 145: EH. Nauck, Zeitschr. Wiss. Zool. xxxiv. 1880, p. 56 (gastric teeth) :
Haswell, Cat. Austr. Crust. p. 41: F. Muller, Verh. Ges. Basel VIII. 1886, p. 473:
de Man, Archiv. f. Naturges. liii. 1887, i, 232, and Zool. Jahrb. Syst. VIII. 1895, p.
496: Ortmann Zool. Jahrbuch., Syst. etc., VII. 1894, p. 469, and in Semon’s Zool.
Forschungsr. (Jena. Denkschr. VIII.) Crust., p. 51: Zehntner, Rev. Suisse Zool. II.
1894, p. 143.
Carpilius lividus, Gibbes, Proc. Amer. Ass. III. 1850, p. 174, is according to A.
Milne Edwards, vide Nouv. Archiv. du Mus. I. 1865, p. 217, the young of Carpilius
convesus. Miers also, Ann. Mag. Nat. Hist. (v) II. 1878, p. 407, considers C. lividus
to be a synonym of Carpilius convexus.
Front vertically deflexed, the prominent median lobe is not de-
cidedly bilobulate—in fact, it is sometimes but obscurely emarginate at
tip—and is separated on either side from the lateral lobes by only a
shallow excavation.
Carapace irregularly marbled with dark red, which in old spirit
specimens sometimes fades entirely.
Seven specimens from the Andamans and Nicobars, |
1898. ] A. Alcock — Carcinological Fauna of India. 81
Carpitopes, Dana, A. Milne Edwards.
Carpilodes, Dana, Silliman’s Amer. Journ. Sci. and Arts, (2) XII. 1851, p. 126, and
Proc. Acad. Nat, Sci. Philad. VI. 1852, p. 77, and U.S. Expl. Exp. Crust. pt. I. p. 192.
Carpilodes, Alph. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 224 (et
synon.)
€arpilodes, Miers, Challenger Brachyura, p. 133.
Carpilozanthus, Alph. Milne Edwards in Maillard’s Vile Réunion, Annexe F,
p. 3. (A. M. E.)
Carapace very broad, convex in both directions, with the regions
generally well demarcated and—especially in the anterior half—sub-
divided into lobular areols; its antero-lateral borders usually sub-
divided into four broad, shallow, rounded lobes; its postero-lateral
borders straight, or a little concave, and strongly convergent.
Front broad (about a third the greatest breadth of the carapace)
obliquely deflexed, grooved and slightly notched in the middle line, but
not distinctly bilobed. |
Orbits small, with entire margins, but usually with the three suture
lines near the outer angle more or less distinct: eye-stalks short and thick.
Antennules folding obliquely, almost transversely. Basal antennal
joint running up between the front and the lower orbital plate much as in
Carpilius; the flagellum rather longer than the major diameter of the orbit.
Anterior edge of merus of external maxillipeds almost transverse.
Chelipeds equal or subequal in both sexes: fingers pointed, but
distinctly grooved or hollowed near the tips.
Abdomen of the male five-jointed, the 8rd—5th somites fused.
Small crabs, easily recognizable by their short broad convex
carapace, with its antero-lateral margins in the form of four broad
shallow rounded lobes, its postero-lateral margins strongly convergent,
and the broad deflexed rather prominent and convex front.
Key to the Indian species of Carpilodes.
I. Surface of carapace quite smooth to the naked eye :—
i. Upper border of meropodites of legs crest-like ... C, lophopus.
ii. Upper border of meropodites of legs not crest-like :—
1. Posterior part of carapace not lobulated :—
a. Gastric region subdivided into three
lobules only ... ade ce, O, tristis.
b. Gastric region subdivided into five
lobules :—=
a. Outer surface of wrist and
hand smooth am ... OO, venosus.
8. Outer surface of wrist nodular,
of hand granular... vee C. stumpsoni.
2. The whole of the carapace divided into a
network of lobules by fine lines wee = C, pediger,
ar. Ll
82 A. Alcock — Carcinological Fauna of India. [No. 1,
II. Part or all of the surface of the carapace covered with
vesiculous granules plainly visible to the naked eye :—
i, Posterior part of the carapace not lobulated :—
1. The whole of the carapace covered with
granules .. are . C. rugatus.
2. Only the mite’ lateral hatte of the carapace
granular ... we C. vaillantianus.
ii. Posterior part of the carapace more or pien divided
into lobules by transverse grooves: the whole surface
of the carapace densely granular :—
1. A single transverse furrow behind the Anat
region ws. .. CO. margaritatus.
2. Two transverse peat (iheikecics of one
that helps to form the raised posterior
margin) behind the gastric region :—
a. Branchial lobules few, long, roll-like... C. monticulosus.
b. Branchial lobules many, small, nodule-
like ae aN -. OC. cariosus.
3. COarpilodes tristis, Dana.
Carpilodes tristis, Dana, U. 8. Expl. Exp. Crust. pt. I., p. 193, pl. ix. figs. 7a-d:
Heller, Novara Crust. p. 17: Alph. Milne Edwards, Nouy. Archiv. du Mus. I. 1865,
p. 225, and 1X. 1873, p. 178: Haswell, Cat. Austr. Crust. p, 56: F. Muller, Verh. Ges.
Basel, VIII. 1886, p. 474: de Man, Notes Leyden Mus. XII. 1890, p. 50: J. R.
Henderson, Tr. Linn. Soc., Zool., (2) V. 1898, p. 853: Ortmann, in Semon’s Zool.
Forschungsr. (Jena. Denkschr. VIII) Crust. p. 51.
Surface of carapace and appendages quite smooth to the naked eye,
but with a dull look due to uniform microscopic miliary granulation.
Gastric region delimited from the front, from the somewhat tumid
supra-orbital margins, and posteriorly, by shallow grooves, and sharply
demarcated from-the branchial regions by fine sharp-cut lines; and
subdivided into three lobules by a fine sharp-cut X shaped median
incision.
Antero-lateral borders divided into four lobes, from the intervals
between which fine sharp lines run obliquely inwards to incompletely
subdivide the hepatic and branchial regions into lobules, Outer part
of hepatic regions on a plane slightly lower than that of the rest of the
carapace.
Colours in spirit: uniform dull brownish-buff, except the fingers and
a large part of the lower border of the hand, which are black.
79 specimens from the Andamans and Nicobars.
4, Carpilodes stimpsoni, A. Milne Edwards.
Carpilodes stimpsoni, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 232,
pl. xi. figs. 2-2c, and IX. 1873, p. 181: de Man, Archiv. fur Naturges. LIII. 1887, i.
p. 284, and Journ. Linn. Soc., Zool., vol. XXII. 1887-88, p. 25.
1898.] A. Alcock —-Carcinological Fauna of India. 83
Differs from Carpilodes tristis Dana in the following more conspicu-
ous particulars :— .
(1) the surface of the carapace is of a shiny smoothness, except
for some irregular pitting on the lobules of the anterior portion :
(2) the gastric region is subdivided into five longitudinal lobules
(as in all the following species) by incisions running almost parallel
with the limbs of the X shaped median incision :
(3) the chelipeds have the upper and outer surface of the wrist
nodular and of the hand granular, and the upper surface of the cor-
responding joints and merus of the legs nodular :
(4) the colour in spirit is light yellowish.
A single specimen from Mergui.
It appears to me very doubtful whether this species is really
distinct from C. venosus.
5. Carpilodes venosus, (Edw.)
Carpilius venosus, Milne Edwards, Hist. Nat. Crust. I. 383.
Carpilodes venosus, A. Milne Hdwards, Nouv. Archiv. du Mus. I. 1865, p. 227,
pl. xii. figs. 2-2b, and IX. 1873, p. 179: Miers, Zool. H. M.S. “Alert,” pp. 183 and
218: Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 358.
Xantho obtusus, De Haan, Faun. Japon. Crust., p. 47, pl. xiii. fig. 5: Krauss,
Sudafr. Crust. p. 31.
Included in the Indian fauna on the authority of Dr. J. R.
Henderson : there are no specimens in the Indian Museum referable to
this species, unless (as, indeed, I believe) C. stimpsoni is synonymous.
From Milne Edwards’ figures this species differs from C. stimpsont
in having the chelipeds and legs perfectly smooth.
6. Carpilodes pediger, n. sp.
Allied to ©. venosus and stimpsoni, from which it differs in having
the whole of the carapace mapped out in lobules.
Most closely allied to 0. ruber A. M. Edw., from which it conspicu-
ously differs in the form of the male chelipeds.
Carapace extremely convex in both directions, its surface, like that
of the appendages, being perfectly smooth to the naked eye though
very finely granular under the lens: it is symmetrically and minutely
subdivided by fine lines into very many little-convex and rather angular-
outlined lobules. The antero-lateral borders are rather deeply four-
lobed, the prominence of the outer angle of the orbit forming a small
fifth lobule.
The chelipeds in the adult male are close upon twice the length of
the carapace and have a very strong tooth on the inner upper border of
84 A. Alcock — Carcinological Fauna of India. [Nosd)
the wrist, and strongly-arched fingers which meet only at the tip, the
movable finger bearing (in addition to the serrations of the hollow tip)
a strong tooth near the base.
In the adult female the chelipeds are very little longer than the
carapace and have only a small tooth on the wrist, and fingers which
are not strongly arched but meet through the greater part of their
extent. |
Colours in spirit, light straw, fingers very light brownish: some-
times the wings of the carapace are light grey and then there is also a
light grey stripe down the middle of the carapace, fore and aft.
Length of carapace 6°5 millim., breadth 10 millim.
Off Andaman Is. 10-41 fms., off Ceylon 263 fms.
Seven specimens,
7. Carpilodes lophopus, n. sp.
All parts are smooth to the naked eye, though under the lens the
surface of the carapace and chelipeds is minutely pitted or eroded. The
regions are demarcated and subdivided by very fine lines; and the
lateral gastric areole (2 M of Dana) and the mid-branchial areole (4
and 5 L of Dana) are particularly, and rather angularly, convex.
The antero-lateral borders are four-lobed, the last two lobes being
rather augular ; the postero-lateral borders are markedly concave.
The front is broad and projects well beyond the orbits.
Chelipeds not very much longer than the carapace: two little
tubercles, one above the other, at the inner angle of the wrist, and
two at the distal end of the upper border of the hand.
The upper border of the meropodites of the legs is distinctly
cristiform, that of the carpopodites is sinuous-cristiform, and both the
upper and the lower edges of the propodites are cristiform—the lower
more distinctly than the upper.
Colours in spirit, yellowish white.
Carapace about 5°5 millim. long, abont 8°5 millim. broad,
Off south-east coast of Ceylon, 34 fms., a male and a female; a
female from off Malabar coast, 29 fms.
8. Carpilodes rugatus, (Latr.) A. Milne Edwards.
Zozymus rugatus, Milne Edwards, Hist. Nat. Crust. I. 385, (A. M. E.)
Zozymus canaliculatus, Lucas, Voy. Astrolabe, Crust. p. 21, pl. iii. fig. 2(A. M. E.)
Carpilowanthus rugatus, A. Milne Edwards in Maillard’s Vile Réunion, Annexe
F, p. 3. (A. M. E.)
Carpilodes rugatus, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 230,
pl. xii. figs. 8, 3b, and IX. 1878, p. 180: Richters in Mobius Meeresf. Maurit. p. 146 :
Miers, Zool. H. M. §. “ Alert,” pp. 517 and 529,
1898. ] A. Aleock— Carcinological Fauna of India. 85
Surface of carapace uniformly covered with granules which are
visible to the naked eye and on the antero-lateral parts of the carapace
are vesiculous: the upper and outer surfaces of the wrist and hand, and
of the corresponding joints of the legs, are closely granular to the naked
eye, the granules of the hand being arranged in longitudinal series.
As in all the Indian species of Carpilodes except C. tristis, the
frontal and supra-orbital borders are cut off from the rest of the cara-
pace by a sinuous groove which also includes the two front lobes of the
four-lobed antero-lateral border, and the gastric region is longitudinally
5—lobular.
Transverse grooves running from the last two intervals between
the lobes of the antero-lateral border cut off, respectively, (1) the hepatic
from the branchial regions, and (2) the first branchial lobule from the
rest of the branchial region. All the lobules are strongly convex.
The cardiac region is not defined, and there is no lobulation of the
posterior moiety of the carapace.
Colours in spirit—pink, fingers purplish-brown with white tips,
3 specimens from the Cocos Islands (Andamans).
9. Carpilodes vaillantianus, A. Milne Edwards,
Carpilozanthus vaillantianus, A. Milne Edwards, in Maillard’s Vile Réunion
Annexe F, p. 3. (A. M. EH.)
Carpilodes vaillantianus, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865,
p. 231, pl. xi. figs. 3-3b. Haswell, Cat. Austral. Crust. p. 57: Miers, Zool. H. M.S,
“ Alert,” p. 529: de Man, Archiv. f. Naturges. LIII. 1887, i. p. 235: Ortmann in
Semon’s Zool. Forschungsr. (Jenaische Denksch. VIII.), Crust. p. 51.
_ This species, if it is really distinct from C. rugatus, differs from the
latter in the following particulars :—
(1) the granulation is confined to the antero-lateral parts of the
carapace :
(2) the lobules of the carapace are less convex :
(3) the furrow that cuts off the anterior branchial lobule does not
meet the furrow that bounds the gastric region.
Among 17 specimens in the Indian Museum there is a good deal of
variation of these characters; so much so, that some of the specimens
might almost be referred to C. rugatus, especially to the “ Astrolabe”
figure.
Five specimens from the Andamans, three from Muscat, two from
Mergui; (the others from Mauritius, Samoa, and Viti).
10. Oarpilodes margaritatus, A. Milne Edwards.
Carpilodes margaritatus, A. Milne Edwards, Nouv. Archiv. du Mus, IX, 1873,
86 A. Aleock— Carcinological Fauna of India. [No. 1,
p. 182, pl. v. fig. 2: Henderson, Trans. Linn. Soc. Zool. (2) V. 1893, p. 353:
Whitelegge, Mem. Austral. Mus. III. 1897, p. 181.
Carapace and legs covered with pearly granules plainly visible to
the naked eye. The carapace is much lobulate, the anterior branchial
lobe being itself trilobulate, and the region behind the gastric region
being crossed transversely by a furrow. The antero-lateral borders are
indistinctly four-lobed. The hands are not longitudinally furrowed.
Colours ; red, fingers black.
No specimens in the Indian Museum collection. Included here on
the authority of Dr. J. R. Henderson.
11. Carpilodes cariosus, n. sp.
Allied to C. margaritatus.
Carapace strongly convex, its whole surface intricately cut up, by
deep grooves, into many small strongly-convex lobules, the surface of
which is pitted and granular, so as to give the carapace as a whole a
somewhat worm-eaten appearance.
The legs also have the extensor surfaces of the long joints granular
and nodular: the outer surface of the wrist is nodular: the outer
surface of the hand is granular and furrowed.
The antero-lateral borders are very distinctly four-lobed.
The space between the gastric region and the posterior border of
the carapace is broken by two (or three, counting the groove inside the
raised posterior border) deep transverse grooves, the space between the
grooves being Cupid’s-bow-shaped. A transverse groove also cuts off a
narrow piece from the posterior extreme of the mesogastric lobule.
Colours in spirit ; whitish with pink spots on carapace, legs pink,
fingers sometimes black with white tips, sometimes pinkish white.
Length of carapace about 5 millim., breadth about 7 millim.
Off Ceylon 26} to 34 fms., 13 specimens including several ovigerous
females: off Andamans 10 to 15 fms., 2 specimens.
12. Carpilodes monticulosus, A. Milne Edwards.
Carpilodes monticulosus, A. Milne Edwards, Nouv. Archiv. du Mus, IX. 1873, p.
181, pl. v. fig. 1: de Man, Archiv. f. Naturges. LIII. 1887, i. p. 2838: Ortmann in
Semon’s Forschungsreisen (Jena. Denkschr. VIII.) Crust. p. 51.
Carapace very broad (not far short of twice as broad as long), its
surface everywhere closely covered with elegant vesiculous granules.
The whole of the carapace is divided, by deep broadish grooves, into
elongate lobules of an elegant smooth roll-like form (quite unlike any
other Indian species). A marrow beaded lobule forms the posterior
limit of the mesogastric lobe (much as in C. cartosus), and two
1898. ] A. Alcock — Carcinological Fauna of India. 87
furrows cross transversely the region between the latter and the
posterior border of the carapace, On the branchial regions, on either
side of the mesogastric lobule, is a small dimpled lobule. ‘The wrist
and hand are closely covered with granules like those on the carapace,
the wrist being dimpled and the hand longitudinally furrowed.
The extensor surfaces of the legs are also closely, but much more
finely, granular, the carpus in all being dimpled.
Antero-lateral borders four-lobed, the lobes narrow, rather shallow,
rounded, and the last three of nearly equal size.
Colours in spirit; dark purplish-red, legs lighter, fingers white
with brownish base.
Two specimens, from Gt. Coco I. (Andamans) and East L,
Andamans are in the Indian Museum.
Carapace not quite 6 millim. long, 10 millim. broad,
Liompra, Dana.
Liomera, Dana Silliman’s Amer. Journ. Sci. and Arts (2) XII. 1851, p. 124;
Proc. Acad. Nat. Sci. Philad. 1852, p. 73; and U. S. Expl. Exp. Crust. pt. I.
p. 160.
Liomera, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 218, and Exp.
Sci. Mex. Crust. p. 239.
Carapace extremely broad, strongly convex in both directions,
transversely barrel-like, either smooth or with the regions very faintly
indicated ; its antero-lateral borders thick, either entire or divided into
four broad shallow rounded lobes, of which the first two are almost
coalescent; its postero-lateral borders very strongly convergent,
straight or a little concave.
Front narrow (from a third to less than a fourth the breadth of the
carapace), obliquely deflexed, grooved and slightly notched in the
middle line, but not distinctly bilobed.
Orbits small, with the three suture lines near the outer angle
usually distinct ; eye-stalks short and thick.
The antennules fold nearly transversely. Basal antennal joint broad
and short, merely touching the front; the flagellum, which is short,
lodged in the orbital hiatus.
Anterior edge of merus of external maxillipeds a little oblique.
Chelipeds equal or subequal in both sexes; fingers somewhat
hollowed at tip. Legs sub-cylindrical.
Abdomen of the male five-jointed, the 3rd—5th somites being
fused.
Small or medium-sized crabs, easily recognized by their short, very
broad, strongly convex, barrel-like carapace.
88 A. Aleock— Carcinological Fauna of India. [No. 1,
13. Liomera cinctimana, (White), Dana.
Carpilius cinctimanus, White, in Jukes Voyage H. M. 8. “ Fly,” Vol. II. p. 336,
pl. ii. fig. 83, and Samarang Crust. p. 37, pl. vii. fig. 4.
Liomera cinctimana, Dana, Silliman’s Journ. (2) XII. 1851, p. 124, and U.S.
Expl. Exp. Crust. pt. I. p. 161: A. Milne Edwards Nouv. Archiv. du Mus. I. 1865,
p. 219, and IX. 1873, p. 176, pl. v. fig. 4, and Exp. Sci. Mex. Crust. p. 240 : Stimpson,
Ann. Lyc. New York, X. 1874, p. 103.
Carpilodes cinctimanus, Miers, Ann. Mag. Nat. Hist. (5) V. 1880, p. 2384:
Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 354.
Liomera lata, Dana, Proc. Ac. Nat. Sci.. Philad. 1852, p. 73, and U. S. Expl.
Exp. Crust. pt. I. p. 161, pl. vii, figs 6a-d: Stimpson, Proc. Acad. Nat. Sci. Phil.
1858, p. 32, and Ann. Lyc. New York, X. 1874, p. 104: Heller, Novara Crust.
p. 9: A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 220, and Exp. Sci. Mex.,
Crust. p. 240: F. Muller, Verh. Ges. Basel. VIII. p. 474.
Carapace extremely broad—its length only about ;' of its breadth
—its surface, like that of the appendages, everywhere smooth and
polished, showing only the faintest indications of a gastro-cardiac region
and of oblique lobulation of the branchial regions: the antero-lateral
border is divided into three coarse lobes, the anterior of which is again
obscurely divided into two. |
Front obliquely deflexed, with a rather prominent convex edge
cleft in the middle line. Orbital margin with three radiating suture-
lines near the outer angle. Chelipeds equal.
Colours in spirit; orange-red, fingers black, hand with a broad
black cross-band merging with the black of the immobile finger.
3 specimens from the Andamans and Muscat (besides. specimens
from Mauritius and South Sea Is.).
14. PLiomera sodalis, n. sp.
Carapace broad (length about 3 breadth) very strongly convex,
perfectly smooth, without any indication of regions, its margins smooth,
entire. Front nearly vertically deflexed, its edge cleft in the middle line.
Eyes large, supra-orbital margin without any suture-lines. Chelipeds
a little unequal; the upper and outer surfaces of the carpus and hand
of the smaller cheliped covered with prickly granules, but in the larger
cheliped the granulation has a very much worn appearance: fingers
hardly hollowed at tip.
Legs (those that are present in the unique specimen) somewhat
hairy ; none of the joints are carinate though some have prickly granules
on the upper surface.
Colours in spirit—of the same blotchy orange and reddish colour as
that of a species of Solenocawlon, in the hollow stem of which the crab
was found,
1898. |. A. Alcock — Carcinological Fauna of India. 89
Length of carapace 6 millim., breadth 9 millim.
Off south-east corner of Ceylon, 32 fms.
This species resembles a small Aftergatis, but has sub-cylindrical
legs and has no margin to the carapace.
Lacunopopvus, Stimpson.
Lachnopodus, Stimpson, Proc. Ac. Nat. Sci. Phila. 1858, p. 32: A. Milne Edwards,
Nouv. Archiv. du Mus. I. 1865, p. 233: Ortmann, Zool. Jahrb. Syst. VII. 1893-94,
p. 452.
‘‘Carapax laevis, regione posticé transversim convexa. Orbita
margine externa trifissa vel trilobata, lobis parvis, obtusis. Antenne
ut in Carpilio (ut in Liomeré? ). Gnathopoda intima lacinia ad apicem
non furcaté. Hectognathopoda ischio longitudinaliter sulcato; mero
superficie versus angulum internum excavata, margine anteriore con-
cava. Chelopoda manu facie externa sulcaté. Pedes ambulatorii valde
setosi, mero compresso, superne spinoso.
“‘ Liomere affinis, sed carapace angustiore, pedibus setosis spino-
sisque.”
This genus is not represented in the Indian Museum.
15. Lachnopodus rodgersi, Stimpson.
Lachnopodus rodgersii, Stimpson, Proc. Ac. Nat. Sci. Phila. 1858, p. 32.
Liomera rodgersii, Miers, Ann. Mag. Nat. Hist. (5) V. 1880, p. 231, pl. xiii. fig. 3,
(orbit and antennz only): de Man, Archiv. fiir Naturges. LIII. 1887, i. p. 237:
J. R. Henderson, Trans. Linn. Soc. Zool. (2) V. 1893, p. 354.
“Carapace transverse, about once and a half as broad as long,
smooth, glabrous, and shining, with the interregional sutures almost
obsolete ; the two posterior teeth of the antero-lateral margins are the
only ones developed, and are very obscurely marked and obtuse. The
front is somewhat produced, and is divided by a median and two lateral
incisions into four lobes, of which the two median are broad and
truncated, and the lateral (or inner orbital lobes) are small and denti-
form. On the upper orbital margin are three small obtuse teeth
(including that of the outer orbital angle); the tooth at the inner
and lower orbital angle is rather prominent. The merus-joint of the
outer maxillipeds is rather small and transverse; and this joint has a
shallow pit on its outer surface. The anterior legs (in the male) are
rubust, smooth; arm or merus-joint with a series of spinules on its
upper margin; carpus smooth, with an antero-internal tooth ; penu!ti-
J, 1. 12
90 A. Alcock — Carcinological Fauna of India. [No. ],
mate joint or palm slightly rugose externally, and with two longitudinal
and parallel grooves on its outer surface ; fingers short, robust, toothed
on their inner margins and with the apices not excavated. The
ambulatory legs are somewhat compressed and clothed with long fulvous
hairs; their merus-joints are spinulose on their upper margins. The
postabdomen of the male is five-jointed, the third to fifth joints being
coalescent. Length 83 lines, breadth nearly 1 inch 1 line.
This species has been hitherto known only from the very short
generic definition of Dr. Stimpson, which, however, embraces all the
characteristic peculiarities of the species, and which agrees exactly with
the example before me, except in one point. Stimpson says (1, c.),
*“ Antennae ut in Carpilio.’ In the specimen now before me the
antennae are of the same structure as in Ivomera, the basal joint being
very short and united at its summit to an inferior prolongation of the
front, and not, as in Carpilius, joined to the front along its inner margin.
I have little doubt that Stimpson erroneously wrote Carpilius for
Liomera, as he does not say that Lachnopodus is distinguished from
Liomera by any peculiarity in the structure of the antenne.
I do not think that Lachnopodus is generically distinct, as Milne
Edwards has described a Liomera (L. longimana) with hairy ambulatory
legs.” | (Miers.)
Genus LIoxaNntTHo.
Carapace broad, moderately or strongly convex in its anterior two-
thirds, flat posteriorly ; the gastric region is fairly or faintly delimited,
and one or two short furrows may pass on to it obliquely from the
intervals between the lobes of the antero-lateral margin, but beyond
this there is no distinct division of the carapace into regions or sub-
regions.
The antero-lateral borders are much like that of Liomera, being
divided into 4 broad blunt lobes, of which the first two are so much
coalescent as to really form but one.
Front somewhat deflexed, bilobed, or sinuous and notched in the
middle line. Fronto-orbital border less than half the greatest width of
he carapace.
Orbital margin and antenne as in Xantho.
Anterior edge of merus of external maxillipeds a little more
oblique than in Xantho.
Chelipeds either subequal or unequal in both sexes, fingers pointed.
Legs subcylindrical.
Abdomen of male five-jointed, the 3rd-5th somites coalescent.
1898. A. Aleock— Carcinological Fauna of India. Sd
Key to the species of the genus Lioxantho.
I. Chelipeds unequal, outer angles of front separated from
the supra-orbital margin by a notch; regional markings
of carapace almost obsolete bb den oo. L. twmidus.
II. Chelipeds equal, outer angles of front fused with supra-
orbital margin :—
i. Carapace chelipeds and legs smoothas wax ... IL. punctatus.
ii. Carapace chelipeds and legs aniformly closely
and finely granular... vow .. L. asperatus.
16. ILnoxantho punctatus, (Kdw.)
_ Xantho punctatus, Milne Kdwards, Hist. Nat. Crust. 1. 396: A. Milne Edwards,
Nouv. Archiv. du Mus. IX. 1873, p. 199, pl. vii. fig. 6: Miers, Challenger Brachyura,
p. 125: de Man, Zool. Jahrb. Syst. 1V. 1889, p. 420, and Notes Leyden Mus. XII.
1890, p. 52, pl. iii. fig. 1.
Liomera punctata, Miers, Zool. H. M.S. Alert, pp. 517, 528: de Man, Archiv. fur
Naturges. LIII. 1887, i. p. 238: J.R. rRedtiendde: Trans. Linn. Soc. Zool., (2) V.
1893, p. 354: Ortmann, Zool. Jahrb., Syst. VII. 1893-94, p. 451.
Carapace moderately convex in the anterior two-thirds, flat post-
eriorly, its surface smooth. The gastric region is fairly well defined
antero-laterally, and the fronto-orbital region marked off, by distinct
grooves ; and two short grooves (of which the anterior is the longer)
pass in obliquely from the notches between the 2nd and 3rd, and 3rd
and 4th lobes of the antero-lateral margin; but this is all the areola-
tion that exists.
Antero-lateral border moulded into four broad shallow lobes, of
which the first two are almost coalescent.
Front bilobed, the outer angle of each lobe fused with the supra-
orbital border, as in Xantho impressus and as in no other Indian species
of Xantho: the width of the front is about a fourth the greatest breadth
of the carapace.
Chelipeds equal in both sexes, perfectly smooth, although a very
indistinct groove sometimes runs about half way along the outer surface
of the hand near the upper border.
Legs thickish, smooth, the dactylus with some fur and a few short
hairs,
Colours in spirit; pinkish yellow or buff, with small red spots on
carapace, and ill-defined pinkish-brown patches on chelipeds and legs ;
fingers black, with light brown tips,
In the Indian Museum are 3 specimens from Cey lon, (as well as 8
from Mauritius and 2 from Samoa).
17. LInowantho tumidus, n. sp.
Carapace in its anterior two-thirds strongly convex from before
92 A. Alcock — Carcinological Fauna of India. [No. I,
backwards and a little convex from side to side, flat in its posterior
third ; perfectly smooth and polished. The limits of the gastric region,
and its division into three sub-regions, are faintly apparent as mere
markings, not grooves; and the fronto-orbital region is marked off by a -
faint groove.
The antero-lateral border is divided into four broad shallow lobes,
of which the first two are almost confluent; from the notch between
the second and third a short groove runs obliquely inwards on to the
carapace, and a still shorter one from the notch between the third and
fourth.
Front much less than a fourth the greatest width of the carapace,
bilobed, the outer angle of each lobe separated from the supra-orbital
margin by a notch and groove.
Chelipeds unequal, smooth and polished.
Legs rather thick, smooth ; a few scattered hairs along the upper
border of the last three and along the lower border of the last two
joints, the dactylus also furred, The upper border of the meropodites
of all the legs, as well as of the arm, is microscopically serrulate or
crenulate.
Colours in spirit pinkish yellow, fingers black with light brown
tips.
In the Indian Museum are 3 specimens from the Andamans, (and
one from Samoa).
This species exactly resembles a quite smooth and strongly-
inflated Xantho bidentatus, and but that I have 4 specimens, represent-
ing both sexes and different ages, I should have regarded it as an
abnormality of that species.
It also has a remarkable resemblance to the Xantho (Lachnopodus )
tahitensis figured and described by de Man in Zool. Jahrb. Syst. IV.
1889, p. 418, pl. ix. fig. 43 but it has not the row of strong spines along
the upper border of the meropodites of the chelipeds and legs, that are
characteristic of that species.
18. ILnoxantho asperatus, n. sp.
Carapace very slightly convex fore and aft in its anterior two-
thirds, quite flat posteriorly and from side to side, very closely sharply
and uniformly granular everywhere except the posterior median portion,
where the granulation is visible only under a lens. The gastric region
is faintly delimited, a short bifurcating groove runs in from the frontal
notch, and two very faint grooves run in obliquely from the two notches
of the antero-lateral margins, but this is all the attempt at areolation
that exists. | ;
1898. ] A. Alcock— Carcinological Fauna of India. 93
Antero-lateral border granular and rather sharp, very obscurely
divided into three most inconspicuous lobes, the first of which hardly
shows a trace of subdivision.
Front not quite a fourth the greatest breadth of the carapace,
obliquely deflexed, emarginate and faintly grooved in the middle line,
its outer angles not separated from the supra-orbital margin.
Chelipeds equal, the upper corner of the outer surface of the arm,
the upper and outer surfaces of the wrists and hands, closely covered
with little pearly granules like those on the antero-lateral parts of the
carapace.
Legs stout: the upper edge of the merus and the dorsal surface of
the next two joints granular like the chelipeds, the dactylus hairy.
Colours in spirit orange-yellow.
In the indian Museum are a male and female probably from
Karachi.
Liacore, De Haan.
Liagore, De Haan, Faun. Japon. Crust. p. 19.
Liagora, Dana, Amer. Jour. Sci. and Arts (2) XII. 1851, p. 124; and U.S. Expl.
Exp. Crust. pt. I. p. 148.
Carapace somewhat approaching the quadrilateral, strongly convex
fore-and-aft, little convex from side to side, smooth, without any indic-
ation of regions.
Antero-lateral border moderately arched, entire; postero-lateral
border very moderately convergent, straight, about as long as the chord
of the antero-lateral ; posterior border long,—about half the greatest
width of the carapace in length, or more.
Fronto-orbital border about half, front about quarter, the greatest
width of the carapace in extent. Front a little deflexed, broadly
bilobed. Orbital margin thin entire, the outer angle of orbit a little
thickened. Eyes on very short thick stalks.
The antennules fold nearly transversely. Basal antennal joint very
short and broad, but passing up between the side of the front and the
inner angle of the orbit; the flagellum, which is about as long as the
major diameter of the orbit, lodged in the orbital hiatus.
Anterior edge of merus of external maxillipeds somewhat oblique.
Chelipeds massive, equal in both sexes, the fingers ume
Legs subcylindrical, rather long, smooth.
Abdomen of male five-jointed, the 3rd—5th somites fused.
19. Inagore rubromaculata, De Haan.
Cancer (Liagore) rubromaculatus, De Haan, Faun. Japon. Crust. p. 49, pl. v.
oe, 1.
94 A. Alcock — Carcinological Fauna of India. [No. 1,
Liagore rubromaculata. Miers, Ann. Mag. Nat. Hist. (5) II. 1878, p. 407 (note) ;
and Challenger Brachyura, p. 111, (footnote),
Carapace transversely somewhat oval, approximating the quadri-
lateral type, with long posterior and only moderately convergent
postero-lateral borders ; its surface devoid of sculpture and perfectly
smooth to the naked eye, microscopically pitted and granular : pterygo-
stomian region somewhat hairy.
Antero-lateral border moderately sharp, entire. Front broadly and
rather faintly bilobed, the outer angles of each lobe pronounced, pro-
minent, and separated from the supra-orbital margin by a short: shallow
groove. A little pimple-like thickening at the outer angle of the orbit.
Chelipeds equal, smooth and polished: both borders of the arm
hairy, the upper border with a few blunt denticles ; both the inner and
the outer angles of the wrist strongly pronounced ; eee long, isa
with the opposed edges strongly but bluntly serrate.
Legs long, subcylindrical, smooth and polished, the dactyli most
elegantly plumed.
Colours in spirit yellowish with numerous large livid red spots.
In the Indian Museum is a single specimen dredged off the
Irrawaddy Delta in 20 fms., (besides 8 from Hongkong).
Alliance II. Zozymorda.
Atergatis. Lophactza,
Zozymus.
Lophozozymus.
ArErGATIs, De Haan, A. Milne Edw.
Atergatis, De Haan, Faun. Japon. Crust. p. 17.
Atergatis, Dana, Silliman’s Journ. Sci. and Arts (2) XII. 1851, p. 124, and U. S.
Expl. Exp. Crust. pl. I. p. 57.
Atergatis, A. Mitng Epwarps, ANN. Scr. Nat. Zoot. (4) XVIII, ieee p. 49, and
Novv. Arcuiv. pu Mus. I. 1865, p. 231.
Atergatis, Miers, Challenger Brachyura, p. 111.
Platypodia, Bell, Trans. Zool. Soc. I. 1835, p. 336.
Carapace externally broad, convex in both directions, regional
boundaries absent or quite inconspicuous, surface either quite smooth
or somewhat pitted ; its antero-lateral borders strongly arched and with
an independent keel-like edge; the postero-lateral strongly convergent,
straight. The under surface of the wings of the carapace is a good deal
hollowed to receive the wrists and hands in flexion.
Front narrow (from a fourth to less than a fifth the greatest
breadth of the carapace) more or less deflexed, its edge shaped’ like
cupid’s bow (7.e., not bilobed).
1898. ] A. Alcock — Carcinological Fauna of India. 95
Orbital margin with the three suture-lines near the outer angle
fine and faint but distinct: eyestalks short and thick, eyes small.
Antennules folding transversely, inter-antennulary septum broad.
Basal joint of antenne short, touching the front only at their
antero-external augle; flagellum lodged in the orbital hiatus, short (less
than the major diameter of the orbit). |
, Merus of external maxillipeds with the anterior border almost
transverse.
Chelipeds subequal in both sexes; fingers pointed, not distinctly
hollowed at tip.
Legs with the upper border of the merus carpus and propus, and
the lower border of the merus and propus, sharply carinate or cristate.
Abdomen of the male five-jointed, the 3-5th somites being fused.
Medium-size and large crabs.
Key to the Indian species of Atergatis.
I, Edge of antero-lateral borders of carapace sharp and
crest-like, forming a tooth or ridge atthe lateral epibran-
chial angle :—
i. Carapace with an even surface, without indications
of regions :—
1. Surface of external maxillipeds almost
devoid of hair; no comb-like tufts of hair
on the legs ae side «. A, integerrimus.
2. Surface of external maxillipeds like a door-
mat; comb-like tufts of thick hair on the
under surface of some of the joints of the
legs sae bt .. A, dilatatus.
ii, Carapace with the surface somewhat lumpy;
variegated with spots and confluent blotches ... A, floridus.
II. Edge of antero-lateral borders of carapace thick and
blunt; no ridge or tooth at the lateral epibranchial
angle a a ait ie .. A, roseus,
20, <Atergatis integerrimus ( Lam.)
Cancer integerrimus, Lamarck, Hist. Nat. Anim. sans Vert. V. Crust. p. 272:
Milne Edwards, Hist. Nat. Crust. I. 874 and in Cuvier’s Régne An. Crust. pl. xi
bis., fig. 1.
Atergatis integerrimus, De Haan, Faun. Japon. Crust. p. 45, pl. xiv. fig. 1:
Dana U. S. Expl. Exp. Crust. pt. I. p. 158: Stimpson, Proc. Ac. Nat. Sci. Phila.
1858, p. 32: A. Milne Edwards, Nouv. Arch. du Mus. I. 1865, p. 235: Miers, Ann.
Mag. Nat. Hist. (5) V. 1880, p. 231: Richters, in Mobius Meeresf. Maurit. p. 145:
F. Muller, Verh. Ges. Basel, VIII. 1886, p. 474: de Man, Archiv. fur Naturges. LIII.
1887, i. p. 244, and Journ. Linn. Soc., Zool., XXII. 1888, p. 24, and Zool. Jahrbuch.
Syst. VIII. 1895, p. 496: A. O. Walker, Journ, Linn. Soc., Zool., XX. 1890 p. 109:
96 A. Aleock—Carcinological Fauna of India. [No. 1,
J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 352: Ortmann, Zool.
Jahrbuch Syst. VII. 1894, p. 462.
Atergatis subdivisus, White, Ann. Mag. Nat. Hist. (2) 1848, p. 284, and
Samarang Crust. p. 38, pl. viii. fig. 3. .
? Atergatis subdentatus, De Haan, Fann. Jap. Crust, p. 46, pl. iii. fig. 1: A. Milne
Edwards, Nouv. Archiv. du Mus. I. 1865, p. 236.
Carapace, length from about 4+ to about 2 the breadth; its surface
in the anterior third or half irregularly and rather distantly pitted,
especially near the front and antero-lateral. borders: except for two
faint creases that partly show the cardiac region, there are no other
traces of regional divisions.
The crest-like edge of the antero-lateral border turns in at the ©
lateral epibranchial angle to form a stout ridge there: this edge some-
times shows traces of two or three fissures.
The front, which is little prominent, meets the antero-lateral borders
at a wide but very distinct angle.
Orbits very small, their width being much less than a third the
width of the front. .
Surface of the external maxillipeds either quite smooth or with
short and scanty hair. Sternum smooth to the naked eye, or with a
little scattered pitting.
Chelipeds equal ; the upper edge of the merus sharply, the upper
edge of the hand and finger strongly but more bluntly, crested; the
upper outer surface of the hand with some scale-like roughening.
The outer surface of the legs is hardly pitted ; the claw, in all the
legs, is hairy, and there is a little tuft of hair near the far end of the
lower edge of the propodite, but all the other joints are generally free
of hair. |
Colours in spirits, pinkish ochre, fingers blackish brown, with whitish
tip and teeth.
30 specimens from Mergui, the Andamans, Ceylon and Singapore.
21. Atergatis dilatatus, De Haan, A. Milne Edwards.
Atergatis dilatatus, De Haan, Faun. Jap. Crust. p. 46, pl. xiv. fig. 2: A. Milne
Edwards, Nouv. Archiv. du Mus. I. 1865, p. 238, and Nouv. Arcuiv. pu Mus. IX.
1873, p. 1838, pl. v. fig. 6: E. Nauck, Zeits. Wiss. Zool. XXXIV. 1880, p. 57, pl. i.
figs. 19-21 (gastric teeth) : F. Miller, Verh. Ges. Basel, VIII. 1886, p. 474: Henderson,
Trans. Linn. Soc, Zool. (2) V. 1893, p. 353.
Closely resembles A. integerrimus, but easily recognized by the
following constant differences :—
(1) the carapace is even broader, and has a sharper edge:
(2) thesurface of the carapace is much more closely and extensively
1898.] A. Alcock — Carcinological Fauna of India. 97
pitted, no part being free from pitting except a small area in the mid-
gastric region: the outer surface of the chelipeds and legs also is quite
rough, from pitting :
(3) the creases that serve to show the cardiac region are much
larger and deeper :
(4) the crest that bounds the endostome in front is higher :
(5) the external maxillipeds are closely covered, like a door-mat,
with long thick bristles; and remarkable comb-like tufts of long stiff
bristles are found along the front border of the ischium of the chelipeds
and along the lower border of the ischium and merus of all the legs:
(6) the surface of the sternum is closely covered with confluent
granules visible to the naked eye.
All these differences are to be noted in a large male with a carapace
119 millim. broad from Ceylon, in a medium-sized male (70 millim.
broad) from the Andamans, and in two small females (59 and 44 millim.
respectively ) from widely different parts of the Andaman group — these
four specimens being in the Indian Museum collection.
22. Atergatis roseus (Ruppell).
Carpilius roseus and marginatus Ruppell, 24 Krabben roth. Meer. p. 13, pl. iii
fig. 3 and pl. vi. fig. 7; p. 15, pl. iii. fig. 4.
Cancer roseus and marginatus, Milne Edwards Hist. Nat. Crust. I. 374, 375.
Atergatis roseus and marginatus, De Haan, Faun. Japon. Crust. p. 17 (names
only):
Atergatis roseus, Heller SB. AK. Wien, XLIII. 1861, p. 309: A Milne Edwards,
Nouv. Archiv. du Mus. I. 1865, p. 239: Kossmann, Reise roth. Meer. Crust. p. 19:
Richters in Mébius Meeresf. Maurit. p. 145: Haswell, Cat. Austr. Crust. p. 42:
Cano, Boll. Soc. Nat. Nap. III. 1889, p. 189: Ortmann, Zool. Jahrb., Syst., VII.
1894, p. 461.
Atergatis marginatus, Krauss, Sudafr. Crust. p. 28: Dana U. 8. Expl. Exp. Crust.
pt. I. p. 158: A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 240.
Atergatis laevigatus, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 241,
pl. xv. figs. 4-4a: Henderson, Trans. Linn. Soc. Zool. (2) V. 1893, p. 352.
' Atergatis scrobiculatus, Heller, Abhand. zool.-bot. Ges. Wien, XI. 1861, p. 5, and
SB. AK. Wien, XLIII. 1861, p. 310: A. Milne Edwards, Nouv, Archiv. du Mus. I.
£865, p. 242.
Carapace of much the same proportions as A. integerrimus, but
with a perfectly smooth dull surface ; no indication whatever of regions ;
the crest of the antero-lateral borders blunt and ending smoothly,
without any ridge or tooth, at the lateral epibranchial angle.
Front, orbits, external maxillipeds and legs as in A. integerrimus,
Fingers fluted, but upper edge of hand rounded, not crested.
Colours in spirit, brownish yellow, fingers blackish brown with
whitish teeth and tips.
22 specimens from Karachi, aia Madras coast.
J. mu. 13
98 A. Alcock — Carcinological Fauna of India. [No. l
23. Atergatis floridus, (Rumph).
Cancer floridus, Ramph, Amboinsch. Rariteitk. p, 16, pl. viii. fig. 5: Linnzeus,
Syst. Nat. (xii) p. 1041.
Cancer ocyroe, Herbst, Krabben, III. ii. 20, pl. liv. fig. 2: Milne Edw. Hist. Nat,
Crust. I. 375.
Atergatis floridus, De Haan, Faun, Jap. Crust.. p. 46: Krauss, Sudafr. Crust.
p. 27: Dana, U. S. Expl. Exp. Crust. pt, I. p, 159, pl. vii. fig. 4: Stimpson, Proc.
Ac. Nat. Sci. Phila. 1858, p. 32: Heller, Novara Crust. p. 8: A. Milne Edwards,
Nouv. Archiv. du Mus. I. 1865, p. 248; and IX. 1873, p. 186: Targioni Tozzetti,
Magenta Crost. p. 24: Miers, P. Z. 8. 1877, p. 133, and Ann. Mag, Nat. Hist. (5) V.
1880, p. 231, and Zool. H. M. 8. Alert, pp. 182, 207, and Challenger Brachyura,
p. 112: Haswell, Cat. Austral. Crust. p. 41; F. Muller, Verh. Ges. Basel VIII. 1886,
p. 474: de Man, Arch. fur Naturges. LIII. 1887, i. 245, and Journ. Linn, Soc. Zool.,
XXII. 1888, p, 24, and Weber’s Zool, Ergeb. Niederl. Ost.-Ind. II. 1892, p. 277:
A. O. Walker, Journ. Linn, Soc., Zool., XX. 1890, p. 109: J. R. Henderson, Traus.
Linn. Soc., Zool., (2) V. 1898, p. 852: Ortmann, Zool. Jahrb. Syst. VII, 1894,
p. 460, and in Semon’s Forschungsr, (Jena. Denkschr. VIII) Crust. p. 51.
Carapace, length about 5%, breadth ; its surface of perfectly smooth
texture, but rendered te by broad shallow depressions that faintly
define and subdivide the regions; the crest of the antero-lateral border
is sharp and ends at a very distinct tubercle at the lateral epibranchial
anyle.
The front forms with the antero-lateral borders a semicircle. The
orbits are rather large, their width being more than one-third that of
the front.
Hxternal maxillipeds free from hair on the surface: the sternum
and the surface of all parts of the appendages except the hands (which
have some rough reticulations on the outer surface) are smooth,
Chelipeds equal, the upper edge of the merus and hand _ strongly
and sharply carinate; the fingers fluted as usual. Legs with crested
edges to the long joints, as in A. integerrimus.
Colours in spirit, yellow; carapace covered with symmetrically
disposed brown spots and confluent blotches, chelipeds and legs with a
few faint brown spots, fingers blackish with whitish teeth and tips,
86 specimens from the Andamans, Mergui, Ceylon, Laccadives and
Karachi.
24, Atergatis sp.
There is, in the Indian Museum, a little specimen of an Atergatis,
which may perhaps be the A. asperimanus insufficiently characterized
by White in the P. Z. 8S. 1847, p. 224 and in the Annals and a
of Natural History, 1848, Vol. II. p. 285, as follows :—
“‘ Carapace with its latero-anterior sides with a cutting edge, part
1898.] A. Alcock — Carcinological Fauna of India. 99
“of carapace behind this punctate; the rest of the surface almost quite
“smooth with three or four impressed lines in front. |
‘“ Hands rugose, especially above; fingers, both movable and _ fixed,
“deeply channelled.
“* Pale yellowish red ; feet darker ; fingers pale horn-coloured.
“ Philippines.”
Our little specimen, from off Ceylon, 34 fms., agrees with this des-
cription; but the ‘‘impressed lines,” which define the gastro-cardiac
region, are so faint as to be only just visible.
It has the crested legs of Atergatis.
Loppacraa, A. Milne Edwards.
_ Lophactewa, A. Milne Edwards, Ann. Sci. Nat. Zool. (4) XVIII. 1862, p. 43; and
Nouv. Archiv. du Mus. I. 1865, p. 245, and IX. 1873, p. 187.
Lophactea, Miers, Challenger Brachyura, p. 113.
“The Lophacteas are distinguished from Atergatis by their narrow-
er and always deeply lobulated carapace.”
Carapace moderately broad, convex in both directions, with the
regions generally well delimited and subdivided into lobes, and the
surface generally (not always) granular; the antero-lateral borders
have an independent crest-like edge, generally thin and sharp and dis-
tantly fissured ; the postero-lateral borders are rather concave.
Front a little deflexed, about a fourth the greatest breadth of the
_ carapace in extent, grooved and emarginate in the middle line, but not
distinctly bilobed, Orbits large, the three suture lines near the outer
angle distinct. Eyes on short thick stalks,
Antennules folding nearly transversely, inter-antennulary septum
broad. Basal joint of antenne short, touching the front only; the
flagellum lodged in the orbital hiatus. Merus of the external maxil-
lipeds with the front edge a little oblique.
Chelipeds equal in both sexes; fingers not hollowed at tip. Long
joints of legs with sharp cr est-like upper borders much as in Atergatis,
Abdomen of the male five-jointed, the 3rd—5th somites being
fused.
Small crabs. |
Lophacteea, except that the fingers are pointed instead of broad and
hollowed-out at tip; appears to me to be as closely as possible related to
Zozymus. In Lophactza, besides the difference in the fingers, the cara-
pace is more convex and less cut up into lobules, and its surface is
generally granular.
100 A. Alcock— Oarcinological Fauna of India. [ No. f,
Key to the Indian species of Lophacteea.
I. Regions and sub-regions of the carapace very distinct ;
postero-lateral borders slightly concave, but not de-
finitely marked off from the rest of the carapace :—
i. Surface of carapace more or less covered with
pearly granules :—
1. Hand sharply crested along upper border :—
a. Pearly granules over the whole of the
carapace, and over the outer surface
of the carpus and propus of the walk-
Ing lege <syids Gah iarehvenea oe .. DL. cristata.
b. Pearly granules whacnt fom pace “of
the post-cardiac region and from the
walking legs.. Voaees sesesecssee L. Semigranosa-
2. Upper border of hand das aeaae —
a. Crest-like margin of antero-lateral
border of carapace simply cleft ...... L. granulosa.
[b. Crest-like margin of carapace deeply
CTOdOG) i y.nehs.ce. sues dadbaeswn) ig fUSiely
ii. Surface of carapace and of S hciidages pefecty
SIMIC... :..<0nessane>ns shawsnascanwew nner we ee 6, anaglypta.
II, Regions and mn aoe of ‘tne carapace so faint as to
be visible only on close inspection ; postero-lateral
borders remarkably concave, and defined by a row of
sharp beads or teeth .........sesecsssscsseeserersetersesssscervee L. corallina.
25. Lophactea cristata, A. Milne Edwards.
Lophactza cristata, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1869, p. 246,
pl. xvi. fig. 1: de Man, Notes Leyden Mus. III. 1881, p. 95, and Arch. fur Naturges.
LIII. 1887. i. p. 246: F. Miller, Verh. Ges. Basel VIII. p. 474: Ortmann, Semon’s
Forschungsr. (Jena. Denkschr. VIII.), Crustacea, p. 50.
Carapace symmetrically intersected by broad smooth rather deep
furrows, which delimit and subdivide the regions, the strongly marked
convexities of the regions and subregions being closely studded with
pearl-like granules : similar, but larger, granules occur in linear series
on the outer surface of the wrist and hand; and similar, but smaller,
granules are found on the outer surface of the corresponding segments
of the legs. The under surface of the carapace is finely granular and
more or less furred,
The whole supra-orbital border is tumid, with a row of pearly
granules.
The crest of the antero-lateral border is divided into four broad
segments by three narrow fissures.
Upper border of the arm and hand strongly and sharply crested,
fingers fluted.
1898. ] A, Alcock — Carcinological Fauna of India. 101
Legs with a few scattered bristles on most of the joints, and with
the claws covered with short fur: the upper edge of the merus carpus
and propus is strongly crested, as are the lower edges of the merus.
Colours in spirit, yellowish or greenish brown, fingers blackish
brown.
One specimen from the Madras coast is in the Indian Museum
collection. (There are other specimens from Mauritius).
2€. Lophactsxa semigranosa, (Heller) A. M. Edw.
Atergatis semigranosus, Heller, Abhand. zool.-bot. Ges. Wien, XI. 1861, p. 6, and
SB. AK. Wien, XLIII. 1861, p. 313.
Lophactza semigranosa, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865,
p. 248: Miers, Zool. H. M.S. Alert, pp. 517 and 527: de Man, Archiv. fur Naturges.
LIILI. 1887, i. p. 246, pl. viii. fig. 4: J. R. Henderson, Trans. Linn. Soc., Zool., (2) V.
1893, p. 355: Ortmann, Zool. Jahrb. Syst. VII. 1894, p. 459.
Closely resembles LZ. cristata Heller, from which it is, perhaps, not
specifically distinct. It differs chiefly in having the pearly granules not
only less sharply sculptured aud less closely studded, but quite absent
from a part of the post-cardiac region, from the supra-orbital border,
and from the outer surface of the walking legs. Its appearance, in
short, is that of DL. cristata with the sculpturing worn.
7 specimens from the Andamans, Mergui, and Ceylon.
27. Lophactzxa granulosa, Riippell, A. M. Edw.
Xantho granulosus, Riippell, 24 Krabben roth. Meer. p. 24, pl. v. fig. 3.
Aegle granulosus, De Haan. Faun. Japon. Crust. p. 17 (name only).
Cancer limbatus, Hdw., Hist. Nat. Crust. I. 377, pl. xvi. fig. 14.
Atergatis limbatus, Dana, U. 8. Expl. Exp. Crust. pt. I. p. 157: Heller, Novara
Crust, p. 8: Streets, Bull. U. S. Nat. Mus. VII. 1877, p. 105.
Lophactza granulosa, A. Minne Epwarps, Nouv. Arcuiv. pu Mus. I. 1865,
p. 247, and 1X. 1873, p. 187: Brocchi, Ann. Sci. Nat. (6) II. 1875. Art. 2, p. 71,
pl. xvii. fig. 138 (male appendages): Hilgendorf, MB. AK. Berl. 1878, p. 787:
de Man, Notes Leyden Mus. III. 1881, p. 95, and Archiv. fur Naturges. LILI. 1887, i.
p. 246: Haswell, Cat. Austr. Crust. p. 43: Miers, Challenger Brachyura, p. 114:
Cano, Boll. Soc. Nat. Nap. III. 1889, p. 190: J. R. Henderson, Tr. Linn. Soc., Zool.,
(2) V. 1893, p. 354: Ortmann, Zool. Jahrb. Syst. &c. VII. 1894, p. 459.
Closely resembles the two preceding species, from which it differs
most conspicuously in having no crest to the upper border of the hand:
the granulation of the carapace is not nearly so sharp-cut and pearl-like.
Iu the Indian Museum are specimens from Australia and Samoa, but
none from India. It is included in the Indian Fauna on the authority
of Dr. J. R. Henderson.
102 A. Alcock — Car'cinological Fauna of India. [ No. 1,
28. Lophactxa anaglypta (Heller), A. M. Edw.
Atergatis anaglyptus, Heller, Abhandl. zool.-bot. Ges. Wien, 1861, p. 6, and SB.
Ak. Wien, XLIIL. 1861, p. 312, pl. ii. figs. 11, 12.
Lophactxa anaglypta, A. Milne Edwards, Nouv. Archiv..du Mus. I. 1865, p. 251,
and IX. 1873, p. 190: Ortmann, Zool. Jahrb, Syst. VII. 1893, p. 459: de Man, Zool.
Jahrb, Syst. VIII. 1895, p. 498.
Carapace with the regions separated and symmetrically subdivided
by broad but well cut grooves, but with the texture of the surface—as
of the appendages—perfectly smooth, the only roughness of any sort
being a few lines and impressions on the outer surface of the hand..
Crest of the antero-lateral border narrow, divided into four lobes
by three insignificant notches or dents, Supra-orbital border not tumid
throughout its extent. Crest of the upper border of the hand low and
rather blunt: crests of the leg-joints distinct but rather low.
One specimen from Galle, one from the Persian Gulf.
29. Lophactxa corallina, n. sp.
Carapace broadly semioval, with remarkably concave postero-lateral
borders, the crest of the antero-lateral border very thin and sharp and
a little angular, the postero-lateral and posterior borders bounded by a
line of sharp beads or teeth. Frout obliquely deflexed, with a sharp
broadly-bilobed edge.
The whole surface of the carapace is very finely granular, but the
division and subdivision of the regions, though undoubtedly existent,
is hardly perceptible, so faint are the inter-regional depressions : some
long stiff hairs occur here and there.
The under surface of the carapace and the surface of the external
maxillipeds and male sternum is finely granular. |
The chelipeds and legs are rather hairy and are beautifully sculp-
tured: at the distal end of the arm is a petal-like crest, and three series
of larger petaloid granules or crests traverse the outer surface of the.
wrist longitudinally : the outer surface of the hand is closely granular,
the granules becoming linear in arrangement and laminar in a to-.
wards the upper part.
The outer surfaces of the legs are covered with granules and teeth,
two crests on the carpopodites of all being very distinct.
Colours in spirit, yellowish or whitish with a pink blush: fingers
with a black cross-band at the base. |
Length of carapace 6 millim., breadth 9 iii,
A male and female from off Gevinee 34 fms.
1898. ] A. Alcock — Carcinological Fauna of India. 103
30. Lophactzxa fissa, Henderson.
- Lophactwa fissa, Henderson, Trans, Linn. Soc., Zool., (2) V. 1893, p. 355, pl.
xxxVi. figs. 8, 8a.
It appears to me possible that this, which seems to be founded on
a single specimen, is only an individual variation of DL, granulosa.
Lophactzxa sp,
From Inglis I. (Andamans) a single small specimen, not agreeing
with any described species, which in the circumstances I forbear to
describe,
It belongs to the UL. cristata and semigranosa group, but has the
inter-regional furrows much shallower and less distinct, and the pearly
granules absent from all but the front part of the gastric region and
the lateral parts of the epibranchial region: those on the chelipeds are
also much fewer aud more scattered. The legs are very hairy.
Zozymvus, Leach.
Zozymus, Leach, [Dict. Sci. Nat. XII. p. 75. Miers]: and in Desmarest, Consid.
Gen. Crust. p. 105.
Zozymus, Milne Edwards, Hist. Nat. Crust. I. 383 (part).
Zozymus, Dana, U. 8. Expl. Exp., Crust. pt. I. p. 189.
Zozymus, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX. 1863, p. 302.
Zozymus, Miers, Challenger Brachyura, p. 134,
Carapace moderately broad, moderately convex in both directions,
with the regions well delimited and subdivided into numerous lobules,
the surface of which is not usually granular.
The antero-lateral borders are sharp and crest-like, and are cut into
lobes (usually four in number) ; the postero-lateral borders are straight
and strongly convergent.
Front about a fourth the greatest breadth of the carapace, obliquely
deflexed, grooved and emarginate in the middle line: orbits large, the
tumid edge with the three suture lines near the outer angle distinct ;
eyes on short thick stalks.
Antennules folding nearly transversely, inter-antennulary septum
broad. Basal joint of antenne short, touching the front only at the
(produced) antero-internal angle; the flagellum short (less than the
major diameter of the orbit), lodged in the orbital hiatus.
Merus of the external maxillipeds with the front edge a little
oblique.
Chelipeds equal in both sexes; fingers with broad hollowed-out
104 A. Alcock — Carcinological Fauna of India. [No. 1,
tips. Long joints of legs with sharp crest-like upper borders much as
in Atergatis.
Abdomen of the male five-jointed, the 3rd—5th somites being fused.
Rather large crabs.
Key to the Indian species of Zozymus.
I. All parts of carapace rugose : inter-regional and inter-
lobular furrows smooth and naked except, perhaps,
near the margin of the carapace oe 4, aeneus,
II. Posterior third of carapace hardly at all rugose: inter-
regional and inter-lobular furrows for the most part
full of short close hair SH sate .. Z. pilosus.
31. Zozymus aeneus, (Linn.)
Cancer incomparibilis, Seba, Thesaurus IIT. 48, pl. xix. fig. 18.
Cancer aeneus, Linn., Mus. Lud. Ulr. p. 451, and Syst. Nat. (ed. xii) p. 1048.
Cancer floridus, Herbst, Krabben, I. ii. 132, 264, pl. iii. fig. 39, pl. xxi. fig. 120.
Cancer amphitrite, Herbst, Krabben, IIT. ii. 5, pl. liii. fig. 1.
Cancer aeneus and floridus, Fabricius, Ent. Syst. II. 455, 445, and Suppl.
p. 335, 338.
Cancer aeneus, Latreille, Hist. Nat. Crust. V. 375: Lamarck, Hist. Nat. Anim.
sans Verteb. V. 271: Desmarest, Consid. Gen. Crust. p. 104: [Quoy et Gaimard,
Voy. Uranie, pl. lxxvi. fig. 1. Edw.].
Zozymus aeneus, Milne Edwards, Hist. Nat. Crust. I. 385,
Aegle aeneus, De Haan, Faun. Japon. Crust. p. 17.
Zozymus aeneus, Dana, U.S. Expl. Exp. Crust. pt. I. p. 192, pl. x. fig. 3:
Stimpson, Proc. Ac. Nat. Sci. Phila. 1858, p. 32: Heller, SB. AK. Wien, XLIII.
1861, p. 326: A. Milne Edwards, in Maillard’s Vile Réunion, Annexe F. p. 4, and
Nouv. Archiv. du Mus, IX. 1873, p. 207: Miers, Ann. Mag. Nat. Hist. (5) II. 1878,
p. 407, and Phil. Trans. Vol. 168, 1879, p. 486, and Ann. Mag, Nat. Hist. (5) V. 1880,
p. 234, and Challenger Brachyura, p. 184: Richters in Mébins Meeresf. Maurit.
p. 146: Haswell, Cat. Austral. Crast. p. 58: F. Muller, Verh. Ges. Basel VIII.
p. 474: de Man, Archiv. fur Naturges. LIII. 1887, i. p. 273: Cano, Boll. Soc. Nat.
Napoli III. 1889, p. 199: J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893,
p. 859: Ortmann, Zool. Jahrb. Syst. VII. 1893-94, p. 458, and Semon’s Forschungsr.
(Jena. Denk. V1II) Crust. p. 50: Whitelegge, Mem. Austral. Mus. IIL. 1897, p. 181.
Carapace with the regions well delimited and very strongly rugose,
the rugosities being small in the postero-lateral regions but every where
smooth and polished, and everywhere symmetrical. The crest-like
antero-lateral borders are 4-scalloped, the three anterior lobes being
rounded and the fourth being dentiform.
The tumid orbital margins are marked by four suture lines. The
front hardly projects beyond the level of the orbit.
The wrist has its outer surface made rugose by meandering
furrows, one of which runs fore and aft, the others transversely. The
1898. | A. Alcock —Carcinological Fauna of India. 105
hand carries a blunt but well-marked crest along its upper border,
below which the surface is rugose much as the wrist: the lower part
of the outer surface of the hand is tuberculous, the tubercles tending
to a linear arrangement. The fingers are fluted, bear strong molariform
teeth and tufts of hair on their cutting edge, and have blunt-pointed,
hollowed out (spoon-like) tips. The furrows of the wrist and hands, as
well as those of the legs, are filled with close short fur.
The merus carpus and propus have the upper edge strongly cari-
nate, the inner surface of each crest bearing a thick fringe of long
somewhat silky hair: the dorsal surface of these joints is furrowed
longitudinally, with many more or less plain transverse impressions
also: the dactyli are hairy up to the claw.
In life the animal is beautifully spotted and ocellated with chocolate
brown on a bluish-grey ground. In spirit the animal has a chinaware
look and a dull yellowish-white colour, with darker yellow and dall
brownish spots and markings.
In the Indian Museum are 30 specimens from the Andamans and
Laccadives.
32. Zozymus pilosus, A. Milne Edwards.
Zozymus pilosus, A. Milne Edwards, Ann. Soc. Ent. France (4) VII. 1867, p.
271; and Nouv. Archiv. du Mus. IX. 1878, p. 208, pl. vii. fig. 2.
Carapace having the regions and lobules well defined in its anterior
two-thirds only : the lobules have a flattened semi-imbricate look, wavy
edges, and a rough or granular surface; and the grooves that separate
them are filled with small short close-set bristles, especially along the
anterior contours of the lobules,
All four lobes of the antero-lateral borders are rounded and not
dentiform. The orbital margin is not very tumid and is marked by
three suture lines. The front projects beyond the orbit.
The wrist and hand are closely nodular: the nodules (those on the
hand especially) have a granular surface, and the grooves that separate
them are full of short close hair: the upper edge of the hand is not
crested. Fingers short, stout, blunt-pointed, hollowed at tip: they are
strongly fluted, the ridges being beaded in their basal half.
The legs have the upper edge of the merus, carpus, and propodite
strongly crested: the crest of the merus and carpus may be subserrate,
and is always notched near the distal end. The dorsal surface of the
carpus and propodite is grooved and nodular—the nodules having a flat,
subimbricate look.
Colours in spirit—yellowish-white, with a faint bluish or purplish
blush ; the crest-like margin of the carapace lighter than other parts;
fingers dark brown with white tips.
J. ui, 14
106 A. Aleock — Careinological Fauna of India, { Nowa;
In the Indian Museum is a specimen from Port Blair Harbour
(Andamans), and one from the Angrias Bank (Malabar Coast) in
15 fms.
Although the chelipeds and antero-lateral margins make this
species easily recognizable from Lophozozymus incisus (Kdw.) de Man, I
am inclined to suspect that this is the young of L. incisus.
Lopnozozymus, A. Milne Edwards.
Lophozozymus, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX. 1863, p. 276 ;
and Ann. Soc. Entom. France (4) VII. 1867, p. 272.
Lophozozymus, Miers, Challenger Brachyura, p. 114,
Differs from Zozymus in having (1) the crest of the antero-lateral
border sharp-edged and (2) the fingers not spooned at tip. To avoid
unnecessary disturbance of accepted nomenclature the name is here
maintained as a subgenus of Zozymus.
Key to the Indian species of the subgenus Lophozozymus.
I. Front lobe of the antero-lateral border confluent and
fused with the orbit: [size moderate or small ]:—
. Regions and lobules of carapace well defined ; the
lobules pitted or dented, the grooves between
them hairy: chelipeds and legs shaggy ............ DL. wmecisus.
i. Regions and lobules of carapace ill defined and
faint: surface of carapace smooth and bare: legs
with a few lank scattered hairs ... - .... wsseee Le. dodone.
II, Front lobe of antero-lateral border soph indl Sesith the
orbit by a gap; carapace smooth, the regions (but not
the subregions) fairly well defined : [size large ]:—
i. Hands smooth and bare .., ....... cecovcoseesees L, octodentatus.
ii. Outer surface of hand granular es ie Eth ae L. cristatus.
33. Lophozozymus octodentatus, Edw.
Cancer saxatilis, Ramph, Amboinsch. Rariteitk. p. 9, pl. v. fig. M.
Cancer rumphii, Guérin, Icon. Régne An. pl. ii. fig. 1, (nec Herbst.)
Xantho octodentatus, Milne Edwards, Hist. Nat. Crust. 1. 398: Lucas in
Jacquinot’s Voyage Astrolabe, Zool., Crust., p. 23, pl. ix. fig. 1: E. Nauck, Zeits.
Zool. XXX1V. 1880, p. 51 (gastric teeth): Haswell, Cat. Austral. Crust. p. 58.
Lophozozymus epheliticus Linn., Miers, Ann. Mag. Nat. Hist. (5) V. 1880, p. 231,
and Zool. H. M.S. Alert, pp. 182, 207: A. O. Walker, Journ. Linn. Soc., Zool., XX.
1890, p. 109: de Man, Zool. Jahrb., Syst., VIII. 1895, p. 518.
Carapace perfectly smooth and polished; the gastric region de-
limited on all sides and partly subdivided, and the hepatic separated
from the branchial regions, by broad smooth shallow depressions.
Underside of carapace hairy,
1898.] A. Aleock —Carcinological Fauna of India. 107
Front gently convex beyond the orbits, finely cleft in the middle
line. Orbital border sharp, somewhat puckered by three sutures near
the outer angle. The crest-like antero-lateral border is sharp and is
cut into four teeth, of which the first is separated from the orbit by a
gap, the third and fourth are keeled, and the third strongly accuminate.
Chelipeds equal, their surface perfectly smooth: the upper edge
of the arm is strongly crested, the crest at its distal end being foliace-
ously expanded and deeply cleft; wrist with a strong double tubercle
at its inner angle; upper edge of hand crested, but rather coarsely ;
fingers large, long, pointed. —
Legs smooth: upper edge of merus carpus and propodite strongly
crested, the inner face of the crest with tufts of long hair; dactylus
furred up to the claw.
Colours in spirit: a bright orange-red network on a dull yellow-
ochre ground, fingers black.
In the Indian Museum are two specimens from Singapore.
34. Lophozozymus cristatus, A. Milne Edwards.
Lophozozymus cristatus, A. Milne Edwards, Ann. Soc. Entom. France (4) VII.
1867, p. 272, and Nouv. Archiv. du Mus. IX. 1873, p. 203, pl. vi. fig. 4: J. R. Henderson,
Trans. Linn. Soc., Zool., (2) V. 1893, p. 361: Ortmann, Zool. Jahrb., Syst., VII.
1893-94, p. 456.
This species appears to differ from LD. octodentatus only in having
the first lobe or tooth of the antero-lateral margin acute instead of
rounded and the outer surface of the hands granular and hairy.
There are no specimens in the Indian Museum; and the species —
if it be distinct —is included in the Indian fauna on the authority of Dr.
J. R. Henderson.
35. Lophozozymus incisus (Edw.) Haswell, de Man.
Xantho incisus, Milne Edwards, Hist. Nat. Crust. I. 397: Hess, Archiv. f. Nat.
XXXI. 1865, i. p. 183: F. Muller, Verh. Ges. Basel, VIII. 1886, p. 474.
? Xantho superbus, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 74, and U. S. Expl.
Exp. Crust. pt. I. p. 167, pl. viii. figs. 5a-b, (nec A. Milne Edwards) de Man.
Lophozozymus incisus, Haswell, Cat. Austral. Crust. p. 58: de Man, Archiv. fiir
Naturges. LIII. 1887, i. p. 268, pl. x. fig. 1: Thallwitz, Abh. u. Ber. Mus. Dresden,
1890-91, no. 3, p. 48.
Carapace with the regions well delimited, and having the branchial
regions (and to a less extent, the gastric region also) subdivided into
lobules which have their anterior margins sinuous and sharply under-
mined so as to have a semi-imbricate look. The surface of these lobules
is a little dented and uneven, and the grooves that separate them are
full of hair.’
108 A. Alcock — Carcinological Fauna of India. [No. ,
The crest-like antero-lateral border is cut into 4 lobes, of which
the first is confluent with the orbit, and the last two are pointed and
strongly keeled. Front little convex beyond the orbits, distinctly bilob-
ed. Orbital border sharp, salient, with three suture-lines.
Chelipeds equal: upper edge of arm with a strong crest, which is
foliaceously expanded and cleft at its distal end ; upper border of hand
and dactylus crested ; outer surface of wrist and hand covered with large
granules, which stand in more or less distinct linear series and are a
good deal concealed by long shaggy hairs. Fingers stout, of good
length, pointed. |
Legs with the upper edge of merus carpus and propodite strongly
crested and shaggy, and the surfaces of the dactylus and of most of
the propodite shaggy.
Colours in spirit yellow with many orange-red patches; fingers
dark brown. 7
In the Indian Museum are 2 specimens, one from the Orissa Coast,
15-35 fms., the other from the Angrias Bank (Malabar Sea) 15 fms.
36. Lophozozymus dodone (Herbst) Hilgendorf, de Man.
Cancer dodone, Herbst, Krabben, III. ii. 37, pl. lii. fig. 5. :
Lophozozymus dodone, Hilgendorf, MB. Ak. Berl. 1878, p. 789: Miers, Zool.
H. M.S. Alert, pp. 517, 527: de Man, Archiv. fiir Naturges. LIII. 1887, i. p. 270,
pl. x. figs. 2, 2a: J. R. Henderson, Trans. Linn, Soc., Zool., (2) V. 1893, p. 361 :
Ortmann, Zool. Jahrb., Syst., VII. 1893-94, p. 457.
Xantho radiatus (? C. dodone Herbst) Milne Edwards, Hist. Nat. Crust. I. 398::
A. Milne Edwards, in Maillard’s l’ile Réunion, Annexe F. p. 4.
Atergatis lateralis, White P. Z. S. 1847, p. 225; Ann. Mag. Nat. Hist. (2) II. 1848,
p. 285; and Samarang Crust. p. 39, pl. viii. fig. 1.
Xantho lamelligera, White, ll. cc. p. 225, p. 285, p. 40 (fide A. Milne Edwards
infra).
Xantho nitidus, Dana, Proc. Ac. Nat. Sci. Phila. 1852, p. 74, and U. S. Expl.
Exp. Crust. pt. I. p. 166, pl. viii. figs. 4a-b.
Atergatis elegans, Heller, Novara Crust. p. 7, pl. i. fig. 4 (fide de Man).
Lophozozymus radiatus, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873,
p. 206.
Carapace smooth with the regions very faintly indicated and with
very few and faint traces of lobulation : sometimes a few lank hairs on
the antero-lateral border.
The crest-like antero-lateral border is trenchant and somewhat
cockled, and is cut into 4 shallow scallops, the last two of which are
acuminate and carinate, and the first of which is confluent with the
orbit. Front slightly convex beyond the orbits and a little emarginate
in the middle line. Orbital border sharp with the suture lines faint
and indistinct,
1898. ] A. Alcock— Carcinological Fauna of India. 109
Chelipeds equal; outer surface of wrist and hand finely granular
or rugose under the lens; upper edge of arm crested but not foliaceously
expanded; both upper and lower edge of hand crested. Fingers very
short and stumpy, pointed.
_ Legs smooth: upper edge of merus carpus and propodite crested
and having a few scattered hairs; also a few scattered hairs on the
surface and lower edge of propodites.
Colours in spirit yellow, with diffused orange-red patches; fingers
brown, white at tip.
In the Indian Museum are three specimens from the Andamans.
Alliance III. Huaanthoida.
EKuxanthus. Hypocoelus.
Evxantaus, Dana.
_ Euzanthus, Dana, Silliman’s Amer. Journ. Sci. and Art. (2) XII. 1851, p. 125;
Proc. Ac. Nat. Sci. Phila. 1852, p. 75; and U.S. Expl. Exp. Crust. pt. I. p. 178.
Melissa, Strahl, Archiv. fur Naturges. XX VII. 1861, i. p. 101.
Euzanthus, A. Milne Edwards, Nouv. Archiv, du Mus. I. 1865, p. 289,
Carapace very broad, strongly convex in both directions, with the
regions well delimited and subdivided into convex lobules.
The antero-lateral borders are sharp and somewhat irregularly
scallopped, the lobes often subpyramidal or dentiform: they do not
terminate at the orbit, but are prolonged, beneath the orbit, to the
buccal cavern. The postero-lateral borders are very short and very
concave.
The front is of no great breadth (about a fifth the greatest breadth
of the carapace), bilobed, and prominent. The supra-orbital border and
the inner angle of the lower border of the orbit are tumid, and the rest
of the orbital margin is very low and forms an unbroken curve, with
only one closed suture line. The eyes have short thick stalks.
The antennules fold nearly transversely. The basal antennal joint
is prolonged right into the orbit, and the short flagellum is therefore
placed inside the orbit. The outer border of the merus of the external
maxillipeds is oblique.
The chelipeds are equal in both sexes, and are relatively small and
light. The fingers are rather long-pointed, and have the tip slightly
but distinctly hollowed out.
| Abdomen of the male five-jointed, the 3rd—5th somites being fused.
Crabs of medium size, easily recognized by the peculiar form of the
basal joint of the antenna and the course of the antero-latera] margin
of the carapace. é
110 A. Alcock — Carcinological Fauna of India. [No. 1,
Key to the Indian species of Kuxanthus.
1. Lobules of carapace almost smooth; outer angle of
orbit not marked by # denticle ......ccccceccesecesecoeens E. melissa,
2. Lobules of carapace rough; outer angle of orbit
marked by @ denticle ...cceccc.secccceescoceees seeececes eo. E. sculptilis,
37. Huxanthus melissa, (Herbst).
Cancer exsculptus, Herbst, Krabben, I. ii, 265, pl. xxi. fig. 121.
Cancer melissa, Herbst, Krabben, III. ii. 7, pl. li. fle. 1.
Euasaanthus melissa, Stimpson, Proc. Ac. Nat. Sci. Phila, 1858, p. 33: A. Milne
Edwards, Nouv. Archiv. du Mus, I. 1865, p. 293: Targioni Tozzetti, Magenta Crust.
p. 27, pl. iii. figs. 1-7: F. Muller, Verh. Ges. Basel, VIII. p. 474: J. R. Henderson,
Trans. Linn. Soc., Zool., (2) V. 1893, p. 359: Ortmann, Zool. Jahrb. Syst. VII.
1893-94, p. 466, pl. xvii. fig. 9.
Cancer mamillatus, Miine Edwards, Hist. Nat. Crust. I. 376.
Melissa mamillata, Strahl, Archiv. fir Naturges. XX VII. 1861, i. p. 103.
Euzanthus mamillatus, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 292,
pl. xv. figs. 2-2b; and IX. 1873, p. 196: Haswell, Cat. Austral. Crust. p. 48: de Man,
Journ. Linn. Soc., Zool., XXII. 1887-88, p. 30.
Eusanthus nitidus, Dana, Proc. Ac. Nat. Sci. Phila. 1852, p. 75; U.S. Expl.
Exp., Crust. pt. I. p. 174, pl. viii. figs. 9a-b. (young).
' Melissa nitida, Strahl, Archiv. fiir Naturges. XXVIT. 1861, i. p. 103.
Cancer exsculptus, Hoffmann in Pollen and Van Dam, Faun. Madagasc. Crust,
p- 38.
?Huxanthus exsculptus var. rugosus, Miers, Zool. H. M. 8S. Alert, pp. 517, 527
(? young.)
The lobules of the carapace are extremely convex, and though
some of them may be a little dimpled, especially in the young, they are
commonly smooth.
The antero-lateral borders are cut into five teeth, but there is often
a tubercle—which may be incompletely double—between the 4th and
Sth teeth ; between the 3rd and 5th teeth the margin is finely granular.
The curve of the orbit is unbroken by any denticle at the outer
angle, and is smooth, not granular.
The outer surfaces of the wrist and hand, as of the corresponding
joints of the legs, are nodular, the nodules and the hollows between them
being smooth: on the lower outer surface of the hand are two longi-
tudinal wrinkles which also have a smooth surface. The fingers have
their surfaces smooth, and their cutting edges strongly toothed, with
the tip distinctly hollowed out.
Colours of good spirit specimens: stone grey or yellowish, with
numerous tiny chocolate-brown or purplish specks, and some large
blotches of the same colour on the gastric, hepatic and branchial regions.
These markings have faded in spirit specimens that have been preserved
1898. ] A. Aleock — Carcinological Fauna of India. 11}
over ten years, Fingers blackish brown, this colouration extending
along the lower border and inner surface of hand.
In the Indian Museum are 17 specimens from the Andamans,
Mergui, and Ceylon (besides a specimen from Samoa).
38. Huxanthus sculptilis, Dana.
Euzanthus sculptilis, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 75, and U.S.
Expl. Exp. Crust. pt. I. p. 173, pl. vii. figs. 8a-d: A. Milne Edwards, Nouv. Archiv.
du Mus. I. 1865, p. 291: Ortmann, Zool. Jahrbnch., Syst., VII.{1893-94, p, 466.
Cancer huonii, Lucas in Jacquinot’s Voy. Astrolabe, Crust. p. 16, pl. iv. fig. 1.
Eucanthus huonit, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 290, pl.
xv. figs. 1-lc: Hasweil, Cat. Austral. Crust. p. 47: Miers, Zool. H. M.S. Alert, pp.
182, 204: de Man, Archiv. fiir Naturges. LIII. 1887, i. p. 263.
Lobules of carapace moderately convex, their surface so much
wrinkled aud dented transversely as to give them an almost scaly look.
Antero-lateral borders cut into six teeth, the edge between all the
teeth being granular.
The orbital margin is granular, and there is a denticle to mark the
outer angle of the orbit,
The nodules of the wrists and hands—and, to a less strongly
marked extent, those of the corresponding joints of the legs—are
granular, as are the hollows between the nodules; and the two wrinkles
along the lower outer surface of the hand are granular. The fingers
resemble those of H. melissa, except that their surfaces are strongly
granular.
Colours of well-preserved spirit specimens : yellowish with purplish
spots and blotches, many of which are confluent; fingers and hand
coloured as in H. melissa.
In the Indian Museum are 3 specimens from Persian Gulf and
Andamans, (besides one from Samoa).
Hypocauvs, Heller.
Hypocelus, Heller, Abh. zool-bot. Ges. Wien, 1861, p. 7; and SB. AK. Wien,
XLIII. 1861, p. 319.
Hypocelus, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865. p. 295,
Differs from Hu«anthus chiefly in having a large oval or reniform
cavity excavated in either pterygostomian region.
It is not represented in the Indian Museum.
39. Hypocelus rugosus, Henderson.
Hypocelus rugosus, Henderson, Trang. Linn. Soc., Zool., (2) V. 1893, p. 358,
pl. xxxvi. figs. 9-11.
112 A. Aleock—Carcinological Fauna of India. [No. 1,
Alliance IV. Xanthoida.
Xantho. Etisus. Orphnoxanthus.
Leptodius. Ktisodes.
Medzeus.
Cycloxanthus.
Hoploxanthus.
XAntTHO, Leach.
Xantho, Leach, Malac. Pod. Britt. pl. xi and text, 1815; and Trans. Linn. Soc,
XI. 1815, p. 820.
Xantho, Desmarest, Consid. Gen. Crust., p. 104.
Xantho, (part) and Eudora (part), De Haan, Faun. Japon. Crust. pp. 18 and 22.
Xantho, (part) Milne Edwards, Hist. Nat. Crust. I. 887.
Xantho, (part) Dana, U. 8S, Expl., Exp., Crust. pt. I. p. 166.
Xantho, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX. 1863, pp. 275 and 294
and Miss. Sci. Mex., Crust. p. 251.
Xantho, Miers, Challenger Brachyura, p. 124.
[Type XanrHo Fioripus, Leach. ]
Carapace broad, moderately convex anteriorly, flat in the posterior
half; the regions generally well delimited and fairly well lobulated in
the anterior two-thirds, but not posteriorly.
Antero-lateral borders arched, usually cut into four teeth or lobes:
postero-lateral borders moderately convergent, not concave.
Extent of fronto-orbital border half, or less than half, the greatest
width of the carapace: front about a fourth the greatest breadth of the
carapace. |
Front little deflexed, rather prominent, usually sublaminar, notched
in the middle line, usually separated from the supra-orbital margin by a
notch or groove.
Orbital margin with two (often indistinct) suture lines above and
one (more distinct) just below the outer angle: usually ‘a prominent
tooth at the inner angle of the lower edge of the orbit, Eyes on short
thick stalks.
Basal antennal joint short, meeting the front at the inner angle:
the flagellum, which is about as long as the orbit, lodged in the orbital
hiatus.
Anterior edge of merus of external maxillipeds nearly transverse,
with commonly a small tooth near the antero-internal angle.
Chelipeds either unequal in both sexes, or less commonly equal in
both sexes (Xantho impressus, Xantho scaberrimus); fingers pointed.
Legs subcylindrical, with the upper edges often sharp (crested in
Xantho scaberrimus.)
1898. ] A. Alcock — Carcinological Fauna of India. 113
Abdomen of male five-jointed, the 3rd—5th somites fused; (in
X. wmpressus the sutures are so distinct that the abdomen may appear
7-jointed),
| Key to the Indian species of the Genus Xantho.
I. Chelipeds equal, or almost equal, in both sexes :—
i. Legs crested, the crest sharp, or serrate, or
crenate: length of carapace a good deal
more than two-thirds the greatest breadth :
lobules of carapace covered with convex
subsquamiform tubercles .........-0000. seerereee Xantho (Lophowanthus)
scaberrimus.
F ii. Legs thick, sub-cylindrical, length of carapace
less than two-thirds the greatest breadth:
lobules of carapace Smooth ........s000c0e. Xantho (Eudora) im-
pressus,
1I. Chelipeds unequal in both sexes: length of carapace
two-thirds, or a little more than two-thirds, the
greatest breadth: the'legs may have sharp, but
never distinctly crested edges :—
i. First two teeth of the antero-lateral margin
faint, obsolescent ; carapace and chelipeds
smooth (non-granular)..,........0..06 Shaloecdueney’ Xantho bidentatus.
ii. Hither the last three, or all four, teeth of
the antero-lateral margin distinct; a large
part of the carapace and of the exposed
surfaces of the chelipeds wrinkled and
STANDIAL .....ccerecsessesserteecscevsceecerccesssessces Aantho distinguendus,
40. Xantho distinguendus, De Haan.
Cancer (Xantho) distinguendus, De Haan, Faun. Japon. Crust, p. 48, pl. xiii.
fig. 7: Heller, SB. Ak. Wien, XLIII. 1861, p. 323.
Chlorodius distinguendus, Stimpson, Proc. Ac. Nat. Sci. Phila., 1858, p. 34.
Xantho macgillivrayi, Miers, Zool. H. M.S. Alert, pp. 183, 211, pl. xx. fig. c.
Lophozozymus (Lophoranthus) bellus, var. leucomanus, Miers, Challenger
Brachyura, p. 115, pl. xi. fig. 1.
Medzus distinguendus, de Man, Journ, Linn. Soc., Zool., XXII. 1887-88, p. 31:
J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 359.
Regions of carapace well delimited, fairly well divided into lobules :
the anterior and lateral parts of the carapace are covered with granular
transverse wrinkles which have almost a scaly look.
Front cleft into two rather prominent, square-cut, sub-laminar
lobes. Antero-lateral margin cut into four sharply granular teeth,
‘which may be all sharply acuminate, or the first may sometimes be
rather indistinct. '
i, LS
L14 A. Alcock — Carecinological Fauna of India. [No. 1,
The under surface of the carapace, below and external to the orbit,
is, like the first lobe of the antero-lateral margin, eroded and granular
Chelipeds unequal in both sexes: upper part of the outer surface
of arm with some fine transverse granular wrinkles; upper and outer
surface of wrist and hand closely granular, the wrist and the upper part
of the hand being also eroded or pitted, most usually in a peculiar
honey-comb. fashion ; fingers stout, fluted.
Legs rather thin: upper edge of merus sharp (almost subcristi-
form), often finely granular: carpus and propodite usually grooved and
ridged longitudinally (the propodite most distinctly so, and on both
surfaces): dactylus covered with close short fur. The sculpture of the
carpus and propodite, as of the chelipeds, is variable, even in specimens
from the same locality.
Abdomen of male 5-jointed, the sutures between the 3rd—5th
somites nearly or quite obliterated.
Colours in spirit : light yellow, fingers blackish brown with whitish
tips.
In the Indian Museum besides specimens from Hongkong, there
are 16 specimens from Mergui, Persian Gulf, and Karachi. Fourteen
little specimens from the Malabar Coast, 28 fms,, are also probably
referable to this species.
Tf this species is to be removed to Medseus on account of the erosion
and consequent indefiniteness of the orbital end of the antero-lateral
margin, Xantho floridus and more certainly Xantho tuberculatus must
share the same fate, and Medzxus must then be absorbed in Xantho.
41. Xantho bidentatus, A. Milne Edwards.
Xantho bidentatus, A. Milne Edwards, Ann. Soc. Ent. France (4) VII. 1867,
p. 266: Miers, Challenger Brachyura, p. 126, pl. xi. fig. 4: Ortmann, Zool. Jahrb.
Syst., VLI. 1893-94, pp. 44.4, 449.
Surface of carapace and appendages smooth (non- -granular) :
gastric region well defined by fine shallow grooves, and very faintly
Jobulated: branchial regions imperfectly separated from the hepatic
regions and very faintly and imperfectly areolated.
Of the four lobes of the antero-lateral margin the first two are
faint, broadly-rounded and coalescent, and the last two possess a small
acumination.
Front prominent, notched in the middle line, to form two lobes, which
have the edge a little concave and the outer angle well pronounced,
Under surface of carapace smooth to naked eye: the side wall.
above the articulations of the legs hairy. ’
oe Poe
1898. } A. Alcock — Carcinological Fauna of India. 115
Chelipeds unequal in both sexes, smooth like the legs.
Abdomen of male five-jointed.
Colours in spirit : dull yellowish brown, fingers almost black.
In the Indian Museum are four specimens from the Andamans.
42. Xantho impressus, (Lamk.) Edw.
Cancer impressus, Lamarck, Hist. Nat. Anim. sans. Vertebr. V. 272.
Xantho impressus, Milne Wdwards, Hist. Nat. Crust. I. 393: A. Milne Edwards,
Nouv. Archiv. du Mus. IX. 1873, p. 198, pl. vi. fig. 2: F. Muller, Verh. Ges. Basel
VIII. 1886, p. 474: de Man, Journ. Linn. Soc., Zool, XXII. 1887-88, p. 30:
J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 359: Ortmann, Zool.
Jahrb., Syst. VII. 1893-94, pp. 444, 449.
Eudora impressa, De Haan, Faun. Japon. Crust., p. 28: A. Milne Edwards in
Maillard’s Vile Réunion, Annexe F. p. 4: Richters in Mébius Meeresf. Maurit.
p. 146, pl. xv. figs. 15, 16.
Carapace very short and broad, little convex in the anterior half,
quite flat in the posterior half.
Gastric and cardiac regions separated from the wings of the cara-
pace by very broad and deep furrows, the wings of the carapace being
thrown into massive smooth lobules by furrows not quite so deep, but
the gastric region being slightly and imperfectly areolate except quite
anteriorly.
Front somewhat declivous, bilobed, not laminar. The four lobes
of the antero-lateral margin are thickened and blunt, the first lobe
leing on a level lower than that of the orbit. Close in front of the
thickened posterior border is a smooth transverse wrinkle.
The tooth and notch at the antero-internal angle of the merus of
the external maxillipeds are very distinct.
Chelipeds equal in both sexes: outer surface of arm with a groove
following the contour of the distal border; outer surface of wrist with
a faintish Y-shaped dimple, and a strong double-crowned tubercle at
the inner angle of the wrist ; upper surface of hand with an incomplete,
longitudinal, pitted furrow; fingers with cutting-edge coarsely crenu-
late.
Legs thick, subcylindrical, smooth ; both edges of the dactylopodites
covered with thick short fur. |
Abdomen of male 5-jointed, with the sutures between the 3rd—5th
joints persistent.
Colours in spirit: waxy white, fingers blackish brown. The whole
animal has a smooth waxy look.
In the Indian Museum are two specimens, from Mergui and the
Andamans (besides three from Mauritius. )
This is a curious form, connecting Xantho with several other genera.
116 A. Alcock — Carcinological Fauna of India. [No },
Sub-genus LopHoxantuus, A. Milne Edwards,
Lophowanthus, A. Milne Edwards, Miss. Sci. Mex. Crust. p. 256.
Difters from typical Xantho (Xantho floridus etc.) chiefly in having
at least the upper edge of the legs distinctly crested: the carapace,
moreover, is longer and narrower.
43. Xantho (Lophoxanthus) scaberrimus, Walker.
Xantho seaberrimus, A. O. Walker. Journ. Linn. Soc., Zool., XX. 1886-90,
pp. 109, 115, pl. vii. figs, 1-4.
Carapace about % as long as broad, moderately convex im the
anterior two-thirds ; regions and subregions strongly defined by broad
deep smooth channels, convex, and covered with smooth well-defined
tubercles that are pea-like in the posterior third, somewhat scale-like
in the anterior two-thirds, and pointed along the antero-lateral border.
Fronto-orbital border less than half the width of the carapace :
front two-lobed, the lobes having an oblique and slightly concave
margin and a well-defined external angle.
Antero-lateral border four-lobed, the first lobe blunt and non-
prominent, the other three prominent and acuminate, all four with the
edges serrulate. Postero-lateral margin not concave, granular ; posterior
margin beaded.
The whole under surface of the carapace, and the surfaces of the
external maxillipeds, male sternum and male abdomen, are closely
covered with large granules,
Chelipeds equal, uniformly closely covered (except upper. surface
of arm and inner and outer surfaces of fingers) with sharpish tubercles,
which are largest on the hand, where they fall into raised longitudinal
parallel series, most marked on the lower part of the outer surface : fingers
fluted in continuation of the ridges on the hand, the ridges of the
dactylus being rough in their basal part. Two tubercles, the anterior
of which is the larger, at the inner angle of the wrist: and two some-
what foliaceous excrescences terminating the crest-like upper edge of
the arm.
First three pairs of legs with the edges of the merus (but especi-
ally the upper edge) sharply crested, the upper edge of the carpus and
propodite strongly serrated, and the dorsal surface of the carpus and
propodite furnished with squamiform granules in series parallel with
this serrated crest. The last pair of legs resembles the others, except
that the crest of the merus is serrated, and the dorsal surface of the
merus is granular.
Colours in spirit uniform ashy white.
1898. | A. Alcock — Carcinological Fauna of India: 117
In the Indian Museum besides a specimen from Japan is one from
off the Orissa coast, 11 fms.
Xantho (Lophoawanthus) scaberrimus var. baccalipes.
Differs from the type in the following particulars :—
(1) the characteristic tubercles have everywhere a worn appear-
ance, especially in the middle of the carapace and on the chelipeds and
the ischium of the external maxillipeds:
(2) the dorsal crest of the arm and of the meropodites of the legs
have each become a row of berry-like teeth, and the serrated crest and
granular ridges of the | carpopodites and propodites of the legs have
become merely low rough elevations,
In the Indian Museum are three large males from Ceylon: the
largest has the carapace 47 millim. long and 6] millim. broad.
Xantho (Lophoranthus) scaberrimus var. cultripes.
Differs from the type in the following particulars :—
(1) the characteristic tubercles are still more “ worn,’’ especially
on the mesogastrium, and near the inner angle of the wrist, and near the
base of the thumb, where they are almost worn away :
(2) the dorsal crest of the arm (with its foliaceous terminal
lobes), and the crests of the merus carpus and propodite in all the legs,
are greatly developed sharp and entire, and tlie raised rows of granules
ou the dorsal surfaces of the leg joints have almost disappeared.
In the Indian Museum is a single male from Singapore, with a
carapace 50 millim. long and 64 broad.
Sub-genus Lepropius, A. Milne Edwards.
Leptodius, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX. 1863, p. 284: Nouv.
Archiv. du Mus. IX. 1873, p. 221: Miss. Sci. Mex., Crust. p. 267, wba synon.
Leptodius, Miers, Challenger Brachyura, p. 136.
Leptodius (e.g. Leptodius exaratus) resembles Xantho (e.g. Xantho
floridunus) in general form and proportions, but differs most conspicu-
ously in having the fingers hollowed out “‘en cuillére” at tip. But this
divergence is almost bridged by Leptodius crassimanus, in which the
spooning of the fingers is indistinct.
Leptodius further differs from the type of Xantho (1) in the
greater convergence of the postero-lateral borders, (2) in the often—
but not always—more than four-lobed antero-lateral border, and (3) in
the often more extensive contact of the basal antennal joint with the
front.
118 A. Alcock — Carcinological Fauna of India. [ No. 1,
Key to the Indian species of the sub-genus Leptodius.
I. Carpus (and sometimes the propodite also) of the four
last pairs of legs strongly bicarinate dorsally —the
crests enclosing a trough-like cavity .........ccecseceerreee DL. cavipes.
II. Carpusand propodite of last four pairs of legs te —
i. Four teeth (exclusive of the orbital angle) on
the antero-lateral border: postero-lateral border
not or hardly shorter than the chord of the
antero-lateral border ...5..4 dees LV ddesscseess cee) MUP eRRUIEEE
ii. More than four teeth on the antero-lateral border:
postero-lateral border distinctly shorter than the
chord of the antero-lateral border :—
1, Front bilaminar, the lobes having a slight-
ly concave edge: 5 teeth on the antero-
lateral margin 4.2 ...000¢s eveoeeee DL. sanguineus.
2. Front bilaminar, the labiis sO eilebiole con-
cave as to make the front almost quadri-
dentate :—
a. “Five teeth on the antero-lateral
margin: finger-tips often rather in-
distinctly hollowed out :—
a. Carapace cut up into numer-
ous strongly convex lobules:
upper surface of wrist and
hand strongly and sharply
rugose and nodular,,,............ L. euglyptus.
B. Lobules of carapace not very
numerous, not very convex,
smooth: upper surface of wrist
and hand somewhat rough ... JL. crassimanus.
b. Eight to ten irregular teeth on the
antero-lateral margin .......see00...008 DL. nudipes.
44. Xantho (Leptodius) exaratus (Hdw.) A. M. Edw.
Chlorodius eraratus, Milne Edwards, Hist. Nat. Crust. I. 402; and in Cuvier
Réune An. Crust. pl. xi. fig. 3: Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 79; and
U. S. Expl. Exp. Crust. pt. I. p. 208: Stimpson, Proc. Ac. Nat. Sci. Philrd. 1858,
p. 34.
Leptodius ewaratus, A. Milne Edwards, Nouv. Archiv. du Mus. IV. 1868, p. 71;
and IX. 1873, p. 222: Kossmann, Reise roth. Meer. Crust. p. 32, pl. i. fig. 1-6:
Hilgendorf MB. Ak. Berl. 1878, p. 790: Richters in Mobius Meeresf. Maurit. p. 148:
Haswell, Cat. Austral. Crust. p. 60: Miers, Zool. H. M.S. Alert, pp. 183 and 214:
de Man, Archiv. f. Naturges. LII1. 1887, i. p. 285, and Journ. Linn. Soc., Zool.,
XXII. 1887-88, p. 33; and in |Weber’s Zool. Ergebn. Niederl. Ost. Ind. II. 1892
p. 278, and Zool. Jahrb., Syst., VIII. 1894-95, p. 521: Gano, Boll. Soc. Nat. Napoli,
III. 1889, p. 202: J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 362;
Whitelegge, Mem. Austral. Mus. 111. 1897, p. 137.
1898.] A. Alcock — Carcinological Fauna of India. 119
Xantho afinis, De Haan, Faun. Japon. Crust. p, 48, pl. xiii. fig. 8: Krauss,
Sudafr. Crust. p. 30.
Xantho lividus, De Haan, 0.c,, l.c. fig. 6: Miers, Zool. H. M. 8. Alert, pp. 183,
214.
Cancer inequalis, Audonin and Savigny Descr. Egypte pl. v. fig. 7 (fide A. M. E.)
Xantho ezaratus var. typica, Ortmann, Zool. Jahrb., Syst., VII. 1893-94, p. 445:
and in Semon’s Forschungsr. (Jen. Denk. VIII) Crust. p. 50,
Carapace moderately broad, moderately convex in the anterior two-
thirds, nearly flat in the posterior third, where also it is not areolated.
Gastric region well-defined, convex, fairly well areolated anteriorly, the
wings of the carapace on either side of it divided into about 5 low
lobules, three of which follow the contour of the antero-lateral border.
The surface of the carapace is non-granular, except sometimes in the
young.
Front not very prominent, but projecting beyond the inner angle
of the orbit, from which it is separated by a notch; bilaminar, the lobes
cut square, but with a slightly concave margin.
Antero-lateral border cut into 4 acuminate teeth, not including the
outer angle of the orbit, or a small denticle below it. Postero-lateral
border equal in length to the chord of the antero-lateral border.
Side wall of carapace, edges of upper surface of arm, and edges of
legs—but especially the upper edge of the meropodites— with a good
deal of hair.
Chelipeds unequal in both sexes. Upper and outer surface of wrist
more or less dimpled or wrinkled; a strong tubercle at inner an gle of
wrist. Hands usually smooth, but the upper surface has, very com-
monly, some low fine transverse or reticulating wrinkles. Fingers large,
thick, more or less fluted, not strongly toothed, meeting at tip only (in
the adult) where they are broadened and hollowed out.
Legs with merus subcylindrical and smooth, carpus and propus
nearly smooth and sometimes very faintly grooved, dactylus granular
and furred along both edges as far as the claw.
Abdomen of male five-jointed.
Colours in spirit: dirty yellow or dirty green, sometimes mottled;
fingers black.
In the Indian Museum are more than 130 specimens, chiefly from
the Andamans, Mergui, Karachi, also from the Persian Gulf, Bombay,
Ceylon, Akyab and Penang.
45, Xantho (Leptodius) sanguineus (Hdw.) A. M. Edw.
Chlorodius sanguineus, Milne Edwards, Hist. Nat. Crust. I. 402: Dana Proc. Ac.
Nat. Sci. Philad. 1852, p. 79, and U. 8. Expl. Exp. Crust. pt. I. p. 207, pl. xi. figs.
lla—d: Heller, Novara Crust. p. 18: Strects, Bull. U. S. Nat. Mus. VII. 1877.
p- 108.
120 A. Aleock — Carcinological Fauna of India. [No. 1;
Leptodius sanguineus, A. Milne Edwards, Nouv. Archiv. du Mus, IV. 1868, p. 7],
and IX. 1873, p. 224: Richters in Mobius, Meeresf. Maurit. p. 147: Haswell, Cat.
Austral. Crust. p. 60: F. Muller, Verh. Ges. Basel, VIII. 1886, p. 474: de Man,
Zool. Jahvb., Syst. VIII. 1894-95, p. 521: Whitelegge, Mem. Anstral, Mus, III.
1897, p. 137.
Leptodius exaratus, var. sanguineus, Miers, P. Z. S. 1877, p. 184; Ann. Mag. Nat.
Hist. (5) V. 1880, p. 234; Challenger Brachyura, p. 138: Cano, Boll. Soc. Nat
Napoli, TIT. 1889, p. 203.
Lagostoma nodosa, Randall, Journ. Ac. Nat. Sci. Philad. 1839, p. 111.
Chlorodius nodosus, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 79, and U. S.
Expl. Exp. Crust. pt. I. p. 210, pl. xi. figs. l4a—g.
Chlorodius edwardsi, Heller, Abh. zool.-bot. Ges. Wien, 1861, p. 10, and SB. Ak.
Wien XLIII. 1861, i. p. 336: Hilgendorf in v. d. Decken’s Reis. Ost. Afr. III, i.
p. 74.
Xantho exaratus var. sanguinea, Ortmann, Zool. Jahrb. Syst. VII. 1893-94,
p. 447.
Differs from Xantho (Leptodius) exaratus in the following particu-
lars :—
(1) the carapace is more convex anteriorly, and the branchial
lobules also are more convex :
(2) there are five teeth on the antero-lateral margin, not includ-
ing the external orbital angle and a small denticle below it:
(3) the postero-lateral border is a good deal shorter than the
chord of the antero-lateral border :
(4) the front is distinctly narrower.
In the Indian Museum are 123 specimens chiefly from the Andamans
and Laccadives, also from the Nicobars, Ceylon, and Persian Gulf.
46. Xantho (Leptodius) crassimanus, A. M. Edw.
Xantho crassimanus, A. Milne Edwards, Ann. Soc. Ent. France (4) VII. 1867
p. 267.
Leptodius crassimanus, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873,
p- 226, pl. xi. fig. 4: Haswell. Cat. Austral. Crust. p. 61: Muller, Verh. Ges. Basel,
VIII. 1886, p. 474: de Man. Archiv. fiir Naturges. LIII. 1887, i. p. 287, and Notes
Leyden Mus. XV. 1893, p. 284, and Zool. Jahrb., Syst. VIII. 1894-95, p. 522.
Xantho exaratus var. crassimana, Ortmann, Zool. Jahrb. Syst. VIL. 1893-94,
p. 448.
Differs from both exaratus and sanguineus in the following particu-
lars :—
(1) the two lobes of the front have the free edge not merely
emarginate, but deeply concave, so that the front appears to be formed
of four little teeth :
(2) the carapace, anteriorly, is much more convex, the regions
are more convex and their areole are more convex :
(3) the fingers are not so broad at tip and not so sharply hollowett .
out :
1898. ] A. Alcock — Carcinological Fauna of India. 121
(4) the upper surface of the wrist and hand is more rugose, It
resembles sanguineus in having 5 teeth on the antero-lateral margin, but
differs from it further in having
(5) the front even narrower, it being less than one-fifth the
breadth of the carapace.
It can at once be distinguished by the very narrow quadridentate
front.
In the Indian Museum are 22 specimens, from the Andamans,
Karachi, Galle (and Australia).
47. Xantho (Leptodius) nudipes (Dana), A. M. Edw.
Chlorodius nudipes, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 79, and U. S.
Expl. Exp. Crust. pt. I. p. 209, pl. xi. figs. 12a-c.
Leptodius nudipes, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873, p, 225:
Miers, Cat. Crust. New Zealand, p. 17: Filhol, Crust. New Zealand, p. 374: de Man,
Journ. Linn. Soc. Zool., XXII. 1887-88, p. 33, and Zool. Jahrb. Syst. 1894-95, p. 523.
Xantho exaratus var. nudipes, Ortmann, Zool. Jahrb. Syst. VII. 1893-94, p. 447.
The whole surface of the carapace is very finely pitted or granular.
The antero-lateral border is divided into four acute lobes or teeth, but
each of the first three teeth have, at base, either one or two (one on
either side) small additional cusps, and the fourth tooth is generally
double, so that altogether there are from 8 to 11 teeth on the antero-
lateral margin. The fingers are broad and deeply hollowed at tip. The
upper surface of hand and wrist is granular and rugose. The lobes of
the front are deeply concave.
In the convexity of the carapace and of its regions and subregions
it resembles sanguineus ; but the front is much broader than in sanguineus,
being more than one-fourth the breadth of the carapace, and the fingers
are typical spoons.
In the Indian Museum are 17 specimens from the Andamans and
3 from Mergui.
48. Xantho (Leptodius) euglyptus, n. sp.
Form of carapace much resembling that of sanguineus, but much
more convex.
Carapace 3 as long as broad, rather strongly convex in its anterior
two-thirds, flat posteriorly: its regions well delimited, convex, and as
completely areolated as any Actz#a—the areole being strongly convex
and somewhat pitted transversely.
Front projecting beyond the orbit, from which it is separated by a
notch, cut into two lobes of which the outer angle is prominent much as
in crassimanus ; its breadth is not quite a third that of the carapace.
J u. 16
122 A. Alcock — Carcinological Fauna of India. [No. 1,
Antero-lateral border cut into five conical teeth between which are
v-anules or little denticles ; postero-lateral borders strongly convergent,
shorter than chords of antero-lateral borders.
Chelipeds unequal: upper and outer surfaces of wrist strongly
wrinkled and pitted ; upper surface of hand nodular, upper half or more
of outer surface of hand longitudinally ridged and transversely wrin-
kled: fingers short, stout, hollowed (but not broadened) at tip.
Legs with carpus and propodite longitudinally ridged and grooved
above —the carpus more distinctly so—and dactylus furred. .
Sidewall of carapace, edges of upper surface of arm, and edges of
legs — but especially upper edge of meropodites — hairy.
Colours in spirit: yellow, fingers and front lower corner of hand
blackish brown.
Length of carapace 10°5 millim., breadth 16 millim.
In the Indian Museum are 45 specimens from Galle and 1 from
Mergui (Marine Survey).
This species, though strongly resembling mudipes and crassimanus,
is at once recognized by the sharp-cut Actzea-like sculpture of the cara-
pace. It is possible that it may be the Chlorodius eudorus of Milne
Edwards. It has the closest possible resemblance to the Xantho quin-
quedentatus of Krauss, Sudafr. Crust. p. 30, pl. i. fig. 3, but that species
is described and figured as having sharp fingers.
49. Xantho (Leptodius) cavipes (Dana).
Chlorodius cavipes, Dana, Proc. Ac. Nat. Sci. Phila., 1852, p. 79; and U.S.
Expl. Exp. Crust. pt. I. p. 212, pl. xii. figs. la-b: Stimpson, Proc. Ac. Nat. Sci.
Phila., 1858, p. 34. |
Leptodius cavipes, de Man, Journ. Linn. Soc., Zool., XXII. 1887-88, p. 34.
Carapace convex in anterior two-thirds, flat behind. Gastric region
convex, well delimited and areolated by fine smooth rather deep
channels ; wings of carapace divided into about five lobules by less deep-
cut and less smooth channels: the whole carapace (except the channels)
covered with miliary granules, which on the lateral lobes of the gastric
region are arranged in lines that have an imbricate look.
Front bilaminar, the fore edge of the lobes thickened and granular.
Antero-lateral border thickened and granular, cut into small irregular
teeth — 8 or 9 in number — which fall into 4 sets. Undersurface of
carapace with short fur.
Chelipeds unequal: upper and outer surface of wrist»and hand
wrinkled and granular, outer surface of hand covered with granules in
more or less distinct lines : fingers little toothed, incurved, blunt-pointed
and hollowed (but not broadened) at tip.
1898. ] A. Alcock — Carcinological Fauna of India. Vee
Legs rough, but not very hairy: upper edge of meropodites finely
serrated, distally sharply notched: wpper surface of carpopodites wilh two
high longitudinal crests enclosing a trough-like space; the propodites are
similarly sculptured, but the sculpture is a good deal concealed by fur :
dactyli furred.
Colours in spirit: dirty yellow or dirty greenish, fingers nearly
black in distal ? only.
In the Indian Museum are 4 specimens, from the Andamans, Mergui
and Ceylon.
Mepaus, Dana.
Medzus, Dana, Silliman’s Amer. Journ. Sci. and Arts, (2) XII. 1851, p. 1253
Proc. Ac. Nat. Sci. Philad. 1852, p. 76; U. S. Expl. Exp. Crust. pt. I. p. 181.
Medzus, A. Milne Edwards, Ann. Sci. Nat. Zool. (4) XX. 1863, p. 279; Miss.
Sci. Mex. Crust. p. 249.
Medzus, Miers, Challenger Brachyura, p. 116.
Carapace not very broad, hexagonal, little convex, the regions well
defined and well areolated.
Antero-lateral borders cut into teeth and very distinctly continued
beneath the orbits to the angles of the buccal cavern.
Fronto-orbital border half, or a little more than half, the greatest
breadth of the carapace.
Front about a fourth, or a little more, the greatest breadth of the
carapace, horizontal, rather prominent, square-cut, notched in the middle
line, separated from the supra-orbital margin by a notch.
Orbits, eyes, basal antennal joint and antennary flagellum as in
Xantho.
Chelipeds either unequal or subequal, the wrists and hands com-
monly covered with large nodules, the fingers pointed.
The abdomen of the male consists of five pieces, the 3rd—5th somites
being fused.
Medzus closely resembles Xantho, but is distinguished by the
narrower carapace and by the relations of the antero-lateral border. In
some species of Xantho (e.g., X. distinguendus) the antero-lateral border
is broken and eroded near the orbit, so that it may be imagined to be
continued to the angle of the buccal cavern, but in Medzus there is no
ambiguity whatever.
50. Medeus nodosus, A. M. Edw.
Medzus nodosus, A. Milne Edwards, Ann. Soc. Ent. France, (4) VII. 1867,
p- 271; Nouv. Archiv. du Mus. IX. 1873, p. 212, pl. viii. fig. 2: Haswell, Cat.
Austral, Crust. p. 52.
x
Carapace shaped much as in Polycremnus, hexagonal, more than
124 A. Alcock— Careinological Fauna of India. [ No. I,
as long as broad, the regions well demarcated, well areolated, finely and
closely granular.
Front horizontal, square-cut, prominent, sublaminar, notched and
grooved in the middle line.
Antero-lateral borders cut into four blunt-pointed rather coarse
granular teeth, of which the two posterior are the strongest.
Chelipeds subequal: wrist and hand covered with granular fungi-
form tubercles, which are arranged in regular longitudinal series on the
hand.
Upper border of meropodites of legs without spinules.
In the Indian Museum is a single small male from off the Ganjam
coast, 73-93 fms.
Henderson (Trans. Linn. Soc. (2) V. 1893, p. 860) appears to consi-
der this species to belong to the genus Halimede, but it has not —if my
identification be correct—the curious male abdomen which distinguishes
that genus from every other Xanthoid except Polycremnus,
Cyctoxantuus, A. Milne Edwards.
Cycloranthus, A. Milne Edwards, Ann. Sci. Nat., Zool. (4) XX. 1863, p. 278;
Nouv. Archiv. du Mus. IX. 1873, p. 209; Miss. Sci. Mex., Crust. p. 258.
Carapace relatively long: front horizontal, prominent, and divided
by a median fissure into two lamellar lobes, and separated from the »
internal orbital angles by a deepish notch.
Orbits small: two fissures in the supra-orbital margin: external
orbital angles inconspicuous, continuous with the antero-lateral borders.
Antero-lateral borders very long, strongly curved, extending far
backwards.
Basal antennal joint short, but touching the front at its inner
angle: the flagellum inserted in the orbital hiatus.
Merus of the external maxillipeds subquadrilateral.
The abdomen of the male consists of five movable pieces.
This genus is not represented in the Indian Museum,
51. Cyeloranthus lineatus, A. Milne Edwards.
Cyclowanthus lineatus, A. Milne Edwards, Ann. Soc. Entom. France, (4) VIF.
1867, p. 269, and Nouv. Archiv. du Mus. IX. 1873, p. 209, pl. vi. fig. 5: Miers, Zool.
H. M.S. Alert, pp. 183, 212: J. R. Henderson, Trans. Linn. Soe., Zool., (2) V. 1893,.
p. 360. _
Carapace broad, very depressed, smooth; the regions little defined ;
antero-lateral borders prolonged far backwards and obscurely divided
into four deutiform lobes.
1898.] - A. Alcock —Carcinological Fauna of India. 125
Outer orbital angle inconspicuous, orbits small, the upper margin
with a narrow cleft.
Front very prominent, lamellar, a little sinuous at the sides, pro-
minent towards the middle where there is a linear fissure.
Chelipeds unequal: the hand has the inner surface flattened, the
upper border obtusely crested, and the outer surface rugose: the wrist,
which has its outer border much dilated, is equally rugose.
The legs are weak and smcoth, the dactylus being a little com-
pressed.
Carapace yellowish, very symmetrically marked with numerous
reddish-brown lines, some of which run obliquely from the antero-
lateral borders towards the front and towards the middle line, while
others run from the posterior border forwards to the cardiac region.
There are no specimens in the Indian Museum collection.
HoPLoxaNTHUs, 0. gen.
Carapace hexagonal, moderately broad, moderately convex fore and
aft, its regions all well defined, and to a certain extent subdivided.
The antero-lateral borders are thin and erest-like and are ona
much lower plane than the rest of the carapace, they end either in both
sexes or in the female only, in a large horizontal lateral epibranchial spine,
and may either be cut into large triangular teeth or may be only
obscurely notched.
Postero-lateral borders straight, moderately convergent, about as
long as the antero-lateral.
Front lamellar, prominent, horizontal, notched in the middle line,
about a fourth to two-sevenths the greatest breadth of the carapace.
Orbital margin with two faintish notches or suture lines above, and
with a small triangular gap just below the outer angle: the inner angle
of the lower orbital margin forms a strongly-projecting tooth: the outer
-orbital angle is confluent with the antero-lateral margin.
The antennules fold obliquely. The basal antennal joint is rather
slender and meets the front: the flagellum, which is about as long as
the major diameter of the orbit, is lodged in the orbital hiatus.
The anterior edge of the external maxillipeds is almost transverse.
Chelipeds unequal, fingers sharp pointed : legs rather slender.
No ridges, defining the efferent branchial channels, on the plate.
The abdomen of the male consists of seven separate segments, the
last segment being no longer than the longest of the others.
This genus is closely allied to Xantho and Cyclozanthus, it is also
related not distantly to Halimede and to Lophozozymus.
126 A. Alcock — Oarcinological Fauna of India. [No. l,
52. Hoploxanthus heatit, n. sp.
The whole of the carapace, legs, and outer surface of chelipeds is
covered with a dense, darkish, extremely short, velvety or branny
pubescence.
Carapace hexagonal, moderately broad, moderately convex, the
regions well defined, tumid, their convexities granular.
Front prominent beyond the orbits and separated from them by a
notch, square-cut, bilaminar.
Antero-lateral border thin sharp, cut into four triangular laciniate
teeth, the last of which—in the female but not in the male—is an
acute salient spine.
The edges of all the teeth, of the front, and of the orbit are finely
granular.
The postero-lateral border is elegantly granular and quite straight:
dorsal to it the wall of the carapace forms a distinct postero- -lateral
facet, sharply marked off from the general surface of the carapace.
Chelipeds a little unequal in both sexes: wrist with a small some-
what cristiform expansion at the outer angle and a tooth at the inner
angle: upper surface of hand with two or three longitudinal raised
sculptured lines, the innermost of which consists of a blunt cristiform
lobule followed by one or two blunt denticles, the outer one or two
being simply crenulate and granular; the outer surface of the smaller
hand is everywhere granular, that of the larger hand is granular in
part—in both cases some of the granules form slightly-raised longi-
tudinal lines.
Legs long, slender.
Colours in spirit yellowish brown.
Carapace of male 10 millim. long, 13 millim. broad; of female,
11] millim. long, 17 millim. broad.
In the Indian Museum are 3 specimens from the east coast of India
and 2 from the Nicobars.
53. Hoploxanthus cultripes, n. Sp.
Carapace hexagonal : the three gastric subregions (lateral and post-
medial), the cardiac region, and two (smaller) median epibranchial regions
stand out as very prominent granular bosses, and the convexity of the
lateral epibranchial spine, and the postero-lateral border» and_ its
neighbourhood are granular, —otherwise the carapace is quite smooth.
Front prominent, sublaminar, with a curved convex finely granular
edge, faintly notched ir the middle line and hardly separated from the
supra-orbital angles.
1898. ] A. Aleock — Carcinological Fauna of India. 127
Antero-lateral borders very thin and sharp, obscurely divided by
faint notches and fainter grooves into 3 broad inconspicuous lobes, and
ending in a strong horizontal pyramidal lateral epibranchial spine.
The chelipeds in the unique specimen are lost; but the legs are
remarkable in having the upper edge of the merus and carpus sharply
carinate, the carpal joints having a second blunter and lower keel along
the dorsal surface.
The legs and the undersurface of the body are covered with the
same dense extremely short pubescence as occurs in H. heextit.
The single imperfect male in the Indian Museum comes from
Karachi, and is 9 millim. long and 13 millim. broad,
ORPHNOXANTHUS, n. yen.
Carapace, owing to the inflation of the branchial regions almost
quadrilateral in outline and almost concave from side to side, but very
decidedly convex fore and aft, broad, the regions well defined but not
to any great extent areolated.
Fronto-orbital border a little more than half the greatest breadth
of the carapace in extent. Front about a third the greatest width of
the carapace, lamellar, projecting horizontally beyond the orbits, broadly
and faintly bilobed. Orbital margin entire: orbits and eyes small.
Antero-lateral border cut into four teeth; postero-lateral borders
convergent only in the posterior half; posterior border long.
The antennules fold almost transversely. The basal antennal joint
is very short and only just touches the turned down edge of the front;
the flagellum which is very long (between 2 and 3 times the length of
the orbit) is lodged in the narrow orbital hiatus.
Owing to the bulge of the outer wall of the efferent branchial canal
and the consequent puffing out of the pterygostomian regions, the front
edge of the merus of the external maxillipeds is quite transverse or
even slightly oblique from without inwards. _ |
The chelipeds are massive and unequal; the fingers are compressed
aud pointed, The legs are very slender... |
The abdomen of the male consists of 5 segments, the 3rd-5th
somites being fused.
Owing to the inflation of the pterygostomian regions the efferent
branchial channels are permanently open, but the low crests that define
them are confined to the posterior part of the endostome.
This genus appears to represent one of the links between Galene and
Xantho. ‘The single known species comes from the Bay of Bengal, 105-
350 fms.
128 A. Alcock — Carcinological Fauna of India. [ No. 1,
54. Orphnowanthus microps, Alcock and Anderson,
Xanthodes microps, Alcock and Anderson, J. A. 8. B. LXIII. pt. 2, 1894, p, 183.
Carapace about § as long as broad, almost quadrilateral in outline,
strongly convex fore and aft, but, owing to the inflation of the branchial
regions, a little concave from side to side; it is rather closely covered
with a very fine short fur, beneath which the surface may be granular
or nearly smooth, but the margins are always granular. The regions
are all well defined and are slightly tumid: the gastric region is divided |
into 3 gently tumid subregions, the branchio-hepatic regions are sub-
divided transversely into three areas, and the fronto-orbital margin is
also marked off.
The antero-lateral border is thin and sharp and is cut into four
sharp finely granular teeth, the first of which runs by a long nearly
transverse margin, into the (undefined) angle of the orbit. The front is
laminar and projects beyond the supra-orbital margin; it is square-cut
and is slightly notched in the middle line, so as to form two broad shallow
lobes. The eyes are small and are to a variable extent deficient in
pigment.
The chelipeds are unequal—very much more so in the male than in
the female: the arm to a variable extent, the entire surface of the
wrist, and the upper border of the hand are scabrous and more or less
hairy; the other surfaces of the hand may be smooth and polished,
or the outer surface may be to a variable extent granular: the fingers
are large, compressed and pointed.
In the male the larger cheliped is about 25 times the length of the
carapace (the hand and fingers forming slightly more than half the
length) and nearly half the arm projects beyond the carapace in repose.
The legs are long slender and finely and sparsely hairy: the upper
edge of the meropodites is scabrous or closely spinulate.
Colours in spirit; chestnut brown with blackish fingers. Length of
carapace (average) 11 millim., breadth 15 to 16 millim.
In the Indian Museum are 29 specimens from the Bay of Bengal,
105-350 fms. ) ,
Etisus, Milne Edwards.
Etisus, Milne Edwards, Hist. Nat. Crust. I. 410.
Etisus, Dana, Silliman’s Amer. Journ. Sci. and Art. (2) XII. 1851, p. 126; and
U.S. Expl. Exp. Crust. pt. I. p. 183.
Etisus, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX. 1863, p. 291; and
Nouv. Archiv. du Mus. IX. 1873, p. 233.
Etisus, Miers, Challenger Brachyura, p. 131.
Carapace broad, moderately convex in both directions, with the
-1898.] way: Alcock — Oarcinological Fauna of India. 129
regions delimited by broad shallow rather indistinct depressions and
having a slightly uneven but not definitely lobulated surface. |
The front is laminar and narrow ; it projects well beyond the supra-
orbital border from which it is separated by a deep notch, and is split
by a suture in the middle line. The orbital margin is broken by three
sutures or actual fissures, and the tooth at the inner angle of the lower
border is very prominent.
The antero-lateral borders, which are a good deal longer than the
postero-lateral, are cut into from 4 to 8 lobes or procurved spines.
) The basal antennal joint has its outer angle produced and tightly
wedged into the internal orbital gap, which it fills; but the flagellum,
which is of good length, does not arise within the orbit but at the base
of this process.
) The outer border of the merus of the external maxillipeds is
oblique.
The chelipeds, which are very massive and rather long, are a little
unequal in the male: the fingers are very stout and strongly arched,
and they meet only at the tip, which is broad expanded and hollowed
out almost like a horse’s hoof.
The abdomen of the male is five-jointed, the 8rd—-5th somites being
fused.
Key to the Indian species of Ktisus.
I. More than four teeth on the antero-lateral border, exclud-
ing the external angle of the orbit: free edge of front
not convex: the process of the basal antennal joint com-
pletely separates the lower from the upper inner angle of
the orbit : legs spiny :—
i, Seven or eight uneven unequal-sized claw-like teeth
on the antero-lateral border ............ sesecssoecse H, dentatus,
ii. Seven evenly arranged broad Meal. homered
teeth of almost uniform size on the antero-lateral
PAROS ic. 0ese DU PR aS oe ale w. HH. utilis.
II. Four teeth (excluding the external Hei of the orbit) | on
the antero-lateral border : free edge of front bow-shaped:
the tooth at the inner canthus of the orbit in contact
with the eave of the orbit beyond the tip of the process
of the basal antennal joint: legs not spiny ....s.s.cccseeeees E, levimanus.
55. Htisus dentatus, (Herbst) Edw.
Cancer dentatus, Herbst, Krabben, I. ii. 186, pl. xi. fig. 66.
Etisus dentatus, Milne Edwards, Hist. Nat. Crust. I. 411: Dana, U. 8S. Expl.
Exp. Crust. pt. I. 185, pl. x. figs. 2a-b : A. Milne Edwards, Nouv. Archiv. du Mus.
IX. 1873, p. 233: Miers, P. Z. 8. 1877, p. 1384: Richters in Mobius, Meeresf. Maurit.
we tl. 17
130 A. Alcock —Carcinological Fauna of India. [No. 1,
p. 146: Lenz and Richters, Abh. Senck. Ges. XII. 1881, p. 421: Haswell, Cat.
Austral, Crust. p. 53.
Etisodes dentatus, Ortmann, Zool. Jahrb. Syst. VII. 1893-94, p. 472.
Carapace smooth (non-granular): gastric region well defined except
at cardiac end, its surface broken, but not definitely lobulated ; similarly
with the branchio-hepatic regions.
The antero-lateral border bears 7 or 8 (exclusive of the external
orbital angle) pro-curved claw-like teeth, uneven both as to size and
place, though four of them—which correspond to the 4 lobes of so
many other Cancroids—are much of one size and much larger than the
other 3 or 4.
The front is lamellar with the free edge slightly and angularly
emarginate, and is cleft in the middle line by a fine sharp groove that
extends well on to the gastric region. The orbital margin has three
teeth, separated by fissures, in its outer part. The tooth at the inner
angle of the Jower edge of the orbit is sharp, and does not come into
contact with the eave of the orbit.
Chelipeds in the adult male a little more, in the adult female a
little less than twice the length of the carapace: the arm has a few
spinules and a good deal of hair along the upper border, and some
granules or blunt spinules along its lower and its distal borders; the
wrist has a strong spike at its inner angle; the hand may or may not
have a few pimple-like granules on its upper outer surface; and the
fingers are fluted, the ridges on the dactylus being crenulate or dentate.
Otherwise the chelipeds are smooth.
In the legs, the merus has both the dorsal and the ventral edges
thickly fringed with long stiff hairs, the dorsal edge being also granular ;
the carpus has at least three series of spinules along its dorsal surface;
the propodite has a rather granular surface with about four (dorsal)
series of spinules, and has much of its luwer edge fringed with long stiff
hairs; and the dactylus is spiny above and hairy below.
Colours of a specimen 7 years in spirit: upper surface bright
maroon fading to yellow near the posterior border ; fingers black,
In the Indian Museum are a young female from Port Blair (Anda-
mans), and three large males (carapace 72 millim. by 111 millim.) from
Great Coco I. (Andamans) and Hast I. Andamans.
56. Htisus utilis, Lucas.
Etisus utilis, Lucas in Jacquinot, Voy. Astrolabe, Crust. p. 27, pl. ii. fig. 6:
Heller, Novara Crust. p.16: A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873,
p. 233: E. Nauck, Zeits. Wiss. Zool. XXXIV. 1880, p. 58 (gastric teeth): Lenz and
Richters, Abh. Senck. Ges. XII. 1881, p. 421: Ortmann, Zool. Jahrb. Syst. VII,
1893-94, p. 472.
1898. ] A. Alcock — Carcinological Fauna of India. 131
Carapace smooth (non-granular), distantly pitted in the anterior
and lateral parts. Gastric region fairly well defined, except at the
cardiac end, its surface showing indistinct traces of lobulation : branchio-
hepatic regions with the surface a little uneven.
Antero-lateral border with 7 (exclusive of the external orbital
angle) regular, even, nearly uniform, compressed, procurved teeth.
Front as in Htisus dentatus, but with the angles a little sharper cut.
The edge of the orbit is trenchant, and near the outer angle are 3 not
very distinct suture-lines: the tooth at the inner angle of the lower
border of the orbit is prominent, and does not come into contact with
the eave of the orbit.
Chelipeds in the adult male twice to twice-and-a-half, in the adult
female once-and-a-half to once-and-two-thirds the length of the cara-
pace: upper and lower edges of arm coarsely and unevenly granular,
much of the upper edge also hairy ; distal end of wrist with 4 or 5 teeth,
the inner two of which are long and large; the hand has, along its
upper border, a double crest of strong teeth, continued in blunter form
along the finger, and on the outer surface of the hand there may be a
few pimple-like granules.
In the legs, both the upper and lower edges — but most the upper
edoe—of the merus, carpus and propodite are thickly fringed with long
bristles, as also is the lower edge of the dactylus; the upper edge of the
merus has also a row of small spines, and the upper edge of the carpus,
propus and dactylus a double row of increasingly larger spines: the
lower edge also of the propus and dactylus is spiny.
Colours in spirit: dull yellowish pink, fingers black.
In the Indian Museum are a male and a female from the Singapore
Museum, and supposed to have come from Singapore. (Heller l.c,
records this species from the Nicobars).
57. Btisus laevimanus, Randall,
Etisus laevimanus, Randall, Journ. Acad. Nat. Sci. Philad., 1839, p. 115: Dana.
Proc. Ac. Nat. Sci. Phila. 1852, p. 76, and U. S. Expl. Exp. Crust. pt. I. p. 185, pl. x.
figs. la-b: A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873, p. 234: Kossmann,
Reise roth Meer. Crust. p. 30: T. Tozzetti, Magenta Crust. p. 29: Streets, Bull,
U.S. Nat. Mus. VII. 1877, p. 105: Hilgendorf, MB. Ak. Berl. 1878, p. 791: Richters
in Mobius Meeresf. Manrit. p. 146: de Man, Notes Leyden Mus. III. 1881, p. 99;
and Archiv. fiir Naturges. LIII. 1887, i. p. 289; and Zool. Jahrb. Syst. VIII. 1894-95,
p. 527: Haswell, Cat. Austral. Crust. p.54: Miers, Zool. H. M. S. Alert, pp. 183,
217; and Challenger Brachyura, p. 132: F. Muller, Verh. Ges. Basel VIII. 1886,
p. 474: J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 362: Ortmann,
Zool. Jahrb. Syst. VII. 1893-94, p. 473: Whitelegge, Mem. Austral. Mus, III, 1897,
p. 131,
132 A. Alcock —Oarcinological Fauna of India. [No. 1,
Etisus macrodactylus, Lucas in Jacquinot’s Voy. Astrolabe, Crust. p. 30, pl. ix.
fig. 2, (A. M. E.)
Etisus convesus, Stimpson, Proc. Ac. Nat. Sci. Phila. 1858, p. 31.
Etisus maculatus, Heller, Abh. zool.-bot. Ges. Wien XI. 1861, p.9; and SB. Ak.
Wien, XLIII. 1861, p. 332: de Man, Notes Leyden Mus. II. 1880, p. 1738.
Gastric region well defined on all sides, its anterior part distinctly
lobulated ; branchio-hepatic regions with three lobules following the
curve of the antero-lateral margins.
Antero-lateral border with 4 broad teeth (exclusive of the external
orbital angle), the last two of which culminate in procurved points.
The front is cleft in the middle line by a groove; its free edge is
bow-shaped. ‘The orbital margin has, in its outer half, three lobular
constrictions defined by three grooves.
The tooth at the inner angle of the lower edge of the orbit is blunt,
and it comes into contact with the eave of the orbit beyond the tip of
the process of the basal joint of the antenna.
Chelipeds in the adult male about twice and a half, in the adult
female a little less than twice the length of the carapace ; the wrist has
a blunt spine at the inner angle, otherwise they are smooth and un-
sculptured.
Legs with both edges of all the long joints hairy, most so on the
lower edge of the dactylus and on the upper edge of the other joints :
the upper edge of the propodite and dactylus is also sharply granular,
but: there are no spines.
Colours in spirit variable: dull yellow, or. dull greenish-brown, or
sea-green, often with cinnamon coloured patches or small spots.
In the Indian Museum are 20 specimens, from Persian Gulf,
Karachi, Bombay, Laccadives, Andamans, and Singapore, (besides
specimens from Celebes and Mauritius).
Etisopss, Dana.
Etisodes, Dana, Silliman’s Amer. Journ. Sci. and Arts, (2) XII. 1851, p. 126
(footnote) ; Proc. Ac. Nat. Sci. Phila. 1852, p. 77; and U.S. Expl. Exp. Crust. pt. I.
p. 184.
Etisodes, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX. 1861, p. 291; and
Nouv. Archiv. du Mus. IX. 1878, p. 235.
The genus Hiisodes resembles Htisus in the characteristic form and
lie of the basal joint of the antennz, and in the characteristic relation
of the front to the orbits ; but it differs in the following particulars :—
The carapace is much longer and narrower; it is less convex, especially
in its posterior third; its regions are clearly defined, and are definitely
sculptured into lobules in the anterior two-thirds of the carapace: the
;
1898, ] A. Alcock — Carcinological Fauna of India. 133
chelipeds are much shorter, the difference in length being chiefly in the
arm; and the fingers though well hollowed out at tip are not so hoof-
like.
58. LHtisodes anaglyptus, (Edw.)
Cancer anaglyptus, Milne Edwards in Cuvier, Régne An. Crust. pl. xi. fig. 4,
Etisus anaglyptus, Milne Edwards, Hist. Nat. Crust. I. 411: Hess, Archiv. fiir
Naturges. XX XI. 1868, i. p..184: de Man, Notes Leyden Mus. XIII. 1891, p. 7.
_Etisodes anaglyptus, A. Milne Edwards, Nonv. Archiv. du Mus. IX. 1873, p. 235:
Haswell, Cat. Austral. Crust. p. 55: Miers, Zool. H. M.S. Alert, pp. 183, 218:
Ortmann, Zool. Jahrb. VII. 1893-94, p. 471.
Length of carapace nearly three-qnarters the breadth,
The regions are all convex and well defined, and the gastric and
branchio-hepatic regions are subdivided into convex lobules, the surface
of which is somewhat dented transversely.
The antero-lateral border is cut into four (excluding the external
angle of the orbit) procurved teeth, the last two of which are claw-like.
The front projects strongly, and is divided into two dorsally-convex
lobes, of which the free edge may either be cut obliquely inwards, or be
so excised as to give the front a four-pronged look.
The orbital margin has, in its outer half, three grooves separating
three blunt teeth: the tooth at the lower inner angle does not come in
contact with the eave of the orbit.
Chelipeds in the male not much more than half again as long ag
the carapace: upper and anterior borders of arm hairy ; upper surface
of wrist nodular, with two teeth (one large) at the inner angle; upper
outer surface of hand with rather irregular longitudinal series of little
nodules and granules ; dactylus fluted, the ridges being crenulated.
Legs very shaggy, the hairs almost concealing some lines of sharp
granules or spinules on the propodite and dactylus. _
In the Indian Museum is a specimen from the Persian Gulf (be-
sides one from Samoa).
59. LHtisodes electra (Herbst), Miers.
Cancer electra and ? metis, Herbst, Krabben, III. ii. 34 and 36, pl. i. fig. 6, and
pl. liv. fig. 3.
Etisus rugosus, Lucas in Jacquinot’s Voy. Astrolabe III. Crust. p. 33, pl. iv
fig. 2 (fide A. M. E., infra).
? Chlorodius dentifrons, Stimpson, Proc. Ac. Nat. Sci. Phila. 1858, p. 34,
Etisodes sculptilis, Heller, Abh. zool.-bot. Ges. Wien XI. 1861, p. 10, and SB. Ak.
Wien XLIII, 1861, p. 333: A. Milne Edwards, Nouv. Archiv. du Mus, IX. 1873,
p. 236, pl. ix. fig, 2: Kossmann Reise roth. Meer., Crust. p. 80,
134 A. Alcock — Carcinological Fauna of India. [No, 1,
Chlorodius samoensis, Miers, Ann. Mag. Nat. Hist. (4) XVI. 1875, p. 341 (Miers
infra.)
Etisodes electra, Miers, Zool. H. M. S. Alert, pp. 183, 217, 517, 532: de Man,
Archiv. fiir Naturges, LILI. 1887, i. p. 290: J. R. Henderson, ‘l'rans. Linu. Soc., Zool.,
(2)3 V. 1893, p. 362.
Closely resembles E. anaglyptus, from which it is distinguished by
the following characters :—
(1) the carapace is even longer and narrower, the length being
quite ? the breadth ;
(2) the whole shietdi of the lobules of the carapace and of the
nodules of the wrist aud hand is closely granular ;
(3) the front is cut into four teeth of nearly equal size ;
(4) the legs, though hairy, are not so shaggy.
In the Indian Museum are 4 specimens from the Andamans and
Nicobars, (besides three from Upolu and Mauritius).
Alliance V, Halimedoida.
Halimede. Polycremnus.
PoLycrRemMNnus, Gerstaecker.
Polycremnus, Gerstaecker, Archiv. fur Naturges. XXII. 1856, p. 120.
Carapace approaching the pentagonal, not very much broader than
long, distinctly convex fore-and-aft, slightly convex from side to side,
the regions rather indistinctly defined and to a certain extent subdivid-
ed by broad shallow depressions.
The antero-lateral border is elegantly four-lobed and is continued
beneath the orbits to the outer angle of the buccal cavern. The postero-
lateral borders are moderately convergent and are about equal in length
to the antero-lateral borders and also to the posterior border.
The fronto-orbital border is less than half the greatest width of the
carapace in extent. The front is narrow (less than a fourth the greatest
breadth of the carapace), sublaminar or hood-like, bilobed, and projects
well beyond the orbits. The three grooves in the vicinity of the outer
angle of the orbit are distinct. Eyes on short thick stalks. The inner
angle of the lower edge of the orbit is strongly produced.
The antennules fold obliquely. The basal antennal joint is long,
rather slender, and well in contact with the front: the flagellum is fous 5
(a good deal longer than the major diameter of the orbit) and is lodged
in the narrow orbital hiatus.
Anterior edge of merus of external maxillipeds almost transverse.
Chelipeds unequal in both sexes. Legs stout.
Abdomen of the male with all 7 joints distinct and separate: the
last segment unusually long and acute. :
1898. | A. Alcock —Carcinological Fauna of India. 135
60. Polycremnus ochtodes, (Herbst) Gerstaecker.
Cancer ochtodes, Herbst, Krabben, I. ii. 158, pl. viii. fig. 54: Fabricius, Ent.
Syst. II. 455, and Suppl. p. 337.
Galene ochtodes, Adams and White, Samarang Crust. p. 43, pl. x. fig. 2.
Polycremnus ochtodes, Gerstaecker, Archiv. fur Naturges. XXII. 1856, p. 121:
A. O. Walker, Journ. Linn. Soc., Zool., XX. 1886-90, p. 110: Henderson, Trans.
Linn. Soc., Zool., (2) V. 1893, p. 359.
Carapace oval-pentagonal, its surface smooth, a little lumpy owing
to the broad shallow depressions that somewhat indistinctly separate
and to a certain extent subdivide the regions,
All the borders are of about equal length: the antero-lateral is
divided into four rounded deep-cut lobes, decreasing in size from behind
forwards, and is continued beneath the slightly tumid lobe of the
orbital angle to the angle of the buccal cavern: on the postero-lateral
border just behind the junction with the antero-lateral are usually a
few granules.
The front projects horizontally forward beyond the orbits and
consists of two unguiform lobes separated in all their extent by a deep
narrow groove: it is a distinct rostrum.
The chelipeds are unequal, most markedly so in the male. The
upper border of the arm is elegantly cut into teeth or pisiform or pearl-
like tubercles: two similar tubercles stand, one below the other, at the
inner angle of the wrist, and the upper and outer surfaces of the wrist are
more or less covered with papule-like or pustulous tubercles: the upper
border of the hand, and of the basal half of the finger bears a row of
pisiform tubercles, and there are numerous pustulous tubercles on the
upper surface and on the proximal part of the outer surface of the
hand: fingers sharp pointed.
The legs are smooth, but the upper border of the meropodites of
all, or of the first three pairs, is distantly serrate or spinulous: the
dactylus and the neighbouring part of the lower border of the propodite
is furred.
Colours in spirit leaden grey, or yellowish with livid markings.
In the Indian Museum are 2 specimens from the Madras Coast and
one from Penang.
HatimepeE, De Haan.
Halimede, De Haan, Faun. Japon. Crust. p. 35: Dana, Amer. Journ. Sci. and
Arts, (2) XII. 1851, p. 125, and U. S. Expl. Exp. Crust. pt. I. p. 149.
Closely allied to Polycremnus, having the same form of male
abdomen.
The genus is not represented in the Indian Museum.
136 A. Alcock — Carcinological Fauna of India. [No. 1,
61. Hualimede (?) thurstonit, Henderson.
Halimede thurstoni, Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 360,
pl. xxxvi. figs. 13, 14.
It appears to me doubtful whether this is a true Halimede.
Alliance VI. Galenoida. [or Subfamily Gateninz].
GaLENt, De Haan.
Galene, De Haan, Faun. Japon. Crust. p. 19.
Galene, Miers, Challenger Brachyura, p. 118 (footnote.)
Car apace approaching the quadrilateral, strongly convex fore and
aft, little convex from side to side; its surface granular in parts, or
nearly smooth, and with the regions more or less distinctly shown by
broad shallow rather vague depressions.
Antero-lateral border moderately arched, ind istinctly four-lobed —
the last 2 or 3 lobes in the typical species being marked by spine-like
teeth : postero-lateral borders very slightly convergent, rather longer
than the chord of the antero-lateral: posterior border long.
Fronto-orbital border less than half the greatest width of the cara-
pace. Front obliquely deflexed, less than one-fifth the greatest width
of the carapace in extent, bilobed or quadridentate. Orbital margin
with the three grooves in the vicinity of the outer angle distinct: eyes
on thick stalks of moderate length. The antennules fold nearly trans-
versely.
Basal antennal joint broad, extremely short, not nearly reaching
the front; flagellum longish (longer than the major diameter of the
orbit) lodged in the broad orbital hiatus.
Anterior edge of merus of external maxillipeds a little oblique.
Chelipeds massive, unequal in both sexes, fingers pointed. Legs long,
stoutish. .
Abdomen of male with all 7 joints separate and distinct.
No crests, delimiting efferent branchial canals, on the endostome.
62, Galene bispinosa (Herbst) De Haan.
Cancer bispinosus, Herbst, Krabben, I. ii. 144, pl. vi. fig. 45, and III. ii, 11,
pi. liv. fig. 1: Fabricius, Ent. Syst, IT. 446, and Suppl. p. 337.
Cancer (Galene) bispinosus, De Haan, Faun. Japon. Crust. p. 49, pl. v. fig. 2.
Galene bispinosa, A. O. Walker, Journ. Linn. Soc., Zool., XX. 1886-90, p. 110,
- Carapace moderately broad, somewhat pentagonal, its surface for
the greater part smooth, but usually scabrous near the borders—espe-
cially the postero-lateral borders; its surface is also somewhat lumpy,
1898.] A. Alcock — Carecinological Fauna of India. 137
owing to the very broad depressions which somewhat vaguely delimit
and to a certain extent subdivide the regious. Pterygostomian region
more or less hairy.
The antero-lateral borders are very indistinctly 4-lobed, the first
lobe being almost obsolete, the second being usually marked by a
granular denticle, aud the third and fourth by two coarse granular
spines. The postero-lateral borders, which are little convergent, are
slightly longer than the chord of the antero-lateral. The posterior
border is about half the greatest width of the carapace.
Front really bilobed, but with both the inner and the outer angles
of each lobe so equally prominent as to appear 4-dentate.
Chelipeds unequal: exposed surfaces of arm either smooth, or more
or less scabrous, both borders of arm uneyen and hairy, the distal end
of the upper border with two strong teeth: both the inner and the
outer angles of thé wrist well pronounced, or even spiniform; the
exposed surfaces of the wrist may be almost smooth, but are usually
studded, to a variable extent, with sharp little tubercles; the upper
outer and lower surfaces of the hand may be almost smooth, but are
usually studded, in the proximal third to three-quarters, with similar
tubercles, in more or less distinct lines: fingers long, sharp-pointed,
the apposed edges with strong molariform teeth.
Legs long, stoutish ; upper border of the meropodites scabrous and
spinulate, upper border of last 3 joints, and lower border of last 2, plumose.
Colours in spirit, leaden white or yellowish.
In the Indian Museum are 3 specimens from the Vizagapatam
coast and 1 from Tennasserim, (besides one from Hongkong): the
amount of granulation of the borders of the carapace and of the cheli-
peds is different in all.
Subfamily II. ACTAEHINAI.
Actm#a, De Haan, A, Milne Edwards.
Actza, De Haan, Faun. Jap. Crust. p. 18.
Actxa and Actxodes, Dana, U. S. Expl. Exp. Crust. pt. I. pp. 162, 194.
Actexa, Heller, Crust. Sudl. Europ. p. 69.
Actza, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, pp. 259, 260.
Psaumis, Kossmann, Crust. roth. Meer. p. 26.
Actza and Actzxodes, Miers, Challenger Brachyura, pp. 118, 135.
Carapace convex fore and aft, slightly convex or flat from side to
side, usually broad, the regions well demarcated by deep grooves, and
again subdiyided into lobules, which are usually convex and granular.
Antero-lateral borders usually four-lobed, but the lobes are shallow and
often indistinct. Postero-lateral borders usually concave, always short,
not strongly convergent.
air. 18
138 A. Alcock—Carcinological Fauna of India. [No. 1,
Front between a third and a fourth the greatest width of the
carapace, deflexed, cleft in the middle line into two lobes. Upper edge
of orbit tumid, usually with two fissures or sutures; a third below the
outer orbital angle: eyestalks short and thick.
Antennules folding obliquely or nearly transversely. Basal anten-
nal joint usually stopping at the angle of the deflexed front, but often
prolonged beyond this, towards or nearly into the orbit; the flagellum
is about as long as the orbit, and is lodged in the orbital hiatus.
Merus of the external maxillipeds with the anterior border little
oblique,
Chelipeds equal in both sexes; fingers usually blunt-pointed, some-
times hollowed-out at tip.
Abdomen of the male five-jointed, somites 3, 4, 5 fused.
Small crabs, distinguished by the elaborate lobulation of the cara-
pace, and by the form of the front, which is usually deep-cleft in the
middle line to form two prominent round-pointed lobes.
Key to the Indian species of Actzea.
I, Legs of ordinary form :—
i. The lobules of the carapace, and the legs, when granular,
bear miliary or vesiculous granules of nearly uniform
size, not tubercles :—
1. Length of the carapace two-thirds or less than
two-thirds the breadth, postero-lateral borders ex-
tremely short and concave :—
a. Carapace and legs covered witha short dense
fur, which does not, however, conceal the
lobules or their granules :—
a. Fingers hollowed at tip, fur black ... A. tomentosa.
B. Fingers long and pede fur light
DEOWDjictepoh os a ~. A. areolata,
b. Carapace and legs wie numerous Peeties
which do not form a coat; fingers blunt-
pointed, but not appreciably hollowed at tip. A. hirsutissima,
2. Length of the carapace rather more than two-
thirds the breadth, postero-lateral borders slightly
concave :—
a. Legs and chelipeds lobulated in the same
style as the carapace :—
a. Lobules of the carapace very marked-
ly isolated and very convex, inter-
lobular grooves very broad and deep,
ane hamy |... 44,. Samons .. <A, rufopunctata,
8B. Lobules of the Sema pti me Pe hearich
ably isolated, the grooves with a short
almost Invisible FUL... s..cessnseenter yeeces .- Ae SMECIORRE
1898. | A. Alcock — Carcinological Fauna of India.
b. Wrist and hands sublobular, corresponding
joints of legs only a little dimpled (areola-
tion of carapace complete) . ........:s00 esses:
c. Chelipeds and legs with a atin peer se
surface, areolation of carapace faint ante.
riorly, incomplete posteriorly :—
a lLobulation of antero-lateral border
' very indistinct, no hairs on the cara-
pace, which is thick and convex ..... :
B. Lobulation of antero-lateral border
fairly distinct the lobules being granu-
lar ; a thin coat of hair :—
w. Carapace of ordinary Actza
ROEURE! Ades torancvs” ageusless eunaesites
y. Carapace more than ? as long
as broad ; its posterior half re-
markably flat..... ;
ii, Carapace covered with tubercles, legs with hakGtaies’ or
spines :-—
1. Carapace with plain isolated tubercles :—
a. Carapace with pearly tubercles and granules ;
front bilobed, but each lobe so deeply ex-
cised as to appear itself bilobed............ CAME
b. Carapace with coarse spine-like tubercles ;
front broadly) bilobed:s,.s06s..decseachsceccees
c. Carapace and chelipeds with pedieledt pisi-
form tubercles, legs with thorns .........
d. Carayace, chelipeds and legs with se
flat-topped tubercles which at the margins
become petaloid; front bilobed, each lobe
cut into four petaloid teeth... ....0.....sc-scseceee
2. Carapace closely covered with cemiinee edhe
the surfaces of which are themselves formed of
confluent granules :—
a. Tubercles of carapace very rough, raspberry-
like, some of those on the legs often spiny:
carapace about seven-ninths as long as broad.
b. Tubercles of zarapace smooth though pitted,
those of the legs never spiny: carapace
about two-thirds as long ag broad....... aa opeicne
II. Propodites and carpopodites of legs dorsally bicarinate in such
a way that the space between the crests appears like a trough
or a series of cups :—
i, Propodites and carpopodites each with one trough;
lobules of carapace granular; front not projecting much.
ii. Carpopodites with at least two cups ; lobules of carapace
pitted as well as granular; front projecting far beyond
Semana anele Of LG OLD srercetensadeaisensrysevsedsensnpenncas
139
A, ruppellii.
A, obesa.
A. pulchella,
A. parvula,
A. nodulosa.
A. echinus.
A. peronii.
A, flosculata,
A, granulata.
A, calculosa,.
A. cavipes,
A. fossulata,
140 A. Alcock — Carcinological Fauna of India. [No. 1,
63. <Actzxa tomentosa, (Edw.) A. Milne Edwards.
Zozymus tomentosus, Milne Edwards, Hist. Nat. Crust. I. 385, and in Cuvier.
Réene An. Crust. pl. xi. bs, fig. 2.
Actza tomentosa, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 262, and
IX. 1873, p. 191: A. Targioni Tozzetti, ‘‘ Magenta” Crost. p. 35, pl. iii. figs. 14 &.:
Hilgendorf, MB. Ak. Berl. 1878, p. 788: Richters in Mébius Meeresf. Maurit.
p. 145: Haswell, Cat. Austral. Crust. p. 44: Ortmann, Zool. Jahrbuch., Syst. &e.,
VIT. 1898-94, p. 453, and in Semon’s Zool. Forschungsr. (Jena. Denkschr. VIII)
Crust. p. 50.
Actzodes tomentosus, Dana U. 8. Expl. Exp. Crust. pt. I. p. 197: Stimpson,
Proc. Ac. Nat. Sci. Philad. 1858, p. 32: Heller, SB. Ak. Wien, XLIII. 1861, p. 328,
and Novara Crust. p. 17: Miers, P. Z. S. 1877, p. 184; and 1879 pp. 20 and 30; and
Phil. Trans. Vol. 168, 1879, p. 486; and Ann. Mag. Nat. Hist. (5) V. 1880, p. 234;
and Zool. H. M. 8. Alert, pp. 517 and 530; and Challenger Brachyura, p. 135:
de Man, Archiv. fur Naturges. LIII. 1887, i. p. 252; and in Weber’s Zool. Ergeb,
Niederl. Ost. Ind. II. 1892, p. 278; and Zool. Jahrb., Syst. &c., VIII. 1894-95, p. 499:
Cano, Boll. Soc. Nat. Napol. III. 1889, p. 199.
Carapace ovoid and very broad, its greatest length less than two-
thirds its greatest breadth, its dorsal surface—like that of all the
surfaces of the chelipeds and legs that are exposed in repose —covered,
as closely and evenly as possible, with a dense short blackish felt
through which peep the shiny tops of very numerous large vesiculous
granules. This felt is not so long as to obscure the areolation of the
carapace which is very perfect and in bold relief, but it obscures the
fact that the deep-cut grooves that separate the lobules are smooth.
The lobules—excluding those of the antero-lateral and supra-
orbital margins and those on the front—are 21 in number, the anterior
8 with the long diameter fore-and-aft, the posterior 5 with the long
diameter transverse.
The front, which is vertically deflexed and does not break the wide
even sweep of the antero-lateral borders, appears nearly equally four-
lobed, the outer lobe on either side being formed by the tumid supra-
orbital border.
The antero-lateral borders are long and beautifully arched; when
undenuded they look entire, but when denuded they are seen to be cut
by narrow clefts into four very shallow lobes of unequal size,—the
clefts being continued as grooves on to the under surface of the cara-
pace. The very short postero-lateral borders are extremely concave.
The tumid supra-orbital border is cleft into lobules by two fissures
similar to the grooves of the carapace, and there is a third fissure at
the outer angle of the orbit.
The whole under surface of the carapace, and the surfaces of the
sternum and external maxillipeds and abdominal terga, are covered
with a dense felt that obscures all the granulation that exists.
1898. ] A. Alcock — Carcinological Fauna of India. 141
The basal antennal joint is broad, and its outer angle does not fall
very far short of the inner angle of the floor of the orbit.
The legs, besides the felt and the granules (which are conical
rather than vesiculous) already spoken of, have their edges — but chiefly
the anterior edge—fringed with coarse tufted hair: similar hair
occurs on the edges of the arm.
The fingers are short, with broadly-rounded hollowed-out tips.
Colours in spirit, as in life, blackish.
In the Indian Museum are 115 specimens from the Nicobars, Anda-
mans, Palk Str., and Laccadives (besides 31 from Mauritius, Australia
and the South Sea Is.).
64. Actxa areolata, Dana.
Actzxa areolata, Dana, Proc. Acad. Nat. Sci. Philad. 1852, p. 73, and U. S. Expl.
Exp. Crust. pt. I. p. 162, pl. viii. figs. la-b: A. Milne Edwards, Nouv. Archiv. dn
Mus. I. 1865, p. 264: EH. Nauck, Zeits. Wiss. Zool. XXXIV. 1880. p. 54 (gastric
teeth): ? Miers, Zool. H. M.S. Alert, pp. 182, 209: de Man, Journ. Linn. Soc., Zool.,
XXII. 1887-88, p. 25: Zehntner, Rev. Suisse Zool. II. 1894, p. 147.
Carapace in proportions and outline almost exactly similar to
A. tomentosa. Its surface and the exposed surfaces of the chelipeds and
legs is also covered as densely as possible with felt; but this felt is
of a lighter colour, and it almost conceals the granulation, excepting 3
or 4 lines of granulation on the lower outer surface of the hands: the
felt also obscures, though it does not conceal, the areolation of the
carapace, owing to the lobules being less convex.
Unlike A. tomentosa the upper surface of the wrist and hand are
nodular (as well as granular), and the fingers are long and pointed,
without any hollowing of the tip.
In other respects this species closely resembles the preceding.
One specimen from Meregui.
65. Actsa hirsutissima (Rippell), De Haan, Dana.
Xantho hirsutissimus, Ruppell, 24 Krabben roth. Meer, p. 26, pl. v. fig. 6: Milne
Edwards, Hist. Nat. Crust. I. p. 389.
Actwa hirsutissima, De Haan, Faun. Japon. Crust. p. 18: Dana, U, S. Expl. Exp.
Crust. pt. I. p. 164: Heller, SB. Ak. Wien, XLIII. 1861, p. 314; and Novara Crust.
p- 9: A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 268, and IX. 1873,
p. 191: Kossmann, Reise roth. Meer., Crust. p. 23: Targioni Tozzetti, Magenta
Crost. p. 37, pl. iii. fig. 26: Richters in Mébius, Meeresf. Manrit. p. 145: de Man,
Notes Leyden Mus. II. 1880, p. 173, and III. 1881, p. 96: Cano, Boll. Soc. Nat.
Napoli, ITI. 1889, p. 189: Ortmann, Zool. Jahrb., Syst., VII. 1893-94, p. 453.
Length of carapace = § breadth.
142 A. Alcock — Carcinological Fauna of India. [No. 1,
Carapace of much the same proportions and outline as A. tomentosa,
but the frontal outline is more convex, and the postero-lateral borders
are a little less concave. |
The surface of the carapace is very completely areolated by deep
smooth grooves, the lobules being exceedingly numerous, strongly convex,
and closely covered with pearly granules; and between and around the
bases of the granules are many short black bristles which do not form
a coat or conceal the texture of the carapace.
The exposed surfaces of the chelipeds and legs are granular and
bristly, like the carapace ; and the carpal joints, and to a less extent
the propodites are dimpled, but not distinctly nodular, above.
Under surface of carapace granular, hairy, and furrowed by grooves
continued from fissures that subdivide the antero-lateral borders into
four shallow lobes. The surfaces of the external maxillipeds and distal
abdominal terga are bristly, those of the sternum and proximal ab-
dominal terga are hairy.
Fingers bluntly pointed but not hollow at tip.
Colours in spirit, yellowish, fingers and greater part of hand black.
In the Indian Museum are a specimen from Samoa, a specimen
without locality, and a specimen from the Andamans or Nicobars.
66. <Actea rufopunctata, (Edw.) Heller.
Xantho rufopwnctatus, Milne Edwards, Hist. Nat. Crust. I. 389: Lucas, Expl..
Sci. Algerie, Anim. Artic. p. 11, pl. ii. fig. 1: A. Milne Edwards in Maillard’s Vile
Réunion Annexe F, p. 4.
Actxa rufopunctata, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 268,.
pl. xviii. figs. 1, la: Richters in Moébius Meeresf. Maurit. p. 145: de Man, Notes
Leyden Mus. IT. 1880, p. 172 and III. 1881, p. 96: Miers, P. Z. S. 1881, pp. 68, 68 ;
and Zool. H. M.S. Alert, pp. 517, 528; and Challenger Brachyura, p. 122: Carus,
Prodr. Faun. Medit. I. p. 518: R. I. Pocock, Ann. Mag. Nat. Hist. (6) V. 1890, p. 75:
J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 357: Ortmann, Zool.
Jahrb., Syst., VII. 1893-94, p. 454; and in Semon’s Forschungsr. (Jena. Denk. —
VIII.) Crust. p. 50. ‘
Actxa nodosa, Stimpson, Ann. Lyc. Nat. Hist. N. Y. VII. 1862, p. 203; and
Bull. Mus. Comp. Zool. II. 188: A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865,
p. 266, pl. xvii. figs. 6-6c ; and Exp. Sci. Mex., Crust. p. 245; and Bull. Mus. Comp.
Zool. VIII. p. 11: Desbonne and Schramm, Crust. Guadaloupe, p. 25: J. S. Kingsley,
Proc. Acad. Philad. XX XI. 1879, p. 393.
Carapace broad, ovoid, its extreme length not quite ¢ but more
than 2 its extreme breadth: its surface is broken, by deep and broad
grooves, into numerous (about 27 excluding those round the orbits and
the front) very convex lobules, which are covered very closely with
large vesiculous granules; the grooves are filled with a dense short
1898. | A, Alcock — Carcinological Fauna of India. 143
felt — with longer hairs sometimes interspersed —against which the
lobules stand out like islands. (Occasionally there are some tufts of
jong hair on the edge of some of the lobules).
The exposed (dorsal) surfaces of the carpal and propodal joints of
the chelipeds and legs are lobulated in the same style as the carapace,
the lobules being granular and being isolated by deep felted grooves.
The front is strongly deflexed, but somewhat prominent, and is
rather sharply bilobed. The tumid supra-orbital margin is broken by
two cross grooves, and is separated from the lower margin of the orbit
by a fissure. The antero-lateral borders are cut into four rounded
lobules of nearly equal size, by deepish fissures. Postero-lateral borders
not appreciably shorter than the antero-lateral, and little concave.
The parts seen on the under surface are not conspicuously granular
or hairy.
The basal antennal joint has its outer els almost in contact with
the inner angle of the lower edge of the orbit.
The edges of the legs (especially the upper edge) are fringed with
coarse hair—as also of the arm.
The lower outer surface of the hand has the granules arranged in
lines, as is the case with most species of Actwa. Fingers blunt-pointed,
hollowed out at tip.
Colours of well-preserved spirit specimens, yellow with the con-
vexities of some of the lobules orange-red; the felt in the grooves
brown ; fingers dark brown with white tips.
Five specimens from Ceylon, up to 34 fms., and four from the
Andamans, up to 36 fms.
67. Actexa speciosa (Dana), Ortmann.
Actxodes speciosus, Dana, U. S, Expl. Exp. Crust. pt. I. p.198, pl. xi. figs. 4a-c:
Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 32.
Actxa speciosa, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 274:
Ortmann, Zool. Jahrb. Syst. VII, 1893-94, p. 455.
Actzxodes nodipes, Heller, Abhandl. zool.-bot. Ges. Wien, XI. 1861, p. 9, and
SB. Ak. Wien, XLIII. 1861, p. 329, pl. ii. fig. 19, and Novara Crust. p.17: A. Milne
Edwards, Nouv. Archiv. du Mus. I, 1865, p. 274: de Man, Notes Leyden Mus. II.
1880, p. 172.
? Psaumis glabra, Kossmann, Reise roth. Meer., Crust, p. 27, pl. i. fig. 4.
This species has a general resemblance to A. rufopunctata, but the
carapace is relatively longer and narrower, its lobulation is much less.
complete and bold, and it is devoid of hairs.
Length of carapace = { the breadth.
Surface of carapace broken up by shallow grooves into numerous
144 A. Alcock — Carcinological Fauna of India. [No. 1,
lobules, which fall into series that appear to radiate from the midcardiac
region. The lobules are closely covered with miliary granules, they
are nowhere very convex, and on the posterior third of the carapace
they are indistinct. Although the carapace looks quite bare to the
naked eye, yet its whole surface —both between the granules and in
the grooves between the lobules—is covered with a fine, extremely
short and inconspicuous felt.
The exposed surfaces of the carpal and propodal joints of the legs
and chelipeds (except the lower outer surface of the hand, which is
granular in lines) has exactly the same style of sculpture and texture
as the antero-lateral part of the carapace.
Two fissures in the upper edge of the orbit, but none between this
and the lower edge of the orbit.
Antero-lateral borders four-lobed, —the lobes subequal and shallow,
Postero-lateral borders distinctly shorter than the antero-lateral and
distinctly concave.
The basal antennal joint falls far short of the inner angle of the
floor of the orbit.
Kidges of legs and chelipeds quite free from hair.
Fingers pointed, very slightly hollowed at tip.
Colours in spirit, yellow.
3 specimens from the Persian Gulf, Ceylon, and Andamans are in
the Indian Museum.
68. Actza ruppellii (Krauss) Hilgendorf.
Aegle ruppellii, Krauss, Siidafr. Crust. p. 28, pl. i. fig. 1. 1843.
Actxa ruppellii, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 270:
Hilgendorf in v. d. Decken’s Reisen Ost Afr, III. i. p. 73, and MB. Ak. Berl. 1878,
p. 787: Miers, Ann. Mag. Nat. Hist. (5) V. 1880, p. 232, and Zool. H. M. S. Alert,
pp. 183, 209: A. O. Walker, Journ, Linn. Soc., Zool., XX. 1886-90, p. 109:
Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 358: Ortmann, Zool. Jahrb.,
Syst., VII. 1893-94, p. 454: de Man, Zool. Jahrb., Syst., VIII. 1894-5, p. 499.
Aegle rugatu, Adams and White, Samarang Crust. p. 43, pl. viii. fig. 5.
Actaea rugata, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 269, and
TX. 1878, p. 192: R. Etheridge jr., Mem. Austral. Mus. No. 2, 1889, pp. 33, 35:
de Man, Notes Leyden Mus. XIII, 1891, p. 1, and in Weber’s Zool. Ergebn. Niederl.
Ost-Ind. II. 1892, p. 277: Whitelegge, Mem. Austral. Mus. III. 1897, p. 129.
Actza rufopunctata, de Man (nec Edw.) Archiv. fur Naturges. LIII. 1887, i.
p- 261, and Journ. Linn. Soc., Zool., XXII. 1887-88, p. 26 (see Notes Leyden Mus.
XIII. 1891, p. 1).
Carapace ~ as long as broad; it and the exposed surfaces of the
legs covered with a shaggy coat—consisting of a sponge-work of short
bristles amid which are numerous long silky tangled hairs—which has
1898.]} A. Alcock— Carcinological Fauna of India. 145
to be removed before the sculpture and texture of the carapace can be
' properly made out.
On the denuded carapace the lobules are numerous, are arranged
in series which appear to radiate from the cardiac region, and are
somewhat indistinct quite posteriorly, but are elsewhere distinct, moder-
ately convex, and separated by broad smooth furrows.
Front obliquely deflexed, rather sharply bilobed. Supra-orbital
margin moderately tumid, narrow, cut by two fissures and separated
from the lower edge of the orbit by a fissure. Antero-lateral borders
four-festooned, the first and last lobes much smaller than the others.
Postero-lateral borders shorter than the antero-lateral, moderately
concave.
_ Exposed (dorsal) surfaces of the chelipeds and legs granular and
shaggy: the wrist and upper surface of the hand are also subnodular,
but the corresponding joints of the legs are but indistinctly grooved.
Lower outer surface of hand with granules in lines that are not so
definite as usual. Fingers blunt-pointed, slightly hollowed at tip.
Basal antennal joint broad: it falls short of the inner angle of the
orbit.
Colours in spirit yellow, fingers dark brown with white tips: in
some specimens faint orange-red spots exist on some of the lobules of
the carapace. |
Carapace markediy more convex in the female than in the male. .
In the Indian Museum are 30 specimens from Malacca Str., Anda-
mans, Mergui, Ceylon, and Persian Gulf,
69. ? Acteea obesa, A. Milne Edwards.
Acta obesa, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 272, pl. xvii.
figs, 2-2b.
This species, if my identification be correct, differs from Actxa
ruppellu, which it closely resembles, in the following characters :—
(1) the carapace and legs are not shaggy, and though they bear
some hairs these do not in any way conceal the texture of the carapace :
(2) the lobulation is quite absent from the posterior third of the
carapace; and elsewhere though quite distinguishable, is extremely
faint, owing to the fineness of the’ grooves :
(3) the entire surface of the carapace—grooves as well as lobules
—is covered with crisp granules, which are largest in tle middle of the
branchial regions : |
(4) the lobulation of the antero-lateral borders, though distinguish-
able, is extremely indistinct, especially in the case of the first lobe;
J. -u1. 19
146 A. Aleock — Carcinological Fawna of India. [No. },
(5) the postero-lateral borders are more concave :
(6) the dorsal surfaces of the chelipeds and legs are crisply granu-
lar, but the carpal joints show almost no dimpling and the propodal
joints none at all.
In the Indian Museum are 2 specimens from the Malacca Straits
and one from Bombay.
70. ? Actxa pulchella, A. Milne Edwards.
Actxa pulchella, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 273,
pl. xvii. figs. 5-5b.
? Actzodes modestus, de Man, Archiv. fiir Naturges, LITI. 1887, i. p. 257, pl. ix.
fig. 3,
This species, if my identification be correct, resembles Actwa
ruppellii and obesa, but has the following differential characters :—
(1) the carapace is altogether less convex :
(2) the lobulation of the carapace is defective posteriorly, and is
faint anteriorly owing to the fineness of the grooves :
(3) the entire surface of the carapace and dorsal surface of legs
and chelipeds is covered (though not crowded) with crisp granules,
most of which carry a short bristle— but these bristles are not close
enough to form a coat:
(4) the antero-lateral borders are sharply granular, but their
lobulation is very indistinct :
(5) except for a furrow across the wrist parallel with the articula-
tion of the hand, the sharply granular surface of the chelipeds and legs
is unbroken.
In the Indian Museum are three specimens from Mergui, the Anda-
mans, and Ceylon.
71. Acta parrula, (De Haan), de Man.
Menippe parvulus, De Haan, Faun. Japon, Crust. p. 21: Krauss, Sudafr. Crust;
p. 34, pl. ii. fig. 2.
Actxa parvula, de Man, Journ. Linn. Soc., Zook, XX. 1887-88, p. 27.
Carapace more than ¢ as long as broad, with the posterior third or
more depressed, perfectly flat, and almost or quite devoid of areolation ;
its surface is everywhere covered with vesiculous granules, which
become very small posteriorly, and with a fine short velvety hair: the
areolation is fairly profuse and quite distinct in the anterior two-thirds.
Antero-lateral borders divided into four granular lobes: postero-
lateral borders less convergent than in any other species of Actxa,
giving the flattened posterior part of the carapace a most abnormal look
1898. | A. Alcock — Carcinological Fauna of India. 147
for an Actza. Front deeply cleft into two round-pointed lobules. The
three grooves near the outer angle of the orbit are fairly distinct.
Upper and outer surface of wrist and hand covered with pearly
granules and velvet: fingers pointed, not hollowed at tip.
The exposed parts of the dorsal surface of the legs are also covered.
with velvet which conceals their sharply granular sculpture. Last pair
of legs rather short.
Colours in spirit yellow or brownish, fingers brown.
In the Indian Museum are 3 specimens, from the Andamans and
Mergui., ; |
This species may be distinguished from all its congeners by the
very moderate difference between the two diameters of the carapace
which also has its posterior part quite flat.
3
72. Actxa cavipes, (Dana), A. Milne Edwards.
Actzodes cavipes, Dana, Proc. Acad. Nat, Sci. Philad., 1852, p. 78, and U. SQ.
Expl. Exp., Crust. pt. I. p. 199, pl. xi. figs. 5a—b.
Actza cavipes, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 280, and
IX. 1873, p. 193: Ortmann, Zool. Jahrb. Syst. VII. 1893-94, p. 456, and in Semon’s
Zool. Forschungsr. (Jena, Denk. VIII) Crust. p. 50.
Carapace about 2 as long as broad, completely lobulated, the lobules
being covered with miliary granules and being separated by broad but
not very deep grooves: the posterior part of the carapace sometimes
has a worm-eaten appearance.
Front obliquely deflexed, with a cupid’s-bow-shaped edge, hardly
projecting beyond the prominent inner angle of the lower edge of the
orbit. Orbital margin unfissured and unbroken. Antero-lateral borders
4, or indistinctly 5, lobed, the lobes granular and uneven, but not pitted.
Postero-lateral borders very much shorter than the antero-lateral,
concave.
Outer surface of wrist with numerous pits and craters, upper outer
surface of hand worm-eaten.
The upper edges of the carpal and propodal joints of the legs have
each a double longitudinal crest, and in every joint the ends of the crests
meet so as to leave a trough-like space between them.
The basal antennal joint almost touches the inner angle of the
orbit.
Fingers long, pointed, slightly hollow at tip.
In the Indian Museum are four specimens from the Andamans,
Mekrdan Coast, and Persian Gulf (besides specimens from Upolu and
Mauritius). fae
L438 A. Aleock — Carcinological Fauna of India. [No. ],
73. Acta fossulata (Girard) A. M. Edw.
Cancer fossulatus, Girard, Ann, Soc. Entom. France (3) VIT, 1859, p. 149, pl. iv.
figs. 2—2b.
‘ Actzwa schmardx, Heller, Abh. zool.-bot. Ges. Wien, 1861, p. 6 and SB. Ak.
Wien, XLIII. 1861, p. 318, pl. i. fig. 13.
Actzxa fossulata, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 279, and
1V. 1868, p. 71: Richters in Mébius Meeresf. Maurit. p. 145.
Psaumis fossulata, Kossmann, Reise roth. Meer., Crust. p. 27, pl. i. fig. 3.
Closely resembles Actxa cavipes, but has the following difference:—
(1) the front projects far beyond the inner angle of the orbit:
(2) the lobes of the carapace have their convexity distinct but
boundaries somewhat indistinct; and in addition to being granular,
they are deeply pitted, and this gives the whole carapace a worm-eaten
look:
(3) the antero-lateral borders are four-lobed, but the first. lobe is
very indistinct, and the lobes are marked with rather large pits :
(4) the upper edge of the hand is bluntly crested and the neighbour-
ing surface is pitted rather than eroded :
(5) the crest of the carpal joints of the legs do not only meet at
their ends, but are also more or less completely joined across the middle
by dissepiments, so that instead of enclosing a single trough they form
at least two irregular cup-like cavities. =
In the Indian Museum are two specimens from Great Coco I.
(Andamans), and Hast I., Andamans.
74. Actzea nodulosa, White.
Actza nodulosa, White, P. Z. S. 1847, p. 224: Ann. Mag. Nat. Hist. (2) II, 1848,
pe 224; and Adams and White, Samarang Crust. p. 389, pl. viii. fig. 4: A. Milne
Edwards, in Maillard’s ile Reunion, Annexe F, p. 5; and Nouv. Archiv. du Mus.
I. 1865, p. 277: Miers, Challenger Brachyura, p. 120: Henderson, Trans. Linn. Soc.,
Zool., (2) V. 1893, p. 356.
Carapace 3 as long as broad, much subdivided by smooth well cut
grooves into numerous small lobules, These lobules are rather irregular-
ly studded with pearly tubercles and granules, the slight irregularity
in size and distribution of which gives the lobules themselves a some-
what irregular look. On several of the lobules of the gastric cardiac
and branchial regions are, sometimes, symmetrically disposed tussocks
of long coarse whitish hair,
The obliquely deflexed front is sharply four-lobed or four-toothed,
the outer lobe on either side (standing at the orbital angle) being small.
The beaded supra-orbital margin is broadly fissured twice and is
separated from the infra-orbital margin by a fissure. The antero-
1898. | A. Aleock — Carcinological Fauna of India. 149
lateral borders are sharply four-lobed, each lobe being rasp-like. The
postero-lateral borders are a little concave. The posterior border is
formed by a row of bead-like granules, in front of which is another row
— broken in the middle—of larger beads. The sternum, and the under
surface of the carapace as far as the beaded epimeral suture, are covered
with vesiculous granules.
Those surfaces of the chelipeds and legs that are exposed in repose
are closely and crisply granular, many of the granules being pearl-like
or bead-like, and those along the dorsad border being spine-like: the
edges of the legs, especially the upper edge, are hairy. The granules
on lower outer surface of hand are arranged as usual in lines.
Fingers short, pointed, not hollow at tip.
The basal antennal joint falls far short of the inner angle of the
floor of the orbit. .
: Colours in spirit, white.
In the Indian Museum are 3 specimens from off the Malabar Coast,
98-29 fms., one from the Persian Gulf, and one from the Andamans.
- Acteea nodulosa var. bullifera.
In this well-marked and very ornamental variety the lobulation of
the carapace—both of its surface and of its antero-lateral borders—is as
deeply cut, as convex, and as regular as it is in Actw#a rufopunctata ; the
tubercles are more of one size, and have a distinct constricted base and
a swollen spherical pearl-like top; the front row of pearly granules of
the posterior border is unbroken ; the front is more bilobed with sinuous
edges than four-lobed, and its edge is elegantly denticulated or beaded;
and all the parts of the under surface of the body are finely granular,
except the sternum, which has a pitted worm-eaten look,
Length of carapace 10 millim., breadth 14°5 millim.
A single female from the Audamans. —
7d. Actsea echinus, n. sp.
Closely resembles Actwa nodulosa White, but has the following
difference :—
(1) instead of pearly tubercles we find coarse conical tubercles or
tooth-like spines with denticulated tops; and on the chelipeds and
autero-lateral borders of the carapace are coarse serrated spines:
(2) the front is broadly bilobed, the angles of the lobes being sharp :
(3) the lobulation of the antero-lateral borders is irregular and
indistinct :
(4) the legs are thin and compressed, the dorsad border of the
carpal and propodal joints forms a serrated crest, and the slightly
150 A, Alcock — Carcinological Fauna of India. [No. 1,
grannlar sculpture of the upper surfaces of these joints and of the
meropodites is concealed by a close short spongy growth of hair.
Colours in spirit yellow, fingers brown.
Length of carapace 17°5 millim., breadth 26 millim.
A single male from off the Malabar Coast 29 fms. |
This species (P) may perhaps be only a variety of 4. nodulosa. I
have noticed it separately, and have figured it, on account of the
resemblance it bears to Herbst’s Cancer ee (Krabben {IT. i. 33,
pl. li. fig. 2).
76. Actwa peronii,, (Edw.)-Haswelh...
Xantho peronii, Milne Edwards, Hist. Nat. Crust. I. 892: Hess, Archiv. fir
Naturges. XX XI. 1865, i. pp. 133, 171.
Xantho spinosus, Hess, Archiv. fiir. Nat. XXXI, “1865, Jaa 132, V7: de Man,
Zool. Jahrb., Syst., II. 1887, pp. 690, 692.
Actza peronti, Haswell, Cat. Austral. Crust, Pp. 46 : Miers, Challewaee Brachyura,
p. 122: de Man, Zool. Jahrb. Syst., II. 1887, pp. 690, gee J. a5 Henderson, Trans.
Linn. Soc., Zool., (2) V. 1893, p. 357.
? Chlorodius polyacanthus, Heller, Abh. zool.-bot. Ges. Wien, 1861, p. 11;
SB. Ak. Wien, XLIII, 1861, p. 339, pl. ii. fig. 21.
Carapace § as long as broad, only moderately convex, the lobulation
distinct but not convex, covered with strongly convex well-isolated
smooth polished tubercles, which are largest on the branchial regions
and are smallest posteriorly, where also they become somewhat squami-
form. On the antero-lateral borders are four (excluding a tubercle
of the supra-orbital series) large tubercles, much similar to those on
the branchial regions, but more prominent (almost stalked) and nen
larger tops (almost pisiform).
Exposed surfaces of wrist and hand covered with tubercles much
like those of the branchial regions, but more prominent: on the lower
outer surface of the hand they become almost squamiform: a few
tubercles at the distal inner corner of the arm.
Exposed surfaces of carpal and propodal joints of legs covered
with stout thorns: smaller thorns on upper edge of meropodites, and
still smaller ones on surface of dactyli except on the claw. :
Front broadly bilobed: each lobe with an S-shaped curve to the
edge and with the outer angle pronounced.
Fingers short, blunt pointed, hardly hollow at tip.
The basal antennal joint stops far short of the 1 inner r angle of the
floor of the orbit.
Colours in spirit light yellow, oe. dark Goon
In the Indian Museum are 3 specimens from Australia but none
from India. It is included here on the authority of Dr. J. R. Henderson.
1898.] A. Alcock — Carcinological Fauna of India. 151
77. Actxa flosculata, n. sp.
Nearest to Actza acantha, A. M. E., and A. hystrix, Miers.
Characterized by the close ey ae (carapace, chelipeds and
legs) of peculiarly ornamental fungiform tubercles which become petaloid
at the ee
Carapace ? as long as broad, convex; all the regions well defined
by conspicuous grooves, and convex ; the regions again subdivided into
few convex lobules by less conspicuous grooves. The whole carapace,
except the broader grooves between the regions, closely covered with
very elegant tubercles which have constricted stalk-like bases and thin
broad oval or kidney-shaped tops. The exposed surfaces of the cheli-
peds and legs are covered with ornaments similar to those on the
carapace, except at the edges, where they become petaloid.
Front broadly bilobed, the edge of each lobe being deeply cut into
four projecting petals. Supra-orbital margin deeply scallopped : antero-
lateral borders ornamented like the surface, obscurely lobed: postero-
lateral borders about equal in length to the antero-lateral, straight.
The terminal abdominal terga and the sternum of the male pitted
and worm-eaten, but with a glazed appearance: under surface of cara-
pace, as far as the epimeral suture, covered with pearly granules.
The basal antennal joint stops far short of the inner angle of the
floor of the orbit.
Fingers short, blunt pointed, slightly hollow at tip.
Colours in spirit ight yellow, fingers brown with white tips.
In the Indian Museum are two specimens from off Ceylon, 34 fms.
and one from off Maldive Is. 28 fms. The carapace of the larger one
is 8 millim. long and 12 millim broad.
The ornamentation of this species gives it a strong resemblance to
Chlorodius fragifer White, with which it may probably prove “0 be
identical.
78. Actsea granulata (Audoin).
Cancer granulatus, Savigny and Audouin, Description de l’Egypte, Crust. pl. vi.
fig. 2.
Cancer savignyi, Milne Edwards, Hist. Nat. Crust. I. 378.
Cancer (Actea) granulatus, De Haan, Faun. Japon. Crust. p. 47.
Actxa carcharias, White, P. Z. S. 1847, p. 224, Ann. Mag. Nat. Hist. (2) IT.
1848, p. 284: A. Milne Edwards, Nouv. Archiv. du Mus. 1. 1865, p. 276,
Actxa pura, Stimpson, Proc. Acad. Nat. Sci. Philad. 1858, p. 32.
Actea granulata, A. Milne Edwards, Nouv. Archiv. du Mus, J. 1865, p. 275, and
IX. 1873, p. 192: Miers, Cat. Crust. New Zealand, p. 16; and P. Z. 8S. 1879,
pp. 20, 30; and Challenger Brachyura, p. 120: Haswell, Cat. Austral. Crust. p. 44:
152 A. Alcock — Carcinological Fauna of India. [No. 1,
Filhol, Crust. New Zealand, p. 373: J. R. Henderson, Trans. Linn, Soc., Zool,
(2) V. 1898, p. 356: Ortmann, Zool. Jahrb., Syst. VII. 1893-94, p. 455.
Actxa Savignii, Kossmann, Reise roth.-Meer., Crust. p. 25: Hilgendorf, MB. Ak.
Berl. 1878, p. 787: Cano, Boll, Soc. Nat. Napol. III. 1889, p, 189,
Carapace nearly as long as broad, of a mulberry-like appearance,
owing to its entire surface being covered with rough tubercles in the
closest possible contact with one another by the base. Hach individual
tubercle again has a mulberry-like appearance, since it is formed of a
number of facetted granules confluent by their bases.
The lobulation of the carapace is very complete, but is almost lost
in the polygonal mosaic of tubercles.
The 4-lobulation of the antero-lateral borders is inconspicuous.
The postero-lateral borders are shorter than the antero-lateral, and
are markedly concave.
Front sharply bilobed, the lobes projecting far beyond the wall
pronounced orbital angle. Orbital margin with three closed sutures.
The exposed surfaces of the chelipeds are covered with the same
strong many-facetted tubercles as the carapace; but on the legs the
tubercles have sharper points and are many of them spiny, especially
those on the dactyli. |
The abdominal terga and the greater part of the sternum are
covered with a mosaic of smooth-worn tubercles: the under wall of the
carapace, as far as the epimeral suture, is granular.
Fingers short, blunt pointed, hardly hollew at tip.
Basal antennal joint prolonged between frout and orbit almost to
the inner angle of the orbit, very much as in Carpilius etc.
Colour in spirit light reddish brown, fingers black with white tips:
in life the colour is uniform purplish black.
In the Indian Museum are 10 specimens from the Persian Gulf,
Karachi, Pedro Shoal, Ceylon, Ganjam Coast, Mergui, and Malacca
(besides 14 specimens from Australia and Hongkong).
79. Actsa calculosa, (Kdw.) A. M. Edw.
Cancer calculosus, Milne Edwards, Hist. Nat. Crust. I. 378.
Actxa calculosa, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 276, pl.
xviii. figs, 3-3a: Haswell, Cat. Austral. Crost. p. 45: J. R. Henderson, Traus. Linn.
Soc., Zool. (2) V. 1893, p. 356.
Not very easily distinguishable from Actsxa granulata at first sight,
but the following difference is constant :—
(1) the carapace is shorter and broader, its length being only about
two-thirds its breadth :
(2) the tubercles of the carapace and chelipeds are much smoother
1898. ] A. Alcock — Carcinological Fauna of India. 153
and are hardly facetted, owing to the granules of which they are formed
being more intimately confluent; and on the posterior part of the cara-
pace the tubercles themselves are confluent, small, and little convex:
the tubercles of the legs are never spiny :
(3) the regions and Jobules of the carapace are much more dis-
tinctly delimited, and the 4-lobulation of the autero-lateral borders is
more distinct.
Colours in spirit much as in A. granulata.
In the Indian Museum are 8 specimens from the Persian Gulf,
Kardchi, and Mergui.
Bawareta, A. Milne Edwards.
Banareia, A. Milne Edwards, Ann. Soc. Hntomol. France (4) IX. 1869, p. 168,
and Nouv. Archiv. du Mus. IX. 1873, p. 198.
Strongly resembles Acta in all points but has the following
difference :—
(1) in the fore edge of the buccal cavern is, on either side, a deep
gap, not a mere suture or fissure such as is seen in some species of Actea:
(2) the fingers are compressed and extremely trenchant, resem-
bling shears.
80. Banareia armata, A. Milne Edwards.
Banareia armata, A. Milne Edwards, Ann. Soc. Ent. Fr. (4) IX. 1869, p. 168,
pl. viii; and Nouv. Archiv. du Mus. IX. 1873, p. 193: Ortmann, Zool. Jahrb., Syst.,
VII. 1893, p. 456: de Man, Jahrb. Hamb, Wiss. Anst, XIII. (Brachyuren des Hamb.
u. Paris Mus.) 1896, p. 75.
Outwardly, from the dorsal view, might almost be mistaken for
Actea ruppellu.
All exposed parts of the carapace and appendages, except the fingers
and lower outer surface of the hand are concealed by a dark shaggy
covering consisting of a dense under-fur with numerous tufts of long
hair.
The carapace is a little more than 3 as long as broad, and when
denuded, is seen to be divided into very numerous small lobules by
broad smooth grooves. The lobules are convex and closely covered
with pearly granules.
The front is formed of two small pointed lobes which do not break
beyond the common curve of the antero-lateral borders: the supra-
orbital border is fissured twice, and is separated from the lower border
of the orbit by a fissure: the antero-lateral borders, when denuded, are
seen to be divided into four granular lobes of unequal size, and a
ay it. 20
154 A. Alcock —Carcinological Fauna of India. [ No. 1,
(fifth) granular tubercle exists just behind the orbit: the postero-lateral
borders are very distinctly concave, and are much shorter than the
antero-lateral borders.
The outer surface of the wrist, when denuded, has much the same
sculpture as the carapace. An oblique patch of the lower outer surface
of the hand is smooth and polished and quite devoid of hair, as are the
fingers except the basal half of the upper edge of the dactylus. The
upper outer surface of the hand, when denuded, shows about six
longitudinal lines of granules, the three upper of which are a little
diffuse.
The fingers are compressed and trenchant, resembling shears:
the cutting edge of the dactylus is entire, but that of the thumb has
three incisiform teeth of unequal size at the base. In marked contrast
to all other parts, the fingers are smooth and polished.
In the Indian Museum are 3 specimens from the Andamans.
It appears to me to be quite consistent at present to separate this
species from Actxa while uniting Actwodes with that genus, for the good
reason that in this case there are no known transitional forms.
Datra, De Haan.
Daira, De Haan, Faun, Japon. Crust. p. 18.
Lagostoma, Milne Edwards, Hist. Nat. Crust. I. 387.
Daira, Dana, U. S. Expl. Exp. Crust. pt. I. p. 202. )
Daira, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 297; and Miss. Sci.
Mex., Crust. p. 248.
Carapace broad, strongly convex in both directions, the regions
well delimited and subdivided into very numerous convex lentil-like
lobules ; its antero-lateral borders strongly arched, crenulate ; its postero-
lateral borders concave, very short.
Front deflexed, two-lobed—the lobes being conspicuous and pro-
minent.
Orbital margin thickened and smoothly crenulate; a strong suture
line in the lower margin. LHyes on short thick sub-globular stalks.
Antennules folding obliquely owing to the large size of the basal
joint: interantennulary septum broad.
Basal antennal joint hardly touching the front, the next joint and
the very short flagellum wedged in the gap between the orbit and the
front.
Merus of the external mawillipeds with a wide and deep notch in the
anterior margin.
The chelipeds are unequal in both sexes; the fingers are blunt-
pointed and hollowed-out at tip.
1898. ] A. Alcock — Carcinological Fauna of India. 155
Upper edge of merus of chelipeds and legs crest-like and elegantly
serrated, upper edge of the succeeding joints of the legs with a crest
of stout sharp spines; but all this ornamentation is concealed by a
broad thick fringe of long coarse hair.
Abdomen of male with all seven joints distinct, but the 3rd—5th
segments are not movable on one another.
On either side of the endostome is an oblique septum defining the
efferent branchial canal, but this septum extends only about half-way
across the palate.
81. Daira perlata, (Herbst) De Haan.
Cancer perlatus, Herbst, Krabben, I, ii. 265, pl. xxi. fig, 122,
Cancer daira, Herbst, Krabben, ITI. ii. 6, pl. liii. fig, 2,
Cancer variolosus, Fabricius, Ent. Syst. Suppl., p. 338.
Daira perlata, De Haan, Faun. Japon. Crust. p. 18.
Lagostoma perlata, Milne Hdwards, Hist. Nat. Crust. I. 387.
Daira variolosa, Dana, U. 8. Expl. Exp. Crust. pt. I. p. 202, pl. x. figs. 4a-d.
Daira perlata, Stimpson, Proc. Ac. Nat. Sci. Phila. 1858, p. 32: A. Milne
Edwards, Nouv. Archiv. du Mus. 1.1865, p. 298; and IX. 1878, p. 196: Heller, Novara
Orust, p. 18: Miers, Cat. Crust. New Zealand, p. 18; and Phil. Trans. Vol. 168,
1879, p. 487: Richters, in Mobius Meeresf, Maurit. p. 147: Filhol, Crust. New Zea-
land, p. 374: R. I. Pocock, Ann. Mag. Nat. Hist. (6) V. 1890, p. 74: Ortmann, Zool.
Jahrb., Syst., VII. 1893-94, p. 474; and in Semon’s Forschungsr. (Jena. Denk. VIII.)
Crust. p. 52: Whitelegge, Mem. III. Austral. Mus., 1897, p. 131.
Carapace oval, strongly convex, subdivided into very numerous
polished lentil and pea-shaped lobules which have their surface finely
pitted. Antero-lateral borders 11 or 12-crenulate: postero-lateral
borders very short, concave.
Upper and outer surface of wrist lobulated almost like the cara-
pace; of hand and dactylus covered with coarse sharp tubercles, which
become almost spiniform on the upper edge: upper part of inner surface
of wrist and hand covered with a mosaic of flat markings that look like
ground-down tubercles: much the same sort of mosaic occurs on the
outer surface of the arm and legs, but on the dactyli and propodites of
the legs the tubercles are either spiny or acute, and at the distal end of
the carpus there is a spine,
Fingers short and thick, with blunt hollowed-out tips, those of the
larger cheliped have the cutting-edge toothed, those of the smaller
cheliped have a plain sharp cutting-edge.
Lower edge and surface of meropodites of legs much excavated in
their distal half to receive the bulging distal end of the carpus in
flexion.
Upper edge of legs fringed thickly with coarse long hair which
156 A. Alcock — Carcinological Fauna of India. LNo es
conceals their serrated and spiny sculpture. Two brushes of hair on
the under surface of the dactyli.
Colours in spirit—mottled shades of warm brown,
In the Indian Museum are 11 females and 3 small males from the
Laccadive reefs (besides 5 females and 1 small male from Mauritius and
2 females from Samoa).
Subfamily III. Cutoropina,
Alliance I. Xanthodioida.
XANTHODES, Dana.
Xanthodes, Dana, Proc, Acad, Nat. Sci. Philad. 1852, p. 75; and U. 8. Expl Exp.
Crust. pt. I. p. 175.
Xanthodes, A. Milne Edwards, Ann. Sci. Nat. Zool. (4) XX. 1863, p. 227; Nouv.
Archiv. du Mus. IX. 1873, p. 200; and Miss. Sci. Mex., Crust. p. 259,
Xanthodes, Miers, Challenger Brachyura, p. 127.
[Type Xanthodes lamarckit (Kidw.) ]
Carapace thick but somewhat depressed, moderately broad, some-
what hexagonal, the regions delimited and to a certain extent areolated
in the anterior two-thirds.
Fronto-orbital border considerably more than half the greatest
width of the carapace in extent.
Front broad (contained about 33 times in the greatest width of the
carapace) bilobed.
Antero-lateral border cut into four lobes or teeth. Orbital margin
with the three grooves either fairly distinct, or quite indistinct.
Basal antennal joint broad and very short; the flagellum, which is
rather longer than the orbit, lodged in the orbital hiatus.
Anterior edge of merus of external maxillipeds almost transverse.
Chelipeds either equal or unequal in both sexes; the arm in repose
is nearly or quite hidden beneath the carapace; fingers pointed, not
hollowed at tip.
Legs stoutish, more or less hairy and granular or spiny along the
upper border.
Abdomen of the male five-jointed.
Of the two Indian species included in this genus, one (Xanthodes
lamarckit) has a certain resemblance to Lioxantho punctatus, from which
it is easily distinguished by the breadth of the frouto-orbital margin ; the
other (Xanthodes notatus) has a strong resemblance to Phymodius
sculptus, from which it is at once distinguished by the sharp-pointed
fingers, not hollow at tip.
1898. | A. Alcock — Carcinological Fauna of India. 157
Key to the Indian species of Xanthodes.
I, Chelipeds eqnal, hands and wrists closely granular,
outer surface of hand with three deep parallel longi-
PmdiMad HALLOWS lowaes occ cr bee « geesebuwedt > odeessdesiaccisera. ¢ A lamarebtis
II. Chelipeds markedly unequal, hands and wrists, at any
rate of the smaller cheliped, studded with sharp spine-
PRCT aan 7, oe Ons eaca casas cade o-abanss dang ooce vaipeeicas oe «6X. NoOtatus.
€2. Xanthodes lamarckit (Hdw.).
Xantho lamarckii, Milne Edwards, Hist. Nat. Crust. I. 391: A. Milne Edwards,
in Maillard’s Vile Réunion, Annexe F, p. 4: Heller, Novara Crust. p. 10: Ortmann,
Zool. Jahrb. Syst. VII. 1893-94, pp. 444, 448.
Xanthodes lamarckii, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1878, p. 200,
pl. vii. fig. 3: Hilgendorf, MB. Ak. Berl. 1878, p. 789: Miers, Zool. H. M. S. Alert,
pp. 517, 529: F. Muller, Verh. Ges. Basel, VIII. 1886, p. 474: de Man, Archiv.
fiir Naturges. LIII. 1887, i. p. 263 ; and in Weber’s Zool. Ergebn. Niederl. Ost-Ind.
II. 1892, p. 278; and Zool. Jahrb. Syst. VIII. 1894-95, p. 513: Whitelegge, Mem.
Austral. Mus. IIT. 1897, p. 130.
Xanthodes granosomanus, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 75; and
U.S. Expl. Exp. Crust. pt. I. p. 175, pl. viii. figs. 10a~c.
Xantho granosomanus, Heller, Novara Crust. p. 11.
The anterior and antero-lateral parts of the carapace are to a
variable extent granular, the posterior part is generally quite smooth.
Fronto-orbital region marked off by a sinuous groove, gastric region
well delimited and fairly distinctly divided into 3 sub-regions, branchio-
hepatic regions incompletely traversed by 2 grooves proceeding from
the 2nd and 8rd intervals of the antero-lateral border: no other grooves
on the carapace.
Outer angle of front not very pronounced, separated from the
supra-orbital margin by a faintish groove. The grooves of the orbital
margiu are almost indistinguishable.
Antero-lateral border divided into four broadish granular lobes,
the last two of which are more acuminate (but bluntly) than the others.
Chelipeds equal in both sexes, stout, rather short (less than twice
the length of the carapace) ; arm hidden beneath the carapace in repose,
its anterior and posterior edges hairy, the upper part of its posterior
surface granular: upper and outer surfaces of wrists and hands as
closely as possible covered with pearly granules, the wrist also has a
few indistinct dimples, and the outer surface of the hand is deeply
scored by three parallel longitudinal furrows: fingers rather long,
pointed.
Upper edge of meropodites of legs very finely serrulate : surfaces of
next three joints closely granular in the vicinity of the upper (anterior)
158 A. Alcock — Carcinological Fauna of India. | ore
edge: some longish hairs scattered along the upper border of the last
four, and also along the ventral (posterior) borders of the last two joints.
Colours in spirit: yellowish white, fingers blackish-brown. In well
preserved spirit specimens the legs are banded with bluish green, and
large confluent bluish green blotches occur on the carapace.
In the Indian Museum are 15 specimens, from the Andamans,
Madras coast and Ceylon, (besides 6 from parts outside India).
83. Xanthodes notatus, Dana.
Xanthodes notatus, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p..76, and U. S. Expl.
Exp. Crust. I. p. 178, pl. viii. figs. 12a-b: A. Milne Edwards, Nouv. Archiv. du Mus.
1X. 1878, p. 201: Haswell, Cat. Austral. Crust. p. 49: de Man, Archiv. f. Naturges.
LIII. 1887, i. p. 264.
Xantho notatus, Heller, Novara Crust. p. 10,
Surface of carapace, except for the characteristic deep cut areola-
tion, quite smooth to the naked eye. Deep well-cut grooves separate
the fronto-orbital region, define the gastric region, and subdivide the
branchio-hepatic regions into 3 or 4 lobules; and the gastric region is
divided into 3 sub-regions by fine but well-cut lines.
Front bilobed, the outer angle of each lobe being well defined and
separated from the supra-orbital margin by a notch and groove. The
3 grooves of the orbital border are distinct.
Antero-lateral border cut into 4 teeth, of which the last two are
procurved and spine-like.
Chelipeds markedly unequal, the larger one more than twice the
length of the carapace: the arm in both is not quite concealed by the
carapace, is devoid of hair, and has the distal end of the upper border
spinate: in the smaller cheliped, the upper and outer surfaces of the
wrist and the upper and a large part of the outer surface of the hand
are studded with sharp spine-like tubercles; but in the larger cheliped
the tubercles are larger, less numerous, and are low and worn, not
spine-like: the fingers are pointed, not hollowed at tip, and in the
smaller cheliped are fluted, the ridges of the dactylus having a few
sharpish tubercles at the basal end.
The upper edge of the meropodites of the legs is spiny, with a few
long fine bristles: the upper borders of the next two joints have each
two rows of spines and a good many long bristles, the carpus having
also a third row of sharp granules: the dactylus is granular and bristly,
and the lower edge also of the propodite has some bristles.
Colours of well-preserved spirit specimens: purplish brown, the
purplish tinge very distinct on the chelipeds, the last 3 joints of the
legs are greenish.
1898. ] A. Alcock — Carcinological Fauna of India. 159
In the Indian Museum are 17 specimens, from the Andamans, Palk
Str. and Ceylon.
This species has a strong likeness to Phymodius sculptus.
Alliance II. Chlorodioida.
Chlorodius. Chlorodopsis.
Phymodias. Cyclodius.
Cutoropius, A, Milne Edwards.
Chlorodius, (part) Milne Edwards, Hist. Nat. Crust. I. 399.
Chlorodius, (part) Dana, Silliman’s Amer. Journ. Sci. and Arts (2) XII. 1851,
p- 126, and U.S. Expl. Exp. Crust. pt. I. p. 204.
Chlorodius, A. Milne Edwards, Ann. Sci. Nat. Zool. (4) XX. 18638, p. 283 ; Nouv.
Archiv. du Mus. 1X. 1873, p. 212; Miss. Sci. Mex., Crust. p. 265.
[Type Chlorodius niger (Forsk.) |
Carapace depressed, flat, hexagonal, the regions faintly or not at
all demarcated, the surface smooth and almost unbroken, except some-
times on the branchio-hepatic region, near the antero-lateral border,
where there may be some broad transverse wrinkles.
Fronto-orbital border more than three-fourths the greatest breadth
of the carapace. Front almost straight, faintly emarginate in the
middle line, extremely broad (between a third and half the greatest
breadth of the carapace), its outer angles separated from the supra-
orbital margin by a groove.
Autero-lateral borders cut into four lobes or teeth, the first being
in very close approximation to the angle of the orbit, Postero-lateral
borders rather longer than the antero-lateral.
Orbit with two suture lines above, and one at the outer angle:
eyes on short thick stalks.
Basal antennal joint large, extending upwards and outwards into
the gap between the front and the orbit; the flagellum situated in the
erevice-like orbital hiatus.
Merus of the external maxillipeds with the anterior border almost
transverse.
Chelipeds unequal, long, more than twice the length of the carapace,
half or more of the arm projecting beyond the edge of the carapace ;
fingers large, broadened and deeply hollowed at tip (horse-shoe shaped).
Legs never spiny, though the upper edge of the meropodites may
have a few spinules distally, and that of the following joints is sharply
granular.
Abdomen of the male consisting of 5 joints, the 3rd-5th somites
being fused.
160 A. Alcock — Carcinological Fauna of India. [ No. 1,
Key to the Indian species of Chlorodius.
I. Carapace with the regions faintly marked, transversely
wrinkled near the antero-lateral borders ; four distinct
teeth (exclusive of the orbital angle) on the antero-
lateral, border ....j:.:. t¢iascowcdi seis dal veniaehi deeb @ernPer Mileeuclece sont) ame amm
II. Carapace with a perfectly smooth unbroken surface;
first lobe of antero-lateral margin almost obsolete, the
third the most distinct Of all ......ccc.ssscnsaqssccesroreesvcsece On LERDEGSIIUINME
84. Chlorodius niger (Forsk.) Riippell, A. M. Edw.
Cancer niger, Forskal, Descr. Anim. p. 89.
Chlorodius niger, Ruppell, 24 Krabben roth. Meer. p. 20, pl. iv. fig. 7 and pl. vi.
fig. 14: Milne Edwards, Hist. Nat. Crust. I. 401: Dana, U. S. Expl. Exp. Crust.
pt. I. p. 216, pl. xii. figs. 5a-c: Stimpson, Proc. Ac. Nat. Sci. Phila., 1858, p. 33:
Heller, SB. Ak. Wien, XLIII. 1861, i. p. 335, and Novara Crust. p. 18: A. Milne
Edwards, Nouv. Archiv. du Mus. IV. 1868, p. 71, and 1X. 1873, p. 214: Kossmann,
Reise roth. Meer. Crust. p. 34: Miers, Ann. Mag. Nat. Hist. (5) V. 1880, p. 234;
and P. Z. 8. 1884, pp. 10,11; and Zool. H. M.S. Alert, pp. 183, 215, 517, 531:
de Man, Notes Leyden Mus. II. 1880, p. 174, III. 1881, p. 98, and Archiv. fiir
Naturges. LIII. 1887, i. p. 279, and Journ. Linn. Soc., Zool., XXII. 1887-88, p. 32 ;
and Zool. Jahrb. Syst. VIII. 1894-95, p. 519: Richters in Mébius Meeresf. Maurit.
p. 147: Haswell, Cat. Austral. Crust. p. 62: J. R. Henderson, Trans. Linn. Soc.,
Zool., (2) V. 1898, p. 861: Ortmann, Zool. Jahrb. Syst. VII. 1893-94, p. 465, and
in Semon’s Forschungsr. (Jena. Denk. VIII.) Crust. p. 61.
Chlorodius hirtipes, White, P. Z. 8S. 1848, p. 226; Ann. Mag. Nat. Hist. (2) IT.
1848, p. 286 ; and Adams and White, Samarang Crust. p. 49, pl. xi. fig, 4.
Chlorodius cytherea, Dana, Proc. Ac. Nat. Sci. Phila. 1852, p. 79, and U. S.
Expl. Exp. Crust. pt. I. p. 213, pl. xii. figs. 2a-c: Stimpson, Proc. Ac. Nat. Sci.
Phila. 1858, p. 33.
Chlorodius nebulosus, Dana, Proc. Ac. Nat. Sci. Phila. 1852, p. 80, and U. 8S.
Expl. Exp. Crust. pt. I. p, 214, pl. xii. fig. 3.
Chlorodius depressus, Heller, Abh. zool.-bot. Ges. Wien, 1861, p.11; and SB. Ak.
Wien, XLIII. 1861, p. 338: Hilgendorf in v. d. Decken’s Reisen Ost-Afr. III. i. p. 74.
Carapace hexagonal, depressed: gastric region delimited by faint
grooves and subdivided into from 3 to 5 indistinct areole by still
fainter grooves; anterior part of branchio-hepatic region, just inside
the antero-lateral margin, with one or two low broad transverse wriuk-
les : the surface of the carapace quite smooth (non-granular),
First lobe of the antero-lateral border small, almost confluent
with the rounded external orbital angle; last two lobes usually ending
in procurved spine-like points. |
Chelipeds, in both sexes, twice or more the length of the carapace,
usually quite smooth to the naked eye; a tubercle or spine on the
anterior edge, and a little crenulation (not always present) on the
posterior edge of the arm; inner angle of wrist strongly pronounced ;
fingers stout, a good deal arched, markedly spoon-like at tip.
1898, ] A. Alcock — Carcinological Fauna of India. 161
Legs with a good deal of hair, and long fine bristles interspersed,
on the dorsal aspect of the last three joints.
Colours in spirit: yellowish brown to bluish or purplish brown,
sometimes mottled ; fingers black.
In the Indian Museum are 64 specimens, frem the Andamans,
Nicobars, Mergui, and Mekran coast, (besides 52 from other parts of
the Indo-Pacific).
85. Chlorodius levissimus, Nana,
Chlorodius levissimus, Dana, Proc. Ac. Nat. Sci. Phila. 1852, p. 80, and U. S.
Expl. Exp. Crust. pt. 1. p. 215, pl. xii. figs. 4a-g.
Carapace hexagonal, a little tumid, its surface perfectly smooth
without trace of regions or areole.
First lobe of antero-lateral border almost obsolete, last tooth very
small, the third tooth much the largest and most prominent.
Anterior edge of arm without a spine.
Last three joints of the legs with a few scattered long fine
bristles—no hair.
Fingers very strongly arched, a character which—as Dana has
noticed—at once distinguishes this little species from the young of
Chlorodius niger, which it otherwise closely resembles.
Colours in spirit, white, fingers brown.
In the Indian Museum are 9 specimens, from the Andamans and
Ceylon, (aud 1 from Mauritius).
Paymopius, A. Milne Edwards.
Chlorodius, (part) Milne Edwards, Hist. Nat. Crust. I. 399: Dana. U.S. Expl
Exp. Crust. pt. I. p. 204. |
Phymodius, A. Milne Edwards, Ann. Sci. Nat., Zool. (4) XX. 1863, p. 283; and
Nouv. Archiv. du Mus. [X. 1873, p. 217; and Miss. Sci. Mex. Crust. p. 266.
Phymodius, Miers, Challenger Brachyura, p. 139.
Carapace moderately flat, hexagonal, all the regions well delimited,
and broken up into numerous convex areole which have a smooth bare
surface.
Fronto-orbital border not quite two-thirds the greatest breadth of
the carapace. Front distinctly bilobed, with the outer angle of each
lobe forming a distinct little lobule; its breadth is about a third the
greatest breadth of the carapace. Orbital margin with 2 grooves above
and one at the outer angle: eyes on short thick stalks.
Antero-lateral borders cut into four lobes or teeth: postero-lateral
border nearly equal in length to the antero-lateral.
Pir. 21
162 A. Alcock — Carcinological Fauna of India. [No. 1,
Basal antennal joint large, extending outwards and upwards into
the gap between the front and the orbit; the flagellum situated in the
orbital hiatus.
Anterior edge of merus of external maxillipeds almost transverse.
Chelipeds unequal, twice or more the length of the carapace, about
half the arm projecting beyond the edge of the carapace: fingers large,
strongly arched, broadened and deeply hollowed at tip.
Legs with the meropodite carpopodite and propodite sharply spinous
along their upper border.
Abdomen of the male five-jointed.
Phymodius is distinguished from Chlorodius by the extensive and
distinct areolation of the carapace, by the narrower front, and by the
spiny armature of the dorsal border of the legs.
Key to the Indian species of Phymodius.
I. Lobules of carapace smooth but dull; chelipeds rough
with nodules or tubercles ; legs with scattered hairs that
do not hide the spines :—
i. Sculpture of carapace sharp cut; chelipeds with
postule-like tubercles extending as far as fingers. P. ungulatus.
ii. Sculpture of carapace worn; chelipeds with irre-
gular nodules that do not are reach more
than halfway along the hand . etme wees P. monticulosus.
II. Lobules of carapace smooth and enauee as Wide: are the
chelipeds; legs with a stiff fringe of hair along the
anterior border concealing the spines there ......... -.... P. sculptus.
86. Phymodius ungulatus (Hdw.) A. M. Edw.
Chlorodius wngulatus, Milne Edwards, Hist. Nat. Crust. I. 400, pl. xvi. figs. 6-8 :
Dana, U. S. Expl. Exp. Crust. pt. I. p. 205, pl. xi. figs. 8a-b: Hess, Archiv.
fiir Naturges. XXXI. 1865, pt. i. pp. 185, 171: Streets, Bull. U. S. Nat.. Mus. VII.
1877, p. 105.
Phymodius ungulatus, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1878,
p. 218: Hilgendorf, MB. Ak. Berl. 1878, p. 790: Kossmann, Reise roth. Meer.
Crust. p. 34: Haswell, Cat. Austral. Crust. p. 59: Miers, Challenger Brachyura,
p. 139: Cano, Boll. Soc. Nat. Napol. III. 1889, p. 201: J. R. Henderson, Trans.
Linn, Soc., Zool., (2) V. 1893, p. 362: Ortmann, Zool. Jahrb. Syst. VII. 1893-94,
p. 464: de Man, Zool. Jahrb. Syst. VIII. 1894-95, p. 524,
? Xantho de Haanii, Krauss, Sudafr. Crust. p. 29, pl. i. fig. 2: Heller, SB. Ak.
Wien, XLIII. 1861, p. 337, and Novara Crust. p. 19.
Chlorodius areolatus, Adams and White, Samarang Crust. p. 41, pl. xi. fig. 3.
The regions and numerous sub-regions of the carapace are all
convex and sharply defined by clean-cut furrows; their surface is
smooth but dull, owing to close microscopic granulation.
1898. ] A. Alcock — Carcinological Fauna of India. 163
Front bilobed, the outer angle in each lobe forming a distinct little
lobule.
The four teeth of the antero-lateral border are sharply conical.
Chelipeds unequal, but not greatly so, very finely granular; arm
with the anterior border rather strongly serrated, and with numerous
pustule-like tubercles along the posterior border; upper and outer
surface of wrist, aud upper as well as a small part of inner and a larger
part of outer surface of hand, covered with well-spaced pustule-
like tubercles, those on the outer surface of the hand being in
longitudinal series; inner angle of wrist strongly pronounced, with
bifid tip. )
Legs with finely granular surface, sharply granular on the dorsal
aspect, where there are some long scattered hairs: upper edge of
meropodites with 1 row of spinules, upper border of carpopodites with
3 rows, of propodites with 2 rows.
Colours in spirit, yellowish brown, or greenish; fingers black.
In the Indian Museum are 12 specimens, from the Andamans and
Ceylon, (as well as 19, from Mauritius and Samoa).
87. Phymodius monticulosus (Dana), A. M. Edw.
Chlorodius monticulosus, Dana, Proc. Ac. Nat. Sci. Phila. 1852, p. 79; and U.S.
Expl. Exp. Crust. pt. I. p. 206, pl. xi. figs. 9a-f: Stimpson, Proc. Ac. Nat. Sci. Phila.
1858, p. 34.
Chlorodius obscurus, Lucas in Jacquinot’s Voyage Astrolabe, Zool. Vol. IIT.
Crust. p. 26, pl. iii. fig. 4.
Phymodius monticulosus, A. Milne Edwards, Nouv. Archiv. du Mus. IV. 1868.
p. 71 (name only): Richters in Mébius, Meeresf. Maurit. p. 148: Miers, Challenger
Brachyura, p. 139: Muller, Verh. Ges. Basel, VIII. 1886, p. 474: Cano, Boll. Soc.
Nat. Napoli, III. 1889, p. 201: J. R. Henderson, Trans. Linn. Soc., Zool., (2) V.
1893, p. 363: de Man, Zool Jahrb., Syst. VIII. 1894-95, p. 524: T. Whitelegge, Mem.
Austral. Mus. III. 1897, p. 136.
Phymodius obscurus, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873, p. 220:
(?) de Man, Notes Leyden Mus. II. 1880, p. 174: Richters in Mobius, Meeresf.
Maurit. p. 148.
Closely resembles P. ungulatus, but is at once distinguished by the
more convex arch of the front and antero-lateral borders, by the greater
dorsal convexity of the carapace, by the ‘‘ worn” look of the sculpture
of the carapace, and by the much less rough hands.
Carapace thick, and distinctly convex in its anterior two-thirds ;
the regions and subregions are all distinct and convex, but the depres-
sions that separate them are broad and not.sharp cut, and this gives the
sculpture a worn or moulded appearance.
Front bilobed, the outer angle of each lobe well defined.
164, A. Aleock — Carcinological Fauna of India. [No. 1,
The four teeth of the antero-lateral margin are blunt and rounded
the first two being very much worn.
Chelipeds unequal—more so than in P. wngulatus: arm with two
or three coarse denticles on the anterior border, the posterior border
being rugose; upper and outer surface of wrist nodular; a certain part
of the upper, as well as of the inner and (more so) of the outer surfaces
of the hand nodular, but except in very young specimens, the nodules do
not extend beyond, and often not so far as, halfway along the hand, so
that the greater part of the hand is often smooth.
Legs as in P. ungulatus, but the spinules are coarser and blunter.
Colours in spirit—dark chestnut brown, sometimes mottled with
grey; fingers blackish brown.
In the Indian Museum is a single specimen from the Nicobars (in
addition to 21 from Australia, the South Seas, and Mauritius).
88. Phymodius sculptus, (A. Milne Edwards).
Chlorodius sculptus, A. Milne Edwards, Nouv. Archiv. da Mus. IX. 1873, p. 217,
pl. viii. fig. 4: de Man, Notes Leyden Mus. III. 1881, p. 98; Archiv. fiir Naturges.
LIII. 1887, i. p. 279; and Journ. Linn. Soc., Zool., XXII. 1887-88, p. 82: Ortmann,
Zool, Jahrb. Syst. VII. 1893-94, p, 466.
Regions of the carapace well defined and subdivided, by broad
and deepish grooves, into smooth, polished, convex but flat-topped
lobules : those of the branchio-hepatic regions are disposed transversely :
the antero-lateral sub-regions of the gastric area are not longitudinally
subdivided.
Front bilobed, the outer gente of each lobe distinct but not very
prominent.
The antero-lateral border is cut into four smooth lobes and is
rather shorter than the postero-lateral.
The chelipeds are unequal: the arm has several sharp teeth on the
anterior border and several pearly tubercles on the distal end of the
posterior border, and the inner angle of the wrist is calere but the
surface of the chelipeds is smooth and polished.
The most characteristic feature of the legs is the dense stiff fringe
of long greenish-yellow bristles that clothes the anterior border of the
last four joints, concealing the sharp spines with which these borders
are armed.
Colours in spirit, body and legs green with brownish points, cheli-
peds brownish, fingers black.
In the Indian Museum are 10 specimens, from the Andamans,
Mergui and Ceylon.
1898. | A. Alcock — Carcinological Fauna of India. 165
Cuioropopsis A. Milne Edwards.
Chlorodopsis, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1878, p. 227.
? Pilodius, (part) Dana, U. 8. Expl. Exp. Crust. pt. I. p. 217.
Carapace flat, more or less hexagonal, the regions well delimited
and well areolated, the areole being granular or hairy, or both.
Fronto-orbital border about two-thirds the greatest breadth of the
carapace. Front bilobed, the outer angle of each lobe usually well
defined and forming a distinct little lobule; its breadth is about a third
the greatest breadth of the carapace.
Antero-lateral border almost always cut into four teeth. Postero-
lateral border commonly a little longer than the antero-lateral.
Orbital border with the three grooves or notches very distinct.
Basal antennal joint large, extending up between the front and the
orbit, the outer angle being prolonged into the orbital hiatus.
Anterior edge of merus of external maxillipeds almost transverse.
Chelipeds either unequal or subequal, their length being generally
under twice the length of the carapace; the arm short and not project-
ing very much beyond the carapace.
Fingers strong, arched, broadened and hollowed at tip, but not so
hoof-like as Chlorodius.
Legs dorsally almost always hairy and spinous.
Abdomen of male 5-jointed.
Chlorodopsis is distinguished from Chlorodius and Phymodius, (1) by
the prolongation into the orbital hiatus of the outer angle of the basal
joint of the antenna, and (2) by the granular and hairy or furry carapace.
Key to the Indian species of Chlorodopsis.
I. The entire carapace cut into strongly-convex, isolated
areole, the surface of which is uniformly covered with
pearly granules: the deep smooth grooves between the
areolz, and the spaces between the granules, covered
with a dense, dark, extremely short fur sseerseceevases, Os Areotata.
II. Only the anterior % to $ of the carapace areolated : three
or four of the lobules just inside the antero lateral
border either bear spines or are themselves spine-like :—
i. Sculpture of carapace and legs almost concealed
by bristles and long hairs :—
1. Posterior fourth of carapace slightly con-
cave; chelipeds rather slender, little unequal,
the black colouration of the thumb hardly
involves the hand at all :—
a, Antero-lateral margin armed with four
large spines (not including the orbital
ANIC) .......c0r00 sescvereccososersscsccesses CO, pilumnoides.
166 A. Alcock — Carcinological Fauna of Indva. [No. 1,
b. Antero-lateral margin divided into
four blunt spinuliferous lobes ......... C. nigrocrinita.
2. Posterior fourth of carapace flat; the
antero-lateral margin consists of four lobes
each capped by several spinules: chelipeds
markedly unequal, the black colouration of
the thumb involves the greater part of the
lower surface of the hand ... ..........se0.... O. melanochira,
ii. Carapace with a few scattered hairs which do not
in the least conceal its sculpture. The groove
that cuts off the fronto-orbital margin from the
rest of the carapace is very distinct :—
1, All four spines of the antero-lateral margin
equal, the 2nd and 3rd commonly with an
accessory spinule near the tip.........eecee0e.. OC. wood-masoni.
2. First spine of the antero-lateral margin
small or obsolescent ; last three spines large,
ClaweliKe ‘easel oi dis dedeScaves soclobe Sockeunaddesene) NOME EnO IEEE
89. Chlorodopsis areolata, (Kdw.) A. M. Edw.
Chlorodius areolatus, Milne Edwards, Hist. Nat. Crust. I. 400: Hess, Archiv.
fir Naturges. XXXI. 1865, pp. 135, 171.
Chlorodius perlatus, Macleay, Ill. Zool. S. Afr., Annulosa, p. 59: Krauss, Sudafr
Crust. p. 31.
Chlorodopsis areolata, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873,
p. 231, pl. viii. fig. 8: Hilgendorf, MB. Ak. Berl. 1878, p. 790: Richters in Mobius.
Meeresf. Maurit. p. 148: Haswell, Cat. Austral. Crust. p. 54: Miers, Zool. H. M. S,
Alert, pp. 517, 532: F. Muller. Verh. Ges. Basel, VIII. 1886, p. 474: de Man, Notes
Leyden Mus. XII. 1890, p. 54: Ortmann, Zool. Jahrb. VII. 1893-94, p. 470.
Carapace flat, but thick, as completely lobulated as any Actza ; the
lobules strongly convex, isolated by broad deep smooth channels, their
convexities.as closely as possible covered with pearly granules, the
dividing channels lined by an extremely short dense dark fur, which
also extends between but does not cover the granules of the lobules.
The front is deeply and broadly cut into two granular lobes, the
outer angle of each of which forms a separate lobule. The three
fissures of the orbital margin are so deep as to give a lobed appearance.
The antero-lateral border is divided by broad notches into four
rounded granular lobes.
Chelipeds unequal, the longer one about twice the length of the
carapace; the upper part of the outer surface of the arm, the nodular
or wrinkled surface of the wrist, and the upper and outer surface of
the hand are all closely covered with pearly granules, which are largest
on the hand: fingers strongly arched, smooth except for some grooving
and granulation at base of dactylus.
1898.] A. Alcock — Carcinological Fauna of India. 167
The exposed surface of the legs is as closely as possible covered
with a dense spongy fur from which the tops of numerous conical or
subspinous granules peep out: the dorsal edge of the legs is also clothed
with a thick shaggy fringe of hair, as also the ventrad edge of the last
two joints.
Colours in spirit yellowish brown to blackish brown; fingers black,
the colouration extending along the lower border and on to both surfaces
of the hand.
In the Indian Museum are 19 specimens, from the Andamans,
Nicobars and Ceylon (in addition to 12 specimens from the South Seas
aud Mauritius).
90. Chlorodopsis pilumnoides, (White).
Chlorodius pilumnoides, White, P. Z. S. 1847, p. 226; Ann. Mag. Nat. Hist.
(2) II. 1848, p. 286; Adams and White, Samarang Crust. p. 41, pl. ix. fig. 3.
? ? Pilodius pilumnoides, Dana, U.S. Expl. Exp. Crust. pt. I. p. 221, pl. xii.
fig. 10a-—c.
Chlorodopsis pilumnoides, de Man, Journ, Linn. Soc., Zool., XXII. 1887-88, p. 34,
Archiv. fiir Naturges. LIII. 1887, i. p. 281: Cano, Boll. Soc. Nat. Napol. III. 1889,
p. 204: Ortmann, Zool. Jahrb. VII. 1893-94, p. 470.
Carapace, chelipeds and legs granular, beneath a copious covering
of short black bristles among which are scattered numerous long white
club-shaped hairs.
Carapace flat, its regions, in the anterior two-thirds, plainly marked
and subdivided by broadish shallow furrows, but not convex; its
posterior third flat, or even a little concave, between two raised trans-
verse beaded lines.
Front cut rather deeply into two granular or denticulate lobes, the
outer angle of each of which forms a little lobule. The three fissures
of the granular orbital margin are distinct.
On the antero-lateral margin are four red-tipped claw-like spines
not including the orbital angle, the middle two, at least, of which have
a pair of spinelets at base: on the carapace just inside either antero-
lateral margin is a scattered group of 5 or 6 similar, but rather smaller,
spines.
Chelipeds subequal, rather slender, not longer than the legs (less
than twice the length of the carapace) ; both edges of the upper surface
of the arm spinulate ; numerous spines on the wrist, the one (or two)
at the inner angle the largest; rows of spines along upper surface, rows
of sharp granules along lower part of outer surface, of hand; fingers
strongly fluted, the ridges being sharply and elegantly serrate or
spinate.
168 A. Alcock— Carcinological Fauna of India. [No. 1,
In the legs, all the edges of the meropodites are more or less spinate
and the carpopodites and propodites are dorsally more or less spinulate.
Colours in spirit: yellowish, or mottled green; legs yellowish with
purplish-brown cross-bands, or light green with dark green cross-bands ;
fingers black, the colouration not extending along the hand.
In the Indian Museum are 7 specimens from the Andamans and 1
from Mergui.
91. ? Chlorodopsis nigrocrinita, (Stimpson).
? Pilodius nigrocrinitus, Stimpson, Proc. Acad. Nat. Sci. Philad. 1858, p. 34.
Differs from 0. pilumnoides in having the antero-lateral margin
cut into 4 blunt lobes which when denuded and examined under a lens
are spinuliferous: only the distal end of the upper edge of the arm is
spinulate.
Four specimens from the Andamans are in the Indian Museum.
It is at once distinguished from C. melanochira, to which it also
bears a strong resemblance, by the altogether different form of the
chelipeds and fingers. The chelipeds, like those of C. pilumnoides, are
slender and of equal size, and the black colouration of tlie fingers does
not extend on to the hand.
i 92. Chlorodopsis melanochira, A. M. Edw.
Chlorodopsis melanochira, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873,
p. 228, pl. viii. fig. 5: Haswell, Cat. Austral. Crust. p. 55: de Man, Archiv. fiir
Naturges. LIII. 1887. i. p. 281, and in Weber’s Zool. Ergebn. Nieder]. Ost-Ind., II.
1892, p. 278; and Zool. Jahrb. Syst. VIII. 1894-95, p. 520: Ortmann, Zool. Jahrb,
Syst. VII. 1893-94, p. 471.
Carapace, chelipeds and legs covered with short black bristles and
long yellow hairs, the yellow hairs being sparse on tle carapace but
exceedingly long and numerous on the legs, and the bristles being
embedded each in a curious little white ball of felt.
On the denuded carapace the regions are all well defined and well
areolated by well-cut smooth grooves, the convexities of the areolee
being granular: the posterior third or fourth of the carapace forms a
flat granular surface.
Front cut into two elegantly denticulated lobes, the outer angle of
each of which forms an independent lobule. The three fissures of the
finely denticulate orbital margin are distinct.
The antero-lateral margin is divided into four lobes, each of which
is crowned with several spinules: two or three-of the lobules of the
carapace just inside either antero-lateral margin are capped with
similar spinules.
1898.] A. Alcock — Carcinological Fauna of India. 169
The chelipeds are markedly unequal, the larger one being less than
twice the length of the carapace. The anterior and posterior edges of
the arm are granular; the wrist has the upper and outer surfaces
studded with granules and conical spine-like tubercles; similar tuber-
cles and pearly granules stud the upper and more or less of the outer
surface of the hand; the finger has a few spinule-like tubercles at base.
The legs have the upper border of the meropodite, carpus, and
propodite denticulate.
Colours in spirit; brownish yellow or mottled green, the mottling
on the legs forming indistinct cross-bands ; fingers black, the coloura-
tion involving the greater part of the lower border and both surfaces of
the lower outer corner of the hand.
In the Indian Museum are 35 specimens from the Andamans.
This species is at once distinguished from C. pilumnoides, (1) by
the smaller size, (2) by the better defined areolation of the carapace, (3)
by the cap of spinelets—instead of a large claw-like spine—on each of
the 4 lobes of the antero-lateral margin, (4) by the marked inequality
of the chelipeds, and (5) by the black colouration of the thumb extending
far back along the hand.
93. Chlorodopsis spinipes (Heller) A. M. Edw.
Pilodius spinipes, Heller, Abh. zool.-bot. Ges. Wien, 1861, p. 11, and SB. Ak.
Wien, XLIII. 1861, i. p. 340, pl. ii. fic. 22.
Chlorodopsis spinipes, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873,
p. 230, pl. viii. fig. 6: de Man, Notes Leyden Mus. III. 1881, p. 98; Archiv. fiir
Naturges. LIII. 1887, i. p. 282; and in Weber‘s Zool. Ergebn. Nieder]. Ost-Ind.
II. 1892, p. 278: J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 361:
Ortmann, Zool. Jahrb. VII. 1893-94, p. 471.
Carapace and chelipeds with a few scattered hairs, legs with
numerous long stiff brown and yellow hairs that a good deal conceal
the sculpture.
The regions and subregions of the carapace in its anterior ¢ are
most remarkably well defined by broad smooth deep-cut grooves, and
are coarsely and unevenly granular.
The front is cut into two lobes which have their free edge entire or
slightly crenulate, and their outer angle isolated and spine-like. The
orbital margin is sharp-cut and almost smooth: the 3 fissures are distinct.
The antero-lateral margin is cut into four teeth, of which the first
is always small and often obsolescent, while the other three are large,
procurved and claw-like. The three or four lobules of the carapace
just inside and parallel with the antero-lateral border have the form of
salient conical tubercles.
J. m1. 22
170 A. Alcock —Carcinological Fauna of India. [No. 1,
The chelipeds are unequal, the larger one being not quite twice the
length of the carapace: the arm has 2 or 3 spines at the distal end of
the anterior and posterior borders; the upper and outer surfaces of the
wrist and hand are covered with sharp spine-like tubercles which
become blunt and pearl-like in the lower part of the hand, and one or
two of the spines at the inner angle of the wrist are enlarged; fingers
with some sharp tubercles at base.
Legs with numerous long sharp spines (which are a good deal
concealed by long stiff hairs) along the upper border,—a single series
on the meropodites, 2 or 3 series on the carpopodites and propodites.
Colours in spirit: yellowish or greenish brown, somewhat mottled
on the carapace and somewhat banded on the legs; fingers black, the
colouration not extending to the hand.
In the Indian Museum are 20 specimens, from the Andamans and
Mergui.
This species is suspiciously like the Pilodius pugil of Dana.
94. Ohlo rodopsis wood-masont, 0. sp.
Carapace with a few rather long scattered hairs, legs with similar
but more numerous hairs, not in any way concealing the sculpture,
chelipeds almost free from hairs.
The carapace is thick, and has the regions and subregions well
defined, in its anterior ?, by broad smooth grooves, and coarsely and —
unevenly granular.
The front is cut into two sharply denticulate lobes, the outer angle
of each of which is very distinctly isolated and spine-like. The orbital
margin is denticulate and has the three fissures distinct.
The antero-lateral margin has four large procurved spines, some
of which (almost constantly the second one) may have an accessory
spinule near the tip. Three or four of the lobules just inside either
antero-lateral margin bear each a somewhat similar spine.
The outer angle of the basal antennal joint is prolonged into the
orbital hiatus.
The chelipeds are unequal, the larger one being not quite twice the
jJength of the carapace. The arm has one or two spine-like teeth at
the distal end of both the anterior and the posterior border; the wrist
is studded with spine-like tubercles and has a pair of strongish spines
at the inner angle; the hand has spine-like tubercles along the upper
surface, and close-set pearly granules along the outer and lower sur-
faces ; fingers with spine-like denticles at base only.
Meropodites of legs with the upper border spinulate ; carpopodites
1898. } A. Aleock —Carcinological Fauna of India. 171
and propodites each with two or three rows of spinules and sharp
granules.
Colours in spirit—yellowish or reddish brown ; fingers black, the
colouration stopping sharply at the base of the thumb.
Carapace 8 millim. long, 13 millim. broad.
In the Indian Museum are 19 specimens from the Andamans.
This species is very closely related to C. melanodactylus, A. M. Edw.
(of which we have in the Museum specimens from Samoa) but differs
in having (1) only a few scattered hairs on the carapace, (2) the front
deeply bifid and elegantly denticulate, with the outer angle isolated
and spine-like, (3) the sculpture of the carapace much sharper and
bolder, (4) the pearly granules and spine-like tubercles of the chelipeds
more numerous and close-set.
From C. spinipes it differs in having (1) the front sharply spinu-
late, (2) the first spine of the antero-lateral border almost as large and
well spaced as the other three, the 2nd and 3rd spines moreover having
almost always an accessory spinule near the tip, (3) the spines of the
legs not so large and acicular and not so much concealed by hairs,
It may very possibly be the Pilodius scabriculus of Dana.
Sub-genus Crcropius, Dana.
Cyclodius, Dana, Silliman’s Amer. Journ. Sci. and Arts, (2) XII. 1851, p. 126;
and U.S. Expl. Exp. Crust. pt. I. p. 222.
Cyclodius agrees in every particular with Chlorodopsis, excepting
only that the carapace is longer and narrower, being, in fact, almost as
much sub-circular as hexagonal.
In general form, as in the relations of the basal antennal joint,
Cyclodius much resembles Htisodes, from which, however, the form and
breadth of the front at once distinguishes Cyclodius.
95. Chlorodopsis (Cyclodius) ornata, Dana.
Cyclodius ornatus, Dana, Proc. Acad, Nat. Sci. Philad. VI. 1852, p. 80; and
U.S. Expl. Exp. Crust. pt. I. p. 223, pl. xii. figs, lla-g.
Carapace flattish, about + as long as broad, almost as much sub-
circular as hexagonal, its regions and subregions delimited by well cut
grooves, the subregions being numerous and having a microscopically
granular surface.
Front a little more than half the greatest breadth of the carapace,
bilobed, the outer angle of each lobe well pronounced. Orbital margin
with two grooves above and one at the outer angle.
Antero-lateral margin cut into four teeth (exclusive of the orbital
angle) the last three of which are procurved and claw-like.
172 A. Alcock — Carcinological Fauna of India, [No. 1,
Basal antennal joint prolonged into the orbital hiatus—and filling
it—on the same extensive scale as in Chlorodopsis areolata.
Chelipeds very little unequal, not much longer and stouter than
the legs, about 1} times the length of the carapace: arm with several
spinules along the posterior border and two large ones on the anterior
border ; wrist and hand with numerous sharp spine-like tubercles, which
fall into longitudinal series on the outer suiface of the hand; fingers
with some coarse spinules at base, rather strongly arched, broadened
and hollowed at tip.
Legs granular, somewhat furred, the upper border of the mero-
podites carpopodites and propodites spinate.
The grooving of the under surface of the carapace, found in all
the species of Chlorodius, Chlorodopsis, &c., is particularly elegant.
In the Indian Museum is a male from the Andamans (and one
from Mauritius).
Alliance IIL. Cymorda.
Cymo, De Haan.
Cymo, De Haan, Faun. Japon. Crust. p. 22.
Cymo, Dana, Amer. Journ. Sci. and Arts, (2) XII. 1851, p. 126; and U. 8S. Expl.
Exp. Crust. pt. I. p. 224.
Carapace about as long as broad, subcircular, or less commonly
elongate-pentagonal ; not, or little, convex; depressed, with regions and
subregions faintly or not at all shown.
Fronto-orbital border from about 3 to? the greatest breadth of
the carapace in extent. Front from about $ to about 4 this measure,
horizontal, bilobed, with the outer angle of each lobe prominent and
separated from the supra-orbital margin by a notch and groove. The
grooves of the orbital margin are either indistinguishable or distinct.
Hyes on short thick stalks.
The antennules fold obliquely. The basal joint of the antenne has
its outer angle produced into the orbital hiatus, and the flagellum,
which is short, is situated between this process of the basal joint and
the front.
The chelipeds are remarkably unequal in both sexes, the larger
cheliped, in adults, being more than half again as long and more than
twice aS massive as the smaller: the fingers of the larger cheliped are
short, thick, blunt-pointed (beak-like) aud hollowed at tip; those of the
smaller hand, though also hollowed-out, are long and slender.
The legs are invested and fringed with a thick shaggy fur that en-
tirely conceals their sculpture: they are short and massive.
1898. ] A. Alcock —Carcinological Fauna of India. 173
The abdomen of the male consists of five joints, the 3rd-5th somites
being fused.
The species of this genus are at once recognized by the subcircular
carapace, which even in the male leaves the first two and part of the
third abdominal terga exposed in a dorsal view; and by the remarkable
inequality and dissimilarity of the chelipeds.
Key to the species of Cymo.
I. Carapace subcircular, depressed, but not quite flat :—
i. Wrists and hands studded with sharpish gra-
nules only: front bilobed, the edge of each
lobe denticulate :—
L. Fingers Whitd......0ccoccecre: covtasseseceves’ O. andreossyi:
2. Fingers black pene at oa, wees CO. melanodactylus
ii. Wrists and hands with large granular sau as
well as granules :—
1. Front bilobed, the edge of each lobe
concave and the angles in the form
of granular tubercles, so that the front
appears four-lobed...........seeseessereseeee. C. quadrilobatus.
[2. Front not four-lobed.. ............ geek C. tuberculatus. |
II. Carapace more elongate-pentagonal than subcircular,
PTE GLY) HA Uiiina eno tigen sa Vall «oo vesisia=ancaa'cs eines sede oniged ne ales «. OC. deplanatus,
96. Cymo andreossyt, (Audouin) De Haan.
Pilwmnus? andreossyi, Audouin on Savigny’s Descr. de lEgypte, pl. v. fig. 5,
p. 86.
Cymo andreossyi, De Haan, Faum. Japon. Crust. p. 22: Dana, U.S. Expl. Exp.
Crust. pt. I. p. 225, pl. xiii. figs. 2 a-b : Stimpson, Proc. Ac. Nat. Sci. Phila. 1858,
p. 34: Heller, SB. Ak, Wien, XLIII. 1861, p. 346, and Novara Crust. p. 20: A.
Milne Edwards, Nouv. Archiv. du Mus, IX. 1873, p. 252: Kossmann, Reise roth.
Meer. Crust. p. 35: Miers, Phil. Trans. Vol. 168, 1879, p. 487, and Zool. H. M.S,
Alert. pp. 517, 582: de Man, Archiv. fur Naturges, LIII. 1887, i, p. 291: J. R.
Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 363: Ortmann, Zool. Jahrb.
Syst. VII. 1893-94, p. 443.
Carapace almost circular, its greatest length being inappreciably
less than it greatest breadth, closely covered with a spongy fur on
removal of which can be seen (1) faint depressions demarcating the
gastric and cardiac regions and incompletely separating the gastric
region into three sub-regions, others subdividing the branchio-hepatic
regions into faintly convex areole, and (2) a few granules on the
anterior part of the gastric region (in a transverse line) and on some
of the lobules of the branchio-hepatic regions.
Frout rather more than 2 the greatest breadth of the carapace in
extent, bilobed, separated rise the dentiform supra-orbital angle by a
174 A. Alcock — Carcinological Fauna of India. [ No. 1,
groove, its free edge irregularly denticulate. Orbital margin entire,
Lateral borders of the carapace somewhat granular.
Chelipeds markedly unequal, covered with fur, but not so much as
to entirely conceal their sculpture. The arm has both borders fringed
with fur and the distal corner of the upper surface granular; the
upper aud outer surfaces of the wrist are studded with sharpish
granules, as are the upper, outer and part of the inner surfaces of the
hands and the basal half of the finger — those towards the upper part
of the hand having a linear arrangement. ‘The fingers of the larger
cheliped are stout, truncated, blunt-pointed and strongly hollowed at
tip; those of the smaller cheliped, though also hollowed, are thin,
slender and pointed.
The legs are covered with a thick shaggy coat of fur, which is
specially long and adherent along the borders. When this is removed
the upper edge of the meropodites is finely granular, and the upper
borders of the following joints are traversed by several rows of sharpish
granules.
Colours in spirit, brownish yellow or fawn-colour, fingers white.
In the Indian Museum are 11 specimens, from Mekran coast,
Ceylon, Andamans and Nicobars (besides 11 from other parts of the
Indo-Pacific).
97. Cymo melanodactylus, De Haan.
Cymo melanodactylus, De Haan, Faun. Japon. Crust. p. 22: Dana, U. 8S. Expl.
Exp. Crust. pt. I. p. 225, pl. xiii. fig. 1: Stimpson, Proc. Acad. Nat. Sci. Philad.
1858, p. 34: A. Milne Edwards, Nouv. Archiv. du Mus, IX. 1873, p. 252: Ortmann,
Zool. Jahrb. Syst. VII. 1898-94, p. 442.
Cymo andreossyi var. melanodactylus, Miers, Zool. H. M. 8S. Alert, p. 533
de Man, Journ. Linn. Soc. Zool. XXII. 1887-88, p. 35.
Differs from C. andreossyi in the following particulars :—
(1) the carapace is more lumpy; (2) the anterior half of the
lateral borders shows more distinct indications of three lobules,
some of which may even bear a spinelet; (3) the fingers are black,
except at tip.
In the Indian Museum are 8 specimens, from the Andamans,
Mergui and Ceylon, (besides 2 from other parts of the Indo-Pacific.)
98. ? Cymo deplanatus, A. Milne Edwards.
? Cymo deplanatus, A. Milne Edwards, Journ. Mus. Godeffr. I. 1873, p, 257.
This species, if I am correct in my identification, differs from
Cymo andreossyt only in the following particulars :—
The carapace is less subcircular and more elongate-pentagonal ;
1898. | A. Alcock —Carcinological Fauna of India. 175
it is as flat as a coin and is either quite smooth or has only a trans-
verse row of granules in the anterior part of the gastric region: the
somewhat pentagonal outline is due to the antero-lateral borders being
convergent from a distinct, though obtuse, angle of union with the
postero-lateral.
99. Cymo quadrilobatus, Miers.
Cymo quadrilobatus Miers, Zool. H. M.S. Alert, p. 533.
General form as of 0. andreossyt.
The carapace is covered with a fine close down which does not,
however, in the least conceal its sculpture.
On either side of the carapace just behind the front are two
granular transverse elevations (=Dana’s areole 2 F and 1M): the
branchio-hepatic regions are distinctly areolated, the areole having the
form of elevated clusters of pearly granules.
The front is really bilobed, but as each lobe has a deeply concave
edge and both angles surmounted by a granular tubercle, it appears
four-lobed.
The three grooves near the outer angle of the (beaded or crenulate
and somewhat tumid) orbital margin are very distinct. The anterior
half of the lateral margin of the carapace is divided into three granular
lobes. |
The chelipeds have the same general form and proportions as in
C. andreossyi: they are more or less invested with a fine down, which
does not conceal their sculpture: the upper surface of the arm is
covered with pearly granules: the upper and outer surfaces of the
wrist, and the upper, outer and much of the inner surfaces of the
hands, are covered with pearly granules, many of which, on the wrist
and in lines along the upper surface of the hand, unite to form
large wart-like tubercles: the fingers of the larger cheliped are stout,
truncated, blunt pointed and somewhat hollowed at tip; those of the
smaller cheliped, though hollowed, are thin and pointed.
The legs are thickly covered with fur and long adherent silky
hairs, beneath which the whole dorsal surface of the last four joints is
granular.
Colours in spirit: light yellow, with either livid or rich chestnut
brown mottled markings on the carapace: fingers of the larger hand
whitish, usually with a black base, those of the smaller hand black with
white tips; in both cases the distal half of the lower border of the hand
is black.
Carapace of largest specimen in the Indian Museum collection 15°5
millim. long, 16 millim. broad.
176 A. Alcock — Carcinological Fauna of India. [No. 1,
Tn the Indian Museum are 5 specimens, from Palk Straits, 5-7 fms.,
and off Little Andaman 12 fms.
100. Cymo tuberculatus, Ortmann.
Cymo tuberculatus, Ortmann, Zool. Jahrb. Syst. VII. 1893-94, p. 443.
This species, from the Maldive Islands, resembles C. quadrilobatus
in the characteristic sculpture of the chelipeds, and may perhaps be
identical with Miers’ species,
It is not represented in the Indian Museum.
Section II, Hyperomerista.
The efferent branchial channels are defined by a ridge on either
side of the palate, the ridges extending right up to the epistomial
edge.
Sub-family [V. Mentppina.
Carapace broad, transversely oval; front a fourth, or less than a
fourth the greatest breadth of the carapace. The basal antennal joint
does not nearly reach the front. The abdomen of the male has all 7
segments distinct and separate.
Alliance I. Menrpetorpa. Carapace convex, its antero-lateral
borders longer than the postero-lateral: ridges of the eudostome
faint.
Alliance II. Psrupoztota. Carapace flat, its antero-lateral borders
shorter than the postero-lateral: ridges of the endostome strong and
sharp.
Sub-family V. Ozina.
Carapace broad, transversely oval; front broad, about a third the
greatest breadth of the carapace. The basal antennal joint is broadly
in contact with the front. All 7 segments of male abdomen distinct
and separate. The efferent branchial channels very distinct and cir-
cumscribed.
Alliance I. Oztomwa. The orbital hiatus is open and is occupied
by the antennary flagellum.
Alliance Il. Ruepuettiorsa. The orbit is a completely closed
cavity.
Sub-family VI. Pitumnina.
Carapace moderately broad; front about a third the greatest
breadth of the carapace: the antero-lateral borders of the carapace
1898. ] A. Aleock— Carcinological. Fauna of Fndia. 177
are not longer than, and are often shorter than, the postero-lateral.
The basal antennal joint does not touch, or only just touches, the
front. :
Alliance J. Pinumnompa. Carapace commonly densely tomentose,
its regions commonly well defined and areolated.
Alliance Il. Hereropanopiotpa. Carapace smooth, its regions
either not at all, or not very well defined.
Sub-family VII. Enrieatna.
Carapace sub-quadrilateral, the antero-lateral borders not forming
an arch but meeting the postero-lateral borders at a very open and
inconspicuous angle. Front very broad, half or more the greatest
width of the carapace, and, with the orbits, occupying the whole anterior
border of the carapace. Basal antennal joint not touchiug the front.
Abdomen of the male either with all 7 segments distinct or with the
3rd, 4th and 5th fused.
Alliance I. Eripuioipa. The gastric region, at least, is well
defined: basal antennal joint short and thick: orbits deep: arms stout
and short.
Alliance II. Trapsztoipa. Carapace perfectly smooth, without
trace of regions: basal antennal joint slender: orbits shallow, affording
little concealment to the eyes: arms long or very long, projecting in
large part or entirely beyond the carapace, in repose.
Alliance III, Domectorpa, No trace of regions; orbits shallow ;
arms short; legs, chelipeds, and frontal and antero-lateral borders of
carapace strongly spinate. Merus of external maxillipeds more than
twice as broad as long.
Alliance IV. Metiorpa. Carapace hexagonal, the regions either
absent or fairly well defined ; basal antennal joint slender; orbits
shallow. The chelipeds are very much shorter and slenderer than the
legs.
Subfamily IV. Menippina.
Alliance I. Menippioida.
Menippe. Myomenippe.
Mentrre, De Haan.
Menippe, De Haan, Faun. Japon. Crust. p. 21.
Pseudocarcinus, Milne Edwards, Hist. Nat. Crust. I. 407.
Menippe, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX, 1863, p. 280; and
Exp. Sci. Mex. Crust. p. 262.
Carapace broad, transversely oval, moderately convex fore and aft,
J. u, 23
178 A. Aleock — Carcinological Fauna of India. [ Noed;
very slightly so from side to side; the regions, except the gastric, little
defined.
Antero-lateral borders long, strongly arched, cut into four teeth ;
postero-lateral borders slightly shorter than antero-lateral, convergent ;
posterior border short.
Front narrow, less than a fifth the greatest uneaaih of the carapace,
rather prominent, almost horizontal, cut into two prominent lobes, the
outer angle of each of which forms a distinct tooth.
Orbit with the three grooves near the outer angle well marked:
inner orbital angles—both upper and lower—well pronounced. LHyes
on short thick stalks.
The side edges of the front are not turned down and the short
basal antennal joint does not nearly reach the front, so that the cavities
of the orbits and antennules are not properly separated : the next antennal
joint just reaches the front, and the long antennary flagellum stands in
the orbital hiatus. The antennules fold nearly transversely.
The anterior edge of the merus of the external maxillipeds is
oblique and a little sinuous but not excised.
The ridges of the endostome, defining the expiratory channels, are
complete, but low and faint.
Chelipeds massive, a little unequal in both sexes; fingers stout,
pointed, not hollowed.
Abdomen of male singularly broad, all seven segments distinct.
101. Menippe rumphii, Fabr., v. Martens.
Cancer rumphii, Fabr., Ent. Syst. Suppl. p. 836: Herbst, Krabben, III. i. 63,
pl. xlix. fig. 2.
Menippe rumphii, v. Martens, Archiv. fiir Naturges. XXXVIII. 1872, p. 88:
de Man, Journ. Linn. Soc., Zool., XXII. 1887-88, p. 36: Henderson, Trans, Linn.
Soc., Zool., (2) V. 1893, p. 363.
Pseudocarcinus bellangeri, Milne Edwards, Hist. Nat. Crust. I. 409, pl. xiv bis,
fig. 15.
Menippe bellangeri, Heller, Novara Crust. p. 15: Muller, Verh. Ges. Basel, VIII.
1886, p. 474,
Gastric region distinct and fairly distinctly. subdivided into three
lobes; between it and the front are four pimple-like tubercles standing
ina square. Two low indistinct somewhat granular elevations, nearly
parallel with the curve of the antero-lateral border, traverse either
branchial region; the first, which is the more distinct, can generally be
traced across the gastric’ region also. The surface of the carapace is
finely pitted antero-laterally, but elsewhere is smooth. |
The front, which is not quite a fifth the greatest breadth of the
carapace, consists of two prominent round-pointed lobes, outside of
1898, ] : A. Alcock— Carcinological Fauna of India. 179
each of which is a prominent: rounded tooth separated from the supra
orbital margin by a groove.
The antero-lateral border is fairly sharp and is divided into four
broad lobes, of which the last two are distinctly, the first two indis-
tinctly, acuminate.
-Chelipeds massive, a little unequal, smooth with some fine and
distant pitting: inner angle of wrist bluntly prominent; fingers stout,
rather short.
Legs stout, smooth, except the upper border which is sometimes
microscopically granular: upper border of carpopodites sparsely, both
borders of propodites and dactyli more thickly, hairy.
‘Colours in spirit reddish or brownish yellow with sometimes a fine
network of darker markings ; fingers black.
In the Indian Museum are 100 specimens, from Penang, Tavoy,
Mergui, Madras coast, Ceylon, Laccadives, Karachi and Persian Gulf.
Subgenus Myomenippe, Hilgendorf.
Myomenippe, Hilgendrof, MB. Ak. Berl. 1878, p. 795.
Closely resembles Menippe in all respects, but differs (1) in the
orbit being a completely closed cavity, owing to the contact of its upper
and lower inner angles; hence the long antennary flagellum is quite
excluded from the orbit, and (2) in the front being rather broader
(nearly a fourth the greatest breadth of the carapace) and six-lobulate.
102, Menippe (Myomenippe) granulosa, A. M. Edw.
Menippe granulosa, A. Milne Edwards, Ann. Soc, Ent. Fr. (4) VII. 1867,
p. 275.
Myomenippe duplicidens, Hilgendorf MB. Ak. Berl. 1878, p. 796, (fide de
Man.)
Myomenippe granulosa, de Man, Journ. Linn. Soc. Zool., XXII. 1887-88, p. 40,
pl. ii. fig. 1; and Zool. Jahrb., Syst., VIII. 1894-95, p. 525.
The gastric region is fairly well demarcated and subdivided into
three areas, the two antero-lateral of which have the surface broken
up into low granular convexities : the lateral regions of the carapace are
also rugose, the wrinkles being granular and falling into two broken series
almost parallel with the curve of the antero-lateral borders. Every
margin of the carapace is granular, as is also—besides the rugosities
already mentioned—but more finely, a good deal of the surface near
the margins.
The antero-lateral border is thin and rather sharp and is cut into
four teeth, the first three of which are broad and anteriorly acuminate,
the last narrow and carinated.
180 A. Alcock — Carcinological Fauna of India. [No. 1,
The front, which is nearly a fourth the greatest breadth of the
carapace, is prominent, is separated from the orbit by a deep notch,
and is bilobed, each lobe being cut into three teeth. The inner lower
angle of the orbit is of the same size and form and as prominent as
the innermost (largest) lobule of the frontal lobes.
Chelipeds massive, a little unequal; upper and outer surfaces of
wrist and upper (and sometimes in the case of the smaller cheliped the
greater part of the outer) surface of hand granular; fingers stout and
rather short : inner angle of wrist sharply prominent, somewhat upcurved,
a finely beaded line passing from its summit, backwards, along the whole
length of the wrist.
Legs stout, with a rough and furred or scurfy surface, the upper
border of the last four joints and the lower border of the last two
rather abundantly fringed with fine stiff hairs.
Colours in spirit light brown, or greenish brownish yellow ; fingers
black.
In the Indian Museum are 6 specimens, from Mergui, Arakan,
Diamond I., Singapore,
Alliance II. Pseudozioida.
Psgvupozius, Dana.
Pseudozius, Dana, Silliman’s Journal (2) XII. 1851, p. 127; Proc. Ac. Nat, Sci.
Philad. 1852, p. 81; and U. S. Expl. Exp. Crust. pt. I. p. 232,
Pseudozius, Miers, Challenger Brachyura, p. 141.
Carapace broad, transversely oval, little convex or quite flat, the
regions not demarcated.
Antero-lateral border arched, shorter than postero-lateral border,
obscurely divided into four very shallow lobes.
Front rather broad, much more than a fourth the greatest breadth
of the carapace, separated from the orbit by a notch, excised in the
middle line and having the outer angles pronounced,—and so, obscurely
four-partite.
Orbital margin entire, the upper and lower inner angles almost in
contact. The antennules fold nearly transversely.
Basal antennal joint very short, the next joint reaches the front;
the flagellum, which is hardly as long as the major diameter of the
orbit, lodged in a notch between the front and the orbital wall, but
quite outside the latter.
The crests of the endostome, defining the expiratory channels, are
strong, and the anterior border of the merus of the external maxillipeds
is notched to assist in forming a permanent expiratory orifice.
1898,] A. Alcock — Carcinological Fauna of India. 18L
Chelipeds massive, unequal in both sexes, the fingers pointed, not
hollowed.
Abdomen of the male with all 7 segments distinct.
103, Pseudozius caystrus (Ad. and White) Miers.
Panopeus caystrus, Adams and White, Samarang Crust. p. 42, pl. ix. fig. 2.
Pseudozius planus, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 81; and U.S.
Expl. Exp. Crust. pt. I. p. 233, pl, xiii, figs.6a-h: Richters in Mobius, Meeresf.
Maurit. p. 148.
Pseudozius caystrus, Miers, Challenger Brachyura, p. 142: Ortmann, Zool.
Jahrb., Syst., VII, 1893-94, p. 434; and in Semon’s Forschungsr. (Jena. Denk.
VIII), Crust. p. 49: de Man, Zool. Jahrb., Syst., VIII, 1895, p. 525: Whitelegge,
Mem. Austral. Mus. III, 1897, p. 136.
_Carapace transversely oval, depressed, smooth, almost flat behind
the deflexed finely granular fronto-orbital region: no distinct regional
boundaries.
The antero-lateral border is fairly sharp and is obscurely divided
into four shallow lobes, the first two of which are rounded and almost
confluent.
Front between a third and a fourth the greatest breadth of the
carapace, bluntly four-partite.
Orbits with the margins entire, eyes small.
The buccal cavern is distinctly narrower anteriorly than posteriorly.
Chelipeds unequal, very massive, quite smooth to the naked eye ;
two strongish tubercles at the inner angle of the wrist; fingers arched,
pointed, in the adult male they meet only at tip.
Legs smooth, dactyli furred, a few fine scattered silky bristles on
the propodite.
Colours in spirit brownish yellow, fingers darker.
In the Indian Museum are 63 specimens, mostly from the An-
damans, but also from the Mekran (Baluchistan) coast, the Laccadives,
and Bombay or Aden. (Also 2 from Samoa and 1] from Bantam).
Subfamily V. Ozina.
Alliance I. Ozioida.
Ozius. Epixanthus,
Ozius, Edw.
Ozius, Milne Edwards, Hist. Nat. Crust. I. 404.
Ozius, Dana, Silliman’s Journ, (2) XII. 1851, p. 127; and U. S. Expl. Exp.
Crust. pt. I. p. 229.
Ozius, A. Milne Edwards, Ann. Sci. Nat. Zool. (4) XX. 1863, p. 289; and Nouv.
Archiv, du Mus. 1X. 1873, p. 237; and Miss. Sci. Mex., Crust. p. 276.
182 A. Aleock— Carcinological Fauna of India. [ Notiy
Carapace broad, transversely oblate-oval, moderately convex fore
and aft, slightly convex or nearly flat from side to side; the regions,
except the gastric, little defined ; the surface smooth, or granular, often
rugose anteriorly.
Antero-lateral borders of good length, strongly arched, usually
broadly crenate or lobulate: postero-lateral borders convergent, usually
about as long as the antero-lateral.
Front rather broad (considerably more than a fourth the greatest
breadth of the carapace) obliquely deflexed, cut into four lobules or
teeth of about equal size, separated from the orbit by a notch.
Orbits deep, rather small, the grooves near the outer angle in-
conspicuous: eyes on short thick stalks. The antennules fold nearly
transversely.
Basal antennal joint prolonged between the side of the ay and
the orbital plate ; the flagellum, which is very small (about half the
major diameter of the orbit in length), stands in the orbital hiatus.
The ridges of the endostome, defining the expiratory channels,
are very strong, and the opposed margin of the merus of the external
maxillipeds is notched, usually very deeply, so that a permanent
expiratory orifice results.
Chelipeds massive, unequal in both sexes; the fingers of good
length, pointed not hollowed. In the Indian species there is a very
large tooth at the base of the dactylus of the larger hand.
The abdomen of the male consists of 7 segments.
Key to the Indian species of Osius.
I, Carapace more than 3? as long as broad, scabrous, more
or less studded — like the wrists and hands—with salient
Oar ly MMDEFGlES vaca, part eodheee de cee nee clies eesnseors ww... O, tuberculosus,
II. Carapace 3 as long as bean ‘sui datl to feel, no titer bess ;
surface of wrists and hands—all or part—reticulate
PBESMORS! ith. 1. hsb is sduvles tec gee ddd Lal's See Ggited ba} Dos ew h'siveiaue'n st!) GQ MiaMmnaR IEE
104. Ozius rugulosus, Stimpson.
Ozius rugulosus, Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 34: Heller,
Novara Crust. p. 22, pl. iii. fig. 1: A. Milne Edwards, Nouv. Archiv. du Mus. 1V.
1868, p. 71, and IX. 1873, p. 240, pl. xi. fig. 3: Miers, P. Z. S. 1877, p. 185: Haswell,
Cat. Austral. Crust. p. 63: Cano, Boll. Soc. Nat. Napol. LII. 1889, p. 204: Ortmann,
Zool. Jahrb., Syst., VII. 1893-94, p. 477, and in Semon’s Forschungsr. (Jena. Denk.
VIII.) Crust. p. 53.
Carapace two-thirds as long as broad, its surface everywhere finely
pitted but not rough, a good deal rugulose and finely eroded just inside
the antero-lateral borders: gastric region fairly well defined and in-
1898. ] A. Aleock — Carcinological Fauna of India. 183
completely subdivided into three areole: branchial regions traversed
by two ridges, which run respectively from the 8rd and 4th lobes of
the antero-lateral borders, obliquely upwards and inwards to the gastric
region.
Front cut into 4 equidistant teeth. Antero-lateral border rather
faintly divided into 5 lobes, of which the first two are broad and round-
ed and the last three are bluntly acuminate. Orbital margin slightly
tumid, well marked off from carapace; faint traces of two grooves
near the outer angle; the inner angle of the lower margin a little
prominent.
The upper and outer surfaces of both wrists and of the smaller
hand, and the upper surface of the larger hand are reticulate-rugulose.
Legs stout, the last three joints and part of the under surface of
the meropodites of all are tomentose.
Efferent branchial foramen large, sub-quadrangular.
Colours in spirit, dark violet brown or dark bluish brown, fingers
black.
In the Indian Museum are 5 specimens, from the Andamans and
Arakan.
105. Ozius tuberculosus, Edw. |
Ozius tuberculosus, Milne Edwards, Hist. Nat. Crust. I. 405: Heller, Novara
Crust. p. 23: A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873, p. 238, pl. xi.
fig. 2: Muller, Verh. Ges. Basel VIII. 1886, p. 474: de Man, Journ. Linn. Soc., Zool.,
XXII. 1887-88, p. 45: J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 364.
Carapace more than two-thirds as long as broad, studded (except
sometimes in the central and posterior parts) with small sharp pearly
tubercles; gastric region well defined and imperfectly divided into
elongate areole; branchial regions crossed transversely by two cres-
centic furrows, which have a common starting-point at the fourth tooth
of the antero-lateral margin; post-orbital furrow well defined.
Front sunk below the level of the orbits, cut into four equidistant
scabrous teeth. Antero-lateral border granular, cut into five teeth
(exclusive of the orbital angle), of which the last is tuberculiform and
the first four are broad and anteriorly-acuminate.
The lower edge of the orbit is separated from the tumid acch of the
upper edge by a small gap, and is deeply concave between the dentiform
external and internal angles.
Basal antennal joint massive, sinuous, granular. Efferent branchial
foramen large, subcircular. |
Chelipeds and legs with rough harsh surfaces: the upper and outer
surfaces of the wrists and hands are for the most part studded with
184 A. Alcock — Carcinological Fauna of India. [No. ],
sharp pearly tubercles like those on the carapace: the inner angle of
the wrist is somewhat produced and forms a double-crowned tubercle.
The whole animal has a harsh feel, due partly to the roughness of
the surface and partly to the presence of very short, stubbly, scattered
bristles.
Colours in spirit, light red or madder, fingers darker, dactyli of
legs blackish.
In the Indian Museum are two specimens, from Mergui and the
Nicobars.
Epixantnos, Heller.
Epizanthus, Heller, SB. Ak. Wien, XLIII. 1861, i. p. 323.
Epivanthus, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX. 1863, p. 290; and
Nouv. Archiv. du Mus. IX. 1873, p. 240.
Epizanthus, de Man, Journ. Linn. Soc. Zool. XXII. 1887-88, p. 45.
Carapace very broad, transversely oval, either moderately convex
or almost flat, the regions very obscurely marked.
Antero-lateral borders long, strongly arched, with a thin sharp
edge, usually regularly fissured or dentate: postero-lateral borders
strongly convergent.
Front broad (from one-fourth to nearly one-third the greatest
breadth of the carapace), slightly deflexed, separated from the supra-
orbital margin by a notch, cut into four teeth or lobes. Hither a suture
or a gap beneath outer angle of orbit.
Antennules folding transversely, inter-antennulary septum broad.
Basal antennal joint very broad and short, largely in contact with the
front ; flagellum very short (less than half the major diameter of the
orbit), lodged in the orbital hiatus.
The ridges of the endostome, defining the expiratory canal, are
very strong, but the anterior border of the merus of the external
maxillipeds is either not at all or only very slightly notched.
Chelipeds massive, unequal in both sexes; fingers long, pointed,
those of the smaller hand being remarkably long and slender.
Abdomen of the male with all 7 segments distinct.
Key to the Indian species of Epixanthus.
I. Carapace nearly flat, nearly smooth; antero-lateral
border divided by very short narrow fissures into four
BVORM BHALMOW LODGI ics vc so coiscy.scccecrsedeedicco.cossavesicgores dina RGR MmmEE
II. Carapace convex, scabrous; antero-lateral border deep-
' ly cut into five sharp thin teeth..,.......cccscscscossserseesee 2, dentatus.
1898. ] A. Aleock — Carcinological Fauna of India. 185
106. Hpixanthus frontalis, (Edw.) Heller.
Ozius frontalis, Milne Edwards, Hist. Nat. Crust. I, 406: Krauss, Sudafr.
Crust. p. 31: Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 34: Hilgendorf, in v. d.
Decken’s Reisen Ost-Afr. III, i. p. 75.
Epizanthus frontalis, Heller, Novara Crust. p. 20: A. Milne Edwards, Nouv.
Archiv. du Mus. IX, 1873, p. 241: Kossmann, Reise roth. Meer., Crust. p. 36:
Richters, in Mébius, Meeresf. Maurit. p. 148, pl. xvi. fig. 16: Lenz and Richters,
Abh. senck. Ges. XII, 1881, p. 421: Miers, Zool. H. M. S. Alert, pp. 517, 534:
F. Muller, Verh. Ges, Basel, VIII, 1886, p. 474: de Man, Journ. Linn. Soc. Zool.,
XXII, 1887-88, p. 46; and Archiv. fur Naturges. LIII, 1887, i. p. 292; and Zool.
Jahrb., Syst. 1894-95, p. 525; Cano, Boll. Soc. Nat. Napol. III, 1889, p. 205; J. R.
Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 364: Ortmann, Zool. Jahrb.,
Syst., VII, 1893-94, p. 477.
Epivanthus kotschii, Heller, SB. Ak. Berl. XLIII, 1861, i. p. 325, pl. i. fig. 14
( fide Heller),
Carapace transversely oval, depressed, almost flat, its length a little
over 3 its breadth, granular and finely and faintly rugulose just inside
the frontal and antero-lateral borders, smooth elsewhere. The gastric
region and its three subregions are faintly indicated, and a low fine
sinuous ridge completely traverses each branchial region from tle last
tooth of the antero-lateral border.
The front, which is a good deal less than a third the greatest
breadth of the carapace, and has a double edge, is cut into four low
teeth, Below the outer angle of the orbit there is a suture, not a gap.
Antero-lateral border thin and sharp, divided by short, narrow
notches into four very broad shallow lobes, of which only the last two
are at all acuminate.
Chelipeds massive, remarkably unequal—in the adult male espe-
cially. They are practically smooth. The fingers of the larger hand
of the adult male are strongly arched and meet only at tip.
Legs almost smooth; the borders of the dactylus and of the distal
half of the propodite, in all, are covered with a short stubbly fur.
Colours in spirit, dirty yellowish or greenish brown, fingers blackish.
In the Indian Museum are 60 specimens, from the Andamans,
Mergui, Akyab, Orissa coast, Ceylon, Makran coast (besides 22 speci-
mens from localities outside India).
107. Hpixanthus dentatus, (White).
Panopeus dentatus, White, P. Z. S. 1847, p. 226; Ann. Mag. Nat. Hist. (2) IT,
1848, p. 286 ; Adams and White ‘Samarang’ Crust. p. 41, pl. xi. fig. 1.
Heteropanope dentatus, Stimpson, Proc. Ac, Nat. Sci. Phila. 1858, p. 35: A,
Milne Edwards, Nouv. Archiv. du Mus. IV. 1868, p. 71.
Epixanthus dentatus, Miers, Ann. Mag. Nat. Hist. (5) V. 1880, p. 233: de Man,
Journ, Linn. Soc., Zool., XXII. 1887-88, p. 46: Henderson, Trans, Linn, Soc., Zool.,
J. u. 24
186 A. Alcock — Carcinological Fauna of India. [No. 1,
(2) V. 1893, p. 364: Ortmann, Zool. Jahrb., Syst., VII. 1893-94, p. 478 ; and in
Semon’s Forschungsr. (Jena. Denk. VIII.) Crust. p. 53.
Epizanthus dilatatus, de Man, Notes Leyden Mus. I. 1879, p. 58 (fide de Man).
Panopeus acutidens, Haswell, P. L. 8., N.S. W., VI. 1881-82, p. 542; and Cat.
Austral. Crust. p. 51, pl. i. fig. 2.
Carapace transversely oval, convex fore and aft, slightly so from
side to side; its length about 5% its breadth; its surface granular and
somewhat tuberculous anteriorly, the tubercles being almost squami-
form and fringed with short stubbly hair. The gastric region and its
three subregions are very faintly indicated: the branchial regions are
traversed by a low, sinuous, finely granular ridge.
The front, which is somewhat less than a third the greatest
breadth of the carapace, has a rather indistinctly double edge and is
cut into four lobes. There is a distinct gap in the orbital margin just
below the outer orbital angle. 3
The antero-lateral border is deeply cut into five very thin sharp-
edged teeth.
The exposed surfaces of the arms wrists and hands are finely reti-
culate-rugulose (most strongly marked on the hands) the reticulating
wrinkles being covered with a very short stubbly or scurfy tomentum.
Similar reticulating lines and patches of the same stubbly or scurfy
growth also closely cover the surfaces of the leg joints.
Colours in spirit, dull earthly brown or yellowish, the carapace and
chelipeds commonly mottled or marbled.
In the Indian Museum are 5 specimens, from Mergui and the
Andamans (besides 2 from the South Sea Is.).
Alliance II. Ruppellioida.
Kuruppellia. Baptozius.
Subgenus HurvpPELLIA.
Ruppellia, Milne Edwards, Hist. Nat. Crust. I. 420 (part).
Ruppellia, Dana, Silliman’s Journal (2) XII. 1851, p. 128, and U. S. Expl. Exp.
Crust. pt. I. p. 245.
Ruppellia, A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX. 1863, p. 291.
Euruppellia, Miers, Zool., H. M.S. Alert, p. 534.
Differs from Ozius only in the form of the orbits. The upper and
lower inner angles of the orbit are in contact, so as to close the orbit
and to completely exclude the antennary flagellum.
I do not think this character is of generic importance, and I agree
with Kossmann that the type of this genus should be included with
Oxzus.
1898. ] A. Alcock — Carecinological Fauna of India. 187
There is however one species, Rwppellia vinosa, Edw., that is
entirely different from any of the species (with, perhaps, the exception
of Ruppellia lata, A. M. E.) with which it has hitherto been supposed
to be congeneric.
This species I have separated as the type of a new genus Baptozius.
If, however, the other species of H. Milne Edwards’ genus Ruppellia
are referred to Ozius, then the name Huruppellia must be retained, in
a different sense, for Ruppellia vinosa Hdw., and the name Baptozius
must lapse.
108. Ozius (Huruppellia) tenax, Ruppell.
Cancer tenaz, Riippell, 24 Krabben roth. Meer., p. 11, pl. iii. fig. 1, pl. vi. fig. 5.
Endora tenazx, De Haan, Faun. Japon. Crust., p. 22.
Ruppellia tenaz, Milne Edwards, Hist. Nat. Crust. I. 421: Kossmann, Reise
roth. Meer., Crust., p. 40.
Carapace transversely oblate-oval, two-thirds as long as broad,
rugulose and granular antero-laterally, smooth to the naked eye else-
where. Gastric region well demarcated in its anterior two-thirds, and
broken up into five incompletely separated but rather convex lobules:
branchio-hepatic regions divided into two transverse somewhat convex
areas, independent of the rugosities inside the antero-lateral margin.
Front sunk below the level of the orbits, cut into four equidistant
rounded granular teeth. Supra-orbital margin tumid, well delimited
from carapace, with two distinct grooves near the outer angle. Infra-
orbital margin separated from the supra-orbital by a notch, deeply
concave between the prominent dentiform internal and external angles.
Antero-lateral border granular, cut into five teeth, the first four of
which are broad and anteriorly-acuminate, the fifth tuberculiform.
Chelipeds markedly unequal; upper and outer surfaces of wrist
very finely granular, upper and more or less of outer surface of hand
granular and studded with larger pustulous granules: inner angle of
wrist bluntly bicuspid: fingers pointed, those of smaller hand long and
rather slender, as in typical Ozius; those of larger hand stout, the
movable finger with a huge tooth at base, as in Indian species of
Ozius. ,
Legs stout, finely granular under a lens, but smooth to naked eye ;
the dactyli covered with velvet up to the claw.
Colours in spirit reddish yellow, the reddish tinge darkest on
carapace.
In the Indian Museum is a fine specimen from the Mekran (Balu-
chistan) coast.
188 A. Aleock— Careinological Fauna of India. [No. 1,
109. Ozius (Huruppellia) annulipes, Edw.
Ruppellia annulipes, Milne Edwards, Hist. Nat. Crust. I. 422: Dana, U.S. Expl.
Exp. Crust. pt. I. p. 246, pl. xiv. figs. 4a—-c: Stimpson, Proc. Acad. Nat. Sci. Phila.
1858, p. 37: A. Milne Edwards, Nouv. Archiv. du Mus. IV. 1868, p. 71: Haswell,
Cat. Austral. Crust. p. 73: Ortmann, Zool. Jahrb,, Syst., VII. 1893-94, p. A479:
Whitelegge, Mem. Austral. Mus. ITI. 1897, p. 137.
Euruppellia annulipes, Miers, Zool. H. M. 8. Alert, pp. 517, 523: de Man, Archiv.
fur Naturges. LIIT. 1887, i. p. 293, pl. xi. fig. 4 (hand only).
Closely resembles Ozius (Huruppellia) tenax, but differs as fol-
lows:—(1) the front is cut into four broader, shallower and much less
prominent teeth: (2) the five teeth of the antero-lateral border are
much sharper, and the margin of the first three is sharp and crest-like :
(3) the supra-orbital margin is, practically, entire, the grooves near the
outer angle being hardly visible even with a lens: (4) the infra-orbital
margin is separated from the supra-orbital only by a shallow groove, is
not concave, and has its outer angle hardly prominent.
In the Indian Museum is a specimen from Muscat (besides one
from Samoa).
BaPTozivs, n. gen.
Ruppellia (part) Milne Edwards, Hist. Nat. Crust. I. 420.
Type Roupprniia vinosa, Edw. (Op. cit. I. 422).
Carapace broad, transversely oval, moderately convex fore and aft,
slightly so from side to side, with no indication of regions,
Front very broad, about two-fifths the greatest breadth of the
carapace, obliquely deflexed, with a thin almost straight edge.
Antero-lateral border short, not two-thirds the length of the
postero-lateral, thin, cut into four sharp-edged teeth.
Orbits large, with a sharp, prominent, entire edge: the upper and
lower inner angles are in contact, so as to completely exclude the anten-
nary flagellum.
Antennules folding nearly ‘transversely, the inter-antennulary
septum very broad.
Basal antennal joint massive; the flagellum of good length (about
three-quarters the major diameter of the large orbit), lodged beneath
the front and quite outside the orbital wall.
The crests of the endostome that define the expiratory canals are
very strong, and a permanent orifice is formed not, as in Ozius, by a
notch in the anterior border of the merus of the external maxillipeds —
for the anterior border of the merus of the external maxillipeds is
1898. | A. Aleock — Carcinological Fauna of India. 189
entire— but by a deep emargination of the prolonged foliaceous oper-
cular process of the first maxillipeds.
Chelipeds fairly massive, unequal in both sexes; fingers of good
length, pointed.
Abdomen of male with all seven segments distinct.
110. Baptozius vinosus, (Kdw.)
Ruppellia vinosa, Milne Edwards, Hist, Nat. Crust. I. 422.
Eurwppellia vinosa, de Man, in Weber’s Zool. Ergebn. Niederl. Ost-Ind. IT.
1892, p. 278, pl. i. fig. 1.
Carapace broad, transversely oval, with a shiny frosted surface due
to extremely close fine granulation, the granules becoming vesiculous
and plainly visible to the naked eye near the frontal, orbital, and lateral
borders.
The orbits are marked off by a fine groove, a short shallow
A-shaped groove bisects the front and ends on the anterior part of the
gastric region, and a fine sinuous crease passes from the interval
between the third and fourth tooth of the antero-lateral margin inwards
towards the gastric region; otherwise the surface of the carapace is
unbroken.
Front nearly two-fifths the greatest breadth of the carapace, almost
straight, with a fine double edge, the upper rim of which runs on to the
orbit while the lower turns obliquely downwards to rest on the basal
antennal joint, — both rims finely beaded.
The antero-lateral borders are cut into four thin sharp-edged teeth,
the first three of which are somewhat angular, while the last is elegantly
procurved : the edges of all are finely beaded.
The orbits are large and almost subtubular: the finely-beaded edge
is entire, and the inner angle of the lower border is bluntly prominent :
quite inside the orbit, where the cornea comes into contact, is an elegant
fringe of eye-lashes.
The upper and outer surfaces of the wrists and hands are finely
frosted: the inner angle of the wrist has the form of a sharp spine.
The last three joints of the legs are more or less covered with a
harsh tomentum, thickest along the upper surface.
Colours in spirit: carapace dark purple above, dark greenish below,
legs greenish, chelipeds greenish daubed with red and purple, fingers
red. In very old spirit specimens the carapace and chelipeds are of a
rosy madder.
In the Indian Museum are 4 specimens from the Andamans and
one from an unrecorded (Indian) locality.
190 A. Alcock — Carcinological Fauna of India. [ No. 1,
Sub-family VI. Pitomnina.
Alliance I. Piluwmnoida.
Pilumnus. Actumnus.
Pitumnus, Leach.
Pilumnus, Leach, Trans. Linn. Soc. XI. 1815, p. 321; and Malac. Podophth.
Brit.: Latreille, Encycl. Meth. X. p. 124: Desmarest, Consid. Gen. Crust. p. 111 :
De Haan, Faun. Japon. Crust. p. 19: Milne Edwards, Hist. Nat. Crust. I. 415:
Dana, Silliman’s Amer. Journ. Sci. and Arts, (2) XII. 1851, p. 127, and U. S. Expl.
Exp. Crust. p. 229: Milne Edwards, Ann. Sci. Nat. Zool. (4) XX. 1863, p. 285 :
Kossmann, Reise roth. Meer. p. 37: Milne Edwards, Miss. Sci. Mex. Crust. p. 280:
Miers, Challenger Brachyura, p. 145.
In the numerous species of this genus the carapace and legs are
generally thickly covered with hair.
Carapace transversely oval or subquadrilateral, declivous anteriorly ,
flat posteriorly, not greatly broader than long; the regions, as a rule,
but moderately plainly demarcated and areolated.
Antero-lateral borders not longer, but commonly shorter, than the
postero-lateral, and cut into teeth which, very commonly, are spiniform.
The front is usually about a third the greatest breadth of the cara-
pace, but is sometimes broader: it is cut into two lobes, the outer angle
of each of which commonly forms an independent dentiform or spini-
form lobule separated from the supra-orbital angle by a groove or notch.
The orbits generally have a gap or fissure just below the outer
angle, and one or two gaps or notches in the upper border: the imner
lower orbital angle is commonly sharp and prominent, The eyestalks
are moderately long and slender.
The antennules fold transversely. The basal antennal joint is
short, either not quite touching the front, or just touching it by its
inner angle; the flagellum, which is planted in the orbital hiatus, is
long, usually very much more than the major diameter of the orbit.
The ridges of the endostome, defining the expiratory channels, are
usually plain but not very high: the anterior border of the merus of the
external maxillipeds is almost transverse and is not notched.
The chelipeds are stout, the fingers coarse, short and pointed,
Legs usually stout and of moderate length.
The abdomen of the male consists of seven separate segments.
Key to the Indian species of Pilumnus.
1. Abnormal species :—
Carapace and chelipeds quite smooth and devoid of hair P. levis.
Carapace covered with symmetrically disposed, raised,
curved or sinwated TIGER: slsee fh dedicvese! | thi eeeha athe vas ewiee P, labyrintiices.
1898. | A. Alcock — Carcinological Fauna of India.
2. Normal species in which the carapace (like the legs and
the greater part of the chelipeds) is covered with a more
or less thick coat of hair, and is without raised ridges :—
I. Front about a third the greatest breadth of the
carapace :—
i. Carapace declivous anteriorly, flat posterior-
ly ; the outer orbital angle is not a spine,
though it may be sharp :—
1. Upper margin of orbit with two, very
distinct, triangular gaps or notches :—
a. Asubhepatic spine just below
outer orbital angle...... weadais canes
b. No subhepatic spine :—
#. Free edge of front, and
upper margin of orbit, finely
denticulate: front very pro-
191
P. vespertilio.
WAINGWG | odeas dees apvindddpocess Ls LONGICOTNIS.
y» Free edge of front, and up-
per orbital margin, smooth
or nearly so: front not
PP OURIIONG Sia cies sax par eee von
2. Upper margin of orbit with one or
both of the two notches indistinct or
absent :—
a. Regions and areolz of carapace
convex, uniformly granular, and
separated by smooth, deepish,
clean-cut grooves ..........
b. Regions etc. of carapace faintly
demarcated and not uniformly
granular :—
xz. Notches in the denticulated
upper orbital margin faint
but distinguishable .........
y. Only one notch in the
smooth upper orbital mar-
gin, and that faint: legs
long and slender ......e00.0.
ii. Carapace uniformly convex, or globose; the
outer orbital angle isa spine like those of
the antero-lateral border :—
1. Regions of carapace fairly distinct :
whole outer surface of larger hand
covered with saa prominent spine-
like tubercles..
2. Regions of carapace faints tafoae aah
of outer surface of hand smooth......
II. Front nearer half than a third the greatest
breadth of the carapace, nearly straight, finely
denticulated, emarginate in the middle line..,.,....
P. andersoni.
. P. cerulescens.
P. sluiteri.
P. cursor.
. P. dorsipes.
P. hirsutus.
P. dehaanii.
192 A. Alcock — Carcinological Fauna of India. [No. 1,
111. Pilumnus vespertilio, Fabr.
Cancer vespertilio, Fabricius, Ent. Syst. II, 463, and Suppl. p. 338.
Pilumnus vespertilio, Desmarest, Consid. Gen. Crust. p. 112: Latreille, Encyc.
Meth. X. p. 125: Milne Edwards, Hist. Nat. Crust. I. 418, and in Cuvier’s Régne
An., Crust, pl. xiv. fig. 8: Dana, U. S. Expl. Exp. Crust. pt. I. p. 236: Heller,
SB. Ak. Wien, XLIII. 1861, p. 343: A. Milne Edwards, Nouv. Archiv. du Mus. IX.
1873, p. 242: Miers, Crust. New Zealand, p. 19; and Ann. Mag. Nat. Hist. (5) V.
1880, p. 234; and Zool. H. M.S. Alert, pp. 183, 219: Tozzetti, Magenta Crust. p. 55,
pl. iv. figs. 25, 27, 32: Hilgendorf, MB. Ak. Berl. 1878, p. 793: E. Nauck, Zeits.
Wiss. Zool. XXXIV. 1880, p. 53 (gastric teeth): Richters in Mobius Meeresf. Maurit.
p- 148: Haswell, Cat. Austral. Crust. p. 65: Filhol, Crust. New Zealand, p. 374,
pl. xlv. fig. 5: de Man, Journ. Linn. Soc., Zool., XXII. 1587-88, p. 58 ; and Archiv.
fiir Nat. LILI. 1887, i. p. 295; and in Weber’s Zool. Ergebn. Niederl. Ost-Ind. II.
1892, p. 288; and Zool. Jahrb., Syst., VIII. 1894-95, p. 537: Cano, Boll. Soc. Nat.
Napol. II. 1889, p. 206: A. O. Walker, Journ. Linn. Soc., Zool., XX. 1886-90,
p. 110: J. R. Henderson, Trans. Linn. Soc., Zool. (2) V. 1893, p. 365: Ortmann,
Zool. Jahrb., Syst., VII. 1893-94, pp. 436, 438, and in Semon’s Forschungsr. (Jena.
Denk. VIII) Crust. p. 49: Zehntner, Rev. Suisse Zool. II. 1894. p. 154.
Pilumnus ursuluws, Adams and White, Samarang Crust. p. 45, pl. ix. fig. 6:
Hess, Archiv. fir Nat. XXXI. 1865, i. pp. 137, 171, pl. vi. fig. 2: Kossmann, Reise
roth. Meer., Crust. p. 39: F. Muller, Verh. Ges. Basel, VIII. 1886, p. 475.
Jarapace, legs and chelipeds (with the exception of the fingers and
the lower corner and lower border of the hand, which are bare) entirely
concealed by a thick, dark, shaggy coat of coarse, tufted and somewhat
matted hair. The hairs are of two kinds, longer and shorter, the longer
being most numerous on the legs and on the borders of the carapace.
The following description (and the descriptions of all the species men-
tioned in this paper) applies to the denuded animal.
Carapace transversely oval, nearly 2 as long as broad, flat pos-
teriorly, a good deal deflexed anteriorly, the regions fairly distinctly
delimited and areolated, the surface studded with small well-separated
clusters of granules, from which the hairs spring.
Front obliquely deflexed, about a third the greatest breadth of the
carapace, cut into two lobes, each of which consists of a large prominent
convex inner division and a small receding semi-independent, but not
dentiform, outer angle, lying nearly in front of the inner upper angle of
the orbit.
The orbital margins, like the edge of the front, are smooth or
obscurely crenulate; in the upper margin are two broad triangular
gaps: the outer angle of the orbit is sharp but not spiniform, and imme-
diately below it is a fissure or gap in the infra-orbital margin.
The antero-lateral border is a little shorter than the postero-lateral,
and is cut into three spiniform teeth, besides which there is a sub-
hepatic denticle behind and below the outer orbital angle.
1898. | A. Aleock—Carcinological Fauna of India. 193
The chelipeds are unequal: the inner angle of the wrist may be
sharp, but is never spiniform: the upper and outer surfaces of the
wrists, of the smaller hand, and of all but the lower border and lower
outer corner of the larger hand (which is quite bare and usually quite
smooth) are covered with clusters of granules, some of which, on the
smaller hand—and sometimes also on the larger hand —are arranged
in longitudinal series,
The carpopodites and propodites of all the legs, and the meropodites
also of the last pair, have the anterior and dorsal aspects granular. The
longest legs are not much more than half again as long as the carapace.
In the Indian Museum are 64 specimens, chiefly from the Anda-
mans, but also from Mergui and Palk Straits; (besides 10 specimens
from other parts of the Indo-Pacific). i
Pilumnus vespertilio, var.
Differs from the above only in having (1) the fur stiff, fine, bristly,
and golden-yellow in colour, and (2) the whole of the outer surface —
but not the lower border—of the larger hand granular.
In the Indian Museum are 9 specimens from Karachi and 1 from
Tavoy.
112. Pilumnus longicornis, Hilgendorf.
Pilumnus longicornis, Hilgendorf, MB. Ak. Berl., 1878, p. 794, pl. i. figs. 8, 9.
Carapace covered with a fine and very short fur, amid which —
especially anteriorly —are numerous long silky bristles. Legs and
chelipeds — except the larger hand, the greater part of which is quite
bare — covered with similar fur and fringed with similar bristles.
Carapace somewhat quadrilateral or hexagonal, about } as long as
broad, anteriorly deflexed, posteriorly flat; the regions fairly distinctly
defined and areolated, the surface granular near the frontal and antero-
lateral margins, elsewhere smooth to the naked eye.
Front obliquely deflexed, about a third the greatest breadth of the
carapace, deeply cut into two lobes, each of which consists of a promi-
nent angularly-convex inner portion and an independent spiniform outer
angle; the free edge finely and evenly denticulate.
Two triangular gaps in the finely denticulated upper orbital margin
and a fissure in the denticulated lower margin, just below the outer
angle, which is not dentiform or very conspicuous.
Antero-lateral margin a good deal shorter than the postero-lateral,
cut into three longish procurved spiniform teeth the bases of which are
granular. No denticle—at most only a slightly-enlarged granule—
below the outer angle of the orbit.
J. 11. 25
194 A. Alcock —Carcinological Fauna of India. [No. I,
Antennary flagellum considerably more than half the greatest
length of the carapace, fringed with some long silky hairs.
Chelipeds very unequal; anterior border of ischium and arm spinu-
late or spinate, both the other borders of the arm spinulate or granular ;
upper and outer surfaces of wrist sharply granular, the inner angle of
the wrist prolonged into a stout spine; the whole upper outer and
lower surfaces of the smaller hand sharply granular, with several rows
of enlarged spiniform granules, fingers of smaller hand fluted; the
larger hand and fingers are smooth, except for a granular patch quite
at the near end of the outer surface and extending a variable distance
along the upper border of the hand, and for a small patch of granules
at the base of the dactylus.
The upper border of the meropodites of the legs is usually sharply
spinate.
Colours in spirit yellow, fingers dark brown.
In the Indian Museum are 21 specimens, from Mekran, Karachi,
Bombay, Nicobars, and Malacca Strait.
113. Pilwmnus anderson, de Man.
Pilumnus andersoni, de Man, Journ. Linn. Soc., Zool., XXII. 1887-88, p. 59),
pl. iii. figs. 5, 6, and Zool. Jahrb. Syst., VIII. 1895, p. 552.
Closely resembles P. longicornis from which it differs in the follow-
ing particulars :—
(1) the carapace appears broader owing to the front being less
prominent :
(2) the free edge of the front and the upper margin of the orbit
are nearly or quite smooth; the lobes of the front are much less _pro-
minent, the notch between them is not so wide and deep, and the outer
angles are dentiform, not spiniform:
(3) the outer angle of the orbit is sharper :
(4) the granular patch at the base and along the upper border of
larger hand is larger.
In the Indian Museum are 5 specimens, from Mergui, Ceylon,
Karachi (and 5 from Gaspar Strait).
14, Pilwmnus sbuitert, de Man.
Pilumnus sluiteri, de Man, in Weber's Zool. Ergebn. Niederl. Ost-Ind. II.
1892, p. 283, pl. i. fig. 2: Ortmann, Zool. Jahrb., Syst., VII. 1893-94; pp. 486, 438.
Pilumnus forskalii, de Man (nec Edw.), Archiv. fir Naturges. LIII. 1887. i
p. 295, pl. xii. fig. 1.
Carapace, legs and chelipeds (except the fingers} covered with a
1898. } A. Alcock —Carcinological Fauna of India. 195
harsh coat of short bristles with longer bristles interspersed, the latter
being most numerous on the legs.
Carapace about £ as long as broad, deflexed anteriorly, nearly flat
in the postericr two-thirds, the regions fairly distinctly delimited, the
surface rather profusely studded with little pits, from which the tufts
of bristles arise; some granules near the antero-lateral borders and on
the front part of the gastric region.
Front cut inte two lobes, each of which is again subdivided by a
deep triangular gap into a large square-cut internal lobe and an acute
triangular external lobule.
Orbital margin granular: there are two gaps in the upper margin,
but the inner one is narrow and indistinct ; there is also a small gap just
below the outer angle of the orbit, which is not very prominent.
Antero-lateral margin not much shorter than the postero-lateral,
cut into three somewhat granular spiniform teeth. No tooth below the
outer angle of the orbit.
Antennary flagellum not quite a third the length of the carapace,
not fringed with hairs, though there may be one or two at its base.
Chelipeds very unequal: upper and outer surfaces of both wrists
and hands and bases of dactyli covered with granules or small pearly
tubercles, which are larger and more numerous and more prominent on
the hands than on the wrists; sometimes a small patch of granules on
inner surface of hands: inner angle of wrists strongly pronounced and
dentiform. 7
Legs stout, the longest pair are about two-thirds again as long as
the carapace.
Colours in spirit: carapace yellow copiously overspread with brick-
red, chelipeds and legs yellow blotched and sometimes banded with terra-
cotta-red.
In the Indian Museum are 7 fine specimens from the Andamans
(besides one from Samoa).
115. ? Pilumnus cursor, A. Milne Edwards.
? Pilumnus cursor, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873, p. 244,
pl. ix. fig. 4: Haswell, Cat. Austral. Crust. p. 67: Miers, Zool. H. M. S. Alert,
pp. 183, 223: de Man, Archiv. fiir Naturges. LIII. 1887, i. p. 299.
Carapace etc. covered with a short fur with long hairs interspersed,
the latter most numerous on the legs.
Carapace subquadrilateral, flat in the posterior half or more,
declivous anteriorly, the regions faintly marked, finely and sparsely
granular.
Front nearly two-fifths the greatest breadth of the carapace, not
196 A. Alcock —Carcinological Fauna of India. [No. 1,
very prominent, divided into two lobes, each of which consists of a
convex inner part and an independent though not very prominent exter-
nal angle.
The upper orbital margin is little prominent and has only one
notch and that indistinct: outer orbital angle not prominent.
Antero-lateral border very much shorter than the postero-lateral,
cut into three sharp teeth: no subhepatic tooth.
Chelipeds unequal: inner angle of wrists sharply pronounced,
upper and outer surfaces of hands granular. :
Legs slender, the longest pair are more than twice the length of
the carapace.
Colours in spirit, carapace reddish-yellow, legs yellow.
In the Indian Museum is a single specimen, from the Andamans.
T identify this species with P. cursor on account of the long slender
legs, the subquadrilateral carapace, the very short antero-lateral borders,
and the broad front.
116. ? Pilumnus cerulescens, A. M. Edw.
? Pilumnus cerulescens, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873,
p. 242, pl. ix. fig. 3: L. Zehntner, Rev. Suisse Zool. II. 1894, p. 153.
Carapace etc. covered with short fur, with long hairs interspersed,
about = as long as broad, subquadrilateral, convex in anterior half, flat
posteriorly: the regions very distinctly defined by well cut grooves, the
areols convex and studded with granules of good size.
Front a third the greatest breadth of the carapace, deflexed, cut
into two lobes the outer angles of each of which form independent
dentiform lobes.
The upper orbital margin shows very faint traces of two shallow
notches: a small triangular gap below the sharp, but non-spiniform,
outer orbital angle.
Antero-lateral borders a good deal shorter than the postero-lateral,
cut into three sharp teeth, in addition to which there is a small denticle
behind and below the outer orbital angle.
Chelipeds unequal; upper and outer surfaces of wrists and of both
hands closely and sharply granular, fingers very short, inner angle of
wrists dentiform.
Legs stout, the longest pair not much more than two-thirds again
the length of the carapace.
Colours in spirit, dull blue with a brownish tinge in places, fingers
blackish brown.
In the Indian Museum isa single specimen from the Andamans.
I judge this species to be P. cerulescens by the Xanthodes-like
form and sculpture of the carapace mentioned by Milne Edwards.
1898. ] A. Aleock—Carcinological Fauna of India. 197
117. ? Pilumnus hirsutus, Stimpson.
? Pilumnus hirsutus, Stimpson, Proc. Ac. Nat. Sci. Philad., 1858, p. 37: Miers,
P. Z. 8. 1879, pp. 20, 31: Haswell, Cat. Austral. Crust. p. 69: Ortmann, Zool.
Jahrb., Syst., VII. 1893-94, pp. 435, 437.
Carapace etc. covered with stiff hairs of two kinds—long: and
short, the former most numerous on the legs.
Carapace nearly ~ as long as broad, convex in both directions,
smooth when denuded, the regions hardly marked.
Front about a third the greatest breadth of the carapace, deflexed,
cut into two lobes much like those of P. vespertilio in shape.
Upper orbital margin smooth, with two very faint and shallow
notches ; lower margin denticulate, witha gap just below the outer
angle. No subhepatic tooth.
Antero-lateral border much shorter than the postero-lateral, with
4 spiniform teeth, one of which is the orbital angle,
Antennary flagellum of moderate length, without hairs, except at
base.
Chelipeds unequal; borders of arm finely granular or denticulate,
wrists with the upper and outer surfaces rough and the inner angle
sharply pronounced; lower part of outer surface of larger hand smooth,
the rest of this surface—like that of the smaller hand—sharply granu-
lar, the granules becoming spiniform towards the upper border.
Legs rather slender, the longest pair about two-thirds again as
long as the carapace.
Colours in spirit, yellow, fingers light brown.
In the Indian Museum are 11 specimens, from the Andamans,
Mergui, and the Malacca Str.
118. ? Pilumnus dorsipes, Stimpson.
? Pilumnus dorsipes, Stimpson, Proc. Ac. Nat. Sci. Philad., 1858, p. 37.
Carapace globose, extremely deep, not very much broader than long,
covered—like the chelipeds and legs—with soft, though stiff, hair
fairly well areolated, finely granular under a lens.
Front about a third the greatest breadth of the carapace, cut into
two convex, rounded, finely denticulate lobes, of which the outer angles
form dentiform lobules.
Upper orbital margin not prominent, the two notches are very
faint and shallow (especially the inner one), but are recognizable: lower
orbital margin with a deep narrow cleft just below the outer angle.
Antero-lateral margin cut into four denticulate spiniform teeth,
one of which is the outer orbital angle. No subhepatic tooth.
?
198 A. Aleock —Carcinological Fauna of India. [No. ],
Antennulary flagellum about a third the length of the carapace,
not hairy, except at base.
Chelipeds unequal, both hands covered, on the outer surface and
upper and lower borders, with prominent spiniform granules, which
also extend some way along both fingers.
Legs rather short, the longest pair being about half again as long
as the carapace.
Colours in spirit yellow.
The body is of such depth that the last pair of legs, even in the
male, lie, in the normal inclination of the body vertically over the first
pair.
In the Indian Museum is a single male from the Andamans.
119. Pilumnus de Haanii, Miers.
Pilumnus de Haanii, Miers, P.Z.S. 1879, pp. 20, 32; and Challenger Brachyura
p. 155, pl. xiv. fig. 1: A. O. Walker, Journ. Linn. Soc., Zool., XX. 1886-1890, p, 110,
Carapace covered with a very fine and short, but dense, fur: legs
and chelipeds with a similar fur mixed with long fine hairs on outer
surface of hands and on borders of legs.
Carapace transversely oval, not three-quarters as long as broad,
the regions (when carapace is denuded) fairly well marked and areolat-
ed, granular towards the antero-lateral margins and near the front.
The front is nearer half than two-fifths the greatest breadth of the
carapace, is nearly straight, not at all prominent, is finely denticulate,
and emarginate in the middle line.
Orbital margin very finely denticulate, the upper border with two
very inconspicuous notches, a fissure below the acute outer orbital
angle.
Antero-lateral border a good deal shorter than the postero-lateral,
cut into three shallow anteriorly-acuminate teeth.
Chelipeds unequal: the outer surface of the wrists with a few
granules anteriorly and along the inner border: upper and outer sur-
faces of hands closely covered with acute spiniform tubercles which
also extend far along the fingers.
Legs stout, unarmed.
Colours in spirit, golden yellow.
In the Indian Museum are 7 specimens from Palk Str. 28 speci-
mens from off Ceylon, 263 to 34 fms., only differ from the typical form
in having the front more deeply emarginate in the middle line.
This species, but for the broader straighter front, and for the
broader carapace, more nearly resembles an Actwmnus than a Pilumnus,
1898. ] A. Alcock—Carcinological Fauna of India. 199
120. Pilumnus labyrinthicus, Miers.
Pilumnus labyrinthicus, Miers, Zool. H. M. §. ‘‘ Alert,’’ pp. 183 and 224, pl. xxii.,
fig. C: and “Challenger” Brachyura, p. 161: A. O. Walker, Journ. Linn. Soc.
Zool. XX, 1836-1890, p. 110; Henderson, Trans. Linn. Soc. Zool. (2), V, 1893, p.
365.
“Tn this curious form the surface of the carapace is everywhere
covered with raised curved or sinuated ridges, which are separated by
wide depressions ; the body and legs are covered with a dense close
brown pubescence; from most of the ridges and from the teeth of the
antero-lateral margins of the carapace spring longer sete, and the
margins of the ambulatory legs are also fringed with longer hairs.
The frontal lobes, which are scarcely separated as usual by a median
notch, are rather broad, straight, and but little prominent ; the antero-
lateral margins are somewhat shorter than the postero-lateral, and are
armed with three distinct teeth, that of the exterior orbital angle being
obsolete. The orbital margin is somewhat thickened ; the epistoma
rather longer in proportion to its breadth than is usual. The basal
antennal joint is short, scarcely attaining to the sub-frontal process,
and not nearly reaching to the apex of the very prominent lobe at the
inner suborbital angle. The chelipeds are rather small and (like the
carapace) are densely pubescent, besides being clothed with longer
hairs; the outer surface of the wrist or carpus is tuberculated beneath
the hairy coat; the palm is clothed externally with long dense hairs ;
the upper margin of the palm bears three distinct tubercles; the fingers
are slaty coloured, dentated on their inner margins and acute at their
apices. The ambulatory legs are densely hairy and of moderate
length.”
Not in the Indian Museum collection.
121. Pilwmnus (?) levis, Dana.
Pilumnus levis, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 82, and U. S. Expl.
Expd. Crust. pt. i, p. 238; de Man, Journ. Linn. Soc. Zool. XXII, 1887-1888, p. 66,
pl. iv. figs. 1 and 2: and Zool. Jahrb. Syst. VIII. 1895, p, 553.
“ Near P. levimanus, but broader. Carapace smooth and shining,
not areolate, rather convex: front emarginate, antero-lateral margin
three-toothed, the teeth minute and like spines, the posterior much the
smallest, outer angle of orbit not raised into a tooth. Anterior feet very
unequal, carpus smooth, not even faint tuberculate ; larger hand wholly
smooth, smaller sparsely hirsute, not at all tuberculate. Posterior
eight feet slender, somewhat hirsute.”
A single specimen from Mergui. It appears to me doubtful that
this species belongs to the genus Piluwmnus.
200 A. Alcock —Carcinological Fauna of India. [No. J,
122. Piluwmnus seminudus, Miers.
Pilumnus seminudus, Miers, Zool. H. M.S. “ Alert” pp. 183 and 222, pl. xxi.
fig. C: “Challenger” Brachyura, p. 161: de Man, Journ. Linn, Soc. Zool. XXII,
1887-1888, p. 65.
“This species resembles P. semilanatus in having the gastric,
cardiac, and branchial regions of the carapace smooth and naked; but
it may be at once distinguished by the following characters :—The
carapace is broader in proportion to its length, and its anterior parts
clothed with a close velvety pubescence, which also extends over the
upper and outer surface of the wrist and palm of the chelipeds; the
two posterior teeth of the antero-lateral margins are more distinctly
spiniform, the basal antennal joint does not nearly reach to the sub-
frontal process; the granulations of the wrist and palm are much more
inconspicuous, those of the outer surface of the palm appear, through
the pubescence, to be arranged in four distinct longitudinal series ;
the ambulatory legs are slenderer.”’
A single small specimen in the Indian Museum, from Mergai, has
been referred by Dr. de Man to this species.
Actumnvs, Dana,
Actumnus: Dana, Silliman’s Amer. Journ. Sci. and Arts, (2) XII. 1851, p. 128;
and Proc. Acad. Nat. Sci. Philad. VI. 1852, p. 82; and U.S. Expl. Exp. Crust. pt. i.
p. 243: A. Milne Edwards, Nouy. Archiv. du Mus. I. 1865, p. 284.
Carapace very little broader than long, convex, fairly well or very
well areolated: the antero-lateral borders short, cnt into teeth; the
postero-lateral longer than the antero-lateral, concave.
Front from about a third to about two-fifths the greatest breadth
of the carapace ; cleft or notched in the middle line, or bilobed with
the outer angles independent, usually separated from the supra-orbital
angle by a notch or groove.
Orbits rather large, with one or two notches or fissures or suture-
lines (which often, however, are indistinct) in the upper margin, and
one (often, also, very indistinct) in the lower margin near the outer
angle. The inner lower angle of the orbit is prominent, and often
comes so near to the supra-orbital angle as to almost exclude the
antennary flagellum from the actual orbital hiatus.
The basal antennal joint touches or nearly touches the front; the
flagellum which is of moderate length (longer than the major diameter
of the orbit) sometimes springs from the orbital hiatus, but is some-
times almost excluded from the hiatus.
The crests of the endostome, defining the expiratory channels, are
s
1898. | A. Alcock —Careinological Fauna of India. 201
not very strong, and the anterior border of the merus of the external
maxillipeds is not notched.
The chelipeds are stout, and are unequal in both sexes: the fingers,
which are short and stout, are commonly defined as spoon-shaped at
tip: they are not really so, but have the tips curved and blunt pointed.
The legs are stout and not very long.
The abdomea of the male consists of seven separate segments, and
the first tergum is unusually long and narrow in all the typical species.
Most of the species of this genus, but not all, are densely tomen-
tese : all, however, ‘have hairy or tomentose legs.
Key to the Indian species of Actumuus.
I. Carapace tomentose :—
i, Carapace areolated : front separated from the supra-
orbital angle by a notch ; antero-lateral border cut
into 3 teeth (exclusive of outer angle of orbit):—
1, Front normally bilobed: supra-orbital margin
granular, with two distinct notches :—
a, Carapace moderately convex, rather
faintly areolate ; outer angles of front
hardly independent ...........sseesesesovee A. Lomentosus.
&. Carapace strongly convex, strongly
areolate ; the outer angles of the front
i are small distinct little lobules :—
x. Lateral gastric areole semi-
‘ ISI Nok ec ccudmageccskvdncrs teh, SOM OKs
y. Lateral Reais rete tenehe nod A, verrucosus.
2. Front broadly triangular, with a deep
button-hole cleft (the hole at posterior end)
in the middle line: supra-orbital margin
thin sharp, with a single deep very narrow
fissure........ siebaee’ age Sabiageedsaabaed MNduas veya biees aden A pUSOELVOTLE,
ii. Carapace not areolated ; foo ait aieenied feorin
the supra-orbital angle; antero-lateral border with
7 spinuliform granules (3 pairs) and an odd one
RUMORS so csiisnigen ace «ss seacedoanedmtenttartathsens thaval de. /tae COCQSs
iI, Carapace perfectly bare :—
i. Carapace not areolate, front broadly bilobed ; legs
Gabo gt PATO 2205561..)0seigacantdrery ts dccccccccsssooe Ae MUGUS.
ii, Carapace very Haltoatle Bethe ect with two
median lobes and two (external) lobules: legs
tomentose :—
1. Surface of carapace (and of parts of chelipeds)
formed of a mosaic of smooth flat polygonal
granules in the closest contact .....6..-s0....0. A, tessellatus.
2. Surface of carapace, etc. covered with sharp
crystalline granules in the closest contact... A. arbutum,
wit. 26
202 A. Aleock —Carcinological Fauna of India. (No. 1,
123. Actumnus tomentosus, Dana.
Actumnus tomentosus, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 82; and U.S.
Expl. Exp. Crust. pt. i. p. 243, pl. xiv. figs. 2a-c: A. Milne Edwards, Nouv. Archiv.
du Mus. I. 1865, p. 285, and 1X. 1873, p. 194: ? Tozzetti, Magenta Crust. +. 56, pl. iv.
figs, 22, 24, 26, 29: Haswell, Cat. Austral. Crust. p. 73: Htheridge, Mem. Austral.
Mus. 1889, pp. 34, 36.
Carapace subcircular, rather more than { as long. as_ broad,
moderately convex, covered with a very dense short smooth tomentum.
Much the same tomentum covers the exposed surfaces of the legs and
chelipeds (except the lower and distal part of the outer surfaces of the
hands), and the legs are also fringed with long fine hairs.
The regions are fairly well delimited and areolated, the areole
being moderately convex: on the undenuded carapace the areole are
faint. ;
Front about two-fifths the greatest breadth of the carapace, cut
into two finely denticulated lobes, the outer angles of each of which,
though sharply separated from the supra-orbital margin, do not form
distinct lobules.
Orbital margin finely denticulate, the lower more markedly so than
the upper; in the upper margin are two broad notches, the outer the
more distinct ; in the lower margin, just below the outer angle, is a
narrow fissure ; outer orbital angle dentiform.
Antero-lateral borders about two-thirds the length of the concave
postero-lateral, very regularly cut into 3 uniform teeth similar to the
outer orbital angle.
Chelipeds unequal: arm smooth; inner angle of wrists sharp,
their inner border finely beaded, a few scattered granules on their
upper and outer surfaces; upper and outer surfaces of hands covered
with pearly granules which become obsolescent or obsolete near the
lower border of the larger hand; dactyli longitudinally grooved, beaded
at base.
Denuded legs nearly smooth. |
In the Indian Museum are 53 specimens, from the Andamans, the
Orissa Coast up to 30 fms., Palk str. and Cheduba.
Our specimens completely agree with Dana’s figure, and are easily
distinguished from A. setifer by the less convex and less distinctly
areolated carapace.
124. Actumnus setifer, (De Haan), A. M, Edw.
Pilumnus setifer, De Haan, Faun. Japon. Crust. p. 50, pl. iii. fig. 8 (Xantho).
Actwmnus setifer, A. Milne Edwards, Nouv. Archiv. du Mus. I. 1865, p. 287,
pl. xv. figs. 5-56: Richters in Mobius’ Meeresf. Maurit. p. 148: Miers, Zool.
1898. | A. Alcock — Carcinological Fauna of India. 203
H. M.S. Alert, pp. 183, 225, 517, 533: de Man, Journ. Linn. Soc., Zool., XXII.
1887-88, p. 47, and Archiv. fiir Naturges. LIII. 1887, i. p. 262: Walker, Journ. Linn.
Soc., Zool. XX. 1886-90, p. 110: Pocock, Ann. Mag. Nat. Hist. (6) V. 1890, p. 74:
Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 364: Ortmann, Zool., Jahrb.
Syst. VII. 1893-94. p. 474.
Closely resembles A. tomentosus, from which it is distinguished by
the following characters :—
The carapace is subglobular: the regions are very distinctly
delimited and areolated, the areole being strongly convex and often
uniformly granular.
The outer angles of the front form distinct little lobules: the
fissure in the lower orbital margin, just below the outer angle, is indis-
tinct.
The granules on the wrist are more numerous.
The more convex carapace and the more numerous and more convex
areole, at once distinguish this species.
In the Indian Museum are 32 specimens from the Andamans,
Ceylon up to 34 fms., Persian Gulf, Pedro shoal, and Mergui (besides
13 from Hongkong and 1 from Samoa).
The Indian specimens, especially those from deep water, have the
lobules of the carapace more convex than those from Hongkong. ©
125. Actumnus verrucosus, Hndrsn.
Actumnus verrucosus, Henderson, Trans. Linn. Soc. Zool. (2), V, 1893, p. 364,
“« The carapace is very convex, covered with a short brown pubes-
cence, and provided with a series of remarkable granulated lobes.
The frontal margin is granulated and four-lobed, the rounded prominent
submedian lobes separated by a narrow median fissure, the outer lobes
of small size. The antero-lateral margin has four prominent, sub-
equal, granulated or subspinose lobes, while the postero-lateral margin
is smooth and deeply excavated ; the upper orbital margin is granulated
and has two well-marked fissures. The granulated lobes on the carapace
are arranged as follows:—On the anterior gastric region, behind the
front, two pairs, of which the posterior is much larger ; on the posterior
gastric region three lobules, one median and anterior, two posterior ;
on each protogastric or lateral gastric region a peculiar V-shaped
lobule ; on the cardiac region two lobules which are slightly excavated
in the centre; on the branchial region three lobules, anterior, postero-
external (which is the largest of the three), and a postero-internal
one placed external to and between the posterior gastric and cardiac
lobules.”
“The right cheliped is slightly larger than the left in both sexes ;
204. A. Aleock — Carcinological Fauna of India. [No. l,
both are clothed with a short pubescence on the outer surface of the
carpus and hand, except towards the base of the immobile finger. The
carpus is sparingly tuberculate externally, with a sulcus running
parallel to the articulation with the hand, and separated from the latter
by a tuberculated strip: the outer surface of the hand is strongly
tuberculate, the tubercles with more or less acute apices, rather closely
crowded and without any definite arrangement. The fingers are short,
with white and obtuse tips, and the immobile one is placed in a straight
line with the lower border of the hand; the dactylus is tuberculated
superiorly on its proximal half, and a prominent tooth is present on
either finger. The ambulatory legs are simply pubescent. The abdo-
men is smooth and seven-jointed in both sexes. The external maxil-
lipeds are smooth, with a faint impressed line in the middle of the
proximal two-thirds of the ischium. The basal joint of the antennal
peduncle is joined to the sub-frontal process, and the terminal joints lie
in the orbital hiatus.”
“The largest specimen (a male) has the carapace 18°5 mm. long
and 25°3 mm. broad.”
126. <Actumnus fissifrons, n. sp.
Carapace and legs covered with a not very dense coat of hairs of
two kinds—long and short—the long hairs most numerous on the legs;
chelipeds with very little hair.
Carapace strongly convex in all directions, ? as long as broad, the
regions distinctly delimited and areolated by smooth shallow grooves,
the areole being slightly convex and more and less granular.
Front not quite a third the greatest breadth of the carapace,
deflexed, broadly triangular, the apex with a deep button-hole fissure
(the hole at the posterior end), the outer angles separated from the
supra-orbital angles by a deepish notch.
Supra-orbital margin thin, sharp, very prominent, deeply fissured
near the middle : infra-orbital margin thin, concave, fissured just below
the outer angle.
Antero-lateral margin a little shorter than the postero-lateral, cut
into three sharp-edged anteriorly-acuminate teeth (exclusive of the
outer orbital angle) : postero-lateral margin deeply concave.
Chelipeds markedly unequal: upper and outer surfaces of wrists
with a few granules, most numerous anteriorly ; upper and outer surfaces
of both hands—including a great part of the fingers—studded with
granules, of which many are enlarged conical or pearl-like, and those
along the upper border are spiniform.
Colours in spirit bright orange yellow.
1898. | A. Alcock — Carcinological Fauna of India. 205
Off Ceylon, 263-34 fathoms. Four specimens.
The carapace of the largest specimen is 21 millim. long and 28
millim. broad.
127. Actumnus tessellatus, n. sp.
Legs tomentose and hairy, chelipeds inconspicuously tomentose in
parts, carapace bare.
The entire dorsal surface of carapace, the upper and outer surfaces
of the wrists and the upper surface of the hands, have the form of an
elegant mosaic of smooth polygonal tile-like granules in the closest
possible contact everywhere.
Carapace strongly convex, # as long as broad, regions well defined
and subdivided by broad depressions, the areole strongly and somewhat
angularly convex.
Front much less than a third the greatest breadth of the carapace ;
deeply cut into two prominent subfoliaceous median lobes, each of
which is flanked externally by a small dentiform lobule.
Orbital margins smooth, not fissured, though there are narrow
inconspicuous depressions where the notches exist in other species. The
antennary flagellum springs from the orbital hiatus.
Antero-lateral margins thin, sharp, cut into 3 teeth (not including
the orbital angle) the last 2 of which are subfoliaceous : postero-lateral
margins a little longer than the antero-lateral, markedly concave.
Chelipeds little unequal : in addition to the mosaic ornamentation
there are a few scattered pustulous granules on the wrist and upper
surface of hand, and all the lower half of the outer surface of the hand
is studded with pearl-like or bead-like granules, which are also found
on the bases of the fingers.
The legs when denuded are smooth to the naked eye.
Colours in spirit: lavender grey, a good deal suffused with orange-
pink, fingers cinnamon.
— Carapace 15 millim, long, 20 millim. broad.
A male and a female from the Persian Gulf.
128. <Actwmnus arbutum, n. sp.
Legs with a somewhat scanty growth of hair not concealing their
sculpture, chelipeds slightly hirsute in places, carapace bare.
The whole dorsal surface of carapace covered with sharp angular
crystalline granules in the closest possible coutact: much the same
ornamentation is found on the upper and outer surfaces of the wrists
and on the upper surface of the hands, the lower half of the outer
surface of the hands being studded with pearly and bead-like granules.
206 A. Alcock — Carcinologicul Fauna of India. [No. ],
Carapace ? as long as broad, strongly convex, profusely foenty and
symmetrically puckered-areolate.
Front much less than a third the greatest breadth of the carapace,
shaped as in A, tessellatus, but the edges of the lobes and lobules are
crenulate.
Orbits and relations of antenne as in A. tessellatus, but the edges
of the orbits are sharply crenulate.
Autero-lateral margin cut into three sharply crenulate granular
teeth —not including the orbital angle : postero-lateral margin shorter
than the antero-lateral, concave.
Chelipeds a little unequal: fingers granular in the basal half or
more.
Carpopodites and propodites of legs, and meropodite of last pair,
sharply granular as to the dorsal surface.
Colours in spirit pink, fingers brownish,
Carapace 13°5 millim, long, 18 millim. broad.
A single male from off the coast of Sind, 51 fms.
129. Actwmnus elegans, de Man.
Actumnus elegans, de Man, Journ. Linn. Soc., Zool., XXII. 1887-88, p. 47.
Carapace and exposed surfaces of legs and chelipeds covered with
a thickish bright-yellow tomentum, with longer hairs on chelipeds and
legs and near frontal margin.
Carapace not much more than $ as long as broad, with some
scattered comparatively large granules, but with almost no indication of
regions ; convex fore and aft, slightly so from side to side.
Front about a third the greatest breadth of the carapace, broadly
triangular, notched at the apex, not separated from but confluent with
the supra-orbital angles. There is a suture line in the lower orbital
margin just below the outer angle.
Antero-lateral borders not shorter than the very concave postero-
lateral, armed with 7 acute spinuliform granules, in 3 pairs, with an odd
one between the first pair and the orbital angle.
Chelipeds unequal: the upper and outer surfaces of wrists ana
both hands, including a large part of the fingers closely studded with
conical white granules.
In the Indian Museum are 2 specimens, one from Mergui the other
from Kyuk Phyu Harbour.
This species seems to me to be better placed with Pilwmnus than
Actumnus: it and Pilumnus scabriusculus White, seem to be very closely
related.
1898. ] A. Alcock — Carcinological Fauna of India. 207
130. Actumnus nudus, A. M. Edw.
Actumnus nudus, A. Milne Edwards, Ann. Soc. Entomol. France, (4) VII. 1867
p- 265: de Man, Journ. Linn. Soc., Zool., XXII. 1887-88, p. 49, pl. ii. figs. 3, 4.
Carapace almost completely bare, legs with only a few scattered
hairs.
Carapace subcircular, convex, regions hardly indicated, studded
with pearl-shaped granules in its anterior and antero-lateral parts,
twelve of these granules are arranged in an arched line—convex for-
wards—on either side of the posterior end of the gastric region,
Front much advanced, divided into two rounded oblique lobes, the
outer angles of which are hardly separated from the supra-orbital
angles.
Antero-lateral border divided into four teeth (not including the
outer orbital angle).
Chelipeds unequal; upper and outer surfaces of hand covered with
pearly granules, which also exist on the upper surface of the wrist.
Found at Pondicherry and Mergui.
Not represented in the Indian Museum collection. ,
This species seems to me to be improperly referred to Actumnus.
Alliance II. Heteropanopioida.
Heteropanope. Kurycarcinus. Nectopanope,
Hereropanops, Stimpson, de Man.
Heteropanope, (part) Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 35.
Heteropanope, (part) A. Milne Edwards, Ann. Sci. Nat., Zool., (4) XX. 1863,
pp. 288, 289.
Pilumnopeus, (part) A. Milne Edwards, loc. cit.; and Ann. Soc. Entomol.
France, (4) VII. 1867, p. 277.
Heteropanope, de Man, Journ. Linn. Soc., Zool., XXII. 1887-88, p. 52.
Carapace moderately broad, moderately or little convex, with the
regions little or hardly demarcated. |
Antero-lateral borders shorter than the postero-lateral, cut into
four lobes or teeth, of which the first is confluent with the outer angle
of the orbit: postero-lateral borders moderately convergent, posterior
border rather long.
Front moderately broad, between a fourth and a third the greatest
breadth of the carapace, cut into two lobes, the outer angle of each of
which is dentiform and separated from the supra-orbital margin by a
notch.
A small triangular gap in the orbital margin just beneath the
outer angle. The antennules fold nearly transversely.
208 A. Alcock —Carcinological Fauna of India. [No. I,
Basal antennal joint short, not reaching the front; the flagellum,
which is about equal in length to the major diameter of the orbit,
lodged in the rather broad orbital hiatus.
The ridges of the endostome, defining the expiratory canals, are
well marked, but the anterior border of the merus of the external
maxillipeds is not notched. The buccal cavern is broader anteriorly
than posteriorly.
Chelipeds unequal in both sexes; fingers rather short, pointed,
not hollowed.
The abdomen of the male consists of seven separate segments.
Heteropanope closely resembles Panopeus (e.g. P. herbstw), but
differs in having the crests of the endostome much more distinct, and
all seven segments of the male abdomen separate.
Key to the Indian species of Heteropanope.
I. Carapace decidedly convex, both rae ate Paeut smooth
to the naked eye .....0.. scores cavesce scene enn annnE
II. Carapace almost flat, wit aaa’ natal of ‘Gihle cheliped
studded with vesiculous or pearly granules....c.sccscssecssereeses H. indica.
131. Heteropanope indica, de Man.
Heteropanope indica, de Man, Journ. Linn. Soc., Zool., XXII. 1887-88, p. 53,
pl. iii. figs. 1, 2. .
Carapace more than two-thirds as long as broad, very little convex,
surface somewhat granular and scantily tomentose near the margins.
Gastric region and its three sub-regions faintly indicated. Two series,
starting respectively from the 3rd and 4th teeth of the antero-lateral
margins, of discontinuous wavy finely granular ridges cross the
carapace transversely, fairly parallel with the common curve of the
frontal and antero-lateral borders.
The finely granular orbital margin has the two grooves near the
external angle, and the gap just below the external angle, distinct.
Antero-lateral border cut into four teeth, of which the first two are
broad thin and compressed and the last two pointed and subpyramidal ;
the edges of all are finely granular.
Chelipeds and legs more or less tomentose. Chelipeds very un-
equal; a curved spine-like tooth at distal end of upper border of arm,
and a spine at inner angle of wrist: upper and outer surface of smaller
hand and wrist studded with vesiculous granules; larger hand quite
smooth, very large, little shorter than the greatest breadth of the
carapace.
1898. | A. Aleock — Carcinological Fauna of India. 209
Colours in spirit, dull earthy brown with a greenish tinge.
In the Indian Museum is a single specimen, from Mergui.
132. Heteropanope levis (Dana),
Panopzxus levis, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 76, and U. S. Expl.
Exp. Crust. pt. I. p. 180, pl. viii. figs. 13u-c: J. E. Benedict and M. J. Rathbun,
P. U. 8. Nat. Mus, XIV. 1891, p. 380.
Carapace two-thirds as long as broad, decidedly convex fore and
aft, its surface perfectly smooth to the naked eye, and bald. The gastric
region and its three sub-regions are as faintly as possible indicated, and
the two broken series of transverse elevations present in H. indica are
also present, but are much blunter smoother and fainter. The orbits
are as in H. indica, but the margin is but microscopically granular.
The antero-lateral border is cut into four teeth, all of which are thin
and compressed, and all but the first are sharply acuminate forwards.
The chelipeds are extremely unequal, and are perfectly smooth and
bare: there is a denticle at the distal end of the upper border of the
arm, and a stout sharp tubercle (often double-crowned) at the inner
angle of the wrist, The greatest length of the larger hand, in the
male, is about equal to the greatest breadth of the carapace, and its
greatest height more than three-quarters the greatest length of the
carapace: in the female this hand is not quite so large.
The edges of the last four joints of all the legs are scantily hairy
in the male, but more profusely so in the female.
Colours in spirit ; brownish yellow or dull green.
In the Indian Museum are 17 specimens from Karachi and one
from Bombay. |
133. Heteropanope eucratoides, Stimpson.
Heteropanope eucratoides, Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 35:
de Man, Journ. Linn. Soc., Zool., XXII. 1887-88, p. 56, pl. iii. figs. 3, 4.
This species is included by de Man in the Mergui fauna. There
are no specimens in the Indian Museum. According to de Man it
chiefly differs from H. indica in having the antero-lateral margins much
shorter, and the 3rd tooth of the antero-lateral margin smaller than
any of the others,
The chelipeds have a smooth surface.
Kourycarcinus, A. Milne Edwards.
Eurycareinus, A. Milne Edwards, Ann. Soc. Entomol. France (4) VII. 1867,
p. 276.
Eurycarcinus, de Man, Journ. Linn. Soc., Zool., XXII. 1887-88, p. 43.
Carapace broad, convex, perfectly smooth, without trace of regions,
wn it. 27
210 A. Alcock — Carcinological Fauna of India. [No. 1,
Antero-lateral borders very much shorter than the postero-lateral,
cut into four lobes or teeth, of which the first is confluent with the
outer angle of the orbit: postero-lateral border moderately convergent,
posterior border rather long.
Front broadish, nearly a third the greatest breadth of the carapace,
obliquely deflexed, projecting a little beyond the orbits, straight and
square cut, commonly emarginate in the middle line.
Orbits shallow, affording little concealment to the eyes, the upper
margin entire, a gap in the lower margin, below the outer angle. The
antennules fold quite transversely.
Basal antennal joint short, not reaching the front, the flagellum,
which is long (much longer than the major diameter of the orbit),
lodged in the orbital hiatus.
The ridges of the endostome, defining the expiratory canals, are
well pronounced, but the anterior border of the merus of the external
maxillipeds is not notched. Buccal cavern wider anteriorly than
posteriorly.
Chelipeds unequal in both sexes, fingers pointed, not hollowed.
The abdomen of the male is seven-jointed.
Eurycarcinus is very closely related to Heteropanope, but is easily
distinguished by the broad smooth convex carapace, the shallow and
rather elongate orbits, and the very short antero-lateral margins.
Key to the Indian species of HKurycarcinus.
TI, Thumb of the larger cheliped with a much-enlarged tooth
at basal end :—
1, Antero-lateral border less ie 3 the length of the
postero-lateral | ©. ...:.cse+s..1 000s es dadennseceesnvocenceases one i. DR aemINMIneD
2. Antero-lateral border at ioe aie ieee of the
postero-lateral .......s0..0+6 sontecccvececoscenes Me MOCHIatiS,
II. Thumb of the larger cheliped Hee a ie Bee |
TOOEH Ab DASE ... + +..ccceccsecsseccsssonrsesttegn ben paceessnenss corioen.-a:) siientnmnnne nn
134. Hurycarcinus orientalis, A. Milne Edwards.
Eurycarcinus orientalis, A. Milne Edwards, Ann. Soc. Entom. France (4) VII,
1867, p. 277: de Man, Notes Leyden Mus. XIV. 1892, p. 226.
Carapace rather over two-thirds as long as broad, perfectly smooth
(except for an extremely fine and faint granular ridge that runs
transversely inwards towards the gastric region from the last tooth of
the antero-lateral margin), decidedly convex fore and aft and slightly
so from side to side.
Front cut quite straight and square, slightly emarginate in the
middle line.
1898. ] A. Aleock— Carcinological Fauna of India. 211
Antero-lateral border cut into four thin shallow teeth, of which
the first two are rounded and the last two are anteriorly acuminate,
the first being the least prominent of all and the last being the smallest
of all. The antero-lateral border is extremely short, a good deal less
than two-thirds the length of the postero-lateral.
Supra-orbital margin entire, the infra-orbital finely denticulate.
Chelipeds markedly unequal, perfectly smooth, iuner angle of
wrist rather strongly pronounced ; the hand and fingers are rather short
and stout and the thumb of the larger cheliped is a good deal shorter
than the hand and has a very strong tooth at its base,
The legs and under surface of the body are covered with a dense,
extremely short scurfy tomentum.
Colours in spirit yellowish brown.
In the Indian Museum are four specimens, from Karachi, Bombay
and the Andamans.
This species agrees in all respects with the descriptions and figures
of Hurycarcinus maculatus, except in respect of the antero-lateral
borders. These are so short that a line joining their posterior ex-
tremities would divide the carapace into two halves, of which the
anterior would be much the smaller: the teeth of the antero-lateral
border are also much shallower and less salient.
135. Hurycarcinus grandidiert, A. Milne Edwards.
Eurycarinus grandidiert, A. Milne Edwards, Ann. Soc, Entom. France, (4) VIT.
1867, p. 277; and Nouv. Archiv. du Mus. IV. 1868, p. 80, pl. xix. figs. 13-16.
Carapace about two-thirds as long as broad, strongly convex fore
and aft, slightly so from side to side, perfectly smooth to the naked eye.
Front cut square, emarginate in the middle line, the fore edge
straight but sloping a little obliquely from the outer angles to the
middle line. Supra-orbital margin entire, the infra-orbital obscurely
denticulate.
Antero-lateral border as in H. orientalis but rather longer, its
length being at least two-thirds that of the postero-lateral; the edges
of all the teeth are a little thickened and granular.
Chelipeds unequal, perfectly smooth, inner angle of wrist pro-
nounced: the hand is more elongate and narrower and the fingers are
slenderer than in FH. orientalis, and the thumb of the larger cheliped has
xio enlarged tooth at the base. The legs, the smaller cheliped, and the
under surface of the body are covered with a dense, extremely short
and fine tomentum.
Colours in spirit, yellowish brown.
In the Indian Museum is a single specimen from the Nicubars.
212 A. Alcock —Carcinological Fauna of India. [No. 1,
The chief difference between this species and EH. orientalis and
maculatus appears to be in form of the hand and fingers of the larger
cheliped.
136. Eurycarcinus maculatus, (A. M. Edw.) de Man.
Pilumnopeus maculatus, A. Milne Edwards, Ann. Soc. Entom. France, (4) VII.
1867, p. 277; and Nouv. Archiv. du Mus. IV. 1868, p. 82, pl. xix. figs. 17-19.
Eurycarcinus maculatus, de Man, Journ. Linn. Soc, XXII. 1887-88. p. 44, pl. ii.
figs. 2 and 3 (not 3 and 4).
The Mergui specimen described by de Man does not appear to be in
the Indian Museum.
This species agrees with LH. orientalis in the form of the chelipeds
(hand and thumb), and appears to differ from that species only in having
a longer and more deeply cut-up antero-lateral border.
NECTOPANOPE, Wood-Mason.
Nectopanope Wood-Mason, Ann. Mag. Nat. Hist. March, 1891, p. 261.
Carapace broad, approaching the quadrilateral, convex fore and
aft, the branchial regions so inflated and convex dorsally as to make the
transverse plane of the carapace strongly concave in the middle line,
the other regions obscurely defined, the surface smooth.
The antero-lateral borders are very much shorter than the postero-
lateral, are very thin and sharp, and are cut into teeth of which the
first is confluent with the outer orbital angle.
Front broad, a third the greatest breadth of the carapace, straight,
square cut, slightly projecting beyond the supra-orbital angle, from
which it is sharply cut off by an angular notch, on either side.
Orbits large, with a thin, sharp, prominent margin; a notch
internal to the middle of the upper margin, the notch breaking this
margin into two curves, one corresponding to the eye-stalk the other to
the cornea: eyes large, reniform, on moderately stout stalks.
Antennules folding transversely. The basal antennal joint is very
short, but almost touches the turned down side-edge of the front: the
flagellum, which is considerably longer than the major diameter of the
large orbit, springs from the rather broad orbital hiatus.
The buccal cavern is broader anteriorly than posteriorly, and the
mouth parts do not nearly reach its front edge, so that a wide and
permanent gap is left: the crests of the endostome are not very strong,
but the free edge of the endostome corresponding to the efferent
- branchial channel, on either side, is deeply excavated. The outer wall —
of the efferent branchial canal forms a strong angular bulge in the
pterygostomian region,
1898. ] A. Alcock — Carcinological Fauna of India. 213
The chelipeds in the female are equal; the fingers are compressed
and pointed, not hollowed.
The legs are long and slender, the propodite and dactylus of the
last pair strongly compressed and a little broadened.
This form is most nearly related to Hurycarcinus.
137. Nectopanope rhodobuphes, Wood-Mason.
Nectopanope rhodobaphes, Wood-Mason, Ann. Mag. Nat. Hist. March, 1891,
p. 261.
Carapace about ¢ as long as broad. Front extremely obscurely
grooved in the middle line. Antero-lateral border cut into three thin
sharp-edged ‘teeth, of which the first is broad and rounded and con-
fluent with the orbit, the second is broad and anteriorly acuminate, and
the third almost spiniform.
Chelipeds smooth, in the female they are equal and are a little over
1; times the length of the carapace: arm with an acute spine near the
far end of the upper border; inner angle of wrist acute, spiniform ;
fingers thin, compressed, each and hooked at tip, armed with thin
laciniate teeth, the thumb very broad.
Legs thin, the first three pairs not much shorter than the chelipeds,
with long compressed-styliform dactylus: the last pair a good deal
shorter, with thin blade-like propodite and dactylus closely fringed
with hair.
Colours in spirit uniform yellowish white: in life pink, witha
dotted, V-shaped, white mark between the gastric and branchial regions.
In the Indian Museum is a single female specimen from off the
Godavari coast 98-102 fms.
Nectopanope longipes, which was referred provisionally to this genus
by Wood-Mason, who had insufficient material for examination, turns
out, now that numerous good specimens have been dredged by the
“‘ Investigator,” to be a Catometope.
Subfamily VII. Heipnina.
Alliance I, Hriphiowda.
Eripaia, Latr.
Eriphia, Latreille, Cuvier Régne An. (1) III. 18,
Eriphia, Desmarest, Consid. Gen. Crust. p. 125.
Eriphia, De Haan, Faun. Japon. Crust. p. 22.
Eriphia, Milne Edwards, Hist. Nat. Crust. I. 425.
Eriphia, Dana, Silliman’s Journ. (2) XII, 1851, p. 128; and U. S, Expl. Exp.
Crust. pt. I. p. 246.
Eriphia, Heller, Crust. Sudl. Kurop, p. 74.
214 A. Aleock — Carcinological Fauna of India. [No. 1
~
Eriphia, A. Milne Hdwards, and Miss. Sci. Mex. Crust. p. 337.
Eriphia, Miers, Challenger Brachyura p. 162.
Carapace thick and deep, approaching a quadrilateral shape, very
little convex or nearly flat, not remarkably broader than long, the
regions except the gastric not demarcated.
Antero-lateral borders slightly curved, much shorter than the
postero-lateral and meeting the latter, not at a strong angle as in most
Cancrids, but at a very open and imperceptible angle; though spinate
they are not cut into lobes.
The fronto-orbital border is extremely broad, much more than
three-quarters the greatest breadth of the carapace ; the front, which is
therefore broad also, is strongly deflexed, is almost straight, and is cut
into two broad lobes the outer part of each of which is broadly in
contact —far beyond the limits of the antennal base —with a singular-
ly broad prolongation of the infra-orbital plate. The orbits, which are
deep and oval, are therefore completely closed and widely separated
from the antenne.
Tlfe basal antennal joint is very small short and broad ; the flagel-
lum is long, more than the major diameter of the large orbit. The
antennules fold transversely.
The crests of the endostome, defining the expiratory canals, are
strong, and the canal is completed below by the foliaceous process of the
first maxillipeds, the anterior edge of that process being coneave. The
oblique anterior border of the merus of the external maxillipeds is not
notched.
Chelipeds massive, unequal in both sexes; fingers stout, pointed,
not hollowed.
The abdomen of the male has all 7 segments separate.
Key to the Indian species of Hriphia.
I, Carapace nearly 4 as long as broad, devoid of hair dorsally ;
front cut into blunt teeth :—
1. Chelipeds smooth to the naked eye ...............002... EH. levimana.
2. Hand and wrist of the smaller = nee sindiled with
tubercles .....0-.0.5. +6 signees sd senecesreegenes + die GaMmmN te
II. Carapace only $ as fee as ena: with numerous scattered
hairs; front not cut into teeth ......... Sean ssuiies pias steeiat sea E. scabricula.
138. LHriphia levimana, Latr. Edw.
Eriphia levimana, Guérin, Icon. R. A., Crust. pl. iii. fig. 1: Milne Edwards,
Hist. Nat. Crust. I. 427: Dana, U. S. Expl. Exp. Crust. pt. I. p. 249, pl. xiv.
figs. 7a-c: Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 37: A. Milne Edwards, in
Maillard’s Vile Réunion, Annexe F. p. 5, and Nouv. Archiv. du Mus. IV. 1868, p. 71,
and 1X. 1873, p. 255: Heller, Novara Crust. p. 24: Hilgendorf in v. d. Decken’s
1898. ] A. Aleock — Carcinological Fauna of India. 215
Reisen Ost-Afr. III. i. p. 75, and MB. Ak. Berl. 1878, p. 797: Miers, P. Z. S. 1877,
p. 135, and Ann, Mag, Nat. Hist. (5) V. 1880, p. 237, and Zool. H. M.S. Alert,
pp. 517, 5384, and Challenger Brachyura, p. 162: Tozzetti, Magenta Crost. p. 60,
pl. v. figs. la-c: HE. Nauck, Zeits. Wiss. Zool. XXXIV. 1880, p. 58 (gastric teeth) :
Richters in Mébius Meeresf. Maurit. p. 151: Haswell, Cat. Austral. Crust. p. 75:
Muller, Verh. Ges. Basel VIII. 1886, p. 475: de Man, Journ. Linn. Soc., Zool., XXII.
1887-88, p. 68, and Archiv. f. Naturges. LITI. 1887, i. 327, and Zool. Jahrb. Syst. &c.
VIII. 1894-95, p. 555: Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 367:
Ortman, Zool. Jahrb., Syst., VII. 1893-94, p. 480, and in Semon’s Forschungsr.
(Jena. Denk. VIII) Crust. p. 54: Zehntner, Rev. Suisse Zool. II. 1894, p. 161:
Whitelegge, Mem. Austral. Mus. III. 1897, p. 137.
Eriphia trapeziformis, Hess, Archiv. fiir Naturges. XX XI. 1865, i. pp. 135, 171,
pl. vi. fig. 4 (see de Man, Zool. Jahrb., Syst., II. 1887, p. 695).
Carapace nearly # as long as broad: gastric region large, well
demarcated and subdivided into three large subregions, its anterior
part, like the anterior part of the branchio-hepatic regions, covered
with small pearly and subsquamiform tubercles ; the rest of the carapace
smooth, but closely covered with very small vesiculous granules not
plainly visible to the naked eye: the post-orbital groove is distinct,
and behind it, parallel with the gastric region, on each side a small
narrow areola is marked off.
The free edge of the frontal lobes is bluntly spinate: there is a
blunt spine also at the lower inner angle of the orbit, and two or three
at the outer angle of the orbit: and there are 5 or 6 blunt spines or
spinules of decreasing size along the antero-lateral border.
Chelipeds almost smooth to the naked eye, though closely covered
with small depressed vesiculous granules under the lens: upper border
of arm denticulate at its distal end, where also the granules on the
neighbouring part of the outer surface are plainly visible without a
lens; the anterior border of the arm denticulate at its proximal end.
The upper part of the inner surface of the wrist forms a distinct facet,
the proximal angle of which is pronounced and the distal end of which
is bounded by two or three blunt spines.
Legs stout, smooth; upper edge of merus denticulate and some-
what hairy, the lower edge with tufts of stiff hair: similar tufts of hair
along upper edge of carpus ‘and on all the edges and surfaces of the
propodite ; the greater part of the dactylus covered with short stiff
hairs and longer bristles.
Colours in spirit dull maroon, with a bluish-green tinge on the
postero-lateral parts of the carapace and on the walking-legs.
In the Indian Museum are 53 specimens, from the Andamans,
Arakan coast, Mergui, Ceylon and Laccadives (besides 2 from
Samoa).
216 A. Alcock — Carcinological Fauna of India. [No. 1,
139. Priphia levimana var. Smithii, Macleay, Hilgdf.
Eriphia smithii, Macleay, Ill. Ann. 8S. Afr. p. 60: Krauss, Sadafr. Crust. p. 36,
pl. ii. fig. 3: Dana, U. 8S. Expl. Exp. Crust. pt. I. p. 251: Stimpson, Proc. Ac. Nat.
Sci. Philad. 1858, p. 37: A. Milne Edwards, Nouv. Archiv. du Mus. IV. 1868, p. 71:
Hoffmann in Pollen and Van Dam, Faun. Madagasc., Crust. p. 6, pl. i. figs. la-c:
Lenz and Richters, Abh. Senck. Ges. XII. 1881, p. 422: Ortmann in Semon’s
Forschungsr. (Jena. Denk. VIII) Crust. p. 54.
Eriphia levimana var. smithii, Hilgendorf MB. Ak. Berl. 1878, p. 797: Miers,
Ann. Mag. Nat. Hist. (5) V. 1880, p. 237, and Zool. H. M. S. Alert, pp. 517, 535:
de Man, Archiv. f. Naturges. LIII. 1887, i. p. 327: P ? Cano, Boll. Soc. Nat. Napoli,
III. 1889, p. 210: A. O. Walker, Journ. Linn. Soc., Zool., XX. 1886-90, p. 110:
Ortmann, Zool. Jahrb., Syst., VII. 1893-94, p. 481.
The variety Smithw differs from the typical Eriphia levimana only
in the sculpture of the chelipeds.
The upper and outer surfaces of the wrists and hands of the smaller
cheliped are closely covered with miliary granules and are profusely
studded with salient and subsquamous tubercles, which, on the lower
half of the hand are arranged in longitudinal series.
The wrist and hand of the larger cheliped may be nearly smooth
or may have a few scattered pustulous tubercles (as they are in most
Indian specimens), or they may more nearly resemble the smaller
cheliped in sculpture.
In the Indian Museum are 15 specimens from Karachi, and one
from the Mekran coast.
140. Hriphia scabricula, Dana.
Eriphia scabricula, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 82, and U. 8. Expl.
Exp. Crust, pt. I. p. 247, pl. xiv. figs. 5a-b: Stimpson, Proc. Ac. Nat. Sci. Philad.
1858, p.'37: A. Milne Edwards, Nouv. Archiv. du Mus. 1X. 1873, p. 256: Hilgendorf,
MB. Ak. Berl. 1878, p. 798: Richters in Mobius Meeresf. Maurit. p. 151: Lenz and
Richters, Abh. Senck. Ges. XII. 1881, p.422: Miers, Zool. H. M. S. “ Alert,” pp. 518,
535: de Man, Notes Leyden Mus. XII. 1890, p. 66, and Zool. Jahrb., Syst., VIII.
1895, p. 555: Ortmann, Zool. Jahrb., Syst., VII. 1893-94, p. 480: Whitelegge, Mem.
Austral. Mus. III. 1897, p. 137.
Eriphia gonagra, Krauss (nec Edw.) Sudafr. Crust. p. 36.
8 °
Carapace ~ as long as broad, grooved on the surface as in
E. laxvimana, closely covered anteriorly and laterally with sharpish
subsquamiform granules among which are numerous soft but stiffish
hairs.
Free edge of frontal lobes entire, microscopically beaded. A sharp
tooth at the outer angle of the orbit only. Antero-lateral border with
4 or 5 sharp teeth of gradually decreasing size.
Upper and outer surfaces of wrists and hands closely covered with
vesiculous granules and sharpish pearly tubercles with numerous hairs
1898. ] A. Alcock — Carcinological Fauna of India. 217
between them, the tubercles on the smaller hand being in longitudinal
series and the hairs thick there.
Legs smooth; the borders of the last four joints, specially the
upper border, fringed with longish hairs.
Colours in spirit, warm light brown, the legs in good specimens
cross-banded alternate dark and light brown.
In the Indian Museum are three specimens, from the Laccadives,
the Andamans and Ceylon, (also one from Samoa)
Alliance II. Trapezioida.
Trapezia. Tetralia. Quadrella. Sphenomerus.
TRAPEZIA, Latreille.
Trapezia, Latreille, Fam. Nat. p. 269, and Encyclop. Meth. x. 695.
Trapezia, Milne Edwards, Hist. Nat. Crust. I, 427.
Trapezia, Dana, Silliman’s Journ. (2) XII. 1851, p. 128, and U. S. Expl. Exp.
Crust. I. p. 252.
Trapezia, A, Milne Edwards, Nouv. Archiv. du Mus. IX. 1878, p. 257 and Miss.
Sci. Mex., Crust. p. 341.
Trapezia, Miers, Challenger Brachyura, p. 163.
Trapezia, Ortmann, Zool. Jahrb., Syst., X. 1897, p. 202.
Grapsillus, Macleay in Smith’s Ill. Zool. S. Afr. p. 67.
Carapace approaching the quadrilateral, little convex, not much
broader than long, smooth and without any trace of regions.
Antero-lateral borders much shorter than the postero-lateral, run-
ning backwards almost straight and parallel with one another, not
therefore meeting the convex curved and convergent postero-lateral
borders at any angle.
Fronto-orbital border extremely broad, about as extensive as the
greatest breadth of the carapace. Front broad, horizontal, lamellar
separated from the supra-orbital angle by a notch; cut into two lobes,
of which both the inner and outer angles are pronounced: so that with
the supra-orbital angle the front usua!ly appears 6-toothed.
The orbits, which afford no concealment to the eyes and are large,
are cut out of the antero-lateral angles of the carapace: their dentiform
upper and lower inner angles are broadly in contact, so that the
antenns are widely excluded from the orbit: their margins are without
fissures or sutures.
The antennules fold nearly transversely, but in most spirit specimens
are extended beyond their fosse. The basal antennal joint is slender
and very short and does not nearly reach the front: the flagellum is
very long, much longer than the major diameter of the orbit.
J. 11. 25
218 A. Aleock— Careinological Fauna of India. [No. I,
The crests of the endostome, defining the expiratory canals, are
well developed and the canals are closed in below by the foliaceous
process of the Ist maxillipeds: the anterior edge of the merus of the
rather slender external maxillipeds is not notched.
The chelipeds are long and very massive and are sub-equal or not
very unequal in both sexes: the arm usually projects a long way beyond
the carapace, and has its anterior edge sharp and crest-like and serrate :
the fingers have usually a thin and sharp cutting-edge, best marked on
the immobile finger. Legs stout, of moderate length.
The abdomen of the male consists of 5 segments, the 3rd—5th being
fused.
The species of Trapezia are found in the crevices of coral-stocks.
Key to the Indian species of Trapezia.
I, A distinet spine or tooth at the junction of the antero-
lateral and postero-lateral borders of the carapace :—
i. Lower border of hand sharp, entire :—
1. Outer surface of hand, in its upper part
at least, covered with a mass of fine
tangled downy hairs ...es«..... .... J. cymodoce.
2. Outer surface of hand smooth me fine —
a. Carapace and appendages plain
yellowish or reddish brown ......... TT. ferruginea.
b. Carapace (and sometimes also the
upper surface of the hands) covered
with an elegant meshwork of fine
dark brown lines (a scurfy pubes-
cence on outer surface of hand,
occaSionally): ..cecauss ssc asec ctuseeree ee QUOORMEIE
e. Carupace and sannatne ee: every
where covered with roundish red
BPOGE ciesice (pve asigah de dees eeis bois slanions due ine | La ARRON MmmE
d. Duranens espe with faintish
brown spots, upper surface of hands
with a network of brown Hines ... T. intermedia.
ii. Lower border of hand granular or bluntly ser-
rulate ;: carapace, etc. covered with roundish red
BPOtS ... ccccccece Sites es .... 1. rufopunctata.
41, Nothing more tia an 1 dhidistinct: anton at the sstetiion
of the antero-lateral and postero-lateral borders:
colours, in spirit, blackish brown..,........5.00 00+ .essereees-. Le digitalis.
With the species of Trapezia the citations of the various writers:
are so extremely uncertain that I have given up the attempt to make
them complete.
Ortmann, in Zoologische Jahrbicher, Abth. fiir Systematik, ete. X. 1s
1898.} A. Alcock — Carcinological Fauna of India. 219
1897, pp. 201-216, has published a little monograph of the group, in
which full lists of citations will be found.
141. Trapezia cymodoce, (Herbst) Miers, de Man, Ortmann.
Cancer cymodoce, Herbst, Krabben ITI. ii. 22, pl. li. fig, 5.
Trapezia cymodoce, Savigny and Audouin, Descr. de Egypte, Crust. p. 85, pl. v.
fig. 2: Miers, Ann. Mag. Nat. Hist. (5) II. 1878, pp. 408, 409: de Man, Notes Leyden
Mus. II. 1880, pp. 177, 178, and Journ. Linn. Soc., Zool., XXII. 1887-88, p. 69:
Ortmann, Zool. Jahrb., Syst. X. 1897, pp. 2038, 204.
Trapezia hirtipes, Lucas in Jacquinot’s Voy. Astrolabe, Zool. III. Crust. p. 44
pl. iv. fig. 14.
Trapezia cerulea, Heller, SB. Ak. Wien, XLIII. 1861, p. 348.
Trapezia dentata, A. Milne Edwards, Nouv. Archiv. du Mus. 1X. 1873, p. 261.
Carapace four-fifths as long as broad, slightly convex in both
directions in the adult female, almost flat in the male, smooth and
polished.
Front prominent beyond the supra-orbital angle, rather deeply cut
into two lobes, each of which has the inner angle dentiform and still
further promineut, and the outer angle (though rounded) sharply
marked and separated by a deep notch from the dentiform supra-orbital
angle.
Inner angle of lower edge of orbit acutely spiniform : outer angle of
orbit acute.
Antero-lateral borders nearly parallel with one another or very
slightly curved outwards, an acute procurved spine marks their junction
with the postero-lateral borders.
Chelipeds sub-equal in both sexes: more than 2¢ times the length
of the carapace in the adult male but not quite so long in the female:
the arm, which projects far beyond the edge of the carapace, has the
anterior border foliaceous and cut into numerous sharp teeth: inner
angle of wrist sharp and prominent, but not usually spiniform: hands
long and compressed, the upper and lower edges (especially the lower)
both sharp, the upper part of the outer surface of the hand (and wrist
also, in many cases) covered with silky wool; fingers compressed, the
cutting-edges thin sharp and not much toothed.
Legs smooth, the dactylus with rather numerous silky bristles,
which are also found scattered along both edges of propus and upper
edge of carpus.
Colours in spirit yellowish or reddish brown, often very dark or
livid on the carapace ; distal two-thirds of fingers commonly dark brown.
In the Indian Museum are 37 specimens, from the Andamans,
Nicobars, Mergui, Palk Straits and the Mekran coast (besides 16 from
other parts of the Indo-Pacific).
220 A. Alcock — Carcinological Fauna of India, [ No. 1,
In some specimens the free edge of the frontal lobes is more or less
crenulate: the outer angle of the orbit and the lateral epibranchial
spine are sometimes blunt: the whole of the outer surface of the hand
is sometimes pubescent, and the arm also.
The species can, however, always be recognized by the uniform
colouration, the hairy outer surface of the hands, and the very promi-
nent front.
142, Trapezia ferruginea, Latr., Miers, de Man, Ortmann.
Trapezia ferruginea, Latreille, Encycl. Meth. X. p. 695: Milne Edwards, Hist.
‘Nat. Crust. I. 429: Heller, SB. Ak. Wien, XLIII. 1861 p. 349, pl. iv. fig. 40: Miers,
Ann, Mag. Nat. Hist. (5) II. 1878, pp. 407, 408: de Man, Notes Leyden Mus. IT.
1880, pp. 178, 179: Ortmann, Zool. Jahrb. Syst., X. 1897, pp. 202, 205.
Grapsillus subinteger, Macleay in Smith’s Ill. Zool. S. Afr., Annulosa, p. 67. ;
Trapezia cymodoce, Dana, U. S. Expl. Exp. Crust. pt. I. 257, pl. xv. fig. 5,
A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873, p. 260, and Miss. Sci. Mex.
Crust. p. 342.
Trapezia miniata, Lucas in Jacquinot’s Voy. Astrolabe, Zool. III. Crust. p. 438,
pl. iv. fig. 10.
Trapezia subdentata, Gerstaecker, Archiv. fiir Naturges. XXII, 1856, i. p. 127.
Differs from T. cymodoce, which it closely resembles in form and
colour, in the following particulars :-—
(1) the front as a whole is not so prominent, and its constituent
teeth, as well as the supra-orbital angle, are not so prominent, and
deep-cut: the tooth at the lower inner angle of the orbit is not so
sharp :
(2) the outer angle of the orbit and the lateral epibranchial spine
are not nearly so spiniform in the adult:
(3) the upper border of the hand is not so sharp, and the outer
surface of the hand is smooth, polished and quite hairless.
In the Indian Museum are 25 specimens, from the Andamans,
Nicobars and Ceylon.
Trapezia ferruginea var. intermedia, Miers.
Trapexia rufopunctata var. intermedia, Miers, Challenger Brachyura, p. 168,
pl. xii. fig. 2, 1886.
Trapezia, sp. Richters in Mébius, Meeresf. Maurit. p. 152, pl. xvi. fig. 13, 1880.
Differs from T. ferruginea only in colouration.
The carapace, legs, arms and wrists are covered with light brown
rather blotchy spots, while the upper surface of the hands is marked by
a network of fine brown lines.
a
Trapezia ferruginea var, guttata, Riipp.
Carapace light brown, edge of front brick-red: chelipeds with a network
of fine pinkish-brown lines: legs with small pink spots.
Twelve specimens have just been dredged by Dr. A. R. S. Anderson of the
“Investigator,” off Great Coco I. (Andamans).
1898. | A. Aleock — Carcinological Fauna of India. 221
A faded specimen could not be distinguished from T. ferruginea.
In the Indian Museum are 3 specimens, from Diamond Island (off
C. Negrais, Burma).
The Museum also possesses one of the ‘“Chailenger” duplicates
from Honolulu.
143. Trapezia ferruginea var. areolata, Dana.
Trapezia areolata, Dana, Proc. Acad. Nat. Sci. Philad. 1852, p. 83, and U.S.
Expl. Exp, Crust. pt. I. p. 259, pl. xv. figs. 8a-b and 9: Heller, Novara Crust. p. 25:
de Man, Archiv. fiir Naturges. LIII. 1887, i. p. 317, and Zool. Jahrb. Syst. VIII.
1894-95, p. 556: Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 366: Ortmann,
Zool. Jahrb. Syst. VII. 1893-94, p. 485.
Trapezia reticulata, Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 37.
Trapezia areolata var. inermis, A. Milne Edwards, Nouv. Archiv. du Mus. IX.
1873, p. 259, pl. x. fig. 6: Miers, Challenger Brachyura, p. 167: Zehntner, Rev.
Suisse Zool. V. 1894, p. 157.
Trapezia ferruginea areolata, Ortmann, Zool. Jahrb., Syst., X. 1897, pp. 208, 206.
This species also differs from 7’, ferruginea only in colouration.
The carapace, and sometimes also the upper surface of the chelipeds,
is covered by a very elegant honeycomb network of fine brown (in
spirit) lines. Even in old spirit specimens this network can be made
out, with a lens, on the carapace, though not on the chelipeds.
In many specimens of T. areolata, the upper part of the outer
surface of the hand is covered with a very fine scurf-like pubescence.
In the Indian Museum are 52 specimens, from the Audamans,
Nicobars, Ceylon and Mergui (besides 3 from other parts of the Indo-
Pacific).
144. Trapezia maculata (Macleay) Dana.
Grapsillus maculatus, Macleay in Smith’s Ill. Zool. 8S. Afr., Ann. p. 67.
Trapezia tigrina, Kydoux and Sonleyet, Voy. Bonite, Vol. I. p. 232, pl. ii. fig. 4.
Trapezia maculata, Dana, U.S. Expl. Exp. Crust. pt. I. p. 256, pl. xv. figs. 4a-d:
Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 37, and Ann. Lyc. Nat. Hist., N. York,
VII. 1862, p. 219: Streets, Bull. U. S. Nat. Mus. VII. 1877, p. 106: de Man, Archiv.
fiir Naturges. LIII. 1887, i. p. 318, pl. xiii. fig. 2: J. R. Henderson, Trans. Linn.
Soc., Zool., (2) V. 1893, p. 366.
Trapezia rufopunctata var. maculata, Miers, Phil. Trans. Roy. Soc., Vol. 168,
1879, p. 487 : Ortmann, Zool. Jahrb., Syst. VII. 1893-94 p. 484.
Trapezia ferruginea maculata, Ortmann, Zool. Jahrb., Syst., X. 1897, pp. 203, 206,
Differs from 17’. cymodoce in the following particulars :—
(1) the front as a whole is not so prominent, nor are its constituent
teeth and the supra-orbital angle quite so deep-cut:
(2) the chelipeds are not much more than twice the length of the
carapace in the male, the arm being shorter than in 7’. cymodoce and
ferruginea ; there is a strong spine at the inner angle of the wrist; the
outer surface of the hand is smooth, polished and hairless :
222 A. Aleock— Carcinological Fauna of India. [ No. 1,
(3) the carapace, chelipeds, legs, etc., are everywhere covered with
well defined roundish red spots.
N.B. ‘the lower border of the hand is sharp and entire (non-
granular, non-serrulate).
In the Indian Museum are four specimens from Table Island (north
of the Andamans).
145, Trapezia rufopunctata (Herbst) Latr., Ortmann.
Cancer rufopunctatus, Herbst, Krabben III. i. 54, pl. xlvii. fig. 6.
Trapezia rufopunctata, Latreille, Encyclop. X. p. 695: Dana, U. S. Expl. Exp.,
Crust. pt. I. p. 255, pl. xv. figs. 3a-b: Lucas in Jacquinot’s Voy. Astrolabe, Zool.
III. Crust. p. 41, pl. iv. fig. 8: Gerstaecker, Archiv. fir Naturges. XXII. 1856, i.
p. 128: Heller, SB. Ak. Wien, XLIII. 1861, p. 350: A. Milne Edwards, Nouv.
Archiy. du. Mus. IV. 1868, p. 71, and IX. 1873, p. 258, and Miss. Sci. Mex. Crust.
p. 342: Hilgendorf in v.d. Decken’s Reisen Ost-Afr. III. i. p. 75, pl. ii, fig. 3:
Kossman, Reise roth. Meer., Crust. p. 42: Miers, Challenger Brachyura, p. 167:
de Man, Archiv. fiir Naturges. LIII. 1887, i. p. 318, pl. xiii. figs. 1,2: J. R.
Henderson, Trans. Linn. Soc., Zool., (2) V. 18938, p. 366: Ortmann, Zool. Jahrb.,
Syst. VII. 1893-94, p. 484, and X. 1897, pp. 205, 207, and in Semon’s Forschungsr.
(Jena. Denk. VIII.) Crust. p. 52: Zehntner, Rev. Suisse Zool. IL. 1894, p. 157.
Trapezia acutifrons, A. Milne Edwards, Ann. Soc. Eutom, France, (4) VII. 1867,
p. 281.
Differs from 7’. cymodoce as follows :—
(1) though the front is of the same general form, the edge of each
frontal lobe is somewhat angularly excised and the outer angle is
angularly acute (not rounded) and is produced to or even beyond the
level of the dentiform inner angle of each lobe:
(2) the inner angle of the wrist is more acute and spiniform, . the
upper border of the hand is rounded and the outer surface smooth
polished and hairless, and the lower border of the hand is granular or
bluntly serrulate :
(3) the carapace, chelipeds and legs are covered with rather large
red spots.
In the Indian Museum are 5 specimens from Ceylon.
146. Trapezia digitalis, Latr.
Trapezia digitalis, Latreille, Encycl. Meth. X.696: Milne Edwards, Hist. Nat.
Crust. I. 429: Heller, SB. Ak. Wien, XLIII. 1861, p. 352: Kossmann, Reise roth.
Meer., Crust. p. 42: de Man, Notes Leyden Mus. II. 1880, p. 177: Ortmann, Zool.
Jahrb. Syst. X. 1897, pp. 203, 208.
Trapezia leucodactyla, Riippell, 24 Krabben roth. Meer. p. 28,
? Trapezia fusca, Lucas in Jacquinot’s Voy. Astrolabe, Zool. III. Crust. p. 45,
pl. iv. fig. 17.
Carapace about five-sixths as long as broad, but having a broader
look, owing to the less marked projection of the front and the greater
1898. | A. Alcock — Carcinological Fauna of India. 225
eurvature and convergence of the postero-lateral borders; its surface
smooth and burnished.
The front is slightly notched in the middle line, and is separated
from the hardly-dentiform supra-orbital angles by a shallow notch: it
is thus rather obscurely divided into two lobes, each of which has the
free edge finely denticulate. Outer angle of orbit acute, as is also the
inner angle of the lower margin.
There may be a slight notch at the junction of the antero-lateral
and postero-lateral borders, but there is never a spine.
Chelipeds subequal mm both sexes, about twice the length of the
carapace, smooth and burnished. The arm is much shorter than it is in
T. cymodoce and ferruginea, being broader than long, its foliaceous
anterior border dentate or crenate; inner angle of wrist acute ; upper
border of hand rounded, lower border sharp.
Legs smooth, dactylus with a few bristles, which are almost absent
from the other joints,
Colours in spirit, blackish-brown, fingers, lower edge of hand and
distal ends of leg joints lighter.
In the Indian Museum are six specimens from Ceylon and Palk
Straits.
TeTrRaLiaA, Dana.
Tetralia, Dana, Silliman’s Journ. Sci. and Arts (2) XII. 1851, p, 128, and Proc.
Ac. Nat. Sci. Phila., 1852, p. 83, and U. S. Expl. Exp. Crust. pt. I. p. 261.
Tetralia, Heller, SB. Ak. Wien, XLIII. 1861, p. 358.
Tetralia, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873, p. 261.
Closely resembles Trapezia in form, and only differs in the following
characters :—
The front is hardly separated from the hardly-dentiform supra-
_ orbital angle by a small and very mconspicuous notch, and has its free
edge very slightly convex, very faintly sinuous or straight (instead of
being divided into lobes or teeth), and finely denticulate.
The antero-lateral borders are usually continued into the postero-
lateral without any trace of a spine or notch to mark their junction.
The eyes are smaller.
The chelipeds are usually remarkably unequal ; the arms are shorter
and their expanded anterior edge is not denticulate throughout.
The meropodites of the legs are short and broad, almost foliaceous.
The abdomen of the male consists of seven separate segments.
147. | Tetralia glaberrima (Herbst. ),
Cancer glaberrimus, Herbst, Krabben I. ii. 262 pl. xx. fig. 115.
Trapezia integra, Latreille, Wncycl, Meth. x. p. 696.
224 A. Alcock — Carcinological Fauna of India. [ No. ],
Trapezia glaberrima, Krauss, Sudafr. Crust. p. 35.
Tetralia glaberrima, Dana, U. S. Expl. Exp. Crust. pt.i. p. 263, pl. xvi. fig. 3:
Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 38: A Milne Edwards, Nouv. Archiv.
du Mus. IX. 1873, p. 262: Kossmann, Reise roth. Meer. Crust. p. 46: Lenz and
Richters, Abh. senck. Ges. XII. 1881, p. 422; de Man, Archiv. fiir Naturges. LIII. 1887,
i. p. 821: J. R. Henderson, Trans. Linn. Soc., Zool., (2) V. 1893, p. 366: Ortmann.
Zool. Jahrb., Syst., VII. 1893-94, p. 485, and X. 1897, p. 209, and in Semon’s
Forschungsr. (Jena. Denk, VIII.) Crust. p. 58: Zehntner, Rev. Suisse Zool. II. 1894,
p. 157.
Tetralia nigrifrons, Dana, Proc. Ac. Nat. Sci. Philad. 1852, p. 83, and U. S.
Expl. Exp. Crust. pt. i. p. 262, pl. xvi. figs. 2a-d: A. Milne Edwards, Nonv. Archiv.
du Mus. 1X. 1873, p. 262: Hilgendorf, MB. Ak. Berl. 1878, p. 798.
Trapezia serratifrons, Lucas iu Jacquinot’s Voy. Astrolabe, Zool., LII. Crust. p. 47,
pl. iv. fig. 20.
Tetralia levissima, Stimpson, Proc. Ac. Nat. Sci. Philad. 1858, p. 38.
Tetralia cavimana, Heller, Abh. zool.-bot. Ges. Wien. XI. 1861, p. 14, and SB.
Ak. Wien. XLIII. 1861, p. 353, pl. iii. figs. 24, 25, and Novara Crust. p. 26: Miers,
Phil. Trans. 168, 1879, p. 488, and Zool, H. M.S. Alert, pp. 518, 537: de Man.
Notes Leyden Mus., II. 1880, p. 180: R. I. Pocock, Ann. Mag. Nat. Hist. (6) V. 1899,
p. 73: Whitelegge, Mem. Austral. Mus, ITI. 1897, p. 138.
Tetralia heterodactyla, Heller, Abh. zool.-bot. Ges. Wien. XI. 1861, p. 14, and
SB. Ak. Wien. XLIII. 1861, p. 354.
Carapace about five-sixths as long as broad, flat smooth and shiny,
with occasionally a faint short and distant pubescence near the frontal
and lateral margins. |
The front is finely and evenly denticulate, is almost straight, and
is generally but not always separated from the similarly denticulate
supra-orbital angle by a slight and inconspicuous break. The lateral
borders are very slightly curved in their anterior half and are moderate-
ly convergent in their posterior half: they show no trace of a spine or
notch, at least in the adult. — ,
The chelipeds are very unequal both in length and bulk in both
sexes, but even more so in the male than in the female. \
In the male the larger cheliped is a good deal more and the smaller
a good deal less than twice the length of the carapace: in the female
the larger is about 1f times and the smaller about 14 times the length
of the carapace.
The arm has the distal end of its anterior border expanded and
finely denticulate: a little down and a few hairs are present on the
outer surface of the wrist hand and finger, especially in the larger
cheliped. |
At the base of the larger hand, on the upper part of the outer
surface, is a roundish pit of variable size and depth and usually full of
hair.
The legs are rather short and stout and end in a curious little
1898. ] A. Alcock — Carcinological Fauna of India. 225
coarse blunt claw: the meropodites are singularly broad and flat: the
-dactyli and propodites have both edges, and the carpopodites the upper
edge, somewhat hairy.
Colours in spirit rather variable: sometimes uniform yellow or
brown, usually the edge of the front and of the anterior part of the
lateral margin is darker —almost black; occasionally the ends of some
of the leg-joints have a black spot, and sometimes the legs are broadly
banded yellow and blackish-brown.
In the Indian Museum are 78 specimens, from the Andamans,
Mergui, Ceylon, the Maldives and the Mekran coast.
In some but not in all young specimens there is a small lateral
spine placed far forward on either lateral border of the carapace.
QUADRELLA, Dana.
Quadrella, Dana, Silliman’s Amer. Journ. Sci. and Arts (2) XII. 1851, p. 128, and
Proc. Ac. Nat. Sci. Philad. 1852, p. 84, and U.S. Expl. Exp. Crust. pt. i. p. 265.
Quadrella, A. Milne Edwards, Miss. Sci. Mex. Crust. p. 344.
Carapace squarely hexagonal, nearly as long as broad, moderately
convex, perfectly smooth without trace of regions.
The antero-lateral borders, which are about equal in length to the
postero-lateral, are straight, slope very slightly outwards, and join the
postero-lateral at a very wide, but distinct, angle, marked usually by a
spine. |
The fronto-orbital border is about equal in extent to the greatest
breadth of the carapace, and the broad almost horizontal front is cut
into four acute spines, external to which, on either side, is seen the acute
spiniform internal angle of the lower edge of the orbit projecting
beyond the acute supra-orbital angles ; so that the front is commonly
spoken of as six-spinate.
The orbits, which are small and are cut out of the antero-lateral
angles of the carapace, afford no concealment to the eyes: their upper
and lower inner angles are in contact so as to exclude the antenne.
The antennules fold almost transversely. The basal antennal joint
is slender and does not nearly reach the front; the flagellum is slender
and long—nearly half the length of the carapace.
The crests of the endostome are distinct and the expiratory canals
are closed in as in Trapezia, ete.
The chelipeds are massive but are of great length, the whole of
the long arm projecting beyond the edge of the carapace: they are
subequal, or not markedly unequal, in both sexes.
Legs long and slender, the dactyli strongly and evenly serrated
along the inner edge.
J. 1. 29
226 A. Aleock — Carcinological Fauna of India. [No. 1,
The abdomen of the male consists of 5 segments, the 3rd—5th
being fused together.
The species of this genus inhabit stocks of corals and Alcyonarians.
148. Quadrella coronata, Dana.
Quadrella coronata, Dana, Proc. Acad. Nat. Sci. Philad., 1852, p, 84, and U.S.
Expl. Exp. Crust. pt. i. p. 266, pl. xvi. figs. 5a-d: Ortmann, Zool. Jahrb. Syst. X.
1897, p. 210.
Trapezia sp. Miers, Zool. H. M. S. Alert, p. 536, footnote, (see Challenger
Brachyura, p. 163 footnote).
Carapace squarely hexagonal, moderately convex, perfectly smooth,
polished, about as long as broad. Anu acute procurved spine at the open
angle of junction of the antero-lateral and postero-lateral borders.
Front with 6 horizoutal spines, the four larger of which belong to
the front proper, the other two being at the lower inner angles of the
orbits which are much more prominent than the also spiniform upper
inner angles.
Outer angle of orbit acute: eyes small, the diameter of the cornea
about a tenth the length of the carapace.
Chelipeds smooth and polished, about 2? times the length of the
carapace, the arm and the palm each being nearly as long as the carapace.
The whole arm, as well as the end of the ischium, visible, from above,
beyond the carapace; an acicular spine at the inner angle of the
ischium and from six to ten such spines along inner (anterior) border
of arm; one, or two, little spines sometimes, but not always, present at
inner angle of wrist : lower border of hand quite smooth.
Legs long slender, about 1 times the length of the carapace: a
few silky hairs on dactylus and propodite, and sometimes a very few on
the carpus also: the inner edge of the dactylus strongly toothed.
Colours in spirit, milkwhite.
In the Indian Museum are 9 specimens from various parts of the
Indian coasts and islands and from depths of 28 to 88 fathoms (one
specimen from ? 7 fathoms).
Quadrella coronata var. maculosa, nov.
Differs from the typical form in the following particulars :—
(1) the greatest breadth of the carapace is distinctly more than
the greatest length (including frontal spines) :
(2) the chelipeds, under a lens, are frosted over with tiny granules :
the anterior border of the arm is finely denticulate, with 2 or 3 spines
at the distal end only ; the mner border of the hand and thumbis finely
denticulate :
1898.) A. Alcock — Carcinological Fauna of India. 227
(3) the carapace is closely covered with tiny purple (in spirit) dots,
except for a very distinctly defined W-shaped white area stretching
across its posterior half: the legs, sternum and abdominal terga are
less closely covered with similar purple specks: chelipeds white.
Length of carapace 7 millim., breadth 8 millim. |
In the Indian Museum area male from off Table I. (Andamans)
15-35 fms., and a female from off the Andamans, 20 fms.
Quadrella coronata, var. reticulata, nov.
Differs from the typical form in the following particulars :—
(1) the carapace is distinctly broader than long :
(2) the chelipeds in the male are only about 25 times the greatest
length of the carapace, and under the lens are more or less frosted over
with granules: the anterior border of the arm is serrate, the iuner
border of the hand and thumb is finely denticulate :
(3) the carapace is symmetrically traversed by several fine purple-
brown lines which intersect to form a regular and wide meshwork, and
there is an irregular meshwork of similar coloured lines on the hands.
Length of carapace 7 millim., breadth 8 millim,.
In the Indian Museum are a male from the Andamans, taken on a
Spongodes, and two from off Ceylon 34 fms.
149. Quadrella boopsis, n. sp.
Differs from Q. coronata in the following particulars :—
(1) the chelipeds in the male are only about twice the greatest
breadth of the carapace :
(2) the arm is stout, is only about three-fifths the greatest length
of the carapace, and has its anterior border serrated, not spiniferous :
(3) the eyes are large, their diameter being about one-fifth the
greatest length of the carapace:
Colours in spirit uniform yellowish.
Length of carapace equal with the breadth, which is 5 millim.
In the Indian Museum are a male and a female from the Arakan
coast 20-30 fms.
This species is not the young of Q. coronata, which has the long
slender arm and small eyes of the adult.
SpHENOMERUS, Wood-Mason.
Sphenomerus, Wood-Mason, Ann. Mag. Nat. Hist. March 1891, p. 263.
Carapace transversely oval or subcircular, the front and antero-
lateral margius forming together a semicircle; markedly convex in
both directions, perfectly smooth, without trace of regions.
228 A. Aleock— Carcinological Fauna of India. [ No. 1,
Antero-lateral borders shorter than the postero-lateral—a spinule
at their point of junction.
Front somewhat deflexed, broad and broadly bilobed. Orbits
affording little or no concealment to the eyes, without fissures or
sutures: there is a gap between the front and the inner angle of the
orbit in which the antennary flagellum is lodged. The fronto-orbital
border, in the adult, is not quite # the greatest breadth of the carapace.
The antennules fold nearly transversely: the basal antennal joint
does not reach the front, the flagellum is a good deal longer than the
major diameter of the orbit.
The buccal cavern is a little narrowed anteriorly. The crests of
the endostome are very faint, but to make up for this the anterior edge
of the buccal cavern is puffed out and is very deeply excised on either
side of the middle line; the anterior margin of the foliaceous process
of the lst maxillipeds is also excised to correspond, and so a permanent
expiratory orifice is formed, which is very large and prominent beyond
the almost transverse anterior edge of the merus of the external
maxillipeds.
The chelipeds are stout, very long and not very unequal; the whole
of the arm projects beyond the edge of the carapace: the fingers are
somewhat compressed and are pointed.
The legs are rather slender.
The abdomen of the male consists of five pieces, the 3-5th somites
being rigidly united but without obliteration of sutures.
150. Sphenomerus trapezioides, Wood-Mason.
Sphenomerus trapezioides, Wood-Mason, Ann. Mag. Nat. Hist. March 1891,
p. 263: Ill. Zool. Investigator, Crust. pl. v. fig. 2 (where the carapace ig drawn a
little too broad).
Carapace about # as long as broad, convex in all directions, smooth,
polished.
The front is about 3 the greatest breadth of the carapace, is obli-
quely deflexed, and is divided into two rather shallow broadly-rounded
lobes the free edge of which is entire.
The supra-orbital angle is not defined, but the dentiform or spini-
form angle of the lower edge of the orbit can be seen from above.
The antero-lateral margins form with the front a semicircular
curve, each carries three sharp spinules, namely, one at the outer angle
of the orbit, one at the junction with the postero-lateral border and one
exactly intermediate between the other two.
The chelipeds are a little, but not very remarkably, unequal: the
larger one is about 25 times the length of the carapace. Their surface
is smooth and polished. The arm, the whole of which is visible beyond
1898. | A. Alcock — Carcinological Fauna of India. 229
the carapace, has much the same shape as in Trapezia, but its anterior
border, though serrated, is not expanded ; the lower border of the hand
is sharp and somewhat dilated posteriorly, as in Trapezia : the inner angle
of the wrist is rounded, but sometimes carries a small spinule.
The legs are slender smooth and polished, and have a few hairs
distally.
Colours in spirit yellowish white, fingers sometimes blackish in
their basal half.
Length of carapace of largest specimen 9 millim., breadth 11 millim.
In the Indian Museum are 11 specimens from the Andaman Sea
at depths between 130 and 290 fms.
Alliance TI. Domecioida.
Domecia, Eydoux and Souleyet.
Domecia, Eydoux and Souleyet, Voy. Bonite, Crust. Zool. vol. i. p, 234: Lucas
in Jacquinot’s Voy. Astrolabe, Zool. vol. iii. Crust. p. 48.
Domaecius, Dana, Silliman’s Amer. Journ. Sci. and Arts, (2) XII. 1851, p. 128,
and U.S. Expl. Exp. Crust. pt. i. pp. 230, 251.
Domecia, A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873, p. 263, and Miss.
Sci. Mex. Crust. p. 345,
? Nelews, Desbonne and Schramm, Crust. Gandaloupe, p. 35.
Carapace somewhat oval transversely but much contracted pos-
teriorly, flat, somewhat hairy, with no trace of regions.
The fronto-orbital border is not much less than the greatest breadth
of the carapace. The front is profusely spinate, the spines being sharp,
a little curved, and falling into about six tufts or groups separated by
more or less well-marked intervals.
The antero-lateral borders pass backwards with but little outward
slope: they are a little shorter than the concave and convergent postero-
lateral borders, and are armed with numerous sharp curved spines.
The orbits are at the antero-lateral angles of the carapace and do
not conceal the eyes, their edge shows no fissures or sutures: their
upper and lower inner angles are broadly in contact, or almost in con-
tact, so as to exclude the antenne.
The antennules fold nearly transversely. The basal antennal joint
hardly reaches the front, though its outer angle is produced towards
the front: the flagellum is short—hardly as long as the orbit.
The buccal cavern is broad: the crests of the endostome are not
very strong; nor is the foliaceous process of the lst maxillipeds pro-
duced far forwards: the external maxillipeds are very large, and the
merus is remarkably broad and short.
The chelipeds are somewhat unequal, and are short and not very
massive: the arm is almost entirely hidden by the carapace: fingers
compressed, pointed.
230 A. Alcock — Carcinological Fauna of India. [ No. },
dni @
The legs are stout, especially the meropodites.
The abdomen of the male has all 7 segments distinct and separate.
151. Domecia hispida, Eydoux and Souleyet.
Domecia hispida, Eydoux and Souleyet, Voy. Bonite, Zool. vol. i. p. 285, pl. ii.
figs. 5-10: Dana, U. S. Expl. Exp. Crust. pt. I. p. 251: Lucas in Jacquinot’s Voy.
Astrolabe, Zool. vol. iii. Crust. p. 50, pl. iv. fig. 8-7: Stimpson, Bull. Mus. Comp.
Zool. II. 1870-71, p. 145: A. Milne Edwards, Nouv. Archiv. du Mus. IX. 1873,
p. 263, and Miss. Sci. Mex. Crust. p. 345, pl. Iviii. fig. 2 (not good): de Man, Archiv.
fiir Naturges. LIII. 1887, i. p, 326: Ortmann, Zool. Jahrb., Syst. VII. 1893-94,
. 478.
4 ? Neleus acanthophorus, Desbonne and Schramm, Crust. Guadaloupe, p. 35.
? Eupilumnus websteri, Kingsley, Proc. Ac. Nat. Sci. Philad. 1879, p. 397,
pl. xiv. fig. 3.
Carapace covered with light-coloured hairs: antero-lateral border
with five or six (including the orbital angle) acute dark-tipped spines,
and several similar spines on the carapace just inside the antero-latera!
border, and also just inside the spiny fronto-orbital border. The orbital
margin and the prominent edge of the epistome are finely denticulate,
Merus of the external maxillipeds extremely broad and short, with
an elevated patch of denticles on its outer surface.
Chelipeds a little unequal, the larger one is not very much longer
than the carapace: the arm, wrist, hand and dactylus are all studded
with acute spines.
Legs stout, not very much shorter than the chelipeds: the anterior
surface of the last four joints fringed with hairs, and the anterior edge
of the merus spinate, as also, but much less distinctly, is the anterior
edge of the carpus and propus.
Colours in spirit, yellow with brown blotches on the carapace and
chelipeds and indistinct dusky cross-bands on the legs.
In the Indian Museum are a male and female from off Little
Andaman I., 10 fms,, and two females from Great Coco I.
Alliance IV. Melioida.
Me ia, Latreille, Edw.
Melia, Latreille, Encycl. Meth. X. 705.
Melia, Milne Edwards, Hist. Nat. Crust. I. 431.
Melia, Dana, Silliman’s Journal (2) XII. 1851, p. 128, and U. S. Expl. Exp.
Crust. pt. I. p. 242.
Carapace rather depressed and narrow, hexagonal, not concealing
the first 25 or 3 abdominal terga even in the male, the regions not, or
fairly distinctly, delimited.
Fronto-orbital border more than ? the greatest breadth of the
carapace. Orbits very shallow, affording little concealment to the eyes.
Antennules folding obliquely.
1898.] A. Alcock —Carcinological Fauna of India. 231
Basal antennal joint slender, of good length but yet hardly touch-
ing the front ; the flagellum very long (half the length of the carapace,
or more), lodged in the orbital hiatus.
Chelipeds slenderer and much shorter than the walking-legs, the hand
often hidden in a matted tuft of hair.
Walking-legs long and stout, the third pair the longest of all.
External maxillipeds somewhat slender and almost subpediform.
Both the Indian species of this genus differ from Melia tessellata
(of which there are several specimens in the Indian Museum collection)
in having the carapace rugulose, the antero-lateral border crenulate,
the front more prominent, and the regions fairly well delimited and
areolated.
152. Mela cestifer, n. sp.
Carapace hexagonal, about as long as broad, rugulose or tubercu-
lous, somewhat pubescent posteriorly and laterally, the regions fairly
well defined and areolated.
Front broad, sublaminar, square-cut, horizontal but on a lower
plane than the gastric region.
Antero-lateral border cut into three blunt lobes, the first of which
is confluent with the outer orbital angle.
Antennary flagellum very long.
Chelipeds extremely slender ; hand hidden in a tuft of adherent hair,
which has to be removed before the slender hooked fingers can be seen,
First pair of legs somewhat more slender than the others, and
shorter than the last pair; the second and third pair stouter and
longer than the others, the third pair being the longest and the stoutest
(especially as to the merus) of all. All the legs are more or less
pubescent.
The abdomen of the male consists of 5 segments, the 3rd—5th being
fused.
Colours in spirit, white, the bases of all the rugosities or tuber-
cules defined by more or less circular very fine dark lines.
Length of carapace barely 4 millim., breadth hardly over 4 millim.
In the Indian Museum are a male and a female from off Ceylon,
34 fms.
153. Melia pugil, n. sp.
Differs from M. cexstifer (females compared) in the following
characters :—=
(1) the carapace is distinctly broader than long :
232 A. Aleock — Carcinological Fauna of India. [ No. I,
(2) the regions though as well defined are not nearly so much
broken up into tubercles :
(3) just behind the 3rd tooth of the antero-lateral margin is a
distinct indentation, making the anterior end of the postero-lateral
border dentiform :
(4) the chelipeds are distinctly stouter and the hand is concealed
in a fleshy glove :
(5) the first pair of legs is as stout as the fourth.
Length of carapace 5 millim., breadth 7 millim.
Colours in spirit, white, with a wider and more angular network
of fine dark lines.
In the Indian Museum is a single female from off Ceylon, 265 fms.
Appendix to Hyperolissa ?
PLaTyPILUMNUS, Wood-Mason.
Platipilumnus, Wood-Mason MS., Alcock, Ann. Mag. Nat. Hist. May 1894,
p 401.
Carapace hexagonal—the prominent bilaminar horizontally-pro-
jecting front forming the shortest side of the hexagon—thin, depressed,
perfectly flat, with the regions and subregions very faintly impressed :
the antero-lateral borders are spinate, the postero-lateral are slightly
convergent, and the posterior border is long.
Front about a third the greatest breadth of the carapace. Upper
margin of orbit spinate, the inner angle of the lower margin acutely
spiniform.
The antennules fold transversely. The basal antennal joint, though
of fair length, does not reach the front; the next joint lies loosely in
the wide orbital hiatus; the antennary flagellum is long, about twice
the major diameter of the orbit,
Buccal cavern quadrangular, very well defined anteriorly; the
external maxillipeds do not nearly cover it, but leave the efferent
branchial channels permanently widely open; the endostomial ridges
that define these last are well defined posteriorly, bnt do not reach the
anterior border of the buccal cavern. |
Chelipeds in the female, markedly unequal, fingers long, pointed.
Legs long, slender, compressed, spiny.
As there is only a single female in the Indian Museum, I cannot be
sure of the place of this genus in the system. It probably belongs to
the Cancroidea, and should be placed near Galene.
|
1898. | A. Aleock — Carcinological Fauna of India. 233
Platypilumnus gracilipes, Wood-Mason.
Platypilumnus gracilipes, Wood-Mason MS., Alcock, Ann. Mag. Nat. Hist. May,
1894, p. 401: Ill. Zool. Investigator, Crust. pl. xiv. fig. 6.
Carapace much depressed, perfectly flat above, with the surface
nearly smooth centrally and very finely and closely granular laterally,
and with the regions indistinctly defined. The front has the form of a
horizontally projecting bilobed lamella, with the free edge sharply and
very evenly spinate and the sides turned abruptly downwards. The
margins of the orbit are spinulate, the upper margin the more distinct-
ly so, and the lower margin terminates internally in a strong oblique
spine, the point of which inclines towards the sharply vertical tooth
formed by the already mentioned downfolding of the lateral edge of
the frontal lamella.
The antero-lateral borders of the carapace which are arcuate and
are shorter than the postero-lateral, are armed with three large spines,
in front of, between, and behind which are several spinules.
The pterygostomian regions are large and inflated, and the branchial
apertures, especially the efferent aperture, are large and patulous.
The eye-stalks are large and are of moderate length; the corneal
region is rather small.
The antennules are long and are transversely folded, their basal
joint is large and inflated.
The antenne are long, their basal joint is slender and free; the
second joint lies loosely in the internal orbital hiatus.
The inner edge of the meropodite of the external maxillipeds is
convex, with a pair of little spines at the summit of the convexity ;
the succeeding joint arises at the antero-internal angle.
The thoracic legs are furnished with many spines and long hairs.
The chelipeds, which are robust, are unequal; their prismatic arm
has all its borders spiny; the short inflated wrist is sharply granular
and spinulate in the distal half of its dorsal surface and along the
outer edge, while the inner edge bears a pair of rather large spines ;
the hand is spinulate everywhere in the smaller cheliped, but only in
the proximal third of its outer surface in the larger; the fingers also
of the smaller cheliped are spinulate on the outer surface, while those
of the larger cheliped are smooth ; the cutting-edges of the fingers are
finely and unevenly toothed.
The other thoracic legs are long, compressed, and slender, and have
the meropodite spiny along both edges, the carpopodite and propodite
spiny along the front edge, and the dactylopodite styliform.
Colour in the fresh state yellowish red.
Andaman Sea, 188-220 fms. A single female.
Ar. 30
234 C. B. and J. B. Bhaduri— On Double Thiosulphates [Now d,
Contribution from the Chemical Laboratory, Presidency College, Calcutta.
On Double Vhiosulphates of Copper and Sodiuwm.—By CHANDRA
BuusHan Bnuapuri, B.A., and Jyrotr BuusHan Buapuri, M.A.
Prem Cuanp RoycHanp Scuonar, Communicated by ALEX. PEDLER,
F.R.S.
CONTENTS.
Page.
Introduction ... sa ste a wee 234
Historical note .. ate oa wo. 24
Yellow double Ericenigistes, a wes COD
Action of cuprous oxide on sodium thiesnipliees ae a. 244
White double thiosulphate with sodium chloride ... .. 246
Yellowish white double thiosulphate ... Bes .. 246
When working with the preparation of a few normal sulphites and
particularly during an attempt to prepare normal cuprous and cupric
sulphites, we had occasionally to use sodium thiosulphate instead of
sodium sulphite or sulphur dioxide dissolved in water. It was soon
found that a beautiful yellow salt separates, on allowing a mixture of
sodium thiosulphate and copper sulphate solutions to stand for some
time, in microscopic needles which however decomposes readily in a day
or two unless special care is taken to get it perfectly dry. By varying
the concentration and proportion, with or without addition of alcohol,
the colour of the salts obtained seemed to change considerably. We
undertook to analyse some of them, and the result of the analyses forms
the subject matter of the present paper.
But before proceeding to describe our work a brief notice of the
work done by previous chemists on the subject may not be out of place.
The following list includes all the salts known at present.
Cuprous Sodium thiosulphate :—
(1) 2Cu,8,0, 7Na,S,0, 2H,O (Jochum C. C. 1885, 642).
(2) 20u,8,0,, 7 Na,S,0, 12H,O (Jochum).
(3) Cua, S 'O,;0 Na: S te 2 H ‘@. (Rammelsberg Poge. 56, 321).
(4) Cu,S,0, 3 Na, S, 0. 6H 0 (Jochum).
(5) 3Cu,S,0, 2Na,8,0, 8 H,0 (Vortmann, M. Ch. 9, 165).
(6) 3Cu,S,0, 2Na,8,0, 5H,O (Lenz, A. 40, 99).
(7) 5Cu,8,0, 4Na,S,0, 8H,O (Jochum).
(8).5 Cu,8,0, 4Na,8,0, 6H,O (Jochum).
(9) Cu,S,0, Na,S,O, H,O (Russel Ch. Ztg. 9, 233).
(10) Cu,S,0, Na,S,O, 3H,0 (Vortmann M. Ch. 9, 165).
(11) 5Cu,S,0, 3Na,8.0, 2Na,SO, H,O (Jochum).
1898. ] . of Copper and Sodium. 235
Cuprous Sodium thiosulphate Cupric Sulphide :—
(1) Cu,8,0, Na,S,0, CuS 4H,O (Lenz. A. 40, 99),
(2) Cu,S,0, Na,S,O, 2CuS (Kessel B. 11, 1585).
Cuprous Sodium thiosulphate Sodium Chloride :—
3Cu,8,0, 2Na,8,0, 4 Na Cl 8 H,O (Siewert, Zeit. ges.
Naturwiss. 26, 486).
Cuprous thiosulphate sodium dithionate :—
(1) Cu,S,0, 4Na,8,0, 4H,O (Vortmann),
(2) 2Cu,S,0, Na,8,0, 2Na,O (Jochum).
See Dictionary of Chemical Solubilities, Inorganic by A. M.
Comey, 1896, page 474, and Dammer’s Handbook IJ, 2 pages, 688-89.
The yellow salt to which Lenz and Siewert assigned the formula
Cu,S8,0,, Na,S,O,, Cu S,4H,O, was found by Kessel to have the
above composition when prepared at 10°C, and to vary in composition
with variation of temperature. Kessel further noticed that the yellow
salt is obtained when the copper sulphate and sodium thiosulphate are
mixed in the proportion of one molecule of the former to two molecules
of the latter. He further says that the reaction takes place in two
stages. According to him the yellow salt is decomposed by cold strong
hydrochloric acid forming a white mass which contains neither free
sulphur nor copper chloride, is permanent when dry, but in contact with
moisture decomposes with separation of copper sulphide and evolution
of sulphur dioxide. He is, however, of opinion that the sodium chloride
is present as an impurity having had nothing to do with the composi-
tidn. , |
Vortmann on the other hand, disagreeing with Kessel describes two
salts, one a greenish yellow salt Cu,S,0, Na,S,O0, H,Oand the
other an intense citron yellow salt 3Cu,S,0, 2Na,8,0, 8H,O. The
former according to him is obtained in microscopic needles when saturated
solutions of copper sulphate and sodium thiosulphate are mixed together
in the proportion of one molecule of the former to two molecules
of the latter. When, however, the solutions are previously heated
to about 40°C and then mixed together, the temperature of the mixture
rises about 5°—7° and the salt with the composition 3 Cu,S,O, 2Na,S,0O,
8 H,O separates out, also in microscopic needles. The yellow salt is
unstable and decomposes thus—3 Cu,8,0, 2 Na,S,0O, 8H,O=3Cu,S
+2Na,S0,+H,S0,+58,4+280,4+7H,0.
The salt we are going to describe agrees in some respects with the
yellow salt described by Lenz and Siewert and confirmed by Kessel, as
also with that described by Vortmann. That there should be copper
sulphide present in a salt with such an intense yellow colour seemed to
us strange. This led us to analyse the salt once again. To start with,
236 C. B. and J. B. Bhaduri— On Double Thiosulphates _[No. ],
it was found that the same yellow salt is invariably obtained at all
temperatures between 20°C and 34°C provided the green colour of the
mixture is not discharged by an excess of sodium thiosulphate. Even
with fairly dilute solution, the same salt is obtained. In one or two
cases, however, the sodium thiosulphate was so much in excess that the
liquor left after the separation of the crystals was yellow. Sometimes
the precipitate was allowed to remain in contact with the mother liquor
overnight. It may be mentioned here that the salt separates out soon
when prepared from concentrated solutions. If, however, dilute solu-
tions are used it takes a much longer time for the salt to crystallise
out, but it is richer in colour and more distinct in crystalline structure.
In contact with the mother liquor decomposition sets in generally
after a day or two, and a dark reddish brown precipitate, more soluble in
water than the yellow salt, is formed which finally changes into black
insoluble copper sulphide, the supernatant liquid becoming perfectly
clear and transparent. In the first few preparations the salt was
simply washed with water, in which it seemed to dissolve to a slight
extent, until free from sulphuric acid, pressed between filter paper,
powdered and dried by exposure to theair. It was found, however,
that the salt so treated did not keep well for a few days, decomposition
setting in, sometimes, even in course of a few hours. The colour
slowly changed to black, and sulphur dioxide was evolved. It was
subsequently found that a fairly stable salt could be obtained if it were
washed at first with water until free from sulphuric acid and finally
repeatedly with rectified spirit over filter pump, dried by exposure to
the air, powdered, and carefully sifted in a fine sieve. The salt thus
prepared did but lose a trifling when kept in a desiccator over calcium
chloride for weeks together. The result of a determination of the loss
is given here using about (7) seven grams of the salt.
22nd December, 1896. Platinum basin and salt =31:2436 grams.
23rd ” ” ” ” = 31°2425 33
9th January, 1897. oa ‘, =O) ‘240 2) roe
26th ss ms * = 31°2412
99
Thus about 7 grams of the salt lost only 2°5 milligrams in three
weeks. This loss may easily be accounted for from a small quantity of
alcohol still present in the air dried sample. In connection with the
decomposition of this salt at ordinary temperature, it may conveniently
be pointed out, here, that more dense the precipitate is and the richer
the colour, the better it keeps. Moisture is perhaps the most important
factor in bringing about the decomposition, as appears from a little of
the above sample, kept sealed in an ordinary glass tube. While the
substance kept quite well in a desiccator over calcium chloride for about
1898. ] of Copper and Sodium. 237
a month, the salt in the sealed tube began to change in the course of @
few days. It is also important to mention here that the salt does not
keep in contact with alcohol as was often noticed during specific gravity
determination. Alcohol therefore appears to have a twofold action
when used in washing the moist salt. Firstly, owing to its strong
affinity for water, it removes easily the last trace of moisture from the
salt, and secondly being in itself very volatile entirely disappears when
exposed to the air.
A series of analysis was made with samples prepared on different
occasions under varying conditions, and the result is tabulated on next
page.
From the percentage composition we obtain the following atomic
ratio :—
Ga: Nagy S:/O ‘= 514 +) 869 < 874): 1-984
14 : Dis 24: 56
oa (ie is) hav SPO
The composition obtained from the analysis of different samples agree
so well among one another that there is no reason to assume it to be a
mixture. On the contrary it may fairly be assumed to be a compound
‘with definite composition. It has also been found that different crops
of the salt obtained from the same mixture at different intervals have
identical composition (see samples B. 1 and B. 2).
On adding caustic soda toa mixture of cupric sulphate and sodium
thiosulphate, a blue precipitate is obtained indicating the presence of
bivalent copper. If, however, the caustic soda is added after some time
when the formation of the yellow salt begins, the yellowish red cuprous
hydrate is obtained. On adding caustic soda to the yellow salt sus-
pended in water, the latter is decomposed, and the same reddish yellow
cuprous hydrate is obtained. This indicates that the first stage of the
reaction consists in the reduction of the cupric copper to cuprous
copper, and it is during the second stage that the precipitation of the
yellow salt commences. The formation of the yellow salt is neither
sudden nor rapid, one or two days being sometimes necessary for
complete precipitation. It appears as probable that a portion of the
copper in solution undergoes oxidation with the reformation of copper
sulphate and reproduction of a blue solution.
The salt thus prepared was subjected to a careful qualitative
analysis .for all other possible sulphur compounds, as for instance sul-
phuric acid both free and combined, a sulphide, a sulphite, and the
thionates. It was found, however, that none of these compounds are
present in the salt even as an impurity. Butit gave all the reactions
[ No. 1,
On Double Thiosulphates
_“~
C. B. and J. B, Bhadur
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of Copper and Sodium. 239
1898. ]
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240 C. B. and J. B. Bhaduri— On Double Thiosulphates [| No. 1,
of thiosulphuric acid. Hence it was concluded that the substance was
a double thiosulphate of copper and sodium, and we confirmed it by
quantitative estimation of the acid. We estimated the thiosulphuric
acid both by the direct action of iodine on the salt acidulated with
hydrochloric acid, and by decomposing the salt with caustic soda,
filtering and titrating the filtrate by iodine after acidification. The
results are tabulated below :—
1 ¢.c. IODINE SOLUTION = ‘01516 Gram I. = ‘01365 Gram H,S,Ox.
A.
Table of direct titration with todine solution.
Wt. of Salt used Vol. of Thio- Vol. of Thio. Porane? f
(Sample VII). | sulphate required.| calculated for | Mean. Se 2
1 gr. Salt. cat ee
0 2909 gram. 10°6 c. c. 36°6 c.c.
01880 ,, 6:9 c.c. 367 c.¢. 36°63 50°
0°3467.— ,, 12°7 c. ¢. 36°6 c.c.
B.
Table of titration of the filtrate after decomposing the salt by NaOH and
filtration.
. Vol. of Thio.
Vol. of Thio- Percentage of
. of Sal d. = calculated f Mean.
Wt. of Salt used. | .uiphate required. = ees Salt. sees H.S,03.
+ |
S |
6 ( 1:0637 gram. 38°25 c. c. 36° Cc. ¢.
"8, [ 15558, 53°5 «C.c. 36'3 ¢. c.
a |
a | 36°2 49°3
O'6199. -.,, 220 c. ¢. 36°3 c. c.
1:9320 ,, 67°9 c.c. 35:1 c. c.*
On comparing the two tables it is found that the result of table A
is slightly higher than that of B. The total sulphur 27°95 °/, if caleu-
lated as thiosulphuric acid would give 49°8°/,; so that there is fair
agreement between the calculated result and that obtained by the
ti? a pha
* In this case the result is too low as the salt was slightly decomposed and
discoloured.
1898. ] of Copper and Sodium. _ 241
method A. This proves that the whole of the copper is present in the
cuprous state, otherwise the iodine required for direct titration would
be less, owing to the liberation of iodine by the cupric salt from potas-
sium iodide. The little discrepancy between A and B will be explained
later on.
Another important point should be mentioned here in connection
with the determination of water. It will be seen on going through the
table that when the water is estimated by heating the substance at
temperatures up to 250° C. in a currentof carbon dioxide and causing
the sulphur dioxide evolved to be absorbed by lead dioxide, we get
9°46 °/, of water. On the other hand when it is heated rather strongly
and the evolved sulphur dioxide is absorbed by red hot lead chromate,
the result is 10°5°/,. It was further noticed that the major portion of
the water was liberated before the evolution of any considerable amount
of sulphur dioxide began. During the decomposition of the salt a
small quantity of sulphuric acid is formed which with the sodium
sulphate, another product of decomposition, may form sodium hydrogen
sulphate, thus accounting for the increased percentage of water ata
very high temperature. We propose to take up the subject in a future
communication in connection with the decomposition of the salt.
It may hence be concluded that the substance is a double thio-—
sulphate of copper and sodium having the formula 7 Cu,S,0, 5 Na,S,O,
16 H,O with the following percentage composition :—
Copper = 32°24 °/,. Sulphur pa it ag ae
‘ Sodium 8°37 °/,. Oxygen (for complete oxidation) = 32°05 °/,.
Water = 10°48°/..
The salt with the composition as given above should however take
32'04.°/, oxygen for complete oxidation of the salt to copper sulphate
and sodium sulphate. There seems to be a slight difference between
the calculated percentage of oxygen and that obtained as a mean of
several analysis. This difference however disappears if the result of
the analysis of the last preparation, the most stable in the batch, is
considered.
V. Hauer says that an acid cuprous -thiosulphate of the formula
Cu,H, (S,0,), may be obtained by adding a concentrated solution of
copper sulphate to a strong solution of sodium thiosulphate until the
colour is deep yellow and gently heating the mixture, as yellow pris-
matic crystals. We have repeatedly tried to prepare the salt, but as
often have we failed. The crystals obtained, invariably contained
sodium thiosulphate, and when the heating was carried beyond 35-40°C.
the mixture in some cases changed colour from yellow to a dark reddish
J, u, 31
24:2 GO. B. and J, B. Bhaduri— On Double Thiosulphates [No.1,
brown in the course of a few minutes, and finally deposited a dark-brown
precipitate readily soluble in water. If, however, thoroughly saturated
solutions of both copper sulphate and sodium thiosulphate, previously
heated to about 50-60°C., be used, similar yellow crystaline precipitate
is obtained which settles down easily. Care should however be taken
in washing the salt, as a comparatively large quantity of sodium
sulphate is formed in this case which has to be removed. But the
freshly precipitated salt is also sensibly soluble in water. It should
therefore be drained as completely as possible, washed once or twice
with water and then with dilute alcohol gradually increasing the
strength of alcohol and finally with alcohol alone. Different samples
have been thus prepared with slight modification in the process, and the
result of the analysis of the same are included in the annexed table.
On examination it will however be found that although the samples
appear to be identical in physical properties, they all differ more or less
im composition and that therefore different formule should be given
to them. (See Table p. 243).
As we had reason to believe that the percentage of sodium in F
was a little too high the atomic ratio appears to be as Cu: Na: S: O:
H,O=4: 3: 7: 16: 6 corresponding to the formula 4 Cu, 8S, O,, 3 Na,
S,0,9H,O. With the following percentage composition :—
Cu=31'84 °/, S=28:17 °/,
Na= 868 °/, O=32°20 °/,
In E the ratio of copper to sodium is as 7: 6.
By heating therefore with sodium thiosulphate a part of copper is
separated, thus increasing the percentage of sodium and sulphur. But
whether at any particular temperature a compound of perfect definite
composition is formed or not, it is impossible to say positively.
We next proceeded to consider what takes place when copper acetate
is used instead of copper sulphate, thinking that acetic acid being a
weaker acid might give the normal copper thiosulphate or the acid salt
of V. Hauer. Copper acetate is less soluble in water than copper
sulphate, and accordingly we had to work with comparitively dilute
solutions. Thinking that concentration and relative proportion might
produce corresponding difference in composition standard solutions of
H,O=102 °/,
copper acetate and sodium thiosulphate (1 ¢.c. Thio. =10°835. S Thio. =
10°835 x -0248 gms. Thio. = :2687 gms. and 1 c.c. copper acetate=
"0182 gms. Cu) were prepared. Kessel and Vortmann state that the
yellow salt described by them can only be obtained when the two solu-
tions are mixed in the ratio of one molecule of copper salt to two
243
of Copper and Sodium.
1898. ]
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a Fe a g, <s - o
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24,4. C. B. and J. B. Bhaduri— On Double Thiosulphates [No. 1,
molecules of sodium thiosulphate. It was soon found that no preci-
pitate is obtained when the mixture is made in the ratio of one molecule
of the copper salt to two molecules of the sodium salt, and that the
same yellow salt is obtained when they are mixed in any other propor-
tion less than the above.
When the yellow salt is treated with a solution of sodium hydrate,
an orange coloured precipitate of cuprous hydrate is obtained. If,
however, the precipitation is effected in presence of a large volume of
water the bright reddish yellow dense precipitate becomes voluminous,
turns dirty brown and can easily be filtered. From concentrated solu-
tion the whole of the copper cannot be completely precipitated and
separated, a portion invariably going into solution. The light brown
precipitate after being thoroughly washed, dissolves in dilute hydro-
chloric acid; a small quantity possessing a black colour however remain
undissolved. No sulphur free or combined can be detected in the
solution. The insoluble residue can only be dissolved in aqua regia and
on analysis was found to contain copper and sulphur only. The residue
therefore consists of copper sulphide. It becomes now of importance
to decide whether the sulphide is present as such in a soluble form in
the original salt or that it isa product of a secondary reaction. We
are of opinion that the latter supposition is the correct one and we shall
presently give reasons for it.
As already stated the bright orange red precipitate cannot be
filtered easily. If, however, the mixture be very gently heated, the
precipitate settles down easily and can therefore be filtered readily.
Should it however be heated nearly to boiling the precipitate darkens
in colour. The residue after careful washing, on treatment with dilute
hydrochloric acid leaves a considerably greater quantity of the black
copper sulphide than that obtained in the cold. It is therefore clear
that the sulphide of copper which is found along with cuprous hydrate
is a product of a secondary reaction between sodium thiosulphate and
cuprous hydrate. That this is the true explanation of it will be evident
from the following consideration. Sodium thiosulphate is neutral to
test paper; so also is cuprous hydrate or oxide. But when cuprous
oxide is treated with a solution of sodium thiosulphate, a strongly alka-
line liquid is obtained even in the cold. According to Field (Quarterly
Journal of the Chemical Society xvi. 28.) cuprous hydrate dissolves in
sodium thiosulphate solution at the ordinary temperature and is repre-
cipitated on heating. We found, however, that with the application of
heat a considerably greater quantity of cuprous hydrate dissolves. To
examine this reaction quantitatively weighed quantities of cuprous
oxide containing 98'5°/, of real cuprous oxide were dissolyed in excess
1898. ] of Copper and Sodium. 245
of sodium thiosulphate with the application of gentle heat and the
sodium hydrate produced was estimated by standard acid, using phenol-
phthaliene as an indicator. The result is tabulated below :—
Vol. of the standard
hydrochloric acid re-
Vol. of standard
Weight of CugO used. ka aie acid quired for 1 pram Mean.
‘ of the oxide.
0°3162 grams. 21°7 G.c. 68°6 cc.
02558 _—=Cé=, 17°4 c.c. 68°4 c.c.
02928 .-.;, 20°3 c.c. 69°3 c.c. 68'8 c.c.
0°2539 _—sé=黓; 17°5 c.c. 68°92 c.c.
e76ss” ;, " © 921°9 ¢.c. 68°94 c.c.
1 c.c. of the standard acid used contained 000734 gram of true
hydrochloric acid gas.
...0°985 grams of real cuprous oxide liberated a quantity of caustic
soda which required 68°8 c.c. of the standard acid: or one gram of real
68°8 x ‘00734 x 40:
‘rehome 68°8 x *( KO" Ok 4
cuprous oxide liberates 365 0985 0562 grams of caustic soda.
1 562
“, Cu,0 : NaOH : : Tap i757 = 701 : 1405
— Ana el
or one molecule of Cu,O liberates two molecules of sodium hydrate
according to the following equation :—
Cu,0+H,0+Na,.S,0,=2 NaOH +Cu,8,0,
The cuprous thiosulphate is kept in solution by the excess of sodium
thiosulphate. This solution is very unstable under ordinary circum-
stances and black copper sulphide soon separates out. Here we have
the explanation of the formation of the black sulphide when the yellow
salt is treated with caustic soda, The reaction may therefore be repre-
sented by the following equation ;:—
7 Cu,8,0, 5 Na,S,0, 16H,0 +14 NaOH =12 Na,S,0,4+7 Cu,O + 23 H,0.
During the first stage of the reaction the whole of the copper is
obtained as cuprous oxide, but the bright redish yellow cuprous oxide
being, in contact with an excess of sodium thiosulphate in course of
time gradually darkens in colour and quite rapidly on heating as the
result of the other equation stated above, and subsequent conversion of
the cuprous thiosulphate into copper sulphide. It is for this reason
again copper cannot be precipitated as oxide by means of caustic soda
from a solution containing one mol. of copper sulphate to two or more
molecules of sodium thiosulphate. Here we have also the explanation
246 C. B. and J. B. Bhaduri— On Double Thiosulphates [No. ],
of the little discrepancy noticed between the thiosulphuric acid deter-
mined by direct titration (method B) and that obtained from the total
sulphur calculated as thiosulphuric acid.
The yellow salt described before dissolves easily in dilute hydro-
chloric acid without any apparent change. Concentrated hydrochloric
acid on the other hand precipitates a white crystalline salt which settles
down at once. After awhile, however, sulphur dioxide is given off freely
from both, and the supernatant liquid turns brown. The white salt if
washed immediately with a little water and finally with alcohol is a
rather stable one. It can in fact be heated in steam chamber without
decomposition. In contact with the mother liquor containing strong
hydrochloric acid it decomposes so soon that the washing of the salt
becomes difficult and at times impracticable. Besides in presence of a
large quantity of strong hydrochloric acid a proportionally large
quantity of sodium chloride is precipitated and cannot therefore be com-
pletely removed by alcohol. If instead of hydrochloric acid, acetic acid
be used, sodium acetate is formed and can be easily removed by washing
and filtration with dilute alcohol in which menstruum it is rather freely
soluble. We accordingly prepared at first a saturated solution of
copper chloride in dilute acetic acid and then gradually added to it a
strong solution of sodium thiosulphate, when a perfectly white salt was
precipitated. Subsequent experience proved that even acetic acid is
not necessary, and the white salt is obtained when a strong solution
of sodium thiosulphate is gradually added to an equally strong solution
of copper chloride. The precipitate is obtained in fine impalpable
powder which settles down easily and can therefore be filtered and
washed without any considerable difficulty. The result of the analysis
of the different samples is given in the annexed table. From the mean
atomic ratio Cu: Na: S: Cl:O0: HO=8: 3°01: 4'68 : 1:29
10°78: 1:91 the salt with the formula 9 Cu,8,0;. 5 Na 8,0, 8 Na Cl
12 H,O is obtained having the following percentage composition :—
Cu = 31:46 S = 24°78 = term
Na = 11°45 Cl = 7°85 H,O = 5:97
If strong solutions of copper sulphate or acetate and sodium thio-
sulphate be mixed nearly in the proportion of one molecule of the
former to two molecules of the latter and allowed to remain for some
time as in other cases, no yellow salt separates out. If on the other
hand a large excess of concentrated acetic acid be added to the mixture
a white or faintly yellowish white precipitate is obtained which is more
soluble in water than the yellow salt. Copper acetate should therefore
be used in preference to copper sulphate. It can then be washed with
1898. | of Copper and Sodium. 24:7
dilute alcohol instead of with water until free from sodium acetate and
finished as before. It appears as probable that the acetic acid plays
the same part in this case as alcohol does in Vortmann’s salts. The
white or faintly yellowish white salt turns yellow in contact with water
especially if the salt be previously dried. On comparison of the result
of analysis of the different preparations as given in the annexed table
we find that this salt again is a double thiosulphate of copper and
sodium having the formula Cu,S,0, Na,S,.0, 23 H,O. Russel describes
(Ch. Ztg. 9233) a salt Cu,S,0, Na,S,0, H,O whose properties and
method of preparation are altogether different from the one described
here. This salt when once perfectly dry, also keeps well even in con-
tact with the atmosphere.
248 C. B. and J. B: Bhaduri— On Double Thiosulphates [No. J,
Table of analysis of the white salt obtained by the action
rahe 5 5 ‘o © re zr)
m m
HS | Fi 3 A =n
£5 S| FS s be Fy
wo a0 on an fel) an an
Darr. fe | # ka Wile: Wich 2 ee
° » = » »
2| 32 | 24 | 22 | 29 | 25 | e4 | 25
pee pee pues | se Pee | ee
al Sh eee. ie E e E Ss
Jan. 14, 1897... D. i 0°214 ove coe 0°3898 Ty) | eee oe
| 0°3235 | 0°1281 | 0°1175 ee
D.2.| 0:3371 | 0°1402 oe 0°6296 ' : 2
0'2220 | 0:0798 ms a 0:061 us
‘|D.3.) 1:1468 | 0°:4592 | 0°4036 23 ‘ Br
0°7384 ms te 1'3463
0°4.602 . vee 3°1193 :
0'5071 one “ 0°1527
1°6106 ove . eve eee w- | 0°0931
D.4,| 0°9890 | 0°3869 | 0°3541 toe °
0°4552 eee “A . 3°042
0°8380 as sae 15119 *
0°7392 toe oo a 0°2352 «
15640 | 06160 | 0°5574.| ... ‘ 00909
15015
i i a
1898.]
of Copper and Sodium. 249
_ of hydrochloric acid (concentrated) on the yellow salt.
PERCENTAGE OF
Cu. | Na. Gi
a
25°05
31°53) 11°75
32°99 25°65
aoe | avs 68
32°08) 11°4
25°06
32°09
31°19) 11°6
cee «. | 248
31°43) 11°55
ATOMIC RATIO OF
REMARKS.
The yellow salt
was dissolved
in dilute hydro-
chloric acid and
then _ precipi-
tated by strong
hy droc hloric
acidjand washed
with alcohol.
3°00 | 3:06) 4°7
eve
The result is high
owing to thesalt
being — slightly
decomposed.
=
3:0 | 2:93! 4°63) 1-24) 10°77
|
eee
28°86 Deducting the
weight of Ag Cl.
Perfectly white
sample.
3°05| 47 | 1°34) 10°78] 1:93] Deducting the
\. weight of Ag Cl.
28°45
os oO OL
5°96
30
Mean 3:01) 4°68} 1-29] 10°78 1-91
50 (.B.and J. B. Bhaduri— On Double Thiosulphates, Sc. |No. 1, 1898.
_
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JOURNAL
OF THE
ASIATIC SOCIETY OF BENGAL
=e R Ge
Vol. LXVII. Part IIL.—NATURAL SCIENCE.
No. II.—1898.
An Annotated List of the Butterflies of the Ké Isles—By Lronet
pe Nickviuuz, F.E.S., C.M.Z.8., S§c., and Hetnrica Kian.
(With Plate I.)
[ Received November 25th, 1897 ; Read January 5th, 1898. ]
The Ké, Key, or Kei Isles,* called by the natives the Evar Isles, have
a total area of about 680 square miles, and have been in the possession of
the Dutch since A.D. 1645. They le south of the equator, also to the
south of Dutch New Guinea, to the south-east of the large island of Ceram
and the nearer small Banda group of isles, to the west of the Aru group
of isles, and to the north of the Timor Laut or Tenimber Islands. They
are placed between 5° '0’-6° ‘5’ S. Lat. and 131° -50’-133° -15’ E. Long.
They may be divided into four parts:—I. Great Ké Isle or Noehoe Ioet,
II. Little Ké or Noehoe Roa, III. the Tiandoe islets, IV. the Koer
islets. The last, according to Professor K. Martin, formed once a part of
the eastern extension of the continent of Asia, while the first three ap-
pertain to the Australian region. Great Ké has a tertiary formation,
consisting of limestone rocks; the surface is hilly, rising to nearly
3,000 ft. elevation. Little Ké and the other islands are all of coral forma-
tion, and are port-tertiary or quaternary. Wherever the soil and situation
are favourable, the islands are planted with cocoa-nut and bread-fruit
trees. The islands to the south of Little Ké are from the evidence of
the rocks of an older formation of the quaternary period. All the
* Pronounced “kay,” or exactly as the letter ‘‘K,” says Dr. A. R, Wallace in
“The Malay Archipelago, ”
J. u. 32
252 L. de Nicéville— List of the Butterflies of the Ké Isles. [No. 2,
islands shew in different places distinct old strand and beach lines, which
prove that they have been periodically raised by volcanic action. Ac-
cording to the natives, about sixty years ago one small island was so
raised above the level of the sea on the western side during the occur-
rence of a great earthquake combined with a tidal wave. Also during
recent years slight earthquakes are felt from time to time. On Little
Ké is found the largest area under cultivation, “batatas,’ a kind of
pea, and also beans, indian-corn and cocoa-nuts being largely grown.
In some places in Little Ké are yet found forests of iron-wood, “ ling-
goo-ah” and “loriah”” wood; also on the mountains of Great Ké. But
the forests are being cut down year by year, and become rapidly thin-
ner, the more so as the Ké islanders every year increase the building of
- “nraus”’ (native boats), which are sold to the people of the islands close to
the Ké archipelago. The natives live principally on sago, which is largely
cultivated ; but rice has to be imported by traders. The exports from
the Ké isles consist of logs of iron-wood; ‘‘ praus,” “ tripang”’ ( béche-de-
mer), turtle-shell, black-shell, green snail-shell, “copra” (dried cocoa-
nuts), and some “‘ beng koedoe”’ (Morinda bracteata, Roxb.), used for dyeing.
The whole population consists of about 24,000 souls, residing chiefly on
Great Ké, of whom about 14,000 are heathens, 9,500 Muhammadans, and
about 500 Roman Catholics, the latter resulting from a mission settled
there for about the last ten years. The people are of very mixed
blood, and are Malaio-Polynesians. Their colour varies from light to
dark brown and is of all shades. The hair is black, mostly long and
slightly curly. There are of course many with short frizzly hair des-
cended from immigrants from New Guinea. They divide themselves
into four classes:— I. The “ mel-mel” are the aristocracy, and are
chiefly descended from immigrated traders from Ternate, Luang, Ceram,
and Macassar in Celebes. From this class are mostly elected the chiefs,
of whom there are usually three or four in each village. II. The
““jam-ah,” who are perhaps the original native aristocracy, of whom
also different chiefs exist. III. The ‘renn-renn,’’ who are free citi-
zens. IV. The “tri-ri” or slaves, or better, bondsmen, Though
there are so many chiefs, they have very little authority over their
people, the family being the unit of government. The religion of the
heathen portion of the population is a very simple one. They have one
or two wooden idols erected in places they consider to be sacred, but
believe much more in ghosts than in their gods; to the former they pray,
and make offerings to them when commencing the cultivation of their
fields, going a journey, in times of distress, &c. In general the people
ate very lazy, and live from hand to mouth, never thinking of the mor-
row. A little sago and a fish suffices for each meal, and of fish there is
1898.] L. de Nicéville—List of the Butterflies of the Ké Isles. 253
an abundance in the sea for the trouble of catching. If they want
a new cloth they can obtain it without trouble on credit from the
numerous traders, paying for it hereafter with ‘“ copra,” ‘ tripang,” &c.
The Ké people consider themselves to be very courageous, but are in
reality cowardly. In the old days before the Dutch took over the gov-
ernment of the islands, the natives were often fighting amongst them-
selves. These little wars, often extending over a considerable period,
came to an end at last when one side or the other had killed a few of
their enemies, seldom more than four or five. Under enemies, women
and children were included as wellas men. The tribe or village losing
the fight and coming to the conclusion that peace was desirable, made
terms with the opposite side, and paid a war indemnity, sometimes in
land, but usually in old cannon (“‘lilas”’), rifles, gongs (“ tom-toms”’),
bracelets and chains of gold, plates, &c., &e.
The islands being very small have also a very poor fauna. The
mammals consist of wild pigs, two species of Cuscus, a flying squirrel, a
flying fox, some species of bats, rats and mice. On Great Ké a kangaroo
and a marsupial badger are found. Birds are more common, but no
paradise-birds occur.
The west monsoon blows from December till April, which is the
wet-season; the east monsoon also from May to November brings much
rain, alternating with fine clear days. The driest months are from July
to November. The mean humidity of the atmosphere of the Ké Isles
in 88°5°/,, and the mean temperature is 80°9° F, (27:4 Cels.), according
to the observations of Mynheer H. C. W. Plauten, an officer of the
Dutch navy.
Great Ké possesses many small rivers, which form here and there
beautiful waterfalls in their course from the mountains to the sea. In
the east monsoon they are usually dry. On Little Ké, in which there
are no hills, there is only one river, which is about a mile and a half
long, but which has an outflow of about two cubic meters a second, and
what is most strange, has a greater flow of water in the dry- than in the
wet-season, no doubt from springs at its source or in its course. Besides
the rivers there are in Great Ké one and in Little Ké two small lakes,
which have no outlet, and seem to be merely accumulations of rainfall
in natural depressions in the ground.
From what has been said above, it will be understood that the en-
tomological fauna of the Ké archipelago is poor, but strangely enough
(at any rate as far as the butterflies are concerned) it has more distinct
endemic species than some of the neighbouring islands which are larger.
For instance, three out of the six known species of Muplea found on
the islands are endemic to them, and three of them are very unusually-
254. L. de Nicéville— List of the Butterflies of the Ké Isles. [No, 2,
coloured animals, being more or less pure white on the upperside in
both sexes, and are mimicked by the females of two peculiar species of
Hypolimnas (H. polymena, Felder, and H. hewitsoni, Wallace). H. M.S.
“Challenger” in her memorable scientific expedition round the world,
called at Ké Dulan, and Dr. A. G. Butler, in Ann. and Mag. of Nat.
Hist., fifth series, vol. xiii, p. 188 (1884), enumerates thirteen species
of Butterflies from thence obtained on that occasion, all of which are
noted below. The late Herr C. Ribbe in his paper on the Butterflies
of Great Ceram (Iris, vol. 1, p. 187 (1889), mentions many species
from the Ké Isles, all of which are referred to herein. Herr J. Rober
of Dresden in Tijd. voor Ent., vol. xxxiv, pp. 261-334 (1891), vol. xxxv,
pp: 85, 86, pls. iii—vi (1892), has written a most interesting paper on
the Lepidoptera of the group of Malayan islands which include Ceram,
Goram, Maumerie, Key, Flores, Letti, Alor, Luang, Bonerate, Timor,
Timor Laut, Kisser, Wetter, and Tanah Djampea, recording 42 (not 41
as stated at page 262) species from the Ké Archipelago. All these are
mentioned below, and those not seen by us have a ™ prefixed to their
names. The present paper is based on materials obtained by Kithn
during the nine years (from 1889) he has resided on the islands, and
he has written the introduction and notes on the habits of the various
species, while de Nicéville is responsible for the rest of the paper, and
has seen it through the press. It is probable that the list is fairly
complete, there are probably only a very few small species left to be
recorded. We record 128 species from the Archipelago, out of which
we have not seen only 17; of these latter most of the names are doubt-
less incorrect identifications, and appear in our list under other names.
Family NYMPHALIDA.
Subfamily Danaina.
1. Hustra (Nectaria) D’URVILLEI, Boisduval.
In Great Ké this species is usually seen flying in the forest at a
great height amongst the trees. It is very rare in Little Ké.
2. Danais (Tirumala) wamata, Macleay.
Very seldom seen in Little and Great Ké, common on Koer Isle.
De Nicéville finds it difficult to distinguish D. septentrionis, Butler,
from this species. Macleay’s species is much the older,
3. Danats (Limnas). prtiuia, Stoll.
Key, Rober, as Danaus chrysippus, Linneus. The Ké Isles examples
agree with those from Australia. It is very rare on all the islands.
ES OO , OOD ee en er nS
eS a
1898.] lL. de Nicéville— List of the Butterflies of the Ké Isles. 255
4, Danats (Salatura) varatensis, Butler.
Rober as Danaus plexippus, Linneus. Originally described from
Larat in the Timor Laut Islands. Single specimens are seen every-
where on Great and Little Ké Islands all the year round, mostly on
open spots covered with “alang-alang” grass (Imperata arundinacea,
Cyrill.).
In de Nicéville’s collection is an old male specimen of D. intensa,
Moore, described from Java, Lombok and Borneo. As the species is un-
known to Kihn, and is represented by D. laratensis in the Keé Isles, it is
probable that the specimen did not come from the Ké Isles.
5. Damais (Salatura) arrinis, Fabricius.
Key (Réber), Ké Dulan (Butler). Very common on all the islands.
6. Danas (Asthipa) cirrina, Felder.
Rober as Danaus gloriola, Butler. Common on Great and Little
Ké Islands amongst bamboo clumps. Mr. Kirby has given some notes
on this species in Ann. and Mag. of Nat. Hist., sixth series, vol. iv,
p. 157 (1889), but they are difficult to follow without seeing the speci-
mens about which he wrote.
7. TrLieRvo zorLus, Fabricius.
Rober as Hamadryas nais, Guérin. Butler as H. niveipicta, Butler,
Ann. and Mag. of Nat. Hist., fifth series, vol. xiii, p. 191, n. 6 (1884),
from Ké Dulan. This “new species” hardly seems to differ from
T. zoilus. Common everywhere in the bush in all the islands. The male
secondary sexual characters in the genus Tellervo (= Hamadryas, Bois-
duval, nec Hamadryas, Hiibner) do not appear to have been described.
They are found in the forewing, and consist of a greater sinuosity of
the inner margin than iu the female; with a large clump on the upper
surface of modified dull (instead of intensely black like the rest of the
ground-colour) black scales from the inner margin commencing at the
base of the wing but not reaching the outer margin, and extending for-
wards as far as the first median interspace, these modified scales being
entirely absent in the female.
8. EKurrm@a (Vadebra) nuryPon, Hewitson.
Moore. Rober. Ké Dulan (Butler). Common on all the islands in
October and November, but single specimens are found all the year round.
Mr. F. Moore in his monograph of the subfamily (Proc. Zool. Soc.
Lond., 1883, p. 284, n. 2) places this species in his genus Chirosa, in which
256 L. de Nicéville— List of the Butterflies of the Ké Isles. [No. 2,
he is followed by Dr. Butler in 1884and by Col. C. Swinhoe (Journ, Linn.
Soe. Lond., Zoology, vol, xxv, p. 342 (1896), that genus coming into
Mr. Moore’s second section of the Hupleina with * One ‘sexual mark’ or
scent-producing organ on the forewing.” FH. ewrypon has no male
secondary sexual characters whatever, and therefore comes into Mr.
Moore’s first section “ No ‘sexual mark’ or scent-producing organ on
the forewing,” and seems to fall best into the subgenus Vadebra.
9. Evueta@a (Chanapa) sacervos, Butler.
' Originally described from Larat in the Timor Laut Islands. Dr.
Butler says that the discal series of white spots on the hindwing are not
[ posteriorly ] notched, but in some of our specimens though not in others
as many as five are occasionally notched. The species is very Sommon
on the Tiandoe and Koer islets, but only two specimens have been taken
on Great Ké, and none on Little Ké,
10. EHvria@a cALLitHo#, Boisduval.
Mr. Moore in his Monograph of the Lupleina, p. 305, places this
species under the genus Salpinx, but according to our specimens and
Dr. O. Staudinger’s figure of the male from New Guinea in Iris, vol. viii,
p- 159, pl. iv, fig. 1, male (1895), it is a true Huplea. It occurs only
on Great Ké, never found on Little Ké or the adjoining islets.
11. EHurre@a (Calliplea) HoprreRI, Felder.
Rober. This species has been figured by Col. Swinhoe in Journ.
Linn. Soc. Lond., Zoology, p. 342, pl. xvi, fig. 1, male (1896). Found
on all the islands everywhere. It is the rarest species of the genus,
except the species which next precedes.
12. Evrie@a. (Calliplea) visenpa, Butler.
Originally described from Maroe Island of the Timor Laut group.
Found in the Ké Islands group only on Tiandoe and Koer, where it is
very common and extremely variable in the size and extent of the white
markings, no two specimens being exactly alike. Some specimens are
very small, perhaps the smallest in the genus, expanding only 2°0 inches
in alar expanse.
13. Hurpn@a (Hirdapa) asstmiuata, Felder. Plate I, Fig. 1, larva.
Moore as H. fraterna Felder, from Ké Island. Rober. Butler as
H. fraterna, from Ké Dulan. This species was originally described
from the “ Arru Islands,” while HL. fraterna, Felder, was described from
a female only from the same islands, on the same page. Probably the
1898.] LL. de Nicéville— List of the Butterflies of the Ké Isles. 257
two species are really one. It is very common on nearly all the islands
of the Ké group, but it seems rare on Koer. Mr. Kihn has bred it.
The larva is black; each segment bears several yellow stripes, the first
of which extends lower than the others and encloses the black spiracle.
The head, abdomen and legs are entirely black. The body bears
four pairs of long fleshy subdorsal tentacles tapering to a point, each
is basally carmine-red, becoming black towards the apex, the first pair
is placed between segments two and three, the second between three
and four, the third between four and five, the fourth between ten and
eleven. The pupa is greenish and metallic.
The Hupleas of the Ké Archipelago are very interesting, and form
four distinct groups as regards coloration and markings :—I, H. ewrypon,
Hewitson, H. hopfferi, Felder, and H. assimilata, Felder, which are
mimicked by the females of Hypolimnas polymena, Felder, and H. hewit-
sont, Wallace; II, H. sacerdos, Butler; III, #. callithoé, Boisduval ; IV,
EF. visinda, Butler.
Subfamily Satyrinz.
14, Myocauesis (Calysisme) Perseus, Fabricius.
Occurs on Great and Little Ké in fields of “alang-alang” grass,
but is not common. The ocellated wet-season form appears to be the
only one found on the islands, In both sexes the ocellus in the first
median interspace of the forewing on the upperside has a small pure
white pupil. |
15. Mycauusis (Mydosama) strivs, Fabricius.
Little and Great Ké Isles. Much rarer than the preceding species.
“ Sutyrus” manipa, Boisduval, and Mycalesis daidis, Hewitson, are
synonyms.
16. Yprnima arctous, Fabricius.
Rober. Occurs commonly everywhere on all the islands, and fre-
quents meadows.
17. Hipto consrantiaA, Cramer,
Rober and Ribbe as Melanitis crameri, Butler, described from New
Britain, equals “ Cyllo” amabilis, Boisduval, from New Ireland. The
type of the genus Hipio of Hiibner is the present species. It is doubt-
fully distinct from the genus Melanitis, Fabricius. H. constantia is
rather rare in all the islands, keeping chiefly to bamboo and sago scrub,
Colonel Swinhoe has described a single male of this species from “ Ké
258 L. de Nicéville — List of the Butterflies of the Ké Isles. [No. 2,
Island” as M. gylippa in Ann. and Mag. of Nat, Hist., sixth series, vol.
xii, p. 255 (1893), collected by Halliburton. Swinhoe does not refer
to H. constantia in his description. Our females from the Ké Isles
agree very well with Cramer’s original figure of that sex, which was
described from the Molucca Isles.
Subfamily Etymnuna.
18. Enymntas (Dyctis) mMELANE, Hewitson.
Rare on all the islands, usually seen flying about swampy places in
the jungle.
Subfamily NympHatina. N
19. CupnHa cCRAMERI, Felder.
Common everywhere on Little Ké, but more so in forests than
elsewhere, flying close to the ground between low bushes. Our speci-
mens agree fairly well with Cramer’s figures of this species from
Amboina in Pap. Ex., vol. 11, pl, exlviii, figs. D, D (I, F in text), male
(1777), as “ Papilio” lampetia. Dr. Aurivillius says that these figures
equal the “ Messaras”’ cramert of Felder, nec “ Papilio” lampetia of
Linneus, as indeed was pointed out by Dr. Felder when renaming
Cramer’s figures.
20. ATELLA EGISTA, Cramer.
From Little and Great Ké. Flies high and is seldom caught ;
keeps to the tops of bushes.
21. Cerrnosta cypALima, Felder. Plate I, Figs. 2, larva; 2a, head
of larva; 2b, 2c, pupa.
Rober. Ké Dulan, Butler, as C. insulata, Butler. Ribbe as
CO. cydippe, Linneeus, var. damasippe, Felder. Dr. Butler in Cist. Ent.,
vol. i, p. 165, n. 37 (1873), described C. insulata from Ké Island. He
does not give the sex of the types. It is probably the same as C. cyda-
lima, originally described from the Arru Islands. The species on Little
and Great Ké is not common, but is less rare at the beginning and end
of the wet than at other seasons. Mr. Kiihn has bred it, the larva being ~
of the usual form, with six compound black spines on each segment, the
dorsal pair twice as long as the two lateral pairs; the ground-colour of
the larva deep black, each segment posteriorly with a pair of sulphur-
yellow bands reaching below the spiracles. The pupa is whitish, clouded
with pale brown, with four pairs of golden spots on the dorsum.
1898.] lL. de Nicéville— List of the Butterflies of the Ké Isles. 259
22. CRrTHOSIA LAMARCKII, Godart.
Occurs only on Tiandoe and Koer Islets, not found on Great and
Little Ké Isles. |
23. Cyntuia crcnta, de Nicéville.
C. cycnia, de Nicéville, Journ. A. S.B., vol. Ixvi, pt. 2, p. 547, n. 4, pl. iii,
figs. 19, male; 20, female (1897).
Little and Great Ké. Single specimens are occasionally caught
on flowers.
24. Precis 1pa, Cramer.
Rober. Very common on all the islands all the year round.
25. Precis zetima, Fabricius.
Kihn has sent one male specimen of this species to de Nicéville. In
the collection of the latter are several examples of both sexes from
Mackay in North Australia, with which the one from the Ké Isles
agrees exactly. The species may be known from P. ida, Cramer, by
having on the underside of the hindwing a decreasing series of two,
three, or four creamy-white spots commencing on the middle of the
costa posterior to the costal nervure. It is a little doubtful in
de Nicéville’s opinion if this character is really of specific value, as in
the common P. iphita, Cramer, of India, the same variation is of fre-
quent occurrence.
26. JUNONIA EXPANSA, Butler.
Precis expansa, Butler, Proc. Zool. Soc. Lond., 1883, p. 367, n. 5.
Rober as J. erigone, Cramer. Described from females from Larat
in the Timor Laut Islands. In the Ké Isles it is rather variable, the
tone of the ground-colour of both wings on both surfaces being much
darker in some specimens than in others. The male is much darker
coloured than the female, especially so on the underside. In Little
and Great Ké it is very common all the year round.
27. *Junonta oriTHyi4, Linneus.
Rober as J. orithya [sic!]. This is we think a very doubtful
record ; at any rate it will not be the typical form of the species which
is found in the Keé Isles.
28. JuNoNIA VILLIDA, Fabricius.
Very rare on Little and Great Ké.
J. u. 33
260 L. de Nicéville— List of the Butterflies of the Ké Isles. [No. 2,
299. *JUNONIA TIMORENSIS, Wallace.
Ké Dulan (Butler). We have not seen this species from the Ké
Isles.
30. Nepris (Rahinda) consiminis, Boisduval. Plate I, Figs. 3,
young larva on food-plant ; 3a, full-grown larva.
Not very rare on Little and Great Ké, single specimens being
here andthere met with. Mr. Kihn has bred this butterfly. The ovum
is laid singly on the underside of the leaves of a papilionaceous shrub,
it is shaped like a raspberry, is pale yellow, and slightly hairy, and
furnished with raised knobs or tubercles in seven rows, the largest row
having 14 tubercles. Larva dark olive-brown, shagreened ; head bifur-
cated in the dorsal line; the second, third and fifth segments furnished
with a pair of processes, the pair on the third segment the longest ;
there is a pale dorsal line from the third to the thirteenth segment, and
a similar spiracular line. For protection against its enemies the larva
has developed the following curious habits :—It bites half through the
middle of one of the bipennate leaves of its food-plant, which is pro-
bably a species of Acacia, and also bites through entirely the small
leaves at the end of the stem beyond the half-bitten-through part, but
fixes each of these bitten-off leaves to the stem by a thread. The larva
rests on the underside at the extreme end of the stem, which has bent
over at a right-angle from the unbitten portion, and feeds on the faded,
dried-up, brown leaves, which very quickly become of the same shade of
colour as the larva. When touched, the larva shakes the leaves. The
larva is very sluggish, and moves very slowly, step by step.
ol. Nepris tacraria, Butler.
Athyma lactaria, Butler, Ann. and Mag. of Nat. Hist, third series, vol. xvii,
98, n. 1 (1866); Neptis lactaria, de Nicéville, Journ. A. 8S. B., vol. Ixvi, pt. Z,
p. 585, n. 2, woodcut of female (1897).
Rare on Little and Great Ké Isles.
1898.] L. de Nicéville— List of the Butterflies of the Ké Isles. 261
32. Neptis (Phedyma) necteEns, de Nicéville.
N. (Phedyma) nectens, de Nicéville, Journ. A. S. B., vol. Ixvi, pt. 2, p. 548, n. 6,
pl. i, fig. 8, female (1897).
Very rare on Little and Great Ké, not known from the other
islands.
33. *NeEPTIS VENILIA, Linneus.
Ribbe records this species from the Key Isles as “ Athyma ” venilia,
but we have never seen it from thence..
o4. Hypotimnas poutna, Linneus.
_ Rober. Common in December and January on all the islands, but
single specimens are met with all the year round. ‘he form of the
female occurring in the islands is that called Papilio tphigenia by
Cramer, Pap. Ex., pl. lxvii, figs. D, E (1775), from Batavia in Java.
The male is normal.
35. HypoLimnas poLyMENA, Felder.
Rober as H. alimena, Linneus, var. heteromorpha, Rober. The
male of this species is typical H. alimena, Linneus, but the female,
which gives its name to the species, has been named H. polymenu by
Felder, from the Arru Isles, and H. heteromorpha by Rober. It is a mimic
of Huplea assimilata, Felder, and the other two similarly coloured
Hupleas found in the Ké Archipelago. It is common in the wet season,
and has been figured by Swinhoe in Journ, Linn. Soc. Lond., Zoology,
vol, xxv, p. 342, pl. xvi, fig. 2, female (1896).
36. HyPoLiIMNas HEWITSONI, Wallace.
Apparently rare in the Ké Archipelago, Kiihn having only obtained
three or four worn male specimens on Tiandoe. Both sexes of this fine
species were described and figured from the Ké Islands by Hewitson in
Proc. Zool. Soc. Lond., 1858, p. 464, pl. liv, figs. 2, male; 1, female, as
Diadema pandarus, Linneus, and Wallace in Trans. Ent. Soc. Lond.,
1869, p. 282, n. 8, named it Diadema hewitsoni from the same place. The
female, which we have not seen, is heavily marked with white on both
wings on the upperside, and doubtless mimics the three similarly
marked species of Hupleas found on the Ké Isles.
37. PAaRTHENOS BRUNNDA, Staudinger.
Rober. This is quite distinct from P. sylvia, Cramer, from Java,
and P. sulentia, Hopffer, from Celebes. Single specimens on Little Ké
and Great Ke are found all the year round.
«
262 L. de Nicéville— List of the Butterflies of the Ké Isles. [No. 2,
38. Evuraria (Leaias) zropus, Linneeus.
Ribbe as Symphedra aeropa. Occurs only on Great Ké on the tops
of hills, and is very rare, Not found on Little Ké Island.
39. SaLAMIs SABINA, Cramer.
Sometimes rare, at other times common, on all the islands.
40. DOoLESCHALLIA AUSTRALIS, Felder.
Ké Dulan (Butler). At times scarce, sometimes common, on Little
and Great Ké Islands.
41. DoLESCHALLIA POLIBETE, Cramer.
Commoner than the preceding species on Little Ké and Great Ké
Isles. It is very interesting that two such distinct species should be
found together on one small group of islands. D. polibete appears to be
found in the Himalayas, Assam, Burma, South India, Ceylon, the Anda-
man and Nicobar Islands, and again in Lombok, Amboina, and the Ke
Islands. It was originally described by Cramer from Amboina in Pap.
Ex., vol. iii, pl. ccxxxiv, figs. D, E, female (1779) ; Cramer’s figure
under the same name on pl. ccxxxv, figs. C, D, male, also from Amboina,
has been named D. cramert by Distant in Ent. Month. Mag., vol. xxii,
p- 41 (1885), and is a quite distinct species. The Lombok, Amboina and
Ké Islands form differs from the form from the other localities named
above as D. polibete in having the four subapical white dots on the fore-
wing much more strongly developed, the more western form having them
more feebly developed and sometimes entirely abseut, but this solitary
character is one on which it is hardly sufficient to base a distinct
species. D. bisaltide, Cramer, and D. pratipa, Felder, seem to be one
species, which is found in the Malay Peninsula, Sumatra, Nias, Java,
Bawean, Borneo, Bali, Lombok and the Philippines.
42, CHARAXES KEIANUS, Rothschild. Plate I, Figs. 4, 4a, 4b, pupa.
C. pyrrhus keianus, Rothschild, Nov. Zool., vol. iv, p. 508, n. 2 (1898).
A rare butterfly on Little and Great Ké Islands. Mr. Kihn has
bred the larva, which feeds on Albizzia sp., and also Mesua ferrea (Iron-
wood). The pupa is of the usual shape, very broad, rounded, smooth,
with some small knobs only round the cremaster. In colour it is
pale green, with snow white stripes and dashes, This species is des-
cribed by de Nicéville in Journ. Bomb. Nat, Hist. Soc., vol. xii, p. ,
n. 8, pl. Z, figs. 13, male; 14, female (1898), When describing it
de Nicéville did not know that it would subsequently be named by
the Hon. Walter Rothschild.
1898.] L. de Nicéville—-List of the Butterflies of the Ké Isles. 263
Family LEMONIIDA.
Subfamily LisytHmaina.
43. LipYTHEA ANTIPODA, Boisduval.
Herr Kiihn has taken a single specimen of this species, and saw
one other. The identification is his, de Nicéville has none from the
Ké Archipelago.
Family LYCASNID.
44, GepRyDUS acragas, Doherty.
Not very rare in Little Ké Island. Most frequently caught on the
young leaves of a species of Sambucus at rest amongst black ants, the
butterfly as well as the ants probably feeding on the sap of the leaf-buds.
The male has a small oval whitish patch surrounding the swollen base of
the third median nervuie on the upperside of the forewing; no other
markings whatever on the upperside. The female has the markings on
the upperside exactly as described for the species by Doherty. Both
sexes on the underside of the forewing have the markings exactly as
described by Mr. H. H. Druce in Proc. Zool. Soc. Lond., 1895, p. 561,
pl. xxxi, figs. 9, male; 10, female, for G. vincula from Borneo. Doherty
unfortunately does not say if the submarginal band of confluent mark-
ings extends from the apex to the outer angle or not. G. vincula differs
from G. acragas in the female being uniformly dull brown without
markings on the upperside of the forewing. The “ Miletus” chinensis,
var. ceramensis, Ribbe, from South and East Celebes, Amboina, Saigun,
Buru and Borneo (Iris, vol. 11, p. 247, n. 95, pl. v (nec i), fig. 2, female
(1889), appears to be quite the same as Geryden acragas, and has two
years’ priority.
45. PITHECOPS BASSARIS, de Nicéville.
P. bassaris, de Nicéville, Journ. Bomb. Nat. Hist. Soc., vol. vii, p. 327, n. 4,
pl. H, figs. 4, male; 5, female (1892).
Rober as Hupsychellus (n. g.) dionisius, Boisduval. Ribbe as Plebejus
dionysius, Boisduval. It is found commonly on Little and Great Ke
Islands during the wet season on roads and paths through swampy
bush country. The genus Hupsychellus is a synonym of Pithecops.
46. *NeropirHEecops zALMORA, Rober.
Plebeius lucifer, Rober, Iris, vol. i, p. 61, pl. iv, fig. 5 (1886).
Not seen from the Ké Isles by us. It was described from the Aru
and Key Islands as P. lucifer by Rober. Herr Rober kindly sent
264 L. de Nicéville— List of the Butterflies of the Ké Isles. [No. 2,
de Nicéville a specimen of this species named by himself from the Aru
Isles, which enables de Nicéville to say confidently that Plebeius
lucifer is a synonym of Neopithecops zalmora. In vol. 11 of Dr. O.
Staudinger’s and Dr. EH. Schatz’s Exotische Schmetterlinge, p. 273,
n. 32, pl. xlviii (1892), reference is made to Herr Rober’s genus
Papua, the type of which is based on Plebeiws lucifer. Consequently
Papua falls to Neopithecops.
47. MercaispA MALAYA, Horsfield,
Mr. de Nicéville has seen four specimens only from the Ké Isles
of this widely distributed species, none of which have tails. It keeps
chiefly to the tops of fruit trees, and is apparently rare, but is probably
less often seen than it would be if larger, brighter coloured, or
haunting lower stations. It occurs only as far as is known on Little
Ké Island of the Ké Archipelago.
48, CYANIRIS KUEHNI, Rober.
Plebeius kuhni, Rober, Iris, vol. i, p. 60, pl. iv, fig. 29, male (1886).
Described by Réber from Hast Celebes and the Key Islands. It is
very close to the widely-spread C. puspa, Horsfield. Especially found
on Little but also on Great Ké Island. It is partial to the flowers of
Leguminose.
49. *CYANIRIS cAGAYA, Felder.
Rober as Plebeius cagaya. We doubt the occurrence of two distinct
species of Cyaniris in the Ké Islands. C. cagaya was described from
the Philippine Isles.
50. ZizERA OTIS, Fabricius.
Common on Little and Great Ké Isles on roads and paths flying
amongst the grass and low-growing herbs.
51. ZizerRa GaikA, Trimen.
Very rare, found only on Little Ké, though probably often over-
looked. It is the smallest butterfly found in the islands, some of our
specimens expanding only ‘6 of an inch.
02. ‘ PLEBEIUS”’ TUALENSIS, Rober.
P. tualensis, Réber, Iris, vol. i, p. 61, pl. v, fig. 26 (1886).
Originally described from the Key Islands, where it is very rare
on Little and Great Ke, and usually caught on flowers of the Legu-
minose. The male on the upperside is coloured like a typical species
\
a
1898.] Li. de Nicéville— List of the Butterflies of the Ké Isles. 265
of the genus Nacaduba, being dark dull purple, with a narrow external
black border. The female on the upperside is dull plumbeous, not dull
purple like the male. The markings on the underside are more similar
to those of the genus Zizera than to those of any other genus known to
us. It has no tails. The neuration differs from both the above-named
genera in that the first subcostal nervule is entirely separated from the
costal nervure in the forewing. It has a near relation in “ Lycena”
meerens, Rosenstock, from tropical Northern Australia, but the male of
that species has the wings narrower, the apex of the forewing more
acute, and the outer margin straighter, the ground-colour of both wings
on the underside darker, and all the markings consequently less promi-
nent. Lyceena merens has been placed by Messrs. Anderson and Spry
in “ Victorian Butterflies,” p. 92, woodcut of male (1894), in the genus
Holochila.
53. Pspupopipsas tras, Felder.
Decidedly rare on Little and Great Ké Isles. Our specimens agree
very well with “ Holochila”’ intensa, Butler, from the Aru Isles and
New Guinea. Unfortunately the description is not comparative with
P.ilias, It has a very quick flight, and sits in the hot sun on the tops
of certain bushes with rounded leaves. Rober records it from Key as
Philiris (n. g.) tlias, Felder. But ‘‘ Thecla” ilias appears to be conge-
neric with Pseudodipsas eone, Felder (1860), the type of the genus
Pseudodipsas, so Philiris would appear to fall to Pseudodipsas. The
genus Holochila, Felder (1862), cannot stand, being preoccupied in
Mammals, and Erina (1832-33), Swainson, cannot be used, as it is
based on the typical species, ‘‘ Papilio” erinus, Fabricius, from Aus-
tralia. Polycyma, Scott (1890), appears on plate xii of Scott’s “ Aus-
tralian Lepidoptera” for Polycyma carythx, itself asynonym of Papilio
erinus, and is a year older than Rober’s name Philivis. All these
names appear to be synonymous with Pseudodipsas. The genus Canda-
lides, Hiibner (1816), type Rusticus Adolescens xanthospilos, Hibner, is
perhaps the oldest name for this group of butterflies.
54, THysonitis TrioPus, de Nicéville, n. sp.
Hasitat: Great and Little Ké Islands.
Expanse: ¢,1°8to 2:0; 92, 2°0 inches.
Description: Femate. Nearest to T. apollonius, Felder, var. supous,
H. H. Druce and Bethune-Baker, Proc. Zool. Soc. Lond., 1893, p. 542,
pl. xlv, fig. 7, female, from Wammo Dobbo in the Aru Isles ; differs in
being “4 of an inch greater in alar expanse; and the uppuRsIpE of
266 L. de Nicéville— List of the Butterflies of the Ké Isles. [No. 2,
both wings having the white discal band of twice the width.
Unpversipe, both wings have the white discal band of the same width as
on the upperside, in var. supous the band is wider than on the upperside,
though not as wide as in T. friopus. Differs from T. hermes, Grose
Smith, Rhop. Ex., pl. Oriental Lycenide iv, figs. 7, 8, female (1895),
from Korrido and Biak Islands, on the upPERSIDE of both wings in having
the discal band pure white instead of heavily irrorated with black
scales. UNDERSIDE, both wings have all the metallic green or blue
(according to the light) markings much more extensive. Differs from
7’. apollonius from several localities in New Guinea in my collection on
the UPPERSIDE of both wings in having the discal white band much
broader and pure white, instead of narrow, dusky and obscure. On
the UNDERSIDE of both wings the discal band is wider, twice as wide on
the hindwing. Matz, Indistinguishable from the same sex of
T. apollonius. Both sexes tailless.
The description above shews that this species is based on the
female sex only, It has been described from three males and one
female. It occurs rarely on Great Ké, and flies about the trees along
the banks of mountain streams; it is found also in high forest in Little
Ké Isle.
55. Taysonitis Korion, H. H. Druce and Bethune-Baker.
T. korion, H. H. Druce and Bethune Baker, Proc. Zool. Soc. Lond., 1893, p. 547,
pl. xlvi, fig. 3, male.
Originally described from the Kei Islands, where it occurs on
Little and Great Ké Isles, on the tops of bushes along the roads and
paths. The remaLE (hitherto undescribed) has on the uppERsIDE of the
forewing the costa widely black, extending into the middle of the
discoidal cell, the outer margin broadly and increasingly black, the rest
of the wing shining bluish-purple, with a white dash on the disc com-
mencing on the disco-cellularnervules. Hindwing with the costa widely
fuscous mixed with whitish, the outer margin more broadly black than
on the forewing, the middle of the wing shining blue. UnneErsipeE, both
wings exactly as in the male. This species has no tails.
56. THYSONITIS ILLUSTRIS, Rober.
Plebeius illustris, Rober, Iris, vol. i, p. 53, pl. iv, fig. 6, female (1885).
We have this species from Great Ké only, where it is very rare.
Messrs. H. H. Druce and G. T. Bethune-Baker in writing their mono-
graph of the genus in Proc. Zool. Soc. Lond., 1893, p. 552, note that
“The male only is described,” but Herr Rober says his type specimen
1898.} Li. de Nicéville—List of the Butterflies of the Ké Isles. 267
was a female. We have both sexes. The hindwing bears a thin black
tail tipped with white.
o7. *THysonitis caHiius, Felder.
Réber as Plebeius celius, Felder, from Key. It is probable that
Herr Rober so identified the species which has been described sub-
sequent to his 1891 paper as 7. korion, H. H. Druce and Bethune-Baker.
_ The latter gentlemen spell the word celius. As figured, this species has
no tails.
58. LycmNestHes emouus, Godart.
Not rare on Little Ké Island. Caught on flowers and bushes in
open places. It is highly probable we think that “ Plebeius”’ seltuttus,
Rober, Iris, yol. i, p. 67, pl. v, fig. 24, male; pl. iv, fig. 24, female (1886),
from the Aru Isles, and East and North-West New Guinea, is the same
species as L. emolus.
59. LYCHNESTHES TURNERI, Miskin.
L. turneri, de Nicéville, Journ. Bomb, \at. Hist. Soc., vol. xii, p. , 0. 13, pl. Z,
figs. 24, male ; 25, female (1898).
Much rarer than the preceding species on Little Ké and found with
it. It was originally described from Australia, and Australian speci-
mens in de Nicéville’s collection agree exactly with both sexes from the
Ké Isles.
60. EvERES ARGIADES, Pallas.
Somewhat rare on Little and Great Ké, flies near the ground,
amongst low plants.
. 61. Nacaposa HERMUS, Felder.
Rober and Ribbe as Plebeius unicolor, Rober, from Key, described
from Ceram, Key and East Celebes. Herr Rober has sent a male to
de Nicéville from Ceram, which proves that P. wnicolor is a synonym of
N. hermus. We have both sexes from the Ké Isles.
62. NAacaDUBA MEIRANGANUS, Rober.
Very rare on Little and Great Ké Islands. Originally described
from the Aru Isles. Has a very quick flight, and settles on the leaves
of bushes by the roadside. In coloration and markings the female of this
species very closely resembles that sex of Lyceenesthes turnert, Miskin.
J. 1, 34
268 L. de Nicéville — List of the Butterflies of the Ké Isles. [No. 2,
63. Nacapupa ANCYRA, Felder.
Found not commonly on all the islands on the flowers of Leguminose.
64.: NacaDUBA PERUSIA, Felder.
Little Ké Isle, rare, mostly found sitting on the tops of dry bushes.
N. laura, Doherty, is very close to N. perusia, if indeed actually separ-
able, except perhaps in the female.
65. NAcapDUBA ATRATA, Horsfield.
Rather rare on Little Ké Isle.
66. Nacapusa nora, Felder.
Little Ké Isle. The tailed form appears alone to occur, and is
perhaps better known as N. ardates, Moore.
67. JAMIDES PURPURATA, Grose Smith.
J. purpurata, Grose Smith, Nov. Zool., vol.i, p. 574, n. 219 (1894).
Ribbe as Plebejus astraptes, Felder. J. purpurata was originally
described from New Guinea. If we have correctly identified the Ké Island
species the male is distinguished by its bluish-purple colour on the upper-
side, which is not nearly as brilliantly metallic as it is in J. bochus, Cramer,
and its allies, and the outer black border to both wings is about 1°5 mm. in
width. It is near to J. cephion, H. H. Druce, Proc. Zool. Soc. Lond.,
1891, p. 367, pl. xxxi, fig. 19, male, from the Solomon Isles, but has the
black border to both wings on the upperside about twice as broad, and
it is not ‘brilliant morpho blue.” It differs from J. astraptes, Felder,
as figured by Semper from the Philippines, in having the black border
to the forewing on the upperside in the male less broad, and on the hind-
wing about twice as broad. In the Ké Isles it is rare, and is found on
Little Ké Island only.
68. Lampwes Aratus, Cramer.
Butler, Ribbe and Roéber as Lampides xtherialis, Butler, from Ké
Dulan (Butler), and Key (Ribbe and Rober). The commonest species
of the genus on all the islands of the Ké Archipelago.
69. LamprpEs cELENO, Cramer.
Butler as L. xlianus, Fabricius, from Ké Dulan, and Rober as
Plebewus selianus, from Key. Also very common on all the islands.
1898.]. Li. de Nicéville— List of the Butterflies of the Ké Isles. 269
70. LaAMPIpEs aMPuHISsA, Felder.
Rare on Little Ké Island. The DZ. areas, H. H. Druce, from the
Solomon Isles, Proc. Zool. Soc. Lond., 1891, p. 368, pl. xxxii, figs. 7,
male; 8, female, is very near to L. amphissa, which was originally des-
cribed from Batjan.
71. Lampipes nyias, Cramer.
Rober as Plebetus euchylas, Hiibner, Not rare on damp spots on
pathways in Little Ké, common on Great Ké.
72. CatTocHRYsops sTRABO, Fabricius.
Ribbe as Plebeyus kandarpa, Horsfield. Occurs commonly on legu-
minous plants on all the islands.
73. CATOCHRYSOPS LITHARGYRIA, Moore.
This species is found on Koer Island. The male is distinctly blue
on the upperside, while C. strabo, Fabricius, is as distinctly purple.
74. CATOCHRYSOPS CNEJUS, Fabricius.
Extremely common on Leguminosze. It occurs on Koer and the
other islands of the Archipelago,
75. Potyvommatus Beticts, Linnzus.
Ribbe as Plebeyus beticus from Key. Occurs rarely on Great Ké
Island.
76. AMBLYPODIA sp.
Herr Kiihn has sent to de Nicéville a somewhat rough coloured
drawing of a species of Amblypodia which the latter is unable to
identify. Herr Kihn caught a single specimen on Great Ké. The
drawing appears to represent a female; the upperside is dark umber-
brown, the forewing has a pale blue basal patch occupying about a
third of the wing, and extending from the subcostal nervure to the
inner margin. The hindwing has a similar basal patch, wedge-shaped,
narrow at the base of the wing, wide outwardly, and occupying the
whole of the discoidal cell. The underside of both wings is as usual
in the genus of various shades of brown, more or less mottled with
white. No species of true Amblypodia has, we believe, hitherto been
recorded as far east as the Papuan region.
270, L. de Nicéville— List of the Butterflies of the Ké Isles. [No. 2,
77. ARRHOPALA HELIUS, Cramer.
Rare on Little Ké, not seen at Great Ké. Found on young
Djamboe trees (Jambosa aqueea, Rumph.).
78. ARRHOPALA AMYTIS, Hewitson.
Roéber as Amblypodia micale, Blanchard, from Key. Common on
Little Ké Island on Djamboe and other fruit trees. Mr. Bethune-
Baker has kindly identified this species for us.
79. CURETIS sp.
Very rare, Kiihn has only one male specimen from Little Ké Island
in his collection, of which he has sent a coloured drawing to de Nicéville.
Without knowing its female, it is difficult to identify the species. It
agrees fairly well with OC. tagalica, Felder, from the Philippines.
80. HyYpoLYcHNA DANISOIDES, de Nicéville.
H. danisoides, de Nicéville, Journ. A. S. B., vol. Ixvi, pt. 2, p. 558, n. 13, pl. iii,
fig. 21, female (1897).
Very rare. Kiihn has bred it, the larva feeding on orchids. The
male differs only from the female in being smaller (1:1 inches in alar
expanse), it has both wings narrower (less rounded), and the apex of
the forewing more acute. The markings are precisely similar. It has
no secondary sexual characters whatever.
81. DsvupoRIx EPIJARBAS, Moore.
Found on Little Ké, and is very rare on flowering trees.
82. RaApaLA PHRANGA, Hewitson.
Originally described from Batchian. According to the figure the
Ke Isle form has rather more green-blue coloration on the upperside of
the forewing in the male than the typical form. The rema.e (hitherto
undescribed) has the forewing on the uppeRsIDE wholely greenish-blue
except the costa, apex and outer margin. Otherwise as in the male,
save, of course, that the male secondary sexual characters are wanting.
“ Deudoria”’ sumsont, Miskin, from Northern Australia, differs but slightly
from the Ké Isles form. It is found commonly on the leaves of trees
and bushes along the roads, especially on Little Ké Island.
83. BINDAHARA ISABELLA, Felder.
Very rare, only a few specimens from Little Ké Island obtained.
1898.] J.. de Nicéville— Lust of the Butterflies of the Ké Isles. 271
84. LipHyRA BRASSOLIS, Westwood.
A single male only obtained on Little Ké Island. Kiihn caught it
“one evening at the lamp. The Hon. Walter Rothschild in Nov. Zool.,
vol. v, p. 97, n. 5 (1898), has described Liphyra brassolis major from
females from Northern Australia. This may be the species named above.
Family PAPILIONIDA.
Subfamily Pirrina.
85. Leprosita xtpHia, Fabricius.
Common on all the islands, softly flying along close to the ground.
86. ELopINA EGNATIA, Godart. |
The specimens from the Ké Isles agree with Hewitson’s figure of
“ Pieris” padusa from Australia, except that the forewing is not nearly
so produced at the apex, the costa consequently being shorter; the
hindwing also is broader. Mr. W. H. Miskin in his Cat. Rhop. Aust.,
p- 8 (1891) gives P. padusa as a synonym of F. egnatia, but it is pro-
bably quite distinct. H. egnatia is not rare on Little Ké Island, but is
a butterfly very difficult to catch. The two sexes are almost exactly
alike. We have absolutely similar specimens from Wetter and
Northern Australia.
87. CavoPsILIA CROCALE, Cramer. Plate I, Figs. 5, larva; 5a, 5b,
pupa. |
Wallace as C. alemeone, Cramer. Rober as C. pomona, Fabricius,
C. crocale, Cramer, and ab. flava, Butler. “ Papilio” crocale, Cramer,
and “ Papilio” pomona, Fabricius, were described in the same year
(1775). We prefer to use the former name as it was accompanied by
a figure. There are several forms of this protean species occur-
ring on all the islands of the Ké Archipelago. Taking those without
ocelli on the underside, we have males with the yellow coloration
evenly suffused over both wings on the upperside, with females to
match them, the maJes agreeing with Butler’s figure of “ Callidryas”’
flava, from the Moluccas, &c., but the females are not nearly so heavily
marked with black on the upperside as in Butler’s figure of that
sex of C. flava. Taking those with ocelli on the underside, we have
males with the yellow coloration evenly suffused over both wings
on the upperside like C. flava, aud others with the yellow coloration
confined to a well-marked basal area on both wings, beyond which the
wing is white ; the females are like the other form. It is very common,
and Kuhn has frequently bred it on “ Djohur” trees, the larva being
272 L. de Nicéville— List of the Butterflies of the Ké Isles. [No. 2,
shagreened, above grayish-green, with a shining steel-blue line above
the spiracles, then a spiracular white stripe, which becomes yellow
from the second to the fourth segments, the infra-spiracular region is
light green, beneath it is bluish-green. Hach segment bears six
folds or creases, and all the segments are minutely dotted with black.
The pupa is pale green, with the head produced into a long pointed
process, the thorax humped in the dorsal line, there is a lateral yellow
line running from the extreme apex to the posterior end of the
pupa.
88, TRIAS HECABE, Linneus.
Rober as Hurema hecabe from Key. Butleras 7. photophila, Butler,
from Ké Dulan. He describes the male only, and says it has no sub-
apical brown patch on the underside of the forewing, this patch, however,
is present in females from Ké. In his latest revision of the genus
(Ann. and Mag. of Nat. Hist., seventh series, vol. 1, p. 75, n. 51 (1898),
he places J. photophila as a synonym of 7. sulphurata, Butler, which
“Ranges from Northern Australia northwards to Timor Laut, Aru, and
New Guinea, and thence eastwards to New Ireland, appearing just to
touch the Solomons; more to the south it ranges eastwards to the
Loyalty, New Hebrides, and Fiji Islands.’ Common on all the islands,
89. TERIAS sp.
Only one specimen from Little Ké. It is allied, from Kiihn’s
description, to T. leta, Boisduval.
90. Tertas CANDIDA, Cramer.
Not as common as JZ’. hecabe, Linneus, on Little and Great Ké
Islands. Dr. Butler gives Amboyna and Ceram only for this species.
Our male specimens agree with Cramer’s figure, and have the abdominak
margin of the hindwing on the upperside broadly black.
91. *Appras tyNncipa, Cramer.
Rober as Tachyris lyncida from Key. We have seen this species
only from Java, Bali and Lombok.
92. APPIAS CELESTINA, Boisduval.
Rare on Little Ké, somewhat more plentifully found on Great Ké
Island. Our females are Form I, bluish-white, not rich yellow on the
upperside.
1898.] L. de Nicéville— List of the Butterflies of the Ké Isles. 273
93. Appras cLAvis, Wallace.
Rober as Tachyris ada, Cramer, from Key. Originally described
from Ké Island. It is rather rare on Little Ké, not sorare on Great
Ké Island.
94. APpPIAS ALBINA, Boisduval.
Decidedly rare on Little and Great Ké Isles. The Form I of the
female, ground-colour white on both surfaces, appears to be the only
one found.
95, HUPHINA RACHEL, Boisduval.
Rober as Pieris pitys, Godart, from Key. ‘ Pieris” rachel was ori-
ginally described from Java, but we have seen no specimens from
thence. Our examples agree very well with the description, except
that the small yellow spot at the external angle (apex) of the hindwing
on the underside is often absent, and when present almost obsolete.
“ Pieris” pitys, Godart, is also a closely allied species, described from
Java, also never seen by us, which is said to have four or five white
spots arranged transversely in addition to the apical white spot on the
upperside of the forewing, whilst our species has one or at most two
subapical white spots only. P. pitys is figured by Lucas from Java, but
he does not shew the four or five white spots mentioned above, and in
other respects his figure does not agree with our specimens from the Ké
‘Isles. Perhaps ‘ Pieris” perictione, Felder, described from the Arru
Islands, is nearest to our species, but it has never been figured, and a
description alone is inadequate to enable one to discriminate between
very closely allied species, though the description agrees very well
with our specimens. H. rachel is very common throughout the year
on all the islands.
Subfamily PaprmLionina.
96. TROIDES PRIAMUS POSEIDON, Doubleday.
Rober as Ornithoptera priamus, Linneus. The Hon. Walter
Rothschild in “A Revision of the Papilios of the Eastern Hemisphere,
exclusive of Africa,” Nov. Zool., vol. ii, p. 191 (1895) records this
species as 7’. priamus poseidon, Doubleday, (& #): 9—ab. hecuba, Rober
(Tijd. voor Ent., vol. xxiv, p. 263 (9, nec &) (1891), from the Key
Islands. On Little Ké, Great Ké and Koer (Kiihn got none from
Tiandoe) this butterfly is always to be seen on the wing, but is never
abundant. The larva feeds on a species of Aristolochia.
274 L. de Nicéville — List of the Butterflies of the Ké Isles. [No. 2,
97. Paptnio poLypoRUS THESSALIA, Swinhoe. Plate I, Figs. 6,
larva; 6a, 6b, pupa.
Rober as P. polydorus, Linneus. The local race thessalia was
originally described from Ké Island (Halliburton). Itis usually com-
mon on Little and Great Ké Isles, but is sometimes rare, flying in
the open forest. ‘The larva is very similar to that of Troides priamus
poseidon, Doubleday, and feeds on the same plant, a species of Aristolo-
chia: it is dark violet in colour, with tentacular fleshy pale red processes
on the second, third, sixth, seventh, tenth and eleventh segments;
those on the fourth, fifth, eighth and ninth are brownish-red; the
process on the sixth segment is based on a pinkish-white spot; that on
the seventh segment has a pale base; there is also a series of supra-
spiracular processes, The pupa is pale brown, mottled with darker
brown, with a pair of red spots on the middle of the back above; it is
furnished with numerous foliaceous processes on the abdominal segments.
98. *PAPILIO FUSCUS ROTALITA, Swinhoe.
Rober as P. beccarii, Oberthtir.. This local race was described from
Ké Island (Halliburton). We have not seenit. Mr. Kihn notes that
he does not believe that this species ever came from the Ké Islands, as
from 1889 to the present date no collector has been on the islands except
himself; also that he (Kiihn) up to 1896 has sent butterflies from the
Ké Archipelago only to Dr. O. Staudinger, except one very small col-
lection to Herr J. Rober, so from whom could Col. Swinhoe have obtain-
ed it? In the original description Col. Swinhoe gives Halliburton as
the collector.
99. Papiio aALBINUS THOMSONII, Butler. Plate I, Fig. 7, larva.
Originally described from Ké Dulan. Mr. Rothschild in Nov. ©
Zool., vol. iii, p. 322, n. 3 (1896), has described an ab. mordingtoni from
Little Kei Island, from one female, captured by the late Capt. H. Cayley
Webster. Mr. Kihn writes to de Nicéville that had he gone on shore
on New Guinea the day Capt. Webster was murdered by the natives, he
(Kiihn) would have shared the same fate. It was quite an accident
that on that day he, for the first time durmng the expedition, remained
on board their vessel. We have not seen this aberration. P. thomsoniz
is a very variable butterfly: in some specimens there is a well-defined
oblique subapical white band on the upperside of the forewing which
often dwindles away to nothing; the large discal white patch on the
upperside of the hindwing varies greatly in size, in some specimens its
outer edge is even, and in the other extreme it is highly irregular,
being continued along the veins towards the margin; sometimes the
1898. | L. de Nicéville —- List of the Butterflies of the Ké Isles. 275
patch ends posteriorly on the first median nervule, sometimes it reaches
the submedian nervure; on the underside of the forewing the subapical
band is sometimes present, usually absent; on the hindwing there is
sometimes a discal series of large irrorated white spots from the costa
to the first median nervule, usually entirely absent; the blue and
orange submarginal markings are also more or less developed, some-
times both series are absent. It is very common on both Little and
Great Ké Isles. The larva is as variable as the butterfly, and feeds on
various trees, especially those of the Natural Order Aurantiacee. The
larva when young resembles a bird’s dropping. When full-grown it is
smooth with no processes; the head is pale green, the body is dark
green, becoming yellowish-green laterally, the second, third and fourth
segments beneath and the legs are brown, the fifth to thirteenth segments
beneath and the prolegs milky-white; the fifth segment bears pos-
teriorly a broad transverse black band, and the eighth segment bears
an oblique short black band on each side just above the spiracles; on
the fourth segment are two subdorsal milky-blue points on each side
one above the other, with a black spot just anterior to the spiracle; on
the ninth segment are two subdorsal obliquely-placed milky-blue points
on each side one above the other; and on the eleventh segment there is
one similar point on each side. Mr. Kiihn notes that the spots are some-
times greenish-brown.
100. #* PAPILIO HGHUS ORMENUS, Guérin.
Rober as P. ormenus, Guérin. Mr. Rothschild gives (b #): &—ab.
pandion, Wallace, also (f*):°9—ab. polydorinus, Haase, also (g *):
2—ab. amunga, Boisduval, from the Key Isles, but they are unknown
to us from the Ké Archipelago.
101. PapiLio mGEus KeIANuS, Rothschild. Plate 1, Fig. 8, larva.
Mr. Rothschild has described this local race from the Little Kei
Island (Kei Toeal) ; and also (a): 9—f, amaranta, and (b): ?—f.
blanca from the same island (Nov. Zool., vol. ii, p. 422, n. 4 (1896).
All our specimens appear to belong to this local race (not to P. ormenus,
Guérin), as in the males the discal white band ends on the first median
nervule, and does not reach the submedian nervure as it does in P. orme-
nus. In one of our male specimens the orange-red anal spot on the
hindwing on the upperside is absent, typically it is present, and Mr.
Rothschild has seen no specimen in which itis absent. It is rather rare
on Little and Great Ké, the female var. blanca especially so, and seen on
the wing only at the end of the wet season. The larva feeds on
orangeaceous trees. It is brownish-green, with lighter very fine lines
J. ut, 35
276 L. de Nicéville — List of the Bulterflies of the Ké Isles, [No. 2,
on each segment; the head and legs are black, with two short conical
yellow processes on the anterior edge of the second segment, the third,
fourth and fifth segments have each three pairs of similar processes,
the sixth, seventh, eighth, eleventh and twelfth segments have each two
pairs of similar processes ; on the fifth segment is a broad black band
ending on each side on the spiracular region; on the ninth segment
arising from the spiracular region and extending backwards over the
tenth segment is a broad oblique lateral band, which terminates in
the subdorsal region in a black conical process ; on the eleventh segment
is a much shorter posteriorly oblique broad black band, not reaching
the two black conical processes one on each side of the subdorsal
region; the thirteenth segment is white marked with large black.
patches ; posterior to the spiracles the body and the prolegs are white,
more or less interrupted with black lines.
102. *PapiLio DEIPHOBUS, Linneeus.
Herr J. Rober in Tijd. voor Ent., vol. xxxiv, p. 275 (1891), described
P. deiphobus, ab. hypoxanthos from the Key Islands. Neither Mr.
Rothschild nor we have seen this species from thence. Mr. Rothschild
spells the name “ hypoxanthus.”
103. PapiLio EUCHENOR OBSOLESCENS, Rothschild.
Rare in Little Ké, not so rare in Great Ké Island. Mr. Rothschild
suggests that P. ambrax epirus, Wallace, may occur in the Key islands,
but we have not obtained it there. It is found on the Aru islands.
104. PapiLio CODRUS TOEALENSIS, Rothschild. Plate I, Figs. 9, 9a,
larva ; 9b, 9c, pupa.
Described from Little Kei Island (Kei Toeal). It is not very rare
on Little Ké, but is difficult to catch. Kiihn has bred it, but does not
know the name of its food-plant. The larva is smooth, rapidly in-
creasing in width to the fourth segment, thence decreasing in width to
the anal segment, the head, body and legs are yellowish-green ; on the
dorsal area of the fifth segment are eight short indigo-blue marks which
form two diamond-shaped figures; on the twelfth and thirteenth seg-
ments are five similar marks, one in the middle with four around the
central mark forming an oblong figure; the second, third and fourth
segments each bears at the side a short, bluntly-conical, pale red, fleshy
process; the thirteenth segment with a pair of diverging similar pro-
cesses ; the spiracles are indigo-blue. The pupa is violaceous light
greenish-gray, sometimes yellowish-green ; the anterior portion is broad,
and produced into a rather high process dorsally ; from the apex of this
1898.] L. de Nicéville— List of the Butterflies of the Ké Isles. 277
process (which is dark brown at the tip) descends on each side to the
spiracle at about the middle of the pupa a fine indigo-blue line, with a
large round blue spot in its middle; the spiracles are dark brown; the
pupa ends in a somewhat sharp point.
105, Papruio EURYPYLUS MELAMPUS, Rothschild.
Described from Little Kei Island (Kei Toeal). Mr. Rothschild
(Nov. Zool., vol. iii, p. 425 (1896), has also described an ab. rufinus from
the same island. P. melampus is very rare, and is found on both Little
and Great Ké Islands.
106. PaPInio SARPEDON CHOREDON, Felder.
Rare on Little and Great Ké Isles. Very quick on the wing.
107. PAPpiILio AGAMEMNON ARGYNNUS, Druce.
Recorded by Wallace from the Ké Island as P. agamemmnon, local
form b, and described from the Ké Island by Druce as P. argynnus. It
is rather common on Little and Great Ké Islands. The larva feeds on
Anona muricata, Linnezeus, Malay name “ Surakajah.”
Family HESPERIID &.
108. Taciapes JAPETUS, Cramer. Plate I, Figs. 10, larva; 10a, 108,
pupa.
Rober, Ribbe and Butler. Very common on all the islands. The
larva is greenish-white, the skin transparent, very finely striped
with white, yellowish between the segmental folds; between the
eighth and ninth segments are two yellowish coloured organs visible
through the skin; the head is dark brown, heart-shaped, strongly
indented above in the dorsal line. Pupa attached openly to a leaf
by a few threads and by the cremastral hooks; pale yellowish,
streaked throughout with reddish-brown; each wing-cover bears two
large irrecularly-quadrangular china-white spots, one at the base of the
wing, which is the smaller, tle other at the anal angle of the expanded
wing, about twice as large ; between the eyes in front are two white
spots like a pair of spectacles; on the third, fourth and fifth abdominal
segments is a triangular white spot on each side; the head is produced
into a long thin pointed process. The larva lives on sweet potatoes.
109. * AmpirriA MARO, Fabricius.
Ribbe as Pamphila maro. This is probably a wrong identification.
278 L. de Nicéville — List of the Butterflies of the Ké Isles. [No. 2,
110. *Norocrypra WAIGENSIS, Plotz.
Rober as Plesioneura waigensis. We have been unable to recognise
this species from the description only. It was originally described from
Waigou.
111. Norocrypta FEISTHAMELII, Boisduval.
Not common on Great Ké, rare on Little Ké. Frequents wet places.
Specimens from the Ké Isles and Northern Australia are peculiar in
having the discal diaphanous band on the upperside of the forewing
produced almost to the costa anterior to the costal nervure, instead of
ending anteriorly on the subcostal nervure as usual.
112. *Texicota aucias, Linneeus.
Rober as Pamphila augias. We have not seen it from the Ké
Archipelago, but its occurrence there is not improbable.
113. Txricora BAMBUSA, Moore.
Apparently not rare. Kihn has sent de Nicéville seven male
specimens.
114. *TeLicota mMoseLeyi, Butler.
Pamphila moseleyi, Butler, Ann. and Mag. of Nat. Hist., fifth Series, vol. xiii,
p. 198, n. 50 (1884).
Ké Dulan (Butler). Capt. E. Y. Watson in Proc. Zool. Soc. Lond.,
1893, p. 103, places this species in the genus Telicota. It may be the
same species as the next, with which it agrees insize. It was described
from a male, while our Padraona augiades, Felder, is represented by a
single female; the description of P. moseleyi does not agree with our
specimen, but the differences may be due to sex.
115. Papraona avciapes, Felder.
This species was described from a male from Amboina, which was
subsequently figured in the “Reise Novara.” Of the six distinct
orange and black Skippers we possess from the Ké Islands this is by
far the largest. It appears to be rare, we have a single boos onl
which is very similar to examples of the same sex of P. RE a,
Moore, from India, but is much larger, and is more tawny, less neues
coloured. :
116. Papraona procieEs, de Nicéville.
P. procles, de Nicéville, Journ. Bomb Nat. Hist
’ ’ . , LU. ist. S ; i ee
pl, J, figs. 7, male; 8, female (1892), oc., vol. vii, p. 358, n, 2],
1898.] LL. de Nicéville — List of the Butterflies of the Ké Isles. 279
Originally described from the Ké Isles, where it appears to be a
common species on all the islands.
117. *Papraona suntas, Felder,
Dr. Butler records this species with a query from Ké Dulan. It
was originally described from Amboina, but has not been figured. We
have not been able to recognise it from the description only. Messrs.
Hlwes and Edwards place it as a synonym with a query of Telicota
[Padraona| dara, Kollar (Trans. Zool. Soc. Lond., vol. xiv, p. 254
(1897).
118. *Papraona masa, Moore.
Rober as Pamphila mesa. We have no species of the genus from
the Ké Archipelago which agrees with P. mesa.
119. Papraona, sp. Ll.
Hasitat: Little Ké Isle.
Expanse: 6, 1:'4; 92, 15 inches.
Description: Mare. Upprrsipe, both wings black, with a slight
purplish gloss; all the markings yellow; cilia yellow. Forewing with
a short costal streak from the base of the wing ; a similar streak in the
discoidal cell; the space between these two streaks slightly irrorated
with yellow scales ; three conjoined subapical elongated dots, of equal
length, the middle one placed slightly nearer the base of the wing than
the others ; four obliquely-placed discal spots, the anteriormost in the
lower discoidal interspace small and quadrate, the second spot occupying
the base of the second median interspace, its outer end excavated, the
third spot larger than the second, occupying the middle of the first
median interspace, its outer end excavated, the fourth spot crossing the
middle of the submedian interspace, quadrate, its outer end excavated,
its inner side not in a line with the spot anterior to it, being placed
nearer the outer margin; from the posterior inner angle of this last-
named spot runs a line of yellow scales along the submedian nervure to
the base of the wing ; a streak in the sutural area, its outer end in a line
with the outer edge of the spot anterior to it. Hindwing with a rather
small indistinct clump of sete at the base; a discal transverse band,
with even edges, and of equal breadth throughout, occupying the middle
of the wing; some yellow sete in the submedian interspace. Unpsr-
siDE, both wings with all the veins narrowly black; a narrow black
anteciliary line, ending on the hindwing at the first median nervule.
Forewing black, but the costa and apex to just posterior to the first
280 L. de Nicéville— List of the Butterflies of the Ké Isles. [No. 2,
median nervule, where it fines away to nothing, dull brownish-yellow ;
the markings much as on the upperside, but those on the dise broader.
Hindwing dull brownish-yellow ; the discal band much narrower than:
on the upperside. AbpomMEN narrowly ringed with yellow. Femate.
Differs from the male only in being a little larger. .
This description has been drawn up by de Nicéville from three
males and three females. It is a very distinct species, but he has not
ventured to name it, asit has probably been already described from New
Guinea or Australia, from whence he possesses but few species of the
genus. It is very near to P. autoleon, Miskin, from Northern Australia.
120. Papraona, sp. 2.
Hasitat: Keé Isles.
Expanse: @,9, 1°0 inch.
Description: Mate. Uprsrsipe, both wings black, with golden-yellow
markings ; cilia yellow, marked with black at the ends of the veins,
broadly on the forewing, narrowly on the hindwing. Forewing more
yellow than black, there being a large basal triangular yellow patch
from the costa to the submedian nervure and first median nervule, a
short yellow streak in the submedian interspace, and a longer one in the
sutural area ; a discal yellow band extends nearly across the wing from
close to the costa to the submedian nervure, its anterior portion shifted
inwardly and out of line with the posterior portion, consisting of three
conjoined increasing spots, the rest of the band consists of five spots,
their inner edge straight, their outer edge toothed. Hindwing with a
small yellow spot in the middle of the discoidal cell, with some yellow
setee anterior to it; a broad curved discal band, commencing just
posterior to the costa by an oval spot, posterior to which is a small
triangular spot placed nearer the margin than the spot anterior to if,
and joined to the discal portion of the band, which latter crosses the
middle of the wing, and is of equal width throughout, with slightly
irregular edges. UNDbERSIDE, forewing. marked somewhat similarly to
the upperside, but the base of the wing is black; there is a small patch
of yellow scales posterior to the three anterior spots of the discal band ;
also a submarginal yellow band beyond the discal band from the costa
to the first median nervule. Hindwing yellow ; the discal band defined
on both sides with a narrow black line; a prominent anteciliary black
thread. ApBpomen black, narrowly ringed with yellow. Fsmaun, very
similar to the male, but all the yellow markings on the upperside
reduced in size, leaving more of the black ground-colour visible.
This description has been drawn up by de Nicéville from twelve
Sl e—————— SSL
er SS Slr,
1898.] _ L. de Nicéville — List of the Butterflies of the Ké Isles. 281
male and two female examples, which were received from Kihn as
Hesperia flavovittata, Latreille, described from New Holland, with which
description as far as it goes the specimens agree fairly well, but
Mr. W. H. Miskin in his Syn. Cat. of the Lep. Rhop. of Australia places
H. flavovittata in the genus TYaractrocera, distinguished by its short
round-clubbed antennee, while the species described above has the longer
pointed-clubbed antenne of the genus Padraona. It appears to be quite
a distinct species, but has probably already been named, indeed, in
de Nicéville’s collection from Mackay in North Australia are exactly
similar specimens, which agree with Hewitson’s description of
“ Ancylowipha” agraulia, from the same region. The correct spelling
of the genus is Ancyloxypha. Miskin places the latter species in the
genus Apaustus, Hiibner, and gives Pamphila sunias, Felder (see no. 117
above) as a synonym, though it has eight years’ priority.
121. PapRAona, sp. 3.
Kasirat: Ké Isles.
Hxpanse: &,°8; 9, °9 of an inch.
Description: Mate. Uppersipe, both wings black, with yellow
markings; cilia of the forewing black anteriorly, becoming yellow
posteriorly, of the hindwing yellow, the terminations of the median
nervules streaked with black. Forewing with a triangular yellow
streak from the base of the wing to beyond the middle, not quite reach-
ing the costa anteriorly, bounded by the submedian nervure and second
median nervule posteriorly, its outer edge inwardly oblique; three con-
joined subapical spots, the anteriormost a mere dot, the second
_ twice as large as the one before it, the third twice as large as the
second ; a discal band consisting of five portions, separated from the
subapical spots, the two anteriormost portions very small, the third
elongated, occupying the base of the second median interspace, the
“fourth of the same length as the third, occupying the middle of the first’
median interspace, the fifth very narrow and linear, the outer edge of
this band is nearly even, the inner edge very irregular, owing to the
third and fourth spots being so much broader than the others; a
yellow streak occupying the basal half of the sutural area. Hindwing
with a small yellow spot in the middle of the discoidal cell; a broad
discal band occupying the middle of the wing, both edges irregular.
Unversips, forewing marked as above, but the apex is broadly yellow.
Hindwing yellow throughout, except a broad black streak in the sub-
median interspace; the spot in the cell and the discal band defined
outwardly by a narrow black line; an anteciliary black thread.
282 L. de Nicéville — List of the Butterflies of the Ké Isles. | No. 2,
AspomEN yellow, narrowly banded with black. Frmate differs from
the male only in having the wings broader, and the apex of the forewing
less produced.
Described by de Nicéville from six male and one female specimens.
It is an easily recognised species, is the smallest of the yellow and
black species occurring in the Ké Archipelago, and is not named here as.
it is probably known already from neighbouring islands. It is near to:
P. mexsoides, Butler, described from Malacca, and figured by Moore from
Ceylon, but is much smaller, and differs a good deal in the details of
the markings.
122. Baorts (Chapra) matutas, Fabricius.
The form of this species occurring in the Ké Isles has in the male
only two most minute dots in the median interspaces in the forewing,
on the underside of the hindwing the discal spots are also very small ;
the female is normal. (. agna, Moore, has the spots smaller than
C. mathias, the present form has them smaller still. It is a common
species on Little Ké Island.
123. Baorrs (Parnara) paiiippina, Herrich-Schiffer,
This species has no spots in the discoidal cell of the forewing,
there is a discal series normally of seven spots, but several of the
anterior ones are often wanting, in an extreme form there are only two
spots in all placed in the median interspaces. It is a common species
on Little Ké Island.
124. * Baoris (Parnara) LARIKA, Pagenstecher.
Rober records this species from Key as Pamphila larika. It was
described from Amboina (Jahr. des Nass. Ver. fiir Natur., vol. xxxvii,
p. 207, pl. vii, fig. 1, female (1884). It is evidently closely allied to the
preceding species, but has all the spots on the upperside of the forewing
larger. Doubtless Herr Rober did not know Parnara philippina, Herrich-
Schaffer, or he would have so named his specimens from Ké. It is very
doubtful if two such closely allied species as P. philippina and P. larika
are found in the Ké Archipelago, even if they be really distinct.
125. Hasora DOLESCHALLU, Felder.
Réber as Ismene doleschalli [sic!]. This beautiful species was
originally described from the Moluccas. It is rather rare on Little and
Great Ké Islands. The opposite sexes do not differ in markings and
coloration. Kiihn has bred it, but has not sent de Nicéville a descrip-
tion of its transformations.
1898.] UL. de Nicéville— List of the Butterflies of the Ké Isles. 283-
126. Hasora piscontor, Felder.
This species has been identified by de Nicéville from a coloured
drawing of an unique specimen in Kiihn’s collection, taken on Little Ké
Island, and which appears to be a female. The species was originally
described doubtfully from Java. Itis one of the handsomest “ Ismenes”
known. In de Nicéville’s collection are both sexes from Northern
Australia.
127. Hasora (Parata) matayana, Felder.
Ribbe as Ismene malayana. Identified by de Nicéville from three
females which agree exactly with Felder’s figure of the species, and with
some female specimens in de Nicéville’s collection from the Andaman
Iles. The Ké examples have no transparent spots whatever on the fore-
wing. Messrs. Elwes and Edwards in their recent monograph, p. 301,
place this species as a synonym of H. chromus, Cramer, which latter
according to Dr. Aurivillius is a synonym of H. alexis, Fabricius (Ent.
Tids., vol. xviii, p. 150, n. 68 (1897).
128,. BaDAMIA EXCLAMATIONIS, Fabricius.
Not uncommon on Little and Great Ké Islands. It is found from
India to Australia.
EXPLANATION OF Prater I.
a
ge
, larva, Huplea (Hirdapa) assimilata, Felder, p. 256.
, larva; 2a, head of larva; 2b, 2c, pupa, Cethosia cydalima,
Felder, p. 258.
, young larva on food-plant; 3a, full-grown larva, Neptis
(Rahinda) consimilis, Boisduval, p. 260.
» 4a, 4b, pupa, Charaxes keianus, Rothschild, p. 262.
, larva; 5a, 5b, pupa, Oatopsilia crocale, Cramer, p. 271.
, larva; 6a, 6b, pupa, Papilio polydorus thessalia, Swinhoe,
p. 274.
, larva, Papilio albinus thomsonii, Butler, p. 274.
, larva, Papilio wegeus keianus, Rothschild, p. 275.
, 9a, larva; 9b, 9c, pupa, Papilio codrus toealensis, Roths-
child, p. 276.
» 10, larva; 10a, 10b, pupa, Tagiades japetus, Cramer, p. 277.
Co bo
o> or
os 8
C OO «7
wo. 1h OG
284 G. King and D. Prain— Descriptions of some New Plants [No. 2,
Descriptions of some new plants from the North-Eastern Frontiers of India.
—By G, Kine and D. Prati,
[ Received January 28th; Read March 2nd, 1898. ]
While dealing with various collections received from the North-
Eastern Frontiers of the Empire in connection with the Botanical
Survey of India, the writers have had to dispose of a number of species
that appear to be new to science and that are unprovided with names
in the Herbarium of the Royal Botauic Garden, Calcutta; the present
paper contains descriptions of a few of the more notable of these.
A considerable proportion of them it was necessary to have compared
at Kew in order to make certain that they were unknown or unrepre-
sented in the unrivalled collection there; our thanks are due to
Mr. Thiselton-Dyer, the Director, and to Dr. Stapf, the Assistant for
India in the Herbarium there, for kind assistance in connection the
examination of these.
ANONACE.
1. GonioTHALAMUS pEDUNCULARIS King & Prain; frutex P, ramulis
gracilibus glabris. Folia tenuiter coriacea, oblonga, plus minusve oblan-
ceolata, breviter acuminata, basi cuneata; utrinque glabra, hebetia,
subtus ex sicco pallide brunnea; nervis secundariis 10-12-jugis curvis
intra marginem inosculantibus subtus plus minus prominentibus supra
obsoletis ; petiolis brevibus ‘25-3 poll. longis, laminis 6°5-9 poll. longis,
his 1:5-2°5 poll, latis. Flores solitarii erecti parum supra-axillares
1:5 poll. longi, pedicellis plus quam uncialibus adpresse puberulis, basin
versus minute bracteolatis. Sepala carnosa, libera, ovata subacuta
patentia utrinque puberula, ‘3 poll. longa. Petala carnosa, seriei
exterioris oblique ovato-lanceolata, acuminata, basi augustata et in-
crassata ubi intus excavata, extus adpresse pubescentia intus basi
puberula ceterum glabra 1°5 poll. long, °75 poll. lata; petala seriei
interioris ovata, acuta basi angustata utrinque sed praesertim extus
pubescentia, dimidio summo in calyptram basi 3-fenestratam cohaerentia.
Antherae oo, sessiles lineares apice capitate. Pzstillia circa 20, germina
linearia, stylis linearibus pubescentilus duplo breviora. Fructus nondum
communicatus.
In Burma svUPpERIORE: in montibus Kachin nuncupatis, Kingiz
mercenar. !
Of all the Indian species of this genus G. peduncularis most
resembles the Ceylonese G. Gardneri H. f. & T. and G. Thwattesii
Ae ear aN a
1898. | from the North-Hastern Frontiers of India. 285
STERCULIACEA.
2. STERCULIA cocnaTA Prain; arbuscula ramulis gracilibus parce
puberulis cortice brunneo obtectis. Folia glabra brevissime petiolata vel
sessilia auguste lanceolata medio versus basin sensin attenuata apice
anguste ovato-acuminata, margine integra, chartacea, subtus prominen-
tius 25-30-nervia simulatque reticulato-venosa. lores albi pedicellati
pedicellis filiformibus glabris, in racemis quam folia dimidio brevioribus
dipositi; calyce campanulato laevi intus glabro extus parcissime
puberulo, lobis linearibus erecto-patentibus tubo multo longioribus.
Follicula oblonga acuta breviter pedicellata extus velutina intus glabra
utrinque rubra ; semina nigra nitida subsphaerica.
In montibus Kachin nuncupatis; Kingii mercenar. !
Folia 8-12 poll. longa, *75-2°5 poll. lata, petiolis nunquam ‘2 poll.
saepius omnino absentibus; racemis 4 poll. longis, pedicellis capillaribus
*3 poll. longis; floribus ‘6 poll. longis ; folliculis 2°5 poll. longis, *75 poll.
latis ; seminibus *5 poll. diam.
Nearest §. Roxburghti, S. parvifolia, and S. striatiflora but easily
distinguished by the sessile leaves and the other characters mentioned.
CONNARACEAE.
3. TAENIOCHLAENA BIRMANICA Prain; frutex ramulis puberulis
teretibus cortice minute lenticellatis. Folia imparipinnata, foliolis 2-3-
jugis, coriaceis, nervo mediano supra puberulo excepto glabris, oblongo-
lanceolatis apice emarginato-caudatis basi inaequaliter cuneatis breviter
petiolulatis, rachide puberulo. lores in racemis brevibus vel paniculis
congestis axillaribus dispositi, bracteis minutis, pedicellis longiusculis.
Calyx basi hemisphaericus, laciniis valvatis oblongis acutis fructu
revolutis. Petala ... Stamina 10, alterna paulo breviora, filamentis basi
vix connatis. Oarpella 5, sessilia, styli parum elongati, puberuli.
Capsulae 1-3, sessiles, ovoideae, parum apiculatae, extus intusque
glaberrimae. Semen oblongum basi arillo adnato dimidiato suffultum,
testa nitida ; cotyledones amygdalinae,
Burma: in montibus Kachin nuncupatis, Kingit mercenar. !
Folia 6-8 poll. longa, rachide 3-4 poll., lamina terminali 3-5 poll.
longa 1:5 poll. lata, lateralibus 1:5-3 poll. longis, petiolulis -15 poll.
Racemis 1°5-2 poll. longis, pedicellis *3-"4 poll. longis. Capsulis *6 poll.
longis, °3 poll. latis. .
Much resembles the only other known species, 7’. Griffithii Hook. fil.,
from Malacca, but with differently shaped leaflets and with fruits that
are glabrous externally instead of pubescent.
986 G, King and D. Prain— Descriptions of some New Plants [No. 2,
LEGUMINOSAE.
4. INDIGOFERA NIGRESCENS Kurz MSS.in Herb. Calcutta; fruticosa
ramulis adpresse brunneo-setosis, foliolis minoribus 17-21, oppositis
membranaceis, atro-viridibus, utrinque sparse adpresse puberulis. Flores
in racemis angustis, elongatis, breve pedunculatis dispositi, bracteis
linearibus alabastris longioribus, pedicellis brevissimis, calyce oblique
campanulato, dentibus lanceolatis tubum excedentibus. Corolla rosea,
Legumen lineare turgidum, adpresse puberulum, rectum, minute apicu-
latum, suturis parum incrassatis ; semina 6-8.
In montibus Kuasta; apud Shillong, G. Mann! Clarke n. 5848!
In montibus Kacurn, piope Myitkyina, Kingii mercenar.! In valle
Taping, YunNnaN austro-occidentalis, apud Momien, J. Anderson! In
montibus SHan, ad Maymyo, Kingit mercenar. !
Folia 3-4 poll., foliola ‘5-8 poll. longa, ‘3-4 poll. lata. Racemz
4-8 poll. longi, pedunculis 1-pollicaribus, bracteis ‘2 poll., calyce °05
poll. longo, corollis ‘2 poll. longis tantum. Legumen ‘75-85 poll.
longum ; °1 poll. crassum.
Very closely related to Indigofera atropurpurea with which it has
been often identified and under which name it is usually met with in
collections. The smaller flowers with longer calyx-teeth, shorter faintly
apiculate and puberulous instead of glabrescent pods, as well as the
very dark green leaves with more numerous and much smaller leaflets
amply distinguish it.
5. SpaTHoLosus Porrincert Prain; frutex scandens robusta,
ramis cylindraceis gracilioribus cortice brunneis, pilisque reflexo-ad-
pressis sparse pubescentibus. Jolia pinnatim 3-foliolata rachide pilis
patentibus pallide fusco-hirsuta, foliolis subcoriaceis ambitu ovatis
margine sinuatis supra uniforme subtus praesertim nervis pilis adpressis
sparse hirsutis, nervo mediano subtus petiolulisque brevibus pilis
patentibus strigoso, foliola terminali aeque basi late cuneata apice late
truncata, nervis prominentibus lateralibus 9-jugis mediano in acumine
subulato producto; foliolis lateralibus paullo minoribus basi parum
inaequali intus cuneatis extus ovato-rotundatis nervis lateralibus 7-8-
jugis, mediano ultra apicem abrupte acutam producto, stipulis deciduis
lanceolatis stipellisque persistentibus subulatis adpresse hirsutis. Flores
racemosi in paniculis terminalibus iterum terve ramosis ramis angulatis
parcius pubescentibus dispositi, bracteis brateolisque caducis minutis,
pedicellis capillaribus calyce brevioribus. Calya dense olivaceo-veluti-
nus, dente summo apice emarginato ceteris anguste deltoideis omnibus
tubo parum brevioribus. Corolla purpurea, vexillo ungue anguste
cuneato apice emarginato, petalis ceteris unguibus gracilibus calycem
subacquantibus, alarum laminis angulo inferiore utrinque barbellatis.
EE Ee
1898. | from the North-Eastern Frontiers of India. 287
Stamina 2-adelphia filamentis vaginae carinalis alternis longioribus,
filamento libero vexillari quam vaginam multo breviore. Ovariuwm
puberulum breviter stipitatum, ovulis 2. Legumen ignotum.
In montibus KacHin nuncupatis, apud Namli, 2000 p. s. m.,
Pottinger !
Ramulis floriferis ‘25 poll. diam. ; folvis 8 poll. longis, rachide 2:5
poll. parte terminali partem petiolarem fere aequante, petiolulis ‘15 poll.
stipellis ‘2 poll. brevioribus, stipulis *25-°3 poll. longis; lamina ter-
minali 5°5 poll. longa 4 poll. lata, lateralibus 4°5 poll. longis his 3 poll.
latis, acuminibus apicalibus °3 poll. longis. Panicula tota pyramidali
1:5—2-pedalis, paniculis secundariis 8-10 poll. longis, iisque ordine tertio
4-6 poll. longis, racemis singulis 1*5-2-pollicaribus, 12-20-floris, pedi-
cellis ‘12 poll., calyce ‘2 poll., corolla ‘3 poll. longis.
A very fine plant nearest to the Malayan species S. gyrocarpus and
S. ferrugineus but abundantly distinct from these and from all the other
Indian species by the sinuate finely apiculate leaflets.
CRUDDASIA Prain.
Calycis lobi acuti, 2 superiores in unum apice minute 2-dentatum
connati, caetera aequilata triangulares infimo tamen lateralibus parum
longiore, Vexillum suborbiculatum, basi nec inflexum; alae oblongo-
ovatae basi carinae adhaerentes ; carina cymbiformis erostris alis aequi-
longa. Stamer vexillare caeteris arcte connatum, antherae uniformes.
Ovarium sessile o-ovulatum; stylus filiformis incurvus sub stigmate
terminali summo apice parce penicillatus ceterum glaber. Legumen.
elongatum, 2-valve, planum, coriaceum, compressum intus inter semina
tenuiter farctum. Semina plano-compressa, suborbicularia, hilo ovato,
estrophiolata. Caulis alte volubilis, foliis pinnatim 5-foliolatis, foliolis
anguste ovatis, stipellatis. Stipulae spinuloso-setaceae caducae, basi-
fixae, Flores purpurascentes, in pedunculis elongatis fasciculato-race-
mosi, fasciculorum rhachide nodiformi. Bracteae bracteolisque caducae.
Calyx vexilloque extus sericeus.
Species singula, montium Kachin incola. Genus _ subtribubus
Galacticarum, praesertim sectioni Collzae generis Galactiae cui etiam
stamen vexillare cum ceteris medio connatum, vel Diocliearum praeser-
tim generi Puerariw fere aeque recte attribuendum; statim tamen ab
ambabus stylo more Huphaseolarum nonnullarum circa stigma barbato
differt, ideoque potius pro genere distincto Dioclieis uti Clitoria Glycinets
relato habenda.
6. Cruppasia InsIGNis Prain.
In montibus Kacuin nuncupatis, 5000 p. s. m., Kingit mercenar. !
Peticli 2-4 poll. rachides communi 1°5 poll. supra canaliculati,
2988 G. King and D. Prain— Descriptions of some New Plants [No. 2,
retro-setosi, laminae ovato-lanceolatae 3-6 poll. longae, 1°5-2°5 poll.
latae, supra glabrae, subtus adpresse pubescentes, inter nervis 12-15-
jugis subtus prominentibus reticulato-venosae; stipellae filiformes *1
poll. longae, petiolulae 1°5 poll. longae; stipulae rigidae *2 poll. longae.
Racemt 8-12 poll, longi pedunculis 3 poll. longis, nodis vix ‘2 poll.
remotis, pedicellis ‘12 poll. longis. Calyx ‘2 poll. tubo campanulato
dentibus aequilongo. Corolla *4 poll. longa, vexillo orbiculato °35 poll.
lato, extus dense sericeo. Legumen 3 poll. longum, ‘3 poll. latum,
seminibus 10-12, :25 poll. longis, ‘2 poll. latis.
7. PueRARIA BELLA Prain ; volubilis ramis gracilibus glabris, foliis
3-foliolatis stipulis caducis stipellis capillaribus, foliolis chartaceis
ovatis longe acuminatis basi cuneatis utrinque petioloque glabris,
petiolulis parce puberulis. Flores in racemis elongatis simplicibus
vel parce ramosis dispositi, rachide parce puberulo, pedicellis brevis-
simis, bracteis deciduis bracteolis 2 sub calycem persistentibus cordato-
ovatis parce puberulis. Calyx campanulatus glabrescens, segmentis
2 summis in labium apice emarginato truncatum connatis, caeteris ovatis
obtusis subaequilongis omnibus tubo parum brevioribus. Corolla pur-
purea calyce plus duplo longiore, vexillo basi auriculis inflexis appendi-
culato, carina rectiuscula alas subaequante. Stamen vexillare omnino
solutum, antherae uniformes. Ovariwm subsessile oo-ovulatum stylus
filiformis superne inflexus,. imberbis; stigma capitatum. Legumen
ignotum.
In montibus Kacutn nuncupatis, prope Myitkyina, Kingii mer-
cenar. !
Foliola 6 poll. longa, 3 poll. lata, stipellis filiformibus ‘25 poll.
longis, petiolulos aequantibus. Bracteolae ‘1 poll. longae. Calyx
25 poll. longus. Corolla ‘7 poll. longa.
This very distinct species belongs to the subgenus Neustanthus, which
is marked by having entire leaves that are contemporaneous with the
flowers. The general appearance of the plant most readily recalls that.
of Pueraria Thunbergiana but its stipules are not persistent as in that
species and the calyx is widely different. The fact that the vexillary
stamen is quite free marks it as an aberrant Pueraria to be placed
near P. peduncularis which exhibits the same character. The flowers—
though not the bracts—recall those of Mastersia assamict and till ripe
fruits are reported it can not be quite certain that it should not be
referred to that genus. Meanwhile it is most satisfactorily located in
Pueraria of which it has all the facies.
8. Derris LattroLia Prain; arbor alta, foliis magnuis, foliolis 5-7
chartaceis ovato-lanceolatis apice acuminatis basi cuneatis vel subrotun-
datis. Flores in paniculis amplis thyrsoideis axillaribus dispositi
1898. | from the North-Hastern Frontiers of India. 289
rachide ramisque glabris angulatis nodis nec tumidis nec productis,
pedicellis distinctis approximatis; calyce glabrescente campanulato
margine truncato; corollae vexillo erecto orbiculari, basi ecalloso
rotundato; filamenta 2-adelphia glabra; ovario puberulo, ovulis 3.
Legumen ligulatum tenue glabrum suturis utrinque alatis nec sinuatis.
In montibus Kacutn, apud Namli, 4000 p. s. m., Kingii mercenar. !
Folia 18-24 poll. longa, foliolis 8-10 poll. longis 3:5-4 poll. latis
utrinque glabris petiolulis ‘25 poll. longis. Paniculae 20-25 poll. longae,
ramulis 2—4 poll. longis ; pedicellis ‘1 poll. longis ; calyce 12 poll. longo
"15 poll. Jato; corolla °3 poll. longa, vexillo °85 poll. lato; legumine
3°5 poll. longo, 1 poll. lato; alis 15-2 poll. latis subaequilatis.
This species is very nearly related to D. thyrsiflora which it much
resembles in flowers and pods except in having them distinctly pedicel-
led ; in this latter respect 16 more nearly approaches D. Wallichit of
which we were at first inclined to treat it as a large flowered variety.
The much larger leaflets however and the fact that this is a tall tree
makes it preferable to treat it as a distinct species.
9. DavperciA Kinerana Prain; frutex scandens lignosa, cortice
lenticellato, foliolis 5-7 anguste ovatis apice breviter acuminatis, basi
cuneatis supra glabris subtus sparse puberulis, coriaceis, Flores pani-
culati, paniculis axillaribus, foliis brevioribus, ramulis subcorymbosim
dispositis, pedicellis brevissimis; calyce dense ferrugineo, dentibus 3
inferioribus anguste triangulis, summos breviores latioresque excedenti-
bus ; petalorum unguibus calyce aequilongis; staminibus 9 monadel-
phis ; ovario glabro 2-ovulato; stylo subulato. Legumen ignotum.
In montibus Kacuin nuncupatis, Kingit mercenar. !
Foliola 2°5-8 poll. longa, 1-1'25 poll. lata; rachide 3 poll. longo
petiolulis *2 poll. longis; paniculae 3°5 poll. longae, ramulis 1-15 poll.
longis, floribus °25 poll. longis.
Very near to Dalbergia Benthami Prain, (D. rubiginosa Benth. Flor.
Hong-Kong 93, not of Roxb.) from Hong-Kong, but with quite different
leaflets and with rather longer panicles of similar flowers. Also near
D. rubiginosa Roxb., from Western India, but again with different
leaflets and rather larger flowers. In general appearance D. Benthami
resembles D. rubiginosa, to which Mr. Bentham has referred it, but the
leaflets of the Chinese plant are thicker, narrower towards the tips, and
have a different pubescence beneath; the flowers too of the Chinese
plant are very like those of the present Kachin species and are con-
siderably larger than there of D. rubiginosa.
10. BavuwiniaA Portincert Prain; robusta scandens, ramulis lenti-
-cellatis, glabris, cortice brunneis; forsan cirrhifera. Folia quam lata
parum longiora, basi cordata, quadrante antico sinu angusto apiculato
290 =G. King and D. Prain— Descriptions of some New Plants [No. 2,
2-loba; crasse coriacea, supra nervis parce hirsutis exceptis glabra subtus
parcissime ferruginea, nervis 9-11, petiolo glabro, stipulis caducis. Flores
racemosi, racemis terminalibus ferrugineo-velutinis bracteis lanceolatis
pedicellos erecto-patentes fere aequantibus, bracteolis bracteis similibus,
alabastris clavatis parte superiore oblonga basin ampullaeformem exce-
dente, Calyx ferrugineo-pubescens, limbo 5-partito segmentis oblongo-
lanceolatis tubo basin versus parum dilatato. Petala 5, subaequalia,
oblanceolata obtusa, longe unguiculata magnopere exserta, utrinque
dense sericea. Stamina 3 fertilia, antheris lineari-oblongis, filamentis
medio parum incrassatis. Ovariwm distincte stipitatum, dense ferrug-
ineum, stylo crasso ferrugineo, stigmate obliquo peltato. Legumen
ignotum.
In montibus Kacuin nuncupatis, inter Namlao et Bansparao,
Pottinger !
Folits 2°5-4 poll. longis, his 2°25-3 poll. latis, petiolo 1-1°3 poll.
longo. Racemis 6 poll. longis, 4°5 poll. latis, pedicellis *75 poll. longis ;
alabastris 1°75 poll. longis. Calycis tubo ‘75 poll., limbo 1 poll. longo.
Petalis 2 poll. longis. Filamentis 2°5 poll. longis.
A very fine species, nearest to B. nervosa, a Khasia plant, from
which it differs in its leaves with fewer nerves, its shorter pedicels, and
its rather larger petals silky instead of rusty externally. In the size of
its flowers its only rival in the group to which it belongs is B. excelsa
BL, from Borneo; the shape however of the petals is different, those of
B, eacelsa being narrower and more acute at the apex.
SAXIFRAGACE AL. ©
11. Hypranewa Porrinesri Prain; fruticosa, ramis novellis pube-
scentibus ; foliis oblongo-lanceolatis acuminatis margine basi cuneato
excepto serratis, utrinque nervis adpresse puberulis ceterum glabris,
nervis 9-10-paribus ascendentibus; cyma ampla dichotoma ramis
pedicellisque pubescentibus ebracteata, florum radiantium sepalis 3
breviter unguiculatis late ovatis subacutis versus apicem grosse serratis
ceterum integris venis prominulis utrinque reticulatis glabris, fertilium
dentibus calycinis triangulis tubo brevioribus; petalis . . . 3; stam-
amibus . « « .« . 3 styls 3 erecto-patulis ovario globoso parum
brevioribus,
In montibus Kacuin nuncupatis, 4100 p. s. m., Pottinger !
Folia 3-4 pollicaria, 1°25 poll. lata, petiolis ‘5-6 poll. Cyma_ pe-
dunculo 1:25 poll. longo, pedunculis secundariis 1-pollicaribus, pedicellis
florum radiantium gracillimis 1°5 poll., pedicellis fertilibus ‘2-25 poll.
Capsula ‘08 poll. diam.
ra tig
—— >, wes
a I i i so
1898. ] from the North-Eastern Frontiers of India. 291
| POTTINGERIA Prarn.
Calycis tubus brevis late campanulatus basi ovarii adnatus, lobis 5
ovatis acutis persistentibus sinubus latis. Petala . . . Stamina 5
erecta ad marginem disci perigyni affixa, filamentis sursum kita
basi parum explanatis ibique extus glandula mediana ornatis ;* antherae
. . .3 Ovarium semisuperum. Capsula supera oblonga! longi-
fadinaliter parum 3-sulcata per stylos 3-partibilis stigmatibus cohaeren-
tibus septicide 3-valvis, placentis a marginibus introflexis carpellorum
simulac secedentibus persistentibusque, singulis utroque margine semina
circa 10 triente summo tantum fertili gerentibus. Semina anguste
fusiformia testa crustacea parum reticulata utrinque parum producta ;
embryo cylindrica majuscula in axe albuminis carnosi.—Folia alterna
brevipetiolata, 5-nervia. Cymae multiflorae axillares. Species singula
montium Kachin incola.
12. Porrrncgerta ACUMINATA Prain; folia ovato-acuminata crasse
coriacea utrinque glabra subtus punctata, breve petiolata, margine
integra nervis 5 mediano proximisque subaequalibus marginalibus
tenuioribus omnibus plus minus subtus prominentibus; cymae axil-
lares pedicellis gracilibus calyce paullo longioribus, bracteis parvulis,
In montibus Kachin, 3,000 p.s.m.; Pottinger !
Folia 2-3°5 poll. longa °75-1'5 poll. lata, petiolis :2 poll. longis.
Cymae 1‘5-2 poll. longae 1 poll. latae, pedicellis ‘15 poll. longis; capsu-
lis ‘2 poll. longis °15 poll. latis.
A very distinct genus of the Tribe Hscallonieae, apparently best
located near Jtea which it resembles in having a similarly partible style
with a similarly half-superior ovary but from which it differs markedly
in having 3 instead of 2 carpels and in the capsule so dehiscing that
when the valves fall away the three filiform placentas persist. The
cinnamon-looking or Melastomaceous-like leaves and the very different
inflorescence also help to give it a quite distinct facies.
Unfortunately our solitary specimen has been collected just as the
plant was passing out of flower so that the petals and anthers have all
dropped, and it is not therefore possible to state whether the former
are valvate or imbricate.
COMBRETACE ZA.
13. TERMINALIA ARGYROPHYLLA King § Prain; arbor magna,
ramulis gracilibus foliisque utrinque dense tomento adpresso persistente
argyreis; foliis suboppositis petiolatis ovatis basi rotundatis apice
acutis, nervis 8-9-jugis astendentibus subtus prominulis, petiolis apice
2-glandulosis ; floribus parvis, lutescentibus, spicatis, spicis in paniculis
terminalibus dipositis, bracteolis lanceolatis deciduis quam flores duplo
d. 1. 37
292 =G. King and D. Prain— Descriptions of some New Plants [No. 2,
brévioribus; calyce extus glabro limbo late campanulato lobis 5 parvis
acutis, intus dense argyreo-villoso, tubo ovato tereti; fructus.
Tn montibus KacuHIn nuncupatis ; Kingit mercenar. !
Folia petiolis °75 poll., laminis 4 poll. longis 1°75 poll. latis. Spicae
singulae #8°5—4 poll. longae, paniculis 8 poll. longis, 6 poll. latis.
This is very different in foliage from any species of J'erminalia in
Herb. Calcutta or in Herb. Kew; it is reported by the native collector
(Shaik Mokim) to be a “timber tree.” The fruits sent as belonging
to it are drupes shaped like those of 7. Chebula but much smaller, being
only ‘5 in. long; as however there are none of them attached to leaf-
specimens it must remain for the moment doubtful if they really belong
and if therefore the species is really referable to § Catappa, which
must be the case if the fruits in question be those of this tree.
CUCURBITACEA.
14. AtsomiTRA puBIGERA Prain; folvis breve petiolatis, pedato-5-
foliolatis, foliolis petiolulatis, membranceis ovatis, acutis subobtusis
vel retusis, margine integris puberulis; basi, terminali excepto, parum
obliquis, membranaceis, supra nervis densius ceterum parcissime
puberulis ; subtus, nervis exceptis, glabris, penninerviis, petiolulisque
dense puberulis; cirrhis apice bifidis; fructw puberulo; seminibus
stramineis utrinque spinuloso-rugosis.
In montibus Kacutn nuncupatis, Kingit mercenar. !
Alte scandens; rami graciles elongati, ramosi, puberuli sulcati,
Petiolus vix striatus °4-"5 poll. longus; petioluli, terminalis °3 poll.,
laterales ‘15 poll. longi; foliola utrinque intense viridia, 1°5—4 poll.
longa, °75-2'0 poll. lata. Czrrhi graciles sulcati pubernli. Paniculae
majusculae valde pluriflorae. Pedwnculus communis lateralis termina-
lisve, gracilis parum sulcatus dense puberulus 2-4 poll. longus,
pedicelli capillares puberuli ‘4 poll. longi, bracteolae subulate. Calyx
puberulus segmentis lanceolatis, linearibus, acutis, corolla glabrius-
cula, segmentis ovatis acutis ‘l poll. longis. ructws subcylindricus
densius velutino-puberulus, ab apice ad basin leviter attenuatus, apice
truncatus, basi subacutus 2°25 poll. longus, "5 poll. crassus. Semina
ambitu subtriangularia, margine profunde lobata basi oblique attenuata,
‘3 poll. longa, ‘25 poll. lata, °15 poll. crassa ; ala obliqua alba translucens
anguste oblonga, apice rotundata‘75 poll. longa, *25 poll. crassa, utrinque
areola clypeata spinuloso-rugosa exsculpta.
This very distinct species is most nearly related to A. clavigera, the
fruits, except for being densely puberulous, being very like those of that
species. But it differs very markedly in its pedate leaves and in its
spinulose-rugose seeds. By an oversight a number of flowering speci-
1898. ] from the North-Lastern Frontiers of India. 293
mens of this species have been distributed to various European Herbaria -
under the name Gynostemma pedatum ; recipients of these specimens are
hereby requested to correct the name. ‘These flowering examples were
received in November, 1897, the fruiting ones in January, 1898.
ARALIACEA.
15. PENTAPANAX STELLATUM King; scandens, novellis digito
minimo fere crassis, cortice pallido glabro lenticellis ornato. Folia
pinnata, rachide gracile glabra basi parum dilatata; foliolis 5, jugis 2
cum terminali, late ovatis vel ellipticis, apice abrupte acutis, basi rotun-
datis margine integris parum recurvis; supra glaberrimis, subtus pilis
longioribus stellatis dense obtectis; nervis 4—5-jugis parum incurvis
subtus prominulis supra distincte impressis; petiolulis maequilongis
lisque jugi summi fere 0. Panicula terminalis basi bracteis lanceolatis
pluribus obsita, sparse ferrugineo-puberula, ramis inferioribus patentibus
remotis, singulis umbellas plures 15-25-florales pedunculatas ferenti-
bus, parte summa umbellas simplices ferente. lores late oblongi;
calycis tubo late campanulato, limbo 5-dentato, dentibus latis obtusis ;
petalis late ellipticis. Fructus ovoideo-globosus prominenter 5-costatus,
glaber. .
Burma: in montibus Shan nuncupatis, apud Fort Stedman; Kingit
mercenar. !
Folia 9-15 poll. longa, foliolis 3-5 poll. longis, 2°5-3'25 poll. latis
petiolulis folioli terminalis 1-1°5 poll., lateralium inferiorum ‘1-15
poll. longis, lateralium summorum fere obsoletis. Panicula 12-18 poll.
longa, ramis inferioribus 3-4 poll. Flores *1 poll. lati. Fructus ‘15 poll.
longus.
Peutapanax is a small genus of which hitherto only six species
have been described ; and of these only one has hitherto been recorded
from Burma. The species now for the first time described differs from
all the others in the dense and very peculiar pubescence by which the
under surfaces of the leaves are clothed. The hairs are long, flexuose,
and united to form large stars of a pale yellowish-brown colour.
16. Hepraptevrum (§Agalma) Lawranceanum Prain; arbor ?,
novellis parce puberulis. Folia digitata foliolis 7-9, late ellipticis apice
mucronulatis, margine integris utrinque glaberrimis, crasse coriaceis ;
nervis 20-30-juzis obscuris petiolulis glabris. Panicula ramosa, ramis
puberulis ramulos fere farinoso-puberulos umbelliferos distinctes emit-
tentibus, umbellis 12-—20-floris, pedicellis aequilongis floribus parum
brevioribus. Calyx margine truncatus. Petala valvata, circa 7, trian-
gula, intus glabra extus pilis coactis dense obtecta. Stamina petalis
isomera antheris oblongo-ovatis sursum parum angustatis. Lructus
294 G. King and D. Prain— Descriptions of some New Plants [No. 2,
turbinatus apice truncatus, medio columna cylindrica (stylis connatis)
coronatus, 7-angulatus.
BuRMA SUPERIOR: in montibus Kachin ; Pottinger !
Foliola 10 poll. longa, 6 poll. lata. Paniculae rami pedales, ramulis
1:25 poll. longis, pedicellis °2 poll. longis. Flores ‘25 poll. longis, ‘2 poll.
latis.
17. Denpropanax Lisrert King; arbuscula glabra parva, novel-
lis cortice grosse lenticellatis ex sicco pallide brunneis. Jolia simplicia,
tenuiter coriacea, late elliptica breviter acuminata, basi cuneata, mar-
gine integra vel dentibus paucis remotis minutis irregulariter serrata ;
utrinque glaberrima supra hebetia; subtus reticulato-venosa, nervis
secundariis distinctis; costa mediana subtus prominente a basi venas
2 ceteris crassiores fere ad apicem ascendentes saepius etiam 2
tenniores marginales emittente, lateralibus supra laminam mediam
3-4-jugis curvatis; petiolis gracillimis inaequilongis. Panicula axil-
laris, ramis paucis umbellatis, umbellis 4-5-floris, floribus subglobosis
pedicellis gracillimis. Calycis tubus subglobosus limbus angustatus
margine minute 5-dentatus. Fructus sphaericus stylis brevibus basi
connatis apice recurvis coronatus.
In montibus Dapruua nuncupatis, apud Torupati, 5,500 p. s. m., J. L.
Lister!
Arbuscula 20-pedalis. oliorwm laminis 3°5-8 poll. longis, his
1'65-4 poll. latis; petiolis 15-7 poll. longis. Flores ‘15 poll. diam.,
pedicellis *3 poll. longis. Fructus *2 poll. diam.
This very distinct Dendropanaz was collected by Mr. J. L. Lister,
in whose honour it is named, when accompanying the Daphla Hill ex-
pedition of 1874,
CORNACEAE.
18. Aanaium Kinatanum Prain ; frutex scandens, inermis, novellis
puberulis; jfolia membranacea, oblongo-ovata, basi truncato-cuneata,
apice rotundato demum breviter acuminata utrinque nervis puberulis
ceterum puncticulata, basi sub-trinervia nervo mediano robustiore
nervos ascendentes 5-6-jugos emittente ; flores in cymis laxis axillaribus
foliis multo brevioribus dispositi, pedunculis pedicellisque puberulis ;
calyce dense puberulo breviter 7-dentato, petalis lutescentibus extus
puberulis saepissime 7, anguste linearibus apice subacutis; staminibus
14, filamentis brevissimis pubescentibus, antheris linearibus; drupis
parcissime adpresse puberulis, parum compressis, longitudinaliter 14-
decim lineatis basi roundatis apice subacutis.
In montibus Kacuin nuncupatis, apud Agata Kedan, etc., Kingii
mercenar. !
Folia 4-6 poll, longa, 1:5-2°5 poll. lata, petiolis ‘25 poll. longis.
1898. | from the North-Eastern Trontiers of India. 295
Cymarum pedunculis ‘5-75 poll., cymis 1 poll. latis 8-12-floris,
pedicellis ‘25 poll. longis. Flores ‘3 poll. tantum longis. Drupae ‘4
poll, longae, °25 poll. latae.
A very distinct species not before represented in Herb. Calcutta or
in Herb. Kew; the affinity, Dr. Stapf has been kind enough to note at
Kew, is with Alangiwm Faberi Oliv., a species not present at Calcutta.
19. MastixiA nvonyMoIDES Prain; arbor, foliis oppositis longius-
cule petiolatis, laminis ovatis apice acuminatis basi cuneatis, margine
integris, crasse coriaceis, supra intense viridibus subtus prasinis, utrinque
glaberrimis, nervis 6-8-jugis subtus distinctioribus parum ascendenti-
bus; thyrsus laxus dichotomus, pedunculo ramisque glabris; flores
ignoti; fructus anguste ellipsoideus.
In montibus Kacuin; Kingw mercenar. !
Folia petiolis °75 poll. longis, laminis 4 poll. longis, 1°75-2 poll.
latis ; pedunculis 2°5 poll. longis, thyrsis 2—-2°5 poll, latis; fructus calycis
limbo 4-dentato coronatus, °6 poll. longus, °3 poll. crassus. —
A very distinct species.
RUBIACE A.
20. Opniorraiza LAwranceana King § Prain; caulis brevis basi
radicans adscendens vel 0; folia elliptico-oblonga, apice acuta basi
cuneata, petiolis brevibus parce puberulis, laminis nervis subtus parce
puberulis, ceterum utrinque glaberrimis, stipulis e basi trianguli fili-
formibus, cymae longe pedunculatae congestae glabrae, bracteolis spatu-
latis obtusis glabris persistentibus, calycis dentibus brevibus triangulis,
corolla brevis tubo cylindrico, limbo angustato; capsula glabra.
In montibus Kachin nuncupatis, Kingii mercenar .!
Folia 1:25-3°5 poll. longa, °75-1°5 poll. lata, lurida. Cymae ‘3-'5
poll. latae, pedunculis gracilibus 3 poll. longis. Corolla ‘2 poll. longa,
tubo angustato.
Very similar to O. lurida Hook. fil. from the Hastern Himalaya in
size and habit, and no doubt related to that species. The bracts are
however different and the corollas are smaller and much narrower.
21. ParpertA Cruppasiana Prain; volubilis corolla excepta omnino
glaberrima; folia opposita petiolata ovata basi truncata apice acuta;
flores congeste cymosi in paniculis axillaribus terminalibusque laxis per
paribus distantibus dispositi; calycis dentibus 5 brevibus triangulis, tubo
campanuluto, bracteolis subulatis ; corolla extus puberula tubularis, intus
dense tomentosa ; fructus oviformis calycis dentibus coronata.
In montibus Kacuin nuncupatis, Kingii mercenar. !
Folia 2-5 poll. longa, 1-2°5 poll. lata, petiolis 1-1:25 poll.
longis; paniculae 4-8 poll. longae, ramis ‘5-1'5 poll. longis, cymis
296 G. King and D. Prain— Descriptions of some New Plants [No. 2,
singulis terminalibus ‘5-75 poll. latis. Corolla °5 poll. longa. Fructus
"25 poll. longus, ‘3 poll. latus.
This very distinct species belongs to the group characterised by
having the fruits uncompressed and differs very markedly from the
other species of that group in having the fruits egg-shaped, narrowed
upwards from the middle, and not subglobose rounded at the top as in
P. tomentosa. From P. linearis, the other Indian species referred to the
group, it differs much in foliage—its general facies, except for the
fruit, being very much that of the common P. foetida.
VACCINIACE A.
22. Acapetes Porrincert Prain ; frutex epiphytica, ramis adpresse
puberulis et pilis rigide setaceis patentibus ferrugineis simulac obsitis.
Folia sessilia ovato-lanceolata a basi fere rotundata sensim ad apicem
longius acuminatam attenuata, margine integra, coriacea, utrinque
glabra, nervis 8-10-jugis supra distinctioribus. Inflorescentia corymbosa
ramiflora pedunculo pubescenti bracteis rigidis cincto, pedicellis gracili-
bus pubescentibus basi bracteatis, bracteis majusculis ovato-lanceolatis
rigidis striato-reticulatis margine puberulo excepto glabris. Cualycis
tubo globoso cum apice pedicelli parum ampliati articulato extus pilis
longis fulvis setaceis apice glandulosis patentibus obsito, limbo cam-
panulato margine 5-dentato prorsus reticulato, dentibus triangulis quam
partem limbi connatam brevioribus, intus glabro extus pilis flaccidis
longis sparse pubescente. Corolla tubulosa recta medio parum ampliato ~
sub limbo breviter 5-lobo lobis late triangulis subobtusis parum
contracta, extus parce pilis flaccidis pubescente, lobis viridibus ceterum
rubris nec lineis notatis. Stamina 10, epigyna, libera, filamentis antheras
fere aequantibus, antherisque glabris; antherarum tubulis corallae
linbum vix attingentibus, dorso 2-calcaratis. Ovariwm 5-loculare stylo
filiformi apice brevissime 5-lobulato ovulis numerosis.
In montibus Kacuin nuncupatis, 4100 p. s. m., Pottinger !
Folia 5-7 poll. longa, 1°5-2°25 poll. lata. Corymbis paucifloris 1°5
poll. longis, pedunculis ‘2 poll., pedicellis ‘5 poll. longis, bracteis -2-25
poll. longis. Calycis tubo ‘1 poll., limbo ‘35 poll. longo, dentibus ‘15 poll.
longis. Corolla ‘65 pall. longa.
This remarkably distinct species is separable from all hitherto des-
cribed Agapetes by its large bracts and its large calyx-limb, the teeth of
which are not partite to the disk as in our other species. It may be
taken therefore as the type of a distinct section (§ Holocalyx) to be
distinguished as follows from the other sections defined in the Genera
Plantarum it. 571 :—
§ Hotocatyx. Corolla elongata recta parum ventricosa breviter
1898. | from the North-Eastern Frontiers of India. 297
d-fida lobis erectis. Stamina recta antherae dorso ecalearatae. Bracteae
majusculae, calycis limbus in dentibus 5 prorsus haud solutus.
DESMOGYNE Kine & Prarn.
Calycis tubus teres pedicelli apice ampliato crateriformi involutus
subglobosus ; limbus magnopere ampliatus late campanulatus, margine
integer, persistens. Corolla tubulosa anguste infundibularis, elongata,
tubo terete recto breve 5-lobo, lobis erectis. Stamina 10, epigyna,
corolla aequilonga filamentis glabris basi inter se et a tubo corollae
liberis; antherae elongatae rectae liberae dorso muticae tubulis connatis
tenuibus strictis membranaceis, loculis extus muriculatis. Ovarium
5-loculare ; stylus filiformis stigmate lobulato; ovula in loculis singulis
numerosa placentis angulo interiori adnatis.—Frutices epiphytici, foliis
alternis persistentibus breviter petiolatis coriaceis integris, Species
singula, montium Indiae transgangeticae incola.
23. DESMOGYNE NERIIFOLIA King & Prain; frutex epiphytica,
ramis glabris gracilioribus, olia alterna oblongo-lanceolata apice longe
caudato-acuminata basi rotundata breve petiolata margine integra, crasse
coriacea nervis mediano subtus prominente supra impresso excepto
obscuris, Inflorescentia corymbosa pedicellis versus apicem pedunculi
sursum incrassati in axillis bractearum triangularum approximatis ;
pedicellis sursum incrassatis apice cupularibus. Calycis tubus globosus
in fundo epicalycis articulatus eoque involutus, limbus inflatus late cam-
panulatus margine integer prorsus prominenter reticulatus. Corolla
elongata recta infundibuliformis 5-loba, lobis brevibus trianeularibus.
Stamina 10, epigyna libera filamentis brevibus antheris elongatis erectis
apicibus corollae limbum attingentibus, dorso muticis. Ovartwm 5-locu-
lare, stylo filiformi apice breviter 5-lobulato; ovulis numerosis. Fructus
ovatus calycis limbo persistente coronatus.
Borma: in montibus Chin etiam in montibus Kachin, Kingii
mercenar. ! .
Folia 4-6 poll. longa ‘7-9 poll. lata; petiolis ‘2 poll. Pedunculi
axillares ad 2°59 poll. usque longi, bracteis triangulis ‘1 poll. longis,
pedicellis 1:2 poll.; calycis tubo *15 poll. longo, limbo °3 poll. longo
latoque. Corollae tubo 1°5 poll. longo, limbo °5 poll, diam., lobis ‘2 poll.
longis. ructus *4 poll. longus, °35 poll. diam.
A very distinct species of the group of Vacciniaceae that constitutes
the ‘‘ genera” Agapetes G. Don., and Pentapterygium Klotzsch. From
all Pentapterygia it differs in having the calyx neither winged nor
ridged and while agreeing with Agapetes in having a smooth calyx
tube it differs from all the known species of that genus in having
the calyx- limb large, entire and widely campanulate, and, Sarthe,
298 G. King and D. Prain— Descriptions of some New Plants [No. 2,
in having the calyx-tube enveloped by the expanded cupular apex of
the pedicel. With reference to this last character the name Desmogyne
(Secpds, a bandage ; yvv7) has been applied to the plant. We feel in-
clined to think that Agapetes and Pentapterygiwm, which are distin-
guished solely by the absence from the former, the presence in the
latter, of ribs or wings to the calyx, are hardly separable as genera. If
these two could be united our plant would then be the type of a section
Desmogyne within this enlarged Agapetes. But seeing that in the three
most authoritative treatises on the genera of plants—the Genera Plan-
tarum, the Histoire des Plantes, and the Natiurlichen Pflanzenfamilien,—
Messrs Bentham and Hooker, Baillon, and Drude have considered it
necessary to keep Pentapterygium apart from Agapetes, we are constrained
to give our Desmogyne, at least for the present, the rank of a genus;
it differs more markedly from either Agapetes or Pentapterygium,
than these two differ from each other. The undivided calyx-limb,
at first sight a more remarkable feature than the ball and socket
arrangement at the apex of the pedicel is not really so important a
difference as it appears; the species immediately preceding this
(Agapetes Pottingert Prain) stands intermediate as regards calyx-limb
between this and the other Agapetes since the limb though 5-lobed at
the margin is there also campanulate and gamophyllus below.
Dr. Stapf who has examined and kindly compared one of our
specimens at Kew agrees with us in thinking that so long as Pentuptery-
gium is kept apart from Agapetes our plant had better receive generic
rank, If Pentapterygiwm could only be reduced to Agapetes the present
species would probably have to be included in this enlarged genus as
Agapetes Desmogyne King & Prain.
PRIMULACEA.
24. Liystmacuta EvaLvIs Wall. in Roxb. Flor. Ind. ed. Carey §&
Wall, 11. 27 var. grandifolia Prain ; folia 6 poll. longa, 2°25 poll. lata;
peduncult 2 poll. longi.
In montibus Kacuin, apud Neochawng 2,500 p. s. m., Pottinger !
This may prove, when more completely represented, to be a distinct
species.
SOLANACEA.
25. Sovanum Frerox Linn. Sp. Pl. ed. ii. 267 var. inermis Prazn ;
omnino nisi aculeis absentibus cum S. feroce convenit.
In montibus Kacutn nuncupatis, Kingit mercenar. !
The collector’s note is ‘ Moima villge; flowers white.” There is
not a character whereby the plant can be separated from 8. ferox except
1898. ] from the North-Eastern Frontiers of India. 299
the complete absence of any trace of prickles whether on leaves or
stems, and in the less numerous needle-like hairs on the fruits.
It is well-known that many forms of S. Melongena under cultivation
entirely lose their armature; it is interesting to find that the same
may apparently happen with a wild species like S. ferow when growing
as a weed in the rich soil that characterises the vicinity of an Indian
hill-village.
GESNERACE.
26. AESCHYNANTHUS GRANDIFLORA Spreng. Syst. Veg. iv. 238 var.
longiflora Prain ; floribus 2°25 poll. longis, ceterum typi.
In montibus Kacuin, Kingii mercenar. !
27. ABSCHYNANTHUS MICRANTHA Clarke in Flor. Brit. Ind. iv. 340
vAR. Pottingeri Prain; capsulis 10-pollicaribus, ceterum omnino typi.
-In montibus Kacuin, Kingit mercenar. !
Quite possibly both these Aeschynanthi—of which the first is only
reported in flower, the second only in fruit—may prove when fully re-
presented to deserve specific rank.
28. AESCHYNANTHUS PUSILLA Prain ; rami elongati gracillimi, sparse
pilosi, nodibus prorsus radicantes, foliis ternatis parvis ovatis apice
acutis, basi cuneatis, °3 poll. longis ‘2 poll. latis utrinque pilis albidis
multicellularibus villosis, nervis obscuris, margine integris petiolis dis-
tinctis (‘L5 poll. longis), villosis; florzbus paucis terminalibus et in
axillis superioribus, ad nodos singulis, pedicellis gracilibus, °2 poll.
longis, dense pilosis; calyce 5-partito, segmentis lanceolatis tuboque
anguste campanulato pilis patentibus dense villosis; corolla extus
parce puberula, pallide flava limbum versus viridescente, lobis ipsis
intus tantum purpurascentibus, tubo 1 poll. longo dimidio inferiore
peranguste cylindrico, dimidio superiore anguste infundibuliformi,
limbo ‘2 poll. lato; filamentis inclusis filiformibus glabris; capsula adhuc
ignota.
In montibus Kacuin nuncupatis; Kingii mercenar. !
This graceful little species may be tentatively referred to the sec-
tion Haplotrichium, its general facies indicating that it is probably
closely allied to Aeschynathus gracilis; till fruiting specimens are ob-
tained, however, its precise position must remain problemetical.
29. Dripymocareus ELATIOR Prain; suffrutescens; rami juniores
pilis divergentibus rufescentes demum glabrati; folia ovato-lanceolata
basi cuneata apice acuminata margine minute serrata, petiolata, petiolis
rufo-pubescentibus, laminis supra parce adpresse puberulis subtus prae-
sertim nervis pubescentibus: cymae pauciflorae axillares pedunculis
gracilibus elongatis puberulis; bracteae lanceolatae ; pedicelli calyce
J. 1. 38
300 G, King and D. Prain— Descriptions of some New Plants [No. 2,
parum longiores saepius singuli; calye campanulatus ad medium usque
fissus dentibus ovato- acutis tubum aequantibus; corolla tubulosa fere
recta extus parcissime puberula, subsymmetrica, pallide purpurea;
capsula in pedicello erecta.
In montibus Kacuty, apud Sim, 5,000 p. s. m., prope rivulis ; Kingiw
mercenar. !
Foliorum laminis 3 poll. longis, 1°25-1'5 poll. latis, petiolis °75 poll.
longis; pedunculis 1°5 poll. longis saepius 3-floris, nonnunquam
(floribus lateralibus geminis) 5-floris; pedicellis *4 poll. longis ; calyce
‘25 poll. longo; corolla 1:2 poll. longa; capsula 1-1:25 poll. longa “lo
poll. lata.
A very distinct species, in habit most resembling D. corchortfolia
Wall., from Penang and Malacca, |
ACANTHACE.
30. RaHINACANTHUS caLcARATUS Nees in Wall. Pl. As. Rar. ii. 109
VAR. maxima Prain; foliis utrinque parcissime puberulis, panicula
condensata ; corollae tubo 1°5 poll. longo; capsula 2 poll. longa.
In montibus Kacuin, Kingtt mercenar. !
This will probably have to be considered a distinct species, Rhin-
acanthus maximus, when full material of the original species is obtained ;
as yet the fruit of Nees’ plant has not been collected. The present
plant has leaves that are exactly like those of the type in size and
texture; they only differ in being faintly puberulous on both sides,
those of Nees’ plant being glabrous ; its calyx and corolla are exactly
like those of Wallich’s Pl. As. Rar. t. 113 except that they are distinctly
larger.
HAEMODORACE.
31. OPHIOPCGON cOoRDYLINOIDES Prain; caule rigide erectiusculo,
crasso, nodis nec radicante, vaginibus ovatis viridibus margine late
scariosis mox deciduis suffulto, folsis late lanceolatis acuminatis 15-17-
nervis, petiolis angustis brevibus basi vaginis scariosis expansis, scapis
quam folia fere dimidio brevioribus, bracteis scariosis, pedicellis vix
longioribus, flortbus fasciculatis, albidis, quam pedicellos brevioribus,
segmentis ovato-oblongis, antheris lanceolatis filamentis brevissimis,
stylo filiformi ; fructw orbiculari.
In montibus Kacuiy, apud Namli, 2000 p. s, m., Pottinger ! Kingii
mercenar. !
Caulis penna cygni crassus, foliis 8-12 poll. longis, ‘75-1 poll. latis,
petiolis vix ] poll. longis. Racemi 3-4 poll. longi, bracteis ‘3 poll. longis,
pedicellis 25 poll. poets perianthio *2 poll. longo, ‘3 poll. lato; fructw
*2 poll. diam.
1898. | from the North-Hastern Frontiers of India. 301
A very distinct species with an elongated stem as in OC. dracaenoides,
but without roots at the nodes whence arise the tufts of leaves; the
vaginal sheaths are exactly as in ©. dracaenoides but the leaves proper
are longer and narrower and have shorter petioles. The flowers are
much as in C. dracaenoides but are slightly smaller ; the fruits are con-
siderably smaller.
LILIACEA.
32. Disporum putLuM Salish. Trans. Hort. Soc. i. 330. var. oblan-
ceolata Prain; foliis oblanceolatis, acuminatis, 8 poll. longis 2°5 poll.
latis, fasciculis circa 15-floris, pedicellis 1:25 poll. longis, fructibus ovatis
subacutis ‘2 poll. longis.
In montibus Kacuin, apud Lammuk, Pottinger !
This is unfortunately only represented by one specimen which is
without flowers. It is obviously most nearly related to D. pullum of
which it is for the moment treated as a variety, though there is hardly
room for doubt that when more fully represented it must be considered
a distinct species.
COMMELYNACE.
33. STREPTOLIRION VOLUBILE Edgew. Trans. Linn. Soc. 90 t. 2.
vAR. setosa Prain; caulibus, petiolis, foliorum marginibus, pedicellis,
bractearumque marginibus, pilis fuscis rigidioribus densius setosis ;
ceterum typi.
In montibus Kacuin ; Kingit mercenar. !
The setose stems, petioles and leaf margins give this plant a very
- distinctive facies, but it cannot be separated by any other character
from typical S. volubile, which is likewise sent by the same collector
from the Kachin Hills, and it will probably be found unnecessary to
accord this more than varietal rank.
AROIDE.
34. TypHoniumM inoprnatuM Prain ; foliorum petiolis quam lamina
triplo longioribus ; lamina ovata apice acuta base sinu latiore cordata ;
pedunculo petioli partem vaginalem fere aequante; spathae tubo sub-
orbiculari quam lamina prorsus suberecta sensim acuminata sexties
breviore ; tubo utrinque viridi, limbo viridi extus basi lineatim obscure
purpurascente intus basi lineis sursum maculis purpureis notato; in-
florescentia foeminea fertili quam mascula multo breviore, pistillodiis
perpaucis simplicibus vel bifurcatis parum recurvis ; spadicis appendice
tereti anguste conoidea vix stipitata reliquam spadicis partem parum
excedente.
In Burma superiore prope Myitkyina, Kingw mercenar. !
302. =G. King and D. Prain— Descriptions of some New Plants [No. 2,
Foliorwm petioli 15-2 dm., pars vaginalis 4-5 em., lamina aeque
8 cm. longa lataque, sinu aeque 2 cm. lato altoque. Pedunculus 4-5 em.
longus, spathae tubus 2 cm. longus, 1°5 cm. latus, limbus 9-10 cm.
longus, triente imo 4°5 cm. latus apice subacutus. Inflorescentia mascula
cylindrica pallidiflora 1 cm. longa ‘45 cm. lata, antherarum thecae
rimis porosis apertae; pars foeminea pallide punicea *2 cm. longa ‘5
cm. lata, pistillodiis basi purpareis medio albis apice viridescentibus
explauatis 5 cm. longis. Inflorescentia tota 8 cm. longa, parte sterili
5 om. longa. Fructus baccatus viridis pallide rubro-suffusus, ovalis, °5
em. longus, ‘4 cm. diam.
This interesting species was first noticed in Oct. 1895, in one of
ithe flower-beds in the Royal Botanic Garden, in which it had appeared
spontaneously. A drawing was made and sent with specimens to Kew,
where it was examined by Mr. N. E. Brown, of the Kew staff, a very able
student of Aroidee; Mr. Brown agreed with us in thinking it new.
No light could be thrown, at the time of its first being noticed, on its
original habitat, though its introduction had apparently not been recent,
seeing that it was subsequently found, when looked for, in almost every
part of the Botanic Garden. The communication of specimens from
Myitkyina in Northern Burma seems at last to definitely settle the
original source of the species. Itis noteworthy that the commonest
of the Typhonia in and around the Royal Botanic Garden, Typhoniwm
trilobatum Schott (Arum trilobatum Linn.=Arum orixense Rowb.), is
evidently a plant introduced here during or since Roxburgh’s incum-
bency as Superintendeut (1793-1815). Roxburgh did not collect it
in Bengal and we have been unable to find it in Bengal ourselves, or to
learn that any one has found it in Bengal except in the immediate neigh-
bourhood of these Gardens—where it is scarce, and inside them—
where it is abundant. Next most common in these Gardens is 7’. inopina-
tum, the species just described, while fairly plentiful but less common than
either is a species that during Roxburgh’s superintendentship was acci-
dentally introduced from the Moluccas, and that he has described as Arwm
trilobatum in Flora Indica, 111. 505, but that is not the Arum trilobatum of
Linnaeus, Sp. Pl. ed. princeps, though it was included by Linnaeus with
the true 7. trilobatum in his Systema ed. x. and his Sp. Pl. ed. ii. The
true Typhonium trilobatum is based on a figure by Hermann (Par. Bat.)
of the Ceylon “ Panuala” which Thwaites, Trimen and others identify
with Arum orizense Roxb. whereas Roxburgh’s plant is the same as
Rumphius’ Arisarum amboinicum (Herb. Amboin. V. t. 110, f.2). It is
usual to give the name T'yphorium Roxburghit to Roxburgh’s plant, on
the authority of Schott, but there is a slight objection to this in the
fact that Schott gives a figure of the plant which he names 7. Roxburghi
1898. ] from the North-Eustern Frontiers of India. 303
and which he takes to be Roxburgh’s one, but which differs altogether
from Roxburgh’s in habit, in length of peduncle (twice instead of half as
long as vaginal portion of leaf-stalk), size of spathe, nature of pis-
tollodes, distance between male aud female portions of inflorescence and
space between male inflorescence and barren appendix. This misiden-
tification is the more inexplicable since Roxburgh has left a very
accurate coloured drawing of the plant intended by him, a drawing
that has been copied by Wight as his Ic. t. 803, and since Schott himself
expresses a doubt whether the plant which he figures as T. Roxburghii be
the same as the Arisarwm amboinicum Rumph. V., t. 100 f. 2, with which
Roxburgh identified his plant. The coloration of the plant figured by
Saunders (Ref. Bot. t. 283) closely approximates to the true Roxburgh-
ian plant, but the tip of the spathe does not twist asin 7. Roxburghii,
the plant which Roxburgh figures. The tip of the spathe does not
twist in the figure given by Rumphius though the account given of the
colour in the Herb. Amboinense agrees well enough, and for the matter
of that, the tip does not always twist in the plant as it grows; the
chief objection to Rumphius’ plant being ours is that its peduncle is
much too long. What makes matters more complicated is that we have
yet another species of Typhoniwm which grows, as if wild,in the Royal
Botanic Garden, and which has all the characters of the plant that
Schott figures. This species, for the writer is inclined to treat it
as a distinct plant, has a white barren appendix in place of the
bright red or terracotta coloured appendices of TZ. trilobatum and
T. invpinatum or the dark purple very long and slender appendix of
Roxburgh’s plant from the Moluccas. Perhaps tbe simplest solution of
the tangle is to quote the Moluccas plant as T'yphonium Roxburghii
Schott (as to citation 7. trilobatwm Roxb.) Arozd. i. 12 (excl. t. 17),
Prodr. 106 (in part); Saunders, Ref. Bot. t. 283= Arum trilobatum Roxb.
Flor. Ind. iii. 505; Wight, Ic. t. 803; and to cite the hitherto unnamed
and undescribed Botanic Garden species which Schott has figured, as a
new species, Typhoniwm Schottit Prain=T. Roxburghii Schott Aroid. t.
17 (excl. descript.). A reference to the original works will show that in
his Prodromus, Schott describes the coloration of T. Roxburghit in terms
that are only applicable to Roxburgh’: Moluccan plant, and says that
the description is based on dried specimens and drawings; in his Avozd-
dec, Schott does not venture to describe colours and it may be safely
assumed from this that both the description and the drawing are from
dried specimens only. Evenif in bothinstances the description may be held
to include Roxburgh’s plant, yet the drawing is certainly that of another
species. As yet we have been unable to find where 2’. Schottii is really
wild. The only truly wild and unintroduced species in Lower Bengal is
304 G. King and D. Prain — Descriptions of some New Plants [No. 2,
T’. cuspidatum Bl., and curiously enough this happens to be mnch the
rarest of the five that are to be found within the limits of the Gardens.
We have been unable to find it noted that the otherwise excellent figure
which Blume gives of 7’. cuspidatum makes the curious mistake of revers-
ing the position of the lower pistillodia. These are cymbiform organs
with the concavity directed upwards in the natural state; in Blume’s
figure the concavity is made to look downwards.
Before leaving this subject it may be pointed out that though
Roxburgh has cited Loureiro’s Arum trilobatum as equivalent to his
A. orixense, this is by no means clearly the case. The pistillodia of
A. orixense (the true A. trilobatum) are, as Roxburgh describes them,
yeliow; those of A. ¢rilobatwm Loureiro, are described, on the other
hand, as red. Thetruthis that the genus Typhonium requires more
careful and extended study, from living plants, than it has yet received.
35. TypHonium Listert Prain; foliorum petiolis quam lamina
dimidio longioribus ; lamina pedatisecta 5-foliolata, segmentis mediano
sessili reliquis per paria breve petiolulatis omnibus anguste ovatis
basi cuneatis apice sensim acuminatis; pedunculo purpureo brevissimo
bracteis cataphyllariis obtecto; spathae tnbo subcylindrico quam
spatha subito refracta quadruplo breviore; tubo extus laete viridi
intus lutescenti, limbo extus margine purpurea excepta laete viridi
intus purpurascente; inflorescentia foeminea fertili quam mascula
parum tantum breviore, pistillodiis paucioribus majusculis ligulatis
deflexis purpureis; spadicis appendice tereti parum stipitata basi
plus minus obliqua concolore lactea, abrupte refracta et spathae limbo
involuta reliquam spadicis partem triplo longiore.
In prov. Cuirracone ; Inster! in Assam; Watt!
Foliorum petioli 2°5-3 dm. longi, pars vaginalis 1:5 cm., cata-
phyllis spathaceis 3-7 cm. longis; laminae segmentis 1:4 dm. longis,
6 cm. latis. Pedunculus vix 1 cm. longus, spathae tubus 4 cm. longus,
2 cm. latus, limbus angulo angulum rectum parum excedente refractus
1-4. dm, longus, 7 cm. latus apice acutus. Injflorescentia mascula cylin-
drica 1'7 cm, longa, ‘9 cm. lata, pars foeminea conica 1 cm. longa, basi
15 cm. lata; parte sterili 1*2 dm. longa, 1°2 cm. lata,
A very distinct species.
36. TypHontum Porrincert Prain; foliorum petiolis quam lamina
duplo longioribus, lamina profunde tripartita partitionibus subaequi-
longis intermedia oblongo-elliptica acuta, lateralibus oblongo-lanceolatis,
lobo triplo breviore oblongo-obtuso subretuso auctis ; pedunculo quam
petioli pars vaginalis duplo breviore ; spathae tubo ovato vel oblongo
quam lamina imo tertio ovata erecta sursum recurva et sensim longe
acuminato-caudata octies breviore ; tubo extus viridi intus rubescente,
1898. | H. N. Ridley — New species of Entada. 305
limbo basi tantum intus rubescente supra puniceo-maculata extus con-
colore pallide viridi; inflorescentia foeminea fertili quam mascula. multo
breviore, pistillodiis numerosissimis varie flexis; spadicis appendice
tereti anguste conoidea breviter stipitata reliquam spadicis partem
magnopere excedente.
In montibus Kacuin, prope Myitkyina, Kingit mercenar. !
Foliorum petioli 2-2'5 dm., pars vaginalis 2°5-3 cm., laminae parti-
tiones ‘8-12 dm. longae ‘75-1 dm. latae, lobi lateralium basales 4-6
em. longi, 3-4. cm. lati. Pedunculus 1-1'5 cm. longus; spathae tubus
3 cm, longus, 2°5 cm. latus limbi pars triens inferior 7 cm. longa, 9 cm.
lata pars summa caudata reflexa basi 3°5 cm. lata, 1°4 dm. longa, sen-
sim apice longe acuminata attenuata. Inflorescentia mascula cylindrica
rubra 1 cm. longa ‘7 cm. lata antherarum thecae rimis porosis apertae ;
pars foeminea alba ‘5 cm. longa, ‘85 cm. lata, pistillodiis albis expla-
natis 1:2-1°5 cm. longis. Inflorescentia tota 4 dm. longa, appendice
sterili 16 dm. longa.
In addition to the forgoing Aroidex, an undescribed Amorphophallus,
which has been included in a List of Kachin Plants, published in the
Records of the Botanical Survey of India as A. Cruddasianus, should be
here alluded to. Complete material has been sent by our Garden
Collector, but for the moment we prefer to withhold a detailed descrip-
tion till living flowers are available, when an accurate account of the
coloration can be given. Our collector has sent also a number of living
tubers, but during the past season these have sent up leaves only. The
tubers alone, however, furnish characters that are sufficiently
diagnostic; in place of being oval or depressed, as in other Amor-
phophallt hitherto described, these in A. Cruddasianus are long and
parsnip-shaped, 6-10 in. long, 2-3 in. across the top.
New species of Entada from Singapore and Borneo—By H. N. Rivtey,
Hsq., M.A., F.L.8., Director, Botanic Gardens, Singapore. Com-
municated by SurceON-Masor D. Prain.
The genus Entada is represented in the Malay Peninsula by the
well-known E. pursaetha, and a species very common in Singapore
which seems to have been entirely overlooked, although it is very con-
spicuous here from its very remarkable fruit. To this plant I propose
to give the name Hntada spiralis.
HE. SPIRALIS, n. sp. A woody climber with twisted somewhat
flattened spiral stems about 2 in. wide and 1-2 in. thick in the thickest
part in large specimens; thinner on the outer edge of the curve
Leaves alternate three inches long (excluding the tendril), petiole one
306 H. N. Ridley — New species of Kutada, [No. 2,
inch, much swollen at the base, pinne four alternate with petiolules
one inch long swollen at base, leaflets alternate rarely opposite, some-
what distant, with very short stalks elliptic to obovate-oblong retuse
inaequilateral, 2 to 3 in. long, 1 in. or less wide, dark green un-
polished above, glaucous beneath, tendril long, bifid at the apex.
Flower-spikes axillary 6 in. long, péduncle 2 in. long, swollen at
base, rachis purplish-brown covered with short stiff hairs. Bracts
minute lanceolate hairy. Flowers copious densely crowded, shortly
pedicellate. Calyx campanulate with five teeth, hairy, green. Corolla-
lobes 4 or 5, oblong obtuse glabrous green, 2 mm. long. Stamens 8 to 10,
at first white soon becoming yellow, filaments ; inch long filiform ;
anthers globose terminal. Ovary cylindrical oblong; style about as
long as the stamens, stigma concave.
The pods are crowded together, three or four being produced on a
peduncle. They are contorted into a spiral all coiled together, with
five to eleven seeds in each pod. The margins are undulate, and not
thickened, and the walls are not woody and remain always green,
When ripe the pod breaks up into joints which as they fall delisce.
The seeds vary much in size; they are obscurely triangular in
outline or heart-shaped. The larger ones are about two inches long
and broad and an inch thick. The testa is chocolate-brown, dull and
less woody than in H. scandens. The plant is very abundant in Singa-
pore and I have also seen it in Province Wellesley. It usually grows
in loose scrub on the edges of woods, or amcng secondary growth. It
constantly throws up shoots from the roots, and is a troublesome plant
to eradicate. The shoots are of a purple black colour. The flower-
spikes are usually produced immediately after the fall of the leaves,
in December or January, but by the time the flowers are open the plant
is clad again in leaves, The period of flowering, however, is rather
irregular and flowers may often be met with at other seasons. The flowers
are fertilized by Diptera, chiefly Syrphidx. A good proportion of them
possess no pistil, but are entirely male. The seed is dispersed chiefly
by monkeys which eat portions of the pods, and throw the seeds about.
The plant is called Akar Beluru by the Malays, as is also H. Pur-
saetha DC.
KE. pursAETHA, DC. described by Dr. Prain (J. A. S. B. Ixvi. 2. 242)
under the name of H. scandens Benth. is a much less common plant in
the Malay Peninsula. JI have only met with it in Pahang. The plant
described and figured by Scheffer under the name Hntada Rumphii (Nat.
Tijds. Ned. Ind. xxxii. t. xvii. xviii. B) seems to me to differ in the
form of the pod only. The foliage of H. Pursaetha seems to vary very
much as does that of H. spiralis, and were it not for the fruit I should
1898. ] H. N. Ridley— New species of Entada. 307
certainly refer the plant from Pahang to H. Rumphit as the fewer large
leaflets quite resemble those of Scheffer’s figure; the pod, however,
exactly resembles one, from the Andamans collected by Dr. Prain and
as also exactly like the figure of H. Pursaetha given by Scheffer.
In examining specimens of H, Pursaetha in the Herbarium of the
Botanic Gardens, Singapore, and these lent me kindly by Dr. Prain,
I note a form in which the rachis of the inflorescence and the petiole
and midrib of the leaf are covered with a rather conspicuous tementum,
reddish in the dry specimen, while the rachis in the ether forms is
much more glabrous, though by no means completely so. These speci-
mens were collected in the Chittagong Hill Tracts (Lister 175), Manipur
(Watt 6726), Silhet (Wallich), and Sikkim (Zhomson), and probably
represent a local form. However, I have seen no fruit.
The speeies from the Indian region and Malay Peninsula then are
HH. Pursaetha DC, and #H. spiralisn.sp. Further east we get LH. Rumphit
Scheff. distinguished by its straight pod with oblong not rounded
joints, with a straight and not indented thickened margin, oblong seeds,
and thinner endocarp. I have alse an evidently distinct species collect-
ed by Dr. Haviland in Sarawak on two occasions of which I have seen
no fruit, which I will deseribe under the name of LH. boerneensis.
H. BORNEENSIS, n, sp. Leaves 4in. long, with 6-7 pairs of leaflets
opposite or in the lower part of the leaf, alternate, oblong retuse, slightly
oblique, coriaceous dark shining above, when dry glaucous beneath,
petiole 4th in. long, pubescent, midrib on both sides of the leaf
pubescent, 1 in. long, and 3ths in. wide; tendrils two on each leaf.
Inflorescence, one foot to one foot and a half long covered with
closely appressed pubescence. Flowers densely crowded, much smaller
than in the other species. Calyx cupulate with very short teeth,
pubescent. Corolla 5-lobed, lobes lanceolate acute. Stamens ten, fila-
ments as in HL. Pursaetha ; anthers eblong. Pistil very rudimentary ;
female flowers and fruit not seen. Borneo, Sarawak, at Penkulu
Ampat (374), Saribas (1563, Haviland).
This species is certainly most nearly allied to EH. spiralis in its
opposite leaflets glaucous beneath, but is very distinct in having from
six to seven pairs, smaller, and pubescent on both sides of the midrib,
in the very muck longer spikes of flowers which are net more than
half the size of those of H. Pursaetha or H. spiralis and the flower spikes
being unisexual. Dr. Haviland notes also that all the specimens are
male. The corolla in this species opens out quite flat starwise, not
merely reflexing its petals as in the ether kinds, which gives it a very
different appearance. It is to be hoped that further explorations in
Sarawak will produce female flowers and fruit.
J. I. 39
~
308 L. de Nicéville — On a small collection of [No. 2,
Ona small collection of Butterflies from Buru in the Moluccas.—By Lionen
pe Nickyitur, F.E.S., C.M.Z.8., &c.
[ Received 25th June; Read 3rd August, 1898. ]
As far as I am aware, no list has been published on the butterflies of
Buru. As local and in especial insular lists of fauna are of particular
interest for distributional and other.reasons, I have thought it advis-
able to prepare the list given below. ‘The material on which it is based
is small, and consists of but few examples of each species. It is not
known to me into whose hands the bulk of the collection has fallen.
The specimens were collected early in 1897 (he visited the island a few
years previously) by Mr. William Doherty, aided by native (Indian)
collectors, and are labelled as from Kayeli. This place is marked
Kajeli in my Dutch maps, and hes to the north-east of the island, and
there is a district, fort, and deep bay of this name. In English the
fort is sometimes written Cayeli. The island of Buru (Boeroe in
Dutch, Bouru in German, Bourou in French) is one of the largest of the
Moluccas or true Spice Islands, and it lies a little south of the equator,
between 38° and 4° S. Lat., and 126°-127:20 E. Long. East of Buru is
the large island of Ceram, with the small Amboina or Ambon group of
islands to the south-west of Ceram again; west of Buru is the very
large island of Celebes. Buru has the Ceram Sea to the north and the
Banda Sea to the south. In shape it is a very regular oval with its
longer axis lying parallel with the equator, it is about 85 miles in
length by 40 in breadth, and has an area of nearly 2,000 square miles.
The northern portion of the island produces the plant from which is
extracted the far-famed Cayaput or Cajaput-oil; and that curious
mammal, the Babirusa, is found in the island as well as in Celehes.
Reference is made below to all such species recorded from Buru of
which I haye been able to find records. In the entomological portion
of ‘ The Voyage of the Astrolabe’ Dr. Boisduval records 23 species from
Bourou. Dr. A. R. Wallace in his Piertde of the Indian and Australian
Regions gives 16 species, and in his Notes on Hastern Butterflies records
one species of the subfamily Elymniine and five of the Nymphaline from
Bouru. ‘Those species not seen by me have an asterisk prefixed to their
names. The present collection consists of 93 species only, of which
J have not seen 29. .
Family NYMPHALIDA.
Subfamily Danainaz.
1}. *Hestia (Nectaria) aza, Boisduval.
Bourou (Boisduval), Bouru (Moore). In my collection from Ternate
and Gilolo, |
1898. } Butterflies from Buru in the Moluccas. 309
2. *Dawnats (Radena) soprina, Boisduval.
Bourou (Boisduval); Recorded only from the Aru Isles and New
Guinea by Moore. | I have it from T'ernate and New Guinea.
3. Danats (Radena) MEGANIRA, Godart.
In my collection from Ceram and Amboina.
4. *Danats (Radena) suvENTA, Cramer.
Bourou (Boisduval). A doubtful record I think.
5. Danais (Nasuma) 1smare, Cramer.
In my collection from Amboina.
6. Danats (Ravadeba) LutEscens, Butler,
__ Described from Ceram and Bouru by Dr. Butler. Recorded also
from Batchian by Moore.
7. *TELLERVO ASSAR(CA, Cramer.
Bourou (Boisduval as Hamadryas assaricus, sic !).
8: *Eurta@a (Vadebra) meuina, Godart.
__ Bourou (Boisduvat). Recorded from Ceram and the Aru Isles by
Moore.
9. Huria@a (Gamatoba) MELANCHOLICA, Butler.
Originally described from Botiru and Amboina. My specimens
agree very well with Mynheer P.C.T. Snellen’s figure of this species
in Tijd. voor Ent., vol. xxxii, p. 381, pl. viii, fig. 2, male (1889), as
E. melancholica, var. ethiops, Butler, described by Butler as a distinct
Species from Waigiou. Sriellen’s specimens were from Roon Island.
My Buru specimens are fairly constant, none of them Have any spots
‘on the upperside, this feature being characteristic of the var. sxthiops.
Unfortunately Dr. Butler did not figure either species. Mr. Moore
keeps them distinct.
10. *EHurtwa (Gamatoba) aucatuok, Godart.
Bourou (Boisduval). Mynheer P. C. T. Snellen in Tijd. voor Ent.,
vol. xxxv, p. 1, n. 1 (1892), has pointed out that the HF. alcathoé of
Moore is not the same species as the H. alcathoé of Godart, the true
E. alcathoé probably being an older name for E. melancholica, Butler.
*‘ Danatis”’ alcathoé was originally described from Amboina.
1l. *Kopta@a (Gamatoba) seicuLirersa, Moore.
Bouru (Moore).
310 L. de Nieéville — On a small collection of [No. 2,
12. Evpnma (Betanga) puroncHettt, Boisduval.
Originally described from Bourow. In my series of specimens
from Buru the spots on the underside vary greatly in size and number,
one extreme has four spots on the forewing and ten on the hindwing,
the other has twenty and thirty-one respectively. I have this species
also from Ceram.
13. *Evrie@a (Chirosa) LAPEYROUSEI, Boisduval.
Described from Vanikoro by Boisduval, recorded from Bouru by
Moore, who has: examined the type specimen in M. Charles Oberthiir’s'
collection.
14, Evur.@a semicrrcuivs, Butler.
Recorded from Bouru, Amboina and Gilolo by Moore. I have ae
females from Buru. They are very richly shot with purple on the
upperside in some lights.
15. *Evrrea (Salpinz) bevcostictos, Gmelin.
Boisduval from Bourou as H. eunice, Godart.
16. Evena@a (Salpinz) pourvans, Moore.
Bouru (Moore). i good series of both sexes. As usual the number
and size of the spots shews much variation. The maue (hitherto unde-
scribed) differs from the female in having the usual secondary sexual
characters of the subgenus Salpine; the UPPERSIDE of the forewing has
the bowed-out portion of the inner margin paler than the rest of the
wing, the blwish-white spot in the swbmedian interspace is smaller,
single and round, in the female it is dowble; the hindwing has the
abdominal margin broadly paler than the rest of the wing, the sub-
marginal series of spots greatly reduced in size or absent altogether ; on
the UNDERSIDE of the forewing the spots on the disc are rich violet-blue, as.
are all the spots on the hindwing, in the female these spots are white.
17. *Kurt@a (Salpine) nismz, Boisduval.
Bourou (Boisduval). Mr. Moore records it from the Aru Islands:
only, while Dr. Boisduval gives it only from Bourou,
18. *Kurpia@a (Stretoploa) watsont, Moore.
Bouru (Moore.
Subfamily Satyrina.
19. Mycatusis (Orsotriana) mEDuS, Fabricius;
Hewitson as Myealesis hesione, Cramer, from Bourv. My specimens
1898. ] Butterflies from Buru in the Moluccas. 3lf
have the discal white line across both wings on the underside very
narrow, becoming obsolete ; all the ocelli well-developed,
20. Mycatests (Calysisme) perseus, Fabricius.
A single quite normal male received, of the wet-season ocellated form.
21. Mycarests (Mydosama) RemuLiA, Cramer.
Bouru (Hewitson and Moore).
22. Mycatesis (Mydosama) sirius, Fabricius,
Described from Amboine, Bourou and Offack by Boisduval as:
Satyrus manipa. Hewitson records it from Bouru as Mycalesis manipa.
Moore as Mydosama manipa from Bouru.
23. Letue arere, Cramer.
Hewitson as Debits europa, Fabricius, var. arete, Cramer, from
Bourou. Mr. Moore in Lep. Ind., vol. i, pp. 256, 270 (1892) says that
EL. arcuata, Butler, from Celebes, is quite distinct from L. areta from Sula
and Amboina. My Buru specimens of both sexes have the upperside
beautifully glossed with plumbeous coloration, which doubtless quickly
fades in cabinet specimens.
24. YprHiMA ASTEROPE, Klug.
The wet-season form, with well-developed ocelli on the wnderside,
only obtained.
25. MeELANITIS LEDA, Linneeus.
Hewitson records this species from Bouru as Cyllo leda. I have
six males, which are remarkably constant. The upperside of both
wings is deep ferruginous, darker on the hindwing’; the ochreous band
within and anterior to the subapical ocelli of the forewing is well-defined ;
there are two ocelli towards the anal angle of the hindwing; the under-
side is richly coloured and is uniform in all the specimens; the ocelli
are large; the forewing has three, the hindwing one, prominent dark
bands crossing the wings.
26. Hreso consrantra, Cramer.
A good series of both sexes of this fine species.
27. Hero amasiwts, Boisduval.
Hewitson as Oyllo amabilisfrom Bouru. One male and two' females
received. The male has two meditm-sized ocelli on the underside of
the forewing at the apex, the lower twice as large as the upper; the
hindwing has first a very large, then two minute, then three nearly
312 L. de Nicéville — On a small collection of [ No. 2,
equal-sized and rather large ocelli. One female has on the underside
of the forewing at the apex one medium-sized ocellus, in the other
specimen it is wanting; the hindwing has the ocelli as in the male,
except that the one in the discoidal interspace is wanting in one specimen,
and in the other that that ocellus and the one in the lower subcostak
interspace also is absent.
Subfamily Eiymnuna.
28. Enymntas (Dyctis) vimtnALis, Wallace.
Originally described from Bouru.
Subfamily NYMPHALINA.
29. CurHa LAMPETIA, Linnseus.
Both sexés received.
80. ATELLA EGISTA, Cramer.
Recorded from Bouru by Wallace. ©
31. CyYNnTHIA ARSINO#, Cramer.
Males only received.
32. Precis HEDONIA, Linnezus.
A single very brightly-marked male received, with all the ocelli
large and conspicuous, as in Blanchard’s figure of ‘ Vanessa hedonia,
Fabricius,” from Banda, in the Voy. Pole Sud, pl. ii, fig. 10 (1853).
33. JUNONIA ERIGONE, Cramer.
Males only. Cramer’s figure of this species with its bright red
markings on the upperside is very bad.
34. Nertis (Phedyma) HELIODORA, Cramer. 3
“« Papilio” heliodora is the type of Felder’s genus Phedyma. My
single female from Buru has the inner edge of the discal white band
on the underside of the hindwing quite straight and even, while
Cramer’s figure of the species, probably also taken from a female,
shews the band highly irregular and uneven. Unfortunately I have no
specimens from Amboyna, from whence P. heliodora was described,
tocompare with my Buruexample. Mr. W. F. Kirby in the Supplement .
to his Syn. Cat. Diurn. Lep., p. 742, n. 30 (1877), gives N. heliodora
as a synonym of “Papilio” pellucidus, Goeze, but both species were
described in the same year (1779), and as Cramer figured his species,
1898. | Butterflies from Buru in the Moluccas. 313
I prefer te use his name. Herr C. Ribbe in Iris, vol. ii, p. 237, n. 73
(1889), alters the spelling to ‘ pellucides.” Goeze’s work is in none
of the Calcutta libraries.
35. Nepris veNnrLia, Linneus.
Boisduval as “ Limenitis’’ venilia from Bourou. My female from
Buru differs from males from the same island in having all the white
markings on the upperside larger and more conspicuous, and the blue
coloration reduced. Dr. Staudinger’s ‘ Athyma” venilia, var. (ab. ?)
evanescens, from Batjan, which he has kindly sent me, appears to be a
very good local race, with all the white markings much reduced, vide Ex.
Schmett., vol. i, p. 147, pl. li, male (1886).
36. HyYPoLIMNAs BOLINA, Linneeus.
Boisduval as “ Diadema” lasinassa, Cramer, from Bourou. Wallace
records it from every island in the Archipelago. Buru males are
normal. I have three distinct forms of the female from Buru :—I,
the upperside almost uniformly fuscous with no discal macular white
or violet band between the second median nervule and the costa on
the forewing; II, as in I, but with a deep orange patch in the submedian
interspace of the forewing ; III, as in I, but with the above-mentioned
macular band, which is sometimes white, sometimes violet, and the
outer half of the hindwing more or less bright brown ; no orange patch.
None of these forms have been named and figured by Cramer, though
Ii approaches his “ Papilio” antigone, aud III his “ P.” melita.
37. HyYPoLIMNAS ALIMENA, Linneus.
Boisduval as ‘“‘ Diadema” alimena from Bourou. Wallace as
* D.” alimena from Bouru. Males only received.
38. Hypotimnas panpora, Wallace.
The female only is described from Bouru by Wallace. The male has
a large discal yiolet patch on the upperside of the forewing divided into
four portions by the black veins, the posteriormost portion in the
submedian interspace is small, the two portions in the median inter-
spaces are large, the anteriormost portion in the lower discoidal inter-
space as small as the first-mentioned portion; except for this violet
patch the forewing is unmarked; the hindwing is like that of the
female, except that the discal series of oval black spots are blind, in
the female they are pupilled with violet. The hindwing differs from
that of H. pandarus, Linneus, in entirely lacking the large discal
violet patch of that species, the orange area being consequently much
larger. ‘The underside differs but slightly from that of the female ; in
314 L. de Nicéville — On a small eotlection of [No. 2,
the forewing the outer-discal series of spots is incomplete, those in the
upper median and lower discoidal interspaces being absent, and the spots
are blue instead of white. Boisduval records this species from Bourou
as “ Diadema” pipleis, Linneus, which is the female of Hypolimnas
pandarus, Linneus, restricted to Amboyna and Ceram by Wallace.
39. HypoLIMNAS ANTILOPE, Cramer.
Wallace from Bouru. I have a good series of both sexes from
Buru. The typical form figured by Cramer has a broad submarginal
pale ochreous-white band on the upperside of the hindwing, in some
specimens this band is of a deeper colour, being quite ochreous, in other
examples the band is ebsolete, the hindwing being almost uniformly
coloured. In some specimens the submarginal series of small white
spots on the upperside of the forewing is obsolete towards the apex,
in others the two anteriormost spots are large and wedge-shaped. On
the underside the same variations occur as on the upperside. This
species is quite distinct from H. anomala, Wallace, from the Malay
Peninsula, Sumatra, Nias, Java, Bali, Lombok, Celebes, and the Philip-
pines. The Sambawa local race has been named H. swmbawana by
Dr, Pagenstecher in Ent. Nach., vol. xxiv, p. 81 (1898).
40. *ParrHenos Noprica, Boisduval.
Boisduval records this species from Boureu and New Guinea as
“ Ménetra”’ nodrica. Ihave it only from the latter island; its occur-
rence in Buru is I believe more than doubtful.
Al, *Hurngaria (Lextas) mropus, Linneus.
Recorded from Bourou by Boisduval as Lexias xropus, doubtless
correctly.
42. SYMBRENTHIA HIPPOCLUS, Cramer.
This species was originally described from Amboyna, and is record-
ed from the Moluccas by Wallace. Cramer’s figure is not very good, as
it shews the outer-discal series of spots on the underside of both wings,
but especially on the hindwing, white instead of pale violet. The
female in Buru is yellow.
43. SALAMIS SABINA, Cramer.
Both sexes received, apparently common in Buru.
44. CyRESTIS THYONNEUS, Cramer.
Boisduval as Oyrestis thyoneus [sic!] from Bourou.
1898. ] Butterflies from Buru in the Moluccas. 310
45. DOLESCHALLIA CRAMERI, Distant.
Mr. W. L. Distant has so renamed Cramer’s figure of “‘ Papilio”
polibete from Amboyna, Ex. Lep., vol. iii, pl. cexxxv, figs. C, D, male
(1779). My Buru specimens agree very well with this figure, except
that the spots on the upperside of the hindwing are black, the posterior
one sometimes with a white pupil, instead of blue as figured.
Family LYCAINIDA.
46. GERYDUS LEOS, Guérin.
Both sexes of this species are described, and the female is figured,
from Bouron, as “‘ Simeethus” leos. On the plate the generic name is
written “ Symethis.” I have one male only from Buru, but both sexes
from South Celebes. It seems to be quite a distinct species, and has
been described by Doherty from Sumba and Sambawaas Clerydus teos
in Journ. A. S. B., vol, lx, pt. 2, p. 185, n. 92 (1891).
47. GERYDUS CERAMENSIS, Ribbe.
Miletus chinensis, var. ceramensis, Ribbe, Iris, vol. ii, p. 247, n. 95, pl. v (nec 1),
fig. 2, female (1889); Gerydus boisduvaliit [sic!], Moore, var. acragas, Doherty,
Journ. A. S. B., vol. lx, pt. 2, p. 186, n. 93 (1891).
Described by Herr C. Ribbe from South and East Celebes, Ambon,
Saigun, Buru and Borneo. I have both sexes from Buru. The female
agrees exactly with the figure of G. ceramensis, both sexes with the
description of G. acragas, so there is but little doubt that the two species
are synonymous. G. acragas was described from Sumba and Sambawa,
and I have recorded it under this name from the Ké Isles, vide p. 263,
n. 44, ante.
48. PitTHEcoPs DIONISIUS, Boisduval.
Common in Buru. The late Herr C. Ribbe recorded it in Iris,
vol. ii, p. 250, n. 105 (1889) from Great Ceram as Plebejus [sic!]
dionysius [sic!]; Dr. A. Pagenstecher in Jahr. des Nass. Ver. fir
Natur., vol. xxxvii, p. 192 (1884) from Amboina as Cupido dionisus
[sic !].
49, LAMPIDES ARATUS, Cramer.
Buru specimens are quite typical.
50. Lampipus evpis, Godart.
Males only received.
J. i. 40
316 L. de Nicéville — On a small collection of [No. 2,
51. Lamprpes Hybas, Cramer.
Herr Ribbe recorded it (1. ¢., p. 249, n. 102) from Great Ceram as
Plebejus euchylas, Hiibner, but Cramer’s name is the older.
52. ARRHOPALA HELIUS, Cramer.
Males only received.
53. ARRHOPALA FULLA, Hewitson.
Described from Boirou [sic!]. Males only received.
Family PAPILIONIDA.
Subfamily Prerina.
54. *TEerRras ZORAIDE, Felder.
Dr. A. G. Butler in Ann. and Mag. of Nat. Hist., seventh series,
vol. i, p. 59, n. 4 (1898), says that 7. zoraide “ Ranges from Bourou
southwards to Australia.” I have not received this species from Buru,
but Australian specimens appear to me to be quite the same as
T. libythea, Fabricius, which is much the older. Wallace records T. drona,
Horsfield, from Bouru, that species being a synonym of T. libythea.
Wallace’s 7. drona from Buru probably now stands in the collection of
the British Museum as 1’. zorazde.
55. TERIAS CANDIDA, Cramer.
Recorded by Wallace from Bouru, and by Dr. Butler (1. c., p. 61,
n. 11) from Amboyna and Ceram. I have females only from Buru,
which agree absolutely with females from the Ké Isles.
56. TeRIAS BIFORMIS, Butler.
Originally described from both sexes from Amboina. I have three
females only from Buru, which are creamy-white on both surfaces. It
is described and figured by Mr. Distant from Singapore as 1’. lacteola,
but the Malay Peninsula is not given as one of the habitats of the
species by Dr. Butler in his latest revision of the genus. Such white
females occur sporadically in India, and in my opinion are only occa-
sional “‘ sports” of J’. hecabe, Linnzeus. As, however, I have no yellow
females from Buru, I retain Dr. Butler’s name for the species, which he
says “ Ranges from Nias through Borneo eastwards to Ternate and
Batchian, extending south to Amboyna and Ceram, and _ probably
crossing New Guinea, to reappear in the Louisiade and Solomon
groups” (Ann. and Mag. of Nat. Hist., seventh series, vol. i, p. 76,
n. 53 (1898). The male as described is “bright lemon-yellow ”’
coloured. Wallace probably recorded this species from Bouru as
1898: | Butterflies from Buru in the Moluccas. 317
T. blanda, Boisduval, which is given by Butler as a synonym of T’.. hecabe,
Linneus.
Wallace describes 7. diversa from Bouru and six other localities.
Butler (1. ¢., p. 74, n. 50) restricts it,to the Philippines.
Dr. Butler (I. c, p. 76, n. 52) says that 7. brevicostalis, Butler,
is probably found in Bourou, but I have not received it from thence.
57. CATOPSILIA CROCALE, Cramer. |
Recorded by Wallace in Trans. Ent. Soc. Lond., third series,
vol. iv, p. 413 (1867) from Bouru as “ Callidryas” alemeone, Cramer.
One very heavily-marked female only received, which, however,
is not as darkly coloured as Dr. Butler’s figure of the same sex of his
“ Callidryas” flava (Lep. Ex., p. 23, n. 2, pl. ix, fig. 5, female (1869).
Dr. Butler does not say whence the specimen he figured came, but he
gives Celebes (Macassar), Ceram and Batchian for C. flava. I quite
agree with his remark ‘“‘ This species may eventually turn out to be an
extreme form of C. crocale.”
58. *HErBomMoIA LEUCOGYNIA, Wallace.
Described from Bouru by Wallace as “‘ Iphias” leucogynia, and
apparently confined to that island. Dr. Adolf Fritze discusses this
species in his interesting paper on the genus Hebomova (Zool. Jahr.,
vol. xi, p. 278 (1898).
59. *ELODINA BOURUENSIS, Wallace.
Originally described from Bouru.
60. *HuPHINA RACHEL, Boisduval.
Recorded from Bouru by Wallace as “ Pieris” rachel.
61. *Huputna saet, Wallace.
Described from Bouru by Wallace as “ Pieris” jael.
Boisduval describes a ‘“‘ Pieris” theodice from Bouron, but this locality
is probably incorrect. Mr. Kirby records it from Chili and Peru.
62. *Appras apa, Cramer.
Recorded from Bouru by Wallace as “ Tachyris” ada.
63. *APPIAS ALBINA, Boisduval.
Males and white females (Form I) recorded from Bouru by
Wallace as “ Tachyris” paulina, Cramer, the latter being found only in
Ceylon.
318 L. de Nicéville — On a small collection of [No. 2,
64. *Appras LEIS, Hiibner.
Recorded from Bouru by Wallace as “ Tachyris” jacquinotit, Lucas,
which species is, in my opinion, a synonym of A. leis.
65. *Appras cynisca, Wallace. |
Described by Wallace from Bouru from the female sex only as
“ Tachyres” cynisca.
66. *APPIAS BOURUENSIS, Wallace.
Described from a unique female from Bouru by Wallace as
“ Tachyris’’ bowruensis.
67. Derias PHILOTIS, Wallace.
Both sexes described from Bouru by Wallace. My male specimens
are a little variable, in some there is a small white spot at the posterior
end of the discoidal cell of the forewing on the underside, in others it
is absent. This is probably the species Boisduval records from Bourou
as “ Pieris” phillyra [sic! ], Godart, which equals D. ceeneus, Linneus.
68. Detias EcHO, Wallace.
Described from Bourou by Wallace.
Subfamily PapiLionina.
69. TROIDES OBLONGOMACULATUS BOURUENSIS, Wallace.
Males only received. This must be the species Boisduval recorded
from Bourou as Ornithoptera hellen.
70. . ParrLto PoLypDoRuS, Linnzeus.
Apparently the commonest species of the genus in Buru.
71. Papinio ruscus, Goeze.
A variable species even in a comparatively small island like Buru.
Herr Ribbe records it from Buru under its synonym P. cinereomaculatus,
Goeze.
72. *PAPILIO GAMBRISIUS, Cramer.
Recorded from Buru by Wallace, Oberthiir and Rothschild.
73. PaPpiLio perPHOBUS, Linneeus.
Apparently not rare in Buru, from whence it has been recorded by
many authors.
74, *PAPILIO DEIPHONTSS, Felder.
The Hon, Walter Rothschild draws especial attention to the fact
1898. | Butterflies from Buru in the Moluccas. 319
‘that Mr. Doherty on his first visit to Buru obtained both P. deiphobus,
Linneus, and P. deiphontes on that island (Nov. Zool., vol. ii, p. 326
(1895). I have only received three males of the former species, all of
which have long tails; the latter species, which I have only from
Ternate and Halmahera, has no distinct tail, only a tooth.
75. *PAPILIO POLYTES ALPHENOR, Cramer.
Not received by me.
76. *PapiLfo EURYPYLUS, Linneus.
Not received by me.
77. *PAPILIO SAKPEDON ANTHEDON, Felder.
Recorded from Bourou by Boisduval as P. sarpedon, Linnzus.
78. PAPILIO AGAMEMNON PLISTHENES, Felder.
Females only received. All the species in this subfamily given
above have been recorded from Buru by the Hon. Walter Rothschild.
Family HESPERIIDA.,
79. TaGIaDES MEeTANGA, Ribbe.
T. neira, Plétz, var. metanga, Ribbe, Iris, vol. ii, p. 265, n. 143, pl. v, fig. 8 (1889).
I do not know T. neira, Pidtz, which was described from the Aru
islands. The var. metanga was described from Great Ceram. My two
females agree very well with the figure of the latter.
80. TAGIADES JAPETUS, Cramer.
Both sexes of this widely-distributed species received.
81. NorocrypTa FEISTHAMELII, Boisduval.
Described as “ Hesperia” feisthameliz, Boisduval, from Amboina
and Bourou. In the text of ‘‘ Voyage de |’Astrolabe,”’ Entomologie,
part 1, p. 159 (1832), the reference to the plate and figure is incorrect,
it should be plate iii instead of ii, and figure 6 instead of 7. The late
Herr C. Ribbe in Ivis, vol. 11, p. 263, n. 142, pl. v, fig. 7 (1889), describes
and figures what appears to bea female of this species from Great Ceram
as “ Plesioneura”’ varians. My females from Buru agree exactly with
Herr Ribbe’s figure. The “ Plesioneura” chimera of Plotz, Berl, Ent.
Zeitsch., vol. xxvi, p. 262, n. 4 (1882), described from “ India,” is also
a synonym. Herr G. Weymer has kindly sent me a coloured drawing
of # specimen of P. chimera identified by the late Herr Plotz himself.
On the underside of the forewing as drawn the discal white fascia ends
anteriorly at the subcostal nervure as in N. restricta, Moore.
320 L. de Nicéville — On a small collection of [ No. 2,
82. TrnicoTa BAMBUSm, Moore.
The male specimens from Buru have the sexual brand on the
upperside of the forewing very prominent, whitish and shining. The
late Herr C. Ribbe sent me a female of this species from Ceram named
“ Pamphila ” ahrendtt, Plétz, and under this name the species stands
in Iris, vol. ii, p. 261, n. 136 (1889). It was originally described as
“ Hesperia” ahrendti from Manila in the Philippines, but is not
mentioned in Herr Georg Semper’s “ Schmetterlinge Philippinischen
Inseln.” On 1. c. pl. v, fig. 5, Herr Ribbe figures a Pamphila arendtr
[sic], but this figure does not in the least agree with 7. bambuse.
83. Tritcota auGiAs, Linneeus.
Males only received. The discal oblique black fascia bearing the
sexual brand on the upperside of the forewing is unusually narrow in
these specimens, consequently the golden-yellow band beyond is un-
usually broad.
84, Trxicota (Padraona) prusias, Felder.
Females only received, which agree exactly with specimens of the
same sex in my collection from the Philippines and Celebes.
85. Trxicota (Padraona) patmarum, Moore.
Males only received. These specimens do not agree with Felder’s
figure of “ Pamphila”’ augiades, male, from Amboina. That species,
which I have never succeeded in obtaining, is very closely allied to
T. palmarum.
86. OcyYBADISTES MARNAS, Felder.
My male specimens from Buru and Waigiou (Waijiu or Waigeu)
agree very well with Messrs. Elwes and Edwards’ description of “ Pam-
phila” marnas, Felder (described originally from Amboina), in Trans.
Zool. Soc. Lond., Zoology, vol. xiv, p. 256 (1897). Messrs. Elwes and
Edwards ignore Mr. Heron’s genus Ocybadistes, although it was described
in 1894, and place the present species in the genus Telicota, though Capt.
Watson pointed out in 1893 that P. marnas and some other species
were not true Telicotas.
87. Baoris (Chapra) MatHias, Fabricius.
My male specimens from Buru have no translucent spots whatever
in the forewing, though the usual discal spots appeur on the underside
but are opaque, while the females are normal.
1898. | Butterflies from Buru in the Moluccas. o21
88. Baoris (Parnara) puinippina, Herrich-Schaffer.
Both sexes received. It seems to be common in Buru.
89. Baorts (Parnara) conticua, Mabille.
Two males of this ‘variable and widely-spread species from Buru
have been received. In both examples the two spots in the discoidal
cell of the forewing are well separated, but this is a very inconstant
feature ; in one specimen there is a single spot, in the other two spots
only, on the disc of the hindwing, the normal number being four.
90. Hasora (Parata) cELZNuUs, Cramer.
Originally described from what appears to be a male (though the
sexual brand is not figured) from Amboyna. I have both sexes from
Buru. The female shews in a good light on the upperside of both
wings a very magnificent plumbeous gloss, which is entirely wanting
in the male. The coloration of both wings on the underside is in the
female of a much paler and lighter shade than in the male, and green
instead of purple. Hasora violacea, Klwes and HKdwards, Trans. Zool.
Soc. Lond., vol. xiv, pp. 297, 299 (1897), from Amboina, is said to be
allied to H. celeenus, but the male differs from that species as identified
by me in having no sexual brand.
91. Hasora (Parata) MaLayana, Felder.
Two females only received from Buru. Like examples from the
Ké Isles, these Buru specimens are without spots on the forewing.
92. Hasora (Parata) HuRAMA, Butler.
Both sexes received from Buru, which differ the one fran the other
only in the presence in the male of the sexual brand on the upperside
of the forewing, this being absent in the female.
93. *Hasora THRIDAS, Boisduval.
Described from Bourou as ‘“ Thyméle” thridas by Boisduval. As far
as Boisduval’s short description goes it agrees with the specimens [|
have identified as H. malayana, Felder. But Watson says in Proc.
Zool. Soc. Lond., 1893, p. 128, that H. thridas comes into group A of
the genus in which the sexual brand in the male is either inconspicuous
or entirely absent, while the male of H. malayana has it conspicuous,
and comes into his group B.
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H Pie
INDEX.
Names of New Genera and Species have an asterisk (*) prefixed.
Acacia, 260
ACANTHACEA, 300
Acanthocyclus, 68
Actxa, 73, 74, 121, 187, 143, 146, 147,
_ 153, 154, 166
», acantha, 151
» areolata, 138, 141
» calculosa, 139, 152
»» carcharias, 151
5, cavipes, 189, 147, 148
» ecninus, 189, 149
* ,, flosculata, 139, 151
» fossulata, 139, 148
» granulata, 139, 151, 152, 153
» hirsutissima, 138, 141
» hystrix, 151
», nodosa, 142
», nodulosa, 139, 148, 149, 150
53 var. bullifera, 149
a obesa, 139, 145, 146
5» parvula, 139, 146
A peronii, 139, 150
», pulchella, 139, 146
» pura, 151
» reéfopunctata, 138, 142, 143, 144,
149
» rugata, 144
» ruppellii, 1389, 144, 145, 146, 153
» Savignit, 152
ts schmarde, 148
» speciosa, 138, 143
tomentosa, 138, 140, 141, 142
Act xiNa, 70, 78, 137
Actzxodes, 137, 154
nd cavipes, 147
_ modestus, 146
os nodipes, 143
fA speciosus, 143
tomentosus, 140
Actumnus, 75, 190, 198, 200, 206, 207
an arbutum, 201, 205
e elegans, 201, 206
~ 7, jesstfrons, 201, 204
3 nudus, 68, 201, 207
A setifer, 201, 202
ry iy, tessellatus, 201, 205, 206
a tomentosus, 201, 202, 293
i verrucosus, 68, 201, 298
Aechmandra Blumeana, 29
e indica, 39
Aegle aeneus, 104
» granulosus, 101
» rugata, 144
» ruppellit, 144
een ynentnirs gracilis, 299
3 grandiflora, var. longiflora
299
* a micrantha, var. Pottingert
299
* pusilla, 299
* Agalma Lawranceanum, 293
» redivivum, 51
Be gts 296, 297, 298
Desmogyne, 298
<oe Pottingeri, 296, 298
rine chia Faberi, 295
Ee Kingianum, 294
Fe 262
Aeoniira, 25, 42
a Capricornica, 42
Me cissoides, 40
ij clavigera, 42, 292
; lava, 40
dh ay? pubigera, 292
Amtlypodia, 269
¥ micale, 270
Ameletia indica, 3
Ammannie, 2
es acutidens,
si baccifera, 3
x debilis, 4
PP elongata, 3
9 indica, 4
= latifolia, 3
a nana, 3
. peploides, 3
. polystachya, 3
ie repens, 3
ie verticillata, 4
vesicatoria, 3
AMMANNLA, 2
epror phage tts, 305
Cruddasianus, 305
meee maro, 277
Ancylovipha agraulia, 281
Ancyloxypha, 281
Anona muricata, 277
ANONACKA, 284
Antidesma, 19
Apaustus, 281
Appias ada, 317
albina, 273, 317
bowruensis, 318
celestina, 272
clavis, 273
cynisca, 318
leis, 318
lyncida, 272
Arachnecthra asiatica, 64, 65
a zelonica, 64, 65
Aralia, 43
armata, Ad
digitata, 52
dubia, 58
ferox, 44, 45
Thomsoni, 44
ARATIACES, 1, 42, 45, 59, 293
Aralidium, 43, 45
5 pinnatifidum, 45
Araliopsis andamanica, 62
Arisarum amboinicum, 302, 308
Aristolochia, 273, 274
AROIDEA, 301, 302, 305
Arrhopala amytis, 270
fulla, 316
: helius, 270, 316
Arthrophyllum, 43, 59, 60
Blumeanum, 59
diversifolium, 59
ellipticwm, 59
javanicum, 59
ovalifolium, 59
pinnatum, 59
Arum or arense, 302, 304
trilobatum, 302, 303, 3804
Asthipa citrina, 255
Atelecyclus, 69
Atella egista, 258, 312
Atergatis, 71, 89, 94, 98, 99, 104
anaglyptus, 102
asperimanus, 98
dilatatus, 95, 96
ie elegans, 108
floridus, 95, 98
integerrimus, 95, 96, 97, 98
levigatus, 97
lateralis, 108
limbatus, 101
marginatus, 97
roseus, 95, 97
scrobiculatus, 97
semigranosus, 101
subdentatus, 96
subdivisus, 96
Athyma lactaria, 260
venilia, 261
9 var. evanescens, 313
Meiictik caseolaris, 11
AURANTIACE®, 275
Badamia exclumationis, 283
Banareia, 73, 153
32 |, Index.
Banareia armata, 153
Baoris contigua, 321
larika, 282
mathias, 282, 320
philippina, 282, 321
(Chapra) mathias, 282,320 »
(Parnara) contigua, 321
larika, 282
philippina, 282, 321
* Baptozius, 76, 186, 187, 188
me vinosus, 189
Bauhinia excelsa, 290
» nervosa, 290
* , Pottingeri, 289
BEGONIACEA, 1
Benincasa, 25, 34
he cerifera, 35
* hispida, 34
Betanga duponchelii, 310
Bindahara isabella, 270
bP] ”
- Blackwellia, 22
caryophyllacea, 22
dasyantha, 22
longiflora, 22
macrostachya, 20
propinqua, 21
, spiralis, 21
Brassaiopsis, 43, 61
conflwens, 58
Hainla, 62
plamata, 61
3”?
” 29
62
Bryonia Blumei, 39
cheirophylla, 34
cochinchinensis, 32
» epigza, 39
geminata, 39
grandis, 32
heterophylla, 39
marginatu, 39
palmata, 29
tenella, 39
umbellata, 27
Callidryas alcmeone, 317
frava, 271, 317
Callslonn hopfferi, 256, 257
3 visenda, 256, 257
CALYCIFLORA, lL
Calysisme perseus, 257, 311
Cancer, 67, 69, 70
» adspersus, 80
5» weneus, 104
» amphitrite, 104
» anaglyptus, 133
» bispinosus, 1386
» calculosus, 152
3 converus, 80
»» cymodoce, 219
» daira, 155 —
» dentatus, 129
var. andamanica,
Cancer distinguendus, 113, 123
dodone, 108
electra, 133
exsculptus, 110
floridus, 98, 104
fossulatus, 148
glaberrimus, 223
granulatus, 151
huoni, 111
impressus, 115
inequalis, 119
incomparibilis, 104
integerrimus, 95
limbatus, 101
maculatus, 79
mamillatus, 110
marginatus, 97
marmarinus, 80
melissa, 110
metis, 183
niger, 160
ochtodes, 135
ocyroe, 98
perlatus, 155
polydora, 150
roseus, 97
ruber, 79
rubromaculatus, 93
rufopunctatus, 222
rumphii, 106, 178
savignyt, 151
sazatile, 79
saaatilis, 106
tenaz, 187
variolosus, 155
vespertilio, 192
(Actza) granulatus, 151
(Galene) bispinosus, 136
(Liagore) rubromaculatus, 93
(Xantho) distinguendus, 118, 123
CANCERIDES, 69
CANCERIENS, 69
”?
ARQUEs, 69
QUADRILATERES, 69
Cancripa, 67, 69, 70
CANCRINA, 69
CANCROIDEA, 68, 232
CYCLOMETOPA, 68
PoRTUNINEA, 68
Candulides, 265
CaRPILioipa, 77, 78
Carpilius, 72, 78, 81, 90, 152
cinctimanus, 88
convexus, 79, 80
lividus, 80
maculatus, 79
marginatus, 97
roseus, 97
venosus, 83
Carpilodes, 72, 81, 85
* »”
cariosus, 82, 86
Index.
Cargnedes cinctimanus, 88
lophopus, 81, 84
329
margaritatus, 67, 82, 85, 86
monticulosus, 82, 86
pediger, 81, 83
ruber, 83
rugatus, 82, 84, 85
stimpsoni, 81, 82, 83
tristis, 81, 82, 83, 85
vaillantianus, 82, 85
venosus, 67, 81, 83
Carpilow anthus, 81
9
rugatus, 84
vaillantianus, 85
Casearia, 14
*%
albicans, 15, 16, 18
andamanica, 15, 16
Championii, 17
cinerea, 16
Clarkei, 15, 18
esculenta, 15, 16
grewrefolia, 15, 16
Kunstleri, 15, 17
levigata, 17
Lobbiana, 15
macrocarpa, 15, 16, 18
subcuneata, 16
variabilis, 16
varians, 17
Zeylanica, 17
CaTAMETOPEs, 69
Catappa, 292
Catochrysops cnejus, 269
”»
bP]
lithargyria, 269
strabo, 269
Catopsilia alemeone, 271
99
crocale, 271, 283, 317
au
» ab. flava,
pomona, 271
Cerasiocarpum Maingayt, 39
penangense, 39
Cethosia cydalima, 258, 283
cydippe, var. damasippe, 258
insulata, 258
lamarckii, 259
Chanapa sacerdos 256, 257
Chapra agna, 282
mathias, 282, 320
Charaxes keianus, 262. 283
99
pyrrhus keianus, 262
Cheirantha, 58
Chirosa, 255
lapeyrousei, 310
CHLORODINA#, 70, 78, 156
CuioropiorpA, 78, 159
Chlorodius, 74, 159, 161, 162, 165, 172
areolatus, 162, 166
cavipes, 122
cytherea, 160
dentifrons, 133
depressus, 160
326
Chlorodius distinguendus, 1138
o edwardsi, 120
” eudorus, 122
“ exaratus, 118
- fragifer, 151
ai hirtipes, 160
= levissimus, 160, 161
ss monticulosus, 163
Bs nebulosus, 160
- niger, 159, 160, 161
a nodosus, 120
a nudipes, 121
& obscurus, 163
As perlatus, 166
- polyacanthus, 150
i samoensis, 134
“ sanguineus, 119
s sculptus, 164
ungulatus, 162
Chlorodopsis, 74, 159, 165, 171, 172
6s areolata, 165, 166, 172
Es hy OES 166, 168
* melanodactylus, 171
th nigrocrinita, 166, 168
5 ornata, 171
% pilumnoides, 165, 167, 168,
169
! spinipes, 166, 169, 171 -
* rf wood-masoni, 166, 170
(Cyclodius) ornata, 171}
CINNYRIDA, 64.
Clitoria, 287
Goereba, 66
- cyanea, 64, 65
CorREBIDE, 66
CoLuma, 287
COMBRETACE, 291
CoMMELYNACES, 301
CoNNARACEA, 285
Cordylanthus frutescens, 22
CorRNACES, 1, 45, 294
Corystoidea, 69
*Cruddasia, 287
*
pa insignis, 287
Crypteronia, 2, 4
7 glabra, 5, 6
Sf Griffithii, 5, 6
os paniculata, 5
pubescens, 5, 6
Cr yptotheca apetala, 4
CUCUMERINE®, 24
Cucumis xgyptiacus, 34
» %mtegrifolius, 31
» Missionis, 27
Cucurbita alba, 35
“A farinosa, 35
“4 hispida, 35
. melopepo, 29
- Pepo, 35
- villosa, 35
CUCURBITACEA, 1, 24, 34, 40, 41, 292
Indez.
Cupha crameri, 258
», lampetia, 312
Cupido dionisus, 315
Curetis, 270
» tagalica, 270
Cuscus, 253
Cyaniris, 264
+ cagaya, 264
5 kwehni, 264
“4 puspa, 264
Cyclodius, 74,78, 159, 171
a ornatus, 171
CycLoMETOPA, 68, 69
Cyclowanthus, 74, 112, 124, 125
lineatus, 68, 124
Cyllo annabilis, 257, 311
» eda, 311
Cymo, 74, 78, 172
» andreossyt, 178, 174, 175
var. melanodactylus,
174:
» deplanatus, 173, 174
», melanodactylus, 173, 174
» quadrilobatus, 173, 175, 176
,, tuberculatus, 68, 173, 176
Cyrmoipa, 78, 172
Cynthia arsine, 312
* cycnia, 259
Cyrestis thyoneus, 314
Daira, '71, 154
» perlata, 155
» variolosa, 155 —
Delbergis Benthami, 289
te Kingiana, 289
bs rubiginosa, 289
DANAIN®E, 254, 308
Danais affinis, 255
», alcathoé, 309
» cttrina, 255
» hamata, 254
» imtensa, 255
» wmare, 309
» guventa, 309
» laratensis, 255
» lutescens, 309
» meganira, 309
» petilia, 254
» septentrionis, 254
3 sobrina, 309
» (Asthipa) citrina, 255
», (Limnas) petilia, 254
» (Nasuma) ismare, 309
» (Radena) juventa, 809
fs a meganira, 309
- sobrina, 809
, (Ravadeba ) lutescens, 309
» (Salatura) affinis, 2- 5
is - lavatensis, 255
(Tirumala) hamata, 254
Danaus chrysippus, 254
» glorola, 255
9 99
Index. 327
Danaus plexippus, 255 .
Debis ewropu, var. arete, 311
Delias ceneus, 318
a Cehg; 31S
» philotis, 318
pe ottopanng, 43, 58, 294
at Listeri, 294
: a Maingayi, 58
parviflorum, 58
Dendrophis pictus, 67
*Derris latifolia, 288
» thyrsiflora, 289
» Wallichii, 289
*Desmogyne, 297, 298
* _ ner iifolia. 297
Deudori 1a epijarbas, 270
» simsoni, 270
Diadema minena, 313
» hewitsoni, 261
., lasinassa, 313
= pandarus, 261
w pipleis, 314
pe wncceryys corchorifolia, 300
elatior, 299
a. 287
DipTeRA, 3806
ey porian pullum, 3801
‘ var. oblanceolata, 301
ee atin australis, 262
a bisaltide, 262
ve crameri, 262, 815
4 polibete, 262
be pratipa, 262
Domecius, 229
Domecia, 76, 229
s hispida, 230
Domecioipa, 177, 229
Dryophis mycterizans, 66
Duabanga, 2, 9
Ks grandiflora, 10
sonneratioides, 10
Dyctis melane, 258
» viminalis, 312
Elodina bouruensis, 317
4 egnatia, 271
Elymnias melane, 258
a viminalis, 312
A (Dyctis) melane, 258
Z viminalis, 312
ELYMNrn”, 258, 308, 312
Enkylia digyna, 40
7" trigyna, 40
area, 305
» borneensis, 307
3 pursetha, 305, 306, 307
vs Rumphii, 306, 307
“A scandens, 306
4 spiralis, 305, 307
Epilobium fruticoswm, 12
Epizanthus, 75, 181, 184
“ ‘dentatus, 184, 185
arneanthais dilatatus, 186
aa frontalis, 184
. kotschii, 185
Erina, 265
Eriphia, 76, 213
a gonagra, 216
¥ levimana, 214, 216
- A var. smithii, 216
aa seabricula, 214
3 smithii, 214, 216
trapeziformis, 215
ERIPHIIDS, 67, 69, 70
HripHiin&, 70, 177, 218
WRIPHIOIDA, 177, 213
EscaLLoniga, 291
Etisodes, 73, 77, 112, 182, 171
» anaglyptus, 133, 134
» dentatus, 180
» electra, 133
» sculptilis, 133
Etisus, 72, 112, 128, 182
» anaglyptus, 133
45 convexus, 1382
» dentatus, 129, 131
» levimanus, 129, 131
» macrodactylus, 1382
» maculatus, 182
» rugosus, 133
utilis, 129, 130
Eudora impressus, 118, 115
» tenax, 187
EUPHASEOLER, 287
Eupilumnus websteri, 280
Euplea, 253, 256, 257, 261
is alcathoé, 309
» assimilata, 256, 261, 283
» dbouruana, 310
» callithoé, 256, 257
“9 duponchelit, 310
a ewnice, 310
5 eurypon, 255, 257
» fraterna, 256
a hisme, 310
» hopfferi, 256, 257
be lapeyrousei, 310
» leucostictos, 310
» melancholica, 3809
E " var. zthiops, 309
» melina, 309
» sacerdos, 256, 267
os semicirculus, 310
» spiculifera, 309
» visenda, 256, 257
: watson, 310
», (Betanga) duponchelit, 310
+ (Calliplea) hopferi, 256, 257
ia a visenda, 256, 257
»» | (Chanapa) sacerdos, 256, 257
», (Chirosa) lapeyrousei, 310
3, (Gamatoba) alcathoé, 809
melancholica, 809
oe ??
328
Euplea (Gamatoba) spiculifera, 309
(Hirdapa) assimilata, 256, 283
(Salpinw bowruana, 310
hisme, 310
ms - leucostictos, 310
(Stictoplea) watson, 310
(Vadebra) eurypon, 255, 257
Ht melina, 309
EUPL@INA, 256
Eupsychellus, 263
dionisius, 263
” +P
Eupteron, 59
Eurema hecabe, 272
Euruppellia, 75, 186
annulipes, 188
tenaw, 187, 188
vinosa, 189
Eurycar cinus, 75, 207, 209, 213
grandidieri, 210, 211
maculatus, 68, 210, 211,
212
orientalis, 210, 211, 212
Euthalia exropus, 262, 314
Ht (Lewias) xropus, 262, 314
EvuxantHoipa, 77, 109
Euzxanthus, 72, 109, 111
exsculptus, var. rugosus, 110
huonw, 111
mamillatus, 110
melissa, 110, 111
me nitidus, 110
sculptilis, 110, 111
Bveres argiades, 267
FICOIDEa, 1
Galactia, 287
GALACTIER, 287
Galene, 70, 74, 77, 127, 136, 232
» bispinosa, 136
» ochtodes, 135
GALENINA, 136
GaLENOIDA, 77, 136
Gamatoba alcathoé, 309
+ melancholica, 309
“ spiculifera, 309
Gastonia palmata, 58
Gecko. 67
Gerydus acragas, 263, 315
» boisduvalit, var. acragas, 315
» ceramensis, 315
a). 1208, SiS
» teos, 315
» vincula, 263
GESNERACEA, 299
Gilibertia palmata, 58
Glycine, 287
ke Nall Gardneri, 284
s pedunculari is, 284
Thwaitesii, 284
Grapsillus, ory.
- maculatus, 221
“4 subinteger, 220
”?
”»
Index.
Gymnopetalum, 24, 31
5 cochinchinensis, 31, 32
- heterophyllum, 33
os antegrifolium, 31
5 quinquelobum, 32
¥s quinqueloculare, °
septemlobum, 35
Gynostemma, 25, 40
& cissoides, 40
* lara, 41
> pedata, 40
es pedatum, 293
GYNOsSTEMMEA, 25
H#MODORACEA, 300
Halimede, 71, 124, 125, 134, 135, 136
2 thwrs/ont, 68, 186
HALIMEDOIDA. 77, 134
Hamadryas, 225
Be assaricus, 309
Me nais, 225
sy niverpicta, 225
Hapalocarpum indicum, 4
5 vesicatorium, 4
HAPLOTRICHIUM, 299
Hasora alexis, 283
» celenus, 321
ot chromus, 283
i discolor, 283
3 doleschallii, 282
By hurama, 321
» malayana, 283, 321
R thridus, 321
» vtolacea, 321
» (Parata) celenus, 321
ss + hurama, 321
* malayana, 283, 321
Habomei 317
oa leucogynia, 317
Hedera cheirantha, 58
» ferruginea, 58
» heterophylla, 56
» Jdgackiana, 59
ovata, 59
» palmata, 58
» polycantha, 62
venosa, 52
Hederopsis Maingayi, 62
Henslovia, 6
» . affinis, 5
glabra, 5
Hookeri, 5
5 leptostachys, 5
pubescens, 5
Heptapleurum, 43, 46
3 affine, 47, 54
Ba avene, 46, 47
. biternatum, 47, 56
= capitatum, 49
Cephalotes, 46, 49, 50 |
* ‘% Curtisir, 47, 56
" dvarieata, 54
Iudex. 329
Heptapleurum ellipticum, 47, 49, 53, 54
heterophyllum, 47, 56
- = Hullettii, 47, 54
* _ latifoliolatum, 47, 51
il a Lawranceanum, 293
- longifolium, 54
* a luridum, 46, 48
* e mervosum, 47, 55
= ~ Ridleyi, 47, 54
Me rigidum, 50
5 scandens, 47, 58
* * Scortechinii. 46, 49
# Singalangense, 47, 51
fh subulatum, 47, 51
* a sub-racemosum, 46, 49
be terebinthacea, 52
5 tomentosum, 46, 50
* 3 triste, 46, 48
if venulosum, 47, 52, 58
_ ” var. macro-
phylla, 52
* 3s Wrayi, 47, 55
* eo (Agalma) Lawranceanum,
293
Hesperia ahrendti, 320
feisthamelii, 319
» flavovittata, 281
HesPERUD&, 277, 319
Hestia aza, 308
@Murvillei, 254
* (Nectaria) aza, 308
ae @urvillei, 254
Heter opanaw, 43, 61
es fragrans, 61
Heteropanope, 75, 207, 210
dentatus, 185
eucratoides, 68, 209
indica, 208, 209
* levis, 208, 209
HETEROPANOPIOIDA, 177, 207
Heteropsis, 43, 62
Hipio, 257
amabilis, 311
» constantia, 257, 311
- Hirdapa assimilata, 256, 283
» fraterna, 256
Hodgsonia, 24, 25
heteroclita, 25
- macrocarpa, 26
Ho.LocaLyx, 296
Holochila, 265
- intensa, 265
Homalium, 14, 19, 24
caryophyllaceum, 22
cordylanthus, 22
fetidum, 22
frutescens, 20, 22
grandiflorum, 20, 23
Grifithianum, 20, 21
* Kunstleri, 20
longifoluum, 20, 21
Homalium obovale, 23
< propinquum, 20, 21
* i undulatum, 20, 23
qoplowanthus, 73, 112, 125
. cultripes, 126
* hextit, 126, 127
Huphina jael, 317
E rachel, 273, 317
* Hydrangea Pottingeri, 290
HyperRo.issa, 70, 76, 77, 233
HypeRoM_eristA, 70, 176
Hypocelus, 71, 109, 111
_ rugosus, 68, 111
Hypolimnas, 254
a alimena, 261, 313
“A 9 var. heteromorpha,
261
ws anomala, 314
4 antilope, 314
A bolina, 261, 313
re heteromorpha, 261
a hewitsoni, 254, 257, 261
a pandora, 313
x polymena, 254, 257, 261
subawana, 314.
Hypolyezena danisoides, 270
Hypopeltarium, 69
Imperata arundinacea, 255
Indigofera atropurpurea, 286
a4 nigrescens, 286
Involucraria Wallichiana, 30
e Wallichii, 29
Iphias leucogynia, 317
Tsmene, 283
» doleschalli, 282
» malayana, 283
Ttea, 291
Jambosa aquxa, 270
Jamides astraptes, 268
a bochus, 268
as cephion, 268
a purpurata, 268
Junonia erigone, 259, 312
o expansa, 259
» orithya, 259
» timorensis, 260
» villida, 259
Jussiza, 12
» angustifolia, 12
» Burmanni, 12
», decumbens, 12
» evaltata, 12
» Jruticosa, 12
on longipes, 12
» octophila, 12
» repens, 13
» scabra, 12
» suffruticosa, 12
i villosa, 12
Lachnopodus, 73, 89, 90
+ rodgerst, 67, 89
330 Index.
Lachnopodus tahitensis, 92
Lagenaria dasystemon, 35
Lugerstremia, 2, 6, 9
celebica, 8
floribunda, 6, 7
Flos-Reginz, 6, 7, 8, 9
grandiflora, 10
hexaptera, 8
hypoleuca, 6, 7
macrocarpa, 9
Munchhausia, 9
ovalifolia, 7, 8
Regine, 9
: speciosa, 9
Lagostoma, 154
modosa, 120
in perlata, 165
Lampides xlianus, 268
extherialis, 268
amphissa, 269
aratus, 268, 315
areas, 269
celeno, 268
elpis, 315
hylas, 269, 316
LEGUMINOSA, 264, 268, 269, 286°
LEMONIIDA, 263
LEPIDOPTERA, 254
Leptodius, 74, 112, 117
cavipes, 118, 122
crassimanus, 117, 118, 120,
121, 122
* euglyptus, 118, 121
exaratus, 117, 118, 120
var. sanguineus,
120
nudipes, 118, 121, 122
bs sanguineus, 118,119, 120, 121
Leptosia xiphia, 271
Leptospartion grandiflorum, 10
Lethe arcuata, 311
» arete, 311
Lexias xropus, 262, 314
Liagora, 74, 93
.. rubromaculata, 93
LIBYTHAINA, 263
Libythea antipoda, 263
Liniaceas, 301
Limenitis venilia, 313
Limnas petilia, 254
Liomera, 72, 87, 90
cinctimana, 88
pe lata, 88
as longimana, 90
a punctata, 91
- rodgersti, 89
* sodalis, 88
- iomanthas 74, 90
» asperatus, 91, 92
ss punctatus, O1, 156
timidus, 91
» >P)
Liphyra brassolis, 271
i. major, 271
Lophactea; 71; 99
ws anaglypta, 100, 102
Pak corallina, 100, 102
* cristata, 100, 101, 103
95 fissa, 67, 100, 103
4 granulosa, 100, 101, 103
semigranosa, 100, 101, 103
Lophowanthus, 116
: bellus, var. leucomanus,
113.
o> scaberrimus, 1138, 116
9 », var. baccalipes, 117
> 99 ~Cultripes, 117
Lophozozynius, 73, 94, 106, 125
£ bellus, var. leucomanus, 113
: cristatus, 67, 106, 107
‘ dodone, 106, 108
5 epheliticus, 106
incisus, 106, 107
octodentatus, 106, 107
3 radiatus, 108 '
1" (Lophoxanthus) bellus, var.
leucomanus, 113
Ludwigia, 12, 13
diffusa, 14
fruticulosa, 14
lythroides, 13
a parviflora, 13, 14
o prostrata, 13
Luff, 25, 33
os acutangula, 34.
» egyptiaca, 33
cattu-picinna, 34
», clavata, 34
cylindrica, 34
Gosa, 34
», hederacea, 34
» Parvala, 34
pentandra, 34
Petola, 34
racemosa, 384
Satpatia, 34
» Wightiana, 40
Lycena merens, 265
ER eae emolus, 267
sarees 267
LyYca&NIDs, 263, 266, 315
* Lysimachia evalvis, var. grandifolia, 298
LYTHRACEA, 1
LYTHRE, 2
Lythrum Perpiis: 4,
Maclellandia Grifithiana, 4
MACROZANONIA, 42
Mastersia assamica, 288
* Mastizia ewonymoides, 295
Medzus, 73, 77, 112, 114, 123
si distinguendus, 113
Ks nodosus, 123
Megisba malaya, 264
Index.
Melanitis, 257
" crameri, 257
» gylippa, 258
a leda, 311
MELASTOMACE, 1
Melia, 76, 230
* ,, cestifer, 231
*, pugil, 231
» tessellata, 231
Metiorpa, 177, 230
Melissa, 109
» mamillata, 110
» mitida, 110
Melotheiat 25, 37
a afinis, 38
a indica, 38
i marginata 38, 39
ee = var. heterophylla,
39
Rumphiana, 39
Meninpe, 75, 177, 179
% bellangeri, 178
=f granulosa, 179
m parvulus, 146
A rumphii. 178
» (Myomenippe) granulosa, 179
MeniPPina#, 70, 176, 177.
MENIPPIOIDA, 176, 177
Messaras cramer, 258
Mesua ferrea, 262
Miletus chinensis, var. ceramensis, 263,
315.
Minetra nodrica, 314
Momordica, 25, 35, 39
” zgyptiaca, 34
's anthelmintica, 37
a Charantia, 35, 37
2) .e2 5. Clarkeana, 35
9 Cochinchinensis, 35, 36
- dioica, 37
i humilis, 37
a Luffa, 34
: macropetala, 37
a mista, 37
mn muricata, 37
" Rozburghiana, 37
és subangulata, 35, 36
tubsflora, 32
Morinda bracteata, 252
Munchhausenia speciosa, 9
Muricia Cochinchinensis, 37
Mycalesis daidis, 257
mt hesione, 310
os manipa, 311
a medus, 310
pe perseus, 257, 311
33 remulia, 311
52 sirius, 257, 311
- (Calysisme) perseus, 257, 311
a (Mydosama) remulia, 311
sirius, 257, 311
”? 3)
MyYRTACEm, 1
_Nacaduba, 265
Mycalesis (Orsotrizna) \medus, 310
_ Mydosama manipa, 311
a remulia, 311
a sirius, 257, 311
Myomenippe, 75, 177, 179
* duplicidens, 179
granulosa, 179
a; ancyra, 268
45 ardates, 268
hs atrata, 268
Ss hermus, 267
i laura, 268
3 meiranganus, 267
A nora, 268
¥ perusia, 268
Nasuma ismare, 309
Nectaria aza, 308
ad Vurvillei, 254
Nectopanope, 75, 207, 212
" longipes, 213
Fi rhodobaphes, 213
Neleus, 229 .
» acanthophorus, 230
Nematopyzis fruticulosa, 14
ss prostrata, 14
3 pusilla, 14
Neopithecops, 264 —
re zalmora, 263
Neptis consimilis, 260, 283
» heliodora, 312
» lactaria, 260
» nectens, 261
» venilia, 261, 318
», (Phedyma) heliodora, 312
+s nectens, 261
( Rahinda) consimilis, 260, 283
Meustaniias 288
Nothopanazx pinnatum, 60
Notocrypta feisthamelii, 278, 319
ss restricta, 319
waigensis, 278
NyMPHatipa, 254, 308
NYMPHALINE, 258, 308, 312
Ocybadistes, 320
marnas, 320
Gthra, 67, 68, 69, 70
CETHRINZ, 69
ONAGRACES, 1, 11
*Ophiopogon cordylinoides, 300
dracznoides, 301
*Ophiorrhiza Lawranceana, 295
lurida, 295
Ornithoptera hellen, 318
priamus, 273
ag cane 75, 112, 127
microps, 128
Orsotra ipne medus, 310
Osmelia, 14, 18
a Maingayj, 19
331
332
Oxyrhyncha, 68, 69
Ozun#, 70, 176, 181
Oz1o1pa, 176, 181
Ozius, 75, 181, 186, 187, 188
» annulipes, 188
» frontalis, 185
» rugulosus, 182
», tenaw, 187, 188
», tuberculosus, 182, 183
» (Hur upypellia) annulipes, 188
tenax, 187, 188
Padraona, 279, 280, 281
ye dugiades, 278
cs autoleon, 280
i dara, 279
a mesa, 279
- mesoides, 282
v palmarum, 278, 320
is procles, 278
Me prusias, 320
sunias, 279
* Paederia Cruddasiana, 295
t. foetida, 296
a linearis, 296
ie tomentosa, 296
Pamphila ahrendti, 320
Ge arendti, 320
x augiades, 320
4 augias, 278
a! larika, 282
mesa, 279
a marnas, 320
ie maro, 277
:, moseleyi, 278
sunias, 281
Panaz armatum, 45
» fragrans, 61
» dackianum, 59
» palmatum, 62
» pinnatum, 60
» polycarpum, 59
secunda, 60
Panopeus, 208
* acutidens, 186
3 caystrus, 181
J dentatus, 185
a herbstii, 208
Papilio, 273
4 zgeus keianus, 275, 283
” ”? bP)
275
” > ” ”
ormenus, 275
” ” 9
) ” ”
” ” 9
275
» agamemnon, 277
argynnus, 277
” 9 plisthenes, 319
», albinus thomsonii, 274, 283
”» PP]
var. amaranta,
ab. amanga,
» pandion, 275
» polydorinus,
Index.
blanca, 275
275,
Papilio albinus thomsonui, ab, mordingtoni,
274
5» ambraz epirus, 276
» antigone, 313
» — argynnus, 277
» vbeccarii, 274
», cinereomaculatus, 318
», codrus tocalensis, 276, 283
» erocale, 271
» deiphobus, 276, 318, 319
99 99
», deiphontes, 318
» erinus, 265
5, euchenor obsolescens, 276
» eurypylus, 319
melampus, 277
9 9)
9 9 »
» . fuscus, 318
» gambrisius, 318
» heliodora, 312
» tphigenta, 261
» lampetia, 258
» melampus, 277
» melita, 313
» ormenus, 275
» pellucides, 313
» pellucidus, 312
» polibete, 315
» polydorus, 274, 318
ty Ms thessalia, 274, 283
» polytes alphenor, 319
5» pomona, 271
» sarpedon, 319
anthedon, 319
‘ a choredon, 277
» thomsonit, 274
PaPILIONIDA, 271, 316
PAPILIONINA, 273, 318
Papua, 264
Parata celenus, 321
» hurama, 321
» malayana, 283, 321
Paratropia brachybotrya, 53
- elliptica, 53
- heterophylla, 56
99 » 9%
<5 scandens, 53
¥ Singalangense, 51
5 Singalense, 51
. subulata, 52
if tomentosa, 50
venulosa, 52
Parnara contigua, 321
- larika, 282
philippina, 282, 321
PARTHENOPIDA, 69
Parthenos brunnea, 261
Pa nodrica, 314
salentia, 261
z sylvia, 261
PASSIFLORACEA, 1
ab. hypoxanthos, 276
ab. rufinus,
277
Index. 333
Pemphis, 2, 4 PIRIMELIDEs, 69 —
a acidula, 4 Pithecops, 263
ey angustifolia, 4 s bassaris, 263
« setosa, 4 dionisius, 315
tenance, 293 Platypilumnus, 70, 76, 232
sd stellatum, 293 gracilipes, 233
Pentapterygium, 297, 298 Platypodia, 94
- Peplis indica, 3 Plebeius xlianus, 268
Pestolozzia lata, 40 » cxelius, 267
ie pedata, 40 » cagaya, 264
Phedyma, 312 » euchylas, 269
a heliodora, 312 » tWlustris, 266
nectens, 261 » kuhni, 264
Philiris, 265 » lucifer, 263, 264
% tlias, 265 » seltuttus, 267
Phymodius, 74, 159, 161, 165 » tualensis, 264
ae monticulosus, 162, 163 unicolor, 267
is obscurus, 163 Plebejus astraptes, 268
- sculptus, 156, 159, 162, 164 ” beticus, 269
ungulatus, 162, 163, 164 af dionysius, 263, 315
PIERINA, 271, 316 % euchylas, 316
Pieris jael, 317 ‘a kandarpa, 269
» padusa, 271 Plesioneura chimera, 319
» perictione, 278 * varians, 319
» phillyra, 318 2 waigensis, 278
» pitys, 273 : Polycremnus, 71, 123, 124, 184, 135
» rachel, 273, 317 bs ochtodes, 135
theodice, 317 Polycyma, 265
Pierrea dictyoneura, 24 " carythx, 265
Pilodius, 165 PoLYDECTINA, 69
“s nigrocrinitus, 168 Polydectus, '70
i pilumnoides, 167 Polyommatus beticus, 269
pugil, 170 PortTUNIDa, 69
a scabriculus, val * Pottingeria, 291
spinipes, 169 * ms acuminata, 291
PILUMNINA, 70, 176, 190 Precis expansa, 259
PitumNorpa, 177, 190 », hedonia, 312
Pilumnopeus, 207 » tda, 259
> maculatus, 212 » tphita, 259
Pilwmnus, 75, 190, 198, 199, 206 » zelima, 259
% andersoni, 191, 194 PRIMULACE, 298
:, andreossyi, 173 Psawmis, 137
a cerulescens, 191, 196 if fossulata, 148
BS? cursor, 191, 195 Hs glabra, 143
3 dehaanii, 191, 198 Pseudocarcinus, 177
7 dorsipes, 191, 197 i bellangeri, 178
Ps forskalii, 194, Pseudodipsas, 265
os hirsutus, 191, 197 » eone, 265
» labyrinthicus, 68, 190, 199 0 ilias, 265
rs. levis, 190, 199 PsSEUDOZIOIDA, 176, 180
5 levimanus, 199 Pseudozius, 75, 180
z, longicornis, 191, 198, 194 9 caystrus, 181
z scabriusculus, 206 ” planus, 181
‘a semilanatus, 200 Pueraria, 287, 288
A seminudus, 200 ge bella, 288
re setifer, 202 ” peduncularis, 288
zy sluiteri, 191, 194 ” Thunbergiana, 288
cs ursulus, 192 Punica, 2
ee vespertilio, 191, 192, 197 Pyranga rubra, 66
” 9 var. 193 Quadrella, 76, 217, 225
Pirimela, 67, 69, 70 + boopis, 227
334
Quadir ella coronata, 226, 227
” ”
” ’
Radena juventa,
meganira, 309
». sobrina, 309
Rahinda consimilis, 260, 283
Rapala phranga, 270
Ravadeba lutescens, 309
* Rhinacanthus calcaratus, var. maxima,
300
* " maavmus, 300
Rhizophora caseolaris, 11
RvuBIACER, 295
Ruppellia, 186, 187, 188
annulipes, 188
lata, 187
. tenar, 187
e vinosa, 187, 188, 189
RupPeE.t.ioipa, 176, 186
Rusticus Adolescens santhospilos, 265
Salamis sabina, 262, 314
Salatura affinis, 255
i laratensis, 255
Salping, 256, 310
nt bouruana, 310.
< hisme, 310
a leucostictos, 310
Sambucus 263
Samypace@, 1, 14
SaMYDE&, 16
SATYRINA, 257, 310
Satyrus manipa, 257, 311
SAXIFRAGACER, 290
Sciadophyllum avene, 48
~ ellipticum, 53
scandens, 53
tomentosum, 50
Seotanthus Porteanus, 33
7 tubiflorus, 32
Simethus leos, 315
SoLaANAcEz, 298
gt cite feroz, 298, 299
Es » var. inermis, 298
= Melongena, 299
Solenocaulon, 88
Sonneratia, 2, 10, 11
7 acida, 10, 11
3)
bb] ”
us alba, 10, 11
es evenia, 11
a Griffithii, 11
ee lanceolata, 11
os Mossambicensis, 11
e neglecta, 11
a obovata, 11
7 ovalis, 11
Pa gapat, it
Spatholobus ferrugineus, 287
i gyrocarpus, 287
* a Pottingeri, 286
var. maculosa, 226,
+ , », reticulata, 227
var. Griffithit, 10, 11
Index.
Sphenomerus, 76, 217, 227
trapenioides, 228
Spongodes, 227
* Sterculia cognata, 285
» parvifolia, 285%
m Rozxburghii, 285
fay striatiflora, 285,
STERCULIACES, 285
Stictoplea watsoni, 310
Bre olaon volubile, 301
var. setosa, 30D
a Rae hippoetus, 314
Symethis, 315
Symphedra xropa, 262
SYRPHIDA, 306
| Tachyris ada, 273, 317
24 bouruensis, 318
KS cynisca, 318
» jacquinotii, 318
» byncida, 272
» paulina, 317
* Taeniochlena birmanica, 285
Grifithii, 285
Tagiades japetus, 277, 283, 319
M4 metanga, 319
oe neira, 319
», Var. metanga, 319
Taractrocera, 281
Telicota, 278, 320
4» augias, 278, 320
» bambuse, 278, 320
. dara, 279
» moseleyt, 278
és palmarum, 320
oF prusias, 320
(Padraona) dara, 279
7 Telicota (Padraona) palmarum, 320
is prusias, 320
Tellervo, 255
a assarica, 309:
e zoilus, 255
_ TELPHUSIDA, 68, 69
Terias, 272
5 odiformis, 316
» blanda, 317
» brevicostalis, 817
», candida, 272, 316
» diversa, 317
» drona, 316
» hecabe, 272, 316, 317
» lacteola, 316
» leta, 272
», libythea, 316
»» photophila, 272
» sulphurata, 272
zoraide, 316
Terminalia, 292
* Ae argyrophylla, 291
- chebula, 292
Tetralia, 76, 217, 223
r cavimana, 224
Index. 335
Tetralia glaberrima, 223
heterodactyla, 224
levissima, 224
yy nigrifrons, 224
Thecla ilias, 265
THIID#, 69
Thyméle thridas, 321
Thysonitis apollonius, 266
var. swpous, 265,
»”»
>
? ”
celius, 267
hermes, 266
iilustris, 266
korion, 266, 267
RE gs triopus, 265
Tirumala hamata, 254
Trapa, 11, 12
Trapezia, 76, 217, 223, 225, 226, 229
acutifrons, 222
areolata, 218, 221
BA re var. inermis, 221
ceruba, 219
cymodoce, 218, 219, 220, 221,
222, 223
dentata, 219
digitalis, 218, 222
ferruginea, 218, 220, 221, 223
var. areolata, 221
guttata, 220
ys “4 » wmtermedia, 220
areolata, 221
maculata, 221
by 9
9 ” ”?
fusca, 222
glaberrima, 224
hirtipes, 219
integra, 223
intermedia, 218
leucodactyla, 222
maculata, 218, 221
miniata, 220
reticulata, 221
rufopunctata, 218, 222
var, intermedia,
220
maculata,
221
” 9 ?
serratifrons, 224
subdentata, 220
- tigrina, 221
TRAPEZIOIDA, 177, 217
Trevesia, 43, 57
palmata, 57
SY var. liwm, 59
Trichosanthes, 24, 26
anguina, 29
aspera, 29
bracteata, 29
Celebica, 27, 30
cordata, 29
costata, 32
cucumerina, 26, 27, 32
“a ichosanthes dioica, 26
Fe Fatoa, 32
* grandibracteata, 30
Ht graniiflora, 26
a Hearni, 26, 28
35 herpetospermum, 28
i: heteroclita, 26
hexusperma, 26
7 integrifolia, 32
laciniosu, 27, 29
macrocarpa, 26
multiloha, 30
Re officinalis, 32
palmata, 27, 29
pilosa, 27
gubera, 29, 30
reniformis, 28
tricuspidata, 27, 28
tricuspis, 29
» | Wallichiana, 27, 29
Wawrexi, 27, 30
i ipodanthera cochinehinensis, 32
Troides oblongomaculatus bouruensis,
318
» priamus poseidon, 273, 274
# i, ab. hecuba, 273
Tupidanthus, 43, 63
calyptratus, 63
Typhonium, 302, 303, 304
cuspidatum, 304
* - inopinatum, 301, 302, 303
i Ni Listeri, 334
* H Pottingeri, 304
4 Roxburghit, 302, 303
* Schottii, 303
trilobatum, 302, 303
UMBELLIFERA, 1
VAaccINIACEm, 296, 297
Vadebra, 256
eurypon, 255, 257
fe melina, 309
Vanessa hedonia, 312
Vitis atroviridis, 40
», trichophora, 40
* Wardenia, 43, 60
* ee simplex, 60
XANTHIDE, 67, 69, 70, 71, 77
XantTHINA, 70, 77, 78
XANTHINI, 69
Xantho, 74, 90, 91, 112, 114, 115, 116,
117, 123, 125, 127
affinis, 119
bidentatus, 92, 113, 114
cavipes, 122
crassimanus, 120, 121, 122
deHaanii, 162
distinguendus, 113, 123
* euglyptus, 121
exaratus, 118, 120
var. crassimana, 120
nudipes, 12]
3
” »
”? 9? ”
336
Xantho exaratus, var. sanguinea, 120
” ” typica, 119
floridanus, 1.17
floridus, 112, 114, 116
granosomanus, 157
granulosus, 101
hirsutissimus, 141
impressus, 91, 112, 118, 115
incisus, 107
lamarekii, 157
lamelligera, 108
lividus, 119
macgillivrayt, 113
nitidus, 108
notatus, 158
nudipes, 121, 122
obtusus, 83
octodentatus, 106
peronit, 150
punctatus, 91
quinquedentatus, 122
radiatus, 108
rufopunctatus, 142
sanguineus, 119, 120, 121
scaberrimus, 112,118,116
te var. baccalipes, 117
_ », cultripes, 117
spinosus, 150
superbus, 107
tahitensis, 92
tuberculatus, 114
(Eudora) impressus, 118
(Lachnopodus) tahitensis, 92
(Leptodius) cavipes, 122
ae 9 crassimanus, 120, 121,
” 122
oS rs euglyptus, 121
Index.
Xantho (Leptodius) exaratus, 118, 120
nudipes, 121, 122
sanguineus, 119, 120,
121
»» (Lophoranthus) scaberrimus, 118,
116
2 var.
baccalipes, 117
scaberrimus, var.
cultripes, 117
” 9
) +P]
» ”
Xanthodes, 74, 156
* granosomanus, 157
ss lamarckti, 156, 157
A microps, 128
ss notatus, 156, 157, 158
XANTHODIOIDA, 78, 156
XANTHOIDA, 77, 112
Ypthima arctous, 257
» asterope, 311
ZANONES, 20d
Zanonia, 25, 40, 42
fe cissoides, 40
=) Oe Clarkei, 41
Fr indica, 41, 42
ag lara, 40
an pedata, 40
5 sarcophylla, 42
Zizera, 265
» gaika, 264
n>. 9 0808, 264
ZOZYMOIDA, 77, G4
Zozymus, 72, 94, 99, 108, 106
a aeneus, 104
Ke canaliculatus, 84
* pilosus, 104, 105
Fe rugatus, 84
is tomentosus, 140
=
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Or THE
*
[TLE PAGE AND INDEX
be
1 janet pana
NEW SERIES. VOL. LXVII. CCCLXX.
JOURNAL |
OF THE |
ASIATIC SOCIETY OF BENGAL,
Vol. LXVII. Part-Il,-No. 1.—1898.
EDITED BY
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2 Issued August 5th, 1898,
Materials for a Careinological Fauna of India. No. 3. _
CONTENTS.
Zz
Materials for a Flora of the Malayan Peninsula.—By Gucndee
K.C.1. E., M. B., LL. D., F. R.S., Superintendent of the}
Botanic Garden, Calcutta ... ae Serpe
Note on the Seasonal change of Plumage in the males of the
Honey-sucker (Arachnechthra asiatica) and of an an
American bird (Coereba cyanea.—By F. Fry, B. A, B.
Deputy Superintendent, Indian Musewm ove .
Note on the Long-Snouted Whip-Snake (Dryophis mycterizans)
F. Fiyy, B. A., F. Z.S., Deputy Superintendent, Indian M
Brachyura COyclometopa. Part I. The Family Xanthid
By A. Atcocg, M. B., C. M. Z. 8., Bannan of the
Museum _ pee seo eee see ty ee
Contribution from the Dicwiea Laboratory, Preston
Calcutta. On Double Thiosulphates of Copper and Sodiun
Cuanpra Bausaan Buavurt, B. A., and Jyott BausHan Baap
M.A., Prem Cuanp RoycHanp. ‘Scuonar. Oommunicat 2 by \
Pepuer, F.R.S. _ San iis ia oer ee
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issued regularly every year. From 1894 an additional volume, Part III,
has been issued. |
The Proceedings up to the year 1864, were bound up with the
_ Journal, but since that date have been separately issued every year.
The following is a list of the Asiatic Society’s publications relating
to Physical Science, still in print, which can be obtained at the Society’s
House, No. 57, Park Street, Calcutta, or from the Society’s Agents in
London, Messrs. Luzac & Co., 46, Great Russell Street, W. C.; and from
Mr. Otto Harrassowitz, Leipzig, Germany.
Asiatic Researcues. Vols. VII, Vols. XIII and XVII, and
Vols. XIX and XX @10/ each Rs. 80 O
Proceepines of the Asiatic Society from 1865 to 1869 (incl.) @
/6/ per No. ; and from 1870 to date @ /8/ per No.
JourNAL of the Asiatic Society for 1843 (12), 1844 (12), 1845
(12), 1846 (5), 1847 (12), 1843 (32), 1850 (7), 1851(7), 1857 (6),
1858 (5), 1861 (4), 1862 (5), 1864 (5), 1866 (7), 1867 (6),
1868 (6), 1869 (8), 1870 (8), 1871 (7), 1872 (8), 1873 (8),
1874 (8), 1875 (7), 1876 (7), 1877 (8), 1878 (8), 1879 (7),
1880 (8), 1881 (7), 1882 (6), 1883 (5), 1884 (6), 1885 (6),
1886 (8), 1887 (7), 1888 (7), 1889 (10), 1890 (9 and 2 Sup-
plts.), 1891 (7), 1892 (7 and Supplt.), 1893 (11), 1894
(8), 1895 (7), 1896 (8), 1897 (8), @ 1/8 per No. to Members
and @ 2/ per No. to Non-Members.
N. B.—The figures enclosed in. brackets give the number of
Nos. in each Volume.
Centenary Review of the Researches of the Bovis, from 1784 to
1883 as - Rs.
Theobald’s Catalogue of Pontiens in ee Acaiseain df ‘the Asiatic
Society (Hxtra No., J. A. S. B., 1868) ... .
Catalogue’of the Mammals and Birds of Burmah, vt HK. Blyth
(Extra No., J. A. S. B., 1875) .
Catalogue of Fossil Vertebrata
Catalogue of the Library of the Asiatic Bobiaby Benbil se
Moore and Hewitson’s Descr iptions of New Indian Teaeinpheee
Parts I-III, with 8 coloured Plates, 4to. @ 6/ each
eu
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18
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NEW SERIES. VOD. Bak VEP: CCGES= ik
JOURNAL
OF THE
ASIATIC SOCIETY OF BENGAL,
Vor-LXVIT. Part ll, No. 2.—189s.
EDITED BY
JHE Natura ge ais
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“ The bounds of its investigation will be the geographical limits of Asia: and
within these limits its inquiries will be extended to whatever is performed by
man or produced by nature.”—Sir WiiuiamM JONEs.
*,* Communications should be sent wnder cover to the Secretaries, Asiat. Soc.,
to whom all orders for the work are to be addressed in India; or care of
Messi's. Luzac ¥ Co., 46, Great Russell Street, London, W. C., or Mr.
Otto Harrassowitz, Leipzig, Germany.
CALCUTTA:
PRINTED AT THE BAPTIST Mission PREss,
AND PUBLISHED BY THE
psiatic Sociery, 57, FARK STREET,
1898.
oi
Price (exclusive of postage) to Members, Re. 1-8.—To Non-Members, Ra 2.
Price in England, 3 Shillings.
Issued September 15th, 1898,
CONTENTS.
ee - Tie
Page
An Annotated List of the Butterflies of the Ké Isles—By LioNEL DB»
Nictvitie, F.E.S., C.M.Z.S., &c., and Heinricu me (with
Plate I)
Descriptions of some new ee ae the Nor th- Hasler n Fr ontiers of
India.—By G. Kine and D. Prain
New species of Entada from Singapore and Borneo. in 12 N.
Ripney, Hsa., M.A., F.Z.8., Director, Botanic Gardens, Singa-
pore. Doinbantaaeted by SurGeon-Masor D. Pratn
On a small collection of Butterflies from Buru in the Moluccas. 5
Lionet pe Nictyiiiez, F E.S., C.M.Z.8., &c.
Plate I will be issued with a subsequent number.
IN A ed) BA
251
284.
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_-—~—-«*' The Proceedings of the Asiatic Society are issued ten times a year
1s soon as possible after the General Meetings which are held on the first
Jednesday in every month in the year except September and October ;
they contain an account of the meeting with some of the shorter and
_ less important papers read at it, while only titles or short resumés of the
oe longer papers, which are subsequently published in the Jowrnal, are given.
ee. The Journal consists of three entirely distinct and separate volumes :
Part I, containing papers relating to Philology, Antiquities, etc.; Part IT,
containing papers relating to Physical Science; and Part III devoted to
Anthropology, Ethnology, ete. 3
Each Part is issued in four or five numbers, and the whole forms
a "previous to which the papers read before the Society were published in
pte a quarto periodical, entitled Asiatic Researches, of which twenty volumes
issued regularly every year. From 1894 an additional volume, Part III,
has been issued.
Journal, but since that date have been separately issued every year.
; The following is a list of the Asiatic Society’s publications relating
zy cae to Physical Science, still in print, which can be obtained at the Society’s
— House, No. 57, Park Street, Calcutta, or from the Society’s Agents in
; London, Messrs. Luzac & Co., 46, Great Russell Street, W. C.; and from
ao ees e = ° -
“ae, ie Mr. Otto Harrassowitz, Leipzig, Germany.
sig Sage Bie <3
= a tate yt os
__ Astantc Researcuzs, Vols. VII, Vols, XIII and XVII, and
oe | Vols. XIX and XX @10/ each Rs. 80 O
7 “ed -___- Procerpinas of the Asiatic Society from 1865 to 1869 (incl.) @
4g ie /6/ per No.; and from 1870 to date @ /8/ per No.
Tee Journat of the Asiatic Society for 1843 (12), 1844 (12), 1845
_ (12), 1846 (5), 1847 (12), 1848 (32), 1850 (7), 1851(7), 1857 (6),
£. FF
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The Proceedings up to the year 1864, were bound up with the ©
1858 (5), 1861 (4), 1862 (5), 1864
1868 (6), 1869 (8), 1870 (8), 1871
1874 (8), 1875 (7), 1876 (7), 1877 (8), 1878 (8), 1879 (7),
1880 (8), 1881 (7), 1882 (6), 1883 (5), 1884 (6), 1885 (6),
1886 (8), 1887 (7), 1888 (7), 1889 (10), 1890 (9 and 2 Sup-
plts.), 1891 (7), 1892 (7 and Supplt.), 1893 (11), 1894
(8), 1895 (7), 1896 (8), 1897 (8), @ 1/8 per No. to Members
and @ 2/ per No. to Non-Members.
(5), 1866 (7), 1867 (6),
(7), 1872 (8), 1873 (8),
N. B.—The figures enclosed in brackets give the number of
Nos. in each Volume.
Centenary Review of the Researches of the Society from 1784 to
1883 eg Rs.
Theobald’s Catalogue of abiiled i in ila Meee of ce Asiatie
Society (Extra No., J. A. S. B., 1868)
Catalogue of the Mammals and Birds of Burmah, by E. Blyth
(Extra No., J. A.S. B., 1875) .
Catalogue of Fossil Vertebrata
Pca of the Library of the Asiatic Séciobd: Bangali
vore and Hewitson’s Descriptions of New Indian Lepidoptera,
Parts I-III, with 8 coloured Plates, 4to. @ 6/ each
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