THE
NAUTILUS
A QUARTERLY JOURNAL
DEVOTED TO THE INTERESTS
OF CONCHOLOGISTS
VOL. XXXVIII
JULY. 1924, to APRIL, 1925
KDIT0R8 AND PUBLIHHERS
HENRY A. PILSBRY
Curator or the Department of Mollusca, Academy ot Natural Scieiicea
Philadelphia
CHARLES W. JOHNSON
Curator of the Boston Society of Natural llUioiy
Boston
//7>
ff')
INDEX
TO
THE NAUTILUS, XXXVIII.
INDEX TO TITLES, GENERA AND SPECIES.
PAGE
Acanthiiia from California, notes on 18, 104
Acanthiua lugiibris Sowb 22
Acanthina paucilirata Steams 21
Aeanthina spirata. var. aurautia Dall 21
Acanthina spirata var. unicarinata Sowb 104
Aehatinella belhila Smith 79
Achatiuella lorata Fer 78
Aehatinella stewardii producta Reeve 78
Aehatinella vulpina olivaeea Reeve 78
Acroluxus 84
Agnathomorphous aulacopoda 86
Alaska, marine sliells of Drier Bay, Knight Island,
Prince William Sound 22
Aleutian Islands, land shells of 122
Amastra nucleola Gould 82
Amblema neisleri nest located 105
Anatina undulata Gould 99
Ancylastnmi 85
Aneylida? 74
Ancylus fluviatilis Mull., the type of Ancylastnira 83
Annandale. Dr. Nelson (obituary) 30
Anodontites flucki Walker, n. sp. (pi. 1, figs. 3, 4) 53
Auriculella auricula Fer 79
Austrobalea Pilsbry, n. gen 6
Balea africana, the affinities of 6
Bermuda, land shells of Admirals Cove 6
Borcherding, Friedrich 106
Buffalo Conchological Section Anniversary 144
Bulimulus alternatus hesperius Fei*ri.ss, n. subsp 40
Bulimulus dealbatus pecosensis P. and F 39
Bulimulus dealbatus ragsdalei Pils 41
Caecum (Meioceras) lennondi Dall, n. sp 7
r,
IV THE NAUTILUS.
Califomian Helices 54
Canada, freshwater inollusea from Mackenzie River Basin. 8
Carelia dolei Ancey 82
Carychium. bermudense Gul 7
Cellana radians (Gmel.) 121
Cephalopods or Elephants on the Mayan Stelae 103
Cepolis bonplandi 59
Cerion agassizii 58
Cerion incanum Binn 59
Collecting in Southern Florida, the Bahamas and Cuba. . 56
Colorado land shells 65
Conchologists in the making 90
Coquina bouillon 32
Donax variabilis Say 32
Drive for The Nautilus 108
Euglandina singleyana W. G. Binn 38
Eulota maackii, a Siberian snail 62
Eulota similaris Fer 80
Fisherola 73
Florida, a remarkable Caecid from 7
God win- Austen, Henry Haversham (obituary) 30
Golisch, W. H. (obituary) 31
Goniobasis clenchi Goodrich, n. sp. (figs. 2-4) 46
Goniobasis dooleyensis Lea 48
Goniobasis livescens Menke 43
Goniobasis plebeius Anth 45
Goniobasis potosiensis Lea 45
Gonyodiscus cronkliitei anthonyi Pils 92
Gonyodiscus patula Desh 86
Gonyodiscus shimeki cockerelli Pils 65
Happia (Happiella) guildingi Bid 87
Hawaiian Islands, a visit to the 76
Hedley, Charies 36
Helix capillata Pease 82
HelLx patula Desh 86
Helix perspectiva of Mergerle von IMuhlfeld versus that
of Say 85
Helix solaria Menke 85
Helminthoglypta fcrrissi Pilsbrv, n. sp 54
Helmiiitlmglyj)ta unil)ilicata Pilsbry 55
Helminthoglypta uiiibilicata cayucosensis Pils., n. subsp. 104
Holospira goldfussi anacachensis Bartseh 39
Holospira roenieri Pfr 42
Japan, new shells from 95
Lacuna from the (California coa.st, notes on 16
THi: NAUTILUS. V
Lacuna divaricata Fab !<>
Lacuna niariiiorata Dall 17
Lacuna ponvcta Cpr 17
Lacuna soliila Loven 16
Lacuna unifasciata Cpr 17
Lacuna varie^ata C'pr 1'^
Lake Titicaca mollusks 108
Lainj)silis superiorensis Marsh 11
Lancidrt' distinfruislied from the Ancyclidae 78
Laiifrfonliella Dall. n. gen 96
Lan«rfonliclla japonica Dall, ii. sp 96
Lanx nuttalli llald 75
Leptachatina fragilis Gulick 81
Leptachtiiui oryza Pfr 81
Lymna'a auricularia in California 125
Lymnft'a peregra geiserieola Beck 92
Mactras of the Pacific coast, notes on 98
Margaritana niarfraritifera L., in Labrador 32
Microcystis chamissoi Pfr 82
Molliisca of the Bridgeport, Connecticut, region 49
Monoceros brevidens Conr 19
Mouoceros engonatum Conr 19
Monoceros lapilloides Conr 19
;Muskrats in Czechoslovakia destroy Margaritana 103
Mya arenaria, an abnormal form 89
Naiades from Nicaragua, two new 51
Nephronaias elvae Walker, n. sp. (pi. 1, figs. 1, 2) 52
Nesophila capillata Pease 82
Newfoundland shells 92
Notes 32, 65, 103, 144
Ostrea ^^rginica Gmel 121
Opeas javaniciun Reeve 80
Oxystyla floridensis Pils 59
Oxystyla undata Bmg 57
Patella tenuilirata Carpenter 121
Pecten acanthodes Dall, n. sp 120
Pecten gibbus Linn 113
Pecten gibbus var. amplicostatus Dall 113
Pecten grandis Sol 112
Pecten heliacus Dall, n. sp 119
Pecten irradians Lam 114
Pecten irradians var. coneentricus Say 114
Pecten liocymatus Dall, n. sp 119
Pecten magellanicus. a large bed of 32
Pecten nucleus Born 115
VI THE NAUTILUS.
Pecten ornatus Lam 118
Peeten raveneli Dall 116
Pecten seiitis Reeve 118
Pecten tereiniis Dall, n. sp 115
Philomycus australis Bergh 80
Physa bottimeri Clench, n. sp. (fig. 4) 12
Physa cupraeonitens and Physa sayi var. warreniana in
hot springs 105
Physa marci Baker, n. sp. (fig. 5) 15
Pleurocera acuta Raf 44
Pleurocerids, some old and a new one 43
Polygyra profunda, the varietal names of 33
Polygyra profunda var. efasciata Walker, n. var 33
Polygyra profunda var. unicolor Walker 34
Polygyra texasiana texasensis Pils 40
Pseudancylus 85
Publications received 34, 66, 106, 143
Purple dye of moUusks 65
Pyramidula (Gonyodiscus) patula Desh 86
Radula technique for Physa 13
Rectartemou jessei 86
Rio Grande, on the 37
Schizothaerus nuttallii Conr 102
Scolodonta eudiscus 87
Sea gull drops clam on policeman 104
Shells of Sanibel, Florida, an additional list 93
Spisula (Hemimactra) catilliformis Conr 101
Spisula (Hemimactra) hemphilli Dall 101
SpisiUa (Hemimactra) voyi Gabb 101
Spisula (Symmorphomactra) falcata Gould 102
Spisula (Symmor])homactra) planulata Conr 102
Streptaxis glaber iiormalis Jouss 86
Stilifer castaiieus Dall, n. sp 97
Stilifer perdepressa Dall, n. sp 97
Strombifonnis langfordi Dall, n. sp 97
Succinea caduca Migh 80
Sweden, a visit to 1
Tanganyika l)eacli 109
Tivehi stultorum Mawe 68
Tornatellides procerulus Ancey 79
Unionida', soiiu' unpublished plates of Conrad's 28
Velletia 84
Veronieella leydigi Simrotli 80
Walkerola . . '. 73
Zalophancylus 75
THE NAUTILUS. Vll
INDEX TO AUTHORS.
Baker. Frank C 15
Baker, II. Burrington 86
Berr\-, S. Stillman 68
Blake, J. Ilenrv 33, 89
Clench. William .1 12, 13, 93, 127
C'cK'kerell, T. I ). A 62. 76
Dall. Wui. II 7, 95, 107, 112, 121
Emery, D. L 56
Eyerdara, Walter 27
Ferriss, James H 37
Gootlrieh, Calvin 43
Hand, Edwin E 90
Ilanna, G. Dallas 122, 125
Hedley, Charles 109
Henderson, Jiiiuns 105
Jaeot, Ai-thur P 49
Johnson, C. W 32, 65, 108
Kenuard, A. S 83, 85
Nylander. Olof 0 1
Pilsbr>% H. A 4, 35, 54, 65, 73, 103, 108
Remingrton. Jr., P. S 127
Sehleseh, Hans 32
Strong. A. M 16, 18, 98, 104
Tomlin, J. R. le B 121
Vanatta, E. G 4, 6, 92, 144
Van Hyning, T 105
Walker, Brvant 33, 51
Wliittaker, E. J 8
Woodward, B. B 83, 85
The Nautilus.
Vol. XXXVIII JULY, 1924. No. 1
A VISIT TO SWEDEN
BY OLOF O. NYLANDER
I left my home iii the woods of Maiue Sept. 10, 1923, and
stopped one day in Boston to see some friends. Leaving
Hoboken, X, J., Sept. 13 on the Scandinavian-American Line
steamer "Hellig Olav," I arrived at Christiansand, Norway,
on the 2-ith, and proceeded by rail to Gothenburg, Sweden.
My object was to see the Tercentennial Exposition of
Gothenburg and to look over the Museum, especially the col-
lection of MoUusca.
Tlie Biological Museum in Gothenburg is a new building
built on the top of a hill in a beautiful park — Slats Skogen.
The collection contains many fine specimens and several large
panoramic views. Prof. S. A. Jagerskiold and his assistant,
Dr. Paul Ilenrici, spent much time in showing me the collec-
tions. The collection of Bohus Lane is most extensive; the
mollusca are arranged in small glass-covered drawei-s with a
fine series of extended animals in alcohol, the best I have seen.
In this museum is also the large collection of land and fresh-
water shells used by C. A. Westerlund in his publications.
Westerlund described many new species that were never illus-
trated. Prof. Jagerskiold said that any American malacolo-
gist would be given every oj^portunity to study the collection.
With Dr. Ilenrici 's assistance, part of two days were spent in
looking through the collection.
iJ THE NAUTILUS.
Oil Oct. 2 I left Gothenburg and went to Skara, a city best
known for its old cathedrals and fine buildings, seminary and
museum of anticjuities. In the park near the museum I col-
lected 230 Helix hortensis Mull., 44 with five bands on a yel-
low ground, 123 plain, varying from a bright yellow to pale
and nearly transparent, representing the varieties lutea uni-
color and alhida. Seventy were the variety p(dlid4i castan^a
Westerlund ; a fine series of this beautiful brown form were
compared with Westerlund types. I spent six days exploring
tlie country about JSkara and then returned to Gothenburg.
I had accepted an invitation from Prof. Jagerskiold to
attend a meeting of the Biological Society of Gotlienburg, at
the Palace Hotel on the 9th, and speak on some American
mollusks — the use of Mija arenaria as food, Busycan perver-
siini and the Indians' use of shells, the fresh-water mussels of
the Mississippi River and the manufacture of buttons ; also the
abalone shells of California. The meeting was attended by
over forty members. Luncheon was served. Professor Carl
Skogsberg presided at the meeting.
On Oct. 11th I went to Boras, one of the large manufactur-
ing cities, to meet Herr Berthold Sundler, an enthusiastic
collector, who has a fine collection, and has done excellent
work in this part of Sweden. In his paper, "Sniickor och
Musslor I Viskana ovre Vattenomrade, " he ably explains the
ecological conditions bearing on the destruction of species. It
might be of interest to note tliat he considers rianorhui {Ar-
miger) crista L. and F. (Armiger) iiautileus L. distinct
species, and seems to have good reasons for doing so. In com-
pany witli lierr Sundler I visited several of his favorite col-
lecting places, but it was loo hite in the season for gooil
collecting. I also visited tlie Historical Museinn.
On Oct. 13th 1 reached Stockholm, the most l)eautiful city
I have seen. The "lliksmuseet" and many other Government
institutions are in new buildings sojne distance out of the city,
known as " Veteiuskaps-staden." Dr. Nils lly. Odhner spent
several days taking me tiirougli the niuscuiu and showing me
the large collection of arctic mollusks collected during the
last century by the various Swedish explorers. I received in
THK NAUTILUS. 3
exchaiijjt* a iiuihIht of iiifrrestiiij^ airtit* species. Dc. ()(liin('i'
is well known in America for his many valuable papers. Dr.
A. (I'Ailly, who spends mueh time working on the lar<;e eol-
loetion of land shells, showed me many of the rare anil beau-
tiful species. 1 am under great obligation to both of these
gentlemen for their many kindnesses and assistance.
October 21 to 23 1 spent in the city of Upsala, famous for
its cathedral, university, library and botanical garden. Prof.
Carl Wiman kindly showed me the collection of fossils in the
nuiseum, and Prof. Erik Stensia assisted me in tiie library,
showing me some rare books, and gave me much valuable in-
formation.
I next visited Lund. This city has nuuiy historical land-
marks and nuiseums. The floors of the Natural History Mu-
seum had just been varnished and I had no chance to see the
collections, but 1 had a pleasant time with Ansamunseu
"Hans Lohmander," who has worked on the raollusks of Swe-
den and has a large privaite collection.
From here 1 went to the old city of Ystad on the Baltic.
In the harbor I collected small specimens of Cardium edule
and Mdcoma halthua. I next visited the old fishing village
Abfekas. Here I found Mya arenuria, rare in the Baltic. I
then went to Oremolla, where, in a ditch near the school house,
I collected Helix hortensis (10 plain and 2 five-banded). This
was a surprise to me, for fifty years ago when I went to this
school I never saw a si)ecimen at this place. In the old church
yard at Skifvarp I also obtained //. hortensis, 2 plain and 12
five-bandeil. I then went to "Ilarte Skag" in company with
an old friend. On the tree trunks I collected 2 plain and 8
five-banded //. hortensis; there were hundreds of young shells
on the trees from 10 to 15 feet from the ground. A short dis-
tance east of here, on the shore of the Baltic, I collected in
1878, 16 specimens of Neritina fliwiatilis- that arc still in my
colleetion.
My next stopping j)lace was "Smyge Fiskclage, " the most
southern part of Sweden. Here I collected Maiomu balthica
and Mi/tilus edulvi. Visiting the church in Ostra Torp I
foiuid on the gravestones 35 //. hortcns^us, all the fivc-I>;i tided
4 THE NAUTILUS.
form. In this graveyard ray father's ancestors are laid at
rest. At "BUagstorp" I visited one of my school teachers,
and together we went to the churchyard, and on the monu-
ment and evergreens collected 10 plain and 15 banded //. )ior-
ten^is. This was my last collecting; it was really too late in
the year to accomplish much. On Nov. 7th I bid my friends
in Ostra Torp farewell and went to Trelleborg and then to
Malmo, my last stop in Sweden.
November 10 to 13 was spent in Copenhagen visiting the
museums. I met Dr. Theodor Mortensen. He said the fauna
on the east coast of Denmark was poor, o\\'ing to the great
amount of fresh water from the Baltic ; he also told me about
his expedition to Kei-Oarna, where he made a large collection.
Magister of Science R. Sparck showed me the collection of
Denmark Mollusca, and kindly gave me some specimens from
the deep water of the Baltic and from Iceland and Greenland.
Leaving Copenhagen on Nov. 14th, I stopped in London on
the 16th and had a glimpse of the British Museum. Arriving
in New York on the 23rd, I spent the next day in the Amer-
ican IMuseum of Natural History, the 26th at the Boston Soci-
ety of Natural History, and was glad to be so near home
again after ten busy weeks abroad.
POLYGYRA APPRESSA LINGTJIFERA AND SANCTA GEORGIENSIS
]JY 11. A. I'lLSBRY AND E. G. VANATTA
An examination of the history of the Bermudan form of
Poljjgyra appressa called Ilclix satuta Georgiotsis led to the
investigation of Helix linguifera of Ferussac and Lamarck.
This name has long rested as a synonym of apprcsaa Say.
The name //. linguifrrn was fii*st listed ])y Ferussiic in
1821, without a description, but with the locality "Les en-
virons de Nogeville, t'tat de Tenesse." This is evidently
Knoxvilie. Two years later, Fc'-nissac ])ul)lisho(l three figures.
Meaiitinic Lamarck had j)ul)lislie(i a (h^scriptioii giving the
same locality. Fcriissjic's figures and Lainarck's description,
so far as it goes, agree very well with shells collected by one
THE NAUTILUS. 5
of US in 1899 at Knoxville, on the south side of the river.
These speoimens hail been referred to /*. apprcssa sculptior
Chaiiw. Tliey are not typical of that subspecies as described
from Scott Co., Virgfinia, tlie periphery being more distinctly
angular in front, and the striation a little stronger and less
close. Both of these charactei"s are seen in Fcrussiic's figures.
The granular sculpture, however, is similar to sculplwr. We
doubt whether there is sutTicient difference to call for two
subspecies. In tliis view, the subspecies will be called Poly-
CYRA APPRESSA LiXGUiFERA (Lamarck), with the following
synonymy :
Ildir linguifera Fenissac, Tableaux Sy.st. de la Famille des
Lima(;ons, 1821, p. 33, no. 95. Les environs de Nogeville, etat
de Tenesse. Nude name.
Helix linguifera Lamarck, Anim. sans Vert. VI, pt. 2, 1922,
p. 90. Nogeville. etat de Tenesse.
Helix linguifera Fer., Hist. Nat. des Moll., pi. 49A, fig. 3
(1823).
[Poli/gi/ra appresfta] sculptior Chadwick, Nautilus XIII,
Sept. 1899, p. 54. Scott Co., Virginia.
Helix sancta Oeorgiensis Terai)le Prime, The Bermuda
Pocket Almanac, 1853, p. 55. Bermuda. Nude name.
Helix sancta Georgiensis W. G. Lane, Five Essays, etc.. The
Spiral Snails of Bermuda (no date, but about 1892), p. 2.
Nude name.
Poli/gijra apprrssa, with the synonym Helix sanctO' georgi-
ensui, Verrill, Trans. Conn. Acad. Sci. XI, 1902, p. 732, text
figure 75 d, e, f on p. 730.
Tiie Bermudan form cannot be said to be quite linguifera
or aculptior. It has their characteristic granulation, but is
slightly heavier in shell and lip, and in the average smaller
than either. Strictly speaking, both sculptior and sancta-
georgiensis have their own slight but recognizable racial char-
acters; but they appear to be subvarieties of linguifera. All
of the Bermudan ''appressa" we have seen, many specimens
from several collectors are of the sanctageorgicnsia form.
Typical Knoxville P. a. linguifera are 16 to 17 mm. tliam-
eter. The type lot of form sculptior measure 15 to 18 mm.
The form sanctageorgiensis measures 12 to 15.2 mm. diameter.
Though the name //. sancta Georgiensis was used as early
THE NAUTILUS.
as 1853, it was first defined in 1902 by Verrill's figures. The
prior notices gave no clue to its characters beyond the associa-
tion with appressa and the locality Bermuda.
THE AFFINITIES OF BALEA AFRICANA MELVILL AND PONSONBY.
BY H. A. PILSBRY
In the course of an attempt to estimate the relations of the
South African fauna to other divisions of the Ethiopian Re-
gion I had occasion to examine Balea africana.^ It has little
in common with the European genus Balea, being much more
like the Japanese Reinia - in shape, small number of whorls
ajid texture; but it differs by the further reduction of aper-
tural armature. Just where the African species belongs in
the system of Clausiliidte remains uncertain. The shell is so
much simplified that recourse must be had to the soft anat-
omy ; but it is certainly not related to Balea.
AUSTROBALEA, DCW gCUUS.
Shell shortly Clausilia-shaped, of few (6-7) whorls. Aper-
ture toothless, with slightly expanded, incomplete peristome.
There is a well-developed, wliolly immersed subcolumellar
lamella within the dorsal side, but no other lamella or plicae.
No clausilium.
Type Balea africana M. & P. Specimens examined from
Karkloof, Natal, collected by H. C. Burnup.
LAND SHELLS OF ADMIRAL'S CAVE, BERMUDA.
BY E. G. VAXATTA
Tile following species were picked from shell-dirt collected
by Mr. Hiram C. Iloyt in 1922. Mr. Arthur Haycock writes:
"Admiral's Cave has an opening in the rock in the roof of
the cave large enough to put your hand through. It is imme-
1 Mclvill :in(l I'oiisoiihy, Ann. M;i^. N. II. IV. is'.Ht. p. 19S.
2 See Pilsl.ry, i'roc ,A. .\'. S. I'liiln. l!Mt], ,,. 471.
THK NAUTILUS. 7
diately below this wIuto tlu* deposit of shells is, and of course
n'cent ones have easy access. Admiral's Cave is between the
roaii leading to the causeway and the road leading to the old
Ferry — an Yg mile from the causeway and nearer this road
than the Ferry one. All the caves and caverns in the rock
ai'ound here are likely places for fossil shells."
Species which have been found fossil only are marked
thus t.
Thifsanophora hypolepia ('Siuitt.', Pils.)
Gasfrocopta rupkola margiiuilba (Pfr.)
Vertigo nuniellata Gul.
Sfrohilops huhbardi Brown
t PoecUozonites nelsoni (Bid.)
Poeciloz&nites bermudensis (Pfr.)
PoecUozonites reinianus (Pfr.)
t PoecUozanites blandi heilprini Pils.
t PoecUozanites vanuttai Pils.
PoecUozanites circumfirmatus Redf.
Zonitoides bermudensis P. & V.
Zonitoides minuscula (Biun.)
t Punctum bristoli (Gul.)
t Carychium bermudense Gul.
The specimens of Carychium bermudense are quite variable
in size ; three measure :
Length 2.25, diameter 1.25 mm.
Length 1.9, diameter 1.1 mm.
Length 1.6, diameter 0.9 mm.
A REMARKABLE CAECID FROM FLORIDA.
i;V W. ir. DALL
Among .some shells sent from Florida by Mr. N. ('. Ijer-
mond were eleven specimens of a Caecum collected at Boca
Ciga Bay, which present some unusual characters.
Caecum (Meioceras) lermondi n. sp.
Shell minute, smooth or with faint incremental wrinkles,
8 THE NAUTILUS.
subarcuate with an obscure medial swelling ; color lucid whit-
ish with four longitudinal rows of small dark bro^\^l spots
articulated with opacjue white spots; two lines are most con-
spicuous on each side of the convex back of the shell, the other
two on each side of the concave portion of the shell are
fainter, and sometimes obsolete; in one specimen the space
between the upper and lower lines on each side is pale bro\ni ;
to these color lines are added more or less evident, transverse,
equally spaced, brownish lines which give the shell a seg-
mented aspect curiously like that of a small maggot ; the colors
differ in strength in different specimens; the posterior end of
the shell is attenuated and decurved, tlie posterior aperture
small, with a minutely mucronate plug; the anterior end of
the shell is larger, the aperture very oblique, almost hori-
zontal, its margin simple, not expanded, nor is there any con-
striction behind it ; the operculum is brownish ; length, 2.5 ;
maximum diameter at girdle, 1.0; diameter of apei'ture, .05
mm. U. S. Nat. Mus. Cat. No. 333531.
When color is present in Caecidie, it is usually either uni-
form over the whole shell or nebulously distributed. The
present species is the first I have been able to trace which has
developed a color pattern. It is quite possible that these
colors are fugitive, at any rate they differ considerably in
strength in different specimens. There is also some difference
in the size of different individuals. The measurements given
are those of the largest of the lot.
Besides the types in the National Collection, others are in
that of Mr. Lenuond.
FRESHWATEK MOLLUSCA FROM MACKENZIE RIVER BASIN. CANADA.
BY K. J. WHITTAKER ^
While engaged in work for tlu^ Geological Survey for sev-
eral sccusons between 1917 and 19212, llie writer, as occasion
permitted, made collections of the freshwater molluscs of the
1 Pul>lisho<l l)y permission of llio Dirt'etor vi the Geological Suiroy
of Canada.
THK NAUTILUS. V
area traverse*!. This iiu-hulod the soiilli shore of Great Slave
lake, the Maekenzie river as far as Fort Simpson, the Liarcl
rivor from its mouth to the South Xahaniii river, and Hay,
Horn, and Kaivisii rivei*s, all tributary to the Maekenzie river.
The writ-er is greatly indebted to ^Ir. F. C. Baker for identifi-
cation and ehecking of species of these molluscs, exclusive of
Spluvriida', and to Doctor W. 11. Dall and Doctor V. St^rki
for identification of members of the above mentioned family.
Through the kindness of Doctor E. ^I. Kindle the writer has
been permitteil to include collections made by him from Fort
Wrigley and Big Island, and additional material from MUls
lake and Hay river.
Collections were made from thirteen localities in all, which
are listed in order below and designated by number. In the
succeeding table the number at the top of each column refers
to the locality to which that number has been assigned.
Li:it of Localities with Number of Species Taken at Each.
1. South shore. Fawn lake, Horn river. 8 species.
2. South shore. Second lake, Horn river. 11 species.
3. Southwest side of Mackenzie river, 30 miles above Fort
Providence. 16 species.
4. Western end of Lake Kakisa. 10 species.
5. South shore of Lake Kakisa near mouth of Kakisa
river. 13 species.
6. Head of Lower Rapids, Liard river, 36 miles above Fort
Simpson. 4 species.
7. At junction of South Nahanni and Liard rivers. 2
species.
8. Mills lake at mouth of Horn river. 6 species.
9. At mouth of Hay river. 10 species.
10. South shore near Sulphur Point, Great Slave lake. 5
species.
11. Big Island, head of Mackenzie river. 2 species.
12. Near Fort Wrigley. 2 species.
13. South side. Mills lake. 14 species.
From the above localities a total of 43 species were obtained
which supplemen/ts considerably the excellent lists with de-
10 THE NAUTILUS.
scriptions given by Dall in the Harriman Alaskan Expedition
series.^ Seventeen forms of pelecypods and twenty-six species
1 Dall, W. H., Harriman Alaska Expedition, Vol. XIII, Land and
Fresh-Molluscs, 1910.
of gastropods, including six terrestrial forms, were recog-
nized. The following list gives their distribution at the dif-
ferent localities :
Felecypoda.
Family Unionid^.
Anodonta grandis footiana Lea. 4.
Anodonta grandis var. imbricata Anthony. 5.
Anodonta var. 9.
Strophitus edentiilus var. 9.
LampsUis superiorensis Marsh. 4, 9.
Family Sphaerud^,
Sphaerium vermontanum Prime. 12.
Sphaerium canadense Sterki ms. 5, 9, 12, 13.
Sphaerium solididum Conrad. 4.
Sphaerium striatinum Lamarck. 4,
Sphaerium tenue Prime. 2, 3, 5, 13.
Sphaerium tumidum Baird. 10 ?, 12.
Musculium jayense Prime. 1.
M usculium rosacenni Prime. 13.
Pisidium campress^im Prime. 13.
risidium indumcnse Sterki. 2, 9.
Pisidium scutellatum Sterki. 1, 2, 3, 5, 13.
Pisidium ventrico^i?n Siterki. 2.
Pisidium, two sp. 5.
Gastropodu.
Family Zokitid.i;.
Euconulus fulvus Muller. G.
Zonitoides nitidus Muller. 4.
Family Endodontid.k.
Pyramidula crankheiti Newcomb. 3, 6, 7, 13.
THE NAUTILUS, 11
Family Succinid-«.
iSuccin^a iivani Say. 3, 13,
l^uccinea chrifsLs Westerluiul. (5, 7.
Suci'inca ntusa Lea. 3.
Family LvMXAKin.K.
Li/nutaea cata-scopimn Say. 1, 4, T), 11.
Lj/ninaca cafascopinni var. 4.
Lymna-ca ohrussa dccampi var. strcugi. 3, 8.
Lymna^a palustris Miiller. 2, 3, 8, 9, 10.
Lymnafa palustris alpoicnsis Baker. 13.
Lymna-ea stagnalis apprcssa Say. 3, 9.
Lymnaea stagnalis var. 1, 4, 8, 11.
Lymnaea vahli (Beck) Miiller. 8.
Plutiorbi-s arcticus Miiller. 1, 2, 4, 5, 13.
Planorbis d^flectiis Say. 9.
Phinorhis exacuous Say. 2, 5.
Pknwrhis hirsutus Gould. 1, 6.
Planorbis parvus Say. 2, 3, 5, 13.
Planorbis trivolvis Say. 3.
Planorbis subcrenatus Cpr. 1, 3, 9.
Scg/nentina armigcra Say. 9.
Segmentina christyi Dall. 1, 3, 8, 13.
Family Amnicolid^.
Amnicola emarginata Kiist<^r. 3, 5, 10, 13.
Family Valvatid.^^.
Valvata lewisii Currier. 1, 2, 3, 5, 10.
Valvata lewisii helicoidea Dall. 2, 4, 8, 13.
Valvata tricarinata Say. 2, 3, 4, 5, 10, 13.
As noted above, there is a considerable variation both in
numbers and species at the different localities. Seventeen
species are added Ix) those listed by Dall from the Mackenzie
basin.
Lampsilis supcriorensis is exceptionally abundant in Hay
and Horn rivers. Thousands are stranded and die on the
banks in early midsumrner as the floo<l waters recede, and
piles of the dead shells are to be seen everywhere. On the
12
THE NAUTILUS.
Other rivers of the area, though conditions seem exactly the
same, this species is rare or absent.
The species recorded from south shore of Great Slave lake
really come from a small pond fifty yards back from the beach
but connected with the lake proper at high water. Shells are
very rare along the beach owing to the fact that, in the break-
up of the ice, the latter is driven up on the beach, destroying
a large part of the moUuscan life.
The fact that casual collecting undertaken wliile the ^^Tite^
was engaged in other work residted in increasing the distri-
bution area of seventeen species should encourage all collec-
tors who have an opportunity of visiting northern latitudes.
However, omng to the boreal and semi-boreal character of the
climate, which imposes a certain amount of uniformity on the
fauna, the number of new species found in any one area is
not likely to be large.
A NEW SPECIES OF PHY8A FBOM TEXAS.
IJY WILLIAM J. CLENCH
The following description is based upon four specimens
sent to me by L. J. Bottimer, of Liberty, Texas.
Physa bottimeri n. sp.
Shell: sinistral, small, subglobular, imperforate, rather thin,
corneous. Color: light horn with a rather dull surface.
Whorls: 41^., the last shouldennl and
very large. Spire : very sliort, the
imclear whorl dark reddish bro\vu in
color. Aperture: large, five-sixths of
the total length of the shell, the outer
edge slightly straightened. Lip : lower
half sligiitly fiariiig. A well-developed
white callus formed along the edge
contiinious willi the l)aso of the colu-
mella. Columella: straight, with a well-defined, slightly thick-
ened fold. Suture: well impressed. SiMilj^ture: composed of
Pliysa liottinu-ri, X
THK NAUTILUS. 18
&ie gTOWtJi lilies, a few coarser lines spaced at irregular in-
tervals proihieing a somewhat costated appearance. Cross
stria' absent.
Tj']>e: Length 6.6, width 4.8, aperture length 5.5 nun. (U.
of M. No. 31617.)
Cotype: L<Mi{]rth 6.3, width 4.8, aperture length 5.5 mm.
(Walker, No. 75895.)
Cotype: Length 6.4, width 5.0, ai)erture length 5.8 mm.
(Clench, No. 1378.)
Cotype: Length 5.5, width 4.4, aperture length 4.8 mm.
(A. N. S. Phila.)
Type locality : Comanche Spring, Fort Stockton, Pecos Co.,
Texas.
Remarks: This spcies of Physa is rather small as compared
with other species of the genus, especially among the shoul-
dered forms. It agrees with Physa lordi Baird in general
outline but ditfers in being smaller in size, in having a white
lip callus, and less thickening and folding of the columella.
A well-formed callus is found on the type and two of the co-
types ; this callus is lacking on the fourth, a juvenile specimen.
Type deposited in the Museum of Zoologj% University of
Michigan ; cotypes in the collection of Dr. Bryant Walker of
Detroit, in my ow^l collection, and in that of the Academy of
Natural Sciences of Philadelphia.
EADULA TECHNIQUE FOR PHYSA.
BY WILLIAM J. CLENCH
University of Michigan
The following method has been found excellent for the ex-
traction, staining and mounting the radula of Physa. As a
preliminary preparation in the field the mollusks were killed
by boiling and were then preserved in 10'/, alcohol. Placing
the Physa in a strainer and then dipping in boiling water for
thirty seconds was found to be the best metiiod. This allowed
easy extraction of the aninuds from the shells. When time
14 THE NAUTILUS.
was available for the study of the radulae, the head and foot
were cut from the rest of fthe body and dropped into a test-
tube containing a solution of potassium or sodium hydroxide
(20 to 40%). The animals remained in this solution for two
or three days, macerating slowly witli the aid of radiator heat.
Usually at the end of forty-eight hours the radulae were free
from the animal tissue and were then transferred to distilled
water. From the water the radulas were transferred first to
chrom-osmic acid, next to crude pyroligneous acid, and then
to 96% alcohol. Only a small amoumt of each acid, sufficient
to cover the radulae, is necessar>^
Euparol was found to be excellent for mounting, much
superior to balsam, inasmuch as complete dehydration and
clearing is unnecessary. Furthermore, the refractive index of
euparol is considerably less than that of balsam, hence fine
details are better revealed under high power. The radidae
can be transferred directly from the 96%: alcohol to the
mounting medium. In the following outline the general pro-
cedure for the method is stated in brief form.
20-40% potassium or sodium hydroxide, 48-72 hours
Distilled water, 15 minutes
Chrom-osmic acid, 2-3 hours
Crude pyroligneous acid, 12-24 hours
96% alcohol, 15 minutes
Euparol.
The chrom-osmic acid is made by dissolving 1 gram of osmic
acid in 50 cc. of a 1% solution of chromic acid. Best results
will be obtained by using thin Euparol. Diaphane or Moun-
tene may be ustnl in place of Euparol, as these three mount-
ing mediums are the same but sold under ditt'erent trade
names.
This is an imi)regnation methoil of staining and results
in a uniform opaijue brown stain.
THK NAUTILUS. 15
A NEW PHY8A FROM CALIFORNIA.
15Y FRANK COLLINS BAKER
Of tho I'liiversity of lllinoiw, Urbana, Illinois
Physa MAiici, new spwies.
Shell small, sinistral, elongate-ovate; whorls 4-5, flatly
rounded ; spire short, acutely dome-shaped, one-half as long as
aperture ; body whorl compressed, flatly rounded ;
color yellowish or greenish horn ; sculpture of
coarse lines of growth %nth no (or but very fijie)
si)iral imi)ressed lines; sutures well impressed,
deeply so between body and penul'iiraate whorl ;
aperture elongate ovate, widely rounded below,
acutely angled above, occupying two-thirds of the
p/iysa marci. length of the shell ; peristome thin, sharj^, with
rarely an indication of internal rib ; columella
covered witli a thick, very wide callus, whicli is especially
wide near the umbilical region, either completely closing the
umbilicus or leaving a. very small, narrow chink ; color of in-
terior of aperture whitish.
Length 6, width 4; aperture length 4.1, width 2 mm. Type.
Length 4.5, width 3.3; aperture length 3.2, width 1.5 mm.
Paratype.
Type locality: Little Valientia Spring (hot sulphur spring),
Santa Barbara National Forest, California. Collected by Mr.
Marcus Dall.
This litrtle Physa differs from all other species at present
known from the southwestern part of the United States. It
somewhat resembles small forms of the eastern Physa qyrina
Say, but differs wholly in sculpture as well as in size. Physa3
from chemical springs are apparently rare. The small size
and form of this species are probably due to the environment
which seems to have inhibited growth.
Type, Cat. No. 359544, United States National Museum.
Paratype, No. Z14210, Natural History Museum, University
of Illinois. Cat. No. 359545, U. S. N. M., contains a paratype,
and Cat. No. 359546, U. S. N. M., 18 specimens, all from the
type locality.
16 THE NAUTILUS.
NOTES ON LACUNA FEOM THE CALIFOENIA COAST.
BY A. M. STRONG
Dr. Dall in his List of Northwest Coast Marine Shellbear-
ing Mollnsca gives nine species and varieties of Lacuna as
occurring on the California Coast. The ranges and references
given indicate a considerable revision in the nomenclature of
the genus. The following notes give a brief review of the
California species.
Lacuna divaricata Fab., 1780 (Fauna Gronlandica, p. 392),
is a very variable circumboreal shell with a wide range. In
ithe Pacific it is listetl as ranging from Norton Sound, Alaska,
to Santa Barbara, California. Tryon's ]\Ian. Conch., vol. 9,
p. 265, gives a long list of synonyms including L. vineta Mtg.,
L. solidula Loven, and L. ?var. compacta Cpr. The first of
these is mentioned by Carpenter and others from Alaska but
it is now considered by Dr. Bartsch to be a distinct Atlantic
species. (Nautilus, vol. 35, p. 89). The second is also ac-
cepted as a distinct species occurring in both the Atlantic
and Pacific but the position of the third is left uncertain.
The typical form of the species is figured in Tryon's ]\Ian.
Conch., vol. 9, pi. 50, fig. 75.
Lacuna solidida Loven, 1846 (Index Moll. Scand., p. 23), is
listed by Dr. Dall as occurring in the Pacific from Puget
Sound to San Diego, Dr. Baker adds EUamar, Alaska (Nau-
tilus, vol. 24, p. 44). It is figured in Tryon's Man. Conch.,
vol. 9, pi. 50, fig. 69. Dr. Arnold recognizes it as a Pleistocene
fossil in Southern California (Pal. San Pedro, p. 303), and
gives as synonyms Littorina pedroana Conr. and Modelia stri-
ata Gal)]).
Dr. Could described a sliell from Puget Sound under the
name of Lacuna carina ta (Proc. Boston Soc. Nat. Ilist., 1848,
p. 75), which Dr. Carpenter consitlered to be the same as L.
solidida Loven, from the Atlantic. Dr. Arnold described a
Pleistocene fossil from San Pedro as L. compaita Cpr. (Pal.
San Pedro, p. 302), and for a figure referred to Tryon's Man.
Conch., vol. 9, pi. 50, fig. 71. This is the figuiv for L. caritiata
THK NAUTILUS. 17
GouM. Dr. Carpenter's shell was described as Ixuumi ( ? suli-
duUi var.) vompacta Cpr., from tlic Swan collect ion at Noah
Bay, Wash. (Aiu». Ac Mag. Nat. Hist., vol. 14, lbU4, p. 42!ij.
If the descriptions correctly represent the type specimens
theso three would all seem to be the same shell, which should
taivc the name of Lacuna solidulu var. carinala Gould. For
range, Dr. Arnold gives, living, Vancouver district (Carpen-
ter). Specimens from Dana Cove, Orange County, California,
have been identitied by Dr. Dall as L. {solidula var.j carinata
Gould.
Lacutia porrecta Cpr., 1864, and varieties effitsa Cpr. and
exa-equata Cpr., were described from the Swan collection at
Neali Bay, Wash. (Ann. & Mag. Nat. Hist., vol. 14, p. 429).
The species is listed as ranging from Bering Sea to San Diego
and is figured in Tryon's Man. Conch., vol. 9, pi. 50, fig. 55,
and in Bull. U. S. Nat. Mus., No. 112, pi. 14, fig. 2. Variety
effusa Cpr. is listed as ranging from Fuca Straits to San Fran-
cisco, while variety exaequata Cpr. is only known from the
type locality. Both forms are figured with the typical in
Tryon's Man., figs. 56 and 57. A third variety, puteoloides
(Cpr. MS.) Dall, 1919 (Proc. U. S. Nat. Mus., vol. 56, p. 349),
is listed from Lobitas, California, and remains unfigured. Dr.
Arnold recognizes the species as a Pleistocene fossil in South-
ern California (Pal. San Pedro, p. 303).
Lacuna unifasciata Cpr., and variety aurantiaca Cpr., were
described from Col. Jewett's collecting at Santa Barbara
(Proc. Zool. Soc, London, 1856, p. 205). The name unifasci-
ata Cpr. has been applied to a group of California shells which
Dr. Dall seemingly divides into three species. He gives the
shell figured in Tryon's Man. Conch., vol. 9, pi. 50, fig. 74, as
the true unifasciata of Dr. Carpenter. The range is given as
Santa Barbara, California, to Magdalena Bay, Lower Cali-
fornia. Variety aurantiaca Cpr. has not been figured, the
range is given as from Santa Barbara to Point Abreojos,
Lower California.
Lacuna marmorata Dall, 1919 (Proc. U. S. Nat. Mus., vol.
56, p. 348) is the shell <lescribed in Keep's West Coast Shells
and listed in Packard 's Molluscan Fauna from San Francisco
18 THE NAUTILUS.
Bay as L. nnifasciata Cpr. It is figured in Tryon's Man.
Couch., vol. 9, pi. 50, fig. 58, as L. variegata Cpr. The range
is given as from Saginaw Bay, Alaska, to San Diego, Cali-
fornia. This seems to be the most common species in Cali-
fornia.
Lacuna variegata Cpr., 1864, was described from the Swan
collecting at Neah Bay, Wash. (Ann. & Mag. Nat. Hist., vol.
14, p. 429). Of the figures given in Tryon's Man. Conch.,
vol. 9, pi. 50, Dr. Dall considers fig. 64 to come nearest to it.
This figure is given as L. fusca Binney, a form of L. divari-
cata Fab. The range is given as from Neah Bay, Wash, to
Monterey, but it has been collected at several points in South-
ern California as far south as Reef Point, Orange County.
The last three species and var. aurantiaca Cpr. are quite
similar in form and texture, but in typical specimens the
color pattern is quite different. L. iinifasciata Cpr. is uni-
formly light brown with a narrow brown line on the keel of
the body whorl. Var. aurantiaca Cpr. is orange w^hite \\'ithout
color markings. In L. marmorata Dall, the keel of the body
whorl is marked by alternating white and brown spots and a
spotted pattern frequently extends over more or less of the
shell. The colors of L. variegata Cpr. are clouded or in zigzag
stripes and the keel of the body whorl is often marked by
white splashes. The details of the color pattern vary consid-
erably and fade in dead shells and cabinet specimens. If the
separation into different species is to be based on this color
pattern, var. aurantiaca Cpr. should be given specific rank.
It might just as well be a variety of one species as of the other.
NOTES ON ACANTHINA FROM CALIFORNIA.
nv A. M. STRONG
The proper arrangcTnent of the various forms belonging to
the genus Acanthi na, formerly Monooeros, wliieli are to be
found in California has been a puzzle to all tiie writers. The
few specimens wliich roacliod the early writers were seeiuiiigly
Cfisily separated into distiiR't species and a number were
THK NAUTILUS. 19
naineil. As larger series became available for study many iii-
tor«j:rii(los were found ami the (inestion of varieties and later
of priority of names became a ilitlieult one. Dr. Dall in linl-
letin U. S. National Museum, No. 112, 1922, rec(^nize8 Acaii-
thina spinita Blainville. 1S33, as tiie first name used and
therefore the typical form. All othor California foruLs, with
the exception of two species from the southern fauna found as
far north as San Dietro, are reduced to varieties.
Conrad in 1837 described three species from Santa Barbara,
Monoceros engonatum, Monoceros brevidens, and Moiwceros
lapUloidcs. At about the same time Sowerby described
Monoceros punctulata and Monoceros unicarinatum. At vari-
ous later dates the specimens in the early European collections
were redescribed and fignretl by Reeve, Deshayes, Gray and
Keiner, seemingly without much regard to the use of the
names. Dr. Carpenter in the Proc. Zool. Soc., 1856, p. 228,
gives the synonymy as follows:
Monoceros engonatum Conrad
= Monoceros unicarinatum Reeve, non Sowerby nee.
Deshayes
= Monoceros unicarinatum pars, Deshayes
Comp. Purpura spirata Blainv.
Monoceros brevidens Conrad
= Monoceros unicarinatum Sowerby no)i Reeve tiec.
Deshayes
Non Monoceros brevidentatum Gray
Monoceros l.vpilloides Conrad
= Monoceros punctulatum Sowerby
= Monoceros punctatum Gray
=^ Monoceros punctatum Reeve
Dr. Carpenter's comments in this connection are of interest.
Under M. eiujoimtum Conrad he says: "The shell figured by
Conrad and found in Mr. Nuttall's collection is very trian-
gular, with a pointed base. The P. spirata of Blainville is a
Sandwich Island shell, brought by M. Botta, very obtuselj
angulated, with a swollen base, scarcely acanthoid, and a canal
long enough for Chorus, Gray. It is remarkable for the scaly
keel of the upper whorls. This shell is reproduced by Keiner
20 THE NAUTILUS.
ill a different form, who affiliates Sowerby's species to that in
the Paris Museum. Deshayes, copying this error, and not
even adopting Blaiuville's earlier specific name, gives the
name and reference of Sowerby, with a description in the
main belonging to the Bhiinville species, although perhaps
with some additions from Sowerby's figure. Mr. Reeve com-
pletes the confusion by describing a shell, 'anfr. superne
angulatis,' very probably the true enganatum of Conraxl,
which he quotas; at the same time quoting the two different
shells above named (one of them under two names, P. spicata
and P. spirata), and figuring a very different shell, not angu-
lated at all. To mere learners, like the author of the present
paper, such differences are exceedingly perplexing," Under
Monoceros hrevidens Conrad he says: "The exact date of
Sowerby's species, which is generally referred to P. engonata
of Conrad, but differs from the figure of that shell, and agrees
much better with the description of this, is difficult to deter-
mine. The volume bears the date 1841. It differs from P.
enganata in being swollen at the base, with less sculpture and
angulations." Finally, in regard to all three forms appears
this: "The differences between the specimens of California
Monoceros are so numerous, and similar specimens from other
quarters are so variable, that the three species here rep)eated
from Conrad are given with ver>' great hesitation. That the
forms figured by Sowerby and Reeve are conspecific. is by no
means im])robable; the form engonata is the most aberrant
but it is by no means unapproaclied."
Later Dr. Carpenter evidently changed his mind to some
extent in regard to the different forms, as in his Supplemental
Report, 1863, p. 663, he lists three forms from the California
coast, as follows:
M(»io<cros cngonafum Conrad ^^ loiicorinatuni Sowerby.
Brown-dotted, with sharj) j)ostorior keel, snioothish.
Monoceros ? var. s])lniluni (lihiinv.). Light colored,
.scaly ; horn not develoi)ed.
Monoceros UipiUoides Conrad = punctatum Gray -|-
hrevulens Conr. Not shouldered, shape of l-apUlus.
For many years writers on California shells and the collector8
THK NAUTILUS. 21
have been followiiif? Carpenter in tlie use of etigonotum Con-
rad antl hipilloidrs' C'onnul for tlie two coiunion forms.
iSpirata Blainville lias been rej^'anled as a variety of ( ngo-
fuita Conrail by nearly all writers. Arnold mentions it in his
Paleontolofry of !San Pedro, p. 24G, as follows: "Specimens
showin<r the scaly surface of var. spiratum grade over into the
smooth form ; the sharp-keeled forms merfje into those which
approach very near to M. lapUloides. Some specimens have
thin and smooth out«r lips, while otliei's are strong^ly dentate.
Of fifty sjiecimens from the San Peilro Pleistocene only three
have the characteristic tooth developed, thus showing that in
the earlier form this distinguishing character was only occa-
sional. The scaly specimens are generally the strongest
keeled. This later form is the var. spiratum of Blainville."
Stearns, in the Am. Journ. Couch., vol. 7, 1872, applies the
name of var. spirata Blainv. to a local form founil at San
Diego, while it is stated in Tryon's Man. Conch, that spiratum
Blainville and cnqonatum Conrad are undoubtedly the same.
If we are to take the light-colored scaly variety describeil by
Carpenter and Arnold as the typical form for the species, the
darker and smoother form which has so long been recognized
as having at least a varietal ditt'erence should take the name
of Acanthina spirata var. en^&nata Conrad, 1837.
It is pretty generally recognized that all gradations are to
be found between the forms long known as Moiioceros eiigo-
twta and Monoceros lapUloid-es. Dr. Dall gives the date for
Sowerby's Monoceros punctulata as 1835 and Monoceros lapU-
loides becomes Acanthina spirata var. punctulata Sowerby,
1835. It is the fonn found on the exposed rocks of the outer
coast and under a different environment takes on a distinct
form.
Another variety, Acanthina spirata var. aurantia Dall, was
described in Proc. U. S. Nat. Mus., vol. 34, p. 248. This is a
color form of variety enqonata Conrad. Similar orange or
lemon tinted specimens of quite a luimber of California shells
have been given varietal names.
Acanthina paucUirata Stearns, 1871, wa.<9 described from
San Diego in Conch. Memo. No. 6, p. 1. The species is stated
22 THE NAUTILUS.
by Lowe (Nautilus, vol. 27, p. 26) to be "In their prime'' at
Cape Coluette, Lower California. In general shape they are
quit« similar to small specimens of variety punctidata Sowerby
but the color pattern is quite distinct. They are " externally
painted with longitudinal broad black and narrow whitish
streaks, interrupted by the white dental groove,'' thus cutting
the black colored streaks up into large squares. In variety
pioictulata the light color preilominates and the dark is re-
duced to small square dots. A large series recently collected
near Lagiina, Orange County, show markings intermediate
between the two. It is quite probable that further collecting
and study will show that paucilirata is only a southern form
of variety punctukita and it will also have to be classed as a
variety of Acanthina spirata Blainville.
The Lower California species Acanthina luguhri^ Sowerby,
1822, is reported from San Diego on the strength of a single
specimen found there some years ago. Orcutt reports it quite
common as far north as the International Boundary Line. It
has been stated that Acanthina paucilirata Stearns forms the
connecting link beween higuhris and engonata. If further
collecting along the northern portion of the coast of Lower
California, where the ranges of the different forms overlap,
shows the full series of intergrades, still further changes in
the nomenclature will have to be made. In groups of this
kind where the forms are so variable it is dit!icult to draw the
line between variety and species, and the (luestion of possible
hybrids is always present.
YARIWE SHELLS OF DEIER BAY. KNIGHT ISLAND, PRINCE WILLIAM
SOUND, ALASKA.
BY \V.\LTER EVERDAM
During the summer of 1923, while engaged at my trade as
cooper iji the herring saltory of tlie Knight Islaiul Packing
Co. in Drier Bay, Prince William Sound, I utilized much of
my spare time in dredging and searching for shells along the
beaches.
THE NAUTILUS. 28
Drior Hay is about 35 miles by boat from La-Touclie; it in
an indentation of Knij^ht Island, about 10 s<|uare miles in
area. !St<M."i) mountains surrouml the bay, causinf? steep rocky
beaehes in most places and often fairly deep water in close to
the shoH'. In Mallard Ray, Cat-head liay and Miner's Cove,
ai> well as several other coves and small bays belonging to
Drier Bay proper, the beaches are covered with fucua and at
low tiile with abundance of eelgrass, and tiie water is quite
sliallow.
The best collecting was obtained by washing the roots and.
leaves of eelgress, which oft<?n grew to the length of 12 feet.
About a ton of eelgrass was washed in tubs and the mud
screened and panned, after which the remaining debris and
shells were carefully examined and separated, and many rare
species of small shells were thus obtained.
Most of the Odostomias and several other small rare species
were found among stones and sea-moss at low tide in [Mallard
Bay. About a dozen species of Chitons were found, some of
them from a depth of 200 feet.
Dredging was rather poor considering the large amount of
physical effort that was performed. About 150 hauls were
made at depths of 10 to 40 fathoms. Most of the dredging
was done near the herring dock by rowing out about 200 feet
and dropping the dredge and then rowing back to tie the boat
to a piling, after which I proceeded to haul in the dredge
rope by hand. It was generally chockfull of rocks and debris
and often weighed over 200 lbs. The results w^ere not always
gratifying considering the meager returns that I generally
got, but whenever I found a new shell I would try again.
Sometimes in the fall of the year when there was very little
work to do in the shop 1 would spend most of a day out in a
small rowboat dredging shells. Three times during the season
I lost my whole dredging outfit in waters ranging from 35 to
200 feet in depth, but each time T succeeded, with patience
and perseverance, to snag it up again.
Many rare small shells were dredged, including two new
species: TurhonHln {Pyrcjolampros) eyerdami (Bartsch) and
run<:turfUa eyerdami (Dall), and twenty-six spei-ies were
found beyond their geographical ranges.
24 THE NAUTILUS.
I\Iost of the shells were sent to Doctor Bartsch for identifi-
cation, to whom I am greatly indebted.
The follo^\•ing shells, comprising 139 species, were collected
in Drier Bay :
Nucula tenuis (Montagu). Scarce, in soft mud; dredged.
Leda fossa (Baird). Not common, in mud, mostly dead shells;
dredged.
Leda minuta (Fabr.). Scarce, in mud; dredged.
Yoldia my alls (Couth.). 2 or 3, in soft mud; dredged.
Yoldia montereyensis (Dall). Rare, in soft mud; dredged.
Yoldia species. In soft mud, dead shells ; dredged.
Chlamys hericeus (Gould). Fairly common on stony bottom.
Propeamiisium alaskense (Dall). Not uncommon; dredged.
Limatida subauriculata (Mont.). Quite common; 30 fathoms.
Monia macrochisma (Desh.). Common on rocks at low tide;
very thin.
Mytilus californianus (Conrad). Rare, at low tide.
MytUus edidis (Linn.). Very common.
Modiolus modiolus (Linn.). Very common.
Musculus niger (Gray). Rare; dredged.
Muscidus vernicosus (Midd.). Very common on eelgrass to
which it is attached by a byssus.
Crenella decussata (Mont.). Common on roots of eelgrass.
Thracia curia (Conrad). Rare, in fine mud; 25 fathoms.
Kennerlia hilirata (Conrad). Not common, fine iiuul.
Cuspidaria heringensvi (Leche). One specimen.
Cuspiduria pla)heti<:a (Dall). Two specinu'ns, nuiddy. 25 fra.
Astarte alaskensi^ (Dall). Common; 10 to 40 fathoms.
Astartc ricfoci/ma rsquimalti (Baird). Common; dredged.
Veiiericardia crehricostata- (Krause). Rare.
Thyasira gouldii (Philippi). Common; dredged.
Diplodonta alrutini (Dall). Under stones at low tide.
rarvilucina tcnui-sculpta (Cpr). Common; dredged.
Serripes groenlandicus (Gmelin). Rare; dredged.
Sd.ridomus gigantcus (Desh.). Very common on sandy
beaches.
Marcia kennerlyi (Reeve). A few specimens; 20 fatlioms.
THE NAUTILUS. 25
Mania suhdiaphana (Cpr.)- Rare; dreilgeci.
I'rotothaca stominea (Conrad). Comuiou on muddy, stony
l)eaches.
I'stphidui lonli (Baird). Not common; dredged.
M a*: 0 ma sit k ana (Dall). On muddy beaches.
Macoma tio.futa. Muddy beaches.
Maioma balthica (Linn.). Very common on muddy beaches.
Spisiila {Pohjnyma) alaskana (Dall). Rare on beach.
3/1/0 truncata (Linn.). Not common, on muddy beach.
Mya intermedia (Dall). Fairly common on muddy beach;
young.
Soxicava arctira (Linn.). Very common on old piles and sub-
merged wood and also floating wood.
Saxicava pholadis (Linn.). Same habitat but less common.
Bankia setacea (Tryon). Destroys all wharves and wootl in a
few years.
Cadulus h( phurni (Dall). Common below low tide mark.
Acteocina culcitella (Gould). Dredged; not common.
Acteocina eximia (Baird). Common on eelgrass roots.
Retusa semen (Reeve). Dredged frequently.
Retusa harpa (Dall). Dredged; not eoimnon.
Ilaminoea olgae (Dall). Very common on eelgrass.
Lara quadra (Dall). Dredged; two specimens.
Lora morchi (Leche). Dredged; one specimen.
Lara harpularia (Couthony). Dredged; a few specimens.
Lora solida (Dall). Dredged; six specimens.
Lora chiachiana (Dall). Dredged; not uncommon.
Lora rosea (M. Sars.). Dredged; about a dozen specimens.
Lora heckii (Moller). One specimen.
Admete couthauyi (Jay). Dredged; quite common.
Olivella boetica- (Cpr.). Dredgeil ; one specimen.
Searlesia dira (Reeve). Very common on rocky beaches.
Sehizopyga mendica (Gould). On beaches, not connnon.
Tritomdia interfosaa (Cpr.). Dredged; not common.
Neptunra ienuviculpta (Cpr.). Rare, under stones at low tide.
Thais lamellosa. (Gmelin). Very common on rocks at high
tide mark.
Thais lima (Martyn). Common on rocks.
26 THE NAUTILUS.
Thais enmrginata (Deshayes). Common, rocks.
MelaneUa micans (Cpr.). Dredged; one specimen.
Turhonilla {Mormula) eschscholtzi (Dall & Bartech). Rare.
TurhonUla {Pyrgolumpros) eyerdanii (Bartseh). 3 specimens.
New species. Dredged from 20 fathoms, shelly bottom.
Odostamia. (Evalea) quadrae (Dall and Bartseh). Among
stones on beach, rare.
Odostomia (Evalea) skidegatensis (Bartseh). Among stones.
Odostomia (Evalea) wiUetti (Bartseh). Among stones on
beach.
Odostomia (Evalea) stephensae (Dall & Bartseh). Among
stones on beach.
Odostomia (Amaura) sanjuatienses (Bartseh). Rare, among
stones at low tide.
Odostomia (Amatira) talpa (Dall & Bartseh), Among stones
at low tide.
Cypraeolina pyriformi^ (Cpr.). Fairly common on seaweeds
at low tide in Mallard Bay.
Cerithiopsis charlottensis (Bartseh). Not common.
Cerithiopsis onealcnsis (Bartseh). One specimen.
Bittium (semibittittm) vancouverense (Dall & Bartseh). Com-
mon at low tide.
Trichotropis canccllata (Hinds). Dredged; common.
TricKotropis insignis (Midd.). Dredged; common.
Michanellum ore.gonense (Bartseh). Dredged; not common
Fartulum occidentale (Bartseh). Dredged; not common.
Tachyrhynchus erosus major (Dall). Dredged; common.
Tachyrhynchus lacteolus (Cpr.). Dredged; not common.
Littorina sUchana (Philippi). Very common on rocks.
Littorina rudis (Donovan), Very common on rocks.
Littorina sctttvlata (Gould). Common on rocks.
Lacuna divaricata (Fabr.), Very common on eelgrass.
Lacuna sp. On eelgrass,
Cingula aleutica (Dall), On eelgrass; six specimens,
Alvania reticulata (Cpr,), Common on eelgrass roots,
Crepidula ling u lata (Gould), Not eoimnoii ; dredged,
Crrpidula nummarius (Gould). Dredged; not common.
Natica (Cryptonatica) aleutica (DaJl). Dredged; common.
THR NAUTILUS. 27
Natica (Euspira) pallida (Brod. & 8owb.). Dredged; not
coiiuuon.
Natica {Eits-pira) monterana (Dall). Dredged ; one specimen.
Vflutina laevigata (Linn.). A few specimens from stones.
Lcpeta concentrica (Midd.). Common at low tide on rocks.
Lcpeta caecoides (Cpr.). Common at low tide under stones.
Acmaea nitra (Esch.). Two specimens.
Acmaca cassis pelta (Esch.). Common.
Acm-aea scutum (Esch.). Not common.
Acmaea pframahilis (Dall). Three young specimens.
Cidarina cidaris (A. Adams). Three specimens, mud; 40 fm.
Machacroplax varicosus (IMigliels & Adams). Dredged; one
specimen.
PupiUaria pupUla (Gould). Very common at low tide.
PupUlaria cinerea (Couth.). Dredged; one specimen.
Pupillciria pupUla. Several varieties.
Margarites helicinus (Phipps), Very common on eelgrass.
Margarites marginata (Dall). Very common on eelgrass.
Diadora aspera (Esch.). Scarce, on rocks at low tide.
Puncturellaeyerdami (Dall). New species. Three specimens.
25 fathoms.
Puncturella galeata (Gould). Dredged ; not common.
Puncturella cucullata (Gould). Dredged; not common.
Siphonaria thersites (Cpr.). Common on fucus.
Kellia laperousii (Deshayete). Among stones on the beach.
Hemithrys psittacea (Lam.). Dredged; not common.
Terehratalia transversa (Sowb.). Dredged; not common.
Laqueus calif ornicus (Koch). Dredged; young; common.
Terehratulina unguicula- (Cpr.). Dredged; not common.
Terehratalia caurina (Gould). Dredged ; common.
Spirorhis (species). Very common on fucus and eelgrass.
Columhelld (Nitidella) gouldii (Cpr.). Dredged; 2 specimens.
Roche fort is tumida (Cpr.). On eelgrass.
Turtania minuta (Fabr.). On eelgrass.
Crijptomya calif ornica (Conrad). Dredged; two specimens.
Cylichnella alba (Brown). Dredged ; not common.
Argobuccinum oreganense (Redf.). Only one mature speci-
men. This species is very abundant in some parts of
Prince William Sound.
28 THE NAUTILUS.
Bucciniim morchunium (Fischer). Common under rocks.
Track ydermon ruber (Linn.). Dredged; not common,
Trachyderman albiis (Linn). Dredged ; not common.
Tracliydermon (Cyanoplax) hartwegii (Cpr.). Under stones.
Tonicella suhmarmorca (Midd.). Dredged; common.
Ischnochiton interstinctus (Gould). Dredged; common.
lachnocliiton {Trioplax) trifidus (Cpr.). Dredged; rare.
Mopalia sinuata (Cpr.). Dredged; rare.
Katheriim tunicata (Wood). On rocks; scarce.
Mopalm (ciluita) woss-nessenskii (Midd.). Under rocks; not
common.
Tonicella lineaia (Wood). Under stones; common.
Mopalia muscosa (Gould). Under stones.
Cryptochiton stelleri (Midd.). On rocks; not common.
Besides the above list of 141 species, about 50 other species
of invertebrates were collected. Among these are some very-
rare starfishes and crustaceans. Two very rare crabs, Chio-
nectes hairdi (Rathbun) and Pinnixa schmitti (Rathbun),
were retained by the U. S. Nat. Museum, and also a number
of a new species of Bryozoa, Discocylis (Bassler), the last spe-
cies being the first of its kind from Alaskan waters.
Two land shells were found, being fairly common in one
place under old boards and building paper — Polygyra Colum-
biana (Lea) and Circiuarui sporfcJhi (Gould).
SOME UNPUBLISHED PLATES OF CONRAD'S "UNIONIDAE".
BY WILLLVM J. FOX
Academy of Natural Scieiii-os of Phila(li'I])liia
The abrupt ending (witli the iiicoinph'te word beauti- on
I)age 118) of the work "Monograi)hy of the Family Union-
idte, or Naiades of Lanuirck " by T. A. Conrad, Phila-
delphia, 1836 [1835-18401, indicated tliat fnrther parts were
then contemplated. In all, Ihirteon a|)i)c;ir('(I, with five plates
each, a total of sixty-five plates.
THK NAUTII.LH. 29
The work was presumably cliscoiitinued on ac-eoiint of ill-
ness aiid lack of fiuaiu-ial suijport (see Popukir Science
Monthly ^ XLVII, p. 260, 1895) ; and it is interestin*!: to note
that the library of the Academy of Natural Sciences of Piiila-
delphia has just come into j)Ossession of seven unpublished
plates. These were obtained, with a miscellaneous lot of nat-
ural history pamphlets, from a seconil-hand book store in
Philadelpiiia about twenty-five years ago, and have remained
unnoticed until their donation to the Academy. They are
inscribed in lead pencil by Conrad as follows: Unio ligamen-
tina 1 Lam., pi. G9 ; ihiio infucatus Conrad, pi. 70; Unio
nasutus Say, pi. 71; Unio Jaycnius Lea [the name Jayanus
crossed out and Jayensis substituted], pi. 72; U. verucos-us
Raf., pi. 73; U. TubercuUitus Raf., pi. 76; and a plate, with
no inscription, containing three figures of what is known to
modern systematists as Quadrula cylindrii'a Say.
These plates are colored lithographs, and are on a paper
differing in quality from that used in the book as issued; it
would seem that they are the lithographer's proofs, colored as
samples to foUow. There is also a proof copy of U. plicatus
Say, pi. 65, which is the last plate in the book as published.
If the parts intended to follow No. 13 were to have five
plates each, as in the preceding numbers, it would indicate
that we have here part of the plates for Numbers 14, 15
and 16.
The plato 72 is inscribed (also in lead pencil) C. A. P. jr.
del., which suggests the name of the artist, Charles Poulson,
Jr., who was elected a member of the Philadelphia Academy
ill Ls41, and was no doubt a relative (son perhaps) of Charles
A. Poulson, to whom Conrad's book is dedicated. It has been
taken for granted that Conrad himself was the delineator of
these illustrations.
30 THE NAUTILI':
DR. NELSON ANNANDALE.
The death of Dr. Annandale, Director of the Zoological
Survey of India, in Calcutta on April 10, at the age of forty-
eight, is a severe loss to science. Dr. Annandale became
Deputy Superintendent of the Indian Museum in 1904. Two
years later he was elected Superntendent. Annandale at one
time or another worked on most groups of animals. He be-
came known as an authority on freshwater si)onges, polyzoa,
eltc, and on stalked barnacles, also publishing on fishes, rep-
tiles and insects. In the last few years he worked with great
industry and genius on freshwater mollusks, his papers cover-
ing a wide field, from Mesopotamia to Japan, and ({uite trans-
fonning this branch of Indian zoology. Most of these studies
were published in the Records of the Indian Museum, Calcutta,.
HENEY HAVEBSHAM GODWIN-AUSTEN.
Lieutenant-Colonel Godwin-Austen died on December 2nd
last, in his ninetieth year. He went to India in 1851 as a
subaltern in the 24tii Foot, but after six years' service, joined
the Indian Survey Department as topographical assistant.
He was the pioneer explorer of the great mountain ranges of
Baltistan, north of the Indus and Kashmir, and fixed the
position of the second highest mountain in the world, s\ibse-
<inently named IMount Godwin-Austen. He carried out mucli
survey work in the northeastern hill regions, bordering
Assam. During this service God win- Austin collected birds,
ui)on which he j)ul)lislie(l many papers, and land snails, upon
wliich he finally specialized. Besides many i)apers on mol-
lusks in periodicals, he published The Land and Freshwater
Mollusca of India, an epoch-making work on Indian land mol-
lusks, also indispensable in the study of all Orient^tl and Ethi-
opian faunas. The various groujjs of Zonitida^ and the land
oi)erculat€s especially were worked out with great acumen and
illustrated j)rofusply with his own drawings. He also con-
THE NAlTII.rs. 31
tributeii the volume on Testacellidii' and Zonitidie in 77u;
Fauna of Jiritish In din.
He retired from the Survey in 1877 and spent tiic hist
forty yeare at Noro, near Godalmiufr. Enf^hmd.
"Of a most attractive disposition and sin<jridar charm of
manner. Goihvin-Austen \\}is much beloved by his contem-
poraries, most of whom lie lon<; oul lived." '
W. H. GOLISCH.
California collectors have been greatly saddened by the
death of William li. Goliseh, which took place in his 54th
year at his home in Los Angeles, April 18th, 1924. Mr. Go-
liseh had always been of a frail constitution and for the past
three yeare had found his activities increasingly limited by
physical weakness. About eight years ago he and his wife
gave the large conehological collection which bears their name
to the Southwest ^luseum, and since that date much of their
time and strength have been devoted to so installing and
arranging their shells that they might be both a pleasure to
tiie hundreds of casual visitors to the Museum and a help and
satisfaction to students. That they were succeeding is the
testimony of those who have had an opportunity to compare
their collection with those in other cities.
Mrs. Goliseh expects to continue the work which her hus-
band was obliged to leave unfinished, feeling that there could
be no better memorial to this beloved collector than the com-
pletion of the work which he loved and began.
To say that Mr. Goliseh was a man of high Christian char-
acter, holding both the respect and affection of all who knew
him, is perhaps supei*f1uous, but those who have been asso-
ciated with him for so long are unwilling to pass over this
last ojiportunity to pay tribute to his personality and to his
achievements in spreading conehological knowledge and in-
terest.— Southwest Museum Conchological Club.
1 .\ fuller sketch of Godwin-Austen's life, with ywirtrait, may be
fouml in The Ibi<i, for April, 19i!4.
32 THE NAUTILUS.
NOTES
CoQuiNA Bouillon. — "When at Uaytona, Florida, a man re-
marked in the course of conversation that coquina made a
delicious soup. Having lived for several years at St. Augus-
tine, Florida, Avhere cocjuina was a term applied only to the
shell-rock, of which Fort Marion and some of the older houses
are built, the recipe sounded far from appetizing. I had
heard of bouillon made from Danax variabilis, and this was
really what the man meant, for he said they were tiny clams,
found very abundantly on the ocean beach at low tide. On
my return I looked into the matter and found that the Span-
ish word coquina means shell-fish in general, Avith no reference
to the shell-rock. Therefore the use of the word for the shell-
rock at St. Augustine and other parts of Florida is purely a
local term. On the other hand, the word Donax has, for one
of its meanings, a kind of shell-fish.
There was an old man of Daytona
Who said they made soup of coquina
It gave us a shock
When we thought of the rock
Tlien we learned that Donax was coquina. — C. W. J.
MaRGARITANA MARGARITLPERA LiNNE IN LABRADOR. — FrOm
Rev. W. W. Perrett of the Moravian ^Mission in Ilopedale in
Labrador I duly received a number of Marijarilaiui niargariti-
fera L., found in a river in the neighborhood of Hopedale.
The specimens are of tlie common type; the biggest shell is
120 mm. in length. 1 tliink tliis is the fii*st record for Labra-
dor. Hopedale is situated in 55.5° N. lat., 60"' W. long. —
Hans Schlesch, Copenhagen.
A Ti.\K(;i-: Pkctkn P>Ki).-^ri may be of interest to the readers
of the Nautilus to Icai-n of the recent discovery of a bed of
Pcctcii ni(i(ff]Ui)ii(us ((iiiieU fi-oni which fishermen witli
flounder dredges are taking hundreds of barrels a day. Tlie
locality is about ten miles northea.st of Race Point, Cape Cod,
on the eastern edge of Sf illwagen 's Hank", in about "20 fathoms.
THK NAUTILUS. 33
Tlie fisliennon say tlic bed is less than a mile scpiare hut tiie
shells are so plentiful they are oblijiri^l to tie up a part of the
net to prevent getting!: too many at one haul, dragf^ing only
ten or fifteen minut<?s. Two specimens sent me measured G'/o
inches each, and were from a lot of 100 barrels, taken by one
boat that day. One barrelful in the shell yields about two
gallons of "eyes," as the fishermen call them, the so-called
eyes being the adductor muscle. These they sell for $3 per
gallon. There are some twenty boats engaged in this industry.
They get from 50 to 150 bbls. a day, and at this rate miLst
soon exhaust the supply. This is the first large bed of P.
niagellanicus discovered north of Cape Cod, although beds of
this species are found south of Cape Cod, even as far as off
the coast of New Jersey. Single specimens are often pulled
up on fishing lines along the coast, indicating beds not yet
exploited. — J. Henry Blake.
The varietal names of Polygyra profunda (Say). — In
1899 (Terr. Moll. .Alieh., p. 12) 1 described the form of this
species without bands as "var. albcr' and incidentally re-
ferred to the form with coalescent bands as var. '' unicolor''.
In 1876 Walker & Beecher (Proc. Ann Arbor Sci. Ass., p.
44) listed without description a ''Helix multUineata var.
albina".
In 1878 Witter (Quar. Jour. Conch., I, p. 384) described
the bandless form of muliUinrata as "var. olh(r\ Walker &
Beecher's "ulhina" being without any description was a mere
"nomen nudum" and cannot be recognized. Witter 's desig-
nation is therefore entitled to stand.
Under these circumstances my use of "alba" as a varietal
name for P. profunda was invalid and I would therefore pro-
pose for the bandless form of that species the varietal name
of cfasciata.
In 1868 Currier (Shell-bearing Moll. Mich., j). 5) and in
1881 DeCamp (Shell-bearing Moll. Mich., p. 6) li.sted "J/rso-
don multUineata unirolor,^^ and in 1894 (R<^v. IMoll. Fauna
Mich., p. 16) I cat-alogued a '^ Polygyra inullilinrata unicolor."
In none of these papers was there any description of the fonn
34 THE NAUTILUS.
thus designated, and being nude names they cannot be held
to prevent the subsequent use of " unkolor'' as a varietal
name in Polygyra. It follows therefore that Polygyra pro-
funda unkolor is entitled to stand as the name for the form
of that species with eoalescent bands. — Bry.vnt W.vlker.
PUBLICATIONS RECEIVED.
Notes on Some Austrauax Cassis. By C. Hedley. (Rec-
ords Australian Museum, vol. 14, pp. 46-48, pi. 8, 1923.)
Ox A Thal.\ssoid Element in* the Australlvn ^Iolluscan
Fauna. By C. Hedley. communicated by C. Oke. (Victorian
Xatur., vol. 40. pp. 75-77. 1923.)
Studies on Australian Mollusca. Pt. 14. By C. Hedley.
^Proc. Linn. Soc. X. S. Wales, vol. 48. pp. 301-316. pi. 30-33,
Oct. 1923.1 Three new species are described, with beautiful
illustrations of Umbrae uliim hotankutn n. sp. and Tdhy:^
angasi Sowerby.
Gymnosomatous Pteropoda from Friday Harbor, "W.vsh-
ENGTON. By H. P. K. Agersborg. (Ann. Sci. Nat. Zool., 10
ser., VI. pp. 391-401, 1923./ Two new species are described.
The Morphology of the Xudibr-vnchiate Mollusc Melibe
LE<:iNiNA (GoL'LD). By H. P. K. Agersborg. (Quart. Joum.
Micr. Sci.. vol. 67, pp. .507-592. Dec. 1923.) A ver>- exhaus-
tive treatise of this interesting nudibranch, proving that Chio-
raera Gould is a sv-nonj-m of Mdibe Rang.
New Pe.\rly Fresh-water Mussels from Mexico xsd
Urugu.vy. By Wm. B. Marshall. (Proe. U. S. Nat. Mus.,
vol. 63, pp. 1^, pis. 1-3, 1923.)
Tekti.vry Mollusks of the Genus Orthaulax from the
Republic of Haiti, Porti* Rico and Cuba. By Wendell R.
Woodring. (Proe. U. S. Nat. Mus., vol. 64, pp. 1-12. pis. 1-2,
1923.)
THE .\ALTILL-5. i-J
Two New Freshwater Snails fsom Michigan. L.. ^^^^
L. Winslow. (Occas. Paper Mas. ZooU Uoiv. ilich_ no. 145,
pp. 1-4. pi. 1, Dec- 1923.;
Xew Molll"Skan Names. By Truman H. Aldrich. (Pitw.
Biol. Soc. Wash., vol. 36. p. 199. Dee. 19*2:3. ' Uyi
n. n. for C. d<illi Aldr. 1594. not Cossmann 1^ --'jca
eo^mitki n. n. for C. smitki Aldr. 1886, not Sowerby 1n?1.
Two New Santo Domixgax Land Shells. By E. G. Van-
atta. f Proe. Acad. Nat. Soi. Phila.. vol. 75. pp. 359, 360.
1923. >
Notes ox Molluscax Nomi:ncl.4.ture. By W. H. DaU.
(Proc. Biol. Soc. Wash., vol. 37. pp. 87-90. Feb. 1924.
Teee Attachmext of Oyster Larvjc By Thorlow C. Nel-
son. Biological Bulletin. March. 1924, p. 143. 1 The foUow-
ing summary is given : " " Full-grown larvae of the oyster
shortly before attachment move over an appreciable area of
solid surface, testing it oat with the foot. Actual attachment
is preceded by circling movements, the larva finally coming to
rest with the left valve held in contact with the substratum
by the foot. The suggestion of Stafford. 1913. that the foot
is the organ of attachment has be«i confirmed by direct ob-
ser\-ation. The use of the foot and of the mantle during fixa-
tion is here described. The circling movements of the larva
while crawling over a substratum not only aid it in obtaining
a favorable foothold, but probably are also instrumental in
producing a fairly even distribution of the spat."
FossiLR-ii Cat-vlogus. I, Animalia, editiis C. Diener. Gas-
tropoda extramarina tertiaria, 1923. In 6 parts«. pp. 1862.
By W. Wenz. The literature of fossil non-marine m: - - -
so widely scattered that great labor is required to lea: .. --:
is published on the Tertiary histon.- of any family or genus.
There has been no general guide to the subject since Sand-
bergers monumental work, now about fiftj- years old. In the
FossQium CcUaloffus the references and synonymy of each
species are given in full, with its horizon and localities. How-
36 THE NAUTILUS.
ever, it is far from being a mere compilation of names ; only
those acquainted with the subject will recognize how much
excellent work has been done in the synonymy, in modernizing
the classification and in correcting generic references. Dr.
Wenz was well equipped for the task by his previous studies
on the same subject, and Me congratulate him on the comple-
tion of this standai'd work, which will be of value not only to
palaeontologists, but also to all interested in non-marine mol-
lusks.— H. A. P.
Land Mollusca from the Soloman Islands. By William
F. Clapp. (Bull. Mus. Comp. ZooL, Cambridge, LXI, No. 11.)
1923. Based upon specimens collected by Dr. W. ]\I. Mann.
About half of the 48 species found proved to be new. Among
the novelties are three species of Placostylus, several Crystal-
lopsis, TrochomorpJia, etc. Perhaps the most interesting is the
snail for which the new genus Cryptacgis is proposed. The
large, thin, subglobose shell is completely coveretl by the
mantle, which is eoncrescent over it. It is regarded as a
highly specialized offshoot of Papuina or some closely related
genus. In many cases anatomical details are described and
figured, and there are five excellent plates of the shells. —
H. A. P.
Mr. Charles Hedley has resigned his position as Principal
Keeper of Collections, Australian Museum, after thirty years'
service. He has been appointed Scientific Director of the
Great Barrier Reef ('onnuittee, a body formed under the
auspices of the Royal Geographical Society of Australasia,
Queensland Branch, to engage upon research jiroblems in rela-
tion to that great natural feature of the Queensland coast.
Mr. Iledley has also been recently elected an Honorary Fellow
of the New Zealand Institute. — Austnilian Zoologist.
THE NAUTILUS. XXXVIll
1.2. NEPHRONAIAS EI.V/E WAI.KKK
3.4. ANonoNTITKS Fl.UCKI WAI KI-.K
Fhe Nautilus.
Vol. XXXVIII OCTOBER, 1924. No. 2
ON THE EIO GRANDE
BY JAMES H. PERRISS
In the last days of Febniaiy, this year, snow-balling led all
Texan sports iu the Peeos-Rio Grande region. To the native-
bom it was their first snow — thus a warm game and full
teams. Occasionally the banks were too deep for the Fords.
At Alpine, however, by tlie time our car had been refreshed
and tried out, the roads were open, and for the next two
months the weather-man was kind and genial, from the Big
Bend to the northeastern corner of Texas.
This was an expedition of only one. Mollusks, cacti and
ferns were the real purj^ose ; pleasure, health, and a few other
things merely incidental. Eleven rattlesnakes were included
in theee incidental features, and some good things botanical
for a public park collection. The journey from Alpine to
Terlingua, in the depths of the Big Bend of the Rio Grande,
was over the route scratched by Dr. Pilsbry and the present
writer in a former (1922) expedition, an account of which we
have in preparation. No investigations were made this time
until new ground was invaded southwest of Terlingua. Here
the bones of Bulimuliis alternatus hesperiiis were very plenti-
ful in the sands of a low plain frequently and recently flooded ;
in its nakedness quite another Death Valley.
In the north slope of a mountain which to the south forms
38 THE NAUTILUS.
the north wall of the Grand Canyon of the Rio Grande, dweUsj
one of the larger Holospiras. Sad to relate, that particular
catch was lost in the mail, thus not identified. Again Buli-
mulus a. hesperius was collected in the brush of the south wall
of the river at Langtry, and upon the higher plains of the
Pecos.
Eastward from Marathon (on the Southern Pacific R. R.)
the El Paso-San Antonio highway, for sixty or seventy miles,
is confined to an ancient valley or channel between ranges of
low mountains, and thus one of the most pleasing prospects.
It was only twenty to forty minutes from car to mountain-top.
Twenty live Lysinoes in one gulch, the very one the collectors
have been searching for for some forty years, because a
"bone" had been found near a prairie-dog hole at Alpine.
Holospira, Bidimulus, and ferns were plentiful.
At Sanderson, in this valley, Polygyra and Helicina broke
in, and continued the remainder of the journey. At the mouth
of the Pecos Euglandina was abundant (and dead). Only
two living ones were found in three half-days of especial dig-
ging. While gathering other shells, the dwelling place of this
member of a wet-land fauna was accidentally discovered in the
highest and drj'est hills. Where the upper layer of limestone
is separated from the main floor one or two feet, the space
filled with broken stone, soil and rubbish, will be found the
home of Euglandina singlcyana. No bones or fragments of
this species were scattered about tlie surface witli the remains
of other species, and the shells in their home were unbroken.
Thus evidently the animal is not toothsome for mice or the
rock cat. Death comes in tlie natural way. The live shells,
agate-yellow in color, polislied, transparent, witli faultless
lines of moUusean areliitecture, a full-grown s})ecinien lightens
the heart, bestirs the energy, of the collector.
Both Pilsbry and I had a profitable experience here at
the Pecos' mouth in 1903. The limestone bluffs, around five
hundred feet above the river bed, present an excellent eleva-
tion for rarities in conchology and botany. Here also a close
acquaintance with our largest rattlesnake was enjoyed.
Nolhol(r7ia srhaffticri, one of the rarest American ferns, thrives
THE NAUTILUS. 89
here under the upper, overhanging sheets of limestone. A
delioato phmt. it prospers best in tlie tine diiKl accinnnlating
upon the tloor of its oave-like iionie. Posing upon a common
level with liis victims, the collector upon hands and knees
crawls about in the cacti and prickly brush with l)ecoming
humility and caution. Thus came about this introduction to
a rattler of seven inches girth. Happily, although the rattler
was coiled for emergencies and, nose and nose, we were less
than two feet apart, his snakeship was exceedingly fat, thus
good-natured and patient. However, to get him permanently
into an army knapsack required the better part of an hour's
diplomacy. It was worth it. With this, my collection contains
all of the American poisonous snakes except three.
A warm shower in the middle of April was a grand spring
opening for the Bulimulus tribe. Then B. alternatus hesperms
climbed tlie fence-posts of the Pecos, the cacti and the brush.
Some of the shrubbery looked like stands of the white snow-
berry in fruit. A few days later B. dealbatns pecosensis ap-
peared at the High Bridge, and during the remainder of the
journey BulimuLiis bushes were in full fruit, all species of the
region being represented. While the highway from the Pecos
River is fairly level for southern Texas, on into Waco, and
one brushy tract like another, each collecting ground contained
a Bulimulus species, variety or color unlike its predecessor.
A mile perhaps below the Laguna postroffice the hills upon
the south bank of the Nueces, high and well timbered, with
precipices and slides, seemed to warrant a little wading.
CheUanthes leucopoda, another of the rarest ferns, had been
found within a hundred miles or so of that beautiful scenerj'.
In fact, since returning home, the original discovery of the
species was found to be Uvalde Canyon, a familiar name in
that part of Texas. Not only this fern thrives in that par-
ticular Nueces bluff, but also Atiemia mexicaaia, another Texas
rarity in ferns ; also the recently described Holospira goMfussi
anacacJietisis Bartsch ; and three species of cacti heretofore
unknown to me.
Possibly in the headwaters of the Neuces and the low range
of mountains from New Braunfels, Fredericksburg and Keer-
40 THE NAUTILUS.
\TLlle, with a western terminus somewhere near the Pecos
drainage, both botany and conchology may contain something
of great interest. The Great Bend of the Rio Grande also
seems tempting. Our two raids there scratched the west side of
the Chisos mountains, and a sight-seeing trip to the Grand
Canyon only. The region is as accessible as any, considering
desert conditions. From Marfa, Alpine and Marathon via
Terlingua to the river is about seventy miles. From the Pecos
to Marfa a little over two hundred miles. From ]\Iarfa to
El Paso another two hundred. Here are mountains 9000 feet
in altitude, and desert conditions quite unlike other better
known deserts. Limestone almost everywhere, the soil fertile,
well covered with vegetation ; population American stock-men
and Mexican gardeners; a few villages and quicksilver mines.
With a Ford or pack train is not this about the best of our
unexplored regions?
In the following list only the species taken in the Big Bend
and along the highroad (which roughly parallels the Southern
Pacific Railway) as far east as the Nueces River are recorded.
The few lots taken in central Texas are from well-known
places.
List of Species, by H. A. Pilsbry and J. H. Ferriss.
Polygyra texasiana texasensis (Pils.). Alpine, Langtrj',
mouth of the Pecos, Devil's River, east of Brackettville and
on the Nueces River. In the western counties this form, which
is smoothish on the upper surface as well as the base, replaces
the typical P. texasiana, which is ribl>ed above. It was first
described as P. texAisensis Pilsbry, Nautilus XVI, p. 31 ; later
we found a smaller and even smoother form in the Devil's
River region, which we described as P. tcxasiuna hjfpcrolia,
Proc. A. N. S. Phila., 1906, p. 128. The large series t^iken
this year shows intergrades in size, scul])tiire and size of um-
]>ilicus between these forms, which cause us to unite them.
Thi/s'(mo]>hora hornii (Gabb). Devil's River and Nueces
River; scarce in tlie drift.
Bulimulus altemaiics hespcriits, u. subsp. {Bulwudus alter-
THE NAUTILUS. 41
nut US ytuirur. Western Form, Pilsbry and Ferriss, Proc. A. N.
S. PhUa., 1906, p. 140, pi. 7, figrs. 1-12; p. 132, %. 5a.)
The sliell averages larger than B. a. nuiruc; spire more
strictly conic; white or pale brown, uniform or with brown
streaks wliich are never ragged; aperture very dark; colu-
mella not toothed. Length 30 to 44 mm.
In working over our collection of the expedition of 1903
the characters of this race were noted, but we did not then
name it as separate from marue. A considerable amount of
additional material from localities mentioned below, and larger
series of the true moruv from the lower Rio Grande valley,
collected by one of us, serve to emphasize the constant differ-
ences between the form of the high western country and maruE
of the lower valley. We take as type of hesperius a specimen
from the east side of the Pecos at the High Bridge, measuring
34 mm. long, 17.4 mm. diameter. Additional localities of 1924
are : a valley just north of the canyon of the Rio Grande, W.
of S. from Terliugua; Langtry; west of and at the highway
bridge over the Pecos, near its mouth.
Bulimulus dealhatus pecosensis P. & F. Quite abundant,
and variable in shape and color. Mouth of the Pecos in sev-
eral places, and about halfway between there and Del Rio;
near, west of, the Nueces River below junction of two dry
branches, and in the river drift.
Bulimulus dealhatus ragsd<ilei Pils. Alpine, Housetop Mt.,
Sanderson, small hills west of St. John ranch, Langtry, Devil's
River, and about 3 miles east of Del Rio.
Locally variable in shape and degree of striation; in some
cases approaching B. d. pecosensis. Sometimes very small,
do\\ni to 16 mm. long.
Bxdimulus, n. sp. A peculiar rather long Bulimulus, very
smooth and glossy, brown with many whitish streaks, was
taken at Sanderson, but the single example is broken, and just
what its characters are remains uncertain. It must be looked
for again.
Holospira goldfussi anacachensis Bartsch. Nueces River.
42 THE NAUTILUS.
Occurs both larger and smaller than the measurements given
in the original account; from 10.7x3.5 mm. to 15x3.9 mm.
Holospira roemeri (Pfr.). Alpine and Housetop Mt., slen-
der specimens; mouth of the Pecos; Devil's River; Nuec«s
River. Specimens from the mouth of the Pecos measure :
Length 14, diam. 4.3 mm. Length 17.7, diam. 3 mm.
And from the Nueces River :
Length 12.5, diam 5 mm. Length 14, diam. 4 mm.
Length 12.7, diam. 4.9 mm. Length 18, diam. 4.3 mm.
Length 13, diam. 4.6 mm.
The longest of these is strictly cylindric ; the shortest is
widest in the middle, while the others are widest in the upper
part.
Euglandina svngleyan<i (W. G. B.). Found on both sides
of the Pecos, near the highway (see above). One broken
specimen measures 20.3 mm. in diameter, and a fragment
indicates a still larger size, thus surpassing the central Texas
specimens.
Helicodiscus eigenmanni Pils. Drift of the Pecos, Devil's
and Nueces Rivers.
Punctmn pygmceum (Drap.). Nueces River.
Polita indentata umhilicata Ckll. Pecos, Devil's and Nueces.
Polita dalliana roemeri (Pils. & Ferr.). Nueces drift.
Pseudohyalitm minuscula (Binn.), P. singleyaim (Pils.)
and P. niimmus (Van.). Devil's and Nueces drift.
Eucoiiulus cher sinus trochulus (Reinh.). Devil's and Nue-
ces river drift.
Succinea lutcola Gld. Alpine, Langtry, mouth of the Pecos
and Devil's Rivers.
Succinea avara Say. Devil's and Nueces Rivers.
Gastrocopta cantracta (Say). Devil's and Nueces Rivers.
G. pentodmi (Say). Nueces River.
G. pcUucida- hordracrUn (Pils.) and G. criftaia (P. & V.).
Pecos, Devil's and Nueces drift.
Pupoidcs niargiiiata (Say). Same localities.
Vertigo ovata (Say). Nueces drift, one specimen.
TIIK NAUTILUS. 43
VaUania j)erspcctiva Sterki. Peeos, Devil 's and Nueces drift.
Lxjmmra {Galha) parva Lea. Devil's and Nueces Rivers.
Planorbis trivolvis Say and P. antrosus Con. Nueces River.
Plunarbis dU<itatus Gld. Nueces drift.
Pl-atwrbis cams Pils. & Ferr. Devil's and Nueces drift.
Planorbis Ikbm^nni Dkr. Pecos, Devil's and Nueces drift.
PUmorbula obstructa (Morel.). Same localities.
Physa sp. Pecos and Nueces Rivers; small specimens only.
Paludestrina protea (Gld.) and P. seenuinni (Fdd.). Nue-
ces drift.
Cochliopa riograndensis Pils. & Ferr. Pecos and Nueces
Rivers.
Helicina orbiculata tropica Pfr. Pecos, Devil's and Nueces
drift; also east of Brackettville.
SOME OLD PLEUROCERIDS AND A NEW ONE
BY CiVLVIN GOODRICH
In the summer of 1923 Mr. W. J. Clench and Mr. L. E.
"Wehmeyer made a collecting trip by automobile that began at
Ann Arbor, Mich., and took in parts of Indiana, Illinois, Mis-
souri, Arkansas, Tennessee, Mississippi, Alabama and Florida.
The Pleuroceridae taken on the journey — described by the
travelers as the Voyage of the Asthma — were placed in my
hands for identification.
GoNiOBASis LiVESCENS (Menke). Wabash River, Ind., two
miles west of Huntington, also two miles west of Peru ; Pipe
Creek, seven miles west of Peru.
This is a stout, bulbous form, varying little in the upper
"Wabash drainage. I have it from the Wabash, Logansport;
Little Wabash, Huntington; Salamanie, Montpelier; Eel,
North Manchester; Tippecanoe, Warsaw; Deer Creek, near
Delphi ; Coal Creek, Veedersburg. This is on a line running
southwest across Indiana. The form is the common one of
the Maumee River system of Lake Erie and occurs in the Rai-
sin, Huron and Clinton rivers of eastern Michigan ; also in
44 THE NAUTILUS.
the more protected parts of the Niagara River and in several
of the streams of Ontario. The form is noticeably different
from the species in Lake Erie, the type locality of livescem.
Anthony described a melanian from the Wabash as M. cuhi-
coides.^ Material from Huntington was identified by Call ^
as this species. The types, which I have examined, are two
specimens sharply angled on the body whorl. I am sure that
as regards livescens of the streams this is only an oddity, rare
and of no specific value. However, in Lake Erie are deep-
water forms that are carinate to the last whorl, the character
being constant. Such shells w^ere dredged a few years ago by
students of the Ohio Biological Station at Put-in-Bay and
there is a beach in Ottawa County, Ohio, upon which these
carinated forms are thrown, doubtless by the severer storms.
Call says he did not fiiid cuhicoides {livescens) in the Wabash
below Lafayette. It is rather strange that Daniels, who col-
lected several times at Lafayette, did not find any Goniobases
there at all.
Other synonyms of livescens, whose tji^es I have exara-
iried, are:
Melania niagarensis Lea, March, 1841.
Melania napella Anth., Dec, 1850.
Melania cuspidata Anth., Dec, 1850.
Melania elata Anth., Dec, 1850.
Melania occulta Anth,, Feb., 1860.
Go-niohasis milesii Lea, June, 1863.
Goniobasis lithasiodcs Lea, June, 1863.
Pleubocera acuta Raf., Wabash River, two miles west of
Huntington, Ind. ; Pipe Creek, seven miles west of Peru, Ind.
This is a common associate of Gouiobasis livescens in north-
em waters, though apj)arently it docs not go as far north as
livescens. Call speaks of finding this species (commonly
known as P. suhulare I^ea) by thousands at Lafayette "on
the muddy banks of the river within the city limits." It is
a large form here and specimens t^iken by Daniels were sent
' Proc. Acad. Nat. Sci., Phila., Feb., 1860, p. 60.
* Indiana Geological Report for 1899, p. 482.
THE NAUTILUS. 4")
out as elevatum Say. There is some question as to what the
true elevatum is inasmuch as ISay himself appears to point
out its resemblance to canaliculatiitn. This would read most
of the elevatum out of the cabinets. A complex of Pleuro-
ceridffi occurs at Lafayette. It is extremely difficult to tell
where acuta leaves off and forms of the canaliculatuin-
undulatum group begin. Possibilities are hybridism and a
merging of species hitherto unsuspected.
GoNiOBASis POTOSiENSis (Lea), stream three miles south and
a second stream four miles south of Potosi, Washington
County, Mo. ; creek two miles west of Shepard, Iron County,
Mo. All tliree localities in the drainage of the Meramec River.
Nearly all the specimens taken are juvenile, but agree with
Lea's meager description and one individual has exactly the
measurements set down for the types. The species is close to
pleheius Anth., common iu the Ozarkiau region, differing from
it principally in having a much more delicate texture, in
being deeply and constantly cariuate and in having the outer
lip regularly rounded with no angulation whatever. Even
quite old specimens would probably be less heavy than the
average pleheius. A possible synonym is Goniobasis ozar-
kensis Call, 1886.
GoNiOB.vsis PLEBEius (Auth.), two miles southwest of Hous-
ton, Texas County, Mo. (Gasconade River drainage) ; two
miles southeast of Hardy, Sharp County, Ark, (White River
drainage).
These forms are identical with those taken by CaU, Hinkley
and Brown in the same region. The "center of population"
of this species appears to be in Missouri, south of the Missouri
River. It is plentiful in northern Arkansas, and there appar-
ently it reaches its largest growth. Certain Ganiobasis from
Des Moines, Iowa, may be as.signed to plcbeius. Dr. Brvant
Walker has specimens from Fort Ilill, 18 miles south of Ft.
Gibson, Indian Territory. This would be in Muskogee County,
Okla., the drainage of the Arkansas River. I take the types
of Melania haleiana Lea, coming from Alexandria, La., and
now in the National Museum, to be young Pleurocera acuta
46
THE NAUTILUS.
Raf. But vnth. this type lot are some juvenile Goniohases,
almost indeterminate but nearer to pleheius than to anything
else. If they are this species, the range of pleheiiis is from
Iowa to Louisiana, westward into Oklahoma and probably into
Kansas.
GoNiOBASis CLENCHi, new species.
Shell: Elongate, rather thin, Dresden-brown to chestnut-
brov^^l, the type without color bands. Sutures deeply im-
pressed. Apex eroded. Tlie first of 6% whorls remaining is
flat in the space between a carinated periphery and a carina
just under the suture; the other wliorls become increasingly
convex to the last one. Sculpture striate-undulate, reticulate ;
low plicae are crossed by revolving lines, a tiny point of shell
substance rising at each junction of plicae and horizontal lines.
This sculpture continues to the peripheiy of the last whorl,
below which are four sharply-defined revolving lines and one
microscopic line. Growth lines are fine, not very regular.
Aperture ovate and of moderate size, showing the exterior
sculpture through the shell substance. Columella slate-colored,
thin, narrow, with a twist or fold near the center. Outer lip
decidedly incurved from suture to below the periphery. Sinus
not large, but distinct.
^^^
2
Fios. 1-3.
Goniobasis ch'iichi
Fig. 4.
Teeth of G. clenehi.
Operculum: Tliin, reddish-brown, roughly triangular; alti-
tude 31/i mm., diameter 3 mm. Nucleus (leju'essed. Nuclear
whorls sharply-rnarkcd. widcly-coih'd. Striie dee]^ly iiioisod
and sliowiiig tiie iTgular growtb and clumge of i)osition of tlie
nuclear whorls with rehit.ion to the apex. Apex blunt. Left
margin curving, very sliglitly sinuous, thickened ; right margin
broadly curved, exceedingly thin ; ])asal margin rounded.
THE NAUTILUS. 47
MeasTirements of shell : Altituile, 24 mm. ; diameter, 8 mm.
Aperture — altitude, 7:^4 mm.; diameter, 4 mm.
Type Uvality : Clioctawhatchoe River, Newton, Dale County,
Ala. Shells taken "on rocks and alon^? bank on ledges; when
in middle of river on rocks in one to tliree feet of water; the
Pleurocerida^ crawling there on a tine deposit of silt or very
soft mud." Collected by W. J. Clench, July 23, 1923.
The number of whorls of an uneroded adult would probably
reach ten. On young shells the first whorls have no sculpture
save two carina?, one close to the suture, the other at the
center. The upper carina sometimes resembles a row of
minute beads. The periphery of the whorls of juveniles is
angular, the base tending to flatten. In a ([uite old shell the
columella is pinkish-white rather than slate-colored and, be-
cause of the addition of callus, the fold is apparent only
under the hand-glass. Most of the shells are bright and shin-
ing. Variation in color, form and sculpture is slight. Of 78
shells that were cleaned of foreign material, six have a nar-
row, dark band on the base ; all of these are young.
The radula appears to differ slightly from that of Melania
depygis Say £is pictured by Troschel and from that of Plenro-
cera elevatiim Say, drawn by F. C. Baker. The formula of
the denticles is :
[l2^7J^ 4^3 — 4-1-1 + 3 — 4^4 ^[7 ^ 12J
Goniohasis clenchi, belonging to the group of catenaria Say,
has some of the aspect of that species, notably the sculpture.
It is, however, a more narrow shell, is not so strongly cari-
nate ; the lip usually is more deeply incurved ; the body whorl
is broadly rounded whereas in catenaria it is subangulate at
the center. Clenchi has the distinctive operculum of catemiria,
which jdso I have found in G. arachno-idca (Anth.) and G.
troostiana Lea, probably geographical outliers of the catenaria.
group.
Herbert H. Smith recognized this species as new. His
localities for it were Choctaw hatchee River, near Pinckard,
Dale County, and the same river near (Geneva, Geneva County,
Alabama.
48 THE NAUTILUS.
The pictures of the shell and operculum were drawn by
Miss Miua Winslow ; that of the radula by Mr. Clench,
GoNiOBASis DOOLEYENSis Lea, Choctawhatchee River, New-
ton, Dale County, Ala.
The Alabama specimens are somewhat smaller than indi-
viduals from Georgia and the plicate sculpture is less pro-
nounced, but in all essentials the shells are the same. The
types of dooleyensis are from Dooly County, Ga. Walker^
cites it from Early and Baker counties, Ga. ; from Chipola
River, Fla. ; all in the Apalachicola drainage ; also from Econ-
fina Creek, Fla., flowing direct in the Gulf. Every one of
these streams, including the Choctawhatchee River, come close
to one another near their mouths, and probably there has
been interchange of fauna.
GoNiOBASis sp., Choctawhatchee River, Newton, Dale County,
Ala.
I am unable to recognize this species -vnth the help of de-
scriptions and illustrations published. It may possibly be the
form which Lea included under G. elliottii as coming from
Uchee and Little Uchee rivers, Ala. The shell is slender,
smooth, mostly greenish-brown, sometimes dark yellow or
nearly black. The apical whorls are carinate, sometimes with
two or three or four incised, microscopic, revolving lines.
The outer lip is slightly sinuous, the aperture elliptical, the
sinus distinct and extended. The shell has a number of nar-
row bauds more often than not.
GoNioBASis CATENARIA (Say), Itchtuckuee River, one mile
eeist of Hildreth, Fla. (Suwanee River drainage).
This is the form which appears in ohl collections as M.
2)apUlosa Anth.
•Nautilus, XVllI, Aj.ril, 1905, p. 134.
THE NAUTILUS. 49
MABINE MOLLUSCA OF THE BRIDOEPOBT, CONNECTICUT, BEOIOH
BY ARTHUR JACOT
The following Mollusca were collected at the various beaches
near the city of Bridgeport, Connecticut. Although the habi-
tats are varieti, only a small number of species Avas secured.
This meagemess of tlie fauna may be ascribed, though prob-
ably not entirely, to proximity to the city. Both Outsell and
Mitchell have reported (Bureau of Fisheries Reports) on re-
sults to oysters and other marine animals of oil, copper and
tar contamination of waters even some distance from the cities.
Slightly lower salinity is probably also responsible — though
but slightly so. ]\Iilford Beach is close to the mouth of the
Housatonic River.
Area (Scaph-arca) campechiensis pexata Say. Long Beach,
Bridgeport; generally distributed, fairly common.
Area {Sea pharca) transversa Say. Long Beach ; less common.
Osirea virginica Gmelin. Common everywhere, the dominant
species, forming banks, as at Milford Beach.
Peeten {Plagioctenium) gihhus horealis Say. Long Beach;
not common.
Ancnnia simplex d'Orbigny. Generally common.
Mytilus edulis Linne. Abundant everj-^vhere, especially so at
Stratford Beach.
Mytilus edulis pelhccidits Pennant. Occasional among the
others.
Modiolus {Brachydontes) demissus plieatulus (Lamarck).
Long Beach, especially on the marsh side.
Pandara {Clidujphora) g&uldiaiui Dall. Fairfield Beach, Fair-
field. Occasional.
Lyonsia hyalina (Conrad). As the preceding.
Cardium (Laevicardium) mortoni Conrad. Very thinly scat-
tered.
Calloeardia {Agriopoma) nwrrhuaim (Linsley). Occasional.
Venus merccnaria Linne. Very few.
Venus merceiiaria notata Say?.
50 THE NAUTILUS.
Gemma gemm/i (Totten). Long Beach.; fairly common.
Gemma gemma- purpurea (H. C. Lea). Far more common.
Petricola plwladiformis Lamarck. Occasional.
Tellina (Angiilu^) tenera Say. Fairfield Beach; occasional.
Macoma, balthica (Linne). Long Beach, around the point.
Ensis directus (Conrad). Long Beach; fragments.
Spisida (Hemimactra) solidisslma (Dillwyn). Silver Beach,
Milford.
Mtdinia lateralis (Say). Milford Beach; fairly common.
My a arenaria Linne. Common most everywhere.
Pyramidella fusca (C. B. Adams). Not uncommon; IMilford
and Long Beach.
Odostomia (Menestho) trifida (Totten). Common generally.
Odostomia {Meiiestho) hisuturalis (Say). Milford and Long
Beach.
Polinices (Neverita) duplicata (Say). Few.
Polinices {Euspira) heros (Say). Long Beach; occasional.
Polinices (Euspira) triseriata (Say). Milford Beach; occa-
sional.
Crepidula fornicata (Linne). Generally distributed.
Crepidida glauca convexa Say. Common, all beaches.
Crepidula plana Say. Occasional, all beaches ; mouth of whelk
shells.
Litorina ohtusata palliata (Say). !Milford Beach; abundant,
common everywhere.
Litorina rudis (Donovan). Similar to previous,
Litorina rudis tenehrosa (Montagu). Long Beach, on marsh
side.
Lacuim viticta (Montagu). Milford Beach; occasional.
Lacuna vincta fusca Gould. Milford Beach ; common.
Euplcura caudata (Say). Fairly common and of general dis-
tribution.
Urosalpinx cvierca (Say). Commoner than the preceding.
Columbella (Anachis) avara similis Ravenel. Fairly common,
Milford Beach.
Columbella (Astyrv;) hnuiia (Say ) . Milford Beach ; abmidant.
Alcctri&ii [Hinia) vibcx (Say). Occasional.
THE NAUTILUS. L)l
Alecirian (Ilyatwssa) obsolcta (Say). Long Beach ; especially
around tlie west end, and eastward of Milford Beach.
Alectrion {2'ritia) trivittata (Say). Stratford Beach, abun-
dant, common generally.
Busycon carica (Gmelin). Silver Beach, Milford, and others;
occasional.
Busycan canaliculatum (Linne). Commoner than previous.
Melampu^ litwatiis Say. Abundant on salt marsh&s.
A comparison of this fauna with that found on the south or
ocean shore of Long Island shows a conspicuous absence of
the following genera: Nucula, Yoldia, Astarte, Divaricella,
Donax, Tagelus, SiliqiM, Corhula, TurhoniUa, Epitoivium,
Triphoris, Bittium, Actean, Tornatina, Cyclina. Although
some of these may later be found in this general vicinity, the
majority of them are species demanding conditions of full
salinity, clean ocean water, and clean sand. At Beaufort,
N. C, for instance, Donux versicolor was most plentiful on the
inlet shores. Once the inlet banks had turned inward far
enough to face the sound this species was lacking or rare.
There is, therefore, considerable difference in habitats whose
substratum conditions are identical but which are situated in
a sound or bay and on the ocean proper. In this connection
it would be interesting to determine how far eastward beyond
Long Island Donax may be found, as at Block Island, Martha's
Vineyard and Nantucket, before the cold stream from the
north would be felt sufficiently to change the temperature
conditions found along the south shore of Long Island.
TWO NEW NAIADES FROM NICASAGUA
BY BRYANT WALlKER
The following species were submitted to me by Rev. W. II.
Fluck of Great Kills, Staten Island, N. Y., for examination
and description, if they proved to be new. Both of them ap-
pear to be undescribed.
52 THE NAUTILUS.
Nephraiiaias elvce u. sp. PI. I, figs. 1-2.
Shell solid, broad oval, rounded before and behind, sub-
compressed, with a very low, nearly obsolete posterior ridge,
which scarcely affects the posterior margin and which is out-
lined by a shallow groove along its posterior side; beaks but
slightly elevated, eroded in the specimens seen; surface
slightly sulcate towards the beaks, much more so towards the
margins and covered with numerous fine striae very close
together, which are cut by radial impressed lines extending
from the beaks to the margins, which are deeper and further
apart anteriorly and posteriorly, giving them a festooned ap-
pearance, which is almost granular in the center of the disk;
epidermis dark brown, shading into yellowish around the mar-
gins, rayless; left valve with two strong, erect pseudocardinals,
which are nearly parallel, and two slightly cum^ed laterals;
interdentum rather long, narrow and rounded; right valve
with a single, strong, elevated pseudocardinal, the sides of
which are about parallel with a small, compressed tooth above
it, and one lateral; anterior muscle scars deeply impressed
and roughened, posterior scars much less so; pallial line dis-
tinct, with radiating sculpture especially on the upper side;
nacre bluish white, slightly iridescent posteriorly ; dorsal scars
numerous, deep and situated luider the interdentum.
Length 62, alt. 44.2, diam. 25.2 mm.
Type locality, the Wanks River or Rio Coco, Nicaragua.
Type in the collection of Rev. W. H. Fluck.
Mr. L. S. Frierson, to whom a photograph was submitted,
suggested that it was very close to Nephronaias rctkid^ata
Simp., if not identical with it. On sending similar photo-
graphs to Mr. W. B. Marshall of the U. S. National Museum
for comparison with Simpson 's type, he replied as follows :
"At first I thought your shell surely was Nephranaias reti-
culata Simpson, but when I came to examine the sculpture I
found that, although Simjison's shell is only about two-thirds
as large as yours, the sculpture on his is two or three times as
coarse. This put me on the fence as to whether yours is or
is not a new species, so I called Dr. Bartsch to my assistance
THE NAUTILUS. 53
and, between iis, we came to the conclusion that yours is prob-
ably different from Simpson's. Your shell is much more
round than Simpson's. Simpson's shell is very light chestnut
in color. I should hardly call the nacre of Simpson's siiell
lurid. It is really a dirty white witli a little iridescence near
the posterior portion."
It would seem, therefore, that the present form is well en-
titled to a name, even though a larger series might afford
connecting links.
There was originally another specimen, a little smaller than
the type, but otherwise exactly the same, which was unfor-
tunately lost in the mail.
At the request of Mr. Fluck, I have named the species after
his niece, Miss Elva M. Smith.
Anodantites flucki n. sp. PI. I, figs. 3-4.
Shell obovate, thin, inflated, slightly compressed by a faint
depression ext-ending from the beaks to the anterior basal
margin ; inequilateral ; slightly gaping towards the anterior
margin, but not posteriorly ; beaks moderately full, but little
elevated, the tips decidedly incurved; hinge line nearly
straight, angled at both ends where it passes into the anterior
and posterior margins; anterior end narrowed and rounded;
base slightly incurved at about the anterior third, correspond-
ing to the depression across the disk ; posterior end regularly
and somewhat broadly expanded, the greatest height being
behind the middle of the shell ; surface smooth with irregular
growth lines; epidermis greenish yellow, dark green on tlie
posterior slope and towards the anterior end, with faint nar-
row, green rays over the center of the disk; nacre light blue,
slightly salmon-color within the beak cavities, very iridescent,
faintly radiately sculptured.
Length 87.3, alt. 46.5, diam. 31.3 mm.
Type locality, the Wanks River or Rio Coco, Nicaragua.
Type No. 74610, Coll. Walker. Cotypes in the collection of
Rev. W. II. Fluck.
As suggested by Mr. Frierson from an inspection of a photo-
graph of the type, this species is evidently rather closely re-
54 THE NAUTILUS.
lated to A. nicaragiue Phil, described in 1848 (Zeits. fur Mai,,
XV, p. 130). It has never been figured or found since. Von
Martens (Biol. Cent, Am., 1900, p. 53G) believed it to be the
same as A. hridgesi Lea. If so, it is quite different from A.
fiucki.
Philippi gives the dimensions of his shell as 65 x 37 x 10 mm.
As von Martens (1. c.) suggests there is probably a t^-po-
grapliical error in the diameter as stated, which is quite in-
consistent with the diagnosis, which says "tumida". So far
as the length and height are concerned, the relative propor-
tions of Philippi 's type and that of fiucki are substantially
the same.
But Philippi states that his shell is gaping at both ends
and that the posterior end is "paullo altiore, oblique suhtru)i-
cata," neither of which specifications apply to this species.
Philippi applied the term "oblique truncata" also to two
other species, A. incarurn and V. valdivianus, both of which
are figured by Pfeiffer in Nov. Conch., Ill, pis, 103 and 105,
and these show distinctly what Philippi meant by this expres-
sion and that it does not apply to the species here described.
CALIFOENIAN HELICES
BY H, A, PILSBRY
HELMiNTnoGLYPTA FERRissi new species.
The shell is depressed, thin, umbilicate, the width of umbili-
cus contained between 7 and 8 times in that of the shell. Of
a pale greenish -buff tint with streaks of deeper green (mig-
nonette-green of Kidgway) ; above the periphery a rather
narrow chestnut-brown band without light borders. The
spire has a faintly pinkish tint. Surface is very glossy. The
embryonic whorls, after the first nearly smooth half-whorl,
are very densely, irregularly granulose. Later whorls have
somewhat sharp but very fine retractive striae. On the last
whorl these are (juite weak except near the suture. The
TIIK NAUTILUS. 55
whorls increase slowly at first, but the last widens rapidly, is
roundetl peripiierally and descends to the aperture. The aper-
ture is large, oval-lunate ; peristome thin, the outer and basal
margins slightly expanded, columellar margin dilated, cover-
ing part of the umbilicus.
Alt. 15 mm., diam. 27.2 mm. ; 5% whorls.
Above Tehipite Valley, middle fork of King's River, Fresno
Co.. California. Type 134224 ANSP. Found on going up a
zigzag trail northward several miles after leaving Tehipite
Valley and Dome, once in forest rubbish among rocks, again
at a crumbling clilf ; elevation probably around 7000 ft.
About 35 specimens were taken, but very few were in good
condition. It is a beautiful species, of a brighter green color
than any form of traski, and differing in the sculpture, both
of embryonic and later stages. The largest one taken meas-
ures 27.2 mm. diam., the smallest adult 24 mm.
Helminthoglypta UMBiLicATA (Pilsbry).
Epiphragm-aphora tudiculata umhUicata Pilsbry, Nautilus,
XII, June 1898, p. 22.
Epiphragmophora dupetithouursi cuastatm Edson, Nautilus,
XXVI, August 1912, p. 37 ; XXVII, August 1913, p. 37, pi.
3, figs. 13, 14.
Helix dupetiihmiursi var. concursus HemphiU, in collection
(Mus. No. 112425 ANSP.).
This is a form apparently related to H. dupetithouarsi
(Dh.), but lighter colored and more depressed, with a wider
umbilicus. Just w^hat relation it bears to other species will
doubtless be clearer w^hen it can be dissected, but for the
present I am leaving it as a species.
It was collected by Hemphill in the vicinity of Morro, San
Luis Obispo Co., thus at a quite low elevation. The unpub-
lished name he gave to the form is mentioned above as it
occurs in many collections. Edson 's locality was eastward, in
the Santa Lucia Range, at an elevation in the neighborhood
of 1500 ft. The original set of umhUicata was not localized
further than San Luis Obispo County. It has not been found
beyond the limits of this county.
66 THE NAUTILUS.
COLLECTING IN SOUTHERN FLOEIDA, THE BAHAMAS AND CUBA
BY D, L. EMERY
During the past suimner, with about two months of spare
time, I decided on a trip to the West Indies. I left St. Peters-
burg on July tenth for Miami, where I was joined by Mr. C.
C. Allen, a most ardent and thorough collector.
The east coast of Florida is a valuable field and a great
many species of both land and marine shells were added to
our collections. The hammocks in the vicinity of Miami are
rich with the Liguus, Di-jmiaeus and Helicina, while the so-
called reefs abound ydth. Polygyra, Urocoptis, Chondropoma,
Thysanophora, Microceramus etc. The Atlantic beach fur-
nishes us with a goodly number of Spirula and lanthina, and
the shores of the outlying keys jdelded Lucina, Loripes, Iphi-
genia, Strombus, Modulus, Cerithiimi and Neritina. The
shrubbery near Miami Beach fairly teemed with Cepolis, Dry-
maeus, Helicina and Cerion ijicamim, while on the grass we
found Succinea and in the canal Perna. On the jetties at the
mouth of the harbor we collected several species of Thais and
Nerita, and farther up were Tectarius, Echinella, Plauaxis,
Littorina and Siphonaria. One side trip from Miami was by
st^ge to Fulford, where in the sand thrown up by the dredge
in a new subdivision were may varieties of marine forms in a
fine state of preservation, which had been buried for many
years. Among tliem were some of the largest and finest
Lucina janiaicensis I have seen. The periostracum on them
was almost perfect while the hinge ligament was practically
gone. On the higher beach we foimd the same as at Miami
Beach, several good lanthina and Spirula. In the marshy
land were a quantity of the Auriculastriim peUucen^, mostly
dead, and Cyrcnoidca floridunu.
After spending all the time we thought we could spare, we
took passage on the S. S. Nassauvian of the Allan line for
Nassau. We spent about three weeks on the island of New
Providence with headijuarters in Nassau. I ^rill sUite here
THK NAUTILUS. 57
that the trip was not so much for all the species we could
gatlier as for those from half aii inch up wliich we could col-
lect in quantity for our exchange lists.
Nassau proper, with its rocky and sandy beaches, its rising
sunny slopes, lime rock reefs, and at the rear the dense
thickets, furnishes all the varieties of collecting one could
wish. On the rocky shores we found Nerita, Neritina, Leuco-
zonia, Acmaea, Columbella, Strombus, Livona, Tectarius, Echi-
nella, Siphonaria and many small forms not yet identified.
Chitons, of two species, were most abundant. On the Sea
Grape and other trees along the shore road we gathered
several varieties of Cerions, Drj'maeus, Cepolis and a verj' few
small Oxystylu- inidata. This species was one we had set our
hearts on, and look as we would, for two weeks not a good
live specimen came to our notice. Finally one day while out
to Waterloo after Drymaeus I called my partner's attention
to a ver>' large one up in the crotch of a gumbo-limbo tree,
supposing it to be inhabited by a soldier crab as all the others
we had found. What was my astonishment upon dislodging
him to find a beautiful live specimen. The next morning,
looking over the side of the piazza over Mr. Allen's store, I
had to call him up from below to see another almost as large
on the side w^all of the house. This proved to be alive also,
making two to my credit. Of course my friend could not hide
his disappointment and I could not let the opportunity for a
"good jolly" get by, so had to remind him several times that
it was too bad to live for two years in a place and get a little
wild fellow domesticated and then have a friend come over
and grab it. In some places on the island the soldier crabs
are so thick that on a dull morning one can scrape them up
by the bucketful. Among these we managed to find about
three dozen each of fine Oxystylas.
The Drj-maeus proved to be quite scarce until one slightly
foggy morjiing on a pair of sai)odilIa trees in a yard of one of
the natives we gathered over 150 fine specimens. We thought
the find and the liberty of climbing the trees worthy of a six-
pence from each of us. In the limestone walls of the quarries
and some of the street cuts we found a number of perfect
68 THE NAUTILUS.
fossils of the Cerian agass^izii. On the ^ass at Fort Charlotte
Succinea barhadensis was very plentiful. In the cut at Union
street were Bulimulus sepulchralis, Succinea ochraciym, Poly-
gyra cereolus, microdonta and a small Thysanophora. One
day we walked across the island to the south beach, and among
the palmettos were Chondropoma revinctum and Cerion agres-
tinc: On the shore which was here both sand and rocks we
found Lucina, Strombus, Asaphis, Siphonaria and Cerithiums.
Here was a landing place for a conch fisherman, and in front
of his cabin were heaps of thousands of the Strombus gi-gas,
all with a hole on one side of the spire, where they loosen the
part of the animal which is attached. These so-called conchs
are a staple article of food with the natives, at the market, in
the little cafes, and on all kinds of dry-goods box stands.
Along the street, night and day, one can buy the little conch
fritters for a penny and the smaller size for a halfpenny. As
a novelty these fritters are O.K. but for a regular diet they
are the limit, as I found out on my former visit when all the
hotels and restaurants were closed.
Jamaica was our objective point, aiid after pro\'iding our-
selves \\ath a chart of the island, folding cots and mosquito
bars, we waited several days for the steamer, running on a
three-week schedule. At the last minute we were refused
tickets on account of not havdng passports. A regulation war-
time passport is required going from one British province to
another in the same sea. We decided on a trip to Cuba, and
the next sailing of the steamer for JMiami saw us on the way.
After another trip to Brickel's hammock and steaming out
about 400 more Liguus we took the Fla. East Coast R. R. for
Key West. The trip doA\Ti is one certainly to be remembered,
especially if one tries to study the country and ask a few
questions to get a conception of the immense undert^iking of
constructing and maintaining a railway over these keys and
the sea. It is one of the greatest engineering feats of the
modem age.
At Key West the collecting is what may be termed slim.
We managed to obtain a number of Cerithium, Columbella,
Siphonaria, Succinea, Cerion, Polygj'ra and Littorina, The
THE NAUTILUS, 59
shore yieldetl about as much as could be expected with a mean
tide of less than two feet.
After what I had heard of the city of Key West, I was
greatly disiippoiuted to see the sailing boats and launches
which were going to ruin in the harbor, and the large number
of warehouses, stores and residences with the windows and
doors boarded up, cigar factories closed and the whole place
with a look of decline. While here we made the ac(iuaiutance
of the station master at Cud joe's Key, and from our descrip-
tion of 0.r\jstyla fioridensis he thought they were found there.
Consequently when he returned we went up with him for a
day, and our search resulted in two badly broken and bleached
specimens that looked as though the storms brought them.
Our day was not wasted, however, as we found a goodly num-
ber of the two varieties of Ceriati incanum which neither of
us had seen before, and a Succinea not yet identified.
At Havana everything had a look of prosperity — in fact we
found it the same wherever we went on the island. We made
Havana our headquarters and took trips every day to some
of the outside points for collecting. Santiago de los Vegas,
Guanajay, Guayabal, Guanabacoa and Playa all proved good
collecting ground, and at Matanzas, where we found abundant
species. We visited the museum connected with the high
school. This, as all other school museums I have seen, was
but poorly represented in the conchological line, while the
mammalia and fishes were much better. lu the vicinity of
some of the caves we had fine collecting, especially Liguus
and Cepolis hanplandi. Everywhere we went near the shore
we found Cerions in profusion. At Guayabal is a high ledge
or bluff with a thick growth of hard woods and vines j here
we found the best collecting. The Megalomastoma, Chondro-
poma and Helicina, three species, were abundant, as well as
many smaller forms.
60
THE NAUTILUS.
List op Species Collected, as Far as Determined.
a, Florida; b, Bahamas; c, Cuba.
omatula Mayn. . . .
phoenicea M. & C. .
Acmaea leucopleura Gmel.
var. ? b
punctulata Gmel. ... b
Aynpullaria miamiensis Pils. a
Area barhata Linn b
unibanata Lam. b
Astraea tuber Linn a
americanum Gmel. . . a
longispinum Lam. ... a
Auriculastrum peUiicens
Mke a
Bulimidus sepulchralis
Poey b
Bullaria amygdalis Dillw. a
occidentalis A. Ads., a
media Phil b
striata Bnig a
Cepolis bonplandi Lam. . . c
cwbensis Pfr e
milleri Pfr b
multistriata Desh. . . c
troscheli Pfr b
variant Mke a, b
Cerion agassizvi Dall (fos-
sil) b
agresthui Mayn b
eurystoma Mayn. . . b
gl<in.<! Kstr b
carlotta Mayn b
incanum Binn a
fasciata W. G. B a
saccharimeta P. & V.. a
niumia Bnig c
vnimiola Pfr c
mumia chrysalis Fcr. c
repitita MajTi b
santesoni I\Iayn b
sculpt a Mayn b
Cerithium ebunieum
Brug a, b
minimum Gmel. vars. a, b
m^uscarum Say a
variahile C. B. Ads. . a
Echinella nodulosa Gmel. a, b
Euglandinu minor Fer. ? . a
Eutrochatella sloanei Orb. c
Fasciolaria tulipa Linn. . . b
Fissurella barbad^nsis
Gmel b, c
nodosa Born c
viridula Lam b
Helicinu adspersa Pfr. . . c
clappi Pils a
orbiculata Say a
submarginata Gray. . c
lanthinn fragilui Lam. . . .a, b
Iphigenia brazUiatia Lam. a
Leucozania cingulifcra Linn, b
occll-ata Gmel b
Liguus crenaius castanco-
zonaius Pils a
cburncus Simp a
marmoratu^ Pils a
miamiensis Simp a
mosicri Simp a
fasciatiis'MnW. (Typi-
cal) c
castancus Simp a
THE NAUTILUS.
61
livingstoni Simp a
ornatus Simp a
rosea tu^ Pils a
testudineus Pils a
versicolor Simp a
ve.rillum Lam c
Lima scabra Dilhv a
Littorina lincata Phil. ... b
mespilum Phil b
ziczac Chem a, b
litterata Phil a
Livona pica Liiiu b
Lucina edcniula- Linn. ... a
Phacaides janwicensis ham. a
pennsylvanica Linn, .a, b
Codakia tigrina Linn a, b
Margaritiphora radiata
Lam a, b
Megalomastoma mani Poey c
Chama macrophyll-a Gmel. a
Chandroponia dentatum Say a
ottanis Pfr c
pfeigerianum Poey . . c
pictum Pfr c
poeyanum Orb c
revinctum Poey b
tenuiliratum Pfr c
Columbella mercatoria Linn, b
Comu^ nnts Hwa.ss b
Crepidula convexa Say ... a
plana Say a
Cypraea exanthema Linn., a
Cylindrella poeyana Orb., b
jejuna Pfr a
Cyrenaidea floridnna Dall. a
Detracia. bulloides Mont. . b
Drymaeus hahamensis Pfr. b
dominicus Rve a
inultilincatus Say ... a
OxystyUi undata Bnig. ... b
Feet en ornatus Lam a
Pedalion hicolor A. Ads. a, b
chenDiitzianuni Orb. . a
Melina obliqua Lam a
Physa acuta Drap b
Planasis lineatu^ Cocta. . .a, b
nucleus Wood a
Pleurodonte auricoma Fer. c
minor Fer c
provisoria Pfr b
Po^ygy^d auriculata Say. . a
microdanta Desh. ... b
septemvolva Say .... a
Praticolella griseola Pfr. . c
Proserpina depressa Orb. . c
Rumina decollata Linn.. . . c
Siphanaria alternata Say . a
brunnea Hani b
naufragitm Stimp. ... a
Sistrum, nodulosum, C. B.
Ads a, b
Melampus bidentatus Bsiy. a
coffeus Linn a
gundlachi Pils b
Microceramus gosse'i Pfr.. b
pontificus Gld a
Modulus modulus Linn. . . a
floridanus Conr a
Muricidea hexagona Lam. a, b
Murex poffium Gmel b
Nerita peloronta Linn. . . .a, b
tessellata Lam a, b
versicolor Gmel a, b
Neriiina pupa Linn b
virginca Linn a
viridis Lam a
62
THE NAUTILUS.
Oleaciim oleacea stramiiiea
Desh c
solidula Pf r a, b
Ope-as micra Orb a, b
Opisthosiphon hahamensis
Shutt b
Ostrea elongata Linn. ? var. a
Spirilla peronii Lam a, b
Spondylus echinatus Mart, a
Strombus hituherciUatus ....
Lam a
gigas Linn a, b
Subemarginula octoradiata
Gmel b
Succmea harhadensis Guild, b
campestris Say a
luteola Gld a
ochracinu Gundl b
Tcctarius tnuricatus Linn.
a, b, c
Thais deltoidea Lam a, b
floridatia Conr a
ha^mastoma Linn. ... a
patula Linn a, b
iindata Linn a, b
Thysanophora vortex Desh. a
Truncatella hUahiata Pfr. a, b
carihaensis Sowb a, b
Urocoptis hahame}isis provi-
dentia Pils b
elegans Pfr c
Vermicularia spirata Phil, a, b
Vivipara georgiana Lea . . a
EULOTA MAACKII, A SIBEBIAN SNAIL
BY T. D. A. COCKERELL
About the middle of the last century R. Maack explored
the Amur and Ussuri districts in eastern Siberia. This scien-
tific pioneer made the most of his opportunities, and the re-
sults were published in St. Petersburg in tliree large volumes,
1859 (Amur) and 1861 (Ussuri). Consequently, when my
wife and I recently visited Okeanskaja, on tlie coast near
Vladivostok, we noted that the most magnificent butterfly of
the region was FapUio maackii Menetries, and the largest and
handsomest snail was called Eulota muttcUii Gerstfeldt. Among
the plants, Rupreclit and IMaximowicz described from the Us-
suri district a leg\iminous genus Maackia, which unexpectedly
proved inseparable from CUidustrv<, based on a single species
growing in Kentucky, Tennessee and North Carolina. There
are also species of plants bearing the name of Maack ; thus
Delphinium maackiatium, L&nicera niau^kii and Pru n us inaackii
THK NAUTILUS. G3
are reeo«i:nize(l as valitl tixlay. Grerstfeldt describeil Maack'a
inollusca, Diplopoda and Chilopoda; finding, as might be ex-
pected, a gootl proportion of new species. Helix viaackii, then
introiluced as new, was said to be found rarely on the middle
of the Amur, and also between Sungari and Ussuri. The
original specimens had diam. maj. 30, rain. 26.5, alt. 24-30 mm.
Dr. Leopold von Schrenck (1859-1867), describing the mol-
lusca of the Amur country, was able to record a long series of
this Helix maackii, of which he recognized a form elatior,
long. 25, lat. 31 mm., and a form deprcss-ior, long- 21, lat. 30
mm. The elatior form was really the type of Gerstfeldt, but
the differences are those of individual variation, high and low
spired shells occurring in the same colonies. When I first
found Eulota maackii at Okeanskaja, early in July, I noted
that the animal was dark slate-color, nearly black; no dorsal
band ; sole dull white, suffusedly greyish anteriorly ; mantle
brownish-whit« or pale grey. {Eulota middendorffii Gerst.
differs in the color of the animal, which is pale, but also with
no dorsal band.) This was a matter of interest, showing that
there was no particular affinity with the common large Japa-
nese species of Eulota, which have a conspicuous and hand-
some dorsal baud. Anatomical examination confirmed the ap-
parent divergence from the Japanese series, and indicated that
E. maackii belong to true Eulota, and was not very remote
from the European E. fruticum. The stout and short penis-
sac passes into a very long and slender epiphallus (about 30
mm. long and 1.7 mm. thick in an Okeanskaja specimen), from
the end of which arises the retractor, without any sign of a
flagellum. The dart-sac is long and cylindrical, while the
mucous glands constitute an amorphous sacculated mass. The
vas deferens is about 35 mm. long. The sperm duct is long,
simple, s^'oUen at the ba.se, but beyond that filiform, with
hardly any bulb. The long and slender uterus has a cylin-
drical pale yellow albumen gland, about 26 mm. long. These
characters are all derived from fresh material.
The discovery of E. maackii at Okeanskaja, taken with the
previously published records, indicated a ^\'ide distribution.
It was nevertheless a little disappointing to find the snail quite
64
THE NAUTILUS.
unmodified about 400 miles up the coast, iu the valley of the
Amagu River, u. lat, 46°, The snails we collected were on the
Kudia River, a tributary of the Amagu, and although there
was much individual variation iu the height of the spire, I
could not detect any tendency to the formation of a distinct
race. There was also no basis for the separation of these coast
snails from those of Maack, which were found inland. Aft^r
this, it was distinctly surprising to find that at Kongaus, a
place in the hills no great distance east of Vladivostok, the
E. maackii were unifonnly of a distinct, gigantic race, wdth a
maximum diameter of 33.5 to 34.5 mm., rarely as small as 32.5.
The difference in measurement involves a great difference in
actual bulk, and the form deserves to rank as a new sub-
species, Eidota mauckii optima.^ A specimen of optima which
I dissected differed from the Okeanskaja maackii in having
the retractor arising from the vas deferens, about 5 mm. be-
yond the well-defined end of the epiphallus. Whether this is a
constant character is perhaps doubtful. The jaw of Okean-
skaja maackii has five strong thick ribs, in the manner of E.
duplocincta Mart. That of optima has the five ribs well sep-
Anatomy of Eulota muackii, from Okeanskaja, and jaw, sperm
duct, dart sac, and ephiphallus, etc., of optima from
Kongaus.
1 Ann. Mag. Nat. Hist,, June, 1924, p. 580.
THE NAUTILUS. 65
arated, after the style of E. platyomphalu MIklff. My assist-
ant, Mr. A. I. Lavrushin, was so fortunate as to liiul near
Kongaus a beautiful albino shell of optitna, clear yellowish-
white without nuu-k injurs, max. diam. 32 mm. This I will call
mut. alhidu. 1 w<us interested to learn that there is a nephew
of Maack's livin}^: in Vladivostok at the present time — an
engineer.
NOTES
The Purple Dye of Mollusks. — Any one desiring infor-
mation on this siibject can find a most interesting and com-
prehensive account in a paper entitled, " The Dyeing of
Purple in Ancient Israel," by Rev. Isaac Herzog, published
in the Proceedings of the Belfast Natural History and Phil-
osophical Society, session 1919-1920, No. 2, pp. 21-33. As a
summary the author says: — " The art of purple-dyeing in]
general, which dating from hoary antiquity — the mention of
tekelet and arganum in the cuneiform texts occurs already
about 1600 B. C. — passed through a long and checkered
career, finally becoming extinct, at least in the Old World, on
the fall of Constantinople 1453."— C. W. J.
Copies of Brj'ant Walker's recent paper on the Ancylidae
of South Africa can be had, gratis, on applying to him at
1306 Dime Bank Building, Detroit, Mich.
Colorado Land Shells. — Professor C. R. Crosby of the
Department of Entomology, N. Y. State College of Agricul-
ture, recently sent in PupUla 'blandi Morse, Vertigo modcsta>
Say and Succin^a avara Say from Pingree Peak, Colorado.
Also G(myodiscus shimeki cockerelli Pils. from Stormy Peaks
at 11,000 ft. The genitalia and teeth of the la.st are much as
in G. perspectiva, but the strongly arched, narrow jaw has
widely spaced, vertical lines, not close striae as in the eastern
species. There are 18.1.18 teeth. Some of the shells show a
few, broad, indistinct reddish flammules. — H. A. Pilsbry.
66 THE NAUTILUS.
PUBLICATIONS EECEIVED.
Freshwater Mollusks of Eagle Lake California. By
G. Dallas Hanna. (Proc. Calif. Acad. Sei., vol. 13, 4 ser. pp.
131-136, pi. I, 1924). Two new species Carinifex occidentalis
and Parapholyx mailliardi are described and fibred.
Rectifications of Nomenclature. By G. Dallas Hanna.
(Proc. Calif. Acad. Sci., vol. 13, 4 ser. pp. 151-186, 1924).
This paper deals vrith. the nomenclature of some 84 species,
59 new names are proposed to replace preoccupied names.
Two new generic names are also proposed. RinieteUa in place
of EctinochUus Cossm. 1889 not Ectinochila Chand. 1883. It
is an interesting case ; ordinarily the two names might possibly
stand, but Cossmann's name is a section or probably a sub-
genus of Rimella, therefore the genus and subgenus should
agree in gender as well as the species. Smithoceras is pro-
posed in place of Folycylus Mojsisovics 1889, not Lam. 1815.
— C. W. J.
Description of a New Genus and Species of Freshwater
Gastropod Mollusk from the Etchegoin Pliocene of C-vli-
fornia. By G. D. Hanna and E. G. Gaylord, (Proc. Calif,
Acad. Sci., vol. 13, 4 ser. pp. 147-149, 1924). This new form,
Scalez petrolia was obtained from the Midlands Oil Company's
No. 1, at a depth of 3306 ft. Based on operculums, which
the authors consider belongs to gasteropods in or near the
family Amnicolidae; the shell unknown. — C. W. J.
The Ancylid.e op South Africa. By Bryant Walker.
Published by tlie author. Full descriptions and illustrations
are given of the South African species, several of them new
forms, others doscrilx-d in a former number of this journal,
together with those of earlier authors. The genera Burnupm,
Perrissia and Oundlachia are represented. The paper is im-
portant to those concerned witli other faunas for the illustrated
synopsis of the new classification of all Ancylida? and other
general matter forming the Introduction. — II. A. P.
THK NAUTILUS. 67
The genus Gyiu)TOMA. By Calvin Goodrich. Univ. Michi-
gan, Mils. Zool., Misc. Pub. No. 12, 1924. The genus Gyrotama
{Schu:ost(>iua Lea) has received no systematic treatment in
fifty years. The older authors worked with very few speci-
mens, and to a considerable extent duplicated one another'a
species. Mr. Goodrich's work is based on the unrivaled col-
lections of H. H. Smith, with a study of the earlier types.
He recognizes some 13 species disposed in five groups. Several
of these groups he believes were derived from various so-
called Goniobases — impressa Lea, lacta Jay, showaltcri Lea
and others, which should be transferred to Gyrotama. Thei
species are illustrated on two excellent plates. G. hendersoivi
and G. walkeri are new species which Goodrich credits to H.
H. Smith. It is hoped that the author will extend his work
to other genera of this intricate family. We cannot have too
many papers of such workmanship. — H, A. P,
On the value of nuclear char.vcters in the classifica-
tion OP m.vrine g.vstropods. By W. H. Dall., Jour. Wash.
Acad. Sci., vol. 14, pp. 177-180, 1924, Various types of
nucleus, Sinusigera and others are specified, and the author
concludes that their differences are due to adaptation for a
floating larva or the reverse, and should not be regarded as
genetically fundamental.
MOLLUSCA op Colorado, Montana, Idaho and Wyoming.
By Junius Henderson. Univ. of Colo. Studies, XIII, No 2.
This work of 160 pages is a revision of the author's illustrated
catalogue of Colorado mollusca (1907-12). with additional
material from tlie other mountain states. The records for this
region have been compiled with great care, and prove to be
far more numerous than one would have supposed. Keys are
provided, and most of the species are figured. The discussion
of the OrcoJiclix group is of particular interest. Good prog-
ress has been made in dealing with records of species from
this region which are of long standing in the literature, but
not recently confinne<], and sometimes intrinsically improb-
able.—H. A. P.
68 the nautilus.
On a Thal.\5S0U) element in the Austr^mian Molluscan
FAUNA. By Charles Hedley. Victorian Naturalist, XI, No. 4.
The genus Coxiella, inhabiting saline pools and lakes some-
times over one hundred miles from the sea, is regarded as a
marine type of gastropod which has become adapted to these
inland waters; hence thulassoid in the sense of Bourguignat,
Smith and others. — H. A. P.
The Pliocene Mollusca of Grkvt Britain. By F. W.
Harmer, (Palaeontographical Soc., vol. 2, pt. 3, pp. 705-856,
pis. 57-64, 1923). Another part of this work has recently
been received, of which parts 1 and 2 were reviewed in The
Nautilus, Jun. 1922. The author died Apr. 11, 1923. This
part contains the Trochidae, Capulidae, Fissurellidae etc., and
the Tectibranehia, Scaphopoda etc. The biological sequence
of the genera seem badly disarranged. The illustrations are
excellent.— C. W. J.
ZuR Kenntnis der Plioc^nen Cragformation von Hall-
bjarnarstadur, Tjornes, Nordisland und ihrer IMollus-
kenfauna. von Hans Schlesch. (Abhand. Archiv fiir
MoU. Band 1, S. 309-370, Taf. 1-12, 1924). An account of
the Pliocene raollusks of northern Iceland, of which 157 species
are recorded, with tables showing their distribution both in
geological times and recent. — C. W. J.
The Life-History and Growth of the Pismo Clah
(TiVELA STULTORUM Mawe) . By Frank W. WeJ^nouth.
Stat€ of California Fish and Game Commission, Fish Bulletin
No. 7, p. 120, 15 figs., 1923.
In this small but meaty volume, the scope of which is indi-
cated rather than described by the title, there undoubtedly
comes to hand one of the most important contributions to our
knowledge of the West American Mollusca which has ap-
peared in many years. In fact, by weight of this paper alone,
Tivcla siuliorum becomes perhaps the most coraplet4?ly known
of any raollusk indigenoiLS to our Pacific coast, not even ex-
cepting the oft^n-dealt-with abalones. The importance of such
THE NAUTILUS. 69
a mass of careful data considered in the broad pliilosophical
spirit which manifests itself in this paper is diflicult to ex-
aggerate and renders tlie contribution a sif^nificant one from
the standpoint of tlie general biologist and physiologist quite
aa much as from the more circumscribed coign of the specialist
on Mollusca or the conservationist.
But a hint of its richness of content can be suggested here.
The interesting natural history of the Pisrao clam, save for
the embr>-onic and early life st-iiges — still, alas, incogniti — are
treated at length, even so prosaic a phenomenon as the mere
manner of distribution of the young and adult clams on the
beach being analyzed with such acumen as to afford clues of
real consequence to an understanding of the essential bio-
nomics. Perhaps the most important as well as the larger
part of the monograph is that which deals with the phenomena
of gro\nh (pp. 14-71). Although more than a decade ago
Belding showed convincingly that certain strong growth-lines
visible on the shell of the mature scallop were seasonal in for-
mation and therefore essentially of the nature of the annual
rings in trees, most other students of molluscan life-histories
have been unable to offer much in the way of confirmatory
evidence, or, even where they admitted its validity, to extend
greatly the practical application of the principle. Wey-
mouth's conclusive demonstration of the truly annual nature
of the major growth-lines in the Pismo clam and his ingenious
working-out of practicable methods of distinguishing them in
instances where other influences have operated to obscure
them, should give a great impetus to similar work among other
lamellibranchs and, be it hoped, among gastropods as well.
These records of a strictly seasonal arrest in growth are shown
usually to be readily distinguishable from the numerous less
significant growth-lines by their heavier pigmentation, by their
coincidence with certain concentric thickenings or depressions
in the shell, and, very curiously, by their translucence when
the shell is held against a strong light — this last alone being
well indicative of the fundamental difference in formation of
the two types of concentric lines. To the minor intra-seasonal
striae it seems that the old and convenient term "incremental
70 THE NAUTILUS.
lines" might now well be restricted. Weymouth found that
these discoveries enabled him to calculate the age of the thou-
sands of individuals examined with a remarkably small inci-
dence of error or doubt, the oldest specimen recorded having
attained the hoary age of 26 years, while in several instances
tlie nearly constantly prevailing peculiarities of the rings for
particular years were found, as in the well-known case of
rings of trees, to correspond to certain outstanding weather
conditions of the years in question — an exceptionally mild
winter, a season of extraordinarily high surf, and so on. The
heaviest clam noted measured 17.75 cm. in length and weighed
1.415 kg. (3 lbs., 21/2 oz.).
Recognition of the facts outlined also made possible some
interesting discoveries concerning variation in the size of
broods or the surviving fractions of same for different years.
As favorable conditions during the spaN\ming and larval his-
tory would appear to be more infrequent tlian distinctly un-
favorable circumstances, the yearly fluctuaton in numbers of
young, and therefore of the commercial clam crop in succeed-
ing years, is frequently very large. This greatly complicates
the question of increase or depletion, and leads the author to
recommend only very moderate legcd measures of conser\'a-
tion, chiefly directed toward the protection of the young, to
which floating oil and the depredations of local clammers
appear to be the worst menace, until such time as more pro-
longed investigation gives a conclusive lead toward some more
positive effort.
A discovery which will be new to most students is that the
very yoiuig Tivelas possess a long slender byssus with an
anchor of attaclied sand griuns at the distal end, wliich serves
to aid them in holding their own in the loose and constantly
moving sand of their habitat luitil their own weight becomes
suflicient, at a length of 2 cm. or over, to enable them to
maintain themselves without it. Incidental obsers-^ations upon
Paphia staminca and Macoma nusuta show that juveniles of
these species likewise possess a byssus. The }"oung of the
Pismo clam are found to be confined to the intertidal zone,
with no less than 90 i)er cent "concentrated in a strip 13
THE NADTILU8. 71
metres Nvide" exposed at nearly every tide. The normal
rajige of the adults is botli greater in total vertical extent
and pitoheii at a considerably lower level than this (low tide
to a depth of several fathoms).
The Pismo clam is shown to be hermaphroditic, both eggs
and sperm being pro<.luced coincidently, though not as a rule
at identical periods of maxima, from the same gonad.
The author does not consider commercial farming of open
beach species of this tj'pe possible.
Unlike so many modern papers in the field of ecology and
bionomics, this one is written in pure English. The lack of
either technical verbiage or of words coined to cover phe-
nomena simply demonstrable without them is a blessed relief
in a work of this nature and makes reading it a pleasure. It
is a pity that so little of the terribly needless mass of extant
literature on the taxonomy of our west American moUusks
emulates the care and thoroughness of this one in the bionomic
field. "We systematists should take the lesson to heart. — S. S.
Beery, Redlands, California.
The Kentucky River and its Mussel Resources. By
Ernest Danglade. Bureau of Fisheries Document no. 934.
1922. This brief article gives some account of the river, with
a list of 40 species of mussels. The following summary may
be of interest :
"The Kentucky River is approximately 400 miles long and
contains many valuable mussel beds. In the upper reaches of
the stream these number about two per mile of channel. They
have well-defined and characteristic locations easily marked.
"The upper Kentucky River is practically an unknown
and unworked mussel-bearing stream and contains an abun-
dance of mussels of commercial value, possessing good nacre
and texture. Of these the mucket constitutes about 90 per
cent. This shell has a desirable color, texture, and uniformity
of thickness throughout. It appears probable, therefore, that
this stream may be particularly useful in the near future as
a source of remunerative employment for the mussel fisher-
man and of desirable raw material for the button manufac-
72 THE NAUTILUS.
turer. The pearls of this river, as a by-product of mussel
fishing, are of small consequence, both in the quantity and
quality of the pieces found.
' ' The railroad and steamboat shipping facilities of the main
river, of North Fork, and of lower Middle Fork are satisfac-
tory. On the upper Middle Fork and on South Fork there
are no railroad or steamboat connections and shipments must
be handled by small boats. In particularly dry seasons of the
year transportation must be made by hauling over rough
roads. This is especially true of the South Fork.
"The method of shell fishing in the Kentucky River is
limited principally to hand picking or to the use of the shell-
fork. A stiff bottom in which the mussels bury themselves
deeply makes implements commonly used elsewhere in shell-
ing, useless in this river.
"Of 40 species of mussels observed as indigenous to the
river, 22 are commercially usable, but only 9 are of relative
importance. This number includes as the most common shells
suitable for button manufacture : the mucket, the pocketbook,
the pimplebacks, the pistol-grip, the long niggerheads, the
maple leaf and the fat mucket. ' '
ARiONiDiE OP Norway. By Fridthjof Okland. (Videnskaps.
Skrifter, 1922, I, no. 5.) In this excellent account attention
is particularly drawn to the diagnostic value of the location
of the shield bands, to the variation of the internal organs,
and to the distribution. It is fully illustrated with 46 text
figures, 5 maps and two colored plates. Arutn subfu^cus is
found to the northern extremity of Norway; the big A. ater
to the seventieth parallel. A. hortensi.<t is found only in the
south, and possibly was introduced by man. Two of the
species treated of, A. circurtiscripUcs and A. hxyrtensis, have
been introduced in the United States. — H. A. P.
The Nautilus.
Vol. XXXVIII JANUARY, 1925. No. 3
THE FAMILY LANCID5: DISTINGUISHED FROM THE ANCYLID.E
BY HENRY A. PILSBRY
The large West American fresh-water Umpets now forming
the genus Lanx Clessin have always been associated with the
Ancylidie. Hannibal, in his classification of 1912 ' placed Lanx
in the subfamily Ancylinte and its subgenus Walkerola in the
Laivapecina\ Subsequently he proposed a subfamily Lancinje,*
without stating what genera it contained. His classification of
Ancyli, however, does not appear worth criticism. It was not
based upon any new observations or data.
In 1918 Walker' made a subfamily Lancinse for the genera
Lanx and Flsherola.
Recently in the course of looking into the characters and
position of the adaptive gill or pseudobranch in some New
York Ancylidffi I took the occasion to examine several species
of LanXy and was surprised to find them without any trace of a
gill (pseudobranch). This was confirmed, together with other
characters mentioned below, in L. patelloides, L. subrotundata
and L. (Fisheroln) lancides.
' Proc. Malac. Soc, London, X, p. 147.
'Nautilus, XVIII, 1914, p. 24.
'Univ. of Michigan Mus. Zool. Misc. Pub., No. 6, p. 17. Walker cred-
ited the suhfamily Lancinsu to Hannibal, but as the latter author neither
defined it or stated its contents, he can hardly be author of the group defined
by Walker.
74 THE NAUTILUS.
As Lanx has no lung, merely an open furrow between mantle
and foot, its respiration is probably aquatic and effected by the
whole of the integuments exposed. It appears to be a river
snail, probably in running, well oxygenated water. In this
connection, information on its station is urgently needed.
It is strange that no conchologist has noticed hitherto that
in Lanx the apex of the shell, though generally very near the
center, is slightly before it, never behind, as in all patelliform
Ancylida?. This anterior position is strongly marked in such
species as L. nutialU and L. lancides. This last form was said
by Hannibal to have the apex " subterminally posterior"!
The characters of shell, pallial region jaw and teeth demon-
strate that Lanx has no direct or near relation to the Ancylidte.
It belongs to a separate family, related to L^-mntieidfe somewhat
as the Ancylidse are to the Planorbida^.
The more conspicuous characters are here contrasted :
Ancylid^ Lancid.5
Shell limpet-like,' the apex Shell limpet-like, the apex
behind the middle and more in front of the middle and on
or less inclined to the right or the median longitudinal line,
left.
Agill (pseudobranch) hang- No pseudobranch or other
ing in the mantle cavitji- (on appendage in the mantle cav-
the right side in Ancylus, on ity. Animal dextral.
the left in all other known
genera).
Jaw in form of a high arch Jaw composed of a large
of many small, subequal ele- upper piece and two lateral
ments. appendages as in Lymna^a.
Radula various, but not Radula Lymnand.
Lymna>id.
' At li'ast in the comparable kpik'I""-
The number of species of Lnm: is uncertain, as the distribu-
tion and variation are still imperfectly worked Dut. It may be
that L. ])a(ellui(lex intergradcs with L. neivl>crnji, l>nt s(im(^ large
lots I have seen do not show intcrgiadation. The distinction
THK NAUTILUS. 76
of Fishcrola as a genus appears to me quite uncalled for, as L.
nutUilli and kootaniensis have intermediate positions of the apex
between F. laucide.-i and the species with nearly central apices.
In fact, I am much disposed to unite all the described forms of
the Columbia river system under the oldest name, Lanx nuttalli
(Hald.); F. Inncidcs, L. kootanieasls and another unnamed form
from the Spokane river being subordinated as subspecies.
While I have seen many specimens, they are from few, well
separated localities.
What grounds, if any, exist for the distinction of the Plio-
cene Zalophancylus Hannibal have not been made clear. A list
of the species of Lancid.T? follows.
Lanx patelloides (Lea). Sacramento drainage, California.
Syn., A. patelloides Binney.
Lanx newberryi (Lea). Sacramento drainage.
Lanx subrotundata (Tryon). Umpqua drainage, Oregon.'
Syn., L. subrotundus Walker, 1918.
Lanx alta (Tryon). Klamath River, Appears to me dis-
tinct from patelloides and neivberryi, but nearer subrotundata.
For the present it had better stand as a species.
Lanx klamathensis Hannibal. Upper Klamath Lake, Ore-
gon, Well distinguished by the depressed form and thin tex-
ture. It is the type of s. -g, IValkerola Hannibal.
Lanx nuttalli (Hald.). Columbia drainage. As will be
seen by comparing the measurements, locality and collector,
Haldeman's A. crassus was described from the same specimen.
Probably he forgot his former name. The northern A. kootani-
ensis Baird appears to be a subspecies W'ith less anterior apex;
I have not seen topotypic material. Fisherola lancides is an-
other subspecies of the Snake River, in which the apex is a
little more anterior, but some of the original lot before me run
close to nuttalli.
' Said by Hannibal to belong to the "Colunibiii system,'" but the reason for
this grouping is not apparent.
76 THE NAUTILUS.
A VISIT TO THE HAWAIIAN ISLANDS
BY T. D. A. COCKERELL
" For we have seen the golden sun
Sink in the sea,
The moonlight on the palms, and heard
God's melody."
It is not uncommon for modern writers, in discussing the
islands of the Pacific, to begin by stating that they have little
sympathy with the sentimental nonsense of previous authors,
but propose to state the facts, and so forth. It is no doubt true
that the expression of feeling concerning the islands has often
been trifling or insincere, and at times wanting in judgment or
good taste; but it should be recognized that language is bareh'
adequate to do justice to the beauty, mystery, and romance of
the subject. To the naturalist, at any rate, the splendid blue
of the sea, the graceful curves of the cocoanut palms, the varied
foliage of the mountain forests, are but the setting for a drama
which at once stimulates and baffles his intellect. Whence
came this marvellous fauna and flora? What influences have
governed its evolution? Why this extraordinary multiplica-
tion of species, often with almost unbelievably restricted ranges?
In spite of many learned tomes, we are still groping for light on
these problems, but hopefully, convinced that their solution
wnll furnish a ke}" to some of the most perplexing questions
confronting the biologist. With the facilities for research now
established and lieing developed in Honolulu, we may expect
to see the riddles of the Hawaiian biota attacked anew, with
modern intensive methods, likely to give results of fundamental
iin]iortance.
The naturalist of today is indeed fortunate, wlien he a])-
proaches the Hawaiian problem, in finding his path made com-
paratively easy by magnificent monographic works. The Fauna
Hawaiiensis, the volumes of the Manual of Conchology, Jor-
dan's lists of the fishes, give at first the impression tliat nearly
all has been done, not only tiio fuuiia, but also tlio (lora has
THE NAUTILUS. 77
received elaborate treatment. Yet in spite of all this, it is still
easy to make discoveries. liast summer, while I was in Hono-
lulu, Dr. D. S. Ji)rdan obtained a number of fishes new to the
islands (some new to science) by visiting the market each morn-
ing. Perkins, in his classic Introduction to the Fauna Hawaii-
ensis, stated that in his opinion only about half the endemic or
truly native species of insects had been discovered and de-
scribed. Dr. C. Montague Cooke tells me that he could now
about double the species of LcpUichatuKi described in the mono-
graph, and he has numerous undescribed snails of other genera.
Thus the present investigator is in the doubly happy position
of finding plenty of important work to do, while the founda-
tions have been securely laid by masters of the science.
In still another respect the worker of today has advantages.
He finds paved roads around the islands, and facilities for get-
ting about are vastly superior to those of former times. Dr.
Perkins writes me (October, 1924): " It is hard to realize now
the conditions under which I worked, from 1892 to 1897 par-
ticularly. Roads were bad, often impassible in wet weather,
and I did most of my work on foot. . . I cut many paths
myself single-handed through the densest wet forests, sometimes
8 or 9 miles in length, and lumped tent, guns, and all ap-
paratus and food on my back in such places."
In the following account of my own brief explorations, I am
enabled to give the names of the snails through the kindness of
Dr. C. Montague Cooke, who identified nearly all my speci-
mens. I am responsible for the slugs. The hours I spent at
the Bishop Museum, looking at the wonderful collection of
Hawaiian shells and hearing Dr. Cooke discuss the problems
involved, are never to be forgotten. May all his learning and
enthusiasm find full expression in print, where it will reach
those in other lands, and of later days ! He speaks at times of
organizing collections and data so that posterity may find it
easy to proceed. This he has done and is doing daily, but who
can expect that a successor of equal ability will be found ? The
history of science shows that unfinislied work is too often
neglected, or falls into incompetent hands, as indeed is true of
human activities in general. Let any one familiar with modern
78 THE NAUTILUS.
zoology name a dozen or twenty of the leading men now grow-
ing old, and ask himself who will step into their shoes. In too
many cases one has sadly to say that no candidate is yet in
sight, and none is likely to appear.
I had scarcely more than arrived in Honolulu when (July
16) Mr. 0. H. Swezey, of the Sugar Planters Experiment Sta-
tion, kindly offered to show me Mt. Tantalus and the upper
part of the Manoa Valley. Mrs. Faus, formerly a student in
our department at the University of Colorado, took us up the
mountain in her car, and we circled round through the forest
on an obscure trail known to Mr, Swezey, who pointed out the
characteristic plants, insects and molluscs. Having been famil-
iar with the literature of Hawaiian natural history since 1891,
when I helped Dr. A. R. Wallace collect data for the second
edition of "Island Life," it was with feelings readily to be im-
agined that I saw the Achatinella snails on the trees, caught the
bees of the insular genus NesopTosopis, observed the splendid
butterfly Vanessa tammeamea circling through the glades, found
the singular longicorn beetle Plagiihmystis pulverulentiis in its
native home. The entomological results of the trip might form
the subject of another article, but we are now concerned with
the snails. The species collected on Mt. Tantalus were as fol-
lows:
Achatinella vulpina olivacea Reeve. This is the most widely
spread form of vulpina, and ought to have been the type of the
species. As is shown in Man. Conch., it occurs in ten or twelve
valleys, the typical vulpina apparently only in three. My
olivacea shells vary from green and pale j'ellowish to reddish-
brown; usually bandless, but rarely with a ]>eripheral black
band.
A. stewartli producta. Reeve. These are sinistral; remarkable
for their large size. The type ol producta is dextral, and one
wonders whether the sinistral rare on Tantalus (referred to in
the Manual) is not an indcjiendent dcveloinuent. It is more
robust than the dextral producta iron\ the head i)f Manoa
Valley, and looks like a different thing.
A. lorata Fer. A beautiful peg-top-f^iiaped siiell; some of the
phase called pallida *' Nuttall," Reeve; others pure white, the
mut. alba, also named, but not described, by Nuttall (1839).
THK NAUTILUS. 79
Amiistni lextilis h'vv. One specimen.
Ldviinflla (/niviila F6r. I got only dead shells.
Aurirulella diaphana Smith. Sinistral.
Torudtcllidf-'^ procerulus Ancey. This minute species originally
Jesorihcd from Maui, is widely distributed over the islands. Is
it a very ancient, conservative form, or has it some unusual
means of travel ? On Kauai I found another Tornntellides
{vKicromphala Ancey) which has a similar wide distribution.
Endodonta (or Nesophila) apparently a new species, which
will be described by Dr. Cooke. It is a minute form, with
rounded periphery and strong ribbing. I found several in a
dead and dry tree-fern stem. It was astonishing to find an ap-
parent novelty in such a well-worked localit}', within sight of
Honolulu.
Si(ccinea rotayidata and a small Helicina complete the set.
From near the top of Mt. Tantalus we crossed a marshy de-
pression and came down a zigzag trail at the head of Manoa
Valley. Although the distance was short, we found different
snails, as follows:
Achatiaella hcllidn Smith, of the pale, straw-3'ellow, slightly
olivaceous, form, wholl}^ without bands, except a very faint
brownish one below the suture. This may be called mut.
straminea, basing the name on the shell figured in Man. Conch.,
plate 41, f. lOd, but my shell is exactly the same.
A. stewart'd Green. Beautiful green shells, usually with two
black bands, but one of my shells is bandless. All are sinis-
tral, whereas the original type was dextral.
A. stewartii producta Reeve. A long dextral shell, with nar-
rower aperture than in my stewart'd or the Mt. Tantalus ^^ pro-
dvda.''
Amastra turritella Fer. Varying from warm reddish-brown
to practically black.
Auriciddla auricida Fer. Both sinistral and dextral shells of
this neat little species, and the body whorl varying from white
through fulvous to deep reddish-brown. One sinistral shell is
entirely very pale yellowish.
Also Phlhnesin and Opeas,
80 THE NAUTILUS.
At a later date I went with the Trail and Mountain Club to a
waterfall at the head of Manoa Valley, in a deep gulch. Here
I found the introduced Opeas javanicuni Reeve, and what was
much more interesting, a series of the slug PhUomycus (nistralis
Bergh. This is a rather small pale grey slug, with three longi-
tudinal bands, and irregular mottling both between and below
the bands. I have investigated the synonymy, with special
reference to the findings of Collinge (Journ. of Malacology,
Dec, 1899), and reach the following conclusions:
(1) It is quite probable that the slug was introduced from
China, and it may be, as Collinge claimed, identical with P.
bilineatus (Benson). It has a jaw with numerous sharp ribs,
which associates it with bilineatus and separates it from the
Japanese P. confusus (Ckll.), which also differs in other respects.
It must be remembered, however, that Benson's slug came
from Chusan I., off the coast of China, and may be a local en-
demic species. A Chusan specimen in the British Museum is
reddish-brown, and superficially appears rather unlike the
Hawaiian animal. Thus it is premature to assume identity,
and the Hawaiian species must continue to stand as P. australis.
(2) It is certain that P. australis is distinct from the Japa-
nese P. confusus (Ckll.) and the Formosan P. formosensis
(Ckll.). P. chinensis (Ckll.), found 1300 miles up the Yang-
tse, is more uncertain, but I now believe it is not audralis.
It remains, of course, for some one to ascertain what forms
of PhUomycus occur in various localities in eastern Asia. One
may surmise that when these are properly collected and studied,
many species will be found.
Much lower in the ^lanoa Valley, within the limits of Hono-
lulu, the introduced slug VeronirrUa Icydigi (fc?imroth) is
common.
After returning from the expedition to Kauai, described be-
low, I was much occupied with the meetings of the Pan-Pacific
Food Conservation Conference, and had little time to hunt for
snails on Oahu. At Kawaihapai, on the north side of the
island, I found it extremely dry, and on the mountain side
overlooking the sea could find only Svcrinea caduca Mighels and
the introduced-, £t</o<a sirnilaris. 1 had been brought uj) in
•nil-: .\ A I 111. IS. '^1
England to hiok upon Succiiicit as almost scniiaciuatic, and it
was surprising to find the genus in the dryest possible places.
In Oahu, and particularly at Waimea on Kauai, I found hill-
slopes with Oinintid, looking exactly like places in Madeira and
Porto Santo which were most prolific in endemic Helicidae.
But in the Hawaiian Islands one has to go to the moist forest
for the native snails, xeroi)hytic regions proving extremely bar-
ren, with rarely anything better than some common introduced
form. There are some endemic insects which inhabit xero-
phytic stations, but probably such areas were extremely re-
stricted prior to the changes brought about by man. At Sacred
Falls, overlooking the northeast coast, I obtained Amastra in-
flata Pfr. and Leptachatina fragilis Gulick. The latter appears
in the Manual as a synonym of L. ginnmea Gulick, but I infer
that Dr. Cooke now considers it distinct.
In the fossil bed along the face of Diamond Head, where col-
lecting is uncomfortable on account of the burs of the grass
Ctiicfirus, I found Leptachatina oryza Pfeiffer, and L. subcylin-
dracea Cooke, the latter evidently distinct, but first published
as a variety of L. oryza.
When I svent to the islands, one of my chief ambitions was
to visit Kauai, the northernmost and in some ways most inter-
esting island. On making enquiries, I was discouraged, being
told that the journey would be both expensive and difficult.
Very fortunately, I learned at this point that the Trail and
Mountain Club was arranging their first expedition to the
island, and had no difficulty in making arrangements to go
along. As the Trail and Mountain Clubs have a system of
interchangeable membership, the fact that I was a member of
the Colorado Club at once gave me good standing in that of
Honolulu. The trip, with a company of sixteen thoroughly
congenial people, men and women, was made under the leader-
ship of Mr. R. J. Baker, a well-known photographer of Hono-
lulu. For fifty dollars each we had five glorious days on the
island, with everything found, including a large truck to take
us about, and also the steamer journeys back and forth. There
was even enough left over for a celebration at the Outrigger
Club at Waikiki, and a ride in an outrigger canoe. Certainly
82 THE NAUTILUS.
the visitor who cares for out-of-door things cannot do better
than get acquainted with the Trail and Mountain Club, which
has a permanent office in the Young Hotel.
At a point on the eastern coast of Kauai, we left the road and
took the trail leading inland, eventually reaching the Upper
Anahola Valley. At different points along the trail I found
snailSj including Lepiachatina cylindrata Pease, L. striatula
Gould, Microcystis chammoi Pfr., Eulota sbnilaris Fer., Opeas
clavulinum var. haivaiiense S3'kes, and Helicina knudseni Pilsbry
and Cooke. Mia'ocystis chamissoi, except for its larger size, is
extremely similar to M. venosa Pease, which my wife collected
last summer on the Island of Rarotonga.
There is a waterfall at the head of the Anahola Valley, which
the part}'^ hurried on to see, but I stopped at a sharp turn in the
trail, where the vegetation was dense and conditions looked
promising for snails. I was richly rewarded by the discovery
of several specimens of Nesophila capiUata (Pease), which had
not been found since Pease (Am. Journ. Conch., II, 1866, p.
292) recorded it as Helix capillata from the "Sandwich Islands."
Dr. Cooke has one of Pease's original specimens, so there is no
doubt about the identity. The species is omitted from the lists
An Manual of Conchology.
On the north side of Kauai are the famous Hacna caves, and
in this vicinity I met with Succinea lumbnli^ Gld., Microcystis
chamissoi and Eulota similaris. The Eulota were of the bandless
form (f. nnicolor F6r. ), while that from the Anahola Valley
shows a brown band (f. zonulata Fer.). Such variation doubt-
less occurs throughout the range of the species.
Near the Haena caves, in the sand hills, I was able, follow-
ing Dr. Cooke's directions, to find the deposit of fossil shells,
containing great quantities of the variable Cariiia dolei Ancey,
which appears to be now extinct, all the known specimens
being presumably from this deposit. The genus Onrlia, witl)
many species, is confined to Kauai and the neigliboring island
Niihau. Along with the Cardia I obtained the following:
Avtnslra nvchola Cld. Possibly also extinct, as the Manual
says that although the species is common in collections, all the
.specimens have the apiiearance of having been collected dead.
TlIK NAUTILUS. 83
Leptachntiiia brevicula Fease? The name queried l>y Dr.
Cooke, ajul in fact it ai)[)ear.s to he a difforeiit species, the aper-
ture heiiig conspicuously cliiTercnt from that of a cotypc figured
in the Manual.
L. hrei'icula micra Cooke. This is known in a living state
from altitudes of 1500 to 1700 feet.
L. leucochila Gulick.
L. antlqua Pease? Name queried by Dr. Cooke. It is like
the Koloa beach shells tigured in the Manual, but very differ-
ent from the more slender shell figured by Pease. Surely it
should receive a new name.
Tornatellides macrumphala Ancey. One juvenile.
Endodonta laminata Pease. A typical Endodonla^ with keel.
NesophUa sp., apparently undescribed, but Dr. Cooke had 3
already, his MS. species "No. 8."
Hclicina berniceia Pilsbry and Cooke. Only known fossil.
It would appear that these sandhill deposits are of Pleistocene
age, reminding one very much of the deposits in the sandhills
at Cani§al, Madeira, and with apparently about the same pro-
portion of extinct species. I greatly regretted that I could not
spend more time collecting in this locality.
THE TYPE OF ANCYLASTRUM IS ANCYLUS FLUVIATILIS, MULLEE
BY A. S. KENNARD, A. L. S. AND B. B. WOODWARD, F. L. S.
But for the fact that more pressing work engaged us, Mr.
Bryant Walker's reading of the facts governing the selection of
the type of Ancylastrum^ would have been challenged before.
The Rule (Art. 80, 11 e) is quite clear: "Species are ex-
cluded from consideration in determining the type of genera
. . . which were not included under the generic name at the
time of its original publication."
When Anryltistrina was created^ the species cumingianics had
' Nautilus, XXXV, 1921, p. 5.
*Journ. de Conchy!. , iv, 1853, p. 63. The name was employe*!, as the
author admits, in afjrcemenl witli Mo(iuin-Tandon, who, however, claims it
as his own (Hist. Moll. France, ii, p. 483).
84 THE NAUTILUS.
not even been conceived; it first appears as a nomen midum in
what Mr. Walker admits was a later paper {torn. ciL, p. 170),
but it was not born until July, 1854 (Proc. Zool. Soc. Lond.,
1853 [1854], p. 91). It is quite obvious, therefore, that by no
amount of sophistry can cumingianus be made the type of An-
cylastrum.
The case is reall}"^ quite simple. Although Beck in 1837 had
divided the genus and created Aa-oluxus for the Incustris group
he gave no reasons for so doing. It was Gray in 1840' who
made the division on anatomical grounds. He did not, as
Bourguignat erroneously asserts, make at that time two genera,
but as his numeration of his genera and species shows, sepa-
rated off his Velletln as a subgenus, without, however, giving it
that distinction. His arrangement was:
3. Ancylus
1 . Ancylus fluviatilis
Velletia
2. Velletia lacustris.
His two divisions were each for a single species and therefore
even then monotypical, but all doubt as to the intended types
is set at rest by his article in the Proc. Zool. Soc. Lond., 1847
(p. 181), where the same arrangement obtains and the two
species are definitely named by him as types.' This last paper
has entirely escaped Mr. Walker's observation. It was not
until 1857 in his second edition of Turton's Manual (pp. 210
and 220) that Gray made these divisions into two distinct
genera.
Expressed in modern manner Gray's classification in 1840
and 1847 was:
Genus Ancylus
8ubg. Avryliis {s. s. )
Subg. Velletln.
Bourguignat in 1853 (lor. rlL, p. 03) accepted Gray's sub-
divisions, but, as the fjishion then was, instead of retaining the
' Turlon'H Manual, mw cd.. p. 217.
' Here also he admits that Acrohrua and Velletia are synonyms.
THE NAUTILUS. 85
name of the genus for one section as "s. s." he gave a distinc-
tive name for each section, thus:
Genus Aurylm
S. g. Anci/lfistrum.
S. g. Velletiu.
Bourguignat'R AiicijUistnim was therefore merely a nom. nov.
for Aiin/lus as Hmited by Gray.
Now by the Rules (Art. 30, II, f . ) "In case a generic name
without originally designated type [in this case Ancyln.struin']
is proposed as a substitute for another generic name, with or
without type [in this case Ancylus, type A. fluvintilis] the type of
either, when established, becomes ipso facto type of the other."
Hence the type of Ancyhistrum is by the Rules beyond question
A. flumatilu (Miiller), and Mr. Walker's Pseudancylus^ goes to
swell the ever-lengthening list of unwanted synonyms.
ON THE HELIX PERSPECTIVA OF MEGERLE VON MUHLFELD VERSUS
THAT OF SAY
BY A. S. KENNARD A. L. S. AND B. B. WOODWARD, F. L. S.
Megerle's paper "Beschreibung einiger neuen Conchylien "
was published in the Mag. Gesell. Naturf. Freunde Berlin, Vol.
viii (pp. 3-11), the title page of which is dated 181S. It
seems, however, that the volume in question was published in
four parts: Pt. i, 1816; pt. ii, 1817; pt. iii, 1817, and pt. iv,
1818,' and tliat Megerle's paper appeared in the first part.
Say's name came out the following year in the Journ. Acad.
Nat. Sci. Philad., i, p. 18. Consequently Megerle's name
holds, and the Helix solaria, of Meuke becomes its synonym,
' Nautilus, XXXV, p. 58.
^See Isis, 1818. col. 144S, 1707 and 1809 [The refemu-e to Megerle's
paper on pape or column 14S3 — I'xl.] also M(?in. .\cad. .Sci. .St. IVtersbourg,
vi-ix. — ('. Davics Sherborn (Index Aniinalium). This appears to have been
known, in part at all events, to i'inuey and Bland, for they give the correct
date for Helix eereolus Megerle, whicli api)ears in the same paper.
86 THE NAUTILUS.
whilst Say's species will have to take the later name Helix
patida of Deshayes, and following the nomenclature of the
Pilsbry- Johnson "Catalogue" (Nautilus, XI, p. 141) be
known as Pyramidula {Gonyodisciis) i>atula (Desh.), or yet bet-
ter as Goniodiscus patula (Desh.).
AGNATHOMORPHOUS AULACOPODA
BY H. BURRINGTON BAKER
In a recent paper (1924, Occ. Papers Mus. Zoo. Univ. Mich.,
no. 156), a provisional arrangement of the " Streptaxidae "
was presented. This was mainly based on radulae, although
certain other features of the anatomy of Scolodonta (Systrophiella)
eudiscus and of Redartemon jessei (one incomplete specimen)
were included.
Since then, I have dissected Strqytaxis (Odontartemon) glaber
normalis Jousseaume and reexamined the other two species.
These new data, which will be figured in a future paper, force
a quite different conception of the relationships than that indi-
cated in the article cited above.
The animal of Streptaxis glaber normalis is very similar to that
of Bedartemon jessei. Foot: strictly holopod; i. e. , without
pedal grooves. Pallial complex: described and figured in this
or a related species by Dr. Pilsbry (1907, Man. Con. XIX, fig.
lii-5); lung venation much weaker than in R. jessei. Penis:
shorter and stouter, but fundamentally similar in structure;
lower half surrounded by a heavy, muscular sheath, which at-
taches along the vas deferens; penial retractor inserted at apex
and attached to diaphragm on the right of spermoviduct
("uterus"). Vas deferens: arises from base of spermoviduct,
runs straight across to penial sheath, encircles the penis, and
finally passes up along tlio side of that organ to enter at its
apex; arrangement i-iniilar in R. jessei {op. cit., fig. x-55), and
portion above penial sheath shown in my figure (although I
did not recognize it at the time). Free ocular retractors: short;
THK NAUTILUS. 87
right one fused postcriad with the left to form a common hand,
which is about half as long as either free muscle and joins the
tail retractors near the level of the mantle edge. In R. jessei,
both eye retractors join a broad band of muscles that retract the
snout; in both species, the right ocular is completely free from
the genitalia. Jaw: absent, only represented as a low, rounded
fold above the buccal aperture.
Further study of the animal of Scolodonta eudiscus brings out
the following additional features. Columellar muscle: receives
the long penis retractor near the base of the liver, the pharyn-
geal muscle at about the middle of the body cavity, then the
two right oculars and finally the two left ones, which unite
with the tail retractors near the foot; right ocular bands pasB
between penis and vagina. Jaw: represented by a definite,
sharp-edged, crescentic fold, which looks like a true jaw but is
quite plastic; examination under high magnification shows that
this structure is covered with a thickened epidermis composed
of polygonic cells. Thus, although a coherent jaw is absent, a
vestigial, homologous development is certainly recognizable.
Foot: margined on either side by a distinct pedal groove, both
of which terminate at a definite, caudal mucous pore below a
small, dorsal projection; above each pedal groove, a rounded
ridge is marked off, except near the caudal end, by another,
less regular furrow; each longitudinal ridge terminates anteriad
as a small, suprapedal lobe under the snout. The foot of
Htqqna (Happiella) guildingi (Bland) is fundamentally similar,
but the j)rojection above the mucous pore is very much longer,
as figured by Bland himself (1805, A. Lye. Nat. Hist., N. Y.,
VIII, 157-9, figs. 3).
From the above descriptions, it will be seen that Scolodonta
(subgenus Systrophielln) and Happia (section Happiella) cannot
be included in the Streptaxidae as they are apparently much
niore closely related to the Aulacopod families than to the
Holopod ones. In fact, they appear to have certain characters
in common with the superfamily Agnatha, but Scolodonta, at
least, differs in the close approximation of tlie cerebral ganglia,
which are practically fused as in the Agnathomorpha. For
these reasons, a separate family is necessary for at least Scolo-
05 THE NAUTILUS.
dontn and Happia; from the radulae and shells, I believe that
this should also include Austroselenites, Zuphos (?), Drepanostom-
ella, MartineUa (?) Miradiscops, Tamayoa and Guestieria.
The American families of Sigmurethra with purely carnivor-
ous dentition, all of which have short cerebral commissures,
may be separated by the following provisional key:
A' Foot aulacopod, bounded by pedal grooves and with a
caudal mucous pore; penial retractor a branch of
columellar muscle Scolodontidae
A" Foot holopod; penial retractor usually attached to dia-
phragm
B' Kidney oblong, parallel to pericardium; jaw well de-
veloped; radula with Zonitid laterals
Haplotrematidae
B" Kidney not parallel to pericardium; jaw obsolete;
radula with strictly aculeate laterals
C Kidney oval, transverse; right ocular retractor typ-
ically not associated with genitalia; shell helicoid
to discoid Streptaxidae
C" Kidney triangular, oblique; right ocular retractor
passes between penis and vagina; shell usually
elongate, often with truncate columella
Oleacinidae
The Scolodontidae may be further defined by the following
sets of characters. Shell: thin, transparent to translucent,
helicoid to discoid, with simple or emarginate peristome.
Kidney: very long; parallel to pericardium as in the Haplo-
trematidae. Jaw: vestigial (to functional?). Laterals of
radula: anterior ends narrow and elongate; cusps aculeate or
fimbriate.
The Streptaxidae certainly include Streptnxi.^ and Rcdartemon
(Artemon of authors); the other American groups are probably
Polygyratia^ Systrophia, Entodlna and Ridleya (?). Shell: heav-
ier and more opaque, usually with reflected peristome, often
with teetii or lamellae. Jaw: truly absent. Laterals of radula:
anterior ends expanded to form large areas for attachment;
cusps strictly aculeate. Although the venation of the lung is
THK NAUTILI'S.
89
very weak in the smaller species, it resembles that of the
Oleacinidae in at least one of the larger forms (Rectartanoii
lessei).
AN ABNORMAL CLAM
BY J. HENRY BLAKE
During my visit to Provincetown, Cape Cod, in August, there
was given to me a curiously deformed clam, Mya armaria Linn.
This specimen was dug with other normal clams from the
'* flats " at the western end of the harbor by Mr. Newcomb, a
native of the town. It was alive when taken and apparently
as " happy " as its neighbors.
Its striking abnormality attracted my attention and I have
90 THK NAUTILUS.
thought it of sufficient interest for the readers of The Nautilus.
The two illustrations here given, natural size, will show the ap-
pearance of the specimen better than any words I may offer.
The right valve is much smaller, quite flat, in one part a
little concave, and about half covered with a yellowish epi-
dermis. The left valve is extremely convex and thicker than
the right, its umbone projecting over the latter, and with slight
radiations on the anterior part.
One can easily contemplate the volition of some enthusiastic
paleontologist who found such a shell in a fossil state, and it
suggests a lesson to the hasty so-called species maker.
C0NCH0L0GIST8 IN THE MAKING
BY p:dwin e. hand
Hyde Park High School, Chicago, Illinois
As aftermath of the World War, it seems that very little can
be accomplished without "a drive." There's a drive for this
and a drive for that, a lot of advertising is done, a day set apart
and the trick is turned.
Since no one has started a drive for future conchologists we
are pegging away in the good old-fashioned way. If results
are inadequate, why not have a Drive ?
Given 150 young boys and girls about 14, to be taught gen-
eral science, how do we get them interested in the lowly snails
and their shells.
A tea strainer securely tied to a stick six feet long accom-
panies us to .Jackson Park, just south of the beautiful art build-
ing of World's Fair fame. On the bottom of the clear water
lagoon, within easy reach, lie thousands of Goniofxisia and Pleu-
rocera. On the bank, with eager, ex]>ectant faces, stand 40 of
these boys and girls seeing a snail for the first time in their
lives.
A few sweeps of the tea strainer and ( lutugh snails arf^ gath-
ered to place in their outstretclied hands all the specimens they
wish to take bonif and ])\i\ro in their "glass of water" acjuar-
riiK .NArTiLii.s. 91
iuins. Those are watched a few days and ])roudly introduced
to little brother or sister or friends. Afterwards the poor snails
are killed by dropping into boiling water, then in a few seconds
removed from their shells, the shells cleaned and mounted in
the cigar-box apartments. The names and recjuired data are
written in corresponding spaces on inside of cover; other kinds,
Plaiwrbis, land shells, etc., are added; trades are made, gifts
from friends begin to come in; a few sea shells are bought, and
Boon the boxes are on exhibition.
A new game has been tackled, and by following directions a
touchdown is scored. The collections are judged by votes of
their makers, and prizes are awarded. The tirst prize this time
was a Xenophora — the original shell collector — whose story
would have been unbelievable without the experience of the
senses of sight and touch.
Then comes the question of correlation of this work with
their other studies. What can be more noticeable than the
"Aa«</7j<^" in its three-fold complex; the shell, natural and
sectioned; the poem, so technically exact and so marvellously
musical, from which always some bright boy or girl can quote,
and the magazine (ours, not "new thought") absolutely new
to all ?
Yes, and books: Baker's "Molluscaof the Chicago Area";
Ward's " Catalog of Mollusca"; Rogers' "Shell Book," to be
ordered for one of their Christmas presents, and other books too
numerous to enumerate.
We correlate with history. The money cowry and its story
is in the boxes. Peden hnslatm "dredged from 50 fathoms off
San Pedro," gift of the Oldroyds, with its " bristling spears "
and coupled with its cousin jncohnevs^ carries us back to the
times of the crusaders.
The bleeding tooth, olives galore, Cypraea anmilns, with its
ring of gold, from the Bnsines.=? Manager of The Nautili's, and
a perfect shower of shells from Becker, A. J. Brown, Chace,
Farrer, Ferriss, Remington and others to come when they read
this — connect us with the greatest of all sciences — generosity or
is it Love? Many of these children are surprised when I tell
them that these shells are sent (if I am wrong, please correct
92 THE NAUTILUS.
me), because the donors want others to feel the joys they have
felt, not onl}' in receiving but in giving.
Finally, Purpura connects us with royalty and the Shell on
the Stamp of Travancore, to Philately.
Polygyra texasiana texasensis, obtained by the author at San-
derson, Texas, this summer, reposes in the boxes as a constant
reminder of Dr. Pilsbry's tussle with the naming problem.
Thus it goes on 3'ear by year.
Friend Higgins reminds us that "naturalists are born, not
made." Friend Johnson tells of Remington and Clench who
were drawn into the field of conchology by visiting the Boston
Society of Natural History. Our Mr. Conrad sent forth Dr.
Zetek, from his school and mayhap there be many others.
I may not have a single famous collector to my credit, but I
claim the honor of having given thousands of Chicago boys and
girls a new outlook on life; a love of nature's beauties from
stars to snails; a knowledge of great names in many realms, and
of books for their own libraries. But I suppose if we want real
rabid conchologists or anything else, we shall have to get up a
drive.
Hear, ye high schools, let's have a drive !
NEWFOUNDLAND SHELLS
BY E. G. VANATTA
Mr. Bayard Long collected the following species of Mollusca
while on a botanical exj^edition in Newfoundland. He noticed
Arion nler (L. ) at many places near I)uy Bulls. This is the
only American locality for the species, except one specimen
found in a garden at Detroit, Michigan, as reported by Dr.
Bryant Walker. The Arions were identified from dried speci-
mens by the jaws and radnloi and Mr. Long's notes. The
single (tonyodiscus a'onkhitei is not quite so angular as tyjucal
var. anthonyi l*ils. LymiKva pei-egrn gciserkola Beck, described
from Iceland, is also an addition to tiie American fauna. It
has been compared carefully with Iceland specimens.
THE NAUTILU8. 93
Vertigo hoppii Bk. (?) Brig Bay and Flower Cove.
PoliUi hammonis (Strom), Brig Bay.
Avion nter (L. ), Bay Bulls.
Arion fasciatus Nils., Trepassey and Whitbourne.
Oonyodiscics cronkhitei anthoni/i Pils.. Brig Bay.
Succinea avara Say, Brig Bay and Flower Cove.
Succinea oralis Say, Flower Cove.
Lymncea pahistris (Miill.), Sandy Cove and Flower Cove.
Lymnaea peregra geisericola Bk. , Junction Pond, Whitbourne.
Anodonta marginnta Say, Junction Pond, Whitbourne.
ADDITIONS TO THE LIST OF MARINE SHELLS OF SANIBEL, FLORIDA
BY WILLIAM J. CLENCH
University of Michigan
During the summer of 1923, a second trip was made to Sani-
bel, Florida. Unfortunately, this trip was made during the
same season of the year as the previous trip, and, as a conse-
quence, seasonal differences in the fauna were not observed.
However, a few additional species were obtained and more notes
on some of the species collected on the first trip was secured.
(Naut. Vol. 37, pp. 52-56, 1923.)
General conditions on the island were similar to those of
1921, except for a deposit of mud and silt along the gulf side of
the island, some four or five hundred feet off shore. Heavy
rains in the fall of 1921 caused the Caloosahatchee River to carry
an unusual amount of sediment into San Carlos Bay, which was
carried by the outgoing tides around the point of the Island and
deposited off shore. This caused a condition unfavorable to
many sand-loving species and, compared with their abundance
on the former trip, their absence was noticeable.
Many more trips were taken during 1923 to the northwest
end of the island in the region of Blind Pass and Clam Bayou.
Very large specimens of Bmycon perversus were collected in the
Pass, the species being especially plentiful around the beds of
Venus campechienais. Three large specimens of Pinna muricata
94 THE NAUTILUS.
and a single specimen of Conus proteus were found alive in shal-
low water at the entrance of Clam Bayou. The Pinna were
buried, hinge down, in the sand, with about an inch of the
shell protruding into the water, and were covered with a growth
of algae which concealed them very effectivel}'. A new part of
Clam Bayou was visited which yielded a very rich harvest of
Barnea costatn. These shells were located by small holes in the
sand which occurred usually in groups of from three to twenty.
They were buried about eight inches in the sand, their siphons
extending about halfway to the surface.
The following list is composed of new records and notes not
recorded on the previous list:
Pecten nodosus (L.). — A single small valve only, I have seen
specimens of this species accredited to Sanibel 5} inches long.
Mytilus recwfus Raf. — Tarpon Bay. A single valve only.
This was found with some debris on an oyster bar near the
wharf landing.
Venus campechiensis Gmel. — On my last list this species was
noted only from Clam Bayou. This year a few specimens were
taken in Tarpon Bay in about five feet of water. A large bed
of this species was found in Blind Pass, a strip of water sepa-
rating Sanibel from Captiva Island to the north.
Cyrend floridond Conr. — Very common. Tliis species was
only collected dead in 1923. However, on this trip, a small
bed was found in Tarpon Bay, located near high-water line
among the mangrove roots. It was peculiar to note that these
were especially abundant in the sand (1 to 2 inches deep) in
the little rivulets draining this swamp area. Similar beds were
located all around the bay, always near high-water line, under
the circumstances noted.
Tellina nlternata Say. — Three dead specimens of this species
were picked up at low water along the beach.
Cerithium Jluridanvvi ^locrch. — Gulf side of island. A single
specimen only.
Cantharus tindus Conr. — Tarpon Ba}'. On my previous list I
reported this species only from the gulf side of the island, and
inhabited by hermit cral)K. On this trip several specimens were
taken alive in llic hay on the oyster bar.
THE NAUTILUS. 95
Valuta junonia Hwass. — A splendid specimen of this species
was presented to nie by a retrident of the island. This was one
of eight specimens collected along the beach by him during the
winter of 1922-1923.
Crq)idula aculeala Gmel. — Gulf side. Two specimens only.
This seems to be a deeper-water form than the other species of
Crepidula collected on the island. Both of these specimens
were attached to the base of a sponge which was brought in by
a storm.
A great many more species undoubtedly exist on and about
this island. During both trips only a short time was devoted
to marine collecting, and more intensive work would reveal
many species not recorded so far, especially if it extended over
sections of the island along San Carlos Bay. During the winter
months many specimens are cast up on the beach that are never
seen at any other season of the year. A local collector had a
specimen of Dolium collected during the past winter (I believe
Dolium galea L.), and he described many other shells, but his
descriptions were too meagre in detail for specific determination.
NEW SHELLS FKOM JAPAN
BY WM. H. BALL
Mr. J. B. Langford of Honolulu, long a noted collector of
Hawaiian marine mollusca, has of late been domiciled in .Japan
where his activities have continued. His collections have been
chiefly littoral, and, since the earthquake, have been diminished
by the dislocation of the shoreline and consequent disappear-
ance of many of its normal population of mollusca. Most of
the species sent were familiar, but a few appear to be unde-
scribed, especially a remarkable Chiton, which, having been
submitted to Dr. Pilsbry, seems to typify a new group as well
as a new species. In his opinion it is most closely related to
the Placiphorella group.
96 THE NAUTILUS.
Langfordiella, new genus.
Form broad and short, little elevated, the girdle produced in
front as in PlaciphoreUa ; insertion plates short, thick, and
pectinated, the tail valve without slits. Type, Langfordiella
jcqjonica n. sp. U. S. Nat. Mus. Cat. No. 333541.
Langfordiella japonica n. sp.
Valves narrow and wide, much eroded in the specimens, a
young individual shows no defined jugal area, the general upper
surface of the valves smooth except for incremental lines and
two strong ribs extending to the margin; insertion plates in the
adult short, thick, strongly pectinated; in the anterior plate
about 12 groups of pectinations; intermediate valves with one
slit between coarse pectinations; tail valve heavy, with no slits
and feeble indications of pectination; sinus very shallow, hardly
defined. Interior of valves white, with a greenish tinge on the
anterior valve. Girdle brownish, leather}'', produced in front,
irregularl}^ studded with short stout brown spinules, often with
a swollen apex, and slender microscopically scaly hairs not dis-
tributed with any apparent relation to the sutures; gill rows
ambient, muzzle triangular. In a dry specimen the shelly
portion measures 32 mm. ; over all, with the contracted an-
terior portion of the girdle 39 mm. ; breadth of valves 31 mm.
The coloration of the upper surface is difliicult to determine,
owing to erosion, but in the youngest specimen there are red-
dish longitudinal streaks with pale intervals while the distal
portions of the valves are nearly black. One of the larger
specimens is more generally reddish, and another is almost
wholly blackish.
Locality. Kmi Busliiu, .Japan; others are from Nemoto
Bashiu.
Mr. Langford notes that the coast where these specimens were
collected was raised during the earth(|uake about ten feet. The
bottom is a soft rock in which the chitons made little circular
holes to roost in. These holes are very numerous but at pres-
ent all empty.
THK NAUTILUS. 97
Slilifer C(v>tanei'3 n. sp.
Shell small, smooth, naticoid with a short mucro; very dark
chestnut brown; whorls about three, exclusive of the mucro,
the last much the largest, somewhat appressed, the suture not
deep; surface with microscopic spiral striulae, and faint incre-
mental lines, base rounded, imperforate, pillar thin, arcuate;
aperture ovate, the margin thin. Height, 4.5; max. diameter,
3.3 mm. Type, U. S. Nat. Mus. Cat. No. 333545.
Locality, Emi Bashiu, Japan, Langford. Attached to an
echinoderm of undetermined species, which has disappeared
from the locality since the earthquake.
The color of this and the following species is unusual, the
greater number of the members of the genus being pure white.
Stilifer perdepressus n. sp.
Shell very dark chestnut, of about three whorls with prom-
inent mucro; the last whorl much the largest, much depressed,
suture appressed, surface smooth except for lines of growth;
base rounded, pillar thin, the upper part reflected over an im-
pressed umbilical dimple; aperture ovate, the outer lip obliquely
retractive. Height, 2.75; max. diameter, 3.0 mm. Type, U.
S. Nat. Mus. Cat. No. 333546.
Locality. On echinoderms at Chikura Bashiu, Japan, Lang-
ford.
Strombiformis langfordi n. sp.
Shell minute, yellowish, except the last whorl which is pale
with a peripheral yellow-brown narrow line; surface polished,
suture appressed: hardly evident: whorls about nine, the apical
whorl blunt, the shell slightly dextrally bent; the whorls flat-
tened, the base somewhat produced, imperforate; aperture sub-
ovate with the lip and pillar thin. Height, 5.0; max. diameter,
1.5 mm. Type, U. S. Nat. Mus. Cat. No. 333555.
Locality. Shimoda Izu, Japan, on Hcliocidaris cr(u>siirpina
(A. Agassiz), Langford.
98 THE NAUTILUS.
NOTES ON THE MACTEAS OF THE WEST COAST
BY A. M. STRONG
The Mactras of the Pacific Coast of North America have
always been a difficult group to collectors and students. Al-
most all the species are known to have a very wide range but as
they are seldom found above low-tide line and the thin, brittle
shells do not last long when washed ashore, but few good speci-
mens are to be seen in the collections. Even now some of the
species are very imperfectly known and there has been much
confusion in the application of the names given by the early
writers whose meager descriptions were based on poor material.
The literature on the group is not extensive in the works
commonly quoted. Gray, in the Mag. Nat. Hist., 1837, de-
scribed Mactra exoleta from the Gulf of California. In the same
year Conrad in the Journ. Acad. Nat. Sci., Phila. , vol. 7, de-
scribed Mactra planulata, Mactra californica and Cryptodon nuttallii
from shells collected by Nuttall on "muddy marshes bare at
low tide" at Santa Barbara. Three species were described by
Gould, Mactra falcata from Puget Sound in the Proc. Boston
Soc. Nat. Hist, for 1850 and Mactra nasuia from San Pedro and
Lutraria nndulata from La Paz in the Proceedings for 1851.
Gould in his Otia, p. 210, says of nasuta Gld., "This species
agrees so nearly with M. falcata Gld. , that until a direct com-
parison is made it may be considered synonymous." Car-
penter, in his review of these different species (Report of 1863,
p. 613) says: " Conrad's types being lost, and his species im-
perfectly described from very young specimens, a difficulty at-
tends their identification. Dr. Cooper found very large valves
(resembling Schizothaerus) in abundance, but much deformed
by the entrance of sand, and apparently killed by the fresh
waters of a great (lood. The large shells belong to two very
distinct species, wliich are probably those of Conrad; among
the small shells is perhaps a third, which may be Dr. Gould's
su]»j>resscd rja.st/to." Car|»('iitcr's application of the earlier
names was unfortunate and resulted in much of the later con-
fusion, particularly as seen in the labels in the older collectionB.
THE NAI'TILIS. 99
Connul in the Amcr. Journ. Conch., vol. 3, p. 193 (18G7)
<U'8criheti Mdctra dolahriformvi and Mactra catilliformis from
Panama. Both of these species are now also credited to Cali-
f(^rnia waters. Dr. Dall in the Nautilus, vols. 7 and 8 (1894),
redescribed several of the species and added Mactra heinphilli
from San Diego. In the same connection he guggested the
name of Mactra alaskana for the Alaskan shell ])reviously re-
ferred to Mactra falcata Gld. The next work on the group
appears in a review of the nomenclature of the Family Mac-
tridae by Dr. Dall published in the Trans. Wagner Inst. Sci.,
vol. 3, part 4 (1898), where these various species are listed
under genera, subgenera and sections with a number of changes
from the previously used arrangement. Finally in Bull. Dept.
Geol., Univ. of Ca!., vol. 9 (1916), Earl L. Packard reviewed
all the West Coast species, both living and fossil. The results
of this last work does not appear in the list of West Coast
Shells in Bulletin 112 of the United States National Museum.
We now have the following species recognized from the West
Coast whose ranges are reported to reach as far north as San
Diego.
Anatina undulata Gould, 1851.
This is the Lutraria undulata of Gould, the Raeta undulata of
Carpenter, and the Labiosa undulata of Arnold and Keep. The
range is given by Dall as San Pedro to Panama and it is fully
descril)ed in Arnold's Paleontology of San Pedro, p. 177. It is
a southern shell of which a few valves have been found washed
up on Southern Califoi'nia beaches and it is easily recognized
by the "concentrically undulated " surface.
Mactra (Mactrotoma) californica Conrad, 1837.
This shell is fully described by Packard and Arnold as well
as by Dall in the Trans. Wagner Inst, of Sci., vol. 3, part 4, p.
876. It is not the Standella californica of Carpenter or the
Mactra californica of Cooper but seems to be in part at least the
Standella nasuta of Carpenter. Dall gives the range as Neah
Bay to San Diego but Packard gives Mactra anguMa Desh. as a
synonym and as this was a Panama shell the range can be
100 THE NAUTILUS.
greatly extended. The species is not uncommon in Southern
California and living specimens are sometimes found on the
mud flats at low tide. It is the smallest of the Coast species,
an inch and a half or less in length and according to Arnold is
distinguished by its thicker shell, nearly central beaks and more
obtusely rounded anterior end.
Mactra (Mactrotoma) dolabriformis Conrad, 1867.
This species was redescribed by Dall in the Nautilus, vol. 7,
p. 138, and the description is copied in Orcutt's Molluscan
World, No. 1627. Dall gives the range as from Lobitas, Cali-
fornia, to Topolabampo, Mexico, but Conrad's type is given as
from Panama. It is a rare species of which a few specimens
have been dredged in Southern California waters. The shell is
much compressed, resembling Spisula falcata Gld., but with
more central beaks, more elongated shape, and larger, the adult
three and a half to four inches in length.
Mactra (Mactrotoma) nasuta Gould, 1851.
There has been a great deal of difficult}' in fixing Gould's
shell. He gives the localities in " Otia " as San Pedro and La
Paz. The shells to which Carpenter, Cooper, Keep, and others
applied the name are now considered to belong to other species.
Dall gives the range as San Diego,? West Columbia, and Old-
royd has recently. Nautilus, vol. 31, p. 97, reported the species
from Puget Sound. Packard in his paper, p. 278, lists the
shell and gives as synonyms M. californica Desh., M. fragilis
Cpr. , and M. deshnyesii Conr., all of which were from the Gulf
of California. Packard says of the shell, "The lack of sulcate
beaks, the pronounced gape, the more ventricous form, besides
several details of hinge, will serve to separate this species from
M. cnlifornicd Conrad with which it is often associated."
Mactra (Mactrella) exoleta Gray, 1837.
This is the Mactra exoleta of Gray, Carpenter, and Arnold and
the Mactra veritricosa of Gould and C. B. Adams. The range is
given by Packard as San Diego to Panama. There may be
some (question as to the accuracy of the San Diego record for
THE NAUTILUS. 101
tlie livinj; shell tlioujih it is well uutluMiliraud from there as a
Pleistocene fossil. Arnold gives ii full description, j). 175, and
says that it is easily distinguislied by "the sharply angular
trigonal shape, great convexity, elevated umbo, and sharp suit-
marginal posterior ridge."
Spisula (IIhmimactra) voyi Ciabb, 18GG.
While this species was first described from a fossil by Gabb
in his Pal. Cal. vol. 2, p. 24, Packard states, p. 283, that it is
represented by a living form in Dall's S. (polynyma Stimpson,
var. ) ala.-<kana. This synonymy would include in part the
Madra falcatn of authors. Dall gives the range for the living
shell as Arctic Ocean to Puget Sound and he described it in
Nautilus, vol. 7, p. 138, and vol. 8, p. 40.
Spisula (Hemimactra) catilliformts Conrad, 18G7.
This species was redescribed by Dall in the Nautilus, vol. 7,
]). 137, and the description is copied in Orcutt's Molluscan
World, No. 1625. It is the Standella califomica of Carpenter
and the Madra califomica and Mactra plamdata of Cooper. The
shell is well figured in Keep. The large, thin valves, four
inches or more in length, are not infrequently washed up on
the California beaches during storms. Arnold says of it, " dis-
tinguished by its large size, broad, almost oval outline, promi-
nent cartilage pit, long depressed area in front of the umbo, and
nearly equally rounded ends." Dall gives the range as Neah
Bay to San Diego but Conrad in his original description states
" Inhabits Panama."
Spisula (Hemimactra) hemphilli Dall, 1894.
This species was described from San Diego by Dall in the
Nautilus, Vol. 7, p. 137, and the description is copied in Or-
cutt's Molluscan World, No. 1626. The range is given by Dall
as San Pedro to San Diego. It is the largest of the Coast
species, reaching a length of six inches, and according to Pack-
ard is distinguished from catilllformis by the more elongated
shape, longer laminae, and more oblique pallial sinus.
102 THE NAUTILUS.
Spisula (Symmorphomactra) falcata Gould, 1850.
Dall in the Trans. Wagner Inst. Sci., vol. 3, part 4, gives this
species as the type of the subgenus Symmorphomactra, though in
the later Bulletin 112 of the United States National Museum he
lists it with the Hemimactras. Carpenter and Cooper included
in the species shells which are now considered to belong to
other species and Cooper considered M. nasiila Gld. a synonym.
It is a rather small shell, about two inches in length, and is
sometimes found at low tide near the entrances to the California
bays. The range is given by Dall as Puget Sound to Coronado
Islands. The species is described by Arnold, p. 176, who
states, "distinguished from the other species by narrower,
more acutely rounded anterior end, and interior radiating
ridges. ' '
Spisula (Symmorphomactra planulata Conrad, 1837.
This is a little known species. It is the Spisula j)/a?n(/a<a of
Dall in the Nautilus, vol. 8, p. 40, but not the Standella plan-
ulata of Carpenter or the Madra planulfita of Gabb, Cooper or
Stearns. Packard in his paper, p. 293, describes the species
and states that it is of the general size and shape of S. falcata
Gld. but flatter and more inequilateral. Dall gives the range
as Monterey to Cape San Lucas.
ScHizoTHAERUs NUTTALLii Courad, 1837.
This the Cryptodon tniUallii of Conrad, the Tresiis maximns of
Middendorf and Gray, and the Lutraria capax of Gould. It is
a well-known Coast shell, ranging from Wrangell, Alaska, to
San Diego, and has been fully described many times.
The synonymy and localities listed would give all these
species a vcr}' great range and raises a question either as to tlie
accuracy of the type localities or as to the correctness of apply-
ing the names to the California shells. In spite of the fact that
five of the species appear in the California lists as ranging to
Panama none of them appear in Zetek's recent list of Panama
shells. Until more is known of the off-shore fauna of the West
Coast of Mexico and Central America it will be very difficult to
answer these questions.
THE NAUTILI'S. lOJ
NOTES
Ckphalopods or Elephants on the Mayan Stel.k. — In
Nature for August 2, Dr. II. 0. Forbes discusses the supposed
elephant sculptures on the !?telic of Copan and Tikal, conclud-
ing that they are really conventionalized cephalopoda. The
illustrations given appear to support his contention, which
seems the more probable as squids are already known to enter
into the raytholog}' of the natives of Ecuador.
The muskrat in Czechoslovakia. — Fiber zibethicus was intro-
duced in Bohemia in 1906 as a fur animal. It bred with as-
tonishing rapidity, and now threatens the balance of the fresh-
water fauna. It is a serious enemy of the mussels, especially
the pearl mussel, Margaritana margnritifera, which it destroys
by gnawing through the shell over the posterior adductor
muscle. H. Prell, in Die Natunoissenschaften, Sept. 12, 1824^
has given an account of its ravages.
Friedrich Borcherding, — A sketch, bibliography and por-
trait appeared in Archiv fur Mollluskenkunde, 1924, pp. 58-66.
Borcherding died March 4th, at the age of 75. He was known
to American conchologists chiefly for his work on the Achati-
nellidre of Molokai, and his monograph of Carelia.
Lake Titicaca mullusks. — Since the publication of my paper
on Mollusks of Lake Titicaca (Proc. A. N. S. Phila., 1924, i)p.
49-54) I have seen a work in which certain species are illus-
trated: Eine Priihistorische Metropole in Siidamerica, by Arthur
Posnansky (Berlin, 1914). On plate II, of Vol. I, there are
photographic figures said to represent Palucleslrina culmineay
P. andecola, Ancylus tilicacensis and PlanoHm titicaccnsi.^, no
authorities given. The first two are the species so named by
d'Orbigny. The Planorbis is probably P. titicacensis of Clessin.
The '■'■ Ancylus^ ^ titicacemis is a Pisidium ! Whether it is ident-
ical with my Pisidium tiiicacense can hardly be decided, but it
is at least quite similar. The shells are mentioned on pages
104 THE NAUTILUS.
33 and 34, but without descriptions or references. A species of
Hippocampus is also figured from the lake. — H. A. Pilsbry.
AcANTHiNA. — In preparing my "Notes on the Acanthina
from California" published in the Nautilus for July, 1924, I
overlooked a recent paper published in the Proceedings of the
Malacological Society of London, August, 1918. In this paper
"On the Radula of the genus Acanthina, G. Fisher", by Rev.
A. H. Cooke, a new subgenus Acanthinucella is jjroposed for
the California shells and it is pointed out that Monoceros uni-
carinata Shy., has priority over Monoceros engonata Conr. Our
old friend Monoceros engonata Conr. therefore becomes Acanthina
(Acanthinucella) spirata Blainv., var. unicarinata Shy. — A. M.
Strong.
Sea Gull Drops Clam on Policeman. — The habit of sea gulls
to carry clams in their talons to a considerable height and drop
them on a hard surface to break, so that the bird can feed on
the bivalve, nearly proved disastrous to Abe Loche, a former
policeman of Atlantic City, N. J.
Loche was walking along the Boardwalk when one of the
gulls flew high above him and dropped the clam directly on
the man's head. He fell and had to be carried into a nearby
drug store for treatment. — Boston Globe.
Shells Wanted.— Mr. E. E. Hand, of the Hyde Park High
School, Chicago, would like to exchange with other High
Schools. Also to receive as gifts or will ])ay for common
species in lots of 50 or 100.
Helminthoglypta umbilicata cayucosensis. — A subspecies
or local race of H. vmhU'irata has been found by Mr. E. P.
Chace and Mrs. Chace on bluffs six miles north of Cayucos, San
Luis Obispo Co., Cal. "The exact locality is the face of the
ocean bluff almost within reach of the spray, near 'China
Camp,' A. R. Storni ranch. The snails were living mider the
beach aster and hen-and-chickcn plants. The blulT at this
THK NADTILUS. 105
point is about 40 ft. high, the lower half rock, tl)c upper clayey
loam."
This race clifTers from luiibilicaln by the smaller uml)ilicuR,
contained 11 to IG times in the diameter of shell, by the dark
color, bet^Yeen russet and cinnamon-brown of Ridgway (while
the typical //. umhilicnUi is from old gold to Isabella color).
When denuded it is very close to pecan brown. The banding
is similar, the dark band being darker than Ridgway's carob
brown. It is profusely malleate, as in umbilicata. Height 16,
diam. 25.5 mm., 6 whorls (type, No. 134843), or somewhat
smaller, height 14.7, diam. 22.3 mm. — H. A. Pilsbry.
Amblema neisleri nest located. — In 1918 Mr. L. S. Frier-
son sent me some money to be used toward finding the rare
Amblema neisleri (Lea). This was after he had examined a
specimen the writer sent him from Florida for identification,
and I think he stated it was the first specimen he had ever
seen. Well, I used his money several times over since then in
the endeavor to locate them, and it is only within the last few
weeks that a nice little lot of choice specimens was received
from a fisherman up on the Chipola River in northwest Florida.
These came accompanied with Amblema boykiniana (Lea) and
Elliptio incrassatus (Lea), very fine specimens of both. As long
as they last I will send a few specimens of the neisleri to anyone
desiring them, — T. Van Hvning, Director, The Florida State
Museum, Gainesville, Florida,
Physa cupraeonitens and Physa sayi warreniana in Hot
Springs. — Physa cupraeaaitens was described by Professor T. D.
A. Cockerell in 1889, from Wellsville Hot Springs, Colorado.
Since then it has been "lost" to science. Last September I
was able to stop for a few moments in passing the type locality
in an automobile, and found the species still abundant both in
the spring where it issues from the tunnel in the rocks and in
the rivulet that comes from the bathing pool just below the
spring. The water is quite warm, but, though it is called a hot
spring, I would hardly speak of it as really hot.
106 THE NAUTILUS.
One might easily suppose the small size of this species to be
due to its warm-water habitat, but that seems to be discredited
by another species in much hotter water that we found on the
same trip. In a spring a short distance above the bathhouse at
Mount Princeton (formerly Hortense) Hot Springs, southwest
of Nathrop, about twenty miles up the river from Wellsville,
we found in fair abundance a species determined by Dr. Bryant
Walker as Physa sayi loarreniana. They are smaller, but not
much so, than specimens from elsewhere. The water of this
spring was quite uncomfortably hot. In the still hotter water
of the big spring at the bath house we found no mollusks at all.
Unfortunately we had no way of taking the temperature of
any of these springs, hence cannot make as definite a report as
the records of mollusks in hot springs, especially in Europe,
cited by Dr. Brues in his recent paper on animal life in the
thermal waters of Yellowstone Park (Proc. Amer. Acad. Arts
and Sciences, LXI, 371-437, 1924). — Junius Henderson.
PUBLICATIONS RECEIVED.
Systematische Index der rezentex Cypr.f.id.e, Dr. Phil.
F. A. Schilder; Archiv. fur Naturgeschichte, 90 Jahrg. heft 4,
pp. 179-214, Berlin.
Kritisches Verzeichnis der rezenten und fossilen
Cypr.'EEN, von F. A. Schilder. Abh. d. Archiv. fiir Molkis-
kenkunde, Bd. 1, heft 2, pp. 1-192, Frankfurt am Main, 1924.
In the first paper above noted, Dr. F. A. Schilder has re-
viewed the whole family of Cypraiida^, in which he includes
also the Amphiperasidte and Triviidic. He divides this group
into five subfamilies, twenty-three genera, and fourteen sub-
genera. He includes Pedkidaria in the Amphiperasinre. He
gives a useful list of the fossil forms, the earliest of which,
Cypraorlns gemmelaroi, appears in the Jura. This is followed
by a condensed systematic Catalogue with the synonymy ap-
proved by tlie author.
THK NAUTILUS. 107
The second paper, after eight pugcs of introduction, give3
seventy-two pages to a bibhography of tlie literature relating to
the family, followed by an nlj)ha!)etical .synonymic list of species
and a two-page addendum of additional matter whicth r,iinc too
late to be incorporated in the original paper.
It will be seen from this brief summary that the aullior's
point of view is similar to that of Cossmann, in that it sub-
ordinates everything to a strictly conchological arrangement,
without reference to distinctions indicated by anatomical char-
acters, basing the classilication entirely upon the form of the
ehell.
Notwithstanding the fact that man}'' malacologists will take a
difYerent view, the data brought together with so much labor
cannot fail to be of service to any one interested in the classifi-
cation of the group, and to earn their gratitude to the compiler.
— W. H. Dall.
Marine Shells of Puget Sound and Vicinity. By Ida
Shepard Oldroyd (Publications Puget Sound Biol. Sta., vol. 4,
271 pages, 49 plates, 1924). This will prove to be one of the
most useful publications on the West Coast shells ever pro-
duced. Correlating descriptions and bringing the nomenclature
up to date of the fauna of a given area, is a useful and commend-
able piece of work, and will do much to increase the general
interest in conchology. The illustrations for the most part are
excellent. — C. \V. J.
New Species of Mollu.sks of the Genus Chilina. By
Wm. B. Marshall (Proc. U. S. Nat. Mus., vol. 66, art. 13, pp.
1-5, pi. 1, 1924). Seven new species are described.
New Mollusks from Santa Elena Bay, Ecuador, By Paul
Bartsch (Proc. U. S. Nat. Mus., vol. 66, art. 14, i)p. 1-9, pis.
1-2, 1924). Fourteen new species of Pyramidellidae and
Melanellidae are described.
New Species of Ferrissia from Lower California. By
Bryant Walker (Jour. Wash. Acad. Sci., vol. 14, pp. 430-432,
1924). Two new species, F. hayacnlifornica and F. occidentalism
both figured.
108 THE >fAUTILUS,
A DRIVE FOB THE NAUTILUS.
Mr. Hand's suggestion in his article about a Drive, seems
appropriate also to The Nautilus, for certainly "results are
inadequate." The Nautilus, like all scientific journals, was
"hard hit" by the Great War. We lost most of our foreign
subscribers, and the increase in the cost of printing making a
raise in the subscription price necessary, had unfortunate re-
sults at home: we now have fewer subscribers than in 1915.
In one of our appeals we spoke of discontinuing the paper
unless there was some improvement in conditions. After this
we noticed that several subscribers did not pay their subscrip-
tions. Finally when they did, one apologized for his delay,
saying, "I would have sent it before but thought you might
stop publishing." Compare this with an English friend who
always sends three years in advance. He has faith in The
Nautilus and in American Science.
There seems to be for Conchology a great field for missionary
work. There are seventeen states which are not represented on
the subscription list of The Nautilus. The figures for our
large cities are still more surprising: 41 of the cities in the
United States which have over 100,000 inhabitants, not one of
whom is sulliciently interested in Conchology to feel the need
of our valuable paper.
For the past thirty-five years the present editors of The
Nautilus have done the work and paid the deficits clieerfully.
These^deficits would have been larger but for the generosity of
a few contributors who have paid the bills for their illustrations.
Our last appeal, October, 1922, was written in rather a jocular
vein, to]^keep up our spirits. It was evidently taken as a joke,
few subscribers responding. The question of continuing The
Nautilus is now up to you. The Drive is on. IIV muxt futve
50 new mbacribers before May Id. The editors will do their part,
but all those who love to study the Mollusca vinst do theirs too,
otherwise the Nautihis nmcricaniiii cannot Iniild its 39th septum.
— II. A. P. AND C. W. J.
The Nautilus.
Vol. XXXVIII APRIL. 1925. No. 4
ON A TANGANYIKA BEACH
BY CHARLES HEDLEY
It is not the sea, one told oneself; but the other self insisted
that it was, pointing to the level horizon where sky and water
meet, to the waves rippling with the familiar song upon the
sand, to the wide beach overgrown with sea-convolvulus and
even to the sea gulls flying overhead. A glimpse of reeds and
papyrus growing off shore is a help to appreciate that this is
indeed a lake.
Not being the sea, to what can it be compared ? The lake
front at Chicago is recalled by the beach, the waves and the
horizon. But beyond the Tanganyika beach rise range beyond
range till the eye rests on an eight thousand foot mountain in
the blue and hazy distance. Happily does Burton's phrase of
"the lovliest glimpses of the Mediterranean" express it. But
against the Mediterranean background is set a foreground which
might belong to New Guinea or Tahiti, a group of blacks at
play upon the shore, hills sloping steeply to the sea, green with
banana plantations, dotted with brown thatched huts and
flagged with palms.
In clear weather, for now is the rainy season, the hills may
be seen across the lake which is only forty miles wide, though
it stretches north and south for 450 miles. The surface is
110 THE NAUTILUS.
2500 feet above the sea while the depth is incredible; four thou-
sand feet is a published record, but a Belgian officer informs
me (I know not on what authority) that it extends half as far
again to two thousand metres. The former record exceeds the
depth of any lake in the world except Lake Baikal in Siberia.
Lake Tanganyika has the narrowest zone of internal drain-
age, the few small streams entering it being incapable of car-
rying down much sediment. The exit is a small stream which
runs west to the Congo. This lake then has a promise of im-
mortality beyond any other lake whose destiny is either to be
filled in by tributaries or breached and drained by exit streams.
Yet it was once larger, for all around are inland cliffs composed
of lake deposit and separated by a level terrace from the water.
Lake Tanganyika lies in the Great Rift Valley of Africa.
This is a huge crack in the crust of the earth which dates back
at least to the Cretaceous epoch. It runs for several thousand
miles from the Jordan Valley in Syria through the Red Sea,
Lakes Rudolph, Tanganyika and Nyassa to Cape Corrientes in
South Africa.
Water in the lower strata of the lake is said to be salt. Since
more water is received and evaporated by the lake than is
spilled into the Congo basin, the salinity of the lake is probably
increasing. My neighbor at the hotel table tells me that even
the surface water is unwholesome and should not be drunk un-
less boiled. He also cautioned me to be careful when collecting
along the beach l)ecause there are numerous and ferocious croc-
odiles which often snatch and kill the negroes. So impatient
was I to see at last the wonderful shells of Tanganyika in their
home, that I could scarcely wait till I had secured a room at
the hotel and despatched a meal, before making for the lake
side. When I reached it there ai)peared along the water side a
long swathe of storm-tossed shells. These were, if I remember
right, the Neothaiima, large and solid gasteropods, some purple,
some yellow, some ovate conical, sonie with a broad sutural
furrow that recalled Kl)urna. This, which perhaps is a deriva-
tive of Vivipara, is a dominant form and continued in great
abundance as far as my promenade extended. It was obvious
that the molluscs lived close by within the sweep of the waves.
THE NAUTILUS. Ill
The exit of the Lukuga River was not far off and as it seemed
promising I walked across to it. Over a broad flood plain at
the river bank were scattered great quantities of shells. Among
them a large river mussel, Irldina, was conspicuous. On pick-
ing it up, there appeared a taxodont hinge, something strange
to find in a river mussel. Where the shells were thickest I sat
down to make a closer search. As I did so, I remembered
being warned at the hotel that just here was where the biting
flies might inflict the sleeping sickness on their victims and
resolved not to linger long. The first thing I found was a large
solid Melania-like shell, Paramelania, which brought the pic-
ture of it in Smith's paper at once to mind. Then I was re-
warded with one after another of its associates. A Nassoid
shell, Edgaria, was the most abundant; this seemed to exhibit
endless variation of color, form and sculpture. There may
have been several species in a handful that I gathered, but a
hasty glance could distinguish no consistent features. A glossy
white Naticoid was the next to attract attention, this and a
rarer Crepiduloid species (Spekia) seemed the most un-lacustrine
shells imaginable. Then I found a brown shell with a white
zone, for all the Avorld like a Planaxis, that on a coral beach
would have appeared quite conformable. Most eccentric of all
was a tiny white acicular shell (Syrnolopsis) like Eulhna but
with the plait of Pyramidella. These were quite common when
the proper place was reached. Not one Tiphohla did I see; its
absence is accountable by the deeper water that it is said to
inhabit. Mixed with the thalassoid shells were a few ordinary
fluviatile types, Lymnfea, Planorbis and Ampullaria. There
were also a few land shells carried down by the storm water.
This hasty glimpse at the Thalassoid fauna of Tanganyika
was ample reward for much wearisome travel. An impression
remained of the extraordinary dissimilarity between this and
any other fauna, and hence of its probable extreme antiquity.
The lake presents itself as a refuge where biological conditions
may have remained unchanged for a prolonged period. While
the thalassoids slept in Tanganyika their contemporaries may
have perished elsewhere through foreign competition and change
of temperature and of other physical conditions. Elsewhere liv-
112 THE NAUTILUS.
ing fossils like Nautilus and Trigonia linger here and there, but
in Tanganyika we seem to see not odd members but a whole
association of species preserved beyond the common span.
Albertvillej Tanganyika, Congo-Beige, Central Africa.
February 9, 1926.
NOTES ON THE NOMENCLATURE OF SOME OF OUR EAST AMERICAN
SPECIES OF PECTEN WITH DESCRIPTIONS OF NEW SPECIES
BY W. H. BALL
There has been in the past some confusion in regard to the
names of our more common species of Pecten, and in revising
the series in the National Museum, it seemed desirable to put
them on a sound basis by scrutinizing carefully the original
sources. This has led to the cleaning up of some doubtful
points as will be seen by the following data.
Peden (Placopecten) grandis Solander
Ostrea grandis Solander, Portland Cat., p. 50, No. 486; p. 99,
No. 2168, 1786.
Ostrea magellanica Gmelin, Syst. Nat., p. 3317, 1792.
Pecten tenuicostatus Mighels, Proc. Boston Soc. Nat. Hist. IV,
p. 41, pi. 4, fig. 7, 1842 (young shell).
Pecten fusais Linsley, Am. Journ. Sci., vol. 48, p. 276, 1845
(nude name); Gould, Am. Jour. Sci., 2nd ser. VI, p. 235,
fig. 6, Sept., 1848 (young shell). Not/u«cusof Gmelin, 1792.
Pecten brunneus Stimpson, New England Shells, p. 48, 1851
(new name for P. fusciis Linsley).
Pecten (Placopecten) clintonins Verrill, Trans. Conn. Acad. VI,
p. 261, 1884 (not of Say).
Pecten (Placopecten) grandis Dall, Trans. Wagner Inst. Ill, pt.
4, p. 727, 1898.
This species has been dredged in rather deep water from
Labrador to points off Cape Hatteras, North Carolina.
There is a marked difference between varieties of this species;
the young form has the radiating .''culpture minutely iml)ricate,
a feature which is sometimes jiersistent in tlie adult, while in
the majority of specimens the imbrication is wanting after ado-
THE NAUTILUS. 113
lescence. In color the species varies from pale to dark pur-
plish, rare individuals show dark and pale radiations.
Fecial (^Plagioclenium) gihhvs Linne
Ostrea gihba Linne, Syst. Nat. ed. X, p. 698, 1758.
Ostrm jiabellum Gmelin, Syst. Nat., p. 3321; 1792 (after Regen-
fuss. Conch. I, pi. 9, fig. 33).
Pcden lineolaris Lamarck, An. s. Vert. VI, p. 166, No. 12, 1819.
Pecten didocatus Say, Journ. Acad. Nat. Sci. Phila., 1st ser. II,
p. 260, 1822; Am. Conch., pt. 6, pi. 56, figs. 2, 2a, 1834
(Encycl. M^th. pi. 213, fig. 3, not named), Binney's wSay, p.
214, pi. 56, f. 2, 2a 1858.
Pecten purpuratxts Conrad, Am. Marine Conch., p. 10, pi. 2, fig.
1, 1831; not of Lamarck, An. s. Vert. VL p. 165, 1819.
Pecten (Plagiorlenium) gibbus Dall, Trans. Wagner Inst. Ill, pt.
4, p. 745, 1898.
Found in comparatively shallow water from North Carolina
to the Antilles.
This is the common Pecten of the southern shores from the
Carolinas to the Antilles. It varies in color to a remarkable
extent, but never reaches the size of its northern relatives.
Pecten sowerbi Guilding, 1826, is supposed to be one of the An-
tillean color varieties. The number of ribs in the typical form
averages twenty, none examined had less than eighteen and
only one out of 154 had as many as twenty-three. A specimen
from West Africa is exactly matched by one from Florida,
though the African form seems to show few of the brilliant
colors common with the Antillean varieties.
Pecten gibbm, variety amplkostatus Dall
Pecten gibhua, var. amplicottatna Dall, Trans. Wagner Inst. III.
pt. 4, p. 747, 1898. Texas Coast to Carthagena.
This is a larger and heavier form than the typical gibbits. It
has from twelve to fifteen strong flattish ribs and all the speci-
mens seen have the lower valve white and the upper valve of a
grayish color. It is also found in the Florida Pliocene.
After study of a multitude of specimens, I am now inclined
to separate specifically from P. gibbus the forms formerly re-
garded as variety nxtcleus and irradians. This was also Stimp-
114 THE NAUTILUS.
son's opinion, expressed in his unpublished manuscript in my
possession.
Pecten (Plagioctenium) irradians Lam.
Pecten irradians Lamarck, Ann. s. Vert. VI, p. 173, 1819;
Chenu, 111. Conch. Ill, p. 8, pi. 39, figs. 10, 11, 1843;
Sowerby, Thes. Conch., Pecten, p. 62, pi. 17, figs. 139, 140,
1842; Reeve, Conch. Icon., Pecten, pi. 6, fig. 26, 1852; W.
G. Binney's Gould's Invert. Mass., fig. 496, 1870. Not of
Potiez et Michaud, Gal. de Douai, II, p. 87, pi. 91, fig. 6,
1844.
Pecten concentrkus var. borealis Say, Journ. Acad. Nat. Sci.
Phila. II, p. 260, 1822.
Pecten concentricm Conrad, Am. Marine Conch., pt. 1, p. 8,
pi. 1, figs, 2, 3, 1831; Gould, Invert. Mass., p. 134, fig. 88,
1841; not of Say.
Pecten turgidus Sowerby, Genera, Rec. and Fossil Shells, pt. 31,
fig. 1, 1825; not of Gmehn, 1792.
Pecten (Plagioctenium) gibbus borealis Johnson, Fauna of New
England, Moll., p. 29.
Ranging from Nova Scotia to Florida and Texas, but abun-
dant only to the northward.
This is the characteristic New England form, large, thin
Bhelled, rather compressed, gray and white with rarely a dull
red, yellow or brownish variation, the ribs averaging 17 to 20,
the basal margin with prominently acute scalloping.
Potiez and Michaud describe this shell as having an imbricate
elevated thread in the channels between the ribs, which never
occurs in this species. Their text says it has twenty ribs, but
their figure shows twenty-eight.
In the south, perhaps by hybridization, the valves sometimes
take on a more inflated form and heavier shell, intermediate
between the northern form and the variety concentricus, usually
with a larger number of ribs than in the latter form.
Pecten irradians var. concentricus Say
Pecten concentricus Say, Journ. Acad. Nat. Sci. Phila., 1st ser.
II, p. 259, 1822.
Coast of New Jersey to Hog Island, Eastern shore of Virginia,
and South Carolina.
This form which agrees closely with Say's description, differs
THK NAUTILUS. 116
from the typical irradimis by being more inrtated, more solid,
with rather uniformly seventeen ribs, the lower valve is yellow-
ish, sometimes slightly zoned with gray, the upper pretty uni-
formly bluish gray and somewhat flatter, the incremental lines
prominent; the valves are white inside, sometimes blotched to-
ward the margin with reddish brown. Full grown specimens
have a length parallel to the hinge line of about 70 mm. , and a
diameter of 45 mm. This variety does not seem to have been
figured.
Pecten (Plagioctenium) nucleus Born
Ostrea nucleus Born, Index Mus. Caes, p. 92, 1778; Test. Mus.
Vind., p. 107, pi. 7, fig. 2, 1780.
Ostren turgida Gmelin, Syst. Nat. p. 3327, 1792 (after Chem-
nitz, Conch. Cab. VIII, p. 323, pi. 65, figs. 621, a-h, 1784).
(Not of Sowerby, Genera, pt. 31, pi. 1, fig. 1, 1825).
Pecten micleiis Sowerby, Thes. Conch., Pecten, pi. 14, fig. 25,
1842; Reeve, Conch. Icon., Pecten, pi. 22, fig. 89, 1853.
Florida Keys to Venezuela.
Having examined a very large number of these shells, I find
them exceedingly uniform in size and coloration, showing none
of the bright colors of young P. gihbus, but maculated gray and
white, sometimes both valves, but the lower valve is usually
pure white. The shells are well inflated and thinner than
5'oung gihbus of the same size.
Pecteni (Euvola) tereinus n. sp.
Shell small for the group, depressed, thin, pale, the colors
when present delicate; the lower valve with about twenty-five
low flattened ribs, with shallow not channeled interspaces, or
with the ribs obsolete altogether, or ill defined; concentric
sculpture weak; the ears subequal, hardly sculptured, the bys-
sal notch shallow, with no ctenolium; internally the valve is
white, the grooves corresponding to the ribs, extend over the
whole disk and are well defined; upper valve flat or slightly
concave, with about 19 narrow elevated not dichotomous ribs
with wider interspaces; the concentric sculpture well marked;
this valve is usually more effectively colored than the other;
internally it is grooved like the other; the outline of the shell
116 THE NAUTILUS.
is nearly circular, the hingeline short, the cartilage pit very
small; height 28; length of valves 30; of hingeline 19; diam-
eter 4 mm. U. S. Nat. Mus. Cat. No. 333632.
Miami Florida, to the Bahamas and south to Cape Roque,
Brazil. The type specimen was dredged off Key West at station
3318 of the U. S. Fish Commission, in 45 fathoms, coral, bot-
tom temperature 75° F.
This species differs from P. raveneli, its nearest relative, in
the obsolescence of the external ribbing, the absence of chan-
nelling and color in the spaces between the ribs; in the de-
pressed form, and in the production of the internal grooving
over the whole disk, while in P. raveneli it is confined to the
more marginal portion. This latter feature is probably due to
the thinner shell in P. terdnus. The coloration is uniformly
more delicate and paler.
Pecten {Euvola) raveneli Dall
Pecten (Euvola) raveneli Dall, Trans. Wagner Inst. Ill, pt. 4, p.
721, pi. 29, fig. 10, 1898 (Pliocene).
Pecten niedius of American catalogues.
The Antillean shell which has been commonly identified with
P. medius Lamarck, is really a different species.
Ostrea (Pecten) medius Gmelin (Syst. Nat. 3330), based on
Gualtieri's figures N, O, on plate 74, is really unidentifiable,
but bears some resemblance to Pecten fricatus Reeve. Pecten
medius Lamarck (An. s. Vert. VI, p. 163, 1819), is basel on
Chemnitz (Conch. Cab. VII, pi. 60, figs. 586-7) who states
that his shell was received from Forskal who collected it in the
Red Sea. The name medius in this connection is preoccupied
by Gmelin. Sowerby in the Thesaurus states that his P.
medius Lamarck, was labeled P. latico»tatus in the British
Museum collection. But this name is })reoccupied by T>amarck
for a fossil species, "the largest known Pecieu," a giant eight
inches in breadth. None of the figures attributed to P. medius
Lamarck can be identified with the true West Indian shell;
Reeve especially (Conch. Icon., fig. 44) gives a large figure
which is neither Lamarck's P. medius nor a West Indian species.
Apparently none of the authors who have copied references
THE NAUTILUS. 117
from one another and applied them to various different species
were acquainted with our shell, at any rate they have not fig-
ured it. A description follows, since the Pliocene form has lost
its color.
Shell of moderate size, thin, the lower valve much inflated,
the upper valve slightly concave or nearly flat; the latter with
a straight hingeline and almost exactly equal ears, each with
four or five subequal minutely imbricate radial ridges; radial
sculpture of the disk about twenty-five low-rounded ribs with
sub-eijual interspaces in which toward the margin traces of in-
cipient intercalary threads may appear; the concentric sculpture
is of prominent fine low lamellae overrunning the whole sur-
face; internally the ears are prominent, the margin deeply radi-
ally grooved, the grooves extending about halfway over the
disk, the wider interspaces flattish, with paired elevated lines
at their margins; the hingeline is finely cross-striated, with a
groove on each side of the cartilage pit to receive a slender
lateral ridge from the opposite valve. The lower valve is pale
with about twenty-six strong flattish radial ribs with narrower
channeled interspaces faintly marked with concentric incre-
mental lines; the ribs toward the margin are in adult shells
divaded by a shallow median groove; in many specimens the
interspaces between the ribs are marked with a conspicuous
brown line; the ears are subequal, somewhat reflexed and feebly
sculptured; there is only a shallow byssal notch without a
ctenolium; the internal grooving resembles that of the other
valve but extends somewhat further toward the umbo; the
basal margin is notched by the sculpture; height of shell, 40;
length, 47; length of hingeline, 24; diameter, 12 mm. U. S.
Nat. Mus. Cat. No. 333634.
The shell described was collected by Captain Jefferson Moser,
U. S. N., at St. Martin's Reef, Florida, in three fathoms. The
range of the recent form is indicated by specimens from North
Carolina and Bermuda to Porto Rico, in the Museum collection.
The coloration is mostly white and pinkish, often with
brownish lines in the grooves of the lower valve. The interior
is usually white, with more or less color around the margin.
The Pliocene form to which the name raveneli was first ap-
118 THE NAUTILUS.
plied is somewhat coarser than its recent representative, but I
think there can be little doubt of their specific identity.
Pecten (Chlamys) omatus Lamarck
'iOstrea sauciata Gmelin, Svst. Nat, p. 3328, 1792, after Chem-
nitz, Conch. Cab. VII, pi. 69, fig. H, 1783. Dillwyn, Rec.
Shells 1, p. 161, 1817.
"IOstrea crenvlata Gmelin, Syst. Nat. p. 3329, 1792, after Lister,
pi. 175, fig. 12a. (left).
Pecten omatus Lamarck, An. s. Vert. VII, p. 176, No. 50, 1819
Conrad, Am. Marine Conch, pt. 1, p. 12, fig. 3, 1831
Sowerby, Thes. Conch., Pecten pi. 14, figs. 77-78, 1842
Reeve, Conch. Icon., Pecten, pi. 19, fig. 68, 1853.
Chemnitz's figure 69 is poor and the species can hardly be
regarded as identifiable. Ostrea crenulata Gmelin is referred by
Dillwyn to the P. radula of authors, the largest figure on Lister's
plate: but Gmelin' s reference is apparently for the left lower
figure of Lister, which seems to represent our shell. However
some doubt must attach to this identification.
The first positively identifiable name for this species is
Lamarck's omatus for which he cites the same figure of Lister
as that used by Gmelin for 0. crenulata.
This shell is white, elegantly sparsely spotted with purple,
the ribs are usually worn smooth and rather strong, though
originally they were covered with extremely fragile cup-like
iml)ricating lamellae which are very rarely intact. Of the 20
ribs, three are usually untouched by the purple color. The
posterior ear is almost obsolete. Our largest specimen is about
26 mm. in height.
Pecten (Cfdamys) senlis Reeve
? Pecten sentis Reeve, Conch. Icon., Pecten, pi. XXIX, fig. 125,
1853.
Pecten omatus (var.) most lists of Antillean Pectens.
This is a compjion south Florida and West Indian shell, usu-
ally referred to as Pecten ornatii.s though perfectly distinct when
carefully studied. The shell is usually bright scarlet or dull
purjde, i)aler toward the umbones, very rarely white or white
with maculations or rays. When fully adult it has about forty
THE NAUTILUS. 119
fine ribs, more or less alternated, which when unworn carry
minute erect low scales which bridge the interspaces and con-
nect laterally with those on the adjacent ribs to a greater or less
extent. This sculpture is lost, but leaves traces on most beach
specimens. The posterior ear is hardly developed, the anterior
much larger. The interior is white, grooves reflecting the ex-
ternal sculpture. The byssal notch is wide and shallow. Both
valves are usually similarly colored. Our largest specimen is
36 mm. high, 80 mm. wide, and the j)air has a diameter of
about 10 mm.
Reeve's shell is obscurely figured and insufficiently described,
without a habitat, but as far as the characters given go, they
agree with those of this species.
Pecten (Chlamys) liocymatus n. sp.
Shell (right valve) small, oblique, moderately convex, with
about 20 low rounded entire ribs with narrower shallow inter-
spaces; the surface somewhat worn, is smooth except for con-
centric lines of growth which may originally have been more
prominent; profile subcircular, the ears small, subequal, with a
few radiating threads on the anterior ear; the byssal notch
shallow; the interior partly grooved in harmony with the ex-
ternal ribs, the edges of the flat interspaces threaded; hinge
line straight, in the right valve with a groove each side of the
resilifer to receive a corresponding ridge from the opposite valve,
these grooves are sharply cross-striated; height, 30; width of
shell, 28; of hinge line, 16; diameter (of right valve), 5 mm.
U. S. Nat. Mus. Cat. No. 62290.
Dredged by the U. S. Fish Commission steamer Albatross at
station 2604, in 34 fathoms, sand, bottom temperature 79° F. ,
39 miles off Cape Hatteras, North Carolina; two right valves.
These specimens, perhaps originally white, have been dis-
colored by the action of the sea water. I have not been able to
find any species recent or fossil which is at all closely related to
this one.
Pecten (Aequipecten) hdiaaui n. Kp.
Shell of moderate size, more or less mottled and zoned in
120 THE NAUTILUS.
pinkish and crimson on a white surface, outline subcircular,
verj-^ shghtly oblique; ribs about 20, not dichotomous distally;
narrow, almost keeled, with wider interspaces; valves equal, in-
equilateral, the ears subequal, the byssal notch wide and shal-
low, with a ctenolium of four teeth; the right valve somewhat
paler than the other; the microscopic surface sculpture com-
prises a row of small pointed scales on top of the ribs, a smaller
series on each side of each rib and 3 or 4 much finer prickly
threads in the bottom of the channels between the ribs; the in-
terior of the valve is more or less clouded with brownish purple,,
grooved to correspond to the external ribs, the ends of the tlat-
tish interspaces slightly excavated, the resilifer rather large;
height, 38; length of valves, 40; of hinge line, 41; diameter,
11 mm. U. S. Nat. Mus. Cat. No. 333635.
Bahamas, fide W. F. Webb.
This species has the general appearance of P. phrygium Dall,
with which it had been placed in the collection, but the micro-
scopic surface is quite different.
Pecten (Aequipecten) acanthodes n. sp.
Shell of moderate size, subcircular, olivaceous brown with
small paler motthngs; with 22 minutely spinose ribs, the spines
in three subequal rows, with two or three smaller prickly
threads in the narrower interspaces, the spines are excavated on
the dorsal surface, more prominent laterally and on the upper
border of the hinge-line; ears subequal, the anterior slightly
larger, both with about five radiating spinose threads; byssal
notch shallow, the ctenolium with eight teeth; internally the
color and sculpture reflect that of the exterior; the flat inter-
spaces between the grooves marginally threaded; hinge-line
cross-striated and feebly longitudinally grooved; height, 32;
width of shell, 32; of hinge-line, 26; diameter, 11 mm. U. S.
Nat. Mus. Cat. No. 333697.
Range from Southern Florida to the Antilles.
Than P. exasperatus this shell is less inflated, wider, and with
less prominent spines. All the si)ecinHMis seen are uniformly
colored.
THE NAUTILUS. 121
NOTE ON THE NAME OF OUB COMMON OYSTER
BY W. H. DALL
It seems that Solander's name of dongaia is preoccupied by
Born for a different species, so that we can fall back on the
familiar virginica. The synonymy is as follows:
Oatrea virginica Gmelin, Syst. Nat. p. 3336, 1792.
Ostrea elongaUi Solander, Portland Cat., p. 55, No. 1303, p.
151, No. 3312, 1786.
Ostrea virginiana 'Bolien, Mus. Bolt., p. 169, 1798.
Not Ostrea elongata Born, Index Mus. Caes., p. 92, 1778;
Test. Mus. Vind., p. 103, pi. 6, fig. 2, 1780.
IDENTITY OF PATELLA TENITILIRATA CARPENTER
BY J. R. LE B. TOMLIN
Carpenter described this limpet as P. (? toreuma) tenvUirata
in the Proc. Zool. Soc. London for 1855, p. 233, without figure,
on the strength of specimens in the Cuming collection said to
have been collected by Hartweg at Monterey.
The type-lot of three specimens is now before me, and if the
shell had been figured there would probably have been little
hesitation on the part of subsequent monographers in assigning
it to its true position. Carpenter's locality and his tentative
reference of his supposed new species to P. toreuvm Reeve have
completely camouflaged the situation.
The three shells are very ordinary, rather worn examples of
Cellana radians GmeVm, the common New Zealand limpet. The
British Museum has more than one series of the real P. toreuma
Reeve, ex coll. Cuming, also labeled Monterey. This of course
is equally erroneous.
122 THE NAUTILUS.
SOME LAND SHELLS FROM THE ALEUTIAN ISLANDS, ALASKA
BY G. DALLAS HANNA
The California Academy of Sciences has two fairly extensive
collections of land shells from the Aleutian Islands, Alaska,
among which there are several species not hitherto recorded
from the region. Since their distribution has an important
bearing on some biological studies the list seems worth publish-
ing.
One collection was made by Dr. E. C. Van Dyke, entomolo-
gist of the University of California. In 1907 he spent consider-
able time on Unalaska Island and to the westward, and his
success in finding minute species is not often duplicated among
professed conchologists.
The other collection was made by me in the vicinity of Un-
alaska at various times when enroute to or from the Pribilof
Islands, 1913-1920.
The northwestward extension of what Van Dyke* has called
the Vancouveran fauna is particularly worthy of note. What
he has found among the insects appears to be true also of the
land shells.
It is true that Dr. Dall * suspected the occurrence of Pohjgyra
Columbiana and Circinaria vancouverensis at Unalaska to be due
to the transplantation of spruce trees from southeast Alaska,
but the wide distribution of these snails and other species in
that vicinity makes this explanation seem doubtful. If a sim-
ilar situation did not exist among the insects the accidental
planting theory would suffice, perhaps; but it seems very im-
probable now that so large a number of species of animals
would have been thus introduced with the few trees which were
brought.
Special search among the small groves of spruce trees on
' The DiRtribution of Insects in Western North America, by Edwin C.
Van Dyke, Anns. Knt. Soc. Araer. Vol. 12, 1919, pp. 1-12. Proc. Calif.
Acad. Sci. 4th ser. Vol. 11, 1921, p. 157.
' Dall, Nautilus, Vol. 31, 1917, p. 13.
THE NAUTILUS. 123
several different occasions failed to reveal any land shells ex-
cept Succineti, Vitrimi and Vertigo. The others were found
ahout the roots of herbaceous vegetation and grasses of the roll-
ing hill country. Dr. Van Dyke's finding of the Polygyra at
Makushin Bay many miles removed from the transplanted
trees strengthens the belief that this and the other species have
lived on the Aleutians for a long period of time.
Succinea chrytns Westerlund.
Makushin Bay, Unalaska Island; Glacier River, Unalaska
Island; Unalaska town; Amoknok Island near Dutch Harbor;
Nazan Bay, Atka Island (Van Dyke).
At these localities the species has not attained the large size
and beautiful golden color of the shells from St. Paul Island,
200 miles to the northward.
Vertigo modesta Say.
Makushin Bay, Unalaska Island; Unalaska town; Unalaska
Island; Dutch Harbor, Amoknok Island; Nazan Bay, Atka
Island (Van Dyke); Expedition Island near Unalaska; Hog
Island in Broad Bay near Dutch Harbor (G. D. H.).
The large number of specimens from these localities shows
considerable variation and a strict identification would probably
necessitate putting some of them under the named varieties but
my study does not seem to warrant this disposition at present.
Columelln dltwola (Ingersoll).
Near Unalaska, Unalaska Island (G. D. H. ). This appears
to be the large boreal form rather than the shorter edentula of
the eastern United States,
Cirdnaria va-ncouverenMs (Lea).
Near town of Unalaska, Unalaska Island; Makushin Bay,
Unalaska Island; Amoknok Island (Van Dyke); Expedition
Island near Unalaska; across straight from Unalaska wharves,
Amoknok Island and near town of Dutch Harbor, same island
(G. D. H.).
My attention was first called to this species as a resident of
124 THE NAUTILUS.
the Unalaska region by Dr. Alex Wetmore, then of the U. S.
Biological Survey, who found it there in abundance in 1912.
Precise directions led me to the same spot in 1916 where large
numbers of the species were found. Many of them had been
broken and the animal removed, presumably by fieldmice.
Subsequently the species was found widely distributed in the
region and abundant in favorable localities.
Pyramidula cronkhitei (Newcomb).
Unalaska town, Unalaska Island; Dutch Harbor, Amoknok
Island; Nazan Bay, Atka Island (Van Dyke); Hog Island in
Broad Bay near Amoknok Island (G. D. H.).
The species is very abundant in some places and appears to
be the same as the form found in the higher mountains of Cali-
fornia and northward.
Punctum conspectum (Bland).
Found sparingly on Amoknok Island, at numerous places by
both Dr. Van Dyke and me. It has been listed from Unalaska
previously.'
Polygyra columbiana (Lea).
Unalaska Island, several places about Bay; Makushin Bay,
Unalaska Island; Amoknok Island, several places (Van Dyke)
(G. D. H.).
Many live specimens collected do not appear to differ from
those found far southward in the range of the species.
Euconolus fuhus alaskciisis Pilsbry.
Unalaska Island, at town; Amoknok Island, near Dutch
Harbor (Van Dyke) (G. D. H.); Makushin Bay, Unalaska
Island; Nazan Bay, Atka Island (Van Dyke); Hog Island in
Broad Bay near Amoknok Island (G. D. H.).
Polita binntyana (Morse).
Unalaska Island, near town; Amoknok Island, near Dutch
Harbor (G. D. H.).
' Dall, Harriman Alaska Expedition, Vol. 13, 1910, p. 53.
THE NAUTILUS. 125
Vitrina ala^kana Dall.
Unala-skji Island, near town; Amoknok Island, near Dutch
Harbor (Van Dyke) (G. D. H.); Makushin Bay, Unalaska
Island; Nazan Bay, Atka Island (Van Dyke); Hog Island in
Broad Bay, near Amoknok Island (G. D. H. )•
Pristiloma arctica (Lehnert).
Near Dutch Harbor, Amoknok Island (Van Dyke) (G. D.
H.); Near town, Unalaska Island; Expedition Island near
Unalaska (G. D. H.).
Zoo genius harpa (Say).
Near Dutch Harbor, Amoknok Island; near Unalaska town,
Unalaska Island; Hog Island in Broad Bay near Amoknok
Island (G. D. H.).
Prophyaaon andersoni (J. G. Cooper).
Near town of Unalaska, Unalaska Island; many places on
Amoknok Island (G. D. H.).
Two color phases of what appears to be this slug are very
abundant on both islands mentioned. In the early spring they
were collected in large numbers crawling over the dead vegeta-
tion where the snow had melted.
LTHN£A AUBICULABIA (LINH.) IN CALIFORNIA
BY G. DALLAS HANNA AND H. WALTON CLARK
The finding of Lymvsen anri/ndarin in California during 1024
extends the range of this exotic form several hundred miles to
the westward. Its first occurence in the state was noted in a
pool in the Japanese tea garden, in Golden Gate Park, San
Francisco (H. W. C. ). Later it was found in abundance in
the fountain pool at the resort known as Byron Hot Springs,
Contra Costa County, California (G. D, H.).
Both of these localities are such to indicate that the molluska
were " planted " in some manner but there is no way to trace
positively the source of the original stock.
126 THE NAUTILUS.
A clue to the manner of introduction may be found in the
fact that it is one of the favorite aquarium snails, and is sold in
considerable numbers by dealers in aquariums and aquarium
supplies under the name of "African or Paper-shelled Snail. "
The same people supply aquatic plants, and snail eggs might
be easily distributed by these.
An analogous case is furnished by the discovery of several
small "Cumberland Terrapin" Chrysemys elegans, one of the
handsomest of our aquatic turtles, and native to the central
states drained by the Mississippi (Indiana, Kentucky, etc.).
Large examples of these are found in the markets of Chicago;
smaller ones are sold at San Francisco by dealers in pets, under
the name of "Japanese" or "Coin" turtles. Many purchas-
ers, finding their pets not thriving in captivity, or becoming
tired of them, sometimes bring them as donations to the Stein-
hart Aquarium, or apparently release them near or at the pools
in the park where they can often be seen basking on stones.
All that have been collected thus far had not reached sexual
maturity, so that the species may be said to be introduced but
not naturalized.
It is probable that Lyimi«a aiiricidaria may have been carried
from place to place in the same manner and for the same
reasons. The species is sufficiently known to attract atten-
tion still. It is worthy to note that all of the occurrences are of
the species living under artificul surroundings. Who will be the
first to find it established in a natural body of water? Our
specimens are large and handsome, being over 25 mm. in alti-
tude. The periostracum is a light straw color and the shells are
much thinner and more diaphanous than the specimens com-
pared from England.
Records of this species in North America are scarce. Baker '
in 1911 listed it from a greenhouse in Chicago, Illinois, and
from Brooklyn, New York. Hannibal in 1912 ' recorded it
from Utah but })robably through errors in synonymy. In the
same year Henderson * did Ihul the species in an artificial pond
* LyranKidre of North America, 1911, p. 182.
M'roc. Mai. Soc. Ix)ndon, Vol. 10, 1912, p. 140.
> Nautilus, Vol. 26, 1912, p. H4.
THE NAUTILUS. 127
in Colorado Springs, Colorado. Later ' he added Dotson Reser-
voir near Fowler, Colorado, to the known hHalitiff.
California Acadetny of Science,'',
San Francisco, California.
VAGABONDING FOR SHELLS
BY P. S. REMINGTON, JR. AND W. .1. CLENCH
The authors have been impelled to write this article because
they feel that they have accomplished something of value to
conchology, and because they hope that their account of it may
induce others to attempt similar trips. The " expedition " was
made under the auspices of the Museum of Zoology at the Uni-
versity of Michigan, the expenses of the trip being borne by the
Museum and by Dr. Bryant Walker of Detroit. The greater
part of the material collected will eventually go to the collec-
tions of the above named. Thanks are due to Dr. Walker and
to Mr. Calvin Goodrich for help in the way of advice, sugges-
tions as to the best places to collect, and gifts of maps and
other equipment. A formal report of the species collected will
come out later.
The purposes of the trip were several. In the first place, we
wanted to collect as large a series of Physa as possible to pro-
vide more material for Clench's doctoral thesis, which is on the
Physa of the U. S. east of the Rocky Mts. We wished to get
our Physa in alcohol for anatomical study, for there is a scarcity
of such material. Secondly, we were anxious to collect through
the Tennessee River system, particularly the upper portion,
which neither of us had ever seen, and which is one of the
richest collecting fields in the country. We also had a list of
various places, furnished us by Mr. (ioodrich, which was desir-
able to collect in either because they were new territory, or be-
cause they were the locality of certain doubtful species. Most
of these we cleaned up. Last, but not least, we were out to get
'Nautilus, Vol. 32, 1918, p. 71, and Univ. Colo. Studies, Vol. 13, No. 2,
1924, p. 162.
128 THE NAUTILUS.
some los. The authors collected shells together as boys, and
at that time one of the things we promised ourselves to do some
time was to go on a hunt for los. This trip promised an oppor-
tunity of doing that and of renewing our boyhood memories.
Before leaving, Clench prophesied that this year would see the
last of the los! It wasn't quite so bad as that, but we did suc-
ceed in taking several hundred. There are undoubtedly a few
left in the French Broad River, which we did not touch. We
had carte-blanche as to where we might go, but were guided
largely by the above considerations. We left Ann Arbor on
July 19 and returned Sept. 8.
In our choice of equipment we were able to profit by the re-
sults of a similar trip made the preceding summer by Clench.
We bought a 1922 Ford touring car in excellent condition and
proceeded to load up the running boards and back seat until it
looked like a gypsy outfit. We were handicapped by the fact
that we had to carry full camping equipment and supplies as
well as the collecting kit. We had a very handy auto tent,
two folding cots, two folding chairs, and a folding table. This
last was bought because of Clench's experience the summer be-
fore, and it was certainly useful. In our grub box we carried
cooking utensils and canned food, mainly beans. Both the
authors were originally from Boston. We had another large
case containing wading boots, sieves, jars, boxes, vials and
bags, of which one can't have too many on such a trip. A
duffle bag for each of us, a suit case of "civilian" clothes, a
dispatch case for papers, a case for toilet articles, a lantern,
blanket rolls, water jug, emergency can of gasolene and one of
oil, and a container of alcohol filled up the back. We had a
very fine two-mantle gasolene lamp which was invaluable.
Although the museum appropriation for this trip came largely
from the conchology de])artment, we received contributions
from the insect and reptile departments, so had to carry along
equipment for collecting such specimens. The insect equip-
ment was limited to nets and cyanide jars, hut we had to take
a large milk can partly filled with formaldehyde to preserve
rejjtiles. The reptiles were put in this to harden and preserve
them, then injected and wrapped in gauze and shipped back.
THE NAUTILUS. 129
We also had a plant press and a minnow seine. All these
things gave us license, when asked our business, to say we were
collecting '' plants and animals " for the University of Michigan.
This was all right till one obliging fellow offered us a pet wood-
chuck! Then we hit upon the happy phrase " we are making
a Biological Survey." They knew what a survey was anyway.
With all this mass of equipment aboard you would say we
must have spent half the day packing and unpacking. It is
true that it did take us an hour or more for this operation at
first, but before long we evolved a system, so that by the end
of the trip we could pack in less than fifteen minutes and make
camp in less time than that. We were surely a spectacle and
got many a kick out of the reactions of the beholders. Vaga-
bonds indeed!
One of the most enjoyable incidents of the whole trip came at
the very start. We left Ann Arbor about nine in the morning
•'heading south." We collected a little, mainly Physa, and
reached Toledo in the afternoon, where we hunted up Mr. Cal-
vin Goodrich. He very kindly insisted on our putting up with
him, so we had an opportunity to inspect his large collection of
Pleuroceridae and "talk shop." As the catch where we were
going would be largely of that family, we wanted all the
" dope " we could get. We had a very enjoyable visit, all too
short, for, as Mr. Goodrich remarked, it isn't often that three
shell collectors get together.
The next day, Sunday, we set out bright and early bound
southwest for Bluffton, Ind. We had heard unfavorable news
from Mr. Goodrich that all the rivers in the south were swollen
from heavy rains, and that unless they went down we would
see very little good collecting. Our original intention was to
strike south through Cincinnati and Cumberland Gap to the
headwaters of the Tennessee River, then work down to Hunts-
ville, Ala. and then north to Louisville. Upon receipt of this
unwelcome news, we decided to reverse the itinerary, so as to
reach the heavy collecting last, in the hope that by then the
rivers would be low. This turned out to be a wise move.
We camped the first night near Hicksville, Ohio, a half-mile
ofif the main road. We can hear the ringing reply, even now,
130 THE NAUTILUS.
of the farmer on whose land we asked permission to camp.
" You bet your life you can! " he said. We want to say right
here that if the camper is neat-appearing and respectful, and
leaves no rubbish or disorder of any kind behind him, he will
have no trouble finding a place to camp, and we were not once
refused in the whole 1700 miles we traveled. It is usually
better to go a little off the main road before camping. One
should be very careful about water, and we carried a bottle of
Halozone tablets which we usually added to the water. Coun-
try folk are always hospitable and often brought us out cake,
pie and other dainties.
We were sorry to leave the wonderful concrete roads of Ohio,
but we had little to complain of in Indiana. There was very
little collecting from Toledo to BlufiPton. On the banks of the
St. Mary's River at Fort Wayne we found some fine Succinea
avara Say and a few other land shells. At Bluflfton we hunted
up Mr. E. B. Williamson, the dragon fly expert, and a mem-
ber of the museum stafif. We were cordially invited to make
his home our headquarters for as long as we cared to stay in
Bluffton. We were glad to stay two days while we rearranged
our outfit, supplied what we found lacking, and collected in
the vicinity. We found quite a few Flanorbis, Physa, Polygyra,
Succinea along the Wabash and near by. The river itself was
disappointing, because it was in high water. The collection of
dragonflies which Mr. Williamson and his brother have gotten
together is one of the best in the world, and we learned many
pointers from observing their laboratory and field methods.
Such precision, accuracy and care of detail are well worthy of
copy.
After bidding a cordial forewell to Mr. Williamson and his
family, we headed south again. We collected our first lot of
Gnniobasift in the Nolund River, two miles west of Centerville,
Ind. They were probably G. seinicarinatn. Say. After that,
Plenroceridae formed the greater part of our catch. We arrived
at Bloomington on the night of the 25th in a pouring rain. It
being too wet to make camp and cook a meal in any degree of
comfort, we put up at a hotel and after supper took in the local
movie and wrote up our field notes. This latter was always a
THE NAUTILUS. 131
part of Dur evening duties, with a study of maps and a plan for
the next day's trip.
It may be well here to mention how we kept tield notes and
cleaned our catch. As soon as we finished collecting at one
spot, the entire catch was put in a can or hag and a slij) of
paper bearing a number put with it. Under the same number
this entire lot of shells was entered in a field note-book, with
full particulars as to ecologic data, etc. Here is a specimen
entry :
"No. 103. Pleurocera, Cawpeloma, Angitrema and Ancnlosa.
Little Tennessee River, Morganton Ferry, Tenn. Along muddy
banks and on rocks and shoals. 8/22/24."
That, by the way, was one of the finest lots we ever collected.
Our lot numbers ran serially from 1 to 145, the last collected on
Sept. 4. On our next trip we shall start with No. 146. Every
80 often we filled the cans of the last few days' catch with alco-
hol, usually doing it in the evening. Then, next morning, one
of us would stay in camp, empty out the alcohol and dry the
lots separately in the sun, while the other collected. At the
same time full labels were written out and the shells wrapped
up and sent back to the museum as soon as enough for a box
had accumulated. We used denatured alcohol, which, while
not as good as pure alcohol, is cheap and readily obtainable
locally. Later, if it is desired to examine the animal for radula,
it can be soaked out. This end of the collecting was not much
fun, but very necessary. We are setting it down because some
one else may want to adopt it and improve on it. In accounts
of collecting trips the authors usually neglect to state how the
shells were cared for and sent back, and we have often won-
dered what methods were followed. John B. Henderson in his
" Cruise of the Thos. Berrera " is one of the few exceptions.
Our methods of collecting were very simple. When you see
the shells, go in after them. At first we used to pack them up
along the edge, walking out on boards and so forth. But after
a while we got so that we waded right in after them, shoes,
puttees, and all, and our feet and lower limbs were continually
wet from one stream to the next and from one day to the next.
Yet we never once took cold on the whole trip. Back in camp
132 THE NAUTILUS.
we used to put on dry clothes, but pull the wet ones back on in
the morning. If the shells were pretty far out and were plenti-
ful, we put on swimming trunks and old boots and went on out.
We struck no really good collecting until about ten miles
south of Bloomington, Ind., and then every stream was thick
with Fleurocerids, Phyaa, and often naiades. We took ten lots
of Pleuroceridae between Clifty Creek, Columbus, Ind., and the
Ohio River at Louisville. We stopped at Mitchell, Ind. , a day
to inspect the curious cave formations there. In an icy cold
stream issuing from Donaldson Cave, we took a shell which
resembled Goniobasis semicarinata, Say. In the cave itself we
hoped to find blind fish, but saw only one and it got away.
We did collect two small salamanders. From Mitchell south,
the country became very rugged and beautiful, quite a contrast
to the extreme flatness of central Indiana. The good ship
"Asthma", however, never failed us, in spite of the load she
was carrying. In the whole 1700 miles she never gave us any
real trouble, although we did change tires seven times one day !
But that was on the " boulevards " of Kentucky. It was with
real regret that we sold the gallant little car in Knoxville. She
should have been put to pasture like all noble steeds.
We collected a fine series of Goniobasis indianemis Pils. in the
Blue River, taking lots at several points three or four miles
apart. This is the type locality for that species. Here we also
collected three specimens of what looks very much like an
Ancv,losay but will be determined later. The first stage of our
trip ended at Louisville, where we got our accumulated mail,
stocked up on beans again, and pulled out for Cave City, Ken-
tucky.
For the first fifteen miles after leaving Louisville we struck
concrete roads and were beginning to have a high opinion of
Kentucky roads and the Dixie Highway in particular. Then
we hit a detour, then more detour, worse than the last, and
more of it for about sixty miles. When the}' repair a road in
Kentucky they take the whole stretch at once. Words cannot
express the meanness of those roads, and we were so busy
" praising " them that we forgot to take pictures of them. We
passed near Lincoln's birthplace, and now we know why he
THE NAUTILUS. 133
left Kentucky. Furthermore we struck no collecting for about
forty miles after leaving liOuisville. In a way, we are grateful
for those detours, though, because in taking one we were led to
one of the richest collecting spots of the whole trip. As we
were pounding along in the midst of the hills, we decided to
stop a while and cool the engine. Right where we stopped we
noticed a sign which said " Visit the dam and eat at Glenbrook
Hotel," and an arrow pointing down. "Dam" sounded like
water (though it was what we had been saying for many miles),
and water meant shells, so down we went, and we certainly
found shells. The dam and brook below were plastered with a
small black Goniobam and we also took several hundred fine
Phjjsa microstoma Say out of the pond above. They have dam-
med up a big spring there for power purposes. As there were
also indications of land shells, we drove down and camped be-
side the hotel for two days while we collected. In the spring
above and below the dam and in a neighboring spring we took
several quarts of shells, taking care to keep the lots separate.
I have never seen shells more plentiful in one spot except in
First Creek at Knoxville. We also discovered to our surprise
that we were only a hundred yards above Rio on the Green
River, so we hastened to get a boat and collect in the river.
The Green River, and it is green, was one of the places we
wanted to hit, because it has not been collected in above Mam-
moth Cave to any extent. We took half a dozen species of
Pleuroceridae, including Anculosa, and a fine lot of naiades.
Also along the sides of the glen above the spring we found
many land shells, Chivphalma^ Polygyra of many species, and
others. So the detours were not an unmixed blessing; indeed,
we found that to get the good shells, one must get ofif the good
roads. So it was with a sense of time well spent that we left
Rio.
At Cave City we sent our first shipment north, two big boxes,
and then we went out of our way a few miles to see Mammoth
Cave. It turned out that this word should be in the plural, for
we found on arriving that there were half a dozen caves or
more, many said to be more splendid than Old Mammoth.
Acting on advice we had received, we spent a morning explor-
134 THE NAUTILUS.
ing Great Onyx Cave and count it time well spent. Here also
we collected a salamander, mole crickets, and a blind beetle.
It was in one of these caves that Call discovered Carychium
stygium, but we saw none. We also collected again in the
Green River here, and then pulled out and set off for Nashville,
Tenn.
There was not much collecting till we struck the Cumberland
River just above Nashville, and here we took a fine lot of shells,
finding Angitrema for the first time, big, spiny ones that looked
like young lo. This was the only day we really suffered with
heat and we finally stopped collecting to run into town and
cool off at a soda fountain and read our mail. This ended the
second stage of our trip.
From Nashville south to Huntsville, Ala. the collecting be-
came very rich, every stream being loaded. It seems that the
forecasts and reports of high water were not borne out, for in
practically every stream we could wade across and see bottom.
In fact, only once on the whole trip did rain so muddy a stream
that we could not collect in it. It was an ideal summer from a
conch ologist's standpoint, for it rained scarcely once throughout
the belt we collected in for the whole two months.
The Duck River, which w'e crossed at Shelbyville proved a
big disappointment, for we found only one shell, a naiad.
Doubtless this stream is polluted right there. But one mile
south of Shelbyville, in a large tributary of Duck River, we
found very good collecting. Here we took our first Anculosa in
quantity, a nodulated species. They were in what we soon
found to be their characteristic situation — on the edges or
underside of fiat rocks in very swift water, usually at the head
of a riffle.
The Elk River, one mile south of Fayetteville furnished the
best collecting we had so far found. Here from the river itself
and from a small slough close by we took about three gallons
of Pleurocera and Anculosa. We also began to get Campelomn
from now on. A few days before, we took the first Vn'iparn of
the trip along the marshy edge of a river one mile north of
Murfreesboro. Shortly after leaving this locality we crossed
the line into Alabama, and came on into Huntsville. This
THE NAUTILUS. 135
town has been made famous l)y the work of many other col-
lectors, notably H. E. Sargent, H. H. Smith and H. E.
Wheeler. The latter in two excellent papers in the Nautilus
gave a list of the shells of Monte Sano, a high plateau overlook-
ing the town. Before going up Monte Sano we collected in the
Big Spring in the heart of the town. This spring contained a
puzzling species of Goniohnms, puzzling because Sargent had
distributed it as G. •perstriata. Goodrich recognized them as G.
nasi^uld and thought the two names might be synonymous.
The shells we found were G. nassula, but we turned up the true
G. perstriata in Spring Creek. It happened that the day we
visited the spring men were cleaning out water weeds, so we
simply sat down and leisurely picked a box full of the little
fellows. In the afternoon we drove up Monte Sano through
one of the two toll-gates left in Alabama and camped on top.
Next day we devoted to scrambling over the mountain, and
found many of the shells listed by Wheeler, but not all, for it
had been an exceedingly dry summer and land shells seemed to
be very scarce. For the first time in our experience we took
Helicina. This ended the third stage of our journey.
Our next objective was Chattanooga, Tenn. It was a ques-
tion whether to take the road up the north bank of the Ten-
nessee, or the south bank. We finally decided on the latter
because the road was reported much better. So we turned
south till we hit the river at Whitesburg Ferry Landing. Here
we found a great quantity of naiades embedded in new wash of
the river, all clean, in perfect condition, nacre and hinge un-
harmed. We took a large series of each species. A little far-
ther on we crossed the Flint River where we spent several hours
collecting large numbers of Vivipara, Campeloma, Anculoaa, and
several species of Goniohasis and Pleiirocera. The river was
very muddy, but normally so, no doubt, because there was no
trouble getting the shells. We put up for the night at New
Hope and collected in Paint Rock River next day. In the
deepest parts the river was only breast-high, rock bottom with
a layer of silt or gravel, so we put on trunks and covered every
foot of the bottom for a half mile. We are quite sure we got
every species there, and a nice lot it was. As usual there were
136 THE NAUTILUS.
two distinct species of Pleurocera, probably more; one, a squat,
heavy form found on rocks, the other a longer, slender species
found on mud and on logs. We recognized among the naiades
such old friends as Proptera alata Say and Leptodea fragilis Raf.
We crossed the Tennessee at Guntersville, turned west over
Bear Mountain and then headed north. Albertsville, Ala., was
the farthest south we reached. There was very little further
collecting till we reached Chattanooga, and in this respect we
were sorry we did not take the northern road. The only
notable collecting was a good series of a fine, large Pleurocera
from Lookout Creek about four miles north of Valley Head.
On Aug. 13 we rounded Lookout Mountain and entered Chat-
tanooga. This ended the fourth stage of our trip.
At this point Paul J. Adams of Knoxville, Tenn. joined the
expedition as an additional collector and stayed with us till the
end of the trip. His knowledge of the country around Knox-
ville and the Great Smoky Mountains was extremely useful to
us and enabled ue to cover much more ground that we other-
wise would have. On the other hand we were able to show him
wrinkles about the "shell game," which were new to him.
We now began to strike more of the localities which Mr. Good-
rich had suggested as desirable to visit, either because these
were type localities or represented only by puzzling and inade-
quate material. We laid out our course especially to hit such
places. The first of these was Chickamauga Creek at Lee and
Gordon's Mill, just outside the military park. The reason for
visiting this was to check up with material from Crawfish
Springs in the park which was the type locality for Pleurocera
planicostata. We found shells plentiful there and collected
several quarts. We then drove west to reach the Conasauga
River, which is the type locality for eight species of Goniobasis
alone. We were also anxious to collect in this river because it
belongs to the Gulf drainage and for that reason presents a
different fauna from that of the Tennessee. The shells did in-
deed look different from any we had so far seen and we took
large lots from two places on the Conasauga, as well as Mill
Creek, Coahuila River and several other streams. We recog-
nized among these Gonivhtu'iis niurrayensis Lea — this is its terri-
THE NAUTILUS. 137
tory. By accident, while buying postcards in Dalton, CJa., we
saw a picture of "Country Club Lake", which was not on the
map. Arriving there we found a fine large Pleiirocera of the
lake type. Mr. Goodrich is now working at identifying all this
material.
A few miles beyond Dalton we turned north again into Ten-
nessee and went out of our way to visit the Ocoee River dam at
Parksville. This dam has backed up the river for ten miles
and made a large inland lake; we were interested to see what
the shell life would be and we also wanted to collect in the
Ocoee, as no shells have been reported from there recently, if at
all. We found out why; there are no shells in the river. We
learned that farther up the river near Ducktown, sulfuric acid
was produced as a waste in zinc smelting, and run off into the
river. This has completely killed all forms of water life, not
even a fish being found. This affects even the Hiwassee River,
into which the Ocoee flows, so that we found no shells in either.
It was a somewhat new experience, for we had grown so used
to finding shells in every creek that we had acquired the habit
of piling out of the car, grabbing an empty can and wading
right out. At times it almost got monotonous. We climbed a
small mountain. Sugar Loaf Hill, beside the dam and found
quite a few of the smaller land shells at 1100 feet.
From the dam we drove west to Cleveland, Tenn. There
was a Ganioba»is reported many years ago from a "spring near
Cleveland on road to Ducktown " which we wanted to look up,
and we also wanted to investigate a small lake, also called
" Country Club Lake ", shown on the map near the town. We
reached the lake just before dark and pitched camp in a grove
of pines, the most ideal camping spot of the trip. Next morn-
ing we collected thoroughly in the lake and in its inlets and
outlets, and took two fine Goniohasis and a beautifully-ribbed
Pleurocera in great quantity. We found a few naiades also.
Our catch here was particularly interesting as the lake drains
into the Conasauga River. In fact, the ridge just back of the
lake was the dividing line between the Conasauga and the Ten-
nessee drainages. About noon we started a hunt for the elusive
spring and found it without much trouble. There were a few
138 THE NAUTILUS.
of the Goniobasis, which we gathered, but did not linger long,
for the water was icy cold and quite deep, and typhus was re-
ported by the local hospital.
We now headed north for the Tennessee River at Loudon,
which we reached at dusk. Here, at Sam's Island, Dr. C. C.
Adams reported large numbers of /o, form lovdenensh^ and we
were all on fire to diminish the local race. Next morning we
borrowed a boat and went down river. At once we began to
find Sti'ephobaxis, Angitrema, Anculosa, and several Pleurocera
and Ancylus. But no lo ! The Tennessee was still a little
high, however, and we might have had better luck a month
later. Finally, one of us did find the first lo of the trip, a fine,
big loudenensis] and he at least will always remember the thrill
he felt when he put his hand on those big spines. The thrill
never wears off with lo even when we later collected several
hundred in a day. Although we felt the bottom barehanded
for many yards around that spot, a swift rocky shoal, we found
no more. So a little after noon we drove north again to Lenoir
City where our mail awaited us, stocked up on beans again,
and drove northward till we struck the Clinch River at Riley
Ferry, not many miles above the mouth. Here again we took
only a lone lo, but a wonderful catch of naiades, many species,
some of which were old friends. Although we saw ideal situa-
tions for lo, we could find none. Perhaps this is because we
had not yet learned how to look for them, or because until the
water is very low, they are quite dispersed. There is a trick to
lo collecting. Later we found the most plentiful on fairly swift
shoals, at the head of the riflles and at the end, just under the
edges of rocks and smooth ledges, usually spire pointing down
stream. Never in water over three feet deep, sometimes partly
out of water. This is surely the queen of fresh-water shells.
During the next day we made a loop to the north and col-
lected Goniohnsln and Physa in Bear Creek, Poplar Creek, and
several sjjrings, coming out on the Clinch again at Emory
River. We camped here and made extensive collections on the
shoals, again l)agging a lone lo, but many Anculosa, some
knobbed, Amjitreina, StrcjtltolHisi^, etc. The Ancidosa were par-
ticularly large and fine here.
THE NAUTILUS. 139
leaving Kingston, we struck west to run down another
doubtful locality, Cane Creek and Toco Creek near the North
Carolina line. We eventually found both of ihese and neither
had any shells at that point. The rapid deforestation has
changed the character of these creeks and given them a shifting,
sandy bottom unfavorable to shells. Such is liable to be the
fate of many other streams; and many others are being dammed
for power, also changing the shell life. So they should be sur-
veyed as soon as jwssible. In getting to these creeks, we fol-
lowed the Little Tennessee and were amply repaid for our
trouble by the rich collecting we got along it. At Coytee Shoals
we found Angitrema, Physa, Unios, Sphaermm, and a very docol-
lated Pleurocera, while a few miles farther at Morgantown Ferry
we made one of our richest hauls — 700 very large, fine Campe-
loma, probably C. coardatum Lea, over 1000 specimens of a
giant Pletirocera nearly two inches long, and the usual run of
Anculosa, etc. Although we hunted carefully we found no trace
of lo in this river, though conditions seemed ideal and some of
the natives reported finding "spiny shells" on the shoals.
Dr. C. C. Adams and his collectors reported the same lack of
success. It would be interesting to follow this up and see if
there are any in the head waters.
We examined the Tellico River at two places near Vonore and
found a few Pleurocera, then headed west again for Sweetwater.
Sweetwater Creek was one of the localities we wanted to check
up on. It is polluted below the town, but we struck it just
above the town, near the pumping station, and not only took a
large series of Pleurocendxie from the creek itself, but also col-
lected several kinds of Goniobasia from Cannon Spring and a
small pond close by. This was a good haul.
We now continued north to Lenoir City again, collecting in
Turkey Creek en route, and ferried over the Tennessee. We
wanted to run into Cades Cove, and climb one of the peaks of
the Great Smoky Mountains, This would enable us to cut
Little River in many places. We certainly did survey the
latter river thoroughly, for we took eight lots out of the river
from points extending from near the mouth almost to the
source. Our trip into Cades Cove and the hike up Thunder-
140 THE NAUTILUS.
head is one of the high lights of the summer. The road into
the Cove, while in good shape, is surely the crookedest road in
the world, for its length. Coming out we counted 226 turns in
ten miles, many of the hairpin variety. This was Paul J.
Adams' old stamping ground, and we left our machine in the
barn of a friend of his, made up our packs and started on the
hike up. These mountains are the highest east of Rockies, and
though this one was only about 5,500 feet elevation (Thunder
Head), it seemed much more before our tired legs reached the
top. The going was made a little easier by picking up an oc-
casional Polygyra andreivsi Binn. A dozen of these will fill your
pack. Next to P. chilhoiveensis Lewis, also found in these
mountains, this is the queen of the Polygyras. Adams assured
us that in a wetter season, land shells would have been much
more plentiful. We saw far too few. We made the top in a
cold fog, and hastened to seek shelter in the cabin of a cattle
herder, old Tom Sparks, another friend of Adams. Early in
the morning we tumbled out and went up the crown of the
mountain just over the North Carolina line where we got a
superb view of the rugged peaks. Then back for a hasty break-
fast and the long pull down which seemed even harder on the
legs than the hike up. Shortly after noon we got back to the
car, packed up and pulled out. In packing we missed a bag
containing two dozen eggs we had just bought and concluded
the farmer's dogs must have gotten them. About a week later,
Clench, after much complaint from the other two on the con-
dition of his duflfle bag, decided to clean it out and found the
whole two dozen where the farmer had put them to be out of
reach of his dogs! Curtain! We are still laughing at that.
We now turned north to Knoxville, stopping just once to
collect several quarts of big Pleurocera from Nale Creek. We
were in a hurry now because Clench felt an attack of his old
malaria, contracted the previous summer, coming on. When
we reached Adams' home in Knoxville, Clench was just able to
wobble up to bed. Strong doses of quinine brought him around,
however, and two days later he was out collecting again. We
cannot express enough gratitude for the kindness and hospital-
ity which the Rev. Mr. and Mrs. Adams showed us then and
later.
THE NAUTILUS. 141
As soon as Clench was over his attack we made a trip to the
Holston River in company with the veteran collector Manley
1). Barber, a resident of Knoxvillc and Rev. Mr. Adams. Mr.
Barber promised us some lo on a shoals about twelve miles up
the Holston, and was able to make good on his promise. For
the first time we took lo in some quantity, nearly 200, as well
as many other shells. These los were big spiny fellows, possi-
bly typical form spinona. On the way back we collected in
Swan Pond Creek and had very good success. While waiting
for Clench to get well, trips were made to the Tennessee River,
with poor success, and to First Creek with wonderful luck. At
the latter place, although the stream was quite cluttered with
rubbish, the water was literally crowded with Pleurocerids.
Never have we seen shells in such quantity. We filled \x\) all
our cans and buckets in a short time — we could have filled
barrels — and then turned home. It would pay to make a com-
plete survey of that stream, it is so rich, and we hope Adams
will do it. Another bit of collecting we did in Knoxville was
a search for land shells at Cherokee Blufifs and for Hdix nemo-
ralis Linn., introduced near the campus of the University of
Tennessee. About 100 live specimens of the latter were carried
to St. Louis by Remington and "planted" on the grounds of
the Principia School.
On Aug. 30 we left Knoxville on the last leg of the trip.
Our intention was to collect farther up on the Clench and the
Powell and perhaps run into Virginia. We were going after lo
in quantity now. Leaving Knoxville we collected in Second
Creek, Knob Fork Creek and Beaver Creek, and struck the
Clinch at Clinton. Here we collected a big series of Campeloma
and Pleicrocera. A few miles beyond we again hit the Clinch
at the old Moore's Ferry and found the usual run of shells, in
quantity, and about 100 lo, small, of the form brevis. We now
headed for the mouth ef the Powell River where we wanted to
get a big series, as there is some talk of a dam being built there.
We found a few lo here in both the Clinch and Powell, as well
as some fine StrepkobasiSy Anculoaa, naiades, and the river forms
of Pleurocern.
Another stop was made on the Clinch near Maynardsville,
142 THE NAUTILUS.
Teun. where we secured a fine catch of mussels. Then we
made a detour to strike the Powell near Cumberland Gap, and
had very good success there. For the first time we took over
100 lo which were nearly smooth. It would be hard to say
what these shells are, as they intergrade, but they range between
form lyttonensis and form powelle^isk. We kept zigzagging be-
tween the Powell and the Clinch here, until we turned northeast
at Tazewell toward Kyle's Ford on the Clinch. A collection
made in the Little Sycamore Creek was remarkable for having
a few Anculosa in it, as well as some forms of Pleurocerids new
to us. Anculosa were not usually found in small creeks. This
day we made camp just before a pelting rain caught us. We
saw the storm coming and stopped at the first handy spot, but
quickly moved after the storm when we found we were camped
close to a very dead calf ! Once before we found we had
camped near a departed mule. We favor a law in Tennessee
requiring farmers to bury deceased animals.
We came out on the Clinch about six miles below Kyle's
Ford, Tenn. and proceeded to collect all the way up to the
ford. The river here is a series of shoals, quite clear, and we
could see the los on the ledges. At Kyle's Ford the}' were
particularly thick, as many as ten being brought up in one
scoop of the hand. These los are from the type locality for the
form brevis. It was thrilling work, though back-breaking, to
grope for shells and keep a footing in the swift water. But our
long-awaited dream of collecting lo had come true and we
chalked off another aim achieved in conchology. We still have
a few left. This was the hist collecting we did, for our funds
were almost exhausted and our time was about up too. Indeed,
we had stretched the appropriations for two so as to cover ex-
penses for three, and we considered that good managing. On
Sept. 5, we drove back to Knoxville. All that remained now
was to dispose of the car, pack our outfit and take the train
back to Ann Harbor. The Rev. Mr. Adams kindly took the
car off our hands and we were glad to know it was with friends.
We feel very tender toward that car. On Sept. 8 we reached
Ann Arbor again, having stop])ed in Detroit a few hours to re-
port to Dr. Walker and spent a pleasant hour with him.
THK NAUTILUS. 143
Our work, while very scattering, did jipproach completeness
on the Little and Clinch Rivers — we collected at eleven stations
on the latter river. As the upper reaches of the Clinch have
been explored by Goodrich and Ortmann, the Clinch may be
said to be thoroughly explored now. We estimated that we
brought back over 250,000 Bpecimens, Dr. Walker says no
such volume of work has ever been done on the Plevroceridae at
one time. The whole work was really in the nature of a sur-
vey, and when the results are made clear, will furnish a basis
for further work later. We want however, to give the los time
to come out of their hiding places again. It is reported that
the canned-bean industry had a big year! A wonderful trip,
vagabonding for shells!
PUBLICATIONS RECEIVED
Pelecypoda from the Marine Oligocene of Western North
America. By Bruce L. Clark (Univ. Calif. Pub., Bull. Dept.
Geol. Sciences, Vol. 15, No. 4, pp. 69-136, pis. 8-22, 1925).
Forty-four new species are described and figured with notes and
figures of many others.
The fossils of the Lower San Pedro Fauna of the Nob Hill
Cut, San Pedro, California. By T. S. Oldroyd (Proc. U. S.
Nat. Mus., Vol. 65, Art. 22, pp. 1-39, pis. 1-2, 1924).
Twenty-two new species and subspecies are described from this
Pleistocene formation which contains some 242 species of shells.
Discovery of a Balkan Fresh-water Fauna in the Idaho Forma-
tion of Snake River Valley, Idaho. By W. H. Dall (U. S.
Geol. Survey, Prof. Paper 132-G, pp. 109-115, pi. 26, Nov.,
1924). Seven new species, including four of the remarkable
genus Orygocei-as are described. A new genus Payettia is pro-
posed, type Lntia dalli White.
Opisthobranchiate Mollusca. By Frank M. MacFarland
(Proc. Cal. Acad. Sci., 4 Ser., Vol. 13, pp. 389-420, pis. 10-
12, Nov., 1924). One new species Aglajn hakeri and a new
genus Tridachiella, type Tridachid ? diomedea are described and
figured.
144 THE NAUTILUS.
The Boring Mechanism of Teredo. By Robert C. Miller
(Univ. Cal. Publications Zool., Vol. 26, pp. 41-80, pis. 3-6,
1924). An interesting paper on a much discussed subject.
WooD-BORiNG MoLLUSKS FROM THE Hawahan, Samoan and
Philipine Islands. By Robert C. Miller (Univ. Cal. Pub.
Zool., Vol. 26, pp. 145-158, pis. 8-11, 1924). Three new
species — Teredo furcillatm, T. samoaemis, and 7\ trullifermis are
described and figured.
Conditions under which Goniobosis livescens occurs in
THE Island region of Lake Erie. By Frederick H. Krecker
(Ohio Jour. Sci., Vol. 24, pp. 299-310, 1924).
NOTES
On the Type Locality of Polygyra townsendiana brunnea. —
Mr. Walter Eyerdam of Seattle, Washington, writes that he
collected the types of Polygyra townsendiana brunnea Vanatta at
Coal Creek, a small branch of Cowlitz River, which flows into
the Columbia River about ten miles south of Kelso, Cowlitz
County, Washington. It was described in the Proc. Acad. Nat.
Sci. Philada, Vol. 76, 1924, p. 25, as being from along the
Columbia River near Kelso. — E. G. Vanatta.
Professor T. D. A. Cockerell and Mrs. Cockerell expect to
visit Argentina during the summer. Fossil insects will be their
primary object, but mollusks are also to be collected.
The Twenty-fifth Anniversary of the Conchological Section of
the Buffalo Society of Natural Sciences was celebrated on Tues-
day, October 23, 1924. Covers were laid for sixty guests. The
President of the Section, Miss Mary E. Walker, has served in
that office most of the quarter century; the Section having been
founded by Miss Elizabeth J. Letson (Mrs. William Alanson
Brvan). The Rev. George F. Williams acted as toastmaster.
Responses dealing with conchological topics of general interest,
reminiscenses of early days and adventures of the club, poems
in appreciation of shells and other topics appropriate to the
occasion were made by Messrs. Chauncey J. Hamlin, Henry R.
Rowland, John E. Andrews, Dr. W. L. Phillips and Mrs. Burr
H. Nicholls. The chief address of the evening was by Dr.
Bryant Walker, whose subject was "Mollusks and Disease."
The Section has 29 Active Meml)ers, 7 Associates and 5 Hon-
orary Members. Monthly meetings are held from September to
May, with papers on conchological and other natural history
and cognate subjects. Long life to the Buffalo Conchological
Section!
$2.00 per Year. ($2.10 to Foreign Countries.) 50 cts. a copy
THE
NAUTILUS
a quarterly
1)i:\»)Tp:d to the interests
vV^ '^ OF CONCHOLOGISTS
• — \y EDITORS AND PUBLISHERS:
K^
A. PiLSbRV, Special Curator of the Department of Mollusca,
Academy of Natural Sciences, Philadelphia.
C. W. Johnson, Curator of the Boston Society of Natural History.
T-ol. XXX Vni. JULY, 1924. No. 1
CONTENTS:
A Visit to Sweden. By Olof 0. Nylmder 1
Polygyra appressa linguifera and sanctageorgiensis By //. A. I^hhnj
and £. d. Vanalla .... ...... 4
The affinities of Balea africana Melv. and Pons. By I/. A. Pilsbry , . 6
Land shells of Admirals Cave, Bermuda. By E. G. Vanatla ... 6
A remarkable Ciecid from Florida. By W. II. Dall. .... 7
Freshwater Mollusca froni Makenzie River basin, Canada. By E. ./.
miittaker 8
A new species of Physa from Texas. By U. J. Clench . .12
Radula technique for Physa. By W. J. Clench 13
A new Physa from California. By Frank C. Baker . . . .15
Notes on Lacuna from the California Coast. By A. M. Strong . .16
Notes on .Vcanthina from California. By A. M. Strong . . . .18
Marine shells of Drier Bay. Knight Island, Alaska. By Walter Ef/erdam. 21
Some unpublished plates of Conrad's " Unionidie ". By Wm. J. Fox . 28
Dr. Nelson Annandale. Henry Haversham Godwin-Austen. W. H.
Golisch 30
Notes 32
Publications Received 34
C. W. JOHN.SON, Business Manager,
Boston Society of Natural History,
234 Berkeley Street,
Boston, .Mass.
Entered u Secoad-Clase matter at the Uomon Pott-Office.
ii THE NAUTILUS.
A New Work on the Cerionidae
By CHARLES J. MAYNARD
This book is the result of fifty years' study in field aud laboratory, with
200,000 Cerions in hand. Many new species are described and valuable facts
given. Nos. 1 to 9 now issued, 150 pages, 37 hand-colored plates. Catalogue
of Cerions for sale now ready, over 100 species. For Circulars, address the
author
457 Crafts St., "West Newton, Mass.
SHELLS
Wanted : Fine American Land and Fresh Water Shells.
Send list of what you have to e.xchange, or will collect.
Offered : Fine Philippine Land Sliells.
WALTER r. we:bb,
202 'Westminster Road.
ROCHESTER. N. Y.
Otia Conchologica
By A. A. GOULD
Shells of the Wilkes, U. S. Explor. Fxped. ; Shells of the N. I'acific E.xplor.
Exped., and descriptions of shells reprinted from various publications with
rectifications, ] 839-' 62. 256 pages, containing descriptions of 18 new genera
and about 1290 new species. A library in itself. A few copies of this rare
publication are available at tlie unusually low rate of #2.50.
Address, SECRETARY, Boston Society of Natural History
234 Berkeley St., Boston, Mats.
Secondhand Books on Art, History, Science, Geology,
Botany, Conchology, Entomology, Zoology, &c.
Send for Tandy's NfW List.
Address, M. TANDY, Dallas City, Illinois.
EXCHANGE NOTICES
For Exchange: Marine, Land and Fresh-water shells of Cal-
ifornia, for other shells not in my collection,
Wkndell 0. (Jregg, M. D.,
304 N. Boyle Ave., Los Angeles, Calif.
Wanted: Either by exchange or purchase, Mexican and Cen-
tral American shells. J. K. LeB. Tomlin,
23 Boscobel Road, St. Leonard-on-Sea, England.
TUK NAUTILUS. Ill
ITe-w Shell Catalog-ue
We have cumi'U'ted llio iiroparaliou ami publiciitiou of Uio fourth
etlitiou of our complete catalof^uo of MoUusca. This edition contains
one hundred and twenty-four paj^es and two liundred and twenty-three
illustrations. Due to the considerable expense of j)ublication, we are
unable to send the catalogue out free of charge. The price of the cata-
logue bound with heavy paper cover is 55c.; bound in paper and inter-
leaved with blank pages, 70c.; and interleaved and cloth bound, 90c.
We will credit the amount paid for catalogue on first order for shells
amounting to live dollars.
Ward's Natural Science Establishment,
76-104 COLLEGE AVE, ROCHESTER, N. Y.
Florida and West Indian Shells
For Sale and Exchange
Send lists of duplicates and desiderata.
N. "W. LERMOND
P. O. Address: Gulfport, Fla., until May 1
After that date, Thomaston, Me.
FOR SALE
Back Volumes of The Nautilus
Vols. 4 to 20, $1.00 eacli; Vol. 21, S3. 00; Vols. 22 to 27,
$1.00 each; Vols. 28 to 30, $1.50 each; Vols. 31 to 30, $2.00
each. Vol. 21 will be sold only with a complete set. We can-
not furnish Vols. 1 and 2 ( " Conchologist Exchange") and
Vol. 3 of The N.autilus.
Also for Sale a Number of Interesting Works on Conchology.
Address:
THE NAUTILUS
234 Berkeley Street,
Boston, Mass.
IV THE NAUTILUS.
THE MANUAL OF CONCHOLOGY.
The volume now in progress contains a monograph of the
Pupillidae, Pupoides, Pupilla, etc.
Issued. quarterly in parts of which four form a volume.
Plain Edition, per part, $3.00. Colored Edition. $5.00.
S. RAYMOND ROBERTS, Treasurer,
p. 0. Address, GLEN RIDGE. NEW JERSEY, U. S. A.
SOWBRBT & FULTON
(Proprietor, HUGH C. FULTON)
Tbe Largest and Finest Stock of Stiells in the Wcrld
Specimens Sent on Approval
SALE PURCHASE EXCHANGE
Collections and Specimens
Named and Arranged
27 Shaftesbury Road
Ravenscourt Park, London "W. 6. England
$2.00 per Year. ($2.t0 to Foreign Countries.) 50 cts. a copy
THE
NAUTILUS
A QUARTERLY
DEVOTED TO THE INTERESTS
OF CONCHOLOGISTS
EDITORS AND PUBLISHERS:
H. A. PiLSBRY, Special Curator of the Department of Mollusca,
Academy of Natural Sciences, Philadelphia.
C. W. Johnson, Curator of the Boston Society of Natural History.
Vol. xxxvnr. ootober, 1924. No. 2
CONTENTS :
On the Rio Grande. By James II. Femas ....... 37
Some old Pleurocerids and a new one. By Calvin Goodrich . . .43
Marine Mollusca of the Bridgeport, Connecticut, Region. By Arthur Jacot 49
Two new Naiades from Nicaragua. By Bryant Walker . . . .51
California Helices. By H. A. PiUbry ....... 54
Collecting in Southern Florida, the Bahamas and Cuba. By D. L. Emery 56
Eulota Maackii, a Siberian Snail. By T. D. A. Cockerell . . . .62
Notes ............. 65
Publications Received 66
C. W. JOHNSON, Business Manager,
Boston Society of Natural History,
234 Berkeley Street,
Boston, .Mass.
tCnlered m Hecond-OlaiK matter at the Boaton Post-oaicc.
11 THE NAUTILUS.
A New Work on the Cerionidae
By CHARLES J. MAYNARD
This book is the result of fifty j-ears' study in field and laboratory, with
200,000 Cerions in hand. Many new species are described and valuable facts
given. Nos. 1 to 9 now issued, 150 pages, 37 hand-colored plates. Catalogue
of Cerions for sale now ready, over 100 species. For Circulars, address the
author
457 Crafts St., West Newton, Mass.
SHELLS
"Wanted » Fine American Land and Fresh Water Shells.
Send list of what you have to exchange, or will collect.
Offered » Fine Philippine Land Shells,
WALTER r. WEBB,
202 ^Westminster Road,
ROCHESTER, N. Y.
Otia Conchologica
By A. A. GOULD
Shells of the Wilkes, U. S. Explor. Kxped. ; Shells of the N. Pacific Explor.
Exped., and deKcriptions of shells reprinted from various publications with
rectifications, 1889-'62. 2-56 pages, containing descriptions of 18 new genera
and about 1290 new species. A library in itself. A few copies of this rare
publication are available at the unu.sually low rate of f2.50.
Address, SECRETARY, Boston Society of Natural History
234 Berkeley St., Boston, Mais.
EXCHANGE NOTICES
For Exchange: Marine, Land and Fresh-water shells of Cal-
ifornia, for other shells not in my collection.
Wendell 0. Gregg, M. D.,
816 Montrose Ave., Montrose, Calif.
Wanted: Unionidae and Mutelidae, not in collection. Send
list. J. R. LeB. Tomlin,
23 Boscobel Road, St. Leonard -on -Sea, England.
For Exchange: Southern California marine shells, for other
shells not in my collection. Morris E. Caruthers,
120 E. Chestnut St., Glendale, Calif.
TlIK NAUTILUS. Ill
JfTew Sliell Catalogr^e
We have ci'iupleied the prcpiualiou unci publiouUou of tlie fouitb
editiuu uf our complete catalogue of MoUusca. This edition contaiuB
>ue hundred and twenty-four pages and two hundred and twenty-three
illustrations. Due to the considerable expense of publication, we are
unable to seud the catalogue out free of charge. The i>rice of the cata>
loyue bound with heavy i>aper cover is 55c. ; bound in paper and inter-
leaved with blank pages, 70c.; and interleaved and cloth bound, 90c.
We will credit the amount paid for catalogue on first order for shells
amounting to five dollars.
Ward's Natural Science Establishment,
76-104 COLLEGE AVE. ROCHESTER, N. Y.
Florida and West Indian Shells
For Sale and Exchange
Send lists of duplicates and desiderata.
N. W. LERMOND
P. O. Address: Gulfport, Fla., until May 1
After that date, Thomaston, Me.
FOR SALE
Back Volumes of The Nautilus
Vols. 4 to 20, $1.00 each; Vol. 21, $3.00; Vols. 22 to 27,
$1.00 each; Vols. 28 to 30, 81.50 each; Vols. 31 to 36, $2.00
each. Vol. 21 will be sold only with a complete set. We can-
not furnish Vols. 1 and 2 ( " Conchologist Exchange") and
Vol. 3 of The Nautilus.
Also for Sale a Number of Interesting Works on Conchology.
Address:
THE NAUTILUS
234 Berkeley Street,
Boston, Maes.
IV THE NAUTILUS.
THE MANUAL OF CONCHOLOGY.
The volume now in progress contains a monograph of the
Pupillidae, Pupoides, Pupilla, etc.
Issued I quarterly in parts" of which four form a volume.
Plain Edition, per part, $3.00. Colored Edition. $5.00.
S. RAYMOND ROBERTS, Treasurer,
p. 0. Address, GLEN RIDQE, NEW JERSEY, U. S. A.
SOWBRBY & PULTON
(Proprietor, HUGH C. FULTON)
The Largest and Finest Stock of Shells in the World
Specimens Sent on Approval
SALE PURCHASE EXCHANGE
Collections and Specimens
Named and Arranged
27 Shaftesbury Road
Ravensoourt Park, London "W. 6. England
$2.00 per Year. ($2.'0 to Foreign Countries.) 50 cts. a copy
THE
NAUTILUS
A QUARTERLY
DEVOTED TO THE INTERESTS
OF COXCHOLOGISTS
EDITORS AND PUBLISHERS:
H. A. PiL^BRY, Special Curator of the Department of Moliusca,
Academy of Natural Sciences, Philadelphia.
C. W. Johnson, Curator of the Boston Society of Natural History.
Vol. XXXVIII. JANUARY, 1925. No. 3
CONTENTS :
The family Lancida? distinguished from the Ancylidiv. By //. A. Pilsbry 73
A visit to the Hawaiian Islands. By T. D. A. Cockerell . . . .76
The type of Ancvlastrum is Ancylus fluviatilis Miiller. By ^4. S. Kennard
and B. B. W'oodward 83
On the Heli.\ perspectiva of Meperle v. Miihlfeld vs. that of Say. By A. S.
Kennard and B. B. Woodivard ........ 85
Agnathoraorphous Aulacopoda. By //. Burrington Bakn . . .80
An abnormal clam. By ./. Henry Blake ....... 89
Gonchologista in the making. By Edwin E. Iland . . . . .90
Newfoundland Shells. By E. O'. Vanalln 92
Additions to the list of Marine Shells of Sanibcl, Florida. By William J.
Clench 93
New Shells from Japan. By W. TT. Ball 95
Notes on the .Mactras of the West Coast. By A. M. Stron;/ .98
Notes lOH
Publications Received 106
C. \V. JOHNSON, Business Manager,
Boston Society of Natural History,
234 Berkeley Street,
Boston, Mass.
Kntered as Becoad-Clasfi matter at the Boiton Poit-Office.
u- ^
11 THE NAUTILUS.
A New Work on the Cerionidae
By CHARLES J. MAYNARD
This book is the result of fifty years' study in field and laboratory, with
200,000 Cerions in hand. Many new species are described and valuable facts
given. Nos. 1 to 9 now issued, 150 pages, 37 hand-colored plates. Catalojiue
of Cerions for sale now ready, over 100 species. For Circulars, address the
author
457 Crafts St., West Newton, Mass.
SHELLS
'Wanted : Fine American Land and Fresh ^V^ater Shells.
Send list of what you have to e.xchange, or will collect.
Offered : Fine Philippine Land Shells.
WALTER r. WilBB,
202 "Westminster Road,
ROCHESTER, N. Y.
EXCHANGE NOTICES
For Exchange: Marine, Land and Fresh-water shells of Cal-
ifornia, for other shells not in my collection.
Wendell 0. Gregg, M. D.,
815 Montrose Ave., Montrose, Calif.
Wanted: Unionidae and Mutelidae, not in collection. Send
list. J. R. LeB. Tomlin,
23 Boscobel Road, St. Leonard-on-Sea, England.
For Exchange: Southern California marine shells, for other
shells not in my collection. Morris E. Caruthers,
120 E. Chestnut St., Glendale, Calif.
For Exchange: West American shells and others. Will also
exchange shells for stamps desired in his collection or stamps
for shells.
Wanted: Copy of Keep's West American Shells.
W. L. Brown, Route No. 1, Box No. 140, Anaheim, Calif.
TlIK NAI'TII.L'S. HI
XTevT Sh.ell Catalog-ae
Wo liiivo coiuplctetl the iiri'imratioii and iiuhlifalinn of the foiutl)
edition of our complete catalo)i;ue of Mollusca. This edition coutalDB
one hundred and twenty-four pa^es and two hundred and twenty-three
illustrations. Due to the considerable expense of publication, we are
unable to send the catalogue out free of charge. The price of the cata-
io^;ue bound with heavy pai>er cover is 55c. ; bound in paper and inter-
leaved with blank pages, 70c.; and interleaved and cloth bound, 90c.
We will credit the amount paid for catalogue on first order for shells
amounting to five dolUvrs.
Ward's Natural Science Establishment,
76-104 COLLEGE AVE, ROCHESTER, N. Y.
Otia Conchologica
By A. A. GOULD
Shells of the Wilkes, U. S. Explor. Kxped. ; Shells of the X. Pacific Explor.
Exped.. and descriptions of shells reprinted from various publications with
rectifications, 1889-'G-. 256 pages, containing descrij)tions of 18 new genera
and about 1290 new species. A library in itself. A few copies of this rare
publication are available at the unusually low rate of ;f 2. 50.
Address, SECRETARY, Boston Society of Natural History
234 Berkeley St., Boston, Mats.
FOR SALE
Back Volumes of The Nautilus
Vols. 4 to 20, $1.00 each; Vol. 21, S3.00; Vols. 22 to 27,
$1.00 each; Vols. 28 to 30, $1.50 each; Vols. 31 to 30, 82.00
each. Vol. 21 will be sold only with a complete set. We can-
not furnish Vols. 1 and 2 (" Conchologist Exchange") and
Vol. 3 of The Nautilus.
Also for Sale a Number of Interesting Works on Conchology.
Address:
THE NAUTILUS
234 Berkeley Street,
Boston, Mass.
IV THE NAUTILUS.
THE 3IANUAL OF COXCHOLOGY.
The volume now in progress contains a monograph of the
PnpilHdae, Pupoides, Pupilla, etc.
Issued quarterly in parts of which four form a volume.
Plain Edition, per part, $3.00. Colored Edition. $5.00.
S. RAYMOND ROBERTS, Treasurer,
p. 0. Address. GLEN RIDGE. NEW JERSEY, U. S. A.
The Largest and Finest Stock of Shells in the World
We have just published a new edition of our MARINE AMD FRESH-
"WATER GASTROPODA CATALOGUE, consisting of 71 pages
and a full index. This issue has been made still more useful as a check-list and
for reference by the addition of the families and also of many additional generic
names. It has the usual wide margins fur adding species not noted therein.
Price, $1.00, post free
In Hand.— A New Catalogue of the Land Shells
Spechnetis Identified. Exchnnge Invited
HUGH C. FULTON
(SOWERBY & FULTON)
27 Shaftesbury Road, London, W. 6, England
rpHE undersigned desires assistance in obtaining lots of Pisid-
ium from all parts of N. America ; also alcoholic specimens
of Succinea, Vertigo, Lymnaea (Radix) and Planorbis (Gyraidus).
HANS SCHLESCH
14 Qu3tav Adolfsgade, Copenhagen, Denmark
$2.00 per Year. ($2.10 to Foreign Countries.) 50 ct«. a copy
THE
NAUTILUS
A QUARTERLY
DEVOTED TO THE INTERESTS
OF CONCHOLOGISTS
EDITORS AND PUBLISHERS:
H. A. PiLSBRT, Special Curator of the Department of MoUusca,
Academy of Natural Sciences, Philadelphia.
C. W. Johnson, Curator of the Boston Society of Natural History.
Vol. XXXVni. APRIL, 1925. No. 4
CONTENTS :
On a Tanganyika Beach. By Charles Hedley . . . . . .109
Notes on the nomenclature of our East American species of Pecten. By
W. II. Dall 112
Note on the name of our common oyster. By W. II. Dall . . .121
Identity of Patella tenuilirata Carpenter. By J. R. LeB, Tomlin . 121
Some land shells from the Aleutian Islands. By G. Dallas Ifanna . .122
Lymniea auricularia (L. ) in California. By O. Dallas Ilanna . . . 125
Vagabonding for Shells. By /*. 5, Remington, Jr., and H'. J. CUnch . 127
Publications Received .......... 143
Notes 144
C. W. JOHNSON, Business Manager,
Boston Society of Natural History,
234 Berkeley Street,
Boston, Mass.
Eatered MSecoad-Cl&ss matter atthe Boston Po«t-Office.
11 THE NAUTILUS.
A New Work on the Cerionidae
By CHARLES J. MAYNARD
This book is the result of fifty years' study in field and laboratory, with
200,000 Cerions in hand. Many new species are described and valuable facts
given, Nos. 1 to 9 now issued, 150 pages, 37 hand-colored plates. Catalogue
of Cerions for sale now ready, over 100 species. For Circulars, address the
author
457 Crafts St., "West Newton, Mass.
SHELLS
Wanted s Fine American Land and Fresh'Water Shells.
Send list of what you have to exchange, or will collect.
Offered : Fine Philippine Land Shells.
WALTER r. WEBB,
202 "Westminster Road.
ROCHESTER. N. Y.
EXCHANGE NOTICES
For Exchange: Marine, Land and Fresh- water shells of Cal-
ifornia, for other shells not in my collection.
Wendell O. Gregg, M. D.,
815 Montrose Ave., Montrose, Calif.
Wanted: Unionidae and Mutelidae, not in collection. Send
list. J. R. LeB. Tomlin,
23 Boscobel Road, St. Leonard-on-Sea, England.
For Exchange: Southern California marine shells, for other
shells not in my collection. Morris E, Caruthers,
120 E. Chestnut St., Glendale, CaHf.
For Exchange: West American shells and others. Will also
exchange shells for stamps desired in his collection or stamps
for shells.
Wanted: Copy of Keep's West American Shells.
W. L. Brown, Route No. 1, Box No. 140, Anaheim, Calif.
THK NAUTILUS. HI
ZTevr Sliell Catalogue
We have completed the preparation and publication of the fourth
edition of our complete catalogue of MolluKca. This edition contains
one hundred and twenty-four jjagcs and two hundred and twenty-three
illustrations. Due to the considerable expense of publication, we are
unable to send the catalogue out free of charge. The price of the cata-
logue bound with heavy paper cover is 55c.; bound in paper and inter-
leaved with blank pages, 70c.; and interleaved and cloth bound, 90c.
We will credit the amount paid for catalogue on first order for shells
amounting to five dollars.
Ward's Natural Science Establishment,
76-104 COLLEGE AVE, ROCHESTER, N. Y.
Otia Conchologica
By A. A. GOULD
Shells of the Wilkes, U. S. Explor. Exped. ; iShellsof the N. Pacific Explor.
Exped. , and descriptions of shells reprinted from various publications with
rectifications, 1839-'62. 256 pages, containing descriptions of 18 new genera
and about 1290 new species. A library in itself. A few copies of this rare
publication are available at the unusually low rate of f2.50.
Address, SECRETARY, Boston Society of Natural History
234 Berkeley St., Boatoo, Matt.
FOR SALE
Back Volumes of The Nautilus
Vols. 4 to 20, $1.00 each; Vol. 21, 83.00; Vols. 22 to 27,
$1.00 each; Vols. 28 to 30, SI. 50 each; Vols. 31 to 3t), 82.00
each. Vol. 21 will be sold only with a complete set. We can-
not furnish Vols. 1 and 2 ( " Conchologist Exchange") and
Vol. 3 of The Nautilus.
Address :
THE NAUTILUS
234 Berkeley Street,
Boston, Mass.
IV THE XACTILUS.
THE MANUAL OF COXCHOLOOY.
The volume now in progress contains a monograph of the
Pupillidae. Pupoides. Pupilla. etc.
Issued quarterly in parts of which four form a volume.
Plain Edition, per part $3.00. Colored Edition. $5.00.
S. RAYMOND EOBEETS, Treasurer,
p. 0. Address, GLEN BIDGE. NEW JEBSEY, U. 8. A.
Ibe Largest ana Finest Stock of Stiells In the World
We have iust published a new edition of our MARINE AMD FRESH-
"WATER GASTROPODA CATALOGUE, consisting of 71 pages
and a fuli index. This is5ue has been made sull more useful as a check-list and
for reference by the addition of the families and also of many additional generic
names- It has the usual wide margins for adding species not noted therein.
Price, $1,00, post free
In Hand.— -A New Cataiogue of the Land Shells
Speritnett.< Identified. Exrhange Invited
HUGH C. FULTON
SOWERBY & FULTON;
27 Shaftesbury Road, London, W. 6, England
rPHE undersigned desires assistance in obtaining lots of PiMd-
■^ iuia from all parts of X. America ; also alcoholic specimens
of Siuxinea, Vertigo, Lymnaea (Radix) and Planorbi^ (Gyraulus).
HANS SCHLESCH
14 Gustav Adolfs^ade, Copenhagren, Denmark
Mill N^ Ml >l I IHH Vl(>
UH 17XS ^