580,5

FB

V, 20:5-7

V, 22:6-10

1942

cop, 2

'. -.V*

D

13.

NEW SPECIES OF CROTON

FROM GUATEMALA

BY

LEON CROIZAT

TECHNICAL ASSISTANT

ARNOLD ARBORETUM OF HARVARD UNIVERSITY

BOTANICAL SERIES

FIELD MUSEUM OF NATURAL HISTORY

VOLUME 22, NUMBER 8

JUNE 15, 1942

PUBLICATION 510

NEW SPECIES OF CROTON

FROM GUATEMALA

BY

LEON CROIZAT

BOTANICAL SERIES

FIELD MUSEUM OF NATURAL HISTORY

VOLUME 22, NUMBER 8

JUNE 15, 1942

PUBLICATION 516

NEW SPECIES OF CROTON FROM GUATEMALA

LEON CROIZAT

The species of Croton here discussed or described as new were

collected by Paul C. Standley during two Field Museum botanical

expeditions to Guatemala in 1938-39 and 1940-41.

Croton botryocarpus, sp. nov. — Frutex gracilis ca. 2-metralis.

Innovationibus inflorescentiisque sordide hispidulo-argillaceis vel

hispido-tomentosis demum glabratis. Foliis ovato-ellipticis vel

ovato-lanceolatis, breviter acuminatis vel cuspidatis, basi rotundatis,

margine pustuloso-asperatis pilisque simplicibus auctis, subtus tri-

chomatibus late stellatis lanulosis, supra novellis aeque ac adultis

glabris, in sicco brunneis, 6-3 cm. longis, 2.5-1.5 cm. latis, venis

primariis pinnatim ca. 8-jugis adscendentibus; petiolo lanuloso-

stellato, 1-0.5 cm. longo, glandulis 2 subcylindricis, stipulis minimis

vel subnullis. Inflorescentiis simplicibus, vulgo foliis verticillatim

dispositis fultis, basi conferte 9 , ad 5-6 cm. longis. Floribus cf1

ignotis. Floribus 9 : perianthio (sub fructu) ca. 3-3.5 mm. lato,

lobis 5 discretis, glabratis integerrimis linearibus, ad 1.5-2 mm.

longis, petalis videntur nullis; capsula ellipsoidea, apice gibbose

3-cocca, ca. 6 mm. longa, 5-5.5 mm. lata, indumento detergibili

matura glabrata, stylis 3 persistentibus ad 2-2.5 mm. longis, quove

fere ad basim partito, columella fructu delapso ca. 4.5 mm. longa,

apice incrassata, semine ellipsoideo 4 mm. longo, 3-3.5 mm. lato,

apice truncate, caruncula minima, testa brunnea obscure pinnatim

costulata, arillo tenuissimo subnitido. — Guatemala, Dept. Jalapa,

vicinity of Jalapa, alt. about 1,360 meters, "damp thicket, slender

shrub 2 meters tall," Paul C. Standley 77519 (type in Herb. Arnold

Arb.; duplicate in Herb. Field Mus.); Dept. Huehuetenango, north-

west of Malacatancito at km. 8 of the highway from Huehuetenango,

alt. about 1,950 meters, "dry open rocky slope; shrub 2 meters tall,"

Standley 82206, Jan. 1941.

Standley 82206 is a poor specimen, as I have it, but probably

belongs here. This new species suggests C. decalobus Muell. Arg.,

but can easily be separated from it on account of the entire perianth

lobes of the 9 flower. Croton lasiopetaloides Croiz., described in

this same paper, has a different kind of foliage and a somewhat

different 9 flower. The congested capsules, the lanulose-stellate

445

446 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

indumentum, the venation, and the peculiar margin of the blade

are characteristic.

Croton callistanthus Croizat in Journ. Arnold Arb. 21: 84.

1940. — Standley 86784, collected along the old road between Finca

Pirineos and Patzulin, alt. 1,200-1,400 meters, Dept. Quezaltenango,

Guatemala, belongs to this species, and is in fruit. Woodson, Allen

& Seibert 1006, collected at Bajo Mona, Prov. Chiriqui, Panama,

also is in fruit, and despite the original erroneous determination,

C. gossypiifolius, certainly belongs to C. panamensis Muell.-Arg.

The fruiting parts of these two plants differ as follows:

Croton callistanthus. Capsule small, the dehisced cocci delicate,

not over 4.5 mm. long on the average, the yellowish epicarp finely

granular and scarcely tomentose; columella at dehiscence about

3 mm. long; seed scaraboid, about 3 mm. long and broad (at the

small caruncle), the testa irregularly granular-rugose.

Croton panamensis. Capsule fairly large for a species of this

group, the dehisced cocci up to 6.5 mm. long, the epicarp manifestly

verruculose and brown-orangish-tomentose; columella about 5.5 mm.

long; seed (not as yet perfectly ripe but apparently measuring full

size) scaraboid, about 5 mm. long and 3.5 mm. broad, the testa

gross-costulate.

Croton callistanthus and C. panamensis are so similar in the

characters of their vegetative parts and cf flowers that they can not

be certainly identified as distinct unless ripe capsules and seeds are

available. It is characteristic of Croton, especially in regions that

have enjoyed the same climate from ages past, that large groups of

species retain the same foliage and the same cT flowers, while speciat-

ing in the 9 perianth and in the capsule, a fact emphasized else-

where in this paper under C. guatemalensis. Thus, it proves practi-

cally impossible to dispose of critical forms unless very good collec-

tions are forthcoming, the risk attending the publication of new

species on anything less than perfect material being very great. It

is not possible as yet to decide whether C. Steyermarkianus Croiz.

is nearer to C. callistanthus than to C. panamensis. On the other

hand, C. triumfettoides Croiz. might be represented by Austin Smith

A407, collected in the region of Zarcero, Costa Rica, a specimen on

which appear the hispid pubescence and the slender stipitate glands

of the type.

A substerile specimen, Standley 88211, Dept. Retalhuleu, region

of Ajaxa, east of Santa Cruz Mulua, Guatemala, belongs apparently

to C. callistanthus.

NEW SPECIES OF CROTON 447

Croton guatemalensis Lotsy in Bot. Gaz. 20: 353, pi. 25, 1-6.

1895. — Croton is a genus which speciates by modification of the 9

flower and fruit rather than in other manners. In Malaysia not

less than in Central America the species of the same affinity are

apt to retain the same foliage, and can not be recognized as distinct

unless the fruit is seen. It is often found that plants that appear

to be identical at first differ very materially in the size and indument

of their capsule.

In Malaysia and Australasia a great many different forms center

around C. argyratus Bl. and C. Verreauxii Baill., up to six or seven

manifestly different forms having been identified for years under

Blume's or Baillon's binomial. In the same manner, several are the

forms in the vicinity of C. niveus Jacq. which range from Venezuela

to Mexico and are confused today as one species. Jacquin's

binomial, as now used, is practically meaningless.

Lotsy's descriptions of C. guatemalensis and C. eluterioides are

such that only one species seems to be involved. The locality of

collection is approximately the same if not the same (Dept. Santa

Rosa, Guatemala), and little in the diagnoses seems to justify the

belief that the species are two rather than one. Notwithstanding

this, it is quite probable that C. eluterioides and C. guatemalensis

are distinct. Lotsy states that the capsule of C. guatemalensis is

unknown, and that the largest fruiting specimen of C. eluterioides

which he saw had a capsule about 21 mm. long and 15 mm. wide.

Heyde & Lux 3035, isotype of C. guatemalensis, and Heyde & Lux

3470, isotype of C. eluterioides, are available here. I find no 9

flowers on the former, but nearly ripe capsules about 13 mm. long

and about 10 mm. wide on the latter. The vegetative parts are

practically identical in the two specimens cited.

A very important character which Lotsy does not mention is the

armature of the capsule of C. eluterioides. The epicarp in this fruit

is coarsely granular if not bluntly muricate, very rough to the

touch and easily seen without a lens. So far, I have seen no other

specimen that I can match with Heyde & Lux 3470, this indicating

that C. eluterioides is apparently local or rare. I have found, on

the contrary, numerous specimens which well agree with Lotsy's

illustration of C. guatemalensis and have a much smaller capsule

than the isotype of C. eluterioides. This capsule is roughened some-

what, but its processes are smaller and more loosely arranged than

on the fruit of C. eluterioides; the size is also smaller, the ripe capsule

448 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

measuring on the specimens in question about 9 mm. in length and

about 8 mm. in thickness, the seed being 7 mm. long and 5 mm. broad.

Thus, two different species seem to exist in the flora of the

Department of Santa Rosa, one with a coarse and large capsule,

the other with a sparingly granular and smaller fruit. The former

is C. eluterioides, the latter I accept here as C. guatemalensis. These

two species can not be told apart unless in fruit. Neither one is

C. niveus Jacq., despite their having been so identified in several

cases. The classic locality of C. niveus is "Carthagenae in sylvaticis"

(Colombia), and I believe that this species is well represented by

such specimens as H. H. Smith 356, Colombia, Santa Marta; and

Brother Elias 327, Colombia, Barranquilla and vicinity. Croton

niveus would seem to be an altogether smaller plant than C. guate-

malensis, with lesser cf flowers, and is probably not found north of

the eastern coast of Nicaragua. I have never seen a specimen of it

in fruit, which is puzzling.

I determine as C. guatemalensis the following specimens, all from

Guatemala, subject to a later revision with material in full fruit

and larger collections: Bequaert 25, near Pacayal, in 1931; Skutch

894, Dept. Quezaltenango, in 1934; Standley 58086, Dept. Sacatepe"-

quez, in 1938; Standley 59582, Dept. Guatemala, in 1938; Standley

62318, Dept. Chimaltenango, in 1939; Standley 86715, Dept. Quezal-

tenango, in 1941; Standley 88108, Dept. Retalhuleu, in 1941; Standley

88174, Dept. Retalhuleu, in 1941; Standley 88918, Dept. Sacatepe*-

quez, in 1941; Standley 89546, Dept. Escuintla, in 1941. Since ripe

or nearly ripe capsules are available on Bequaert 25, Skutch 894,

Standley 62318, 86715, 88918, and 89546, my concept of C. guate-

malensis rests in the main upon these collections.

This manuscript was in type when I received three specimens

which very closely match Heyde & Lux 3470, the isotype of C.

eluterioides Lotsy available here, namely: Dodge & Goerger s.n.,

Costa Rica: Prov. Cartago, slope of Cerro Carpintera above La

Union de Tres Rios, 1,350-1,500 meters, July 1936; P. H. Allen 661,

Costa Rica: Vicinity of El Alto Station on road to Cartago, 1,550

meters, "tree 20 m., flowers white," December 1937-January 1938;

Davidson 865, Panama: Prov. Chiriqui, Boquete, 1,200 meters,

"large tree," July 1938. The Davidson collection bears a fruit

which has dehisced cocci about 19 mm. long with a much roughened

epicarp, and ellipsoid seeds, pointed at the chalazal end, strongly

keeled on the back, very shallowly costulate, about 19 mm. long and

9 mm. broad. The caruncle is remarkably small and the aril brown-

NEW SPECIES OF CROTON 449

ish, with darker mottlings nearly parallel with the conspicuous raphe.

On Allen 661 is found a 9 flower perfectly comparable with flowers

of the same sex on Standley 86715. The styles of the Allen plant are

manifestly longer and much more intricate (up to 5 mm. long) than

those of the Standley collection (not over 2.5 mm. long) ; the pedicel

is also shorter (1 cm. long) and somewhat stouter in the former than

in the latter (ca. 2 cm. long). Barring these differences, the two

flowers can not be told apart with certainty.

On the strength of the cited material I now believe that C.

eluterioides is a more southern plant than C. guatemalensis, ranging

mostly in Panama and Costa Rica and reaching in Guatemala its

northern distributional limits. All the specimens listed have been

distributed as C. niveus Jacq.

Croton jalapensis, sp. nov. — Frutex vel arbuscula ad 6 m.

alta. Innovationibus inflorescentiisque pube stellato-hispidula

citrina indutis, citius glabratis. Foliis pro more ovatis haud longe

acuminatis, margine grosse dentato-serratis caeterum ambitu figura-

que ludentibus, interdum elliptico-lanceolatis vel ovato-lanceolatis

subcuspidatis, basi rotundatis vel rotundato-truncatis, 10-3 cm.

longis, 6-2.5 cm. latis, firme chartaceis Celtidis spp. texturam in

mentem vocantibus, novellis subtus griseo-tomentellis venis pallide

lutescentibus, supra laxius tomentosis; adultis subtus valde glabratis,

supra indumento longe radiato sparsissimo papillisque scabridis,

in sicco olivaceis; venis primariis ad 5-jugis, primo jugo tertium

laminae superum attingente valde ramoso, margine inter dentes

glandulis stipitatis sub lente obviis ornato; petiolo luteo-tomentello

2-2.5 cm. longo, apice glandulis 2 anticis vel subanticis stipitatis

ornato. Floribus c? ignotis. Floribus 9 suppeditantibus valde

mancis: pedicello brevissimo, 1.5 mm. .longo vel breviore, eadem

ratione ac ovario induto; ovario globuloso tricocco, ca. 2 mm. magno,

indumento citrino conferto toto occultato, columella fructu delapso

ca. 5 mm. longa, coccis maturis vix tomentellis ca. 9 mm. longis,

duris; semine scaraboideo haud applanato, 5.5 mm. longo, ca. 5

mm. lato, apice more proprio producto-carinato, caruncula estipitata

1 mm. longa, 1.5 mm. lata, testa grosse at haud profunde rugoso-

costata, arillo subnitido tenui, brunneo-maculato. — Guatemala,

Dept. Jalapa, vicinity of Jalapa, alt. about 1,360 meters, "damp

thicket; shrub 2-3 meters tall," Paul C. Standley 76414, Nov. 1940

(type in Herb. Arnold Arb.; duplicate in Herb. Field Mus.); same

locality and altitude, "damp thicket. Shrub or small tree to 6 meters,

scarce," Standley 77378, Nov. 1940.

450 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

Strongly suggesting C. quercetorum Croiz., but differing from it

in the seed, the capsule, the glands, the margin of the leaf, and the

indument, the last especially on young leaves. Its affinities suggest

both C. corylifolius Lam., of the West Indies, and C. malvaviscifolius

Millsp., of Yucatan.

Croton jutiapensis, sp. nov. — Frutex 2-metralis. Innovationi-

bus inflorescentiisque pube hispida pallide ochracea tomentosis,

citius glabratis. Foliis ovato-lanceolatis vel lanceolatis, apice

breviter cuspidatis, basi rotundato-cuneatis, 6-3 cm. longis, 3.5-1.5

cm. latis, supra brunneo-olivaceis, subtus brunneo-griseis, primum

utrinque tomentosis, adultis supra quam subtus laxius indutis,

margine manifeste duplicato-serratis vel dentato-serratis (dentibus

ad 3 per 1 cm. marginis longitudine), inter crenas sub lente stipitato-

glandulosis, venis primariis utrinque ca. 6-jugis, adscendentibus,

bene anastomosatis, penninerviis; petiolo laete ochraceo-tomentoso,

0.5-1.5 cm. longo, glandulis stipitatis 2 lateralibus, stipulis setaceis

integris ad 0.5 mm. longis. Flore 9 unico viso: perianthii lobis

anisomeris, more Julocrotonis spp., primo intuito insigni, ad 12 mm.

longo, totidem lato, lobis ovarium valde excedentibus 5, majoribus

spathulatis, ad 8 mm. longis, 2.5 mm. latis, integris, minoribus

linearibus ad 5-6 mm. longis, vix 0.3 mm. latis, integris, perianthio

ipso profunde parti to, petalis (videtur) subnullis, glandulis obscuris;

ovario albocitrino, hispido-tomentoso, ad 4 mm. lato, 3 mm. longo,

stylis 3 filiformibus, hispidulis, quove fere ad basem in lacinias

2 ca. 4 mm. longas partito. — Guatemala, Dept. Jutiapa, vicinity of

Jutiapa, alt. about 850 meters, "brushy rocky slope. Shrub 2 meters,

common," Standley 74971, Oct.-Nov. 1940 (type in Herb. Arnold

Arb.; duplicate in Herb. Field Mus.).

Suggests a tomentose form of C. Guildingii Griseb. of Grenada

in the West Indies, but is immediately characterized as distinct by

the anisomerous lobes of its 9 perianth. Resembles in habit and

foliage C. macrodontus Muell. Arg., of Veracruz in Mexico, but has

an altogether different 9 flower.

Croton lasiopetaloides, sp. nov. — Frutex ad 1.5 m. altus.

Innovationibus inflorescentiisque pube stellata brunnea vel cinerea

indutis. Foliis ovatis vel ovato-lanceolatis breviter acuminatis,

basi rotundatis vel levissime cordatis, primo utrinque tomentosis

serius supra glabratis vel glabris, subtus semper cinereo-tomentellis,

in sicco e lutescentibus (foliis emorientibus) olivaceo-brunneis, ad

marginem quam ad laminam confertius pustulosis, subintegris, 7-5

NEW SPECIES OF CROTON 451

cm. longis, 5.5-3.5 cm. latis, nervis primariis ad 5 late patentibus,

supra impressis cum trabeis, primo jugo laminae dimidium attin-

gente; petiolo ca. 2 cm. longo, glandulis 2 anticis parvis, stipulis

glandulosis valde obscuris. Inflorescentiis subspicatis brevibus.

Floribus cf ignotis. Floribus 9 subsessilibus vel sessilibus, perian-

thio ca. 5 mm. lato, lobis 5 discretis lineari-acuminatis ca. 2 mm.

longis, 0.75 mm. latis, basi quove glandula sessili sat magna obsito,

integerrimis, dorso parcius tomentosis, ovario hispido-albicante baud

tomentello ad 2.5 mm. longo, 3.5 mm. lato; stylis 3 ad 3.5 mm.

longis, quove fere ad basem partito, columella fructu delapso ca.

6 mm. longa, apice manifeste triloba incrassata. — Guatemala,

Dept. Huehuetenango, mountains west of Aguacatan on the road

to Huehuetenango, alt. about 1,950 meters, "in oak forest, shrub

1.5 meters tall," Paul C. Standley 81219, Dec. 1940 (type in Herb.

Arnold Arb.; duplicate in Herb. Field Mus.).

In certain details of its flowers and, so far as seen, in the conspic-

uous leaf buds, C. lasiopetaloides suggests C. botryocarpus Croiz.,

but actually differs from it in the following characters: (1) The

ovary is definitely hispid, not tomentellous or glabrate; (2) the

perianth lobes are somewhat narrower, acuminate, and slightly

tomentose, neither ligulate nor glabrous; (3) the foliage is not

pinnatinerved and has a lesser number of primaries, the over-

development in length of the first pair causing the venation to appear

as if 3-nerved; (4) the leaves at unfolding are tomentose on both

faces and remain so a while; in C. botryocarpus the leaves, even at

unfolding, are scarcely hairy above and become there almost im-

mediately glabrous; (5) the columella after dehiscence is stouter

and somewhat longer.

The specific name is derived from Lasiopetalum Sm., a genus of

the Sterculiaceae, and in particular from Lasiopetalum Schulzenii

Benth., an Australian species which C. lasiopetaloides appears to

suggest in habit and in foliage. It is worth noticing here that the

Sterculiaceae Lasiopetaleae are closely related with Croton, the

evidence to be derived from floral morphology and general somatic

characters indicating that both Croton and Lasiopetalum arose con-

temporaneously from a common malvoid ancestor in the same vicinity.

Croton limnocharis, sp. nov. — Frutex 2-4-metralis. Innova-

tionibus inflorescentiisque pube stellata albida tomentellis, demum

glabratis. Foliis optime ovato-cuspidatis vel ovato-lanceolatis, basi

plus minusve obvie cordatis, 12-6 cm. longis, 7.5-3 cm. latis, in

sicco supra olivaceis subtus pube tenui stellato-flocculosa detergibili

452 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

griseo-tomentellis, tenuiter chartaceis, margine vix repandulis vel

subintegris, venis primariis utrinque ca. 6-jugis, primo jugo laminae

tertium inferum vel (frequentius) dimidium attingente, ramoso;

petiolo sat gracili, tomentello, ca. 2.5 cm. longo, glandulis caute

inquirendis 2 patelliformibus sessilibus posticis, stipulis nullis vel

subnullis. Floribus cf ignotis. Floribus 9 : perianthio delicato

vix 3 mm. lato, pedicello ca. 2 mm. longo, lobis triangularibus vix

1-1.5 mm. longis, ovario ellipsoideo toto albicante tomentello ad

2 mm. longo, 1.5 mm. lato; stylis 3, ad 2-2.5 mm. longis, quove

fere ad basem parti to; capsula matura globulosa, ca. 7 mm. longa

lataque, leviter trigona, pedicello fulta ca. 2 mm. longo, indumento

facillime detergibili; semine scaraboideo applanato, ad chalazam

rotundato-acutato, ad carunculam subtruncato-apiculato, 4.5 mm.

longo, fere totidem lato, caruncula valde depressa ad 2 mm. lata,

testa sublevi, arillo tenui subnitido lutescente. — Guatemala, Dept.

Jutiapa, vicinity of Jutiapa, alt. about 850 meters, "damp thicket,

shrub 2-4 meters," Paul C. Standley 75215, Oct.-Nov. 1940 (type

in Herb. Arnold Arb.; duplicate in Herb. Field Mus.).

This new species is closely allied with C. pungens Jacq. and C.

sarcopetalus Muell. Arg., that is to say, with a group having pantropic

American range. It may be easily confused at sight with C. pseudo-

xalapensis Croiz., from Honduras, which it resembles in habit, but

it plainly differs from this species on account of the venation, which

definitely tends to be 3-nerved, not pinnate. The 9 flower and fruit,

moreover, are pediceled and not sessile, and the seed is smoothish,

not manifestly costate-rugose.

The extensive material of C. xalapensis H.B.K. and C. pseudo-

xalapensis Croiz. that I have seen in the past three years makes me

doubt that it will be possible to maintain these two binomials as

distinct in a final revision of the group. Croton xalapensis was

originally reported from the vicinity of Veracruz in Mexico, and

C. pseudoxalapensis is endemic to western Honduras. So far, my

experience has been that the species of Euphorbiaceae with a wide

range occur preferably on the eastern coast of Central America

and Mexico, the western coast, on the contrary, being exceedingly

rich in localized endemics. Under the circumstances, it may easily

prove possible that the same species occurs in Honduras and in the

state of Veracruz, Mexico.

Croton quercetorum, sp. nov.— Frutex vel arbuscula ad 8 m.

alta. Innovationibus inflorescentiisque argillaceo-tomentosis sordi-

dis vel brunnescentibus, indumento serius deciduo. Foliis ovato-

NEW SPECIES OF CROTON 453

ellipticis vel triangulari-ovatis vel lanceolatis, acuminatis vel cuspi-

datis, basi rotundatis vel cuneatis, 14-7 cm. longis, 9.5-3.5 cm.

latis, firme chartaceis, in sicco brunneo-olivaceis discoloribus, novellis

utrinque tomentosis, citius glabrescentibus, adultis supra glabris,

subtus trichomatibus stellatis longe radiatis parcissime indutis,

margine irregulariter dentato-serratis, dentibus sub lente glandulosis,

venis primariis utrinque ca. 5-jugis, primo jugo laminae ipsius

dimidium attingente obscure ramoso; petiolo graciliore 2.5-4.5 cm.

longo, tomentello, apice glandulis ca. 4 patelliformibus vulgo adaxia-

liter ornato, stipulis lentis, simplicibus vel 2-3-ramosis, gracilibus,

ad 1.5 cm. longis, sat persistentibus. Inflorescentiis terminalibus

simplicibus, habitu spicatis ad 20 cm. longis. Floribus d" ignotis.

Floribus 9 (sub fructu tantum notis): perianthio ad 7 mm. lato,

pedicello vix 2 mm. longo, lobis integris, triangularibus, ca. 4 mm.

longis, 2 mm. latis, dorso indumento rufescente grosso indutis;

coccis delapsis ad 12 mm. longis, subligneis, glabratis; semine 10 mm.

longo, 6.5 mm. lato, scaraboideo, caruncula vix 1.25 mm. magna,

applanata, testa utrinque pinnatim costato-rugosa, arillo subnitido

tenui lutescente. — Guatemala, Dept. Jalapa, mountains about

Chahuite, northwest of Jalapa, alt. 1,650 meters, "damp oak forest;

shrub or tree to 8 meters; abundant," Paul C. Standley 77460, Nov.

1940 (type in Herb. Arnold Arb.; duplicate in Herb. Field Mus.).

A species in the general affinity of C. corylifolius Lam., of the

West Indies, strongly characterized by its stipules, indument, and

capsule. It resembles certain states of C. jalapensis Croiz., but can

easily be distinguished from it on account of its larger capsule and

the seed, which is grossly rugose-costate on both its faces.

UNIVERSITY OF ILUNplS-URBANA

III III 1 III HI (I ' 'I' " ' * •"

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