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PROCEEDINGS
OF THE
SCIENTIFIC MEETINGS.
OF THE
ZOOLOGICAL SOCIETY
OF LONDON
FOR THE YEAR
1878.
AND SOLD AT THEIR HOUSE IN HANOVER SQUARE,
LONDON:
MESSRS, LONGMANS, GREEN, READER, AND DYER,
PATERNOSTER ROW,
Lik-s80
OF THE
COUNCIL AND OFFICERS
OF THE
ZOOLOGICAL
SOCIETY OF LONDON.
1878.
COUNCIL.
Tue Marauis or TweEeppaLr, F.R.S., President.
Tue Lorp ABINGER.
Georez Busx, Esq., F.R.S.
Masor-Gen. Hunry Crerx, R.A.,
F.R.S.
Rozert Drumuonp, Esq., Trea- |
surer.
ProFessor Frower, F.R.S.,
President.
Joun P. Gassrorr, Jun., Esq.
Rear Apurrat Tue Count Guer- |
cHEn, R.N.
F, DuCanz Govan, Esq.
Arruur GRore, Esq.
Epuounp W.H., Horpsworta, Esq.
Vice- |
| Ropert Hupson, Esq., F.R.S.,
Vice-President.
| Proressor Mivart, F.R.S.
| Proressor Newton, F.R.S., Vice-
President.
| Henry Poxtocs, Esq.
Lievt.-Cot. Str Cartes Rus-
sELL, Bart., M.P.
| | Ospert Satvin, Esq., F.R.S.
Pate Luriey ScLaTER, Esq.,
M.A., Pa.D., E.RS., Secretary.
_ Josrpn Travers Suitu, Esq.
_ Ricnarp H. &. Vxvyan, Esq.
| Tur Lorp WALsinenan.
PRINCIPAL OFFICERS.
Office, 11 Hanover Square, W.
P. L. Scrarer, Esq., Ph.D., F.R.S., Secretary.
Mr. F. H. Warernouse, Librarian.
Mr. W. J. Wrt1aMs, Clerk.
Gardens, Regent's Park, N.W.
Mr. A. D. Barrrerr, Superintendent.
Proressor Garrop, F.R.S., Prosector.
LIST
OF THE
CONTRIBUTORS,
With References to the several Articles contributed by each.
Aston, Epwarp R., F.L.S., F.Z.8., &e. ae
Note on the Dentition of Cuscus .... 2... ee cece cece es 274
Exhibition, on behalf of Dr. Elliott Coues, C.M.Z.S., of
two specimens of Synaptomys cooperi, Bd. ay See . 633
On the Squirrels of the Neotropical ae (Plate XLI. 1) 656
Supplementary Note on the Neotropical Squirrels ...... 954
AnpeErson, ANDREW, F.Z.S.
Letter from, concerning Falco babylonicus, figured in the
Baccigs- Proceedings 106 157 O% rac ia «0! «5 accls's oo 0,2 ae
On a new Species of Indian Prinia. (Plate XIX.) .... 370
Exhibition of a Bamboo driving-stick taken from the nest
pen teria Pineda Fe td toe a Ses ts ese as tors O9S
9
a
Aneas, Grorce Frencu, F.L.S., F.R.G.S., C.M.Z.S., &e.
Descriptions of seven new Species of Land-Shells recently
collected in Costa Rica by Mr. Adolphe Boucard. (PlateV.) 72
Description of a new Species of Latiavis. (Plate V.) .. 74
Description of a new Genus of Land-Shells belonging to
tip Faunly Cyclophorides,.. scsi. 5 hos fo eens sea alo
iv
Descriptions of nine new Species of Land and Marine
Shells from various localities. (Plate XVIII.).. ....
Description of a new Species of Tudicula.........0.00+
Descriptions of six Species of Bivalve Shells in the Collec-
tion of Mr. Sylvanus Hanley, F.L.S., and of a Helix from
Page
~ volt
the Solomon Islands. (Plate LIV.) ............... soa Bae
Descriptions of ten Species of Marine Shells from the Pro-
vince of South Australia. (Plate LIV.)........ < 361
A List of additional Species of Marine Mollusca to be in-
cluded in the Fauna of the Province of South Australia, with
Notes on their Habitats and Local Distribution ..........
Barpoza pu BocaGe, Professor Jost Vicente, Director of
the National Museum, Lisbon, F.M.Z.S.
Liste des Antilopes d’ Angola
Bartiett, A. D., Superintendent of the Society’s Gardens.
Letter from, relating to the alleged existence of Anoa de-
pressicornia in the Philippines . on... coeesejsec0 0% os sere
Bates, H. W.., F.L.S., F.Z.S.
On new Genera and Species of Geodephagous Coleoptera
from-{Genttal AMeres .'c.04, 150 cerse eee: eee eee ee
On new Species of Coleopterous Insects (Geodephaga and
Longicornia) taken by Dr. Stoliczka during the Forsyth Ex
pedition to Kashgar in 1873-74 .............. we Paes ta
Brppomg, Lieut.-Col. R. H., C.M.Z.S.
Description of a new Genus of Tree-Lizards from the higher
Ranges of the Anamallays. (Plate XIV.) ...... ......
Descriptions of new Uropeltidze from Southern India, with
Remarks on some previously described Species
Description of a new Genus of Snakes of the Family Cala-
maride, from Southern India ..............ec06
Description of anew Batrachian from Southern India, be-
longing to the Family. Phryniscide ....,...........00.-
864
741
882
713
153
154
v
Description of six new Species of Snakes of the Genus
Silybura, Family Uropeltidz, from the Peninsula of India. .
Bock, Cart, F.G.S. &c.
Descriptions of two new Species of Shells from China and
Japan: (Plate SLM L., figs. 1—Si)i pes 5.0%) stedles latte nese e
Bovucarp, Apourue, C.M.Z.S8.
On Birds collected in Costa Rica. (Plate IV.) ........
Notes on some Coleoptera of the Genus Plusiotis, with
Descriptions of three new Species from Mexico and Central
Ameriés. (Plate XVI)" 05 oar. Saige writ sete eins easels
Brooke, Sir Victor, Bart., F.Z.S.
Onileazelle grantees os ates. SS Saeieas sleet Meaecees<. Os
On the Classification of the Cervide, with a Synopsis of
the existing Species. (Plate LV.) .........
On a new Species of Gazelle from Eastern Africa.
Pence Leh oa tte ites a's aks sco we, oc apenas elena e Slane
+
Burmeister, Dr. Hermann, Diretor of the National Museum,
Buenos Aires, F.M.Z.S.
Notes on Conurus hilaris and other Parrots of the Argen-
tine Republic......
Butter, ArTHuR Garpin_er, F.L.S., F.Z.S., Assistant, Zoo-
logical Department, British Museum.
On a small Collection of Lepidoptera obtained by the Rev.
J. S!'Whitmee‘at the Ellice Islands. .:. 0.0 .... 0.0.05...
Description of new Lepidoptera of the Group Bombyecites,
in the Collection of the British Museum ................
On a small Collection of Lepidoptera from Jamaica......
Description of a new Species of the Orthopterous Genus
Phylloptera from Madagascar .......
Deseription of a Remarkable new Spider from Madagascar
Page
800
727
293
723
79
296
648
799
vi
Page
Cuersemay, S. T., F.L.S., Curator of the Auckland Museum.
Descriptions of three new Species of Opisthobranchiate
Mollusca from New Zealand. (Plate XV.)..........-..- 275
Cuark, Joun W., M.A., F.Z.S.
Notes on three stuffed Specimens of the Sea-lion of the Pri-
bilov Islands (Otaria ursina). (Plate XX.) ............ 371
Couuett, Rosert, C.M.Z.S.
On Latrunculus and Crystallogobius, two remarkable Forms
of Gobioid Bishes: i"). “aes. Tia Aa ae sw ee eee 318
Cougs, Dr. Exurott, C.M.Z.S. (See Atston, Epwarp R.)
Day, Surgeon-Major Francis, F.Z.8. &e.
Remarks on Mr. Whitmee’s paper on the Manifestation of
Rearend. Anger in’ Bishes ..\ 2: Ba he > + tent ogee ean leeeaee eee
Exhibition of, and remarks upon three jaws of Indian
Sharlene =< i015 So eee es os Seem ccs ener es oe Pate Coe
<=
Dosson, G, E., M.A., M.B., F.L.S., &e.
Additional Notes on the Chiroptera ct Duke-of-York Island
and the adjacent Parts of New Ireland and New Britain.... 314
Notes on Myxopoda aurita, Milne-Edw. .............. 871
Notes on recent Additions to the Collection of Chiroptera
in the Muséum d’ Histoire Naturelle at Paris, with Descrip-
tions of New and Rare Species.................. So Gon oe 873
Druce, Hersert, F.L.S., Gopman, Frepericx Du Cane,
and Savin, Ossert, M.A., F.R.S.
Catalogue of Lepidoptera collected by Mr. S. N. Walter
in the Island of Billiton. (Plate XL.) ........ 0.02 000. 637
Ex.iot, Danrex G., F.R.S.E., F.L.S., F.Z.8., &e.
A Study of the Pteroclide or Family of the Sand-Grouse 233
On the Fruit-Pigeons of the Genus Pfilopus. (Plates
NUTT MLV) dete waaies «wane ++. cence OOM
Vil
Everett, A. H.
Letter from, addressed to the Marquis of Tweeddale, con-
cerning the occurrence of Anoa depressicornis in the Philip-
TER ycteap-o at aati plates dete 2 ac a ea So ag = ay ee
Finscu, Otto, Ph.D., C.M.Z.S., &c., Director of the Bremen
Museum.
On a new Species of Finch from the Feejee Islands. (Plate
2 LMI EN RSC TEL te ee Ae eee Bee
Fiower, Witu1am Henry, LL.D., F.R.S., F.L.S., V.P.Z.S.,
Conservator of the Museum of the Royal College of
Surgeons, and Hunterian Professor.
On the Skull of a Rhinoceros(R. lasiotis, Scl.?) from India
Forses, Witit1am ALEXANDER, F.Z.S.
Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘ Challenger.’—No. VII. On the Birds of
Cape York and the neighbouring Islands (Raine, Wednesday,
Ste OU Y EN HOS ts cienn sid pws. algae ag Moines ae res Lae
On a small Collection of Birds from the Samoan Islands
and the Island of Rotumah, Central Pacific... ............
Garrop, Atrrep Henry, B.A., F.R.S., F.Z.S., Fellow of
St. John’s College, Cambridge, Prosector to the Society.
On the Systematic Position of the Momotide..........
Page
440
712
634
120
351
100
Note on the Gizzard and other Organs of Carpophagalatrans 102
Note on the Anatomy of the Binturong (Arctictisbinturong) 142
Notes on the Anatomy of Passerine Birds.—Part IV..... 143
Notes on the Anatomy of Tolypeutes tricinctus, with Re-
marks on other Armadillos ...... . oo 222
Notes on the Visceral Anatomy of Zycaon pictus, and of
Nyctereutes procyonides ........ Ree JA: «. Seme 373
On the Trachea of Tantalus loculator and of Vanellus
MUMPIRMIMENG No ae Soe Ce tae Loew 0s ga ea sane h ameee oe 625
On the Anatomy of the Maleo (Megacephalon maleo).... 629
Vili
Page
Notes on Points in the Anatomy of Levaillant’s Darter ;
CP lots Lop allonis) -0\<,..5 +: exaltsn tip mtr» wiblegeel oer eee 679
Notes on the Anatomy of Indicator major . . 930
Garrop, Atrrep Henry, B.A., F.R.S., and iter
WiuuiaM, M.B., F.R.S.
On the Gravid Uterus and Placenta of Hyomoschus aqua-
ticus. (Plate XLIV.)...... . 682
Geppes, P.
Notice of a memoir on the Mechanism of the Odontophore
in certain Mollusca .. . 212
Gopman, Freperick DuCang, F.L.S., F.Z.S., and Satvin,
Ospert, M.A., F.R.S., F.Z.S.
Descriptions of nineteen new Species of Diurnal Lepido-
pteraifrom CentralAmeriog: .\.. . (tecdsist Ale. liad ac 264
Descriptions of new Species of Central-American Butter-
flies of the Family Erycinide ......... sinha’ so spe tre ae ean
List of the Butterflies collected in ext New Guinea
and some neighbouring Islands by Dr. Comrie during the
Voyage of H.M.S. ‘Basilisk.’ (Plate XL.) .... 643
Descriptions of some apparently new Species of Butterflies
from New Ireland and New Britain, sent by the Rev.G. Brown 733
GopmaNn, Freprerick Du Canez, Satvin, Ospert, M.A.,
F.R.S., and Drucr, Hersert, F.L.S.
Catalogue of Lepidoptera collected by Mr. S. N. Walter in
the Island of Billiton. (Plate XUL.)...< 2... .s.2e0 bangs 637
GintuHer, ALBERT, M.D., Ph.D., F.R.S., V.P.Z.S., Keeper
of the Zoological Department, British Museum.
On Reptiles from Midian collected Ly Major Burton.
(Eine eye: ee ete ae re eee eee 977
Gurney, Joun Henry, F.Z.S.
Letter on a supposed example of Falco babylonicus 2
Notes on a Specimen of Polydorus lately living in the
Society’s Gardens .......... 230
Remarks upon the Habits of Asturinula monogrammica.. 791
Harrravs, Dr. G., F.M.Z.S.
On a new Species of Notauges (N. hildebrandti, Wap.) 72)
Hotmwoop, F.
Remarks upon a young specimen of Temminck’s Manis
RMUEINCMMITICNE Ye ee, ke ee =a), G32
Huxtry, Tuomas H., LL.D., Sec.R.S., V.P.Z.S.
On the Classification and the Distribution of the Crayfishes 752
Irpy, Col. L. H. Loyn, F.Z.S.
Exhibition of nests, eggs, and young of Cypselus pal-
OI a Fa ns 976
Jacosy, Martin.
Descriptions of new Species of Phytophagous Coleoptera. 144
Descriptions of new Species of Phytophagous Coleoptera
from Central and South America...................... 982
Jerrreys, J. Gwyn, LL.D., F.R.S., F.Z.S.
On the Mollusca procured during the ‘ Lightning’ and
‘ Porcupine’ Expeditions, 1868-70. (Part I.) (Plates XXII.
Lanxester, Prof, E. Ray, M.A., F.R.S.
Notice of a memoir on the Hearts of Ceratodus, Proto-
plermevand, Ohinterar’ i0n5 < da ijecennedd bokuTht oo. ee 634
Layarp, E. L., C.M.G., F.Z.8S., &c., H.B.M. Consul in New
Caledonia. :
Letter from, addressed to the Secretary, concerning Glycy-
phila fasciata and Petreca kleinschmidti .............. 655
Lover, Eomunp Giuus, FZ.S.
Exhibition of a head of a peculiar variety of Bos americanus 392
x
5 Page
Meyer, Dr. A. B., Director of the Royal Zoological Museum,
Dresden, C.M.Z.S.
Letter from, addressed to the Secretary, respecting a new
BeEG-GL PRGA.) ica oc0.0.4 » 0-005 5) wi mto aur = tn aie 9 792
Letter from, concerning the supposed existence of the Anoa
(Anoa depressicornis) in the Philippines.............- -. 881
Miers, Epwarp J., F.L.S., F.Z.S., Assistant in the Zoological
Department, British Museum.
Notes on the Penegide in the Collection of the British
Museum, with Descriptions of some new Species. (Plate
pA | ee a ee ae age eR TIES, “MS eo 298
Mivart, St. Gzorer, Ph.D., F.R.S., F.Z.S., M.R.L., &e.
Notice and abstract of a memoir on the Fins of Elasmo-
branchs, with Considerations on the Nature and Homologies
OLAVIELEG Oatley IGIMUS aJsrcrs sets ceyee's os reves: olsnerd ous aucrese ean
Moorer, Freperic, F.Z.S., Assistant Curator, India Museum,
London.
A Revision of certain Genera of European and Asiatic Li-
thosiide, with Characters of new Genera and Species.
Plates TU) 9 fk hegih 3s Dal kh gucinirem - caret 3
Descriptions of new Asiatic Hesperidee. (Plate XLV.) .. 686
List of Lepidopterous Insects collected by the late R.
Swinhoe in the Island of Hainan ..................02-- 695
A List of the Lepidopterous Insects collected by Mr. Ossian
Limborg in Upper Tenasserim, with Descriptions of new
Species, “(Plates Gil LTE.) esis iis ss oe ne yoo
Newron, ALFRED, M.A., F.L.S., F.Z.S., Professor of Zoolog
and Comparative Anatomy in the University of Cam-
bridge.
Exhibition of, and remarks upon, a stone supposed to be
from the gizzard of Pezophaps solitaria ................ 291°
Exhibition of, and remarks upon, a supposed hybrid between
the Red Grouse and the Ptarmigan .....-.....e.2--00-. 7938
xl
Page
Nicuouson, Francis, F.Z.S.
On a Collection of Birds from Abeokuta. (Plate X.) ... 128
A List of the Birds collected by Mr. E. C. Buxton at
Darra-Salam, on the coast of Africa opposite Zanzibar - 353
On an apparently new Species of American PE ce atone 390
OvustaeT, Emits, C.M.Z.S.
Description d’une nouvelle espéce de Casoar (Casuarius
eaoran).-"- (Piste 2 XP, yey oO? MeN hd. be ouaele te 389
Owen, Prof. Richard, C.B., D.C.L., F.R.S., F.LS., F.Z.S., &e.
On the Relative Positions to their Constructors of the
Chambered Shells of Cephalopods (Plate EX.) ielshera cen we 955
Parker, T. Jerrery, Assoc. R.S.M.
Remarks upon the stridulating organ of the Common
ae ase am usin MA SS cs «stung Abeer BOD
Note on the Stridulating Organ of Palinurus vulgaris.
(Plate XXX.) ....:. aA citis o.9'e chest HE nas wid he ta Bote 442
Satvapor1, Counr Tuomas, M.D., C.M.Z.S.
Reports on the Collections of Birds made during the Voyage
of H.M.S. ‘Challenger.,—No. VI. On the Birds of Ternate,
Amboyna, Banda, the Ké Islands, and the Aru Islands . 78
Satvin, Ossert, M.A., F.B.S., F.ZS.
Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘Challenger. —No. XII. The Procella-
RE eo hse hehe a th ee mf 735
Satvin, Ossert, M.A., F.RB.S., F.Z.S., and Gopman, Fre-
DERICK Du Cang, F.L.S., F.Z.S.
Descriptions of nineteen new Species of Diurnal Lepido-
ptera from Central America .. . -» 264
Descriptions of new Species of Central-American Butter-
flies of the Family Erycinidee ...........0.... cece eee 360
xii
Page
List of the Butterflies collected in Eastern New Guinea and
some neighbouring Islands by Dr. Comrie during the Voyage
of BMS, * Bastiek.” ~ (Plate JEL.) ... a. ance 2 eee
Descriptions of some apparently new Species of Butterflies
from New Ireland and New Britain, sent by the Rev. G.
733
Satvin, Ossert, M.A., F.R.S., Gopman, Freperick Du
Cane, and Drucr, Hersert, F.L.S.
Catalogue of Lepidoptera collected by Mr. S. N. Walter
in the/lslandyof Billitons- (Plate Xalis)s =. ee eee 637
Satvin, Ossert, M.A., F.R.S., F.Z.S., and Scuater, P. L.,
M.A.) PH.D, FRB:
On the Collection of Birds made by Prof. Steere in South
Msdericn.- (Plates Sete NUT ye i Sie ie ois Ete 135
Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘Challenger.,—No. IX. On the Birds
GE TIAN CLD AMI EPICA Zee ti "se eeet = ales sale Sale Se oe te a
Descriptions of three new Species of Birds from Ecuador.
(Plates RVD. & MX VIET). enn wre ses s caver lwlaene ae
Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘Challenger..—No. XI. On the Stega-
nopoiles ahd: Thapenmes! 9 e575. 0 eee ent aed 92 2
SaunpeErRs, Howarp, F.L.S., F.Z.S.
On-the:Zarmcor Gals: 1.2. feet ees Sen ee ee eae
Exhibition of, and remarks upon, some Indian Eggs .... 976
Sciater, Purie Lutiey, M.A., Ph.D., F.R.S., Secretary to
the Society.
Report on the additions to the wits Menagerie in
December 1877 . SAMeE : ce I
Report on the adiiGans to Te Society’: 8 eee in
Januar lS7S. paso. cos sews ese so od ae ee he onee 115
Exhibition of, and remarks upon, a skin of a Cassowary in
the collection of the British Museum .................. 212
Report on the additions to the hacek s Menagerie in
Meviuary sha tsi. seca al ee ek ee oe Te ee 289
xiii
Page
Exhibition of, and remarks on, a Second Collection of
Birds from Duke-of-York Island, New Britain, and New
Ireland, received from the Rev. G. Brown, C.M.Z.S....... 289
Exhibition of, and remarks upon, the typical specimen of
Ninox solomonis, Sharpe ..........-+...- aie chek gs ois < ae
Exhibition of a Coot, probably the typical specimen of
rater GaerOT eR i). bs Sa rod dea cing tlt eo) oS bee 291
Exhibition of, and remarks upon, the type specimen of
Dicrurus marginatus of Blyth ........02.. 22.048 ihe tee 339
Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘ Challenger..—No. VIII. On the Birds
pf the ard wont Petandae sad. oie ecnie aicien so wisid's 08 clare 346
Report on the additions to the Society’s Menagerie in
Makeb S78 ycral ees deeaes ah ae AE ahs dh ome 4 det OER
Exhibition of, and remarks upon, a typical specimen of
a new Box( Vulpes Cae) aia igw iain ies aE 0'n,0 neat alsi me binyaty S92
Report on the additions to the Society’s Menagerie in
Papeitee Wage se Pee oe oc a. aS eer ode AA)
Further Remarks on Fuligula nationi. (Plate XXXII.).. 477
Reports on the Collections of Birds made during the Voyage
of H.M.S. ‘Challenger’—No. X. On the Birds of the At-
lantic Islands and Kerguelen’s Land, and on the Miscella-
MOE HICCORE (on. ne as eR ae Oe atte t oes 6 ~- 976
Report on the additions to the Society’s Menagerie in
May 1878, and Remarks on Tapirus roulini. (Plate
a Fakta hss hg Cs ua inlay. Sees Aaa aes «- 631
Remarks upon the living examples of Ciconia maguari and
Ciconia episcopus now in the Society’s Gardens .......... 633
Note on a second specimen of Felis lanea ............ 655
Notice of a supplementary memoir on the Curassows (Cra-
RE Sram a4 hS onias ©. daakes wee Sas
On a third Collection of Birds made by the Rey. G.
Brown, C.M.Z.S., in the Duke-of-York Group of Islands
mid! Morvicuatey. “(Plate A MIT.)--.... o. 0. seats eerewad dete: 670
xiv
Page
Report on the additions to the Society’s Menagerie in
June, July, August, September, and October 1878........ 788
Notice of some hybrid Monkeys lately born in the Society’s
Menpsenieter: ein oe oe Perea oe eats eae 791
Exhibition of, and remarks upon, a specimen of Sasicola
stapazina killed in Lancashire .........+..+.ee00++-.00 S81
Report on the additions to the Society’s Menagerie in
November 1876. > (Plate LAR oo rns cect alnvewaciem penne
Further remarks on the occurrence of Sazicola stapazina
AN RARE aos di wor 's aS 8 Ge ws Bes we SS ge yg go
Notice of an error in the last Part of the Society’s Pro-
CERGERE i sire Soper wins, s anit aad Gop? oo cies 6s ea
Scuater, P. L., M. A., Ph.D., F.R.S., and Satvin, Ospert,
M.A., F.R.S., F.Z.S.
On the Collection of Birds made by Prof. Steere in South
Amervaor€P lites SA RL) 05), PG a8. Veh. FI 135
Reports on the Collections of Birds made during the Voyage
of H.M.S. ‘Challenger..—No. IX. On the Birds of An-
HEE ATA CRIED 3 2-2 a Saad a chps ano seas da) manana: eee 431
Descriptions of three new Species of Birds from Ecuador
(Plates RV UR) GOR EE! ee es
Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘ Challenger.’-—No. XI. On the Stega-
hopades jandInspennes\! 2.4. st. SI Ee ted ee ce 650
Srrsoum, Henry, F.Z.S.
Exhibition of and remarks upon a large series of skins of
the Hooded and Carrion Crows.......... es. ee ee cies 976
On a new Species of Sylvia from Abyssinia, and on some
other A DyssMIMG RS YIVIUNE . Sac. vee csi le mes we a a oe pe
On the Identity of Horornis fortipes, Hodgs., Neornis
assimilis, Gray, Horeites robustipes, Swinhoe, Horeites
pallidus, Brooks, and Horettes brunnescens, Hume,....,.. 980
Xv
Page
Saarer, R. Bownter, F.L.S., F.Z.S., &c., Senior Assistant,
Zoological Department, British Museum.
On a new Species of Indicator, with Remarks on other
Bpectesan tne Gin. oy Sah ye os. wines ones oe oy so 5.0% 793
A Note on Peoptera lugubris, Bp. (Plate XLIX.)...... 802
SHarpr, R. Bownter, F.L.S., F.Z.S., and Wairmer, Rev.
S. J., C.M.Z.S.
On a small Collection of Birds from the Ellice Islands,
with a Note on other Birds found there ................ 271
Smiru, Enear A., F.Z.S., Zoological Department, British
Museum.
Description of a new Species of Helix from Japan ...... 105
Descriptions of new Land-Shells from Japan and Borneo.. 495
Descriptions of five new Shells from the Island of Formosa
and the Persian Gulf, and Notes upon a few known Species.
(Pinte SLYI. fies. 4-14.) x, ose Hoe wed sepedits Gacteind ales geo
On a Collection of Marine Shells from the Andaman
SEs) eA MMMM a oboe 6. annie (v0. 5: \)) nspg-ininlo duwoleparn Susy 804
Sowersy, G. B., Jun.
Descriptions of ten new Species of Shells. (Plate XLVIII.) 795
Turner, Wiitiam, M.B., F.R.S., and Garrop, A.H., M.A.,
F.R.S.
On the Gravid Uterus and Placenta of Hyomoschus aqua-
Rae RS ALUN REED MLR V 8) SS crc iae vinnie. ealdiiaie’s «pes. Wea ge epee g 682
TwEEDDALE, ArTHUR, Marquis of, F.R.S., President of the
Society.
Contributions to the Ornithology of the Philippines.—No.
IV. On the Collection made by Mr. A. H. Everett in the
Islands of Dinagat, Bazol, Nipah, and Sakuyok. (Plates
We AVE arti eo shies fn He ae BR ee 106
Xvi
On anew Philippine Genus and Species of Bird. (Plate IX.)
On anew Species of the Genus Buceros ........-.+-+-
Contributions to the Ornithology of the Philippines.—No.
V. On the Collection made by Mr. A. H. Everett in the Island
of Negros: sortie SS oan we
Contributions to the Ornithology of the Philippines.—
No. VI. On the Collection made by Mr. A. H. Everett in the
Psi Oy Ges et. eee wit ere = ne piaan es thee eosin ees
Contributions to the Ornithology of the Philippines.—No.
VII. On the Collection made by Mr. A. H. Everett in the
Island of Panaon
Contributions to the Ornithology of the Philippines.—
No. VIII. On some Luzon Birds in the Museum at Darm-
atadt.. (Plate, XV.) ers eoalid tsi. eA. INS
Contributions to the Ornithology of the Philippines.—
No. IX. On the Collection made by Mr. A. H. Everett in
the Island of Palawan. (Plates XXXVII. & XXXVIII.) ..
Contributions to the Ornithology of the Philippines.—
No. X. On the Collection made by Mr. A. H. Everett in the
Esland.of Beliol p44 tea ee SE
Correction of an error in his last paper on Philippine Birds
Contributions to the Ornithology of the Philippines.—
No. XI. On the Collection made by Mr. A. H. Everett at
Zamboanga, in the island of Mindanao. (Plates LVII.-LIX.)
Watson, M., M.D., Professor of Anatomy, The Owens College,
Manchester.
On the Male Generative Organs of Hyena crocuta.
(Plates: XIV Ds gee ae ens vi me tRNA ase
On the Male Generative Organs of Chlamydophorus
truncatus and Dasypus sexcinctus. (Plate XLIII.)
Page
114
.. 280
. 339
429
611
936
.-. 416
Xvil
Westwoop, Professor J. O.
Notice of a memoir containing a synopsis of the Uraniide,
a family of Lepidopterous Insects, together with a complete
monograph of Coronilia, one of the genera of which it is
Page
composed mols
Wuire, F. Bucuanan, M.D., F.L.S.
Contributions to a Knowledge of the Hemipterous Fauna of
St. Helena, and Speculations on its Origin. (Plate XXXI.) 444
Wuitme:E, Rev. S. J., C.M.Z.S.
On the Manifestation of Anger, Fear, and other Passions
in Fishes, and on the Use of their Spines...... sureate oe. 132
Reply to Mr. Day’s remarks on his paper on the Manifes-
tation of Fear and Anger in Fishes .................... 221
Wuirmesg, Rey. S. J., C.M.Z.S., and SHarpg, R. Bowpter,
F.L.S., F.Z.8., &e.
On a small Collection of Birds from the Ellice Islands, with
a Note on other Birds found there .................... 271
Woop-Mason, J., F.Z.S.
Oa new and little-known Mantide. (Plates XXXV. &
Sc Bins SEE a ah ae iia a 580
On the Structure and Development of the Trachea in the
Indian Painted Snipe (Rhynchea capensis). (Plate XLVII.) 745
har Says 9
“ibaa he hie er gree i he etheen f) j igs 7a
=o oe tact S Foci ing sais Ae ii e ae svigees one
4
es aire Speed yin ae ee ie ny thee Mi ¢ Gh ant aad
aia een ery gee XM ste eee uy ine cae
MY
eee sr ee
per oi ia a) alia a & id nensieges ve aul we) ie : ES
essa Pi ee i ea i] cit if fie Ene a eta
,
e Ki pees eee
‘oapedas% dis we ie Sait ae nie
oe fe anes hice ats a yk att ae
en Rey SR Bek» ee ne oR ALE ae
wan
é 7.)
Sgt ea My,
>
= t Ass
hae oes ont) Fain can oi aes _
“ca eben ons OF pee iene mn
PY etre se .
- obs
Foie tbeau on ‘igo bine
Bik MATA oaths eR DAY M, ay
ees ee ae ae meme ie
et je 5 ats
ahi? “ite
LIST OF PLATES.
1878.
Plate Page
I.
ML Pr ASIAUICHIGUEAOSTIEN Pe terete ot es etals chelates Satats Ritter a a o'eb ors 3
Il.
VA eZ OM OL CIAOULCANE rae st lelshd siaid ao toes Mate = ote 37
V. New Shells from Costa Rica .........:.s0.e0000. 72
WAN S Coys argentabi oases eta ee ee Catal oss
Vil. Fig.l. Hypothymis celestis. Fig. 2. Mivornis capitalis 106
Vil. Fig.l. Diceum schistaceum. Fig. 2. Diceum everetti
Fig. 3. Prionochilus olivaceus ......0.+.ss+eee00
UN PD Gsycroraphi, speciosa «3... shins «rein eee oleh 114
XS AMONG SLATE, iota tack is dete eisai oh es 128
XD. Myiarchus)semirufus. . 2c. oie vos tases s+
DY MCI) NGO Ie IRINA opt nao ie SO cig Dera ac 135
USP Criplerus evans /ASCratUs. . outdo ach delo siete snes
XIV. Lophosalea anamallayand........cseere eee eeeees 153
XV. New Opisthobranchiate Mollusca ............2. 55 275
MVE New. Species, of Plustatis . «seis «a se oleisln oly sin hie 293
AVI. . New Species of Penzeidse: sien}. cain stowoioiers » le, oe 298
XVIII. New Shells from Kurrachi, Madagascar, Ecuador, ee) 311
ISTE OGR USA BlOS UN coon EG dowOonbaond sib e se
ONE TIT NOMOCEDNALO . chtaie da iclela aleitte 42) aiusaieys 370
XX Otariaunsinag = Vs, Gy Sy Pins JUV. once ire ers ness 371
KMS Casuariusvedwardsy vicc. ccc ssn sve naec sees s+ es 389
XXII, } Brachiopoda of the ‘Lightning’ and ‘Porcupine’ 393
XXIII. } EXPEGIGION Ts cistct ois ols) efelaniatete n> « ayer ela a's! sfay sleeps) si) 0
oh } Anatomy Of Hy@Nd CrOCUuta.. ccs ee eee sneeeeaees 416
PRON y SPSPEE KOCIE, cfe,staicisieleie'sic'o's'e)o\elo) ol, view slope Sela, tisinals 429
XXVII. Neomorphus radiolosus ......00 sere esse ceecnes | 438
XXVIII. <Aramides calopterus..........0++ aisfelnfae sharers nlite
XXIX. Amblynura kleinschmidti......+.+05 os Pialaeue ere oraksi ane eee
XX
Plate Page
XXX. Stridulating Organ of Palinurus .............0000: 442
MK, Hemiptera of St- Helena 5.5 ucscco x pele oece cis ais olene we 444
RONG LUE Ot A BATLONG otajoissalejaletels, 01 <16xeVoreioiste stele Pano tO ES oC 477
ROMA, | PLO DUS PICHEVENUIIS ele) s1e1610 «1s (altel s o/s. \s)e)e) one felelelenstater } 500
EO OMULV Se PEEL ODS IQ ESLNOE trator stash sole fevenetn ayn /oXs/ oie ieto faealouai el ater eer
Patty | Mow Mautilio sige caus? sc vdspe sae i a
ROKER VN EG eben ehtinme Mawes Mie Gath vaseb dee ae Re CO 611
BS LL, Tachostome Tuff TONS wcn.055 3 sie. 4 sve 9: 4 sno’ ao Paes
RIN, PUGS FORINT 4 Ving totes os cp ee ue iss aomaane ene 632
Slee New due pidapterasartscics otis over cretone ates Wosonbo- 637
PTs SSEVUTUS PUSHLUUS) coe ce oie 0,5 on 310 0ie aie tesa lore theater 656
MOUS “Carpophaga melanochrod. oe os. cio sss o> pv ns alee 670
XLIII. Anatomy of Chlamydophorus truncatus............++ 673
PTV. “Anatomy of yomoschusi.....csls csi > svi ce os ener ones 1682
MEVED New AsiaticHHlespentdce..y en's sjejasd acer ot oitrrae heirs 686
XLVI. New and other Shells from various localities.......... 727
XLVII. Trachea of Rhynchea bengalensis ..........00.e0008 745
ME VIM. New, Shells .24.02002% 0000 A OF esti ncaa eS IEA O a foo
MLK, Peoptera lugubris aise waded weet tee eats ails. 0e 802
i) Shells of the Andaman, Islands <eiitsieetes sates = oe 804
LI.
LII.} Lepidoptera from Upper Tenasserim.......... 00065. 821
LII
LIV. New Shells. .....c.8.004 Rieeetoay oteretahe tate tarereh yes Mees cae
LV. Classification of the Cervid@ ......cccceccecuveees . 883
LVI. Skull of Gazella walleri ........ say eaters SAO ta ae erste 929
EVI... Accigiier-stevensonts «610 ics n00 cde reece «olkees
LVITI.. . Pseudoptyna gurneyt ovo. dese ie ve ve dls ve ewe ee 936
LUX.'. -Chetura gieititcig.. 0 DRE tee VOPR RET IN os cae
LX. Chambered shells of Cephalopods............ Sones 8 lls,
LXI.. Ara, spitiscc.sicies oe aaleletare ects Fong Oia Conic 5 a aoa
LXII. Zamenis elegantissimus ......0.cecccceccccesccees 977
LIST OF WOODCUTS.
1878.
Page
ihinx. of Afomoter leasones (a via oo 0 WWE PRN alee) o yah Bef 101
Interior of the gizzard of Carpophaga latrans .........0. ccc 103
Syrinx of Carpophaga latrans ...... 66. e es SNA Me ME Ae 104
PLC Her \((CAMEN A) CONGEREI a wea sconsn sn starr ss 2, See olale We delobn des reels oe 106
Heads of Oryzoborus crassirostris and O. atrirostris ... 0. cece ene 1386
Three outer primaries of Larus nove-hollandiey jr. .... 2.4545 WSs) LBS
Three outer primaries of Larus nove-hollandia, ad........ cee eee 186
Three outer primaries of Larus nove-hollandia@, old........eee0es 187
Three first primaries of Larus scopulinus, old ....... cece eee ene 188
Three first primaries of Larus hartlaubi, ad...c. sce cere eee eee 189
Three outer primaries of Larus bullert, old vi... cee cece eee ee . 190
Three outer primaries of Larus bullert, nearly ad. .......... Cie! aS)!
Three outer primaries of Larus gelastes, JUV... 1.60.0 eee ees me elo?
Three outer primaries of Larus serranus, Ad. oc. cece e sees eens 196
Three outer primaries of Larus brunneicephalus, ad. ... sss vee aes 197
Three outer primaries of Larus melanocephalus, jX. . 1... e see ees 199
Three outer primaries of Larus ridibundus, J. 0 ccc cece eevee ens 201
Three outer primaries of Larus maculipennis, ld ..... 240.00 eae 202
Three outer primaries of Larus glaucodes, old ..... cee ce eee eeees 208
Three outer primaries of Larus saunderst, ad. v6.0.1. c eects 205
Three outer primaries of Larws
[email protected]. seve even nes 207
Head of Caswarius salvadorti oi... cece eens Papa Paahtes Aes 213
Cephalic shields of Tolypeutes tricinctus, T. conurus, and T. muriet. 223
EyeAIOL, Volypentes’ UiCMectus. »..\.\. i ane Bean eaten celdleleleie oe 8 226
PEFR AITE. OLX CIID UMICUTCLUS 6 cn aos 5'a 0 sett tere iets « weeletsl se «eat 229
LGA Ol B1cer OS ANAANENSMS > o'..c caves -< DRO eee eo 278
Head of Buceros, semigaleatus: . . Mey Nw er wweiee ec eens | OPE so lale 279
lead tot PAyllor hind) COrvind ss). «<n «da cama aa 0 so dye et cletesiereiie © 317
PUGH ALOR PAY HON UNG GALEN: <r uc, x pr-'ain «isinrs hehehe ate ele! Metal edelot 317
PEPTIC eee OCHMUCTUS Suds w eestabete ee Site dala c lols Ghee aleve eee 319
Head of Latrunculus pellucidus, mature male, magnified .,........ 823
Crystallogobius nilssoni, mature Male ... cc rice eee ees 331
Xxil
Page
Snout of mature male of Crystallogobius nilssoni .... 66. e eee 335
Cerscum of Wyctereutes procyontdes iy Hi. ne. lens 6 88 ve on ss 375
Wangsrof Barsiielld mnirausles cs ciaansce «2 +-saxis ainjo in ee eaasee Pepe pele 385
Mra ches vot chyigul a NALONGG —. <0 <, 21a 9s) a: oer 4.05) tear eee 478
VIOLIN COUTZ OO elt i eae ee RE ei ET SO ahs) 7 496
TEC OAMENE) CORVENIL 5 =i. xiv. [sie ietectagnsine Calo adesice ele 497
Cyclophor us (Myxostoma) bathy raphe, er Ce oo ot Sot 498
First primaries of the genus Péelopus:... 0c. cae e clee te eme eens 508
First primary and foot of Ptilopus togaster.... 0.66. c cee eee aes 510
First primary and foot of Ptilopus fasciatus ........ 0c eee evens 510
First primary and foot of Ptilopus geelvinckianus ..........0000++ 511
First primary and foot of Ptilopus coronulatus 0.00.0. cec eevee 511
Wing of Drepanoptila holosericea 62s. ccc cue ne neve ee renee 513
UCR A ETT ia S ssicce aioe ete RN Baad chante eA Gaak hae pS ATES 610
Lower end of the trachea of Tantalus loculator $1.6... 20s eee 626
Windpipe of Vanella casennensts. co. cic s.e oe ced coe ss oneness 628
Lower larynx of Megalocephalon maleo ........0 cece een eees 630
Phiyllopter a: segonoudes. 00.0, AeA OG OTS AEE 649
Stomach of Plotus levawlants ce sc. is ceaials wine Sake oe ee 680
Head of Gazella granti (side View)...........seceeessevccueces 724
Head of Gazella granti (front view) .....0..edecceveseeeeeeees 725
Trachea of Rhynchaa capensts; SD vewsis sc'iivda'ds eaale iis Sajs's @ dels 748
Podobranchia of the antepenultimate thoracic limb of Astacus fluvia-
BES Ja, oa sina. 5 xc0' © Oia BU ed Rr SPRITE Ie 757
Podobranchiz of the first maxillipede of Astacus fluviatilis, Asta-
coides madagascariensis, and Parastacus brasiliensis ............ 759
Branchiostegite of Astacus fluviatilis 60.0... ccc cece eee eens 760
Branchiostegite of Astacopsis franklinti ....0 60.0 c cece cece ees 760
Podobranchia of the antepenultimate thoracic limb of Astacopsis
PP ONCE aos ¢.o, 0, 9 « on: waite oT SHES, SARs BS HURT CE 765
Podobranchia of the antepenultimate thoracic limb of Cheraps. . 767
Podobranchia of the antepenultimate thoracic limb of Aatosvatin
PUOCROG ASCOT LCDS i. «0 5 pew SORT ENY DIKDIae «elie DEORURI BE los 772
Antler ot: Ceruellies nung, 2 sensi tee ke SE 898
Antler of Cervus (Rusa) aristotelts .......s ccc cee ee ccccececues 900
Antler of Cervus (Rusa) hippelaphus ...... Se ita at See py te 903
Antler of Cervus (Rucervus) duvauceli .... ccc ccc cece eee ee eee 905
Antler of Cervus (Elaphurus) davidianus ..... 06060 cc cece eens 906
Antler of Cervus (Axis) axis ............ Beinctajein © ghstoe gestae 907
Antler of Cervus ( Pseudarts)! slit Fo... RRR ae 908
Antler of Cervus (Cervus) elaphus. . 10.0 cses cs vewiuw sence eres 910
Antler of; Ceres (Dart Wada oo. 5c Vals «SOO INS DTI 914
Antler of Cervus (Dama) mesopotamicus .......2000 0.0 cece ee 914
Wutler of Alves Wate re tern tn oc’ ees vo BNO NS. 915
Antler of Ca ‘apr FONE ME ces tris sais ae RE NAS ee 917
Antler of Cariacus (Cariacus) virginianus ....... 00000000 eee 918
Antler of Cartacus (Cartacus) macrotis ......0 ccc cece eee eee 921
Antler of Cartacus (Blastocerus) paludosus oo. .6 60. cee ee ec ee 922
XXxili
Antler of Cariacus (Blastocerus) campestris .....000 ccc ccc eens
Antler of Cartacus (Furcifer) antisiensis oo. 0.0... 0 0 ccc cues
wuitler of Cariaeus (Compsus) rufees os... cscs vce sc cunsevqevencs
MADE OL LENG CL CAP ORAUB . oe vii veges vane cule etaeneans
ES PLIES see ie CRS pect aaah ek atc sak esas ¢
Palatal aspect of the truncated vomer of Ramphastor ariel........
etree FQMICRE MAIO. oe oe ks eas wo vis anise eelcne fs
Section of chambered part of shell of Vermetus gigas ............
Section of chambered part of shell of Spondylus varius... 0.00.2...
Section of part of shell of Nautilus striatus .....6... 6.00 cee eee
Section of part of shell of Spirula australis ...... 00.0.0. c eee
Protoconch and protosiphon, with following chambers, of Spirula
MAOH MNS Cast hehe G Sreereen atte nS fone NST oe kine eh axon e
ERRATA.
Page 666, line 7, for rufo-niger read brunneo-niger.
Page 667, line 2, for rufo-niger read brunneo-niger.
Page 670, line 18, for are not yet quite adult read are quite adult.
Page 723, line 2 from bottom, for p. 535 read p. 527.
Page 929, line 1, for Western read Eastern,
PROCEEDINGS
OF THE
SCIENTIFIC MEETINGS
OF THE
ZOOLOGICAL SOCIETY
OF LONDON
1878.
(PLATES)
PRINTED FOR THE SOCIETY,
AND SOLD AT THEIR HOUSE IN HANOVER SQUARE,
LONDON:
MESSRS. LONGMANS, GREEN, READER, AND DYER,
PATERNOSTER ROW.
: Bons PAAR ine 3
E TAA
1878.
Plate Page
I.
De ABinte TRO cso ite afer Haein s ela wR ace 3
Il. ;
LY.) Zonpirichis OUlcane..0'36 55-85 si iain bs Qa sess 32 2 37
N. ~ New Shella fram Costa: Rieai\ ‘aja. ia eet «ol 72
Niles Conte: ang ental auvedintoiacts Sates st xv Sore BO teers oe
VI. Fig.l. Hypothymis celestis. Vig. 2. Mixornis capitalis 106
VIII. Fig. 1. Diceum schistaceum. Fig. 2 . Diceum everetti
Fig. 3. Prionochilus olivaceus ........0+ 0000 eee
EX. Dasyerotaphacspectosey i. cs «<3 Se aeipawaesniseo se, o- 114
Xs Amadinasharpee \ cidc detain iexem Semaine pattems! stor sie 128
MN) Matar clus Semin uUfuss »| 2) <iagebeusie os ols ove Wake oh si al tecwreu os
IT Capito: SCC gia seca casaisi oss 6 eakeeions 135
MU, Crypiurus Transp aseiatus soci ccnp 00 vl ois nein cia ble oe
XIV. Lophosalea anamallayand...... 0.2. e eee eevee ees 153
XV. New Opisthobranchiate Mollusca .........0+.+2++ 275
VIS | New Species Of; Plustozts. =. sc :cc'e > 6 obs cjeleusi mela * lee 293
XVII. New Species of Penzide ............:.00sceeeees 298
XVIII. New Shells from Kurrachi, Madagascar, Ecuador, ‘abhi 311
PALMER ALT May gratesel ciietmis aiigerege 317i stelelahoe (erage) ar stevases afer os
EUG, = Pirined POMOCEDRGALG — a sce’) va eis: cin oA eysttreb aks sete es ed oie 370
RON COLAO UPSUN Ms Vs Ss Dae? se Sa) |UVueaiatsfaiavet st gs cictetels 371
RON MC ASIATIUS COM OTOSI «chars sists «s) stouniensiersyefele-| slecpela leks‘ 389
ae ee of the ‘Lightning’ and ‘Porcupine | 393
PRONOUN! CXS ONS yieyciatesstee Shs, ne Fsveve Wie eeyeceminnayeeaeierl 81a /-a)s ;
AY. } Anatomy OL Hy CNM CHOCULA = cyelelavislere s «os ess) 0's + = 416
PRONG V(MCR AE SEED KOHL. 4) steteysicie (or sicicenisrofel e.g «)e dhs oP ehsaste, este alt, asciote 429
DOV LL pNCOIOTPIUS NUMOIOSIE oe ca. eee es ve alin eves 458
Reeve Aramedes colopterus. . oo. ceige ce x00 eseey ohivesiae
XXIX. Amblynura kleinschmidti .......00ccessvevecseees 440
LIST OF PLATES.
Plate
XXX,
XXXII.
XXXII.
XXXII.
XXXIV.
XXXY.
XXXVI.
XXXVII.
XXXVIII.
XXXIX.
XL.
XLI.
XLII.
XLII.
XLIV.
XLV.
XLVI.
XLVII.
XLVIII.
XLIx.
Page
Stridulating Organ of Palinurus ......4. SSacumn foc 442
Hemiptera of St. Helena ..... BiieoaG oGanos haobe 444
ENE GUL MEAELONG. c\etorsioielalere(ave oltre ole) stolarel ster ste seioleliateleae 477
Piilopus pictwwentris —.. 0. vecccussssnsenvrsccesncs
Ptilopus Gestr0t 2. v0 ene vinyew sents a ceeverecncrns } ae
| Wey Wiantidlee re cu tiaittes cetera esas viadeachonale ualerse/otehehetoeets 580
TEER Sot OSS Sn osiban pmo aOnbSoo UES boo ou. }u
Trichostoma rufifrons .... That ei tecet ais onieaat
PAP US TOMBTE 0% sexys sc, pials «tsa ale ele bi ee oa 632
New Lepidopteriice wie cesis'ee.cn0 tie as stein) efelsceeto tiene 637
AD Re DG DE 3 hoieeo Molo Grace nomerapo-conn aoc” 656
Carpophaga melanochrod... i... se ese essen neeees 670
Anatomy of Chlamydophorus truncatus..........0+4+ 673
Amatomy: Of ea) OMOSCHIUS si. voile s aicis\e “hee cee sete 682
New Asiatic Hespertde =... 0+ 002 eos viele eens 686
New and other Shells from various localities.......... 727
Trachea of Rhynchea bengalensis ......0600ceeeeeee 745
New-Shells :caaewdascorsas teres cores «eee cele 795
Peoptera lugubris ........+... ar gene era c rotate eis avers 802
Shells of the Andaman Islands ..........-.0000 000: 804
Lepidoptera from Upper Tenasserim.......... 24+... 821
iNew tSbiellsisviccrenes sys vaicietese tele o's = foreplay Vevers sl etal 859
Classification of the Cervide...... cbinodemRAetnatan a+ 883
Skull of Gazella walleri ........ Riertogeies ee otne terete 929
ACCHBILEN SLCREN SOIT Neve sorte sola aie mate ore tha tersie Taare
PISCUMOPLO TD TH UTIEGE Tatareysteleis cafe 2 celica ie nine hielo) tale 936
CU HAGTAOS TEED oh 331. Ov anid SOR OIA Oo MICE. OING NC vio
Chambered shells of Cephalopods..............+05 . 955
TAGEN GS TE kOe ODIO AED DU DU COTO OA Ce MIONO ATI Od an 976
ZGMENTS CLEGUNTESSIMUS we vie ee oe! «1 ws eos es 60 pte 977
LIST OF THE PUBLICATIONS
OF THE
ZOOLOGICAL SOCIETY OF LONDON.
Tue scientific publications of the Zoological Society are of two
kinds — “ Proceedings,” published in an octavo form, and
“Transactions,” in quarto.
According to the present arrangements, the “ Proceedings”
contain not only notices of all business transacted at the scien-
tific meetings, but also all the papers read at such meetings
and recommended to be published by the Committee of Pub-
lication. From fifty to seventy coloured plates and engravings
are attached to each annual volume of the “ Proceedings,” to
illustrate the new or otherwise remarkable species of animals
described in them, Amongst such illustrations, figures of the
new or rare species acquired in a living state for the Society’s
Gardens are often given.
The “Proceedings” for each year are issued in four parts, on the
first of the months of June, August, October, and April, the part
published in April completing the volume for the preceding
year. They may be obtained with black or coloured illustrations.
The “Transactions” contain such of the more important
communications made to the scientific meetings of the Society
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MADE FOR
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SECRETARY TO THE SOCIETY.
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PROCEEDINGS
OF THE
SCIENTIFIC MEETINGS
OF THE
ZOOLOGICAL SOCIETY OF LONDON.
January 15, 1878.
Robert Hudson, Esq., F.R.S., in the Chair.
The following report on the additions to the Society’s Menagerie
during the month of December 1877 was read by the Secretary :—
The total number of registered additions to the Society’s Mena-
gerie during the month of December 1877 was 103, of which 17 were
by birth, 28 by presentation, 24 by purchase, 21 by exchange, and
13 were received on deposit. The total number of departures during
the same period, by death and removals, was 101.
The most noticeable additions during the month were :—
1. A Family of Gelada Baboons (Cynocephalus gelada), deposited
by Mr. C. Hagenbeck, December 7th, consisting of an adult male,
three young males, and four females of this singular and little-known
Monkey. A single example of it was once before exhibited for a
short time in the Society’s Menagerie ; but this is the first occasion
on which we have had an opportunity of watching their curious
habits and observing their grass-eating propensities.
2. A pair of Musk-deer (Moschus moschiferus), presented by Sir
Richard Pollock, K.C.S.1., H.M.Z.S., Commissioner at Peshawur,
N.W.P., and arrived December 15th. This is believed to be the
first pair of Musk-deer that have reached Europe alive. They have
been placed temporarily in one of the outer cages of the New Lion-
house, and seem likely to do well.
Proc. Zoou. Soc.—1878, No. I. l
2 ON THE BIRDS OF THE N.W. PROVINCES. (Jan. 15,
The Secretary read the following letter from Mr. A. Anderson,
F.Z.S., containing a correction of his last paper on the Raptorial
Birds of the N.W. Provinces :—“I find that Mr. Hume is right in
his conjecture ' that the bird figured at pl. xxii. P. Z. 8. for 1876,
as Falco babylonicus, is an abnormally coloured F. barbarus.
«‘ Mr. Gurney, who has kindly (at my request) reexamined the bird,
and compared it with specimens at the British Museum, has autho-
rized my publishing the following memorandum which he has drawn
up on the subject.
*¢ Memorandum.
«At the request of Mr. Anderson I inspected the Falcon, figured
in the Society’s ‘ Proceedings’ for 1876, on pl. xxiil., as a specimen of
Falco babylonicus, before that figure was drawn. The specimen
struck me as smaller than any 7. babylonicus I had previously seen,
and as having rufous edgings to a larger proportion of the feathers of
the mantle; but it certainly did not occur to me that it was any thing
else than a small male of F. babylonicus, with the male of which
species I was not, however, well acquainted, the specimens of that
Falcon in the Norwich Museum being apparently, by their size, all
females.
«At p. 140 of ‘Stray Feathers’ for 1877 this Falcon is referred
to; and the Editor suggests that it is too small for F. babylonicus,
and ‘ might perhaps be /. barbarus.’
“In consequence of this remark, I recently took the skin, at Mr.
Anderson’s request, to the British Museum for comparison, and
found that it closely agrees with a Falcon from Nubia in that collec-
tion, both in size and in coloration; this Nubian Falcon has been
identified by Mr. Sharpe with F. darbarus, and is specimen a, in
the enumeration given at p. 387 of vol. i. of his British- Museum
Catalogue.
“*T believe that Mr. Sharpe’s identification of this specimen is cor-
rect, and that both it and Mr. Anderson’s Etawah Falcon are abnor-
mally coloured specimens of /. barbarus.
“J. H. Gurney.
“6th December, 1877.
“In expressing my regret that the mistake in question should
have occurred, I may remark that I believe the present is the first
recorded occurrence of an adult /. daréarus within the limits to
which I have restricted my researches, whereas the capture of F,
babylonicus has been of comparative frequency ; the addition there-
fore of the former (f. barbarus) to my plains Catalogue is of
greater importance than if it had been the latter (7. babylonicus).”
The following papers were read :—
1 Cf. ‘Str. Feath.’ for 1877, p. 140.
Hanhart imp.
ASIATIC LITHOSHDAS .
.
.
< % a
Aa 3
\
’
3
“a ge!
re F
n.
J '
a
R. Mintern, del. et lith.
ASIATIC LITHOSIIDE.
R. Mintern. del, et lith.
ASIATIC LITHOSIIDE.
1878. ] MR. F. MOORE ON THE LITHOSIID®. 3
1. A Revision of certain Genera of European and Asiatic
Lithosiidz, with characters of new Genera and Species.
By F. Moors, F.Z.S., Assistant Curator, India Museum,
London.
[Received October 29, 1877.]
(Plates I.-III.)
Family Lirnosirp2.
Subfamily Hypsinz.
All the genera in this subfamily possess a peculiar stridula-
tory apparatus in both sexes. This is distinctly visible on the
upperside of the posterior margin of the fore wing, and as viewed
from the underside of the wing appears as a short oval nacreous
cavity, situated between the submedian vein and the extreme margin,
along the upper edge of which projects a raised longitudinally oblique
fold, which is smooth in some, and transyersely scabrous in others ;
and on the upperside of the hind wing is a patch of scabrous scales,
or a short transverse outwardly-curved raised scabrous bar, which,
by the motion of this wing, evidently plays upon the raised fold over
the edge of the cavity, and produces a stridulating sound.
Hypsa, Hiibner, Verz. bek. Schm. p. 172.
HyYpsa CANARAICA, 0. sp.
Female. Upperside—fore wing cinereous brown ; veins white ; base
of wing ochreous and spotted with black ; a whitish clavate streak
from lower end of the cell; cilia cinereous: hind wing white, with
broad black marginal band crossed by white veins ; four black discal
spots and a short anal streak; cilia whitish. Thorax ochreous,
with small black spots; abdomen brighter ochreous, with a dorsal
and lateral row of black spots. Underside cinereous brown, basal
half of fore wing white, with two broad black spots within, and a third
below the cell ; hind wing as above, with the marginal band extend-
ing along the costa. Near to H. heliconia.
Expanse 2 inches.
Canara (Ward). In coll. F. Moore.
HyYPpsA SERICEA, D. sp.
Male. Upperside pale silky ochreous-yellow ; cilia whitish: fore
wing with the base brighter and dotted with black; an indistinct
whitish spot from lower end of the cell: hind wing with costal border
whitish ; a broad cinereous brown marginal band and two confluent
longitudinal discal streaks. Body pale ochreous-yellow; abdomen
brighter with dorsal and lateral row of black spots; palpi with
black terminal point and two lateral spots ; legs grey, streaked with
black. Underside cinereous brown; fore wing with yellow hind
margin and basal costal streak ; hind wing with marginal band and
1*
4 MR. F. MOORE ON THE LITHOSIIDZ. [Jan. 15,
discal streaks almost confluent, being separated only by a narrow
yellow division.
Expanse 12 inch.
Canara (Ward). In coll. F. Moore.
Near to H. heliconia.
Damattis, Hitibner.
Damalis, Hiibner, Verz. bek. Sch. p. 172.
DAMALIS CONCANA, 0. sp.
Near to H. alciphron, Cram. pl. 133. f. E.
Upperside ochreous ; cilia white: fore wing at base bright ochre-
ous and dotted with black; a small whitish streak from lower end
of the cell: hind wing with black marginal band crossed by pale
veins, and two irregular longitudinal discal streaks. Thorax with
black dots; abdomen with dorsal and jateral row of black spots ;
legs whitish, streaked with black. Underside—fore wing pale
ochreous-brown; hind margin yellow; cell white, bordered with
black and enclosing a black spot : hind wing yellow, with paler black
marginal band, and streaks as above.
Expanse 22 inches.
Canara (Ward). In coll. F. Moore.
Dieama, Moore.
Digama, Moore, Catal. Lep. E.I. C. ii. p. 298 (1858).
Fore wing—first branch of subcostal long, free ; second at a con-
siderable distance before end of the cell, trifureate; fifth from near
end of the cell, curved upward and looped to third at one third its
length ; radial from end of the cell; discocellulars nearly erect ;
median vein four-branched, upper from end of the cell, second and
third from angles near its end. Hind wing with two subcostal
branches beyond the cell; discocellulars curved outward, oblique ;
four median branches, three upper from end of the cell. Below the
submedian vein on the underside of the fore wing, is a short longi-
tudinal cavity, along the upper edge of which projects a raised trans-
versely scabrous narrow bar ; and on the upperside of the hind wing
is a short transverse outwardly curved raised scabrous bar, so placed
that, by the motions of this wing, it evidently plays across the longi-
tudinal bar on the fore wing and produces a stridulatory sound.
DiGAMA HEARSEYANA.
D. hearseyana (part), Moore, Catal. Lep. E.I. C. ii. p. 298. pl. 7a.
f. 3a, 2 (1858).
Male and female. Upperside—fore wing greyish brown, with four
basal transverse series of black spots, and an intermediate spot in
the cell, with paler interspaces along the veins and wavy bands across
the disk: hind wing and abdomen ochreous yellow, the latter with
dorsal row of black spots. Thorax, head, and palpi grey-brown ;
front and middle of thorax black-spotted ; joints of palpi with
terminal black band; legs black-banded above, pale ochreous
1878. ] MR. F. MOORE ON THE LITHOSIID. 5
beneath. Underside—fore wing uniform pale grey-brown ; hind
wing with a small black discocellular spot at upper end of cell.
Expanse, d 1,2;, 2 13 inch.
N.W. India (Allahabad, Manpuri, Deyra, Simla).
In the Catalogue Lep. East-India Company, above quoted, is
described and figured (fig. 3) a male insect from Ceylon, which I
then considered to pertain to this species. The Ceylon specimens,
however, are very distinct ; and the species has been named by Felder
as D. insulana.
DIGAMA INSULANA.
Digama insulana, Feld. Verh. zool.-bot. Gesch. 1868, p. 285.
D. hearseyana (part), Moore, Catal. Lep. E.I. C. ii. p. 298, pl. 7.
f. 3, 2 (1858).
Male and female. Upperside—fore wing pale grey-brown, basal
spots prominent : hind wing ochreous yellow, male with a grey-brown
marginal streak, and female with a short marginal spot, situated near
the middle. Head and thorax pale grey-brown, black-spotted ; ab-
domen ochreous, with dorsal black spots; antennz brown in male ;
legs black-banded above ; palpi with terminal black bands. Under-
side paler ; both wings with a black spot at end of the cell.
Expanse 14 inch.
Ceylon. In coll. India Museum.
DIGAMA SIMILIS, Nn. sp.
Male and female. Allied to D. hearseyana, but of smaller size ;
fore wing similarly marked ; hind wing in the male with a dusky
brown marginal streak from near anal angle, and another near apex,
the latter only present in the female.
Expanse 3 1, 9 154, inch.
N.W. Himalaya (Simla). In coll. F. Moore.
DIGAMA FIGURATA, 0. sp.
Male. Upperside—fore wing greyish white, with very prominent
brown basal spots, two median, transverse, irregular, sinuous, partly
confluent bands, and a partly confluent marginal band; a large black
spot in middle and a lunule at end of the cell : hind wing pale ochreous
yellow, with prominent black marginal marks. Thorax and head
white, spotted with black; antennze brown; palpi white, with
broad terminal black bands; abdomen ochreous-yellow, with dorsal
and lateral black spots ; legs white, with black bandsabove. Under-
side—forewing grey-brown, spots in cell visible; hind wing with
black spot at end of the cell, and on middle of outer margin.
Expanse 1,3, inch.
Burmah. In coll. F. Moore.
Caurenta, Moore.
Calpenia, Moore, P. Z. 8. 1872, p. 571.
CALPENIA KHASIANA, 0. sp.
Female. Upperside—fore wing brownish fawn-colour vinous-tinged,
6 MR. F. MOORE ON THE LITHOSIID. [Jan. 15,
paler along the veins; a broad pale yellow band extending lon-
gitudinally from base below the cell and upward to near the costa,
being crossed by the veins; some yellow spots on the costa; a small
spot and streak at base of cell, and two outer submarginal rows of
small dentate spots of chrome-yellow: hind wing chrome-yellow,
with four transverse rows of vinous brownish black spots and a rayed
basal streak, the spots broadly lunate, irregular in size, the outer
row being marginal and the smallest. Antennze, front of head, and
palpi black; thorax dark greyish brown, longitudinally streaked
with chrome-yellow; abdomen chrome-yellow, with dorsal and
lateral row of spots. Underside same as above; legs dark grey.
Expanse 34 inches.
Khasia hills (Atkinson). Coll. Dr. Staudinger.
Neocuera, Hibner.
Neochera, Hiibner, Verz. bek. Schmett. p. 173.
NEOCHERA HELICONIOIDES, 0. sp.
Male. Upperside—fore wing purplish cinerous ; all the veins nar-
rowly lined with white; base of wing black-spotted, and with a
subbasal orange-yellow streak with zigzag borders: hind wing grey-
black, washed with purplish blue ; three grey streaks from the base
and a prominent submarginal grey narrow undulating band; cilia
white. Body orange yellow; a black spot on each tegula, one on
hind part of thorax, and a dorsal and lateral row of spots on abdomen.
Underside grey-black ; both wings with white external veins and
undulating submarginal band; the basal area broadly white and
with a large blackish cell-spot; front of head with a central black
spot ; palpi and legs white, streaked with black.
Expanse 22 inches.
Luzon, Philippines. In Coll. Dr. Staudinger.
Similar in pattern to Hypsa heliconia.
Subfamily Lirnosiinz.
The genera are here arranged in the following sectional groups,
viz. :—
A. Fore wing with subcostal vein five-branched, median vein four-
branched :
Eligma, Macrobrochis, Tripura, Paraona, Atolmis, Sidyma, Chu-
ringa, Vamuna, Mahavira, Korawa, Hesudra, Ghoria, Chryseglia,
Conistes, Lithosia.
B. Fore wing with subcostal vein five-branched, median vein three-
branched :
Simareea, Tarika, Brunia, Gandhura, Collita, Katha, Eilema,
Manulea, Systropha, Chrysurabdia, Capissa, Dolyoma, Mithuna,
Cossa, Ranghana, Tegulata.
Ba. Hind wing with a recumbent plumose lappet :
Nishada.
“I
1878. ] MR. F. MOORE ON THE LITHOSIID.
Bes. Fore wing with a recumbent plumose lappet :
Teulisna, Macotasa, Zadadra, Prabhasa, Gampola.
C. Fore wing with subcostal vein fowr-branched, median vein
three-branched :—
Pelosia.
D. Fore wing with a plumose, lappet-covered, sac-like cavity on
the costal border ; five subcostal and three median branches :
Doliche, Bizone, Cyana.
E. Aberrant group: fore wing with five subcostal and four median
branches :
Baroa, Agrisius, Lyclene, Barsine, Miltochrista, Setinochroa,
Setina, Nudaria, Gimene.
Group A. Fore wing with subcostal vein rive-branched, median
vein FouR-branched.
Exrema, Hiibner.
Eligma, Hiibn. Verz. bek. Schmett. p. 164 (1816).
Panglima, Moore, Catal. Lep. E.1.C. ii. p. 297 (1858).
Surina, Walk. Tr. Nat.-Hist. Soc. Glasgow, i. p. 9 (1869).
Fore wing—first subcostal branch free; second from near end of
the cell, trifureate ; fifth from end of the cell, bent up and with a short
spur looped to third close to its base ; radial from end of the cell;
discocellulars bent, upper longest ; median vein four-branched, angled
at end of the cell, two upper branches from the front and third from
angle below the end, fourth from beyond half towards the base.
Hind wing—two subcostal branches, and three upper median branches
from end of the cell.
Type, Z. narcissus (P. narcissus, Cram. Lep. Exot. i. pl. 73. f.
E, F).
China, Java, Penang, S. India.
Macrosrocuis, H.-S.
Macrobrochis, H.-Scheeff. Lep. Exot. Sp. Nov. pl. 72. fig. 531
(1856).
Fore wing—first subcostal branch free, curving upwards at its
base towards costal vein ; second from near end of the cell, trifur-
cate ; fifth from end of the cell, bent upward and with a short spur
looped to third near its base; radial starting from fifth subcostal
branch beyond the cell ; upper and lower discocellulars bent inward
at very oblique angles ; median vein bent upward near end of cell,
four-branched, two upper from point at end of the cell, third from
angle below its end. Hind wing—two subcostal and two upper
median branches from beyond end of the cell, third median below
its end.
Type, M. gigas (Lithosia gigas, Walk. Cat. Lep. Het. B. M. ii.
p- 494, 1854; WM. interstitialis, H.-S. 1. c. f. 531).
8 MR. F. MOORE ON THE LITHOSIIDE. (Jan. 15,
Macrosrocuis LEUCOSPILOTA, Nl. Sp.
Male and female. Differs from MV. gigas (Lithosia gigas, Walk.
Catal. Lep. Het. B. M. ii. p. 494) in the fore wing having more
elongated white spots, and in the hind wing having the marginal
band much narrower and traversed by a series of elongated spots on
both upper and underside.
Expanse 2? inches.
Cherra Punji, Assam. In coll. India Museum, and F. Moore.
MacrosROCHIS NIGRESCENS, 0. sp.
Male. Allied to M. gigas. Differs in the spots on the fore wing
being much less distinct ; the basal and discal spots partly covered
with black scales and totally obsolete on the underside ; on the hind
wing the base is only slightly dusky-white on both sides,
Expanse 2? inch.
N.E. Bengal. In coll. F. Moore. .
Tripura, Moore.
Tripura, Moore, Catal. Lep. E.I. C. ii. p. 298 (1858).
Fore wing—first subcostal branch free ; second from before end of
the cell, trifurcate ; fifth from end of the cell, bent upward and
touching third near its base; radial from end of the cell; disco-
cellulars bent in the middle ; median vein four-branched, two upper
from angles at end of the cell, third from angle before its end. Hind
wing—two subcostal branches beyond cell, three upper median
branches at its end.
Type, 7’. prasena, Moore, /. c. p. 299, pl. vil. a. fig. 6 (1858).
Himalayas.
5 PARAONA, 0. g-
Fore wing long, narrow, costa slightly arched, apex acute, exterior
margin oblique, posterior margin slightly convex towards the base ;
subcostal vein five-branched ; first branch oblique, free; second arising
before end of the cell; third starting from above base of second, tri-
furcate towards its end ; radial from end of the cell, curved upward
and joined to third subcostal branch, at one fourth its length beyond
the cell; discocellulars slightly bent in the middle; median four-
branched, two upper at end and third before end of the cell. Hind
wing long, somewhat narrow, apex slightly produced, exterior margin
and angle convex; two subcostal branches one third beyond the
cell ; median four-branched, two upper beyond and third from end
of the cell. Body stout. Antenne in the male serrate, the teeth
finely pectinate, setose in the female. Palpi small, ascending, pilose
beneath, apical joint very minute. Legs stout, short ; femora slightly
pilose beneath ; tibize thickened, middle and hind tibize spurred.
PARAONA SPLENDENS. (Plate I. fig. 1.)
Crambomorpha splendens, Butler, Trans. Ent. Soc. Lond. 1877,
p. 357.
Bombay (Dr. Leith). In coll. British Museum, and F. Moore.
1878. | MR. F. MOORE ON THE LITHOSIID. 9
Atrotnis, Hibner.
Atolmis, Hiibn. Verz. bek. Schmett. p. 164 (1816).
Gnophria, Steph. Brit. Lep. ii. p. 98 (1829).
Fore wing—first subcostal branch free ; second from before end of
the cell, trifurcate ; fifth from end of the cell, bent upward and touch-
ing third at half its length ; radial from end of the cell; discocellu-
lars bent inward ; median four-branched, two upper from end of the
cell, third from angle below its end. Hind wing—two subcostal
branches beyond the cell; three upper median branches from its
end.
Type, A. rubricollis (N. rubricollis, Linn).
Europe.
Sipyma, Walker.
Sidyma, Walk. Catal. Lep. Het. B. M. vii. p. 1686 (1856).
Fore wing—first subcostal branch long, free ; second from near end
of the cell, trifurcate ; fifth from end of the cell and touching third
near its base ; radial from fifth subcostal branch at some distance
beyond the cell ; discocellulars bent ; median vein angled near end,
four-branched, two upper from end, third from angle below end of
the cell: hind wing— two subcostal and three upper median branches
from end of the cell.
Type, S. albifinis, Walk. Catal. Lep. Het. B. M. vii. p. 1686.
Masuri, N.W. Himalayas (Capt. Lang).
SIDYMA APICALIS, n. sp. (Plate I. fig. 2.)
Male. Upperside purplish-black ; fore wing with a narrow tri-
angular white apical patch. Collar, thorax beneath, and anal tuft
orange-red. Underside as above.
Expanse 13 inch. .
Darjiling (Atkinson). In coll. Dr. Staudinger.
Smaller than S. aldifinis, which is known only from Masuri. Dis-
tinguished by having the apical white patch on the fore wing only.
CHURINGA, 0. g.
Male and female. Wings ample, broad, somewhat short in male.
Fore wing with the costa slightly arched towards end, apex pointed,
exterior margin oblique, posterior angle rounded; first subcostal
branch very oblique, free; second from near end of the cell, trifur-
cate ; fifth from end of the cell, with a loop-spur to the base of the
third ; radial starting from below the fifth subcostal beyond end of
the cell; discocellulars of equal length, bent inward at the middle ;
median vein four-branched, two upper from end of cell, next at some
distance before and lower from half length of the cell. Hind wing
convex at the apex; exterior margin rounded, abdominal margin
short ; two subcostal branches at one fourth beyond end of the cell ;
median four-branched, two upper from end, next before the end, and
lower from half length of the cell. Body large; thorax with long
pilose tegule. Antennze in male broadly pectinate, setose in female.
Palpi slender, ascending, basal joint pilose beneath, second and third
10 MR. F. MOORE ON THE LITHOSIID. [Jan. 15,
joints very long, tip blunt. Legs long, femora slightly pilose
beneath.
Type, C. rufifrons. Will also include Lithosia beema, Moore,
P.Z. 8. 1865, p. 798.
CHURINGA RUFIFRONS, n. sp. (Plate I. fig. 12.)
Male and female. Upperside—fore wing pale purplish ochreous-
brown, costal and posterior margins narrowly edged with ochreous
yellow; hind wing and abdomen pale ochreous-yellow. Thorax
brown ; tegule edged with yellow; head, collar, and palpi reddish
ochreous ; thorax beneath and legs bright ochreous; tip of palpi,
antennze, fore legs above, and tarsi, brown. Underside paler.
Expanse, 317%, 2 23% inches.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
VAMUNA, 0. g.
Fore wing long, narrow ; costa slightly arched beyond the middle ;
apex acute ; exterior margin short and slightly truncate below the
apex, oblique hindward, the angle slightly convex ; posterior margin
convex towards the base in male. Hind wing broad; apex slightly
produced ; exterior margin convex in the middle. Veins similar to
those in Churinga (C. rufifrons), excepting that in the fore wing the
lowest or fifth branch of the subcostal is emitted from upper angle
of the cell, the median branches nearer together at their base, and
the lowest nearer the end of the cell. Body moderately stout. An-
tennze in both sexes setose. Palpi moderately long, ascending, pro-
jecting beyond the head, second joiut long, third jomt very short.
Legs long, naked; middle and hind spurred.
Type, V. remelana (Lithosia remelana, Moore, P. Z.S8. 1865, p.
798).
Darjiling.
VAMUNA MACULATA, uu. sp. (Plate I. fig. 5.)
Male. Upperside—fore wing greyish ochreous; hind wing pale
dull ochreous at base and whitish externally, with three marginal
black spots, the two upper larger, the lower small and at some dis-
tance from anal angle. Body ochreous. Underside dull ochreous
basally, whitish externally ; fore wing with a brownish patch at the
apex, a black longitudinal basal streak, and a broad large transverse
subapical patch ; hind wing with only the lower and middle marginal
black spots. Female much paler, the apex of fore wing slightly
brownish-ochreous; hind wing above and beneath with smaller
median and lower black spots ; the fore wing with the basal black
streak less defined and the subapical spot small.
Expanse, 313, ? 2 inches.
Darjiling (Atdinson). Incoll. Dr. Staudinger and F. Moore.
VAMUNA BIPARS, n. sp. (Plate I. fig. 11.)
Male. White, costal edge of the base and apical half of fore wing
fuliginous black ; hind wing with a fuliginous-black submarginal band,
1878. | MR. F. MOORE ON THE LITHOSIIDE. 11
the middle portion of which is very broad and blackest. Antenne,
front of head, and tip of palpi black ; fore and middle legs above, a
terminal spot on hind tibize and their tarsi, black ; tarsal joints with
a white band; base of palpi, thorax and abdomen, legs beneath,
and anal tuft ochreous. Underside duller white ; the black band on
fore wing confined to the disk, the apical border being pale brownish-
ochreous ; upper portion of band on hind wing obsolete.
Expanse 2 inches.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
MABAVIRA, 2. g.
Male. Fore wing long, narrow, costa arched, apex acute, exterior
margin oblique, posterior margin recurved; subcostal vein five-
branched, first and second arising before and near end of the cell,
third trifurcate, the two lower branches at equal distances from end
of the cell; discocellulars long, straight, upper obliquely inward,
lower obliquely outward ; radial from upper end of the cell ; median
vein four-branched, the two upper branches from lower end of the
cell, third at some distance before its end; submedian vein curving
towards hind margin: hind wing longer than broad, exterior mar-
gin convex ; subcostal vein two-branched beyond end of cell ; dis-
cocellulars as in fore wing; median vein four-branched, the two
upper branches from end of cell and contiguous at their base, third
immediately before end of the cell. Body slender. Antenne ser-
rate, with long and delicate pectinations. Legs slender, femora
slightly pilose beneath ; middle and hind tibiee spurred. Palpi slender,
porrect, basal joint pilose, apex pointed.
MaHAVIRA FLAVICOLLIS, n. sp. (Plate I. fig. 3.)
Male. White, costa slightly edged with brown at the base; collar
yellow ; antennz yellow; fore femora with a blackish streak on the
inner side.
Expanse 12 inch.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
Korawa, 0. g.
Male and female. Fore wing long, narrow ; costa slightly arched ;
apex somewhat acute, exterior margin very oblique; subcostal vein
five-branched, first branch ascending and touching the costal but
free at its end, second starting before eud of the cell, third at end of
the cell, trifurcate, lowest branch at one third beyond end of the
cell; upper discocellular angled outward at its middle, lower oblique ;
radial from angle of upper discocellular ; median vein four-branched,
the two upper from lower end of the cell; submedian slightly re-
curved. Hind wing moderately short, apex slightly produced and
convex, exterior margin rounded; subcostal two-branched beyond
end of the cell; discocellulars of equal length, bent inward ; median
vein four-branched, two upper beyond end of the cell, middle branch
from end of the cell; submedian nearly straight. Body moderate.
12 MR. F. MOORE ON THE LITHOSIID. [Jan. 15,
Palpi small, porrect, pilose. Antennze minutely and finely bipecti-
nate. Legs pilose, sparsely in male.
KorawA PALLIDA, 0. sp.
Male and female. Semidiaphanous ; fore wing pale fleshy-yellow ;
hind wing white ; thorax slightly ochreous-yellow ; abdomen white ;
palpi black-tipped ; legs white, fore tibize with a black longitudinal
line in front.
Expanse ¢ 14, 2 2 inches.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
HeEsupDRaA, 0. g.
Male. Wings short, rather broad. Fore wing with the costa nearly
straight ; apex acute, exterior margin very oblique; subcostal vein
five-branched, first branch very oblique, free, running close to costal,
second arising before end of the cell, trifurcate, fifth from end of
the cell, looped to third near its base; radial from upper end of cell;
discocellulars bent in the middle; median vein four-branched, two
upper from one fourth beyond the cell, third near its end. Hind wing
produced at the apex; exterior margin very oblique, nearly straight ;
abdominal margin long; two subcostal branches from one fourth
beyond the cell; median four-branched, two upper from one third
beyond the cell, third close to its end. Antenne broadly pectinate.
Palpi slender, curved upward, slightly pilose at base. Legs slender.
Hesupra Divisa, n. sp. (Plate I. fig. 4.)
Male. Upperside pale testaceous: fore wing witb a broad dark
purplish-grey band occupying the posterior half of the wing ; base
of the costa also purplish grey ; costal border yellowish. Thorax,
head, side of palpi, and legs above, purplish grey; abdomen pale
greyish, testaceous beneath, tip ochreous; base of palpi and legs
beneath pale ochreous ; antennz purplish-brown. Underside—fore
wing pale purplish-grey, costal border yellowish; hind wing narrowly
edged along the anterior margin with greyish black.
Expanse 1,3; inch.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
GuoRI, 0. g.
Fore wing long, narrow, costa slightly arched beyond the middle,
apex acute, exterior margin oblique and slightly convex, posterior
margin long ; first subcostal branch very oblique, free; second tri-
furcate; fifth from end of the cell and bent upward, looped to the third
near its base ; radial starting from below the fifth subcostal branch at
one third beyond the cell; discocellulars obliquely concave; median
four-branched, the two upper from end of cell, third immediately
before its end, fourth at half its length. Hind wing broad, apex
slightly produced, exterior margin convex in the middle ; two sub-
costal branches at one third beyond the cell; median four-branched,
two upper beyond the cell, third at its end. Body slender, extending
1878. } MR. F. MOORE ON THE LITHOSIIDE. 13
beyond hind wing. Antenne setose. Legs slender, squamose. Palpi
slender, porrect, base pilose beneath.
GHORIA ALBOCINEREA, n. sp. (Plate I. fig. 10.)
Male. Upperside—fore wing silky white, with a cinereous-brown
band along posterior margin: hind wing cinereous; cilia white.
Thorax and abdomen white; middle of thorax and streak on tegulee
cinereous brown; collar and front of head, and antenne yellowish ;
anal tuft pale brownish ochreous, Underside cinereous brown, palest
on hind wing, abdominal margin cinereous white ; costal edge of fore
wing yellowish towards the apex. Legs cinereous brown above,
yellowish beneath ; palpi yellowish, tip brown.
Expanse 1,4; inch,
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
GHORIA SERICEIPENNIS, Nl. sp.
Male. Upperside—fore wing silky white, with a broad dark
cinereous brown band along posterior margin ; hind wing white, pale
cinereous brown at the apex and along anterior margin. Middle of
thorax and tegulz, and front of head, dark cinereous brown; abdo-
men above and beneath white, anal tuft brown; collar yellowish ;
palpi ochreous yellow ; legs above cinereous brown, whitish beneath.
Underside—fore wing, anterior border, and apex of hind wing brown.
Expanse 12 inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
Curyseuia, Butler.
Chryseglia, Butler, Trans. Ent. Soc. 1877, p. 556.
Veins of the fore wing similar to those in @onistis entella. Those
in the hind wing differ in the median having but three branches, two
only starting from the end of the cell.
Type, C. magnifica (Lithosia magnifica, Walk. Journ. Linn. Soc.
vol. vi. p- 103), from Borneo.
CHRYSEGLIA FERRIFASCIATA, n. sp. (Plate I. fig. 6.)
Male and female. Upperside golden yellow: fore wing glossy,
with a steel-blue band along the costa, a broad transverse median band
and a large triangular spot near base of posterior margin; also a
blackish outer marginal band, which is broadest in the male ; median
band dilated hindward and angled on itsinner margin below the cell :
hind wing with the margin and cilia at the apex slightly blackish.
Tegulee and legs steel-blue. Underside paler yellow ; fore wing with
pale dusky brown bands as above.
Expanse, ¢ 1585, 2 24% inches.
Nepal (Ramsay), Darjiling (Grote). In coll. F. Moore.
Contstis, Hiibner.
Conistis, Hiibn. Verz. bek. Schmett. p. 165 (1816).
Fore wing—first subcostal branch long, free; second arising from
near end of the cell; third from its end, trifurcate; radial curving
14 MR. F. MOORE ON THE LITHOSIIDE. [Jan. 15,
from end of the cell; discocellulars bent in the middle ; median vein
four-branched, two upper from end, third from angle before end of
the cell. Hind wing—two subcostal branches immediately beyond
end of the cell, two upper median branches from beyond, and third
from end of the cell.
Type, @. entella (P. entella, Cram. Pap. Exot. pl. 208. f. D).
India, Ceylon,
Will also include Gnophria ceramensis, Voll. Tijdsch. voor Ent.
1872, iii. p. 246, pl. 12. f. 1.
Lirnosta, Fabr.
Lithosia, Fabr. Ent. Syst. Suppl. p. 459 (1798).
Sexes dissimilar in colour and pattern. Fore wing with first sub-
costal branch free, second starting near end of the cell, trifurcate,
fifth curving up from end of the cell and touching third near its base ;
radia] from end of the cell in male, from angle of upper discocellulars
in female; discocellulars curved in male, angled at both ends in
female ; median four-branched, three upper from end of the cell.
Hind wing with one subcostal branch in male, two in female beyond
the cell ; two median branches beyond tke cell in male, and from its
end in female.
Type, Lithosia quadra of Europe.
Will also include Lithosia dives (Monistis dives, Butler, Ann.
Nat. Hist. Nov. 1877, p. 398, from Japan.
Group B. Fore wing with subcostal vein r1vE-branched ; median
vein THREE-branched.
SIMAREEA, 0. g.
Similar in form, colour, and pattern to the male of the European
L. quadra, but of different venation in the fore wing; and both sexes
are alike, the discocellulars not being bent at the upper and lower end
of the cell (as in female Z. qguadra) ; and the median vein has only
two branches in both sexes proceeding from end of the cell (Z. quadra
having three in the male). In the hind wing the costal vein has two
branches beyond the cell in both sexes, Z. guadra having one only ;
the median branches also start at some distance from the cell in both
sexes. The palpi are more pilose, have the second and third joints
much longer, the apex blunt, and projecting much beyond the head.
SIMAREEA BASINOTA. (Plate I. fig. 16.)
Lithosia basinota, Moore, P. Z. S. 1865, p. 798.
Darjiling.
TARIKA, 0. g.
Fore wing long, broad ; costa arched; exterior margin convex,
posterior margin rounded at the base; first subcostal branch short,
oblique, anastomosing with the costal, second from near end of the
cell, third from near base of second, bifurcate, fifth from end of the
cell, bent upward and touching the third at one fourth its length;
discocellulars bent inward, upper angled close to subcostal, lower
1878. ] MR. F. MOORE ON THE LITHOSIIDZ. 15
shortest ; radial from angle of upper discocellular ; median three-
branched, two upper branches at one third beyond the cell; sub-
median nearly straight. Hind wing broad, apex slightly produced ;
subcostal branches at one fourth and median branches at one third
from end of the cell. Palpi small, short, slightly decumbent. Legs
stoutish, naked. Antenne minutely pectinate. Body slender, not
extending beyond hind wing.
TARIKA VARANA. (Plate I. fig. 8.)
Lithosia varana, Moore, P.Z.S8. 1865, p. 797.
Darjiling.
TARIKA NIVEA, 0. sp.
Lithosia nivea, Walker, MS.
Male and female. Fore wing silky white, costal edge pale yellow ;
head, front of thorax, paipi, body and legs beneath, yellow ; tip of
palpi brown; fore and middle legs brown above.
Expanse, ¢ 13, Q 12 inch.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
BRvnIA, 0. g.
Fore wing long, narrow; first branch of subcostal vein short, oblique,
anastomosed to the costal, second starting from angle close before end
of the cell, trifurcate, fifth from end of the cell and looped to third
branch at its base; discocellulars ‘bent below the middle, upper
angled close to end of the cell; radial from angle of upper disco-
cellular ; two upper median branches at half length beyond end of
the cell. Hind wing long, apex produced ; two branches of subcostal
vein at one third beyond the cell, and two median branches at two
thirds beyond. Palpi ascending, pilose beneath.
Brunia antica. (Plate I. fig. 9.)
Lithosia antica, Walker, Catal. Lep. Het. B. M. ii. p. 505.
Ceylon.
Will also embrace Zithosia natara, Moore, from Java, and L.
sarawaca, Butler, from Borneo.
GANDHARA, lh. g.
Fore wing broad, somewhat short ; costa much arched beyond the
middle ; apex acute; exterior margin slightly oblique, rounded in
middle ; hind margin slightly rounded at the base; first branch of
subcostal vein curving upward and extending free to beyond end of
the costal, second and third branches starting together from end of
the cell, third trifurcate; discocellulars zigzag, the angles equidistant ;
radial from upper angle; two upper median branches from lower
angle of cell. Hind wing broad, short.
GANDHARA sERVA. (Plate I. fig. 7.)
Iithosia serva, Walker, Catal. Lep. Het. B. M. ii. p. 506.
Darjiling.
16 MR. F. MOORE ON THE LITHOSIID&. [Jan. 15,
CoLuira, n. g.
Wings very broad. Fore wing with the costa arched; subcostal
five-branched, first short, oblique, anastomosed to costal, second
starting at some distance before end of the cell, third trifurcate, the
fifth or lowest branch nearest the base ; upper discocellular slightly
angled close to subcostal, whence starts the radial; median three-
branched, two upper at one fourth beyond the cell. Hind wing very
broad, apex produced, exterior margin rounded in the middle, ab-
dominal margin long and extending beyond abdomen; two subcostal
branches and two upper medians from one third beyond the cell.
Palpi short, porrect.
Type, C. griseola (Lithosia griseola) of Europe. Will also
include ZL. vetusta, Walk., from China, and the European L. compla-
nula and L. stramineola, Dbleday.
COLLITA LILACINA, 0. sp.
Fore wing pale lilac-grey, with pale yellowish costal border ; hind
wing pale greyish ochreous; thorax grey; head pale yellow; palpi
black-tipped ; fore legs brownish above.
Expanse 14 inch.
W. Yunan (Anderson). In coll. F. Moore.
Similar in appearance to Gandhara serva (Lithosia serva, Walk.),
from Sikkim, but of entirely different venation.
Courts PARvA, n.sp. (Plate I. fig. 13.)
Female. Fore wing greyish ochreous-brown; hind wicg and
abdomen paler; fore wing yellowish along the costal border. Un-
derside uniform ochreous brown, palest on lower part of the hind
wing. Legs cinereous brown above.
Expanse 355 inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
Allied to C. vetusta (Lithosia vetusta, Walk.), from North China,
having the same venation, but is of a much smaller size.
KaTHA, n. g.
Wings clothed with stout compact imbricated scales.
Male. Fore wing long, very narrow ; costa nearly straight, apex
rounded ; exterior margin slightly oblique ; posterior margin re-
curved, convex near the base; slightly folded longitudinally below
the median vein ; cell long, narrow; discocellulars bent outward at
upper end and inward at lower end; first branch of subcostal vein at
some distance before end of the cell, short, oblique, and anastomosed
to costal; second short, arising beyond end of the cell; third trifur-
cate, its lowest branch at equal distance between second and fourth ;
radial starting from upper angle of discocellular ; two upper branches
of median vein at half distance from end of the cell; submedian
recurved. Hind wing long, broad; two subcostal branches at one
third from cell; discocellulars slightly bent inward, two median
branches at one half distance from the cell, and wide apart. Palpi
1878. ] MR. F. MOORE ON THE LITHOSIID®. 17
rather short, pilose at base, porrect. Antennz minutely pectinate.
Legs long, rather stout. Body long, extending beyond hind wing.
Karua nierirrons. (Plate I. fig. 15.)
Lithosia nigrifrons, Moore, P. Z. 8. 1872, p. 572.
North India.
This genus will also include Lithosia apicalis, Walk., from Borneo,
L. intermixta, Walk., L. brevipennis, Walk., from South India, and
L. helveola of Europe.
KaTua TERMINALIs, n. sp. (Plate I. fig. 14.)
Male. Ochreous; fore wing darkest, with a distinct curved pur-
plish band across exterior margin. Antenne, front of head, tip of
palpi, middle of thorax, and streak on tegulz purplish black ; legs
above purplish brown.
Expanse 1}, inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
Allied to K. apicalis (Lithosia apicalis, Walk. Journ. Linn. Soc.
Zool. vi. p. 104) from Borneo, but differs on the fore wing in the
apical band not extending upward onto the costa, and in the absence
of the slight apical patch on the hind wing, which is present in
Bornean examples.
KaTHa CUCULLATA, 0. sp.
Male. Upperside—fore wing pale purplish grey, costal edge at
apex and cilia yellowish ; an indistinct greyish-brown outer mar-
ginal band: hind wing and abdomen pale ochreous grey ; anal tuft
ochreous. Thorax, top and front of head, and palpi except at base
beneath, black; legs black, middle and hind legs pale beneath ;
antennze black. Underside—fore wing brownish ochreous; hind
wing pale ochreous.
Expanse 1+), inch.
Andamans. In coll. F. Moore.
Allied to K. terminalis and K. apicalis.
Ertema, Hibner.
Eilema, Hiibn. Verz. bek. Schmett. p. 165 (1816).
Wings long, narrow. Fore wing with the subcostal five-branched,
first short, oblique, and anastomosed to costal, second free and widely
separated from third, third trifurcate, the lowest or fifth nearest the
base, the two upper near apex ; radial from upper end of the cell;
discocellulars inwardly oblique; median three-branched, two upper
beyond half distance from the cell, lower long, from near its base.
Hind wing—two subcostal branches at one third, and two upper
median branches at two thirds beyond the cell.
Type, Z. caniola, Hiibner, of Europe.
Manutea, Wallengren.
Manulea, Wallengren, Ent. Monats. vii. p. 145.
Wings narrow. Fore wing with the costa not arched ; subcostal vein
Proc. Zoou. Soc.—1878, No. II. 2
18 MR. F. MOORE ON THE LITHOSIID™. (Jan. 15,
five-branched, first very oblique and free, running close to costal near
its end, second arising at some distance before end of the cell, third
starting and curving downward from second at half its length, bifur-
cate, fifth from end of cell and touching the third near its base ;
radial in a line with subcostal from end of the cell; discocellulars
bent in the middle; median three-branched, two upper from one
fourth beyond the cell. Hind wing narrow; abdominal margin
short; two subcostal branches close to end of the cell; two upper
median branches at nearly half length beyond.
Type, WM. gilveola (Lithosia gilveola) of Europe.
MANULEA CALAMARIA, 0. sp.
Male. Pale glossy straw-yellow ; costal edge of fore wing at base
dusky. Head, thorax, body beneath, and anal tuft brighter yellow;
palpi black-tipped ; legs above, and tarsi entirely, black ; antennze
yellow. Underside darker; fore wing pale brownish luteous.
Expanse 11 inch.
N.W. Himalaya. In coll. F. Moore.
This species is allied to the European M. gilveola.
SystropHa, Hiibn.
Systropha, Hiibn. Verz. bek. Schmett. p. 166 (1816).
Wings small. Fore wing short, narrow; subcostal five-branched,
first branch short, oblique, anastomosing partly with costal near its
end, second at some distance before end of the cell, third trifurcate,
lowest branch at some distance beyond the cell; upper discocellular
shortest, lower oblique and bent inward near end; radial from angle
of upper; two upper median branches at half distance beyond the
cell. Hind wing broad.
Type, S. aureola (Lithosia aureola) of Europe.
SysTROPHA DORSALIS, 0. sp.
Female. Fore wing straw-yellow, slightly ochreous along posterior
margin: hind wing yellowish white. Head, thorax, legs, and abdo-
men at sides and beneath ochreous yellow; abdomen above lilac-
grey.
Expanse 1,2; inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
SYSTROPHA AURIFLUA, n.sp. (Plate II. fig. 7.)
Fore wing glossy straw-yellow ; hind wing paler yellow. Thorax,
head, legs, and anal tuft brighter yellow.
Expanse 119 inch.
Manpuri, N.W. India (Horne). In coll. F. Moore.
CurysorapspiA, Butler.
Chrysorabdia, Butl. Trans. Ent. Soc. 1877, p. 357.
Fore wing—first subcostal branch contiguous to costal, second from
before end of the cell and bent onto the third above its end, third
trifurcate, curved upwards ; radial from end of the cell; discocellulars
1878.] MR. F. MOORE ON THE LITHOSIID. 19
bent in the middle ; median vein three-branched, two upper from end
of the cell. Hind wing—two subcostal branches; discocellulars very
slender, scarcely visible, two upper median branches.
CHRYSORABDIA VIRIDATA. (Plate II. fig. 1, 3.)
Lithosia viridata, Walk. Catal. Lep. B. M. Suppl. p. 225.
Darjiling.
CaPissA, 0. g.
Fore wing long, narrow ; costa arched before the apex, exterior
margin oblique and convex, hind margin slightly convex near the
base. Male with a Jongitudinal fold below the cell’, where the scales
are raised and rough; cell short, narrow; discocellulars of equal
length, bent inward; first subcostal branch very short, oblique,
anastomosed to costal; second immediately before end of the cell ;
third at its end, and trifurcate ; radial from beyond end of the cell,
starting from the third branch of subcostal near its base ; median
vein straight, three-branched, two upper from beyond end of the cell ;
submedian slightly recurved. Hind wing long, broad, apex produced,
the male having the scales on the disk raised and rough ; two sub-
costal and two median branches beyond end of the cell ; discocellulars
bent inward, upper the longest. Body long, slender, extending be-
yond hind wing. Antennz minutely pectinate in male, setose in
female. Palpi prominent, pilose at base, apex slender. Legs long,
slender.
Carissa 1nNoTaTa. (Plate II. fig. 2, 3.)
Lithosia innotata, Butler, Trans. Ent. Soc. 1877, p. 352.
N.W. Himalayas.
This genus will also include Z. vagesa, Moore, from the Khasia
hills, Z. insolita, Walk., from China, L. nigripars, Walk., from N.
India, and Z. sambara, Moore, from Java.
CapIssA AURIFLAVA, ND. Sp.
Female. Fore wing deep golden-yellow ; hind wing and abdomen
paler. Underside of fore wing greyish at the base; femur, tibia,
and tarsal joints with a black terminal band; palpi black at tip ;
antennee yellow.
Expanse 1,8; inch.
Katmandu, Nepal (General Ramsay). In coll. F. Moore.
Allied to C. vagesa (Lithosia vagesa, Moore, Catal. Lep. E.I. C.
ii. p. 304), but is of a deeper colour, and the legs are banded only
with black, whereas in C. vagesa the legs are entirely black.
CaPissA PALLENS, n.sp. (Plate II. fig. 3.)
Female. Pale whitish ochreous; fore wing glossy, costal edge
ochreous. Thorax, palpi, and abdomen beneath ochreous; legs
dusky brown above; palpi brown at tip; antenne brown. Un-
derside—costal border of both wings brighter-coloured ; middle of
fore wing pale ochreous brown.
1 Probably a sound-producing apparatus.
20 MR. F. MOORE ON THE LITHOSIID. [Jan. 15,
Expanse 12 inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
Somewhat allied to C. insolita (Lithosia insolita, Walker, Catal.
Lep. Het. B. M. ii. p. 497).
CaPISSA FLAVENS, 0. sp.
Female. Glossy straw-yellow; hind wing and abdomen above
slightly paler; apex of palpi, fore legs above, a terminal spot on
middle and hind femora, and tarsi dusky brown.
Expanse 1+ inch.
Saidabad, Cashmere (Hellard). In coll. F. Moore.
CapissA FASCIATA, n. sp. (Plate II. fig. 4, 3.)
Male. Upperside ochreous yellow; fore wing with a brown-
speckled fascia along the posterior border and about the disk ; hind
wing and base of abdomen pale yellow; antenne, apical joint of
palpi, and legs above brown.
£Expanse 1,4; inch.
Ceylon. In coll. F. M. Mackwood.
Do.goma, n. g.
Male and female. Wings moderately long and broad. Fore wing
with the subcostal vein bent at its second branch, five-branched,
first branch curving upward but free from costal, second starting
before end of the cell, curved upward, third trifureate, lowest branch
at one third its length beyond the cell; upper discocellular erect,
lower oblique and bent inward; radial starting from the middle; two
upper median branches at less than one fourth from end of the cell,
lower branch angled and with a slight spur near its base. Hind
wing—subcostal branches at one third and median branches at half
distance beyond the cell.
Type, D. reticulata (Lithosia reticulata, Moore, P. Z. 8. 1865,
p- 798).
Darjiling.
DOLGOMA ANGULIFERA.
Lithosia angulifera, Feld. Nov.-Reise Lep. pl. 106. f. 12 (1868).
Male and female. Luteous ; fore wing sparsely brown-speckled,
crossed by a brown-speckled discal band, which is bent outward
beyond end of the cell; hind wing pale luteous white. Underside
paler-coloured.
Expanse, ¢ 14, 9 14 inch.
Darjiling. In coll. F. Moore.
DoLGoMA BRUNNEA, n. sp. (Plate II. fig. 8.)
Male. Cinereous brown; fore wing uniformly dark-coloured ;
hind wing paler. Underside paler than above ; costal border of fore
wing, and legs beneath, yellowish.
Expanse | inch.
Darjiling (Atkinson). In coll, Dr. Staudinger and F. Moore.
1878. | MR. F. MOORE ON THE LITHOSIID2. 21
MITHUNA, 0. g.
Wings short, rather broad. Fore wing arched, apex pointed ; first
branch of subcostal vein arising near end of the cell, short, oblique,
anastomosing with costal but free at itsend; second branch quadrifid,
the upper and lower branches starting together at one third beyond
the cell; upper discocellular very short, lower curved inward ; radial
from lower end of upper discucellular ; two upper median branches
starting at half distance beyond the cell, lower branch straight from
one third before its end. Hind wing with subcostal and median
branches at nearly one half length beyond the cell. Body short.
Palpi small, pilose beneath. Antenne setulose. Legs smooth.
MiTHUNA QUADRIPLAGA, n.sp. (Plate II. fig. 9.)
Fore wing luteous brown, with a distinct dusky-brown median
transverse band, which is angled outward at end of the cell, a similar
band also crossing the disk ; outer margin with a series of dusky
spots, which also indistinctly cross the cilia: hind wing pale luteous
brown. Underside paler. Head and thorax brown.
Expanse | inch.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
Cossa, Walker.
Cossa, Walk. Catal. Lep. Het. B. M. pt. xxxii. p. 232 (1864).
Male and female. Wings somewhat broad. Fore wing arched along
the costa; first subcostal branch short, very oblique, anastomosed to
costal, second free, third from end of the cell, trifurcate ; discocel-
lulars very oblique, angled at ends ; radial from upper angle; two
upper median branches from one third beyond the cell.
Type, C. basigera, Walk. 1. c.
S. India.
Cossa QUADRISIGNATA, n.sp. (Plate II. fig. 10.)
Male. Fore wing dark purplish brown, with slightly paler streaks
along the veins; costal border pale purplish cinereous, with two
prominent short black streaks ; a small spot at end of the cell: hind
wing and abdomen above pale brownish cinereous ; anal tuft slightly
ochreous ; thorax, head, abdomen beneath, and legs brown. Un-
derside pale cinereous brown; middle of fore wing brown.
Expanse 11 inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
Cossa PALLIDA, D. sp.
Male. Pale greyish ochreous: fore wing with two prominent
short black broad streaks on the costa, the first subbasal, the other
slightly beyond the middle; a short, narrow, black streak at end of
the cell: hind wing with a very indistinct dusky submarginal fascia.
Abdomen greyish, tuft ochreous. Underside pale yellowish ochreous ;
middle of fore wing ochreous brown. Legs greyish brown above.
Expanse 1,1; inch.
Dharmsala (N.W. Himalaya). In coll. F. Moore.
22 MR. F. MOORE ON THE LITHOSIID. (Jan. 15,
Cossa BRUNNEA, n.sp. (Plate II. fig. 11.)
Female. Fore wing dark purple-brown, with a small black spot at
end of the cell, and a short black streak on the costa near the
middle: hind wing paler. Thorax, head, and legs dark brown.
Underside with the margins paler.
Expanse 14 inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
RANGHANA, 0. g.
Female. Fore wing very long, narrow; costa arched, apex pointed,
exterior margin very oblique, angle and posterior margin rounded ;
subcostal vein five-branched, first branch short, arising immediately
above end of the cell, and joining the costal, second and third at
equal distance from first, third trifurcate; radial from below sub-
costal beyond the cell at equal distance between first and second
branches ; cell short ; discocellulars straight ; median vein straight,
three-branched, two upper at one third from the exterior margin,
lower recurving from below end of the cell; submedian veins
extending to posterior angle. Hind wing long, narrow ; subcostal
branches at half length beyond the cell; cell short; discocellulars
deeply curved ; median straight, two-branched. Body short. Palpi
very short, stout. Legs long, slender; middle and hind tibize
spurred. Antenne setose.
RaNGHANA PuNCTATA, n. sp. (Plate II. fig. 12.)
Female. Pale ochreous-buff colour ; fore wing with a marginal and
apical series of nine small black spots. Underside paler, without
marks.
Expanse 11 inch.
Calcutta (Atkinson). In coll. Dr. Staudinger.
TreGuuata, Walker.
Tegulata, Walk. Journ. Proc. Linn. Soc. 1862, p. 110.
Female. Fore wing with a convex protuberance on middle of
the costa ; apex pointed ; first subcostal branch short, anastomosed
to costal, second from angle before end of the cell, third trifurcate,
lowest or fifth branch nearest base; discocellulars angled at each
end ; radial from upper angle below end of the cell; median vein
three-branched, two upper one-third beyond the cell.
Type, 7. tumida, Walk. l. c., from Borneo.
TEGULATA BASISTRIGA, n.sp. (Plate II. fig. 5.)
Female. Upperside pale luteous brown: fore wing brightest at the
apex, slightly brown-speckled ; a black streak along base of costal -
edge and along base of posterior margin: hind wing pale brownish
fawn-colour externally. Underside darker. Legs brown above.
Expanse 1 inch.
Ceylon. In coll. Dr. Staudinger.
Allied to 7. tumida, Walk.
1878. | MR. F. MOORE ON THE LITHOSIIDE. 23
TEGULATA PROTUBERANS, n.sp. (Plate II. fig. 6.)
Female. Upperside pale luteous brown; fore wing with numerous
brown speckles and a dark brown prominent costal spot ; hind wing
and abdomen pale luteous-yellow. Underside paler; middle of fore
wing brownish.
Expanse 1,2; inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
Group B. a. Hind wing with a recumbent plumose lappet.
NisHADA, 0. g.
Male and female. Fore wing somewhat short, broad, costa consi-
derably arched, apex very acute; exterior margin oblique and
slightly convex; posterior margin short, in the male convex and
fringed to near the base; subcostal vein five-branched, first branch
arising at half length of the cell and slightly touching the costal
vein, second at one third before end*of the cell, third at some distance
beyond the cell, fourth bifid at half its length; cell narrow ; disco-
cellulars very slender, upper short, lower obliquely curved inward ;
radial starting from lower end of upper discocellular ; median vein
three-branched, the two upper at nearly half distance beyond end of
cell, lower before its end; submedian vein recurved upwards from
the base. Hind wing in the male very short and broad, somewhat
quadrate, anterior margin produced upward from the base and folded
over onto the underside, and there provided with a long recumbent
fan-like plumose covering or appendage’ ; the hind wing in the female
is longer, being of the same length as posterior margin of fore wing,
its anterior margin nearly straight, and is without the folded plumose
appendage; costal vein following the anterior margin beneath the
fold in the male; subcostal vein two-branched, the upper arising
near base of the wing; cell broad; discocellulars recurved ; median
vein straight, two-branched, the branches very close together, lower
branch from before end of the cell; submedian straight. Antenne
minutely and finely pectinate in male, setose in female. Body
slender, extending beyond hind wing. Legs stout, smooth; fore femora
thickened ; middle and hind tibiz spurred. Palpi small, porrect,
projected slightly beyond the head, pilose beneath, apex small, pointed.
NISHADA FLABRIFERA, N. sp.
Male and female. Yellowish ochreous, palest in female ; fore wing
in male brighter ochreous, and slightly ferruginous at the base;
thorax, head, and antennez brownish ochreous.
Expanse 1 inch.
Calcutta District (Zarr). In coll. F. Moore.
Allied to the Bornean Lithosia rotundipennis, Walk. Journ. Linn.
Soe. Zool. vi. p. 104, which species is also probably identical with
L. chilomorpha, Snellen Van Voll., Tijd. voor Ent. 1877, p. 67, pl. 5.
fig. 1, from Sumatra. Both of these species will come into the
genus Nishada.
1 Probably a sound-producing apparatus,
24 MR. F. MOORE ON THE LITHOSIID. [Jan. 15,
Group B. B. Fore wing with a recumbent plumose lappet.
Trutisna, Walk.
Teulisna, Walk. Journ. Proc. Linn. Soc. 1862, p. 109.
Male. Wings somewhat short and broad. Fore wing—costa
depressed at the base, arched in the middle ; apex pointed, exterior
margin oblique, angle acute ; posterior margin very convex towards
the base; first subcostal branch short, anastomosed to costal; second
free; third from end of cell, trifurcate ; radial from third submedian
branch at a considerable distance beyond the cell; cell folded over at
base and overlapped for half its length by a recumbent plumose
appendage’ ; discocellulars very oblique ; median three-branched, two
upper at some distance beyond the cell.
Female without the fold and plumose tuft.
Type, T. plagiata, Walk. Journ. Linn. Soc. 1862, p. 109, from
Borneo.
TEULISNA TENUISIGNA, n. sp. (Plate II. fig. 13.)
Male. Yellow. Differs from the same sex of T. tetragona in its
smaller size, being also narrower in both wings. Fore wing with a
lengthened black band beneath the cell, the upper end of which is
straight, and the basal end very oblique and truncated ; a small spot
on costa before the apex ; cilia slightly blackish at the apex. Middle
of thorax, tegule, and front of head black; fore legs dusky above.
Expanse 1;% inch.
Sikkim. In coll. F. Moore.
MAcotTasa, n. g.
Male. Fore wing narrow ; costa slightly raised beyond the middle ;
costal vein raised, with a recumbent plumose tuft overlapping basal
half of the folded cell beneath’; subcostal five-branched, first short,
oblique, anastomosed to costal, second starting at some distance be-
fore end of the cell, third starting from second near its base and bent
downward and touching end of the cell, bifurcate near its end, fifth
starting in a straight line from end of the cell ; discocellulars straight,
nearly erect ; radial starting in a straight line from lower end of the
cell; median three-branched, the two upper at some distance apart
below the radial and parallel to it, the base starting from below end
of the cell in a line with the discocellulars; lower median branch
from near base of cell, deeply curved. Hind wing somewhat narrow,
apex pointed ; two subcostal branches at one third beyond the cell ;
two upper median branches near its end. Antenne minutely setu-
lose. Palpi small, ascending. Legs slender.
Female. Fore wing not folded or plumed; radial starting from
upper end of the cell; two upper median branches straight from
beyond end of the cell.
1 Probably a sound-producing apparatus.
1878. | MR. F. MOORE ON THE LITHOSIID £. 25
MACOTASA BIPLAGELLA. (Plate II. fig. 14.)
Gcophora biplagella, Walk. MS.
Upperside pale whitish luteous: fore wing with a dark brown
median costal streak, which is indistinctly continued as a narrow band
across the wing, the streak more prominent and larger in the male ;
apex also brown-streaked : hind wing whitish at the base.
Expanse, ¢ 15, Q 1, inch.
Borneo. In coll. British Museum and F. Moore.
ZADADRA, N. g.
Male. Fore wing shortened, distorted, and crumpled at the apex ;
costa arched beyond the middle, apex very convex, exterior margin
very oblique hindward, posterior margin shortened and very convex ;
cell very long, folded and overlapped above with a short tuft of
recumbent plumes its entire length’ ; subcostal and median veins con-
tiguous, the branches being near apex of the wing ; discocellulars very
convex ; subcostal at some distance from costal, first branch recurved
upward from end of the cell, second starting from near base of the
first, bifurcate near the end, fourth before end of the cell and touching
the second near its base, fifth branch and radial from eud of cell;
median vein three-branched, two upper beyond end of the cell, lower
recurving from end near and parallel to upper branch ; submedian
boldly recurved. Hind wing long, broad; apex much produced ; ex-
terior margin recurved ; cell long; upper discocellular long, inwardly
oblique, lower very short and outwardly oblique; subcostal bifid from
end of cell ; median three- branched, two upper beyond end of cell,
lower at half its length ; submedian nearly straight. Body slender,
longer than hind wing. Antenne minutely and finely bipectinate.
Legs long, smooth, spurred. Palpi slender, porrect ; base pilose ;
apex long.
Female. Wings longer ; cell of fore wing not folded or tufted, and
the apex not crumpled ; hind margin much less convex, and the
submedian less recurved.
ZADADRA DisTorRTA. (Plate II. fig. 15.)
Lithosia distorta, Moore, P. Z. 8. 1872, p. 572.
Darjiling. In coll. F. Moore.
PRABHASA, 0. g.
Wings long. Fore wing narrow, very slightly arched before the
apex; exterior margin oblique, slightly convex ; posterior margin
long, nearly straight ; subcostal vein at some distance from costal,
first branch short, curving upward to costa before end of the cell,
second straight from end of the cell, third contiguous at base to the
second, trifurcate, looped to second near base ; cell long, in the male
folded and tufted with recumbent plumes above to beyond half its
length’ ; discocellulars long, convex ; radial from lower part ; median
vein three-branched, two upper curving hindward from lower end of
the cell, lower curving from below the cell at nearly half its length ;
1 Probably a sound-producing apparatus.
26 MR. F. MOORE ON THE LITHOSIID. (Jan. 15,
submedian recurved. Hind wing produced at the apex; exterior
margin recurved ; abdominal margin short; subcostal two-branched
beyond the cell; discocellulars concave; cell short, broad; median
vein three-branched, two upper branches at half distance beyond
the cell, lower from before half its length; submedian nearly
straight. Body slender, longer than hind wing, tufted in male.
Antenne slender, with very fine delicate pectinations. Legs slender,
long, smooth, spurred. Palpi slender, long, slightly ascending be-
yond front of head.
PRABHASA VENOSA, n. sp. (Plate II. fig. 16, 3.)
Male and female. Upperside pale luteous brown: fore wing with
the veins darker brown ; an indistinct brown transverse discal band,
which is bent outward beyond the cell; male with a greyish-brown
plumose tuft overlapping and extending half the length of the cell;
anal tuft in male ochreous. Underside paler; band across fore wing
not visible.
Expanse, 3 154, 2 153; inch.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
PRABHASA FLAVICOSTA, n. sp. (Plate II. fig. 17.)
Female. Upperside uniformly dark luteous brown: hind wing and
abdomen purplish brown: fore wing with the costal border pale
yellow ; discal band most prominent at costal end, broad, but not
bent outward. Front of head, thorax at sides, palpi and abdomen
beneath, yellow; legs above cinereous brown, yellowish beneath.
Underside uniform brown ; costal border on both wings yellowish.
Expanse 1,3; inch.
Cherra Punji (dékinson). In coll. Dr. Staudinger and F. Moore.
PRABHASA COSTALIS, 0. sp.
Male and female. Whitish testaceous : fore wing with the costal
border whitish, and a brown spot one third before the apex; lower
part of wing brown-speckled, most distinct along the veins. Un-
derside—fore wing testaceous brown, dark-speckled along the base
of costa and subcostal vein. Legs dusky brown above.
Expanse, dg 1, 2 1,%; inch.
North China. In coll. F. Moore.
GAMPOLA, 0. g.
Male. Wings short, broad, truncate at the apex. Fore wing arched
towards the end, apical angle acute, exterior margin nearly erect,
posterior angle convex and fringed ; costal edge at the base folded
over and fringed for a short distance’; cell long and narrow, folded
beneath the median vein; subcostal at some distance from costal,
five-branched, first short, anastomosing with costal, but free at its end,
second arising at some distance before end of the cell, third trifurcate,
the upper branch nearest to and at one third beyond the cell; disco-
cellulars bent below the middle, outwardly oblique, upper longest ;
radial from the angle ; median vein bent upward near end of cell,
} Probably a sound-producing apparatus.
1878. | MR. F, MOORE ON THE LITHOSIID&. 27
top branch from upper angle, second from lower, third from before
half length of the cell. Hind wing quadrangular ; two subcostal and
two upper median branches from one third beyond the cell, lower
median from half its length. Antenne finely pectinated. Palpi
small, porrect. Legs slender, naked, middle and hind spurred.
Body slender, short.
Female. Wings longer, not truncate at apex; fore wing not
folded ; veins as in male, excepting that the hind wing has, appa-
rently, only one upper median branch.
GamPpoLa FasciaTA, n. sp. (Plate II. fig. 18.)
Male. Upperside—fore wing pale brownish ochreous, brown-
speckled, the speckles composed of dark stout scales forming a fascia
below the cell and across the disk. Head, thorax, and abdomen
fuliginous brown ; anal tuft slightly ochreous ; palpi brown at tip ;
legs slightly brown above. Hind wing and underside paler.
Female. Fore wing not brown-speckled, with an indistinct trans-
verse dusky maculated discal band and a small spot at posterior
angle. Underside dusky at apex of fore wing.
Expanse, ¢ 14, Q 1), inch.
Ceylon. In coll. F, M. Mackwood and Dr. Staudinger.
Group C. Fore wing with subcostal vein rour-branched, median
vein THREE-Oranched.
Pexosra, Hiibn.
Pelosia, Hibn. Verz. bek. Schmett. p. 165 (1816).
Samera, Wallengren, Wien. ent. Mon. vii. p. 146.
Male and female. Wings rather short and broad. Fore wing with
subcostal and costal veins wide apart; subcostal four-branched ;
first branch short, oblique, anastomosing with costal but free beyond
its end; second arising at some distance before end of the cell,
straight, widely separated between first and third; third bifid, at
half distance from end of the cell; discocellulars bent inward, lower
shortest, upper angled close to subcostal; radial from the angle of
upper; two upper median branches from half distance beyond the
cell, lower from half before its end. Hind wing—two branches
of subcostal at one fourth, and median branches at two thirds
beyond the cell.
Type, P. muscerda (Lithosia muscerda) of Europe.
Group D. Fore wing with a plumose lappet-covered sac-like cavity’.
Bizone, Walker.
Bizone, Walker, Catal. Lep. B. M. ii. p. 548.
BiZONE BELLISSIMA, N. sp. (Plate III. fig. 13.)
Male. Upperside—fore wing with a transverse basal and two
oblique median red bands, a broad outer marginal ochreous-yellow
band, and three black spots at end of the cell: hind wing and abdo-
' Probably a sound-producing apparatus.
28 MR. F. MOORE ON THE LITHOSIID. (Jan. 15,
men red; tuft white. Thorax white, with two red bands. Under-
side uniform red; cell-spots in fore wing and discocellular streak in
hind wing blackish. Fore and middle legs with blackish bands.
Body beneath and legs white. Palpi black at the side.
Expanse 1,8; inch.
Masuri, N.W. Himalaya (Hutton). In coll. F. Moore.
Bizone coccingEA, n.sp. (Plate III. fig. 14.)
Male. Upperside crimson-red, the base of fore wing slightly
yellow: fore wing with a subbasal and discal black band, the former
bent inward and the latter outward at the costal end; three black
spots at end of the cell. Thorax yellowish, banded with red ; anal
tuft and abdomen beneath yellow; legs yellow, with red bands.
Underside of wings uniform red.
Expanse 1,5, inch.
Sikkim. In coll. F. Moore.
BIZONE ALBA, N. Sp.
Male. Upperside white: fore wing with a small basal red spot
and narrow streak on base of costal edge ; a black spot at end of the
cell. Underside pale red, broadly on anterior margin of fore wing
and narrowly on hind wing; palpi at sides and fore and middle legs
with blackish bands.
Expanse ;®; inch.
North China. In coll. F. Moore.
Group E. dberrant. Fore wing with rive subcostal and rour
median branches.
BaRoa, n. g.
Fore wing long, moderately broad; costa slightly arched ; apex
somewhat pointed ; exterior margin oblique and convex ; cell short,
narrow ; subcostal vein five-branched, first branch free, second tri-
furcate, the forks ascending, fourth from end of the cell; disco-
cellulars angled at the ends; radial from upper angle; submedian
four-branched, two upper from end and third from angle before end of
the cell. Hind wing somewhat short, broad ; apex pointed ; exterior
margin convex; two subcostal branches from end of the cell; three
submedian branches from end and fourth before end of the cell.
Body extending beyond hind wing. Palpi slender, long, pilose
beneath, ascending and extending beyond the head; terminal joint
blunt. Legs slender. Antenne minutely pectinate in male, setulose
in female.
Type, Baroa punctivaga (Cycnia punctivaga, Walk. Catal. Lep.
Het. B. M. iii. p. 682).
Java (Horsfield).
BarsineE, Walker.
Barsine, Walk. Catal. Lep. Het. B. M. ii. p. 456.
BARSINE FLAMMEALIS, n. sp. (Plate III. fig. 15.)
Fore wing yellow, mostly covered with bright-red intervening
1878. ] MR. F. MOORE ON THE LITHOSIID. 29
streaks between the veins, crossed by three outwardly recurved
grey-brown bands; some spots of same colour at base of the wing,
and longitudinal discal streaks joining the outer band between the
veins: hind wing pale pinky red. Body red; thorax with grey-
brown spots. Underside—fore wing mostly red, slightly yellow on
outer veins, and brighter yellow on outer margin; legs red.
Expanse 2 inches.
India. In coll. F. Moore.
Nearest allied to B. mactans, Butler.
BarsINE GLorrosa, n.sp. (Plate III. fig. 16.)
Female. Upperside—fore wing ochreous red, veins broadly lined
with yellow; a broad blackish well-defined subbasal cross band, a
narrow discal recurved band, and an apical series of short black lon-
gitudinal streaks ; two small black dots at base of wing; costal edge
and cilia black: hind wing and abdomen pale pink. Thorax red,
with a narrow black middle streak and two anterior spots; palpi
and legs red. Underside —fore wing red, with short black costal and
apical streaks ; hind wing paler, apex dusky black; middle tibize
with a black terminal streak.
Expanse 1,4; inch.
Khasia hills (Adkinson). In coll. Dr. Staudinger.
Nearest allied to B. eruciata, Walker, from Borneo.
BaRSINE PUNICEA, 0. sp.
Female. Upperside—fore wing ochreous yellow, slightly reddish
along the costal border ; with a blackish subbasal cross band, a waved
curved discal band, and an outer series of alternated long and short
longitudinal streaks ; cilia and apical edge of costa black : hind wing
pale ochreous red, with black marginal band. Abdomen blackish,
tip yellow. Underside dull ochreous red; apex of fore wing and
outer margin of hind wing blackish; middle tibie with a terminal
black streak.
Expanse 7 inch.
Darjiling (A¢kinson). In coll. Dr. Staudinger and F. Moore.
This species is nearest allied to B. lineata, Walk. Cat. Lep. Het.
B. M. iii. p. 760, from Borneo.
BaRSINE INFLEXA, n. sp. (Plate III. fig. 17.)
Male. Upperside—fore wing ochreous red, veins lined with yellow ;
costal edge, posterior margin, and cilia black ; a black subbasal cross
band somewhat thickened within the cell, the outer lower arm of
which is bent inward; a narrow discal transverse maculated band,
and an outer series of short streaks, one on each vein: hind wing and
abdomen pale pink. Thorax and head red, streaked with yellow ;
legs red. Underside—fore wing red, with black apex and margins ;
hind wing yellowish, with dusky black apex.
Expanse { inch.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
30 MR. F. MOORE ON THE LITHOSHD. (Jan. 15,
BARSINE FLAVIVENOSA, n.sp. (Plate III. fig. 18.)
Male. Upperside—fore wing red, with all the veins very broadly
lined with yellow; a blackish subbasal imperfect cross band, with the
lower portion of the outer arm obsolete; a curved discal band and
an incurved apical series of short black streaks : hind wing, abdomen,
and legs very pale pink. Underside pink, hind wing palest.
Expanse € inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
LycLene, Moore.
Lyclene, Moore, Catal. Lep. E. I. C. ii. p. 390.
Cyllene, Walk. Catal. Het. B. M. il. p. 544.
LycLENE RUBRICOSA, n. sp. (Plate III. fig. 1.)
Maie and female. Ochreous red: fore wing with dusky-black
subbasal broad zigzag band, an oblique median band and zigzag outer
discal band, the latter slightly confluent about its middle in the
female ; a small spot also at base of wing, a curved spot at end of
cell, and a small series along exterior margin. ‘Thorax black-spotted.
Underside with markings on fore wing indistinct ; fore and middle
tibize with a blackish streak.
Expanse ;; inch.
Bombay (Dr. Leith). In coll. F. Moore.
Allied to L. artocarpi.
LYCLENE ARTOCARPT, 0. sp.
Male and female. Fore wing ochreous, with a dusky-grey subbasal
series of short longitudinal streaks; an upright median band and
outward discal oblique irregular band, both confluent on middle of
the hind margin; a small blackish spot at base of wing, and an
indistinct spot at end of the cell: hind wing and abdomen pale
ochreous. Thorax with black spots. Underside paler, markings
on fore wing less distinct ; an indistinct median dusky fascia across
hind wing in the female.
Expanse 4 inch.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
Nearest allied to L. humilis, Walk. Catal. Lep. Het. B. M. ii.
p- 554. ‘Larva feeds on the Jack tree (Artocarpus incisa),
January.’ —A. Grote, MS. note.
LYCLENE RADIANS, n. sp. (Plate III. fig. 2.)
Male. Fore wing yellowish white, with a black basal spot, three
transverse series of spots, each composed of three, and the middle
series curved; a prominent spot at end of the cell, and a marginal
series of broad black lines, one on each vein, their inner ends con-
fluent and forming a bordered line, which is bent outward at its
middle: hind wing with a less prominent series of short black mar-
ginal lines, which do not extend to the anal angle. Body ochreous
yellow. Underside similarly marked, the basal spots on fore wing
1878. ] MR. F. MOORE ON THE LITHOSIID2. 31
less distinct ; hind wing with two indistinct spots from middle of the
costa; legs black-streaked.
Expanse | inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
LYCLENE DELICATA, n. sp.
Female. Pale yellow: fore wing with some small black subbasal
spots, a median transverse narrow line, a lunule at end of the cell, a
discal recurved series of small spots, one on each vein, and an outer
marginal series of very narrow black streaks, one on each vein: hind
wing with a marginal series of very narrow black streaks, one on each
vein. Underside with the outer marginal series of streaks and median
line as above, the median line extending partly across hind wing ;
femora, tibize, and tarsi with black terminal streak.
Expanse 1+ inch.
Darjiling (Grote). In coll. F. Moore.
Allied to Z. radians. Differs in having all the markings much
narrower, in addition to the median transverse line on the fore wing.
From L. prominens it also differs in the more slender markings, and
in the absence of the sharp dentate discal line.
LYCLENE PROMINENS, n. sp. (Plate III. fig. 3.)
Male and female. Upperside ochrey yellow: fore wing with four
black basal spots, a bent transverse subbasal line, a wavy median line,
a narrow oblique spot at end of the cell, an irregular discal band
with two sharply dentate points beyond the cell, and an outer mar-
ginal series of longitudinal streaks, one on each vein : hind wing with
a marginal series of short black longitudinal streaks. Two black
spots on middle of thorax. Underside marked as above, the median
and discal lines broader, the former extending across hind wing ;
abdomen with black bands beneath ; legs with black bands.
Expanse, g 1, 9 1,3, inch.
Cherra ; Khasia hills (G.-Austen). In coll. F. Moore.
LYCLENE ZEBRINA, n. sp. (Plate III. fig. 4.)
Male. Upperside yellowish white: fore wing with brown basal
linear spots, a median transverse line, a small spot at end of the cell,
and a wide outer marginal series of narrow streaks: hind wing with
a brown marginal band. Underside with broader markings than
above, the middle line crossing the hind wing ; marginal streaks in
place of the band.
Expanse 55; inch.
Calcutta District (Farr). In coll. F. Moore.
LycLene PALMata, n.sp. (Plate III, fig. 5.)
Female. Upperside pale ochrey yellow: fore wing with two bluish
basal spots, a transverse linear series of three spots, a median macu-
lated purple line, anda broad fan-shaped discal series of longitudinal
purple streaks decreasing in length to middle of hind margin: hind
wing with some short purple apical streaks. Underside marked as
32 MR. F. MOORE ON THE LITHOSIID®. [Jan. 15,
above, the basal spots less distinct; tibiee with black terminal
band.
Expanse 1,3, inch.
N.E. Bengal. In coll. F. Moore.
LYCLENE INTERSERTA, n. sp. (Plate III. fig. 6.)
Female. Upperside—fore wing ochrey yellow, with a short black
streak at the base, narrow subbasal transverse bent line, a discal line
which is convex anteriorly and bent posteriorly ; between these is an
oblique-angled line, the point being opposite to the angle of the sub-
basal line ; an upper and lower longitudinal line, the former along
the cell, the latter below it and furcate, the forks bent backward ;
also an outer marginal confluent looped line; cilia black, with white
border: hind wing pale yellow; cilia at apex black. Thorax and
head yellow, with black streaks; palpi and legs above blackish ;
abdomen paler yellow. Underside uniform yellow, markings in-
distinct.
Expanse 1,3; inch.
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Moore.
Near to L. euprepioides, Walk.
LycLENE OBSOLETA, n. sp. (Plate III. fig, 7.)
Female. Yellow: fore wing slightly ochreous yellow externally ;
with several purplish-brown subbasal spots, a deeply sinuous discal
band, and outer contiguous series of spots: hind wing and abdomen
yellow. Underside paler; markings on fore wing indistinct ; hind
wing with a short costal streak before the apex; fore tibiee with a
blackish terminal streak.
Expanse 5% inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
May be distinguished by the absence of the median transverse band
between the subbasal spots and sinuous band.
LYCLENE DISCISTRIGA, 0. sp.
Male and female. Fore wing yellow, with black basal speckled
spots, a median transverse recurved narrow band, an oblique spot at
end of the cell, a discal series of narrow longitudinal streaks, and a
marginal series of small spots: thorax black-spotted: hind wing
paler. Underside pale yellow; markings on fore wing very in-
distinct.
Expanse, ¢ 145, 2 1%; inch.
Darjiling. In coll. F. Moore.
Allied to the Bornean ZL. strigipennis, H. S. Lep. Spec. Nov.
f, 437.
LYCLENE INCONSPICUA.
Male. Pale ochreous red, semidiaphanous: fore wing with oblique
transverse basal and median indistinct dusky band, and discal streaks,
one along each vein and terminating in a marginal spot: hind wing
1878.] MR. F. MOORE ON THE LITHOSIIDE. 33
with a slight dusky apical streak. Underside—fore wing with the
discal markings confluent. Legs with dusky streaks,
Expanse +9 inch.
Northern India (Farr). In coll. F. Moore.
LYCLENE TERMINATA.
Male. Upperside pale yellow; fore wing with black irregular
basal streaks, an oblique recurved narrow median transverse line, an
oblique streak at end of the cell, and a discal series of narrow elon-
gated streaks, one on each vein and terminating ina spot at the outer
margin. Underside of fore wing as above. Legs black-streaked.
Expanse 49 inch.
Khasia hills (G.-Austen). In coll. F. Moore.
LycLene assamica. (Plate III. fig. 8.)
Male. Pale yellow. Upperside—fore wing with a purple-brown
basal spot, two subbasal transverse series of spots, a spot at end of
the cell, and a discal dentate band, each point having a terminal
spot which forms a submarginal series. Thorax black-spotted.
Expanse ;8; inch.
Dibrughur, Assam (A¢kinson). In coll. Dr. Staudinger.
Allied to both Z. undulosa and the Bornean L. cuneigera, Walk.
LYCLENE SPILOSOMOIDES, N. sp.
Male. Upperside yellowish white ; fore wing with two black spots
at the base and one on the costa, a subbasal and discal recurved series
of smaller linear spots, one on each vein. Body yellowish, tuft
brighter; a black spot on each tegula; palpi and band on lower
part of vertex brown. Underside uniformly paler, without mark-
ings.
Expanse 1,% inch.
N.W. India. In coll. F. Moore.
LycLeNneE INDISTINCTA, n.sp. (Plate III. fig. 9.)
Female. Fore wing dull yellow, with two very indistinct subbasal
curved series of pale brown spots, a bent discal series of spots, and
an outer series of short longitudinal streaks: hind wing and under-
side much paler ; fore tibise cinereous-brown above.
Expanse 144 inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
JRMENE, Walker.
Aimene, Walker, Catal. Lep. B. M. it. p. 542.
ZEMENE MACULIFASCIA, n. sp. (Plate III. fig. 10.)
Male. Fore wing white, crossed by six black maculated bands ;
the first basal; second curved; third and fourth discal, irregular,
and with dusky-brown suffused interspace; the other two marginal ;
two spots within the cell, the one at the end largest ; cilia spotted
near apex: hind wing ochreous-white. Thorax black-spotted ;
Proc. Zoo. Soc.—1878, No. III. a
34 MR. F. MOORE ON THE LITHOSIID. (Jan. 15,
palpi black ; fore legs black-streaked. Underside—fore wingbrown,
spots slightly visible; hind wing with a brown spot at end of the
cell, an indistinct submarginal fascia and spots on cilia near apex.
Expanse 1,4, inch.
Darjiling (Atkinson). In coll. Dr. Staudinger.
EMENE SINvATA, 0. sp. (Plate III. fig. 11.)
Male. Upperside ochreous white: fore wing with five narrow
black sinuous bands, a spot within and a lunule at end of the cell,
and an outer marginal row of small spots; cilia with a black line
and three widely separated spots: hind wing pale brownish-grey ;
cilia brown-lined ; a black terminal spot on tegule. Underside
brownish grey, markings on fore wing indistinct; hind wing with
pale brown spot at end of the cell, and a submarginal fascia ;
palpi black ; fore legs black-streaked.
Expanse 1 inch.
Cherra Punji (Atkinson). In coll. Dr. Staudinger.
EMENE SUBCINEREA, D. Sp.
Allied to 4. taprobanis ; the fore wing greyish-ochreous, with
markings more like those in 4. guttulosana, but smaller and
narrower ; hind wing and abdomen cinereous. Underside cinereous-
brown.
Expanse 7 inch.
N.W. India. In coll. F. Moore.
ARMENE MODESTA, 0. sp.
Male. Nearest to 4. guttulosana, Walk., from Bombay, but
smaller in size, darker in colour, and the hind wing pale fuliginous ;
the markings on the fore wing much smaller, the spot at end of the
cell narrow and lunular, and the sagittate mark below it nearly
obsolete.
Expanse 55; inch.
Formosa (Swinhoe). In coll. F. Moore.
ABMENE TENEBROSA, D. Sp.
Male and female. Fore wing hoary black, the grey scales pro-
minent, with some black spots on the costa within and below the cell,
and on the exterior margin ; hind wing and abdomen cinereous-grey.
Thorax hairy, black. Underside dark cinereous-grey ; apex of fore
wing ochreous; legs blackish above.
Expanse, ¢55;, 25% inch.
Bombay. In coll. F. Moore.
Allied to 4. sordida, Butler, Trans. Ent. Soc. 1877, p. 372, but
is a very much darker insect.
Setina, Schrank.
SETINA PUNCTILINEA, D. sp.
Male. Upperside—fore wing yellow; a black streak on base of
costal margin, a spot below it, two small very indistinct subbasal
1878. | MR. F. MOORE ON THE LITHOSIID£. 35
spots, one at end of the cell, and a more prominent marginal row of
spots: hind wing and abdomen pale yellow. Thorax black-spotted ;
legs black, streaked above.
Expanse +9 inch.
Ceylon. In coll. F. M. Mackwood and F. Moore.
Allied to the European S. irrorella.
SETINA DISCISIGNA, N. sp.
Female. Upperside pale yellow: fore wing tinged with ochreous
along hind margin; a black spot near base, a smaller spot at base of
costa, and a more prominent spot at end of the cell: hind wing suf-
fused with purplish-brown uear the apex and along abdominal margin.
Thorax and head yellow, both with black spots ; abdomen and legs
purplish-brown, tip yellow. Underside—middle of fore wing dusky
brown.
Expanse | inch.
Cherra (Atkinson). In coll. Dr. Staudinger and F. Moore.
SETINA NEBULOSA, N. Sp.
Female. Upperside—fore wing yellow ; with a black basal spot, a
broad irregular-bordered discal ochreous-brown band and confluent
subbasal irregular-shaped band: hind wing cinereous white, with
broad dusky indistinct submarginal fascia. Thorax yellow; abdo-
men cinereous-white. Underside and legs paler.
Expanse 1,2; inch. .
Darjiling (Atkinson). In coll. Dr. Staudinger and F. Movre.
SetinocHroa, Felder.
Setinochroa, Feld. Nov. Voy. iv. pl. 106.
SETINOCHROA AURANTIACA, n. Sp. (Plate III. fig. 12.)
Male. Upperside bright orange-yellow: fore wing with a black
basal spot, two subbasal, a large spot at end of the cell, anda
recurved submarginal series of four or five spots : hind wing ochreous-
brown externally. Thorax black-spotted ; abdomen greyish-brown,
tip yellow. Underside—middle of the fore wing ochreous brown,
base of costa and subapical streaks black ; hind wing black-bordered
at apex.
Expanse 19 inch.
Khasia hills (G.-dusten). In coll. F. Moore.
Allied to S. infumata, Feld. Nov. Voy. iv. pl. 106. fig. 16, but
differs in being of a different yellow in colour, in the fore wing having
the subbasal spots, and the hind wing having only a marginal blackish
border.
SETINOCHROA PALLIDA, 2. sp.
Male. Upperside—fore wing pale yellow, with small black spot at
base, one at end of the cell, and a submarginal series of spots: hind
wing pale ochreous-brown externally. Underside—fore wing pale
ochreous-brown in the middle ; cell and apical spots very indistinct.
Expanse ;"; inch.
3%
36 MR. F. MOORE ON THE LITHOSIIDZ. [Jan. 15,
Manpuri, N.W. India (Horne). In coll. F. Moore.
Differs from S. infumata in the wings being narrower, and of a
paler colour both above and beneath.
SETINOCHROA POSTICA, 0. Sp.
Male. Upperside bright ochreous-yellow : fore wing with a small
black spot at base, a prominent spot at end of the cell, and a recurved
submarginal series: hind wing slightly brownish-ochreous, with a
distinct black streak at end of the cell. Underside—fore wing
ochreous-brown in the middle, with black apical spots; cell-streak
in hind wing.
Expanse 14 inch.
Dhurmsala, N.W. Himalayas. In coll. F. Moore.
Nuparis, Haw.
Nudaria, Haw. Lep. Brit. p. 156.
NvuDARIA FASCIATA, 0. Sp.
Male and female. Pale yellow, diaphanous: fore wing with a
broad purple-brown band from the costa, decreasing to middle of
hind margin, and lunular streaks on exterior margin; hind wing
paler, slightly suffused with purple-brown at the base. Body yel-
lowish-white ; legs purple-brown above.
Expanse +9 inch.
Darjiling (Atkinson). In coll. F. Moore.
Allied to N. margaritacea, Walk. ; but that species differs in
pats the fore wing crossed by four distinct irregular sinuous
bands.
EXPLANATION OF THE PLATES.
Puate I.
. Paraona splendens, p. 8.
. Sidyma apicalis, n. sp., p. 9.
. Mahavira flavicollis 8, un. sp., p. 11,
. Hesudra divisa 3, n. sp., p. 12.
. Vamuna maculata 3, n. sp., p. 10.
Chryseglia ferrifasciata 3, un. sp., p. 13.
Gandhara serva, p. 15.
Tarika varana, p. 15.
. Brunia antica, p. 15.
10. Ghoria albocinerea, n. sp., p. 13.
11. Vamuna bipars 3, n. sp., p. 10.
12. Churinga rufifrons 3, n. sp., p. 10.
13. Collita parva, un. sp., p. 16.
14, Katha terminalis, n. sp., p. 17.
nigrifrons, n. sp. p. 17.
16. Stmareea basinota, p. 14.
Prate II.
. Chrysorabdia viridata 8, p. 19.
Capissa innotata g, p. 19.
pallens, n. sp., p. 19.
—— fasciata 3 n. sp., p. 20.
. Tegulata basistriga, n. sp., p. 22.
protuberans 2, n. sp., p. 28.
—
Fig.
CO OAD CrP OD tO
duxt TaPuUe LT
1878.] | MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 37
Fig. 7. Systropha auriflua, un. sp., p» 18.
8. Dolgoma brunnea, n. sp., p. 20.
9. Mithuna quadriplaga, n. sp., p- 21.
10. Cossa quadrisignata, n. sp., p. 21.
brunned, n. sp., p. 22.
12. Ranghana punctata, n. sp., p. 22.
13. Teulisna tenuisigna, n. sp., p. 24.
14. Macotasa biplagella 3, n. sp., p. 25.
15. Zadadra distorta 3, p. 25.
16. Prabhasa venosa gn. sp., p. 26.
flavicosta 2, n. sp., p. 26.
18. Gampola fasciata S$, n. sp., p. 27.
Prats III,
1. Lyclene rubricosa, n. sp., p 30.
2. —— radians, n. sp., p. 30.
3 prominens, n. sp., p. dl.
4 zebrina, n. sp., p. dl.
5. —— palmata, nv. sp., p. 31.
6
ul
8
9
. — meterserta, n. sp., p. 32.
obsoleta, n. sp., p. 32.
assamicd, 0. sp., p. 30.
indistincta, n. sp., p. 30.
10. Aimene maculifascia, un. sp., p. 33.
11. —— sinuata, n. sp., p. 34.
12, Setinochroa aurantiaca, n, sp., p. 35.
13. Bizone bellissima 3, n.sp., p- 27.
coccinea S, un. sp., p. 28.
15. Barsine flammealis, n. sp., p. 28.
gloriosa, n. sp., p. 29.
17. —— inflexa, n. sp., p. 29.
18, —— flavivenosa, n. sp., p. 30.
2. On Birds collected in Costa Rica.
By Apotprue Bovcarp, C.M.Z.S.
[Received Nov. 15, 1877.]
(Plate IV.)
Costa Rica, the most southern of the five Republics known together
under the name of ‘‘ Central America,”’ is said to contain 150,000 in-
habitants, of which 100,000 live in the valleys of San José and Car-
tago. The remainder of the population is widely distributed in small
villages, some of them lying at great distances from each other.
Thence the difficulty of travelling in the interior of the country.
I arrived at Puntarenas, the Pacific port, on the 29th of Decem-
ber 1876, and remained in the country till the 30th May 1877—in
all, five months.
My head quarters were San José, the capital. It is situated in the
centre of a very fertile valley, about midway between the Atlantic
and the Pacific, at the altitude of 4100 feet. It is said to contain
15,000 inhabitants. My intention was to take a house; but the
rent, furniture, and servants were so dear that I stopped all the time
at the Hotel Victor, where I was recommended to go.
38 MR, A. BOUCARD ON BIRDS FROM COSTA RICA. [Jan. 15,
My hunting-ground during the first weeks was in the valley of
San José; but I was rather disappointed, birds being very scarce,
which was due chiefly to the poverty of the vegetation, all the soil
for many miles having been turned into coffee-plantations. The
only places for hunting were the roads from San José to the towns
and villages of Heredia, Alajuela, Tres Rios, Desamparados, &c.
On the trees and shrubs surrounding the coffee-plantations small
flocks of birds were flying about. These flocks consisted of the
following species—Pyrgisoma cabanisi, Zonotrichia pileata, Buarre-
mon chrysopogon, Saltator grandis, Catharus melpomene, Turdus
grayi, Cyclorhis flavipectus, Hlainea pagana, Myiarchus crinitus,
and several other species of ‘l'anagride, Fringillidz, and Tyrannide.
Among the Trochilide Pyrrhophena riefferi, Saucerottia sophie,
and Chlorolampis osberti were the only three species to be seen at
that time.
In the gardens and maize-fields several species of Spermophila were
found feeding on seeds.
In the barrancas (deep ravines occasioned by the large quantity of
water which falls in the rainy season) the vegetation was more
varied, and several species of Columbidee were met with. In all
parts of the valley you always find the same species.
At the end of January, I made an excursion to the valley of San
Carlos, in company with MM. Koschny and Meil de Fontenay, who
proved to be very good travelling companions. The valley of San
Carlos divides the Republic of Costa Rica from that of Nicaragua.
It lies on the Atlantic slope, and extends as far as the river San
Juan, which is the boundary of the two countries. It is quite a
tropical climate, and considered very unhealthy. The forests are of
the most magnificent description. There are scarcely any roads or
inhabitants, except four or five settlers who are trying to raise cattle,
and some few Nicaraguans who are engaged in extracting the juice
of the india-rubber tree (Siphonia elastica).
To reach San Carlos we had to pass Sarzero and Laguna, two
small villages lying at the summit of the Cordillera, near the volcano
of Poas. Thence we had to descend gradually about 5000 feet
through a very picturesque country.
We arrived the third day, at twelve, at the rancho where we
intended to stop. From this rancho you can reach Greytown by
water in little more than half a day. The return takes about six
days, as you have then to go against the current.
The animal life is prodigious; flocks of birds are, constantly
passing to and fro. Early inthe morning and at sunset large quan-
tities of Howling Monkeys (Mycetes palliatus) are heard all around.
They make a fearful noise; and one who has not heard them before
may well fancy that the forest is thronged with Jaguars. These
last-mentioned animals used to be very common; but now they are
rather scarce, having been much persecuted.
Large flocks of Ara macao, Chrysotis farinosa, Pionus senilis,
Conurus astec, and other species of Parrots are constantly passing in
all directions.
1878.] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 39
This excursion lasted a fortnight. During that time I made a
good collection of bird-skins, amongst them a female of the rare
Cotingine bird, Carpodectes nitidus, quite new to science ; also many
other good species, as Myrmotherula melena, Urospatha marti,
Prionirhynchus carinatus, Crypturus boucardi, &e.
All the time I had to work at a great disadvantage. On the trunk
of a tree I prepared my skins, with an empty box for a seat. Pigs
and ants were always ready to do mischief; and I lost several good
specimens of birds by pigs devouring them when they were drying
in the sun.
My other excursions were to Cartago, Aguas Calientes, Navarro,
Orozi, the Volcano of Irazu, Candelaria, Naranjo, &e.
Cartago is a town of about 10,000 inhabitants. It was formerly
the capital; but owing to the frequent earthquakes felt there, and
the great destruction of property occasioned by the same, the govern-
ment was transferred to San José. The town is built at the base of
the Volcano of Irazu, in a beautiful valley which varies from three
to six miles in width and is about ten in length.
Aguas Calientes is a very prosperous village, only two miles and
a half from Cartago. Although at such a short distance, it is much
rae, being well sheltered from the north wind, and about 250 feet
ower.
A few miles further, going north-east, are the small villages of
Navarro and Orozi, the localities for Phainoptila melanoxantha, Chlo-
rophonia callophrys, Geotrygon costaricensis, Tetragonops frantzii,
&c.; and if you still go further north, you reach Tucuriqui, the
locality where Arcé procured the type-specimen ( ¢ ) of Carpodectes
nitidus. All these localities are from 500 to 2500 feet lower than
Cartago, and the fauna is very rich.
Naranjo is the name of a very important hacienda, belonging to
to the Tenoco family. Lately a small village of the same name has
been formed by the workmen of the hacienda. It stands on the
road from San José to Limon (the Atlantic port).
This locality is very good for a naturalist, lying between the
voleanoes of Irazu and Turrialba. In a few hours you can change
your altitude from 1500 to 8000 feet or more if you choose; con-
sequently there is a great variety of species to be got. The forests
are luxuriant, due chiefly to the climate, which is very wet. It rains
nearly all the year round. I found there many of the same species
met with at San Carlos.
From (artago I made several excursions to the Volcano of Irazu.
Up to the altitude of 6000 feet you only see maize- and wheat-
plantations, occasionally some potreros (plains) where they fatten
cattle, until you reach a small village called Potrero cerrado, where
they cultivate potatoes extensively. A little further on begins the
forest, where a species of oak tree is very abundant. Then the
fauna changes entirely. You do not meet any more with the species
of birds found in the valley, but such others as Ptilogonys caudatus,
Setophaga aurantiaca and S. torquata, Parula gutturalis, Pezopetes
capitalis, &e. Further on, at the altitude of from 7000 to 8000 feet
40 MR. A. BOUCARD ON BIRDS FROM CosTa RICA. [Jan. 15,
the forest is more dense, and the animal life is very abundant. This
is the chief locality for Pharomacrus cestaricensis, Panterpe in-
signis, Dorycha bryante, and Turdus nigrescens.
At the altitude of 8000 feet the forest ceases altogether, and you
walk on scorize and ashes sent from the voleano of Irazu a long time
ago. Many small trees and aromatic plants grow in the soil up to
the summit of the mountain.
This is the locality for Selasphorus flammula and S. scintilla, which
are seen sucking the flowers of mistletoes and heaths. Here also I
met with my new species Zonotrichia vulcani. I reached the summit
of the volcano, and slept there. On the other side, looking north-
west, is the crater, which I have thoroughly explored.
Rancho redondo is a farm situated on the western slope of the
Volcano of Irazu. It is also a good locality for Phainoptila melano-
xantha, Eugenes spectabilis, Chasmorhynchus tricarunculatus, and
other rare species.
Atenas is about thirty miles west from San José, on the road to
Puntarenas.
San Mateo is on the Pacific slope, at the foot of the Aguacate moun-
tain (rich in gold-mines), about thirty-eight miles from the port. The
climate is quite tropical. Some of the species found at San Carlos
are also met with here. It is also a rich locality for reptiles and
insects.
Barranca is only about half a mile from the sea, and ten miles
from Puntarenas. The village is built near the river of the same
name.
Puntarenas is the port. Its name is very appropriate, as it stands
on a sandy peninsula extending for about six miles. On one side is
the sea, on the other a river; sometimes the width of this isthmus
between the river and the sea is only fifty yards, and never more
than one mile. It is excessively hot and unhealthy for a new comer.
The vegetation is poor; but four miles from the town is a fine forest,
where animal life is plentiful.
Altogether Costa Rica has been well explored by MM. Warzewicz
Frantzius, Hoffmann, Ellendorf, Carmiol, father and son, Endrés,
Arcé, Zeledon, Cooper, and others.
Up to this day 520 species of birds (some doubtful) are recorded
as having been collected in that country ; and I feel well satisfied
when I think that in a country so well worked up I have been able
to obtain several new species and some great rarities—such as both
sexes of Phainoptila melanoxantha, a new genus and new species just
described by Mr. Osbert Salvin, Carpodectes nitidus ( 9 ), Catharus
Srantzii, Catharus gracilirostris, Turdus nigrescens, Turdus obso-
letus, Parula gutturalis, Pezopetes capitalis, Melozone leucotis,
Eugenes spectabilis, Selasphorus scintilla, Selasphorus flammula,
Oreopyra hemileuca, Oreopyra cinereicauda, Panterpe insignis,
Conurus hoffmanni, Geotrygon costaricensis, Dendrortyx leucophrys,
Crypturus boucardi, &e. &e.
The total number of species which I collected is about 250; but
I worked very hard to obtain this result, and until the time of my
1878.] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 4]
departure I did not lose a single day, and often I had no repose
from morning till night. I could easily have obtained a larger
number of specimens; but I did not collect very well-known and
common birds which were already well represented in my collection.
I consider Costa Rica one of the worst places for making col-
lections, in consequence of the difficulty of transport, the bad
roads, and the great expense which these deficiencies occasion.
You must carry every thing with you, live and sleep as you can and
where you can, sometimes in the most miserable huts, sometimes in
the forest.
At first I was not able to find help at any price. A few hunters,
willing to work for me at high pay, made such bad skins that I
was obliged to dismiss them.
Others to whom I offered good prices for birds in the flesh never
came again after I gave them powder and shot. At last, a little
before my departure, I met with two good hunters, whom I
employed a few days, and did very well with them; but it was
rather too late, the time of my departure having arrived, and the
best season for birds being over.
The classification which I have followed in the following list is
that of my ‘ Catalogus Avium.’
Ordo Crypruri.
Family TrnamMipz.
1. NorHocEeRcus BONAPARTI, Gray.
Tinamus frantzii, Lawr.
Native name “Gallina del monte.’’ Only one specimen procured,
in May 1877, at Rio Navarro, at the base of the Candelaria Moun-
tains, twelve miles from San José.
This seems to bea rare species ; the typical specimen of 7’. frantzi,
Lawr., was collected at Cervantes by J. Zeledon. Cervantes is not
far from Rio Navarro.
Like all Tinamide, it is found in the dense parts of the forest,
where it feeds on insects and seeds. They can be easily detected by
the noise they make when scratching the ground in search of food.
They go in pairs, repeatedly calling one another. When fearing
danger they run with much rapidity.
2. CrYPTURUS BOUCARDI, Scl.
The type of this species was from Mexico, and not from Guate-
mala as stated in Gray’s ‘ Hand-list of Birds.’ As well as I can
remember, the first specimen (the type of this species) was obtained
by me at Playa Vicente, a small village up the river Papaloapam,
the mouth of which is at Alvarado, on the Atlantic, between Vera
Cruz and Minatitlan.
As far as I know, all specimens of this bird have been obtained
in the tropical forests a few hundred feet above the sea-level ; but
it is an interesting fact that this species has been obtained also in
Guatemala and Costa Rica.
42 MR, A. BOUCARD ON BIRDS FROM CosTA RICA. [Jan. 15,
Only one specimen ( ) collected, at San Carlos, where it was
rather abundant in February. They are rather easy to detect, in
consequence of a plaintive cry which they repeat frequently, as if
calling one another.
The flesh is white and delicious to eat.
I have also one specimen, obtained at San Carlos, which is much
darker than C. bowcardi. It has not any of the black and yellow
spots, either on the back or on the secondaries and tertiaries, which
are so conspicuous in the former. Instead of that the secondaries
are nearly black, bordered with rufous grey, the tertiaries are rufous
grey, the back is rather dark rufous, darker on the rump, the
upper tail-coverts are rufous, with an indication of yellow and black
spots, as in C. boucardi. On the underside the abdomen is nearly
of the same grey colour as the breast, slightly rufous on each side.
Ordo GaLuInz.
Family Cracip2.
3. PENELOPE PURPURASCENS, Wagl.
Native name “ Faisan.”’
Hab. Volcan de Irazu.
Rather common in the forests on the slopes of the volcano. It is
also found in San Carlos, showing that it lives indifferently in low or
high altitudes.
4. CHAMZPETES UNICOLOR, Saly.
Hab. Volcan de Irazu.
They are in great demand for food in San José, where they sell at
from four to eight shillings.
Subfamily Cracinz.
5. Crax GLOBICERA, L.
Native name “ Pavo.”’
Hab. San Carlos, Volcan de Irazu, Naranjo.
Rather common.
Subfamily OpontTorHoRIN&.
6. OponrorHORus GuTTATUS, Gould.
Native name ‘‘ Perdiz.’’
Hab. Curridabat, near San José.
Two specimens, one male and one female, killed in March.
7. DENDRORTYX LEUCOPHRYs, Gould.
One specimen, from the Volcano of Irazu, May. Killed at the
altitude of 7000 feet. It was scratching the ground in search of
food.
8. Ortyx LEYLANDI, Moore.
Valley of San José. Common in the coffee-plantations during the
1878. | MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 43
rainy season, from May to December, rare in the other months ; then
it disappears completely from the valley. They are also met with
in small flocks in the plains. They run very: fast.
Ordo CorumBz.
Family CoLUMBID4.
Subfamily CoLuMBIN2.
9. CoLUMBA ALBILINEATA, Gray.
Several specimens from the Volcano of Irazu, May. Is found in
flocks of ten or twelve, feeding on seeds. I have seen them coming
from all directions to drink at a spring of mineral water near Desam-
parados.
10. CotumBA sUBVINACEA, Lawr.
Only one specimen, from Candelaria, May. Very rare.
11. ZENAIDURA CAROLINENSIS, L.
Several specimens, from San José, January to May.
Subfamily Gourin#.
12. CHAM#PELIA PASSERINA, L.
Common near San José, January to May.
13. CHAMHPELIA RUFIPENNIS, Gray.
Very abundant at Puntarenas in May.
14, Leproprita vERREAUXI, Bp.
Several specimens from San José, January to March.
15. Leproprtiua cassini, Lawr.
Only one specimen, from San Carlos, February. Inhabits the
forest. I also found two eggs of this species on a small tree. They
are uniform white. The male was sitting on the eggs.
16. GEOTRYGON MONTANA.
San Carlos, February.
17. GEOTRYGON COSTARICENSIS, Lawr.
Several specimens killed on the mountain of Candelaria, at the
altitude of from 3000 to 4000 feet.
Feeds on seeds, and very likely also on insects. The crop of one
of the specimens dissected, and which I have here, is full of small
seeds. I have always seen them on the ground, going singly. The
sexes are exactly alike.
18. GEOTRYGON CHIRIQUENSIS, Sclat.
Geotrygon ceruleiceps, Lawr.
Native name ‘“‘ Paloma del Monte.”
Volcano of Irazu and Candelaria.
This is a scarce species in Costa Rica, Like the former, it has the
44 MR. A. BOUCARD ON BIRDS FROM Costa RICA. [Jan. 15,
same habits as the Tinamidz and Odontophorinee of going about on
the ground. I have never seen it perched.
Ordo LimicoLz.
Family CHARADRIIDA.
19. Oxyecuus vocireRvus, L.
Valley of San José.
Excessively common in the plains, where they are seen constantly
running about searching for food, and even catching grasshoppers
and small insects on the wing. When surprised they run very fast,
at the same time erying, as if to advise one another of some danger.
Family ScoLoracip2.
20. TRINGOIDES MACULARIUS, L.
Several specimens shot in the valley of San José. They are found
along the streams, flying from one stone to another, or searching for
food close to the water. When surprised they take to flight, emitting
a sharp cry, similar to pit, pit, pit.
Ordo HERODIONES.
Family PLATALEID.
21. PuaTauea agasa, L.
One specimen near San Jos¢, April.
Ordo ANSERES.
Subfamily ANATINA.
22. DENDROCYGNA AUTUMNALIs, L.
Native name “ Pato.”
Common at the laguna of Cartago, where the division of the waters
takes place, either for the Atlantic or the Pacific. ,
Ordo AccIPITREs.
Family FaLconrip2&.
23. MICRASTUR SEMITORQUATUS, Vieill.
One specimen, killed in the forests of Candelaria Mountains, May.
24. Burro LaAtissimus, Wils.
Buteo pennsylvanicus, Wils.
One specimen, killed at San José in January.
25. URUBITINGA GHIESBREGHTI, Du Bus.
One specimen, from Naranjo, April. Rare.
26. TuHrasatrus HARPYIA, L.
Native name “ Aguila.”
I have seen one specimen alive in San José. It was caught in the
1878.] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 45
Candelaria Mountains. It was kept in a large cage, and seems quite
at home. When you approach it, it erects its crest, and presents
then a very imposing sight.
27. ELANOIDES FURCATUS, L.
One specimen from Naranjo. I saw several in a place cleared for
sowing. They sat motionless for hours at the top of the highest
trees, probably watching for their prey.
28. Lepropon uNncINATUS, Temm.
Native name “ Gavilan.”
One specimen, from Naranjo, April. This bird has a strong cry,
which sounds like ova, ova, oua, repeated several times in succession.
It resembles very much the cry of the Peacock when asking for rain
(as vulgarly said). It is so loud that you can hear it at a long
distance. The capture of my specimen was due toit. It was perched
on the top of a high tree in the forest ; and if it had not been for its
ery, I donot think I should have detected it. Several of these birds
were at about half a mile one from the other, and crying in succes-
sion ; and it occurs to me that each one takes possession of a certain
part of the forest and admits of no intruder.
29. CERCHNEIS SPARVERIUS, L.
Common in all altitudes, January to May.
Family VuLtruripz.
30. CATHARTES PAPA, L.
Native name “ Rey de los Zopilotes.”’
Seen at San Mateo in May.
31. CATHARISTES ATRATUS, Barth.
Native name “ Zopilote.”
Common everywhere all the year round.
32. Ginops aura, L.
Native name ‘‘ Zopilote de cabeza colorada.”
San José, January to May.
Ordo SrricsEs.
Family BuBonipz.
33. Scops BRASILIANUs, Gm.
Native name ‘‘ Lechuza.”
One specimen from San José. It was perched on a small tree in
company with two adult specimens. I killed the young and the
female with the same shot.
34. LopHosTRIX STRICKLANDI, Sclat. & Saly.
San Carlos, February.
46 MR. A. BOUCARD ON BIRDS FROM Costa RICA. [Jan. 15,
Ordo Psrrract.
Subfamily Arainz.
35. Ara Macao, L.
Native name ‘ Guacamaya.”
Common at San Carlos. Every morning they fly about in large
flocks. In the daytime you can see them in the forest eating fruit ;
they are easily detected by the noise they make, and by the rejected
pieces of fruit constantly falling down from the trees on which they
are perched.
36. ARA mitiITARIs, L.
San Carlos, Sarzero, February.
37. ConURUS HOFFMANNI, Cab.
Native name “‘ Perico.””
Several specimens killed at the foot of the Candelaria Mountains.
I have also seen small flocks at Aguas Calientes, near Cartago. When
flying they appear excessively beautiful: this is due principally to
the conspicuous colour of their wings.
38. Conurus astec, Souancé.
Native name “ Perico.”’
One specimen killed at San Carlos, amongst a large flock eating
fruit and seeds in the forest, February.
39. BroroGerys Tovi, Gm.
Native name “ Periquito.”
Common at San Carlos, February.
40. Pronus sENILI1s, Spix.
Native name ‘‘ Cotora.”’
Common at San Carlos in February. Found in company with
Conurus astec.
41. Curysotis rartnosa, Bodd.
Chrysotis pulverulenta, Gro.
Native name “ Loro.”
Common at San Carlos in February.
Ordo Coccyees.
Subordo ZyGopacTyLa.
Family RampHastip@.
42. RAMPHASTOS TOCARD.
Native name “ Pico de canoa.”’
Common at San Carlos in February.
43. PrEROGLOSSUS TORQUATUS, Gm.
Common at San Carlos in February.
1878.] | MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 47
44, PreroGLossus FRANTZII, Cab.
Common at San Carlos in February.
45. SeELENIDERA SPECTABILIs, Cass.
Naranjo, April.
46. AULACORAMPHUS C@RULEOGULARIS, Gould.
Naranjo, April.
All these species are found in the forests in small flocks of from
ten to fifteen. They live chiefly on fruit.
Family Carironip2&.
Subfamily Carrronin&.
47. TETRAGONOPS FRANTZII, Sclat.
Several specimens from Navarro, May. Found in the forests near
the streams.
48. Capito BourcieERt, Lafr.
Two specimens, from Orozi and Navarro.
Family Bucconip2.
49. Mawtacorpriua PANAMENSISs, Lafr.
Two specimens, male and female, killed at San Carlos in February.
Always seen in the forests.
Family GaALBULIDX.
50. GALBULA MELANOGENIA, Sclat.
Common at San Carlos in February.
Family Cucu.ip.
Subfamily CroropHAGIN.
51. CRoTOPHAGA SULCIROSTRIS, Sw.
Very common in the valley of San José on the edges of the coffee-
plantations. They go in flocks of from ten to fifteen, flying from one
bush to another. They do not notice the presence of man, and are
commonly called “‘Garapatero”’ (eater of Garapatas, a common para-
site which lives principally on cattle, and also on men). They are
often seen on the back of animals, eating the insects which infest
them. The cattle seem to enjoy very much the close company of
this auxiliary, which does such good service for them. I have one
egg of this species, which is of the size of a Pigeon’s, all white.
Subfamily Coccyzinz.
52. Coccyzus minor, Gm.
One specimen, from Puntarenas, May. This is the first time that
this species is recorded from Costa Rica.
48 MR. A. BOUCARD ON BIRDS FROM Costa RICA. [Jan. 15,
53. PiayA MEHLERI, Bp.
Several specimens from San José. It is called commonly “ Bobo
de pecho aplomado.” ‘The expression “ Bobo” means imbecile,
because it is very tame, and you can shoot it very easily. I have
always seen them on the edges of the coffee-plantations.
54. Morococcyx eryTHropyGus, Less.
One specimen, from Atenas, May. It was on the side of the road,
where I observed it for several minutes before I killed it. It
jumped from one stone to another, then came down and scratched
the ground for insects, then perched on a low branch, and so on,
without taking any notice of me, although I was only a few yards
distant.
Subordo HETERODACTYL.
Family TROGONID2.
55. PHAROMACRUS MOCINNO, Lave, var. costaricensis, Gould.
Native name “ Quetzal.”
A large number of specimens, from the Volcano of Irazu, Navarro,
Candelaria, Naranjo, in fact from all the mountains around San
José and Cartago. Very difficult to get. It is only by imitating
the cry of the female that the men of the country can get a shot
at the males. They keep in the denser parts of the forest, chiefly
in the places called “‘barrancas”’ (deep ravines, very difficult of
access). They feed on seeds and fruit. In May (the season of
acorns) they are found in small flocks of ten to twelve on the robble
trees, and can then be easily killed. During the breeding-season
they go in pairs, male and female, constantly calling one another.
For further details on the habits of this bird, see part xi. of ‘Or-
nithological Miscellany’ ’.
56. TrRoGon MASSENA, Gould.
San Carlos, February. I have always seen these birds in the
forests. They perch high, and remain a long time stationary.
57. TrRoGon PUELLA, Gould.
Several specimens killed at Navarro and in the Candelaria Moun-
tains in May.
Subordo ANISODACTYL2.
Subfamily ALCEDININ &.
58. CmerYLE CABANIsI, Tsch.
Native name “ Martin pescador.”’
Common everywhere, January to May.
Family Momorip.
59. Momorus LEssonlI, Less.
Four specimens, from San José, January, March. It is vulgarly
? T have sent a full account of my observations on this species to Mr. G. D.
Rowley, which he has published in his interesting work.
1878. | MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 49
called ‘ Bobo,”’ because it is quite tame and easy to kill. I saw
these birds chiefly in the coffee-plantations near the streams. Its
cry has rather a hollow sound, sometimes like the word ‘‘mou mot,”
or “mot mot,” from which the name of the genus is derived.
60. UrospaTHa MARTH, Spix.
Several specimens from San Carlos, killed in February. Always
seen in the forest, where they make a fearful noise calling one
another. Its note has some resemblance to the cry of the former
species, but is much stronger and more hollow. They are easily
killed at first, but they are wilder after a few shots. I have met
with them in small flocks.
61. PrRioNIRHYNCHUS CARINATUS, Du Bus.
One specimen, a female, collected at San Carlos, February. Rare.
Goes by pairs in the forest. This is the first time that this species
has been found so far south.
62. EUMOMOTA SUPERCILIARIS, Sw.
One specimen, from Barranca, near Puntarenas, May. Like Mo-
motus lessoni and other Momotide, it will remain for a long time
perched on the same branch, and the only sign of its being alive is
the frequent movement of its tail.
Ordo Prct.
Family Picrpx.
63. Dycriopicus JARDINEI, Malh.
Native name ‘‘ Carpintero.”
Two specimens from the Volcano of Irazu and from Navarro, Jan-
uary and May 1877.
64. CenTURUS HOFFMANNI, Cab.
Two specimens, from San José, January, and Cartago, ‘April;
killed in the coffee-plantations. Also one specimen from Puntarenas
which I cannot possibly separate from this species, although it is a
much smaller bird.
65. CenTURUS PUCHERANTI, Malh.
Two specimens, one from San Carlos, February, the other from
Naranjo, April.
66. MELANERPES FORMICIVORUS, Sw.
Several specimens from the Volcano of Irazu, May 1877. I have
seen some also at Naranjo.
Proc. Zoou. Soc.—1878, No. IV. 4
50 MR. A. BOUCARD ON BIRDS FROM CosTA RICA. [dan. 15,
Ordo PassEerREs.
Subordo Oscrnegs.
OSCINES DENTIROSTRES.
Family Turpvip&.
67. Turpus Gray, Bp.
Native name “ Giguira.”’
Several specimens from San José, where they are found in large quan-
tities during the fruit-season. They are particularly fond of a small
fruit called in the country ‘“‘cereza.” My specimens differ widely
from the ordinary form of T. grayi in their general coloration :
instead of being rufous, they are ashy grey with a rufous reflection,
but so different in coloration that it is impossible to mistake one for
the other; the sides of the abdomen are of the same colour as the
underside plumage, whereas in T. grayi they are rufous, nearly red.
I have two eggs of this bird, found in one nest built in the
centre of an aloe growing on the road. They are green, with rufous
spots.
68. Turpus PLEBEIUS, Cab.
One specimen, from Navarro, April.
69. TuRDUS NIGRESCENS, Cab.
Several specimens from the Volcano of. Irazu.
I never saw this species anywhere else. They are found in small
flocks in the oak-tree region, at the altitude of from 6000 to 10000
feet. I found a nest containing two eggs. They are of a pale
uniform green.
70. TuRDUS LEUCAUCHEN.
Only one specimen, from Orozi, May.
71. Turpvus oBsoLetus, Lawr.
Only one, female specimen, from Naranjo, April. This species
seems to be rare. It lives in the forest.
72. CATHARUS MELPOMENE, Cab.
Native name “ Paxaro ingles” (English bird) ; I could not learn
why.
Many specimens, from San José and Cartago, where they are found
on the edges of the coffee-plantations. They are often seen on the
ground searching for insects. They sing beautifully.
73. CATHARUS FRANTZI, Cab.
Two specimens from Navarro, May. Very rare.
74. CATHARUS GRACILIROSTRIS, Salyv.
Two specimens from the Voleano of Irazu, April. Very rare.
1878.] | MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 51
Family Syiviip#.
75. MyItADESTES MELANOPS, Salv.
Several specimens, from Navarro, May 1877. Always found in
the forest, where they are easily detected in consequence of their
beautiful and clear song. It is short; but nothing could be more
perfect in tone. Occasionally they are brought alive for sale to
San José; but they are not easily domesticated.
Family TROGLODYTID£.
76. TROGLODYTES INTERMEDIUS, Cab.
Troglodytes hypaédon, Sclat.
Many specimens, from San José, Cartago, and Naranjo. It is
vulgarly called ‘¢ Salta pared,” which means “ jump-the-wall.” They
are common in the gardens from January to May.
77. THRYOPHILUS MODEsTUS, Cab.
Many specimens, from San José, Cartago, and San Mateo. _I have
always found them on the edges of the coffee-plantations. January
to May.
“78. THRYOPHILUS CASTANEUS, Lawr.
Several specimens, from San Carlos ; killed in February. Found in
. the forests near the habitations.
“79. THrRYOPHILUS PLEUROSTICTUS, Sclat.
Two specimens, from Puntarenas, May. Killed in the forest close
to the sea.
80. THRYOPHILUS THORACICUS, Salv.
One single specimen, from San Carlos, February. Rare.
81. CAMPYLORHYNCBUS ZONATUS, Less.
Only one specimen, killed at San Mateo in May. They go in
small flocks, making a great noise as they fly from one tree to another
in the open spaces in the forest.
82. HeniIcorHINA LEUCOSTICTA, Cab.
Several specimens, from San Carlos and Orozi. Always found in
the forest on the ground.
83. HeNnIcORHINA LEUCOPHRYS, Tsch.
Only one specimen, from Navarro, May.
Family Mnrori,tip2.
84. S1uRUS NOVEBORACENSIS, Gm.
One specimen, from San José, March.
85. Srurus aurocaPILuus, L.
Two specimens, from San José, January and March.
| 4*
52 MR. A. BOUCARD ON BIRDS FROM Costa RICA. [Jan. 15,
86. Mniotiuta vaArtA, L.
San Carlos, February.
87. ParuLA GUTTURALIS, Cab.
Several specimens, from the Volcano of Irazu, April and May.
Killed on the oak trees at an altitude of 6000 feet.
88. HELMINTHOPHAGA PEREGRINA, Wils.
Several specimens, from San José, January, March.
89. DeNDRG@CA BLACKBURNIA, Gm.
One specimen, from Naranjo, April.
90. DENDR@CA PENSYLVANICA, L.
San Carlos, February.
91. DeENpDR@CcA &sTIVA, Gm.
Several specimens, from San José, January, March.
92. DENDRECA VIEILLOTI, Cassin.
One single specimen, from Puntarenas, killed in May on the
mangrove trees, in close proximity to the river.
93. GroTHLYPIs TRICHAS, L.
Several specimens, from San José and Cartago, January to May.
94, GroTHLYPIS POLIOCEPHALA, Baird, var. caninucha, Ridgw.
Only two specimens, from San José, killed in January and
February.
95. Myoptocres pusiLuus, Wils.
Several specimens, killed at San José in January.
96. BASILEUTERUS MESOCHRYSUS, Sclat.
Many specimens, from San José and Cartago, January to May.
97. BASILEUTERUS CULICIvORUS, Licht.
San José, January to May.
98. BASILEUTERUS UROPYGIALIs, Sclat.
Only one specimen, from San Carlos, killed in February. Itis a
very rare species, living along the streams in the forest. It never
rests, but is constantly going up or down the river from one place to
another, sitting on stones or scratching the ground. I saw several
specimens in the same place, but could only get one, as it is a very
wild bird.
99. SETOPHAGA RUTICILLA, L.
Several specimens, from San José and Cartago, January and April.
1878. ] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 53
100. SrropHAGA AURANTIACA, Baird.
Several specimens from Cartago and Naranjo. ‘This species in-
habits the forests; in the middle of the day they go along the
streams, flying from one tree to another, feeding on insects. I have
always seen them going in pairs.
101. SerorpHaGa ToRQUATA, Baird.
Several specimens of this rare species, all of them killed in the
forest on the Volcano of Irazu. Always seen in pairs. The female
differs very little from the male.
Family VirEONID.
102. VrrEOSYLVIA FLAVOVIRIDIS, Cass.
One specimen, from San José, April.
103. VrrEOsYLVIA OLIVACEA, Linn.
San José, January to May.
104. VirrOSYLVIA PHILADELPHICA, Cass.
Several specimens, from San José, January.
105. VirEOSYLVIA FLAVIFRONS, Vieill.
Several specimens, from San José, January, March.
106. VIREOSYLVIA JOSEPH, Sclat,
Only one specimen, from Naranjo, April.
107. HyLorpuitus pEcurtatus, Bp,
One specimen, from San José, May.
108. CycLoRHIS FLAVIPECTUS, Sclat.
Cyclorhis subjiavescens, Cab.
Several specimens, from San José, February, March, and April.
Family AMPELIDA.
109. PriLoGonys caupatus, Cab.
Several specimens, killed in the forest of oak trees on the Volcano
of Irazu ; also some specimens from Navarro, May.
110. PHAINOPTILA MELANOXANTHA, Saly.
Several specimens of both sexes, from Navarro and Rancho Re-
dondo, January to May. Fouud in pairs along the streams—not in
the region of the oak trees, as suggested by Mr. G. Dawson Rowley,
but much lower down, at an altitude of about 4000 feet, and in
semitropical forests. The two sexes of this rare species have been
beautifully figured, from specimens obtained by me, in the ‘Orni-
thological Miscellany,’ part x.
54 MR. A. BOUCARD ON BIRDS FROM CosTA RICA. [Jan. 15,
OscINES TENUIROSTRES.
Family CareBip2.
111. DigiuossA PLUMBEA, Cab.
Several specimens, from Cartago, Navarro, Naranjo, Voleano of
Trazu, and La Laguna; all of them killed at the altitude of from
4000 to 8000 feet.
112. Dacnis vENusTA, Lawr.
Several specimens, from Candelaria, May.
113. CHLoROpPHANES sPrIza, var. guatemalensis, Sclat.
One specimen, from Naranjo, April.
OscINES CONIROSTRES.
Family TanaGrip&.
114. CHLoroPpHoniA CALLOPHRYs, Cab.
Several specimens of this fine species, from Navarro and Canda-
laria, May.
115. EUPHONIA ELEGANTISSIMA, Bp.
Several specimens, from San José. Found on the road feeding on
the fruit of the mistletoe. These birds sing beautifully, and are
easily domesticated.
116. EvpHonia CRASSIROSTRIS, Sclat.
Only one specimen, from Cartago, April.
117. Evpnon1a Gou.nt, Sclat.
Two specimens, male and female, from San Carlos, February.
118. CALLISTE ICTEROCEPHALA, Bp.
Calliste frantzii, Cab.
Two specimens, from Naranjo and Orozi, June, May. Rather
abundant on the edges of the forests where clearings have been
opened.
119. CaLuisTeE GyroLorpes, Lafr.
Navarro, May.
120. Cauuisre LARVATA, Du Bus.
Calliste francisce, Latr.
Several specimens, from Naranjo, Orozi, and San Carlos.
121. CaLuisrE bowl, Salv.
Several specimens, from Navarro, May. Rare.
122. TANAGRA CANA, Sw.
Tanagra diaconus, Less.
Several specimens, from San José, January. Rather common on
the trees bordering the coffee-plantations.
1878. ] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 55
123. TANAGRA PALMARUM, Max.
Tanagra melanoptera, Hartl.
Common at Naranjo, April; San José, March to May.
124. RAMPHOCELUS PASSERINI, Bp.
San Carlos, February ; Naranjo, April. Found commonly in the
“rastrojos”’ (abandoned plantations near the habitations). Always
seen in pairs; but several pairs are often met with near one another
in the same locality.
125. PHLOGOTHRAUPIS SANGUINOLENTA, Less.
Orozi, May. I have seen them in pairs, like the above species ;
but sometimes five or six may be seen eating fruits or seeds on the
same tree.
126. Pyranca ruBRA, L.
San José, April.
127. PyRANGA ERYTHROMELENA, Licht.
Naranjo, April ; Candelaria, May. In the forest.
128. PyraNGA BIDENTATA, Sw.
Several specimens, from the forests of Navarro and Volcano of
Trazu, May.
129. PH@NICOTHRAUPIS FUSCICAUDA, Cab.
Two specimens, male and female, from San Carlos, February.
Found in the forest, perching on branches near the ground. Feeds
on insects.
130. TacuypuHonvus Luctuosvus, Lafr. & D’Orb.
San Carlos, February. Feeding on insects in the forest.
13]. TACHYPHONUS DELATTRII, Lafr.
San Mateo, May. I have always found these birds in company
with the above species, and many others, where large quantities
of migratory ants were travelling. They went in the same direction
as the ants; and now and then I saw them hopping to the ground,
and perching on low branches. Doubtless they were feeding on
these ants.
132. CHLOROSPINGUS ALBITEMPORALIS, Lafr.
Several specimens collected at La Laguna, Naranjo, and Navarro,
February, April, and May. This species seems to be widely dis-
tributed. I have seen them in the forest, chiefly at places where
openings had been made for sowing-purposes.
133. BuARREMON CAPITALIS, Cab.
Pezopetes capitalis, Cab.
Several specimens from the Volcano of Irazu, April and May. The
two sexes are exactly alike. They go in pairs; but several pairs
56 MR. A. BOUCARD ON BIRDS FROM CosTA RIcA. [Jan. 15,
may be seen at a short distance from one another. They perch on
the lower branches in the forest, but are most of their time busy
scratching the ground in search of insects. They make such a noise
in doing so, that at first I thought they were Odontophori or Tv-
nami. The sexes are constantly calling each other; and as soon as
you see one appearing, the other follows shortly after. They run
fast. I have never seen this bird anywhere else.
134. BUARREMON BRUNNEINUCHUS, Lafr.
Several specimens from Cartago, Navarro, and Rancho Redondo,
April, May. Rather a rare species in Costa Rica ; all my specimens
were obtained on the edges of abandoned plantations at the altitude
of 4500 feet.
135. BUARREMON CHRYSOPOGON, Bp.
Many specimens, from San José and Cartago.
136. BuarREMON TIBIALIs, Lawr.
Several specimens, from Navarro and the Volcano of Irazu, May.
Rather scarce ; found in the forest, principally along the streams.
137. ARREMON AURANTIIROSTRIS, Lafr.
Arremon rufidorsalis, Cass.
One specimen, from San Carlos, February.
138. SALTATOR MAGNOIDES, Lafr.
Several specimens, from San José and Cartago. Common in March
and April.
139. SatTaToR GRANDIS, Lafr.
Several specimens, from San José. Common in March, April, and
May.
Family FRINGILLIDE.
140. CuHRYSOMITRIS BRYANTII, Cass.
One male specimen, from the Volcano of Irazu, May. Rare.
Subfamily PasseRELLINAE.
141. EMBERNAGRA STRIATICEPS, Lafr.
One specimen, from San Carlos, February. Killed close to the
rancho. I have also seen this species at Puntarenas.
142. Pyreisoma CABANISI, Sclat. & Saly.
Many specimens, from San José and Cartago, January to May.
Killed on a sort of hawthorn edging the coffee-plantations.
143. PyrGisoMa LEUCOTE, Cab.
Melozone leucotis, Cab.
Several specimens, from San José, February, and Cartago, April.
Very rare.
1878.] | MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 57
144, ZonorRicHIA VULCANI, sp.n. (Plate IV.)
Supra cinereo-olivacea, pileo, dorso alis et cauda pallide nigris,
plumarum marginibus olivascentibus, secundariorum dorso proxi-
morum marginibus externis rufis ; subtus plumbescenti-cinerea
unicolor, rostro et pedibus pallide flavis.
Forehead olivaceous grey, with the central extremity of each fea-
ther nearly black ; back and wing-coverts of the same colour, but
the black is darker and very conspicuous in each feather ; tail-coverts
olivaceous ; upper surface of the tail almost black, with olivaceous
fringes ; underside uniform plumbeous grey, slightly darker on the
breast and sides of the abdomen ; under tail-coverts olivaceous grey ;
underside of the wings and tail about the same grey as the abdomen,
with silky reflections; primaries and tertiaries black, with olivaceous
fringes ; secondaries with the fringes wider and rufous in the upper
part ; the upper mandible of the beak nearly red, then black, with
white tip; under mandible fleshy yellow ; toes of the same colour.
Full length 62 inches, wing 34, tail 3.
There is no difference whatever between the male and female,
except that the latter is a trifle smaller.
The young is very much like young of Junco, Peucea or Melo-
spiza; the upperside and underside are olivaceous grey, with the
centre of each feather black ; the fringes of the upper wing-coverts
are wider than in the adult specimens and more rufous.
Several specimens, from the Volcano of Irazu, obtained at the alti-
tude of 10,000 feet.
145. Zonorricu1A PILEATA, Bodd.
Many specimens, from San José, Cartago, Sarzero, and Volcano of
Trazu. Vulgarly called “Come-maiz” (Eat-maize), because they are
very injurious to the maize-plantation crops. It is met every where,
even in the gardens, and takes the place in America of the Passer
domesticus in Europe. I have seen them at nearly all altitudes.
The eggs of this bird are green, with rufous spots all over at the
broader end.
Subfamily Cyanospizinz.
146. VoLATINIA JACARINA, L.
San José, March.
147. Cyanospiza ciris, L.
One specimen from Tres Rios, January. Obtained in a curious
way. Having shot what I thought to be Cyanospiza cyanea, I could
not find my bird where I thought I saw it fall, but during my search
I found this specimen of Cyanospiza ciris, quite dead. It was so
thin that it had evidently died from cold and hunger ; and I attribute
to this fact the body not being decomposed, as I have no doubt that
it had been dead for some time. I took great care in skinning the
bird; and I had the satisfaction to make a fine skin of it, which
is now exhibited.
58 MR. A. BOUCARD ON BIRDS FROM costa RICA. [Jan. 15,
148. Cyanospiza CYANEA, L.
San José, March.
149. PHONIPARA PUSILLA, Sw.
Several specimens, from San Jos¢, January and February. Found
in the rastrojos.
Subfamily SPERMOPHILIN.
150. HepyMELES LupoyicrANa, L.
One specimen, from San José, January.
151. PuHeructicus TIBIALIs, Baird.
Several specimens, from Navarro and the Volcano of Irazu, May.
152. SPERMOPHILA MORELETI, Pucher.
Several specimens, from San José, March and April. Very com-
mon in the maize-plantations in the time of the harvest.
153. SPERMOPHILA CORVINA, Sclat.
Two specimens, male and female, from San Carlos, February.
154. SpeRMOPHILA AURITA, Bp.
Spermophila semicollaris, Lawr.
One specimen, male, from San Mateo.
Subfamily EMBERIZIN&.
155. Evspiza AMERICANA, Gm.
Several specimens, from San José and Potrero cerrado (Volcano of
Irazu). Excessively common in March, flying about on the roads
in large flocks. It is then very fat, and delicious eating.
OscINES CULTRIROSTRES.
Family IcrERID&.
156. OcyaLus wAGLeRI, Gray and Mitch.
Orozi, May. I have also seen many specimens at San Carlos and
Naranjo.
157. Ostrnorps MONTEZUMA, Less.
Very common at San Carlos and Naranjo, February. The sight
of their nests hanging close together at the extremity of the branches
on the treesis very peculiar. They select the highest trees left where
the forest has been cleared ; so that they cannot escape detection. I
have seen as many as thirty nests in one single tree. They make a
fearful noise when they come and go out of their nests.
158. Casstcus pREvosTI, Less.
One specimen, from San José, March. Killed on the bushes
forming the edges of coffee-plantations. I have seen many other
specimens.
1878. ] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 59
159. IcrERUS BALTIMORE, L.
Several specimens, from San José, March and April. Called vul-
garly ‘‘ Cacique de Naranjo” (Cacicus of orange-trees) ; because they
are very fond of this fruit.
160. IcTERUS PROSTHEMELAS, Strickl.
One specimen, from Naranjo, April.
161. MoLorurus zNevs, Wagl.
Very common at San José, February to May. They go about in
large flocks, and are often seen in the gardens of the town, as well
as on the pavement in the streets.
162. SruRNELLA LUDOVICIANA, L.
Very common in the “ potreros”’ (plains), principally at Cartago,
April and May.
Family Corvip.
163. Psttoruinus morio, Licht.
Several specimens, from San José. Very common in the coffee-
plantations. Called vulgarly “Pepe;” because their cry is an imi-
tation of that Spanish word, in English “papa.” As soon as they
perceive any body, man or beast, moving about they fly, making a
fearful noise with their cries of pa, pa, pa. You can depend upon
it that when you hear these birds crying some one is near at hand.
Subordo TRACHEOPHONZ.
Family DENDROCOLAPTIDE.
164. SyNALLAXIS ERYTHROPS, Sclat.
Two specimens, from the Volcano of Irazu, March, and Navarro,
May. In the forest.
165. SynaLLaxis Pupica, Sclat.
Synallaxis nigrifumosa, Lawr.
One specimen, from Naranjo, April; killed on the edge of the
sugar plantations.
166. PsEUDOCOLAPTES BOISSONEAUTI, Lafr. (?).
One specimen from Navarro, May. It differs from the other
specimens which I have of P. doissoneauti in the sides of the throat
being yellow instead of pure white; however, for the present, I do
not consider this difference sufficient to make another species of it.
Subfamily Poityporinz,
167. PHILyDOR RUFO-BRUNNEUS, Lawr.
One specimen, from Navarro, May.
168. ANABAZENOPS VARIEGATICEPS, Sclat.
Several specimens, from Candelaria, April, May.
60 MR. A. BOUCARD ON BIRDS FROM CosTA RICA. [Jan. 15,
Subfamily DENDROCOLAPTIN&.
169. MARGARORNIS RUBIGINOSA, Lawr.
Two specimens, from Navarro, May.
170. GuypHORHYNCHUS CUNEATUS, Licht.
One specimen, from Naranjo, April. Killed in the forest on the
trunk of a tree. This bird is constantly flying from one tree to
another. It is one of those which travel in company with many
other species.
171. DENDROCOLAPTES SANCTI-THOMA, Laft.
One specimen, from Navarro, April.
172. DENDROCOLAPTES PUNCTICOLLIS, Sclat. and Salv.
Two specimens, g and @, from Naranjo, April. I killed the two
on the same tree. They were busily engaged in building their nest
in a hole of the trunk.
173. DENDRORNIS SUSURRANS, Jard.
One specimen, from San Carlos, February.
174. Prcotarpres compressus, Cab.
Two specimens, from Navarro, May.
175. X1IPHORHYNCHUS PUSILLUS, Sclat.
One specimen, from Naranjo, April. This species seems to be rare.
My specimen was killed on the trunk of a tree when going up and
down on the same.
Family Formicariip&.
Subfamily THAMNOPHILINZE.
176. THAMNOPHILUS NIGRICRISTATUS, Lawr.
Thamnophilus punctatus, Cab.
One specimen, from San Carlos, February. Rare. All the species
of this genus go about in the forest in company with many other
birds.
177. THAMNOPHILUS N&VvIUS, Gm.
Two specimens, from San Carlos, February.
178. THAMNOPHILUS DOLIATUS, L.
One specimen, from San Mateo, May.
179. DysirHAMNUS STRIATICEPS, Lawr.
One specimen, from San Carlos, February. A very rare species.
180. DysIrHAMNUS SEMICINEREUS, Sclat.
Several specimens, Naranjo, April,
1878. MR, A. BOUCARD ON BIRDS FROM COSTA RICA. 61
Subfamily Formicrvorin#&.
181. MyRMOTHERULA MELZNA, Sclat.
Only one specimen of this fine species. It seems to be very rare.
I killed it in the forest perched near the ground. It formed part of
a large flock of birds which was not a single moment at rest. As I
said before, it is a very common occurrence to see large quantities of
ants where these birds are. I have always observed that the Tana-
grids (genus Buarremon), Turdidee (several genera) and Dendrocolap-
tidee (all the genera) and a few others keep to the ground, feeding on
ants and other insects. At the same time birds of other genera, as
Tanagra, Calliste, Pipra, and of many genera of Tyrannide, Coere-
bidz, and Ampelide, are perched on the trees above, some of them
feeding on fruit, others on insects. All of them are constantly flying
about all the time.
But the most interesting fact is that every species follows the same
route. They travel rather fast ; and it has always been an object of
admiration to me to see how far they could travel in one day, taking
their food at the same time.
I was in the habit of searching for one of these flucks. Once
found, it was only a matter for me of selecting what species I wanted ;
but you must follow the same route as they, and occasionally you
may lose yourself in the forest. These remarks apply to species
found in the tropical forests from the level of the sea to the altitude
of 4000 feet. But I have also observed the same thing with the
species which inhabit higher altitudes, whether they live in the
forests or on the plains.
182. MyrRMOTHERULA MENETRIESI, d’Orb.
One specimen, from Naranjo, April.
183. ForMIcIvVORA BOUCARDI, Sclat.
San Carlos, February.
Rather rare.
184, RAMPHOC@NUS SEMITORQUATUS, Lawr.
One specimen, from San Carlos, February. Very rare. Perch on
small trees in the forest. It was with the flock spoken of above.
185. CERCOMACRA TYRANNINA, Sclat.
San Carlos, February.
186. Myrmeciza immacuLatTa, Sclat. and Salv.
Several specimens, from San Carlos, February, and San Mateo,
May. Rare.
This species is found on both sides of the country—I mean, in the
tropical forests of the Atlantic and Pacific plains. It is therefore
very probable that tropical species travel from one sea to the other
where they have the facility to do so, without the necessity of pass-
ing to a higher altitude; and this is the case in Costa Rica. In a
certain part of the country you can pass from one sea to the
other without leaving the tropical forests.
62 MR. A. BOUCARD ON BIRDS FROM CosTa RICA. [Jan. 15,
Subfamily ForM1IcaRriInz.
187. Pirays picotor, Lawr.
One specimen, from San Carlos, February.
188. FoRMICARIUS HOFFMANNI, Cab.
One specimen, from San Carlos, February, killed in the forest,
on the ground. This bird is constantly going about, and has a
peculiar way of raising and lowering its tail when running along,
with which I was very much amused. I saw several other speci-
mens always on the ground. ‘The same thing occurs with the birds
of the genus Siurus.
189. GRALLARIA PERSPICILLATA, Lawr.
One specimen, from San Carlos, February. Very rare.
Family TyrRaANNID&.
190. Savornis AQUATICA, Sclat. & Saly.
Several specimens, from San José, January, March. This bird has
the curious habit of staying all day long on the stones or bushes grow-
ing in or near the streams. It will always be seen at the same
place for hours and days one after another. It lives on all the
insects flying about over the stream. They are always seen in pairs,
g and @. Another pair may occasionally be seen at a short
distance.
Subfamily PLatyRHYNCHINZ&.
191. PuaryRHYNCHUS ALBIGULARIS, Sclat.
One specimen, from Navarro, May.
192. TopIROSTRUM CINEREUM, L.
Several specimens, from Cartago and San José. Rather rare.
193. EuscAaRTHMUS SQUAMICRISTATUS, Lafr.
Two specimens, from Naranjo, April. A rare species and difficult
to detect. Goes about in the forest on the small trees.
194. SERPOPHAGA CINEREA, Strickl.
From Naranjo, April. Excessively rare and remarkable for its habits.
It has exactly the same mode of life as Sayornis aquatica, living
along the streams and sitting on the stones lying in or near the
water.
First I killed the 2, in the morning, and for hours could not get
a shot at the ¢, and went away. In the afternoon I went again
to the same place, and after a little while managed to kill the g
bird. The two sexes are exactly alike.
My specimens differing from the Columbian bird, which I have
under the name of S. cinerea, I give a description of them :—
Forehead and cheeks quite black, with the inner webs of the
central feathers white, back and upper coverts of the tail ashy grey ;
1878.] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 63
throat, breast, and abdomen of the same colour, but much paler,
nearly white on the throat ; primaries and secondaries black, tertiaries
black with white ends or fringes ; upperside of the tail black, under-
side dark grey, nearly black; under wing-coverts of the same colour
as the abdomen ; bill and toes black.
Length 4 inches, wings 24, tail 13. The two sexes are exactly
alike. Closely alliedto S. cinerea, but a much smaller bird altogether ;
besides, the colour of the back has not the olive tint conspicuous in
S. cinerea.
Subfamily Evaineinz.
195. Mionecres oLEaGinus, Cab.
One specimen, from San Mateo, May.
196. Tyranniscus pAaRvus, Lawr.
One specimen, from Navarro, May.
197. ELAIneA pAGana, Licht.
Elainea subpagana, Sclat. & Salv.
Several specimens, from San José, January to May. On the
edges of the coffee-plantations.
198. Exarnra rrantzu, Lawr.
One specimen, from the Volcano of Irazu, March.
199. LeGatus ALBICOLLIs, Vieill.
Several specimens, from Naranjo, April, all of them killed on a
large tree standing by itself in a “ potrero” near the habitations.
200. Myiozeretes TEXENs!s, Giraud.
Two specimens, from San José, March, and Naranjo, April.
Common.
201. Ruyncnocycius suLPHURESCENS, Spix.
Only one specimen, from San Carlos, February.
202. MytopyYNastTEs LUTEIVENTRIS, Sclat.
Two specimens, from Orozi, May.
Subfamily Tyranninz.
203. MEGARHYNCHUs PITANGUA, L.
Native name “ Chorchita.”
One specimen, from San José, March. Not common here.
204. Muscivora MEXICANA, Sclat.
Native name “‘ Rey de los Traga moscas.”’
Two specimens, male and female, from San Ramon. Lives in the
forest only. When in love it shows its beautiful crest with much
effect,
64 MR. A. BOUCARD ON BIRDS FROM CosTA RICA. [Jan. 15,
205. Myr1osrus sULPHUREIPYGIUS, Sclat.
Two specimens, from Naranjo, April.
206. Myrosius ERyTHRURUS, Cab.
One specimen, from San Carlos, February.
This seems to be a rare species. I wanted more specimens, and
could only succeed in seeing another, which I could not shoot.
207. MirREPHORUS AURANTIIVENTRIS, Lawr.
Several specimens, from La Laguna, February Naranjo, April,
and Candelaria, May. I killed these in the forest at the altitude of
4000 to 8000 feet. They used to stay at the same place for a long
time ; they select a branch, and you see them there ail day long.
All the time they fly about catching insects, and returning to the
same spot. I missed one of my specimens twice ; notwithstanding,
it came back to its favourite branch. There is no appreciable
difference between the two sexes. I have also one specimen with
the abdomen of the same colour as the throat and the breast ;
otherwise there is no appreciable difference. It was obtained at
San Mateo, altitude 1000 feet.
208. Emprmponax FLAVESCENS, Lawr.
Several specimens, from Naranjo, April.
209. Conropus virENs, L.
Several specimens, from Cartago, April.
210. My1arcaus CRINITUS, L.
Native name ‘‘ Traga mosca.”
Several specimens, from San José, January, and Cartago, April.
Rather rare in Costa Rica. They used to perch on a selected branch
of the trees edging the coffee-plantations, and remained there all day
long catching the insects passing by.
211. My1ArcHUS PANAMENSIS, Lawr.
Three specimens, from Punta Arenas, May. Killed on the mangle-
trees near the river.
212. My1arRcHUS LAWRENCEI, Giraud.
Myiarchus nigricapillus, Cab.
Many specimens, from San José, Cartago, and Naranjo, January
to May.
Native name “‘ Tonto vivo.”’
213. My1arcuus CINERASCENS, Lawr.
Two specimens, from Punta Arenas, May. Killed at the end of
the sandy point near the town.
214. TyRANNUS MELANCHOLICUS, Vieill.
Many specimens, from San José, January to May. Common
everywhere, especially on the trees surrounding the coffee-plan-
tations.
1878. ] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 65
215. Mitvuuus rorricatus, Gm.
Native name “ Tijerillo.”’
Several specimens from San José, January to May. Very common
in the Savana, a beautiful plain about one mile and a half from the
town. They are usually perched on the small shrubs in the plain.
Family Corineip 2.
216. Tiryra personata, Jard. & Selby.
Several specimens from Orozi. Always seen perched on the highest
trees, where they remain sometimes for hours, as if they were on the
watch. They are easily mistaken for Carpodectes nitidus, which they
resemble very much when seen at a long distance.
217. PacnyRHAMPHUS VERSICOLOR, Hartl.
Only one specimen, from Candelaria, May. Very rare.
218. PacHyrRHAMPHUS CINEREIVENTRIS, Sclat.
One specimen, from San Mateo, May.
219, Lipaucus HOLERYTHRUS, Sclat. & Salv.
Several specimens from San Carlos, February ; and Naranjo,
April.
220. CarPoDEcTES NITIDUs, Salv.
One adult female, from San Carlos, February. This sex, being
quite new to science, I give a description of it.
Head and back of a dark bluish grey, rather lighter on the fore-
head ; the upper feathers of the tail nearly black, the underside ashy
grey ; wing-coverts of the same colour as the back ; primaries black ;
the secondaries and tertiaries have a white border on their inner webs ;
cheeks, throat, and round the eyes white ; breast and abdomen pale
bluish grey ; under wing- and under tail-coverts white: the lower
part of the abdomen white ; thighs white ; toes black ; bill blue.
Total length 82 inches, wing 52, tail 28,
I procured this bird quite accidentally. During our stay at San
Carlos we used to bathe every afternoon in a stream close by the
rancho, and I always carried my gun with me. The third day after
our arrival at San Carlos we were quietly enjoying our bath when
my friend Meil saw this bird alighting on a tree close by. Imme-
diately he told me I came out of the water and took my gun; but
the bird was perched so high that I could not make out with cer-
tainty what species it was. I was struck by its colour, and I thought
immediately of Carpodectes, of which I had seen a figure in the
‘Proceedings.’ I had a shot at the bird; and it fell down, but far
away in the forest amongst Agave plants growing near the stream.
My friends saw the direction where it fell ; and although I was quite
naked, I went into the forest, and, after twenty minutes of the most
active search, I had the pleasure of finding it quite dead. Several
times I was near giving up the search ; but now I feel very happy
Proc. Zoou. Soc.—1878, No. V. 5
66 MR. A. BOUCARD ON BIRDS FROM COSTA RICA. ([Jan. 15,
that I did not do so, as I consider this bird one of the most valuable
additions to my collection.
Every day after that. I went to the same place and in the neigh-
bourhood to look for more; but I was disappointed, until the day
before my departure, when I saw what I think was the mate of my
female; but, to my great annoyance, I could not get a shot at it.
It sat for more than two hours, without moving, at the top of a
tree more than one hundred feet high, where I could not kill it.
It kept me until nightfall with my head lifted up in such a manner
that I had a stiff neck when I left. I think it is the greatest punish-
ment for a naturalist to be looking at a very rare bird above his
head for several hours without being able to get a shot at it.
When I went away I recommended the people living in the rancho
to kill some specimens for me. I offered them a high price ; but all
for nothing: they said they never saw another. If it was not neglect
of theirs, this must be a very rare bird indeed.
221. CHASMORHYNCHUS TRICARUNCULATUS, Verr.
One specimen from the Volcano of Irazu, January. This seems
to be arare species. My specimen was killed at the altitude of
6000 feet.
222. CEPHALOPTERUS GLABRICOLLIs, Gould.
San Carlos, February ; Volcano of Irazu, May. This singular bird
is found at all altitudes up to 10,000 feet in the forests. Were it
not for its cries it would be excessively difficult to detect, as it
always keeps in the densest parts. Its ery is something extraor-
dinary. When I heard it first I thought it was a new species of
Crax or some other similar large bird. Being very anxious to see
what it was, I went into the forest ; and after a great deal of trouble
I found it. They perch rather high, but are easily killed when
once found.
Family Pirripz.
223. PrpRA MENTALIS, Sclat.
One specimen from San Carlos, February. Perches usually low,
and keeps in the forest.
224. PirrA LEUCORRHOA, Sclat.
Cervantes, Naranjo.
225. CHIROXIPHIA LINEARIS, Bp.
Several specimens, from Navarro, May.
226. CHIROMACHAERIS CAND&I, Parz.
One specimen, from San Carlos, February. Perches on the small
trees in the forest; and the male has the particular habit, as have
some other species of Pipridz, of making a great sound when flying
from one branch to another. It imitates exactly the sound produced
by the children’s toy rattle. I think this noise is produced by its
1378.] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 67
beak. It resembles the sound of mandibles opening and shutting with
noise several times in succession.
Ordo Macrocuires.
Family Caprimuteip2.
227. NYCTIDROMUS ALBICOLLIs, Gm. -
Two specimens, male and female, from Cartago. They were ina
small forest, on the ground. They usually fly away as you get quite
close to them, and settle again a little further on.
Family Hirunpinip2.
228. ATTICORA CYANOLEUCA, Vieill., var. monTANA, Baird.
Several specimens from San José, March; Cartago, April. During
these months they are building their nests in the roofs of the houses.
They appear in February, and leave when the rainy season comes.
229. STELGIDOPTERYX FULVIPENNIS, Sclat.
Stelgidopteryx fulvigula, Baird.
Several specimens, from San José, March to May. They are
principally seen flying near the streams, sometimes in large quanti-
ties.. The first time I had a shot at them I killed six ; there were
about fifty on a small tree. Although I have killed a good many,
I never found a female amongst them. I suppose the females must
have been in their nests somewhere ; but where, I was never able to
find out.
Family Cypseip&.
230. Hemiprocne zonaris, Shaw (?).
In one of my excursions, at the Volcano of Irazu I saw a large
quantity of birds which, I think, must have been this species. I am
certain about the genus; but not having been able to get a specimen,
I leave the question of species for the future.
Ordo Trocurtt.
231. PHaETHORNIS LONGIROsTRIS, Delattre.
Naranjo, April. This bird is always seen in the denser part of the
forest. One of the favourite flowers on which it feeds is that of the
plane-tree.
232. PHAETHORNIS EMILLE, Bourc.
Several specimens from Tres Rios, January ; Naranjo, April.
233. Pyemornis ADOLPHI, Gould.
San Carlos, February. This is a very common species—but very
difficult to detect, as it only comes out of the forest early in the
morning and late in the afternoon. In the middle of the day they
are very abundant in certain parts of the forest, many specimens being
seen at a few yards from one another. They are perched on small dry
5®
68 MR. A. BOUCARD ON BIRDS FROM CosTA RICA. [Jan. 15,
branches near the ground. In the pairing-season they sing all day
long; the note sounds like ‘‘bebe, bebe,” repeated a great many times
in succession. Although it cannot be heard at a long distance, it is
perfectly amazing and amusing to hear them. As soon as one finishes,
another commences; and it is a puzzle to find where they are,
although they may be only at two or three yards distance, on account
of their colour and their diminutive size. When singing they move
their head and tail in the most pretty way. Now and then they fly
about, but soon return to their favourite branch. I have usually
heard the song of these birds in the breeding-season, the males
singing while the females are on their nests.
When in Mexico I have heard many species of Humming-birds
sing, and I could distinguish by its song what species it was.
Their songs are very pretty, and quite distinct from one another,
especially when they are of birds of distinct genera. I have heard
Pygmornis adolphi, Campylopterus hemileucurus, Sphenoproctus cur-
vinennis, Eugenes fulgens, Celigena clemencie, Lamprolema rhami,
Delattria henrici, Petasophora thalassina, and several other species
sing.
234. CAMPYLOPTERUS HEMILEUCURUS, Licht.
Several specimens from Tres Rios, January ; and Rancho redondo,
May. Rather rare in Costa Rica. Inhabits the forests; but takes
its food as well on the flowers of trees growing close by in the open
spaces. My specimens were obtained when sucking the flowers of
an acacia tree which is found abundantly in the coffee-plantations.
It has a special sharp cry when flying about. It commonly con-
structs its nest in the barrancas, on the plants growing between the
rocks, and generally uses moss for its construction.
235. EUGENES SPECTABILIS, Lawr.
One male and one female of this very rare species, from the Volcano
of Irazu, April and May. Altitude from 6000 to 8000 feet. It is
very fond of a parasitic plant something like mistletoe, which bears a
beautiful red flower. No description having been published of the
male of this species, I give it now.
Forehead dark metallic blue ; back black ; rump metallic green ;
tail bronzy black, with the two central feathers more greenish ;
wings black, with the upper coverts metallic green ; throat metallic
dark green ; breast and abdomen greyish, mixed with green feathers ;
feet and beak black.
Length 52 inches, wing 32, tail 18, beak 14. — (Mus. Boucard.)
The chief distinctions between this species and Eugenes fulgens are
in its much larger size, its longer beak, its grey breast and abdomen,
its back entirely black, and the colour of the crown and throat,
which are quite of a different green and blue. ‘,
236. OREOPYRA CINEREICAUDA, Lawr.
One specimen of this rare species, from Navarro, May. Inhabits
the forest. ;
1878. ] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 69
237. OREOPYRA CALOLAMA, Salv.
Many specimens of both sexes from Naranjo, April; Navarro and
Rancho redondo, May. I have not the least doubt that the bird
described as Anthocephala castaneiventris, Gould, is. the female of
this species. These birds are only met with in the forest.
238. OREOPYRA HEMILEUCA, Saly.
Two specimens, from Naranjo, April. It seems to be a very rare
species.
239. PerasopHoRA cyANoTISs, Bourc.
Petasophora cabanisi, Lawr.
Several specimens from the Volcano of Irazu, April. Rather
common at the altitude of 8000 feet, feeding on the flowers of a
small shrub. They take possession of a certain space containing
several of these shrubs; and when not feeding on these flowers,
they perch on a dry branch near the place, and fight all the other
Humming-birds that dare to intrude.
240. Hetropoxa sAcuLA, Gould.
Heliodoxa henryi, Lawr.
Several specimens from Naranjo, April; Volcano of Irazu, May.
One male, although it is not in very fine plumage, has a bright front,
and the throat exactly like the specimens from Columbia. Rather
rare in Costa Rica.
241. HELIOMASTER CONSTANTI, Delattre.
Only one young male, from San José, January ; killed while he was
sucking the flower of an acacia tree,
242. THALURANIA CoLOMBiIcaA, Bourc. & Muls., var. veNusta,
Gould.
Several specimens from San Carlos, February; and Naranjo,
April. It perches on very high trees in the forests, and is dithi-
cult to get at. It is a rare species in Costa Rica.
I spent many hours before I could get the pair which I have,
from San Carlos. I could not recognize the bird, because it was
standing and feeding on the flowers of a parasitic plant very high
on a large cedar tree. I thought it was a new species of Microchera,
so small was its appearance. When I killed the first I was quite
disappointed, and was sorry to have lost so much time for it.
243. Kuais cuimetti, Boure. & Muls.
Klais merriti, Lawr.
Only two specimens, from Navarro, May: a very rare species.
244. MicrocHERA PARVIROSTRIS, Lawr.
Only one specimen of a young male, given to me by Mr. Juan
Zeledon. Excessively rare. Although I knew the exact locality
where found, I could never find one.
70 MR. A. BOUCARD ON BIRDS FROM COSTA RICA. [Jan.15,
245. PapHiosa HELEN#, Del.
I saw several male specimens of this species at San Carlos; but I
was never able to get a shot at them. I have one female specimen
from Turrialba.
246. Dorycua BRYANTA, Lawr.
San José, March; Volcano of Irazu, April. A very rare species.
It appears as soon as it is daylight, and is very quick. It never rests
long in one place. When on the wing it makes a great noise, similar
to that of a large Coleopteron (Scarabeidee) when flying.
247. TROCHILUS CoLUBRIS, Linn.
One specimen, from San José, January. Seems to be very rare in
Costa Rica.
248. SELASPHORUS SCINTILLA, Gould.
Several specimens from Cartago, April; and Volcano of Irazu,
May. Seen also at the high altitude of 10,000 feet in company
with Selasphorus flammula. Feeds on the flowers of small plants.
When I say feeds on the flowers, I mean that it takes from the flowers
not only the honey, but, above all, the minute insects which are inside.
This is usual with all the Humming-birds. I have no doubt that
they can live a certain time on honey only; but it is not sufficient,
and their principal food is insects. It is very seldom that you skin
a Humming-bird without finding the crop filled partially with insects
of all descriptions. I have brought home a certain number of
crops in alcohol. All of them contain insects, which I intend to
determine as soon as possible. SS. scintilla makes a very slight noise
when flying ; it is not more than that of a drone-bee.
The place where I killed my specimens had been taken possession of
by a certain number of Sancerottia sophie and Pyrrhophena riefferi.
Each one of these had its assigned place, and used to drive away any
trespasser. I noticed that S. scintzlla used to go only to the small
plants near the ground; and I have no doubt that the little noise
made by the bird was to escape its enemies S. sophie and P. riefferi.
Flying very low was another precaution; however, the sight and
the hearing of these little birds is so subtle that poor 8. scintilla was
soon detected and obliged to fly away. I think this also is the reason
why this species was rather scarce.
249. SELASPHORUS FLAMMULA, Sally.
Several specimens, male and female, of this very rare species.
All of them were killed at the summit of the Volcano of Irazu, on
parasitic plants (mistletoe) growing on small alpine trees in a soil
composed principally of ashes and scoriz. There, at the altitude of
10,000 feet and more, it takes possession of a locality where flowers
are abundant, and defends its property against S. scin¢il/a and other
birds,
1878. ] MR. A. BOUCARD ON BIRDS FROM COSTA RICA. 71
250. THAUMATIAS CUPREICEPs, Lawr.
One specimen, from Naranjo, April, killed in the forest. I have
also several other specimens of this species sufficiently good for
skeletons.
251. EuPHERUSA EGREGIA, Sclat. & Salv.
One specimen, from Navarro, May.
252. AMAZILIA CINNAMOMEA, Less.
Several specimens, from San Mateo and Puntarenas, May. This
species seems to be found only on the Pacific coast. I have some
specimens from Tehuantepec; and I remember having seen this
species near Juquila (Mexico). The male and female are exactly
alike. They are fond of the flowers of orange-trees.
253. PyRRHOPHZANA RIEFFERI, Boure.
Several specimens, from San José and Cartago, January to May.
Very common species, and quite domestic in its habits. “Quantities
of them may be seen in private gardens.
254. SAUCEROTTIA SOPHLE, Bourc.
Several specimens, from San José and Cartago, January to May.
They are constantly fighting with P. riefferi; but this species is
seldom seen in the gardens of San José.
255. PANTERPE INSIGNIS, Cab. & Heine.
Several specimens of this fine species, from the Volcano of Irazu,
April and May. I never saw this bird anywhere else. It is very
rare. It feeds on the mistletoe growing on the robble (oak tree).
The female is a little smaller than the male ; but otherwise they are
exactly alike. This is one of the few species in which the female is
a bright-coloured bird, nearly as fine as the male. I have several
females, and I am quite sure about the sex.
256. Damopuita AMABILIS, Gould.
Several specimens, from San Carlos, February. A very rare species ;
found only in the forest.
257. SAPPHIRONIA BOUCARDI.
Arinia boucardi, Muls., Ann. Soc. Linn. de Lyon, 12 Oct. 1877.
Several specimens, from Puntarenas, May. Male and female.
Mus. Boucard.
258. CHLOROLAMPIS CANIVETI, Less.
Chlorolampis salvini, Cab.
Several specimens, from San José and Cartago, January to May.
Commonly found in the gardens, where it makes its nest.
“I
bo
MR. G. F. ANGAS ON [Jan. 15,
3. Descriptions of seven new Species of Land-Shells recently
collected in Costa Rica by Mr. Adolphe Boucard. By
Grorce Frencu Aneas, C.M.Z.S., F.L.S., &.
[Received November 17, 1877.]
(Plate V.)
Hewix BovucarpI, n.s. (Plate V. figs. 5, 6, 7.)
Shell narrowly and profoundly umbilicated, depressedly conical,
moderately thin, very finely radiately striated, white, the upper surface
ornamented with two fawn-coloured bands passing into chocolate to-
wards the apical whorls, or entirely fawn-colour, with a narrow effuse
chocolate band next to the periphery ; whorls 43, somewhat convex,
strongly bluntly keeled at the periphery, with an impressed groove
below the keel ; apex obtuse, tinged with violet ; sutures impressed,
bordered with a narrow chesnut line; base convex, white; umbilicus
dark brown; umbilical region tinged with lemon-yellow ; aperture
very oblique, narrowly quadrately ovate, pale violet within; outer
lip a little expanded and reflexed, slightly sinuate and projected
above, arcuate below.
Diam. maj. 13 lines, min. 10, alt. 7.
Hab. Navarro, Costa Rica; on leaves of trees. (Mus. Boucard.)
Heurx ADELA, n.s. (Plate V. figs. 8, 9, 10.)
Shell umbilicated, discoidal, flattened, very finely radiately striated,
pale straw-colour, ornamented with a single narrow black band just
above the periphery of the last whorl ; whorls 33, somewhat convex ;
spire very much depressed; base flatly convex, with an excavated
area around the umbilicus which is shallow and non-perspective ;
aperture horizontal, narrowly oblongly ovate; outer lip slightly ex-
panded and reflexed.
Diam. maj. 12 lines, min. 9, alt. 4.
Hab. Novarro, Costa Rica, on trees. (Mus. Boucard.)
Herrix zsorus, n.s. (Plate V. figs. 11, 12.)
Shell umbilicated, depressedly conoidal, moderately solid, very finely
obliquely striated and finely granulated, the granules being denser
on the last whorl and obsolete towards the apex, brown, with a pale
suffused band at the periphery of the last whorl; whorls 5, some-
what convex, the last bluntly keeled, contracted and deflected behind
the outer lip; apex obtuse; base rather flattened; aperture hori-
zontal, ear-shaped, brown, with a prominent, erect, slightly undulating,
lamelliform, transverse ridge within upon the wall of the body-whor! ;
peristome white, somewhat thickened and reflexed, with a short blunt
tooth at the base of the columella and a longer compressed triangular
ove incurved within the aperture opposite the ridge on the body-
whorl; margins united by an erect lamella.
Diam. maj. 12 lines, min. 10, alt. 6.
GB Sowerby lith Hanhart imp.
NEW SHELLS FROM COSTA RICA &c.
1878. ] LAND-SHELLS FROM COSTA RICA. 73
Hab. Buena Vista, Costa Rica, at an altitude of 3000 feet.
(Mus. Boucard.)
This shell belongs to a group of Helices remarkable for their
curiously toothed apertures, having as its nearest allies H. quadri-
dentata, Brod., and H. auriculina, Petit.
Cycorus Bovcaror, n.s. (Plate V. figs. 3, 4.)
Shell widely and perspectively umbilicated, globosely turbinate,
white, with a broad olive-chestnut band above and-a narrower one
just below the periphery, very finely longitudinally striated and
crossed with rows of rugged irregular undulating oblique malleated
ridges; whorls 6, convex, rapidly increasing, the last large and
rounded ; aperture nearly circular, white within 3 peritreme simple,
scarcely thickened, with a semilunar notch at the junction of the body-
whorl.
Diam. maj. 16 lines, min. 13, alt. 12.
Hab. San Carlos, Costa Rica.
A fine species with an excavated notch where the peritreme touches
the body-whorl, as in Cyclotus mexicanus, Menke, from Vera Cruz.
(Mus. Boucard.)
BuLIMUs JOSEPHUS, 0. s. (Plate V. figs. 13, 14.)
Shell perforate, elongately ovate, moderately solid, very finely
obliquely striated, whitish, ornamented with somewhat distant irre-
gular slightly undulating narrow olive-brown stripes, which become
obsolete near the umbilical region; spire elevately conical; sutures
impressed ; apex brownish yellow; whorls 63, flatly convex ; aper-
ture ovate; somewhat effuse below ; outer lip expanded and flattened;
columella triangular and flattened inwards, ending in a blunt rounded
callus internally.
Diam. 7 lines, alt. 16.
Hab. San José, Costa Rica, on the trunks of acacia trees.
(Mus. Boucard.)
Buximvus 1razueEnsis, n.s. (Plate V. figs. 17—2().)
Shell somewhat elongately ovate, rimate, moderately thin, longi-
tudinally irregularly striated, shining, more or less longitudinally
striped or freckled with black, and ornamented with numerous smal}
white spots; whorls 6, rather convex ; spire a little shorter than the
aperture ; aperture oblong-ovate ; lip thin, simple, tinged inside with
rose-colour.
Diam. 6 lines, alt. 12.
Hab. Volcano of Irazu, Costa Rica, on low aromatic bushes on
the eastern slope of the volcano.
A variety of this pretty species occurs in which the prevailing
colour is light brown, profusely sprinkled with little white spots.
(Mus. Boucard.)
BuLimus NAVARRENSIs, n.s. (Plate V. figs. 15, 16.)
Shell narrowly rimate, more or less elongately ovate, thin, irre-
74 MR. G. F. ANGAS ON A NEW LATIAXIS. [Jan. 15,
gularly longitudinally striated, shining, whitish, tinged with pale
straw-colour towards the base, variously painted with irregular more
or less undulating longitudinal greyish black stripes or quadrilateral
blotches, frequently taking the form of a confused band towards the
centre of the whorl; spire conical, shorter than the aperture ; apex
acute ; whorls 53, rather convex, the last somewhat inflated ; aperture
elongately ovate ; peristome simple, thin ; outer lip slightly expanded
outwards.
Diam. 6 lines, alt. 12.
Hab. Navarro, Costa Rica; a somewhat smaller and more inflated
variety was obtained by Mr. Boucard at Cartago.
This species somewhat resembles B. iodostylus of Pfeiffer ; but it
is generally more inflated, with the outer lip expanded, and wants the
violet colouring on the columella characteristic of the latter.
Abundant on acacia trees. (Mus. Boucard.)
EXPLANATION OF PLATE V.
Figs. 1, 2. Latiaxis elegans, p. 74. Figs. 11,12. Helix esopus, p. 72.
3,4. Cyclotus boucardi, p. 73. 13, 14. Bulimus eae? 2M p- 73.
5-7. Helix boucardi, p. 72. 15, 16. navarrensis, p. 73.
8-10.
adela, p. 72. 17-20.
irazuensis, p. 73.
4. Description of a new Species of Latiazis. By GuorcE
Frencu Aneas, C.M.Z.S., F.L.S., &e.
[Received November 17, 1877.]
(Plate V.)
LaTIAXIS ELEGANS, 0.8. (Plate V. figs. 1, 2.)
Shell rimate, ovately turreted, solid, white; length of spire and
aperture equal ; whorls 54, irregularly distantly longitudinally ribbed,
flattened and excavated above, with a wide projecting keel, curving
upwards and scalloped at the edge, running round the periphery of
the whorls, which are encircled by scabrous ridges, fine and close-set
above the keel, stouter, wider apart, and more squamate below, one
ridge (intermediate between the keel of the body-whorl and the
base) being larger and more elevated than the others ; aperture semi-
oval; outer lip sulcate within; columella nearly straight ; umbilical
region bordered by an elevated squamose ridge; canal very short,
slightly twisted and recurved.
Diam. 11 lines, alt. 1 in. 2 lines.
Hab. Unknown.
This beautiful species of Latiawis has been placed in my hands
for description by Mr. Robert Geale.
1878. | DR. H, BURMEISTER ON CONURUS HILARIS. 75
5. Notes on Conurus hilaris and other Parrots of the Argen-
tine Republic. By Dr. H. Burmeister, F.M.Z.S.
[Received November 19, 1877.]
During my residence at Tucuman, in the years 1859-60, I saw
several flights of six or eight of a long-tailed parrot, of which I was
not able to obtain a specimen, although the birds came every week
during the maturing-time of the Indian corn to the garden of the
house in which [ lived, and did much damage to the seeds of the
plants cultivated there. This bird, while it had the same habit of
plundering the gardens as the other Parrots inhabiting this country
(for example Conurus patagonus and C. murinus), was also, like its
fellows, very cautious, some of them keeping watch on the top of the
highest plants, and, as soon as any one appeared with gun in hand,
giving a quick cry of alarm, upon which the whole flock flew away.
On this account I was unable to obtain a single specimen of this
beautiful bird, which I have not seen in any other part of this country,
and which I recognized at once, even ai a distance, as new to my
collection, from its large red front. At last I found a living specimen
in the possession of a shop-keeper, who would not part with it,
having reared it from the nest and made a great favourite of it.
But I was able to observe more carefully the specific distinctness of
this bird than from a distance; and from this examination I made
the description published in my ‘ Reise durch die La Plata-Staaten,’
vol. li. p. 442, as follows :—
Size and colour of Conurus murinus ; whole plumage bright green,
only the forehead at the base of the bill blood-red. Bill, in life,
pale rose-colour, or quite white in old individuals. Carpal region
green; the wing-feathers blackish, becoming bluer towards the
margins. Tail narrow and cuneiform, greenish towards the margins,
red at the base and below. Legs light flesh-colour. Iris orange.
Whole length 10 inches, wing 5, tail 43.
This short description, the only one I was able to give at the time,
was not sufficient to make the species thoroughly known. I there-
fore wrote to my friends at Tucuman, as soon as I returned to
Buenos Aires in the year 1862, to send me specimens of this
interesting Parrot ; but all my solicitations, though repeated many
times, were without effect. I never obtained a single specimen,
although the bird is not rare in the vicinity of the large forests on
the western sides of the town, near the slopes of the mountains, and
sometimes very common.
At last my friend Dr. Bruland, physician at Tucuman, an excel-
lent collector who had forwarded me many interesting objects for
my collection, brought me two well-preserved skins; and some
months before, Mr. Holmberg, of Buenos Aires, who made a
journey to collect zoological objects in the northern provinces,
brought home from Salta a skin of the same species, which I was
able to compare with the other two now in the Public Museum of
Buenos Aires.
76 DR. H. BURMEISTER ON CONURUS HILARIS-_ [Jan. 15,
These three skins enable me to give the following full description.
Beginning with the size, I find that the living specimen I examined
must have been very young, and by no means fully adult, as the
three skins now under my inspection are alllarger. The two largest
measure from the front to the tip of the tail 13 inches, and the
smaller specimen (from Salta) 12 inches, tail 63 inches, and the
wings 73-7}.
The predominating colour is pure green, somewhat darker on the
upperside, more especially on the head and neck, but clearer on the
underside ; the wing- and tail-feathers are of the same colour outside,
both with black shafts, and the former with a blackish interior margin
inclining to blue. The inside of these above-mentioned feathers is
yellowish grey, darker on the margins, and the yellow colour clearer
on the tail-feathers towards the base, which in some specimens are
of an orange appearance.
There is no red on these feathers in any of the three specimens ;
but it is possible that when alive it may have had this colour, if not
very distinct, at least visible. I find in one of these specimens a
red spot on two of the tail-coverts, which are generally completely
reen.
5 The red colour is regular only on the front of the head, beginning
from the base of the upper mandible, extending up to and behind the
eye; but a small space of green appears over the middle of the eye,
and descends on the cheeks to the ear and the under mandible,
with some green feathers near the angle of the mouth.
At first these red feathers are very small and of a dark red-
brown, and thence the others become blood-red. Besides this regular
frontal patch there are other red feathers on other parts of the
body, but without any constant regularity in different specimens of
the same species. I find in my three examples great difference in
the distribution of these red feathers.
There generally seem to exist some red feathers on the neck,
front of the throat, and breast ; also there is always a circle of red
feathers at the end of the tibia.
As to the habits and manner of living of this bird, I cannot add
any thing to what I have already said at the commencement of this
communication. It lives only in the northern mountainous dis-
tricts of the Argentine Republic, never coming to the plains of the
Pampas.
I take this occasion to correct some mistakes made by Dr. Finsch
in his work, respecting my previous communications on different
species of Parrots of this country and of Brazil.
The author says (vol. i. p. 174) that he knowns nothing of the
spines on the sides of the tongue of the Ar, mentioned by me in
my work on the Animals of Brazil. In that work (vol. ii. p. 152)
I described the tip of the tongue of this genus as like a stalk in
iorm (sténgelférmig). But this German word was a mistake of my
printers. I wrote “ stémpelformig”’ (shaped like a pestle), com-
paring the thick fleshy tongue of the Are (although not entirely
1878.] AND OTHER ARGENTINE PARROTS. 77
cylindrical) tothat instrument, but by no means to that of a stalk
of a flower or fruit, to which the tongue has no resemblance.
I have at this very moment under my inspection the tongue of a
large Macrocercus macao (Sittace chloroptera, Gray), which recently
died in the house of one of my friends. This tongue has on each
side, behind the two fringed lobes, with which the tongue of all
parrots terminate, several large spines.
In the tongue of Bolborhynchus murinus I find no spines in the
same place.
In volume i. of the same work, Dr. Finsch thinks it surprising
(auffallend) that I have not mentioned that Conurus nanday also
inhabits the Argentine Republic, although the specimen in Mr.
Sclater’s collection came from Buenos Aires. This latter fact may
be true; but I am still more surprised to find that Dr. Finsch thinks
that all specimens coming from Buenos Aires must necessarily have
lived in this country. The species alluded to is by no means an
inhabitant of the Argentine Republic, but comes only from Paraguay,
and is often brought to Buenos Aires alive, as the bird is here con-
sidered a great rarity. That it lives also at Bolivia I have never
heard here in Buenos Aires, though I have had a living specimen in
my room for a long while, which was very tame, and would stand on
my shoulder and remain in this position while I took long walks.
At last I gave it toa friend of mine who was going to Europe, with
the instruction that after its death it was to be put in the collection
at Halle.
The description of this bird given by Dr. Finsch is not entirely
correct. The head has no blackish brown on the front and vertex,
but is entirely black, the brown colour being the consequence of the
skin being old and dry.
In vol. ii. part 1, p. 116, of Dr. Finsch’s work, the Bolborhynchus
monachus is named “calita.’ The orthography of this term is
erroneous, the bird, which is very common in all parts of the Argen-
tine Republic, being called always catita. Again, Dr. Finsch says
the two sexes do not differ; but I find the breast of the male of a
much clearer grey than that of the female, and that it has very dis-
tinct, darker and clearer transverse stripes, while the latter are
wanting in the female, which has an obscure grey-coloured breast,
with a trace of brown-grey, which is not seen at all in the male.
In the young state both sexes are alike.
In vol. i. p. 126, Dr. Finsch describes my Bolborhynchus
rubrirostris from the young specimen sent by myself to the Bremen
Museum, and blames me for not mentioning the blue-grey-greenish
colour on the throat and breast. But this colour is not persistent,
but is only found in the young bird (of the first year). The adult
bird is entirely clear green on the breast, the same as on the neck
and the whole body. From the young state of the specimen ex-
amined by Dr. Finsch results also the discrepancy in the length of
the wing, which measures 5 inches in the old bird, and not 3” 10'”
as he says.
78 COUNT T. SALYADORI ON THE {Jan. 15,
6. Reports on the Collection of Birds made during the
Voyage of H.M.S. ‘ Challenger.—No.VI. On the Birds
of Ternate, Amboyna, Banda, the Ké Islands, and the
Aru Islands. By T. Satvaport, C.M.Z.S.
[Received November 26, 1877.]
The collection of birds made at the above-mentioned localities is
not very large, numbering only 131 specimens, belonging to 79 species.
None of these are new to science ; but some are of special interest, as
having been described quite recently or as being new to the locality
where now they have been found.
I propose to treat of them separately, according to the localities.
I. Birds from Ternate.
[The ship arrived at Ternate on the evening of the 14th October,
1874, and sailed from Ternate at 11 a.m. on the 17th Oct. 1874.
—J. M.)
The birds are only 12 in number, and belong to 10 species; a
specimen of Lorius cyanauchen, although bought alive at Ternate,
must have been brought there from a very different and far-away
locality.
1. Scors moroTeEnsis, Sharpe.
Scops morotensis, Sharpe, Cat. Strig. p. 75, pl. vii. f. 1 (1875),
(type examined).
[No. 325. Male: eyes yellow; stomach contained large grass-
hoppers.—J. M.]
One “male” (No. 325) in the dark rufous plumage.
All the specimens from Ternate which I have seen agree better
with the form from Morty than with that from Halmahera, S. /euco-
spilus (G. R. Gr.).
I know of other instances in which the birds inhabiting the small
islands surrounding Halmahera, although very far one from the
other, agree better inter se than with the form inhabiting the main-
land.
2. TANYGNATHUS MEGALORHYNCHUS (Bodd.).
Psittacus megalorhynchus, Bodd. Tabl. Pl. Enl. p. 45 (1783)
(ex Buffon, Pl. Enl. 713).
Eclectus megalorhynchus, Schleg. ; Finsch, Die Papag. ii. p. 351
(1870).
[No. 337. Male: eyes white; bought alive at Ternate.—J. M.]
3. Eciectus roratus (Miill.).
Psittacus roratus, Mill. S. N. Suppl. p. 77 (1776) (2); Cass.
Pr. Ac. Nat. Sc. Philad. 1864, p. 240. no. 52.
Psittacus grandis, Gm. S. N. i. p. 335. no. 102 (1788) (2).
Eclectus grandis, Wagl.; Finsch, Die Papag. ii. p. 340 (1868).
1878. ] BIRDS OF TERNATE, AMBOYNA, ETC. 79
Eclectus polychlorus, ex Batchian, Halmahera et Ternate, auctt.
(nec Scop.) (¢); Finsch, Die Papag. ii. p. 393 (partim) (1868).
(336. Male: eyes yellow. 361. Female: this is the female of
No. 336 ; it was purchased along with others at Ternate and kept
alive on board till the 28th Oct., when it was killed and skinned, as
it proved a nuisance while alive.—J. M.]
Two specimens bought alive at Ternate, a “ male” (No. 336)
green and a “ female ’’ (No. 361) red; this one is very interesting,
having some green feathers among the scapularies.
Both specimens are much smaller than the birds which I have
seen from Halmahera.
4. Lorius pomice Lia (Linn.).
Psittacus domicella, Linn. S. N. i. p. 145. n. 26 (1766).
Domicella atricapilla, Wagl.; Finsch, Die Papag. ii. p.763 (1868).
[No. 325. Male: eyes red.—J. M.]
One “male” (No, 335) not fully adult, bought alive at Ternate }.
6. CyANALCYON pDiops (Temm.).
Alcedo diops, Temm. Pl. Col. 272 (1824) (type examined).
Halcyon diops, Steph. ; Sharpe, Mon. Alced. pl. 77 (1868-187 y:
[No. 334. Male: eyes black ; stomach had insects.—J. M.]
7. PACHYCEPHALA MENTALIS, Wall.
Pachycephala melanura, part., Bp. Consp. i. p. 388 (ex Moluccis)
(1850) ; G. R. Gr. (nee Gould), P. Z. S. 1860, p. 353; Schleg.
Ned. Tijdschr. voor de Dierk. iv. p. 44 (1871).
Pachycephala albicollis, Temm. Mus. Lugd. (nec Lafr.) (Bp. l. ¢.).
Turdus armillaris, Temm. (Bp. J. ¢.).
Lanius cucullatus, Licht. (Bp. J. ¢.).
Pachycephala mentalis, Wall. P. Z.S. 1863, p. 30 (type examined).
Pachycephala nigrimentum, G. R. Gr. Hand-list, i. p. 388. sp.
5880 (1869), descr. nulla (type examined).
[No. 333. Male: stomach had insects.—J. M.]
8. Cyrrostomus FRENATUS (S. Miill.). :
Nectarinia frenata, S. Mill. Verh. Land- en Volkenk. p- 173
(note) (1843).
Cinnyris frenatus, Shelley, Mon. Cinnyr. pt. iii. (1877).
Cyrtostomus frenatus, Salvad. Atti R. Ac. Sc. Tor. xii. p- 317
(1877).
? 5. Lortus cyanaucuen (Miill.).
Psittacus cyanauchen, 8, Mill, Verhandl. Land- en. Volkenk, p. 107 (1839-
1844) (type examined).
Domicella cyanauchen, Finsch, Die Papag. ii. p. 773 (1868).
Domicetla lory mysorensis, Meyer, Sitzb. k. Ak. Wiss, Wien, xx. p. 233 (1874).
[No. 335. Male: eyes red.—J. M.]
One ‘“‘male” bought alive at Ternate, but certainly brought there from the
Island of Mysore, in Geelvink Bay, where only L. cyanauchen lives in the
wild state. :
80 COUNT T. SALVADORI ON THE {Jan. 15,
[327. Male: eyes brown; stomach had insects. 329. Female.
—J. M.)
9. Hermorimia auricers (G. R. Gr.).
Nectarinia auriceps, G. R. Gr. P. Z.S. 1860, p. 348.
Hermotimia auriceps, Salvad. Atti R. Ac. Se. Tor. x. p. 228
(1874).
Cinnyris auriceps, Shelley, Mon. Cinnyr. pt. v. (1877).
[No. 326. Male: eyes brown; stomach had insects.—J. M.]
10. ToTanus GLAREOLA (Linn.).
(No. 330. Female: eyes brown; stomach contained shells.—
J. M.)
Il. Birds from Amboyna.
[The ‘Challenger’ arrived at Amboyna on Sunday 4th Oct., 1874,
and sailed from Amboyna on the 10th Oct., 1874, at 4 p.m..—J. M.]
During the stay 23 birds were procured, representing 15 species,
as follows’ :—
1. Urospizias 10GasTer (S. Mill.)
Falco hiogaster, S. Miill. Verh. Land- en Volkenk. p. 110, note
(1839-1844) (type examined).
Astur hiogaster, Sharpe, Cat. Accipitr. p. 104 (1874).
[No. 302. Male: eyes coral-red ; legs orange; stomach had fish-
bones and crabs.—J. M.]
2. Grorrroyus RHOoDOPS (G. R. Gr.).
Psittacus fuscicapillus, Vieill. N. D. xxv. p. 316, Java! (1817),
(descr. falsa; type examined) (9).
Psittacus rhodops, G. R. Gr. P. Z. 8. 1861, p. 436, Waigiou ! and
Mysol! (descr. nulla; type examined).
Eclectus rhodops, Schleg. Mus. P.-B. Psittaci, p. 43 (1864),
Ceram, Amboyna, and Bouru.
Pionias rhodops, part., Finsch, Die Papag. ii. p. 380 (1868).
Geoffroyus schlegelit, Salvad. Ann. Mus. Civ. Gen. x. p. 29. no.
40, Buru, Boano, Ceram, Amboyna, Arouko, Goram, Monawolka
(1877).
[No. 304. Male: eyes white; bill, upper part reddish, lower
blackish.—J. M.]
This species resembles G. aruensis (G. R. Gr.); but it is larger,
has the under wing-coverts of a darker blue, and the female has. a
darker brown head.
The type of Psittacus fuscicapillus, Vieill., is certainly a female of
this species ; but the description is wrong, as also the locality.
Psittacus rhodops, G. R. Gr. (1. ¢.), was said to be from Waigiou
and Mysol; but knowing that in these islands lives a representative
1 [Besides the skins examined by Count T. Salvadori, two Cassowaries were
procured alive, and the skins afterwards preserved. They appear to be imma-
ture examples of Caswarius galeatus, or of a very closely allied species.—P. L. 8.]
1878. ] BIRDS OF TERNATE, AMBOYNA, ETC. 81
form, Geoffroyus pucherani, Bp., I thought that it could not be the
same bird as Eclectus rhodops, Schleg., from the Ceram group ;
accordingly I gave to Schlegel’s species the name of Geoffroyus
schlegelit. Having, however, quite recently seen Gray’s types in the
British Museum, I find that they really belong to the Ceramese
species; and I am quite sure that the localities Waigiou and Mysol
are wrong. Those specimens no longer bear Wallace’s original labels,
which circumstance may explain how it was that wrong localities
have been attributed to them.
3. Cryx Lepipa, Temm.
Ceyx lepida, Temm. Pl. Col. 595. f. 1, Amboyna (1836) ; Sharpe,
Mon. Alced. pl. 46 (pt. ii. 1868).
[No. 303. Female: bill and legs red; eyes black ; stomach had
insects.— J. M.]
4, MacropTeryx MysTacea (Less).
Cypselus mystaceus, Less. Voy. Coq. Zool. Atlas, pl. 22 (1826)
(type examined).
Macropteryx mystacea, Salvad. Ann. Mus. Civ. Gen. x. p. 311.
sp. 1 (1877).
[298. Male; 299. Female: eyes black; stomach had insects.
—ZJ. M.]
Two specimens—one ‘‘ male’’ (No. 298), with the brown spot on
the ear-coverts, and one female (No. 299), without the brown spot.
5. CyrTosToMus ZENOBIA (Less.).
Cinnyris zenobia, Less. Voy. Coq. Zool. Atlas, pl. 30. f. 3 (1826),
i. 2, p. 679 (1828)*.
Cinnyris clementie, Less. Dict. Se. Nat. i. p. 18 (1827).
Cyrtostomus melanogaster, G. R. Gr. Hand-list, i. p. 112. sp.
1390 (1869), descr. nulla.
Cyrtostomus zenobia, Salvad. Atti R. Ac. Sc. Tor. xii. p. 319
(1877).
[No. 308. Male; 312. Female; 313. Male: eyes black; stomach
contained insects.—J. M.
Three “ males” (No. 308, 312, 313), apparently fully adult; all
three have the feathers of the forehead tipped with metallic blue;
one of them (No. 313) has the upper parts more greenish than the
other two, which have the same parts of a rich olive-brown.
6. Hermorimia aspastorpes (G. R. Gr.).
Nectarinia aspasioides, G. R. Gr. P. Z.S. 1860, p. 348, Am-
boyna.
Hermotimia aspasioides, Salvad. Atti R. Ac. Sc. Tor. x. p. 208,
216 (1874), xii. p. 306 (1877).
Cinnyris aspasioides, Shelley, Mon. Cinnyr. pt. v. (1877).
‘ T have not been able to find the type of this species in the Museum of Paris.
Proc. Zoo. Soc.—1878, No. VI. 6
82 COUNT T. SALVADORI ON THE [Jan. 15,
(314. Male: eyes black. 312. Female: eyes black ; stomach
had insects.—J. M.]
Two specimens—one “ male” (No. 314) not fully adult, with the
green cap mixed with blue, and one “‘ female” (No. 312).
7. Dic#uM VULNERATUM, Wall.
Diceum vulneratum, Wall. P. Z. 8. 1863, p. 32, note (Ceram ;
type examined).
[No. 315. Female: eyes brown; stomach had insects and seeds.
Small bird.—J. M.]
This sex has not yet been described.
Fem. Superne cinereo-fusca, alis et cauda obscurioribus, paullum
nitentibus ; pileo presertim frontem versus brunneo tincto ;
supracaudalibus rubris ; inferne cinerea; gastreo medio seu
gula, colli antici, pectoris abdominisque parte media et subcau-
dalibus albis ; rostro et pedibus fuscis.
8. BupyTes viripis (Gm.).
[305. Male: eyes brown; legs and bill black; stomach had
insects. 310. Female: eyes brown. 319. Male: eyes brown ;
stomach had insects.—J. M.]
Three specimens—one “ male” (No. 305) and one “female” (No.
310) scarcely different, and one ‘‘ male” (No. 319) with the under-
parts of a paler yellow, and with the back rather greyish.
9. MacropyGiA AMBOINENSIS (Linn.).
[301. Male: eyes red; feet red; bill black ; fruit in stomach.
320. Female: eyes orange; stomach had seeds; legs red.—J. M.]
Two specimens—one fully adult “male” (No. 301), and one
“female” (No. 320). The female has the cap of a uniform cinna-
mon colour, the breast more reddish than the male, and no green
or purplish reflections on the nape, which is finely mottled with
dark brown and reddish fulvous.
10. CHaraApRIvs FuLVUs (Gm.).
[No. 317. Male: eyes hazel; stomach contained worms and
land-shells.—J. I. ]
11. EvpromiAs veREDUus (Gould).
Charadrius veredus, Gould, P. Z. 8S. 1848, p. 38; id. B. Austr.
vi. pl. 14.
Eudromias veredus, Harting, Ibis, 1870, p. 209, pl. vi.
[No. 318. Male: eyes hazel; stomach contained worms and land-
shells.—J. M.]
12. AlGIALITIS GEOFFROYI (Wagl.).
Charadrius geoffroyi, Wagl. Syst. Av. gen. Charadrius, sp. 19
(1827).
Ayialitis geoffroyi, Harting, Ibis, 1870, p. 378, pl. xi.
[No. 311. Male: eyes brown.—J. M.]
1878.] BIRDS OF TERNATE, AMBOYNA, ETC. 83
13. TRINGOIDEs HYPOLEUCUs (Linn).
[No. 306. Female: eyes hazel; stomach had insects.—J. M.]
14. GALLINAGO sTeNURA (Kuhl).
“‘Scolopax stenura, Kuhl,” Bp. Ann. Stor. Nat. iii. fase. x (1830).
Gallinago stenura, Schleg. Mus. P.-B. Scolopaces, p. 12 (1864) ;
Salvad. Cat. Syst. Ucc. di Borneo, p. 334. sp. 353 (1874).
[No. 316. Male: eyes black ; stomach had worms.—J. M.])
15. Burorimes savanica (Horsf.).
Ardea javanica, Horsf. Trans. Linn. Soc. xiii. p- 190 (1820) ;
Schleg. Mus. P.-B. Ardea, p. 43 (1863).
Butorides javanica, Salvad. Cat. Uce. di Borneo, p. 351. sp. 370
(1874).
[No. 300. Male: eyes orange; legs yellow; bill black; crustacea
in stomach. 309. Female: eyes yellow; legs green.—J. M.]
Two specimens—one “male” (No. 300) fully adult, and a young
“female”? (No. 309).
Ill. Birds from Banda.
[The ‘ Challenger’ arrived at Banda on the 29th September, 1874,
and sailed from Banda on the evening of the 2nd October, 1874.—
J. MJ
Only 10 specimens, belonging to 7 species, are in the collection,
which, although a very small one, is of some interest, as birds from
Banda are rather scarce in Museums.
1. SauropaTis cHLorts (Bodd.).
[No. 293. Male; 294. Female: eyes black; legs have a dull
brown tinge; base of the lower mandible white, the rest black.—
J. M.)
2. Monarcua rnornatus (Garn.). *
Muscicapa inornata, Garn. Voy. Coq. Zool. Atlas, pl. 16. f. 2
(1826), & i. 2, p. 591 (1828), New Guinea (type examined).
Drymophila cinerascens, Temm. Pl. Col. 430. f. 2 (1827), Timor
(type examined).
Monarcha fulviventris, Hartl. P.Z.S. 1867, p. 830, Echiquier
Islands (type examined).
[No. 292. Male: eyes brown; legs and bill slate-blue; the bill
lighter than legs ; stomach contained insects.—J. M.]
After having examined the types mentioned above and having
compared together many specimens from Timor, Banda, Amboyna,
Goram, Tijoor, Ternate, Halmahera, Batchian, Tifore, Sula, Ké
Islands, Aru Islands, Mysol, New Guinea (Sorong, Dorei, and Man-
nam), Mafor, the Admiralty Islands, and the Echiquier group, I
have arrived at the conclusion that they all belong to one species.
The Drymophila cinerascens, Temm., represents the fully adult bird ;
6*
84 COUNT T. SALVADORI ON THE (Jan. 15,
and the Muscicapa inornata, Garn., is the young stage of the same,
differmg from the adult in the reddish colour of the outer edgings
of the remiges. Under the name of M. fulviventris Hartlaub has
described a specimen from the Echiquier group, with the abdomen
rather pale, exactly like others from the other localities mentioned
above.
3. Rurprpura squamarta, S. Mill.
Rhipidura squamata, S. Mill. Verh. Land- en Volkenk. p. 189
(1839-1844), Banda (type examined).
Rhipidura griseicauda, Salvad. Ann. Mus. Civ. Gen. vii. p. 924.
sp. 20 (1875), Waigiou.
[No. 289. Female: eyes hazel; stomach contained insects.—
J. M.]
This species is allied to R. rufifrons (Lath.) from Australia, from
which it differs in having the forehead black in the middle and
white on the sides ; the tail is grey, white-tipped, and with scarcely
any reddish colour near the base. S. Miller described this species
with the tail brown-black, instead of grey; and this mistake
made me believe that the bird from Waigiou, described by me, was
different.
4. PachycerHALa pH xonoTa (Miill.).
Myiolestes phajonotus, Mill. Mus. Lugd. (type examined) ; Bp.
Consp. i. p. 358 (1850), Banda.
Pachycephala brunnea, Wall. P. Z.S. 1865, p. 478, Banda, Sal-
watty (type examined).
Pachycephala senex, Pelz. Verh. k. zool.-bot. Gesellsch. Wien,
1872, p. 429, Aru (type examined).
[No. 290. Male; 291. Female: eyes light hazel; legs flesh-
colour ; stomach contained insects.—J. M.]
The sexes are exactly alike.
The types in the Museum of Leyden are marked “ Borneo,” which
is a mistake ; Bonaparte has given the exact locality.
This species has a very extensive range ; I have seen specimens
from Banda, Ceram-laut, Mysol, Salvatty, Waigiou, Mafor, Ternate,
Dammar, Mareh, Motir, and Tifore. It is not quite certain that
the bird described by Von Pelzeln (/. ¢.) was really from the Aru
Islands.
5. Myzome a Borel, 8. Mill.
Nectarinia (Myzomela) boiei, 8. Mill. Verh. Land- en Volkenk.
p- 172 (1839-44) (type examined).
[No. 295. Female: eyes brown ; stomach had insects. The
male of this bird had a red head ; but no specimen was shot.—J/. M.]
6. ZosTEROPS CHLORIS, 8. Mill.
Zosterops chloris, S. Mill. in Mus. Lugd. (type examined) ; Bp.
Consp. i. p. 398 (1850); Hartl. Journ. f. Orn. 1865, p. 24.
[Ne. 287. Male; 288. Female: eyes brown; legs slate-colour ;
1878.] BIRDS OF TERNATE, AMBOYNA, ETC. 85
stomach contained insects. In flocks on trees near the shore.—
J. M.}
The sexes are exactly alike.
To this species have been referred also specimens from Ternate
and Bouru ; but those from Ternate agree better with Z. intermedia,
Wall., from Celebes ; and one specimen from Bouru in the British
Museum, collected by Wallace, belongs to a species not yet described.
7. CARPOPHAGA CONCINNA, Wall.
Carpophaga chalybura, G. R. Gr. (nee Bp.), P. Z. 8. 1858, pp.
187, 196 (Aru).
“Carpophaga chalybea, Bp.” (errore), Rosenb. Journ. f. Orn. 1864,
p. 132. sp. 226 (Aru).
Carpophaga concinna, Wall. Ibis, 1865, p- 383. no. 52, Matabello
Island, Aru (a small island west of ) (type examined) ; Salvad. Ann.
Mus. Civ, Gen. ix. p. 200. no. 39 (1876).
Carpophaga roseinucha, patt., Schleg. Ned. Tijdschr. Dierk. iii.
p- 197 (1865).
[No. 296. Female. This bird was sent on board alive as a present
to the Professor.—J. M.]
IV. Birds from the Ké Islands.
[The ship arrived at Ké Doulan, Ké Islands, on the morning of
the 25th September, 1874, and left on the morning of the 26th
September.—J. M.]
The birds from Ké Islands are 25, representing 14 species; they
have been all collected at the little island named Dulan’.
The Ké Islands were quite unknown to naturalists before Wallace
made a short stay there of six days in January 1857. He only
obtained thirteen species in all?, which were catalogued, together with
those from the Aru Islands, by Mr. G. R. Gray*. Von Rosenberg
made a longer stay on the Ké Islands, and collected there rather
extensively ; but no complete catalogue of his collection has been
published. He gave an account of his visit to the Ké Islands, and
of some of the birds collected there by him, in his ‘ Reis naar de
zuidoostereilanden gedaan en 1865,’ pp- 67-83.
Dr. Beceari in 1873 made a rather large collection of birds in
the Ké Islands. Of this some of the new species have been already
described by me‘; but I propose to give a complete account of it as
soon as my occupations permit.
* Cosmos di Guido Cora, ii. tay. ii. (1874).
? Ann. & Mag. Nat. Nist. (2), xx. p. 473 (1857).
° P.Z. 8. 1858, pp. 169-198.
+ “Nuove specie di uccelli delle Isole Aru e Kei, raccolte da Odoardo Beccari,”’
Ann. Mus, Ciy. Gen. vi. pp. 73-80 (1874) ; “Altre nuove specie di uccelli raccolte
nella Nuova Guinea dal Signor L. M. D’Albertis e nelle Isole Aru e Kei dal Dott.
O. Beceari,” op. cit. vi. pp. 308-314; “Deserizione di sei nuove specie di
uccelli delle Moluche, delle Kei e delle Isole Aru,” op. cit. vii. pp. 983-990
(1875).
86 COUNT T. SALVADORI ON THE [Jan. 15,
1. PANDION LEUCOCEPHALUs, Gould.
Pandion leucocephalus, Gould, P. Z. S. 1837, p. 138; id. B.
Austr. i. pl. 6 (1848); Sharpe, Cat. Acciptr. p. 451 (1874).
[No. 268. Male: eyes yellow; legs whitish ; bill black ; stomach
contained fish. Shot by Chauner; no others were seen. Shot far
in the bush from the shore.—J. 27/.}
2. GEOFFROYUS KEYENSIS, Schleg.
Psittacus capistratus, G. R. Gr. (nec Bechst.), P. Z. S. 1858,
p- 183 (@), Ké Islands (type examined).
Eclectus personatus (part.), Schleg. Ned Tijdschr. Dierk. iii. p.
332 (A. grande forme (partim), p. 333) (1866); id. Mus. P.-B.
Psittaci, Revue, p. 18 (partim) (1875).
Pionias rhodops (part.), Finsch, Die Papag. ii. pp. 388, 955
(1868).
Geoffroyus keyensis, Schleg. MS. (Finsch, Die Papag. ii. p. 956) ;
Salvad. Ann. Mus. Civ. Gen. x. p. 29. sp. 37 (1877).
[No. 271. Male; 269, 270. Females : eyes white ; upper mandible
red in the male, both black in the females ; stomach contained seeds
and fruit.—J. M.]
Three specimens—a young “male”? (No. 271) in transitional
plumage, an adult “female” (No. 270) with the head pale brown
olive, and a not fully adult “female” (No. 269) with the head
conspicuously tinged with green.
3. Eos rusra (Gm.)
Domiceila rubra, Finsch, Die Papag. ii. p. 786 (1868).
Psittacus (Eos) bernsteinii, Rosenb. Nat. Tijdschr. v. Ned. Ind.
xxv. p. 145 (1863), Ké Islands.
Lorius squamatus, Schleg. (nec Bodd.), Mus. P.-B. Psittaci, p.
124. no. 1, 2 (1864) (types examined).
Domicella schlegelit, Finsch, Die Papag. ii. p. 792 (1868),
Ké Islands (type examined).
[No. 272. Male; 273. Female: eyes light red ; bill dullred ; legs
black ; stomach contained seeds with some insects. Only two were
shot. The Professor has one of the same species alive on board,
bought at Ké Doulan.
No. 324. Female: eyes brown; legs black; bill rose-coloured.
This bird was purchased alive at Ké Doulan, Ké Islands, and kept
till today (Oct. 12, 1874) on board, and was killed on account of
the noise it made.—J. M.]
Three specimens—one “male” (No. 272) and one “ female”
(No. 273) fully adult and alike, and a “female” (No. 324) not
fully adult, with some of the feathers of the underparts edged with
blue and turning to greenish in the hidden part.
The Domicella schlegelit, Finsch, l. ec. (= Lorius squamatus,
Schleg.), was established on some young birds, which have the
feathers of the underparts edged with blue ; the same thing happens
in the young of Z. rictniata and of E. cyanogenys.
1878. | BIRDS OF TERNATE, AMBOYNA, ETC. 87
4. Rurpipura AssIMiLis, G. R. Gr.
Rhipidura assimilis, G. R. Gr. P. Z. 8. 1858, p. 176, Ké Islands
(type examined).
[No. 286. Male: insects in the stomach; eyes black.—J. M. |
This species is allied to R. isura, Gould, from Australia, and to
R. setosa (Q. & G.) from New Ireland and New Guinea; it
differs from the first in the larger dimensions, in the paler band
across the breast, and in the spot at the tip of the two outer tail-
feathers less extended and light fulvous instead of white; from R.
setosa it is easily distinguished by the upper tail-coverts rufescent.
5. GRAUCALUS POLLENS, Salvad.
Graucalus pollens, Salvad. Ann. Mus. Civ. Gen. v. p. 75 (1874).
[No. 280. Female: eyes black; stomach contained seeds; feet
and bill black.—J. JL]
This specimen agrees entirely with the typical females described
by me. This species comes next to G. caledonicus (Gm.) from New
Hebrides and New Caledonia; but it differs in the dimensions being
somewhat smaller and in the darker colour. _ It is also allied to Gr.
personatus, Mill., from Timor; but it is larger, darker, and also
conspicuously different in the under wing-coverts, which are not
white, but dark like the rest of the plumage.
In the British Museum I have seen a Graucalus, said to have been
collected by Wallace in the Sula Islands, and wrongly named G.
temmincki, which is very much like the females from the Ké Islands,
except that it is a little smaller.
6. LauacE Karu (Less.).
Lanius karu, Less. Voy. Coq. Atlas, pl. 16 (1826).
Ceblepyris karu, Less. Voy. Coq. Zool. i. 2, p. 633 (1828) (type
examined).
Lalage karu, Hartl. Journ. f. Orn. 1865, p. 168.
[Nos. 282, 283. Males: eyes black; stomach contained in the
one seeds, in the other insects.—J. M.]
Two specimens, one “male” (No, 282) fully adult, with the
upper parts shining black, the under wing-coverts pure white, and
the breast and the abdomen barred across; this is exactly like
another specimen from Naiabui, collected by Signor D’Albertis and
marked female (!) (Ann. Mus. Civ. Gen. ix. p. 28. sp. 108, 1876).
The second specimen (No. 283) is also marked “ male:” it is not
fully adult ; it has the upper parts grey, and the under wing-coverts,
as well as the underparts, tinged with reddish.
I suspect that to the same species must be referred the Ceblepyris
rufiventris, Jacq. & Pucher., and I question whether it really was
from Raffles Bay. The type of this species, examined by me in the
Museum of Paris, agrees entirely with two specimens from the Aru
Islands, collected by Beccari, aud which are exactly like the type of
Lalage polygrammica, G. R. Gr., also from the Aru Islands.
88 COUNT T. SALVADORI ON THE [Jan. 15,
7. Dicruropsis MEGALORNIs (G. R. Gr.).
Dicrurus megalornis, G. R. Gr. P. Z. 8. 1858, pp. 199, 193,
Ké Islands (type examined).
Chibia megalornis, Sharpe, Cat. Birds in the Brit. Mus. iii. p. 242.
no. 9 (1877).
[Nos. 274, 275. Both females: eyes red; legs and bill black ;
stomach contained insects.—J. M.]
This bird and the other allied Austro-Malayan species have been
united by Mr. Sharpe to the genus Chibia, Hodgs., the type of
which, Chibia hottentotta (Linn.), has a much longer and more acute
bill, and is also very conspicuous for the tuft of silky hairs spring-
ing from the forehead—a feature which, among the Austro-Malayan
species, is only to be found in Dicrurus pectoralis, Wall., from the
Sula Islands’.
8. PHILeEMON PLUMIGENIS (G. R. Gr.).
Tropiderhynchus plumigenis, G. R. Gr. P. Z. 8S. 1858, pp. 174,
191, Ké Islands (type examined).
[No. 276. Male; 277. Sex uncertain: eyes brown; stomach
contained seeds.—J. M.]
Both are young birds, with the feathers of the sides of the breast
with broad yellow edges, and with the outer edges of the remiges
tinged with olive.
9. CALORNIS METALLICA (Temm.).
Lamprotornis metallicus,Temm. Pl. Col. 266, 1824, Amboyna
(type examined).
Calornis viridescens, G. R. Gr. P. Z. 8S. 1858, p. 181, Aru
Islands (type examined).
‘Calornis nitida, G. R. Gr. P. Z. S. 1858, p. 181, New Ireland
(type examined).
Calornis amboinensis, G. R. Gr. P. Z.S. 1858, p. 182, Amboyna
(type examined).
Calornis gularis, G. R. Gr. P. Z. S. 1861, p. 431, Mysol (type
examined).
Calornis purpurascens, G. R. Gr. Hand-list ii. p. 26. no. 6377,
(1870), Goram (type examined).
[No. 284. Female: eyes red; stomach contained seeds ; caught
by Suhm in a spider’s web.—J. M.]
One fully adult “female” (No. 284), resembling a fully adult
male.
This species has been found in nearly all the Moluccan and Pa-
puan Islands, from the Sula Islands westward to the Soloman Islands
eastward, and lives also in the northern part of Australia.
Temminck, in the text of the ‘ Planches Coloriées,’ says that his
1 T think that the Austro-Malayan species may be more properly referred to a
peculiar genus, which I propose to call Dierwropsis, characterized as follows :—
Rostrum crassum, rostro specierum generis Dissemuri simzle, sed culmine minus
acuto, crasstusculo ; cauda bifida divaricata, rectricibus eatimis superne introrsum
paullum convolutis.
1878. ] BIRDS OF TERNATE, AMBOYNA, ETC. 89
L. metallicus was from Timor and Celebes ; but this seems to be a
mistake, as the typical specimens in the Museum of Leyden, ex-
amined by me, are from Amboyna. In Celebes and in Timor lives
another species of the genus Calornis, and not C. metallica.
Specimens from the different localities do not differ specifically :
those from Cape York are somewhat more purplish, but not always
in the same way ; one from Goram, a small island near Amboyna (C.
purpurascens, Gray), is exactly like some specimens from Cape York.
Some specimens have the throat more purplish than others; one
from Mysol (C. gularis, Gray) cannot be separated from others from
Halmahera and Cape York.
10. CARPOPHAGA CONCINNA, Wall.
[Nos. 263-264. Males; 265, 266, 267. Females: eyes between
orange and a coral-red ; the stomach in all contained fruit and seeds ;
legs coral-red; bill black. There were a great many of these Nut-
meg Pigeons shot, enough for lunch for the whole ward-room. They
were very good eating.—J. M.|
11. Myristricrvora BICOLOR (Scop.).
Myristicivora bicolor, Salvad. Ann. Mus. Civ. Gen. ix. p. 276
(1877).
[No. 278. Male: white Pigeon ; eyes brown; stomach contained
fruit; feet blue; bill black.—J. M.]
This bird has the under tail-coverts entirely white.
In a recent paper of mine’ I have shown that M. dzcolor (Scop.),
M. spilorrhoa (G. R. Gr.), and M. melanura, G. R. Gr., are per-
fectly distinct species, and that M. bicolor (Scop.) most likely is a
migratory bird in the Moluccas, as it is in Celebes, and that, while
in Celebes it is to be found together with M. luctuosa (Reinw.), in
the Moluccas it lives together with M. melanura, Gray.
12. MacropyG1a KEYENSIs, Salvad.
Macropygia, sp., Wall. Ann. & Mag. Nat. Hist. (2) xx. p. 473
(partim), 1875, Ké Islands.
Macropygia phasianella, part., G. R. Gr. (nec Temm.), P. Z. 8.
1858, pp. 187, 196, Ké Islands; Wall. Ibis, 1865, p. 389. sp. 81
(partim) ; Rosenb. Reis naar zuidoostereil. p. 81 (1867).
Macropygia turtur, part., Schleg. Mus. P.-B. Columéa, pp. 110,
113, 114 (specimina Ins. Key), 1873.
Macropygia keyensis, Salvad. Ann. Mus. Civ. Gen. ix. p. 204.
sp. 64 (1876).
No. 279. Male: eyes hazel; stomach contained seeds; legs red ;
bill black.— J. M.]
I named this Ké-Islands species (J. s. c.) before seeing a fully adult
bird, feeling quite confident, from Schlegel’s description, that it was
really distinct. I have recently seen the adult specimens con-
tained in the Leyden Museum, and I am satisfied that the Ké-
1 « Qsservazioni intorno alle speci del genere Myristictvora, Rehb.” (Ann,
Mus. Ciy. Gen, ix. pp. 265-277).
90 COUNT T. SALVADORI ON THE {Jan. 15,
Islands bird belongs to a peculiar form. I shall now give a full
description of the adult bird collected during the ‘ Challenger’ expedi-
tion, and shall proceed after to point out the principal differences
from the allied forms.
Capite rufo-vinaceo, sincipite et genis pallidioribus ; vertice et
occipite grisescentibus ; collo postico viridi et purpureo nitente,
conspicue cinerascente ; dorso, uropygio, supracaudalibus alarum-
que tectricibus exterioribus brunneo-cinnamomeis, punctulis
nigris obsoletis adspersis ; gula rufescenti-albida ; collo antico
et pectore summo vinaceis, fasciolis transversis nigris notatis ;
pectore imo et abdomine albido-fulvescentibus, fasciolis trans-
versis nigris notatis; abdominis lateribus et subcaudalibus
brevioribus pallide cinnamomeis, nigro transfasciolatis ; sub-
caudalibus longioribus et tectricibus alarum inferioribus cinna-
momeis unicoloribus ; remigibus fuscis, exterius cinnamomeo
limbatis ; cauda superne brunneo-grisescente, inferne palli-
diore et magis grisescente ; rectricibus tribus utrinque latera-
libus partim lete rujis, fascia lata subapicali nigra notatis ;
rostro nigro; pedibus rubris.
Long. tot. circa 0™370; al. 0"-018; caud. 07190; rostr.
0™-018 ; tarsi 0"-021.
This species is intermediate between M. amboinensis (Linn.) and
M. doreya (Bp.).
It differs from UW. amboinensis (which it much resembles in the
light underparts) :—
Ist. In the greyish tinge of the vertex, of the occiput, and of the
hind neck ;
2nd. In the more vinaceous colour of the frout neck ;
3rd. In the more whitish underparts ;
4th. In the darker hue of the back ;
5th. In the larger dimensions.
From M. doreya, Bp., to which it bears a great resemblance in
the upper parts, it differs :—
lst. In the more greyish hind neck ;
2nd. In the much less vinaceous front neck and upper breast ;
3rd. In the more whitish underparts ;
4th. In the lighter and more greyish tail ;
5th. In the larger dimensions.
13. TRINGOIDES HYPOLEUCUS (Linn.).
[No. 285. Eyes black; stomach contained portion of shells.—
J. M.]
14. NumMEnrvus vRopyGistis, Gould.
[No. 281. Female: eyes black; stomach contained crustacea
(crabs).—J. M.]
In my Catalogue of the Birds of Borneo I have referred this
eastern form to N. ph@opus (Linn.); but now I think that N.
uropygialis, Gould, with the uropygium constantly mottled with
dark grey, is-entitled to specific rank.
1878.] BIRDS OF TERNATE, AMBOYNA, ETC. 91
V. Birds from the Aru Islands.
[The ship arrived at Dobbo, Aru Islands, about noon on the 16th
September, 1874. The same afternoon some birds were shot near
the village of Dobbo; these were all put in spirits. On the 17tha
large party landed on the island of Wokan and shot thirty-two
specimens. On the 19th September thirteen specimens were shot at
Wanumbai, and on the afternoon of the 22nd and morning of the
23rd seventeen more. The ship left Aru on the morning of the
23rd September, 1874, at 6 p.m.—J. M.]
The birds collected in the Aru Islands are 61, belonging to
40 species; they are from three different localities—Wokan, Wa-
numbai, and Dobbo, already visited by Wallace, Von Rosenberg, or
Beccari.
Only one species, Drymacedus beccarii, lately described by me from
the Arfak Mountains, is new to the islands.
1. CuncuMA LEUCOGASTER (Gm.).
Haliaétus leucogaster, Sharpe, Cat. Accipitr. p. 307 (1874).
[No. 262. Female: eyes yellow; stomach contained fish. Shot
on Wokan on 21st September. Skinned by Crosbie.—J. M.]
This bird is in transitional plumage, like the specimen figured by
Schlegel, Valkvog. pl. 4. f. 2.
2. HaLIASTUR GIRRENERA (Vieill.).
Haliaétus girrenera, Vieill. Gal. Ois. i. p. 31 (syn. emend.), pl. x.
(1825) (type? examined).
[No. 235. Male: eyes red; stomach contained crabs and cuttle-
fish. Shot on Wokan.—J. M.]
I have found in the Museum of Paris a specimen marked “ Papous
(Freycinet, 1820),” which seems to be the type of Vieillot.
3. Urospizias POLIOCEPHALUs (G. R. Gr.).
Accipiter poliocephalus, G. R. Gr. P.Z.S. 1858, pp. 170, 189,
Aru Islands (type examined); Sclat. Ibis, 1859, p. 276, 1860,
p- 322, pl. x. (fig. avis adulte tantum) ; Rosenb. Nat. Tijdschr.
Ned. Ind. xxv. p. 228. n. 47 (1863); id. Journ. f. Orn. 1864,
p- 116. n. 47; Kaup, P. Z.S. 1867, p. 177; Wall. Ibis, 1868,
p- 12 (partim); Salvad. Ann. Mus. Civ. Gen. vii. p. 75, sp. 7,
Ansus (1875).
Nisus poliocephalus, part., Finsch, New Guin. p. 155 (1865); |
Schleg. Ned. Tijdschr. Dierk. iii. p. 326 (partim) (1866); id. Mus.
P.-B. Revue Accipitr. p. 88 (1873); Gieb. Thes. Orn. ii. 711
(1875).
Leis poliocephalus, G. R. Gr. Hand-list, i. p. 34. sp. 336
1869).
: Astur poliocephalus, Rosenb. Reis naar zuidoostereil. p. 36 (1862);
Sharpe, Cat. Accipitr. p. 117 (specim. a, 6 tantum) (1874).
Urospizias poliocephalus, Gurn. Ibis, 1875, p. 365.
92 COUNT T. SALVADORI ON THE [Jan. 15,
Astur contumax, Rosenb. Reis naar zuidoostereil. p. 36 (Aru),
descr. nulla (1867).
Trospizias spilothoraz, Salvad. Ann. Mus. Civ. Gen. vii. p. 900.
sp. 1 (1875), Arfak and Jobi, x. pp. 115, 117. sp. 5 (1877).
Accipiter etorques, part., Sclat. P. Z. S. 1877, p. 109. n. 45 (young
specimen).
[No. 253. Male, from Wanumbai.—J. M.]
This bird is young.
I have found in the museums of Leyden and Berlin two specimens
which are very much like my U. spilothorax; but they have only a
few longitudinal dark spots on the lower part of the breast, showing
that these spots are not a permanent character, and that my U. spilo-
thorax represents only a stage of U. poliocephalus.
4. MicroGLossus aTERRIMUS (Gm.).
[No. 206. Wokan. Male: eyes black; cheek-membrane red;
stomach contained many of the insides of kernels of nuts. This was
shot on the top of one of the highest trees. Only one seen at the
time.—J. M. |
I have seen many specimens from the Ara Islands, and these are
in general much smaller than those from New Guinea; and I think
that the type of Psittacus alecto, Temm., which I have seen in the
Museum of Leyden, must have come from the Aru Islands’. I do
not think that there is more than one species of the genus Micro-
glossus.
5. CYCLOPSITTACUS MELANOGENYs (Rosenb.).
*‘Psittacula melanogenia, Rosenb.” Schleg. Ned. Tijdschr. v.
Dierk. iii. p. 330 (1866), Aru (type examined); Rosenb. Nat. Tijdschr.
Ned. Ind. xxix. p. 142 (1867).
Psittacula melanogenys, Finsch, Die Papag. ii. p. 626 (1868).
Cyclopsittacus melanogenys, Salvad. Ann. Mus. Civ. Gen. x. p. 29,
sp. 33 (1877).
[No. 207. Wokan. Female: eyes black; legs with a greenish
tinge.
No. 208. Wokan. Probably a male; but the specimen was too
much injured to determine.—J. M. |
The example, without indication of the sex, is most likely a female,
haying the ear-coverts orange.
6. EcLectus potycHLorus (Scop.).
Psittacus polychlorus, Scop. Del. Flor. et Faun. Insubr. p. 87,
n. 27 (1786) (ex Sonnerat), 3.
Eclectus linnei, Wag). Mon. Psitt. p. 571, t. xxi. (1832), 9.
Eclectus polychlorus, var. aruensis, G, R. Gr. P. Z. 8. 1858, p.
183.
' The specimen in the Berlin Museum, which has been regarded as a second
one of M. alecto, looks smaller than the New-Guinea specimens from having
the primaries cut off.
1878.] BIRDS OF TERNATE, AMBOYNA, ETC. 93
Eclectus polychlorus, Salvad. Ann. Mus. Civ. Gen. x. p. 31. sp. 7
(1877).
[No. 212. Wokan. Male: eyes orange; upper mandible red
towards the base, and whitish-yellow towards the tip; lower one jet-
black; stomach contained inside-coating of cocoa-nut, and some
species of comminuted red coral (tubular coral) ; feet black.—J. M.|
Mr. G. R. Gray considered the Aru specimens as forming a pecu-
liar race of Z. polychlorus, on account of the yellowish band at the
tip of the tail being more conspicuous and tinged with reddish; but
the differences are very slight and not constant.
7. CHALCOPSITTACUS SCINTILLATUS (Temm.).
Psittacus scintillatus, Temm. Pl. Col. p. 569 (1835), Lobo, juv.
(type examined).
Chalcopsitta rubrifrons, G. R. Gr. P. Z. 8. 1858, p. 182. pt. 135
(ad.), Aru (type examined).
Domicella scintillata, Finsch, Die Papag. ii. p. 752 (1868).
Chaleopsittacus scintillatus, Salvad. Ann. Mus. Civ. Gen. x.
p- 34, sp. 75 (1877).
[No. 213. Wokan. Male: eyes dull red; feet black; both man-
dibles black ; stomach quite empty.
No. 229. Wokan. Male: eyes orange; feet and bill black;
stomach contained insects and leaves.—J. M.]
Both specimens are fully adult.
8. TRICHOGLOSSUS NIGRIGULARIS, G. R. Gr.
Trichoglossus nigrigularis, G. R. Gr. P. Z. 8. 1858, p. 183, Aru
(type examined).
Trichoglossus cyanogrammus, part., Finsch, Die Papag. ii. 830
(1868).
[No. 237. Male, Wokan: eyes red, bill red; legs slate-colour ;
stomach contained small seeds. Shot at Wokan.—QJ. M.]
This species is confined to the Aru Islands and to the Ké Islands,
and it is quite distinct from TJ’. cyanogrammus (Wagl.), with which
it has been assimilated by Wallace, Schlegel, and Finsch.
I have seen many specimens of both species, and I have found that
constantly T. nigrigularis differs from T. cyanogrammus :—
Ist. In the red of the breast inclining to orange, paler and with a
most conspicuous yellow tint near the dark edges of the feathers.
2nd. The dark edges of the breast-feathers are much narrower,
as in 7. massene.
3rd. The head above has only two colours, blue in the anterior
half, greenish in the other half; while in 7’. cyanogrammus the head
above has three colours, blue on the sinciput, green on the vertex,
and purplish-brown on the occiput, like 7. massene.
4th. The middle of the abdomen is generally blackish.
5th. The dimensions are larger.
The Ké-Islands specimens are a little larger than those from the
Aru Islands.
94 COUNT T. SALVADORI ON THE (Jan. 15,
9. Sauropatis cHLorts (Bodd.).
Halcyon sordida, var., G. R. Gr. P. Z.S. 1858, p. 172 (Aru).
[No. 244. Female, Dobbo: legs slate, with slightly bluish tinge ;
eyes black. Shot on Dobbo on the 20th September, 1874.—J. M.]
This bird has the upper parts very dark, and in that respect ap-
proaches S. sordida (Gould), to which Mr. Gray thought that the
Aru birds should be referred ; but I have seen other specimens from
the Aru Islands with the upper parts of a much brighter colour,
exactly like others from Amboyna, Bouru, and other localities.
The true S. sordida (Gould) is a much larger bird.
10. Savropatis sancta (Vig. & Horsf.).
Halcyon sanctus, Sharpe, Mon. Alced. pl. 91 (1868-1871).
Sauropatis sancta, Salvad. Ann. Mus. Civ. Gen. x. p. 306, sp. 33
(1877).
[No. 219. Male (young), Wokan: eyes black; upper mandible
and edge of lower black, under base of lower light-coloured; feet
brown.—J. M.]
11. SauRoMARPTIS GAUDICHAUDI (Q. & G.).
Dacelo gaudichaudii, Quoy & Gaim. Voy. Uran. Zool. p. 112,
pl. 25 (1824), Waigiou, Guebé (type examined); Sharpe, Mon.
Alced. pl. 116 (1868-1871).
Sauromarptis gaudichaudi, Salvad. Ann. Mus. Civ. Gen. x. p. 306.
sp. 35 (1877).
[203. Male; 204. Female, Wokan: eyes black; legs greenish,
yellow tinge ; stomach contained insects and portions of leaves.—
J. M.
The male has the tail blue; and the female has the tail chestnut.
12. Ssuromarptis Tyro (G. R. Gr.).
Dacelo tyro, G. R. Gr. P. Z. 8. 1858, p. 171, pl. 133, Aru (type
examined) ; Sharpe, Mon. Alced. pl. 117 (1868-1871).
Sauromarptis tyro, Salvad. Ann. Mus. Civ. Gen. x. p. 307. sp. 36
(1877).
[No. 205. Young male? Wokan: eyes black; legs greenish-
yellow tinge ; stomach contained insects and portions of leaves.—
J. M.)
This bird differs from the adult in having the underparts entirely
reddish, and in the blue of the wings inclining to greenish.
13. AAGOTHELES WALLACE, G. R. Gr.
Aigotheles wallacei, G. R. Gr. P. Z. S. 1859, p. 154, Dorey,
(type examined) ; Schleg. Ned. Tijdschr. Dierk. iii. p. 340 (1866),
Aru; Salvad. Ann. Mus. Civ. Gen. x. p. 310. sp. 7 (1877).
Caprimulgus brachyurus, Rosenb. Nat. Tijdschr. Ned. Ind. xxix.
p- 143 (1867), Aru (type examined).
[No. 259. Female, Wanumbai.—J. M.]
I have compared this specimen with the type in the British Mu-
1878. ] BIRDS OF TERNATE, AMBOYNA, ETC. 95
seum and with two other specimens from New Guinea, collected by
D’Albertis and Bruijn’s men. From all these it differs in the finer
vermiculations, and in the somewhat smaller dimensions, and perhaps
may belong to a different species; but we must see some more spe-
cimens from the Aru Islands before we can decide the question.
The Caprimulgus brachyurus, Rosenb., is a young bird with the
tail incompletely developed.
14. Macropreryx MYSTACEA (Less.).
[No. 254. Male, Wanumbai: eyes black.—J. M.]
This bird has a brown spot on the ear-coverts.
15. Hrrunpo savantca, Sparrm.
Hirundo frontalis, Quoy & Gaim. Voy. Astrol. Zool. i. p. 204,
pl. 12. f. 1 (1830), Dorey (type examined).
[No. 225. Female, Dobbo: eyes black.—J. M.)
16. HyLocHELipon NIGRICANS (Vieill.).
Hirundo nigricans, Vieill. N. D. xiv. p- 523 (1817), New Holland
(type examined).
[No. 226. Male, Dobbo: eyes black. Shot by the Professor at
the village of Dobbo.—J. M.]
17. Macha2RoRHYNCHUS XANTHOGENYs, G. R. Gr.
Macherirhynchus xanthogenys, G. R. Gr. P. Z. 8. 1858, p. 176,
Aru (type examined) ; Salvad. Atti R. Acc. Sc. Tor. x. p. 375. sp. 2
(1875).
[No. 234. Male, Wokan: eyes and bill black ; feet slate. Shot
at Wokan.—J. M.]
18. Gravcatus mexanors (Lath.).
Campephaga melanops, var., G. R. Gr. P.Z.S. 1861, p. 435.
[No. 228. Female, Dobbo: eyes brown; stomach contained in-
sects ; legs and bill black. Shot by the Professor at the village of
Dobbo.—J. M.]
The specimen agrees entirely with black-faced Australian males,
and with male birds collected’ by Signor D’Albertis at Yule Island
(Ann. Mus. Civ. Gen. vii. (1875), p. 820. sp. 41).
19. EpoLiosoma MUELLERI, Salvad.
Ceblepyris plumbea, part., 8. Miill. (nec Wagl.), Verh. Land- en
Volkenk. p. 189 (1839-1844), New Guinea only (type examined).
Edoliosoma miilleri, Salvad. Ann. Mus. Civ. Gen. vii. p. 928
(1875).
[No. 260, Wanumbai: male. No. 245. Female: eyes hazel ;
bill black ; legs dark with brownish tinge; stomach had insects.
Shot on 20th, Dobbo.—J. M.]
These specimens agree in every respect with New-Guinea specimens.
Edoliosoma tenuirostre (Jard. et Selby), from Australia, is scarcely
different from the New-Guinea bird ; perhaps it is a little larger.
96 COUNT T. SALVADORI ON THE [Jan. 15,
S. Miiller, under the name of Cedlepyris plumbea, has united a
specimen from Utanata and a young bird from Timor ; but this, ac-
cording to me, belongs to a much lighter species, not yet named, of
which I have seen adult specimens in the British Museum. With the
same name of C. plumbea there is in the Leyden Museum a specimen
marked ‘‘ Borneo :” it is a young bird exactly like that from Timor ;
and I think that the locality is wrong. On the evidence of this
specimen Hartlaub (Journ. f. Orn. 1865, p. 155) has asserted the
existence of C. plumbea, Miill., in Borneo.
20. Dicruropsis assimitis (G. R. Gr.).
Dicrurus assimilis, G. R. Gr. P. Z.S. 1858, p. 129, Aru (type
examined ).
Chibia assimilis, Sharpe, Cat. B. Brit. Mus. iii. p. 239 (1877).
[No. 214. Female, Wokan: eyes dull red; feet and bill black ;
stomach contained insects.—J. M.]
21. CoLLURICINCLA MEGARHYNCHA (Q. & G.).
Muscicapa megarhyncha, Quoy & Gaim. Voy. Astrol. Zool. i.
p- 172, pl. 3. f. 1 (1830), Dorey (type examined).
Napothera eleoides, Mill., Mus. Lugd. (type examined).
Myiolestes aruénsis, G. R. Gr. P. Z. S. 1858, p. 180, Aru (type
examined).
Pinarolestes megarhynchus, Sharpe, Cat. B. Brit. Mus. iii. p. 295
(1877).
[No. 240. Male: eyes hazel; legs slate. Shot at Wanumbai.—
J. M.|
I quite agree with Mr. Sharpe in uniting the Aru bird with the
New-Guinea one.
22. Recres arvénsis, Sharpe.
Rectes dichrous, G. R. Gr. (nec Bp.), P. Z. 8. 1858, p. 179, Aru.
Rectes aruénsis, Sharpe, Cat. B. Brit. Mus. iii. p. 285 (1877),
Aru (type examined)’.
[No. 238. Female, Wanumbai: eyes dull red; feet slate-colour
with a violet tinge; stomach contained land-shells and remains of
insects.—J. M.]
255. Female, Wanumbai: eyes hazel.—J. M.]
These specimens, both marked females, and a female collected by
Beccari in the Aru Islands, agree in having the head blackish, the
front neck blackish tinged with reddish brown, the uropygium and
the upper tail-coverts blackish brown, the tail blackish above, and
decidedly reddish brown underneath. These three specimens, all
1The type of Rectes dichrous ceramensis, Meyer (Sitzb. Ak. Wien, Ixix.
p. 208), which I haye lately seen in the Museum of Vienna, is a specimen of
R. uropygialis, Gray, and not of R. aruénsis,as Mr. Sharpe suspects. The
locality, Ceram, is, no doubt, wrong. Rectes draschi, Pelzelu, Verh. z.-b. Gesell.
Wien, 1876, p. 218, is a female of Hdoliosoma schisticeps, G. R. Gr. (ex Hombr.
& Jacq. Voy. Pole Sud, Atlas, Zool. pl. 10. f. 1, 2).
1878. ] BIRDS OF TERNATE, AMBOYNA, ETC. 97
marked females, differ from two other specimens collected by Beccari,
marked males. These have the head, the neck, the tail, and the
upper tail-coverts jet-black ; and the breast in one specimen is tinged
with black more conspicuously than in the others. Ought we to
believe that the plumage of the first three specimens, described by
Sharpe as that of the young bird, is also the plumage of the adult
female, or that by a strange combination all the three specimens
marked females are really young ones?
23. Cracricus cassicus (Bodd.).
[No. 218. Female, Wokan: eyes dark, deep blue; base of the
mandibles light-coloured, with a tinge of blue, tips of mandibles
black ; stomach with leaves and seeds.—J. M.]
24. CyrTosToMuS FRENATUS (S. Miill.).
|No. 222. Male, Wokan: eyes black ; bill and feet black—J. If.]
25. Myzometa nieriTA, G. R. Gr.
Myzomela nigrita, G. R. Gr. P. Z.S. 1858, p. 173, Aru (type
examined).
Myzomela erythrocephala, Meyer (nec Gould), Sitzb. k. Ak. Wiss.
zu Wien, Ixx. p. 204 (1874), 2 and young ¢ (specimens examined).
[No. 232. Wokan, male: eyes hazel ; bill and feet black ; stomach
had insects.—J. M. |
Gray describes the male as “ entirely shining deep black,’’ omit-
ting to mention that it has the under wing-coverts and the inner edges
of the remiges white.
26. XANTHOTIS FILIGERA (Gould).
Ptilotis filigera, Gould, P. Z.S. 1850, p. 278, pl. 34, Cape York.
[No. 261. Wanumbai, male.—J. M.]
This bird has the light spots on the nape scarcely visible. Two
specimens from Utanata, collected by S. Miiller, in the Museum of
Leyden, belong to this species and not to the northern form X.
chrysotis (Less.).
27. DRYMAG@DUS BECCARII, Salvad.
Drymacdus beccarit, Salyad. Ann. Mus. Civ. Gen. vii. p. 963. sp.
52 (1875), Arfak.
[No. 257. Wanumbai, male.—J. W.]
This example agrees completely with the type from the Arfak
Mountains. ‘This is a new species for the Aru Islands.
28. CALORNIs METALLICA (Temm.).
[Nos. 215, 216,217. Males, Wokan: eyes coral-red ; bill and legs
black ; small black seeds in the stomach of the adult males. The
above were the most abundant birds we saw on the 17th September
1874.—J. M.]
Proc. Zoou, Soc.—1878, No. VII. 7
98 COUNT T. SALVADORI ON THE [Jan. 15,
29. Mino pumontyI, Less.
[No. 250. Wanumbai, male: eyes yellow. No. 233. Wanumbai,
female: eyes yellow; legs and bill yellow ; stomach contained seeds.
30. ParapisEA ApopA, Linn.
[Nos. 248, 249. Males: eyes yellow. No. 247. Female: eyes
brown. No. 202. Young male: eyes pale yellow; legs light fawn-
colour; the stomach wasempty. This bird was shot by Mr. Abbot ;
he saw two others at the same time, but noticed none of them with
plumes. The Professor and Captain Nares saw a large flock ; and,
again, none of them were noticed with plumes. No other specimens
were seen by the party. No. 246. Female: eyes brown.—J. M.]
Four specimens from Wanumbai, and one from Wokan.
Ist. An adult “male” (No. 249), but not in full dress, wanting the
long yellow feathers on the sides.
2nd. A “ male”’ (No. 248) not fully adult, having the two middle
tail-feathers with very large webs, like the lateral ones, and longer
than these, one about two inches, the other only one inch; they
are of a lanceolate shape.
3rd. An adult “ female ”’ (No. 247) with yellowish nape.
4th. A young “male” from Wokan (No. 202).
5th. A young “female” (No. 246). The two last are exactly alike in
colour; but the male is larger than the female. Both have no yellow
on the nape.
31. CicinnuRUS REGIUS (Linn.).
Cicinnurus regius, Salvad. Ann. Mus. Civ. Gen. ix. p. 192. sp. 24
(1876); Sharpe, Cat. B. Brit. Mus. iii. p. 171 (1877).
[No. 201. Male: eyes dull red; legs a delicate sky-blue; the
stomach contained a few remains of insects. This bird was shot by
Mr. Abbot ; two others were in company with it at the time. These
were the only specimens seen, either by Mr. Abbot or any other of
the party. They were not apparently very shy. A gun is said to
have been fired, close by, just before they were observed. No. 241.
Male, shot at Wokan: eyes dull red; legs delicate sky-blue or cobalt
blue. Nos. 242, 243. Males, shot at Wanumbai. Nos. 250, 251.
Males: eyesyellow. No. 252. Male (young): eyes yellow.—/. M.]
Seven specimens—one fully adult “ male” (No. 201) from Wokan,
and five adult “‘males”’ (Nos. 241, 242, 243, 250, 251), and a young
“male” (No. 252) from Wanumbai.
32. Manucopia atTRa (Less.).
Manucodia atra, Salvad. Ann. Mus. Civ. Gen. ix. p. 189. sp. 4
(1876); Sharpe, Cat. B. Brit, Mus. iii. p. 183 (1877).
[Nos. 209, 210. Male, female: eyes orange; legs black ; the
stomach contained fruit and seeds.—J. M.]
The male is larger than the female.
1878.] BIRDS OF TERNATE, AMBOYNA, ETC. 99
33. AALURGDUS MELANOTIS (G. R. Gr.).
Ptilonorhynchus melanotis, G. R. Gr. P. Z. S. 1858, p. 181, Aru,
(type examined).
Ailureedus melanotis, Gould, B. of New Guinea, pt. i. (1875) ;
Salvad. Ann. Mus. Civ. Gen. ix. p. 193, sp. 28 (1877).
[No. 211. Eyes red; legs with a bluish tinge. Appears to be a
young bird; could not make out the sex.—J. W.]
Mr. Gould has expressed some doubts about 4. arfakianus,
Meyer, being really different from this species; but the greater
amount of black on the head and on the neck of 4. arfakianus, the
triangular shape of the light spots, with the acute angle turning
downwards, on the upper part of the breast, and the smaller dimen-
sions, distinguish at once the New-Guinea bird from its Aruan
relative.
34. Pritorus rozonus, G. R. Gr.
Ptilonopus (!) iozonus, G. R. Gr. P. Z.S. 1858, p. 186, Aru (type
examined),
Ptilopus ionozonus, Salvad. Ann. Mus. Civ. Gen. ix. p. 198. sp. 24
(1876).
[No. 289. Wanumbai, female: legs red; base of upper mandible
red, tip yellow ; lower green, with the tip lighter ; stomach empty.
—J. M.)
35. Pritorus coronuLatus, G. R. Gr.
Ptilonopus coronulatus, G. R. Gr. P. Z. 8. 1858, p. 185, pl. 138,
Aru (type examined).
Ptilopus coronulatus, Salvad. Ann. Mus. Civ. Gen. ix. p. 198. sp.
26 (1876).
[Nos. 220, 221. Wokan, males: eyes red; bill green; feet red
(dull) ; stomach empty.—J. M.]
36. CARPOPHAGA ZOEH (Less.).
Columba zoée, Less. Voy. Coq. Zool. Atlas, pl. 39 (1826), i. pt. 2,
p- 705 (1828), Dorey (type examined).
Carpophaga zoée, Salvad. Ann. Mus..Civ. Gen. ix. p. 201. sp. 47
(1876).
[No. 236. Wanumbai, male: eyes white; legs red; beak black ;
stomach contained seeds. Shot at Wanumbai.—J/. WV. ]
37. Macropyer1a poreEyA, Bp.
Macropygia doreya, Bp. Consp. ii. p. 57 (1854), New Guinea
(type examined); Salvad. Ann. Mus. Civ. Gen. ix. p. 204. sp. 65
(1876).
Macropygia phasianella, part., G. R. Gr. (nec Temm.), P. ZS.
1858, p. 187, Aru Islands only; Wall. Ibis, 1865, p. 389. sp. 81
(partim). ”
[No. 254. Wokan, female.—J. J/.]
100 MR. A. Hs GARROD ON THE MOMOTIDZ. (Jan. 15,
38. Mecapropius pUPERREYI, Less. & Garn.
Megapodius duperreyi, Less. & Garn. Bull. Se. Nat. viii. p. 113
( 1826), Dorey (type examined); Less. Voy. Coq. Zool. i. pt. 2,
p- 700, 703, pl. 36 (1826-1828).
Megapode reinwardt, Temm. Pl. Col. livr. 37, genre Megapode
(1823).
Megapodius rubripes, Temm. Pl. Col. 411 (25th October 1826'),
Celebes! Amboyna!
Megapodius reinwardtii (sic), Temm. apud Wag}. Syst. Av. Gen.
Megapodius, note (1827), descr. nulla.
Megapodius reinwardtii, Wagl. Syst. Av. Gen. Megapodius, Ad-
ditamenta, sp. 4 (1827).
Megapodius rufipes, 8. Miill., Verh. Land- en Volkenk. pp. 23,
109 (1839-1844), Utanata.
Megapedius, sp., Wall. Ann. & Mag. Nat. Hist. (2) xx. pp. 473,
477 (1857), Ké, Aru.
[No. 230. Female: eyes dull red; legs orange; bill horn-colour.
—J. M.)
A female (No. 230) from Wokan.
39. NUMENIUS UROPYGIALIS, Gould.
[No. 258. Wanumbai, male.—J. M.]
40. TRINGOIDES HYPOLEUCUs (Linn.).
[No. 227. Dobbo, female: legs light green; billdark green; eyes
hazel; stomach contained insects.—J. M.]
7. On the Systematic Position of the Momotide.
By A. H. Garrop, M.A., F.R.S., Prosector to the Society.
[Received December 3, 1877.]
In my paper on certain muscles of birds and their value in classifi-
cation *, I have made an error, which I desire to correct, with refer-
ence to the systematic position of the Momotide. I there included
them with the Coraciide as part of a single family, characterized
among the Anomalogonatee by the possession of a nude oil-gland,
together with colic czeca. Since the time my paper appeared, I have
had the opportunity of dissecting several species of Momotide, thanks
to the kindness of Mr. O. Salvin, including Momotus lessoni, M.
equatorialis, M. brasiliensis {a specimen which had lived in the
Society’s Gardens), Hylomanes gularis, and Eumomota superciliaris ;
and I find that in none of these are colic czeca present.
Further, in Hylomanes gularis and Eumomota superciliaris I find
' I doubt whether this date, given by Crotch (Ibis, 1868, p. 500), for the pub-
lication of the 69th livraison of Temminck’s ‘ Planches Coloriées,’ is exact ; my
doubts rise from the fact that Mf. rubripes is not mentioned by Wagler in his
‘Systema Avium,’ published in 1827.
2 P.Z.8. 1874, p. 128.
ia>
1878. ] MR. A. H. GARROD ON THE MOMOTIDS. 101
a minute tuft on the apex of the oil-gland, although in the several
species of Momotus there is no trace of any tuft; in fact they have
lost it, evidently since the family was differentiated off.
Such being the case, the Momotidz must be placed with the Pici-
formes, as defined by me, instead of with the Passeriformes ; and the
amended arrangement may be thus tabulated, the Todide and Mo-
motide almost certainly forming a single family, as has been sug-
gested by many, and which is confirmed by the observation made by
Dr. Murie that in the Todidze the beak is serrate’.
ANOMALOGONATE.
Aves Piciformes. Aves Passeriformes.
Bucerotide. Coraciide.
Alcedinide. Steatornithide.
Momotide. Caprimulgide.
Todide. Galbulide.
{ Ramphastide. Meropide.
Capitonidee. Trogonide.
Pici. Bucconidez (?).
Passeres.
In further favour of the inclusion of the Momotide with the Pici-
formes may be mentioned the pterylographic peculiarity found in
them all, namely that the outer pectoral branch of the inferior tract
is separated almost entirely from the inner branch, with which it
blends for nearly its whole distance, or entirely, in the Passeriformes.
From a skin, I have been able to determine that the deep flexor
tendons of the leg of Todus viridis are arranged on exactly the same
plan in it as in the Momotide’, and that its tensor patagii brevis
also terminates in exactly the same manner as it does in them?.
The syrinx of the Momotidz has never been fully described, so far
as I am aware. I therefore exhibit a figure of it as it appears in
Syrinx of Momotus lessoni.
Momotus lessoni, which resembles that of the other species which I
have examined. Fig. A is the anterior view; B, the posterior.
1 P.Z.8. 1872, p. 671.
2 Vide P. Z. 8, 1875, p. 344, 3 Vide P.Z. 8. 1876, p. 511.
102 MR. A. H. GARROD ON CARPOPHAGA LATRANS. [Jan. 15,
The large cartilaginous three-way piece, in which the trachea
terminates inferiorly, is compound, being formed of several fixed
rings. It is complete in front, being represented behind by a
hooked process on either side, extending inwards towards the middle
line, where the two nearly meet. The lateral muscles of the trachea
extend down to the upper margin of this peculiar syrinx ; and a few
of their anterior fibres continue onwards to the surface of the carti-
laginous box, where they terminate, sometimes higher and sometimes
lower, but always before reaching its inferior margin.
8. Note on the Gizzard and other Organs of Carpophaga
latrans. By A. H. Garrop, M.A., F.R.S., Prosector to
the Society.
[Received December 3, 1877.]
In the collection of birds preserved in spirit by H.M.S. ‘ Chal-
lenger’ is the body, after the skin had been removed, of a single
specimen of Carpophaga latrans, together with the gizzard of a second
individual of the same species, obtained at Kandavu, Fiji. These
form the material for the present communication.
In his note-book Mr. John Murrray makes the following remarks
on the species':—‘*Stomach contained the fruit of some tree un-
known to me. The coat of the stomach had hard _papilla-like
ossifications of a circular form, two or three rows. . . . These indura-
tions are composed of a horny substance”—from which it is seen
that Mr. Murray was the first to recognize the existence of the
strange arrangement to be here described.
The thin-walled and capacious crop contained only one thing in
its interior—a complete fruit, which has been identified for me
by Mr. W. T. Thiselton Dyer, as that of Oncocarpus vitiensis.
In the gizzard was also found a portion of a second example of the
same fruit.
Oncocarpus vitiensis is a tree belonging to the natural order Ana-
cardiaceze, which, according to Dr. Seemann’, is ‘ about sixty feet
high, bearing large oblong leaves and a very curious corky fruit, some-
what resembling the seed of a walnut.” The tree is included
among those which are poisonous by the Fijians ; and its sap produces
an intense itching of the skin, when brought into contact with it,
whence the native name Kau Karo or itch-wood.
For the crushing of this very hard fruit a special anatomical
modification of the gizzard-walls of this Fruit-pigeon is developed,
which is peculiarly interesting and tends to prove the plasticity of
organs when aberrant forces come into play.
The gizzard is not developed to any thing like the extent that it is
‘1 Vide P. Z.8. 1877, p. 737.
2 Seeman’s Mission to Viti, p. 334,
1878.] MR. A. H. GARROD ON CARPOPHAGA LATRANS. 103
in the Gallinze or Anseres, but, as in most Carpophagine birds, is
small, and has its muscular- walls comparativaly ill-developed. No
stones of any kind were found in it.
It is the epithelial lining which is so peculiar in Carpophaga
latrans. Instead of being smooth, or folded into plications, as is
usually the case, its surface is raised into horny cones which closely
resemble in appearance the tubercles for the attachment of the spines
of the Echinoderm genus Cidaris. In fig. 1, a, the interior of the
gizzard is represented; fig. 1, 6, is a section of one of the cones
resting upon the muscular gizzard-wall.
These conical processes are corneous throughout, are erect, and
are quite transparent when cut into sections. There are twenty-
Fig. 1.
Interior of the gizzard of Carpophaga latrans.
three of them, large and small, in each of my specimens. The larger
cones, which are the more numerous, average seven millimetres in
diameter at their bases, their axial length being about four milli-
metres; the smallest cone is four millimetres across and of nearly
the same height.
The cones are arranged, close to one another, in a fairly regular
manner upon the two muscle-masses, being distributed in rows of
three, counting either transversely or longitudinally. This disposes of
eighteen of the twenty-three cones. The remaining five are found
on the tendinous intermuscular walls of the organ, in longitudinal
rows, two in one row, three in the other.
A section of any one of the cones demonstrates that it is not
in the least ossified, but corneous throughout, and of about the
density of ox-horn. It is also seen that the attached surface of
the epithelium does not participate in the undulations of the free
surface, being quite smooth. Neither does it send any processes into
the cones. Between the cones the epithelium is yielding, and only
semicorneous.
A still further exaggeration of this abnormal condition of the
epithelium of the gizzard of Carpophaga latrans has been described
104 MR. A. H. GARROD ON CARPOPHAGA LATRANS. (Jan. 15,
by MM. Jules Verreaux and O. Des Murs in Phenorhina goliath’
ot New Caledonia, which ‘se nourrit de graines de semicarpum.”
In this bird “le gésier, déja on ne peut plus musculeux par lui-méme,
a sa surface intérieure réguliérement recouverte . . . de pointes véri-
tablement osseuses, rappelant la forme de celles quise voient a la sur-
face ducorps de la Raia bouclée, ou Clavel, ou Clavelade. Ces
pointes, en céne aplati, ont leur base plane de 5 millim. de diamétre,
d’une hauteur de 5 a 6 mill., sont légérement inclinées sur elles-
mémes, et quelquefois recourbées par la dessication, lextrémité en
étant mousse.” A central fibrous peduncle is also said to run
through each osseous element.
Phenorhina goliath, from what has been said above, therefore
differs from Carpophaga latrans in having the cones of the gizzard
proportionally longer, at the same time that they are ossified
(which necessitates the presence of vessels in the ossification, which
appear after death as the fibrous cord) and oblique. There is,
however, a great similarity between the two organs.
I am informed by the Rev. S. J. Whitmee that Carpophaga pacifica
in the Samoan Islands feeds on nutmegs, from which it is highly
probable that in that species the gizzard-epithelium is modified in a
manner similar to that of the Fijior New Caledonian species. Speci-
mens of C. pacifica preserved would therefore be of special interest
for the determination of this point.
Syrinx of Carpophaga latrans.
With reference to the other parts of Carpophaga latrans, the
intestine is very capacious, only nine inches long, and transversely
sacculated from the contraction of its outer longitudinal muscular
coat, this producing the appearance of thirty bold transverse folds on
the mucous surface. There are no colic ceca; and, as in the genus
Carpophaga generally *, the gall-bladder is well developed. The
liver-lobes are equal in size.
* Revue et Magasin de Zoologie, 1862, p. 168, * Vide P.Z.S, 1874, p. 258.
1878.] MR. E. A. SMITH ON A HELIX FROM JAPAN. 105
The syrinx (fig. 2, p. 104) is Columbine, with its lateral muscles
attached inferiorly to the membrane between the penultimate and
antepenultimate tracheal rings. The trachea is composed of rings
which are very yielding in the middle line posteriorly. As can be
seen in the figure, the musculi sterno-tracheales, which are inde-
pendent of the intrinsic muscles, are not quite symmetrically attached.
The furcula, as is the rule in the subfamily, is very slender, but
complete, wherein this species differs from Phenorhina goliath, in
which it is cartilaginous at its symphysial end, according to MM.
Verreaux and Des Murs’.
Myologically, the ambiens is to be found, not large ; the femoro-
caudal with its accessory head are well developed ; the semitendinosus
and its accessorius are the same.
There are two carotids; so that in this as well as all the other
features above mentioned Carpophaga latrans agrees with my
definition of the division of the Columbide into which it naturally
falls, its gizzard differing, however, from that of all but one of the
species which have been examined. iv
9. Description of a new Species of Helix from J apan.
By Enear A. Suirnu, F.Z.S.
[Received December 4, 1877.]
Hewrx (CaMENA) CONGENER, n. sp.
Testa convewiusculo-orbiculata, spira breviter conica, subaperte
umbilicata, tenuis, sordide pallido-virescenti-albida, fasciis
tribus nigrofuscis, media latissima, infima umbilicum pingente ;
anfractus 54, conveviusculi, lineis incrementi obliquis flexuosis,
striisque spiralibus confertis insculpti, ultimus subtus satis con-
vecus, prope aperturam leviter breviterque descendens; apertura
perobliqua, semilunaris, intus fasciata ; peristoma tenue, rosa-
ceum vel liliaceum, marginibus conniventibus, supero breviter
expanso, basali et columellari latius reflexis.
Diam. max. 30 mill., min. 25, alt. 18 ; aperture long. 15, lat. 14.
As this form approaches very closely to H. peliomphala of
Pfeiffer, I will give a comparative description as the best means of
showing its distinctness.
It is uniformly smaller, more narrowly umbilicated and of a paler
ground-colour ; its spire is constantly more elevated, aperture smaller;
the banding in the fourteen specimens examined invariably follows
the same arrangement, and not one of them exhibits any trace of the
opaque yellow stripes or blotches which are almost invariably ob-
servable in a greater or less degree (on the bands most conspicuously)
in H. peliomphala.
-This is a very pretty, delicate species, and of a thin semitransparent
texture. On the front of body-whorl, about two thirds of its whole
extent from the lip, there is an obliqne strongly marked line which is
? Loc. cit, p. 140.
106 THE MARQUIS OF TWEEDDALE ON (Jan. 15,
suffused with a pinkish tint, evidently marking a period of cessation
of growth, the colour having stained that part of the shell which then
formed the lip of the aperture.
The uppermost dark band of the last whorl is situated a little above
the middle, and passes just above the suture, and can be traced in part
almost to the apical whorl. Adjoining this thin line there is a second
thread-like one which borders the sutural edge of the whorls. The
median band is very broad, frequently more than twice as wide as
that above it, and encircles the whorl somewhat below the periphery ;
and the basal one, equally broad with the preceding, winds around
the rounded margin of the umbilicus.
Helix (Camena) congener.
Other allied forms of this species are H. luhuana, Sowerby, which
seems to be distinct from H. peliomphala, H. simode, Jay, and H.
herklotsit, Martens.
Whether these five are good and distinct species or only local forms
of one and the same, of which intermediate gradations have not yet
been observed, remains yet to be ascertained ; but, so far as our
present knowledge extends, the differences of form, texture, banding,
&c., warrant the presumption that they are specifically distinct.
The series of specimens upon which the above description is based
has been kindly placed in my hands for examination by Mr. G. B.
Sowerby, junior, who has presented two of them to the national
collection.
10. Contributions to the Ornithology of the Philippmes.—
No. IV. On the Collection made by Mr. A. H. Everett
in the Islands of Dinagat, Bazol, Nipah, and Sakuyok.
By Arruur, Marquis of TweErpparz, F.R.S., President
of the Society.
[Received December 11, 1877.]
(Plates VI.—VIII.)
The four islands in which Mr. Everett collected the birds of
which I propose to give an account are situated to the north of the
[Smit lith
TIVDOATUIVMWIC OMT PaTIa
it ti Es I ce Ue LS he a (ELE Dp Lilo
2 MIXORNIS CAPITALIS
J.Smit lith
Neo
G3
DICAAUM &
DICAUM EVERE
PRIONOCHILUS
Hanhart imp
_ 1878.) THE ORNITHOLOGY OF THE PHILIPPINES. 107
shores of Mindanao, and are only separated from that island by
narrow channels. Through these channels the wind blows with
considerable force throughout the greater part of the year, being
either the gales of the S.W. or the N.E. monsoon.
Mr. Everett’s observations on the ornithological features of these
islands, coming as they do from a personal observer, merit transcrip-
tion. He writes :—“ No district visited so far has equalled the vici-
nity of Manilla in the number and variety of its birds. The progress
of the work shows that a very close general similarity exists between
the islands of Mindanao and Luzon, though there are some marked
local differences. This was to be expected; and it is rather sur-
prising that the differences are as marked as they are, when we con-
sider that not only are the islands separated from another by very nar-
row spaces of sea, but that they are situated in an area where strong
winds blow in various directions, with little cessation during the major
part of the year, accompanied with violent squalls and typhoons, so
that any high degree of specialization in the local avifauna would
seem, @ priort, out of the question.”
In the island of Dinagat, Mr. Everett writes, he found birds to be
excessively rare ; still he obtained 39 species, 7 of which were unde-
scribed :—
Ceyx argentata. Diceum everetti.
Hypothymis celestis. Prionochilus olivaceus.
Mizxornis capitalis. Aithopyga dubia.
Diceaum schistaceum.
Only 13 species were obtained in the small islands of Nipah,
Bazol, and Sakuyok; and they are all known inhabitants of Min-
danao.
1, Cacarua H&MATUROPYGIA (1).
[Nipah, 2, July. ]
2. LoricuLus HARTLAUBI (7).
[a. Dinagat, ¢,2, June. 6. Bazol, o, July.]
The example figured, P. Z.S. 1877, p. 819 (¢), is in full adult
plumage, the occipital feathers being orange-red, grading into orange
on the nape, which colour suffuses and descends the back, losing
itself in the pure green of the lower back. There is no blue about
the face, chin, or throat ; the dark blue of the under surface of the
rectrices fades into pale blue at their apices. The orange hue on
the back of the female figured is less intense.
3. HaRPacTES ARDENS (34).
[Dinagat, ¢, June. Iris brown; bill yellow, base green. j
A single example, in adult plumage, and which has a more massive
bill than is found in Luzon individuals. Otherwise alike, and di-
mensions equal.
4. MEROPS PHILIPPINUS (35).
[Bazol, ¢, July.]
108 THE MARQUIS OF TWEEDDALE ON [Jan. 15,
5. EurRYSTOMUS ORIENTALIS (37).
[a. Dinagat, g, June. 6. Nipah, 2, July.]
6. PELARGOPSIS GIGANTEA,
Pelargopsis gigantea, Walden, Ann. & Mag. Nat. Hist. (4) xiii.
p. 123, P. Z.S. 1877, pp. 541, 822.
[Dinagat,?, ¢, June. ]
Neither example in full plumage ; and both are not separable from
the Butuan male observed upon/. c. Butuan, Pasananca, Malanipa,
and these Dinagat individuals appear to belong to a small race of
P. gigantea.
7. CEYX ARGENTATA. (Plate VI.)
Ceyx argentata, Tweeddale, Ann. & Mag. Nat. Hist. (4) xx.
p- 533, 2 (1 December 1877).
[Dinagat, 2, June. Iris brown; bill black ; feet pale reddish.]
Q. Chin, throat, upper breast, abdomen, ventral region, major
under wing-coverts, and thigh-coverts pure white. Cheeks, sides
of head, and ear-coverts black. Lower breast and flanks black, washed
with deep blue. Rectrices and under tail-coverts black. Plu-
mage above black; a bold supercilium commencing above the eye
and running into the occipital crest, formed by the component
plumes being tipped with pale silvery blue; many of the feathers of
the vertex minutely tipped with silvery blue. Occipital crest tipped
with pale silvery bluish green, or silvery blue, or the two together.
Dorsal plumage and lengthened upper tail-coverts black at base, then
silvery bluish green, then pale silvery blue. Wings black. Major
coverts each with a terminal silvery-green spot. Loral spot white ;
an isolated tuft on the sides of neck, white tipped with creamy fulvous.
Bill black.
Wing 2°62 inches, tail 1°37, tarsus 0°45, culmen 1°50.
8. SAUROPATIS CHLORIS (47).
[Dinagat, 9, June.]
9. PyRRHOCENTOR MELANOPS (65).
[Nipah, 2, July.]
10. PENELOPIDES AFFINIS.
Penelopides affinis, Tweeddale, Ann. & Mag. Nat. Hist. (4) xx.
p-. 534; P.Z.8. 1877, p. 824.
[Dinagat, ¢, June: iriscrimson. a. Q, June: iris rather dark
brown-orange. |
11. GRAUCALUS sTRIATUS (74).
[Nipah, ¢, 9, July: iris Naples-yellow. |
12. Latace pomrnica (76).
[Dinagat, ¢, 2, June. ]
1875. ] THE ORNITHOLOGY OF THE PHILIPPINES. 109
13. HyLorerre PHILIPPINENSIS (78).
[Dinagat, g, June; 4. Q: iris dark brown; bill black; legs
bluish grey. ]
Sexes alike—not to be distinguished from the type.
14. DicrurRvus STRIATUS.
Dicrurus striatus, Tweeddale, P. Z.S. 1877, p. 545, no. 20.
[Nipah, ¢, July.]
15. LevcocEeRCA NIGRITORQUIS (83).
[Dinagat, 2, June.]
16. CyorNIS PHILIPPINENSIS.
Cyornis philippinensis, Sharpe, Tr. Linn. Soc. ser. 2, Zool. 1.
p- 325, “ Panay ” (1876).
{Dinagat, ¢, June. ]
A single example, hardly separable from Zebu individuals, and
agreeing with them in dimensions, consequently larger than Luzon
and North-Mindanao specimens. Wing 3°66 inches, as against
2°95 ; tail, 3-00, as against 2°75.
17. Hypotaymis cexestis. (Plate VII. fig. 1.)
Hypothymis celestis, Tweeddale, Ann. & Mag. Nat. Hist. (4)
xx. p. 536, 2 (1 Dec. 1877).
{Dinagat, 2, June: iris dark-brown; bill blue tipped and
margined black ; interior of gape yellow-green ; orbital ring light
green; legs grey blue. |
9. Azure. Feathers of the head grey at their insertion, and
each tipped with glistening azure; a lengthened occipital crest
similarly marked. Space before the eye and lores pale blue. Chin,
cheeks, throat, sides of neck, and upper breast rich velvety turquoise-
blue, darker than the blues of the rest of the plumage. Back and
upper tail-coverts with grey bases to the feathers, the extremities
being blue, which is paler on the upper tail-coverts. Wing-coverts
of the same shade of blue as the head. Quills dark brown externally,
margined with a paler shade of blue. Lower breast, abdomen, vent,
flanks, and under tail-coverts white, tinged with pale greenish blue,
most intense on the flanks. Axillaries grey, tinged with pale blue.
Rectrices above like the quills; underneath dark brown like under
surface of the quills. Shafts of the rectrices brown above, almost
white underneath. Shoulder-edge blue.
Wing 2°82 inches, tail 3-00, tarsus 0°62, culmen 0°56.
A form nearly allied to H. azurea, but differing in its lengthened
crest, its paler tints of blue, and in the absence of any black mark-
ings. The blue shade of the throat and upper breast is about equal
in tint to that of the predominant colour of the older known species.
18. HrruNpDO JAVANICA.
Hirundo javanica, Sparrm., Mus. Carls. fase. iv. t. 100 (1789).
[Dinagat, ¢, June.]
110 THE MARQUIS OF TWEEDDALE ON (Jan. 15,
19. SARCOPHANOPS STEERII.
Eurylemus steerii, Sharpe, ‘ Nature,’ vol. xiv. p. 297. “ Basilan.”
Sarcophanops steerit, Sharpe, Tr. L.S. ser. 2, Zool. i. p. 344,
t. liv.
[Dinagat, 2, June: iris brilliant sap-green ; bill, orbital skin,
and legs fine blue, paler on the tarsal scales ; nails light grey.]
20. BropERIPUS ACRORHYNCHUS (90).
[Dinagat, 5, June.]
21. Mrxornis capitauis. (Plate VII. fig. 2.)
Mizxornis capitalis, Tweeddale, Ann. & Mag. Nat. Hist. ser. 4,
xx. p. 535 (Dee. 1, 1877).
[Dinagat, 2, June: iris orange; bill blackish grey; legs greenish
lead-grey ; soles and nails ochreous yellow. ]
Q. Forehead, vertex, and occiput rufous, each feather with a
pale fulvous central streak. Space before the eyes and lores dull
grey. Cheeks, ear-coverts, and sides of neck ashy, each feather
with a pale albescent central streak. Nape, back, and wing-coverts
the same, the pale striations on the back being very prominent in
consequence of the abnormal length of the dorsal plumage. Uropy-
gium and upper tail-coverts olive rusty grey and unstriated. Rec-
trices brown ; outer pair with the whole outer and apical third of
the inner web pale yellowish white; remaining pairs, middle
excepted, broadly tipped on both webs with white, middle pairs
slightly tipped with white. All the quills brown, narrowly margined
externally with fulvous grey, the tertiaries with pale rusty. Basal
portion of quills margined on their inner webs with white. Chin
and throat rufous and rufous-white. Breast, under wing-coverts,
abdomen, vent, and under tail-coverts white, faintly tinged with very
pale yellow.
Wing 2°75 inches, tail 2°50, tarsus 0°70, culmen 0°62.
22. MACRONUS STRIATICEPS.
Macronus striaticeps, Sharpe, Tr. L. 8. ser. 2, Zool. i. p. 331,
‘* Basilan”’ (1876).
[Dinagat, 9 : iris white. ]
The type specimen is not quite so rufous below, otherwise
identical.
23. Ixus GOIAVIER (99).
[Nipah, 2, pairing, July. |
24. PotioLopHus uRostictus (101).
[Dinagat, ¢, June: orbital ring lemon-yellow. 6, 2, June.]
Sexes alike.
25. CRINIGER EVERETTI.
Criniger everetti, Tweeddale, Ann. & Mag. Nat. Hist. ser. 4, vol.
xx. p. 535, et P. Z.S. 1877, p. 827, pl. Ixxxiv.
[Dinagat, ¢, 9, June.]
1878. } THE ORNITHOLOGY OF THE PHILIPPINES. 111
26. ORTHOTOMUS FRONTALIS.
Orthotomus frontalis, Sharpe, Ibis, 1877, p. 112, t. ii. f. i.;
Tr. L. S. (2) Zool. i. p. p. 336.
[Dinagat, ¢, 2, June.]
27. ZOSTEROPS EVERETTI.
Zosterops everetti, Tweeddale, P. Z.S. 1877, pp. 762, 829.
[Dinagat, ¢, June: iris yellowish brown; bill blackish; legs
pale grey. 0b, 9, June.]
Dinagat, North-Mindanao, and Zebu birds do not differ.
28. Dic#uUM RUBRIVENTER (120 partim).
Diceum rubriventer, Lesson, Tr. p. 303; Tweeddale, P.Z.S.
1877, p. 763.
[Dinagat, 3, June. }
The Dinagat birds in no respect differ from Luzon, Zebu and
North-Mindanao examples.
29. Diczum scuisTaceuM. (Plate VIII. fig. 1.)
Diceum schistaceum, Tweeddale, Ann. & Mag. Nat. Hist. ser. 4,
vol. xx. p. 537 (December 1, 1877).
[Dinagat, ¢, June. ]
3. Above fuliginous ashy; below ashy white. Remiges and
rectrices above dark ashy brown, underneath paler. Under wing-
coverts and axillaries pure white. Base of mandible (in dry speci-
mens) yellowish white. Apex and the maxilla pale ruddy brown.
Wing 2°25 inches, tail 1°37, tarsus 0°62, culmen 0°35.
A large species with a short stout bill.
30. Dic#um EvERETTI. (Plate VIII. fig. 2.)
Diceum everetti, Tweeddale, Ann. & Mag. Nat. Hist. ser. 4, vol.
xx. p. 537 (December 1, 1877).
[Dinagat, ¢, June: iris greyish brown; bill brown, base orange;
legs light grey. |
dg. Above dark olive-green, paler on the cheeks. Chin, throat,
and breast pale grey. Remainder of underparts albescent grey
tinged with palish yellow. Major coverts narrowly margined with
greenish yellow. Quills above dark brown, almost black, with very
narrow pale olive-green external margins. Rectrices above like the
quills, narrowly tipped with albescent fringe. Under wing-coverts
and axillaries pure white. Bill (dry specimen) brown at tip, yellow
at base.
Wing 1-94 inch, tail 1:00, tarsus 0°50, culmen 0°37.
31. PrronocuiLus otivaceus. (Plate VIII. fig. 3.)
Prionochilus olivaceus, 'Tweeddale, Ann. & Mag. Nat. Hist. ser. 4,
vol. xx. p. 536 (December 1, 1877).
[Dinagat, 2, June: iris sienna-brown ; bill grey-black, base of
mandible paler ; legs light grey. |
112 " THE MARQUIS OF TWEEDDALE ON (Jan. 15,
Q@. Entire upper surface, wing-coverts, and surface of closed
wing uniform pure olive-green, which colour encircles the eye and
covers the sides of the neck. Space before the eye, cheeks, sides of
throat, and the upper breast grey. Lower breast, abdomen, flanks,
and under tail-coverts the same grey washed faintly with olive-green,
the mesial line being albescent. Chin and throat white, slightly
greyish, forming a broad well-defined greyish white stripe, com-
mencing at the chin and descending to the breast. Axillaries and
under wing-coverts pure silky white. Remiges and rectrices dark
brown, margined with the olive-green hue of upper plumage.
Wing 2°12 inches, tail 1:25, tarsus 0°62, culmen 0°37.
Since I described this species (/. c.) I have received the male, which
in no way differs.
32. NECTAROPHILA SPERATA (122).
[Dinagat, ¢, June; 6, 9: iris brown; bill very dark brown;
legs black. c, June: iris dark brown; bill black, mandible brown ;
legs black. d, ¢ (juv.), June: iris dark brown. e, Nipah, ¢,
July. ]
I cannot separate example d ( ¢juv.) from examples 4 and ¢ (2).
A slight supercilium, chin, throat, cheeks, and whole lower surface
pale uniform greenish-yellow, above olive-green. They differ from
a supposed female, ex Surigao, described (P. Z.S. 1877, p. 830) by
having the chin, throat, and cheeks uniformly coloured with the
breast and abdomen, but agree in the ruddy colouring of the wing
when closed. A fourth example (sex not stated on label) is undis-
tinguishable from the Surigao female previously described (J. ¢.).
33. ARACHNECHTHRA JUGULARIS (123).
[Dinagat, 2, June. ]
+ 34. ALTHOPYGA DUBIA, N. sp.
[Dinagat, ¢, June: iris yellow-brown; bill brown; legs dark
grey ; feet ochreous. 6, ¢, June: bill and legs very dark brown.
c, 2, June: iris bright Indian-red ; bill black; legs light brown. |
3 (juv.?). Above, wing-coverts, and exposed surface of quills
olive-green. Uropygium bright sulphur-yellow, forming a broad
yellow band. Rectrices black, edged with olive; laterals tipped
with pale yellow-olive. Feathers of the lower surface ashy at their
base, and all tipped pale yellow; lores the same; cheeks rather
albescent-ashy. Under wing-coverts white; carpal edge very pale
yellow. Quills dark brown.
@. Like the male above described.
Wing. Tail. Tarsus. Culmen.
6 1°82 1-12 0°62 0:75
2 1°80 1-00 0°62 0°75
The three examples on which this species is founded are difficult
to distinguish, by their plumage alone, from 4. bella 2 vel 3
1878. | THE ORNITHOLOGY OF THE PHILIPPINES. 113
juv. ; but the dimensions are considerably greater :—culmen 0-7 5, as
against 0°50 ; wing 1-80, as against 1-62. Doubtless the adult male
will closely resemble that of 2. bella.
35. ANTHOTHREPTUS GRISEIGULARIS.
Anthothreptus griseigularis, Tweeddale, P. Z. S. 1877, p. 830.
[Sakuyok, ¢, July.]
36. RHABDORNIS MYSTACALIS (124).
[a. Dinagat, ¢, June: iris crimson ; bill black. 6. ¢: legs
yellowish brown. c. ¢: legs dark grey. |
The only female in a series of seven examples wears the same
dress as the males. Luzon individuals do not differ.
37. Corvus PHILIPPINUS (125).
[Dinagat, 3, 9.]
Female smaller than male.
38. CALORNIS PANAYENSIs (128).
[a. Nipah, 3,9, July. 5. Sakuyok, 2, July. |
39. Sarcops caLvus (129).
[Dinagat, ¢, June.]
40. OsMOTRERON AXILLARIs (136).
| Dinagat.)
A single example (¢), which cannot be separated from typical
individuals.
41. PHABOTRERON AMETHYsTINA (139).
[Dinagat, 2, June: iris yellowish brown. ]
This, Luzon, and N. Mindanao (Butuan) examples do not differ.
42. PHABOTRERON BREVIROSTRIS.
Phabotreron brevirostris, Tweeddale, P. Z.S. 1877, p. 549,
[Dinagat, 2, May.]
Undistinguishable from N. and S.W. Mindanao examples.
43. CARPOPHAGA £=NEA (141).
[ Dinagat. ]
44. Myristricivora BICcoLor (143).
[a. Nipah, 9, July: iris dark brown; bill blackish blue ; feet
light lead blue. 6. Sakuyok, ¢, July. |
45. HEMIPHAGA POLIOCEPHALA (144).
[Dinagat, 2, June. Iris Indian-yellow, passing into red on outer
circumference; orbital ring and patch crimson; bill black; feet
carmine ; nails dark grey. ]
Proc. Zoou. Soc.—1873, No. VIII. 8
114 ON A NEW PHILIPPINE BIRD. [ Jan. 15,
46. ORTYGOMETRA CINEREA (172).
[Dinagat, 2, June: iris crimson. |
47. HypoTznipi1a TorRQvATA (177).
{Dinagat, 9, June. |
48. BuroripEs JAVANICA (197).
[a. Dinagat, 9; 6. Nipah, ¢ 9, July: ins bright yellow; legs
bright chrome-yellow. |
No date on label of Dinagat example, but killed in June.
49. Dysporvus suta (214).
[Dinagat, ¢, June: iris white; feet light leaden-green ; bill
whitish green. |
In full white and rich brown plumage. Wing 15-30, tail 10-00,
culmen 3°90, tarsus 1°60.
11. On a new Philippine Genus and Species of Bird. By
Artuur, Marquis of Tweeppate, F.R.S., President of
the Society.
[Received December 12, 1877. |
(Plate IX.)
Until Dr. Steere some three years ago discovered in the island of
Basilan two species’ belonging to the Timelitde, this natural family,
so characteristic of the Indian region, was supposed to be unrepre-
sented in the Philippines. Since then Mr. Everett has discovered
a third species’ in the island of Dinagat, and a fourth, the subject
of this note, in the island of Negros. This last bird is so anomalous
in its structure that I propose to constitute it the type of a new genus,
DASYCROTAPHA’.
Base of maxilla densely clothed with short feathers, space behind
the eye naked. Bill about the length of head, and formed as in
Mixornis. Wing lengthened, longer than tail, first primary half the
length of the second, which is a fifth shorter than the third, third a
little shorter than the fourth, which is equal to fifth and sixth. Tail
moderate and square. Tarsus strong; hallux with claw stout
and long; digits short, slender; outer a little shorter than inner.
DasycRoTAaPHA SPECIOSA, sp.nov. (Plate IX.).
Head crested ; forehead with dense short plumes covering the
base of the maxilla; circle round the eye, whole space before the eye,
tuft on the side of the base of mandible, chin and uppermost
‘ Macronus striaticeps, Sharpe, and Dendrobiastes basilanica, Sharpe.
2 Mixornis capitalis, Tweeddale.
3 dacs, villosus, et kpdradot, tempora capitis.
ee. eo 1 te,
J.Smit hth. Hanhart imp.
DASYCRGTAPHA SPECIOSA.
aes $
= Pe
“, i ? ay (
: | os 4p we
:
fy
1878.] THE SECRETARY ON ADDITIONS TO THE MENAGERIE. 115
part of throat pure lemon-yellow ; crown of the head black ;
postoccipital plumes yellow ; nape yellowish green tipped with black ;
a lengthened tuft of plumes springing from above the eye bright
orange ; a line immediately below this tuft black ; a tuft of stiff
decomposed feathers springing from below the eye and extending over
the ears white or greyish white ; an irregular band across the throat
black ; dorsal feathers grey with light olive-green tips and white shafts;
uropygium yellowish green ; upper tail-coverts the same, tinged
with rufous; lower throat and upper breast bright yellow, most of the
feathers with black terminal drops ; lower breast and rest of under
plumage duller yellow tinged with green on the flanks ; quills brown
margined with yellowish olive-green, inner margin of quills pale
yellow ; wing-coverts dull olive-green, carpal edge and wing-lining
yellow ; tail dull rufous.
Wing 2°62, tail 2°37, tarsus 0°87, culmen 0°75.
Hab. Valencia, Island of Negros, ¢, August. Iris crimson;
bill orange-yellow. (Everett.)
The hereditary affinity of this new form with Macronus striati-
ceps and Miwornis capitalis is betrayed by the colouring and
markings of the dorsal plumage.
7
February 5, 1878.
Prof. St. George Mivart., F.R.S., V.P., in the Chair.
The Secretary read the following report on the additions to the
Society’s Menagerie during the month of January 1878.
The total number of registered additions to the Society’s Mena-
gerie during the month of January was 91, of which 43 were by pre-
sentation, | by birth, 41 by purchase, and 6 were received on deposit.
The total number of departures during the same period, by death and
removals, was 78.
The most remarkable additions during the month of January were
as follows :—
1. A Japanese Wild Dog, presented by Harry Pryer, Esq., of
Yokohama, January lst—an animal apparently allied to the “Dhole”
of India and the “Dingo” of Australia. Mr. E. W. Janson has
kindly favoured me with the following extracts from two letters which
he has received from Mr. Pryer on this subject.
“Yokohama, Oct. 8th, 1877.
“T have sent by the ‘ Loudoun Castle,’ via New York, a fine speci-
men of the Japanese Wild Dog. This is an animal quite new, and is
neither a Wolf nor an ordinary Dog. Its principal distinguishing
features are its long narrow feet and its head ; it most resembles the
dog used by the hunters here, one of which is figured in Siebold. Its
habits are totally different from those of any domesticated Dog. When
pleased it has a most extraordinary way of laughing, and also, when
pleased or very angry, has a curious dancing gait. ie specimen
116 PROF. MIVART ON THE FINS OF ELASMOBRANCHS. | Feb. 5,
was taken a puppy on the slopes of Fujisan last year. I bought him
for $11 in November last. He has learnt a sort of bark from the other
dogs which he did not know when I first had him; he comes out
sometimes with a regular wolf’s howl, but not often. In his muzzle
he resembles more the mainland Wolf, the northern Wolf being, like
the Siberian one, very long in the nose, and those on the mainland
(C. hodophylax) much shorter. My first impression was that this dog
was a cross between a wolf and the hunter’s dog ; but both its parents
then would have had very Jarge round feet, which this has not. A
friend told me that it is very like the Indian Dhole. In winter it has
a thick coat ; but in summer all the long thin hair comes out, and it
then has only its coarse wiry hair on. It is a much scarcer animal
than the Wolf on the mainland, which latter is rather common in the
mountains, anywhere where the sheep-faced Antelope is found.”
“Yokohama, Noy. 17th, 1877.
“T am trying to get a Southern Wolf (Canis hodophylax) to
send as a present to the Society. I have sent the Wild Dog in
the ‘ Loudoun Castle,’ and should like to hear what they say about
him. He is a complete puzzle to me. At one time I would look at
him and think that he was only an abnormal form of the common
Dog, run wild, and at other times feel as perfectly convinced that
he had nothing in common with it. He has very narrow feet, clean
limbs, very long canines, coarse hair in summer and plenty of long
thin hair in the winter, eyes and ears like a Wolf. I heard of
the animal wherever I went from the hunters, but only succeeded in
getting this one, as they say it is very difficult to catch, more so than
the Wolf, the capture of which is no easy task, as I know, as I have
been in places where they were plentiful, but only saw one and
poisoned another, although I have heard half a dozen at a time
howling quite close to me in the forests of Yamato.
“« My Wild Dog differs considerably from C. lupus, which is found
in Yesso, and C, hodophylax, which is confined to the Main Island,
although in his shorter muzzle he resembles the latter. When we
know more about the cave and other recent bone-deposits of Japan,
we shall be able to speak more definitely about him.”
2. A young Penguin of the genus Spheniscus, purchased January
24th. This bird is said to have been obtained in Chili, and is
probably the young of Spheniscus humboldti. lt is very tame, and
has been for the present placed in a compartment of the Fish-house,
where it seems likely to do well.
Prof. St. George Mivart, F.R.S., read a Memoir entitled, ‘“ Notes
on the Fins of Elasmobranchs, with Considerations on the Nature
and Homologies of Vertebrate Limbs,” of which the following is an
abstract :—
In this paper I describe (from dissections made for the purpose)
the skeleton of the paired and azygos fins, especially the dorsal fins,
of Zygena malleus, Mustelus antarcticus, Notidanus cinereus, Scyl-
1878.] PROF. MIVART ON THE FINS OF ELASMOBRANCHS. 117
lium canicula, Ginglymostoma cirratum, Chiloscyllium ocellatum,
Acanthias blainvillei, Spinax niger, Pristiophorus japonicus, Pristis
cuspidatus, Rhynchobates djeddensis, Trygonorhina fasciata, and
Callorhynchus antarcticus ; and I add notes made from preparations
of Lamna cornubica, Cestracion phillippi, Squatina angelus, Polyodon
foliosus, and Polypterus bichir.
Besides wishing to ascertain the positive conditions of skeletal
structures in these forms, the examinations were partly undertaken
with a view to the questions :—
(1) What is the nature of Vertebrate limbs generally ?
(2) What is the relation of Piscine to other limbs?
To these questions four others are subordinate :—
A. Are the paired-limb structures of a nature distinct from that
of azygos fins ?
B. Are paired limbs essentially axial structures which have become
more or less detached from skeletal axis, or peripheral structures which
have become secondarily more or less connected with it ?
C. What is the nature of limb-girdles ?
D. What is the line of genesis of the cheiropterygium.
I then note and discuss the opinions on these matters of Oken,
Carus, Cuvier, Owen, Maclise, Goodsir, Humphrey, Gegenbaur,
Macalister, Huxley, Balfour, and Parker.
I have myself arrived at the conclusion that the nature of the
paired and azygos limbs is fundamentally the same. I have, in fact,
been brought to this conviction by finding various degrees of coales-
cence between the cartilaginous rays supporting the dorsal fins, and
various degrees of connexion or continuity between such fin-supports
and the subjacent axial skeleton. I have noted coalescence amongst
the rays in Seyllium canicula, Ginglymostoma cirratum, and especially
in Notidanus cinereus, where it is carried to such an extent that the
rays are supported by one continuous basal cartilage. Continuity with
the axial skeleton is described as existing in the dorsal fin-cartilages in
several forms, but especially in Pristis and Pristiophorus; and I would
suggest that the lateral pressure of its saw-like rostrum must be more
or less aided by very firm fixation to the vertebral column of the
cartilages supporting the dorsal fin.
I have found much resemblance between the skeleton of the ventral
and the dorsal fins, as, for example, in Notidanus, in Chiloscyllium and
Raia; also between the anal and ventral fins, as again in Notidanus.
But the ventral fin of Polyodon is the most striking, presenting as it
does a longitudinal double series of simple parallel rays, quite like the
simplest form of the skeleton of the dorsal fins. Now, as the ventral
and pectoral fins are admitted on all hands to be of the same nature,
if the ventrals are of the same nature with the azygos fins, the pectoral
ones must be also of that same nature.
As to the objection which may be drawn from the attachment of
the pectoral fins to the axial skeleton by a shoulder-girdle instead
of by a direct continuous longitudinal adhesion, as in some dorsal fins,
I observe :—(1) The pectoral-fin support could not continuously adhere
to the axial skeletun longitudinally without impeding the flexure of
118 PROF. MIVART ON THE FINS OF ELASMOBRANCHS. [Feb. 9,
the body in swimming ; (2) the pectoral fins join the body at too low
a level to abut directly on the vertebral column; (3) such direct
connexion is prevented by the existence of the body-cavity.
I believe that the limb-girdles are lateral ingrowths from the
paired-fin skeleton (an idea already suggested by Mr. J. K. Thacher of
New Haven); and I believe such skeleton is the modified remnant of
a longitudinal series of similar rays formed primitively in a continuous
longitudinal lateral fold.
Similarly, I believe that the skeleton of the azygos fins is a structure
also formed primitively in a continuous median fold, and that the
dorsal rays are not outgrowths from the vertebral column. If, how-
ever, neural spines are homologous with the cartilaginous rays of
dorsal fins, then it seems to follow that the spinous processes of higher
animals and of man are essentially exoskeletal parts which have
adhered to, and grown to be connate with the axial skeleton.
As to the caudal fin, I note a constant difference both as to the
number and form of the skeletal parts of its dorsal and ventral por-
tions, seeming to point to some genetic difference needing investi-
gation.
I then proceed to the question of the homologies of piscine and
digit-bearing limbs, adopting the view that the preaxial margin of
the pectoral fin is that turned obliquely dorsad,—a matter which
seems established by its innervation, apart from other proof.
I oppose the view which adopts the Ceratodus type of limb as the
representative of the archipterygium (Elasmobranch ventrals certainly
lend no vestige of support to the theory), and then consider and criti-
cise the successive modifications of opinion expressed by Gegenbaur
and Huxley.
I regard some Rays as having pectorals hypertrophied indeed, yet
most like the true archipterygium (7. e. the least-modified by coales-
cence) of all Elasmobranch pectorals. As to Teleosteans, not only
do I think with Giinther that the arrangement of the limb-skeleton of
Ceratodus is foreshadowed in the pectoral fin of Acipenser, but I
think it probable that the Teleostean form of limb as seen in Anguilla
and Blennius affords us indications of a very primitive type, whence
the pectorals of Teleosteans on the one hand and of Elasmobranchs
on the other may both have been derived.
I cannot think that the metapterygium has developed into the limb-
axis of the digit-bearinglimb ; I believe such axis to be rather derived
from the mesopterygium, or (as this is sometimes absent, as in Chi-
loscyllium, or imperfect as in Polypterus) from the propterygium.
The conclusions I have arrived at as to the probable genesis of
Vertebrate limbs are as follows :—
(1) Two continuous lateral longitudinal folds were developed
similar to dorsal and ventral median longitudinal folds.
(2) Separate, narrow, solid supports in longitudinal series, and
with their long axes directed more or less outwards at right angles
with the long axis of the body, were developed in varying extents in
all these four longitudinal folds.
(3) The longitudinal folds became interrupted variously, the
1878.] PROF. MIVART ON THE FINS OF ELASMOBRANCHS. 119
lateral fold so as to form two prominences on each side, i.e. the
primitive paired limbs.
(4) Each anterior limb increased in size more rapidly than the
posterior limb.
(5) The bases of the cartilaginous supports coalesced as was
needed according to the respective practical needs of the different
separate portions of the longitudinal folds, 7. e. the respective needs
of the several fins.
(6) Occasionally the dorsal radials coalesced (as in Notidanus &c.)
and sought centripetally (as in Pristis &c.) adhesion to the skeletal
axis.
(7) The rays of the hinder paired limbs did so more constantly,
and ultimately prolonged themselves inwards by mesiad growths from
their coalesced bases, till the piscine pelvic structure arose as we see
it in Squatina '
(8) The pectoral rays with increasing development also coalesced
proximally, and, thence prolonging themselves inwards to seek a
point @ appui, shot dorsad and ventrad to obtain a firm support and
at the same time to avoid the visceral cavity. Thus they came to
abut dorsally against the axial skeleton and to meet ventrally together
in the middle line below.
(9) The lateral fins, as they were applied to support the body on
the ground, became elongated, segmented and uarrowed, so that
probably the line of the propterygium, or possibly that of the meso-
pterygium, became the axis of the digit-bearing limb.
(10) The distal end of the incipient cheiropterygium either pre-
served and enlarged preexisting cartilages or developed fresh ones to
serve fresh needs, and so grew into the developed cheiropterygium ;
but there is not as yet enough evidence to determine what was the
precise course of this transformation.
(11) The pelvic limb acquired a solid connexion with the axial
skeleton, a pelvic girdle, through its need of a point d’uppui as a
locomotive organ on land.
(12) The pelvic limb became also elongated; and in those cases
where its function was quite similar to that of the pectoral limb its
structure became also quite similar (e.g. Icthyosaurus, Plesiosaurus,
Chelydra, &c.); but for the quadrupedal mode of progression it
became segmented and inflected in a way generally parallel with, but
(from its mode of use) in most cases in part inversely to, the inflec-
tions of the pectoral limbs.
The amount of apparently spontaneous change needed to effect
these transformations may appear excessive ; but I believe that the
excessive plasticity of the animal organism is generally too little ap-
preciated—a plasticity which results in and is evidenced by the many
instances we now know of the independent origin of similar structures.
The plasticity of animals might be expected to be great ; for plasticity
is bodily reaction in response to external stimuli. The response which
ig most rapid and complete is sensation ; and an animal is a creature
' This view has been put forward by Mr. J. K. Thacher of New Haven,
Connecticut, See Connecticut Trans. vol. iii.
120 MR. W. A. FORBES ON BIRDS [Feb. 5,
the essence of which is ‘‘impressibility,” and may be described as
a more or less complex arrangement for carryiug about, nourishing,
and propagating a plexus of sensations.
The two initial questions may then, in conclusion, be answered as
follows :—
(1) Vertebrate limbs are differentiations of continuous lateral folds.
They are therefore not limited to four, and are for locomotive conve-
nience. There might apparently be several successive paired limbs on
each side, just as there are often several successive dorsal fins paired ;
and azygos fins and limbs being of the same nature, each separate
part may be called a pterygium, and for the sum total of the whole
the term sympterygium may be employed.
(2) Piscine limbs are related to digit-bearing limbs as structures which
have diverged less from the primitive condition, a natural consequence
of fishes making use of their fins in that medium in which the primi-
tive continuous lateral folds were first developed.
This paper will be puplished entire in the Society’s Transactions.
The following papers were read :—
1. Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘Challenger.—No. VII. On the
Birds of Cape York and the neighbouring Islands (Raine,
Wednesday, and Booby Islands). By W. A. Forsss,
F.Z.8.
[Received December 18, 1877.]
The collection of Birds made by H.M.S. ‘Challenger’ at Cape
York and in its neighbourhood, of which the following is an account,
comprises 61 skins, referable to 37 species, As might have been
expected, all, or nearly all, belong to well known Australian forms,
one or two only being left uncertain for want of more materials and
on account of the immature condition of the specimens. Most of
the skins are in excellent condition; and their value is much increased
by the notes in Mr. Murray’s journal as to the colour of the soft
parts &c. Besides Cape York, Raine Island (at the end of the
Barrier Reef), Wednesday Island (in Torres Straits), and Booby
Island (also in Torres Straits) were visited, and collections made. I
copy the following extracts from Mr. Murray’s journal as regards
the localities where birds were obtained :—
‘Raine Island, Barrier Reef, Australia. Ship landed two boats
for nearly three hours. The following birds were taken” (several
sea-birds, Rallus pectoralis, and Strepsilas interpres).
**Cape York, Somerset. Ship arrived on evening of Ist Sept.,
1874; left Cape York on Sept. 8th.” (44 skins were obtained here).
*‘Wednesday Island, Torres Straits. Parties landed the same
day (Sept. 8). Most of the birds seen were the same as those shot
1878. ] COLLECTED BY THE ‘CHALLENGER.’ 121
about Cape York. The following two, however, have not been pro-
cured at Cape York ; they were shot on shore (TYotanus incanus and
4igialites inornatus). A great many flocks of Bee-eaters were
noticed making passages between the islands.”’
* On 9th Sept., 1874, a party landed on Booby Island, a very
small island, with only a few shrubs on it. The following land birds
were shot or brought on board; and in addition a Rail was seen, a
Megapodius, and one other land bird.” (The species obtained were
Todirhamphus sanctus, Merops ornatus, Zosterops luteus, Pachy-
cephala sp., Ptilopus superbus, and Syncecus cervinus.)
In the following list I have, with a few exceptions, followed the
arrangement and nomenclature of Mr. Gould’s ‘Hand-book to the
Birds of Australia.’
1, Nrnox Boosook (Lath.).
Ninozx boobook, Sharpe, Cat. of B. ii. p. 168.
Spiloglaux marmoratus, Gld. Handb. B. Austr. i. p. 73.
One male skin of this species from Cape York. No. 167.
** Stomach contained insects”? (Murray, MS.).
The specimen sent agrees generally with two skins in Mr. God-
man’s collection from the same locality, and with Mr. Gould’s de-
scription of Spiloglaua marmoratus, which Mr. Sharpe, in the
second volume of his catalogue, treats as “the adult of the large
form of N. boobook”’ (l.c. p. 170).
2. PoDARGUS PAPUENSIS, Quoy & Gaim.
Podargus papuensis, Quoy & Gaim. Voy. de |’ Astrol. Ois. t. 13 ;
Gould, B. of Austr. Supp. pl. ; id. Handb. B. Austr. i. p. 91.
“No. 186, 2. Eyes red, feet and bill light yellow. Stomach
contained insects.’”” Cape York, one specimen.
3. Merops ornatus, Lath.
Merops ornatus, Gld. B. of Austr. ii. pl. 16; id. Handb.i. p. 117.
Tn all four specimens of this common Australian species. Two
females from Booby Island (Nos. 199, 200), and a pair (¢ & 2)
from Cape York (nos. 147 and 178). Of all the eyes are stated to
be “red,” and the feet and legs black, except the female from Cape
York, which is described as having the legs “with violet tint.’
The three females differ considerably from one another in the length
of the produced middle tail-feathers ; in the male the produced part
is thinner and at least twice as long as in the other sex.
4, TopIRHAMPHUs SANCTUs (Vig. & Hors.).
Todirhamphus sanctus, Gld. Handb. B. Austr. i. p. 128 ; Sharpe,
Alced. pl. 91.
One specimen from Booby Island. ‘No. 192, ¢. Eyes black ;
mandibles black, except base of lower one, which is whitish.
Stomach had remains of a crab.”
122 MR. W. A. FORBES ON BIRDS [Feb. 5,
5. Hatcyon macreayt, Jard. & Selby.
Haleyon macleayi, Jard. & Selby, Il. Orn. vol. iii. pl. 101; Gld.
B. of Austr, ii. pl. 24; Sharpe, Ale. pl. 78.
Cyanaleyon macleayi, Gld. Handb. B. Austral. i. p. 133.
One male specimen from Cape York. ‘No. 161. Eyes, bill, and
legs black. Stomach contained insects.”
6. ARTAMUS LEUCOPYGIALIS, Gld.
Artamus leucopygialis, Gld. B. of Austr. ii. pl. 33; id. Handb.
i. p. 154.
One specimen from Cape York. “No. 177, ¢. Bill blue tipped
with black, feet black. Stomach contained insects.”’
7. GRAUCALUS HYPOLEUCUS, Gld.
Graucalus hypoleucus, Gld. B. of Austr. ii. pl. 57; id. Handb.
i, p. 196.
One skin of this species from Cape York. “No. 151, 2. Legs,
bill, and eyes black. Stomach contained insects.”
8. PacHYCEPHALA, sp. ine.
“No. 196, g¢. Pachycephala. Eyes brown, bill and legs horn-
colour. Stomach had insects.”
A single skin of a -Pachycephala from Booby Island, immature,
still retaiming some of the rufous feathers characteristic of imma-
turity in its wings. It is certainly one of the species which, when
adult, are yellow beneath, the under tail-coverts being bright gam-
boge-yellow. The species of this group are so hard to determine,
unless males and in adult plumage, that it seems better to leave this
bird, although not exactly like any Pachycephala I have been able
to examine in the British Museum or in Mr. Godman’s collection,
without a name for the present, more particularly as Mr. Sharpe is,
I believe, now working at this group for the next volume of his
catalogue.
9. PINAROLESTES RUFIGASTER (Gld.).
Colluricincla rufigaster, Gld. B. of Austr. i. p. xxxvii; id. Handb.
B. Austr. i. p. 226.
Pinarolestes rufigaster, Sharpe, Cat. of B. iii. p. 296.
No. 170, 9. Snrike.
One specimen, from Cape York, of this difficult genus. Mr. Gould,
to whom I showed the specimen, was inclined to identify it with his
Colluricincla parvissima* (Ann. & Mag. N. H. ser. 4, x. p. 114), and
has kindly lent me the type of that species (which Mr. Sharpe, ¢. ¢.
p- 297, treats as the young of P. rufigaster) for comparison. I find,
however, that the Cape-York bird has a distinctly shorter wing, a
stronger and more arched bill, and is less rufous on the underside.
Mr. Sharpe, who has recently worked up this group for his cata-
logue, has examined this specimen, and refers it to P. rufigaster.
Count Salvadori suggests that this really=P. megarhynchus of New Guinea.
1378. ] COLLECTED BY THE ‘CHALLENGER.’ 123
10. Manucop1a Goutnt, Gray.
Manucodia gouldi, Gld. Handb. B. Austr. i. p. 236; Sharpe,
Cat. of B. iii. p. 181.
Manucodia keraudreni, Gld. B. Austr. Suppl. pl. 9.
Two female specimens. ‘No. 152. Bower-bird. Eyes orange,
bill and legs black. Stomach contained small seeds. For curious
loop in the wind-pipe see body. his bird was shot on the island
opposite Somerset by Moseley.” The colours of the soft parts and
contents of the stomach are the same in the other specimen.
The two skins sent agree well with Mr. Sharpe’s description and
differ from Mr. Elliot’s figure of the New-Guinea species (M. he-
raudrent) in the points noticed in Gray’s original description of the
species. The curious conformation of the trachea in M. heraudreni
has long been known, having been originally described by Lesson.
Beccari (Ibis, 1876, p. 252) says he thinks it probable that this
is a peculiarity of the male sex. Mr. Murray’s notes, however,
would seem to contradict this view, unless, as is hardly likely, two
species otherwise so nearly allied should differ in this peculiarity,
11. Pritoruis ALBERTI, Elliot.
Ptiloris alberti, Ell. P. Z.S. 1871, p- 583; id. Mon. Parad. pl.
xxiv.; Sharpe, Cat. of B. iii. p. 156.
Ptiloris magnificus, Gould, B. Austr. Supp. pl. 51.
Craspedophora magnifica, id. Handb. i. p. 595.
Six skins of this Rifle-bird from Cape York, four of which are
adult males in full plumage, and one a female. Another, marked Oe
is either a young male coming into full plumage, or more probably
an aged female assuming male plumage, as some of the feathers on
the top of the head show traces of a metallic blue colour, as also
does the shoulder, whilst more or fewer of the primaries on each
side have acquired the black colour characteristic of the male. Of
Nos. 164 and 165 it is noted in Mr. Murray’s journal :—<“ Males:
eyes, bill, and legs black, soles of the feet yellow. The stomach
contained a red fruit, with a large stone about the size of a pea.
Some parts of insects in 165.” Of No. 184, “Female: eyes black;
stomach contained ants and grubs.” Besides the specimens sent,
several others seem to have been procured. An interesting account
of the shooting of these specimens will be found in Lord George
Campbell’s ‘ Log Letters from the ‘ Challenger,’’ p. 185.
12. Mimeta virints (Lath.).
Mimeta viridis, Gld. Handb. B. Austr. i. p. 462.
Oriolus viridis, Gld. B. Austr. iv. pl. 13; Sharpe, Cat. of B. iii.
p. 212.
A young male from Cape York, agreeing generally with Mr.
Sharpe’s description (/. ¢c.) of the young bird. “No. 169, ¢.
Thrush.”
13. Mimera rLavocrncrtus (King).
Mimeta flavocineta, Gid. Handb. B, Austr. i. p. 466.
a
124 MR. W. A. FORBES ON BIRDS [Feb. 5,
Oriolus flavocinctus, Gld. B. of Austr. iv. pl. 14.
Oriolus flavicinctus, Sharpe, Cat. of B. iii. p. 206.
An adult male from Cape York, agreeing with skins in Mr. God-
man’s collection, collected by Cockerell. “No. 189, ¢. Shrike.
Eyes red, bill red, feet bluish.”
14. SPHECOTHERES FLAVIVENTRIS, Gld.
Sphecotheres flaviventris, Gld. B. of Austr. Suppl. pl. 37; id.
Handb. B. Austr. i. p. 468; Sharpe, Cat. of B. ili. p. 225.
Three specimens from Cape York, of which two are males in full
plumage, agreeing with specimens in Mr. Godinan’s collection.
“Nos. 172, 173. Eyes black, feet flesh, bill black.” The third
specimen is marked female (‘‘No. 174, 2. Eyes black, bill horn,
legs brownish. Stomach contained insects and seeds’’), but, as there
are traces of yellow on the breast, is more probably a young male
assuming the adult plumage. The orbits in this bird (in the skin)
are dark brownish black, not yellow, though this is probably acci-
dental. In other points, too, particularly in the plainly striated under
tail-coverts, and the colouring of the upper surface of the head, the
specimen hardly agrees with Mr. Gould’s description of the female
S. flaviventris. Mr, Gould, to whom I have shown it, would give
no decided opinion on it; but Mr. Sharpe has examined it, and pro-
nounces it to be of this species.
15. Pritotis curysotis, Lewin.
Ptilotis chrysotis, Gld. B. of Austr. iv. pl. 32.
Ptilotis lewinii, Swains.; Gld. Handb. B. Austr. i. p. 503.
Three skins of this common Australian species, which is not re-
corded from N. Australia in the ‘ Hand-book’—two from Cape York,
one from Cape-York Island. ‘No. 150, 2. Eyes brown, legs
slate, bill black. Stomach contained insects.’ ‘No. 157, ¢.
Eyes brown, bill black, legs blue, different from No. 156 (P. filigera).
Stomach contained insects, same as 156.”
16. PritoTis FILIGERA, Gld.
Ptilotis filigera, Gld. B. of Austr. Suppl. pl. 42; id. Handb.
B. Austr. 1. p. 522.
A single example, from Cape-York Island, of this distinct Honey-
eater, originally described by Mr. Gould from this district. ‘“ No.
156, ¢. Eyes brown, bill black, legs bluish. Stomach contained
insects and small brown bodies like seeds.”
17. PHILEMON BUCEROIDES, Swainson.
Tropidorhynchus buceroides, Gld. B. of Austr. Suppl. pl. 44; id.
Handb. B. Austr. i. p. 547.
One specimen from Cape York, agreeing with specimens in Mr.
Godman’s collection. ‘‘No. 160, 2. Leatherhead. Eyes dark
red, bill and skin about the head black. Stomach contained beetles
and other insects.”
1878. } COLLECTED BY THE ‘ CHALLENGER.’ 125
18. Myzometa osscura, Gld.
Myzomela obscura, Gld. B. of Austr. iv. pl. 67; id. Handb. B.
Austr. i. p. 559.
A single skin from Cape York. ‘‘No. 159, ¢. Eyes brown'.’’
19. DicauM HIRUNDINACEUM (Shaw).
Diceum hirundinaceum, Gld. B. of Austr. ii. pl. 34; id. Handb.
B. Austr. i. p. 581.
A single specimen in full plumage, from Cape York. “No. 149,
3. Eyes black, legs and bill black. Stomach contained green
fruit.”
20. NecraRINiA FRENATA, Mill.
Nectarinia australis, Gld. B. of Austr. Suppl. pl. 45; id. Handb.
B. Austr. i. p. 584.
Three specimens from Cape-York Island (two males in full
plumage, and a female) of this Sun-bird, which Mr. Sclater has re-
cently shown to be the Nectarinia frenata of Miiller (see P. Z.S.
1877, p. 104). ‘No. 153, 3; 154, ¢; 155, 9. Eyes brown, legs
and bill black. Stomach contained small insects.”
21. ZosreRoprs LuTevs, Gld.
Zosterops luteus, Gld. B. of Austr. iv. pl. 83; id. Handb. B.
Austr. i. p. 590.
One specimen from Booby Island (“ No. 197, 2. Eyes brown,
feet and bill with a bluish tinge. Stomach had small seeds”), which
agrees with a specimen of this species collected by Macgillivray,
kindly lent to me by Mr. Gould for comparison.
22. CACOMANTIS FLABELLIFORMIs (Lath.).
Cacomantis flabelliformis, Gld. Handb. B. Austr. i. p. 618.
Cuculus cineraceus, Gld. B. of Austr. iv. pl. 86.
A female, from Cape York, of this Cuckoo, agreeing with specimens
in Mr. Godman’s collection, except in having the breast and under-
parts much duller rufous, and faintly freckled and banded with
dusky— indications, probably, of a young bird. “‘No.146, 2. Cuckoo.
Eyes brown, legs yellow on hind aspect, brown on front, bill brown.
Contained insects in stomach.”
23. CENTROPUS PHASIANUS (Lath.).
Centropus phasianus, Gld. B. of Austr. iv. pl. 92; id. Handb. B.
Austr. i. 634.
A female from Cape York. ‘No. 163. Legs bluish, bill horn.
Stomach contained insects.”
24. MicRoGLossuM ATERRIMUM (Gmel.)
Microglossum aterrimum, Gld. B. of Austr. Suppl. pl. 61; id.
Handb. B. Austr. ii. 27; Finsch, Pap. i. p. 370.
A pair of specimens from Cape York. ‘No. 185, ¢: eyes
2 Mr. Gould says “bright red.”
126 MR. W. A. FORBES ON BIRDS [Feb. 5,
black ; stomach contained seeds and fruit. No. 188, 2: stomach
contained fruit, same as 185.” The male has the beak larger, and
feathers of the crest more developed than the female.
25. TRICHOGLOSSUS SWAINSONII, Gld.
Trichoglossus swainsonii, Gld. B. of Austr. v. pl. 28.
Trichoglossus multicolor, Gld. Handb. B. Austr. ii. p. 93.
A pair of this well-known species from Cape York. ‘No. 145,
3. Legs and eyes grey. Bill orange, with a darker tinge. Green
fruit.” ‘No. 166, ¢. Same as 145.”
This species of Trichoglossus is not recorded from the north part
of Australia in Mr. Gould’s Hand-book, though, as noticed by
Finsch (Papag. ii. p. 822 and 824), it extends northwards as far as
Cape York.
26. Pritopus suPeRBus (Temm.).
Ptilonopus superbus, Gld. B. of Austr. v. pl. 57.
Lamprotreron superbus, id. Handb. B. Austr. ii. p. 108.
Two males and a female, from Booby Island, of this Fruit-pigeon.
“No. 193, 2; 194, g. Eyes yellow, bills greenish, feet red.
Stomachs had nothing in them.” ‘‘ No. 198, ¢. Dove; same as 194.
Stomach contained nothing.”
27. CARPOPHAGA ASSIMILIS, Gld.
Carpophaga assimilis, Gld. B. Austr. Suppl. pl. 67.
Megaloprepia assimilis, Gld. Handb. B. Austr. il. p. 111.
A male from Cape York, agreeing in coloration and size with
specimens in Mr. Godman’s collection. ‘‘No. 180: Feet green,
eyes red, bill greenish yellow, red at base and about nostrils.”
28. GEOPELIA HUMERALIS (Temm.).
Geopelia humeralis, Gld. B. of Austr. v. pl. 72.
Erythrauchena humeralis, id. Handb. B. Austr. ii. p. 142.
One from Cape York. “No. 171, ¢. Eyes red, feet purple,
bill black. Stomach contained fruit.”’
29. GEOPELIA TRANQUILLA, Gld.
Geopelia tranquilla, Gld. B. Austr. v. pl. 73; id. Handb. B.
Austr. ii. p. 144.
A single specimen, apparently immature, of this little ground-
dove from Cape York. ‘No. 148, 2. Eyes white, cere blue, bill
slate, legs pale flesh-colour. Stomach contained small seeds.’’
30. MreGaropius TuMuLuvs, Gld.
Megapodius tumulus, Gld. B. of Austr. v. pl. 79; id. Handb.
B. Austr. ii. p. 167.
A single specimen, unsexed, of a Megapode, from Cape York,
agreeing with Mr. Gould’s description of M. tumulus. “No.
168.”
1878. ] COLLECTED BY THE ‘CHALLENGER.’ 127
31. TaLeGaLua LaTHAMi (Gray).
- Talegalla lathami, Gld. B. of Austr. v. pl. 77.
A male from Cape York, a district from which it is not recorded
by Mr. Gould.
“No. 187. Eyes yellow’, head red. Stomach contained seeds.”’
32. TuRNIx MELANOTA, Gld.
Turnix melanotus, Gld. Handb. B. Austr. ii. p. 182.
Hemipodius melanotus, B. of Austr. v. pl. 84.
Two females of this species, originally described by Mr. Gould
from Moreton Bay, agreeing perfectly with the description in the
‘Hand-book.’ ‘No. 176. Cape York, Albany Island. Eyes
white, feet yellow, bill yellow and black. Stomach contained in-
sects.” “No.195. Booby Island. Eyes white, legs yellow, base of
bill yellow, tip black. Stomach contained small round seeds.”
33. Syna@cus cervinus, Gld.
Syneecus cervinus, Gld. Handb. B. Austr. ii. p. 195.
A Quail, from Cape-York Island, which agrees with specimens of
this species in Mr. Gould’s collection. ‘No. 175, 3. Eyes brown,
feet yellow, (bill?) bluish black.”
34. ANGIALITEs INORNATUs (Gld).
Hiaticula inornata, Gld. B. of Austr. vi. pl. 19.
Ochthodromus inornatus, Gld. Handb. B. Austr. ii. p. 237.
A female, from Wednesday Island, agreeing with specimens in
Mr. Godman’s collection, but with the markings on the face and the
pectoral band somewhat darker. ‘‘No. 191. Wednesday Island.
Eyes, legs, and bill black.”
35. ToTanus INCANUS (Gm.).
Totanus pulverulentus, Mill. Natuurk. Verhand. Land- en Volkenk.
p. 152.
Totanus griseopygius, Gld. B. of Austr. vi. pl. 38.
Gambetta pulverulentus, Gld. Handb. B. Austr. ii. p. 268.
One from Wednesday Island. ‘‘No. 190, 2 Sandpiper. Legs
yellow, bill blackish, eyes black. Stomach contained remains of
Crustacea.”
36. STREPSILAS INTERPRES, L.
Three specimens from Raine Island. “No. 128, ¢. Eyes brown,
feet light red with black claws, bill black. Stomach contained small
calcareous particles.” ‘No. 129, 9. Sameas 128.” “No. 142, 9.
Same as 128.”
37. Raxuvus pecroratis, Gld.
Rallus pectoralis, Gld. B. Austr. vi. pl. 76.
Two males and a young female, from Raine Island, of this well-
t “ Brown,” according to Mr. Gould; his was probably a younger bird.
128 MR. F. NICHOLSON ON BIRDS FROM ABEOKUTA. [Feb. 5,
known species. The young bird, No. 130, resembles the old one
generally in coloration, but has all the colours duller, the transverse
barring of the lower surface much less conspicuous and altogether
wanting on the breast, the white spots of the scapularies absent,
those on the wing-coverts smaller, and those on the tertiaries also
smaller, and tinged with rufous; the primaries also are less con-
spicuously banded with rufous.
‘Nos. 126, 127, males. Eyes red, upper mandible dark, the
lower with a slight red tinge, feet light brown. Stomach contained
beetles, egg-shells, and small calcareous particles.”
2. On a Collection of Birds from Abeokuta. By Francis
NicHoxson, F.Z.S.
[Received December 20, 1877.]
(Plate X.)
I have lately received from my friend, Mr. Henry Robin, a small
collection of birds made by him in the neighbourhood of Abeokuta.
Although preserved in spirits, the birds arrived in very good con-
dition, and have made fair specimens. On showing them to my
friend Mr. R. Bowdler Sharpe, he strongly advised me to publish a
short list of the species, inasmuch as we have never received any
natural-history specimens from Abeokuta before. This collection
also adds a species or two to the avifauna of Western Africa ; and
one bird appears tobe new. I have deposited a series of the skins in
the British Museum ; and Mr. Sharpe has confirmed my identifica-
tions. I am indebted to him for a comparison of some of the more
obscure species.
1, Scors teEvcotis (Temm.); Sharpe, Cat. B. ii. p. 97.
One specimen, which has apparently died in captivity. I am not
aware of the occurrence of this species in West Africa below Senegal]
having been noted.
2. MeLm=NORNIS EDOLIOIDES (Sw.) ; Sharpe, Cat. B. ili. p. 315.
One specimen, somewhat extending the range of this. species in
Western Africa.
3. Lantus smiTuit, Fraser; Sharpe, Cat. Afr. B. p. 50. no. 478.
Two adults, and three young birds in changing plumage; all ap-
parently males, with no red on the flanks.
4. Dryoscopus GamBensis (Licht.); Sharpe, Cat. Afr. B.
p. 48. no. 453.
One male specimen.
5. Dryoscopus masor, Hartl.; Sharpe, Cat. Afr. B. p. 47.
no. 450.
An adult specimen.
or
Zot fase ls Xe
C. Davenport del et lith,
|
| AMADINA SHARPEI.
Mintern Bros imp
1878.] MR. F. NICHOLSON ON BIRDS FROM ABEOKUTA. 129
6. Burais crisoxa (L.) ; Sharpe, Cat. Afr. B. p. 42. no. 391.
One specimen.
7. Exmry1a Loncicaupa (Sw.); Sharpe, Cat. Afr. B. p. 42.
no. 400.
Four specimens.
8. Bras musicus (V.); Sharpe, Cat. Afr. B. p. 43. no. 403.
An adult male.
9. Puarystrra cyANEA (Miil.); Sharpe, Cat. Afr. B. p. 44.
no. 412.
A large series of specimens, containing both sexes and the young
birds.
10. Hirunpo xruropica (Blanf.) ; Sharpe, Cat. Afr. B. p. 46,
no. 432.
The first occurrence of this Swallow in Western Africa, I
have compared it with several specimens from North-eastern Africa
in the British Museum, and I have no hesitation in pronouncing it
to be H. ethiopica. It is not quite so purple as Bogos-Land skins ;
but this is probably due to the fact of its having been preserved in
spirits.
11. Turpus ICTERORHYNCHUS, Cab.
T. pelios, Sharpe, Cat. Afr. B. p. 21. no. 187 (nec Bp.).
Four specimens.
12. CrinigeR stmpLex (Temm.); Sharpe, Cat. Afr. B. p. 22.
no. 197.
Four specimens.
13. Pycnonotus BARBATUS (Desf.); Sharpe, Cat. Afr. B. p. 23.
no. 2]4.
Five specimens.
14. CossypHa veRTICALIs, Hartl.; Sharpe, Cat. Afr. B. p. 26.
no. 237.
Three specimens.
15. Hypouars opaca (Licht.); Dresser, B. Eur. part xxviii.
One specimen.
16. Crinnyris FuLIGINosus (Shaw).
Nectarinia fuliginosa, Sharpe, Cat. Afr. B. p. 39. no. 362,
One specimen.
17. CrnNyRIS SPLENDIDUs (Shaw).
Nectarinia splendida, Sharpe, Cat. Afr. B, p. 38. no. 357.
One specimen.
Proc. Zoou. Soc.—1878, No. IX. i)
130 MR. F. NICHOLSON ON BIRDS FROM ABEOKUTA. [Feb. 5,
18. Crnnyris VENusTUus (Shaw).
Nectarinia venusta, Sharpe, Cat. Afr. B. p. 40. no. 376.
Three specimens.
19. PHoLipAUGES LEUCOGASTER (Gm.); Sharpe, Cat. Afr. B.
p. 04. no. 514.
Two pairs, the males exhibiting the very extreme of the bluish
and reddish purple shades prevalent in this species.
20. HyPHANTORNIS BRACHYPTERA (Sw.); Sharpe, Cat. Afr. B.
p- 59. no. 561.
One example.
21. Hypaanrornis TexTor (Gm.); Sharpe, Cat. Afr. B. p.
59. no, 257.
Two adult males.
22. NiGRiTa CANICAPILLA(Strickl.) ; Hartl. Orn. W.-Afr. p. 130,
The single specimen sent is more ashy and not of such a clear
cinereous grey as Gaboon examples, and the spots on the wings are
smaller ; the shade, however, may be owing to the specimen having
been preserved in spirits, The conspicuous white spots on the
wings and the white rump show that the bird belongs to N. cani-
capilla and not to N. emilie.
23. EKUPLECTES FRANCISCANA (Isert) ; Sharpe, Cat. Afr. B. p. 62.
no, 988.
Four adult males in full plumage.
24. Vipva princrpatis (L.); Sharpe, Cat. Afr. B. p. 63. no. 600.
A female bird.
25. HypocuEera cHatyBeEata (Miull.); Sharpe, Cat. Afr. B.
p- 64. no, 604.
26. PyrRENEsTES CaPITALBUS (Temm.) ; Sharpe, Cat. Afr. B.
p. 68. no. 650.
A female bird, doubtless belonging to this species.
27. SPERMOSPIZA HZMATINA (V.); Sharpe, Cat. Afr. B. p. 68.
no, 653.
One specimen, not in full plumage.
28. CrirHaGRa curysopyGa (Sw.); Sharpe, Cat. Afr. B. p. 67.
no. 638.
One specimen.
29. AMADINA SHARPII, n. sp. (Plate X.).
Supra olivascenti-flava, tectricibus alarum dorso concoloribus ;
remigtbus fuscis, extts dorsi colore marginatis, secundartis in-
1878.] MR. F. NICHOLSON ON BIRDS FROM ABEOKUTA. 131
ternis dorso concoloribus: caudd olivascenti-flavd, rectricibus ex-
terioribus intus fuscis ; pileo summo colloque postico cinereis ;
Sronte angustd, loris et facie laterali albis; lined angustd
pone-auriculari nigrd; guld totd nigrd; jugulo et pectore
summo cinereis ; abdomine, subcaudalibus tibiisque cinerascen-
tibus; corpore laterali leté flavo; subalaribus fulvis, flavo
lavatis ; margine alari flavo; remigibus infra brunneis, intis
albicanti marginatis ; rostro nigro ; pedibus brunneis. Long.
tot. 4°3, culm. 0°4, al@ 2°35, caude 1°95, tarsi 0°65.
The accompanying figure will give a good idea of this beautiful
little Finch, which is not allied to any other African species, and
appears to be quite new to science.
30. CypsELus pArvus (Licht.); Sharpe, Cat. Afr. B. p. 2.
no. 13.
One specimen,
31. Merors surerci.iosus (L.); Sharpe, Cat. Afr. B. p. 3.
no. 2).
One specimen.
32. Isprptna Picta (Bodd.); Sharpe, Cat. Afr. B. p. 7. no. 53.
Five specimens.
33. HaLcyon SENEGALENSIS (L.); Sharpe, Cat. Afr. B. p. 8-
no. 66.
One specimen.
34. Curysococcyx KLAAst (V.); Sharpe, Cat. Afr. B. p. 13.
no. 112.
Ten specimens, in all states of plumage.
35. BARBATULA SUBSULPHUREA (Fras.); Sharpe, Cat. Afr. B.
p- 16. no. 139.
Three specimens.
36. Denpropicus GorRT# (Miill.) ; Sharpe, Cat. Afr. B. p. 18,
no. 165.
One specimen.
37. PerisTERA TYMPANISTRIA (Temm.); Hartl. Orn, W.-Afr.
p. 197.
One specimen.
38. Acritrs nypoLEeucus (L.); Hart]. Orn, W.-Atr. p. 235.
One specimen.
9*
132 REY. S. J. WHITMEE ON THE HABITS OF FISHES. [Feb. 5,
3. On the Manifestation of Anger, Fear, and other Passions,
in Fishes, and on the Use of their Spines.* By the Rev.
S. J. Warmer, C.M.Z.S.
[Received January 2, 1878.]
In his remarks on the erection of the dermal appendages by
various animals under the excitement of anger and terror Mr.
Darwin (‘ Expression of the Emotions in Men and Animals’) confines
himself to “three of the great Vertebrate classes,” viz. mammals,
birds, and reptiles (p. 95). When I first read the passage referred
to, I tried to think of some reference to the manifestation of anger
by fishes in books with which I am familiar, but I failed to call any
to mind. And as Mr. Darwin has not brought forward any ex-
amples as to how fishes manifest their ‘emotions,’ I am almost
compelled to believe no observations have been recorded. I read,
too, in the article ‘ Ichthyology,’’ in the ‘ Encyclopzedia Britannica’
(vol. xii. p. 221, Sth ed.), after a fanciful comparison between birds
and fishes, to the great disadvantage of the latter, that “the silent
dweller in the deep knows few attachments, expresses no language,
cherishes no affections, . . . the cravings of hunger seem alone
to exercise a frequent and influential action over his monotonous
movements ;” and that “by whatever scenes in nature fishes are
surrounded, their perceptions are probably indistinct and dull” *.
Notwithstanding that so little appears to be written on the mani-
festation of feeling by fishes, it strikes me many persons who have
kept an aquarium must have noticed that they show anger quite as
plainly as any other animals.
For some time past I have kept a good sized aquarium for the
instruction and amusement of myself and my family. My house is
situated on the coast, and has a belt of shallow and smooth water
before it, formed by the coral-reef which runs along in front of it
nearly a mile out at sea. This lagoon is a fine place for fishes; and
I encourage the natives to bring me any thing remarkable they obtain
when fishing there. I have therefore often had fishes belonging to
several genera together in my aquarium. Amongst other things I
have had opportunities of watching their quarrels, which are by no
means infrequent among the individuals of the same species, and
which are of constant occurrence between different species and genera.
The signs of anger in most fishes are so obvious that one of my boys,
between three and four years old, who is fond of making his own
observations in natural history, knows an angry fish as well as most
people know an angry bull or an angry boar.
1 This paper was written in Samoa, in October 1875, and is given in its
original form. §. J. W., December 27, 1877.
2 There are other remarks in the article mentioned which, I presume, will be
corrected in the new edition of the ‘ Encyclopedia ’—such, e. g., as the statement
that, the elegant and diversified colours of fishes are probably for the special
gratification of man. Unfortunately these quickly disappear when man gets
possession of the fish.
1878.| REV. S.J. WHITMEE ON THE HABITS.OF FISHES. 133
When swimming in a placid condition, the anterior portion of the
dorsal fin in fishes is seldom elevated ; hence the spines which occupy
this position are laid back more or less closely, and are often scarcely
visible’. The anal fin is also often not fully expanded ; but I have
not usually seen this so marked as in the dorsal. On the least fright
up goes the dorsal, and the spines are at once elevated. As an ex-
ample of how slight a cause will suffice to produce this change, I
may mention that, before writing the last sentence, I rose from my
desk and lightly tapped the front of my aquarium. The anterior
dorsal fins of its inhabitants (previously laid back) were instantly
elevated, and all assumed the attitude of attention. Under the
influence of great anger or fear, such as would be caused by the
presence of a carnivorous fish in the aquarium, the dorsal fin is
raised to its extreme height, and the spines, both of the dorsal and
anal fins, are very prominent. The scales all over the body are
also more or less raised, and with them, of course, any other dermal
appendages the particular fish may possess. This causes it to
assume larger proportions than under ordinary circumstances.
I believe the spines are elevated under the influence of fear as well
as by anger. I was once trying to catch a Tetrodon nigropunctatue
which was in my aquarium, when it inflated itself and elevated the
fine spines with which its body was covered, and which were pre-
viously buried in its loose and flabby skin. This of course was
under the influence of fear. And this appears to give a hint as to
the purpose for which this fish and those of the genus Diodon inflate
themselves.
When I have observed fishes chasing each other, apparently in
playfulness rather than anger, I have often seen them swim at each
other with open mouths as if to bite. But when they have appeared
to intend a serious attack they have always turned suddenly round
and lashed at one another with the caudal fin. I believe serious
fighting is always done with the tail. And from their anatomy
one would naturally expect this to be the chief mode of attack in
most fishes.
I incidentally mentioned this way of fighting by fishes in a paper
recently sent to the Society, in which I described an attack made
by some small fishes on an Antennarius*. These little things were
evidently in great dread of their carnivorous neighbour. But,
like small birds in presence of a bird of prey, they could not keep
at a distance, but continually tried to torment it. In attacking it
they always took care to strike at its posterior part, although this
was protected by a block of coral. I said, in the paper referred to,
that this mode of attack by the tail ought to be very effective in the
case of an Acanthurus, and that it may account for the armature of
the Acronuridee.
In connexion with this subject I will mention a thought which
has often occurred to me as to the chief purpose served by fishes’
' Drawings of fishes usually represent them with al/ the fins fully expanded
and the spines prominent ; but this is not their ordinary condition,
? See P. Z. S. 1875, p. 545.
134 REY. S, J. WHITMEE ON THE HABITS OF FISHES. [ Feb. 5,
spines, viz. protection against the attacks of those of the class which
are carnivorous. I have never seen a fish try to use its dorsal spines
actively to strike with. But I have seen a Balistes swim rapidly
past an antagonist, and graze its side with its file-like lateral spines.
I am strongly inclined to believe the dorsal and anal spines are used
for defence only.
There is little doubt that most carnivorous fish capture their prey
by outswimming them, as most carnivorous quadrupeds capture theirs
by outrunning them. In the shallow waters within our reefs it is a
very common thing to see shoals of fish leaping out of the water
when chased by the larger ones which prey upon them. They pass
along with a series of springs at a great rate; and the noise they
make is heard at a long distance. Most fishes thus chased would, if
captured, be seized from behind. If so, the strong spines on the
anal and dorsal fins, inclined as they are backwards, would often be
of immense value to their possessors in preventing them from being
swallowed. I am inclined to believe this is the chief, if not the sole,
use of these spines’.
If this view be correct, those fish most exposed to the attacks of
others which are carnivorous ought (other things being equal) to be best
protected. I have not given sufficient attention to the subject to say
whether this be so or not. But from a merely superficial examina-
tion I fancy further investigation would prove that spiny fishes are
more frequently found in confined and shallow waters, where they
have little opportunity of escape: and that slow-swimming fishes are
most frequently protected with peculiar defensive armature. The
Diodon and Tetrodon may serve as examples of the latter. They
swim slowly, and if unprotected would be specially liable to be
preyed upon by carnivorous fish. Unless alarmed neither look for-
midable. In both of them the spines are ordinarily concealed in the
soft and loose skin ; and then they present a very different appearance
from the stuffed specimens or plates by which they are chiefly known.
But the individuals of both genera possess the power of very rapidly
inflating their loose skins, and thus erecting the spives with which
they are more or less completely covered. In the case of the Diodon
there can be no doubt that its inflated and bristling appearance would
serve to protect it from attack; and I imagine the most voracious
monster would think twice before attempting to feast on the less-pro-
tected Tetrodon.
1 Since the above was written I have seen a proof of the correctness of this
view as to the use of spines. I was one day passing along the lagoon on the
shores of Upolu, when my boatmen noticed a great splashing in the water at a
distance. Knowing the cause, they pulled for the place. There we saw a large
fish with a smaller one sticking in its jaws, caught apparently by the erected
spines. They got separated just as we approached, and before my crew could
secure them; but the natives told me they not unfrequently take fishes in that
way. Sometimes they find both dead, one firmly sticking by its spines in the
jaws of the other.— December 1877.
“Ne
eee nme ee nme en Eno mMEnere
Gort PIEYWEL
ies &;
ii oy
(4 ‘
7 4 ia
r af
g)
Rar nist
. ————" DPIGRBESOLIdivid
“UHL FAAS L
ue}
1878.] ON BIRDS FROM SOUTH AMERICA. 135
4. On the Collection of Birds made by Prof. Steere in South
America. By P. L. Sctarer, M.A., Ph.D., F.R.S.,
and OsperT Satvin, M.A., F.R.S.
[Received January 3, 1878.]
(Plates XI.—XIII.)
At the request of Prof. J. B. Steere, of the University of Michi-
gan, U.S. A., we have had great pleasure in examining the collection
of birds which he made during his recent transit across South Ame-
rica, from Para to Callao, and in determining and naming the speci-
mens.
In Nov. 1870, Prof. Steere went from Para first to Vigia, on the
south bank of the Amazons, near the mouth, above fifty miles below
Para. Returning to Para he next visited Arare, on the south side
of the island of Marajo. On finally leaving Para for the interior,
in June 1871, he ascended the river to Santarem, and from Santarem
visited Obidos on the north bank. From Santarem he continued
upwards to Manaos, or Barra, as it was formerly called, and thence
made an excursion up the Purus river to Ituchy and Marrahan.
Leaving Manaos in Oct. 1871, he ascended the main stream to
Pebas, in Peru, where several skins, having the well-known “make” of
Mr. Hauxwell (amongst which is a skin of Porzana hauxwelli, nobis),
were procured. From Pebas Prof. Steere continued the ascent of
the river to Barrancas and Pongo Manseriche, at the foot of the
Andes, but returned downwards to the mouth of the Huallaga, and
then ascended that stream to Yurimaguas, where he arrived in
March 1872.
Here the voyage by steam came to an end, and Prof. Steere pro-
ceeded across country, via Tarapota, Moyobamba, Rioja, Chacha-
poyas, and Caxamarca, to the Pacific coast at Truxillo. From
Truxillo Prof. Steere kept along the coast, via Pacasmayo to
Sorritos} and Tumbez, where, as will be seen by our notes, many
of the local and interesting species of Western Peru were met with.
From Tumbez Prof. Steere went to Guayaquil, and thence up to
Quito and back. Next he proceeded to Lima by Callao, and made
an excursion from the Peruvian capital up the Rimac and over the
Andes, to the vicinity of Cerro de Pasco, returning finally to Callao.
From Callao Prof. Steere sailed, in May 1873, for China and the
Philippines, where he made those great discoveries in ornithology
which have already rendered his name well known to ornitholo-
gists *.
! Sorritos I do not find marked in the maps; but Professor Steere tells us it
is on the coast, about 20 miles south of Tumbez.
’ 2 See Mr. Sharpe’s memoir in Trans, Linn. Soe, n. s. Zool. yol. i. p. 807 e¢
seqg. (1877).
136 MESSRS. SCLATER AND SALVIN ON BIRDS [Rebs Ss
During his journey across South America Prof. Steere made a
collection of 911 skins of birds, belonging to about 362 species of
the following groups :—
ex. sp. ex, sp.
1. Pagseres .... 503. 145 9. Herodiones .. 13 9
2. Cypseli...... 108 52 10. Anseres 4,..°5 <2 5 4
38 Biches eek 23 17 M=*Columbx, ..>: 27. 14
4. Coceyges.... 93 45 12. Galline ...... acl 5
Oe PSittacke erik OD 16 Poe AaTHEs fee ac Te 4
6. Striges ..-...; 14 5 14. Limicole .... 41 18
7. -Aceipitres... .;..: 23 16 15. Gavi. :....'. 7 5
8. Steganopedes. 38 3 16. CrypturL ... 36. 9 4
As might have been expected from the fact of Prof. Steere not
having made a lengthened stay at any of the localities visited, the
greater number of the specimens which he collected belong to
well-known species, which we have named according to our ‘ No-
menclator Avium Neotropicalium.’ But there are examples of
twenty-two species amongst them, either unknown to us or other-
pik of interest, concerning which we have the following notes to
offer.
1. ORYZOBORUS ATRIROSTRIS, Sp. NOV.
Ater eneo-nitens unicolor ; remigibus primariis ad basin albis,
speculum parvum teetricibus absconditum efficientibus ; rostro
erasso, nigro; caudd paulum rotundata: long. tota 60,
ale 2°9, cauda 2°7, tarsi 0°75.
a. Head of Oryzoborus crassirostris.
b. Head of O. atrirostris.
Hab. Moyobamba, Peru (Steere).
Obs. Ab O. crassirostri et affinibus rostro robustiore et nigro
diversus.
This Oryzoborus, of which Prof. Steere only obtained a single
example, now in the Museum of the University of Michigan, is
quite new to us’. It is readily distinguishable from O. erassiros-
1 Sporophila othello, Bp. Consp. i. p. 498, ex Am. Centr. (Mus. Berol.) is
unknown to us and indeterminable, unless the type is existent.
1878.] COLLECTED BY PROF. STEERE IN SOUTH AMERICA. 137
tris and its representative forms (O. maximiliant, O. occidentalis,
and O. melas) by its black bill, which, as will be seen by the out-
line given, is also more robust than in O. crassirostris.
There is a small white speculum, which does not extend onto the
outer web of the three outer primaries, and is concealed by the
greater coverts when in their natural position. On the under sur-
face of the wing the white extends barely half an inch beyond the
black under wing-coverts, and then passes into grey, and at the
extremities of the feathers into black.
2. PirzorRHINA CINEREA, Laft.
Guiraca cendre, Préyv. et Des Murs, Voy. ‘ Venus,’ Ois. p- 209.
Gutraca cinerea, Lafr. Mag. de Zool. 1843, pl. 20 (subg. Piezo-
rhina).
Camarhynchus cinereus, Gray and Mitch. Gen. B. ii. p- 369 ;
Bp. Consp. i. p. 542; Salvin, Trans. Zool. Soc. ix. p- 491; Tacz.
P. Z.'S. 1877, p: 321.
MM. Jelski and Stolzmann first discovered the true habitat of
this curious Finch, which, through an error of the Naturalists of the
Voyage of the ‘ Venus,’ had been assigned to the Galapagos,
Prof. Steere’s series contains a single skin obtained in Dec. 187 2,
at Sorritos, in Western Peru. It is marked “ male: eyes light
hazel.” It agrees in every respect with an example from Tumbez
(Jelski) in Sclater’s collection.
3. Ha@MopuILa sTOLZMANNI, Tacz.
Hemophila stolzmanni, Tacz. P. Z. 8. 1877, p. 322, pl. xxxvi.
fig. 2.
Of this recently described Finch, from the same district, Prof.
Steere likewise obtained a single example at Sorritos, in Dec. 1372.
It is marked “male: eyes hazel.” The discovery of a typical
Hemophila (of which the six previously known species are entirely
confined to Central America) in Western Peru is a fact of the highest
interest.
4, GNATHOSPIZA RAIMONDH, Tacz..
Gnathospiza raimondii, Tacz. P. Z. 8. 1877, p. 320, pl. xxxvi.
fig. 1.
Of this peculiar Fringilline form Prof. Steere obtained four exam-
ples during his sojourn at Sorritos.
5. IcTERUS GRACE-ANN&, Cassin.
Ieterus grace-anne, Tacz. P. Z. 8. 1877, p. 323.
Prof. Steere’s series contains two skins of this well-marked Icterus,
described in 1867 by the late Mr. Cassin, from a single specimen in
the museum of the Academy of Philadelphia, of which the exact
locality was uncertain. The western sea-board of Ecuador and Peru,
however, is its undoubted habitat, as the late Prof. James Orton
138 MESSRS. SCLATER AND SALYIN ON BIRDS [ Feb. 5,
obtained specimens at Machala, near Guayaquil, as recorded by
Salvin (Ibis, 1874, p.323). Jelski and Stoltzmann also found it at
Tumbez, as recorded by Taczanowski, /. s. c.; and Prof. Steere col-
lected two examples—one at Pacasmayo on the coast north of
Truxillo, and the other at Tumbez.
6. CYANOCORAX MYSTACALIS.
Pica mystacalis, Geoffr. Mag. de Zool. 1835, pl. 34.
Cyanocorax mystacalis, Sel. et Salv. Nomencl. p. 39, et P.Z.S.
1876, p. 272; Tacz. P. Z.S. 1877, p. 323.
Cyanocorax ortoni, Lawr. Ann. Lyc. N. Y. xi. p. 166.
Of this fine species Prof. Steere obtained one example at Tumbez,
where MM. Jelski and Stoltzmann also procured it. Having now,
through Mr. Lawrence’s courtesy, had the opportunity of examining
the type of his C. ortoni, we are able to state that our anticipation -
as to its identity with C. mystacalis was quite correct. Prof. Orton’s
skin is labelled as having been obtained at “ Zicapa, Peru, Oct. 22,
1874.” It may be remarked, however, that, whereas in Prof. Steere’s
specimen the five outer pairs of rectrices are wholly white, in Prof.
Orton’s the pair next to the middle pair have the basal half blue
and indications of like colour in the centres of the adjoining pairs.
It is probably on some such specimen as this that C. dellus of
Schlegel * was established.
7. EUSCARTHMUS ZOSTEROPS
A skin of a Tyrant, obtained by Prof. Steere at Moyobamba, so
nearly agrees with FZ. zosterops, Pelzeln, Orn. Bras. p. 173, that we
are unwilling to separate it on the faith of a single specimen. But
it is certainly larger in its dimensions *, has a rather shorter bill,
and the edgings of the wing-coverts and secondaries are paler and
more distinct.
8. OrcHiLus EcAuDATUS (Lafr. et D’Qrb.).
A skin of this species from Rioja, Peru, seems to agree well with
Sclater’s example from San Esteban (P. Z.S. 1868, p. 631).
D’Orbigny’s original type was from Yuracares, Bolivia.
9. MyIARCHUS SEMIRUFUS, sp. nov. (Plate XI.)
Supra fuscescenti-cinereus ; uropygio et alarum caudeque mar-
ginibus castaneis, remigum omnium et rectricum mediarum parte
media mgricante ; subtis ochraceo-rufus unicolor ; rostro et
pedibus nigris; long. tota. 7°5, ale 3°4, caude 3°3, tarsi 1:0.
H1ab. Pacasmayo, Peru (Steere).
1 Cf. Sel. et Salv. P. Z. 8. 1876, p. 272.
Long. tota, ale, caude.
4 Fix. ex Borba (Natt.). ........:.20 40 2:0 LZ
», Moyobamba (Sfcere)...... 50 2°2 2:1
1878.] COLLECTED BY PROF. STEERE [IN SOUTH AMERICA. 139
Although so abnormal in colour, we cannot arrange this bird, of
which Prof. Steere unfortunately obtained but a single specimen, as
otherwise than a typical Myiarchus. The wings are rather short,
the third, fourth, and fifth primaries being nearly equal and longest.
The wing-end, embracing the first five or six primaries, is black, as
are the centre portions of the other remiges ; but the latter are bor-
dered above and below with rufous. In the tail the black colour is
confined to the inner webs of the three middle pairs of rectrices, the
two outer pairs being wholly rufous, and the next pair having only a
slight trace of the black patch.
10. MACHZROPTERUS PYROCEPHALUS, Sclater.
Macheropterus pyrocephalus, Sclater, Ibis, 1862, p. 176.
Of this beautiful Manikin two examples were procured by Prof.
Steere—one at Rioja, and the other at Moyobamba.
11. ATTILA TORRIDUS, Sclater.
Altila torridus, Sclater, P. Z.S. 1860, p. 280.
A single skin from Santa Rosa agrees with Sclater’s types of his
A. torridus, except in being rather brighter below and in being
rather shorter in the wing. We do not think it distinguishable.
12. FuRNARIUS PILEATUS, Sp. nl.
Supra castaneus, superciliis albis; pileo obscure brunneo; subtis
albus, lateraliter ochraceo perfusus; remigibus nigris rufo
bifasciatis ; rectricibus castaneis, macula in pogonio interiore
nigra preditis; rostro corneo, mandibuld inferiore ad basin
albicante ; pedibus nigris : long. tota 6°8, ale 3°5, cuude 2:4,
tarsi 1:1.
Hab. Santarem, Amazons (Steere).
Obs. F. figulo affinis, sed pileo brunneo ed pedibus majoribus
diversus.
A single skin of a Furnarius obtained at Santarem, on the south
bank of the Amazons, seems to be referable to a representative
form of F. figulus, of S.E. Brazil, distinguishable by its dark brown
cap. In F. figulus the head is of exactly the same colour as the
back. *
13. THAMNOPHILUS SIMPLEX, Sclater.
Thamnophilus simplex, Sclater, Ibis, 1873, p. 387.
A single skin obtained at Vigia exactly agrees with Sclater’s type
of this little-known species, now in his collection. It is marked
‘‘male: eyes red.”
14. STEATORNIS CARIPENSIS.
A skin of this bird in Prof. Steere’s collection is labelled as having
been obtained at Caxamarca, in the Peruvian Andes. This, so far as
we know, is much further south than any previously recorded locality
140 MESSRS. SCLATER AND SALVIN ON BIRDS (Feb. 5,
for this wonderful bird. Since we examined this skin, Mr. C. Buckley
has sent an example of Steatornis from Sarayacu on the Rio Bobo-
naza, a confluent of the Pastaza, in Ecuador, thus connecting Prof.
Steere’s Peruvian locality with those hitherto known for Steatornis
in Columbia and Venezuela.
15. Picumnus scLaTeRt, Tacz. P. Z. 8S. 1877, p. 327.
We are inclined to refer a skin of a Picumnus, obtained at Tumbez
in December 1872, to this recently described species, although in
this difficult group an actual comparison of specimens is necessary to
establish identity.
Prof. Steere’s specimen is marked ‘* male,”’ but agrees with a bird
described as a female by M. Taczanowski in having the head spotted
with white. If, therefore, the sexes are rightly determined in both
cases, it would seem that this species differs from most of its con-
geners in not having the head spotted with red or yellow in the male
Sex.
16. CHLORONERPES SEDULUS.
Campias sedulus, Cab. et Hein. Mus. Hein. iv. p. 153.
Chloronerpes sedulus, Scl. et Salv. Nomencl. p. 99.
A “male” of this little known species was obtained by Prof.
Steere at Obidos, on the Amazons. Sclater’s collection contains a
single female from Guiana; and the type described by Cabanis and
Heine was of the same sex. The male, which is hitherto undescribed,
differs from the female in having the feathers of the cap terminated
with red instead of yellow.
The species is easily recognizable by the large well-defined pale
yellow spots on the wing-coverts. Prof. Steere notes the eyes as
* blood-red.”
17. CoLapres RuPICOLA (Lafr. et D’Orb.).
Colaptes rupicola, Scl. et Salv. Nomenel. p. 101.
A “female” skin from Cajamarca, which we refer to this species,
differs slightly from skins obtained in Southern Peru by Mr. Whitely
(see P. Z.S. 1869, p. 154) in having the spots on the chest produced
laterally into narrow transverse bands. The rump and lower surface
are likewise rather deeper in colour.
The label states that the bird is found ‘‘in the Upper Andes, among
the rocks; nests in mud walls; eyes sulphur yellow.”
18. Capiro sTEERI, sp. nov. (Plate XII.)
Supra let viridis, pileo vivide coccineo, nuchd ce@ruled ; subtis
pallidé limonaceo-flavus, plagd gulari et alterd subpectorali
coceineis ; ventre viridi flavoque variegato; rostro plumbe-
scente, tomiis et apice flavis ; pedibus obscure plumbeis: long.
tota 5°8, ale 2°8, caud@ 1°8, tarsi O'S.
Hab. Moyobamba, Peru (Steere).
1878.] COLLECTED BY PROF. STEERE IN SOUTH AMERICA. 141
Obs. Affinis C. richardsoni, sed colore corporis superioris_palli-
diore viridi, nucha cyane, et pectore limonaceo-flavo sané diversus.
Of this beautiful new Capito, which we have the pleasure of dedi-
to its discoverer, there is, unfortunately, but one specimen in the
collection, now in the Museum of the University of Michigan. The
species is closely allied in form and disposition of colour to C.
richardsoni, but may be distinguished by the differences pointed out
above.
Among some birds lately sent by Mr. G. N. Lawrence of New
York to Sclater for determination, we find an imperfect skin, evi-
dently of the same species. It was obtained by Walter S. Church
at the ‘* head-waters of the Huallaga,” during his journey up that
river, and belongs to the collection of the Smithsonian Institution,
19. CoLuMBULA CAMPESTRIS (Spix).
Five specimens of this species are in Prof. Steere’s collection, all
of them obtained at Arare, in the island of Marajo. The species,
though long ago deseribed by Spix, has seldom been met with by
more recent collectors. Natterer, however, secured several examples,
mostly in the Brazilian provinces of Goiaz and Cuyaba,
20. Limosa FEDOA (Linn.).
Limosa fedoa, Scl. & Salv. Nomencl. p. 146.
One of two specimens of this Godwit was obtained at Santa Rosa,
Ecuador ; the other is without precise locality. Though the allied
L. hudsonica ranges throughout South America, this species, so far
as we know, has not hitherto been noticed south of Central America.
21. Xema SABINII, J. Sab.
Xema sabinii, Coues, Birds of the North-west, p- 660.
Prof. Steere’s collection contains a single example of the Fork-
tailed Gull, shot at Tumbez, Western Peru. This is the first instance
of the occurrence of the species in any thing like so southern a locality,
the limit of its southern range having been hitherto supposed to be
the Great Salt Lake, Utah. The specimen is in adult'winter plumage,
the head being white, the nape and back of the neck ashy black,
each feather with a very narrow white margin; the mantle is pure
grey, and the primaries coloured as in the adult bird.
22. CRYPTURUS TRANSFASCIATUS, sp.n. (Plate XIII.)
Supra ochraceus, dorso toto et alis extius nigro transfasciatis ;
cervice posticd murino-brunned ; pileo nigro, ochraceo brunneo
transfasciato ; superciliis, guld et abdomine medio albis ; cervice
umd et pectore griseis ; hypochondriis et tectricibus caude@ infe-
rioribus cervinis nigro transfasciatis ; rostro corneo, mandibule
basi flavicante, pedibus flavis: long. tota 12, ale 6:2, caude 2"1,
tarsi 2°9, rostri a rictu 1°5.
Hab. Santa Rosa, Ecuador (Steere).
142 PROF. A. H. GARROD ON THE BINTURONG, [Feb. 5,
Obs. C. variegato et C. bartletti forsan affinis, sed statura majore
et pectore griseo distinguendus.
This apparently undescribed Tinamou has the back conspicuously
transversely barred as in C. variegatus, but is much larger than that
species and has the whole of the lower part of the neck brownish
grey instead of cinnamon. C. bartletti, desczibed by us from speci-
mens obtained by Mr. E. Bartlett in Eastern Peru (P. Z. 8. 1873,
p. 311), is also a smaller bird, and has the whole under surface
except the throat fawn-colour ; it also wants the light supercilium
and the transverse marks on the occiput. In size the new species
approaches C. sallzi; but the totally different colour of the under
surface of this species renders further comparison unnecessary.
Prof. Steere’s collection contains two specimens of this species,
exactly resembling one another in colour, but differing slightly in
size, the bird described above being the larger of the two.
5. Note on the Anatomy of the Binturong (Arectictis
binturong). By A. H. Garrop, M.A., I’.R.S.
[Received January 7, 1878. ]
In an earlier communication ' I was able to confirm the observa-
tions of others, and to add fresh details, with reference to the anatomy
of Arctictis binturong. Since that paper was published, two other
specimens of the species have passed through my hands, the earlier
of which differed in no respect from the one which I had previouly
described. The last, however, which died on January 4, 1878, pre-
sented a peculiarity which I feel to be deserving of record. It was
a female, apparently adult, having lived in the Gardens of the
Society since October 19, 1875. The abnormal feature which it pre-
sented was the total absence of any trace of the colic cecum, which,
as is shown in a drawing accompanying my former paper, is normally
extremely small.
The line of separation between the small and large intestines is
well defined ; and there is no valvular constriction between the two
tubes, as is the case in the Arctoidea generally. There is a large
Peyer’s Patch quite close to the termination of the smaller gut.
The non-constancy of the presence of the diminutive cecum in
Aretictis binturong, and its total absence in Nandinia binotata’,
makes it evident that the existence of the cecum is a less import-
ant diagnostic character than was inferred by earlier investigators.
1 P. Z. 8. 1873, p. 196.
Vide Prof. Flower’s “ Note,” P. Z. 8. 1872, p. 683.
1878.] PROF. A. H. GARROD ON PASSERINE BIRDS. 143
6. Notes on the Anatomy of Passerine Birds. Part IV.
By A. H. Garrop, M.A., F.R.S.
[Received January 7, 1878.]
In my “Notes on the Anatomy of Passerine Birds,” Part II.,
whilst discussing the systematic position of the Eurylemide, I
mention that the method of insertion of the tensor patagii brevis
muscle of the arm is unknown’. Through the kindness of Lieut.-
Colonel Godwin-Austen, who has placed at my disposal carbolized
specimens of Psarisomus dalhousie and Serilophus rubropygius, I
have recently had the opportunity of dissecting these species, and
am now able to state that in both of them the tendon of the tensor
patagii brevis is perfectly passerine, agreeing precisely with the
typical arrangement previously described by me in the Order >—the
two tendons which traverse the distance between the point where
the main tendon joins the extensor metacarpi radialis longus and
the elbow running independently and parallel as they course to
their humeral attachment.
Tn neither specimen was I able to dissect out the plantar tendons,
because they were so dry and brittle that no amount of soaking
would render them fit for observation. I was, however, able to
determine some other anatomical points, including the existence of
a single carotid artery (the left), a normal disposition of the vessels
of the thigh (the main artery of the leg being the sciatic), the con-
‘siderable size of the femoro-caudal, of the semitendinosus, and of
the accessory semitendinosus, and the absence of the accessory
femoro-caudal and of the ambiens muscles.
Previous evisceration, I regret to say, prevents my adding any thing
with reference to the syrinx and the abdominal viscera ; nevertheless
I think that now the vomer and the tensor patagii brevis are known
to conform exactly with the Passerine type, all shadow of doubt as
to the perfectly Passerine structure of the Eurylemide may be dis-
missed, notwithstanding the aberrant vinculum joining their deep
flexor tendons.
So far as its sternum is concerned, Psarisomus dalhousie so
closely resembles Eurylemus javanicus, as figured by Mr. Sclater in
‘ The Ibis’, that no further remark with reference to it is neces-
sary, except to draw attention to the unforked condition of the
manubrium. My specimen of Serilophus was too much shot for me
to make out its sternal peculiarities.
1 P, Z. 8. 1877, p. 449.
2 P. Z, 8. 1876, p. 508.
$ Third Series, vol ii., 1872, p. 179.
144. MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. [Feb. 5,
7. Descriptions of new Species of Phytophagous Coleoptera.
By Martin Jacosy.
[Received January 10, 1878.]
Family EumM oLPip&.
Genus CHALCOPHANA.
1. CHALCOPHANA RUFIPENNIS, Sp. Nov.
Above, shining bright rufous; below, black; legs testaceous ;
head with a median depression, impunctate, labrum and antennz
light testaceous ; thorax finely and very sparingly punctate, together
with the head rufous; elytra of the same colour, distinctly and
regularly punctate-striate on the disk, more irregular towards the
sides, where also three distinct ribs or coste are visible running
parallel to each other. Underside black, with a metallic green tint;
legs and tarsi yellowish or testaceous.
Length 4 lines.
Hab. Costa Rica.
Collected by Mr. Rogers.
This species is closely allied to C. discolor of v. Harold, but
differs abundantly in the coloration of the antennze, the underside,
and legs.
2. CHALCOPHANA COSTATIPENNIS, Sp. nov.
Upper and underside rufous testaceous ; antennee black, the basal
three joints ferruginous ; femora red; tibiee and tarsi black ; head
minutely punctured, with a fovea between the eyes; thorax with a
few minute punctures ; scutellum smooth ; elytra each with about
seven distinct costz, all of which, with the exception of the fifth
and sixth, commence below the transverse and deeply marked basal
depression; the fourth, fifth, and sixth costz united at a little dis-
tance from the apex ; the interstices deeply punctate, the punctures
united in pairs.
Length, 3 to 33 lines.
Hab. Chontales, Nicaragua.
This species, although closely allied to C. discolor, Harold, and
C. rufipennis above described, is distinguished from both by the
strongly raised costee on the entire disk of the elytra and by the
entirely black tibiz.
Genus CoxasrorpeEs, Castelnau.
3. CoLASPOIDES VIRIDICOLLIS, sp. nov.
Oblong-ovate, convex, fulvous above ; head, thorax, and a broad
margin round the elytra bright metallic green ; head sparingly but
deeply punctate, with a deep longitudinal groove; antennz with
the first six joints, the labrum, and palpi fulvous, the last four joints
of the antennze much depressed and widened, of a fuscous colour ;
thorax with the anterior and posterior margin gradually produced
in the middle, the sides evenly rounded, with a shallow but distinct
1873.] MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 145
oblique depression at a little distance from the anterior angles,
surface distantly but distinctly punctate ; scutellum smooth, metallic
green. Elytra scarcely wider at the base than the thorax, evenly
rounded towards the apex, finely punctate-striate, the interstices
plane; the entire lateral margin, with the apex to a greater extent,
is of a bright, shining, metallic green, while the rest of the surface
is of a light fulvous colour, where the punctures are also surrounded
by a piceous spot. Underside (with the exception of the thorax and
part of the breast) and legs entirely fulvous.
Length 2 to 23 lines,
Hab. Amazons.
Collection Jacoby.
Family CoRYSOMELID.
Genus DorypHora.
4. DoryPHORA FLAVOGUTTATA, Sp. nov.
Ovate, convex, above dark brown with a slightly greenish tint,
each elytron with three fulvous spots; head greenish black, spa-
ringly but distinctly punctured, the punctures ou the clypeus more
strongly marked; antenne black, with the underside of the first
joint testaceous, as well as the apex of the last joint ; thorax with
the middle of the anterior margin nearly straight, the lateral margins
a little thickened and evenly rounded anteriorly, surface but little
shining, of the same colour as the head, deeply punctate, more
closely so towards the sides; scutellum triangular, smooth; elytra
much wider than the base of the thorax, convex till their middle,
but rather suddenly declining towards the apex, surface punctured in
the same way as the thorax, and partly arranged in double lines of
striee, each elytron with an oval patch near the scutellum, a trans-
verse one in the middle, and another smaller near the apex, the
latter spot sometimes obsolete, of a reddish fulvous colour. Under-
side and legs greenish, shining ; the mesosternal process straight and
very acute.
Length 5 lines.
Hab. Volcano of Irazu, Costa Rica.
5. DoRYPHORA PUNCTIPENNIS, Sp. nov.
Oblong, convex, above brown ; base, lateral margins, a transverse
vitta, and a subsutural stripe near the apex, of the elytra, flavous ;
head light brown, with a slight greenish reflection, moderately close
and distinctly punctured, labrum lighter-coloured ; antennz larger
than the thorax, with the first five joints shining brownish green,
the basal joint testaceous below, the rest blackish, hairy ; thorax
narrowed at the base, from there diverging to the apex, the anterior
angles acutely produced, of the same colour as ihe head, very
deeply and coarsely punctate towards the sides, on the disk a little
less closely, and leaving a longitudinal middle line smooth ; scutellum
impunctate ; elytra much wider at the base than the thorax, closely
variolose-punctate, the punctures arranged in strize towards the
suture, and everywhere distinctly visible to the naked eye, of the
Proc. Zoou. Soc.—1878, No. X. 10
146 MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. [Feb. 5,
same colour as the head and thorax, shining, the base and the
lateral margins (with the exception of a small space a little before
the middle, where the margin is interrupted by a brownish black
spot, which also extends across the elytral seam below), a transverse
vitta from one side to the other, and a short streak running from
the apex parallel with the suture flavous. Underside brownish
black.
Length 54 lines.
Hab. Volcano of Irazu, Costa Rica.
Collected by Mr. Rogers.
6. DoryYPHORA FLAVOMARGINATA, Sp. nov.
Oblong-ovate, subdepressed, greenish or brownish ; head, lateral
margin of thorax broadly, elytral one narrowly testaceous ; head
and mouth testaceous, the former closely punctate in front of the
eyes, nearly smooth in the middle, with a greenish V-shaped mark ;
antennee reaching to the base of the thorax, with the first four
joints testaceous, the rest more or less stained with greenish black ;
thorax with the front margin slightly convex in middle, and the
sides evenly rounded, surface closely and distinctly punctured at
the base only, the rest sparingly and very minutely ; a subquadrate
greenish black patch narrowed towards its base occupies the middle
of the disk, leaving the sides testaceous ; scutellum flavous, more or
_less stained with fuscous, smooth ; elytra very little wider at the base
than the thorax, semiflattened, of a greenish or brownish tint,
irregularly and rather closely punctate-striate, with fine transverse
and longitudinal scratches, the lateral margin from the shoulder to
the apex testaceous, only punctured near the inner edge, the outer
half perfectly smooth with the exception of a single row of rather
deep punctures near the extreme margin. Body below, with the
exception of the mesosternal process, the tibie externally, and the
margins of the abdominal segments (the latter of which are testa-
ceous) fuscous, shining.
Length 43 lines.
Hab. Peru.
Collection Jacoby.
7. DoRYPHORA DORSOMACULATA, Sp. nov.
Ovate, regularly convex, dark fuscous ; elytra light green, with a
pear-shaped sutural patch, fuscous ; head very finely punctate,
~ fuscous, labrum flavous, antenne with the first joint light, the rest
darker brown; lateral margins of the thorax rounded throughout,
slightly sinuate in middle, more dispersedly but not coarser-punctured
than the head, and of the same colour; scutellum subtriangular,
impuncetate ; elytra strongly punctate-striate (visibly to the naked
eye), the strize arranged in single rows, of which the first is short
and the seventh disturbed by a few extra punctures near its middle ;
the interstices also extremely finely punctate, of a light olive-green
colour, with the suture narrowly, the lateral margins more distinctly,
part of the extreme base, and, connected with it, a common sutural
1878.] MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 147
pear-shaped patch extending to nearly the middle of the length of
the elytra, fuscous brown; a narrow light-flavous stripe accom-
panies the marginal and sutural one through its entire length.
Entire underside and legs fuscous brown.
Length 5 lines.
This species was discovered by Mr. Rippon in Nicaragua.
8. DoRYPHORA INSULARIS, sp. nov.
Ovate, regularly convex, above greenish black, with a submarginal
broad longitudinal vitta light testaceous ; head sparingly and finely
punctate, labrum fulvous; antenne rather long, the last five joints
flattened and broad, of a greenish black colour ; thorax of the same
colour, throughout finely punctured with seven or eight deep punc-
tures parallel with the extreme lateral margin; scutellum smooth,
triangular ; elytra slightly wider at the base than the thorax, punc-
tate-striate towards the suture, the rest of the surface confused-
punctate of a more or less dark shining green colour, interrupted
only by a broad longitudinal stripe of light testaceous colour com-
mencing a little below the shoulder and extending not quite to the
apex. This stripe runs parallel with the extreme lateral margin,
but has the inner margin concave a little before the middle, but
nearly straight thence to the apex. Underside and legs of a little
darker green, shining.
Length 63 lines.
Hab. Peru.
Collection Jacoby.
Genus CaALLIGRAPHA, Erichs.
9. CALLIGRAPHA VIOLACEO-MACULATA, Sp. nov.
Oblong, convex ; head and legs ferruginous ; thorax and under-
side greenish, very shining ; elytra light flavous, the suture, three
transversely placed irregular spots at the posterior half, a median
one at the lateral margin, and an oblique, large, bifurcate mark con-
nected with the suture below the base, shining purplish; another
short branch of the same colour is sent off from the suture below
the middle of each elytron ; head very sparingly and minutely punc-
tured, with a triangular-shaped mark at the back greenish eneous;
antenne gradually thickened towards the apex, entirely ferruginous ;
thorax shining greenish, sides parallel at the base, rounded ante-
riorly, surface with a shallow fovea near each anterior margin,
minutely and sparingly punctate on the disk, with a few deeper
punctures near the base; scutellum triangular, the sides a little
rounded, greenish black, smooth. Elytra wider at the base than the
thorax, of a bright yellow hue, punctate-striate near the suture, the
rest of the punctures only plainly visible within the darker mark-
ings, the suture broadly, the lateral margins narrowly, purplish ; the
first is greatly widened in the middle, and connected with an
obliquely placed two-branched mark below the base, and another,
shorter streak with an upward direction below the middle; besides
10*
148 MR.M.JACOBY ON PHYTOPHAGOUS COLEOPTERA. [ Feb. 5,
this, and two transverse markings at the apex, and placed laterally,
is a distinct “~-shaped spot placed directly below the middle of each
elytron. Legs and apex of abdomen ferrugineous.
Length 45 lines.
Hab. Costa Rica.
Collection Jacoby.
The absence of all smaller spots usually found in the species of
this genus, and the peculiar pattern of the elytral markings, as well
as the purplish hue of the latter, will distinguish this species from
others.
Family GALLERUCID2.
Genus Diasrortica, Chevrol.
10. DiABROTICA SEXPUNCTATA, SD. nov.
Elongate, subparallel, above dark brick-red, each elytron with
three black spots ; head flavous, front impressed with a short longi-
tudinal groove, which terminates in a deep fovea, between the eyes,
vertex very minutely punctured ; antennz more than half the length
of the body, very light testaceous; thorax subquadrate, the lateral
margins bulging out slightly before the middle, with all the angles
obtuse, surface impunctate, but little convex, of a flavous colour,
stained with darker irregular patches ; scutellum smooth, triangular :
elytra wider at the base than the thorax, and increasing in width
towards the apex, distinctly margined at the sides, and closely punc-
tate, the interstices here and there subrugose; they are of a brick-
red colour, with a small round spot on the shoulder, another below
the base near the sutural margin, and a third behind the middle
black. Underside blackish, finely yellowish-pubescent ; tibia tes-
taceous ; claws darker.
Length 5 lines.
Hab. Volcano of Irazu, Costa Rica.
This insect closely resembles in colouring the D. tripunctata of
Oliv., but differs in the want of the impression on the thorax,
besides the different shape of the elytral spots.
11. DiaBROTICA VENTRICOSA, sp. nov.
Elongate-ovate, very convex and widened posteriorly, black ; head
and thorax ferrugineous ; elytra with two basal spots, a transverse
band in middle and another at the extreme apex flavous ; head finely
punctate, with two crescent-shaped elevations between the antenne ;
the latter very light testaceous, with the apical joints a little darker ;
thorax subquadrate, very convex, the posterior margin sinuatein middle,
and the sides very deflexed near the posterior angles, disk obliquely
foveolate on each side, finely punctured everywhere; scutellum
transversely impressed near the base, black ; elytra very swollen
posteriorly, and much widened from before the middle, distinctly
punctured, the punctuation partly arranged in strize, especially so
towards the apex, shining black; a large round spot near the
scutellum, another small one directly below the humeral callus, a
1878.] MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 149
transverse narrow band in the middle extending from one side to the
other, and a still more narrow and shorter vitta at the apex flavous.
Body beneath black ; legs and tarsi testaceous.
Length 3 lines.
Hab. Panama.
12. DiaBrorica FUSCOMARGINATA, sp. nov.
Elongate, convex, light brown above; head, disk of thorax, and
the elytral margin dark fuscous or black; head entirely black, im-
punctate, with a deep fovea between the eyes, the latter very promi-
neut and black, base of the antenne divided only by a sharp longi-
tudinal ridge ; palpi and antennz entirely flavous ; thorax scarcely
wider than long, quadrate, distinctly emarginate at the sides, the
latter slightly widened before the middle, from there straight to the
base ; surface very obscurely punctate, shining black or fuscous, with
the lateral margins flavous; scutellum also of this colour; elytra
much wider at the base than the thorax, slightly widened towards
the apex, with a narrow but very distinct margin, surface closely
and irregularly punctured, of a light brown colour, assuming a
flavous hue towards the apex, the base to a greater or lesser extent,
and a narrow stripe near the extreme lateral margins not reaching
the apex, dark fuscous. Underside, with the exception of the breast
(which is black), and legs entirely testaceous.
Length 42 lines.
Hab. Volcano of Irazu, Costa Rica.
Collected by Mr. Rogers.
13. DIABROTICA MULTIPUNCTATA, sp. nov.
Head, thorax, and a narrow sutural and lateral margin of the
elytra rufous ; elytra flavous, each elytron with eight small black
spots ; head impunetate, with a deep fovea in the middle; antennz
robust, dark rufous or fuscous; thorax transversely quadrate, disk
bifoveolate, with a black spot in front of each fovea ; scutellum black,
impunctate ; elytra subdepressed, widened behind, flavous, the spots
placed as follows—one immediately below the base near the scutellum,
a second one below the first but near the lateral margin, three
placed transversely in the middle, a triangular-shaped spot directly
below, and two others, transversely placed, at a little distance from
the apex, the sutural and lateral margins rufous. Body below
black, closely pubescent ; femora testaceous, with a black line above;
tibize and tarsi dark fuscous or black.
Length 3 lines.
Hab. Mexico.
Allied to D. mezicana, Harold, but principally differing in the
rufous emargination of the elytra.
14. Diasrortica NIGROLINEATA, Sp. nov.
Elongate, light flavous ; head and two longitudinal vittee on each
elytron black ; head impunctate, with a deep fovea in the middie;
antennz longer than half the length of the body, with the third
150 MR. M.JACOBY ON PHYTOPHAGOUS COLEOPTERA. [Feb. 5,
joint scarcely double the length of the second, the middle joints
fuscous, the first and three last joints almost. white, extreme apex
fuscous; thorax quadrate, obsoletely impressed on the disk, the
latter impunctate, shining flavous ; scutellum black ; elytra irregu-
larly and closely punctate, of a little lighter colour than the thorax,
the base, suture, a submarginal longitudinal stripe from the base to
the apex, and another stripe on the disk of each elytron, not reaching
the apex, black. Body below and femora testaceous ; breast, tibiee,
and tarsi black.
Length 3 lines.
Hab, Guatemala.
Collected by Mr. Salvin.
15. D1ABROTICA VIRIDIMACULATA, Sp. Nov.
Above black; thorax, a round spot near the base, a transverse
vitta behind the middle, the apex, and the extreme lateral margin of
each elytron greenish ; head black, shining, impunctate, with a deep
fovea in the middle; antennz long, filiform, with four middle joints
fuscous, the others light testaceous ; thorax quadrate, obsoletely
impressed on the disk with two foveze, and a smaller one near the
base ; scutelium obscure fuscous ; elytra widened behind, narrowly
margined, surface closely and finely punctate, with a short longitu-
dinal depression near the sides, black, the margins, a large round
spot occupying the apex, another smaller one below the base, and
a transverse band behind the middle, not quite touching the suture,
light greenish. Underside light flavous; breast, tibize, and tarsi
fuscous.
Length 3 lines.
Hab. Cayenne.
Collection Jacoby.
16. DIABROTICA GRACILIS, Sp- nov.
Oblong, greeu, each elytron with the base and a spot near the
apex fulvous; head black, impunctate ; antennz with the second
and third joints very short, equal, the first joint light green, the
second to the eighth fuscous, ninth and tenth joints light flavous,
and the apical one black ; thorax quadrate, narrowed near the base,
disk bifoveolate, bright green, obsoletely punctured ; scutellum
black, smooth; elytra closely punctate, a little rugose near the
sides, of the same green colour as the thorax, the base in shape of a
transverse band, and a large round spot near the apex, bright fulvous,
the suture in the neighbourhood of the scutellum, and extending to
the end of the basal vitta, narrowly edged with black. Underside
green; breast, tibize, and tarsi black.
Length 23 lines.
Hab. Brazil.
17. D1aBROTICA NIGROMACULATA, Sp. NOV.
Elongate, green; head and five markings on the elytra black ;
head shining, with the usual fovea in the middle; antennse dark
1878.] MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. 151
ferruginous ; thorax of the same shape as that of D. gracilis, obliquely
bifoveolate, green, impunctate; scutellum black; elytra a little
widened behind the middle, distinctly longitudinally suleate below
the shoulder, surface closely punctured, green, an irregular blotch
on the shoulders, a small elongate, triangular sutural mark near the
base and below the scutellum, and a transverse narrow spot near the
apex of each elytron, not touching the margin or the suture, black.
Underside and legs exactly as in D. gracilis.
Length 23 lines.
Hab. Ecuador.
Two specimens, collected by Mr. Buckley, in my collection.
18. DraBROTICA VARIOLOSA, Sp. nov.
Oblong, convex, widened behind, entirely dark green ; head and
disk of thorax dark fuscous; head impunctate, with a fovea in
middle and a highly raised ridge between the antennz ; the latter
with the second and third joints very short, the three basal joints
light green, the following three flavous, and the rest ferruginons ;
thorax quadrate, rather convex, the disk with two foveze and an
indistinct transverse fuscous mark; scutellum triangular, black ;
elytra convex, much widened towards the apex, and coarsely and
irregularly variolose and punctured, of adark green colour. Under-
side pitchy black, femora green; the breast on each side covered
with a longitudinal patch of golden-yellow hair, and the abdominal
segments margined with long greenish hair, which also covers the
legs.
Length 3 lines.
Hab. Ecuador.
Oue specimen in my collection.
This species is allied to D. marginata of Sturm, but is devoid of
the brownish margin round the elytra, and is distinguished by the
colour of the underside and legs as well as of the antenne.
19. DIABROTICA PERUANA, Sp. Nov.
Elongate, convex, much widened behind, black beneath ; thorax
flavous ; elytra flavous, with the base and a large spot near the apex
bluish black ; head impunctate, with two distinct tubercles in front
of the usual middle fovea; autenne long, filiform, entirely black ;
thorax subquadrate, impunctate on the disk, with two fovez near the
sides light flavous ; scutellum black; elytra of the same colour as
the thorax or a little lighter, with a broad band at the base, not
touching the lateral margins, and a large subquadrate spot behind
the middle bluish black ; the punctuation on the disk very distinct
and even coarse towards the base. The entire underside with the
legs and tarsi black, finely covered with yellowish hair.
Length 33 lines.
Hab. Peru.
20. DIABROTICA SEXPLAGIATA, Sp. nov.
Elongate, parallel ; head and thorax testaceous, the latter with a
152 MR. M. JACOBY ON PHYTOPHAGOUS COLEOPTERA. [Feb. 5,
longitudinal stripe on each side fuscous; elytra dark brown or
fuscous, the lateral margins and six roundish spots, 2, 2, 2, whitish ;
upper part of head finely punctured, in some specimens obscure, of
light brown colour, lower part light testaceous ; thorax transversely
quadrate, the sides rather broadly margined, surface bifoveolate,
impunctate, lighter or darker testaceous, with a longitudinal black
stripe near the lateral margins ; scutellum testaceous ; elytra closely
rugose-punctate, dark fuscous, with the margins, the apex, and 6
large roundish spots, placed transversely in pairs, whitish. Underside
and femora testaceous, with the tibize and tarsi fuscous, in one
specimen entirely testaceous.
Length 2 to 23 lines,
Hab. Peru, Panama.
Genus Ca@Lomera, Chevrol.
21. C@&LOMERA ATRO-CERULEA, Sp. Nov.
Broadly ovate, widened posteriorly; head, thorax, and apex of
abdomen rufous ; elytra bluish black ; antennz and legs black ; head
more or less deeply and longitudinally depressed, with a narrow mesial
groove through its entire length, finely punctate and flavous-pubescent;
antennze with the third joint double the length of the fourth, black,
the extreme base of the first joint flavous; thorax broadly rounded
at the sides, about twice as broad as long, surface deeply depressed
on each side, with a longitudinal groove in the middle of the disk,
throughout closely punctate, and covered with short flavous
pubescence ; scutellum rufous, the apex broadly rounded; elytra
much widened towards the middle, distinctly marginate, the sides
suddenly deflected from the shoulder, so as to form a distinct lon-
gitudinal ridge, which limits the disk of each elytron laterally to the
extent of its entire length, surface extremely closely punctate-rugose.
Underside dark fuscous, the last abdominal segment rufous ; legs
and tarsi black.
Length 4 to 43 lines.
Hab. Peru, Panama.
Notes on previously described Species.
CRIOCERIS AUSTRALIS, Jacoby, P. Z.S. 1876, p. 807.
This species, described by me in 1876, is (as a comparison with the
type in the British-Museum collection, of the existence of which I was
previously not aware, has convinced me) the true C. nigripes of
Fabricius. Lacordaire’s description of that species (vol. i. p. 573),
which I took as a guide, and which made me believe that the insect
before me was a different species, is totally at variance with the insect
of Fabricius ; and my description is therefore the right one, although
the name australis must stand as a synonym.
RHYPARIDA MADAGASCARIENSIS, P. Z. S. 1877, p. 512.
RHYPARIDA COSTATIPENNIS, P. Z.S. 1877, p. 513.
vg
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LOPHOSALEA ANAMALLAYANE
1878.] COL. R. H. BEDDOME ON A NEW TREE-LIZARD. 153
RuyPaRIDA NIGRICOLLIs, P. Z.S. 1877, p.513.
These three species, described by me under the generic name of
Rhyparida, belong, as subsequent studies have convinced me, to the
genus Syagrus of Chapuis.
8. Description of a new Genus of Tree-Lizards from the
higher Ranges of the Anamallays. By Lieut.-Colonel
R. H. Beppome, C.M.Z.S.
[Received January 12, 1878.]
(Plate XIV.)
LoPHOSALEA, nov. gen.
Gen. Char. No femoral pores, no lateral wings, tympanum naked ;
crest of the back and anterior portion of the tail very high, the lobes
separate ; scales of the back and sides large, irregular, imbricate, sub-
equal in size, but mixed with a few small scales, strongly keeled, the
tips directed backwards ; a few trihedral or spinous scales above the
tympanum ; a large gular sac; tail of moderate length, very much
compressed ; all the scales keeled, the lower ones very prominently.
LOPHOSALEA ANAMALLAYANA, Sp. nov. (Plate XIV.)
Three large scales between the base of the orbit and the top of
the tympanum ; the last of which is much raised and subtrihedral ;
a spine behind the tympanum, and another between it and the
commencement of the nuchal crest; crown of head concave; the
rostral shield is subvertical and over the first labial only (not hori-
zontal and over the first and second labials as in Salea) a few large
trihedral scales form a semicircle across the occipital region in front
of the nuchal crest ; seven or eight upper labials, the last very small ;
seven lower labials ; tympanum large ; eye rather small, eyelid large
and scaly as in Salea; longitudinal series of quadrangular scales on
each side of the gular sac, nuchal crest composed of two small and
four large scales, the interspace between this and the dorsal crest
very small and occupied by three small crest-scales ; the dorsal and
subcaudal crest is composed of twenty-eight large thin erect some-
what crimped scales one half an inch in length, and extends along
more than one half the length of the tail. Head much narrowed
in front, somewhat elongate and pointed as in Cophotis ; the hind leg,
if laid forward, only reaches the shoulder ; toes with carinated trans-
verse plates below, the third and fourth toes are very long, the
fourth being a little the longer ; length of body 42 inches, of tail 73
inches ; colour of body fawn with irregular transverse bars of white,
edged and intermixed with black, three across the body and eight
across the tail, a white band along the labials and from the angle of
the mouth to the shoulder ;_ belly white.
Hab. Anamallay Mountains, 6000 feet—a very rare lizard.
154 COL. R. H. BEDDOME ON NEW UROPELTIDA. [Feb. 5,
This genus is allied to Salea, from which it differs in its gular sac ;
it has more the aspect of the Ceylonese Cophotis, from which, how-
ever, it is further removed by its naked tympanum.
9. Descriptions of new Uropeltide from Southern India,
with Remarks on some previously-described Species.
By Lieut.-Colonel R. H. Buppomr, C.M.Z.S.
[Received January 12, 1878.]
SILYBURA NIGRA, 0. sp.
Snout short, slightly pointed, rostral not nearly separating the
nasals; eye very small, in the front part of the ocular shield, no
supraorbital ; scales all in nineteen rows, abdominals 180-185,
caudal disk convex, not well defined, scales slightly keeled, sub-
caudals six to nine pairs, terminal scute of tail bicuspid; body black,
very iridescent, with irregular transverse rows of yellowish-white
spots, a yellowish streak commences at the fourth labial and ex-
tends along the sides for 1 to 2 inches, sides and belly more or
less blotched with broad transverse patches of yellow, the first one
or two inches of the belly generally uniform black.
Hab. Pulney Mountains, 4000 ft. elevation.
SILYBURA NITIDA, 0. sp.
Snout obtusely pointed; rostral very small, not nearly separating
the nasals ; eye small, in the front part of the ocular shield ; no supra-
orbital ; fourth upper labial reddish yellow, caudal disk convex, in-
distinct ; scales three to six; keeled terminal scute rather large, not
laterally compressed, bicuspid, the points side by side, scales in
seventeen rows, abdominals 188 to 194. Colour nearly jet-black,
but very iridescent ; the belly with very distant large reddish-yellow
blotches, the black colour much predominating.
Hab. Anamallays, 4000 to 5000 ft.
STILYBURA PETERSI, N. sp.
Snout obtuse ; rostral very small, not separating the nasals; tail
laterally compressed, caudal disk not defined ; the scales nearly quite
smooth or faint traces of keels, the terminal seute shovel-shaped, not
bicuspid ; eye very small, in the front part of the ocular shield; no
supraorbital ; abdominals 155 to 158 ; subcaudals ten to twelve pairs ;
scales in seventeen rows ; above uniform brown; sides with indistinct
yellowish white spots or narrow transverse bars, which sometimes
extend across the belly ; a broad yellowish patch across the vent.
Hab. Anamallays, 4000 ft. Rare; a small species.
SILYBURA MACULATA, N. sp.
Snout obtuse ; rostral very smali, not separating the nasals, which
are as large as the frontals ; eye rather large, in the front part of the
ocular shield ; no supraorbital; tail laterally compressed, not well
defined ; caudal disk without keels, but a few of the terminal scales
a little rugose, terminal scale slightly bicuspid, the points side by
1878.] MR. H. SAUNDERS ON THE LARIN&. 155
side. Scales round the body in seventeen rows, abdominals about
156, subcaudals eight to thirteen pairs. Colour of a uniform brown
or sometimes quite black, with several deep-red blotches along the
sides of the anterior portion of the trunk and about the tail, and
rarely several of these are present along the sides of the trunk.
Hab. ?
The tail of this species is more like that of Plectrurus than that
of Silybura; but I prefer to restrict Plectrurus to those species
which have a supraorbital shield and only fifteen rows of scales.
The snake I formerly described as Plectrurus trilineatus has been
placed by Dr. Giinther in a new genus, “ Platyplectrurus:’’ this has
the supraorbital and only fifteen rows of scales, but it differs from
Plectrurus proper in the presence of an elongated temporal shield over
the fourth labial, besides in other poiats, as noted by Dr. Giinther ;
and I think the genus may stand. Plectrurus sanguineus (which I
find, on a reexamination of the species, has always fifteen rows of
scales only round the centre of the body, not seventeen) will belong
to this genus; and I have described two other species, P. hewstoni
and P. madurensis.
My Plectrurus pulneyensis has been placed by Dr. Giinther in
Rhinophis ; but it should, I think, rather be referred to Silybura.
My Silyéura canarica has been referred to Plectrurus: it has only
fifteen rows of scales (but it wants the supraorbital); the tail, however,
is exactly that of the type of Plectrurus, with the points of the
terminal scutes one above the other. It may form a distinct section
in the genus, or have a new genus constituted for it. The only
other St/ybura described with fifteen rows of scales is Silybura
macrolepis of Peters, which I have never seen, and its locality is
unknown ; it has evidently the typical caudal disk of Stlybura. Dr.
Giinther omits all mention of this species in his synopsis of the
species published in the Society’s ‘ Proceedings’ in March 1875.
I wish that I could attach the same importance as Dr. Giinther
does to the number of the abdominal plates; if he is correct in this,
I have several new species of the ocellated Silybure like S. ocellata,
hiura, and melanogaster.
Specimens of all the species described in this paper have been
sent to the British Museum.
10. On the Larine or Gulls.
By Howarp Saunpers, F.L.S., F.Z.S.
[Received December 31, 1877.]
A revision of the Larine is a more difficult task than that of the
Sternine, the members of the present subfamily being subject to far
greater variations in plumage on their passage from youth to age
than is customary with the Terns. The latter, although offering
differences in individual size and graduation of tint which are often
extremely puzzling, may be said to have only two, or at most three,
stages of plumage, viz. the immature, the winter-, and the adult
156 MR. H. SAUNDERS ON THE LARIN&. [ Feb.+4,
breeding-state, in the last of which the majority assume a black head
or crest; but with the Gulls these conditions are more complex.
Even in those species which are destitute of hood at all seasons there
is a seemingly endless variation in the pattern of the primaries, the
general tendency being to an increase in the lighter anda diminution
in the darker portions of the webs with the advancing age of the indi-
vidual—a rule which also holds good with many of those species the
adults of which bear a hood in the breeding-season, whilst, on the
other hand, there are others which exhibit the apparent anomaly of
having a hood in the immature stage, and losing it in the adult plu-
mage. The individual variations in size are even greater than in the
Terns; and the range of the Gulls being, as a rule, less extensive,
there are to be found several remarkable isolated and specialized
forms, side by side with others, which are little more than climatic
varieties of a general type. These circumstances have led to the
establishment of a multiplicity of genera and of species, many of
them exceedingly ill defined ; and it was not until I had examined
a considerable series of specimens here, and had visited the Museums
of Paris, Leiden, Mainz, Berlin, and Copenhagen, for the purpose
of identifying the types with the descriptions, that I could hope to
clear up some of the more obscure questions.
The literature of this group has been rendered especially intricate
through the perverted ingenuity of two systematists who have
undertaken its revision. Boie and Brehm are not guiltless in the
matter of genera- and species-making; but their labours were
chiefly confined to sorting the European Gulls backwards and for-
wards into fanciful groups, and to splitting up each species into
three or four, which can, for the most part, be easily referred back
to their origin. But when Bonaparte and Bruch undertook the
revision of the Larine of the whole world, they speedily enveloped
the question in a perfect fog of synonymy, their only object being,
apparently, to make as mauy genera and species as possible. Even dis-
tinct genera were erected for one and the same species in different plu-
mages; the most closely allied forms were placed far apart, and widely
divergent ones were united ; whilst it seemed to be accepted as an
axiom that a different geographical habitat was sufficient to constitute
a species. Revision followed revision; and to the work of the declining
days of both these authors we owe at least half of the synonymy
which encumbers these pages. It was their intention to perform
a similar office for the Terns; but death cut their plans short,
and to this is owing the comparative simplicity of the synonymy of
the Sternine.
The result of their labours appears in Bonaparte’s last completed
list (for that in the ‘Conspectus Avium’ was never finished), in the
‘Comptes Rendus,’ xlii. p. 770 (1856), in which he makes 68
“undoubted”’ species and 22 genera of Larine alone, besides 5
more species which he considered doubtful—with justice, as re-
gards four of them, two being his own, one Bruch’s, and one
Wagler’s, whilst the fifth, Larus fuliginosus, is an excellent species
with which he was evidently unacquainted. To this succeeded the
1878. | MR. H. SAUNDERS ON THE LARIN&. 157
reaction of common sense in the shape of one of Professor Schlegel’s
admirable Catalogues raisonnés of the Mus. d’Hist. Nat. des Pays-
Bas, a monographical review which, in the words of Prof. Blasius,
“tis destined to be the foundation of all true Gull-knowledge for
those whose ideas of what really constitutes a species are not sacri-
ficed to an arbitrary whim or a geographical sport.’”” The remarks
which I have thus translated are to be found in a critical review of
great merit in the J. f Orn. (1865, p. 369, and 1866, p. 73). In
this Blasius reduces Bonaparte’s species to 35—a diminution which
to some degree falls into the other extreme, partly owing to the fact
that the author was then unacquainted with several perfectly good
species, of which series have since become available, and also to his
having united some closely allied forms which, in my opinion, are
more conveniently treated by giving them specific rank.
As regards the North-American Larine, Dr. Elliott Coues has
contributed two important reviews (Proc. Ac. Nat. Se. Philad. 1862,
p- 291, and ‘Birds of the North-west ’ (1874); and these, coupled
with the excellent Revised List of the Neotropical Laride by Messrs.
Sclater and Salvin (P. Z. 8. 1871, p. 564), make us better acquainted
with the American species than with those of any other great divi-
sion of the globe. But although there are probably no undiscovered
forms, there still remain many details to be learned respecting the
haunts, nidification, and various plumages of the American Larine,
especially those of the Pacific-coast islands ; and, indeed, there are
two from the Galapagos Islands so rare that the one, L. fuliginosus,
is only to be found in three or four collections, whilst of the other,
Xema furcatum, only two examples are known to exist, the one in
the Paris, the other in the British Museum.
With regard to several of the Old-World species there are also
some important gaps to be filled up ; but with one solitary exception,
viz. Rhodostethia rosea, the least-known species and the most
interesting forms are those which are found on the shores and islands
of the Pacific, on both the Asiatic and American sides. These will
be noticed under their respective heads; and I will now pass on to
consider the genera and subgenera amongst which the species have
been divided. Most of these have been based upon colour, geogra-
phical distribution, or upon the mere caprice of the systematist—upon
any thing in short except those structural differences which afford
a valid reason for their employment; but as many of these genera
are either used erroneously or in a perverted sense by those who
have not studied the question, I will give a full synopsis of them with
remarks. Those genera which appear worthy of retention are printed
in small capitals.
GENERA.
Larus, Linn. Syst. Nat. i. p-. 224 (1766).
For all Gulls.
Xena, Leach; J. Ross, App. ii., Ross’s Voy. p. 57 (1819).
The generic characters are given as defined dy Leach; these
158 MR. H. SAUNDERS ON THE LARIN&. [Feb. 5,
are afterwards stated more fully by Stephens in Shaw’s Gen. Zool.
xili. p. 176 (1825). Type and sole representative then kuown, Xx.
sabinii. Principal characteristic, the forked tail.
Xema, Boie, Isis, 1822, p. 563.
For the European hooded Gulls ; but
Xema, Boie, Isis, 1844, p. 192,
Includes all those Gulls, either with or without hoods, which he
has not otherwise located under Gavia or Larus.
(N. B. Gavia, Moehring, 1752, Gen. Av. is prior to the 12th ed.
Linn., and need not be considered ; and Gavia, Brisson, is undefined.)
Gavia, Boie, Isis, 1822, p. 563.
No description, but is applied to two such structurally different
species as L. eburneus and L. tridactylus.
Gavia, Boie, Isis, 1826, p. 980,
Is limited to LZ. eburneus only.
Gavia, Boie, Isis, 1844, p. 191,
Contains the two former and L. audouini.
Gavia, Kaup, Nat. Syst. Eur. Thierw. pp. 99, 196 (1829).
For L. ridibundus and ‘“‘ L. capistratus.”
Gavia, Macgill. Man. Brit. Ornith. p. 239 (1842).
For all the hooded Gulls, including Xema sabinit.
Gavia, Bruch, J. f. Orn. 1853, p. 106.
For the small grey-mantled Gulls without hoods.
Rissa, Leach, Stephens in Shaw’s Gen. Zool. xiii. pt. 1. p. 180
(1825).
Type, R. brunnichii=L. tridactylus. Character—hind toe absent
or rudimentary.
Cheimonea, Kaup, Nat. Syst. Eur. Thierw. pp. 84, 196 (1829).
Type, RB. tridactyla.
Pacoruita, Kaup, op. cit. pp. 69, 186 (1829).
Type, Larus eburneus.
Leucus, misprint Lencus, Kaup, op. cit. pp. 86, 196 (1829).
For L. marinus, glaucus, and fuscus ; but
‘“Leucus, ex Kaup,”’ Bp. Consp. Av. ii. p. 215 (1857),
Omits the black-mantled species and includes the larger grey-
backed Gulls.
1878. ] MR. H. SAUNDERS ON THE LARIN&. 159
Hydrocoleus, Kaup, op. cit. pp. 113, 196 (1829).
For L. minutus and “L. plumbiceps,” “Gulls with black heads
and white eye-streak.”’
Ichthyaétus, Kaup, op. cit. pp. 102, 196 (1829).
Type and sole representative, L. ichthyaétus.
Laroides, Brehm, Vig. Deutschl. p- 738 (1831).
Includes most of the European hoodless Gulls.
Chroicocephalus, Eyton, Brit. Birds, p- 53 (1836).
Based upon the coloured hood, small size, and more naked tibia.
The latter characteristic only holds good with regard to a limited
number of hooded Gulls, and is by no means confined to them ;
whilst none of the other peculiarities adduced seem to be sufficient
for generic distinction.
The spelling of this word has been altered to
Kroicocephalus, Jameson, Journ. Asiatic. Soc. viii. p- 243 (1839),
Chroiocephalus, Reichenbach, Nat. Syst. Vég. p. v.,
Chrecocephalus, Strick], Ann. Nat. Hist. p. 40 (1841),
and to
Chroocephalus, Scl. & Salv. P. Z. S. 1871, p. 576 (note)*.
Rossia, Bonap. Comp. List B. Eur. & N. Am. p. 62 (1838).
For ZL. roseus. No description of generic character ; and the name
had already been employed otherwise by Owen.
Ruoposreruta, Macgill. Man. Brit. Orn. pt. ii. p. 252 (1842).
Type, L. roseus. Generic character described.
Cetosparactes, Macgill. Man. Brit. Orn. pt. ii. p- 251 (1842).
Type, Pagophila eburnea. Generic character described. Name
altered to
Catosparactes, Gray, Gen. Birds, iii. p. 655, note (1845).
*“‘Plautus, Klein,” Reichenbach, Nat. Syst. Vog. Longip. p. 5
(1852).
(NV. B. Klein’s Hist. Av. Prodromus, pp. 146-148 (1750), is out
of date ; and his Plautus includes Auks, Gulls, and Petrels.)
Glaucus, Bruch, J. f. Orn. 1853, p. 101.
For the large and medium-sized grey-mantled species.
“Gabianus, Bp.,” Bruch, J. f. Orn. 1853, p. 100 (description) ;
Bonap. Naumannia, 1854, pp- 211, 215; Consp. Av. ii. p. 212 (1857)
Type, L. pacificus.
|» Whilst these sheets are passing through the press, Mr. H. T. Wharton
(‘Zoologist,’ March 1878, p. 105) has pointed out the existence of an adjective,
Xpwikds, meaning coloured ; so that Eyton’s error (if any) appears to have been
merely the omission of the marks of diseresis over the second vowel.—H. 9.
160 MR. H. SAUNDERS ON THE LARINZE. [Feb. 5,
Gavina, Bp. Naum. 1854, p. 212.
For L. canus and allies, and Z. audouini; but in
Gavina, Bp. Consp. Av. ii. p. 222 (1857),
the type and sole representative is L. audouint.
Dominicanus, Bruch, J.f.Orn. 1853, p. 100; id. op. et¢. 1855, p. 280.
For the large dark-mantled Gulls, including the author’s idea of
what L. cachinnans of Pallas should be.
“‘ Leucopheus, Bp.,’’ Bruch, J. f. Orn. 1853, p. 108 (descrip-
tion).
Type and sole representative, L. scoresbii; but
Leucopheus, Bonap. Naumannia, 1854, p. 211,
also includes Z. heermanni; and in
Leucopheus, Bp. Consp. Av. ii. p. 231 (1857),
are substituted for the latter L. fuliginosus and L. belcheri.
‘* Blasipus, Bp.,”’ Bruch, J. f. Orn. 1853, p. 108 (description).
Type and sole representative, LZ. modestus, Tsch.; but
Blasipus, Bonap. Naumannia, 1854, p. 211,
also includes ZL. crassirostris, Vieill., and, in the Consp. Av., further
includes LZ. heermannt.
‘‘Adelarus, Bp.,” Bruch, J. f. Orn. 1853, p. 106, “Hdelméwen.”
For those species which have both a dark mantle and a hood.
Gelastes, Bonap. Naumannia, 1854, p. 212 (deser. nulla).
For L. gelastes and the small unhooded southern Gulls.
Atricilla, Bonap. Naumannia, 1854, p. 212.
For L. atricilla and the three other pretended species evolved
from it.
Creagrus, Bonap. Naumannia, 1854, p. 213 (descript. nulla) ;
Bruch, J. f. Orn. 1855, p. 292 (descr.).
Type, L. furcatus.
Subgen. Melagavia :
Gavia, Br. Gavia ae migra
Cirrhocephala 94, pp. , .
For certain hooded Gulls.
Cirrhocephalus, Bruch,
For a species which had a grey hood !
Bruchigavia, Bp. Consp. Av. ii. p. 228.
A genus playfully made, because Bruch’s Gavie were not the
1878.] MR. H. SAUNDERS ON THE LARIN&. 161
same as the author’s Gavie! Its only claim to remembrance is its
adoption by Mr. W. L. Buller as a genus for a New-Zealand species.
Procellarus et Epitelarus, Bonap. Naumannia, 1854, pp. 211,
213.
Genus defined. Type and sole representative, P. neglectus, which
is an immature L. scoresbii. This species the author had already
located in the genus Leucopheus.
Clupeilarus, Bonap. Consp. Av. ii. p. 220 (1857).
For L. fuscus, cachinnans, and verreauxti. This genus has not
even the merit of consistency ; for it contains such different species
as above, whilst it omits Z. dominicanus (of which L. verreauwi is
only the African form) and L. marinus.
Of the rejected genera one of the best is Gabianus, Bp., of which
the sole representative, L. pacificus, has a remarkably deep, strong
bill. But it differs in no other structural point from other typical
species of Larus, whilst even in the form of the bill it is at times
closely approached by old males of LZ. dominicanus ; so that I think its
adoption would be inexpedient. Lewcopheus, Bp., has been confused
between the author and Bruch until it includes species which Bona-
parte himself has almost simultaneously located in two other genera ;
and I can see no structural difference sufficiently marked to make it
desirable to employ either it or Blasipus, which, according to Bona-
parte’s latest view, includes two species differing considerably in the
form of the bill. Adelarus, Bp., appears to be the result of an attempt
to Latinize the compound word “ Edelméwen,” and should rank with
his Bruchigavia and kindred genera.
The arrangement of the species of Larus is matter of considerable
difficulty. The plan adopted by Schlegel of dividing the Gulls into
Lari marini, for unhooded species, and Lari cucullati, for those which
at one time or another bear a hood, will not stand the test of later
experience,—almost all of those which have a hood in their immature
stage being emphatically Sea-gulls, as are also a few of those which
have a hood in the breeding-season; whilst at least two of the un-
hooded species are partial to inland waters, and present, in conse-
quence, the slight modifications of form shown by many of the
hooded marsh-breeding Gulls. Under these circumstances any
ascending or descending arrangement must necessarily be artificial ;
but I have endeavoured to group the species in the most natural
manner which seemed to me to be practicable.
It may be as well to observe that by an “adult” bird I mean one
which has lost the mottlings, barred tail, and other signs of imma-
turity ; but an “old”? bird is often subject to important alterations
in the coloration or “pattern” of the webs of the primaries, although
the general plumage may undergo no material change. The distine-
tion between the age (in years) of the individual and the age (in
months) of the primary and other feathers should also be held in
Proc. Zoou. Soc.—1878, No. XI. 11
162 MR. H. SAUNDERS ON THE LARIN&E. [Feb. 5,
mind, to prevent confusion, although I have endeavoured to avoid
any ambiguity on this point.
Genus Pacopuina.
The short stout bill, coarse rough feet with serrated membranes,
much excised webs, and strong curved claws appear to entitle this
species to generic separation. The hallux is connected on the inside
of the foot by a serrated membrane with the imner toe, a peculiarity
which I do not recollect seeing noticed elsewhere. The name has
been in use for nearly half a century, and is of general acceptance.
]. PaGorpHILA EBURNEA (Phipps).
Larus eburneus, Phipps, Voy. N. Pole, App. p. 187 (1774) ; Gm.
Syst. Nat. i. p. 596 (1788); Scoresby, Arct. Voy. i. p. 535 (1820)
(Spitzbergen) ; Schl. Mus. P.-Bas, Lavi, p. 6 (1863).
Larus candidus, Miller, Prodromus, p. viii. (1776); O. Fabr.
Faun. Green. p. 103 (1780).
Larus niveus, Bodd. T. des Pl. Enl. p. 58, no. 994 (1783) (nee
Pallas).
Larus albus, Schiff. Mus. Orn. p. 65, tab. 42 (1789).
Gavia eburneus, Boie, Isis, 1822, p. 563; Brehm, Vog. Deutsch.
p- 765 (1831).
Pagophila eburnea, Kaup, Nat. Syst. eur. Th. pp. 69, 196 (1829);
Gray, Gen. Birds, iii. p. 655 (1849); Newton, Ibis, 1865, p. 507
(Spitzbergen, breeding); P. Wright, Ibis, 1866, p. 217 (Polynia
Island, breeding); Dresser, B. of Eur. pts. lix. lx. May 1877.
Gavia nivea, Brehm, Vog. Deutsch. p. 766 (1831).
Cetosparactes eburneus, Macgill. Man. Brit. Orn. pt. ii. p. 252
(1842); Brit. Birds, vol. v.
Larus brachytarsus, Holb. Fn. Greenl. p. 52 (1846).
Pagophila eburneus, Bruch, J. f. Orn. 1853, p. 106.
Pagophila brachytarsa, Bruch, J. f. Orn. 1853, p. 106; Lawr. B.
N. Am. p. 856 (1858); vide Reinht. Ibis, 1861, p. 18.
Pagophila nivea, Bp. Compt. Rend. xlii. p. 772 (1856); Consp.
Ay. ii. p. 230 (1857).
Larus (Pagophila) eburnea, Coues, B. N.W. Am. p. 648 (1874).
Hab. Arctic regions, from Novaya Zemlya to Baffin’s Bay, and
the eastern portion of Arctic America, but not as yet found in the
North Pacific. Straggles down the western coast of Europe and
Eastern America in winter. I can see no reason for considering
Holboll’s L. drachytarsus to be a distinct species.
Genus Rissa.
The principal characteristic assigned to this genus by Leach is
the rudimentary character, or absence, of the hind toe. As this
is not always constant, and as certain rare individuals from
the North Pacific out of many hundreds, are occasionally to be
found with a visible hind claw, and even with a nail, it has been
urged by some systematists that it is not a valid genus. The
1878. ] MR. H. SAUNDERS ON THE LARIN&. 163
strongest evidence against it is that brought by Dr. Coues in his
‘Birds of the North-West,’ p. 646, where he says that whereas ‘a
part of the Kittiwakes from the North Pacific are not distinguishable
tn any way from the North-Atlantic bird, others have the hind toe as
perfectly formed and proportionately as large as in any species of
Larus! And there is a gradation between them.” He goes on to
cite an extreme example from Plover Bay, with a hallux, including
the nail -2 in., with a perfect claw. Whilst writing this paper I have
received, through the kindness of the authorities of the Smithsonian
Institution at Washington, a similar extreme form with a hallux and
nail -2 in., the claw being well formed and curved, although the whole
is certainly but small for the size of the bird, as may be seen by a com-
parison with a species of about the same size, Z. canus, in which the
hallux and nail measure ‘5 in. Mr. O. Salvin has also lent me two
specimens, both, as well as my own, from Alaska, in the one of
which the nail is somewhat less developed, whilst in the other it
is absent, as in the Atlantic bird. It is probable that this ex-
treme form is both rare and local; at least I have never been able to
find any but these two examples amongst the many Kittiwakes,
Pacific and others, which I have examined. However, there it is; and
if the genus Rissa depended solely upon the absence of the hind toe,
it would have to be given up. There are, however, other structural
characteristics, which, when united, seem to me to have weight. The
tarsus is remarkably short, being only 12 against 1-9 in. in length
of middle toe and claw, proportions unknown in any other group of
Gulls ; the shape of the curved bill is also peculiar ; the fail is visibly
although not deeply, forked ; whilst these structural differences are
supplemented by such minor characteristics as the peculiar livery of
the immature bird, totally unlike that of the adult, or of the young
of any other species, and by its exclusively crag-nesting habits. Bear-
ing all these points in mind, I think that, although it is no longer
absolutely correct to say that the genus Rissa depends upon the
absence of the hallux, yet it is advisable to retain it as, on the whole,
a valid natural division, sanctioned, moreover, by general use during
upwards of half a century.
2. Rissa TRIDACTYLA (Linn.).
Larus rissa, Linn. Syst. Nat. i. p. 224 (1766); Phipps, Voy.
N. Pole, App. p. 187 (1774); Leach, Syst. Cat. Brit. Mus. p. 40
(1816); Scoresby, Arct. Voy. i. p. 534 (1820).
Larus tridactylus, Linn. Syst. Nat. i. p. 224 (1766); O. Fabr.
F. Greenl. p. 98 (1780) ; Gm. Syst. Nat. i. p. 595 (1788); Schl.
M. P.-Bas, Lari, p. 31 (1863) ; Godman, Ibis, 1872, p- 222 (Ca-
naries),
Larus albus, P. L. 8. Miiller, Natursystem, p. 108 (1776) (based
on Buffon’s Mouette cendrée tachetée).
“Larus cinerarius, Linn.” O. Fabr. F. Greeal. p. 101 (1780), nee
Linn. (winter-plumage, from description).
Larus riga (mispr.), Gm. Syst. Nat. i. p. 594 (1788).
Larus nevius, Schiff. Mus. Orn, p. 64 (1789).
ii
164 MR. H. SAUNDERS ON THE LARINZ. [Feb. 5,
Larus torquatus et Larus gavia, Pallas, Zoogr. Rosso-As. ii. pp.
328, 329 (1811).
Gavia tridactylus, Boie, Isis, 1822, p. 563.
Rissa brunnichii, Stephens, ex Leach, in Shaw’s Gen. Zool. xiit.
pt. i. p. 181, pl. 21 (1825), type of genus Rissa.
Cheimonea tridactylus, Kaup, Nat. Syst. eur. Th. pp. 84, 196
(1829), type of genus Cheimonea.
Laroides tridactylus, Brehm, Vog. Deutschl. p. 754 (1831).
Laroides rissa, id., op. cit. p. 755 (1831).
Laroides minor, id., op. cit. p. 756 (1831).
Rissa cinerea, Eyton, Cat. Brit. B. p. 52 (1836).
Rissa tridactyla, Macgill. Man. Brit. B.ii. p. 249 (1842) ; Hist.
Brit. B. v. p. 515; Gray, Gen. Birds, iii. p. 655 (1849); Bruch, J.
f. Orn. p. 103 (1853) ; Bp. Consp. Av. il. p. 225 (1857).
“Rissa brachyrhynchus (Gould),”’ Bruch, J. f. Orn. 1853, p. 1038,
nec Gould.
Rissa borealis et gregaria, Brehm, Naum. 1855, p. 294.
* Rissa niveus (Pall.),’’ Bruch, J. f. Orn. 1855, p. 285, nec Pallas.
Rissa kotzebui, Bp. Consp. Av. ii. p. 226 (1857).
Larus (Rissa) tridactylus, Coues, B. N.W. Am. p. 644 (1874).
Larus tridactylus, var. kotzebui (Bp.), ibid. p. 646 (1874).
Hab. Arctic region, and along the sea-coasts of the subarctic
region, down to about 40° N. lat., breeding perhaps even in the
Canaries (Godman) ; in winter it is abundant about the Azores,
Canaries, and opposite coast of Africa. In America it is found on
both Atlantic and Pacific coasts, but does not seem to extend far
down the latter, nor to Japan or China even in winter.
In treating of the genus, I have already noticed that examples
are occasionally found about Alaska and the Aleutian Islands with
a minute but tolerably developed hind toe, and, at times, with a
visible nail, a variation which is rare and not always equal in extent,
even on both feet of the same individual. I have therefore treated
var. kotzebui as a synonym, there being no other difference between
this and the ordinary Kittiwake, and a gradation between them
existing.
3. Rissa BREvIROSTRIS, Brandt.
Larus brachyrhynchus, Gould, P.Z.S. 1843, p. 106; Zool. Voy.
of “Sulphur,” p. 50, pl. 34 (1844), nee Richardson (1831).
“ Rissa nivea (Pall.),’’ Gray, Gen. Birds, iii. p. 655 (1849), nec
Pallas; Lawr. B. N. Am. p. 855; Elliot, B. N. Am. pl. 54.
“Rissa brevirostris, Brandt,” Bruch, J. f. Orn. 1853, p. 103;
id. 1855, p. 285; Dall and Bann. Tr. Chic. Ac. i. 1869, p. 305.
“Gavina citrirostris, Schimper,” Bruch, J. f. Orn. 1855, p. 284
(Kamtschatka).
Rissa brachyrhyncha, Bp. Consp. Av. ii. p. 226 (1857); Coues,
P. Ac. N. Se. Philad. 1862, p. 306.
Larus warnecki, Coinde, Rey. et M. Zool. 1860, p. 401 (Aleutian
Islands).
1878. | MR. H. SAUNDERS ON THE LARIN&. 165
Larus brevirostris, Coues, Key Am. B. p. 315 (1872); Elliot’s
Prybilov Is. Birds, no. 553.
Larus (Rissa) brevirostris, Brandt ; Coues, B. N.W. Am. p. 646
(1874).
Hab. The North Pacific, between Alaska and Kamtschatka,
where it is very abundant, breeding in thousands on the Prybilov
Islands.
This is a very distinct species, and may at once be recognized by
its very short stout bill, orange-red legs and feet, and dark mantle ;
the ground-colour of the primaries is also dark grey. I am indebted
to the liberality of the authorities of the Smithsonian Institution for
a fine specimen of this species, which is as yet rare in collections.
A variation in the hind toe and nail, similar to that in R. tridac-
fyla, although in a smaller degree, is observable in this species. My
own specimen has no claw on the right hind toe, and only a minute
black speck on the left ; of two others, from Alaska (e Mus. Salvin
& Godman), the one has no hind nail whatever, whilst the other has
small black nails, unequal in size, on both hind toes.
Genus Larus.
4, Larus Guaucus, Fabr.
Larus glaucus, O. Fabricius, Faun. Greenl. p. 100 (1780, ex
Briinn.); Gm. Syst. Nat. i. p. 600 (1788) ; Scoresby, Arct. Voy. 1.
p. 535 (1820); Middendorff, Sib. Reise, il. p. 241 (1853); Newton,
This, 1865, p, 509 ; Schlegel, Mus. P.-Bas, Lari, p. 4 (1863) ; Coues,
B. N.W. Am. p. 620(1874) ; Swinhoe, Ibis, 1874, p. 165 (Japan) ;
Seebohm & H. Brown, Ibis, 1876, p. 453 (Lower Petchora) ; Dresser,
B. of Europe, pts. lix., lx. (1877).
“Larus giganteus, Temm,”’ Benicke, Ann. Wetterau. Gesellsch.
il. p. 140 (1814).
Larus leuceretes, Schleep, N. Ann. Wetterau. G.i. p. 314 (i819).
Larus consul, Boie, Wiedemann’s Zool. Mag. p. 126.
Larus islandicus, Edmonst. Mem. Wern. Soe. iv. (1822) p. 185
(nee Edmonst. op. cit. p. 506=L. leucopterus).
Larus glacialis, Macgill. Mem. Wern. Soc. v. pl. i. p. 270 (1824).
Leucus glaucus, Kaup, Natiirl. Syst. p. 86 (1829).
“ Larus glacialis, Benicke,” Brehm; Vig. Deutschl. p. 732 (1831).
Larus hutchinsii, Richards. F. Bor.-Am. ii. p. 419, note (1831) ;
Cassin, P. Philad. Ac. 1862, p. 290; Coues, ibid. p. 294; Elliot,
B.N. Am. ii. pl. 53; Dall. & Bann. Tr. Chic. Ac. p. 304 (1869).
Plautus glaucus, Reich. Nat. Syst. Av. Longip. p. 5 (1852); Ie.
Ay. pl 47. fig. 316-318, pl. 50. fig. 2640.
Glaucus consul (Boie), Bruch, J. f. Orn. 1855, p. 101.
Laroides glaucus, Bruch, J. f. Orn. 1855, p. 281.
Hab. Arctic regions; seldom breeding much to the south of the
arctic circle. In winter it goes southwards, and has been known to
straggle as far as the Mediterranean, to Long Island on the Atlantic
coast of America, also on to the coasts of Japan, whence I have seen
specimens obtained at Hakodadi by Capt. Blakiston.
166 MR. H. SAUNDERS ON THE LARINZE. [Feb.5,~ -
L. hutchinsii 1 consider to be an immature L. glaucus in the stage
where the mottled brown of immature plumage has passed away, and
the pearl-grey mantle has not yet begun to show. This stage lasts
but a short time, which will account for the fact that this supposed
species has so rarely been obtained; but I have always observed in
young specimens in captivity that at this stage they are nearly, and
sometimes quite, white. A fine example in this state, obtained off
Japan by Capt. St. John, H.M.S. ‘Sylvia,’ is in the Marquis of
Tweeddale’s collection. Mr. Collett obtained one in Norway in
September 1871; and several have been recorded from America.
The feet and legs in adults of this Gull are bright flesh-pink, and
not lemon-yellow as depicted by an extraordinary freak of the
colourist in Mr. Dresser’s ‘ Birds of Europe.’
5. LaARuS LEUCOPTERUS, Faber.
Larus argentatus, E. Sabine, Tr. Linn. Soc. xii. p. 546 (nee
auctt.).
Larus leucopterus, Faber, Prod. Isl. Orn. p. 91 (1822) ; Sw. &
Rich. F. Bor.-Am. ii. p. 418 (1831) ; Schl. Mus. P.-Bas, Lari, p. 5
(1863); Dall. & Bann. Tr. Chic. Ac. i. 1869, p. 304 (Alaska and
Lower Yukon); Coues, B. N.W. Am. p. 622 (1874); Dresser, B.
of Europe, pt. xlix (1876).
“ Larus glaucoides, Temm.’’ Meyer, Taschenb. iii. p. 197 (1822);
Boie, Isis, 1822, p. 562.
Larus glaucoides, Temm. Pl. Col. 77° live. Introd. Larus (1828).
Larus islandicus, Edmonst. Mem. Wern. Soe. iv. p. 506 (1823)
(nec Edmonst. op. cit. p. 185).
Larus arcticus, Macgill. Mem. Werner. Soc. v. no. xii. p. 268
(1824) (large specimen).
Larus minor, Brehm, Vog. Deutschl. p. 736 (1831).
Laroides glaucoides, Brehm, op. cit. p. 744.
Laroides leucopterus, Brehm, op. cit. p. 745; Bruch, J. f. Orn.
1855, p. 281.
Laroides subleucopterus, Brehm, op. cit. p. 746.
Glaucus leucopterus, Bruch, J. f. Orn. 1853, p. 101.
Glaucus glacialis, Bruch, op. cit. p. 101.
Larus chalcopterus, Licht. Nomencl. Av. Mus. Berol. p. 99
(1854), sine descr. (type examined, H.%.).
? Laroides chalcopterus (Licht.), Bruch, J. f. Orn. 1855, p.22.
Laroides glacialis, Bruch, op. cit. 1855, p. 282.
Leucus chalcopterus, Bp. Consp. Av. ii. p. 216 (1857).
Leucus arcticus, Bp. op. eit. p. 216.
Leucus leucopterus, Bp. op. cit. p. 217.
Hab. Even more thoroughly arctic, during the breeding-season,
than L. glaucus; straggling southward in winter as far as the coast
of France. It is not even authenticated as breeding in Iceland or
Spitzbergen ; but it does so within the arctic circle from Greenland
to Behring’s Straits, It also breeds in Alaska; and I have examined
a specimen which was obtained in Japan by Capt. Blakiston.
1878.] ~ MR. H. SAUNDERS ON THE LARINA. 167
6. Larus GLaucescens, Licht.
Glaucus glaucescens (Licht.), Bruch, J. f. Orn. 1853, p. 101 (type
examined, H. S.).
“Glaucus glaucopterus, Kittlitz,’ Bruch, J. f. Orn. 1853, p.
101.
Larus glaucopterus, Licht. Nomencl. Av. Mus. Berol. p. 99, sine
descr. (1854), Behring’s Straits, Chamisso (type examined, H. 8.).
Laroides glaucescens, Bruch, J. f. Orn. 1855, p- 281.
Leucus glaucescens, Bp. Compt. Rend. xlii. (1856) p. 770 ; Consp.
Av. il. p. 216 (1857).
Larus glaucescens, Licht.; Lawr. B. N. Am. p. 842 (1858); Coues,
Proce. Philad. Ac. 1862, p. 295; id. B. N.W. Am. p- 622; Swinhoe,
Ibis, 1874, p. 165 (Japan).
Larus chalcopterus, Lawr. B. of N. Am. p- 843, 1860; Coues, Proe.
Philad. Ac. 1862, p. 295 (nec Licht. ).
Hab. Pacific coast of North America up to Behring’s Straits, and,
on the Asiatic side, Kamtschatka, and as far south as Hakodadi,
Japan, whence 1 have seen a specimen obtained by Capt. Blakiston.
This apparent link between the large Gulls with white primaries
and those with barred primaries may be roughly described as a
Herring-Gull with the black portion of the primary-pattern nearly
washed out. It is quite unmistakable. The changes of plumage in
its progress to maturity show, however, that its relationship to L.
glaucus is closer than to L. argentatus.
7. Larus ARGENTATUs, Gm.
Larus fuscus, Penn. Brit. Zool. ii. p. 131 (1768), nee Linn. ; Mont.
Orn. Dict. i. (1802).
Larus argentatus, Gm. Syst. Nat. i. p. 600 (1788), ex Briinn. ;
Schl. M. P.-Bas, Lari, p. 16 (includes allies) ; Gundlach, J. f. Orn.
1857, p. 236 (Cuba) ; B. du Bocage, Jorn. Soe. Lisb. 1868, pp. 149,
330 (Angola); Hartlaub, Syst. Orn. W.-Afrik. p. 251 (1857),
Senegal; Dresser, B. of Europe, pt. xxii. (Oct. 1873); Coues, B.
N.W. Am. p. 625, 1874.
Larus marinus, var. 3, Latham, Ind. Orn. ii. p. p. 814 (1790).
Larus glaucus, Retzius, F. Suec, i. p. 156 (1800), nee Briinn. ;
Meyer & W. Taschenb. ii. p. 471 (1810).
Larus cinereus, Leach, Syst. Cat. Brit. Mus. p. 40. (1816).
Larus argentatoides, Brehm, Beitr. Vigelkunde, iii. pp. 791, 799
(1822).
Larus argenteus, Macgill. Mem. Wern. Soc. vy. p. 264 (1824).
“Larus argentatoides (Bonap.),’ Sw. & Richards. F. Bor.-Am.
Birds, p. 417 (1831), nec Brehm, nee Bonap. ;
Laroides major, argentatus, argenteus, urgentatoides, et argenta-
ceus, Brehm, Vog. Deuischl. pp. 738-743 (1831).
Glaucus argentatus, Bruch, J. f. Orn. 1853, p. 101.
Glaucus aryentatoides, Bruch, op. cit. p. 101.
Larus marinus, Gundl. J. F. Orn. 1857, p- 236; Lembeye, Aves,
de Cuba, p. 122 (1850), cf Gundl. J. f. Orn. 1871, p. 291.
168 MR. H. SAUNDERS ON THE LARINZ. [Feb. 5,
Larus smithsonianus, Coues, Pr. Ac. N: Se. Philad. 1862, p. 296
(North America).
In this species the amount of white on the primaries increases with
the age of the individual. Mr. Dresser (B. of Europe, xxii. L. ar-
gentatus, p. 3) describes an adult male from the Orkneys in summer,
in my collection, as having ‘the outermost primary almost entirely
blackish, white towards the tip, and crossed by a subapical black
band; the next two grey at the base, black towards the tip, being
finally terminated by a large white spot.” This is correct, so far
as any breeding bird with unspotted pearl-grey mantle may be termed
adult; but the example in question is far from being an old bird.
Bearing in mind that the extreme white tip diminishes by abrasion
with the age of the feather, the following are the patterns of the
outer primaries with the increasing age of the bird :—QOn the outer
primary the white spot, or “ mirror,” absorbs the black bar till the
latter wholly disappears, leaving the primary pure white from the
tip to more than two inches upwards; whilst from above, a grey
“* wedge’ along the inner web gradually eats into the black portion,
reducing the width of the black along the inner web to only two
inches. In the second primary a white “mirror”? appears, which
also increases with the age of the bird; but in this feather, so far
as I have yet seen, it does not wholly absorb the black bar and
unite with the white tip ; what it does, however, is to eat round the
black above it, so as to cut off the black from the inner web, and
thus unites with the grey wedge, which has been gradually increasing
its dimensions downwards. _ It is needless to give a minute description
of the remaining primaries; it will suffice to say that, as a rule, the
encroachment of the light portions upon the dark ones increases with
the age of the bird, and there may easily be stages of further pro-
gression with which I am not yet acquainted. This grey ‘‘ wedge”
on the upper portions of the primaries should be borne in mind, as
it is an important distinction between some closely allied species.
These observations equally apply to the Yellow-legged Herring-Gull
(L. eachinnans) and to the American bird which Dr. Coues formerly
distinguished as L. smithsonianus, Dr. Coues, although he has
given it up as a species, even now maintaias (B. of N.W. Am. p. 628)
that if a subapical spot (or ‘‘ mirror’’) is present on the second pri-
mary of the American bird, it is small; but in two examples before
me, of the correctness of whose locality I am well assured, the one
from Grand Manan, in June, has it well developed, whilst in another,
from Long Island, the mirror extends right across the feather, and
on the inner web has nearly eaten through the black and effected a
junction with the grey wedge above. Indeed only one European
bird in my collection has the mirror still more developed. The
average of American may possibly be a little larger than the average
Old-World specimens ; but I have not examined a sufficient series of
the former to speak with the same confidence upon this point that Dr.
Coues does ; at any rate that difference is admittedly unworthy of
specific distinction. The mané/e in the true adult L. argentatus is,
as every one knows, pearl-grey, the legs and feet being flesh-
coloured ; and the ring outside the eye is of a pale yellow. Taking
1878. ] MR. H. SAUNDERS ON THE LARINA. 169
these as the characteristics to distinguish it from its congeners, its
range may be defined as the north-west of Europe from the Varanger
Fiord, the Baltic, the western coasts down to North Africa, the
Azores (where it breeds), Madeira, and the Canaries (Godman). To
Greenland it is a very rare straggler; but it was obtained at Winter
Islands, near Melville Peninsula, occurs in Hudson’s-Bay territory
as far as the Mackenzie River, and probably reaches right across to
the Pacific coast, where it certainly occurs, a specimen from Kodiak,
collected by Wosnesensky, in the St.-Petersburg Museum, being, as
Mr. Seebohm informs me, much lighter in the mantle than Z. ca-
‘chinnans ; and several specimens from the west coast of Mexico are in
my collection. Dall and Bannister also record what seems to be this
species from Alaska and from the Upper Yukon. There can be no
doubt that examples from northern latitudes have a somewhat lighter
mantle than those from more temperate regions, although the trans-
ition is very gradual; and this light form has received the name of
L. argentatoides. From Labrador this species ranges down the coast
and along the great rivers and inland lakes as far as Texas; it also
visits Cuba and Bermudas. Prof. Barboza du Bocage, in his “ List
of Birds in the Lisbon Museum” (J. f. Orn. 1876, p. 291),
cites an example obtained on the Angola coast, and another at Porto
Alexandre, Benguela, more than 15° south of the equator, and in
the latitude of the island of St. Helena. ‘lhis is indeed an exten-
sion of its range, provided there is no error in the identification of
the specimens.
8. Larus CACHINNANS, Pall.
Larus cachinnans, Pallas, Zoogr. Ross.-As. ii. p. 318 (1811).
Larus argentatus, Bp. tconogr. F. Ital. Uccelli, Introd. (1832-41) ;
Middend. Sib. Reise ii. p. 242 (part.), (1853); Schlegel, Mus.
Pays-Bas, Lari, p. 17 (part.), 1863 ; Blasius, J. f. Orn. 1865, p. 380
(part.) ; Hume, Yarkand Exp. Zool. p. 299 (1873), Kashmir; id.
Stray Feath. i. p. 270 (1873), Scinde.
Larus cachinnans, Pall.; v. der Mihle, Orn. Griechenl. p. 143
(1842).
Glaucus leucopheus, (Licht.) Bruch, J. f. Orn. 1853, p. 101 (Red
Sea).
Glaucus michahellesit, Bruch, tom. cit. p. 101.
“ Glaucus borealis, Brandt,” Bruch, tom. cit. p. 101.
Laroides michahellesti, Bruch, op. cit. 1855, p. 282.
Laroides cachinnans, (Pall.), Bruch, tom. cit. p. 282.
“ Laroides borealis, (Brandt)”’ Bruch, tom. cit. p. 282.
Larus epargyrus, Licht. Nomencl. Av. Mus, Berol. p. 99 (1854),
sine descr. (type examined, H. S.).
Larus leucopheus, Licht. Nomencl. Av. Mus. Berol. p. 99 (1854),
descr. nulla (type examined, H. 8.) ; Salvadori, Cat. Ucc. Sard. p.
129 (1864) ; Finsch & Hartl. Vog. Ost-Afrik. p. 818 (1870);
Dresser, B. of Europe, pt. xxii. (Oct. 1873).
Larovdes leucopheus, Bp. Naumannia, 1854, p. 212; id, Consp. ii.
p. 219 (1857).
170 MR. H. SAUNDERS ON THE LARINE. [ Feb. 5,
Larus fuscescens, Sclater, P. Z. S. 1867, p. 315, et Rev. List of
Vert. (1872), p. 316.
? Larus fuscescens, Licht.; Bruch, J. f. Orn. 1853, p. 100,
(part.). One of the two specimens so labelled in the Berlin
Museum is of this species ; the other is a L. fuscus.
After much consideration and the examination of a very. large
series of specimens from various localities, I have come to the con-
clusion that this form, or species, is sufficiently distinct to be treated
apart from L. argentatus. The distinguishing characteristics of L.
cachinnans are the darker mantle, yellow legs and feet, and the deep
orange-red ring round the outside of the eye. These colours are:
naturally much more apparent in life than in dried skins; but the
colour of the mantle is enough to enable any one with an ordinary
perception of shades to separate the two birds at a glance. In the
pattern of the primaries, and in the individual variations in size both
are alike. With regard to the name which I have adopted, it seems
to me that there cannot be the slightest doubt as to the species
Pallas meant by his Larus cachinnans from the Caspian and the
Steppes: he describes it fully ; and, to avoid any ambiguity as to the
shade of colour of the mantle, he uses precisely the same term that
he does for the mantle of ZL. ichthyaétus, which exactly suits this
species, whilst it is too dark for LZ. argentatus, and too light for L.
affinis. Yet more, my friend Mr. Seebohm, on his return from
Siberia, examined the Larine in the St.-Petersburg Museum; and,
thanks to him, I am able to state from absolute comparison that L.
cachinnans, and L. leucopheus of the Mediterranean are the same,
Pallas’s name having the priority.
It appears, indeed, to be a form which, whether from living in a
more brilliant atmosphere, or from frequenting inland seas as dis-
tinct from great oceans, or from other causes with which we are not
acquainted, has acquired a greater intensity of coloration than its
congener ; but it is not altogether easy to indicate its precise range.
The most northern example that I have examined is from Havre,
an adult male, the oldest, to judge by the primaries, of any grey-
backed gull in my collection ; so that it appears to straggle up the
French coast. It is not, however, till the Mediterranean is reached
that L. cachinnans replaces L. argentatus; thence it ranges through-
out that inland sea, breeding on its shores and islands ; goes up the
Black Sea, across the steppes and the low-lying marshy and salt-
lake districts of Russia from the mouths of the Volga and the shores
of the Caspian, as far as Vologda, across the Ural river and the
Kirgish steppes, to the Irtich and as far as Lake Baikal. The above
seems to be, roughly, its breeding-range ; for Meves’s description of
the ‘‘ Z. eachinnans’’ obtained at Cholmogory on the Dwina applies
better to the next species: it was so dark in the mantle that he at
first took it to be L. fuscus. The species found in Kashmir by the
Yarkand expedition was probably L. cachinnans. It goes down the
Red Sea, and in winter visits the Persian Gulf, and the Mekran coast
as far as Kurrachee. It is also found along the coasts of China and
Japan in winter, and is the species recorded by Swinhoe under the
1878. | MR. H. SAUNDERS ON THE LARIN®. 171
names of LZ. cachinnans and L. occidentalis ; but his birds are most
decidedly not the true L. occidentalis of Audubon, which has never
as yet been obtained anywhere on the Asiatic shores. All Asiatic
birds show very distinctly the grey wedge in the outer primaries,
which L. occidentalis never does: besides, their mantles are not
nearly dark enough for that species. I possess, or have examined,
a large series of specimens from almost all the localities I have
indicated, and have taken very little from descriptions ; indeed it
is necessary to rely in this matter on one’s own observations, as the
confusion respecting this group is inconceivable to any one who has
not studied the question. In Japanese and Chinese specimens, all
obtained in winter, my collection is especially rich, owing to the
efforts of Capt. St. John, of H.M.S. ‘Sylvia,’ and Lieut. Stanley
Muggeridge, of H.M.S. ‘ Kestrel ;’ Professor ‘Taczanowski has sent
me specimens from Lake Baikal, and Mr. W. Muloch and Mr.
Blanford examples from Kurrachee and the Mekran coast, whilst
as regards Russia and the Mediterranean Mr. Seebohm’s and other
collections have been available. I may observe that I think it quite
possible that in individuals from more northern localities the feet may
not be so distinctly yellow as in examples breeding in the Mediter-
ranean, believing, as I do, that certain atmospheric conditions exercise
a considerable effect upon coloration.
9. Larus AFFINIS, Reinh.
Larus afinis, Reinhardt, Vidensk. Meddel. 1853, p. 78, et Ibis,
1861, p. 17 (type examined, H. 8.); Seebohm and H. Brown, Ibis,
1876, p. 452.
Larus argentatus (partim), Midd. Sib. Reise, ii. p. 242. (The
description of the bird from Sea of Okhotsk clearly applies to this
species.)
Larus cachinnans, Licht. Nomencl. Av. Mus. Berol. p. 99, nec
Pallas (specimens examined, H. 8.).
Larus fuscus, Jerdon, B. of India, ii. p.-830 (1864).
Larus cachinnans, Meves, Ofv. k. Vetensk. Ak. Férh. 1871, p. 786
(Cholmogory) ; Heugl. Orn. N.O.-Afr. Bd. 2. Abth. ii. p. 1392 (nee
Pallas).
ian sp.? No. 39, Heuglin (nec Pallas), Ibis, 1872, p. 65
(Novaya Zemlia and Waigats): description can only apply to this
species.
eae occidentalis, Hume, Stray Feath. 1873, p. 273 (nec
Audub.).
Larus heuglini, Bree, B. Eur. 2nd ed. v. p. 58 (1876).
Professor J. Reinhardt, of Copenhagen, was the first to discrimi-
nate and to confer a name upon an individual of this species which
had straggled to Greenland; but for a knowledge of its habitat and
distribution we are indebted to Messrs. Seebohm and Harvie Brown,
who found it breeding upon the Petchora, where, however, it only
comes about 11th May, and retires southward on the approach of
winter. The series of specimens brought home by these energetic
naturalists threw a light upon many points which had hitherto been
obscure, and showed the real position of numerous specimens from
a
172 MR. H. SAUNDERS ON THE LARINE. [Feb. 5,
the Red Sea and the Beloochistan coast, which had formerly been a
great trouble, most of them being in immature plumage, and not
suiting either LZ. cachinnans or L. fuscus.
Mr. Hume’s description of the pattern of the primaries of the birds
found in winter about Kurrachee (Stray Feathers, 1873, p. 273)
shows clearly that his LZ. occidentalis is this species, and by no
means the true American bird, the occurrence of which, as I
have said before, has never yet been authenticated on the coasts of
Asia.
Heuglin’s dark-mantled bird from Novaya Zemlia is clearly L.
affinis ; and Middendorff’s description of a variety of L. argentatus
found round the southern shores of the Sea of Okbotsk also applies
to this species.
It may appear strange at first sight that this species should have
been first described from Greenland; but that is merely due to the
unusually careful attention which the fauna of that country has
received from Dr. Reinhardt, whose watchful eyes not even a straggler
could escape. I have examined the type, and am satisfied that it is
of this species. It connects with L. fuscus rather closely (although
quite distinct) in the length of its foot as compared with that of the
tarsus, it having a proportionally smaller foot than either L. argen-
tatus, L. cachinnans, or L. occidentalis, but larger than L. fuscus.
From the last it may also be distinguished by its larger size and the
distinct “pattern” of the outer primaries, the grey wedge being
quite marked in this species, whilst it is absent in the outer feather
of Z. fuscus. The mirror on the second primary is moreover ouly
to be found in very old birds (not one of thirteen breeding-birds
obtained on the Petchora had it); whilst in old Z. fuscus this
mirror is always present. The present species is in fact a Herring-
Gull which passes the whole of the year in a brilliant atmosphere ;
and I cannot help thinking that to this, and to other conditions of
existence with which we are as yet unacquainted, its intensity of
coloration is mainly attributable.
10. Larus occrpentatis, Audubon.
Larus occidentalis, Aud, Orn. Biogr. v. p. 320 (1839) ; Lawr. B.
of N. Am. p. 845 (1858); Elliot, B. N. Am. il. pl. lii.; Coues,
P. Ac. N. 8. Philad. 1862, p. 296; Schl. M. P.-Bas, Lari, p. 15
(1863).
Giaucus occidentalis, Bruch, J. f. Orn. 1853, p. 101.
Laroides occidentalis, id. op. cit, 1855, p. 282; Bp. Consp. Av. ii.
p- 219 (1857).
Larus argentatus, var. occidentalis, Coues, Key N. Am. Birds,
p- 312 (1872); id. B. of N.W. Am. p. 633 (1874).
Larus fuscus?, Saund. P. Z.S. 1875, p. 158 (Lower California).
Hab. Pacific coast of North America down to Magdalena Bay,
Lower California.
It seems to me that this is a very recognizable form, and fully
deserving of consideration as a species. Its nearest ally is, on the
whole, L. afinis; but in the wing-pattern the grey wedge is absent in
1878. | MR. H. SAUNDERS ON THE LARIN#. 173
the two outer primaries; and their ground-colour is rather darker
than in most examples of L. fuscus. In the colour of the mantle
many specimens are quite as dark as L. fuseus, especially those
from Southern California, one of which I should have referred to
that species but for its long coarse foot ; for at that time I had only
northern specimens of L. occidentalis available, and these are con-
siderably lighter on the mantle and in general tint. Since then [
have had the opportunity of inspecting more examples and of knowing
the species better ; and it seems to me that its large deep bill will
generally, and its large coarse foot, longer than the tarsus, will
always, suffice to separate it from L. Juscus, which has a delicate
foot, much shorter than the tarsus. ‘As a rule L. occidentalis is a
stouter bird; but some males of L. fuscus from the south of Europe
run very long in the wing and as large in the bill, whilst on the other
hand the Magdalena-Bay ZL. occidentalis is a female, and has an
unusually slender bill. In one example I have found a tiny sub-
apical spot on the second primary on one side, but not on the other ;
so that it is probably a mark ‘of extreme age; but in fully adult
L. fuscus this is common. Although I have laid stress upon the
characters which distinguish this Gull from L. Juscus, yet it is rather
more closely related to the Herring-Gull group, as shown by its
generally larger size, stout bill, and large feet, which are flesh-coloured
in this species, although, as in many other cases, they are sometimes
rather yellow when dried.
11. Larus ruscus, Linn.
Larus fuseus, Linn. Syst. Nat. i. p. 225 (1766); Scop. Ann. i.
Hist. Nat. p. 80 (1769); Gmel. Syst. Nat. i. p. 999 (1788); Lath.
Ind. Orn. p. 815 (1790); Temm. Man. @’Orn. p. 496 (1815); Schl.
M. Pays-Bas, Lari, p. 15 (1863); Finsch & Hiartl. Vog. Ost-Afr.
p- 820 (1870) ; Sharpe and Dresser, B. of Europe, pt. xvi. (Feb-
ruary 1873).
Larus flavipes, Meyer, Tasch. Vig. Deutsch. ii. p. 469, pl. front.
(1810).
Larus cinereus, Leach, Syst. Cat. Mamm. &c., Brit. Mus. p- 401
(1816).
“Larus argentatus, Mont.” Bewick, Brit. B. Supp. p. 39 (1821).
Leucus fuscus, Kaup, Natiirl. Syst. pp. 86 and 196 (1829).
Laroides melanotos, harengorum et fuscus, Brehm, Voég. Deutschl.
pp. 747-749 (1831).
Dominicanus fuscescens (Licht.) partim, Bruch, J. f. Orn. 1853,
p- 100. Lichtenstein’s example from Arabia is this species; the
other bearing this name is L. cachinnans (specimens examined, H. S.).
Dominicanus fuscus (Linn.), Bruch, J. f. Orn. 1853, p. 100; id.
op. cit. 1855, p. 281, including his L. fuscescens of 1853.
Clupeilarus fuscus, Bonap. Consp. Av. ii. p- 220 (1857).
“* Larus medius, Hempr. & Ehr.” in Mus. Berol. “Gumfudde.”
An immature specimen of this species, H. S.
Hab. The north of Europe, the Faroes, the Baltic, Russia as far
east as Archangel, the British Islands, the French coast, and the
174 MR. H. SAUNDERS ON THE LARIN&E. [Feb. 5,
Canaries (probably its south-west breeding-limit, although it is
known to go as far south as Senegal, whence [ have examined spe-
cimens obtained in May). It visits the Portuguese and Spanish coasts,
goes up the Mediterranean to the Black Sea, ascends the Nile to
Nubia, and is found throughout the Nile country, whilst on the Red
Sea, as far as Aden, it is stated to be sedentary (Finsch § Hartlaub).
Jerdon’s solitary immature specimen procured at Jaulna, in the
Deccan, was probably LZ. affinis, the length of the wing (18 to 19
inches) being greater than that of any L. fuscus I ever saw, 1737
inches being the very extreme for an old and fresh-moulted bird;
and, indeed, Mr. Dresser gives only 15:7! Mr. Dresser states that
Dybowski found it in Dauria ; but Prof. Taczanowski, in his “ Faune
de la Sibérie Orientale” (Bull. Soc. Zool. de France, i.) does not
mention it amongst that traveller’s collection; and with regard to
the quotation by the above author of Mr. Swinhoe’s authority for its
occurrence on the coast of China, a reference to the revised “ List
of the Birds of China,” P. Z.S. 1871, p. 421. no. 656, shows that
the species referred to (‘Ibis,’ 1860, p. 68) as L. fuscus was really
L. crassirostris, Vieill. (LZ. melanurus, Temm.), thus considerably
circumscribing the eastern range attributed toit. As regards America,
Dr. Coues considers that there is no good evidence of its occurrence
on the coasts of the United States.
The distinguishing characteristics of the adult of this species are
its dark slate-coloured mantle, chrome-yellow legs and feet, and the
shortness of the foot as compared with the tarsus. The outer pri-
maries are very dark, and may be termed black, with a subapical
patch or mirror on the first; and in old birds there is a small
mirror on the second primary ; but even in these there is xo sign of
the grey wedge on the upper part of the inner web of the outermost,
scarcely so on the second, and but rarely even on the third—a
marked difference from the LZ. argentatus group. In L. fuscus the
inner web merely fades into a lighter shade towards the edge; and
this dark ground of the primaries, and the absence of distinct
‘pattern,’ coupled with its average smaller size, and especially the
smaller foot, will generally be sufficient to distinguish it from either
L. affinis or L. occidentalis. But for the large coarse foot, I should
have assigned a specimen of L. occidentalis, from Magdalena Bay,
California, to this species ; for at the time I had never seen so dark a
form of the American bird. In shade of mantle there is much
variation, some being in this respect quite as light as LZ. afinis, whilst
the blackest are Egyptian specimens, in which the yellow feet .are
also brightest, probably due to climatic influences. ‘These very dark
birds when old have certainly a white subapical spot on the second
primary, although Blasius imagined that this was confined to northern
and lighter-mantled birds. The examples which run closest to L.
affinis are some from Malaga and Tangiers, two of which I have with
the subapical spot on the second primary, showing considerable age ;
but even with these the difference between them and L. affinis is very
marked, L. afinis having the grey wedge so much more defined.
The adults, therefore, are quite distinguishable ; but some years ago,
1878.1 MR. H. SAUNDERS ON THE LARIN&. 175
before Messrs. Seebohm and Harvie Brown’s explorations on the
Petchora had made me acquainted with a series of ZL. affiinis, I was
much puzzled by Mr. W. Blanford’s immature birds from the
Baluchistan coast, obtained in December 1871 to January 1872, and
I led him into the error of ascribing them to L. fuscus. By my
note-book of specimens examined I find that I was in much doubt
even at that time, owing to their size and the relative dimensions of
the feet and tarsi; but according to my lights I could then ascribe
them to nothing else. I now consider them to have been Z.
affinis.
12. Larus cattrornicus, Lawr,
? Larus niveus, Pall. Zoogr. Rosso-As. ii. p: 320, pl. 86 (1811),
(Kamtschatka), nee Bodd. T. Pl. Enl. 994 (1783).
? Laroides americanus, Brehm, Vég. Deutsehl. p- 743 (1831).
Larus californicus, Lawr, Aun. Lye. New York, vi. 1854, p. 79;
id. Birds N. Am. p. 846 (1858); Coues, Pr. Ac. N. S. Phil. 1862,
p- 300; id. B. of N.W. Am. p. 634 (1874).
Laroides californicus, Bp. Consp. Av. ii. p. 220 (1857).
Larus delawarensis, var. caltfornicus, Coues, Key N.-Am. B.
p- 313 (1872). é
Hab. Pacific coast of North America from Vancouver’s Island to
Lower California, and the interior as far as the vicinity of Great
Slave Lake; also Japan coast, whence I have examined an adult
specimen in the Marquis of Tweeddale’s collection, obtained by Capt.
St. John, H.M.S. ‘Sylvia,’ off Kali, Japan, in January 1872.
The name L. niveus, Pallas, is not available, having been previously
employed by Boddaert for Pagophila eburnea; but I have long been
of opinion that the bird described by Pallas under that name was
really this species—a conviction strengthened by the sight of an un-
doubted example from J apan, proving that it dues cross the North
Pacific. It has generally been supposed to apply to a larger race of
L. canus ; but Pallas knew L. canus perfectly well, and he describes
his Z. niveus as a somewhat scarce visitor to the northern and
Kamtschatkan seas, and not in the habit of going far up the rivers,
also as “‘magnitudo corvi coracis.”? Now the largest ZL. canus could
hardly be described as of the size of a Raven ; and, except in the wing,
which is proportionally long in that Gull, the other measurements
are too large for it; but they exactly suit ZL. californicus, and
Pallas’s figure is a perfect portrait of a specimen recently sent to me
through the kindness of the authorities of the Smithsonian Institu-
tion. Perhaps these remarks may assist in laying the ghost of that
Larus niveus which for nearly a century has been haunting the lists
of systematists.
Dr. Coues inserts with a ? DL. argentatordes, Bonap. Synopsis,
Ann. Lye. Nat. Hist. New York, p- 360 (1828), as the earliest name
for this species, whilst disavowing any desire to supersede Lawrence’s
title; but as the name had already been applied by Brehm in 1822
for L. argentatus, it could not be used for this species, even if correct.
But Bonaparte says of his species :—‘ Common near New York and
176 MR. H. SAUNDERS ON THE LARIN&, [ Feb. 5,
Philadelphia ; we have also shot it on the southern coasts of Eng-
land.” Now I am not aware of this species having been found within
2000 miles of New York ; and the description and measurement (20
inches in length) suit L. delawarensis, the length of which Dr. Coues
gives as 19°75 inches, better than any other. Dr. Coues goes on
to identify L. argentatoides of Richardson with Bonaparte’s species :
but this cannot be ; for Richardson’s birds were from 23 to 25 inches
in length, with a mantle of the same shade as the Iceland Gull—both
too large and too light for Z. ealifornicus. Richardson got his bird at
Melville Peninsula, and speaks of it as found at Hudson’s Bay. Dr.
Coues says the Smithsonian Institution possesses specimens ‘ from lo-
calities not far distant from those of Richardson ;” but it seems to me
that Great Slave Lake is a very considerable distance from Melville
Peninsula, although nearer to it than to New York. Dr. Coues gives
the length of L. ealifornicus as 20 inches; and as Richardson’s 23
to 25 inches do not suit him, he quite gratuitously suggests that
Richardson drew up his measurements from the largest specimens ;
whilst as for the colour of the legs, which are described as “ flesh-
coloured,” whereas in L. californicus they are olivaceous, his assump-
tion is that Richardson described them from dried skins! There can
be very little doubt that Richardson’s birds were examples of L. ar-
gentatus; for his measurements and descriptions suit that species
better than any other.
I am glad to see that Dr. Coues, in his ‘Birds of North-West
America,’ has reconsidered his previous hasty determination that this
species was merely a large variety of L. delawarensis. To judge
from the examples I have examined, L. californicus, although cer-
tainly the connecting-link between the Z. argentatus and the L. canus
group, is perfectly distinct from either. In the pattern of the
primaries it rather goes with L. argentatus, in the colour of the soft
parts with Z. delawarensis ; in the colour of the mantle it is much
darker than either, though not so dark as L. occidentalis.
13. Larus DELAWARENSIS, Ord.
Larus delawarensis, Ord, Guthrie’s Geogr., 2nd Am. ed., i. p. 319
(1815) fide Lawr. B. N. Am. p. 846 (1858); Coues, B. of North-
West Am. p. 636 (1874); Wheeler, Rep. Exp. and Surv. W. of
100th Mer. p. 485 (1876); Reid, Zoologist, 1877, p. 489 (Ber-
mudas).
Larus canus, Bp. Speechio Comp. p. 69 (1827), nec auctt.
? “ Larus argentatoides, Brehm,” Bp. Synopsis, p. 360 (1828),
nec Brehm.
Larus zonorhynchus, Richardson, F. Bor.-Am. ii. p. 421 (1831) ;
Audubon, B. Am. viii. p. 35, pl. 446 (1839) ; Schlegel, Mus. P.-
Bas, Lari, vi. p. 22 (1863); Blasius, J. f. Orn. 1865, p. 380; Bp.
Consp. Av. ii. p. 224 (1857); Gundlach, J. f. Orn. 1857, p. 236
(Cuba).
Glaucus zonorhynchus (Rich.), Bruch, J. f. Orn. 1853, p. 102.
“Glaucus occidentalis (Audub.),” Bruch, J. f. Orn. 1853, p. 101,
taf. ii. fig. 20, nec Audubon. ’
1878. | MR. H. SAUNDERS ON THE LARIN&. 177
Gavina zonorhynchus (Richards.), Bp. Naum. 1854, p. 212;
Bruch, J. f. Orn. 1855, p. 282.
Gavina bruchi, Bp. Naumann. 1854, p. 212; Bruch, J. f. Orn.
1855, p. 283.
“Laroides occidentalis (Audub.),” Bruch, J. f. Orn. 1855, p. 282,
nec Audubon
Larus zonorhynchus, var. mexicanus et var. bruchi, Bp. Consp. Ay.
il. p. 224 (1857).
Hab. Interior and coasts of North America from the Saskatche-
wan and Labrador to Great Salt Lake (breeding), and in winter to
the middle and southern States, Cuba, and the Bermudas. An im-
mature bird in my collection obtained by Mr. H. Whitely at Hako-
dadi, Japan, 14th December, 1864, seems from its size and stout
barred bill to belong to this species.
The adult is easily distinguished from Z. canus by its larger size,
stout, double-zoned bill, and lighter mantle; it is smaller than L.
californicus, its wing-pattern is different, and the mantle is much
lighter.
14. Larus canus, Linn.
Larus canus, Linn. Syst. Nat. i. p. 224 (1766); Gm. Syst. Nat.
1. p. 596 (1788) ; Schl. Mus. P.-Bas. Lavi, p. 23 (1863) ; Sharpe
& Dresser, B. of Eur. pt. xvii. (1873) ; David & Oust. Ois. de la
Chine, p. 517 (1877).
Larus cinereus, Scop. Ann. i. Hist. Nat. p. 80 (1769).
Larus hybernus, Gm. Syst. Nat. i. p. 596 (1788).
Larus procellosus, Bechst. Orn. Tasch. p. 373 (1802).
Larus cyanorhynchus, Meyer, Tasch. Vog. Deutschl. ii. p. 480
(1810).
Laroides procellosus et L. canescens, Brehm, Vog. Deutschl.
pp- 750-753 (1831).
Larus canus, var. major, Middendorff, Sib. Reise, ii. p. 243 (1853).
Larus heineit, Homeyer, Naumannia, 1853, p. 129.
Glaucus canus, Bruch, J. f. Orn. 1853, p. 102.
“Glaucus lacrymosus (Licht.),” Bruch, J. f. Orn. 1853, p. 102,
nee Licht.
Gavina kamtschatchensis, Bp. Naumanuia, 1854, p. 212.
Gavina heinei, Bruch, J. f. Orn. 1855, p. 283.
Gavina canus, id. op. cit. p. 284.
? Rissa nivea, Bp. Cat. Parzudaki, p. 11 (1855).
“Larus niveus, Pall,’ Bp. Consp. Av. ii. p. 224 (1857); Swinhoe,
P.Z.S8. 1871, p. 420; David & Oust. Ois. de la Chine, p. 518,
1877, nec Pallas.
Larus delawarensis, Coues, Pr. Ac. N. Se. Phil. 1861, p. 246.
Larus canus major, Schl. Mus. P.-Bas, Lari, p. 26 (1863).
“Larus suckleyi, Lawr.”’ Schl. M. P.-Bas, Lari, p. 27 (1863), nec
Lawr. (Japan).
Larus audouini, Tristram, Ibis, 1868, p. 330, nee Payr.
Hab. Throughout the Palearctic region, but very rare in Ice-
land ; once in Labrador.
Proc. Zoou. Soc.—1878, No. XII. 12
178 MR. H. SAUNDERS ON THE LARIN&. [ Feb. 5,
A specimen bearing the label of the Labrador Expedition of Dr.
E. Coues and Mr. J. W. Dodge in 1860, obtained at Henley Harbour
on the 2lst August, and marked ZL. delawarensis, came into my
hands indirectly from Mr. Krider, of Philadelphia. I have very little
doubt as to its being really Z. canus; but the American naturalists
will have an opportunity of disputing or confirming my view, as I
have sent it to the Smithsonian Institution.
Few species differ so much in individual size as L. canus; and I
cannot admit the specific validity of the large race found throughout
Northern Russia and Siberia. Off Japan all sizes are found ; and the
colour of the mantle is also very variable, being lightest in Scotch
breeding-birds. It appears to be a species which attains its greatest
development in the north and east, and deteriorates in size as it
ranges south and west. I have already pointed out that I consider
I. niveus of Pallas to be really the earliest name of L. ealifornicus,
Lawr.
15. Larus BRACHYRHYNCHUS, Rich.
Larus canus, Richardson, F. Bor.-Am. ii. p. 420 (1831), nee Linn.
nec auctt. (adult).
Larus brachyrhynchus, Rich. F. Bor.-Am. ii. p. 421 (1831), juv.
(nec Gould, P.Z.S. 1843), type described, Great Bear Lake, May
23rd, 1826; Coues, P. Ac. N.S. Philad. 1862, p. 302; Elliot, B.
N. Am. ii. pl. 53; Dall & Bann. Tr. Ch. Ac. 1869, p. 305.
Larus suckleyi, Lawr. Ann. Lye. New York, 1854, p. 264; id.
B. N. Am. p. 847 (1858) ; Schl. M. P.-Bas, Lari, p. 27 (1863).
Rissa septentrionalis, Lawr. Ann, Lye. New York, 1854, p. 266;
id. B. N. Am. p. 854 (1858),
Larus canus, var. brachyrhynchus, Coues, Key N. Am. B. p. 313
(1872); id. B. N. W. Am. p. 638 (1874).
Hab. North-Pacific coast of America, from Sitka downwards, and
the interior to Great Bear Lake.
I can refer to no other species the specimens obtained by the late
Mr. Hepburn at San Mateo, California, and one very old and fresh-
moulted bird in the Copenhagen Museum from Sitka. They are
certainly not L. delawarensis ; and they are smaller than any L. canus
in my collection except one, a quite abnormally small female from
Orkney. The bill is slender and weak; and the foot with the middle
toe and nail is nearly as long as the tarsus, which in LZ. canus is con-
siderably longer. The bill is olive-green to mandible, in front of
which it is yellow, the former colour being much more predominant
than in old Z. canus, There is much more grey from the base of
the primaries downwards than in LZ. canus ; and on the third primary
the wedge descends to the level of the tip of the fifth primary, whilst
there is a broad subapical mirror on the third primary, which, again,
is seldom, if ever, the case in L. canus; also the ends of the prima-
ries are much more broadly tipped with white.
So far, I think, I am in accord with Dr. Coues, who has had the
advantage of examining Richardson’s type, which, however, is a
young bird ; but as regards the adult, of which he has seen far more
1878. ] MR. H. SAUNDERS ON THE LARIN#E. 179
examples than I have, there is a slight discrepancy to be explained.
He says that the colour of the mantle of L. brachyrhynchus is rather
lighter than in LZ. canus; my specimens are certainly rather darker
than the darkest £Z. canus. Perhaps this is a slip of the pen, or an
inversion of his reference; otherwise I cannot understand it. I
retain this species as it has aiready been described, because the
specimens before me have a general appearance so different from L.
eanus that they are distinguishable at a glance, although it is rather
difficult to define the differences on paper ; but I expect that a larger
series will throw light upon the subject. Schlegel’ bird from
Japan assigned to this species seems to me to be merely a large L.
canus,
16. Larus aupouint, Payr.
? Larus quadricolor, Scop. Ann. i. Hist. Nat. p. 81. no. 109 (1769).
Larus audouini, Payraudeau, Ann. Sc. Nat. viii. p. 462 (1826) ;
Temm. Pl. Col. livr. 81, pl. 480 (1826); Gould, B. Europe, v.
pl. 438 (1837) ; Schl. M. P.-Bas, Lari, p. 22 (1863); Degl. & Gerbe,
Orn. Europ. ii. p. 420 (1867) ; Lilford, Ibis, 1875, p. 31.
Larus payraudei, Vieill. Faun. Frang. Ois. p. 396 (1828), fide
Deg. & Gerbe, loc. cit.
Gavia audouinii (Payr.), Boie, Isis, 1844, p. 191.
Glaucus audouini (Payr.), Bruch, J. f. Orn. 1853, p.. 102.
Gavina audouini (Payr.), Bp. Naum. 1854, p. 212; Consp. Av.
li. p. 222 (1857).
Laroides audouini (Payr.), Bruch, J. f. Orn. 1855, p, 282 ; Brehm,
Naum. 1855, p. 294.
Hab. Mediterranean, especially about Corsica and Sardinia, and
the neighbouring coasts of Italy, the Balearic Islands, and North
Afriea. Mr. Gould quotes Natterer (7m litt.) as having shot three
specimens outside the Straits of Gibraltar, between that place and
Tarifa. Lord Lilford found it breeding on the island of Toro, off
Sardinia. Canon Tristram’s reputed Z. audouini from Palestine are
all Z. canus.
This Gull is one of the most unmistakable species, owing to its
length of wing, dark primaries, lead-coloured or black legs, and, when
adult, its cherry-red double-zoned bill, From the description given
by Scopoli of his Z. guadricolor it is probable that it is this species ;
but in the absence of certainty it is undesirable to use that name,
17. Larus Marinus, Linn.
Larus marinus, Linn. Syst. Nat. i. p. 225 (1766); O. Fabr.
Faun. Greenl. p. 102 (1780); Meyer, Tasch. Vég. Deutschl. pl. ii.
p- 465; Temm. Man. d’Orn. p. 490 (1815); Macgill. M. Wern.
Soc. vol. v. p. 255 (1824); Schlegel, M. Pays-Bas, Lari, p. 10
(1863); Sharpe and Dresser, B. of Eur. pt, xv. (1872); Coues, B.
of North-West (America), p. 624 (1874); Swinhoe, Ibis, 1874,
p- 165 (Japan) ; Reid, Zoologist, 1877, p. 489 (Bermudas),
Larus nevius, Linn. Syst. Nat. p. 225 (1766).
12*
180 MR. H. SAUNDERS ON THE LARIN. [Feb. 5,
Larus maculatus, Bodd. Tab. Pl. Enl. p. 16 (1783).
Larus maximus, Leach, Syst. Cat. Brit. Mus. p. 40 (1816); Brehm,
Vég. Deutschl. p. 728 (1831).
Leucus marinus, Kaup, Natiirl. Syst. pp. 86, 196 (1829).
Larus miilleri, Brehm, op. cit. p. 729 (1831).
Larus fabricii, id. op. cit. p. 730 (1831).
Dominicanus marinus, Bruch, J. f. Orn. 1853, p.100; id. ib, 1855,
p. 280.
Larus albus, P. L. 8. Miller, Natursystem, p. 108 (1776), has
generally been quoted by copyists as a synonym of this species ; but
investigation shows that it.is based upon Buffon’s ‘‘ Mouette cendrée
tachetée”’ (vol. vii. p. 424; Pl. Enl. 387), which represents a young
Rissa tridactyla.
Hab. Northern and temperate Europe and Iceland (breeding) ;
visiting the Mediterranean in winter, as far as Greece; the Canaries,
and probably the Azores. In Northern Greenland Prof. Reinhardt
assures me that it is very rare; it breeds in Labrador, occurs on the
great lakes of North America, and visits Florida in winter. Lem-
beye’s specimen, recorded from Cuba, turned out to be Z. argen-
tatus (vide J. f. Orn. 1871, p. 290); but it has occurred at the Ber-
mudas (Reid). No record from the American side of the Pacific ;
but I have examined undoubted specimens from Japan collected by
Capt. Blakiston. This is a very great extension of its previously
known range.
18. Larus pominicanvs, Licht. .
Larus dominicanus, Licht. Verz. Doubl. p. 82 (1823); Darwin,
Zool. “ Beagle,” Birds, p. 142 (1841); Cassin, Orn. U.S. Expl.
Exp. p. 377 (1858), Callao?; Schlegel, M. P.-Bas, Lari, p. 12
(1863); Layard, B. S. Africa, p. 367 (1867) ; Durnford, Ibis, 1877,
p. 45 (Chuput, Patagonia) ; id. tom. cit. p. 201 (prov. B. Ayres).
Larus littoreus, Forster, Deser. Anim. p. 46 (1844), Cape of
Good Hope.
Larus antipodus, Gray, Cat. Anseres Brit. Mus. p. 169 (1844),
New Zealand. j
Dominicanus antipodus, Bruch, J. f. Orn. 1853, p. 100.
“Dominicanus pelagicus Anglor.,” Bruch, J. f. Orn. 1853, p. 100
(India and Oceania) ; id. op. cit. 1855, p. 280; Bp. Consp. Av. ii.
p. 214 (1857).
Dominicanus vetula, Bruch, J. f. Orn. 1853, p. 100, 1855, p.
281 (Cape Good Hope); Bp. Consp. Av. ii. 214 (1857).
“‘ Dominicanus vociferus Anglor.,” Bruch, J. f. Orn. 1853, p. 100,
1855, p. 281 (South America).
Dominicanus antipodum, Bruch, J. f. Orn. 1855, p. 281; Bp.
Consp. Av. ii. p. 214 (1857).
Dominicanus fritzei, Bruch, J. f. Orn. 1855, p. 281 (Straits of
Sunda, near Java?) (type inWiesbaden Mus. examined, H.S.); Bp.
Consp. Av. ii. p. 214 (1857).
Larus vociferus, Burm. Syst. Uebers. Th. Bras. p. 448 (1856) ;
id. La Plata-Réise, ii, p. 518 (1861).
1878. ] MR. H. SAUNDERS ON THE LARIN&. 181
Clupeilurus antipodum, Bp. Compt. Rend. xlii. p. 770 (1856).
Larus verreauxii, Bp. Rev. et Mag. Zool. vii. 1855, p. 16.
Dominicanus verreauxii, Bp.; Bruch, J. f. Orn. 1855, p.281(Chili).
Dominicanus azare (Less.), Bp. Consp. Av. ii. p. 214 (1857).
Lestris antarcticus (!), Ellman, Zool. 1861, p. 7472.
Lestris fuscus, id. Zool. 1861, p. 7472.
Larus vetula, Gurney, Andersson’s B. Damara Land, p. 357
(1872) ; Shelley, Ibis, 1875, p. 86 (Natal).
Hab. New Zealand, Kerguelen Island, and the other islands
between it and Cape of Good Hope, African coast to 22°S. lat., the
opposite coast of South America, the Falkland Islands, Patagonia,
the coast of Chili, and the island of Juan Fernandez. Iam sceptical
as to the locality assigned to L. fritzei, whilst equally unable to
accept Bonaparte’s version of Sunda being a mistake for Sund[ Lund? |
in Sweden!
In a large series of specimens from the above localities I can detect
no specific differences, individuals from the same localities often
varying quite as much in the dimensions of the bill as do those
from widely remote places. The absence or presence of the white
mirror near the tip of the first primary is of no specific value what-
ever, being entirely dependent upon the age of the individual ; it
does not appear till after the bird has assumed the full black mantle,
and increases in size with age.
The deep drown-black of the mantle, as distinct from the slate-
black of Z. fuscus, and its strong bill and larger size, will distin-
guish Z. dominicanus from that species; it is smaller than LZ. ma-
rinus, has a different pattern of primaries, and has olivaceous-coloured
legs and feet.
Messrs. Sclater and Salvin state (P. Z. S. 1871, p. 576) that the
examples then living in the Society’s gardens had flesh-coloured
legs and feet ; but this is either a slip of the pen, or else they must
have been looking at a ZL. marinus, the only ‘ Black-back” which
when adult has those parts of that colour.
19. Larus paciricus, Latham,
Larus pacificus, Latham, Suppl. Ind. Orn. p. 68 (1891); Gould,
B. of Austral. vol. vii. pl. 19; Schlegel, M. P.-Bas, Lari, p. 7
(1863).
Larus frontalis, Vieillot, in Nouv. Dict. H. Nat. 2nd ed. t. xxi.
p- 505 (1818), im. ad. (Tasmania).
Larus leucomelas, Vieillot, N. Dict. H. Nat. 2nd ed. t. xxi. p. 509
(1818), adult (Tasmania).
Larus bathyrinchus (sic), Macgill. Mem. Wern. Soc. vy. (1823-4),
« 253.
. Larus georgii, King, Surv. Intertrop. Australia, ii. p. 423 (1826)
(King George’s Sound, S.W. Australia).
Gabianus pacificus (Lath.), Bruch, J. f. Orn. 1853, p. 100, et
1855, p. 280; Bonap. J. f. Orn, 1854, p. 211; Rev. et Mag. Zool.
1855, p. 13; Consp. Av. ii. p, 212 (1857).
182 MR H. SAUNDERS ON THE LARINE. [Feb. 5,
Gabianus bathyrhynchus, Bruch, J. f. Orn. 1855, p. 280; Bp.
Consp. Av. ii. p. 212 (1857). se
Gabianus georgii (King), Bp. Consp. Av. ii. p. 213 (1857).
Hab. South-west portions of Australia, Bass’s Straits, and Tas-
mania; not included by Mr. Buller in his ‘ Birds of New Zealand,’
but there are three specimens labelled from that locality in the British
Museum, obtained by the Antarctic Expedition.
Mr. Gould’s plate hardly gives a correct idea of the dimensions and
great depth of bill in this fine species, which may easily be distin-
guished by this feature in all stages. In the adult the tail is
crossed by a black band; and this peculiarity in the plumage, coupled
with the stout bill, seems to place this species midway between the
typical Gulls and those of the next group. The value of Gabianus
_ as a genus for this species has been already discussed.
20. LARus BELCHER], Vigors.
Larus belcheri, Vigors, Zool. Journ. iv. p. 358 (1829); id. Zool.
Beecher’s Voy. ‘“ Blossom,” p. 39; Schlegel, M. P.-Bas, Lari, p. 9
(1863), excl. syns.; Scl. & Salv. P. Z.S. 1871, p. 575.
Adelarus belcheri, Bruch, J. f. Orn. 1853, p. 107; id. 1855,
p- 279; Bp. Naum. p. 212 (1854).
Leucopheus belcheri, Bp. Consp. Av. ii. p. 232 (1857).
« Larus fuliginosus, Gould,” Cassin, U.S. Expl. Exp. Orn. p. 378
(1858), nec Gould (Cape Horn to Callao).
Larus frobeenii, Phil. & Land. Wiegm. Archiv, 1861, p. 292.
Larus frobeni, iid. Cat. Aves Chil. An. Univ. Chil. tom. xxxi.
p- 288.
Hab. West coast of S. America, from Callao southwards to Chili,
the western portions of the Straits of Magellan, and down to Cape
Horn.
I have the fully adult bird with pure white head and underparts
from Chorillos, near Callao, Peru; but I observed the immature birds
with dark hoods in far greater numbers. Although several of these
Pacific Gulls have a hood in the immature stage, which is lost in the
adult, in none of them is the change so remarkable as in this species.
I confess that I cannot see any adequate reason for giving it generic
rank ; but Bonaparte thought differently, for he made it the type of
his genus Procellarus, being quite unaware that it was absolutely
the same species as the bird which he had already located in the
genera Leucopheus and Adelarus!
Dr. Coues, in a general notice of American Gulls, under the head
of L. heermanni (B. of N.W. p. 642), says that L. belcheri is “ not
a white-headed Gull at all,’ though he afterwards says that he
should not be surprised if, in the adult state, it lost its hood. In
this last surmise he is quite right; the adult L. belcheri isa perfectly
white-headed Gull, at the first glance being like a stout LD. fuscus
with a black band on its tail.
21. LARUS HEERMANNI, Cassin.
Larus heermanni, Cassin, Proc. Ac. Nat. Se. Philad. vi. p. 187
1878.} MR. H. SAUNDERS ON THE LARIN&E. 183
(1852) ; id. B. Californ. p. 28, pi. 5; Sel. & Salv. P.Z.S. 1871,
p- 574; iid. Nomencl. Av. Neotrop. p. 148.
Adelarus heermanni, Bruch, J. f. Orn. 1853, p. 107, et 1855,
. 279.
3 Leucopheus heermanni, Bp. Naumannia, 1854, p. 211.
Blasipus heermanni, Bp. Consp. Av. ii. p. 211 (1857).
Larus ( Blasipus) heermanni, Coues, B. N.W. Am. p. 641(1874).
Hab. Pacific coast, from Vancouver’s Island and California (breed-
ing), down to Panama in winter.
The nearest ally of this species is perhaps Z. crassirostris of the
opposite Asiatic coast and islands; but its red bill and the lead-
coloured neck and underparts will always serve to distinguish the
adult ; in Z. crassirostris the neck and underparts are white.
22. Larus crassrrostris, Vieill.
Larus crassirostris, Vieill. N. Dict. H. Nat. 2nd ed. p. 508 (1819),
ex Krusenstern (Nagasaki, Japan); Schlegel, M. P.-Bas, Lari, p. 8
(1863); Swinh. P.Z.S. 1871, p. 421; David & Oust. Ois. de la
Chine, p. 518 (1877).
Larus melanurus, Temm. Pl. Col. 77 liv. pl. 459 (1828); Temm.
& Schl. Faun. Japon. Aves, p. 132, pl. 88 (1850); Taczan. Bull.
Soc. Zool. France, i. p. 264 (1876).
Adelarus melanurus, Bruch, J. f. Orn. 1853, p. 107, et 1855,
p-. 279,
Blasipus crassirostris, Bp. Naumannia, 1854, p. 211; id. Consp.
Av. li. p. 212 (1857).
Larus fuscus, Swinhoe, Ibis, 1860, p. 68 (nee Linn.).
Hab. Coasts of Japan and China, and large lakes and rivers of
latter ; breeding in colonies, generally on islands.
I have already pointed out roughly the distinctions between this
species and L. heermannt ; superficially the present bird is not unlike
the adult of L. beleheri ; but the mantle is much lighter, and the tail
is white with a black bar, whereas in L. belcheri the greater portion
of the tail is black.
23. Larus mopestus, Tsch.
Larus modestus, Tschudi, Wiegm. Arch. 1843, pt. i. p. 389; id.
Fauna Peruana, Aves, p. 306 (1845-6), pl. 35; Scl. & Salv. P. Z.S.
1871, p. 573; id. Nomencl. Av. Neotrop. p. 148 (1873).
Larus bridgesi, Fraser, P. Z. 8. 1845, p. 16; id. Zool. Typ. t. 69
(1849), type in Brit. Mus.
Blasipus bridgesi (Fraser), Bruch, J. f. Orn. 1853, p. 108, et
1855, p. 280; Bp. Rey. Zool. 1855, p. 21; Consp. Av. ii. p. 212.
“ Blasipus polios (Natt.),’ Bp. Rev. Zool. 1855, p. 21; Consp.
Ay. ii. p. 212 (1857).
Leucophaius modestus, G. R. Gray, Hand-l. Birds, iii. p. 116
(1871).
Hab. Pacific coast of South America, from Callao to Valparaiso,
and probably further south.
This species, which in its immature plumage bears an indication
184 MR. H. SAUNDERS ON THE LARINZE. [Feb. 5
of a dusky hood, has been confused with L. fuliginosus of the Gala-
pagos Islands; but its much _slenderer bill, tarsi, and feet will
at once distinguish it from the latter, even in youth; whilst in
the adult the clear grey of the underparts and the white blending
into grey of the forehead and head, distinguish it from any other
known species of Gull. Mr. Fraser’s plate above cited gives a very
fair idea of the adult; but very old birds are much lighter about the
upper parts than his example.
24. Larus ruLicinosvs, Gould.
Larus fuliginosus, Gould, Zool. “ Beagle,” iii. p. 141 (1841) ;
Sel. & Salv. P. Z.S. 1871, p. 573; Salvin, Trans. Zool. S. ix. p.
505, pl. Ixxxvil.
Leucopheus fuliginosus et Adelarus neptunus, Bp. Rev. Zool.
1855, p. 20; et Consp. Av. ii. p. 232 (1857).
Procellarus heermanni! (part.), G. R. Gray, Hand-l. Birds, ii.
p- 116 (1871).
Hab. Galapagos Islands,
This stoutly built and well-marked species appears to be restricted
to the Galapagos group. Mr. Salvin’s illustration above cited will
probably prevent further confusion between this bird and ZL. mo-
destus ; but it is most extraordinary that the late Mr, G. R. Gray
should have confounded it with Z. heermanni, when he had the
type of the present species before him. Specimens are exceedingly
rare in collections ; and of the eggs and nestlings no examples are
known.
Although this species differs from the preceding ones in having
a hood in its adult plumage, yet its other affinities seem to indicate
that its proper place is here and not with the ordinary hooded
Gulls.
25. Larus scoressi, Trail.
Larus scoresbii, Trail, Mem. Wern. Soc. iv. p. 514 (1823), (New
South Shetland Islands) ; Pelzeln, Orn. Novara Exp. p. 151; Abbott,
Ibis, 1861, p. 165 (Falkland Islands).
Larus hematorhynchus, Vigors, in lett. King’s Zool. Journ. iv
p. 103 (1828-9); Jard. & Selby, Ill. Orn. ii. pl. 106 ; Gould, Zool.
“* Beagle,” Birds, p. 142.
Leucopheus hematorhynchus, Bruch, J. f. Orn. 1853, p. 108,
et 1855, p. 287; Bp. Naum. 1854, p. 211.
Procellarus neglectus, sive Kpitelarus neglectus, Bp. Naumann.
1854, pp. 211, 213; id. Rev. et Mag. Zool. p. 13 (1855); id. Consp.
Ay. ii. p. 211 (1857), type and sole representative of Procellarus.
Leucopheus scoreshii, Bp. Consp. Av. ii. p. 231 (1857); Blasius,
J. f. Orn. 1865, p. 378; Scl. & Salv. P. Z. S. 1871, p. 579; iid.
Nomencl. Av. Neotrop. p. 148.
Larus seoresbyi, Schl. M. P.-Bas, Lari, p. 33 (1863).
Hab. Patagonia, east coast, south of about 45° S. lat., down to
New South Shetland Islands, in about 63° S.; the Falkland Islands,
the Straits of Magellan, and up the coast of Chili as far as Chiloe.
1878. ] MR. H. SAUNDERS ON THE LARINA. 185
In spite of the somewhat peculiar shape of the bill in this species
I hardly think it desirable to place it on that account in a distinct
genus, especially as Bonaparte’s genus Leucopheus also includes
such dissimilar species as the present and L. heermanni of North
America. His Procellarus is founded on a young bird of the same
species! Larus scoresbii, however, is a very well-marked species,
from its short, stout, crimson bill, and coarse legs and feet, the webs
of the latter being a good deal incised. In the immature stage this
bird has a sooty hood ; but in the adult the upper parts are grey.
26. LaARUS NOV#-HOLLANDIA, Steph. (Fig. 1.)
Larus nove-hollandia, Stephens, Shaw’s Gen. Zool. xiii. pt. i.
p- 196 (1826), ex Latham.
Larus scopulinus, var. major, Forst. Deser. Anim. p- 106 (1844),
Figat:
is 2. 3.
Three outer primaries of L. nove-hollandie, jr. (from the type of Gavia
pomarre, Bruch, of 1853, not of 1855).
Larus jamesonii, Wilson, Il. Zool. pl. xxiii. (1831),
Xema jamesonii, Gould, Birds of Australia, vol. vii. pl. xx (1848).
aoe Jamesonii,. Wils. Bruch, J. f. Orn. 1853, p. 102; 1855,
p. 285.
186 MR. H. SAUNDERS ON THE LARINZ. [Feb. 5,
Gavia andersonii, Bruch, J. f. Orn. 1853, p. 102, et 1855, p. 285
(type examined, H. S.).
Gavia pomarre, Bruch, J. f. Orn. 1853, p. 103, not Gavia pomare
of 1855, p. 285, which is L. bulleri (type in Mainz Mus. examined,
H.S8.), Society Islands.
Gelasies gouldi, Bp. Naumann. 1854, p. 216.
Gelastes corallinus, Bp. Naumann. 1854, pp. 212, 216 (type in
Paris M. examined, H. 8.).
Gelastes andersonii, Bp. tom. cit. p. 212.
Gavia gouldii, Bp. Bruch, J. f. Orn. 1855, p. 285.
Fig. 2.
is 2. 3.
Three outer primaries of L. nove-hollandie, ad. (from the type of Gavia
andersonit, Bruch),
Bruchigavia gouldi, Bp. Consp. Av. i. p. 228 (1857).
Bruchigavia pomare, Bp. tom. cit. p. 229.
Bruchigavia jamesonii, Bp. tom. cit. p. 228 (1857); Gould, Handb,
B. Austral. ii. p. 387 (1865).
Bruchigavia corallinus, Bp. op. cit. p. 228 (1857).
Larus scopulinus major, Schlegel, M. P.-Bas, Lari, p. 29 (1863),
Hab. Australia from Raine Island, Torres Straits, to Bass’s Straits,
and Tasmania ; also New Caledonia, and perhaps the Society Islands.
1878. | MR. H. SAUNDERS ON THE LARIN#. 187
Although very close to LZ. scopulinus of New Zealand, I think
this species may fairly be distinguished by its larger size throughout,
and by the greater amount of white mirror in the pattern of the
three outer primaries. In old birds of L. nove-hollandieé there is
always a mirror on the third primary as well as on the first and
second ; in a large series of LZ. scopulinus 1 have never found this.
The amount of white is also greater in the Australian bird, and the
shape of the mirror is different, as is shown in the accompanying
fignres. L. corallinus of Bonaparte has been supposed to be LZ. macu-
lipennis, Licht., on the strength of the assertion in the Conspect. Av.
that the type was obtained in Brazil by Castelnau. I have examined
the type, which bears no indication of locality, nor do I for a moment
Fig. 3.
Three outer primaries of L, nove-hollandie, old.
believe that it ever was killed in Brazil ; it is, however, of this species,
and a large-billed example.
To make matters plainer, I have had figures prepared of the three
outer primaries in three different specimens of this species. In
fig. 1 (p. 185) is given the pattern in a young bird, taken from a
drawing by Dr. O. Finsch of the type of Gavia pomarre of Bruch,
1853 (but noé his G. pomare of 1855, which latter =Z. dulleri), in
the Mainz Museum. Fig. 2 (p.-186) represents the primaries of
Bruch’s G. andersonii, from the same source. Fig. 3 (p. 187)
188 MR. H. SAUNDERS ON THE LARINZ, [Feb. 5,
shows the pattern of an old L, nove-hollandie, obtained in Tasmania
by Capt. V. Legge, R.A., and, if compared with fig. 4 (p. 188), will
show the difference between this species and old L. scopulinus.
27. Larus scoputinus, Forst. (Fig. 4.)
Larus scopulinus, Forster, Descr. Anim. p. 106 (1844), New
Zealand ; Schlegel, M. P.-Bas, Lari, p. 28 (1863); Finsch, J. f.
Orn. 1870, p. 360; Buller, B. of N. Zeal. p. 273 (1873).
Fig. 4.
1.
)
Three first primaries of L. scopulinus, old.
Larus nove-hollandie, Gray, Voy. Ereb. & Terr. Birds, p. 18
(1844), New Zealand (nec Stephens).
Lestris scopulinus, Ellman, Zoologist, 1861, p. 7472.
Hab. New Zealand.
Under the head of the preceding species I have pointed out the
differences which seem to me to separate this form from L. nove-
hollandie ; but the drawing (p. 188) of the first primaries of an old
bird will show the principal point of distinction better than any
description.
28. Larus HartLaust (Bruch). (Fig. 5.)
Gavia hartlaubi, Bruch, J. f. Orn. 1853, 102, et 1855, p. 268,
Cape G. Hope (type examined, H. S.).
1878. ] MR. H. SAUNDERS ON THE LARINE. 189
“Larus poiocephalus, Sw.,’, Layard, B. 8. Africa, p. 368 (1867),
nec Sw.
Larus hartlaubi, Saunders, P. Z. 8S. 1874, p. 293.
Hab. Southern coast of Africa, especially about Table Bay, Cape
of Good Hope, where Mr. Layard obtained many specimens and also
eggs, some of which he presented to me.
For along time this species was supposed to be the winter or
hoodless dress of the grey-capped L. pheocephalus, Sw.; but, as I
have already pointed out (P. Z. S. 1874, p. 293), the present species
Fig. 5.
1 z: 3.
Three first primaries of L. hartlaubi, ad.
never has a hood at all. It is quite distinct from its close allies L.
scopulinus and L. nove-hollandia, and may be recognized by its
smaller size, proportionally longer and slenderer bill, which is of a
rich crimson, and by the more sooty colour of the under wing-
coverts, especially along the carpal joint. There is a small elongated
mirror on the first and second primaries ; but the remaining portions
of those feathers are black almost to the roots. The drawing (fig. 5)
shows the pattern of the primaries.
190 MR. H. SAUNDERS ON THE LARINZ. (Feb. 5;
29. Larus BuLLERI, Hutton. (Figs. 6 & 7.)
Gavia pomare, Bruch, J. f. Orn. 1855, p. 285 (not of 1853,
which = L. nove-hollandie).
Bruchigavia melanorhynehus, Buller, Ibis 1869, p. 43 (nee L.
melanorhynchus, Temm.).
Larus (Bruchigavia) melanorhynchus, Finsch, Ibis, 1869, p. 381 ;
Travers, Trans. N.Z. Inst. 1871, p. 209.
Larus bulleri, Hutton, Cat. Birds New Zeal. 1871, p. 41; Potts,
Ibis, 1872, p. 38; Buller, B. New Zeal. p. 276, et fig.
Hab, This species appears to be restricted to New Zealand, and
perhaps to the South Island,
I have examined the type of Bruch’s Z. pomare of 1855; anil it is
undoubtedly of this species; but the type of his Z. pomarre of 1853
Fig. 6.
3.
Three outer primaries of ZL. dulleri, old, from the type.
is as certainly L. nove-hollandi@ ; and it is to the latter that Mr.
Buller alludes as having been examined by him previous to the pub-
lication of his ‘ Birds of New Zealand.’ The third specimen entitled
L. pomare in the Mainz collection is a young L. ridibundus! Bruch’s
name, therefore, cannot be employed, having been previously applied
to another species; and this species must stand as Larus bulleri,
1878. | MR. H. SAUNDERS ON THE LARINZ. 191
Hutton. It is very distinct from L. scopulinus, and appears to fre-
quent inland lakes and river-beds in preference to the sea-shore.
The drawings (pp. 190, 191) showing the distinctive markings of the
three outer primaries will be better than any description,
i Oe 3
Three outer primaries of L, bulleri, nearly ad., from the type of Gavia
pomare, Bruch, of 1855.
During my recent visit to Bremen I went into the question of this
and the three preceding species with Dr. Finsch, who had previously
studied the subject and had made numerous and careful drawings of
the primaries of Bruch’s types of Z. pomare in the Mainz Museum,
and of many other specimens. These drawings he most generously
placed at my disposal ; and, thanks to his liberality, I am enabled to
figure the primaries of two of Bruch’s types, bearing the same name,
but belonging to two totally distinct species,
39. Larus Getasrrs, Licht, (Fig. 8.)
Larus gelastes, Licht. in Thienem. Fortpflanz. Vog. Eur. pt. v.
p. 22 (1838), type in Berlin Mus. ; Keys. & Blas. Syst. Verzeichn.
Europ. Siiug. p. 95 (1840); Degl. Orn, Europ. ii. p. 318 (1849) ;
Bree, B. Eur. 2nd ed. v. p- 72 (1876); Blanford, East Persia, ii.
p- 291 (Makran coast),
“Larus leucocephatus, Boissonneau,”’ fide Keys. & Blas. op. cit,
p. 22.
192 MR. H. SAUNDERS ON THE LARINZ. | Feb. 5,
Larus genei, De Bréme, Rev. Zool. ii. (1839) p. 321.
Larus tenuirostris, Temm. Man. d’Orn. 2nd ed. pt. iv. p. 478, pl.
(1840) ; Crespon, Faune Mérid. ii. p. 126 (1844).
Xema lambruschinii, Bp. Icon. Faun. Ital. i. Uce. p. 135, pl. (livr.
xxvii. 1840).
Xema gelastes (Licht.), Boie, Isis 1844, p. 192.
Xema genei (De Bréme), id. loc. cit.
Gavia gelastes, Bruch, J. f. Orn. 1853, p. 102, et 1855, p. 286.
Larus columbinus, Golowatschow, Bull. Soc. Imp. Nat. Mose.
1854, t. i. p. 435.
“« Gelastes rubriventris (Vieill.),”’ Bp. Naum. 1854, p. 216.
Fig. 8.
a 2. 3.
Three outer primaries of L. gelastes, juv.
Gelastes lambruschini, Bp. Cat. Parzud. p. 11 (1855) ; id. Consp.
Av. ii. p. 227.
Larus subroseus et Larus brehmii, Heug). Syst. U. Vog. N.O.-
Afr. Sitz. Ak. Wiss. Wien, 1856, p. 321; cf. Heugl. Orn. N.O.-Afr.
ii. Abth., Band 2, p. 1412.
Gelastes columbinus. Bp. Consp. Av. ii. p. 227 (1857).
“ Gelastes leucocephalus, Brisson,” Bp. Consp. Av. ii. p. 227
(syn.).
1878. } MR. H. SAUNDERS ON THE LARIN®. 193
Larus lambruschini, Bp., Schl. M. P.-Bas, Lari, p- 28 (1863) ;
Hume, Stray Feath. i. p. 274 (1873).
Chroicocephalus gelastes, Licht. Nomencl. Av. M. Berolst. p. 98.
“Larus arabicus, Hemp. & Ehr.,” Mus. Berolst. (fide H. S.),
Hab. South coast of Spain, the Mediterranean and Black Sea
(breeding); the Red Sea, Persian Gulf, and the coasts as far as
Kurrachi; also West Africa to Senegal, whence there is a specimen
in the Paris Museum.
This species seems to have no very near allies. It never hasa hood ;
yet its structure and wing-pattern remind us of L. ridibundus. In
appearance it is like ZL. scopulinus; but its wings are very long, its
bill is slender, its flight is Tern-like, and its eggs are singularly like
those of Sterna cantiaca and S. media. I locate it here because I
do not know of any better position for it.
31. Larus LevcopHTHaumus, Licht.
Larus leucophthalmus, Licht.; Temm. Pl. Col. liv. 62, pl. 366
(1825); id. Man. d’Orn. 4™° pt. p. 486 (1840); Heugl. Ibis, 1859,
p- 349; Konig-Warth. Ibis, 1860, p- 129; Finsch & Hartl. Vog.
Ostafr. p. 821 ; Finsch, Trans. Zool. Soc. vii. pt. vi. p. 302; Schl.
M. P.-Bas, Lari, p. 32.
Iris (mispr.) leucophthalmus, Lesson, Tr. d’Orn. p- 618 (1831).
Xema lecophthalmum, Bp. Ucc. Eur. p- 78 (1842).
Xema leucophthalma, Gray, Brit.-M. List, Anseres, p. 171 (1844).
Adelarus leucophthalmus, Bruch, J.f. Orn. 1853, p. 106, et 1855,
p. 278; Bp. Compt. Rend. xlii. p. 771 (1856) ; Blasius, J. f. Orn.
1865, p. 378.
Chroicocephalus leucophthalmus, Brehm, Naumannia, 1855, p- 295.
Larus masauanus, Heugl. Peterm. Geogr. Mittheil. 1861, p. 31.
Hab. Temminck states that this species visits Greece and the
shores of the Bosphorus; but no recent travellers in those parts
have ever met with it; and although two energetic naturalists, Dr.
Kriiper and M. Alléon, have for years been resident in the neighbour-
hood and have thoroughly explored the Greek Archipelago and the
Bosphorus, they have never met with it, either there or in the Eubcean
channel, which Lindermayer (Vog. Griech. p. 177) so particularly
indicates. Even in the Red Sea it is, according to Von Heuglin,
very scarce north of the tropic, though frequent more to the south ;
it does not, however, appear to go beyond the Gulf of Aden and the
Somali coast ; for neither Blanford nor Hume mention it as occurring
along the Mekran coast ; and its range is therefore much less exten-
sive than that of the allied species LZ. hemprichi.
32. Larus HempPricut (Bp.).
Xema crassirostris (Licht.), Boie, Isis. 1844, p- 192.
Larus crassirostris, Licht. Nomencl. p. 99 (1854) (nec Vieill.),
Mus. Berolst.
Adelarus hemprichii, Bp.; Bruch, J. f. Orn. 1853, p- 106 (deser.),
et 1855, p. 278; Bonap. Naum. 1854, p- 212; Blasius, J. f. Orn.
1865, p. 378.
Proc. Zoou. Soc.—1878, No. XIII. 13
194 MR. H. SAUNDERS ON THE LARIN. [Feb. 5,
Larus hemprichi, Heugl. Ibis, 1859, p. 350 ; Konig-Warth. Ibis,
1860, p. 129 (nidif.); Schleg. Mus. P.-Bas, Lari, p. 32; Finsch,
Trans. Zool. S. vii. pt. vi. p. 302, pl. xxvii.; Finsch & Hartl.
Vig. Ostafr. p. 823; Blanford, Abyssinia, p. 441 (Red Sea) ;
Hume, Stray Feath. i. p. 279; Blanford, East Persia, il. p. 292
(Aden to Kurrachee harbour); Hume, Stray F. 1876, p. 414; Butler,
op. cit. 1877, p. 296.
Hab. This Gull appears to have a wider range than its congener L.
leucophthalmus, as it is not confined to the Red Sea south of the
tropic, but also frequents the Persian Gulf and the coast eastward as
far as Bombay. Von Heuglin and Baron Ko6nig von Warthausen
(Joc. cit.) have given an excellent account of the habits and nidifi-
cation of both these species; and more recently in ‘Stray Feathers’
Capt. Butler has given a description of the great colony on the island
of Astola.
In the Bulletin de la Société Zoologique de France, 1"° partie, 1877,
p. 32, M. Jules Vian gives an account of two immature specimens,
one of L. leucophthalmus and one of L. hemprichi, which were
supposed to have been obtained on the coast of Nice. M. Vian
showed me these specimens when I was last in Paris; and they are
undoubtedly genuine examples ef the respective species. M. Vian
thinks that the opening of the Suez Canal may have caused their
appearance upon the shores of France; I cannot prove the contrary,
but, from the make-up of the skins, I must confess I am very sceptical
as to their having got so far as Nice alive. They had passed through
at least one, if not two dealers’ hands before M. Vian saw them.
33. Larus arricitua, Linn.
Larus atricilla, Linn. Syst. Nat. i. p. 225 (1766), nec Pallas (ex
Catesby) ; Gm. Syst. Nat. i. p. 600 (1788); Temm. Man. d’Orn.
ed. 2, pt. ii. p. 779 (Mediterranean, in error); Montagu, Orn.
Dict. Rennie’s ed. p. 259 (1833) (Winchelsea) ; Schl. M. P.-Bas,
Lari, p. 44 (1863); Scl. & Salv. P. Z. 8. 1871, p. 576; Coues,
B. N.-West Am. p. 650 (1874); Reid, Zoologist, 1877, p. 489
(Bermudas).
Larus ridibundus, Wilson, Am. Orn. ix. p. 89, pl. 74. fig. 4
(1814), nee Linn. ; Léotaud, Ois. de Trinidad, p. 532.
Xema atricilla, Boie, Isis, 1822, p. 563; Cab. in Schomb.
Guiana, ili. p. 761.
Gavia atricilla, Macgill. Man. Brit. Orn. ii. p. 240 (1842).
Chroicocephalus atricilla, Bruch, J.f. Orn. 1853, p. 106; Lawr.
B. N. Am. p. 850 (1858).
Chroicocephalus serranus, Bruch, J. f. Orn. 1853, p. 106 (nec
Tschudi).
Atricilla catesbyi, Bruch, J. f. Orn. 1855, p. 287.
*‘Atricilla megalopterus,” Bp.; Bruch, J. f. Orn. 1855, p. 287.
*Atricilla micropierus,’ Bp.; Bruch, tom. cit. p. 288.
Hab. America, from Maine, on the east coast, down to the mouth
of the Amazons and to the West-Indian Islands; on the west coast,
1878. | MR. H. SAUNDERS ON THE LARIN&. 195
California, Mexico, Guatemala, and as far south as Tumbez, the
northern frontier of Peru (Jelski). Once obtained by Col. Montagu
at Winchelsea, Sussex. The specimen in the British Museum, from
his collection, is undoubtedly of this species ; but there seems to be
no warrant for its reputed occurrence in Southern Europe and the
Mediterranean.
This species may always be known by its black primaries.
34. LARUS FRANKLINI, Sw. & Rich.
Larus atricilla, J. Sabine, App. Franklin’s Polar Sea, p. 695
(1823), nec Linn. nec auctt. (the description clearly applies to this
species).
Larus franklini, Sw. & Rich. F. Bor.-Am., Birds, p. 424, pl. 1xxi.
(1831); Schlegel, Mus. Pays-Bas, Lari, p. 36 (1863); Scl. & Salv.
P. Z. 8. 1871, p. 577; Newton, P. Z. S. 1871, p. 57, pl. iv. fig. 4,
ege (Manitoba) ; Coues, B. N.-West Am. p. 653 (1874).
Larus cucullatus, Licht. MSS. (Mexico), type in Berlin Mus.
(examined, H. S.).
Larus pipixcan, Wagler, Isis, 1831, p. 515.
Xema franklini, Bp. Comp. L. B. Eur. & N. Am. p. 62 (1838) ;
Boie, Isis, 1844, p. 194.
Xema pipixcan, Boie, loc. s. cit.
Chroicocephalus franklini, Bruch, J. f. Orn, 1853, p. 104, et 1855,
p- 289; Lawr. B. N. Am. p. 851 (1858).
Chroicocephalus cucullatus, Bruch, J. f. Orn. 1853, p. 104, et 1855,
p. 290; Lawr. B. N. Am. p. 851 (1858) ; Coues, Proc. Phil. Ac.
1862, p. 309.
Chroicocephalus kitlitzii, Bruch, J. f. Orn. 1853, p. 104 (described
from a drawing), nec Swinhoe, P. Z.S. 1860, which=ZL. saundersi.
Chroicocephalus schimperi, Bruch, J. t. Orn. 1853, p. 104 (nec
Schlegel, 1863, which = Z. saunders?).
Larus cinereo-caudatus, Ph. et Landb. Wiegm. Arch. 1861], p. 293.
Hab. Interior of North America, west of the Mississippi; breeds
in Manitoba; seldom visits the Atlantic coast, but has once occurred
at St. Bartholomew’s, West Indies (Sund.) ; goes down the Pacific
coast as far as Chili, whence I have a fully adult example with partial
hood, collected by Mr. E. Reid, of Santiago, and also one from
Callao, Peru.
The primaries of this species undergo much alteration with the
age of the bird; and in time the subapical mirror on the first extends
over the greater part of the webs.
From the description there can be little or no doubt that Bruch’s
L. kitlitzi and L. schimperi are referable to this species. Both
were suppressed in his second review (1855), when he corrected a
few of his more glaring errors; and all trace of the second name has
vanished from the Mainz Museum, where J especially looked for it, as
Schlegel had adopted it for a totally different Chinese species. The
type was said to have come from New Zealand, had a bright red bill,
dark hood, and black primaries with white tips. i
13
196 MR. H. SAUNDERS ON THE LARINE. [Feb.5,
35. Larus serrANus, Tsch. (Fig. 9.)
Larus serranus, Tschudi, Wiegm. Arch. 1844, pt. i. p. 314;
Fauna Peruana, Aves, p. 307 (1845-6) ; Scl. & Salv. P. Z. S. 1871,
3 GGT
7 Chroicocephalus personatus (Natt.), Bruch, J. f. Orn. 1853, p. 104,
et 1855, p. 289.
Xema cirrhocephalum, Peale ; Cassin, U.S. Expl. Exp. Orn. p. 381
(1858).
Larus glaucotis, Cassin, 7. e. p. 381 (1858).
Gavia serrana, Bp. Rey. Zool. 1855, p. 19.
Larus personatus, Natt.; Schl. M. Pays-Bas, Lari, p. 35 (1863).
Gavia personata, Blasius, J. f. Orn. 1865, p. 372.
Larus bonapartit, Scl. & Salv. P. Z. 8. 1868, p. 178.
Hab. This fine black-hooded species is found throughout the Cor-
dillera and the Andes of Northern Chili, Bolivia, Peru, and as far
north as Ecuador, whence Mr. Salvin has a specimen. It breeds
Fig. 9.
ir 2. 3.
Three outer primaries of L. serranus, ad.
in colonies on the shores of mountain-lakes, and is well known to
the Quichua-speaking Indians under the name of ‘ Quiulla,” doubt-
less an imitation of its ery. During the bad weather it descends to
the west coast. The eggs and nestlings are as yet unknown to me.
1878.] MR. H. SAUNDERS ON THE LARIN&. 197
Messrs. Sclater and Salvin (P. Z. S. 1871, p. 577) quote Bur-
meister as an authority for the occurrence of this species near Mendoza
in the Argentine Republic; but the deseription which he gives
clearly applies to L. maculipennis; indeed the dimensions cited
being smaller than those of his L. maculipennis, Licht. (which,
again, is not that species, but L. cirrhocephalus, Vieill.), show
clearly that his bird cannot possibly be the real Z. serranus, which
is the largest of the group of the true Hooded Gulls in America.
On the other hand, the dimensions assigned by Peale and Cassin to
their respective X. cirrhocephalum and L. glaucotis (18 inches long),
and the descriptions of the markings of the primaries, go to prove that
those names must be referred to this species. The figure (p. 196)
shows the pattern of the primaries in an adult and tolerably old bird.
36. LARUS BRUNNEICEPHALUs, Jerdon. (Fig. 10.)
Larus brunneicephalus, Jerdon, Madras Journ. xii. p- 225 (1840);
Schl. Mus. P.-Bas, Lari, p- 35 (1863).
Fig. 10.
Three outer primaries of Z, brunneicephalus, ad.
Xema brunnicephala, Gray, List of B. in Brit. Mus. iii. p- 172
(1844); Jerdon, B. of India, iii. p. 832 (1864); Holdsworth, P. Z. 8,
1872, p. 480 (Ceylon).
198 MR. H. SAUNDERS ON THE LARIN. [Feb. 5,
Chroicocephalus brunnicephalus, Bruch, J. f. Orn. 1853, p. 105.
Chroicocephalus brunniceps, Cabanis, J. f. Orn, 1853, p. 105
(note); Bruch, J. f. Orn. 1855, p. 291. :
Chroicocephalus brunneicephalus, Bp. Compt. Rend. Ac. Se. xli.
p. 73 (1856) ; Swinhoe, P. Z.S. 1871, p. 421; Dav. et Oust. O. de
la Chine, p. 521. c
Xema brunneicephalum, A. David, N. Arch. Mus., Bull. vii. no.
460 (1871).
Larus lacrymosus, Licht. (Bengal, imm.), type in Berlin Mus.
(examined, H. S.).
Gavina lacrymosus (Licht.); Bp. Naum. 1854, p. 212.
Gavia brunnicephalus (Jard.); Bp. Naum. 1854, p. 213.
Chroicocephalus tibetanus, Gould, P. Z.S. 1864, p. 54. (Tibet,
Major Hay. |
Xema brunneicephala, Hume, Yarkand Exp. Orn. p. 300, pl.
xxall. (1873); Blyth, J. A.S. B. 1875, pt. ii. p. 162 (Burma).
Hab. Henderson found this Gull very abundant in full breeding-
plumage in July, at an elevation of 15,000 feet, near the Pangong
lake ; and in winter it occurs both on the inland waters and along the
coast of India, as far west as Sind. Eastward it occurs in Burma and
visits Ceylon, where it is abundant. David obtained it abundantly in
Mongolia and China, apparently as near to the sea-board as Pekin.
Taczanowski, however, does not cite it amongst the species found in
any part of Siberia; and it is therefore doubtful if it can be
Middendorff’s L. ridibundus, var. major. As regards its reported
occurrence in Japan, Cassin’s bird obtained in Parry’s expedition is
clearly L. ridibundus.
37. Larus 1cuTHyAETUuS, Pall.
Larus ichthyaétus, Pallas, It. ii. App. no. 27 (1776), Caspian ;
Gm. Syst. Nat. i. p. 599 (1788); Pallas, Zoogr. Ros.-As. ii. p. 322
(1811) ; Riipp. Reise N. Afr. Atlas, p. 27, pl. 17 (1826), Red Sea;
Cassin, Perry’s Exp. Japan, ii. p. 232 (1856) (?); Tristram, Ibis,
1868, p. 330, Palestine; Shelley, B. of Egypt, p. 307, pl. 13
(1872); Schlegel, M. P.-Bas, Lari, p. 34 (1863) ; Dresser, B. of
Europe, pt. xviii. (June 1873).
Ichthyaétus, Kaup, Nati]. Syst. pp.102,169 (1829), type of genus
Ichthyaétus.
Xema ichthyaetum, Bp. B. Eur. & N. Am. p. 62 (1838).
Larus kroicocephalus, Jameson, Journ. As. Soe. viii. p. 242 (1839).
Kroikocephalus ichthyetus, Jerdon, B. of India, iii. p. 831
(1864).
Chroicocephalus icthyaétus, Bruch, J. f. Orn. 1853, p. 104;
Swinhoe, P. Z. 8. 1863, p. 327.
Larus ichthyaétus minor, Schl. M. P.-Bas, Lavi, p. 34 (1863).
Hab. Caspian Sea and inland lakes, Lake of Galilee (Tristram),
Mediterranean coast of Egypt and up to Nubia, the Red Sea, thence
to Bombay and down the coast to Madras, and on the lakes and
Jarge rivers of Northern India. As a straggler, in the Black Sea and
the Greek Archipelago, and once at the mouth of the Exe, in Eng-
1878. | MR. H. SAUNDERS ON THE LARINZE. 199
land Cassin states that Perry’s expedition obtained two specimens
in Yedo Bay, Japan, where it is described as abundant ; but both the
supposed examples were young birds without a trace of a hood, only
striated on the head; and the other points of the description would
equally apply to young Herring-Gulls, which I am inclined to think
they were. At all events the reported occurrence of this Gull in
Japanese waters remains unconfirmed; Capt. Blakiston has never
met with it; and Capt. St. John, H.M.S. ‘Sylvia,’ during the years
he was surveying those coasts, never saw it.
38. Larus MELANocEPHALUS, Natt. (Fig. 11.)
Larus melanocephalus, Natt. Isis, 1818, p. 816; id. in Temm.
Man. d’Orn. 2nd ed. ii. p. 777 (1820); Schl. M. P.-Bas, Lari,
Fig. 11.
Three outer primaries of L. melanocephalus, jr.
p. 43 (1863); Degl. & Gerbe, Orn. Europ. ii. p. 437 (1867) ;
Bree, B. of E. 2nd ed. y. p. 81 (breeding in Black Sea) ; Saund. Ibis,
1872, p. 79 (Thames).
Xema melanocephalus, Boie, Isis, 1822, p. 365.
Xema melanocephalon, Brehm, Vig. Deutschl. p. 757 (1831).
200 MR. H. SAUNDERS ON THE LARIN2. [Feb. 5,
Xema caniceps, Brehm, Vog. Deutschl. p 758.
Chroicocephalus melanocephalus, Bruch, J. f. Orn. 1853, p. 104.
Melagavia melanocephalus, Bp. Naum. 1854, p. 213.
Gavia melanocephala, Bp. Compt. Rend. 1856, xhi. p. 771.
Hab. Mediterranean and Black Sea ; outside the Straits of Gib-
raltar I observed it, apparently breeding, in the marshes of Huelva;
and it regularly ascends the west coast of the Iberian peninsula and of
France as far as Bordeaux, as is proved by the specimens annually ob-
tained there. It is therefore not at all astonishing that an immature
specimen should have been shot out of a flock of ZL. ridibundus at
Barking Creek in January 1866. This example is in the British
Museum.
In old birds the primaries are white, excepting a very black streak
down the outer web of the first primary only ; but it is not unusual
to find birds with the full black hood and white tail indicative of
maturity, but with a good deal of black on both the outer and inner
webs of the outer five primaries, the black crossing both webs near
their extremities and forming a subapical bar. The drawings (fig.
11, p. 199, and fig. 12, p. 201) show the differences between the
pattern of the three outer primaries of a bird of the year, shot in
March, and that of a L. ridibundus of about the same age.
39. Larus r1ipiBuNDvs, Linn. (Fig. 12.)
? Larus cinerarius, Linn. Syst. Nat. i. p. 224 (1766).
Larus ridibundus, Linn. Syst. Nat. 1. p. 225 (1766) ; Schl. M. P.-
Bas, Lari, p. 37 (1863).
Larus erythropus, Gm. Syst. Nat. i. p. 597 (1788).
Larus cinerarius, Schaff. Mus. Orn. p. 63 (1789); Pall. Z.
Rosso-As. ii. p. 326.
Larus canescens, Bechst. Orn. Tash. p. 370 (1802).
Larus atricilla, Pallas, Zoogr. Rosso-As. ii. p. 324 (1811), nee
Linn.
Larus nevius, id. tom. cit. p. 327.
Larus capistratus, Temm. Man. d’Orn. 2nd ed. pt. ii. p. 785
(1820).
Xema ridibundus, Boie, Isis, 1822, p. 563.
Xema capistratus, Boie, Isis, 1822, p. 563, et 1844, p. 192.
Xema ridibundum, Brehm, Vog. Deutsch. p. 760 (1831).
Xema pileatum, id. op. cit. p. 761.
Xema capistratum, id. op. cit. p. 762.
Chroicocephalus capistratus, Eyton, Hist. Rarer Brit. B. p. 63
(1836); Bruch, J. f. Orn. 1853, p. 105.
Chroicocephalus ridibundus, Eyton, Cat. Brit. Birds, p. 53 (1836) ;
Bruch, J. f. Orn. 1853, p. 105; Swinhoe, P. Z. 8. 1871, p. 421;
David & Oust. Ois. de la Chine, p. 520 (1877).
Gavia ridibundus, Bp.. Naumannia, 1854, p. 213.
Gavia capistratus, Bp. op. cit. p. 213.
Chroicocephalus pileatus, Brehm, Naum. 1855, p. 295.
Larus brunneicephalus?, Cassin, Perry’s Exp. Japan, ii. p. 233
(1856), clearly this species in winter dress.
1878. ] MR. H. SAUNDERS ON THE LARIN&. 201
Larus eahirinus, “ Hemp. & E., Syria,’ Mus. Berolst (fide H. S.).
Larus eahiricus, “ Ehr, Arabia,” id. (fide H. S.).
Hab. Northern and temperate Europe, breeding; the Mediter-
ranean coast to Egypt and Asia Minor, the Hed Sea, Arabian coast,
and the coast, interior waters, and rivers of India; ascending for 500
miles up the Burrampootra (Godwin-dusten), Burma, China, and
Three outer primaries of L. ridibundus, jr.
Japan in winter; also throughout the more temperate portions of
Siberia (breeding). It is said by Schlegel to visit “‘South Africa ;”
but I remain rather suspicious of the accuracy of the collectors who
are responsible for that somewhat vague locality, so long as this state-
ment is unconfirmed from other sources.
40. Larus macutirennis, Licht. (Fig. 13.)
Larus maculipennis, Licht. Verz. Doubl. p. 83 (1823), Monte
Video (type examined, H. S.); Scl. & Salv. Nom, Av. Neotrop. p.
148 (1873); Durnford, Ibis, 1877, p. 43 (Chuput valley, Patagonia) ;
id. tom. cit. p. 202 (prov. B. Ayres).
Xema cirrhocephalum, Gould, Zool. Beagle, iii. p. 142 (1841),
partim.
202 MR. H. SAUNDERS ON THE LARIN&. [Feb. 5,
Xema cirrhocephala?, Gray, List Birds Brit. Mus. iii. p. 173 (1844),
partim (East Patagonia).
Chroicocephalus maculipennis, Bruch, J. f. Orn. 1853, p. 105.
Gavia maculipennis, Blasius, J. f. Orn. 1865, p. 374.
“* Larus serranus, Tsch.,” Burmeister, La Plata-Reise, ii. p. 519
(1861), nee Tschudi (Entre Rios, Mendoza).
Larus cirrhocephalus, Hudson, P. Z.S. 1870, p. 802; id. P. Z.S.
1871, p.4, nee Vieillot.
Larus glaucodes, Saunders, P. Z. 8. 1874, p. 294 (partim).
Hab. Chuput valley, North-east Patagonia (breeding), up to the
La Plata, the Argentine Provinces, Mendoza, Entre Rios, Uruguay,
and the south coast of Brazil.
Fig. 13.
Three outer primaries of L. maculipennis, old,
Burmeister’s description of the bird which he calls L. serranus
clearly applies to this species; the dimensions (length 12 in., wing
11 in., tarsus 2 in.) exactly suit it, whilst the wing of true L. serranus
is over 14 in. long.
The distinction between this species and the next, Z. glaucodes,
seems to rest upon the patterns of the primaries, as shown in the
drawings (figs. 13 and 14). In L. maculipennis the black forms a
bar ; in L. glaucodes it is a mere border: and this is perfectly constant.
1878. ] MR. H. SAUNDERS ON THE LARINE. 203
T have selected the primary of the oldest L. maculipennis I could
find as presenting the least amount of black, and therefore closest to
L. glaucodes ; but, in spite of that, the difference is quite recognizable.
41. Larus euaucopes, Meyen. (Fig. 14.)
Larus glaucodes, Meyen, Obs. Zool. p. 115, pl. xxiv. ; id. Beitr.
Zool. p. 239, pl. xxxiv. (1834); Gay, Faun. Chil. Aves, i. p. 480
(1847); Cassin, Birds U.S. Astronom. Exp. p. 204 (1855); Scl. &
Salv. Nom. Av. Neotrop. p. 148 (1873).
Larus glaucotes, Cabanis, Ibis, 1861, p. 312.
Larus glaucotis, Schlegel, Mus. P.-B. Lari, p. 42 (1863).
Xema cirrhocephalum, Gould, Zool. Beagle, ili. p. 142 (1841),
part., nec Vieill.
Xema cirrocephala, Gray, List Birds Brit. Mus. iii. p. 173 (1844).
Fig. 14.
1. yep 3:
Three outer primaries of L. glaucodes, old.
Xema glaucodes, Boie, Isis, 1844, p. 192 ?
Larus albipennis, Peale, Zool. U.S. Expl. Exp. p. 288 (1848) ;
fide Cassin, in Orn. U.S. Expl. Exp. p. 379 (1858) ; Chili.
Larus albipennis, Licht. M.S.; Gray, List B. Brit. M. p. 173
(1844), type in Berlin Mus. (examined, H. 8.).
Chroicocephalus glaucotes, Bruch, J. f. Orn. 1853, p. 105, et 1855
p- 291.
204 MR. H. SAUNDERS ON THE LARIN&. [Feb. 5,
Gavia roseiventris, Gould, P. Z.S. 1859, p. 97 (Falkland
Islands).
Larus roseiventris, Sclater, P. Z. S. 1860, p. 391; Abbott, Ibis,
1861, p. 166 (Falkland Islands).
Gavia glaucotis, Blasius, J. f. Orn. 1865, p. 374.
Chroicocephalus glaucodes, G. R. Gray, Hand-l. Birds, iii. p. 114.
Hab. Falkland Islands, Straits of Magellan, Western Patagonia,
and coast of Chili. I caunot find any evidence of its occurrence in
Eastern Patagonia; and certainly at the Chuput river, lat. 43° S., its
place is taken by L. maculipennis.
Specimens from the Falklands seem to be smaller on the average
than Chilian examples; but there is no other point of difference.
42. Larus CIRRHOCEPHALUS, Vieill.
Larus cirrhocephalus, Vieillot, in 2nd ed. Nouv. Dict. Hist. Nat.
t. xxi, p. 502 (1818), Brazil; id. Gal. des Ois. ii. p. 223, pl. 289
(1834); Schlegel, M. P.-Bas, Lari, p. 37 (1863); Scl. & Salv. P. Z.S.
1871, p. 578; iid. Nom. Av. Neotrop. p. 148 (1873) ; Saunders, P.
Z.8. 1874, p. 292; Durnford, Ibis, 1877, p. 201.
Larus poliocephalus, Temm. M. d’Orn. ii. p. 780 (1820); Max.
v. Wied, Beitr. iv. p. 854 ; vide Salv. Ibis 1874, p. 320.
Chroicocephalus cirrhocephalus, Bruch, J. f. Orn. 1853, p. 106.
Cirrhocephalus plumbiceps, Bruch, J. f. Orn. 1855, p. 288.
“Larus maculipennis, Licht.,’’ Burm. Syst. Ueb. Th. Brasil. iii. p.
448 (1856), nec Licht. (Brazil coast, especially the small islands) ;
id. La Plata-Reise, ii. p. 518. no. 256 (1861) (R. Parana).
Gavia cirrhocephala (part.), Blasius, J. f. Orn. 1865, p. 376.
Hab. Coast of Brazil, bays and lakes, down to Rio de la Plata,
and up the Parana. Mr. Durnford did not observe it south of Buenos
Ayres. On the Pacific coast it occurs near Callao, one specimen
from Chorillos being in my own collection, and another, obtained by
M. Grec of Lima at the Chinchas Islands in 1855, being in the
Bordeaux Museum.
43. Larus PHHZOCEPHALUS, Sw.
Larus poiocephalus (sic), Swains. B. W. Afr. ii. p. 245, pl. 29
(1837) : ad. ex. believed to be the type, in Cambr. Mus. from Sene-
gambia, examined, H. S.
Xema pheocephala, Strickl. & Scl. B. of Damar., Contrib. Orn.
1852, p. 160.
Gavia cirrhocephala (part.), Blasius, J. f. Orn. 1865, p. 376.
Larus poiocephalus, Swains., Layard, B. 8. Afr. p. 368 (1867),
partim ; Barboza du Bocage, J. f. Orn. 1876, p. 293 (Cunene, Ben-
guela).
Larus pheocephalus, Hart]. Orn. W.-Afr. p. 252 (1857); id. J. f.
Orn. 1861, p. 273; Finsch & Hartl. Vég. Ost-Afr. p. 825 (1870) ;
Saunders, P. Z. S. 1874, p. 292.
Cirrhocephalus poiocephalus, Gurney, Anderss. B. Damaral. p. 358
(1872).
Hab. West Coast of Africa from the Gambia down to Walvisch
1878. ] MR. H. SAUNDERS ON THE LARIN&. 205
Bay, Damaraland, and at Lake Ngami (Chapman). I possess or
have examined specimens from the above localities. The species may
extend as far as the Cape of Good Hope and to the south-east coast ;
but of this I have no positive information.
This species is very closely allied to L. cirrhocephalus; but it is
smaller, and the bill and feet are orange-red, whilst in the larger
American species those soft parts are of a deep lake-colour ; and the
feathers in this species do not come down so close to the base of the
nostrils—differences which are quite sufficient to separate the two
forms. The African species was long confounded with L. hartlaubi
of the Cape of Good Hope, a bird belonging to a totally distinct
group, and which never has a hood at all.
44. Larus saunprrs! (Swinhoe). (Fig. 15.)
“Gavia kittlitziit, Bruch,’ Swinhoe, Ibis, 1860, p. 68 (not of
Bruch, which is described from a drawing and is L. franklini).
Larus schimperi, Schlegel, M. P.-Bas, Lari, p. 40, 1863 (not of
Bruch, 1853, nor of Bp., which =Z. franklini).
Three outer primaries of L. sawnders?, ad.
Chroicecephalus kittlitzit, Swinh. Ibis, 1863, p. 428, et P. Z.S.
p. 328.
Xema kittlitzii, David, N. Arch. Mus. Bull. vii. 1871, no. 461.
Chroicocephalus saundersi, Swinhoe, P. Z. S. 1871, pp. 273, 421,
pl. 22; David & Oustalet, Ois. de la Chine, p. 523 (1877).
Hab. The coasts of China, especially about Amoy, in winter
206 MR. H. SAUNDERS ON THE LARINA. [ Feb. 5,
(Swinhoe), and the interior waters, and those of Mongolia (David).
Nidification unknown.
This well-defined species, with which my friend the late R. Swinhoe
did me the honour to associate my name, is, to judge by the struc-
ture of its feet, an inland species or river-Gull during a great part
of the year. The tarsi are slender, the hind toe elevated and long,
and the webs of the feet are much scalloped; indeed the foot is
almost that of a Marsh-Tern. Had Bonaparte or Bruch been
acquainted with it, they would doubtless have created a genus for it.
The bill is very stout and corvine-looking ; the hood, in breeding-
plumage, is of a deep metallic black; and the pattern of the pri-
maries (see fig. 15, p. 205) is also peculiar, these being principally
white with a black bar near the tip, and a black border to the edge
of the inner web.
45. Larus minutus, Pallas.
Larus albus, Scop. Aun. i. Hist. Nat. p. 80. no. 106 (1769).
Larus minutus, Pallas, Reise Russ. Reichs, iii. p. 702, App. no.
35 (1776); Gm. Syst. Nat. i. p. 595 (1788); Pallas, Zoogr. Rosso-
As. ii. p. 331 (1811); Schlegel, M. P.-Bas, Lari, p. 42 (1863);
Sharpe and Dress. B. of Eur. pt. iv. (1871).
Larus atricilloides, Falk. Itin. iii. p. 355, t. 24, fide Gm. Syst.
Nat. i. p. 601 (1788).
Xema minutus, Boie, Isis, 1822, p. 365.
Larus @ orbignyi, Audouin, Hist. Nat. de I’ Egypte, pl. 9. fig. 3,
Expl. p. 271 (1825).
Hydrocoloeus minutus, Kaup, Nat. Syst. p. 113 (1829).
Larus nigrotis, Lesson, Tr. d’Orn. p. 619 (1831).
Chroicocephalus minutus, Eyton, Hist. R. Brit. B. p. 61 (1836) ;
Bruch, J. f. Orn. 1853, p. 105.
Gavia minuta, Macgill. Hist. Brit. B. v. p. 613 (1852).
Hab. European coasts and occasionally inland (on passage and in
winter); breeding in the marshes of Russia, and formerly in Gottland.
Middendorff obtained it in May on the Lena, to the south of Jakusk,
and as far east as the sea of Okotsk. Once in north India (Irdy) ;
North Africa to Egypt in winter.
Scopoli’s description of Z. albus applies fairly to this species, but
it is not sufficiently precise to enforce the adoption of that name to
the prejudice of a long accepted one like that of Pallas.
46. Larus PpHILADELPHLE (Ord). (Fig. 16.)
Sterna philadelphia, Ord, Guthrie’s Geogr. 2nd Am. ed. ii. p. 319
(1815), fide Lawr., B. N. Am. p. 252.
Larus minutus, J. Sabine, App. Franklin’s Polar Sea, p. 696
(1823); Sw. & Rich. F. Bor.-Am. Birds, p. 426 (1831), nec Pallas.
“Larus capistratus, Temm.,”’ Bp. Specch. Comp. p. 69 (1828),
nec Temm.
Larus melanorhynchus, Temm. Pl. Col. livr. 85, tab. 504 (1830),
Chili?
Larus bonapartii, Sw. & Rich. F. Bor.-Am., Birds, p. 425, pl. 72
1878. ] MR. H. SAUNDERS ON THE LARINZ. 207
(1831); Thompson, B. of Ireland, iii. p. 317 (1851), near Belfast ;
Schlegel, Mus. P.-Bas, Lari, p. 41 (1863).
Xema bonapartii, Bp. Birds of Eur. and N. Am. p. 62 (1838).
Chroicocephalus bonapartit, Bruch, J. f. Orn. 1853, p. 105, et
1855, p. 292.
Chroicocephalus subulirostris, Bp.; Bruch, J.f. Orn. 1853, p. 105
(type in Mainz Mus. examined, H. 8.).
Gavia bonapartii, Bp. Naumannia, 1854, p. PAGE
Gavia subulirostris, id. tom, cit. p. 213.
Fig. 16.
il 2. 3.
Three outer primaries of L. philadelphie, jr.
Chroicocephalus philadelphia, Lawr. B. N. Am. p. 852 (1858);
Dall & Bann. Tr. Chicago Ac. 1869, p. 305 (Alaska) ; Newton,
P. Z. S. 1871, p. 57, pl. iv. fig. 6, egg (Arctic America).
Gavia bonapartei, Blasius, J. f, Orns,.1865, p.371.
Larus philadelphia, Allen, Am. Nat. iii. p. 643 (Salt Lake) ;
Harting, Handbk. Brit. B. p. 172 (1872); Reid, Zoologist, 1877,
p. 489 (Bermudas).
Hab. British North America and Alaska (in summer), breeding
on the Yukon and neighbouring localities. In autumn it descends
the coasts of America, as far as California on the west and North
Carolina on the east; as a straggler it has visited the Bermudas and
also the British Islands.
The drawing (fig. 16) of the outer primaries in the young shows
wherein the pattern differs from those of allied species. In the adult
the distinctions are yet more marked.
208 MR. H. SAUNDERS ON THE LARIN&E. [ Feb. 5,
Genus Ruopostetuts, Macgill.
The distinguishing characteristic of this genus is the cuneate tail,
in which respect the sole representative species is unlike every other.
On account of this elongation of the central feathers of the tail,
some systematists have placed it next to the Stercorariine; but it
should not be inferred frem this solitary point of resemblance that
the two genera are at all closely related, their representatives being
in other respects far apart. It is much to be desired that the ster-
num of the next specimen obtained should be preserved, as I believe
this part of its structure has never been critically examined.
47. Ruoposteruia RosEA, Macgill.
Larus roseus, Macgill. Wern. Soe. Trans. v. no. xiii. p. 249 (1824),
descr. of sp. from Melville Peninsula; Jard. & Selby, Ill. Orn. vol. i.
l. xiv.
: Larus rossii, Richards. App. Parry’s 2nd Voy. p. 359 (1825),
Melville Peninsula; J. C. Ross, App. Ross’s 2nd Voy., Nat. Hist.
p- 36 (1835), Felix Harbour, Boothia ; Sw. & Rich. F. Bor.-Am. ii.
Birds, p. 427 (1831); J. C. Ross, App. Parry’s Narr. p. 195 (1828).
Rossia rosea (Macgill.), Bp. Comp. List, p. 62 (1838).
Rhodostethia rossi, Macgill. Man. Brit. Orn, pt. ii. p. 253 (1842).
Rhodostethia rossii, id. Brit. Birds, v. p. 618.
Rhodostethia roseus, Bruch, J. f. Orn. 1853, p. 106.
Rhodostethia rosea, Saunders, Ibis, 1875, p. 484 (jr.) ; Payer,
Austrian Exp. ii. p. 91 (English transl.) ; Dresser, B. of Eur. pt.
i. (1877)
Larus collaris,’’ MS., Schreibers, in Mus. Vindob.
Hab. Melville Peninsula, 693° N. lat., and Boothia, straggling to
Greenland, once to the Faroes, once to Heligoland, and (on very
questionable authority) once to England. This last specimen, which
I have examined, has every appearance of having been mounted from
a skin and not from the flesh. Ross and Parry state that it was
seen to the north of Spitzbergen in about 82° N. lat.; but they did
not obtain specimens, and no subsequent visitors te that district have
observed it; more recently Lieut. Payer says that it was obtained
about Franz-Josef Land.
There can be no doubt of the prior claim of Macgillivray’s name
for this species; but its imposition, in anticipation of that which
Richardson intended to bestow on it, gave rise to a good deal of bad
feeling at the time.
In ‘The Ibis,’ 1875, p. 484-487, I gave a description of the im-
mature plumage of this Gull from two specimens in the Mainz
Museum, and enumerated the eleven examples known to exist; to
these may be added one more in Copenhagen, and one, of which Mr.
O. Salvin has recently informed me, in the museum of Vienna: total
thirteen specimens.
In reply to inquiries respecting the Vienna example, Hr. von
Pelzeln informs me that it formed part of the collection made by
Giesecké during his seven years’ residence in Greenland, and came
1878. ] MR. H. SAUNDERS ON THE LARINZ. 209
into the possession of the Imperial Museum in 1818, when it re-
ceived from Professor Schreibers the MS. and unpublished name
of Larus collavris. In the interval between the publication of
the 4to edition of Ross’s ‘ Voyage to Baffin’s Bay’ and the later
8vo edition (both of which bear the date of 1819), Ross, or Leach
(for, although under Ross’s name, Leach was probably the real
authority) heard of Schreibers’s name, and, jumping at the con-
clusion that Schreibers’s bird was LZ. sabini, inserted the synonym
of X. collaris (Schr.) for that species in the 8vo edition. Had
Schreibers’s description been published, his name would have con-
siderably antedated the present one.
Genus Xena, Leach.
The real distinguishing character of this genus, as instituted by
Leach, is the forked tail; but the name has been improperly
employed by Boie and others for many other species. Leach,
however, defined it most clearly ; and a generic name should never
be used in any other sense than that of the founder.
48. Xema saBrinir (Sabine).
Larus sabini, J. Sabine, Tr. Linn. Soc. xii. p. 520, pl. 29 (1818);
J. Wilson, Ill. Ornith. pl. iii. (1831).
“Xema sabini, Leach,” J. Ross, App. Ross’s Voy. Baff. Bay, p.57
(1819), 4to ed. ; Steph. in Shaw’s Gen. Zool. xiii. pt. i. p. 177, pl. 20
(1826); Eyton, Rarer Brit. B. p. 54 (1836).
“‘Xema collaris (Schreibers),” Ross in App. Ross’s Voy. Baf.
Bay, il. p. 164 (1819), 8vo ed. (not in 4to ed.), nec Schreibers.
Gavia sabini, Macgill. Man. Brit. Orn. ii. p. 241 (1842).
Larus sabini, J. C. Ross, App. Ross’s 2nd Voy. p. 37 (1835).
Larus sabinii, Richardson, App. Parry’s 2nd Voy. p. 360 (1824) ;
Sw. & Richs. F. Bor.-Am., Birds, p. 428 (1831); Middendorff, Sib.
Reis., Zool. ii. p. 244, pl. xxiv. fig. 5, xxv. fig. 1 (young and egg),
(1853).
Xema sabinii, Bruch, J. f. Orn. 1855, p. 292; Lawr. B. of N.
Am. p. 856 (1860); Newton, P. Z.S. 1871, p. 57, pl. iv. fig. 5
(egg); Dresser, B. Europe, pt. xxxi. August 1874.
Larus sabinei, Schl. M. P.-Bas, Lari, p. 44 (1863).
Xema sabinei, Coues, B. of N.W. Am. p. 660 (1874-5) ; Reid,
Zoologist, 1877, p. 420 (Bermudas).
Hab. Arctic America, breeding to the north of Upernavik, in
Greenland, and then across to the west, breeding in Alaska (Dall) ;
not rare in Plover Bay, Eastern Siberia (Dall), and breeding on the
tundras of the Taimyr, north of 74° (Middendorf’). In autumn it
migrates southward ; and many specimens have from time to time
been obtained on the British coasts and those of the continent, as
far east as Holstein, and on the French coasts. Most of these are
birds of the year; but Dr. L. Bureau has an adult, with full black
hood, captured on the coast of Brittany on August 25th, 1872. In
America its southern range, as until now recorded, was down to New
York on the east, and to Great Salt Lake, Utah, on the west—
Proc. Zoou. Soc.—1878, No. XIV. 14
210 MR. H. SAUNDERS ON THE LARIN&. [Feb. 5,
roughly speaking, as far as 40° N. lat. ; but Messrs. Sclater and Salvin
have recently shown me a specimen from Prof. Steere’s collection’,
Mus. University of Michigan, labelled “ Macabi, September 1872,”
and on the card-label ‘‘ Tumbez,’’ Macabi Island being a little to the
north of Huanchacho on the coast of Peru, in nearly 8°S. lat.—an
enormous extension of its previously known limits. It is in adult
plumage, with the exception of some dusky feathers on the nape;
the forehead and head are white and devoid of the hood, which it
was reasonable to suppose would only be worn during the breeding-
season, although this is the first absolute proof I have had of this;
the bill as in adults; the tail white and well-forked; feet rather
faded. I imagine this to be a bird of the second year. This species
has once occurred at the Bermudas.
The name of X. collaris was applied to this species as a synonym
in the 8vo edition of Ross’s ‘ Voyage’ (1819), under the erroneous
impression that this was the LZ. collaris to which Prof. Schreibers,
then Director of the Vienna Museum, had given that MS. and un-
published name; but as a matter of fact that name belongs to an
example of Rhodostethia rosea. For the elucidation of this point I
am indebted to the kindness of Herr A. von Pelzeln, who also in-
forms me that the Vienna Museum is the possessor of what is pro-
bably the oldest specimen of this species in any European collection,
it having been received, without any published name or description,
from the Ornithological Institution of that city about 1806.
49. Xema FuRcATUM (Neboux).
Mouette a queue fourchue, Neboux, Rev. Zool. 1840, p. 290.
Larus furcatus (Neboux), Voy. ‘ Vénus,’ Atlas, pl. x. (1846);
Prévost & Des Murs, Voy. ‘ Vénus,’ v. Ois, p. 277 (1855).
Xema furcatus, Bruch, J. f. Orn. 1853, p. 103.
Creagrus furcatus, Bp. Naumannia, 1854, p. 213; Bruch, J. f.
Orn. 1855, p. 292; Salvin, Tr. Zool. 8. ix. p. 506 (Galapagos).
Hab. The type of this species in the museum of Paris is said to
have been obtained at Monterey, California, in November, by Dr.
Neboux of the French frigate ‘ Vénus.’ The only other specimen
known to exist is in the British Museum, and was obtained during the
voyage of H.M.S.’s ‘ Herald’ and ‘ Pandora,’ at Dalrymple Rock,
Chatham Island, Galapagos group. The ‘ Herald’ appears to have
visited Chatham Island between the 11th and 16th January, a date
which is worth bearing in mind, as the British-Museum specimen
seems to be in fully adult plumage ; the grey tint which pervades the
lower part of the neck and breast in the Paris specimen, and which is
probably a sign of comparative immaturity, having disappeared, leaving
the hood well defined. But for the absence of the distinct collar at
the bottom of the hood, and the narrow white band of feathers at
the base of the upper mandible, this bird might be briefly described
as a gigantic Sabine’s Gull, the tail being rather more forked in
proportion than in that species. It is certainly remarkable that, in
spite of the researches of the American naturalists, no other speci-
1 See aboye, p. 141.
1878. ] MR. H. SAUNDERS ON THE LARIN&. 211
men of this Gull should ever have been obtained in any part of
California; and Mr. Salvin inclines to think the Californian locality
an erroneous one, in view of similar mistakes known to have been
made with the birds collected during the voyage of the ‘Vénus.’ It
is on the other hand somewhat strange that if the head quarters of
this Gull, of which the appearance is sufficiently striking to attract
attention, are in the Galapagos group, none of the other visitors or
naturalists should have brought any news of it.
Creagrus is one of Bonaparte’s arbitrary and undefined genera ; and
there seems to be no structural difference to warrant the generic
separation of this species from Xema sabinii.
In this paper I have from time to time acknowledged with much
pleasure the assistance I have received from my friends and cor-
respondents ; and I have now to render my especial thanks to the
authorities of the British and Cambridge Museums, and of the
Smithsonian Institution, Washington, and to Dr. Brewer of Boston,
to Professor Peters of Berlin, Professor Reinhardt of Copenhagen,
M. Oustalet, M. Bouvier, Mr. O. Salvin, Mr. H. Seebohm, .Mr.
Harvie-Brown, Mr. Gervase Matthew, R.N., and Mr. E. Hargitt,
for the opportunities they have severally afforded me of examining
examples of rare species and also series of specimens.
P.S. (April 1st).—I take this Opportunity of correcting an error
in my paper on the Sterning (P. Z. 8.1876, p. 671) where two
species are united under the name of Anous ceruleus. At that
time had not geen a Specimen of the real A. ceéruleus, Bennett ;
and that species and 4. cinereus, Gould, are so very much alike
that, until examples were available for comparison, the descriptions
and plates might easily have been taken to refer to the same species
in different stages of plumage. Seen by the light of further expe-
rience they appear to be distinct ; and the following is their synonymy
and habitat :—
ANOUS CHRULEUS (Bennett).
Sterna cerulea, F. D. Bennett, Narr. Whaling-Voy. round Globe,
li, App. p. 248 (1840), Christmas Island, Pacific.
“Sterne cendré,” Neboux, Rev. Zool. Oct. 1840, p. 291, et
Stolida cinerea. id. Voy. ‘ Vénus,’ Atlas, pl. 9 (1846); nee Anous
cimereus, Gould, P. Z. 8.1845 (Pacific, N. of Equator),
Sterna teretirostris, Lafresnaye, Rev. Zool. 1840, p. 242, et
Procelsterna tereticollis, id. Mag. de Zool. 1842, pl. 29: no lo-
cality; both described from a single specimen purchased from a
dealer at Havre.
Anous parvulus, Gould, P. Z. S. 1845, p. 104 (deseribed from a
Specimen obtained by Bennett at Christmas Island); Cassin, U.S,
Expl. Exp., Birds, p. 393 (Honden Island, Low archip, 14° S.,
138° W.).
Megalopterus plumbeus, Peale, U.S. Expl. Exp. p. 285 (1848),
Honden Island.
14*
212 MR. P. L. SCLATER ON CASUARIUS SALVADORII. [Feb. 19,
Anous cinereus (Neb.), Finsch and Hart]. F. Central- Polynesiens,
p- 239 (1867), Pheenix group, 3° 8’S., 171° W., nee Gould.
Hab. Pacific, from a little north of the Equator; Christmas Island;
the Ellice group, 9° S., 179° E., whence there are two specimens in,
the British Museum recently obtained by the Rev. S. J. Whitmee ;
the Pheenix group; and Honden Island, Low archipelago.
ANOUS CINEREUS, Gould.
Pelecanopus pelecanoides, G. R. Gray, L. Birds Brit. M. iii. p. 180
(Australia, presented by Sir T. Mitchell).
Anous cinereus, Gould, P. Z.S. 1845, p. 104 (N.E. Australia) ;
id. B. Australia, vii. pl. 76 (1848), Norfolk I. and N.E. coast Aus-
tralia.
Procelsterna albivitta, Bp. Compt. Rend. xlii. 1856, p. 773 ; Gould,
Hand-b. B. Austr. ii. p. 420 (1865); Gray, Hand-l. ili. p. 128
(1871).
Sterna cinerea, Schlegel, M. P.-Bas, Sterne, p. 38 (1863), Aus-
tralia.
Anous albivittatus, Fisch, P. Z. 8S. 1877, p. 776 (Eua, Friendly
group).
Hab. Norfolk Island; N.E. Australia and the Tonga or Friendly
group, in about 22°S., 175° W. It is presumably the species
observed by Mr. E. L. Layard in the Fiji group.
The range of these two species appears to be nearly parallel, that
of A. ceruleus bemg the more northerly. A. ce@ruleus is smaller
than A. cinereus, Gould, and is darker all over, especially on the
underparts, which are blue-grey, whereas in 4. cinereus they are
nearly white. The differences are too great to be explained away as
being due to age, and I admit the distinctness of the two species ;
but they are very closely allied. The fact of their being found in
such close proximity within so limited an area is very remarkable.
February 19, 1878.
Prof. Mivart, F.R.S, V.P., in the Chair.
Mr. P. Geddes read a memoir on the mechanism of the odonto-
phore in certain mollusca. In this paper the view of Cuvier that
the movements of the radula depend upon those of the under-
lying cartilages was substantially revived. Arguments were adduced
against the more recent theory of Prof. Huxley that it runs like a
chain-saw, the cartilages merely forming a pulley-block. The
use of bacteria as food by Limneus was also described in the
memoir, which will be published in the Society’s ‘ Tranasctions.’
Mr. Sclater exhibited the skin ofa fine adult Cassowary, which had
recently been acquired for the collection of the British Museum.
The specimen was labelled ‘‘ Wandammen, May, 1876,” and was
1878.] MR. P.L. SCLATER ON CASUARIUS SALVADORII. 213
Head of Casuarius salvadorii.
214 MR. F. DAY GN FEAR AND ANGER IN FISHES. [Feb. 19,
believed to have been obtained by the collectors of Herr Bruijn, of
Ternate. Wandammen is situated on the western coast of the
Bay of Geelvink, near the southern extremity’.
Mr. Sclater stated that the only Cassowary of the present form
known from this locality was the species shortly indicated by Bec-
cari in his ‘‘ Lettera Ornitologica” (Ann. Mus. Civ. Genova, vii-
p- 717) as Casuarius tricarunculatus’; but there were no traces
whatever of a third wattle in the present specimen.
The form of the casque in the present bird was nearly that of C.
australis, the summit being elevated high above the head, and much
compressed laterally, but forming almost a point at the summit
instead of a longitudinal ridge (see fig., p. 213). The wattles were
two, one on each side of the median line, closely approximating at
their bases, but divided down nearly to their origin, and about three
inches in length.
The species, which Mr. Sclater considered to be new to science,
must, he said, be placed in the first section of his arrangement as
given in P. Z. 8. 1875, p. 87, next to C. galeatus, from which the
form of the helmet distinguished it. From C. Jeccarii of the Arru
Islands it might be known by the more completely divided neck-
wattles.
The whole length of the body of the bird, from the tip of the
casque to the tip of the tail, was about five feet ; the tarsus measured
12°5 in., the middle toe 85 in. The distance from the commis-
sure of the mandible to the top of the casque was 7°5in., the dis-
tance from the gape to the extremity of the bill 7°5 in.’
A communication was read from Mr. F. Day, F.Z.8., containing
the following remarks on Mr. Whitmee’s paper on the manifesta-
tion of fear and anger in Fishes, read at the last meeting.
** At the last meeting of the Society (February 5th), Mr. Whitmee
read a paper on the manifestation of fear, anger, &c., by Fishes,
observing that a recent author, ‘On the Expression of the Emo-
tions,’ had entirely omitted allusion to this class of Vertebrates,
whilst, as regards anger, he believed that no observations had been
recorded. I propose offering a few remarks upon the foregoing,
as well as upon some other subjects touched upon by Mr. Whitmee.
«The means of expression in animals adverted to by Mr. Darwin
(excluding those of the ears, which would be out of place in
fishes) are :—sounds, vocally or otherwise produced; the erection
1 See the chart of Geelvink Bay in ‘Cosmos,’ yol. viii. tabb. 3 & 10.
2 The only known specimen of C. tricarunculatus was, as Count T. Salvadori
kindly informs me, still living in Ternate in July last, where it was seen by
M. Laglaize.
3 [Since these remarks were made I find that an example of this same
Cassowary was exhibited by M. Oustalet at a meeting of the Association Scien-
tifique de France on February 23rd, and the species named C. salvadorii (see
Bull. Ass. Se. de France, no. 539, vol. xxi. p. 649). I have therefore withdrawn
my name in favour of that of M. Oustalet. I think it probable that the Casso-
wary from Southern New Guinea, formerly living in the Society's Gardens,
which I referred (P. Z. 8. 1875, p. 527) with some doubt to C. beecarit. is a
younger individual of this same species,—P. L. 8. |
1878. | MR. F. DAY ON FEAR AND ANGER IN FISHES. 215
of dermal appendages under the influence of anger or terror, which
last would be analogous to the erection of scales and fin-rays in the
class Pisces. Regarding special expressions, as those of joy, pain,
astonishment, &c., we could hardly expect to find such so well
marked in fishes as in some of the higher animals, in which the
play of the features often affords us an insight into their internal
emotions. Eyes’ destitute of movable eyelids, cheeks covered
with scales, or the head enveloped in dermal plates, can scarcely
mantle into a smile or expand into a broad grin. We possess,
however, one very distinct expression in fishes, which is absent or
but slightly developed in many* of the higher animals—namely,
change of colour. All are aware that when a fish sickens its
brilliant colours fade, but less so how its colour may be aug-
mented by anger, and a loss of it be occasioned by depression
the result of being vanquished by a foe. Some forms also emit
sounds when actuated by terror, and perhaps in times of anger ; but
of this last I possess no decided proofs.
“As regards manifestations of anger, Dr. Cantor makes the
following remarks on the Macropodus pugnaz * :-——‘ When the fish
is in a state of quiet, with the fins at rest, the dull colours present
nothing remarkable. But if two are brought within sight of each
other, or if one sees its own image in a looking-glass, the little
creature becomes suddenly excited, the raised fins and the whole
body shine with metallic colours of dazzling beauty, while the pro-
jected gilt membranes, waving like a black frill round the throat,
add something grotesque to the general appearance. In this state
it makes repeated darts at its real or reflected antagonist. But
both, when taken out of each other’s sight, instantly become quiet.
. . . . The Siamese are as infatuated with the combats as the Malays
are with their cock-fights, and stake considerable sums, and some-
times their own persons and their families. The license of exhibit-
ing fish-fights is farmed, and affords a considerable annual revenue
to the King of Siam.’
“The foregoing extract shows anger in fishes demonstrated by
change in colour and the erection of the fins and gill-membranes. I
will now give an observation respecting the three-spined Stickle-
back*. After a fight between two examples ‘a strange alteration takes
place almost immediately in the defeated party : his gallant bearing
forsakes him ; his gay colours fade away ; he becomes again speckled
and ugly; and he hides his disgrace amongst his peaceable com-
panions, who occupy together that part of the tub which their
tyrants have not taken possession of ; he is, moreover, for some
time the constant object of his conqueror’s persecution.’ In fact we
here perceive how the disgrace of defeat affects the spirits of the
1 Couch (Ilustrations, &e., p. 305) says, “ The faculty of giving forth brilliant
light from the eyes, as in a cat, is said to have been observed by fishermen in
the Blue Shark.”
2 Numerous examples of the Saurians are exceptions.
3 Cantor, Catal. Mal. Fish, 1850, p. 87. Sir J. Bowring, in his account of
Siam, p. 155, gives a very similar account of the battles of these fishes.
* Couch, ‘ British Fishes,’ 1865, vol. iv. p. 172.
216 MR. F. DAY ON FEAR AND ANGER IN FISHES. [Feb. 19,
vanquished ; this reacts on his health, and as a result his brilliant
hues fade away. The conqueror, on the other hand, exulting in
his victory becomes more resplendent ; he does not forget his former
triumph, and considers it no disgrace to occasionally lord it over his
beaten foe.
‘Fear is shown by fish in many ways. ‘When hooked or netted
they sometimes empty their stomachs by an instinctive act of fear,
or to facilitate escape by lightening their load? (Owen, Comp.
Anat. i. p. 419). There is not an angler unacquainted with the
natural timidity of fishes, nor a keeper in charge of a salmon-pass,
who does not know how easy it is for poachers to deter the salmon
from venturing along the path raised expressly for his use.
« Amongst the coral-reefs of the Andaman Islands I found the
little Heliastes lepidurus abundant. As soon as the water was
splashed they appeared to retire for safety to the branching coral,
where no large fish could follow them; so frightened did they
become, that on an Andamanese diving from the side of the boat,
they at once sought shelter in the coral, in which they remained
until it was removed from the sea. In Burma I observed, in 1869,
that when weirs are not allowed to stretch across the rivers (which
would impede navigation), the open side as far as the bank, is
studded with reeds; these, as the water passes over them, cause
vibration, and occasion a curious sound alarming the fishes, which,
crossing to the weired side of the river, become captured.
“Under the influence of fear or anger the well-known Climbing
Perch of India (Anabas scandens) not only erects its spiny-rayed
fins and its gill-covers, but also the scales on its body, even down to
those along the base of the caudal fin’; this to a less extent,
pethaps, appears common to spiny-rayed forms.
“Hooker, alluding to Gulls, Terns, Wild Geese and Pelicans in the
Ganges valley, observes, ‘These birds congregate by the sides of
pools and beat the water with violence, so as to scare the fish,
which then become an easy prey—a fact which was, I believe, first
indicated by Pallas during his residence on the banks of the Caspian
Sea’?, Fishes, under the influence of terror, dash about with their
fins expanded, and often run into places which must destroy them.
Thus droves and droves of Sardines in the East, impelled by the
terror of pursuing Sharks, Bonitoes, Seir, and other voracious fishes,
frequently throw themselves on the shores in enormous quantities.
Friar Odoric, who visited Ceylon about 1320, says, ‘There are fishes
in those seas that come swimming towards the said country in
such abundance, that for a great distance into the sea nothing can
be seen but the backs of fishes, which, casting themselves on the
shore, do suffer men for the space of three daies to come, and to
take as many of them as they please, and then they return again
into the sea’ °.
“Fishes frequently show distinet signs of affection. Yarrell
(‘British Fishes’) mentions how a person who had kept two
1 | Rishes of Malabar’ (1865), p. 133.
2 Himalayan Journals, yol. i. p. 80. > Hakluyt, vol. ii. p. 37.
1878. ] MR. F, DAY ON FEAR AND ANGER IN FISHES. 217
small ones together in a glass vessel gave one away; the other
refused to eat, and showed evident symptoms of unhappiness until
his companion was restored: Pennant, how the River-Bullhead
‘deposits its spawn in a hole it forms in the gravel, and quits it
with great reluctance: General Hardwicke, how the Gouramy
(Osphromenus olfax), in the Mauritius, forms a nest amongst the
herbage growing in the shallow water in the sides of tanks. Here
the parents continue to watch the place with the greatest vigilance,
driving away any interloping fish. The amphibious walking fish of
Mysore (Ophiocephalus striatus) appears to make a nest very
similar to that of the Gouramy, and over it the male keeps guard ;
but should he be killed or captured, the vacant post is filled by his
partner (Colonel Puckle). When very young the fishes keep witb,
and are defended by their parents, but so soon as they are suffi-
ciently strong to capture prey for themselves they are driven away
to seek their own subsistence (see ‘ Fishes of India,’ p. 362). But
it is not only these monogamous amphibious fishes which show an
affection for their eggs and also for their fry, but even the little
Etroplus maculatus has been observed to be equally fond of its ova.
‘The eggs are not very numerous, and are deposited in the mud at
the bottom of the stream, and, when hatched, both parents guard
their young for many days, vigorously attacking any large fish that
passes near them’ ’*.
** Although the proceedings of the members of the marine and
estuary genus Arius and its allies show not quite so distinctly signs
of affection, still it must be a well-developed instinct which induces
the male to carry about the eggs in his mouth until hatched, and to
remove them in this manner when danger is imminent. I have taken
the ova just ready for the young to come forth out of the mouth and
fauces of the parent (male) fish ; and in every example dissected there
was no trace of food in the intestinal tract.
“At many temples in India fishes are called to receive food by
means of ringing bells or musical sounds. Carew, in Cornwall, is said
to have called his Grey Mallet together by making a noise like chop-
ping with acleaver. Lacépéde relates that some fishes, which had
been kept in the basins of the Tuileries for more than a century,
would come when called by their names, and that, in many parts of
Germany, Trout, Carp, and Tench are summoned to their food by
the sound of a bell. These instances are doubtless mostly due to
the fishes having learnt by experience that on the hearing certain
sounds they may expect food. But Lacépéde mentions that some
were able to distinguish their individual names; and the same occurs
in India. Lieutenant Conolly remarked upon seeing numerous fishes
coming to the Ghaut at Sidhnath to be fed when called; and on
‘expressing our admiration of the size of the fish, “‘ Wait,” said a
bystander, “until you have seen Raghu.” The Brahmin called out
his name in a peculiar tone of voice; but he would not hear. I
threw in handful after handful of ottah (flour) with the same
success, and was just leaving the ghaut, despairing and doubting,
* Jerdon, ‘Madras Journal of Literature and Science,’ 1849, p. 143.
218 MR. F. DAY ON FEAR AND ANGER IN FisHES. [Feb. 19,
when a loud plunge startled me. I thought somebody had jumped
off the bastion of the ghaut into the river, but was soon undeceived
by the general shout of “ Raghu, Raghu,” and by the fishes, large
and small, darting away in every direction. Raghu made two or
three plunges, but was so quick in his motions that I was unable to
guess at his species’’.
« Although I have alluded to an increased brilliancy of colour being
caused by anger, and a dull hue occasioned by terror or illness, there
appears to be still another cause at work. During the breeding-
season some species, such as Periophthalmus schlossert, appear to
become more brilliant, especially the males, as I have observed in
my report on the freshwater fishes of Burma.
“ Pallegoix states that in Siam the Dog’s-tongue is a fish shaped
like a Sole; it attaches itself to the bottom of boats, and makes a
sonorous noise, which is more musical when several are stuck to
the same boat and act in concert (vol. i. p. 193). These noises
can scarcely be due to anger or fear. Sir J. Bowring (vol. i.
p- 276) also remarks upon having heard this fish, ‘which sticks
to the bottoms of the boats, and produces a sound something like
that of a Jew’s harp struck slowly, though sometimes it increases in
loudness, so as to resemble the full sound and tones of an organ.
My men have pointed me out a fish about four inches long as the
author of the music.’
‘««Some years since, at Madras, I obtained several specimens of a
freshwater Siluroid fish (Macrones vittatus) which is termed ‘ the
fiddler’ in Mysore. I touched one which was on the wet ground,
at which it appeared to become very irate, erecting its dorsal
fin, making a noise resembling the buzzing of a bee, evidently a
sign of anger or terror. Having put some small Carp into an
aquarium containing one of these fishes, it rushed at a small example,
seized it by the middle of its back, and shook it like a dog killing a
rat ; at this time its barbels were stiffened out laterally like a cat’s
whiskers”.
“Many fish, when captured, make noises, perhaps due to terror.
Thus the Caranx hippos, Tetrodon, and others grunt like a pig.
Darwin (Nat. Journ. vol. vii.) remarks on a Silurus found in the
Rio Parana, and called the Armado, which is remarkable for a harsh
grating noise when caught by hook and line; this noise can be
distinctly heard when the fish is beneath the water.
‘« Aristotle and Ailian were aware of this faculty in some of the
fishes of the Mediteranean *.
“The Cuckoo-Gurnard (Trigla pini) and the Maigre (Sciena
aquila) utter sounds when taken out of the water*; and Herrings,
when the net has been drawn over them, have been observed to do
1 “Observations on the Past and Present Condition of Onjein,” ‘ Journal
of the Asiatic Society of Bengal,’ vi. p. 820.
? «Fishes of India,’ p. 449.
3 See ‘De Animal.’ lib. iy. cap. ix.; lian, lib. x. cap. xi.; Pliny, lib, ix.
cap. Vili. and lib. xi, cap. xviii. ; Atheneeus, lib. vii. cap. i. & vi.
4 Yarrell, ‘ British Fishes,’ i. pp. 44-107.
1878. | MR. F. DAY ON FEAR AND ANGER IN FISHES. 219
the same: ‘this effect has been attributed to an escape of air from
the air-bladder; but no air-bladder has been found in the Coftus,
which makes a similar noise.’
«The Lesser Weaver buries itself in the loose soil at the bottom of
the water, leaving only its head exposed, and awaits its prey. If
touched, it strikes upwards or sideways; and Pennant says it
directs its blows with as much judgment as a fighting-cock (Yarrell,
vol. i. p. 26). Fishermen assert that wounds from its anterior
dorsal spines are more venomous that those caused by the spines on
its gill-covers.
* As regards fighting, I should suppose that, unless some portion
of the body is peculiarly adapted for this purpose, as the rostrum of
the Sword-fish, or the spine on the side of the tail in the Lancet-
fishes, we must chiefly look to the armature or covering of the
jaws for weapons of offence.
“And this naturally leads us on to ask if, as suggested by Mr.
Whitmee, ‘the chief purpose served by fishes’ spines is protec-
tion against the attacks of those of the class which are carnivorous.’
The first inquiry is, Do the carnivorous forms most abound amongst
Acanthopterygians or Malacopterygians? The spiny-rayed forms
(speaking as a whole) have the teething far better developed than
the soft-rayed Carps of the fresh waters, or the Herrings of the sea,
the former (spiny-rayed forms) being the most carnivorous, the
latter (or soft-rayed fish) being preyed upon by the former, and by
the Elasmobranchii.
TI observed’, ‘ It may not be amiss to point out that the Acantho-
pterygian or spiny-rayed fishes appear to be most numerous in the
ocean (preying upon their articulated-rayed neighbours, the Clupeidee,
&c.); but as we examine waters more inland, the Salmonide or Cypri-
nidee usurp their place, these latter not being possessed of spinate,
but merely articulated rays. A maritime residence appears most
adapted for the Acanthopterygian or spiny-rayed fishes, a fresh-
water inland one to the Malacopterygians or spineless forms.’
**Spinate dorsal and anal fins being much more frequent in the
carnivorous and voracious forms than in their weaker neighbours
leads me to suppose that they must have some other function than
protection of the fish from its enemies. One of these is probably
to guard the fin-membrane from injury, for which purpose spines
are much better adapted than rays in fast-swimming species. One
has only to witness how commonly the spineless anal fins of Clupeoids
and Siluroids are found injured in the tropics, to feel sure that much
of this is owing to the absence of spinate protection. We do not
see the same injury existing to any thing like a similar extent in
Teleosteans whose fins are armed with strong spines; but in the
intermediate forms, as the first dorsals of Sier-fishes (Cyécum), the
weak spines of the fins are frequently injured.
““Mr. Whitmee remarked that in serious attacks fish ‘always
turned suddenly round and lashed at one another with the caudal
fin ;’ and he continued that he believed serious fighting is always
«Journal of the Linnean Society, Zoology,’ vol. xiii. p. 111.
220 MR. F. DAY ON FEAR AND ANGER IN FISHES. [Feb. 19,
done with the tail. ere I am unable to concur. Fish, the same
as other animals, when they commence to fight, employ that portion
of the body most suited to such a purpose. Thus the Perch will
employ his teeth, the Sword-fish his elongated snout, the Lancet-
fish the spine at the side of its tail, the Siluroid may use his dorsal
or pectoral spine. Instances of all these modes of attack may be
referred to.
‘Continuing Couch’s illustration of the Stickleback, he observes,
‘The bite of these little furies is so severe that I have frequently
known it, when inflicted on the tail, produce mortification and,
consequently, death. They also use their lateral spines (ventral
fins) with such fatal effect that, incredible as it may appear, I have
seen one during a battle absolutely rip his opponent quite open, so
that he sank to the bottom and died.’
“That the Sword-fish employs his sword-like projection for this
purpose is well known ; frequent examples occur of his driving that
formidable weapon into ships, whilst, according to Swainson, he is
very fierce, and attacks a Whale whenever he encounters one ; other
observers, however, represent the Sword-fish as gentle and inoffensive
except to Whales.
“The Lancet-fish as it swims past its enemy tears up an open wound.
“« As regards the Siluroids we have more than one mode of attack;
but I do not know of any Siluroid that employs its tail for this
purpose. In one of the Siluroids of the Ohio ‘the first ray of the
dorsal is formed of a very strong, sharp spine, which the animal uses
to kill others of a smaller size ; for this purpose it gets beneath the
fish it intends to attack, and then, suddenly rising, wounds it re-
peatedly in the belly.’
‘“‘Mr. Whitmee supposes that most carnivorous fish capture their
prey by outswimming them; but to this there are numerous excep-
tions. The Angler, or Fishing Frog (Lophius piscatorius), ‘ while
crouching close to the ground, by the action of its ventral and
pectoral fins, stirs up the sand and mud; hidden by the obscurity
thus produced, it elevates its appendages (situated on the upper
surface of the head), moves them in various directions by way of
attraction as a bait, and the small fishes approaching either to
examine or to seize them, immediately become the prey of the
Fisher’ (Yarrell). In India we find a freshwater Siluroid (Chaca
lophioides) which ‘ conceals itself among the mud, from which, by
its lurid appearance, and a number of loose filamentous substances
on its skin, it is scarcely distinguishable; and with an immense
open mouth it is ready to seize any small prey that is passing along’
(Ham. Buchanan). In March 1868 I obtained a fine example of
Ichthyscopus inermis (‘ Fishes of India’ p. 261), which I placed
in water having a bed of mud: into this it rapidly worked itself,
first depressing one side and then the other, until only the top of
its head and mouth remained above the mud, whilst a constant
current was kept up through its gills. It made a noise, half snap-
ping and half croaking, when removed from its native element.
This sound I consider most probably due to fear. Some fish, in
1878. ] MR. F. DAY ON FEAR AND ANGER IN FISHES. 221
short, invariably seize their prey with their mouths, and that without
calling the caudal fin into play. In fact, a stroke with the tail appears
sometimes to denote contempt in fishes; it is not rare that anglers
find fishes sometimes swim up to their bait, which they not only
refuse, but, giving it a lash with their tail, decline to rise any
more. This may, however, be a symptom of curiosity, which is
largely developed in the finny tribe.
“I might multiply instances from many authors, but consider those
adverted to are sufficient to show that various ichthyologists have
remarked upon the emotions of anger and terror in fishes being
shown by the erection of their dermal appendages and gill-covers,
as well as by changes of colour, whilst terror induces some species
to emit sounds that are not commonly perceived; that fishes some-
times show affection for their partners in captivity, mourning their
removal ; that they may be tamed sufficiently to come to a recognized
sound, even to individual names that have been bestowed upon them ;
and that some species have an instinctive affection for their eggs and
young, which they guard against intruders with the greatest deter-
mination.
‘At the present time, in the Royal Westminster Aquarium, is a
live example of the Electric Eel (Gymnotus electricus) which has
in its electric organs the means of showing when it is affected by
anger or terror. Some consider this curious property is for protec-
tion against Alligators ; it is certainly used against fishes for the
purpose of obtaining food ; but when we remember how, when the
Indians drive in horses and mules to the waters infested by the Eels,
they immediately attack them, we must admit that such cannot be
for the purpose of preying upon them, but is due to anger or terror
at being disturbed.”
Mr. Whitmee being unable to attend, the Secretary read the
subjoined reply to Mr. Day’s remarks.
“* By the courtesy of the Secretary I have seen Mr. Day’s comments
on my paper. As I cannot attend the meeting to-morrow evening,
I crave the liberty of presenting two or three written observations.
“‘1. My paper was written in Samoa in 1875; and my position
there, of course, prevented me from having access to the whole
literature of this subject. I was aware of the conduct of the Stickle-
back in guarding its nest, and also of similar conduct in some other
fishes. But my object was to show, in opposition to a view quoted,
that fishes, as a class, manifest as much feeling as most other
animals. In stating that I had not met with observations showing
this, I did so as an excuse for presenting a paper which I feared
possessed little intrinsic value. I am glad the paper has led Mr,
Day to bring forward so much evidence in confirmation of the view
I advocated.
«2. Mr. Day gives a more general application than I intended to
my observations about the mode of fighting with the tail. I stated
an observed fact, viz. that certain fishes in my aquarium, when
fighting, lashed at each other with their tails. From this I inferred
222 MR. A. H.GARROD ON TOLYPEUTES TRICINCTUs. [Feb. 19,
that such a mode of attack might be common among most, not all,
fishes. Of course, those which possess more effective weapons in
another part of the body would not resort to such a mode of
warfare.
«*3. As to the way in which carnivorous fish capture their prey, I
said, ‘doubtless most do it by outswimming them,’ and not that ad/ do
this. In P.Z.S. for 1875, p. 545, I mentioned an observed fact of
an Antennarius multiocellatus angling for small fish. Of course no
one would ever imagine that the slow-moving fish mentioned by
Mr. Day would outswim their prey.”
1. Notes on the Anatomy of Tolypeutes tricinctus, with
Remarks on other Armadillos. By A. H. Garrop,
M.A., F.R.S., Prosector to the Society.
[Received January 14, 1878.]
The Society purchased, on the 3rd of October 1877, a young
female living specimen of the Apar (Tolypeutes tricinctus), which
was the first example of the species exhibited alive in the Gardens,
and probably the first ever brought alive to this country. It was in
bad health on its arrival, and, never recovering, died, without any
visible lesion, on the 27th of December following. Its death has
given me an opportunity of determining some of the most important
points in its anatomy, which may be accepted as a supplement to Dr.
Murie’s valuable and elaborate memoir on T'olypeutes conurus in the
‘Transactions of the Linnean Society’ *
The measurements of the specimen under consideration are as
follows :—
inches
From tip of nose to base of tail........ 12°55
Lienpthinae head wot hora t ss aed Poe ees 2°8
Lengthinfstailtindte: alt: Sh tara 2°1
from which it is evident, on comparison with the table of measure-
ments of the specimens in the national collection given below, that
the individual was not adult, but fairly grown.
The differences between the two known species of the genus Toly-
peutes are so slight that it may be worth while referring to them
before going further. It is to M. Geoffroy Saint-Hilaire that we
owe the discovery of the second species (7. conurus) ; and his most
lucid description is to be found in the ‘Comptes Rendus’ for 1874?.
Therein the history of the Apar is fully expounded, references being
given to all previous important accounts of the animal.
It is in the central portion of the cephalic shield that the most
important peculiarities are to be seen. ‘The marginal plates of the
posterior two thirds of this shield form a regular series, and enclose
other larger plates—namely, a posterior median plate, followed ante-
© Volkiscex, pn 71: 2 Vol. xxiv: p. 572.
1878.] MR. A. H. GARROD ON TOLYPEUTES TRICINCTUS. 223
riorly in 7. tricinetus by a pair of plates, in front of which, again, is
another smaller pair.
In T. conurus the posterior median plate is followed by a single
larger plate, and that by a pair of plates the transverse breadth of
which is greater than that of the second median plate behind and
in contact with it. In fig. 1, a and 4 exhibit these features, as well
as their effect in causing the characteristic difference in the general
shape of the cephalic shield of the two species—that in T. tricinctus
being triangular, with its greatest breadth opposite the posterior of
the pairs of plates, that in T. conurus being oval, with its greatest
breadth opposite the single pair of plates.
a. T. tricinctus. b. T. conurus. - e. T. muriei.
Cephalic shields of Tolypeutes.
On referring to the beautiful illustrations given by Dr. Murie of
the animal dissected by him', the cephalic shield of which is here re-
produced in outline (fig. 1, ¢), it is evident that it does not corre-
spond with either of those above described; and I may mention that
there is a skin in the national collection (Brit. Mus. spec. 140 a)
which agrees with it. In these the posterior median plate is followed
by a second larger median plate, as in 7. conurus, and that, more
forward, by a small median plate in association with two small lateral
plates. The difference makes me feel justified in establishing a third
species of Tolypeutes, based upon Dr. Murie’s figure, together with
_ the skin in the national collection above referred to, for which I
would suggest the name Tolypeutes muriei, after the author who has
* Traus. Linn, Soc. vol. xxx. pl. xxxi. fig. 3,
224 MR. A. H.GARROD ON TOLYPEUTES TRICINCTUS. [| Feb. 19,
so well described its structure. Dr. Murie’s figure, being based upon
a photograph of the specimen, is thoroughly reliable.
M. Is. Geoffroy St.-Hilaire has so well differentiated 7’. tricinctus
and 7’. conurus that it will be necessary for me only to mention that
in the former there are five toes on the fore feet (as in the Society’s
specimen in my possession), whilst in 7’. conurus, following the
account of the discoverer of the species, there are ‘‘ trois doigts seu-
lement, avec le rudiment d’un quatriéme, aux pattes antérieures (ce
rudiment represente le doigt interne). Point de doigt externe.”
The same remarks apply to 7’. muriei; for in the specimen in the
British Museum there are four toes on each fore foot, whilst in Dr.
Murie’s example the pollex was not present.
The tail, with its infundibuliform armature, is distally covered
with four pairs of small rows of plates, arranged in longitudinal lines,
there being one superior and one inferior pair, one supero-lateral and
one infero-lateral. In 7’. tricinctus the whole organ is flattened from
above downwards; in Z’. conurus and 7’. muriei it is not so. Two
and a quarter inches appear to be its extreme length along its dorsal
curve in all but the largest individuals, where it may reach two and
a half inches. I do not find that the different species differ in the
length of this appendage, which is correlated, as far as its length
goes, with the length of the head, on the right side of which it
always lies when the animal is rolled up'.
I can find no important differences among the species in the ante-
rior portion of the carapace. In the three free transverse semizones
T. conurus and T', muriei agree, and differ from T’, tricinctus in that
the terminal or marginal lateral scutes of each zone (which with the
scute above on each side of each zone are less tuberculated than the
rest, to reduce the friction when the animal rolls itself up) are more
detached from the second scute, are more rounded, and are smaller
proportionately.
In the posterior moiety of the carapace of 7’. conurus and T. mu-
rie. fifteen, sixteen, seventeen, or eighteen rows of scutes can be
counted along the middle line from before backwards if the marginal
scutes are included in the numeration; in 7’. évicinctus I never find
more than thirteen or fourteen rows.
In 7. tricinctus only there is a triangular area composed of small
scutes, with its apex directed downwards, and basally in contact
with the cephalic shield, which interpolates itself between the eye
and ear on each side.
The following measurements of the specimens of the different
species in the national collection may prove of service in determining
the proportions of each. They demonstrate that the head is decidedly
shorter than the anterior moiety of the carapace in 7’. éricinctus,
whilst in 7’. conurus and ZY. muriei it is nearly always longer; that
in 7. tricinetus the head is more than half the length of the poste-
rior moiety of the carapace, whilst in the two other species it is not
so much as half that measurement.
1 Dr. Murie figures the tail on the left of the head (/, c.)—the photograph
from which the drawing was made not haying been reversed, most probably.
1878.] MR. A. H. GARROD ON TOLYPEUTES TRICINCTUS.
Length of cephalic
shield ...........+
Length of ante-
rior shield .....
Length of poste-
rior shield ......
Length of tail
(upper aspect).
T. tri-
cinctus,
in.
2-25
on
oO
BM.
. Ino. 1403.
=
T. conu-| T. conu-| T. conu-
rus. rus. rus.
in, in. in.
2-75 2°75 2°75
2°75 2°55 2°65
5D 5°75 58
2'2 2-2 2:2
B.M. | B.M. 52, B.M. 46,
. 140 2.| 8, 24,10. |9, 28, 35.
225
T.muriei,| T. muriei,
in. in
31 [;
2'8 2°65
63 6:3
? ?
B.M. | Dr. Mu-
no. 1404] rie’s
specimen.
It should also be mentioned that in T. fricinctus there is much
more hair on the carapace at the posterior margins of the scutes than
in the two other species, and that it has only eight teeth on each side
of each jaw, whilst both in 7. conurus and in T. muriei there are
nine.
All the specimens of the genus Tolypeutes agree, as far as my ex-
perience goes, in the manner in which their cervical vertebrze anky-
lose—the atlas,
free.
together with vertebrae 5, 6, and 7 alone being
Concerning the geographical distribution of the genus Tolypeutes,
the localities whence both Dr. Murie’s and my specimen were ob-
tained cannot be determined, they having been both purchased of
dealers. :
The specimen 140 a in the British Museum, of 7. muriei, was
presented by Burnett and Fitzroy, and therefore must have pro-
bably come from some part of the coast of La Plata or Patagonia.
A national specimen of 7. conurus is marked as having come from
Bolivia. Another of the same species, in the museum of the Royal
College of Surgeons, was presented by Mr. Darwin.
The type specimen of 7. conurus was obtained in the province of
Santa Cruz de la Sierra, Bolivia; and for a cast of its cephalic shield,
from which I have been able to determine the correctness of the
nomenclature above adopted, I have to express my very best thanks
to Professor Alphonse Milne-Edwards.
The species described by Burmeister’ from Buenos Ayres is 7’.
conurus.
Azara found a four-toed species in Paraguay ; it was therefore not
T.. tricinctus.
The brain of no species of Tolypeutes has been described, Dr.
Murie having met with an accident with his specimen. By Gervais a
cast is figured of the interior of the skull, which demonstrates the
Proc. Zoou. Soc.—1878, No. XV.
* Anales del Museo Publico de Buenos Ayres 1871 tom. ii. p. 117.
226 MR.A.H.GARROD ON TOLYPEUTES TRICINCTUS. [Feb. 19,
large size of the olfactory lobes’, together with its general propor-
tions, and but little more.
Our knowledge of the brain of the Dasypodide has been much
increased by a paper from the pen of Prof. Turner in the first
volume of the ‘Journal of Anatomy and Physiology’* on the brain
of Dasypus sexcinctus, in which the bibliography of the subject is
fully given.
The general appearance of the brain of Tolypeutes tricinctus differs
but little from that of Dasypus sexcinctus, whilst in surface-markings
it much more closely resembles that of Tatusia peba, as figured by
Rapp *, being comparatively simple in its gyration. There is not
that difference in the breadth of the anterior and posterior portions
of the cerebral hemispheres which is found in Dasypus sexcinctus.
The Sylvian fissure is only represented by an open and very shallow
angle, above which, on the surface of the hemisphere, is a slight,
partly encircling, shallow groove of some length, separated from it
by a very short distance (wide fig. 2, c).
Brain of Tolypeutes tricinctus.
The large olfactory lobes are are much broader than they are deep.
On the surface of each cerebral hemisphere there are two sulci.
One of these, the anterior, is horizontal, and divides the frontal lobe
into an upper and a lower part. Its outer extension is nearly to the
lateral margin, not going backwards more than one fifth the length
of the hemisphere. Internally it ceases on the flat interhemispheral
surface as far backward as its outer end (wide fig. 2, a).
The second sulcus is upon the superior cerebral surface, running
from behind forwards and inwards. Posteriorly it commences at the
outer posterior angle of the hemisphere a short distance from its ex-
1 «Ties formes cérébrales des Edentates,’ Nouv. Archiy. du Muséum, 1869,
tom. v. pl. ii. fig. 8.
2 186%, p: 313. 3 Hdentaten, tab. viii. fig. 3.
1878.] MR. A. H. GARROD ON TOLYPEUTES TRICINCTUS. 227
tremity. It does not, therefore, turn round the posterior end of the
hemisphere to become continuous with the fissure of the hippocam-
pus, as it does in Dasypus sexcinctus. Anteriorly it ceases in the
middle of the superior surface of the frontal lobe, a short distance
behind the convex margin of its upper moiety, and at about the
transverse level of the ends of the anterior sulcus. The general
direction of this sulcus is horizontal when viewed from the side, it
being slightly curved, with its convexity downwards.
The sulcus present on the superoparietal cerebral surface of Dasy-
pus sexcinctus is entirely absent in Tolypeutes tricinctus.
The internal cerebral surface very closely resembles that of Dasy-
pus sexcinctus as figured by Prof. Turner. The corpus callosum is
short, pointed in front, inclined obliquely upwards and backwards,
being rounded and thick posteriorly, where it becomes continuous
with the psalterium of the fornix. The anterior commissure is well
developed. The sides of the corpus callosum are very much
upturned. The calloso-marginal sulcus is feebly represented, being
of about the length of the corpus callosum, and situate at a level a
little anterior to it (vide fig. 2, d).
The corpora quadrigemina are large, and separated by a deep lon-
gitudinal fissure, the pineal gland being extremely small. In the
natural position of the spirit-preserved specimen it was quite impos-
sible to cover the testes by the posterior margins of the cerebral
hemispheres, which leads me to think that Prof. Turner is hardly
justified in his correction of Tiedemann, Cruveilhier, and Stannius,
who all say that the corpora quadrigemina are uncovered by the
cerebrum (in part at least) (wide fig. 2 a).
The median and lateral lobes of the cerebellum are nearly equal in
size. The medulla is very large, and the fifth nerve is enormous.
The optic nerves and commissure are insignificantly small. I could
not see any corpora albicantia.
With reference to the visceral anatomy of Tolypeutes tricinctus,
the tongue was 23 inches long in my specimen, and °4 inch broad at
the base. It is soft and elongate-triangular (lanceolate) in shape,
flattened above. No circumvallate papille are visible, the others
being inconspicuous and uniformly distributed. On the palate
there are nine larger and smaller transverse ridges on each side, the
interval between the last pair of molar teeth being smooth. The
number of teeth is eight above and seven below, on each side.
The epiglottis is slightly indented by a notch in the middle line ;
and the soft palate embraces it, so that the rima glottidis opens into
the posterior nares, as in so many mammals, during ordinary respi-
ration. A tonsilitic pit exists on each side of the fauces.
The Jungs are divided into four lobes on the right side, and two
on the left. In most Dasypodide there are three lobes on the left
side; but in the animal under consideration the two upper were
blended. Three is said by most authors to be the number of lobes
of the right lung; but I always find a fourth azygos lobe as well,
hidden behind the heart, in the genera Dasypus and Xenurus, absent
in Tatusia. Dr. Murie’s figure of the lungs ee eS ex-
15*
228 MR. A.H.GARROD ON TOLYPEUTEs TRICINCTUS. [Feb. 19,
cept in the absence of division of the left upper lobe, from my speci-
men.
The stomach is of the ordinary shape, with but a short lesser cur-
vature. It is muscular, but not powerfully so, at its pyloric end.
Along its greater curvature, when laid out flat, it measures 63 inches.
The liver has the lateral fissures deep, whilst the umbilical fissure
is comparatively insignificant, The gall-bladder is partly, but not
deeply, embedded in the abdominal surface of the right central lobe.
The Spigelian lobe forms a short rounded cone, very broad at its
base. According to the method elsewhere suggested by me’, the
formula of the bulk of the lobes is thus expressed :—
L.L. 13>R.C.>R.L. 2>L.C. 2>C.
The small intestines measure 75 inches, the large intestine 6°5
inches. There are no ceca; but there is an abrupt change in the
diameter of the alimentary canal at the ileo-ceecal valve, as in T’.
conurus and in the genus Tatusia.
The uterus is triangular, the fundus being perfectly straight when
viewed from in front, and the Fallopian tubes joining it at the ex-
treme upper and outer angles. The conical clitoris is an inch long ;
and the genito-urinary orifice is a longitudinal slit 0°3 inch from its
apex.
vom the various papers on the visceral anatomy of the Dasy-
podidee I may refer to Hunter’s description of Tatusia peba*, Pro-
fessor Owen’s account of the same species, and of Dasypus sexeinc-
tus*, and Hyrtl’s monograph of Chlamydophorus truncatus*. To
these I may add my own notes on Xenurus unicinctus, together with
those upon the other species which have passed through my hands,
as an assistance towards the determination of the affinities of Z'oly-
peutes.
In Xenurus unicinctus the gall-bladder is so deeply embedded in
the tissue of the right hepatic lobe that it nearly penetrates to its
diaphragmatic surface. This I find to be the case in Dasypus vil-
losus, D. sexcinctus, and D. vellerosus; whilst in Tatusia hybrida
and Tolypeutes tricinctus it is much less sunk. The cystic duct is
very much twisted in a corkscrew manner. The proportional bulk
of the hepatic lobes is almost exactly the same as in Tolypeutes tri-
cinctus. In Tatusia peba and T. hybrida the right central lobe is
the largest, not the left lateral. In Tatusta the umbilical fissure is
less significant than in Dasypus, Xenurus, or Tolypeutes.
The junction of the large and small intestines in Xenurus is as in
Tatusia peba and T. hybrida, there being no cecal dilatations, as
there are in Dasypus villosus, D. sexcinctus, D. minutus, and D.
vellerosus.
The following measurements of the intestines demonstrate their
relative lengths :—
1 P. 7.8. 1875, p. 57.
? «Essays and Observations on Natural History,’ 1861, vol. ii. p. 182.
° P.Z. 8. 1831, pp. 141 and 154, and P. Z. 8. 1832, p. 180.
* Denkschr. der kais. Akad. Wien, ix. 1855.
1878.] | MR. A. H. GARROD ON TOLYPEUTES TRICINCTUS. 229
SY PS] Ces iene are
Sto | Sor] 8. | Seo} So] § .
sxi|sx |] 8s/8s/e¢)] 98
BeF/ Ss] SS] SS] ss] SX
Ee ee ieee rae ay “hs
ft ft ft tt: ft. ft.
Small intestine...... 16:3 | 146 |} 185] 115] 5-1 43
|
Large intestine...... 15 15 16 1°25) 7-0 0:66
|
In Xenurus unicinctus the vesiculz seminales are not present, the
distal ends of the ureters being enlarged. Cowper’s glands and a
prostate are present; and the apex of the glans penis is not trilobed
as in Tatusia, but simple, as in Dasypus and Tolypeutes. The
penis itself is huge. In Tolypeutes, according to Dr. Murie, the
vesiculee seminales are also absent.
In Tatusia peba and T. hybrida the vesiculze seminales are large,
and there is a pair of lateral lobes to the apex of the glans penis.
The whole penis is proportionally smaller in Twtusia than it is in
Dasypus, as is the clitoris. In these respects Tolypeutes and Xenu-
rus agree with Dasypus, my experience being, as far as the male
organs of Dasypus are concerned, confined to D. sexeinctus.
In Dasypus, Tolypeutes, and Xenurus there are only a pair of
nipples, in J'atusia there are two pairs. In NXenurus unicinctus
the uterus is triangular, as in Dasypus and Tolypeutes.
Fig. 3.
Brain of Xenurus unicinctus.
The brain of Xenurus is intermediate in its form and surface-
markings between Dasypus and Tolypeutes. The hemispheres, when
230 MR. J.H. GURNEY ON A SPECIMEN OF POLYBORUS. [Feb. 19,
viewed from above, are considerably broader posteriorly than an-
teriorly, as in Dasypus. The olfactory lobes are large; and the
corpora quadrigemina are completely covered. The oblique sulcus
on the back part of the superior surface of each hemisphere, which
runs forwards and outwards, is present, as in Dasypus, but it is very
short. The two other sulci are well developed, that in the frontal
portion being longer than in either Dasypus or Tolypeutes. The
upper or second sulcus runs from in front—near the anterior internal
angle of the superior surface of the hemisphere—backwards and
outwards for some distance, but not nearly as far as the posterior
margin of the cerebrum, it ceasing a little behind the level of the
outer extremity of the posterior oblique sulcus. In this respect it
resembles Tolypeutes and differs from Dasypus (vide fig. 3).
It is to Prof. Owen that we are indebted for our knowledge of
the great differences between the colic ceca and female generative
organs of the genera Tatusia and Dasypus. These facts, when
taken in association with other characters above brought forward and
referred to, may be thus tabulated :—
Dasypus.
Ears apart.
Nipples 2.
Uterus triangular.
External genitalia
enormous.
|Glans penis simple.
Vesiculz seminales
absent.
Lateral colic cxeea
present.
Azygos lobe of lung
present.
Tolypeutes.
Ears apart.
Nipples 2.
Uterus triangular.
External genitalia
enormous.
Glans penis simple.
/Vesicule seminales
absent.
No colic exea.
|Azygos lobe of lung
present.
Aenurus.
Ears apart.
Nipples 2.
Uterus triangular.
‘External genitalia
enormous.
Glans penis simple.
Vesiculz seminales
| absent.
‘No colic ezeca.
present.
Chlamydophorus.
Ears apart.
|Nipples 2.
Uterus pyriform.
External genitalia
small in female.
?
?
Lateral colie cca
present.
Azygos lobe of lung) Azygos lobe of lung
present (Hyrtl).
Tatusia.
Ears approximate.
Nipples 4.
Uterus pyriferm.
External genitalia
not excessive.
Glans penis tri-
lobed.
Vesiculz seminales
present.
No colie cxea,
Azygos lobe of ling
absent.
Thus we may infer that the genera Dasypus, Tolypeutes, and
Xenurus resemble one another more than any of them do Tatusia.
Chlamydophorus, in other respects so different from them all, is
like Dasypus alone in its colic ceea, like Dasypus, Tolypeutes, and
Xenurus in most respects, and like T'atusia only in the shape of its
uterus.
2. Notes on a Specimen of Polyborus lately living in the
Society’s Gardens. By J. H. Gurney, F.Z.S.
[Received Jan. 17, 1878.]
The Society’s ‘ Progeedings’ for 1876 contain at p. 333 a plate
and a concise notice of two remarkable Polyéori which were said to
have been obtained in Patagonia, and which were placed in the
Gardens on the 2nd of March of that year.
1878.] | MR. J. H. GURNEY ON A SPECIMEN OF POLYBORUS. 231
On the 27th of May, 1877, one of these birds unfortunately died ;
and on examining the skin of this specimen, which proved on dis-
section to be a male, I have noticed the following points.
The wing measures 15°6 inches; but the ends of the primaries are
worn considerably, and except for this the measurement would, I
think, have exceeded 16 inches ; the length of the tarsus is 3°7, and
that of the middle toe s. u. 2°3 inches.
I have had the opportunity of comparing these measurements with
those of four specimens of Polyborus tharus which have been kindly
lent to me by Messrs. Salvin & Godman, and with those of two
specimens in the Norwich Museum; also with the measurements
given by Mr. Sharpe, at p. 32 of the first volume of his Catalogue,
of two others in the British Museum.
Unfortunately in none of these eight examples is the sex recorded ;
but the result appears to prove that Polyborus tharus is subject to
considerable variation of size, which further investigation may show
to be incident, in part, to the different localities from which the
specimens have been obtained.
The following are the measurements alluded to :—
Tn the collection of
Messrs. Salvin & Godman. Wing. Tarsus. Middle toe 8. «.
Bahia 14°8 3°0 1-9
of 15:1 35 2°2
156 3°5 21
33
Supposed to be from
_ Bolivia 16:0 3°5 2°2
In the Norwich Museum.
Locality unknown
17°2 3°7 2°3
» 17-4 3°9 PN
In the British Museum,
as given by Mr. Sharpe.
Locality not quoted 15°5 3°8
» 17°9 3°8
It will be seen from the above measurements that there is nothing
in the dimensions of the specimen which died at the Gardens to show
that it is not an abnormal example of P. ¢harus ; and although the
extraordinary paleness of the ground-colour of its plumage presents
a curious difference from the coloration of the same portions of
the plumage in P. tharus, the form and disposition of the markings
in great measure agree with those of that species in its immature
stage.
The following notes refer to the details in which the pale specimen
agrees with the ordinary P. ¢harus in its immature stage :—The upper
part of the head, including the occipital crest, is decidedly darker
than the adjacent plumage ; the ear-coverts are whole-coloured and
of a castaneous tint; the feathers of the throat and upper breast
have strongly marked longitudinal shaft-marks ; and similar shaft-
marks, but narrower and fainter, are perceptible on the lower breast
232 MR. J. H. GURNEY ON A SPECIMEN OF POLYBORUS. [Feb. 19,
and on the interscapular feathers. All these shaft-marks resemble
greatly in shape those on the corresponding portions of the plumage
in the normal immature P. ¢harus; but in the latter the shaft-marks
are paler than the sides of the feathers, while in the pale Patagonian
specimen they are darker and more rufous. In the last-named bird
faint transverse marks of a similar rufescent tint, but very pale, are
perceptible on the abdomen and under tail-coverts ; similar transverse
marks, but darker, less rufous and more numerous, are apparent in
a specimen of P. tharus in Messrs. Salvin & Godman’s collection,
marked “ Bolivia ?,’? which seems to me to be passing from the im-
mature to the adult stage. Transverse bars alternately dark and pale
and of a similar character are found on the lower part. of the back in
Polyborus tharus in all its stages; and these markings, though
modified in their coloration, especially by the substitution of dull
fulvous bars for those of blackish brown in ordinary specimens, are
unmistakably present in the pale specimen under consideration.
The latter bird was moulting its primaries when it died; and I
observe that the transverse bars on the new quill-feathers are broader
than on the old, the latter agreeing in general character with the
corresponding bars in immature specimens of the normal P. tharus,
and the former with the same bars in adult birds. ‘These bars are
much more rufous, especially on the newly acquired feathers, than
is the case in normal specimens; in the latter they are of various
shades of brown, but not tinged with rufous.
The narrow transverse bars on the upper surface of the tail are
less distinct in the Patagonian pale bird than in ordinary specimens
of P. tharus ; but they are similar in character and of about the same
number (from 15 to 16). The broad terminal blackish-brown band at
the end of the tail in the ordinary bird is represented in the pale
one by a dull fawn-coloured band of similar dimensions tinged with
rufous on its upper edge and on the external webs of the outer pair
of rectrices ',
During a recent visit to the Zoological Society’s Gardens, I observed
that the bare skin on the face of the survivor of the two pale birds,
changed rapidly from carmine-red to pale pink ; but Mr. Bartlett
informed me that in neither of the pale specimens did these occasional
and transient changes assume a wider range, and that in them the
bare skin never became yellow, as may be sometimes seen to be the
case (even to the extent of a decided lemon-colour) in the normal
specimens of Polyborus tharus inhabiting the adjacent cages.
On the whole, I incline to agree with the opinion expressed by
Mr. Sclater in his original notice of this curious pair of birds, that
they are “ young individuals of P. ¢harus in an abnormal phase of
plumage,” or, as I would rather say, of colouring generally.
* The tail in the pale birds was imperfect when they were drawn for the
plate in the Proce. Zool. Soe, ; but the specimen under consideration obtained its
full tail subsequently,
1878.] MR. D. G. ELLIOT ON THE PTEROCLIDE. 233
3. A Study of the Pteroclide or Family of the Sand-Grouse.
By D. G. Exxon, F.R.S.E. &e.
[Received January 18, 1878. ]
The materials for the present paper were supplied by the
splendid collection in the Paris Museum, kindly placed at m
disposal by Prof. A. Milne- Edwards, who afforded me every facility
for investigating this interesting group of birds, and to whom, as
well as to Dr. E. Oustalet, Aide-Naturaliste, I desire to express my
thanks. With the exception of P. decoratus, the type of which is,
I believe, unique, every known species of the Pteroclidz is repre-
sented in that Museum, many of them by series of both sexes in
different plumages ; and certain Specimens are the types of the older
writers. In the treatment of my subject I have followed the same
method as that adopted for my “ Review of the Thidinee”’+, and give
first the Literature of the Family, then the Classification, followed
by some remarks upon the different genera proposed for the group ;
LITERATURE OF THE Famity.
Linneus is the first author to claim our attention.
1766. Linnavs, « Systema Nature,’
In the list of Species comprised in the genus Tetrao, this author
Places among those characterized by « pedibus nudis” a single
species belonging to the Pteroclide, called by him Teérao alchata,
Species 1,
1771. Linnzus, ‘ Mantissa.’
In this work the P, senegalus is described as Tetrao senegalus.
Species 2.
1773. Pauxas, ‘Reise durch Verschied. Provinz. Russisch. Reichs.’
Syrrhaptes paradoxus is described as Tetrao paradoxus. Species 3.
1774. Pauas, ‘Nov. Comm. Petropol.’
Pierocles arenarius first made known as Tetrao arenarius.
Species 4,
1786. Scoroxt, ‘ Deliciz Faun et Flore Insubrice.’
Pterocles fasciatus first described as Tringa Jasciatus, Species 5,
1788. Gme tin, ‘ Systema Nature,’
Five species of this family are given by this author, all placed in
innzus’s genus Tetrao, viz. T. alchata, Linn., 7. namaqua, de-
scribed for the first time, 7. indicus (=T7. fasciata, Scop.), 7.
arenarius, Pall., and T. paradoxus, Pall. Species 6.
* P. Z. 8. 1877, p. 477.
234 MR. D. G. ELLIOT ON THE PTEROCLIDE. [Feb. 19,
1815. Temminck, ‘Histoire Naturelle Générale des Pigeons et
Gallinacés,’
A general review of the Pteroclide is here given, as known to the
author, with some species renamed that were previously described.
They are as follows :—P. arenarius, Pall., P. bicinctus, P. quadri-
cinctus, both described for the first time, P. setarius (=T. alchata,
Linn.), P. tachypetes (=T. namaqua, Gmel.), and Syrrhaptes
pallasii (=T. paradowa, Pallas). Species 8.
1817. Virrixot, ‘ Nouveau Dictionnaire d’ Histoire Naturelle.’
In his genus Gnas are here included such species of this family
as were known to the author, viz. @. cata (=T. alchata, Linn.),
. bicineta, Gi. indica (=T. fasciata, Scop.), Gi. namaqua, and
CE. arenaria. No new species are described.
1822. Burcue tt, ‘Travels in South Africa.’
Pterocles variegatus first described. Species 9.
1823. LicuTEeNstTrErIn, ‘ Verzeichniss der Doubletten des zool.
Museum zu Berlin.’
Pterocles coronatus first described. Species 10.
1825. Temminck, ‘Planches Colorices.’
Pterocles exustus and P. lichtensteini first described and figured.
In an earlier portion of the work, in the 9th livraison (1823), P.
arenarius is figured ¢ and 9; and in the 57th livraison (1825) a
representation of both sexes of P. coronatus is given. Species 12.
1837. Swainson, ‘ Birds of West Africa.’
Pterocles quadricinctus, Temm., renamed P. tricinctus.
1843. Gou.p, in the ‘ Voyage of the Sulphur, Birds.’
Pterocles personatus described. Species 13.
1849. Smiru, ‘Illustrations of South-African Zoology.’
Pterocles yutturalis described. Species 14.
1850. Gouxp, in ‘ Proceedings of the Zoological Society of
London.’
Syrrhaptes thibetanus described. Species 15.
1856. Bonaparte, in ‘Comptes Rendus.’
In his Table des Galinacés, Bonaparte proposes the genus Péero-
clurus for certain species of this family, of which the 7’. alchata,
Linn., may be considered as the type.
1870. Casanis, in ‘Von der Decken’s Reise.’
Pterocles decoratus described. Species 16.
CLASSIFICATION.
The natural position of the Pteroclide (called by Prof. Huxley
Pteroclomorphe), in the Class Aves, is between the Columbide
(Peristeromorphz) on the one hand, and the Tetraonine series of the
1878. | MR. D. G. ELLIOT ON THE PTEROCLIDE. 235
Gallinaceous birds (Alectoromorphz) on the other, supplying the link
which connects these two great divisions, and with them constitutes
the Gallo-Columbine series. The Pteroclide resemble the Alecto-
romorphe in their skull, palatines, maxillo-palatines, and bill; but
the pterygoid and basipterygoid processes, sternum, furcula, coracoid,
and fore limb are like those of the Pigeon. The foot, with its
almost absent hallux (entirely wanting in Syrrhaptes), and tbe short
tarso-metatarsus are, however, very unlike a Pigeon’s. The vocal
organs, as shown by Mr. Parker’, are the same as a Pigeon’s; the
trachea is cartilaginous, and possesses a pair of laryngeal muscles
at its bifurcation; but the crop, gizzard, gall-bladder, and small
intestines are like those of Gallinaceous birds. The czeca coli are
very voluminous, but have twelve continuous longitudinal folds in
their mucous membrane, instead of seven as in Lagopus. The
proventricular glands, however, are Pigeon-like. The pterylosis of
the Saud-Grouse, although very similar to that of a Pigeon, differs
somewhat on its dorsal surface. The lateral neck-spaces are shorter,
only reaching to the beginning of the neck; the superior wing-space
is absent ; the lumbar tracts coalesce with the posterior part of the
dorsal tract, which latter also joins the plumage of the tibia. In
some species the narrow spinal tract is wanting. The Pteroclide
also differ from the Pigeons in possessing an after-shaft on the
contour feathers, and from the Gallinaceous birds in having a
naked oil-gland.
From the above it will be clearly seen how intermediate these
singular birds are of the two great groups between which they are
placed. In some of their characters they are also pluvialine, and
their flight is especially Plover-like; but in their method of drinking
they resemble the Pigeons, as the bill is thrust up to the nostrils
into the water, and retained there until the bird’s thirst is allayed.
The members of the two genera adopted for this family closely
resemble each other in their outward shape, being generally rather
heavy of body, with long pointed wings and extremely short legs
and toes, apparently quite disproportionate in size to the rest of
the bird. The species, therefore, are awkward when on the ground,
run clumsily, and appear to great disadvantage; but on the wing
they move rapidly and gracefully. The bill, though small, is strong
and Grouse-shaped ; in Pterocles the tarsi are feathered in front ; in
Syrrhaptes both tarsi and toes are com pletely covered with feathers,
as in the winter coat of the Ptarmigan. I have arranged the
members of Pterocles in two divisions, distinguished by the pre-
sence or absence of a conspicuous band across the breast. ‘The first
of these, without a pectoral band, contains five species, all but one
having the breast of a uniform coloration.
The second division, with a pectoral band, possesses nine species,
having, as a rule, a very different style of coloration from those in
the other section of the genus. In the remaining genus Syrrhaptes,
but two species are as yet known, separated in their outward appear-
ance from their relations in Pterocles by the feathered leg and foot
1 Trans. Zool. Soe. vol. y. p. 204,
236 MR. D. G. ELLIOT ON THE PTEROCLID. [ Feb. 19,
and the entire absence of a hind toe. One character by which
Syrrhaptes was formerly distinguished was the extremely lengthened
filamentous first primary observed in S. paradoxus; but since the
discovery of the second species, which does not possess a similar
formation, this peculiarity will not answer as a means of generic
distinction. Both species of Syrrhaptes have the median rectrices
much lengthened beyond the rest; but as this also occurs with
several species of Péerocles, it cannot be regarded as of generic
importance.
GENERA.
But few generic terms fortunately have been proposed for these
birds. Usually placed by the older authors, who were misled by
their feathered tarsi, in Yetrao, they were first separated from
the Grouse by Temminck, who, in 1815’, created for them the term
Pterocles. In 1811 Illiger proposed Syrrhaptes for the Tetrao
paradora of Pallas, separating it from the other species of the
family. In 1812 Fischer instituted the term Mematura for the
same bird, followed in 1816 by Vieillot with Heteraclitus, and also
with Gnas for the other members of the family. Lastly Bona-
parte, in 1856, created the term Pteroclurus. Two generic forms
are evidently represented among the Pteroclide, for which the terms
Syrrhaptes, of Iliger, to contain the two species S. paradoxus and
S. thibetanus, and Pterocles of Temminck to include all the rest of
the family, have the claim of priority, and are amply sufficient for
the recognition of the generic divisions now existing. In the
osteology of the two genera there are considerable differences obser-
vable, as has been well shown by Mr. Parker, some of which are the
following :—The skull of Syrrhapées is weaker, less gallinaceous,
and more pigeon-like than that of Pterocles; the upper frontal
region is narrower between the eyes, and the alze of the ethmoid are
less swollen between the crura of the nasal. In Pterocles the bones
of the face are as strong as is observed in a Pigeon; the lower
jaw is stronger than in Syrrhaptes, and bends further back; the
postorbital and squamosal processes are stronger, as is also the
malar arch; in some characters of the skull and skeleton they are
neatly alike. Péerocles has the scapula more like that of a Grouse,
and also possesses one more caudal vertebra than Syrrhaptes, and the
styliform and sacral ribs have no appendage; but both genera have
a rudiment attached to the last hemapophysis. The sternum is
stronger, with the episternum and hyosternal processes as in Syr-
rhaptes ; but the external hyposternal processes are shorter.
Of fossil remains of Pteroclidee but two tarso-metatarsal bones have
been found, which have been described by Prof. A. Milne-Edwards, in
his great work ‘Les Oiseaux Fossiles,’ as Pterocles sepultus. By the
kindness of their possessor I have had an opportunity of examining
1 In his ‘Manuel d’Ornithologie’ (1820), 2nd part, p. 474, in a footnote,
Temminek states that he published an account of these birds under the generic
name of Pterocles in 1809. I have not been able to find this publication, nor
any record of it, and 1815 is the earliest date I can assign to the genus.
1878.] MR. D. G. ELLIOT ON THE PTEROCLID&. 237
these interesting specimens. ‘There is a slight difference observable
in their size, one being rather stouter than the other; and they evidently
belong to two individuals, perhaps different sexes of the same species ;
and, as stated by Monsieur A. Milne-Edwards’, represent a species
between the P. alehata, Linn., and the P. gutturalis, Smith.
The following are the genera proposed for these birds, with the
type of each added :—
Type.
(1811) Syrrhaptes, Illiger, Prodr. p. 243 ......... Tetrao paradoxus, Pall.
(1812) Nematura, Fisch. Mém. Soc. Natur. Moscou Tetrao paradoxus, Pall.
(1815) Pterocles, Temm. Man. Ornith, p. 299...... Tetrao alchata, Linn.
(1816) Gnas, Vieillot, Analyse, p. 52 ............08 Tetrao alchata, Linn.
(1816) Heteraclitus, Vieillot, Analyse, p. 53 ...... Tetrao paradoxus, Pall.
(1856) Péeroclurus, Bon. Compt. Rend. slii. p. 880 Tetrao alchata, Linn.
GEOGRAPHICAL DISTRIBUTION.
The Pteroclide have been found in only three of the zoogeogra-
phical divisions of the earth’s surface, viz. the Palearctic, Oriental,
and Ethiopian regions; and in these they are chiefly restricted to
tracts devoid of cultivation, or actual deserts. From the fact that
of the sixteen known species twelve are met with in the Ethiopian
region, we may safely regard the Pteroclidz as having originated in
Africa, from which they spread into the other two regions named
above. Of the twelve Ethiopian species, seven (or a little over half)
are not found in the other zoogeographical divisions. In the
Oriental region, which is the next richest in the birds of this family,
seven species are found, one of which is peculiar to it ; and, lastly,
the Palearctic region is tenanted by five species, two of which,
constituting the genus Syrrhaptes, are not found elsewhere. Com-
mencing with the Ethiopian region, where the species appear confined
mainly to the eastern side of the continent, we find the Pé. gut-
turalis extending its range from Abyssinia to Kurrichane, just north
of the Cape Colony, where it was procured by Sir A. Smith. In
Egypt and Nubia P¢. coronatus is found, not going south of the
last-named country on this coast. It is also met with in the
southern part of the desert of Sahara, and again on the plains of the
Sinaitic peninsula, and probably occurs throughout Arabia, as, together
with the three species next mentioned, it is found in Sindh of the
Oriental region. In the region extending from and including
Egypt to the Somali country Pt. evustus, Pt. senegalus, and Pt.
lichtensteint are found ; while Pt. quadricinctus appears to be re-
stricted to Abyssinia and Sennaar, although given by Swainson as
a bird of West Africa under the name of Pt. tricinctus. Passing
southwards along the same coast, we find Pt. variegatus, Pt.
namaqua, and Pt. bicinctus inhabiting the Trans-Vaal, whence
they range across this part of the continent into Damaraland.
Somewhere in the interior of Eastern Africa Pé. decoratus dwells,
according to Cabanis; but its precise distribution does not appear
1 Oiseaux Fossiles, tom. ii. p. 295, pl. 141. figs. 1-9.
238 MR. D. G. ELLIOT ON THE PTEROCLIDA. [ Feb. 19,
to be exactly known. The type and only known specimen was
obtained near Lake Jipe. It is most probable that the range of
some of the species mentioned above may be much greater; but
before this can be definitely determined our knowledge of the
interior of the continent and its avi-fauna must be greatly increased.
Pt. exustus is given by Hartlaub, in his ‘System der Ornith. West-
Afrika’s,’ upon the authority of a specimen in the Berlin Museum,
as an inhabitant of Senegal. I have not seen this corroborated by
any other writer; and the species seems to be more particularly a
bird of East Africa, although it is not unlikely that it may skirt the
southern border of the Sahara quite across the continent. Finally,
according to Tristram, P¢. arenarius is found everywhere in the
Sahara, except in the extreme south. In the Oriental region,
the next in order, we find that in Sindh, with the exception of
Pt. decoratus and Pt. quadricinctus, all of those species already
mentioned as dwelling in the Ethiopian region are met with, two
of which (Pé. coronatus and Pt. lichtensteini) do not seem to be
found in any other part of India. P¢. senega/us is met with along
the shores of the Runn of Cutch and in Northern Guzerat ; while
Pt. arenarius goes north into the Punjaub and thence through the
North-west Provinces; and P¢. alehata occurs in the Peshawur
valley. Pt. exustus and Pt. fasciatus are distributed generally
throughout India, the last-named species not being found in Sindh.
The last zoogeographical region in which these birds occur is the
Paleearctic ; and commencing in the far east we have the members
of the genus Syrrhaptes; of which S. paradoxus ranges from the
plains of Pekin and Tientsin, through Mongolia and the Great
Gobi desert into the Kirghis steppes, occasionally wandering into
parts of Western Europe in more or less extensive numbers. The
year 1863 was a uotable one for a great migration of this species,
flocks of considerable size having been observed as far west as
Ireland. The only other known species, S. tibetanus, is dis-
tributed from the Kokonor basin on the east, through Northern
Thibet to Ladakh. In this region also, Pé. senegalus is a native
of Palestine, where, according to Tristram, it is numerous. P2.
arenarius is common throughout Turkestan, and is not rare in the
eastern steppes of New Russia and the Caucasus. It is also met
with in Persia and about Erzeroom, not uncommon in Palestine,
and is distributed generally throughout south-western Europe, being
common in Spain and Portugal. It is also a dweller of the Algerian
deserts, whence it extends into the Ethiopian region as already
enumerated, and has also been obtained in Greece. Its most
western locality that I have seen recorded is the Canary Islands,
where it has been observed in the Gran Canaria. The only re-
maining species to be noticed in the Palearctic region is Pé. alchata,
which ranges from the Kirghis steppes into southern Russia ;
it is common in Asia Minor and Palestine, and occurring through-
out Southern Europe, is found in France where Pt. arenarius is
not known. It is also common in Spain, and frequently met with
in Portugal, and is abundant in the Algerian deserts. It has been
1878. | MR. D. G. ELLIOT ON THE PTEROCLID. 239
obtained at Malta, and also in Greece, where, according to Von der
Miihle, a specimen has been procured in Eubecea.
Genus PTEROCLEs.
Body rather long and heavy; bill rather stout; culmen curved ;
groups ascending; sides of mandibles compressed. Nostrils basal,
open, oblong, partly covered by a membrane; frontal feathers pro-
jecting over, but not intruding upon, the nasal aperture. Wings long,
pointed ; first primary longest. Tail cuneate; median rectrices
sometimes lengthened heyond the rest. Tarsi short, covered an-
teriorly with downy feathers; feet small, bare, the hallux extremely
short, almost obsolete, not reaching the ground.
Key to the Species.
A. Without pectoral band. .
@, Bront and: chin) black y «ac .4.% es -ccasesvcss oe cveviot 1. Pt. personatus.
6. Loral space black; throat yellowish, bordered
on its lower edge with a band of black ; breast
greenish-grey ; abdomen chestnut ............... 2. Pt. gutturalis.
c. Stripe on each side of forehead, from nostril to
above the eye, chin, and centre of throat black 3. P¢. coronatus.
d. Lores and band encircling back of head pearly-
grey; cheeks, ear-coverts, and throat orange
yellow; centre of abdomen black ............... 4, Pt. senegalus.
e. Top of head yellowish-brown, streaked with
black; throat pearly-grey; underparts rufous,
spotted with white........... siccSine yoberibackcscogebcs 5. Pt. varicgatus.
B, With pectoral band.
a. Without black bar on forehead.
a’, Median rectrices not lengthened much be-
yond the rest.
Upper part of throat and sides of neck
rufous; lower portion of throat black ;
band on lower part of breast; abdomen
and flank black ............ BPR told. RE 6. Pt. arenarius.
b'. Median rectrices greatly lengthened be-
yond the rest.
a’, Throat yellow; black band across
breast ; abdomen and flanks chest-
Wulf dacs tasesasbederrededaneemenere nr siaaee 7. Pt. exustus.
b". Throat and stripe behind the eye
black; sides of throat rufous; centre
of breast chestnut, bordered above
and below with black ; rest of under-
DANUSLWOMLON Mesitssec rade: ssectk aes tee 8. Pt. alchata.
oe”. Throat orange-yellow; breast crossed
by a white and a chestnut band;
abdomen rufous brown............ vee «9, Pl. namaqua,
6. With black bar across forehead.
wd. Breast uniform greenish buff.
a”, Lower part of breast bordered with a
chestnut band, succeeded by a white
one; rest of underparts yellowish-
white, barred narrowly with black ;
wing-coverts with two black bands,
margined on the upper side only
with white .......... Belk cat Thome « cpeeee 10. Pt. fasciatus,
240 MR. D. G. ELLIOT ON THE PTEROCLID. [Feb. 19,
6”. Lower part of breast bordered with a
rather broad white band, succeeded
by a broad black one; rest of under-
parts yellowish-white, irregularly
barred with black .............c.seeee- 11. Pt. bicinetus.
ec”. Lower part of breast crossed by four
bands, first a maroon one, succeeded
by a white one, then a black, and
lastly a white one, alternating ;
wing-coyerts with only one black
band, margined on both sides with
WILE Aisne rte tian: Ba ctkeieacteeea dase 12. Pt. quadricinctus.
4’, Throat pale buff; upper part of breast buff,
crossed with numerous narrow black
bars; middle of breast uniform buff,
crossed in centre by a narrow black bar,
and another of the same hue on its
lower edge; rest of underparts yellow-
ish-white, barred narrowly with black... 18, Pé. lichtensteini.
e. Black bar on forehead, divided in the centre by
a white spot; throat and abdomen black ; black
bar crosses the breast, between which and the
abdomen is a broad white band .............0-00s 14, Pt. decoratus.
1. PreROCLES PERSONATUS.
Pterocles personatus, Gould, Voy. ‘Sulphur,’ Zool. pl. 30 (1843) ;
id. P.Z.S. (1843), p. 15; Gray, Gen. B. vol. iii. p. 519 (1845) ;
Reich. Syn. Av. Gallinze (1851), pl. ceviii. figs. 1821, 1822.
Hab. Madagascar.
This is one of the largest species of the genus, readily recognized
from its relatives by the entirely black forehead and chin. M.
Grandidier informs me that it is very abundant in Madagascar,
where he has shot many, and that its flesh is palatable. In its
economy and habits it does not differ from the other members of the
genus.
Male. Forehead and chin jet black. Lower part of throat and
sides of face buffy-white. Top of head, occiput, upper part of back
and breast dark isabelline or vinous. Middle of back dark brown,
greyish tinge on some feathers; rest of back, rump, and upper tail-
coverts black, barred narrowly with light buff. Wings uniform,
like the breast ; a few of the greater coverts tipped with brownish
black, forming three irregular bars upon the wing. Inner secon-
daries dark grey, tipped with vinous. Primaries dark chocolate-
brown, with light edges on the inner webs. Abdomen, flanks, and
vent barred narrowly with black and rufous; the feathers nearest
the breast barred with black and buffy white. Under tail-coverts
light chestnut. Tail brownish black, barred with white (these
broadest and most numerous on the outer webs), the feathers largely
sped with white. Bill apparently bluish, with dark tips. Feet
ack.
Total length 133 inches; wing 93, tail 5, bill along gape 3.
Female. Throat and sides of face buffy white; breast vinous.
Entire upper parts and wings pale buff, barred irregularly with black.
Primaries dark brown, with light edges and tips. Flanks and
1878. ] MR. D. G. ELLIOT ON THE PTEROCLIDA. 241
chest barred with white and black; abdomen and vent barred with
rufous and black. Under tail-coverts ochraceous. ‘Tail blackish
brown, barred and largely tipped with yellowish white. Bill and feet
blue-black.
Total length 113 inches; wing 83, tail 47, bill along gape 3
A young eed Hite frome the female in having the breast vinous
in the centre, pale buff on the sides, each feather with two narrow
brown bars. Abdomen, flanks, and vent chestnut, barred sparingly
with black. Wings very pale buff, barred with black. Rest of
plumage as in the female.
2. PrEROCLES GUTTURALIS.
Pterocles gutturalis, Smith, Ill. 8.-Afr. Zool. (Birds), pl. 3 3,
pl. 31 2 (1849); Gray, Gen. B. vol. iii. p. 519 (1849); Reich.
Syn. Av. Galline (1851) pl. cex. figs. 1829, 1830; von Heugl.
Syst. Uber. Vig. N.O.-Afr. (1855) p. 304; Parker, Proc. Zool.
Soc. (1862) p. 258; id. Journ. fiir Ornith. (1862) p. 286; Layard,
Birds S. Afr. (1867) p- 278; id. Ibis (1868) p. 269; Ayres, Ibis
(1869) p. 297; Blanf. Geol. & Zool. Abyss. (1870) p. 421; Von
Heugl. Ornith. Nordost-Afr. Band ii. (1873) p. 862; Buckle, Ibis
(1874) p. 385.
Hab. Kurrichaine, South Africa (Smith); Trans-Vaal (Ayres,
Buckley) ; Adoa and Axum to Shoa, Abyssinia (Heuglin); Agula,
Antalo, Abyssinia (Blanford).
This fine species of Sand-Grouse was first described by Sir A.
Smith (/. c.), who obtained it about eighty miles to the eastward of
Latakoo, his attention having been drawn to the bird by its cry,
which was very different from that of P. namaqua, and resembled
the syllables tweet-tweet. Like all the other species they resorted
to the water in large flocks, but when feeding were dispersed and
observed only singly or in pairs. Grass-seeds and abundance of
gravel were found in the stomachs of the individuals that were killed.
The eggs, which are two or three in number, are laid upon the bare
ground; and the young take to a wandering life almost immediately
after they are hatched.
In Abyssinia Mr. Blanford considers this species to be local, as
he only saw it in two or three places in the highlands, at Agula
and Antalo; and Riippeil mentions it only at Shoa, though Von
Heuglin procured it near Adowa.
Mr. Ayres states that in the neighbourhood of Potchefstrom, in
the territory of the Transvaal, this species is abundant towards the
end of winter and beginning of spring. Their flight is strong, re-
sembling that of Pigeons ; and when alarmed, they do not run, but
lie close to the ground, from which they take wing at once. Their
notes, uttered only on the wing, are harsh, and may be heard at a
considerable distance. They feed upon a species of tare or pea, and
upon bulbous roots of a species of grass; from the aromatic pro-
perties of these last the birds, after feeding freely upon them, are
thoroughly scented. Iris is dusky-brewn. The egg is “14% in.
by +3 in. ; the ground-colour is dusky-tawny, marked with lines and
Proc. Zoou. “Soe. —1878, No. XVI. 16
242 MR. D. G. ELLIOT ON THE PTEROCLIDE. [ Feb. 19,
blotches of umber, especially towards the base of the egg, where
these marks form an encircling zone.”
Male. Black stripe from nostrils to the eye; sides of head, neck,
and throat pale buff; a black line encircles the lower part of the
throat, and extends onto the back of the neck, where it is broadest.
Top of head dark brown; back of neck pale greenish yellow.
Breast dark greyish buff, with a wash of green. Entire rest of
underparts chestnut. Back and inner secondaries greenish grey.
Wing-coverts brownish grey, the greater ones tipped with ochra-
ceous. Primaries dark brown, with pale edges. Median rectrices
greenish grey; lateral ones black, tinged with ochraceous on edge
of outer web, and tipped with light rufous. Under tail-coverts
bright chestnut. Iris deep brown. Bill and feet black.
Total length 11% inches; wing 9, tail 4%, bill along gape 3.
Female. Top of head black; feathers margined with light buff.
Back of neck buff, with a central streak of black. Feathers of back,
wings, and breast black, with a large buff spot on each web at the
tip, those on lower part of back and wings being rather circular in
form, giving those parts a spotted appearance, while the breast and
upper part of back seem to be streaked with broad black lines on a
buff ground. Secondaries black, scalloped on the edges, and tipped
with light buff. Primaries blackish brown. Flanks, abdomen,
thighs, and vent deep rufous, narrowly barred with black. Tail
black, barred with deep buff along the margins of the webs, and
tipped with reddish buff. Upper tail-coverts like the back ; under
coverts deep chestnut, barred on their basal half with black. Bill
and feet black.
Total length 11 inches, wing 9, tail 4, bill along gape $.
3. PTEROCLES CORONATUS.
Pterocles coronatus, Licht. Verz. d. Doubl. (1823) p. 65; Temm.
Planch. Color. (1825) nos. 339, 340; Riipp. Neue Wirbelth. (1823-
1835) p. 114; Wagl. Syst. Av. (1827) Pterocles, sp.4; Gray, Gen.
B. vol. ili. p. 519 (1849); Reich. Syn. Av. Gallinze, pl. ceviii. figs.
1815, 1816 (1851); Heugl. Syst. Uber. N.O.-Afr. (1855) p. 304;
Tristr. Ibis (1860) p. 71; Von Heugl. Journ. fiir Ornith. (1862)
p- 415; Hartm. Journ. fiir Ornith. (1863) pp. 303, 307; Adams,
Ibis (1864) p. 27; Allen, Ibis (1864) p. 240; Wyatt, Ibis (1870)
pp: 9, 16; Shelley, Ibis (1871) p. 144; id. B. Egypt (1872) p.
221; Hume, Str. Feath. (1873) vol.i. p. 224; Von Heugl. Ornith.
Nordost-Afr. (1873) Band ii. p. 863; Wise, Str. Feath. (1876)
vol. iv. p. 230.
Hab. Rare in Egypt and Nubia (Shelley) ; southern parts of the
Sahara (Tristram) ; El Kab, Egypt (Allen); Nubia and Kordofan
(Heuglin) ; Nubia and Arabia (Riippel/); Plain of El Goa, Sinai
(Wyatt); Jacobabad, N.W. Sindh (Hume); Kurrachee district
(Wise).
According to Tristram this Sand-Grouse is confined to the more
southern portions of the Sahara, where it supplants P. arenarius.
He only met with it in parties of four or five; and the egg was of an
1878. ] MR. D. G. ELLIOT ON THE PTEROCLID RE. 243
ashy white with a few almost obliterated pale-brown markings. In
Egypt Capt. Shelley found this species to be rare, and also in Nubia ;
and he does not think it goes into the Delta. Dr. Adams met with it
at the Second Cataract; and when at El Kab, Mr. Allen killed a
pair out of a flock that came down to drink at a pool. Their ery
was hoarser than that of the common species (P. exustus?),
Male. A line on each side of the forehead, from the nostril to
above the eye, chin, and a narrow line down the centre of the throat
Jet black. A pearly grey band on each side of the occiput, meeting
at the back of the neck. Centre of forehead white. ‘Top of head
vinous. Sides of neck, throat, and lower part of neck behind
orange. Upper parts and wings vinous, with the tips of the wing-
coverts pale buff, surrounded by a dark grey margin on the inner
side. Primaries dark brown, with the tips of the 5th to 9th buffy-
white; shafts of first two pure white, of the rest brownish white.
Entire underparts pale sandy-buff, darkest on the middle of abdo-
men and vent. Median rectrices not extending beyond the rest,
vinous. Lateral feathers pale rufous, with a subterminal bar of
black, and white tips. Bill bluish. Feet dark brown.
Total length 10 inches ; wing 73, tail 34, bill along gape 3.
Female. Throat and sides of the neck orange, palest on the chin.
Entire upper parts sandy buff with a vinous tinge, barred narrowly
with black. Wings paler buff, irregularly barred with fine black
lines, broadest and most conspicuous on the scapulars and some of
the inner secondaries. Breast and underparts very pale buff, almost
white upon the flanks, each feather margined with pale brown.
Under tail-coverts yellowish white. Tail vinous, with a subterminal
black bar, and white tip.
Total length 10 inches, wing 73, tail 34, bill along gape 3.
4. PrEROCLES SENEGALUS.
Tetrao senegalus, Linn. Mantissa (1771) p. 526; Lath. Ind. Orn.
vol. ii. p. 642.
La Gélinotte de Sénégal, Buff. Planch. Enlum. p- 130.
Pin-tailed Grous, var. A, Lath. Gen. Syn. (1783) p. 748.
Senegal Grous, Lath. Gen. Hist. B. vol. viii. (1823) p. 253.
Pterocles guttatus, Licht. Verz. d. Doubl. (1823) p. 64; Riipp.
Neue Wirbelth. (1823-1835) p. 114; Wagl. Syst. Av. (1827), Pée-
rocles, sp. 8; Less. Trait. Ornith. (1831) p. 517. sp. 8; Allen, Ibis
(1864) p. 240; Gould, B. Asia, pt. ui. pl. 6; Hume, Ibis (1872)
p- 468; Von Heugl. Ornith. Nordost-Afr. Band ii (1873) p. 860.
Pterocles senegalensis, Shaw, Nat. Misc. vol. iv. pl. 933 ; Taylor,
This (1859) p. 50, (1867) p- 67; Heugl. Syst. Uebers. Vig. Nordost-
Afr. (1855) p. 304; Speke, Ibis (1860) p. 247; Tristr. Ibis (1868)
p- 211; Taczanow. Journ. fiir Ornith. (1870) p. 51; Shelley, Ibis
(1871) p. 144.
Pterocles senegalus, Gray, Gen. B. vol. iii. p- 519 (1845); Tristr.
Ibis (1860) p. 71.
Pteroclurus senegalus, Bon. Compt. Rend. (1856) vol. xlii, p-
880; Reich. Syn. Av. Galline (1851) pl. cex. figs. 1831, 1832.
16
244 MR. D. G. ELLIOT ON THE PTEROCLIDA. [Feb. 19,
Pterocles senegalus, Shelley, B. Egypt (1872) p. 220; Hume,
Str. Feath. (1873) vol. i. p. 221; James, Str. Feath. vol. iti. (1875)
p. 418; Butler, Str. Feath. (1875) vol. iii. p. 4, (1877) vol. v.
p- 231.
Fukee, Somali (Speke).
Hab. Egypt (Shelley) ; extreme south of the Sahara (T'ristram) ;
Somali country, East Africa (Speke); Egypt and Nubia ( Riippell) ;
Palestine (Tristram) ; Sindh, Northern Guzerat, along the shores
of the Runn of Cutch (ume).
Canon Tristram states that this species is confined to the extreme
south of the Sahara, but is more plentiful there than P. coronatus.
He obtained one nest; and the egg had a ground-colour similar to
that of P. alchata, but was scarcely more than half the size, and
had very faint brown spots. .
In the Somali country Capt. Speke met with this Sand-Grouse on
the plateau in large flocks. In Sindh Mr. Hume found this species
in certain localities to be very numerous. Their habits are very
much like those of other Sand-Grouse, keeping together in parties
of from fifty to sixty ; and very often in winter a flock consists of one
sex only. They trot about on the dry soil, picking up seeds and
insects, or squat and enjoy the warmth of the morning sun. They
seemed to be less pugnacious than some of the other species, as,
although he watched them closely, he did not observe the perpetual.
skirmishing among the males, such as he had noticed took place with
those of P. arenarius in the Punjaub. It is probable that this
species breeds in Sindh. '
Male. General plumage sandy buff. Lores and a band encircling
the back of the head pearly grey. Underparts sandy-buff, washed
with grey on the breast. Middle of abdomen black. Basal portion
of scapulars, greater wing-coverts, and secondaries dark brown,
tips buff. Primaries pale buff, graduating into brownish black
towards the tips, which are buff. Central rectrices lengthened
beyond the rest, pale buff, apical half black; lateral ones greyish
brown, with a subterminal black band and white tips. Bill and feet
lead-colour.
Total length 122 inches, wing 73, tail 3, bill along gape 2.
Female. Throat and sides of neck orange, lighter on the chin.
Entire upper parts and wings sandy~buff, covered with black spots.
Primaries sandy-buff, graduating into dark brown near the ends, and
tipped with pale buff. Underparts bright buff, spotted on the breast
with dark brown. Centre of abdomen black. Upper tail-coverts
like the back. Median rectrices buff, faintly barred with black,
their elongated tips brownish-black. Lateral feathers pale brown,
lightest on the outer webs, with a subterminal brownish-black bar,
and white tips. Bill dark horn-colour. Feet dark brown.
Total length 113 inches, wing 8, tail 44, bill at gape 3.
5. PreRocLes VARIEGATUS.
Pterocles variegatus, Burch. Trav. 8. Afr. vol. ii. p. 345 (1822) ;
Gray, Gen. B. vol. iii. p. 519 (1849); Reich. Syn. Av. Gallinze
1878. | MR. D. G. ELLIOT ON THE PTEROCLIDS. 245
(1851) pl. cex. figs. 1833, 1834; Strickl. & Sclat. Contr. Ornith.
(1852) p. 157; Lay. B. S. Afr. (1867) p. 279; id. Ibis (1868)
p- 269, (1869) p. 375; Ayres, Ibis (1871) p. 262; Anderss. B.
Dam.-1. (1872) p. 242.
Pterocles maculosus, Smith, Ill. 8.-Afr. Zool. vol. ii. p. 345 (1822);
Gray, Gen. B. vol. iii. p. 519 (1845).
Hab. Between Latakoo and the Tropics, South Africa (Smith) ;
banks of the Limpopo and Mariqua (4yres); Damaraland (An-
dersson).
Male. Throat and sides of face pearly-grey. Top of head and
occiput deep buff streaked with black. Upper parts and wings
brownish-black, tipped with yellowish-ochre, and spotted with white ;
the tips of the greater coverts, however, are reddish-ochre. Primaries
brownish black, with light edges. Breast and entire underparts
bright rufous, each feather with a subterminal white bar. Tail
brownish-black, barred with light buff, and tipped with yellowish
ochre. Bill black, feet yellowish-brown.
Total length 93 inches, wing 62, tail 34, bill along gape 2.
Female. Top of head bright buff, streaked with dark brown.
Sides of head and throat bright uniform buff. Feathers of the back
and wings blackish-brown, tipped with light yellowish-brown, and
with two or three white bars on the webs, which do not, however,
reach the shaft, and give to the back the appearance of being spotted
with white. Primaries blackish-brown, edged at the tip with white.
Breast reddish-buff, indistinctly barred with white. Underparts
dull white, barred with pale brown. Upper tail-coverts like the back.
Tail brownish-black, barred and tipped with light buff. Under tail-
coverts uniform light buff. Bill black; feet yellowish-brown.
Total length 93 inches, wing 63, tail 34, bill along gape 4.
6. PrmRocLEs ARENARIUS,
Tetrao arenaria, Pall. Noy. Comm. Petropol. vol. xix. p. 418,
tab. 8 (1774); id. Itiner. vol. iii. App. p. 699, no. 271; id. Zoogr.
Rosso-Asiat. vol. ii. p. 73, pl. (1831); Lath. Ind. Orn. vol. ii.
p- 642 (1790); Bonnat. Ency. Méth. tom. i. p. 200 (1790).
_ Sand-Grous, Lath. Syn. vol. ii. pt. 2. p. 751 (1783) ; id. Gen. Hist.
B. vol. viii. (1823), p. 254.
Tetrao fasciatus, Desfont. Mém. de Y Acad. des Se. p. 502 (1787).
Aragonian Partridge, Lath. Syn. Suppl. p. 223 (1787).
Perdiw aragoniea, Lath. Ind. Orn. p. 645 (1790); id. Gen. Hist.
B. vol. viii. (1823). p. 256 B.
_ Pterocles arenarius, Temm. Pig. et Gallin. vol. iii. (1815) pp. 240,
712; id. Planch. Col. nos. 52, 53; id. Man. Ornith. (1820)
vol. ii. p. 476, vol. iv. p. 330; Wagl. Syst. Av. (1827) Péerocles ;
Less. Trait. Ornith. (1831) p. 516, sp. 1; Fras. Proc. Zool. Soe.
(1839) p. 121; Gray, Gen. B. vol. iii. p. 518 (1849) ; Blyth, Cat.
B. Mus. Asiat. Soc. (1849) p. 249; Reich. Syn. Av. Gallinze (1851)
pl. cevii. figs. 1809-12; Adams, Proc. Zool. Soc. (1858) p. 501;
Salv. Ibis (1859) p. 352; Tristr. Ibis (1860) p. 69, (1868)
p- 211; Irby, Ibis (1861) p, 223; Jerd. B. Ind. vol. iii. p. 496;
246 MR. D. G, ELLIOT ON THE PTEROCLID&. [Feb. 1
Bree, B. Eur. vol. iii. p. 226, pl. (1862) ; Tristr. Proc. Zool. Soc.
(1864) p. 449; Lilford, Ibis (1866) p. 379; Drake, Ibis (1867)
p. 428; Smith, Ibis (1868) p. 450; Taezanow. Journ. fiir Ornith.
(1870) p.51; Saund. Ibis (1871) p. 223; Rey, Journ. fiir Ornith.
(1872) p. 154; Godman, Ibis (1872) p. 219; Garrod, Proc. Zool.
Soe. (1873) pp. 468, 639, (1874) p. 254; Hume, Str. Feath. vol. i.
(1873) p. 219; Kriper, Journ. fiir Ornith. (1875) p. 282; James,
Str. Feath. vol. ii. (1875) p. 418; Butler, Str. Feath. (1876) vol.
iv. p. 4, (1877) vol. v. p. 231; Dresser, Ibis (1876) p. 322; Scha-
low, Journ. fiir Ornith. (1876) p. 186; Scully, Str. Feath. (1876)
vol. iv. p. 1792; Danf. Ibis (1877) p. 273.
Tetrao subtridactyla, Hasselgq. It. p. 250.
Tuturuk, Boora-kurra, in Afghanistan (Jerdon). Buht-titar,
Bakht-tit, Buklit, also Ban-chur-Kurmoor, at Peshawur (Jerdon).
Ll Koudhre, Arabic (Tristram). Corteza, near Seville (Saunders).
Ortega Churra, in Spain (Lilford). Barrigra negra in Portugal
(Smith). Beghitak, Yarkand (Scully).
Hab. Desert regions of Asia, Southern Europe, and North Africa
(Blyth) ; plains of Fuerteventura, Gran Canary (Godman); Harak-
ta plains of the Eastern Atlas, Djendeli, Madracen (Sa/vin) ; Sahara
desert everywhere, except in the extreme south (Tristram); Erze-
roum (Dickson and Ross) ; Asia Minor (Danford) ; Palestine (Tvis-
tram); very common in Jodhpoor, not uncommon in Sindh, fairly
plentiful in eastern portions of Cutch, all along the eastern shores of
the Runn, and in the north-eastern portions of Kattiawar (Hume) ;
Punjaub, N.W. Provinces, Bengal (Adams), Besharik, Yarkand, and
Koshtak in Eastern Turkestan (Scully) ; Western Persia (De Filippi).
When in Yarkand Mr. Scully states (/. c.) that he heard of a bird
called Beghitak by the natives, which lived in the desert ground,
was yellowish brown in colour, and had feathered legs, and long
pointed wings. Its blood was reputed a specific for consumption.
Although he did not succeed in obtaining a specimen, yet once near
Besharik he saw a pair at a distance, and the next day he saw three
more, which appeared to be yellowish brown above, breast dove-
colour, abdomen dark or black, and lower tail-coverts white. They
were so wild and wary that he could not get a shot at them. Al-
though he refers these birds to P. arenarius, it is more probable, as
suggested by Mr. Hume, they were Syrrhaptes paradoxus.
Jerdon, in the ‘ Birds of India,’ says that this Sand-Grouse is only
found in the north-west provinces and Sindh, rarely extending so
low as Allahabad, is tolerably abundant in the Punjaub, and stated
to be very abundant on the edges of the great desert. He had also
heard of its having been killed near Nusserabad and in Khandeish.
It arrives at the end of September, and leaves in March, and is there-
fore only a winter visitor in India. It frequents sandy plains, flies
in vast flocks, and, like others of the family, it regularly visits twice
a day certain drinking-places, and is fond of basking in the sun and
rolling in the sand. The flight of this species is very strong and
swift ; and sometimes it traverses great distances. It is a very shy
bird, difficult of approach, and much esteemed as an article of food.
1878. | MR. D. G. ELLIOT ON THE PTEROCLID. 247
In flocks it is stated that one or other sex greatly preponderates ;
sometimes seven females are obtained to one male, and vice versi,
Canon Tristram states (/. c.) that this species is universally dis-
tributed throughout the Sahara, except in the extreme south, where
it gives place to P. senegalus. It has so much of the Plover-cha-
racter that he took the first covey that he saw for large Plovers.
He did not find it so wary as P. alehata; but when approached
closely it would suddenly rise to a great height and fly for a long
distance. The species lays three eggs of a pale fawn-colour, with
obsolete pale brown blotches ; they are elliptical in shape, and are
placed two in a line, and the third lengthwise outside, in a depression
in the ground. He observed that when the hen was sitting one wing
was spread out to cover the eggs ; and she thus presented a grotesque
lopsided appearance. The flesh, although extremely white, was very
poor and dry, without any flavour. In support of this testimony
Capt. Irby says, in his paper “On the Birds of Oudh and Kuma-
on,” that this bird, obtained by him near Hurdue, was uneatable
and tended to confirm what the natives say, “‘that they live upon
sand.”
Male. Top of head and occiput dark brownish grey, with a pearly
tinge, becoming a lighter grey on loral space, around the eye, and
ear-coverts. Upper part of throat and sides of neck chestnut, be-
* coming deep orange towards the back of the neck. Lower part of
.
throat in front black. Breast pearly grey, crossed on the lower part
by a rather broad black band, succeeded by another of rich buff,
which is widest on the sides near the wings. Rest of underparts
jet-black. Feathers of’ the back and wing-coverts are rich buff, each
feather with a subterminal greyish-black bar, and tipped with orange-
yellow. Secondaries deep buff, the terminal third of outer web
bright orange; some of the inner ones with a subterminal greyish-
black bar like the coverts, and some with this colour only along the
shaft. Greater wing-coverts orange-yellow on their apical third.
Primaries greyish black. Upper tail-coverts like the back; under
coverts black, tipped with white. Rectrices pale rufous, crossed
with narrow black bars, and tips white. Thighs black; tarsi
covered in front with buffy white feathers. Bill bluish. Feet
brownish. Iris brown. Total length 114 inches, wings 94, tail 41,
bill along gape 2.
Female. 'Vop of head and occiput rufous, streaked with black.
Upper part of throat and sides of head yellowish white, the feathers
of the lores and ear-coverts with a fine central black streak ; lower
part of throat crossed by a blackish band. Breast rich buff, becoming
buffy white near the abdomen, the feathers of the upper part, with
a central streak and spot at the tip, black. A black band crosses
the lower part, beneath which there are no spots. Rest of under-
parts black. Back and wings deep buff, crossed irregularly with
narrow black lines; the greater coverts tipped with orange. Secon-
daries deep buff, barred narrowly with black, and all but the inner-
most ones with a broad band of deep orange on the outer webs at
their tips. Primaries greyish black, edges whitish. Rectrices very
248 MR. D. G. ELLIOT ON THE PTEROCLIDZ. [ Feb. 19,
dark brown, crossed with narrow indistinct brownish-black lines, and
tipped with buffy white. Under tail-coverts black, tipped with
white. Thighs brownish black; tarsi covered in front with short
buffy-white feathers. Bill brownish, with a black tip. Total length
11} inches, wing 9, tail 33, bill along gape 3.
A young bird from Erzeroum has the head, back, wings, ard breast
hight buff, with a central line and one or more bars on each feather
black, some crossing in a direct line, some in the shape of arrow-
heads. Primaries greyish black, with buffy-white tips. Abdomen
and flanks black. Middle rectrices light buff, crossed with nume-
rous narrow curved black bars ; the lateral ones greyish black along
the shafts, buff on the outer margin, barred with black and tipped
with white, this last increasing in extent as it goes towards the ex-
ternal feather. Under tail-coverts brownish black, and tipped with
white.
7. PTEROCLES EXUSTUS.
Bahtah Grous, Lath. Gen. Hist. Birds, vol. viii. (1823) p. 258.
Pterocles senegalensis, Licht. (nec Shaw), Verz. Doubl. p. 64
(1823).
Pterocles exustus, Tem. Pl. Col. (1825) nos. 354, 360; Wagl.
Syst. Av. (1827), Pterocles, sp. 9; Riipp. Neue Wirbelth. (1823-35)
p- 114; Less. Trait. Ornith. (1831) p. 517, sp. 9; Sykes, Proc.
Zool. Soc. (1832) p. 154; Gray, Gen. B. vol. iii. p. 519 (1849) ;
Blyth, Cat. B. Mus. Asiat. Soc. (1849) p. 249; Reich. Syn. Av.
Gallinee (1851) pl. ecix. figs. 1823-24; Jerd. Madr. Journ. vol. xii.
p- 3; Hutt. Journ.. Asiat. Soc. Beng. vol. xvi. p. 785; Burgess,
Proc. Zool. Soc. (1855) p. 30; Von Heugl. Syst. Uebers. Vog. N.O.-
Afr. (1855) p. 304; Hartl. Syst. Ornith. W.-Afr. (1857) p. 205;
Adams, Proc. Zool. Soe. (1858) p. 502; Taylor, Ibis (1859) p. 50,
(1867) p. 67; Irby, Ibis (1861) p. 235; Jerd. B. Ind. vol. iii.
(1864) p. 502; Tristr. Proe. Zool. Soc. (1864) p. 449; Gould,
B. Asia, pt. 11. pl. 13; Adams, Ibis (1864) p. 27; Allen, Ibis
(1864) p. 240; Tristr. Ibis (1868) p. 211; Beavan, Ibis (1868)
p- 378; Blanf. Geol. & Zool. Abyss. (1870) p. 4i0; Shelley, Ibis
(1871) p. 158; id. B. Egypt (1872) p. 218; Lloyd, Ibis (1873)
p-415; Hume, Str. Feath. (1873) vol.i. p. 225; Adams, Str. Feath.
(1873) vol. 1. p. 392; Von Heugl. Ornith. Nordost-Afr. Band ii.
p- 855 (1873); id. Journ, fiir Ornith. (1875) p. 57; Butl. Str.
Feath. (1875) vol. iil. p. 4; Fairb. Str. Feath. (1876) vol. iv.
. 262.
‘ Pteroclurus exustus, Ball, Str. Feath. (1874) vol. ii. p. 426.
Bar-titar, Bakt-titar, Kumar-tit, Hind.; Kuhar, N.W. Prov.;
Jam polanka, Tel. ; Kal-Kondari, Tam. (Jerdon). Gutta in Egypt
(Adams).
Hab. N.E. Africa and Arabia (Heuglin); Egypt and Nubia
(Shelley) ; Upper Egypt (d4dams) ; Hindustan, Middle and Western
Asia, Southern Europe, North Africa (Blyth); Abyssinia (Blan-
ford); Senegal, Mus. Berl. (Hartlaub) ; Casamanze (Verreauz) ;
Nubia and Abyssinia (Riippell) ; Palestine (Tristram) ; India, gene-
1878. | MR. D. G. ELLIOT ON THE PTEROCLID. 249
rally in desert places, unknown in Malabar, in wooded districts of
Central India, in Lower Bengal, and in Assam, Sylhet, and Burmah
(Jerdon); Dukhun (Sykes) ; Punjaub (Adams) ; Ahmednagar dis-
trict, Khandala (Fairbanks); Chotah Nagpur (Ball); Oudh
(Jrby) ; common in Sindh (Hume) ; Sambhur lake (ddams) ; Um-
ballah and Morar (Beavan) ; Kattiawar (Lloyd).
This species, Capt. Shelley states (/. ¢.), is the most abundant of
the various kinds of Sand-Grouse met with in Egypt, and ranges
throughout that country, as wellas Nubia. It goes in small flocks,
uttering, when on the wing, a peculiar loud note which is heard for
a considerable distance in the morning and evening, when on their
way to their drinking-places. The localities in which he found them
most abundant were Fayoom, Golosaneh, Karnak, and between
Silsilis and Kom Ombo. Tts nest is simply a hole in the sand, lined
with dried grass ; and it breeds in April.
According to Jerdon this is the most common Sand-Grouse found
in India, and is met with in every part of the country, except in the
wooded districts. It frequents the open plains and ploughed lands
or fallow fields. This species feeds early in the morning, and be-
tween eight and nine o’clock goes to drink, and again visits the water
at about four o’clock in the afternoon. At such times thousands
congregate at these places; and they come from all quarters, flying
at a great height, and announcing their presence by a peculiar loud
piercing call. They stay but a moment at the water-side, and then
fly off. . They are difficult to approach, and when on the ground can
hardly be distinguished from it, the colour of their plumage assimi-
lating so closely. This Sand-Grouse feeds on various hard seeds,
especially of the Alysicarpi, Desmodium, &c., as well as on grass-
seeds and grain. It breeds in the Deccan and Southern India from
December to May, and still later in Central India. The eggs are
laid on the bare ground, and are of a greenish colour, thickly spotted
with grey and brown, and nearly of the same thickness at both ends.
If kept some time after being killed, this bird is very good as an
article of food. In Oudh, says Capt. Irby, this species is very
common on the sandy plains from January to July. Two eggs,
which he found in June, resembled in size and markings those of
Caprimulyus europeus; and if there had been any trees near the
place, he would: have. referred them to some species of Goat-
sucker. Dr, Adams says that this bird, when wounded, spreads out
its tail and wings, hiding its head under the breast, and emits a series
of chuckles, like a fowl. As with the same Species in India, he
found the flesh very dry, and the only possible way of making them
fit for the table was to remove the skin before cooking them.
Male. Head, neck, entire upper parts, and breast sandy buff,
lightest on the breast, and having a greenish shade on the back.
Throat orange. Across the breast at the shoulders is a narrow black
band, occasionally edged on its upperside with white. Rest of
underparts bright chesnut, with the centre of the abdomen black.
Wings like the back, each feather margined at the tip with a narrow
line of blackish chestnut, and the outer webs of secondaries and
250 MR. D. G. ELLIOT ON THE PTEROCLIDE. [ Feb. 19,
greater coverts yellow on the terminal third. Some of the feathers
also have a smail white spot just beneath the dark line on the tip.
Primaries brownish black ; the inner ones tipped with white, broadest
on the inner web. Central rectrices much lengthened beyond the
rest, deep buff, indistinctly mottled with dark brown near the base,
with an irregular scalloped subterminal black bar, then buffy white,
and the lengthened narrow apical portion black. Lateral ones with
the inner webs dark rufous brown; outer ones buff, barred with
blackish brown, a subterminal black bar, and white tips. Under
tail-coverts brownish black, with white tips. Thighs chestnut, tarsi
covered in front with short buffy-white feathers.
Total length 92 inches, wings 7, tail 33, bill along gape 3.
Female. Top of head buff, darkest in the centre, each feather with
a central line of black. Throat and loral space pale orange-yellow.
Sides and back of neck and breast bright buff, each feather with a
wedge-shaped central black line, the widest part being at the tip,
and also with another black bar crossing it in the centre. The fea-
thers on the lower part of the breast are tipped with black, margined
on the upperside with white, forming an irregular band across,
below which there is an unspotted light buff space. Flanks, abdomen,
thighs, and vent black, narrowly barred with rufous. Upper parts
bright buff, barred narrowly with jet-black. Wing-coverts like the
back, the lesser ones tipped with blackish chestnut, the greater ones
broadly tipped with light yellow, forming a broad band on lower side
of the wing when closed. Secondaries light buff, barred with black,
the tips of outer webs of some of the outer secondaries light yellow.
Primaries chocolate-brown, broadly tipped on inner webs with white.
Upper tail-coverts like the back. Median pair of rectrices extended
beyond the rest, attenuated for their terminal third, which is brownish
black, the remaining portion being bright buff, barred narrowly with
black ; lateral feathers dark brown on inner webs, bright buff on
outer, with numerous bars of black, and tips white. Under tail-
coverts yellowish white, barred with black. Bill brownish black.
Total length 11? inches, wing 7, tail to end of central rectrices
42, bill along gape 2.
8. PreROCLES ALCHATA.
The Little Pin-tailed Grouse, Edw. Glean. p. 84, pl. 249 (1758).
Tetrao alchata, Linn. Syst. Nat. (1776) vol. i. p. 276; Gmel.
Syst. Nat. (1788) vol. i. p. 754; Lath. Ind. Orn. vol. ii, p. 641
(1790) ; Bonnat. Ency. Méth. (1790) tom. ii. p. 203, pl. 92.
fig. 3.
Pin-tailed Grous, Lath. Gen. Syn. (1783) vol. ii. pt. 2nd,
». 748.
Pterocles setarius, Temm. Pig. et Gallin. vol. iii. pp. 256, 714
(1815); id. Mon. Ornith. vol. ii. p. 478 (1820), vol. iv. p. 330; Wagl.
Syst. Av. (1829) Pterocles, sp. 6; Less. Trait. Ornith. (1831) p.
516. sp. 6; Parker, Proc. Zool. Soc. (1864) p. 258; Taczanow. Jour.
fiir Ornith. (1870) p. 51.
Gnas cata, Vieill. Nouv, Dict. Hist. Nat. vol. xii. p. 418 (1817),
or
_
1878. ] MR. D. G. ELLIOT ON THE PTEROCLID. 2
Pin-tailed Grous, Lath. Gen. Hist. B. vol. viii. (1823) p. 250.
Pterocles alchata, Licht. Verz. Doubl. p. 64 (1823) ; Tristr. Proc.
Zool. Soc. (1864) p. 449; Jerd. B. Ind. vol. iii. p. 500 (1864) ;
Gould, B. Eur. pl. 258; Blyth, Cat. B. Mus. Asiat. Soc. (1849) p.
249; Gray, Gen. B. vol. iii. p. 518; Reich. Syn. Av. (1851) Gallinee,
pl. cevii. figs. 1813-14; Heugl. Syst. Uebers. Vog. Nordost-Afr.
(1855) p. 303; Salv. Ibis (1859) p. 352; Tristr. Ibis (1860) p. 70;
Powys, Ibis (1860) p. 239; Wright, Ibis (1864) p. 139; Lilford,
Ibis (1866) p. 379; Bartl. Proce. Zool. Soc. (1866) p. 78; Drake
Ibis (1867) p. 428; Beaven, Ibis (1868) p. 377 ; Smith, Ibis (1868)
p- 451; Hume, Str. Feath. (1873) vol.i. p. 221; Garrod, Proc. Zool.
Soe. (1873) p. 468; Severzov, Journ. fiir Ornith. (1873) pp. 385,
386; Von Heugl. Ornith. Nordost-Afr. (1873) vol. ii. p. 853.
Tetrao chata, Pall. Zoogr. Rosso-Asiat. vol. ii. p. 73 (1831).
Pterocles caspius, Ménétr. Cat. Rais. de Zool. p. 47 (1832).
Pteroclurus alchata, Bon. Compt. Rend. vol. xlii. p. 880 (1856).
El-Guettha Arab (Tristram). Ganga inSpain (Lilford). El Koudre
in Tangiers (Drake). Corticol in Portugal (Smith).
Hab. Middle Asia, Afghanistan, Syria, Spain, N. Africa (Blyth) ;
Portugal (Smith) ; common in Spain (Lelford) ; Tangier (Drake) ;
Harakta Plains of Eastern Atlas (Salvin) ; Malta (Wright); Pales-
tine (Tristram) ; Punjaub and Sindh (Jerdon); Jacobadad, Sindh,
Nundan, near Altock, Peshawur valley, Abbotabad, and isolated
localities in Huzara (Hume); Western Persia (De Filippi).
Jerdon says that this is a rare bird in India, only a few finding
their way across the Sutlej. It flies in flocks, and is shy and wary.
The specific appellation is taken from its Arabic name El-chata
or Ll-katta; and in India there does not seem to be any special
native name for the bird.
Tristram found this Sand-Grouse in vast flocks in winter in the
Mzab and Tonarick country ; and, except in the breeding-season,
it was very difficult to approach, and when packed it is necessary
to be well-mounted to get a second shot. It is even more vigo-
rous upon the wing than P. arenarius, and when on the ground
is very garrulous. Its breeding-habits are exactly the same as
those of the species just named; but the egg has a much richer
fawn-coloured tint, and is covered and sometimes zoned with large
maroon-red blotches.
In Spain, as stated by Lord Lilford (J. c.), this species, as well as
P. arenarius, is common in many parts of the Castilles, La Mancha,
Murcia, Andalusia, Estremadura, and in some districts of Aragon
and Catalonia. They delight in bare and stony ground; and even
in “hungry Spain” they are considered ‘ muy male carne.”
Male. Top of head and nape dark greyish brown ; superciliary
stripe and sides of the throat deep orange; ear-coverts yellowish;
a black line goes from behind the eye and down the sides of the
nape. Throat black; upper part of breast greenish buff; rest of
breast chestnut, bounded above and below with a narrow black line ;
remainder of underparts white. The back has the feathers vinous at
base, rest pale yellowish green, with a yellowish spot at tip.
252 MR. D. G. ELLIOT ON THE PTEROCLID. [Feb. 19,
Rump and upper tail-coverts buff, barred narrowly with black.
Lesser wing-coverts pale vinous at base, then a narrow line of white
followed by a broader one of pale chestnut, and then a narrow line
of yellow, and tipped with pale chestnut. Some of the outer
larger wing-coverts were like those just described ; but the imer
ones are dark, have the base dark ash, then rich yellow, and narrowly
bordered with black. Inner secondaries rufous at base, barred
with black, tipped with pale green, and a conspicuous yellow spot
in the centre. Rest of secondaries pale yellowish-green, with
yellow on their outer webs near the tips. Primaries ashy grey,
margined on inner webs with white; the secondaries and tertiaries
inclined to buff along the shaft on inner webs; shafts black. Tail-
feathers. buff, barred with black and tipped with white; the two
central ones entirely black on their apical half, are extremely
narrow, and extend far beyond the other rectrices.. Under tail-
coverts buff, barred with black, and tipped with white. Feathers of
the tarsi. white. Bill horn-brown. Bare space around the eye lead-
colour; iris dark brown.
Total length 1314 inches, wing 8, tail to end of longest rectrices
6, bill along gape +4.
Female. Top of head and back of neck deep buff, barred with
black. Sides of face and neck orange-yellow. Chin and throat
white. A black band around the lower part of neck, but not
extending onto the back of the neck, beneath which is a broad
light chestnut band bordered above and below with black as in the
male. Enutire underparts pure white. Upper parts buff, narrowly
barred with black. Wing-coverts buff, barred with black, and with
a grey spot on inner web near tip, this spot on some of the
greater coverts being enlarged into a bar crossing the feathers near
the tip. Secondaries buff, barred with black, and having a sub-
terminal grey bar. Primaries like those of the males. Upper tail-
coverts yellowish buff, irregularly barred with black. Rectrices like
those of the male. Bill horn-colour.
Total length 13 inches, wing 73, tail 53, bill at gape 3.
9. PreEROCLES NAMAQUA.
Namaqua Grous, Lath. Gen. Syn. B. (1783), vol. ii. pt. 2, p. 750 ;
id. Suppl. p. 215; id. Gen. Hist. B. vol. viii. (1823) p. 251.
Tetrao namaqua, Gmel. Syst. Nat. (1788) vol. i. p. 754; Lath.
Ind. Orn. vol. ii. p. 642, sp. 19; Sparrm. Voy. vol. i.p. 153; Bonnat.
Enc. Méth. (1790) tom. ii. p. 204.
Pterocles tachypetes, Temm. Pig. et Gall. (1815) pp. 274, 715;
Wagl. Syst. Av. (1827), Pterocles, sp. 7; Less. Trait. Ornith. (1831)
p. 517, sp.7; Layard, B. S. Afr. (1867) p. 277; Ayres, Ibis (1871)
p. 262.
Gnas namaqua, Vieill. Nouv. Dict. Hist. Nat. (1817) vol. xii.
», 422.
Pterocles simplex, Less. Trait. Ornith. (1831) p. 517.
_ Pterocles namaqua, Gray, Gen. B. vol. iil. p. 518; Bocage, Journ.
fiir Ornith. (1876) p. 303.
1878. ] MR. D. G. ELLIOT ON THE PTEROCLID#. 253
Pteroclurus namaqua, Reich. Syn. Av. Galline (1851) pl. CCix. ;
Anderss. Dam.-l. (1872) p. 242.
Hab. Palestine (Tristram) ; Transvaal, between Potchefstroom
and Marico (Ayres) ; Damara-land (Andersson).
Mr. Ayres first met with this species in the Transvaal between
Potchefstroom and Marico, and afterwards along the river Limpopo,
but nowhere very abundant. They squat on the approach of danger,
and rise suddenly, with a quick beat of the wings; and their flight
is very rapid. The iris is dusky, and eyelids yellow.
According to Layard, the Namaqua Grouse is very abundant on
the Karroo plains throughout the Cape colony and in Namaqua-
land. It congregates in small parties, runs with considerable
swiftness, and the flight is very rapid. Its note, when on the wing,
resembles the piping of a Plover, to which bird this species bears a
considerable resemblance. It sometimes, during droughts, extends
its migrations to the Cape Flats. It lays from seven to ten eggs,
light cream-colour, spotted with -brown and purple, axis 17",
diameter 12'’, Andersson states that this bird is common in some
parts of Damara-land, and generally comes to the water about eight
or nine o’clock in the morning. They circle around at a consi-
derable height, and frequently do not attempt to descend until
directly over the spot, when they shoot down with great velocity,
describing more or less of semicircles before they alight. They feed
on seeds, berries, and small bulbs, and swallow gravel to assist the
digestion.
Male. Top of head dark brown. Throat orange. Upper part of
breast vinous, crossed on the lower part by a band of white, followed
by another of reddsh chestnut. Flanks dark vinous grey. Middle
of abdomen chestnut ; vent and under tail-coverts yellowish white.
Upper parts dark brown, lighter on the rump and upper tail-
coverts, and washed with green. Scapulars dark brown, with a
subterminal light buff bar and publish rey tips. | Wing-coverts
ashy brown, subterminal buffy white bar, and chestnut tips. Secon-
daries dark vinous brown, inclining to light buff near their tips.
Primaries rufous brown; shafts rufous, except those of first and
second, which are white, and the sixth to tenth broadly margined
with white on their inner webs towards their tips. Median rectrices
extending slightly beyond the rest, rufous brown, graduating into
dark brown on their attenuated portion ; rest of tail rufous brown,
and tipped with yellowish white. Bill horn-colour. Feathers on
tarsus buffy white.
Total length 104 inches, wing 7, tail 44, bill along gape 4.
Young male. Differs in having the entire underparts very deep
vinous buff, inclining to rufous on the abdomen, and ochraceous on
the vent and under tail-coverts. Feathers of the back and wings
barred with rufous and black irregularly, and tipped with greyish
purple on the scapulars and some of the inner secondaries. Tail
buff, barred narrowly with dark brown. Head, throat, and back of
neck as in the adult males.
Female. Top of head, back, and sides of neck and breast deep
254 MR. D. G. ELLIOT ON THE PTEROCLID. [ Feb. 19,
buff ; feathers of the first with a central streak of black ; those of the
other parts barred irregularly with the same colour. Rest of under-
parts buffy white, graduating into light buff upon the flanks and
lower tail-coverts, and all, except the last-named, barred narrowly
with black. Upper parts buff, barred irregularly with narrow lines
of black, and some of the scapulars tipped with greyish purple as
in the male. Primaries same as the male. Median pair of rectrices
slightly elongated beyond the rest, reddish-buff, barred narrowly
with black, and with black tips; the lateral feathers blackish brown,
with irregular reddish-buff bars on the outer webs, and with yellowish-
white tips. On the inner webs the buff bars are very faintly
indicated, and on some of the feathers they are obsolete. Bill
horn-brown.
Total length 94 inches, wing 63, tail 44, bill along gape 4.
10. PrEROCLES FASCIATUS.
Tringa fasciata, Scop. Delic. Faun. et Flor. Insubr. vol. ii. p. 92
(1786).
Indian Grous, Lath. Gen. Syn. (1783) p. 752; id. Gen. Hist.
B. vol. viii. (1823) p. 260.
Tetrao indicus, Gmel. Syst. Nat. (1788) vol. i. p. 755 ; Bon-
nat. Ency. Méth. (1790) tom. i. p. 201.
Perdix indica, Lath. Ind. Orn. vol. ii. p. 650, sp. 23 (1790).
Ginas indicus, Vieill. Nouv. Dict. Hist. Nat. (1817) p. 422.
Pterocles quadricinctus (nec Temm.), Less. Trait. Ornith. (1831),
p. 526, sp. 3; Sykes, Proc. Zool. Soc. (1832) p. 155; Burgess,
Proc. Zool. Soc. (1855) p. 30.
Pterocles fasciatus, Gray, Gen. B. vol. iii. p. 518; Blyth, Cat.
Birds Mus. Asiat. Soc. (1849) p. 249 ; Jerd. Ind. Ornith. pls. 10,
36; Reich. Syn. Ay. Gallin. (1851) pl. ceviii. figs. 1815-16 ;
Adams, Proc. Zool. Soc. (1868) p. 502; Jerd. B. Ind. vol. iii. p.
498 (1864); Gould, B. Asia, pt. ii. pl. 14; Beavan, Ibis (1868) p.
378 ; Lloyd, Ibis (1873) p. 415; Adams, Str. Feath. (1873) vol. i.
p- 391; Ball, Str. Feath. (1873) vol. ii. p. 426; Fairb. Str. Feath.
(1876) vol. iv. p. 262; Wise, Str. Feath. (1876) vol. iv. p. 230;
Butler, Str. Feath. (1877) vol. v. p. 231.
Handeri in the south ; Boot-bur in the north-west ; Sunda polanka
Tel. (Jerdon).
Hab. India generally, except Malabar and Lower Bengal (Jerdon);
Dukhan (Sykes) ; Punjab (Adams); Deccan (Burgess); Khandesh
(Wise); Morar, near Gwalior, Umballah (Beavan); Kattiawar
(Lloyd); Hindustan generally (Blyth) ; Sambhur lake (4dams) ;
Chota Nagpur (Ball); Ahmednagar district, Kandalia (Fair-
bank).
Jerdon (/.c.) states that this species is found over the greater part
of India, but not in Malabar and Lower Bengal, and is not abundant
anywhere. He had seen it in the Carnatic, the Deccan, and in
Central India; and Adams records it as pretty common in the
jungles around the base of the Sewalik range in the Punjaub. Un-
like other Sand-Grouse, it frequents thickets, and chiefly bushy and
1878. | MR. D. G. ELLIOT ON THE PTEROCLIDE. 255
rocky hills. ~ It is generally met with in pairs, towards the end of
the rains in parties of eight or ten, and when flushed rises with a low
chuckling call, and after a short low flight drops again into cover. The
eggs, cylindrical in form, are a dull earthy green, with a few dusky
spots. It was procured in Kattiawar, Western India, by Capt. Lloyd,
in the Than district, where one evening a small party, coming to
drink after dusk, alighted in front of his tent. He thought they
were Goatsuckers ; and on firing at them was surprised to find he
had secured a specimen of this Sand-Grouse. In the vicinity of
Sambhur lake, Central India, Mr. Adams found this species common
about all the low ranges of hills. The nest was simply a hollow scraped
in the ground, with anumber of small pieces of stone round the edge,
and some loose grass for a lining. The number of eggs varied from
two to three ; in one instance, however, four were found. He gives
their colour as very different from that mentioned by Jerdon, viz. a
deep to pale salmon-colour when fresh ; but, on being blown, in a
few days this tint changes to a rich cream-colour, and the spots are
light lavender and rusty. They are of a blunt oval form, 1:4 inch in
length by 1 inch in breadth.
Male. A conspicuous black band crosses the forehead, below which,
covering the nostrils and also over each eye, is a pure white spot.
Top of head and occiput buff, streaked with narrow black lines.
Sides of head, throat, neck, and upper part of breast uniform sandy
buff. Upper part of back similarly coloured, but darker. Across
the breast, extending from the shoulders of the wing, are, first, a
band of deep chestnut, widest in the centre, succeeded by a broader
one of light buff, beneath which is an irregular one of black. Rest
of underparts black, narrowly banded with white or buffy white.
Feathers of back and rump generally crossed by three black bars,
and tipped with sandy buff; and between the first and second black
bar from the tip is one of white. Shoulders uniform sandy buff.
Wing-coverts and outer secondaries crossed by several black and
white bars, forming two or three conspicuous bands across the wing,
the tips of all the feathers being fulvous. The inner secondaries
are buff, barred only with black, no white. Primaries brownish
black, edged on both webs with whitish. Tail and under tail-
coverts fulvous, barred with black. Bill red, orbital skin lemon-yel-
low ; irides dark brown. Feet yellowish. Total length 94 inches,
wing 7, tail 37, bill at gape 3.
Female. Throat and cheeks pale buff. Head reddish buff, streaked
with black. Entire upper and under surface sandy buff, barred
irregularly with brown. Tips of wing-coverts pale buff. Primaries
_ and secondaries dark brown, edged on inner webs with white. Bill
red. Dimensions about the same as the male.
11. Prerocies BIcINCTUus.
La Gélinotte des Indes, Sonner. Voy. aux Indes Orient. et & la
Chine, vol. ii. pl. 96 (1782).
Péerocles bicinctus, 'Temm. Pig. et Gall. tom. iii. pp. 247, 718 ;
Gray, Gen, B. vol. iii, p. 518.
256 MR. D. G. ELLIOT ON THE PTEROCLIDS. [Feb. 19,
Gnas bicincta, Vieill. Nouv. Dict. Hist. Nat. vol. xii. (1817) p.
421; id. Gal. Ois. vol. ii. (1834) p. 60 (text).
Double-banded Grouse, Lath. Gen. Hist. B. vol. viii. (1823) p.
259.
Pterocles bicinctus, Wag). Syst. Av. (1827) Pterocles, sp. 2; Less.
Trait. Ornith. (1831) p. 516, sp. 2; Reich. Syn. Av. Galline
(1851) pl. ceviii. figs. 1819-20 ; Strickl. and Sclat. Contr. Ornith.
(1852) p. 157; Layard, Birds S. Afr. (1867) p..278 ; Ayres, Ibis
(1869) p. 298; Gurn. Anderss. Birds Dam. L. (1872) p. 241;
Buckley, Ibis (1874) p. 385; Bocage, Journ. fiir Ornith. (1876)
p. 303.
Hab. Transvaal, near the Limpopo (Ayres) ; Limpopo, to the
Matabili country (Buckley); Damara and Great Namaqua Lands
(Andersson).
This is the most common of the Sand-Grouse found near the
Limpopo, according to Mr. Ayres, with perhaps the exception of P.
gutturalis. The greater number in June were in flocks, although
some few had paired and were breeding. The eggs are laid on the
bare ground, and are brownish pink, spotted and blotched all over,
particularly at the thick end, with a lighter tint of the same colour.
Size 155 by 12 inch. This species was also found to be very abun-
dant from the Limpopo to the Matabili country by Mr. Buckley ;
and in the evening they come down in immense flocks to the water-
holes to drink. During the day he met with them chiefly in pairs,
or three individuals together ; and when they rose from the ground
they uttered a creaking note. According to Andersson, this species
is very abundant in Damara and Great Namaqua Lands, and can
always be met with during the dry season at any of the permanent
watering-places to which they resort at morning and evening. When
dispersed they usually go in pairs, and feed chiefly onthe seeds of
grasses and berries, mingling with their food considerable quantities
of coarse sand.
Their flesh, though very white, is excessively tough, and it is best
to skin the bird before cooking. The iris is deep red, the skin round
the eye chrome-yellow.
The figure given by Sonnerat in his ‘ Voyage aux Indes Orien-
tales et A la Chine’ is evidently referable to this species ; from having
only two bands upon the breast, a white one succeeded by a black one,
it cannot possibly represent P. fasciatus, Scop., to which it has
been referred by some authors, In his text Sonnerat only mentions
one band, the white one, which does not describe any species of
Sand Grouse at present known. The locality, ‘‘ céte de Coromandel,”
however, is not one of the habitats of the present species, so far as I
am aware, as it is apparently solely found in Africa.
The specimens described by Temminck were given to him by Le
Vaillant, who procured them in Namaqualand.
Male. Black bar across the forehead, in front of which and over
each eye is a white spot, these latter sometimes coalescing and form-
ing a white bar behind the black one. Top of head and occiput
rufous, streaked with black. Throat, sides of neck, upper part of
1878. ] MR. D. G. ELLIOT ON THE PTEROCLID &. 257
back, breast, and shoulders of the wing uniform sandy buff, with a
greenish tinge. Lower part of breast crossed by two rather broad
bands, the upper one pure white, the lower jet-black. Rest of under-
parts barred narrowly with buffy white and brownish black. Feathers
of the back rufous, barred with greyish black, those on the rump
tipped with yellowish white. Greater wing-coverts and secondaries
brownish grey, with a white Spot at the tip of each feather. Pri-
maries rufous-brown, all but the first two tipped with white. Tail,
with the upper and under coverts, light buff, barred with brownish
black. Bill reddish ; feet yellow.
Total length 92 inches, wing 63, tail 34, bill (along gape) 2.
Another, younger male specimen differs in having a narrow black
line over each eye, between it and the white mark, and also by
having the back and inner secondaries deep rufous,i rregularly barred
with black, the wing-feathers tipped with buffy white. Outer secon-
daries blackish brown, like the primaries, some tipped on outer web
with light buff, others with a subterminal bar of the same colour.
The greater wing-coverts are blackish brown tipped with yellowish
white.
Female. Head, back, neck, and breast buff, irregularly barred
with brownish black, and tipped with buffy white on.the back and
breast. No band across the breast. Flanks, abdomen, and vent
barred with white and black. Wings irregularly barred with black
and reddish buff, the feathers tipped with white ; primaries choco-
late-brown. Tail and its coverts reddish buff, barred with black and
tipped with fulvous. Bill reddish, tip black ; feet yellowish.
Total length 92 inches, wing 7, tail 31, bill along gape 2.
12. Prerocies quapRicinctus.
Pterocles quadricinctus, Temm. Pig. et Gallin. (1815) vol. iii.
pp. 252, 713; Wagl. Syst. Av. (1827), Péerocles, sp. 3; Strickl.
Proc. Zool. Soe. (1850) p. 220; id. Ann. & Mag. Nat. Hist. 2nd ser.
vol. ix. p. 347; Heugl. Syst. Uebers. Vig. N.O.-Afr. (1855) p. 304;
Hartl. Syst. Ornith. W.-Afr. (1857) p. 205.
Gnas bicinctus, Vieill. (nec T emm.) Gal. Ois. pl. 220, nec text
(1834).
Pterocles tricinctus, Swains. Birds W. Afr. (1862) vol. ii. pl. 23 ;
Von Heugl. Ornith. Nordost-Afr. (1873) Band ii. p. 867.
Hab. Sennaar, Abyssinia (Heuglin) ; West Africa (Strickland).
Some confusion has been created in the synonymy of this species
by Vieillot having figured it in his ‘Galerie des Oiseaux’ as bicinctus
(a name given by Temminck to quite a different species), while in
the text he has correctly described the true dicinctus and not the
bird represented in his plate. Although Vieillot’s figure only shows
three bands across the breast, yet Temminck asserts, in the ‘ Planches
Coloriées’ (genre Ganga, footnote), that it is his quadricinctus ;
and as it is natural to suppose he knew his own species when he saw
it, I have taken his determination as the correct one, more especially
as I have never been able to find a specimen of Pterocles with four
well-defined bands across the breast. So far as I can judge from the
Proc. Zoou. Soc.—1878, No. XVII. M7
258 MR. D. G. ELLIOT ON THE PTEROCLIDA. [Feb. 19,
specimens I have seen, the name t¢ricinctus, given to this species by
Swainson, is by far the most appropriate one ; and itis to be regretted
that it cannot be employed.
Temminck also refers to the present species the bird figured and
described by Sonnerat, who, he states, was the first to make it known
as La Gélinotte des Indes. In this determination I believe Tem-
minck to hare been mistaken, and that he confounded Sonnerat’s
bird with the P¢é. bicinctus of Temminck. The species figured by
Sonnerat has but two bands on the breast, a black and a white one ;
and I consider it to be the Pt. bicinctus, Temm., and have accord-
ingly referred it to that bird.
The only habitats I can find for Pt. guadricinctus are Abyssinia
and Sennaar on the east coast, while Swainson gives West Africa
as the place whence his specimen came. It has never, so far as I am
aware, been found in India. The examples described by Temminck
were in the cabinet of M. de Breukelewaert at Amsterdam.
Male. Spot on the forehead, covering the nostrils, and one over
each eye white. A broad black band crosses the forehead just in
front of the eyes. Top of head and occiput fulvous, streaked nar-
rowly with black. Throat yellowish white. Nape, sides of neck,
and breast uniform sandy buff. Across the lower part of the breast
are three bands, the first dark chestnut, the second white, and the
third black. Temminck says there is a fourth one, of white. I do
not perceive this in the specimens before me; but I am inclined to
think individuals vary in the number of these bars. | Underparts
black, narrowly barred with white. Back deep buff, barred with jet-
black. Wings fulvous, the shoulders uniform, not spotted. Wing-
coverts and secondaries crossed by a single black bar, margined on
both sides with white. The innermost secondaries have numerous
black bars, and xo white ones. Tail and under-coverts deep buff,
barred with black. Bill red; feet yellow.
Total length 91 inches, wing 74, tail 33, bill (along gape) 2.
Specimen described from the White Nile.
Female. The entire top of head fulvous, streaked with black, and
the back of neck spotted with the same. Throat white. Breast
bright fulvous, without any bars, and the underparts barred with
white and narrow lines of brownish black. Feathers of back and
inner secondaries rufous, barred with black and margined with fulvous.
Wings and outer webs of outer secondaries pale buff, crossed by
several narrow jet-black bars. Inner webs of outer secondaries and
primaries brownish black. Tail and its under-coverts deep buff,
barred with black. Bill reddish horn-colour ; feet yellow.
Total length 91 inches, wing 741, tail 34, bill (along gape) 3.
13. PrerRocLes LICHTENSTEINI.
Pterocles lichtensteinit, Temm. Planch. Col. (1825) no. 355, 361
Rupp. Neue Wirbelth. (1823-38) p. 114; Wagl. Syst. Av. (1827),
Pterocles, sp. 8; Less. Trait. Ornith. (1831) p. 516. sp. 8; Gray,
Gen. B. vol. iii. p. 518 (1849) partim; Reich. Syn. Av. Gallin.
(1851) pl. ccix. figs. 1827-28; Heugl. Syst. Ubers. Vog. Nord-
1878. | MR. D. G. ELLIOT ON THE PTEROCLIDE. 259
ost-Afr. (1855) p. 304; id. Ibis (1859) p. 343; Speke, [bis (1860)
p. 247; Blanf. Geol. & Zool. Abyss. (1870) p. 419; Hume, Ibis
(1872) p. 468; Finsch & Hart]. V6g. Ost-Afr. p. 563; Hume, Str.
Feath. (1873) p. 219; Von Heugl. Ornith. Nordost-Afr. (1873)
Band ii. p. 865; Wise, Str. Feath. (1876) vol. iv. p. 230.
Pterocles bicinctus, Licht. (nee Temm.) Verz. Doubl. (1823)
p- 65.
Hab. Very common in the Danakil and Somali countries (Heuglin,
Speke); Abyssinia (Blanford); Egypt and Nubia (Rippell); Gool
Mahomed Mehur, Upper Sindh (Hume); Kurrachee district (Vise).
Blanford (J. c.) states that the habits of this bird in Abyssinia
are the same as those of the Pt. fasciatus of India. It is but rarely
met with in the open plains, but keeps to bush- or tree-jungle, and
in such places is usually found in pairs. The drinking-hours are at
daybreak and at dusk; and the scene at a spring in the evening, he
says, is very interesting, the birds coming in with a constant rush until
dark. Single birds and small flocks, uttering a note like ‘“ queep-
queep,” would fly up and down the watercourse, keeping but a
few feet above the bushes, the noise of their wings announcing their
presence in the dusk before the birds themselves could be seen. Mr.
Blanford says they are excellent eating, the flesh, though rather
hard, being of a delicious flavour. The species appears to be con-
fined to the coast region. In the Lebka valley, at Mohabar, only
2000 feet above the sea, but few were seen at the spring in the even-
ing; and at higher elevations none were met with.
Mr. Hume relates, in ‘ Stray Feathers’ (1873), that he met with
this species at Gool Mahomed Mehur, Upper Sindh. Only one pair
were seen and procured.
Male. Forehead white, crossed by a circular jet-black line, com-
mencing at the opening of the mouth; over each eye a black spot.
Throat pale buff. Centre of breast fulvous, with a narrow black bar
crossing it in the middle, and another bordering it on the lower side.
Rest of plumage of body buff or buffy white, barred narrowly with
black. Wings barred with black and white and tipped with buff.
Scapulars in some specimens dark buff or rufous, barred with black,
in others like the rest of the wing-feathers just described. Pri-
maries dark brown, edged with yellowish white. Tail pale buff,
barred with black and tipped with yellowish buff, as are also the
upper and under coverts. Iris brown; orbit lemon-yellow. Bill
red ; feet flesh-colour.
Total length 10 inches, wing 74, tail 34, bill along gape 2.
Female. Throat pale buff, spotted minutely with blackish brown.
Breast, wings, and back light buff, inclining to rufous on the secon-
daries, with all the feathers crossed by numerous narrow black lines.
Primaries chocolate-brown. Abdomen, flanks, and vent dull white,
barred narrowly with black. Tail with its coverts buff, barred with
black. Bill light horn-colour ; feet yellow.
Total length 10 inches, wing 71, tail 31, bill along gape 3.
a
17%,
260 MR. D. G. ELLIOT ON THE PTEROCLIDZ. (Feb. 19,
14. PTEROCLES DECORATUS.
Pterocles decoratus, Cab. Journ. fiir Ornith. (1868) p. 413 ; id.
Von der Deck. Reis. vol. iii. p. 43, pl. 13 (1869); Finsch & Hartl.
Vog. Ost-Afr. p. 565; Von Heugl. Ornith. Nordost-Afr. (1873)
Band ii. p. 870.
Hab. Vuterior of East Africa, Lake Jipe.
I have never seen a specimen of this bird, and only know it from
Cabanis’s description and figure. He compares it to ¢ricinctus,
Swains. (=quadricinctus. Temm.), but says it is smaller, and
gives no description of the wings. It is, however, a very distinct
species, and can easily be recognized by the characters given below,
most of which are not found in any other known species.
Male. Centre of forehead white, a large spot on each side and
throat black. Top of head, neck, and back reddish grey, crossed
with narrow lines of black. Superciliary stripe buffy white. Sides
of head and breast sandy buff, with a greenish gloss, beneath which
is an irregular black bar extending upwards to the shoulders of the
wing, succeeded bya broad white band. Abdomen and flanks black,
the feathers margined with rufous.
“Total length 9; bill along gape 7!" ; wing 6°5''; tail 2! 7'"’.
Genus SyRRHAPTES.
Body large, comparatively heavy. Bill rather small, delicate,
pointed at tip ; culmen slightly curved, gonys ascending. Nostrils
basal, hidden in feathers. Wings very long, pointed, first primary
longest ; primaries in one species filamentous. Tail cuneate, median
pair of rectrices always lengthened beyond the rest, apical portion
attenuated and pointed. ‘Tarsus short, entirely covered by short
feathers. Feet tridactyle ; hallux obsolete; toes very short, com-
pletely hidden in feathers above ; the soles bare and scaly. Claws
moderate, curved and acute.
Key to the Species.
A. Median rectrices prolonged far beyond the rest.
a. Primaries filamentous. Breast uniform vinous, tinged
with green on upper part, and crossed in the centre
by an irregular bar formed of the narrow black
lines at the tips of the feathers. Abdomen blackish... 1. S. paradorus.
6. Primaries not filamentous. Upper part of breast buffy
white, irregularly barred with black ; rest of under-
parts pale buff, becoming white on the under tail-
COVORES Siird.ceectehey al Rin daaee-s shen -Roas tues eengerennie stent 2. S. tibetanus.
1. SyRRHAPTES PARADOXUS.
Tetrao paradozus, Pall. Reis. Russ. Reichs, vol. ii. App. p. 712,
tab. F (1773); Gmel. Syst. Nat. (1788) vol.i. p. 755. sp. 30 ; Lath.
Ind. Ornith. vol. ii. p. 643. sp. 20 (1790) ; Bonnat. Ency. Méth.
(1790) tom. ii. p. 205, pl. 93. fig. 1 ; Pall. Zoogr. Rosso-Asiat. vol. ii.
p. 74 (1831),
Heteroclite Grous, Lath. Gen. Syn. Birds, vol. iv. p. 753 (1783);
id. Gen. Hist. Birds, vol. viii, (1823) p. 261.
1878. | MR, D. G. ELLIOT ON THE PTEROCLID2. 261
Syrrhaptes paradocus, Ilig. Prodr. (1811) p. 243.
Syrrhaptes pallasit, Temm. Pig. et Gallin. vol. iii. pp. 282, 716
(1815) ; id. Planch. Col. 95.
Heteroclitus tartaricus, Vieill. Nouv. Dict. Hist. Nat. vol. xiv. p.
453 (1817).
Syrrhaptes paradoxus, Licht. in Evers. Reise nach Buch. p. 134
(1823); Gray, Gen. B. vol. iii. p. 519; Reich. Syn. Av. Gallin.
(1851) pl. cevii. figs. 1807-8 ; Moore, Ibis (1860) pl. iv. p. 105;
Swinh. Ibis (1861) p. 341; Newt. P. Z.S. (1861) pp. 203, 397 ;
Wortley, P. Z.S. (1861) p. 196; Parker, P. Z. S. (1862) p. 257,
(1863) p. 516; Swinh. P. Z. S. (1863) p. 306 ; Radde, Reis. S. O.-
Siber. (1863) vol. ii. p. 287, pl. xiv. fig. 3 (egg); Newt. Ibis (1864)
p. 185; Salvad. Ibis (1864) p. 228; Von Droste, Journ. fiir Orn.
(1868) p. 406 ; Mill. Journ. fiir Orn. (1869) p. 118; Von Homey.
Journ. fiir Orn. (1870) p. 170; Malmgr. Journ. fiir Ornith. (1870)
p- 295; Fritsch, Journ. fiir Orn. (1871) p. 312; Swinh. P. Z. S.
(1871) p. 398; Severzov, Journ. fiir Orn. (1873) p. 380; Von
Heugl. Journ. fiir Orn. (1874) p. 399 ; Taczanow. Bull. Soc. Zool.
France (1876) vol. i. p. 241 ; Dress. Ibis (1876) p- 322; id. B.
Eur. pt. 53 (1877) pl. ; Prjevalsky, in Rowl. Ornith. Misc. pt. ix.
p- 382; David & Oust. Ois. de la Chine (1877) p. 389.
Tetrao arenaria, Pall. Zoogr. Rosso-Asiat. (1831) pl. 53, nec
text.
Syrrhaptes heteroclita, Vieill. Gal. Ois. vol. ii. p. 64, pl. 222
(1834).
Stepnaya kuritza, Mongolia (Prejevalsky).
Hab. Tientsin, China (Wortley) ; Pekin and Tientsin (Swinhoe);
Turkestan (Severtzov) ; Mongolia (Prjevalsky); Kirghis steppes,
Donaria, plains of Petchely (David ¢ Oust.).
Pallas’s Sand-Grouse is essentially an Asiatic species, making its
home in the arid Kirghis steppes, the great Gobi desert, and east-
ward through Mongolia to the plains between Pekin and Tientsin in
North China, where it is especially numerous during the winter. The
year 1853 is the earliest in which this bird’s presence is recorded in
Europe, Moschler having met with it at Sarepta, on the Volga
(Naum. vol. iii. p. 305). In 1859 a few specimens were obtained
in various parts of Western Europe ; but in 1863 a great number of
this species, estimated at many hundreds, suddenly appeared in many
parts of Europe, constituting a veritable ornithological “Tatar”
invasion, as it was aptly called by the historian of this remarkable
event. The birds proceeded as far as Ireland, in the west, to the
Feeroes, in the north, and to Perpignan in France, to the south. Great
numbers of them were killed; and it is doubtful if many survived
to return to their eastern resorts. The cause of this singular migra-
tion has never been ascertained.
About the plains between Peking and Tientsin, according to
Swinhoe, flocks of hundreds of this species constantly pass, flying
swiftly like the Golden Plover. The natives call them “ Sha-chee,”’
or Sand-fowl. They are generally caught in nets; and after a fall
of snow a space is cleared, upon which the net is placed and
262 MR. D. G. ELLIOT ON THE PTEROCLIDZ. [Feb. 19,
the place strewed with small green beans. Upon these catching
the eyes of the birds, the flocks descend into the snare and are
trapped. Great numbers could be taken in this way. Although
shy when on the ground, yet on the wing they would dart within
a few yards of aperson. Their cry wasa kind of melodious chuckle.
According to native testimony, they are numerous on the great plains
of Tartary beyond the Great Wall, where they breed in the sand.
In Mongolia, according to Prjevalsky, as given in Rowley’s ‘ Orni-
thological Miscellany,’ this is one of the most characteristic birds,
and is found both in the steppes and deserts. In summer they
go beyond Lake Baikal in the north to breed, but spend the winter
in the Gobi desert and in Ala-shan. They go in enormous flocks,
and feed chiefly upon the seeds of Agriophyllum gobicum. They fly
rapidly, with a peculiar noise of the wings; and at a distance the
sound a flock makes is like the sighing of the wind. When flying
they utter a note like “ truck-turuck, truck-turuck;’ and small
flocks are accustomed to rise in the air and swoop down towards the
ground, as Rooks do when migrating. On the ground they run
clumsily, and their tracks much resemble those of small mammals.
Like the species of Pterocles, these birds visit some drinking-place
every morning and evening, but never settle without first describing
a circle to assure themselves that there is no danger. Prjevalsky also
met with this species in the Hoang-ho valley in South-east Mongolia,
and also about Kalgan. This species deposits its eggs in the sand ;
and the female does not sit very closely, but leaves her charge when
approached, and also every morning and evening exposes the eggs to
the weather, in order to visit the drinking-places. It is a very shy
bird, and takes wing generally before one can get near enough to
oe Falco hendersoni persecutes them, but cannot always catch
them.
Male. Top of head, occiput, back of neck, a line behind the ear-
coverts, and breast grey ; on the forehead an ochraceous tinge ; chin
pale buff. Throat, ear-coverts, and a line on each side of the upper
part of the hind neck dark orange. Across the breast a band of dull
white, each feather margined narrowly with black. Back and wings
vinous, the former barred with black. Outer webs of the greater
coverts chestnut, forming a conspicuous bar along the wing when
closed. Spurious wing-feathers buff, with a central black line. Pri-
maries light ashy grey on outer webs, dark brown on inner ; the fifth
to ninth conspicuously margined on both webs with light buff. The
first primary considerably elongated and attenuated. Abdomen jet-
black. Vent and under tail-coverts pure white. Tail grey, like the
primaries, scalloped on the edges of the feathers with buff and tipped
with white. The median pair elongated, filamentous, their lengthened
portion brownish black. Tarsi and toes thickly covered with short
buffy-white feathers. Bill yellowish brown; iris dark brown ; nails
black.
Total length 144 inches, wing to end of first primary 10, tail to
end of median rectrices 74, bill along gape 4.
Young. Top of head buff, streaked with black. Throat buffy
1878. ] MR. D. G. ELLIOT ON THE PTEROCLID. 263
white, with a narrow black line bounding its lower edge. Upper
part of breast grey ; abdomen and a spot on the centre of flanks
dark chestnut; chest and upper part of flanks, rest of underparts,
and under tail-coverts pure white. Entire upper parts buff; secon-
daries and lesser wing-coverts barred irregularly with black. Greater
wing-coverts chestnut on their outer webs, Spurious wing-feathers
buff, with a central black line. Primaries grey on outer webs, dark
brown on inner, and margined on the innermost ones with buff. Tail
dark grey, tipped with white and barred on the inner webs with buff,
the outer webs having only a slight buffedging. First primary and
median rectrices filamentous, the lengthened portion brownish black.
Tarsi and toe: covered with greyish feathers. Bill brown.
Total length 104 inches, wing to end of lengthened primary 832,
tail to end of median rectrices 53, bill along gape 3.
Ihave not seen this stage of plumage described by any writer.
There are two specimens in the Paris Museum—one a little more
mature than the other, with the bar across the breast commencing
to show as in the adult male.
Female. Crown and neck buff, striated with black. Throat ochra-
ceous yellow, feathers edged with black. The upper parts like those
of the male, but barred more closely with black; and the wing-coverts
are spotted with black. Across the throat is a black band; and the
sides of the neck and breast are spotted with black. Rest of plumage
like that of the male; but the primaries and median rectrices are
notso lengthened. ‘The dimensions are about the same as those
of the male.
2. SyRRHAPTES TIBETANUS.
Syrrhaptes tibetanus, Gould, Proc. Zool. Soc. (1850) p. 92 ; id.
B. Asia, pt. ii, pl. 12; Reich. Syn. Av. Gallin. (1851) pl. cevii. B.
figs. 3020-2] ; Adams, Proc. Zool. Soc. (1858) p. 502, (1859) p.
186; Hend. & Hume, Lahore to Yark. (1873) p. 279; Prjevalsky
in Rowl. Orn. Mise. pt. ix. p. 384 (1877).
Kaling, Tibet and Ladak (Henderson), Stepnaya Kuritza, Tibet-
skaya (Prjevalsky).
Hab. Salt Lakes of Ladakh (Adams); Karakash valley, Yarkand
(Henderson); Northern Tibet, steppes of Kokonor (Prjevalsky).
Mr. Henderson (J. ¢.) says that this species was met with by the
Yarkand Expedition in the Karakash valley towards the end of July,
also at a lake twenty miles south-west of Mallik Shah, and was very
abundant at Chagra, above the Pangong lake, where it flew up and
down the stream in flocks of several hundred individuals, uttering
all the time a ery like “ Yak, yak.” It was very tame; and a great
many were shot and, unfortunately, eaten.
Lieut. Prjevalsky (J. c.) first met with this Sand-Grouse in the
Kokonor steppes in flocks of from ten to thirty, but in Northern
Tibet saw it in flocks of about a hundred individuals. They were
generally on the sandy plains, and fed on the seeds of various grasses
and beans. When so engaged they could be approached quite
close, and seldom flew above a hundred yards before settling again,
264 MESSRS. GODMAN AND SALVIN ON NEW [Feb. 19,
even when shot at. When flying they utter a note like ‘‘caga, caga,”
and on the ground run clumsily and slowly, generally forming a line.
In Northern Tibet they were frequently heard flying about at night ;
but no cause could be found to account for this strange behaviour.
The Kokonor basin, according to Prjevalsky, forms the northern and
probably also the eastern limit of its geographical distribution.
Male. Head and back of neck barred alternately with black and
white, chin and front yellowish white. Sides of neck and throat
orange-yellow. Breast buffy white, barred irregularly with brownish
black. Upper part of back and wings buff, lower part of back and
rump buffy white; all the feathers minutely mottled with black,
hardly perceptible on the wings. Inner webs of scapulars black.
Primaries and secondaries brownish black ; the fourth to eighth
primary brownish white at the tip, mostly on the inner web. Median
rectrices like the back, lengthened and filamentous, their apical half
black ; lateral feathers reddish buff, barred with black, and tipped
with white. Under surface white; under tail-coverts reddish buff,
barred with black and tipped with white. Legs and feet covered
with buffy-white feathers ; bill and nails black.
Total length 153 inches, wing 10, tail 73, bill along gape 4.
4. Descriptions of nineteen new Species of Diurnal Lepi-
doptera from Central America. By F. DuCanz GopMan
and OsBerT SALvIN.
[Received Jan. 18, 1878.]
Nearly half of the species described in this paper have been sent
us during the past year by Mr. H. Rogers from Costa Rica, where
he has been collecting on the slopes of the voleano of Irazu. The
rest of the species are selected from unnamed specimens in our col-
lection, some of them from Guatemala and some from Mexico.
Figures of most of them have been prepared and will be published
shortly. Specimens of all the species are in our collection.
SATYRID.
1. EvurrycHIA PHILODICE.
3g. Exp.1:9in. Margin entire, anal angle of secondaries slightly
produced ; rich brown, central portion of the primaries slightly
paler, middle of the secondaries rich reddish fulvous with a very
distinct tawny-yellow patch, the inner edge slightly sinuate, the
outer extending to two black submarginal spots between the radial
nervules; a third nearly obsolete black spot between the median
branches : beneath reddish brown, a dark band crossing both wings
through the middle of the cell of each, strongly sinuate on the
secondaries ; beyond the cell of the primaries are two subparallel
transverse bands including towards the inner margin a tawny yellow
band; a conspicuous tawny yellow band crosses the secondaries
1878.] DIURNAL LEPIDOPTERA FROM CENTRAL AMERICA. 265
beyond the cell, five argenteous spots surround the margin of the
secondaries, whereof those between the lower radial nervule and the
third and second median branches are the pupils of black ocelli ;
that between the first and second median branches is sagittate.
Q. Exp.1°95. Similar to the male, the colour somewhat paler and
the wings more rounded.
Hab. Costa Rica, Irazu (H. Rogers).
Obs. This well-marked species belongs to the Z.-phocion group of
the genus (cf. Butler, Journ. L. S. Zool. xiii. p. 123), and comes
next H. argentella and #. pyracmon, having the wings similarly
shaped. The peculiar tawny-yellow marks of the secondaries render
the species very distinct from its allies. This marking on the upper
surface is subject to some variation, being more restricted in some
specimens and divided by the reddish brown of the inner portion of
these wings running through it longitudinally. Mr. Rogers has sent
us a good series of this fine species.
2. EvprycHIA ROGERSI.
3S. Exp. 1°9. Form as in &. argentella: uniform brown above,
with a single conspicuous black spot near the margin of the second-
aries, between the third median branch and lower radial nervule:
beneath paler, a dark red-brown sinuate band crosses both wings
through the cell of each; another, nearly straight on the primaries,
sinuate on the secondaries, also crosses both wings beyond the cell,
outside of which is a pale-brown common band margined exteriorly
by a red-brown band hardly traceable on the secondaries; two
argenteous spots near the apical angle of the secondaries, and an
elongated submarginal one between the first and second median
branches; a conspicuous ocellus corresponding to the black spot of
the upper surface, the outer ring of which is fawn-yellow, and in the
black pupil is a concentric silver spot.
Q. Exp. 1:9. Similar to the male but rather paler and the wings
more reunded ; the dark bands of the underside appear as indistinct
reddish bands on the upper surface.
Hab. Costa Rica, Irazu (H. Rogers).
This species also belongs to the Z.-phocion group of the genus.
The presence of a single black submarginal spot on the secondaries,
as well as other characters, distinguishes it from its allies #. argen-
tella, E. pyracmon, and E. philodice.
3. EUpTyYCHIA VETONES.
¢. Exp.1‘8in. Primaries slightly faleate ; outer margin of second-
aries sinuate, most produced at the end of the second median branch :
uniform brown above: beneath brown mottled with paler scales; an
obscure narrow band crosses the cell of both wings ; a broad sub-
marginal pale reddish-brown band, expanding towards the outer
margin, on the primaries; a distinct, yellowish, narrow, nearly
straight band traverses the secondaries from inside the apical to in-
side the anal angle; a distinct ocellus with an argenteous crescent
266 MESSRS. GODMAN AND SALVIN ON NEW [Feb. 19,
inside between the second and third median branches ; argenteous
spots on either side of this ocellus.
Q. Exp. 1:95in. Like the male, but the upperside crossed with
reddish bands corresponding to the markings of the underside ; the
ocellus of the underside shows above as an indistinct black spot.
Hab. Costa Rica, Irazu (H. Rogers).
Obs. A close ally of EH. ithama, Butler (Lep. Ex. p. 9, t. iv. f. 4),
from which it differs in the much clearer definition of the markings
of the underside, the extracellular transverse cross band of the
secondaries being yellowish and not reddish as in H. ithama. The
male of FZ. ithama is not known to us; but, comparing the females of
the two species, the red markings of the upper surface of the southern
race are much more conspicuous than in the northern. Mr. Butler’s
figure of HZ. ithama (the type of the species being before us) is much
too pale in general tint, and the cross bands far too conspicuously
shown.
4. LYMANOPODA EUVOPIS.
6. Exp. 2:2in. Primaries with apical angle rather pointed, outer
margin of secondaries rounded ; rich brownish-black ; black ocelli
with white pupils between the radial nervules and the second and
third median branches, halfway between the cell and the outer mar-
gin; a transverse row of obsolete black spots beyond the cell of the
secondaries: beneath reddish-brown, yellowish beyond the cell; a
bicurved row of five distinct black ocelli with white pupils between the
cell and the outer margin of the primaries, on either side of this row
a sinuate black band, the distal one reddish towards the apical angle ;
at the end of the cell a dusky yellowish spot surrounded by a black
line: secondaries beneath with dark red-brown bands from costal
margin across the cell, and beyond the cell from the third median
branch to the inner margin; a submarginal band made up of out-
wardly concave concentric marks ; across the wing beyond the cell
runs a band of seven small white-pupillated black ocelli, each on a
patch of rich red-brown.
Q . Exp. 2°22in. Brown ; primaries with six black white-pupillated
ocelli arranged in a double curve beyond the cell on a tawny-yellow
ground; secondaries with distal half (except the margin) tawny-
yellow, a band of six black spots crosses the wings beyond the cell:
beneath tawny-yellow, the base of the primaries rather darker, ocelli
and markings of primaries as in the male ; secondaries almost uniform
tawny-yellow, the ocelli and markings almost obsolete.
Hab. Costa Rica, Irazu (H. Rogers).
Obs. A very distinct species of this Andean genus, the first as yet
found north of the Isthmus of Panama.
5. PEDALIODES TRIARIA.
3. Exp. 2°45 in. Allied to P. praaithea (Hew. Ex. Butt. Prono-
phila, pl. v. f. 28, 29), differing in the more pointed primaries ; the
orange band of the same wings is narrower and of nearly equal
width ; in the secondaries the orange spot, instead of being restricted
1878.] DIURNAL LEPIDOPTERA FROM CENTRAL AMERICA. 267
to the apical angle, extends halfway along the outer margin. Be-
neath the difference between the two species is more obvious; the
whole under surface of the secondaries in P. triaria is dark brown
mottled with black, two broken black bands crossing the wings, one
through the cell, the other beyond it; a row of six submarginal
ocelli with white pupils, that between the first and second median
branch being much the largest and most conspicuous ; the mottling of
the apical area of the secondaries is upon a yellowish ground-colour ;
the light ground-colour at this part, and over the distal third of the
secondaries in P. praxithea is wanting.
Hab. Costa Rica, Irazu (H. Rogers).
6. PEDALIODES CREMERA.
Q. Exp.2°9in. Primaries entire, apical angle somewhat produced,
outer margin of secondaries sinuated ; dark brown; primaries crossed
by a broad submarginal band of obscure tawny, the inner edge of
which is concave, and the outer serrate running parallel to the outer
margin: beneath mottled with blackish brown, in the tawny band
of the primaries, between the first and second median branches is a
white-pupillated ocellus; an irregular dark line follows the outer
margin of the secondaries halfway between the outer margin and
the cell; between the first and second median branches near the
margin is a large black ocellus with a white pupil, and two smaller
ones near the anal angle.
Hab. Costa Rica, lrazu (H. Rogers).
7. OXEOSCHISTUS ROGERSI, Sp. 0.
Q. Exp. 3in. Allied to 0. phaselis (Hew.), differing above in
having a conspicuous row of tawny oval spots, separated by the
nervules running parallel to the outer margin, the three spots sepa-
rated by the first and second median branches, and that between the
middle and upper radials bearing large dark brown spots of the same
brown coloui as the rest of the wings: beneath, the markings re-
semble those of O. phaselis, except that the marginal region of the
primaries is strongly marked with fulvous corresponding with the
markings of the upper surface.
Hab. Costa Rica, Irazu (H. Rogers, type mus. nostr.) ; Chiriqui
(Ribbe: mus. O. Staudinger).
Dr. Staudinger’s Chiriqui specimen exactly resembles ours, except
that it is rather paler, being not so fresh.
O. phaselis has been placed in the genus Pronophila by Mr.
Hewitson, and in that of Dedalma by Mr. Butler (Cat. Satyr. p.
184); but, so far as we can see, its proper place is in the genus Oweo-
schistus, with the members of which it agrees in neuration, and with
some of them in style of coloration.
NyYMPHALID.
8. PHyYCIODES THEBAIS, Sp. n.
Exp. 1:5 in. Allied to P. orseis, Edwards, ex California, as to the
position of the marks of the upper surface, and as to the coloration of
268 MESSRS. GODMAN AND SALVIN ON NEW [Feb. 19,
the under surface of the wings: the markings above are pale yellowish
instead of deep fulvous, and the submarginal row of ocelli of the
secondaries is almost obsolete ; the outer margin of the primaries is
more sinuate, being slightly concave about the middle, instead of
following a simple convex outline: beneath the dark marks near the
outer margin and inner angle of the primaries are more clearly
defined.
Hab. Mexico, Mountains of Oaxaca (Fenocchio, type) ; Guate-
mala, Highlands near Duenas, 4800 ft. (S. & G.).
Mus. nostr. Two examples.
9. PHYCIODES BOUCARDI, sp. n.
Exp. 1:3 in. Form of P. elada (Hew.), which it somewhat re-
sembles on the upperside, but from which it differs completely on
the underside. The tawny spots are all of nearly equal size, that
halfway between the end of the cell and the apex of the primaries
(divided into three by the nervules) being slightly the largest. The
spots on the secondaries beyond the cell are arranged in three con-
centric lines, the submarginal one consisting of a series of lunules,
the middle one consisting towards the anal angle of obsolete ocelli.
Beneath nearly uniform tawny, the secondaries being paler; the basal
two thirds of both wings crossed irregularly with fine dark lines ;
the outer third of the primaries submarginally dark, enclosing round
tawny spots; secondaries with a dark patch about the middle of the
outer margin, and a row of black spots corresponding to the obsolete
ocelli of the upperside.
Hab. Mexico, Putla (Rébouch).
Mus. nostr. One example.
We are indebted to Mr. Boucard for the possession of this
species.
10. PuyciopEs sUBOTA, sp. n.
$. Exp. 1°7in. Allied to P. ardys (Hew.), from which it differs
on the upperside in the submarginal lunate lines of the secondaries
being obsolete, and in having the bases of both wings without the
faint marking seen in P. ardys. Beneath, the deep-tawny spots
and pale fulvous wash are replaced by deep brown and pinkish grey ;
a well-defined band of pale spots crosses the middle of the second-
aries.
@. Exp. 1°95 in. Similar tothe ¢, but larger, and all the marks,
especially the band crossing the secondaries, better-defined.
Hab. Guatemala, valley of the Rio Polochic (S. §& G.); Costa
Rica, Irazu (Rogers).
Mus. nostr.
The Costa-Rica specimens are slightly aberrant, the base of the
wings beneath being darker and the cross band of the secondaries
narrower.
1]. PuycioDEs DRYM#A, sp. n.
3.14 in. Allied to P. ardys, Hew., and the last-described
1878.] DIURNAL LEPIDOPTERA FROM CENTRAL AMERICA. 269
species ; from both it differs in the absence of the band crossing
the secondaries, its place being occupied by a row of very small
spots: beneath, the markings of the secondaries are very indistinct,
and consist chiefly of dark marks near the region of the anal angle,
and one projected from beyond the middle of the costa halfway
across the wing.
Hab. Guatemala; hacienda “las Nubes ;” valleys of the rivers
Chisoy and Polochic.
Mus. nostr.
12. Paycroves cynras, sp. n.
3. Exp. 1:45 in. Allied to P. leanira, Feld., ex California, from
which it differs in having a well-defined submarginal row of red spots
on the secondaries above, and also some red spots near the middle of
the outer margin of the primaries, but none on the inner part of the
wing : beneath, the dark markings of P. deanira across the cell and
towards the costa of the secondaries are wanting ; but, as in that
species, the nervules are black.
Hab. Mexico, mountains of Oaxaca (Fenocchio).
Mus. nostr.
13. PuyctopEs NeBuLosa.
3. Exp.1-5in. Belongs to the P.-ardys (Hew.) group. Markings
of wings above deep tawny red; wings more elongated than in P. ardys
and its allies; base of the secondaries with obsolete tawny marks;
secondaries beneath pale greyish, region of the outer margin dark,
and a dark subtriangular mark from the costa halfway across the
wing.
Hab. Guatemala, hacienda “las Nubes ” (0. S.).
Mus. nostr.
Taken in the beginning of February 1874.
14. EResta MECHANITIS, Sp. n.
Q. Exp.2-2in. Like E. eunice (Hiibn.), but differing in having
the apical half of the primaries crossed with two well-defined yellowish
bands: the inner one is but slightly indented and not distinetly
broken as in B. eunice ; the outer one crosses the apex as a distinct
band instead of being an obsolete spot.
Hab. Nicaragua (Janson).
Probably a northern race of Z. eunice, but differing slightly from
all the forms of that species found in the Amazonian region.
15. ErEsta DRYPETIs, sp. n.
3. Exp. 2-2in. Differs from E. eunice and the last-described
species chiefly in the narrowness and direction of the transverse band
of the secondaries. The dark mark over the median nervure of the
primaries is obsolete ; the dark band crossing the primaries is con-
tinuous.
Hab. Panama (MecLeannan, type) ; Guatemala, interior (§. s- G.).
270 ON LEPIDOPTERA FROM CENTRAL AMERICA. [Feb. 19,
16. TIMETES PHIALE, Sp. 0.
g. Exp. 2°5in. Closely allied to 7. corita, Bates, from which
species the male hardly differs : the region of the costa of the secon-
daries is suffused with the reddish tawny colour of the apical angle ;
and beneath the general tint is paler.
Q. Exp. 2°5 in. Differs from 2 7’. corita in general tint of the
wings being brick-red instead of rich brown, the transverse markings
of the wings being much more clearly defined ; beneath the markings
are all very indistinct, both wings being crossed by a common rufous
band.
Hab. Guatemala, Volcano of Atitlan (Salvin).
Mus. nostr. Three examples.
Taken at the end of January 1874, in forest, at an elevation of
about 4000 feet.
17. ADELPHA FALCATA, Sp. n.
3. Exp. 22 in. Primaries elongated, outer margins concave,
dark brown ; both wings with a common white band extending from
the space above the second median branch of the primaries to the
anal angle of the secondaries, where the usual tawny spot is absent.
A tawny spot extends curving outwards from about the middle of
the distal half of the subcostal nervure of the primaries towards the
margin; the outer margin of this spot is deeply sinuated, apex nar-
rowly white. Beneath rich tawny ; the white band as above, but
with a dark margin on either side; the tawny spot is paler and
rounded ; a submarginal row of glaucous spots, obsolete about the
middle of the margin of the primaries; two glaucous spots in the
cell and one beyond it ; two transverse series across the base of the
secondaries, all with dark margins.
Q. Similar to the ¢, but larger (exp. 2°55 in.); the primaries
being less falcate.
Hab. Guatemala, valley of the Rio Polochic.
Mus. nostr.
Obs. In pattern of coloration this species somewhat resembles J.
iphicla (L.); but it may be at once distinguished by the strongly
falcate wings, the absence of the tawny spot at the anal angle of the
secondaries, and the coloration of the under surface.
18. ADELPHA DIOCLES, p. n.
3. Exp. 2:1in. Dark brown, with three darker submarginal
bands to both wings; a common white band crosses both wings from
above the second branch of the median nervure of the primaries to
the anal angle, near which is a tawny spot; a tawny spot runs from
the costa of the primaries, parallel to the margin, nearly as far as the
first median branch, and is separated from the white band by an
oblique dark line ; two dark-margined tawny spots cross the cell, one
at the end and one in the middle. Beneath the bands are as on the
upperside ; the margins of both wings are bluish white, with a mar-
ginal and central tawny line ; this marginal band is separated from
1878. | ON BIRDS FROM THE ELLICE ISLANDS. 271
the central bands by a tawny dark-edged band ; the cell of the pri-
maries is whitish at the base and crossed by a dark-margined spot of
the same colour near the end; beyond it is a similar mark running
into the central white band.
Hab. Voleano of Chiriqui (drcé).
Mus. nostr.
A beautiful and distinct species, coming perhaps nearest A. syma
(Hiibn.) of S. Brazil.
PAPILIONID.
19, PapILio SYEDRA, sp. n.
o. Exp. 5 in. Allied to P. abderus, Hopff., and P. asclepius,
Hiibn., the yellow submarginal lunules of the secondaries being absent
as in the former of these two species. From it the present species
differs in the row of black lunules which traverse the secondaries be-
neath, beyond the yellow band, being distinctly margined on their inner
convex edges with maroon, the outer concave edges enclosing an area
of blue; these marks in P. abderus are respectively grey and yellow:
the submarginal lunules beneath on the secondaries are deeper ma-
roon than in P. abderus.
Hab. Volcano of Chiriqui (Arcé § Ribbe).
Mus. nostr. et O. Staudinger.
We have long possessed a single specimen of this species. Having
recently seen others in the collection of Dr. Staudinger, we have been
able to test the stability of the characters by which it differs from
P. abderus. ‘
5. Ona small Collection of Birds from the Ellice Islands.
By R. Bowprer Suarpe, F.L.S., F.Z.S., &. With a
Note on other Birds found there, by the Rev. S. J.
WHITMEE.
[Received January 21, 1878.]
The Rev. S. J. Whitmee has been so kind as to submit to me a
small parcel of birds obtained for him by a collector in the Ellice
Islands. The few skins do not give materials fora large paper ; but
I trust that the details connected with the synonymy of Anous ceru-
leus will be found interesting.
1. Arpra sacra, Gm.; Finsch & Hartl. Faun. Central-Polyn. p.
201 (1867).
Two specimens in changing plumage, one with the white dress
nearly complete, the other still with plentiful remains of ashy-brown
feathers.
2. ANOUS CHRULEUS.
I must acknowledge the assistance which I have received from Mr.
272 MR. R. B. SHARPE ON BIRDS [Feb. 19,
Howard Saunders’s paper on the Sterninz in the ‘ Proceedings of
this Society for 1876; and it is solely through want of material that
he was unable to discriminate the two Grey Noddies of the Pacific,
as I am able to do from the receipt of the present specimens.
Two examples are in the collection now sent ; and I saw at once
that they belonged to a different species from the bird marked as A.
ceruleus in the national collection. The following is a description
of one of these specimens :—
Supra cinereus, alis cauddque quam dorsum vix saturatioribus :
primariis duobus externis extiis magis nigricantibus : pileo col-
loque undique purius canis, fronte et facie anticd albicantibus :
maculd parvd anteoculari nigrd : corpore reliquo subtis cinereo,
subalaridus eaternis et margine alari canescentibus : rostro nigro.
Long. tota 9°3, rostri a fronte 0°95, alee 6:6, caudze 2°75, rectricis
penultimee 3°85, tarsi 0°9.
It appears that this small species is unquestionably the true Sterna
cerulea of Bennett, and that Mr. Gould was perfectly right when
he separated the North-Australian bird as Anous cinereus, and that
he was wrong in sinking this name, published in 1845, in favour of
Néboux’s, which was not published until 1846. The following
appears to be the true synonymy of the two species.
ANOUS CEHRULEUS.
Sterna caerulea, F. D. Bennett, Narr. of a Whaling Voyage, ii.
App. p. 248 (1840).
Sterng cendré, Néboux, Rev. Zool. 1840, p. 291.
Sterna tereticollis, Lafr. Rev. Zool. 1841, p. 242.
Procelsterna tereticollis, Lafr. Mag. de Zool. 1842, pl. 29.
Anous parvulus, Gould, P. Z. 8. 1845, p. 104; Cassin, U.S.
Expl. Exp. Birds, p. 393 (1858); Gray, Cat. B. Pacifie Isl. p. 60
(1859).
Anous gracilis, Gray, Gen. B. iii. p. 661 (1846, lapsu).
Stolida cinerea, Néboux, Voy. Vénus, Atlas, pl. 9 (1846, nec
Gould).
Megalopterus plumbeus, Peale, U.S. Expl. Exp. p. 285 (1848).
Anous cinereus, Prévost et Des Murs, Voy. Véuus, v. p. 276 (1855);
Finsch & Hartl. Faun. Central-Polyn. p. 239, taf. xiii. fig. 4, taf. iv.
figs. 4 & 5 (1867).
Procelsterna cinerea, Bp. C. R. xiii. p. 773 ee
Anous cinerea, Gray, Hand-l. iii. p. 123. no. 11083 (1871).
Diagn. : Minor, schistaceo-cinereus, pileo et corpore subtus toto vix
pallidius cinereis ; secundariis externis angustissimé ad apicem
albo limbatis ; subalaribus schistaceo-cinereis.
Hab. in insulis orientalibus maris Pacifici.
ANOUS CINEREUS. nboes Kis,
Pelecanopus pelecanoides, Gray, List of Anseres, p. 180 (1844).
Anous cinereus, Gould, P. Z.S. 1845, p. 104; id. B. Austr. fol.
pl. 46 (1848): Reichenb. Vég. Neuholl. p. 181. no. 536 (1850).
ns)
1878.] FROM THE ELLICE ISLANDS. 273
Anous tereticollis, Gray, Gen. B. iii. p. 661 (1846).
Procelsterna albivitta, Bp. C. R. xlii. p. 773 (1856); Gould,
Handb. B. Austr. ii. p- 420 (1865).
Sterna cinerea, Schleg. Mus. P.-B. Sterne, p. 38 (1863).
Anous albivitéa, Gray, Handl. B. iii. p- 123, no. 11089 (1871).
Anous ceruleus, Saunders, P. Z. 8. 1876, p. 671 (nee Bennett).
Diagn. : Major, supra cinereus ; pileo et collo undique et corpore
subtis toto albicanti-cinereis 3 secundariis conspicue albo termi-
natis ; subalaribus albis.
Hab. In Australia septentrionali et orientali usque ad insulas Pa-
cificas ‘ Friendly ’ dictas,
Dr. Finsch, in his paper read before the Society on the 20th of
November last, records this Species from Eua in the F riendly group.
He recognizes the distinctness of the two species; but there is no
need to restore the name of Anous albivitta, as the title cinereus
was given by Gould in 1845, and Néboux’s .A. cinereus was not pub-
lished till 1846.
3. ANOUS sTOLIDUS (L.) ; Saunders, P. Z. S. 1876, p. 669.
Two specimens.
4. Anous LEUCOCAPILLUS, Gould; Saunders, P. Z. S. 1876,
p- 670.
A single example of a Noddy apparently referable to this species.
5. STERNA ANESTHETA (Scop.); Saunders, P. Z. S. 1876,
p- 664.
One specimen.
6. Gyers canpipa (Gm.) ; Saunders, P. Z. S. 1876, p. 667.
One specimen.
Besides the above-named Species there are some eggs, two of a
Noddy Tern, sent with the skins of Anous stolidus and A. leucoca-
pillus and probably belonging to the former, and three eggs appa-
rently of a Duck. In one box were also two wings of a small
Parrot.
Note on the preceding Communication. By the Rev. S. J.
Wuirmer, F.R.G.S., C.M.Z.S.
During 1876 Mr. Fritz Jansen, a Danish Botanical student who
had been residing with me in Samoa for a year, collecting the flora
of those islands, went on a cruise through several groups of islands to
collect for me the flora and as much as possible of the fauna existing
there. In addition to the birds included in the foregoing list, he
saw a Carpophaga in the Ellice Islands ; and the Frigate-bird (Fre-
gaia aquila) also occurs there. In fact, the latter bird is domesti-
cated by the natives ; and when I was in those islands in 1870 I saw
scores of them about the villages sitting on long perches erected for
Proc. Zoou. Soc.—-1878, No. XVIII. 18
274 MR. E.R. ALSTON ON THE DENTITION OF CusCUS. [Feb. 19,
them near the beach. The natives procure the young birds and tie
them by the leg and feed them till they aretame. Afterwards they let
them loose, and they go out to sea to get their food, and return to
their perches in the villages at intervals.
I cannot say to what species the Carpophaga is referable, not
having seen it myself. Mr. Jansen procured young ones in May and
June ; but he, thinking they were the same as the Pigeon found in
Samoa (C. pacifica), did not preserve any specimens. Natives of the
Ellice Islands who were in Samoa when [I left there, told me their
pigeon is like the Samoan species, ‘‘ except that it is smaller, owing
to its food being less plentiful.” That is their own explanation of
the cause of difference. They say it feeds almost entirely on the fruit
of the native fig (Ficus, sp.). They also tell me it is not gregarious
in those small islands as it is in Samoa during part of the year. As
far as they know it does not migrate, but may always be found
sparsely distributed over the islands.
So far as I have been able to learn, no Pigeon occurs in the Gilbert
Tslands ; but a Carpophaga occurs in the Union or Tokelau Islands.
Mr. Jansen tells me that all the birds he procured in the Ellice Is-
lands, which are included in the foregoing list, are found also in the
Union Islands. This may probably be considered nearly a complete
list of the avifauna of those small atolls.
6. Note on the Dentition of Cuscus.
By Epwarp R. Atsron, F.L.S., F.Z.S., &c.
{Received January 25, 1878.]
In reporting last year on the Rev. G. Brown’s collection from
New Ireland and the vicinity’, I observed that the two specimens of
Cuscus contained in it differed from the characters usually given of
C. orientalis in haying but two small teeth behind the large lower
incisors, and in wanting the small extra upper premolar. At the
same time I corroborated Mr. Waterhouse’s remark® that the num-
ber of functionless teeth is not always trustworthy in this group,
and observed that it must be left to future investigation to show
whether the New-Ireland Phalangers were or were not constant in
the peculiarity of their teeth.
Mr. Sclater has now kindly placed in my hands a second small
collection received from Mr. Brown, consisting of one example of
Mus browni, Alst., one of Uromys rufescens, Alst., ten of Belideus
ariel, Gould, and eight of Cuscus orientalis, Pall. All these species
were represented in Mr. Brown’s former consignment ; but the series
of the last-named now available enables me to decide the question as
to its dentition.
In four of these eight specimens the extra upper premolar is
1 P. Z.8. 1877, p. 126. * Nat. Hist. Mamma, i. p. 265.
Hanhart
TSmit lith
W
NEW OPISTHO-BRANCHIATE MOLLUSCA
1878.] MR. CHEESEMAN ON NEW OPISTHOBRANCHIATES. 275
present, being in one case nearly as large as the constant first pre-
molar, while in another it is a mere rudiment ; in the other Sour
examples it is altogether absent. In the number of the functionless
teeth of the lower jaw (which I carelessly called small incisors in
my previous paper, but which Professor Owen regards as the homo-
logues of the canines and anterior premolars") there is the greatest
possible variety, one example having three in one ramus and none
at all in the other; three, however, appears to be the normal num-
ber. These differences have nothing to do with the age of the in-
dividual ; for the extra upper premolar is absent in a half-grown
animal, and the small lower teeth are present in one of evidently
great age.
It appears, therefore, that the dentition of this genus is even more
variable than has been believed, and that the existence of the addi-
tional teeth must be dropped from the specific characters of C.
orientalis. I may add that in several of these specimens, which are
all females, the dark dorsal stripe is absent; this also has been re-
garded as a diagnostic character,
7. Descriptions of three new Species of Opisthobranchiate
Mollusca from New Zealand. By S. T. Currseman,
F.L.S., Curator of the Auckland Museum.
[Received January 29, 1878.]
(Plate XV.)
From a number of new species of Opisthobranchiate Mollusca
collected in or near Auckland Harbour I have selected for descrip-
tion the three following prominent forms,
1. PLEUROBRANCHUS ORNATUS, 0. sp. (Plate XV. figs. 1, 2.)
Body 3-4 inches long, broadly elliptical, depressed, nearly equally
rounded at both ends, colour varying from pale buff to a clear
reddish brown, with irregularly disposed blotches of a rich dark
red-brown ; mantle large, extending over and concealing both head
aud foot, quite smooth, margin thin, entire ; dorsal tentacles short,
stout, abruptly truncate, finely transversely wrinkled, approximate
at their origin, but gradually diverging at their apices ; colour
reddish brown tipped with white ; eye-specks black, placed a little
distance behind the tentacles, embedded in the integument, but
appearing through it; oral tentacles united in front by a thin semi-
circular expansion which forms a veil concealing the mouth, and
which is carried in advance of the foot ; mouth roundish, with
fleshy lips; buccal plates two, regularly reticulated ; odontophore
with numerous rows of similar unciform teeth. Branchial plume
placed in the groove between ‘the foot and the mantle, very large,
* Odontography, i. p. 382.
[3*
276 MR. CHEESEMAN ON NEW OPISTHOBRANCHIATES. [Feb. 19,
composed of about 22-24 pectinations; foot oblong, thin and
flexible, pale waxy white.
Shell internal, 3 to ? inch long, squarish oblong, thin and mem-
branous, semitransparent, slightly iridescent, closely marked with
somewhat irregular concentric strie or folds; colour varying from
nearly white to pale pinkish or tawny brown. Spire minute, ob-
scure, mouth occupying the whole of the undersurface.
My first specimens of this handsome species were obtained from
under stones between tide-marks in Auckland Harbour; where,
however, it is by no means common. Near Waiwera and in some
other localities on the Hauraki Gulf it is much more frequently met
with. It is easily kept alive in an aquarium, but is very sluggish in
its movements.
2. PLEUROBRANCH4&A NOVH-ZEALANDIA, nu. sp. (Plate XV.
fig. 3.)
Body oval, convex, thick and fleshy, smooth and lubricous to the
touch, but the whole surface nevertheless covered with minute
puckers and folds. Colour light grey, copiously streaked with
irregular anastomosing lines of dark greyish-brown, and sprinkled
with numerous minute and almost microscopic white dots. Mantle
smooth, not nearly so long as the foot, and not concealing the
branchize, rather broader on the right side; oral veil broad, extend-
ing over and concealing the mouth, in front semicircular, and with a
delicate fringed margin, but at each side produced into a short
tentacle-like lobe ; mouth large, round, in a state of rest concealed
in the sulcus between the oral veil and the foot, but capable of being
greatly protruded in a proboscidiform manner ; buccal plates two,
large, finely and regularly reticulated or faceted; odontophore
broad, with numerous rows of similar unciform teeth ; tentacles
dorsal, wide apart, short and stout, projecting outwards, folded down
the outer side, tips obliquely truncate; eyes minute, black, placed
within the integument at the inner bases of the tentacles, quite in-
ternal, and not to be seen without dissection ; foot long, extremely
flexible, sole pale ashy grey ; branchial plume often over an inch in
length, and free for half that distance; pectinations about 17,
finely ciliated ; shell none ; length 2°5 to 3°25 inches.
This species is very abundant in Auckland Harbour, usually
affecting sandy or muddy localities. In the winter and spring
months large numbers are often exposed at neap tides, having pro-
bably come into shallow water to deposit their ova. Capt. Hutton,
of the Otago Museum, informs me that he has collected the same
species at Port Nicholson. It is hardy and not easily killed, and
may be kept in confinement for a long time. When in a healthy
state it is by no means inactive, crawling along by means of its
muscular foot much more quickly than might be expected. It has
a curious habit of floating in a reversed position, just under the
surface of the water; and I have also observed it swimming by
means of rather violent vertical undulations of its body.
1878.| THE MARQUIS OF TWEEDDALE ON A NEW HORNBILL. 277
3. ACLESIA GLAUCA, n. sp. (Plate XV. fig. 4.)
Body from 3 to 5 inches long, about ovate when at rest, but
capable of considerable extension, a little contracted behind the
head, then elevated, and suddenly sloping to a point posteriorly ;
entirely covered with numerous simple and branched tentacle-like
processes, the largest of which are sometimes eight lines long.
Colour on the sides pale greyish-brown, passing on the back into a
dull sea-green; the whole surface with numerous irregularly shaped
black blotches that are longest on the back. Along the back there
is also a double row of from 8 to 12 emerald-green specks, each sur-
rounded with a zone of umber. Dorsal tentacles # inch long, folded
down the outer side so as to appear tubular, beset with filiform ap-
pendages. Labial tentacles similar in shape, but rather larger.
Branchial cavity large, protected by the folded-in edges of the
mantle, branchiz quite internal ; foot long and narrow, pointed be-
hind, without side-lobes as in Aplysia, sole pale sea-green ; mouth
roundish, placed under the head; odontophore with very numerous
rows of simple hooked teeth ; gizzard strengthened with large trian-
gular calcareous plates; shell none.
Like many of the species of the allied genus Aplysia, this animal
possesses the power of emitting a purple fluid from the edges of the
mantle, but only in small quantity ; and it may often be handled
without any thing of the kind being observed. All my specimens
are from Auckland Harbour, and were obtained from rather sandy
localities near the extreme verge of low-water mark.
EXPLANATION OF PLATE XV.
Fig. 1. Pleurobranchus ornatus.
2. Shell of the same.
3. Pleurobranchea nove-zealandie.
4, Aclesia glauca.
8. On a new Species of the Genus Buceros. By Arruur,
Marquis of Twerppatz, F.R.S., President of the Society.
[Received February 2, 1878.]
In a collection of birds made at Amparo (in the extreme south of
the Philippine island of Leyte), and sent to me by Mr. Everett,
are some examples of a Hornbill of the genus Buceros, which differ
from the two other known Philippine species’ sufficiently to require
description. The characters which differentiate the large Buceros
of Mindanao, B. mindanensis, from the one which inhabits Luzon
were stated some mouths ago before this Society (see P. Z. S. 1877,
543). But in those two species the form and general contour of
the bill and casque are alike, whereas in this second representative
form of B. hydrocorax the form of the casque is very different.
The colouring of the bill resembles that of B. mindanensis ; and in
© B. hydrocorax and B. mindanensis.
278 THE MARQUIS OF TWEEDDALE ON A NEW HORNBILL. [Feb. 19,
the general colouring of the plumage there is little or no difference.
It is the form of the casque that provides the differential specific
character. Instead of being produced forward as in B. hydrocorar
and B. mindanensis, with an abrupt, compressed, and elevated
anterior margin, the superior plane of the casque loses itself and
dies away on the culmen. The superior plane of the casque is not
flat but arched along its length, the crown of the arch during the
course of its auterior half forming an almost acute ridge. In the
Fig. 1.
Head of Buceros mindanensis, from above.
two known species the contour of the superior flat plane cf the
casque is that of an oval, the posterior end being rounded and the
anterior pointed, In this Leyte species the posterior end is also
rounded ; but the sides, instead of gradually expanding to the greatest
breadth of the superficial area and then graduaily contracting to the
apical point, continue in almost parallel lines for about two thirds of
the length of the major axis of the casque, and then terminate ab-
ruptly, forming corners from which the sides of the remainder of the
casque recede until they reach the culmen.
[S. Leyte, g (adult), September. Iris light yellow; orbital skin
darkest sepia, almost black; gular skin dark Indian-yellow ; feet
coral-red ; nails dark brown-grey.— Everett. |
1878.| THE MARQUIS OF TWEEDDALE ON A NEW HORNBILL. 279
The iris of an adult male from the island of Panaon is recorded by
Mr. Everett as being ‘pale blue.’
Distinct slightly corrugated basal plates are present on the walls
of the mandibles of the adult birds.
The first plumage has little resemblance to that of maturity. The
space between the rami of the mandible and the patch at the base of
the sides of the mandible is dirty brown. The head, neck, breast,
Side view of head of Buceros semigaleatus,
Fig. 4.
Head of Buceros semigaleatws, from above.
and abdomen are dirty greyish fulvous-white, the feathers of the
head, neck, and upper breast being ferruginous at their insertions
and grey at their tips ; and such is the character of the thigh-coverts
and ventral plumage. The upper tail-coverts are also dirty greyish-
fulvous white, with either brown or ferruginous bases. The dorsal
plumage and the quills and other wing-feathers are brown, the quills
dark brown, with broad terminal dirty greyish-fulvous white marks
or edgings. The middle pair of rectrices are for two thirds of their
length pale brown, the apical third being white or dirty creamy
white. All the other rectrices have the basal part to a greater or
less extent pale brown, or pale rusty-brown. In some, probably the
new feathers, the whole rectrix is white. In the plumage described
280 THE MARQUIS OF TWEEDDALE ON [Feb. 19,
the bill is black with a yellow tip, and the casque is not formed.
The colouring of the soft parts of the young bird is noted by Mr.
Everett as follows :—‘*S. Leyte, 2, September. Iris dark brown ;
bill black ; orbital skin greenish yellow; gular skin yellow; legs and
feet dark orange. 6. S. Leyte, 2, October. Iris grey-brown ; bill
jet-black, tip orange; feet dull orange.’
The bill remains black after the bird has assumed the completely
mature plumage and after the casque is almost perfectly formed.
Dimensions. Casque.
a= =F A... = ——— — -—--eeoreneee
Wing. Tail. Tarsus. Bill Length. Breadth. Sides.
from
gape.
in. in. in. in. in. in. in.
fof 15°80 15-00 2-20 5°85 4:90 2-00 3°35
2 14:30 14:50 2:20 5:30 4-75 1°65 3°30
3 juy. 14:50 14:00 2-20 5:20 4:80 1:40 0-00
For this Hornbill I propose the title of Buceros semigaleatus.
9. Contributions to the Ornithology of the Philippimes.—
No. V. On the Collection made by Mr. A. H. Everett in
the Island of Negros. By Artuur, Marquis of Twrrp-
DALE, F.R.S., President of the Society.
[Received January 30, 1878.]
Mr. Everett has been continuing his zoological researches in
the Philippine Islands unremittingly, and, by the consignment of a
collection of birds made during the month of August last in the
neighbourhood of Nueva Valencia and Dumaguete, situated at the
extreme south of Negros, has enabled me to continue these contri-
butions to the ornis of the archipelago.
“The southern extremity of Negros,” Mr. Everett writes,
“‘which is the most mountainous part of the island, and where I
hoped to find accessible virgin forest, is simply one vast field of maize,
sugar-cane, and hemp, perfectly cleared, even far up the steep sides
of the mountains, and is a very poor district indeed for birds.”
Notwithstanding, Mr. Everett in the space of one month secured
56 distinct species, 24 of which are now recorded for the first time
from Negros. Of these 24 there are 6 new to the Philippine area,
3 of which are new to science.
Species not hitherto known as being inhabitants of the Philip-
pines :—
Collocalia francica. Dasycrotapha speciosa,
Butalis latirostris. Zosterops nigrorum.
Limosa egocephala. Macropygia eurycerca.
The last three are new species.
1878. | THE ORNITHOLOGY OF THE PHILIPPINES. 281
The exact habitat of one previously recorded Philippine bird has
been determined by Mr. Everett,
Eudromias geoffroy? ;
and there now remain 38 Philippine birds of which the precise
habitat still continues undecided.
The number of species known to inhabit the island of Negros
previous to the date of Mr. Everett’s visit was about 59'; and to
this number he has added 24, and raised the total of known Negros
species of birds to 83.
The remarkable species Dasycrotapha speciosa, a new type of
Timeliine genus, and the discovery of a second Philippine species of
Collocalia are the most interesting results of Mr. Everett’s ornitho-
logical researches in Negros.
1. CacatuaA HZMATUROPYGIA (1).
[Valencia, ¢, August. ]
2. TANYGNATHUS LUZONENSIS (3).
[Valencia, 3, 2, August.]
Of two examples (¢) shot in August, one has the crown and
occiput blue, the otber green; in both the uropygium is green.
3. LorRIcULUS REGULUS (6).
Loriculus regulus, Souancé; Rowley’s Ornith. Mise. pt. vii.
p- 238, pl.
(Dumaguete, ¢, 9, August: bill orange-red; cere orange ;
feet lighter orange; claws black. @Q (juy.), August: feet dirty
ochre-yellow. |
The series consists of eight examples, four marked as belonging to
males, and four to females.
The males are in the adult typical plumage of the species. Two
of the females have the occiput faintly golden, and traces of the
orange nuchal band. The two other females have the whole head
green, and the crimsen upper tail-coverts covering only half the
length of the rectrices. No blue about the chin, cheeks, and throat.
The male, as figured by Keulemans (/. c.), only exhibits a red
band across the throat, whereas the lower throat and breast possess
a large orange-red plastron.
4. Limnaétus PHILIPPENSIS (14).
[Valencia, 2, August: iris pale brownish grey ; bill black ; lores
and cere dull greenish ; feet light dirty yellow; claws black. ]
Pure white underneath, the thigh-coverts being very faintly barred
1 Dr. Steere added 21 species to the 38 recorded by me (Tr. Z. 8. ix. p. 252)
as known from Negros. Dr. Steere collected partly in North Negros, and
partly at Dumaguete in the South; but, from the general term “ Negros” being
frequently all that is given as the locality by Mr. Sharpe (Tr. L. 8. 2nd Series,
Zool. vol. i.), it is impossible in many cases to determine the exact part of the
island in which Dr. Steere’s specimens were obtained.
282 THE MARQUIS OF TWEEDDALE ON [Feb. 19,
with irregular lines of pale fulvous. Head and neck pure white, a
few feathers with brown terminal drops. Back brown, each feather
natrowly margined with pale tawny rufous. Uropygium and upper
tail-coverts pale earthy brown, some of the coverts tipped with pure
white. Minor wing-coverts brown, edged with dirty white. Major
coverts pale earthy brown, with paler margins and white terminal
edges. Quills dark brown, more or less blotched with paler brown
and white. Rectrices above rich brown, with five or six very narrow
cross dark brown irregular bands, a broader dark brown subterminal
band. Shafts pure white for three fourths of their length. Ree-
trices underneath albescent, the brown bands being more conspicuous
than above.
Wing 15°20 inches, tail 12°00, tarsus 3°50, culmen 1°50. Nuchal
crest-plumes dark brown, tipped with white; length 3°30.
5. HALIASTUR INTERMEDIUs (17).
[Valencia, 9, August. }
In full plumage.
6. Merops pHiLipPinus (35).
[Valencia, ¢, 9, August. Dumaguete, 2, August. ]
The series (7) consists cf examples ranging from first plumage to
that of the almost adult. None are in full plumage.
7. MERops BICOLOR (36).
[Valencia, ¢, 9, August. ]
In a series of five examples, one is in perfect plumage, the
remaining four represent different stages of immaturity.
8. EURYSTOMUS ORIENTALIS (37).
[Dumaguete, 2, August. ]
9. EnromMostiA GULARIS (44).
[ Valencia, ¢, August. Dumaguete, 2, August.]
10. SauROPATIS CHLORIS (47).
(Valencia, 2, August. Dumaguete, 2, August. |
11. XANTHOLEMA ROSEA (51).
[ Valencia, ¢, August. }
12. CoLLOCALIA FRANCICA.
Hirundo francica, Gm. S. N.i. p. 1017; Walden, Ibis, 1874,
p- 132. no. 70.
[Valencia, d, 9, August. ]
Mr. Everett’s researches enable me to make known the existence
in the Philippines of a second species of Collocalia. He has sent
five examples. Underneath they are mouse-grey ; above fuliginous
brown, faintly tinged with bottle-green. The wings and tail darker
brown than the back, showing a greenish gloss on the new feathers.
1878.] THE ORNITHOLOGY OF THE PHILIPPINES. 283
I can find no character whereby to separate these Negros individuals
from those inhabiting Malacca, the Andamans, Sikim, Ceylon, the
Neilgherries, the Mauritius, and the Fiji Islands. Wing 4:50.
Formerly (7. ¢.) I referred by implication C. troglodytes to the C.-
JSrancica section of the genus ; but a reexamination of a considerable
series of C. troglodytes shows that it more properly belongs to the
section of which true C. esculenta may be taken as being the type.
13. Cenrrococcyx vrrivis (64).
. [Valencia, ¢, August. Dumaguete, ¢, August. ]
14. PENELOPIDES PANINI (68) ?
[Valencia, 9, August: iris dull chestnut-brown ; orbital and
other bare skin white faintly tinged bluish ; bill brown with olive-
green tinge; feet dark lead-grey with a faint greenish cast ; nails
black. |
The single example sent wears the dress of the mature female of
P. panini; but the bill is smooth, without lateral plates, grooves,
or casque. The upper tail-coverts are ferruginous and not. black.
It may belong to a distinct representative form.
15, ARTAMUS LEUCORYNUS (73).
[Dumaguete, ¢, 2, August. }
16. GraucaLus striatus (74).
[Dumaguete, g, August. Iris crimson. ‘alencia, 9, August.
Iris crimson ; bill black ; legs grey-black. |
Of four examples three (3) have the throat and upper breast uni-
form slate-colour, the remainder of the lower plumage, the uropygium
and upper tail-coverts being broadly banded with black and white.
One example (marked 2 ) differs by having the entire lower plumage
banded with black and white.
17. Votvocivora? c#RULESCENS (75)?
[Valencia, ¢, August: iris dark chocolate; bill black ; legs
blackish grey. ]
A single example of a bird (marked ¢ ) which I provisionally
refer to the above species is sent from Negros by Mr. Everett. it
differs from every example in my series ex Luzon and Zebu in
having a longer and larger bill, in being dark ashy grey above, with-
out any dark brown almost black margins to the feathers, the uro-
pygium and upper tail-coverts being of still paler grey, in being
below pale grey and not dark iron-grey, in having the major wing-
coverts pure white and the tertiary quills broadly margined with pure
white, and in having the two outer pairs of rectrices broadly tipped
with white and some of the other rectrices slightly white- tipped. The
under tail-coverts are also almost pure white. The general dimensions
are about equal in all. The adult males of V. cerulescens are jet black;
the adult females are dark plumbeous grey (conf. P. Z. §, 1877, p.
759. no. 23), while this bird is pale-coloured. V. cerulescens is as
284 THE MARQUIS OF TWEEDDALE ON [Feb. 19,
yet only known to inhabit Luzon and Zebu; and this may possibly
belong to a representative form. But as it may also be V. c@rules-
cens in first plumage, I refrain from bestowing on it a distinet title.
18. LALAGE pomrnica (76).
{Valencia and Dumaguete, ¢, 2, August. ]
19. Dicrurus MIRABILIs (81).
[Valencia, 5, 9, August. ]
20. PHILENTOMA ALBIVENTRIS.
Philentoma albiventris, Sharpe, Tr. L.S. 2nd series, Zool. i.
p- 325. no. 49, “ Guimaras” (Nov. 16, 1876).
[Valencia, ¢, August. ]
A single example is sent by Mr. Everett ; and it seems to belong
to a species distinct from P. eyaniceps, and Mr. Sharpe appears to
have been justified in separating it, notwithstanding the doubt he ex-
presses (J. c.). Besides the lower breast and abdomen being pure
white, the dimensions of the rectrices and wings, when compared with
those of an adult Luzon male, are longer ; tail 4:12 as against 3°50) ;
wing 3°25 as against 2°87. These, however, are larger dimensions
than those given by Mr. Sharpe of his type.
21. LeucocERCA NIGRITORQUIS (83).
[Valencia, ¢, August. Dumaguete, 9, August. ]
22. CyORNIS PHILIPPINENSIS.
Cyornis philippinensis, Sharpe, Tr. L. S. 2nd series, Zool. i. p.
325.
Valencia, ¢ (juv.), August: iris dark brown; bill black ; legs
bluish grey. Dumaguete, August. |
The young birds (of which a series of five are sent) have their
plumage, with the exception of the remiges and rectrices, which are
brown, washed with blue or greenish blue, marked with large pale
rufous spots. In one the whole of the body-plumage is so marked ;
in another the abdomen and throat are white ; in another several of
the uniform blue dorsal feathers have come in, and the breast is
turning to pure rufous. The throat-plumes are the last to change
to the adult colouring, while full maturity is indicated by the inten-
sity of the rufous on the throat and breast.
23. HyporHyMIs AzuREA (85).
{ Valencia, ¢ (not adult), August: iris dark brown ; bill black ;
legs dark grey. Dumaguete, ¢, August. ]
24. BuUTALIS LATIROSTRIS.
Muscicapa latirostris, Raffles, Tr. L. S. xiii. p. 312; Walden,
Ibis, 1873, p. 308.
[Valencia, 2, August. |
A single example in first plumage, with pale rufous spots on the
1878. | THE ORNITHOLOGY OF THE PHILIPPINES. 285
sides of the neck and margins to the secondary and tertiary quills
and major coverts ; wing 2°56.
25. BropERIPUS ACRORHYNCHUS (90).
[Valencia, 9 (juv.), August: bill dark sepia-brown. Dumaguete,
3 (juv.), August: bill dark vandyke-brown. |
Judging by the series Mr. Everett sends (seven) and other examples
of the Negros Oriole I have examined, it would appear that the ex-
tent of the yellow on the forehead is more restricted in the Negros
and Guimaras forms than in examples from any of the other Philip-
pine islands.
26. MEGALURUS RUFICEPS.
Megalurus ruficeps, Tweeddale, Ann. & Mag. Nat. Hist. ser. 4,
vol. xx.; P.Z. 8. 1877, p. 94.
[Valencia, 2, August: iris light clay-brown. ]
Males from the same locality are of equal dimensions. New to
Negros.
27. DasyCROTAPHA SPECIOSA.
Dasycrotapha speciosa, Tweeddale, P. Z. 8. 1878, pl. ix. p. 114.
[Valencia, ¢, August: iris crimson ; bill orange-yellow. ]
28. Ixus GoravieR (99).
[Valencia and Dumaguete, ¢ and 9, August. |
A series of five, with ear-coverts and sides of head brown.
29. HysplPETES PHILIPPINENSIS (102),
[Valencia and Dumaguete, 9, August. |
30. CopsyCHUS MINDANENSIS (105).
[Valencia, ¢ and 2, August. Dumaguete, 9, August. |
31. Cisticoxa, sp. ?
[Valencia, ¢, August: iris light grey-brown; bill brown; legs
pale tinged with brown. |
Mr. Everett’s note, above quoted, refers to a single example of
the genus Cisticola which I am unable to determine. The example
is in perfect plumage and belongs evidently to an adult; but the
great variations in plumage the members of the genus exhibit make
it possible that it belongs to some described species; and I refrain
from bestowing what may be a useless title. Its dimensions are
smaller than those of any species known to me. Wing 1°62, culmen
0°31, tarsus 0°62. The first primary is relatively long and broad,
0°53. Underneath the general colouring yellow-white, with a rufous
tinge on the breast; flanks, under tail-coverts, wing-lining, and
thigh-coverts pale ferruginous. Above the colouring is pale ferru-
ginous-brown, each feather dark centred with brown; the nape is
uniform pale ferruginous-brown, the upper tail-coverts being
brighter ferruginous; the quills are brown margined with the
286 THE MARQUIS OF TWEEDDALE ON [Feb. 19,.
same pale ferruginous as the upper plumage. Rectrices above pale
brown, underneath much paler, with bold subterminal dark brown
almost black spots, which show through on the upper surface
as brown marks; the inner margins of the quills are pale ferru-
ginous.
This Negros bird differs from N.-Mindanao, Celebes, and Bangkok
(Siam) examples of C. grayi in its smaller dimensions and longer
and broader first primary, and in the absence of the pure uniform
rufo-fulvous cap.
The description of C. semirufa, Cab., is too meagre for identifi-
cation, and no dimensions are stated; but it is probably the same
bird as C. grayi. O. ruficeps, Gould (Sclater, P. Z. 8. 1877, p. 98),
belongs to C. grayi, the only difference Dr. Brown’s example
exhibits being the absence of the rufous tinting of the breast and
under surface generally.
32. ORTHOTOMUS CASTANEICEPS (113).
[Valencia, ¢ and 9, August. ]
New to Negros and not separable from the type, which inhabits
the island of Guimaras.
33. CorypALLA LuGuBRis (117).
[Dumaguete, ¢ and 2, August. |
34. Parus ELEGANS (118).
[Valencia, ¢ and 9, August. ]
New to Negros.
35. ZosTEROPS NIGRORUM, Ni. Sp.
[Valencia, ¢ and 9, August. ]
Male and female. Above light olive-green ; whole under plumage
light greenish-yellow, almost pure yellow; closed wing hike the
back, outer quill-margins paler ; space before the eye almost pure
yellow; axillaries and under wing-coverts very pale yellow ; a narrow
black mark margins the white feathers of the underside of the
orbit ; rectrices pale brown edged with olive-green. Wing 2°18,
tail 1°87, tarsus 0°62, culmen 0°44.
Closely allied to Z. austent, Walden, ex Karen-nee, this Negros
species differs in being of a darker shade of green above and a
lighter yellow underneath.
36. Dic&uM H&ZMATOSTICTUM.
Diceum hematostictum (Sharpe, ‘ Nature,’ August 1876, p. 297;
Tr. L. S. ser. 2, Zool. i. p. 339).
[Valencia, 3, August. |
37. NECTAROPHILA SPERATA (122).
[Valencia, ¢ and 2, August. ]
38. ARACHNECTATHRA JUGULARIS (123),
Valencia, ¢ and 9, August. ]
1878. | THE ORNITHOLOGY OF THE PHILIPPINES. 287
39. ASTHOPYGA MAGNIFICA.
Athopyga pacifica (Sharpe, ‘ Nature,’ August 1876, p. 297;
Tr. L. S. ser. 2. Zool. i. p. 342; Shelley, Monogr. Cinnyride, pt.
3, pl.
[Valencia, ¢ and 2, August. ]
In one example (¢) the uropygium is deep orange and not
yellow.
40. ANTHOTHREPTUS CHLOROGASTER.
Anthreptes chlorogaster (Sharpe, Tr. L. S. ser. 2, Zool. i. p. 342.
no. 107, ‘‘ Negros,” (1876).
[Valencia, 2, August: iris indian-red.]
41. Corvus pHrLippinus (125).
[Dumaguete, ¢, 2, August. ]
The disproportion between the dimensions of sexes in the Philip-
pine Crow exhibits itself in the Negros examples sent by Mr. Everett,
the wing of the male measuring 12 inches, as against 11 in the
female ; culmen of male 2°37, of female 2.12.
42. CALORNIS PANAYENSIS (128).
(Dumaguete, ¢, 2, August. |
43. Sarcops caLvus (129).
[Dumaguete, 5, 9, August. ]
44. Munia sacort (1382).
[Valencia, ¢, 2, August. Dumaguete, ¢, 2, “ breeding,”
August. |
May not Meyen’s Fringilla minuta (Walden 133) be M. jagori
in first plumage, before the black feathers comein? Otherwise it is
remarkable that a species stated by Meyen to occur in numberless
troops in the Luzon sugar-plantations has not, since he wrote (1834),
been recognized.
45. OSMOTRERON AXILLARIS (136).
[ Valencia, ¢, August. Dumaguete, 2, August. ]
46. LeucorrEeRon GironiERt (137)?
[ Valencia, ¢, August. }
A single example of a Pigeon much resembling L. gironiert, ex
Luzon and Guimaras, and probably representing an immature stage
of its dress, is sent by Mr. Everett. It differs from all the phases
of plumage described by me (/. c.) in having the throat, breast, and
abdomen pale ashy white, much stained pale verdigris green. The
crown and nape rich green like the back. A purple pectoral band
is indicated by a few isolated plumes. The first primary is not
_abruptly attenuated ; but perhaps the attenuated first quill is only
produced later. The species has not heretofore been recorded from
Negros.
288 ON THE ORNITHOLOGY OF THE PHILIPPINES. [Feb. 19,
47, CaARPOPHAGA &NEA (141).
[Dumaguete, dg, 9, August.
48. IANTHG@NAS GRISEIGULARIS (145).
[Valencia, 3, August: iris orange ; orbital region and basal half
of bill dark crimson ; apical half of bill pale yellow ; feet dull car-
mine ; nails yellowish grey. |
49. MacropyGIA EURYCERCA, sp. n. (146, partim).
[Valencia, ¢, August: iris yellow, with outer ring crimson; or-
bital region and base of bill crimson; bill light brown; feet car-
mine. |
Mr. Everett sends a single example, which agrees in every respect
with the example obtained in Negros by Dr. B. Meyer, and described
by me (Tr. Z. S. ix. p. 218. no. 146) when writing on M. tenui-
rostris. A recomparison made with Luzon examples leaves no doubt
that the Negros bird is specifically distinct.
50. TurTuR pussuMIERI (147).
[Valencia, ¢, August. Dumaguete, ¢, August. ]
51. CHALCOPHAPs INDICA (150).
[Valencia, ¢, August. |
52. Eupromias GEoFrrroy! (161).
[Dumaguete, 2, August: bill black; legs very pale greenish
grey; feet darker grey. |
53. RALLINA EURYZONOIDES.
Gallinule euryzonoides, Lafresn. Rev. Zool. 1845, p. 368; Tweed-
dale, P. Z.S. 1877; p. 767.
[ Valencia, ¢, August. |
New to Negros. The Luzon (Manilla) bird (Tr. Z.8. ix. p. 231.
no. 175) has to be compared ; for it is doubtful whether it does not
rather belong to this species than to R. fasciata.
54. Hypora2nipia TorQuAtTA (177).
[ Valencia, 9? , August. ]
New to Negros.
55. Limosa #GOCEPHALA.
Scolopax egocephala, Linn. S. N. i. p. 246. no. 16.
[Valencia, August. ]
56. NycTicoRAX MANILLENSIS (198).
(Dumaguete, ¢, August: iris golden yellow; bill black, the base,
greater part of lower half, and bare orbital skin bright yellow-green ;
legs and feet light yellow-green, clouded with olive-green in front,
and on the upper surface of toes. |
In first plumage.
1878.] ON BIRDS FROM DUKF-OF YORK ISLAND, ETC. 289
March 5, 1878.
Prof. Newton, F.R.S., V.P., in the Chair.
The Secretary read the following report on the additions to the
Society’s Menagerie during the month of February 1878 :—
The total number of registered additions to the Society’s Mena-
gerie during the month of February was 63, of which 27 were ac-
quired by presentation, 17 by purchase, 4 by exchange, 6 were bred
in the Gardens, and 9 were received on deposit. The total number
of departures during the same period, by death and removals, was 93.
The most noticeable additions during the month of February
were
A pair of Leopards (Felis leopardus), obtained from Capt. Phillips,
of the S.S. ‘Mesopotamia,’ February 18th. These Leopards, which
were brought from the Persian Gulf by Capt. Phillips on his last
voyage, are remarkable for their long hairy coats, bushy tails, and
pale body-colour, which reminds one rather of the Ounce (Felis
uncia). They are probably from some part of the mountainous
district of Persia.
Mr. Sclater exhibited a second collection of birds from Duke-of-
York Island, New Britain, and New Ireland, which he had received
from the Rev. G. Brown, C.M.Z.S. These were mostly duplicates
of the collection brought before the Society on the 19th of February
last year’, and consisted of examples of the following species :—
1. Rhipidura setosa. 12. Centropus ateralbus.
2. Monarcha alecto. 13. Helectus polychlorus.
3. Dicranostreptus megarhyn- 14. Nasiterna pusio.
chus. 15. Lorius hypeenochrous.
4. Nectarinia aspasia. 16. Carpophaga rubricera.
5. Srenata. Ly. van-wycki.
6. Philemon cockerelli. 18. Gdirhinus insolitus.
7. Calornis nitida. 19. Chalcophaps stephani.
8. Gracula kreffti. 20. Calenas nicobarica.
9. Corvus orru. 21. Megapodius eremita.
10. Dendrochelidon mystacea. 22. Tringoides hypoleucos.
11. Halcyon sanctus. 23. Tringa acuminata.
Mr. Sclater called special attention to the following specimens, and
read the subjoined notes on them ;—
6. PHILEMON COCKERELLI (/.c. p. 104).
Two additional examples of this Philemon, both from New Britain,
are in the collection. The species is exactly of the same form as P.
plumigenis (Tropidorhynchus plumigenys, G. R. Gray, P.Z.S.
1 See P. Z. 8. 1877, p. 96; also Mr. Ramsay’s notes on the same collection,
Proc. Linn. Soc. New 8. Wales, vol. i. p. 369.
Proc. Zoou. Soc.—1878, No. XIX. 19
290 MR. P. L. SCLATER ON ATHENE VARIEGATA. [Mar. 5,
1858, p. 174) of the Ké Islands, but is a larger and stronger bird,
darker on the head, and paler on the throat and belly.
10. DENDROCHELIDON MystTacea (J. c. p. 105).
Five skins from Duke-of-York Island.
12. CENTROPUS ATERALBUS (/. c. p. 106).
Three examples of this Coucal from New Britain. In one the
head is black, and back of the neck partly so; in the second the
neck is white; and in the third the head is also white,—showing
that, as I suggested (J. s. c.), there is much variation in colour in
this species.
13. EcLecrus poLYcHLoRUsS (I. c. p. 106).
Of twenty skins of this species, mostly labelled ‘‘ Duke-of- York
Island,” but without indication of sex, fifteen are males (green) and
five females (red).
18. CEDIRHINUS INSOLITUs (J. ¢. p. 110).
Of this singular Fruit-pigeon, until lately supposed to be so ex-
cessively rare, there are no less than nineteen skins from Duke-of-
York Island. The sexes are not marked; but there is no variation
in their plumage; so we may consider them identical in this
respect. Some of the specimens have the frontal knob much more
developed, and are probably males.
In Part viii. of his ‘Ornithological Miscellany’ (pl. lxvi.) Mr.
Rowley has given us an excellent figure of this species.
21. Mrearopius EREMITA, Hartlaub, P. Z.S. 1867, p. 830.
Count T. Salvadori informs me that he has satisfied himself that
Megapodius hueskeri of Cabanis and Reichenow, as I had called this
bird, /. s. c., is identical with the species previously described by
Hartlaub under the name above given. He further assures me that
my Megapodius rubrifrons from the Admiralty Islands (P. Z.S.
1877, p. 556) is undistinguishable.
23. TRINGA ACUMINATA, Horsf.
One skin of this species, which was not represented in the first
series, from New Ireland. ‘
Mr Sclater took this opportunity of exhibiting the specimen re-
corded by him (P.Z.S. 1869, p. 123) as Athene variegata, and
subsequently described and figured by Mr. Sharpe (P. Z. 8. 1876,
p- 673, pl. lxii.) as Ninow solomonis. Mr. Sclater had compared
this specimen with the type of Noctua variegata of Quoy and
Gaimard, and had ascertained that his identification was correct, and
that there was therefore no need of Mr. Sharpe’s new name for the
specimen.
1878.] PROF. NEWTON ON PEZOPHAPS SOLITARIA. 291
Mr. Sclater exhibited a stuffed Coot belonging to the Museum of
Science and Art at Edinburgh and believed to be the typical speci-
men of Fulica gallinuloides of King (Zool. Journ. iv. p. 96), and
showed that it was really an example of Fulica leucoptera, Vieillot,
and not of Fulica armiilata, as he and Mr. Salvin (P. Z. 8. 1868,
p- 465 et Ex. Orn. p. 115), misled by Capt. King’s imperfect de-
scription, had supposed.
Professor Newton, M.A., F.R.S., V.P.Z.S., exhibited a stone sent
him by Mr. Caldwell, C.M.Z.S., and remarked :—
«The veracity of the earliest writer who treated at any length of
the Solitaire (Pezophaps solitaria) affords a pleasing contrast with
that of the latest. Many of the statements put forth by Leguat
concerning that bird have already been curiously confirmed; and I
have much pleasure in laying before the Society what appears to be
another proof of his accuracy. Writing of the hen Solitaires he
says :—
«On leur trouve toujours dans le gésier (aussi bien qu’aux males)
une pierre brune de la grosseur d’un Oeuf de poule; elle est un peu
raboteuse, platte d’un coté & arrondie de l’autre, fort pesante, & fort
dure. Nous avons jugé que cette pierre nait avec eux; parce que
quelque jeunes qu’ils soient, ils en ont toujours, & n’en ont jamais
qu’une; & qu’outre cela, le canal qui va du jabot au gésier, est trop
étroit de moitié pour donner passage 4 une pareille masse. Nous
nous en servions préférablement 4 aucune autre pierre, pour aiguiser
“nos couteaux’?.
‘When Mr. H. H. Slater was appointed by the Royal Society
Naturalist to the Transit-of-Venus Expedition in Rodriguez, I
especially drew his attention to this statement; but I have under-
stood from him that, notwithstanding his careful examination of
the caves of that island, he never found any thing bearing out
Leguat’s assertion. Shortly after his return, our Corresponding
Member, Mr. Caldwell, of Mauritius, visited Rodriguez, as the Society
is already aware (Proc. Zool. Soc. 1875, pp. 644-647). He was,
as he has told us, more fortunate, and obtained three or four of what
he believes to be the stones mentioned by Leguat. One of these he
has been so good as to give to my brother, Mr. Edward Newton;
and on behalf of both those gentlemen I now exhibit it. You will see
that in most respects it agrees closely with the description of Leguat.
It is brown, somewhat rough, heavy and hard. It is hardly,
however, flattened on one side ; and in connexion with that fact I may
remark that the bird with whose remains it was associated appears to
have been young. Its mineralogical nature has yet to be determined ;
but before it is cut up for that purpose I thought the Members of the
Society would like to see it. Its weight is a little over 14 oz.
**Mr. Caldwell has also been so kind as to furnish me with some
remarks on this and the other specimens he obtained ; but his state-
l Voyages et Avantures de Francois Leguat &c. Londres: MDCCVIII,
vol. i p. 100,
ise
292 MR. PARKER ON THE COMMON ROCK-LOBSTER. [ Mar. 5,
ments respecting them will possibly be more advantageously published
as an appendix to the account of this bird which, as most (though
apparently not every one) of our Members are aware, was drawn up
more than twelve months ago, at the request of the Royal Society, by
Mr. J. W. Clark and Mr. Edward Newton.”
Mr. T. J. Parker exhibited and made remarks upon the stridu-
lating organ of the Common Rock-Lobster (Palinurus vulgaris).
He stated that the apparatus in question was produced by a peculiar
modification of the antennulary sternum and of the basicerite or
second joint of the antenna. The antennulary sternum forms a pro-
jecting vertical keel, with a rounded anterior border, and slightly
convex sides; the anterior border is grooved in the middle line, the
groove being bounded on each side by a strong smooth ridge; each
lateral surface is marked with a shallow groove; and between this
groove and the border, to which it is parallel, the surface is so smooth
as to have the texture of polished ivory. The basicerite is uncalcified
along a slit-like space on its inner surface; this space is filled up
with a thick chitinous membrane, which is produced superiorly into
a large flap, projecting outwards from the surface of the joint.
Immediately beneath this flap the chitin becomes thickened, takes
on the form of an oval area or pad, about } inch long by 7 inch
wide, and marked by a number of fine parallel ridges. In relation
with the inferior edge of this pad is a small calcified tubercle.
When the antenna is in place, the flap works over the ridge on*
the corresponding side of the anterior border of the antennulary
sternum, the tubercle fits into the groove on its lateral surface, and
the ridged pad is closely applied to the smooth space between the
groove and the border. Under these circumstances, when the
antenna is moved upwards the friction of the pad against the smooth
surface produces a loud grating noise, the principle being similar to
that of the sound produced by the friction of india-rubber against
paper. The apparatus can at any time be thrown out of gear, and
the antenna moved noiselessly, by slightly abducting the latter. In
November last, Mr. Saville Kent remarked in ‘ Nature’ upon the
“shrill squeaking sound”’ emitted by living specimens of Palinurus
when handled, this sound being due, according to Mr. Kent, to the
friction of the abdominal somites; and Mr. Parker suggested that
the noise referred to may possibly have been produced by the
apparatus described. He stated further that the apparatus was
possessed by both sexes, although in the two specimens examined
the noise produced was far louder in the male ; and he also remarked
upon the fact that, notwithstanding the great perfection of the
stridulating organ, the auditory organ of Palinurus is in a very
imperfect condition.
The following papers were read :—
WPurkass hth Hanhart ump
NEW SPECIES OF PLUSIOTIS
1878.] MR. A. BOUCARD ON THE GENUS PLUSIOTIS. 293
1. Notes on some Coleoptera of the Genus Plusiotis, with
Descriptions of three new Species from Mexico and
Central America. By A. Bovcarp, C.M.Z.S.
[Received F ebruary 4, 1878.]
(Plate XVI.)
I have again the pleasure to exhibit to the Fellows of this Society
a fine series of Beetles of the genus Plusiotis, which I have acquired
since I wrote my former paper on these Insects in the ‘Proceedings’ !.
Amongst them are four new species totally distinct from all others.
Three of these have been liberally given to me by my friends MM.
Sallé, Baden, and Rodriguez. The fourth I discovered during my
last voyage in Costa Rica. This increases the number of species
known of these fine insects to twenty, of twelve of which the types
are in my collection.
During my recent Journey in Costa Rica, 1 have also been able
to procure one more specimen of Plusiotis batesii, which agrees com-
pletely with the type, and one specimen of Plusiotis chrysargyrea,
Sallé, varying from the original types of the species, not being quite
of the same colour.
These insects are excessively rare, and appear to confine themselves
to the forests, at a very high altitude.
I was in Costa Rica in the proper season, and at the exact locality
where these insects are found; but I was not able to get more than
three specimens, although I offered a high price tor them to the
natives, and did myself all that possibly could be done; I made
several excursions on purpose, but with no result.
Every ove in the country knew what I meant when I asked for
golden and silvery beetles; but they did not procure me any.
PLusIorIs BATESIL.
Plusiotis batesii, Boucard, P, Z. S. 1875, p. 119, pl. xxiii.
ITab. Volcano of Irazu, Costa Rica.
One specimen of this fine Species, found in May near Potrero
Cerrado, at the altitude of 8000 feet, was alive, but had lost its abdo-
men and some of its legs. It had evidently just been caught by a bird
and dropped, or had disengaged itself, after having lost its abdomen.
The vitality of this insect is so great, that several hours after I had
found my specimen it could still move part of its body and its remain-
ing legs. Although I looked with the greatest care for this insect, I
was uot able to find any more Specimens,
At the suggestion of my friend Sallé, who thought that they ought
to be found abundantly on Mimosa trees, like Pelidnota, I searched
for them during many days on these plants, but without success,
and those which I secured ‘were found in the oak-tree region.’ I saw
another specimen, which I consider to be a very fine variety of this
" See P. Z. 8. 1875, p. 117.
294 MR. A. BOUCARD ON THE GENUS PLusiIoTIs. _[ Mar. 5,
species, in a small collection of insects made by Mr. Rogers. It is
of the most splendid brass-colour ; it was found near Rancho redondo,
on the Volcano of Irazu, by an Englishman who was in the company,
Mr. Rogers.
This insect is now in the collection of Mr. Du Cane Godman.
PLUSIOTIS CHRYSARGYREA.
Pelidnota chrysargyrea, Sallé, Annales de la Soc. Ent. de France,
tome iv. p. 362.
One specimen of this magnificent species, found in April in the
Candelaria Mountains, at the altitude of 5000 feet, in the oak-tree
region.
It is quite perfect, and was taken alive on a roble tree (Tecoma
pentaphylla, Jacq.). Being somewhat different from the types of the
species, I give a description of it :—
Oblongo-ovata ; capite thoraceque aureo-pallidis, marge rubro ;
elytris aureis, tenuiter punctatis, punctis in stris regulariter dis-
positis ; infra aurea, tibiis tursisque cupreis, antennis fuscis.
Length 1 inch 1} line. Mus. Boucard.
Head, thorax, and elytra very slightly punctured, of a pale burnished
gold-colour with coppery-red reflections ; the anterior margin of the
head and the sides of the thorax of a coppery-red colour, which dis-
tinguishes this species from P. bates? ; tibize and tarsi of the same
colour, with blue claws ; abdomen and pygidium of a brilliant golden
colour; antenne rufous.
Its place is near P. batesi, to which it is closely allied.
Puiusiotis Lara, Sturm, Cat. 1843, p. 341.
According to M. Auguste Sallé, of Paris, who possesses the type of
P. leta, Sturm, this species must be abolished, as it was described
from a bad specimen of Plusiotis psittacina, which is from Mexico.
This proves that I was right when I suggested in my former paper
that the locality of Valparaiso for this species was probably wrong ;
as up to this date no species of Plusiotis has been found south of
Columbia.
PLUSIOTIS BADENI, n. sp. (Plate XVI. fig. 2.)
Oblongo-parallela, valde convexa, supra viridi-argentea, cupite
cupreo-violaceo, medio viridi crebre punctulato ; thorace subtiliter
punctulato, margine cupreo-violaceo, scutello violaceo levi; elytris
tenuiter punctatis, punctis in strus profundis regulariter dispo-
sitis; infra viridi-argentea, femoribus viridibus, tibiis cwpreo-
violaceis, tarsis cupreo-aureis, antennis fuscis.
Length 7+ lines.
Head of a coppery purple colour with the centre green, thorax
silvery green, with a broad margin of the same colour as the head,
both slightly punctured ; the seutellum is also of that same colour ;
elytra silvery green, slightly darker than the head and thorax,
strongly punctured, the punctuation disposed in regular striz ; the
1878. ] MR. A. BOUCARD ON THE GENUS PLUSIOTIS. 295
underside is of a paler green with silvery reflections ; tibize coppery
purple, and tarsi coppery metallic; antennze rufous.
Hab. Mexico.
The female of this species is of the same size, but a little wider ;
the tarsi are less coppery ; and the colour of the thorax, elytra, and
pygidinm is slightly paler.
This species must be placed between P. gloriosa, and P. lacordairet.
I have pleasure in naming this fine insect after my friend Doctor
Baden, to whom I am indebted for the specimen now in my collection.
Types, museums Baden and Boucard.
PLUSIOTIS RODRIGUEZI, n. sp. (Plate XVI. fig. 1.)
Oblongo-convewa, supra viridi-aurea ; capite thoraceque subtiliter
punctulatis ; elytris punctatis, punctis in striis regulariter dis-
positis, margine lato aureo splendido; infra pallide viridis,
tibtis tarsisque aureis, antennis rufis.
Length 1 inch.
Head and thorax slightly punctured, the latter twice as long as it
is broad, convex, with areddish gold margin, elytra strongly punctured ;
the punctuation is disposed in regular strie, convex, broader in the
middle and rounded in the base with a magnificent gold margin all
round ; the underside is of a pale uniform green with silvery reflec-
tions ; tibiz and tarsi golden; antennz rufous.
Hab. Guatemala.
I have only one specimen of this charming species, the only one
which was collected in the neighbourhood of Guatemala City by my
intimate friend Mr. Juan Rosé Rodriguez, to whom I have the plea-
sure of dedicating this fine insect. It was found by him in the forests
of oak trees near the capital. Its place is between P. costata and
P. sallei, Bouc.
Type, mus. Boucara.
PLUSIOTIS PRASINA, 0. sp. (Plate XVI. fig. 5.)
Oblongo-ovata, supra viridi-prasina ; capite fortiter punctulato,
thorace subtiliter punctulato ; elytris fortiter punctatis, punctis
in striis irregularitus ; infra viridis, tibiis tarsisque violaceo
micantibus.
Length 1 inch 1 line.
Head and elytra of a dark metallic green, both strongly punctured ;
on the latter the punctuation is disposed in irregular strie ; they are
oblong, a little broader in the middle, with a prominent, thick, and
compressed margin on the sides, which is very conspicuous ; thorax
slightly punctured, of the same colour ; the underside is metallic green;
tibize metallic purple ; tarsi of the same colour, with coppery reflec-
tions ; claws nearly black ; antennee dark rufous.
Hab. Mexico.
Its place is between P. auripes and P. sallet.
I have only one specimen of this insect, which was given to me by
my intimate friend M. Auguste Sallé.
Types, museums Boucard and Sallé.
296 MR. A. G. BUTLER ON LEPIDOPTERA [ Mar. 5,
Puusioris BoucARDI, Sallé. (Plate XVI. fig. 3.)
Oblongo-parallela, valde convexa ; capite thoraceque viridi-aureis,
subtiliter punctatis, margine cupreo; elytris splendide viridi-
aureis, fortiter punctatis ; infra cupreo-rufa, tibiis tarsisque
nigris, antennis fuscis.
Length 1 inch 2 lines.
Head, thorax, pygidium, and elytra of a magnificent metallic golden
green, with red reflections, surrounded by a narrow coppery-red mar-
gin. The elytra are rugose and strongly punctured ; the underside
is of a coppery red; femora, tibize, and tarsi quite black in the ante-
rior legs; in the middle and posterior pairs the femora are coppery
red and the tibiee and tarsi are black ; antennze nearly black.
Hab. Costa Rica.
I have only one specimen of this magnificent species, found in the
mountains of Candelaria, in the forest of robles (oak trees) at the
altitude of 5000 feet.
Its place is near P. victorina.
Type, museum Boucard.
I have seen all the species of this genus described in this and my
previous paper, and am satisfied that they are good and distinct. I
have examples of all of them in my collection, except P. psittacina
and P. gloriosa.
EXPLANATION OF PLATE XVI.
Fig. 1. Plusiotis rodriguezi, p. 295.
2. P. badeni, p. 294.
. P. boucardi, p. 296.
. P. mnizechii, P. Z. 8. 1875, p. 124.
. P. prasina, p. 299.
Ore Co b
2. On a small Collection of Lepidoptera obtained by the Rey.
J.S. Whitmee at the Ellice Islands. By Arruur G.
Butter, F.L.S., F.Z.8., &e.
[Received Feb. 6, 1878.]
The little series of Lepidoptera obtained from the Ellice Islands by
Mr. Whitmee, although it contains only five species of Butterflies
and two of Moths, makes a valuable addition to our knowledge of the
geographical distribution of the forms inhabiting the Australian
region, since we have hitherto been quite ignorant of the insect-fauna
of these islands.
Of the seven species, all of which are described, one is of special
interest to me; I refer to Huploea distincta, a form figured by
Herrich-Schiiffer, from an example taken in the Viti Islands, as a
variety of £. eleutho. To this insect I gavea name in my “ List of
the Diurnal Lepidoptera of the South-Sea Islands” (P. Z. 8. 1874,
pp. 274-291), and was severely censured for so doing by Herr
1878. ] FROM THE ELLICE ISLANDS. 297
Schmeltz, in a paper criticising my determinations of South- Pacific
Butterflies. Herr Schmeltz considered 2. distincta to be an indi-
vidual variety, and regarded it as an injury to science to give ita
name ; it is therefore satisfactory to find that Z. distincta is the
common Euplea of the Ellice Islands, and consequently may be re-
garded as a fixed local form, of equal importance with L. helcita, E.
angasii, and others of the #.-eleutho group.
RHOPALOCERA.
NYMPHALID.
Evrxiaa, Fabricius.
1. EUPL@A ELEUTHO.
Danais eleutho, Quoy & Gaimard, in Freycinet’s Voy. pl. 83.
fig. 2 (1815).
The fact that this species is so rare that Mr. Whitmee only brought
home a single example seems to show that its time of appearance is
not synchronous with that of EL. distincta. It represents the oppo-
site extreme of modification in the group to which it belongs, and
therefore is not likely to be a variety, since forms more nearly ap-
proaching #. distineta in character are known to be locally con-
stant.
2. EUPL@A DISTINCTA.
Euplea distincta, Butler, P. Z.S. 1874, p. 278.
A good series, in both sexes, was obtained by Mr. Whitmee.
Junonia, Hibner.
3. JUNONIA VILLIDA.
Papilio villida, Fabricius, Mant. Ins. p. 35. no. 366 (1787).
Resembles Australian examples, being less suffused with orange
tawny than Samoan specimens.
Diapema, Boisduval.
4. DIADEMA NERINA.
Papilio nerina, Fabricius, Syst. Ent. p. 509. no. 277 (1775).
The whole of the specimens are rather small, resembling those of
Cape York.
5. DIADEMA OTAHEIT.
Diadema auge, Cramer, var. otaheite, Felder, Verh. zool.-bot.
Ges. Wien, xii. p. 492. no. 185 (1862).
This may possibly be a dwarfed race of the preceding species ; it
is the common form of Samoa, where the specimens run eyen smaller
than in the Ellice Islands.
298 MR. E. J. MIERS ON THE PENZIDE. [Mar. 5,
HETEROCERA.
Lirgosiip&.
Deroreta, Stephens.
6. DEropEIA PULCHELLA.
Tinea pulchella, Linneeus, Syst. Nat. |, 1. p. 884 (1766).
Larger than Australian specimens, and with the red dashes on the
primaries broader and longer.
OpHIvusIDa.
Acu#a, Hubner.
7. ACHEA MELICERTE.
Phalena-Noctua melicerte, Drury, Ill. Ex. Ent. i. p. 46, pl. 23.
fig. 1.(1770).
One worn example was obtained of this widely distributed moth.
3. Notes on the Peneide in the Collection of the British
Museum, with Descriptions of some new Species. By
Epwarp J. Miers, F.L.S., F.Z.S., Zoological Depart-
ment, British Museum.
[ Received February 12, 1878.]
(Plate XVII.)
Mr. G. E. Dobson, M.B., F.L.S., lately presented to the Trustees
of the British Museum a remarkable specimen of Peneus, with the
request that I should lay a description of it before the Society. In
studying the affinities of this form, which is described below under
the name of P. dobsoni, I found it necessary to revise the characters
of the whole of the species of this large and difficult genus ; and al-
though the materials existing in the national collection are not rich
enough at present to permit of a complete review of the subject, it is
hoped that the following notes on the species in the collection will
contribute toward the elucidation of the genus, while the synoptical
key to the genus appended will serve to indicate what appear to be
the principal distinctive characters of the different species, and to
mark what are desiderata in the collection of the British Museum.
I have added some remarks upon the Funchalia woodwardi, and on
the Peneus edwardsianus of Mr. J. Y. Johnson, showing that this
latter species must be referred to the genus Aristeus of Duvernoy ;
also a description of a species of Sicyonia not hitherto recorded.
PENZUS CANALICULATUS.
Palemon canaliculatus, Olivier, Encycl. Méth. Hist. Nat. vii.
p. 660 (1811).
Penaeus canaliculatus, M.-Edw. Hist. Nat. Crust. ii. p. 414
(1837); De Haan, Faun. Japon. Crust. p. 190 (1849).
P 2.9.18 76, Pl 2V0
C.Berjeau del et lith.
NEW SPECIES OF PENAZIDA.
Mintern Bros imp.
1878. ] MR. E. J. MIERS ON THE PEN®ID. 299
Penaeus marginatus, Randall, Journ. Ac. Nat. Sci. Phil. viii.
p- 146 (1839).
? Peneus plebejus, Hess. Arch. f. Nat. p- 168, pl. vii. fig. 19
(1865).
The series in the British-Museum collection includes specimens
from the Gulf of Suez, Shanghai, Yokohama (Japan), Torres Straits,
Shark Bay, Houtmann’s Abrolhos, Sir-C.-Hardy Island, and the
Loyalty Islands (Lifu). In both sexes there is a single spine on the
second joint of the first and second pairs of legs, and none on the
third pair of legs. In the closely allied European P. caramote there
is, besides these spines, a spine upon the first joint of each of the
first three pairs of legs. The sixth postabdominal segment is not
sulcate in either species; and in both the terminal segment is armed
with lateral marginal spines.
PENAUS BRASILIENSIS.
Peneus brasiliensis, Latr. Nouv. Dict. Hist. Nat. xxv. p. 256
(1817); M.-Edw. Hist. Nat. Crust. ii. p. 414 (1837) ; Stimpson,
Ann. Lye. Nat. Hist. New York, x, p- 132 (1871).
This species in external characters closely resembles the prece-
ding, but differs in the following particulars :—There is a spine upon
the second and third joints of the first pair of legs, one upon the
second joint of the second pair of legs, and none upon the third pair
of legs, The sixth postabdominal segment is grooved on either side
of the longitudinal median carina on its upper surface. There are
no spines on the lateral margins of the seventh postabdominal
segment.
Specimens from Whydah on the west coast of Africa agree in all
respects with authentic specimens from Brazil presented by the
Paris Museum, and with specimens from the West Indies (Barba-
does) and North America, proving that this, like many other species.
occurs on both sides of the Atlantic.
PEN ZUS SEMISULCATUS.
Peneus semisulcatus, De Haan, Faun. Japon. Crust. p. 191,
pl. xlvi. fig. 1 (1849).
A large series of specimens of both sexes is in the Museum collec-
tion, agreeing with De Haan’s description and figure. It is probable
that under the Fabrician name of P. monodon two closely allied
but apparently distinct species were confounded by earlier authors.
In P. semisulcatus the rostrum is 7-8-toothed above and 3-toothed
below, the dorsal carina, which is faintly or obsoletely canaliculated,
extends to the posterior margin of the carapace, and the gastro-
hepatic sulcus is faintly defined. There are examples from India
(Pondicherry, Calcutta), Formosa, the Philippines, North Australia,
and the Fiji Islands (Mbau), in the Museum collection.
In two specimens from Australia (Shark Bay), and one from Cey-
lon, the rostrum is 6~7-toothed above, the teeth toward the apex
separated by much wider intervals than in the specimens referred to
300 MR. E. J. MIERS ON THE PENEZIDA. [ Mar. 5,
P. semisulcatus, there is a short dorsal carina which does not reach
to the posterior margin of the cephalothorax and is not canaliculated
above, and the gastro-hepatic sulcus is very deep and strongly de-
fined. For this latter form I propose to retain Fabricius’s name of
P. monodon, if the two species be distinct; as it would seem that
the common Indian form must be designated by De Haan’s name of
P. semisulcatus, it having been first distinctly characterized by him,
while it is impossible to ascertain from Fabricius’s brief description
which of the two species was known to that author.
The figure of P. tahitensis of Heller (Reise der Novara, Crust.
p- 121, pl. xi. fig. 2, 1865) resembles this species ; and in it the ros-
trum is represented as 3-toothed below; but it is described as eden-
tate; so I cannot refer it with certainty to P. semisulcatus.
The P. carinatus of Dana(U.S. Expl. Exp. xiii., Crust. i. p. 602,
pl. xl. fig. 2, 1852), from Singapore, is only briefly described, but
appears to be identical with P. semisulcatus, with which it agrees in
the form and number of the teeth on the rostrum. It is not, how-
ever, mentioned in the description whether the longitudinal carina
on the cephalothorax is sulcate or not.
PENZUS HARDWICKI, sp. n. (Plate XVII. fig. 1.)
The cephalothorax is very minutely granulated; the cervical
suture is in its posterior half, and the cardiaco-branchial altogether,
obsolete ; a canaliculated dorsal crest extends from the base of the
rostrum to the posterior margin. The rostrum reaches beyond the
peduncles of the antennules, is 8—9-dentate above, the last tooth
separated by twice the ordinary distance from the preceding; the
inferior margin is entire; it is curved regularly upward toward the
distal extremity, which is acute. There is a spine on the underside
of the second joint of the first and second pairs of legs, and none on
the third pair. The form of the sternum is as in the P. curviros-
tris of Stimpson ; and the third to sixth postabdominal segments are
carinate as in that species. The last postabdominal segment is
deeply longitudinally sulcate above, and, as in P. semisulcatus, acute
at apex, without lateral spines.
Hab. Indian Seas? (Hardwicke).
There are two specimens in the British-Museum collection, of
which the exact locality is not known. This species is most nearly
allied to P. curvirostris, Stimpson, from Simoda, Japan, but differs
in having a canaliculated dorsal crest, and a longer rostrum, which
is acute at the extremity ; the sutures of the cephalothorax are also
much less distinct. From the P. semisulcatus of De Haan, which
resembles this species in having a single longitudinal dorsal sulcus,
it differs in the form of the rostrum, which is edentate below, and
in having only a single spine at the base of the first pair of legs,—
and from the P. sculptilis of Heller, from Java, another unisul-
cate species, in the longer, more curved rostrum, and in the post-
abdomen being keeled only on the third to sixth segments, whereas
in P. sculptilis all the postabdominal segments are said to be
keeled.
1878.] MR. E. J. MIERS ON THE PENEIDE. 301
PEN XUS INDICUS.
Peneus indicus, M.-Edw. Hist. Nat. Crust. ii. p. 415 (1837).
Peneus stylirostris, Stimpson, Ann. Lyc. N. H. N. York x. p. 134
(1871).
In specimens in the British-Museum collection from the Indian
seas, Ceylon, and Chefoo, which I refer to this species, the first pair
of legs are bispinose at base, the second pair unispinose, and the
third pair unarmed ; there are no lateral marginal spines on the last
postabdominal segment. The rostrum in young specimens consider-
ably projects beyond the antennal scale, and the median dorsal carina
reaches nearly to the posterior margin of the cephalothorax ; in adult
specimens, however, the rostrum scarcely projects beyond the anten-
nal scale, and the dorsal carina is nearly obliterated. In one speci-
men, fully adult, from Amoy, which is probably only a variety of
this species, the rostrum is even shorter than the antennal scale, and
4-dentate.
I cannot find any valid characters by which to distinguish from
this species the P. stylirostris, Stimpson, of Panama, of which speci-
mens, from that locality, are in the British- Museum collection.
PENZUS MONOCEROS.
Peneus monoceros, Fabr. Ent. Syst. Suppl. p. 409 (1798) ; M.-
Edw. Hist. Nat. Crust. ii. p. 415 (1837); De Haan, Faun. Japon.
Crust. p. 192, pl. xlvi. fig. 2, P. ensis (1849); Dana, U.S. Expl.
Exp., Crust. i. p. 605, pl. xl. fig. 5 (1852).
Specimens in the British-Museum collection from the Indian seas,
Calcutta, Ceylon, and Penang, agree with M.-Edwards’s short de-
scription, and possess the diagnostic characters indicated by De
Haan, but differ from De Haan’s longer description in having no
lateral marginal spines on the terminal segment. De Haan’s speci-
mens belong perhaps to a distinct species, in which case the name
of P. ensis, as they are designated in the plate, would be applicable
to them.
PENZUS MEMBRANACEUS.
Peneus membranaceus, Risso, Crust. des environs de Nice, p. 98
(1816) ; Hist. Nat. Eur. Mérid. v. p. 68 (1826); M.-Edw. (part.),
Hist. Nat. Crust. ii. p. 417 (1834); Heller, Sitz. Akad. Wissensch.
xlv. i. p. 423, pl. 1. fig. 49 (1862) ; Crust. siidl. Europa, p. 296,
pl. x. fig. 11 (1863).
Penaeus longirostris, Lucas, Explor. Sci. Algérie Crust. p. 46,
pl. iv. fig. 6 (1849).
Peneus bocagei, Johnson, Proc. Zool. Soc. p. 255 (1863), p. 900
(1867).
See Johnson’s remarks (P. Z.S. 1867, p. 900) on the complicated
synonymy of this species. If Dr. Heller’s views are correct, Milne-
Edwards attributes to P. membranaceus some of the characters of
the Solenocera siphonocera (Peneus siphonocerus, Philippi), subse-
quently described by Lucas, Ann. Soe. Ent. France, vill, p. 219
302 MR. E. J. MIERS ON THE PEN 4ZIDS. [ Mar. 5,
(1850), under the name of Solenocera philippii. Besides P. stpho-
nocerus the Peneus distinctus of De Haan (Faun. Japon. Crust.
p- 194, 1849) evidently belongs to this genus, and must be desig-
nated Solenocera distincta. Of it I have seen no specimens.
Perhaps also the Pencus crassicornis of Milne-Edwards (Hist.
Nat. Crust. ii. p. 418, 1837) from India should be referred here.
PreN&US DOBSON, sp. n. (Plate XVII. fig. 2.)
Cephalothorax smooth, not hairy or scabrous, with the cervical,
gastro-frontal, and cardiaco-branchial sutures faintly indicated.
Antennal and hepatic spines present, but no pterygostomian spine at
the antero-inferior angle of the cephalothorax. Rostrum slender,
not quite reaching to the apex of the antennal scale, 7—9-dentate
above, about four of the teeth being posterior to the frontal margin,
the posterior tooth separated by a greater interval than the others,
which are equidistant from one another; there are no teeth on the
anterior third of the upper margin, nor on the inferior margin; the
apex, in the only specimen which appears perfect, is acute. Poste-
riorly the rostrum is prolonged into a short median dorsal crest,
which terminates at some distance from the posterior margin. There
are no longitudinal median dorsal su/c?. The first five postabdomi-
nal segments are rounded at their postero-lateral angles ; the poste-
rior margins of the first, second, fourth, and fifth have a semicir-
cular notch on each side of the body ; there is a longitudinal median
dorsal carina on the third_to sixth segments, which is but faintly
indicated on the third segment, and a faintly indicated lateral carina
on each side of the fifth and sixth segments; the terminal segment
is longitudinally sulcate above, the extremity very slender and acu-
minate, the lateral margins ciliated, but without mobile spines; the
appendages of the sixth segment somewhat oblong-oval, and nar-
rowed at the distal extremities.
The eyes are very large; the antennules with the joints of the pe-
duncle flattened on their inferior surfaces, and with two flagella, of
which the inner is nearly twice as long as the outer, but not as long
as the cephalothorax. Antennee with the flagella very long. The
crowns of the mandibles are slightly concave on their inner surfaces,
the cutting-edges thin, and with a tooth near the apex; the palpus
tomentose and 2-articulate, the terminal joint foliate, triangular and
acute. The second maxillipedes are densely hairy ; the outer maxil-
lipedes are very slender, almost styliform, and their exognathi reach
almost to the extremity of the penultimate joimt. The three ante-
rior pairs of legs are small and very slender, successively slightly in-
creasing in length, but the third pair not more robust than the pre-
ceding ; they are very feebly didactyle, and the fingers are much
shorter than the palmar portion of the hand; the fourth pair of legs
are very feeble, and much shorter than the preceding ; the fifth pair
are obsolete, thickened, and two-jointed, the terminal joint consti-
tuting an indurated corneous lobe; there is also a small lobate pro-
longation at base of each of the fourth legs; and the lobe between
the bases of fifth pair of legs (which resembles in form that usual
1878.] MR. E. J. MIERS ON THE PENEZID. 303
in the female of Peneus, but is not divided by a median fissure) is
of the same corneous consistency. All the legs, except the fifth
rudimentary pair, are furnished with an exopodite. There is a spine
on the under surface of the second joint of the legs of the first three
pairs, that on the third pair of legs very small.
Hab. Mangalur (Mangalore), west coast of India.
Collected by Dr. G. E. Dobson, to whom I have much pleasure
in dedicating the species.
I have examined four specimens of this species, all of which are
certainly females. Oue of these has been presented by Dr. Dobson
to the trustees of the British Museum. The fact that no males
have yet been observed renders it possible that the rudimentary and
indurated condition of the fifth pair of legs may be peculiar to the
female sex—an opinion shared both by Dr. Semper and Mr. Wood-
Mason, who have seen the specimens ; and as the mouth-appendages,
although presenting some peculiarities, do not depart widely in struc-
ture from the type usual in Peneus, I think it advisable until more
specimens shall have been examined, to retain the species in that
genus. Should further researches, however, prove that the rudi-
mentary condition of the fifth legs exists in both sexes, the name
Mangulura may be adopted to designate the genus, which will be
characterized not only by the above-mentioned character, but also
by the triangular shape of the terminal joint of the mandibular pal-
pus (which is less delicate and transparent in texture than is usual
in Peneus), and the slender outer maxillipedes. The species will then
stand as Mangalura dobsoni’.
The form of the fifth pair of legs is not analogous with that cha-
racteristic of the very different genus Sergestes, wherein these
members, although rudimentary, are very slender, and similar in
structure to the preceding pairs.
In the form of the rostrum P. dobsoni resembles the Peneus
sculptilis of Heller, from Java (Voy. Novara, Crust. p. 122, pl. xi.
fig. 1); but, besides the characters derived from the fourth and fifth
pairs of legs, which are of the normal form in P. seulptilis, that
species differs, if the figure be correct, in the much longer flagella of
the autennules, and in the absence of a notch in the posterior mar-
gin on the sides of the first and second postabdominal segments, in
the form of the first postabdominal segment, which is rounded at its
1 Since writing the above, Professor Wood-Mason has discovered, among speci-
mens of P. dobsoni from the same locality (Mangalore), a single male individual,
in which he observed that the fifth legs are fully developed, and which, by his
kindness, I am now enabled to figure (see Pl. XVII. fig. 2, f). It is of very small
size, not much more than half as large as the females described by me, which
it in all respects resembles, except in the fifth legs. These are very slender and
elongated, longer, in fact, than those of the preceding pair, with the dactylus
much shorter than the preceding joint. The fact, therefore, thatthe rudimen-
tary condition of the fifth legs is a sexual character peculiar to the female may
be regarded as definitely established ; nor does it seem that this can possibly
be accidental, or due to the loss of the legs and subs2quent induration of the
joint, as the indurated terminal lobe is of the same shape on both legs in all
four of the specimens seen by me, and I have never obseryed a similar condition
of the joint in any other specimen of the genus.
304 MR. E. J. MIERS ON THE PENZIDS. [ Mar. 5,
antero-inferior angle, &c. The sex of the specimens of P. sculptilis
is not stated.
In the P. avirostris, Dana (U.S. Expl. Exp. xiii. Crust. i. p. 603,
pl. xi. fig. 3, 1852), from Singapore, which is but shortly charac-
terized, the rostrum is somewhat of the same form, but straighter,
shorter, and only 6-toothed above, the legs of the two posterior pairs
much more slender than the preceding (sex not stated). The
cephalothorax is not dorsally carinate in its posterior half.
PEN ZXUS AFFINIS.
Peneus affinis, M.-Edw. Hist. Nat. Crust. ii. p. 416 (1837);
De Haan, Faun. Japon. Crust. p. 192, pl. xlvi. fig. 3, P. barbatus
(1849).
Penaeus velutinus, Dana, U.S. Expl. Exp. xiii. Crust. i. p. 604,
pl. xl. fig. 4 (1852).
Whether this species be the P. afinis of Milne-Edwards must remain
uncertain, as his type specimen has not been figured, and his short
description leaves many particulars unnoticed ; but it is certainly the
one figured by De Haan under the name of P. barbatus, and referred
by him in the text to P. affinis. P. velutinus of Dana (U.S. Expl.
Exp. xiii. Crust. i. p. 604, pl. xl. fig. 4, 1852) from Lahaina, Sand-
wich Islands, seems to be also referable to the same species. His
description applies to it ; and although his figure, when compared with
that of De Haan, presents some slight differences, they are probably
due to inaccuracies of drawing. Examples of this species are in the
British-Museum collection from Japan, Sandy Cape, Sharks’ Bay,
West Australia, and the Gulf of Suez; hence it would appear to
have a wide geographical range.
P. constrictus, Stimpson (Ann. Lye. Nat. Hist. New York, x.
nos. 4, 5, p. 135, 1871), from the coast of Carolina, U.S., of which
there is one specimen in the British-Museum collection, presented
by the Smithsonian Institution, is nearly allied to the above. The
teeth of the rostrum are stated to be equidistant by Stimpson; but in
the specimen I have examined, the last tooth is (as in P. affinis)
separated by a wider interval than the preceding. It differs, how-
ever, in being much less pubescent (the postabdomen is naked and
glossy), and in having a very deep longitudinal median sulcus on the
last segment, which is margined by lateral carinee.
P. pubescens of the same author (/. c. p. 133), based upon a single
female example with imperfect rostrum from St. Thomas, would also
seem from the description to be closely allied to P. affinis, and, in
the absence of specimens for comparison, I am unable to say how it
may be separated from that species. It will probably prove to be
distinct, on account of the widely distant locality ; therefore I do not
quote it as a synonym of the Indo-Pacific form. ‘
PEN ZUS STYLIFERUS.
Peneus styliferus, M.-Edw. Hist. Nat. Crust. ii. p. 418 (1837).
There is a single specimen, a male, in bad condition, which I refer
1878. ] MR. E. J. MIERS ON THE PEN XID. 305
with some hesitation to this species, in the Museum collection. The
rostrum is of the form described by Milne-Edwards, but reaches but
little beyond the peduncle of the antennules, and scarcely at all be-
yond the antennal scale. The fifth pair of legs is considerably
longer than the preceding, and very slender; the dactylus small,
of the usual form. This species cannot be the male sex of P.
dobsoni, described above, as it differs in the form of the postabdo-
minal segment, which is rounded at its antero-lateral angle, in the
existence of spines on the lateral margins of the last postabdominal
segment, &c. The median dorsal carina is prolonged to the poste-
rior margin of the cephalothorax ; and there isa raised lateral line on
each side, extending from a point immediately above the antennal
spine halfway to the posterior margin.
If the genus Xiphopeneus of Smith (Trans. Conn. Ac. ii. p. 26,
1871) be sustained, it will probably be necessary to refer this
species to it, as it agrees in the form of the rostrum, and in the length
of the legs of the fifth pair, although the dactylus is quite short.
PENZXUS KROYERI.
Peneus kroyeri, Heller, Wien. Akad. Wissensch. Sitzungsb. xlv.
(Abth. 1.), p. 425, pl. ii. fig. 51 (1862).
Aiphopeneus harttii, Smith, Trans. Conn. Ac. ii. p. 28, pl. i.
fig. 1 (1871).
I have little hesitation in referring the Xiphopeneus harttii of
Smith, from Caravelhas, Bahia, to the previously described P. kréyeri
of Heller, from Rio Janeiro. A comparison of the figures and de-
scriptions will, I think, suffice to establish their identity: and there
is a specimen from the West Indiesin the British-Museum collection
which appears to belong to the same species. The genus Xipho-
peneus was characterized by its author mainly by the long and slender
antennules (which have a small lamelliform appendage which is not
foliaceous and expanded over the eye as in Peneus), the form of the
cephalothoracic sulci, aud the great length of the fourth and fifth
thoracic legs (which have the terminal joints slender and flagelliform
or, rather, styliform). The mouth-organs, antenne, &e. are not
different from those of Peneus.
It will be seen that these characters are not of greater value than
those which distinguish the Peni of Milne-Edwards’s first and second
sections (which have never been considered of generic importance).
Moreover the length of the posterior legs is certainly a character
that varies in the different species ; and in one (P. styliferus) already
referred to, which agrees in most of its characters with Xiphopeneus,
the dactylus of the fifth pair of legs is quite short. Nor do we kaow
at present how far these peculiarities are dependent upon the sex of
the animals (the specimens of P. Ardyert and P. styliferus in the
Museum collection are both males).
If the genus Xiphopeneus be retained as distinct, it will be neces-
sary to constitute more than one other new generic division for species
hitherto included in Peneus; and as the materials at present existing
Proc. Zoou. Soc.—1878, No. XX. 20
306 MR. E. J. MIERS ON THE PEN ZID&E. [ Mar. 5,
in the British-Museum collection do not enable me to adopt this
course, as many species are not as yet represented in it, I have, in
the synopsis of the species given below, and until a more complete
revision of the genus shall have become possible, included the species
in Penaeus.
Synoptic Table of the Species of Penzus.
The object of the following analytical Table is to indicate the prin-
cipal external characters by which the species of Peneus may be
distinguished. There are certain species which have not been
described with sufficient detail to allow of their incorporation in the
list ; these are either previously referred to, or are mentioned by Milne-
Edwards in the ‘ Histoire Naturelle des Crustacés.’ The species of
which there are no specimens in the British-Museum collection, and
which are therefore unknown to me, are distinguished by an asterisk,
I, Antennules with both flagella extremely short. Two deep longitudinal
sulci extending along the whole length of the cephalothorax, adjacent to
the rostrum and the median carina, which is itself deeply sulcate. (A
supraorbital spine present.)
First and second pairs of legs bispinose, third pair unispinose at base.
Seventh postabdominal segment with lateral marginal spines.
1. P. caramote, Risso.
European.
First pair of legs bispinose, second pair unispinose, third pair unarmed at
base, Terminal segment without lateral marginal spines.
2. P. brasiliensis, Latr.
Atlantic region.
First and second pair of legs unispinose, third pair unarmed at base. ‘Ter-
minal segment with lateral marginal spines.
3. P. canaliculatus, Oliv.
Indo-Pacific region.
II. Antennules longer. No lateral sulci on the cephalothorax, bordering the
rostrum and median carina. (Supra-orbital spine usually absent.)
A. Rostrum much longer than the eyes, reaching nearly to, or beyond, the end
of the antennal scale.
a. Carapace with a ridge extending backward from the origin of the rostrum
to, or nearly to, the posterior margin of the cephalothoraz.
§ The carina between the base of rostrum and posterior margin sulcate,
and reaching to the posterior margin.
First pair of legs bispinose, second pair unispinose, third pair unarmed at
base. Rostrum ‘dentate, third to sixth postabdominal segments keeled.
4. P. semisulcatus, De Haan,
Japan, Indo-Pacific region.
First and second pairs of legs unispinose, third pair unarmed at base.
Rostrum = dentate, curved upward toward the apex; third to sixth post-
abdominal segments keeled.
5. P. hardwickii, Miers,
Indian Seas.
Rostrum 2 dentate, slightly ascending toward the apex. All the postab-
dominal segments keeled.
*6, P. sculptilis, Heller.
Java.
1878. ] MR. E. J. MIERS ON THE PEN EIDE. 307
§§ The carina between the base of rostrum and posterior margin of the cephalo-
thorax not sulcate, and often not reaching to the posterior margin.
* Rostrum with the inferior margin dentate. (No lateral spines on last post-
abdominal segment.)
9-10 c . . -
Rostrum 5-5-dentate. Terminal segment ending in a long spine.
*7, P. setiferus, Linn.
West Indies, east coast of North America.
Rostrum a dentate, not reaching beyond the antennal scale. ‘Terminal
segment acute.
8. P. monodon, Fabr.
Indian and Australian Seas.
Rostrum yery slender and styliform in its distal half, qe-dentate and reach-
ing slightly beyond the antennal scale.
9. P. indicus, M.-Kdw.
Indo-Malayan Seas; Panama ?
** Rostrum with the inferior margin entire.
1. Rostrum not reaching far beyond the end of the antennal scale.
Rostrum ensiform, straight, 8-10-dentate above, the teeth extending along
nearly the whole upper margin. Terminal postabdominal segment without
lateral marginal spines.
10. P. monoceros, Fabr.
Indo-Malayan Seas.
Rostrum curyed upward toward apex, 8-dentate above, dorsal carina reach-
ing nearly to the posterior margin. Terminal segment as in P. monoceros.
*1. P. curvirostris, Stimpson.
Japan, Simoda.
Rostrum robust, curved slightly upward, 8-9-dentate above, the last tooth
situated on the dorsal carina at considerable distance from the preceding.
Terminal segment with a pair of lateral spines.
12. P. membranaceus, Risso.
Mediterranean, Madeira.
Rostrum styliform at apex, nearly straight, 7-9-dentate above in its proximal
half. Terminal segment without lateral marginal spines; 5th legs (in the
female) rudimentary. ;
13. P. dobsoni, Miers.
West Coast of India.
Rostrum slender, styliform and produced at apex, 6-9-dentate above at its
base. Terminal segment with prominent lateral marginal spines; 5th legs
(in the male) long and slender, dactylus small.
14, P. styliferus, Edw.
Bombay.
2. Rostrum reaching far beyond the end of the antennal scale.
Rostrum 9-10-dentate above; the 3-4 last teeth placed upon the slender
styliform distal half. Last postabdominal segment with lateral spines.
*¥15. P. foliaceus, Risso.
Mediterranean.
Rostrum 6-dentate above, the styliform distal portion without teeth. Ter-
minal segment without lateral spines.
16. P. Aroyeri, Heller.
Brazil West Indies.
b. Cephalothorax without any median dorsal ridge. (Rostrum entire below.)
Rostrum straight, dentate nearly to its apex; first tooth separated by a
20*
308 MR. E. J. MIERS ON THE PENHID. { Mar. 5,
wider interval than the preceding. Cephalothorax and postabdomen tomentose.
Terminal segment not carinate, with minute lateral spines.
17. P. affinis, Edw.
Indo-Pacific region.
Rostrum and cephalothorax as in P, affinis. Postabdomen naked. Termi-
nal segment with the deep longitudinal median sulcus defined by lateral carine,
and with lateral spinules.
18. P. constrictus, Stimpson.
East Coast United States.
Rostrum somewhat sinuated, 9-dentate above, with the teeth equidistant.
Terminal segment without lateral spines.
*19. P. tenuis, Dana.
North Patagonia.
Rostrum styliform toward apex, nearly straight, 6-dentate above near its
base; the last tooth separated by a wider interval. Terminal segment without
lateral marginal spines.
*20, P. avirostris, Dana.
Singapore.
B. Rostrum very short, not reaching beyond the eyes.
* Rostrum slender.
Eye-peduncles long. Rostrum not half as long as the eyes, 7-dentate above.
*21. P. podophthalmus, Stimpson.
Hong Kong.
Eye-peduncles of moderate length. Rostrum a little shorter than the eyes,
5-dentate above.
*22. P. gracilis, Dana.
Sooloo Sea.
Eye-pedunceles thick. Rostrum scarcely reaching beyond them, 8-9-dentate
above. (5th? legs greatly elongated, with the dactylus nearly half as long as
the cephalothorax.)
*23. P. stenodactylus, Stimpson.
Hong Kong.
** Rostrum laminiform, high, and laterally compressed.
Rostrum 9-10-dentate above. Flagella of the antennules half as long as
the peduncle.
*24, P, lamellatus, De Haan.
Japan.
Rostrum 6-dentate above. Flagella of the antennules about as long as the
peduncle,
* 95. P. brevicornis, M.-Edw.
Indian Seas.
ARISTEUS EDWARDSIANUS. (Plate XVII. fig. 3.)
Peneus edwardsianus, Johnson, P. Z. 8. 1867, p. 897.
The typical example of this very fine species is in the collection
of the British Museum; and I have little hesitation in referring it
to the genus dristeus of Duvernoy (Ann. Sci. Nat., sér. 2, Zool. ‘xv.
p- 101, 1841). The character principally relied upon by Duvernoy
for the distinction of the genus was derived from the ramose structure
of the branchize, which he conceived were formed upon a different
type from that found in other genera of the Caridea ; but I am able,
from an examination of the branchiz of 4. edwardsianus, to con-
firm Dana’s observation that they do not differ essentially in struc-
1878. ] MR. E. J. MIERS ON THE PENEZIDS. 309
ture from those of the species of Peneus. There are, however,
other points in which the genus Aris¢eus differs from Peneus, and
which seem to warrant the retention of Duvernoy’s generic name.
The two flagella of the antennules are very unequal ; one is extremely
short, the other remarkably long, as long as the animal itself; the
palpi of the mandibles are robust, not foliaceous and membrana-
ceous, as in Peneus ; the fifth pair of legs is very long and slender,
not shorter than the preceding pair; the first pair of postabdominal
appendages terminate in a single ramus, the second ramus being
rudimentary or entirely absent. I may add that the anterior margin
of the first postabdominal segment does not form an angle with the
lateral margins, as usual in Peneus, but the antero-lateral angles
are rounded.
Duvernoy established the genus upon specimens from Nice, which
he referred, no doubt correctly, to the Peneus antennatus of Risso
(Hist. Nat. Eur. Mérid. v. p. 68, 1826).
P. edwardsianus resembles in general appearance the figure of
P. antennatus given by Duvernoy ; but the carapace is marked
with strong ridges upon the hepatic, pterygostomian, and branchial
regions, the last postabdominal segment is not sulcate above, and
the rami of the postabdominal appendages, especially of the first
pair, are much longer. There are also some marked differences in
the mouth-appendages which can scarcely be due to inaccurate
drawing of the details of Aristeus antennatus; the palpus of the
mandible in A. edwardsianus is much more robust than in the
Mediterranean species, is densely hairy, and the third joint is
bilobate; the exognath of the second pair of maxillipedes is very
long, reaching beyond the end of the antennal scale, and more than
twice the length of the endognath itself, while that of the third
pair of maxillipedes is quite short.
The specimen is a female.
The genus Xiphopeneus of Smith (referred to above) resembles
Aristeus in the form of the carapace and rostrum, and length of the
posterior legs and postabdonimal appendages; but there does not
exist the marked disparity in length in the flagella of the antennules,
nor isit stated that the mandibular palpi and rami of the first pair of
postabdominal appendages differ from those of the ordinary Penei.
FUNCHALIA WOODWARDI.
Funchalia woodwardi, Johnson, P. Z.S. 1867, p. 896.
The mouth-organs of the unique example of this remarkable form
(which is in the British-Museum collection) have not been dissected ;
but the long, corneous, and somewhat sickle-shaped processes of the
mandibles are plainly visible, and suffice to establish its generic
distinctness ; the ‘‘ broad lamellar appendage” at each side of the
mouth, referred to by Mr. Johnson, is evidently the foliaceous
palpus of the mandible, the terminal joint of which is large and
truncated at its distal extremity.
The specimen is a female.
310 MR. ANGAS ON A NEW GENUS OF LAND SHELLS. __|[{ Mar. 5,
Sicyonia Furcata, sp.n. (Plate XVII. fig. 4.)
This species, in the form of the body, closely resembles Sicyonia
ocellata, Stimpson (Proc. Ac. Nat. Sci. Phil. p. 43, 1860), of which
young specimens from Hong Kong, presented by the Smithsonian
Institution are in the British-Museum collection, and also one fully
adult from Ceylon, presented by E. W. H. Holdsworth, Esq. As in
S. ocellata, the dorsal carina is very convex and strongly dentate,
the cephalothorax tomentose, and the first to sixth postabdominal
segments deeply channelled and without spines on the lateral margins.
The terminal segment, however, is of very different form, being fur-
cate at the extremity, the lobes of the fork divergent, acute at apex
and deeply channelled above, the inner and outer margins carinate
and ending in small subterminal spines on each lobe of the fork.
In S. ocellata the last postabdominal segment is simple and acute at
apex.
Hab, Sooloo Islands.
A single adult example is in the collection. This specimen, and
also those of S. ocellata that I have seen, appear to be of the
female sex.
EXPLANATION OF PLATE XVII.
Fig. 1. Peneus hardwichii (p. 300), lateral view, natural size.
Ja, Terminal segment and uropoda of the same, natural size.
2. Peneus dobsoni (p. 302), lateral view of cephalothorax, natural size.
2a. Mandible, magnified.
26. Second maxilla, magnified.
2¢. First maxillipede, magnified.
2d. Second maxillipede, magnified.
2¢, Third maxillipede, magnified.
2f. Part of the ventral surface of the cephalothorax of a male individual,
showing the form and position of the fifth pair of legs (from a
specimen lent by Prof. J. Wood-Mason, see note p. 308).
2q. The same view of a female, showing the rudimentary and indurated
condition of the fifth pair of legs.
3. Exognath of the mandible of Aristeus edwardsianus (p. 308), natural
size.
4. Sicyonia furcata (p. 310), lateral view, natural size.
4a, Terminal segment and uropoda of the same, magnified.
4. Description of a new Genus of Land Shells belonging to
the Family Cyclophoride. By Guorcr Frencu ANGas,
C.M.Z.8:, F..8i, &e:
[Received Feb. 16, 1878.]
Genus Mascara, Angas.
Operculum horny, ovate, pointed above ; whorls few ; nucleus situ-
ated near the base of the columellar margin.
Shell rimate, elongately ovate, conically turreted, nearly smooth
or faintly longitudinally striated ; aperture ovate, rounded below and
angulate above ; peristome continuous, single; outer lip thickened
and slightly expanded.
Hanhart imp.
GSowerby lth
NEW SHELLS FROM KURRACHI, MADAGASCAR,
ECUADOR & AUSTRALIA
1878. ] ON NEW LAND AND MARINE SHELLS. 311
J have established this genus for the reception of certain species
of land shells that have hitherto been included in the genus Megalo-
mastoma of Guilding, a natural West-Indian group, from which,
however, they appear to me to differ so greatly in several important
characters that I do not hesitate to separate them from the Megalo-
mastomata as a distinct natural group, having its habitat in Madagas-
car and the Mascarene Islands.
The type of my genus Mascaria is the Cyclostoma croceum of
Sowerby, which has been placed in the genus Megalomastoma by
H. & A. Adams, Pfeiffer, Troschel, and other authors. With it
must be included a species recently described by M. Morelet, in the
‘Journal de Conchyliologie’ (tome xvii. No. 2, April 1877), under
the name of Megalomastoma litteratum.
Mascaria crocea is peculiar to the Mauritius, where it was found
alive by Sir David Barclay on the hills in the neighbourhood of
Black River. M. litterata is from Madasgascar, and was collected
there in some abundance by Mr. Waters.
5. Descriptions of nine new Species of Land and Marine
Shells from various localities. By Grorcs Frencu
Aneas, C.M.Z.S., F.L.S., &c.
[Received February 16, 1878. ]
(Plate XVIII.)
Buimus waters, n. sp. (Plate XVIII. fig. 1.)
Shell imperforate, elongately turreted, rather thin, shining, finely
irregularly longitudinally striated, the striee here and there forming
tessellated rows and patches, pale olive, lighter towards the apex,
mottled with small brown spots and irregular markings, with indica-
tions of two paler bands, one below the suture, and the other towards
the base of the last whorl ; spire acuminate, somewhat obtuse at the
apex ; whorls 9, rather convex, the last of equal length with the
spire ; sutures impressed and slightly crenulate ; aperture subovate,
one third the length of the shell, pale violet within ; outer lip simple,
scarcely thickened at the edge; columella with a slight callus below,
a little arcuate towards the base.
Alt. 3 inches, diam. 11 lines; length of aperture 1 inch.
Hab. Madagascar. :
This species is allied to B. obtusatus, Gmel., and B. moreleti,
Desh., with which, including perhaps B. clavator, Petit, and B.
balstoni, Angas, it combines to form a natural group of Madagas-
car Bulimi. The specimen from which my description is taken is
in the collection of Sir David Barclay. There is a second example
in that of Mrs. De Burgh.
I would here correct the name of a species in the list I gave in the
Society’s ‘ Proceedings’ for 1877 (p. 527), of certain land shells from
the Island of Madagascar that had been submitted to me by Mr.
312 MR. G. F. ANGAS ON NEW | Mar. 5,
Edward Bartlett. For Helix cornu-giganteum, Chemn., read H.
questieriana, Crosse (see Journ. de Conchyliologie, 1868, p. 269, pl.
ix. fig. 4). A large series of specimens of this latter species were
sent home lately from Madagascar by Mr. Waters; and although I
observed certain peculiarities about them on comparing them with
typical specimens of H. cornu-giganteum, I then hesitated to sepa-
rate them from the latter species, regarding them as possibly a local
variety, and not being aware at the time that they had been elevated
to the rank of a species by M. Crosse. I have now great pleasure
in adopting that gentleman’s name.
Helix guestieriana differs from H. cornu-giganteum, Chemn.
(=H. vesicalis, Lam.), in being somewhat smaller, paler in colour,
and more tumid, with the base less flattened and the umbilicus con-
cealed by a callus, and especially by having the inner lip constantly
more or less granulated—a character that I do not see referred to in
the description of the shell in the French Journal.
Butimus (OrosroMus) QUADRIFASCIATUS, 0. sp. (Plate XVIII.
figs. 2, 3.)
Shell elongately ovate, rimate, moderately solid, shining, finely
longitudinally striated, white, transversely banded with light brown,
the last whorl with four bands, the central one being the widest ;
spire conical, apex obtuse; whorls 6, somewhat convex; aperture
oval, eyual to the spire; outer lip flattened and expanded at the
base. 7
Alt. 1 inch 3 lines, diam. 64 lines.
Hab. Ecuador.
Butimus (Orostomus) NAPo, n.sp. (Plate XVIII. figs. 4, 5.)
Shell rimate, ovately fusiform, moderately solid, shining, very
finely and irregularly striated, pale fawn-colour, with a narrow white
band next below the suture and a similar white band surrounding the
perforation ; spire sharply conical, somewhat obtuse at the apex ;
whorls 6, slightly convex ; aperture ovate, same length as the spire ;
outer lip expanded and flattened at the base, white behind, bordered
by a narrow suffused orange band.
Alt. 1 inch 3 lines, diam. 5} lines.
Hab. Ecuador.
Buutimus (Eurytus) Eros, n. sp. (Plate XVIII. figs. 6, 7.)
Shell imperforate, oblong-ovate, thin, very finely and closely sha-
greened all over, the apical whorls marked with irregular longitudinal
strize, light-greenish olive, ornamented with small spots and cloudy
patches of a darker colour; spire very short, apex obtuse, pink ;
whorls 4, rather convex, the last very large; aperture ovate, more
than two thirds the length of the shell, effuse below; outer lip
slightly expanded and reflexed, which, together with the columella
and the interior of the aperture, is of a bright rose-colour.
Alt. 1 inch 54 lines, diam. 8 lines.
Hab. Ecuador.
1878. | LAND AND MARINE SHELLS. 313
A beautiful species, characterized by its closely granulated sculp-
ture, and the rosy coloration of the mouth.
ROSTELLARIA LUTEOSTOMA, n. sp. (Plate XVIII. figs. 8, 9.)
Shell fusiform, solid, light brown, paler below the sutures ; spire
attenuated, apex sharp-pointed; whorls 15, the first 9 or 10
longitudinally strongly plicate, those nearest the apex being cancel-
lated with fine transverse ridges, the lower whorls nearly smooth,
with a few concentric grooves near the base or the last whorl; colu-
mella covered with a strong tumid polished callus; outer lip with 5
digitations on the lower half of the margin ; posterior canal extend-
ing upwards beyond the centre of the third whorl; beak moderate,
nearly straight in the young shell, somewhat curved outwards in
the adult ; aperture golden yellow, deepest on the lower portion of
the columella.
Length 4 inches 3 lines, diam. 1 inch 3 lines.
Hab. Kurrachi, near the mouth of the Indus.
This beautiful shell forms an interesting addition to the restricted
genus Rostellaria (Gladius of Klein), of which hitherto only about
half a dozen recent species have been described.
All the known species inhabit the Asiatic seas, none having been
met with in any part of the New World. The two examples (one
young and the other in the adult state) from which I have taken my
description, are from the collection of Sir David Barclay, and are, so
far as I am aware, unique in this country.
Mirra (CosTELLARIA) LINCOLNENSIS, n. sp. (Plate XVIII.
figs. 10, 11.)
Shell acuminately fusiform, solid, whitish, tinged with irregular
longitudinal chestnut flames, with a narrow band of interrupted spots
encircling the centre of the whorls, the lower half of the Jast whorl
chestnut, with a faint band of reticulated brown and white spots in
the middle ; whorls 7, a little convex, with numerous, stout, rounded
longitudinal ribs, which are slightly nodulous below the sutures ;
interstices smooth, sutures impressed ; spire longer than the aperture;
outer lip simple, a little contracted below ; columella with 4 strong
plaits, the posterior one the largest: basal canal short, slightly re-
curved.
Length 7, breadth 2 lines.
Hab. Port Lincoln, South Australia.
Mirra sCHOMBURGKI, n.sp. (Plate XVIII. figs. 12, 13.)
Shell ovately fusiform, moderately solid, strongly longitudinally
ribbed, the ribs becoming nearly obsolete towards the lower portion
of the last whorl, livid brown, with a broad pale suffused band on
each whorl and four narrow dark brown lines encircling the last
whorl, one above and three below the band; whorls 64, convex ;
spire turreted ; sutures impressed; aperture subovate, equal in
length to the spire; outer lip simple, arcuate, sulcate internally ;
314 MR. G. E. DOBSON ON BATS { Mar. 5,
columella with three stout plaits, which are carried round the base
of the last whorl. é
Alt. 5, diam. 2 lines.
Hab. South Australia.
SIPHONARIA ALBIDA, n. sp. (Plate XVIII. figs. 14, 15.)
Shell elongately ovate, subsymmetrical, sharply conical, rather thin,
white ; ribs very numerous, rounded, unequal, crossed more or less
with squamose ridges ; apex subcentral, prominent, smooth, recurved ;
siphuncle inconspicuous ; interior white, shining, almost pearly.
Long. 8, lat. 6, alt. 34 lines.
Hab. St. Vincent’s Gulf, South Australia.
A pure-white conical species, with the ribs very numerous and
crossed with irregular squamose ridges.
Lepa (ApRANA) NEWCOMBI, n.sp. (Plate XVIII. figs. 16, 17.)
Shell equivalve, very inequilateral, flattened, lanceolate, the poste-
rior side obtusely angled, the anterior terminating in a point slightly
curved upwards, thin, shining, white, sculptured with very fine, re-
gular, close-set concentric strize, which on the posterior side extend
nearly to the margin of the valves, whilst anteriorly they cease
abruptly, leaving about one third of the surface smooth, faintly
marked with the lines of growth only; front dorsal margin very
slightly concave, two thirds the length of the shell ; ventral margin
somewhat arcuate behind, flattened below, and again slightly arcuate
towards the anterior extremity ; beaks very small, approximate, in-
curved.
Length 1 inch 4 lines, alt. 4 lines, lat. 24 lines.
Hab. Dredged in Navy Bay, Aspinwall, in 8 fathoms (Dr. W.
Newcomb).
I have much pleasure in dedicating this new species of Adrana to
my old friend and correspondent Dr. W. Newcomb, of Ithaca, a
distinguished American conchologist.
EXPLANATION OF PLATE XVIII.
Fig. 1. Bulimus watersi, p. 311. | Fig. 10, 11. Mitra lincolnensis, p. 313.
2, 3. quadrifasciatus, p. 312. 12, 13. schomburgki, p. 313.
4, 5. —— napo, p. 312. 14,15. Stphonaria albida, p. 314.
6, 7. eros, p. 312. | 16,17. Leda newcombi, p. 314.
8, 9. Rostellaria luteostoma, p, 313.
6. Additional Notes on the Chiroptera of Duke-of-York
Tsland and the adjacent Parts of New Ireland and New
Britain’. By G. E. Dozson, M.A., M.B., F.L.S., &c.
[Received Feb. 18, 1878.]
The following notes are derived from an examination of a second
collection of Chiroptera, consisting of forty-five specimens, recentl
y I ’ y
1 See P. Z. 8. 1877, p. 114.
1878. | FROM DUKE-OF-YORK ISLAND. 315
sent home by the Rev. G. Brown, C.M.Z.S., and kindly placed in
my hands by Mr. P. L. Sclater, Secretary of the Society.
This second collection contains no species which were not repre-
sented in the first; but the opportunity afforded of examining ad-
ditional specimens of certain species enables me to correct and add
to the notes published in my first paper, and to show that the Chiro-
pterous fauna of Duke-of-York Island is even more characteristic of
the Australian region than there stated.
Preropus MELANOPOGON, Var. NEOHIBERNICUS.
Pteropus melanopogon, var. neohibernicus, Peters, Monatsb. Akad.
Berl. 1876, p. 317.
Three specimens, an adult female with forearm 7'-3, one younger
with forearm 7'':0, and one quite immature with forearm 6’°5. All
have the ear one inch in length, and quite similar in shape to that of
typical examples of P¢. melanopogon from the Malay archipelago.
The index finger appears to be remarkably long, probably propor-
tionally longer than in any other species of the genus. In the
largest specimen (with forearm 7°3 inches) it is 5:8, the metacarpal
bone of the middle finger being 5:1, while in a full-grown specimen of
Pt. melanopogon from Bouru (with forearm 7'*75), in which the meta-
carpal of the middle finger is 5''"2, the index finger measures 5!'*5
only. Jn all the specimens the second upper premolar is separated
by a rather narrow space from the canine, and the premolars and
molars are very large, but with scarcely developed internal cusps,
and no trace of basal projections. In the older female the fur is very
short, and almost bright yellowish brown throughout, the back of the
head and across the shoulders brighter than the rest of the body, on
the thighs brown, the fur of the face and throat of a reddish brown
shade, but no trace of a black patch of hairs beneatlf the chin and
throat. In the younger female the fur is much longer and darker,
but, as in the older specimen, it is reduced to a narrow line about an
inch wide along the centre of the back ; the face about the eyes, the
cheeks, and under the muzzle and throat reddish brown, the whole
under surface paler reddish brown ; the back of the head bright yel-
low ; the neck above and on the sides light yellowish brown, shoulders
brighter; the narrow line along the centre of the back bright yellow-
ish, edged with dark brown, which colour extends upon the hairs
clothing the thighs. The immature specimen rather resembles the
older one; but the back of the head is paler yellow, the face and chin
darker brown, and the fur along the back is much broader and
almost uniformly light brown.
PrreRoPus CAPISTRATUS.
Pteropus capistratus, Peters, Monatsber. Akad. Berl. 1876,
p. 316 (with a coloured plate); Dobson, Proc. Zool. Soc. 1877,
p- 116.
A single female specimen, not differing from those previously de-
scribed; the black ring surrounding the muzzle between the ears
316 MR. G. E, DOBSON ON BATS [Mar. 5,
and eyes very conspicuous where it passes round the lower jaw in
front of the throat.
CyNONYCTERIS BRACHYOTIS.
Cynonycteris brachyotis, Dobson, P. Z. 8.1877, p. 116.
A large number of specimens representing all ages of this animal ;
so that the full size of the species can be determined with considerable
certainty. This scarcely differs from that given in my original de-
scription. In the largest specimen, an adult male, the forearm is
2°9 inches, or only one twentieth of an inch longer than that of the
type. In all, the peculiar nakedness of the shoulders is present,
and the fur of the sides of the neck radiates from a central point
near the place of origin of the antebrachial membrane, corresponding
to the position of neck-glands, which, in the adult male referred to
above, are covered on each side by a circular tuft of coarse yellow
hairs, as in most of the species of the genus Pteropus. The colour
of the fur is very similar in all, being greyish yellow-brown, the base
of the hairs darker ; in the immature specimens the fur is longer
and darker throughout. In all, except the oldest individuals, there
is a minute first upper premolar between the approximated canine
and second premolar, generally so small as to be seen with diffi-
culty.
CEPHALOTES PERONII.
Cephalotes peronii, Geoffroy, Ann. du Muséum, xv. p. 104
(1810).
More than one third of the whole collection consists of specimens
of this species, which, therefore, appears to be very abundant in these
islands.
From the yeung with milk-dentition to the aged with worn teeth,
all have the back equally naked from the shoulders backwards.
They correspond closely in measurements and other respects with
specimens from other parts of the Austro-Malayan subregion, of
which this species is eminently characteristic.
MELONYCTERIS MELANOPS.
Melonycteris melanops, Dobson, P. Z.S. June 1877, pp. 119-121,
figs. 4—7, and pl. xvii.
Pteropus (Cheiropteruges) alboscapulatus, Ramsay, Proc. Linn.
Soc. New South Wales, July 1877, p. 17.
An adult female, scarcely differing in the colour of the fur from
those previously described, having also the white spot near the place
of origin of the antebrachial membrane from the shoulder. The
canines appear to be as large as in the male. The wings are at-
tached posteriorly, as in other specimens, to the base of the third
toe, or to the space between the bases of the second and third
toes.
This is undoubtedly the species described under the name of Pte-
ropus (Cheiropteruges) alboscapulatus by Mr. Pierson Ramsay in
1878. | FROM DUKE-OF-YORK ISLAND. 317
the publication referred to above, as may be seen by comparing the
descriptions. In my original description of this species, I did not
notice the white spot on the shoulder, very correctly described by
Mr. Ramsay, as it was completely concealed from view in the speci-
mens examined by me (owing to the manner in which the dry skins
were preserved, the wings being brought forward in front of the
head); nor have I referred to the yellow spots ou the wings, which,
as they do not appear in this specimen, may be accidental.
PHYLLORHINA TRICUSPIDATA.
Rhinolophus tricuspidatus, Temm. Monogr. Mammal. ii. p. 20,
pl. 32. figs. 11, 12 (1835-41).
An immature specimen with light-brown fur, of a dull shade
throughout, very different from the bright reddish brown fur of the
adult individual from the same locality referred to in my former
paper.
PHYLLORHINA CERVINA.
Rhinolophus cervinus, Gould, Mammals of Australia, iii. pl. xxxiv.
(1853).
A careful examination of the additional specimens in this collec-
tion has shown me that I was mistaken in referring to Ph. galerita,
Cantor, of the Oriental region, a single specimen in the first collec-
tion, which really belongs to the Australian representative of that
Phyllorhina cervina. Phyllorhina galerita.
species. Recent specimens, and specimens preserved in alcohol, of
this Australian species (Ph. cervina, Gould) are readily distinguished
from the closely allied Ph. galerita by the much larger frontal
glandular sac in adult males, by the much narrower anterior portion
of the horizontal horseshoe-shaped nose-leaf (compare the woodcuts
above), and by the much greater length of the calcaneum. In
dried skins, however, these parts become so altered in appearance
that it is almost impossible to distinguish the species.
PHYLLORHINA CALCARATA.
Phyllorhina calcarata, Dobson, P. Z. 8. 1877, p. 122, fig. 8.
A specimen of an immature individual with (as in the type) ver-
tical ridges dividing the concave front surface of the transverse erect
318 MR. R. COLLETT ON CERTAIN GOBIOID FISuES. [ Mar. 5,
portion of the nose-leaf, and having also the last caudal vertebra
free.
Ph. coronata, Peters, from the Philippine Islands, approaches this
species in size and in the remarkable length of the calcaneum ; but the
very different form of the nose-leaf and ears at once distinguishes it.
7. On Latrunculus and Crystallogobius, two remarkable
Forms of Gobioid Fishes. By Roserr Corer, of
Christiania, C.M.Z.S.
[Received February 18, 1878.]
Upon different occasions, at meetings of the Scientific Society of
Christiania, I have communicated my observations on a singular
group of our Gobioid fishes, which differ from the typical genus
Gobius not only in the almost perfect transparent body, but also in
differences in the structure of the scales, fins, and the form of the
body’.
As the said group in several points offers much of interest, I will
endeavour to relate the general results of these researches, all of
which have been made in the Christiania fjord, a most favourable
locality for marine fishes and invertebrates. From these researches I
find :—
1. Each of the genera, Latrunculus, Gunth., 1861, and Crystallo-
gobius, Gill, 1863, include one single species, L. pellucidus (Nardo)
1824, and C. nilssoni (Diib. & Kor.) 1844.
2. Latrunculus pellucidus (Nardc) from the Mediterranean, L.
pellucidus (Kessler) from the Black Sea, L. albus (Parn.) from
Scotland, and LZ. stuwitzii (Diib. & Koren) from Norway are all
one and the same species.
3. The males of both species havea dentition totally different from
that of the females; in Z. pellucidus these teeth are only developed
during the spawning-season.
4. The female of C. nilssonii has the first dorsal and the ventral
fins rudimentary ; and the ovaries of this species reach back far
behind the vent.
5. Both species are annual vertebrates, and accomplish their life
in the course of one year.
Genus Latruncuuus, Giinth.
Genus Latrunculus, Giinth. Cat. Acanth. Fishes, Brit. Mus.
vol. ili. p. 80 (1861).
Body transparent, elongate, rather compressed, covered with large,
deciduous, rounded scales attached to parallel muscular impressions.
Both sexes dentiferous, the mature male with a series of long teeth,
1 Sleeten Latrunculus og dens nordiske Arter (Forh. Vid. Selsk. Christ. 1872).
Norges Fiske, med Bemezrkunger om deres Udbredelse (Tillegsh. til Forh. Vid.
Selsk. Christ. 1874). Om Slegterne Latrunculus og Cryscallogobius (Forh. Vid.
Selsk. Christ. 1876).
1878.] MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 319
and with strong canine teeth, which form the hinder series in the
upper jaw. Two separated dorsal fins, the first composed of 5
spines. Ventral fins united, gill-openings rather wide. Pseudo-
branchiz present.
Latrunculus pellucidus 3 ad., in the breeding-season, 2.
L. pELLuciDuS (Nardo).
1824. Gobius pellucidus, Nardo, Giorn. Fis. Chim. Stor. Nat.
Pavia, tom. iii. Osserv. p. 7 (teste Giinther).
1826. ? Aphia meridionalis, Risso, Hist. Nat. Eur. Mér. tom. iii.
p- 287 (Paris, 1826).
1837. Gobius albus, Parn. Trans. Roy. Soe. Edinb. vol. xiv. p-
137, Feb. 1837 (Edinb. 1840).
1844. Gobius stuwitzii, v. Diib. & Kor. Kel. Vet. Akad. Férhandl.
1844, p. 51, tab. ii. fig. 4 (Stockh. 1846).
1859. Gobius pellucidus, n. sp., Kessl. Bull. Soc. Nat. Mose. 1859,
tom. xxxii. No. 2, p- 260 (Mose. 1859).
1861. Latrunculus albus, Ginth. Cat. Acanth. Fish. Brit. Mus.
vol. iii. p. 80, and p. 556 (Lond. 1861).
1861. Latrunculus pellucidus, Giinth. Cat. Acanth. Fish. Brit.
Mus. vol. iii. p. 556 (Lond. $861).
1861. Gobiosoma stuvitzii, Giinth. Cat. Acanth, Fish. Brit. Mus.
vol. iii. p. 86 (Lond. 1861).
1863. Boreogobius stuvizii, Gill, Proc. Acad. Nat. Sci. Philad.
1863, p. 269 (Philad. 1864).
1872. Latrunculus stuwitzii, Coll. Forh. Vid, Selsk. Christ. 1872,
p- 9 (Christiania, 1873).
Teeth in the mature male long and cylindrical, and with strong
curved canine teeth in both jaws; teeth in the female and the im-
mature male minute. The ventral fins united to a funnel, with the
longer rays towards the body. The ovaries do not project behind
the vent.
D'5, D’ 13 (12); C. 10417410; A. 14; V.6; P. 16;
Lin. lat. 25; Vertebr. 27; M. Br. bi
A. Remarks on the Synonymy.
The first northern form of the transparent Gobioids was discovered
by Parnell in the Scottish friths, and shortly described in Feb. 1837,
in the | 4th volume of the Trans. of the Royal Society of Edinburgh,
under the name of Godius albus. His description (which, by the way,
320 MR. R. COLLETT ON CERTAIN GOBIOID FISHES. [ Mar. 5,
is rather incomplete) bas afterwards, without any further addition, been
repeated in the various editions of Yarrell’s History of Brit. Fishes.
In May, 1844, Von Diiben and Koren in Ofv. Kgl. Vet. Akad. Forh.
for the same year introduced two other species of the same group,
both of which had been previously found, in Dec. 1834, in the district
of Bergen, Norway, by the late naturalist, Mr. P. Stuwitz, after whom
one of them was named Goédius stuwitzii. The last-named species
had certainly, as } ointed out by the authors, a decided similarity to
the Scottish G. albus. Still the difference in the construction of the
body, as well as of the teeth, was so obvious that they could not
easily be mistaken. Instead of being plump, like ZL. albus, with the
head thick, the cleft of the mouth wide, the teeth long, the interorbital
space wide, the second dorsal and the anal fins even, G. stuwitzi
was slender, with the head thin, and the cleft of the mouth short,
the teeth extremely feeble, the interorbital space narrow, and the
second dorsal and the anal fins diminishing behind. In 1861, Dr.
Malm, in Bohuslin, Sweden, found both the named species in fully
developed state, one of them (G. stuwitzii) with ripe spawn; and
he therefore gave, in July 1863, in the Foérh. Skand. Naturf. 9. Mote,
comparative diagnoses of them, chiefly based upon the structure of
the teeth and the vertical fins.
When Dr. Giinther, in 1861, issued the volume of his Catalogue
of the Fishes of Brit. Museum, in which he treats of the Gobioids,
G. albus was made the type of the new genus Latrunculus'; but,
from the description of . s¢uwitzii, he did not venture to include
this under the same genus; for want of a better place he inserted it
under the genus Godiosoma, Gir., as it was still undecided that it
possessed scales, which, for the first time, was proved by Dr. Malm
in 1863. Prof. Gill, in the Proc. Acad. Nat. Sci. of Philadel-
phia for 1863, even made it a type for a new genus Boreogobius,
without giving even asingle word of reason for this alteration; and
Dr. Bleeker, in 1874, followed in his steps in the Synopsis of
the Gobiide, which he communicated in the 9th volume of Arch.
Neer]. Sci. Exact. et Nat.
Since the autumn of 1871, I have found the form Z. stuwitzii in the
Christiania fjord in great numbers, and have collected innumerable
specimens during the autumn months. All these specimens have
fully agreed with the original description given by Diiben and Koren,
and consequently are constantly different from that of Parnell’s LZ.
albus. Having provided myself during the first autumn with what
I thought sufficient materials, consisting of several thousand speci-
mens, I gave, in the Forh. Vid. Selsk. Christiania, 1872, a short de-
scription of the species, comparing it with the descriptions at hand
of L. albus, in which I endeavoured as much as possible to keep the
two species separate. During the succeeding years I tried amongst
the great quantities of L. stuwitzii to find L. albus, but without
1 The name Latrunculus was employed by Dr. Gray as early as in 1847, for
a Gasteropod (Proce. Zool. Soc. Lond. 1847, p. 159). But, as I was informed
by my friend the late Dr. Mérch, of Copenhagen, this name is quite superfluous,
and will hardly ever be reemployed for that mollusk.
1878.] MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 321
success. In the winter months the fjord is covered with ice; and in
the spring aad summer I had no opportunity of tracing them. In
August 1874 I certainly found a couple of breeding females, together
with some others which had already spawned ; but none of these
differed in any point from the typical LZ. stuwitzii; and in September
again the young of the year solely were seen. Therefore I could
add nothing of importance to my previous researches published in
the ‘ Norges Fiske’ in 1874.
It was almost accidentally that, in June 1875, I found one of
the spawning-places ; and during a couple of excursions I succeeded
in collecting a considerable number of fully developed specimens of
both sexes. It then became quite evident that L. s¢uwitzii was based
on the female and the immature males of the same species, the mature
males of which were Z. albus. On several occasions examples of
typical Z. albus, which all, without exception, proved to ve old males,
were taken from the same shoal as ZL. stuwitzi, all of which were
females ready for spawning. Another lot consisted of somewhat
younger specimens in various stages of transformation between both
forms, all full-grown, but with the eggs and milt not yet perfectly
ripe.
Von Diiben and Koren described their species from four specimens
taken in winter (in the month of December), and consequently out of
their spawning-season; but Parnell, on the contrary, had for the
description of his Gobius albus 50 specimens caught in June in
Solway Frith, thus just during their breeding-time. From Parnell’s
description we may conclude that he had only mature males at his
disposal ; nor does Dr. Giinther, in compiling his catalogue (1861),
seem to have had at his disposal any females,—as may be seen
from the fact that both these authors speak only of individuals
with long teeth. By the kindness of Dr. Giinther I have received
three specimens of the Scottish form, collected by Parnell himself ;
and one of these, curiously enough, proves to be a female with the
minute teeth and the sloping fins, in other words, a typical L.
stuwitzii. Thus Parnell had overlooked female individuals, perhaps
few in number, and had drawn his description solely from the
males.
Having endeavoured to identify the two species of Western
Europe, it remains to discuss the third and fourth species of this
genus, the LZ. pellucidus of Southern Europe.
In 1824, according to Dr. Giinther, a description was given by
Nardo of a Mediterranean form under the name of Gobius pellucidus.
I have not had an opportunity of seeing the periodical in which
Nardo gave his original description (Giorn. Fis. Chim. di Stor.
Nat. d. Pavia); but from the title of the paper (“ Osservazioni ed
ageiunte all’ Adriatica Ittiologia”’) it seems that his specimens were
taken in the Adriatic. The species since Nardo’s time has often
been mentioned by those naturalists who have treated upon the fauna
of the Mediterranean, and, as it appears, under somewhat different
names. In Hist. Nat. Eur. Mérid. tom. iii. p. 287 (1826), Risso
mentions a species under the name Aphia meridionalis, n. gen. et sp.,
Proc. Zooxu. Soc.—1]878, No. XXI. 21
322 MR. R. COLLETT ON CERTAIN GOBIOID FISHES. [ Mar. 5,
which perhaps may be the same as Risso’s, although it can hardly be
proved with any certainty, as the description is, in the first place, too
incomplete, and, secondly, disagrees in several respects with our
species. It is therefore scarcely advisable to adopt the generic name
given by Risso; butif this naturalist really had in view LZ. pellucidus
(which is very probable), his description, besides being incorrect’, is
also incomplete, from no notice having been taken of the mature
male. Another generic name, which is ascribed to Nardo, is Brachio-
chirus (ef. Ronap. Cat. Method. Pesci Europ. p. 64, Napoli, 1846,
and Atti Ist. Veneto, tom. v. ser. 3, p. 796, Venezia, 1859-60); but
Ihave had no opportunity of examining the work wherein that genus
is established*. Canestrini mentions the species, in his memoirs on
the Gobioid Fishes found in the Gulf of Genoa (Arch. Zool. Anat.
Fisiol. fase. ii. vol. i. p. 152), under the name of Godius albus, Parn.
This author shows the difference in the structure of the teeth be-
tween the sexes in the Mediterranean form, without, however,
bringing his description to bear upon the young male.
Last year I received from Prof. G. O. Sars a couple of specimens
of the Adriatic form, collected by him during a stay ia Trieste in
the spring of 1876. As it seems to abound there, and is well known
to all fishermen, it is very possible that it is Nardo’s G. pellucidus;
and it has therefore been of much interest to me to be able to make
a comparison between this and the northern form, with which it
agrees in development (ZL. stuwitzii). This comparison has been
thoroughly conclusive of the fact that they are identical. The speci-
mens from Trieste were collected in the month of March, and had
then not yet reached their fully mature state. Their total length
varies between 42 and 44 millims.; otherwise the length of the
head, the size of the eyes, the general structure of the body, the
number of rays and of the transverse muscular strie—further, the
teeth—in short, every thing, down to the most minute spots of the
coloration, completely agrees with specimens of the same size and
state of development from Christiania.
Finally, Kessler, in Bull. Soc. Imp. Nat. Mosc. tom. xxxii. (1859),
gives a tolerably detailed description of a form from the Black Sea
(Odessa), which he calls Godius pellucidus, n. sp., being apparently
ignorant of Nardo’s previous description of a species of the same
name. )
With regard to this form, it does not seem to differ at all from ZL.
albus (or the male in spawning-season). The distinction between
them, which Dr. Giinther thinks may be made, that Kessler’s
species has a larger diameter of the eye, namely } of the length of
1 This is especially the case with regard to the description of the fins and
their rays ; the number of the rays in the first dorsal is also omitted, which is a
point of great importance.
? If in Risso’s Aphya and Nardo’s Brachiochirus we can recognize Latrun-
culus pellucidus, both these names will have priority over Latrunculus, Ginther.
The decision of this, as well as other similar questions of priority, I leave to
those who haye greater opportunities of examining the literature of Southern
Europe than can be obtained here in the library of the University of Chris-
tiania.
1878.] MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 323
the head, and equal to the snout, whilst, on the contrary, the eye of
L. albus is contained 5 times in the length of the head, and is
shorter than the length of the snout, is of little consequence, as in
fact both the original specimens of Parnell and those from the
Christiania Fjord have rather the proportions of length given by
Kessler. Although this author expressly remarks that he knows
the mature females, he must have overlooked (as well as Parnell)
the differences between these and the males, and in his description
(for instance, of the structure of the teeth) only have taken notice
of the latter.
B. General Description.
Before entering into a more detailed description of the species in
general, I will briefly point out some of the principal peculiarities
which separate the individuals in the different periods of life. From
this it will be seen that the difference in the sexes of the species in the
fully developed state is considerably greater than with any of our
northern species of the typical genus Gobius.
Out of the spawning-season the males and the females (which at
this time only exist in the shape of young specimens) are exactly alike
in appearance, although they soon reach the size of the old ones.
The body is slender, the head pointed ; the jaws, in consequence, are
less oblique, and seem, therefore, to be shorter. The teeth are all
uniform, minute, and form a close single row. The caudal fin is
slightly emarginated, the second dorsal and the anal diminishing
behind. In this stage they are all typical L. stuwtzie.
The nearly mature males, the milt of which is not yet quite ripe,
have the head considerably more thickened, and longer jaws ; besides
the minute teeth, there is become visible an outer row of cylindrical
teeth, yet not full-grown, but having reached a greater length than
the others. The posterior rays of the second dorsal and the anal
fins are still lower than the anterior ; the caudal is slightly emargi-
nated, or square.
Fig. 2.
Head of Latrunculus pellucidus, mature male, magnified.
The mature males have an average total length (the caudal in-
cluded) of 42—43 millims. (a few have the milt ripe, with a total
length of 40 millims.), but reach a size of 50-51 millims. The body
is plump ; the head thick and tumid ; the interorbital space broad ;
the cleft of the mouth wide, with fully developed wel only one
324 MR. R. COLLETT ON CERTAIN GOBIOID FISHES. [ Mar. 5,
kind, few in number, but long and cylindrical. The posterior rays
of the anal and second dorsal fins are, as a rule, not shorter than the
anterior, making the fins about even in their whole length. The caudal
fin is slightly square, with rounded edges in the young individuals,
but truncated in the elder ones.
The nearly mature females, not yet quite ready for spawning,
agree in general with the individuals in the immature state, from
which they differ in the somewhat swollen belly.
The females ready for spawning have an average length of about 42
millims. ; many having the spawn ripe are of a length of 58 millims. ;
none reach a length of more than 46 millims. (the caudal included).
The head is not visibly thickened, but is about the same as during the
whole young period; it is rather more pointed than in the male;
the cleft of the mouth is considerably smaller, although the jaws
may possibly have become a trifle longer than before. The teeth,
however, have not undergone any change, but are uniform, exceed-
ingly fine, and placed in the same row as in the young individuals.
The caudal fin is emarginated, and the anal and the second dorsal
are tapering behind, as is the case in their younger state.
The construction of the body is slender and elongated, laterally
compressed ; and it is only during the spawning-season that the body
of the male becomes thicker and more plump.
A specimen of medium size out of the spawning-season has a
total length of 40 millims., of which the head is 9 millims. ; and this
is contained about 4°4 in the total length (the caudal fin included).
The greatest height is 5:5 millims., or contained 7:2 in the total.
The vent is situated about 2 orbital diameters nearer the snout than
the root of the caudal.
In the spawning-season the body becomes fully developed,
although the total length does not increase. In a male with a
total length of 40 millims. the length of the head is 9°5 millims.
(4:2 in the total) ; in the female about the same as before.
The eyes have a diameter of 2 millims., and occupy 2 of the
length of the head. The orbital space, which, as a rule, is narrow,
and does not reach the breadth of an orbital diameter, has in the
mature male fully the last-named breadth.
The number of the gill-rays is 5; the gill-openings are wide, and
stretch forwards to about the middle of the orbit.
The mouth is oblique, and the lower jaw longer than the upper,
the maxillary extending to below the middle of the orbit. During
the spawning-season the jaws of the male become considerably
larger and more oblique ; at the same time, the head and all parts
of the body become swollen, the jaws extending much deeper under
the eye.
The number of the vertebrze is 27 ; the processus spinosi are very
long, and reach almost to the dorsal and ventral line. ‘The medulla
spinalis extends as far back as to the middle of the 22nd vertebra,
The teeth—Before the spawning-season the structure of the teeth
is exactly alike in both sexes. In each of the jaws is found on
both sides a single series of 20-25 minute and very closely fitted
1878.] MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 325
teeth, which project very little beyond the jaws. Only the inner-
most part of the lower jaw is entirely toothless. These teeth make
their appearance very early in the young. In a young specimen
of scarcely 14 millims. length, some very minute (microscopical )
teeth are already visible in the front part of the lower jaw, although
they cannot with certainty be seen in the upper. In specimens
of the total length of 23 millims. there are some small teeth visible
in both jaws ; and on being dried, one may see that some of them are
already developed; still they continue to be minute until the speci-
mens have reached their full length.
The teeth of the female undergo no special alteration the whole
of their life through, but continue daring the spawning-season in the
same state as before; but in the males, on the other hand, these
minute teeth of young specimens begin to disappear towards the
spawning-season, and are replaced by a set of long and cylindrical
ones. ‘These long teeth do not proceed from the minute teeth, but
constitute an entirely new set, which shoot forth at the same time
as the orignal set disappears. In the specimens wherein the organs
of generation are still not fully mature, both sets of teeth are visible
for a short period at the same time, the new set only half deve-
loped.
Finally, during the spawning-season all the minute teeth of the
male have almost entirely disappeared, and are replaced by the
new, long, and almost straight teeth, situated at a remarkable dis-
tance from each other, and chiefly to be found on the front part
of the jaws. In the lower jaw their number is 4, or sometimes but
3, on each side; they are not particularly pointed, but almost cylin-
drical, only slightly curved, and placed almost vertically upon the
outer edge of the jaw. At a little distance below and inside the
last of these teeth, on each side, is found a sharp, curved tooth
(canine) directed backwards and almost in the form of a claw. In
the posterior half of the jaw the prolonged teeth are missing, a few
of the minute teeth of the young period still remaining here.
In the upper jaw the number of the long teeth is 5 on each side,
of which a few (1-2) are also to be found behind the middle of the
jaw. In the upper jaw there is also found, inside the ordinary row
on each side, the curved canine, which is directed backwards; this
canine is placed further in front than the opposite one in the lower,
or nearly at the point of the jaw. There may further be found in
the posterior part more or less remaining of the original minute teeth,
whilst these only exceptionally are found at the point of the jaw.
The fins.—The first dorsal fin has 5 rays, and is a little higher
in the full-grown males than in the females, without, however,
reaching the height of the body. . All the rays are about evenly
high, undivided, feeble, and rather closely set. This is one of those
fins which appear latest, and is entirely wanting in specimens of a
total length under 16 millims., wherein all the remaining fins have
already got their rays. In individuals of the total length of about
22 millims, this fin is found with extremely feeble and short spines,
which are often difficult to detect.
326 MR. R. COLLETT ON CERTAIN GOBIOID FISHES. [Mar. 5,
The distance between the two dorsals is at least so great that
the rays of the first can never reach to the commencement of the
second.
The second dorsal fin has 13, rarely only 12 rays, the last of
which is a double one, although not divided in many young indivi-
duals. In the full-grown female, as well as in immature individuals
of both sexes, it strongly decreases posteriorly: the first is only half
as long as the second and third, which are the longest ; after this the
length diminishes successively, and the last (the double one) has
scarcely the length of the first. The first ray is not articulated ; the
remaining are all articulated; the first and the second are single, the
remaining divided at the point (the double ray only in its innermost
branch). In the mature male the posterior rays are lengthened,
the fin becoming almost even; the point of the posterior rays
reaches a distance of the diameter of the eye from the root of the
caudal fin, this distance being considerably larger in the females
and young individuals. The caudal fin has 17 articulated rays,
besides a row of 10 short, feeble, unarticulated, supplementary rays
on each side. All the articulated rays (with the exception of the
two outermost on each side) are twice divided in the outer two thirds
of the fin. The length of the fin does not exceed considerably the
height of the body. Its form in young specimens, as well as in
the female is always slightly emarginated, which is also the case
with a great number of the males during the spawning-season ; other
males have the caudal fin square, with rounded corners ; and it is
only in the case of the older that the middle rays are longer than
the other ones, by which means the form of the fin becomes
rounded.
The anal fin has 14 rays, the last of which is a double ray. In its
construction and position it agrees in general with the second dorsal ;
and having a ray more, it stretches a little further back than this.
The pectoral fin hes 16 articulated rays, all of which, with the
exception of the outermost on each side, are divided in the outer
third of the fin, and after the fourth ray doubly divided. The
form is somewhat pointed; the point is scarcely the distance of
one diameter of the eye from the vent. In accordance with the rule
in the generality of fishes, this fin is one of the latest developed ; in
a specimen with a total length of 19 millims., it consists only of a
rounded flap, in which there is a vestige of a few rays.
The ventral fins have their origin beneath the root of the pectorals,
and consist each of 6 rays. These together form the more or less
developed funnel, which is typical of the subfamily Gobiine. When
expanded, the opening of the funnel forms an oblong ring, which
runs parallel with the ventral line. All the rays are three times
divided down to the root, and are very movable. In the full-grown
male the extreme point reaches almost as far back as the pectorals.
In the female and the immature males the ventrals are very much
shorter, the funnels and the outer division of the rays indistinct ; the
oxen point is distant more than half the length of the head from
the vent.
1878.] MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 327
The mucous glands’, the true nature of which has perhaps not
yet positively been proved, are to be found on all parts of the head
in longer or shorter rows, which vary comparatively little in the
different specimens. To give a more detailed description of them
would require too much space ; but a drawing (see fig. 2, p. 323) has
been taken from a comparison of several specimens. Two parallel
series across the hinder part of the interorbital space are easy to be
observed, as may also two others, running along the upper side of the
snout, and sending forth some shorter series down to the nostrils.
The rest of these glands reach as far back as to the first dorsal fin ;
and shorter rows are found immediately behind the veutrals.
The scales and the skin.—The whole body, with the exception of
the head and fins, as well as the neck, throat, and belly, down to the
vent, is covered with large and very easily deciduous scales. These are
fastened to transverse muscular impressions, which bend together in
the lateral line of the body, where the vertebrae are connected. The
scales are arranged along the sides of the body thus:—First a
series of 25 scales are fixed along the lateral line; above and
below this row run two other rows; so that together there are 5 rows
between the dorsal and ventral line. The scales are perfectly trans-
parent, membranous and cycloid ; in the lateral line they are a trifle
smaller than in the other rows ; they are comparatively large, having
a diameter of nearly 1 millim., and are few in number. Accord-
ing to the above statement the total number of scales would be hardly
more than 100 on each side of the body. The period in which the
scales are developed I have not been able to observe to a certainty, as
the scales in young specimens seem to be still more easily deciduous
than in the older. In the smallest specimen I have found, with
a total length of 13 millims., the transverse impressions, to which the
scales are fastened, are already visible ; but I have not detected the
scales themselves before the specimens have attained about half their
size.
The colour.—When alive the whole body is almost perfectly trans-
parent, having only a few minute stellulate pricks inthe skin. One
may without difficulty count the vertebree from the outside, and
clearly distinguish the brain with its different parts, as well as the
pulsation and the blood-corpuscles. The most coloured part of the
body is the eye, and the innermost part of the black bladder. When
preserved in spirits, or in a frozen state, they soon become white and
opaque, whilst if preserved in glycerine they remain half transparent ;
then the minute pricks become fainter, whilst the iris always retains
its silvery colour.
In the breeding-season the minute pricks become more numerous,
especially in the males. A row of these pricks, yellowish red in
colour, stretches along the sides, corresponding in number to the
vertebree. Along the dorsal line another row of brown pricks is
found. Along the anal fin, on each side of the root of the rays, is a
row of black pricks, which continues behind the end of the fin in
' “Stripes of minute warts,” Giinther, Catal, vol. iii. 1861 ; “ Bxternal
papille of taste,” Winther, Naturh. Tidsskr, 3. R., 9. B., p. 181, Kbhyn. 1874.
328 MR. R. COLLETT ON CERTAIN GOBIOID FISHES. [ Mar. 5,
the shape of a short black stripe ; this is the most conspicuous and
durable of all the rows, and is never wanting. The caudal fin has
a brownish shade; sometimes this is also the case with the anal fin,
whilst all the other fins are hyaline. The iris is silver-coloured,
with black pupil; the upper margin of the eye is golden, which,
after death, changes to a black. ‘The younger individuals are more
sparingly pricked; the row along the anal fin is the first which
shows itself, and is already visible in the young ones, with a total
length of 13 millims., where all the other pricks are wanting.
In the males in full breeding almost every part of the body has a
faint brownish light, although this is nowhere collected into larger
dots, but is most distinct along the transverse impressions of the
muscles.
C. Habits and Distribution.
Generation.—It is probable that no specimen lives more than one
year, and after the close of the breeding-time dies without living
through another spawning; consequently these fishes are really
annual vertebrates.
The spawning-season occurs in the Christiania Fjord in the summer
months. The latest date at which I have found females with
ripe spawn was on August 2, 1872, although the greater number of
them had then already spawned. The normal period of spawning
is the end of June or the beginning of July. On the 10th and 14th
of August, 1875, I found them in the middle of their breeding,
although several were still not quite mature. When the spawning is
finished all the specimens disappear entirely, and become apparently
extinct.
The first young ones I took on August 28, 1875, when their
average size was 22 millims.; a few specimens were rather larger
(attaining about 30 millims.), or less (down to 13 millims.). At the
same time there is not a single specimen of a greater size to be found
at those places where they, in the middle of June, were found in
numbers and only in a full-grown state.
A month later the median length may be estimated at 34 millims.,
in the end of October at 38 millims. The development proceeds
tolerably even with them all; and specimens are very rarely to be
found which differ to any degree from the medium size. Most of the
individuals attain their full size as early as December or January, in
order that, when the organs of generation begin to develop themselves
in the following spring, they may acquire greater plumpness and
strength in all parts of the body—a change which, as we have indicated
above, chiefly takes place in the males. Amongst the thousands of
specimens which I have examined during the autumn and winter,
I have not found a single specimen with the slightest trace of the
long teeth that. characterize the mature male; and during the summer
time all the males, without exception obtain these teeth.
In the young females, at the age of only four months, the ovaries
may be seen through the transparent body in the form of two elon-
gated strings, filled with clear eggs; in the spawning-season these
1878.] MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 329
strings take up the greater part of the belly, without, however, as is
the case in Crystallogobius nilssonii, reaching behind the vent along
the root of the caudal fin. ‘The number of the eggs is between 1800
and 2000.
Habits.—This species, as well as the following differs in its habits
in several respects from most of the typical Gobiide. Above the
compressed body the laterally placed eyes indicate that they do not
live at the bottom, as is the case with the greater number of
Gobies, but swim about freely. Neither do they keep themselves
dispersed or in single numbers, but in large and dense shoals,
where their perfect transparency, in connexion with their. inferior
size, enables them to avoid immediate observation. Sometimes I have
taken them from the stomachs of other fishes; but by far the
greatest number are brought up to the surface by the nets set out
for different other fishes, especially for the sprat and the young of
the common herring. These fishes are caught in the Christiania
Fjord in September and so long as the fjord is free from ice—as a
rule, until the beginning of the new year. In the spring and summer,
when the nets are drawn for mackerel, they are also to be found,
and then with the organs of generation fully developed. These nets
are drawn at a depth varying from one to fifteen fathoms. An
examination of the stomachs of these fishes shows that they contain
chiefly pelagic copepods and the fry of mollusks in their swimming-
stages; these animals principally live in the upper strata of the sea
some few fathoms under the surface. The bottom consists, in most
of the inner parts of the Christiania Fjord, of clay and mud, which,
in the more shallow water, is covered with Zostera.
These fishes seem always to keep together in enormously large
shoals ; and when the meshes are very fine they are sometimes caught
in great masses. The 30th of October 1875 (which was a par-
ticularly successful year for their development), I found in a single
draught (which, besides, also brought in some young specimens of
two or three species of Gadus, several Pleuronectes flesus, and Cteno-
labrus rupestris, some Gobii of common species, and herrings) such
large masses of them that I collected between two and three thousand
specimens, whilst possibly more than double this number had already
escaped through the meshes. In the same year I was not able to
notice any diminution of their numbers, and they were always found
in the same places; still they are not at all equally numerous every
year, and they have been rather scarce both in 1876 and 1877, es-
pecially in the latter.
In the Mediterranean they must be produced in still greater
numbers, as they are caught in very fine nets and sold as food.
In several of the fishing-markets in the towns by the Adriatic and in
Sicily Prof. G. O. Sars saw vessels filled to the brim with these
fishes alone, forming a complete jelly-like mass. Kessler also men-
tions their use as food in Odessa.
Upon the least touch the scales fall off; and after the nets are
drawn, scarcely a single specimen will have its scales complete, and the
greater number will have lost almost every scale. The proportionally
330 MR. R. COLLETT ON CERTAIN GOBIOID FisHES. [ Mar. 5,
large swimming-bladder, which occupies a considerable part of the
belly, expands still further when the individuals are brought up
to the surface, the somewhat fatigued fish not being able to com-
press it so as to make the specific weight of the body to decrease
sufficiently to permit it again to descend under the water. The
weight of the body is also diminished by the loss of the scales;
consequently a great number of such individuals are driven about
on the surface of the water in a still living state, till they are
carried away by the stream or picked up by sea-birds and other
enemies, They swim, however, not without a certain degree of
rapidity.
Their enemies are almost all other fishes who occupy the same
locality. Especially the different kinds of Gadus devour large masses ;
and even in the case of small specimens of Gadus morrhua, G.
esmarkii, and G. merlangus, I have occasionally found the stomach
crammed with them. When they float about in their helpless state
on the surface of the water, Gasterosteus aculeatus and Spinachia
vulgaris snatch large pieces out of their brittle bodies.
As mentioned above, their food consists chiefly of microscopic
copepoda and other small pelagic animals. In the specimens that I
have examined in the Christiania Fjord, these principally consisted of
the copepods Dias longiremis, Lilljeb., Temora velox, Baird, together
with the larvee of decapods (Hippolyte and Palemon), and mollusks
in their swimming-stage (Cardium). The stomach contains this
kind of food both in the summer and the winter.
Distribution.—Taking for granted, that the forms from Southern,
Western, and Northern Europe are identical, L. pe/lucidus has a very
wide distribution, although our information touching this species is
at present very incomplete. In this case it is met with (possibly in
certain spots only) from the western coast of Norway, about 60° N.
long., down along Western Europe to the Mediterranean, in the
Adriatic, and right up to the innermost parts of the Black Sea.
In Norway it is met with outside of Bergen, and in the Christi-
ania Fjord. At the first-named place the late naturalist Mr. Stuwitz
obtained four specimens in December 1834; it has, however, not since
been found in this locality. In the Christiania Fjord I have found
it in several places inside Doobak, chiefly amongst the islands outside
Christiania; most likely itis to be found in the outer parts of the
fjord also, and undoubtedly at other places along our southern coast,
not mentioned here.
In Sweden it has been found by Dr. Malm in Bohuslen.
In Scotland Mr. Parnell has taken it in Solway Frith, June 1836 ;
and in his ‘ Catalogue’ Dr. Giinther states that the British Museum
also has specimens from the Frith of Forth.
In the Mediterranean it is known, through Prof. Canestrini, from
the Gulf of Genoa; and Prof. G. O. Sars found it in the beginning
of 1876 at most of the places visited by him—e. g. in Sicily, at
Messina, and Syracuse—everywhere in enormous masses. In the
Adriatic, whence it was already described in 1824 by Nardo, and
afterwards mentioned by several anthors, he found it at Trieste.
1878.] | MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 331
In the Black Sea it was taken in the harbour of Odessa by
Kessler in 1859.
Genus CrysTaLLocosivs, Gill.
Crystallogobius, Gill, Proc. Acad. Nat. Sci. Philad. 1863, p- 269
(1863).
Body transparent, elongate, strongly compressed, naked, with
parallel muscular impressions. The male only dentiferous, the teeth
in a single series; strong canine teeth in the lower jaw. Two
separate dorsal fins, the first composed of two spines; ventral fins
united, gill openings wide ; pseudobranchiz present.
Fig. 3.
Mature male of Crystallogobius nilssoni, in the breeding-season 4.
C. nizssont (Dib. & Kor.).
1844. Gobius nilssonii, Diib. & Kor. Kgl. Vet. Akad. Handl.
1844, p. 53, tab. 2. fig. 3 (Stockh. 1846).
1861. Gobiosoma nilssonii, Giinth. Cat. Acanth. Fish. Brit. Mus.
vol. iii. p. 86 (Lond. 1861).
1863. Crystallogobius nilssonii, Gill, Proc. Acad. Nat. Sci. Philad.
1863, p. 269 (Philad. 1864).
1872. Latrunculus nilssonii, Coll. Forh. Vid. Selsk. Christ. 1872
p- 10 (Christiania, 1873).
1874. Latrunculodes nilssonit, Coll. Forh. Vid. Selsk. Christ. 1874,
p- 151 (1875).
The intermaxillary bones in the male very prominent, short;
small teeth in front of both jaws, long canine teeth in the lower
jaw. The ventral fins united into a funnel, with the shorter rays
towards the body. The first dorsal and the ventral fins in the
female rudimentary; the ovaries project far behind the vent.
Number of muscular impressions 27. The lower jaw the longer.
D’ 2, D? 19 (20); C. 8 (9 or 10) + 15 (14) + 8 (gori10); A. 21;
V.6; P.30 (31). Lin. lat. 27. Vertebr. 29. M.B. 5.
>
A. Remarks on the Synonymy.
This species was described by von Diiben and Koren in the Vet.
Akad. Handl. for 1844, under the name of Godius nilssonii, the de-
scription having been taken from half a dozen specimens found off
the Bergen coast in 1834—43, although it had earlier, during the same
332 MR. R. COLLETT ON CERTAIN GOBIOID FISHES. [ Mar. 5,
year, been mentioned, but without any accurate description, and
under the temporary name of Godius linearis, by the same author
in a letter which was published in (fv. Kgl. Vet. Akad. Forh.
May 1844. In Giinther’s ‘ Catalogue,’ vol. ii. (1861) it is placed
under the genus Gobdiosoma, although the correctness of this is
doubted. As early as in 1863 Prof. Gill (Proc. Acad. Nat. Sci.
Philad.) established for this species a new generic name, Crystallo-
gobius. Synonymous herewith is the name Latrunculodes, by which,
in 1874, not knowing the remarks of Prof. Gill, I entered it in a
treatise on the Gobioid Fishes of Norway (Forh. Vid. Selsk. Christ.
1874), and later in the same year in apaper on the Fishes of Norway
(Supplementary part of the Forh. Vid. Selsk. Christ.).
Since the species was entered in the system in 1874, it has not
been the subject of detailed description. Nilsson, in his ‘ Skan-
dinavisk Fauna’ (1855), has taken his description from that of
Diiben and Koren, as is also the case with the few other authors
who mention the species.
The same applies to this species as to Latrunculus pellucidus, that
the characters mentioned in the descriptions only belong to one of
the sexes, in Crystallogobius nilssonii to the fully developed male.
The female of this species, which shows a constant and remarkable
difference from the male, has not hitherto been described.
In June 1875 I found the species in the Christiania Fjord in mature
state, together with Latrunculus pellucidus, though in very much
smaller numbers ; and more lately I have on a few occasions collected
young specimens out of the spawning-season. Although the number
of specimens which at present stand at my disposal, namely 33, do
not represent an unbroken series of all the different ages of both
sexes, I can still give a descriptive account of their principal stages.
B. General Description.
The mature males have a total length of between 43 and 47
milllims.; still maturity is evidently attained in some specimens
with a length of 38-40 millims. The first dorsal fin has long rays
with a wide membrane ; the head is thickened, with a short snout ;
the jaws long, and singularly bent towards each other; the canine
teeth in the lower jaw high and curved; and the intermaxillary
bones form on each side a strong projection, almost in the form of
a nail. A little more pointed is the head, and the body on the
whole is more slender, in the males which have not reached a total
length of 70 millims; neither have the jaws nor the teeth in this
case attained their full length, though the testicles are filled with
ripe milt.
The young males, whose testicles do not yet contain ripe milt,
although their total length is only a trifle less than the smallest of
the mature specimens, namely 36 millims., have the body thin and
slender, the head pointed, almost as in the females; the teeth,
however, are already visible, and all the fins normally developed.
The mature females, with the eggs ready for spawning, are con-
siderably smaller, and of a more slender construction, than the
1878.] | MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 333
males. Although the total length is almost the same as in many of
the mature males (36-37 millims., in a single specimen 39 millims. ),
their body is so slender and their weight so inconsiderable, that two
full-grown females, with the ovaries filled with ripe eggs, were not
able to weigh down one large full-grown male. The head is
pointed, with short jaws, which are straight and perfectly toothless,
and are wanting the males’ prominent projection on the intermaxillary
bone. The first dorsal and the ventral fins are rudimentary (at a
superficial glance they appear to be entirely wanting); the pectoral
fins are certainly developed, but very short. The ovaries project far
behind the vent on each side of the anal fin.
The young females, with a total length of 30-33 millims., and
with the ovaries filled with unripe eggs, have the body very slender
and pointed: as in the old female the jaws are toothless, the first
dorsal and the ventral fins apparently wanting.
The young during growth.—The few young with a total length
of about 25 millims. that have been at my disposal have all been
males. This is shown by the first traces of teeth, by the want
of the long ovaries, as well as by the presence of all the fins; cer-
tainly the first dorsal and the ventrals are short, but never wanting
or even rudimentary. These fins may already be seen in the young
with a total length of 21 millims., the smallest specimen I have
found ; in this one, however, there was not yet any trace of the
pectorals.
It may therefore be supposed that in the males the first dorsal
and the ventral fins spring forth ata very early age, whilst the
females never have these fins developed ; the pectorals, on the other
hand, are present in both sexes (although in the female in a less-
developed state), but are always late in appearance.
As compared with Latr. pellucidus, the difference in the exterior
between the sexes in the present species appears in several ways
already at a very early period, whilst this difference in Latr. pellu-
cidus, as stated above, only becomes apparent in mature specimens.
The construction of the body is thin and elongated, very com-
pressed, and does not, as is the case in Latr. pellucidus, practically
increase in plumpness in the spawning-season.
In a full-grown specimen, ready for spawning, with a total length
of 46 millims. (caudal included), the length of the head is about
10 millims, and is contained 43 times in the total length. The
greatest height (53 millims.) continues almost unaltered from the
occiput to about a head’s length from the caudal fin, and is con-
tained 82 times in the total. The vent, on the posterior margin of
which there is a small papilla, is situated halfway between the snout
and the root of the caudal, and in some individuals projects in the
form of a short tube.
In the female, which, as stated above, is considerably smaller
than the male, the head is somewhat more pointed. Ina specimen
with the eggs ripe the head is contained about 5 times or a little
more in the total length.
The diameter of the eye is one fourth of the length of the head.
334 MR. R. COLLETT ON CERTAIN GOBIOID FISHES. [ Mar. 5,
The eyes are situated perfectly laterally, so that the lower margins
do not seem to project more than the upper. ‘The interorbital space
is, in the female and young specimens, not broader than*the diameter
of the eye, in the full-grown male a little larger. The branchio-
stegals are 7 in number ; the gill-openings are wide.
The mouth is oblique, with the lower jaw the longer. In the
female and the young males the lower jaw is almost straight, or
only faintly bent, and the intermaxillary bones are quite even in
their outer margin; but in the full-grown male the jaws receive a
special development during the spawning-season. The inter-
maxillary bones then become broad, and form on each side of their
margin a distinct projection, almost in the form of a nail; at
the same time that the teeth reach their full development the
lower jaw becomes long and strongly curved upwards, and when the
mouth is shut it reaches a little in front of the intermaxillary bones,
the projecting nail of which encloses the almost angularly curved
upper part of the lower jaw. The point of the latter jaw is square,
and, as is the case with the intermaxillary bones, in the male den-
tiferous. In these old males the gape reaches backwards to the
posterior margin of the eye, whilst in the female and the young
males it scarcely reaches beyond the middle of the eye. The length
of the lower jaw in proportion to the total length isin the old males
as 1:72, in the female as 1:10.
The number of vertebre is 29.
The teeth.—Whilst during its immature state Latr. pellucidus has
a construction of the teeth common to both sexes, which continues
permanent in the female, whilst the male towards the spawning-
season in a comparatively short time changes these teeth for another
set of a totally different construction, this is not the case with
Crystallogobius nilssonii. First, the female is perfectly toothless
all her life; secondly, the teeth of the male, which are com-
posed of both small and acute teeth and long canine teeth, are
developed very early, and all reach their full size during the spawning-
season.
The exact time when the teeth make their appearance in the young
is probably when they are ‘about two months old. A young one,
with a total length of 21 millims., and perhaps about one month and
a half old, had no traces of teeth.
In the young males, which have reached a total length of 26-28
millims., the teeth begin to show themselves in the intermaxillary
bones and the point of the lower jaw; at this period all the teeth
are about equal in size.
In the male during the spawning-season, when the teeth have
reached their full size, the projecting intermaxillary bones have on
their upper and prominent portion a single row of closely placed,
small, acute, and curved teeth, 13-14 in number, on each side
(see fig. 4); on the lower square margin one or two teeth directed
straight downwards or backwards are found. The maxillary bones
are without teeth, as well as the vomer. In old males the outer
part of the lower jaw runs out in a long upward curve, which at
1878.) MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 335
its point is dentigerous, while otherwise the jaw has no teeth;
the innermost part of this jaw forms a sharp edge, which, as the
jaws meet one another, places itself inside the corresponding part
of the upper jaw, whilst the point of the lower jaw, as men-
tioned above, is encompassed by the outward-bent intermaxillary
bones. The point of the lower jaw is square and broad, and has a
remarkable similarity to the jaw of a carnivorous animal. On
each side of the square point is found a long and curved canine
tooth, and between them a row of, together, eight small and acute
teeth of the same construction as those in the intermaxillary bones,
Fig. 4.
Snout of mature male of Crystallogobius nillsoni in the breeding-season,
strongly magnified.
The female must be pronounced toothless, as such teeth as can
be found are partly irregular and partly perfectly microscopical.
In a fresh or soft specimen it is scarcely possible to find even a
trace of teeth ; but in a dried specimen a couple of very minute and
straight isolated teeth may be seen, with the aid of the microscope,
on the middle of the lower jaw ; these, however, are sometimes en-
tirely wanting on one side. In a full-grown female the length of
these teeth is about 54; to +4 of a millim.
The fins.—The first dorsal fin, which has two rays, becomes
only fully developed in the mature or the nearly full-grown male,
but is much shorter in the young (not half-grown) male, and
rudimentary in the female. In the males it has strong but not stiff
rays, which are placed at comparatively large intervals, and con-
siderably sloped backwards, and have the point almost bent down-
wards, the membrane, which begins already a little in front of the
first, continuing behind the last, and running along the dorsal line
towards the beginning of the second dorsal fin; in some specimens
it even connects both fins completely. The first ray is a little
shorter than the second, the length of which does not much exceed
the greatest height of the body. In the young males the rays are
shorter, and the membrane between them incomplete. In the female
they are shortened to such an extent that they can only be discovered
by very minute examination ; sometimes only one ray seems to be
developed. In the female the length of the rays seems never to
exceed the length of the eye, and is generally shorter, sometimes
having only the length of the crystalline lens.
336 MR. R. COLLETT ON CERTAIN GOBIOID FisHEs, [Mar. 5,
The second dorsal has nineteen, sometimes twenty rays, the last
of which is a double ray. By means of a strong membrane the rays
are directed strongly backwards, and appear, therefore, lower than
they really are; they are all even, and have a length equal to half
the height of the body. The posterior rays are not elongated even
in the male, and reach to an orbital diameter’s distance from the
root of the caudal fin. All the rays are indistinctly articulated, and
undivided ; and it is only by the aid of the microscope that in some
individuals a trace may be seen of an indistinct division at the point.
Still shorter and more delicate are the rays of the female, their length
being scarcely equal to an orbital diameter.
The caudal fin has fifteen articulated middle rays, sometimes only
fourteen; also on each side eight, sometimes nine or ten short con-
necting rays, which are widely divergent in their points, especially the
foremost, so that they can even point forwards, giving the root of
the fin an increased breadth. The middle rays are articulated,
and all of them divided from about the middle (except the two
outermost on each side); bnt neither of the branches is further
divided. The first ray is short and thin, and the articulation is not
very plain ; the fourth and fifth rays are the longest, while the middle
rays again become shorter. One consequence of this is, that the
fin becomes more or less emarginate, with rounded edges; it is
rather short, and scarcely attains the greatest height of the body.
The anal fin counts twenty-one rays, the hinder of which is a
double ray. It takes up nearly the whole space between the vent
and the caudal, and agrees very much in its form and construction
with the second dorsal. It is only by very favourable light that any
trace of articulation in the points of the rays can be discovered, and
not in all of them. The last ray reaches almost to the root of the
caudal; in some undamaged individuals it is even connected with
the last-named fin by a membrane, whilst this membrane is wanting
in others.
The pectoral fin counts thirty, sometimes thirty-one rays. Twelve
to fourteen of the middle rays are singly divided from about the
middle; the remainder are undivided. All are finely articulated.
The form of the fin is broadly rounded. The pectoral fins are not
very long ; in mature males they have about two thirds of the head’s
length, and do not reach to the vent. In young males and‘in the
female they are much shorter, although they are never rudimen-
tary.
The ventral fins, which are attached close to the root of the pec-
toral fins, and just behind the lower end of the gill-openings, are
well developed in the mature male, shorter in the younger male, and
rudimentary in the female.
When present they form a closed and deep funnel, which is square-
cut behind, and, contrary to what is found in the other Gobiide,
have their innermost rays (nearest to the belly) shorter than the
outer ones. The number of the rays is six in each fin: of these the
first is divided from about the middle; the remainder are more
slender, undivided, and not articulated. On the whole, the faint
{878.] MR. R. COLLETT ON CERTAIN GOBIOID FISHES. 337
tendency to division of the rays is characteristic of this species as
compared with Latrunculus pellucidus, in which division of the rays
takes place especially in the ventral fins. The funnel is comparatively
short; when closed it is, even in the male, removed from the vent
to a distance about equal to its own length. In young males the
ventral fins are very small, and their funnel-like shape is almost
imperceptible, the connecting membrane being incomplete.
In the female the ventral fins are completely rudimentary, being
only present as a couple of two disconnected and very short rays
scarcely the diameter of the eye in length.
The mucous glands, which in L. pellucidus form so extensive a
system, seem to be almost entirely wanting in Crystallogobius nils-
sonii ; in none of the specimens which I have examined have I
been able, with certainty, to find a single complete row of such
glands, either on the head or on any other part of the body.
The skin—No trace of scales has hitherto been detected in
any of the specimens examined. As is the case in L. pellucidus,
the whole side of the body shows a row of transverse impressions ;
when examined in the microscope, no trace is found of follicles to
which lost scales may have been attached; and it is therefore
searcely to be doubted that the body is altogether naked in this
species. The number of these muscular impressions is twenty-seven,
twenty of which have their place behind the vent; they bend to-
gether into the form of an angle in the middle line of the body, and
denote exactly the number and size of the vertebree. An actual
lateral line, with open pores, does not exist.
The colour.—As is the case in L. pellucidus, the body during
life is transparent, with a faint bluish tinge, thereby enabling the
vertebree, the apple‘of the eye, &c. easily to be seen. The colora-
tion of the skin is very faint, and does not vary in this respect to
any degree, even during the spawning-season. The eyes are strongly
coloured, their upper edge being of a shining metallic lustre (black
after death) ; the iris is silvery. A well-defined blackish spot is
found at the point of the lower jaw; another black stripe stretches
along the throat towards the ventral fins. At the root of each
ray in the anal and the caudal fin there is a black point; these
together form a continuous and distinct row, which is present in
both sexes and at all ages; along the dorsal fin there is found a
somewhat fainter row of brownish points, which is indistinct, or
nearly wantivg, in the female. Moreover all the rays are covered
with minute blackish points. The body itself is almost unspotted
(whitish when preserved in spirits), except on the sides of the belly,
where a row of fine brownish spots stretches itself; these spots are
largest and most distinct in the breeding female, their number being
3-5 on each side.
C. Habits and Distribution.
Generation.—With regard to the time of breeding and its dura-
tion, nothing conclusive can be said at present on account of the very
Proc. Zoou. Soc.—1878, No. XXII. 22
338 MR. R. COLLETT ON CERTAIN GOBIOID FISHES. [Mar. 5,
few materials at my disposal. In all probability Cryst. nilssonii reaches
(as is the case with Latr. pellucidus) maturity at the age of one
year, after which it disappears. ‘The spawning-season is probably
the same as in the other species, consequently during the summer
months; on the whole, however, it appears to have a somewhat
longer duration.
From the researches I have been able to make among col-
lected specimens, compared with those preserved in the Museum
of the University, it appears that the spawning probably begins in
some individuals as early asin March or April ; perhaps the greater
number attain their maturity in the last part of June; whilst in other
specimens the spawning does not take place before the beginning of
August, or perhaps later. After the conclusion of the spawning the
specimens no doubt perish, their life having thus lasted only one
year.
" Habits.—In its habits this fish probably agrees perfectly with
Latr, pellucidus. The extraordinarily compressed body denotes that
it swims freely about in rather deep water, and not, like most of the
true Gobii, resting itself at the bottom; in fact this would be almost
impossible, on account of the peculiar construction of the ventrals of
the male, and, in the case of the female, from its entire want of these
fins. Most of the specimens mentioned by other authors have been
taken in the dredge at a depth of 30 fathoms. Von Diiben and
Koren state that some of their specimens crept into empty tubes of
Chetopterus norvegicus, in which, however, they had only taken
refuge whilst the dredging was going on.
In the Christiania Fjord I have taken these fishes, together with
Latr. pellucidus in nets set out for mackerel, Clupea sprattus, and
young of Cl. harengus. The nets bring them to the surface in the
same manner as is the case with L. pel/ucidus. As a rule they lose
the capability of again descending, from the swimming-bladder
becoming strongly distended when they reach the upper strata of the
water. This fish is met with in far less numbers than the last-named
species. In the same manner it is taken as food by other fishes;
and I have taken it from the stomach of small young examples of
Gadus morrhua.
Food.—Upon examining a number of stomachs of individuals
taken in the month of June, I have found them filled with digested
remains of Entomostraca, without, however, being able to recognize
any species of them. Probably they were Calanidz, or, perhaps,
pelagic Copepoda of the same species as are found in Laér. pellu-
cidus.
Distribution.—Besides being found in Norway, Cr. nilssonii has
been taken more lately on the coast of Bohuslen (Sweden) ; and
perhaps it also occurs on other parts of the coast of Northern Europe.
In Norway its habitat extends from the Christiania Fjord along
the western coast, nearly to the Trondhjems Fjord, in about 63° N.
latitude.
The most northern point at which it has been found was at Chris-
tiansund, in July 1843, by v. Diiben, and also about the same
1878.] ON THE ORNITHOLOGY OF THE PHILIPPINES. 339
time at Askwold, in Sond-fjord, by Mr. Christie—in all only six
specimens.
In the neighbourhood of Bergen five specimens were taken
by Mr. Stuwitz, December 1834; in 1850 one specimen was found
at Bongestrémmen, near the town: in all six specimens.
One specimen was dredged by Prof. G. O. Sars at Espevar, in the
mouth of the Hardanger Fjord in August 1873.
In the mouth of the Christiania Fjord two specimens were dredged
in two different localities about 1860; finally, in the years 1875-77
I took twenty-eight specimens in nets between the islands off
Christiania.
In Sweden two specimens were found (in a dead state) in Bohuslen
by Dr. Malm, in June 1877,
March 19, 1878.
Arthur Grote, Esq., V.P., in the Chair.
Mr. Sclater exhibited the type specimen of Dicrurus marginatus
of Blyth (Ibis, 1865, p. 46), belonging to the Derby Museum,
Liverpool, and pointed out its identity with Muscipipra vetula
(Licht.) of Southern Brazil, of the family Tyrannide.
The following papers were read :—
1. Contributions to the Ornithology of the Philippines.—No.
VI. On the Collection made by Mr. A. H. Everett in
the Island of Leyte. By Artur, Marquis of Twrxp-
DALE, F.R.S., President of the Society,
[Received February 25, 1878.]
The island of Leyte is about 130 miles long, and has an average
breadth of 35 miles. It lies S.S.W. of Samar, its northern part
lying in close contiguity to Samar. Zebu is situated to the west,
and Dinagat to the south-east, while Mindanao is due south. Water-
channels of unimportant breadth separate Leyte from these other
islands. A peculiar ornis is therefore not to be expected, and Mr.
Everett’s researches have not made known any special feature. That
gentleman, however, procured all the birds of which I propose to
give a list at Amparo (on the south-west coast of Leyte) and in its
vicinity. But Mr. Everett writes that he does not believe the collec-
tion he sends at all adequately represents the avifauna of Leyte,
and he hopes to revisit it and explore the mountains in the centre
and south of the island.
The most interesting species is Buceros semigaleatus, a purely local
form, distinguishable from the two previously known Philippine
22%
~
340 THE MARQUIS OF TWEEDDALE ON [Mar. 19,
species by the contour of the casque. Thriponax pectoralis is
another species, the deviation of which from 7’. javensis of Luzon is
probably owing to isolation. In Arachnothera flammifera Mr.
Everett has added another Malayan genus to the Philippine area.
1. CACATUA H&MATUROPYGIA (1).
{[Amparo, ¢ 2, July. ]
2. TANYGNATHUS LUZONENSIS (3).
[S. Leyte, ¢, September. ]
3. LoricuLUS HARTLAUBI (7)?
[Amparo, 53 2, July. ]
I provisionally identify a series of nine examples of the South-Leyte
Loriculus with L. hartlaubi until further research establishes the
permanency of the characters whereby these Leyte birds differ from
those of North Mindanao and Dinagat. Five individuals are exam-
ples of the male bird in apparently adult plumage. They differ from
true L. hartlaubi in having a much smaller orange-red pectoral
plastron, and in its not running up to the naked chin, but stopping
short some way below, so that the upper throat is uniform green.
Above, the back is green, and not golden, nor is that of the female,
The dimensions are equal.
4. AsTuR TRIVIRGATUs (11).
[ Amparo, ?, July. |
In Mr. J. H. Gurney’s opinion this Leyte example is in immature
plumage, and probably belongs to a male.
5. SPILORNIS HOLOSPILUS (16).
[S. Leyte, ¢ 2, September. |
6. Haiastur INTERMEDIUS (17).
[S. Leyte, ¢ 2, September. ]
7. THRIPONAX PECTORALIS, 0. Sp.
[S. Leyte, ¢ 2, September. ]
Like 7. javensis, but differs in having the feathers of the chin,
throat, sides of head and neck white, broadly centred with a median
black stripe, and in the fulvous-white colouring of the abdomen
covering the breast, each pectoral plume being centred with black.
8. MeERops PHILIPPINUS (35).
[S. Leyte, ¢ 2, September. ]
9. MEROPsS BICOLOR (36).
[S. Leyte, ¢ 9, September. }
Two examples in mature plumage, two immature; the chestnut
head and back in the latter mixed with green, and the middle pair of
rectrices not fully developed.
1878. } THE ORNITHOLOGY OF THE PHILIPPINEs. 341
10. Eurystomus orIENTALIS (37).
[Amparo, ¢ 2, August. ]
11. ALCEDO BENGALENSIS (38).
[S. Leyte, g 2, October. ]
12. Ceyx aArRGENTATA.
Ceyx argentata, Tweeddale, Ann. & Mag. N. H. ser. 4, vol. xx.
p- 533.
[S. Leyte, g, October. |
13. Enromosra Guuaris (44).
[Amparo, 3, August ; S, Leyte, September. |
14. SavROPATIS CHLORIS (47).
[Amparo, ¢ 2, August. ]
15. PELARGOPSIS GIGANTEA.
Pelargopsis gigantea, Walden, P. Z. 8. 1877, p. 541).
[ Amparo, 5 °, July ; S. Leyte, ¢ 9 , September and October. ]
‘fen examples, all of which belong to the smaller North-Mindanao
and Dinagat race, which I have hitherto provisionally referred to
P. gigantea.
16. CentRococcyx viripis (64).
[S. Leyte, ¢ 2, October: iris brown. 6. 3, September: iris
yellowish brown; bill black ; legs steel-grey. |
17. Bucrros sEMIGALEATUS.
Buceros semigaleatus, Tweeddale, P. Z. S, 1878, p. 277.
18. PENELOPIDES AFFINIS,
Penelopides affinis, Tweeddale, Aun. & Mag. N. H. ser. 4, vol. xx.
p. 934; P.Z.S. 1877, pp. 824, 825 (fig. 1)",
[Amparo, g ?, July; 8. Leyte, ¢ 2, September and October, 2
juv.*: iris raw sienna-brown; bill greenish brown; mandible green-
ish; legs dark greenish grey. @. Iris in living bird rich dark
burnt-sienna-orange. ]
The new rectrices in both sexes of this species are pure white, and
not pale rufous, the apices always remaming black. In one ex-
ample (2) the two middle rectrices have a dark rufous terminal
band, black being indicated where this band meets the paler rufous
of the basal portion. Ina young male this also occurs,
19. Lantus nasutus (70).
[S. Leyte, ¢ 9, September and October.
* The passage in which I differentiate P. afinis from P. panini and P. ma-
nille (J. c.) should read ‘by the absence of the channelling on the lateral plates
of the maxilla.”
* Marked ¢ by collector, but in complete black plumage,
342 THE MARQUIS OF TWEEDDALE ON [ Mar. 19,
20. Lanius LUCIONENSIS (72).
[S. Leyte, ¢ 2, September. |
The males are in adult dress, with pearl-grey foreheads. The
female has all the gular and pectoral plumage narrowly edged with
brown, as well as the feathers of the flanks and thighs. The entire
upper plumage is uniform liver-brown, without a trace of grey on the
forehead.
21. ARTAMUS LEUCORYNUS (73).
[Amparo, g, August; S. Leyte, 2, October. |
22. LaLAGE DOMINICA (76).
[Amparo, ¢, July. |
23. CyORNIS PHILIPPINENSIS.
Cyornis philippinensis, Sharpe, Tr. L. 8. 2nd series, Zool. i. p. 325.
[Amparo, ¢, July.]
24. HrruNDO JAVANICA.
Hirundo javanica, Sparrman, Mus. Carls.
[S. Leyte, g, September, ¢ juv., October. ]
25. BRODERIPUS ACRORHYNCHUS (90).
[Amparo, 9, July ; S. Leyte, ¢ 2, September.]
26. Mrxornis CAPITALIS.
Mizxornis capitalis, Tweeddale, P. Z. 8. 1878, p. 110, pl. vii. fig. 2.
[S. Leyte, 2, October: iris white. ]
27. Ixus GOIAVIER (99).
[Amparo, 2, July; S. Leyte, ¢ 2, September. }
28. HypsipeTes PHILIPPINENSIS (102).
[S. Leyte, ¢, September. |
29. Monticoua souitarivs (103).
[S. Leyte, ¢ 2, October. |
Three examples are in dull plumage, the feathers, especially on
lower surface, broadly margined with brown ; above, a bluish tinge.
These are marked female by the collector. A fourth example (marked
2 ) is passing over from this stage, the blue tinge above being much
more marked, while most of the abdominal plumage and the under
tail-coverts are rich chestnut. A fifth example (marked g) is in
nearly full blue and dark rich chestnut plumage.
30. CopsycHUS MINDANENSIs (106).
[S. Leyte, ¢ 2, September. ]
31. PHYLLOSCOPUS BOREALIS.
Phyllopneuste borealis, Blasius, Naumannia, 1858, p. 313.
[S. Leyte, ¢, October. ]
1878.] THE ORNITHOLOGY OF THE PHILIPPINES, 343
32. CALOBATES MELANOPE (115).
[S. Leyte, °, September. }
33. MyzanTHe pygmma (121).
[S. Leyte, 3, October : iris chocolate. }
34. ARACHNOTHERA FLAMMIFERA, 0. sp.
[S. Leyte, 3, September : iris salmon-red ; bill black; legs dark
lead-grey.]
Above uniform rich olive-green ; chin, throat, sides of head, and
neck pale grey ; upper breast pale grey, tinged with yellow ; lower
breast a purer pale yellow; abdomen, flanks, ventral region, and
under tail-coverts deep yellow; Wing-lining silky pale grey, almost
white; pectoral tufts (which measure some three quarters of an
inch long) bright vermilion-red ; quills brown, margined with clive ;
rectrices brown, all but the middle pair boldly tipped on the inner
webs with white.
Wing 2°45, tail 1°75, culmen 1-25, tarsus 062.
The discovery of this species by Mr. Everett adds another charac-
teristic Malayan genus to the Philippine avifauna.
35. NECTAROPHILA SPERATA (122).
[S. Leyte, ¢, September. ]
36. ARACHNECHTHRA JUGULARIS (123).
[Amparo, J 2, J uly, August. ]
37. Corvus PHILIPPINUS (125).
(S. Leyte, g 2, July. }
38. CALORNIS PANAYENSIS (128).
[Amparo, ¢, August; S. Leyte, 3 2, September and October, ¢,
Juv., September : iris brownish ochre-yellow. |
The young bird is in brown and striated plumage.
39. SARcops caLvus (129).
[Amparo, 3 9, July.]
These two examples have the whole back, from the nape to the
uropygium, and also the scapulars dark brown. This is also the case
in my Zebu series, and in some individuals from South Negros. In
one Negros example the back is dark brown, many of the feathers
being fringed with various shades of grey. In another the amount
of grey and brown on the back is about equal ; and it would seem that
the grey colouring of the back belongs to the fully adult bird, and
that the nape, uropygium, and upper tail-coverts assume the full
grey colouring of the adult before the dorsal plumage commences to
turn grey ; the whole of the upper plumage is brown at the base,
the tips alone changing to grey.
40. Munta sacorr (132).
[Amparo, ?, July; 8. Leyte, ¢, September. ]
344 THE MARQUIS OF TWEEDDALE ON { Mar. 19,
41, PHABOTRERON BREVIROSTRIS.
Phahotreron brevirostris, Tweeddale, P. Z. 8. 1877, p. 549.
[Amparo, 9, July. ]
42. CARPOPHAGA ZNEA (141).
[Amparo, ¢ 2, July. |
43. TurTUR DUssUMIERI (147).
[Amparo, ¢ 2 .]
44. CHARADRIUS FULVUS (159).
[S. Leyte, ¢ 2, September and October. }
In non-breeding plumage.
45. EupRoMIAs GEoFrFrroy! (161).
[S. Leyte, ¢ 2, September and October. }
46. Eupromias MonGoticus (163).
[S. Leyte, 2, September. |
47. AGIALITES DUBIUS (162).
[S. Leyte, 2 ¢, September and October. ]
All the examples, nine in number, are of birds in full breeding-
plumage.
48. ANGIALITES PERONII.
Charadrius peronii (Temm.) ; Schlegel, Mus. Pays-Bas, Coraces,
p- 33; Walden, Tr. Z. S. viii. p. 90. no. 142, pl. x, f. 2.
[S. Leyte, 2 , October. |
In immature plumage ; first primary only two thirds grown ; sides
of breast and narrow connecting band brown, mixed with rufous ;
broad frontal patch pure white, bounded posteriorly by a narrow
brown band ; crown and occiput rufous brown; tarsus one inch.
49. MELANOPELARGUS EPISCOPUS.
[Amparo, ¢, July. |
In full plumage.
50. GALLINULA CHLOROPUS (169).
[Amparo, ¢ 2, July.]
51. ORTYGOMETRA CINEREA (172).
[Amparo, ¢ 9, July; S. Leyte, 3, September. ]
52. Porzana Fusca (174).
[S. Leyte, ¢, September : iris brilliant red ; orbital ring crimson ;
bill olive-green, culmen black ; legs light dull carmine. |
A single example, which belongs to the small race, 7. e. true P.
JSusca, and not to P. erythrothorax.
1878. | THE ORNITHOLOGY OF THE PHILIPPINES. 345
53. RALLINA EURYZONOIDES.
Gallinula euryzonoides, Lafr. Rey. Zool. 1845, p. 368.
[Amparo, ¢ 9, July and August ; S. Leyte, ¢ 2, September. |
54. AMAURORNIS OLIVACEA (176).
[S. Leyte, ¢ 2, Jnly.]
55. Hypor2nipia rorquata (177).
[Amparo, d, July; S. Leyte, 2, July. ]
The female example differs from every other individual of this
Philippine Rail I have examined in having the broad pectoral band
coloured like the back.
56. Hypor#nrpia sTriaTA (179).
[Amparo, d 9, July; S. Leyte, ¢ 2, September. ¢ juv., Sep-
tember, bill purple brown, the culmen dark brown. |
57. TOTANUS INCANUS.
Scolopax incana, Gm. S. N. i. p. 658.
[S. Leyte, 2, October: iris dark brown ; bill dark brown, nearly
black ; legs light greenish ochre. |
58. TRINGOIDES HYPOLEUCUS (183).
[S. Leyte, 2 , September. |
59. GALLINAGO SCOLOPACINA (187).
[S. Leyte, ¢ 2, September. ]
60. GaLLInaGo MEGALA (188).
[S. Leyte, 2, September. |
61. RayNCH#XA CAPENSIS (189).
[S. Leyte, ¢, September. ]
62. ARDETTA CINNAMOMEA (192).
[Amparo, ¢, July; S. Leyte, ¢ 9, September. |
63. Heropias GARZETTA (195).
[S. Leyte, 2, September: iris light yellow ; orbital skin yellow-
ish, clouded with dark grey ; bill black, base of mandible whitish ;
legs black ; feet greyish green ; upper surface of toes tinged with
purplish. ]
64. Buroripes savanica (197).
[Amparo, g, July; S. Leyte, September. |
One example in full, another in immature plumage.
65. NycricoRAxX MANILLENsIS (198).
[Amparo, 2, July; S. Leyte, g , September: iris golden-yellow ;
orbital skin yellow-green; bill black; legs pale ochreous white
but brown in front and on upper surface of toes. ]
The male is in full breeding-plumage. The lengthened occipital
346 MR. P. L. SCLATER ON THE [Mar. 19,
plumes are black throughout their length, and not only at their tips,
as described by Vigors and figured by Fraser.
66. STERNA BERGII.
Sterna bergit, Licht. Verz. d. Doubl. Mus. Berl. p. 80.
[S. Leyte, ¢, October. ]
67. DENDROCYGNA VAGANS (203).
[Amparo, 6 2, July; S. Leyte, ¢, September. ]
2. Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘ Challenger”—No. VIII. On the
Birds of the Sandwich Islands. By P. L. Scuatzr,
M.A., Ph.D., F.B.S.
[Received February 26, 1878.]
The ‘Challenger’ arrived in Hilo Bay, Owhyhee, on the 14th of
of August, 1875, and left on the 19th of the same month.
The skins of birds collected were 24 in number, belonging to 13
species. They were all obtained “ within eight miles of the town.”
The collection, although small, and containing nothing absolutely
new except a single species of Anas, is of interest, as it enables us
to record the actual island of the Sandwich group upon which the
species contained in it were found, and as including an example
of the little-known Buteo solitarius of Peale.
The best account of the avifauna of the Sandwich-Island group,
which is still very imperfectly known to us, is that given by Dole in
18691. On this I have commented at some length in ‘ The Ibis’ for
1871°, and given my views on the subject pretty fully. Lately M.
Oustalet has described a most remarkable new form * from the same
group ; but I am not aware of any other more recent contributions
to this subject *.
1. CHASIEMPIS SANDVICENSIS.
Sandwich Flycatcher, Lath. Syn. ii. p. 343, undé
Muscicapa sandvicensis, Gm. 8. N. 1. p. 945.
Chasiempis sandvicensis, Cab. Wiegm. Arch. 1847, i. p. 208.
Cnipolegus, sp. 1238, Scl. Cat. A. B. p. 203; vide P. Z. 8. 1873,
p- 595.
No. 528. Hilo, Owhyhee. 9.
BOD Aoge cnn: Cogn rhs ‘tok
“Flycatcher: eyes black ; stomachs contained insects.” —J. MW.
1 «« Synopsis of the Birds hitherto described from the Hawaiian Islands. By
8. B. Dole,” Proc. Boston Soc. N. H. xii. p. 294.
2 «Remarks on the Avifauna of the Sandwich Islands. By P. L. Sclater,”
Ibis, 1871, p. 356.
2 Lowioides bailloni: vide Bull. Soc. Phil. de Paris, 7™° sér. t. i. p. 99.
4 Dr. Pucheran’s somewhat peculiar views on the avifauna of the Sandwich
Islands will be found explained in the Procés-Verbaux Soc. Phil. de Paris,
1858, p. 85.
1878.] BIRDS OF THE SANDWICH ISLANDS. 347
2. PH#ORNIS OBSCURA.
Dusky Flycatcher, Lath. Syn. vol. ii. pt. i. p. 344, undé
Muscicapa obscura, Gm. 8. N.i. p. 945.
Tenioptera obscura, Cassin, B. U.S. Expl. Exp. Atlas, pl. ix.
fig. 3.
Pheornis obscura, Scl. Ibis, 1859, p. 327, et 1871, p. 360.
No. 534. Hilo, Owhyhee. <¢.
535. ” 2 2 :
536. e ‘
537. ”? 3) ”
“ Eyes brown; bill and feet black.’””-—J. M.
3. Mowoa NOBILIs.
Gracula nobilis, Merrem, Av. Descr. et Ic. p. 8, pl ii.
Mohoa nobilis, Cass. Pr. Ac. Sc. Phil. vii. p. 440.
Yellow-tufted Bee-eater, Lath. Syn. i. p. 683, undé
Merops niger, Gm. 8. N. i. p. 465.
No. 524. Male. Hilo.
** Eyes black ; stomach contained fruit and insects.”—J. WM,
4, PsiTTIROSTRA PSITTACEA.
Parrot-billed Grosbeak, Lath. Syn. ii. p. 108. pl. 42, undé
Lowxia psittacea, Gm. S. N. i. p. 844.
Psittirostra psittacea, Sw. Class. B. ii. p. 295.
538. Male. Hilo.
545. Male. Hilo.
** yes red ; stomach contained fruit.”—J. WM.
As I have already stated (Ibis, 1871, p. 360), this bird has, in
my opinion, no relationship to the Fringillide, but is merely an
exaggerated form of the same type as Hemignathus and its allied
genera.
5. Drepanis CoccinEa (Merrem).
Mellisuga coccinea, Merrem, Av. Descr. et Ic. p. 14, pl. iv.
(1786).
Drepanis coccinea, Cassin, Zool. U.S. Expl. Exp. p. 177.
521. Female adult.
522. Young female. } Hilo.
523. ” ”
‘Eyes red ; bill and legs orange; stomach contained insects and
seeds.”—J. MV,
6. DREPANIS SANGUINEA (Gm.).
Certhia sanguinea, Gm. S. N. i. p. 479.
Drepanis sanguinea, Bp. Consp. 1. p. 404.
530. Female. Hilo.
348 MR. P. L. SCLATER ON THE [ Mar. 19,
7. DreEPANis FLAVA, Bloxham.
Drepanis flava, Bloxh. Voy. Blonde, p. 249 ; Dole, Proc. Boston
Soc. N. H. xii. p. 298.
525. Male.
526. Male. . r
527. Female. Be Oe ares
533. Female.
‘« Eyes black ; stomachs had seeds and insects.’—J. M.
The females are much less yellowish than the males, especially
below.
8. Munra nisorrA (Temm.); Walden, Trans. Zool. Soe. viii. p. 73.
543. Male. “Sparrow, introduced; native who shot it said he
had never seen one before.”-—J. MV.
An imperfect skin of this species (or of the nearly allied M. punce-
tularia), no doubt introduced.
Prof. Baird a short time ago sent me a skin of a young or female
of another Eastern Finch from the Sandwich Islands (probably
Munia malabarica) for determination; also without doubt introduced.
9. Orus BRACHYOTUS (Forster).
Strix sandwichensis, Bloxam, Voy. Blonde, p. 250.
Asio galapagoensis, Cassin, Mamm. et Orn, U.S. Expl. Exp.
p- 207.
Asio accipitrinus, Sharpe, Cat. B. ii. p. 234.
544. “Male; eyes yellow; brought on board alive.”—J. M.
10. Burro soLiTaRivs.
Buteo solitarius, Peale, Zool. U.S. Expl. Exp. Birds, p. 62 (1848).
Pandion solitarius, Cassin, Mamm. & Orn. U.S. Expl. Exp.
p. 97; Atlas, pl. iv.
Polioaétus solitarius, Sharpe, Cat. B. i. p. 452.
No. 540. Female.
« Eyes brown; cere and legs yellow; bill black.”—J. M.
Mr. J. H. Gurney has kindly examined this interesting bird for
me (of which two specimens were obtained, but one of them was sub-
sequently lost), and supplied me with the following notes :—
«The female Buzzard brought from Owhyhee by the ‘ Challenger’
Expedition (No. 540 in that collection) appears to be of the same
species, though in a different stage of plumage, as the previously
unique specimen from the same island, which is preserved in the
Museum of the Academy of Natural Sciences at Philadelphia, and
which was described by Peale, in the first edition of the ‘ Zoology of
the United-States Exploring Expedition,’ published in 1848, under
the name of Buteo solitarius.
“In the subsequent edition of this work, published in 1858, and
edited by the late Mr. Cassin, the generic name of Buteo, as applied
to this species, was dropped, and that of Pandion substituted, with
the following remark :—‘ This bird is strictly a member of a subgenus
1878. | BIRDS OF THE SANDWICH ISLANDS. 349
of the generic group Pandion designated Polioaétus by Dr. Kaup.’
(Vide op. cit. p. 98.)
“ Mr. Ridgway, on the contrary, after au examination of the type
specimen, wrote to me that he considered it ‘a Buteonine form dif-
fering from the true Buteones ouly ...... in the system of coloration,
which reminds us somewhat of Milvago chimachima.’
‘In the specimen brought home in the ‘ Challenger’ there is much
less resemblance to the coloration of M. chimachima, and nothing,
in my opinion, to justify the removal of this species from the genus
Buteo.
«The following is a comparison of the measurements of the Chal-
lenger example with those of the type specimen as given by Mr. Cassin
in the work to which I have referred :—
Type specimen ‘Challenger’ specimen
(sex unknown). (female).
inches. inches.
Total length of skin........ 17 (above) Wir
Ree Fue, OE Ae pie aD 12
MEANS. £87290 WS BL SOR: 75 72
Bill from gape ............ 15 1°5
Datsngy 212A A 2°75 2°8
Hind toe and claw ........ 1*75 2°2
Middle toes. u. .......... a8 Leg
Claw of ditto.............. 0:8
Culmen (exclusive of cere) .. I'l
Gere yest ons DER: 0:4
**Mr. Cassin describes the type specimen as having ‘the third,
fourth, and fifth quills longest and nearly equal,’ which agrees with
the ‘Challenger’ specimen, as does his description under the head
of ‘form’ generally ; but I think that, in speaking of the toes as
having ‘their under surfaces strongly corrugated or pustulated,
claws very large, long, and curved,’ he uses rather stronger language
than is strictly applicable to the ‘ Challenger’ specimen, though in
this the under surface of the feet is certainly somewhat corrugated
and roughened, and the claws are slightly longer than in Buteo vul-
garis or in Buteo desertorum.
*« As regards the difference of coloration in the two specimens, Mr.
Cassin describes the type as having the ‘ head and entire underparts
and upper tail-coverts yellowish white, all the feathers being pure
white at their bases and widely terminated with pale yellowish, more
distinct on the breast and tibize.’
“In the ‘Challenger’ bird the upper part of the head is dark
wood-brown like the mantle, except that some of the feathers, espe-
cially on the crown and forehead, are partially edged with yellowish
white; on the sides of the head these white edgings are more con-
spicuous ; the throat is white; the breast, abdomen, under tail-
coverts, flanks, and wing-linings are white, slightly tinged with buff,
especially on the tibize, and more or less mingled with wood-brown
throughout, except on the lower abdomen and tibize, the brown pre-
350 ON THE BIRDS OF THE SANDWICH ISLANDS. [ Mar. 19,
dominating over the buffy-white on the flanks, and almost entirely
covering the sides of the breast.
‘Mr. Cassin further describes the type specimen as having the
‘occiput and neck behind with oblong longitudinal spots of umber-
brown, which is the colour also of all the upper surface of the body,
wings, and tail; shafts of quills reddish chestnut on their upper
surface, white beneath ; inner webs of quills widely edged with white ;
tail beneath silky-white tinged with yellowish.’
“The ‘Challenger’ specimen agrees with this portion of Mr. Cassin’s
description, with the following exceptions, viz.:—the occiput and
hinder part of the neck are white-coloured and of the same hue as the
mantle; the upper surface of the shafts of the quills is brown, but
without any ‘ reddish-chestnut’ tint ; the upper surface of the tail
is brown, with transverse bars of a darker brown; on the middle pair
of rectrices these are eleven in number, the lowest bar being subtermi-
nal; on all the rectrices the edge of the outer web and the tip are ot
a darker brown than the remainder of the feather, but beyond the
dark tip there is at the extreme end of the feather a small central
spot of white, which is more conspicuous on a new rectrix only par-
tially grown than in the older feathers, from some of which it has
been worn off by use ; the inner webs of the lateral rectrices are also
more or less mottled with white towards the edge. The dark trans-
verse bars are indistinctly apparent on the under surface of the tail.
“The figure of the type specimen (U.S. Explormg Expedition,
pl. iv.) agrees with the description given by Mr. Cassin, and also shows
a larger bare space behind the eye than exists in the ‘Challenger’
specimen; but, notwithstanding this difference and those of coloration
to which I have alluded, I think that the coincidence of size and form
marks the two specimens as belonging to the same species, and that
the example obtained by the ‘Challenger’ Expedition must be con-
sidered as a specimen of Buteo solitarius of Peale.”
11. ANAS WYVILLIANA, Sp. nov.
«Anas boschas,”’ Dole, Pr. Boston 8. N. H. xii. p. 8305; Finsch
et Hartl. Orn. C. P. p. xxxix. (7)
541. Male.
542. Male.
Supra nigra, plumis fusco limbatis ; pileo nigro, fusco minute pune-
tato ; subtus pallide fusca, in gutture et pectore magis rufescens,
in ventre magis ochracea, plumis nigro punctatis et maculatis ;
alis extus brunnescenti-cinereis ; speculo amplo purpureo, mar-
gine supra et infra albo inde nigro ornato; awillaribus albis ;
rostro superiore nigro, inferiore carneo; pedibus aurantiacis.
Long. tota 15°0, ale 9°3, caude 3°0, rostri a rictu 2°0, ejusd.
lat. sub naribus 0°7, tarsi 1-5.
Hab. Inss. Sandwich, maris Pacifici.
This Duck belongs to the section of true Anas which embraces 4.
obscura of North America, A. superciliosa of Australia, and other
species, and in which both sexes somewhat resemble the female of
Anas boschas. The head is dark, finely pointed with brown, and
} Eyes brown; legs and feet orange.
1878.] MR. FORBES ON BIRDS FROM THE SAMOA ISLANDS, 35]
bears no trace of a superciliary stripe; the speculum is shining
purple with a black border above and below ; beyond the black
border below the coverts terminate in a broad margin of pure white ;
above the upper black border, which is formed by the ends of the
small coverts, there is a narrow ashy-white margin,
The description is taken from specimen 541; the other (542) is
apparently a young bird and is rather smaller in its dimensions.
12. Furica aut, Peale.
Fulica alai, Peale, Zool. U.S. Expl. Exp. Birds, p- 224; Cassin,
Mamm. & Orn. U.S. Expl. p. 306, Atlas, pl. xxxvi.
531. Male. ‘“Eyesred; bill flesh-coloured with a yellowish tinge ;
feet and legs pale slate-colour.”°—J. M.
13. Toranus IncaNnus (Gm.).
532. Female. Eyes hazel ; bill brown ; legs yellow.
3. On a Small Collection of Birds from the Samoan Islands
and the Island of Rotumah, Central Pacific. By W. A.
Forzss, F.Z.S,
[Received March 1, 1878.]
Mr. Sclater has lately put into my hands for determination a small
collection of birds from the above localities, made by the Rev. G.
Brown, C.M.Z.S., of the Wesleyan Mission. It consists of 47 skins
referable to 19 species. Of these all but 4 skins, of two species, are
from the Samoan Islands of Upolu and Sayaii, and are well known
already as inhabitants of these islands. They require no further
notice here, except one.
PACHYCEPHALA ICTEROIDES, Peale; Finsch & Hartl. Faun.
Centr.-Polyn. p. 76.
Three examples of this species, in different stages of plumage,
though none are quite adult, traces of the rufous plumage of imma-
turity remaining to a greater or less extent. All are marked “from
Upolu ;” and the native name given is “ Vasavasa.” Mr. Layard
(P. Z. S. 1876, p. 494) doubts the occurrence of this species in
Samoa, and refers the bird from those islands to P. flavifrons (Peale),
which, as Dr. Finsch has shown (J. f. O. 1872, p- 39), is the adult
male of P. icteroides.
The island of Rotumah is, as far as I am aware, entirely unknown
ornithologically ; and the three skins in this collection from that
locality are, I believe, the first that have ever been received from the
island. It is a small island, 4 or 5 miles in extent from north to
south, and was discovered by Captain Edwards in his search for the
‘Bounty’in1791. It lies nearly midway between the Fijisand the Ellice
group, in about long. 177° E., lat. 122° S. Some account of it will
352 ON BIRDS FROM THE SAMOA ISLANDS. [ Mar. 19,
be found in the ‘South-Pacific Directory,’ 3rd edition, p. 627 (London,
1871). The two species of birds of this island represented are,—
Aptonis vitrensis, Layard, P. Z. 8. 1876, p. 502.
Aplonis tavuensis, F. & H. Faun. Centr.-Polyn. p. 103, t. x. f. 2
(nec Gmelin).
One skin of this species, agreeing with skins from Fiji in the
British Museum, and others from the same islands in the ‘ Chal-
lenger’ collections, determined by Dr. Finsch as “‘ Aplonis vitiensis,
Lay.” (conf. Finsch, P. Z.S. 1877, p. 735). The native name is
given as “ Husila.”’
MyzoMELA CHERMESINA, G. R. Gray, G. B. i. pl. 38; id. Cat.
B. Trop. Islands, p. 11.
This species was figured by Mitchell in the ‘ Genera of Birds,’
but not described ; nor was any locality mentioned for it in the list
of species of Myzomela in the same work. Bonaparte, however, in
his ‘ Conspectus,’ gave ‘‘ New Guinea” as the habitat, though what
reason for this he had, other than that at that time ‘‘ Nova Guinea”’
was a convenient “refuge for the destitute,” is not known. The
figure in the ‘ Genera’ represents a bird with a uniform scarlet under
surface; but fortunately the original specimen is still in existence,
mounted in the Bird Gallery of the British Museum. ‘Two speci-
mens of a Myzomela, certainly distinct from any other known as
inhabiting the Pacific Islands, were contained in Mr. Brown’s collec-
tion from Rotumah; and a look at Gray’s figure sufficed to indicate
considerable differences between the two birds. On examining, how-
ever, Gray’s type of his M. chermesina, it became evident at once
that the two birds were really identical, and that the apparent differ-
ence, consisting in the belly and vent being of a uniform red in the
plate, instead of a brownish-black, was due to a mistake on the artist’s
part. As Myzomela chermesina has not yet been described, and is
irrecognizable from Gray’s figure, I herewith proceed to give a
diagnosis and description.
MyzoMELA CHERMESINA, G. R. Gray, Gen. B. i. pl. 38; id. B.
Trop. Isl. Pacif. p. 11; Bp. Consp. Av. i. p. 394. sp. 3 (‘ex Nova
Guinea”!) ; Gray, Handl. B.i. p. 154. no. 1989 (“New Guinea? ”’}.
3 (ad.?). Fusco-nigricans, alis caudaque nitore nonnullo metal-
lico ; mento, gula, pectore lateribusque abdominis, eum dorso
uropygioque, nitide coccineis, plumis basi nigris ; rostro nigro,
pedibus brunneo-corneis. Long. tot. circa 43, rostr. 2, al. 3,
caud. 2 (poll. Angl.).
Male. Fuscous-black ; interscapulars and wing-coverts darker, and
with a slight metallic gloss, which extends onto the tail; primaries
browner, the internal web narrowly margined with whitish, except
at tip; wings beneath pale greyish-brown; chin, throat, breast,
sides of belly till near legs, back, rump, and upper tail-coverts shi-
ning crimson-scarlet, the feathers black at base; beak black ; legs
dark horn-colour.
1878.] MR. F. NICHOLSON ON BIRDS FROM DARRA-SALAM. 353
A second specimen, marked female, resembles in general that
described, but is rather smaller, and all the colours are duller, par-
ticularly the red of the throat and chest, so that the black-brown of
the back forms a broad ring between the chin and breast. The
under wing-coverts are whitish, and the pale internal margin of the
primaries more conspicuous. It is probably a younger bird.
Habitat. Island of Rotumah, Central Pacific. Native name
““Aramea.” I ought to mention that Mr. Sharpe has lately re-
ceived, in a collection from Erromango, one of the New Hebrides,
a specimen of this bird almost identical with mine in every respect.
This is very curious ; for Erromango is far removed from Rotumah,
and the neighbouring island of Tanna is inhabited by a distinct
species (Myzomela cardinalis (Gm.), figured in Latham’s Synopsis,
vol. i. pl. xxxiii. fig. 2).
4, A List of the Birds collected by Mr. E. C. Buxton at
Darra-Salam, on the coast of Africa opposite Zanzibar.
By Francis Nicuotson, F.Z.S.
[Received March 4, 1878.]
My friend Mr. Buxton, whose ornithological collection from
Sumatra was described by the Marquis of Tweeddale last year
(‘ Ibis,’ 1877, p. 283), and whose name is therefore already well
known to naturalists, has recently returned from a short visit to the
Zanzibar coast, and has brought with him a small collection of birds.
These he has lent to me for description; and I give a list of the
species in the present paper, merely adding my regret that it is not
a longer one. Owing, however, to a sharp attack of fever, Mr.
Buxton was only able to remain on the coast a little more than a
fortnight ; and this accounts for the smallness of the collection. I
feel sure that ornithologists will agree that, considering the col-
lecting of birds was not the primary object of Mr. Buxton’s expedi-
tion, the number of species is not a meagre one for a fortnight’s
shooting.
Our knowledge of East-African ornithology is not very extensive
at present ; and in the present paper I have referred to the only two
essays which have dealt with the birds of the Zanzibar district.
These are as follows :—
1. Hartlaub, G. Report on a Collection of Birds formed in the
\sland of Zanzibar by Dr. John Kirk.—P. Z. S. 1867, p. 823.
2. Sharpe, R. B. On a Collection of Birds from Mombas, in
Eastern Africa.—P. Z. S. 1873, p- 710.
There remains also the list of birds procured by the late Baron
von der Decken, and described by Dr. Cabanis in the third volume
of his ‘ Reisen in Ost-Afrika,’ and the inestimable work of Drs. Finsch
and Hartlaub, the ‘ Vogel Ost-Afrika’s.’ I have followed the order
and classification adopted in the Jast-mentioned book.
Proc. Zoou. Soc.—1878, No. XXIII. 23
354 MR. F. NICHOLSON ON [Mar. 19,
I have only to add that the locality where this collection was
made is Darra-Salam, described by Mr. Buxton as a splendid har-
bour, on the main coast, S.E. of Zanzibar: the country is agricul-
tural for a mile, and then covered with bush and low timber. The
birds were procured in July 1877.
Besides the birds brought by Mr. Buxton I am indebted to Dr.
Giinther for permission to examine a small parcel of skins sent to
him by Messrs. I. and F. Moir of Zanzibar, and collected by them
also at Darra-Salam. Although most of the birds had already been
obtained by Mr. Buxton, there are one or two not included in that
gentleman’s captures.
1. ASTUR TACHIRO. =
Astur tachiro (Daud.), Sharpe, Cat. B. i. p. 99.
Nisus tachiro, Finsch & Hartl. Vog. Ostaf. p. 78.
A young female in Messrs. Moir’s collection.
2. ASTURINULA MONOGRAMMICA.
Asturinula monogrammica (Temm.); Finsch & Hartl. Vog. Ost-Afr.
p. 59; Sharpe, P. Z. S. 1873, p. 711; id. Cat. B.i. p. 275.
Two specimens. It is quite evident from the way in which these
two birds differ in the character of the bars on the under surface,
that there is some variation in this respect to be accounted for, and
A. meridionalis, Hartl. (Cf. Sharpe, Cat. i. p. 277) cannot be sepa-
rated as a species.
3. GLAUCIDIUM CAPENSE.
Glaucidium capense (Smith) ; Sharpe, Cat. B. ii. p. 223.
Athene capensis, Finsch & Hartl. ¢. c. p. 98.
Two specimens of this rare little Owl, agreeing with Mr. Sharpe’s
description of the type in the British Museum. Dr. Finsch and
Hartlaub considered it probable that this species would be more
plentiful on the east coast of Africa than it is in South Africa; and
this seems likely to prove the case.
4. EuRYSTOMUS AFER.
Eurystomus afer (Lath.) ; Finsch & Hartl.¢. ¢. p. 150; Sharpe,
Ibis, 1871, p. 274; id. P. Z. S. 1873, p. 712.
Two specimens: wing 6°8 inches.
5. CoRACIAS CAUDATA.
Coracias caudata(L.); Hartlaub, P. Z. S. 1867, p. 824; Finsch &
Hartl. ¢. c. p. 154; Sharpe, Ibis, 1871, p. 194; id. P. Z, S. 1873,
p- 712.
6. HALCYON CHELICUTENSIS.
Halcyon chelicutensis (Stanl.); Finsch & Hartl. ¢. ¢. p. 162;
Sharpe, ¢. ce. p. 712.
Halcyon striolata, Hartl. t. ¢. p. 824.
Two specimens.
1878.] BIRDS FROM DARRA-SALAM. 355
7. CERYLE RUDIS.
Ceryle rudis (L.); Finsch & Hartl. ¢. c. p. 175.
Two specimens of this widely distributed species, which, however,
does not seem to be so common in E. Africa as elsewhere, as this is
the first record of it from Zanzibar.
8. Mrrops sUPERCILIOSUS.
Merops superciliosus, L.; Finsch & Hartl. ¢. c. p. 178.
Three specimens, all of the brown-headed type, the true M. super-
eiliosus.
9. Mrrors PUSILLUs.
Merops pusillus (Mill.); Sharpe, ¢. c. p. 712.
M. minutus, V.; Finsch & Hartl. ¢. c. p. 188.
Three specimens ; the eyebrow in each being very distinctly blue.
10. MERoPs ALBICOLLIS.
Merops albicollis, V. ; Finsch & Hartl. ¢. c. p. 185; Sharpe, €. ¢.
p- 712.
Several specimens of this very common Bee-eater.
11. DicroceRcUS HIRUNDINACEUS.
Dicrocercus hirundinaceus, Sharpe, ed. Layard, B. S. Afr. p. 101.
Merops hirundineus, Finsch & Hartl. ¢. ¢. p. 193.
On comparing this specimen with other African individuals, I
notice that the colour of the blue on the abdomen and under tail-
coverts is paler, and I can find no trace of a blue eyebrow in Mr.
Buxton’s specimen.
12. InRISOR ERYTHRORHYNCHUS.
Irrisor erythrorhynchus (Lath.); Hartl. ¢. ¢. p. 824; Finsch &
Hartl. ¢. c. p. 202; Sharpe ¢. ¢. p. 712.
One specimen.
13. CINNYRIS MICRORHYNCHUS.
Cinnyris microrhynchus, Shelley, Monogr. Cinnyridz, part 1.
Nectarinia jardinii, Hartl. P. Z. S. 1867, p. 824; Finsch &
Hartl. ¢. c. p. 218, pl. ii. fig. 1; Sharpe, ¢. ec. p. 713 (nee Verr.).
Cinnyris jardinii, Cab. in Von der Decken’s Reis. iii. p. 29.
An adult male fully bearing out the distinctions put forward by
Captain Shelley.
14. CrnnyRIS GUTTURALIS.
Cinnyris gutturalis (L.); Shelley, Monogr. part 1; Sharpe, ed.
Layard, B. S. Afr. p. 311.
N. gutturalis, Hartl. t. c. p. 824; Finsch & Hart. ¢. c. p. 216 ;
Sharpe, ¢. c. p. 713.
Cinnyris gutturalis, Cab. t. ¢. p. 28.
Several specimens.
23%
356 MR. F. NICHOLSON ON [Mar. 19,
15. ANTHODIEZTA ZAMBESIANA.
Anthodieta zambesiana, Shelley, Monogr. Cinnyr. part 1; Sharpe,
ed. Layard, B. S. Afr. p. 321.
N. collaris, Hartl. t. c. p. 824; Finsch & Hartl. ¢.c. p. 223;
Sharpe, ¢. c. p. 713 (nec V.).
N. corallis, Cab. t. c. p. 28.
An adult male individual, confirming the characters given by
Captain Shelley.
16. DRYMG@CA AFFINIS.
Drymeca affinis, Smith ; Sharpe, ed. Layard, B. S. Afr. p. 259.
One example. Cf. Sharpe, /. c.
17. CisTICOLA ISODACTYLA.
Cisticola isodoctyla, Peters; Sharpe, ed. Layard’s B. S. Afr.
p- 269.
Drymeca isodactyla, Finsch & Hartl. t. ¢. p. 236.
Three specimens.
18. MAcRONYX STRIOLATUS.
Macronyz striolatus, Heugl. J. f. O. 1863, p, 164.
M. croceus, Finsch & Hartl. ¢. c. p. 276 (nec V.).
Two specimens. I think that Heuglin was wrong in reuniting his
M. striolatus with M. croceus. It seems to me to be a distinct bird,
with the sides of the neck streaked with white near the black collar,
whilst JZ. croceus has no white streaks on the side of the neck.
19. ANTHUS RAALTENI. —
Anthus raalteni, 'Temm.; Hartl. ¢. c. p. 825; Cab. ¢. e. p. 22;
Finsch & Hartl. ¢. ¢. p. 274.
Three specimens agreeing with Zanzibar skins of Dr. Kirk’s in the
British Museum, and identified as above by Drs. Finsch & Hartlaub.
20. CRATEROPUS KIRKI.
Crateropus kirkii, Sharpe, Layard’s B. S. Afr. ed. 2, p. 213.
One specimen of a Babbling Thrush in bad order. It appears to
belong to Mr. Sharpe’s new species.
21. PYCNONOTUS NIGRICANS.
Pycnonotus nigricans (V.) ; Finsch & Hartl. ¢. ¢. p. 297 ; Sharpe
P. Z. S. 1873, p. 712.
Izus nigricans, Hartl. t. c. p. 825.
Three specimens.
22. ORIOLUS LARVATUS.
Oriolus larvatus, Licht. ; Sharpe, P. Z. S. 1873, p. 714; id. Cat.
B. iii. p. 217.
An adult in Messrs. Moir’s collection.
1878. } BIRDS FROM DARRA-SALAM. 357
23. BUCHANGA ASSIMILIS.
Buchanga assimilis (Bechst.) ; Sharpe, Cat. B. iii. p. 247.
Dicrurus divaricatus (Licht.) ; Finsch & Hartl. ¢. ec. p. 323;
Sharpe, ¢. c. p. 714.
Two specimens.
24. TELEPHONUS ERYTHROPTERUS.
Telephonus erythropterus (Shaw); Finsch & Hartl. ¢. c. p. 336°
Sharpe, ¢. c. p. 714.
Two specimens.
25. DRYOSCOPUS AFFINIS.
Dryoscopus affinis, Gray; Hartl. ¢. c. p. 824.
Laniarius affinis, Finsch & Hartl. ¢. c. p. 349.
L. salime, iid. t. ec. p. 349.
Dryoscopus cubla et D. salime, Sharpe, ¢. ec. p. 714.
I forwarded these birds to my friend Mr. R. Bowdler Sharpe, who is
working at the genus, and am indebted to him for the synonymy as
above, and for the following note:—“I am glad to see that Dr.
Hartlaub, in his new work on the Birds of Madagascar, unites D.
salime to D. affinis. The two specimens now sent by Mr. Buxton
represent these two supposed species—the male being D. affinis, and
the female with white lores and grey rump D. salime. . A large
series of dated and sexed specimens must be examined before we can
understand for certain the plumages of these little Bush-Shrikes.
In the present pair the female has a totally black wing, and has
grey bases to the scapulars, while the male has a narrow white edging
to some of the wing-coverts, and very distinct white bases to the
seapulars.”
26. BRADYORNIS PALLIDUS.
Bradyornis pallidus (Miller) ; Sharpe, Cat. B. iii. p. 310.
B. subalaris, Sharpe, ¢. c. p. 713, pl. lviii. fig. 1.
Two specimens.
27. SIGMODUS GRACULINUS.
Sigmodus graculinus (Cab.) ; Sharpe, Cat. B. iii. p. 325.
Prionops graculinus, Cab. t. c. p. 24, taf. iii.
A specimen in changing plumage, but without any white bar on
the lower surface of the wing.
28. PHOLIDAUGES VERREAUXI.
Pholidauges verreauxi, Bocage in Finsch & Hartl. Vog. Ostafr.
p- 867; Sharpe, ¢. c. p. 714.
P. leucogaster, Finsch & Hartl. ¢. c. p. 376.
An adult male, with the characteristic white edging to the outer tail-
feather. It was included in the ‘ Vogel Ost-Afrika’s’ as P. leucogaster,
from its being met with in Mozambique by Commander Sperling.
There can be little doubt, however, that P. verreauxi of South Africa
358 MR. F. NICHOLSON ON [Mar. 19,
goes up the east coast, and extends at least as far as Angola on the
west coast.
29. EsTRELDA CYANOGASTRA.
Estrelda cyanogastra (Daud.) ; Sharpe, Cat. Afr. B. p. 65. no.
616.
A single specimen in Messrs. Moir’s collection.
30. HyPHANTORNIS NIGRICEPS.
Hyphantornis nigriceps, Layard; Finsch & Hartl. ¢. e. p. 392;
Sharpe, ¢. c. p. 715.
One specimen of this Weaver-bird was included in the collection
sent by Messrs. Moir.
31. PyTELIA MELBA.
Pytelia melba (L.); Finsch & Hartl. ¢. c. p. 441.
A female specimen, and of course difficult to determine correctly
from this sex only ; but it appears to agree with examples from South
Africa in the British Museum. Drs. Finsch and Hartlaub, more-
over, record it from South Mozambique, where Dr. Peters ob-
tained it.
32. EUPLECTES NIGRIVENTRIS.
Eupiectes nigriventris (Cass.) ; Sharpe, ¢. ¢. p. 715.
Pyromelana nigriventris, Finsch & Hartl. ¢. e. p. 415.
An adult male specimen.
33. EuUPLECTES CAPENSIS.
Euplectes capensis (L.).
Pyromelana capensis, Finsch & Hartlaub, ¢. ¢. p. 416.
An adult male specimen.
34. PasseR DIFFUSUS.
Passer diffusus, Smith ; Hartl. t. e. p. 826.
P. swainsoni, Finsch & Hartl. ¢. c. p. 451.
Two specimens.
35. CoLrus LEUCOTIS.
Colius leucotis, (Riipp.) ; Finsch & Hartl. ¢. ¢. p. 472; Sharpe,
tvenp: 714.
One specimen.
36. Tockus MELANOLEUCUS.
Tockus melanoleucus (Licht.); Sharpe, ed. Layard, B.S. Afr.
p. 127.
Buceros melanoleucus, Finsch & Hartl. ¢. e. p. 485.
Two specimens.
37. Pa@ocEPHALUS FUSCICAPILLUS.
Peocephalus fuscicapillus (Verr.}.
1878.] BIRDS FROM DARRA-SALAM. 359
Psittacus fuscicapillus, Hart). t. ec. p. 826.
Pionias fuscicapillus, Finsch & Hartl. ¢. ec. p. 499.
Peeocephalus Suscicapillus, Sharpe, ¢. ¢. Dei Alle
Three specimens.
38. Poconoryuyncuus MELANOPTERUS.
Pogonorhynchus melanopterus (Peters); Finsch & Hart]. ee
p- 904 ; Marsh. Monogr. Capit. pl. vii.
An adult specimen of this rare Barbet.
39. BarBaTuta pusiLua.
Barbatula pusilla (Dum.) ; Marshall, ¢. c. p. 117, pl. xlviii.
I have carefully compared Mr. Buxton’s specimen with two others
in the British Museum from Abyssinia (Blanford); and I find them
quite identical in coloration, the Zanzibar bird being the least bit
smaller (wing 2°05 in.). This species is new to the avifauna of Eastern
Africa.
40. Coccystrs JAcoBINvus.
Coccystes jacobinus (Bodd.) ; Sharpe, P. Z. S. 1871, p. 597, et
1873, p. 711.
One specimen in moult.
41. Cenrropus superciirosvs,
Centropus superciliosus (Riipp.) ; Hartl. ¢. ¢. p. 826.
Several specimens,
42, TRERON DELALANDII.
Treron delalandii (Bp.); Hartl. ¢. ¢. p- 827: Finsch & Hartlaub,
t.c. p. 535.
One specimen agreeing with a South-African example of the above-
named species, excepting that the size is rather smaller (wing 7-2
inches), and the bill rather long. The markings and colour of the
tail appear to be identical.
43. FRANCOLINUS GRANTI.
Francolinus grantii, Hart). P. Z. S. 1865, p. 665, pl. 39. fig. 1;
Finsch & Hartl. ¢. ¢. p. 589.
Identical with typical specimens of Francolinus rovuma, Gray
(Cat. Gallinze Brit.’ Mus. 1867, p. 52), which is identified with F.
granti by Drs, Finsch and Hartlaub.
44. AcTiITIs HYPOLEUCUS.
Actitis hypoleucus (L.); Hartl. ¢. ec. p. 827; Finsch & Hartl.
é. c. p. 752.
One specimen.
360 MESSRS. GODMAN AND SALVIN ON | Mar. 19,
5. Descriptions of new Species of Central-American But-
terflies of the Family Erycinide. By F. DuCane Gop-
MAN and OsBert SALVIN.
[Received March 7, 1878.]
The species of Erycinide described in the present paper are some
that we have been unable to find names for when examining the
Central-American representatives of the family. Specimens of the
greater part of them are in our collection; but some have been
described from examples in that of Mr. H. Druce, whilst others
have been lent to us by Dr. O. Staudinger of Dresden, and by Mr.
T. Belt. We hope, before long, to publish figures of the whole of
them.
1. EuRYGONA HYPOPH A.
g. Exp. 1:15in. Outer margin of secondaries slightly angulated ;
above nearly uniform dark brown with a slight rufous tint in the
centre of each wing; beneath brown; across the primaries beyond
the cell a broad, indistinct, dark band; across the middle of the
secondaries a similar band; a submarginal row of nearly obsolete
black spots on the secondaries.
2. Exp. 1:2in. Outer margin of secondaries more strongly angu-
lated in the middle than in the male; above dull brown, slightly
paler on the distal half of the secondaries ; three white spots on the
primaries, one at the end of the cell and two beyond it, one on either
side of the second median branch: beneath, much paler, nearly
uniform ; an indistinct, narrow, dark band passes from the costa
of the primaries inside the two outermost of the white spots and
across the secondaries, curving round to near the middle of the
inner margin ; a submarginal row of black spots with minute white
ones associated with them on the secondaries.
Hab. Veragua, Chiriqui (Ribée).
Mus. nostr. (3) et Dr. O. Staudinger ( 2 ).
2. EuRYGONA LEUCORRHOA.
3. Exp. 1°25 in. Secondaries slightly produced, margin entire ;
above deep red-brown; costa, apex, and outer margin pure dark
brown: beneath pale earthy brown, distal third of the primaries
and the apex of the secondaries darker ; distal half of the secondaries
silvery grey ; a narrow brown line crosses the middle of the median
branches of the primaries, and crossing to the secondaries curves
within the space between the first median branch and the submedian
nervure and passes to the inner margin; the secondaries have a
submarginal row of black spots surrounded by silvery white, that
between the lower radial and second median branch being the
largest ; the margin itself at its most prominent part is rufous.
Hab. Veragua, Chiriqui (Ribde).
Mus. nostr. et Dr. O. Staudinger.
1878. ] CENTRAL-AMERICAN BUTTERFLIES. 361
3. EuRYGONA AMPHIDECTA.
d. Exp. 1*3 in. Secondaries slightly produced, outer margin dis-
tinctly dentate; above brownish red; costa and outer margin of
both wings (the secondaries only as far as the middle of the outer
margin) deep brown: beneath nearly uniform brown; a common
narrow band runs beyond the cell across the primaries, and curving
gradually on the secondaries passes to the inner margin; a black
spot with white outer edge near the margin of the secondaries
between the lower radial and second median branch; between it
and the anal angle are five similar smaller spots; margin at the
anal angle rufous ; inside the spots is an indistinct band parallel to
the margin. é
¢. Exp. 1‘5in. Earthy brown, each wing paler in the middle,
with a central spot as dark as the margin ; beneath paler, nearly
uniform, the dark bands more distinct than in the male, and the
secondaries with a row of submarginal nearly obsolete dark spots,
and a similar row of less defined spots round the outer margin of the
primaries.
Hab. Veragua, Chiriqui (Rise).
Mus. Dr. O. Staudinger.
4, EURYGONA INCONSPICUA.
9. Exp. 11 in. Secondaries rounded; upper surface uniform
pale earthy brown, rather darker about the apex of the primaries ;
through this colour the bands of the underside are faintly seen:
beneath whitish tinged with fawn-colour ; a rufous band crosses both
wings through the end of the cell in each, and turning sharply
between the first median branch and submedian nervure of the
secondaries passes to the inner margin; a second band crosses the
ptimaries between the end of the cell and the apex ; the outer margin
of the same wings is dark but less rufous than the bands; on the
secondaries is a submarginal row of black spots, those on either side
of the median nervure being the largest (the lower one larger than
the other) and nearly round; those near the anal angle are linear
and surrounded by white; the margin of the secondaries is
narrowly rufous, and a dark band runs between the large black
spots and parallel to the inner band from the costa to where the
inner band turns to the inner margin.
Hab. Costa Rica, Cache (H. Rogers); Veragua, Chiriqui (Ridée).
Mus. nostr. et Dr. O. Staudinger.
5. EuRYGONA RUSSATA.
Q. Exp. 1°5in. Above brown, a large fulvous spot on the median
branches of the primaries ; secondaries, except the margin next the
apex, red-brown, darker than the spot on the primaries: beneath
greyish white ; marginal fourth of the primaries darker ; a narrow
dark band crosses the primaries beyond the cell, and following irre-
gularly the curve of the margin of secondaries passes to the inner
margin ; a submarginal row of black subtriangular spots edged with
362 MESSRS. GODMAN AND SALVIN ON {Mar. 19,
white on the secondaries, that between second median branch and
lower radial the largest.
Hab. Veragua, Chiriqui (4rcé).
Mus. H. Druce.
6. LimNAas MELANOCHLORA.
3. Exp. 1:3 in. Above black, with a broad central stripe of rich
orange passing longitudinally through both primaries and secondaries
from base to outer margin: beneath, as above, but somewhat more
subdued in colour.
Hab. Veragua, Chiriqui (Ribde).
Mus. nostr. et Dr. Staudinger.
Obs. Allied to L. bryazis, but the wing-stripes narrower and their
edges parallel; their colour, too, is orange instead of yellow.
7. LIMNAS CRRULEATA.
3. Exp. 1°55in. Above dark, with a rich blue gloss ; across the
primaries to the costa, a little beyond the middle, a distinct orange-
yellow band with parallel edges: beneath dark blue-black, with the
band of the primaries as above ; between the submedian and median
nervures, and between the median branches of the secondaries are
blue-grey streaks, which become more distinct near the anal angle.
Hab. Veragua, Chiriqui (Rzbbe).
Mus. nostr.
It will probably be necessary to remove this species, L. bryaxis, and
L. melanochlora from Limnas and place them in a distinct genus.
8. CYRENIA PYRIPPE.
3. Exp. 1°8in. Allied to C. martia. Secondaries shorter, and
outer margin less produced and rounded; the white and red spots
of the secondaries above nearly obsolete ; beneath the white spot of
the same wings is oval instead of round, and placed nearer the outer
margin.
Hab. Veragua (Arcé).
Mus. nostr.
9. ITHOMEIS IMITATRIX.
Ithomeis eulema, Butl. & Druce, P. Z. 8. 1874, p. 353 (nec Hew.).
dé. Exp. 2°15 in. Primaries and margin of the secondaries black ;
a curved row of six ochraceous spots nearly parallel to the outer
margin of the primaries, the three nearest the costa elongated ; three
similar spots, one near the end of the cell and one on each side of
the second median branch; an obsolete streak between the median
and submedian nervures, and the secondaries (except the margin)
tawny red; four small yellow spots within the black margin of the
secondaries: beneath as above, the spots in the margin of the
secondaries being larger and seven in number; this black margin
passes on to the base of the wing, leaving the basal half of the costa
tawny red.
Hab. Veragua and Costa Rica.
1878. ] CENTRAL-AMERICAN BUTTERFLIES. 363
Mus. H. Druce.
Obs. Allied to I. eulema, from which it differs in the primary
wings being black almost to the base, and in the deeper tint of the
tawny red of the secondaries.
10. SymmacuiA RHACOTIS.
3. Exp. 1:2in. Allied to S. domitianus, from which it differs
above in having a green patch near the anal angle of the secondaries
instead of a submarginal line of the same colour ; beneath the colour
is paler and more ochraceous, and the metallic Spots more numerous
and more regularly dispersed over the whole under surface of the
wings.
Hab. Guatemala, Polochic valley (Hague); Honduras, S. Pedro
(G. M. Whitely).
Mus. nostr.
11. MrseneE IGNICAUDA.
3. Exp. 1:0 in. Above black, a spot on the inner margin of either
secondary and the abdomen red; beneath dark brownish, profusely
Spotted with black grey-edged spots.
2 like the ¢; but the red spot extends across the secondaries
and nearly to the end of the cell of the primaries ; beneath the red
spot of the primaries appears as above; abdomen dark.
Hab. Veragua (Arcé).
Mus. nostr.
Obs. Allied to M. @rope, but the red of the secondaries more
restricted, and the abdomen red instead of black.
12. MEsENE sILarIs.
3. Exp. 1:05 in. Margins of both wings (except the inner broadly)
brownish, enclosing a pale lemon-yellow patch common to both
wings ; beneath exactly as above.
Hab. Nicaragua, Chontales (Belt).
Mus. T. Belt et nostr.
Perhaps allied to 1, pyrippe, but differing both from it and all
its immediate congeners by the paleness of the inner portion of the
wings (in M. pyrippe these are red) and by the breadth of the dark
border.
13. MESENE TYRIOTEs.
dg. Exp. 1'lin. Above brownish black, a broad band of orange
red crossing the middle of both secondaries and the body; beneath
uniform brownish black, rather paler than above.
Q dark brown, a band of orange-red (paler than in the male)
crossing the primaries from the middle of the costa to the anal
angle; beneath as above, but rather paler in colour.
Hab. Veragua, Chiriqui (Ribbe), Calobre (Arcé),
Mus. nostr.
Obs. A close ally of M. sagarts, in which the orange band in the
364 MESSRS. GODMAN AND SALVIN ON [Mar. 19,
male passes onto the primary wings. In the present species this band
is restricted to the secondaries.
14. PAcCHYTHONE GIGAS.
2. Exp. 1°65 in. Black, with a large ovate spot in the apical third
of the primaries, another, pointed, at the base of the wing, and ex-
tending along the inner margin, but not reaching to the costa, and
the secondaries, except the margin, pale yellow; beneath as above,
the margin of the secondaries not extending over the whole of the
costa.
Hab, Panama (Ridéde).
Mus. Dr. O. Staudinger.
Oés. In coloration this species resembles the female of Aricoris
gelasine, described by Westwood as 4. myrtis. The very minute
palpi and the short antennee indicate its proper position in the genus
Pachythone. It is, however, much larger than any member of the
genus hitherto described.
15. CHARIS VELUTINA.
o. Exp. 1:15in. Dark brown above, with darker obsolete bands
on the central portion of both wings, arranged more or less concen-
trically, two subparallel submarginal metallic bands on both wings,
the inner one on the primaries irregular and interrupted: beneath
rich dark purple with obsolete black spots corresponding in position
with the indistinct bands of the upper surface ; a single narrow sub-
marginal metallic band much interrupted on the anterior wings ;
cilia of both wings dark with small white equidistant spots.
Hab. Guatemala, Polochic valley (H. Hague).
Mus. nostr.
Obs. Allied to C. iris, Staudinger ; but the under surface differs in
being purple instead of rich indigo-blue. In C. iris the fringe of
both wings is pure white; in the present species it is dark with but
few white spots.
16. CHARIS CROCEA.
3. Exp. 1:15in. Above rich orange-red, with dark spots along
the costa, outer margin of the primaries, and outer and inner margins
of secondaries; those on the outer margin of the primaries almost
confluent, and with the marginal row of the secondaries containing
the metallic spots characteristic of the genus ; the inner area of the
wings devoid of spots : beneath, as above, but the spots of the costa
more confluent and those of the outer margin of the secondaries less
so; the spots, too, are generally dispersed over the wings.
Hab. Veragua, Calobre (Arcé).
Mus. nostr.
Obs. Allied to C. perone, which it resembles as to colour; but the
spots above are fewer in number and not so generally distributed.
17. CHARIS HOLOSTICTA.
2. Exp. 1°08in. Above lemon-yellow sprinkled with small dark
1878.] CENTRAL-AMERICAN BUTTERFLIES. 365
spots nearly evenly distributed over the whole area of the wings; a
submarginal metallic line round the outer margin of both wings:
beneath as above, the yellow colour being a little paler and the spots
slightly larger. ;
Hab. Veragua (Arce).
Mus. nostr. et H. Druce.
Obs. Also allied to C. perone, but the ground-colour lemon instead
of orange-yellow, and the spots on the wings rather fewer in
number,
18. CHARIS PH@CILOPTERA.
Q. Exp. °95 in. Above pale greyish brown, profusely spotted
with yellowish-white spots, those on the primaries gradually becom-
ing larger towards the outer margin, the margin itself being broadly
unspotted, except a submarginal row of minute spots, joining which
on their outer sides are small black spots ; the submarginal metallic
spots, though present, are scarcely visible: beneath, the primaries as
above, but the secondaries are silvery-white with a few irregular
dark spots chiefly in the region of the costa.
Hab. Veragua, Chiriqui (Ride).
Mus. Dr. O. Staudinger.
19. CHARIS OCHRIAS.
g. Exp. 1-2in. Above white with a slight yellowish tint ; pri-
maries with four black spots along the costa, one at the apical angle,
two on the outer margin, one at the anal angle, two on the inner
margin, and one within the area of the wing beyond the cell;
secondaries with six black spots round the edge and three within
the area of the wing; submarginal metallic line (as usual in the
genus) but feebly developed: beneath precisely as above.
Hab. Costa Rica, Irazu (H. Rogers).
Mus. nostr.
Obs. Allied to C. sulphurea, R. Felder (with the type of which it
has been compared), from Mexico, from which it differs in being
paler in colour and in having all the spots on the outer margin
larger ; there is also a spot on the inner margin of the primaries,
wanting in C. sulphurea.
20. THAROPS PURPURATA.
¢. Exp. 1°5 in. Above rich purplish blue, apical portion of pri-
maries and apex of secondaries brownish black ; some dark elongated
transverse spots on the distal two thirds of the primaries: beneath
greyish white, apex of primaries and outer margin of both wings
brownish ; small brownish spots over the whole under surface, those
nearest the apical and anal angles of the secondaries slightly the
largest.
8 like the male, but the wings more obtuse, and the secondaries
above, like the primaries, crossed with dark marks.
Hab. Guatemala, Polochie valley (Hague) ; Veragua (Arcé).
Mus. nostr.
366 MESSRS. GODMAN AND SALVIN ON [Mar. 19,
Obs. Allied to T. menander, but more purple in tint above;
beneath purer white, and the spots at the apical and anal angles of
the secondaries smaller. In colour it resembles 7. coruscans and
T. nitida, but differs from both in the arrangement of the spots of
the under surface.
21. THAROPS ISTHMICA.
g. Exp. 1-5in. Allied to T. splendida, but smaller, and the anal
angle of the secondaries less produced : beneath the ground-colour
is nearly uniformly sprinkled with pinkish scales, the central area
of both wings in 7’. splendida being decidedly whiter; there are no
conspicuous spots at the apical and anal angles of the secondaries,
and the spot on the costa is much smaller.
Hab. Veragua, Calobre (4rcé).
Mus. nostr.
92. LEMONIAS PELARGE.
3. Exp. 1°33 in. Primaries, base, and narrow outer margin of
secondaries dark brown; some indistinct blue spots near the apex
of the primaries ; a spot near the anal angle of the primaries and the
inner area of the secondaries reddish orange: beneath grey sprinkled
with dark scales; proximal half of both wings with distinct black
spots; outer marginal area of both wings darker, variegated with
lighter marks, especially at the apical angle of both wings and the
anal angle of the primaries.
Hab. Guatemala, Polochic valley (H. Hague).
Mus. nostr.
Obs. Allied to L. parthaon, in which the blue spots of the pri-
maries are very distinct; in the present species the orange of the
secondaries extends almost to the costa: beneath the primaries have
much more blue-grey, and the submarginal spots of the secondaries
are obsolete, being large in L. parthaon.
23. LEMONIAS THEAGES.
3. Exp. 1:05in. Above dark brown, central area of secondaries
to the inner margin, and numerous spots over the margin of the
secondaries and the whole area of the primaries (two, one at the end
of the cell and one below it, the largest), white ; beneath as above,
except that the base of the secondaries is white, and bears several
black spots.
Hab. Veragua (Arcé) ; Costa Rica (Rogers).
Mus. nostr.
Obs. Allied to L. orpheus; but the white spots of the primaries
are much larger, and the dark margin to the secondaries broader.
24. LEMONIAS DEBILIS.
3g. Exp. 1:2in. Dark brown above irrorated with ochraceous
scales, dark brown spots with ochre margins irregularly dispersed over
both wings, except along the outer margin of both, where they lie
contiguous to one another, forming a submarginal row: beneath as
1878.] CENTRAL-AMERICAN BUTTERFLIES. 367
above; but, the ochraceous scales being more numerous than the
brown ones, the under surface has a paler tint.
2 like the male, but with the wings less pointed.
Hab. Veragua, Chiriqui (Ribée) ; Nicaragua, Chontales (Belt).
Mus. nostr. et T. Belt.
25. LEMONIAS HYPOGLAUCA.
?. Exp. 1°15 in. Outer margin of primaries curved outwardly ;
the apex slightly faleate: above greyish brown with blackish spots
dispersed over the surface of both wings, those within and below the
cell of the primaries the largest ; the spots on the secondaries are
arranged in two submarginal rows which follow the curvature of the
outer margin; and an irregular cluster in and about the end of the
cell: beneath whitish, with spots corresponding to those of the
upper surface, those on the primaries dark, those on the secondaries
rufous.
Hab. Mexico.
Mus. H. Druce.
Besides this specimen in Mr. Druce’s collection, we have seen
others in the Berlin Museum.
26. NyMPHIDIUM HZMATOSTICTUM.
dé. Exp. 1-6 in. Costa and outer margin of primaries dark brown,
a triangular patch with its base on the inner margin and apex
towards the apical angle, an isolated spot near the apex, and the
whole central portion of the secondaries white; a submarginal
row of black spots on both wings, bounded on either side by narrow
blue-grey waved lines; margin of the secondaries inwardly red for
two thirds of its length from the anal angle, the rest and a narrow
margin dark brown: beneath with the margins as above, but with
the inner irregular line encircling the black submarginal row of spots
nearly white ; two irregular white spots on the outer margin of the
primaries, one near the middle and one near the apical angle ; the
red of the secondaries more restricted.
Hab. Panama (Ridbe).
Mus. Dr. O. Staudinger.
27. NYMPHIDIUM ICTERICUM.
d. Exp. 1:6 in. Base of secondaries and primaries ochre-brown ;
a broad band, constricted in the middle, passes from the middle of
the costa towards the middle of the outer margin, a large irregular
spot from the anal angle halfway along the inner margin, an irre-
gularly defined double submarginal row of spots, and the whole of
the remainder of the secondaries ochre-yellow ; across the cell and
the region of the median nervure are some obsolete dark transverse
bars: beneath as above, but slightly paler.
@ like the male, but less ochraceous in tint, the lighter markings
being purer yellow.
Hab. Veragua (Arcé) ; Chiriqui (ibde).
Mus. H. Druce (3), Dr. O. Staudinger ( ).
368 MESSRS. GODMAN AND SALVIN ON | Mar. 19,
28. NyMPHIDIUM SICYON.
¢. Exp. 1-6 in. Costa of primaries, base, and outer margin of
both wings dark brownish-black ; inner margin and central area of
both wings yellowish ochre; in the dark margin of the secondaries
are two parallel yellowish submarginal lines, between which are a
series of black spots: beneath paler than above; the submarginal
lines in the border of the secondaries are absent; but the black spots
are present.
Hab. Guatemala, Polochic valley (H. Hague).
Mus. nostr.
Obs. Allied to N. agle, but the outer margin darker, and the
margin of the primaries and secondaries are without the rufous band
contained in that of N. agile.
29. NYMPHIDIUM ADELPHINUM.
3. Exp. 1°85in.; 9, exp. 2°15 in. Closely allied to WV. lycorias,
of which it is a more southern race; it differs in having the white
band in the dark margin of the secondaries almost obsolete in the
male instead of a biconvex conspicuous spot ; the white spot near
the middle of the outer margin of the primaries is much smaller:
beneath, these spots, though more distinct, are not nearly so large as
in the allied species.
Hab. Costa Rica, Irazu (Rogers); Veragua, Calobre (Arcé).
Mus. nostr.
30. NYMPHIDIUM VELABRUM.
3. Exp. 1°8in.; 92, exp. 1:95. Allied to WN. phliasus; but the
white band common to the two wings is broader, and on the pri-
maries extends just into the cell instead of merging in the fulvous
band of the same wings ; this fulvous mark, in the present species,
is restricted to an isolated spot in the apical area of the primaries.
In the relative position of the white band and the fulvous spot this
species resembles Adelpha iphicla and its allies, whereas N. phliasus
resembles d. cocala.
Hab. Veragua, Calobre (Arcé).
Mus. nostr.
31. THEOPE ISIA.
36. Exp. 1°25 in. Costa, apex (narrowly), outer margin of pri-
maries, and cilia of secondaries dark brownish-black ; secondaries
and a subtriangular patch occupying from the inner margin to the
end of the cell of primaries blue: beneath uniform orange.
Hab. Guatemala, Polochic valley (H. Hague).
Mus. nostr.
Obs. The distribution of the blue of the upper surface resembles
that of 7. hypoxanthe ; the colour of the under surface is paler and
purer yellow, being almost exactly the tint of the under surface of
T. eudocia.
1878. ] CENTRAL-AMERICAN BUTTERFLIES. 369
32. THEOPE BAREA.
d. Exp. 1'15in. Cell of the primaries nearly to the end, the
inner margin (broadly), and the whole of the secondaries dark shining
blue with a slight purple tinge; rest of the primaries dark brown
with a slightly lighter central patch: beneath uniform rather dark
lemon-yellow.
2. Exp. 1-lin. Primaries dark brown, a patch of blue covering
from the inner margin to half the cell, and passing beyond it, but
leaving the outer margin and costa broadly dark ; secondaries the
same blue colour, apex and outer margin (narrowly) dark brown;
aaa the same as the male, the yellow colour being hardly so
right.
Hab. Veragua (Arcé); Chiriqui (Ribde).
Mus. nostr. ( 3), et Dr. O. Staudinger (2).
Obs. In the colour of the upper surface this species is like 7’.
aureonitens, but the blue is darker, as well as the apex of the
primaries ; the colour beneath is much paler yellow.
33. THEOPE CENINA.
3. Exp. :95 in. Inner margin of primaries and cell nearly to the
end and whole of secondaries dark shining blue slightly tinged with
purple ; rest of the primaries dark brown; beneath uniform pale
dirty ochraceous yellow.
Hab. Veragua, Chiriqui (Ridée).
Mus. nostr.
Obés. Allied to 7. barea, described above; but the colour of the
under surface is more ochraceous in tint, paler and not nearly so
pure in tone.
34. THEOPE DECORATA.
¢. Exp. 1°0in. Primaries above and margin of the secondaries
black; a spot on the inner margin of the primaries near the anal
angle, a few scattered scales near the base of the same, and the whole
central area of the secondaries blue: beneath dark brown, the apex
of the primaries lighter, nearly the basal half of the costa of the
primaries and the base of the secondaries yellow; a curved band of
iridescent blue, seen only in a certain incidence of light, occupies
the remainder of the costa of the primaries nearly to the apex, it
then crosses the wing to about the middle of the outer margin.
Hab. Nicaragua, Chontales (Bel¢).
Mus. nostr. et T. Belt.
Obs. Belongs to the T. janus group, from which it differs in many
respects.
Proc. Zoou. Soc.—1878, No. XXIV. 24
370 MR. A. ANDERSON ON A NEW SPECIES OF PRINIA. [Mar. 19,
6. On a new Species of Indian Prinia.
By AnpRew AnpzRson, F.Z.S.
[Received March 18, 1878. ]
(Plate XIX.)
It had been my intention to write a connected account of the birds
of Northern Kumaon, the result of a two months’ ornithological excur-
sion into the Alpine regions of this part of the North-western Hima-
layas during the summer of 18751. Time, however, has failed me,
to say nothing of bad health and heavy official duties ; so that I must
leave all details for another time, and content myself for the present
with offering to the Society a description of a Prinia, which I feel
confident is new to science, and which was obtained in the Bagesur
valley on the 18th June, 1875, at an elevation of between 3000 and
4000 feet.
PRINIA POLIOCEPHALA, sp. n. (Plate XIX.)
Similis P. stewarti, et fronte cinerea pileo concolori, sicut in hac
specie; sed dorso brunneo nec cinereo diversa; rostro nigro ;
pedibus pallide brunneis; iride pallide flavida; palpebris
pallide stramineis.
Long. tot. 3°8, culminis 0°45, alee 1:7, caudze 1°8, tarsi 0°8.
Hab. Kumaon, India.
This species is of the same group as P. cinereocapilla and P.
stewarti, but is distinguished from both by trenchant characters,
which may be expressed in the following synoptic table : —
a. Back grey, as also the entire head................ssceseeeseenerees stewartt.
6. Back rufescent brown, contrasting with the head, which is
prey.
a’. Forehead grey, like the Crown ......ccecsecesserescuveeecees poliocephala.
6'. Forehead fulvous, crown of head grey...... .......-++-e0 cinereocapilla.
This discovery of an apparently new species of Prinia in a com-
paratively speaking well-explored country like Kumaon is an event
of some interest.
I should not, however, omit to mention that I am indebted to my
friend Mr. W. E. Brooks for having brought to my notice that the
bird in question is apparently undescribed, and that it is not ‘ P.
cinereocapilla,”’ by which name I had it catalogued in my note-book.
Unfortunately, I only obtained one specimen of the new bird, a
male ; but at the same place a fair series of P. hodgsoni, all males, was
also added to my collection. Both species of Warblers were in beau-
tiful plumage; an examination into their sexual organs indicated
that the latter end of June and July must be their breeding-season ;
and hence probably the reason why no female examples were met with.
1 Tn an article contributed by me to ‘ Stray Feathers’ for 1875, vol. iii., en-
titled “ Little or unknown Himalayan Oology,’ I have already given an ac-
count of the nidification and habits of some of the birds I met with on this
oceasion.
JSmit kth.
Hanhart imp
PRINTA POLIOCEPHALA.
bth.
Smit
*
_ >
1878. ] MR. J. W. CLARK ON OTARIA URSINA. 371
As might be expected, the subject of the present paper has all
the habits of its near ally P. stewarti, for which, indeed, I mistook
it on the wing. When shot it was wending its way very adroitly
through dense nettle-jungle (Girandinia heterophylia), climbing up
and down the stalks in search of mosquitoes and other insect food.
The allied species to P. policephala, viz. P. stewarti and P. cinereo-
capilla, do not apparently occur in Kumaon.
In concluding my remarks on the subject, I am glad to be able to
add that the Marquis of Tweeddale concurs with me in considering
the Prinia new to science. The Marquis writing to me respecting
this species, gives his opinion in the following terms, which by his
permission, I here transcribe :—“ Your Prinia seems to be a good
and distinct species.”
7. Notes on three stuffed Specimens of the Sea-lion of the
Pribiloy Islands (O¢aria ursina). By J. W. Cuarx,
M.A., F.ZS.
[Received March 19, 1878.]
(Plate XX.)
The specimens I have the pleasure of exhibiting this evening
were sent to the Museum of the University of Cambridge in October
1877, by the Alaska Commercial Company. Some time before that
date the Company had most obligingly acceded to my request that
they would endeavour to obtain for me some specimens of this Sea-
lion, which is the most abundant of the two species found upon the
islands ceded to them for trading-purposes by the Government of the
United States, the other being Steller’s Sea-lion (O¢aria stelleri).
The “set of Seals,” as the Secretary’s letter terms them, consists
of a male, a female, anda pup, from St. Paul’s Island, Alaska. Those
who have read Elliott’s ‘ Report’? will remember that this is the
island which is most thickly peopled by this species of Otaria and
yields the greatest number of skins in each year. The skull and
the bones of the arm and leg accompanied each skeleton ; so that
the age of the animal can be approximately ascertained. From the
condition of those portions of the skeleton, and from the size of the
animals as compared with the measurements given in the work
above quoted and in Allen’s ‘ Monograph’ * (pp. 73-108), it is clear
that they are not full-grown. This is to be regretted, as the oppor-
tunity so seldom occurs of obtaining an O¢aria in that condition: and
from the vast numbers that exist on the Prybilov Islands, I had
hoped that the specimens, when they arrived, would prove to be
adult. Notwithstanding this drawback, however, they are a most
interesting group, and illustrate extremely well the difference in
1 Report on the Prybilov Group or Seal Islands of Alaska. By Henry W.
Elliott, 4to. Washington, 1873.
2 On the Eared Seals (Otariadw). By J. A. Allen,” Bull. Mus. Oomp. Zool.
Harvard College, Cambridge, vol. ii, p. 1. aie
372 MR. J. W. CLARK ON OTARIA URSINA. [Mar. 19,
size and build between the male and female. It is the first time,
moreover, so far as I am aware, that a pair of any species of Otaria
has been obtained from the same locality at the same season of the
year, and also the first time that this particular species has been
seen in this country; and I feel most grateful to the Company for
their generous present. |
The male is 5 feet 10 inches long, measured in a straight line from
the tip of the nose to the root of the tail. The general colour of the
pelage is a black inclining to grey; and the hairs are 1 inch long.
A closer examination shows that the grey tint is due to the
presence of white hairs intermingled with the black ones. On the
neck, right across the whole superior surface, from the back of the
head to the shoulders, the hair is much coarser, and individual
hairs are exactly twice as long as on the back. There are a great
many more white hairs on this part. The hair is equally long
under the throat, but not along the sides of the neck. The shaggy
appearance presented by this part of the pelage in an animal no
older than our specimen gives an indication of what it must be
in full-grown or aged individuals, and fully accounts for the
**mane”’ so often described by the early voyagers. The muzzle is
of a light brown, with very short hair. There is a patch of light-
brown hair near the ear. The arm is destitute of hair near the
distal end of the radius and ulna. There is a patch of a slightly
rufous tinge behind it. A thick rufous under-fur covers the back,
neck, and sides of the body, but it nowhere extends further than
halfway down. The hair under the belly is thinner and shorter;
indeed it is extremely short on that part, and generally rather
lighter in colour than on the rest of the body, especially between
the hind legs. The enormous length of the cartilaginous flaps
that project beyond the toes on the hind feet is characteristic of this
species, and has been already noted and figured by Allen. Exami-
nation of the skull shows that the occipital crest has not yet been
developed, and that the teeth are unworn. The canines have not
reached their full size. The epiphyses of the bones of the arm and
leg are still quite distinct.
The female measures 3 feet along the same line as in the male.
The pelage is generally much closer and finer, and the under-fur
thicker, and longer in proportion to the hair. No white hairs are
admixed with the dark ones; but these latter have light-yellow tips,
which become larger on the breast, so that that part is quite yellow.
The colour under the belly is rufous, edged with a band of a lighter
shade, which has a wavy outline, ascending higher just behind the
arm and in front of the hind leg. The rufous tint is darkest between
the fore legs. The skull shows that the animal is quite young, much
younger probably than the male—an unfortunate circumstance, as we
are unable to appreciate the difference in size between the sexes with
the accuracy that could be wished. It will be seen that this female
is just half the length of the male. Allen (¢. c. p. 76) gives the
length of an adult male as from seven to eight feet, of an adult female
as four feet.
1878. ] MR. A. H. GARROD ON LYCAON PICTUS. 373
The cub (the sex of which was not noted) is of a glossy black
all over the back part of the body, inclining to brown on the nose,
throat, and chest. The parts under the belly, especially between
the hind legs, are of a light brown; and there is a yellow patch
behind the fore limb.
The differences in colour, of which it is almost impossible to give
an intelligible description on paper only, are extremely well illustrated
in the accompanying figures (Plate XX.).
8. Notes on the Visceral Anatomy of Lycaon pictus, and
of Nyctereutes procyonides. By A. H. Garrop, M.A.,
F.R.S.
[Received March 18, 1878.]
Having had the opportunity in my prosectorial capacity of dis-
secting a male specimen of Lycaon pictus, as well as several, both
male and female, of Nyctereutes procyonides, I take the present
opportunity of giving the results at which I have arrived.
Lycaon pictus.—This canine animal, so different from its allies
in its digitation, is not at all aberrant in its visceral anatomy, which
has not been previously described, so far as I am aware. The fol-
lowing are some of the most important details :—
The anterior portion of the palate is black, the pigment extending
back as far as half an inch, onto the soft palate, of which the
posterior one and a half inch is unpigmented. There is no uvula,
a median shallow notch occupying its position. The tonsils are
elongate, lunate, and vertical in position.
On the tongue the filiform papille are all small ; and among them
small papillze fungiformes are sparsely scattered. Three circumval-
late papillee on each side, increasing in size from before backwards,
and converging posteriorly, form the normal V. There is no trace
of a lytta.
Of the salivary glands the compact submaxillaries are slightly
larger than the irregularly shaped parotids. The zygomatic glands
are as big as small chestnuts. The accessory submaxillary (or sub-
lingual) glands are situated nearly in contact, in the middle line of
the floor of the mouth.
The thyroid gland is formed of two parts, each of the size of a
sheep’s kidney, these being joined at the inferior internal angle by a
narrow isthmus of thyroid tissue. The superior thyroid artery is
enormous.
The stomach presented no differences from that of Canis famili-
aris. The following are the lengths of the intestines :—
ft. in.
Small intestine.............. 9 1
Cecuminhes Sey SR 0 7
374 MR. A. H. GARROD ON LYCAON PICTUS. [Mar. 19,
The czecum is quite caniform, its curves being exactly represented
in that of Canis familiaris’.
The liver is deeply fissured, upon the same plan as in all the Canidee
—and all the Carnivora, in fact,—the cystic fissure being very deep,
which allows the fundus of the gall-bladder to appear on the
diaphragmatic surface of the organ. The left lateral lobe is the
largest, the right central and right lateral being slightly smaller.
These last are half as large again as the caudate and left central,
which are at least four times the bulk of the Spigeliau, upon which
latter there is a small accessory lobule.
In the generative organs, as in all Canidee, the prostate is large,
whilst Cowper’s glands and the vesiculz seminales are absent. The
os penis is large, quite straight, four inches in length, and deeply
grooved, as in all the Canide, along its lower surface.
In the lungs there are three lobes to the left, and four to the
right, one of the latter being azygos. The median lobes of both
sides are the smallest ; the inferior the largest. The fissures between
the lobes are all deep.
The brain of Lycaon pictus is perfectly dog-like, resembling that
of Canis lupus (as figured by Leuret and Gratiolet ?) in almost every
detail, the division of the posterior limb of the gyrus third above
the Sylvian fissure extending as far forward on the superior cerebral
surface as in that species, or even further, the anterior superior angle
of the gyrus next below it being rather more strongly developed.
The sulcus between the uppermost (or fourth) gyrus and the
third is parallel to the great longitudinal fissure between the hemi-
spheres.
In Nyctereutes procyonides the tongue is covered with filiform
papillee smaller in size than in Lycaon pictus, allowing the pro-
portionally larger fungiform papillz to appear more conspicuously
among them. These latter posteriorly become the papillee circum-
vallatz, five on each side, larger posteriorly, and arranged in a V-
manner.
There is no uvula; and the soft palate embraces the upper end
of the larynx with facility.
The stomach is not peculiar, except that it is more than usually
muscular at its pyloric end.
In an adult male which died on the 2nd of February last, the
father of a litter of six born on May 2nd 1877, the small intestine
measured eight feet; the large, one foot and an inch, the ceecum
being two inches long, rounded at the end, and slightly turned to the
left side apically. It is figured in the accompanying sketch.
In three other specimens, not adult, the following were the
intestinal lengths :—
1 Vide P, Z. 8. 1873, p. 748, fig. 18.
. * Anatomie comparée du Systeme Neryeux (Paris: 1839-1857), pl. iv. fig.
oup.
1878.] | MR. GARROD ON NYCTEREUTES PROCYONIDEs. 375
a a a i aa.
2, half-grown. 2, a month old. 2, a month old.
res |e seer SE eS IER egret as
Small intestine ...... 4:25 feet. 575 feet. 73 feet.
Large intestine ...... 6 inches, 8 inches, 8 inches,
Cxeum) few 1:25 inches, 1-5 inches, 2 inches.
There is evidently not much constancy in the length of the
viscera, even in specimens of ‘the same age and sex.
Cxeeum of Nyciereutes procyonides.
The liver differs from that of Lycaon pictus and other Canidee in
the great size of the Spigelian lobe. In this the accessory lobule,
referred to above, is enlarged to form part of the lobe itself, which
is, by its presence in a semi-independent condition, rendered bifid
apically. In the depth of the cystic fissure, and all other respects,
it is quite caniform.
The lungs are not peculiar, the fissure between the left upper
and middle lobes only being less developed than in many of its allies,
The azygos lobe is present on the right lung.
The prostate is well developed ; Cowper’s glands are absent, as
are the vesiculz seminales. The os penis is three inches: in length,
straight, and deeply grooved inferiorly to transmit the urethra.
The glans penis is bluntly conical, the ‘urethra opening terminally,
much as in the American Cervidee,
In Nyctereutes procyonides the brain is perfectly caniform,
The posterior limb of the third convolution is bifureate, the bifur-
cating suleus not being lengthy, going upwards and forward with-
376 MR. GARROD ON NYCTEREUTES PROCYONIDES. [Mar. 19,
out having any extension directly onwards to the anterior extremity
of the hemisphere. There is scarcely any tendency in the second
gyrus to form an antero-superior angle; and the sulcus between gyri
three and four is not quite parallel to the great longitudinal fissure,
it diverging slightly from the middle line as it goes forward. In
these respects the brain more resembles that of Canis vulpes’ than
those of Canis familiaris or C. lupus.
In the peritoneal cavity of the adult male Mycéereutes (which,
like the half-grown female, had excessive atheroma of all its larger
arteries) I found an immense number of parasitic worms, collected
especially about the abdominal surface of the liver and the stomach.
These worms had ‘‘ heads”? much like those of the Bothrio-
cephali, but larger. My friend Mr. F. G. Penrose has most kindly
made sections of them, and has demonstrated the existence of a
most peculiar cavity in each. This cavity is coiled up within the
ovate ‘head ;” its lumen is small; and its walls are plicated very
extensively, the magnitude as well as the number of the folds being
great. It opens externally at its proximal extremity by one of its
ends only. The “body” is teenioid in its proportions, and is not
segmented. It is about two and a half inches in total length, the
“head”? being about the size of a hemp-seed or a little smaller.
There are a few general remarks suggested by the above recorded
facts.
First, with reference to the colic cecum in the Canide, I have
on a previous occasion noticed the aberrant form of that appen-
dage in Canis cancrivorus*, where it is nearly straight. Two other
specimens of the species have since passed through my hands,
which have been entirely confirmatory of my earlier observation.
In Nyctereutes procyonides the czecum is slightly more caniform
than in C. eancrivorus; it is a little broader also.
From the examination of other Canide, I find that the czecum,
in its twistings, resembles that of Canis familiaris in being turned
about twice and a half upon itself in C. laniger, C. lagopus *, C.
anthus, C. fulvus, C. antarticus, CO. azare, Otocyon lalandii, and
Lycaon pictus.
In Canis aureus I have found the terminal twist wanting, the apex
of the ceecum turning down as in C. famelicus.
In Canis cancrivorus and in Nyctereutes procyonides the cecum
is nearly straight.
Secondly, with reference to the brain, Prof. Flower has done
much to condense and classify the facts to be arrived at from the
study of the convolutions *, which latter, in my estimation, throw
much light upon.the mutual affinities of the Fissiped Carnivora.
It seems to me that the typical major convolutions of the Car-
nivorous brain form three complete and uniformly broad gyri round
1 Leuret and Gratiolet, Joc, cit. pl. iv. fig. 2, Renard.
2 P. Z. 8. 1873, p. 748.
8 Vide Flower. ‘‘Hunterian Lectures,” ‘ Medical Times and Gazette.’ Lon-
don, June Ist, 1872, p. 622.
4 P.Z. 8. 1869, p. 482.
1878.] THE SECRETARY ON ADDITIONS TO THE MENAGERIE. 377
the Sylvian fissure, which in the Mustelide and in the Genets remains
as such, notwithstanding that these two latter groups had other-
wise diverged before the brain began to modify. From the Mus-
teline animals (the Arctoid ancestral type) the Urside seem to
have diverged, the superior or third cerebral convolution broadening
and tending to divide, whilst the others persist unmodified.
Those Viverridz which are more modified than the genus Genetta,
acquire a broadening of the lowest or first cireum-Sylvian convolu-
tion, especially in its posterior limb, in which a perpendicular
sulcus is formed ; and this peculiarity is more strongly marked in
Hyena, as well as in Proteles. Inthe Felide the anterior as well
as the posterior limb of this first cireum-Sylvian gyrus broadens,
and becomes perpendicularly bisected to such an extent that if in
them there were a longitudinal sulcus developed in the upper median
portion of the gyrus, a complete secondary gyrus would appear.
Such a gyrus, evidently thus originating, is found in the Canide,
in which the extra convolution is therefore a reduplication of the
first, dependent on the differentiation off of its outer moiety.
On the assumption of the correctness of this hypothesis, the
classification of the Fissiped Carnivora might be represented thus:—
Ursidz. Mustelide. Hyenide. Viverridz. Felide. Canide.
7 @ @ @ @ @
eoounh arenas
Yee
= aA
|
@
Ancestral Type.
By Prof. Flower’, after a most careful analysis of their cranial
and other peculiarities, the Canidz are placed between the Arctoidea
and Ailuroidea ; but from the same facts Mr. H. N. Turner” placed
the three major groups in the same order of sequence that the
brain-markings indicate, namely Urside, Felide, and Canide,
which makes it evident that such an arrangement is not opposed to
the teaching of the parts other than the cerebral hemispheres.
April 2nd, 1878.
Prof. Newton, F.R.S., V.P., in the Chair.
The Secretary read the following report on the additions to the
Society’s Menagerie during the month of March 1878.
The total number of registered additions to the Society’s Mena-
gerie during the month of March was 74, of which 32 were by pre-
? P. Z. 8. 1869, p. 4. 2 P. Z.S. 1848, p. 83.
378 THE SECRETARY ON ADDITIONS TO THE MENAGERIE. [Apr. 2,
sentation, 7 by birth, 7 by purchase, 4 were received on deposit,
and 24 by exchange. The total number of departures during the
same period, by death and removal, was 105.
The most noticeable additions during the month of March were
as follows :—
1. An Isabelline Bear (Ursus isabellinus, Horsf.), received in
exchange from the Zoological Gardens of Calcutta, March 7th.
The acquisition of this animal, which has been placed in the Bears’
dens next to its near relative the Syrian Bear (Ursus syriacus),
renders the series of Bears now living in the gardens nearly perfect.
It consists of 21 individuals, referable to 11 species, as follows :—
List of Bears living in the Menagerie.
2 Polar Bears 5,2 (Ursus maritimus).
4 Brown Bears (U. arctos).
1 Hairy-eared Bear (U. piscator).
2 Grizzly Bears (U. feroz).
1 Syrian Bear (U. syriacus).
1 Isabelline Bear (U. isabellinus).
3 Himalayan Bears (U. tibetanus).
2 Black Bears (U. americanus).
3 Malayan Bears (U. malayanus).
1 Spectacled Bear (U. ornatus).
1 Sloth Bear (Melursus labiatus).
2. A Le Vaillant’s Darter (Plotus levaillanti), purchased March 9.
This African species is new to the collection, although we have had
several specimens of its American representative (Plotus anhinga),
and have at present one individual of the latter species living in the
fish-house. The present example is probably from Senegal; but
this Darter likewise occurs all over Southern and Eastern Africa up
to 12° N. lat.’
3. Two examples of the very singular Water-tortoise of the
Amazons, generally known as the Matamata (Chelys matamata),
remarkable for the long pendent filaments on its neck. The larger
of the two measures about 137 inches in length.
This species is new to the collection, and, so far as I know, has
not been previously brought alive to Europe.
I take this opportunity of recording the fact that the large female
Anaconda (Eunectes murinus), which was purchased on the 15th
February, 1877, and which (as I stated P. Z.S. 1877, p. 303) began
to produce young (dead) the 2nd April, 1877, continued to produce
young Snakes (mostly in a more or less decomposed condition) up to
about three months since, when fifteen finally came forth. In all
sixty-two young were produced.
This Snake is further remarkable for its'long fast. Up to the
15th March, when she killed and ate a single duck, all offers of food
were refused ; so that she must have fasted thirteen months.
1 Of. Finsch & Hartl. Vogel Ost-Afrika’s, p. 841.
1878. ] ON THE ORNITHOLOGY OF THE PHILIPPINES. 379
It is impossible to say how long she may have been without food
previous to her arrival at the gardens. One thing is certain, how-
ever, that she could not have taken food while in the box in which
she arrived from South America, as she was so closely packed as to
be barely able to move.
The following papers were read :—
1. Contributions to the Ornithology of the Philippines. No.
VII.—On the Collection made by Mr. A. H. Everett in
the Island of Panaon. By Arruur, Marquis of Tweed-
dale, F.R.S., President of the Society.
[Received March 6, 1878.]
The small collection, of which it is proposed to give an account,
was made by Mr. Everett at San Francisco, on the west coast of
the small island of Panaon. This island is situated to the south-
east of Leyte, from which it is separated by a narrow channel.
Nor is its southern extremity distant from the most northerly part
of Mindanao.
The number of species obtained is too small to permit me to
draw any general conclusions as to the affinities of its ornis.
But the occurrence of the Leyte species of Buceros and Thriponaz,
B. semigaleatus and T. pectoralis, rather than those of Mindanao,
indicates a closer affinity to the northern than to the southern
island.
Panaon has never hitherto been visited by an ornithological col-
lector.
1. CacaTUA BZMATUROPYGIA (1).
[Panaon, 9 , October. ]
2. PRIONITURUS DISCURUS (2).
[Panaon, 3, October.]
3. THRIPONAX PECTORALIS.
Thriponaz pectoralis, Tweeddale, anted, p. 340.
[Panaon, ¢, 2, October. ]
4, CHRYSOCOLAPTES LUCIDUs (32).
[Panaon, 2, October. ]
Crest and crown dark brown, with ruddy fulvous spots.
5. EuRYSTOMUS ORIENTALIS (37).
{Panaon, 3, 2, September. ]
6. Enromosia Guuaris (44).
{Panaon, 3, 2, October. ]
380 ON THE ORNITHOLOGY OF THE PHILIPPINES. [Apr. 2,
7. BUCEROS SEMIGALEATUS.
Buceros semigaleatus, Tweeddale, anted, p. 277.
[Panaon, ¢, 2, October. ]
8. LANIUS LUCIONENSIS (72).
[Panaon, 5, 9, October. |
Several of the examples from Panaon have the whole head above
coloured like the back, uniform liver-brown. There are females
with brown-freckled breasts and flanks.
9. GRAUCALUS STRIATUS (74).
[Panaon, 5, 2, October. ]
10. DicRURUS STRIATUS.
Dicrurus striatus, Tweeddale, P. Z. S. 1877, p. 545.
[Panaon, ¢, 2, October. |
11. BropERIPus ACRORHYNCHUs (90).
[Panaon, ¢, October.]
12. Mixornis CAPITALIS.
Mixornis capitalis, Tweeddale, anted, p. 110, pl. vii. f. 2.
[Panaon, ¢, September. |
13. PotioLopuus urRostictus (101).
[Panaon, 2, October. ]
14. HypsireTEs PHILIPPINENSIS (102).
[Panaon, 9 , October. ]
15. CRINIGER EVERETTI.
Criniger everetti, Tweeddale, Ann. & Mag. Nat. Hist. ser. 4, vol.
Xx. p. 535.
[Panaon, 3, September and October.]
16. Monticoxa souitarius (103).
[Panaon, ¢, 2, October. ]
17. CoryDALLA LuGUBRIS (117).
[Panaon, ¢, October. |
18. DiczuM MODESTUM, Nn. sp.
[Panaon, ¢, October. ]
Above dull olive-brown. Upper tail-coverts and outer margins of
the wing-feathers a brighter olive-green. Rectrices above dark
brown, some narrowly edged with olive-green. Cheeks greyish
brown. Chin and throat greyish white. Breast pale grey. Abdo-
men, ventral region, and under tail-coverts faint yellowish white.
1878. | MR. A. G. BUTLER ON NEW LEPIDOPTERA. 381
Axillaries, wing-lining, and shoulder-edge pure white. Bill, in
dried skins, black.
Wing 1°95, tail 1:00, tarsus 0°37, culmen 0°37.
This Flower-pecker closely resembles Myzanthe pygmea Q. It
differs in wanting the lively olive-green colouring of the upper
plumage of M. pygmea @, more especially on the uropygium, and
the pale yellowish-green lores. The wing too is longer, that of I.
pygmea measuring 1°70. If the sex noted on the label is correct,
this Panaon Diceum is very distinct from M. pygmea S$. If the
genus Myzunthe, as distinct from Diceum, is to be retained, D.
modestum should belong to it.
19. CoRVUS PHILIPPINUs (125).
[Panaon, 3, October. ]
20. PHABOTRERON AMETAYSTINA (139).
[Panaon, 2, September. ]
2. Description of new Lepidoptera of the Group Bombycites,
in the Collection of the British Museum. By Arruur
G. Butter, F.L.S., F.Z.S., &e.
[Received March 8, 1878.]
The following species are believed to be new to science :—
AGARISTID&.
Evusemt1a, Dalman.
1, EvsEMIA INCONGRUENS, 0. sp.
3. General aspect of E. africana, but the primaries with the
first ochreous spot interrupted in the centre ; three ochreous dots in
the form of a triangle behind them; the abbreviated subapical belt
much broader, and not notched at its infero-anterior edge; the spot
at external angle of twice the size; secondaries reddish orange, with
the usual black border. Expanse 2 inches 8 lines.
Q. Spots of primaries paler than in the male; secondaries stra-
mineous, with a much narrower black border. Expause 2 inches
8 lines. Abyssinia,
ARCTIID&.
Pron1A, Walker.
2. PIONIA CALOPTERIDIA, 0. Sp.
Primaries purplish black, crossed by two broad yellowish bands,
the first basal, the second discal and laterally bisinuate ; secondaries
hyaline-white, tinted towards the borders with fuliginous brown ;
382 ; MR. A. G. BUTLER ON NEW LEPIDOPTERA. [Apr. 2,
veins towards the base yellow; head, collar, and thorax purplish
black, tawny or ochraceous at the sides ; abdomen fuliginous brown,
ochraceous at the sides; wings below more clearly defined than
above, the costal borders alternately black and yellow; palpi and
legs black and yellow; venter yellowish. LExpanse, ¢ 1 inch 3
lines, 2 1 inch 2 lines. do, Para; 2, Venezuela (Dyson).
The species of this genus are marvellous mimics of the Coleopte-
rous genus Calopteron.
Axuprenus, Walker.
3. ALPENUS PURUS, 0. sp.
Wings snow-white; primaries with one black dot at base, one
near the base of the median vein, two before the middle of costal
margin, three in a triangle on interno-median interspace, and five
in a curved series beyond the middle; secondaries with a spot at
the end of the cell, one near the outer margin, and two near the
anal angle; thorax snow-white, with a black dorsal line ; collar and
tegulze spotted with black; abdomen ochreous, with a dorsal series
of black spots, anus and ventral surface cream-coloured : primaries
below without spots. Expanse 1 inch 7 lines. Abyssinia.
This species much resembles Spilosoma menthastri, but with the
smooth black-spotted thorax of Alpenus.
TERACOTONA, 0. gen.
Allied to Phragmatobia, but with longer and narrower primaries,
the last subcostal branch with a single instead of a double furca;
discocellulars distinctly angulated ; secondaries more acuminate at
apex ; subcostal branches emitted from a very short footstalk ; dis-
cocellulars more transverse, less distinctly angulated ; radial emitted
further from the third median branch; body much less woolly.
Type, Aloa rhodophea of Walker.
4, TERACOTONA ROSEATA, 0. Sp.
Primaries above reddish testaceous speckled with brown; a black
discocellular spot and a subcostal spot near apex; secondaries rose-
red, brightest.on the abdominal area; a large black discocellular
spot; thorax grey ; head, front of collar, and inuer edge of tegulee
brownish buff; a black spot on each shoulder, and one on each
tegula ; abdomen black, rosy at the base, a dorsal stripe, the pos-
terior margins of the segments and the anus ochreous : wings below
rosy, with black discocellular spots; borders testaceous, speckled
with pale brown; primaries with an irregular transverse dusky discal
belt; body below whity-brown; anterior femora above carmine,
knees black, tarsi banded. with black ; venter ochreous at the sides,
with two lateral series of black spots. Expanse 2 inches 2 lines.
Natal (G@ueinzius).
Allied to T. submacula (Spilosoma submacula, W|k.,=Spilosoma
obscurum, Wlk.), but with longer wings, and differently coloured.
1878. | MR. A. G. BUTLER ON NEW LEPIDOPTERA. 383
Puaos, Walker.
5. PHAOS VIGENS, N. sp.
Primaries dark chocolate-brown with cream-coloured margins ; a
discoidal streak broken into three nearly equal spots, an apical
costal spot, and an angulated stripe beyond the middle (its inferior
extremity terminating at the centre of the interno-median interspace
in an acute spur) cream-coloured: secondaries ochreous, whitish
upon costa, rosy at abdominal border; a black discocellular spot and
a broad regular external black border ; fringe cream-coloured: head
blackish, with cream-coloured frons, collar pale ochreous; tegule
blackish-brown fringed with pale ochreous; thorax blackish ; abdo-
men rose-red, with a dorsal row of black dots; anus ochreous;
wings below with all the paler areas broader and pale clear ochreous ;
body below pale ochreous, varied with blackish ; anterior femora
above carmine. Expanse | inch 4 lines. Tasmania.
We have four examples of this species, which Walker confounded
with his P. fascinula, as he did also the four following.
6. PHAOs NIGRICEPS, n. sp.
6. Allied to the preceding ; primaries black-brown, the base, a
furcate streak from the base, the apical two-thirds of costal margin,
a triangulated discal stripe, an interrupted submarginal line, and the
fringe cream-coloured; base of costa with a tuft of rosy hairs;
secondaries rosy, with black discocellular spot, a broad black ex-
ternal border and ochreous fringe; head and thorax black, collar
and anterior margins of tegule dull buff; abdomen rose-red, with
transverse dorsal black bars and lateral black spots: under surface
with the pale portions buff; anterior femora above carmine. Ex-
panse 1 inch 6 lines.
Q . Basal furca of primaries only represented by a basal dash and
a spot in the cell, discal stripe and submarginal line much more
slender ; pale portions below rosy. Expanse | inch 4 lines.
3. Victoria (Bakewell) ; Sydney (Doubleday).
Readily distinguished from P. fascinula by its black head; in
marking it is nearer to P. vigens.
7. PHAOS NOTATUM, 2. sp.
6. Primaries above dark chocolate-brown ; a longitudinal dash
from the base to near the middle of the cell, a broad streak from the
base to the centre of the inner margin, a subquadrate spot at the
middle of the costal margin, a spot below the latter intersected by
the origins of the median branches, a curved macular discal stripe,
the apical portion of costal margin, the fringe, and a submarginal
series of spots (of which the first, second, and fifth are small, the
others elongated and extended to the margin) cream-coloured :
secondaries rosy with whitish costal area; a large black discocellular
spot ; a broad irregular black submarginal belt; thorax black, head,
collar, and borders of tegule clothed densely with testaceous hair:
body bright rose-red, with black dorsal and lateral spots: under
384 MR. A. G. BUTLER ON NEW LEPIDOPTERA. [Apr. 2,
surface yellower than the upper, the dark areas and markings paler.
Expanse | inch 6 lines. Sydney.
As might be expected, this species is nearly allied to P. fascinula,
from the Hunter River ; it may, however, readily be distinguished
by the broad testaceous fringes to the tegulee and the marking of the
primaries.
8. PHAos NEXUM, 0. sp.
do. Primaries black-brown; the base, an abbreviated internal
streak, a quadrate spot in the cell, an abbreviated acutely angulated
central band from the costa to the first median branch, an angulated
diseal stripe, and a broad external border enclosing a submarginal
series of black spots cream-coloured; secondaries rosy, a black
discocellular spot and a broad black submarginal belt, outer margin
ochreous ; thorax black; head, collar, and fringes of tegule buff ;
abdomen rose-red with dorsal and lateral black spots: wings below
much yellower than above, the black portions slightly paler. Ex-
panse 1 inch 4 lines. Victoria (Bakewell).
This species is chiefly distinguishable by the abbreviated angulated
central belt, and the discal stripe almost uniting with the external
border.
9. PHAOS LACTEATUM, 0. sp.
Primaries cream-coloured ; a nearly central angulated broad black
belt, uot reaching the margins, and interrupted by an irregular spot
upon the median vein; a black sinuous costal stripe from the base,
only separated from the black band by the subcostal vein ; an oblique
blackish basal streak ; four black spots in a discal series before apex,
and three in a pyramidal form near external angle: secondaries
rose-red, with ochraceous borders; a black discocellular spot and a
confluent series of unequal discal spots parallel to the outer margin :
thorax black ; head, collar, and fringes of tegulz sordid buff; ab-
domen rose-red, with dorsal and lateral series of black spots. Under
surface with the costal and internal borders tinted with rose-colour ;
the broad belt of primaries greyish ; body below cream-coloured
spotted with black; femora carmine in front, tibize and tarsi grey.
Expanse | inch 6 lines. Moreton Bay (Damel).
This is a very distinct-looking species, and more like a typical
Arctia than any of the other forms in the genus.
LITHOSIID&.
CisTHENE, Walker.
10. CistHENE NIVEATA, D. sp.
Primaries snow-white, sprinkled sparsely with dark brown scales ;
a dark brown spot at base, and a second on inner margin: secondaries
with the basal half rosy whitish at costa, external half greyish
brown: head and thorax white, abdomen rose-coloured: primaries
below greyish brown, body below white. Expanse 8 lines.
Espiritu Santo.
1878. ] MR. A. G. BUTLER ON NEW LEPIDOPTERA. 385
Mapua, Walker.
11. Mapua DITIS, n. sp.
Primaries bright yellow, with the veins and two marginal lines
carmine; base and outer border sparsely sprinkled with black
scales ; two parallel irregular transverse black lines, the first central,
the second discal; secondaries pink with white fringe; head and
thorax bright yellow, tegulee edged internally with carmine ; abdo-
men pink: primaries below rosy with yellow borders and traces of
two greyish transverse belts; body below cream-coloured, anterior
legs and palpi rose-coloured. Expanse 6 lines. Espiritu Santo.
BaRSINELLA, 0. gen.
Allied to Barsine (section of Miltochrista), but the wings com-
paratively much broader, sborter, and more densely scaled ; the
costal margin of the primaries angulated, the under surface of these
wings crossed at the end of the cell by a perfect fence of long con-
vergent hairs ; the costal margin of primaries with a central fringe
of long hairs, the last two or three segments of the abdomen densely
fringed with hair. ©
Type, B. mirabilis.
12. BaRsINELLA MIRABILIS, N. Sp.
Primaries bright cadmium yellow, two black spots at the base,
a <-shaped marking near the base, two irregular parallel central
lines, and a discal series of dots black, a G-shaped marking at the
end of the cell and the outer margin carmine, fringe golden yellow ;
Barsinella mirabilis.
secondaries rose-red; head and thorax cadmium-yellow, a black
spot on each shoulder; abdomen whitish: primaries below yellow,
the basal half of costal border and a discal series of dots black ;
secondaries paler than above; body ochraceous. Expanse 7 lines.
Espiritu Santo.
NYCTEMERID24.
Auetis, Hibner.
13. ALETIS TENUIS, 0. sp.
Wings fulvous, semitransparent; primaries with the costal and
external margins and a broad subquadrate apical patch enclosing a
Proc. Zoox. Soc.—1878, No. XXV. 25
,
386 MR. A. G. BUTLER ON NEW LEPIDOPTERA. [Apr. 2,
large oblique oval white spot black ; secondaries with the apex and
a more or less confluent marginal series of dots at the terminations
of the nervures black ; body yellow, banded with black. Expanse
1 inch 9 lines. Dar-es-Salem, Zanzibar (J. & F. Moir).
A delicate little species, resembling the New-world genus
Stenele.
14. ALETIS VARIABILIS, 0. sp.
. Ochreous; primaries with the apical two fifths blackish,
enclosing a large transparent white patch beyond the cell, and two
smaller pyriform spots on the median interspaces ; secondaries with
a regular blackish border enclosing nine transparent white spots,
most of which touch the outer margin; body greyish, spotted with
white. Expanse 2 inches 3 lines. Ambriz (Monteiro).
Q@. Var. a. Primaries with the pyriform spots elongated into
ellipses which distort the blackish border, also a third bifid spot
below them completely interrupting the border near the external
angle; secondaries with the white marginal spots confluent and
breaking through the border at apex. Ambriz (Monteiro).
¢g. Var. 6. Primaries with all the white spots confluent, forming
a large irregular hyaline patch which almost covers the disk;
secondaries with the inner edge of the marginal border only trace-
able near the anal angle, the white spots expanded into a broad
hyaline belt, which widens gradually to the costa.
Angola, Bembe Mines (Monteiro).
A. variabilis is much like Stenele lutescens.
The other species of dletis in the Museum were all placed by
Mr. Walker under A. helcita, to which they are allied. They are as
follows :—
1. A. libyssa, Hopffer, from Natal and the Zulu country.
2. A. fascelis, Linnzeus, from the Congo.
3. A. helcita, Clerck, from Western Africa (exact locality un-
known).
4, A. druryi = A. helcita, Drury, Ml. iii. pl. 29. f. 4. Sierra
Leone.
5. A. macularia, Fabricius, from Ashanti.
Of these forms A. /ébyssa is the most decidedly distinct, the others
are mere local modifications of one type differing from one another
in the size and number of the white spots of primaries and the width
of the black border of secondaries. Cramer’s A. helcita is the same
as that figured by Drury.
De1LeEMERA, Hiibner.
15. DEILEMERA SIGNATA, N. Sp.
Allied to D. artemis, Boisd., and D. pellex, Linn. (Nyctemera
separata, Wlk.), but smaller than either, with the broad white
postmedian patch of primaries reduced to a quarter the size, and
with the dark border of secondaries of double the width. Expanse
1 inch 7 lines. Darnley Island (Dr. W. Y. Turner).
1878. ] MR. A. G. BUTLER ON NEW LEPIDOPTERA. 387
CHALCOSIIDS.
Pompeton, Walker.
16. Pomprton AMPLIATUM, 0. sp.
2. Near to P. marginatum from Penang, but larger, the pri-
maries with narrower and duller blue apical border, its inner margin
dentate-sinuate ; blue apical patch of secondaries extending further
up the costal area; yellow spot on under surface of primaries oval
and of double the size; the metallic green costal shot only traceable
upon the veins, and even there of a duller bluish tint; ventral sur-
face of abdomen with the black spots about eight times as large.
Expanse 2 inches 8 lines. Celebes, near Macassar (Wallace).
Liparips.
Morasa, Walker.
17. Morasa LORIMERI, 0. sp.
Allied to M. modesta (Sarothropyga rhodopepla of Felder), but
the primaries black mottled with whity brown, with the outer
border and a biangulated discal belt of whity-brown scales, veins
spotted with orange; secondaries pink, with a broad greyish outer
border not quite reaching the anal angle, fringe sordid rosy ; head
and thorax black ; abdomen rose-red with lateral tufts of ochreous
hair: primaries below smoky grey, with the base ochreous ; second-
aries rather paler than above ; body blackish, clothed with testa-
ceous hairs ; femora rose red. Expanse 1 inch 9 lines,
Tugela river, Natal.
This species was sent to me in a letter by my cousin Miss J. K,
Lorimer, who caught it near the Gordon Mission station.
Notropvontip2.
Rosrma, Walker.
18. RosrMa FULVIPENNIs, n. sp.
Q Primaries dull emerald-green with orange costal margin;
fringe white, tipped with plum-colour and with two apical spots of
the same colour, a snow-white spot at the end of the cell; second-
aries bright fulvous (almost orange) ; thorax green, frons white, with
two fulvous spots ; palpi ochreous ; collar chocolate-brown ; abdomen
fulvous: under surface bright yellow with broad reddish-orange
costal borders; primaries with the outer border pale shining pinky
brown; secondaries with the apical half reddish orange ; legs above
brownish. Expanse 1 inch 9 lines, St. Paulo (Bates).
Allied to R. dorsalis, zelica, myops, and costalis.
SATURNIIDA.
Copaxa, Walker.
19. Copaxa GeMMIFERA, n. sp.
General pattern of C. expandens from Venezuela, but smaller and
25
*»
388 MR. A. G. BUTLER ON NEW LEPIDOPTERA. (Apr. 2,
with more acuminate primaries; above bright stramineous, the
external area densely irrorated with red-brown scales, but leaving
clear yellow patches or spots behind a submarginal series of small
black spots, which are partly united by a zigzag dusky line; a
straight dark brown stripe from the apex of primaries to the middle
of the abdominal margin of secondaries ; a very irregular dark brown
line across the basal area; a small tricoloured ocellus (white, black,
and yellow) with hyaline centre in the middle of each wing; pri-
maries with the basal half of costal border lilacine grey, a broad
dark brown longitudinal belt from the base to the straight oblique
discal stripe, an irregular dentate-sinuate brown line from the costa
to the inner margin just beyond the ocellus, a curved dark brown
streak from the costa to the oblique stripe, two grey-edged apical
white spots ; secondaries with a regular dentate-simuate brown line
just beyond the ocellus; head, collar, and tegule grey; palpi and
prothorax plum-coloured: wings below altogether paler than above,
the markings less defined, the dark longitudinal belt of primaries
obsolete ; body below pale yellow, the anterior tibie and tarsi plum-
coloured. Expanse 3 inches 8 lines. Ambriz (Monteiro).
We also have what seems to be a faded example of this species
from Lake Nyassa; the latter, however, differs somewhat in mark-
ing, and may prove to be locally constant.
Arracus, Linneeus.
20. ATTACUS PRYERI, 0. Sp.
Allied to A. walkeri of Felder from N. China, but darker than
any of the species of the A. cynthia group ; olive brown, with paler
borders and the usual submarginal lines; the pale belt (bounding
the dark angulated central line externally) white inwardly, pinky
whitish and diffused outwardly, with no defined intersecting stripe
as in all the allied species; the maggot-like markings, basal white
belts, and the apical markings of primaries as in A. walkeri. Ex-
panse, ¢ 5 inches 10 lines, 2 6 inches 2 lines.
Yokohama (Jonas).
This species is the most undeniably distinct of all the forms allied
to A. cynthia. I name it after Mr. H. Pryer, of Yokohama.
Abttacus cynthia, as figured by Drury, does not agree with Felder’s
Chinese species, but is certainly nearest to the form occurring in
Java, to which Mr. Moore has given the name of A. insularis. It
may eventually prove to be distinct from the latter, from which it
appears to differ in the more angulated central transverse line ; in
all other characters, such as colour, the strongly dentate pale belt,
and the bent form of the maggot-like marking of the primaries, it is
extremely like the Javan form.
1878. M. E. OUSTALET ON CASUARIUS EDWARDSI. 389
3. Description d’une nouvelle espéce de Casoar (Caswarius
edwardsi). Par M. E, OustTater.
[Received March 12, 1878.]
(Plate XXI.)
Parmi les animaux rapportés par MM. Raffray et Maindron de
leur récent voyage en Nouvelle Guinée et acquis par le Muséum
@histcire naturelle de Paris se trouvent deux Casoars parfaitement
adultes. Le premier de ces oiseaux, tué A Salwatty, en juillet 1877,
est certainement le Casuarius uniappendiculatus, Blyth; Vautre, pris
& la méme époque aux environs de Dorey, ressemble au premier abord
au Casuarius papuanus, von Rosenb., auquel javais été primitive-
ment tenté de le rapporter, mais il me parait différer essentiellement
de cette espéce par Ja forme de son casque. M. Schlegel dit en effet
(Nederl. Tijd. v. d. Dierk. iv. 53) que le Casuarius papuanus a le
casque modelé comme le Casuarius uniappendiculatus. Or dans ce
dernier dont jai, comme je l’ai dit tout-4-l’heure, un individu adulte
sous les yeux, le casque est. en pyramide assez élevée, la face posté-
rieure, ovale, aplatie, étant fortement inclinée, les faces latérales
planes étant disposées en toit, le point culminant du casque se trou-
vant bien en avant de I’ wil, et l’aréte antérieure descendant brusque-
ment de ce point vers le bec. Le méme type de casque s’ observe égale-
ment dans le Casuarius kaupi et dans le Casuarius westermanni
figurés par M. Sclater (Proc. Zool. Soc. 1872, p. 148 et pl. ix., et
1875, pl. xix.). Du reste, dans une lettre qu’il m’a fait ?honneur
de m’écrire il y a peu de jours, M, Salvadori affirme que le Casuarius
westermanni ne différe point du Casuarius papuanus qui habite les
environs de Dorey; et, d’un autre coté, M. Sclater a réuni récem-
ment (Proc. Zool. Soc. 1875, p. 85, note) son Casuarius kaupi
(qui n’est pas celui de M. von Rosenberg) a son Caswarius wester-
manni.
Si, comme le dit M. Schlegel, et comme cela semble probable
d’aprés P opinion exprimée par M. Salvadori, le Casuarius papuanus
a le casque de la méme forme que le Caswarius uniappendiculatus et
le Casuarius westermanni, il est tout-a-fait impossible de rapporter
a la premiére de ces espéces (C. papuanus) Vindividu provenant du
voyage de MM. Raffray et Maindron. Dans ce Casoar en effet, dont
je donne une figure aussi exacte que possible, exécutée d’aprés un
croquis fait par M. Maindron 4 la Nouvelle Guinée, immédiatement
apres la mort de l’animal, le casque, relativement peu élevé, et fort
étroit, se confond presque par sa base avec la téte sur laquelle il
repose: sa face posterieure, au lieu d’étre aplatie et fortement
inclinée, est légérement excavée et s’éléve presque verticalement ; elle
est en outre un peu échancrée 4 son bord supérieur, au point ot
aboutit l’aréte formée par la jonction des deux faces latérales.
Celles-ci sont faiblement bombées et se rencontrent suivant une
ligne qui partant de la dépression signalée sur le haut de la face
390 MR. F. NICHOLSON ON A NEW AMERICAN PIPIT. [Apr. 2,
postérieure, remonte jusqu’a un point situé au-dessous de l’oreille et
redescend ensuite, en décrivant une légére sinuosité, mais en pente
trés-douce, vers le bec, qui est séparé du casque par un petit sillon
transversal. I] résulte de cette. disposition que le point culminant
du casque n’est placé non plus comme dans le Caswarius uniappendi-
culatus en avant de l’ceil, mais notablement en arriére. On peut
noter encore que les faces latérales du casque se prolongent en pointe
sur les cétés de l’aréte supérieure du bec, comme on le voit dans
la figure ci-jomte. Somme toute, le casque présente 4 peu prés le
méme aspect que dans le Casuarius bennetiti et, vu en-dessus, offre
deux arétes que se coupent a angle droit. La coloration des parties
nues n’est pas non plus exactement celle qui est indiaué2 par M.
Schlegel (Mus. des Pays-Bas, Struthiones, et Neder]. Tijds. loc. cit.)
pour son Casuarius papuanus, les taches rouge-carmin qui occupent
les cOtes du cou, au-dessous des yeux et des oreilles, ne s’étendent pas
jusqu’a la tache rouge-orangée qui couvre la nuque, et sont separées
de celle-ci par une teinte bleue; en outre Ja tache rouge-orangée se
prolonge inférieurement sous forme de bandes latérales, lisérées de
bleu enavant. Mais ce sont la des différences plus légéres que celles
que l’on constate dans laformeducasque. Ces derniéres me paraissent
suffisantes pour autoriser la création, en faveur de ce Casoar, d’une
espéce nouvelle que je proposerai de nommer Casuarius edwardsi.
Le type de ma description est un mile, adulte, tué, m’a-t-on
affirmé, auprés de Dorey, sur la céte septentrionale de Ja Nouvelle
Guinée.
En terminant je ne puis m’empécher d’exprimer le désir de
voir un ornithologiste publier une monographie compléte du
groupe si intéressant des Casoars, dont les espéces sont encore si mal
définies, et n’ont pas été complétement figurées jusqu’a ce jour.
4. On an apparently new Species of American Pipit.
By Francis Nicuozson, F.Z.S. &c.
[Received March 28, 1878.]
During a recent examination of the American Pipits in the British
Museum I came across two skins from Peru, labelled Anthus chit,
which appear to me to be specifically distinct from that bird; and I
therefore propose to name the species
ANTHUS PERUVIANUS, 0. sp.
A. Similis A. chii auct.(A.rufo, Gm.) sed dorsi plumis et tectricibus
alarum late albo nec fulvescenti marginatis ; corpore subtus albido
nec flavicante, et precipue caude colore distinguendus; rectricibus
duabus externis pallidé fumoso-brunneis, intus oblique saturalius
brunneis ad apicem ipsum albidis et versus apicem pogonii externi
saturatius brunnescentibus.
Long. tot. 5:1, culmen 0-5, ale 2°45, caude@ 1°75, tarsi 0°8.
Adult male. Lark-like. Above dark brown, the feathers exter-
1878.] MR. F. NICHOLSON ON A NEW AMERICAN PIPIT. 391
nally edged with dull sandy buff; the dorsal feathers dull whitish on
their inner webs, produci.g a distinctly varied appearance: scapu-
lars more uniform, the longer feathers with a subterminal shade of
dark brown, the edges and tips being dull sandy buff; lower back
and rump also rather more uniform than the rest of the back, only
slightly mottled with dark brown subterminal spots: tail-feathers
dark brown with nearly obsolete edges of sandy-buff, the two middle
feathers margined with ashy whitish ; the two outside feathers smoky
brown with a faintly indicated oblique mark of darker brown on the
inner web, the tips white, and the outer web distinctly darker brown
towards the tip: least wing-coverts like the back ; median and
greater series blackish brown, very broadly edged with dull whitish ;
primary-coverts and primaries dark brown, narrowly edged with
yellowish buff, the first long primary dull white for nearly the entire
length of the outer web; secondaries brown with a tiny white tip
and externally edged with dull white and sandy buff: lores and a
distinct eyebrow dull whitish ; ear-coverts brown, washed with sandy
buff, tipped with darker brown, with whitish shafts; cheeks and
under surface of body dull white, tinged with buff on the cheeks and
breast ; throat and abdomen uniform, the chest marked with trian-
gular spots of brown; a distinct moustachial streak of brown; sides
of body washed with fulvous brown and longitudinally streaked with
dark brown; thighs and under tail-coverts buffy white ; under wing-
coverts and axillaries white, the edge of the wing mottled with
tiny brown spots; quills brown below, whitish along the inner web ;
bill horn-brown, the under mandible paler ; legs waxy yellow.
Hab. Peru.
Syn. Anthus rufus, Sclater & Salvin, P. Z. S. 1868, p. 173.
Anthus chii, Taczanowski, P. Z.S. 1874, p. 506.
This species is of the same small size as the bird called in the
British Museum and by most ornithologists 4. chii, but more cor-
rectly by Messrs. Sclater and Salvin A. rufus. It is not so yellow
below ; and this isa point that strikes the observer at once. Oncom-
paring the upper surfaces the white edgings of the wing-coverts and
dorsal feathers in A. peruvianus are very noticeable. But the chief
difference lies in the outer tail-feathers, which are white in 4. chit,
smoky brown in A. peruvianus. The oblique mark of dark brown
on the inner web is distinct in the former, nearly obsolete in the
latter. Both species have a brown mark near the tip of the outer
web.
The typical specimens were collected by Mr. Henry Whitely in
Peru, the male in the Cataridon valley on the 12th of February 1867,
the female at Islay on the 17th of the same month.
392 MR. E. G. LODER ON BOS AMERICANUS. [Apr. 16,
' April 16, 1878.
E. W. H. Holdsworth, F. Z. 8., in the Chair.
Mr. Sclater exhibited a typical specimen of the new Fox lately
described by Mr. W. T. Blanford (J. A. S. B. xlvi. pt. ii. p. 321) as
Vulpes cana, from Baluchistan, which Mr. Blanford had forwarded to
him for examination. Mr. Blanford having stated (J. s. c.) that his
species appeared to be the same as the small Bushire Fox which in his
Zoology of Persia (‘ Eastern Persia,’ vol. ii. p. 41) he had referred
with doubt, after examining a young specimen living in the Society’s
Gardens, to Canis famelicus, Riipp., had requested Mr. Sclater to
investigate this point. But the comparison of the skin sent by Mr.
Blanford with the animal referred to by him (‘ List of Vert.’ 1877,
no. 213 4, p. 62), which was still living in the Society’s Gardens,
showed that the two animals must belong to quite different species.
Mr. Sclater was indebted to Mr. E. R. Alston for the subjoined
note upon these differences :—
“ Blanford’s Vulpes cana is quite different from your C. famelicus
at the Gardens, which is much. larger and lighter in colour. The fur
of the back is dark with light tips, forming a nearly uniform grey
grizzled with black on/y on the shoulders. The nape is like the
back ; and the outsides of the ears appear nearly black ; the limbs are
fulvous. The difference in size is shown in the following measure-
ments of the skeleton of the specimen of the same animal that died’,
taken to correspond with those given by Blanford.
inches
Length of skull from occipital crest.......... 5°45
Length of skull from foramen magnum ...... 5°0
Leneth of nasal wbones) piss ers) on sald s'ee Se op 1:80
Breadth; of. nagal portion |: 6 enero wt eloinwe ase escao
Breadth: of frontal, region pi)..3). << ow) aplyjal «eae
Breadth of btain-case’:. . 3.) |. -.:crdsseries ace) LOU
Breadth of zygomatic arches ........-..... 2°95
Length of palate)... sae ssrs <apaaayeuin> jhe apie 2°60
Beeadth,of palates: arositgai-iaah 44tuep ss Reese, Go
Length of last upper prem. ................ 52
Length. of first. molar. 6. 24.4). sd:sfeasis oe tas qh 940
Breadth of first molar ...... dae" sas phals Sapdtonnier "45
Lengthiot-luimerus -)uc, gant. <)ca ieivie wee sone 4:40
Rieneth .ofradis J tesaqttnay teem qarnarionsw 4 5:0
Rencehrot Tomiie hj. 2 tesla. stsiey emdicl done} ori 1s 4:90
Length of fibia,. went owir add cine i pts}. we
Mr. E. G. Loder, F.Z.S., exhibited a mounted head of the
peculiar variety of Bos americanus, called the ‘‘Rocky-Mountain
Bison” *, remarkable for its soft, dark and long hair on the forehead.
1 No 218 ¢ of ‘List. of Vert.’ (1877) p. 62.
? Allen’s Memoir on the American Bison’ pp. 39, 40,
avidson aelvet lth
BRACHIOPODA OF THE
‘LIGHTNING’ &" PORCUPINE’ EXPEDITIONS.
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BRACHIOPODA OF THE
"LIGHTNING & “PORCUPINE” EXPEDITIONS.
N LIN
1878.] DR. GWYN JEFFREYS ON MOLLUSCA. 393
This specimen had been obtained about 75 miles from Denver, Colo-
rado, in 1875.
The Secretary exhibited, on the part of Mr. A. Anderson, F.Z.S.,
a Bamboo driving-stick with leather thong attached, which had been
taken out of the nest of an Indian Fish-Eagle (Haliaétus leucory-
phus) in December 1876.
A communication was read from Prof. J. O. Westwood containing
a synopsis of the Uraniide, a family of Lepidopterous Insects, to-
gether with a complete monograph of Coronilia, one of the genera
of which it is composed.
This memoir will be published entire in the Society’s ‘‘ Transac-
tions.”
The following communications were read :—
1, On the Mollusca procured during the ‘ Lightning’ and
- Porcupine’ Expeditions, 1868-70. (Part I.) By J.
Gwyn Jerrreys, LL.D., F.R.S., F.Z.S.
[Received April 5, 1878. ]
(Plates XXII., XXIII.)
These expeditions were conducted under the auspices of the Royal
Society ; and preliminary Reports of them were published in the
‘Proceedings ’ of that Society for 1869 and 1870.
I subjoin lists of the dredging-stations, to which I shall have occa-
sion to refer in giving the habitats of the species hereafter described
or mentioned. The stations are extracted from the Reports and
accompanying Charts.
‘LIGHTNING’ ExpEpiTIon, 1868.
North of the Hebrides to the Faroe Isles.
No. of | No. in North West Depth in Bottom
Station. | Chart. | Latitude. Longitude. | Fathoms. |Temperature.
a bu aes, o Fahy.
ie VII. 60 10 5 59 550 32°8
2. IX. 60 24 6 38 170 41-7
3. x 60 28 6 55 500 33°2
4, XII. 59 36 ft 20) 530 47-3
5. XIII. 59 =~5 7 29 189 49-6
6. IIT. 60 31 9 18 229 48:0
i XVI. ol 2 12 4 650
8. Off the Faroe Isles. 164 & 208
The first column shows the stations at which dredgings were made,
394 DR. GWYN JEFFREYS ON MOLLUSCA OF THE _[Apr. 16,
and the second shows the corresponding stations in the chart; other
stations, at which soundings only were taken, are omitted.
‘ PorcuPINE’ ExpepiTion, 1869.
First Cruise.
Off the West of Ireland.
No. of North West Depth in Bottom
Station, Latitude. Longitude. Fathoms. | Temperature.
|
o 4 GO) fees oFahr,
dle 51 42 bk zit 85-110 49:0
Dingle Bay. 30-40
2 51 22 12. 25 808 41-4
3. 51 38 12 50 370-722 43:0
4 51 56 13 39 251-539 49°5
5 O22 7 12 52 364 48:8
6 52 25 11 40 90 50:0
7 52 14 ll 48 159 50:4
8 53 15 11 51 106 51-2
9 53 16 12 42 165 49-7
10 53 23 13 29 85 49:5
Galway Bay. 15-20
13. 53 42 13 55 - 208 49°6
14 53 49 13 15 173 49°6
15 bt 5 12 NE 422 47-0
16 54 19 11 50 816 39°5
17 54 28 ll 44 1230 37°8
18 54 15 Lea 183 49-4
Donegal Bay. 25-40
19. 54 538 10 56 1360 374
20. 55 11 ll 31 1443 370
21 55 40 12 46 1476 36°9
22, 56 8 13 34 1263 373
23. 06) ail 14 19 630 435
23.4 56 13 14 18 420 46-4
24 56 26 14. 28 109 46-4
25. 56 41 13 39 164 46°5
27. Rockall Bank. 54 48°3
28. 56 44 12 52 1215 371
30. 56 24 ll 49 1380 37:1
31. 56 15 Il 25 1360 37-2
Lough Swilly. 3-13
Lough Foyle. 10
North Channel. 40
Dredgings were made at nearly all the sounding-stations in this
cruise.
1878.] ‘LIGHTNING’ AND ‘PORCUPINE’ EXPEDITIONS. 395
Second Cruise.
Off the South of Ireland.
|
No. of North West Depth in Bottom
Station. Latitude. Longitude. Fathoms | Temperature.
i ie} U °
33. 50 38 9 27 74 49°6
34. 49 51 10 12 Wb 49°6
35. 496 10 -5¢ 96 51:3
36. 48 50 Te 29) 725 43°9
37. 47 38 12 8 2435 36°5
38. 47 39 iso 2090 363
39. 49 1 1l 56 557 47-0
40. 49 1 12° +5 517 47-7
41, 49 4 12 22 584 46:5
42. 49 12 12 52 862 39:7
43. 5OveL 12 26 1207 377
44, 50 20 11 34 865 39°4
45. rap | ll 21 458 48:1
Dredgings were made at all the sounding-stations in this cruise.
Third Cruise.
North of the Hebrides to the Faroe Isles.
4 fe] i °
47 59 34 i - AS 542 438
51 59 «54 7 52 355 46°2
52. 60 25 8 10 384 30°6
55. 60 4 Gr 19 605 29:8
56. 60 2 6, 11 480 30°7
57. 60 14 Gr 17 632 380°5
58. 60 21 6 51 540 30°8
60. 63 5 58 167 44:3
61. 62 1 5 19 114 45:0
62. 61 59 4 38 125 446
64 61 21 3 44 640 30:0
65 61 10 2 21 345 30°0
East Longitude.
68 60 23 0 75 44:0
69. 60 1 0 18 67 43:8
70 60 4 0 21 66 45-1
89 59° 38 [7 46 445 45°5
90. 59 41 7 34 458 45-2
The Minch. 60-80
Little Minch. 45-50
Near Belfast. 70
Loch Torridon. 40
Off Lerwick. 10-66
l
Not many deep-sea dredgings were made during the third cruise,
although soundings were taken at all the stations.
396 DR. GWYN JEFFREYS ON MOLLUSCA OF THE [Apr 16,
‘Porcupine’ ExPeEpDITION, 1870.
First Cruise.
Falmouth to Gibraltar.
No. of North West Depth in Bottom
Station. Latitude. Longitude. Fathoms. | Temperature.
fo] i i °o
12 43 10 15 567
2. 48 37 10 9 305 48°5
3. 48 31 10 3 690
3a. 500
6. 48 26 9 44 358 50°3
8. 48 13 9 il 257 50°0
9. 48 6 9 18 539 48:0
10. 42 44 9 23 81 531
11% 42 32 9 24 332 SES
12 42 20 oy 17 128-232 525
Vigo Bay. 20
Off Cape Mondego. 100-380
13; 40 16 CLES 220 52-0
14 40 6 9 44 469 51°5
16 39° 55 9 56 994 40'3
17 39 42 9 43 740-1095 39'7
17 a. 39 39 9 39 795 49°3
17 4. 785
19. 39! Q7 9 3d 248 StF,
Setubal Bay. 64
Off Cape Espichel. 740
22, 38 15 9330 718 52°0
24. 37 «19 OF 13 292 §2°7
25. 37) S11 9 374 535
Off Cape Sagres. 45-58
26. 36 44 & 8 364 52-7
27. 36 37 q 33 822 51:3
28. 36 29 in 16 304 53°3
28 a, 286
29, 36 20 6 47 227 550
30. 36 15 6 52 386 52°7
31. 35 56 ie 6 477 50°5
32 385 41 ty ue 651 50-0
33. 35 633 6 54 554 49-7
34 385 44 6 53 414 50:0
36 35 35 6 26 72-128 550
Tangier Bay. 30
Gibraltar Bay. 3
I dredged at nearly all the stations in this cruise.
1878.] ‘LIGHTNING’ AND ‘ PORCUPINE’ EXPEDITIONS. 397
Second Cruise.
Mediterranean.
No. of North West: Depth in Bottom
Station. Latitude. Longitude. Fathoms. | Temperature.
Algesiras Bay, Gibraltar. 1-15
fe) r] oO i oO
40. 36 s= 10) 4 40 586 | 55:0
41. 35) DT 4 12 730 550
45. 35 36 2 29 207 54:7
Capo de Gata. | 40-69
Cartagena Bay. | 60-84
“ie } Algerine Coast. { ackg
Off Jijeli. 40-80
E. Long
51. 36) "5b 1 10 1415 54-7
54. 387 «41 6 27 1508 | 55:0
55. 37 330 6 51 1456 | 55:0
| Gulf of Bona. 25
Benzert Road. 40-65
Rasel Amoush. 45
Gulf of Tunis. 25-85
Adventure Bank. 30-92 |
Off Rinaldo’s Chair. 60-160
58. 36 43 | 13 36 266 56°5
Very few productive deep-sea dredgings were made during this
second cruise.
Some of the new and peculiar species of Mollusca, which were
procured in these expeditions, have been already described and
noticed by me in the fifth volume of ‘British Conchology,’ the
* Proceedings of the Royal Society,’ and the ‘ Annals and Magazine
of Natural History.’ I will now endeavour to complete the work,
and to record all the species, with particulars of their geographical,
bathymetrical, and geological distribution. Figures of the more
remarkable species will also be given.
I commence with the class usually regarded as lowest in the scale
ot organization, or the least-specialized, among the Mollusca, viz.
the
BRACHIOPODA.
Notwithstanding the long and persevering labours of that monarch
of Brachiopodists, Mr. Davidson, as well as of Professor King, Mr.
Dall, and other excellent zoologists, the natural arrangement and
coordination of this difficult class cannot be said to be yet satisfac-
torily established. I prefer to steer a middle course between extreme
systematists, and not to follow Professor King in making Terebratula
cranium the type of another genus (his Macandrevia), nor to com-
press all the family of Terebratulide into the single genus Terebra-
398 DR. GWYN JEFFREYS ON MOLLUSCA OF THE [Apr. 16,
tula, as Mr. G. B. Sowerby, Junr., and Mr. Reeve have done. As
to Terebratulina and Waldheimia, Mr. Davidson some years ago
considered both subgenera of Terebratula; but he is now of opinion
that Waldheimia is a distinct genus, because (to use his own words)
‘the recent or living species are but a handful, in comparison with
the fossil species, and these last must be taken into account when
making divisions of the class, and it would bea very great advantage
to retain such divisional groups, which have so many distinguishing
characters, and are so numerous.” I fully admit the force of his
reasoning on the first mentioned ground, and that all fossil species
must be included with recent species in the same scheme of classifica-
tion ; but the latter ground, which depends on the validity of distine-
tive characters and the relative number of species, is not, in my opinion,
equally tenable. Itis notorious that Terebratulina and Waldheima
gradually pass one into another, as well as into the main or typical
genus Terebratula; and I contend, with great deference to Mr.
Davidson’s more experienced judgment, that it would be more ad-
vantageous to science not to multiply unnecessarily the generic
names, when fewer would answer every purpose. It cannot be pre-
tended that such names have any value beyond being mere symbols
or signs to distinguish and recognize certain groups; they are, and
must be, quite artificial and arbitrary. The question of number of
species in a genus does not appear to me of much importance. Take
for instance the genera Unio, Helix, and Bulimus, each of which,
even in a restricted sense, contains several hundred species. It may,
of course, be possible to divide these and other well-stocked genera
into more genera, each containing an equal number of species; and
the same method might be adcpted with respect to the species, and
even with respect to the individuals of every species. But that would
hardly be a scientific proceeding; and it might cause much con-
fusion or needless trouble. All genera ought to be equivalent
as regards distinctive characters, not as regards the number of
species.
I will now venture to suggest an arrangement of the few Brachio-
poda which are at present known to inhabit the European seas, and
which comprise the species procured in the ‘ Lightning’ and ‘ Por-
cupine’ Expeditions. The following Table gives not only the names
of the species and principal varieties, but also the range of depth
and the places where they have been found fossil in the Tertiary
formation.
1878.] ‘LIGHTNING’ AND ‘ PORCUPINE’ EXPEDITIONS. 399
BRACHIOPODA OF THE EUROPEAN SEAS.
>
Localities where
Names of species and :
fossil.
varieties.
| No. of Species.
‘ Lightning’ or
‘ Porcupine
Range of depth
in fathoms.
* Jointed.
A. Perforated or tubuliferous.
Family I. TEREBRATULIDA.
Genus I, TEREBRATULA.
Subgenus I. Terebratulina.
— |caput serpentis, Linné.............0000 0-1180 |Coralline Crag; Scotland;
Scandinavia ; Belgium ;
South Italy ; Azores.
gob
SSS?
— |tuberata, Jeffreys .......cccecseceececees 340-795
EPIOMA NCH: Cadesdacecess.saqdeseccens 500
Typical.
subquadrata, Jeff. ...cecsssessvcsvensee 500-600
sep SPVALECH,, BOM Mesattatenseteercscese) ccoceses 5-1456 |S. Italy.
var. minor, Philippi .......0.04+ 40-1000 |S. Italy ; Morea.
var. sphenoidea, Ph. ..........+- 100-994 |S. Italy.
Subgenus 2. Waldheimia.
LONOKE, IEP ene cece sts scamasiinaeiemereys 1450
ste ioraniunn, Deller! ola ccccvs Cte atectases 5-690 |Scandinavia; S. Italy.
=) | Septata,) Phar de issccsvescecdvceststrocsens 64-725 |Norway; S. Italy;
Rhodes.
Genus IT. TeReBRATELLA.
spitzbergensis, Davidson .........000005 20-690 |Scandinayia; Canada.
Genus III. Areropn.
Typical.
— | decollata, Chemmitz......ccccccecesseees 18-364 |S. W. France; Austria;
Italy ; Malta; Rhodes.
Subgenus 1. Cistella.
neapolitana, Scacchi ..eccssecseeeveees 30-130 |S. W. France; 8. Italy ;
Rhodes ; Austria.
cistellula,.S. V. Wood. ..........0..000+- 20-45 |Coralline Crag; Austria;
Sicily ; Rhodes.
cuneata, Piss .....serseserrasgneesereee 28-200 |Austria; S. Italy; Rhodes.
Subgenus 2. Gwynia.
Capa, SEPT ahs «dais sneglsaieese ssVeaxei 8-25 |Norway.
Genus IV. Puatypia.
=— | |aflOMIOld ey SCG? EM. v.ctcp eect essats oc 40-600 |Sicily ; Austria.
Genus V. Mrceriia.
eA n ICE beat L1G Sep Gaiters seasons sae seee ee 20-292 |Italy; Austria.
var. monstruosa, Sc, ..........4. 20-292 |Sicily.
400 DR. GWYN JEFFREYS ON MOLLUSCA OF THE [Apr. 16,
BracuHriopopa OF THE European Szas (continued).
| No. of Species.
| el
ma
ae fos
og | Bios
"er rae
oe 5 S sys
a2 Names of species and Sa Localities where
aE varieties, 2 fossil.
ae a
5 ra
Genus VI. TurcmEa.
mediterranea, R7SSO .........cceceeee eens 30-300 Turin ; Sicily.
B. Imperforate.
Family Il. RHYNCHONELLID/.
Genus I, Arrerta.
1S hacTrar || PROMI MALT. saecs.~d-abimwen acess. amp] 650-1443
Genus II. Ruyncnone.ua.
19, psittacea (rostrum-psittaci), C2. .... 10-690 |G. Britain and Ireland ;
| Scandinavia ; Canada.
ZO" ‘Netculay SEG wEnZG © asses eseesarvceseeeees 690 Sicily.
**Hingeless. |
Family I. CRANIIDA.
Genus Cranta.
"ammonia, MRE, .Jocitebss benaccinocece= see 3-808 Sicily.
|
| Family II. DISCINID®.
| Genus Discrva.
| .— atlantica, Kimg ......-.ceeessserre 690-2400 Coralline Orag.
| |
Two new and remarkable species of Terebratula (viz. T. trigona
and 7. subquadrata) were dredged by Mr. Saville Kent in 1871 off
the coast of Portugal ; and these species I propose to describe and
figure, because my dredgings in the ‘ Porcupine’ Expedition of 1870
were nearly on the same ground. Mr. Kent’s discovery confirms my
previous statement and unchanged impression, that we know ver
little of the deep-water fauna, and that it is an inexhaustible field of
research. Terebratula tenera, from the ‘ Valorous’ Expedition, is
also figured.
I will notice very briefly the other Brachiopods of the European
seas, which were not procured in the ‘ Lightning’ or ‘ Porcupine’
Expedition.
For the illustrations accompanying this paper, and for other valu-
able assistance, I am indebted to my friend Mr. Davidson.
1878.] ‘LIGHTNING’ AND ‘ PORCUPINE’ EXPEDITIONS. 401
* Jointed.
A. Perforated or tubuliferous.
Family I. TerReEsRATULID2.
Genus I. TEREBRATULA.
Subgenus I. Terebratulina. Loop, short, annular in its adult state.
1, TEREBRATULA CAPUT-SERPENTIS, Linné, (Plate XXII. fig. 1.)
Anomia caputserpentis, L. S. N. ed. 12, p. 1153: T'. caput-serpen-
tis, ‘ British Conchology,’ ii. p. 14, pl.i. f. 1; v. p. 164, pl. xix., f. 2.
‘ Lightning’ Expedition: Station 4, 530 fathoms.
‘ Poreupine’ Exp., 1869: St. 2, 808 f., on Oculina prolifera ;
6, 90 f.; 14,173 f.; 16, 1180 f.; 23a, 420f. ; 25, 164 f.; 54, 363 f.;
57, 632 f.; 61,114 f.; 65, 345 f.; 74 and 75, 203-250 f. ; 77, 560
f.; 78, 290 f.; 84, 155 f.; Loch Torridon, 40 f.; off Belfast, 70 f.
1870: Atlantic, 24,292 f.; Mediterranean; Adventure Bank,
92 f.; off Rinaldo’s Chair, 60-160 f.
Spitzbergen and Davis Strait to Malta and the Adriatic; Jamaica ;
North-East America ; Japan ; Korea; Australia; New Zealand: shore
to 150 f.
Extremely variable with respect to the proportional length of the
shell and the number of ribs or strize. One variety is the 7’. septen-
trionalis of Couthouy; and another may be called mediterranea,
which is longer and more slender, more compressed or flatter than
usual, and more or less cloven in front. In a monstrosity from the
Adventure Bank the upper valve is extraordinarily convex and boat-
shaped, and there are no longitudinal ribs or striz on the outer
margin.
I have figured (Pl. XXII. f. 1) the interior of a young specimen to
show that the loop is not in that stage of growth completely annular
or closed at the top, which is the principal characteristic of the sub-
genus Terebratulina.
The synonyms are very numerous; I have noted seventeen.
The localities for this and other Brachiopods in a fossil state will
be found in the table of species.
2. TEREBRATULA TUBERATA', Jeffr. (Plate XXII. fig. 2.)
SHELL, upper valve ovately triangular, somewhat convex ; lower
valve squarish, compressed; it is solid, opaque, and lustreless :
sculpture, about 20 fine longitudinal ribs, besides a few intermediate
strize, all of which radiate from the beak, and are closely covered
with short tubercles, some of which at the sides are raised and
become prickly: cecal tubuli very minute and numerous: colour
brownish white: margins of upper valve in front rounded and
slightly scalloped by the ribs, at the sides sloping with a gentle
curve, behind acute-angled : beak pointed : foramen or byssal pas-
sage oval, incomplete: del¢idium broad, slightly excavated or con-
cave: hinge-plate thick: teeth in upper valve small, strong, and
2 Covered with prickles.
Proc. Zoou. Soc.—1878, No. XXVI. 26
402 DR. GWYN JEFFREYS ON MOLLUSCA OF THE [Apr. 16,
triangular : skeleton or apophysary process in lower valve consisting
of a simple loop, which is attached to the hinge by short and stout
crura, ending in proportionally large triangular upright points or
spurs, from which proceeds the obtusely angular or nearly semicir-
cular portion of the loop at the base; the outer surface of the crura
is minutely and closely ridged across: inside furnished with about
10 ribs, but not with any intermediate strize ; edge notched to corre-
spond with the furrows. L. 0:1, B. 0°095.
‘Porcupine’ Exp., 1870: St. 17a, 795 f. Asingle (upper) valve
only.
I have also, through the kindness of Professor Lovén, a perfect
specimen from the Josephine Bank off the Strait of Gibraltar, which
was dredged in 340-430 fathoms.
The peculiar shape and sculpture serve to distinguish this tiny
species from the young of 7. caput-serpentis, and from the next
species 7’. trigona.
3. TEREBRATULA TRIGONA|, Jeffreys. (Plate XXII. fig 3.)
SHELL triangular, much compressed, solid for its size, semi-
transparent between the ribs, and glossy: sculpture, 8 rounded
longitudinal ribs, which radiate from the beak, and become gradually
broader towards the front ; those in the middle of each valve are the
largest ; the ribs are strongly wrinkled across by the lines of growth ;
the interstices or furrows are narrow: cecal tubuli numerous and
close-set: colour clear white: margins gently curved and scalloped
in front, abruptly sloping at the sides, broad or having somewhat
ear-like expansions behind: Jeak short : foramen oval and propor-
tionally large, incomplete: deltidium small: hinge-plate thick :
teeth in the upper valve short and strong: skeleton in the lower
valve consisting of a semicircular loop, which is attached on each
side by short and strong crura, and terminate in small pointed spurs:
inside of both valves smooth and polished, exhibiting through the
shell the thick outside ribs. L. 0:1, B. 0-1.
A single living specimen was dredged by Mr. Kent, in Mr. Mar-
shall Hall’s yacht the ‘ Norna,’ off the coast of Portugal, in about
500 fathoms. He very obligingly gave it to me with a specimen of
the following species. It may perhaps be immature; but I cannot
identify it with the young of any other species.
This pretty little shell differs from the young of 7. caput-serpen-
tis in shape, colour, having fewer ribs and a smooth interior, and in
the loop being rounded instead of incurved at the base. In T.
caput-serpentis of the same size as this (Pl. XXII. fig. 1) the inside,
especially in front, has strong and conspicuous ridges, which become
notches in the full-grown shell.
Typical. Loop short, semiannular.
4, TEREBRATULA SUBQUADRATA’, Jeffreys. (Plate XXII. fig. 4.)
SHE tt broadly triangular or oval, with a squarish outline ; upper
Triangular, ? Somewhat square.
1878.] ‘LigHrNING’ AND ‘PORCUPINE’ EXPEDITIONS. 403
valve gibbous ; lower valve somewhat compressed ; it is rather solid,
Opaque, and lustreless: sculpture, slight and fine longitudinal strize,
which radiate from the beak (50 to 60 in a larger, and half that
number in a smaller Specimen) ; these are crowded at the sides, and
are occasionally or partially intervenient ; concentric marks of growth
distinct, besides humerous intermediate and irregular lines: eweal
tubuli white, minute, and close-set: colour white, with an ochreous
tinge: margins curved in front, or slightly indented there in the
middle of the upper valve, more or less truncated at the sides,
rounded and somewhat shouldered behind in the lower valve: beak
extended or produced : Soramen large, circular, complete, thickened,
and reflected at the lower side: deléidium long and narrow, slightly
grooved across: hinge-plate thick and broad: teeth in the upper
valve short and incurved : loop narrow, keyhole-shaped, attached to
the hinge by short crura, which terminate halfway in small blunt
spurs, obtusely angular below in the smaller, and flexuous in the
larger specimen: inside smooth, slightly grooved lengthwise in the
smaller specimen, to correspond with the outside striz, and having
the inner edges of the larger specimen thickened and notched or
crenate, as in Astarte sulcata and in many species of Venus, Nucula,
and other genera. L, 1:05, B. 1 (larger specimen) ; L. 0°65,
B. 0°55 (smaller Specimen ).
Hab. Off the coast of Portugal, in 500-600 fathoms (Kent).
Of this new and interesting species Mr. Kent dredged two speci-
mens and a couple of valves, which he presented to Mr. Davidson
and myself.
I at first thought it might be the 7’. orbiculata of Seguenza
(Mem. Soe. Ital. Sc. Nat. t. i. no. 4, 1865, p. 7, tav. ii. f. 14—
16), judging from the description and figures; but he has lately,
with his usual courtesy, sent me for inspection the type of his species.
This shows that the smaller valve is not shouldered (so as to give
the shell a squarish outline), and that the strize are much finer and
more numerous than in 7’. subguadrata. He, indeed, now suspects
that his species may be the young of a large species, such as 77
ampulla or T. grandis; but that Species is not striated lengthwise.
I should be inclined to refer Seguenza’s species to the variety sphe-
notdea of T. vitrea. The form of the skeleton or apophysis depends
to some extent on the shape of the specimen in a species so variable
as T. vitrea.
¥ 5. TEREBRATULA viTREA, Born. (Plate XXII. fig. 5.)
Anomia vitrea, Born, Test. Mus. Czes, Vind. p. 104,
‘Porcupine’ Exp., 1870: Atlantic, St. 13, 220 f.; 16, 994 f, ;
17a, 740 f.; 24, 292 Ff. ; 25, 374 f.; off Cape Sagres, 45-58 f, ;
26, 364 f.; 27, 322 £5 28,8041); 29,'227. £3; Mediterranean,
45, 207 f.; Capo de Gata, 40-69 f.; 55, 1456 f. ; Adventure Bank,
92 f.
Atlantic coast of Spain to the Adriatic and fEigean : depths
5-310 f.
Bopy clear white and gelatinous: mantle lining the valves, and
26*
404 DR. GWYN JEFFREYS @N MOLLUSCA OF THE [Apr. 16,
furnished at its edges with short and very delicate close-set cilia of
different lengths and sizes, which project somewhat beyond the
valves, as in 7’. caput-serpentis.
The tubercles which close the tubular perforations in the shell
are extremely small and numerous. In some large and unusually
thick valves dredged by Dr. Carpenter in H.M.S. ‘Shearwater,’ off
the coast of Tunis, the perforations are irregular or unsymmetrical,
as in 7’. grandis.
Var. 1. minor, Philippi, =T. affinis, Calcara.
‘ Porcupine’ Exp., 1X70: Mediterranean, St. 58, 266 f.
Off Jan Mayen’s Isle, 263 f. (Friele, as T. arctica) ; Mediter-
ranean, Adriatic, and Agean, 30-310 f.; ‘Challenger’ Exp., off
Cape of Good Hope, 150 f., and Azores, 1000 f. ; ‘ Josephine’ Exp.,
off Villa Franca, Azores, 320-600 f.; Japan, 55 f. (A. Adams, as
T. davidson).
In a specimen of this variety from the Bay of Naples the front
margin is cloven or deeply indented, apparently in consequence of
the mantle having been injured in that part.
Var. 2. sphenotdea, Philippi. (Plate XXII. fig. 6.)
‘Porcupine’ Exp., 1870: Atlantic, St. 16, 994 f., 24, 292 f.;
25, 374 f.
Gulf of Florida, 100-270 f. (Pourtales, as T. cubensis).
After a protracted and very careful examination of my specimens,
which I had considered the 7’. sphenoidea of Philippi, and having
compared them with fossil specimens sent me by Professor Seguenza
as Philippi’s species from the Sicilian Tertiaries, as well as with a
series of 7. cubensis which I received from Count Pourtales and Pro-
fessor Alexander Agassiz, and also after a close comparison of all these
specimens with the description and figures given by Philippi,
Seguenza, Pourtales, and Dall, Iam convinced that 7’. sphenoidea
and 7’. cubensis are the same, and constitute a well-marked variety
of T. vitrea. The loop in 7’. sphengidea and T. cubensis is precisely
similar. The shape and proportions of the loop depend on the
shape of the shell in 7’. vitrea, and, I believe, in every other species
of Terebratula. Some recent and fossil specimens of this variety
exhibit more or less distinct, although slight, longitudinal striz,
which radiate from the beak and are especially visible at the sides ;
and the same kind of sculpture is occasionally observable in speci-
mens of the typical form of 7”. vitrea.
As to synonyms, I cannot recognize any valid character that would
serve to distinguish Seguenza’s Sicilian Tertiary species, 7”. elliptica
and 7. michelottiana from T.. vitrea, or his 7’. miocenica, T. orbicu-
lata, and 7’. benoitiana from the variety sphenoidea. TT. manticula
of Fischer (Journ. Conch. t. xvii. p. 81, pl. iii. fig. 4), a Miocene
fossil of the south-west of France, seems to be closely allied to the
variety minor, the only difference being that the deltidium in the
fossil is larger. 7. cernica of Crosse, from the Mauritius (Journ.
Conch. t. xxi. p. 285, and xxii. p. 75, pl. i. fig. 3), may be also the
present nearly ubiquitous species.
Costa, in his ‘Fauna del Regno di Napoli,’ refers the T.
1878.] ‘LIGHTNING’ AND ‘PORCUPINE’ EXPEDITIONS. 405
sphenoidea of Philippi to the variety ¢rigona of his own 7’. irregu-
laris, which is represented as having the skeleton of 7. septata or
some other septigerous species of the subgenus Waldheimia.
Subgenus II. Waldheimia. Loop long, reflected.
6. TeREBRATULA TENERA, Jeffreys. (Plate XXII. fig. 7.)
T. tenera, Jeffr. in Ann. & Mag. N. H., Sept. 1876, p. 250.
*Valorous’ Exp.’ 1450 f.
7. TEREBRATULA CRANIUM, Miiller.
LT. cranium, Mill. Zool. Dan. Prodr. p. 249, no. 3006: Br.
Conch. ii. p. 11; v. p. 163, pl. xix. fig. 1, la.
‘ Lightning’ Exp.: St. 2, 170 f.; 4, 530 f.; 5, 189 f.; 7, 650f. ;
off the Farves, 164 and 208 f,.
‘Porcupine’ Exp., 1869: St. 14, 173f.; 15, 422 f. ; 23a, 420 f. ;
25, 164 f. ; 46,374 f.; 50, 355 f. 357, 632 f.; 61, 114 f.; 65,345f.;
74, 203 f.; 75, 250 f.; 78, 290 f.; 83, 262 f. ; 84, 155 f.; 85, 190
f. 1870: St. 1, 567 f.; 2, 305 f.; 3, 690 f. (and var. oblonga) ;
Vigo Bay, 30 f.; 24, 292 f.
Greenland (Wallich, Mobius), and Norway (Miiller and others),
to the south-west of France (De Folin, Fischer), Vigo (McAndrew) :
5-650 f. St. Margaret’s Bay, Nova Scotia (Willis)? Northern
Asia and Japan (A. Adams),
The tubercles or czcal extremities of the canals which permeate
the shell are more numerous in this than in the next species (a
septata), and are arranged in wavy lines. The gradual develop-
ment of the skeleton or apophysary process in both species is very
remarkable, and has been admirably shown by Herr Friele of Bergen
in a series of figures which accompany his papers on the subject in
the ‘ Vidensk. Selsk. Forh.’ for 1875, pl. i. fig. 9, a, and * Archiv
for Mathematik og Naturvidenskab’ for 1877, tab. i.—iii. Mr. Charles
Moore had also, in ‘the Geologist’ for 1860, described and illus-
trated certain modifications in the loop of a fossil species, Terebra-
tella buckmanni, which are not less important in respect of the
classification of the Brachiopoda. The skeleton of a young Megerlia
truncata, and probably of every other jointed Brachiopod, is different
from that of an adult specimen.
In the second volume of iny work on British Conchology, p- 14,
I stated that the lower valve in the young of 7. cranium is “ fur-
uished with a very distinct and prominent crest or ridge, placed
inside and nearly in the middle of this valve,’ and that the same
character “ likewise occurs in 7. septata, Philippi, a Sicilian fossil
(2. septigera, Lovén), and is remarkably developed in that species ;
but the foramen is incomplete in 7. cranium, and entire in T. sep-
tata.’ I now find, on further examination of my Shetland speci-
mens, that some of those which I had taken to be the young of 7’.
cranium really belong to 7’. septata; and consequently the latter
species is an inhabitant of our own coasts as well as of the Norwegian
406 DR. GWYN JEFFREYS ON MOLLUSCA OF THE [<Apr. 16,
and other seas. The septum in the very young of 7’. cranium consists
of a mere upright point, as in Megerlia; and the loop is then round
and small, becoming elliptical and extended in the adult. The
foramen is incomplete in the young of 7. septata, and resembles
that of 7’. cranium in every state ; and this is likewise the case as to
the young of 7’. vitrea.
Mr. Dall, in the ‘ American Naturalist’ for March 1871 (vol. v.
p- 55), indicated a new genus of Brachiopods, founded on some
young specimens of 7’. cranium which I had sent him from the
‘Porcupine’ dredgingsin 1869. He said the loop “ has no secondary
attachment to the hzmal valve, and the latter is destitute of a
septum.” This genus he proposed to name Frenula, and the
species jeffreysiz. Inthe American Journal of Conchology for 1871
(vol. vii. pt. 2, p. 65, pl. ii. figs. 7-10) he gave a more detailed
account, with figures, of the same species as [smenia? jeffreyst.
His figures agree with those published by Herr Friele to illustrate
the development of the skeleton in 7. cranium. Mr. Dall remarks
as to his species, ‘‘ Without a septum in either valve. Loop un-
attached except by the hzemal processes to the hinge plate,’ and
further on, “ None of the specimens had the slightest trace of a
septum.” He then alludes to having subsequently received from me
other young Brachiopods, of which he says, ‘ All of them possessed
a filiform but rather stout septum;” and, although with some
doubt, he was of opinion that all the specimens belonged to the same
species, and that in those first mentioned ‘the septum must have
been broken away.” He has since, at my request, kindly sent me
for my inspection a specimen of his species, which he now calls
Megerlia jeffreysi, and which he says he received from me as the
young of 7’. cranium; and he has presented me with some valves
dredged by him at Port Etches in Alaska, at a depth of about 15
fathoms, and which according to him were “probably” the same
species, although he carefully explained that he would not vouch
for it. The valves in question were worn and apparently old, but
all of them showed the septum most distinctly; and they may
belong to the species which Mr. Dall has very lately described anew
as Megerlia jeffreysi. These valves are not unlike Terebratella (?)
frontalis of Middendorff in shape and texture; but they differ
essentially in the position of the septum as well as in the deltidium.
In Mr. Dall’s imperfect specimens the septum resembles that of
Terebratella spitzbergensis, viz. in being a long narrow and thread-
like lamina extending from the middle of the deltidium for about
two thirds of the inside towards the front ; in 7. frontalis (judging
from the type most obligingly sent me by Dr. v. Schrenck from the
St. Petersburg Museum for my inspection) the septum is that of
Argiope, being gnomon-shaped or triangular, and placed near the
front. The apophysis of Megerlia truncata does not correspond
with that of any of the above-named species. I would here remark
that the septum is not at all likely to be broken away and entirely
disappear, because it is formed in the protected part of the lower
valve. It is persistent in the most worn and even fragmentary spe-
1878.) ‘Ligutning’ AND ‘PORCUPINE’ EXPEDITIONS. 407
cimens of 7’. septata, Terebratella spitzbergensis, and T. (?) frontalis,
and of other similarly furnished Brachiopods. Mr. Dall has now
redescribed Megerlia Jeffreysi in his ‘ Scientific Results of the Ex-
ploration of Alaska’ (June 1877, p. 48), and Says as to the apophysis
“ Megerlie sanguinec simillima,” the habitat being thus given,
ON. E. Atlantie, 155-345 fathoms, Jeffreys, with Waldheimia
cranium, Semidi Islands, Port Etches, Dall ; Victoria, V. Id, J.
Richardson.” Mr. Davidson tells me that Herr Friele was mistaken
im saying that the former considered M. jeffreysi the young of
Megerlia sanguinea.
Now I fear there must have been some mistake about the Atlantic
and Pacific specimens. I have no doubt that the shell described
last year by Mr, Dall as Megerlia jeffreysi (see Plate XXIII. fig. 3),
and which by his permission I have examined, is a perfectly good
Species of Megerlia, having a distinct Septum and the loop three
times attached; and I thank him sincerely for the compliment he
as paid me in associating my name with it. But at the same time
I firmly believe that the specimens described by him in 1871 as
Frenula jeffreysii and Ismenia? Jeff reysi belong to a different species
and another genus, and that such last-mentioned species was 7’.
cranium. The other specimens which I afterwards sent him, and
which had a “rather stout septum,” may have been the young of
T. septata.
Synonyms: Anomia terebratula, L. ; Terebratula plicata, Philipp-
son; T. euthyra, Philippi; 7. subvitrea and 7. glabra, Leach ; and
perhaps, in the younger state, Waldheimia davidsoniana of Seguenza.
Mr. Davidson in 1853 and 1855 placed Waldheimia as a sub-
genus of Terebratula ; but in 1861 he raised it to generic rank.
v 8. TeREBRATULA sEprata, Philippi. (Plate XXIII. figs. 1,
1 a, 6, ¢; loop.)
T. septata, Ph. Moll. Sic. ii. p- 68, t. xviii. fig. 7 (1844).
* Porcupine’ Exp., 1869: St. 23a, 420 f. ; 36, 725 f.; 39, 557 f.;
54, 363 f.: 65,345 f.; 67, 64 f.; 68, 75 f.; 74, 203f.; 75,250 f. 1870:
1, 567 f.; 3,690 f. ; 6, 358 f.; 9,539 f. A perfect cast or petrifaction
of apparently this species was also dredged by me, with other Ter-
tiary fossils, at Station 24, 292 f., off the Atlantic coast of Spain,
Norway, 100-300 f. (Lovén and others) ; Shetland, 80-90 fz,
young, with 7. cranium (J. G. J ia
A Japanese Brachiopod, Terebratella marie of A. Adams, may
be a dwarf variety of the present species, corresponding with the
variety minor of T\. vitrea. Waldheimia raphaélis of Dall, also
from the North Pacific, looks like a gigantic variety of 7. septata,
the author having kindly permitted myself and Mr. Davidson to
compare his specimen of W. raphaélis with some of T. septata, my
largest specimen of which measures an inch and three tenths in
length.
I cannot detect any distinctive character of importance in Wald-
heimia floridana of Pourtales, from the Gulf of Florida, 100-200
fathoms, to separate it specifically from 7. septata. He says (Bull.
408 DR. GWYN JEFFREYS ON MOLLUSCA OF THE [Apr. 16,
Mus. Comp. Zool. No. 7, 1869, p. 127), ‘* Very young specimens are
flatter, rounded, and have a strait margin; they could scarcely be
distinguished from the young of Terebratula cubensis, if it was not
for the loop and septum seen by transparency. There is also some
variety of form in the old; in some specimens the length is greater
than the breadth, and there is considerable diversity in the sinuosity
of the frontal margin.” 7’. septata is not less variable in shape and
in the flexuosity of the front margin; a young specimen which I
dredged in the ‘ Porcupine’ Expedition of 1870, attached to a branch
of Oculina prolifera, was mistaken by me for Terebratella spitzber-
gensis, having the same oval and laterally compressed outline, and
a similar septum visible through the shell.
In young and half-grown specimens of 7. septata, as well as in
other allied species, the side flanges or flaps of the loop are closed
by a membrane, which is removable; the openings then appear.
The inner layer of the shell is microscopically and closely striated
lengthwise. The gradual development of the skeleton or apophy-
sary process, indicated by the changes which take place in the course
of growth, is very remarkable, and is shown in the accompanying
figures (Plate XXIII. figs. 1 a, 6, ¢). Mr. Davidson and I agree
that the loop in 7. septata, when quite young, resembles that of a
Megerlia, and is three times attached; at a rather more advanced
age it assumes the form of a 7'eredratella, and is twice attached; in
the full-grown state it has all the characteristics of the subgenus
Waldheimia, and is attached only to the hinge-plate. Mr. Friele
has also, to some extent, demonstrated these retrograde changes
in his papers above referred to.
This species is unquestionably the 7’. septigera of Loven (1846),
and equally without doubt (so far as my opinion is worth any thing
in either case) the 7’. septata of Philippi, 1844. Seguenza, how-
ever, considers Philippi’s species to belong to the genus T'erebratella ;
and he has named Lovén’s species Waldheimia peloritana. Although
the skeleton represented in Philippi’s figure was imperfect, there is
no appearance of the cross bar to which the loop is attached in
Terebratella, and which cross bar is not less persistent than the sep-
tum in broken and worn specimens of so many species of that genus
as well as of Megerlia. Indeed his description and figures agree as
closely with imperfect specimens of 7. septigera, as those of his 7’.
euthyra with imperfect specimens of 7’. cranium. Seguenza may
have found in the Sicilian Tertiaries not only his W. peloritana,
which he now refers to 7’. septigera, but also a species of Terebra-
tella which he regards as Philippi’s species. For this last-mentioned
species, if not known in a recent or living state, he might give
another name. Were not priority of publication an essential stand-
point for scientific nomenclature, septigera might be retained for the
recent species, and septata for Seguenza’s fossil species of Terebra-
tella ; but in the absence of better data it may be more advisable
*‘qnieta non moveri.” W. peloritana and W. floridana appear to
be the same variety of 7’. septata.
I cannot refrain from expressing my admiration of the careful and
1878.] ‘LIGHTNING’ AND ‘ PORCUPINE’ EXPEDITIONS. 409
valuable work of Professor Seguenza, notwithstanding that my notions
of the distinctive character of certain species do not accord with
his.
Genus IT. TEREBRATELLA.
TEREBRATELLA SPITZBERGENSIS, Davidson, (Plate XXIII.
fig. 2; loop.)
Terebratella spitzbergensis, Dav. in Ann. & Mag. N.H. 1855,
xvi. p. 442, pl. x. fig. 3: B. OC. ii. p. 2; v. p. 164, pl. xcix. fig. 3.
Spitzbergen, Iceland; ‘ Valorous’ Expedition, Wellington Channel,
Davis Strait, Gulf of St. Lawrence, North Japan, and Shetland :
depths 20-690 fathoms,
Genus III. Arciopr.
Typical. Having three septa.
“ 1. ArRGIoPE DECOLLATA, Chemnitz.
Anomia decollata, Ch. Conch. Cab. yiij. p. 96, pl. 78. figs.
705 a-d.
Argiope decollata, B. C. ii. p. 18; v. p. 164, pl. xix. fig. 3.
‘ Porcupine’ Exp., 1870: Atlantic, St. 24, Qua f.; 36, 364 f.
Mediterranean, 58, 50 f.; Adventure Bank, 92 f.
Guernsey, South-west of F rance, Atlantic coasts of Spain, Medi-
terranean, Adriatic, Egean, Madeira, Canary Isles: 18-130 f.
In some Specimens, recent and fossil, the ribs are evanescent or
Searcely perceptible. Size variable.
Synonyms: Anomia detruncata, Gmelin; Terebratula aperta, De
Blainville; Z. cardita and Vis urna-antiqua, Risso; T. ungula,
Retz; 7. pectiniformis, O. G. Costa; 7. dimidiata, Scacchi,
Subgenus Cistella. With a Single septum.
2. ARGIOPE NEAPOLITANA, Scacchi.
Terebratula neapolitana, Sc. Oss. Zool. 1833, ii. p. 18,
Mediterranean, Adriatic, Egean, and Canary Isles: 30-320 f.
The number of ribs varies from two to eight, some specimens
being quite ribless; and the shape and size are also very variable
characters, as in 4. cuneata. From that species 4. neapolitana
may be distinguished by the inner margin in the latter being fur-
nished with a numerous row of small and short ridges, which become
tubercular in front, by the anterior margin being indented in the
. middle, and by the septum in the lower valve being notched across,
In some Specimens, which are more transparent than usual, the
marginal ridges can be seen outside.
Philippi has, in the figures given in his first volume, evidently
confounded the present species (his Terebratula seminulum) with
Platydia anomioides, which it seems he did not then know.
Terebratula cordata, Risso; T, seminulum, Ph.; Orthis bifida,
Costa; A. forbesi, Davidson.
410 DR. GWYN JEFFREYS ON MOLLUSCA OF THE [Apr. 16,
3. ARGIOPE CISTELLULA, 8. V. Wood.
Terebratula cistellula, S. V. Wood, in Ann. & Mag. N. H. vi,
p- 293.
A. cistellula, B. C. ii. p. 19, pl. 1. f. 2; v. p. 164, pl. xix. f. 4.
Norway, Shetland to Guernsey, Ireland, Normandy, S.W. France :
20-45 f.
I was wrong in referring this species to the Orthis lunifera of
Philippi, which is (at any rate partly) the young of Megerlia trun-
cata. I likewise mistrust the Mediterranean habitat cited by me for
a specimen of the present species, and have now omitted it ; the spe-
cimen was more probably the young of 4. cuneata.
4. ARGIOPE CUNEATA, Risso.
Terebratula cuneata, Risso, Hist. Nat. de l’ Eur. mérid. iv. p. 388,
fig. 179.
Mediterranean, Adriatic, Augean, and Canary Isles: 28-200 f.
Probably also Guadeloupe, 111-150 f. (Schramm, fide Crosse and
Fischer as A. schrammt).
The shape varies from triangular to broadly oblong; ribs two to
ten, more or less vanishing in some specimens, and altogether want-
ing in others.
I have been favoured by Professor Seguenza with an opportunity
of examining his unique specimen of A. biplicata from the Straits of
Messina, and of comparing it with an extensive series of A. cuneata
and its colourless variety. The result of such examination and com-
parison, as regards both the outside and inside, compels me to con-
sider the specimen an abnormal form of A. cuneata.
Terebratula soldaniana, Risso ; Anomia pera, v. Mihlfeld ; T. sco-
binata, Cantraine, not -Anomia scobinata of Linné.
Subgenus Gwynia. No septum.
5. ArGIorE capsuLa, Jeffreys.
Terebratula capsula, Jeffr. in Ann. & Mag. N. H. ser. 3, 11. p. 125,
pl. v. f..4; iii. pl. ii. f. 7, 8.
A. capsula, B.C. ii. p. 21; v. p. 164, pl. xix. f. 5.
East and south of Ireland, Plymouth, Weymouth, Guernsey,
Etretat ; Sluys-kill, Zealand, near the Belgian frontier (Colbeau)? :
8-25 f.
Ihave lately opened and carefully examined more specimens of this
minute Brachiopod, as well as very young specimens of Terebratula
caput-serpentis, T. cranium, T. vitrea, Argiope decollata, A. neapo- .
litana, A. cistellula, A. cuneata, Platydia anomioides, and Megerlia
truncata, of nearly similar dimensions. All of these last are cer-
tainly distinct from A. capsula in respect of the internal apparatus or
skeleton, which is nearly wanting in A. capsula. The young of
Platydia anomioides has, besides the septum, a very large byssal
orifice in the lower valve; otherwise that species might be the adult
of A. capsula: but it has never been found with the latter species,
1878.] ‘LIGHTNING’ AND ‘PoRCUPINE’ EXPEDITIONS. 411
nor even in the same parts of the ocean. It is clearly unscientific to
suppose that, because 4. capsula is tiny, it cannot be a good species,
in whatever genus it ought to be placed. Until there is some proof
or fair presumption that it is embryonic or immature, it would be
preferable to retain it in the subgenus of Argiope, for which Dr. King
proposed the generic name Gwynia,
Genus IV. Prarypra.
¥ Puatyp1a anomroipzs, Scacchi and Philippi.
Orthis anomioides, Sc. & Ph., Moll. Sie. ii. p. 69, t. xviii. f. 9.
‘ Porcupine’ Exp., 1869.: St. 78,290 f. One of my specimens is
twice the length of any I have seen from the Mediterranean ; the
other is minute, of an oval shape, and apparently abnormal.
Coast of Portugal, 600 f. (Kent)!; Mediterranean and /Egean,
40-120 f.; Gulf of Florida, 237 f. (Dall) ; ‘ Challenger ’ Exp.,
“attached to a specimen of Wauldheimia kerguelensis,” St. 145,
lat. 46° 40'S., long. 37° 50! E., off Prince-Edward Isles, 150 f.
(Davidson) !
One of my Tunisian specimens is acutely triangular. The lower
valve is sometimes more convex than the upper, as in Anomia ephip-
pium, owing to the habit this Brachiopod has of attaching and
moulding itself by the under surface to corals and other irregular
substances.
This is the Terebratula appressa of Forbes, who published it in
the same year as Philippi. Mr. Dall referred to this Species the
Orthis eusticta of Philippi; but that is a species of Megerlia, and
very different from this. According to Professor Seguenza the
young of P. anomioides is the Terebratula seminulum of Philippi.
Mr. Davidson proposed the genus Morrisia for the present Species ;
but, with true scientific candour, he gave it up in favour of Platydia,
which had been described by O. G. Costa a few months previously
in the same year. In the ‘Annals’ for December 1855, he appa-
rently considered Morrisia and Megerlia subgenera of Terebratella;
and in July 1861 he queried Morrisia as a distinct genus. However,
his present scheme of classification is much more satisfactory ; and
I will not presume to criticise it.
Genus V. Mrerrtia, (Megerlea) King.
/ MEGERLIA TRUNCATA, Linné.
Anomia truncata, L. 8. N. p- 1152.
M. truncata, B.C. ii. p- 22; v. p. 164.
‘Poreupine’ Exp., 1870: Atlantic, St. 24, 292 f. (and var.
monstruosa); 36, 128 f. Mediterranean, 45, 207 f. (and var.),
Capo de Gata, 40-69 f.; 58, 266 f. (and var.) ; Adventure Bank,
92 f. (and var.),
Atlantic coasts of France and Spain, Mediterranean, Adriatic, and
Aigean: 10-120 f. Cape of Good Hope, 224 f. (and var.). Canary
Isles, 60 f. ‘ Challenger’ Exp., off Teneriffe, 70-75 f. Isle of
Bourbon, 200 f. (Deshayes, as Morrisia gigantea). Australia (Angas,
412 DR. GWYN JEFFREYS ON MOLLUSCA OF THE [Apr. 16,
fide Dall). The habitat given by Linné, “in pelago Norvegico,” is
erroneous, so far as we at present know or believe, as well as
“Torquay ” on Dr. Turton’s authority.
Very common and not less variable. Some specimens, especially
the young, are more or less acutely triangular; others are nearly
ribless or smooth ; the very young have merely a moderately large
byssal orifice in the lower valve, the beak of the upper valve being
pointed ; and the loop is annular as in the subgenus Terebratulina.
This is a case of progressive development, and the reverse of what
takes place in Terebratula septata. The skeleton is distorted in the
monstrous form or variety. The size given by Deshayes for his
Morrisia gigantea scarcely exceeds one half of some ‘ Porcupine’
specimens from 266 fathoms.
There are six other synonyms, but obsolete. Terebratula (after-
wards Orthis) lunifera of Philippi, Forbes, and Costa is the young ;
the skeleton being imperfect or broken causes the appearance denoted
by that specific name. Mr. G. B. Sowerby, jun., figured in his
‘Thesaurus’ a ribless variety of Argiope cuneata as Philippi’s species.
The distorted variety, which Scacchi at first described as Terebra-
tula monstruosa, but subsequently called ‘‘ Antecedentis varietas”’
with reference to the present species, is the Morrisia davidsoni of
Eudes Deslongchamps; it not unfrequently occurs with the normal
and intermediate forms.
Seguenza in 1865 classified ‘‘ Megerlea” as a subgenus of Tere-
bratella.
Genus VI. THECIDEA.
THECIDEA MEDITERRANEA, Risso.
T. mediterranea, Risso, Hist. Nat. de Eur. mérid. iv. p.394, f. 185.
Mediterranean, especially the African coasts, 30-300 f. Jamaica,
60 and 150 f. (Barrett, fide Davidson). Mauritius (Sir Henry
Barkly)!
Thecidea spondylea of Scacchi, and probably 7’. testudinaria of
Michelotti.
B. Imperforate.
Family Il. RoyNCHONELLID&.
Genus 1. ATRETIA.
ATRETIA GNOMON, Jeffreys. (Plate XXIII. fig. 4.)
A. gnomon, Jeffr. Proc. R. 8S. no. 121, 18th Noy. 1869, p. 121 ;
Ann. & Mag. N. H. Sept. 1876, p. Db):
‘Porcupine’ Exp., 1869: St. 20, 1443 f.; 30, 1380 f.
‘Valorous’ Exp.: Davis Strait, 1100-1750 f.; North Atlantic,
1450 f. Norwegian Arctic Exp., 1877: about thirty miles west of
«‘Tromsd,” on the slope of the banks, “cold area,” 650 f. (Friele) !
According to Mr. Dall, Atretia is a synonym of Zittel’s genus
Dimerella (Dunker and v. Meyer, Palzont. 1870, p. 220), which is
a Jurassic Brachiopod; but, although it belongs to the same family,
1878.] ‘LIGHTNING’ AND ‘ PORCUPINE” EXPEDITIONS. 413
the internal apparatus is different, and the beak is prominent in
Atretia and incurved in Dimerella. The septum in A¢fretia is short,
gnomon-shaped, and central ; in Dimerella it extends from below the
beak to the front of the lower or smaller valve. The genus Manna
of Dewalque (Soc. Malac. Belg., 1874), from the “sable noir,” or
Upper Miocene of the Antwerp Crag, is also a Rhynchonellidan ; but
the internal apparatus is likewise different from that of Atrefia.
Each of the curved processes in the lower or smaller valve of Aéretia
gnomon has the outer end trifid, or is divided into three symmetrical
digitate points.
Genus II. RuyncHone ta.
1. RHYNCHONELLA PSITTACEA (ROSTRUM-PsITTACI), Chemn.
Anomia rostrum-psittaci, Ch., viii. p. 106, tab. 78. f. 4 a, 8, ¢.
R. psittacea, B.C. ii. pp. 22, 23; v.p. 164, pl. xeix. f. 4.
Arctic and northern seas in both hemispheres, from Spitzbergen to
Drontheim, and from Franklin-Pierce Bay in 79° 25! N. lat. to
Maine, U.S., in the North Atlantic; and from Behring Strait to Japan
in the North Pacific: 10-690 f. Australia, with R. nigricans (J. W.
Flower)! Shetland, 90-100 f.; apparently recent, but possibly
relics of the “ glacial era”? (J.G. J.). North Sea, S.E. of Yarmouth
(Metzger) ; possibly also semifossil. Several spurious or more than
doubtful habitats on the Scotch and English coasts have been re-
corded.
J 2. Rayncuonenia sicuLa’, Seguenza, MS. (Plate XXIII. figs.
5, 6.)
Bopy whitish, gelatinous : man¢éle not furnished at its edge with
setee or bristles, as is the case in the Terebratulidee: arms fringed with
short cirri, which are unequal in length and curl inwards at their
extremities : byssal plug small, cylindrical, and slender; its outer
case or sheath is chitinous, and resists the action of liquor potasse.
SHELL ovately triangular, having the upper valve convex and the
lower valve somewhat compressed, rather thin, semitransparent, and
glossy: sculpture, numerous and close-set, minute, very fine and
regular longitudinal striz ; lines of growth slight but distinct : colour
pale brownish-yellow: margins nearly straight in front (instead of
being flexuous as in R. psitéacea), irregularly sloping at the sides,
broadly angular behind: beak very short, incurved : foramen very
small, oval, incomplete: deléidium broad, slightly concave: hinge-
plate narrow : teeth in the upper valve triangular and long ; the pro-
cesses in the lower valve approximate at the base, forming an in-
verted V, and are irregularly oblong, rather thin, and concave in the
middle; each of these processes extends outwards as a thin and
narrow lamina, which curves upwards, and is divided at its outer end
in a pectinate or comb-like manner, as in Atretia gnomon, but has
more digitations or points; the sockets of the teeth are grooved
transversely : septum ridge-like, short, placed in the centre of the
1 Sicilian.
414 DR. GWYN JEFFREYS ON MOLLUSCA OF THE [Apr. 16,
large muscular scar below the beak: inside smooth and glossy, not
contracted behind, nor grooved like R. psittacea. L. 0°9, B. 0°75.
‘Porcupine’ Exp., 1870; St. 3, 690 f.: Chops of the English
Channel. One perfect and living specimen, besides two not quite
perfect valves and some fragments.
The shape of the Sicilian fossil, for a specimen of which (see Plate
XXIII. fig. 6) I am indebted to Professor Seguenza, is rather more
broadly triangular, and is uniformly convex; but it agrees with the
recent specimen above described in its peculiar sculpture, want of
flexuosity in the front margin, short beak, and small foramen. The
fibrous texture of the shell is much softer and looser than that of 2.
psittacea'. Attached to the living specimen of R. sicula is a remark-
able species of Balanus or an allied genus, which will be elsewhere
noticed.
** Hingeless.
Family I. CRANIID2.
Genus CRANIA.
CraniA ANOMALA, Miller.
Patella anomala, Mill. Zool. Dan. Prodr. p. 237.
C. anomala, B. C. ii. p. 24, pl. i. f. 3; v. p. 165, pl. xix. f. 6.
‘Lightning’ Exp., St. 2,170 f.; 4, 530f.; 5, 189 f. (var. alba).
‘Porcupine’ Exp., 1869, St. 2, 808 f.; 6, 90 f.; 7,159 f.; 12,
670f.; 14,173 f., on Oculina prolifera; The Minch, 170f.; 74 & 75,
203-250 f.; 78, 290 f., with Platydia anomioides. 1870, Atlantic,
1, 567 f.; 3, 690 f., upper valve coloured with reddish-brown streaks ;
24, 292 f.: Mediterranean, 45, 207 f.; 58, 266 f.; Adventure Bank,
92 f.
Spitzbergen to the Mediterranean, Adriatic, and Aigean : 3-255 f.
Jamaica (Barrett, as C. personata, in coll. M‘Andrew); North Sea,
0-106 f. (Metzger).
Some of the specimens procured by Dr, Carpenter in H.M.S.
‘Shearwater’ off the coast of Tunis, in from 40 to 120 fathoms, are
unusually thick, especially at their edges, which are closely notched
inside ; and they belong to the variety which Seguenza has described
and figured as C. /amellosa from the Sicilian Pliocene beds. In two
or three of these ‘ Shearwater’ specimens the hinder margin of the
upper valve is nearly straight; and the posterior muscular scars
project somewhat beyond the sides, so as to give a hinge-like cha-
racter to that part of the valve. The interior of the upper valve in
one specimen is covered with crowded oblong tubercles, which are
arranged in the line of growth. The inside margin of the lower
valve is generally more or less raised, as I pointed out in ‘ British
Conchology,’ vol. ii. p. 25.
There are several synonyms, viz.:—Criopus jfimbriatus for the
animal or soft parts of this and other Brachiopods, and Anomia
1 In R. psittacea the curved processes in the lower valve are rather long, ir-
regularly triangular and twisted ; the lamina is furnished underneath with five
or six slight obliquely transyerse ridges, which form notches at the inside edge.
1878.| ‘LIGHTNING’ AND ‘ PORCUPINE’ EXPEDITIONS. 415
turbinata for the shell of the present species (Poli) ; Patella distorta
(Montagu); Orbicula norvegica (Lamarck) ; C.ringens and C. rostrata
(Heeninghaus); Discina ostreoides (Turton); Criopus orcadensis
(Leach), and Crania personata (Sowerby and others, not Lamarck) ;
var. alba=C. suessia (Reeve, in coll. M‘Andrew) !
Mr. Dall was quite right in ascribing the original authorship of
the generic name crania to Retz, instead of to Philippson, as I had
erroneously stated. See Schrift. Berlin. Ges. naturf. Fr. Bd. ii.
p- 72, tab. i. f. 2-7, 1781. Philippson published the same name in
1788.
Family II. Discrinipa.
Genus Discrna.
v Discina aTLANTICA, King. (Plate XXIII. fig. 7.)
D. atlantica, King in Proc. N. H. Soc. Dublin, 1868, vol. v.
p- 170: Jeffr. in Ann. & Mag. N. H., Sept. 1876, p. 202.
‘Porcupine’ Exp., 1869, St. 19 a, 1366 f.
Off the west coast of Ireland, 1240 f. (Capt. Hoskyns). North
Atlantic, while fishing up the deep-sea telegraph cable in about
2400 f. (Sir James Anderson), ‘ Valorous’ Exp., 1450 & 690 f.
(J. G.J.). ‘Challenger’ Exp., near St. Paul, 1850 f.; N. Pacific,
1875 f. & 2050 f.; off Bermuda, 2180 f. (Davidson) !
I have also to thank Mr. Davidson for telling me of a mistake in
my short description of the animal in the ‘ Annals’ above cited ;
and I gladly take this opportunity of correcting it. The word mantle
ought to be substituted for “ arms.”
D. atlantica is probably the same species as the Coralline-Crag fossil
which Mr. 8. Wood at first doubtfully named D. norvegica and after-
wards D. fallens, and which Mr. Davidson at first doubtfully named
Orbicula lamellosa and afterwards D. fallens. But O. norvegica of
Sowerby (=O. lamellosa, Broderip) is a very different and tropical
species of Discina.
The genus Discina appears to be the connecting link, at least analo-
gically, between the Brachiopoda and Conchifera through Anomia,
both having a similar plug of attachment in the lower valve.
EXPLANATION OF THE PLATES.
Puate XXII.
Fig. 1. Terebratula caput-serpentis, young; to show the loop: p. 401.
2. T. tuberata, p. 401.
3. T. trigona, p. 402.
4. T. subquadrata, p. 402.
5. T. vitrea; loop: p. 403.
6. T. vitrea, var. sphenoidea; loop: p. 404.
7. T.tenera, p. 405.
Puate XXIII.
. Terebratula septata, p. 407.
. Terebratella spitzbergensis; loop: p. 409.
. Megerlia jeffreyst; loop: p. 407.
. Atretia gnomon, p. 412.
Hm OS bo
416 DR. M. WATSON ON THE MALE { Apr. 16,
Fig. 5. Rhynchonella sicula (recent), p. 413.
6. R. sicula (fossil), p. 414."
7. Discina atlantica, p. 415.
Observation—In the enlarged figure of Terebratula vitrea, var. sphenoidea,
the strise are much too strongly marked. In the figure of Discina atlantica the
setz are represented too thick.
2. On the Male Generative Organs of Hyena crocuta. By
M. Warson, M.D., Professor of Anatomy, Owens Col-
lege, Manchester.
{Received April 5, 1878.]
. (Plates XXIV. & XXV.)
In a previous communication ’ I laid before this Society a deserip-
tion of the female organs of Hyena crocuta, and having lately had
an opportunity of examining those of an adult male of the same
species, have thought that a description of these would not be alto-
gether uninteresting, more especially as, so far as I can ascertain, no
complete account of them has hitherto been published. For the
sake of comparison I shall follow the plan pursued in the paper
referred to, describing in the first place the external, and secondly
the internal organs of generation.
External Parts.
The anal and perineal regions of the male resemble so closely those
of the female that without a very accurate examination it is impos-
sible to distinguish between the sexes. In both there is a well-
marked glandular pouch above the anus; in both there are cutaneous
elevations corresponding in appearance to that of the scrotum in the
males of allied species ; and in both there is an elongated, pendulous,
penis-like body, surrounded by a prepuce, and perforated at its
extremity by a single aperture of small size.
Penis.—This organ is directed forwards, and lies along the middle
line of the perineum as in the dog. The free portion, consisting of
a central glans surrounded by a thick fleshy prepuce, projects for-
ward and downward from the level of the abdominal wall, and
measures one inch and a half in length. The glans is conical in
form and without distinct corona, the urethral aperture being situated
on the apex of the cone. The inferior lip of this aperture is pro-
longed forward to some extent, so that seen from the side the glans
presents the appearance of being obliquely truncated from before
backward and upward as in the female, although the truncation is
not so well marked as in that sex. In the flaccid condition of the
organ the glans is for the most part concealed by the prepuce, its
apex only being exposed. The latter portion is of a dark blue colour,
and its surface is much corrugated, whilst so much of the glans as is
concealed by the prepuce is of a purplish colour and presents no trace
of corrugation. Its basal two thirds are moreover covered by small
7 Proc. Zool. Soc. May, 1877.
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1878. ] GENERATIVE ORGANS OF HYZNA CROCUTA. 417
recurved spines, which diminish in size toward the margin of the
prepuce and are altogether absent on its free corrugated portion.
The urethral aperture is smaller and less dilatable than the corre-
sponding opening in the female. The prepuce, consisting of a thick
double fold of integument of a bluish colour externally and flesh-
coloured internally, is thrown into well-marked circular rugz and is
attached by a distinct freenum to the under surface of the base of
the glans. When retracted it is continuous with the surface of the
latter. In the female the freenum is attached, not to the dase of
the glans, but to the inferior lip of the urino-genital canal.
Perineum.—The perineum, from the inferior border of the scrotum
to the base of the free portion of the penis, measures 4 inches in
length, a measurement which agrees exactly with that of the corre-
sponding region of the female. The integument, like that of the
prepuce is of a bluish colour, smooth, and devoid of hair with the
exception of a space immediately behind the free portion of the penis,
which space is covered with down of a light yellowish colour, the
hairs being directed forwards. It is worthy of note that this area
in the male corresponds exactly with that which in the female is
studded with the little depressions described in my paper on the
genital organs of that sex. And here I would take the opportunity
of correcting an error into which I unwittingly fell in that paper. I
refer to the statement contained therein that the integument of the
female perineum is thrown into longitudinal ruge. These, as the
accompanying drawing shows, are altogether absent in the male in
the recent state of the parts, and only manifest themselves subsequent
to their immersion in aleohol, As the female organs had been sub-
jected to this treatment previous to their examination, I have little
doubt that in their natural condition these parts present the uniformly
smooth appearance I have just described in the male animal. The
presence of these folds in the female, moreover, induced me to hazard
the conjecture that they might represent in H. erocuta the labia of
other carnivora. This supposition is now no longer tenable, as the
examination of the male perineum has led me to conclude that the
homologues of the labia are not to be sought in the longitudinal
perineal folds just referred to, but rather in the cutaneous elevations
described in my former paper under the name of “ scrotal pouches.”
Scrotum.—Yhe scrotum is non-pendulous, and resembles that of
the Felidee proper. It is situated one inch below the margin of the
anus ; and its division into two is indicated externally by a shallow
groove. Each half is about the size of a small walnut; and the
surface of each is covered with hair of alight brown colour. It is
interesting to observe that this description corresponds almost exactly
with that of the so-called “‘ scrotal pouches”’ of the other sex, the
only difference being that in the male the scrotum is slightly more
prominent than the “scrotal pouches ”’ of the female.
Anus and Anal-gland Pouch.—The anus is situated one inch above
the scrotum ; and between the former and the root of the tail is a
pouch similar to that which is found in the female. The interior of
the pouch is smooth and devoid of hair, except the roof, which is
Proc. Zoou. Soc.—1878, No. XXVII. 27
418 DR. M. WATSON ON THE MALE (Apr. 16,
clothed with down of a lightish colour. The down at the external
angles of the pouch is longer than elsewhere, and forms two tufts
which project into the pouch. Opening into the bottom of each
extremity of this pouch is the duct of an anal gland ; whilst extending
between these larger apertures is a belt of small perforations, which
indicate the ducts of a number of small isolated follicles lying in
relation to the fundus of the pouch.
Nipple.—Upon either side of the middle line, and half an inch
external to the free portion of the penis, is a single rudimentary
nipple. It measures one eighth of an inch in height, and lies con-
cealed in the thick hair of this region.
Internal Organs.
Rectum and Anal Glands.—The rectum occupies the usual posi-
tion in the pelvis, and is provided with walls of unusual thickness,
the external or longitudinal muscular bundles being particularly well
developed. It is attached to the lower surface of the sacrum by a
double fold of peritoneum or meso-rectum, which measures between
two and three inches in breadth, and consequently allows of consi-
derable movement of the rectum within the cavity of the pelvis.
The terminal portion of the rectum is surrounded by two muscles,
the levator and sphincter ani. 1. The levator ani is of considerable
strength, and arises from the fascia lining the pelvis. Its fibres pass
backward and upward, and terminate on the walls of the rectum
under cover of the sphincter ani. 2. The sphincter ani is very
powerful, and arises from the under surface of that caudal vertebra
which is situated directly above the anus, its origin being interposed
between those of the depressors of the tail. The muscle is two
inches in breadth, and invests not only the extremity of the rectum
but also the anal glands together with the pouch into which these
open. It is inserted into the skin midway between the anus and
scrotum. By means of this arrangement the sphincter ani acts not
only as such upon the anus and anal-gland pouch, but also as a com-
pressor of the anal glands. The gland-pouch resembles the corre-
sponding structure in the female, being transversely oval in form.
The anal glands are two in number, one being placed on each side of
the rectum. They are oval in form and nodulated on the surface.
Each measures 1j inch in length and 3 inch in greatest breadth.
They are thus considerably smaller than the corresponding glands in
the female, in which they measure 24 inches in length and 13 inch
in breadth. The gland consists of a secreting wall 3 inch in thick-
ness, which limits a central cavity. From the posterior extremity of
this cavity a duct passes off and opens into the corresponding angle
of the anal pouch. ‘The eavity of the gland was filled with a thick
secretion, which differed from that of the female in being of a
blackish colour and in being destitute of the very offensive odour
characteristic of that sex. In addition to these larger glands a belt
of isolated follicles extends across the bottom of the anal pouch.
They pour their secretion into the latter by means of the little aper-
tures above described.
1878. ] GENERATIVE ORGANS OF HYZNA CROCUTA. 419
Bladder and Urethra.—The bladder is regularly pyriform and of
small size, measuring 4 inches in length. As in the female, it passes
gradually into the urethra, so that it is difficult to say where the
one ends and the other begins. It is completely covered by peri-
toneum as far back as the openings of the ureters, and is attached
to the pelvic and abdominal walls by reflections of that membrane.
The ligaments thus formed are three in number, two being superior
and one inferior. Each superior ligament is attached by one extre-
mity to the superior and supero-lateral pelvic wall, and by the
other to the lateral aspect of the bladder. Each consists of a double
fold of peritoneum, in the anterior or free margin of which is placed
the fibrous cord formed by the obliterated hypogastric artery ; and
crossing the inner side of the latter from before backward as they
enter the pelvis are the ureter and the vas deferens, the ureter being
uppermost. The inferior vesical ligament is sickle-shaped, and con-
sists of an elongated fold of peritoneum, which extends between the
ventral surface of the bladder and the anterior abdominal wall as far
forward as the umbilicus. The ureters are remarkably thick and
muscular, and open into the base of the bladder half an inch from
the orifice of the urethra. The urethra extends from the bladder to
the extremity of the penis, and consists of two portions, a membra-
nous and a spongy. ‘The membranous portion lies within the pelvic
cavity, and reaches from the neck of the bladder to the bulb of the
urethra. It measures 3 inches in length, and is surrounded between
the openings of the vasa deferentia and the bulb by a thick layer of
circularly arranged muscular fibres, the superficial fibres being
attached to the peritoneal fold which intervenes between the bladder
and rectum, whilst the deeper fibres are confined to the urethral wall.
On slitting open this portion of the canal a well-marked longitudinal
fold of mucous membrane is seen to extend along its floor from the
neck of the bladder as far as the middle, in length, of the membranous
part of the urethra, where it gradually disappears. Upon the summit
of this fold, and 1 inch in front of the neck of the bladder, is a cir-
cular opening + of an inch in diameter. This opening is the mouth
of a very minute recess, on the fundus of which the two vasa defe-
rentia terminate. The recess evidently represents the vesicula pro-
statica, but does not, so far as I could ascertain, form a central cul-de-
sac extending beyond the openings of the vasa, as it does in another
species of this genus. The prostate gland is altogether absent—a
fact difficult to reconcile with Professor Flower’s observation, that in
a preparation of the male organs of H. crocuta in the Museum of
the Royal College of Surgeons this gland measures “ half an inch in
length and rather less in breadth”’’.
The spongy portion of the urethra lies within the penis, and mea-
sures 8 inches in length. It is surrounded by a layer of erectile
1 Since the above was in type I have had, through the kindness of Professor
Flower, an opportunity of examining the organs from which his description was
taken, and can bear testimony to the truth of his observations regarding the
presence of a prostate gland. It consisted of two distinct masses, each mea-
suring half an inch in length and a quarter of an inch in are the duets of
2
420 DR. M. WATSON ON THE MALE [Apr. 16,
tissue, which is of considerable thickness where it forms the bulb, but
is thin elsewhere. Opening into the bulb are the ducts of Cowper’s
glands.
Testicles and Vasa deferentia.—The testicles are lodged in the
non-pendulous scrotum, which is divided into two distinct compart-
ments by a median septum. Each compartment is lined by a tunica
vaginalis, which is reflected over the testicle. The cavity of the sac
is continuous at the abdominal ring with that of the peritoneum, the
communication between them remaining pervious. The testicle is
oval in form, and measures 17 inch in length. The epididymis lies
along its lower side, and is expanded above and below to form the
globus major and minor. The latter is nearly as large as the former ;
and the so-called body of the epididymis is consequently reduced to
a mere rudiment. The vas deferens, continuous with the globus
minor, passes through the inguinal canal, enters the pelvic cavity,
where it lies between the layers of the superior ligament of the
bladder, and finally sinks into the muscular wall of the membranous
portion of the urethra to terminate along with its fellow on the floor
of the vesicula prostatica as already described. There are no vesi-
culz seminales.
Cowper’s Glands.—These are two in number and of large size.
Each is placed alongside of the membranous portion of the urethra,
in the interval between the levator ani and ischio-cavernosus muscles.
Each gland is pyriform, and measures 1} inch in length, and 2
of an inch in greatest breadth. The base is directed forward; and
from the apex is given off a duct 2 of an inch in length, which
opens into the roof of the bulbous portion of the urethra. The
glands are solid ; and each is provided with a capsule of pale muscular
fibres.
Penis.—Tbis organ is composed of two corpora cavernosa together
with a corpus spongiosum. The former consist of two bands of
erectile tissue flattened from side to side as in the clitoris of the
female, and united along the middle line. Each is attached to the
ischium just in front of the tuberosity of that bone, to form the crus,
and unites with its fellow opposite the posterior extremity of the
pubic symphysis. The corpus spongiosum is perforated by the
urethral canal, and expands anteriorly and posteriorly to form the
glans and the bulb, With the exception of these portions, this mass
of erectile tissue is thin and contributes no great thickness to the
wall of the urethra. The bulb is large and prominent, and is in-
vested by the fibres of the bulbo-cavernosi muscles. The corpus
spongiosum of the male therefore differs from the corresponding
structure in the female, inasmuch as in the latter this body is divided
into two lateral halves which are situated altogether above the urino-
genital canal, whilst in the former they have coalesced to form a
which open into a little recess on each side of the utriculus. As this animal
was known to be at least twenty-four years of age, whilst my specimen was not
adult, it is possible that by reason of their small size the prostate glands of the
latter may have escaped my observation. In Mr. Flower’s specimen the other
generative glands were each twice as large as in my own.
1878.] GENERATIVE ORGANS OF HYZNA CROCUTA. 421
single mass surrounding the canal of the urethra. Further, in the
female the corpora spongiosa take no part in the formation of the
glans clitoridis, whilst in the male the glans penis is formed by the
spongy body. The penis, formed as just described, measures
oan in length from the bulb of the urethra to the apex of the
glans.
Muscles of Penis.—Of these there are three which are bilateral,
and one which is common to both sides. (1) The tschio-cavernosus
or erector penis is very strong, and arises from the ischium imme-
diately in front of the tuberosity. The fibres pass forward and are
inserted into the corresponding corpus cavernosum close to the junc-
tion of the latter with its fellow on the opposite side. The muscle
covers the crus penis. (2) The dulbo-cavernosus is also strong, and
arises from a median tendinous band which separates it from its
fellow. The fibres pass transversely outward so as to cover the
bulb of the urethra, and are inserted into the corpus cayernosum
along the inner border of the preceding muscle. (3) Retractor
penis. In my paper on the female organs of H. erocuta I expressed
my belief that the retractors of the clitoris probably arise from either
the ischium or pubis. I am now satisfied from my examination of
those of the male that such is not the case, but that in both sexes
they arise from the lower surface of the sacrum. In the male each
retractor is a broad but thin muscle which, arising from the pelvic
surface of that bone, passes backward and downward and reaches the
side of the rectum. Some of the fibres terminate on the gut, where
they lie under cover of the levator ani; but the greater number form
a narrow riband-like muscle, which passes downwards and extends
along the under-surface of the penis in contact with its fellow to be
inserted, as in the female, into the corpus cavernosum just behind
the glans. (4.) Elevator urethre. The muscle so named is very
strong, being nearly as thick as the erector. It is a flattened band,
and arises from the inner surface of the ischium 1 inch in front of
the tuberosity. The fibres pass downward and forward, and termi-
nate below the angle formed by the junction of the membranous and
spongy portions of the urethra on a tendon which is common to it
and to the muscle of the opposite side. Lying, as it does in the
normal position of the animal, Je/ow the urethra, this tendon when
the muscular fibres contract will tend to elevate and at the same
time to compress the urethral canal. A large artery, vein, and
nerve derived from the pudics pass together along the outer side of
each cavernous body as in the female, and terminate close to the
glans. Another artery of large size, also derived from the pudic
trunk, passes into the crus penis of each side, and apparently supplies
the erectile tissue.
Comparison of the Male Organs of allied Species —Having now
described the male organs of H. crocuta, it may be well to compare
them with those of closely allied species. The number and arrange-
ment of the anal glands is similar in both sexes of this species ; and
therefore there is little to add to what I have already said regarding
them in my communication on the female organs, In my descrip-
422 DR. M. WATSON ON THE MALE [Apr. 16,
tion of these glands in the latter sex I pointed out that in respect of
them H. crocuta agrees more closely with Profeles than with any
species of its own genus, and may merely add that the present dis-
section has enabled me to confirm Dr. Murie’s observation * that in
the male animal as in the female an anal-gland pouch is present.
Mr. Busk? had previously convinced himself “that no trace of a
pouch between the root of the tail and the anus exists, at any rate
in the male of this species ;’”? and Dr. Murie, in deference to Prof.
Kaup’s® attempt to classify the Hyzenas in accordance with the
presence or absence of this pouch, was not prepared to deny that
it might occasionally be absent in the male. I think this matter
may now be considered settled, and may merely observe that Gold-
smith * appears to have been aware of the presence of this pouch in
every species of Hyzena before any of the authors referred to had
written on the subject.
With reference to the male organs of allied species, I find that
Daubenton ° describes and figures those of H. striata ; and these are
also referred to by John Hunter® and Owen’. Cuvier* gives an
account of the male organs of Hyzena without indicating the species
to which they belong ; but from the agreement between his observa-
tions and those of the authors just named, I presume that H. striata
formed the subject of his investigations. Of the male organs of H.
brunnea, so far as I can ascertain, we know nothing; but of those
of the closely allied genus Proteles, Prof. Flower® gives an accu-
rate description. H. striata and Proteles agree with H. crocuta
with respect to the form and position of the scrotum, the form of
the testicle, and the absence of vesicula seminales, whilst they differ
from the last-named species in the possession of a prostate gland.
Mr. Flower, it is true, refers to the presence of a prostate gland in
H. crocuta® ; but of the absence of this in the specimen I examined
I was careful to convince myself; and until further observations are
made, it is impossible to reconcile these opposing statements regard-
ing a matter of fact. In H. striata the prostate is large and con-
sists of two kidney-shaped lobes, whilst in Proteles it consists of a
single mass of large size having ‘‘ the appearance of a bilobed disk.”
Cowper’s glands are large and of the same form in all three species,
which likewise agree in the form of the bladder and the very mus-
cular character of the ureters. With respect to the vesicula pros-
tatica, Leuckart™ describes and figures it in H. striata as a minute
flask-shaped cul-de-sac projecting beyond the urethral wall and
lying in the interval between the lobes of the prostate. He says,
moreover, that in that animal it does not open into the urethra.
1 Trans. Zool. Soe. vii. p. 503.
2 Journ. Proc. Linn. Soe. vol. ix. p. 71. 3 Tsis, 1828, p. 1144.
4 Animated Nature, ‘‘ Hyena.”
® Buffon’s Hist. Nat. tom. ix.
6 « Wssays and Observations,’ by Owen, vol. ii. p. 58.
7 ‘ Anatomy of Vertebrates,’ vol. iii. p. 671.
8 * Anatomie Comparée,’ vol. v.
» Proc. Zool. Soc. 1869, p. 491. 10 Thid. p. 493.
1‘ Cyclopedia of Anatomy,’ vol. iv. Art. “ Vesicula prostatica.”
1878.] GENERATIVE ORGANS OF HYEZNA CROCUTA. 423
Mr. Flower found no distinct uterus masculinus in Proteles; but as
he refers to a median ridge on the floor of the urethra, in which he
could detect a very minute aperture, the arrangement appears to be
very similar to that described above in H. crocuta, In respect
therefore of the form of the vesicula prostatica, I. crocuta agrees
more closely with Proteles than with striata, differing from the
latter inasmuch as the vesicula does not project beyond the urethral
wall, and in the fact that it communicates with the canal of the
same. According to Leuckart it is an extremely rare occurrence
that the vasa deferentia open into the vesicula prostatica ; but this
is certainly the case in H. crocuta. The penis of the Spotted
Hyeena closely resembles that of H. striata and of Proteles, differing
only from these in the absence of a conical body of cartilaginous
consistence which has been described by Cuvier in the glans penis
of the former and by Professor Flower in that of the latter. The
glans penis in all of them is invested by small recurved spines.
Comparison of the Male and Female Organs of H. crocuta—
When describing the female organs, I remarked that “ the arrange-
ment of these is such as to give them a great resemblance to those
of the males in other species,” and that, if we supposed the vesicula
prostatica of the latter enlarged to the size of the female uterus, we
should in the absence of a prostate gland, have an almost identical
arrangement of the excretory passages of both sexes. The absence,
as a matter of fact, of the prostate gland in the male H. crocuta
makes the resemblance between the male and female organs of this
species even greater than I anticipated. And here I may be per-
mitted to observe that nowhere in the group of mammals is the truth
of the conclusions at which embryologists have arrived respecting
the homologies of the various parts of the sexual apparatus in the
two sexes so beautifully shown asin the animal under consideration,
Did any doubt remain regarding the similarity of plan upon which
these are built up, it would be at once dispelled by an inspection of
the sexual organs of Hyena crocuta. In both sexes there is a short
urethra opening close to the mouth of the uterus, which organ, in
accordance with functional requirements, is of large size in the female,
but is reduced to a minimum in the male. In both there is a urino-
genital canal extending from the junction of the urinary and sexual
canals to the extremity of the penis or clitoris, according to sex; and
in both there are two Cowperian glands of large size Opening into
that canal close to the root of the penis or clitoris. A comparison
of the internal genital organs of the two sexes proves conclusively
that the lower part of the so-called prostatic, together with the
membranous portion of the male urethra, are homologous with
the commencement of the urino-genital canal of the female, which
canal, in the majority of mammals, is so metamorphosed to form
the vestibule, that its homologue in the other sex is by no means
readily recognizable. This homology is further substantiated, in
the animal we are considering, by the size and form of Cowper’s
glands, and the points of entrance of the ducts of these into the
urino-genital canal in each sex. Lastly, the homology of the so-
424 DR. M. WATSON ON THE MALE [Apr. 16,
called spongy portion of the male urethra with that portion of the
urino-genital canal which, in the female, lies in relation to the clitoris,
is manifest. The erectile organ is of the same size, and constructed
upon the same plan, in both sexes, the only difference being that in
the female the urino-genital canal is not surrounded by the erectile
tissue of the corpus spongiosum as is the case in the male, this dif-
ference being due to the fact that in the latter the essentially bilateral
spongy bodies have coalesced in the middle line, whilst in the female,
in accordance with physiological requirements, they remain distinct
throughout life, and do not surround the sexual canal. In the
female, moreover, and associated with this arrangement, there is a
complete absence of the elevator urethra and _ bulbo-cavernosi
muscles.
It may be well now to ascertain whether the examination of this
animal throws any light upon the subject of the probable homo-
logues of the vagina and uterus in the malemammal. The inajority
of embryologists (among whom I may mention the names of Leuckart,
Kdlliker, and Allen Thomson) are now agreed that the so-called
utriculus of the male mammal represents both the uterus and vagina
of the other sex, these organs being formed by the coalescence of
the ducts of Miiller. I have, however, in a previous paper remarked
concerning the corpus uteri of the female H. crocuta, “‘'That the
whole of this is to be regarded as corpus uteri, and not as consti-
tuting any portion of the vagina, is proved by the absence of any
constriction in its interior which might correspond to an os uteri,
the tubular body of the uterus remaining of the same calibre, and
having the walls of uniform thickness down to its opening into the
urino-genital canal.’’ It is evident therefore that in the female H.
erocuta, the vagina being altogether absent, we must conclude that
in the male of this species the utriculus represents the uterus alone,
and not the uterus and vagina together. The same remark holds
good, so far as I can ascertain, of only one other placental mammal—
that is, of the Indian Elephant, in the female of which, as Mayer’
pointed out, the vagina is altogether absent, and the uterus opens
directly into the urino-genital canal. In the male, therefore, of that
animal, as in that of H. crocuta, the utriculus, as shown by Leuckart?,
clearly corresponds to the uterus, and to the uterus alone, of the
female. With reference to the homologue of the prostate gland in
the female mammal, this, according to Prof. Allen Thomson*, is to
be looked for in tissue uniting the urethra with the vagina. We
might therefore be justified in expecting that in those animals in the
female of which the lower ends of the Miillerian ducts unite to form
the uterus and not the vagina, the prostate gland would be absent in
the male. Such is certainly the case in A. crocuta; but that it
would be erroneous to accept this as a general law is proved by a
reference to the Elephant, in which, although the vagina is absent in
the female, the male nevertheless possesses prostate glands of con-
? Nova Acta Acad. Cxs. Leop.-Car. tom. xxii. p. 38.
« Cyclopedia of Anatomy,’ vol. iv., Art. “ Vesicula prostatica.”
° Quain’s ‘ Anatomy,’ 8th edit. vol. ii. p. 826,
1878. ] GENERATIVE ORGANS OF HYENA CROCUTA. 425
siderable size. There appears, indeed, upon consideration, to be no
reason why these glands should not be present in the males of those
species the females of which want the vagina, as well as in those in
which that organ is present, seeing that in both cases we have to do
with the tissue surrounding the lower ends of the Miillerian ducts,
Turning uow to the external genital organs, I have before directed
attention to the very close resemblance which in respect of these
the male bears to the female, the so-called scrotal pouches and clitoris
of the latter closely simulating the scrotum and penis of the other
sex. After a very minute examination of these parts in both sexes
I have only been able to recognize the following by no means very
evident points of distinction between them - (1) In the female the
diameter of the free portion of the clitoris, including the prepuce, is
greater than in the male, measuring 13 inch in the former, and 1]
inch in the latter. (2) The aperture at the extremity of the clitoris
of the female is slightly larger and much more dilatable than the
corresponding aperture in the male. This difference depends on the
fact that the corpus spongiosum in the male forms the glans penis
and completely surrounds the urethral aperture, whilst in the female
the upper portion only of the glans is composed of erectile tissue, its
lower portion being formed by the walls of the urino-genital canal.
(3) The glans penis of the male is covered with recurved spines,
whilst the glans clitoridis of the female is devoid of these. (4)
In the male the cutaneous surface immediately behind the free
portion of the penis is covered with hair, whilst in the female the cor-
responding space is dotted with the little cutaneous depressions I
formerly described, and is devoid of hair.
It will at once be.seen that these points of distinction are slight,
and are not such as would enable one to decide the sex of the animal
in the absence of such an examination as is well nigh impossible so
long as the animal is alive.
Bisexual Nature of the Hyena according to the Ancients.—And
this leads me to observe, what I formerly stated, that the consideration
of these facts may serve to explain the ideas which were common
among the ancient Greeks regarding these animals, namely that
they were hermaphrodites. Since the date of my former communi-
cation, I have endeavoured to come to some conclusion regarding
the origin of this apparently extraordinary belief, with the result
that it appears to me that a belief which in our day would rightly
be considered altogether absurd, by no means deserves to be so cha-
racterized when viewed in the dim light of the knowledge possessed
by the ancients respecting these animals. ‘These people had appa-
rently, either directly or indirectly, ascertained certain facts regarding
the sexual organs of the Hyena, which, in the absence of more de-
finite knowledge, almost compelled them to adopt what I may call
the bisexual theory regarding them—a view which, as I formerly
stated, still obtains among the imperfectly educated Boors of South
Africa. Among the Greeks, Aristotle! (B.c. 384) appears to have
been the first who really investigated the sexual organs of the Hyzena;
* ‘Historia Animalium,’ yi. 32.
426 DR. M. WATSON ON THE MALE | Apr. 16,
but that the genus was well known before his time is proved, first,
by the fact that Herodotus’ (B.c. 484) speaks of the animal as being
met with in the Libyan desert, and, secondly, that the view in ac-
cordance with which each individual was bisexual, was current before
the time of Aristotle, who takes pains to show its absurdity. At a
later date Pliny? reasserts the bisexual nature of these animals; and
his assertion is repeated by Hlian*. As, however, with one exception,
to which I shall by-and-by refer, the observations of the last two
historians do not contain any thing of importance which had not
been previously stated by Aristotle, we may confine ourselves to a
criticism of the writings of the latter ; and by doing so I think we shall
throw some light on the origin of the view in question. Aristotle
says*:— “The Hyzena resembles a Wolf in colour, but is more shaggy,
and its back is provided with a mane. It is said that it has the
genital organs of both sexes at once; but this is not the case. Its
male organ resembles that of the Wolf and Dog; and it has what re-
sembles a female organ under the tail. But this last, although
similar to the female organ in form, is imperforate. Underneath this
again is the feecal passage. The female Hyzena has this same so-
called female organ situated as in the male, just under the tail; but
it isimperforate. Next to this is the feecal passage ; and under this
again is the true female organ. The female Hyena, moreover,
possesses an uterus like other female animals. The female Hyzena
is rarely caught. Hunters maintain that for ten males they catch
but one female.” With regard to this passage, the first question
that arises is as to the particular species of Hyena which Aristotle
himself examined. There can, I think, be no doubt that it was
either H. brunnea or H. striata, but most probably the latter. His
description of the female organs is such as enables us to decide that
it was certainly not H. erocuta, inasmuch as he says that the gland-
pouch had been mistaken for the female organ, showing that the
former had some resemblance to the latter. But the female organ
of H. crocuta bears no resemblance whatever to the gland-pouch ;
therefore it must have been the female organ of another species to
which he referred. In favour of H. striata is (1) his cbservation
that the animal is provided with a mane, which, as we know, is
more evident in the Striped Hyzena than in H. brunnea; and
(2) the geographical distribution of the two species—the Striped
Hyzena being found in abundance in Northern Africa, with the
topography and productions of which the Greeks were well acquainted,
whilst H. brunnea is confined to the central and southern districts of
that continent, districts with which the Greeks were by no means
so familiar. But if we come to the conclusion that either H. striata
or H. brunnea formed the subject of Aristotle’s investigation, I would
point out that in denying, as the result of his observations, the bi-
sexual character of either of these animals, he is attempting to refute
a hypothesis the origin of which (as referring to them) it is impos-
sible to explain, seeing that the male and female external organs of
' Rawlinson’s Herodotus, vol. iii. p. 172. 2 Pliny, viii. 30.
* «Historia Animalium,’ i. 25. 4 Loe. cit.
1878.] GENERATIVE ORGANS OF HYANA CROCUTA. 427
both species differ widely in appearance, and that consequently the
sexes of each are readily distinguishable one from the other.
If this be granted, we need not further attempt to explain an
assertion, the origin of which has no foundation in fact. But that
this hypothesis did originate with reference to either of these species
I am not prepared to admit, as it appears to me that the bisexual
character attributed to the Hyzena aliows of a rational explanation if
we regard that character as referring not to either H. striata or 7,
brunnea, either of which Aristotle may have examined, but to H.
crocuta, which he certainly did mo¢ examine. And as further showing
that in all probability Aristotle unwittingly mixed up and confounded
Teports regarding totally distinct animals, I would direct attention
to two of the sentences above quoted, in one of which he says, “ It
is said that it (the Hyzena) has the genital organs of both sexes at
once ;” and in another ‘‘ The female hyzena is rarely caught. Hunters
maintain that for ten males they catch but one female.” Passing
over the evident inconsistency of these two statements, I have already
shown that the first is inexplicable when applied to either H. striata
or H. brunnea ; and, in like manner, the second is equally devoid of
meaning when regarded as having reference to either of these species,
inasmuch as we have no reason to believe that the proportion in num_
ber of the sexes is different in either of them from what it is in other
carnivorous mammals, or that the female of either is more difficult of
capture than the male. We conclude, therefore, that if the bisexual
theory of the ancients regarding the Hyzena had any foundation in
fact at all, such fact had reference not to either H. striata or H.
6runnea ; and by a process of exclusion we are compelled to regard it
as having reference to H. crocuta. I have already shown that it is
almost impossible to distinguish the female of this species from the
male by the mere inspection of the external genital organs; and in
this fact lies, as I believe, the explanation of the views held by the
ancients regarding these animals. This explanation is moreover
borne out by Aristotle’s observation that « the female Hyzena is
rarely caught,’’ a fact which Pliny also records. The latter historian
makes a further, with reference to the present inquiry important
observation which is not to be found in the writings of Aristotle. He
says, “It is a matter of common belief that the Hyena is bisexual,
and that it is male and Jemale in alternate years.’ It is evident
here that Pliny, like Aristotle, mixes up distinct reports concerning
the animal, as it requires no elaborate argument to show that were
the animal bisexual it did not require to change its sex every year.
Be this as it may, it appears to me that if we consider these various
statements as referring to H. crocuta, the origin of all of them admits
of a rational explanation. It might well be that the ignorant traders
who trafficked with the natives of the interior of the African con-
tinent, observing that all, or at least the majority, of the Hyznas
which they encountered were apparently males, should have ex-
perienced some difficulty in accounting for the continuance of the
species, and have had recourse to the view that each animal was
bisexual. It is not improbable that, as Aristotle explains, this view
428 ON THE GENERATIVE ORGANS OF HYZNACROCUTA. [Apr. 16,
may have been borne out by the presence of an anal pouch, which,
according to him, was mistaken for;the female organ—an explanation
which is rational enough if we consider that H. crocuta formed the
subject of the hypothesis, but one which, as I have already shown,
loses all its force if we regard it as referring to either of the other
species of Hyzena. I would add that the other statements of the his-
torians already quoted, to wit that the Hyzena changes its sex every
year, and that the female is rarely caught, are equally explicable in
accordance with this view. The first of these I regard as another
attempt to solve the problem of generation, seeing that the animals,
to the eye of the observer, were uniformly of the male sex; whilst
the latter is merely a statement of fact. It may be urged that the
geographical distribution of H. crocuta, confined as that animal is
to the region of Africa lying south of the Sahara’, and consequently
beyond the ken of the Greeks, is against this view. But I may be
permitted to point out that itis almost certain that in their time, as
now, trading caravans from the coast visited these districts and con-
veyed to the Greeks information regarding the animals which in-
habited them ; otherwise it is impossible to account for the accurate
knowledge which Herodotus undoubtedly possessed regarding the
African continent as far to the south-west as the valley of the
Niger’. In all probability the statements of travellers regarding the
bisexual nature of the Hyzena, originating with H. crocuta, were
transferred by the Greeks to H. striata, with which they were better
acquainted—statements which when taken as referring to the latter
species are absolutely devoid of meaning, but which, when regarded
as applying to the former, admit, as it appears to me, of a rational
explanation.
I have, in conclusion, to acknowledge the kindness of Dr. Alfred
Young in preparing the illustrations for this paper.
EXPLANATION OF THE PLATES,
Puate XXIV.
View of the male perineum and penis of Hyena crocuta. From above
downwards are seen the orifice of the anal-gland pouch, and anus’;
below the latter is the scrotum, together with the bare perineal inte-
gument ; lowest of all is the penis, surrounded by its prepuce.
Puate XXV.
Fig. 1. Male generative organs and extremity of rectum of Hyena croeuta.
G.P, glans penis; Pr, prepuce; S, scrotum seen from the side; P,
enis; C.P, crura penis, covered by the ischio-cavernosi muscles ;
B. U, Bulb of urethra, covered by the bulbo-cavernosi muscles; R. P,
retractor penis muscle; B, bladder; UR, urethra; T, T, testicles,
that on the right side lying in its serous sac; V. D, vasa differentia ;
R, rectum; A. G, anal gland; L. A, levator ani muscle.
Fig. 2. Intrapelvie portion of urethra laid open, showing elevated fold of
mucous membrane on its floor, together with the opening of the utri-
culus on the summit of this fold.
Fig. 3. Glans penis and prepuce of Hyena crocuta. 'The prepuce is retracted
to show the small recurved spines which invest the glans.
1 «Geographical Distribution of Mammalia,’ by Andrew Murray.
* Heeren’s ‘ Ancient Nations of Africa,’ yol. i. p. 192.
vey ae
1878. ] ON THE ORNITHOLOGY OF THF PHILIPPINES. 429
3. Contributions to the Ornithology of the Philippines.—
No. VIII. On some Luzon Birds in the Museum at
Darmstadt. By Arruur, Marquis or TWEEDDALE,
F.R.S., President of the Society.
[Received March 27, 1878.]
(Plate XXVI.)
Professor Koch, of Darmstadt, through the obliging interven-
tion of Dr. Briiggemann, has kindly sent to me for examination a
small number of birds collected in the vicinity of Manilla by Herr vy.
Othberg. Though few in number, some of them belong to species of
which the Philippine habitat has not hitherto rested on indisputable
evidence, while two of them have not before been known as being
migrants to the archipelago.
COLLOCALIA FUCIPHAGA.
Hirundo fuciphaga, Thunb. Act. Holm. xxxiii. p. 151, t. iv.,
* Java ;” Walden, ‘ Ibis,’ 1874, p: 185.
Collocalia linchi, Horsf. & Moore, Cat. E.I. C. Mus. i. p- 100.
no 123, “ Java.”
A single example of a species of the genus Collocalia was sent by H.
vy. Othberg to the Darmstadt Museum, and, I am assured by Dr.
Briiggemann, was obtained in Luzon. It is not separable from the
Javan species, C. linchi, H. & M., i. e. the true H. Suciphaga of
Thunberg. No species of the genus has hitherto been known to
inhabit Luzon ; nor has this species been found in any other Philip-
pine Island.
Turpvus onscurvs (92).
The Philippine habitat of this Thrush has hitherto rested solely on
Prince Bonaparte’s authority (Coll. Delattre, p.28). Three exam ples
from the vicinity of Manilla are sent to me by Professor Koch.
‘TURDUS CHRYSOLAUS (93).
The certainty of this Thrush being an inhabitant of the Philip-
pines turned on the authenticity of a skin in Mr. Gould’s collection
(of. Sclater, ‘Ibis,’ 1863, p. 197). Four examples from Manilla
are in the Darmstadt Museum.
TURDUs VARIUS.
Turdus varius, Pallas, Zoogr. R.-Asiatica, i. p- 449.
Professor Newton (Hist. Brit. Birds, pt. iv. p. 254) has remarked
that Mr. Gould had received an example of White’s Thrush from
Manilla. While there could be no possible doubt of the strict accu-
racy of Professor Newton’s determination of the species, I did not
430 ON THE ORNITHOLOGY OF THE PHILIPPINES. [Apr. 16,
feel quite confident that the origin of the specimen was indisputable
(for I had also examined it) ; and the species was therefore not in-
cluded by me in my list of Philippine Birds. Professor Koch, how-
ever, sends me three examples from Manilla.
ErytTHropitta kocut. (Plate XXVI.)
Pitta kochi, Briiggemann, Abhandl. naturw. Ver. zu Bremen, v.
p.65, t. ii. Ff. 6,
Professor Koch has kindly sent me the type specimen of this Pitta,
from which the accompanying figure is taken. The species is un-
doubtedly distinct from all the other known members of the sub-
genus. It is almost as large as Hydrornis nipalensis. The tarsus
measures nearly two inches. Its specific characters and points of
difference are fully detailed by its describer (/. c.).
ACROCEPHALUS FASCIOLATUS.
Acrocephalus fasciolatus, G. R. Gray, P. Z.S. 1860, p. 349.
Acrocephalus insularis, Wallace, ‘ Ibis,’ 1862, p. 350.
Calamoherpe fumigata, Swinhoe, P. Z.8. 1863, pp. 91, 293.
Calamoherpe subflavescens, Elliot, P. Z. S. 1870, p. 243.
A single example from Manilla in the yellow under-plumage of
A. fasciolatus, with which species I identify 4. insularis. New to
the Philippines.
ANTHUS GUSTAVI.
Corydalla gustavi, Swinhoe, P. Z.S. 1871, p. 366; Briigge-
mann, Abhandl. naturw. Ver. zu Bremen, v. p. 67 ; Sharpe, Tr. Linn.
Soe. ser. 2, Zool. i. p. 338.
Dr. Briiggemann, who first made known the occurrence of this
bird in the Philippines (/.c.), correctly identified the same example
Professor Koch has sent to me. It was obtained by H. v. Othberg
near Manilla. Some months later, Mr. Sharpe (/. c.) made known
that Dr. Steere had discovered the same species in the island of
Basilan,
ANTHUS MACULATUS.
Anthus maculatus, Hodgs., Gray, Zool. Misc. p. 83; Tweeddale,
P. Z.S. 1877, p. 696.
Two examples from Manilla are sent by Professor Koch. Mr.
Everett also procured it at Monte Alban (cf. Tweeddale, /. c.).
1878. ] ON THE BIRDS OF ANTARCTIC AMERICA, 431
4. Reports on the Collections of Birds made during the
Voyage of H.M.S. ‘ Challenger.,-—No.IX. On the Birds
of Antarctic America. By P. L. Scuarer, M.A., Ph.D.,
F.R.S., and Ospurr Satvin, M.A., F.R.S.
[Received April 10, 1872.]
In this report the birds collected at the island of Juan Fernandez,
at various points during the subsequent passage along the coast of
Patagonia, and at the Falkland Islands are noticed.
The ‘Challenger’ was at Juan Fernandez from the evening of
the 13th to the evening of the 15th of November, 1875, and reached
the coast of Patagonia on the 3lst December, at Port Otway, east
of Cape Tres Montes. On the Ist of January, 1876, a party
landed on Penguin Island in Messier’s Channel; and the same
evening the ship arrived in Cove Bay. On the 2nd January
parties were landed on Middle Island, on the 3rd at Gray Har-
bour, on the 4th at Grappler Harbour, on the 5th and 6th at
Tom Bay, near the Trinidad Passage ; and small collections were
made at each place. On the 8th of January the ship arrived at
Porto Bueno, and remained there till the 10th. The same evening
a party was landed at Isthmus Bay, and a few birds were obtained.
From the 11th to the 13th the ‘Challenger’ remained at Port
Churrucha, and proceeded thence to Port Famine. From the 14th
to the 17th she was at anchor off Sandy Point, whence she proceeded
through the straits, and anchored about 10 a.m. on the 18th off
Elizabeth Island. Leaving the Magellan’s Straits, the © Challenger’
arrived at Port Stanley, Falkland Islands, on the 23rd of January,
1876, and remained altogether about two weeks at the Falklands.
During this entire period 169 specimens of birds were collected,
which are referable to 41 species.
The ornithology of this part of the world is well known to us
from the researches of Darwin, Cunningham, Philippi and Landbeck,
and other naturalists’. It was not to be expected, therefore, that
any novelties would be found during such a cursory passage along
the coast.
1, TURDUS FALKLANDICUS, Q. et G.; Scl. et Salv. Nomencl.
p- 2.
619-621, males,
Bod san fenialed: } Juan Fernandez.
“« Eyes brown, bill and legs yellow, feet a little darker yellow.”
647, male,
648, female,
** Kyes brown; stomachs had berries.’’
685, male,
Gia demale, } Port Churrucha.
** Eyes brown; stomachs had berries.”
* See my address on Geographical Zoology in Rep. Brit. Assoc. 1875, p. 105,
and my list of papers on the Birds of Chili, P, Z. 8. 1867, p. 108.
Gray Harbour, Messier’s Channel.
432 MESSRS. SCLATER AND SALVIN ON THE (Apr. 16,
The occurrence of this Thrush in Juan Fernandez is new to us'.
The skins from that island do not differ from continental specimens.
Specimens 647, 648, from Gray Harbour, Messier’s Channel, and 685,
from Port Churrucha (obtained in the month of January), are in
nestling-plumage. The lower surface is thickly spotted with black ;
and the back-feathers carry light fulvous shaft-stripes, and are tipped
with black.
2. Hirunpo meyent, Cab.; Scl. et Salv. Nomencel. p. 14.
le Be ic \ Sandy Point.
710, 711, females,
“* Byes black ; stomachs had insects.”
3. PuryeiLtus Gay (Eyd. et Gerv.); Scl. et Salv. Nomencl:
p. 3l.
662, 663, females, Porto Bueno.
‘* Kyes red; stomachs had sand and grubs.”
4. PHRYGILUS MELANODERUS (Quoy et Gaim.); Scl. et Salv.
Nomencel. p. 31.
727, male, Falkland Islands.
** Eyes black ; stomach had seeds.”
5. ZONOTRICHIA CANICAPILLA, Gould; Sclater, ‘Ibis,’ 1877,
p- 47, pl. i. fig. 1.
702, 703, females, Sandy Point.
** Eyes brown; stomachs had insects.”
These skins are of young birds, and in a stage in which they
cannot be distinguished from Z. pileata; but from their locality they
should belong to Z. canicapilla.
6. Curzus ATerRIMuS (Kittl.) ; Scl. et Salv. Nomenel. p. 38.
645, male, Gray Harbour.
«* Eyes black ; stomach had insects.”
7. CentTrRiTEs NiGER (Bodd.) ; Scl. et Salv. Nomencl. p. 44.
705-707, males, | :
708, 708a, females, { S247 Point.
“ Eyes black ; stomachs had insects.”
8. AN&ZRETES PARULUs (Kittl.) ; Scl. et. Salv. Nomencl. p. 47.
665. female, Porto Bueno.
** Eyes grey.”
9. ANHERETES FERNANDEZIANUS, Philippi; Scl. ‘ Ibis,’ 1871,
p- 179, pl. vii. fig. 1 ; Sel. et Salv. Nomencl. p. 47.
606-613, males, Juan Fernandez.
** Eyes white ; insects in stomachs.”
1 See article on the Land-birds of Juan Fernandez in ‘ Ibis,’ 1871, p. 178.
1878. ] BIRDS OF ANTARCTIC AMERICA. 433
10. Exatnea avsicers (Lafr. et D’Orb.); Scl. et Saly. No-
mencl. p. 48.
688, male, Port Churrucha.
** Eyes black.”
11. Taunioprera pyrope (Kittl.); Scl. et Saly. Nomencl. p. 42.
704, male, Sandy Point.
“ Eyes black ; stomach had inseets.’’
12, CincLopes paTacHonicus (Gm.); Scl. et Saly. Nomencl.
p- 62.
629, male,
630, female, } Port Otway.
‘* Eyes hazel, feet horn-colour.
641, male, | Messier’s Channel, Cold Harbour.
642, young male,
** Eyes brown.”
No. 642, obtained Jan. 2, is a nestling lately flown.
13. ScyraLopus MAGELLANICUS (Lath.); Sel. et Salv. No-
mencl. p. 76.
675, male,
676, female, \ Porto Bueno.
Eyes black ; stomachs had insects.”
680, male, Isthmus Bay.
‘* Eyes brown.”
14. PyGARRHICUS ALBIGULARIs (King) ; Scl. et Salv. Nomencl-
ps. 67.
674, male, Porto Bueno.
“* Eyes grey ; stomach had insects.”
15. Oxyurus sprntcaupa (Gm.) ; Scl. et Salv. Nomencl. p. 62.
631, female, Port Otway,
‘* Eyes black ; stomach had insects.”
664, female, Porto Bueno.
** Eyes brown.”
687, male, Port Churrucha.
“* Eyes brown.”
16. EusrepHANUS GALERITUS (Mol.); Scl. et Salv. Nomencl.
p. 90; Sel. ‘Ibis,’ 1871, p. 181.
ee ee } Juan Fernandez.
“* Eyes black, legs black.”
677, male, Porto Bueno.
*« Eyes black.”
The occurrence of this Humming-bird on Juan Fernandez is now
confirmed.
Proc. Zoou. Soc.—1878, No. XXVIII. 28
434 MESSRS. SCLATER AND SALVIN ON THE [Apr. 16,
17. EusTrPHANUS FERNANDENSIS (King); Scl. et Salv. No-
mencl. p. 90; Scl. ‘ Ibis,” 1871, p. 180.
575-580, 582, 583, 585, 585, 588-592, males,
593, 594, males (young), Juan Fernandez.
595-600, 603, 605, females,
Of 575 to 592 it is noted by Mr. Murray—“ All males, eyes
black, feet and bill black ; insects in stomachs. This was the most
abundant bird on the island apparently. It would seem to be the
male of the next lot (595-605).”
Of the last-mentioned lot, Mr. Murray noted, “ I think there are
strong reasons for believing that these are the females of the pre-
ceding. They were several times shot together on the same tree.”
There can be no question of Mr. Murray’s view being correct.
See my notes on this subject in ‘ Ibis,’ 1871, p. 181.
18. CAMPEPHILUS MAGELLANICUS (Gm.); Scl. et Salv. No-
mencl. p. 98.
667, male,
668, female, } Porto Bueno.
«« Eyes yellow, feet and bill black; stomachs had insects.”’
19, CeryxLe sTELLATA (Meyen) ; Scl. et Salv. Nomencl. p. 103.
628, male, Port Otway.
“ Byes black, feet yellowish green; stomach was empty.”
638, male, Corr Harbour.
“ yes brown ; stomach had fish-bones.”’
649, male, Gray Harbour.
‘« Kyes black ; stomach was empty.’
20. Conurus sMARAGDINUS (Gm.); Scl. et Salv. Nomencel.
p- 112.
696, male, HEE
697-699, females, } Sandy Point.
* Eyes red ; stomachs had seeds ; bill and legs horn-colour.”’
21. Orus sracuyortus (Forst.); Sel. et Saly. Nomencl. p. 116.
712, male, Sandy Point.
** Byes yellow, bill black ; stomach had a mouse in 102}
717, male, Elizabeth Island.
* Eyes yellow, cere yellow, bill black; stomach had remains of
mouse.”
22. GrRANOALTUS MELANOLEUCUs (Vieill.); Scl. et Salv. No-
mencl. p. 119.
713, female, Elizabeth Island.
“Eyes brown; stomach had goslings; cere yellow, tip of bill
black, legs yellow.”
23. TINNUNCULUS SPARVERIUS (Linn.) ; Scl. et Salv. Nomencl.
p: 121.
700, male, Sandy Point.
1878. | BIRDS OF ANTARCTIC AMERICA. 435
“* Eyes brown, feet yellow, bill bluish; stomach had remains of
mouse,”
701, female, Sandy Point.
“Eyes brown, legs yellow, cere yellow, bill blue; stomach had
remains of small birds.”
715, 716, females, Elizabeth Island.
“ Eyes brown, legs yellow, cere yellow.”
These specimens all belong to the form kept distinct by Mr.
Sharpe (Cat. Birds, i. p- 439) under Swainson’s title eianamominus.
But when a large series is examined, we find so many intermediate
forms between 7. cinnamominus on the one hand, and T. isabellinus
on the other, that it seems better to keep these three continental
forms all under one specific designation. On the other hand, the
insular forms 7’. leucophrys and T. sparverioides of Cuba, and T.
dominicensis of the other Antilles, seem to be fairly distinct.
24. Mitvaco cuimanco (Vieill.) ; Scl. et Salv. Nomencl. p. (22.
669, 670, females, Porto Bueno.
*« Eyes brown, feet bluish grey.”
695, male, Sandy Point.
‘* Kyes brown, bill yellowish tinge, legs blue.”
25. Potysorus THARUS (Mol.); Scl. et Salv. Nomenel. p. 123,
679, male, Isthmus Harbour.
“Eyes brown, bill blue, cere flesh-coloured, and legs flesh-
coloured.”
684, male, Port Churrucha.
“Eyes brown, cere orange, legs yellow, bill bluish tinge.”
26. CaTHARTES AURA (Linn.) ; Scl. et Salv. Nomencl. p- 123.
nops falklandica, Sharpe, Cat. Birds, i. p- 27, pl. ii.
731, female, Falkland Islands.
“‘ Eyes brown ; had remains of birds in the stomach.”
We consider Mr. Sharpe’s generic and Specific names for this
Vulture alike unnecessary.
Illiger, in his « Prodromus ’ (1811), gave two types for his genus
“ Cathartes,” Vultur papa and V. aura. In 1816 Vieillot made
two genera, Gypagus for V. papa and Catharista for Vultur aura
and V. atratus. Cathartes, however, has been generally used for the
latter group; and there is, in our opinion, no reason whatever for
transferring it back to the former, which has an excellent and
appropriate name in Gypagus. There seem to us no sufticient
characters to separate the red-headed and black-headed species of
Cathartes into two genera.
As regards the Falkland-Island form of C. aura, which Mr.
Sharpe designates Gnops falklandica, the differential character
relied upon, which is, apparently, merely the slight greyish-brown
margination of the secondaries, appears to us to be insufficient.
436 MESSRS. SCLATER AND SALVIN ON THE (Apr. 16,
27. NycTicorax osscurRvs, Bp.; Scl. et Salv. Nomencl. p. 126.
655, female, Tom Harbour.
«* Ryes orange, feet greenish yellow ; stomach had fish.
666, female, Porto Bueno.
«* yes orange.”
737, female, Falkland Islands.
** Kyes yellow ; stomach had crustacea.”’
28. THERISTICUS MELANOPIS (Gm.); Scl. et Salvy. Nomencl.
p. 127.
691, male, é
692, Tancle, f Sandy Point.
Female, eyes red, feet pink, bill black.
Male, eyes yellow, bill green, legs pinkish with black scales.
29. BeRNICLA ANTARCTICA (Gm.) ; Scl. et Salv. Nomenel. p. 128.
632, male,
633, female,
634, young,
635, young,
‘** Eyes brown, feet yellow; bill of male black, of female flesh-
colour ; bill of young dark ; feet grey-brown ; stomachs had pieces
of small stones, shells, and sea-weed.””
736, female, Falkland Islands.
The two goslings (obtained January 1) are just changing their
down, and present the black back, brown head, and barred wings
and undersurface of the adult female partially developed. The sex
is not noted in either of them.
Penguin Islands, Messier Channel.
30. CHLO#PHAGA POLIOCEPHALA, Scl.; Scl. et Salv. Nomencl.
p. 128.
643, male,
G4 deiniehes } Gray Harbour.
‘* Eyes brown, feet and legs yellow and black ; stomachs had grass
and berries.”
658, female,
659, female,
658 a, young,
658 6, young,
‘* Eyes brown, bill black, feet yellow and black.”
Tom Harbour.
31. Marca siBiLaTRIx (Poeppig) ; Scl. et Salv. P. Z.S. 1876,
p- 395.
719, male, Elizabeth Island.
** Byes grey, bill black, feet black ; stomach had sand &c.”
32. SPATULA PLATALEA (Vieill.) ; Scl. et Salv. Nomencl. p. 130.
694, male, Sandy Point.
“ Eyes white, bill black, feet yellow; stomach had pebbles, &c.”’
1878. ] BIRDS OF ANTARCTIC AMERICA. 437
33. TACHYERES CINEREUS (Gm.).
Tachyeres cinereus, Scl. et Salv. P. Z. S. 1876, p. 402.
Micropterus cinereus, Cunningham, Trans. Zool. Soe. vii. p- 493.
651, female (young), Messier’s Channel.
** Eyes brown; stomach had crabs.”
656, young, Messier’s Channel.
‘* Eyes brown, feet and bill black.”’
657, female, Tom Harbour.
“Eyes brown, bill yellow tipped with black, feet yellow; stomach
had small pieces of sand.”
678, male, Straits of Magellan.
681, 682, females, Port Churrucha.
“‘ Weighed 8 lb. and 83 lb.”
725, 726, males (young), Falkland Islands.
‘Stomach had shells trom the kelp chiefly ; bill black, feet
yellow-brown.”
There is nothing in the present series which would induce us to
question Dr. Cunningham’s view, that the “ Flying Loggerhead ” is
the young of the ordinary species before it gets too heavy for flight.
Such a specimen as “No. 681, young female,” could evidently
accomplish flight, which in large fully adult birds would be probably
impossible.
In all the immature birds the bill is uniform black.
34. CotumBaA @nas, Linn.
626, male, J Tend
627, female, ( 242" Fernandez.
*« Eyes orange, feet red, bill black.’’
No doubt an introduced bird; the two specimens vary in plumage.
35. Ratuus anrarcricus, King; Scl. et Salvy. Nomencl. p- 139.
693, female, Sandy Point.
“Eyes red, bill black, feet yellow ; stomach had pebbles.”
36. VANELLUs occIDENTALIS, Harting.
Vanellus occidentalis, Harting, P. Z. 8. 1874, p- 451.
V. cayennensis, Sclater, P. Z. 8. 1867, p. 331; Scl. et Salv. No-
mencl. p. 142 (partim).
722, male, Elizabeth Island.
“ Eyes pink, legs pink, bill black at tip, pink at base.”
The characters given by Mr. Harting to separate this western
form from V. cayennensis are slight, but, we think, on the whole,
sufficient to justify the employment of his name.
37. Ha/MATOPUS LEUCOPUS (Garnot); Scl. et Saly. Nomencl.
p. 143.
636, male, Penguin Islands.
“* Bill red, legs flesh, eyes orange.”
438 MESSRS. SCLATER AND SALVIN ON (Apr. 16,
ae a } Tom Harbour.
04, young,
«* Eyes orange ; stomach had mussels.’
“ Eyes of young were brown, bill of adult red, feet flesh-coloured ;
bill of young brown.”
661, female, Tom Harbour.
720, male, Elizabeth Island.
“Eyes yellow, feet flesh, bill red, eyelids red; stomachs had
mussels.”
38. Hazmarorus ater (Vieill.) ; Scl. et Salv. Nomencl. p. 143.
721, female, Elizabeth Island.
“Byes yellow, feet flesh, bill red, eyelids red; stomach had
mussels.”
39. Eupromias mopesta (Licht.) ; Scl. et Salvy. Nomencl.
p. 143.
646, female, Gray Harbour.
*‘ Ryes brown ; stomach had insects.”’
660, female, Tom Harbour.
«* Byes black ; stomach had insects.”
673, male, Porto Bueno.
*« Kyes brown ; stomach had seeds and sand.”
690, male, Port Fanurie.
** yes brown; stomach had sand.”
728, male, Falkland Islands.
‘Eyes brown ; stomach had sandy particles &c.”’
40. GALLINAGO PARAGUALE (Vieill.); Scl. et Salvy. Nomencl.
p- 144.
671, 672, females, Porto Bueno.
** Eyes brown, feet bluish ; in No. 672 the feet are yellowish.”
729, male,
730, belie Falkland Islands.
“‘ Eyes brown ; stomachs had worms &c.”’
41. Trinea Fruscico. iis (Vieill.) ; Scl. et Salv. Nomenel. p. 145.
733, male, Falkland Islands.
«Eyes brown; stomach had sand &c.”
5. Descriptions of three new Species of Birds from Ecuador.
By P. L. Sctatzer, M.A., Ph. D., F.R.S., and Osserr
Satvin, M.A., F.R.S.
[Received April. 11, 1878.]
(Plates XXVII. & XXVIII.)
Mr. ©. Buckley, during his recent second expedition into the
eastern valleys of the Andes of Quito, has obtained a series of birds
EEE EEE EEE ne ee
1878. | THREE NEW BIRDS FROM ECUADOR. 439
many of which are of great interest, such as Oreothraupsis arre-
monops, Jardine, Cyanocitta pulchra, Lawrence, and others, Three
species, which appear to be new to science, we Propose now to
characterize.
1. Buarremon Levcoris, sp. noy.
Oleagineo-viridis, supra valde saturatior Jeré in nigrum trans-
tens ; alis extus, caudd totd et capitis lateribus nigris, dorso
Sere concoloribus ; piteo castaneo; oculorum ambitu angusto
albo ; rostro nigro, pedibus corneis : long. tota 7-0, ale 3-4,
caud@ 3°3, tarsi 11.
Hab. Yauayaca, Ecuador (Buckley).
Mus. Salvino-Godmannico.
Obs. Species a B. castaneicipite, Scl., cui affinis, dorso obscuro
et ciliis albis prorsus distinguenda.
2. NEoMoRPHUS RADIOLOSUs, sp. nov. (Plate XXVII.)
Cristatus, supra eéneo-niger, frontis et dorsi superioris necnon
tectricum minorum plumis fasciis angustis albis aut fulvis
transradiatis ; tectricibus superioribus et secundariis totis extis
cum dorso inferiore saturate castaneo-rufis ; tectricibus caude
superioribus et caudd supra éneo-viridibus, caudé parum |
Ppurpureo tinctad ; subtis omnino sordidé albus, nigro omnino et
Srequenter transradiatus, crisso nigricante; rostro pallide
corneo, pedibus obscuré corylinis: long. tota 19:0, ale 66,
caude rectr. med. 10:0, ext. 6°8, tarsi 2°7, rostri a rictu lin.
dir. 2-0.
Hab. Intaj, Ecuador (Buckley).
Mus. Salv.-Godm.
Mr. Buckley sends one specimen of this remarkable Cuckoo,
which, though a typical Neomorphus, belongs to a very distinct
species of the genus, and differs from the four previously known
members of this group? in being conspicuously banded on the lower
surface of the body and round the back of the neck. In its bright
chestnut-red wing-coverts it approaches to N. rufipennis (P. Z. S.
1849, Aves, pl. x.), but must constitute a third and separate section
of the genus, as arranged by Sclater.
3. ARAMIDES CALOPTERUS, sp.nov. (Plate XXVIII.)
Supra saturate olivaceus, collo postico et pileo concoloribus ;
dorso inferiore et caudé nigricantibus ; fascia colli utringue
latera occupante et tectricum alarium externarum marginibus
saturate rubris; subtis plumbeus, gula albé; subalaribus
nigris albo transfasciatis ; primariis intis nigricantibus ;
rostro viridi, pedibus rubris : long. tota 13-0, ale 6°5, caude
2°3, tarsi 2°4, rostri a rictu 2-0.
Hab. Sarayacu, Ecuador (Buckley),
Mus. Salv.-Godm.
' Cf. Sclater, P. Z. S. 1866, p. 59, et Lawrence, Ibis, 1873, p. 287.
440 DR. O. FINSCH ON A NEW FINCH. [Apr. 16,
Obs. Proximus A. saracure, sed colli alarumque pictura rubra,
subalaribus albo-nigris neque rufo-nigris, et primariis nigricantibus
nec rufescentibus sané diversus.
This fine Aramides, of which Mr. Buckley has sent two specimens
obtained at Sarayacu on the Rio Bobonassa, may be readily dis-
tinguished from its nearest ally, 4A. saracura, by the characters given
above. Unfortunately, in our “Clavis specierum” of this genus
(P. Z. S. 1868, p. 447) we have used the black and white under
wing-coverts for the primary division of the genus into two sections.
In this respect, however, our new species would come into the
second division, although it is certainly most nearly allied to A.
saracura of the first section.
6. On a new Species of Finch from the Feejee Islands. By
Orto Frnscu, Ph.D., C.M.Z.S., Director to the Bremen
Museum.
[Received March 27, 1878.]
(Plate XXX.)
AMBLYNURA KLEINSCHMIDTI, sp. nov.
Supra viridis, subtus letius tincta ; regione parotica viridi-flava ;
capistro nigro; pileo obscure cyanescente ; tectricibus caude
superioribus lete rubris; subalaribus isabellinis ; rostro et
pedibus corneo-rubris.
Front, fore part of cheeks to the posterior edge of eye, and chin
black ; vertex obscure blue; upper surface dark grass green, the
same as the outer edge of the remiges, which are blackish brown,
like the tail-feathers ; upper tail-coverts splendid scarlet-red ; under-
parts grass-green, much brighter and lighter than the upper parts,
ear-region bright greenish-yellow ; under wing-coverts isabelline ;
bill and feet horny yellow in the skin, in the living bird apparently
flesh-coloured, nails dark brown.
rostr. a
fronte,
c. 100 63 29 14 21 15 French millim.
«ae gsm yn yam gi 82" ¢. 6" English inches.
Hab. Viti-levu, Figi Islands.
Mr. Kleinschmidt, the indefatigable collector of the Museum
Godeffroy, discovered this beautiful new species in the interior of
Viti-Levu, in November 1877 ; and I have great pleasure in naming ©
it after its discoverer. The single specimen sent to me by the
Museum Godefroy is, no doubt, an old bird: the sex is not marked.
This bird belongs to the short-tailed group of Finches, which
Reichenbach separated generically, s. n. Amblynura, but has a more
elongated bill than other species of the genus.
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