1 I E> RARY

OF THE

UNIVERSITY

OF ILLINOIS

5&O.5

.\\

The person charging this material is re-

sponsible for its return on or before the

Latest Date stamped below.

Theft, mutilation, and underlining of books

are reasons for disciplinary action and may

result in dismissal from the University.

University of Illinois Library

DEC21M70

NOV 0 <

L161— O-1096

276 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Schistocarpha oppositifolia (Kuntze) Rydb. — This essentially

tropical species is widely distributed in tropical America, and

according to Rydberg in his account of the genus (N. Amer. Fl.

34: 306. 1927) it extends northward to San Luis Potosi. A recent

collection shows that it occurs, rather unexpectedly, also in Tamau-

lipas: Jaumave, stony river valley, September 30, 1931, H. W.

von Rozynski 76.

Vernonia angusta (Gleason), comb. nov. Eremosis angusta

Gleason, N. Amer. FL 33: 98. 1922.

BOTANICAL SERIES

FIELD MUSEUM OF NATURAL HISTORY

FOUNDED BY MARSHALL FIELD, 1893

VOLUME XI NUMBER 6

REVISION OF THE GENUS COREOPSIS

BY

EARL EDWARD SHERFF

RESEARCH ASSOCIATE IN SYSTEMATIC BOTANY

B. E. DAHLGREN

CURATOR, DEPARTMENT OF BOTANY

EDITOR

PUBLICATION 366

CHICAGO, U.S.A.

OCTOBER 20, 1936

BOTANICAL SERIES

FIELD MUSEUM OF NATURAL HISTORY

FOUNDED BY MARSHALL FIELD, 1893

VOLUME XI NUMBER 6

REVISION OF THE GENUS COREOPSIS

THE LIBRARY OF THE

QCT281936

UNIVERSITY OF ILLINOIS

BY

EARL EDWARD SHERFF

RESEARCH ASSOCIATE IN SYSTEMATIC BOTANY

B. E. DAHLGREN

CURATOR, DEPARTMENT OF BOTANY

EDITOR

PUBLICATION 366

CHICAGO, U.S.A.

OCTOBER 20, 1936

PRINTED IN THE UNITED STATES OF AMERICA

BY FIELD MUSEUM PRESS

REVISION OF THE GENUS COREOPSIS

EARL EDWARD SHERFF

PREFACE

The revision of the genus Coreopsis as set forth in the following

j pages is the outcome of a suggestion received in 1912. Early in

that year Dr. Jesse M. Greenman, then Assistant Curator of

- Taxonomy at Field Museum (now Curator of the Herbarium at

: the Missouri Botanical Garden), selected from the Compositae the

closely related genera Bidens, Coreopsis, and Cosmos as offering

« great promise for the student who should study them painstakingly.

- In turn, each of these genera has been studied by me since then.

3 The results of my investigations in Cosmos have been summarized

I in large part in an earlier paper (Revision of the Genus Cosmos,

I Field Mus. Bot. Ser. 8: 401-447. 1932) and will be concluded in

a subsequent treatment to appear in the North American Flora.

- My studies in Bidens and Coreopsis have been detailed in many

articles which have appeared during more than two decades, particu-

- larly in the Botanical Gazette and the American Journal of Botany,

but also in the Journal of the Washington Academy of Sciences and

the Journal of the Pan-Pacific Research Institution. In the present

state of our knowledge, it is impossible, of course, to render a complete

^ account of all the species in these two genera. Perhaps many,

I especially in Bidens, remain to be discovered. I have, however,

I attempted a revisional survey of each genus which should be as

complete for the entire world as the many facilities offered me for

research would permit. Treatments of the species growing in North

^ America are to appear soon in the North American Flora. A com-

v prehensive revision of the entire genus Bidens will appear in separate

' volumes of the Field Museum Botanical Series. The present paper

attempts a revisional summary of the entire genus of Coreopsis.

No attempt has been made to present an extended history

; of the genus Coreopsis. This history has been somewhat interlocked

5 with that of Bidens and in my monograph of that genus I have

- presented a historical re"sum£ to which readers are here referred. In

addition, numerous references given accompanying the various sec-

^ tions (infra) may be consulted with profit.

Many of the world's principal scientific institutions have assisted

- by lending materials or by permitting access to their herbaria,

279

280 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

libraries, or gardens. My very deep gratitude should be expressed

here to the staffs of all these institutions and particularly to the

following individuals: Dr. Ivan Murray Johnston, Research Asso-

ciate, Arnold Arboretum; Dr. Adolph Engler (deceased), former

Director, Dr. Ludwig Diels, present Director, and Dr. Johannes

Mattfeld, Curator, Botanical Garden of Berlin; Dr. Alfred Barton

Rendle, former Keeper of Botany, and Dr. John Ramsbottom,

present Keeper of Botany, British Museum of Natural History;

Dr. E. De Wildeman, former Director, Dr. Walter Robyns, present

Director, and Dr. P. Staner (until recently Attach^ at the Museum

of the Belgian Congo at Tervueren but now on the staff), of the

National Botanical Garden at Brussels; Dr. Enrico Carano, former

Director, Dr. Giovanni Negri, present Director, and Dr. Renato

Pampanini, former Adjutant and Conservator at the Institute of

Botany of the University of Florence (now Professor of Botany and

Director at the Botanical Institute of the Royal University, Cagliari,

Italy); Dr. John Isaac Briquet (deceased), former Director of the

Conservatory and Botanical Garden of Geneva; Dr. Benjamin

Lincoln Robinson (deceased), former Curator, and Mr. Charles A.

Weatherby, Assistant Curator, Gray Herbarium of Harvard Univer-

sity; Dr. Edgar Irmscher, Curator of the Herbarium of the Botanical

Institute at Hamburg; Sir David Prain, former Director, Sir Arthur

W. Hill, present Director, Dr. Otto Stapf (deceased), former Keeper

of Herbarium, Mr. A. D. Cotton, present Keeper of Herbarium, and

Miss M. L. Green, Botanist, Royal Botanical Gardens of Kew; Dr.

Benjamin Daydon Jackson (deceased), former Secretary of the

Linnean Society of London; Dr. Jesse More Greenman, Curator of

the Herbarium, Missouri Botanical Garden; Dr. Nathaniel Lord

Britton (deceased), former Director, New York Botanical Garden; Dr.

H. LeComte, honorary Professor, Dr. Henri Humbert, Professor, Mr.

Louis Anfray (deceased), former Preparator, and Dr. Paul Danguy,

Vice-Director, Museum of Natural History of Paris; Dr. Philip Munz,

Professor of Botany, Pomona College; Dr. Ira L. Wiggins, Stanford

University; Dr. William R. Maxon, Associate Curator, United States

National Museum; Dr. Richard Wettstein-Westersheim (deceased),

former Director, Botanical Garden and Institute of Vienna.

My work has been carried on from time to time in most of the

above institutions and in numerous others, but principally in the

Field Museum of Natural History. To the authorities of Field

Museum am I indeed indebted for the many courtesies and invaluable

aid extended to me over a long period of time. Especially do I

REVISION OF THE GENUS COREOPSIS 281

thank Mr. Stephen C. Simms, Director, Drs. Charles F. Millspaugh

(deceased), former Curator of Botany, and B. E. Dahlgren, present

Curator of Botany, and Mr. Paul C. Standley, Associate Curator of

the Herbarium, for their assistance in many ways.

Much effort has been made to eliminate typographic errors in

the text. Throughout the preparation of my original manuscript

and final text I have been aided greatly in the proof reading by my

wife, Fern R. Seeley Sherff. It is a pleasure to acknowledge here

her cooperation.

During the progress of my work, I have photographed several

hundred of the more important specimens studied, such as types

and cotypes. Complete sets of these photographs are in my private

collection and in the Herbarium of the Field Museum of Natural

History. Throughout the text the term "cotype" is used to connote

a duplicate of the type, as is usually shown, for example, by the use

of the same collection number.

Genus COREOPSIS: Descriptio

L. Gen. PI. ed. 5. Num. 879. 1754; Sp. PL 907. 1753 (maxima

pro parte); Acispermum Neck. Elem. Bot. 1: 34, No. 64. 1790.

(For other synonyms see under names of various sections.)

Plantae (familiae Compositarum) herbaceae vel saepe fruticosae,

glabrae vel pubescentes. Folia opposita vel rarius alterna, indivisa

atque integra dentatave, vel tripartita, vel semel bis terve ternatim

vel pinnatim dissecta. Capitula mediocria vel majuscula, manifeste

pedunculata vel pedicellata, solitaria vel laxe corymboso-paniculata,

radiata. Involucri bracteae plerumque in 2 raro in 3-4 seriebus

dispositae, basi plus minusve connatae, exteriores plerumque

herbaceae vel submembranaceae adpressae vel saepe patentes,

interiores plerumque majores brunneae vel flavae membranaceae.

Receptaculum planum vel convexiusculum. Flores radii ligulati,

1-seriati, neutri vel rarius styliferi ac fertiles vel steriles, lamina

patente integri vel paucidentati, plerumque flavi rarius rosacei

vel discolores. Paleae planae vel concaviusculae membranaceae

striatae flores tubulosos subtendentes. Flores tubulosi flavi vel

superne colorati, hermaphroditi ac fertiles vel intimi steriles, regulares,

corollae limbo parum ampliati apice breviter 4- vel saepissime 5-

dentati, saepe ad gutturis basim anulo glabro pilosove circumscripti ;

antheris basi integris vel (auriculis minutis) sagittatis; stylorum

ramis apice truncatis vel conicis vel breviter caudato-appendiculatis.

Achaenia obcompressa, orbiculata vel oblonga vel plus minusve

>

•

282 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

oblengo-linearia, saepe 2-alata alls membranaceis vel indurato-

crassatis integris vel fractis vel pectinato-dentatis planis vel incur-

vatis, glabra vel (marginibus praesertim) villosa, epapposa vel

aristis lanceolatis linearibusve glabris erecto-hispidulisve vel 2

dentibus vel 2 paleis papposa (interdum medio apice poculo minuto

coronata), numquam aristis retrorsum hamosis munita.

Type species: Coreopsis lanceolata L.

SECTIONES GENERIS

Sect. I. Electro, (DC.) Blake, Proc. Amer. Acad. n. ser. 41: 337.

1913; pro genere DC. Prodr. 5: 630. 1836. Nos. 1-4. (This and the

next four sections included by Blake under the subgenus Leptosyne

[DC.] Blake, loc. cit. 336. 1913; pro genere DC. Prodr. 5: 531. 1836;

etiam A. Gray, Proc. Amer. Acad. 17: 218. 1882; etiam A. Gray,

Syn. Fl. N. Amer. 1, pt. 2: 299. 1884; pro sect. Coreopsidis 0. Hoffm

in Engler & Prantl, Nat. Pflanzenf. 4, pt. 5: 243. 1890 [exclud

Epilepidem, Coreocarpum, et Acomam].)

Suffruticosae, foliis oppositis, indivisis vel ternatim partitis,

coriaceis, lamina segmentisve ovatis vel lanceolatis. Capitula

solitaria vel paniculato-corymbosa. Involucri bracteae exteriores

circ. 5, plus minusve oblongae; interiores circ. 8, longiores, plus

minusve ovato-oblongae. Flores ligulati circ. 5, plerumque 2-3-

dentati, ligula oblongi vel elliptici, fertiles, tubo pubescentes.

Flores tubulosi limbo raro 4- plerumque 5-dentati, anulo deficiente,

tube pubescenti quam gutture cylindraceo-infundibuliformi brevi

ore; styli ramis apice subulato-appendiculatis. Achaenia valde

obcompressa, glabra, marginata, exteriora lata interiora multo

angustiora, omnia epapposa vel interiora raro jugo aristarum

glabrarum angustarum munita. Plantae mexicanae, centrali

americanae, et haitienses. (Type, Electra mexicana DC. = Coreopsis

mutica var. leptomera.)

Sect. II. Anathysana Blake, op. cit. 339. Nos. 5-7.

Herbae perennes caulibus pluribus e radice lignea foliis oppositis

integris vel pinnatiformibus lobis paucis filiformi-linearibus. Invo-

lucrum ut apud Sect. I sed interius 8-12-bracteatum. Flores ligulati

fertiles. Flores tubulosi saepius exanulati; styli ramis apice incras-

satis breviter appendiculatis. Achaenia ut apud Sect. I, epapposa.

Plantae mexicanae (num. 7 ex insula Socorro). (Type, Leptosyne

mexicana A. Gray=Coreopsis cyclocarpa.}

Sect. III. Tuckermannia (Nutt.) Blake, loc. cit. 340; pro genere,

Nutt. Trans. Amer. Phil. Soc. ser. 2. 7: 363. 1841; pro sect. Lep-

REVISION OF THE GENUS COREOPSIS 283

tosynei, A. Gray, Bot. Calif. 1: 356. 1876; etiam Syn. Fl. N. Amer.

l,pt. 2:300. 1884. Nos. 8 and 9.

Perennes, robustae, foliis alternis carnosis 2-3-pinnatim dissectis,

capitulis magnis. Involucri bracteae subaequales, exteriores 5-8,

lanceolato-oblongae; interiores circ. 12, oblongae. Flores ligulati

magni, fertiles. Flores tubulosi anulo subglabro circumscripti.

Achaenia obcompressa, glabra, anguste alata, epapposa vel raro

marginibus in dentes breves vel aristas currentibus aegre papposa.

Plantae e California, Baja-California, et proximis insulis Oceani

Pacifici. (Type, Tuckermannia maritime, Nutt.=Coreopsis maritima.)

Sect. IV. Pugiopappus (A. Gray) Blake, loc. cit.; Agarista DC.

Prodr. 5: 569. 1836 (nee alior.); pro genere, A. Gray, Pacif. R. Rept.

4: 104. 1857; etiam Proc. Amer. Acad. 8: 659. 1873; pro sect. Lep-

tosynei A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 300. 1884. Nos. 10 and 11.

Annuae, e basi ramosae, foliis 2-3-pinnatfidis plerumque basali-

bus, capitulis mediocribus solitariis, pedunculis elongatis subnudis.

Involucri bracteae exteriores 5-7, interiores circ. 8. Flores ligulati

plerumque styliferi ac fertiles, interdum neutri vel cum stylis brevibus

inclusis, ligula lati ac multistriati. Flores tubulosi anulati, anulo

hirsute. Achaenia dimorpha; radii epapposa, cortice marginata et

ad facies quidem plus minusve costata; disci marginibus longe

villosa, apice jugo aristarum lineari-lanceolatarum sursum setulo-

sarum munita. Plantae Californiae australis. (Type, Agarista

Calliopsidea DC.=Coreopsis Calliopsidea.)

Sect. V. Euleptosyne (A. Gray) Blake, op. cit. 341; pro sect.

Leptosynei A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 299. 1884. Nos. 12

and 13.

Habitu involucroque sect. Pugiopappo similis sed bracteis exte-

rioribus linearis. Flores ligulati glabri, fertiles. Flores tubulosi

anulati ; styli ramis apice incrassatis, breviter appendiculatis. Achae-

nia phelloptera, apice poculo minuto coronata. Plantae Arizonae,

Californiae, et Baja-Californiae borealis. (Type, Leptosyne Doug-

lasii DC.=Coreopsis Douglasii.')

Sect. VI. Pseudo- Agarista A. Gray, Proc. Amer. Acad. 22: 428.

ISGl; Epilepis Benth. PI. Hartweg. 17. 1839. Nos. 14-40.

Plus minusve fruticosae, foliis saepius decompositis segmentis

angustis. Receptaculi paleae cum fructu saepe deciduae, ad hujus

faciem exteriorem applicatae, apice saepius eroso-denticulatae.

Flores tubulosi exanulati, styli ramis apice plus minusve conicis vel

caudato-appendiculatis. Achaenia plus minusve lineari-oblonga,

284 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

exalata sed marginibus sursum villosa, apice biaristata aristis

linearibus vel lanceolatis antrorsum hispidis. Plantae mexicanae et

austro-americanae. (Type, Coreopsis petrophila.)

Sect. VII. Eucoreopsis Nutt. Trans. Amer. Phil. Soc. n. ser. 1:

357. lS4l;Coreopsoides Moench, Meth. PI. 594. 1794; Anacis Schrank,

Denkschr. Akad. Munch. Math. Nat. 5: 5. 1817; Leachia Cass.

Diet. Sci. Nat. 25: 388. 1822; Chrysomelea Tausch, Hort. Canal.

1823; Chrysostemma Less. Syn. Gen. Compos. 227. 1832; sect.

Chrysomelea (Tausch) Nutt. loc. cit.; sect. Gyrophyllum Nutt. op.

cit. 358; sect. Leachia (Cass.) A. Gray, Syn. Fl. N. Amer. 1, pt. 2:

291. 1884. Nos. 41-91.

Herbae, foliis oppositis integris vel pinnatim partitis plerumque

petiolatis. Ligulae plus minusve cuneatae, apice 3-5-lobatae vel

-dentatae. Receptaculi paleae basi latae apicem versus attenuato-

filiformes. Styli rami apice abrupte cuspidati vel conici. Achaenia

fere orbiculata, ad maturitatem incurvata, nonnulla vel omnia

demum papillata vel muriculata, saepe ventris apice basique callum

(hoc quidem in unico capitulo saepe valde polymorphum) ferentia;

pappo 2 dentibus minutis paleaceis vel deficiente. Plantae boreali-

americanae et africanae, rarissime (C. lanceolata) in Asia. (Type,

Coreopsis coronata Hook.=Coreopsis nuecensis.)

Sect. VIII. Silphidium Torr. & Gray ex A. Gray, Syn. Fl. N.

Amer. 1, pt. 2: 294. 1884; pro parte sect. Eucoreopsidis Torr. & Gray,

Fl.N. Amer. 2:341. 1843. No. 92.

Folia opposita, magna, indivisa, serrata. Ligulae integrae.

Styli rami apice ovato-triangulati, breviter mucronati. Achaenia

obovato-oblonga, exalata, apice obsolete bidentata. Plantae e

Georgia ad Boreali-Carolinam. (Type, Coreopsis latifolia.)

Sect. IX. Calliopsis (Reichenb.) Nutt. Trans. Amer. Phil. Soc.

n. ser. 7: 360. 1841; pro genere, Reichenb. Mag. Aesthet. Bot. No.

70. 1823; etiam DC. Prodr. 5: 568. 1836; Diplosastera Tausch, Hort.

Canal. 1823; cf. Flora 1: 83. 1824; sect. Coreoloma (pro minima parte

ex exemplo C. Leavenworthii) et subsect. Calliopsidium (omnino)

Torr. & Gray, op. cit. 346. Nos. 93-100.

Herbae, foliis oppositis raro integris plerumque 1-2-pinnatim

divisis lobis integris. Involucrum exterius parvum. Ligulae plerum-

que bicolores vel quidem rosaceae, apice dentatae. Styli rami apice

truncati ac subpenicillati. Achaenia apice calva vel biaristata,

alata vel exalata. Plantae boreali-americanae, rarissime ex aliqua

parte (C. lanceolata et C. tinctoria) sinenses. (Type, Calliopsis bicolor

Reichenb. = Coreopsis tinctoria.}

REVISION OF THE GENUS COREOPSIS 285

Sect. X. Eublepharis Nutt. Trans. Amer. Phil. Soc. n. ser. 7:

359. 1841; sect. Rabdocaulis Nutt. loc. cit.; sect. Coreoloma Torr.

& Gray, Fl. N. Amer. 2: 346. 1843 (exclud. C. Leavenworthii) ; sect.

Cosmella Torr. & Gray, op. cit. 348 (exclud. C. roseam). Nos. 101-111.

Herbae, foliis oppositis vel alternis, plerumque integris. Ligulae

flavae vel rosaceae, dentatae. Flores disci atro-purpurei vel flavi.

Styli rami truncati vel obtuse conici. Achaenia alata alis fimbriatis

vel pectinato-dissectis. Plantae boreali-americanae praesertim e

parte austro-orientali. (Type, Coreopsis gladiata.)

Sect. XI. Euprestinaria Schz. Bip. in Walp. Repert. 6: 163. 1846.

Nos. 112-114.

Herbae annuae vel perennes. Florum tubulosorum tubus glaber,

styli ramis apice cono hispido superatis. Achaeniorum oblongorum

vel oblongo-lanceolatorum aristae paleaceae lanceolato-lineares utrin-

que ciliis adrectis hispidae. Plantae africanae praesertim abyssini-

cae. (Type, Coreopsis Prestinaria.)

CLAVIS

a. Plantae boreali-americanae antillanaeque.

6. Flores ligulati fertiles.

c. Herbae annuae.

d. Disci achaenia exaristata, phelloptera.

e. Achaenia setis clavellatis numerosis obsita.

12. C. Douglasii.

e. Achaenia non clavellato-setosa 13. C. Stillmanii.

d. Disci achaenia biaristata, longe ciliata.

e. Involucri bracteae exteriores lineares . . 10. C. Bigelovii.

e. Involucri bracteae exteriores late ovatae.

11. C. Calliopsidea.

c. Herbae perennes vel frutices.

d. Folia simplicia vel ternatisecta, lamina vel foliolis sub-

anguste lanceolatis vel latioribus.

e. Capitula numerosa cymoso-paniculata; foliis vel foliolis

nunc plus minusve lanceolatis nunc oblongo-ovatis.

1. C. mutica.

e. Capitula pauca vel solitaria; foliis simplicibus.

/. Folia 2-6 cm. longa.

g. Folia cuneate oblanceolata vel obovata.

2. C. cuneifolia.

286 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

g. Folia anguste lanceolato-oblonga .... 4. C. Buchii.

f. Folia 1-2 cm. longa, ovalia 3. C. parvifolia.

d. Folia vel segmenta linearia vel angustiora.

e. Folia integra vel raro lateraliter 2-lobata, 2-6 cm. longa.

5. C. cyclocarpa.

e. Folia principalia pinnatim lobata.

/. Flores ligulati usque ad 11 mm. longi.

g. Involucrum circ. 6-8 mm. altum, foliorum lobis

lateralibus plerumque 3 jugis ... 6. C. pinnatisecta.

g. Involucrum circ. 4-5 mm. altum, foliorum lobis

lateralibus saepius 1 jugo 7. C. insularis.

/. Flores ligulati saltern 2 cm. longi.

g. Pedunculi dispersi, plerumque 1.5-4 dm. longi.

8. C. maritima.

g. Pedunculi corymbose congregati plerumque 1-2

(saepius usque ad 1.5) dm. longi. .9. C. gigantea.

b. Flores ligulati steriles.

c. Fruticosae.

d. Achaenia glabra apice calva 51. C. cordylocarpa.

d. Achaenia ciliata, biaristata.

e. Folia simplicia 15. C. petrophiloides.

e. Folia 1-3-pinnatisecta.

/. Involucri bracteae interiores 5-6 mm. longae.

14. C. petrophila.

f. Involucri bracteae interiores 6-9 mm. longae.

g. Plantae glabrae vel foliis vix pilosiusculae.

h. Foliorum segmenta ultima 0.6-0.8 mm. lata;

involucri bracteis exterioribus lineari-oblongis.

18. C. Pringlei.

h. Foliorum segmenta ultima 1-5 mm. lata; involu-

cri bracteis exterioribus linearibus.

17. C. rhyacophila.

g. Plantae hispidae 19. C. rudis.

c. Herbaceae.1

d. Stylorum rami termino acuti vel cuspidati.

1 For convenience in making comparisons, the plants under c are keyed as

closely as feasible according to the characters utilized by F. E Boynton in his

treatment of the species of Coreopsis in the southeastern United States (J. K. Small,

Fl. S. E. United States 1274. 1903).

REVISION OF THE GENUS COREOPSIS 287

e. Achaeniorum alae dissecte fimbriato-pectinatae.

101. C. saxicola.

e. Achaeniorum alae angustae vel latae, integrae (interdum

fractae vel scissae) vel (C. congregata) supra quidem

rotundo-lobatae sed numquam fimbriato-pectinatae.

/. Foliorum laminae vel segmenta margine integra.

g. Stylorum rami termino cuspidati; paleis basi latis

superne attenuato-filiformibus; foliorum laminis

simplicibus vel pinnatim lobatis.

h. Perennes; ligulis omnino flavis.

i. Achaenia alata alis patulis vel minime incur-

vatis.

j. Achaeniorum corpora (alis exclusis) circ. 2

mm. longa et circ. 1 mm. lata. 43. C. debilis.

j. Achaeniorum corpora (alis exclusis) 2.5-3.2

mm. longa et saltern 1.4-2 mm. lata.

k. Caules basim versus foliosi, gerentes ramos

longos nudos pedunculoideos.

L Involucri bracteae exteriores angustis-

sime lineares, 5-11 mm. longae.

46. C. heterolepis.

L Involucri bracteae exteriores lanceolatae

vel oblongo-lineares veloblongo-ovatae,

3-8 mm. longae.

m. Folia petiolis inclusis plerumque 3-8

cm. longa, involucri bracteis exteri-

oribus 3-5 mm. longis.

42. C. corninsularis.

m. Folia petiolis inclusis plerumque

usque ad 1.5 dm. longis, involucri

bracteis exterioribus 4-8 mm. longis.

41. C. lanceolata.

k. Caules fere usque ad summam foliosi;

pedunculis moderate brevibus.

I. Foliorum laminae latae, simplices vel

lateraliter 1-5-lobatae.

w. Involucri bracteae subaequales 7-10

mm. longae 45. C. pubescens.

288 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

m. Involucri bracteae exteriores 4-7

(raro -8) mm. interiores 12-14 mm.

longae 44. C. intermedia.

I. Foliorum laminae in segmenta angusta

pinnatim partitae. . . 47. C. grandiflora.

i. Achaenia alata alis valde incurvatis integris

demum calloso-incrassatis. . .48. C. auriculata.

h. Annuae.

i. Ligulae discolores 50. C. nuecensis.

i. Ligulae omnino flavae 49. C. congregata.

g. Stylorum rami termino acute conici; paleis lineari-

bus vel termino vix dilatatis; foliorum laminis

plerumque palmato-lobatis vel -divisis (pro C.

major e var. Oemleri simplicibus).

h. Foliorum laminae petiolatae. . . .91. C. tripteris.

h. Foliorum laminae sessiles.

i. Foliorum laminae infra medium non vel parce

sectae 86. C. palmata.

i. Foliorum laminae usque ad basim partitae.

j. Flores tubulosi corollis flavi.

k. Foliorum segmenta filiformia vel lineari-

filiformia 0.3-1 mm. lata.

87. C. verticillata.

k. Foliorum segmenta latiora. . .90. C. major,

j. Flores tubulosi corollis purpureo-brunnei.

k. Foliorum principalium segmenta pauca,

plerumque 1.5-3.5 mm. lata; internodiis

quam foliis saepius longioribus.

89. C. Delphinifolia.

k. Foliorum principalium segmenta numero-

sa, 0.5-1.5 mm. latis; internodiis quam

foliis plerumque brevioribus.

88. C. pulchra.

f. Foliorum laminae regulariter dentatae.

92. C. latifolia.

d. Stylorum rami termino truncati vel obtuse conici.

e. Achaenia exalata.

/. Ligulae basi rubro-brunneae alibi flavae; disci floribus

corolla atro-rubris.

REVISION OF THE GENUS COREOPSIS 289

g. Involucri bracteae exteriores quam interiores multo

breviores; achaeniis anguste oblongis, tenuibus,

subplanis , 93. C. tinctoria.

g. Involucri bracteae exteriores quam interiores non

plerumque breviores; achaenia obovata, crassa,

convexa 94. C. basalis.

f. Ligulae roseae; disci floribus corolla flavis.

95. C. rosea.

e. Achaenia alata.

/. Achaeniorum alae integrae.

g. Achaenia apice calva vel minute dentata.

h. Ligulae discolores.

i. Achaenia circ. 2 mm. longa, anguste vel late

alata 96. C. Cardaminefolia.

i. Achaenia 2.4-2.8 mm. longa, angustissime alata

(vel interdum tantum marginata).

99. C. Atkinsoniana.

h. Ligulae omnino flavae 97. C. stenophylla.

g. Achaenia perspicue aristata aristis interdum in

maturitate deciduis.

h. Ligulae discolores (specie texana) . . 98. C. similis.

h. Ligulae omnino flavae (specie floridana).

100. C. Leavenworthii.

/. Achaeniorum alae dissecte fimbriato-pectinatae.

g. Folia non juncoidea; ligulis flavis.

h. Foliorum majorum laminae plus minusve oblongae

vel lanceolatae.

i. Folia inferiora quam internodia breviora vel

paulo longiora, opposita. . .103. C. Linifolia.

i. Folia inferiora quam internodia multo longiora.

;'. Foliorum imorum laminae lineares vel lanceo-

latae, apice plerumque mucronatae.

k. Foliorum laminae simplices integraeque;

achaeniis elliptico-oblongis, circ. 4 mm.

longis, ala quam corpore multo angus-

tiore 104. C. longifolia.

k. Foliorum laminae saepe lobatae; achaeniis

anguste oblongis, circ. 5 mm. longis, ala

corpori latitudine aequali.

105. C. falcata.

290 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

y. Foliorum imorum laminae oblongae vel

ellipticae, apice obtusae. . . 106. C. gladiata.

h. Foliorum majorum laminae plus minusve ovatae.

i. Folia opposita; petiolis (ac laminis juvenilibus)

ciliatis 108. C. integrifolia.

i. Folia alterna; petiolis (ac laminis quidem

juvenilibus) eciliatis. . .107. C. Helianthoides.

g. Folia juncoidea, laminis veris deficientibus; ligulis

rosaceis 109. C. nudata.

a. Plantae austro-americanae.

6. Folia indivisa.

c. Folia 0.5-2 cm. lata.

d. Folia oblanceolata, 0.5-1.5 cm. lata.

e. Folia integra, 5-7.5 mm. lata 25. C. oblanceolata.

e. Folia supra medium acriter paucidentata, 0.8-1.5 cm.

lata 25a. C. Woytkowskii.

d. Folia anguste ovato-lanceolata, 1-2 cm. lata.

16. C. Irmscheriana.

c. Folia angustiora.

d. Folia lineari-flagellaria tantum 0.5-1 mm. lata, totam

longitudinem aequaliter angusta 26. C. longula.

d. Folia angustissime spathulato-linearia, 1-2 mm. lata.

27. C. venusta.

b. Folia divisa.

c. Folia primaria minuta, tantum 7-9.5 mm. longa.

d. Folia cum fasciculis axillaribus, quam internodia plerum-

que paulo brevioria; involucri bracteis exterioribus

biserialibus circ. 18 29. C. senaria.

d. Folia sine fasciculis axillaribus, quam internodia saltern

duplo longiora; involucri bracteis exterioribus uniseri-

alibus circ. 8 28. C. nodosa.

c. Folia primaria majora.

d. Capitula pansa ad anthesin circ. 1.1-2 cm. lata; foliorum

laminis vel segmentis 1.5-4 mm. latis.

e. Partes novellae glaucescentes; involucri bracteis exte-

rioribus linearibus quam interioribus dimidio breviori-

bus. 20. C. glaucodes.

e. Nullae partes glaucescentes.

REVISION OF THE GENUS COREOPSIS 291

/. Involucri bracteae exteriores minutae, longitudine

tantum circa tertiae interiorum.

g. Involucri bracteae exteriores ovatae; foliorum seg-

mentis lateralibus cuneatis vel elliptico-obovatis

vel spathulatis, 2-6 mm. latis . . . 21. C. microlepis.

g. Involucri bracteae exteriores lineari-oblongae; foli-

orum segmentis lateralibus anguste linearibus,

1-1.7 mm. latis 22. C. Macbridei.

/. Involucri bracteae exteriores lineari-oblongae, quam

interiores dimidio breviores 24. C. parviceps.

d. Capitula pansa ad anthesin 2.2-5 cm. lata.

e. Folia densissime congregata, ramis fere abditis.

/. Foliorum segmenta lineari-oblanceolata, apicaliter

subobtusa; involucri bracteis exterioribus circ. 8,

oblongis, apice rotundatis, circ. 5 mm. longis et

1.5 mm. latis, uniseriatim dispositis . . 30. C. foliosa.

f. Foliorum segmenta lineari-subulata, apicaliter acuta;

involucri bracteis exterioribus circ. 11, ovatis,

circ. 6 mm. longis et 3.3-4.8 mm. latis, biseriatim

dispositis 31. C. polyactis.

e. Folia plerumque laxius (C. trilobae interdum dense)

disposita, ramis plus minusve manifestis.

/. Foliorum segmenta ultima lanceolata vel oblonga;

involucri 6 bracteis exterioribus 2-3 mm. longis et

1-1.6 mm. latis 32. C. notha.

/. Foliorum segmenta ultima angustiora.

g. Foliorum triloborum segmenta ultima flagellaria,

apicaliter acuta, 1-3.5 cm. longa et plerumque

0.3-0.6 mm. lata.

h. Folia laxissime patentia vel etiam subreflexa.

33. C. capillacea.

h. Folia dense adgregata, suberecta. .34. C. triloba.

g. Foliorum segmenta ultima diversa.

h. Involucra glaberrima 35. C. spectabilis.

h. Involucra (saltern infra) hispida.

i. Kami foliaque hinc illinc resinosa, suaveolentia.

36. C. suaveolens.

i. Kami foliaque diversa (petiolis raro inferne

resinosis).

292 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

j. Involucri bracteae exteriores biseriales, circ.

16 38. C. imbricata.

j. Involucri bracteae exteriores uniseriales,

multo pauciores.

k. Foliorum primariorum multa 2.5-4.5 cm.

longa.

I. Achaenia villoso-ciliata dorso sub palea

glabra sed ad ventrem villosissima.

37. C. Pickeringii.

1. Achaenia villoso-ciliata sed faciebus

glabra 39. C. Townsendii.

k. Folia primaria 1-2.5 cm. longa.

1. Internodia superiora multo folia super-

antia, ligulis siccis brunneo-flavis.

23. C. Killipii.

I. Internodia superiora non vel moderate

folia superantia, ligulis siccis claro- vel

subclaro-flavis 40. C. fasciculata.

a. Plantae africanae.

b. Achaenia valde obcompressa, marginibus saepissime tenuia

alatave.

c. Aristae achaeniorum filiformes ac corporibus breviores, vel

deficientes.

d. Plantae annuae (vel verisimiliter annuae).

e. Folia superne hispida.

/. Achaenia (alis inclusis) circ. 2-3.5 mm. lata.

g. Involucri bracteae exteriores 6-11 mm. longae.

h. Flores ligulati circ. 2.5-3 cm. longi.

76. C. oligoflora (vide etiam 72. C. multifloram).

h. Flores ligulati circ. 1-2 cm. longi. . . 79. C. vulgaris.

g. Involucri bracteae exteriores 3-6 mm. longae.

h. Gapitula numerosa, ad anthesin circ. 2.5-3.3 cm.

lata, demum cum achaeniis maturis ±1.2 cm.

alta et plerumque paulo angustiora, achaeni-

orum corporibus plerumque 7-10 mm. longis,

aristis perspicue densissimeque erecto-hispidis

supra paleas valde manifestis.

80. C. Giorgii (vide etiam 78. C. Goffardii}.

REVISION OF THE GENUS COREOPSIS 293

h. Capitula pauca, ad anthesin ±4 cm. lata, demum

cum achaeniis maturis sub 1 cm. alta et ±1.3

cm. lata, achaeniorum corporibus 3.5-6 mm.

longis, aristis minutissimis ac non supra paleas

manifestis 77. C. Quarrei.

h. Capitula subnumerosa, ad anthesin circ. 4-4.5

cm. lata, demum cum achaeniis maturis circ.

1 cm. alta et parce latiora, achaeniorum cor-

poribus 4.5-6.5 mm. longis, aristis in paleis

subabditis 82. C. injucunda.

/. Achaenia (alis inclusis) 4-5 mm. lata.

81. C. Mattfeldii.

e. Folia superne glabra vel glabrata.

/, Achaenia exaristata 56. C. Negriana.

f. Achaenia biaristata.

g. Foliorum caulinorum saepius 2-3-pinnatisectorurn

segmenta anguste linearia; capitulis pansis ad

anthesin 2.5-5 cm. latis.

h. Foliorum segmenta plerumque circ. 1 mm. lata;

involucro hispido 71. C. leptoglossa.

h. Foliorum segmenta plerumque 2-4 mm. lata;

involucro glabrato vel subglabrato.

69. C. Feruloides.

g. Foliorum 2-pinnatisectorum segmenta principalia

oblongo-lanceolata; capitulis pansis ad anthesin

2.5-4 cm. latis.

61. C. Prestinariaeformis var. /3 incisa.

g. Foliorum caulinorum indivisorum laminae vel 3-5-

partitorum segmenta late linearia; capitulis pansis

ad anthesin 5-7 cm. latis 83. C. Borianiana.

g. Foliorum 3-5-lobatorum foliola ovata vel ovato-

sublanceolata 61. C. Prestinariaeformis.

d. Plantae perennes (vel verisimiliter perennes).

e. Foliorum principalium laminae vel segmenta lanceolata

vel latiora.

/. Foliorum segmenta dentata dentibus perspicue elon-

gatis (saepe 1-1.5 cm. longis) . . . 110. C. Mildbraedii.

f. Foliorum laminae vel segmenta diversa.

. Folia indivisa.

294 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

h. Folia scabra, saepius 10-16 cm. longa; involucre

hispido 85. C. scabrifolia.

h. Folia glabra, 1.5-5 cm. longa; involucre glabro.

62. C. Barteri.

g. Folia divisa.

h. Achaenia exaristata.

i. Folia plerumque pinnata vel pinnatifida seg-

mentis principalibus saepius late oblongo-

lanceolatis vel ovatis, inter se saepe propin-

quis, supra scaberulis subtus tomentellis.

j. Caulis primo tomentellus mox glabratus.

66. C. Whytei.

y. Caulis perpetuo tomentellus.

65. C. pinnatipartita.

i. Folia 2-3-pinnatisecta, foliolis vel segmentis

principalibus saepius anguste oblongo-lance-

olatis, glabris 67. C. Lupulina.

h. Achaenia biaristata.

i. Capitula pansa ad anthesin 5.5-9 cm. lata.

y. Folia pubescentia.

73. C. bella (vide etiam. .72. C. multifloram).

j. Folia faciebus glabra 84. C. togensis.

i. Capitula pansa ad anthesin usque ad 4 cm. lata.

y. Folia molliter denseque pubescentia; achae-

niis plumbeo-nigris minutis corpore tantum

3-4 mm. longis 68. C. Neumannii.

y. Folia glabra vel subglabra.

k. Pauca folia divisa. ...... .62. C. Barteri.

k. Folia plerumque divisa.

Z. Involucrum moderate vel valde hispidum.

m. Involucrum moderate adpresso-his-

pidum; pedunculo tenui glabra-

toque; floribus tubulosis extrinsecus

glabris 75. C. exilis.

m. Involucrum valde hispidum; pedun-

culo subrobusto hispidoque; floribus

tubulosis inferne pilosis.

64. C. oblonga.

I. Involucrum non nisi basi hispidum.

REVISION OF THE GENUS COREOPSIS 295

m. Caulis rigidus superne circ. 2.5-3.5

mm. crassus, foliis principalibus pe-

tiolo adjecto circ. 4-5 cm. longis,

capitulis pansis 4-5.5 cm. latis,

planta kamerunensi . 63. C. monticola.

m. Caulis gracilis superne 1-2 mm.

crassus, foliis principalibus petiolo

adjecto 5-8 cm. longis, capitulis

pansis 3-3.5 cm. latis, planta ango-

lensi 74. C. Curtisii.

e. Foliorum principalium laminae vel segmenta late vel

anguste linearia.

/. Foliorum dentes saepe in setas elongatas desinentes;

lamina indivisa, usque ad 2 dm. longa et 4-10 mm.

lata 111. C. bracteosa.

f. Foliorum dentes non setigerae.

g. Folia inferiora 1-2 dm. longa; involucri bracteae

exteriores circ. 12, lineari-subulatae, 1-1.5 cm.

longae 57. C. camporum.

g. Folia inferiora raro 1 dm. longa.

h. Frutices.

i. Achaenia exaristata.

;'. Segmenta ultima 2-3 mm. lata.

52. C. elgonensis.

j. Segmenta ultima multo angustiora.

53. C. Chippii.

i. Achaenia biaristata 54. C. scopulorum.

h. Herbae.

i. Capitula pansa ad anthesin 4-8.5 cm. lata.

j. Involucri bracteae exteriores elongatae, lat-

eraliter lobatae; ligulis sulphureis.

58. C. Ellenbeckii.

j. Involucri bracteae exteriores normales; ligulis

aurantiacis 70. C. ochracea.

i. Capitula pansa ad anthesin circ. 2-2.5 cm. lata.

;'. Folia atro-viridia, segmentis plerumque

1.2-2.5 mm. latis 59. C. lineariloba.

j. Folia pallida, segmentis plerumque circ. 1

(raro 2) cm. latis 60. C. Schimperi.

296 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

c. Aristae achaeniorum subulatae.

d. Herbae annuae.

e. Involucri bracteae interiores glabrae.112. C. Prestinaria.

e. Involucri bracteae interiores pilis subplanis obtectae.

61. C. Prestinariaeformis et var. /3 incisa.

d. Herbae perennes 113. C. macranlha.

b. Achaenia tantum moderate vel aegre obcompressa.

c. Achaenia exaristata; foliis indivisis spathulato-oblanceolatis-

55. C. Jacksonii-

c. Achaenia biaristata aristis lanceolato-subulatis; foliis bipin<-

natipartitis 114. C. pachyloma et var. 0 inanis.

ABBREVIATIONS USED FOR HERBARIA CITED

Berl. Herb. Berlin Botanical Garden, Berlin-Dahlem.

Boiss. Herb. Boissier, Geneva.

Brit. Herb. British Museum of Natural History, South

Kensington.

Bruss. Herb. National Botanical Garden, Brussels.

Calif. Herb. University of California, Berkeley.

Del. Herb. Delessert, Geneva.

Field Herb. Field Museum of Natural History, Chicago.

Flor. Herb. Institute of Botany, University of Florence,

Italy.

Gray Herb. Gray, Harvard University, Cambridge,

Massachusetts.

Hamb. Herb. Hamburg Botanical Garden, Germany.

Kew Herb. Royal Botanic Gardens, Kew.

Linn. Herb. Linnaeus, Linnaean Society, London.

Mo. Herb. Missouri Botanical Garden, St. Louis.

Mun. Herb. Munich Botanical Garden, Germany.

Mus. Cong. Herb. Musei Congoensis, Tervueren, Belgium.

(Recently merged with the Herbarium of the

National Botanical Garden, Brussels.)

Mus. V. Herb. Museum of Natural History, Vienna.

N. Eng. Herb. New England Botanical Club, Cambridge,

Massachusetts.

N. Y. Herb. New York Botanical Garden, New York City.

REVISION OF THE GENUS COREOPSIS 297

Oxf. Herb. University of Oxford, Oxford.

Par. Herb. Museum of Natural History, Paris.

Phila. Herb. Philadelphia Academy of Natural Sciences,

Philadelphia.

Pom. Herb. Pomona College, Claremont, California.

Stanf. Herb. Stanford University, California.

Stockh. Herb. National Natural History Museum, Stock-

holm.

U. S. Herb. United States National Museum, Wash-

ington.

U. V. Herb. University of Vienna, Austria.

1. Coreopsis mutica DC. Prodr. 5: 571. 1836. ElectraGaleottii

A. Gray, PI. Wright. 110. 1852. C. Galeottii (A. Gray) Hemsl. Biol.

Centr. Amer. Bot. 2: 195. 1881. C. mutica var. genuina Blake,

Proc. Amer. Acad. 52: 55. 1-917.

Foliorum segmenta vel laminae tenuiter lanceolatae vel etiam lineari-

lanceolatae var. 7 leptomera.

Foliorum segmenta vel laminae latiores.

Folia petiolis adjectis tantum circ. 2.5-4 cm. longa.

var. 8 holotricha.

Folia 4-17 cm. longa.

Folia utrinque glabrata C. mutica sensu stricto.

Folia infra dense pubescentia var. /3 subvillosa.

Fruticosa, plus minusve glabra, 0.3-1.3 (etiam usque ad 2) m.

alta. Folia petiolata petiolis anguste alatis 0.6-3 cm. longis, petiolo

adjecto 4-17 cm. longa, simplicia vel trisecta vel saepe tripartita,

lamina segmentisve oblonge lato-lanceolatis vel lanceolato-ovatis

(terminali apice saepe acuminato) utrinque glabratis vel inferne

praecipue ad venas sparsim hispidis, acriter serratis, superne saepe

subcoriaceis. Capitula paniculato-corymbosa, nunc pauca (5-15

pro unica inflorescentia) nunc numerosa (usque ad circ. 60), tenuiter

pedicellata pedicellis saepe glabris 1-4 cm. longis, radiata, pansa

ad anthesin 2-5 cm. lata et 8-17 mm. alta. Involucrum saepe

glaberrimum, bracteis exterioribus circ. 5, nunc oblongis nunc late

angusteve spathulato-oblanceolatis, apice plus minusve mucronatis,

5-12 mm. longis, quam interioribus ovato-oblongis vel rotundato-

ovatis demum saepe dimidio brevioribus. Flores ligulati circ. 5,

lutei, fertiles, ligula oblongi, tubo pilosi, apice 2-4-denticulati, 1-2

cm. longi. Paleae glabrae, oblongo-lineares. Florum tubulosorum

298 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

stylorum rami apice subulati hispidique. Achaenia valde obcom-

pressa, corpore nigra alls stramineo-brunnea, exteriora late oblanceo-

lata alia angustiora, glaberrima, 6-13 mm. longa et 1-3 mm. lata,

apice emarginato calva vel interiora rarissime biaristata.

Type specimen: Collected by F. W. Keerl at Tlapujahua (Tlapa-

coya), northern Puebla, Mexico (Bruss.). DeCandolle cited Martius'

herbarium for his type specimen, but a good-sized and representative

fragment is in DeCandolle's Prodromus Herbarium (Del.).

Distribution: State of Hidalgo, Mexico, southeastwardly through

Oaxaca and Chiapas into Guatemala.

Specimens examined : Baites, Mexico, 1864 (Gray) ; Gust . Bernoulli

196, in thickets, Bano de los Padres, Guatemala, November, 1865

(N.Y.) ; Cassiano Conzattii 1431 , alt. 1,650 meters, Cerro San Antonio,

State of Oaxaca, Mexico, June 26, 1906 (Gray) ; Ehrenberg 354 pro

parte, Mexico (Pom.); Galeotti 2086, Cordilleras of Mexico (Gray,

fragment; one of two types of Electro, Galeottii A. Gray); Ghies-

bregkt 133, State of Chiapas, Mexico (Gray); idem 539, growing 5-6

ft., cold and temperate region, November, 1864-1870 (Gray; Mo.);

Sutton Hayes, large shrub, near Amatitlan, Department of Amatitlan,

Guatemala, July 20, 1860 (Gray); Heyde & Lux 3792, alt. 2,400

meters, Laguna de Ayarza, Department of Jalapa, Guatemala,

1892 (Berl.; Gray; N.Y.); E. W. D. Holway 5, alt. 1,500 meters,

Guatemala City, Guatemala, January 1, 1915 (Gray) ; idem 52, alt.

2,100 meters, San Rafael, Department of Guatemala, Guatemala,

January 9, 1915 (Gray); idem 154, alt. about 1,950 meters, Solola,

Guatemala, January 30, 1915 (Gray); idem 613, Guatemala City,

December 21, 1916 (Gray); W. A. Kellerman 6296 , alt. 1,465 meters,

San Rafael, Department of Guatemala, Guatemala, February 3, 1907

(Field) ;C. G. Pringle 8218, alt. 2,850 meters, Sierra de Pachuca, State

of Hidalgo, Mexico, September 14, 1899 (Berl.; Field; Gray; Mo.;

N.Y. ; Pom.) ; idem 13041 , alt. 2,490 meters, dry, calcareous bluffs near

Metepec Station, State of Hidalgo, September 20, 1904 (Berl. ; Field ;

Gray); C. A. Pur pus 1550, sunny, rocky slopes, Pachuca, State of

Hidalgo, September, 1905 (Field; Gray; Mo.; N.Y.); J. N. Rose,

J. H. Painter, & J. S. Rose 8837, near Tulancingo, State of Hidalgo,

July 22, 1905 (Gray; Mo.; N.Y.; U.S.); Osbert Salvin, alt. 1,590

meters, Volcan del Fuego, Guatemala, August, 1873 (Berl.); H. Von

Tuerckheim II. 2043, alt. 1,400 meters, San Cristobal, Department

of Alta Verapaz, Guatemala, December, 1907 (Field, 2 sheets; Gray;

Mo.; N.Y.).

REVISION OF THE GENUS COREOPSIS 299

Gray wrote, concerning material referred here to this one species

(PL Wright. 110. 1852): "We appear to have two species, which,

however, are not as well marked as could be desired, viz.: 1. Electro,

Mexicana 2. E. Galeottii." Gray's use of "foliis lato-

lanceolatis" for Electro, mexicana indicates that he mistook simple,

broad-leaved forms of Coreopsis mutica (E. Galeottii) for DeCandolle's

Electro, mexicana, which latter had narrow leaves and would best be

designated today as a variety (vide C. mutica var. 7 leptomera, infra).

Coreopsis mutica var. 0 subvillosa DC. Prodr. 5: 571. 1836.

C. mexicana var. hyperdasya Blake, Proc. Amer. Acad. 49: 338. 1913.

Folia infra ubique dense pubescentia, supra venis exceptis glabrata

vel scabriuscula, caulibus et gemmis et inflorescentia piloso-tomen-

tosis lente subglabratis.

Type specimen: Collected by Wilhelm Friedrich von Karwinski

in Mexico (Mun.).

Distribution: State of Hidalgo and southeastwardly through

Puebla into State of Oaxaca, Mexico.

Specimens examined: C. Conzatti 1955, alt. 1,700 meters, Alturas

de San Pablo Hintzo, State of Oaxaca, August 25, 1907 (Field) ; idem

2074, alt. 1,900 meters, Las Sedas, State of Oaxaca, October 20, 1907

(Field) ; idem & V. Gonzalez 545 et 546, alt. 2,000 meters, Cerro de San

Felipe, State of Oaxaca, September 1, 1897 (Gray) ; Ehrenberg 354 pro

parte, between Mineral del Monte and Pachuca, State of Hidalgo

(Berl., 2 sheets); E. W. Nelson 1718, alt. 1,800-2,250 meters, near

Reyes, State of Oaxaca, October 17, 1894 (Gray) ; Nicolas, State of

Puebla, October, 1905 (Field) ; C. G. Pringle 4896, alt. 1,800 meters,

ravines of hills near City of Oaxaca, State of Oaxaca, September,

1894 (Field; Gray, 2 sheets; Mo.; N.Y.; Pom.; U.S.); Charles L.

Smith, alt. 2,100-2,400 meters, Sierra de San Felipe, State of Oaxaca,

October 11, 1894 (Mo.; N.Y.); Lucius C. Smith 25, San Juan del

Estado, State of Oaxaca, June 18, 1894 (Gray); idem 808, alt.

2,130 meters, La Carbonera, State of Oaxaca, September 20, 1895

(Gray).

Coreopsis mutica var. 7 leptomera Sherff, Bot. Gaz. 88: 300.

1929. Electro, mexicana DC. Prodr. 5: 630. 1836. C. mexicana

(DC.) Hemsl. Biol. Centr. Amer. Bot. 2: 196. 1881. C. chrysantha

Sess£ & Moc. Fl. Mex. ed. 2. 194. 1894 (non alior.). Silphium

ternatum Sess£ & Moc. op. cit., 195.

Glabrata; e specie foliorum glabrorum laminis vel segmentis

tenuiter lanceolatis vel etiam lineari-lanceolatis differt.

300 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: Collected by Cyrus Guernsey Pringle, No. 9895,

at altitude of 2,040 meters, clay banks, Dublan, State of Hidalgo,

Mexico, October 15, 1902 (Field).

Distribution: State of Guanajuato eastwardly through Quere*taro

into State of Hidalgo and thence southward into Federal District,

Mexico.

Specimens examined: commun. Alfredo Duges sub num. 472,

Guanajuato, State of Guanajuato, 1895 (Gray); Hort. Hulier 8,

December 5, 1866 and 101, January 9, 1867 (N.Y.); Mendez, about

Villalpando beyond City of Guanajuato, State of Guanajuato (Gray;

cotype oi Electro, mexicana DC.); Pringle 9895 (type, Field; cotypes,

Berl.; Gray; Mo.; N.Y.; U.S.); idem 13547, growing 5 to 6 feet high

at alt. 2,550 meters, barranca above Santa Fe, Federal District,

September 1, 1905 (U.S.); C. A. Purpus 1339, Ixmiquilpan, State of

Hidalgo, September, 1908 (Berl.); J. N. Rose, J. H. Painter, &

J. S. Rose 9717, near Cadereyta, State of Quere"taro, August 22,

1905 (Gray; N.Y.; U.S.); Uhde 627, Mexico.

DeCandolle's Electro, mexicana came from Villalpando, near the

City of Guanajuato, and was a form with simple, narrow leaf blades.

The specimen sent by Duges from the same general locality had

mainly tripartite leaves, a distinction of no diagnostic value in this

species, and, what was indeed significant, the segments were likewise

narrow. A study of these and other collections shows that the form

with narrow leaf blades or leaflets constitutes a rather well-marked

variety occupying a range of greater western but lesser southern

extent than does the species proper.

Coreopsis mutica var. 5 holotricha Blake, Proc. Amer.

Acad. 52: 55. 1917. C. mexicana var. hyperdasya f. holotricha Blake,

op. cit. 49: 338. 1913.

Folia parva circ. 2.5-4 cm. longa, utrinque cinerea pube densa

subscabra.

Type specimen: Collected by Carl Albert Purpus, No. 3099,

vicinity of San Luis Tultitlanapa, State of Puebla, Mexico (Gray).

Distribution: Known only from type locality in State of Puebla,

Mexico.

Specimens examined: Purpus 3099 (type, Gray; cotypes, Berl.;

Field; Mo.; U.S.).

2. Coreopsis cuneifolia Greenm., Proc. Amer. Acad. 40: 43.

1904.

REVISION OF THE GENUS COREOPSIS 301

Fruticosa, verisimiliter 5-9 dm. alta, caulibus rubris griseisve

aegre angulatis, ramulis primo antrorsum subadpresso-pubescentibus

demum nodis exceptis glabratis. Folia sessilia vel subpetiolata,

simplicia, cuneata vel oblanceolato-cuneata, dimidio superiore

dentibus patentibus mucronatisque paucidentata, infra medium

sensim cuneateque usque ad basim angustata, utrinque glabra vel

infra parce pubescentia, glandulo-punctata, 1.5-5 cm. longa et

0.6-2 cm. lata. Capitula solitaria vel paniculato-cymosa pedunculis

tenuibus saepe 1-2 dm. longis, radiata, pansa ad anthesin 1-2.5 cm.

lata et 8-11 mm. alta. Involucri glabri bracteae exteriores circ. 5,

lineari-oblongi, adpressae vel parce patentes, 2-5 mm. longae;

interiores oblongo-ovatae, 5-8 mm. longae. Flores ligulati 5 vel 6,

fertiles, flavi, ligula late oblongi vel cuneato-obcordati, apice nunc

vix nunc profunde dentati, 0.6-1.3 cm. longi, tubo brevi parce

pubescenti. Paleae glabrae, apice obtusae. Florum tubulosorum

(circ. 12) stylorum rami terminaliter angustato-appendiculati.

Achaenia valde obcompressa, circumambitu naviculoidea, glaber-

rima, 7-10 mm. longa et 3-4 mm. lata, fusca vel alis viridi-brunnea,

nitida, apice emarginato calva.

Type specimen: Collected by Joseph Nelson Rose, No. 2344,

State of Durango, Mexico, August 16, 1897 (Gray).

Distribution: State of Durango southward into State of Jalisco,

Mexico.

Specimens examined: C. R. Barnes & W. J. G. Land 165, alt.

1,680 meters, Sierra de San Esteban, State of Jalisco, September 28,

1908 (Field); iidem 313, alt. 1,800 meters, dry hills along road to

Santo Domingo Mine, Etzatlan, State of Jalisco, October 6, 1908

(Berl.; Field); C. G. Pringle 8781, dry, rocky mountains above

Etzatlan, State of Jalisco, October 2, 1903 (Field; Gray; N.Y.);

idem 11900, alt. 1,800 meters, Sierra de San Esteban, near Guada-

lajara, State of Jalisco, October 21, 1903 (Berl.; Field; Gray);

Rose 2344 (type, Gray; cotype, U.S.).

3. Coreopsis parvifolia Blake, Proc. Amer. Acad. 49: 338. 1913.

Fruticosa, juventate adpresse pubescens denique glabrata, veri-

similiter 5-9 dm. alta. Folia subpetiolata, rigida, parva, tantum

circ. 1-2 cm. longa, ovalia, supra adpresse pubescentia infra paulum

crinita vel glabrata, supra partem inferiorem integram late cuneatam

utroque circ, 5-mucronato-dentata, summa imminuta subintegra.

Capitula solitaria ramos elongatos superne subnudos terminantia,

radiata, pansa ad anthesin 2.5-3 cm. lata et 1-1.5 cm. alta. Involu-

302 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

cri bracteae exteriores circ. 5, subcrinitae, oblongo-spathulatae,

obtusae, 4-6 mm. longae; interiores oblongo-ovatae, obtusae, apice

fimbriatae, 8-11 mm. longae. Flores ligulati circ. 5, lutei, ligula

ovales vel oblongi, apice denticulati, circ. 1.2-1.5 cm. longi. Paleae

lineari-oblongae, nitidae. Flores tubulosi 4-dentati, stylorum ramis

apice tenuiter hispido-caudatis. Achaenia immatura plana, oblance-

olata, nitida, alato-marginata, glaberrima, apice calvo emarginata,

8 mm. longa et 2.2 mm. lata.

Type specimen: Collected by Carl Albert Purpus, No. 2581, on

dry, rocky hillsides, Esperanza, State of Puebla, Mexico, August,

1907 (Gray).

Distribution: Known only from type locality in State of Puebla.

Mexico.

Specimens examined: Purpus 2581 (type, Gray; cotypes, Field;

U.S.); idem 2581a, eodem loco, October, 1911 (Berl.).

4. Coreopsis Buchii (Urb.) Blake, Contr. U. S. Nat. Herb.

22: 641. 1924. Selleophytum Buchii Urb. in Fedde, Repert. Sp. Nov.

13: 484. 1914.

Fruticosa, erecta, glabra, 3-10 dm. alta, ramis teretibus obscure

brunneo-rubris plus minusve pruinatis, internodiis 2-4 cm. longis.

Folia opposita, sessilia, 4-7 cm. longa et 1-1.5 cm. lata, anguste

lanceolato-oblonga, glabra, subcoriacea, pallide viridia, subnitida,

margine integra sed interdum aegerrime undulata, basi lata leviter

cordata, supra medium sensim angustata, apice acuta, nervo medio

supra prominente, lateralibus utroque latere 12-16, minoribus inter-

mixtis, utrinque reticulato-conjunctis et prominentibus. Capitula

pauca (solitaria vel terna umbellata), radiata, recte pedunculata

pedunculis circ. 4-6 cm. longis, in juventute globosa nigrescentiaque,

fructifera circ. 2 cm. lata et circ. 1.5 cm. longa. Involucri glabrati

bracteae exteriores 4-5, laxe accumbentes, subcoriaceae, e basi circ.

2-2.5 mm. lata sensim angustatae, apice indurate subacutae, tergo

tenuiter vel obsolete 3-5-nervatae, circ. 8-10 mm. longae; interiores

lanceolato-oblongae, inferne coriaceae superne tenuiores, circ. 12-13

mm. longae et circ. 3-4 mm. latae, apice obtusiusculae. Flores

ligulati flavi, tubo 2.5 mm. longi et (extrinsecus) pilosuli, limbo 8

mm. lati (apice non viso) tergo breviter pilosi. Paleae apice acumi-

natae, pleraeque 3-nervatae, circ. 11-12 mm. longae et circ. 1-1.5

mm. latae. Achaenia oblongo-linearia, plano-obcompressa vel sub-

tetragona, griseo-brunnea, exalata, apice et marginibus et minus

costa interiore breviter denseque setosa, unaquaque facie ±8-sulcu-

lata, corpore marginalia ±7 mm. longa et usque ad 1.3 mm. lata alia

REVISION OF THE GENUS COREOPSIS 303

usque ad 10 mm. longa et 0.6-0.7 mm. lata, apice biaristata aristis

filiformibus antrorsum hispidis circ. 4-5 mm. longis, 1-2 squamellis

interdum interjectis.

Type specimen: Collected by WilhelmBuch, No. 1137, at altitude

of 1,800 meters, in open forests, Morne la Selle, Haiti, January

(Berl.).

Distribution: Known only from Haiti.

Specimens examined: Emery C. Leonard 4103, on cliff near Fond

Parisien, Etang Saumatre, Haiti, May 5-13, 1920 (U.S.).

5. Coreopsis cyclocarpa Blake, Proc. Amer. Acad. 49: 339.

1913. Leptosyne mexicana A. Gray in Wats. Proc. Amer. Acad.

22: 429. 1887.

Herba perennis, 6-7 dm. alta; caulibus pluribus e radice lignea,

infra parce pubescentibus et 1-2 mm. crassis. Folia opposita,

integra vel prope medium rarissime trilobata, lamina (vel segmentis)

lineari-filiformia, basi ciliata, apice acuta, 2-6 cm. longa. Capitula

corymbose disposita, non numerosa, tenuiter pedunculata pedunculis

glabris vel superne sparsim hispidis plerumque 2-7 cm. longis, pansa

ad anthesin 1.5-2.5 cm. lata et circ. 6-8 mm. alta. Involucri bracteae

exteriores 5 vel 6, glabrae, late lineari-oblongae vel ovato-lanceolatae,

crassiusculae, apice acutae vel mucronatae, 2-3 (-3.5) mm. longae;

interiores ovatae vel ovato-lanceolatae, glabratae vel apice fimbriato-

ciliatae, 5-7 mm. longae. Flores ligulati 8-10, flavi, fertiles, ligula

oblongi, apice subprofunde dentati, circ. 1.1-1.3 cm. longi. Paleae

oblanceolatae, apice rotundato-obtusae, 4-5 mm. longae. Flores

tubulosi exanulati, styli ramis apice incrassatis breviter appendi-

culatis. Achaenia valde obcompressa, epapposa; extima (ex floribus

ligulatis) obovata, purpureo-nigrescentia, anguste alato-marginata,

circ. 3.5 mm. longa et circ. 2.2-2.8 mm. lata; alia minora.

Type specimen: Collected by Edward Palmer, No. 568, along a

rivulet in a grassy bottom, Rio Blanco, State of Jalisco, Mexico,

September, 1886 (U.S.).

Distribution: State of Jalisco, Mexico.

Specimens examined: Palmer 568 (type, U.S.; cotype, Gray);

C. G. Pringle 3570, State of Jalisco, September 10, 1890 (Gray);

idem 3841, by streams, etc., near Guadalajara, State of Jalisco,

September 24, 1891 (Berl.; Field, 2 sheets; Gray; N.Y.); idem

11546, alt. 1,500 meters, dry bluffs near Guadalajara, State of

Jalisco, October 4, 1903 (Field; Gray); J. N. Rose & J. H. Painter

7480, near Guadalajara, September, 1903 (N.Y.).

304 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

6. Coreopsis pinnatisecta Blake, Proc. Amer. Acad. 49: 339.

1913. Leptosyne Pringlei Robins. & Greenm. Amer. Journ. Sci.

3. 50: 155. 1895.

Perennis, e radice brevi-oblonga crassiuscula lignosaque, 3-6 dm.

alta, caulibus pluribus, adscendentibus, simplicibus vel subsimplici-

bus, infra foliosis supra subnudatis, striato-angulatis, glabratis vel

infra moderate hispidis. Folia opposita, petiolata petiolis 0.5-1.2

cm. longis, petiolo adjecto 2-3 cm. longa, subrigida, pinnata, adscen-

dentia, sparsim hispida; rhachi et (plerumque 5 vel 7) foliolis linea-

ribus, acriter apiculatis, 0.4-1 mm. latis, foliolis 6-12 mm. longis

inferioribus sensim longioribus. Capitula solitaria vel 1-4-congregata

inflorescentia corymbosa, tenuiter pedunculata pedunculis 5-15 cm.

longis, radiata, pansa ad anthesin 1.5-1.9 cm. lata et circ. 6-7 mm.

alta. Involucri glabrati bracteae exteriores 5 vel 6, late oblongae,

supra saepe latiores, apice obtusae, 3-4 mm. longae; interiores

ovato-oblongae, apice puberulenti subobtusae, circ. 6 mm. longae.

Flores ligulati circ. 6, flavi, ligula oblongi, apice 3-denticulati dente

mediano minore, fertiles, circ. 9-11 mm. longi. Paleae glabrae,

lineari-oblongae, apice subobtusae. Flores tubulosi 5-dentati, infra

anulo hispido instructi, stylorum ramis apice incrassatis breviter

appendiculatis. Achaenia obovata, atra, glabra, circ. 4 mm. longa

et circ. 1.3 mm. lata, apice epapposo rotundata atque emarginata.

Type specimen: Collected by Cyrus Guernsey Pringle, No. 4871,

at altitude of 2,100 meters, on the Sierra de San Felipe, State of

Oaxaca, Mexico, August 7, 1894 (Gray).

Distribution: State of Oaxaca and reaching northward barely

into State of Puebla, Mexico.

Specimens examined: Pringle 4871 (type, Gray; cotypes, Berl.;

N.Y.; Pom.; U.S.); C. A. Purpus 4098, Cerro de Paxtle, vicinity

of San Luis Tultitlanapa, State of Puebla, September, 1909 (Field;

Gray; N.Y.).

7. Coreopsis insularis (Brandeg.) Blake, Proc. Amer. Acad.

49: 340. 1913. Leptosyne insularis Brandeg. Erythea 7: 5. 1899.

Herbacea, probabiliter perennis, decumbens et diffuse ramosa,

fere glabra, forsitan 3-6 dm. alta. Folia opposita, petiolata petiolis

±6 mm. longis, petiolo adjecto 1-2.5 cm. longa, pinnatim plerumque

3- (rarius 5-) lobata, lobis anguste linearibus 0.4-1 mm. latis.

Capitula pauca, tenuissime pedunculata pedunculis glabris saepius

0.8-2 dm. longis, radiata, pansa ad anthesin circ. 1.3 cm. lata et circ.

5-6 mm. alta. Involucri bracteae exteriores circ. 5 vel 6, lineares

REVISION OF THE GENUS COREOPSIS 305

vel lineari-lanceolatae, subglabrae, apice obtusae vel rotundatae,

circ. 2.5-3.5 mm. longae; interiores ovatae vel oblongo-acuminatae,

4-5 mm. longae. Flores ligulati ±5, flavi, fertiles, ligula elliptico-

oblongi, apice 2-lobulati, circ. 7 mm. longi. Paleae lineares. Flores

tubulosi minute 2-5 mm. longi, exanulati, stylorum ramis termino

anguste elongato-conicis. Achaenia extima (ex floribus ligulatis

orta) valde obcompressa, elliptico-obovata, late alata, glaberrima,

3.5-4.2 mm. longa, epapposa; interiora angustiora, anguste marginata.

Type specimen: Collected by A. W. Anthony, No. 394, on Socorro

Island, off coast of Baja California, Mexico, March-June, 1897

(Calif.).

Distribution: Known only from Socorro Island, off coast of Baja

California.

Specimens examined: Anthony 394 (type, Calif.; cotypes, Gray;

U.S.); F. S. Bachelew 223, Socorro Island, May 27-July 3, 1903

(Gray; N.Y.).

8. Coreopsis maritima (Nutt.) Hook. f. Bot. Mag. pi. 6241.

1876. Tuckermannia maritima Nutt., Trans. Amer. Phil. Soc. 2.

7: 363. 1841. Leptosyne maritima (Nutt.) A. Gray, Proc. Amer.

Acad. 7: 358. 1868; etiam Noblet, Rev. Hort. 1873: 330 et pi. (sine

num.). 1873. L. maritima var. gigantea Hort. ex Noblet, loc. cit.

Leptogyne maritima (Nutt.) A. Gray ex Hook. f. loc. cit. (sphalm).

Caules carnoso-herbacei, e radice crassa profunda recta sublig-

neaque, patentes, ramosi, glabri, 3-8 dm. alti. Folia nunc alterna

nunc opposita, petiolata petiolis latis vittatis striatis basi dilitatis

1-7.5 cm. longis, petiolo adjecto saepius 1-1.5 cm. longa, 2-3-

pinnata, segmentis oblonge linearibus, planis, obtusis, 1-2.5 mm.

latis et usque ad 3 cm. longis. Capitula non numerosa, speciosa,

elongate pedunculata pedunculis robustis glabris plerumque 1.5-4

dm. longis, radiata, pansa ad anthesin 6.5-8 (raro usque ad 13.5)

cm. lata et ±1.5 cm. alta. Involucri bracteae exteriores 6-8, oblongae

vel ovato-lanceolatae, obtusae, glabrae vel apice minutissime pubes-

centes, 1-2 cm. longae; interiores oblongae, apice pubescentes, saepius

paulo longiores. Flores ligulati 14-20 (plerumque 16-18), flavi,

ligula elliptico-oblanceolati, apice circ. 3-denticulati, 2-3.5 (-5.5)

cm. longi. Paleae glaberrimae, oblongo-lineares, apice acutae,

demum circ. 1-1.2 cm. longae. Florum tubulosorum stylorum rami

abrupte conico-incrassati vix appendiculati, anulo glabro. Achaenia

anguste oblongo-obovata, plana, nigra nisi alis atro-brunneis, circ.

6-7 mm. longa et alis inclusis 2-3.5 mm. latis, apice glabra vel

interdum alis productis breviter 1 -2-aristata.

306 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: Collected by Edward Tuckerman, Jr., on shelving

rocks, near sea, San Diego, California. Type material not examined,

perhaps still extant among the Nuttall plants (Brit., Kew, or Phila.).

Distribution: South westernmost California; rare northwest-

wardly as far as Monterey County and southwardly in Baja (Lower)

California as far as All Saints' Bay and San Martin Island.

Specimens examined: A. W. Anthony 216, San Martin Island,

Baja (Lower) California, March-June, 1897 (Field; Gray; Pom.);

Margaret Armstrong 741, San Diego, California (N.Y.); T. S.Brande-

gee, San Martin Island, Baja California, April, 1897 (Pom.); idem,

San Diego, California, June, 1900 (Pom.); idem, eodem loco, May,

1906 (N.Y.); Harley P. Chandler 5123, La Jolla, California, April

16, 1904 (N.Y.);F. E. &E. S. Clements 226, cliff edge, dune strand,

eodem loco, March 14, 1914 (Field; Gray; N.Y.); D. Cleveland, San

Diego, California (Gray); R. B. Cowles, moist slope, Coronado

Islands, Baja California, March 29, 1921 (Pom.); G. W. Dunn,

eodem loco, April 17, 1891 (Field) ; Alice Eastwood 2565, Point Loma,

California, April 6, 1913 (Gray) ; Miss F. E. Fish, All Saints' Bay,

Baja (Lower) California, May, 1882 (Gray) ; G. L. Fleming & W. W.

Eggleston 19676, alt. 50 meters, Torrey Pines Reservation, San Diego

County, California, April 10, 1924 (Gray; Pom.); Hort. Bot. Harvard

Univ. cult, annis 1851 et 1852 e sem. a Fremontio lect. (Gray) ; Marcus

E. Jones, Todos Santos Bay, Baja California, April 11, 1882 (Pom.,

2 sheets) ; idem, ranch 29 miles southwest of Tia Juana, Baja Cali-

fornia, April 13, 1925 (Pom., 2 sheets); idem, Coronado Islands,

Baja California, June 10, 1926 (Pom.); idem 3134, Soledad, Cali-

fornia, March 29, 1882 (N.Y., 2 sheets; Pom., 2 sheets); A. J.

McClatchie, San Diego, California, April, 1893 (N.Y.); Philip A.

Munz 7955, sea bluffs, Cardiff, California, May 9, 1924 (Gray; Pom.) ;

C. R. Orcutt, Point Loma, California, March 24, 1884 (Field) ; idem

& W. S. Bowne, San Diego, 1884 (N.Y.) ; C. C. Parry, eodem loco,

1850 (N.Y., 2 sheets, one from the Torrey Herbarium and constitut-

ing the basis of a drawing by Riocreux); idem, eodem loco, 1883

(Gray) ; C. G. Pringle, bluffs of the seashore, eodem loco, April 25,

1882 (Field); F. M. Reed 6147, Potrero Canyon, Baja California,

February 5, 1929 (Pom.); M. S. Snyder, seashore, La Jolla, Cali-

fornia, February 25, 1895 (Field, 2 sheets) ; Mary F. Spencer 86, in

beach sand, vicinity of San Diego, California, April 16, 1916 (Gray;

N.Y.); H. Strauss, San Diego and nearby islands, September 24,

1908 (Berl.) and February 3, 1912 (Berl.) ;L. Street, San Elejo Lagoon,

Baja California, May 12, 1917 (Pom.).

REVISION OF THE GENUS COREOPSIS 307

9. Coreopsis gigantea (Kell.) H. M. Hall, Univ. Calif. Publ.

Bot. 3: 142. 1907; cf. S. B. Parish, Muhlenbergia 8: 133-134. 1913;

etiam C. F. Millspaugh, Field Mus. Bot. 5: 284, pi. 9 and pi. 13, f. 1.

1923. Leptosyne gigantea Kellogg, Proc. Calif. Acad. 4: 198. 1872.

Tuckermannia gigantea (Kell.) Jones, Contrib. West. Bot. No. 15:

74. 1929.

Perennis, erecta, e radice magna plus minusve lignosa, caule

robusto carnoso vix lignoso 5-12.5 cm. crasso et plerumque 3-12

rarius usque ad 20 rarissime usque ad 30 dm. alto, durante per 3-8

annos, ramis glabris, primariis distantibus horizontalibus vel adscen-

dentibus terminos versus foliosis. Folia saepius alterna, petiolata

petiolis glabris carnosis basi plus minusve dilatatis 2-9 cm. longis,

petiolo adjecto saepius 0.8-2 dm. longa, glabra, plerumque 2-3-

pinnata, segmentis nunc moderate (0.8-1.3 mm. latis) nunc

angustissime (0.25-0.4 mm. latis) lineari-filiformibus, integris vel

lateraliter subemarginatis, saepe 3-nervatis, usque ad +5 cm.

longis. Capitula pedunculata pedunculis (ramulis ultimis) sub-

robustis stramineis nudis vel subnudis corymbose congregatis

plerumque 1-2 dm. longis, plerumque radiata, pansa ad anthesin

4-8 cm. lata et circ. 10-13 mm. alta. Involucri glabri bracteae

exteriores 5-7, oblongo-lineares vel lineari-lanceolatae, apice trun-

catae, 1-2 cm. longae, quam interiores 12-15 ovato-oblongae vel

obovatae apice pubescentes nunc paulo breviores nunc vix longiores.

Flores ligulati 10-16, flavi vel interdum apice pallido-flavidi, ligula

anguste vel late oblanceolati, apice circ. 3-dentati, 2-3.8 cm. longi.

Paleae oblongo-oblanceolatae, glaberrimae, apice obtusae, demum

±7 mm. longae. Florum tubulosorum stylorum rami abrupte

incrassati non appendiculati, anulo setis capitatis hispido. Achaenia

subplana, obovato-oblonga vel oblanceolato-oblonga, glabrata, nigra

vel marginibus latis subincrassatis atro-brunnea, omnino (alis

inclusis) 5.5-6.8 mm. longa et 2.5-3.5 mm. lata, apice calva vel alis pro-

ductis obsolete bidentata, saepe poculo minutissimo centrali coronata.

Type specimen: Collected by William George Washington Har-

ford (under auspices of Captain S. Forney, United States Coast Sur-

vey}, near Cuyler Harbor, San Miguel Island, about 40 miles off

coast of Santa Barbara, California.

Distribution : San Luis Obispo to Ventura counties, southwestern

California; also on islands off shore and extending southwardly

as far as Guadalupe Island, off coast of Baja (Lower) California.

Specimens examined (all from California or islands nearby except

where otherwise noted [Palmer 41])' Anon., mouth of Santa Maria

308 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

River, February 7, 1880 (Gray) ; T. S. Brandegee, Island Santa Cruz,

April, 1888 (Gray); Dr. Coulter 316, San Miguel to Santa Barbara

(Gray); M. B. Dunkle 1857, alt. 4.5 meters, shallow soil on rock,

Bird Rock, Santa Catalina Island, April 11, 1928 (Pom.); Alice

Eastwood, Point Sal (near boundary between Santa Barbara and

San Luis Obispo counties), May 12, 1896 (Gray); W. W. Eggleston

19577, alt. 30 meters, Concepcion to Jalama, Santa Barbara County,

March 29, 1924 (Gray; Pom.); A. D. E. Elmer 3634, Surf, Santa

Barbara County, May, 1902 (Field; Gray; Pom.); Roxana S. Ferris

7572, ocean side of sand dunes, 5 miles south of Surf, Santa Barbara

County, April 4, 1929 (Pom.); Grant & Wheeler 680 and 2228,

Catalina Island, April 21-26, 1904 (Field; N.Y.; Pom.); John

Thomas Howell 8220, San Nicolas Island, March 13, 1932 (Pom.);

Marcus E. Jones, Santa Cruz Island, April 13, 1924 (Pom., 2 sheets) ;

idem, Malibu Hills, north of Santa Monica Mountains, April 26,

1926 (Pom.); idem, Santa Cruz Island, March 25, 1929 (Pom.);

Philips Mills Jones, Santa Rosa Island, April, 1901 (Calif. };Ezra C.

Knopf 56, in rocky soil on hillsides, Pebble Beach Road, Santa

Catalina Island, March 26, 1921 (Field) ; idem 258, Bird Rock, Santa

Catalina Island, November 20, 1921 (Field) ; C. F. Millspaugh 4630,

eodem loco, January 22, 1920 (Field); Philip A. Munz 11378, sand

dunes, Surf, May 17, 1929 (Pom.); idem & E. Crow 11553, Santa

Cruz Island, April 7, 1930 (Pom.); iidem 11691, seedlings on shaded

wall of Water Canyon, Santa Rosa Island, April 8, 1930 (Pom.);

iidem 11750, shaded canyon wall northwest of ranch house, Santa

Rosa Island, April 9, 1930 (Pom.); Munz & J. Voss 11877, high

exposed cliff, San Miguel Island, April 10, 1930 (Pom.); L. W.

Nuttall 571, on sea cliffs, near Avalon, Santa Catalina Island, May

14, 1920 (Field); Edward Palmer 41, Guadalupe Island, off Baja

(Lower) California, 1875 (Berl.; Gray); Parry, San Diego (Gray);

Miss Plummer, mountains above Santa Barbara, 1878 (Gray);

Huron H. Smith 5034, road to Pebble Beach, Santa Catalina Island,

May 30, 1912 (Field) ;Blanch Trask 76, San Nicolas Island, April, 1901

(Gray, 2 sheets) ; W. G. Wright, Ventura County, March, 1894 (Field).

10. Coreopsis Bigelovii (A. Gray) Voss in Sieb. & Voss, Vil-

morin Blumeng. ed. 3. 1: 488. 1894 (where spelled C. Bigelowii).

Pugiopappus Bigelovii A. Gray, Pacif. R. Rept. 4: 104. 1857.

P. Breweri A. Gray, Proc. Amer. Acad. 8: 660. 1873. Leptosyne

Bigelovii A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 300. 1884. L. Hamiltonii

Elmer, Bot. Gaz. 41: 323. 1906. Coreopsis Bigelovii (A. Gray)

H. M. Hall, Univ. Calif. Publ. Bot. 3: 141. 1907.

REVISION OF THE GENUS COREOPSIS 309

Herba annua, erecta, 1-4 dm. alta, caulibus saepius pluribus vel

numerosis, simplicibus vel inferne ramosis, basim versus foliosis,

supra multum elongatis ac nudis pro pedunculis, glabris. Folia

saepe dense congregata, plus minusve alterna, petiolata petiolis

usque ad 6 cm. longis, petiolo adjecto 5-11 cm. longa, plerumque

1-2-pinnata, segmentis ultimis linearibus, planis, obtusiusculis, sub-

membranaceis, glaberrimis vel rarius vix scabrido-hispidis, plerum-

que 0.6-1.7 mm. latis. Capitula caules (pedunculos) terminantia,

radiata, pansa ad anthesin 2-4.2 cm. lata et 0.8-1 cm. alta. Involucri

glabri bracteae exteriores circ. 8, lineares, apice obtusiusculae ac

nitido-induratae, 4-9 mm. longae; interiores oblongo-ovatae,

moderate longiores. Flores ligulati 8, flavi (interdum infra intense

supra aegre), ligula elliptico-oblanceolati vel anguste obovati, apice

dentati, fertiles, 1-2 cm. longi. Paleae lineari-oblongae vel navi-

culoideae, obtusae vel subacutae, glabrae, apice minute eroso-ciliatae,

demum circ. 7-9 mm. longae. Flores tubulosi anulo glabri, stylorum

ramis abrupte terminantibus et vix appendiculatis. Achaenia

dimorpha exteriora (e floribus ligulatis) valde obcompressa, cuneato-

obovata, rubro-nigra, glabrata, faciebus valde rugosa, marginibus

anguste calloso-alata, apice exaristata, 4-5 mm. longa et 2-3 mm.

lata; interiora (e floribus tubulosis) angustiora, oblongo-linearia,

faciebus nitidis nigra, tergo glabrata ventre erecte albido-villosa,

marginibus densissime erecto- et albido-villosa, corpore 4-6.2 mm.

longa et ±1.5 mm. lata, apice biaristata aristis lanceolatis planis

densissime erecteque albido-ciliatis circ. 2-3 mm. longis, inter aristas

poculo membranaceo vix manifesto coronata.

Type specimen: Collected by John Milton Bigelow, Mojave

Creek, California, 1853-1854 (Gray).

Distribution : California.

Specimens examined: R. Bacigalupi 1209, alt. 540 meters, just

above Oak Grove on the Mineral King Road, Tulare County, April

23, 1925 (Pom. ; forma elatior) ; Bigelow (type, Gray) ; W. H. Brewer

241, San Buenaventura, 1860-1862 (Gray; sub nom. Pugiopappo

Breweri} ; Ira W. Clokey & B. Templeton 4520, alt. 350 meters, dry

open hillside, Mandeville Canyon, Santa Monica Mountains, March,

1929 (Pom.) ; Frederick V. Coville & Frederick Funston 742, alt. 800-

2,000 meters, Shepherd Canyon, Argus Mountains, April 30, 1891

(Berl.; Field; Gray); Craig, Newsom, & Hiland 80, at 35 miles south

of Randsburg, Mojave Desert, April 5, 1927 (Pom.); W. R. Dudley

340, region of Tehachopi Peak, Tehachopi Mountains, June 25,

1895 (N.Y.); Alice Eastwood, Kaweah, April 27, 1895 (Gray); H. C.

310 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Ford, Santa Barbara County, 1889 (Gray) ; G. B. Grant 5424, Saugus,

April 23, 1903 (Pom., 2 sheets) ; H. M. Hall 3074, alt. 1,050 meters,

vicinity of Elizabeth Lake, Antelope Valley, southern California,

May 1-3, 1902 (Pom.); A. A. Heller 7662, Kern Canyon, Kern

County, April 12, 1905 (Field; Gray); C. L. Hitchcock 12340, near

brush, open desert, 10 miles southwest of Shoshone, April 17, 1932

(Pom.); Marcus E. Jones, alt. 1,380 meters, Darwin, April 28, 1897

(Pom.); idem, alt. 1,380 meters, Shepherd's Canyon, Argus, Moun-

tains, April 30, 1897 (N.Y.; Pom.); idem, alt. 1,800 meters, Pleasant

Canyon, Panamint Mountains, May 6, 1897 (Pom.); idem, alt.

1,140 meters, Cajon Pass, May 16, 1903 (Pom., 2 sheets); idem,

Cave Spring, March 12, 1924 (Pom.); William L. Kennedy, Fort

Tejon, spring of 1876 (Field) ; J. G. Lemmon, top of Mount Diablo

(Field); idem 138, Mojave Desert, 1880 (Gray); Philip A. Munz

4466, alt. 1,140 meters, dry slope near south side of summit, Cajon

Pass, southern California, April 27, 1921 (Pom.); idem 9136, alt.

900 meters, dry slope, Tule River, Tulare County, March 22, 1925

(Pom.); idem 9186, alt. 900 meters, disintegrated lava, north of

Parkfield, Monterey County, March 24, 1925 (Pom.); idem 10042,

sandy desert, 5 miles south of Willow Springs, April 4, 1926 (Pom.);

idem, R. D. Harwood & L M. Johnston 4077, alt. 1,200 meters,

clearing in chaparral, north side of Cajon Pass, May 25, 1930 (Pom.);

Munz & D. Keck 7842, gravelly wash, 7 miles east of Daggett, April 6,

1924 (Pom.); iidem 7879, sandy wash, 10 miles southwest of Garlic

Springs, Mojave Desert, April 8, 1924 (Pom.); Samuel B. Parish,

Marengo, April, 1882 (Field) ; idem & W. F. Parish 226, Whitewater

River, March, 1881 (Berl. ; Field) ; C. C. Parry, Pacific Coast of North

America, 1880 (Field); idem 115, eodem loco, 1881 (Field); idem &

J. G. Lemmon 184, southern California, etc., 1876 (Field); C. G.

Pringle, hills bordering the Mojave Desert, May 11, 1882 (Field;

Gray); C. A. Purpus 5063, alt. 300-600 meters, sunny hillsides,

Middle Tule River, April-September, 1897 (Gray); idem 5531,

alt. 1,200-1,500 meters, hillsides near Erskin Creek, southeastern

California (Gray); Marjorie Shaw 1941, Upper Cajon Pass, May,

1917 (Pom.); eadem, Edna Spaulding & Mrs. C. L. Walton, alt.

900 meters, Antelope Valley, Mojave Desert, April 6-8, 1917 (Pom.);

Mary F. Spencer 407, summit of Cajon Pass, San Bernardino

County, April 27, 1917 (Gray; Pom.); eadem 1954, alt. 960 meters,

Mojave Desert, May 1, 1922 (Gray; Pom.); eadem 2040, rocky

places, Maillard Canyon, near Banning, April 23, 1922 (Pom.);

William Trelease, Hesperia, April 10, 1892 (Mo., 2 sheets) ;G. R. Vasey

REVISION OF THE GENUS COREOPSIS 311

290, Branch of Mojave River, 1880 (Field; Gray); W. G. Wright,

San Bernardino, 1880 (BerL); idem 172, Colorado Desert, March,

1881 (Gray).

11. Coreopsis Calliopsidea (DC.) A. Gray in Torr. Bot. Mex.

Bound. 90. 1858. Agarista Calliopsidea DC. Prodr. 5: 569. 1836;

Regel, Gartenflora 23: 356, pL 816. 1874. Pugiopappus Calliopsidea

(DC.) A. Gray, Proc. Amer. Acad. 8: 660. 1873; Bot. Calif. 1: 355.

1876. Leptosyne Calliopsidea (DC.) A. Gray, Syn. Fl. N. Amer. 1,

pt. 2: 300. 1884. L. Calliopsidea var. nana A. Gray, loc. cit.

Herba annua, erecta, glabra, 1-6 dm. alta, caule tereti simplici

vel ramoso nunc usque ad medium (vel ultra) nunc tantum ad basim

folioso. Folia alterna, petiolata petiolis alatis usque ad 2 (raro -5)

cm. longis, petiolo adjecto 1-7 cm. longa, 1-2-pinnatifida rhachi

lobisque planis linearibusque plerumque 0.6-1.7 (raro -5) mm. latis.

Capitula ramos elongates aphyllos terminantia, solitaria, radiata,

pansa ad anthesin 2-5 cm. lata et circ. 9-12 mm. alta. Involucri

bracteae exteriores 4-6, oblongo-ovatae vel saepius ovato-cordatae,

apice obtusae, angustissime pallido-marginatae, tergo longitudinaliter

4-13-striatae, 4-7 mm. longae; interiores circ. 8, oblongo-ovatae,

apice acutae vel pubescentes, 6-13 mm. longae. Flores ligulati

plerumque 8, flavi, tubo clavo-setosi, ligula late cuneato-obovati,

apice plus minusve truncati et irregulariter multidenticulati, 1-2.5

cm. longi. Paleae lineari-oblanceolatae, longitudinaliter nitido-

striatae striis interrupte brunneis, supra minutissime eroso-ciliatae,

plerumque 4-7 mm. longae, cum achaenio deciduae, ad faciem

hujus exteriorem applicatae. Flores tubulosi corolla 5-dentati,

basim versus clavo-setosi, anulo pubescentes, stylorum ramis termi-

naliter brevi-conici. Achaenia plana dimorpha, exteriora (ex floribus

ligulatis) obovato-oblonga faciebus nitidis purpureo-nigrisque glabra

vel sparsim pulverulenta, margine subanguste alata ala parce in-

crassata, apice calva, omnino ±4.5 mm. longa et ±2.2 mm. lata;

interiora (ex floribus tubulosis) lineari-oblanceolata vel cuneato-

oblonga, nigra, tergo nitida glaberrimaque, marginibus ventreque

erecte elongate- villosa, corpore 4-5 mm. longa, apice biaristata

aristis circ. 3-4 mm. longis densissime antrorso-hispidis.

Type specimen: Collected by David Douglas in California (Del.).

Distribution: Southern half of California.

Specimens examined: Anon., loco ignoto (BerL); Townsend S.

Brandegee, Alcalde, Fresno County, April, 1891 (Field); Craig,

Newsom & Hiland 129, Randsburg, April 5, 1927 (Pom.); David

312 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Douglas, California (Gray); idem (distrib. Herb. Soc. Hort. Lond.),

California, 1833 (Berl.; doubtless a cotype) ; Alice Eastwood, Cuyama,

White Hills, May 2, 1896 (Gray) ; eadem 3187, Mojave, Kern County,

May 12, 1913 (Gray) ; J. B. Feudge 1382, alt. 750 meters, open desert,

Kramer, Mojave Desert, May 25, 1926 (Pom.); Harvard Univ.

Bot. Gard., cult., 1871 (Gray); H. E. Hasse, Mojave Station, Kern

County, March 15, 1889 (Field); A. A. Heller 7723, gravelly hill-

sides, Sunset, April 20, 1905 (Field; Gray); Ivan M. Johnston,

rather common, sandy soil, near Mojave, May 5, 1920 (Pom.);

idem, common, 1 mile south of Crutts Post Office, Mojave Desert,

May 14, 1922 (Pom.); Marcus E. Jones, Cuyama Canyon, April 28,

1926 (Pom.); idem, Adelanto, May 12, 1927 (Pom.); idem, alt. 900

meters, Mojave, May 20, 1903 (Pom.); A. Kellogg & W.G. W. Har-

ford436, California (Gray) ; J. G. Lemmon (137?), California (Gray;

type of var. nana A. Gray) ; idem 4520, Lemmon's Ranch, Cholame,

June, 1887 (Gray); S. B. Parish 9771, Mojave Station, Kern County,

April 26, 1915 (Gray); Parry, California (Gray); W. M. Pierce,

desert, 20 miles east of Victorville, April 12, 1922 (Pom.); C. G.

Pringle, Mojave Desert, May 14, 1882 (Field; Gray).

12. Coreopsis Douglasii (DC.) H. M. Hall, Univ. Calif.

Publ. Bot. 3: 140. 1907. Leptosyne Douglasii DC. Prodr. 5: 531.

1836. L. calif ornica Nutt. Trans. Amer. Phil. Soc. ser. 2, 7: 363. 1841.

L. Newberryi A. Gray, Proc. Amer. Acad. 7: 358. 1868.

Herba annua, glabra, 1.5-4 dm. alta, caule brevi erectoque

ramoso ramis (pedunculis) paucis basim versus foliosis aliter nudis

vel raro folio valde imminuto instructis, apice monocephalicis.

Folia (subradicalia) petiolata petiolis plerumque 2-5 cm. longis,

petiolo adjecto 3-10 (-15) cm. longa, filiformi-linearia integraque

vel saepius 2-5-partita lobis lineari-filiformibus apice obtusis.

Capitula (solitaria, elongatissime pedunculata) radiata, pansa ad

anthesin 2-3.3 cm. lata et 6-8 mm. alta. Involucri glabrati bracteae

exteriores 5-16, lineari-attenuatae vel lineari-oblongae, apice obtusis-

simae, 4-8 mm. longae; interiores ovatae, apice acutiusculae, paulo

longiores. Flores ligulati circ. 8, fertiles, flavi (vel apicem versus

saepe aibidi), ligula oblongi vel ovato-subrotundati, apice circ.

3-dentati dentibus saepe magnis rotundatisque, plerumque 1-1.8

cm. longi. Paleae lineares vel lineari-oblongae, apice obtusae,

demum 3-6.5 mm. longae. Flores tubulosi anulo plerumque hispido

instructi, stylorum ramis cono hispido lato brevique terminatis.

Achaenia obcompressa, cuneato-obovata, diverse rubro-brunnea vel

nigro-rubra, primo (sed non demum) nitida, marginibus crasso-

REVISION OF THE GENUS COREOPSIS 313

alata, duabus faciebus setis clavo minimo similibus instructa, 2-3

mm. longa et 1.2-1.5 mm. lata, apice exaristata sed poculo minuto

membranaceoque coronata.

Type specimen: Collected by David Douglas in California (Del.).

Distribution: Southern Arizona westward into southern Cali-

fornia and reaching southward slightly into Baja (Lower) California,

Mexico.

Specimens examined (all from California except where noted to

the contrary) : LeRoy Abrams 3342, Moro Hills, Fallbrook, April 28,

1903 (Gray); idem 3628, dry hills between Campo and Jacumba,

May 27, 1903 (Gray; Pom., 2 sheets); idem 11788, sandy areas in

hills between Rosamond and Mojave, April 30, 1927 (Pom.); idem

11915, between Red Rock Canyon and Walker Pass Road, El Paso

Range, May 1, 1927 (Pom.); C. F. Baker 4090, Claremont, April 15,

1903 (Field; Gray; Pom.); H. N.Bolander, Julian City near Golden

Cherry Mine (Gray); Katharine Brandegee, San Diego, April 25,

1900 (Calif.) ; T. S. Brandegee'3368, common on the mesas, San Diego,

April 26, 1903 (Field; Gray; Pom., 2 sheets);*1. E. & E. S. Clements

224, La Jolla, March 7, 1914 (Field; Gray); iidem 225, eodem loco,

March 17, 1914 (Calif.; Gray); D, Cleveland, San Luis Rey, 1877

(Gray); Mrs. A. L. Coombs, Warner's Springs, San Diego County,

spring of 1911 (Gray) ; Dr. Coulter 339, California (Gray) ; Douglas,

California (cotype, Gray) ; AliceEastwood 2574, Warner's Hot Springs,

San Diego County, April 9, 1913 (Gray); eadem 3194, Mojave, Kern

County, May 13, 1913 (Gray); eadem 4377, Blochman's Ranch,

Mariposa County, April 21, 1915 (Gray); W. W. Eggleston 19748,

alt. 850-950 meters, Jacumba Springs, San Diego County, April

11-16, 1924 (Pom.) ; Roxana S. Ferris 7466, sandy flat between Coso

Hot Springs and Coso Junction, Kern County, April 30, 1928

(Pom.); eadem 7710, about 5 miles from Mojave or the Willow

Springs Road, Kern County, May 3, 1929 (Pom.); George B. Grant

681, Altadena, April, 1902 (Field, 2 sheets) ; idem 682, eodem loco et

tempore (Gray); Harvey M. Hall 3973, northwest corner of Baja

California, Mexico, April, 1903 (Field) ; Harvard University Botan-

ical Garden, cult., 1882 (Gray) ; H. E. Hasse, Los Angeles, April 22,

1888 (Field); A. A. Heller 7674, Kramer, April 13, 1905 (Field;

Gray); C. L. Hitchcock 12224, sandy wash, 10 miles southeast of

White Tanks, East Riverside County, April 9, 1932 (Pom.) ; Edmund

C. Jaeger 140, Cabazon, April 11, 1920 (Pom.); Ivan M. Johnston,

among creosote bushes in open sandy areas, near Mojave, May 5,

1920 (Pom.); idem, on hillside, Victorville, May 15, 1920 (Pom.);

314 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

idem, alt. 870 meters, in sandy wash, 5 miles southeast of Victorville,

May 17, 1920 (Pom.); idem 1251, alt. 600 meters, field above

Claremont, March 29, 1917 (Pom.); Marcus E. Jones, Pasadena,

February 22, 1882 (Pom.); idem, Fallbrook, March 25, 1882 (Pom.);

idem, San Diego, April 3, 1882 (Pom.) ; idem, alt. 300 meters, White-

water, May 11, 1903 (Pom., 2 sheets) ; idem, alt. 880 meters, Victor,

May 17, 1903 (Pom., 2 sheets); idem, San Diego, April 14, 1906

(Pom., 2 sheets); idem, alt. 1,200 meters, Julian, April 17, 1906

(Pom.); idem, near Santa Ana Canyon, March 10, 1926 (Pom.); idem

3361 pro parte, Pasadena, May 2, 1882 (N.Y.; Pom., 2 sheets);

idem 3373, eodem loco et tempore (Berl.); William Kellogg, Rancho

Encinal, Monterey County, April 15, 1903 (Gray) ; J. G. Lemmon,

San Bernardino, March, 1880 (Gray) ; Philip A. Munz 2148, foothills,

Corona, April 27, 1918 (Pom.); idem 2538, alt. 750 meters, desert,

Barstow, April 12, 1919 (Pom.) ; idem 3814, alt. 300 meters, dry slope,

Santa Ana River Canyon near Rincon, April 11, 1920 (Pom.); idem

9357, alt. 480-510 meters, dry bottom of Cobal Canyon, north of

Claremont (Gray; Pom.); idem 12415, open desert, 20 miles north

of Box S Ranch, San Bernardino County, April 21, 1932 (Pom.);

idem & D. Keck 7930, sandy desert 20 miles west of Barstow, April

10, 1924 (Gray; Pom.); Philip A. Munz & R. D. Harwood 3385, alt.

360 meters, gravelly wash, Yemescal Canyon, March 20, 1920 (Pom.) ;

iidem 3475, alt. 900 meters, sand, Victorville, March 28, 1920

(Pom.); Philip A. Munz, L. Street, & G. Williams 2362, alt. 535

meters, roadside, San Gorgonio Pass, near Cabazon, March 28,

1919 (Pom.) ; iidem 2388, eodem loco et tempore (Pom.) ; Newberry,

Sitgreaves Pass, southern Arizona, March 26 (Gray; type of Lep-

tosyne Newberryi A. Gray) ; Thomas Nuttall, sine loco vel tempore

(Gray) ; Charles R. Orcutt, Point Loma, San Diego, March 12, 1884

(Calif.); idem, northern Baja California, April 17, 1886 (Field);

Edward Palmer 126, Camp Grant, May 10, 1867 (Gray; the specimen

cited by A. Gray for his Leptosyne Newberryi); idem 217, Mojave

River, southeastern California, May, 1876 (Field); idem 677,

San Quentin Bay, Baja California, January, 1889 (Field; Gray);

S. B. Parish 2211, Whitewater Canyon, April, 1880 (Gray); idem

5191, alt. 300-750 meters, vicinity of San Bernardino, April 13,

1903 (Field); idem 10415, Mojave Station, Kern County, March

25, 1915 (Gray) ; idem & W. F. Parish, San Bernardino, April, 1881

(Field); iidem, foothills of San Bernardino Mountains, May, 1888

(Field); C. C. Parry, San Bernardino (Gray); idem 116 (Gray);

idem & J. G. Lemmon 185, southern California, etc., 1876 (Field;

Gray); W. M. Pierce, desert, 20 miles east of Victorville, April 12,

REVISION OF THE GENUS COREOPSIS 315

1922 (Pom.); M. S. Snyder, La Jolla, March 15, 1895 (Field); Mary

F. Spencer 1 72, in hollows of granite rocks, San Diego County, April

8, 1917 (Gray; Pom.); eadem 563, desert sand, Hesperia, Mojave

Desert, May 4, 1917 (Gray; Pom.); eadem 653, alt. 750 meters, in

rocky places, Maillard Canyon, April 22, 1922 (Gray) ; eadem 653a,

in rocky places, Snow Creek near White Water, Mojave Desert,

April 10, 1922 (Gray); eadem 2123, alt. 750 meters, in rocky places,

Maillard Canyon near Banning, April 23, 1922 (Gray); Wallace,

Los Angeles, 1854 (Gray).

The exterior involucral bracts, when few and more oblong, would

seem to connote a variety. Too many apparently transitional forms

have been encountered in herbaria, however, to permit as yet

definite conclusions to be reached.

13. Coreopsis Stillmanii (A. Gray) Blake, Proc. Amer. Acad.

49: 342. 1914. Leptosyne Stillmanii A. Gray in E. Durand, Journ.

Acad. Nat. Sci. Phila. 3: 91. 1855; Torr. Bot. Mex. Bound 2: 92. 1859.

Involucri bracteae exteriores 5-8, oblongae, 4-8 mm. longae; achae-

niis 3.5-4 mm. longis. .-.' C. Stillmanii.

Involucri bracteae exteriores 8-16, tenuissime lirieares superne usque

ad apicem sensim angustatae, 7-12 mm. longae; achaeniis

4-4.5 mm. longis var. /8 Jonesii.

Herba annua, erecta, glabra, 1.5-4.5 dm. alta, infra ramosa ac

foliosa, ramis (interdum numerosissimis) in pedunculos nudos

erectos monocephalicos elongatis. Folia petiolata petiolis usque ad

5 cm. longis saepe latis planisque, petiolo adjecto 4-10 cm. longa,

pinnatim 3-7-partita foliolis membranaceis anguste lineari-spathu-

latis apice obtusis supra medium plerumque 1-2.5 mm. latis interdum

rursus lobatis. Capitula (solitaria, longe pedunculata) radiata, pansa

ad anthesin 2-3 cm. lata et 7-9 mm. alta. Involucri bracteae exteri-

ores 5-8, oblongae, glabrae, obtusae, 4-8 mm. longae; interiores

late ovatae, apice pubescentes, interdum paulo breviores. Flores

ligulati circ. 8, flavi, ligula oblongi vel obovati, apice grosse circ.

3-dentati, fertiles, usque ad 1.5 cm. longi et 1 cm. lati. Paleae glab-

rae, oblongae, plus minusve acutae, 4-6 mm. longae. Flores tubulosi

anulo plus minusve glabri, stylorum ramis termino abrupte crasso-

conicis et vix apiculatis. Achaenia cuneato-orbiculata vel anguste

cuneato-obovata, plana, dorso glabrata ventre plus minusve papil-

losa vel aegre setosa setis interdum clavellatis, duabus faciebus nigra,

margine crasso-alata rugosaque, circ. 3.5-4 mm. longa et (alis

idjectis) 1.4-2.3 mm. lata, apice exaristata sed poculo membranaceo

coronata.

316 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: Collected by Dr. Stillman in valley of the upper

Sacramento River, northern California (Gray).

Distribution: California.

Specimens examined: H. N. Bolander 4520, hillsides, Auburn,

April 10, 1865 (Calif.; Gray; N.Y.; U.S.); Thomas Bridges 260,

California (N.Y.); Mrs. C. C. Bruce 1987, Iron Canyon, Butte

County, May, 1897 (Pom., 2 sheets); J. W. Congdon, Mariposa

Water Ditch, Mariposa County, March 28, 1901 and April 4, 1903

(U.S.); Harvard University Botanical Garden, cult., 1873 and 1882

(Gray); A. Kellogg & W. G. W. Harford 439, dry sandhills, Antioch,

April 16, 1868-1869 (Gray) and April 8, 1869 (N.Y.); Dr. Stillman

(type, Gray; cotype, N.Y.).

Coreopsis Stillmanii var. /3 Jonesii Sherff, Bot. Gaz. 97: 605.

1936.

Herba ± 2 dm. alta, basi multum ramosa. Capitula ad anthesin

3.5-4 cm. lata et circ. 1 cm. alta. Involucrum glabratum; bracteis

exterioribus 8-16, angustissime linearibus, sensim usque ad apicem

obtuse truncatum angustatis, 7-12 mm. longis, demum saepe

patentibus subreflexisve, quam interioribus oblongo-ovatis paulo

brevioribus. Achaenia 4-4.5 mm. longa; corpore ipso nigro, circ.

1 (alis stramineis inclusis circ. 2) mm. lato, glaberrimo vel secundum

costam medianam sparsim papillato-hispidulo et alibi interdum

paucissimis setulis armato, his apice nunc acribus nunc capitatis.

Type specimen: Collected by Marcus E. Jones, No. 3361 pro

parte, Pasadena, California, May 2, 1882 (Pom.).

Distribution: Known only from type locality in California.

Specimens examined: Jones 3361 pro parte (type, Pom.).

14. Coreopsis petrophila A. Gray, Proc. Amer. Acad. 22: 428.

1887.

Fruticosa, 6 dm. forsitan quidem 1 m. alta, ramosissima,

glabrata, caulibus gracilibus obscure tetragonis, saepe rubescentibus.

Folia tenuiter petiolata petiolis 1-2.5 cm. longis, petiolo adjecto 3-6

cm. longa, pinnatipartita, segmentis 5-9 subulato-linearibus, acriter

indurato-apiculatis, inaequalibus, majoribus saepius trifidis vel

1-2-dentatis, ultimis 0.6-5 mm. latis. Capitula graciliter brevius-

culeque pedicellata (pedicellis pubescentibus) et subcymosa, radiata,

pansa ad anthesin 2-2.5 cm. lata et circ. 7-8 mm. alta. Involucri

campanulati subcriniti bracteae exteriores lineari-oblongae, minutae,

tantum circ. 1-2 (-3) mm. longae; interiores oblongae vel oblongo-

REVISION OF THE GENUS COREOPSIS 317

ovatae, 5-6 mm. longae. Flores ligulati 5, flavi, integri, ligula

oblongi, 8-12 mm. longi, tubulo hispido. Paleae scariosae, angusto-

oblongae vel oblanceolatae, dorso longissime villosissimae, apice

trilobatae, quam achaeniorum corpora paulo longiores. Flores

tubulosi circ. 6-10, stylorum ramis apice tenuiter hispido-appendi-

culatis. Achaenia recta, subturgida, obovato-oblonga, nigra, dorso

glabra, marginibus intusque longissime villosissima, corpore 3-4.5

mm. longa et 0.7-0.9 mm. lata, apice biaristata aristis tenuibus

antrorsum barbellatis 3-4.5 mm. longis.

Type specimen: Collected by Edward Palmer, No. 530, hanging

loosely about rocks at the entrance of canyons, Rio Blanco, State of

Jalisco, Mexico, September, 1886 (U.S.).

Distribution: State of Durango and southward into State of

Jalisco, Mexico.

Specimens examined: C. R. Barnes & W. J. G. Land 158, alt.

1,680 meters, Sierra de Esteban, State of Jalisco, September 28, 1908

(Field); Edward Palmer 489< Tejame"n, State of Durango, August

21-27, 1906 (Gray; N.Y.); idem 530 (type, U.S.; cotypes, Gray;

N.Y.); C. G. Pringle 2197, cool ledges of barrancas, near Guadala-

jara, State of Jalisco, November 20, 1888 (Gray) ; idem 2303, rocky

bluffs of Rio Grande de Santiago, near Guadalajara, September 30,

1889 (Field; Gray; N.Y.) idem 11507, alt. 1,500 meters, Sierra de

Esteban, near Guadalajara, September 30, 1903 (Berl.; Field; Gray).

15. Coreopsis petrophiloides Robins. & Greenm. Proc. Amer.

Acad. 29: 388. 1894.

Fruticosa, forsitan 7-9 dm. alta, caulibus plus minusve glabratis,

superne foliosis; ramulis angulatis, paulo pubescentibus. Folia

subsessilia vel petiolata petiolis marginatis usque ad 2 cm. longis,

omnino 4-9 cm. longa, rigida, simplicia, rhomboideo-lanceolata,

acuta, dimidio superiore inciso-serrata, glandulo-punctata, supra

paulo pubescentia, infra glabra ac pallidissima. Capitula in inflo-

rescentias terminales corymbosas congregata, tenuiter pedicellata

pedicellis bracteatis plerumque 2-6 cm. longis, radiata, pansa ad

anthesin 1.5-2.5 cm. lata et 8-10 mm. alta. Involucri bracteae

adpresse hispidae, exteriores circ. 6-8, lineari-oblongae, apice

rotundatae ac cartilagineae, tergo 3-nervatae, circ. 2-3 mm. longae;

interiores ovali-oblongae, 5-7 mm. longae. Flores ligulati ± 8,

flavi, ligula oblongi, apice denticulati, ± 1 cm. longi. Paleae angus-

tissime rhomboideo-lanceolatae, tergo valde erecto-villosae ventre

glaberrimae, apicem acrem versus lateraliter acriterque 3-lobatae

318 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

lobo terminal! multo longiore, ± 6-7 mm. longae. Flores tubulosi

5-dentati, stylorum ramis abrupte breviterque angusto-appendi-

culatis. Achaenia oblongo-linearia, plana, brunneo-nigra, nitida,

longitudinaliter obscuro-sulcata, tergo glabrata, marginibus et

ventris nervo mediano perspicue erecto-villosa pilis brunneo-albidis,

corpora circ. 5-6.5 mm. longa et pilis exclusis 0.5-1 mm. lata, apice

biaristata aristis tenuissimis stramineis antrorsum hispidis circ.

4-5.5 mm. longis.

Type specimen: Collected by Cyrus Guernsey Pringle, No. 5508,

at altitude of 2,400 meters, Nevado de Colima, State of Jalisco,

Mexico, May 22, 1893 (Gray).

Distribution: Nevado de Colima of southernmost Jalisco, south-

eastwardly through Michoacan into State of Guerrero, Mexico.

Specimens examined: Brother G. Arsene, alt. 2,100 meters, Cam-

panario, vicinity of Morelia, State of Michoacan, December, 1910

(U.S.); idem 3185, alt. 2,100 meters, Cerros San Miguel, vicinity of

Morelia, November 18, 1909 (Gray; Kew; Mo.; N.Y.; Stockh.;

U.S., 2 sheets); idem 5295, eodem loco, December, 1910 (Gray;

Mo.; U.S.); idem 6026, alt. 2,200 meters, Cerro Azul, vicinity of

Morelia, 1910 (Gray; Mo.; N.Y.; U.S.);«tem 6060, alt. 2,200 meters,

Cerros San Miguel, vicinity of Morelia, November 15, 1911 (Gray;

Kew; Mo.; U.S.); E. W. Nelson 2185, alt. 2,100-2,550 meters, north-

east slope of Sierra Madre, near Chilpancingo, State of Guerrero,

December 24, 1894 (U.S.); idem 2249, alt. 2,700-3,060 meters, top

of Sierra Madre, eodem loco et tempore (U.S.); Parkinson, Mexico

(Kew; Oxf.); Pringle 5508 (type, Gray).

16. Coreopsis Irmscheriana Bruns, Mitteil. Inst. Allgem. Bot.

Hamburg 8: 81. f. 11, nos. 6-9. 1929.

Herba (ex auctore perennis) 2-3 dm. alta, e radice primaria

parce fibrosa 5-15 cm. longa; caule erecto, satis dense folioso,

rubescenti, tereti, pilis strictis brevibus aspero, laxe ramoso; ramis

decumbentibus vel adscendentibus, teretibus vel quadrangularibus,

vix sulcatis, rubescentibus, hirtis, 1-2 dm. longis. Folia caulina

opposita, brevissime petiolata vel sessilia, anguste ovato-lanceolata,

irregulariter crenato-serrata, usque ad 4 cm. longa et 1.5 cm. lata;

ramulorum alternantia, saepius sessilia, utrinque pilis rigidis sparse

adpresseque hirsuta, plerumque sub 2.3 cm. longa et 0.7 cm. lata.

Capitula solitaria ad apices ramorum superne subnudorum dense

hirsutorum, radiata, pansa ad anthesin 3-3.5 cm. lata et saltern

1 cm. alta. Involucri bracteae ± 16 irregulariter biseriales, lineari-

REVISION OF THE GENUS COREOPSIS 319

oblongae, dense hirsutae, usque ad 8 mm. longae et 2 mm. latae.

Flores ligulati ± 12, aurei, ligula oblongi, circ. 1.4-1.6 cm. longi et

db 4 mm. lati. Paleae margine serratae, apice longe acutae, circ.

6 mm. longae. Flores tubulosi lutei, ± 7 mm. longi; tubo inferne

piloso; limbi lobis intus pilosis. Achaenia obcompressa, nigra, pilosa,

biaristata aristis ciliato-pilosis.

Type specimen : Collected by Ernesto Guenther and Otto Buchtien,

No. 55, at altitude of 30 meters, in the Ravine ("Schlucht") of

Chule, Mejia, Lomas, Peru, November 12, 1923 (Hamb., 2 sheets).

Distribution: Known only from type locality in southern Peru.

Specimens examined: Guenther & Buchtien 55 (2 type sheets,

Hamb.; cotype, Mun., from its photograph in Field).

17. Coreopsis rhyacophila Greenm. Proc. Amer. Acad. 35:

313. 1900.

Suffruticosa, erecta, glabra, 3-10 dm. alta, caulibus teretibus,

griseo-brunneis vel brunneo-rubescentibus, foliosis. Folia opposita

petiolata petiolis tenuibus usque ad circ. 6 cm. longis, petiolo adjecto

usque ad 11 cm. longa, plerumque 2-3-pinnatisecta, utrinque glabra,

glanduloso-punctata, segmentis linearibus vel anguste oblongo-

lanceolatis, apice acutis vel acerrimis, membranaceis, saepius 2-5

mm. latis. Capitula corymbosa, non numerosa, tenuiter pedicellata

pedicellis glabris nudis vel 1-3-bracteatis saepius 5-7 cm. longis,

radiata, pansa ad anthesin 2.5-3 (rarius -4.5) cm. lata et circ. 8-11

mm. alta. Involucri bracteae exteriores circ. 8, patentes vel reflexae,

lineares, apice subacutae, basi saepe sparsim hispidae, 4-6 mm.

longae; interiores late oblongo-ellipticae, apice pubescentes, 7-9 mm.

longae. Flores ligulati 7-10, flavi, ligula elliptico-oblongi, apice

plerumque integri, 1.4-2.3 cm. longi. Paleae lineari-oblongae, cum

fructu deciduae, ad hujus faciem exteriorem applicatae, apice acres

et saepe 1-2-lobatae, dorso ac margine erecto-villosissimae, ventre

glabrae, 5-6 mm. longae. Florum tubulosorum styli ramosi, ramis

conico-appendiculatis. Achaenia obcompressa, cuneate lineari-

oblanceolata, nigra, dorso glabra, margine ac ventris costa mediana

erecto-villosissima, corpore circ. 4 mm. longa et circ. 0.8 mm. lata,

apice tenuiter biaristata aristis albidis antrorsum hispidis circ. 3-4

mm. longis.

Type specimen: Collected by Cyrus Guernsey Pringle, No. 7866,

at altitude of 2,500 meters, in lava fields near Cuernavaca, State of

Morelos, Mexico, February 9, 1899 (Gray).

320 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Distribution: States of Michoacan and Morelos, southern

Mexico.

Specimens examined : Brother G. Arsene, Cerro Azul, District of

Morelia, State of Michoacan, October, 1909 (Field); Pringle 7866

(type, Gray); idem 8332, alt. 2,400 meters, lava fields above Cuer-

navaca, State of Morelos, May 5, 1900 (Berl.; Field; Gray; N.Y.;

Pom.); idem 8431, alt. 2,550 meters, eodem loco, October 13, 1900

(Field; Gray; N.Y.; Pom.); idem 9886, eodem loco, November 19,

1902 (Berl.; Field; Gray; N.Y.).

18. Coreopsis Pringlei Robins. Proc. Amer. Acad. 43: 41. 1907.

Fruticosa, ramosa, ramis glabris teretibus cortice ochraceo-griseo

obtectis, ramulis striatis viridibus plus minusve 6-angulatis foliosis.

Folia opposita petiolata petiolis ± 1 cm. longis ac saepe marginatis,

petiolo adjecto 2-4 cm. longa et 1-3 cm. lata, bipinnatifida, pallide

viridia, glaberrima vel vix pilosiuscula, segmentis patentibus angus-

tissime linearibus leviter acutatis integris vel cum lobis secundariis

paucis instructis 4-18 mm. longis et 0.6-0.8 mm. latis. Capitula

ramulos terminantia solitaria vel corymbose 3-5-aggregata, tenuiter

pedunculata pedunculis 1-5 cm. longis nudis vel in mediana parte

cum bractea unica lineari instructis, erecta vel nutantia, radiata,

pansa ad anthesin 2-3 cm. lata et 7-9 mm. alta. Involucri campa-

nulati bracteae exteriores circ. 8, lineari-oblongae, apice rotundatae

basi pilosiusculae, 3-5 mm. longae et 1 mm. latae; interiores oblongo-

ovatae, circ. 6-8 mm. longae. Flores ligulati circ. 8, juventate supra

aurei subtus flavidi maturitate laete flavi, ligula oblongi, apice

integri, circ. 1.2 cm. longi et 4-6 mm. lati. Paleae lineari-oblongae,

cum fructu deciduae, ad hujus faciem exteriorem applicatae, apice

eroso-ciliatae, circ. 5 mm. longae. Stylorum rami pro floribus

tubulosis anguste conico-appendiculati. Achaenia vix matura

obcompressa, cuneate lineari-oblanceolata, dorso obscure circ.

4-costata et glabra vel sparsim erecto-villosa, facie ventrali et

marginibus erecte villosissima, corpore nigro circ. 5 mm. longa, apice

tenuiter biaristata aristis albidis antrorsum hispidis circ. 3-4 mm.

longis.

Type specimen: Collected by Cyrus Guernsey Pringle, No. 10050,

at altitude of 2,040 meters, on dry ledges, San Juan del Rio, State

of Quere*taro, Mexico, September 8, 1905 (Gray).

Distribution: Known only from type locality in State of Quere*-

taro, Mexico.

Specimens examined: Pringle 10050 (type, Gray; cotypes, Berl.;

Field; N.Y.; U.S.).

REVISION OF THE GENUS COREOPSIS 321

19. Coreopsis rudis (Benth.) Benth. & Hook, ex Hemsl. Biol.

Centr. Amer. Bot. 2: 196. 1881. Epilepis rudis Benth. PL Hartweg.

17. 1839.

Fruticosa, erecta, hispida, foliosa, forsitan 7-9 dm. alta. Folia

opposita petiolata petiolis anguste alato-marginatis circ. 1-2 cm.

longis, petiolo adjecto circ. 4-7 cm. longa, vix rigidula, pinnatisecta,

segmentis cuneatis lanceolatisve mine integris mine grosse dentatis

pinnatifidisve, saepius 3-10 mm. latis. Capitula corymboso-pani-

culata pedicellis tenuibus plerumque 0.5-1.5 cm. longis, radiata,

pansa ad anthesin circ. 2.5 cm. lata et circ. 8-10 mm. alta. Involucri

campanulati bracteae exteriores circ. 6, oblongae, obtusae, circ. 3-4

mm. longae, interioribus late oblongo-obovatis dimidio breviores.

Flores ligulati circ. 5, flavi, ligula ovali-oblongi, neutri, apice sub-

integri vel plus minusve denticulati. Paleae cum fructu deciduae,

ad hujus faciem exteriorem applicatae, dorso valde longeque erecto-

villosae, ventre glaberrimae, apice trifidae, circ. 5-6 mm. longae.

Flores tubulosi 5-dentati, stylorum ramis cono superatis. Achaenia

obcompressa, anguste lineari-oblanceolata, dorso circ. 4-costato

glabra, marginibus villosissima, ventre praeter lineam villosam

glabra, corpore subnigro 4-5 mm. longa et circ. 0.7-0.8 mm. lata,

apice tenuiter biaristata aristis stramineis antrorsum hispidis circ.

2-3 mm. longis.

Type specimen: Collected by Theodor Hartweg, No. 116, in pine

forests, Bolanos, Canton of Colotlan, State of Jalisco, Mexico, 1836-

1838 (Kew).

Distribution: Known only from type locality in Canton of Colot-

lan, State of Jalisco, Mexico.

Specimens examined: Hartweg 116 (type, Kew; cotypes, Berl.;

Gray).

Bentham described the ray flowers as neuter, but Bentham and

Hooker (Gen. PI. 2: 386, sub Epilepide. 1873) described them as

fertile. In the fruiting heads before me they are completely sterile.

20. Coreopsis glaucodes Blake & Sherff ex Sherff, Bot. Gaz.

80: 369. 1925.

Frutex glaberrimus, 1 m. altus, partibus novellis glaucescenti-

bus, caule tereti subgriseo, ramis ramulisque striatis, parce angulatis,

subbrunneis, glaucescentibus, internodiis 1-6 cm. longis. Folia

opposita tenuiter petiolata petiolis basi in poculum circ. 1 mm. altum

connatis, 4-10 mm. longis, petiolo adjecto circ. 1.5-3.5 cm. longa

et 6-18 mm. lata, circumambitu saepius cuneata vel rhombica

322 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

rarius deltoidea, ad medium vel ultra 3- (vel etiam 4-) lobata et

in petiolum cuneate decurrentia, lobis cuneatis vel oblanceolatis

vel lineari-lanceolatis vel lanceolatis, acutis, integris (interdum

terminal! rarius etiam lateralibus 2-3-dentatis), membranaceis, pal-

lido-viridibus, glaucescentibus, plerumque 6-16 mm. longis et 1.5-

3.5 mm. latis. Capitula parva 1-4 ad apices ramorum, tenuiter

pedunculata pedunculis 1.5-5.5 cm. longis, radiata, pansa ad anthesin

1.2-2 cm. lata et 5-7 mm. alta. Involucri glabri glaucescentisque

bracteae exteriores 8, anguste triangularibus, obtusae, 3-nerviae,

2-2.5 mm. longae et 0.2-0.5 mm. latae; interiores oblongae, obtusae,

apice obscure ciliolatae, multinerviae, flavo-marginatae, glauces-

centes, 5-7 mm. longae et circ. 1.8 mm. latae. Flores ligulati 8,

aurei, ligula oblongi, subintegri, circ. 8-nervii, ± 7.5 mm. longi et

± 3.2 mm. lati. Paleae oblongae, subacutae vel obtusae, circ.

5-nerviae, ad lineam medianam pilosae, apice eroso-spinulosae,

± 4.5 mm. longae. Disci florum stigmata deltoidea, hispidula, api-

culata. Achaenia oblanceolato-oblonga, obcompressa, griseo-nigra,

dorso glabra, marginibus piloso-ciliata, apice ciliata, ventre ad

costam medianam sparsim pilosa, corpore 3.5-4 mm. longa et 1.8

mm. lata, biaristata aristis sursum ciliatis, 1.5-2 mm. longis.

Type specimen: Collected by August Weberbauer, No. 4048, in

the open formation, at altitude of 2,600 meters, below Hacienda La

Tahona, near Hualgayoc, Department of Cajamarca, Peru, May 15,

1904 (Berl., 2 sheets).

Distribution: Known only from type locality.

Specimens examined: Weberbauer 4048 (2 type sheets, Berl.).

21. Coreopsis microlepis Blake & Sherff ex Sherff, Bot. Gaz.

80: 370. 1925.

Frutex + 4 dm. altus, caule tenui sub inflorescentia plerumque

simplici raro ramis paucis erectis simplicibus ramoso, usque ad 1.8

mm. crasso, hexagono, glabro vel in lineis posita sub nodis plus

minusve hispidulo, internodiis 1.3-5 cm. longis, quam foliis plerum-

que longioribus. Folia opposita tenuiter petiolata petiolis superne

submarginatis inferne hirsuto-ciliatis 4-10 mm. longis basi in poculum

connatis, lamina circumambitu deltoidea paene ad costam trilobata,

8-21 mm. longa et 7-22 mm. lata, segmentis membranaceis, glabra,

margine saepe revolutis, acriter apiculatis, lateralibus cuneatis vel

elliptico-obovatis vel spathulatis, integris vel 2-3-dentatis, 5-11

mm. longis et 2-6 mm. latis, terminali 3-4-lobato saepius usque circ.

ad medium lobis integris vel 2-3-dentatis. Capitula mediocria 3-15

REVISION OF THE GENUS COREOPSIS 323

cymam vel paniculam nudam efficientia rarius solitaria, pedicellata

pedicellis pilosis 1.5-5 cm. longis, radiata, pansa ad anthesin circ.

1.5-2 cm. lata et 6-7 mm. alta. Involucri glabri vel basi ipso his-

piduli bracteae exteriores 8-9, ovatae vel oblongo-ovatae, sub-

obtusae, 1.5-2 mm. longae et 0.6-0.8 mm. latae; interiores elliptico-

oblongae, obtusae, facie subnigrae margine angusta flavidae, 6-7 mm.

longae et circ. 2.2 mm. latae. Flores ligulati verisimiliter 8, aurei,

ligula oblongo-ovati, subintegri, circ. 7-nervii nervis principalibus

dorsaliter sparsimque hirsutulis, db 6.7 mm. longis et 3 mm. latis.

Paleae obovatae, obtusae vel acutae, circ. 6-nerviae, ad lineam

medianam pilosae, circ. 4.5 mm. longae. Disci florum stigmata

incrassata, hispidula, abrupte apiculata. Achaenia obovata vel

oblonga, griseo-nigra, longe piloso-ciliata et duabus faciebus pilosa,

corpore 2.8-3.5 mm. longa et circ. 1.2 mm. lata, biaristata aristis

subpaleaceis, sursum piloso-ciliatis, 0.8-1.2 mm. longis.

Type specimen: Collected by Alexander Mathews, No. 1418,

Province of Chachapoyas, Department of Amazonas, Peru, 1835-

1836 (Kew, 2 sheets).

Distribution: Known only from type locality.

Specimens examined: Mathews 1418 (2 type sheets, Kew); A.

Raimondi 1774, road between Chacapoyas and Cheto, Peru, March,

1869 (Berl.).

22. Coreopsis Macbridei Sherff, Bot. Gaz. 89: 369. 1930.

Frutex glabratus, ramosus, caulibus subteretibus. Folia laxe

disposita, opposita, petiolata petiolis tenuibus circ. 1-1.5 cm. longis,

primaria petiolo adjecto etiam 4-4.5 cm. longa, pinnatim 5-7-secta,

segmentis anguste linearibus, glaberrimis, apicaliter acribus, 1-2.5

cm. longis et 1-1.7 mm. latis. Capitula corymboideo-paniculata,

numerosa, tenuissime pedunculata (vel pedicellata) pedunculis

saepius 3-7.5 cm. longis glaberrimis vel apicem versus vix pubes-

centibus, radiata, pansa ad anthesin tantum circ. 2 cm. lata et

6-9 mm. alta. Involucri glabrati sicci subnigri bracteae exteriores

circ. 8, lineari-oblongae, obtusae, tantum circ. 1.5-2 mm. longae;

interiores oblongo-ovatae circ. 5-6 mm. longae. Flores ligulati circ.

8, fusco-flavi, ligula lineari-elliptici, apicaliter integri vel rarius

denticulati, tantum circ. 8-10 mm. longi et 1.5-3 mm. lati. Paleae

lineari-oblongae, circ. 5-striatae, tergo villosissimae ventre glab-

rae cum achaenio deciduae. Disci florum stigmata incrassata et

minutissime caudato-appendiculata. Achaenia lineari-oblanceolata,

valde obcompressa, dorso sub palea glabra, ventre ad costam medi-

324 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

anam villosa, marginibus longe perspicueque villoso-ciliata, apice

biaristata aristis sursum villosis circ. 2 mm. longis.

Type specimen: Collected by J. Francis Macbride, No. 3504, neat

compact half-shrub (or shrub) of southwestern rock outcrops, alt.

about 2,100 meters, Huanuco, Peru, April 26, 1923 (Field).

Distribution: Peru.

Specimens examined: Macbride 3152, ragged shrub, river canyon

slopes, alt. about 2,100 meters, Ambo, April 4, 1923 (Field; U.S.);

idem 3504. (type, Field; cotype, U.S.).

The specimens examined had been determined by Dr. S. F. Blake

as a form of Coreopsis Townsendii Blake, with more leaf lobes than

on the type. A careful comparison with the type of that species,

however, shows many striking differences. C. Macbridei has a more

scraggly habit; its leaves are fewer, their petioles narrower, and their

divisions 5-7 rather than usually 3; the capitula are more numerous

(more than 30 on one sheet), their peduncles and involucres almost

completely glabrous (not conspicuously tomentose), their external

bracts about 1.5-2 mm. (not 4-4.5 mm.) long, their diameter at

flowering about 2 cm., not 2.5-3 cm., the ligules deep yellow (not

light yellow) and only about 8-10 mm. long and 1.5-3 mm. wide,

not 15-18 mm. long and 5-7 mm. wide, etc.

23. Coreopsis Killipii Sherff, Bot. Gaz. 94: 594. 1933.

Fruticosa, glabra, 6-12 dm. alta, ramosa ramis tenuibus angulatis

internodiis quam foliis longioribus. Folia petiolata petiolis moderate

ciliatis circ. 4-13 mm. longis basi in poculum ± 1.5 mm. altum

connatis, petiolo adjecto circ. 1.5-2.5 cm. longa, pinnatim vel sub-

bipinnatim divisa, segmentis ultimis saepius oblongo-linearibus

crassiusculis eciliatis planis vel vix revolutis apice acutis subacutisve.

Capitula pauca, in pedunculis ramos elongates superne nudatos

terminantibus ± 1-1.2 cm. longis disposita, radiata, pansa ad

anthesin ± 2.6-2.8 cm. lata et ± 0.8-1 cm. alta. Involucrum

tantum ad basim ipsam hispidum, bracteis exterioribus circ. 8,

oblongis, eciliatis, apice obtusis vel rotundatis, 3-5 mm. longis et

1-1.7 mm. latis; interioribus ovato-oblongis, siccis plus minusve

atris, apicem versus vix ciliatis, 7-10 mm. longis. Flores ligulati

forsitan 5 vel 6, brunneo-flavi, ligula elliptico-oblongi, apice integri

vel parce denticulati, circ. 1.2 cm. longi. Paleae spathulato-oblance-

olatae, dorso (praecipue ad costam medianam) hispidae ventre

glaberrimae, apicem acutum versus ciliatae, circ. 5-6 mm. longae ad

corpus achaenii strictim applicatae. Flores tubulosi subbrunneo-

flavi, stylorum ramis incrassatis apice subito breviterque conico-

REVISION OF THE GENUS COREOPSIS 325

appendiculatis. Achaenia valde obcompressa, cuneato-oblanceolata,

nigra vel fere purpurascenti-nigra, dorso glaberrima, ventre margini-

busque perspicue erecto-pilosa pilis elongatis sericeis, corpore 4-5

mm. longa, apice biaristata aristis linearibus stramineis antrorsum

hispidis 1.5-2.5 mm. longis.

Type specimen: Collected by Ellsworth P. Killip and Albert C.

Smith, No. 21823, straggling shrub 2-4 feet high, open hillside at

altitude of 3,000-3,200 meters, Tarma, Department of Junin, Peru,

April 20-22, 1929 (Field).

Distribution: Known only from type locality in Peru.

Specimens examined: Killip & Smith 21823 (type, Field).

24. Coreopsis parviceps Blake & Sherff ex Sherff, Bot. Gaz. 80:

368. 1925.

Frutex glaberrimus, caule tereti, striatulato, griseo, ramis ramu-

lisque tenuibus striato-angulatis subbrunneis, internodiis 1.5-4 cm.

longis. Folia opposita petiolata petiolis tenuibus basi in pocula

brevia et vix ciliolata connatis 3-8 mm. longis; laminis inferioribus

circumambitu deltoideis, circ. 2.2 cm. longis et usque ad 2.8 cm. latis,

irregulariter 2-4-partitis, crassiusculis, minutissime punctulatis,

segmentis linearibus vel lineari-ellipticis 9-16 mm. longis et 1.8-3

mm. latis perlucidule pinnato-venosis apice acutis; superioribus

integris vel interdum trilobatis, lineari-ellipticis, 8-19 mm. longis

et 1.5-3 mm. latis. Capitula parva, solitaria, tenuiter pedunculata

pedunculis glabris db 2.7 cm. longis, radiata, pansa ad anthesin

circ. 1.8 cm. lata et circ. 6 mm. alta. Involucri bracteae exteriores

6-8, lineari-oblongae, glabrae, apice obtusae, circ. 3 mm. longae et

0.8 mm. latae, interiores late lanceolato-oblongae, apice obtusae et

eroso-denticulatae, circ. 6 mm. longae et 2.5 mm. latae. Flores

ligulati circ. 8, aurei, ligula late oblongo-ovati, circ. 12-nervii, apice

subintegri ac subemarginati, circ. 9 mm. longi et 4 mm. lati. Paleae

oblongo-obovatae, dorso sparsim pubescentes, circ. 5-nerviae, apice

truncatae vel emarginatae plus minusve spinuloso-ciliatae, circ. 3.5

mm. longae. Disci florum stigmata deltoideo-triangularia, acuminata,

hispidula. Achaenia valde immatura oblonga, piloso-ciliata, ventre

ad costam medianam sparsim pilosa, apice ciliata ac biaristata aristis

sursum piloso-ciliatis, 2 mm. longis.

Type specimen: Collected by Const. De Jelski (distrib. Ign.

Szyszylowicz), No. 765, Tambillo, Peru, August 19, 1878 (Berl., 2

sheets).

Distribution: Known only from type locality.

Specimens examined: De Jelski 765 (2 type sheets, Berl.).

326 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

25. Coreopsis oblanceolata Blake, Contr. U. S. Nat. Herb.

22: 642. 1924.

Frutescens, caule gracili, glabro, internodiis inferioribus mod-

erate brevioribus superioribus multo longioribus quam foliis. Folia

opposita, oblanceolata, Integra, coriacea, glabra, apice acutiuscula

obtusave et obtuse calloso-mucronulata, de loco diametri maximi

sensim in basim petiolo saepe similem angustata, parce revoluta,

manifeste unicostata venis lateralibus obscuris, utrinque viridia,

circ. 2.5-3.8 cm. longa et circ. 5-7.5 mm. lata. Capitula

solitaria vel perpauca, pedunculata pedunculis subtenuibus usque

ad 11 cm. longis, radiata, pansa ad anthesin ± 4 cm. lata et

± 1.2 cm. alta. Involucri bracteae exteriores circ. 8, anguste ob-

longae, basi parce angustatae et sparsissime pubescentes, apice

rotundatae vel obtusae, marginibus breviter hispidae, 3-vittatae,

circ. 5.5 mm. longae et circ. 1.8 mm. latae; interiores oblongo-

obovatae, acutae acuminataeve, apice eroso-ciliatae, circ. 7.5 mm.

longae. Flores ligulati ± 8, flavi, ligula elliptico-oblongi, apice

bidentati, ± 2 cm. longi et ± 6 mm. lati. Paleae elliptico-ovatae,

acuminatae, inferne ciliatae, apicem versus eroso-ciliatae, ± 5 mm.

longae. Flores tubulosi flavi, tubi apice pilosi; styli ramis termino

breviter triangulatis et acutis. Achaenia immatura valde obcom-

pressa, oblongo-elliptica, ventre pubescentia, dense longo-ciliata,

apice setosa, corpore circ. 4 mm. longa, biaristata aristis lanceolato-

linearibus, subaequalibus, dense antrorsumque hispidis, circ. 2.2

mm. longis.

Type specimen: Collected by Charles Henry Tyler Townsend, No.

A211, at altitude of 2,400-2,850 meters, on top of the western

Cordillera opposite Huancabamba, Peru, September 26, 1911 (Field).

Distribution: Known only from type locality in northernmost

Peru.

Specimens examined: Townsend A211 (type, Field).

25a. Coreopsis Woytkowskii sp. nov.

Fruticosa, usque ad 7 dm. alta; ramulis sulculatis albido-tomen-

tosis. Folia opposita subtenuiter petiolata petiolis marginatis con-

duplicatis hispido-ciliatis et faciebus plus minusve setosis usque ad

circ. 1.5 cm. longis, petiolo adjecto usque ad 5.5 cm. longa et 0.8-1.5

cm. lata, oblongo-oblanceolata, supra medium vel apicem versus

pauciserrata (1-3 dentibus pro utroque latere), apice acuta rarius

subtruncata sub medio usque ad petiolum sensim vel curvescente

angustata, submembranacea, utrinque minutissime numerosissime-

REVISION OF THE GENUS COREOPSIS 327

que glandulo-papillata sed non vel secundum venulas hispidula,

suaveolentia. Capitula solitaria ad ramulorum apices disposita,

pedunculata pedunculo nitide albo-lanato circ. 2.5-4 cm. longo,

radiata, pansa ad anthesin 4-4.5 cm. lata et 1.3-1.8 cm. alta. In-

volucrum extus basi perspicuissime alibi moderate sparsimve albo-

lanatum; bracteis exterioribus ±6, oblongis, supra medium dilatatis,

apice rotundis, tergo 3-nervatis et minutissime glandulo-papillatis,

circ. 8 mm. longis et 2-2.6 mm. latis; interioribus oblongo-ovatis

rfcdimidio longioribus. Flores ligulati 8, intense lutei, ligula obovati,

apice obscure 2- vel 3-denticulati, ±2 cm. longi, tubo ovarioque plus

minusve albo-pilosis. Paleae oblongo-lineares, dorso longissime

erecteque sericeo-pilosae, ventre glabrae, apice eroso-emarginatae,

±7 mm. longae. Achaenia matura non visa; valde immatura plana,

cuneate linearia, dorso glabra, ventre marginibusque elongate

erecteque sericeo-pilosa, apice biaristata aristis tenuissimis antror-

sum hispidis corollis subaequalibus.

Type specimen : Collected by Felix Woytkowski, No. 24, on rocky

hills at altitude of 2,625 meters, vicinity of Celendin, Department of

Cajamarca, Peru, June 5, 1936 (Field).

Distribution: Known only from type locality in Peru.

Specimens examined: Woytkowski 24- (type, Field).

The collector's notes state that this species "grows in profusion

in some places." It is very easily distinguishable in its foliage from

all other South American species of Coreopsis. Fortunately, the

type, though collected but very recently, was received in time to be

treated in this work. To Mr. Paul C. Standley, Associate Curator

of the Herbarium of Field Museum of Natural History, I am in-

debted for the privilege of examining it.

26. Coreopsis longula Blake, Contr. U. S. Nat. Herb. 22: 642.

1924.

Frutex glaber, verisimiliter circ. 1 m. altus, caule ramisque

tenuibus et multiangulato-striatis, internodiis quam foliis saepius

brevioribus. Folia opposita interdum cum fasciculis axillaribus,

simplicia, flagellari-linearia, basi in poculum circ. 1 mm. altum

connata, crassiuscula, subrevoluta, sicca minutissime numerosis-

simeque albido-punctulata, eciliata, apice obtusiusculo cartilagineo-

indurata, saepius 4-7.5 cm. longa et 0.6-1.1 mm. lata. Capitula

non pauca, pedunculata pedunculis tenuibus 1-4 cm. longis, radiata,

pansa ad anthesin ± 2.4 cm. lata et circ. 8 mm. alta. Involucri

glabrati bracteae exteriores circ. 8, oblongo-lineares, obtusae,

328 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

2.5-4 mm. longae et 0.4-0.6 mm. latae; interiores oblongo-lanceolatae,

flavo-marginatae, apice obtuso subtruncatae et eroso-denticulatae,

7-8.5 mm. longae et 2-2.5 mm. latae. Flores ligulati circ. 8, flavi,

ligula elliptico-oblanceolati, apice integri vel obscure denticulati,

circ. 1.2 cm. longi et 2.5-3.5 mm. lati. Paleae oblongo-lineares,

obtusiusculae, 5-7-striatae, glabrae vel dorsaliter ad costam medi-

anam hispidae, 7-8.5 mm. longae et 0.6-1 mm. latae. Disci florum

stigmata incrassata, breviter obscureque crasso-apiculata. Achaenia

lineari-oblonga, obcompressa, atra, corpore circ. 7 mm. longa et circ. 1

mm. lata, dorso leviter ventre marginibusque graviter sursum pilosa,

apice sursum pilosa et vestigio floris minutissimo coronata et bia-

ristata aristis sursum hispidis 2.5-3 mm. longis.

Type specimen: Collected by Alexander Mathews, in the Province

of Chachapoyas, Peru, 1835-1836 (Gray).

Distribution: Known only from type locality.

Specimens examined: Matheivs, Province of Chachapoyas, Peru

(type, Gray); idem, eodem loco, sine anno (Kew) et 1835 (Kew) et

1836 (Kew, 2 sheets).

27. Coreopsis venusta HBK. Nov. Gen. et Sp. 4: 180 (229).

1820.

Frutex ramosus, erectus vel procumbens, ramis teretibus glabris

striatis, ramulis tetragonis, striatis; omnibus internodiis plerumque

0.6-3 cm. longis. Folia opposita, indivisa, anguste spathulato-

linearia, apice acutiuscula, basi angustata, integerrima, exsiccata

subcoriacea, subuninervia, eciliata, supra glabra, infra ad costam

medianam hispida, 1.5-4 cm. longa et 0.8-1.5 mm. lata. Capitula

solitaria, tenuiter pedunculata pedunculis glabratis 5-15 cm. longis,

radiata, pansa ad anthesin circ. 2.5 cm. lata et 6-8 mm. alta. Invo-

lucri plus minusve hispidi bracteae exteriores circ. 8, oblongae,

obtusae, circ. 3.5-4 mm. longae et 1-1.5 mm. latae; interiores ovato-

ellipticae, obusae, multinerviae, margine angusto flavidae aliter atro-

brunneae, 5-6 mm. longae. Flores ligulati 6-8, aurantiaci, ligula

oblonga, apice obsolete tridentata vel subintegri, circ. 12 mm. longa

et 5-6 mm. lata. Paleae oblanceolatae, tenuissime membranaceae,

flavi, perspicue atronervatae circ. 6 nervis, supra laciniato-ciliatae,

circ. 6 mm. longae. Disci florum stigmata incrassata, hispida,

apicaliter brevi-caudata. Achaenia lineari-oblonga, obcompressa,

brunneo-atra, tergo sub palea glabra, margine piloso-ciliata, ventre

hispida saepe plus minusve tuberculata, corpore circ. 5 mm. longa

et 1-1.2 mm. lata, apice biaristata aristis sursum piloso-ciliatis

circ. 3 mm. longis.

REVISION OF THE GENUS COREOPSIS 329

Type specimen: Collected by Alexander Humboldt and Aime

Bonpland, near Loja (Loxa), Province of Loja, Ecuador (Par.).

Distribution: Ecuador and Peru.

Specimens examined: A. S. Hitchcock 21564, alt. 2,200-3,100

meters, Paramo between San Lucas and Ona, Province of Loja, Ecua-

dor, September 7, 1923 (Gray; N.Y.; U.S.); Humboldt & Bonpland,

near Loja, Ecuador (cotype, Berl.); iidem, Paramo de Saraguru,

Ecuador (Berl.); A. Raimondi 6463, Callacate, Department of

Cajamarca, Peru, May, 1879 (Berl.); idem 12060, Huambo, Province

of Chota, Department of Cajamarca, Peru, June, 1868 (Berl.).

28. Coreopsis nodosa Sherff, Bot. Gaz. 94: 595. 1933.

Fruticosa, forsitan 6-8 dm. alta, ramosissima ramulis nunc sub-

teretibus nunc valde angulatis glanduloso-puberulis internodiis

numerosissimis plerumque 1-4 rarius usque ad ±10 mm. longis et

circ. 1-1.5 mm. crassis. Folia numerosissima opposita, petiolata

petiolis plus minusve hispido-ciliatis basi in poculum minutum

connatis 1-5 mm. longis (parte inferiore ad ramulos juncta manente

post casum foliorum), petiolo adjecto tantum circ. 7-9.5 mm. longa,

glabra, pinnatim 3-7-partita segmentis crassiusculis oblongis vel

ovato-oblongis sparsim hispidulis apice subobtuso-mucronatis

plerumque tantum 2-3.5 mm. longis et 0.4-0.7 mm. latis. Capitula

tenuiter pedunculata pedunculis (supra summa folia) circ. 1 cm.

longis, radiata, pansa ad anthesin ±1.8 cm. lata et 7-8 mm. alta.

Involucri basaliter tomentuloso-hispidi bracteae exteriores circ. 8,

cuneato-oblongae, longitudinaliter 3-vittatae, glabratae, apice ob-

tusissimae vel rotundatae, 2-3 mm. longae; interiores cuneato-

oblongae apice obtusissimae 4-7 mm. longae. Flores ligulati 6-8,

claro-flavi, ligula elliptico-oblongi, apice integri, ±8 mm. longi.

Paleae lineari-oblongae, dorso marginibusque erecto-hispidae ventre

glabratae, apice acutae, 5-6 mm. longae, ad corpus achaenii strictim

applicatae. Achaenia valde obcompressa, cuneate lineari-oblonga,

atra, duabus faciebus praesertim ventrali ac marginibus valde

erecto-setosa setis longis albidis sericeis, corpore circ. 4 mm. longa,

apice biaristata aristis tenuibus stramineis antrorsum setosis 1-1.8

mm. longis, inter aristas poculo minuto coronata.

Type specimen: Collected by Francis W. Pennell, No. 13646,

open, rocky slope, at altitude of 2,900-3,100 meters, Ollantaitambo,

Department of Cuzco (Cusco), Peru, April 26, 1925 (Field).

Distribution: Known only from type locality in Peru.

Specimen examined: Pennell 13646 (type, Field).

330 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

29. Coreopsis senaria Blake & Sherff ex Sherff, Bot. Gaz. 80:

367. 1925.

Frutex ramulis hexagonis, lineatim (2 lineis) hispidulis, infra

foliorum basibus persistentibus vestitis, internodiis usque ad 10 mm.

longis. Folia opposita cum fasciculis axillaribus, brevissime petiolata

petiolis tantum circ. 1-1.5 mm. longis glabris omnino connatis;

laminis 3-partitis, segmentis anguste lineari-oblanceolatis, apice

acutis callosisque, margine integris, carnosis, atro-viridibus, glabris,

1-nerviis, circ. 6-8 mm. longis et 0.6-0.9 mm. latis. Capitula solitaria

in pedunculis brevissimis (usque ad 1 cm. longis) hirtellis insidentia,

radiata, pansa ad anthesin circ. 2 cm. lata et 7 mm. alta. Involucri

bracteae exteriores manifeste biseriales, omnino circ. 18, subaequales,

herbaceae, lanceolatae vel lineari-lanceolatae, obtusae vel subacutae,

saepe apiculatae, 3-nerviae, infra medium ciliatae, dorso glabrae,

4-5 mm. longae et 1 mm. latae; interiores 8-9, membranaceae,

oblongae, obtusae, margine angusto flavidae aliter brunneae, cilio-

latae sed faciebus glabrae, circ. 2.5 mm. latae. Flores ligulati circ.

8, aurei, ligula oblongo-ovati, circ. 11-nervati, circ. 6 mm. longi et

3 mm. lati. Paleae lanceolatae, acuminatae, ciliatae, ad lineam

medianam pilosae, 5-nerviae, circ. 4 mm. longae. Achaenia valde

immatura obovato-oblonga, margine apiceque piloso-ciliata, facie

exteriore glabra interiore aegre pilosa, biaristata aristis sursum

piloso-ciliatis, circ. 2.2 mm. longis.

Type specimen: Collected by A. Stubel, No. 35 pro parte, on

punas, on an excursion from Pacasmayo to Moyobamba, Peru,

April to June, 1875 (Berl.).

Distribution: Peru.

Specimens examined: A. Raimondi 436, heights between Balsas

and Celendin, departments of Amazonas and Cajamarca, Peru, April,

1869 (Berl.); Stubel 35 pro parte (type, Berl.); Felix Woytkowski 23,

growing up to 6 dm. tall, common, on rocky hills, alt. 2625 m., vicinity

of Celendin, Department of Cajamarca, Peru, June 5, 1936 (Field).

The leafy branchlets have a juniperous aspect. The numerous

outer bracts of the involucre also afford a ready means of determi-

nation.

30. Coreopsis foliosa A. Gray, Proc. Amer. Acad. 5: 125. 1861.

Frutex ramosus, caule ramisque subquadrilateralibus striatis

hispidis fere occultis. Folia numerosissima opposita glabra petiolata

petiolis planis circ. 3-9 mm. longis, petiolo adjecto 1.2-2 cm. longa,

lamina nunc digitato-ternata nunc pinnatim 4-7-partita, segmentis

REVISION OF THE GENUS COREOPSIS 331

oblanceolatis, crassiusculis, apice obtusiusculis, basim versus angus-

tatis, faciebus minutissime densissimeque albido-punctulatis, mar-

gine saepe revolutis, 4-8 mm. longis et 1-1.6 mm. latis. Capitula

terminalia, breviter pedunculata pedunculis hispidis tenuibus 0.5-2

cm. longis, radiata, pansa ad anthesin circ. 2.8-3.4 cm. lata et 1.2

cm. alta. Involucri hispidi bracteae exteriores circ. 8, oblongae,

apice rotundatae, circ. 5 mm. longae et 1.5 mm. latae; interiores

lanceolatae apicaliter obtusae et interdum suberosae circ. 8 mm.

longae. Flores ligulati circ. 8, ligula anguste obovati, apice plus

minusve denticulati, 1.5-1.9 cm. longi. Paleae late lineari-oblongae,

apice obtuso minute erosae, circ. 6 mm. longae et 1 mm. latae.

Disci florum stigmata rotundo-incrassata, breviter apiculata.

Achaenia late oblanceolata, nigra, piloso-ciliata, una facie perspicue

altera leviter pilosa, corpore circ. 5 mm. longa et 1.8 mm. lata, apice

biaristata aristis sursum hispidis circ. 3 mm. longis.

Type specimen: Collected by Alexander Mathews, No. 1376,

Taulia, Peru, in 1835 (Kew).

Distribution: Known only from type locality.

Specimens examined: Mathews 1376 (type and cotype, Kew).

31. Coreopsis polyactis Blake & Sherff ex Sherff, Bot. Gaz.

80:372. 1925.

Frutex densissime foliosus, 1 m. altus, ramis subteretibus ac

sordido-hirsutulis, internodiis 4-14 mm. longis. Folia opposita

petiolata petiolis plus minusve ciliatis ad basim in poculum hirsutu-

lum 1 mm. altum connatis, petiolo adjecto 1-1.6 cm. longa, in speci-

minibus siccis plerumque subflavescentia, tripartita, segmentis

integris vel terminali tripartite, linearibus vel anguste lineari-

oblanceolatis, 3-6 mm. longis et 0.5-1 mm. latis, acutis, crassis,

sparsim hirsutulis praecipue ad margines; axillis plerumque fasci-

culis obsitis. Capitula saepius solitaria, brevissime pedunculata

pedunculis crassiusculis dense flavescenteque subtomentoso-pilosis

1-1.8 cm. longis demum fistulosis, radiata, pansa ad anthesin 3-3.5

cm. lata et 8-10 mm. alta. Involucri bracteae dense flavescenteque

subtomentoso-pilosulae exteriores circ. 11, manifeste biseriales,

foliaceae, ovatae, obtusae, multinerviae, subaequales, 5-7 mm.

longae et 3.3-4.8 mm. latae; interiores circ. 13, crasso-membranaceae,

elliptico-oblongae, supra usque ad apicem subobtusum vel subacutum

angustatae, anguste pallido-marginatae, multinerviae, quam ex-

teriores dimidio longiores. Flores ligulati circ. 12, aurei, ligula

obovati, subintegri, circ. 10-nervii, circ. 1.4-1.6 cm. longi et 6.5 mm.

332 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

lati. Paleae lineari-oblongae, acutae, circ. 7-nerviae, supra sparsim

ciliolatae, ad lineam medianam pilosae, circ. 6 mm. longae. Disci

florum stigmata deltoidea, hispidula, apiculata. Achaenia immatura

lineari-oblonga, obcompressa, piloso-ciliata, dorso glabra, ventre

pilosa, apice eciliata sed biaristata aristis sursum ciliatis, 3-4 mm.

longis.

Type specimen: Collected by August Weberbauer, No. 6995, on

grass steppes with scattered shrubs, at altitude of 3,500-3,600 meters,

between Hacienda Llague*n and Succhabamba, Province of Otuzco,

Department of Libertad, Peru, June 28, 1914 (Berl., 2 sheets).

Distribution: Known only from type locality.

Specimens examined: Weberbauer 6995 (2 type sheets, Berl.).

Distinct from all other South American species of Coreopsis in

its broadly oval-ovate, biseriate outer involucral bracts.

32. Coreopsis notha Blake & Sherff ex Sherff, Bot. Gaz. 80:

373. 1925.

Folia plerumque 3-4 cm. longa C. notha sensu stricto.

Folia circ. 1.5 cm. longa var. /3 parvulifolia.

Frutex 1 m. altus, caule striatulo subtereti glabrescente,

ramis striato-angulatis sparsim hispidulis, internodiis plerumque

1-4.5 cm. longis. Folia opposita, tenuiter petiolata petiolis sparsim

ciliatis 1-1.5 cm. longis ad basim in poculum plus minusve hispidulum

1.5 mm. altum connatis, petiolo adjecto plerumque 3-4 cm. longa,

ternatisecta lamina circumambitu deltoidea, foliolis lateralibus

breviter stipitatis vel subsessilibus 3-5-sectis terminali longiusculo

stipitato pinnatim 5-secto, segmentis ultimis lanceolatis vel oblongis,

acutis, membranaceis, obscure pubescentibus et ciliatis, plerumque

3-8 mm. longis et 1-2.5 mm. latis. Capitula longe tenuiterque

pedunculata pedunculis monocephalicis sordide crispo-pilosulis

praecipue apicem versus 7-21 cm. longis, radiata, pansa ad anthesin

3.4-3.8 cm. lata et circ. 7-8 mm. alta. Involucri bracteae exteriores

6, oblongae, apice obtusae vel rotundatae, basi plus minusve ciliatae,

dorso non nisi ad basim pilosulae ventre sordido-pilosulae, 2-3 mm.

longae et 1-1.6 mm. latae; interiores 8, oblongo-ellipticae, obtusae,

anguste flavescenti-marginatae, apice minute ciliolatae aliter glabrae,

9-12 mm. longae et 3-4.6 mm. latae. Flores ligulati, aurantiaci,

ligula oblongo-obovati, apice subintegri, 13-15-striati, 1.7-2.3 cm.

longi et circ. 9 mm. lati. Paleae oblongae, apice truncate hispidulo-

ciliatae, ad dorsi lineam medianam pilosae, circ. 12-nerviae. Disci

florum stigmata breviter deltoidea, hispidula, apiculata. Achaenia

REVISION OF THE GENUS COREOPSIS 333

lineari-oblanceolata, obcompressa, brunneo-nigra, marginibus piloso-

ciliata, apice ciliolata, ad ventris costam medianam pilosa, corpore

circ. 6 mm. longa et 1.7 mm. lata, bi- (raro tri-) aristata aristis

sursum piloso-ciliatis, 2.7-3.4 mm. longis.

Type specimen: Collected by August Weberbauer, No. 3812, at

altitude of 2,400-2,700 meters, above San Pablo, Province of Caja-

marca, Department of Cajamarca, Peru, April 26, 1904 (Berl., 2

sheets).

Distribution: Known only from Department of Cajamarca, Peru.

Specimens examined: A. Raimondi 7732 and 7914, alt. 1,920

meters, Cerro de Catache, Cascas, Province of Contumaza, Depart-

ment of Cajamarca, Peru, May 21, 1875 (Berl.); Weberbauer 3812

(2 type sheets, Berl.).

Leaves similar to those of the Mexican C. rhyacophila Greenm.,

but smaller and somewhat less dissected. Vernacular name is given

as "pul," and the heads are said to be used for dyeing.

Coreopsis notha var. ft parvulifolia Sherff, Amer. Journ. Bot.

22: 707. 1935.

A specie foliis tantum circ. 1.5 cm. longis differt.

Type specimen: Collected by A. Raimondi, No. 6695, Con-

tumaza, Province of Contumaza, Department of Cajamarca, Peru,

1875 (Berl.).

Distribution: Known only from type locality in northern Peru.

Specimen examined: Raimondi 6695 (type, Berl.).

Sharply separable from C. notha by the smaller leaves and perhaps

smaller stature of the plant. The more or less filiform leaf petioles

offer a ready distinction from the otherwise deceivingly similar C.

fasciculata var. laevigata of southern Peru.

33. Coreopsis capillacea HBK. Nov. Gen. et Sp. 4: 180

(230). 1820.

Fruticosa, adscendens vel erecta, caule glabro demum ±8 mm.

crassa, forsitan 6-8 dm. alta, ramosa ramis lignosis glabris tetragonis-

que, ramulis glabratis vel ad terminos hispidulis subdense foliaceis

foliis quam internodiis multo longioribus. Folia opposita, petiolata

petiolis usque ad 2 cm. longis, petiolo adjecto, 3-6 cm. longa, pin-

natim 3-5-partita segmentis filiformibus 0.5-1 mm. latis, glabratis

vel rarius minute hispidulis subplanis integerrimis apice acerrimis

lateralibus quam terminali paulo brevioribus. Capitula solitaria

vel perpauca, tenuiter pedunculata pedunculis pubescentibus circ.

334 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

5-10 cm. longis, radiata, pansa ad anthesin circ. 2.5-3.8 cm. lata

et 8-11 mm. alta. Involucri bracteae exteriores circ. 8, lanceolato-

lineares vel oblongo-lineares, facie dorsali longitudinaliter valde

3-nervatae, apice subobtusae, marginibus et facie ventrali albido-

hispidae; interiores late oblongo-lanceolatae, longiores, glabratae,

acutiusculae. Flores ligulati circ. 8, aurantiaci, ligula oblongo-

elliptici vel oblongo-oblanceolati, apice denticulati, ±1.5 cm. longi.

Paleae lanceolatae vel sursum abrupte ad apicem et deorsum sensim

ad basim angustatae, apice saepe eroso-dentatae, dorso hispidae

ventre glabrae, ±5 mm. longae. Florum tubulosorum styli rami

apice breviter acriterque conici. Achaenia atra, oblonge lineari-

lanceolata, valde obcompressa, longitudinaliter multistriata, margini-

bus et ventre (saepe etiam ad tergi costam medianam) dense erecto-

hispida pilis albidis elongatis sericeis et basi papillatis, corpore circ.

4.5-5.5 mm. longa et circ. 1.2-1.5 mm. lata, apice plus minusve

setoso biaristata aristis linearibus antrorsum hispidis circ. 2 mm.

longis.

Type specimen: Collected by Alexander Humboldt and Aime

Bonpland, locality uncertain but suspected by Kunth as being in

the Peruvian Andes. Type should be among the Humboldt and

Bonpland specimens at Paris.

Distribution: Southernmost Ecuador and perhaps adjacent Peru.

Specimens examined: Ed. Andre 1^387, cold region, Catamayo,

Province of Loja, Ecuador, October 31, 1876 (Kew); William Jame-

son, hills near Loja, Ecuador, commun. anno 1865 (Kew) ; J. N. Rose,

A. Pachano, & George Rose 23146, vicinity of Zaragura, Ecuador,

September 28, 1918 (Gray; N.Y.; U.S.);Berthold Seemann 675, Loxa

(Loja), Ecuador, August, 1847 (Kew).

34. Coreopsis triloba Blake, Contr. U. S. Nat. Herb. 22:

643. 1924.

Fruticosa, erecta, caule ramisque gracilibus dense foliatis terminos

versus primo piloso-tomentosis mox glabratis, internodiis plerumque

2-8 mm. longis interdum longioribus (quidem 2 cm.). Folia opposita,

petiolata petiolis 1-2 cm. longis basi in poculum 0.5-1 mm. altum

connatis, petiolo adjecto 2-4 cm. longa, glabra, raro integra lineari-

filiformiaque plerumque tripartita segmentis coriaceis lineari-fili-

formibus apice subulatis integris vel mediano interdum 3-partito,

ultimis 0.3-0.8 mm. latis. Capitula pauca (plerumque 1-3 ad

unicum ramulum), tenuiter pedunculata pedunculis primo piloso-

tomentosis demum glabratis, 1-6 cm. longis, radiata, pansa ad

REVISION OF THE GENUS COREOPSIS 335

anthesin circ. 1.5 cm. lata et 6-8 mm. alta. Involucri bracteae circ. 8,

lanceolatae vel lanceolato-ovatae, obtusiusculae, crasso-herbaceae,

adpressae vel parce laxae, longitudinaliter 3-vittatae, glabrae, circ.

2.5-3 mm. longae et 0.8-1.2 mm. latae; interiores oblongo-ellipticae,

obtusae, glabrae, circ. 5-6 mm. longae. Flores ligulati forsitan 8,

flavi, ligula elliptico-oblongi, apice denticulati, ±6 mm. longi.

Paleae immaturae acutiusculae, apice spinuloso-ciliatae. Achaenia

(valde immatura) obcompressa, ciliolata, biaristata aristis paleolis

similibus antrorsum hispido-ciliatis 0.8 mm. longis.

Type specimen: Collected by William Jameson in Ecuador (U.S.,

2 sheets).

Distribution: Ecuador and Peru.

Specimens examined: Const. De Jelski 705, Cutervo, Peru, May,

1879 (Berl., 2 sheets); Jameson, Ecuador (2 type sheets, U.S.);

E. P. Killip & A. C. Smith 22171, open rocky hillside, alt. 3150

meters, Mantaro Canyon, south of Huancayo, Department of Junin,

April 29, 1929 (Field; forma foliis plus minusve atypicis) ; A. Raimondi

3022, Cutervo, Province of Cutervo, Department of Cajamarca, Peru,

May, 1879 (Berl.).

35. Coreopsis spectabilis A. Gray, Proc. Amer. Acad. 5: 125. 1861.

Suffrutex saltern 3-4.5 dm. altus, caulibus glaberrimis, teretibus,

striatis, internodiis 2-6 cm. longis et circ. 1.5-2 mm. crassis. Folia

opposita, tenuiter petiolata petiolis 0.5-2 cm. longis basaliter in

poculum hyalinum circ. 1-1.3 mm. altum connatis, petiolo adjecto

3-6 cm. longa, 2-3-pinnatisecta, segmentis ultimis planis, glabris,

acriter indurato-apiculatis, plerumque 0.5-0.8 mm. latis. Capitula

perspicue pedunculata pedunculis glaberrimis (inferne 1-bracteatis

vel saepius foliatis 2-3 jugis foliorum) 2.2-2.7 dm. longis, radiata,

pansa ad anthesin ±4 cm. lata et 6-8 mm. alta. Involucri bracteae

glaberrimae, exteriores 6-8, late lineari-oblongae, apice obtusae,

4-5.5 mm. longae et 0.9-1.5 mm. latae; interiores anguste ovatae,

1-1.2 cm. longae. Flores ligulati (forsan 8), flavi, ligula obovati,

apice obscure denticulati, circ. 1.5-1.8 cm. longi et ±1 cm. lati.

Paleae oblanceolatae, apicaliter obtusae, dorsaliter (ad margines

glabrae aliter) sursum sericeo-pilosae, ventraliter glabrae, 6.5-8 mm.

longae, unaquaque cum achaenio decidente. Achaenia valde obcom-

pressa, nigra, oblanceolata, dorsaliter glaberrima, ventraliter (secun-

dum costam medianam) ac marginaliter pilis sericeis albidisque

perspicue surso-pilosa, ad latera subtilia, corpore 5.5-6.5 mm. longa

et 2-2.3 mm. lata, apice biaristata aristis linearibus, stramineis,

sursum sericeo-hispidis, parce patentibus, 2.5-3 mm. longis.

336 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: Collected by McLean, in the Andes Mountains

of Peru (Kew).

Distribution: Peru.

Specimen examined: McLean, Andes Mountains, Peru (type,

Kew).

36. Coreopsis suaveolens Sherff, Bot. Gaz. 89: 369. 1930.

Frutex ramosus, verisimiliter 5-8 dm. altus, ramis subteretibus

plerumque glabratis sed hac iliac glutinosis odore Covilleae divari-

catae (Cav.) Vail suaveolentibus. Folia opposita fasciculata, pri-

maria 1.2-1.6 cm. longa, pinnatim 3-5-secta, plus minusve glutinosa,

segmentis petiolis similibus vel rursus sectis, ultimis linearibus,

glanduloso-hispidis, carnosis, apicaliter mucronatis, tantum 0.3-0.6

mm. latis. Capitula ramos superne nudos (pedunculos tenues)

terminantia, solitaria, radiata, pansa ad anthesin ±2.3 cm. lata et

7-9 mm. alta. Involucri hispidi bracteae exteriores circ. 8, oblongae

vel late lanceolatae, obtusae, 3-5 (-7) mm. longae, quam interiores

oblongo-lanceolatae circ. 8 mm. longae saepius dimidio breviores.

Flores ligulati circ. 6, lutei, ligula oblongo-elliptici, apice integri vel

(etiam profunde) bifidi, 7-11 mm. longi. Disci florum stigmata

brevia, abrupte incrassata truncataque. Achaenia ignota. Habitu

C. fasciculatae Wedd. valde similis.

Type specimen: Collected by Erich Werdermann, No. 1114, at

altitude of about 3,800 meters, Cordillera de Lallinca, Province of

Tarapaca, Department of Tarapaca, Chile, March, 1926 (Gray).

Distribution: Known only from type locality in Chile.

Specimens examined: Werdermann 1114 (type, Gray; cotypes,

Field; Stockh.; Calif.). The label on the Field Museum sheet

says "Cord. Co. Columfusca, Apacheta, Prov. Tarapaca. . . ."

37. Coreopsis Pickeringii A. Gray, Proc. Amer. Acad. 5: 124.

1861. Coreopsis Boliviana Blake, Contr. U. S. Nat. Herb. 22: 644.

1924.

Frutex fere glaberrimus vel praecipue caule ramisque plus minusve

hispidus, 6-10 dm. altus, internodiis plerumque 1-4 cm. longis.

Folia opposita petiolata petiolis tenuibus usque ad 2 cm. longis, basi

in poculum membranaceum saepe 4 mm. altum connatis, petiolo

adjecto 2-3.5 cm. longa, ternatisecta, foliolis 3-5-sectis, segmentis

glabris, crassiusculis, linearibus, eciliatis, acriter apiculatis, usque

ad 1 cm. longis et 0.6-1.5 mm. latis, quam rhachi tenui vix latioribus.

Capitula longe tenuiterque pedunculata pedunculis usque ad 1.5

REVISION OF THE GENUS COREOPSIS 337

(rarius 2.3) dm. longis, radiata, pansa ad anthesin 3.5-4 (-5) cm.

lata et 0.9-1.1 cm. alta. Involucri hispidi bracteae exteriores circ. 8,

late lineari-oblongae, apice obtusae, dorso glabrae hispidaeve,

marginibus longe ciliatae, circ. 5 mm. longae et 1-2 mm. latae;

interferes oblongo-lanceolatae, flavo-marginatae, multistriatae, circ.

1 cm. longae et 3-3.5 mm. latae. Flores ligulati 8 vel 9, flavi, ligula

anguste obovati, apice rotundati et integri vel obscure denticulati,

1.5-2.4 cm. longae et 4-9 mm. lati. Paleae oblongae, apice obtusis-

simae vel truncatae et saepe eroso-dentatae, exteriores dorso villosae,

circ. 7-8 mm. longae. Disci florum stigmata ovoidea, breviter

apiculata. Achaenia oblongo-oblanceolata, plana, atra, dorso sub

palea plus minusve glabra ventre et praesertim marginibus sursum

villosissima, corpore 5.5-6.5 mm. longa et circ. 1.1-1.3 mm. lata,

apice biaristata aristis sursum hispidis circ. 2 mm. longis.

Type specimen : Collected by the United States Exploring Expe-

dition under Captain Wilkes, high in the Andes above Obrajillo,

Department of Lima, Peru (U.S.).

Distribution: Peru and Bolivia.

Specimens examined: Otto Buchtien 97, alt. 3,750 meters, dry

slopes, La Paz, Bolivia, June 18, 1906 (Field) ; J. Francis Macbride

2932, alt. about 2,400 meters, on southeastern rock slide, Matucana,

Peru, March 14-18, 1923 (Field; U.S.); idem & Feather stone 210, alt.

about 2,400 meters, rock outcrop on northern slope, Matucana, Peru,

April 12-May 3, 1922 (Field) ; Francis W. Pennell 14342, forming

large clumps on open, rocky slope, alt. 2,700-2,900 meters, Canta,

Department of Lima, Peru, June 11-19, 1925 (Field) ; H. H. Rusby

1685, alt. 3,000 meters, near La Paz, Bolivia, October, 1885 (type

collection of C. boliviana Blake; Field; Gray; N.Y.); U. S. Explor.

Exped. under Capt. Wilkes, above Obrajillo, Peru (type U.S.).

38. Coreopsis imbricata Sherff, Bot. Gaz. 89: 370. 1930.

Frutex 6-12 dm. altus, caule ramisque glabris, striatis, ramulorum

internodiis numerosis saepius tantum 2-10 mm. longis. Folia

opposita, petiolata petiolis tenuibus basi vix connatis circ. 1 cm.

longis, petiolo adjecto circ. 2-2.5 cm. longa, ternatim bipinnatisecta,

segmentis linearibus, crassiusculis, glabris, eciliatis, acriter apiculatis,

2-8 mm. longis et circ. 0.3-0.6 mm. latis. Capitula ramulos termi-

nantia (vix manifeste pedunculata), radiata, pansa ad anthesin

3.5-4.5 cm. lata et 1-1.2 cm. alta. Involucri villosi bracteae exteri-

ores circ. 16, biseriales, late lineari-oblongae, apice obtusae, 6-7

mm. longae et 1.1-1.5 mm. latae ; interiores ovatae, flavo-marginatae,

338 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

obtusae, 1-1.1 cm. longae. Flores ligulati circ. 8, flavi, ligula oblongo-

elliptici, apice integri vel subintegri, 1.8-2.4 cm. longi et 6-7 mm.

lati. Paleae lineares, circ. 5-striatae, apice obtusae et denticulatae,

±8 mm. longae et ±0.5 mm. latae. Disci florum stigmata caudato-

apiculata. Achaenia lineari-oblanceolata, plana, atra, dorso glabra,

ventre ac marginibus ac apice sursum villosa, corpore ±5 mm.

longa et ±1.4 mm. lata, biaristata aristis sursum hispidis ±2.7

mm. longis.

Type specimen: Collected by R. Pearce, altitude 3,600 meters,

Cordillera Huanta (midway between Lima and Cuzco), Peru,

February, 1867 (Kew).

Distribution: Known only from type locality.

Specimens examined: Pearce, Cordillera Huanta (type, Kew).

Close to Coreopsis Pickeringii Gray, from which it differs in its

numerous, much shorter internodes, its lack of elongate peduncles,

the caudate-tipped stigmas of its disc florets, its double outer in-

volucre the bracts of which are larger, etc.

39. Coreopsis Townsendii Blake, Contr. U. S. Nat. Herb.

22: 643. 1924.

Fruticosa, forsitan 4-7 dm. alta, caule gracili, tetragono, sub-

viridi, glabro, ramis erectis. Folia opposita, petiolata petiolis basi

connatis 1-2.5 cm. longis, petiolo adjecto 2-5 cm. longa, pinnatim

3-5-partita segmentis integris linearibus acutis coriaceis unicostatis

glabris vel inferne ad costam sparsim obscureque pubescentibus

7-23 mm. longis et 1-1.8 mm. latis. Capitula solitaria vel perpauca,

tenuiter pedunculata pedunculis laxe (ad capitulum dense) pilosis

circ. 1-5.5 cm. longis, radiata, pansa ad anthesin 2.5-3.4 cm. lata

et ±7 mm. alta. Involucri bracteae exteriores circ. 6, lineari-

oblongae, obtusae vel rotundatae, 3-vittatae, inferne ciliatae aliter

glabrae, circ. 4-4.5 mm. longae et circ. 1-1.2 mm. latae; interiores

elliptico-oblongae, apice obtuso eroso-ciliatae, ±8 mm. longae et ±2.5

mm. latae. Flores ligulati 8, flavi, ligula anguste oblongo-obovati,

±14 mm. longi et ±7 mm. lati. Paleae oblongae, apice eroso-ciliatae,

dorso infra mediane erecto-pilosae, circ. 3.3-4 mm. longae. Flores

tubulosi flavi, tubi apice glandulo-puberuli, gutture campanulato-

infundibuliformi superne sparsim pilis multiloculatis piloso; stylorum

ramis termino deltoideis et obtusiusculis. Achaenia submatura

oblonga, valde obcompressa, atra, ventris costa mediana plus minusve

erecto-pilosa, tergo glabra, dense longeque erecto-ciliata, corpore

circ. 3.5 mm. longa, apice biaristata aristis (immaturis) lanceolatis,

trigonis, dense antrorsumque hispidis, ±1.3 mm. longis.

REVISION OF THE GENUS COREOPSIS 339

Type specimen: Collected by Charles Henry Tyler Townsend,

No. A192, at altitude of 1,950-2,250 meters, Huascaray, Peru,

September 10, 1911 (Field).

Distribution: Bolivia and Peru.

Specimens examined: K. Fiebrig 3476, alt. 3,200 meters, Jaipa,

Bolivia, February 20, 1904 (Berl., 2 sheets); Townsend A192 (type,

Field).

40. Coreopsis fasciculata Wedd. Chlor. And. 1: 71. 1855.

Coreopsis Matthewsii A. Gray, Proc. Amer. Acad. 5: 124. 1861.

Involucra plus minusve tomentosa C. fasciculata sensu stricto.

Involucra glaberrima vel basi vix sparso-hispida . . .var. 0 laevigata.

Frutex circ. 7.5 cm. altus, caule ramisque glaberrimis vel inter-

dum pubescentibus, angulato-striatis, parce subtetragonis, internodiis

saepius 1-3 cm. (ramulorum saepius 2-7 mm.) longis. Folia opposita

(ilia ramulorum sterilium subfasciculata), petiolata petiolis tenuibus

6-10 mm. longis, inferne saepe ciliatis raro resinosis basi in poculum

1-3 mm. altum connatis, petiolo adjecto 1.3-2 cm. longa, tripartita,

foliolis 2-3-fidis, segmentis linearibus, vix membranaceis, plerumque

glabratis, apice apiculatis, 1-7 mm. longis et 0.4-1.2 mm. latis.

Capitula ramulos (in pedunculos superne tomentosos 1-12 cm. longos

productos) terminantia, radiata, pansa ad anthesin 2.3-2.8 cm.

lata et 8-10 mm. alta. Involucri plus minusve tomentosi bracteae

exteriores 6-8, spathulato-lanceolatae vel lineari-lanceolatae, apice

obtuso saepe rotundatae, 1.3-5 mm. longae et 0.5-2 mm. latae;

interiores lanceolatae, saepius flavo-marginatae, circ. 1 cm. longae.

Flores ligulati circ. 7 vel 8, flavi, ligula elliptico-oblongi vel anguste

obovati, apice subintegri, 1-1.5 cm. longi. Paleae lineari-oblance-

olatae, apice laciniato-denticulatae, dorso villosae, circ. 6 mm.

longae. Disci florum stigmata parce caudato-apiculata. Achaenia

lineari-oblanceolata, plana, dorso sub palea glabra, ventre margini-

busque sursum villosa, corpore circ. 5 mm. longa et circ. 1 mm. lata,

biaristata aristis sursum villosis circ. 2 mm. longis.

Type specimen: Collected by Claude Gay in Peru (Par.).

Distribution: Ecuador (where apparently rare) to Peru and

Bolivia.

Specimens examined: J. Ball, alt. 3,600-3,900 meters, rocky places

near Chicla, Peru, April 21-23, 1882 (Kew); A.Bonpland, equatorial

America (Par., sub nom. Coreopside Matthewsii A. Gray); Otto Buch-

tien 595, alt. 3,300 meters, lower half of Obrajes, La Paz, Bolivia,

May 13, 1919 (Field, 2 sheets; Gray); A.E. Douglas, Arequipa, Peru,

340 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

1892 (Gray); Mr. & Mrs. F. E. Hinkley 51, ravines at about 2,700

meters, southern slopes of Chachani Mountain, north of Arequipa,

Peru, March, 1920 (Gray); Hrdlicka, alt. 2,400-3,000 meters,

western Cordillera, vicinity of San Damian, Peru, January-February,

1913 (U.S.); E. P. Killip & A. C. Smith 21536, straggling shrub,

2-4 ft. high, alt. 3,000-3,500 meters, open hillside, Rio Blanco,

Department of Lima, Peru, April 15-17, 1929 (Field) ; iidem 23304,

up to 3 ft. high, alt. 4,200 meters, open hillside between Huanta and

Hacienda Pargora, Department of Ayacucho, Peru, May 2-19,

1929 (Field) ; Macbride & Featherstone 436, alt. about 2,400 meters,

rocky, grassy summit slope, Matucana, Peru, April 12-May 3, 1922

(Field) ; iidem 690, alt. about 3,600 meters, rocky places, Rio Blanco,

Peru, May 8-19, 1922 (Field); iidem 1237, alt. about 3,000 meters,

rocky southwestern canyon slope, Yanahuanca, Peru, June 16-22,

1922 (Field); Alexander Mathews 571, Purrochuca and Obrajillo,

Peru, April to July (Kew; the basis of C. Matthewsii A. Gray);

F. W. Pennell 13244, alt. 2,800-2,900 meters, rocky ravine above

Arequipa, Peru, April 7-16, 1925 (N.Y.); idem 14396, alt. 2,800-

3,200 meters, open, rocky slopes, along Rio Chillon, above Obrajillo,

Department of Lima, Peru, June 13-23, 1925 (N.Y.) ; idem 14710, alt.

3,300-3,500 meters, Huaros, Department of Lima, Peru, June 23, 1925

(N.Y.) ;C.S. Sargent 27, above Chicla, Peru, December 26, 1905 (U.S.) ;

C. Troll 3157, Paychama, Chile, March 9, 1927 (Berl.); Warszewicz

37, Ecuador (Berl.); August Weberbauer 203, rocky places above

3,000 meters alt., Huillacachi, southwest from Matucana, Peru,

December 28, 1901 (Berl.); idem 2394 pro parte, alt. 3,300-3,500

meters, Tarma, Department of Junin, Peru, February 10, 1903

(Berl.); idem 5764, alt. 3,100 meters, between the Rio de Lomas and

the Rio Yauca (15° 10'-15° 20' S. Lat.), May 15, 1911 (Berl.; Field);

idem 5782, alt. 3,300 meters, south from Chavina, Province of Tarina-

cochas, Department of Ayacucho, Peru, May 16, 1911 (Berl.; Field).

Coreopsis fasciculata var. ft laevigata Sherff, Amer. Journ.

Bot. 22: 707. 1935.

Foliorum segmenta saepius lineari-spathulata vel anguste ob-

longo-oblanceolata, involucris glaberrimis vel basi vix sparso-hispidis.

Type specimen: Collected by Fortunato L. Herrera, No. Ill, at

altitude of 3,400-3,600 meters, Cuzco, Peru (Berl.).

Distribution: Southern Peru.

Specimens examined: Herrera 111 (type, Berl.); idem 391, alt

3,200-3,600 meters, environs of Cuzco, November, 1924 (Berl.;

REVISION OF THE GENUS COREOPSIS 341

planta tinctoria, fide lectoris); idem 1186, alt. 3,500-3,600 meters,

heights of Cuzco, November, 1926 (Berl.); idem 1481 pro parte,

alt. 3,350 meters, environs of Cuzco, March, 1927 (Gray; cum

Bidente andicola var. decomposita 0. Ktze. commixta).

41. Coreopsis lanceolata L. Sp. PI. 908. 1753; Sims in Curtis,

Bot. Mag. pi. 2451 . 1823. Bidens Caroliniana, florum radiis latissimis,

insigniter dentatis, etc. Martyn, Hist. PI. Rar. 26 and plate. 1728.

Bidens succisae folio, radio amplo laciniato Dillen. Hort. Eltham.

PL Rar. Icon, et Nom. 55, pi. 48, /. 56. 1732. Coreopsoides lanceolata

(L.) Moench, Meth. 594. 1794. Coreopsis lanceolata var. glabella

Michx. Fl. Bor. Amer. 2: 137. 1803. Leachia lanceolata (L.) Cass.

Diet. Sci. Nat. 25: 389. 1822. Chrysomelea lanceolata (L.) Tausch,

Hort. Canal./. 1 (ex DC. Prodr. 5: 570. 1836). Coreopsis lanceolata

var. Succisaefolia DC. Prodr. 5: 570. 1836. Coreopsis lanceolata

var. angustifolia Torr. & Gray, Fl. N. Amer. 2: 344. 1843. Bidens

caroliniana Hemsl. in Curtis, Bot. Mag. 27, index. 1906.

Glabra vel foliorum basibus ciliolata C. lanceolata sensu stricto.

Pubescens vel hirsuta var. /3 villosa.

Herba perennis, 2-6 dm. alta, caulibus erectis vel adscendentibus,

teretibus vel subangulatis, glabris, ramosis, basim versus foliosis,

superne subnudis ac maxime elongatis (in pedunculos desinentibus).

Folia opposita, glabra vel basi ciliata, plerumque 5-15 cm. longa,

superiora sessilia inferiora saepius longe tenuiterque petiolata petiolis

laminas interdum superantibus, nunc spathulata nunc linearia vel

lineari-lanceolata nunc lineari-oblanceolata, plerumque simplicia

raro 1-2 lobis parvis lateralibus divisa. Capitula radiata, pansa ad

anthesin 3-6 cm. lata et 1-1.4 cm. alta. Involucri bracteae exteriores

8-10, lanceolatae vel oblongo-ovatae, glabratae vel apice pubescentes,

4-8 mm. longae; interiores lanceolato-ovatae vel oblongo-ovatae,

8-12 cm. longae. Flores ligulati circ. 8, flavi, ligula obovati vel

cuneati, apice 3-lobati lobo mediano plerumque 2-3-dentato, 1.3-

3 cm. longi. Paleae inferne oblongo-lineares superne filiformes,

4-6 mm. longae. Florum tubulosorum stylorum rami caudato-

appendiculati. Achaenia obcompressa, alata, circumambitu (alis

paululum incurvatis inclusis) orbiculata, 2.3-3 mm. longa, corpore

atra, ventre gibbis callosis saepe perspicue munita ac saepe tuber-

culata vel setis nunc superne sensim dilatatis nunc apice capitatis

setosa, apice dentibus parvis fimbriolatis bidentata.

Type specimen: No specimen cited. The Linnean concept rested

upon such clear and unmistakable illustrations as those given by

Martyn (loc. cit.) and by Dillenius (loc. cit.).

342 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Distribution: Michigan and Lake Superior southward to Florida,

Alabama, Louisiana, southeastern Texas, and northern New Mexico;

widely cultivated for ornament and occasionally escaping; apparently

becoming established in eastern China.

Specimens examined: Anon., Florida (Gray; var. angustifolia

Ton\ & Gray, an official specimen for the treatments in Torr. &

Gray, loc. cit. and Gray, Syn. Fl. loc. cit.); H. C. Benke 3183, Miller,

Indiana, May 22, 1922 (Field) ; J. M. Bigelow, Fort Smith, Arkansas

to the Rio Grande, along 35th parallel of latitude, 1853-1854 (N.Y.);

Biltmore Herb. 2067a, dry soil near Apalachicola, Florida, May 2,

1883 (Gray); Dr. Boykin, Georgia (Gray; N.Y.); M. A. Brannon,

near Crown Point, Indiana, 1889 (Field); Brendel, Illinois (Berl.,

2 sheets) ; Mason Bross, Hyde Park, Chicago, Illinois, June 3, 1878

(Field) ; S. H. & D. R. Camp, Bay View, Michigan, July 14, 1896

(Field); H. C. Cheo & W. F. Wilson 11 p.p., near house, Mo Kan

Shan, Province of Chekiang, China, June 19, 1926 (Calif., 2 sheets);

Hubert Lyman Clark, Falmouth Road, Woods Hole, Massachusetts,

August 16, 1899 (Gray); A. H. Curtiss, Pensacola, Florida, 1885

(N.Y.); idem 1481, dry pine woods, near Aspalaga, Florida, April

(var. villosae adpropinquans; Berl., 2 sheets; Field, 2 sheets; Gray);

Arthur W. De Selm 140, west of Kankakee, Illinois, June 1, 1913

(Field); Royal A. Dixon 528, vicinity of Huntsville, Texas, May

6-12, 1910 (Field; var. angustifolia Torr. & Gray); idem & Cornelia

Gage 713, Morgan Park Ridge, Illinois, May 27, 1907 (Field); C. W.

Duesner, Pine, Indiana, 1908 (Field) ; F. S. Earle & C. F. Baker 243,

Auburn, Alabama, May 8, 1897 (Field); H. Eggert, rocky hills,

Jefferson County, Missouri, May 20, 1887 (Gray); idem, eodem

loco, May 25, 1896 (Gray); J. H. Ehlers 518, dry, sandy soil, near

Cecil Bay, Emmet County, Michigan, July 14, 1917 (Gray) ; George

Engelmann, prairies, vicinity of East St. Louis, Illinois, May, 1845

(Berl.);FrcmA; C. Gates 508, Ravenswood, Chicago, Illinois, June 23,

1905 (Field); idem 2478, sandy soil, Waukegan, Illinois, June 8,

1908 (Field) ; idem & M. T. Gates 10547, along sandy road, Cecil Bay,

Emmet County, Michigan, July 14, 1917 (Field); Jesse M. Green-

man 2017, sandy soil near Lake Michigan, Beach, Illinois, June 16,

1907 (Gray); idem 2592, Dune Park, Indiana, June, 1910 (Gray);

Dr. Hale, Louisiana, 1842 (Gray); Elihu Hall, sandy barrens,

Athens, Illinois, 1861 (Gray, cum var. villosa commixta, sed alibi

tantum species ipsa); Roland M. Harper 22, Screeton, Arkansas,

May 1, 1923 (Gray); herb. Hexamer & Maier, rich woods, June 16,

1855 (Gray); E. J. Hill, dry grounds, Morgan Park, Chicago,

REVISION OF THE GENUS COREOPSIS 343

Illinois, June 17, 1876 (Field); Albert S. Hitchcock, Madison County,

Florida, June-July, 1898 (Field); Hart. Berol, September, 1832

(Berl.) ; Mary E. Hutchinson, Randolph, Wisconsin (Field) ; L. N.

Johnson, sandy ridges in woods, Chicago, Illinois, June 19, 1891

(Gray); A. B. Langlois, western Louisiana, April, 1880 (Field); 0. E.

Lansing, Jr. 848, along railroad, Clarke, Indiana, May 31, 1900

(Field); idem 935, eodem loco, June 20, 1900 (Field); idem 2715,

swale near railroad, Indiana Harbor, Indiana, June 17, 1908 (Field) ;

idem 2782, eodem loco et tempore (Gray, var. villosae adpropin-

quans); /. A. Lapham, Milwaukee, Wisconsin (Gray); Le Conte,

North America (Berl.); George W. Letterman, top of sandhill near

Franklin County Road Bridge, Missouri, July, 1883 (Mo.); C. 0.

Levine 778, Honam Island, Kwang-Tung Province, China, May 12,

1917 (Gray; Mo.); L. H. Lighthipe, Monticello, Florida, April 5,

1891 (Field); F. Lindheimer, wet woods west of Houston, Texas,

early May, 1840 (Berl.) ; Ray N. Lloyd, rich or damp soil, Ravens-

wood, Chicago, Illinois, June 3, 1887 (Field); Loring, Lake Superior

(Gray); W. H. Manning, Mackinac Island, Michigan, July 11,

1915 (Gray); F. E. McDonald, dry, gravelly slopes, Peoria, Illinois,

June, 1899 (Field); idem, high, dry hills overlooking Horse Shoe

Bottom, Peoria, June, 1904 (Gray) ; idem, high, gravelly hills, Peoria,

June, 1915 (Field) and June, 1916 (Field) ; S.B. Mead, Mason County,

Illinois, June 18, 1845 (Field; var. angustifolia Torr. & Gray); W. S.

Moffatt 105, sandy thickets, Chicago, Illinois, June, 1895 (Berl.;

Gray); H. F. Munroe, Hyde Park, Chicago, Illinois, May (Field)

and June, 1879 (Field); E. J. Palmer 17, cherty barrens, Newton

County, Missouri, July 1, 1906 (Gray); idem 2386, cherty barrens,

Reding's Mill, Missouri, July 1, 1909 (Gray); idem 2386 A, eodem

loco et tempore (Gray); A. S. Pease 17993, damp, sandy beach,

Bailey's Harbor, Door County, Wisconsin, June 18, 1921 (Gray);

S. F. Poole, Sharon, Massachusetts, July, 1905 (Gray); Thomas C.

Porter, banks of Savannah River, near Augusta, Florida, April,

1847 (Gray) ; P. H. Rolfs 309, Tallahassee, Florida, April 20, 1895

(Field); B. F. Saurman, rich, grassy plains, Apalachicola, Florida

(Field; var. angustifolia Torr. & Gray); F. Scammon, Chicago,

Illinois, June, 1859 (Field) and June 3, 1859 (Field) ; A. K. Schindler

345a, alt. 1,100 meters, Lu-shan, Kuling Mountains, Province of

Kiangsi, China, August-September, 1908 (Berl.); J. H. Schuette,

Door, Wisconsin, July 30, 1887 (Field, 2 sheets) ; idem, shore of Lake

Michigan north of Sturgeon Bay Canal, Wisconsin, July 14, 1890

(Field) -EarlE. Sherff5006, cult, in Harvard Univ. Bot. Gard., August

12, 1929 (Berl.; Brit.; Field); H. C. Skeels 304, Joliet, Illinois, June

344 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

21, 1904 (Field); Paul C. Standley 4410, alt. about 2,460 meters,

hillside near Cowles, Pecos River National Forest, New Mexico,

July 19, 1908 (Gray) ; A. N. Steward, open hillside, Lu-shan, Ruling,

Province of Kiangsi, China, July 27, 1922 (Calif.); idem 1518, in

open, Kikungshan, Province of Honan, China, July 24, 1925 (Calif.) ;

idem 1583, damp meadow, eodem loco, July 27, 1925 (Calif.); W. F.

Thurrow, Hockley, Texas, 1890 (Field) ; S. M. Tracy 9454, St. Andrews,

Florida, April 29, 1908 (Gray) ; idem & Lloyd 508, Virginia, August

15, 1910 (Field; Gray);L. M. Umbach, dry sands, Miller, Indiana,

June 5, 1897 (Field); idem, sands, Clarke, Indiana, June 4, 1898

(Field); H. E. Wheeler 82, near Hazen, Grand Prairie, Arkansas,

May 22, 1924 (Field); H. N. Whitford 64 and 203, waste fields, Cold

Spring Harbor, New York, August, 1903 (Field) ; Emile F. Williams,

self-sown and thriving, Concord, Massachusetts, June 20, 1908

(Gray); Wilbur H. Wright 50, Miller, Indiana, July 2, 1908 (Field).

Coreopsis lanceolata var. /3 villosa Michx. Fl. Bor. Amer. 2:

137. 1803. C. crassifolia Dryand. in Ait. Hort. Kew. ed. 1. 3: 252.

1789; non Sess£ & Moc. Fl. Mex. ed. 2. 194. 1894. C. oblongifolia

Nutt. Journ. Acad. Phila. 7: 76. 1834. C. lanceolata var. crassifolia

Ait. ex Heynhold, Nomencl. 219. 1840.

E specie planta omnino vel fere usque ad summam pubescente

differt.

Type specimen: Collected by Andre Michaux in Carolina (Par.).

Distribution: South Carolina, Illinois, Missouri, and western

Arkansas, southwardly to Florida, Alabama, and Louisiana; else-

where probably introduced.

Specimens examined: Anon., Tampa Bay, Florida (Gray); H. H.

Babcock, Chicago, Illinois, June 17, 1874 (Field); J. M. Bigelow,

Fort Smith, Arkansas, June 23, 1853 (N.Y.); Biltmore Herb. 5693b,

dry soil, Spartanburg, South Carolina, May 12, 1897 (Gray) ; B. F.

Bush 84, common in barrens, Eagle Rock, Missouri (Gray) ; William

M. Canby 55, Columbia, South Carolina, May 9, 1899 (Gray);

A. W. Chapman, middle Florida (Field); A. H. Curtiss 6390, old

field near River Junction, Florida, May 2, 1898 (Gray) ; H. Eggert,

stony hills, Jefferson County, Missouri, June 10, 1891 (Gray; Mo.);

idem, Hematite, Missouri, May 25, 1896 (Calif.; Field; U.S.); herb.

George Engelmann, Aiken, South Carolina, April, 1882 (Mo.}; Frank

C. Gates 942, Chicago, Illinois, June 24, 1905 (Field); idem 2817,

Beach, Illinois, July 1, 1908 (Field) ;C. A.Geyer, Beardstown, Illinois

(Mo., 2 sheets); Grossly, Hyde Park, Chicago, Illinois, June 10,

REVISION OF THE GENUS COREOPSIS 345

1877 (Field); Dr. Hale, Louisiana (Field; Gray); E. J. Hill 1108,

top of limestone ledge, Lockport, Illinois, June 14, 1899 (Field);

idem 2746, eodem loco et tempore (Gray) ; G. G. Kennedy, dry pine

woods, Ormond, Florida, March 29, 1897 (Gray) ; 0. E. Lansing, Jr.

24-3, waste ground, Hyde Park, Chicago, Illinois, June 14, 1898

(Field) ; G. W. Letterman, Allenton, Missouri, August, 1883 (Gray) ;

idem, Merely, Missouri, July, 1884 (Mo., 2 sheets) ; F. E. McDonald,

high, dry hills, Peoria, Illinois, May, 1915 (Field); Mrs. J. M.

Milligan, Jefferson County, Missouri (Field); E. J. Palmer 27056,

rocky, open ground, along stream near Berry ville, Arkansas, May 7,

1925 (Gray) ; H. W. Ravenel, San tee Canal, South Carolina (Gray) ;

F. Scammon, Chicago, Illinois (Field); Earl E. Sherff 5013, cult, in

Harvard Univ. Bot. Gard., August 12, 1929 (Brit.; Calif.; Field);

idem 5014., cult., eodem loco et tempore (Field; Kew); idem 5016,

cult., eodem loco et tempore (Calif.; Field; Kew; N.Y.; U.V.);

Mary F. Spencer 1114, alt. 300 meters, ditch near San Bernardino,

California, May 9, 1919 (Gray) ; E. F. Williams, well established,

Wickford, Rhode Island, June 17, 1908 (Gray) ; R. E. Woodson, Jr. &

E. S. Anderson 1545, Birmingham, Alabama, April 20, 1927 (Mo.).

42. Coreopsis corninsularis Sherff, Bot. Gaz. 94: 597. 1933.

Herba perennis, erecta, gracilior, caulibus tenuibus angulatis

glabris 2 vel 3 ex unico basi circ. 3-4 dm. altis. Folia opposita,

plus minusve petiolata petiolis tenuibus usque ad 3 cm. longis basi

sparsim hispido-ciliatis, petiolo adjecto plerumque 3-8 cm. longa

et 1-6 mm. lata, saepius integra anguste vel late spathulato-linearia

crassiuscula faciebus glabra marginibus ciliata apice subacuta vel

subobtusa, interdum 3-5-partita foliolis lateralibus multo minoribus

et tantum 1-2 mm. latis. Capitula solitaria longe pedunculata

pedunculis tenuibus glabris 1-1.8 dm. longis, radiata, pansa ad

anthesin 3-5 cm. lata et 10-12 mm. alta. Involucri glabri bracteae

exteriores 8-10, lanceolatae vel oblongo-lineares, 3-5 mm. longae;

interiores ovato-oblongae, demum circ. 1 cm. longae. Flores ligulati

circ. 8, flavi, ligula oblongo-oblanceolati, apice saepius 3-lobati lobo

mediano multo majore apice rursus in 2 dentes inciso, 1.5-2.5 cm. longi.

Paleae tenuiter lineares, superne filiformes, 3-4 mm. longae. Flores

tubulosi flavi, styli ramis terminaliter caudato-appendiculatis.

Achaenia valde obcompressa, dorso convexa, nigra, corpore ipso

oblongo, 2.5-3.2 mm. longa et (alis exclusis) 1.2-1.5 mm. lata, facie-

bus glabra vel glabrata, marginibus alata alis expansis membranaceis

quam corpore saltern 0.5-0.75 angustioribus, apice papposa 2

fimbriatis squamellis circ. 0.1-0.2 mm. longis.

346 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: Collected by Samuel Mills Tracy, No. 8542,

on Horn Island, Mississippi, May 25, 1903 (Mo.).

Distribution: Known only from type locality on Horn Island,

Mississippi.

Specimens examined: Tracy 8542 (type, Mo.; cotype, Field).

43. Coreopsis debilis Sherff, Bot. Gaz. 89: 366. 1930.

Herba perennis, caulibus saepius 2-6, e radice lignescenti erectis,

tenuissimis, angulatis, glabris, 3-6 dm. altis. Folia opposita,

membranacea, petiolata petiolis tenuibus usque ad 2 cm. longis,

lamina nunc indivisa linearia vel parce lineari-oblanceolata margine

integra ciliataque apice vix acuta basim versus sensim angustata

faciebus saepe aegre hispida petiolo adjecto 2-5 cm. longa et 1-5 mm.

lata; nunc majora, petiolo saepe 3.5 cm. longo adjecto 7-9 cm. longa,

aegre pinnata (raro etiam subbipinnata) segmentis lateralibus ple-

rumque 1-2 jugis, quam terminal! minoribus, linearibus vel subfili-

formibus. Capitula pauca vel numerosa, tenuiter pedunculata

pedunculis 1-2 (-3) cm. longis, radiata, pansa ad anthesin circ.

2-2.5 cm. lata et circ. 7-9 mm. alta. Involucri saepius glabrati

bracteae exteriores circ. 8, lineares lanceolataeve, apice acutae vel

obtusae, margine ciliatae, basim versus saepe scariosae, 3-4 mm.

longae; interiores lanceolatae vel ovato-lanceolatae, 7-8 mm. longae.

Flores ligulati flavi, ligula cuneato-obovati, apice saepius 4-dentati

dentibus acutis, circ. 1 cm. longi. Paleae lineares, superne angustis-

sime elongatae. Disci florum stigmata terminaliter caudata.

Achaenia obcompressa, tergo papillato-rugoso convexa, corpore

ipso atro circ. 2 mm. longa et circ. 1 mm. lata, marginibus anguste

alata, apice 2-squamellata squamellis fimbriolatis; facie ventrali

glabra vel papillato-scabra, supra infraque valde callosa.

Type specimen: Collected by John Donnell Smith, No. 602, in

dry, sandy, old fields along coast of Mississippi Sound, Harrison

County, Mississippi, September 15, 1885 (Field).

Distribution: Florida and Georgia westward to Mississippi.

Specimens examined: A. H. Curtiss, Pensacola, Florida, summer

of 1885 (N.Y.) ;F. E. Lloyd & S. M. Tracy 527a, Gulfport, Mississippi,

September 8, 1900 (N.Y.); Charles Mohr, dry, open, sandy hills,

etc., eodem loco, July 4, 1874 (U.S., 2 sheets); idem, in woods, Cull-

man, Alabama, June 1, 1882 (Gray) ; idem, low, open places, Cullman,

June, 1883 (U.S., 3 sheets); idem, dry hillsides, vicinity of Ashland,

Bibb County, Alabama, June, 1883 (U.S.) ; idem, borders of woods and

fields, dry openings, Mulberry River Valley, Blount County, Ala-

REVISION OF THE GENUS COREOPSIS 347

bama, June 6, 1883 (U.S.) ; idem, sandy soil in open copses, Columbus,

Mississippi, June 4, 1888 (U.S.); idem, dry bald prairies, Gallion,

Alabama, May 25, 1893 (U.S.); Charles L. Pollard & William R.

Maxon 4-96, Lithonia, Georgia, August 12, 1900 (N.Y.) ; John Donnell

Smith 602 (type, Field; cotype, Gray).

A species more closely related to Coreopsis lanceolata L. and C.

grandiflora Hogg. J. Donnell Smith had determined his plant as C.

lanceolata var. angustifolia Torr. & Gray. That variety, however,

has a very different habit, the plants being less branched and the

leaves being basally clustered and almost always simple. Donnell

Smith had shown Asa Gray his Mississippi material, and Gray had

called it "a peculiar form" of C. lanceolata L. Mohr's material from

Pensacola, Florida, likewise had been seen by Gray. Thus, one

sheet bears a note stating: "Dr. Gray remarks: 'I have nothing like

this. I suppose it may be an altered, coast form of C. lanceolata.

But it is very peculiar.' ' (This note is written on the back of a card

which, by strong transmitted light, is seen to have been Donnell

Smith's regular herbarium label. This would seem to indicate that

Mohr and his contemporary had studied their several specimens

mutually and had regarded them as identical.) This same sheet has

Mohr's first determination, Coreopsis lanceolata var. angustifolia,

but this he had subsequently crossed out and replaced with C.

grandiflora. His other foregoing specimens all had been labeled by

him as C. grandiflora. This interpretation was retained by him in

his Plant Life of Alabama (Contr. U. S. Nat. Herb. 6: 805. 1901).

From C. grandiflora, however, our plants differ in their more slender

and wiry, less herbaceous stems, in the tendency for some entire

stems to have minute, simple leaves (smaller than the simple leaves

formed in C. lanceolata), and in the tiny achenes, which have the

body proper about 2 mm. long and about 1 mm. wide, as against

about 2.5 mm. long and about 1.4-1.8 mm. wide in C. grandiflora.

(A similar or even more pronounced achenial difference separates

C. debilis also from C. lanceolata. Furthermore, the achenes of C.

debilis have narrow wings [0.2-0.5 mm. wide] while those of C.

grandiflora and C. lanceolata have wings 1 mm. or so in width.)

44. Coreopsis intermedia Sherff, Bot. Gaz. 88: 299. 1929.

Herba erecta, plus minusve glabra, forsitan perennis, supra parce

ramosa, ±6 dm. alta, caule subtereti vel moderatim angulato,

sulcato. Folia opposita, simplicia, basalia longe tenuissimeque

petiolata petiolis usque ad 4.5 cm. longis, laminis oblongo-oblance-

olatis vel anguste obtuseque obovatis; principalia caulina sessilia,

348 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

late oblongo-lanceolata, ciliata, crassiuscula, apice subacuta, 5-7

(-9.5) cm. longa et 1.2-2 (-3.2) cm. lata. Capitula pedunculata

pedunculis 1.5-2.5 dm. longis, radiata, pansa ad anthesin 3.5-4

dm. lata et ±12 mm. alta. Involucri bracteae exteriores 8-10,

lanceolatae vel lineari-lanceolatae, tergo glabratae, margine saepe

diaphana ciliatae, apice acutae cartilagineaeque, 4-7 (rariter -8)

mm. longae; interiores late lanceolatae, plerumque 12-14 mm.

longae. Flores ligulati circ. 8, flavidi, unicolores, obovati vel late

oblanceolati, apice trilobati lobo mediano valde emarginato, circ.

1.5 cm. longi. Paleae superne elongatae et valde caudato-attenu-

atae. Disci florum stigmata apice caudata. Achaenia suborbi-

cularia, valde obcompressa, dorsaliter convexa, brunneo-atra, alata

alis membranaceis planis vel rarissime parce incurvatis 0.2-0.4 mm.

latis, apice saepe bidenticulata, faciebus perspicue tuberculata,

facie ventrali raro callosa, corpore ipso oblongo-obovato vel late

oblongo-oblanceolato 2-3 mm. longa et 1.3-2 mm. lata.

Type specimen: Collected by Julian Reverchon, No. 2077 pro

parte, in sandy woods, Mineola, Texas, June 12, 1900 (Berl.).

Distribution: Known only from type locality and vicinity, Texas.

Specimens examined: Reverchon, Pine Island, May 5, 1903 (Mo.);

idem 2041, sands, Big Sandy, May 27, 1901 (Mo., 2 sheets); idem

2077 pro parte (type, Berl.; cotype, Mo.); idem 2077 pro parte,

sands, Mineola, June 10, 1900 (Mo.).

The stems of the plants examined are uniformly leafy from bottom

to top as in Coreopsis pubescens and the leaves resemble the glabrous,

undivided ones occasionally found in that species. The elongate

peduncles, however, also the pronounced differentiation between

exterior and interior involucral bracts, are more as in C. lanceolata,

although not typical for that species.

45. Coreopsis pubescens Ell. Sketch Bot. S. Carol. & Georgia

2: 441. 1824. C. auriculata Schkuhr, Handb. ed. 2, pi. 260. 1808

(non L.). C. auriculata var. 7 Torr. & Gray, Fl. N. Amer. 2: 343.

1843. C. auriculata var. 8 Torr. & Gray, op. cit. 344 (excl. syn. quod

C. pubescens var. robusta est).

Plus minusve pubescens vel hirsuta. . . .C. pubescens sensu stricto.

Caulis ramique glabri vel glabrati var. 0 robusta.

Herba perennis, erecta, simplex vel paulum ramosa ramis sub-

erectis, plus minusve pubescens vel hirsuta, 6-12 dm. alta, caule

usque ad summam folioso. Folia opposita, polymorpha, saepius

5-10 cm. longa, inferiora longe (2-5 cm.) tenuiterque petiolata altera

REVISION OF THE GENUS COREOPSIS 349

saepe sessilia, laminis nunc integris ovalibus vel oblongo-ovatis vel

elliptico-lanceolatis nunc (plus minusve irregulariter) 3-5-partitis

foliolis lateralibus plerumque multo minoribus ac oblonge lineari-

lanceolatis. Capitula tenuiter pedunculata pedunculis 1-2 dm.

longis, radiata, pansa ad anthesin 3-5 cm. lata et 8-11 mm. alta.

Involucri bracteae subaequales plus minusve glabratae, exteriores

8-10, lineari-lanceolatae, plurivittatae, 7-10 mm. longae; interiores

ovatae. Flores ligulati circ. 8, flavi, ligula cuneati vel oblongo-

cuneati, apice 3-lobati lobo mediano rursus 2-4-lobulato, 1-2.3 cm.

longi. Paleae inferne latiores superne elongato-filiformes, demum

6-8 mm. longae, achaenia multum superantes. Florum tubulosorum

stylorum rami apicem versus abrupte angustati et lineari-appendi-

culati. Achaenia obcompressa, suborbiculata, circ. 2.8-3 mm. longa,

corpore ipso nigra, alis (±0.5 mm. latis) planis integris saepe pur-

purea, faciebus glabra vel papillata, ventraliter apice basique saepius

gibbo cartilagineo munita, apice vero squamellis fimbriatis plerumque

2-dentata.

Type specimen: None cited, but habitat given as "western dis-

tricts of Georgia."

Distribution: Virginia, Illinois, and Oklahoma southward to

Florida, Alabama, and Louisiana. Sometimes escaped or adventive

elsewhere.

Specimens examined (it may well be that these determinations,

most of which were made many years ago, include a few specimens

which are properly referable to the var. robusta) : Samuel M. Bain 95

p.p., fields, Henderson, Tennessee, May, 1893 (Gray); H. C. Beardslee

& C. A. Kofoid, alt. 900 meters, Great Smoky Mountains, Swain

County, North Carolina, August, 1891 (Field; Gray); Beyrich,

grassy banks of rivers, Georgia (Berl., 2 sheets);/7. Boynton, High-

lands, North Carolina, 1889 (Gray; Mo.); Buckley, Alabama (Gray);

B. F. Bush 172A, uncommon, in dry ground, McDonald County,

Missouri, September 1, 1893 (Gray); idem 6041, woods, Webb City,

Missouri, July 23, 1910 (N.Y.) ; idem 6137, woods, Monteer, Missouri,

August 8, 1910 (Gray); idem 6531, eodem loco, October 22, 1911

(Mo.); idem 7872, open woods, eodem loco, October 8, 1916 (Gray);

Carpenter, Louisiana (Gray); A. W. Chapman, mountains of Georgia

(Field); A. H. Curtiss 17, rich woods, Jackson County, Florida,

June 18, 1886 (Gray; foliis sed non achaenis C. auriculatae adpropin-

quans); H. Eggert, dry woods, Jefferson County, Missouri, July 8,

1879 (Gray) ; George Engelmann, Mountain Creek bottom, Missouri,

1833 (Mo.); idem, St. Louis, Missouri, August, 1863 (Berl.); Henry

350 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

A. Gleason 2749, dry, upland woods, Makanda, Illinois, July 22, 1902

(Gray); idem 2750, cleared uplands along railroad, eodem loco et

tempore (Gray) ; P. Graebner, cult, in Hort. Berol. ex sem. ex Amer.

Boreali missis, August 5, 1910 (Berl.); J. M. Greenman 3771, Shef-

field Mountain, near Arcadia, Missouri, July 22, 1915 (Mo.); H.E.

Hasse, Desoto, Missouri, July, 1887 (Field); Elihu Hall, central

Illinois (Gray); W. Hoffman, Corn Creek, vicinity of Rolla, Mis-

souri, July 15, 1871 (Berl., 2 sheets); Hort.Berol, 1806-1812 (Berl.);

Hort. Lips, ex Amer. septentr. (Berl.); Hort. Paris, 1816 (Berl.);

H. W. Houghton 3864, common, open woods, near Shawneetown,

Oklahoma, May 28, 1916 (Gray); H. D. House 4023, below 900

meters alt., sandy soil along A very 's Creek, Pisgah Forest, North

Carolina, September, 1908 (Gray); John H. Kellogg, Allenton,

Missouri, August 12 and 13, 1884 (Mo.) ; idem, St. Louis, Missouri,

July 15, 1903 (Mo.); George W.Letterman, Allenton, Missouri (Mo.);

Kenneth K. Mackenzie 328, common in woods, Pleasant Grove,

Missouri, July 21, 1897 (Field); Michaux, North America (Berl.);

E. J. Palmer 2317 and 2317 A, open woods, Forest Mill, Missouri,

June 23, 1909 (Gray); idem 2513, barrens, Joplin, Missouri, July 20,

1909 (Gray) ; idem 29347, rich, moist soil, vicinity of Beaver, Arkan-

sas, October 24, 1925 (Gray); Thomas C. Porter, Hawk's Nest on

New River, 9 miles above Kanawha Falls, West Virginia, July 21,

1880 (Field; N.Y.); Ns. Riehl 65, very moist ground, St. Louis,

Missouri, July, 1838 (Berl.); F. Rugel, in rocky, wet places of the

mountains at Broad River, North Carolina, June-July, 1841 (Berl. ;

N.Y.); idem, in open places and along roads, between Quincy and

Aspalaga, Florida, May, 1843 (Mo.); Albert Ruth 655, along railroad,

Valley Head, Alabama, July, 1898 (Mo.); idem 665, Lulu Falls,

Lookout Mountain, Georgia, July, 1898 (N.Y.); E. M. Shepard,

Greene County, Missouri, July, 1880 (Gray); Earl E. Sherff 989,

southeast of Pacific, Missouri, August 7, 1910 (Gray); idem 5005,

cult., Harvard Univ. Bot. Gard., August 12, 1929 (Berl.; Field);

C. W. Short 42, Lexington, Kentucky, 1831 (Gray) ; John K. Small,

alt. 750-1,500 meters, on slopes of Thomas Bald, boundary of North

Carolina and Georgia, August 19, 1893 (Field); idem, alt. 1,560

meters, summit of Thomas Bald, August 19, 1893 (Field) ; idem, base

of Currahu Mountain, Toccoa, Georgia, July, 1896 (N.Y.); Huron

H. Smith 1089, along railway tracks, Des Arc, Missouri (Field);

idem 2456, Blue Ridge Mountains, Fannin County, Georgia, July 22,

1909 (Field) ; idem 2582, eodem loco, August 7, 1909 (Field) ; John

Donnell Smith, alt. 1,410 meters, north face of cliffs, Wildcat Ridge,

REVISION OF THE GENUS COREOPSIS 351

west flank of Whitesides Mountain, 6 miles northeast from High-

lands, North Carolina, August 21, 1882 (Gray); idem, alt. 1,440

meters, face of cliffs, eodem loco et tempore (Field; Mo.); idem,

alluvial banks of river at Cumberland Falls, Whitler County,

Kentucky, August 23, 1883 (Field) ; idem, thickets, alluvial banks of

Soquee River, Habersham County, Georgia, September 13-15,

1883 (Field; Gray); idem, edge of woods, banks of Oconee River,

above Milledgeville, Georgia, August 16, 1884 (Field; Gray); idem,

gullies in high, wooded hills west of Cahawba River, Perry County,

Alabama, September 2, 1885 (Field; Gray); G. W. Stevens 2387,

shady, grassy thicket in woods near Ottawa, Oklahoma, August 27,

1913 (Gray); Roland Thaxter, Cullowhee, North Carolina, June-

July, 1887 (Gray); R. E. Woodson, Jr. 655, Crawford County,

Missouri, July 10, 1926 (Mo.).

Passes into the var. robusta.

Coreopsis pubescens var. /3 robusta A. Gray ex Eames,

Rhodora 18: 239. 1916. C. testudinea Shuttl. in herb.

E specie caule ramisque glabris vel glabratis differt.

Type specimen: Collected by Asa Gray and John Carey, North

Branch of New River, North Carolina, July, 1841 (Gray).

Distribution: Virginia and North Carolina westward to

Kentucky.

Specimens examined : Biltmore Herb. 307c, slopes of Cedar Cliff

Mountain, Buncombe County, North Carolina, August 2, 1897

(Gray; Mo.); John Carey, North Carolina (N.Y.); idem & Asa

Gray, North Branch of New River, etc. (type, Gray; cotype, N.Y.);

iidem, Virginia, July, 1841 (Gray) ; E. H. Eames 8825, growing about

1 meter tall, waste grounds, Bridgeport, Connecticut, July 21, 1914

(Gray) ; M. L. Fernald & Bayard Long 19265, fallow field northeast of

East Brewster Station, Brewster, Massachusetts, July 20, 1919

(Gray); P. A. Rydberg 9445, alt. 1,650-1,800 meters, Craggy Moun-

tains, Buncombe County, North Carolina, July 21, 1925 (N.Y.;

Pom.); John Donnell Smith, growing 4 ft. high, upland meadow,

Richland Gap, Heywood County, North Carolina, August 14, 1882

(Gray; Mo.) ; idem, alluvial soil and sandy bank of river, Cumberland

Falls, Whitley County, Kentucky, August 23, 1883 (Gray; Mo.).

Originally regarded by Asa Gray (Amer. Journ. Sci. 42: 45. 1841)

as a ''larger form of Coreopsis auriculata, with nearly all the leaves

undivided." At times very distinct, again scarcely distinguishable

from the species proper.

352 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

46. Coreopsis heterolepis Sherff, Bot. Gaz. 89: 365. 1930.

Herba perennis, glabra, caule angulata, ramosa, dense foliosa,

3-6 dm. alta. Folia membranacea plus minusve dimorpha: basalia

dense congregata, petiolata petiolis planis angustissime alatis usque

ad 6 cm. longis, petiolo adjecto 8-12 cm. longa, indivisa spathulata

vel oblonga apice obtusa, alia pinnata foliolis nunc linearibus nunc

oblanceolatis nunc oblongis nunc rhomboideo-subovatis ac etiam

1-1.7 cm. latis, lateralibus saepius 1-2 jugis ac multo minoribus;

caulina numerosissima, petiolata petiolis anguste marginatis marginis

basi rarius setis pluriloculatis sparsim positis obsita circ. 1-2.5 cm.

longis petiolo adjecto circ. 6-10 cm. longa, plerumque pinnata, foliolis

anguste linearibus, 2-8 cm. longis et 0.3-2.5 mm. latis, apice sub-

acutis, lateralibus 1-2 jugis. Capitula tenuissime pedunculata

pedunculis 1-2 dm. longis, radiata, pansa ad anthesin 3-4 cm. lata

et 5-7 mm. alta. Involucri glabri vel subglabri bracteae exteriores

angustissime lineares, saepius patentes, apice acutae, 5-11 mm.

longae; interiores ovato-lanceolatae nunc breviores nunc longiores.

Flores ligulati circ. 8, omnino flavi, ligula cuneato-obovati, apice

obtuse 4-dentati, circ. 1.5-1.8 cm. longi. Paleae superne capillares,

circ. 4-5 mm. longae. Disci florum stigmata caudato-elongata.

Achaenia minima plano-convexa, corpore nigro oblongo vel oblongo-

ovato ipso tantum circ. 1.3-1.7 mm. longa et circ. 1-1.5 mm. lata,

faciebus levia vel papillato-rugosa, marginibus anguste alata alis

rubris circ. 0.2-0.4 mm. latis, apice minutissime 2-squamellata

squamellis fimbriolatis.

Type specimen: Collected by Ernest Jesse Palmer, No. 6962A,

on dry, sandy bluffs, Heber Springs, Cleburne County, Arkansas,

October 30, 1914 (Mo., 4 sheets).

Distribution: Known only from type locality.

Specimens examined: Palmer 6962A (4 type sheets, Mo.).

47. Coreopsis grandiflora Hogg ex Sweet, Brit. Fl. Card. 2:

pi. 175. 1826; Nutt. Trans. Amer. Phil. Soc. 2, 7: 358. 1841. C.

Boykiniana Nutt. loc. cit. C. heterophylla Nutt. loc. cit. (nee alior.).

C. grandiflora var. subintegrifolia Torr. & Gray, Fl. N. Amer. 2: 345.

1843.

Caules foliaque pubescentia vel piloso-hispida var. 7 pilosa.

Caules et foliorum facies plerumque glabrata.

Foliorum segmenta omnia anguste linearia vel quidem filiformia.

var. 8 Harveyana.

Foliorum simplicium lamina vel divisorum segmenta latiora.

REVISION OF THE GENUS COREOPSIS 353

Folia caulis basim versus adgregata, pedunculis crassis elongatis

(saepe 2-3 dm. longis) -. var. /3 longipes.

Folia subaequaliter distributa, pedunculis tenuioribus saepius

0.8-1.5 dm. longis C. grandiflora sensu stricto.

Herba perennis vel raro annua, erecta vel adscendens, plerumque

non pubescens nisi petiolis ciliatis, 3-6 dm. alta, saepe ramosa,

plerumque foliosa. Folia opposita, plus minusve petiolata petiolis

saepius 0.5-4 cm. longis, inferiora simplicia vel irregulariter partita

lamina (vel segmento terminali) spathulata vel lanceolata, petiolo

adjecto saepius 5-10 cm. longa; alia plerumque 3-5-partita (raro

simplicia) segmentis interdum rursus 3-5-partitis segmentis ultimis

linearibus vel lanceolato-linearibus, plerumque 1.2-5 mm. latis.

Capitula saepius solitaria, moderate pedunculata pedunculis tenui-

bus saepius 1-1.5 dm. longis, radiata, pansa ad anthesin 3-6 cm. lata

et 8-12 mm. alta. Involucri bracteae exteriores 7-10, saepe dupli-

citer dispositae, lanceolato-subulatae, infra saepe subdilatatae, mar-

gine ciliatae et plus minusve albidae, 5-9 (raro -18) mm. longae;

interiores ovatae, nunc paulo longiores nunc paulo breviores. Flores

ligulati plerumque 8, flavi, ligula cuneato-obovati, apice 3-lobati

lobo mediano majore rursus 2-partito, 1.3-2.5 cm. longi. Paleae

lineares, infra latiores supra attenuatae filiformesque, demum circ.

6-7 mm. longae. Flores tubulosi summa aurantiaci, stylorum ramis

apice perspicue cuspidati. Achaenia obcompressa et dorso convexa,

circumambitu (alis inclusis) orbiculata, circ. 2.5 mm. longa, corpore

nigra, faciebus glabra vel minute papillata, ventraliter apice basique

saepe verruca callosa magna ornata, marginibus alata alis planis vel

vix incurvatis quam corpore dimidio angustioribus, apice minute

biaristata aristis squamellatis fimbriolatisque vel demum calva.

Type specimen: Sent by a Mr. Hogg from New York to England

as a new species and represented by Robert Sweet's type plate

(loc. cit.).

Distribution: Missouri and Kansas southward to Florida,

Louisiana, and New Mexico.

Specimens examined: H. C. Benke 3279, Pensacola, Florida,

March 9, 1922 (Field) ; George D.Butler 16, prairies, 1.5 miles north

of Limestone Gap, Oklahoma, June 13, 1877 (Field) ; F. S. & Esther

S. Earle 73, 4 miles northwest of Auburn, Alabama, July 13, 1899

(Mo.);EsemE. Hall, Texas, 1877 (Gray); W. H. Haller 982, low

ground, Wilson County, Kansas, 1895 (Gray) ; ex Hort. Berol. (Gray) ;

H. W. Houghton 3567, common on hillsides, vicinity of Tishomingo,

Oklahoma, April 15, 1916 (Gray); Dr. Leavenworth, Louisiana

354 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

(Gray); Le Conte, North America (Gray); George W. Letterman,

Gratiot, Missouri, May 30, 1897 (Field) ; idem, along Frisco Railroad

near St. Louis, Missouri, August, 1899 (Mo.)',Lindheimer, Houston,

Texas, April, 1842 (Gray); Kenneth K. Mackenzie 100, frequent,

introduced along railroads, Kansas City, Missouri, June 5, 1898

(Field); Dr. Pitcher, Arkansas (Gray); J. Reverchon, sandy loams,

woods, Dallas, Texas, May, 1874 (Mo.); idem, Dallas, May, 1876

(Gray); idem 513, sandy forests, Dallas, May, 1883 (Mo.); idem

1478, rich sands, Dallas, May (Berl.; Field); Albert Ruth 482, in

rich woods near Arlington, Texas, May 24, 1920 (Field, 2 sheets);

J. H. Schuette 76, cult., Green Bay, Wisconsin, August 28, 1900 (Field) ;

John K. Small, alt. 300-330 meters, on Little Stone Mountain, De

Kalb County, Georgia, July 7, 1893 (Field, 2 sheets; Gray); H. E.

Wheeler 62, near Hazen, Grand Prairie, Arkansas, May 22, 1924

(Field) ; Charles Wright, Texas (Gray).

Coreopsis grandiflora var. ft longipes (Hook.) Torr. & Gray,

Fl. N. Amer. 2: 345. 1843. C. longipes Hook. Bot. Mag. pi. 3586.

1837.

Folia caulis basim versus dense vel moderate adgregata, plerum-

que simplicia vel pinna ta; pedunculis crassioribus elongatis saepe

2-3 dm. longis.

Type specimen: Collected by Thomas Drummond in Texas and

illustrated by Hooker with type plate (loc. cit.).

Distribution: Texas.

Specimens examined : Elihu Hall, Texas, 1877 (Field); idem 342,

wet prairie, Houston, April 18, 1872 (Field) ; Earl E. Sherff 278, cult.,

St. Louis, Missouri, July 7, 1910 (Field).

Coreopsis grandiflora var. 7 pilosa Sherff, Bot. Gaz. 89: 368.

1930.

A specie caulibus foliisque pubescentibus saepe etiam longe pa-

tenteque piloso-hispidis differt.

Type specimen: Collected by Steivardson Brown, Nathaniel L.

Britton, and Peter Bisset, No. 2011, cultivated at Agricultural Station,

Island of Bermuda, May 22-June 6, 1914 (N.Y.).

Distribution: Known in a native state only from northern

Georgia.

Specimens examined : Brown, Britton, & Bisset 2011 (type, N.Y.);

John H. Kellogg, cult., Missouri Bot. Card., May 23, 1911 (Mo.);

John K. Small, alt. 210-300 meters, between Alcovy River and No

REVISION OF THE GENUS COREOPSIS 355

Business Creek, Oconee and Gwinnett Counties, Georgia, July 14,

1893 (Field, 2 sheets; U.S.); idem, alt. 225 meters, on banks of

Yellow River near McGuire's Mill, Gwinnett County, July 11, 1893

(Mo.) ; idem, alt. 255 meters, on banks of Yellow River, near Yellow

River (Store), Gwinnett County, July 20, 1893 (Field).

The two cultivated specimens examined have a very unique aspect

because of their marked hispidity. In spontaneous material the hairs

are shorter and less spreading.

Coreopsis grandiflora var. 5 Harveyana (A. Gray) Sherff,

Bot. Gaz. 94: 593. 1933. C. Harveyana A. Gray, Syn. Fl. N. Amer. 1,

pt. 2: 292. 1884.

Foliorum segmenta anguste linearia vel quidem filiformia;

achaeniis paulo minoribus, circ. 2 mm. longis.

Type specimen: Collected by Francis LeRoy Harvey, No. 22,

on cliffs near Fort Smith, Arkansas, June (Gray).

Distribution: Missouri and Arkansas, perhaps also Kansas and

Oklahoma.

Specimens examined: Anon,, Arkansas National Forest, Arkan-

sas, 1909 (U.S.); 5. F. Bush 158, Sheffield, Missouri, June 12, 1894

(Berl.); Frederick V. Coville 127 Ark., Mountain Park near Little

Rock, Arkansas, July 17, 1887 (U.S.); H. Eggert, Williamsonville,

Missouri, June 11, 1893 (Field) ; F. L. Harvey 22 (type, Gray, cum

C. tinctoria Nutt. pro parte parva commixta); E. J. Palmer 8120,

wet, open hillsides, Malvern, Arkansas, June 23, 1915 (Mo.); H. S.

Reynolds, Judsonia, Arkansas, May 31, 1877 (Field); Huron H.

Smith 1150, railway tracks, Des Arc, Iron County, Missouri, June 8,

1908 (Field).

The type sheet of C. Harveyana Gray bears 3 specimens. The

first at the left is C. tinctoria Nutt., having the oblong, wingless

achenes, the bicolored ligules, the subtruncate style branch tips, the

linear-oblong paleae, etc. of that species. The other two specimens

are very distinct, having orbiculate, winged achenes, caudate-conical

tips to the style branches, upwardly elongate-attenuate paleae, etc.

Gray clearly was misled by the admixture of the small C. tinctoria

specimen, for in his Synoptical Flora he separated C. Harveyana

partly on the basis of the "rays sometimes brown-purple at base."

Had the rays of C. Harveyana really possessed the tendency toward

a bicolored state, a separation of the type as a new species would have

been justified. Since the rays were really yellow, however, the type

had nothing left to separate it from C. grandiflora proper except the

356 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

narrower leaf divisions and the slightly smaller achenes. These

differences Gray himself elsewhere (e.g. C. Drummondii Torr. &

Gray, et var. Wrightii ipsi, Syn. Fl. N. Amer. 1, pt. 2: 291. 1884)

considered as connoting a variety. (A supposedly duplicate sheet of

the Harvey material at St. Louis [Mo.] bears all C. tinctoria Nutt.!)

48. Coreopsis auriculata L. Sp. PL 908. 1753. Chrysanthemum

hirsutum Virginian, auriculato Dulcamaraefolio octopetalon. Chry-

santh. etc. Plukenet Aim. pi. 83, f. 5. 1691; Pluk. Phytogr. pi. 242,

f. 4. 1769. Chrysanthemum trifoliatum minus Virgin, foliis obtusio-

ribus, hirsutis Morison Hist. Univ. Oxon. 3, sect. 6, pi. 3, f. 45 (46).

1715. Cymbaecarpa auriculata (L.) Hort. Reg. Matrit. ex Elench. PI.

Hort. Bot. Destremx 1805: 19. 1806. Leachia trifoliata Cass. Diet.

Sci. Nat. 25: 389. 1822 (ex synon. C. auriculata L.). Chrysomelea

auriculata (L.) Tausch Hort. Canal. 1823. Coreopsis auriculata var.

diversifolia Ell. Sketch Bot. S. Car. Ga. 2: 437. 1824. Chysomeka

auriculata (L.) Tausch ex DC. Prodr. 5: 571. 1836 (sphalm). Core-

opsis auriculata var. a Ell. ex DC. loc. cit. Coreopsis auriculata

var. glabrata DC. loc. cit. Anacis auriculata (L.) Schrank, Denk-

schr. Akad. Muench. Math. Nat. 5: 7. 1817. Coreopsis diversifolia

DC. loc. cit. (nee alior.). Coreopsis auriculata var. /3 Torr. & Gray,

Fl. N. Amer. 2: 343. 1843. Coreopsis oculata Hort. in Handlist

Hardy Rock PI. Harvard Univ. Bot. Gard. 10. 1927 (sphalm).

Herba perennis, erecta vel adscendens, simplex vel ramosa,

pubescens vel hirsuta vel supra interdum glabrata, inferne foliosa,

basi stolonifera. Folia opposita, petiolata petiolis nunc tenuibus

nunc latis saepe elongatis itaque laminas plurimum excedentibus,

petiolo adjecto plerumque 4-12 cm. longa; lamina membranacea,

ovato-orbiculata vel elliptico-ovata, apice obtusa vel subacuta,

lateribus integra vel 1-2-lobata lobis parvis basalibus. Capitula

solitaria ramos elongates (saepius 1-2.5 dm. longos) nudos termi-

nantia, radiata, pansa ad anthesin 3.5-5 cm. lata et circ. 9-10 mm.

alta. Involucri bracteae subaequales, exteriores circ. 8, pallidae,

ovato-oblongae vel lanceolato-oblongae, apice rotundatae vel sub-

acutae, lateribus saepe tenuissime albido-marginatis glabratae vel

hispido-ciliatae; interiores ovatae. Flores ligulati circ. 8, flavi, ligula

obovati vel late cuneati, apice 3-lobati lobo mediano emarginato,

1.5-2.3 cm. longi. Paleae anguste lineares, superne attenuatae,

demum 6-9 mm. longae. Florum tubulosorum ramorum styli

terminis caudato-cuspidati. Achaenia oblanceolata vel obovata,

obcompressa, incurvata, nigra, glabra vel minute papillata, margi-

nibus anguste alata alis demum cartilagineis involutisque, 2.2-2.9

REVISION OF THE GENUS COREOPSIS 357

mm. longa, apice primo minute bisquamellata demum plerumque

calva.

Type specimen: Linnaeus' first citation of literature is that of

"Gron. virg. 105." This refers to a plant collected by John Clayton

in Virginia and perhaps still extant in London (Brit.). Linnaeus cited

also the two illustrations by Plukenet (loc. cit.) and the one by

Morison (loc. cit.); all three are clear and agree unmistakably with

each other.

Distribution: Virginia and Kentucky southward to northern

Florida, Mississippi, and Louisiana. Cited also for Illinois by

Robinson and Fernald (Gray's New Man. 838. 1908).

Specimens examined : Biltmore Herb. 176, rich woods, Biltmore,

North Carolina, May 15, 1896 (Field ; Gray) ; idem 1 76b, eodem loco,

May 20, 1897 (Field; Gray); ex herb. G. W. Bischoff (Gray); S. B.

Buckley, Alabama, June, 1841 (Mo.); idem, Virginia (Gray); Wil-

liam M. Canby 53, Stone Mountain, Georgia, May 3, 1899 (Gray) ;

idem 54, Chattahoochie, Georgia, May 2, 1899 (Gray); A. W. Chap-

man, mountains of Georgia (Field); Miss M. A. Coe, Atlanta,

Georgia, May 6, 1903 (Gray); A. H. Curtiss, Bedford County,

Virginia, June 1, 1871 (Gray); idem, eodem loco, May 31, 1872

(Field); idem 1485, rich woods, Jackson County, Florida, June

(Berl.; Field); F. S. Earle & C. F. Baker, Auburn, Alabama, April 4,

1897 (Field) ; J. Hale, Alexandria, Louisiana (Gray) ; T. G. Harbison,

Highlands, North Carolina, May, 1902 (Gray); R. A. Harper 74,

Talladega County, Alabama, April 16, 1906 (Gray); ex herb. Dr.

A. C. Hexamer & Dr. F. W. Maier, rich woods, Soluda River, Lex-

ington District, South Carolina, June 14, 1855 (Gray); ex Hort.

Berol. (Gray); M. E. Hyams, Statesville, North Carolina, July,

1880 (Pom.); C. Mohr, open woods throughout the mountains,

Lawrence County, Alabama, May, 1880 (Field); ex herb, eiusdem,

Winston County, Alabama, May 1, 1881 (Field) ; P. H. Rolfs 392,

Pensacola, Florida, April 11, 1894 (Field) ; Albert Ruth, open woods,

near Knoxville, Tennessee, May, 1895 (Gray; Pom.); idem 67, rich

woods, Knox County, Tennessee, June 15, 1900 (Gray); idem 652,

copses, Knoxville, May, 1898 (Mo.) ; C. W. Short, Lexington, Ken-

tucky, 1835 (Gray); J. K. Small & A. A. Heller, near R. and D.

Railroad, between Fall Creek and Danville, Virginia, June 3, 1891

(Field); ex Dr. Steetz, ex Lexington, Kentucky (Berl.); J. T. Stewart,

northern Mississippi, 1863 (Field); K. A. Taylor, dry, sandy pine

woods, Columbia, South Carolina, May, 1890 (Field) ; G. R. Vasey,

North Carolina, 1878 (Field) ;E. H. Wharf, North Carolina (Gray).

358 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

49. Coreopsis congregata Blake, Journ. Wash. Acad. Sci.

19:275.1929.

Herba annua, gracilis, erecta, subsimplex, sparsim pubescens,

1-3.5 dm. alta, caule sulcato-striato, sparsim piloso praecipue

superne pilis plerumque adscendentibus, internodiis 2-5 cm. longis.

Folia opposita sessilia vel petiolata petiolis pilosissimis usque ad 1

cm. longis, omnino circ. 2-4 cm. longa, pinnatifida vel inferiora

bipinnatifida, circumambitu triangulata; segmentis ultimis mem-

branaceis, oblongis vel ovatis vel lanceolato-ovatis, acutis vel obtusis,

calloso-mucronulatis, utrinque pilosis, marginibus ciliatis, jugo infimo

paene distincto. Capitula pauca (1-7), nunc solitaria mine corym-

bosa, tenuissime pedunculata pedunculis sparsim pilosis erectis

nudis vel minute 1-2-bracteatis 2-5 cm. longis, radiata, pansa ad

anthesin 1-1.5 cm. lata et 0.5-0.7 cm. alta. Involucri pilosi bracteae

exteriores circ. 8, tenues, anguste spathulatae vel lineares, obtuse

calloso-apiculatae, 2.5-3.5 mm. longae et circ. 0.7 mm. latae, 3-

nervatae nervo mediano multo magis perspicuo; interiores elliptico-

oblongae, apice obtuso ciliolatae, aureae, 5-7-nervatae, 4.5-4.8 mm.

longae et circ. 1.8 mm. latae. Flores ligulati 8, aurei, steriles, ligula

elliptico-oblongi, apice subintegri, tantum circ. 5-nervatae, circ.

6-7.5 mm. longi et 2.5-2.8 mm. lati, tubo glandulis stipitatis instructo.

Paleae lineares, acutae vel obtusae, glabrae, flavae, circ. 4-vittatae,

circ. 3.5 mm. longae. Flores tubulosi 12-20, corollis aureis tubo

glandulis stipitatis instructis, stylorum ramis termino anguste

caudato-appendiculatis. Achaenia exteriora obcompressa, obovata

vel cuneato-rotundata, brunneo-atra, glabra, tergo levia vel superne

plus minusve muricata, ventris mediane unicostati ac crustaceo-

mamillati basi callosa, marginibus incrassatis subbrunneis circ.

0.3 mm. latis longitudinaliter rugulosa infra sinuata supra in lobos

paucos rotundatos divisa, apice parce depressa ac exaristata, 2-2.3

mm. longa et 1.3-1.5 mm. lata; interiora angustiora.

Type specimen: Collected by Ynes Mexia, No. 445, common at

altitude of 1,200 meters, growing in masses, in damp places in

openings in oak and pine forests, trail from El Batel to Pica de

Aguila, Sierra Madre, State of Sinaloa, Mexico, November 14, 1925

(Gray).

Distribution: Known only from type locality in State of Sinaloa,

Mexico.

Specimens examined: Mexia 445 (type, Gray; cotype, Mo.).

50. Coreopsis nuecensis Heller, Bot. Expl. S. Texas 106. 1895.

Coreopsis coronata Hook. Bot. Mag. pi. 3460. 1836 (nee alior.).

REVISION OF THE GENUS COREOPSIS 359

Herba annua, sparsim hirsuta vel fere glabra, 2-6 dm. alta,

caulibus erectis sed debilibus, basim versus ramosis, superne nudis.

Folia opposita jugis saepe remotis, summa subsessilia, alia petiolata

petiolis latis angustissimisve hispido-ciliatis (pilis pluriarticulatis)

usque ad 5 (quidem ad 12) cm. longis, petiolo adjecto 4-10 (-20)

cm. longa, lamina nunc late ovalia vel oblongo-spathulata integra,

nunc pinnatim secta segmentis usque ad 5, ultimo multo maximo,

lateralibus saepius alternis raro lobatis imis saepe petiolulatis.

Capitula ramos nudos saepe 2-3 dm. longos (pedunculos) terminan-

tia, radiata, pansa ad anthesin circ. 3.5-4.8 cm. lata et ± 1 cm. alta.

Involucri bracteae exteriores circ. 8, oblongo-lanceolatae vel anguste

deltoideo-ovatae vel subulatae, tergo glabratae, apice angusto

saepius rotundatae, lateribus saepe diaphanae, marginibus ciliatae

setis pluriloculatis, 5-8 mm. longae, quam interiores ovatae tergo

glabrae vel albido-setosae apice subacutae paulo breviores. Flores

ligulati circ. 8, ligula cuneato-ovati, basim versus maculis magnis

atro-sanguineis vel brunneo-rubris ornati aliter flavi, apice 3-lobati

lobo mediano multo majore plerumque in 2 lobulos hos saepe emar-

ginatos rursus diviso, 1.5-2 cm. longi. Paleae lineares, supra demum

angustissimae ac coloratae, 3-6 mm. longae. Flores tubulosi nunc

flavidi nunc superne purpurascentes, stylorum ramis crasso-truncatis

minute ac abrupte mucronatis. Achaenia obcompressa, circumam-

bitu alis inclusis obovato-orbiculata circ. 4 mm. longa et circ. 4 mm.

lata, dorso (convexo brunneo-nigro) spinulis saepe capitatis dense

ventre (concavo utrinque plerumque perspicue calloso) sparsim

munita, alata alis membranaceis brunneis irregulariter incisis vel

lobatis circ. 1 mm. latis, apice 2 squamellis vel spinulis usque ad 0.4

mm. longis munita.

Type specimen : Raised from seed collected by Thomas Drummond

in Texas, about 1835.

Distribution: Southeastern Texas.

Specimens examined: Berlandier 1802, from Bejar (Bexar) to

Austin, Texas, April, 1828 (Gray). A. H. Curtiss 6362, waste ground,

escaped from cultivation, St. Augustine, Florida, April 13-May 18,

1898 (Gray; Mo.); Thomas Drummond 199, Texas (Gray); Elihu

Hall, cult, e sem. texanis, Athens, Illinois, 1864 (U.S.); Hort.

Cantabr., cult, anno 1848 e sem. texanis a Lindheimero lectis (Gray,

2 sheets); idem, cult., September, 1856 (Mo.); Marcus E. Jones

29511, Kingsville, Texas, March 29, 1932 (Pom.); idem 29512,

Encino, Texas, March 28, 1932 (Pom.};F.Lindheimer, prairies west

of the Brazos River, Texas, April (Mo.); idem 363, Guadeloupe

360 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

bottoms at Victoria, Texas, February, 1845 (Mo.); C. Mohr, cult,

e sem. texanis, Mobile, Alabama, May 10, 1887 (U.S.); H. Ness

2078, moist forest lands, Brazos County, Texas, April, 1900 (Mo.,

2 sheets) ;#. J. Palmer 9144, sandy, open ground, Inez, Texas, March

11, 1916 (Mo.); Earl E. Sherff 5017, cult, in Harvard Univ. Bot.

Gard. (Hort. Cantabr.), August 12, 1929 (Field; Kew);5. C. Tharp

5635, Woodboro, Texas, March 17, 1929 (U.S.); Tracy 8925 p.p.,

Victoria, Texas, April 27, 1905 (Mo.).

Heller's name C. nuecensis goes back by direct synonymy to the

name C. coronata Hook., which it supersedes because of the homonym

C. coronata L. Heller's cited specimen (A. A. Heller 1548, Corpus

Christi, Texas, April 11, 1894), however, was very different from

C. coronata Hook, and was, indeed, at the time an undescribed species.

It was later named C. similis Boynt. (qu. vide).

51. Coreopsis cordylocarpa A. Gray, Proc. Amer. Acad. 22:

428. 1887.

Perennis, fruticosa, multicaulis, foliosa, 1.5-1.8 meters alta,

brevissime adpresso-hispida, caulibus subtetragonis. Folia tenuiter

petiolata petiolis 1-3.5 cm. longis, petiolo adjecto plerumque 0.7-2

dm. longa, submembranacea, pinnatim 3-5-partita foliolis simplici-

bus vel rursus 2-3-fidis, segmentis ultimis linearibus vel anguste

oblongo-lanceolatis saepius 3-11 mm. latis, margine indurato-cras-

siusculis et apicem acutum subacuminatumve versus acriter (inter-

dum inciso-) dentatis. Capitula subcymosa pedunculata pedunculis

tenuibus folia paulo vel multo superantia, radiata, pansa ad anthesin

3-4 cm. lata et circ. 1 cm. alta. Involucri plus minusve hispidi

bracteae subulatae, exteriores lineares 4-6 mm. longae, quam

interiores lanceolatae saepe paulo longiores. Flores ligulati circ.

10, lutei, ligula oblongi, apice integri vel denticulati, 1.5-2.2 cm.

longi. Paleae oblongo-lineares, glabrae, apice acutae, demum 5-7

mm. longae. Florum tubulosorum stylorum rami apice conico-

angustata. Achaenia obcompresso-clavata, glabra, exalata, brun-

neo-nigra, obscure longitudinaliterque striata, circ. 8-12 mm.

longa et ± 1.5 mm. lata, apice contracto disco parvo calvo terminata.

Type specimen: Collected by Edward Palmer, No. 172, at Rio

Blanco, State of Jalisco, Mexico, July, 1886 (U.S.).

Distribution : Known only from State of Jalisco, Mexico.

Specimens examined: C. R. Barnes & W. J. G. Land 155, alt.

1,560 meters, bank of stream, Sierra de San Esteban, State of Jalisco,

September 28, 1908 (Field, 2 sheets); Palmer 172 (type, U.S.;

REVISION OF THE GENUS COREOPSIS 361

cotypes, Gray; N.Y.); C.G. Pringle 2367, river banks, near Guada-

lajara, State of Jalisco, October 7, 1889 (Field; Gray; N.Y.); idem

9885, alt. 1,500 meters, banks of Rio Blanco, near Guadalajara, July

31, 1902 (Berl.; Field; Gray; N.Y.); idem 11506, eodem loco, October

20, 1903 (Field; Gray).

52. Coreopsis elgonensis Sherff, Bot. Gaz. 80: 374. 1925.

Planta erecta, perennis, ramosa, 8-10 dm. alta; ramis angulatis,

glabris, infra ligneis, internodiis numerosis et saepe tantum 3-10 mm.

longis. Folia opposita, numerosa, sessilia, tantum circ. 1-1.8 cm.

longa, ternatim divisa, foliolis membranaceis, margine ciliatis,

faciebus atro-punctatis et interdum sparsissime hispidis, cuneatis,

ternatim lobatis vel integris, segmentis ultimis acerrime apiculatis,

plerumque 2-3 mm. latis. Capitula pauca in corymbis disposita

pedunculis 2-9 cm. longis, radiata, pansa ad anthesin 2.5-3.5 cm.

lata et 8-11 mm. alta. Involucri bracteae exteriores 6-11, lineares,

supra latiores, apice acutae, plerumque glabratae, 5-9 mm. longae;

interiores lanceolatae, hispidae, paulo breviores. Flores ligulati

8-12, flavi, ligula anguste elliptico-oblanceolati, apice rotundati

sed minute plus minusve denticulati, 8-12 mm. longi et 3-4.5 mm.

lati. Paleae oblongo-lanceolatae, brunneae, pluristriatae, apice

saepe longo-attenuatae, circ. 6-7.5 mm. longae. Disci florum

stigmata caudato-attenuata. Achaenia late lineari-oblonga, plana,

omnino atra, glabra, exaristata, 3.5-4.8 mm. longa et 1.1-1.3 mm.

lata, marginibus non vere membranaceis.

Type specimen: Collected by R. A. Dummer, No. 3304, locally

frequent, at altitude of 3,900 meters, in thicket at edge of cliff, west

side of crater, Mount Elgon, Uganda, British East Africa, January,

1918 (Kew).

Distribution: Known only from type locality.

Specimens examined: Dummer 3304 (type, Kew).

Strongly suggestive, in general habit, of such South American

species as Coreopsis polyactis and C. senaria. The many small,

sessile, cuneate, ternate or biternate leaves appear at first glance to

be in whorls rather than in pairs. Correlated with the leaf abun-

dance is the shortness of the internodes. Thus, for example, one

branch of the type is seen to have 28 internodes in a length of 2.5

dm., giving an average length of only about 9 mm.

53. Coreopsis Chippii M. B. Moss, Kew Bull. 196. 1929.

Suffrutex, 1 m. altus, caulibus (glabratis) et ramulis (puberulis)

subteretibus, internodiis 5-12 mm. longis. Folia opposita, tenuiter

362 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

petiolata petiolis circ. 4-6 mm. longis, petiolo adjecto circ. 1.5-2 cm.

longa, 1-2-pinnatisecta, segmentis glabris vel glabriusculis, acriter

apiculatis, crassiusculis, tantum circ. 0.2-0.4 mm. latis. Capitula

subcorymbosa, tenuiter pedicellata pedicellis valde puberulis et

circ. 1-2.5 cm. longis, radiata, pansa ad anthesin ± 2.3 cm. lata et

5-6.5 mm. alta, demum subglobosa et circ. 1 cm. diametro. Invo-

lucri hispidi bracteae exteriores 10-20, lineares, obtusae vel etiam

subtruncatae, 4-5 mm. longae et circ. 0.5 mm. latae, saepe patentes;

interiores late lanceolatae paulo longiores. Flores ligulati (pro-

babiliter circ. 8), flavi, ligula anguste obovati, apice 3-denticulati,

circ. 1 cm. longi. Paleae oblongae, obtusae, glabrae, nitidae, superne

atriores, 4.5-5.5 mm. longae. Disci florum stigmata truncata,

minute appendiculata. Achaenia valde obcompressa, dorso convexa,

anguste obovato-oblonga, nigra, glaberrima, apice truncate exaris-

tata, ad latera crassiusculo-marginata sed non vere alata, circ. 4 mm.

longa et 1-1.4 mm. lata.

Type specimen : Collected by Thomas Ford Chipp, No. 66, grow-

ing in scrub on mountain top, also in forest in ravines, at altitude of

3,125 meters, top of Mount Kinetti (Kineti), Imatong Mountains,

southern Sudan, February 11, 1929 (Kew).

Distribution: Known only from type locality in southern Sudan.

("The Imatong Mountains, the highest point of which is Mt. Kineti,

10,414 feet, are situated between 4° and 4° 30' N. and 32° 20' and

33° E., just north of the boundary between the Anglo-Sudan and

Uganda"; T. F. Chipp, Kew Bull. 177. 1929.)

Specimens examined: Chipp 66 (type, Kew).

54. Coreopsis scopulorum Sherff, Bot. Gaz. 88: 302. 1929.

Frutex +2 dm. altus, glaber, ramosus, ramis foliosissimis, inter-

nodiis saepe tantum 3-5 mm. longis. Folia opposita, petiolata

petiolis 6-10 mm. longis, petiolo adjecto tantum 1.2-2 cm. longa,

plus minusve biternatisecta, segmentis ultimis linearibus, subcarno-

sis, margine saepe revolutis, apice mucronatis, vix 1 mm. latis.

Capitula tenuiter pedunculata pedunculis 3-5 cm. longis ac saepe

ad ramorum terminos 3-5-adgregatis, radiata (radiis in typo defi-

cientibus sed ovariis circ. 8 sterilibus repraesentatis), demum circ.

8 mm. alta et 7-10 mm. lata. Involucri bracteae exteriores circ.

8, anguste lineares, patenti-reflexae, apice rotundatae vel abrupte

cuspidatae, circ. 3 mm. longae, interioribus lanceolatis atque apicem

versus reflexis subaequalibus. Paleae perspicuae, late oblongae,

apice rotundato-obtusae, achaeniorum corpora moderate superantes.

REVISION OF THE GENUS COREOPSIS 363

Disci florum stigmata apice perspicue caudata. Achaenia linearia,

obcompressa, exalata, faciebus marginibusque erecto-setosa, atra,

singula facie circ. 8-striata, corpore 5-6 mm. longa et 0.7-1 mm.

lata, apice erecto-setosa ac perspicue biaristata aristis tenuibus

erecto-hispidis circ. 2.5 mm. longis.

Type specimen: Collected by Edmund Heller, on cliffs at altitude

of 2,130 meters, on summit of Mount Garguez, British East Africa,

August 26, 1911 (U.S.).

Distribution: Known only from type locality.

Specimens examined: Heller, on cliffs at alt. 2,130 meters, summit

of Mount Garguez, British East Africa, August 26, 1911 (type, U.S.).

Apparently nearest Coreopsis elgonensis (from which it differs in

its longer and petiolate leaves, its shorter external involucral bracts, its

aristate achenes, etc.) and C. Chippii (from which it differs in its much

broader leaf segments, etc.). All three species offer a strong resem-

blance to certain low, shrubby South American species of Coreopsis.

55. Coreopsis Jacksonii S. Moore, Journ. Linn. Soc. 35:

347. 1902. Plate 1. Bidens spathulata Sherff, Bot. Gaz. 76: 149,

pi. 13. 1923. B. Jacksonii (S. Moore) Sherff, Bot. Gaz. 81: 45. 1926.

Folia plus minusve glabra C. Jacksonii.

Folia pro parte maxima dense hispida var. /3 arthrochaeta.

Herba procumbens vel adscendens, plus minusve glabra, forsitan

annua, caule subsimplici nonnullis nodis radicante, 3.5-5 dm.

longo. Folia non manifeste petiolata, 4-8 cm. longa et 1.5-2.5 cm.

lata, valde membranacea, spathulato-oblanceolata, apice subobtusa,

inferne sensim angustata, integra vel remotissime minimeque denti-

culata, ciliata, in sicco plus minusve stramineo-viridia. Capitula

perpauca (1-3), pedunculata pedunculis pubescentibus usque ad

6 cm. longis, radiata, pansa ad anthesin circ. 3.5-4.5 cm. lata et

7-9 mm. alta. Involucri bracteae exteriores circ. 6, membranaceae,

oblongae vel oblanceolatae, ciliatae, apice orbiculatae, ad anthesin

circ. 8-10 mm. longae et 2-3 mm. latae; interiores oblongo-ovatae,

apice subacutae, ad anthesin circ. 3.5-4.5 mm. longae. Flores

ligulati circ. 6, lutei, ligula elliptico-oblanceolati, apice dentati,

1.5-1.8 cm. longi et 4.5-6.5 mm. lati. Florum tubulosorum stigmata

termino anguste elongata. Achaenia subatra vel brunneo-atra,

crasso-clavata, plus minusve tumida ac quadrangulata, non mani-

feste striata, circ. 3.2-3.7 mm. longa et 1.1-1.7 mm. crassa, glabrata,

exalata, exaristata sed apice saepe reliquo basali floris hermaphroditi

coronata itaque false rostrata.

364 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: Collected by Frederick J. Jackson, Kikuyu

region, British East Africa, 1889 (Brit.).

Distribution: Kikuyu region and northward to Mount Kenya

(Kenia), British East Africa.

Specimens examined: Frederick J. Jackson, Kikuyu region, 1889

(type, Brit.); Edgar A. Mearns 1291, alt. about 3,630 meters, in

"giant heath" zone, western slopes of Mount Kenya, British East

Africa, September 21-27, 1909 (type of Bidens spathulata Sherff,

U.S.); idem 1722, alt. about 3,000 meters, bamboo zone, western

slopes of Mount Kenya, September 28-October 7, 1909 (U.S.).

A species strikingly different from all other species of Coreopsis

and easily distinguished from other African species by its thin,

spatulate leaves. Its habit of rooting at many of the nodes,

evidently in response to aqueous conditions of the habitat, is almost

as distinctive, being common in only a few species of these two genera.

My examination of the type specimen of Coreopsis Jacksonii S.

Moore (Brit.) showed that it was merely a tiny, dwarfed specimen

(cf. Moore, loc. cit., "unfortunately a mere scrap") of the species

described and illustrated by me under the name of Bidens spathulata.

(The type of Bidens spathulata, it will be noted, was collected upon

Mount Kenya, British East Africa, while the type of Coreopsis Jack-

sonii was collected in the small Kikuyu area immediately to the south-

west.) Because, however, of its very diminutive stature (being only

a few centimeters high), it lacked the spatulate type of leaves that

so strongly characterizes larger plants. The leaves were more

rotund in outline. For these reasons, Moore's description of Co-

reopsis Jacksonii had been, of necessity, so misleading that only a

personal study of his type plant could impart an understanding of

its true specific status. As to its generic status, its complete lack

of mature achenes, as also in the case of the type of Bidens spathulata,

made it necessary to rely upon general habit, which is somewhat

more that of Bidens than of Coreopsis. The recent finding (1) of the

var. arthrochaeta of this species and (2) of Mearns 1 722 of the species

proper makes it possible, however, to refer the species definitely to

Coreopsis.

Coreopsis Jacksonii var. $ arthrochaeta Sherff, Bot. Gaz.

88: 302. 1929.

A specie foliis pro parte maxima dense hispidis, setis albidis

elongatis sensim attenuatis multiloculatis differt.

FIG. 1. Coreopsis Jacksonii (S. L. Moore) Sherff.

365

366 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: Collected by R. L. Piemeisel and Leonard W.

Kephart, No. 166, at altitude of 3,300 meters, vicinity of Camp

Gusisu, Aberdares Mountains, British East Africa, July 29, 1927

(U.S.).

Distribution: Known only from type locality.

Specimens examined: Piemeisel & Kephart 166 (type, U.S.).

The type material of the var. arthrochaeta consisted of at least

six small, entire plants and these offer a strikingly unique appearance

because of their densely hispid leaves. Since endemism is pronounced

among the plants of the Kikuyu region, and since elsewhere even

where endemism is relatively unimportant (in the United States)

Coreopsis displays many forms that are accepted as varieties, it

appears worth while to distinguish the Piemeisel & Kephart plants as

connoting a group of varietal rank.

Explanation of Fig. 1. — Coreopsis Jacksonii S. L. Moore: a,

flowering specimen, X0.66; b, portion of leaf enlarged to show cilia-

tion, X2.65; c, exterior involucral bract, X2.65; d, interior involucral

bract, X2.65; e, ligulate floret, X2.65;/, palea, X2.65; g, disc floret,

X2.65; all from Mearns 1291, type of Bidens spathulata Sherff,

Herb. U. S.

56. Coreopsis Negriana Sherff, Bot. Gaz. 90: 397. 1930.

Herba annua, erecta, ±5 dm. alta, caule gracili, glabro vel ad

summam subhispido, viridi-stramineo. Folia opposita petiolata

petiolis usque ad 12 mm. longis saepe tenuissimis petiolo adjecto

±5 cm. longa, 1-2-pinnatisecta, segmentis principalibus lineari-

oblongis, membranaceis, supra infraque glabratis, marginibus ciliatis,

acriter serratis dentibus interdum in setas elongatas desinentibus.

Capitula graciliter pedunculata pedunculis moderate hispidis usque

ad 14 cm. longis, radiata, pansa ad anthesin 2-2.5 cm. lata et circ.

6-7 mm. alta. Involucri hispidi bracteae subaequales exteriores circ.

8, lineari-spathulatae, acriter indurato-apiculatae, 3-4 mm. longae;

interiores oblongo-ovatae, apicaliter saepe acutae. Flores ligulati

circ. 8, flavi, ligula lineari-elliptici, apice obtuso obscure denticulati,

circ. 1 cm. longi et circ. 2-3 mm. lati. Paleae nitido-hyalinae,

lineari-oblongae vel lineari-lanceolatae, demum conduplicatae ac

achaenium amplectentes, maturae circ. 6 mm. longae. Disci florum

stigmata subito capilliformi-caudata. Achaenia lineari-oblonga,

valde obcompressa, glabra, corpore nigro estriato vel obscurissime

multistriato tantum 3-4 mm. longa et circ. 1 mm. lata, marginibus

anguste alata alis brunneis glabrisque, apice exaristata ac glabra.

REVISION OF THE GENUS COREOPSIS 367

Type specimen: Collected by Giovanni Negri, No. 915 bis, at

altitude of 1,500 meters, Arussi Galla, June 30, 1909 (Flor.).

Distribution: Known only from type locality in Arussi Galla,

Gallaland.

Specimens examined: Negri 915 bis (type, Flor.).

A species close to C. pachyloma 0. & H., from which it differs,

however, in its smaller capitula, also its smaller achenes which are

obcompressed, winged, and glabrous. The general habit is deceivr

ingly like that oiBidens setigera (Coreopsis setigera Schz. Bip.), some

of the foliar teeth even having the setiform extensions so notable in

that species. But B. setigera's achenes are distinctly striate, up-

wardly setose upon the faces and edges, entirely exalate, and (in

the species proper) at the apex conspicuously slender-aristate.

57. Coreopsis camporum Hutch. Kew Bull. 381. 1921.

Herba perennis, erecta, caule 1-1.25 m. alto, basim versus

dense foliato internodiis brevibus, glabro, simplici vel superne corym-

boso-ramoso. Folia opposita, petiolata petiolis anguste marginatis

circ. 1-2.5 cm. longis, petiolo adjecto 0.5-1.5 dm. longa, simpliciter

vel rarius bipinnata, sicca laete viridia, segmentis linearibus vel

lineari-lanceolatis, acutis, revolutis, unaquaque facierum glabris,

margine spinuloso-setulosis et saepe sparsim dentatis (dentibus e

basibus triangularibus et longe setigeris), usque ad 9.5 cm. longis

et 7 mm. latis. Capitula pauca vel numerosa, subcorymbosa,

breviter tenuiterque pedunculata pedunculis superne puberulis,

radiata, pansa ad anthesin 2-3 cm. lata et 6-9 mm. alta. Involucri

bracteae exteriores circ. 12, patulae, lineares, acutae, tergo glabratae

sed marginibus setulosae, 0.8-1.5 cm. longae; interiores adscendentes,

oblongo-lanceolatae, obtuse acuminatae, circ. 6 mm. longae et 2

mm. latae, crebre resinoso-striatae. Flores ligulati circ. 10, auran-

tiaci, fertiles, ligula oblongo-elliptici, apice breviter bilobati, circ.

1.2-1.5 cm. longi et circ. 4 mm. lati, circ. 9-nervii. Paleae oblongo-

oblanceolatae, minute mucronatae, circ. 5.5 mm. longae et circ.

1.5 mm. latae. Achaenia immatura lineari-oblonga, complanata,

omnino brunnea, alata, ciliato-pubescentia, corpore 4-5 mm. longa,

biaristata aristis rigidis circ. 2.5 mm. longis.

Type specimen : Collected by H. V. Lely, No. 383, at altitude of

1,380 meters on plains between Hepham and Ropp, northern Nigeria,

July 5, 1921 (Kew).

Distribution: Northern Nigeria.

368 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

Specimens examined : Lely 383 (type, Kew); idem 449, alt. 1,380

meters, Ropp, July 19, 1921 (Kew); J. Dent Young 153, alt. 900-

1,350 meters, waste land, Vom, Bauchi Plateau, 1922 (Kew).

58. Coreopsis Ellenbeckii 0. Hoffm. Bot. Jahrb. 38: 205. 1906.

Herba perennis, glaberrima. Folia (caulium juvenilium) petiolata

petiolis alato-marginatis basi connatis 5-10 mm. longis, petiolo

adjecto 3-5 cm. longa, bipinnatipartita, rhachi late linearia ac alata

segmentis linearibus acutis apicaliter callosis 1-2 mm. latis. Capi-

tula (tantum unicum terminantem ramum juvenilem vidi) tenuiter

pedunculata pedunculo glabro fere 3.5 cm. longo, radiata, pansa ad

anthesin (bracteis patentibus inclusis) circ. 4.5 cm. lata et circ. 11

mm. alta. Involucri bracteae exteriores circ. 5, perspicue foliaceae,

trifidae, glaberrimae, apicaliter callosae, 1.7-2.4 cm. longae, lobo

terminali multo majore ±1.5 mm. lato et usque ad 1.5 cm. longo;

interiores oblongae membranaceae brunneae margine angusto

luteolo cinctae circ. 7-8 mm. longae. Flores ligulati 8, flavi, ligula

elliptico-oblongi, multistriati, ±1.8 cm. longi et ±4 mm. lati, apice

acriter 3-dentati dente mediano acriter bidenticulato. Disci florum

stigmata terminaliter caudata. Achaenia matura non visa; ovariis

obcompressis, margine hyalino cinctis faciebus glaberrimis sed

marginibus dense erecto-ciliatis ciliis superioribus elongato-sericeis,

biaristatis aristis corolla brevioribus rigide erecto-hispidis.

Type specimen: Collected by Dr. Elknbeck (Exped. Baron von

Erlanger), No. 529, at altitude of 2,500 meters in meadows on moun-

tain slopes, on the Gara Mulata, Harar, State of Harar, eastern

Abyssinia, March 21, 1900 (Berl.).

Distribution: Known only from type locality at Harar, eastern

Abyssinia.

Specimens examined : Ellenbeck 529 (type, Berl.).

Founded upon the lone type specimen, which consists of four

leafy stems 13-20 cm. long, one of them ending in a flowering head.

These come from the base of an old stem some 6 mm. thick

which had died away. The juvenile or second-growth nature of the

parts relied upon for the technical description leave some doubt

as to the degree to which the characters recorded are truly

representative.

59. Coreopsis lineariloba 0. Hoffm. Bot. Jahrb. 30: 430. 1901.

Herba perennis, ramosa, ramis glabris, erectis, internodiis saepius

9-13 cm. longis. Folia opposita petiolata petiolis planis ac anguste

marginatis usque ad 3 cm. longis, petiolo adjecto usque ad 12 cm.

REVISION OF THE GENUS COREOPSIS 369

longa, bipinnatipartita segmentis rhachi subsimilibus saepius 1-1.5

cm. longis et plerumque 1.2-2.5 mm. latis linearibus apice acuto

mucronatis glabris etiam eciliatis. Capitula mediocria tenuiter

pedicellata pedicellis glabris cymam laxam bracteis linearibus

munitam formantia, radiata, pansa ad anthesin circ. 2 cm. lata et

7-9 mm. alta. Involucri bracteae exteriores 10-12, lineares, apice

obtusae acutaeve, basi ciliatae ceterum glabrae, 4-6 mm. longae;

interiores ovatae, acuminatae, apice tomentellae ceterum glabrae,

saepe paulo breviores. Flores ligulati circ. 8-10, flavi, ligula ovati,

apice subintegri, circ. 1 cm. longi. Paleae late oblongae, apice

subacutae. Disci florum stigmata terminaliter caudato-attenuata.

Achaenia matura ignota; ovariis obcompressis glabris vel saepius

margine et inter aristas pappi ciliatis, pappi aristis 2 (raro 3)

subulatis tubum corollae superantibus antrorsum longe pectinato-

ciliatis.

Type specimen: Collected by W. Goetze, No. 790, growing 1

meter high on red laterite at altitude of 1,900 meters on an undulating

plateau, Liangira, Ubena, southwesternmost German East Africa,

March 24, 1899 (Berl.).

Distribution: Known only from type locality in German East

Africa.

Specimens examined: Goetze 790 (type, Berl.).

60. Coreopsis Schimperi 0. Hoffm. Bot. Jahrb. 38: 205. 1906.

Microlecane abyssinica f. elongata Vatke, Linnaea 39: 497. 1875.

Herba perennis, glaberrima, e radice fasciculata radiculis ±1 dm.

longis et circ. 2.5 mm. crassis, erecta vel basim versus procumbens

et nodis radicans, caule gracili subsimplici 6-9 dm. alto. Folia

opposita petiolata petiolis brevibus basaliter dilatatis connatisque

tantum circ. 2-7 mm. longis, petiolo adjecto usque ad 5.5 cm. longa,

quam internodia multo brevioria, circumambitu late deltoideo-

ovata, 2-3-pinnatipartita segmentis linearibus acutis apice callosis

margine raro spinuloso-ciliatis circ. 1 (raro 2) mm. latis. Capitula

in corymbo amplo laxiusculo foliis reductis bracteato disposita

pedicellis gracilibus spinuloso-hispidis vel glabratis apice paululo

dilatatis usque ad 4.5 cm. longis, radiata, pansa ad anthesin 2-2.8 cm.

lata et tantum circ. 5 mm. alta. Involucri bracteae exteriores circ.

8 (raro usque ad 10), anguste lineares, subherbaceae, glaberrimae,

subacriter calloso-apiculatae, 3-3.5 mm. longae et circ. 0.4 mm.

latae; interiores late ellipticae, medio membranaceae purpurascentes

pilos breves crassos gerentes, margine hyalina glaberrimae, apice

370 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

in acumen breve obtusum productae, quam exteriores interdum

paulo longiores. Flores ligulati circ. 8, flavi, ligula oblongo-elliptici,

apice plerumque 3-lobulati, 1-1.5 cm. longi et 4-7 mm. lati. Paleae

hyalinae, nitidae, late lineari-oblongae, apice subobtusae, lineis 3

duplicibus percursae circ. 4-5 mm. longae. Disci florum stigmata

minima terminaliter caudato-angustata. Achaenia leviter obcom-

pressa, nitida, crasso-clavata, glaberrima, aegre striata, exalata,

exaristata, circ. 2.5 mm. longa et fere 1 mm. lata.

Type specimen: Collected by Wilhelm Schimper, No. 1386, in

meadows at altitude of 2,460 meters, Gaffat, Abyssinia, October 15,

1863 (Berl., 2 sheets).

Distribution : Abyssinia.

Specimens examined: Rohlfs & Stecker, Godofelassi, Abyssinia,

January 7, 1881 (Berl.); Schimper 1386 (2 type sheets, Berl.; type

and cotype of Microlecane abyssinica f. elongata Vatke).

The type material had been incorrectly ranked by Vatke as a

forma under Microlecane abyssinica (Schz. Bip.) B. & H. f. (Bidens

setigera [Schz. Bip.] Sherff). As 0. Hoffmann (loc. cit.) has shown,

the two are very different.

61. Coreopsis Prestinariaeformis Vatke, Linnaea 39: 499.

1875. C. Pristinariaeformis Vatke ex 0. & H. in Oliv. Fl. Trop. Afr.

3: 387. 1877 (sphalm). C. heterocarpa Chiov., Ann. Bot. Roma 9:

75. 1911 (pro maxima parte; cf. etiam var. incisa infra).

Folia simplicia vel pinnatim 3-5-tripartita.

C. Prestinariaeformis sensu stricto.

Folia bipinnatisecta var. 0 incisa.

Herba annua, undique plus minusve pilis crassis flexuosis obsita,

caule simplex vel quidem e basi valde ramosa, 1-6 dm. alta. Folia

sessilia vel breviter petiolata, 2.5-7 cm. longa, circumambitu ovato-

lanceolata vel deltoideo-cordata, sicca subvalde membranacea,

simplicia vel plurimum subincise 3- vel etiam sub 5-partita, segmentis

varie ovatis vel rhomboideo-ovatis et margine dentatis vel incisis

lateralibus multo minoribus. Capitula saepius numerosa vel numero-

sissima, tenuiter pedicellata pedicellis ante et per anthesin erectis

postea hamato-recurvis et tune cernua, radiata, pansa ad anthesin

circ. 2.8-3.3 cm. lata et circ. 1-1.2 cm. alta. Involucri bracteae

aequilongae, exteriores 10-12, patulae, lineares vel lineari-spathulatae,

apice obtusae vel acutiusculae, glabrae, circ. 4-6 mm. longae;

interiores ovato-lanceolatae vel oblongo-ovatae, rufidulae, in dorso

notatae ibique pilis longis crassis flexuosis flavescentibus onustae.

REVISION OF THE GENUS COREOPSIS 371

Flores ligulati 6 vel 7, aurei, ligula varie elliptico-obovati oblongo-

elliptici vel late spathulato-oblanceolati, apice denticulati, superne

in centre vel parum subtus macula aurantiaca rotunda elliptica vel

lunulata praediti, 1.5-2.5 cm. longi et usque ad 8 mm. lati. Paleae

glaberrimae, supra rufae. Achaenia valde obcompressa apice erecto-

setosa in 3 vel 4 seriebus exteriora corpore ±8 mm. longa et

±2 mm. lata, marginibus latissime alata alis apice truncatis et

setoso-ciliatis; centralia lineari-oblonga exalata setoso-ciliata corpore

±12 mm. longa et ±2.5-3 mm. lata; omnia cum pappo duobus

aristis linearibus vel demum lanceolato-subulatis acutissimis achaenii

corpori aequilongis vel quidem tantum 3-4 mm. longis antrorsum

hispidis formato.

Type specimen: Collected by Wilhelm Schimper, No. 1173, at

altitude of 2,460 meters in meadows at Gaffat, Abyssinia, September

25, 1863 (Berl., 2 sheets).

Distribution: Abyssinia.

Specimens examined: Emilio Chiovenda 970, Debarek, District

of Semien, Province of Amhara, Abyssinia, July 11, 1909 (Flor.);

idem 1148, Gondar, District of Dembia (Dembea), Province of Am-

hara, Abyssinia, July 30, 1909 (Flor.); idem 1869, vicinity of Gondar,

September 8, 1909 (Flor.); idem 1953 et 1954, District of Dembia,

September 8, 1909 (Flor.); G. W. Grabham, Bahadur Gorges, on

Lake Tana (L. Tsana), Abyssinia (Kew); Plowden, Abyssinia (Kew);

Schimper 1173 (type, Berl., 2 sheets; cotype, Kew).

An important plant in the festal celebrations of some of the

Abyssinians (cf. Chiovenda, loc. cit.).

Chiovenda (loc. cit.) cited seven of his own specimens as a basis

for his Coreopsis heterocarpa: Nos. 1090, 1148, 1869, 1953, 1954, 1955,

and 970 (in the order here given). His No. 1090 is my var. incisa

(infra). The others, unless 1955 (which alone I have been unable

to find at Florence), are C. Prestinariaeformis proper. Chiovenda's

description as drawn fits primarily C. Prestinariaeformis.

Coreopsis Prestinariaeformis var. ft incisa Sherff, Bot. Gaz.

90: 387. 1930. C. heterocarpa Chiov. Ann. Bot. Roma 9: 75. 1911

(pro minima parte).

A specie foliis acriter et perspicue bipinnatisectis segmentis

principalibus oblongo-lanceolatis differt.

Type specimen: Collected by Dr.Ellenbeck (Expedition of Baron

von Erlanger], No. 1436, growing 1 meter high at altitude of 2,600

meters, Djafa, Arussi, Galla, Gallaland, July 21, 1900 (Berl.).

372 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Distribution: Northern Abyssinia southward into Gallaland.

Specimens examined: Emilio Chiovenda 1090, very common,

Gondar, District of Dembia (Dembea), Province of Amhara, Abys-

sinia, July 26, 1909 (Flor.); Ellenbeck 1436 (type, Berl.).

62. Coreopsis Barter! 0. & H. in Oliver, Fl. Trop. Afr. 3: 390.

1877. C. badia Sherff, Bot. Gaz. 76: 90. 1923.

Herba glabra, adscendens, ramosa ramis teretibus striatis, 3-10

dm. alta. Folia opposita vel rarius trina, sessilia vel subsessilia,

membranacea, 1.5-5 cm. longa, nunc cuneato-oblanceolata et apicem

versus subremote acriterque serrata, nunc irregulariter trilobata

lobis lineari-oblongis. Capitula ramos singillatim terminantia,

tenuiter pedunculata pedunculis 3-9 cm. longis, radiata, pansa ad

anthesin 2-2.6 cm. lata et 7-9 mm. alta. Involucri glabri bracteae

subaequales, exteriores 7-10, lineari-oblongae, acriter apiculatae,

demum 5-7 mm. longae et 0.6-0.8 mm. latae; interiores oblongo-

lanceolatae circ. 2.2-2.6 mm. latae. Flores ligulati circ. 8, lutei,

ligula lineari-elliptici, apice obscure denticulati, 7-10 mm. longi et

3-4 mm. lati. Paleae late oblongo-lineares, apice obtusissimae, circ.

7 mm. longae. Disci florum stigmata parce incrassata, hispida,

caudata. Achaenia obcompressa, atra, corpore tenuiter vel sublate

linearia, 5-9 mm. longa et 0.9-1.5 mm. lata, faciebus venuste adpres-

seque erecto-pilosa, marginibus (saltern achaenia exteriora) late et

perspicue membranaceo-alata alis erecto-ciliatis, apice erecto-his-

pido biaristata aristis tenuibus erecto-hispidis 2-4 mm. longis.—

Involucri bracteae interiores, paleae, achaeniorum alae badiae vel

parce purpureo-badiae.

Type specimen: Collected by Charles Barter, No. 870, chiefly in

cultivated ground, Borgu, Upper Guinea (Kew).

Distribution: Northwestern Africa, from Kamerun west to Togo

and northward to Borgu.

Specimens examined: W. B. Baikie, on the Niger River, West

Tropical Africa (Kew) ; Barter, Borgu, Upper Guinea (type, Kew) ;

Mission Gironcourt 256, on mountains of gneiss, Togo, 1908-1909

(type of Coreopsis badia Sherff, Berl.); C. Ledermann 1855, alt.

1,800 meters, swampy place on shore of stream, Babadju, Kamerun,

December 15, 1908 (Berl., 2 sheets).

63. Coreopsis monticola (Hook, f.) 0. & H. in Oliver, Fl.

Trop. Afr. 3: 390. 1877. Verbesina (Prestinaria) monticola Hook,

f. Journ. Linn. Soc. 7: 200. 1864.

REVISION OF THE GENUS COREOPSIS 373

Caulis demum plerumque glabratus. C. monticola.

Caulis laxe vel subdense pilosus var. ft pilosa.

Herba perennis, erecta, rigida, subsimplex, 3-11 dm. alta, caule

vix subtetragono, superne circ. 2.5-3.5 mm. crasso, demum plerum-

que glabrato. Folia opposita petiolata petiolis planis submar-

ginatis 0.6-2 cm. longis, petiolo adjecto circ. 4-5 cm. longa, cir-

cumambitu ovata, subcoriacea, margine et facie inferiore aegre

hispida vel mox glabrata, trisecta, foliolis 3-5-sectis, segmentis

ovatis vel lanceolatis vel anguste oblongis, infra pallidioribus,

dentibus acriter apiculatis. Capitula corymbosa, pedunculata

pedunculis tenuibus 2-8 cm. longis, radiata, pansa ad anthesin

4-5.5 cm. lata et 8-10 mm. alta. Involucri glabri bracteae exteriores

8 vel 9, lineares, acriter indurato-apiculatae, 8-9 mm. longae, quam

interiores oblongo-ovatae nitidae rubro-brunneae vix longiores.

Flores ligulati circ. 8 vel 9, aurantiaci, ligula lineares, circ. 13-15-

nervii, apice minute sed acriter denticulati, circ. 1.8-2.2 cm. longi et

4.5-6 mm. lati. Paleae lineari-oblongae, circ. 5-nerviae, apice

obtusissimae, circ. 9 mm. longae et 2 mm. latae. Disci florum stig-

mata lanceolato-caudata. Achaenia oblonga, obcompressa, nigra,

exalata vel subanguste alata alis stramineis, facie ventrali margini-

busque sursum setosa, corpore 3-5.2 mm. longa et alis inclusis 1.1-2

mm. lata, apice corona setarum erectarum coronata ac biaristata

aristis tenuibus stramineis sursum hispidis circ. 2.5-3 mm. longis.

Type specimen: Collected by Gustav Mann, No. 1922, at altitude

of 1,800-2,100 meters, Kamerun (Cameroon) Mountains, Kamerun,

November 6, 1862 (Kew).

Distribution: Kamerun.

Specimens examined: W. Kalbreyer 125, alt. 2,100-2,550 meters,

open plain, dry places, Kamerun Mountain, Kamerun (Berl.);

Luckhardt 626, Kamerun, December, 1912 (Berl.); Mann 1922

(type, Kew; cotypes, Berl.; Gray);/1. Mildbraed 3392, "half-shrub"

growing 7.5-10 dm. high, furrows between lava streams, lower Fuko

Plateau, Great Kamerun Mountain, Kamerun, June, 1908 (Berl.);

Mintz 40, alt. 2,400-2,800 meters, Great Kamerun Mountain,

October 29, 1910 (Berl.) ; Preuss, alt. 2,000-2,800 meters, Kamerun,

1891 (Berl.); idem 689, Buea, Kamerun, 1891 (Berl., 3 sheets; Gray);

Gust. Simon 3, on lava fields, eodem loco, December 26, 1909 (Berl.).

Coreopsis monticola var. ft pilosa Hutch. & Dalz. Fl. West

Trop. Afr. 2:143. 1931.

374 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

Herba usque ad 3 m. alta, caulibus ramisque laxe vel subdense

pilosis pilis multiloculatis, foliis membranaceis, petiolo adjecto 6-10

cm. longis, superne hispidis; involucri bracteis exterioribus oblongo-

linearibus vel lineari-lanceolatis 6-14 mm. longis et 1.5-3 mm. latis;

interioribus tergo hispidis.

Type specimen: Collected by Dr. Dunlap, No. 204, growing up to

10 feet high in ravine above timber line at altitude of about 2,100

meters, Kamerun Mountain, Kamerun, January 17, 1926 (Kew).

Distribution: Kamerun.

Specimens examined : Dunlap 204 (type, Kew) ; F. W. H. Migeod

211, grassland, alt. 2,250 meters, Kamerun Mountains, November

17, 1927 (Kew).

64. Coreopsis oblonga Sherff, Bot. Gaz. 76: 80, pi. 7. 1923.

Fig. 2.

Herba erecta, verisimiliter perennis ac 6-12 dm. alta; caule

obtuse tetragono, atro, apicem versus pubescenti, alibi glabro.

Folia petiolata petiolis hispido-ciliatis vel brevibus (usque ad 1.5

cm. longis) latisque vel longioribus (usque ad 3 cm.) et parce angustis,

petiolo adjecto 6-7.5 cm. longa, rigida et subcarnosa, pinnata, foliolis

acriter dentatis dentibus calloso-apiculatis, subprofunde incisis

sinibus angustis vel foliolis lateralibus infimis etiam tripartitis,

breviter hispido-ciliatis, setis acribus sparsissime vestitis, terminali

oblongo-lanceolato, lateralibus oblongo-ovatis. Capitula pauca,

breviter pedunculata pedunculis subrobustis ac hie illic foliolo-

bracteatis usque ad 6 cm. longis, radiata, pansa ad anthesin 2.5-3

cm. lata et 9-11 cm. alta. Involucrum setis validis acribusque his-

pidum; bracteis exterioribus 6-9, late lineari-spathulatis, apicem

versus glabratis, apice subobtusis, 8-11 mm. longis, eis capitulorum

juniorum discum manifeste superantibus; interioribus oblongo-

oblanceolatis vel oblongo-obovatis, apicem versus constricta, exteri-

oribus demum subaequalibus. Flores ligulati ± 7, aurantiaci, ligula

elliptico-lanceolati, apice late obtusi et integri denticulative, circ.

15 striis percursi, tubo adjecto 1.5-2.4 cm. longi, tubo circ. 5 mm.

longo. Paleae lineari-oblongae. Flores tubulosi limbo aurantiaci,

infra pilosi (ad medium etiam longe hirsuti), supra perpaucis pilis

interdum conspersi; stylorum ramis terminaliter elongato-attenuatis.

Achaenia matura ignota. Ovaria subnigra, oblongo-cuneata, alata,

plana, faciebus glabra, corpore 2-3 mm. longa et 0.7-1 mm. lata,

apice arrecte setosa et longe biaristata; aristis acribus, supra et ad

medium glabris, basim versus setulis arrectis 1-3 sparsim munitis,

2-2.5 mm. longis.

FIG. 2. Coreopsis oblonga Sherff.

375

376 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: Collected by Alex. Carson, No. 106, Lake Tan-

ganyika, East Equatorial Africa, September, 1893 (Kew).

Distribution: Known only from type locality in East Equatorial

Africa.

Specimens examined: Carson 106 (type, Kew).

A species approaching Coreopsis pinnatipartita 0. Hoffm., Bidens

Grantii (Oliv.) Sherff, and B. Holstii (0. Hoffm.) Sherff. The hairy,

aristate ovaries distinguish it easily from C. pinnatipartita, while

the definite wings of the ovaries distinguish it from B. Grantii and

B. Holstii.

Explanation of Fig. 2. — Coreopsis oblonga Sherff: a, main portion

of flowering specimen, X0.72; 6, lower leaf from a, X0.72; c, exterior

involucral bract, X4.33; d, interior involucral bract, X4.33; e,

ligulate floret, X2.16; /, palea, X4.33; g, disc floret with achene

immature, X4.33; all from type, Herb. Kew.

65. Coreopsis pinnatipartita 0. Hoffm. Bot. Jahrb. 30: 432.

1901.

Fruticosa, perennis, erecta, 1-3 m. alta, ramis erectis, tomentel-

lis, junioribus foliatis. Folia opposita petiolata petiolis anguste

marginatis usque ad 3 cm. longis, petiolo adjecto 9-16 cm. longa,

pinnatipartita, circumambitu ovata, segmentis ovatis vel lanceolatis,

acutis vel obtusis, irregulariter pinnatilobis vel plus minusve pro-

funde serratis, coriaceis, supra scaberulis subtus tomentellis. Capi-

tula in corymbum oligocephalum terminalem congesta, breviter

subvaldeque pedicellata pedicellis tomentellis, radiata, pansa ad

anthesin 5-6.5 cm. lata. Involucri bracteae aequilongae, ovatae,

obtusae, dorso tomentosae, mox reflexae, 8-15 mm. longae, exteriores

circ. 8-10, plus minusve biseriales. Flores ligulati circ. 10-12,

flavi, ligula elliptici vel elliptico-oblanceolati, apice plerumque

acriter denticulati, circ. 3 cm. longi et circ. 6-9 mm. lati. Paleae

perspicue 3-pluri-lineatae lineis brunneis vel brunneo-purpurascenti-

bus, apice obtusae, achaenia multo superantes. Disci florum

stigmata terminaliter attenuato-elongatae. Achaenia lineari-ob-

longa, valde obcompressa, glaberrima, nigra vel subbrunneo-nigra,

faciebus multistriata, alata alis margine exteriore stramineis ac

crassiusculis, 5-6.5 mm. longa et 1.5-1.8 mm. lata, apice exaristato

calloso-indurata.

Type specimen: Collected by W. Goetze, No. 1041, at altitude of

2,300 meters on slopes of Poroto Mountain, southwesternmost

German East Africa, June 17, 1899 (Berl., 2 sheets).

REVISION OF THE GENUS COREOPSIS 377

Distribution: Nyassaland to German East Africa.

Specimens examined : J. Buchanan 380, Nyassaland, 1891 (Berl.);

A. Engler 1266, growing 1 meter high at alt. 2,000-2,400 meters, in

high mountain shrub and eagle-fern formation, "Magamba oberhalb

Kwai," West Usambara, German East Africa, October 4, 1902

(Berl.) ; Goetze 1041 (2 type sheets, Berl.); Ad. Stolz 1306, growing 3

meters high at alt. 1,450 meters, Kyimbila, Nyassaland, May 23, 1912

(Berl., 2 sheets); A. Whyte, alt. 1,200-1,800 meters, Mount Zomba,

Nyassaland, December, 1896 (Berl.).

66. Coreopsis Whytei S. L. Moore, Journ. Linn. Soc. 35: 348.

1902.

Herba verisimiliter perennis elataque, caule erecto subtereti vel

obtuse angulato pubescenti mox glabrato, ad nodos tumido. Folia

opposita, principalia petiolata petiolis saepius 2-3 cm. longis petiolo

adjecto ±8-10 cm. longa, pinnatifida segmentis ovatis acutis den-

tato-lobulatis; summa sessilia lanceolata dentato-lobulata integrave;

omnia supra sparsissime minutissimeque hispida, subtus pubescentia.

Capitula pedunculata pedunculis pubescentibus circ. 2-4 cm. longis,

sine dubio radiata, post anthesin fere vel vere 3 cm. lata et ±1.3

cm. alta. Involucri 3-seriati bracteae extimae et intermediae

anguste obovatae obtusae adpresso-erecte hispidae pilis plurilocu-

latis ±1.2 cm. longae; intimae oblongae obtusae tenuiores et paulo

breviores. Flores ligulati non observati. Paleae oblongo-lanceolatae,

obtusissimae, fere 1 cm. longae. Achaenia valde obcompressa, sub-

cuneate lineari-oblonga, angustissime alata, brunneo-nigra, glabra,

faciebus striata, apice incrassato exaristata, circ. 6-7 mm. longa et

circ. 2 mm. lata.

Type specimen: Collected by Alexander Whyte, No. 35, on Mount

Milanji (Mount Mlange), southernmost Nyassaland, October, 1891

(Brit.).

Distribution: Known only from type locality in southernmost

Nyassaland.

Specimens examined: Whyte 35 (type, Brit.).

67. Coreopsis Lupulina 0. Hoffm. Bot. Jahrb. 30: 432. 1901.

Fruticosa, glaberrima, perennis, ramosa, 2-3 m. alta, ramis

angulatis terminaliter foliatis. Folia opposita tenuiter petiolata

petiolis basaliter connatis usque ad 4 cm. longis, petiolo adjecto

8-12 cm. longa, principalia bipinnatipartita segmentis oblongis ser-

ratis vel pinnatifidis apice acutis, superiora simpliciter pinnatipartita.

378 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Capitula saepius in corymbos densos oligocephalos congesta pedicel-

lis brevibus crassisque, radiata, pansa ad anthesin circ. 6 cm. lata et

1.5-1.8 cm. alta. Involucri bracteae demum valde reflexae, exteri-

ores circ. 7-9 crassiusculae oblongo-ovatae apice obtusae circ. 8

mm. longae et 4-6 mm. latae, interiores membranaceae ovatae vel

lanceolato-ovatae obtuse acuminatae apice minute fimbriatae circ.

12 mm. longae. Flores ligulati circ. 10, flavi, ligula elliptici, apice

circ. 3-denticulati, circ. 2.5 cm. longi et circ. 7-8 mm. lati. Paleae

perspicue brunneo-striatae, lineares, obtusae, corollas superantes,

exteriores apicem versus demum reflexae. Disci florum stigmata

apicaliter elongato-attenuata. Achaenia valde obcompressa, lineari-

oblanceolata, omnino atra, marginaliter alata alis quam corpore

paulo vel etiam dimidio angustioribus, glabra, faciebus multi-

striata 5.5-8 mm. longa et (alis inclusis) 1.7-2.5 mm. lata, apice

calloso-marginata et obtuse emarginata.

Type specimen: Collected by W. Goetze, No. 1069, at altitude of

2,400 meters on slopes of Beya Mountain, Usafua, south westernmost

German East Africa, June 27, 1899 (Berl., 2 sheets).

Distribution: Known only from type locality in German East

Africa.

Specimens examined : Goetze 1069 (2 type sheets, Berl.).

This species and C. pinnatipartita are distinguished from the

very similar Bidens Holstii (0. Hoffm.) Sherff by their winged

achenes.

68. Coreopsis Neumannii Sherff, Bot. Gaz. 76: 85. 1923.

Herba ±1.5 m. alta, caule maximam partem glabro, sulcato,

ramoso, internodiis folia facile superantibus. Folia subsessilia vel

petiolata petiolis usque ad 1 cm. longis, petiolo adjecto 2-4 cm. longa,

plerumque tripartita (inferiora non visa, pinnata ?) foliolis ovatis vel

interdum lanceolatis, dentatis vel lobulatis, pubescentibus, dentibus

acerrime apiculatis. Capitula corymbosa, 5-11 in unaquaque corym-

borum, radiata, pansa ad anthesin 2.2-2.7 cm. lata et circ. 6 mm.

alta, pedunculis tenuibus plerumque 1-3 cm. longis. Involucrum

inconspicue hispidum, bracteis exterioribus circ. 8, flavido-viridibus,

lineari-spathulatis, apice acutis, 3-7 mm. longis, quam interioribus

lanceolatis paulo brevioribus. Flores ligulati plerumque 8, flavi,

ligula elliptico-oblongi, paucistriati, apice integri vel denticulati,

circ. 1.2 cm. longi. Paleae oblongo-lanceolatae, saepius acutae,

nitidae, achaeniorum aristis superatae. Disci florum stigmata

caudato-elongata. Achaenia plumbeo-nigra, circumambitu lineari-

REVISION OF THE GENUS COREOPSIS 379

oblonga vel oblanceolata, valde obcompressa, alata alis coloris

similis, glabra, corpore 3-4 mm. longa et (alis inclusis) 1-1.5 mm.

lata, biaristata aristis brevissime sursumque setosis, circ. 0.5-1 mm.

longis.

Type specimen: Collected by Oscar Neumann (Exped. to Sidamo,

Utadera, and Lake Abbaja), No. 134, at altitude of 2,800 meters,

mountain meadows, Gardulla (incorrectly printed Gandulla at

time of my original description in 1923), Gallaland (southern Abys-

sinia), January 14, 1901 (Berl.).

Distribution: Known only from type locality in Abyssinia.

Specimens examined: Neumann IS 4 (type, Berl.).

The single type specimen has its heads in corymbose clusters, and

thus simulates in habit Bidens squarrosa HBK. of tropical America.

69. Coreopsis Feruloides Sherff, Bot. Gaz. 80: 375. 1925.

Herba erecta, ramosa, forsitan annua, circ. 7-10 dm. alta,

glabra, viridis vel glaucescens, caule subtetragono. Folia sessilia vel

subsessilia, 4-8 cm. longa, bipinnata, costa mediana tenui, segmentis

linearibus vel anguste lanceolatis, crassiusculis, minute ciliatis,

obtusis vel subacutis, plerumque 2-4 mm. latis. Capitula sub-

numerosa, corymbosa, radiata, pansa ad anthesin 2.5-3 cm. lata et

5-7 mm. alta. Involucri subglabrati bracteae exteriores circ. 8,

lineares, apice subobtusae indurataeque, 3-5 mm. longae, interiores

lanceolatae circ. 6-7 mm. longae. Flores ligulati circ. 8, flavi, ligula

lineari-oblongi, apice obtuso saepius emarginati et aegre circ. 3-

denticulati, ±1.5 cm. longi et circ. 4-5 mm. lati. Paleae lineari-

oblongae, longitudinaliter 1-3-nerviae, apice subabrupte acutae,

circ. 5.5-7 mm. longae. Disci florum stigmata caudato-attenuata.

Achaenia matura minuta, non exserta, atra, valde obcompressa,

lineari-oblonga, leviter alata, faciebus subobscure striata, faciebus

marginibusque minute erecto-setosa, corpore tantum 3.5-4 mm.

longa et (alis inclusis) parce 1 mm. lata, apice biaristata; aristis

tenuissimis, stramineis, setulis minutis erectis numerosis instructis,

1-1.6 mm. longis.

Type specimen: Collected by E. Battiscombe, No. 945, at altitude

of 2,250-2,400 meters, Kinabop Plateau, Western Aberdare Moun-

tains, British East Africa (Kew).

Distribution: Known only from type locality.

Specimens examined : Battiscombe 945 (type, Kew).

The general habit simulates very closely that of the Mexican

Bidens serrulata (Poir.) Desf. (B. grandiflora Balb.) and B. Ferulaefolia

380 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

(Jacq.) DC. The leaves suggest the primary lower lateral portion

of the more decompound leaves of Ferula glauca L. Aside from the

habit, the tiny achenes are very distinctive, very few African species

of Coreopsis having such small ones. The achenial wings are of the

same color as the body proper, thus easily escaping notice. The

species is somewhat unwelcome, as it serves further to weaken the

generic distinctions between Coreopsis and Bidens.

70. Coreopsis ochracea 0. Hoffm. Bot. Jahrb. 30: 431. 1901;

Sherff, Bot. Gaz. 80: 375 and pi 19. 1925. C. cosmophylla Sherff,

op. cit. 76: 90 and pi. 9, ff. h-n. 1923. Bidens ochracea (0. Hoffm.)

Sherff, op. cit. 158. Fig. 3.

Herba perennis, e radice crassa caules interdum complures emit-

tens, elatos, parce ramosos, glaberrimos, infra obtuse supra acrius

quadrangulatos, 0.8-2 m. altos. Folia pallida petiolata petiolis

marginatis usque ad 4.5 cm. longis, petiolo adjecto 4-10 vel saepe

etiam usque ad 18 cm. longa, pinnatim 3-5-partita, segmentis sub-

crassis, integerrimis vel iterum 3- (rarius 2- vel 4-) partitis, ultimis

linearibus, plerumque 1.5-5 mm. latis, apice acutis et mucronatis,

supra glaberrimis vel breviter scabro-pilosis, subtus glabris vel saepe

ad venam medianam perspicuam sparsim setosis, margine paulo

revolutis et breviter scabro-ciliolatis. Capitula ramos terminantia,

longe pedunculata pedunculis ebracteatis vel foliis reductis bracteatis

et usque ad 3 dm. longis, radiata, pansa ad anthesin 4-8.5 cm. lata

et 1-1.5 cm. alta. Involucri bracteae exteriores 8-10, basi saepe

pilosae, margine sparsim spinuloso-ciliatae, ceterum plerumque

glabrae, lineari-ellipticae vel late lanceolatae, supra basim paulo

constrictae, latiores saepe ad apicem in acumen breve obtusum

subito angustatae, 8-13 mm. longae; interiores lanceolatae vel

ovato-lanceolatae paulo longiores, glabrae vel apice minute pubes-

centes, paleas aequantes vel paulo superantes. Flores ligulati

plerumque 8, aurantiaci, ligula lineari-elliptici, saepe minute pul-

verulento-tomentosi, apice denticulati, 2-3.5 cm. longi. Paleae

oblongo-lanceolatae, acutae. Disci florum stigmata terminaliter

acuto-subcaudata. Achaenia plana, atra, alata, marginibus erecto-

setosa, faciebus glabra, corpore 6-7 mm. longa et 2.5-3.5 mm. lata,

apice setosa et biaristata aristis tenuibus, calvis vel sparsissime

erecto-setosis, circ. 2 mm. longis.

Type specimen: Collected by W. Goetze, No. 731, at altitude of

about 1,700 meters, in thin bush-growth on red laterite on hilly

plateaus, Bweni, Uhehe, German East Africa, March 11, 1899

(Berl.).

FIG. 3. Coreopsis ochracea O. Hoffm.

381

382 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Distribution: Northern Nyassaland northward through German

East Africa into British East Africa.

Specimens examined :E. Battiscombe 83, alt. 1,050-1,200 meters,

Muhoroni, British East Africa (Kew; forma foliis plus minusve

atypica); Goetze 731 (type, Berl.); Muenzner 159, Msamvia, Lake

Tanganyika District, German East Africa, February 24, 1909

(Berl.; type of Coreopsis cosmophylla Sherff); W. H. Nutt, alt.

about 1,800 meters, between Lake Tanganyika and Lake Rukwa

(L. Rikwa, L. Leopold), 1896 (Kew); Ad. Stolz 764, growing 2 meters

tall at alt. 600 meters, Kyimbila, Langenburg, German East Africa,

June 8, 1911 (Berl.); idem (similiter) 764, alt. 600 meters, eodem

loco, February 27, 1912 (Del.) ; A. Whyte, northern Nyassaland (Kew).

Hoffmann had not seen ripe fruits of this species ("Reife Friichte

fehlen; nach den Fruchtknoten zu erteilen konnten sie gefliigelt sein";

Hoffm., loc. cit.). In my own earlier studies it seemed that the

specimens of this handsome species of the Nyassa region were of two

kinds, differing generically on the basis of the winged character of the

achenes. Further specimens have shown, however, that some materi-

al is merely more tardy than the rest in displaying achenial wings.

As the specimens mature, their achenes become fairly well winged.

Explanation of Fig. 3. — Coreopsis ochracea 0. Hoffm.: a, 6,

flowering branch and portion of stem with foliage, X0.58; c, d,

exterior involucral bracts, X2.33; e, interior involucral bract, X2.33;

/, ligulate floret, Xl.75; g, palea, X2.33; h, disc corolla, X2.33; i,

immature achene, X3.5; all from type, Herb. Berl.

71. Coreopsis leptoglossa Sherff, Bot. Gaz. 76: 88. 1923.

Herba annua, erecta, gracillima, speciminibus observatis parce

ramosis, caule tetragono, glabro, 8-10 dm. alto. Folia petiolata,

petiolis latis hispido-ciliatisque usque ad 7 mm. longis vel angustatis

glabratisque usque ad 2.5 cm. longis, petiolo adjecto 5-9 cm. longa,

glabra, bipinnata, segmentis linearibus, plerumque circ. 1 mm.

rarius 1.5-2 mm. latis, apice acutis, margine non ciliatis sed parce

revolutis. Capitula tenuiter pedunculata pedunculis 2-10 cm.

longis, radiata, pansa ad anthesin 4-5 cm. lata et 6-9 mm. alta.

Involucri bracteae basi dense aliter leviter hispidae setis acribus,

arcuatis, multi(6-12)-loculatis, exteriores 8-10, lineares, apice

acutae, margine maximam partem non ciliatae, 5-7 mm. longae et

0.6-0.9 mm. latae, interioribus lanceolatis plerumque breviores.

Flores ligulati circ. 8, lutei, leptoglossati, ligula lineari-elliptici, apice

integri vel obscure denticulati, 2-2.5 cm. longi et 4-6 mm. lati.

REVISION OF THE GENUS COREOPSIS 383

Paleae ignotae. Disci florum stigmata angustato-caudata. Achaenia

matura ignota; ovariis obovatis, valde obcompressis, late alatis,

supra ad margines et medium erecto-hispidis, biaristatis; aristis

erecto-hispidis, 0.4-0.7 mm. longis.

Type specimen: Collected by T. Kassner, No. 2871, on mountain

slope, Lofuku River, Belgian Congo, May 25, 1908 (Berl.).

Distribution: Known only from type locality in Belgian Congo.

Specimens examined : Kassner 2871 (type, Berl.).

Has flowering heads resembling those of Bidens Steppia (Steetz)

Sherff, especially as to involucre.

72. Coreopsis multiflora Sherff, Bull. Jard. Bot. Brux. 13:

292. 1935.

Herba erecta, forsitan perennis, probabiliter ±1 m. alta, caule

vel ramis glabratis atque acriter tetragonis. Folia principalia petio-

lata petiolis marginatis ±1.5 cm. longis, petiolo adjecto ±1.2 dm.

longa, pinnatim circ. 7-partita, foliolis lateralibus oppositis (jugis

remotis), inferioribus circ. 5 cm. longis; omnibus membranaceis,

circumambitu lanceolatis, cum rhachi confluentes, incise lobulatis

(lobulis plus minusve dentatis dentibus acutis) utrinque subsparsim

setosis (setis tenuibus) marginibus ciliatis. Capitula pauca ad

ramos, pedunculata pedunculis subrobustis inferne glabratis superne

setis albidis crispis patentibus hispidis 2-6 cm. longis, radiata, ad

anthesin cernua et pansa 5-6 cm. lata et ±1.3 cm. alta. Involucrum

basi dense hispidum setis albis crispis acribus pluriloculatis, post

anthesin circ. 2-2.3 cm. latum, bracteis subaequalibus; exterioribus

linearibus, acutis, crassiusculis, extus glabratis vel sparsim hispidis,

intus inferne glabris superne densissime hispidis, 8-11 mm. longis;

interioribus ovato-oblongis, extus subsparsim hispidis. Flores

ligulati 8-10, aurantiaci, ligula lineares vel lineari-oblanceolati,

apice minutissime 2-3-denticulati, circ. 2.5 cm. longi et 4-5 mm. lati.

Flores tubulosi numerosi (±125), corollis extrinsecus glabris circ. 5

mm. longis, stylorum ramis longe tenuiterque appendiculatis.

Paleae oblongae, apicem ciliatum obtusumque versus atriores, ad

anthesin saepius aristas aequantes. Achaenia non visa. Ovaria

plana, ovata, alata, infra medium obsolete supra medium manifeste

ciliata setis saepe erectis, faciebus glabrata, corpore circ. 3 mm. longa

et circ. 1.5 mm. lata, apice biaristata aristis tenuibus antrorsum his-

pidulis 1-1.5 mm. longis.

Type specimen: Collected by Dom Benoit Thoreau, No. 2. 11, Pla-

teau of Bianos, Kansenia, Katanga, Belgian Congo, June, 1927 (Bruss.).

384 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Distribution: Katanga, Belgian Congo.

Specimens examined: Thoreau 2. 11 (type, Bruss.).

73. Coreopsis bella Hutch. Kew Bull. 364. 1907.

Herba erecta, verisimiliter perennis, suffruticosa, ±1 m. alta,

ramis parce pubescentibus. Folia petiolata petiolis ±2-3 cm. longis,

petiolo adjecto 7-15 cm. longa, tenere coriacea, circumambitu trian-

gulata vel ovato-triangulata, pinnatipartita, supra breviter subtus

longe pubescentia segmentis ovato-lanceolatis acutis plus minusve

profunde serratis vel irregulariter pinnatilobis. Capitula tenuiter

pedunculata pedunculis pubescentibus ±2-6 cm. longis, radiata,

pansa ad anthesin circ. 5.5-6.5 cm. lata et circ. 1.4-1.7 cm. alta.

Involucri bracteae obtusae, circ. 1.3 cm. longae et circ. 4 mm. latae,

exteriores oblongae, pilosae; interiores extrinsecus pubescentes.

Flores ligulati circ. 12, flavi, ligula oblongo-elliptici, circ. 3 cm. longi,

apice subacuti vel obscure dentati, tubo brevi extrinsecus pilosi.

Paleae oblongae, glabrae, apice obtusae, apicem versus leviter con-

strictae, circ. 1 cm. longae et circ. 1.5 mm. latae. Achaenia immatura

plana, pilosa, corpore circ. 1 cm. longa, apice biaristata aristis 1.5

cm. longis.

Type specimen: Raised at Kew in 1907 from seeds collected in

British East Africa by R. Diespecker (Kew).

Distribution: British East Africa.

Specimens examined : Cult, at Kew from seeds collected by Dies-

pecker (type, Kew).

Close in general appearance to some African species of Bidens.

Mature achenes are much to be desired.

74. Coreopsis Curtisii Sherff, Bot. Gaz. 96: 146. 1934. C.

Curlissii Sherff, Bull. Jard. Bot. Brux. 13: 289. 1935 (sphalm).

Herba erecta, verisimiliter perennis, gracilis, forsitan circ. 0.5

m. alta, subsimplex, caule angulato glaberrimo plus minusve pur-

pureo-striato. Folia opposita, tenuiter petiolata petiolis glaberrimis

basi connatis usque ad 2 cm. longis, petiolo adjecto usque ad 8 cm.

longa, pinnata vel bipinnatisecta, laminae circumambitu triangulato-

ovata, segmentis primariis circ. 3 vel 5 pallidis membranaceis lanceo-

latis acriter dentatis secundum venas marginesque nunc sparsissime

nunc subconferte albido-setulosis. Capitula pauca, ±3-adgregata et

ad rami finem corymbose disposita pedunculis erectis tenuibus glaber-

rimis 4-8 cm. longis, radiata, pansa ad anthesin circ. 3-3.5 cm. lata

et circ. 11-13 mm. alta. Involucri pallidi bracteae supra reflexae,

REVISION OF THE GENUS COREOPSIS 385

exteriores circ. 8 vel 9, lineari-oblongae, glabrae vel basi extrema

hispidae, apice acutae, ±4 mm. longae, interioribus ovato-oblongis

apice glanduloso-pubescentibus moderate breviores. Flores ligulati

circ. 8, aurantiaci, ligula anguste oblongi, apice integri vel sub-

denticulati, ±1.5 cm. longi. Paleae lineari-oblongae, apicem versus

valde aurantiacae, apice ipso subobtusae. Flores tubulosi extrin-

secus glabrae vel secundum corollae superne valde aurantiacae

dentes pubescentes et ad tubi apicem sparsim pilosi; stylorum ramis

pulcherrime aurantiacis et elongato-appendiculatis. Achaenia ma-

tura non visa. Ovaria oblanceolata, plana, nitido-brunnea, longi-

tudinaliter pluristriata, plus minusve alata, marginibus et faciebus

saltern supra erecte setosa, corpore ±4 mm. longa, apice valde

erecto-setosa et biaristata aristis glaberrimis ±1.5 mm. longis.

Type specimen: Collected by Mrs. Richard C. Curtis from

material cultivated in 1930 from seeds obtained in Angola, Portu-

guese West Africa (Field).

Distribution: Known only from Angola.

Specimens examined: Mrs. Curtis, cult. 1930 (type, Field; cotype,

Gray).

A species offering a superficial resemblance in leaf outline and

dissection to C. oligoflora.

75. Coreopsis exilis Sherff, Bull. Jard. Bot. Brux. 13: 288. 1935.

Herba perennis, erecta, gracillima, simplex, 0.5-1.25 m. alta,

caule subtetragono glabro infra circ. 3 mm. supra circ. 1.2 mm.

crasso. Folia petiolata petiolis tenuibus anguste marginatis glabris

basi vix connatis 1.5-3 cm. longis, petiolo adjecto circ. 7-9 cm.

longa, pinnatim 3-5-partita, foliolis oblonge lanceolatis (vel superi-

orum lineari-oblongis), pauciter acridentatis, membranaceis, pallidis,

ciliatis, utrinque subsparsim adpresso-hispidis, apice acutis. Capi-

tulum (unicum visum) pedunculatum pedunculo tenui glabrato

+11 cm. longo, radiatum, pansum ad anthesin circ. 3.5 cm. latum et

+ 1.5 cm. altum. Involucri bracteae aequilongae, exteriores circ.

8, adpressae, oblonge lineares, basim versus ciliatae ac moderate

hispidae supra sparsim adpresso-hispidae vel glabrae, apice acutae,

longitudinaliter saltern inferne 3-lineata lineis atris, 9-10 mm.

longae; interiores ovato-oblongae, moderate adpresso-hispidae.

Flores ligulati circ. 8, aurantiaci, ligula subovati, apice 2-3-denti-

culati, ±1.7 cm. longi. Paleae subacutae, glabrae vel aegre pubes-

centes, ovariorum aristis superatae. Flores tubulosi extrinsecus

glabri, supra aurantiaci, corolla circ. 8 mm. longi. Achaenia matura

386 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

non visa. Ovaria plana, alata, anguste cuneato-obovata, supra

marginibus faciebusque erecto-setosa, corpore circ. 5.5 mm. longa,

biaristata aristis acerrimis stramineis infra antrorsum paucisetosis

supra glabris circ. 4 mm. longis.

Type specimen: Collected by P. Quarre, No. 419 pro parte, on

farm of H. Droogmans, Katuba, Katanga, Belgian Congo, May 15,

1927 (Bruss.; nom. indig. Kibunyubunyu).

Distribution: Katanga, Belgian Congo.

Specimens examined: Quarre 419 pro parte (type, Bruss.).

Similar in habit to C. Curtisii of Angola but differing in exterior

involucral bracts, which are appressed, moderately hispid below

the middle, and about 9-10 mm. long (not reflexed, not "glabrous

except at very base," and not ±4 mm. long); in ovarial aristae, which

are about 4 mm. long and somewhat antrorse-setose below (not

±1.5 mm. long and not extremely glabrous) ; etc. The median leaves

of the type have the terminal leaflet linear-oblong, about 5 cm.

long, and few-dentate (1-3 teeth on each side). Root described by

Quarr£ as running ("racine tracante").

76. Coreopsis oligoflora Klatt, Leopoldina 25: 107. 1889.

C. oligantha Klatt, Ann. Naturh. Hofmus. Wien 7: 103. 1892.

Involucri bracteae exteriores lineares, acriter apiculatae, circ. 1 mm.

latae C. oligoflora sensu stricto.

Involucri bracteae exteriores late oblongae vel ovatae, apicaliter

obtusissimae, saepius 2.5-4 mm. latae var. /3 robusta.

Herba erecta, annua, 5-10 dm. alta, ramosa ramis angulatis, caule

ramisque sparsim pilis tenuissimis albis elongatis hispidis. Folia

opposita, pilis tenuissimis argenteo-nitidis elongatis plus minusve

hispida, breviter petiolata petiolis alato-marginatis basaliter con-

natis usque ad 1.5 cm. longis, petiolo adjecto 3-6 (-10) cm. longa,

bipinnatisecta, segmentis majoribus ovatis vel lineari-lanceolatis,

membranaceis, aegre atropunctulatis, dentatis dentibus acriter

calloso-apiculatis. Capitula tenuiter pedunculata pedunculis saepius

7-13 cm. longis, radiata, pansa ad anthesin 4-6 cm. lata et circ. 1

cm. alta. Involucri hispidi bracteae exteriores circ. 8, lineares,

nitide acriterque calloso-apiculatae, 6-10 mm. longi et circ. 1 mm.

latae, interioribus oblongis plerumque aequales. Flores ligulati circ.

8, lutei vel aurantiaci, ligula oblanceolato-elliptici, apice integri vel

minutissime denticulati, 2.5-3 cm. longi et usque ad 12 mm. lati.

Disci florum stigmata terminaliter angustata. Achaenia late

lineari-oblonga, nigra, plana, apicem versus minute erecto-setosa,

REVISION OF THE GENUS COREOPSIS 387

lateraliter alata alls 0.6-0.8 mm. latis, marginibus erecto-setosa,

corpore circ. 8 mm. longa et cum alis circ. 3-3.5 mm. latis, apice

erecto-setoso biaristata aristis tenuibus ±2 mm. longis antrorsum

hispidis.

Type specimen: Collected by Dr. Buchner, No. 32, among bushes,

Malange, Angola, Portuguese West Africa, March, 1879 (Berl.).

Distribution: Belgian Congo and south westward across Portu-

guese West Africa.

Specimens examined (this list doubtless includes some specimens

of the more recently described var. robusta) : L. Achten 253B, Luobo-

Kasai, Belgian Congo (Bruss.) ;F. Allard 65, Kuku Wungu, Belgian

Congo, 1909 (Bruss.); idem 330, Kikubungu, Belgian Congo, 1910

(Bruss.); Baum 878, alt. 1,300 meters, between Kulei and Kutsi,

Mambunda Region, southeastern Angola, Portuguese West Africa,

April 29, 1900 (Berl.); Bavicchi 14, Esungume Dolo, Belgian Congo,

1913 (Bruss.); J. Bequaert 113, Bukama, Belgian Congo, May 12,

1911 (Bruss., 2 sheets); Dr. Buchner 32 (type, Berl.); J. Claessens 67,

Lubuku, Belgian Congo, September, 1907 (Bruss., 2 sheets); idem

73, Belgian Congo, August, 1907 (Bruss.); idem 88, Ka-Tembo,

Belgian Congo, January 18, 1921 (Bruss., 2 sheets}; idem 400, Koloko

Konbe, Belgian Congo, January, 1910 (Bruss.); F. Flamand,

Kiamohampa, Belgian Congo, September 20, 1911 (Bruss.); J.

Gillet 806 and 885, Kisantu, Belgian Congo, 1900 (Bruss.); P.

Hamerlinck 1502 and 1512, Wombali, Belgian Congo, 1914 (Bruss.);

Fr. Hens 45, becoming shrubby and growing 1.5 meters high at alt.

270 meters, marsh of Stanley Pool, Belgian Congo, August 14, 1888

(Bruss. ; Kew) ; Em. Laurent, between Lusambo and Lomani, Belgian

Congo, December, 1895 (Bruss.); idem & Marc. Laurent, under-

growths, Kondue, Belgian Congo, November 25, 1903 (Bruss.);

Marc. Laurent 512, Bolobo, Belgian Congo, April 15, 1905 (Bruss.);

E. Lescrauwaet 188, Mokole (Lubi), Belgian Congo, September 12,

1904 (Bruss.); Alexander von Mechow 33, Pungo Andongo, Angola,

Portuguese West Africa, January-April, 1879 (Berl., 2 sheets); idem

459, Malange, Angola, March, 1880 (Berl.); Nelis, Bokala, Belgian

Congo, May, 1913 (Bruss.); P. Pogge 257, Belgian Congo, March,

1876 (Berl.); idem 1282, in cultivations, Mukenge tract (campine),

Belgian Congo, November 6, 1881 (Berl.); idem 1295, eodem loco,

March 15, 1882 (Berl.); idem 1309, prairie (campine) on the Lualaba

River, southeastern Belgian Congo, April 24, 1882 (Berl.); A. Sapin,

N'Dolo, Belgian Congo, May, 1906 (Bruss.); idem, N'Kolokosso,

Belgian Congo, June 28, 1906 (Bruss.); idem, Sokomini Lake,

388 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

Sankuru (Lubilash) River, Belgian Congo, September, 1906 (Bruss.) ;

idem, Katola Msek, Belgian Congo, April, 1908 (Bruss., 2 sheets);

idem, growing on the plains, Munungu, Belgian Congo, April, 1910

(Bruss.); R. Schlechter 12474, Dolo, Belgian Congo, May- June,

1899 (Berl.; Bruss.; Kew); Sparano 25, "odor recalling slightly the

marguerite," Belgian Congo, June 6, 1913 (Bruss.); Hyacinthe

Vanderyst, Ndembo, Belgian Congo, August 20, 1906 (Bruss.);

idem, eodem loco, January 7, 1907 (Bruss.); idem, Station at Inkissi

Falls, Belgian Congo, May, 1907 (Bruss.); idem, Leopoldville,

Belgian Congo, May 23, 1907 (Bruss.) and May 27, 1910 (Bruss.);

idem, Kumpako, Belgian Congo, 1908 (Bruss.); idem, N'Boma,

Belgian Congo, April 28, 1911 (Bruss.); idem B224, Lonzo, Belgian

Congo, July 26, 1906 (Bruss.); idem 700, Wombali, Belgian Congo,

May, 1913 (Bruss.); idem 1099, eodem loco, June, 1913 (Bruss.);

idem 1187, Bandundu, Belgian Congo, June, 1913 (Bruss.); idem

1341, Wombali, Belgian Congo, June, 1913 (Bruss.); idem 1582,

Bokala, Belgian Congo, July, 1913 (Bruss.); idem 4304, Bugunu,

Belgian Congo, May 28, 1914 (Bruss.); idem 4885, Wombali, Belgian

Congo, 1914 (Bruss.); idem 10097, Kamtsha, Belgian Congo, July,

1921 (Bruss.); idem 20963, Lulua-gare, Belgian Congo (Bruss.);

E. Verdick, Lukafu, Katanga, Belgian Congo, March, 1900 (Bruss.) ;

F. C. Wellman, alt. 1,500 meters, Bailundo District, Benguella

(Benguela; Abunda) region, eastern Angola (Kew).

Often confused with Bidens Steppia (Steetz) Sherff, a species

having more elongate exterior involucral bracts, mostly 11 or 12

slender-liguled ray florets instead of about 8 rather wide ones,

leaves larger and more decompound, achenes practically exalate, etc.

Coreopsis oligiflora var. 0 robusta Sherff, Bot. Gaz. 90: 386.

1930.

A specie involucri bracteis exterioribus spathulato-oblongis vel

oblongis vel ovatis saepius 2.5-4 mm. latis apicaliter obtusissimis

differt.

Type specimen: Collected by R. Buttner, No. 408, Bolobo,

Belgian Congo, November 5, 1885 (Berl.).

Distribution: Belgian Congo.

Specimens examined: Joseph Bequaert, Leopoldville, Belgian

Congo, April 9, 1915 (Bruss.); idem 7205, Belgian Congo, March

29, 1915 (Bruss.); Buttner 408 (type, Berl.); DeGiorgil42, Kisengwa,

Belgian Congo, October, 1922 (Bruss.); idem 231, herb of the moist

plain, Kanda-Kanda, Belgian Congo, November, 1922 (Bruss.);

D. H. Linder 1 706, moist places at edge of savanna, Bolobo, Belgian

REVISION OF THE GENUS COREOPSIS 389

Congo, December 16, 1926 (Berl.); P. Pogge 1292, Mukenge tract

(campine), Belgian Congo, April 13, 1882 (Berl.); P. Quarre 2173,

7-9 dm. tall, on plateau, Saeomintra, Belgian Congo, November,

1930 (Bruss., 2 sheets) ; idem 2826, growing 1 meter tall, Pastorale,

Division of Lualu, Belgian Congo, November, 1931 (Bruss., 2 sheets) ;

idem 2921, herb 1.2 meters tall, Lovoi, Kamina, Belgian Congo,

March, 1932 (Bruss., 2 sheets); Walter Robyns 138, much branched

at base, ±1.5 meters tall, right bank of Guhisi, beyond railroad,

Kisantu, Belgian Congo, July 3, 1925 (Bruss.); H. Vanderyst 12366,

between La Labue and La Loange, Belgian Congo, October, 1922

(Bruss.); idem 21227, Belgian Congo (Bruss.); F. Vermoesen 2403,

Belgian Congo, September, 1919 (Bruss., 2 sheets).

Dr. Otto Hoffmann had designated the type varietally under the

name C. oligantha Klatt in the Berlin Herbarium, but I have not as

yet found that he ever published the varietal name.

77. Coreopsis Quarrei Sherff, Bull. Jard. Bot. Brux. 13: 289.

1935.

Herba annua, gracilis, erecta, ±5 cm. alta, caule moderate

erecto-ramoso glabrato nunc 1 nunc ±2.2 mm. crasso. Folia petiolata

petiolis usque ad 1.3 cm. longis setoso-ciliatis setis pluriloculatis,

petiolo adjecto usque ad 5 cm. longa, bipinnatisecta, foliolis mem-

branaceissimis acute dentatis lobulatisve supra moderate infra

(interdum sparsius) adpresso-hispidis. Capitula radiata, pansa ad

anthesin ±4 cm. lata et circ. 1 cm. alta, tenuiter pedunculata pedun-

culis caulem ramosve terminantibus glabris vel summam versus

aegre setosis usque ad 1.2 dm. altis. Involucri demum hemisphaerico-

depressi bracteae exteriores circ. 7 vel 8, glabratae vel aegre hispidae,

lineari-oblongae, supra interdum dilatatae, apice rotundatae vel

subacutae, 3-4 mm. longae, quam interiores ovato-oblongae sparsim

setosae dimidio breviores. Flores ligulati ±8, aurei, lineari-oblongi,

apice integri vel bidenticulati, ±2.3 cm. longi. Paleae oblongae

vel lineari-oblongae, apice subcolorato obtusissimae vel subtrun-

catae, 6-7.5 mm. longae. Flores tubulosi circ. 4 mm. longi, extrin-

secus praecipue ad tubi summam saepe patenti-hispidi. Achaenia

plana, brunneo-atra, obovata vel oblonga, alata, corpore 3.5-6 mm.

longa et alis inclusis 2-2.5 mm. lata, corporis plurisulculati ipsius

facie ventrali erecte setosa setis e tuberculo ortis, facie dorsali non

nisi apicem versus tuberculato-setosa, marginibus pectinato-tuber-

culatis saepius brunneis flavisve numerose erecto-setosa, apice erecte

setulosa ac minutissime biaristata aristis tenuibus antrorsum his-

pidis sub 0.5 mm. longis.

390 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: Collected by P. Quarre, No. 419 pro parte, on

farm of H. Droogmans, Katuba-Elisabethville, District of Upper

Katanga, Belgian Congo, May 16, 1927 (Brass.).

Distribution: District of Upper Katanga, Belgian Congo.

Specimens examined: Quarre 419 pro parte (type, Brass.).

Confused by Quarre with C. exilis, from which it differs in being

an annual, having the exterior involucral bracts 3-4 mm. long and a

half shorter than the inner ones, the achenial bodies varying from

3.5 to 6 mm. long, the aristae less than 0.5 mm. long, etc. C. exilis

is perennial, has involucral bracts subequal and 9-10 mm. long,

achenial bodies about 5.5 mm. long, aristae about 4 mm. long, etc.

78. Coreopsis Goffardii Sherff, Bull. Jard. Bot. Brux. 13:

290. 1935.

Herba annua, erecta, 2.5-8 dm. alta; caule glabro, subtetragono,

purpurascenti, suberecte ramoso. Folia petiolata petiolis marginatis

sparsim hispidis (setis pluriloculatis) 1-3 cm. longis, petiolo adjecto

usque ad 1 dm. longa, 2-3-pinnatisecta rhachi petiolo marginato

simile; segmentis ultimis nunc oblongis nunc obovato-oblongis

nunc plus minusve lanceolatis, utrinque moderate adpresso-his-

pidula et saepe praecipue supra minutissime glandulosa, membra-

nacea, dentibus saepius subabrupte minutissimeque apiculatis.

Capitula ramulos tenues hispidos 1-7 cm. longos terminantia, radiata,

pansa ad anthesin ±3 cm. lata et ±11 mm. alta. Involucrum

hemisphaericum basi dense superne subsparsim adpresso-hispidum,

bracteis exterioribus circ. 8, lineari-oblongis, superne saepe latioribus,

tergo longitudinaliter 1-3-atronerviis, apice acutis vel subacutis,

circ. 4-5 mm. longis, quam interioribus oblongo-ovatis paulum

brevioribus. Flores ligulati circ. 8, aurantiaci, ±1.5 cm. longi,

ligula anguste oblongi, inferne glabrati superne minutissime adpresso-

hispiduli, apice subintegri. Paleae oblongae, apice obtusae rotunda-

taeve ac erecte ciliolatae, ad anthesin quam corollae tubulosae

breviores. Flores tubulosi extrinsecus glabra vel tubi summa

(anulo) hispiduli, dentibus saepius aurantiaci, 4-5 mm. longi.

Ovaria (achaeniis maturis deficientibus) plana, oblongo-obovata,

faciebus glabratis mediane paucistriata, alata, corpore sub 3 mm.

longa, marginibus infra obsolete eroso-denticulata summam versus

erecte setosa, apice cuneato-emarginato erecte setosa ac biaristata

aristis tenuibus antrorsum hispidulis sub 0.5 mm. longis.

Type specimen' Collected by P. Quarre, No. 1616, on Mr.

Goffard's farm, Kipila-Elisabethville, District of Upper Katanga,

Belgian Congo, May, 1929 (Brass.).

REVISION OF THE GENUS COREOPSIS 391

Distribution: Known only from type locality in District of Upper

Katanga, Belgian Congo.

Specimens examined: Quarre 1616 (type, Bruss.).

79. Coreopsis vulgaris Sherff, Bull. Jard. Bot. Brux. 13: 291.

1935.

Herba annua, erecta, ±4 dm. alta, caule glabrato plus minusve

tetragono, apicem versus suberecte ramoso. Folia petiolata petiolis

tenuibus patenti-ciliatis interdum submarginatis usque ad circ. 2

cm. longis, petiolo adjecto circ. 5-8 cm. longa, 2-3-pinnatifida,

segmentis primariis ±5 ovatis vel ovato-lanceolatis, ultimis mem-

branaceis utrinque adpresso-hispidis dentatis dentibus acriter

apiculatis. Capitula pauca (±7) , pedunculata pedunculis subrobustis

sparsissime patenti-hispidis (setis crispis albidisque) 1-4 cm. longis,

radiata, pansa ad anthesin circ. 3-3.5 cm. lata et ±1.2 cm. alta.

Involucrum magnum depresso-hemisphaericum, ±2 cm. latum et

±1 cm. altum, setis acerrimis crispis albidis basi dense alibi sub-

sparsim conspersum; bracteis exterioribus circ. 8, lineari-oblongis,

longitudinaliter circ. 3-lineatis lineis atris, apice obtusis vel acutis,

6-11 mm. longis; interioribus ovato-lanceolatis, apice glanduloso-

hispidulis, saepius 8-10 mm. longis. Flores ligulati circ. 8, flavi,

ligula lineari-oblongi, apice integri vel subintegri, circ. 1-2 cm.

longi. Paleae oblongae, apice atriore subrotundatoque vix apicu-

latae, aristis superatae. Flores tubulosi extrinsecus glabri. Achaenia

non visa. Ovaria plana, oblonga vel obovata, faciebus glabrata,

marginibus (infra breviter supra longe) apiceque erecto-setosa,

alata, corpore alis inclusis 3-4 mm. longa et 1-1.5 mm. lata, biaristata

aristis tenuibus antrorsum hispidis circ. 2.5-4 mm. longis et corollae

tubum dimidio superantibus.

Type specimen: Collected by A. Becquet, No. 62, at altitude of

1,000 meters, common on savanna, flowering entire year except in

dry season, Mukishi-Lomam, District of Middle Katanga, Belgian

Congo, September, 1927 (Bruss.).

Distribution : District of Middle Katanga, Belgian Congo.

Specimens examined : Becquet 62 (type, Bruss.).

The annual habit, large discs, and comparatively inconspicuous

rays combine to give this species the appearance of an unattractive

weed. — Referred to Coreopsis (instead oiBidens) with some hesitancy,

since it is not known that the mature achenes are alate.

80. Coreopsis Giorgii Sherff, Bull. Jard. Bot. Brux. 13:

292. 1935.

392 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Herba annua, erecta, ±1 m. alta, caule glabro subtetragono

tenuiter ramoso. Folia petiolata petiolis tenuibus anguste mar-

ginatis hispido-ciliatis circ. 1-1.5 cm. longis, petiolo adjecto ±7 cm.

longa, bipinnatifida, segmentis secundariis oblongo-lanceolatis vel

ovato-lanceolatis membranaceis infra moderate supra subsparsim

adpresso-hispidis acriter dentatis. Capitula subnumerosa sub-

corymbose ad ramorum apices disposita, pedicellata pedicellis

sparsissime setosis 3-6 cm. longis, radiata, pansa ad anthesin circ.

2.5-3.3 cm. lata et circ. 8-9 mm. alta. Involucrum moderate ad-

presseque albido-setosum, bracteis exterioribus oblongo-linearibus,

apice induratis atque acutis subobtusisve 4-6 mm. longis quam

interioribus ovato-lanceolatis plerumque brevioribus. Flores ligulati

circ. 8, flavi, ligula lineari-oblongi, apice integri vel minutissime

denticulati, circ. 1.5-2 cm. longi. Paleae glaberrimae, cuneate

oblanceolatae, apice atriore rotundatae. Flores tubulosi glabri.

Achaenia valde obcompressa, nitide brunneo-atra, faciebus pluri-

striata et (praecipue secundum medium atque apicem versus) erecto-

setosa, alarum marginibus minutissime numerosissimeque erecto-

setulosa, corpore alis inclusis exteriora ±7 mm. longa et ±3.2 mm.

lata interiora ±1 cm. longa et ±1.7 mm. lata, omnia apice densissime

erecto-hispida et biaristata aristis perspicue densissimeque erecto-

hispidis circ. 1 mm. longis, setulis stramineo-fulvis.

Type specimen: Collected by De Giorgi, No. 170, Katompe,

District of Middle Katanga, Belgian Congo, October, 1922 (Bruss.).

Distribution: District of Middle Katanga, Belgian Congo.

Specimens examined: De Giorgi 170 (type, Bruss.).

Related to C. oligoflora and C. Mattfeldii. From C. oligoflora

it differs at once in having the exterior involucral bracts 4-6 (not

6-10) mm. long, the ligulate florets about 1-2 (not about 2.5-3)

cm. long, etc. From C. Mattfeldii it differs in having the exterior

achenes, wings included, about 3.2 (not 4-5) mm. wide, etc.

81. Coreopsis Mattfeldii Sherff, Bot. Gaz. 76: 83. 1923.

Herba erecta, ramosa, verisimiliter circ. 7-10 dm. alta, caule

subtetragono, glabrato vel disperse piloso, sulcato. Folia petiolata

petiolis latis ±1 cm. longis, petiolis adjectis 4-10 cm. longa, pinna-

tisecta, utrinque moderatim pilosa, margine parce ciliata, laciniis

oppositis indivisis vel pinnatisectis, segmentis ultimis linearibus

(2-6 mm. latis), acutis atque indurato-apiculatis. Capitula corym-

bosa, radiata, pansa ad anthesin circ. 3.5-4 cm. lata et 9 mm. alta.

Involucri bracteae dorso hispidae pilis albis multiloculatis atque ad

basim tumidis, aequilongae (5-8 mm.), exteriores lineari-spathu-

REVISION OF THE GENUS COREOPSIS 393

latae, interiores ovato-lanceolatae. Flores ligulati circ. 8, flavi,

ligula lanceolati vel obovato-lanceolati, striati, apice obscure minu-

teque denticulati, 1.6-2 cm. longi et 4-9 mm. lati. Paleae hyalinae

late oblongae, apice subobtusae, circ. 8 mm. longae. Disci florum

stigmata angustatissimo-caudata. Achaenia valde obcompressa,

nigrescentia, alata alis 1-1.5 mm. latis, faciebus glabra vel ad costas

(et supra sparsim) hispida, margine apiceque hispida, biaristata

aristis tenuibus stramineis sursum hispidis 1-2 mm. longis, exteriora

obovata 6-8 mm. longa et (alis inclusis) 4.25-5 mm. lata, interiora

oblanceolata circ. 8 mm. longa et (alis inclusis) 4 mm. lata.

Type specimen: Collected by Alexander Carson, No. 75, Fwamba,

Lake Tanganyika, German East Africa, 1894 (Berl.).

Distribution: Known only from type locality in German East

Africa.

Specimens examined: Carson 75 (type, Berl.; cotype, Kew).

Named very appropriately after Dr. Johannes Mattfeld, who

most generously assisted in placing at my disposal many of the

Berlin Herbarium specimens of African Coreopsis and Bidens for

detailed study. It bears a strong superficial resemblance to Bidens

Steppia (Steetz) Sherff, but can easily be distinguished by its achenes

and involucral bracts. The many mature achenes on the type differ

remarkably from the ones on the type and other specimens (already

cited in my monograph of the genus Bidens) of Bidens Steppia. For

B. Steppia they are at most widely oblong-linear in outline, 7.5-9

mm. long and 2.5-3 mm. wide; there is merely a thickened lateral

ridge, wings being absent. For Coreopsis Mattfeldii the achenes are

much thinner and wider, the outer ones being obovate, 6-8 mm. long

and 4.25-5 mm. wide, the inner ones being oblanceolate, about 8

mm. long and 4 mm. wide; all are winged, and even on submature

fruiting heads the rounded upper ends of the wings, adjacent to the

aristae, are easily visible to the eye. The external bracts about

equal the inner ones in length in C. Mattfeldii; in Bidens Steppia

they tend to exceed the inner ones by 2-6 mm.

The nearest ally in Coreopsis is C. oligiflora Klatt.

82. Coreopsis injucunda Sherff, Bull. Jard. Bot. Brux. 13:

293. 1935.

Herba annua, erecta, ramosa, 4-10 dm. alta, caule ramisque

subtetragonis glabris. Folia petiolata petiolis tenuibus marginatis

hispido-ciliatis setis pluriloculatis usque ad ±2.5 cm. longis, petiolo

adjecto ±6 cm. longa, bipinnatisecta segmentis linearibus subacriter

394 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

apiculatis membranaceis utrinque marginibusque sparsim albido-

hispidis. Capitula moderate numerosa, pedunculata pedunculis

ramos terminantibus sparsim hispidis tenuibus saepius 1-1.5 dm.

longis, radiata, pansa ad anthesin circ. 4-4.5 cm. lata et 1.2-1.4 cm.

alta. Involucri hispidi bracteae exteriores circ. 8, subulato-lance-

olatae, basi connatae, apice acutae, circ. 4-6 mm. longae, quam

interiores oblongo-ovatae paulo breviores. Flores ligulati circ. 8,

flavi, ligula oblonge lineares, apice integri, circ. 2.5 cm. longi. Paleae

oblongae, apice obtusissimae vel subtruncatae, achaeniorum aristas

superantes. Florum tubulosorum corollae circ. 5 mm. longae,

extus glabrae. Achaenia plana, atra, obovata vel oblonga, corpore

4.5-6.5 mm. longa et alis inclusis circ. 2-2.3 mm. lata, faciebus

pluristriata dorsali parce ventrali moderate erecto-setulosa, margini-

bus apiceque numerose erecto-ciliata, biaristata aristis tenuibus

stramineis antrorsum hispidulis sub 1 mm. longis.

Type specimen: Collected by P. Quarre, No. 1662, on Mr.

Goffard's farm, Kipila-Elisabethville, District of Upper Katanga,

Belgian Congo, May, 1929 (Bruss., 2 sheets).

Distribution: District of Upper Katanga, Belgian Congo.

Specimens examined: Quarre 1662 (Bruss., 2 type sheets).

83. Coreopsis Borianiana Schz. Bip. ex Schweinf. Verh. Zool.

Bot. Ges. Wien 18: 684. 1868. C. Borianiana var. cannabina Schz.

Bip. loc. cit. C. guineensis 0. & H. in Oliver, Fl. Trop. Afr. 3: 390.

1877; etiam in Hutch. & Dalz. Fl. West Trop. Afr. 2: 143, /. 217.

1931. C. Chevalieri 0. Hoffm. & Muschl. Bull. Soc. France 57,

Me"m. 8: 118. 1910. C. chrysopterocarpa Chiov. Ann. Bot. Roma

9: 75. 1911.

Involucri bracteae exteriores 9-12, uniseriales, 0.7-1.5 cm. longae.

C. Borianiana sensu stricto.

Involucri bracteae exteriores 18-22, biseriales, plerumque 1.4-1.8

cm. longae var. /3 multiplex.

Herba annua, erecta, plus minusve glabra, 6-1.5 dm. alta, caule

aegre subtetragona vel etiam fere teres. Folia opposita saepius

breviter petiolata petiolis latis marginatis plerumque 3-12 mm.

longis, petiolo adjecto 5-13 (etiam -18) cm. longa, simplicia vel

2-3-partita, lamina segmentisve remote acriterque denticulatis

(denticulis saepe subcapilliformibus), linearibus, utrinque elongato-

angustatis, plerumque 4-10 mm. latis, membranaceis, minutissime

spinuloso-ciliatis. Capitula non numerosa, tenuiter pedunculata

pedunculis 5-18 cm. longis ad apicem saepe abrupte dilatatis ac

REVISION OF THE GENUS COREOPSIS 395

paucis bracteis bracteae involucri exterioris valde similibus obsitis,

radiata, pansa ad anthesin 5-7 cm. lata et 1.3-2 cm. alta. Involucri

bracteae exteriores 9-12, uniseriales, anguste lineares, tergo glabrae

vel hispidae, ad apicem saepe minute spinuloso-ciliatae, acriter vel

subacriter (saepe indurate-) apiculatae, 0.7-1.5 cm. longae et 0.6-1

mm. latae; interiores lanceolato-oblongae vel ovato-oblongae, apice

plerumque dorso interdum pubescentes, saepius paulo longiores.

Flores ligulati circ. 8, aurei, ligula obovato-oblongi, 8-14-striati,

apice obtuso obscure vel manifeste denticulati, 2.5-3 cm. longi et

8-14 mm. lati. Paleae oblongo-oblanceolatae, demum interdum

crassiusculae, circ. 10-18 mm. longae et 2.5-4.5 mm. latae. Disci

florum stigmata caudato-appendiculata. Achaenia valde obcom-

pressa, alata, circumambitu (alis inclusis) ovato-orbiculata, corpore

ipso nigra lineari-oblonga tuberculato-setosa 9-15 mm. longa

et 1.2-2.3 mm. lata, alis badia vel argentea nitida dense setoso-

ciliata circ. 2.8-3.8 mm. lata, apice setosa et biaristata aristis erectis,

sursum hispidis, circ. 2.5-4 mm. longis.

Type specimen: Collected by Boriani, No. 42, in the Fesoglu,

northeastern Africa, 1839 (Mus. V.).

Distribution: Across Africa from Eritrea to Se'ne'gal and reaching

southward to Kamerun and Togo.

Specimens examined: C. Barter 933, Nigeria, 1857-1859 (Berl.;

Gray); Boriani 42 (type, Mus. V.; etiam fragmentum typi in herb.

Schz. Bip. in Par.); Aug. Chevalier 2083, Koulitsoro, Sudan, 1899

(Berl. ; Bruss.) ; idem 9633, Massenya (Massenia), Bagirmi (Baguirmi),

Sudan, 1902-1904 (Berl.); idem 9721, Kolke'te and Moito, Dar-el-

Hadjer, Baguirmi Nord, Territory of Chari, September 6-9, 1903

(herb. Chevalieri; type of C. Chevalieri 0. Hoffm. & Muschl.); Dr.

Kersting 14, alt. 350 meters, Trogode, Kerkri, Togo, October 28,

1897 (Berl.) ; idem 422, alt. 400 meters, Sokode'-Basari, Togo, October,

1901 (Berl.); Lecard 290, Se'ne'gal (Bruss.); C. Ledermann 5243,

growing 1-1.5 meters high, alt. 380 meters, between Tschamba and

Doreba, Karnerun, September 18, 1909 (Berl., 2 sheets); A. Pappi

7733, alt. 1,000 meters, Baza Cunama-Barentu, Erythrea, November

28, 1908 (Flor., type of Coreopsis chrysopterocarpa Chiov.); T. Pfund,

Dar-Fur, 1876 (Berl.) ; idem 336, Takari on the Rahad, Kordofan,

July, 1875 (Berl., 2 sheets); Dr. Rowland, common in savannas,

western Lagos, 1893 (Berl., 2 sheets) ;F. Schroeder 74, alt. 350 meters,

Sokod£ Farm, Togo, October 19, 1900 (Berl., 2 sheets); G. Schwein-

furth (Flora of Gallabat No. 439), Matamma, Gallabat (Galabat)

region, northwestern Abyssinia, middle of October, 1865 (herb.

396 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Schz. Bip. in Par.); idem (Flora of Gallabat No. 585}, eodem loco

et tempore (Berl., type material of var. cannabina Schz. Bip.).

Coreopsis Chevalieri 0. Hoffm. & MuschL, based on Aug. Chevalier

9721, is seen from its type specimen (which was very kindly obtained

for my study from the Chevalier Herbarium by Dr. H. Humbert

of Paris) to belong here.

Coreopsis Borianiana var. ft multiplex Sherff, Bot. Gaz. 90:

385. 1930.

A specie involucri bracteis exterioribus biserialibus 18-22, plerum-

que 1.4-1.8 cm. longis quam interioribus saepe longioribus differt.

Type specimen: Collected by "Graf. Zech." under No. 168,

Kete Kratschy, Togo, October 21, 1898 (Berl.).

Distribution: Known only from type locality in Togo.

Specimens examined : Zech. 168 (type, Berl.).

84. Coreopsis togensis Sherff, Bot. Gaz. 76: 87. 1923.

Herba perennis, caule glabro, rubro-viridi, sulcato, subtetragono,

usque ad 2 m. alto. Folia petiolata petiolis 0.5-4 cm. longis, petiolo

adjecto 10-18 cm. longa et usque ad 20 cm. lata, pinnatim 3- vel

5-partita, foliolis lanceolatis, apice saepe longe attenuatis, grosse

vel moderatim serratis dentibus acriter apiculatis infra pallidioribus,

margine minute ciliatis, aliter glabris, lateralibus usque ad 11 cm.

longis et 3 cm. latis, terminali usque ad 16 cm. longo et 4 cm. lato.

Capitula pauca, longe pedunculata pedunculis maximam partem

glabris usque ad 18 cm. longis, radiata, pansa ad anthesin 7-9 cm.

lata et circ. 1.5 cm. alta. Involucrum non plerumque pubescens

nisi basi, bracteis exterioribus 7-10, magnis, foliaceis, lanceolatis,

apice angustatis, interdum ciliatis, 1-2 (et etiam usque ad 4) cm.

longis, interiores lanceolatas aequantibus, vel multo superantibus.

Flores ligulati 7-9, lutei, ligula late elliptico-lineares vel ovato-

oblanceolati, apice minute vel grosse profundeque (sinu usque ad

1 cm. alto) dentati, 3-4.5 cm. longi et 0.7-2 cm. lati. Paleae tenuiter

lineari-oblongae pluribus lineis percursae apice subacutae, omnino

plus minusve flavidae, 1-1.5 cm. longae. Disci florum stigmata

angusto-elongata. Achaenia obcompressa, brunneo-atra, oblongo-

linearia, plerumque alata, margine antrorsum ciliata, supra faciebus

et apice erecto-hispida, corpore 8-10 mm. longa et (alis inclusis)

2.5-3 mm. latis, biaristata aristis nudis vel basim versus antrorsum

1- vel 2-spinulosis, circ. 1 mm. longis.

REVISION OF THE GENUS COREOPSIS 397

Type specimen: Collected by R. Buettner, No. 142, on steppe

behind Ketzabenki, Togo, August 29, 1890 (Berl., 2 sheets).

Distribution: Togo.

Specimens examined : Buettner 142 (Berl., 2 type sheets) ; W.Busse

3416, at alt. of about 700 meters, Hausberg, near Misahoehe, Togo,

December 7, 1904 (Berl., 2 sheets); Kling 60, Jege, Bismarckburg,

Togo, 1889 (Berl.); idem 166, eodem loco, October 30, 1889 (Berl.).

On the type specimens the exterior involucral bracts are much

larger than in Busse's specimens, being twice the length of the inner

ones. The type has also larger leaves and ligulate florets, some of

the ligules reaching 2 cm. in width, but all the other characters are

too much alike to warrant specific or probably even varietal separa-

tion of the Busse plants. The achenial measurements are taken from

Busse's plants, as the achenes of the type are immature.

85. Coreopsis scabrifolia Sherff, Bot. Gaz. 76: 86. 1923.

Herba simplex vel parce ramosa, caule quadrangulato, glabro,

sulcato, perenni e basi lignea, verisimiliter circ. 1 m. alto, non

robusto. Folia nunc sessilia, nunc petiolata petiolis alatis et elongatis

usque ad 4 cm. longis, petiolo adjecto 8-15 cm. longa et 1-3 cm. lata,

lanceolata (raro oblanceolato-linearia), indivisa (raro pauca pinnatim

lobata), acriter et interclum grosse dentata, utrinque acuminata vel

tantum acuta, scabra, minute spinuloso-ciliata, infra ad venas

sparsim piloso-hispida et pallidiora. Capitula pauca (1-4), sub-

tenuiter pedunculata pedunculis maximam partem glabris 3-12 cm.

longis, radiata, pansa ad anthesin 5-6 cm. lata et circ. 1 cm. alta.

Involucri bracteae exteriores 8-11, lineares vel lineari-lanceolatae,

dorso basi hispidae, 1-1.5 cm. longae, interiores lanceolatas ae-

quantes vel paulo superantes. Flores ligulati circ. 8, flavi, ligula

elliptici vel ovato-lanceolati, 2-2.5 cm. longi et 5-12 mm. lati.

Paleae tenuiter lineari-oblongae, circ. 5-7-purpureo- vel brunneo-

striatae, circ. 1 cm. longae. Disci florum stigmata incrassata ter-

minaliter caudata. Achaenia (alis inclusis) brunneo-atra, late linea-

ria, valde obcompressa, moderatim sed inconspicue alata, antrorsum

setosa, corpore 9-11 mm. longa et (alis inclusis) 2-3 mm. lata,

biaristata, aristis nudis vel sursum 1-3-spinulosis, 1.5-2.5 mm. longis.

Type specimen: Collected by T. Kassner, No. 2776, under trees,

Kundelungu, Belgian Congo, May 15, 1908 (Berl.).

Distribution: Southeastern Belgian Congo.

Specimens examined: S. De Giorgi, vicinity of Elisabethville,

Belgian Congo, 1923 (Bruss.); Homble 329, bordering stream,

398 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

eodem loco, May, 1912 (Brass., 2 sheets; nom. indig., musombe');

Kassner2776 (type, Berl.: cotypes, Brit., 2 sheets).

The general habit is deceivingly like that of Bidens Baumii (0.

Hoffm.) Sherff, and, were it not for the achenial and involucral

differences, distinction between the two would be at times difficult.

86. Coreopsis palmata Nutt. Gen. Amer. 2: 180. 1818. Calli-

opsis palmata Spreng. Syst. Veg. 3: 611. 1826. Coreopsis pauciflora

Lehm. Ind. Sem. Hort. Hamb. 1833; cf. Linnaea 10: Litt. 76. 1836.

Coreopsis praecox Fresen. Linnaea 13: Litt. 93. 1839; cf. Ind. Sem.

Hort. Bot. Francofurt. 1838.

Herba perennis, erecta, 5-9 dm. alta, caulibus ad basim hori-

zontalem radicantibus glabris vel ad nodos hirsutis, simplicibus

vel paulo ramosis, tetragonis, foliosis. Folia opposita, principalia

(summa plerumque indivisa exclusa) petiolata petiolis planis alatis

0.4-2.5 cm. longis et 3-6 mm. latis, petiolo adjecto 2-7 cm. longa,

rigida, crassiuscula, suberecta, pedato- vel palmato-tripartita seg-

mentis integris vel irregulariter 2-3-lobatis, segmentis (vel lobis)

ultimis petiolo similibus oblongo-linearibus supra atque infra glabra

marginibus scabrido-ciliatis apice saepius obtusis vel subrotundatis,

plerumque 3-6 mm. latis. Capitula plerumque 1-3 (rarius usque

ad 6), breviter pedunculata pedunculis glabratis 1-4 cm. longis,

radiata, pansa ad anthesin nunc 2.5-3.5 nunc etiam 4-6 cm. lata,

0.9-1.5 cm. alta. Involucri bracteae exteriores 8-12, late lineari-

oblongae, pallidae, obtusae, tergo glabrae, marginibus scabrido-

ciliatae, 5-9 mm. longae, interdum subbiseriatim dispositae, quam

interiores oblongo-obovatae irregulariter (et praecipue superne)

ciliolatae paulo breviores. Flores ligulati circ. 8, flavi, ligula oblongo-

obovati, apice plus minusve rotundato obsolete dentati, 1.5-2.7

cm. longi. Paleae angustissime oblongo-lineares (filiformes), supra

moderate dilatatae, apice acutae, demum ±6.5 mm. longae. Florum

tubulosorum stylorum rami anguste conico-appendiculati. Achaenia

valde obcompressa, elliptico-oblonga, basim versus attenuata,

glabra, subnigra, anguste alata, apice alis productis breviter biden-

tata, omnino 5-6.5 mm. longa et 1.8-2.3 mm. lata.

Type specimen: No particular specimen cited. Habitat given by

Nuttall as "on the open plains of the Michigan Territory, Illinois,

and Lower Louisiana."

Distribution: Wisconsin and eastern Manitoba southwardly to

Indiana, Missouri, and Oklahoma.

Specimens examined: J. A. Allen, Iowa (Field); C. F. Baker, St.

Croix Falls, northwestern Wisconsin, August 15, 1900 (Gray);

REVISION OF THE GENUS COREOPSIS 399

C. R.Barnes, Converse, Missouri, June 30, 1877 (Field); M. S.Bebb,

prairies, Winnebago County, Illinois, September, 1859 (Gray) ; idem,

prairies, Marion County, Illinois, 1860 (Field); Robert Bebb 50,

Clarke, Indiana, 1894 (Field); idem 1048, open hillside, Lauderdale,

Wisconsin, July 19, 1902 (Field) ; H. C. Benke 3967, Crystal Lake,

Illinois, June 30, 1923 (Field); 0. W. Blakley 1451, in woods near

Page, Oklahoma, June 20, 1914 (Gray); E. Bourgeau, Winnipeg

Valley, Canada, 1859 (Berl.); Fred Brendel, Peoria, Illinois (Field);

idem, Illinois (Berl.); Buckley, Illinois (Gray); 5. F. Bush 160, Mon-

tier, Missouri, June 30, 1894 (Berl.; Gray); idem 8249, prairies,

Purdy, Missouri, August 17, 1905 (Gray); /. W. Clokey 2428, dry

prairie soil, Niantic, Illinois, June 29, 1915 (Field; Gray) ; George L.

Clothier & H. N. Whitford 1037, prairies, Brown County, Kansas,

July 30, 1897 (Gray); Robert Combs & C. R. Ball 517, Ames, Iowa,

June 25, 1897 (Berl.; Gray); John M. Coulter, field near LaPorte,

Indiana, July 22, 1878 (Field); idem, Kankakee River, Indiana,

1882 (Field, 2 sheets) ; R. I. Cratty, Iowa (Field) ; idem, Armstrong,

Iowa, July 18, 1897 (Gray); idem 16, Emmet County, Iowa, 1879

(Field); Arthur W. DeSelm 275 and 276, south of Monee, Illinois,

July 6, 1913 (Field) ; idem 394, highways near Otto, Illinois, July 19,

1913 (Field);/1. W. Dewart, St. Cloud, Minnesota, September, 1892

(Mo.) ; H. Eggert, St. Louis, Missouri, June 29, 1875 (Berl., 2 sheets) ;

idem, dry ground, eodem loco, June 18, 1886 (Gray) ; idem & Sydow,

prairie and dry hills, eodem loco, July 1, 1874 (Gray) ; W. H. Emig

146, along roads near Duroc, Missouri, July 2, 1913 (Mo.); ex herb.

T. J. & M. F. L. Fitzpatrick, common, woods and prairies, Allamakee

County, Iowa, June 27, 1895 (Field); W. D. Frost, Willmar, Minne-

sota, July, 1892 (Berl.); Frank C. Gates 598, Ravenswood, Illinois,

July 17, 1905 (Field); idem 3148, Zion City, Illinois, July 19, 1909

(Field); H. A. Gleason, in the black oak association, Shirland,

Illinois, June 29, 1908 (Gray); idem, bunch-grass prairies, Milroy,

Illinois, July 6, 1908 (Gray) ; idem 326, Forsythe, Illinois, June 10,

1896 (Gray) ; Elihu Hall, dry, open timber lands, Athens, Illinois,

1861 (Field, 2 sheets); J. R. Heddle 698, Madison, Wisconsin,

August 1, 1907 (Field); W. Hoffman, Corn Creek, Ozark Region,

Missouri, June 21, 1870 (Berl., 2 sheets); J. M. Holtzinger, Lamoille,

Minnesota, June, 1895 (Field) ; E. W. D. Holway, prairies, Decorah,

Iowa, July 16, 1888 (Field); Hortus Lipsiae, ex Illinois (Berl.);

D. Houghton, 200 miles above Falls of St. Anthony, upper Mississippi

River (Gray); idem, dry prairie of upper Mississippi River above

Falls of St. Anthony, July, 1832 (N.Y.); 0. E. Lansing, Jr. 307,

400 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

prairie soil, Hammond, Indiana, June 28, 1898 (Field); idem 2797,

roadside, Chicago, Illinois, July 19, 1910 (Field; Gray); idem 3113,

alt. 445 meters, fallow field east of Mansfield, Missouri, June 5-12,

1911 (Field; Gray); /. A. Lapham, Milwaukee, Wisconsin (Gray);

Leavenworth, Louisiana (Gray) ; ex Lehmannio ipso sub nom. Coreop-

side pauciflora (Gray; N.Y.); Ray N. Lloyd, prairies, Ravenswood,

Illinois, July 11, 1887 (Field) ; W. H. Manning, Lake City, Minne-

sota, August 13, 1883 (Gray) ; F. E. McDonald, dry prairies, Peoria,

Illinois, June, 1891 (Field); idem, eodem loco, 1900-1901 (Field);

idem, eodem loco, June, 1904 (Gray); idem, eodem loco, June, 1915

(Field); Mead, Illinois, 1850 (Gray); Edgar A. Mearns 144, Fort

Snelling, Minnesota, July 16, 1888 (Field); R. T. Morgan, vicinity

of Fountaindale, Illinois, August, 1877 (Berl.);E. J. Palmer 18078,

dry, open woods, near Ironton, Missouri, June 26, 1920 (Gray);

Harry N. Patterson, prairies near Oquawka, Illinois, July (Field);

idem, near Oquawka, June 20, 1872 (Field); A. S. Pease, dry soil

south of Champaign, Illinois, October 9, 1920 (Gray); idem 12394,

Champaign, September 11, 1909 (Gray) ; Donald C. Peattie 8, moist

prairie, East Chicago, Indiana, July 31, 1920 (Field); Ns. Riehl

41, hills, St. Louis, Missouri, June, 1838 (Berl.); Benjamin L. Robin-

son, Normal, Illinois, July, 1887 (Gray) ; C. 0. Rosendahl 656, Spring

Grove, Minnesota, June 30, 1902 (Gray) ; J. H. Sandberg, Hennepin

County, Minnesota, August, 1889 (Field, 2 sheets); idem, copses,

eodem loco, July, 1890 (Field) ; Savage & Stull 60, Ironton, Missouri,

June 16, 1897 (Field) ; J. H. Schuette, St. Anthony Park, Minnesota,

July 2 and 14, 1888 (Field); idem, eodem loco, July 8, 1888 (Field;

Gray); idem, St. Anthony, Minnesota, July 16, 1888 (Field); idem,

Plover, Wisconsin, July 26, 1888 (Gray) ; idem, Green Bay, Wiscon-

sin, August 13, 1899 (Gray) ; idem, Green Lake, Wisconsin, August

13, 1899 (Field) ;E. P. Sheldon, Sleepy Eye, Minnesota, July, 1891

(Field) ; idem, Fort Snelling, Minnesota, June, 1895 (Field) ; Earl E.

Sherff 284, St. Louis, Missouri, July 7, 1910 (Gray); idem 1642, dry

sandbarrens west of Kankakee, Illinois, August 3, 1912 (Field);

idem 1781, dry hill, Elgin, Illinois, August 27, 1912 (Field, 2 sheets);

H. C. Skeels 375, Gougar's Prairie, Joliet, Illinois, July 14, 1904

(Field) ; Ernest C. Smith 510, roadsides, Hinsdale, Illinois, September

2, 1902 (Field); G. W. Stevens 2713, open woods in mountain valley

near Page, Oklahoma, September 9, 1913 (Gray); J. T. Stewart,

bluffs around Peoria, Illinois, 1867 (Field) ; E. B. Uline, Fish Lake,

LaPorte County, Indiana, July, 1891 (Field); idem, Mishawaka,

Indiana, July 1, 1891 (Field, 2 sheets) ; L. M. Umbach, prairies, Du

Page, Illinois, July 6, 1898 (Berl.; Field; Gray).

REVISION OF THE GENUS COREOPSIS 401

87. Coreopsis verticillata L. Sp. PL 907. 1753. Chrysanthe-

mum Marianum, etc. Pluk. Mant. pi. 344, /• -4- 1769. Coreopsis

tenuifolia Ehrh. Beitr. 7: 168. 1792; Schkuhr Bot. Handb. edit. 2.

pi. 260b, fig. ad dextr. 1808. Coreopsis verticillata var. 0 tenuifolia

Michx. Fl. Bor. Amer. 2: 139. 1803. Bidens verticillata (L.) Baill.

Hist. PI. 8: 305. 1886.

Herba perennis, glabra, caulibus e basi plus minusve horizontali

erectis, 5-9 dm. altis, subangulatis, superne ramosis. Folia opposita,

sessilia, 3-6 cm. longa, laminis palmatim 3-partitis, segmentis 1-2-

pinnatis, ultimis lineari-filiformibus, acriter apiculatis, 0.3-1 mm.

latis. Capitula saepius pauca, saepe corymbosa, tenuissime pedun-

culata pedunculis glabris vel sparsissime pilosis, radiata, pansa ad

anthesin 2.5-5 cm. lata et 6-9 mm. alta. Involucri bracteae exteriores

circ. 8, oblongae vel oblongo-lineares, 4-6 (raro -9) mm. longae, sub-

glabrae, apice saepius obtusae, quam interiores elliptico-oblongae

minutissime pubescentes plerumque paulo breviores. Flores ligulati

circ. 8, flavi, ligula elliptico-oblongi, apice integri vel minute denticu-

lati, 1.2-2.5 cm. longi. Paleae lineares vel filiformes, superne latiores,

apice acutae, ±4 mm. longae. Flores tubulosi flavi, pubescentes,

stylorum ramis apice cuspidato-appendiculatis vel acriter conicis.

Achaenia plana, oblongo-obovata, faciebus glabris subatra, margini-

bus anguste flavido-alata, apice exaristata vel (alis parce productis)

vix bidentatis dentibus laceratis, omnino 3-5 mm. longa et 1-1.7

mm. lata.

Type specimen: No type cited by Linnaeus, although Virginia

was given for the habitat. Linnaeus' first citation of literature is

that of Gronovius, Virg. 105, whence the type would be understood

to be a plant collected by John Clayton in Virginia (as then bounded)

and perhaps still extant (Brit.).

Distribution: Maryland to Florida, Alabama, and Arkansas.

Reported by some authors as extending to Kentucky and Nebraska.

Probably not native farther north, although frequently cited for

Ohio, Michigan, Canada, etc. (cf. Torr. & Gray, Fl. N. Amer. 2:

342. 1843; Gray, Syn. Fl. N. Amer. 1, pt. 2: 293. 1884). Occasionally

escapes from gardens.

Specimens examined: Anon. 576, Arkansas, September, 1835

(Berl.); anon., cult., June, 1818 (Berl.); W. W. Ashe, Chapel Hill,

North Carolina, July, 1898 (Field); M. S. Bebb, District of Columbia,

1863 (Field; Gray); ex Biltmore Herb. 437a, open pine woods,

Spartanburg, South Carolina, June 17, 1897 (Field; Gray); Walter

Blair, dry ridges, Prince Edward County, Virginia, June, 1881

402 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

(Gray) ; S. F. Blake 9542, in colony, sandy field, Washington, District

of Columbia, July 11, 1926 (Pom.); ex herb. J. W. Blankinship,

Southern Pines, North Carolina, July 18, 1895 (Gray); Buckley,

southeastern United States (Gray); Agnes Chase 2382, in woods,

Riverdale, Maryland, June 22, 1904 (Field); eadem 2474, rocky

woods, West Chevy Chase, Maryland, July 12, 1904 (Field); A. H.

Curtiss, Virginia (Field); idem, Bedford County, Virginia, June 10,

1871 (Gray) ; Earl Jerome Grimes 3768, border of woods beside rail-

way, vicinity of Williamsburg, Virginia, June 16, 1921 (Gray);

ex herb. D. Gronovii (Berl. ; cotype?) ; James Hall, Virginia, 1828-

1834 (Field) ; R. M. Harper 5, alt. 345 meters, rocky bank of Town

Creek on Sand Mountain, near Chavies, Alabama, November 24,

1905 (Gray; Mo.); A. A. Heller 933, about Suffolk, Virginia, June

8-13, 1893 (Field, 3 sheets; Gray); Theodor Holm, open thickets,

Brookland, District of Columbia, June, 1894 (Gray); Hort. Berol.,

cult., July 25, 1815 (Berl.); idem, cult., September, 1834 (Berl.);

idem, cult., 1837 (Berl.); H. D. House 150, sandy woods, Takoma

Park, District of Columbia, August 3, 1904 (Kew) ; G. MacCarthy,

District of Columbia (Field); E. L. Morris 105, sandy roadsides

along Potomac River, from Chain Bridge to opposite Cabin John

Bridge, Virginia, July 1, 1899 (Field); Thomas C. Porter, 40 miles

south of Lynchburg, Virginia, July 17, 1880 (Field) ;L.F. & Fannie

R. Randolph 150, dry, sandy soil, sterile hillside field, vicinity of

Lorton, Virginia, June 14, 1922 (Gray) ; Arthur Schott, near George-

town, District of Columbia, August, 1857 (Field) ; idem, woods near

Georgetown, June 22, 1860 (Field); Earl E. Sherff 253, cult., St.

Louis, Missouri, July 6, 1910 (Field); idem 5004, cult., Harvard

Univ. Bot. Gard., August 12, 1929 (Field); John K. Small, alt.

75 meters, on banks of Little Long Creek, Albemarle, North Carolina,

August 22, 1894 (Field); idem & A. A. Heller 372, between Gold

Hill and Falls of the Yadkin, Stanley County, North Carolina,

August 18, 1891 (Field, 3 sheets) ; E, S. Steele, Washington, District

of Columbia, August, 1889 (Pom., 2 sheets).

88. Coreopsis pulchra Boynt. in Small, Fl. S. E. United States

1277, 1340. 1903.

Herba perennis, erecta, glaberrima, 4-7 dm. alta, perspicue

foliosa, caulibus moderate angulatis, plus minusve fastigiato-ramosis.

Folia opposita, sessilia, principalia 2.5-4.5 cm. longa, palmatim

circ. 3-partita foliolis mox subpalmatimque 2-3-partitis, segmentis

ultimis anguste oblongeque linearibus, integris, 1-nervatis, mem-

branaceis, eciliatis, apice obtusis, 2-4 cm. longis et 0.5-1.5 mm.

REVISION OF THE GENUS COREOPSIS 403

latis. Capitula numerosa, corymbosa, tenuiter pedunculata pedun-

culis saepius 4-10 mm. longis, radiata, pansa ad anthesin 3-5 cm.

lata et circ. 9-11 mm. alta. Involucri bracteae exteriores 8-10,

lineari-oblongae, superne saepe sensim vel subabrupte latiores,

apice obtusae, glabrae, 5-8 mm. longae; interiores ovatae, apice

pubescentes, plerumque tantum 4-6 mm. longae. Flores ligulati

6-8, intense flavi, ligula elliptico-oblongi vel anguste obovati,

apice integri vel vix denticulati, 1.6-2.4 cm. longi. Paleae anguste

lineares, apice saepius dilatatae, ±6 mm. longae. Flores tubulosi

corollis purpureo-brunnei, stylorum ramis apice acriter conicis.

Achaenia obcompressa, oblongo-obovata, anguste alata, apice corona

humili lacerataque ornata, circ. 4 mm. longa.

Type specimen : Biltmore Herb., No. 14728 of the former Biltmore

Herbarium. Specimen was collected in "northern Alabama," but

doubtless was destroyed, along with many others, by the flood waters

at Biltmore.

Distribution: In region of the Lookout Mountain, northeastern-

most Alabama, and perhaps in adjacent Georgia.

Specimens examined: T. G. Harbison, cult., Highlands, North

Carolina, August 1, 1905 (Gray, 2 sheets) and August, 1906 (Gray) ;

Charles Mohr, Mentone, Alabama (N.Y., 2 sheets); idem, open,

rocky woods, Mentone, near the Falls, Lookout Mountain, September

18, 1892 (U.S.); idem, damp rocks, DeSoto Falls, near Mentone,

September 12, 1898 (U.S.); Albert Ruth 612, sandy soil, Lookout

Mountain, DeKalb County, Alabama, July, 1898 (N.Y.); idem 624,

near DeSoto Falls, Lookout Mountain, July, 1898 (U.S.).

89. Coreopsis Delphinifolia Lam. Encycl. 2: 108. 1786; Bot.

Mag. pi. 156, sub nom. C. verticillata. 1791. Ceratocephalus Delphinii

foliis Vaill. Hist. Acad. roy. sci. 1720: 328. 1722; Ehret PI. et Pap.

Rar. Depict, pi. 9, f. 1749. C. verticillata var. linearis Michx. Fl.

Bor. Amer. 2: 138. 1803 (fide Torr. & Gray, Fl. N. Amer. 2: 342.

1843). C. discolor Link, Enum. Hort. Berol. 2: 353. 1822.

C. Delphiniifolia Lam. ex Ind. Lond. 2: 294. 1930 (sphalm).

Foliorum segmenta magna ex parte 2-3.5 mm. lata.

C. Delphinifolia sensu stricto.

Foliorum segmenta magna ex parte 0.3-2 mm. lata . var. /3 chlooidea.

Herba perennis, caulibus erectis, gracilibus, tetragonis, glabris

vel ad nodos hispidis, ad summam ramosis, 6-15 dm. altis, per totam

longitudinem foliosis, internodiis folia vix aequantibus. Folia

opposita, sessilia, suberecta vel parce patentia, 5-9 cm. longa, summa

404 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

saepe indivisa integraque, alia palmatim 3-partita, foliolo mediano

plerumque rursus in 3-5 segmenta diviso, foliolis lateralibus simplici-

bus vel raro 2-partitis, segmentis ultimis linearibus membranaceis

acutis glabris integris 1-6 (plerumque 2-3.5) mm. latis. Capitula

corymbosa, tenuissime pedunculata pedunculis glabris 2-7 cm.

longis, pansa ad anthesin 3-5 cm. lata et circ. 8-10 mm. alta. Invo-

lucri glabri bracteae exteriores 8-10, oblongo-lineares, obtusae, 4-6

mm. longae, quam interiores ovatae nunc vix longiores nunc vix

breviores. Flores ligulati circ. 8, flavi, ligula anguste elliptico-

oblongi, apice integri vel parce dentati, 1.3-2.5 cm. longi. Paleae

filiformes, ad summam paulo dilatatae, achaenia superantes. Flores

tubulosi corolla brunnei, stylorum ramis apice tenuiter caudatis.

Achaenia obcompressa, circumambitu (alis brunneis 0.5-1 mm. latis

inclusis) elliptico-oblonga vel obovata, 4.5-6 mm. longa et (alis

inclusis) 2-4 mm. lata, glabra, corpore subnigra, apice minute 2-

dentata dentibus fimbriatis.

Type specimen: The species was founded directly upon Cerato-

cephalus Delphinifolius Vaill. as illustrated by Ehret (loc. cit.) and

supported by the material cultivated in the King's Garden at Paris.

The native habitat was given as Virginia.

Distribution: Virginia to Georgia and Alabama.

Specimens examined: Liberty Hyde Bailey, cult., Ithaca, New

York, July 24, 1935 (Field; sub nom. C. microphylla); ex Biltmore

Herb., sub num. 2064A, rare in moist soil, Eastman, Georgia,

June 4, 1900 (N.Y.); A. H. Curtiss 6825, low ground near Millen,

Georgia, July 1, 1901 (Gray); herb. Drs. A. C. Hexamer & F. W.

Maier, wet pine woods, Summerville, South Carolina, May 27,

1855 (Gray); Hort. Berol (Berl.); Hort. Cantabr., 1842 (Gray);

Hort. Lipsiae (Berl.); Hort. Paris., 1820 (Berl.); Michaux, North

America (Berl.); Earl E. Sherff 5003, cult, in Harvard Univ. Bot.

Gard. (Hort. Cantabr.), August 12, 1929 (Field); John K. Small, in

and about the Altamaha River Swamp, Liberty County, Georgia,

June 18-21, 1895 (Field); J. T. Stewart, near Charleston, South

Carolina, June 1, 1865 (Field); K. A. Taylor, pine barren, Summer-

ville, South Carolina, June 10, 1891 (Field).

Coreopsis Delphinifolia var. /3 chlooidea Sherff, Bot. Gaz.

94: 596. 1933.

E specie foliorum segmentis magna ex parte tantum 0.3-2

(nonnullis usque ad 3.2) mm. latis differt.

REVISION OF THE GENUS COREOPSIS 405

Type specimen: Collected by Samuel Bottsford Buckley, in pine

woods, mountains of Alabama (Gray).

Distribution: Georgia and Alabama.

Specimens examined : Buckley, in pine woods, etc. (type, Gray) ;

Roland M. Harper 545, dry pine barrens, Sumter County, Georgia,

August 31, 1900 (Field).

90. Coreopsis major Walt. Fl. Carol. 214. 1788. C. senifolia

Michx. Fl. Bor. Amer. 2: 138. 1803. C. maior Walt, ex Ind. Lond.

2: 295. 1930.

Folia simplicia var. e Oemleri.

Folia principalia tripartita.

Foliola 1-3 (rarius -5) mm. lata var. 8 linearis.

Foliola plerumque 5-25 mm. lata.

Foliola plerumque 5-11 mm. lata var. 7 rigida.

Foliola latiora, plerumque 1-2.5 cm. lata.

Foliola plus minusve pubescentia C. major sensu stricto.

Foliola glabra var. 0 stellata.

Herba perennis, e basi horizontali erecta, gracilis, supra suberecte

ramosa ramis tenuibus, 5-9 dm. alta, caule ramisque angulatis plus

minusve pubescentibus. Folia opposita, sessilia, divaricata vel sub-

erecta, summa simplicia integraque, alia tripartita, segmentis nunc

subrhomboideo-ovalibus nunc elliptico-lanceolatis vel etiam lanceo-

latis (rarissime lineari-lanceolatis), membranaceis, integris, plus

minusve pubescentibus, ciliatis, apice obtusis, terminali 4-7 cm.

longo et 1.2-2.5 cm. lato, lateralibus plerumque paulo minoribus.

Capitula subcorymbosa tenuiter pedunculata pedunculis glabris

vel sparsim hispidis saepius 2-8 cm. longis, radiata, pansa ad anthesin

2.7-4 ( -5) cm. lata et 7-9 mm. alta. Involucri plus minusve hispidi

bracteae exteriores circ. 8, lineari-oblongae, superne saepe dilatatae,

apice obtusae, 4-8 mm. longae, interiores ovato-oblongae vel lance-

olato-ovatae subaequales. Flores ligulati circ. 8, flavi, ligula oblongo-

lineares vel clliptico-oblongi, apice integri vel denticulati, 1.5-2.5

cm. longi. Paleae lineares, supra paulo dilatatae, achaenia multo

superantia. Florum tubulosorum stylorum rami acriter conici.

Achaenia valde obcompressa, elliptico-oblonga vel obovata, sub-

nigra, glabrata, alata alis subbrunneis 0.2-0.7 mm. latis, omnino

4.5-6 mm. longa et 2-4 mm. lata, apice alis productis breviter biden-

tata vel plurisetosa.

Type specimen: No specimen cited. The remnants of Walter's

collection, however, are in London (Brit.).

406 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Distribution: South Carolina and Kentucky southwardly to

Georgia and Mississippi.

Specimens examined: C. F. Baker, Blount Springs, Alabama,

May 5, 1898 (Pom.); S. B. Buckley, Alabama, July, 1841 (Mo.);

ex Cambridge Bot. Gard., Cambridge, Massachusetts, August, 1845

(Berl.); A. H. Curtiss 6471, dry thickets near Stone Mountain,

Georgia, June 12, 1899 (Gray); John Davis 1855, Stone Mountain,

Georgia, June 18, 1921 (Gray; forma foliis glabratis); A. Dedrick,

dry, sandy soils (Field) ; F. S. Earle 2060, Auburn, Alabama, May 28,

1899 (Field, 2 sheets); idem 2061, eodem loco, May 28, 1899 (Pom.);

idem & C. F. Baker, eodem loco, August 17, 1897 (Pom.); H. Eggert,

Sherwood, Tennessee, June 8, 1897 (Field); ex herb. Hexamer et

Maier, along fences, Abbeville County, South Carolina, June,

1855 (Gray); Miss Anna King 116, dry border of woods, Mammoth

Cave, Kentucky, July 2, 1916 (Field) ; B. F. Leeds, Chattanooga,

Tennessee, September 29, 1892 (Field) ; Dr. Mohr, Mobile, Alabama

(Field); Philip A. Munz 1589, dry, open woods, near Montgomery,

Alabama, June 10, 1917 (Pom.); Josephine Skehan 137, Ocean

Springs, Mississippi, June, 1892 (Gray; forma var. stellatae adpro-

pinquans) ; John K. Small, alt. 300-420 meters, on northwest slope of

Stone Mountain, Georgia, July 3, 1893 (Field) ; John Donnell Smith,

grassy pine woods, Orangeburg District, South Carolina, August 9,

1884 (Field).

Coreopsis major var. /3 stellata (Nutt.) Robins. Rhodora 10:

68. 1908. C. stellata Nutt. Journ. Acad. Phila. 7: 76. 1834. C.

senifolia var. stellata (Nutt.) Torr. & Gray, Fl. N. Amer. 2: 342. 1843.

Glabra, foliolis plerumque angustioribus, saepe longioribus, sae-

pius rhomboide lineari-lanceolatis, terminali usque ad 1 dm. longo

(rarissime monstroseque trifido vel tripartito).

Type specimen: No particular specimen cited.

Distribution: Virginia to southernmost Ohio and southwardly to

Florida and Alabama.

Specimens examined: Anon. (Mt. Desert Nurseries), Bar Harbor,

Maine, August, 1906 (Gray); Samuel M.Bain 95 p.p., sandy hills,

Henderson, Tennessee, July, 1893 (Gray); H. C. Beardslee & C. A.

Kofoid, alt. 900 meters, sandy woods, Swain, North Carolina,

August, 1891 (Gray); Beyrich, in thickets, Georgia (Berl.); ex

Biltmore Herb. 175a and 6, light soil in upland woods, North Carolina,

July 7, 1897 (175a, Field; Gray: 175b, Gray); J. R. Churchill, woods,

Lookout Mountain, near Chattanooga, Tennessee, June 2, 1911

REVISION OF THE GENUS COREOPSIS 407

(Gray) ; John Davis 2097, dry, open woods, Tryon, North Carolina,

June 29, 1920 (Gray); idem 7818, dry woods, near Anderson, South

Carolina, August 30, 1917 (Mo. ; forma var. rigidae adpropinquans) ;

A. H. Curtiss, Bedford County, Virginia, July 1, 1868 (Field); idem,

eodem loco, June 30, 1871 (Gray, 2 sheets; forma nonnullorum

foliorum foliolis tripartitis) ; idem, eodem loco, July 3, 1872 (Field) ;

idem 1487, alt. 600 meters, rocky woods. Chilhowee Mountains,

Tennessee, June (Berl., 2 sheets; Field); F. S. Earle & C. F.

Baker, Auburn, Alabama, May 28, 1898 (Field) ; George Engelmann,

Norristown, eastern Tennessee, August 27, 1876 (Mo.); Juliet

Fauntleroy 654, near Huddleston, Virginia, August 9, 1914 (Gray);

Gates, Alabama (Gray); Asa Gray & John Carey, mountains of

'Virginia and North Carolina, July, 1841 (Gray; var. rigidae adpro-

pinquans); James Hall, Montgomery County, Alabama, 1828-1834

(Field) ; T. G. Harbison, Waynesville, North Carolina, 1897 (Gray) ;

idem, Highlands, North Carolina, 1905 (Gray); Roland M. Harper

46, alt. 205 meters, dry oak woods, Athens, Georgia, June 25, 1900

(Berl.; var. rigidae adpropinquans); idem 131, alt. 205 meters,

eodem loco, July 3, 1900 (Gray); A. Arthur Heller 99, vicinity of

Salisbury, North Carolina, June 21, 1890 (Field, 2 sheets); idem

1013, alt. 540 meters, near Hickory, North Carolina, June 23, 1893

(Field; var. rigidae adpropinquans); Hooker, Kentucky (Berl.);

Hortus Cantabr., cult., 1842 (Gray); idem, cult., 1873 (Gray); ex

Hort. Lipsiae (Berl.); Arthur H. Howell 808, Anniston, Alabama,

June 10, 1913 (U.S.); F. W. Hunnewell, 2nd, dry woods, White

Sulphur Springs, West Virginia, July 4-6, 1914 (Gray); T. H.

Kearney, Jr., White Cliff Springs, Tennessee, July 14, 1894 (Field,

2 sheets); B. F. Leeds, Chattanooga, Tennessee, September 29,

1892 (Field, 2 sheets) ; E. E. Magee, Hayward County, North Caro-

lina, September, 1897 (Gray); idem, Waynesville, North Carolina,

September 9, 1897 (Gray); idem, alt. 750 meters, Highlands, North

Carolina, September 2, 1902 (Gray) ; idem, eodem loco; below Satulah

Mountain, August 2, 1902 (Gray);E.L. Morris 1050, alt. 620 meters,

shaded river bank near Spanishburg, West Virginia, July 23, 1900

(Field) ; Thomas C. Porter, middle Georgia, 1846 (Gray) ; L. F. &

Fannie R. Randolph 1066, dry, sterile soil, edge of woods, southeast

of Granite Falls, North Carolina, July 30, 1922 (Gray, var. rigidae

adpropinquans); Riddell, Kentucky (Gray);BenjaminL. Robinson 79,

Asheville, North Carolina, August 2, 1893 (Berl.; Gray);F. Rugel,

Florida, 1844 (Gray; Mo.); ex herb. Margaret P. Russell, Asheville,

North Carolina (Gray); Albert Ruth, sandy woods, Knoxville, Ten-

nessee, June, 1893 (Field); idem, gravelly soil and rocky places,

408 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Mount Nebo, eastern Tennessee, July, 1893 (Mo., 2 sheets); idem,

open woodlands, Knoxville, June, 1895 (Berl.) ; idem, woods, Lookout

Mountain, Georgia, July 20, 1897 (Gray) ; Earl E. Sherff 5007, cult.,

Harvard Univ. Bot. Gard., August 12, 1929 (Berl.; Del.; Field);

W. L. Sherwood, Waynesville, North Carolina, July, 1897 (Gray);

idem, Tuckasiegee Valley, North Carolina, September 20, 1897

(Gray); B. Shimek, Monte Sano, Alabama, July, 1891 (Field);

C. W. Short, hilly lands, Kentucky (Field) ; Howard Shriver, Wythe-

ville, Virginia, July, 1877 (Berl.; Field); John K. Small, alt. 900

meters, on Round Top Mountain, west of Seven Mile Ford, Smyth

County, Virginia, July 2, 1892 (Field; Gray); idem, alt. 660 meters,

on Farmer Mountain, on New River, Carroll County, Virginia, July

12, 1892 (Field); idem, alt. 750 meters, Nick's Creek, base of Pine

Glade Mountain, Smyth County, Virginia, August 5, 1892 (Field);

idem & A. Arthur Heller, alt. 1,350 meters, Green's Hill, near Blowing

Rock Mountain, Watauga County, North Carolina, July 10, 1891

(Field); iidem, between Cranberry and Linville, North Carolina,

July 17, 1891 (Field) ; iidem, southern slopes of Grandfather Moun

tain, North Carolina, July 25, 1891 (Field) ; iidem, alt. 1,050 meters

Rocky Knob, near Blowing Rock Mountain, Caldwell County

North Carolina, July 24, 1891 (Field; Gray); iidem 328, Blowing

Rock Mountain, Watauga County, North Carolina, July 29, 189!

(Field, 3 sheets); iidem (similiter) 328, eastern slopes of Blowing

Rock Mountain, Caldwell County, North Carolina, July 31, 189"

(Field, 3 sheets); John Donnell Smith, Warm Springs, Madison

County, West Virginia, July 29, 1880 (Field); idem, Caesar's Head

South Carolina, July, 1881 (Field); Roland Thaxter, Cullowhee

North Carolina, June-July, 1887 (Gray); S. M. Tracy 9049, Warm

Springs, Georgia, May 20, 1905 (Berl.; Field; Gray); G. R. Vasey

northern Alabama, 1878 (Field) ; idem, North Carolina, 1878 (Gray)

William C. Werner 92, Ironton, Ohio, 1892 (Gray) ; Karl M. Wiegand

& W. E. Manning 3414, sandy bank by roadside, 4 miles west oj

Raleigh, North Carolina, June 27, 1927 (Pom.); iidem 3415, edge of

dry pine woods, 12 miles south of Asheboro, North Carolina, June

30, 1927 (Pom.}; Charles Wright, northern Georgia (Gray).

Coreopsis major var. 7 rigida (Nutt.) F. E. Boynton in Small,

Fl. S. E. United States 1276, 1340. 1903. C. senifolia var. rigida

Nutt. Gen. Amer. 2: 180. 1818. C. Wrayi Nutt. Journ. Acad. Phila.

7: 76. 1834. C. senifolia Hook. Bot. Mag. 63: pi 3484- 1836 (non

Michx.). C. Delphinifolia var. rigida (Nutt.) Torr. & Gray, Fl. N.

REVISION OF THE GENUS COREOPSIS 409

Amer. 2: 342. 1843. C. rigida Nutt. ex Torr. & Gray, loc. cit. ; cf. Ind.

Kew. 1: 617. 1895.

Folia rigidula, tripartita, plerumque suberecta, foliolis plerumque

lineari-lanceolatis, faciebus saepissime glabris marginibus scabridis,

saepius 3-7 cm. longis et 5-11 mm. latis.

Type specimen: No type cited, but habitat was given as Georgia.

Distribution: Virginia and Tennessee southwardly to Georgia.

Specimens examined: Anon., ex Nutallio ipso, loc. ignoto (Gray,

sub nom. Coreopside Wrayi Nutt.); Harley Harris Bartlett 1638,

pine barren, vicinity of Thomson, Georgia, July 23, 1909 (Pom.);

William M. Canby, Florence, South Carolina, July 3, 1878 (Field);

H. Eggert, dry woods on Yellow River, Gwinne County, Georgia,

July 27, 1897 (Mo.) ; Roland M. Harper, sunny, rocky hillside, vicin-

ity of Sawdust, Georgia, June 18, 1927 (Gray; Mo.); Hort.

Cantabr., cult., 1849 (Gray) ; C. G. Lloyd, near Rockwood, Tennessee,

August 10, 1880 (Field); C. Mohr, dry pine woods, Bladon (Bladen),

Alabama, July, 1875 (Field); H. W. Ravenel, Santee Canal, South

Carolina, July, 1846 (Gray); Albert Ruth, dry, sandy soil, Chil-

howee Gap, Tennessee, July, 1893 (Mo.) ; idem, sandy soil, Knoxville,

Tennessee, May, 1896 (Mo.); H. Shriver, Wytheville, Virginia, 1875

(Field); John K Small & A. Arthur Heller, north of Hickory, North

Carolina, June 25-26, 1891 (Field); Huron H. Smith 2526, Blue

Ridge Mountains, Fannin County, Georgia, July 26, 1909 (Field).

Coreopsis major var. 6 linearis Small, Bull. Torr. Bot.

Club 22: 48. 1895.

Parce inconspicueque pubescens, caule 3-6 dm. alta. Folia tri-

partita segmentis linearibus, 3-11 cm. longis et 1-3 (rarius -5) mm.

latis utrinque acuminatis. Capitula etiam ligulae achaeniaque paulo

minora.

Type specimen: Collected by John Kunkel Small, at altitude of

300-330 meters, on Little Stone Mountain, DeKalb County, Georgia,

July 7, 1893 (N.Y., 2 sheets).

Distribution: South Carolina and Central Georgia.

Specimens examined: J. K. Small, Little Stone Mountain, etc.

(type, N.Y., 2 sheets); John Donnell Smith, grassy pine woods,

Orangeburg District, South Carolina, August 9, 1884 (Field; Gray).

Coreopsis major var. e Oemleri (Ell.) Britton, Mem. Torr.

Bot. Club 4: 131. 1894. C. Oemleri Ell. Sketch Bot. S. Carol. &

Georgia 2: 435. 1824.

Var. stellatae similis sed foliis simplicibus differt.

410 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

Type specimen: Collected by Oemler, near the junction of the

Broad and Saluda rivers, vicinity of Columbia, South Carolina.

Distribution: North Carolina to Georgia.

Specimens examined : None.

Asa Gray (Syn. Fl. N. Amer. 1, pt. 2: 294. 1884) treated this as

"the abnormal entire-leaved form" of var. stellata. An examination

of the specimens at Gray Herbarium shows no material of the

"entire-leaved form," however, and it is probable that Gray was

guided largely by the inscription on A. H. Curtiss's label for a

sheet of normal var. stellata at Gray Herbarium. This has the

observation, "with simple leaves not uncommon." Gray's treat-

ment was followed essentially by Robinson and Fernald (Gray's

New Manual, ed. 7. 838. 1908). It may be noted, too, that Boynton

(in Small, Fl. S. E. United States 1276. 1903) made the var. Oemleri

to include var. stellata. Boynton's complete omission of the entire-

leaved plants from his description, however, may well lead to the

suspicion that he had unintentionally overlooked them. Small

(Bull. Torr. Bot. Club 22: 48. 1895) describes "a peculiar state"

of C. major having "leaves undivided, there being two opposite and

entire leaves at each node in place of the normal three-parted ones."

Clearly, he referred to plants of var. Oemleri. He states further:

"It seems to have been first collected by Dr. and Mrs. Britton at

Black Mountain Station, North Carolina. Later Mr. Heller secured

it near Salisbury, N.C., and last season (1893) I came upon it at the

western base of Stone Mountain, Georgia." Apparently the variety

is somewhat less distinct than the other varieties of C. major (even

they, however, pass from one into another by various transitional

forms; cf. A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 294. 1884), but seems

to bear a relation to its parent species comparable to that of var.

Smithii to Coreopsis tripteris.

91. Coreopsis tripteris L. Sp. PI. 908. 1753; Hook. Bot. Mag.

pi. 3583. 1837. Anacis tripteris (L.) Schrank in Miinch. Math, et

Nat. 5: 7 (fide DC. Prodr. 5: 568. 1836). Chrysostemma tripteris

(L.) Less. Syn. Compos. 227. 1832 (fide DC. loc. cit). Coreopsis

tripteris var. /3 Torr. & Gray, Fl. N. Amer. 2: 341. 1843. Coreopsis

tripteris vars. Deamii et intercedens Standl. Rhodora 32: 34. 1930.

Folia divisa.

Foliorum segmenta subrhomboidea var. 7 subrhomboidea.

Foliorum segmenta late elliptico-linearia vel anguste oblongo-

lanceolata C. tripteris sensu stricto.

Folia simplicia var. /3 Smithii.

REVISION OF THE GENUS COREOPSIS 411

Herba perennis, e basi horizontal! erecta, plerumque glabra rarius

pubescens, plus minusve pallida, 1-3 m. alta, caulibus teretibus

summam versus ramosis. Folia opposita, principalia petiolata petiolis

0.5-3 cm. longis, petiolo adjecto 6-12 cm. longa, 3-partita, foliolo

mediano saepe rursus 3-partita, segmentis late elliptico-linearibus vel

anguste oblongo-lanceolatis, apice acutis vel subobtusis, marginibus

aegre revolutis, faciebus glabris vel rarius pubescentibus; folia summa

minora, saepe simplicia et subsessilia. Capitula (contusa odorata)

subcorymbosa, tenuiter pedunculata pedunculis saepius 3-8 cm.

longis, radiata, pansa ad anthesin 3-5 cm. lata et 8-10 mm. alta.

Involucri saepius glabri bracteae exteriores circ. 8, oblongo-lineares,

obtusae, circ. 2-3 mm. longae, interiores oblongo-ovatae 4-6 mm.

longae. Flores ligulati plerumque 7 vel 8, flavi, ligula plus minusve

elliptico-oblongi, apice subintegri vel dentati, 1.2-2.4 cm. longi.

Paleae filiformes, apicem versus dilatatae, striis purpureis percursae,

5-7 mm. longae. Flores tubulosi primo plus minusve flavi demum

brunnei vel purpurei, stylorum ramis apice breviter caudato-appendi-

culatis. Achaenia brunneo-nigra, obcompressa, faciebus glabra,

circumambitu (alis angustis subbrunneis inclusis) cuneato-oblonga

vel cuneato-obovata, omnino 5-7 mm. longa et 2.8-4.2 mm. lata,

apice emarginato setis minimis erectis munita.

Type specimen: No particular type was cited. The habitat was

given as Virginia and the first cited reference was the Coreopsis foliis

subternatis integerrimis of the Hort. Upsal. 269. Other citations were

in the following order: Rudbeckia foliis compositis integris, Roy.

lugdb. 181; Chrysanthemum virginianum, folio acutiore laevi trifoliatof.

anagyridis folio, Moris. Hist. 3. p. 21. s. 6. t. 3. f. 44; Raj. Suppl. 215.

Distribution: Massachusetts, southern Ontario, and Wisconsin,

southwardly to Georgia, Mississippi, western Louisiana, and eastern

Kansas.

Specimens examined: H. C. Beardslee & C. A. Kofoid, alt. 600

meters, sandy banks, Great Smoky Mountains, Swain County,

North Carolina, August, 1891 (Field; Gray); M. S. Bebb, Fountain-

dale, Illinois (Gray); Florence Beckwith 54, Dallas City, Illinois,

September, 1917 (Field); H.C.Benkel841, Elgin, Illinois, September

8, 1916 (Field) ; Biltmore Herb. 173a, moist grounds and bordering

streams, Biltmore, North Carolina, August 1, 1897 (Gray; Pom.);

0. W. Blakley 3420, open woods near Page, Oklahoma, August 27,

1914 (Mo.); J. W.Blankinship, Greene County, Missouri, September

7, 1888 (Gray); Fred W. Brendel, Peoria, Illinois (Field); A. B.

Burgess 433, railroad bank, Flowerfield, Michigan, September 30,

412 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

1903 (Field); idem 513, shrubbery at roadside, Portage, Michigan,

October 5, 1903 (Field); W. W. Calkins 135, prairie, Berwyn, Illinois

(Field); idem 145, common, Berwyn, Illinois, August 26, 1907

(Field) ;S.H.& D. R. Camp, dry ground, Jackson County, Michigan,

August 18, 1897 (Field); iidem, Jackson County, Michigan, August

18, 1898 (Field); Agnes Chase, open woods, Rogers Park, Illinois,

August 27, 1896 (Field); Virginius H. Chase, dry prairie north of

Wady Petra, Illinois, August 12, 1896 (Field); J. R. Churchill, shore

of Lake Michigan, Indiana, September 4, 1893 (Pom.); John Davis,

near LaGrange, Missouri, June 9, 1915 (Mo.); C. C. Deam & H. A.

Gleason, dry upland woods, Bluffton, Indiana, September 1, 1904

(Gray); Arthur DeSelm 499, highways south of Tucker, Illinois,

August 7, 1913 (Field); idem 621, highway near Kankakee, Illinois,

September 7, 1913 (Field) ; W. M. Dick, Lorain County, Ohio, July

31, 1895 (Gray); S. S. Dickey 96, dry roadside, Warren County,

Pennsylvania, August 25, 1923 (Gray); Charles K. Dodge, near

Port Huron, Michigan, August 4, 1895 (Field); C. W. Duesner,

Miller, Indiana, 1908 (Field) ; F. S. Earle, creek bottoms, Auburn,

Alabama, September 4, 1899 (Field); H. Eggert, prairie and dry

hills, St. Louis, Missouri, August 25, 1874 (Gray); idem, woods,

St. Louis, Missouri, August 23, 1887 (Gray); idem 122, central

United States, August 30, 1875 (Berl.) ; James Galen 4%89, woods

and thickets, eastern United States, 1883 (Gray) ; Frank C. Gates 808,

Edgewater, Illinois, August 23, 1905 (Field); idem 10104, prairie,

Carthage, Illinois, September 15, 1916 (Field); H. A. Gleason, in

strip of original prairie, east of Rantoul, Illinois, October 5, 1907

(Gray); idem 812, prairie soil along roadsides, southwest of Dal ton

City, Illinois, August 22, 1899 (Gray); idem 1813, dry upland woods,

Grand Tower, Illinois, August 28, 1900 (Gray); idem 1977, prairie,

Illinois, September 4, 1900 (Gray); idem 2753, dry upland woods,

Herod, Illinois, August 23, 1902 (Gray); Grassly, Miller, Indiana,

August 31, 1880 (Field); J. M. Greenman 544, Tygart Junction,

West Virginia, September 24, 1904 (Field; Gray) ; idem 3991, Webster

Grove, Missouri, September, 1920 (Mo.); Dr. Hale, Louisiana

(Gray); Elihu Hall, barrens, etc., Athens, Illinois, 1861 (Field;

Pom.); idem, eodem loco, August, 1864-1867 (Field); idem, eodem

loco, August, 1868 (Field); Roland M. Harper 1954, alt. 50 meters,

bank of Flint River, near Albany, Georgia, August 27, 1903 (Berl.;

Field; Gray); A. A. Heller, vicinity of Salisbury, North Carolina,

August 21, 1890 (Field); idem, near mouth of Tucquan Creek,

Lancaster, Pennsylvania, September 1, 1890 (Gray); idem & E.

413

Gertrude Halbach, near McCall's Ferry, Pennsylvania, September

9, 1893 (Field); A. A. & E. G. Heller 4129, alt. 90 meters, near

Texarkana, Arkansas, August 23, 1898 (Field; Gray); Albert S.

Hitchcock, wet ground, Iowa City, Iowa, 1888 (Calif.); Hort. Bero-

linensis, cult., September 11, 1876 (Berl.); ex Hort. Goetting. (Berl.);

Hort. Lipsiae, cult. (Berl.) ; Hort. Bot. de I'Ecole de Medecine de Paris,

August, 1820 (Berl.); J. W. Huett, Ottawa, Illinois (Gray); T. H.

Kearney, Jr. 553, Clear Creek, Bell County, Kentucky, September,

1893 (Field; Gray; Pom.; etiam in Field qua varietati Smithii

adpropinquans) ; Miss Anna King 534, dry banks or in prairie land,

Glenwood, Illinois, September 20, 1918 (Field) ; eadem 566, dry soil

in thickets of waste field, eodem loco, September 20, 1918 (Field) ;

W. Krebs, Cleveland, Ohio (Berl.); 0. E. Lansing, Jr. 428, open

woods, Chicago, Illinois, August 14, 1898 (Field); idem 647, prairie

land, West Pullman, Illinois, July 27, 1899 (Field); idem 1646,

sandy woods near dunes, near Porter, Indiana, September 16, 1902

(Field); idem 2832, border of oak woods, Gibson, Indiana, August

13, 1910 (Field; Gray); idem 3822, sandy roadsides, Starved Rock,

Illinois, September 7-8, 1914 (Field); idem 3911, open, sandy woods,

Pine, Indiana, August 22, 1915 (Field) ; Ray N. Lloyd, Ravenswood,

Illinois, August 31, 1887 (Field) ; John Macoun, thickets, Sandwich,

Ontario, August 4, 1892 (Gray) ; F. E. McDonald, open, dry woods,

Peoria, Illinois, August, 1900 (Field) ; idem, eodem loco, September,

1901 (Field) ; idem, frequent in rich woods, eodem loco, August, 1904

(Gray); Michaux, North America (Berl.); Charles F. Millspaugh

3865, railroad ditch, Lakeside, Berrien County, Michigan, September,

1914 (Field); W. S. Moffatt 492, thickets, Pine, Indiana, August 31,

1895 (Field; Gray) ; Albert Hanford Moore 2552 and 2553, sandy east

bank of Tygart's Valley River, near Tygart Junction, West Vir-

ginia, September 24, 1904 (Gray); E. L. Moseley, Oxford, Ohio,

August 21, 1895 (Field) ;L. H. Pammel 621, Ames, Iowa, September

10, 1897 (Gray); H. N. Patterson, dry bluffs, Henderson County,

Illinois, August, 1871 (Field); Donald C. Peattie 164, sandy fields,

Tremont, Indiana, August 13, 1920 (Field); idem 2029, Griffith,

Indiana, August 15, 1925 (Field); Thomas C. Porter, banks and

islands of Susquehanna River, Lancaster County, Pennsylvania,

August 22, 1861 (Gray); Miss Carrie A. Reynolds, Stony Island,

Illinois, August 24, 1909 (Field); Riddell, Ohio (Gray); Robert

Ridgway 60, Olney, Illinois, September 7, 1917 (Field); Benjamin

L. Robinson, rich prairie soil, Hendrix, Illinois, August 31, 1904

(Gray); idem 56, Bloomington, Illinois, September, 1893 (Berl.);

414 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

F. Rugel, in valley of Blue Ridge Mountains, North Carolina,

August, 1841 (Berl.); Albert Ruth 69, woods, Lookout Mountain,

Georgia, July 19, 1897 (Gray) ; H. P. Sartwell, Maumee River, Ohio

(Field); Earl E. Sherff 1038, near Jefferson Barracks, Missouri,

August 11, 1910 (Field) ; John K. Small, north of Marysville, Penn-

sylvania, August 15, 1888 (Field); idem, vicinity of mouth of Tuc-

quan Creek, Lancaster County, Pennsylvania, September 1, 1890

(Field); idem, eodem loco, October 11, 1890 (Field); idem, alt. 480

meters, in canyon at Tallulah Falls, Rabun County, Georgia,

August 4, 1893 (Field) ; idem, alt. 900 meters, about Estatoah Falls

on Mud Creek, Rabun County, Georgia, August 12, 1893 (Field);

idem & A. A. Heller 100, vicinity of Faith Post Office, North Caro-

lina, August 14, 1891 (Field); Frank Thone 232, Starved Rock,

Illinois, May-September, 1921 (Field); S. M. Tracy 8670, Taylor-

ville, Mississippi, August 20, 1903 (Field; Gray); L. M. Umbach,

woods, Clarke, Indiana, August 28, 1897 (Field) ; idem, eodem loco,

August 20, 1898 (Field) ; G. R. Vasey, Lookout Mountain, Tennessee,

1878 (Field) ;L.F. Ward, Potomac flats, Maryland, August 12, 1877

(Field); Roscoe J. Webb, dry soil, Pippin Lake, Portage County,

Ohio, September 6, 1909 (Gray) ; E. F. Williams, vacant lots, Boston,

Massachusetts, September 3, 1903 (Gray); Wilbur H. Wright 221,

Chicago, Illinois, August 22, 1908 (Field).

A tall species of attractive appearance and wide distribution

over the eastern and central United States. Commonly glabrous,

but at times becoming more or less pubescent. Torrey and Gray

(Fl. N. Amer. 2: 341. 1843) designated a pubescent form ("leaves

minutely scabrous-puberulent. . . . Western Louisiana, Dr. Veatch")

as "var. /3." During the four decades following, however, Asa

Gray had many opportunities to see the inconstancy of the pubes-

cence character in C. tripteris. Hence, in his Synoptical Flora of

North America (1, pt. 2: 294. 1884) he modified the description of

C. tripteris from "smooth and glabrous" to read "smooth and

glabrous, or leaves minutely pubescent." Some years ago I myself,

in a purely tentative way, treated the pubescent extremes as con-

stituting a variety, which variety was set off with a proper name

in my manuscript. With the examination of hundreds of herbarium

and field specimens, however, it became apparent that the pubes-

cence character was too capricious to admit of utilization, and that

Gray's reduction of a variety thus segregated, to synonymy, would

needs be accepted. Recently Standley (loc. cit.), apparently over-

looking the treatments by Torrey and Gray and by Gray, has

415

described the var. Deamii to include forms with leaves pubescent at

least underneath and involucres hairy; also the var. inter cedens to

include forms with leaves pubescent at least underneath and

involucres glabrous.

Coreopsis tripteris var. 0 Smithii Sherff, Bot. Gaz. 88: 301.

1929.

E specie foliis omnibus vel fere omnibus integris, lamina tenuiter

oblongo-lanceolata differt.

Type specimen: Collected by John Donnell Smith in low, open

woods near Montgomery, Alabama, August 26, 1885 (Field).

Distribution: Kentucky and Arkansas, southwardly to Florida,

Georgia, and Mississippi.

Specimens examined: Ex herb. A. W. Chapman, Georgia (Mo.);

T. H. Kearney, Jr., along Clear Creek, Bell County, Kentucky,

September, 1893 (Field); George V. Nash 2585, River Junction,

Gadsden County, Florida, September 5, 1895 (U.S.);#. J. Palmer

8468, moist, open ground, Malvern, Arkansas, September 4, 1915

(Mo.); Charles Louis Pollard 1222, Waynesboro, Mississippi, August

8-9, 1896 (Field) ; John Donnell Smith, low, open woods, etc. (type,

Field; cotypes, Gray; Mo.).

According to Smith's additional note on the type sheet, Asa Gray

had pronounced this plant a new variety of Coreopsis tripteris and,

in a letter written in November, 1885, had given it a varietal name.

While Gray would probably have published the name had he not died

shortly afterward, his proposed name was passed over by me in

accordance with the recommendations of the Vienna Rules (Recomm.

XIV. e) and the variety named in honor of the type collector.

Coreopsis tripteris var. 7 subrhomboidea Sherff, Bot. Gaz.

88: 303. 1929.

Folia tripartita foliolis lateralibus lanceolatis terminali rhom-

boideo-lanceolato 1.7-2.3 cm. lato, petiolo adjecto 6-8 cm. longa.

Capitula minora, achaeniis tantum 4-4.5 mm. longis.

Type specimen: Collected by Ernest Jesse Palmer, No. 29421,

sandy, open woods bordering bog, near Texarkana, Bowie County,

Texas, October 27, 1925 (Gray).

Distribution: Known only from type locality in Bowie County,

Texas.

Specimens examined: Palmer 29421 (type, Gray).

416 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

In the more than two hundred collections of the species proper

studied by me, all the leaflets were variously linear, oblong-linear,

or narrowly oblong-lanceolate, and the achenes were commonly

5-6 mm. long. In the Palmer plant the terminal leaflets approach

very distinctly a rhombus in outline and the achenes are smaller,

measuring only about 4-4.5 mm. in length.

92. Coreopsis latifolia Michx. Fl. Bor. Amer. 2: 137. 1803.

Leiodon latifolium (Michx.) Shuttl. Distrib. Rugelii PI., Herb. Berol.,

etc.

Herba perennis, erecta, gracilis, glabra vel pubescens, 1-1.5

m. alta, summam versus erecte ramosa, foliosa usque vel fere usque

ad summam. Folia opposita, subsessilia vel brevi-petiolata, omnino

plerumque 1-2.3 dm. longa et 4-10 cm. lata, valde membranacea,

ovata vel ovalia, basi late cuneata apice graciliter acuminata, mar-

gine ciliata ac serrata dentibus mucronatis. Capitula pauca vel

numerosa, paniculato-corymbosa, tenuiter pedicellata pedicellis

2-10 cm. longis, radiata, pansa ad anthesin ±4 cm. lata et ±1.2

cm. alta. Involucri bracteae exteriores 6-9, perspicue herbaceae,

lineari-oblongae, apice obtusae, patentes, 6-8 mm. longae; interiores

oblongae, apicem versus abrupte angustatae, paulo longiores.

Flores ligulati 4 vel 5, flavidi, ligula elliptico-oblongi, apice integri

vel paululum dentati, circ. 1.5-1.8 cm. longi. Paleae lineares,

obtusae, ±8 mm. longae. Flores tubulosi pauci (circ. 12), flavidi

vel demum parce brunnei, stylorum ramis ad apicem conicis. Achae-

nia obcompressa, oblongo-lanceolata, exalata, apice angusto truncata

ac calva, circ. 7 mm. longa.

Type specimen: Collected by Andre Michaux at higher altitude

in the Carolina Mountains (Par.).

Distribution: North Carolina southward to Georgia.

Specimens examined: Anon., mountains of North Carolina,

September, 1843 (Gray); Biltmore Herb. 5374b, slopes of Craggy

Mountain, Buncombe County, North Carolina, August 12, 1897

(Gray; N.Y.); Buckley, mountains of Carolina (Gray) and Georgia

(N.Y.); herb. A. W. Chapman, mountains of Georgia (N.Y.);

Asa Gray, Hickory Nut Gap, southeastern United States, 1843

(N.Y.); idem & W. S. Sullivant, mountains of Carolina, 1843 (Gray);

Harvard Univ. Bot. Gard., cult, anno 1849 e sem. e plantis in monti-

bus Carolinae lectis (Gray); Hort. Berol., cult. (Berl.); F. Rugel,

on summits of mountains, Broad River, North Carolina, July, 1841

(Berl.; N.Y.; appellata Leiodon latifolium Shuttl.).

417

93. Coreopsis tinctoria Nutt., Journ. Acad. Phila. 2: 114. 1821.

Calliopsis bicolor Reichenb. Mag. Aesthet. Bot. pi. 70. 1823. Dip-

losastera tinctoria (Nutt.) Tausch, Hort. Canalius fasc. 1, icon, et

descript. 1823. Calliopsis tinctoria (Nutt.) DC. Prodr. 5: 568. 1836.

Coreopsis tinctoria var. /3 atropurpurea Hook. Bot. Mag. pi. 3511.

1836. Coreopsis elegans Hort. fide Sieb. & Voss, Vilmor. Blumeng.

ed. 3. 1. 487. 1894.

Among the numerous illustrations of this widely cultivated

species we may cite the following: Bart. Fl. Amer. Sept. pi. 45.

1822; J. Radius, Schrift. Naturforsch. Gesellsch. Leipz. 1: pi. 4-

1822; Bot. Mag. pi. 2512. 1824; Bot. Reg. pi. 846. 1824; Sweet,

Brit. Flow. Gard. 1: pi. 72. 1824.

Capitula pansa ad anthesin 2-3 cm. lata, achaeniis 1.2-4 mm. longis.

C tinctoria sensu stricto.

Capitula pansa ad anthesin 1-1.5 cm. lata, achaeniis 0.9-1.4 mm.

longis var. ft imminuta.

Herba annua, erecta, glabra, 6-12 dm. alta, caulibus subtetra-

gonis vel subteretibus, foliosis, multo ramosis ramis plus minusve

angulatis. Folia opposita, subsessilia vel breviter petiolata, saepius

5-10 cm. longa, 1-2-pinnata vel summa indivisa, segmentis (vel

laminis) anguste linearibus vel lineari-lanceolatis. Capitula nu-

merosa, subcorymbosa, tenuiter pedunculata pedunculis (pedicellis)

plerumque 4-10 cm. longis, radiata, pansa ad anthesin 2-3 cm. lata

et 4-6 mm. alta. Involucri glabri bracteae exteriores circ. 8, plus

minusve biseriales atque inter se imbricatae, lineari-oblongae vel

saepius triangulatae, lateribus plerumque scariosae, ±2 mm. longae;

interiores deltoideo-ovatae vel oblongo-ovatae, circ. 5-6 mm.

longae. Flores ligulati circ. 7 vel 8, inferne brunneo-rubri, alibi

flavi, ligula obovati, apice plerumque 3-lobati, 0.7-1.5 cm. longi.

Paleae subfiliformes, superne coloratae atque angustatae, apice

acutae, demum circ. 4-4.5 mm. longae. Flores tubulosi atro-rubri,

stylorum ramis apice obtusis. Achaenia lineari-oblonga vel deorsum

paulo angustata, valde obcompressa, glabra vel glabrata, exalata,

nigra, apice calva, nunc vix 1.2 nunc etiam 3-4 mm. longa.

Type specimen: Collected by Thomas Nuttall, in flooded meadows

on banks of Red River, Arkansas. Should be in London (Brit, or

Kew). I have seen an authentic, fully labeled cotype (Berl.).

Distribution: Minnesota, Saskatchewan, and Washington south-

wardly to Louisiana, Texas, and California; cultivated everywhere

for ornament and escaping frequently. Apparently becoming

established at various places in China.

418 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

Specimens examined: Anon. 575, Fort Gibson, Arkansas, June,

1835 (Berl.); Carrie Barker 31, low ground, Tonka wa, Oklahoma,

June 27, 1908 (Field) ; J. M. Bates 2958, Red Cloud, Nebraska, July

20, 1903 (Gray); Robert Bebb 1174, sandy soil, Galveston, Texas,

May 18, 1903 (Field) ; H. C. Benke 327, Hutchinson, Kansas, October

5, 1918 (Field); idem 4306, Claflin, Kansas, June 22, 1926 (Field);

idem 4657, Murphysboro, Illinois, July 6, 1928 (Field); Katharine

Brandegee, Redstone Park, California, July 25, 1905 (Calif.); N. L.

Britton, Norfolk, Virginia, June 10, 1892 (Field); B. F. Bush 159,

Sheffield, Missouri, June 27, 1895 (Berl.); idem 3017, introduced,

Sugar Creek, Missouri, June 22, 1905 (Gray): George D. Butler 14,

sulphate flats, fields, and roadsides, north of Limestone Gap, Okla-

homa, June 18, 1877 (Field); William M. Canby 157, plains of

Manitoba, Canada, August 7, 1897 (Gray); Harley P. Chandler

7010, Rio Hondo, Texas, July, 1913 (Berl.; Gray); H. C. Cheo &

W. F. Wilson 11 p.p., Mo Kan Shan, Province of Chekiang, China,

June 19, 1926 (Gray); iidem 29, cult., eodem loco, June 28, 1926

(Calif., 2 sheets; Gray); Mr. & Mrs. J. Clemens 978, Leon Springs,

Texas, May 29, 1911 (Pom.); Fred Clements 2778, Spencer, Nebraska,

July 25, 1893 (Gray); Royal A. Dixon 17, Huntsville, Texas, June

3-12, 1908 (Field); R. R. Dreisbach 1042, escaped, ash dumps,

Olney, Pennsylvania, July 15, 1922 (Field); Drummond, Texas

(Gray); George Engelmann, cult., St. Louis, Missouri, September,

1850, e sem. neo-mexicanis a Fendlero ad Santam Fe. lectis (Mo.);

Henry Engelmann (Captain Simpson's Expedition), Fort Laramie,

July, 1858 (Mo.); B. W. Everman, Lake Pend d'Oreille, Idaho,

August 7, 1893 (Field) ; August Fendler 397 pro parte, prairie hollows

near Pawnee Fork, New Mexico, September, 1847 (Mo.); Emily

F. Fletcher, Westford, Massachusetts (Gray); Dr. Hale, Louisiana

(Gray); Elihu Hall 344 pro parte, wet places, Hempstead, Texas,

May 28, 1872 (Pom., 3 sheets); idem 345, eodem loco, June 3, 1872

(Field) ;F.L. Harvey, bluffs, Little Rock, Arkansas (Mo.); Heiland,

cult., Lychen, Province of Brandenburg, Germany, August 16, 1882

(U.S.); A. Arthur Heller 11752, along railroad south of Redding,

California, August 30, 1914 (Field; Gray); Albert S. Hitchcock 279

and 666, buffalo wallows, Pawnee County, Kansas, August 19,

1895 (Gray); idem 13943, savanna, Schofield Barracks, Leilihua,

Island of Oahu, Hawaiian Islands, July 5, 1916 (U.S.); Hooker,

Arkansas (Berl.); Hort. Paris., 1823 (Berl.); Marcus E. Jones

29474, north of Corpus Christi, Texas, March 30, 1932 (Pom.);

W. W. Jones 227, Middle Verde, Arizona, July 12, 1922 (Gray);

J. B. Leiberg, moist places, Kootenai County, Idaho, June, 1890

419

(Field); idem, shores of Lake Pend d'Oreille, Idaho, July, 1892

(Field) ; F.Lindheimer 102 p.p., prairies, Galveston Island, Texas, 1843

(Berl.; Gray); idem 548, rocky places, New Braunfels, Texas, May,

1851 (Berl., 2 sheets); idem 897, Texas, May, 1851 (Berl., forma

multis achaeniis tuberculatis) ; John Macoun, Winnifred, Assiniboia,

August 12, 1895 (Gray) ; idem 38, Red Deer Lakes, Canada, July 22,

1879 (Gray); F. E. McDonald, along roadsides near East Peoria,

Illinois, July, 1919 (Field); Albert Hanford Moore 1832, escaped, in

dump by Lake Anthony, Cottage City, Massachusetts, July 16,

1904 (Gray); Nealley, Chenates, western Texas, 1889 (Field);

J. C. Nelson 1802, waste ground, Salem, Oregon, July 30, 1917

(Gray); J. B. Norton 1455, cult., alt. 300-900 meters, Kuliang Hills,

near Foochow, Province of Fukien, China, July-August, 1919 (U.S.);

Thomas Nuttall, flooded meadows, etc. (cotype, Berl.);£". J. Palmer,

cherty barrens, Newton County, Missouri, July 15, 1906 (Gray);

idem 2272, cherty barrens, Joplin, Missouri, June 19, 1909 (Gray) ;

J. C. Parlin, North Berwick, Maine, July 4, 1892 (Gray); Dr.

Pitcher, Arkansas (Gray); Birdie T. Powell, Hastings, Nebraska,

July, 1886 (Field); J. T. Rothrock, cult., Philadelphia, Pennsylvania,

June 25, 1877 (Field); idem 317, alt. 1,500 meters, Arizona, July,

1874 (Field; Gray); T. E. Savage, J. E. Cameron, & F. E. Lenocker,

Spokane, Washington, July, 1898 (Field) ; iidem, The Dalles, Oregon,

August, 1898 (Field) ; A. K. Schindler 335, alt. 1,100 meters, Lu-shan,

Kuling Mountains, Kiangsi, China, July-August, 1908 (Berl.);

J. H. Schuette, cult., Green Bay, Wisconsin, September 14, 1888 and

August 23, 1897 (Field); idem, cult., eodem loco, July 31, 1901

(Gray); idem, cult., eodem loco, August 2, 1901 (Field; forma);

E. P. Sheldon, Pike Island, Minnesota, July, 1895 (Pom.); Sherff 250

pro parte, cult., St. Louis, Missouri, July 6, 1910 (Field) ; idem 499,

cult., eodem loco, July 16, 1910 (Field); idem 507, cult., eodem loco

et tempore (Field; Gray); idem 5000, cult., Chicago, Illinois, July

3, 1929 (Berl.; Calif.; Field; ligulis supra brunneo-rubris infra plus

minusve flavis); idem SOOOa, eodem loco et tempore (Berl.; Brit.;

Calif.; Field; Kew; Mun.);5. Shimek, Oklahoma City, Oklahoma,

July, 1892 (Field); G. W. Stevens 784, waste place along roadside,

Cleo, Oklahoma, June 8, 1913 (Gray); idem 1212, waste place,

Mountain Park, Oklahoma, June 23, 1913 (Gray); A. N. Steward

1510, Kikungshan, Province of Honan, China, July 23, 1925 (Calif.);

Tang Chung Chang & Nong Sing Po 3790, campus of Fukien Christian

University, Foochow, Province of Fukien, China, June 8, 1926

(Calif.); W. F. Thurrow, Hockley, Texas, 1890 (Field); To Rang

Peng, Ts'ang Wai Tak, & Ts'ang Un Kin 936 (Canton Christian

420 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

College No. 12935}, Shiu-chau, Province of Kwangtung, China,

July 18, 1924 (Calif .) ; S. M. Tracy 7902, Weatherford, Texas, May

26, 1902 (Berl.; Field; Gray);L. M. Umbach 2752, waste ground,

Clyde, Illinois, July 14, 1900 (Gray); Van Hermann 2686, cult.,

vicinity of Santiago de las Vegas, Province of Havana, Cuba, May

16, 1905 (Calif.; Field; Gray); L. F. Ward, Ford County, Kansas,

October 5, 1897 (Gray); Mark White 218, Summer County, Kansas,

June 17, 1899 (Mo.); Charles Wright 339 and 341, western Texas,

May-October, 1849 (Gray); Cyril Zeller, Ellis County, Kansas,

1908-1911 (Mo.).

A species often confused with C. basalis var. Wrightii and with

C. Atkinsoniana. From both it is at once distinguished by its almost

oblong, entirely wingless achenes. In cultivation, plants occur with

the rays reddish-brown or brownish-red almost or quite throughout.

These are the var. atropurpurea of Hooker. At times their rays are

distinctly yellow on the under surface and brownish-red above.

Coreopsis bicolor Bosse ex Buchenau (Linnaea 25: 630. 1853,

nomen) doubtless belongs with C. tinctoria.

Coreopsis tinctoria var. ft imminuta Sherff, Bot. Gaz. 94:

594. 1933.

Capitula pansa ad anthesin 1-1.5 cm. lata, bracteis exterioribus

±1 mm. interioribus circ. 5-6 mm. longis, achaeniis 0.9-1.4 mm.

longis.

Type specimen: Collected by John Michael Holzinger, Santa Fe,

New Mexico, August 13, 1911 (Mo.).

Distribution: Known only from type locality of Santa Fe, New

Mexico.

Specimens examined: J. M. Holzinger, Santa Fe, New Mexico

(type, Mo.).

94. Coreopsis basalis (Dietr.) Blake, Proc. Amer. Acad. 51:

525. 1916. Calliopsis basalis Dietr. in Otto & Dietr. Allgem. Garten-

zeit. 3: 329. 1835. Coreopsis diversifolia Hook. Bot. Mag. pi. 3474.

1836 (nee alior.). Calliopsis Drummondii D. Don in Sweet, Brit.

Fl. Gard. ser. 2. 4: pi. 315. 1838. Coreopsis Drummondii (D. Don)

Torr. & Gray, Fl. N. Amer. 2: 345. 1843. Coreopsis Drummondii

var. ft Torr. & Gray, loc. cit. Coreopsis picta Hort. fide Sieb. & Voss,

Vilmor. Blumeng. ed. 3. 1: 487. 1894.

Foliorum segmenta linearia vel vix lanceolato-linearia.

var. ft Wrightii.

REVISION OF THE GENUS COREOPSIS 421

Foliorum segmenta nunc lineari-lanceolata nunc elliptico-oblonga

vel quidem orbiculata C. basalis sensu stricto.

Herba annua, erecta, ramosa, nunc fere glabra nunc pilis pluri-

articulatis valde hispida vel tomentosa, 2-4 dm. alta, caulibus

ramisque angulatis ac sulcatis. Folia opposita, petiolata petiolis

1-5 cm. longis, petiolo adjecto usque ad 12 cm. longa, principalia

1-3-pinnata segmentis lineari-lanceolatis vel elliptico-oblongis vel

quidem orbiculatis, membranaceis, undulato-integris, apice obtusis

vel raro acutis. Capitula ramos (terminaliter per 5-15 cm. plerum-

que nudos) terminantia, radiata, pansa ad anthesin 3-4.5 cm. lata

et ±8 mm. alta. Involucri bracteae exteriores 8-10, saepe patentes,

subulatae vel lineari-lanceolatae, apice plerumque acutae, margini-

bus plus minusve hispido-ciliatae, tergo longitudinaliter plerumque

3-vittatae, 5-9 mm. longae, quam interiores ovatae glabrae paulo

breviores. Flores ligulati circ. 8, magna ex parte flavi sed ad basim

brunneo-rubri, ligula cuneate ac late obovati, 1.3-2.3 cm. longi,

apice 3-lobati lobo mediano rursus 2- (raro 3-) lobulato. Paleae

angustissime lineares, demum contortae et bracteas interiores sub-

aequantes. Flores tubulosi saltern superne atro-rubri, stylorum

ramis apice obtuso-conicis. Achaenia obovata, tergo convexo nigra

ac valde papillata, marginibus incurvatis incrassata cartilagineaque,

1.4-1.8 (-2) mm. longa, apice calva.

Type specimen: Cultivated in Berlin Botanical Garden from

seed said to have come from Missouri. If "Missouri" is correct,

then the plants there must have been likewise in cultivation, since

the species is known in the spontaneous state only in Texas.

Distribution: Native to Texas. Sometimes cultivated for orna-

ment and hence occasionally found elsewhere as an escape.

Specimens examined: E. R. Bogusch 994, Gonzales County,

Texas, June 6, 1926 (Pom.); Mrs. Charles C. Deam 1864, near Lake

Helen, Florida, April 26, 1906 (Gray); T. Drummond 69 (commun.

Hooker anno 1835), Rio Brasses (Brazos River), Texas (Gray);

idem 200 (commun. Hooker anno 1835), Texas (Gray);#. H.Eames

8839, waste ground, Fairfield, Connecticut, August 6, 1914 (Gray);

Elihu Hall, Hempstead, Texas, June, 1872 (Field) ; idem 343, sandy

prairie, eodem loco, May 2, 1872 (Field; Gray; Mo.; N.Y.; Pom., 2

sheets); Harvard Univ. Bot. Gard., cult., 1844 (Gray); Hort. BeroL,

July, 1837 (Berl., 2 sheets) ; J. F. Joor, open hills on Galveston Bay,

Texas, June (Mo.) ; John H. Kellogg, cult, in Missouri Bot. Gard.,

June 30, 1905 (Mo.); F. Lindheimer, Galveston Island, Texas,

November, 1842 (Mo.); idem, Texas, 1849 (Mo.); idem 54, Mill

422 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

Creek, Texas, April-May, 1844 (Mo.); idem 101, on dunes, Galves-

ton Island, May, 1843 (Calif.; Field; Gray; Mo., 3 sheets; N.Y.);

B. Mackensen 76, San Antonio, Texas, May 10, 1911 (Field) ; Manz

29, Victoria, Texas (Berl.) ; ex herb. C. Mohr, damp meadows near

coast, eastern shore of Mobile Bay, Alabama, April, 1868 (Field);

E. J. Palmer 7740, nearly pure sand, Liberty, Texas, May 22, 1915

(Mo.); J. Reverchon, White Creek, Brazos, Texas, July 4 (Berl.);

J. Schneck, La Grange, Texas, May, 1894 (Field); J. H. Schuette,

cult., Green Bay, Wisconsin, September 14, 1888 (Field); Earl E.

Sherff 498, cult., St. Louis, Missouri, July 16, 1910 (Field); idem

5015, cult., Harvard Univ. Bot. Gard., August 12, 1929 (Berl.;

Brit.; Calif.; Field; Kew; Mun.); idem 5018, eodem loco et tempore

(Field; Del.; Mo.; Mus.V.; U.S.); W. F. Thurrow, Hockley, Texas,

1890 (Field); S. M. Tracy 8550, Cameron, Louisiana, July 6, 1903

(Field; Gray; Mo.); idem 8925 p.p., Victoria, Texas, April 27, 1905

(Berl.; Field; Gray; N.Y.); S. R. Warner, Willis, Texas, June 1

(Mo.); Charles Wright, Texas (Gray).

Coreopsis basalis var. ft Wrightii (A. Gray) Blake, Proc.

Amer. Acad. 51: 526. 1916. C. Drummondii var. Wrightii A. Gray,

Syn. Fl. N. Amer. 1, pt. 2: 291. 1884; cf. (typo) A. Gray, PL

Wright. 2: 90. 1853; cf. etiam (descriptione sine typo vel nomine)

A. Gray, PI. Wright. 1: 109. 1852.

E specie tantummodo foliorum segmentis linearibus vel vix

lanceolato-linearibus, capitulis paulo minoribus differt.

Type specimen: Collected by Charles Wright, rocky hills on the

San Pedro, western Texas, 1849 (Gray). Gray's name and descrip-

tion (Syn. Fl. N. Amer. loc. cit.) rest upon the variety given in PI.

Wright. 2: 90. 1853. The treatment in that work, however, gives

first the reference to PI. Wright. 1: 109. 1852, concerning a plant

collected by Wright in western Texas, in 1849; and, second, the

citation of Wright 1237, collected on prairies of the Leona and Nueces,

Texas, May, 1851. Fortunately the two collections were identical.

Distribution: Widely distributed across Texas and reaching

westwardly into New Mexico.

Specimens examined: B. F. Biish 696, common in sand, Dallas,

Texas, May 10, 1900 (Mo.) ; George L. Fisher 5007, Houston, Texas,

May 13, 1917 (U.S.); 5. H. A.Groth 66, limestone hills near Bracken,

Texas, July 8, 1903 (Field; Gray); A. A. Heller 1730, alt. 480-600

meters, Kerrville, Texas, May 7-14, 1894 (Field; Gray; Mo.);

Hort. Cantabr., cult, anno 1852 e sem. a C, Wrightio lectis (Gray) ;

REVISION OF THE GENUS COREOPSIS 423

G. Jermy 657, Iron Creek, Gillespie, Texas (Field; Mo.); Marcus

E. Jones 28012, west of Uvalde, Texas, April 26, 1931 (Pom.); idem

28013, east of Carisso Spring, Texas, April 27, 1931 (Pom.); E. J.

Palmer 12115, rocky hills, San Marcos, Texas, July 1, 1917 (Mo.);'

J. Reverchon, western Texas, 1882 (Gray) ; Miss Ellen D. Schulz 867,

Austin, Texas, July, 1920 (U.S.); Charles Wright, Texas (Gray);

idem 217, New Mexico, 1851 (Gray) ; idem 1237, prairies of the Leona

and Nueces, western Texas, May, 1851 (Gray; Mo.; N.Y.; U.S.).

95. Coreopsis rosea Nutt. Gen. N. Amer. PL 2: 179. 1818.

Calliopsis rosea (Nutt.) Spreng. Syst. Veg. 3: 611. 1826. Conopsis

rosea Nutt. ex Less. Syn. Gen. Compos. 228. 1832 (sphalm). Coreopsis

rosea f. leucantha Fern. Rhodora 21: 171. 1919.

Herba erecta, simplex vel ramosa, glabra, sub terra caulibus

horizontalibus repens, caule supra terram tereti vel superne vix

angulato 2-6 dm. alto. Folia opposita axillis saepe foliosis, basi

ciliata parce connata, 2-5 cm. longa, integra vel irregulariter plus

minusve 2- vel 3-partita, lamina vel segmentis linearibus, 0.5-3 mm.

latis, eciliatis, apice parce acutis. Capitula ramulos tenues apicem

versus nudos (pro pedunculis 3-8 cm. longis) aliter foliosos terminan-

tia, radiata, pansa ad anthesin 2-2.5 cm. lata et 4-6 mm. alta.

Involucri glabri bracteae exteriores 6-10, saepe irregulariter positae,

mox patentes, oblonge lineari-lanceolatae, tantum 1-3 mm. longae;

interiores oblongo-ovatae, basim versus connatae, 4-6 mm. longae.

Flores ligulati circ. 8, rosei vel rarius albidi, ligula cuneato-oblongi,

apice plerumque 3-lobati lobo mediano rursus lobulato, 8-13 mm.

longi. Paleae lineares, 2-4 mm. longae. Flores tubulosi flavidi,

stylorum ramis apice abrupte conico-incrassatis. Achaenia obcom-

pressa, anguste elliptico-oblonga, nigra, exalata, glabra vel papillata,

apice exaristata sed vix poculata, circ. 2 mm. longa.

Type specimen: None cited, but "open grassy swamps, from New

Jersey to Georgia" given as the habitat.

Distribution: South westernmost Nova Scotia; Massachusetts to

Delaware and (fide Nutt.) south to Georgia.

Specimens examined : J. W. Adams 480, Hankens Pond, on Petti-

coat Branch of Maurice River, Millville, New Jersey, August 15,

1926 (Gray); anon., Smithfield, Rhode Island, 1871 (Gray); C. F.

Batchelder, in water at marshy edge of pond, Falmouth, Massa-

chusetts, August 14, 1906 (Gray) ; herb. William Boott, Winter Pond,

Winchester, Massachusetts, July 30, 1876 (Gray) ; J. Bernard Brin-

ton, low grounds, Egg Harbor, New Jersey, September 3, 1888

424 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

(Field); idem, Egg Harbor, August 18, 1889 (Field); William M.

Canby, Felton, Delaware, September, 1867 (Gray); idem, swamps,

Delaware, August, 1877 (Field) ; J. Franklin Collins, Spectacle Pond,

Lincoln, Rhode Island, September 11, 1900 (Gray); idem, M. L.

Fernald, & H. H. York 294, wet sand, margin of Sand Pond, War-

wick, Rhode Island, September 8, 1914 (Berl.; Field); Joseph W.

Congdon, Lincoln, Rhode Island, August 22, 1871 (Field); herb.

Carlton C. Curtiss, Martha's Vineyard, August 13, 1892 (Field);

M. L. Fernald, E. B. Bartram, & B. Long 24687, peaty and cobbly

beach of St. John (Wilson's) Lake, Yarmouth County, Nova Scotia,

July 23, 1921 (Gray); iidem 24688, peaty margin of Goven Lake,

Yarmouth County, Nova Scotia, July 23, 1921 (Gray); M. L.

Fernald, C. H. Bissell, C. B. Graves, B. Long, & D. H. Linder 22859,

gravelly margin, northwest side of Tusket (Vaughan) Lake, Nova

Scotia, August 20, 1920 (Gray}; Fernald &Long 10664, damp, sandy

shore of Loon Pond, Lakeville, Massachusetts, August 26, 1913

(Gray); iidem 10669, damp, sandy borders of ponds west of White

Pond, Chatham, Massachusetts, September 9, 1913 (Gray); iidem

17591, forming a broad belt, wet, sandy lower beach and inundated

margin, Buck Pond, Harwich, Massachusetts, August 30, 1918

(Gray) ; iidem 1 7594, eodem loco et tempore (Gray, 2 sheets) ; iidem

24689, wet, cobbly beach of Vaughan (Tusket) Lake, Nova Scotia,

August 13, 1921 (Gray) ; iidem 24690, wet, peaty margin of Vaughan

Lake at Gavelton, Nova Scotia, August 13, 1921 (Gray); iidem

24691, peaty and cobbly margin of Gilfilling Lake, Yarmouth

County, Nova Scotia, August 23, 1921 (Gray); iidem 24692, peaty

and sandy margin of Salmon (Greenville) Lake, Yarmouth County,

Nova Scotia, August 25, 1921 (Gray) ; iidem & D. H. Linder 22860,

cobbly beach of Tusket (Vaughan) Lake, Gavelton, Nova Scotia,

September 4, 1920 (Gray); iidem 22861, wet, peaty shore, East

Branch of Tusket River, Gavelton, September 4, 1920 (Gray);

iidem 22862, sandy and gravelly beach of Butler's (Gavelton) Lake,

Gavelton, September 4, 1920 (Gray) ; Mrs. E. T. Gibbs, South Pond,

Plymouth, Massachusetts (Gray); Jesse M. Greenman 437, Nine

Mile Pcnd, Cape Cod, Massachusetts, September 5, 1898 (Gray);

H. D. House 9660, Artist's Lake, Middle Island, Suffolk County,

New York, August 14, 1923 (Gray) ; Edwin Hunt, Sudbury, Massa-

chusetts, August 12, 1874 (Field); George G. Kennedy, Orleans,

Massachusetts, July 31, 1891 (Gray); idem, Sharon, Massachusetts,

September 2, 1904 (Gray); L. A. Kenoyer & F. W. Pennell 3224,

margin of Mares Pond, Falmouth, Massachusetts, July 29, 1911

REVISION OF THE GENUS COREOPSIS 425

(Gray); Henry A. Lang, Egg Harbor, New Jersey, September 10,

1905 (Gray) ; E. S. Miller, Long Pond, Wading River, New York,

August 30, 1871 (Gray); idem, Wading River, August 18, 1877

(Field, 2 sheets) ; ex herb. Thomas Morong, pond borders, Winchester,

Massachusetts, August 5, 1881 (Field) ; John Murdoch, Jr., Orleans,

Massachusetts, August 22, 1900 (Field) ; Joseph Murdoch, Rochester,

Massachusetts, August 24, 1902 (Field); Oakes, in sandy, flooded

places, Plymouth, Massachusetts (Gray); C. F. Parker, Bristol,

Pennsylvania, August 11, 1865 (Field; Gray); herb. Margaret P.

(Mrs. Robert Shaw) Russell, Snow's Pond, Rochester, Massachusetts,

August 17, 1918 (Gray); F. C. Seymour 1384, in shallow water,

Seth's Pond, West Tisbury, Martha's Vineyard, Massachusetts,

September 7, 1917 (Gray); Harold St. John 2939, grassy shore of

Long Pond, Southampton, New York, August 18-21, 1920 (Gray);

G. Thurber, Cranston, Rhode Island, August, 1844 (Berl.) ; ex herb.

E. Tuckermann, Jr., New England (Berl.); Emile F. Williams, Catau-

met, Cape Cod, Massachusetts, September 15, 1901 (Gray).

96. Coreopsis Cardaminefolia (DC.) Torr. & Gray, Fl. N.

Amer. 2: 346. 1843. Calliopsis Cardaminefolia DC. Prodr. 5: 568.

1836. Coreopsis Cardaminifolia DC. ex Nutt. Trans. Amer. Phil.

Soc. n. ser. 7: 360. 1841 (sphalm). Coreopsis Cardaminefolia var.

angustiloba Torr. & Gray, loc. cit. Coreopsis Cardaminefolia var.

lineariloba A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 291. 1884.

Herba annua, glabra, erecta, gracilis, suberecte ramosa, 2-5 dm.

alta, caulibus subteretibus, foliosis. Folia opposita, remota, quam

internodia breviora, petiolata petiolis plerumque 1-4 dm. longis

(summa sessilia), inferiora medianaque laminis 1-2-pinnata segmentis

elliptico-oblongis vel late angusteve linearibus, superiora nunc

pinnatim paucilobata nunc simplicia. Capitula saepe numerosa,

tenuiter pedicellata pedicellis glabris 1-5 cm. longis, radiata, pansa

ad anthesin tantum circ. 1.6-2.4 cm. lata et 6-8 mm. alta. Involucri

glaberrimi bracteae exteriores 6-9, plus minusve lanceolatae atque

irregulariter positae, saepe subacutae, marginibus diaphanae, 1-2

mm. longae; interiores ovatae 5-8 mm. longae. Flores ligulati circ.

7 vel 8, ligula oblongo-obovati, guttur versus macula brunneo-

purpurea ornati alibi flavi, apice 3-lobati, ±1 cm. longi. Paleae

filiformes, circ. 3-4 mm. longae. Flores tubulosi minuti, numerosi,

corolla atro-purpurei, stylorum ramis apice obtusissime conicis.

Achaenia elliptico-oblonga, valde obcompressa, faciebus glaberrima

vel minute tuberculata, marginibus anguste vel late alata, circ. 2 mm.

longa, apice calva vel minute bidentata.

426 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Type specimen: The species was founded upon four specimens

("pi. exs. 1668, 1956, 1478, et 2312") collected by Jean Luis Berlandier

at various locations in southeasternmost Texas and adjacent Mexico

(Del.). DeCandolle gave three localities: (1) between Bexar (Bejar)

and Trinity River; (2) at Matamoros, State of Tamaulipas, Mexico;

(3) at Lake St. Nicholas, Corpus Christi Bay.

Distribution: Western Louisiana, Kansas, and Arizona south-

ward to States of Tamaulipas, Coahuila, and Chihuahua, Mexico.

Specimens examined : Berlandier 566, Texas, May 15, 1829 (Mo.);

idem 882, about Matamoros, State of Tamaulipas, Mexico, April,

1831 (Gray); idem 1073, from Matamoros northward to Nueces

River, Texas, April, 1834 (Gray); idem 1956, Texas, May 15, 1829

(Gray); idem 2312, about Matamoros, State of Tamaulipas, April,

1831 (Gray) ; idem 2503, from Matamoros to Nueces River, Texas,

April, 1834 (Gray; Mo.); idem 2580, Texas (Field); R. S. Cocks,

Cameron, Louisiana, July, 1903 (Gray); J. F. Collins 8, wastes,

Providence, Rhode Island, July 30, 1892 (Gray) ; Miss M. B. Croft,

San Diego, Texas, 1885 (Field); Thomas Drummond 68, Brazos

River, Texas (Gray); A. Fendler 897 pro parte, Pawneefork, New

Mexico, August, 1847 (Mo.); idem 441, New Mexico, 1847 (Gray;

Kew) ; J. Gregg 901 , prairie west of Matamoros, State of Tamaulipas,

Mexico, June 5, 1847 (Mo.) ; Dr. Hale, Louisiana (Gray) ; Elihu Hall

344 pro parte, wet places, Hempstead, Texas, May 28, 1872 (Field) ;

Herbert C. Hanson A13, alt. 2,100 meters, wet places, Flagstaff,

Arizona, August 22, 1922 (Field); Harvard Univ. Bot. Gard., cult,

e sem. Texanis anno 1849 (Gray); W. W. Jones 160, Verde Valley,

Arizona, September 9, 1920 (Gray);Lindheimer 102 p.p., Texas, 1843

(Gray); D. T. MacDougal 293, alt. 2,100 meters, vicinity of Flag-

staff, Arizona, July 15, 1898 (Field; Gray); E. W. Nelson 6104,

near Colonia Garcia, in the Sierra Madre, State of Chihuahua,

Mexico, August 25, 1899 (Gray); idem 6813, Sabinas, State of

Coahuila, Mexico, May 21, 1902 (Gray) ; E. W. Olive 72, abundant

in buffalo wallows, near Cinnamon City, Oklahoma, July, 1893

(Field); J. H. Oyster 4014, Kansas, September 7, 1884 (Field);

Edward Palmer 635, States of Coahuila and Nuevo Leon, Mexico,

February-October, 1880 (Gray) ; idem 655, San Lorenzo de Laguna

and vicinity, southwest of Parras, State of Coahuila, May 1-10,

1880 (Field) ; C. G. Pringle 8328, alt. 210 meters, Rio Grande Valley

near Diaz, State of Coahuila, April 25, 1900 (Field; Gray; Pom.);

J. Reverchon 83, western Texas, 1882 (Gray); J. T. Rothrock 157,

alt. 2,010 meters, Zuni Village, New Mexico, July, 1874 (Field;

REVISION OF THE GENUS COREOPSIS 427

Gray) ; B. C. Tharp 5627, causeway, Corpus Christi, Texas, March

16, 1929 (U.S.); J. W. Tourney 594, Flagstaff, Arizona, June 30,

1892 (Gray); Townsend & Barber 150, alt. 2,250 meters, near Colonia

Garcia, in the Sierra Madre, State of Chihuahua, Mexico, July 19,

1899 (Field; Gray); W. F. Thurrow, Hockley, Texas, 1890 (Field);

C. Wright 34.0, western Texas, October, 1849 (Gray); idem 1236,

New Mexico, 1851 (Gray) ; M. S. Young, muddy hollow in the plains,

near Paloduro Canyon, Texas, September 7, 1917 (Mo.).

97. Coreopsis stenophylla Boynt. Biltmore Bot. Studies 1:

141. 1902.

Herba glabra, annua vel vix perennis, 3-4 dm. alta, caulibus

rigidis sed gracilibus, subfastigiate ramosis, omnino foliosis. Folia

opposita, rigidula, erecta vel adscendentia, plerumque quam inter^

nodia longiora, simplicia vel irregulariter pinnata, lamina vel foliolis

filiformibus vel anguste linearibus usque ad 2 mm. latis. Capitula

subnumerosa, tenuissime pedicellata pedicellis ±5 cm. longis, radiata,

pansa ad anthesin 2-3 cm. lata et ±6 mm. alta. Involucri bracteae

exteriores circ. 8, lanceolatae, 1-2 mm. longae; interiores ovatae

5-6 mm. longae. Flores ligulati plerumque 8, flavi, ligula obovati,

apice 3-lobati lobis rotundatis, 1-1.4 cm. longi. Flores tubulosi

atro-rubri, stylorum ramis apice plus minusve truncatis. Paleae

lineares, acutae. Achaenia plana, elliptico-oblonga, 1.5-2 mm.

longa, marginibus anguste alata alis integris, apice bidentata dentibus

brevibus crassis.

Type specimen: Collected for the Biltmore Herbarium, in pine

woods near Hammond, Louisiana, September 17, 1900. The type,

now probably destroyed, was in the Biltmore Herbarium. An

excellent duplicate specimen is extant (N.Y.).

Distribution: Louisiana.

Specimens examined: Ex Biltmore Herb., Hammond, Louisiana,

September 17, 1900 (N.Y., cotype); R. S. Cocks 3631, dry, open,

sandy fields, Shreveport, Louisiana, July, 1907 (N.Y.); S. M. Tracy

3456, Lake Charles, Louisiana, August 9, 1897 (N.Y.); ex herb.

C. L. Willich & F. W. Weiss, loco ignoto (Berl.).

98. Coreopsis similis Boynt. in Small, Fl. S. E. United States

1278, 1340. 1903.

Herba annua, glabra, erecta, 1-3 dm. alta, caulibus sulcatis

striatisque, ramosis. Folia opposita, polymorpha, principalia tenuiter

petiolata petiolis usque ad 7 cm. longis, petiolo adjecto 2-8 (-13) cm.

428 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

longa, simplicia vel pinnatim partita, lamina vel foliolis oblonge

linearibus vel elliptico-oblongis (raro oblongo-ovalibus), apice acutis

vel obtusis, membranaceis, 5-12 mm. latis; summa simplicia vel

pinnata, lamina vel foliolis plerumque angusto-linearibus. Capitula

pauca vel numerosa, subcorymbosa, tenuissime pedicellata pedicellis

±5 cm. longis, radiata, pansa ad anthesin 1.5-2.5 cm. lata et 6-8 mm.

alta. Involucri bracteae exteriores 7-9, lanceolatae vel interdum

ovatae, glabratae, apice obtusae, 1.5-2 mm. longae; interiores ovatae,

4-5 mm. longae. Flores ligulati plerumque 8, supra guttur purpureo-

rubri alibi flavi, ligula obovati, apice 3-lobati lobis rotundatis, ±1 cm.

longi. Flores tubulosi numerosissimi, minuti, corolla atro-purpurei,

stylorum ramis obtusissimo-apiculati. Achaenia plana, orbiculata,

faciebus subnigris glabra, marginibus late alata unica ala diametro

corpus (nunc 2 mm. longum et 0.5 mm. latum, nunc 3 mm. longum

et 1 mm. latum) aequante, apice biaristata aristis tenuibus glabris

externe ad alas plus minusve adgregatis tantum circ. 0.5 mm. longis.

Type specimen: Collected by G. C. Nealley at Brazos Santiago,

Texas, 1889 (U.S.). The year is erroneously given by Boynton

as 1899.

Distribution: Along the Gulf of Mexico coast, Texas.

Specimens examined: W. L. Bray 48, Virginia Point, Texas,

April 16, 1899 (U.S.); A. Arthur Heller 1548, sea level to 12 meters

alt., Corpus Christi, Texas, April 9-12, 1894 (Gray; Mo.; N.Y.);

G. C. Nealley, Brazos Santiago, etc. (type, U.S.; cotype, Field);

Robert Runyon 385, Brownsville, Texas, January, 1923 (U.S.).

99. Coreopsis Atkinsoniana Dougl. in Lindl. Bot. Reg. pi.

1376. 1830. Calliopsis Atkinsoniana (Dougl. in Lindl.) Hook. Fl.

Bor. Amer. 1:311. 1833.

Herba annua vel plus minusve perennis, erecta, glabra, 6-12

dm. alta, ramosa ramis arcuato-adscendentibus vel suberectis,

caulibus angulatis, internodiis saepius elongatis. Folia radicalia

l-3-(saepius 2-) pinnatisecta segmentis lineari-oblongis vel anguste

elliptico-lanceolatis terminali multo majore petiolo 2-8 cm. longo,

1-2 dm. longa; caulina plerumque 1-2-pinnatisecta segmentis

anguste linearibus vel lineari-subspathulatis. Capitula subnume-

rosa, tenuiter pedunculata pedunculis glabris plerumque 5-15 cm.

longis, radiata, pansa ad anthesin 3-5 cm. lata et 7-10 mm. alta.

Involucri bracteae exteriores 8-12, saepe irregulariter positae,

lineari-oblongae, glabrae, margine scariosae, apice obtusae, 1-3 mm.

longae; interiores ovatae, saepissime glaberrimae, 6-8 mm. longae.

REVISION OF THE GENUS COREOPSIS 429

Flores ligulati plerumque 8, guttur versus brunneo-purpurei alibi

flavi, ligula cuneate obovati, apice 3-lobati lobo mediano rursus

2-3-lobulato, 1.2-1.8 cm. longi. Paleae tenuiter oblongo-lineares,

marginibus scariosae alibi aurantiaco-rubrae, ±5 mm. longae. Flores

tubulosi corolla purpureo-rubri, stylorum ramis apice abrupte sub-

truncatis. Achaenia valde obcompressa, nigra, oblonga vel oblongo-

oblanceolata, faciebus glabra vel minutissime sparsimque papillatal

marginibus angustissime alata, 2.4-2.8 mm. longa, apice calva ve,

vix bidentata.

Type specimen: Collected by David Douglas on Mewries (or

Menzies, cf. DC. Prodr. 5: 568. 1836) Island in the Columbia River

in 1825.

Distribution: Saskatchewan and British Columbia southwardly

to Oregon, Arizona, and South Dakota.

Specimens examined: L. R. Abrams 9465, Hood River, Hood

River County, Oregon, July 25-28, 1922 (Pom.); Anon., cult, ex

sem. ex herb. DeCand., anno 1839 (Gray) ; E. Bourgeau, rare, near

lake on the prairie, Saskatchewan, September 17, 1857 (Kew);

Townsend S. Brandegee, the Dalles, Oregon, 1882 (Calif.); Douglas,

North America (Gray); idem, Fort Vancouver, British Columbia

(Gray) ; Drake & Dickson, Bradfords Island, Columbia River, June,

1889 (Field) ; A. D. E. Elmer 601, sand-gravel shores of Palmer Lake,

northwest of Loomiston, Washington, August, 1897 (Berl.; Pom.);

Griffiths, Highmore, South Dakota, July, 1897 (Berl.);Elihu Hall,

Oregon (Field) ; idem, cult, from Dakota seed, 1877 (Gray) ; W. G. W.

Harford & George W. Dunn, Cascades, northwestern United States,

May 28, 1869 (Berl.); A. Arthur Heller, Coeur d'Alene River, near

Harrison, Idaho, July 12, 1892 (Field); idem, about Lake Pend

d'Oreille, near Lakeview, Idaho, August 1-10, 1892 (Pom.); Thomas

J. Howell, Oregon, 1881 (Field) ; idem, Klickitat, Washington, June,

1881 (Field); idem, banks of Columbia River, July, 1881 (Berl.);

idem 224, Multnomah County, Oregon, July, 1877 (Gray); Marcus

E. Jones, Sand Point, Idaho, September 18, 1908 (Pom.); Frank 0.

Kreager 315, marshy shore of Lake Calispell, Stevens County,

Washington, July 28, 1902 (Gray); John B. Leiberg 1542, alt. 640

meters, region of Coeur d'Alene Mountains, Idaho, August 17,

1895 (Field; Gray; Pom.); Lemmon 4159, meadow near San Fran-

cisco Mountains, Arizona, September, 1884 (Gray); Dr. Lyall,

Cascade Mountains to Fort Colville, lat. about 49° N., 1860 (Gray) ;

idem, from Fort Colville to Rocky Mountains (Oregon Boundary),

1861 (Berl.); Daniel T. MacDougal 617, Lakeview, Idaho, August,

430 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

1892 (Field) ; Ernest MacKay 25, very abundant in wet sand, head

of Grand Coulee, Douglas County, Washington, July 12, 1902

(Gray; Pom.); J- M. Macoun, between Kettle and Columbia Rivers,

Waneta, British Columbia, July 28, 1902 (Berl.; Field; Gray; Pom.);

J. C. Nelson 1980, sandy shore of Columbia River, Hayden Island,

Oregon, September 8, 1917 (Gray) ; Frank W. Peirson 3900, near

Biggs, Oregon, July 16, 1923 (Pom.) ;C.G. Pringle, banks of Columbia

River, Washington, September 10, 1881 (Field); Margaret P. (Mrs.

Robert Shaw) Russell, Hood River, Oregon, August 15 (Gray);

J. H. Sandberg, D. T. MacDougal, & A. A. Heller 617, on shores

near Farmington Landing, Lake Coeur d'Alene, Idaho, July 11,

1892 (Berl.; Field; Gray; Pom.); E. P. Sheldon 11081, Bonneville,

Oregon, August 10, 1902 (Gray; Pom.); Sherff 250 pro parte, cult.,

St. Louis, Missouri, July 6, 1910 (Gray); Rev. Mr. Spalding, Clear

Water, Oregon (Gray) ; W. N. Suksdorf, bottom lands of the Columbia

River, Washington, August 26, 1882 (Berl.); G. R. Vasey 551,

Washington, 1889 (Gray); Sereno Watson 217, Bed of Columbia

River, Old Fort Colville, Washington, October 3, 1880 (Gray);

idem (similiter) 217, mouth of the Chelan, Columbia River, Okan-

ogen County, Washington, October 12, 1880 (Gray).

100. Coreopsis Leavenworthii Torr. & Gray, Fl. N. Amer.

2: 346. 1843. Coreopsis angustata Greene, Pittonia 4: 239. 1901.

Folia simplicia var. d Lewtonii.

Folia principalia 1-2-pinnata.

Foliorum caulinorum foliola terminalia nunc obovata nunc lanceo-

lato-oblonga, 0.8-1.5 cm. lata var. /3 Garberi.

Foliorum caulinorum foliola terminalia (vel eorum segmenta

ultima) circ. 1-3 mm. lata.

Folia basalia pinnatim circ. 9-partita var. 7 Curtissii.

Folia basalia integra C. Leavenworthii sensu stricto.

Herba annua, erecta, glabra, 5-15 dm. alta, caulibus teretibus

subsimplicibus vel suberecte ramosissimis, omnino vel fere usque ad

summam foliosis. Folia opposita, ima integra elongate lineari-

spathulata, quam internodia longiora, petiolis ±1 dm. longis,

laminis ±5 cm. longis et saepius 3-5 mm. latis; principalia internodiis

longiora, saepius 2-5-partita segmentis anguste linearibus plerumque

1-3 mm. latis et petiolo elongate plus minusve similibus; summa

integra, internodiis breviora. Capitula pauca vel numerosa, pa-

niculato-corymbosa, tenuiter pedicellata, radiata, pansa ad anthesin

2-3 cm. lata et circ. 5-6 mm. alta. Involucri bracteae exteriores

REVISION OF THE GENUS COREOPSIS 431

circ. 6-10, saepe irregulariter positae, lineari-lanceolatae vel ovato-

lanceolatae, saepe coloratae, apice obtusae, margine plus minusve

diaphanae, tergo plerumque scabridae, tantum 1-2 mm. longae;

interiores ovatae, supra pubescentes, 5-7 mm. longae. Flores

ligulati circ. 8, flavi, ligula obovati, apice 3-lobati lobis triangulato-

rotundatis mediano majore saepe emarginato, 0.8-1.5 cm. longi.

Paleae lineares vel subfiliformes, acutae, demum circ. 2-4 mm.

longae. Flores tubulosi numerosi, minuti (circ. 2.2 mm. longi),

stylorum ramis apice lato-conicis. Achaenia obcompressa, glabra,

corpore nigro 2-3 mm. longo et circ. 0.5-0.7 mm. lato lineari-oblonga,

marginibus membranaceo-alata ala subbrunnea corporis latitudinem

aequante vel superante, apice biaristata aristis erectis tenuibus

antrorsum hispidulis circ. 0.3-0.6 mm. longis.

Type specimen: Collected by Melines C. Leavenworth, in Florida.

The first cited specimen came from Tampa Bay (Gray).

Distribution: Florida.

Specimens examined: John H. Barnhart 2715, moist ground near

Ehren, April 26-May 16, 1899 (Field) ;F. S. Blanton 6972, wet ditch,

roadside, south of Lakeland, March 28, 1930 (Field); Mrs. E. G.

Britton 132, prairie west of Cutter, March 23, 1904 (Field) ; Buckley,

Florida (Gray); ex herb. William Canby, shores of Indian River,

February, 1889 (Gray) ; A. H. Curtiss 5375, shore of Lake Worth,

May 8, 1895 (Gray; Pom.); Mrs. Charles C. Deam 1709, border of

Lake Worth, near West Palm Beach, February 22, 1904 (Gray);

A. Fredholm 5^73, prairie, Orange County, August 8, 1902 (Gray) ;

idem 5778, low prairie, Okeechobee region, Brevard County, April

13, 1903 (Gray); idem 6094, swamp, Osceola County, October 12,

1903 (Gray); A. P. Garber, Tampa Bay, 1876 (Gray); idem, Keys of

Florida, 1876 (Gray); idem, Mellonville, March, 1876 (Field); idem,

Lake Monroe, March, 1876 (Field); idem, Tampa, May, 1876 (Field);

idem, Rosewood, June, 1876 (Field); W. Garvens, vicinity of Palm

Beach, February 15-March 15, 1908 (Field); Albert S. Hitchcock,

New River, December 26, 1895-January 11, 1896 (Field); idem,

Palm Beach, eodem tempore (Field); idem, Levy County, June-

July, 1898 (Field); William Kellogg, Owanita, about March 18,

1907 (Gray) ; Dr. Leavenworth, Tampa Bay (type, Gray) ; George V.

Nash 832, hummock land, vicinity of Eustis, May 16-31, 1894 (Berl. ;

Field; Gray); idem 1257, eodem loco, July 1-15, 1894 (Berl.; Gray);

idem 1876, Orange Bend, May 28-June 15, 1895 (Pom.); Edward

Palmer 286, Indian River, 1874 (Gray);Fcmme R. Randolph 136,

moist sand among sand dunes near Lake Worth, Kelsey City,

432 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

January 2, 1921 (Gray; foliis omnibus simplicibus) ; F. Rugel 102,

Florida (Field); idem 142, Florida, 1845 (Mo.); John K. Small &

J. J. Carter, Bull Key, opposite Lemon City, November 6, 1903

(Field); iidem 1219, Fort Lauderdale, November, 1903 (Field);

John Donnell Smith, grassy pine barrens, Sumter County, March

21, 1883 (Field); Miss Jeanette P. Standley 195, in pine land, vicinity

of Fort Myers, May 20, 1916 (Field; Gray); F. C. Straub 97, Port

Orange, April 9, 1895 (Gray); idem 159, eodem loco, May 18, 1895

(Gray); S. M. Tracy 6920, Palma Sola, May 16, 1900 (Mo.); idem

6921, eodem loco et tempore (Field, 2 sheets; N.Y.; U.S.; cotypes of

Coreopsis angustata Greene); idem 7704, Braidentown, June 15,

1901 (Berl.; Field; Gray); H. J. Webber, Sanibel Island, January 29,

1896 (Field); idem 414, margin of Lake Worth, West Palm Beach,

June 7, 1896 (Field).

Coreopsis Leavenworthii var. /3 Garberi A. Gray, Syn. Fl.

N. Amer. 1, pt. 2: 291. 1884.

Folia caulina omnia pinnatim 5-7-partita foliolis brevioribus

latioribusque, terminali subobtuso nunc obovato nunc lanceolato-

oblongo, 0.8-1.5 cm. lato.

Type specimen: Collected by Abraham Pascal Garber, Tampa,

Florida, May, 1876 (Gray).

Distribution: Known only from type locality of Tampa, Florida.

Specimens examined: A. M. Ferguson, pine barrens, Tampa,

June-July, 1898 (Mo.); A. P. Garber, Tampa, May, 1876 (type,

Gray).

Coreopsis Leavenworthii var. 7 Curtissii Sherff, Bot. Gaz.

94: 592. 1933.

Folia 1-2-pinnata; basalia pinnatim circ. 9-partita, foliolis

oblongo-linearibus ad basim sensim angustatis plerumque 2-4 cm.

longis et 2-5 mm. latis, lateralibus (praecipue inferioribus) utrinque

saepe rursus 1-lobatis lobo parvo linearique; caulina quam internodia

nunc multo longiora nunc multo breviora, plerumque 2-pinnata

foliolis lateralibus saepius 3 vel 4 jugis, segmentis ultimis omnibus

0.5-2.5 cm. longis et circ. 1-3 mm. latis.

Type specimen: Collected by Allen Hiram Curtiss, No. 6734, in

dry, pastured ground along the Suwannee River, near Branford,

Florida, October 23, 1900 (Gray, 2 sheets).

Distribution: Known only from type locality in Florida.

Specimens examined: Curtiss 6734 (Gray, 2 type sheets).

REVISION OF THE GENUS COREOPSIS 433

Coreopsis Leavenworthii var. 8 Lewtonii (Small) Sherff,

Bot. Gaz. 94: 592. 1933. C. Lewtonii Small, Bull. Torr. Bot. Club

25: 146. 1898.

E specie foliis simplicibus plerumque quam internodiis breviori-

bus radice saepe perenni differt.

Type specimen: Collected by Frederick Lewis Lewton, Forest

City, Florida, 1894 (N.Y.).

Distribution: Florida.

Specimens examined : Liberty H. & Ethel Zoe Bailey 13047, sands

at Sarasota, September 28, 1929 (Field); A. H'Curtiss 1480, swamps

and shores, Halifax River, June (Berl.; Field, 2 sheets; Gray); A. A.

Eaton 469, Little River, Bade County, December 5, 1903 (Field,

2 sheets); Albert S. Hitchcock 139, around flatwood ponds, Myers,

July-August, 1900 (Field) ; Frederick L. Lewton, Forest City, 1894

(type, N.Y.); idem, Cedar Hammock, Sumter County, August 3,

1894 (N.Y.); George V. Nash 2255, Sanford, July 24, 1895 (Berl.;

Field; Gray); F. Rugel 133, Florida, 1843 (Gray; Mo., 2 sheets);

J. K. Small & J. J. Carter 3025, the Everglades, intersecting Long

Key, January 18-26, 1909 (N.Y.); John Donnell Smith, pine-barren

ponds, Pellicier's Creek, St. John's County, March 4, 1882 (Field,

3 sheets; Gray); S. M. Tracy 7256, Long Key, May 8, 1901 (Berl.;

Field; Gray); idem 7355, Perico Island, December 6, 1901

(Gray; Mo.).

101. Coreopsis saxicola Alex. Torreya 32: 161. 1932.

Herba perennis, erecta, glabra, suberecte ramosissima, saltern

superne foliosissima, 7-10 dm. alta, caule subtereti ramis tetragonis.

Folia opposita, principalia petiolata petiolis eciliatis inferne dilatatis

basi connatis saepius 1-4.5 cm. longis, pinnata vel bipinnata seg-

mentis oblongo-linearibus sparsim minuteque ciliatis 2-9 (rarius

-13) mm. latis (raro multa simplicia spathulataque). Capitula

corymbose disposita, tenuiter pedunculata pedunculis ±1 dm.

longis, radiata, pansa ad anthesin 2.5-4 cm. lata et circ. 1-1.2 cm.

alta. Involucri bracteae exteriores plerumque 7 vel 8, lanceolatae,

apice acutae vel subacutae ac saepe hispido-ciliatae alibi glabrae,

marginibus albidae, 7-9 mm. longae; interiores oblongo-ovatae

paulo longiores. Flores ligulati circ. 8, subaurantiaci, ligula oblongi

vel cuneato-obovati, apice tenuiter lobati lobis plus minusve lobula-

tis, ±1.5 cm. longi. Paleae lineares, superne attenuatae atque

elongatae, demum circ. 6-8 mm. longae. Florum tubulosorum styli

ramorum terminis breviter conici. Achaenia obcompressa, valde

434 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

incurva, circumambitu orbiculata vel obovato-orbiculata, atra,

faciebus glabra vel papillata, circ. 2 mm. longa et (alls brunneis

irregulariter pectinato-laceratis inclusis) circ. 2-2.5 mm. lata, apice

demum calva.

Type specimen: Collected by Francis W. Pennell, No. 4029, on

thin soil over granite, Stone Mountain, DeKalb County, Georgia,

August 2, 1912 (N.Y.).

Distribution: DeKalb County, northern Georgia, westward into

Tallapoosa County, Alabama.

Specimens examined: F. S. Earle 2147, Double Bridges, Talla-

poosa County, Alabama, August 9, 1899 (N.Y.); H. Eggert, Stone

Mountain and vicinity, DeKalb County, Georgia, July 23, 1897

(Mo., 2 sheets; N.Y.);F. W. Pennell 4029 (type, N.Y.); JoJm Kunkel

Small, alt. 300-450 meters, on and about Stone Mountain, Georgia,

August 1-6, 1895 (Field; N.Y.); Karl M. Wiegand & W. E. Manning

S421, sandy soil, northwest base of Stone Mountain, Georgia,

August 17, 1927 (Pom.; forma foliis plerumque simplicia); iidem

8422, turfy rocky slope, base of Stone Mountain, Georgia, August 17,

1927 (Pom.).

An anomalous species, combining the foliar habit, involucre, and

paleae of Eucoreopsis (especially C. grandiflora') with the lacerate-

winged achenes of species like C. Linifolia. The tips of the style

branches of the disc florets are somewhat intermediate, being less

elongate-cuspidate than in the former but more definitely conical

than in the latter.

102. Coreopsis paludosa M. E. Jones, Contr. West. Bot. 12:

46. 1908.

Herba annua, erecta, glabra, subsucosa, simplex vel paniculato-

ramosa, ±6 dm. alta, internodiis elongatis usque ad 1.7 dm. longis.

Folia alterna, nunc simplicia (pro ramis vel plantis parvis), anguste

spathulata, 3.5-7 cm. longa et 2-5 mm. lata, lamina sensim in

petiolum subaequalem decurrente, integra, apice subobtusa; nunc

1-2-pinnata (pro plantis majoribus), majora (radicem versus posita)

1.5-2.25 dm. longa petiolata petiolo ±8 cm. longo foliolis lineari-

spathulatis apice subobtusis (lateralibus circ. 3 jugis, 2-4 cm. longis

et 1.5-3.5 mm. latis terminali circ. 6-8 cm. longo et 7-8 mm. lato)

saepius (ad marginem ad folii terminum spectantem) 1-lobatis lobo

±8 mm. longo; altera minora, 1-pinnata, foliolis lateralibus 2 jugis.

Capitula pauca (±15), tenuiter pedicellata pedicellis usque ad 8 cm.

longis, radiata, pansa ad anthesin ±3.5 cm. lata et circ. 7-8 mm.

REVISION OF THE GENUS COREOPSIS 435

alta. Involucri glabri bracteae exteriores 4-6, oblongae, apice

obtusae, plurinervatae, margine plus minusve hyalinae, adpressae

vel rarius patentes, circ. 2 mm. longae; interiores ovatae, superne

subelongatae, demum 6-8 mm. longae. Flores ligulati circ. 8, ligula

rhomboideo-obovati, ad ligulae basim purpureo-maculati aliter flavi,

supra medium 3-lobati lobis lateralibus parvis terminal! multo

majore, 1.5-2 cm. longi. Paleae anguste lineares, glaberrimae, 4-5

mm. longae. Florum tubulosorum stylorum rami termino truncati.

Achaenia obcompressa, nigra, anguste obovata, basi flavo-callosa,

faciebus glaberrima, marginibus pectinatim cartilagineo-dentata

dentibus brunneis quam 0.1 achaenii latitudinis non longioribus,

apice calva, 2.4-3.2 mm. longa et circ. 1.2-1.4 mm. lata.

Type specimen: Collected by Marcus E. Jones at altitude of

2,100 meters, growing in water in creeks, etc., Marsh Lake, Sierra

Madre, State of Chihuahua, Mexico, September, 1903 (Pom.).

Distribution: Known only from type locality in State of Chi-

huahua, Mexico.

Specimens examined: Marcus E. Jones, alt. 2,100 meters, Meadow

Valley, Sierra Madre, State of Chihuahua, September 17, 1903

(Pom.; an associate type).

103. Coreopsis Linifolia Nutt. Journ. Acad. Phila. 7: 75.

1834. Coreopsis callosa Bertol. Misc. Bot. 7: 42. 1848.

Herba perennis, glabra, pallida, erecta, 5-7 dm. alta, caulibus

gracilibus, supra ramosis, plus minusve tetragonis. Folia opposita

vel inferiora alterna, integra, inferiora laminis spathulata vel oblanceo-

lata quam internodia breviora (vel basalia interdum duplo longiora)

4-9 mm. lata apice rotundata basi petiolata petiolis saepius 1-4 cm.

longis; alia sessilia, linearia, apice saepe truncata vel obtusissima,

multo minora. Capitula pauca, corymboso-paniculata, tenuiter

pedunculata pedunculis saepius 3-7 cm. longis, radiata, pansa ad

anthesin 2-3 cm. lata et circ. 6-7 mm. alta. Involucri bracteae

exteriores circ. 6-8, glabratae, saepe irregulariter positae, ovatae,

longitudinaliter pluristriatae, lateribus scariosae, apice subacutae

vel rotundatae, 1-3 mm. longae; interiores oblongo-ovatae, saepius

glabrae, 5-8 mm. longae. Flores ligulati circ. 7 vel 8, flavi, ligula

obovati, apice 3-lobati lobo mediano emarginato, 1-1.5 cm. longi.

Paleae lineares, apice acutae, circ. 5-6 mm. longae. Flores tubulosi

corolla atro-purpurei, stylorum ramis apice obtuse conicis. Achaenia

plana, oblonga vel ovalia, subnigra, faciebus glabra, marginibus

ornata ala irregulariter lacerata circ. 0.2-0.3 corporis latitudinis

436 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

lata, corpore circ. 3 mm. longa, apice tenuiter biaristata aristis

sparsim antrorsumque hispidis ±1 mm. longis.

Type specimen: Collected in Alabama (Brit.?).

Distribution: North Carolina to Florida, thence westwardly to

Alabama and Mississippi.

Specimens examined (formerly confused by me in herbaria with

C. longifolia Small; for that reason there are cited here only the

few specimens which have been recently reexamined) : A. W. Chap-

man, wet barrens, Florida (Field); ex herb, eiusdem 174b, swamps

in the pine barrens, Apalachicola, Florida, September 22, 1879

(Gray; Mo.) ; A. B. Langlois, Spring Hill, southern Alabama, October

1, 1880 (Field) ; J. Skehan, Ocean Springs, Mississippi, September 16,

1895 (Mo.) ; John Donnell Smith, open tidal marshes on Biloxi Bayou,

Harrison County, Mississippi, September 16, 1885 (Field; Gray);

S. M. Tracy 4778, Koshtaw, Mississippi, October 25, 1898 (Field;

Mo.); idem 8565, Gateswood, Alabama, October 31, 1903 (Field;

Gray; Mo.).

The type of Coreopsis callosa Bertol. has not been seen by me.

Bertoloni's long description, however, fits C. Linifolia. His concluding

observation ("Haec species appropinquaret Coreopsidem angusti-

foliam Ait., si haberet omnia folia alterna, et lanceolato-linearia")

shows beyond doubt that he had our C. Linifolia before him. More-

over, Bertoloni's type had been collected by Dr. Gates in Alabama.

A Gates specimen still extant (N.Y. ; not cited above) was undoubt-

edly a duplicate. I found this latter to be typical C. Linifolia.

104. Coreopsis longifolia Small, Bull. Torr. Bot. Club 22: 47.

1895. C. angustifolia Dryand. in Ait. Hort. Kew. ed. 1. 3: 253.

1789. (Non L. Sp. PI. 908. 1753.)

Herba perennis, glabra, pallida, 7-12 dm. alta, erecta, caulibus

simplicibus vel summam versus parce ramosis, teretibus. Folia

alterna, integra, basalia (petiolis elongatis adjectis saepius 1-2 dm.

longa) quam internodia multo longiora laminis linearia vel lineari-

oblonga vel lineari-lanceolata 2-11 mm. lata apice plus minusve

acuta marginibus saepe subscariosa; superiora sessilia, ad bracteas

lineares plus minusve reducta. Capitula pauca paniculato-corym-

bosa, tenuiter pedunculata pedunculis usque ad 2 dm. longis, radiata,

pansa ad anthesin 3-5.5 cm. lata et ±1 cm. alta. Involucri bracteae

glabrae exteriores circ. 7-9, ovatae vel oblongo-lanceolatae, lateribus

plus minusve diaphanae, apice saepe rotundo-dilatatae, 2-5 mm.

longae; interiores oblongo-ovatae 6-12 mm. longae. Flores ligulati

circ. 8, flavi, ligula cuneate obovati, apice 3-lobati lobo mediano

REVISION OF THE GENUS COREOPSIS 437

rotundato vel emarginato, 1.5-2.5 cm. longi. Paleae oblonge lineares,

acutae, demum usque ad 1 cm. longae. Flores tubulosi corolla atro-

purpurei, stylorum ramis apice obtusissime conicis. Achaenia plana,

corpora ipso circ. 4 mm. longo atra et circumambitu (alis exclusis)

oblonga vel late linearia, faciebus glabra margine pectinato-alata

dentibus brunneis quam corporis diametro 0.3-0.6 brevioribus,

apice breviter biaristata aristis antrorsum setulosis saepe caducis.

Type specimen: Collected by Allen Hiram Curtiss, No. 4489, in

dry, grassy pine woods about Jacksonville, Florida, October 16, 1893

(N.Y.).

Distribution: North Carolina, Georgia, and Florida.

Specimens examined (perhaps several additional specimens

determined by me some years ago for various herbaria as C. Linifolia

Nutt. will be found to belong here): Curtis, North Carolina (N.Y.);

A. H. Curtiss, Florida, November (Field) ; idem, near Jacksonville,

Florida, September (Gray); idem 1470, margin of swamp near Jack-

sonville, Florida, October (Pom.); idem 14.77 pro parte, moist pine

barrens near Jacksonville, November (Field, 2 sheets; N.Y.);

idem 4489 (N.Y., 2 type sheets) ; idem 5266, moist pine barrens near

Jacksonville, October 15, 1894 (Field; Gray; N.Y., 2 sheets; U.S.);

idem 5348 pro parte, thickets bordering swamps near Jacksonville,

October 22, 1894 (N.Y.); idem 5568, moist, grassy pine barrens near

Jacksonville, October 7, 1895 (Field; Gray; Mo., 2 sheets; Pom.);

idem 6278, sides of ditch near Jacksonville, October 31, 1898 (Field;

Gray); Mary E.Francis 155, low grounds, Alva, Florida, November

25, 1917 (U.S.); A. P. Garber, Levy County, Florida, November,

1877 (Field; Gray; U.S.); herb. Otto Katzenstein, southeastern

United States, September 23, 1897 (Gray); L. H. Lighthipe 557 1

South Jacksonville, Florida, October 30, 1896 (N.Y.); F. Rugel 480,

Florida, 1845 (Field; U.S.); J. H. Simpson, Manatee, Florida, 1889

(U.S.) ; J. K. Small & J. J. Carter 1028, pine lands, Fort Lauderdale,

Florida, November 19-25, 1903 (Field); J. K. Small &G. V. Nask,

the Everglades, west of Miami, Florida, November 1-9, 1901 (N.Y.);

Mm Jeanette P. Standley 423, low pine land, vicinity of Fort Myers,

Florida, October 25, 1916 (Pom.); S. M. Tracy 7142, Braidentown,

Florida, October 16, 1900 (Field; Gray; Mo.; N.Y.; U.S.).

Passes into C. gladiata.

105. Coreopsis falcata Boynt. Biltmore Bot. Studies 1: 141.

1902.

Herba perennis, pallida, glabra, 8-12 dm. alta, caulibus teretibus,

summam versus corymbose ramosis. Folia alterna, inferiora petio-

438 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

lata petiolis 1-1.5 dm. longis lamina (aequaliter 1-1.5 dm. longa;

0.6-3 cm. lata) elliptico-oblonga vel linearia vel lineari-oblanceolata

terminaliter nunc obtusa nunc longe attenuata marginibus Integra

vel saepe 1-2-lobata lobis angustis adscendentibusque; superiora

sessilia, multo minora, angustiora, lateraliter saepe 1-plus-lobata.

Capitula pauca vel vix subnumerosa, tenuiter pedunculata pedunculis

±1 dm. longa, radiata, pansa ad anthesin 3-5 cm. lata et 6-9 mm.

alta. Involucri glabri bracteae exteriores circ. 8-10, lanceolatae,

4-7 mm. longae; interiores ovatae vel oblongo-lanceolatae, dimidio

longiores. Flores ligulati circ. 8, aurantiaco-flavi, ligula cuneato-

obovati, apice 3-lobati lobo mediano majore rotundato vel emar-

ginato, 1.3-2.3 cm. longi. Paleae lineares, acutae. Achaenia plana,

oblonga, corpore circ. 4.5 mm. longa, marginibus alata alis latis

pectinatis dentibus corporis diametrum fere aequantibus, apice

biaristata aristis usque ad 1 mm. longis.

Type specimen: Collected for the Biltmore Herbarium, in shallow

water near Pembroke, North Carolina, June, 1901. The type proper

was preserved in the Biltmore Herbarium, where it was doubtless

destroyed later on by flood waters. A sheet of fine duplicate material

is still extant, however (N.Y.).

Distribution: North Carolina to Georgia.

Specimens examined: Anon., Georgia (Gray); anon., Macon,

Georgia (N.Y.) (these first two specimens are doubtless of the same

collection; the first had been studied by Asa Gray and was regarded

by him as C. gladiata in his Synoptical Flora of North America; the

second had come from John Torrey's private herbarium and was

under the name C. gladiata; this latter specimen was probably the

basis for Torrey and Gray's words "sometimes 1-2-ternately-parted"

in describing the lower leaves of C. gladiata [Fl. N. Amer. 2: 347.

1843]); ex Biltmore Herb., shallow water near Pembroke, North

Carolina, etc. (cotype, N.Y.); ex eodem 14?58b, shallow water or wet

places in pine lands, Selma, North Carolina, May 31, 1902 (N.Y.);

Cuthbert, in pool, Georgia, October 10, 1898 (N.Y.); Karl M.

Wiegand & W. E. Manning 3410, dry pine woods, 7 miles north of

Washington, North Carolina, June 21, 1927 (Pom.).

106. Coreopsis gladiata Walt. Fl. Carol. 215. 1788. C. dicho-

toma Michx. Fl. Bor. Amer. 2: 137. 1803.

Herba perennis, glabra, pallida, erecta, 6-8 dm. alta, caulibus

teretibus vel vix angulatis, supra ramosa et subnudata . Folia plerum-

que alterna, integra, inferiora petiolata petiolis elongatis quam inter-

REVISION OF THE GENUS COREOPSIS 439

nodia plerumque multo longiora laminis elliptico-oblonga vel

oblanceolata marginibus scariosa plerumque 1-2.5 cm. lata apice

rotundata atque interdum aegre mucronata; superiora sessilia, multo

minora, linearia vel spathulato-linearia. Capitula pauca, sub-

corymbosa, tenuiter pedunculata pedunculis glabris ±1 dm. longis,

radiata, pansa ad anthesin 3-4 (-6) cm. lata et 8-11 mm. alta.

Involucri glabri bracteae exteriores 4-10, saepe biseriatim dispositae,

ovatae vel lanceolatae, plerumque obtusae, lateraliter scariosae,

2-4 (-6) mm. longae; interiores oblongo-ovatae 8-12 mm. longae.

Flores ligulati circ. 8, flavi, ligula cuneato-obovati, apice 3-lobati

lobo mediano rotundato vel saepius emarginato, 1-2.5 cm. longi.

Paleae lineares, apice acutae, demum ±8 mm. longae. Flores

tubulosi corolla atro-purpurei, stylorum ramis apice late breviterque

conicis. Achaenia plana, elliptico-oblonga, faciebus nigris glabra,

marginibus anguste pectinato-alata, corpore circ. 3.5 mm. longa,

apice biaristata aristis tenuibus antrorsum hispidis.

Type specimen: No specimen cited. The plant in mind was

native to Carolina. My notes, contain no mention of an authentic

(author's) specimen among the Walter specimens in London (Brit.).

Distribution: North Carolina to southern Florida and west-

wardly to southern Mississippi.

Specimens examined: C. Billington 131, low places near salt water,

Lynn Haven, Bay County, Florida, October 15, 1921 (U.S.);

Biltmore Herb, (ex herb. A. W. Chapman) 20661), low pine barrens,

Apalachicola, Florida, September 22, 1879 (Mo.; U.S.); J. Brown-

field, Summerville, South Carolina, October, 1892 (Mo.); William

M. Canby, vicinity of Wilmington, North Carolina, October, 1867

(Field; Mo.); ex herb. A. W. Chapman, southern Florida (Field;

U.S.); A. H. Curtiss, Chattahooche, Florida, 1875 (U.S.); idem

1477 pro parte, moist pine barrens near Jacksonville, Florida,

November (Gray; Mo.); Roland M. Harper 1666 p.p., wet barrens

north of Moultrie, Georgia, September 24, 1902 (Berl.; Mo.; U.S.);

Albert S. Hitchcock 910, New River, Florida, December 26, 1895-

January 11, 1896 (Field) ; Gerald MacCarthy, eastern North Carolina,

1883 (Field) ; Michaux, North America (Berl., sub nom. Coreopside

dichotoma Michx.); Charles Mohr, margins of pine-barren swamps,

Poplarville, Mississippi, October 18, 1894 (U.S.); John H. Redfield

5630, near foot of Blue Ridge, Caesar's Head, South Carolina,

September 3, 1876 (Mo.); F. Rugel, Florida, 1842-1849 (Mo.;

foliorum apicibus C. longifoliae adpropinquans) ; John K. Small &

J. J. Carter 694, in the Everglades, between Cocoanut Grove and

440 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Cutler, Florida, October 31-November 4, 1903 (N.Y.); Udem 954,

in pine lands, Fort Lauderdale, Florida, November 19-25, 1903

(Field; N.Y.); iidem 2731, pine lands between Perrine and Larkin,

Florida, November 16, 1906 (N.Y.); iidem & G. K. Small 3401, in

the Everglades, Cutler to Homestead, Florida, February 15, 1911

(N.Y.); J. T. Stewart, in swamp, near Charleston, South Carolina,

1865 (Field); ex herb. Katherine A. Taylor, Summerville, South

Carolina (Pom.).

Passes into C. longifolia and apparently also into C. Linifolia.

107. Coreopsis Helianthoides Beadle, Bot. Gaz. 25: 448. 1898

(non C. Helianthoides Forst. Prodr. 91, num. 543. 1786; nomen

subnudum).

Herba perennis, erecta, pallida, 5-12 dm. alta, caulibus glabris,

plus minusve teretibus, summam versus erecto-ramosis. Folia

alterna, moderate adpresso-hispida setis pluriloculatis, marginibus

scariosae, apice obtusae vel acutae, inferiora longe petiolata petiolis

glabratis conduplicatis saepius 0.5-1.5 dm. longis, lamina (saepe

rhomboide vel oblonge) ovata vel ovato-lanceolata 6-13 cm. longa

et 2-6 cm. lata; superiora abrupte minora, sessilia, elliptico-oblonga

vel linearia. Capitula pauca, corymbosa, tenuiter pedunculata

pedunculis ±1 dm. longis, radiata, pansa ad anthesin 3-4.5 cm. lata

et 7-9 mm. alta. Involucri glabri bracteae exteriores 6-11, saepe

subimbricatae, ovatae vel deltoideo-lanceolatae, tergo longitudinaliter

pluristriatae, basi plus minusve subauriculatae, lateribus subdia-

phanae, apice obtusae, plerumque 1-5 mm. longae; interiores ovatae

vel interdum oblongo-lanceolatae, 8-12 mm. longae. Flores ligulati

plerumque 8, flavi, ligula cuneato-obovati, apice grosse 3-lobati lobo

mediano saepe emarginato, ±2 cm. longi. Paleae tenuiter oblongo-

lineares, ±4-5 mm. longae. Flores tubulosi corolla atropurpurea,

stylorum ramis apice obtuse conicis. Achaenia oblonge oblanceolata

vel obovata, valde obcompressa, subnigra, faciebus glaberrima,

marginibus subregulariter pectinato-alata dentibus brevibus longi-

tudine circ. 0.1-0.2 corporis latitudinis aequantibus, circ. 3 mm.

longa, apice brevissime biaristata.

Type specimen: Collected by Alvin Wentworth Chapman, Aspa-

laga, Florida, October, 1897. Herbarium not cited but presumably

was the Biltmore Herbarium. The Chapman specimen No. 2066c

in Herb. N. Y. had been distributed from the Biltmore Herbarium.

Distribution: North Carolina southward to Florida and Alabama.

REVISION OF THE GENUS COREOPSIS 441

Specimens examined: Ex Biltmore Herb. 2139, Thomasville,

Georgia, September 21, 1901 (N.Y.); William M. Canby, near

Wilmington, North Carolina, October, 1867 (Field; Mo.; N.Y.);

A. W. Chapman, Aspalaga, Florida, 1897 (U.S.); ex. herb, eiusdem

2066b, low pine barrens, Apalachicola, Florida, September 22, 1879

(Gray); (exBiltmore Herb.) ex eodem herb. 2066c, eodem loco, October,

1897 (N.Y.; undoubtedly cotype material); A. H. Curtiss .1477

pro parte, pine barrens, near Jacksonville, Florida, November

(Berl.; Field; Mo.; U.S.); idem 4-487, near Jacksonville, Florida,

October 21, 1893 (N.Y.; U.S.); idem 5348 pro parte, thickets

bordering swamps, eodem loco, October 22, 1894 (Field; N.Y., 2

sheets; U.S., 2 sheets); idem 10223, Chattahooche, Florida, October,

1875 (Mo.); A. Cuthbert, peat bog, Augusta, Georgia, September 25,

1898 (N.Y.); Roland M. Harper 653, alt. 90 meters, sandy bog

just southeast of Americus, Georgia, September 15, 1900 (N.Y.);

idem 1666 p.p., wet barrens north of Moultrie, Georgia, September

24, 1902 (Gray; N.Y.) ; Charles Mohr, below lake, pine barren swamp,

Yellow Pine, Alabama, August 8, 1895 (U.S.); J. B. Norton, swamp

near mouth of Kilgore Branch, east of Hartsville, South Carolina,

November 5, 1921 (U.S.); H. Ravenel, damp pine land, Santee

Canal, South Carolina, September (Gray) ; F. Rugel 570 pro parte,

Florida, 1845 (Field) ; John Donnell Smith, swampy woods, Columbia,

South Carolina, September, 1883 (Field, 2 sheets); idem 2028,

swampy woods 4 miles northeast of Columbia, September 25, 1883

(Field; Gray; U.S., 2 sheets); Taylor, Thomasville, Georgia, Septem-

ber 30, 1903 (Gray); Otto Vesterland, Marion County, Florida,

November, 1889 (U.S.).

108. Coreopsis integrifolia Poir. Encycl, Suppl. 2: 353. 1811.

Herba perennis, erecta, gracilis, 6-9 dm. alta, caule glabro,

tereti vel subangulato, subsimplici, fere omnino folioso. Folia

opposita, petiolata petiolis hispido-ciliatis usque ad 3 cm. longis (vel

summa sessilia); laminis diverse ovalia vel ovata vel suboblonga,

crassiuscula, utrinque acuta vel subobtusa, marginibus venisque

saltern primo hispida aliter glabra, 1-4 cm. longa, majora 1.5-2 cm.

lata. Capitula pauca, tenuiter pedunculata pedunculis glabratis

±1 dm. longis, radiata, pansa ad anthesin 2.5-4 cm. lata et ±9 mm.

alta. Involucri bracteae exteriores ±6, lineari-oblongae, ciliatae,

obtusae, demum patentes vel reflexae, 2-4 (-6 ?) mm. longae; interi-

ores ovatae 4-8 mm. longae. Flores ligulati plerumque 8, flavi,

ligula cuneato-obovati, apice 3-lobati, 1-1.5 cm. longi. Paleae

tenuiter lineari-oblongae. Florum tubulosorum stylorum rami

442 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

apice obtusissime brevi-conici. Achaenia non visa. Ovaria plana,

subalata, biaristata aristis nudis.

Type specimen: Collected byBosc in Carolina (Par.?).

Distribution: South Carolina and Georgia, also (fide F. E.

Boynt. in Small Fl. S. E. United States 1279. 1903) in Florida.

Specimens examined : Biltmore Herb. 14759a, low, flat land along

St. Mary's River, Folkston, Georgia, August 27, 1904 (N.Y.);

A. W. Chapman, banks of Spring-brook, Decatur County, Georgia

(N.Y.); Ravenel, damp, rich soils, Santee Canal, South Carolina,

September (Gray). In addition, I have seen the fragment (Gray)

obtained by Asa Gray at the DeCandolle Herbarium, doubtless

from a cotype specimen.

A species very rare in herbaria and imperfectly known as to

fruiting characters. Boynton (loc. cit.) classes it, in his key, with

the species having achenial margins fimbriate or dissected-winged.

His specific description (p. 1279) omits, however, all reference to

achenes.

109. Coreopsis nudata Nutt. Gen. N. Amer. PL 2: 180. 1818;

Hook. f. Bot. Mag. pi. 6419. 1929. Calliopsis nudata (Nutt.) Spreng.

Syst. Veg. 3: 611. 1826. Conopsis nudata Nutt. ex Less. Syn. Gen.

Compos. 228. 1832 (sphalm).

Herba perennis, glaberrima, 6-12 dm. alta, caulibus e radice fere

tuberosa erectis, teretibus, flexuosis, summam versus corymbose

pauciramosis, internodiis elongatis. Folia alterna, simplicia, anguste

linearia, plus minusve teretia et juncoidea, basalia elongatissima

saepe 3-4 dm. longa, mediana superioraque ±1 dm. longa, summa

subulata ±1 cm. longa. Capitula pauca, radiata, longe tenuiterque

pedunculata, pansa ad anthesin 4-6 cm. lata et circ. 7-8 mm. alta.

Involucri bracteae exteriores 6-9, plus minusve irregulariter positae,

lanceolatae, medio longitudinaliter vittatae, marginibus diaphanae,

basi subauriculatae, tergo saepe subscabridae, 2-4 mm. longae;

interiores ovatae, saepe subscabridae, 7-10 mm. longae. Flores

ligulati circ. 8, rosei, obovato-cuneati, apice 3-lobati lobis lateralibus

rotundatis mediano plerumque multo majore truncate vel emargi-

nato, 1.5-2.2 cm. longi. Paleae late lineari-oblongae, apice subacutae,

circ. 5-6 mm. longae. Flores tubulosi corolla flavidi, stylorum ramis

obtuse truncato-conicis. Achaenia subplana oblonga, faciebus nigra,

marginibus irregulariter pectinato-dentata dentibus saepe 2-4-

congregatis subbrunneis plus minusve planis apice obtusissimis

quam corporis diametro saltern dimidio brevioribus, corpore circ.

REVISION OF THE GENUS COREOPSIS 443

2.5-3.1 mm. longa et (dentibus lateralibus inclusis) 1-1.4 mm. lata,

apice biaristata (aristis linearibus stramineis antrorsum hispidis

0.6-1 mm. longis) atque inter aristas poculo minuto central! coronata.

Type specimen: Collected by William Baldwin, near St. Mary's,

"west Florida."

Distribution: Southern Georgia and northernmost Florida.

Specimens examined: Biltmore Herb. 2070b, bordering pond in

pine barrens, near Apalachicola, Florida, April 14, 1894 (Gray);

A. W. Chapman, Apalachicola, Florida (Gray; N.Y., 2 sheets);

herb, eiusdem 2070b, bordering ponds in pine barrens, near Apa-

lachicola, Florida, April 14, 1894 (Field; N.Y.; cf. Biltmore Herb.

2070b); A. H. Curtiss 1484, pine barren swamps, near Jacksonville,

Florida, May (Berl.; Field, 3 sheets; Gray) ; idem 4488, near Jackson-

ville, April 29, 1893 (N.Y.); idem 4713, swampy places in pine

barrens, near Jacksonville, April 21, 1894 (Field ; Gray) ; Asa Gray,

Apalachicola, Florida, April, 1875 (Gray); Roland M. Harper 1001,

alt. 70 meters, wet pine barrens near Collins, Georgia, July 4, 1901

(Berl.; Gray; N.Y.); idem 2198, cypress pond near Douglas, Coffee

County, Georgia, May 11, 1904 (Berl.; Gray; N.Y.); Albert S.

Hitchcock, Jacksonville, Florida (Field); B. F. Saurman, wet pine

barrens, Apalachicola, Florida (Field); John K. Small, John B.

DeWinkeler, & Charles A. M osier 11252, Apalachicola prairies,

Florida, April 25, 1924 (N.Y.); John Donnell Smith, near Starke,

Florida, April 9, 1880 (Field) ; C. S. Williamson, Waycross, Georgia,

April 15 (N.Y.).

110. Coreopsis Mildbraedii Muschler, Wiss. Ergebn. Deutsch.

Zentr. Afr.-Exped. 1907-1908, 2: 381. 1911.

Herba perennis vel suffruticosa, erecta, 1-1.5 m. alta; cauli-

bus ramisque subhirtis vel deinde mox glabratis glaberrimisve,

striato-sulcatis, teretibus, plus minusve ochraceis. Folia opposita

petiolata petiolis concavis basi subvaginatis et linea prominente ad

medium adpresse retrorsumque hirsuta conjunctis usque ad 2 cm.

longis, petiolo adjecto 8-17 cm. longa, semel bis vel terne pinnati-

secta, segmentis principalibus circumambitu ovatis vel rhomboideo-

lanceolatis, faciebus dense glanduloso-punctulatis sed glabratis,

marginibus ciliatis ac perspicuissime dentatis (dentibus elongatis

saepius lineari-lanceolatis apicaliter acerrimis saepe 1-1.5 cm.

longis) apice acerrime elongato-acuminatis. Capitula subnumerosa,

tenuiter pedicellata pedicellis (1-3-bracteatis vix ultra 0.75 mm.

crassis, infra glabris supra hispidis) saepius 4-12 cm. longis, radiata,

444 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

pansa ad anthesin ±4.5 cm. lata et 11 mm. alta. Involucri bracteae

exteriores (1-3 proximae vere pedicella genitae exclusae) circ. 6-9,

virides, elongato-lineares, dorso glabratae, marginibus infra plerum-

que glabratae supra spinuloso-ciliatae, apice induratae, saepius

1.3-1.8 cm. longae et circ. 1.5 mm. latae; interiores ovatae, margi-

naliter diaphano-flavae alibi intense purpurascenti-atrae, tergo

pubescentes, apice pulverulentae, tantum 5-8 mm. longae. Flores

ligulati verisimiliter 8-10, flavi, ligula elliptico-oblanceolati, apice

minute denticulati, circ. 2 cm. longi et 4-6 mm. lati. Paleae oblongo-

lineares, pluristriatae, apice saepe subabrupte acutae, 6-8 mm.

longae. Disci florum stigmata terminaliter elongato-caudata.

Achaenia obovato-oblonga vel lineari-oblonga, obcompressa, apicem

versus rarius contracta, atra, unaquaque facierum circ. 8-sulcata et

sparsim erecto-setosa, marginibus anguste alata alis brunneis

antrorsum setosis, corpore ipso 4.5-6 mm. longa et 1.5-2 mm. lata,

apice dense breviterque erecto-setoso biaristata aristis erectis

tenuibus sursum hispidis 1-2.5 mm. longis.

Type specimen: Collected by Mildbraed (Exped. d. Herzogs Ad.

Friedr. z. Mecklenburg Afr. 1907-1908), No. 2539, growing 1-1.5

meters high at altitude of 3,000 meters, at edge of mountain forest

and near the Ericaceae, Butagu Valley, Ruwenzori West, Belgian

Congo, middle of February, 1908 (Bert.).

Distribution: Known only from type locality in Belgian Congo.

Specimens examined: JeanLebrun 4507, alt. 2,475 meters, Valley

of the Nyamuwamba, Ruwenzori, Belgian Congo, November, 1931

(Mus. Cong., 2 sheets); Mildbraed 2539 (type, Berl.).

The achene wings are narrow and thickish, offering an approach

to Bidens. The species is at once distinguished from all other species

in both Bidens and Coreopsis by its (sharply cut) foliage. The

exceptionally elongate exterior involucral bracts are an additional

aid in identification. Lebrun's label describes the plant as having

a strong odor of tansy (Tanacetum vulgare L.).

111. Coreopsis bracteosa Sherff, Bot. Gaz. 76: 88. 1923.

Herba glabra, verisimiliter perennis et 7-10 dm. alta, caulis

internodiis folia superantibus, mediis inferioribusque saltern 18-22

cm. longis. Folia sessilia, indivisa, linearia, 6-20 cm. longa et 4-10

mm. lata, in apicem acutum sensim angustata, sparsim dentata

dentibus saepe ad spinas vel etiam ad pilos spinulosos flexiles reduc-

tis, margine revoluta et interdum minutissime spinuloso-ciliata.

Capitula pedunculata pedunculis maximam partem glabris usque ad

REVISION OF THE GENUS COREOPSIS • 445

5 cm. longis, radiata, pansa ad anthesin 3.5-4.5 cm. lata et 6-9 mm.

alta. Involucri bracteae exteriores numerosae 19-21, anguste

lineares, margine spinuloso-ciliatae, aliter glabrae, apice acerrime

apiculatae, 1-1.7 cm. longae, interioribus lanceolatis dorso hispidis

(pilis inferne squamellato-dilatatis) plerumque dimidio longiores.

Flores ligulati circ. 12, ligula elliptici, lutei, apice denticulati, cire.

1.7-2 cm. longi et 5-7 mm. lati. Paleae oblongo-lanceolatae, apiee

subito angustatae, costa mediana brunnea perspicua. Disci florum

stigmata caudato-apiculata. Achaenia atra, linearia vel oblonga,

valde obcompressa, plerumque alata, margine et supra ad faciem

sursum hispida, corpore circ. 4-5 mm. longa et (alis inclusis) 1-1.5

mm. lata, biaristata aristis antrorsum hispidis 0.6-3 mm. longis.

Planta Bidenti chaetodontae Sherff affinis sed foliis indivisis non

pinnatipartitis, involucri bracteis exterioribus 19-21 non tantum 12,

achaeniis plerumque alatis non exalatis, etc. differt.

Type specimen: Collected by Hans Meyer, No. 532, at altitude of

1,200-1,400 meters, bush and tree-grass-steppe, Jhangiro and

Karagwe, German East Africa, summer of 1911 (Berl.).

Distribution: British and German East Africa.

Specimens examined: Meyer 532 (type, Berl.); A. Whyte, on

5th march from Nandi to Kagamagas, Tropical East Africa, 1898

(Kew).

112. Coreopsis Prestinaria Schz. Bip. in Walpers, Repert. 6:

163. 1846. Prestinaria Bidentoides Schz. Bip. loc. cit. Verbesina

veris A. Richard, Tent. Fl. Abyssin. 1: 407. 1847. C. Prestinaria f.

typica Vatke and f. latisecta Vatke, Linnaea 39: 498. 1875.

Herba annua, subglabra, erecta, 2-6 dm. alta, ramis tenuibus

saepe subnumerosis. Folia opposita, inferiora (ac interdum mediana)

tenuiter petiolata petiolis saepius glabratis usque ad 2 cm. longis,

superiora (ac plerumque mediana) breviter alato-petiolata vel sub-

sessilia, omnia 3-6 cm. longa, circumambitu triangulata vel ovato-

triangulata, bipinnatisecta, lobis membranaceis linearibus vel

oblongo-lanceolatis glabris vel aegre ciliatis apice calloso-mucronatis.

Capitula tenuiter pedicellata pedicellis usque ad 6 cm. longis, sub-

numerosa, radiata, pansa ad anthesin 2-3 cm. lata et circ. 8 mm. alta.

Involucri glabri bracteae exteriores 8-10, uniseriales, lineares vel

lineari-spathulatae, trinerviae, calloso-apiculatae, circ. 6 mm.

longae; interiores oblongo-lanceolatae, badiae, margine diaphanae,

saepe paulo longiores. Flores ligulati circ. 8, flavi, ligula lineari-

elliptici, apice integri, circ. 1-1.4 cm. longi et 3-4 mm. lati. Paleae

446 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

anguste oblongae, nitidae, paucistriatae, usque ad circ. 12 mm. longae

et circ. 1.5 mm. latae. Disci florum stigmata tenuia terminaliter

angustata. Achaenia obcompressa, corpore lineari-fusiformia vel

lineari-clavata nigra faciebus adpresse erecteque papillato-setosa

6-9 mm. longa et 0.9-1.1 mm. lata, lateraliter alata alis angustis

stramineis vel subbadiis superne 0.3-0.5 mm. latis marginibus

exterioribus erecto-ciliatis, apice erecto-setosa et biaristata aristis

subulatis stramineis antrorsum hispidulis 4-6 mm. longis.

Type specimen: Collected by Wilhelm Schimper, No. 62, near

Djeladjeranne (Dscheladscheranne), Abyssinia, 1840 (Par.; nomen

vulg., Gellgelle Maskal).

Distribution: Abyssinia.

Specimens examined: J. M. Hildebrandt 417, during rainy

season, September, 1872 (Berl.); A. Petit, Abyssinia (Berl., 3 sheets;

Par., 3 sheets); Quartin Dillon 612, Abyssinia, November, 1840

(Par., 2 sheets); Schimper 62 (type, Par.; cotypes, Berl., 4 sheets;

Gray; N.Y.; Par., 2 sheets); idem 429, mountains and valleys at

1,860 meters alt., Gowwo Soria, Tigre, Abyssinia, September 26,

1862 (Berl., 2 sheets; type collection of f. latisecta Vatke); H. Steud-

ner 349, Keren, etc., Tigre, Abyssinia (Berl.; nom. tigrense, Ade").

Often confused with C. macrantha but easily distinguished by its

annual habit, lower stature, smaller flowering heads, smooth inner

involucral bracts, more narrowly winged achenes, etc. The f.

latisecta Vatke has leaf divisions slightly wider than in the type speci-

mens but apparently is only a robust state of the species.

Vatke (loc. cit.) confused C. macrantha with this species, as is

shown by his inclusion of Schimper 71 and 1423 under C. Prestinaria.

Oliver and Hiern (in Oliv. Fl. Trop. Afr. 3: 387. 1877) stated incor-

rectly that the aristae exceeded the achene in length. Seldom do

they even equal it, although the coincidence of certain aristae with

others underneath in a mature fruiting head often gives an exag-

gerated effect to the aristal length.

The native names recorded by Schimper and others for this and

closely similar species are: Gellgelle Maskal (Schimp.; spelled

Gueulguel-Makhul, A. Rich. loc. cit.); Ad£ (Steudner; spelled Adde",

A. Rich. loc. cit.); Embobahgad£ (A. Rich. loc. cit.).

113. Coreopsis macrantha Schz. Bip. in Walpers, Repert. 6:

163. 1846. Verbesina macrantha (Schz. Bip.) A. Rich. Tent. Fl. Abys-

sin. 1: 408. 1847. C. macroptera Schz. Bip. ex Schweinf. & Aschers.

in Schweinf. Beitr. Fl. Aethiop. 284. 1867 (nomen subnudum);

REVISION OF THE GENUS COREOPSIS 447

cf. Pirotta, Fl. Delia Colonia Eritrea 1: 184. 1904. C. Prestinaria f.>

elatior Vatke, Linnaea 39: 498. 1875 (excl. Schimperi plantam n. 22).

Prestinaria macrantha Schz. Bip. ex 0. & H. in Oliver, Fl. Trop. Afr.

3: 391. 1877.

Herba perennis, adscendenti-erecta e radice lignescenti fibrosa

multicauli, 3-15 dm. alta, caulibus subtetragonis vel subteretibus,

glabris, internodiis quam foliis saepius multo longioribus. Folia

opposita, plemmque breviter petiolata petiolis glabris late alato

marginatis tantum 2-8 mm. longis, petiolo adjecto 3-8 cm. longa,

bipinnatisecta, lobis linearibus glabris membranaceis, corpusculo

subcorneo terminatis, saepius 1.5-4 mm. latis. Capitula tenuiter

pedicellata pedicellis glaberrimis usque ad 18 cm. longis, radiata,

pansa ad anthesin 3-4 (-5) cm. lata et 8-13 mm. alta. Involucri

bracteae subaequilongae, exteriores 9-13 saepe subbiseriales, lanceo-

lato-lineares vel rarius anguste lineares, 3- (vel sub 5-) nerviae,

omnino virides vel marginibus angustis albescentes, glabrae, apice

subabrupte acutae, circ. 8-13 (rarius -18) mm. longae; interiores

oblongo-lanceolatae, coloratae, marginibus diaphanae, ad tergi

costam medianam hispido-fimbriatae. Flores ligulati circ. 8-10

(12 fide Schultzii), flavi, ligula ovato-oblongi, apice minute denti-

culati, 1.7-2.5 cm. longi et 5-10 mm. lati. Paleae lineari-oblongae,

glabrae, 1-1.4 cm. longae. Disci florum stigmata caudato-appendi-

culati. Achaenia plana, corpore nigra anguste lineari-oblonga ad-

presse erecto-setosa 6-8 (-13) mm. longa et 0.9-1.2 mm. lata;

margine moderate vel latissime alata alis hispidis et densissime

erecto-ciliatis supra interdum 2-4 mm. latis; apice setis numerosis

albescentibus erectis nitidis elongatis ac 2 aristis erectis subulatis

stramineis antrorsum hispidis 3-7 mm. longis coronata.

Type specimen: Collected by Wilhelm Schimper (sect. 2, No. 1236,

Enschedcap, Abyssinia (Par.).

Distribution: Eritrea and Abyssinia.

Specimens examined: Anon., cult, in Hort. Berol., 1860 (Berl.;

Par.); Antinori, Province of Scioa (Shoa), Abyssinia, 1878 (Flor.);

Emilio Chiovenda, District of Semien, Province of Amhara, Abys-

sinia, December 3, 1909 (Flor.); idem 1820, District of Dembia

(Dembea), Province of Amhara, September 3, 1909 (Flor.); idem

2205, Gondar, Province of Amhara, September 25, 1909 (Flor.);

Quartin Dillon, Abyssinia (Par.) ;Ellenbeck (Exped. Baron vonErlanger)

1660, alt. 3,500 meters, Schoa, Province of Schoa (Shoa), Abyssinia,

November 16, 1900 (Berl.); Adriano Fiori 1826, alt. 2,000 meters,

Hamasen region, Eritrea, April 1, 1909 (Flor.) ; G. Negri 56, Province

448 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

of Scioa, Abyssinia, 1909 (Flor.) ; idem 235 bis, eodem loco et tempore

(Flor.); idem 618, alt. 2,650 meters, eodem loco (Flor.); 0. Neumann

75, alt. 1,800-2,300 meters, between Kontschu and Tomata (Gudji),

Abyssinia, December 24, 1900 (BerL; forma involucre exteriore C.

Prestinariae adpropinquans) ; A. Pappi 1194, alt. 2,800-3,000

meters, toward the Torrent of Arigot, Mount Mamahot, Soyra

Mountains, August 23, 1902 (Flor.); idem 1303, alt. 2,600-2,800

meters, Torrent of Arigot, Soyra Mountains, Ocule' Cusae, Eritrea,

August 23, 1902 (Flor.); idem 2642, Eritrea, July 14-15, 1902

(Flor.); A. Petit, Abyssinia (Berl., 2 sheets); V. Ragazzi, Antoto,

Province of Scioa, October 2, 1887 (Flor.); idem, Let-Marefia,

Province of Scioa, September 4, 1886 (Flor.); idem, eodem loco,

September 6, 1886 (Flor.); idem 250, alt. 2,000 meters, Amasen

Quazien west of Dorfu, Eritrea, September, 1892 (Flor.); Rohlfs

& Sleeker, alt. 2,450 meters, Katz, Abyssinia, December 30, 1880

(Berl., 2 sheets) ; Rilppell, between Temben and Siemen, Abyssinia,

June or July, 1832 (Par.); W. Schimper (itin. abyssin. sect. 5. n. 22,

edit. Buchinger, 1854), Abyssinia (Berl.; Del.; Kew; Par., 3 sheets;

type material of Coreopsis macroptera Schz. Bip.); idem 4$, in fields,

alt. 2,790 meters, Dibra Eski (Siemen), Abyssinia, November 6,

1850 (Par., 2 sheets; "blooming in August and September"); idem

71, alt. 1,890 meters, mountains and valleys, Mai Sigamo, Abyssinia,

September 26, 1862 (Berl., 3 sheets); idem 85, Abyssinia (Par.);

idem (itin. abyssin. sect. 5. n. 95, edit. Buchinger, 1854), Abyssinia

(Par.); idem (itin. abyssin. sect. 5. n. 1005, edit. Buchinger, 1854),

Province of Agow, Abyssinia, 1853 (Par., 3 sheets); idem 1236 (type,

Par.; cotypes, Berl., 2 sheets; N.Y.; Par.); idem 1423, alt. 3,150

meters, Mount Gunna, Abyssinia, December 18, 1863 (Berl. ; Kew) ;

Hugh Scott, alt. 2,700-2,900 meters, Mount Zuguala, Abyssinia,

October 21-27, 1926 (Kew) ; H. Steudner 253, Province of Amhara,

May 20, 1862 (Berl.); A. Terracciano & A. Pappi 2474, alt. about

2,000 meters, Alibaret-Sella Mogasas, Bogos, Eritrea, February 5,

1893 (Flor.).

Vatke (loc. cit.) appears to have completely ignored the perennial

habit and proportionately shorter achaenial aristae of this species.

He merged it (as f . elatior) with C. Prestinaria, a course not followed

by Oliver and Hiern, or by Otto Hoffmann, or by other subsequent

students of the group.

C. macroptera was known to Oliver and Hiern (in Oliv. Fl. Trop.

Afr. 3: 391. 1877) only through achenes, and they referred it inter-

rogatively to C. macrantha Schz. Bip. Schultz Bipontinus' private

REVISION OF THE GENUS COREOPSIS 449

herbarium (Par.) contains several sheets of excellent material

labeled C. macroptera in his own writing. These, also several found

elsewhere under his determination C. macroptera, have thicker and

taller stems, mostly different leaves, mostly larger capitula with the

larger, exterior bracts mostly more numerous and with their margins

more conspicuously whitish, etc. (Folia subsessilia, 4-8 cm. longa,

nunc bipinnatisecta lobis linearibus acriter apiculatis 1.5-5 mm.

latis, nunc anguste oblongo-lanceolata lamina ipsa 1-1.5 cm. lata

marginibus dentibus vel lobis linearibus perspicuissime elongatis

[etiam 4.5 cm. longis] munita.) But a study of numerous other

specimens more recently collected, especially by various Italian

collectors, shows all intergradations, unless as to foliage. (In C.

macroptera some of the principal leaves appeared to have a petiole

and midrib grossly expanded into an abnormal blade, this irregularly

toothed with greatly elongate, linear lobes. The appearance was

unnatural and apparently teratological. However, we may con-

sider this character as of no important value, since C. Prestinaria is

known to produce at times a precisely comparable leaf -form. — More-

over, it may be noted that some authors who have retained C. macrop-

tera have overlooked even this one seeming distinction. Thus

R. Pirotta [Fl. Delia Colonia Eritrea 1: 184. 1904] lists C. macroptera

and cites for it Terracciano & Pappi 2474, a plant which is typical

C. macrantha.)

114. Coreopsis pachyloma 0. & H. in Oliver, Fl. Trop. Afr.

3: 391. 1877. C. involucrata Schz. Bip. in Walpers, Repert. 6: 163.

1846 (non Nutt. Journ. Acad. Phila. 7: 74. 1834). Verbesina involu-

crata (Schz. Bip.) A. Rich. Tent. Fl. Abyssin. 1: 409. 1847. C.

callosa Schz. Bip. in Schweinf. & Aschers. Enum. 284. 1867.

Achaenia biaristata C. pachyloma sensu stricto.

Achaenia exaristata var. /3 inanis.

Herba annua, erecta vel adscendens, ramosa, 3-7 dm. alta; caule

ramisque (siccis) viridi-stramineis, perspicue striatis, nunc glabratis

nunc hispidis, nitidis. Folia opposita tenuiter petiolata petiolis ba-

saliter connatis interdum hispidis usque ad 2 cm. longis, petiolo ad-

jecto usque ad 6 cm. longa, circumambitu late oblonga vel deltoideo-

ovata, bipinnatipartita, lobis valde membranaceis, nervis margini-

busque subsparsim setosis, saepe 1 vel 2 dentibus acriter serratis,

abrupte mucronulatis, saepius 2-4 mm. latis. Capitula subcorym-

bosa, pedunculata pedunculis tenuibus hispidis 5-14 cm. longis,

radiata, pansa ad anthesin circ. 2 cm. lata et 6-8 mm. alta. Involucri

bracteae exteriores circ. 8, lineari-spathulatae, valde foliaceae, facie-

450 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

bus glabrae vel raro subhispidae, marginibus hispido-ciliatae, apice

acuto vel subobtuso flavido-albidae ac cartilagineo-induratae, 7-12

mm. longae et superne 1-2.5 mm. latae: interiores ovato-oblongae

dorso hispidae saepe paulo vel etiam fere dimidio breviores. Flores

ligulati 6-8, flavi, ligula elliptico-oblanceolati, 6-12-striati, apice

integri vel denticulati, 8-10 mm. longi et 3-5 mm. lati. Paleae

subalbidae convexe conduplicatae, acutae, 7-9 mm. longae. Disci

florum stigmata terminaliter caudata. Achaenia corpore ipso

dorsaliter lineari-oblanceolata, moderate obcompressa, nigra (nitida),

glabra vel supra erecto-setosa, 4.5-5.5 mm. longa et dorsaliter fere

1 mm. lata, lateraliter alis stramineis crassis erecte hispidis margi-

naliter rotundatis ventraliter fere 0.5 mm. latis bialata, apice biaristata

aristis lanceolato-subulatis, hyalinis, infra valde membranaceis supra

rigidis acerrimisque, marginibus minute erecto-ciliolatis, 4-5.5 mm.

longis.

Type specimen: Collected by Wilhelm Schimper in Abyssinia

(Kew).

Distribution: Abyssinia.

Specimens examined: Anon., under bushes at Gaffat near Debra

Tabor, May 10, 1862 (Berl.); Eduard Ruppell, between Temben and

Siemen, June or July, 1832 (type of C. involucrata Schz. Bip., Par.);

Schimper (itin. abyssin.) sect. 5. nos. 15 (Del.; Par.) and 933 (Par.).

An annual with the foliar habit of Bidens pilosa var. bimucronata

f. odorata (Cav.) Sherff. The achenial characters separate it at once

from other species of Coreopsis.

Coreopsis pachyloma var. ft inanis Sherff, Bot. Gaz. 90:

386. 1930.

A specie achaeniis etiam primo exaristatis differt.

Type specimen: Collected by Wilhelm Schimper, No. 933 pro

parte, Abyssinia, commun. 1853 (Par.).

Distribution: Abyssinia.

Specimen examined: Schimper 933 pro parte (type, Par.).

PLANTAE EXCLUSAE

Coreopsis abyssinica Schz. Bip. ex Walp. Repert. 6: 163. 1846=

Bidens chaetodonta Sherff, Bot. Gaz. 90: 387. 1930.

Coreopsis abyssinica var. bipinnato-partita Chiov. in Pirotta, Fl.

Colon. Eritrea 185. l9Q4:=Bidens setigera (Schz. Bip.) Sherff, op. cit.

90: 390. 1930.

REVISION OF THE GENUS COREOPSIS 451

Coreopsis abyssinica var. glabrior 0. & H. in Oliv. Fl. Trop. Afr.

3: 389. l877=Bidens chaetodonta var. glabrior (0. & H. in Oliv.)

Sherff, op. cit. 90: 388. 1930.

Coreopsis abyssinica f. latisecta Vatke, Linnaea 39: 499. 1875

(nom. subnudum)=5zdews articulata Sherff, op. cit. 94: 591. 1932.

Coreopsis Acmella (L.) E. H. L. Krause, Beih. Bot. Centralbl.

32, pt. 2: 340. l9U=Spilanthes Acmella (L.) Murr. Syst. Veg. ed.

13. 610. 1774.

Coreopsis Acmella var. uliginosa E. H. L. Krause, loc. cit.=

Spilanthes uliginosa Swartz, Prodr. Veg. Ind. Occ. 110. 1788.

Coreopsis alata Cav. Icon, et Descr. 3: 30, pi. 260. 1795= Ver-

besina occidentalis (L.) Walt. Fl. Carol. 213. 1788.

Coreopsis alba L. Sp. PI. 908. l753.=Bidens pilosa var. radiata L

Dondiaefolia (Less.) Sherff, Bot. Gaz. 97: 607. 1936.

Coreopsis alternifolia L. op. cit. 909. YI53=Actinomeris alterni-

folia (L.) DC. Prodr. 5: 575. 1836.

Coreopsis ambacensis Hiern, Cat. Welw. Afr. PL 1, pt. 3: 586.

1898=Bidens ambacensis (Hiern) Sherff, Bot. Gaz. 59: 309. 1915.

Coreopsis ambigua Nutt. Journ. Acad. Phila. 7: 75. lSB4=Bidens

mitis (Michx.) Sherff, op. cit. 81: 43. 1926.

Coreopsis amplexicaulis Cav. Descr. 226. l8Q2=Simsia foetida

(Cav.) Blake, Proc. Amer. Acad. n. ser. 41: 385. 1913.

Coreopsis Anagallis E. H. L. Krause, Beih. Bot. Centralbl. 32,

pt. 2: 344. Vd\±=Erihydra Anagallis Gardn. Lond. Journ. Bot. 7:

409. 1848.

Coreopsis Anthemoides DC. Prodr. 5: 573. 1836=Bidens Anthe-

moides (DC.) Sherff, Bot. Gaz. 56: 493. 1913.

Coreopsis arenicola S. L. Moore, Journ. Linn. Soc. 37: 170. 1905=

Bidens musoziana Sherff, op. cit. 59: 309. 1915.

Coreopsis arguta Pursh, Fl. Amer. Sept. 2: 567. 1814=Bidews

mitis (Michx.) Sherff, op. cit. 81: 43. 1926.

Coreopsis aristata Willd. Sp. PI. 3: 2253. 18Q4:=Bidens aristosa

(Michx.) Britt. Bull. Torr. Bot. Club 20: 281. 1893.

Coreopsis aristosa Michx. Fl. Bor. Amer. 2: 140. 18Q3=Bidens

aristosa (Michx.) Britt. loc. cit.

Coreopsis aristosa var. mutica A. Gray, Syn. Fl. N. Amer. 1, pt.

2: 295. 1884:=Bidens aristosa var. mutica A. Gray ex Gattinger, Fl.

Tenn. 172. 1901.

452 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Coreopsis arizonica (A. Gray) O. Hoffm. in Engl. & Prantl, Nat.

Pflanzenf. 4, pt. 5: 243. 189Q=Coreocarpus arizonica (A. Gray)

Blake, Proc. Amer. Acad. n. ser. 49: 344. 1913.

Coreopsis Artemisiaefolia Jacq. Icon. PI. Rar. 3: pi. 595. 1786-

1793; Coll. Suppl. 155. 1796= Cosmos sulphureus Cav. Icon, et

Descr. 1:56. pi. 79. 1791.

Coreopsis Artemisifolia Sess£ & Moc. Fl. Nov. Hispan. ed. 2.

137. 1893= Cosmos sulphureus Cav. loc. cit.

Coreopsis asperata Hutch. & Dalz. Fl. West Trop. Afr. 2: 141-143.

1931; cf. Sherff, Bot. Gaz. 93: 219. l932=Bidens asperata (Hutch.

& Dalz.) Sherff, loc. cit. 93: 220. 1932.

Coreopsis Aspilioides Bak. Kew Bull. 153. 1898=Bidens Aspilioides

(Bak.) Sherff, op. cit. 94: 590. 1932.

Coreopsis aurea Dryand. in Ait. Hort. Kew. ed. 1, 3: 252. 1789=

Bidens aurea (Dryand. in Ait.) Sherff, op. cit. 59: 313. 1915.

Coreopsis Bacana Heyne ex DC. Prodr. 5: 632. \836=Glossogyne

pinnatifida (Buch.-Ham. in Wall.) DC. in Wight, Contr. 19. 1834.

Coreopsis baccata L. PI. Surin. 14. 1775; Amoen. Acad. 8: 262.

H85=Wulffia stenoglossa DC. 5: 563. 1836.

Coreopsis Baumii 0. Hoffm. in H. Baum, Warb. Kunene-Zambesi

Exped. 419. lW3=Bidens Baumii (0. Hoffm.) Sherff, Bot. Gaz. 59:

309. 1915.

Coreopsis Bidens L. Sp. PL 908. 1753=Bidens cernua L. op. cit. 832.

Coreopsis Bidentoides Nutt. ex Torr. & Gray, Fl. N. Amer. 2:

339. 1843=Bidens Bidentoides (Nutt.) Britt. Bull. Torr. Bot. Club 20:

281. 1893.

Coreopsis biternata Lour. Fl. Cochinch. ed. 1. 508. 179Q=Bidens

biternata (Lour.) Merr. & Sherff ex Sherff, Bot. Gaz. 88: 293. 1929.

Coreopsis Buchingeri Schz. Bip. in Schweinf. & Aschers. Enum.

284, nomen=Bidens rotata Sherff, op. cit. 90: 391. 1930.

Coreopsis Buchneri Klatt, Leopoldina 25: 107. 1889=Bidens

Buchneri (Klatt) Sherff, op. cit. 76: 158. 1923.

Coreopsis Buchneri Klatt, Ann. Naturh. Hofmus. Wien 7: 103.

l892=Bidens grandis Sherff, op. cit. 59: 309. 1915.

Coreopsis chrysantha Vatke, Linnaea 39: 499. 1875=Bidens

Vatkei Sherff, op. cit. 90: 388. 1930.

Coreopsis chrysantha var. simplicifolia Vatke, op. cit. 500=

Bidens Dielsii Sherff, op. cit. 90: 388. 1930.

453

Coreopsis cordifolia (Schz. Bip.) Drake del Cast. 111. Fl. Ins. Mar.

Pacif. 208. 1890; Fl. Polyn. Franc. 109. 1892=Bidens cordifolia

Schz. Bip. Flora 39: 361. 1856.

Coreopsis cordifolia Sess£ & Moc. PL Nov. Hispan. ed. 2. 137.

1893= Dahlia cardiophylla Blake and Sherff, nom. nov.

The type is still extant (Herb. Bot. Card. Madrid). Dr. Sidney

F. Blake, Senior Botanist of the United States Bureau of Plant

Industry, has independently examined it at my suggestion and writes:

"I agree that it is undoubtedly a hitherto unrecognized species of

Dahlia."

Coreopsis coriacea 0. Hoffm. in Engl. Pflanzenw. Ost-Afr. C:

414, 1899=Bidens coriacea (0. Hoffm.) Sherff, Bot. Gaz. 81: 52. 1926.

Coreopsis corymbifolia Buch.-Ham. ex DC. Prodr. 5: 602. 1836=

Bidens pilosa L. Sp. PI. 832. 1753.

Coreopsis Cosmoides A. Gray, Proc. Amer. Acad. 5: 126. 1861=

Bidens Cosmoides (A. Gray) Sherff, Bot. Gaz. 70: 98. 1922.

Coreopsis crassifolia Sesse* & Moc. Fl. Mex. ed. 2. 194. 1894=

Dahlia variabilis Desf. Cat. Hort. Par. ed. 3. 182. 1829. The SessS &

Mocino type came from mountains in Temascaltepec, State of

Mexico. Its leaves are simple (although the lower ones are described

as ternate). The general appearance is thus quite different from

that in Dahlia variabilis Desf. With that species, however, it is

definitely seen to belong from a study of Pringle 3165. The plant of

this number at St. Louis (Mo.) has closely matching foliage, but

the plant at Chicago (Field) has the leaves mostly 5-partite, as

very commonly found in D. variabilis.

Coreopsis Crataegifolia 0. Hoffm. Bot. Jahrb. 30: 431. 1901=

Bidens Crataegifolia (0. Hoffm.) Sherff, op. cit. 76: 158. 1923.

Coreopsis cuspidata Bertol. Misc. Bot. 7: 44. 1848 (ex descript. et

patri&)=Bidens mitis (Michx.) Sherff, op. cit. 81: 43. 1926.

Coreopsis dichotoma Drake del Cast. 111. Fl. Ins. Mar. Pacif. 209.

1890 (nee alior.)=Bidens Degeneri Sherff, op. cit. 85: 3. 1928.

Coreopsis diffusa Jones, Extracts Contr. West. Bot. 18: 73. 1933=

Chrysanthellum mexicanum Greenm. Proc. Amer. Acad. 39: 114.

1903 (forma humilis sed non aliter atypica).

Coreopsis discoidea Torr. & Gray, Fl. N. Amer. 2: 339. 1843=

Bidens discoidea (Torr. & Gr.) Britt. Bull. Torr. Bot. Club 20: 281.

1893.

454 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Coreopsis dissecta (Benth.) Blake ex Ind. Kew. Suppl. 5: 66. 1921

—Coreocarpus dissectus (Benth.) Blake, Proc. Amer. Acad. n. ser. 49:

344. 1913.

Coreopsis Elliotii S. L. Moore, Journ. Linn. Soc. 35: 346. 1902=

Bidens Elliotii (S. L. Moore) Sherff, Bot. Gaz. 59: 309. 1915.

Coreopsis exaristata 0. Hoffm. in Engler, Pflanzenw. Ost-Afr. C:

414. lS99=Bidens microcarpa Sherff, op. cit. 76: 84. 1923.

Coreopsis exaristata var. gracilior 0. Hoffm. loc. cit.=Bidens

gracilior (0. Hoffm.) Sherff, loc. cit. 1923.

Coreopsis Ferulaefolia Jacq. Hort. Schoenbr. 3: 65. 1798=Bidens

Ferulaefolia (Jacq.) DC. Prodr. 5: 603. 1836.

Coreopsis filifolia Hook. Bot. Mag. pi. 3505. 1836=Thelesperma

filifolium (Hook.) A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 301. 1884.

Coreopsis Fischeri 0. Hoffm. in Engler, Pflanzenw. Ost-Afr. C :

414. 18W=Bidens Fischeri (0. Hoffm.) Sherff, Bot. Gaz. 76: 158. 1923.

Coreopsis flammula Banks ex Steud. Nomencl. ed. 1. 108. 1821=

Bidens laevis (L.) Britton, Sterns & Poggenb. Prel. Cat. 29. 1888

(fide syn. B. Chrysanthemoidis in Steud. loc. cit.).

Coreopsis Foeniculacea Sesse" & Moc. ex DC. Prodr. 5: 603. 1836=

Bidens Ferulaefolia var. Foeniculaefolia (DC.) Sherff, Bot. Gaz. 81:

39. 1926.

Coreopsis foetida Cav. Icon, et Descr. 1: 55, pi 77. l79l=Simsia

foetida (Cav.) Blake, Proc. Amer. Acad. n. ser. 41: 385. 1913.

Coreopsis formosa Bonato Pisaura autom. Coreopsis formosa 22,

pi. 2. 1793=Cosraos bipinnatus Cav. Icon, et Descr. 1: 10, pi. 14- 1791.

Coreopsis frondosa 0. Hoffm. in Engler, Pflanzenw. Ost-Afr. C:

414. l899=Bidens magnifolia Sherff, Bot. Gaz. 90: 390. 1930.

Coreopsis fruticosa Forst. Fl. Ins. Austr. Prodr. 91. 1786, nomen=

Bidens australis Spreng. Syst. Veg. 3: 453. 1826.

Coreopsis Georgina Cass. Diet. Sci. Nat. 18: 441. 1820= Dahlia

variabilis (Willd.) Desf. Cat. Hort. Par. ed. 3. 182. 1829.

Coreopsis Georgina var. nuda Cass. op. cit. 442= Dahlia variabilis

(Willd.) Desf. loc. cit.

Coreopsis Georgina var. pruinosa Cass. op. cit. 443= Dahlia vari-

abilis (Willd.) Desf. loc. cit.

Coreopsis glaucescens O. & H. in Oliv. Fl. Trop. Afr. 3: 389. 1877=

Bidens articulata Sherff, Bot. Gaz. 94: 591. 1932.

Coreopsis Grantii Oliv. Trans. Linn. Soc. 29: 98. pi 65. 1874=

Bidens Grantii (Oliv.) Sherff, op. cit. 59: 309. 1915.

REVISION OF THE GENUS COREOPSIS 455

Coreopsis hawaiensis (A. Gray) Drake del Cast. 111. Fl. Ins. Mar.

Pacif. 209. 189Q=Bidens hawaiensis A. Gray, Proc. Amer. Acad. 5:

128. 1861.

Coreopsis heterophylla Cav. Icon, et Descr. 3: 34, pi. 268. 1795;

Descr. 226. l8Q2=Iostephane heterophylla (Cav.) Benth. in Benth.

& Hook. Gen. PL 2: 368. 1873.

Coreopsis heterophylla Bertol. Misc. Bot. 7: 43. 1848 (ex descript.

et patria)=M2ens mitis (Michx.) Sherff, Bot. Gaz. 81: 43. 1926.

Coreopsis Hillebrandiana Drake del Cast. 111. FL Ins. Mar. Pacif.

209. l89Q=Bidens Hillebrandiana (Drake del Cast.) Deg. ex Sherff,

op. cit. 85: 6. 1928.

Coreopsis hirta Raf. Ann. Nat. 15. 1820= Rudbeckia triloba L.

Sp. PL 907. 1753 (fide Torr. & Gray, Fl. N. Amer. 2: 349. 1843).

Coreopsis Hoktii 0. Hoffm. in Engler, Pflanzenw. Ost-Afr. C:

415. 18W=Bidens Holstii (0. Hoffm.) Sherff, Bot. Gaz. 76: 79. 1923.

Coreopsis incisa Ker, Bot. Reg. 1: 7 (pi.}. l8l5=Bidens incisa

(Ker) G. Don in Sweet, Hort. Brit. ed. 3. 360. 1839.

Coreopsis incurva Moench, Meth. Suppl. 245. lSQ2=Bidens

Ferulaefolia (Jacq.) DC. Prodr. 5: 603. 1836 (fide auctorum).

Coreopsis insecta S. L. Moore, Journ. Bot. 46: 42. 19Q8=Bidens

insecta (S. L. Moore) Sherff, Bot. Gaz. 59: 309. 1915.

Coreopsis involucrata Nutt. Journ. Acad. Phila. 7: 74. 1834=

Bidens polylepis Blake, Proc. Biol. Soc. Wash. 35: 78. 1922.

Coreopsis Jardinii (Schz. Bip.) Drake del Cast. 111. Fl. Ins. Mar.

Pacif. 209. 1890; FL Polyn. Franc. 109. l892=Bidens Jardinii Schz.

Bip. Flora 39: 360. 1856.

Coreopsis Jasminifolia Bertol. Misc. Bot. 7: 44. l848=Bidens

mitis (Michx.) Sherff, Bot. Gaz. 81: 43. 1926.

Coreopsis kilimandscharica 0. Hoffm. Bot. Jahrb. 20: 234. 1894=

Bidens kilimandscharica (O. Hoffm.) Sherff, op. cit. 59: 309. 1915.

Coreopsis Kirkii 0. & H. in Oliv. FL Trop. Afr. 3: 390. 1877=

Bidens Kirkii (0. & H. in Oliv.) Sherff, op. cit. 59: 309. 1915.

Coreopsis leucantha L. Sp. PL ed. 2. 1282. H63=Bidens pilosa var.

radiata Schz. Bip. in Bark.-Webb & Berth. Hist. Canar. 3, 2, part 2:

242. 1836-1850.

Coreopsis leucanthema L. Cent. PL 1: 29. 1755; Amoen. Acad. 4:

291. l759=Bidens pilosa var. radiata Schz. Bip. loc. cit.

Coreopsis leucorhiza Lour. FL Cochinch. ed. 2. 622. 1793 (ex

descript. et patr\B.}=Bidens pilosa var. minor (Bl.) Sherff, Bot. Gaz.

80: 387. 1925.

456 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Coreopsis leucorrhiza Lour. op. cit. ed. 1. 508. 1790 (ex descript. et

•pa.tria)=Bidens pilosa var. minor (Bl.) Sherff, loc. cit.

Coreopsis Liebmannii Schz. Bip. ex Hemsl. Biol. Centr. Amer.

Bot. 2: 196. l88l=Verbesina intermissa Blake, Contr. U. S. Nat.

Herb. 23: 1578. 1926.

Coreopsis limensis Jacq. Coll. 2: 299. 1788=Encelia canescens

Lam. Encycl. Meth. 2: 356. 1786 (fide Blake, Proc. Amer. Acad. n.

ser. 41: 369. 1913).

Coreopsis linearifolia DC. Prodr. 5: 570. 1836=Bidens angustis-

sima HBK. Nov. Gen. et Sp. 4: 183 (233). 1820.

Coreopsis linearifolia 0. & H. in Oliv. Fl. Trop. Afr. 3: 390. 1877=

Bidens Schweinfurthii Sherff, Bot. Gaz. 59: 309. 1915.

Coreopsis lineata Klatt, Ann. Naturh. Hofmus. Wien 7: 103. 1892

=Bidens grandis Sherff, loc. cit.

Coreopsis lucida Cav. Descr. 224. \8Q2=Bidens aurea (Dryand. in

Ait.) Sherff, op. cit. 59: 313. 1915.

Coreopsis Macraei A. Gray, Proc. Amer. Acad. 5: 126. 1861=

Bidens Campylotheca Schz. Bip. Flora 39: 359. 1856.

Coreopsis macrocarpa A. Gray, loc. cit.=Bidens macrocarpa (A.

Gray) Sherff, Bot. Gaz. 70: 97. 1920.

Coreopsis Menziesii A. Gray, op. cit. 127=Bidens Menziesii (A.

Gray) Sherff, op. cit. 70: 98. 1920.

Coreopsis micrantha (Gaud.) A. Gray, loc. cit.— Bidens micrantha

Gaud. Voy. Freycinet Bot. PI. 85 (sine descript.). 1830; ibid. 464.

Coreopsis mitis Michx. Fl. Bor. Amer. 2: 140. 18Q3=Bidens mitis

(Michx.) Sherff, Bot. Gaz. 81: 43. 1926.

Coreopsis molokaiensis (Hillebr.) Drake del Cast. 111. Fl. Ins.

Mar. Pacif. 210. l89Q=Bidens molokaiensis (Hillebr.) Sherff, op. cit.

70: 97. 1920.

Coreopsis multifida DC. et var. mutica DC. Prodr. 5: 573. 1836=

Bidens pilosa var. bimucronata f. odorata (Cav.) Sherff, op. cit. 81:

41. 1926.

Coreopsis nitida H. R. M. ex Elench. PI. Hort. Bot. J.-J. Destremx

1805: 10. 1806, nomen (ex synon.)=Bidens aurea (Dryand. in Ait.)

Sherff, op. cit. 59: 313. 1915.

Coreopsis odorata Poir. in Lam. & Poir. Encycl. Suppl. 2: 350.

1811; etiam Lam. & Poir. Atlas pi. 704, f- 1 (sine anno) -Bidens

pilosa var. bimucronata f. odorata (Cav.) Sherff, op. cit. 81: 41. 1926.

Coreopsis odoratissima Cav. ex Pers. Syn. PL 2: 477. 18Q7=Bidens

pilosa var. bimucronata f. odorata (Cav.) Sherff, loc. cit.

REVISION OF THE GENUS COREOPSIS 457

Coreopsis Oerstediana Benth. ex Oerst. Kjoeb. Vid. Medd. 93.

1852=Bidens Oerstediana (Benth. ex Oerst.) Sherff, op. cit. 80: 385.

1925.

Coreopsis ovata Cav. Icon, et Descr. 3: 41, pi. 280. 1795= Acti-

nomeris ovata (Cav.) Nutt. ex DC. Prodr. 5: 576. 1836 (cf. Nutt.

Gen. 2: 181. 1818).

Coreopsis parviflora Jacq. Hort. Schoenbr. 3: 65, pi. 374- 1798=

Cosmos parviflorus (Jacq.) HBK. Nov. Gen. et Sp. 4: 189 (241),

1820.

Coreopsis polycephala (Schz. Bip.) Benth. & Hook, ex Drake del

Cast. 111. Fl. Ins. Mar. Pacif. 217. 1890; etiam Fl. Polyn. Fr. 108.

1892=Bidens polycephala Schz. Bip. Flora 39: 360. 1856.

Coreopsis procera Dryand. in Ait. Hort. Kew. ed. 1. 3: 253. 1789=

Actinomeris alternifolia (L.) DC. Prodr. 5: 575. 1836.

Coreopsis pukhella 0. Hoffm. Bot. Jahrb. 38: 204. l9Q6=Bidens

microphylla Sherff, Bot. Gaz. 90: 390. 1930.

Coreopsis quadricornis Krock. Fl. Siles. 2, pt. 2: 481. 1793=

Bidens cernua L. Sp. PI. 832. 1753.

Coreopsis Remyi Drake del Cast. 111. Fl. Ins. Mar. Pacif. 210.

l89Q=Bidens micrantha Gaud. Voy. Freycinet Bot. PI. 85 (sine

descript.). 1830; ibid. 464.

Coreopsis reptans L. Syst. Nat. ed. 10. 2: 1228. 1759; Amoen.

Acad. 5: 381 (nomen) et 407. l76Q=Bidens reptans (L.) G. Don in

Sweet, Hort. Brit. edit. 3. 360. 1839.

Coreopsis Ridens Gunn. Fl. Norveg. 2: 87. 1772 (sphalm)-

Coreopsis Bidens Li.=Bidens cernua L. Sp. PI. 832. 1753.

Coreopsis Rueppellii Schz. Bip. ex Walp. Repert. 6: 163. 1846=

Bidens Rueppellii {Schz. Bip. ex Walp.) Sherff, Bot. Gaz. 90: 389.

1930.

Coreopsis ruwenzoriensis S. L. Moore, Journ. Linn. Soc. 35: 345.

l9Q2=Bidens coriacea(0. Hoffm.) Sherff, op. cit. 81: 52. 1926.

Coreopsis Sambucifolia Cav. Descr. 225. 18Q2=Bidens Sambucifolia

Cav. Icon, et Descript. 3: 15, pi. 229. 1795.

Coreopsis sandvicensis (Less.) Benth. & Hook, ex Drake del Cast.

111. Fl. Ins. Mar. Pacif. 210. 1890=JBz'dews sandvicensis Less. Linnaea

6: 508. 1831.

Coreopsis scandens Sess£ & Moc. PI. Nov. Hispan. ed. 2. 137.

1893; etiam Fl. Mex. ed. 2. 194. 1894=Bidens squarrosa HBK.

Nov. Gen. et Sp. 4: 187 (238). 1820.

458 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Coreopsis Schaffneri A. Gray, Proc. Amer. Acad. 19: 15. 1883=

Bidens Schaffneri (A. Gray) Sherff, Bot. Gaz. 56: 493. 1913.

Coreopsis Schkchteri (Sherff) Burtt Davy in Kew Bull. 568. 1935=

Bidens Schkchteri Sherff, Bot. Gaz. 76: 146. 1923.

Coreopsis Seretii DeWild. Ann. Mus. Congo, ser. 5. 2: 212. 1907=

Bidens Seretii (DeWild.) Sherff, op. cit. 76: 162. 1923.

Coreopsis serrata DC. Prodr. 5: 595. 1836=Bidens laevis (L.)

Britton, Sterns, & Poggenb. Prel. Cat. 29. 1888.

Coreopsis serrulata Poir. Encycl. Suppl. 2: 352. 1811=Bidens

serrulata (Poir.) Desf. Cat. Hort. Par. ed. 3. 186. 1829.

Coreopsis serrulata (Schz. Bip.) Benth. & Hook, ex Drake del

Cast. 111. Fl. Ins. Mar. Pacif. 210. 1890; etiam Fl. Polyn. Franc. 109.

1892=Bidens bipontina Sherff, Bot. Gaz. 85: 10. 1928.

Coreopsis setigera Schz. Bip. ex Walp. Repert. 6: 163. 1846=

Bidens setigera (Schz. Bip.) Sherff, op. cit. 90: 390. 1930.

Coreopsis simplicifolia (Vatke) Engler, Abh. Preuss. Akad. Wiss.

1891, 2: 435. l892=Bidens Dielsii Sherff, op. cit. 90: 388. 1930.

Coreopsis speciosa Hiern, Cat. Welw. Afr. PI. 1, pt. 3: 585. 1898=

Bidens grandis Sherff, op. cit. 59: 309. 1915.

Coreopsis Steppia Steetz in Peters, Nat. Reise Mossamb. Bot.

496. 1862-1864=Bidens Steppia (Steetz) Sherff, op. cit. 76: 82. 1923.

Coreopsis Stuhlmannii 0. Hoffm. in Engler, Pflanzenw. Ost-Afr.

C: 415. 1899=Mens Stuhlmannii (0. Hoffm.) Sherff, op. cit. 76:

158. 1923.

Coreopsis tanna Forst. ex Steud. Nomencl. ed. 2. 1: 420. 1840=

Glossogyne tenuifolia Cass. Diet. Sci. Nat. 51: 475. 1827.

Coreopsis tannensis Spreng. Fl. Hal. Mant. 53. 18Q7=Glossogyne

tenuifolia Cass. loc. cit.

Coreopsis Taylori S. L. Moore, Journ. Bot. 44: 22. 19Q6=Bidens

Taylori (S. L. Moore) Sherff, Bot. Gaz. 59: 309. 1915.

Coreopsis ternata Chiov. Ann. Bot. Roma 9: 74. 1911=Bidens

ternata (Chiov.) Sherff, op. cit. 90: 391. 1930.

Coreopsis tetragona Cerv. ex La Llave & Lex. Nov. Veg. Descr.

1: 31. 1824=Bidens aurea (Dryand. in Ait.) Sherff, op. cit. 59: 313.

1915.

Coreopsis trichosperma Michx. Fl. Bor. Amer. 2: 139. 1803=

Bidens coronata (L.) Britt. Bull. Torr. Bot, Club 20: 281. 1893; cf.

Sherff, op. cit. 56: 495. 1913.

REVISION OF THE GENUS COREOPSIS 459

Coreopsis trichosperma var. tenuiloba A. Gray, Syn. Fl. N. Amer.

1, pt. 2: 295. lSS4:=Bidens coronata var. tenuiloba (A. Gray) Sherff,

op. cit. 86: 446. 1928.

Coreopsis trifida Poir. Encycl. Suppl. 2: 353. I811=Thelesperma

filifolium A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 301. 1884.

Coreopsis trifoliata Bertol. Fl. Guat. 36. 184Q=Bidens squarrosa

HBK. Nov. Gen. & Sp. 4: 187 (238). 1820.

Coreopsis trilobata Vahl ex Klatt, Leopoldina 23: 90. 1887=

Montanoa pauciflora Klatt, loc. cit.

Coreopsis tripartita M. B. Moss, Kew Bull. 184, 196. 1929=Bidens

Mossii Sherff, Bot. Gaz. 92: 202. 1931.

Coreopsis ugandensis S. L. Moore, Journ. Linn. Soc. 35: 347. 1902

=Bidens ugandensis (S. L. Moore) Sherff, op. cit. 59: 309. 1915.

Coreopsis variifolia Salisb. Prodr. 206. !796=Bidens reptans (L.)

G. Don in Sweet, Hort. Brit. ed. 3. 360. 1839.

SPECIES INCERTAE SEDIS

Coreopsis acuta Pursh, Fl. Amer. Sept. 2: 569. 1814 (Conopsis

acuta Pursh ex Less. Syn. Gen. Compos. 228. 1832 — sphalm).

Georgia. "Is perhaps Actinomeris squarrosa Nutt." (Torr. & Gray,

Fl. N. Amer. 2: 349. 1843.)

Coreopsis aspera Pursh Fl. Amer. Sept. 2: 570. 1814. Maryland.

"Not identified and probably not of the genus" (A. Gray, Syn. Fl.

N. Amer. 1, pt. 2: 290. 1884).

Coreopsis bituminosa Noronha, Verh. Batav. Gen. 5. ed. 1. Art.

IV. 11. 1790. Malaya.

Coreopsis brasiliensis Colla, Herb. Pedem. 3: 479. 1834.

Coreopsis emarcida Noronha, Verh. Batav. Gen. 5. ed. 1. Art.

IV. 11. 1790. Malaya.

Coreopsis flexicaulis Raf. Med. Repos. N.Y. Hexade 2. 5: 361.

1808. Southern New Jersey. "Not identified and probably not of

the genus" (A. Gray, Syn. Fl. N. Amer. 1, pt. 2: 290. 1884).

Coreopsis (per errorem typ. Cereopsis) gracilis Blanco, Fl. Filip.

ed. 2. Suppl. 591. 1845. (Cf. Fernandez-Villar in Blanco, op. cit.

ed. 3. pi. 287. 1878-1880; etiam Merrill, Sp. Blanc. 380. 1918.)

Coreopsis japonica Hort. Dammann ex Wien 111. Gartenzeit. 20:

438, /. 41. 1895. Japan.

Coreopsis lanceolata var. longipes Kew Bull. App. 2: 68. 1931.

Nomen nudum.

460 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Coreopsis minima Hort. ex Steud. Nom. ed. 2. 1: 420. 1840.

North America.

Coreopsis Neaei DC. Prodr. 5: 573. 1836. South America.

Coreopsis perfoliata Walt. Fl. Carol. 214. 1788. Carolina. Cf.

Bidens laevis (L.) Britton, Sterns, & Poggenb. Prel. Cat. 29. 1888.

(The small Walter Herbarium in London (Brit.) lacks a specimen of

C. perfoliata.)

Coreopsis philadelphica L. ex Jacks. Ind. Linn. Herb. 62. 1912.

Nomen nudum.

Coreopsis radiata Mill. Gard. Diet. ed. 8. Coreopsis No. 5. 1768.

South Carolina. Perhaps Bidens laevis (L.) Britton, Sterns, &

Poggenb. Prel. Cat. 29. 1888.

Coreopsis repens Sesse* & Moc. Fl. Mex. ed. 2. 194. 1894. Mexico.

Coreopsis scabra Raf. in Robin & Raf. Fl. Ludov. 72. 1817

(Conopsis scabra Raf. ex Less. Syn. Gen. Compos. 228. 1832 — sphalm).

Louisiana.

INDEX OF COLLECTORS CITED1

Page numbers in italic type

Abrams, L. R., nos. 3342, 3628, 11788,

11915, 313

no. 9465, ±29

Achten, L., no. 253B, 387

Adams, J. WM no. 480, 423

Allard, F., nos. 65, 330, 387

Allen, J. A., Iowa, 398

Andre, E., no. 4387, 334

Anonymous, ex T. Nuttall, 409

no. 575, 418

no. 576, Arkansas, Sept., 1835, 401

Georgia, 438

mts. of N. Carolina, Sept., 1843, 416

Arkansas Nat. Forest, 1909, 355

Bar Harbor, Aug., 1906, 406

Gaffat near Debra Tabor, 1862, 450

Macon, Georgia, 438

Santa Maria R., Feb. 7, 1880, 308

Smithfield, 1871, 423

Tampa Bay, 344

cult., June, 1818, 401

cult, ex seminibus ex herb. DC.

anno 1839, 429

cult, in Hort. Berol. anno 1860, 44?

Anthony, A. W., no. 216, 306

no. 394, 305

Antinori, Prov. Scioa, 1878, 447

Armstrong, Margaret, no. 741, 306

Arsene, Bro. G., Cerro Azul, Oct., 1909,

320

nos. 3185, 5295, 6026, 6060, 318

Ashe, W. W., Chapel Hill, July, 1898,

401

Babcock, H. H., Chicago, June 17,

1874, 344

Bachelew, F. S., no. 223, 305

Bacigalupi, R., no. 1209, 309

Baikie, W. B., Niger River, 372

Bailey, L. H. & E. Z., no. 13047, 433

Bain, S. M., no. 95 pro parte, 349

no. 95 pro parte, 406

Baites, Mexico, 1864, 298

Baker, C. F., Blount Springs, May 5,

1898, 406

St. Croix Falls, Aug. 15, 1900, 398

no. 4090, 313

Ball, J., near Chicla, Apr. 21-23, 1882,

339

Barker, Carrie, no. 31, 418

Barnes, C. R., Converse, June 30, 1877,

399

Barnes, C. R., & W. J. G. Land, no.

155, 360

no. 158,317

nos. 165, 313, 301

Barnhart, J. H., no. 2715, 431

Barter, C., Borgu, 372

no. 933, 395

Bartlett, H. H., no. 1638, £09

Batchelder, C. Foster, Falmouth, Aug.

14, 1906, 423

Bates, J. M., no. 2958, 418

Battiscombe, E., no. 83, 382

no. 945, 379

Baum, H., no. 878, 387

Bavicchi, no. 14, 387

Beardslee, H. C., & C. A. Kofoid,

Great Smoky Mts., Aug., 1891, 411

Swain County, Aug., 1891, 349

Swain, Aug., 1891, .406

Bebb, M. S., District of Columbia,

1863, 401

Marion Co., 1860, 399

Winnebago Co., 1859, 399

Fountaindale, 411

Bebb, R., nos. 50, 1048, 399

no. 1174, 418

Beckwith, Florence, no. 54, 411

Becquet, A., no. 62, 391

Benke, H. C., no. 327, 118

no. 1841, 411

no. 3183, 342

no. 3279, 353

no. 3967, 399

nos. 4306, 4657, 418

Bequaert, Joseph, Leopoldville, Apr. 9,

1915, 388

no. 113, 387

no. 7205, 388

Berlandier, J. L., nos. 566, 882, 1073,

426

no. 1802, 359

nos. 1956, 2312, 2503, 2580, 426

Berlin Bot. Garden (Hort. Berol.), ex,

353

1806-1812, 350

cult., July 25, 1815, 402

Sept., 1832, 343

cult., Sept., 1834, 402

cult., 1837, 402

July, 1837, 421

cult., Sept. 11, 1876, 413

Bernoulli, Gust., no. 196, 298

1 Unnumbered specimens are cited, generally, in the order of exactness as to

geographic location, in alphabetical order as to locality, or in chronological order.

461

462 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Beyrich, H. C., banks of rivers, Georgia,

349

thickets, Georgia, 406

Bigelow, J. M., Ft. Smith, 1853-1854, 342

Ft. Smith, June 23, 1853, 344

Mohave Creek, 1853-1854, 309

Billington, C. B., no. 131, -439

Biltmore Herbarium, Hammond, Sept.

17, 1900, 427

near Pembroke, June, 1901, 438

no. 173a, 411

nos. 175a and b, 406

nos. 176, 1766, 557

no. 307c, 351

no. 437a, 401

no. 2064A, 404

no. 20666, 439

no. 2067a, 342

no. 20706, 443

no. 2139, 441

no. 53746, J>16

no. 56936, 344

no. 147586, 438

no. 14759a, 442

Bischoff , G. W., ex herb., 357

Blair, Walter, Prince Edward Co.,

June, 1881, 401

Blake, S. F., no. 9542, 402

Blakley, O. W., no. 1451, 399

no. 3420, 411

Blankinship, J. W., Southern Pines, 402

Greene Co., 411

Blanton, F. S., no. 6872, 431

Bogusch, E. R., no. 994, 421

Bolander, H. N., Julian City, SIS

no. 4520, 31 6

Bonpland, A., equatorial America, 339

Boott, William, herb, of, Winter Pond,

July 30, 1876, 423

Boriani, no. 42, 395

Bourgeau, Eugene, Saskatchewan, Sept.

17, 1857, 429

Winnipeg Valley, 1859, 399

Boykin, S. B., Georgia, 342

Boynton, F. E., Highlands, 1889, 349

Brandegee, K., San Diego, Apr. 25, 1900,

313

Redstone Park, July 25, 1905, 418

Brandegee, T. S., The Dalles, 1882, 429

Isl. Santa Cruz, Apr., 1888, 308

Alcalde, Apr., 1891, 311

San Martin Isl., Apr., 1897, 306

San Diego, June, 1900, 306

San Diegc, May, 1906, 306

no. 3368, 313

Brannon, M. A., near Crown Point,

1889, 342

Bray, W. L., no. 48, 428

Brendel, Frederick, Illinois, 342

Illinois, 399

Peoria, 399

Peoria, 411

Brewer, W. H., no. 241, 309

Bridges, Thomas, no. 260, 316

Brinton, J. B., Egg Harbor, Sept. 3,

1888, 423

Egg Harbor, Aug. 18, 1889, 424

Britton, N. L., Norfolk, June 10, 1892,

418

Britton, Mrs. N. L., no. 132, 431

Bross, Mason, Hyde Park, June 3, 1878,

342

Brown (S.), Britton (N. L.), & Bisset

(P.), no. 2011, 354

Brownfield, J., Summerville, Oct., 1892,

439

Bruce, Mrs. C. C., no. 1987, 316

Buchanan, John, no. 380, 377

Buchner, no. 32, 387

Buchtien, Otto, no. 97, 337

no. 595, 339

Buckley, S. B., southeastern U. S., 402

Alabama, 349

pine woods, mts. of Alabama, £05

Alabama, June, 1841, 357

Alabama, July, 1841, £06

mts. of Carolina and Georgia, 416

Florida, 431

Illinois, 399

Virginia, 357

Buettner, R., no. 142, 397

no. 408, 388

Burgess, A. B., no. 433, 411

no. 513, 412

Bush, B. F., no. 84, 344

no. 158, 355

no. 159, 418

no. 160, 399

no. 172A, 3.49

no. 696, 422

no. 3017, 418

no. 3249, 399

no. 3416, 397

nos. 6041, 6137, 6531, 7872, 3*9

Butler, G. D., no. 14, 418

no. 16, 353

Calkins, W. W., nos. 135, 145, 412

Cambridge Bot. Gard. (see Hort. Bot.

Harvard Univ.)

Camp, S. H. & D. R., Bay View,

July 14, 1896, 342

Jackson Co., Aug. 18, 1897, 412

Jackson Co., Aug. 18, 1898, 412

Canby, W. M., Felton, Sept., 1867, 424

near Wilmington, Oct., 1867, 441

vicin. Wilmington, Oct., 1867, £39

swamps, Delaware, Aug., 1877, 424

Florence, July 3, 1878, 409

Indian River, Feb., 1889, 431

nos. 53, 54, 357

no. 55, S44

no. 157, 418

Carey, John, North Carolina, 351

INDEX OF COLLECTORS

463

Carey, John, & Asa Gray, North

Branch of New River, etc., 351

Virginia, July, 1841, 351

Carpenter, W. M., Louisiana, 349

Carson, Alex., no. 75, 393

no. 106, 376

Chandler, H. P., no. 5123, 306

no. 7010, 1*18

Chang, Tang Chung, & Nong Sing Po,

no. 3790, 1*19

Chapman, A. W., wet barrens, 1+36

middle Florida, 344

ex herb., southern Florida, 1*39

Georgia, 1*15

mts. of Georgia, 349

mts. of Georgia, 357

mts. of Georgia, 3*16

Apalachicola, W

Spring-brook, 44#

Aspalaga, 1897, 1*1*1

ex herb., no. 1746, 1*36

ex herb., 20666, 1*1*1

ex herb., 2066c, 1*1*1

herb, of, 20706, 1*1*3

Chase, Agnes, Rogers Park, Aug. 27,

1896, 1*12

nos. 2382, 2474, 1*02

Chase, V. H., n. of Wady Petra, Aug.

12, 1896, 1*12

Cheo, H. C., & W. F. Wilson, no. 11

pro parte, 31*2

no. 11 pro parte, 1*18

no. 29, 1*18

Chevalier, Aug., nos. 2083, 9633, 9721,

395

Chiovenda, Emilio, District of Semien,

Dec. 3, 1909, 1*1*7

no. 970, 371

no. 1090, 372

no. 1148, 371

no. 1820, 1*1*7

nos. 1869, 1953, 1954, 371

no. 2205, 1*1*7

Chipp, T. F., no. 66, 362

Churchill, J. R., shore of L. Michigan,

1*12

Lookout Mt., June 2, 1911, 1*06

Claessens, J., nos. 67, 73, 88, 400, 387

Clark, H. L., Falmouth Rd., Aug. 16,

1899, 31*2

Clemens, Mr. & Mrs. Joseph, no. 978,

1*18

Clements, F. E., no. 2778, 1*18

Clements, F. E. & E. S.f nos. 224, 225,

313

no. 226, 306

Cleveland, Daniel, San Diego, 306

San Luis Rey, 1877, 313

Clokey, I. W., no. 2428, 399

Clokey, I. W. & B. Templeton, no. 4520,

309

Clothier, G. L., & H. N. Whitford,

no. 1037, 399

Cocks, R. S., Cameron, July, 1903, 1*26

no. 3631, 1*27

Coe, Mary A., Atlanta, May 6, 1903,

357

Collins, J. F., Spectacle Pond, Sept.

11, 1900, 1*21*

no. 8, 1*26

Collins, J. F., M. L. Fernald & H. H.

York, no. 294, 1*21*

Combs, Robert, & C. R. Ball, no. 517,

399

Congdon, J. W., Lincoln, Aug. 22, 1871,

424

Mariposa Water Ditch, March 28,

1901, 316

Mariposa Water Ditch, Apr. 4, 1903,

315

Conzatti, Cassiano, no. 1431, 298

nos. 1955, 2074, 299

Conzatti, Cassiano, & Gonzalez, nos.

545, 546, 299

Coombs, Mrs. A. L., Warner's Springs,

1 Q1 1 010

lifll, Old

Coulter, J. M., near LaPorte, July 22,

1878, 399

Kankakee River, 1882, 399

Coulter, Thomas, no. 316, 308

no. 339, 313

Coville, F. V., no. 127Ark., 355

Coville, F. V.,&F. Funston, no. 742, 309

Cowles, R. B., Coronado Isls., March

29, 1921, 306

Craig, Newsom, & Hiland, no. 80, 309

no. 129, 311

Cratty, R. I., Iowa, 399

Armstrong, July 18, 1897, 399

no. 16, 399

Croft, Mary B., San Diego, 1885, 1*26

Curtis, M. A., N. Carolina, 1*37

Curtis, Mrs. R. C., cult, in 1930 from

Angola seeds, 385

Curtiss, A. H., Florida, November, £37

Virginia, 1*02

Bedford Co., July 1, 1868, £07

Bedford Co., June 1, 1871, 357

Bedford Co., June 10, 1871, 401

Bedford Co., June 30, 1871, £07

Bedford Co., May 31, 1872, 357

Bedford Co., July 3, 1872, £07

Chattahooche, 1875, 439

near Jacksonville, Sept., 437

Pensacola, 1885, 31*2

Pensacola, summer of 1885, 31*6

no. 17, 349

no. 1470, 437

no. 1477 pro parte, 437

no. 1477 pro parte, 439

no. 1477 pro parte, 441

no. 1480, 433

no. 1481, 342

464 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

no. 1484, MS

no. 1485, 557

no. 1487, 1>07

no. 4487, 441

no. 4488, 445

no. 4489, 457

no. 4713, 445

no. 5266, 457

no. 5348 pro parte, 457

no. 5348 pro parte, Ml

no. 5375, 431

nos. 5568, 6278, 457

no. 6362, 359

no. 6390, 544

no. 6471, 406

no. 6734, 452

no. 6825, 404

no. 10223, 441

Curtiss, C. C., Martha's Vineyard,

Aug. 13, 1892, 424

Cuthbert, Alfred, Augusta, Sept, 25,

1898, 441

Georgia, Oct. 10, 1898, ±38

Davis, John, near LaGrange, June 9,

1915, 412

no. 1855, 406

nos. 2097, 7818, 407

Deam, C. C., & H. A. Gleason, Bluffton,

412

Deam, Mrs. C. C., no. 1709, 431

no. 1864, 421

Dedrick, Allan, dry, sandy soils, 406

De Giorgi, S., vicin. Elisabethville, 1923,

597

no. 142, 231

no. 170, 592

De Jelski, Const., no. 705, 555

no. 765, 525

De Selm, A. W., no. 140, 342

nos. 275, 276, 394, 599

nos. 499, 621, 412

Dewart, F. W., St. Cloud, Sept., 1892,

599

Dick, W. M., Lorain Co., July 31, 1895,

412

Dickey, S. S., no. 96, 412

Dillon, Quartin, Abyssinia, 447

no. 612, 446

Dixon, R. A., no. 17, 418

no. 528, 342

Dixon, R. A., & Cornelia Gage, no. 713,

342

Dodge, C. K., near Port Huron, Aug.

4, 1895, 412

Douglas, David, N. America, 429

California, 312

California, 515

California, 1833, 512

Ft. Vancouver, 429

Douglas, A. E., Arequipa, 1892, 340

Drake, D., & J. H. Dickson, Bradfords

Isl., June, 1889, 429

Dreisbach, R. R., no. 1042, 41 8

Drummond, Thomas, Texas, 41 8

no. 68, 426

no. 69, 421

no. 199, 559

no. 200, 421

Dudley, W. R., no. 340, 509

Duesner, C. W., Miller, 1908, 412

Pine, 1908, 342

Duges, Alfredo, no. 472, 500

Dummer, R. A., no. 3304, 361

Dunkle, M. B., no. 1857, 508

Dunlap, no. 204, 574

Dunn, G. W., Coronado Isls., Apr. 17,

1891, 306

Eames, E. H., no. 8825, 551

no. 8839, 421

Earle, F. S., Auburn, Sept. 4, 1899, 412

nos. 2060, 2061, 406

no. 2147, 454

Earle, F. S. & E. S., no. 73, v555

Earle, F. S., & C. F. Baker, Auburn,

Apr. 4, 1897, 557

Auburn, Aug. 17, 1897, 406

Auburn, May 28, 1898, 407

no. 243, 542

Eastwood, Alice, Kaweah, Apr. 27,

1895, 509

Cuyama, May 2, 1896, 512

Point Sal, May 12, 1896, 308

no. 2565, 506

no. 2574, 515

no. 3187, 512

nos. 3194, 4377, 515

Eaton, A. A., no. 469, 455

Eggert, Heinrich (Henry), St. Louis,

Aug. 25, 1874, 412

St. Louis, June 29, 1875, 599

Jefferson Co., July 8, 1879, 549

St. Louis, June 18, 1886, 599

Jefferson Co., May 20, 1887, 542

St. Louis, Aug. 23, 1887, 412

Jefferson Co., June 10, 1891, 544

Williamsonville, June 11, 1893, 555

Hematite, May 25, 1896, 544

Jefferson Co., May 25, 1896, 542

Sherwood, June 8, 1897, 406

Stone Mt. and vicin., July 23, 1897,

454

Yellow R., July 27, 1897, 409

no. 122, 412

Eggert, H., & Sydow, St. Louis, July 1,

1874, 599

Eggleston, W. W., no. 19577, 308

no. 19748, 515

Ehlers, J. H., no. 518, 542

Ehrenberg, K. A., no. 354 pro parte, 295

no. 354 pro parte, 299

INDEX OF COLLECTORS

465

Ellenbeck, no. 529, 368

no. 1436, S72

no. 1660, 447

Elmer, A. D. E., no. 601, 429

no. 3634, 308

Emig, W. EL, no. 146, 399

Engelmann, George, Mountain Creek

bottom, 1833, 349

vicin. East St. Louis, May, 1845, 342

St. Louis, Sept., 1850, from Fendler's

seeds, 418

St. Louis, Aug., 1863, 349

Norristown, Aug. 27, 1876, 407

herb, of, Aiken, Apr., 1882, 344

Engelmann, Henry, Ft. Laramie, July,

1858, 418

Engler, Adolph, no. 1266, 377

Evermann, B. W., L. Pend d'Oreille,

Aug. 7, 1893, 418

Fauntleroy, Juliet, no. 654, 407

Fendler, August, no. 397 pro parte, 418

no. 397 pro parte, 426

no. 441, 426

Ferguson, A. M., Tampa, 1898, 432

Fernald, M. L., E. B. Bartram, & B.

Long, nos. 24687, 24688, 424

Fernald, M. L., C. H. Bissell, C. B.

Graves, B. Long, & D. H. Linder,

no. 22859, 424

Fernald, M. L., & B. Long, nos. 10664,

10669, 17591, 17594, 424

no. 19265, 351

nos. 24689, 24690, 24691, 24692, 424

Fernald, M. L., B. Long, & D. H.

Linder, nos. 22860, 22861, 22862,

424

Ferris, Roxana S., no. 7466, 313

no. 7572, 308

no. 7710, 313

Feudge, J. B., no. 1382, 312

Fiebrig, Karl, no. 3476, 339

Fiori, Adriano, no. 1826, 447

Fish, Fanny E., All Saints' Bay, May,

1882, 306

Fisher, G. L., no. 5007, 422

Fitzpatrick, T. J. & M. F. L., ex herb.,

Allamakee Co., June 27, 1895, 399

Flamand, F., Kiamohampa, Sept. 20,

1911 387

Fleming, 'G. L., & W. W. Eggleston,

no. 19676, 306

Fletcher, E. F., Westford, 418

Ford, H. C., Santa Barbara Co., 1889,

310

Francis, Mary E., no. 155, 437

Fredholm, Adolph, nos. 5473, 5778,

6094, 431

Frost, W. D., Willmar, July, 1892, 399

Galen, James, no. 4289, 412

Galeotti, H. G., no. 2086, 298

Garber, A. P., keys of Florida, 1876,

431

Tampa Bay, 1876, 431

L. Monroe, March, 1876, 431

Mellon ville, March, 1876, 431

Tampa, May, 1876, 431

Rosewood, June, 1876, 431

Levy Co., Nov., 1877, 437

Garvens, Wolfgang, vicin. Palm Beach,

1908, 431

Gates, F. C., no. 508, 342

no. 598, 399

no. 808, 412

no. 942, 344

no. 2478, 342

no. 2817, 344

no. 3148, 399

no. 10104, 412

Gates, F. C. & M. T., no. 10547, 342

Geyer, C. A., Beardstown, 344

Ghiesbreght, A. B., nos. 133, 539, 298

Gibbs, Mrs. E. T., South Pond, 424

Gillet, J., nos. 806, 885, 387

Gironcourt, Mission of, no. 256, 372

Gleason, H. A., east of Rantoul,

Oct. 5, 1907, 412

Shirland, June 29, 1908, 399

Milroy, July 6, 1908, 399

no. 326, 399

nos. 812, 1813, 1977, 412

nos. 2749, 2750, 2753, 412

Goetze, W., no. 731, 382

no. 790, 369

no. 1041, 377

no. 1069, 378

Grabham, G. W., Bahadur Gorges, 371

Graebner, P., cult. Hort. Berol., Aug. 5,

1910, 350

Grant, G. B., nos. 681, 682, 313

no. 5424, 310

Grant & Walter Wheeler, nos. 680,

2228, 308

Grassly, C. W., Hyde Park, June 10,

1877, 344

Miller, Aug. 31, 1880, 412

Gray, Asa, Hickory Nut Gap, 1843, 416

Apalachicola, Apr., 1875, 443

Gray, A., & John Carey, mts. of

Virginia and N. Carolina, July,

1841, 407

Gray, A., & W. S. Sullivant, mts. of

Carolina, 1843, 416

Greenman, J. M., no. 437, 424

no. 544, 412

nos. 2017, 2592, 342

no. 3771, 350

no. 3991, 412

Gregg, Josiah, no. 901, 426

Griffiths, David, Highmore, July, 1897,

429

Grimes, E. J., no. 3768, 402

466 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Gronovius, D., ex herb., 402

Groth, B. H. A., no. 66, 422

Guenther, Ernesto, & O. Buchtien, no.

55, 319

Hale, Josiah, Louisiana, 3^5

Louisiana, 412

Louisiana, 4-18

Louisiana, 426

Louisiana, 1842, 342

Alexandria, S57

Hall, Elihu, Oregon, 429

central Illinois, 350

Athens, 1861, 342

Athens, 1861, 399

Athens, 1861, 412

cult, from Texas seeds, 1864, 359

Athens, Aug., 1864-1867, 412

Athens, Aug., 1868, 412

Hempstead, June, 1872, 421

cult, from Dakota seeds, 1877, 429

no. 342, 3 5 It

no. 343, ^21

no. 344 pro parte, 418

no. 344 pro parte, 426

no. 345, 418

Hall, E. E., Texas, 1877, 353

Hall, H. M., no. 3074, 310

no. 3973, 313

Hall, James, Virginia, 1828-1834, 402

Montgomery Co., 1828-1834, 407

Haller, W. H., no. 982, 353

Hamerlinck, P., nos. 1502, 1512, 387

Hanson, H. C., no. A13, 426

Harbison, T. G., Waynesville, 1897, 407

Highlands, May, 1902, 357

Highlands, 1905, 407

Highlands, Aug. 1, 1905, 403

Highlands, Aug., 1906, 403

Harford, W. G. W., & G. W. Dunn,

Cascades, May 28, 1869, 429

Harper, R. A., no. 74, 357

Harper, R. M., vicin. Sawdust, June 18,

1927, 409

no. 5, 402

no. 22, 342

no. 46, 407

no. 131, 407

no. 545, 405

no. 653, 441

no. 1001, 443

no. 1666 pro parte, 439

no. 1666 pro parte, 441

no. 1954, 412

no. 2198, 443

Hartweg, Theodor, no. 116, 321

Harvard Univ. Bot. Gard. (reading

Hort. Bot. Harv. Univ. or Hort.

Cantabr. on some labels), 1842, 404

cult., 1842, 407

cult., 1844, 421

ex Cambridge, Aug., 1845, 406

cult., 1848, from Lindheimer's Texas

seeds, 359

cult., 1849, 409

cult, from Carolina seeds, 1849, 416

cult, from Texas seeds, 1849, 426

cult, from Fremont's seeds, 1851,

1852, 306

cult, from Wright's seeds, 1852, 422

cult., Sept., 1856, 359

cult., 1871, 312

cult., 1873, 407

cult., 1873, 1882, 316

cult., 1882, 313

Harvey, F. L., Little Rock, 418

no. 22, 355

Hasse, H. E., Mojave Station, 312

Desoto, July, 1887, 350

Los Angeles, Apr. 22, 1888, 313

Hayes, Sutton, near Amatitlan, July 20,

1860, 298

Heddle, J. R., no. 698, 399

Heiland, Lychen, Aug. 16, 1882, 418

Heller, A. A., vicin. Salisbury, Aug. 21,

1890, 412

near mouth of Tucquan Creek, Sept. 1,

1890, 412

Coeur d'Alene River, July 12, 1892,

429

about L. Pend d'Oreille, Aug. 1-10,

1892, 429

no. 99, 407

no. 933, 402

no. 1013, 407

no. 1548, 428

no. 1730, 422

no. 7662, 310

no. 7674, 313

no. 7723, 312

no. 11752, 418

Heller, A. A., & E. G. Halbach, near

McCall's Ferry, Sept. 9, 1893, 413

Heller, A. A. & E. G., no. 4129, 413

Heller, Edmund, Mt. Garguez, Aug. 26,

1911, 363

Hens, Fr., no. 45, 387

Herrera, F. L., nos. Ill, 391, 1186, 1481,

S£0

Hexamer, A. C., & F. W. Maier,

Summerville, May 27, 1855, 404

Abbeville Co., June, 1855, 406

ex herb., Soluda R., June 14, 1855,

357

rich woods, June 16, 1855, 342

Heyde, E. T., & E. Lux, no. 3792, 298

Hildebrandt, J. M., no. 417, 446

Hill, E. J., Morgan Park, June 17, 1876,

O t 0

nos. 1108, 2746, 345

Hinkley, Mr. & Mrs. F. E., no. 51, 3^0

Hitchcock, A. S., Jacksonville, 443

Iowa City, 1888, -413

New River, 1896, 431

INDEX OF COLLECTORS

467

Palm Beach, 1896, 1*31

Levy Co., 1898, 431

Madison Co., June-July, 1898, 31*3

no. 139, 433

no. 279, 1*18

no. 666, M8

no. 910, 439

no. 13943, 1>18

no. 21564, 829

Hitchcock, C. L., no. 12224, 313

no. 12340, 310

Hoffman, W., Corn Creek, June 21,

1870, 899

vicin. Rolla, July 15, 1871, 350

Holm, Theodor, Brookland, June, 1894,

402

Holway, E. W. D., Decorah, July 16,

1888, 399

nos. 5, 52, 154, 613, 298

Holzinger (Holtzinger), J. M., Lamoille,

June, 1895, 899

Santa Fe, Aug. 13, 1911, WO

Hombl<§, no. 329, 397

Hooker, J. D., Arkansas, 1+18

Kentucky, 407

Hort. Cantabr., see Harvard Univ. Bot.

Gard. (for Hort. Berol., etc. see

Berlin, etc.)

Houghton, Douglass, 200 miles above

St. Anthony's Falls, 399

above St. Anthony's Falls, July, 1832,

399

Houghton, H. W., no. 3567, 853

no. 3864, 350

House, H. D., no. 150, 402

no. 4023, 350

no. 9660, ^It

Howell, A. H., no. 808, 407

Howell, J. T., no. 8220, 308

Howell, T. J., Oregon, 1881, 429

Klickitat, June, 1881, 429

banks of Columbia, July, 1881, 429

no. 224, 1*29

Hrdlicka, vicin. San Damian, Jan.-

Feb., 1913, 340

Huett, J. W., Ottawa, 1*13

Hulier, Hort., no. 8, 300

Humboldt, Alexander von, & Aime"

Bonpland, near Loja, 329

Paramo de Saraguru, 329

Hunnewell, F. W., 2nd, White Sulphur

Springs, July 4-6, 1914, 407

Hunt, Edwin, Sudbury, Aug. 12, 1874,

4*4

Hutchinson, Mary E., Randolph, 343

Hyams, M. E., Statesville, July, 1880,

357

Jackson, F. J., Kikuyu region, 1889, 361*

Jaeger, E. C., no. 140, 313

Jameson, William, Ecuador, 335

hills near Loja, commun. 1865, 334

Jermy, Gustav, no. 657, 423

Johnson, L. N., Chicago, June 19, 1891,

343

Johnston, I. M., near Mojave, May 5,

1920, 312

near Mojave, May 5, 1920, 318

Victorville, May 15, 1920, 313

5 miles s.e. of Victorville, May 17,

1920, 311*

1 mile s. of Crutts P.O., May 14, 1922,

312

no. 1251, 314

Jones, M. E., Pasadena, Feb. 22, 1882,

314

Fallbrook, March 25, 1882, 314

San Diego, Apr. 3, 1882, 314

Todos Santos Bay, Apr. 11, 1882, 306

Darwin, Apr. 28, 1897, 310

Shepherd's Canyon, Apr. 30, 1897,

310

Pleasant Canyon, May 6, 1897, 310

Whitewater, May 11, 1903, 314

Cajon Pass, May 16, 1903, 310

Victor, May 17, 1903, 314

Mojave, May 20, 1903, 312

Marsh Lake, Sept. 17, 1903, 435

San Diego, Apr. 14, 1906, 314

Julian, Apr. 17, 1906, 314

Sand Point, Sept. 18, 1908, 1*29

Cave Spring, March 12, 1924, 310

Santa Cruz Isl., Apr. 13, 1924, 308

ranch 29 miles s.w. of Tia Juana,

Apr. 13, 1925, 306

near Santa Ana Canyon, March 10,

1926, 314

Malibu Hills, Apr. 26, 1926, 303

Cuyama Canyon, Apr. 28, 1926, 312

Coronado Isls., June 10, 1926, 306

Adelanto, May 12, 1927, 312

Santa Cruz Isl., March 25, 1929, 308

no. 3134, 306

nos. 3361 pro parte, 3373, 314

nos. 28012, 28013, 423

no. 29474, 418

nos. 29511, 29512, 359

Jones, P. M., Santa Rosa Isl., Apr.,

1901, 308

Jones, W. W., no. 160, 426

no. 227, 1*18

Joor, J. F., Galveston Bay, June, 421

Kalbreyer, W., no. 125, 373

Kassner, T., no. 2776, 398

no. 2871, 383

Katzenstein, Otto, herb., s.e. U.S., Sept.

23, 1897, 437

Kearney, T. H., Jr., Clear Creek, Sept.,

1893, 415

White Cliff Springs, July 14, 1894,

407

no. 553, 413

Kellerman, W. A., no. 6296, 298

468 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Kellogg, A., & W. G. W. Harford, no.

436, 312

no. 439, 816

Kellogg, J. H., Allenton, Aug. 12-13,

1884, 850

St. Louis, July 15, 1903, 350

cult., June 30, 1905, 421

cult., Mo. Bot. Gard., May 23, 1911,

554

Kellogg, William, Rancho Encinal,

April 15, 1903, 31 1*

Owanita, 1907, 451

Kennedy, G. G., Orleans, July 31, 1891,

4#4

Ormond, March 29, 1897, 545

Sharon, Sept. 2, 1904, 424

Kennedy, W. L., Fort Tejon, spring,

1876, 310

Kenoyer, L. A., & F. W. Pennell, no.

3224, 424

Kersting, nos. 14, 422, 395

Kew., Hort., cult, from R. Diespecker's

seeds, 584

Killip, E. P., & A. C. Smith, no. 21536,

31*0

no. 21823, 325

no. 22171, 555

no. 23304, 540

King, Anna, no. 116, 405

nos. 534, 566, MS

Kling, nos. 60, 166, 597

Knopf, E. C., nos. 56, 258, 508

Krebs, William, Cleveland, 415

Kreger, F. 0., no. 315, 1*29

Lang, H. A., Egg Harbor, Sept. 10,

1905, 425

Langlois, A. B., western Louisiana,

Apr., 1880, 545

Spring Hill, Oct. 1, 1880, 436

Lansing, O. E., Jr., no. 243, 545

no. 307, 599

no. 647, 1*13

nos. 848, 935, 545

no. 1646, MS

nos. 2715, 2782, 5*5

no. 2797, WO

no. 2832, 415

no. 3113, 400

nos. 3822, 3911, MS

Lapham, I. A., Milwaukee, 545

Milwaukee, 400

Laurent, Em., between Lusambo and

Lomani, 587

Laurent, Em. & Marc., Kondue, Nov.

25, 1903, 587

Laurent, Marc., no. 512, 587

Leavenworth, M. C., Louisiana, 555

Louisiana, 1*00

Tampa Bay, 1*31

Lebrun, Jean, no. 4507, 1*1*1*

L4card, no. 290, 595

Le Conte, North America, 55.4

Ledermann, C., no. 1855, 372

no. 5243, 595

Leeds, B. F., Chattanooga, Sept. 29,

1892, 1*06

Chattanooga, Sept. 29, 1892, 1*07

Lehmann, ex, 1*00

Leiberg, J. B., Kootenai Co., June, 1890,

1*18

L. Pend d'Oreille, July, 1892, 1*19

no. 1542, 1*29

Lely, H. V., nos. 383, 449, 568

Lemmon, J. G., top of Mt. Diablo, 310

San Bernardino, March, 1880, 31 J*

no. 137?, California, 312

no. 138, 510

no. 4159, 1*29

no. 4520, 312

Leonard, E. C., no. 4103, 505

Lescrauwaet, E., no. 188, 587

Letterman, G. W., Allenton, 550

near Franklin Co. Road Bridge, July,

1883, 5-45

Allenton, Aug., 1883, 545

Merely, July, 1884, 5-45

Gratiot, May 30, 1897, 351*

near St. Louis, Aug., 1899, 554

Levine, C. 0., no. 778, 5-45

Lewton, F. L., Forest City, 1894, 1*33

Cedar Hammock, Aug. 3, 1894, 455

Lighthipe, L. H., Monticello, Apr. 5,

1891, 545

no. 557, 457

Linder, D. H., no. 1706, 588

Lindheimer, F., w. of Brazos River,

Apr., 559

w. of Houston, early May, 1840, 545

Houston, Apr., 1842, 554

Galveston Id., Nov., 1842, 1*21

Texas, 1849, 1*21

no. 54, 1*21

no. 101, 1*22

no. 102 pro parte, 1*19

no. 102 pro parte, 1*26

no. 363, 559

no. 548, 419 ;

no. 897, 419

Lips., Hort., ex Amer. septentr., 550

ex Illinois, 599

Lloyd, C. G., near Rockwood, Aug. 10,

1880, 409

Lloyd, F. E., & S. M. Tracy, no. 527a,

54ff

Lloyd, R. N., Ravenswood, June 3,

1887, 545

Ravenswood, July 11, 1887, 400

Ravenswood, Aug. 31, 1887, 415

Loring, Lake Superior, 545

Luckhardt, no. 626, 575

Lyall, David, Cascade Mts. to Ft.

Colville, 1860, 429

Colville to Rocky Mts., 1861, 429

INDEX OF COLLECTORS

469

Macbride, J. FM no. 2932, 337

nos. 3152, 3504, 321*

Macbride, J. F., & Featherstone, no.

210 337

nos. 436, 690, 1237, 340

MacCarthy, Gerald, District of Co-

lumbia, 402

eastern N. Carolina, 1883, 439

MacDougal, D. T., no. 293, 426

no. 617, 429

MacKay, Ernest, no. 25, 430

Mackensen, Bernard, no. 76, 422

Mackenzie, K. K., no. 100, 351*

no. 328, 350

Macoun, J. M., Waneta, July 28, 1902,

430

Macoun, John, Sandwich, Aug. 4, 1892,

US

Winnifred, Aug. 12, 1895, 419

no. 38, 419

Magee, E. E., Hayward Co., Sept.,

1897, 407

Waynesville, Sept. 9, 1897, 407

below Satulah Mt., Aug. 2, 1902, 407

Highlands, Sept. 2, 1902, 407

Mann, Gustav, no. 1922, 373

Manning, W. H., Lake City, Aug. 13,

1883, 400

Mackinac Isl., July 11, 1915, 343

Manz, no. 29, 422

Mathews, Alexander, Prov. Chacha-

poyas, 1835, 1836, 328

no. 571, 340

no. 1376, 331

no. 1418, 323

McClatchie, A. J., San Diego, Apr.,

1893, 306

McDonald, F. E., Peoria, June, 1891, £00

Peoria, June, 1899, 343

Peoria, Aug., 1900, 413

Peoria, 1900-1901, £00

Peoria, Sept., 1901, 413

Peoria, June, 1904, 343

Peoria, June, 1904, 400

Peoria, Aug., 1904, 413

Peoria, May, 1915, 345

Peoria, June, 1915, 400

Peoria, June, 1916, 343

Peoria, July, 1919, 419

McLean, Andes, Peru, 336

Mead, S. B., Mason Co., June 18,

1845, 343

Illinois, 1850, 400

Mearns, E. A., no. 144, 400

nos. 1291, 1722, 364

Mendez, 300

Mexia, Ynes, no. 445, 358

Meyer, Hans, no. 532, 444

Michaux, Andr6, North America, 350

North America, 404

North America, 413

North America, 439

Migeod, F. W. H., no. 211, 374

Mildbraed, F., no. 2539, 444

no. 3392, 373

Miller, E. S., Long Pond, Aug. 30, 1871,

425

Wading River, Aug. 18, 1877, 425

Milligan, Mrs. J. M., Jefferson Co., 345

Millspaugh, C. F., no. 3865, 413

no. 4630, 308

Mintz, no. 40, 373

Moffatt, W. S., no. 105, 343

no. 492, 413

Mohr, Charles, Mentone, 403

Mobile, 406

ex herb., eastern shore Mobile Bay,

Apr., 1868, 422

Gulfport, July 4, 1874, 346

Bladon, July, 1875, 403

Lawrence Co., May, 1880, 357

ex herb., Winston Co., May 1, 1881,

357

Cullman, June, 1883, 346

vicin. Ashland, June, 1883, 346

Mulberry R. Valley, June 6, 1883, 346

cult, from Texas seeds, Mobile,

May 10, 1887, 360

Columbus, June 4, 1888, 346

Mentone, Sept. 18, 1892, 403

Gallion, May 25, 1893, 347

Poplarville, Oct. 18, 1894, 435

Yellow Pine, Aug. 8, 1895, 441

DeSoto Falls, Sept. 12, 1898, 403

Moore, A. H., no. 1832, 419

nos. 2552, 2553, 413

Morgan, R. T., vicin. Fountaindale,

Aug., 1877, 400

Morong, Thomas, ex herb., Winchester,

Aug. 5, 1881, 425

Morris, E. L., no. 105, 402

no. 1050, 407

Mosely, E. L., Oxford, Aug. 21, 1895,

413

Muenzner, no. 159, 382

Munroe, H. F., Hyde Park, May, 1879,

3L.3

Hyde Park, June, 1879, 343

Munz, P. A., no. 1589, 406

nos. 2148, 2538, 3814, 314

no. 4466, 310

no. 7955, 306

nos. 9136, 9186, 310

no. 9357, 314

no. 10042, 310

no. 11378, 308

no. 12415, 314

Munz, P. A., & E. Crow, nos. 11553,

11691, 11750, 308

Munz, P. A., & R. D. Harwood, nos.

3385, 3475, 314

Munz, P. A., R. D. Harwood, & I. M.

Johnston, no. 4077, 310

470 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XI

Munz, P. A., & D. D. Keck, nos. 7842,

7879, 310

no. 7930, 311,.

Munz, P. A., L. Street, & G. Williams,

nos. 2362, 2388, Silt

Munz, P. A., & J. Voss, no. 11877, 308

Murdoch, John, Jr., Orleans, Aug. 22,

1900, 425

Murdoch, Joseph, Rochester, Aug. 24,

1902, 425

Nash, G. V., nos. 832, 1257, 1876, 431

no. 2255, 433

no. 2585, 415

Nealley, G. C., Brazos Santiago, 1889,

428

Chenates, 1889, 419

Negri, Giovanni, no. 56, 447

nos. 2356is, 618, 448

no. 9156is, 367

Nelis, Bokala, May, 1913, 387

Nelson, E. W., no. 1718, 299

nos. 2185, 2249, 318

nos. 6104, 6813, 426

Nelson, J. C., no. 1802, 419

no. 1980, 430

Ness, H., no. 2078, 360

Neumann, Oscar, no. 75, 448

no. 134, 379

Newberry, Sitgreaves Pass, March 26,

314

Nicolas, Puebla, Oct., 1905, 299

Norton, J. B., e. of Hartsville, Nov. 5,

1921, 441

no. 1455, 419

Nutt, W. H., between L. Tanganyika

and L. Rukwa, 1896, 382

Nuttall, L. W., no. 571, 308

Nuttall, Thomas, without data, 314

Red River, 419

Cakes, Plymouth, 425

Olive, E. W., no. 72, 426

Orcutt, C. R., Point Loma, March 12,

1884, 314

Point Loma, March 24, 1884, 306

northern Baja California, Apr. 17,

1886, 81k

Orcutt, C. R., & W. S. Bowne, San

Diego, 1884, 306

Oyster, J. H., no. 4014, 426

Palmer, Edward, no. 41, 308

no. 126, 314

no. 172, 360

no. 217, 314

no. 286, 431

nos. 489, 530,317

no. 568, SOS

nos. 635, 655, 426

no. 677, 314

Palmer, E. J., Newton Co., July 15,

1906, 419

no. 17, 343

no. 2272, 419

nos. 2317, 2317A, 350

nos. 2386, 2386A, 343

no. 2513, 350

no. 6962A, 352

no. 7740, 422

no. 8120, 355

no. 8468, 415

no. 9144, 300

no. 12115, 423

no. 18078, £00

no. 27056, 345

no. 29347, 350

no. 29421, 415

Pammel, L. H., no. 621, 413

Pappi, A., nos. 1194, 1303, 2642, 448

no. 7733, 395

Paris., Hort., 1816, 350

1820, 404

1823, 418

Paris, Hort. Bot. de 1'Ecole de M<§d. de,

Aug., 1820, 413

Parish, S. B., Marengo, Apr., 1882, 310

nos. 2211, 5191, 314

no. 9771, 312

no. 10415, 314

Parish, S. B. & W. F., San Bernardino,

Apr., 1881, 314

foothills of San Bernardino Mts.,

May, 1888, 314

no. 226, 310

Parker, C. F., Bristol, Aug. 11, 1865,

425

Parlin, J. C., North Berwick, July 4,

1892, 419

Parry, C. C., Pacific coast of N.

Amer., 1880, 310

California, 312

San Bernardino, 314

San Diego, 308

San Diego, 1850, 305

San Diego, 1883, 306

no. 115, 310

no. 116, 314

Parry, C. C., & J. G. Lemmon, no. 184,

310

no. 185, 314

Patterson, H. N., Henderson Co., Aug.,

1871, 413

near Oquawka, July, 400

near Oquawka, June 20, 1872, 400

Pearce, R., Cordillera Huanta, Feb.,

1867, 338

Pease, A. S., s. of Champaign, Oct. 9,

1920, 400

no. 12394, 400

no. 17993, 343

INDEX OF COLLECTORS

471

Peattie, D. C., no. 8, 400

nos. 164, 2029, 413

Peirson, F. W., no. 3900, 430

Peng, To Kang, Ts'ang Wai Tak, &

Ts'ang Un Kin, no. 936, 419

Pennell, F. W., no. 4029, 434

no. 13244, 340

no. 14342, 537

nps. 14396, 14710, 340

Petit, A., Abyssinia, 446

Abyssinia, 448

Pfund, T., Dar-Fur, 1876, 395

no. 336, 395

Piemeisel, R. L., & L. W. Kephart, no.

166, 366

Pierce, W. M., 20 mi. e. of Victorville,

Apr. 12, 1922, 314

desert 20 mi. e. of Victorville, Apr.

12, 1922, 31 2

Pitcher, Zina, Arkansas, 354

Arkansas, 419

Plowden, Abyssinia, 371

Plummer, Miss, mts. above Santa

Barbara, 1878, 308

Pogge, P., nos. 257, 1282, 387

no. 1292, 389

nos. 1295, 1309, 387

Pollard, C. L., no. 1222, 414

Pollard, C. L., & W. R. Maxon, no. 496,

347

Poole, S. F., Sharon, July, 1905, 343

Porter, T. C., middle Georgia, 1846, 407

near Augusta, Apr., 1847, 343

Susquehanna R., Aug. 22, 1861, 413

40 mi. s. of Lynchburg, July 17, 1880,

402

9 mi. above Kanawha Falls, July 21,

1880, 350

Powell, B. T., Hastings, July, 1886, 419

Preuss, Kamerun, 1891, 373

no. 689, 373

Pringle, C. G., banks of Columbia R.,

Sept. 10, 1881, 430

bluffs of seashore, San Diego, Apr.

25, 1882, 306

hills bordering Mojave Desert, May

11, 1882, 310

Mojave Desert, May 14, 1882, 312

nos. 2197, 2303, 317

no. 2367, 361

nos. 3570, 3841, SOS

no. 4871, 304

no. 4896, 299

no. 5508, 318

no. 7866, 320

no. 8218, 298

no. 8328, 426

nos. 8332, 8431, 320

no. 8781, 301

no. 9885, 361

no. 9886, 320

no. 9895, 300

no. 10050, 320

no. 11506, 361

no. 11507, 317

no. 11546, 303

no. 11900, 301

no. 13041, 298

no. 13547, 300

Purpus, C. A., no. 1339, 300

no. 1550, 298

nos. 2581, 2581a, 302

no. 3099, 300

no. 4098, 304

no. 5004, 316

nos. 5063, 5531, 310

Quarre", P., no. 419 pro parte, 386

no. 419 pro parte, 390

no. 1616, 391

no. 1662, 394

nos. 2173, 2826, 2921, 389

Ragazzi, V., Let-Marefia, Sept. 4, 1886,

448

Let-Marefia, Sept. 6, 1886, 448

Antoto, Oct. 2, 1887, 448

no. 250, 448

Raimondi, A., no. 436, 330

no. 1774, 323

no. 3022, 335

no. 6463, 329

nos. 6695, 7732, 7914, 333

no. 12060, 329

Randolph, Fannie R., no. 136, 431

Randolph, L. F. & F. R., no. 150, 402

no. 1066, 407

Ravenel, H. W., Santee Canal, 345

damp pine land, Santee Canal, Sept.,

441

damp rich soils, Santee Canal, Sept.,

442

Santee Canal, July, 1846, 409

Redfield, J. H., no. 5630, 439

Reed, F. M., no. 6147, 306

Reverchon, Julian, White Creek, July 4,

422

Dallas, May, 1874, 354

Dallas, May, 1876, 354

western Texas, 1882, 423

Pine Isl., May 5, 1903, 348

nos. 513, 1478, 354

nos. 2041, 2077, 348

Reynolds, Carrie A., Stony Is!., Aug. 24,

1909, 413

Reynolds, H. S., Judsonia, May 31,

1877, 355

Riddell, J. L., Kentucky, 407

Ohio, 413

Ridgway, Robert, no. 60, 413

Riehl, Nicholas, no. 41, 400

no. 65, 350

472 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Robinson, B. L., Normal, July, 1887,

1>00

Hendrix, Aug. 31, 1904, 413

no. 56, 413

no. 79, 407

Robyns, Walter, no. 138, 389

Rohlfs & Stacker, Katz, Dec. 30, 1880,

448

Godofelassi, Jan. 7, 1881, 370

Rolfs, P. H., no. 309, 343

no. 392, 357

Rose, J. N., no. 2344, 301

Rose, J. N., A. Pachano, & G. Rose,

no. 23146, 334.

Rose, J. N., & J. H. Painter, no. 7480,

303

Rose, J. N., J. H. Painter, & J. S. Rose,

no. 8837, 298

no. 9717, 300

Rosendahl, C. O., no. 656, WO

Rothrock, J. T., cult., Philadelphia,

June 25, 1877, 419

no. 157, 426

no. 317, 419

Rowland, western Lagos, 1893, 355

Rugel, Ferdinand, Broad R., June-July,

1841, 350

Broad R., July, 1841, 416

valley of Blue Ridge Mts., Aug., 1841,

41.4

Florida, 1842-1849, 439

between Quincy and Aspalaga, May,

1843, 350

Florida, 1844, 407

no. 102, 432

no. 133, 433

no. 142, 432

no. 480, 437

no. 570 pro parte, 441

Runyon, Robert, no. 335, 428

Rtippell, Eduard, between Temben and

Siemen, 1832, 448

Rusby, H. H., no. 1685, 337

Russell, Margaret P., Asheville, 407

Hood R., Aug. 15, 430

herb, of, Rochester, Aug. 17, 1918,

425

Ruth, Albert, Chilhowee Gap, July,

1893, 409

Mount Nebo, July, 1893, 408

Knoxville, June, 1893, 407

Knoxville. May, 1895, 357

Knoxville, June, 1895, 408

Knoxville, May, 1896, 409

Lookout Mt., July 20, 1897, 408

no. 67, 357

no. 69, 414

no. 482, 354

nos. 612, 624, 403

no. 652, 357

nos. 655, 665, 350

Rydberg, P. A., no. 9445, 35.Z

Salvin, Osbert, Volcan del Fuego,

Aug., 1873, 298

Sandberg, J. H., Hennepin Co., Aug.,

1889, 400

Hennepin Co., July, 1890, 400

Sandberg, J. H., D. T. MacDougal,

& A. A. Heller, no. 617, 430

Sapin, A., N'Dolo, May, 1906, 387

N'Kolokosso, June 28, 1906, 387

Sokomini L., Sept., 1906, 387

Katola Msek, Apr., 1908, 388

Munungu, Apr., 1910, 388

Sargent, C. S., no. 27, 340

Sartwell, H. P., Maumee R., 4*4

Saurman, B. F., Apalachicola, 343

Apalachicola, 443

Savage, T. E., J. E. Cameron, & F. E.

Lenocker, Spokane, July, 1898, 419

The Dalles, Aug., 1898, 419

Savage & Stull, no. 60, 400

Scammon, Franklin, Chicago, 345

Chicago, June, 1859, 343

Chicago, June 3, 1859, 343

Schimper, Wilhelm, no. 48, 448

no. 62, 446

nos. 71, 85, 448

no. 429, 446

no. 933 pro parte, 450

no. 1173, 371

no. 1236, 448

no. 1386, 370

no. 1423, 448

sect. 5, no. 15, 450

sect. 5, nos. 22, 95, 448

sect. 5, no. 933, 450

sect. 5, no. 1005, 448

Schindler, A. K., no. 335, 419

no. 345a, 343

Schlechter, Rudolph, no. 12474, 388

Schneck, Jacob, LaGrange, May, 1894,

422

Schott, Arthur, near Georgetown, Aug.,

1857, 402

near Georgetown, June 22, 1860, 402

Schroeder, F., no. 74, 395

Schuette, J. H., Door, July 30, 1887, 343

St. Anthony Park, July 2, 14, 1888,

400

St. Anthony Park, July 8, 1888, 400

St. Anthony Park, July 16, 1888, 400

Plover, July 26, 1888, 400

cult., Green Bay, Sept. 14, 1888, 419

cult., Green Bay, Sept. 14, 1888, 422

n. of Sturgeon Bay Canal, July 14,

1890, 343

cult., Green Bay, Aug. 23, 1897, 419

Green Bay, Aug. 13, 1899, 400

Green Lake, Aug. 13, 1899, 400

cult., Green Bay, July 31, 1901, 419

cult., Green Bay, Aug. 2, 1901, 419

no. 76, 354

Schulz, Ellen D., no. 367, 423

INDEX OF COLLECTORS

473

Schweinfurth, G. (Fl. Gallabat 439),

Matamma, Oct., 1865, 395

(Fl. Gallabat 585), Matamma, Oct.,

1865, 396

Scott, Hugh, Mt. Zuguala, 1926, 448

Seemann, Berthold, no. 675, 33%.

Seymour, F. C., no. 1384, 1*25

Shaw, Marjories, no. 1941, 310

Shaw, Marjorie, Edna Spaulding, &

Mrs. C. Li. Walton, Antelope

Valley, 310

Sheldon, E. P., Sleepy Eye, July, 1891,

ItOO

Ft. Snelling, June, 1895, 400

Pike Island, July, 1895, 1*19

no. 11081, 1*30

Shepard, E. M., Greene Co., July, 1880,

350

Sherff, E. E., no. 250 pro parte, 419

no. 250 pro parte, 430

no. 253, 402

no. 278, 354

no. 284, 400

no. 498, 422

nos. 499, 507, 419

no. 989, 350

no. 1038, 414

nos. 1642, 1781, 400

nos. 5000, 5000a, 419

no. 5003, 404

no. 5004, 402

no. 5005, 350

no. 5006, 343

no. 5007, 408

nos. 5013, 5014, 345

no. 5015, 422

no. 5016, 345

no. 5017, 360

no. 5018, 422

Sherwood, W. L., Waynesville, July,

1897, 408

Tuckasiegee Valley, Sept. 20, 1897,

408

Shimek, Bohumil, Monte Sano, July,

1891, 408

Oklahoma City, July, 1892, 419

Short, C. W., hilly lands, Kentucky, 408

Lexington, 1835, 357

no. 42, 350

Shriver, Howard, Wytheville, 1875, 409

Wytheville, July, 1877, 408

Simon, Gust., no. 3, 373

Simpson, J. H., Manatee, 1889, 437

Skeels, H. C., no. 304, 343

no. 375, 400

Skehan, Josephine, Ocean Springs,

Sept. 16, 1895, 436

no. 137, 406

Small, J. K., n. of Marysville, Aug. 15,

1888, 414

vicin. mouth of Tucquan Creek,

Sept. 1, 1890, 414

vicin. mouth of Tucquan Creek,

Oct. 11, 1890, 414

Round Top Mt., July 2, 1892, 408

Farmer Mt., July 12, 1892, 408

Nick's Creek, Aug. 5, 1892, 408

Stone Mt., July 3, 1893, 406

Little Stone Mt., July 7, 1893, 354

Little Stone Mt., July 7, 1893, 409

banks of Yellow R., July 11, 20, 1893,

355

between Alcovy R. and No Business

Creek, July 14, 1893, 354

Tallulah Falls, Aug. 4, 1893, 414

Estatoah Falls, Aug. 12, 1893, 41 4

Thomas Bald, Aug. 19, 1893, 350

Little Long Creek, Aug. 22, 1894, 402

Altamaha R. Swamp, June 18-21,

1895, 404

Stone Mt., Aug. 1-6, 1895, 434

base of Currahu Mt., July, 1896, 350

Small, J. K., & J. J. Carter, Bull Key,

Nov. 6, 1903, 432

no. 694, 439

no. 954, 440

no. 1028, 437

no. 1219, 432

no. 2731, 440

no. 3025, 433

no. 3401, 440

Small, J. K., J. B. DeWinkeler, &

C. A. Mosier, no. 11252, 443

Small, J. K., & A. A. Heller, near R.

& D. Railroad, June 3, 1891, 357

n. of Hickory, June 25, 26, 1891, 409

Green's Hill, July 10, 1891, 408

between Cranberry and Linville, July

17, 1891, 408

Rocky Knob, July 24, 1891, 408

Grandfather Mt., July 25, 1891, 408

no. 100, 414

no. 328, 408

no. 372, 402

Small, J. K., & G. V. Nash, Everglades,

1901, 437

Smith, C. L., Sierra de San Felipe,

Oct. 11, 1894, 299

Smith, E. C., no. 510, 400

Smith, H. H., no. 1089, 350

no. 1150, 355

no. 2456, 350

no. 2526, 409

no. 2582, 350

no. 5034, 308

Smith, J. D., near Starke, Apr. 9, 1880,

443

Warm Springs, July 29, 1880, 408

Caesar's Head, July, 1881, 408

Pellicier's Creek, March 4, 1882, 433

Richland Gap, Aug. 14, 1882, 351

Wildcat Ridge, Aug. 21, 1882, 350

Wildcat Ridge, Aug. 21, 1882, 351

Sumter County, March 21, 1883, 432

474 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XI

Cumberland Falls, Aug. 23, 1883, 351

Columbia, Sept., 1883, 44*

banks of Soquee R., Sept. 13-15,

1883, 351

Orangeburg District, Aug. 9, 1884,

406

Orangeburg District, Aug. 9, 1884,

^09

banks of Oconee R., Aug. 16, 1884,

351

near Montgomery, Aug. 26, 1885, 415

west of Cahawba R., Sept. 2, 1885, 351

Biloxi Bayou, Sept. 16, 1885, 436

no. 602, 347

no. 2028, 441

Smith, L. C., nos. 25, 808, 299

Snyder, Mrs. M. S., seashore, La Jolla,

Feb. 25, 1895, 306

La Jolla, March 15, 1895, 31 5

Spalding, Clear Water, 430

Sparano, no. 25, 388

Spencer, Mary F., no. 86, 306

no. 172, 315

no. 407, 310

nos. 563, 653, 653a, 315

no. 1114, 345

nos. 1954, 2040, 310

no. 2123, 315

Standley, Jeanette P., no. 195, 432

no. 423, 437

Standley, P. C., no. 4410, 344

Steele, E. S., Washington, Aug., 1889,

402

Steetz, ex, ex Lexington, 357

Steudner, H., no. 253, 448

no. 349, 446

Stevens, G. W., no. 784, 419

no. 1212, 419

no. 2387, 351

no. 2713, 400

Steward, A. N., no. 1510, 419

nos. 1518, 1583, 1922, 344

Stewart, J. T., northern Mississippi,

1863, 557

near Charleston, 1865, 440

near Charleston, June 1, 1865, 404

around Peoria, 1867, 400

Stillman, valley of upper Sacramento R.,

316

St. John, Harold, no. 2939, 425

Stolz, Ad., no. 764, 382

no. 1306, 377

Straub, F. C., nos. 97, 159, 432

Strauss, H., San Diego and near-by isls.,

Sept. 24, 1908, 306

San Diego and near-by isls., Feb. 3,

1912, 306

Street, L., San Elejo Lagoon, May 12,

1917, 306

Stubel, A., no. 35 pro parte, 330

Suksdorf, W. N., Columbia R., Aug. 26,

1882, 430

Taylor, Thomasyille, Sept. 30, 1903, 441

Taylor, Katherine A., ex herb., Sum-

merville, 440

Columbia, May, 1890, 357

Summerville, June 10, 1891, 404

Terracciano, A., & A. Pappi, no. 2474,

448

Tharp, B. C., no. 5627, 427

no. 5635, 360

Thaxter, Roland, Cullowhee, June-

July, 1887, 351

Cullowhee, June-July, 1887, 408

Thone, Frank, no. 232, 414

Thoreau, D. B., no. 2.11, 384

Thurber, George, Cranston, Aug., 1844,

425

Thurrow, W. F., Hockley, 1890, 344

Hockley, 1890, 419

Hockley, 1890, 422

Hockley, 1890, 427

Tourney, J. W., no. 594, 427

Townsend, C. H. T., A192, 339

no. A211, 326

Townsend & C. M. Barber, no. 150, 427

Tracy, S. M., no. 3456, 427

no. 4778, 436

no. 6920, 432

no. 6921, 432

no. 7142, 437

no. 7256, 433

no. 7355, 433

no. 7704, 432

no. 7902, 420

no. 8542, 3*6

no. 8550, 422

no. 8565, 436

no. 8670, 4H

no. 8925 pro parte, 360

no. 8925 pro parte, 422

no. 9049, 408

no. 9454, 344

Tracy, S. M., & Lloyd, no. 508, 344

Trask, Blanch, no. 76, 305

Trelease, William, Hesperia, Apr. 10,

1892, 310

Troll, C., no. 3157, 340

Tuckermann, Edward, Jr., ex herb.,

New England, 425

Uhde, no. 627, 300

Uline, E. B., Fish L., July, 1891, 400

Mishawaka, July 1, 1891, 400

Umbach, L. M., Miller, June 5, 1897,

344

Clarke, Aug. 28, 1897, 414

Clarke, June 4, 1898, 344

DuPage, July 6, 1898, 400

Clarke, Aug. 20, 1898, 4H

no. 2752, 419

U. S. Expl. Exped. under Capt. Wilkes,

above Obrajillo, 337

INDEX OF COLLECTORS

475

Vanderyst, Hyacinthe, Ndembo, Aug.

20, 1906, 388

Ndembo, Jan. 7, 1907, 388

Inkissi Falls, May, 1907, 388

Leopoldville, May 23, 1907, 388

Kumpako, 1908, 388

Leopoldville, May 27, 1910, 388

N'Boma, Apr. 28, 1911, 388

nos. 5224, 700, 1099, 1187, 1341,

1582, 4304, 388

nos. 4885, 10097, 388

nos. 12366, 21227, 389

Van Hermann, no. 2686, WO

Vasey, G. R., northern Alabama, 1878,

408

N. Carolina, 1878, 357

N. Carolina, 1878, 408

Lookout Mt., 1878, 414

no. 290, 311

no. 551, 430

Verdick, E., Lukafu, 388

Vermoesen, F., no. 2403, 389

Vesterland, Otto, Marion Co., Novem-

ber, 1889, 441

Von Mechow, Alexander, nos. 33, 459,

387

Von Tuerckheim, H., no. 11-2043, 298

Wallace, Los Angeles, 1854, 315

Ward, L. F., Potomac flats, Aug. 12,

1877, 414

Ford Co., Oct. 5, 1897, 420

Warner, S. R., Willis, June 1, 422

Warszewicz, no. 37, 340

Watson, Sereno, no. 217, 430

Webb, R. J., Pippin L., Sept. 6, 1909,

414

Webber, H. J., Sanibel Isl., Jan. 29,

1896, 432

no. 414, 432

Weberbauer, A., nos. 203, 2394 pro

parte, 340

no. 3812, 838

no. 4048, 322

nos. 5764, 5782, 340

no. 6995, 332

Wellman, F. C., Bailundo District, 388

Werdermann, Erich, no. 1114, 336

Werner, W. C , no. 92, 408

Wharf, E. H., N. Carolina, 357

Wheeler, H. E., no. 62, 354

no. 82, 344

White, Mark, no. 218, 420

Whitford, H. N., nos. 64, 203, 344

Whyte, Alexander, northern Nyassa-

land, 382

Mt. Zomba, Dec., 1896, 377

no. 35, 377

Wiegand, K. M., & W. E. Manning,

no. 3410, 438

no. 3414, 408

no. 3415, 408

nos. 3421, 3422, 434

Williams, E. F., Cataumet, Sept. 15,

1901, 425

Boston, Sept. 3, 1903, 414

Wickford, June 17, 1908, 345

Concord, June 20, 1908, 344

Williamson, C. S., Waycross, Apr. 15,

443

Willich, C. L., & F. W. Weiss, ex herb.,

427

Woodson, R. E., Jr., no. 655, 351

Woodson, R. E., Jr., & E. S. Anderson,

no. 1545, 345

Woytkowski, Felix, no. 23, 330

no. 24, 327

Wright, Charles, Texas, 354

Texas, 422

Texas, 423

northern Georgia, 408

no. 217, 423

nos. 339, 341, 420

nos. 340, 1236, 427

no. 1237, 423

Wright, W. G., San Bernardino, 1880,

311

Ventura Co., March, 1894, 308

no. 172, 311

Wright, W. H., no. 50, 344

no. 221, 414

Young, J. D., no. 153, 368

Young, M. S., near Paloduro Canyon,

Sept. 7, 1917, 427

Zech., Graf., no. 168, 396

Zeller, Cyril, Ellis Co., 1908-1911, 420

THE LIBRARY OF THE

OCT281936

UNIVERSITY OF IH INOI?

ATIONS

1 OF NATURAL

-

0, U.S.A.

-1936