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62 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

cm. longa 5.5-9.5 cm. lata, apice obtusissima vel subrotundata,

basi profunde (1.5-3.5 cm.) cordata, sinu aperto lato, lobis basalibus

subrotundatis, toto margine undulato-dentato, dentibus brevibus

latis obtusissimis, supra sparse villosa vel in statu juvenili forsan

laxe tomentosa, subtus ad venas nervosque sparsissime breviter

tomentulosa vel fere glabra penninervia; folia caulina multo minora

ovata, petiolo latissime alato atque vaginante, foliis supremis ad

vaginas reductis 2-2.5 cm. tantum longis; inflorescentia ad capitu-

lum singulum multicapitatum 2.5-3 cm. diam. pedunculatum reducta,

bracteis extimis linearibus foliaceis usque 1.5 cm. longis; involucrum

8 mm. altum, phyllariis ca. 8 glabris oblongo-ovatis acutiusculis

prope apicem albo-ciliatis arete imbricatis; corolla glabra 6-7 mm.

longa; achaenia anguste columnaria nigrescentia 4 mm. longa, pappo

albo fere aequilongo. — Mexico: Very wet meadow near San Juanito,

Distrito de Bocoyna, Chihuahua, alt. 2,430 meters, July 26, 1937,

Forrest Shreve 8031 (type in Herb. Field Mus.). Memlichi, Rio Mayo,

Chihuahua, in transition zone among pines, alt. 2,250 meters,

September, 1936, Howard Scott Gentry 2785; aquatic, a small colony

in meadow seep.

Verbesina Gentry! Standl., sp. nov. — Herba erecta inferne

ut videtur simplex 50 cm. alta et ultra, caule crasso dense scabro

vel hirtello fere ad capitula alato, alis viridibus integris usque 4 mm.

latis; folia opposita majuscula sessilia et longe basi in alas decurrentia,

ovata vel rhombico-ovata, caulina usque 14 cm. longa et 8 cm. lata

acuta vel acuminata dentata vel grosse dentata, crassa, utrinque

scaberrima; capitula numerosa corymbosa, inflorescentia usque 30

cm. longa, pedUnculis ultimis graciusculis exalatis brevibus dense

viscido-hirtellis et scabris, capitulis campanulatis radiatis 5-6 mm.

altis 7-10 mm. latis luteis; phyllaria vix ad 3 mm. longa inaequalia

oblonga acuta hispidula; radii pauci lineares 5-7 mm. longi lutei;

paleae disci latae apice apiculato-rotundatae glabrae. — Mexico:

Guasaremos, Rio Mayo, Chihuahua, in valley margins, August 23,

1936, Howard Scott Gentry 2416 (type in Herb. Field Mus.).

The heads are in young flower, and it is impossible to determine

the nature of the achenes. The collector describes the plant as a

tall, slender herb with orange heads. Its relationship is with Ver-

besina pterocaula DC., of southern Mexico, which differs conspicu-

ously in its larger heads as well as in several other respects.

STUDIES OF AMERICAN PLANTS— X

BY

PAUL C. STANDLEY

CURATOR OF THE HERBARIUM, DEPARTMENT OF BOTANY

V

NATURAL

HISTORY

THE LIBRARY OF THE

JUN291940

UNIVERSITY OF ILLINOIS

BOTANICAL SERIES

FIELD MUSEUM OF NATURAL HISTORY

VOLUME 22, NUMBER 2

JUNE 12, 1940

PUBLICATION 472

PRINTED IN THE UNITED STATES OF AMERICA

BY FIELD MUSEUM PRESS

STUDIES OF AMERICAN PLANTS-X

PAUL C. STANDLEY

The following pages are devoted chiefly to descriptions of new

plants, principally from extensive tropical American collections

received for determination during the past few years. Series that

have proved particularly rich in additions to the known floras of

Mexico and Central America are those of Mr. E. Matuda in Chiapas,

especially on the Volcano of Tacana; those of Mrs. M. E. Davidson

in the mountains of Chiriqui, Panama; and those from the high

and isolated peak of Cerro Potosi, in the State of Nuevo Leon,

Mexico, collected by Dr. C. H. Mueller and by Mr. Richard A.

Schneider. It is perhaps worthy of note that the most recent collec-

tions obtained in Costa Rica by Mr. Austin Smith, particularly

those of the San Carlos region, have continued to yield numerous

additions to the published catalogue of the Costa Rican flora.

Among the rather numerous new Rubiaceae here published,

probably most important are those gathered by Dr. A. Ducke

during his latest exploration of the Amazon Valley. In this instance,

as in case of the other series mentioned, further new species still await

description and publication.

CYCLANTHACEAE

Carludovica Killipii Standl., sp. nov. — Epiphytica, caudice

3 cm. crasso dense foliato brevi, vaginis dense imbricatis, inter-

nodiis brevissimis; folia magna longipetiolata coriacea, petiolo

crasso ca. 28 cm. longo, vaginis ca. 14 cm. longis basi fere 6 cm.

latis; lamina profunde bifida ca. 38 cm. longa, segmentis linearibus

3.5 cm. latis longe attenuato-acuminatis trinerviis; pedunculus 10 cm.

longus crassiusculus, nodis 2-3 ; spadix anguste oblongus 4.5 cm. longus

1.3 cm. latus apice rotundatus, densissime multiflorus. — Colombia:

In mangrove swamp, Buenaventura Bay, Dept. El Valle, April 13,

1939, E. P. Killip 34975 (type in Herb. Field Mus.).

Two inflorescences are present, one much weathered, the other

somewhat the worse for excessive moisture while the specimen

was being dried, so that floral details can not be determined readily.

A distinct species appears to be represented, and I have been unable

to associate it with any described one. The collector states that the

65

66 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

leaves, which are conspicuously distichous and apparently numerous,

form large, fan-like clusters. The habitat reported is unusual for

plants of this genus.

ARACEAE

Anthurium Allenii Standl., sp. nov. — Epiphytica, caudiculo

brevi crasso, internodiis brevissimis; petiolus subgracilis 20-26 cm.

longus 10-14 mm. infra apicem geniculatus, nodo non incrassato;

lamina oblongo-lanceolata prope basin paullo latior in sicco sub-

coriacea utrinque fusco-punctata, 30-40 cm. longa 6.5-10.5 cm.

lata, acuminata et subito cuspidata, basi obtusa vel breviter cuneata,

costa gracili utrinque prominente, nervis lateralibus primariis

numerosis gracillimis prominulis secondariis vix crassioribus angulo

semirecto adscendentibus ca. 7 mm. a margine in nervum distinctum

gracillimum collectivum conjunctis; pedunculi graciles 18-35 cm.

longi petiolis longiores vel breviores; spatha oblongo-lanceolata

5 cm. longa ca. 1 cm. lata apiculata; spadix breviter crasse stipitatus

cylindricus vel fere caudiformis, ad anthesin sursum paullo attenua-

tus, 9 cm. longus 5 mm. crassus, in statu fructifero 14 cm. longus

et fere 1 cm. crassus. — Panama: North rim of El Valle, Prov. Code",

June, 1939, P. H. Allen & A. H. G. Alston 1854 (type in Herb.

Mo. Bot. Gard.) ; May, 1939, Allen 1819.

This evidently is closely related to A. chiriquense, described

here, but that has epunctate leaves which are about equally narrowed

to each end, rather than punctate and broadest near the base, as

in A. Allenii.

Anthurium Baileyi Standl., sp. nov. — Planta majuscula

terrestris, caudice brevi vel fere nullo ca. 1.5 cm. crasso, internodiis

brevissimis; cataphylla brevia persistentia subcoriacea; petiolus

50-60 cm. longus gracilis prope basin 7 mm. crassus teres in sicco

brunnescens; lamina deltoideo-ovato-cordata in sicco tenuis atque

fere membranacea ca. 44 cm. longa atque 32 cm. lata, subito brevi-

terque acuminata, supra viridis et saepius dense nigro-punctata,

subtus aliquanto pallidior, lobis posticis magnis, rotundatis vel

latere exteriore obtuso-angulatis, sinu lato aperto triangulari ca.

12 cm. longo, nervis primariis costalibus utroque latere ca. 7 nervum

tenuem irregularem collectivum prope marginem formantibus,

nervis basalibus 5; pedunculus ca. 45 cm. longus gracilis teres brun-

nescens; spatha ut videtur viridis 12.5 cm. longa basi 1.5 cm. lata

lineari-lanceolata longissime attenuata spadice paullo brevior;

spadix 1.5 cm. longe stipitatus 12 cm. longus basi 7-8 mm. crassus

sursum paullo attenuatus obtusus. — Panama: Barro Colorado

STUDIES OF AMERICAN PLANTS— X 67

Island, Canal Zone, June 15, 1931, L. H. & Ethel Zoe Bailey 196

(type in Herb. Field Mus.).

The type specimen is the only one seen with an inflorescence,

but here probably belong the following leaf specimens, all except

one from the vicinity of the Canal Zone: Barro Colorado Island,

Standley 31294, 41104. Hills north of Frijoles, Standley 27440. Rio

Tapia, Prov. Panama, Standley 26159. Rio Tecumen, Prov. Panama,

Standley 26739. Puerto Obaldia, San Bias Coast, Pittier 4277.

The material listed above is rather puzzling. Offhand the leaf

specimens appear to represent a single species, and probably they

do, but there is considerable possibility also that two distinct species

if not two genera are represented. The matter can be decided only

when a larger number of flower-bearing specimens is available for

study. In the type specimen the leaves are glabrous beneath and

the petioles unarmed, but in several of the sterile specimens the

leaves are conspicuously and densely puberulent beneath, at least

along the nerves, and the petioles are decidedly aculeolate.

Anthurium chiapasense Standl., sp. nov. — Epiphytica, caudi-

culo brevi crasso, internodiis brevissimis, cataphyllis ferrugineis in

fibras mox solutis; petiolus crassiusculus subteres 25 cm. longus

fere 2 cm. infra apicem nodosus, nodo vix incrassato; lamina in

sicco subcoriacea subfusca elongato-oblongo-sagittata ca. 30 cm.

longa et prope medium 6.5-7.5 cm. lata, apice longe sensim atten-

uato-acuminata, basi profunde cordata, lobis posticis oblongis

rotundato-obtusis retrorsis vel paullo extrorsis, sinu ca. 7 cm. alto

angusto oblongo aperto, costa gracili utrinque prominente, nervis

lateralibus primariis utroque latere supra basin ca. 9 angulo semi-

recto adscendentibus utrinque prominentibus secundariis paullo

crassioribus remote a margine in nervum distinctum collectivum

plus minusve irregularem conjunctis, lamina basi 5-nervia; pedun-

culus gracilis 30 cm. longus; spatha ut videtur oblonga atque ca. 3

cm. longa; spadix crasse 4 mm. longe stipitatus anguste cylindricus

6 cm. longus prope basin 5 mm. crassus, sursum paullo attenuatus

obtusus. — Mexico: Mt. Ovando, Chiapas, April 4, 1936, E. Matuda

1562 (type in herb. Mo. Bot. Card.; photograph in Herb. Field Mus.).

Anthurium chiriquense Standl., sp. nov. — Epiphytica, caudi-

culo brevi crasso, internodiis brevissimis; petiolus subteres laminam

fere aequans 20-30 cm. longus crassiusculus ca. 8 mm. infra apicem

geniculatus, nodo incrassato; lamina anguste elliptico-oblonga

vulgo 29-35 cm. longa atque 7-10 cm. lata in sicco subcoriacea

epunctata, apice obtusa atque subito cuspidato-acuminata, basi

68 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

subobtusa vel late cuneata, costa gracili utrinque prominente, nervis

lateralibus primariis numerosis vix secundariis crassioribus angulo

semirecto adscendentibus, ca. 5-9 mm. a margine in nervum dis-

tinctum collectivum conjunctis; pedunculi crassi ca. 45 cm. longi

vulgo fere duplo petiolis longiores; spatha purpurascens oblongo-

lanceolata ca. 6.5 cm. longa atque 1 cm. lata acuminata; spadix

purpurascens graciliter cylindricus obtusus, interdum sursum paullo

attenuatus, 12-25 cm. longus, ca. 7 mm. prope basin crassus. —

Panama: In rain forest, Bajo Chorro, Boquete District, Prov.

Chiriqui, alt. 1,800 meters, Mrs. M. E. Davidson 283 (type in

Herb. Field Mus.), 312.

Related to A. turrialbense Engler, of Panama and Costa Rica,

which differs in having relatively much narrower, punctate leaves.

Anthurium garagaranum Standl., sp. nov. — Planta grandis

ut videtur epiphytica, caudice brevi crasso; petiolus crassus 40-60

cm. longus anguste sulcatus subteres ca. 1.5 cm. infra apicem geni-

culatus, nodo 7-8 mm. crasso; lamina profunde trilobata basi sub-

truncata, lobis lateralibus incurvo-falcatis 23-26 cm. longis 10-11

cm. latis obtusis vel subacutis, latere exteriore rotundatis, interiore

fere rectis, costa ca. 4 cm. supra basin in nervos primarios 4 desinente,

nervis crassis prominentibus fere ad apicem lobi productis, seg-

mento medio ca. 36 cm. longo et 9 cm. lato vel ultra, longe angus-

teque acuminate, costa subtus crassa prominente, nervis primariis

costalibus numerosis crassiusculis prominentibus angulo semirecto

vel latiore divergentibus, prope marginem in nervum collectivum

conjunctis; pedunculus ca. 25 cm. longus teres 5-7 mm. crassus;

spatha lanceolato-linearis ca. 17 cm. longa prope basin 1.5 cm. lata;

spadix gracilis caudiformis sessilis vel subsessilis ca. 17 cm. longus

atque prope basin 9 mm. crassus, sursum paullo attenuatus.—

Panama: Foothills of Garagara, Darien, at 30-500 meters, Pittier

5601 (type in U. S. Nat. Herb.).

The type material was determined by Krause as Philodendron

tripartitum (Jacq.) Schott, a rather common Central American

species which this plant does not greatly resemble in leaf form. It

appears to be related closely to Anthurium tripartitum Engler,

described from Barbacoas, Colombia. Engler's description agrees

in most respects with the Panama plant, but there are enough

discrepancies to make it evident that the latter is a distinct new

species.

Anthurium gracilens Standl., sp. nov. — Epiphytica vel

terrestris, caudice gracili ca. 4 mm. crasso brunnescente, internodiis

STUDIES OF AMERICAN PLANTS — X 69

elongatis; cataphylla lineari-attenuata brunnea decidua; petiolus

gracillimus lamina paullo brevior 10-22 cm. longus 1-1.5 cm. infra

apicem geniculatus, nodo gracili; lamina in sicco membranacea et

vulgo laete viridis, subtus paullo pallidior, epunctata, ovato-elliptica

ad ovato-oblonga 17-21 cm. longa 6.5-8 cm. lata cuspidato-acumi-

nata, vix basin versus angustata, basi late rotundata atque emargi-

nata vel subcordata, rare truncata, e basi 5-nervia, nervis primariis

utroque latere ca. 9 angulo semirecto vel latiore adscendentibus

tenerrimis, prope marginem in nervum collectivum tenerum

fere regularem conjunctis; pedunculus ca. 11 cm. longus gracillimus;

spatha lanceolato-oblonga viridis 1 cm. lata vel ultra; spadix graci-

liter 1 cm. longe stipitatus cylindricus obtusus, ca. 4.5 cm. longus

et 5-6 mm. crassus. — Panama: Rio Chiriqui Viejo Valley, Chiriqui,

July 13, 1939, Peggy White 160 (type in Herb. Field Mus.; duplicate

in Herb. Mo. Bot. Card.). Bajo Mona, Rio Caldera, Chiriqui,

1,500-2,000 meters, Woodson, Allen & Seibert 1026. Vicinity of

Cerro Punta, Chiriqui, P. H. Allen 1531.

Somewhat similar to A. pattens Schott, of the same general region,

which differs in its much narrower leaves, obtuse or rounded and

not at all emarginate or subcordate at the base.

Anthurium silvigaudens Standl. & Steyerm., sp. nov.—

Terrestris ca. metralis, caudice brevi ca. 1 cm. crasso cataphyllis

magnis 8-9 cm. longis diutius persistentibus fibrosis dense obtectis,

internodiis abbreviatis; petiolus crassiusculus 27 cm. longus evagin-

atus apice 1 cm. longe geniculatus; lamina ovato-cordata in sicco

tenuis et fere membranacea, ca. 37 cm. longa atque 22 cm. lata,

cuspidato-acuminata, basi profunde (ca. 11 cm.) cordata, lobis

posticis late rotundatis intus protractis, sinu clause angusto, e basi

7-nervia, nervo infimo intra lobum posticum nervos secundarios ca.

8 emittente, nervis primariis costalibus utroque latere ca. 8 ad vel

fere ad marginem prolongatis; pedunculi breves 7 cm. tantum longi;

spatha pallide viridis ovata vel lanceolato-ovata ca. 9.5 cm. longa

atque 4-5 cm. lata longiacuminata, basi late amplexicaulis et cor-

data; spadix subsessilis (perfectus non visus) purpurascenti-viridis

1 cm. crassus. — Guatemala: Terrestrial in wet forest near Tactic,

Dept. Alta Verapaz, alt. about 1,500 meters, April 5, 1939, Paul C.

Standley 70505 (type in Herb. Field Mus.).

In most respects this is similar to A. fraternum Schott, also

Guatemalan, but in that the leaves have 5 rather than 7 basal nerves,

and there are other less important differences.

70 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

MARANTACEAE

Calathea ursina Standl., sp. nov. — Herba perennis acaulis

ca. 35 cm. alta; folia 3-7 erecta, petiolo 13-18 cm. longo gracillimo

ubique dense pilis fusco-brunneis paten tibus longihirsuto fere 1 cm.

infra apicem geniculato, nodo non incrassato; lamina membranacea

elliptico-oblonga vel lanceolato-oblonga 11-17 cm. longa 4-5 cm. lata

longe attenuato-acuminata, basi acuta, utrinque sat dense pilis longis

plerumque patentibus hirsuta; pedunculus ca. 9 cm. longus dense

longihirsutus; inflorescentia condensata ca. 5 cm. longa atque 3 cm.

lata, bracteis luteis paucis tenuibus sublaxis omnibus pilis longis

patentibus brunneis hirsutis, exterioribus late ovatis imbricatis 1.5-2

cm. longis, interioribus solemniter elongatis elevato-venosis usque

3 cm. longis acuminatis erectis. — Peru: On rising ground, Gamita-

nacocha, Rio Mazan, Dept. Loreto, alt. 100-125 meters, February 20,

1935, Jose M. Schunke 294 (type in Herb. Field Mus.).

Local name, Bijahuillo. "Plants solitary."

MORACEAE

Brosimum Mello-Barretoi Standl., sp. nov. — Arbor 12-metralis

ut videtur dense ramulosa, ramulis teretibus brunnescenti-ochraceis

glabris, internodiis brevibus; stipulae attenuato-acuminatae 1 cm.

longae minutissime puberulae vel fere glabrae; folia modica

brevissime petiolata coriacea, petiolo crassiusculo usque 1 cm.

longo glabro; lamina anguste oblonga vel lanceolato-oblonga 9-15

cm. longa 3-5 cm. lata subito breviterque acuminata, acumine

ipso obtuso, basi paullo inaequali acuta ad late obtusa et saepe

breviter contracta, glabra, supra in sicco brunnescens sublucida,

costa venisque conspicuis sed vix elevatis, subtus paullo pallidior,

costa crassiuscula fere plana, nervis lateralibus utroque latere ca. 15

angulo lato saepe fere recto abeuntibus fere rectis tenerrimis planis

prope marginem arcuato-junctis, venulis arete reticulatis sed non

elevatis; capitulum fructiferum globosum breviter pedunculatum

sordide brunneum 12 mm. diam. — Brazil: Matta, Fazenda da Ca-

choeira, Municipio Tombos, Minas Geraes, July 18, 1935, H. Mello

Barreto 1658 (Jard. Bot. Bello Horizonte 16566; type in Herb.

Field Mus.).

Local name, Bainha de espada. The material, although ample,

is not in satisfactory state for study, as regards the inflorescences,

but it seems properly referable to Brosimum, in which genus I have

been unable to find a published name for it. It is a pleasure to name

this tree for its collector, who has made one of the largest and best

STUDIES OF AMERICAN PLANTS — X 71

prepared series of specimens ever obtained by any botanical worker

in Brazil.

Cecropia Garciae Standl., sp. nov. — Arbor 12-metralis; petiolus

crassus 42 cm. longus et ultra subteres dense laxeque albo-lanatus;

lamina magna crasse herbacea digitatim fere ad basin ca. 20-lobata,

segmentis anguste oblongo-oblanceolatis ca. 45 cm. longis et 12 cm.

latis, apice obtusis vel subrotundatis atque subito cuspidato-acumi-

natis, infra tertiam partem sensim ad basin angustatis integris,

supra viridibus glabris, subtus albidis ubique dense tomento minuto

arete adpresso indutis, nervis lateralibus numerosis angulo semi-

recto vel paullo latiore adscendentibus rectis teneris prominentibus

fere in marginem desinentibus apice abrupte sursum curvatis; in-

florescentia feminea crasse 3 cm. longe pedunculata, spicis 15 vel

ultra arete sessilibus 11-14 cm. longis 8 mm. crassis apice obtusis-

simis. — Colombia: In edge of forest along coast, east side of Gorgona

Island, Dept. Nariflo, February 11, 1939, E. P. Killip & Hernando

Garcia 33221 (type in Herb. Field Mus.).

Noteworthy for the very numerous leaf segments, bright green

above and covered beneath with a dense but minute and closely

appressed tomentum.

Coussapoa Duquei Standl., sp. nov. — Arbor 30-metralis,

trunco 1 m. diam., ramulis crassis fusco-ferrugineis in sicco rimosis

sparse praesertim prope nodos hirto-pilosis, internodiis brevibus;

folia modica longipetiolata crasse coriacea, petiolo crassiusculo

5-10 cm. longo hirtello vel glabrato; lamina late ovata vel elliptico-

ovata 15-18 cm. longa 10-11 cm. lata obtusissima, basi rotundata

et brevissime subpeltata, supra sublucida glabra, nervis manifestis

sed non elevatis, subtus brunnescens ubique sat dense laxeque

tomentosa, costa gracili elevata, nervis primariis lateralibus utroque

latere ca. 13 gracilibus prominentibus rectis obliquis angulo semi-

recto adscendentibus in marginem desinentibus, nervis alteris fere

aequicrassis parallelis interpositis, margine leviter undulato-crenato;

capitula (in statu nimis juvenili tantum visa) in cymas breves

parvas axillares petiolis breviores disposita pedunculata ut visa

5 mm. tantum diam., ramis pedunculisque dense molliter brunneo-

pilosis. — Colombia: Palestina, Dept. Caldas, alt. 1,500 meters, J. M.

Duque 1767 (type in Herb. Field Mus.).

The present species is similar to C. Lawrancei Standl., likewise

Colombian, but in that the petioles and branches are copiously

hirsute with long hairs.

72 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

Coussapoa incomitata Standl., sp. nov. — Arbor 20-metralis

in statu juvenili epiphytica, ramulis teretibus crassis fuscis in sicco

striatis primo tomentulosis, serius glabratis, internodiis brevissimis;

stipulae parvae late ovatae subacuminatae 5 mm. tantum longae

extus laxe pilis pallidis sericeae; folia parva petiolata crasse coriacea,

petiolo gracili 2.5-3 cm. longo laxe araneoso-tomentoso, serius

glabrato; lamina oblongo-obovata vel subcuneato-obovata 6-8 cm.

longa 3^4.5 cm. lata apice rotundata, basin versus saepissime paullo

angustata, basi ipsa anguste rotundata et vulgo breviter emarginata,

supra glabra vel tantum ad costam sparse tomentulosa, costa ner-

visque subimpressis, sublucida, subtus grisea vel brunnescens,

primo dense pilis laxis albis araneoso-tomentosa, glabrescens, costa

gracili prominente, nervis lateralibus utroque latere ca. 6 obliquis

angulo semirecto adscendentibus subarcuatis gracilibus elevatis,

venis transversis ultimisque laxe reticulatis prominentibus; pedun-

culi feminei ad nodos aggregati simplices crassi sordido-puberuli

vel glabrati, capitulis subglobosis densissime multifloris 6-7 mm.

diam., perianthio in statu fructifero fere 2 mm. longo obovoideo

apice rotundato sparse vel dense puberulo. — Brazil : Matta, Fazenda

da Cachoeira, Municipio Tombos, Minas Geraes, July 29, 1935,

H. Mello Barreto 1795 (Jard. Bot. Bello Horizonte 16810; type in

Herb. Field Mus.).

An apparently well marked species, noteworthy for the small

leaves and small, clustered fruiting heads. The local name is re-

ported as Mata pau vermelho.

Coussapoa Schunkei Standl., sp. nov. — Arbor 15-metralis,

trunco ca. 50 cm. diam., ramulis crassis sordide ochraceis vel brun-

nescentibus dense minute tomentellis atque ad nodos longipilosis,

internodiis brevissimis; stipulae 7 cm. longae vel ultra longiatten-

uatae dense puberulae et pilis longis pallidis subadpressis indutae;

folia modica petiolata coriacea, petiolo crassiusculo 4-5.5 cm. longo

dense minute puberulo et sparse longipiloso; lamina deltoideo-ovata

12-16 cm. longa 8-10.5 cm. lata apicem obtusum versus sensim

angustata, basi late rotundata et truncata vel subemarginata, supra

in sicco fusco-brunnescens glabra, nervis manifestis sed vix elevatis,

subtus pallide brunnescens, tomento minuto brunnescente densis-

sime obtecta, primo sparse laxe praesertim ad nervos costamque

araneoso-tomentosa, costa gracili elevata, nervis lateralibus utroque

latere ca. 10 rectis angulo semirecto adscendentibus gracilibus in

marginem desinentibus, basi 5-nervia, venis obscuris, margine

integro vel subundulato; inflorescentiae masculae laxe cymosae ca.

STUDIES OF AMERICAN PLANTS — X 73

8 cm. longae axillares fasciculatae, pedunculo gracili usque 4 cm.

longo, ramis late patentibus dense minuteque puberulis atque

sparse longipilosis, capitulis paucis globosis 2-3 mm. tantum diam.

2^4 mm. longe pedunculatis. — Peru: On river bank, Gamitanacocha,

Rio Mazan, Dept. Loreto, alt. 100-125 meters, January 27, 1935,

Jose M. Schunke 130 (type in Herb. Field Mus.).

Local name, Uvilla. "Solitary; flowers cream; fruit brown,

edible." Among Peruvian species, this is most closely related to

C. vellerea Klotzsch, whose type was collected at Macora, Huanuco,

by Ruiz and Pavon.

Pourouma chocoana Standl., sp. nov. — Arbor 8-10-metralis;

folia magna longipetiolata subcoriacea, petiolo crasso 17 cm. longo

dense scabro atque hispido-hirsuto; lamina ca. 25 cm. longa et 28 cm.

lata basi profunde (3.5 cm.) aperte cordata, ca. ad medium trilobata,

lobo medio late elliptico, costa media utroque latere nervos pri-

maries ca. 12 rectos emittente, lobis lateralibus late ovatis breviter

cuspidato-acuminatis, basi late rotundatis, lamina supra scabra et

tactu asperrima in sicco fusco-brunnescente, subtus ubique tomento

arete adpresso albo dense obtecta; inflorescentia feminea cymosa

multiramosa ca. 6 cm. longa atque 9 cm. lata multiflora 8 cm. longe

pedunculata, pedunculo ramisque dense tomento adpresso sordido-

brunneo indutis, pedicellis fructiferis crassissimis usque 5 mm.

longis; fructus maturus elliptico-ovoideus 1.5 cm. longus 1 cm. latus

paullo compressus obtusus dense sordideque brunneo-tomentellus.

—Colombia: Dense forest near junction of Rio Condoto and Rio

San Juan, Intendencia El Choco, alt. 100-150 meters, April 20,

1939, E. P. Killip 35095 (type in Herb. Field Mus.).

POLYGONACEAE

Goccoloba caurana Standl., sp. nov. — Arbor 15-metralis,

trunco 60 cm. et ultra diam., coma angusta, ramis erectis, cortice

brunnescente, ramulis crassis dense foliatis ochreis magnis 2 cm.

longis et ultra dense obtectis glabris; folia inter maxima breviter

petiolata crasse coriacea, petiolo crasso glabro 2.5-4 cm. longo;

lamina glabra oblonga 21-40 cm. longa 9-16 cm. lata apice obtusa

et subito breviterque acuminato-contracta, basi subrotundata vel

obtuse angustata, in sicco brunnescens, costa nervisque supra sub-

impressis, subtus paullo pallidior, costa gracili elevata, nervis latera-

libus utroque latere ca. 11 prominentibus obliquis angulo acuto

adscendentibus prope marginem arcuato-conjunctis, venis incon-

spicuis laxe reticulatis; flores dense racemosi, racemis ante anthesin

74 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

3.5 cm. longis axillaribus et dense fasciculatis, rhachi dense minute

sordido-puberula, pedicellis brevissimis; perianthium glabrum ca.

3 mm. longum, segmentis ovalibus apice rotundatis; stamina ex-

serta. — Venezuela: In forest on steep slope above the playa of Sal to

de Para, Medio Caura, State of Bolivar, alt. 200 meters, in 1939,

Llewelyn Williams 11366 (type in Herb. Field Mus.).

Local name Arahueke. "Trunk branched from the base; flowers

red, produced in March; fruits small and round." No fruits are

present with the type specimen. The species is noteworthy for its

exceptionally large leaves, almost perfectly oblong in outline.

Ruprechtia Tangarana Standl., sp. nov. — Frutex vel arbor

4-6-metralis, trunco 4-6 cm. diam., ut videtur dense ramosa, ramis

crassis fuscis vel sordido-ochraceis lenticellatis, internodiis brevibus

vel aliquanto elongatis glabris; folia breviter petiolata subcoriacea

in sicco fusca, petiolo crassiusculo 7-10 mm. longo glabro; lamina

lanceolato-oblonga vel elliptico-oblonga vulgo 6-21 cm. longa et

2.5-7.5 cm. lata, acuta usque longiacuminata, basi acuta vel sub-

obtusa, lucida glabra, nervis venisque supra prominulis, costa

gracili subtus elevata, nervis lateralibus utroque latere ca. 12 obliquis

angulo semirecto adscendentibus irregularibus teneris prominentibus,

venis prominulis arete reticulatis; flores masculi graciliter breviter

pedicellati parvi racemosi, racemis dense multifloris in axillis vel

ad nodos defoliates solitariis vel fasciculatis plerumque 2.5-4 cm.

longis, floribus glabris, pedicellis filiformibus glabris ochreolis vix

longioribus, staminibus exsertis; flores feminei in racemos breves

densissimos multifloros sessiles usque 3 cm. longos dispositi, pedi-

cellis brevissimis; calyx fructifer 1 cm. longus obtuse trigonus

utrinque sparse minute sericeus, segmentis exterioribus ca. 4 mm.

latis obovato-oblongis apice obtusis vel anguste rotundatis; fructus

acute trigonus 6 mm. longus lucidus brunneus. — Peru: On river bank,

Gamitanacocha, Rio Mazan, Dept. Loreto, alt. 100-125 meters,

January 25, 1935, Jose M. Schunke 115 (in fruit; type in Herb. Field

Mus.) ; February 15, 1935, Schunke 255.

Local name Tangarana blanca. "Flowers white." Only two

other species of Ruprechtia have been reported from Peru, and

neither of these from Loreto. R. Tangarana is more closely related

to R. apetala Wedd., which has shorter leaves, soft-pilose beneath,

elongate staminate racemes, and a fruiting perianth 2.5 cm. long.

CARYOPHYLLACEAE

Stellaria limitanea Standl., sp. nov. — Herba ut videtur pro-

cumbens laxe ramosa, caulibus 30 cm. longis et ultra gracilibus

STUDIES OF AMERICAN PLANTS — X 75

nodosis fragilibus sat dense glanduloso-villosis, internodiis saepe

foliis longioribus; folia petiolata herbacea parva, petiolo gracili

glanduloso-villoso 2-4 mm. longo; lamina ovato-deltoidea 8-14 mm.

longa 6-7 mm. lata (in statu bene evoluto probabiliter multo major)

sensim longiacuminata, basi truncata vel breviter cordata, utrinque

dense pilis longis viscidis villosa; pedicelli axillares solitarii 9-18 mm.

longi glanduloso-villosi; sepala oblongo-ovata 5-6 mm. longa sensim

acuminata tenuiter nervata hyalina viscido-villosula; petala non

visa, nulla(?); capsula late ovoidea obtusa sepalis fere aequilonga

lucida, stylis 3; semina suborbicularia 1 mm. lata ferruginea minute

obtuseque tuberculata. — Mexico: Volcan de Tacana, State of

Chiapas, alt. 1,400 meters, March, 1939, E. Matuda 2775 (type in

Herb. Field Mus.).

A relative of S. nemorum L., a widespread plant of Mexico and

Central America, this is notable for its large flowers and unusually

abundant and viscid pubescence.

MONIMIACEAE

Siparuna macra Standl., sp. nov. — Arbuscula gracilis 4-me-

tralis, ramulis gracilibus teretibus dense minute stellato-puberulis,

internodiis bene elongatis; folia parva petiolata membranacea,

petiolo gracili 1-2 cm. longo minute stellato-puberulo; lamina

oblongo-oblanceolata 5-11 cm. longa 1.5-3 cm. lata subito caudato-

acuminata, basin versus sensim attenuata, remote inaequaliter

denticulata, supra in sicco fusco-viridis sparse pilis minutis sessilibus

stellatis conspersa tactu asperula, subtus fere concolor densius pilis

similibus stellato-puberula, costa gracili prominente, nervis laterali-

bus utroque latere ca. 8 arcuatis angulo lato adscendentibus;

inflorescentiae axillares e basi ramosae ca. 3 cm. longae, ramis simpli-

cibus vel semel dichotomis gracilibus remote paucifloris sparse min-

uteque stellato-puberulis aliquanto scorpioideis, pedicellis usque

2.5 mm. longis; receptaculum 2-2.5 cm. latum et aequilongum sparse

inferne stellato-puberulum vel fere glabrum, apice glabrum. —

Costa Rica: Villa Quesada, Canton de San Carlos, Prov. Alajuela,

alt. 725 meters, edge of Caribbean rain forest, March 29, 1939, Austin

Smith F1828 (type in Herb. Field Mus.).

"Stems arching, the bark light brown; leaves dark green, faintly

shining above, paler beneath; flowers pale yellow." Among Costa

Rican species this is simple of recognition because of the small and

relatively narrow leaves, the very sparse and minute pubescence of

the leaves, and the small receptacles.

76 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

CRUCIFERAE

Thlaspi mexicanum Standl., sp. nov. — Herba perennis caes-

pitosa 4-10 cm. alta ubique glabra, caulibus erectis e radice incras-

sata pluribus; folia basalia parva longipetiolata, petiolo crassiusculo;

lamina ovata, rhombico-ovata vel oblongo-elliptica vulgo 6-9 mm.

longa et 3-5 mm. lata, apice obtusa vel anguste rotundata basi

cuneata, integra vel remote undulato-dentata; folia caulina 5-9 mm.

longa lanceolato-oblonga vel oblongo-ovata obtusissima, basi pro-

funde cordata et amplexicaulia pallida integra; racemi pauci- vel

multiflori sublaxi usque 2 cm. longi, pedicellis rectangule patentibus

crassiusculis 3-4 mm. longis; sepala purpurascentia 1.5 mm. longa

ovali-oblonga apice rotundata; petala alba angusta 2-2.5 mm.

longa; siliculae cuneato-obovatae 5-6 mm. longae 4 mm. latae

apice profunde emarginatae basi cuneatae sessiles angustissime

supra medium alatae; stylus brevis alas terminales siliculae vix

aequans. — Mexico: State of Nuevo Leon, abundant on meadows

above timber line, peak of Cerro Potosi, Municipio de Galeana, July

21, 1935, C. H. Mueller 2244 (type in Herb. Field Mus.). Sparse in

meadow below timber line, peak of Cerro Potosi, Mueller 2248.

In Payson's account of the genus Thlaspi (Univ. Wyo. Publ.

Bot. 1: 145. 1926), this would fall, in his key, under T. parviflorum

A. Nels., a species of Wyoming, which has decidedly narrower pods,

obtuse or truncate at the apex. I have seen no other Mexican

material of the genus, although some of the species might be expected

to occur in the mountains of Chihuahua and Sonora.

ROSACEAE

Potentilla Knoblochii Standl., sp. nov. — Herba perennis

humilis subacaulis; folia digitata pauca vel numerosa ca. 6 cm.

longa, petiolis 2-5 cm. longis pilis longis albidis patentibus vel

adscendentibus sat dense pilosis, alte lateque vaginatis; foliola pler-

umque 5 sessilia anguste oblonga vel oblanceolato-oblonga vulgo

1-2.5 cm. longa et 4-7 mm. lata, apice obtusa vel anguste rotundata

et brevissime obtuse tridentata, basin subobtusam versus paullo

angustata, supra griseo-viridia pilis longis adpresso-pilosa, subtus

albida dense adpresso-tomentosa et pilis longis adpresso-pilosa vel

sericea, supra partem tertiam basalem remote breviterque adpresso-

serrata; pedunculi vix folia aequantes, interdum breviores, 1-3-flori,

foliis 1-2 reductissimis onusti, pilis longis patulis pilosi; calyx dense

pilis longis patulis pilosus fere 5 mm. longus, sepalis oblongo-ovatis

acutis, bracteolis conformibus paullo brevioribus vel fere aequilongis;

STUDIES OF AMERICAN PLANTS — X 77

petala ut videtur pallide lutea 5 mm. longa apice late rotundata vel

subemarginata. — Mexico: Mojarachic, Chihuahua, May 4, 1938,

I. W. Knobloch 5104 (type in Herb. Field Mus.).

Prunus Lundelliana Standl., sp. nov. — Arbor omnino glabra,

ramulis gracilibus ferrugineis vel cinereo-brunnescentibus lenti-

cellatis rimosis; folia petiolata subcoriacea, petiolo 1-1.5 cm. longo;

lamina lanceolato-oblonga vel rarius lanceolata 7-10 cm. longa

2.5-4 cm. lata longiacuminata, acumine obtuso, basi acuta vel sub-

acuta, supra lucida, in sicco fusco-brunnescens, subtus brunnescens

glandulis 3-4 valde remote a basi positis orbicularibus onusta,

integra; racemi axillares vel e nodis defoliatis nascentes laxe pauci-

vel multiflori breviter pedunculati 3-5 cm. longi, pedicellis gracilibus

4-9 mm. longis; cupula calycis late campanulata fere 3 mm. lata et

aequilonga basi obtusa, sepalis brevissimis late rotundatis; petala

lata alba vix 2 mm. longa; stamina longiexserta; stylus crassiusculus

2.5 mm. longus. — Mexico: Hacienda Siltepec, State of Chiapas,

January, 1938, E. Matuda 1906 (type in Herb. Field Mus.). Santa

Rita, Mapastepec, Chiapas, January, 1938, Matuda 2016.

In most respects this agrees well with the description of Prunus

tetradenia Koehne, of Veracruz. In that, however, two of the leaf

glands are inserted near the base of the blade, rather than far remote

from it, and the leaves are described as substantially larger than in

P. Lundelliana.

LEGUMINOSAE

Acacia Gentlei Standl., sp. nov. — Kami crassi teretes fusco-

ferruginei dense elongato-lenticellati minute puberuli vel glabrati;

spinae binatae lucidissimae 4-4.5 cm. longae 5 mm. crassae teretes,

apice anguste attenuatae, glabrae curvae; folia breviter petiolata,

petiolo ca. 12 mm. longo infra apicem glandula elongata paullo

elevata instructo dense minute puberulo, rhachi brevi dense puberula

inter paria pinnarum elongato-glandulosa; pinnae 3-jugae 4-6 cm.

longae, rhachi dense puberula vel pilosula; foliola plerumque 9-18-

juga oblonga 8-13 mm. longa 3-5 mm. lata apice obtusissima mutica,

basi solemniter obliqua rotundata, lucida glabra vel glabrata, mar-

gine tantum minutissime ciliolato, e basi palmatinervia, costa

paullo excentrica utroque latere nervos paucos emittente, nervis

venulisque utrinque prominulis vel prominentibus; flores spicati,

spicis ad nodos solitariis pedunculatis ut videtur ca. 3 cm. longis,

floribus arete sessilibus; legumen graciliter 2-3 cm. longe stipitatum

falcato-lineare ca. 21 cm. longum et 8 mm. latum valde compressum

78 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

apice longissime et fere spiniforme attenuatum, basin versus longe

angusteque attenuatum, glabrum, marginibus incrassatis obtusisque.

—British Honduras: Without special locality, Percy Gentle 185

(type in Herb. Field Mus., received from Yale School of Forestry).

Called locally Red cockspur. This bullhorn acacia is remarkable

for the form and especially the size of its pods, still immature in the

specimen at hand. They are much longer than those of any related

species.

Bauhinia Castillo! Standl., sp. nov. — Ramuli gracillimi pallidi

glabri flexuosi; folia petiolata subcoriacea, petiolo gracillimo 18-22

mm. longo glabro; lamina integra anguste oblongo-lanceolata vel

triangulari-oblonga 8.5-9.5 cm. longa 2.5-3.7 cm. lata apicem

obtusum vel acutum versus sensim angustata, basi truncata vel

rotundata, supra glabra sublucida in sicco griseo-viridis, nervis ve-

nisque prominulis, venulis arete reticulatis, subtus grisea ubique

dense pilis minutissimis adpressis induta, basi 7-nervia, nervis 3

mediis ad vel fere ad apicem protractis elevatis gracilibus, venulis

laxe reticulatis; racemi axillares laxe multiflori ca. 4 cm. longi graci-

liter 3.5 cm. longe pedunculati, rhachi sparse hirtula, pedicellis fere

filiformibus ad 14 mm. longis puberulis; calyx 18 mm. longus acu-

minatus griseo-puberulus ad anthesin uno latere fissus; petala fere

linearia 18 mm. longa ad 2 mm. lata extus hirtella; stamen fertile 1,

anthera 4 mm. longa. — British Honduras: Freshwater Creek Re-

serve, February 17, 1939, A. Castillo 48 (type in Herb. Field Mus.).

Known locally by the name Cow-tongue. An unusually well dis-

tinguished species, because of the simple, entire leaves, which appear

glaucous on the lower surface on account of the dense, appressed

covering of minute, whitish hairs.

Cassia potosina (Britt. & Rose) Standl., comb. nov. Chamae-

crista potosina Britt. & Rose, N. Amer. Fl. 23: 283. 1930.— The

following collection agrees well with the original description of this

species, which was based upon Purpus 4832 from Minas de San

Rafael, San Luis Potosi: Jaumave, Tamaulipas, in 1932, H. W.

von Rozynski 546.

Inga chiriquensis Standl., sp. nov. — Arbor 4.5-9 m. alta,

ramulis crassis subteretibus densissime pilis brevibus brunneis

piloso-tomentosis, internodiis 3.5-4 cm. longis; folia vulgo 6-8-

foliolata, petiolo crasso 3-3.5 cm. longo nudo dense piloso-tomentoso,

rhachi 8-12 cm. longa dense hirtello-pilosa latiuscule alata 6-7 mm.

lata, glandulis majusculis; foliola sessilia inaequalia subcoriacea

STUDIES OF AMERICAN PLANTS — X 79

elliptico-oblonga vel obovato-oblonga 12-15 cm. longa 5-8 cm.

lata obtusa et apiculata, basi inaequali cuneato-obtusa, supra ubique,

ad nervos densius, fulvo-hispidula, costa nervisque prominentibus,

subtus dense molliter pilis lutescentibus patulis velutino-pilosula,

costa nervisque elevatis et insigniter conspicuis; flores spicati, spicis

brevibus densifloris crasse 3 cm. longe pedunculatis; legumen juvenile

subteres multicostatum 9 cm. longum et ultra 5 mm. diam. dense

molliter brunneo-tomentosum. — Panama: Volcan de Chiriqui, Bo-

quete District, Prov. Chiriqui, alt. 2,100 meters, July 13, 1938, Mrs.

M. E. Davidson 928 (type in Herb. Field Mus.).

Inga Davidsoniae Standl., sp. nov. — Arbor 4.5-9 m. alta dense

ramulosa, ramulis crassiusculis brunnescentibus dense molliter

piloso-tomentosis, internodiis brevibus; folia plerumque 6-foliolata,

petiolo ca. 3 cm. longo dense breviter piloso, anguste alato, rhachi

3-5 cm. longa hirtella vel pilosula anguste alata vix ultra 2 mm. lata,

glandulis parvis semiglobosis apice porosis; foliola subcoriacea

oblonga vel ovato-oblonga, saepe lanceolato-oblonga, 5-8 cm. longa

2-3 cm. lata acuta usque anguste rotundata, basi obliqua obtusa

vel anguste rotundata, supra lucida sparse praesertim ad costam

hirtella vel fere glabra, venis prominulis arete reticulatis, subtus in

sicco brunnescentia sparse hispidula vel glabrata, costa nervisque

prominentibus; flores ut videtur capitati; legumen anguste oblongum

paullo curvatum 17 cm. longum 4 cm. latum sessile, apice truncatum

et apiculatum, basi obliqua late rotundatum, subsparse fulvo-

hispidulum vel glabratum, marginibus paullo incrassatis bicarinatis,

seminibus ca. 16. — Panama: Volcan de Chiriqui, Boquete District,

Prov. Chiriqui, alt. 2,250 meters, July 13, 1938, Mrs. M. E. Davidson

943 (type in Herb. Field Mus.).

Lupinus cacuminis Standl., sp. nov. — Herba perennis elata

ca. 35 cm. alta, caule simplici vel pauciramoso erecto crassiusculo

in sicco subangulato vel striato, sat dense foliato, dense pilis brevibus

albidis arete adpressis sericeo; folia modica, petiolo 3-5.5 cm. longo

gracili sericeo 1-2 cm. longe vaginato, lobis liberis stipularum lineari-

attenuatis ca. 8 mm. longis; foliola vulgo 7 lineari-oblanceolata 2-4

cm. longa 4-5 mm. lata acuta vel subacuminata, basin versus sensim

angustata, supra viridia sparse breviter patulo-pilosa, subtus fere

concoloria sat dense sericea; racemi densi multiflori 8-10 cm. longi

4 cm. lati, pedicellis ca. 8 mm. longis dense pilis adscendentibus

pilosis; bracteae foliosae virides lineares persistentes ad 18 mm.

longae persistentes; calyx extus dense sericeus, fere ad basin bila-

biatus, segmentis lanceolato-linearibus, inferioribus 10 mm. longis,

80 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

superioribus paullo brevioribus, tubo brevi basi fere symmetrice

obtuso; vexillum fere 1.5 cm. longum apice late rotundatum, extus in

medio prope apicem sparse laxe sericeum; legumen lineari-oblongum

ca. 5 cm. longum atque 1.2 cm. latum ubique densissime pilis longis

patentibus laxe molliterque pilosum, apice subito acutatum et longi-

rostratum; semina ca. 5. — Mexico: State of Nuevo Leon, Sierra

Madre Oriental, ascent of Sierra Potosi by north hogback, about

20 miles northeast of Galeana, abundant on the alpine meadow and

in upper forest, alt. 3,450 meters, July 26, 1934, C. H. & M. T. Mueller

1257 (type in Herb. Field Mus.). Peak of Cerro Potosi, abundant

in upper pine woods and above timber line, July 21, 1935, C. H.

Mueller 2269.

The flowers are described as having a bluish lavender keel with

a central white area broadest at the apex; or as red in middle upper

lip and edge of lower lip, the ground color blue.

Lupinus LeSueurii Standl., sp. nov. — Herba perennis plus

minusve caespitosa 12-15 cm. alta, caudice apice dense basibus

persistentibus petiolorum obtecto; folia numerosa 4-8 cm. alta,

petiolo gracili 2-5 cm. longo dense pilis longis patulis albidis piloso

ca. 12 mm. longe vaginato, lobis liberis stipularum 4-5 mm. longis

attenuato-linearibus; foliola vulgo 7 anguste obovata vel elliptico-

oblanceolata 5-16 mm. longa 2.5-7 mm. lata, apice obtusa et cus-

pidato-apiculata, basin versus cuneato-angustata, utrinque sat

dense pilis laxis albidis subadpressis pilosa; pedunculus gracilli-

mus dense pilis longis patulis laxe pilosus erectus, racemo 3.5-6 cm.

longo laxe multifloro, pedicellis ca. 3 mm. longis dense pilosis; calyx

late campanulatus 4 mm. longus dense albo-pilosus, basi uno latere

auctus et subcalcaratus, segmentis labii inferioris lanceolato-lineari-

bus; bracteae deciduae lineari-lanceolatae alabastra subaequantes;

corolla ut videtur intense caerulea, vexillo 7 mm. longo extus glabro.

—Mexico: Salto Canyon, Babicora, Chihuahua, July 22, 1937,

Harde LeSueur 1346 (type in Herb. Field Mus.). Mojarachic,

Chihuahua, April 10, 1938, /. W. Knobloch 5070.

Lupinus Mueller! Standl., sp. nov. — Herba perennis elata

superne ramosa ca. 60 cm. alta, ramis gracilibus rigidis teretibus

densissime pilis albidis vel griseis brevibus adscendentibus vel

adpressis sericeis, sat dense foliatis, internodiis brevibus; folia parva,

petiolo gracili 10-14 mm. longo sericeo brevissime vaginato, lobis

liberis stipularum lineari-attenuatis ca. 8 mm. longis; foliola vulgo

7 anguste oblanceolato-oblonga plerumque 13-18 mm. longa et

3-4 mm. lata acuta vel acutiuscula atque cuspidato-apiculata,

STUDIES OF AMERICAN PLANTS — X 81

utrinque dense griseo-sericea; racemus ca. 9 cm. longus et 4 cm.

latus sublaxe multiflorus, pedicellis gracilibus usque 10 mm. longis

dense patulo-pilosis; bracteae lineares inconspicuae deciduae; calyx

extus dense albo-pilosus fere ad basin bilabiatus, segmentis lanceo-

lato-linearibus subaequalibus, inferioribus 7 mm. longis; vexillum

1.5 cm. longum glabrum. — Mexico: State of Nuevo Leon, abundant

throughout the pine woods, Las Canoas, on Cerro Potosi, Muni-

cipio de Galeana, July 17, 1935, C. H. Mueller 2205 (type in Herb.

Field Mus.).

The flowers are described as varying from blue through lavender

to purple.

Lupinus texensis Hook. f. albus Standl., f. nov. — E typo

speciei tantum petalis albis differt. — Texas: Madisonville, April 16,

1932, George L. Fisher 3225 (type in Herb. Field Mus.).

White-flowered plants are not particularly rare among the

lupines, and an albino form of the Texas bluebonnet was to be

expected, although I find no record that it has been reported formally.

Phaseolus phanerophlebius Standl., sp. nov. — Herba volubi-

lis, caule ramoso angulato gracili sparse pilis brevibus reflexis brun-

nescentibus piloso, internodiis brevibus; stipulae lanceolato-ovatae

5 mm. longae acutae erectae striato-nerviae; folia inter minora 3-

foliolata, petiolo gracili 2-3 cm. longo retrorso-piloso; foliola crasse

membranacea trinervia lineari-lanceolata vel anguste lanceolato-

oblonga 3.5-6 cm. longa 8-15 mm. lata acuta vel attenuato-acuta,

basi rotundata vel subtruncata, interdum prope basin subangulata,

supra lucida glabrata, subtus sparse hispidula, pilis brunnescentibus

plerumque patulis, venis prominentibus arete reticulatis; racemi

pauciflori ca. 12 cm. longe pedunculati nodosi ca. 3 cm. longi, brac-

teis striatis oblongo-lanceolatis 4 mm. longis; calycis glabrati tubus

late campanulatus 4 mm. longis, segmentis inferioribus lineari-

attenuatis 5-6 mm. longis; corolla intense caerulea 2.5 cm. longa.—

Panama: Trailing herb on open hillside, Boquete, Prov. Chiriqui

alt. 1,200 meters, June 25, 1938, Mrs. M. E. Davidson 757 (type in

Herb. Field Mus.).

Phaseolus Rimbachii Standl., sp. nov. — Herba volubilis

ramosa, caule gracili subtereti sparse saltern prope nodos pilis albis

laxis brevibus patulis consperso; stipulae virides lanceolatae acutae

striatae 3-4 mm. longae erectae persistentes; folia inter minora,

petiolo gracili ca. 3 cm. longo fere glabro, rhachi 1-1.5 cm. longa,

stipellis filiformibus 2-3 mm. longis; foliola breviter petiolulata

82 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

rhombico-ovata 3-4.5 cm. longa 2-3.5 cm. lata acuta vel acuminata,

basi latissime cuneata, membranacea, utrinque fere glabra, supra

viridia, subtus glaucescentia; pedunculi 7-12 cm. longi graciles fere

glabri, racemis paucifloris laxis usque 5 cm. longis, rhachi sparse

laxe pilosa, pedicellis curvis sparse pilosis usque 8 mm. longis,

bracteolis lanceolatis vix 2 mm. longis; calyx late campanulatus

3.5-4 mm. longus basi late rotundatus glaber subaequaliter breviter

lobatus, lobis latis ciliatis; vexillum 10-12 mm. longum glabrum;

ovarium glabrum apicem versus dilatatum, stylo 8 mm. longo dorso

infra apicem dense barbato. — Ecuador: Interandine highland, alt.

2,800 meters, A. Rimbach 292 (type in Herb. Field Mus.).

"A perennial herb, climbing to a height of 5 meters." The

glaucous or glaucescent under surface of the leaflets is a conspicuous

feature of the species.

Pithecolobium Davidsoniae Standl., sp. nov. — Arbor 6-12-

metralis fere glabra, ramulis teretibus gracilibus dense elevato-

lenticellatis brunnescentibus, internodiis brevibus vel elongatis

glabris vel obscure minute puberulis; folia parva plerumque 6-

foliolata, rhachi petioloque nudis sparse minute puberulis vel fere

glabris, petiolo eglanduloso, rhachi glandulis parvis depressis solis

inter foliola onusta; foliola membranacea brevissime petiolulata

inaequalia oblongo-elliptica vel elliptica 4-7.5 cm. longa 1.8-4 cm.

lata acuta vel obtusa, basi inaequali acuta, glabra vel glabrata,

subtus paullo pallidiora; flores viridescenti-lutei capitati, capitulis

globosis 1.5 cm. diam. axillaribus vel e nodis defoliatis nascentibus,

pedunculis 2-3 cm. longis glabris supra medium minute bracteatis;

calyx glaber 2-2.5 mm. longus tubulosus, dentibus triangulari-

ovatis tubo duplo brevioribus; corolla glabra tubulosa 6-7 mm.

longa gracilis, lobis obtusis vix ultra 1 mm. longis. — Panama: Volcan

de Chiriqui, Boquete District, Prov. Chiriqui, alt. 2,100 meters,

July 12, 1938, Mrs. M. E. Davidson 900 (type in Herb. Field Mus.).

Swartzia phaneroptera Standl., sp. nov. — Arbor 15-30-metra-

lis fere glabra, ramulis gracilibus subteretibus rimosis et lenticellatis;

folia magna pinnata 5-7-foliolata ca. 35 cm. longa vel ultra, petiolo

7-18 cm. longo versus apicem latiuscule alato, paribus foliolorum

4-5.5 cm. distantibus, rhachi latiuscule alata 5-6 mm. lata; foliola

subaequalia crasse membranacea oblonga, obovato-oblonga vel

oblanceolato-oblonga 12-19 cm. longa 5-8 cm. lata subito attenuato-

acuminata, basin versus paullo angustata, basi ipsa obtusa, supra

in sicco olivacea lucida ad costam puberula, aliter glabra, nervis

tenerrimis prominentibus, venulis prominulis arete reticulatis, sub-

STUDIES OF AMERICAN PLANTS — X 83

tus paullo pallidiora glabra, costa tenui elevata, nervis lateralibus

utroque latere ca. 7; inflorescentia (in statu fructifero tan turn visa)

12 cm. longe pedunculata pendula(?), ut videtur pauciflora, pedi-

cellis crassis fere 2 cm. longis; legumen (post dehiscentiam et vacuum

tantum visum) 1.5-2 cm. longe stipitatum teres glabrum lucidum

acuminato-rostratum 9-14 cm. longum, saepe hinc inde inter semina

abrupte constrictum, ca. 1.5 cm. latum. — Honduras: Dept. Colon,

Guarunta, Wispernini Camp, alt. 15-23 meters, in tropical rain

forest, March, 1938, Christine & Wolfgang von Hagen 1394 (type in

Herb. Field Mus.) ; also, from the same locality, but sterile, no. 1370.

Trifolium Schneider! Standl., sp. nov. — Herba pumila dense

caespitosa, caulibus plus minusve elongatis usque 8 cm. longis inter-

textis apice dense foliatis usque 3 mm. crassis, partibus infra apices

dense stipulis vetustis obtectis; stipulae lineari-lanceolatae 3-4 mm.

longae attenuatae; folia parva 3-foliolata, petiolo ad 1 cm. longo

sparse adpresso-pilosulo; foliola subsessilia cuneato-obovata 5-6 mm.

longa 4-5 mm. lata saepissime breviter emarginata glabra costato-

nervia; capitula brevissime pedunculata foliis multo breviora

plerumque 3-4-flora, bracteis brevissimis, floribus breviter pedi-

cellatis; calyx ca. 4 mm. longus basi acutiusculus 10-nervius, sparse

brevissime pilosulus, lobis lanceolato-linearibus rigidis attenuatis

erectis tubo fere duplo longioribus; corolla glabra 5-6 mm. longa

persistens; legumen glabrum turgidum apiculatum ca. 4 mm. longum.

—Mexico: In cracks between rocks, exposed alpine slope of Cerro

Potosi, alt. 3,750 meters, Municipio de Galeana, State of Nuevo Leon,

July 20, 1938, Richard A. Schneider 942 (type in Herb. Field Mus.).

"A plant whose tough, much branched stems grow in the cracks

between the rocks of the mountain top, leaving only the tips of the

branches exposed. The tiny leaves and flowers hug the surface.

Found only locally on the most exposed and rocky areas of the peak.

Not abundant anywhere, flowering and even fruiting plants being

difficult to locate. Observed only above timber line. Apparently

nearly out of bloom at this date. Flowers pure, pale pink."

This is one of the many interesting plants collected in 1938

on Cerro Potosi by the Mexican Biological Expedition of Students

of the University of Illinois. In habit and general aspect the plant

bears no close resemblance to any other Mexican clover, and it is

presumably one of the localized types confined, so far as now known,

to this isolated high peak.

Vicia canifolia Standl., sp. nov. — Herba cirrhifera perennis ca.

60 cm. alta simplex, caule striato dense pilis laxis albis patulis

84 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

piloso, internodiis plerumque ca. 2 cm. longis; stipulae lineares

erectae striatae 5 mm. longae; folia sessilia ca. 6 cm. longa cirrho

terminata, rhachi nuda dense albo-pilosula; foliola ca. 34 remota

linearia vulgo 1-2 cm. longa 1-1.3 mm. lata obtusa et apiculata

trinervia utrinque dense pilis laxis patulis albis pilosula, margine

interdum revoluto; racemi axillares solitarii secundi 2.5 cm. longe

pedunculati ca. 3 cm. longi dense multiflori (flores ad 25), pedicellis

vix 2 mm. longis pilosulis; calyx pallidus pilosulus 3-3.5 mm. longus

basi oblique rotundatus, dentibus subremotis angustis erectis;

corolla ut videtur alba caeruleo tincta 1 cm. longa, petalis glabris. —

Mexico: La Cima, Chihuahua, June 29, 1936, Harde LeSueur 714

(type in Herb. Field Mus.).

This may be only a variety of one of the normally green and

glabrate species of Mexico. The abundant, whitish, lax and spread-

ing pubescence gives the plant a pale coloring that is most distinctive.

RUTACEAE

Amyris tripetala Standl., sp. nov. — Arbuscula 3-6-metralis

omnino glabra, ramulis gracilibus tortuosis nigrescentibus longi-

trorsum rugosis; folia alterna plerumque trifoliolata, interdum

1-foliolata, petiolo gracillimo 1-2 cm. longo, rhachi gracili nuda ca.

2 cm. longa; foliola parva usque 3 mm. longe petiolulata vel sub-

sessilia inaequalia angustissime lanceolata 4-7 cm. longa 8-13 mm.

lata, longissime attenuata, acumine ipso obtuso, basin versus sensim

angustata, fere integra vel apicem versus sparse obscure crenulata,

supra lucida, nervis venisque prominulis, subtus paullo pallidiora,

costa tenui prominente, nervis lateralibus numerosis obliquis angulo

ca. semirecto adscendentibus inaequalibus fere rectis prominentibus,

venulis prominulis laxe reticulatis; inflorescentia terminalis cymoso-

paniculata sublaxe multiflora breviter pedunculata 5.5 cm. longa

atque 9 cm. lata vel minor, ramis primariis gracillimis plerumque

late divergentibus, floribus trimeris cymose aggregatis, bracteis late

ovatis minus quam 1 mm. longis, pedicellis vix ad 2 mm. longis;

sepala rotundo-ovata apiculata 0.5 mm. longa; petala 2.5 mm.

longa ovali-elliptica apice obtusissima 1-costata; stamina 3 petalis

fere duplo longiora, antheris ovalibus 1 mm. longis. — Panama:

Boquete, Prov. Boquete, alt. 1,140 meters, May 12, 1938, Mrs. M.E.

Davidson 621 (type in Herb. Field Mus.).

"Flowers creamy white." In its trimerous flowers the tree is

perhaps anomalous in the genus Amyris, to which it seems best

referred. The narrow leaflets distinguish it from other Central

American species.

STUDIES OF AMERICAN PLANTS — X 85

Zanthoxylum citroides Standl., sp. nov. — Kami inermes

teretes olivacei vel subochracei crassi aliquanto nodosi glabri, inter-

nodiis brevibus; folia modica glabra pinnata petiolata ut videtur

3-5-foliolata, petiolo 2.5-3 cm. longo superne late alato et 6-8 mm.

lato, inferne nudo vel anguste marginato, rhachi inter foliola late

alata et usque 1 cm. lata; foliola membranacea elliptico-oblonga

8 cm. longa atque 3 cm. lata vel minora acuminata, basi acuminata,

sparse grosseque pellucido-punctata, in toto margine minute crenu-

lata vel fere integra; flores parvi in axillis vel ad nodos defoliates

dense aggregati numerosi sessiles vel subsessiles trimeri; sepala

rotunda ca. 0.8 mm. longa apice late rotundata; petala lata fere 2

mm. longa apice rotundata. — Mexico: North slope of Volcan de

Tacana, Chiapas, alt. 2,100 meters, April 2, 1939, E. Matuda 2961

(type in Herb. Field Mus.).

Marked by the combination of few acuminate leaflets, broadly

winged leaf rachis, and densely crowded clusters of sessile flowers.

MELIACEAE

Trichilia Davidsoniae Standl., sp. nov. — Arbor fere omnino

glabra, ramulis crassis teretibus, novellis rugulosis minutissime

strigillosis, cito glabratis, internodiis ut videtur brevibus; folia

5-foliolata 5-5.5 cm. longe petiolata, petiolo rigido supra piano

minute strigilloso, rhachi 4-5 cm. longa ad nodos paullo incrassata;

foliola coriacea glabra 7-14 mm. longe petiolulata cuneato-oblongo-

obovata 7-12 cm. longa 4.5-5.5 cm. lata, apice late rotundata vel

subtruncata, basin versus longe sensim cuneato-angustata, supra

lucidissima, costa prominente, nervis venisque prominentibus atque

laxe reticulatis, margine plus minusve revoluto, subtus paullo palli-

diora opaca, costa elevata, nervis lateralibus utroque latere ca. 8

angulo lato divergentibus fere rectis prope marginem arcuato-

conjunctis prominulis, venis obscuris et non elevatis; inflorescentia

ut videtur parva axillaris pauciflora petiolo brevior, floribus breviter

crasseque pedicellatis; capsula 2 mm. longe crasseque stipitata

subglobosa sparse puberula vel fere glabra lucida in sicco rugosa

obtuse trigona fere 1.5 cm. longa 12 mm. lata, apice basique rotun-

data; semina ca. 9 mm. longa 5 mm. lata. — Panama: In rain forest,

Bajo Chorro, Boquete District, Prov. Chiriqui, alt. 1,800 meters, Feb-

ruary 28, 1938, Mrs. M. E. Davidson 347 (type in Herb. Field Mus.).

The material is incomplete and scant, having been taken from

a branch found on the ground, but it is sufficient to illustrate an

apparently quite distinct Central American species, marked by the

few, cuneate-obovate, coriaceous, very lustrous leaflets.

86 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

DICHAPETALAGEAE

Tapura bullata Standl., sp. nov. — Arbor parva, ramis tereti-

bus rimosis fusco-ferrugineis, internodiis ca. 4 cm. longis ochraceo-

tomentellis vel cito glabratis; folia magna brevissime petiolata

coriacea solemniter bullata, petiole crasso 10-14 mm. longo tomen-

tello vel glabrato; lamina anguste lanceolato-oblonga ca. 22 cm.

longa vel ultra et 5-6 cm. lata subsensim longiacuminata, basi valde

inaequilaterali uno latere rotundata altero acuta, supra in sicco

fusca lucida glabra, nervis venisque sat profunde impressis, subtus

paullo pallidior sparse hinc inde hirtella vel fere glabra, costa promi-

nente, nervis lateralibus utroque latere ca. 8 obliquis curvis pro-

minentibus angulo semirecto vel paullo latiore adscendentibus prope

marginem arcuato-junctis, venis elevatis laxe reticulatis; flores in

petiolo infra basin laminae dense aggregati parvi ca. 2.5 mm. longi,

sepalis anguste oblongis intus glabris extus dense pilis brevibus

ochraceis patulis vel subadpressis dense hirtellis. — Colombia: Vicinity

of Barranca Bermeja, Magdalena Valley, between Sogamoso and

Colorado rivers, Dept. Sur de Santander, alt. 100-500 meters, Janu-

ary 11, 1935, Oscar Haught 1510 (type in Herb. Field Mus.).

Probably the first species to be reported for Colombia of this

small genus, which ranges from the Guianas to Amazonian Peru.

The large, conspicuously bullate leaves distinguish it from all other

species.

EUPHORBIACEAE

Croton Skutchii Standl., sp. nov. — Arbor gracilis 27-metralis,

ramulis crassis in sicco angulatis sparse minute adpresso-lepidotis;

folia magna longipetiolata membranacea, petiolo crassiusculo 10-13

cm. longo dense vel sparse adpresso-lepidoto, apice glandulis 2

orbicularibus flavescentibus depressis onusto; lamina ovato-ovalis

ca. 24 cm. longa atque 14 cm. lata acuta vel breviter acuminata,

basi lata breviter cordata, supra viridis glabra, subtus paullo palli-

dior ubique minute sparse vel densius adpresso-lepidota, costa tenui

prominente, nervis lateralibus utroque latere ca. 17 angulo latiusculo

adscendentibus prominentibus teneris fere rectis prope marginem

dichotomis, venis transversis remotis subparallelis, venulis incon-

spicuis laxe reticulatis; racemi axillares longissimi breviter peduncu-

lati sat dense multiflori subcurvi 20-25 cm. longi ca. 1 cm. lati,

floribus infimis 3-5 femineis, reliquiis numerossissimis dense aggre-

gatis masculis; flos femineus usque 8 mm. longe pedicellatus, sepalis

foliaceis late ellipticis 8 mm. longis ut videtur post anthesin paullo

accrescentibus apice obtusis vel rotundatis utrinque densissime

STUDIES OF AMERICAN PLANTS — X 87

lepidotis; ovarium pilosum, stylis repetite dichotomis; alabastra

mascula ca. 2.5 mm. diam., calyce extus dense lepidoto ad medium

lobato, lobis late ovatis obtusis; petala sepalis paullo longiora ciliata.

— Costa Rica: In clearings, vicinity of El General, Prov. San Jose",

alt. 670 meters, June, 1939, Alexander F. Skutch 4377 (type in Herb.

Field Mus.).

Pera barbellata Standl. Field Mus. Bot. 8: 19. 1930.— When

this species was published, a single collection was known, Schipp

201, from British Honduras. In more recent years several additional

specimens have come to hand, and they may be reported as follows:

British Honduras: Machaca, in forest, alt. 15 meters, a tree of 9

meters, the trunk 22 cm. in diameter, August, 1933, Schipp S578. All

Pines, in secondary forest, at sea level, a tree of 12 meters, Schipp 568.

—Guatemala: Santa Teresa, Subin River, PeteX April, l93S,Lundell

2686. La Libertad, Pete"n, April, 1933, Lundell 3067.— Mexico:

Uvero, Oaxaca, near sea level, May, 1937, L. Williams 9412.

Regarding the Mexican tree Mr. Williams has supplied the

following notes: Local name Palo prieto. A tree 21 m. tall, with

narrow crown; trunk round, moderately straight, 40-70 cm. in

diameter, unbranched for about 16 m.; bark 6-12 mm. thick, choc-

olate-brown with gray patches and with small scales; inner bark

deep yellow. Sapwood white to deep yellow, darkening slightly

on exposure to the air; heartwood distinctly defined, medium brown

to chocolate-brown, composing about one- third of the wood; has a

slightly unpleasant odor when freshly cut; used mostly for railroad

ties. In forest on dry slopes and in low, slightly humid land.

Phyllanthus Millei Standl., sp. nov. — Frutex 2-metralis

distiche ramosus, ramis subteretibus viridibus pallide lenticellatis,

novellis angulatis dense puberulis vel minute villosulis, internodiis

foliis brevioribus; folia parva biseriatim inserta brevissime petiolata

tenuiter membranacea, petiolo tenui vix 1.5 mm. longo; lamina

ovato-ovalis vel rotundato-ovalis 8-13 mm. longa 7-9 mm. lata

apice late rotundata et interdum breviter emarginata, basi obtusa

vel rotundata, utrinque sparse sordido-puberula, nervis lateralibus

utroque latere ca. 4 arcuatis inconspicuis; flores axillares solitarii

vel saepius paucifasciculati, pedicellis filiformibus solemniter in-

aequalibus minute puberulis usque 4 mm. longis; sepala 5 ovalia vel

ovata fere 2 mm. longa apice obtusissima pallido-marginata. —

Ecuador: In regione tropica, ad Bahia, January, 1938, Luis Mille

1061 (type in Herb. Field Mus.).

88 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

AQUIFOLIACEAE

Ilex chiriquensis Standl., sp. nov. — Arbor omnino glabra,

ramis crassis ochraceis teretibus, internodiis brevibus; folia modica

petiolata crasse coriacea, petiolo crasso canaliculate 8-12 mm.

longo; lamina in sicco subfusca rotundato-elliptica vel late cuneato-

obovata 3.5-6 cm. longa 3-5 cm. lata apice late rotundata et sub-

emarginata, basi obtusa vel late cuneata, infra medium integra,

supra medium obscure et brevissime remoteque adpresso-crenata,

crenis utroque latere ca. 4, supra lucida, costa prominula, nervis

manifestis sed non elevatis, subtus opaca sparse nigro-punctata,

costa crassiuscula prominente, nervis lateralibus utroque latere ca.

4 angulo lato divergentibus remote a margine irregulariter con-

junctis, venis vix prominulis laxe reticulatis; flores in axillis vel ad

nodos defoliates fasciculati, pedicellis crassis usque 3 mm. longis

(et ultra?); calyx 1.5-2 mm. altus breviter lobatus, lobis late ro-

tundatis. — Panama: In rain forest, Bajo Chorro, Boquete District,

Prov. Chiriqui, alt. 1,800 meters, February 1, 1938, Mrs. M. E.

Davidson 243 (type in Herb. Field Mus.).

The available material is scant, consisting of a small branch

found upon the ground, with a few leaves and flower buds. The

leaves are quite unlike those of any species previously known from

southern Central America, being unusually wide, broadly rounded at

the apex, and few-nerved.

Ilex Davidsoniae Standl., sp. nov. — Arbor humilis omnino

glabra, ramis subteretibus in sicco fuscis lenticellis pallidis elevatis

conspersis, internodiis aliquanto elongatis; folia modica breviter

petiolata coriacea in sicco fusca, petiolo crasso 5-7 mm. longo canali-

culate; lamina oblongo-elliptica ca. 9.5 cm. longa et 4.5 cm. lata

subabrupte acuminata, acumine ipso obtuso, basi obtusa, fere ad

basin grosse inaequaliter crenata (dentibus utroque latere ca. 14),

supra lucida, costa impressa, nervis fere obsoletis, costa subtus

elevata, nervis lateralibus utroque latere ca. 8 angulo lato divergenti-

bus; flores feminei numerosi umbellati, umbellis in axillis aggregatis

paucifloris breviter pedunculatis, pedicellis fructiferis crassis angu-

latis 3-5 mm. longis; calyx sub fructum patens 2.5 mm. latus in-

crassatus brevissime lobatus, lobis latis rotundatis subapiculatis;

fructus immaturus ellipsoideo-globosus 4 mm. longus stylo magno

depresso coronatus. — Panama: In rain forest, Bajo Chorro, Boquete

District, Prov. Chiriqui, alt. 1,800 meters, January 25, 1938, Mrs.

M. E. Davidson 166 (type in Herb. Field Mus.).

STUDIES OF AMERICAN PLANTS — X 89

RHAMNACEAE

Gouania hypoglauca Standl., sp. nov. — Frutex scandens 5-

metralis, ramis crassis teretibus brunneis sat dense breviterque

pilosis, pilis patulis brunneis, internodiis foliis multo brevioribus;

folia modica petiolata subcoriacea, petiolo crassiusculo 7-8 mm.

longo dense brunneo-hirtello; lamina oblongo-ovata vel ovali-ovata

5.5-8.5 cm. longa 3-4.5 cm. lata subito acuta vel breviter acuminata,

basi truncata vel emarginata, supra in sicco fusca sparse pilis patulis

brunnescentibus pilosa, costa nervisque anguste impressis, subtus

griseo-fulvida ubique tomento arete adpresso densissime obtecta,

ad nervos adpresso-pilosa, e basi trinervia, costa gracili prominente,

nervis lateralibus utroque latere ca. 3 angulo semirecto adscenden-

tibus, margine remote inconspicue adpresso-crenato; flores spicati,

spicis (juvenilibus cum alabastris tantum visis) dense multifloris

breviter pedunculatis racemose paniculatis. — Costa Rica: Villa

Quesada, Canton de San Carlos, Prov. Alajuela, alt. 750 meters, edge

of Caribbean rain forest, April 14, 1939, Austin Smith F1952 (type

in Herb. Field Mus.).

From all other Gouania species of Central America, the present

Costa Rican plant is distinguished by the very dense, yellowish

gray, closely appressed tomentum of the lower leaf surface.

TILIACEAE

Sloanea Picapica Standl., sp. nov. — Arbor 15-metralis ut

videtur dense ramulosa, ramulis sat gracilibus et dense foliatis

fusco-ferrugineis pallido-lenticellatis minutissime sericeis vel fere

glabris; folia parva coriacea breviter petiolata in sicco subfusca,

petiolo subgracili 5-7 mm. longo glabrato; lamina lanceolato-ovata

vel elliptico-oblonga 5-7 cm. longa 2-3 cm. lata subabrupte longe

anguste acuminata, acumine ipso obtuso, basi obtusa vel subacuta,

glabra, supra sublucida, nervis venisque non elevatis, subtus paullo

pallidior, costa gracili prominente, nervis lateralibus utroque latere

ca. 7 teneris prominentibus angulo lato adscendentibus remote a

margine arcuato-conjunctis, venis obscuris laxe reticulatis; flores

axillares solitarii vel breviter racemosi, pedicellis fructiferis rectis

ca. 1 cm. longis minutissime hirtellis; capsula ellipsoidea 4-valvata

monosperma 1.5-2 cm. longa basi et apice obtusa dense purpuras-

centi-tomentosa, ubique dense setis glabris purpurascentibus usque

7 mm. longis induta; semen ovale 1 cm. longum 6 mm. latum.—

Honduras: Tropical rain forest, alt. 15-23 meters, Guarunta, Wisper-

nini Camp, Dept. Colon, March, 1938, Christine & Wolfgang von

Hagen 1390 (type in Herb. Field Mus.).

90 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

Local name, Picapica. In its small, glabrous, long-acuminate

leaves in combination with the form of its fruit this is altogether

unlike any other species of Sloanea known from Central America.

MALVACEAE

Sida turneroides Standl., sp. nov. — Herba perennis ca. 30 cm.

alta, caulibus solitariis vel pluribus gracilibus rigidis simplicibus

vel pauciramosis erectis viridibus minutissime pilis stellatis sessilibus

conspersis, internodiis saepissime foliis paucis longioribus; stipulae

filiformes 6-8 mm. longae; folia parva breviter petiolata membra-

nacea, petiolo gracili 4-7 mm. longo stellato-puberulo; lamina anguste

oblonga usque ovali-oblonga 2-3.5 cm. longa 1-1.5 cm. lata, apice

obtusa vel anguste rotundata atque mucronata, basi obtusa vel

rotundata, grosse inaequaliter serrata, supra laete viridis sparse

setuloso-pilosa vel fere glabra, subtus pallidior pilis minutis stellatis

sparse conspersa; flores axillares et solitarii vel ad apicem caulis

pauci et aggregati, pedunculis 7 mm. longis vel brevioribus crassius-

culis minute stellato-puberulis; calyx late campanulatus 6-7 mm.

longus angulatus extus sparse minute stellato-pubescens, ad costas

adpresso-pilosus, fere ad medium lobatus, lobis late triangularibus

acutis vel acuminatis erectis; petala ca. 12 mm. longa glabra; car-

pella fructus 7 glabrata 3 mm. longa obtusa, apice spinis 2 rigidis

erectis 1.5 mm. longis onusta. — Mexico: Jaumave, Tamaulipas,

in 1932, H. W. von Rozynski 514 (type in Herb. Field Mus.).

In general appearance the plant has some resemblance to Sida

acuta Burm., and the leaves probably are distichous as in that species.

BOMBACACEAE

Hampea latifolia Standl., sp. nov. — Arbor humilis, ramulis

crassiusculis glabratis sed pilis paucis remotis parvis stellatis con-

spersis, internodiis brevibus; folia magna longipetiolata mem-

branacea, petiolo tereti usque 16 cm. longo crassiusculo glabrato;

lamina rotundo-cordata 15-21 cm. longa 15-20 cm. lata acuta,

apice ipso obtuso, basi profunde cordata, obscure 3-angulata, mar-

gine integro vel subundulato, supra fere glabra sed praesertim ad

venas pilis paucis parvis sessilibus stellatis conspersa, subtus paullo

pallidior, subdense pilis parvis stellatis induta, basi 7-nervia; flores

in axillis 2-4-fasciculati, pedicellis crassiusculis usque 1.5 cm. longis

apicem versus paullo incrassatis dense stellato-furfuraceis, pilis

sessilibus vel ex parte stipitatis; calyx 1 cm. longus vel paullo ultra

extus densissime albide stellato-tomentellus in anthesi irregulariter

breviter lobatus; petala ca. 3.5 cm. longa apice obtusa vel late sub-

STUDIES OF AMERICAN PLANTS — X 91

truncata atque ad 2 cm. lata inaequilateralia remote fusco-punctata,

latere altero densissime stellato-furfuracea; stamina numerosissima

petalis paullo breviora. — Guatemala: Finca San Jos£ Nil, Retal-

huleu, August 23, 1936, W. R. Hatch & C. L. Wilson 383 (type in

Herb. Field Mus.).

In the writer's key to the species of Hampea (Journ. Wash.

Acad. Sci. 17: 395. 1927) this would run at once to H. platanifolia

Standl., a plant of the Pacific lowlands of Costa Rica. That is

evidently distinct in its more abundant, coarse, rather than close

and more or less furfuraceous, pubescence.

GUTTIFERAE

Clusia conferta Standl., sp. nov. — Arbor omnino glabra,

ramulis crassis, internodiis brevibus; folia magna fere sessilia coria-

cea, petiolo ca. 1 cm. longo sed late usque ad basin alato; lamina

oblonga vel aliquanto cuneato-oblonga 21-24 cm. longa 9 cm. lata,

apice rotundata, basin versus paullo angustata, basi ipsa late cu-

neata, in sicco brunnescens, nervis supra manifestis sed non elevatis,

subtus concolor, costa crassa elevata, nervis lateralibus utroque

latere numerosis tenerrimis angulo latiusculo abeuntibus; inflores-

centia terminalis capitato-congesta 5 cm. longe pedunculata, floribus

ca. 9 sessilibus, bracteis late ovatis coriaceis concavis arete adpres-

sis ca. 8 mm. longis apice obtusis vel rotundatis; sepala 4 decussata

crasse coriacea rotundata, saepe latiora quam longa, apice late

rotundata; ovarium magnum subglobosum, columna stylari crassa

fere 4 mm. longa 2.5 mm. crassa, stigmatibus 6 ovato-ovalibus 2 mm.

longis minutissime papillosis. — Guatemala: Banks of Rio Dulce,

Dept. Izabal, March 28, 1939, C. L. Wilson 401 (type in Herb.

Field Mus.).

Clusia orthoneura Standl., sp. nov. — Arbor 12-18-metralis,

trunco ut dicitur 5-7.5 cm. diam., ramulis subgracilibus rugosis

ferrugineis, in sicco transverse rugulosis vel rimosis, internodiis

plerumque brevibus; folia inter minora petiolata coriacea, petiolo

crassiusculo 8-12 mm. longo; lamina anguste lanceolato-oblonga

utrinque fere aequaliter angustata 6.5-9.5 cm. longa 1.8-3.5 cm.

lata, apicem versus sensim attenuata, apice obtuso vel acuto, basin

versus sensim attenuata, costa nervisque supra manifestis sed non

elevatis, subtus concolor, costa tenui basin versus elevata sursum

evanescente, nervis lateralibus obscuris et non elevatis angulo

angustissimo adscendentibus, cum costa fere parallelis, utroque

latere ca. 7; inflorescentia terminalis 1-3-flora 2 cm. longe peduncu-

92 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. ^22

lata, pedicellis crassis compressis ca. 1.5 cm. longis, bracteis concavis

latissime ovatis 6-7 mm. longis obtusis; sepala 4, exteriora 13 mm.

longa et bene latiora apice late rotundata arete adpressa, interiora

8 mm. tan turn longa; petala 2 cm. longa et ultra ca. 2.5 cm. lata;

columna staminalis 8 mm. alta 7 mm. crassa, staminibus ca. 20,

filamentis crassis ca. 6 mm. longis, antheris recurvis paullo ultra

2 mm. longis. — Colombia: Region of Chapon, State of Boyaca,

100 miles northwest of Bogota, in low, dense forest, alt. 1,080 meters,

June 2, 1932, A. E. Lawrance 131 (type in Herb. Field Mus.).

"Flowers pinkish white with yellow stamens. An epiphytic

tree, growing high up on the broken trunk of another forest giant."

The species is noteworthy for the unusually narrow leaves with but

few nerves running almost parallel with the costa.

COCHLOSPERMACEAE

Cochlospermum trilobum Standl., sp. nov. — Frutex metralis,

ramulis teretibus longitrorsum in sicco rugosis ferrugineis pilis

patulis brevibus sat dense pilosulis; folia modica longipetiolata

membranacea, petiolo gracillimo 3-6 cm. longo pilosulo; lamina 3-6

cm. longa 4-8.5 cm. lata, basi profunde aperte cordata, profunde

triloba e basi trinervia, supra viridis sparse breviter pilosa, subtus

pallidior dense velutino-pilosa, lobis subaequalibus, posticis late

patentibus ellipticis vel ovato-ellipticis apice obtusis vel rotundatis

inaequaliter adpresso-serratis; pedicelli 7-15 mm. longi minute

puberuli; sepala inaequalia 17 mm. longa late oblonga usque late

elliptica apice late obtusa vel rotundata dense minutissime puberula;

petala ca. 3 cm. longa; antherae ca. 5 mm. longae lineares; capsula

ovoidea 5.5 cm. longa 3 cm. lata acuta, longitrorsum dense striata.—

Bolivia: Robore", alt. 416 meters, in dry soil on pampas, October,

1934, Martin Cardenas 2970 (type in Herb. Field Mus.).

Apparently this is an unusually well marked species, quite

unlike any of which I have seen material or description. Char-

acteristic are the trilobate rather than 5-lobate or digitate leaves.

The stature also may be distinctive, although the common Central

American Cochlospermum vitifolium, while usually a small or rather

large tree, often flowers when only one or two meters high.

THYMELAEACEAE

Daphnopsis Mello-Barretoi Standl., sp. nov. — Frutex metralis,

ramis gracilibus teretibus ferrugineis lenticellis parvis pallidis con-

spersis, novellis densissime pilis brevibus ochraceis adscendentibus

tomentosis, internodiis abbreviatis; folia parva breviter petiolata

STUDIES OF AMERICAN PLANTS — X 93

crasse membranacea vel subcoriacea, petiolo crasso 3-4 mm. longo;

lamina late oblonga vel ovali-oblonga 3-5 cm. longa 1.5-2.5 cm.

lata apice rotundata, basi obtusa vel subrotundata et breviter subi-

toque contracta, supra sat dense pilis plerumque patulis hirtello-

pilosa, nervis inconspicuis, subtus paullo pallidior dense molliterque

pilosa, costa tenui prominente, nervis lateralibus utroque latere

ca. 5 arcuatis angulo lato adscendentibus, venulis prominulis laxi-

uscule reticulatis; inflorescentiae parvae numerosae pauciflorae,

vel elongatae multiflorae et racemiformes, ramis dense ochraceo-

pilosis, floribus breviter crasseque pedicellatis; calyx ad basin fructus

persistens 2.5-3 mm. longus dense tomentulosus, lobis oblongo-

ovatis acutis; stylus brevis, stigmate globoso-capitato; fructus late

ellipticus glaber 1 cm. longus 7 mm. latus basi et apice rotundatus.—

Brazil: Sujo, Jardim Botanico, Bello Horizonte, uncommon, Janu-

ary 8, 1934, H. Mello Barreto 7556 (Jard. Bot. Bello Horizonte

21723; type in Herb. Field Mus.).

Evidently related to D. brasiliensis Mart., which differs normally

in having much larger and more elongate leaves, conspicuously

narrowed at each end.

MYRTACEAE

Calyptranthes Killipii Standl., sp. nov. — Frutex 2.5 m. altus,

ramulis subteretibus tortuosis fuscis vel fusco-griseis, internodiis

brevibus, primo sparse minute ferrugineo-sericeis cito glabratis;

folia modica breviter petiolata coriacea, petiolo crassiusculo 5-6 mm.

longo glabrato; lamina elliptica vel obovato-elliptica 6-7.5 cm.

longa 2.5-3.5 cm. lata apice obtuse contracta et abrupte caudato-

acuminata, acumine usque 13 mm. longo angusto subobtuso, basi

cuneato-acuta vel late acuta, supra in sicco griseo-olivacea glaber-

rima dense minute puncticulata, costa anguste impressa, nervis

venisque occultis, subtus in statu juvenili ferruginea et dense arete

sericeo-tomentosa, cito glabrata vel sparse minuteque griseo-sericea,

costa gracili elevata, nervis lateralibus obsoletis, margine subre-

voluto; flores parvi cymoso-paniculati, cymis paucifloris ca. 1 cm.

longis et fere 2 cm. latis basi trichotomis, floribus sessilibus vel

breviter pedicellatis, bracteis minutis; calyx ante anthesin obo-

voideus 2.5-3 mm. longus apice rotundatus dense minutissime

ferrugineo-sericeus. — Colombia: In tidal thicket, Buenaventura

Bay, Dept. El Valle, May 4, 1939, E. P. Killip 35514 (type in Herb.

Field Mus.).

Eugenia choapamensis Standl., sp. nov. — Arbor 12-metralis,

ramis teretibus fusco-griseis glabris, internodiis brevibus vel rarius

94 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

elongatis; folia majuscula breviter petiolata subcoriacea, petiolo

crassiusculo 5-7 mm. longo late canaliculate; lamina anguste ob-

longa vel lanceolatooblonga 12-16 cm. longa 4-5.5 cm. lata acuta

vel acuminata, acumine obtuso, basi acuta, glabra, supra in sicco

fusca, costa basin versus subimpressa, nervis obscuris, subtus palli-

dior brunnescens, costa gracili elevata, nervis lateralibus utroque

latere ca. 10 angulo lato abeuntibus teneris prominentibus fere

rectis remote a margine arcuato-conjunctis; flores axillares vel

saepius e nodis defoliatis nascentes, interdum ut videtur breviter

racemosi, pedicellis 10-13 mm. longis rigidis, bracteis latis vix 1 mm.

longis patentibus; fructus ellipsoideo-oblongus teres vulgo 2-2.5

cm. longus 8-12 mm. latus glaber, basi et apice obtusus, calycis

lobis apice fructus persistentibus vix 2 mm. longis late deltoideo-

ovatis apice rotundatis. — Mexico: Near stream in arroyo, San

Juanito, Distrito Choapam, State of Oaxaca, alt. 375 meters, April 1,

1938, Ynes Mexia 9254 (type in Herb. Field Mus.). In original

forest, trail from San Juanito to La Trinidad, alt. 460 meters, April

13, 1938, Mexia 9296.

"Flowers white; mature fruit black, edible; bark thin, grayish

outside, reddish within." In most respects E. choapamensis re-

sembles E. pleurocarpa Standl., of the State of Jalisco, but that

has smaller and relatively broader leaves, and conspicuously costate

fruit.

MELASTOMACEAE

Blakea bella Standl., sp. nov. — Arbor, ramulis gracilibus

teretibus pallide brunneis rimosis, internodiis brevibus vel ali-

quanto elongatis, minute sparse vel subdense stellato-furfuraceis,

serius glabratis; folia modica petiolata crasse membranacea, petiolo

1.5-3 cm. longo crassiusculo sparse minuteque stellato-furfuraceo;

lamina triplinervia elliptico-oblonga vel anguste elliptica 10-24 cm.

longa 4-8 cm. lata subabrupte cuspidato-acuminata, acumine ipso

obtuso, basi acuta vel subobtusa, supra glabra densissime pallido-

puncticulata, nervis venisque non elevatis, subtus fere concolor, in

statu juvenili sparse minute stellato-furfuracea, cito glabrata et

in statu adulto fere omnino glabra, nervis primariis teneris promi-

nentibus, venis transversis obscuris; flores axillares solitarii crasse

6-8 mm. longe pedicellati, involucre basi acutiusculo; bracteae

exteriores alte connatae extus sparse minute stellato-furfuraceae

vel fere glabrae 1.5-2 cm. latae fere 2.5 cm. longae, parte libera

ovato-rotundata breviter acuminata, bracteis interioribus omnino

coalitis; calyx ca. 16 mm. longus et apice 2.2 cm. latus, lobis brevis-

STUDIES OF AMERICAN PLANTS — X 95

simis latioribus quam longis apiculatis extus minute stellato-fur-

furaceis; petala ut videtur rosea 3 cm. longa latissima apice truncata

vel late rotundata; antherae late oblongae 5-6 mm. longae, filamentis

crassis aequilongis. — Guatemala: Bank of Rio Dulce, Dept. Izabal,

March 28, 1939, C. L. Wilson 402 (type in Herb. Field Mus.);

February 6, 1939, Wilson 161.

Doubtless this is a showy and handsome tree, because of its

abundance of large flowers. The foliage appears glabrous to the

naked eye, and is almost so in fact when the leaves are fully

developed.

Centronia grandiflora Standl., sp. nov. — Arbor 6-metralis

et ultra, ramis obtuse tetragonis dense breviter setoso-pilosis brun-

neis, internodiis elongatis; folia magna longipetiolata membranacea,

petiolo 3.5-7.5 cm. longo dense brunneo-setoso-piloso; lamina sub-

integra elliptica vel oblongo-elliptica ut videtur 16-25 cm. longa,

7-10 cm. lata, sensim longiattenuata, basi obtusa vel anguste ro-

tundata et breviter emarginata, supra in sicco cinereo-viridis glabra

minute puncticulata, nervis venisque manifestis, ut videtur in vivo

bullata, subtus brunnescens sat dense minute stellato-pubescens et

setoso-pilosa, serius glabrata, 7-plinervia, venis elevatis arete reticu-

latis; inflorescentia terminalis pauciflora 5 cm. longe pedunculata,

pedicellis crassis ca. 7 mm. longis dense ferrugineo-pilosis; calyx

in alabastro 1.5 cm. longus 1 cm. latus densissime pilis longis brun-

neis intertextis indutus, apice breviter mammillatus; petala glabra

late obovata 2 cm. longa apice late rotundata. — Panama: In rain

forest, Bajo Chorro, Prov. Chiriqui, alt. 2,100 meters, March 2,

1938, Mrs. M. E. Davidson 360 (type in Herb. Field Mus.).

The other Central American species of the genus, Centronia

phlomoides Triana of Costa Rica, has a calyx only 11-12 mm. long,

and petals half as large as those of C. grandiflora.

CLETHRACEAE

Clethra obliquinervia Standl., sp. nov. — Ramuli crassi dense

stellato-pilosi et tomentelli, internodiis brevissimis; folia mediocria

petiolata coriacea in sicco brunnescentia, petiolo crassiusculo 1.5-2

cm. longo dense breviter brunneo-piloso; lamina oblanceolato-

oblonga 8-10 cm. longa 2.5-4 cm. lata apice rotundata, basin versus

longe sensim attenuata, basi ipsa cuneata vel subobtusa, supra

sublucida, fere glabra sed pilis paucis minutis stellatis remotis con-

spersa, costa nervisque impressis, subtus albida ubique tomento

minuto stellate densissime obtecta, nervis lateralibus obliquis

96 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

utroque latere ca. 12 teneris prominentibus angulo semirecto

adscendentibus, venis obscuris; racemi fasciculati vel breviter panicu-

lati numerosi usque 11 cm. longi breviter pedunculati laxe multiflori,

rhachi dense breviter brunneo-stellato-pilosa, pedicellis omnibus

brevibus vix ad 3 mm. longis, bracteis lineari-subulatis patentibus

calycem aequantibus; calyx dense tomentellus 3.5 mm. longus,

lobis oblongo-ovatis obtusis; petala calyce bene longiora glabra

ciliata; ovarium dense adpresso-pilosum. — Mexico: Buena Vista,

Escuintla, Chiapas, January, 1938, E. Matuda 1894 (type in Herb.

Field Mus.).

LOGANIACEAE

Buddleia Matudae Standl., sp. nov. — Ramuli obtuse tetragoni

dense brunneo-stellato-tomentelli, internodiis superioribus brevibus;

folia modica petiolata firme membranacea, petiolo crassiusculo

1.5-3 cm. longo dense stellato-tomentello; lamina oblongo-ovata

vel lanceolato-oblonga 8-13 cm. longa 2.5-6 cm. lata sensim vel

subito longiacuminata, basin versus sensim angustata vel interdum

cuneato-acuta, supra in sicco fusca sparse pilis minutis stellatis

conspersa, subtus sordide brunnescens densissime tomento stellato

sublaxo obtecta, fere integra vel inconspicue crenulata, costa subtus

prominente, nervis lateralibus utroque latere ca. 11 obliquis promi-

nentibus prope marginem arcuato-conjunctis; flores parvi in capitula

globosa densissime multiflora 4-7 mm. lata breviter pedunculata

dispositi, capitulis racemosis, racemis paniculatis, paniculis late

pyramidalibus ca. 10 cm. longis atque 15 cm. latis breviter peduncu-

latis, ramis dense stellato-tomentosis; calyx fere 2 mm. longus extus

laxe stellato-tomentellus, lobis oblongis vel ovato-oblongis erectis

obtusis sinubus latis sejunctis tubo fere duplo brevioribus; corolla

extus sparse stellato-tomentella vel fere glabra, calycis tubo lobis

vix longiore, limbo ca. 3 mm. lato, lobis brevibus late ovalibus vel

subrotundatis apice late rotundatis intus glabris.— Mexico: Mount

Ovando, Chiapas, December, 1936, E. Matuda S-085 (type in

Herb. Field Mus.). Mount Pasitar, Chiapas, December 30, 1936,

Matuda S-36.

GENTIANACEAE

In 1932 the writer described (Rhodora 34: 176) a form oiEustoma

Russellianum having pure white corollas as E. Russellianum f.

Fisheri, named for the collector, Mr. George L. Fisher. Later in

the same year Mr. Fisher made further collections to show the color

variations in this showy plant, but the material has lain in the

herbarium without further study until the present time. Two

STUDIES OF AMERICAN PLANTS — X 97

other color variants seem worthy of nomenclatural recognition, and

may be named as follows:

Eustoma Russellianum (Hook.) Griseb. f. bicolor Standl., f.

nov. — Corolla alba, lobis basi purpureo tinctis. — Texas: Willis,

July 4, 1932, C. L. Brock 32114 (type in Herb. Field Mus.).

Mr. Fisher states that this is an easily recognizable form, about

as plentiful as the following one. The three grow together on the

farm of Mr. L. D. Cluxton near Willis, in such abundance that

he sells large quantities of the flowers each year to florists, especially

of the albino form, f . Fisheri. In forma bicolor the corolla is white

except for some purple coloring at the base of the lobes.

Eustoma Russellianum f. roseum Standl., f. nov. — Corolla

rosea, lobis basi sordide rubris.— Texas: Willis, July 5, 1932, George L.

Fisher 32116 (type in Herb. Field Mus.).

In this form the corolla is pink or rose-pink, the lobes dark brick-

red at the base. The collector states that it is locally plentiful with

the type.

EBENACEAE

Diospyros Delgadoi Standl., sp. nov. — Arbor ut videtur dense

ramosa, ramulis patentibus nodosis dense puberulis, internodiis

brevibus; folia inter minora brevissime petiolata coriacea, petiolo

crassiusculo ca. 4 mm. longo; lamina ovalis vel oblongo-elliptica

3-7.5 cm. longa 2-3.5 cm. lata obtusa vel saepe rotundata, basi

obtusa, supra in sicco cinerea glabrata, tantum ad nervos sparse

minute puberula, costa nervisque prominulis, subtus brunnescens

laxe pilis flaccidis intertextis tomentosa, costa gracili prominente,

nervis lateralibus utroque latere 3-5 arcuatis teneris prominentibus

angulo latiusculo adscendentibus, venis obscuris; calyx in statu

fructifero patens ad medium trilobus utrinque sat dense piloso-

tomentosus 1.5-2 cm. latus, lobis latissimis acutis vel subacutis;

fructus globosus 1.5-2 cm. diam. in maturitate glaber et ad apicem

breviter mammillatus sericeus; semina ca. 5 plerumque 1 cm. longa

atque 5 mm. lata. — Venezuela: Boundary between Lara and Tru-

jillo, Trentina, November, 1937, Esteban Delgado 96 (type in Herb.

Field Mus.).

The species is referable to the genus Maba, as treated by Hiern

and other authors, but Maba seems better referable to Diospyros,

where it has been placed by various recent writers.

Diospyros Mexiae Standl., sp. nov. — Arbor 5-metralis gracilis,

ramulis crassiusculis in sicco olivaceis substriatis glabratis, inter-

98 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

nodiis brevibus; folia modica breviter petiolata coriacea in sicco

olivacea, petiolo crassiusculo 1-1.5 cm. longo; lamina oblongo-

lanceolata vel anguste elliptico-oblonga 9-14 cm. longa 3-5 cm.

lata acuta vel acuminata, basi acuta, supra lucida glabra vel tantum

ad costam sparse puberula, costa impressa, nervis subplanis, subtus

ut videtur primo brunneo-tomentella sed cito glabrata, costa gracili

elevata, nervis lateralibus utroque latere ca. 10 tenerrimis aliquanto

irregularibus prominulis angulo latiusculo adscendentibus rectis vel

arcuatis obliquis; flores axillares, pedicellis fructiferis incrassatis

vix ultra 3 mm. longis; calyx 3-(vel 4- ?)lobus 2.5 cm. latus vel

ultra, profunde lobatus, extus brunneo-tomentellus vel serius gla-

bratus, intus dense sericeus, lobis late deltoideo-ovatis acutis vel

acuminatis paten tibus; fructus subglobosus vel paullo altior quam

latus 2 cm. longus vel ultra apice late rotundatus et mammillatus,

ut videtur primo sat dense brunneo-sericeus sed cito glabratus.—

Brazil: About 5 km. north of Fazenda da Gramma, on westward

face of mountain slope, matta virgem, forming part of understory

in dense shade, Districto Carangola, State of Minas Geraes, alt.

920 meters, February 4, 1930, Ynes Mexia 4316 (type in Herb.

Field Mus.).

CONVOLVULACEAE

Ipomoea Davidsoniae Standl., sp. nov. — Volubilis herbacea,

caule gracili dense pilis longiusculis pallidis patentibus piloso, inter-

nodiis elongatis; folia modica longissime petiolata membranacea,

petiolo gracillimo 5-10 cm. longo dense breviter piloso; lamina late

deltoideo-ovato-cordata 5-7.5 cm. longa 3.5-7 cm. lata acuminata

vel cuspidato-acuminata, basi profunde lateque cordata, lobis

posticis late rotundatis, integra vel subundulata, utrinque dense

pilis longiusculis mollibus patentibus pallidis pilosa; inflorescentia

perfecta non visa, floribus delapsis ca. 3 mm. longe pedicellatis;

sepala subcoriacea subaequalia adpressa ovalia vel elliptica ca. 12

mm. longa apice late obtusa et minute mucronata dense pilis longis

pallidis patentibus pilosa; corolla pallide purpurea ca. 4 cm. longa

glabra. — Panama: Bajo Mono, Boquete District, Province of

Chiriqui, alt. 1,350 meters, April 3, 1938, Mrs. M. E. Davidson 595

(type in Herb. Field Mus.).

The available material is not well preserved, but an apparently

distinct and undescribed species is represented, noteworthy locally

for the abundant pubescence of the foliage, and especially for the

densely long-pilose sepals.

STUDIES OF AMERICAN PLANTS— X 99

Ipomoea quesadana Standl., sp. nov. — Herba volubilis omnino

glabra, caule gracillimo, internodiis plerumque elongatis; folia

inter minora longipetiolata membranacea, petiolo gracillimo 2.5-6

cm. longo; lamina ambitu late triangulari-ovata plerumque 4-6 cm.

longa et 3-6 cm. lata, typice breviter vel subprofunde triloba, lobo

medio triangulari vel ovato acuminate, lobis lateralibus medio

multo brevioribus, interdum valde reductis vel nullis, lamina basi

leviter lateque cordata; lamina foliorum superiorum saepe vulgo

reducta et 1-3 cm. tantum longa, rotundo-ovata et interdum apice

rotundata, breviter petiolata; flores parvi, pedunculis axillaribus

crassiusculis 5-18 mm. longis 1-3-floris interdum granulosis, pedi-

cellis crassis usque 5 mm. longis; sepala subaequalia, exterioribus

interdum paullo brevioribus, subcoriacea glabra, apice rotundata

vel subtruncata et saepe brevissime mucronata; corolla rosea glabra

ca. 2 cm. longa. — Costa Rica: Villa Quesada, Canton de San Carlos,

Prov. Alajuela, alt. 850 meters, February 21, 1939, Austin Smith

H1609 (type in Herb. Field Mus.).

"Growing in rain forest, Caribbean watershed; small morning-

glories, rather fragile, the corolla deep begonia rose in the throat,

light pink on the limb, 2.5 cm. long and 2 cm. wide, open at midday."

VERBENACEAE

Clerodendron Moldenkeanum Standl., sp. nov. — Frutex vel

arbor ut videtur inermis, ramis crassiusculis fusco-ochraceis, inter-

nodiis elongatis, ramulis glabris vel ad nodos sparse pilosis, cica-

tricibus foliorum delapsorum plus minusve incrassatis; folia bene

evoluta non visa, juvenilia in sicco fusca modica breviter petiolata,

petiolo crassiusculo ad 7 mm. longo glabro vel sparse piloso; lamina

ovata vel lanceolato-ovata 4.5-6 cm. longa 1.5-2.5 cm. lata acumi-

nata, basi acuta vel obtusa, utrinque puncticulata, supra glabra,

costa nervisque subimpressis, subtus ad costam nervosque sparse

breviter setuloso-pilosa, cito glabrata, costa gracili elevata, nervis

lateralibus utroque latere ca. 5 angulo semirecto adscendentibus

arcuatis prominentibus; flores in cymulas paucifloras axillares

sessiles aggregati, pedicellis gracilibus 1.5-2 cm. longis glabris; calyx

glaber, tubo campanulato basi acuto 4 mm. longo, lobis lineari-

subulatis paullo inaequalibus 5-6 mm. longis erectis glabris; corolla

in alabastro 12 mm. longa obtusa extus glabra, tubo limbo fere

duplo longiore fauce 2 mm. lato. — Mexico: Volcan de Tacana,

Chiapas, alt. 1,400 meters, March, 1939, E. Matuda 2760 (type in

Herb. Field Mus.).

100 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

This species, evidently distinct from the few known previously

from Mexico and Central America, is named for Dr. Harold N.

Moldenke, monographer of this and other genera of the Verbenaceae.

SOLANACEAE

Lycianthes amphidoxys Standl., sp. nov. — Frutex scandens,

trunco 2 cm. crasso, caulibus usque 5 m. longis ramosis glabris,

internodiis brevibus vel aliquanto elongatis; folia inter minora

breviter petiolata crasse membranacea, paris valde inaequalia,

petiolo 3-8 mm. longo sparse pilosulo vel fere glabro; lamina foli-

orum majorum lanceolato-oblonga 5-10 cm. longa 1.5-3.5 cm. lata

longissime attenuato-acuminata, basi plus minusve inaequali acuta,

supra in sicco olivacea sparse breviter scabro-pilosa, subtus fere

concolor ad costam nervosque incurvo-puberula vel fere glabra,

costa gracili elevata, nervis lateralibus utroque latere ca. 6 teneris

arcuatis prominentibus angulo semirecto vel latiore adscendentibus;

lamina foliorum minorum rotundato-ovata 1.5-3 cm. longa obtusa

vel rotundata, basi rotundata; flores axillares solitarii vel saepius

paucifasciculati, pedicellis gracillimis 1.5-2.5 cm. longis glabris;

calyx glaber latissime campanulatus ca. 4 mm. altus atque 5 mm.

latus truncatus, paullo infra marginem appendicibus 5 viridibus

incrassatis obtusis erectis ca. 2 mm. longis onustus; corolla lutea

ca. 8 mm. longa extus glabra; bacca globosa 8 mm. diam. — Ecuador:

Western Cordillera, forest region above Balsapampa, east of Baba-

hoyo, alt. 2,600 meters, July, A. Rimbach 840 (Yale School For.

34191; type in Herb. Field Mus.).

"Flowers 1.5 cm. long, yellow; berry at first whitish, finally red.

Wood white. Bark 1 mm. thick, brownish gray, smooth but some-

what fissured, with elongate lenticels."

Lycianthes chiriquensis Standl., sp. nov. — Arbor ut dicitur

9-12-metralis, fere glabra, ramis crassiusculis ochraceis rimosis,

internodiis brevibus glabris; folia subaequalia petiolata membrana-

cea, petiolis gracilibus glabris 1-2 cm. longis; lamina inaequilateralis

oblongo-elliptica vel oblongo-ovata, rarius late ovata, 5.5-12 cm.

longa 2.5-5.5 cm. lata subabrupte acuminata vel longe attenuato-

acuminata, basi solemniter inaequali acuta vel obtusa, supra in

sicco glabra olivacea, subtus concolor, in axillis nervorum dense

stellato-pilosa, aliter glabra, nervis lateralibus utroque latere ca. 5

arcuatis angulo latiusculo adscendentibus; flores axillares plerum-

que paucifasciculati, pedicellis fere filiformibus vulgo 10-14 mm.

longis, interdum brevioribus, glabris; calyx campanulatus truncatus

STUDIES OF AMERICAN PLANTS — X 101

basi obtusus 3.5 mm. longus et aequilatus; corolla pallide purpurea

extus glabra 8 mm. longa ad basin lobata, segmentis late patenti-

bus lineari-oblongis acutiusculis intus glabris; antherae fere 6 mm.

longae obtusae. — Panama: In forest, Boquete, Prov. Chiriqui, alt.

1,140 meters, May 12, 1938, Mrs. M. E. Davidson 623 (type in Herb.

Field Mus.).

The flowers are described as lavender. The size indicated by

the collector is extraordinary for this genus and family, and may

be erroneous.

Lycianthes luisana Standl., sp. nov. — Herba 30 cm. alta et

ultra ut videtur sparse ramosa, caule crassiusculo villosulo vel

incurvo-pilosulo, internodiis plerumque elongatis; folia majuscula

petiolata membranacea, paris insigniter inaequalia, petiolo gracili

usque 4 cm. longo sed vulgo multo breviore; lamina foliorum ma-

jorum elliptica vel late ovata 10-22 cm. longa 4-9.5 cm. lata breviter

vel longe acuminata, basi plus minusve abrupte et saepe inaequaliter

acuta vel late cuneata, supra in sicco viridis pallido-puncticulata

glabra, subtus concolor puncticulata ad costam nervosque sparse

incurvo-puberula vel fere omnino glabra, nervis lateralibus utroque

latere ca. 6 arcuatis teneris angulo latiusculo adscendentibus; flores

axillares solitarii, pedicellis gracilibus 1.2-3 cm. longis glabris;

calyx glaber turbinato-campanulatus ca. 4 mm. longus et 5 mm.

latus truncatus paullo infra marginem appendicibus 10 viridibus

plus minusve incrassatis 3-4 mm. longis onustus; corolla rotata fere

1 cm. longa glabra patens ut videtur viridescens; antherae 6 mm.

longae obtusae. — Mexico: Tamazunchale, San Luis Potosi, July,

1937, Mary Taylor Edwards 913 (type in Herb. Field Mus.), 520,

535, 496.

The berries are said to be bright red when ripe. The collector

reports the following rather numerous local names for the plant:

Chilequaqua, Flor de mariposa, Tomatillo, Arrete de la Virgen.

Such a large number of distinct names from a single locality leads one

to suspect that they were manufactured for the occasion by the

informant or informants.

Lycianthes obliquifolia Standl., sp. nov. — Frutex ut videtur

scandens ramosus, ramis subgeniculatis crassiusculis brunneo-

ochraceis sparse breviter villosis, internodiis brevibus, nodis plus

minusve incrassatis; folia inter minora breviter petiolata omnia

subaequalia membranacea, petiolo 5-9 mm. longo sparse patenti-

villoso; lamina lanceolato-ovata 5.5-10 cm. longa 2-3.5 cm. lata

102 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

solemniter inaequilatera longe angusteque acuminata, basi oblique

acuta usque subrotundata, supra in sicco olivacea sparse breviter

villosula, subtus concolor sparse breviter villosa, nervis lateralibus

utroque latere ca. 4-5 subarcuatis angulo semirecto adscendentibus

obliquis; flores axillares solitarii, pedicellis saltern fructiferis gracili-

bus rectis 4-5 cm. longis sparse villosis sub calycem paullo incrassatis;

calyx latissime campanulatus ca. 4 mm. altus atque 5-6 mm. latus

truncatus sparse subadpresse villosulus, paullo infra marginem

appendicibus 10 viridibus incrassatis erectis 3-4 mm. longis onustus;

bacca subglobosa ca. 6 mm. lata. — Mexico: Volcan de Tacana,

western slope, alt. 2,800 meters, March 30, 1939, E. Matuda 2938

(type in Herb. Field Mus.).

Physalis sonorensis Standl., sp. nov. — Herba multiramosa ut

videtur fragilis, ramis dichotomis breviter dense viscido-pilosis,

internodiis foliis brevioribus subteretibus interdum tantum puber-

ulis; folia parva longissime petiolata herbacea, petiolo gracillimo

vulgo 1.5-2.5 cm. longo breviter viscido-piloso; lamina late ovata,

elliptico-ovata vel rotundato-ovata 1.3-2.5 cm. longa 1-2 cm. lata

obtusa vel subacuta, basi late rotundata vel obtusa, interdum late

cuneata vel abrupte breviterque contracta, grosse sinuato-dentata

vel undulato-dentata, basin versus Integra, utrinque sparse vel

subdense breviter viscido-villosula; pedicelli solitarii gracillimi ad

anthesin erecti ca. 2 cm. longi dense viscido-villosuli; calyx

ad anthesin campanulatus 5-6 mm. longus medio 3.5 mm. latus sub-

dense breviter viscido-villosulus, dentibus erectis oblongo-ovatis

tubo paullo brevioribus acutis; corolla pallide lutea 1 cm. longa

extus sparse viscido-villosula, limbo breviter lobato, lobis latissimis

obtusis. — Mexico: Bachoco, 12 miles east of Cajeme, Sonora, in

rocks on malpais hill, Lower Sonoran Zone, in thorn forest, February

20, 1937, Howard Scott Gentry 3011 (type in Herb. Field Mus.).

The species is remarkable for the greatly elongate pedicels and

petioles.

Solandra brevicalyx Standl., sp. nov. — Ramuli crassi ochracei

rugosi; folia modica petiolata papyracea, petiolo 1.5-2.5 cm. longo

sparse laxeque stellato-tomentoso; lamina obovata vel elliptico-

obovata 6-9.5 cm. longa 3.5-5.5 cm. lata, apice obtusa vel rotundata

et acuminato-apiculata, basi inaequali acuta, supra sparse pilis

laxis parvis ramosis conspersa, subtus densiuscule stellato-tomentosa,

nervis lateralibus utroque latere ca. 6 angulo lato adscendentibus

teneris subarcuatis prominulis; flores crasse ca. 7 mm. longe pedi-

cellati magni; calyx ca. 3 cm. longus et 1 cm. latus, sparse laxeque

STUDIES OF AMERICAN PLANTS — X 103

stellato-villosus, lobis lanceolatis vel ovatis acutis vel acuminatis

tubo duplo brevioribus; corolla glabra 10-17 cm. longa, limbo 6 cm.

lato vel ultra, tubo inferne gracillimo supra calycem 3 mm. lato,

sursum sensim dilatato. — Mexico: Region of Jaumave, Tamaulipas,

in 1932, H. W. von Rozynski 394 (type in Herb. Field Mus.).

Solandra brevicalyx is probably the northernmost member of

this small genus. From other related species it is well distinguished

by the short calyx.

Solatium boquetense Standl., sp. nov. — Herba vel frutex

metralis, ramis crassis dense aculeatis, densissime pilis magnis laxis

fulvis inaequalibus longissime stipitatis stellatis indutis, aculeis

rectis patentibus gracilibus plus minusve compressis usque 5 mm.

longis fusco-ochraceis; folia inaequalia magna petiolata herbacea,

petiolo crasso foliorum ma jorum usque 9 cm. longo aculeato et dense

pilis stipitatis tomentoso; lamina late ovali-ovata vel late elliptica

usque 26 cm. longa atque 20 cm. lata vel multo minor, apice obtusa

vel subacuta, basi lata breviter cordata usque subacuta, brevissime

angulato-lobata vel subintegra et tantum undulata, utrinque ad

costam nervosque aculeis paucis crassis remotis armata, supra

densissime pilis fulvidis sat longis sessilibus vel breviter stipitatis

intertextis induta, radio centrali pili aliis multo longiore, subtus

pallidior densissime molliter pilis stellatis plerumque stipitatis

tomentosa; flores dense in racemos breves multifloros aggregati

breviter crasse pedicellati extus in omnibus partibus pilis magnis

fulvis longistipitatis sat mollibus stellatim ramosis dense obtecti;

calyx 7-8 mm. longus, tubo late campanulato basi obtuso, lobis

erectis tubo aequilongis obtusis; corolla alba 1 cm. longa vel ultra

fere ad basin lobata, segmentis lanceolato-oblongis acuminato-

apiculatis intus glabris; fructus immaturus globosus densissime pilis

longissimis plurilocularibus hispidus. — Panama: Boquete, Prov.

Chiriqui, alt. 1,140 meters, June 28, 1938, Mrs. M. E. Davidson 817

(type in Herb. Field Mus.).

The plant is perhaps a relative of S. hirtum Vahl, one of the

common, weedy species of Central America.

Solatium Davidsoniae Standl., sp. nov. — Frutex inermis

3-4.5 m. altus ubique glaber, ut videtur sat ramosus, ramis nodosis

sordide ochraceis, internodiis brevibus; folia inter minora breviter

petiolata membranacea paris inaequalia, petiolo crassiusculo 5-9

mm. longo; lamina in foliis majoribus oblique ovata vel oblongo-

ovata 4.5-8 cm. longa 2-4 cm. lata acuta vel subito acuminata,

104 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

basi inaequali acuta vel abrupte contracta, Integra, supra in sicco

viridis, subtus paullo pallidior; lamina foliorum minorum paris

rotundato-ovata vel rotundo-elliptica 1-3 cm. longa et saepe fere

aequilata, apice obtusa vel rotundata, basi obtusa vel rotundata;

flores ut videtur solitarii axillares, pedicellis 1.5 cm. longis gracilibus

sursum paullo incrassatis; calyx 4 mm. longus et aequilatus basi

obtusus, margine breviter obtuse dentato; corolla pallide lutea 12

mm. longa profunde lobata, lobis elliptico-ovatis obtusis. — Panama:

In rain forest, Bajo Chorro, Boquete District, Prov. Chiriqui, alt.

2,100 meters, March 2,1938, Mrs. M. E. Davidson 362 (type in Herb.

Field Mus.).

Solatium pluviale Standl., sp. nov. — Frutex 2-4.5 m. altus

aculeatus, ramis crassis fere glabris sed pilis paucis parvis remotis

sessilibus et stipitatis stellatis conspersis, internodiis elongatis,

aculeis paucis patentibus usque 8 mm. longis compressis basi valde

dilatatis fusco-brunneis; folia magna petiolata herbacea subaequalia,

petiolo usque 7.5 cm. longo aculeis paucis magnis armato sparse

pilis breviter stipitatis parvis rigidis consperso; lamina 16-28 cm.

longa et 15-25 cm. lata ad medium vel brevius pinnato-lobata utrin-

que ad costam nervosque aculeis paucis patentibus rectis magnis

armata, supra aspera sat dense pilis minutis rigidis pauciramosis

fulvidis sessilibus conspersa, subtus fere concolor similiter sed densius

induta, lobis laminae lanceolato-oblongis vel interdum deltoideis

acutis vel acuminatis patentibus vel obliquis; flores in cymas multi-

floras densas pedunculatas dispositi pedicellati, pedicellis dense

rigide pilis brevibus fulvidis eglandulosis indutis; calyx dense stellato-

hispidulus, tubo late campanulato ca. 3 mm. longo et paullo latiore

5-lobato, lobis remotis sinubus late rotundatis segregatis oblongo-

linearibus erectis tubo aequilongis; corolla alba ca. 1 cm. longa extus

densissime pilis parvis rigidis stellatis sessilibus conspersa, fere ad

basin 5-lobata, segmentis lanceolato-oblongis acutiusculis intus

glabris; antherae angustae ca. 6.5 mm. longae sursum sensim attenu-

atae. — Panama: In rain forest, Bajo Chorro, Boquete District,

Prov. Chiriqui, alt. 1,800 meters, January 24, 1938, Mrs. M. E.

Davidson 189 (type in Herb. Field Mus.).

Noteworthy for the stout, glabrate branches, and for the large,

deeply pinnatifid, green, very rough leaves.

Solatium salsum Standl., sp. nov. — Frutex aculeatus ca.

metralis, ut videtur dense ramosus, ramis crassiusculis teretibus

densissime pilis stellatis sessilibus vel breviter stipitatis brunnes-

centibus indutis, internodiis brevibus, aculeis sat numerosis vel

STUDIES OF AMERICAN PLANTS — X 105

interdum paucis rectis patentibus lateraliter subcompressis 5-6 mm.

longis basi valde dilatatis; folia inter minora petiolata herbacea,

petiolo 6-17 mm. longo dense stellato-tomentoso; lamina ovata,

oblongo-ovata vel oblongo-ovalis vulgo 3.5-7 cm. longa et 2-4 cm.

lata apice obtusa vel rotundata, basi rotundata vel late obtusa et

vulgo inaequalis, uno latere breviore, integra vel subundulata, supra

dense pilis stellatis sessilibus rigidis conspersa, costa venisque im-

pressis, subtus paullo pallidior brunneo-ochracea densissime pilis

intertextis sessilibus stellatis tomentosa, costa crassiuscula promi-

nente, nervis lateralibus utroque latere ca. 6 angulo semirecto ad-

scendentibus; inflorescentiae umbelliformes vel breviter racemosae

plerumque 2-7-florae vix ultra 6 mm. longe pedunculatae, pedicellis

gracilibus vulgo 1-2.5 cm. longis pilis stipitatis stellatis dense in-

dutis; calyx ca. 6 mm. longus extus dense stellato-tomentosus, pilis

sessilibus vel substipitatis, fere ad basin lobatus, segmentis lanceo-

lato-oblongis acutiusculis in statu fructifero paullo accrescentibus

et usque 9 mm. longis; corolla alba ca. 8 mm. longa extus dense

stellato-tomentosa stellatim patens, segmentis fere lineari-lanceo-

latis attenuato-acutis; antherae fere 5 mm. longae sursum paullo

attenuatae; bacca globosa glabra apice rotundata 8-9 mm. diam.—

Mexico: Salina Cruz, Oaxaca, on rocky slope above the seashore,

June, 1937, Llewelyn Williams 9706 (type in Herb. Field Mus.);

edge of road in low forest, Williams 9686; side of road, in sandy

soil near the seashore, Williams 9703.

SCROPHULARIACEAE

Buchnera leiantha Standl., sp. nov. — Herba simplex erecta

stricta 25-30 cm. alta, caule gracili sed rigido bifariam incurvo-

puberulo, internodiis foliis duplo brevioribus; folia numerosa an-

guste linearia vel fere subulata erecta vel adpressa sessilia plerumque

1.5-3 cm. longa ad margines arete revolutos scabrella obtusa 1-

nervia; flores spicati, spicis dense paucifloris 2.5-4 cm. longis, bracteis

lanceolatis cymbiformibus acuminatis ciliatis dimidium calycis

aequantibus arete adpressis; calyx glaber 8 mm. longus subtubulosus

sursum paullo dilatatus, dentibus anguste triangularibus subulato-

acuminatis erectis 1.5 mm. longis; corolla ut dicitur alba vel pur-

purascens ca. 1 cm. longa extus glabra, lobis rotundato-ellipticis 3

mm. longis. — Panama: Boquete, Prov. Chiriqui, alt. 1,350 meters,

an herb 10-25 cm. high, January 11, 1939, Mrs. M. E. Terry 1273

(type in Herb. Field Mus.).

From other Buchnera material that I have seen from Central

America, this is readily separable by the glabrous flowers.

106 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

Castilleja bella Standl., sp. nov. — Herba nana perennis dense

caespitosa 6-10 cm. alta, caulibus abbreviatis simplicibus saepe

fere nullis foliis basalibus occultis, sparse laxeque longipilosis vel

villosis; folia radicalia numerosissima 3-6.5 cm. longa basi 4-7 mm.

lata linearia longissime attenuata, saepissime falcata et incurva

vel subtorta, sparse laxe pilosa vel fere glabra ca. 6-nervia; racemi

laxe pauciflori, rhachi laxe albido-villosa; bracteae ca. 2 cm. longae

supra basin purpureo-rubrae, plerumque fere ad medium trilobae,

lobo medio oblongo obtusissimo, interdum utroque latere breviter

lobulato, lobis lateralibus fere aequilongis linearibus obtusis, extus

praesertim ad nervos longivillosae vel puberulae; calyx fere 2.5 cm.

longus vix apice 4 mm. latus dense albo-villosus bilobus, lobis

5 mm. longis obtusissimis purpureo-rubris dense minute puberulis;

corolla calyce paullo longior 2.5 cm. longa vel ultra, tubo hyalino

glabro, galea 1 cm. longa recta viridi superne dorso dense breviter

pilosula rubro-marginata, labio inferiore viridi vix 1.5 mm. longo. —

Mexico: State of Nuevo Leon, abundant in meadows above and

below timber line, peak of Cerro Potosi, Municipio de Galeana,

July 21, 1935, C. H. Mueller 2249 (type in Herb. Field Mus.).

Abundant on rocky alpine meadows of peak of Cerro Potosi, alt.

3,750 meters, July 18, 1938, Richard A. Schneider 935.

Mr. Schneider has supplied the following notes: "Bracts brilliant

red. A very conspicuous plant, abundant on the treeless peak,

and extending down (equally abundant) into the pine forest and

in grassy openings for over 300 meters. In exposed places only

2.5-5 cm. high. In the shelter of the timber at times reaching a

height of 12 cm., but even in such places most of the plants are low.

One yellow-flowered individual was collected." Probably a local,

alpine species, noteworthy for its low, dense habit, the stems bearing

flowers almost or quite to their bases, which are hidden by the very

numerous long, twisted and curled basal leaves.

GESNERIACEAE

Columnea Muenscheri Standl., sp. nov. — Frutex epiphyticus,

ramo unico viso ca. 30 cm. longo crasso cortice ferrugineo obtecto,

internodiis brevibus 1-1.5 cm. tan turn longis; folia subdensa breviter

petiolata subcoriacea, petiolo crassiusculo 3-5 mm. longo; lamina

lanceolato-linearis 5.5-7.5 cm. longa 5-8 mm. lata longe attenuato-

acuminata, basin versus sensim attenuata, 1-nervia, supra in sicco

cinereo-viridis glabra, costa impressa, subtus subsparse pilis longis

laxis sericea, costa gracili elevata rubescente; flores axillares solitarii,

bracteis lanceolato-ovatis 6-8 mm. longis acuminatis laxe sericeis,

STUDIES OF AMERICAN PLANTS — X 107

pedicello ca. 1 cm. longo laxe adpresso-piloso; calyx 2 cm. longus

extus subsparse laxe adpresso-pilosus fere ad basin lobatus, lobis

lineari-lanceolatis longissime attenuatis intus ad costam breviter

pilosis; corolla laete rubra 6.5 cm. longa extus pilis longis patentibus

multilocularibus rubris villosa, tubo 3.5 cm. longo ad orem 1 cm.

lato, limbo insigniter bilabiate, lobo postico fere 3 cm. longo atque

expanse 2 cm. lato apice sub truncate, labio inferiore profunde

trilobo, lobis e basi lata ovato-deltoideo-acuminatis. — Guatemala:

On pine ridge, Finca Moca, Dept. Suchitepe"quez, alt. 990 meters,

May 4, 1937, W. C. Muenscher 12449 (type in Herb. Field Mus.).

Evidently a handsome and showy plant, marked by the very

large corollas and long, narrow leaves.

Kohleria filisepala Standl., sp. nov.— Herba ut videtur erecta,

caule subgracili dense brunnescenti-villoso, internodiis superioribus

brevibus; folia modica petiolata herbacea, petiolo gracili 2.5-5.5 cm.

longo dense villoso; lamina elliptico-lanceolata vel lanceolato-oblonga

10-14 cm. longa 3-5.5 cm. lata anguste longiacuminata, basi acuta,

utrinque sat dense, subtus paullo densius, pilis longis multilocularibus

villosa, nervis lateralibus obliquis angulo semirecto vel angustiore

adscendentibus fere rectis utroque latere ca. 8; inflorescentiae

axillares cymose ca. 5-florae graciliter 3-7 cm. longe pedunculatae

folia interdum fere aequantes, pedicellis brevibus vix ad 3 mm.

longis; calycis tubus anguste turbinatus ca. 3 mm. longus basi

attenuatus, lobis liberis deciduis anguste linearibus et fere filiformibus

inaequalibus 12-24 mm. longis dense villosis; corolla extus sparse

villosa, tubo ca. 4 cm. longo e basi ad faucem ca. 7 mm. latum sensim

dilatato, lobis ovato-triangularibus patentibus acutis 7-8 mm.

longis intus puberulis; discus lobatus glaber, stylo filiformi 4.5 cm.

longo; antherae lineares 8 mm. longae. — Mexico: Mount Ovando,

Chiapas, December 16, 1936, E. Matuda 574 (type in Herb. Field Mus.).

A remarkable plant, especially noteworthy because of the exceed-

ingly narrow and greatly elongate calyx lobes.

RUBIACEAE

Alibertia dolichophylla Standl., sp. nov. — Arbuscula, ramulis

crassiusculis nodosis ochraceis rimosis sparse puberulis vel glabratis,

internodiis abbreviatis; stipulae 1 cm. longae obtusae; folia majuscula

petiolata pergamentacea, petiolo ca. 2 cm. longo; lamina anguste

oblanceolato-oblonga ca. 21 cm. longa et 5-6 cm. lata acuminata,

basin angustam versus sensim longiattenuata, glabra vel glabrata,

in sicco fusca, supra sublucida, costa gracillima subtus prominente,

108 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

nervis lateralibus utroque latere ca. 9 obliquis angulo latiusculo

adscendentibus teneris; inflorescentia mascula terminalis 1 cm.

longe pedunculata 7-flora cymosa et capituliformis, pedicellis brevis-

simis vel nullis crassis minute subadpresso-pilosulis; hypanthium

cum calyce campanulatum 6 mm. longum 4 mm. latum basi obtusum

minute adpresso-pilosulum, calycis dentibus triangularibus obtusis

1 mm. longis erectis; corolla alba in alabastro 1.5 cm. longa, tubo

3.5 mm. lato dense retrorso-sericeo, lobis tubo aequilongis angustis

obtusis breviter denseque antrorso-sericeis. — Brazil: Juruty Velho,

civ. Para, silva terris altis argillosis ad meridiem lacus, July 29, 1927,

A. Ducke 22916 (type in Herb. Field Mus.).

Among the somewhat numerous Brazilian species of Alibertia,

the present Amazonian one is noteworthy for its relatively narrow

and elongate leaves.

Blepharidium mexicanum Standl., sp. nov. — Arbor 5-6-

metralis fere glabra, ramulis crassis saltern in sicco complanatis et

fuscis glabris, internodiis 4-6 cm. longis; stipulae ferrugineae tenues

lanceolato-triangulares 2.5 cm. longae et basi 12 mm. latae subulato-

attenuatae glabrae; folia magna petiolata membranacea, petiolo

2.5-3.5 cm. longo supra sparse piloso; lamina elliptica 19-26 cm.

longa 9-14 cm. lata acuta vel breviter obtuseque acuminata, basi

obtusa vel rotundata, supra ad costam nervosque sparse pilosa,

aliter glabra, subtus paullo pallidior in axillis nervorum albo-barbata,

sparsissime fere ubique breviter pilosa vel glabrata, costa crassiuscula

elevata, nervis lateralibus utroque latere ca. 13 angulo lato adscen-

dentibus arcuatis pallidis teneris prominentibus, venis laxe reticu-

latis; inflorescentiae axillares solitariae 9-17 cm. longe pedunculatae,

simplices et flores 3 vel plures umbellatos gerentes vel breviter

cymosae vel subracemosae, foliaceo-bracteatae, bracteis superioribus

plerumque elliptico-oblongis et 5-6 mm. longis, inferioribus multo

majoribus, pedicellis interdum usque 2 cm. longis, flore centrali

cymulae sessili; hypanthium obovoideum glabrum 4 mm. longum

basi acutatum; calyx 4 mm. longus et 7 mm. latus glaber, lobis

brevibus latissimis convolutis apice late rotundatis ciliolatis; corolla

alba glabra, tubo ca. 4 cm. longo ad faucem 5 mm. lato sursum vix

dilatato, lobis ovalibus apice rotundatis ca. 1 cm. longis patentibus.—

In advanced forest, "Rvina," Palenque, Chiapas, July, 1939, E.

Matuda 3670 (type in Herb. Field Mus.).

The genus Blepharidium, with a single species, B. guatemalense,

was published by the writer in 1918 (Journ. Wash. Acad. 8: 59),

being based upon a single collection made by H. Pittier along

STUDIES OF AMERICAN PLANTS — X 109

the Saklak River below Secanquim, Alta Verapaz, Guatemala.

Further material of the same plant, a shrub 2.5-5.5 meters high,

was collected by the writer in 1922 on the hills above Quirigua,

Dept. Izabal, Guatemala, and the following recent collections

also are in the Herbarium of Field Museum: Near Quirigua, alt.

about 150 m., April, 1939, Standley 72473, 72273, 72275, 72472.

In the original description the fruit was not described, and no

perfect fruit is now available. It is, however, a septicidal and

partially loculicidal capsule 1.5-2 cm. long, obtuse, acutish at the

base, hard and somewhat woody. The shrub is fairly common on

the hot and rather dry hills overlooking Quirigua, growing among

other scrubby vegetation, such as Curatella, Byrsonima, etc.

It was a surprise to receive material of evidently the same genus

from the State of Chiapas. At first it was assumed that it belonged

to the species described from the north coast of Guatemala, but

closer study shows differences that force one to believe that two

species are represented. As compared with the Chiapas plant, the

Guatemalan one has much larger flowers, the corolla tube being

6 cm. long. Moreover, in B. guatemalense the only leaf pubescence

consists of tufts of hairs in the axils of the primary nerves, while

in B. mexicanum these hair tufts are present and there are also sparse

hairs scattered here and there over all the lower leaf surface.

Borreria Bradei Standl. Field Mus. Bot. 11: 184. 1936.—

When described, this species was based upon a single collection,

Brade 9953 from Therezopolis, Pedra Chapadao, State of Rio de

Janeiro, Brazil. The following collections have been received more

recently: Sta. Magdalena, Alto do Desengano, alt. 2,000 meters,

March, 1934, Santos Lima & Brade 13354 (Herb. Inst. Biol. Veg.

24607). Serra dos Orgaos, alt. 2,000 meters, May, 1931, A. C. Brade

10800 (Herb. Mus. Nac. Rio 24308). Sta. M. Magdalena, Serra da

Furquilha, Rio de Janeiro, alt. 1,700 meters, March, 1935, Santos

Lima & Brade 27682.

Borreria evenia Standl., sp. nov. — Herba ut videtur perennis

erecta(?) ramosa, caulibus crassiusculis fusco-ferrugineis glabris

nodosis 9 cm. longis, internodiis abbreviatis; stipularum setae

incrassatae et herbaceae 2-4 mm. longae; folia parva crasse herbacea

subsessilia anguste lanceolato-oblonga 1.5-2.3 cm. longa 3-7 mm.

lata apicem obtusum vel acutiusculum versus angustata, basi sub-

acuta vel obtusa et contracta, glabra, costa supra anguste impressa,

subtus prominula, nervis venisque obsoletis, subtus minute fusco-

puncticulata; flores parvi dense capitati, capitulis terminalibus

110 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

sessilibus ca. 8 mm. latis basi bracteis paucis foliis conformibus

fulcratis; hypanthium glabrum oblongum; sepala 2 erecta triangulari-

oblonga ca. 0.8 mm. longa subobtusa; corolla glabra 1.5 mm. longa

vel paullo ultra, lobis oblongo-ovatis obtusis tubo brevi multo

longioribus; stamina breviter exserta; fructus cocci glabri angusti

1.8 mm. longi. — Trinidad Island (a possession of Brazil), B. Lobos

4875 (type in Herb. Field Mus., received from the Museu Nacional

of Rio de Janeiro).

Borreria nana Standl., sp. nov. — Herba ut videtur perennis

erecta (?) sparse ramosa, caulibus crassiusculis angulatis glabris

10-12 cm. longis, internodiis foliis longioribus vel brevioribus;

stipularum setae suberectae rigidae viridescentes usque 4 mm.

longae; folia sessilia vel breviter petiolata rigide herbacea oblonga,

oblongo-lanceolata vel ovato-oblonga 2-3 cm. longa 7-12 mm. lata

in sicco laete lutescentia, acuta vel obtusa, basi cuneato-acuta vel

subobtusa, utrinque glabra, costa venisque supra anguste impressis,

costa subtus prominente crassa, nervis lateralibus utroque latere

ca. 3 prominentibus angulo angusto adscendentibus; flores glabri

dense capitati, capitulis terminalibus et axillaribus sessilibus densis-

sime multifloris 1 cm. latis vel paullo ultra, basi bracteis 2 vel pluribus

foliis conformibus fulcratis; hypanthium turbinatum paullo ultra 1

mm. longum; sepala 4 triangulari-oblonga 3-4 mm. longa subae-

qualia viridescentia acuta vel subacuta; corolla ca. 5 mm. longa

caerulea vel purpurea, tubo superne late dilatato, lobis patulis

triangulari-ovatis intus villosis; stamina exserta; fructus cocci 3 mm.

longi septicide dehiscentes; semina nigra 2.5 mm. longa minute

transverse rugulosa utrinque rotundatis, facie interiore anguste

sulcata. — Brazil (State of Sao Paulo): Jabaquara, December, 1914,

A. C. Erode 7056 (Herb. Inst. Biol. 6802; type in Herb. Field Mus.).

Santo Amaro, December, 1906, A. Usteri (Herb. Inst. Biol. 11831).

S. Caetanho, October, 1922, A. Gehrt 8045.

The collections cited were determined by Krause as Borreria

asclepiadea Cham. & Schlecht., which Schumann in Flora Brasili-

ensis reduced to synonymy under B. Poaya (St. Hil.) DC. As

described, B. asclepiadea differs from the Sao Paulo plant in having

flowers about twice as large, and probably Schumann was correct

in his disposition of that name.

Chione chiapasensis Standl., sp. nov. — Arbor 8-metralis

omnino glabra, trunco 35 cm. diam., ramulis in sicco fuscis ad nodos

aliquanto compressis, internodiis solemniter elongatis; stipulae 5-6

STUDIES OF AMERICAN PLANTS — X 111

mm. longae oblongo-ovatae 5-6 mm. longae acutae deciduae; folia

petiolata crasse membranacea vel subcoriacea, petiolo 1-2.3 cm.

longo; lamina late elliptica usque elliptico-oblonga 12-15 cm. longa

4.5-7 cm. lata subito breviterque acuminata, acumine obtuso, basi

acuta usque acuminata, interdum subito acutata, supra in sicco

fusca, subtus pallidior et aliquanto brunnescens, costa gracili promi-

nente, nervis lateralibus utroque latere ca. 7 angulo semirecto ad-

scendentibus tenerrimis subarcuatis, venulis obsoletis; inflorescentia

terminalis cymoso-corymbosa ca. 6 cm. longa et 7 cm. lata 7 cm.

longe pedunculata, laxe subpauciflora, pedicellis fructiferis usque

4 mm. longis; fructus teres 2-locularis ambitu anguste oblongus,

basi et apice acutiusculus 15-18 mm. longus 5 mm. latus lucidus

calyce parvo persistente coronatus; calyx paullo ultra 1 mm. altus

breviter obtuse dentatus. — Mexico: River side in virgin forest,

"Rvina," Palenque, Chiapas, July, 1939, E. Matuda 3675 (type in

Herb. Field Mus.).

The fruits are much larger than those of any other species of the

genus. Those of C. mexicana Standl., the only other species known

from Mexico, are only 1 cm. long.

Chione costaricensis Standl., sp. nov. — Arbor 7-metralis

omnino glabra, ramis gracilibus teretibus ochraceis ad nodos ali-

quanto compressis, internodiis brevibus; stipulae deciduae 4-5 mm.

longae triangulari-ovatae acutae; folia petiolata crasse membrana-

cea, petiolo 1-1.5 cm. longo; lamina elliptico-ovata usque lanceo-

lato-oblonga 6.5-14.5 cm. longa 3.5-7 cm. lata breviter acuminata

vel acutata, acumine obtuso, basi acutiuscula usque rotundata et

subito contracta, supra in sicco fusca, subtus paullo pallidior, costa

prominente, nervis lateralibus utroque latere ca. 9 angulo latiusculo

abeuntibus subarcuatis tenerrimis, venulis inconspicuis laxe reticu-

latis; inflorescentia terminalis cymoso-corymbosa ca. 4 cm. longa

atque 6 cm. lata laxe multiflora 3 cm. longe pedunculata, basi folia-

ceo-bracteata, bracteis minutis, pedicellis crassis usque 14 mm.

longis prope medium bracteatis; hypanthium clavatum 3 mm. Ion-

gum basin versus sensim attenuatum; calyx late campanulatus

5 mm. latus vix ultra 2 mm. altus, margine breviter lobato, lobis

rotundatis; corollae tubus 5-6 mm. longus et 4 mm. latus sursum

paullo dilatatus, lobis 4 rotundato-ellipticis apice rotundatis tubo

fere aequilongis intus glabris; stamina exserta, antheris anguste

oblongis 3.5 mm. longis. — Costa Rica: Villa Quesada, Canton de San

Carlos, Prov. Alajuela, alt. 825 meters, edge of Caribbean rain forest,

March 14, 1939, Austin Smith F1778 (type in Herb. Field Mus.).

112 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

This is the first species of Chione to be reported from Central

America. It is similar to the Mexican C. mexicana Standl., which

differs in having smaller, short-pedicellate flowers and relatively

somewhat narrower leaves.

Chomelia triantha Standl., sp. nov. — Frutex gracilis 2-me-

tralis, ramis subteretibus nigrescentibus nodosis, internodiis abbre-

viatis, novellis ferrugineis hirtellis; stipulae persistentes e basi

triangulari longe subulato-attenuatae 4.5 mm. longae; folia parva

breviter petiolata crasse membranacea, petiolo crassiusculo 3-6 mm.

longo sparse strigilloso; lamina lanceolato-oblonga 4.5-6.5 cm. longa

1.3-2.5 cm. lata acuta vel longiuscule acuminata, basi obtusa vel

acuta, supra viridis lucid a glabra, costa prominente, subtus pallidior

brunnescens, in axillis nervorum sparse breviter albo-barbata, aliter

glabra vel glabrata, costa gracillima prominente, nervis lateralibus

utroque latere ca. 6 angulo angusto adscendentibus tenerrimis

prominulis, venulis obscuris laxe reticulatis; inflorescentiae axillares

solitariae triflorae, pedunculo fere filiformi 1.7-3.5 cm. longo fere

glabro, floribus sessilibus, bracteis filiformibus 4-5 mm. longis;

hypanthium anguste oblongum 2.5 mm. longum dense albido-stri-

gosum, calycis lobis subulatis viridescentibus erectis 3 mm. longis;

corolla alba extus sparse strigosa ca. 22 mm. longa, tubo gracillimo

vix ultra 1 mm. lato, lobis ovato-oblongis patentibus cuspidatis 5 mm.

longis intus glabris, dorso paullo infra apicem appendice subulato

ca. 0.5 mm. longo auctis; stylus breviter exsertus. — Colombia:

Departamento de Boyaca, in thickets, Orocu4, alt. 140 meters, May

12, 1939, Oscar Haugkt 2822 (type in Herb. Field Mus.).

Crusea Gentry! Standl., sp. nov. — Herba erecta annua(?)

gracilis sparse ramosa, 35 cm. alta vel ultra, caule paullo ultra 1 mm.

crasso subacute tetragono glabro, internodiis foliis multo longioribus;

stipularum setae utroque latere ca. 5 subulatae usque 3 mm. longae;

folia parva brevissime petiolata membranacea anguste lanceolata

2-3.5 cm. longa 5-8 mm. lata longe angusteque attenuata, basi

obtusa subito contracta, supra in sicco cinereo-viridia enervia scab-

erula, subtus pallidiora glabra, costa tenui prominente, nervis later-

alibus utroque latere 3-4; capitula densissime multiflora fere 2 cm.

lata longissime pedunculata, basi bracteis pluribus foliis conformibus

et usque 2.5 cm. longis fulcrata; flores tetrameri, hypanthio glabro;

calycis lobi lineari-filiformes 5 mm. longi basi paullo dilatati scaber-

ulo-ciliati patuli; corolla extus minute puberula, tubo gracili 5 mm.

longo sursum subdilatato, lobis 4 lineari-oblongis ca. 4 mm. longis

patentibus acutiusculis intus glabris; stamina longiexserta, fila-

STUDIES OF AMERICAN PLANTS — X 113

mentis tenerrimis, antheris oblongis 1 mm. longis; fructus cocci

o vales 2 mm. longi utrinque obtusi laeves glabri. — Mexico: Quiricoba,

Distrito de Alamos, Sonora, November 12, 1933, Howard Scott

Gentry 766M (type in Herb. Field Mus.).

Diodia Mello-Barretoi Standl., sp. nov. — Herbacea vel suf-

fruticosa ut videtur erecta rigida ramosa stricta 40 cm. alta vel

ultra, caulibus crassis nigrescentibus dense scaberulis, dense foliatis,

internodiis foliis multo brevioribus, nodis plus minusve incrassatis;

stipularum setae 1-5 mm. longae rigidae scaberulae; folia parva

oblique adscendentia sessilia rigide coriacea lanceolata vel lineari-

lanceolata 4-14 mm. longa basi 2-5 mm. lata apice subulato-

mucronata cymbiformia enervia lucida glabra, supra late concava,

subtus acute carinata vel rotundato-convexa, ad margines carti-

lagineos minute scaberula; flores in axillis sessiles solitarii, hypanthio

dense scaberulo, dentibus calycis triangularibus erectis 1 mm. longis

vel ultra acutis vel acuminatis; corolla extus furfuraceo-puberula

ca. 5 mm. longa, tubo superne late dilatato, lobis triangulari-ovatis

tubo duplo brevioribus acutiusculis intus glabris; stamina breviter

exserta, antheris oblongis 1.2 mm. longis; fructus non visus. — Brazil:

Campo, Sierra do Cipo, km. 116, Municipio Santa Luzia, January 13,

1934, H. Mello Barreto 3541 (Herb. Jard. Bot. Bello Horizonte

11483; type in Herb. Field Mus.); February 2, 1938, Mello Barreto

9274 (Herb. Jard. Bot. Bello Horizonte 26167).

The plant differs conspicuously from other species in its strict,

erect, branched habit, the conspicuously nodose stems, and especially

in the small, suberect, rigid, cymbiform leaves, in which the costa

is either conspicuous and keel-like or quite obsolete.

Galiutn Cuatrecasasii Standl., sp. nov. — Herba repens vel

adscendens sat dense ramosa, ramis gracillimis intertextis usque

30 cm. longis vel ultra ad angulos minute scaberulis, internodiis

foliis plus quam duplo longioribus; folia plerumque 6-verticillata

oblonga vel superiora paullo latiora, plerumque 3-5 mm. longa

obtusa vix apiculata basin versus paullo angustata atque fere ses-

silia, 1-nervia, ad costam marginesque scaberulo-ciliata; flores parvi

in cymulas trifloras terminales solitarias vel ternatas dispositi,

cymulis vulgo basi foliatis et foliis fere occultis, pedunculis florum

lateralium folia subaequantibus glabris; corolla tetramera paullo

ultra 1 mm. longa glabra, lobis late ovatis obtusis; fructus baccatus

glaber subglobosus 1.5 mm. longus apice late rotundatus. — Colombia:

Sabana de Bogota, Dept. Cundinamarca, alt. 2,600 meters, December

114 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

29, 1938, Jose Cuatrecasas 468 (type in U. S. Nat. Herb. ; fragment

in Herb. Field Mus.).

It is somewhat surprising to receive from the not exactly un-

explored Bogota region a Galium quite unlike the few other species

known from Colombia. This one, however, does not approach

any of those known from Colombia, nor has it close relatives else-

where in the northern Andes.

Galium juniperinum Standl., sp. nov. — Herbacea vel suf-

fruticosa multiramosa, caulibus rigidis gracilibus pallidis 60 cm.

longis vel ultra dense scaberulis, internodiis plerumque foliis longi-

oribus; folia parva quaternata adscendentiavel subadpressa, interdum

subpatula, subulata rigida 2-4 mm. longa sessilia, apice spinoso-

subulata, 1-nervia glabra, marginibus minute scaberulis; cymulae

axillares solitariae vulgo triflorae inconspicuae saepius foliis vix

longiores breviter pedunculatae, floribus breviter pedicellatis, brac-

teis triangulari-subulatis pedicellis brevioribus; corolla luteo-viridis

4-loba 2 mm. lata, lobis ovali-ellipticis obtusissimis; fructus ut

videtur succosus dense vel sparse setulis rectis pallidis patulis ob-

tectus. — Mexico: Earth wall of canyon, Galeana, Nuevo Leon,

alt. 1,620 meters, August 4, 1939, V. H. Chase 7772 (type in Herb.

Field Mus.).

The present plant is one of a large collection obtained in Mexico

during the summer of 1939 by the Mexican Biological Expedition

of Students of the University of Illinois. It is an unusually distinct

species, in habit strongly resembling certain species of Relbunium,

to which genus it is certain to be referred at first glance. The setu-

lose fruit and, of course, the absence of an involucre at the base of

the flower necessitate its reference to Galium.

Gonzalagunia Rojasii Standl., sp. nov. — Arbor 6-metralis,

ramulis crassis subteretibus ochraceis vel pallide brunnescentibus

nodosis, novellis dense pilis longis rigidulis pallidis hispidis, inter-

nodiis saepe valde elongatis; stipulae e basi late triangulari acumi-

natae usque 8 mm. longae persistentes; folia magna breviter petiolata

membranacea, petiolo crassiusculo 1-2 cm. longo hispido; lamina

elliptica bel obovato-elliptica 11-20 cm. longa 5-9.5 cm. lata acuta

vel saepius breviter acuminata, basi acuta vel interdum longiuscule

angustata et plus minusve decurrens, supra viridis sparse pilis bre-

vibus hispidula, subtus vix pallidior ubique ad nervos venasque

breviter adpresso-pilosula, costa gracili elevata, nervis lateralibus

utroque latere ca. 9 angulo latiusculo adscendentibus arcuatis

STUDIES OF AMERICAN PLANTS — X 115

tenerrimis marginem fere attingentibus; inflorescentia spiciformis

pedunculata 16-21 cm. longa, floribus in cymulas parvas paucifloras

sessiles dispositis, rhachi crassa hispida, cymulis numerosis remote

insertis, bracteis minutis subulatis vel anguste triangularibus in-

conspicuis; hypanthium latum basi rotundatum 1.5 mm. longum

sparse puberulum vel fere glabrum, calyce 1 mm. alto brevissime

remote 4-dentato; corolla pallide viridescenti-lutea extus sparse

minute strigillosa, tubo gracili tereti 6 mm. longo, lobis ovato-

rotundatis vix 2.5 mm. longis; bacca 4-locularis depresso-globosa

4-5 mm. lata. — Guatemala: Damp forest, El Muro, below Santa

Maria de Jesus, Dept. Quezaltenango, alt. about 1,200 meters, March

1, 1939, Paul C. Standley 67145 (type in Herb. Field Mus.). Dense

damp forest, Finca Pireneos, below Santa Maria de Jesus, alt.

1,350 meters, March 11, 1939, Standley 68344.

Local name, Cachito de venado. The species is well marked

by the dense pubescence of very long, stiff, spreading hairs that

covers the branches. It is named for Professor Ulises Rojas, Director

of the Jardin Botanico of Guatemala, in whose company and as

whose guest the second collection was made, at Finca Pireneos.

Upon this and other days Professor Rojas was a most competent

and congenial guide to some of the most interesting regions of the

Guatemalan Occidente.

Hamelia barbata Standl., sp. nov. — Frutex vel arbuscula 3-

7.5 m. alta, ramis gracilibus viridibus minute pallide lenticellatis

glabris, internodiis brevibus; stipulae triangulari-acuminatae 3-4

mm. longae; folia opposita petiolata membranacea, petiolo gracili

glabro 1.2-2.5 cm. longo; lamina oblongo-oblanceolata vel oblongo-

elliptica 14-20 cm. longa 4.5-6 cm. lata subito longe angusteque

acuminata, basin versus angustata, basi ipsa acuta vel acuminata,

supra viridis glabra, subtus fere concolor in axillis nervorum dense

pallide villoso-barbata, aliter glabra, costa gracillima prominente,

nervis lateralibus utroque latere ca. 10 tenerrimis obliquis sub-

arcuatis angulo latiusculo adscendentibus; inflorescentia terminalis

cymoso-corymbosa breviter pedunculata laxe multiflora ca. 4.5 cm.

longa et 7 cm. lata, floribus secundis glabris, pedicellis gracilibus

2-3 mm. longis; hypanthium oblongo-turbinatum 3 mm. longum,

calycis dentibus vix ultra 0.5 mm. longis late deltoideis vel rotundatis

et apiculatis; discus valde elevatus 1.3 mm. altus; corolla aurantiaca

tubulosa vel superne dilatata et usque 5 mm. lata, ca. 18 mm. longa,

lobis latissimis ca. 2 mm. longis; stamina inclusa. — Guatemala:

Wet forest, Finca Monterrey, south slope of Volcan de Fuego,

116 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

Dept. Escuintla, alt. about 1,200 meters, February 5, 1939, Paul C.

Standley 64569 (type in Herb. Field Mus.). Panajabal, Dept. Chimal-

tenango, 1,350 meters, Standley 62113. In forest, Finca Moca, Dept.

Suchitepe"quez, alt. 810 meters, January, 1935, Skutch 2058.

The only closely related species is Hamelia longipes Standl.,

of the Atlantic lowlands of Guatemala and Honduras, which differs

in having much longer pedicels, and lacks the barbate nerve axils

characteristic of H. barbata.

Hamelia Haughtii Standl., sp. nov. — Frutex vel arbuscula,

ramulis gracilibus dense pilis longiusculis patulis pilosis, internodiis

elongatis; stipulae lineari-attenuatae 8-10 mm. longae pilosae; folia

magna longipetiolata membranacea opposita, petiolo gracili 3.5-6 cm.

longo patule piloso; lamina oblongo-ovata vel lanceolato-oblonga

16-23 cm. longa 6-8.5 cm. lata longiacuminata, basi acuta et inter-

dum subcontracta, supra viridis, ad costam pilosa, aliter glabra,

subtus ubique dense pilis rigidulis fulvis vel brunnescentibus pilosa,

nervis lateralibus utroque latere ca. 11 arcuatis tenerrimis; inflo-

rescentia terminalis cymoso-corymbosa ca. 5 cm. longa atque 10 cm.

lata 3 cm. longe pedunculata, laxe multiflora, ramis secundifloris,

floribus arete sessilibus; hypanthium 3 mm. longum oblongum basi

rotundatum, dense ferrugineo-pilosum, calycis dentibus brevissimis

triangularibus; corolla laete lutea extus sparse minute pilosula, tubo

2 cm. longo sursum sensim dilatato fauce 6 mm. lato, lobis rotundo-

obovatis ca. 5 mm. longis apice late rotundatis intus sparse puberu-

lis. — Colombia: Puerto Araujo, Dept. Santander, in moist, lowland

forest, April 15, 1937, Oscar Haught 2176 (type in Herb. Field Mus.);

in swamp forest, June 6, 1935, Haught 1751.

The principal distinctive characters are the large corollas and

the abundant, long, soft pubescence of the leaves and inflorescence.

Hoffmannia laxa Standl., sp. nov. — Frutex 2-metralis ut

videtur omnino glaber, ramis teretibus in sicco fuscis, internodiis

plerumque 3-6 cm. longis; folia modica longipetiolata tenuiter

membranacea, petiolo gracillimo 4-5 cm. longo superne longe

angusteque marginato; lamina elliptica vel late ovato-elliptica

12-16 cm. longa 6.5-8 cm. lata sensim vel subabrupte acuminata,

basi abrupte cuneato-acuta vel acuminata et longe anguste decur-

rens, supra intense viridis, costa venisque manifestis sed vix elevatis,

subtus multo pallidior, costa gracili prominente, nervis lateralibus

utroque latere ca. 9 arcuatis angulo lato saepe recto divergentibus

teneris prominulis, venis obscuris et laxe reticulatis; inflorescentiae

STUDIES OF AMERICAN PLANTS — X 117

numerosae axillares fasciculatae laxissime pauciflorae petiolis sub-

aequilongae gracillime ca. 2.5 cm. longe pedunculatae, pedicellis

fere filiformibus 4-7 mm. longis; hypanthium obovoideum fere 2 mm.

longum basi obtusum, calycis dentibus vix 0.6 mm. longis late

triangularibus acutis erectis; corolla in alabastro glabra 4 mm.

longa acuta vel acutiuscula, tubo ut videtur brevissimo, lobis tubo

duplo longioribus. — Costa Rica: Sucre, Canton de San Carlos, Prov.

Alajuela, in Caribbean rain forest, alt. 1,000 meters, March 30, 1939,

Austin Smith F1851 (type in Herb. Field Mus.).

"A shrub with several erect stems, the bark dull, dark green.

Leaves very dark green, faintly shining above, the petiole some-

times dull pink, the under surface pale. Flowers pale green, the

peduncle pale carmine." The species is noteworthy for the extremely

lax, few-flowered inflorescences with elongate pedicels.

Hoffmannia Wilsonii Standl., sp. nov. — Frutex 2-3-metralis,

ramis gracilibus teretibus ochraceis rimosis, internodiis plerumque

elongatis, novellis subdense breviter sordido-villosulis; stipulae vix

ultra 2 mm. longae, e basi late triangulari acuminatae; folia modica

longipetiolata tenuiter membranacea, petiolo gracili 3-5.5 cm. longo

breviter villosulo; lamina ovato-elliptica vel oblongo-elliptica 12-18

cm. longa 6-8 cm. lata subabrupte longiacuminata, basi acuta vel

subobtusa, supra viridis glabra, subtus paullo pallidior ad nervos

venasque adpresso-pilosa, nervis lateralibus utroque latere ca. 8

angulo lato adscendentibus arcuatis, venis laxe reticulatis; flores

in axillis pauci dense cymoso-aggregati sessiles vel breviter pedicel-

lati, cymulis sessilibus vel subsessilibus; hypanthium oblongo-turbi-

natum ca. 2 mm. longum glabrum vel sparsissime pilosulum, calycis

dentibus vix 1 mm. longis triangulari-subulatis erectis acutis;

corolla extus glabra vel hinc inde pilosa 7 mm. longa, tubo cylindra-

ceo, lobis angustis acutiusculis tubo multo longioribus. — Guatemala:

In virgin forest, Finca Los Alpes, Dept. Alta Verapaz, alt. 1,200

meters, March 21, 1939, C.L. Wilson 344 (type in Herb. Field Mus.).

Malanea Duckei Standl., sp. nov. — Frutex scandens, ramis

gracilibus teretibus atque densissime pilis longis adscendentibus

rigidulis ferrugineis vel pallidis pilosis, internodiis brevibus vel

elongatis; stipulae persistentes late ovatae 7-8 mm. longae dense

pilosae; folia majuscula breviter petiolata subcoriacea in sicco

fusco-brunnescentia, petiolo crasso 6-10 mm. longo piloso; lamina

elliptica vel late elliptica 8-14 cm. longa 5-10 cm. lata obtusa vel

acutiuscula, basi subrotundata vel obtusa, supra lucida, costa

nervisque ut quoque venis impressis, ad costam nervosque adpresso-

118 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

pilosa, aliter glabra vel glabrata, subtus brunnescens ubique pilis

longiusculis laxe adpressis pilosa, costa gracili elevata, nervis later-

alibus utroque latere ca. 9 angulo semirecto vel latiore adscendentibus

arcuatis elevatis, prope marginem arcuato-junctis, venulis elevatis

laxe reticulatis; flores parvi glomerulato-spicati, spicis axillaribus

4-5.5 cm. longis usque 3 cm. longe pedunculatis, glomerulis florum

ca. 7 mm. diam. sessilibus oppositis, paribus remotis, rhachi dense

patule pilosa, floribus arete sessilibus, bracteis minutis; calyx cum

hypanthio glaber campanulatus 1.5 mm. longus, margine undulato,

ferrugineus; corolla albida 5 mm. longa extus densiuscule albo-stri-

gosa, tubo cylindraceo superne vix dilatato, lobis 4 triangulari-oblongis

paten tibus obtusis intus albo-lanatis; stamina exserta, antheris oblon-

gis 1 mm. longis. — Brazil: Manaos, circa Cachoeira do Hindu, silva

non inundabili leviter paludosa, July 12, 1937, A. Ducke 34799 (type

in Herb. Field Mus.); same locality, May 12, 1937, Ducke 494.

Manettia Skutchii Standl., sp. nov. — Herbacea volubilis

omnino glabra, caule ca. 3 mm. crasso sulcato, internodiis foliis

longioribus; stipulae latae ca. 2.5 mm. altae apice rotundatae; folia

majuscula petiolata in sicco coriacea, petiolo crasso usque 14 mm.

longo superne marginato; lamina late elliptica vel oblongo-elliptica

6-9 cm. longa 3-5 cm. lata subito longiacuminata, basi obtusa vel

subrotundata atque abrupte contracta et decurrens, supra in sicco

cinerea sublucida, costa nervisque plus minusve impressis, lamina

subbullata, subtus fere concolor, costa crassa, nervis lateralibus

utroque latere ca. 5 teneris prominentibus solemniter arcuatis angulo

lato adscendentibus, venis paucis prominulis laxe reticulatis; inflo-

rescentiae axillares laxe pauciflorae racemose cymoso-paniculatae

foliatae usque 8 cm. longae ut videtur recurvae, pedicellis crassis

5-15 mm. longis, bracteis minutis; hypanthium obovoideum 3.5 mm.

longum; calycis tubus latus 2 mm. longus, lobis 4 ca. 3 mm. longis

late ovalibus vel subrotundatis incrassatis et carnosis, apice late

rotundatis vel subtruncatis, paten tibus; corolla pallido-rosea in

sicco subcoriacea, tubo crasso 13-15 mm. longo sursum solemniter

dilatato et fauce 6 mm. lato, basi 4 mm. lato, in fauce villoso, lobis

ovato-oblongis paten tibus vel subrecurvis ca. 8 mm. longis apicu-

latis et acutiusculis, intus ad apicem villosis; antherae semiexsertae.—

Ecuador: Vicinity of Puyo, Prov. Napo Pastaza, eastern foothills

of the Andes, alt. 750-1,000 meters, September, 1939, Alexander F.

Skutch 4525 (type in Herb. Field Mus.).

A related species is Manettia divaricata Wernham, of Peru and

Bolivia, which differs in having much thinner leaves and conspicu-

STUDIES OF AMERICAN PLANTS — X 119

ously narrower calyx lobes. In M. Skutchii the calyx lobes are

united below to form a distinct tube, as in M. divaricata.

Palicourea Cardenasii Standl., sp. nov. — Frutex 2-metralis,

ramis crassiusculis dense minute puberulis subteretibus, internodiis

2-3.5 cm. longis; stipulae persistentes basi in vaginam brevissimam

connatae, lobis erectis fere foliaceis semiovatis approximatis 5-7 mm.

longis acutis minutissime puberulis vel fere glabris; folia inter minora

petiolata subcoriacea, petiolo gracili 7-12 mm. longo minutissime

puberulo; lamina elliptico-lanceolata vel oblongo-lanceolata 7-9 cm.

longa 2.5-3.5 cm. lata longe angusteque interdum falcate acuminata,

basi cuneato-angustata, supra in sicco nigrescens ubique scaberulo-

puberula, costa nervisque prominentibus, subtus fere concolor ad

nervos costamque breviter pilosula vel adpresso-pilosula, inter ner-

vos glabrata, costa gracili prominente, nervis lateralibus utroque

latere ca. 10 obliquis angulo ca. semirecto adscendentibus leviter

arcuatis gracilibus prominentibus, venulis vix prominulis laxe reticu-

latis; inflorescentia terminalis erecta parva thyrsoideo-paniculata

5-8 cm. longa 3-4 cm. lata sat dense multiflora ca. 2 cm. longe

pedunculata, pedunculo ramisque dense sordido-puberulis, ramis

infimis valde adscendentibus brevibus, floribus sessilibus aggregatis;

hypanthium crassum vix ultra 0.6 mm. altum, calyce campanulari

ca. 0.8 mm. alto subtruncato vel late breviter sinuato-dentato;

corolla lutea anguste tubulosa ca. 7 mm. longa extus sparse minute

puberula, in alabastro apice obscure lobulata, lobis oblongis patenti-

bus vel subreflexis dimidium tubi fere aequantibus intus glabris

obtusis; antherae lineares subexsertae. — Bolivia: Chimore", Cocha-

bamba, at edge of forest, alt. 1,000 meters, November 2, 1937,

Martin Cardenas 2072 (type in U. S. Nat. Herb. ; fragment in Herb.

Field Mus.).

A species with no outstanding characters, but quite unlike any

one known previously from Bolivia.

Palicourea Danielis Standl., sp. nov. — Frutex 3-4-metralis,

ramis sat crassis teretibus, internodiis 2.5-4.5 cm. longis fere omnino

glabris; stipulae crassae in vaginam subtruncatam vix ultra 2 mm.

longam puberulam coalitae, lobis utroque latere 2 usque 3.5 mm. lon-

gis erectis rigidis triangulari-subulatis attenuatis persistentibus;

folia inter minora breviter petiolata crasse coriacea, petiolo crasso

3-4 mm. longo puberulo vel glabrato; lamina ovata vel lanceolato-

ovata 5-7.5 cm. longa 2-3 cm. lata subabrupte longiacuminata,

basi obtusa, supra in sicco lucida glabra, nervis venisque profunde

impressis, subtus ubique dense minute hirtella vel fere tomentulosa,

120 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

costa crassa solemniter elevata, nervis lateralibus utroque latere ca.

12 insigniter elevatis subarcuatis approximatis, venulis crassis

elevatis arete reticulatis sed vix conspicuis, marginibus valde late-

que revolutis; inflorescentia terminalis recta thyrsiformis in statu

fructifero ca. 5 cm. longa atque 4 cm. lata, pedunculo 3.5 cm. longo

dense sordido-pilosulo, ramis infimis radiatim divergentibus brevi-

bus dense sordido-pilosulis, floribus sessilibus vel subsessilibus aggre-

gatis ut dicitur luteis; hypanthium dense hispidulum, lobis calycis

brevibus acutiusculis; corolla perfecta non visa, 5 mm. longa vel

ultra dense hispidula; fructus ellipsoideus 5 mm. longus sparse

hispidulus, pyrenis solemniter costatis. — Colombia: San Pedro,

Dept. Antioquia, December, 1937, Brother Daniel & Brother Tomds

1554 (type in U. S. Nat. Herb.; fragment in Herb. Field Mus.).

The material, unfortunately, is not well preserved, but evidently

it represents an unusually well marked species, as species of this

genus are limited. The most conspicuous feature is found in the

very thick, coriaceous leaves, with the lateral nerves salient and

riblike on the lower surface, and strongly immersed on the upper

surface.

Posoqueria brachyantha Standl., sp. nov. — Arbor parva

omnino glabra, ramis sat gracilibus teretibus in sicco fuscis, inter-

nodiis bene elongatis; stipulae persistentes coriaceae triangulares

obtusae adpressae 7-10 mm. longae; folia modica breviter petiolata

coriacea in sicco fusco-brunnescentia, petiolo crasso 8-10 mm. longo;

lamina elliptica usque late elliptica vel rotundo-elliptica 7.5-11.5 cm.

longa 5-8 cm. lata obtusa vel acuta, interdum subrotundata, basi

obtusa vel rotundata, supra lucida, subtus paullo pallidior, costa

crassa prominente, nervis lateralibus utroque latere ca. 5 angulo

lato adscendentibus teneris prominentibus subarcuatis, venis ob-

soletis; inflorescentia cymoso-corymbosa terminalis sublaxe multi-

flora breviter pedunculata, pedicellis crassis usque 1 cm. longis;

hypanthium turbinatum 5 mm. longum superne ca. 4 mm. latum

basi acutum; calyx late campanulatus 2.5-3 mm. altus ad tertiam

partem lobatus, lobis late rotundatis ciliatis; corolla alba, tubo

gracili 4-6 cm. longo 2-3 mm. lato, lobis ovato-oblongis 8-10 mm.

longis apice rotundatis ciliolatis; antherae lineares lobos corollae

aequantes. — Brazil: Cucuhy, Rio Negro superior, in rupibus ad

basin montis granitici secus rivulum, September 9, 1935, A. Ducke

34992 (type in Herb. Field Mus.).

From all its close relatives the species is distinguished by its

unusually short corollas.

STUDIES OF AMERICAN PLANTS — X 121

Remijia amphithrix Standl., sp. nov. — Arbuscula 3-6-metralis,

ramulis crassis dense breviter pilosulo-tomentosis; stipulae ovali-

ovatae fere 2.5 cm. longae 11 mm. latae obtusae extus densissime

adpresso-pilosae; folia magna breviter petiolata subcoriacea, petiolo

crasso 2-3 cm. longo densissime breviter patenti-piloso; lamina late

elliptica ca. 29 cm. longa et 15 cm. lata obtusa et breviter cuspidata,

basi subrotundata et abrupte breviter contracta, supra in sicco

brunnescens sparsiuscule pilis brevibus patulis vel subadpressis

fulvis conspersa et sparse pilis longis patentibus hirsuta, costa

venisque non elevatis, subtus pallide fulva, ad venas costamque

dense strigosa, aliter pilis brevibus subpatulis densissime pilosa,

costa crassa elevata, nervis lateralibus utroque latere ca. 16 teneris

prominentibus, inferioribus angulo fere recto abeuntibus, superioribus

angulo angustiore usque semirecto adscendentibus arcuatis marginem

attingentibus, venis tomento occultis; inflorescentia spiciformis

interrupta ca. 17 cm. longe pedunculata, inferne aliquanto foliaceo-

bracteata, floribus in cymulas capituliformes densissime multifloras

sessiles vel infimas usque 2 cm. longe pedunculatas dispositis sessili-

bus; cymulae oppositae, paribus infimis usque 8 cm. distantibus;

hypanthium campanulatum basi obtusum 2.5 mm. longum dense

ochraceo-tomentosum, calyce vix ultra 2 mm. longo adpresso-

pilosulo inaequaliter dentato, dentibus late ovatis acutiusculis;

corolla pentamera extus dense adpresso-pilosa, tubo 10-12 mm.

longo sursum vix dilatato fauce 2-2.5 mm. lato, lobis usque 8 mm.

longis patentibus lanceolato-linearibus intus glabris vel sparse

minutissime puberulis apicem subobtusum versus sensim attenuatis;

stamina inclusa. — Colombia: Comisaria El Vichada, edge of forested

area about 70 km. southeast of Orocue", alt. 150 meters, April 25,

1939, Oscar Haught 2789 (type in Herb. Field Mus.).

From all other Colombian species of the genus this is conspicu-

ously different in the large and wide leaves that are broad and

rounded at the base. The collector states that the white flowers

have a peculiar odor that is not very agreeable.

Remijia asperula Standl., sp. nov. — Arbuscula, ramis crassis

subteretibus vel obtuse tetragonis dense hirsuto-pilosis vel glabratis,

internodiis abbreviatis; stipulae magnae oblongo-ovatae 4 cm.

longae et 2 cm. latae vel majores obtusae vel breviter acuminatae

intus glabrae extus densissime pilis longis subadpressis hispido-

pilosae; folia magna opposita breviter petiolata subcoriacea, petiolo

crasso 1-2 cm. longo vel paullo ultra dense hirsute; lamina oblanceo-

lato-oblonga usque oblongo-obovata 22-30 cm. longa 7.5-14 cm.

122 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

lata acuminata vel subobtusa et breviter cuspidata, basi acuta vel

contracta et anguste decurrens, basin versus plus minusve angustata,

supra in sicco fusca sublucida hirsuta et scabro-pilosula, tactu

asperula, costa nervisque non elevatis, subtus pallide brunnescens

ubique dense pilis longis submollibus patentibus fulvis hirsuta, costa

gracili elevata, nervis lateralibus utroque latere ca. 18 teneris promi-

nentibus angulo latiusculo divergentibus fere rectis marginem attin-

gentibus, venis inconspicuis laxe reticulatis; flores majusculi dense

cymoso-aggregati, cymulis dense multifloris saepe capituliformibus,

post anthesin laxioribus, bracteatis paniculatis, paniculis axillaribus

usque 17 cm. longe pedunculatis pauciramosis usque 25 cm. longis,

ramis basalibus usque 4.5 cm. longis sed vulgo brevioribus dense

brunneo-pilosis; bracteae ex parte magnae et foliosae, plerumque

ellipticae vel ovatae et 10-12 mm. longae dense strigoso-sericeae;

pedicelli nulli vel usque 4 mm. longi crassiusculi; hypanthium obo-

voideum 3 mm. longum basi subobtusum densissime longipilosum;

calyx 3-4 mm. longus extus dense adpresso-pilosus ad medium den-

tatus, dentibus triangularibus acutis erectis; corolla alba in ala-

bastro fere 15 mm. longa sursum angustata extus densissime pilis

longis adscendentibus brunnescentibus pilosa; capsula anguste

oblonga vel lanceolato-oblonga 2-3 cm. longa ca. 7 mm. lata sursum

angustata, basi acutiuscula, densissime pilis longis et brevibus inter-

mixtis brunneis vel ferrugineis piloso-tomentosa; semina numerosis-

sima, parte seminifera ca. 2 mm. longa utrinque ala albida tenui

ca. 2 mm. longa aucta. — Brazil: Manaos, civ. Amazonas, silva

secundaria non inundabili loco arenoso humido prope cataractam

altam fluminis Taruma, May 13, 1936, A. Ducke 22854 (type in

Herb. Field Mus.).

The species bears some superficial resemblance to R. Ulei Krause,

apparently a rather common species of the Upper Amazon, but

that has only rather sparse and very inconspicuous pubescence, and

its leaves are relatively much longer and narrower. The capsules of

the two species are similar.

Remijia Duckei Standl., sp. nov. — Arbuscula, ramis crassis

subteretibus dense longipilosis, internodiis abbreviatis; stipulae

tenues ferrugineae oblongo-lanceolatae vel ovato-oblongae 5-10 cm.

longae basi usque 3 cm. latae longiattenuatae extus longipilosae

diutius persistentes; folia magna opposita petiolata subcoriacea,

petiolo crasso 2-3 cm. longo piloso-tomentoso; lamina oblonga,

anguste lanceolato-oblonga vel late elliptico-oblonga 15-55 cm.

longa 5-20 cm. lata, acuta vel breviter acuminata, basi acutiuscula

STUDIES OF AMERICAN PLANTS — X 123

usque rotundata, supra in sicco fusco-brunnescens, primo praesertim

ad costam nervosque laxe pilosa, cito glabrata, sublucida, costa

nervisque non elevatis, subtus pallide brunnescens vel brunneo-

ochracea in areolis intra nervulos minutissime fere microscopice

pallido-tomentella, aliter sat dense laxeque pilosa vel ad nervos

adpresso-pilosa, serius glabrescens, costa gracili elevata, nervis

lateralibus utroque latere ca. 21 teneris elevatis angulo latiusculo

adscendentibus fere rectis juxta marginem abrupte sursum curvatis

et in marginem desinentibus, venulis prominulis arctissime reticu-

latis; flores majusculi in cymulas parvas dense paucifloras bracteatas

dispositi, cymulis ad apices ramorum cymosis paniculam usque

25 cm. longe pedunculatam 7-25 cm. longam racemiformem vel late

pyramidalem efformantibus, ramis laxe subsparse piloso-tomentosis,

infimis obliquis usque 6 cm. longis; bracteae lineari-filiformes usque

2 cm. longae; flores sessiles vel brevissime pedicellati; hypanthium

obovoideum 3 mm. longum dense pilis longis subadpressis pilosum,

calyce 4-5 mm. longo campanulato breviter vel longius dentato,

dentibus triangularibus usque fere subulatis; corolla alba in ala-

bastro 2.8 cm. longa sursum attenuata extus densissime pilis longis

pallidis adpressis sericea, lobis lanceolato-linearibus ca. 15 mm.

longis longiattenuatis; capsula oblongo-clavata 2.5-3 cm. longa

apice obtusa, basi angustata, valvulis ca. 8 mm. latis. — Brazil:

Santa Izabel, Rio Negro, civ. Amazonas, ad ripas Igarap£ Dara,

September 9, 1935, A. Ducke 35058 (type in Herb. Field Mus.).

A relative, apparently, of R. firmula Wedd., also of the upper

Amazon and the Rio Negro, which has capsules only 1 cm. long,

according to description, and much less abundant pubescence.

The type material of R. Duckei consists of three sheets of specimens.

One of these bears a leaf of considerable morphological interest,

since the upper half of the blade is very coarsely and regularly den-

tate. This is the closest approach to a truly dentate leaf that I

have ever seen among American Rubiaceae, and is scarcely sug-

gestive even of the regularly pinnatifid leaves found in some species

of Pentagonia.

Stachyarrhena acuminata Standl., sp. nov. — Arbor parva

glabra, ramulis cinnamomeis, cortice exfoliante, internodiis 6-15 mm.

tantum longis teretibus; folia magna petiolata coriacea, petiolo

crassiusculo 10-15 mm. longo; lamina anguste oblongo-lanceolata

18-28 cm. longa 5-7 cm. lata longiacuminata, basin versus sensim

angustata, basi ipsa acutissima, supra lucida, costa nervisque promi-

nentibus, subtus brunnescens minute papillosa, costa gracili elevata,

124 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

nervis lateralibus utroque latere ca. 15 angulo lato interdum fere

recto abeuntibus teneris prominentibus arcuatis prope marginem

sursum curvatis, venis obscuris; inflorescentia terminalis erecta

2-2.5 cm. longa ca. 3 cm. lata anguste paniculata 1-1.5 cm. longe

crasse pedunculata, floribus in parte superiore spicatis, inferioribus

in cymulas parvas paucifloras dispositis, pedicellis crassissimis vix

ad 3 mm. longis; hypanthium brevissimum, calyce late campanulato

2.5 mm. alto 3 mm. lato lucido truncate coriaceo; corolla coriacea

alba, tubo lato 4 mm. longo, lobis late ovatis obtusis suberectis ca.

3.5 mm. longis intus tomentosis. — Brazil: Borba, Rio Madeira,

civ. Amazonas, silva non inundabili, November 8, 1935, A. Ducke

35053 (type in Herb. Field Mus.).

Doubtless related to S. spicata Hook. f. of the Amazon Valley,

which has relatively shorter and much broader leaves, and normally

a much more elongate inflorescence. The somewhat similar S.

longifolia (Spruce) Hook. f. has obtuse leaves and a simply spicate

inflorescence. The type of S. longifolia is well matched by the

following collection: Rio Negro superior, civ. Amazonas, ad ripas

sabulosas infra Marituba, a shrub with white flowers, March, 1936,

Ducke 35051.

Stachyarrhena Duckei Standl., sp. nov. — Arbor parva omnino

glabra, ramulis gracilibus subteretibus in sicco fusco-ferrugineis,

internodiis 1.5-4.5 cm. longis; vagina stipularis truncata annuli-

formis vix ultra 1 mm. alta; folia majuscula petiolata subcoriacea,

petiolo crassiusculo 1.5-2 cm. longo; lamina lanceolato-oblonga

15-19 cm. longa 6-8 cm. lata sensim longiacuminata, basi acuta,

supra in sicco fusco-brunnescens sublucida, costa prominente, nervis

manifestis sed vix elevatis, subtus fere concolor, costa prominente,

nervis lateralibus utroque latere ca. 12 angulo latiusculo abeuntibus

tenerrimis prominentibus subarcuatis prope marginem arcuato-

conjunctis, venulis vix prominulis laxe reticulatis; inflorescentia

terminalis erecta 2.5 cm. longe pedunculata simpliciter spicata

5.5 cm. longa 1 cm. lata sat dense multiflora, floribus arete sessilibus;

calyx latissime campanulatus 3 mm. latus vix 2 mm. altus truncatus

lucidus; corolla alba in alabastro 5.5 mm. longa obtusissima. —

Brazil: Santa Izabel, Rio Negro, civ. Amazonas, ad ripas inundabiles,

December 9, 1936, A. Ducke 35052 (type in Herb. Field Mus.).

Although similar to S. acuminata, this differs in having a strictly

simple inflorescence, and its leaves also are relatively broader,

besides being widest near the base rather than at the middle. The

leaves are longer and much more acuminate than those of S. spicata.

STUDIES OF AMERICAN PLANTS — X 125

Stachyarrhena reflexa Standl., sp. nov. — Frutex 4-metralis

omnino glaber, ramis crassiusculis ochraceis vel fulvo-cinereis ali-

quanto nodosis, internodiis plerumque abbreviatis; vagina stipularis

ca. 4 mm. longa coriacea breviter lobata, lobis apice rotundatis;

folia petiolata coriacea rigida, petiolo crasso 1-1.5 cm. longo; lamina

anguste oblonga 6-14 cm. longa 2.5-3.8 cm. lata obtusa, basi an-

gustato-acuta, supra in sicco fusca lucida, costa nervisque promi-

nulis, subtus brunnescens sublucida, costa gracili elevata, nervis

lateralibus utroque latere ca. 13 tenerrimis vix prominulis angulo

lato abeuntibus, venis fere obsoletis; inflorescentia simpliciter spicata

ca. 3 cm. longa et 5 cm. longe pedunculata, abrupte reflexa, floribus

in verticillis densis multifloris remotis aggregatis arete sessilibus;

calyx late campanulatus truncatus lucidus ca. 5 mm. latus et 3 mm.

altus; corolla in alabastro 6-7 mm. longa apice obtusa. — Brazil:

Capoeira, Fazenda Sao Bento-Merces de Diamantina, Municipio

Diamantina, Minas Geraes, November 24, 1937, H. Mello Barreto

10012 (Herb. Jard. Bot. Bello Horizonte 22237; type in Herb.

Field Mus.).

Because of its reflexed and pendent inflorescence, this evidently

is related to S. penduliflora Schum., of Para. The plant of that

rather remote region, however, has leaves twice as broad, and

mostly very obtuse or rounded at the base.

VALERIANACEAE

Valeriana cacalioides Standl., sp. nov. — Herba elata simplex

ca. 90 cm. alta, caule 7-8 mm. crasso aliquanto fistuloso in sicco

brunnescente et marmorato-maculato, internodiis usque 18 cm.

longis glabris vel apicem caulis versus sparse breviter hirsutulis;

folia ca. 4-juga longipetiolata membranacea in sicco fusca, petiolo

1.5-6 cm. longo vel in foliis supremis fere nullo sparse breviter

pilosulo; lamina late rotundato-ovata vel fere hastato^reniformis

5.5-7.5 cm. longa 5-8.5 cm. lata acuta, basi profunde aperte cordata,

prope basin hastato-angulata, aliter grosse sinuato-dentata vel

sublobata, sparse praesertim prope basin breviter pilosa vel fere

glabra, basi palmatim ca. 7-nervia; inflorescentia terminalis longi-

pedunculata e cymis 2 capituliformibus dense multifloris ca. 2 cm.

latis 2 cm. longe pedunculatis composita, ramis dense pilosis, brac-

teis foliaceis linearibus usque 5-6 mm. longis; corolla glabra fere

5 mm. longa, lobis subpatentibus late oblongo-ovatis obtusissimis

tubo duplo vel triple brevioribus; stamina longiexserta. — Mexico:

Mount Ovando, Chiapas, April 12, 1937, E. Matuda 1812 (type in

Herb. Field Mus.).

126 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. 22

The large, broad leaves are a distinctive character of the plant,

reminding one somewhat by their form of those of Ipomoea Batatas

or of some other members of the Convolvulaceae.

COMPOSITAE

Actinea ursina Standl., sp. nov. — Herba perennis caulescens

pauciramosa ca. 30 cm. alta erecta, caulibus pluribus gracilibus

capitulis ca. 3 terminatis sat dense et laxissime albido-lanatis, inter-

nodiis foliis longioribus; folia integra, basalia ca. 6 cm. longa, 6-7

mm. lata lineari-oblanceolata apice anguste rotundata vel obtusa,

basin versus longe sensimque attenuata et in petiolum marginatum

decurrentia, utrinque pilis laxis longis lanata; folia caulina linearia

sessilia vel subsessilia 2.5-6 cm. longa 2-3 mm. lata laxe lanata;

pedunculi graciles usque 17 cm. longi laxe denseque lanati; capitula

majuscula, disco 1.5 cm. lato fere 1 cm. alto, involucre dense pilis

longis albis lanato, phyllariis obtusis; radii lutei ca. 1 cm. longi et

3 mm. lati, apice breviter tridentati, extus glanduloso-pilosuli. —

Mexico: State of Nuevo Leon, scattered on drier, more open slopes,

Las Canoas, on Cerro Potosi, Municipio de Galeana, July 17, 1935,

C. H. Mueller 2189 (type in Herb. Field Mus.).

As this group of plants is treated by Rydberg (N. Amer. Fl. 34:

101. 1915), under the generic name Tetraneuris, this seems to be

related to Tetraneuris formosa Greene and T. Ivesiana Greene, of

the southwestern United States, but it is distinct from those and their

relatives by its abnormally long and lax, copious pubescence.

Erigeron chiriquensis Standl., sp. nov. — Herba erecta ramosa

elata 20-50 cm. alta, caule superne pauciramoso crassiusculo

subtereti sparsissime pilis brevibus albidis strigoso vel glabrato,

internodiis foliis brevioribus; folia numerosa sessilia 2-3 cm. longa

4-8 mm. lata acuta, inferiora pinnatilobata, segmentis lateralibus

paucis oblongis acutis, superiora grosse pauciserrata, suprema in-

tegra vel breviter serrata et saepe linearia, basin versus integra et

sensim angustata, sparse utrinque albido-strigosa, crasse membra-

nacea; capitula terminalia solitaria vel aggregata breviter peduncu-

lata, disco ca. 12 mm. lato et 6 mm. alto, pedunculis apice dense

strigosis; phyllaria ca. 6 mm. longa subaequalia linearia longiat-

tenuata hirsutula pallido-marginata; radii albi vel pallide caeru-

lescentes numerosi anguste lineares 3-4 mm. longi; corolla filiformis

glabra usque 7 mm. longa; achaenia paullo ultra 1 mm. longa sparse

hispidula, pappi setis paucis 3.5 mm. longis albis minutissime scab-

erulis. — Panama: Potrero Mulato, Volcan de Chiriqui, Boquete

STUDIES OF AMERICAN PLANTS— X 127

District, Prov. Chiriqui, alt. 3,140 meters, July 18, 1938, Mrs. M. E.

Davidson 872 (type in Herb. Field Mus.).

Erigeron potosinus Standl., sp. nov.— Herba perennis erecta

elata 30 cm. alta vel paullo ultra, caulibus subsimplicibus capitulis

2 longipedunculatis terminatis minute sparse puberulis et sparse

hispidulo-pilosis paucifoliatis, internodiis saepe foliis longioribus;

folia basalia 6-7 cm. longa petiolata, petiolo marginato, lamina ob-

longo-oblanceolata usque 4.5 cm. longa et 1.5 cm. lata integra apice

rotundata fere glabra sed saltern subtus sparse puberula; folia

caulina ca. 8 subaequalia, superiora paullo minora, oblonga vel

lanceolato-oblonga acuta vel obtusa, basi sessilia et subamplexi-

caulia, integra, fere glabra ciliata; pedunculi graciles 9-15 cm. longi

viscido-puberuli et patule pilosi; capitula majuscula, disco ca. 1.5 cm.

lato et 7-8 mm. alto, phyllariis linearibus subaequalibus, exterioribus

viridibus dense minuteque glanduloso-puberulis longiattenuatis

erectis et subadpressis numerosissimis; radii albi purpureo tincti

anguste lineares usque 1.5 cm. longi; achaenia juvenilia ochracea

1.5 mm. longa sparse puberula, pappi setis paucis duplo longioribus

albis mollibus. — Mexico: State of Nuevo Leon, abundant on forest

floor, Las Canoas, on Cerro Potosi, Municipio de Galeana, July 16,

1935, C. H. Mueller 2174 (type in Herb. Field Mus.). Below timber

line, near peak of Cerro Potosi, alt. 3,450 meters, July 20, 1938,

Richard A. Schneider 945.

Regarding the second collection cited, Mr. Schneider has made

the following notes: "Abundant on open mountain side; spreading

by offshoots; rays white or pale pink. A pretty daisy of open,

rather rocky woods, and of open mountain slopes. Usually only one

head to a stem. Just coming into bloom July 20, while the other

Erigeron of this altitude was in full bloom when first observed on

July 1."

Helenium pinetorum Standl., sp. nov. — Herba perennis

robusta usque 60 cm. alta, caule vulgo tricephalo simplici inferne

1 cm. crasso striato glabro vel sparsissime lanato, internodiis foliis

multo brevioribus; folia caulina numerosa sessilia, inferiora et media

anguste oblanceolata usque 14 cm. longa 1.7-2.5 cm. lata attenuato-

acuta, basin versus sensim angustata glabra vel glabrata paralleli-

nervia; folia superiora lanceolata vel anguste ovata 3.5-8 cm. longa

1.5-2.5 cm. lata longiacuminata, basi dilatata et obtusa vel rotun-

data, fere glabra vel primo sparse lanata, integra; pedunculi crassi

3.5-7 cm. longi dense lanati; capitula magna, disco ca. 2 cm. lato

et 1.2 cm. alto, phyllariis numerosis lineari-lanceolatis viridibus

128 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. 22

longiattenuatis herbaceis dense lanatis laxis; radii lineares 2.5 cm.

longi intense lutei; pappi squamellae lineari-lanceolatae 3.5 mm.

longae subulato-attenuatae. — Mexico: State of Nuevo Leon, Sierra

Madre Oriental, ascent of Sierra Potosi by north hogback, about

20 miles northeast of Galeana, abundant in the upper pine forest,

alt. 3,390 meters, July 26, 1934, C. H. & M. T. Mueller 1258 (type

in Herb. Field Mus.). Peak of Cerro Potosi, abundant in upper pine

woods, north side, in loose clumps of 4-10 plants, July 21, 1935,

C. H. Mueller 2274.

The type collection was distributed as Helenium integrifolium

(HBK.) Benth. & Hook., the only similar species, which was de-

scribed from the State of Hidalgo, and grows in the mountains of

Guatemala as well as on the high peaks of southern Mexico. In

H. integrifolium, as indicated by ample material at hand for study

and by the original illustration of Kunth, the plant is densely lanate

throughout, and the upper cauline leaves are all narrow, almost

linear, or else divided into narrow lobes. In H. pinetorum all the

leaves are entire, and the upper ones are broad, especially at the base.

Senecio Austin-Smithii Standl., sp. nov. — Frutex epiphyticus

1.5 m. longus, ramis dense foliatis ochraceis subteretibus glabris,

novellis in sicco fuscis, internodiis abbreviatis; folia inter minora

petiolata in sicco subcoriacea et fusca, petiolo gracili 8-10 mm.

longo; lamina anguste oblanceolato-oblonga 4-7.5 cm. longa 1.7-2.5

cm. lata apice obtusa vel anguste rotundata, basin versus cuneato-

attenuata, integra glabra; inflorescentia terminalis cymoso-corym-

bosa 5 cm. longa et aequilata, pedicellis usque 4 mm. longis sparse

araneoso-lanuginosis vel fere glabris; capitula numerosa condensata

eradiata anguste campanulata; phyllaria ca. 7 et 5 mm. longa an-

guste oblonga obtusissima pallido-marginata glabra vel basi tantum

obscure lanuginosa, bracteolis paucis brevibus; corollae 4-5 mm.

longae glabrae; achaenia immatura glabra, pappi setis longis albis

mollibus. — Costa Rica: Palmira, Prov. Alajuela, alt. 1,860 meters,

February 9, 1938, Austin Smith H299 (type in Herb. Field Mus.).

"An epiphyte with woody stems 1.5 meters long, on forest tree in

semi-shade; bark smooth and pale brown; leaves a little thickened,

flexible, glabrous and shining; inflorescence green."

Solidago Muelleri Standl., sp. nov.— Herba perennis elata

ca. 70 cm. alta ut videtur simplex, caule dense foliato gracillimo

cinereo-puberulo, internodiis foliis multo brevioribus; folia lanceo-

lato-linearia 4-9 cm. longa 3-9 mm. lata longe angusteque attenuata,

STUDIES OF AMERICAN PLANTS — X 129

basin versus longe angusteque attenuata et plus minusve petiolata,

remote minute adpresso-serrata, vel superiora interdum integra,

1-nervia vel obscure trinervia, supra in sicco griseo-viridia sparse

puberula vel fere glabra, subtus praesertim ad costam gracilem

prominentem minute pilosula; capitula numerosa parva secunda in

racemos breves terminales et axillares 2-3 cm. longos disposita,

inflorescentia tota anguste paniculata foliata 15 cm. longa basi 3.5

cm. lata, pedicellis tenuibus ad 3 mm. longis adpresso-pilosulis;

capitula 3 mm. alta campanulata, phyllariis inaequalibus late

linearibus obtusis adpressis stramineis apice ciliatis, dorso glabris;

achaenia 1.3 mm. longa minute hispidula, pappi setis albis rigidulis

ca. 2 mm. longis; radii pallide lutei fere lineares floribus disci vix

longioribus. — Mexico: State of Nuevo Leon, common in moist

crevices on river bank, Canon Marisio Abajo, Rancho Las Adjuntas,

Municipio de Villa Santiago, June 27, 1935, C. H. Mueller 2062

(type in Herb. Field Mus.).

OF THE

JUN 2 9 1940

UNIVERSITY OF IUW

UNIVERSITY OF ILLINOIS-URBANA