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THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY,
ZOOLOGY, BOTANY, ann GEOLOGY.
(SEING A CONTINUATION OF THE ‘ANNALS ’ COMBINED WITH LOUDON AND
CHARLESWORTH’S ‘MAGAZINE OF NATURAL HISTORY. )
CONDUCTED BY
PRIDEAUX JOHN SELBY, Esa., F.I.S.,
CHARLES C. BABINGTON, Esea., M.A., F.R.S., F.L.S., F.G.S.,
JOHN EDWARD GRAY, Ph.D., F.R.S., F.L.S., V.P.Z.S. &c.,
AND
WILLIAM FRANCIS, Ph.D., F.L.S.
Oe en ee t
VOL. XVI.—THIRD SERIES...
~——_——— reer eee oe
LONDON:
PRINTED AND PUBLISHED BY TAYLOR AND FRANCIS.
SGLD BY LONGMANS, GREEN, READER, AND DYER; SIMPKIN, MARSHALL, AND CO.35
KENT AND CO.; BAILLIERE, REGENT STREET, AND PARIS;
MACLACHLAN AND STEWART, EDINBURGH ;:
HODGES AND SMITH, DUBLIN: AND ASHER, BERLIN.
1865.
‘*Omnes res create sunt divine sapientic et potentiz testes, divitiz felicitatis
humane :—ex harum usu bonitas Creatoris; ex pulchritudine sapientia Domini;
ex ceconomia in conservatione, proportione, renovatione, potentia majestatis elucet.
Earum itaque indagatio ab hominibus sibirelictis semper estimata; a veré eruditis
et sapientibus semper exculta; malé doctis et barbaris semper inimica fuit.”—
LINNZUS.
“ Quel que soit le principe de la vie animale, il ne faut qu’ouvrir les yeux pour voir
qu'elle est le chef-d’ceuvre de la Toute-puissance, et le but auquel se rapportent
toutes ses opérations.”—BRUCKNER, Théorie du Systéme Animal, Leyden, 1767.
po6 0600 0n 0 Do WN b EN poe
Obey our summons; from their deepest dells
The Dryads come, and throw their garlands wild
And odorous branches at our feet ; the Nymphs
That press with nimble step the mountain thyme
And purple heath-flower come not empty-handed,
But scatter round ten thousand forms minute
Of velvet moss or lichen, torn from rock
Or rifted oak or cavern deep: the Naiads too
Quit their loved native stream, from whose smooth face
They crop the lily, and each sedge and rush
That drinks the rippling tide: the frozen poles,
Where peril waits the bold adventurer’s tread,
The burning sands of Borneo and Cayenne,
All, all to us unlock their secret stores
And pay their cheerful tribute.
J. Taytor, Norwich, 1818.
FLAMMAM,
CONTENTS OF VOL. XVI.
[THIRD SERIES. ]
NUMBER XCI.
Page
I. Descriptions of new Genera and Species of Diatoms from Hong-
kong. By R. K. Grevitue, LL.D., F.R.S.E. &e. (Plate V.)...... 1
II. Notes on British Lichens. By the Rev. W. A. Le1guron,
BeAeptcboslie m(biate LVa)) icactcccerceeebdcuscastecacemenosceacmee ore Ole
III. On the Gland of the Phyllodium of Acacia magnifica. By
thepReveaWi Ac WEIGH TONG Db Aches S big sasecserconessceaeteoerceeener 12
IV. On the Nomenclature of the Foraminifera. By Messrs.
W. K. Parker, F.R.S., T. R. Jones, F.G.S., and H. B. Brapy,
BES FGos, (Plates T5010.,, DME) sees. ce. 2ce ess ces osacseeeeeeeeinees 15
V. On Ageon Alfordi, a new British Sea-Anemone. By PHILIP
Eien GLOSS eb. tS. (ESL RVI) | cee ease arn sesscscecetensetcactonsees 41
VI. On the Presence of certain Secreting Organs in Nematoidea.
By ALEXANDER MACALIsTER, F.R.C.S.I., Demonstrator of Ana-
tomy, Royal College of Surgeons, Ireland ..........-.cscccesseceesesccoees 45
VII. Description of a new Species of Corvina from the Gambia.
yr AEB Ps GUNTUR Reape os cm ansoeseccicaneactaseeacnaecddeerss peweesenss 48
Proceedings of the Zoological Society .....scscsereceesssecsereee sheets 49—65
On the Pollen-grains of Ranunculus arvensis, by George Gulliver,
F.R.S.; On the Feathers of Dinornis robustus, Owen, by W.S.
Dallas, F.L.S., Keeper of York Museum; On the Metamorphoses
undergone by certain Fishes before acquiring the Adult Form,
by Prof. Agassiz; Description of the Egg of Parra gallinacea,
by John Gould, F.R.S. &c.; On a new Form of Brachiolaria, by
M. Sars; Investigation of the Structure of the Eneephalon of
Fishes, and of the Homological Signification of its different parts,
lv CONTENTS.
by M. Hollard ; Description of a new Species of Rock- Kangaroo
(Pterogale longicauda) fron New South Wales, by Gerard Krefft
Page
65—72
NUMBER XCII.
VIII. On the Homology of the Buccal Parts of the Mollusca. By
Dr. Orro A. L. Mércu, of Copenhagen. (Plate VI.)
IX. Descriptions of recently discovered Spiders collected in the
Cape de Verde Islands by John Gray, Esq. By Jonn BLAcKWALL,
Me S renseaaves cone ssvconcecsaessenans\Jascsisascacsttecsersaseesene-a-heo sete ee
X. Contributions to an Insect Fauna of the Amazons Valley.
CoLeorpTerA: Loneicornes. By H.W. Bares, Esq.
XI. On the Occurrence of Limopsis Belcheri, Corbula sulcata, and
some other recent Shells in the fossil state in Miocene Tertiary Beds
near Melbourne. By Frepertck M‘Coy, Professor of Natural
Science in the University of Melbourne, and Director of the Melbourne
National Museum 80. oa. scssccseeseteucescs asic tvces saceeanasaneevens ease ee
XII. Observations on Raphides and other Crystals in Plants.
By GrornGe (GULLIVER, EOIN.) sscescccdseceasae cess seeasmeartareteastenees
XIII. Gn the Opereculum and its Mantle. By Dr. O. A. L.
MORCH eseeecees eee
POOR e OOH Oe eer SHEE EEE POD HEH EHH EEH HEH EEE HEE EEE EEE EOE OEE EOE EEE
XIV. Notes on some Amphibians. By Joun Hoae, M.A., F.RS.,
mI Seco nace cee csiaeescsvecssa cos cssecmeneescsanoaararncceeceeceecsaveecceaeenees
XV. Remarks on the Histology of two Specimens of Rhynchopora
Geinitziana, De Verneuil, from near the River Oukhta, Province of
Archangel, and belonging to the Collection of the Corps des Mines of
St. Petersburg. By Professor W1LL1AM KiNG
73
101
Proceedings of the Royal Society ; Zoological Socicty .......... 128—142
Note on the Food of the Aye-Aye, by Mr. A. D. Bartlett ; On the His-
tology of the Acalephz, by Prof. Kolliker; On a new Type in
the Group of Ascidians (Chevreulius callensis), by M. Lacaze-
Duthiers ; On some singular Organs appended to the Feet of
certain Crustacea, by MM. Claus and Sars .........s0eseeeee 142—144
NUMBER XCIII.
XVI. On a new Lizard, with Ophidian Affinities, from the Lower
Chalk (Saurospondylus dissimilis). By Harry SEELEY, F.G.S., of
the Woodwardian Museum, Cambridge
eee eUEeR ECE eee eeer er ee eee errr e
CONTENTS. Vv
Page
XVII. Notice of a new Finner Whale from Formosa. By Dr. J.
IE) GRAYS EMRE SSI SCCS) Arse tensinstewGen sd nccsesacsasicates's CSpOORDBOnEE pebaccnoacoC 148
XVIII. On the Male Generative Organs of Phalangium. By Dr.
AC GCRQEUN sesseocss socewelse Mer eaemteceee senses cdariccese tac rsidenadacasndsaideee ores 149
XIX. Descriptions of new Species of Crioceride. By J.S. Baty. 153
XX. Description of a new Species of Cetonia in the Collection of
the British Museum. By Arruur G. Bur er, Assistant, Zoological
Department, British Museum ......... Aecleceacesecdiesccsees Sscaedesseetera 161
XXI. Investigations on new or rare Crustacea of the French
Coastsa Dy MOSS aistececcccomsdeeonaacccccdessesencesecssecsceasasect 162
XXII. Contributions to an Insect Fauna of the Amazons Valley.
CoLreopTEeRA: Loncicornes. By H. W. Bares, Esq. ............ 167
XXIII. A Description of some Fossil Corals from the South-
Australian Tertiaries. By P. Martin Duncan, M.B.(Lond.), See.
Geol-soe! (Plate, VIM.) <::..... Bete oeae sence seadaeneodoe one cea eeneneeeeet 182
XXIV. Notes on the Australian Species of Arripis. By FREDE-
RIcK M‘Coy, Professor of Natural Science in the University of Mel-
bourne, and Director of the National Museum at Melbourne ......... 187
XXV. On Undescribed Fossil Entomostraca from the Brick-earth
of the Nar. By GreorGe Stewarpson Brapy. (Plate IX.) ...... 189
XXVI. Classification of Polyps. (Extract condensed from a
Synopsis of the Polypi of the North Pacific Explormg Expedition
under Captains Ringgold and Rodgers, U.S.N.) By A. VerrIbu... 191
Proceedings of the Royal Society; Zoological Society ......... 197—220
On the Chilian “ Anguilla,” by Dr. R. A. Philippi; On the Parasitic
Nature of the Mistletoe, by Joseph Boehm ; On a Fungus which
is developed in Ivory and Bone, by Prof. Wedl; Note on the
Ammobroma Sonore; On the Intercellular Matter and the Ves-
sels of the Latex in the Root of the Dandelion, by Dr. A. Vogel;
On the Structure of the Luminous Organs in the Male of Lam-
pyris splendidula, by M. Schultze; De Jeude’s Collection of
ME OMI SCaiye «secs ce see neaeses sdecsenecccccnsne-saacenonetseee cere name ee
NUMBER XCIV.
XXVII. On Ammonites from the Cambridge Greensand. By
Harry Seecry, F.G.S., of the Woodwardian Museum, Cambridge.
CERES Pe NU) coco so hacen prs on che aceee meee nt ceemestbowcthinwast weeds seis ABO
XXVIII. Descriptions of New Genera and Species of Gallerucide.
PAY sir SESAME, Uaacarccosc orynmees sod dlc caeAeReeme eee staaes Pests raceaae ae
vi CONTENTS.
Page
XXIX. On the Australian Species of Paludina. By E. von
NUARRENS Mss Cav, Zo) sets epwecoone te eeee sonia ciioe «cic sence lcocemees 255
XXX. An Examination of the Dejeanian Genus Celomera (Coleo-
plera Phytophaga) and its Affinities. By the Rev. WaMLretr CLark,
Weer Lagan eas bestia ons yuemeonerageadtncassbevesiasanesseeesdeesiasesssanas deta 256
XXXI. On the Occurrence of Orcynus alalonga on the Coast of
Devons yeh Wokt. SC OMT... dy -csacacesevessteeacderaese-mases noesee saree 268
XXXII. Proofs of the Animal Nature of the Cilio-flagellate Infu-
soria, based upon Investigations of the Structure and Physiology of
one of the Peridinia (Peridinium cypripedium, n.sp.). By Prof. H.
JANES CLARK, AB. B.o.. GElate NEE:) cds cncswesenaescececescesnmeee 270
New Books :—Travels and Researches in Crete, by Capt. T. A. B.
Spratt, R.N., C.B., F.R.S. &e.—Handbook of British Water-
weeds or Algee, by Dr. John Edward Gray, F.R.S., late President
of the Botanical Society of London. The Diatomacex, by W.
Carruthers, F.L.S. &e.—Natural-History Transactions of North-
umberland and: Durham. Vol. I. Part 1. ....c.ccccscecccssses 279—282
Proceedings of the Zoological Society .........6++0+ sacisaiaisbelocatess 283—297
On Pristolepis marginatus, Jerd., by T. C. Jerdon, Esq.; On the
Constitution of the Fruit in the Cruciferae, by M. E. Fournier ;
On the Male Generative Organs of Phalangium, by Sir John
Lubbock, Bart., F.R.S.; On the Mode in which the Long-eared
Bat captures its Prey, by Wm. Sowerby; On the Habits of the
Water-Shrew (Crossopus fodiens), by N. L. Austen......... 298—302
NUMBER XCV.
XXXIII. On the Microscopie Structure of the Shell of Rhyncho-
nella Geinitziana. By Witutiam Bb. Carrenter, M.D., F.R.S.,
BLS GS tances deone vec nosaccns theca cles cecesacdceseecau cc eeenee nace enee 305
XXXIV. Contributions to an Insect Fauna of the Amazons Valley.
CoLEopTERA: Loncicornes. By H. W. Barss, Esgq.: ............ 308
XXXV. An Examination of the Dejeanian Genus Celomera (Coleo-
ptera Phytophaga) and its Affinities. By the Rev. HAMLET CLARK,
Me blame vonensve cosets POR eer Sonosda cat codpa crepao scence nes 315
XXXVI. On the Muscular Anatomy of the Leg of the Crocodile.
By the Rev. SamureL HaucGuron, M.D., Fellow of Trinity College,
Dublin. (Plate XVIz)) ...d..cccccccccovecmemmoeenemeccets sma acettness seneeees 26
CONTENTS. vil
Page
XXXVII. Observations on Raphides and other Crystals in Plants.
By Georce GuLutver, F.R.S. ...... Gauideloucaaseaneees aaa shaassrwanbuaeeas 331
XXXVIII. Note on the Cretaceous Deposits of Australia. By
Freperick M‘Coy, Professor of Natural Science in the University
of Melbourne, and Director of the National Museum of Victoria...... 333
XXXIX. On a new Growing Slide for the Microscope. By H. L.
SMITH, Kenyon College, U.S. .......sesssssssesesececeeserseesesnsccseseonns 334
XL. Descriptions of recently discovered Species, and Characters of
a new Genus, of Araneidea from the East of Central Africa. By
VOHINE DENG KevWAM olan Saisceianertescemeeneceeeecmcsecinenecdctiseesweoieaneiss 336
XLI. On two new Plesiosaurs from the Lias. By Harry SEELEY,
F.G.S., of the Woodwardian Museum, Cambridge. (Plates XIV. &
DNOVE)) Mitotnls occ seuatecssecccnsccrsossacs rans ssecctes SBidScapuanacnccoretosmccenqaGonWO0 352
New Books :—A History of British Ferns, by Edw. Newman. The
Fourth or School Edition.—Chart of Fossil Crustacea, by J. W.
Salter and H. Woodward. With Descriptive Catalogue.—Genera
Plantarum: auctoribus G. Bentham et J. D. Hooker. Vol. i.
PAYS eanceeerececeecessassssscnnsesnonees dccenscccsccanenccecccenes 359—364
Proceedings of the Zoological Society .......c.secesseeeeeeens sahamenessbites 365
Dr. Sturm’s Collection of Objects of Natural History in Nuremberg ;
On the Existence of Liquid and Solid Matters in the Trachean
Vessels of Plants, by M. T. Lestiboudois; On the Organization
of the Cypridine, by Prof. Claus; On the Anatomy of Tridacna
elongata, by M. Léon Vaillant ; Remarks on the Protective Sheath
and on the Formation of the Stem of the Root, by M. R. Caspary;
Graduation from “‘ Individual Peculiarities”’ to Species in Insects,
by Dr. B. D. Walsh; Note on the Cultivation of Eels, by M. L.
Soubeiran’ cc ccsacccasessenanececwsesdwencoess esiveeeleaneseee sone —oor:
NUMBER XCVI.
XLII. On the Systematic Value of the Organs which have been
employed as Fundamental Characters in the Classification of Mollusca.
By DrvOvACi; MORCH Giccstsssdebeceepcepsoeecuebesaaveenmvemmiededse.nJa0 385
XLIII. Description of four new Species of Butterflies in the Col-
lection of the British Museum. By A. G. Bur er, F.Z.S., Assistant,
Zoabgical Department, British Museumiss. <0. .c.:cccacesatesscccscccasenens SOP
XLIV. Remarks on Prof. H. J. Clark’s Peridinium cypripedium.
Byer des Caner, Wi: GUC: aces csiok a euoaap atest dacs tench cee nseseasvegeen 399
vill CONTENTS.
Page
XLV. Descriptions of new Genera and Species of Gallerucide.
Ppl Ss DAL, PLs.9. wenescescsaeewenuivsanecansadesesesesecsacantncsancwcess 402
XLVI, On the Limits of the Subkingdom Mollusca. By Dr. O.
Ads LU SURG) afer: aaron Gancoeannonuecnoabdaanedc ccrihoceddannonaosoncoccacesncnsaeng6 ww 411
XLVITI. Notes on the Paleozoic Bivalved Entomostraca. No. VI.
Some Silurian Species (Primitia). By Professor T. Rupert Jones,
£-G.s., and Or, BB. Hour, BGS... (Plate S00.) evesescceesasene 414
XLVIII. On the Names of the Genus Mystomys. (In a Letter to
Professor Allman.) By Dr. J. E. Gray, F.R.S., V.P.Z.S., F.L.S. &c. 425
XLIX. Descriptions of new Species of Shells. By E. von Mar-
PENT Sa VD cs sceascnascceteeateretaastessstcccnacednsshaeescanteesteasaces steno: 428
i. On Gliyniodon ornatus. By M;\SERRES) ...0..scssecsessosssocere 432
New Book :—British Conchology. Vol. III. Marine Shells, com-
prising the remaining Conchifera, the Solenoconchia, and Gas-
teropoda as far as Littorina, by JouN Gwyn Jerrreys, F.R.S.,
AGS ASECs. goes c<ictinaiiais deat olaiceleine aut Galine enjeetenneinee ces teceses dec aaaer stare 443
Capture of Muscicapa parva at Scilly, by E. H. Rodd; On the Canine
Teeth of Thylacoleo carnifex, Ow., by Prof. M‘Coy; On Afgeon
Alfordi, by the Rev. D. P. Alford; On the Nest of the Ten-spined
Stickleback, by W.H. Ransom, M.D. .......:s:ccsssecossoene 447— 451
AMGEN eacae ase cei cisseteccine sa conor cectoc secs coos temeltates tls dcnene seaeiee eee 452
PLATES IN VOL. XVI.
1. bo-orvignys Models of Foraminifera.
EET:
IV. New British Lichens.
V. New Genera and species of Diatoms.
VI. Buccal parts of Mollusca.
VII. Ageon Alfordi.
VII. Fossil Corals from South Australia.
IX. Fossil Entomostraca.
4h.
XI }New Ammonites from the Cambridge Greensand.
XII. Peridinium eypripedium.
XIII. Paleozoic Bivalved Entomostraca.
XIV. Plesiosaurus eleutheraxon.
XV. Plesiosaurus cliduchus.
XVI. Muscular Anatomy of the Leg of the Crocodile. -
THE ANNALS
MAGAZINE OF NATURAL HISTORY.
(THIRD SERIES. ]
eel eeseséunstaneseeen per litora spargite muscum,
Naiades, et circdm vitreos considite fontes :
Pollice virgineo teneros hic carpite flores :
Floribus et pictum, dive, replete canistrum.
At vos, 0 Nymphe Craterides, ite sub undas ;
Ite, recurvato variata corallia trunco
Vellite muscosis e rupibus, et mihi conchas
Ferte, Dez pelagi, et pingui conchylia succo.”’
N. Parthenii Giannettasii Ecl.1.
No. 91. JULY 1865.
I.—Descriptions of new Genera and Species of Diatoms from
Hongkong. By R. K. Grevitie, LL.D., F.R.S.E. &c.
[Plate V.]
PaLMERIA, nov. gen., Grev.
Frustules free, hemispherical, having an axis on the plane,
and longitudinal lines, indicative of fissiparous division, on the
convex surface; structure an exceedingly minute hexagonal
cellulation, appearing (except under a high magnifying power)
like a fine close moniliform striation, radiating from a subcentral
nucleus to the margin of the valve, along with a more distant
series of fine coste terminating in intramarginal punctiform
nodules.
This very singular Diatom cannot be satisfactorily referred to
any of the families defined in the most recent general arrange-
ment—that of Mr. Ralfs in the last edition (1861) of Pritchard’s
‘History of Infusoria.?’ With two of them, however (the Cym-
bellee and the Coscinodiscee), there appear to be some points
incommon. The form of the valve agrees with the first of these
families, and the frustule, as in Amphora, presents a ventral and
a dorsal aspect ; but there are no transverse striz, no longitudinal —
line, no nodules. With regard to the second family named, it is
excluded at once by the absence of the requisite disciform struc-
Ann. & Mag. N. Hist. Ser. 3, Vol. xvi.
2 Dr. R. K. Greville on new Genera and
ture and intermediate ring-like zone. Nevertheless the minute
organization is very similar ; for we have an hexagonal cellulation
radiating from a centre, along with a series of other sharply
defined lines, which may be called radiating coste, and which
terminate in submarginal nodules—the analogues of the lines
and minute spines in Coscinodiscus concinnus of Smith. But
our new Diatom seems, at first sight, to have a closer affinity
with Dr. Wallich’s curious genus Hemidiscus, which at present
is placed (I fear erroneously) among the Anguliferee. The
arcuate form, radiating cellulation, and submarginal puncta
certainly show an approximation between the two, Whatever
view, however, may be taken of their family relations, there are
differences which must keep them apart as genera—the chief of
which is the nodule on the ventral margin of Hemidiscus. The
submarginal punecta in that genus are also very different from
the minute nodules which terminate the radiating coste in Pal-
meria; and it is not unworthy of notice that the point of radia-
tion in the latter genus is never exactly centrical, being invariably
somewhat to the ventral side of the centre ; herein in Hemi-
discus it is truly centrical.
In a popular view, the frustule of Palmeria is graphically
compared by its discoverer to the half of a peeled orange, the
lines on the convex surface representing the segments of that
fruit.
In the living state, the endochrome is collected into minute
roundish masses, which dot the whole of the interior of the
siliceous covering. The nucleus in the valve is then very con-
spicuous ; and ‘T have a sketch, from Mr. Palmer, showing
radiating “reticulated cords,” similar to those described and
figured by Professor Max Schultze in Coscinodiscus centralis and
his Denticella regia (Mice. Journ. vol. vu. p. 13, pl. 2. figs. 11-18).
This appearance has also been observed, even in this country, in
specimens received from China, both of Palmeria and of a spe-
cies of Coscinodiscus, by my friend Mr. Laurence Hardman. The
peculiar current-like motions described by Prof. Max Schultze
have been witnessed likewise to some extent by Mr. Palmer,
especially the formation of a more compact mass of active parti-
cles, changing both its shape and position within the cell from
time to time. (Letter, June 1864.)
Palmeria Hardmaniana, n.sp., Grey. Pl. V. figs. 1-4.
Hab. Hongkong Harbour; John Linton Palmer, Esq., Surgeon
R.N., 1863.
A very large Diatom (varying, however, greatly in size), some
examples being as much as *0160" and upwards in diameter.
Species of Diatoms from Hongkong. 3
Frustules nearly, but not fully, hemispherical previous to self-
division ; on one side plane, with an axis through the middle
and a valve on each side; on the convex surface marked with a
middle longitudinal line and two fainter ones, indicating the
direction of future fissiparous division. The valves lunate, with
a straight ventral margin and obtuse ends; the surface covered
with very fine lines of hexagonal cellulation, radiating from a
blank nucleus or roundish vacant space, not situated exactly in
the centre, but a little towards the ventral side. Radiating from
the same point, and at more remote intervals, are also exceed-
ingly fine, but stronger, lines or delicate costee, which terminate
in a continuous row of punctiform nodules a little within the
margin of the whole circumference.
The first gatherings of Diatoms in Hongkong Harbour, sent
me by Mr. Palmer, contained some imperfect frustules of this
Diatom, which, along with his notes and sketches, greatly stimu-
lated my desire for more perfect materials. Fortunately my
friend Mr. L. Hardman had also received some collections from
the same locality; and, as they were rich in this production, he,
at my request, most kindly instituted a careful and minute
examination into its structure ; and it is only due to his friendly
aid and unrivalled manipulation to state that it is mainly from
his notes, sketches, and specimens that my drawings and de-
scriptions were prepared. My valued correspondent Mr. Palmer
subsequently forwarded a cell filled with frustules, preserved in
fluid, along with additional interesting information confirming
the results at which both Mr. Hardman and myself had arrived,
It is with sincere pleasure that I unite with the name of the
discoverer, now so worthily bestowed on the genus, that of the
gentleman who so materially aided me in the investigation of
its structure.
Palmeria Hardmaniana is a very delicate and fragile Diatom,
readily dividing (if arrived at the proper stage) at the slightest
touch—falling asunder as the segments of an orange are sepa-
rated, to use a simile already referred to. In size the frustules
appear to have a great range. Mr. Hardman has observed some,
however, which seem to belong to a second species, only half
the average size of the one now described, and much stronger
and thicker. As far as I can judge from a specimen he has
kindly given me, I am disposed to agree with him ; but it will
be well to examine a larger series of examples before coming to
a positive conclusion. Mr. Palmer informs me that he also has
observed frustules of a much smaller size; but, in the absence
of details, it is impossible to say whether they belong to the
supposed new species or not.
1*
A Dr. R. K. Greville on new Genera and
Genus ASTERIONELLA.
Asterionella Synedreformis, n. sp., Grev. Pl. V. figs. 5, 6.
Frustules exactly linear throughout, truncate, with two minute
apiculi at the apex; valve slightly dilated and rounded at the
base.
Hab. Hongkong Harbour; J. L. Palmer, Esq.
A very fine and distinct species, agreeing only with A. Ralfsii
im the frustules being exactly linear; but the latter is a fresh-
water species, having, besides, the valve of a different shape.
The present species is a large one, the frustules being sometimes
0080" and upwards in length. Towards the apex the margins
appear to be thickened, and terminate in two minute apiculi.
The valve is transversely striated, shortly acuminated at the
apex, becoming more slender below (as in other species), and
then slightly dilated at the base. The late Prof. Smith com-
mitted an oversight in resting the character of the genus partly
upon the frustules being “ inflated towards one or both extremi-
ties ;” for he defines one of his own three species (A. Ra/fsiz) as
“on F. V. exactly linear,” which they are. This is of little
consequence, as the main character depends upon the stellate
arrangement of the frustules, combined with the form of the
valve, which is different at the two extremities. Our new spe-
cies appears to be abundant at Hongkong. I have one specimen
in my cabinet in which only two frustules are absent from a
circle of fourteen.
Genus SuURIRELLA.
Surirella Palmeriana, n. sp., Grev. Pl. V. fig. 7.
Valve ovate-oblong, with obtuse ends ; ale inconspicuous ; costze
very slender, numerous, regular, terminating in a narrow me-
dian line, and having very delicate intermediate striz.
Hab. Hongkong Harbour ; in material communicated by J. L.
Palmer, Esq.
A very fine species, allied to S. gemma, but differing in im-
portant particulars, especially in the total absence of ale, and in
the canaliculi bemg equally distant. It is, besides, a much
larger species. Prof. Smith gives the length of S. gemma at
from 0025" to -0058"; that of S. Palmeriana is 0080". The
canaliculi are 4 in O01", and the intervening spaces are beauti-
fully arched or crenate at the margin. It appears to be exceed-
ingly rare, as Mr. Palmer has not referred to it in his letters,
Genus CRESWELLIA.
Creswellia annulata, n. sp., Grev. PI. V. fig. 8.
Minute; frustules short, cylindrical, truncate, transversely stri-
Species of Diatoms from Hongkong. 5
ated so as to appear annulate ; connecting processes very nu-
merous, very short, linear, obtuse.
Hab. Hongkong Harbour; in material communicated by J. L.
Palmer, Esq.
A fragile little species, seldom found quite entire, and gene-
rally in solitary frustules; but I have repeatedly seen two in
connexion, and have at the present moment a chain of three
under my eye. I have not been able to perceive any cellulation ;
but the annulate character is very obvious. The length of the
frustule is about -0010"; the breadth a little less.
I take the present opportunity of mentioning that in speci-
mens burnt on the cover, and mounted dry, I have observed
indications of an annulate structure in C. cylindracea, to which
our new species appears to be allied.
Genus HEMIAULUS.
Hemiaulus chinensis, n. sp.,Grev. Pl. V. fig. 9.
Frustules rectangular, very minutely punctate; the angles pro-
duced into linear-attenuated horns, tipped with a spine; space
between the horns nearly straight.
Hab. Hongkong Harbour; J. L. Palmer, Esq.
Like Hemiaulus Polycystinorum and others of the genus, the
present Diatom is most inconstant in point of size, not so much
in the length as in the breadth of the frustules. I have them
from ‘0007" to 0030". It is a delicate and transparent species,
and liable to be overlooked, unless burnt on the cover previous
to mounting. At the sutural line the puncta are exceedingly
minute, and gradually increase in size to the surface of the valve
included between the horns. I am not acquainted with any true
species of Hemiaulus in which the punctation is so minute. The
horns are somewhat truncate, and furnished with a terminal,
slender, short spine.
In his memorandum sketch, Mr. Palmer has represented the
valve, a view of which I have not been so fortunate as to obtain.
. Genus AMPHIPRORA.
Amphiprora lepida, n. sp., Grev. Pl. V. fig. 10.
Front view broadly linear, elongated, rounded at the ends, very
slightly constricted in the middle, and with a marginal row
of very minute puncta; middle portion grooved with straight
longitudinal lines.
Hab. Hongkong Harbour ; in material transmitted by Mr. J. L.
Palmer.
Well characterized by its narrow form, almost straight mar-
6 Dr. R. K. Greville on new Genera and
gins, and ribbed appearance of the connecting zone. I cannot
perceive any trace of strie or of supplementary wings. The
length of the frustule is 0065"; breadth in the middle -0010".
Amphiprora hyalina, nu. sp., Grev. Pl. V. fig. 11.
Hyaline ; front view divided, by a deep and sharp constriction,
into two roundish lobes, with a row of extremely minute
marginal puncta, and a second inner row of irregular ones ;
supplementary wings forming a single curve; connecting
zone with longitudinal lines.
Hab. Hongkong Harbour; in material communicated by J. L.
Palmer, Esq.
I offer this species with some little apprehension that it may
turn out to be an extraordinary variety or condition of A. alata.
Assuredly it appears to have absolutely nothing in common with
the figures given by Prof. Smith in his ‘Synopsis of the British
Diatomaceee ;? but it more nearly resembles Kiitzing’s figures in
his ‘ Bacillarien,’ which, although not quoted in Pritchard’s
‘Infusoria,’ are copied in the plates. These figures, however,
are somewhat confused, and, being deficient in details, cannot
be regarded as definitive. There is, if I may be allowed the
expression, a great difference in the physiognomy of the Chinese
Diatom. The supplementary wings also, which sweep down and
intersect, the lines of constriction, constitute an important cha-
racter. I have been unable to observe whether the wing be
continued round the ends, as is said to be the case in A. alata.
The marginal puncta are much more minute than in that spe-
cies ; and I cannot make out any striz. Length -0028", breadth
0018".
Amphiprora venusta, n. sp., Grev. PI. V. fig. 12.
Front view broadly panduriform, truncate at the ends; wings
widely rounded at the corners, and then rapidly constricted ;
supplementary wings narrow, passing in a single curve within
the nodule, striated; margin strong, with a row of minute
puncta.
Hab. Hongkong Harbour; in material communicated by J. L.
Palmer, Esq.
This species possesses a considerable likeness to A. alata
(Edin. New. Phil. Journ. vol. xviii. pl. 1. fig. 13), but differs in
its strong margin, in the row of marginal puncta, and in the
apparent absence of striz in the wings. In general outline it is
also very similar to A. Meneghiniana and A. Brébissoniana (loc. cit.
pl. 4. figs. 7, 8) ; but here the single curve of the supplementary
wings, besides other characters, removes it at once. In the dry
Species of Diatoms from Hongkong. 7
mounted slide the margin appears of a fine blue colour. Length
of frustule -0040".
Amphiprora chinensis, n. sp., Grev. Pl. V. fig. 13.
Front view transversely striated, oblong, rounded at the ends,
moderately constricted at the middle; wings flatly convex ;
supplementary wings narrow, with a marginal row of dark
points; connecting zone with a few narrow, ridge-like lines ;
striz minutely moniliform.
Hab. Hongkong Harbour; in material communicated by J. L.
Palmer, Esq. .
The frustules are sometimes twisted so as to throw a portion
of the wings out of focus; and the disposition to twist seems
almost always to cause more or less undulation in the rib-like
lines of the middle portion. The whole surface is transversely
striated, and the striz of the wings are clearly, although ex-
tremely minutely, moniliform. The flatness of the wing-curves
gives a degree of parallelism to the general outline, which is
very characteristic. The supplementary wings follow the curve
of the primary ones in a somewhat less degree, and pass down
within the nodules, having the appearance of an elevated ridge,
the ridge-line marked with a row of minute dark spots (scarcely
puncta), caused possibly by the intersection of the strie. Length
of frustule 0040" and upwards.
EXPLANATION OF PLATE V.
Fig. 1. Frustule of Palmeria Hardmaniana: front view, magnified 100
diams., showing the lines of future fissiparous division.
Fig. 2. The same, showing the valves.
Fig. 3. The same, showing the valves and frustule in the process of divi-
sion.
Fig. 4. Valve of the same, showing the radiating structure, magnified
150 diams.
Fig. 5. Asterionella Synedreformis, exhibiting two of the frustules and
the bases of others in situ.
Fig. 6. Valve of the same.
Fig. 7. Surirella Palmeriana.
8
Fig. 8. Cresswellia annulata: two frustules in situ, the valves of one of
them separating.
Fig. 9. Hemiaulus chinensis : two frustules in situ.
Fig. 10. Amphiprora lepida.
Fig. 11. as hyalina.
ig. 12. ne venusta.
Fig. 13. or chinensis.
[All the figures are magnified 400 diameters, except figs. 1 to 3, which
are magnified 100, and tig. 4, which is magnified 150 diameters. |
8 Rev. W. A. Leighton on British Lichens.
: II.— Notes on British Lichens.
By the Rev. W. A. Lricuton, B.A.; F.B.S.E.
[Plate IV.]
I purRrosg, in this and subsequent papers, from time to time to
present notes and illustrations of new or recently discovered
British Lichens, or such as have not been figured and described
im Sowerby’s ‘ English Botany’ and Supplement.
Gonrtonema, Nyl.
Thallus filiform; gonidial cells large, filled with granules, con-
catenated into a central axis. Apothecia biatorine or gyalec-
toid.
Gonionema velutinum, Ny]. Thallus dark brown, slender, en-
tangled ; apothecia dark brown, terminal, centre depressed, margin
swollen, pale within ; sporidia in asci 8, ellipsoid, simple, colour-
less ; paraphyses slender.
Gonionema velutinum, Nyl. Prodr. 16 (1857), Syn. 88, t. 1. fig. 11 (1858),
Scand. 23 (1861).
Collema velutinum, Ach. Syn. 329 (1814).
On the northern precipices of Craig Breidden, Montgomery-
shire, June 1864.
This minute lichen grows in scattered or continuous patches
on the face of the rock, and resembles in general appearance a
coarse dense velvety pile, of a blackish-brown colour. It con-
sists of minute, slender, cylindrical filaments, simple or branched,
erect, uniform in height, crowded and entangled into a czespitose
mass. When moistened and viewed under the microscope, these
filaments are found to consist of an outer fleshy or cartilaginous
continuous membrane, of a darkish-brown or olive-tawny colour,
within which is seen a central axis filling the entire external
cylindrical membrane, formed of large globular or spherical cells
concatenated in a moniliform manner, compressed longitudinally
by juxtaposition, and thus giving the cells a transverse dilata-
tion. On the external membrane being ruptured, the central
axis in longer or shorter lengths extrudes itself, and is then seen
to be of a pale dirty glaucous-green colour, and the cells to be
filled with very minute spherical granules, which, on the appli-
eation of diluted sulphuric acid, become of a reddish tinge. I
did not observe any apothecia on the Breidden specimens; but
the structure of the thallus in these so corresponds with an
authentic specimen in fructification, received from Dr. Nylander
himself, as to leave little or no doubt of their identity, notwith-
standing a slight difference in the width of the filaments, most
probably resulting from age and situation. The apothecia on
Rev. W. A. Leighton on British Lichens. 9
Dr. Nylander’s specimen are very minute, appressed, and sessile
on the upper extremities of the thalline filaments, and of a
similar colour, depressed or gyalectoid in the centre, and sur-
rounded with a thickish tumid margin, internally pale, and con-
sisting of narrow linear-oblong asci interspersed among very
slender paraphyses slightly swollen at the apices. Sporidia 8
in each ascus, ellipsoid, hyaline.
Dr. Nylander (/. c.) describes the spermogonia (which I have
not seen) as pale, globular or turbinate, and terminal ; spermatia
oblong, short; sterigmata slender. He also says, the hymeneal
gelatine becomes blue by the action of iodine, and finally of a
vinous red.
PLaTE IV. fig. 1. Portion of filament of thallus, magn. 330 times linear.
fig. 2. Central axis, magn. 330 times linear.
fig. 3. Cells of central axis, magn. 1200 times linear.
fig. 4. Asci and paraphyses, magnified 330 times linear.
fig. 5. Sporidia, magn. 1200 times linear.
fig. 6. Sterigmata and spermatia, after Nylander.
Sprtonema, Born.
Thallus filiform, branched, fruticulose; granula gonima large,
in transverse strata; apothecia lecideine, lenticular.
Spilonema paradoxum, Born. Thallus blackish brown, slender,
cespitose, entangled, branched ; apothecia black, terminal, hemi-
spherical, immarginate; hypothecium nigrescent; sporidia in
asci 8, oblong, simple, colourless; paraphyses thick, articulate.
Spilonema paradoxum, Bornet! Trois Lich. Nouv. p. 4, in Mém. Cherb.
Dec. 1856, tab. 1 & 2; Nyl.! Prodr. 17 (1857), Syn. 89, t. 2. f. 3 (1858),
‘Scand. 23 (1861); Leight.! Lich. Brit. Exs. 347 (1858); Mudd, Man.
35 (1861).
On rocks near the Harlech turnpike, at Barmouth, North
Wales, June 1856, in fructification.
Thallus forming larger or smaller, dense or scattered patches
of a black olive-brown colour, on the bare surface of granitic
rocks, presenting a dense czespitose velvety aspect. Filaments
of thallus erect, flexuose, and curved, entangled, irregularly and
somewhat secundly branched, about 3th of an inch in height.
The extremities of the branches, when moistened and viewed
under the microscope, are found to consist of a continuous outer
membrane, of an olive-tawny colour, within which the large
rounded or oblong gonidial cells are seen arranged in tolerably
regular transverse strata. The older stems exhibit the gonidia
more scattered and irregular, but still disposed in a distinctly
transverse direction, and immersed in a dense cellular tissue.
Apothecia terminal, minute, hemispherical, without any margin,
black; hypothecium nigrescent. Paraphyses short, thick, arti-
10 Rev. W. A. Leighton on British Lichens.
culate, the apical cell slightly enlarged and dark coloured.
Sporidia 8, im narrow asci, linear-oblong, simple, hyaline.
Spermogonia (which I have not seen), according to Bornet and
Nylander, lateral, tubercular, black ; arthrosterigmata articulate;
spermatia “ ovoides, fort petites ” (Bornet), “ breviter cylindrica”
(Nylander).
Even in a sterile state, this plant, which has a general resem-
blance to Ephebe pubescens, may be distinguished by attention
to the regular transverse arrangement of the gonidia, which are
also of much larger size than those of that plant, and altogether
differently grouped together. The outline of the filaments in
Spilonema is uniform, whilst in Ephebe it is crenate or wavy,
arising from the minute rugosities or tuberculations of the sur-
face corresponding with the internal strata of gonidia. I thought
at one time that there was also a chemical distinction,—dilute
sulphuric acid turning the old and young filaments of Spi/onema
to a dark-green colour, whilst in Hphebe the younger branches
were coloured green, and the older stems reddish; but this
character did not seem, after experiments on different specimens,
to be satisfactorily constant.
It is to be feared that, at least in some copies of my ‘ Lich.
Brit. Exsic.,’ specimens of Ephebe pubescens, which grew in the
same locality at Barmouth with Spzlonema paradoxum, have been
inadvertently inserted.
PuaTe IV. fig. 7. Extremity of younger branch, magn. 330 times linear.
fig. 8. Portion of older stem, magn. 330 times linear.
fig. 9. Sporidia, magn. 1200 times linear.
fig. 10. Paraphyses.
fig. 11. Sterigmata and spermatia, after Bornet and Nylander.
Eruese, Fr., Born.
Thallus filiform, branched, fruticulose ; granula gonima smaller,
subtransversely arranged, in little heaps, two, four, or more
together. Apothecia endocarpoid, in thickened portions of
the thallus.
Ephebe pubescens, Fr. Thallus blackish brown, slender, czes-
pitose, entangled, branched, slightly rugulose. Sporidia 8 in
asci, lineari-oblong or subfusiform, 1-septate, hyaline; para-
physes none.
Ephebe pubescens, Fr. S. O. V. 256 (1825); Bornet, in Ann. Se. Nat. sér. 3.
xviii. 170, t. 7; Nyl. Prod. 17, Syn. 90, t. 2. f. 1. & 17-20; Scand. 24;
L. P.1!; Moug. & Nestl. 358!; Heppe, 712!; Fellm. Lich. Lapp.
Or. 2!
Summit of Pen-Maen-Mawr, June 1851. Harlech turnpike,
at Barmouth, Caernarvonshire, June 1856. Rocks at Dartmoor,
Rey. W. A. Leighton on British Lichens. 1]
J. Ralfs, Esq.!_ Rocks at Coachford, west of Cork, J. Carroll,
Esq.!
This plant occurs in dense, entangled, decumbent masses,
loosely attached to the rocks. Filaments rather coarse, and of
a minute, tubercular, or scabrous appearance, dark brown, va-
riously and irregularly branched. When seen moistened under
the microscope, they are of an olive-brown colour; the granula
gonima in the young branches and extremities of the branches
appear transversely arranged, very similarly to those of Spilo-
nema paradoxum; but in the larger and older stems they are
more scattered and distant, and smaller in size, and are arranged
somewhat irregularly transversely, in heaps of several together.
The apothecia are immersed in subfusiform swollen portions of
the thallus, at a little distance from the extremities of the
branches, and are similar to those of Endocarpon, spherical,
with brownish perithecia. Paraphyses indistinct, mucilaginous.
Sporidia 8 in each ascus, oblong, elongated, shortly fusiform,
l-septate, hyaline. Spermogonia in lateral prominences ; sterig-
mata simple; spermatia straight, cylindrical.
For the unravelling of the synonymy, see Bornet, /. c.
PLATE IV. fig. 12. Portion of older stem, magn. 330 times linear.
fig. 13. Sporidia, magn. 1200 times linear.
fig. 14. Spermatia and sterigmata, after Nylander.
I possess a plant from Dr. Nylander, without any locality,
named Pilonema contextum, Nyl., which, from the general ap-
pearance and structure of the thallus, seems allied to the fore-
going. It grows in dense entangled czspitose masses, of a
blackish-brown colour. The filaments are very much branched,
and seem, when moistened under the microscope, to consist of
longitudinal series of small moniliform granula gonima immersed
in cellular tissue. There is no fructification on the specimen.
PuaTe IV. fig. 14. Portion of thallus, magn. 330 times linear.
fig. 15. Granula gonima, magn. 1200 times linear.
Racodium rupestre, Pers., of which I have a specimen from
Dr. Th. M. Fries, gathered at “Smolandia, Femsjo, 1851,”
and which I have gathered on rocks at Sychnant, near Conway,
and on the High Rock, Bridgenorth, Shropshire, and have also
specimens from Llandrindod (Rev. T. Salwey), Leicestershire
(Rev. A. Bloxam), and Cleveland (Mr. W. Mudd), and which,
according to a specimen received from Dr. Guthnich, from the
collections of Scherer, gathered at ‘“‘ Téte Noire,” and labelled
*Collema pannosum, Scher. Enum. p. 248,” would seem to be
included in that species by that author. Of this plant a fair
representation is given in Dillwyn’s ‘ Conferve,’ tab. 101, as
C. ebenea. Viewed under the microscope, it is found to consist
12 Rev. W. A. Leighton on the Gland of
of minute filaments indistinctly septate, over which is spread a
network of longitudinal fibres. No fructification has hitherto
been detected. There can be little doubt of the lichenoid nature
of this plant, the structure being similar to that of Canogonium.
(See Karsten’s Paper in Ann. Nat. Hist. ser. 8. vol. viii. p. 208,
pli.) .
Puate IV. fig. 16. Filament, magn. 330 times linear.
fig. 17. Filament, magn. 1200 times linear.
Chroolepus Arnottit, Hook., of which I have an authentic spe-
cimen gathered “ Kinross-shire, July 7, 1837,” approaches these
plants in external aspect ; but the microscope shows it to consist
of branched filaments of spherical cells, of a rich chocolate-
brown, tapering towards the extremities, where a distinct con-
Jugation may be seen.
Puate IV. fig. 18. Filament, magn. 330 times linear.
igo. oni iee at extremity of filament, magn. 330 times
near.
fig. 20. Conjugation, magnified 660 times linear.
Lichina pygmea, Ag. (Leight. Lich. Brit. Exs. 260) is beauti-
fully represented in Grev. Scott, Crypt. t. 219, and its micro-
scopic details in Tulasne’s Mém. Lich. tab. 9. figs. 1-6.
Mount’s Bay, Cornwall (J. Ralfs, Esq.!) may be recorded as
an additional habitat.
PuaTeE IV. fig. 21. Sporidium, magn. 1200 times linear.
Lichina confinis, Ag. This Lichen is also beautifully given
in Grey. Scott, Crypt. t. 221, and in Tulasne, /. c. tab. 10.
figs. 12-18.
Mount’s Bay, Cornwall (J. Ralfs, Hsq.!), and Black Stones,
Conway Bay, Caernarvonshire! June 1856, are additional
habitats.
PuLaTE IV. fig. 22. Sporidium, magn. 1200 times linear.
Pterygium centrifugum, Nyl. Syn. 92; Arnold, Lich. Juras.
Exs. 159, may probably be found on our limestone-rocks.
The scales on the Plate are the ;,\, >th of an inch, magn. 330, 660, and
1200 times linear.
I11.—On the Gland of the Phyllodium of Acacia magnifica.
By the Rev. W. A. Lerenton, B.A., F.B.S.E.
My attention has been attracted to a plant of Acacia magnifica
when in blossom. On the upper edge of the vertical phyllodia
(for the plant has no true leaves) subtending the showy spikes of
yellow flowers, which proceed from their axils, appeared a pellucid
drop of liquid, varying in size from that of a large pin’s head
the Phyllodium of Acacia magnifica. 13
to that of a grain of mustard-seed. This to the taste was sweet
and sugary. The flowers themselves had no odour, except to-
wards nightfall, when they gave out a weak disagreeable smell,
only perceptible on close contact. On wiping off the sugary
secretion, it was observed that it proceeded from a small sunken
linear-oblong orifice or slit, surrounded by a swollen margin.
The phyllodium itself is attached to the branch by a swollen
base, the surface of which is curiously marked by shallow rime,
alternately arranged, halfway round the base. From this swollen
portion the base tapers gradually, and becomes much narrower,
until, about a quarter or half an inch from the branch, the phyl-
lodium expands into a fusiform swelling, on the centre of which
the above linear-oblong orifice is situated. From this fusiform
swelling the phyllodium tapers to its uniform thickness. These
appearances are seen on looking down on tbe upper edge of
the phyllodium from above, and are represented in fig. 1,
Fig. 3.
LONE]
where a is the swollen rimose base, 6 the fusiform expansion
bearing the orifice. Fig. 2 represents a lateral view of the same,
where 4 is the situation of the glandular orifice, and c the large
bundles of vascular and spiral tissue, which proceed in a parallel
direction to the apex of the phyllodium. On making a vertical
section of this basal part of the phyllodium transversely through
the glandular orifice, the section, in a dry state, shows the ap-
pearances represented in fig. 3. Externally there is the bright-
yellow epidermis, with a layer of large cells immediately under-
neath, containing chlorophyll ; then similar large cellular tissue,
of a white colour and loose in texture; then a denser cellular
tissue of much smaller cells of a white colour, which is continued
towards the central slit of the gland, but becomes of a pale
yellow or slightly tawny colour, probably from the very minute
granular contents. Dilute sulphuric acid and weak solution of
14 Rev. W. A. Leighton on Acacia magnifica.
iodine produced no change of.colour in any of these parts. On
moistening the section with water, the external lips of the orifice
become swollen and partially closed, the slit alone being visible,
as seen, more highly magnified, in fig. 4 (where the same letters
indicate the same parts as in fig. 3, viz. d, epidermis; e, chloro-
phyll-cells ; f, loose white cellular tissue ; g, dense white cellular
tissue ; /, dense yellow cellular tissue ; 7, glandular slit; 4, bun-
dles of vascular and spiral vessels). Here it is seen that the
epidermis ceases somewhat above the base of the slit, where
apparently the cellular tissue is exposed, and from which surface
the pellucid liquid is excreted.
The plant began to blossom on the 27th of March, and was
then removed from the green-house into the drawing-room,
where the secretion immediately attracted my attention. I my-
self watered the plant every morning ; and thus it was daily, and,
indeed, many times every day, under my constant observation ;
and the secretion was pointed out to members of my family and
to many friends almost daily. As I proposed to investigate the
source and cause of the secretion with the microscope, I carefully
watched it day by day, and am thus able to state definitely that
the liquid drop was visible on the upper edge of every phyllodium
subtending a spike of flowers during the whole time the plant
continued in flower, viz. from March 27 to April 22. For a few
days previous to April 22, the secretion appeared to decrease
and partially to cease on some of the phyllodia. On April 23
the blossoms began to wither and fall. On the 24th the blos-
soms fell more rapidly and abundantly; and, to my surprise,
there was an almost total cessation of the secretion, which now
appeared on a very few only of the phyllodia. On the 24th the
plant was returned to the greenhouse, and since that day to the
present time (May 30), although the plant has been watched
carefully for this express purpose, not the least secretion has
taken place, and the orifice of the gland appears to have become
partially filled up or obliterated.
Here observation ends; but, on beholding such a curious
structure, the mind naturally speculates—but in vain— How is
this secretion effected ? Nature does not disclose her vital forces.
We then turn to the probable end to be effected by such a pro-
vision; and here conjecture may be possibly more successful.
The secretion takes place oniy during the period that the plant
is in blossom. So soon as the flowers fade and begin to fall,
the secretion ceases and disappears. It would seem then to be
in some way or other connected with the fertilization of the
flower; and as, when the secretion becomes excessive, it falls
and blotches the lateral expansion of the phyllodium, it is pro-
bably to attract insects to effect this. It is right, however, to
On the Nomenclature of the Foraminifera. 15
confess that no insects were observed to alight on the plant;
but this may be owing in some measure to the early season of
the year at which the plant blooms in this country, or to its
having been taken from the green-house into a drawing-room,
where the windows were generally closed ; or, what is still more
probable, that British insects are not the same as Australian, and
have not the same habits ; for it seems almost evident that it would
require an insect of some considerable size and of some peculiar
structure and habits to remove and apply the pollen, the secre-
tion not being in the blossom itself, but at a short distance from
it, on the phyllodium.
However, none of the flowers were fertilized; but it was re-
marked that the styles became elongated to nearly double the
length of the stamens, particularly towards the time of the fading
and falling of the blossoms. The thought readily arises, Is this
another imstance of dimorphism? and is there another plant,
with short-styled stigmas, or with some other peculiar structure,
adapted and necessary for the perfect fertilization? This, future
and further observation may verify; but it appears highly sug-
gestive of a fine field of research to those who possess or have
access to large collections of Acacieg. The fact of some Acacie
fruiting abundantly in greenhouses, and others rarely or never,
has often attracted attention; and artificial fertilization would
do much towards ascertaining whether it is to the absence of
insect agency that the sterility of the plants is due.
An intelligent nurseryman here informs me that he has never
observed the plant to form legumes, or, at all events, other than
abortive ones. He says the plant was originally raised at Ghent,
from seed from Australia, and that that place is the great mart
where it is propagated by cuttings, and imported into this
country.
The microscopist will find the stamens, and indeed every por-
tion of the floral whorls, beautiful and interesting objects, as,
from their extreme transparency, the cellular tissue and the
spiral vessels are distinctly displayed, without any dissection or
other preparation than being placed in a drop of water.
IV.—On the Nomenclature of the Foraminifera. By W. K.
Parker, F.R.S., T. Rupert Jonss, F.G.S., and H. B. Brapy,
Holly Ube [Plates I., II., III.]
Part X. (continued).—The Species enumerated by D’ Orbigny in
the ‘ Annales des Sciences Naturelles, vol. vii. 1826,
Ill. The Species illustrated by Models.
Previous_y to the publication of his “ Tableau Méthodique des
16 Messrs. Parker, Jones, and Brady on the
Céphalopodes”’ in the ‘ Annales des Sciences Naturelles,’ vol. vii.
1826, D’Orbigny had prepared and published, in part at least,
a hundred models of Foraminifera (at that time regarded as
microscopic Cephalopods), illustrating many of the species for
the first time. These models were made in plaster of Paris,
were about an inch or more in length, and were issued in sets
of twenty-five, arranged in suitable boxes, each box bearing
a label as follows* :—‘ Models of microscopic Cephalopods,
recent and fossil, representing one individual of each of the
principal divisions of a new method based on the mode of
growth of the shells. The Models are from forty to two
hundred times the size of the originals, so as to show their cha-
racters distinctly. By M. Alcide Dessalines D’Orbigny, junior.
There are Four Fasciculi, each comprising twenty-five Models ;
besides, for the first sixty subscribers, three or four shells.
The great rarity of the originals does not allow any more to be
promised. (The specimens are in glass boxes, which must be
opened with great care.) The four Fasciculi will be issued in
the course of the first six months of 1823: the price of each is
twenty francs, payable either at Rochelle to the author (Jardin
des Capucins), or at Paris to M. ———. Letters and money
to be post-free. The First Fasciculus of the Models may be
seen at Paris, at the Museum of Natural History of the Jardin
du Roi, and at M. ————’s._ The subscribers will receive with
* “Modéles de céphalopodes microscopiques, vivans et fossiles, représen-
tant un individu seulement de chacune des principales divisions d’une
nouvelle méthode, basée sur le mode d’accroissement des coquilles. Le
diamétre de ces modéles a été porté de 40 jusqu’a 200 fois celui des
coquilles originales, afin de rendre plus sensibles tous leurs caractéres.
“Par M. Alcide Dessalines D’Orbigny, fils.
‘La Souscription se composera de 4 Livraisons, qui comprendront
chacune 25 Modeéles, et en outre 3 a 4 coquilles pour les 60 premiers
souscripteurs: l’extréme rareté des originaux ne permet pas maintenant
d’en promettre davantage. (Ils sont ici renfermés dans les boites de verre,
qu’on ne doit ouvrir qu’avec la plus grande précaution.)
** Les quatre Livraisons seront expédiées dans le cours des 6 premiers
mois de l’année 1823; le prix de chacune d’elles est 20 frances, et sera
payé, soit a la Rochelle, chez auteur (Jardin des Capucins), soit a Paris,
chez M. , en affranchissant les lettres et largent.
“On peut voir la premiére Livraison de ces Modeles, a Paris, au Muséum
d’ Histoire Naturelle du Jardin du Roi, et chez M. . Les souscripteurs
recevront, avec la quatriéme Livraison, le tableau méthodique de la distri-
bution de ces Céphalopodes, qui indiquera, par numéros correspondant a
ceux des Modéles, le nom des individus envoyés et l’ordre de leur classifi-
cation.”
sé Ame T ivraison.
** Nota.—Les Modeéeles coloriés représentent les coquilles fossiles ; et les
blanes, les vivans. Le leu et la forme des syphons y sont indiqués par
des traces ou par des points noirs.”
Nomenclature of the Foraminifera. 17
the Fourth Fasciculus the Systematic Table of the Distribution
of these Cephalopods, indicating, by numbers corresponding to
those of the Models, the names of the specimens sent, and the
order of their classification.”
With the Fourth Fasciculus the following note was issued :—
“The coloured Models represent the fossil shells ; the white
Models, the recent shells. The place and shape of the siphuncles
are indicated by the marks or black spots.”
In his Introduction to M. D’Orbigny’s memoir in the same
volume of the ‘Annales des Sciences Naturelles,’ p. 99, M. Feé-
russac says that two of the Fasciculi of Models had been pub-
lished, and that the other two would soon follow. The Models
therefore were issued, partly, before November 7th, 1825, when
the Memoir was presented to the Academy of Sciences ; and
though we do not know the exact date of the publication of the
third and fourth sets, we shall here regard them as belonging
to about the same period (1825-26).
M. D’Orbigny’s researches on Foraminifera appear to have
arisen from his father’s attention having been directed to these
Microzoa; for the elder D’Orbigny, who was a physician at
Esnaudes, near Rochelle, wrote to M. Fleurian de Bellevue, in
1819, on his discovering, on the shores of the Atlantic, micro-
scopic Cephalopods, among which he had seen “living Lenti-
culines, Rotalies, Discorbes, Spirolines, &c.” (See Annales de
Physique, vol. xxxviil. p. 187.) The younger D’Orbigny en-
larged his knowledge of these little shells by the collection of
numerous samples of sea-sand and of fossiliferous deposits from
various parts of the world, working perseveringly and methodi-
cally for several years, reducing already published notices and
his own observations to a system, which, though artificial
throughout and otherwise defective, was very useful ; he described
the general characters and external features with care, illus-
trating his descriptions by the Models now under consideration
and the elaborate plates of his noble quarto and folio volumes
on the Foraminifera, and leaving in the end an extensive collec-
tion of material. His several works and their numerous illus-
trations, chiefly relating to the larger specimens which had come
under his notice, are necessarily the groundwork for writers on
Foraminifera up to the present time.
We will now proceed to the examination of the Models
seriatim, in the order in which they were originally numbered
by D’Orbigny. A series of carefully prepared outlmes appended
to this paper (Plates I., II., & III.) will assist in forming a cor-
rect appreciation of our remarks. In these plates the forms
represented by the Models are grouped, as far as may be, ac-
cording to their respective families: —the Minioxipa (Vertebra-
Amn. & Mag. N. Hist. Ser.3. Vol. xvi. 2
18 Messrs. Parker, Jones, and y ady on the
lina, Pl. I. figs. 1, 2; Miliola, figs. 3-15; Fuabularia, fig. 16 ;
Peneroplis, figs. 17-20; Orbiculina, figs. 21, 22; Alveolina,
fig. 23); Lirvonipa (Valvulina, figs. 24-26) ; Lacenipa (No-
dosarina, figs. 27-46 ; Polymorphina, P\. II. figs. 47-53; Uvi-
gerina, fig. 54); Guosicerinipa (Globigerina, figs. 55, 56;
Pullenia, fig. 57; Spheroidina, fig. 58; Teatularia, figs. 59—
63; Bulimina, figs. 64-66; Cassidulina, fig. 67; Discorbina,
figs. 68-71; Planorbulina, figs. 73-77, and PI. III. figs. 78, 79 ;
Pulvinulina, P|. III. figs. 80-82; Rotalia, figs. 88-86 ; Calca-
rina, figs. 87-90); Nummuxtnipa (Amplhistegina, figs. 91, 92 ;
Operculina, figs. 93, 94; Nummulina, fig. 95; Polystomella,
fig. 96; Nonionina, figs. 97-99 ; Heterostegina, fig. 100).
Livratson 1',
Model no. 1. Nodosaria Radicula*, Linn. sp. Annales des
Sciencest, vol. vii. 1826, p. 252, no. 3.
Hab, Adriatic. Pl. I. fig. 27.
The common straight Nodosarian form, with few globose
chambers, smooth and free from any surface-markings.
Model no. 2. Nodosaria (Orthocerina) Clavulus*, Lamk. sp.
Page 255. no. 48.
Hab. Fossil near Paris. PI. I. fig. 25.
A Clavuline variety of Valvulina triangularis. The term
Orthocerina was applied by D’Orbigny in 1839 (Foram. Cuba,
p- 18 of 8vo edition, p. 47 of 4to ed.) to a Foraminifer really
related to Nodosaria, namely, O. quadrilatera (For. Cuba, pl. 1.
figs. 11, 12). An Orthocerina which was described and figured
by Reuss as Triplasia (and afterwards Rhabdogonium) Murchisont
is perhaps the best type of this somewhat peculiar genus. Al-
though Orthocerina is evidently one of the Lagenide, allied to
both Nodosaria and Uvigerina, it must be allowed to stand
apart. (See Carpenter’s Introd. Foram, p. 166.)
Prof. Reuss, in describing the three-sided Orthocerina above
alluded to, from fossil specimens found in the Cretaceous rocks
of the Eastern Alps, made use of the generic term Triplasia for
it (Sitzungsber. Akad. Wiss. Wien, vii. 1854), and subsequently
substituted Rhabdogonium as a denomination for these three- or
four-sided Orthocerine (ibid. 1860). D’Orbigny’s subgeneric
term is misapplied to the Model under consideration, yet it was
evidently used in 1839 on the plan that he originally intended ;
* The species and varieties marked by an asterisk have been already
noticed by us in former papers.
+ The references throughout are to D’Orbigny’s paper im the ‘ Annales
des Sciences Naturelles,’ ser. 1. vol. vii. 1826,
Nomenclature of the Foraminifera, 19
and in the Cuban figures we have examples of the type of the
genus, though in a somewhat exceptional condition,
Model no. 3. Frondicularia rhomboidalis, D’Orb. Page 256, no. 1.
Hab. Adriatic. Pl. I. fig. 31.
A variety of Frondicularia complanata, Defr. The lozenge-
shaped form of Frondicularia in which the lateral wings of the
later chambers reach back only about half the length of the
shell,
Model no. 4. Vaginulina tricarinata, D’Orb. Page 258, no. 4.
Hab. Adriatic. Pl. I. fig. 34.
A three-sided Nodosaria, with somewhat oblique chambers,
and having a keel along each of the three edges. A similarly
modified form is found fossil at Baden, near Vienna, and is de-
scribed and figured by Dr. Karrer as Rhabdogonium pyramidale,
Kar., in his valuable paper on the distribution of the fossil
Foraminifera of the Vienna Basin (Sitzungsber. Math.-nat.
Class. K. Akad. Wiss. vol. xliv. 1861, p. 20, pl. 1. fig. 5.
We have met with V. ¢ricarinata recent im Mediterranean
sands, in which, however, it is very rare. It also occurs in a
fossil condition, sparingly, in Tertiary clay from near Malaga,
and in the Subapennine Tertiary shell-sands.
Model no. 5. Nodosaria (Dentalina) obliqua, D’Orb.
Page 254, no. 36.
Hab. Adriatic. PI. I. fig. 32.
A Dentaline Nodosaria, with broad and oblique chambers, the
aperture being terminal, somewhat towards the convex side of
the slightly curved shell.
This is a common form, both recent and fossil, and is scarcely
separable from D. communis, D’Orb.
Model no. 6. Marginulina Raphanus*, Linn. sp. Page 258, no. 1,
pl lO. tgs. 7, o>
Hab. Adriatic ; fossil at Castel-Arquato, Italy. PI. I. fig. 35.
The Marginuline condition of Nodosarina Raphanus, and the
best type of the group. The straight N. Raphanus is the N.
Rapa of D’Orbigny.
Model no. 7. Textularia pygmea*, D’Orb. Page 263, no. 18.
Hab. Adriatic. PI. II. fig. 59.
Another small variety of Tevtularia aggluiinans, subsequently
described, no. 15 (p. 263), and named 7. aciculata, is identical
with this, and must be included under the same name.
2%
20 Messrs. Parker, Jones, and Brady on the
Model no. 8. Quinqueloculina Lyra, D’Orb. Page 303, no. 45.
Hab. Adriatic and Mediterranean. PI. I. fig. 11.
A thin, narrowish Miliola, with subsigmoid, somewhat cari-
nate chambers.
Model no. 9. Bulimina elegans, D’?Orb. Page 270, no. 10.
Hab. Adriatic, near Rimini. PI. II. fig. 64.
Bulimina Preslii, Reuss, figured in the Verst. Kreid. Bohm?
(1846), pl. 13. fig. 72, and in Haidinger’s ‘Naturwiss. Abhand,
vol. iv. (1850), Kreidemergels von Lemberg, pl. 3. fig. 10, is the
best type of the genus Bulimina. B. eles gans 18 a “delicate va-
riety, narrower and less robust in growth, as well as more im-
bricated in the disposition of the chambers.
Model no. 10. Rotalia Menardii, D’?Orb. Page 273, no. 26.
Hab. Adriatic, near Rimini. PI. III. fig. 81
This must be placed in the genus Pulvinulina, being a good
subspecies of P. repanda. It is found in deep water, and is
seldom abundant, except in Tropical seas. One or two beautiful
specimens have occurred to us on our own coast, in sand dredged
from deep water off the Isle of Man.
Model no. 11. Nonionina Limba, D’Orb. Page 594, no. 14.
Hab. Adriatic, near Rimini. PI. III. fig. 99.
A variety of the subtype N. asterizans, ¥. & M., from which
it ‘differs in the greater development of the stellate sutural
limbation, and in possessing a narrow thick keel, instead of the
rounded edge of the subtype. The same variety, but with
stouter and somewhat encrusted shell, occurs fossil in the
neighbourhood of Bordeaux.
Model no. 12. Rotalia punctulata*, D’Orb. Page 273, no. 25.
Hab. Adriatic, near Rimini. PI. III. fig. 82
This is a Pulvinulina, and not a Rotalia; it is perhaps even
a more fully developed form than the type, P. repanda, F. & M.
We have found fine handsome specimens in dredgings from the
coast of Norway.
Model no. 18. Gyroidina orbicularis, D’Orb. Page 278, no. 1.
Hab. Adriatic, near Rimini. PI. IIT. fig. 85.
The thin-shelled, somewhat globular variety of Rotalia Bec-
cart, inhabiting deepish water, usually much smaller than the
common shallow-water form. It is a widely distributed variety,
and has been found on our own coast, in the Irish Sea and off
the Shetland Islands.
Nomenclature of the Foraminifera. 21
Model no. 14. Robulina virgata, D’Orb. Page 290, no. 17.
Hab. Adriatic, near Rimini. PI. I. fig. 40.
An umbonate Cristellaria, with few chambers, and destitute
of any keel. Thick square bands of shell-substance, covering
the partition-walls of the chamber, radiate in straight lines from
the central umbo to the margin.
Model no. 15. Rotala bisaculeata, D’?Orb. Page 273, no. 20.
Hab. From ballast-sand. Pl. III. fig. 89.
This is rather a subvariety of the Rotaline genus Calcarina,
of which Model no. 84 may be taken as the type. It does not
differ very greatly from Deshayes’s Calcarina rarispina (Lyell’s
Manual, Sth edit. p. 228, fig. 286), but sufficiently, however, to
render a separate trivial name convenient. Its peculiarity con-
sists In possessing a keel, extended into double points at inter-
vals round the margin.
Model no. 16. Peneroplis planatus*, F. & M., sp.
Page 285, no. 1.
Hab. Mediterranean, New Holland (Rawack). PI. I. fig. 17.
This is the well-known widely distributed type, for which, as
we have before stated, Forskal’s specific name Peneroplis per-
tusus takes precedence (Ann. Nat. Hist. March 1865).
Model no. 17. Globigerina bulloides, D’Orb. (young).
Page 277, no. l.
Hab. Adriatic, near Rimini. PI. II. fig. 56.
This is the young shell. See note on the adult, Model 76.
Model no. 18. Adelosina striata, D’Orb. (young).
Page 304, no. 2.
Hab. Fossil at Castel-Arquato, Italy. PI. I. fig. 14.
It has been satisfactorily demonstrated that the delicate Mz-
liole, with retort-shaped chambers, grouped by D’Orbigny
under the generic name Adelosina, are only young specimens of
other well-known species. The Model now under consideration
represents the very young condition of Quinqueloculina Brong-
niartii ; and the Model 97, which is given as the adult of the same
species, 1s only Q. Brongniarti somewhat further developed.
Model no. 19. Cristellaria (Saracenaria) Italica*, Defr. (young).
Page 293, no. 26.
Hab. Adriatic, near Rimini; and fossil near Sienna. PI. I,
fig. 42.
The young condition of the shell. See note on the adult,
Model no. 85.
22 Messrs. Parker, Jones, and Brady on the
Model no. 20. Orbiculina numismalis*, Lamk. Page 805, no. 1.
Hab. The Antilles and the Marianne Isles. PI. I. fig. 21.
Though he adopts this form as the type of the genus Orbicu-
lina, D’Orbigny rightly associates the three forms catalogued by
Lamarck as O. uncinata (adult), O. numismalis (middle-aged),
and O. angulata (young) as the same species in different stages
of growth. O. orbiculus, F. & M., termed O. nummata by La-
marck, should also be included amongst the middle-aged forms.
Subsequently D’Orbigny took the adult form, with the well-
known name “ adunca,” originally conferred on it by Fichtel
and Moll, as the central type; and we find this figured in the
“Cuba” Monograph, pl. 8. figs. 8-16. On the same plate
(figs. 4-7) we find another adult form of Orbiculina, thin, flat,
and orbicular, with the name of O. compressa.
Model no. 21. Dendritina Arbuscula*, D’Orb. Page 285,
no. ly ply lo. figs, 7.
Hab. Fossil near Bordeaux. PI. I. fig. 20.
The thick robust variety of Peneroplis pertusus, with a single
large aperture running into irregular dendritic ramifications.
Found recent in tropical seas.
Model no. 22. Articulina nitida*, D’Orb. Page 800, no. 1.
Hab. Fossil near Paris. PI. I. fig. 2.
An elongated subcylindrical variety of Vertebralina striata,
previously figured and described by Batsch as Nautilus conico-
articulatus. It is an exceedingly variable form, and may be
found im every gradation, from the narrow subcylindrical con-
dition represented in the Model to the broad Renulites opercu-
laria of Lamarck. The specimens taken from the abyssal depths
of the Mediterranean and the deeper portions of the Red Sea
are characteristically small. In the Eocene marl of Baljik, in
Bulgaria, small and still more elongated specimens occur.
Model no. 28. Polymorphina Thouini, D’Orb. Page 265, no. 8.
Hab. Fossil near Paris. PI. IT. fig. 49.
A long narrow variety of P. lactea, W. & J.
Model no. 24. Spirolina cylindracea*, Lamk. Page 286, no. 1.
Hab. Fossil near Paris. Pl. I. fig. 19.
The crozier-like, deep-water form of Peneroplis pertusus, com-
mon in the Grignon Tertiaries, and found recent in the deeper
seas of tropical climates. Notwithstanding the identity with
Peneroplis, the distinctive name Spirolina is convenient, if not
necessary, in separating the long subcylindrical varieties from
the outspread forms found in shallow water.
Nomenclature of the Foraminifera. 23
quae no. 25. ‘Valvulina triangularis*, D’Orb.
Page 270, no. 1.
Hab. Fossil near Paris PI. I. fig. 24.
This is the arenaceou- triserial, three-sided shell which may
be looked upon as the ty ¢ of a very variable group. The two
Clavuline varieties which e represented by Models nos. 2 and
66 show the wide rap.s* on one side; and Prof. Williamson’s
Rotalina fusca (Valvulina Austriaca), Monogr aph, pl. 5. figs. 114,
115, exhibits the opposite extreme in contour.
LivrAIson 2™,
Model no. 26. Lingulina carinata, D’Orb. Page 257, no. 1.
Hab. Antilles; and fossil near Sienna. PI. I. fig. 28.
The compressed, sharp-edged, straight subtype of Nodosarina,
common as a fossil in the Lias and in many Tertiary clays, but
rare in arecent condition. In D’Orbigny’s ‘ Cuba” it is men-
tioned as occurring in the West-Indian seas and in the neigh-
bourhood of the Canaries. It is figured in Williamson’s Mono-
graph (pl. 2. figs. 33, 34), and noted as having been found recent
in three British localities. Dr. Carpenter has it very large from
Japan.
Model no. 27. Planularia Cymba*, D’Orb. Page 260, no. 4,
pl. 10. fig. 9.
Hab. Adriatic. PI. I. fig. 38.
This has been already noticed by us amongst the forms illus-
trated by figures in D’Orbigny’s Memoir.
Model no. 28. Textularia gibbosa*, D’Orb. Page 262, no. 6.
Hab. Mediterranean; and fossil at Castel-Arquato, Italy.
Ply UT. fies 60:
This is one of the largest forms of Textularia, with ventricose
segments, and is found widely distributed. In common with
the other bold varieties, it is at home and most luxuriant in
water from 50 to 100 fathoms.
Model no. 29. Polymorphina Burdigalensis, D’Orb.
Page 265, no. 2.
Hab. Fossil near Bordeaux. PI. II. fig. 48.
A somewhat unsymmetrical Polymorphina, having one side
flattened, and consequently an excentric aperture. The cham-
bers are. numerous, and are compactly put together, the edges
overlapping and giving a general even outline to the shell.
24 Messrs. Parker, Jones, and Brady on the
Model no. 30. Polymorphina (Pyrulina) Gutta*, D’Orb.
Page 267, no. 28, pl. 12. figs. 5, 6.
Hab. Fossil, Castel-Arquato. Pl. II. fig. 51. *
Noticed in our previous paper on the species figured by
D’Orbigny.
Model no. 31. Biloculina aculeata, D’Orb. Page 298, no. 3.
Hab. Fossil in the rocks of Pauliac (Gironde). PI. I. fig. 5.
This seems to be an unusual and somewhat monstrous 57-
loculina, with a number of somewhat pointed prominences,
arranged in two rows on opposite sides of the outer chamber.
It may be regarded as a subvariety of B. ringens, which it closely
resembles in general contour.
Model no. 82. Quinqueloculina Ferussacti, D’Orb.
Page 301, no. 18.
Hab. Fossil near Paris. Pl. I. fig. 12.
A narrow, elongated, somewhat angular Quinqueloculina,
having a few stout cost traversing the chambers from end to
end. Wilhamson’s Miliolina bicornis var. angulata (Monogr.
pl. 7. fig. 196) may be assigned to this species.
Model no. 38. Quinqueloculina Saxorum*, Lamk.
Page 301, no. 1.
Hab. Fossil near Paris. Pl. I. fig. 13.
This is a well-known form, not uncommon in a recent con-
dition on the coral-reefs of tropical seas, and very abundant in
the Calcaire grossier near Paris.
Model no. 34. Calcarina Calcar*, D’Orb. Page 276, no. 1.
Hab. Martinique, Isle of France, Madagascar. PI. III. fig. 87.
This cannot be separated from Calcarina Spengleri, Linn.
The list of localities might be much extended, as there are few
tropical seas in which specimens do not abound.
Model no. 35. Rotalia rosea, D’Orb. Page 272, no. 7.
Hab. The Antilles, the Isle of Martinique, Point Carbet.
Behe. 79.
This belongs to Planorbulina, not to Rotalia (restricted), and
is nearly allied to Planorbulina Haidingeriit. Its pink colour
is very characteristic ; and its tolerably limited distribution gives
it additional claims for distinctive appellation. Coloured figures
are given in the “Cuba” Monograph, pl. 3. figs. 9,11. Small
specimens are not uncommon in West-Indian sponge-sand.
ri)
Or
Nomenclature of the Foraminifera.
Model no. 36. Gyroidina Soldani, D’Orb. Page 278, no. 5
Hab. Adriatic, near Rimini. PI. III. fig. 86.
As we have before stated, there is no sufficient reason for the
separation of either this or G. orbicularis from the genus Rotalia.
It is a small deep-sea variety, somewhat bolder, ‘and having a
thicker shell than G. orbicularis.
Model no. 37. Truncatulina tuberculata*, D’Ovb.
Page 279, no. 1.
Hab. All the shores of European seas. Fossil near Bordeaux,
Paris, and Castel-Arquato. PI. II. fig. 77.
This is Truncatulina lobatula, W. & J.,sp., which is the com-
monest variety of Planorbulina farcta, ¥. & M., sp.
Model no. 88. Rosalina Parisiensis, D’Orb. Page 271, no. 1.
Hab. Fossil near Paris. PI. II. fig. 70.
This is a flat, outspread, thin- edged variety of Discorbina
trochidiformis, Lamk. , Sp.; a good subspecies—the type being
D. Turbo, D’Orb., sp. It is commonly somewhat concave on its
under surface. Discorbina ochracea (Rotalina ochracea, Will.
Rec. For. Gt. Brit. pl. 4. fig. 112, and pl. 5. fig. 113) is a feebler
form, closely allied to the one under adie
Model no. 89. Rotalia rosacea, D’Orb. Page 273, no. 15.
Hab. Fossil, Bordeaux. PI. I. fig. 71.
This is another useful subspecies of Discorbina Turbo, D’Orb.,
sp. It is identical with D’Orbigny’s Asterigerina Planorbis and
Williamson’s Rotalina Mamilla.
Model no. 40. Amphistegina vulgaris, D’Orb. Page 805, no. 8.
Hab. Fossil on the borders of the Lagoon of Tau, and near
Bordeaux. PI. III. fig. 91.
The type of the genus.
Model no. 41. Casstdulina levigata, D’ Orb. Page 281, no. 1,
pl. 15. figs. 4, 5
Hab. From ballast-sand. PI. II. a 67.
One of the species figured by D’Orbigny i in the plates illus-
trating his memoir, and), as such, noticed in our previous paper.
Model no. 42. Anomalina elegans, D’Orb. Page 282, no. 4.
Hab. Fossil near Bordeaux. PI. II. fig. 73.
The generic term Anomalina should properly be confined to
the bold and more or less biconcave forms of the Planorbuline
type. This Model illustrates Discorbina,—a good intermediate
subspecies, passing insensibly into D, vesicularis. The same
26 Messrs. Parker, Jones, and Brady on the
form is figured by D’Orbigny, in the ‘ For. Foss. Vien.’ pl. 10.
figs. 13-15, under the name of Rotala complanata.
Model no. 43. Nonionina sphaeroides, D’Orb. Page 293, no. 1.
Hab. From ballast-sand. . Pl. II. fig. 57.
This is Pullenia spheroides, and bears no relation to the No-
nionine subtype. See Carpenter’s ‘ Introduction,’ p. 184, It is
figured in D’Orbigny’s ‘ For. Foss. Vien.’ pl. 5. figs. 8-10 as
Nonionina bulloides. It is a small deep-water form, found both
recent and fossil.
Model no. 44. Cristellaria Cassis*, F. & M. Page 290, no. 8.
Hab. Adriatic, near Rimini; and fossil at Sienna. PI. I. fig. 45.
The young shell. The adult is represented in Model no. 83.
Model no. 45. Polystomella crispa*, Linn. Page 283, no. 1.
Hab. The Atlantic, the Mediterranean, and the Adriatic.
Pel tie 96.
Model no. 46. Nonionina levis, DPOrb. Page 294, no. 11.
Hab. Fossil near Paris. PI. III. fig. 97.
This is Nautilus incrassatus, F. & M. (Nonionina incrassata :
see Ann. Nat. Hist. ser. 3. vol. v. p. 101).
Model no. 47. Cristellaria levigata, D’Orb. Page 292, no. 19.
Hab. Fossil near Caen (Jurassic). Plate I. fig. 43.
A not uncommon form of Cristellaria, im which the early
chambers are rotulate, and the later chambers strive to take a
rectilinear arrangement. D’Orbigny refers, on the same page,
to other Cristellarie from the Limestone of Caen, namely C.
lamellosa, C. Cadonensis, C. Lituus; and at p. 259 he refers to
Planularia elongata, P. depressa, and P. striata from the same
limestone.
Model no. 48. Peheroplis planatus, F. & M. Page 285, no. 1.
Hab. Mediterranean, New Holland. PI. I. fig. 18.
This is the narrow variety, P. arietinus, Batsch.
Model no. 49. Planulina Ariminensis*, D’Orb.
Page 280, no. 1, pl. 5. figs. 1-3 bis.
Hab. Adriatic, near Rimini. Plate III. fig. 78.
A flat, thin Planorbulina, with raised chamber-walls.
Model no. 50. Alveolina Boscii*, Defrance. Page 306, no. 5.
Hab. Fossil near Paris. Pl. I. fig. 23.
The subtypical fusiform Alveolina, for which Montfort’s trivial
name, A. sabulosa, takes precedence.
Nomenclature of the Foraminifera. a7
LivRaAison 3",
Model no. 51. Nodosaria (Glandulina) Glans*, D’Orb.
Page 252, no. 2.
Hab. Adriatic; rare. PI. I. fig. 30.
The finely striated variety of G. levigata, D’Orb. Batsch
figures this, together with the primary elongated Nodosarian
form, as Nautilus comatus. (Sechs Kiipfertaf. pl. 1. figs. 2 a—d.)
Model no. 52. Nodosaria (Mucronina) Hasta, D’Orb.
‘Page 256, no. 49.
Hab. Adriatic. PI. I. fig. 29.
A carinate, finely striated Nodosarian, of the straight type.
Reuss refers to this Model as a Frondicularia—a judgment in
which we cannot agree. Its straight septa and lateral keels
indicate rather its Linguline affinity ; besides which, it is more
elongated in contour than is usual in Frondicularia. It is quite
out of the question to draw a distinct line between these two
subgenera ; but in this case there is no room for doubt as to the
nearest relationship.
Model no. 53. Rimulina glabra, D’Orb. Page 257, no. 1.
Hab. Adriatic. Pl. 1. fig. 37.
An oblique, robust, somewhat compressed, few-chambered
shell, with the aperture taking the form of a long slit down the
edge of the large terminal chamber. Type, Nodosarina Raphanus.
Model no. 54. Vaginulina elegans, D’Orb. Page 257, no. 1.
Hab. Adriatic. PI. I. fig. 33.
A beautiful limbate Vaginulina, with the septal lines thickened
by exogenous deposit of clear shell-substance.
Model no. 55. Marginulina glabra, D’Orb. Page 259, no. 6.
Hab. Fossil near Sienna. PI. I. fig. 36.
A common form, not coiled enough to be a Cristellaria, and
not well enough nourished, one may say, to be enriched with
thickened shell-matter and ribs such as we see in Marginulina
Raphanus, the well-grown type.
Model no. 56. Pavonia flabelliformis*, D’?Orb. Page 260, no. 1,
pl. 10. figs. 10, 11.
Hab. Madagascar. PI. I. fig. 22.
This is possibly a misprint for Pavonina. We have already
noticed it in speaking of the species illustrated by figures in
D’Orbigny’s Memoir. It may be an Orbiculina.
28 Messrs. Parker, Jones, and Brady on the
Model no. 57. Bigenerina Nodosaria*, D’Orb. Page 261, no. 1,
pl. 11. figs. 9-12.
Hab. Adriatic. PI. II. fig. 62.
A subtypical form of Teatularia, with a flat wide commence-
ment, but taking on a cylindrical uniserial mode of growth in
its later chambers. It is a common form in the Mediterranean
and many other seas, and has been found as far north as Shet-
land.
Model no. 58. Bigenerina (Gemmulina) digitata*, D’Orb.
Page 262, no. 4.
Hab. The Mediterranean. PI. IT. fig. 61.
Narrower in the growth of the early chambers than B. Nodo-
saria, having frequently a curved and compressed contour.
This form and the preceding have usually the subarenaceous
shell-structure of the larger Tewtularie.
Model no. 59. Vulvulina Capreolus*, Defr. Page 264, no. 1,
pl. 11. figs. 5-8.
Hab. Adriatic. Pl. II. fig. 63.
D’Orbigny refers this species to Defrance ; but we are unable
to find any notice of it in Defrance’s works.
As we have before stated, this 1s Grammostomum Pennatula,
Batsch, sp.
Model no. 60. Dimorphina tuberosa, D’Orb.
Page 264, no. 1.
Hab. Mediterranean. PI. II. fig. 53.
Some very dissimilar forms have been grouped under the
name of Dimorphina. As this is the first mention of the generic
term, we propose to confine its application to the Dimorphine
varieties of Polymorphina, of one of which Model no. 60 is a
correct delineation. It is rare. We have specimens of it from
the Crag of Suffolk. A somewhat similar, but really Nodosa-
rine, form occurs not unfrequently in the Mediterranean area,
both fossil and recent ; off Syra (90 fathoms) ; and fossil, from
Tertiary clays near Malaga.
Model no. 61. Polymorphina (Guttulina) Problema, D’Orb.
Page 266, no. 14.
Hab. Fossil, Castel-Arquato, Italy. Pl. II. fig. 50.
A many-chambered Polymorphina, somewhat irregular in the
disposition of the segments, which are long and distinct, and
do not overlap each other to the extent that is seen in most
members of the genus.
Nomenclature of the Foraminifera. 29
Model no. 62. Polymorphina (Guttulina) communis*, D’Orb.
Page 266, no. 15, pl. 12. figs. 1-4.
Hab. Adriatic; fossil near Bordeaux, Paris, Dax, and Castel-
Arquato. PI. II. fig. 47.
One of the forms alluded to in a previous paper as figured by
D’Orbigny. It is the typical Polymorphina lactea, W. & J., sp.
Model no. 63. Polymorphina (Globulina) gibba, D’ Orb.
Page 26, no. 20.
Hab. Atlantic (on the coast near Rochelle), Adriatic (near
Rimini) ; fossil, near Paris, at Grignon, near Dax, near Bordeaux,
at Chavagnes (Maine-et-Loire), and at Castel-Arquato. PI. IL.
fig. 52.
The simplest and Globuline condition of Polymorphina; the
chambers are few in number and embrace each other so as to
leave no inequalities in the surface-contour,
Model no. 64. Virgulina squamosa, D’Orb. Page 267, no. 1.
Hab. Fossil near Sienna. Plate IT. fig. 66.
One of the biserial forms of Bulimina, simulating Textularia
in its mode of growth; much elongated, like the more delicate
Textularie, but having the lengthened and twisted Bulimine
aperture. V. squamosa is subordinate to V. Schreibersit.
Model no. 65. Spheroidina bulloides, D’Orb. Page 267, no. 1.
Hab. Adriatic, near Rimini, Isle of France ; fossil near Sienna.
PRIL: fig: 58.
A good subtype, nearly related to Globigerina, but, in its clear
and white shell, indistinct perforations, and general arrangement
of chambers, bearing great similarity (or isomorphism) to the
Miliole. Its home is in very deep water, and it occurs most
plentifully in the seas of warm regions.
Model no. 66. Clavulina Parisiensis*, D’Orb. Page 268, no. 3.
Hab. Fossil near Paris. PI. I. fig. 26.
Type, Valvulina triangularis. This form is directly interme-
diate between the type (Model no. 25) and the narrow, pointed
condition shown in Model no. 2 (V. Clavulus). See Carpen-
ter’s ‘Introd.’ pl. 11. figs. 17, 18, and D’Orbigny’s “Cuba”
(Valvulina tricarinata), pl. 2. figs. 16-18. The Model appears
to have been taken from a specimen which has accidentally lost
its valve.
Model no. 67. Uvigerina pygmea*, D’Orb. Page 269, no. 2,
pl. 12. figs. 8, 9.
Hab. Fossil near Sienna. PI, II. fig. 54,
A typical form.
30 _ Messrs. Parker, Jones, and Brady on the
Model no. 68. Bulimina caudigera, D’Orb. Page 270, no. 16.
Hab. Adriatic, near Rimini. PI. II. fig. 65.
An acute-ovate Bulimina, with closely investing elongate seg-
ments, the later ones reaching back to the apex formed by the first.
Model no, 69. Rosalina globularis*, D’Orb. Page 271, no. 1,
pl. 138. figs. 1-4.
Hab. Shores of the Atlantic, growing attached to sea-weeds
and corals. Pl. II. fig. 69.
This is Discorbina globularis, a good subspecies, Egger (For.
Miociin-Sch. pl. 4. figs. 1-3) describes and figures it under the
name of Rotalina semiporata. A somewhat similar variety is
given by D’Orbigny (Fos. For. Vien. pl. 11. figs. 4-6) as Rosalina
obtusa, fine specimens of which, with lieato-granulated surface,
are common off Greenland. D. g/obularis is common everywhere,
from the shore down to 50 fathoms,
Model no. 70. Rotalia armata, D’Orb. Page 273, no. 22.
Hab. Cayenne, Martinique; fossil at Chavagnes (Maine-et-
Loire), near Nantes, and near Bordeaux. PI. ILI. fig. 88,
The short-spined variety of Calcarina Spengleri.
Model no. 71. Rotalia pulchella, D’Orb. Page 274, no. 32.
(No locality given.) Pl. III. fig. 80.
This belongs to the genus Pulvinulina, of which it is a
somewhat flat-faced variety. It is the Cidarollus plicatus of
Montfort. The same variety is given in D’Orbigny’s “ Cuba”
(pl. 5. figs. 1, 2, 8) as Rotalina Caribea.
Model no. 72. Rotalia (Discorbis) Gervillii*, D’Orb.
Page 274, no. 36.
Hab. Fossil at Valognes. PI. II. fig. 72.
This is the Discorbites (Discorbis) vesicularis of Lamarck—a
good varietal form of Discorbina Turbo, D’Orb., sp.
Model no. 78. Rotalia (Trochulina) Turbo*, D’Orb.
Page 274, no. 39.
Hab. Fossil near Paris. PI. II. fig. 68.
This is Discorbina Turbo; a type-form, having its fullest
development in D. trochidiformis, Lamarck, sp.
Model no. 74. Rotalia (Turbinulina) Beccarii*, Linn.
Page 275, no. 42.
Hab. The European shores of the Atlantic; Island of Mar-
tinique. PI. III. fig. 83.
Specimens of this somewhat variable species have received
Nomenclature of the Foraminifera. 31
several different names in the course of the ‘ Tableau des Cépha-
lopodes.’ Thus the Model now under consideration (74) is the
ordinary European form; Rotalina Corallinarum (Model no. 75)
is the more fully developed West-Indian variety, which is a still
better type; and R. tortuosat (page 275, no, 40) is the form
common in the Adriatic. Rosalina Parkinsoniana (For. Cuba,
p- 99, pl. 4. figs. 25-27) is another name given to an Atlantic
variety.
Model no. 75. Rotalia (Turbinulina) Corallinarum, D’Orb.
Page 275, no. 48.
Hab. The Atlantic, at Noirmoutier. PI. III. fig. 84.
Rotalia Beccarii, best type.
Livraison 4,
Model no. 76. Globigerina bulloides, D’Orb. (adult).
Page 277, no, 1.
Hab, Adriatic, near Rimini. PI. II. fig. 55.
The common, wide-spread, and essentially deep-sea Globi-
gerina. .
Model no. 77. Truncatulina refulgens*, Montfort, sp.
Page 279, no. 5, pl. 13. figs. 8-11.
Hab. Adriatic, near Rimini; Mediterranean, off Corsica; South
Sea at Rawack, Isle of Madagascar, Cape of Good Hope. PI. II.
fig. 76.
This belongs to the type, Planorbulina farcta, F. & M., sp.
Model no. 78. Planorbulina nitida*, D?Orb. Page 280, no, 1.
Hab. Atlantic, coast of Bellisle. Pl. II. fig. 75.
A convenient subspecies, intermediate between P. Mediterra-
nensis, D’Orb., and P. (Truncatulina) lobatula, W. & J., sp.
Type, Planorbulina farcta, ¥. & M., sp.
Model no. 79. Planorbulina Mediterranensis*, D’Orb.
Page 280, no. 2, pl. 14. figs. 4-6 bis.
Hab. Mediterranean, attached to various bodies. PI. II.
fig. 74,
D’Orbigny subsequently altered the name of this form (For.
Cuba, p. 85, pl. 6. f. 11-13) to Planorbulina vulgaris, because
he found that the species was not peculiar to the Mediterranean.
Such a change seems contrary to rule; so we adhere to the
original trivial name.
t+ Termed Rosalina Beccarii by D’Orbigny in his ‘ For. Cuba,’ p. 100.
32 Messrs. Parker, Jones, and Brady on the
Model no. 80, Operculina complanata*, Defrance.
Page 281, no. 1, pl. 14. figs. 7-10.
Hab, Fossil, Bordeaux. PI. III. fig. 93.
Subtype of Nwmmulina.
Model no. 81. Vertebralina striata*, D’Orb. Page 283, no. 1.
Hab. Mediterranean; Red Sea; South Sea, near Rawack.
Plo. fies:
The robust, compressed, typical form. See also note on
Model no, 22.
Model no. 82. Robulina cultrata*, Montfort. Page 287, no. 1.
Hab. Adriatic; and fossil near Vienna. PI. I. fig. 39.
There seems no necessity for Robulina as a generic term; and
it would be impossible to carry out even the conventional dis-
tinction, depending upon the degree of curvature and minute
differences im the position of the terminal orifice, laid down by
D’Orbigny in separating it from Cristellaria. Cristellaria cul-
trata is a good subspecific name for those carinate varieties of the
subtype, Cristellaria Calcar, Linn., which are without spinous
processes.
Model no. 83. Cristellaria Cassis*, F. & M. Page 290, no. 3.
Hab. Adriatic, near Rimini; fossil near Sienna. PI. I. fig. 44.
The adult shell. The young form is given in Model no. 44.
Model no. 84. Cristellaria costata, D’Orb. Page 292, no. 10.
Hab. Adriatic, near Rimini. PI. I. fig. 46.
A thin, elongated, carinate Cristellaria, with parallel coste
over the body of the shell.
Model no. 85. Cristellaria (Saracenaria) Italica*, Defy.
Page 293, no. 26.
Hab. Adriatic, near Rimini; and fossil near Sienna. PI. I.
fig. 41.
A trihedral keelless variety of Cristellaria Calcar. Model
no. 19 shows the young condition of the same variety.
Model no. 86. Nonionina umbilicata*, D’Orb. Page 293, no. 5,
pl. 15. figs. 10-12.
Hab. Adriatic, near Rimini; Mediterranean ; and fossil near
Bordeaux and Sienna. PI. ILI. fig. 98.
A variety of Nonionina asterizans, F. & M., sp., scarcely dis-
tinguishable from N. pompilioides of the same authors.
Nomenclature of the Foraminifera. 33
Model no. 87. Nummulina planulata, Lamk.* (young).
Page 296, no. 4.
(No locality). Pl. IIL. fig. 95.
The Model has been constructed from a somewhat convex
individual, and the aperture is wrongly placed.
Model no. 88. Nummulina (Assilina) discotdalis, D’Orb.
Page 296, no. 1.
Hab. South Sea, at Rawack. PI. III. fig. 94.
This is an Operculina, one of the thick subvarieties of O. com-
planata, Def. The term Assilina has been applied to those flat
Nummuline which, having no alar flaps to their segments, do
not cover up their umbones, but leave the whorls apparent, with
or without an increased growth of the edges of the septa.
Model no. 89. Siderolina levigata, D’Orb. Page 297, no. 2.
Hab. Fossil, St. Pierre at Maestricht. Pl. III. fig. 90.
A variety of Calcarina Spengleri, Gmel., sp.; a smooth,
quadrangular (or rather bluntly 4-spined), biconvex form, having
all traces of the chambers concealed by the excessive deposit of
shell-substance.
Model no. 90. Biloculina bulloides*, D’?Orb. Page 297, no. 1,
pl. 16. figs. 1-4.
Hab. Adriatic, near Rimini; fossil near Paris and near
Bordeaux. PI. I. fig. 3.
This varies but little in its outline from Biloculina ringens,
Lamk., sp., and B. levis, Defr., sp.
Model no. 91. Biloculina depressa, D’Orb. Page 298, no. 7.
Hab. Adriatic, near Rimini; fossil at Castel-Arquato. PI. I.
fig. 4.
The flattened carinate subvariety of B. ringens.
Model no. 92. Spiroloculina depressa*, D’Orb. Page 298, no. 1.
Hab. Mediterranean ; and fossil at Castel-Arquato. Pl. I.
fig. 6.
In the ‘Annales des Sciences Naturelles’ this Model is referred
to as Spiroloculina perforata—evidently one of the very numerous
misprints which oceur in D’Orbigny’s Memoir. We take the
name given with the Model as the one probably intended, The
form represented is Spiroloculina planulata, Lamk., sp.
Model no. 93. Triloculina trigonula*, Lamk. Page 293, no. 1.
Hab. Fossil near Paris, Soissons, and Valognes. PI. I. fig. 7.
A-good subtype.
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. 3
34 Messrs. Parker, Jones, and Brady on the
Model no. 94. Triloculina tricarinata, D’Orb. Page 299, no. 7.
Hab. Red Sea. PI. I. fig. 8.
This form is closely allied to 7. trigonula, of which it may
be considered a variety. The outer margins of the chambers,
instead of being rounded, as in 7. trigonula, are produced, and
form a sharp keel-like armature to the three angles of the shell.
Model no. 95. Triloculina oblonga*, Montagu. Page 300, no. 16.
Hab. Adriatic; Mediterranean; Atlantic, on the French and
English coasts; the Antilles; fossil near Bordeaux, Soissons,
Dax, and Castel-Arquato. PI. I. fig. 9.
The elongated, compressed form of the same subtype, passing
in many instances into weak forms of Quinqueloculina Seminu-
lum, Linn., sp.
Model no. 96. Quinqueloculina secans, D’Orb. Page 303, no. 43.
Hab. Adriatic and Mediterranean. PI. I. fig. 10.
This is the large outspread form of Quinqueloculina Seminu-
lum, Linn., sp., common in all our littoral sands, and having
sharp edges and obscure irregular transverse markings, never
amounting to costee.
Model no. 97. Adelosina striata, D’Orb. Page 304, no. 2.
Hab. Fossil at Castel-Arquato. PI. I. fig. 15.
The immature form of Quinqueloculina Brongniartii, D’Orb.
See note to Model no. 18, which represents the still younger
condition of the shell.
Model no. 98. Amphistegina Lessoni*, D’?Orb. Page 304, no. 3,
pl. 17. figs. 1-4.
Hab. Isle of France. PI. III. fig. 92.
A thick variety of Amphistegina vulgaris, D’Orb. See Model
no. 40.
Model no. 99. Heterostegina depressa, D’Orb.+ Page 305, no. 2,
pl. 17. figs. 5-7.
Hab. Island of St. Helena. PI. IIT. fig. 100.
A high-class Foraminifer of the Nummuline group, but having
its proper chambers subdivided by secondary septula into nu-
merous chamberlets. See Carpenter’s ‘ Introd. Foram.’ p. 288,
pl. 19. fig. 1. Heterostegina is plentiful in some parts of the
tropical and subtropical seas, and occurs fossil in the Middle
Tertiary Limestones of Malta, Vienna, and the West Indies,
and in Arabia.
+ Inadvertently omitted to be noticed in our former paper on D’ Orbigny’ s
species, Ann. Nat. Hist. ser. 3. vol. xii. p. 439. .
Nomenclature of the Foraminifera. | 35
Model no. 100. Fabularia Discolithes*, Defr. Page 307, no. 1,
pl. 17. figs. 14-17.
Hab. Fossil near Paris ; a much depressed variety at Valognes.
Plt. femG:
Probably, as we have before stated, this specific name is a
misprint for Discolithus. The name applied by Roissy, however,
(F. ovata) takes precedence.
Catalogue of D’ Orbigny’s Models. 1826.
Models,
Type. Corrected name. Pl. | Fig. D’Orbigny’s name. —_a
Livr.| No.
Vertebralina Vertebralina striata, D’O....... i 1. |Vertebralina striata ...| 4. | 81.
striata, D’ O. { —— conico-articulata, Batsch.| ,, 2.|Articulina nitida ...... L222
Biloculina ringens, Lam. ...... ” 3. |Biloculina bulloides ...). 4. | 90.
dopressa, 2)’ O; sccsscscosse| 54. 1|) 4s depressa,..-<..-« | Ee | ONE
—— aculeata, DO. ............ less 5. |—— aculeata ............ 20 ols
Spiroloculina planulata, Lam..| ,, | 6. |Spiroloculina depressa .| 4. | 92.
Triloculina trigonula, Zam. ...| ,, 7. |Triloculina trigonula...| 4. | 93.
tricarinata, D’O. ....,.... a 8. |—— tricarinata......... 4. | 94,
Miliola Semi- } |—— oblonga, Montagu......... later 9. |—— oblonga ............ 4, | 95.
nulum, Linn. | |Quinqueloculina secans, D’O.. ,, | 10.|Quinqueloculina secans| 4, | 96.
Daya ON ey asaas-sg creel a3) (ML Laren Liye cy, aceite, cae li ® 8.
— Ferussacii, D’O. ......... We spuslielzze Ferussacii ......... | 2. | 32.
—— Saxorum, Lam. .........0.. 3», | Ld. i—— Saxoruma,.-......-.. Pe || 3334
—— Brongniartii, D’O. (very) ,, | 14. |Adelosina striata
[young). (YOUN) eee seee elses
\|_—Brongniartii,D’ O.(young)| ,, | 15. /—— striata (adult) am || Oe
CE ype) isccnsAeass> Fabularia ovata, Rotssy. ...... » | 16. Fabularia Discolithes.... 4. 100.
|Peneroplis pertusus, Forsk. ...| ,, | 17.|Peneroplis planatus ...| 1. | 16.
Peneroplis —— arietinus, Batsch. ......... ees eV Se en teen 2. | 48.
pertusus, Fors. ) |Spirolina Lituus, Gmelin....... », | 19. Spirolina cylindracea...) 1. | 24.
Dendritina Arbuscula, D’O....;,, | 20.|Dendritina Arbuseula..| 1. | 21.
Gir pe int veansee Orbiculina adunea, F. f M....),, | 21. |Orbiculina numismalis.. 1. | 20.
Orbiculina?...... Pavonia flabelliformis, D’O....| ,, | 22. |Pavonia flabelliformis...| 3. | 56.
(ype) sae .n Alveolina sabulosa, Monff. ...! ,, | 23. |Alveolina Boscii......... 2, | 50.
Valvulina tri- / |Valvulina triangularis, D’O....) ,, | 24. |/Valvulina triangularis .| 1. | 25.
angularis, Clavulus, Lam. ..........- -| y, | 25, |Nodosaria Clayulus Sai, ey
DQ. | |—— Parisiensis, D’ O............ » | 26. |Clavulina Parisiensis...| 3. | 66.
Nodosaria Radicula, Linn. ...) ,, | 27. |Nodosaria.Radicula ...| 1. I.
Lingulina carinata, D’O. ...,.. oh | lek ‘Lingulina carinata...... 2. | 26;
= Hasta On es cnece ae » | 29. |Nodosaria Hasta......... 3. | 52.
Glandulina Glans, D’O. ...... ae aoOs|=——— | Gans ease eee eee 3 | ol:
| Frondicularia rhomboidalis, | ,, | 31. |Frondicularia rhomboi-
[ DO. dalisieescaweren. [iis
| |Dentalina obliqua, D’O. ...... , | 32, |Nodosaria obliqua ...... Ll. 5.
Vaginulina elegans, D’O. ...... ,», | 33. |Vaginulina elegans...... 3. | 54.
—— tricarinata, D’O, .........| ,, | 34. tricarinata ......... 1; 4.
Nodosarina Marginulina Raphanus, Linn..) ,, | 35.|Marginulina Raphanus} 1. | 6,
Raphanus, { |—— glabra, D’O. ............... », | 36. |—— glabra.,............. 3. | 55.
Linn, Rimulina glabra, D’O. ......... » | 37. |\Rimulina glabra......... 3. | 53.
| Planularia Cymba, D’O. ...... » | 38, |/Planularia Cymba ...... Ps, Ne
Cristellaria cultrata, Montf. ...|,, | 39. |Robulina cultrata ...... 4, | 82.
| —— virgata, D’ O........csceceee » | 40. Virgata:*...5cps...+ | akee
—— Italica, Defr. ............06. 5, | 41. \Cristellaria Italica ...... 4, | 85
—— —— (youing)............... » | 42.— (young) ...} 1. | 19.
—— levigata, DO. ............ » | 45. \—— leevigata............ 2. | 47.
Wassiss HNP scene sate » | 44. Cassis (adult) 4. | 83.
a (YOURE) Nerpereesee.e: » | 40.) —— (young)...... 2, | 44,
\ (lays ih JBMO} concanenceoaone 3 | 46. | Costaty .......0006- 4, | 84:
Messrs. Parker, Jones, and Brady on the
Type. Corrected name.
Polymorphina lactea, W. f J. .| IT.
—— Burdigalensis, D’O. ...... Fs
Polymorphina } |—— Thouini, D'O. ............
lactea, W. § J. Problema, D’O. ......... x
— Gutta, DO. ............4.. Ae
feloloe IOOE” cconcopAsocoase -
Dimorphina tuberosa, D’O. ...| _,,
(CBYPG)idvecsacegee. Uvigerina pygmiea, D’O. ...... a
oe (|
(ype) Beier eseccs Pullenia spheroides, D’O. ...| ,,
(CN 7Tst2) \egadoaccene Spheeroidina bulloides, D’O....| ,,
(\Textularia pygmexa, DO. . 3
Textularia ag- | |—— gibbosa, D’O................ %
glutinans, | Bigenerina digitata, D’O. ...... ys
DiC: i= Nodosarias2))) Oo eccsesas: _
| Grammostomum Pennatula, *
Balin Bulimina elegans, D OO: A Baa;
Prosliienes: = caudigera, D) O.sgevseses sss 8
: Virgulina squamosa, D’O....... ns
(49981) poopadadote Cassidulina levigata, D’ O. of
( Discorbina Turbo, B Osc: ;
F : lobullaris; 2DVO. i c...: sess
Dicer ee ff esas eer aaee DON eicicsnsskiins
Sea pea 1 —— rosacea, DO Leader Ree ‘
: vesicularis, Dam. ......... i
elegans; DUO. .:0..cs00++e. i
Planorbulina Mediterranensis,| ,,
[D’O.
Planorbulina | |—— nitida, D’O. ..............- 5
farcta, F. § ¢ |Truncatulina refulgens, Monzf.| ,,
M. —— lobatula, W. Gf J. .........
Pulvinulina
repanda, £.
Sf M.,
Rotalia Bec-
carl, Linn.
Calearina
Spengleri,
Linn.
Amphistegina
vulgaris,.D’ O.
Nummulina
perforata,
Montfort.
Polystomella
crispa,Linn.
ee ee rr
—— rosea, D’O.
Pulvinulina pulchella, D’O....
———— Men ancien) 1Oneenccsceceae
—— punetulata, DO. 0.00...
Rotalia Beccarii, Linn. (Huro-
[pean form).
Becearii, Linn. (West-In-
[dian form).
—— orbicularis, D’ O.
—— Soldanii, D’O. ...........
Calcarina Spengleri, Linn.
——armata, DO... ..J.c.ei0s:
—— bisaculeata, DIO:
—— levigata, D’O.
Amphistegina vulgaris, D’ 0...
Lessoni, D’ 0.
Operculina complanata, Def. sor
—— discoidalis, D)O. .........
Nummulina planulata, Lam....
Polystomella crispa, Linn.
Nonionina incrassata, /’. ¢° M.
—— pompilioides, /. § MW. ...
|—— Limba, DO. ............00:
Heterostegina depressa, D’O. .
Planorbulina Ariminensis, D” O. TI.
Fig. D’Orbigny’s name.
47. Polymorphina commu-
MNS.) ro acleeisoeianeneee
48. —— Burdigalensis
49, |—— Thouini ............
50. |—— Problema .........
51. | —— Gutta ..
52. PADD Aas, eciicanecnads
53. Dimorphina tuberosa...
99.
100.
. |Planorbulina
. |—— nitida
. Truneatulina refulgens .
. |—— tuberculata.........
. |Planulina Ariminensis .
. |Rotalia rosea ....cecceses
. |\—— Menardii
2. |—— punctulata
. |\—— Beccarii
. |Rotalia armata
)
. Uvigerina pygmea......
5. |Globigerina bulloides...
(young)......
7. |Nonionina spheeroides .
. |Spheeroidina bulloides .
. |Textularia pygmexa
gibbosa
. |Bigenerina digitata......
. |— Nodosaria
. |Vulvulina Capreolus ...
. |Bulimina elegans
caudigera
. |Virgulina squamosa ...
. |Cassidulina leevigata ..
. |Rotalia Turbo
teeter n eens
re
Mediter-
ranensis
teeter ete eee nene
pulchella
eee wneee
See eee aee
eee ennes
5. \Gyroidina orbicularis...
. |—— Soldanii
bisaculeata
ween eens
. |Siderolina levigata
. Amphistegina vulgaris .
92. |—— Lessonii
. Operculina complanata,
. |-—— discoidalis
. Nummulina planulata .
}. |Polystomella crispa
. |Nonionina leevis.........
umbilicata
Heterostegina depressa.
ILM Dace cee eeneeeee
Models.
—_——
Livr. | No.
3. | 62.
3. | 29.
2. | 28.
3. | 61.
2. | 380.
3% || (oy,
3. | 60.
3: Oe
4. | 76.
We |) ily.
2. | 43.
oe G0:
“Tel fe
2. | 28.
3. | Oo:
By |} D7
3. | 09:
iL 9:
3. | 168!
3. | 64.
2. | 41.
3. | 73.
3. | 69.
2. | 38.
2. | 39.
3. | 72.
2. | 42.
AD i:
4. | 78.
AS eitte
De edie
2. | 49.
BA aly
BH | Zflle
ee eos
Ths |) 4,
3. | 74.
3. | 75.
Vo des
BS || olbe
2. | 34.
3. | 70.
ils |) Us
4. | 89.
2. | 40.
4. 8.
4, | 80.
4, | 88.
4, | 87.
oulnetae
2. | 46.
4, | 86.
i jf) We
4, | 99.
Nomenclature of the Foraminifera. 37
APPENDIX.
Reuss and Fritscl’s Models of Foraminifera. 1861.
Whilst speaking of Models of Foraminifera, it may not be
amiss to notice a more recent series than that of D’Orbigny—
namely, a set of one hundred plaster Models prepared at Prague,
under the direction of Professor Reuss and Dr. Anton Fritsch*.
These Models are 5 centim. in length, and are furnished with
printed labels. The species have been selected with a view of
supplying a perfect series, and at the same time completing
D’Orbigny’s suite of Models. Although the aim of the authors
of this set of Models is to give delineations of a more extended
series of types than were known to D’Orbigny, and though the
workmanship expended upon them has the advantage in point
of skill, it may be doubted whether the species chosen for illus-
tration are altogether so apt, or the general result so instructive,
as in the earlier series. Of the forms thus illustrated, some
have been selected from species already known, and many have
been described and figured in one or other of Professor Reuss’s
numerous papers on Fossil Foraminifera. The needless multi-
plication of genera appears in a striking light in reviewing the
nomenclature of this catalogue. We propose, however, merely
to give the list of these Models as arranged in the “ Catalogue,”
with the Number and Locality appended to each; also a few
remarks, when specially called for, and the ¢ype to which the
form belongs. We leave the order and the classification as we
find them, thinking that it may be of interest to some to com-
pare the results with the somewhat similar, but less artificial,
system which we are in the habit of usimg, based upon the
principles laid down in Dr, Carpenter’s “ Introduction.”
We have to thank Mr. 8. V. Wood, F.G.S., for lending us
his set of D’Orbigny’s Models during several years, whilst we
have been engaged on this Memoir.
* Verzeichniss von 100 Gypsmodellen von Foraminiferen, welche
unter der Leitung des Prof. A. Reuss und Dr. Anton Fritsch gearbeitet
wurden. Ausgegeben von W, Fric, Naturalienhandler in Prag, Wasser-
gasse Nro. 736— U.
“ Die Auswahl der Species ist so getroffen, dass die gegenwiirtige Samm-
lung ein vollstiindiges Ganze pildet, und dabei die D’ ‘Orbigni’ sche Suite
kompletirt. Die Exemplare sind 5 Centimétre gross und mit gedruckten
Etiquetten versehen.
“Der Preis (sammt Emballage) 36 fl. 6. W. oder 24 Th. P. C.”
MM. Reuss and Fritsch have liberally given us a setj of these useful
Models,
38
Messrs. Parker, Jones, and Brady on the
Catalogue of the Models of Foraminifera prepared by Professor
Dr. A, E. Reuss and Dr. Anton Fritsch.
[ Prof. Reuss’s Classification is here followed. ]
1861.
I. FORAMINIFERA WITH NON-POROUS SHELLS.
A. Wiru Sanpy SILicrous SHELLS.
I. Lituolidea (Reuss).
- 3. Haplophragmium inflatum ...... Upper Greensand.
(L. nautiloidea, Lam.)
4, irregulare, Roem., sp. ..... Chalk.
(L. trregularis, Roem.)
II. Uvellidea (Reuss).
5. Valvulina triangularis, D’O. ...... Valvulina triangularis, D’ O,...|Eocene.
6. Verneuilina spinulosa, ss. ......) | (Miocene.
7. Tritaxia tricarinata, D’O., sp. ... } Textularia agglutinans, D’ O. ‘Cretaceous.
(Verneuilina tricarinata; old) } |
8, 9. Ataxophragmium variabile, D’ O.,
BPon (Ganon Gay -eameccsrsetasteertnerce Bulimina Preslii, Fss. ......... Chalk.
10. Clavulina communis, D’O. ...... ‘Tertiary.
11. Gaudryina pupoides, D'O. ...... Textularia agglutinans, D’O.< Upper and Middle
12. Bigenerina Nodosaria, D’O....... | Recent. [ Chalk.
3s Conwlinaiconica,,)) Omerncercns ors: '? Lituola nautiloidea, Zam. ... Recent.
14. Chrysalidina gradata, D’O. ......... ?Chrysalidina gradata, D’ O.... Cretaceous.
B. Wirn Compact, Porce,titanovus, CaLcarrous SHELLS.
I. Squamulinidea (Reuss).
Il. Miholidea (Reuss).
15. Cornuspira involvens, Fss. ......... Cornuspira foliacea, Phil. ...... Miocene.
16. Uniloculina Indica, D’O.......... , ( Recent.
(? Young of a striped Quin-
queloculina.)
17. Biloculina Lunula, D’O. ......... | Tertiary.
(B. depressa, D’O. a : 5
18. Spiroloculina dilatata, » OS beeek pg 3 SEONG Eigse Miocene.
(Sp. planulata, Lam.)
19. ‘Priloculitia pibba, DO. ..:......... Tertiary.
(TL. trigonula, Lam.)
20. Quinqueloculina, sp., D'O. ...... | Tertiary.
21. Fabularia discolithes, Defr. ......... Fabularia ovata, Rozssy......... Eocene.
Name.
(Lituola Cenomana, D’O.)
. Haplostiche foedissima, ss. ......
(LZ. Scorpiurus, Montf.)
. Placopsilina irregularis, D’O. ...
Type.
|
Lituola nautiloidea, Lam. ...
|
Locality.
Upper Chalk.
Upper Greensand.
Nomenclature of the Foraminifera. 39
No. Name. Type. Locality.
III. Peneroplidea (Reuss).
22. Peneroplis pulchellus, D’O. ...... Recent.
23. planatus, Montf. ............ : Recent.
24, Dendritina arbuscula, D’O. ...... Peneroplis pertusus, Forssk. . |Recent.
25. Spirolina Austriaca, D’O.......... Middle Tertiary,
26. Vertebralina mucronata, D’O. ...... Vertebralina striata, D’O....... ‘Recent.
27. Hauerina compressa, D’O. ......... Miliola Seminulum, Zinn....... ‘Miocene.
28. Pavonina flabelloides, D’O. ......... ? Orbiculina adunea, fF’. M....|Recent.
IV. Orbitolitidea (Reuss).
29. Cyclolina cretacea, D’O. ............ Patellina concava, Lam.......... Chalk-marl.
31.
. Orbitolites macropora, Laz..........
FORAMINIFERA WITH POROUS SHELLS.
2. Ovulites margaritacea, Lam...
pearenasim plexe i sSieacsestee neste
. Fissurina carinata, Rss.............
. Nodosaria tetragona, Fss. .........
. Orthocerina quadrilatera, D’O.......
. Dentalina acuminata, Pss.
. Vaginulina Badensis, D’ O.
5. Rimulina glabra, D’O. .........+6.
. Frondicularia Lanceola, Rss.
. —— Goldfussii, fss....../.........
. Rhabdogonium acutangulum, Rss.
. Amphimorphina Haueri, Newgel.
. Dentalinopsis semitriquetra, /ss.
. Flabellina oblonga, V.
. Psecadium ellipticum, Fss. ......
6. Lingulina costata, D’O.............
. Lingulinopsis Bohemica, Rss. ...
. Pleurostomella fusiformis, fss, (An
Il.
Orbitolites complanata, Lam...
Mastricht Chalk.
A. WitH HYAtine, FINELY Porous Catcargous SHELLS,
I. Spw
Spirillina punctata, D’O. ............ |
‘ulhdea (Reuss).
?
fmm remem meee rar ener ensersseseseseree
II. Ovulitidea (Reuss).
wos
Ill. Rhabdoidea (Reuss).
waleanies Walls. saiisneseisic sac
(L. levis, Montagu.)
(L. marginata, Montagu.)
inflata, Pss.
(A short NV. Raphanistrum.)
ISTO, TEES. agodencoactasct88c
(A long V. Radicula.)
Hormeina, DO. cesecccccsss
transversalis, Rss. ...........-
cristellaroides, Rss. .........
(Fr. complanata, Defr.)
inbtyenks Pa ais cess odonadcepee
Wikiharaoeiy JES, Sohoncaocesecde
(A triangular Dentalina.)
Miinst. ...
COrdabas elsss oaaeceeeedcee eee
(£7. oblonga, V. M.)
(A globose or Glanduline
Marginulina.)
(A Linguline Marginulina.)
Nodosarina Raphanus, Linn.
Lagena sulcata, W.f- J.......
Nodosarina Raphanus, Linn.
— Se =
Orthocerina Murchisoni, Rss...
———_--.
|
}
a
Orthocerina Murchisoni, Rss...
(
|Tertiary and recent.
|Ovulites margaritacea, Lam. ...|Eocene.
|Upper Chalk,
Recent.
Tertiary.
Gault.
‘Upper Chalk.
Upper Greensand.
|
Recent.
Upper Chalk.
Upper Chalk.
Upper Tertiaries,
Gault.
Lower Greensand.
Recent.
Upper Greensand.
Upper Greensand.
Chalk-marl,
Lower Greensand.
Miocene.
Lower Greensand.
Tertiary.
Upper Chalk,
Nodosarina Raphanus, Linn.
Bulimina Preslii, Rss. .........
extreme form of Virgulina.)
Miocene.
Miocene.
Chalk-marl.
Gault.
AO. On the Nomenclature of the Foraminifera.
No. Name. Type. | Locality.
IV. Cristellaridea (Reuss).
59. Marginulina Ensis, D’O. ......... Upper Chalk.
60. ulllata, 2200770. ..c-..eren-00- Upper Chalk.
aa Carer ll TC ‘Nodosarina Raphanus, Linn. perme
63. Robulina deformis, /?ss. ......... | Tertiary.
(First stage of C. rotulata.) )
V. Polymorphinidea (Reuss).
64, Bulimina pupoides, D’O. (f.) ... Tertiary and recent.
65. Virgulina WEGUUSA LESS te einniee sone Bulimina Preslii, Mss, ...... Tertiary.
(V. sguamosa, D’O.)
66. Uvigerina pygmza, D’O. ............ Uvigerina pygmea, D’O. ...... ‘Tertiary.
67. Poly morphina complanata, D’ O. ~ | Miocene.
68h PyrulinalGutta yO. c..cesc+ ce Tertiary.
69. Globulina equalis, D’O. ......... /Polymorphina lactea, Hg. | Upper Tertiary.
70. Guttulina Austriaca, D’'O. ...... : Tertiary.
71. Spheroidina Austriaca, D'O. ...... Spheeroidina bulloides, D’ O.... Tertiary.
(Sph. bulloides, D’O.)
72. Dimorphina obliqua, D’O. ......... Nodosarina Raphanus, Linn. Tertiary.
(A Dentaline Marginulina.)
VI. Cryptostegia (Reuss).
73. Chilostomella ovoidea, /ss. ...... [ Tertiary.
(A Biloculine Miliola?) PoMialisla Mominul :
74, Allomorphina cretacea, Iss. ...... ? Miliola Seminulum, Linn. Upper Greensand.
(A Quinqueloculine Miliola?)
VIL. Textilaridea (Reuss).
5. Textilaria Conulus, Pss. ......... Upper Chalk.
(T. agglutinans, D’O.) |
76. Proroporus complanatus, /ss. ... > Textularia agglutinans, D’ O. 4 |Gault.
(Thefirststage of Bigenerina.) |
77. Sagraina pulchella, D’O. ......... ) Recent.
78. Vulvulina Gramen, D’O. ............ Bulimina Preslii, ss. ......... Recent.
79. Bolivina Beyrichi, Iss. ............... ‘Textularia agglutinans, D’ O..../Tertiary.
80. Schizophora ING CHOLEME ASS! onc: | sacaexsesoseseossecastoemeesecnese see Miocene.
(Bigenerine Vulvulina Pen natula. )\
VIL. Cassidulinidea (Reuss).
81. Cassidulina crassa, D’O. ......... iI Cassidulinn levioata cDiO Recent.
82. Ehrenbergina serrata, Iss. ...... J 2 oh aga lane ** | | Miocene,
(Cassidulina serrata, Rss.)
B. ForaMIniIFERA WITH VERY Porous CaLcarEous SHELLS,
I. Rotalidea (Reuss).
83. Rotalia Brongniarti, D’O............. \Pulvinulina repanda, F’. gM. .|Miocene.
(Pulvinulina auricula, F.&M.)
84. Ginanclanaeltssiecocsiecesteessens /Rotalia Beccarii, Linn. ......... Tertiary.
(Rotalia Soldanii, D’O.) |
85. —— bulimoides, Rss. ...0........008 (Bulimina Preslii, ss. ......... Tertiary.
(Bulimina clegantissima, Will.)
86. Siphonina reticulata, Rss. ............ Planorbulina farcta, 7’. § M. ..|Miocene.?
(Planorbulina reticulata, Rss.) |
No.
87.
88,
90.
Sue
92.
95. —— bulloides, D’O. .............0006- |
: (Pullenia spheroides, D’O.)
96. Fusulina cylindrica, Fisch. .........
99.
. Amphistegina Quoyi, D’ O.
. Operculina, sp. (after Carpenter)...)
Mr. P. H. Gosse on Aigcon Alfordi.
Name.
Asterigerina Planorbis, D’O. ......
(Discorbina rosacea, D'O.)
89. Siderolithes calcitrapoides, Lam.
Planorbulina Mediterranea, D’ O....
Globigerina bulloides, D’O. .........
Orbulina universa, D’O. ...........-
41
Type. | Locality.
Discorbina Turbo, D’O. ...... ‘Tertiary.
\Calcarina Spengleri, Gm. ...... ‘Chalk of Mastricht.
Planorbulina farcta, #. & M. ..\Recent.
Globigerina bulloides, D’O. ... Tertiary.
(Orbulina universa, D’ O.
Tertiary and recent.
C. Wirn CaLcaAreEous SHELLS TRAVERSED BY AN INTERMEDIATE
CANAL-SYSTEM.
I. Polystomellidea (Reuss).
Polystomella aculeata, D’O. .........
Nonionina communis, DO. .........
(WV. Scapha, F. & M.)
II. Num
Heterostegina,sp.(after Carpenter )
costatas DON. css ws ee cceswer
Polystomella crispa, Linn (Tertiary and recent.
Nonionina asterizans, /’. & M..|Tertiary.
Pullenia spheroides,D’ O. ...... Tertiary and recent.
[ stone.
Fusulina cylindrica, Fésch. ...|\Carboniferous Lime-
mulitidea (Reuss).
Amphistegina vulgaris, D’ O....{Recent.
Nummulina perforata, Monéf. .. Recent.
Heterostegina depressa, D’ O. Recent.
‘Miocene.
'
V.—On AMegeon Alfor
di, a new British Sea-Anemone.
By Puirie Henry Gosss, F.R.S.
[
Fam
Plate VII.]
ily Antheade.
Genus A‘cron, mihi.
Base adherent to rocks w
ith a moderate tenacity ; broader than
the medium diameter of the column.
Column irregularly distensible, not mucous, somewhat versa-
tile, but generally forming a tall, erect, thick pillar, the summit
expanding; the margin tentaculate; the surface longitudinally
fluted (as if composed of a multitude of slender vertical cylinders
placed in contact side by side), each cylinder studded with a
single vertical row of minute warts. No suckers or loopholes.
Substance pulpy, membranous.
Disk expanded, membranous, concave, revolute.
Tentacles numerous, in several rows, long, lax, irregularly
flexuous, scarcely retractile.
Mouth not ordinarily set on a cone, but pouted after the re-
ception of food; lip thin. Gonidial tubercles prominent.
Acontia wanting (?).
42 Mr. P. H. Gosse on Aigeon Alfordi,
geon Alfordi, mihi.
The only known species. Colours and dimensions as follows :—
Basal disk brick-red.
Column pea-green, suffused with a purplish hue, as if from
within the skin, more marked towards the extremities than in
the medial region ; the whole covered with red dots, so minute
as to be distinguished only with the aid of a lens. The warts
have each but one speck of crimson, which is central and
much larger than the other dots. The red hue on the warts
being thus limited to a single speck, the area of the warts
appears of a brighter, clearer green than the rest of the body.
Disk. Inner half purplish grey, abruptly divided from the outer
half, which is of a most lustrous satiny green ; radii faint lines
of grey.
Tentacles lustrous satiny green throughout, each bearing a faint
line of grey along its outer side.
Mouth. Lip and throat grey.
When fully expanded, it is sometimes 4 inches in height by
11 inch in diameter; at other times it will be 2 inches in each
direction. Expanse of flower 64 inches.
Locality. The Scilly Islands.
This very fine Anemone, which might well put in a claim to
be considered pulcherrima, if we had “but a Paris to judge, and
which is by very far the noblest acquisition to our British marine
zoology that Iam aware of since the publication of my ‘Actino-
logia,’ we owe to the researches of the Rev. D. P. Alford, M.A.,
Chaplain of the Scilly Islands. On the 29th of March last,
this gentleman observed ‘‘some very bright green tentacles
reaching out from beneath a large stone,’ which, though he at
first supposed them to indicate an Anthea cereus, proved to be-
long to an unknown treasure. ‘ Here was an Anemone with
high-standing column hke an Aipéasia, but with the surface
warted, and with tentacles like the richest green velvet, throw-
ing into the shade the brightest of Antheas. Moreover the
tentacles were of the same colour to their very tips, without the
least tinge of pink or purple.” ‘I put mto the same bottle,”
continues Mr. Alford, “a Nereis 4 mches long; and by the time
I got home (within a quarter of an hour), the poor worm had
Been seized in the middle by the lovely green tentacles, and only
its head was to be seen protruding from the Anemone’s mouth.
With his good dinner, my friend became active, far more lively
than I have ever found Aiptasia. He soon fastened firmly to
the side of the basin; but his tentacles and column were per-
petually on the move. He shows himself off to most advantage
when he curves his column upwards so. as to present his full
a new British Sea-Anemone. 43
flower just beneath the surface of the water. After referring
carefully to your ‘ History, I found much resemblance in the
column to the Bunodes Ballii, but far more likeness in all other
respects to the Aiptasia. Its column has the crimson specks of
the former; but it has the size and flexibility of the latter. It
also resembles Azptasia in the length of its tentacles, in the fact
that these often become extremely attenuated, in never quite
closing the disk, in its well-marked radu, and also in having the
margin crenate ; for all the finer specimens of Azptasia which I
have observed have the margin crenate, not tentaculate.”* “The
green of the tentacles is very much that of the common Ulva;
and, like that, it keeps its brilliant colour by lamp-light, when
Anthea loses much of its beauty.”
Mr. Alford immediately submitted his prize to me for exami-
nation. We had both thought that it might possibly prove to
be Actinia pustulata of Dana; but a reference to the figures and
descriptions in that author’s great work on ‘ Zoophytes’ at once
set this suspicion at rest}. I have no hesitation in pronouncing
it hitherto unrecognized. With very obvious affinities to both
Aiptasia and Anthea, the character of the column well distin-
guishes it from either. The surface in Aiptasia is minutely cor-
rugated, in Anthea cancellated by the intersection of furrows ;
in Afgeon there are frequently seen transverse wrinkles; but
the warts are very manifest when, from a peculiar curve of the
body, these cross wrinkles are quite obliterated. It has much
more of an erect column than Anthea. After very many pro-
tracted watchings with a powerful lens, when the body was in
the most favourable conditions for observation, I could never
discern the slightest trace of cinclides; nor has any amount of
provocation educed the emission of acontia.
The cylinders of which the skin of the column is built up are
alternately larger and smaller: each of the former terminates in
a short process (the marginal tentacles), the latter are truncate.
This structure is seen to most advantage when the animal is
greatly distended.
With respect to its habits in captivity, several circumstances
indicate that this charming’ species is very eligible for the aqua-
* T have considered these marginal processes as budding tentacles in
both species. Perhaps Iam wrong, but it may suffice to say that in this
respect 4igeon agrees with Aiptasia. They are the projecting summits of
the cylinders or flutes, and alternate with the outermost row of tentacles
proper.
+ A. pustulata, A. veratra, A. clematis, and A. florida of Dana appear
to constitute a genus of Actiniade (as limited in Act. Brit. p. 171), agree-
ing with Actinza in the possession of marginal spherules, and differing
from it im having a warted column. M. Milne-Edwards (Corall. i. 274)
has given the name of Phymactis to this genus.
4A Mr. P. H. Gosse on Aigeon Alfordi.
rium. In the first place, it possesses a wonderful tenacity of
life, as the following facts will show. On the 4th of April,
Mr. Alford enclosed the specimen in a small canister, and sent
it to me by post. On the 12th, an ample missive under the
great seal of the General Post Office informed me that a package
pane to me was detained by the postmaster at Plymouth,
on account of the exudation of water, according to statute in
that case made and provided. Hope pretty nearly died within
me; but I wrote to a friend at Plymouth, who kindly obtained
the offending package, gave the prisoner a twenty-four hours’
bath, and then re-posted him on to me. On the evening of the
17th I turned him out, and, before night, had the pleasure of
seeing him adhering, and expanding in all his beauty, none the
worse for his fortnight’s captivity.
From that day to this (June 10) he has luxuriated in a little
cylindrical vase of sea-water, always displaying his full glories
in the most ungrudging manner. He is always ready for dinner,
and swallows large lumps of raw meat with a very vigorous
appetite. The liveliness and versatility of his movements ereatly
augment the interest which attaches to him as a tenant of the
tank. Mr. Alford’s first evening’s impressions of his character
—<T have had him close beside me all this evening, and he has
never been alike in shape or size for two minutes together”
have been justified by my more lengthened experience.
At present the specimen remains unique ; but my friend is on
the qui vive*, and we may hope that more examples may soon
be discovered among the sea-beaten rocks of those rocky outposts
of England.
As it devolved on me to give the illustrious stranger a pair of
names, I have borrowed one from a haat eepLamne hero of
antiquity —
“ geon qualis, centum cui brachia dicunt,
Centenasque manus ”’} ;
and the other from the fortunate discoverer.
EXPLANATION OF PLATE VII.
AEgeon Alford, of the natural size, in its ordinary state of distention.
* T will add an interesting fact from a letter of Mr. Alford’s :—‘‘ The
great abundance of Aiptasia seems the most marked feature of these
islands, as far as Anemones are concerned. Other species and varieties
are well represented ; but amongst the rocks in Porth Crassa Bay, at the
back of my house, the Aiptasie are innumerable—far more common than
Actinia mesembryanthemum.”
+ Virg. Aineid. x, 565.
Mr. A. Macalister on Secreting Organs ix Nematoidea, 45
VI.—On the Presence of certain Secreting Organs in Nema-
toidea. By AuexanperR Macauister, F.R.C.S.1., Demon-
strator of Anatomy, Royal College of Surgeons, Ireland.
Tuer existence of special secreting organs in the Nematoid
Entozoa is by no means a discovery of very modern date; for
several of the earlier helminthologists have described various
parts of the animals in this class as subservient to the function
of secretion. Of late, however, our knowledge of these struc-
tures has been much extended, mainly through the increasing
perfection of the microscope, which has thrown light upon all
branches of invertebrate anatomy, and has shown us greater
complexities of structure in those creatures which had previously
been regarded as of simpler organization.
Four series of these glandular organs have been already de-
scribed in different Nematoids; and I think that the apparatus
which I am about to notice is entitled to rank as a fifth kind of
secreting organ, separate in function from any of those at pre-
sent known. Those already recognized are—(1) The salivary
ceeca described by Owen in Gnathostoma spinigera, consisting of
four small blind tubes communicating with the mouth: similar
organs Siebold has noticed in Strongylus striatus ; and although
some have doubted the function assigned to them (Bagge, in
Appendix to ‘Thesis de Evolutione Strongylt auricularts, &c.),
yet I think we are justified in adopting Owen’s view as being
correct. (2) Cloquet, in his work on the anatomy of Ascaris
lumbricoides, describes the thickened parietes of the cesophagus
as being glandular, probably secreting a fluid to assist in the
assimilation of the food. (3) There are in many species intes-
tinal ceca with which Owen associates an hepatic function.
Mehlis, in the ‘Isis’ for 1831, figures and mentions several of
these ; and Siebold, in his ‘ Anatomy of the Invertebrata,’ refers
to their occurrence in several species, especially in Ascaris he-
terura, A. semiteres, A. depressa, A. angulata, A. ensicaudata,
A, mucronata, and A. osculata. Leidy, in the ‘Smithsonian
Contributions,’ pt. 5. p. 49, pl. 7, figures and describes one of
these organs in Thelastomum appendiculatum ; and Diesing no-
tices another in a species of Ascaris infesting the Dugong.
The 4th and last-described gland (leaving out of account the
secreting parts of the reproductive apparatus) is the curious
tubular organ described by Siebold (Bagge, loc. cit. supra) in
the Strongylus auricularis and Ascaris brevicaudata, A. acuminata,
A. paucipara, and A. dactyluris, which opens near the middle of
the body on the ventral aspect, and which in the last-named
species I have on several occasions traced with considerable
facility.
46 _ Mr. A. Macalister on the Presence of certain
To these four I think we may add another group of organs
which seem as distinctly glandular as any of those above referred
to. These are present in the Ascaris dactyluris, Rud., a small
white Entozoon, which inhabits in enormous quantities the large
intestine of Testudo greca. In the interior of these parasites,
as I have elsewhere described*, the lowest part of the club-
shaped intestine exhibits a small dilatation, immediately inferior
to which it suddenly contracts into a narrow rectum, that passes
downwards and forwards to the anus, forming an obtuse angle
(re-entrant forwards) with the upper part of the alimentary canal.
Surrounding the constriction which marks the origin of the
rectum, are four small ovate or pyriform bodies, granular in
appearance, usually seeming as though solid, in other subjects
appearing slightly excay ated. Their inner aspect is placed in
very close apposition to the wall of the gut; so that at first it
seemed to me as if they opened directly into the narrowed com-
mencement of the rectum; however, when carefully examined
by reflected light, my friend Dr. Barker has shown to me that,
at least im some specimens, such is but an apparent and not a
real attachment, and that the true connexion between these oval
bodies and the intestine is by means of long fine ducts, which
open into that canal immediately above the anus. Somecmes
these tubes pass from the inner or intestinal side of the glands ;
in other subjects the masses narrow into a somewhat flask-like
shape, and have their attenuated necks continuous with the
duct: in the former case the organs were globular, in the latter
they seemed rather pyriform. In another specimen the sacs
were calcarate, with their curved projecting spurs directed up-
wards and outwards. No appearance of nerves or nerve-ganglia
was visible in connexion with them; and the lateral and antero-
posterior tegumentary lines dipped i inwards to come almost into
contact with their outer coat.
Whether these bodies exist in other species of Nematoids or
not, I cannot say ; but, as far as my observations have extended,
T have not sueceeded in finding either themselves or any notice of
such an organization elsewhere. In the species under consider-
ation, Howeree they are unmistakeably distinct and constantly
present ; for out of many specimens examined by Dr. Barker
and myself, both separately and conjoitly, we were able to de-
tect their existence in ey ery individual.
It would be difficult, if not almost impossible, to predicate as
to the exact nature of these bodies; but I can only conceive of
** On the Anatomy of Ascaris dactyluris,” read before the Dublin
Natural History Society, June 1865.
+ This, however, does not seem to be the invariable mode of attachment ;
for I have failed to find the duct-like processes im many individuals.
Secreting Organs in Nematoidea. 47
two tenable hypotheses regarding them. They have evidently
nothing to do with the reproductive apparatus, as they are
equally present in both males and females, and seem to have no
connexion in either with the sexual organs. They might, how-
ever, be either secreting glands or intestinal ceca. The latter
hypothesis I should be inclined to regard as very improbable ;
for though ceca are described and figured by Leidy, of Phila-
delphia, and others, as I have before mentioned, yet in all those
species in which they occur we find them placed much higher
in the alimentary canal, often at the point where the stomach
or intestine joins the cesophagus: they are usually single or un-
symmetrical, always hollow; and though often communicating
with the intestine by a narrow neck, yet rarely or never is that
structure so suddenly attenuated and duct-like as is constantly
the case in these secreting organs in Ascaris dactyluris. We
are thus led to adopt the last hypothesis, that they are special
glandular structures—an opinion which, I think, is supported by
their numbers, by their thick, solid, granular walls, by their long
ducts, when present, and by their invariably low position with re-
gard to the alimentary canal. This latter point is also of much
importance in relation to the function fulfilled by these bodies, if
glandular; for as their secretion would be poured into the lowest
portion of the rectum, it could not be to any extent excrementi-
tious in its nature, but must be directly evacuated before absorp-
tion could take place ; so we may regard these organs as a means
of evolving effete matter from the system: mayhap they might be
among the earliest examples of a renal apparatus in the animal
kingdom ; and if so, certainly they are the first examples of such
having been found in the Entozoa. Indeed, as a general rule
among the lower departments of animal life, the appearances of
renal organs are more or less equivocal : even in the Insecta the
Malpighian tubes (by far the most distinct urinary apparatus in
the Articulata) were often mistaken for hepatic organs, until
Brugnatelli and Wurtzer proved that these canals contained
urate of ammonia in the Silkworm, as Meckel afterwards demon-
strated in Melolontha (Aychiy fiir Physiologie, 1816, 1818, 1826).
In Myriapoda the same tubular structure obtains with tolerable
distinctness ; and in Crustacea we have the urinary system re-
presented by the tubes traced by Milne-Edwards m Maia, by
Duvernoy in Portumnus, and by Meckel in Palemon and others,
Usually these are ceca, which open sometimes into the pylorus
but occasionally into the rectum. No representative organ has
been, to my knowledge, described in Annelida. In Echino-
dermata Jager has referred to the slightly branching sinuous
tubes of Holothuride as being renal in their nature ; but Miiller,
who describes these structures under the name of the Cuvierian
48 Dr. A. Giinther on a new Species of Corvina.
organs, seems rather doubtful as to their function. Below these
in the scale of nature we meet with no distinct vestiges of urmary
excreting organs; so if my hypothesis regarding the nature of
these above-described bodies be accepted, they will rank as either
the first or the second early traces of such glands as yet found
in the animal kingdom.
VII.— Description of a new Species of Corvina from the Gambia.
By Dr. ALBERT GUNTHER.
Mr. Moore, Curator of the Liverpool Free Public Museum,
has kindly sent for my inspection a Sciznoid Fish collected by
J. Lewis Ingram, Esq., at Bathurst, on the River Gambia, which
proves to be an undescribed species of the genus Corvina, for
which I propose the name of
Corvina Moorii.
). 855. A.2. L. lat. 64, I. transy. 7/z-
This species is distinguished by its broad and obtuse head,
similar to that of Collichthys. The eye is comparatively small,
about one-ninth of the length of the head, and only one-half of
the extent of the snout. Interorbital space very broad, convex,
its width being one-third of the length of the head. Hind mar-
gin of the preoperculum obliquely descending backwards, with
short spmous teeth at the angle and along the margins. Snout
very obtuse ; jaws with narrow bands of short cardiform teeth,
those of the outer series being much larger and conical. Cleft
of the mouth of moderate width, situated at the lower side of
the snout, the maxillary extending to behind the hind margin
of the orbit.
The length of the head is more than the depth of the body,
and one-fourth of the total length (without caudal). Scales of
moderate size, irregularly arranged. Pectoral fin considerabl
longer than the ventral, as long as the post-orbital part of the
head. Dorsal spines of moderate strength, not flexible; the
second is the longest, and rather more than half as long as the
head; the soft dorsal fin of moderate height. Caudal fin convex,
shghtly produced in the middle. The second anal spine strong,
two-thirds as long as the first soft ray, and nearly one-third as
long as the head.
Uniform blackish brown, the centre of each scale being lighter ;
fins black.
The specimen is 20 inches long.
Numerous species of Acanthopterygian fishes, especially from
the west coast of Africa, show osseous tumours in some parts of
Zoological Society. 49
their skeleton. The seat of these tumours is chiefly the neural
or hemal processes, more rarely the interneurals and inter-
hemals. In the typical specimen of Corvina Moorii a date-like
osseous tumour is attached to the spine of the second dorsal fin ;
and a second specimen which we have seen, from the same loca-
lity, has, singularly enough, a perfectly similar tumour on the
same spine.
We have formerly (Fishes, ii. p. 296) expressed our opinion
that these peculiar tumours are anomalous deposits of osseous
matter, and that species founded on such a character (like Cor-
vina clavigera, Cuv. & Val.), are extremely doubtful. Indeed
we have now not the least doubt that this Corvina clavigera is
identical with C. nigrita, of which we have seen an example,
likewise belonging to the Liverpool Museum, which has the
ventral and anal spines excessively thickened, in consequence of
a similarly abnormal deposition of bony substance.
PROCEEDINGS OF LEARNED SOCIETIES.
ZOOLOGICAL SOCIETY.
Jan. 10, 1865.—Dr. J. E. Gray, F.R.S., in the Chair.
On THE ANATOMY AND Hapsits oF THE WATER-OusEL (CINCLUS
Aquaticus). By Epwarps Crisp, M.D., F.Z.S, ere.
I have for a long time been occupied in preparing a work on the
British Birds, more especially in reference to their structure, in con-
nexion with their habits, the nature of their food, &c.; and there is
no bird that has puzzled me so much as the Water-Ousel, and it is
on this account that I bring the subject before the Society, hoping
that I may obtain some information from the members present. I
need not go very minutely into the history of this bird; but it will, I
think, be interesting to compare some parts of its anatomy with those
of the other Merulide. 'The object of my paper will be to endeavour,
first, to ascertain by what means this bird, so unlike all aquatic
birds in form, is enabled to dive and remain some time under water
and capture its prey; secondly, to inquire respecting the nature of
its food, and its supposed depredations on the ova and fry of fishes.
I may premise that I have shot several of these birds in Scotland for
the purpose of ascertaining the character of their food, and that I have
had many opportunities of observing their habits. The three speci-
mens on the table were sent to me recently (Nov. 30) by my friend
Mr. Grierson, of Thornhill, Dumfriesshire ; and I have dissected and
examined them, as I had done on former occasions, in relation to the
two questions above referred to. As the evidence of one inquirer «n
reference to the habits of this or of any other bird is comparatively
valueless, let me quote a few authorities upon the subject.
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. 4
50 Zoological Society :—
Montagu, in his Ornithological Dictionary, says he “ discovered
the nest of this bird in consequence of the old bird flying, with a fish
in its bill, to the young. These were nearly fledged, but incapable of
flight ; and the moment the nest was disturbed, they fluttered out
and dropped into the water, and, to our astonishment, instantly
vanished, but in a little time made their appearance at some distance
down the stream, and it was with difficulty two out of five were taken,
as they dived on being approached. ‘The motion under water,” he
says, “is effected by short jerks from the shoulder-joint, not, as in
all other diving-birds, with extended wings.”
Yarrell dissected this bird, and found nothing in its structure to
account for its diving and remaining on the ground without any
muscular effort.
Mr. Macgillivray (Naturalist, vol. i. p. 105) says, ‘ I have seen the
Dipper moving under water in situations where I could observe it
with certainty, and I readily perceived that its actions were similar
to those of the Divers, Mergansers, and Cormorants, which I have
often watched from an eminence as they pursued the shoals of sand-
eels along the sandy shores of the Hebrides. It in fact flew, not
merely using the wing from the carpal joint, but extending it con-
siderably, and employing its whole extent as if moving in the air.
The general direction of the body is obliquely downwards ; and great
force is evidently used to counteract the effects of gravity, the bird
finding it difficult to keep at the bottom.”
Other observers have given similar testimony, some asserting that
bubbles of air appeared on the surface after the bird was submerged :
but these must have arisen from the disturbance of the earth at the
bottom of the river; for no diving-bird, I believe, emits air from its
lungs when under water. The air is got rid of before the act of
diving takes place. But let me now speak of some parts of the
anatomy of this bird, before I attempt to answer the first question.
The average weight of this bird is said to be 23 oz.; but in four that
I have weighed the average weight has been about 24 0z., the males
being a little heavier than the females; the length 77 inches, and
11 inches from the tip of each wing. The brain weighed 10 grains,
the eyes 12 grains, the skin and feathers 132 grains, the pectoral
muscles 135 grains. The gizzard moderately thick, and lined with
a tough cuticle. The length of the whole alimentary tube was
16 inches ; the cesophagus, as in the other Merulide, not dilated
into acrop. The trachea of nearly uniform calibre, and consisting
of 36 rings; the vocal muscles largely developed, as in the other
members of this family. The tail-glands comparatively of large size.
I have depicted all the above parts in the drawing before the
Society ; but the parts of the anatomy of this bird to which I am
anxious to direct attention are the shortness of the wing and the great
development of the wing-muscles—features which I believe will in
a great measure account for the diving-powers of this bird and its
progress under water. As might be expected, too, from the frequent
motion of the tail, the caudal muscles are much developed. On
comparing the visceral anatomy of this bird with that of the other
~
Dr. E. Crisp on the Water-Ousel. 51
British Merulide, all of which I have dissected, with the exception of
White’s Thrush (Turdus White), very little proportional difference is
observed. The length of the intestinal tube in the Redwing (7’.
iliacus) is 14 inches; the brain weighs 16 grains, the pectoral
muscles 170 grains, the weight of the body being about 25 oz. In
the Fieldfare (7. pilaris), weighing 43 0z., the brain weighs 26 grains,
and the intestinal tube measures 22 inches. In the Ring-Ousel
(T. torquatus), weight 3 oz. 180 grains, the alimentary canal is
13} inches in length, and the weight of the brain is 26 grains; and
these parts in the Missel-Thrush (7. viscivorus), in the Blackbird
(T. merula), and Song-Thrush (7. musicus) are of nearly the same
proportionate length and weight. In the young Water-Ousel that
I have dissected, I observed nothing remarkable in its anatomy.
So that, as regards the visceral anatomy, there is no important
difference between the Water-Ousel and the other members of this
group, although among the British Merules this is the only bird
that feeds exclusively on animal food; but, to show how the habits
of a bird may be altered in this respect, I have mentioned a young
Water-Ousel that was reared under a Bantam, and fed on porridge
(P. Z. S. 1859, p. 200).
Some writers upon this bird have spoken of the claws as being well
adapted for holding on to stones and other objects at the bottom of
the water ; but on comparing the claws of the Water-Ousel with those
of the other Merulida, it will be seen that the bird has no advantage
of this kind, although the comparatively blunted form of the claw
would lead to the inference that it is used for the purpose mentioned.
The bones of the Water-Ousel, like those of the other British mem-
bers of this group, contain no air* ; and it is singular that the skele-
ton of the Fieldfare, Redwing, and Missel-Thrush (birds of passage)
should in this respect resemble that of the short-flighted Water-Ousel.
As regards the food, I am afraid that we cannot entirely acquit this
bird of occasionally destroying the fry of fish ; but I know of no reli-
able evidence to prove that it takes the ova. In the three specimens
before the Society, the gizzards of all contained Entomostraca, and
one of them a Gordian (Gordius aquaticus). In others that I have
dissected, I have discovered chiefly Entomostraca and the larvee of
Phryganea ; indeed I have found that its food is very similar to that
of the young Salmon (Salmo salar).
Mr. Gould, in his present work ‘The Birds of Great Britain’
(part 1), mentions that he examined five of these birds that were
shot on the River Usk, in Nov. 1859, and that no trace of spawn was
found in any of them; their hard gizzards were entirely filled with
the larvee of Phryganea and the Water-beetle (Hydrophilus). One
had a small Bullhead (Cottus gobio), which the bird had doubtless
taken from under a stone. Mr. Gould thinks that, by destroying
insects and their larvee that may attack the ova and fry of fishes,
these birds may do great service.
* I need scarcely say that some of the cranial bones of birds, like those of
mammals, contain air.
4*
v
52 Zoological Society :—
Mr. Macgillivray found beetles and water shells (Lymnea and
Ancylus) and the larvee of Ephemera, Phryganea, and other aquatic
insects.
Sir W. Jardine, in his ‘ Birds of Great Britain,’ says, ‘‘ In one part
of Scotland, sixpence per head is given for these birds. In another
district, 548 were killed in three years.’” He adds, “The ova of any
kind of fish we have never detected in the stomach or intestines ; nor
do we think that they habitually frequent the places where the
spawn would be deposited ; and if they did, we would deem it almost
impossible that they could reach it after it was covered in the
spawning-bed,” &c.
So that I hope we may fairly acquit this interesting little bird of
the depredations of which it has so often been accused; but I hope
that we shall ere long see the Water-Ousel, with the Little Grebe
(Podiceps minor), in the Society’s fish-house, where a better oppor-
tunity will be afforded of learning its habits.
As is well known, this bird has been variously classed by different
writers. Mr. Gould, in the work before quoted, says he regards
Cinclus as one of the isolated forms of ornithology, and that it has
some remote alliance with the genera Troglodytes and Scytalopus
and their allies. -
DescripTion oF A New Species oF ENTOZOON FROM THE
INTESTINES OF THE D1AMOND-SNAKE OF AUSTRALIA (Mo-
RELIA SPILOTES). By W. Barrp, M.D., F.L.S.
Boturipium (SoLeNnorHorvs, Creplin) arcuatum, Baird.
Length of the largest specimen (which, however, is not quite
perfect at lower extremity) 10 inches. Breadth, about the middle of
its length, 4 lines. Head, consisting of its two tubular bothria, about 7
lines in length and 3 lines in breadth. Bothria smooth, cylindrical,
arched outwardly, and connected together throughout their whole
extent, and each of about the same diameter at the top as at the
bottom. Upper openings circular and large; lower openings very
small and quite terminal. Neck none. Articulations at anterior
extremity extremely small, appearing like mere rugze. Articulations
of rest of body, in adult specimens, very numerous, narrow, much
broader than long, and crowded together ; in smaller and apparently
younger specimens (which, however, look as if perfect in length), the
articulations near the posterior extremity are, comparatively speaking,
much larger, longer than broad, and are more like those of B. laticeps
or B. pythonis. The most distinguishing character is the size and
shape of the head.
a Intestines of the Morelia spilotes, from Australia. (Mus.
Brit.).
For the specimens of this species I am indebted to Dr. A. Giin-
ther, who found them attached to the inner surface of the intestines
Dr. G. Bennett on the Lyre-Bird of New Holland. 53
of a specimen of an Australian Python, the Diamond-Snake, Morelia
spilotes.
}
Mo
<
¢ \
\\
Cg: ,
Fig. a. Worm of natural size, attached to inner surface of intestine.
Fig. 6. Bothria, slightly enlarged, showing the upper openings.
Fig. ec. The same, showing lower openings.
Fig. d. Posterior extremity of a young specimen, showing the, comparatively
speaking, larger articulations.
Jan. 24, 1865.—E. W. H. Holdsworth, Esq., in the Chair.
The Secretary read the following extract from a letter addressed
to him by Dr. Bennett, F.Z.S., dated Sydney, Nov. 18th, relating
to a living specimen of the Lyre-bird of New Holland (Menura su-
perba), which the Acclimatization Society of that city were intending
to transmit by the first favourable opportunity to this Society :—
“After repeated trials of keeping this wild and restless bird in
captivity, and having procured and lost in one year numerous living
birds of all ages, from the young bird to the adult, we have so far
succeeded as to preserve one alive and in excellent health, and feed-
ing well, since the 23rd of August last ; to this day it continues
in good health and condition. It is a young bird, at present in im-
54 Zoological Society :—
mature plumage, and the sex cannot yet be determined. It is placed
in a large wire compartment with the Talegallas or Brush-Turkeys,
and it appears to enjoy their society very much. Whether their com-
pany reconciles it to confinement I cannot say ; but, at all events, it
feeds well and thrives, and displays a great amount of activity for a
great part of the day, running about the cage incessantly, scratching
the ground. It feeds on the larva of the Tettigonia or ‘‘ Locust”’ of
the colonists, meat chopped very small, slugs, and worms. This
bird was captured at Broughton’s Pass, Illawarra district. Should
we be fortunate enough to keep it alive till the time of the departure
of the ‘La Hogue,’ it will be sent to the Zoological Society under
Mr. Broughton’s care, when it will have every chance of reaching
England alive.”
February 14, 1865.—Dr. J. E. Gray, F.R.S., in the Chair.
The Secretary read the following letter, addressed by Dr. H. Bur-
meister, of Buenos Ayres (Foreign Member), to Dr. J. E. Gray, con-
taining the description of a new species of Whale, proposed to be
called Balenoptera patachonica, together with some particulars as
to specimens of certain other Cetacea in the Museum of Buenos
Ayres.
as Gray stated, in reference to the new Whale, that it was of
much interest as being the first well-described Fin-Whale from the
southern hemisphere. Dr. Gray considered it evidently a typical
species of the genus Physalus, distinguishable from all the northern
species by the shortness of the lateral rigs compared with the dia-
meter of the bodies of the cervical vertebrze.
“ Buenos Ayres, 22nd December, 1864.
*‘T now send you drawings of the Whale in the Buenos Ayres
Museum, drawn by myself, and, as I believe, exact to nature.
“Fig 1. The skull. We have two specimens—one complete, the
other consisting only of the hinder part, without the jaws. In the
former the upper jaws are no longer in position, but separated from
the cranium, and therefore little importance can be attached to the
width of the opening between the intermaxillary bones in the ante-
rior part of the cleft between them ; it may be somewhat exaggerated.
All the other parts are entirely exact from nature, and well preserved.
“Length of the intermaxillary, 7 feet 2 inches ; length of the max-
illary, 7 feet ; length of the under jaw, 10 feet 2 inches. Breadth of
the frontal bones between the orbits, 5 feet ; breadth of the vertex
behind, 2 feet 8 inches.
«The baleen is entirely black, without any other colour. We have
two kinds in the Museum—one 53 feet and the other 1 foot 8 inches
in length. This last only may be from the Balenoptera; the other
perhaps from a Balena, because it is much more slender and more
fringed. ‘
«Comparing my drawing (fig. 1) with that of Cuvier from the Cape
Balenoptera (Oss. Foss. pl. 26. fig. 2), you will find that the
Dr. H. Burmeister on a new Whale. 55
suture between the frontal bone and the parietal is situated much
more towards the external part of the frental bone, being in my skull
Fig. 1.
Skull seen from above.
exactly in the angle where both bones are united, and therefore not
seen from above in my drawing. Another difference of the species
First cervical vertebra
is indicated by the longitudinal carina in the vertex of the Cape spe-
cies, there being no trace of such carina in either of riy specimens.
56 Zoological Society :—
‘Unfortunately the tympanic bones are wanting in both, and I
can tell you nothing of them. But the zygomatic bone is preserved,
and is of the same form as that figured in Cuvier’s work, figs. 1
and 3, but somewhat smaller than the latter figure.
“The seven cervical vertebree are free, separate from each other,
and the body of every one has the epiphyses on each side, the specimen
being that of a young individual. But in the atlas and front side of the
axis these epiphyses do not exist. I send you drawings of the first
(fig. 2), the second (fig. 3), the fourth (fig. 4), and the sixth (fig. 5)
Fig. 3.
Second cervical vertebra.
vertebrae ; the third exactly resembles the fourth, and the fifth
only differs in a small opening in the lateral arc, indicated in my
drawing of the fourth, on the left side. The seventh has no inferior
Fig. 4.
Fourth cervical vertebra.
Dr. H. Burmeister on a new Whale. 57
process at all, but a much stronger superior one, of the same form.
All the five vertebree after the second are very thin, 2 inches in
Bigs 3.
Sixth cervical vertebra.
diameter, the third being the thinnest of all, and the following ones
somewhat thicker; the seventh is 2} inches in thickness.
“Of costal or dorsal vertebrae we have fourteen, very well indicated
by the flattened end of the transverse processes being united with the
ribs. The first of these dorsal vertebree is very thin, 3 inches in
diameter ; and the second somewhat thicker, 33 inches; after these
the bodies are much stronger, from 6 to 8 inches in diameter. The
three first dorsal vertebree have transverse processes more rounded,
and directed forward. After the third they are more flat and broad,
and directed transversely to the sides. After these fourteen vertebree
foliow twelve others with thinner transverse processes, rounded and
sharp at the end, and with bodies of much larger diameter—from
10 to 12 inches. Then follows a strong vertebra, the thirteenth,
12 inches in diameter, with a smaller and shorter transverse process,
which seems to me the first caudal ; but as the epiphysis is wanting,
there is no attachment for the heemapophysis on its hinder end. In-
deed its body is flattened on the under side, not carinated as the body
of the antecedent; which also seems to me to prove that it is the
first caudal. Of heemapophyses we have four in the Museum, of un-
equal size, the first 5 inches high, the largest 8 inches, and 3 to
4 inches broad between the lamin.
“The ribs are not perfect as regards number, but the first seven
or eight are preserved. I send you drawings of the upper and lower
extremities of the first four (figs. 6, 7, 8, 9).
“The sternum is wanting, and of the os hyoideum we have only
58 Zoological Society :—
the corpus, of precisely the same form as that figured in Cuvier’s
Oss. Foss. pl. 25. f. 14.
Fig. 6. Rick 7. Fig. 8. Fig. 9.
“Of the pectoral fin we have only the scapula, of which I send
you a drawing (fig. 10); both processes are well developed and some-
what compressed.
Fig. 10.
A Uf
Ui
AYN
Y yy)
YY
SS Mi,
Scapula.
‘The animal was found some leagues from Buenos Ayres, on the
banks of the River Plata, where it came ashore some thirty years
ago. It was brought to the gardens of Rosas, at Palermo, where the
skeleton was exhibited a long time, till, after the fall of the tyrant,
it was transferred to the Museum. The parts now deficient were
then lost.
‘«T suppose that the species might be the same as that you have
indicated in your synopsis as Balenoptera australis, Desmoulins
(Voy. Ereb. and Terror, Mamm. p. 20); but as I have never seen
Dr. A. Giinther on the British Salmonoids. 59
that animal, Iam unable to speak concerning its external appearance.
Therefore I believe it is better to describe the species in question
under a new name, and I propose to you, if you please to accept it,
that of Balenoptera patachonica.
“« Since I have received the excellent books you sent me, and for
which I give you my best thanks, I have found in them figures of the
two skulls of Dolphins in the Buenos Ayres Museum. ‘The larger
is your Delphinus Eurynome (p. 38, pl. 17), and the smaller your
Delphinus microps (p. 72, pl. 25). Both are inhabitants of the
Atlantic in our latitude. The new Phocena is wanting in your
list. I propose to give the name Phocena spinipinnis to it, from the
numerous spines on the dorsal fin. We have the entire animal, with
the skull, which I will examine when it is taken from the dry skin
in which it is enclosed. By the next French steamer I will send you
an accurate drawing and complete description of it.”
*¢ P.S.—I have told you nothing of the under jaw of Balenoptera
patachonica, because the surface of the bone is much destroyed by
long exposure to the air, rain, and sun; but the hinder part, with
the coronoid process, is represented in fig. 11.”
A letter was read, addressed to the Secretary by Prof. J. J. Bian-
coni, of Bologna, stating that, in the course of researches upon the
osteology of the extinct genus pyornis, he had come to the con-
clusion that that form belonged to the Vulturidze, and not to the
Struthious birds.
Dr. A. Giinther gave an account of the present state of his re-
searches into the British species of Salmonoid fishes, which he had
undertaken whilst engaged in preparing the catalogue of the speci-
mens of this family in the collection of the British Museum. Dr.
Giinther stated that the genus Salmo was essentially an arctic group,
inhabiting the northern portions of both hemispheres, and becoming
more abundant in species upon receding from subtropical into tem-
perate latitudes. Dr. Giinther was disposed to believe that the
species of this genus to be found within British waters would be
ultimately found to be much more numerous than had been hitherto
suspected. From the materials at present at his command, he had
already been able to distinguish what he believed would turn out to
be four new species of the non-migratory group of true Salmo, be-
sides identifying several others heretofore imperfectly distinguished.
60 Zoological Society :—
Dr. Giinther requested the assistance of the Fellows of the Society
and their friends in furnishing him with series of specimens of our
native Salmons and ‘T'routs from every part of the British islands,
stating that in this difficult group of fishes no certain conclusions
could be arrived at without a large number of specimens for com-
parison. Dr. Gunther exhibited the subjoined table as giving a list
of the British species of Salmo with which he was acquainted :—
Subgenus I. Cuarrs (Salvelini).
IP ULligOtt 7 25.5 as eet Windermere, &c.
DE MRORUSIU. & toidiarits hone A ile Llanberris Lakes, N. Wales.
Se LOPUUB eid on cts oh Scotland.
AIG OY Us a hecho th eco thues ae Lough Melvin, Ireland.
Caleta sr, dolecs| sus cael ohe Lough Eske and Lough Dan, Ireland.
Subgenus II. Satmons (Salmones).
a. Migratory Species.
1A 77 el a ere ieee True Salmon of British rivers.
QUCOMOTACUBY wiles st ons coe **Sewin”’ of South Wales.
BS METUCED bin eet ee eee “Sea Trout ”’ of Scotland.
b. Non-migratory Species.
Gil Ras Be eS ane es England.
DENG QUNGTOAU sche uetegones oe Scotland and N. W. England.
6. nigripinnis, sp. nov. ..Mountain-lochs of Wales (and Scotland).
TeuLOVENENSIS: “ae Aca hotles Loch Leven, Scotland.
BIN C100 yee ft sna an ae «Gt. Lake Trout”’ of Scotlandand Wales.
9. orcadensis, sp.nov... .. Lakes of Orkneys.
10. brachypoma, sp.nov. . . Firth of Forth.
11. stomachicus, sp. nov. . . Ireland. .
Description oF Two New AvstTRALIAN Birps.
By Joun Goutp, Esa., F.R.S., eve.
1. MALURUS LEUCONOTUS.
The entire head, neck, under surface, rump, and tail deep blue ;
back, shoulders, greater and lesser wing-coverts, and secondaries silky
white; primaries brown; bill black; feet brownish black.
Total length 53 inches; bill 5; wing 2; tail 34; tarsi 4.
Hab. Interior of Australia ; precise locality unknown.
Remark.—In size this new species is very similar to M. Laméerti,
while in its colouring it assimilates to M. leucopterus ; from both,
however, it may be at once distinguished by the whiteness of its
back, which has suggested the specific name I have assigned to it.
2. ARTAMUS MELANOPS.
Lores, face, rump, and under tail-coverts black ; stripe over the
eye, ear-coverts, sides of the face, throat, and under surface delicate
vinous grey; two middle tail-feathers black, the remainder black
Mr. A. Newton on some Bones of Didus. 61
largely tipped with white; upper surface of the wings grey, their
under surface white; bill leaden grey, darkest at the tip; feet
blackish brown.
Total length 63 inches ; bill? ; wing 42; tail, 3; tarsi 3.
Hab. Central Australia.
Remark.—This large and fine species is unlike every other known
member of the genus. It is most nearly allied to 4. albiventris, but
differs from that bird in the jet-black colouring of its under tail-
coverts, and from A. cinereus in its smaller size and the greater
extent of the black on the face. The specimen from which the
above description was taken has been kindly sent to me by Mr. S.
White, of the Reed-beds, near Adelaide, South Australia, who
informs me that it was shot by him at St. Becket’s Pool, lat.
28° 30’, on the 23rd of August, 1863, and who in the note accom-
panying it says, “‘I have never seen this bird south. It collects at
night, like 4. sordidus, and utters the same kind of call. It seems
to be plentiful all over the north country. I saw it at St. Becket’s
Pool, feeding on the ground, soaring high in the air, and clinging in
bushes, like the others. The two sexes appeared to be very similar
in outward appearance. The stomachs of those examined were
fleshy, and contained the remains of small Coleoptera.
ON SOME RECENTLY DISCOVERED BoNES OF THE LARGEST
KNOWN Species oF Dopo (Dipus nazAreNnvus, BARTLETT).
By ALFRED Newton, M.A., F.L.S., F.Z.S.
The three bones which I now have the pleasure of exhibiting
have been recently received by me from my brother Mr. Edward
Newton, a Corresponding Member of this Society, who himself found
two of them in a cave on the south-west side of the island of Ro-
driguez, which he visited on the 2nd of November last. The third
was obtained on the same island, about the same time, by Captain
Barkly, a son and aide-de-camp of the Governor of Mauritius. All
three belong, without doubt, to the largest known species of Dodo,
to which Mr. Bartlett (P. Z.S.1851, p. 284) applied the name Didus
nazarenus, and which was so unaccountably overlooked by Messrs.
Strickland and Melville in their excellent monograph of the curious
group Didine. These authors, as Mr. Bartlett showed (oc. cit.), did
not distinguish between this very large bird and the smaller and more
slender “Solitaire”? (Pezophaps solitaria), which, if we are to trust
the evidence before us, was, equally with Didus nazarenus and D.
éneptus, an inhabitant of Rodriguez.
The two bones found by my brother were picked up near the en-
trance of a very dry cave, where little, if any, stalagmitic deposit was
forming, at least at the time of his visit. One is a perfect left tarso-
metatarsus, and the other a left humerus, wanting its extremities, as
is so often the case in specimens of this bone found under circum-
stances which lead to the belief that the bird to which it belonged
had been eaten by men or dogs.
The bone found by Captain Barkly is a right femur. Though
62 Zoological Society :—
nearly perfect, it seems to have been much exposed to the action of
the weather, and, in consequence of its condition, it has sustained a
little damage by the crumbling away of some part of its extremities.
This has probably happened since its discovery ; but one advantage
results from the circumstance—namely, that the cellular structure
of the bone is thereby rendered plainly visible.
I proceed to give the dimensions of these specimens, and, for con-
venience of comparison, I shall, as far as possible, follow Dr. Mel-
ville’s plan of measurement (‘The Dodo and its Kindred,’ page 116).
Fragment of left Humerus.
inches. lines.
iliransverse diameter’ of shaft). sir. ./.:~ 0 o0-te an sin tare
Antero-posterior diameter of shaft ........ vlie cre tote pietalepll amma
Left Tarso-metatarsus.
Length from middle trochlear groove to inter-condyloid
tubercle yetirc eet storia sonia ie sire rete ee ees ore
external trochlear to external condyloid fossa 6 4
—— internal trochlear to internal condyloid fossa 6 = 7
Breadth of upper extremity’... 3. osc sae coe eesti 1 6
Antero-posterior diameter of the same.............. ele ao
Breadth of lower-extremity ........ 60:2... eke alist aaa 7
Projection of ento-calcaneal process .........-+6.2-005 0°. 8
Right Femur.
Length from inter-condyloid notch to upper surface of neck 6 0
upper edge of trochanter major to external 6 9
CONG. 41. ate
Transverse diameter Of Shalt scr: <5 <1) che 1-10) wel «oie 0-10
Antero-posterior............ Pe te BO wie sis siecle OMT:
Transverse diameter of upper extremity ...........-... 2 02
Transverse diameter of lower extremity ...........+.. ee)
All these specimens, unlike those in the Paris Museum, are en-
tirely free from incrustation.
I believe there are no other examples of the humerus and femur
of this species in this country. The specimen of the tarso-metatar-
sus figured in illustration of Mr. Bartlett’s paper, to which I before
referred (P. Z. 8. 1851, Aves, pl. xiv. fig. 1) is, as I learn from Mr.
Gerrard, now in the British Museum, and there are other examples
of it in the Andersonian Museum at Glasgow.
I must here tender my thanks to Mr. W. K. Parker for the kind
assistance he has rendered me in accurately measuring these bones.
And now I wish to make one suggestion. It is well known that
at Oxford there is an old picture of a Dodo, painted by one of the
Saverys, which seems hitherto to have been referred without hesi-
tation to Didus ineptus. Mr. Strickland, in speaking of it, says :—
« A remarkable feature in it is its colossal scale, the Dodo standing
about 3 feet 6 inches high, and being double the size which the
Dr. J. E. Gray on a new Species of Bush-Goat. 63
picture in the British Museum, the description of eye-witnesses, and
the existing remains warrant us in attributing to the bird. It is
difficult to assign a motive to the artist for thus magnifying an
object already sufficiently uncouth in appearance’’ (‘The Dodo,’
&e. p. 31). Is it not possible that the artist may in this painting
have taken a life-sized portrait of the large species (Didus nazarenus,
Bartlett) to which these bones belong ?
In conclusion, I have to state that I should be very glad if these
remarks were the means of exciting further search for the remains
of the Dodo and its allies. In Rodriguez the bones must be far
from scarce, and, as the present instance shows, they may be found
with little trouble. My brother picked up two of them, as I have
said, in a cave during a very hasty visit. It is a matter of the
greatest regret that a regularly organized search is not instituted by
some resident in that island, or by some visitor to whom time is no
object. We may depend upon it that a rich reward awaits the care-
ful explorer of the Masearene caverns and alluvial deposits.
Norice oF THE SKULL OF A New Species or Busu-Goar
(CrEPHALOPHUS LONGICEPS), SENT FROM THE GABOON BY
M. Du CuHaitiv. By Dr. J. E. Gray.
M. Du Chaillu has lately sent to the British Museum several skins
and skeletons of the Gorilla (showing how abundant it must be at
the Gaboon), the skin and skeleton of a Chimpanzee, three skeletons
of the African Manatee, and the head of a Bush-Goat or Cephalo-
phus.
: The skull of the Cephalophus on examination proves quite distinct
from any that has previously occurred to me; and as it indicates the
existence of a large species of the genus, I have sent a notice of it to
the Society in hope that we may before very long have a complete
specimen of the animal to describe.
CEPHALOPHUS.
Section I. Horns decumbent.
CrPHALOPHUS LONGICEBRS.
The skull elongate ; face elongate, compressed in front of the
eyes ; the nose in front of the eyes narrow, sides only very slightly
tapering; nasal bone very long, produced between the frontal be-
hind, much longer than the medial suture of the frontal. The horns
elongate, conical, diverging at the tips, decumbent, in a line with the
forehead ; forehead convex between the orbits.
Length of skull 10 inches 9 lines; width at zygoma 4 inches
7 lines ; length of horn-cores 5 inches ; length of lower jaw 9 inches.
The only species with which the animal can be compared, on ac-
count of its size, is C. sylvicultriz ; but the skull of the latter is
short and ventricose, and that of C. longiceps is elongate and slen-
der. The face of C. sylvicultriz is short, and the nose between the
impression for the suborbital glands broad and tapering; the fore-
64 Zoological Society :—
head is much more convex and rounded. The following are the mea-
surements of the skull of an adult male :—Length of skull 10 inches
1 line; width at zygoma 4 inches 7 lines; length of lower jaw
8 inches 9 lines.
The skull of C. longiceps resembles in general form and some
other particulars the figure of the skull of the male C. altifrons,
figured by Dr. Peters (Reise n. Mossamb. t. 38. f.1). But that skull
is not above half the size of the one here described; and the form
of the core of the horns is different, the one being conical and elon-
gate, and the other angular and converging at the tip.
Skull of Cephalophus longiceps.
The skulls of the larger species of Cephalophi may be divided into
two groups, according to the position of the horns, as compared with
the frontal line.
In some the horns are decumbent and bent back, being nearly in
a line with the forehead, as in Cephalophus coronatus, C. sylvicul-
trix, C. Ogilbyi, C. natalensis (figured in Cat. Ungulata, B.M. t. x.
f. 1), C. longiceps, and C. altifrons, Peters. In others the horns
are ascending, placed at an obtuse angle with regard to the line of
the forehead, as in Cephalophus Grimmius and C. ocularis of Peters
(Reise nach Mossambique, Saugeth. t. 39, 40).
The forehead in all the Cephalophi with decumbent horns is con-
vex and rounded; but in C. Ogilbyi itis very much rounded—more
than in any other species I know; it is much higher than the base
of the horn. In the species which Dr. Peters has called C. altzfrons it
does not appear to be so high as usual in the genus. In C. Grimmius,
with ascending horns, it is flat between the eyes. The following
observation is founded on the comparison of a series of skulls of
males :—The skulls differ in the length of the face, thus:—In C.
natalensis the face is short; the distance from the orbit to the upper
end of the intermaxillary bone is shorter than the length of the in-
termaxillary bone. In C. sylvicultriz, C. Ogilbyi, and C. ocularis
the distance above defined and the length of the intermaxillary are
nearly equal. In OC. Grimmius they are rather longer. In C. longi-
ceps the distance from the front edge of the orbit to the tip of the
Miscellaneous. 65
intermaxillary is much longer than the length of the intermaxil-
lary.
in some skulls the nasal bones are the same length as the upper
suture of the frontal one, as in C. natalensis, C. sylvicultrix, and C.
Ogilbyi. In C. altifrons, according to Dr. Peters’s figure, they are
shorter. In C. coronatus and C. rufilatus they are much shorter—
only about two-thirds the length. In one skull of C. Grimmius they
are longer, and in another skull shorter, and in C. longiceps much
longer.
The above observations are made only on a few, sometimes only on
one specimen of the species; and when I have three or four speci-
mens of the same species, as is the case with C. Grimmius, the skulls
present some variations in the form of the nasal bones and in the
length of the intermaxillaries as above noted.
Dr. Peters figures as the skull of a young female of C. altifrons
a skull of a very different form from that of the skull with the horns
of the male above referred to. I have not observed such a difference
in the skulls of the females of any of the species of Cephalophus that
have occurred to me. I have some doubt if it does belong to the
same species, as the figure of the young female animal is very like
the skull of a female C. Grimmius, which is an animal that has ascend-
ing horns in the male.
MISCELLANEOUS.
On the Pollen-grains of Ranunculus arvensis.
By GeorGe GuLLiver, F.R.S.
FINDING, on reference to my note-book entries (of no less than five
different examinations in the course of four years), that the pollen-
grains of Ranunculus arvensis always appeared to differ remarkably
from those of its British allies, I have recently examined the pollen
of these plants again. The difference now to be described appears
so constant and remarkable as to deserve a place in the descriptions
of this species.
The examinations include all the British yellow-flowered Ranun-
culeze with divided leaves, except R. parviflorus. This species I
have not seen growing. All the others are as common about Eden-
bridge as elsewhere. Even R. hirsutus, which Prof. Babington
marks ‘Waste land and corn-fields, rare,’ grows abundantly in
patches in some of our lanes or by-roads; but happily the very
noxious weed I. arvensis scarcely intrudes into pastures, though it
is a sad pest in some of our stiff arable land, and too well known to
our husbandmen under the name of the “ hedgehog.”
The pollen of each species was repeatedly compared in the same
stage of development—a necessary precaution, the neglect of which
has too often led to perplexing discrepancies in botanical descriptions.
In the following measurements the average sizes only are mentioned,
as made from the pollen shaken out of the anthers on to a dry piece
of glass, and viewed by transmitted light.
Ann. & Mag. Nat. Hist. Ser. 3. Vol. xvi. ay
66 Miscellaneous.
Ranunculus auricomus: pollen-grains round and smooth, and
sioth of an inch in diameter.
R. acris: pollen-grains round and smooth, and =}>th of an inch
in diameter.
R. repens: pollen-grains round and smooth, and
in diameter.
R. bulbosus: pollen-grains round and smooth, and 7},th of an
inch in diameter.
R. hirsutus: pollen-grains smoothish, with three depressed scars,
and <1,th of an inch in diameter.
R. arvensis: pollen-grains round, rough, and so much larger than
those of the other species as to measure ;4,,th of an inch in diameter.
The roughness remains when the pollen-grains are treated either with
dilute acids or water.
Hence the roughness and comparatively large size of the pollen-
grains of R. arvensis are very evident, and this curious difference is
certainly constant in our plants. It may be easily seen under a
magnifying power of fifty diameters. When much more magnified,
some inequalities may appear on the surface of the pollen-grains of
the five preceding species. An examination of the pollen of R. parvi-
florus would be interesting.
" ;
<b zth of an inch
On the Feathers of Dinornis robustus, Owen.
By W.S. Dauras, F.L.S., Keeper of York Museum.
The acquisition by the Yorkshire Philosophical Society of a spe-
cimen of Dinornis robustus, Owen, in so perfect a state of preserva-
tion that it retains even portions of the muscular and integumentary
systems, enables me to describe at least a part of the structure of the
feathery covering of this remarkable bird, and thus to throw some
further light upon its affinities among birds with which we are ac-
* quainted in the living state. The general condition of the skeleton
was described by Mr. Allis in a paper read before the Linnean Society
in June last ; and Professor Owen has since made use of one or two
portions of it for the completion of his description of the species, in a
paper communicated to this Society ; but the fact of the occurrence
of the feathers, however imperfect, of a bird which, as far as our in-
formation goes, has long been extinct, seems to call for some special
notice.
At first sight, indeed, it would seem that the fresh condition of
many parts of this skeleton, and the preservation of traces of the soft
parts, might warrant us in supposing that many years have not
elapsed since the bird to which it belonged wandered over the hills
of Otago; but all possibility of drawing from these circumstances
any conclusions as to the period of its death is set aside by the fact
that other parts of the skeleton are in a state of decay which would
apparently require a free exposure to the weather for many years for
its production.
The portion of skin which bears the remains of feathers covered
the greater part of the flat, rhombic region of the pelvis immediately
above the commencement of the tail, and extended, on the left side,
Miscellaneous. 67
beyond the ridge bounding this part of the pelvis, and for some dis-
tance down the slope of its side, where it has beneath it the aponeu-
rotic portion of some of the great muscles of the thigh. The feather-
bearing portion forms a sort of broad, irregular, transverse band
across this region of the pelvis, encroached upon anteriorly by a wide
semicircular notch, and posteriorly, a little to the right of the centre,
by an irregular worn space exhibiting numerous perforations, indica-
ting the former positions of feathers which have disappeared. The
skin itself is rather thick and coarse. The remains of feathers occur
only on that part of the skin which covered the flat back of the pelvis,
in which their insertions give rise to strongly marked papille. The
skin on the sloping left side of the pelvis bears no feathers, and pre-
sents no traces of their insertion. It appears, however, to have lost
some of its outer layers, and certainly does not furnish evidence suf-
ficient to prove the existence of a featherless space at this part, which
would be opposed to Nitzsch’s description of the pterylography of
the Struthionide.
The feathers are all very imperfect, consisting only of the basal
portions of the shaft and accessory shaft, with here and there some
traces of the barbs. The latter occur most abundantly towards the
left side, and especially in the feathers situated upon the left ridge,
from which the specimen here figured (fig. 1) was taken. The shafts
are always evidently imperfect ; the longest fragment existing in the
skin is only about 2 inches in length. The stem tapers gradually,
the quill being the widest part and about .),th of an inch in dia-
meter. The quill is inserted about ;3,ths of an inch into the skin,
and the webs appear generally to have commenced about ;zth of an
inch from the junction of the quill with the shaft. From these data
it is of course impossible to form any opinion as to the original length
of the feathers.
The accessory shafts are considerably smaller than the main shafts,
but still of sufficient size to constitute an important portion of the
plumage. The longest accessory shaft that I have been able to find
measures 13 inch in length, and is imperfect; there is little doubt
that the accessory shafts were both shorter and more slender than
the true feathers.
The shaft is somewhat convex above, and marked with a fine lon-
gitudinal furrow beneath. It is of a brown colour beneath, but pale
horn-colour above, probably from exposure to external influences.
The accessory shaft is of a pale horn-colour, and appears to be nearly
cylindrical.
The structure of the web is somewhat different from that which
occurs in the Emu and the Cassowary. Towards the base of the
shaft the barbs spring in groups of four or five together from nearly
the same spot, and thus this part of the web assumes a tufted aspect.
As we advance towards the apex this arrangement speedily ceases ;
the number of barbs springing from the shaft gradually diminishes,
until each side bears only a single series of these appendages. The
barbs consist of slender, flattened fibres, bearing long, silky, and very
delicate barbules, without any trace of barbicels, but presenting a di-
5*
68 Miscellaneous.
stinctly beaded appearance when examined by a simple lens. Under
the microscope, with a moderate power, this beaded aspect is lost,
and the barbule appears merely divided by faint transverse partitions
into a series of cells, some of which, towards the apex, exhibit small
tooth-like projections representing the rudiments of barbicels (fig. 3).
All the barbs remaining on the feathers appear to be imperfect.
Fig. 1. The basal portion of a feather detached from the skin, of the natural size :
a. The accessory shaft.
2. Part of a barb with the barbules ; magnified 15 diameters.
3. Apical portion of a barbule ; magnified 150 diameters.
The barbs of the accessory plume are of the same general struc-
ture as those on the main shaft, but they appear to form a single
series on each side from the base.
The barbs nearest the base of the feather, both in the main web
and the accessory plume, are destitute of barbules for some distance
from their base; but this distance gradually decreases until the barb
is furnished with barbules throughout its whole length.
It is evidently impossible to determine from these mere fragments
of feathers what was the precise structure of those organs when per-
fect; we cannot even decide whether the basal barbs possessed the
Miscellaneous. 69
hair-like tips characteristic of those of the Emu and Cassowary, and
still less whether the apical portion of the feather supported simple
barbs such as occupy that position in those birds. The only fact of
importance, indeed, that I can hope to make known by this paper is
that the Dinornithes undoubtedly possessed a large accessory plume,
thus adding another proof of their relationship to the green-egged
Emus and Cassowaries existing in the Australian region, and of their
difference from the white-egged group of Struthiones represented
in Africa and South America.—Proc. Zool. Soc. March 14, 1865.
On the Metamorphoses undergone by certain Fishes before acquiring
the Adult Form. By Proressor AGASSIZ.
I have lately observed in Fishes metamorphoses as considerable as
those known to take place in the Amphibia. Now that pisci-
culture is followed with so much success and on so large a scale,
it is surprising that this fact has not been long since observed ; but
this may perhaps be attributed to the circumstance that these
metamorphoses usually commence after the hatching of the young,
at a period when they die rapidly, if kept in captivity. At this age,
moreover, they are for the most part too small to be conveniently
studied in their natural element. Nevertheless this is the most im-
portant period of their growth, if we wish to study their natural
affinities. I intend shortly to show how certain small Fishes, at first
resembling Gadoids or Blennioids, pass gradually to the type of the
Labroids and Lophioids. I shall also be able to show how certain
embryos resembling the tadpoles of the Frog or Toad, gradually ac-
quire the form of Cyprinodonts,—how certain Apodal Fishes become
transformed into Jugular and Abdominal Fishes, and certain Mala-
copterygians into Acanthopterygians, and, lastly, how we may found
a natural classification of Fishes upon the correspondence existing
between their embryonic development and the complication of their
structure in the adult state.
Quite recently I have discovered that the metamorphoses of certain
members of the Scomberoid family are perhaps still more unexpected
than any of those which I have previously observed. Every ichthyo-
logist knows the characters of the Dory (Zeus faber), and the pecu-
liarities which connect this fish with the family of the Scomberoids.
Another less-known but very curious fish, drgyropelecus hemigymnus
(Cocco), which likewise inhabits the Mediterranean, has been gene-
rally referred to the Salmon family, or placed with the Salmons as a
subfamily. Systematic authors have generally regarded the Scom-
beroids and the Salmons as very different fishes, the former being
referred to the Acanthopterygii and the latter to the Malacopterygil.
Nevertheless <Argyropelecus hemigymnus is neither more nor less
than the young state of Zeus faber.
I expect that all ichthyologists will reject this assertion as erroneous.
Nevertheless nothing can be more true; and therefore, instead of
seeking to prove it by long arguments, I shall, for the present, merely
request my confréres to procure small specimens of the Dory (of 8
to 10 centimétres in length), and to compare them with authentic
spesimens of Argyropelecus, feeling certain that they will admit the
70 Miscellaneous.
identity of the two fishes as soon as they have made the compari-
son.—Comptes Rendus, January 23, 1865, p. 152.
Description of the Egg of Parra gallinacea.
By Joun Gouxp, F.R.S. &e.
The ground-colour of the egg of this species is of a dark shining
raw-sienna tint, over which are traced in various directions a series
of broad and fine hair-like contorted lines of brownish black, which,
by occasionally uniting laterally and crossing each other, form here
and there large blotches. Although these markings are of the same
character on each egg, they are somewhat differently distributed :
thus, on one of the two I possess, they are more numerous at the
larger end, and absent at the smaller; while on the other they are
more abundant at the smaller, and less so at the larger extremity.
The eggs are one inch and an eighth in length by seven eighths of an
inch in breadth. They are, moreover, rendered remarkably conspi-
cuous by the singularly pointed form of the smaller end, and by their
small size as compared with that of the bird, but above all by the
form and disposition of the markings, which are as if traced by the
hand of a person who had amused himself by attempting to cover the
surface with fantastic streaks, blotches, and contorted curves from
end to end.— Proc. Zool. Soc. Dec. 13, 1864.
On a new Form of Brachiolaria. By M. Sars.
M. Sars has discovered a new Echinodermatous larva belonging to
the Brachiolarian type. It presents a greater affinity to the Bipin-
narie than those observed by Johannes Miller. Its development is
also very similar to that of the Bipinnaria,—the Starfish mm course of
formation presenting the same relations of position and union with
the body of the larva. There are, however, some differences. In
the Bipinnarie the rudiment of the ambulacral system makes its
appearance very early, in the form of a rosette of five czeca; in the
Brachiolaria, on the contrary, these cxeca are not brought together
in a group, but distant from each other, and their circle is open on
one side. This condition persists until after the formation of the
perisoma, with its five arms and their spines.
The Brachiolarie are really distinguished from the Bipinnarie
only by the presence of their contractile arms at the anterior extremity.
M. Sars has ascertained that these organs, whose function has hitherto
been doubtful, act as an apparatus of attachment. They may be com-
pared with the very similar organs of attachment of the larvee of
Wchinaster sanguinolentus and Asteracanthion Miilleri. ‘Thus these
various types of larvee, so different in appearance, are united in an
unexpected manner.—Videnskabsselskabets Forhandlingar, 1863 ;
Abstract in Bibl. Univ., May 1865, Bull. Sci., p. 62.
Investigation of the Structure of the Encephalon of Fishes, and of the
Homological Signification of its different Parts. By M. Houuarp.
The type of the encephalon in Fishes is inferior to that prevailing
in Mammalia, not only in its general development, but also in the
absence of several organs. This typeis not only inferior, but it is
Miscellaneous. ra |
also special, and susceptible of numerous modifications. The pecu-
liarities presented by it have given rise to very various interpretations,
often founded on deceptive analogies of form, and most of them in-
capable of demonstration. With the exception of some determina-
tions which are evident at the first glance, most of them are still in
the condition of simple hypotheses, or mere probabilities, or are
completely erroneous. In order to arrive at results such as science
can accept, we must (1) commence by referring the different parts of
the encephalon of Fishes to the divisions which embryogeny furnishes
for this brain as for those of other Vertebrata, and (2) ascertain with
precision the organs composing each of these divisions, and distin-
guish among these organs those which are fundamental and those
which belong to the development of higher types.
By proceeding in this manner, tracing the series of three ence-
phalic regions which correspond successively to the three primitive
cerebral vesicles of vertebrate animals (the epencephalic, mesen-
cephalic, and prosencephalic), and ascertaining that the epencephalon
is divided into two subregions, namely, that of the postcerebrum or
calamus and that of the cerebellum, and that the prosencephalon is
also divided, forming an anterior and an intermediate cerebrum, the
author found no difficulty in determining, at first in a general way,
but afterwards in detail, the cerebral organs which are developed in
each of these regions in Fishes.
For the postcerebrum, or region of the calamus, we have here two
pairs of small grey masses, superposed upon the roots of the fifth,
eighth, and ninth pairs of encephalic nerves. These little lobules,
commonly known as the posterior lobes or lobi vagi, correspond to
the streaks of grey matter which border the fourth ventricle in the
higher Vertebrata, and especially the posterior pyramids, forming
what are called the valves of Tarin.
The posterior brain, which usually forms a large single lobe sup-
ported by two lateral peduncles and emitting two anterior processes,
is a well-characterized cerebellum; and by contrasting with that of the
Batrachia and Reptiles, but more or less resembling that of Birds,
it constitutes one of the features of the cerebral type of Fishes.
The mesencephalon is concealed beneath the posterior part of the
prosencephalon. It is composed of tuberculiform masses seated on
a floor which covers a true aqueduct of Sylvius: it is here that the
processes of the cerebellum terminate; and we may easily recognize
the tubercula geminata in the hollow grey masses which cover this
small ventricular region.
The greatest difficulty of determination is presented by the pros-
encephalon, and especially by the posterior subregion or intermediate
cerebrum. This region, more complex than the others in all Verte-
brata, presents peculiar arrangements in Fishes; and to decipher it
we must recall the constitution of the intermediate cerebrum in the
higher Vertebrata. It is at once the peduncular region, the region
of the third ventricle, of the cerebral nucleus—or at least of the
fundamental portion of that nucleus, which is composed of the pedun-
cular fasciculi, the optic lobes, and the striated bodies.
In Fishes we readily detect this peduncular region, composed
Te Miscellaneous.
inferiorly of two large pyramidal fasciculi, superiorly of the continua-
tion of the other fasciculi of the medulla. Two pairs of lobes, one
inferior, one superior, are attached to it. The superior lobe is com-
posed of a semicircular nucleus, from which originate two superposed
layers, the fibres of which cross, and which give to the lobe a great
superficial development. The outer layer is the principal root of the
optic nerve. These can only be regarded as the optic lobes.
The inferior lobes are more problematical. The inferior pyramidal
cords are divided between these and the preceding lobes ; this was
already known. But the other medullary fibres, after traversing the
optic lobes, also penetrate the inferior lobes, instead of passing
directly, as has been supposed, to the anterior cerebrum. Another
newly ascertained fact is, that it is from the inferior lobes that the
medullary fasciculus originates which spreads in the anterior lobes of
the cerebrum in Fishes; so that the true serial position of the in-
ferior lobes is that of a continuation of the optic lobes, and preceding
the hemispheres. This position is occupied in the higher animals
by the corpora striata, with which the inferior lobes of Fishes may
therefore be identified. —Comptes Rendus, April 17, 1865, p. 768.
Description of a new Species of Rock-Kangaroo (Pterogale longi-
cauda) from New South Wales. By Grrarp Krerrt.
Hair remarkably soft and long (3 inches in length upon the back
and sides), dark grey at the base, tipped with pale yellow and
black, giving the fur a mottled appearance. Head and neck grey,
a lighter patch extending from the base of the ears to the nostrils.
Ears grey at the base, black at the tip ; sides slightly fringed with
yellow. Shoulders and fore legs dark grey, grizzled with white,
which colour extends to about the middle of the body. The hair of
the back and haunches is of much longer growth, silky to the
touch, of a mottled brownish-grey colour, and changing into rusty
yellow near the base of the tail. The tail at its root is sandy-coloured,
but soon changes into dark brown, the hair being very coarse and
long, forming into a broad brush at the end.
Inches
Length from tip of nose to root of tail.......... 294
Dail”, Seder re ees a St ake chats es clea sy ovcre ke Sate neurone 3 Oo
Facerto base Ot, Carts...) are ose trem oeecre ee fet
Arms and ands: 5.4: acc uacite chet dies «0 eaoeiee 6
Marsivand GCS: otc ses eM cele neue euos Ree *, ff
Barts teh fsa? ONE scans aches denies sae ee ee 21
Petrogale longicauda is easily distinguished from all other species by
its remarkably long and bushy tail, which is about a foot longer than
that of any other Rock-Wallaby. A single specimen of this inter-
esting animal has been procured by Mr. George Masters, Assistant
Curator of the Australian Museum, at Dabee Rylstone, 250 miles
N.W. of Sydney. Mr. Masters informs me that this Wallaby is
very quick and difficult to approach, and that, after watching for
two nights, only one specimen could be secured. The skull was
completely broken, so that no description could be given of it.—
Proc. Zool. Soc. March 28, 1865.
THRE ANNALS
MAGAZINE OF NATURAL HISTORY.
) [THIRD SERIES. ]
No. 92. AUGUST 1865.
VIIL.—On the Homology of the Buccal Parts of the Mollusca.
By Dr. Orro A. L. Mércu, of Copenhagen.
[Plate VI.]
Tue oral organs have, throughout the animal kingdom, furnished
some of the most important systematic characters. Lianzeus first
based the system of Mammalia, and Fabricius that of Insects, upon
the structure of these parts. It was, however, not until 1847 that
Prof. Lovén* placed the natural classification of the Mollusca on
a scientific base, chiefly founding it on the dentition of the tongue
(radula). Another part of the oral organs, the mandibles, has
been much neglected. Most authors understand by this name
all hard bodies near the entrance of the mouth. I believe it is
necessary to distinguish two (or perhaps three) different kinds
of oral plates corresponding to the mandibles and maxillie of the
Arthropoda. The maxilla is a median, unequal-sized, corneous
plate attached to the bulbus pharyngeus over the oral aperture,
and serving to divide the food into morsels. It is found in all
Land Pulmonata (Phyllovora, Gray). According to its struc-
ture, I have distinguished the following groups :—Ozygnatha,
with a smooth maxilla; Auwlacognatha, with a closely suleated
maxilla, crenulated at the edge; Odontognatha, with a strongly
ribbed maxilla, forming projecting teeth on the edge; Gonio-
gnatha, with the maxilla composed of oblique plates (genus
Orthalicus)+. Among the Land Pulmonata, the maxille are
wanting in the Agnatha (Vermivora, Gray), which swallow their
prey entire and alive (genera Onciis, Testacella, Helicophanta,
Caffra, Daudebardia, Streptaxis, Urocoptis, Glandina). Among
the marine Mollusca, it is only found in A¥gzrus and, perhaps,
Siphonodentalium.
nF Ofversigt af Kgl. Vetenskaps Akademiens Forhandlingar, 1847.
+ Morch, Malacozoologische Blatter, 1859.
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. 6
74 Dr. Morch on the Homology of
The maxilla of the aquatic Pulmonata is provided with a lateral,
linear, moveable appendage on each side, the real homology of
which is not yet quite ascertained, nor even whether it is a part
of the maxilla or independent of it.
The maxilla is divided into two* in the case of the young Limax
before it leaves the egg. The two halves are afterwards united
by an intermediate piece, which, in a more advanced period of
growth, is indicated by a notch in the projecting middle tooth.
Ido not believe this circumstance shows that the maxilla is formed
by a union of the lateral mandibles.
A superior and inferior maxille are only found in the Cephalo-
poda, and perhaps in the larval state of some Pectinibranchiata ;
but it is not placed beyond doubt that the beak of the Cephalopod
is really homologous with the maxilla in the Pulmonata. The
maxilla of Succinea is not unlike the upper maxilla in the Ce-
phalopoda. Prof. Van Beneden’s figure of the “ machoire supé-
rieure et inférieure ” of Amphipeplea glutinosa (Kixercices Zooto-
miques, f. 6-9) looks very like the beak of a Cephalopod; but
it may be erroneous, because the radula is represented (f. 7) as
forming a part of the lower maxilla. A similar mistake was
committed by Moquin-Tandon, who represented in Neritina as
superior and inferior maxillee what, according to Claparéde, are
only the edges of a corneous membrane lining the oral cavity.
Messrs. Alder and Hancock have shown, in Acanthodoris
pilosa, a solitary inferior maxilla represented by a flat plate with
a split in front protruding from the mouth; but this part seems
to me more likely to belong to the lingual cartilages.
Perhaps the “anterior or lower lip, armed with teeth,” repre-
sented by Macdonald+} in Clio caudata is the edge of an inferior
maxilla.
The mandibule are two lateral concave plates, not unlike a
bivalve shell}, the free edge of which (masticatory edge § of
Bergh) is either tuberculated or denticulated. From the position
of the muscles and from the form of the pectinated edge in
Phyllodesmium, Ehrb. (Bergh, ‘Anatomie’), it appears that the
mandibles cannot be used for mastication, but are only adapted
for use as a pair of forceps to hold the food during the triturating
action of the radula. By the umbones of the mandibles (umbi-
licus, Midd., Bergh) the two halves are united with muscular
ligaments. This kind of lateral mandibles is found in the
Pleurognatha Gymnobranchiata (dolidie, Diphyllidie, &c). In
* Heynemann, Malacozoologische Blatter, 1861, p. 143. The same is
the ease with Vitrina in its young state.
t Trans. Royal Soc. Edinb. vol. xxiii. pt. 2. p. 188, t. 9. f. 3.
+ Described as such by Brown in Binghami radoi
{ Described as such by Brown in Binghamia paradowa.
§ Cutting edge (Alder and Hancock).
the Buccal Parts of the Mollusca. 75
Proctonotus mucroniferus Alder and Hancock have discovered a
“ corneous transverse arch or strap” uniting the two halves, and
reminding us of the middle maxilla in the Pulmonata. The
nature of the mandibles described by Troschel (Gebiss der
Schnecken, t. iii. f. 18) in Pnrewmodermon is, 11 my opinion, very
doubtful.
The cheek-plates*, or immoveable mandibles, are two lateral
plates, without cutting edges, composed of scaly or acicular +
particles, which seem only of use to protect the inside of the
mouth from injury by the spinous tongue. Sometimes the two
plates are united above by a ligament. This kind of mandible is
found in nearly all the Tnioglossata, as well in those provided
with a rostrum (Cyclostomus, Valvata, Rissoa, Jeffreysia, Crepi-
dula, Vermetus, Trichotropis, &c.) as in those with an haustellum
(Marsenia, Natica, Cyprea, Cassis, Triton, Strombus, &c.) : vide
Troschel, Gebiss derSchnecken. These plates are apparently want-
ing in all the Rhachiglossata (Murex, Fusus, Nassa, &c.). The
linear horny plates described in Buccinum undatum by Cuvier
(Anat. des Mollusq. figs. 11, 12) and by Valenciennes (Archives
du Muséum, t.v.p. —, t. 25. f. 6) are probably appendages of
the tongue, and used as a handle in perforating the shells on
which they prey.
The prehensile collar (Hancock) is a circular band composed
of conical, often forked, erectile spies. Sometimes it is entire,
as in Ancula cristata and Goniodoris nodosa; in others it is
divided into two lateral parts, not unlike cheek-shields, as in
Idaha pulchella and Acanthodoris pilosa.
Evertile cheek-cushions (“lames latérales,” Lacaze-Duthiers t),
two large evertile lateral sacs, covered with close-set lanceolate
plates, like pavement, in Pleurobranchus. They seem to be an
intermediate form between the prehensile collar and the evertile
arms of the Gymnosomata and Cephalopoda.
Cheek-hooks or cheek-crooks (harpage $); mandibles, Eschricht,
in Clione; “les csxecums de la bouche, avec un tube corné dans
Pintérieur ” (Van Beneden ||) ; “evertile tubes serving as pre-
hensile organs” (Troschel4]) (this author has not detected these
organs in Clione and Pneumodermon); “ lateral or cheek-pouches”
(Macdonald**, LEschricht++ first showed that these tentacle-
* *Mandible-like parts, indumentum epitheliz, the horn sheaths of
the inner lip” (Bergh).
+ In Thecosomata (Hyalea, &c.) they are composed of four or five
band-lhke plates (vide 'Troschel, Gebiss).
f Ann. des Se. nat. sér. 4. vol. xi. t. 7. f. 4.
§ [intentionally use this form of the word,
|| Exercices zootomiques. { Gebiss d. Schn.
** Loc. cit. and Linn. Trans. xxii. 1858, p. 248.
++ Anatomiske Undersogelser over Clione borealis, 1838.
6%
Ac Te Dr. Morch on the Homology of
like organs, which are always provided with corneous hooks,
are evertile; but this has been established by Eydoux and
Souleyet* in the genera Pneumodermon and Cline.
The long arms of the decapod Cephalopods seem to me with-
out doubt the same organs as the cheek-hooks of Gymnosomata ;
they have the same latero-ventral position, and are retractile
within a cavity or pouch inside the short arms, which may be
considered as a single fissured veil, and perform the same func-
tion in the animal. Prof. Lovén considered the long arms of
Decapoda homologous with the tentacula (vibracula) of the Gas-
teropoda, and compared them specially with the rhinophores of
Doris. I do not believe that this homology is natural, because
the tentacula are always dorsal, and the “ long arms ” of Cephalo-
poda always /atero-ventral. It seems also very doubtful whether
an organ of prehension can be homologous with an organ of
smell. If the Cephalopoda have no tentacula at all, the “ crétes
auriculaires ” (D’Orb.) or “ paupiéres inférieures” (D’Orb.) of
the Myopsid Decapods may be considered as their homologue.
The triangular filaments over the eyes of Octopi are perhaps, too,
a form of tentacula. I believe they are provided with a nervous
ganglion at their base, like those of Doris.
The “organe en pioche,” in Conus, of Quoy and Gaimard is
a tubular sac, provided with arrow-like corneous hooks with a
hollow channel, which are considered venomous. I have had an
opportunity of confirming the exactness of the anatomy repre-
sented in the ‘ Voyage de l’Astrolabe.’ As these hooks are not
situated in the true oral tube, it is still very doubtful whether
they really can be considered to belong to the radula. I
suppose these arrow-like hooks are more allied to the cheek-
hooks of Gymnosomata. As the hooks turn their points in
opposite directions in the two arms of the pouch, it is probable
they are evertile each in a different direction, but how and through
what aperture is not yet discovered.
On the Palpi and Lips.
There are often in the Gasteropoda two different apertures in
the same animal, called mouths. The true mouth is the anterior
opening of the bulbus pharyngeus, frequently prolonged into an
haustellum. This opening is only provided with the hard plates
described above. The outer or false mouth is a simple slit in
the skin, containing the lips or palpi, but never any mandibles;
and it forms a passage for the protrusile haustellum, which, in
a retracted state, is concealed in a cavity behind this slit.
In the Acephala the palpi are represented by two pairs of
* Voyage de la Bonite; Mrs. Gray’s figures of Moll. t. 255. f. 3.
the Buccal Parts of the Mollusca, 77
foliaceous expansions, grooved (sulcated) inside, and destined to
conduct the particles of food to the mouth, and they may thus
be considered passive prehensile organs.
In the Calyptreeide the same organ is composed of a single
pair, which is represented in all drawings of these mollusks as
two intertentacular tubercles. In reality these tubercles are
flat inside, grooved as in the Acephala, and probably used as a
pair of forceps, as appears from their relative position in different
specimens. In Capulus the palpi form a long haustellum-like
tube, with a narrow slit on the upper side. In Dentalium and
Siphonodentalium they form a closed, flat tube.
In the Doride the palpi are situated near the outer oral aper-
ture, and are sometimes of a linear form, with a longitudinal
groove on the middle, sometimes foliaceous (Heaxabranchus) or
meeting together in a semicircle (Lamellidoris), or they become
the oral veil.
In Conus, Terebra, and perhaps Pleurotoma this veil is de-
veloped into a large infundibuliform sucking-cup, which dis-
appears when the haustellum is protruded. This veil was for a
long time regarded as the proboscis, until Dr. Gray* showed
its real nature. In Conus tulipa and C. striatus the edge is
divided into many digitations. (Voyage de l’Astrolabe; Mrs.
Gray’s figures, 1. t. 10. f.6, and 12. f. 2.)
In the gymnosome Pteropoda the oral veil is divided into
several conical ‘‘arms,” provided with numerous suckers, and
probably corresponding to the grooves in the labial palpi of
Acephala and Calyptreeide.
In the Cephalopoda these “arms” are still more developed,
and united by a membrane, which sometimes extends to the tips
(Cirroteuthis, Eschricht).
Under the name “ velum,” as employed by Lovén, very dif-
ferent organs are confounded. I believe three kinds may be
distinguished :—
1. The oral veil, formed by the palpi or lips, an organ
for prehension or locomotion (Cephalopoda, larva of Opistho-
branchia, e. g. Doris), or even partly available as a male organ
(hectocotyl of the Cephalopoda), as in the Spiders. The ‘‘ mem-
brane orale”’ of D’Orbigny, found in the decapod Cephalopoda,
seems to me a kind of suspensorium, lke the “ brids” of the
funnel.
2. The tentacular veil, situated further from the mouth. It is
placed, in Pleurobranchus, above the mouth, and formed by the
union of the anterior tentacula (vibracula in Aplysia). In
b
* “On the Head of the genus Conus,” Ann. Nat. Hist. Aug. 1853, p. 176.
78 Dr. Moreh on the Homology of
Clione its two halves can cover the arms entirely, like a hood
(see Eschricht’s ‘ Anatomie’), and resemble the “ crétes auricu-
laires ” and “ paupiéres inférieures ” of D’Orbigny, above men-
tioned.
3. The post-tentacular veil is only found in the larval state of
Rissoa, Lovén (Ofversigt, 1847), and in the genera Chiropteron
of Sars* and Macgillivrayia of Macdonald.
In analogy with the names of the foot given by Prof. Huxley,
these three kinds of velum may be called Prohistion, Mesohistion,
-and Metahistion.
That the short arms of Cephalopoda cannot be considered
the homologue of the foot, is evident from the circumstance that
the same kind of arms is found in the Pteropoda Gymnosomata
simultaneously with an undoubted foot. Prof. Lovén first
showed that the funnel must be the foot of Gasteropoda, and he
suggests that the interior valvula of most Decapods and of
Nautilus} corresponds to the solea pedis of Gasteropoda—a
proposition which, I believe, is correct. The funnel would thus
correspond to the epipodium in the Gymnosomata; its dorsal
wings, attached to the neck of nearly all Decapod Cephalopoda,
are provided with three cartilages which may be compared to
a tripartite operculum}, thus proving that the lobus operculi-
gerus of Lovén§ is a part of the epipodial line (Huxley), and not
of the true foot (solea).
Pedipes afra, as represented by Adanson (Hist. Nat. du Séné-
gal, tab. 1) and by Lowe (Zool. Journ. vol. v. pl. 18. f. 8, 9), is
the only example of a foot divided into the propodium, meso-
podium, and metapodium of Prof. Huxley.
EXPLANATION OF PLATE VI.
Fig. 1. Pneumodermon: intestinal channel, with the labial suckers, sali-
vary glands, stomach, rectum, and anus. The cheek-pouches
are united at their ends by a muscular band.
Fig.1b. The cesophagus opened, to show the entrance of the cheek-pouches,
with the prehensile hooks in the centre, on both sides of the
radula.
* Chiropteron semilunare, Sars (Beskrivelser og Jagttagelser 1835, t. 14.
f. 38), is probably the larva of Aporrhais.
+ R. Owen, ‘On the Pearly Nautilus,’ t. 3. f. 2 e.
{~ The middle cartilage is articulated to the inside of the front of the
shield or dorsal plate, and the two lateral cartilages to similarly excavated
plates in the inner margins of the mantle.
§ The dorsal part of the metapodium (Huxley, “‘ On the Morphology
of Cephalous Mollusca,”’ Phil. Trans. 1853, p. 29).
the Buccal Parts of the Mollusca. 79
Fig. 1c. Pneumodermon, with everted prehensile hooks (harpagz). (Figs.1,
14, and 1c from ‘ Voyage de la Bonite.’)
Fig. 1d. One of the sacs opened, to show the evertile hooks, with some
detached corneous spines. (From Van Beneden’s ‘ Exercices
zootomiques.’ )
Fig.2. Clione limacina, Phips.: the anterior part of the cesophagus.
The harpaga of the right cheek-pouch is everted; the left re-
tracted, and seen through an opening.
Fig. 2a. The harpaga, with from 24 to 32 corneous hooks. From Esch-
richt’s ‘ Undersdgelsé om Clione borealis.’
Fig. 3. Onychoteuthis, from the ventral side, with protruded arms. It is
doubtful whether the metacarpal suckers are really destined
to act against each other, as represented by Férussac and D’Or-
bigny.
Fig. 4. Conus tulipa: the oral veil expanded at the edge, and opened to
show the haustellum.
Fig. 4a. The oral veil in a contracted state.
Fig. 46. The intestinal channel and its glands, the two liver-lobes, and an
intermediate small so-called “ gland,” perhaps the true stomach.
Over the nervous ring is the ‘“ organe en pioche,” attached to
the cesophagus by muscular bands; but I have not been able, in
a specimen of C. consors, to discover any communication with the
interior of the cesophagus. It cannot, therefore, represent the
papillz of the tongue, which in Littorina and Patella are of enor-
mous length. At the insertion of the ‘ organe en pioche,” close
to the base, is the long vermiform canal ‘of the single ‘‘ glande
salivaire cucumiforme.” The walls of this cylindrical organ are
enormously thick and muscular; and it therefore seems to me
not to be a salivary gland, but a suctorial stomach, like that of
most haustellate imsects. A similar unequal sac, in the same
position, is found in the following Mollusca, all of which are
provided with an haustellum :—
Murex cichoreus, Voyage de l’Astrolabe, t. 36. f. 1, et Suppl.
t. 2. f. 1 d, troisieme glande salivaire.
Dolium olearium, ibid. t. 41. f. 4.
Voluta fusus, Q.& G. ibid. t.44.f.9h, “diverticulum ou cecum
cesophagien trés-considérable ;” and f. 10 and 11.
Ancillaria albisuleata, ibid. t. 49. f. 11 d, “diverticulum,
espéce de czecum.”’
The petiolate cheek-pouch im Lamellidoris, described by Mr.
Hancock, is perhaps, too, a kind of instrument destined to produce
a vacuum, like the suckers of Cephalopoda.
Fig. 4 c. The “ organe en pioche”’ opened, to show the arrows turning the
points in a different direction m each branch.
Fig.4d. Different forms of the arrows, hollow inside.
(Figs. 4-4 d from ‘ Voyage de l’Astrolabe,’ t. 44.)
Fig. 5. Pharynx of Pleurobranchus, showing the “lames latérales” ca
both sides of the tongue. a. The hard parts of the “lames
latérales,”’ strongly magnified. (From Lacaze-Duthier’s ‘ Anato-
mie du Pleurobranche orange,’ J. c.)
80 Mr. J. Blackwali on recently discovered Spiders
1X.—Descriptions of recently discovered Spiders collected in the
Cape de Verde Islands by John Gray, Esq. By Joun Brack-
WALL, F.L.S.
Tribe Octonoculina.
Family Lycosip#.
Genus Lycosa, Latr.
Lycosa helva.
Length of the female ths of an inch; length of the cephalo-
thorax 13, breadth --; breadth of the abdomen 1; length of
a posterior leg +; length of a leg of the third pair =7.
The eyes, which are unequal in size, are disposed in front and
on the sides of the anterior part of the cephalothorax ; four,
much smaller than the rest, form a transverse row immediately
above the frontal margin, the two lateral ones being rather
smaller than the intermediate ones of the same row; the other
four describe a trapezoid, the two anterior eyes, which are the
largest of the eight, forming its shortest side. The cephalo-
thorax is long, convex, clothed with short adpressed hairs, com-
pressed before, rounded in front and on the sides, which are
marked with furrows converging towards a narrow, dark-brown
indentation in the medial line of the posterior region ; it is of a
dull yellow colour, with a broad brown band, mingled with yel-
low, extending along each side, a short brown line directed
backwards from each eye of the posterior pair, and a small dark
brown streak on each angle of the frontal margin. The falces
are powerful, conical, vertical, convex in front, and armed with
teeth on the inner surface; and the lip is somewhat quadrate,
being rather broader at the base than at the apex. These organs
have a very dark-brown hue, the lip being the paler. The
maxille are straight and enlarged and rounded at the extremity;
the sternum is oval, glossy, and thinly clothed with whitish
hairs. The legs are robust, provided with hairs and sessile spines,
and the metatarsi and tarsi have hair-like papille distributed on
their inferior surface; the fourth pair is the longest, then the
first, and the third pair is the shortest ; each tarsus is terminated
by three claws; the two superior ones are curved and pectinated,
and the inferior one is merely rudimentary; the palpi have a
curved, pectinated claw at their extremity. These parts are of a
dull yellow colour. The abdomen is oviform, convex above, and
projects over the base of the cephalothorax; it is clothed with
short hairs, and is of a pale yellow-red colour; at the anterior
extremity of the upper part there is a brown angular mark,
having its vertex directed downwards; and an obscure band of
the same hue, which has a projecting point on each side, near
collected in the Cape de Verde Islands. 81
its middle, and whose posterior extremity is bifid, extends from
within the anterior angle along the middle; between the termi-
nation of this band and the spinners there are several faint,
brown, angular lines, whose vertices are directed forwards, and
numerous minute spots of a similar colour occur on the sides,
the extremity of the spinners being of a darker brown; the
sexual organs are minute, and have a red-brown hue.
Captured in the Island of St. Antonio.
Genus Hersriia, Savigny.
Hersilia versicolor.
Lengtb of the female (not including the spimners) -=%;ths of
an inch; length of the cephalothorax +z, breadth ~,; breadth
of the abdomen +5; length of a leg of the second pair +7; ;
length of a leg of the third pair J.
The cephalothorax is short, broad, slightly compressed and
elevated before, rounded in front and on the sides, moderately
convex, and has an indentation in the medial line ; it is of a
dull brownish-yellow colour, with a brownish-black angular
mark in the middle of its posterior half, whose vertex is directed
forwards, and some large spots of the same hue on the lateral
margins. The falces are conical, somewhat inclined towards the
sternum, provided with long hairs, and are of a dark-brown co-
lour, the extremity having a red-brown hue. The maxillze are
short, powerful, strongly inclined towards the lip, and obliquely
truncated at the extremity, which is pointed on the inner side ;
the lip is somewhat quadrate, being broader at the base than at
the extremity; the sternum is broad and reniform. These parts
have a dull yellowish-white hue. The legs are long, provided
with hairs, and of a dark-brown hue, with narrow whitish an-
nuli; the second pair is longer than the first, and the third pair
is the shortest ; the fourth pair was mutilated, but the femora
were entire, and exactly corresponded in length with those of the
second pair; the slender metatarsi and tarsi have each only one
joint, the latter being terminated by three claws ; the two supe-
rior ones are curved and pectinated, and the inferior one is in-
flected near its base. The palpi, which are long and resemble
the legs in colour, have a curved, pectinated claw at their extre-
mity. The eyes are disposed on the anterior part of the cephalo-
thorax in two transverse curved rows, having their convexity
directed forwards; they are situated high above the frontal
margin, the lateral eyes of the anterior row being very much
the smallest, and the two intermediate ones of the same row,
which are rather wider apart than those of the posterior row,
much the largest of the eight. The abdomen is rather broader
82 Mr. J. Blackwall on recently discovered Spiders
in the middle than at the extremities, convex above, and projects
over the base of the cephalothorax ; on the upper part and sides
it is densely clothed with coarse, adpressed, reddish, yellowish,
and whitish hairs intermixed, and has black and white bristles,
more or less erect, distributed over the surface of those parts; a
brownish-black band extends from the anterior extremity of the
upper part to its middle; this band, which is enlarged about its
middle, is bifid at its extremity, and from each diverging branch
a row of spots of the same hue passes to the spinners ; the sides
are marked with oblique brownish-black streaks, and the under
part, which is downy, has a very pale dull yellowish hue; the
sexual organs are highly developed, prominent, and of a red-
brown colour; the superior spinners are long, triarticulate, and
have the spinning-tubes disposed on the inferior surface of the
elongated terminal joint, which tapers to a point; this joint has
a brownish-black hue on the upper and exterior surfaces of its
base, the colour of the other parts of these spinners and of the
intermediate and inferior pairs being pale dull yellow.
Captured in the Island of St. Jago.
Family Saiticrp”.
Genus Saurticus, Latr.
Salticus simplex.
Length of the male ;3,ths of an inch; length of the cephalo-
thorax 5!;, breadth +; breadth of the abdomen +,; length of
a posterior leg 3; ; length of a leg of the second pair 4.
The legs are robust, especially those of the anterior pair, and
are provided with hairs and a few spines; the fourth pair is
rather the longest, then the third, and the second pair is the
shortest ; they are of a reddish-brown colour, the femora being
the darkest, and the tarsi much the palest ; each tarsus is termi-
nated by two curved claws, below which there is a small scopula.
The palpi are short, and have a dark-brown hue; the radial
jot projects a pointed apophysis from its extremity, on the
outer side ; the digital jot is oval, convex, and hairy externally,
compact at the extremity, concave underneath, at the base,
comprising the palpal organs, which are well developed, not
very complex in structure, and of a red-brown colour. The
minute intermediate eye of each lateral row is nearer to the an-
terior than to the posterior eye of the same row. The cephalo-
thorax is somewhat quadrilateral, sloping abruptly at the base,
and very gradually to the front, which projects a little beyond
the base of the falces ; it is of a dark-brown colour, and is clothed
with reddish-brown hairs; a broad patch of yellowish-white or
pale yellow hairs is situated below the lateral eyes; a spot im-
collected in the Cape de Verde Islands, 83
mediately behind each posterior eye, and a short streak on the
upper part of the posterior slope, are composed of yellowish-
white hairs; and a narrow longitudinal band of white hairs
occurs above each lateral margin. The falces are conical and
vertical ; the maxille are straight, and enlarged and rounded at
the extremity; and the lip and sternum are oval. These parts
are of a dark-brown colour; the sternum, which is the darkest,
is supplied with some long white hairs, and the extremity of
the falces, maxillee, and lip is tinged with red. The abdomen is
oviform, pointed at the spmners (which are prominent), convex
above, and projects a little over the base of the cephalothorax ;
it is of a very dark-brown colour, and is well clothed with hairs,
those in the medial line of the upper part forming a broad, dull
yellowish-brown band that tapers to the spmners ; a spot in the
middle of the anterior extremity, and three spots disposed in a
row on the upper part of each side, of which the anterior one is
the largest, are composed of white hairs; and the under part is
abundantly supplied with hoary hairs.
Captured in the Island of St. Nicholas.
Salticus lepidus.
Length of the male =3,ths of an inch; length of the cephalo-
thorax +, breadth ;4;; breadth of the abdomen +;; length of a
posterior leg +; length of a leg of the second pair 2.
The minute intermediate eye of each lateral row is nearly
equidistant from those constituting its extremities. The cephalo-
thorax is large, glossy, somewhat quadrilateral, sloping abruptly
at the base, and very gradually to the front, which projects a
little beyond the base of the falces; it is of a brownish-black
colour, the cephalic region and narrow lateral margins being the
darkest ; the middle of the posterior half is tinged with red,
and there is a spot composed of white hairs nearly intermediate
between the posterior pair of eyes; the front and a broad longi-
tudinal band extending above each lateral margin are densely
clothed with white hairs. The falees are short, conical, and
vertical; the maxilla are straight, and enlarged and rounded at
the extremity, and the lip is oval. These parts are of a dark-
brown colour, the extremity of the maxillz and the apex of the
lip having a dull-yellow hue ; the sternum is oval and of a pale
dull-yellowish colour. The legs are robust and provided with
hairs and a few sessile spines; the fourth pair is the longest,
then the first, and the second pair is the shortest ; they are of a
dull-yellow colour, with a dark-brown spot at the extremity of
the joints, the tarsi having a slight tinge of red; the anterior
pair, which are the darkest, have a broad brownish-black band
extending along the anterior surface of the femora, genua, and
84. Mr. J. Blackwall on recently discovered Spiders
tibie, and the metatarsi are of the same hue; each tarsus is
terminated by two curved claws, below which there is a small
scopula. The palpi are short, and of a pale yellow colour, the
axillary joint, the base of the humeral joint, and the outer sur-
face of the radial jot having a brown-black hue; the radial
joint projects a curved, finely pointed, black apophysis from its
extremity on the outer side, which is directed obliquely forward
and outward; the digital joint is oval, convex, and hairy exter-
nally, concave within, comprising the palpal organs, which are
moderately developed, with a black filiform spme curved from
the outer side round the base and inner side, and have a pale
brownish-red hue. The abdomen is oviform, pointed at the
spinners (which are prominent), convex above, and projects a
little over the base of the cephalothorax ; it is densely covered
with adpressed hairs, and has a broad yellow-brown band in the
medial line, which tapers to the spinners; on each side of this
band there is a longitudinal band composed of white hairs, which
overlie and almost conceal a black band whose outline is some-
what irregular near its posterior extremity, on the inner side; a
narrower black band passes along the lower part of each side,
and the space between these black bands has a yellow-brown
hue; the under part is of a pale dull-yellowish colour, with a
large oval soot-coloured spot in the middle, and the spinners
have a dark-brown hue.
Captured in the Island of St. Nicholas.
Salticus sedulus.
Length of an immature male }th an inch; length of the ce-
phalothorax +1;, breadth +1, ; breadth of the abdomen ';; length
of a posterior leg .;; length of a leg of the second pair 5.
The cephalothorax is glossy and somewhat quadrilateral
: | SLORY: as
sloping abruptly at the base, and gradually to the front, which
projects a little beyond the base of the falces ; it is of a yellowish-
brown colour, the cephalic region, the narrow lateral margins,
and a fine parallel line situated immediately above them having
a black hue; there is a broad, longitudinal, brown band on each
side of the medial line, and the frontal margin is densely clothed
with long white hairs, which extend to the sides, The falces
are short, conical, and vertical; the maxille are straight, and
enlarged and rounded at the extremity; and the lip and ster-
num are oval. These parts have a yellowish-brown hue, the
falces, maxillee, and lip being the darkest at the base; and the
, lie I S )
sternum, which is the palest, has brownish-black lateral margins.
The legs are robust, provided with hairs and sessile spines, two
5 ae ] : : E y
parallel rows of the latter occurring on the inferior surface of
the tibize and metatarsi of the first and second pairs ; the fourth
collected in the Cape de Verde Islands. 85
pair is the longest, then the third, and the second pair is the
shortest ; they are of a brownish-yellow hue, with some brownish-
black spots and streaks, a longish one of the latter occurring on
the anterior surface of the femora, with the exception of those
of the fourth pair; each tarsus is terminated by two curved
claws, below which there is a small scopula. The palpi are
short, and paler than the legs ; the digital jomts of the specimen
described were very tumid, indicating that it had to undergo its
final ecdysis before it arrived at maturity. The minute inter-
mediate eye of each lateral row is rather nearer to the posterior
than to the anterior eye of the same row. The abdomen is ovi-
form, pointed at the spimners (which are prominent), convex
above, and projects a little over the base of the cephalothorax ;
it is of a yellowish-white colour, with a broad, dark-brown
band, somewhat irregular in outline, extending from the anterior
extremity of the upper part along each side of the medial line
to the spinners; and, parallel to these bands, a narrower one of
the same hue passes along each side ; the spinners are of a dark-
brown colour, and an obscure soot-coloured spot occurs at their
base, on the under part. .
Captured in the Island of St. Nicholas.
Family THomisip&.
Genus Tuomisus, Walck.
Thomisus piger.
Length of an immature female ~,ths of an inch; length of
the cephalothorax ;1;, breadth ;4;; breadth of the abdomen +; ;
length of a leg of the second pair +; length of a leg of the third
pair }.
The eyes are disposed on the anterior part of the cephalo-
thorax in two transverse curved rows, forming a crescent whose
convexity is directed forwards; the eyes of each lateral pair,
which are larger than the intermediate ones, are seated on a
conspicuous tubercle, the anterior ones being the largest of the
eight. The cephalothorax is convex, slightly compressed before,
truncated in front, with a few bristles directed forwards from its
anterior margin, rounded on the sides, and abruptly depressed
at the base; the falces are subconical, vertical, and provided
with a few bristles near the base in front; the maxille are
obliquely truncated at the extremity, on the outer side, and in-
clined towards the lip, which is triangular; the sternum is
heart-shaped ; the legs are very unequal in length, and are pro-
vided with hairs and spines, two parallel rows of the latter oc-
curring on the inferior surface of the tibize and metatarsi of the
second pair, which is much longer and more robust than the
86 Mr. J. Blackwall on recently discovered Spiders
third and fourth pairs, the third pair being the shortest; the
legs of the first pair were missing, but, judging from the coxe,
they probably did not differ materially 1 in dimensions from those
of the first pair; each tarsus is terminated by two curved claws
having one or two minute teeth at their base; the palpi are
short, and have a small curved claw at their extremity. These
parts are of a pale dull-yellow colour; numerous very minute,
slightly raised, dark- brown spots occur on the cephalothorax ; a
pale brown band passes from each lateral pair of eyes towards
its base, and the tubercles on which those eyes are seated have
a soiled white hue. The abdomen is sparingly clothed with
short hairs, convex above, broader towards the posterior than at
the anterior extremity, which has the ¢ appearance of having been
cut in a direct line across, and projects over the base of the
cephalothorax ; it is of a pale dull yellowish colour, obscurely
freckled with dull white; on each side of the upper part there
is a curved series of small soot-coloured spots, which are most
conspicuous in the posterior region; the two series converge
towards their extremities, and deseribe a large oblong oval ;
soot-coloured band passes along the .upper part of each oH
nearly to the spinners, its posterior extremity bemg broken into
irregular spots.
Captured in the Island of St. Antonio.
Family Drassip&.
Genus Drassus, Walck.
Drassus n igromacu latus.
Length of the female (not including the ene)
inch ; length of the cephalothorax =3;, breadth 3; ; ‘breadth of
the abdomen + E length of a posterior leg +; length of a leg of
the third pair =.
The cephalothorax is compressed before, truncated in front,
rounded on the sides, which are somewhat depressed, abruptly
sloped at the base, and has a slight narrow indentation in the
medial line of the posterior region ; it is of a pale reddish-brown
colour, is clothed with yellowish-grey hairs, which are densest
on the sides, and has some long black ones, more or less erect,
distributed over its surface ; three black spots form a row on
each side, the anterior one being the largest and most irregular
in form ; it has a large triangular brown-black mark at its base,
and the narrow later al mar eins are of the same hue. The eyes
are disposed on the anterior part of the cephalothorax in two
transverse, slightly curved rows; the convexity of the anterior
row is directed upwards, and the two intermediate eyes are the
collected in the Cape de Verde Islands. 87
largest and darkest of the eight ; the convexity of the posterior
row is directed forwards, and the two intermediate eyes are
rather nearer to each other than they are to the lateral eyes of
the same row; the lateral eyes of both rows are separated by a
wide interval. The falces are powerful, conical, convex at the
base, and vertical ; the maxille are convex at the base, hollowed
on the inner side, and strongly curved towards the lip, which is
longer than broad and rounded at the apex. These parts are of
a dark-brown colour tinged with red, the falces being the darkest,
and the extremity of the maxille having a yellowish-white hue.
The sternum is short, oval, hairy, and of a yellowish-brown
colour. The legs are robust, provided with hairs and spines,
the latter bemg most numerous on the tibie and metatarsi of
the third and fourth pairs; they are of a yellow-brown colour,
marked with a few obscure soot-coloured spots, the metatarsi
and tarsi, which are the darkest, being tinged with red; the
fourth pair is the longest, then the first, and the third pair is
the shortest; the tarsi are supplied with hair-like papille on
their inferior surface, and are terminated by two curved, pecti-
nated claws. The palpi are short, and resemble the legs in co-
lour; and the digital joint, which is provided with strong spines,
has a curved, pectinated claw at its extremity. The abdomen is
oviform, densely clothed with hairs, convex above, and projects
a little over the base of the cephalothorax ; it is of a dull yellowish-
grey colour, with a broad band of black hairs curved round its
anterior extremity, and extended along each side in an irregular
line more or less interrupted; from the middle of the curved
band an obscure, longitudinal, brown band extends, which tapers
to its posterior extremity; and on each side of it there is a
series of irregular black spots that become confluent as they
approach the spinners, which are cylindrical, prominent, and of
a pale red-brown colour, with a black line on the sides and under
part of their base; the sexual organs are moderately developed,
and have a red-brown hue, that of the branchial opercula being
yellowish brown.
The male is smaller than the female, but it resembles her in
colour. ‘The radial joint of its palpi is rather smaller than the
cubital joint, and projects a straight, pointed, dark-brown apo-
physis from its extremity on the outer side; the digital joint is
oval, convex, and hairy externally, concave within, comprising °
the palpal organs, which are moderately developed, rather pro-
minent, with a slender, curved spine on the outer side, and two
processes at their extremity, the inner one of which is much
the smaller; their colour is red-brown.
One male and four females of this species were taken in the
Island of St. Jago, and three females in the Island of Fogo. It
88 Mr. J. Blackwall on recently discovered Spiders
belongs to Walckenaer’s family Lithophile and first race Luci-
fuge of the genus Drassus.
Drassus assimilatus.
Length of the male (not including the spinners) +}ths of an
inch; length of the cephalothorax ,';, breadth +; breadth of
the abdomen +; length of an anterior leg 3; length of a leg of
the third pair +.
The legs are long, provided with hairs and sessile spines, and
of a dull yellowish-white colour, the metatarsi and tarsi, which
are strongly tinged with brown, having numerous hair-like pa-
pill on their inferior surface ; the first pair is the longest, then
the fourth, and the third pair is the shortest; each tarsus is
terminated by two curved pectinated claws. The palpi resemble
the legs in colour, and the radial, which is much longer than
the cubital joint, has no apophysis at its extremity; the digital
joint is of a narrow oblong-oval form; it is convex and hairy
externally, compact and somewhat pointed at the extremity, and
has a shallow concavity near its base, on the under side, com-
prising the palpal organs, which are small, little complicated in
structure, with a fine, curved, black spine towards the inner
side, and a shorter one at their extremity ; these organs have a
pale red-brown hue. The eyes, which are seated on black spots,
are disposed on the anterior part of the cephalothorax in two
transverse, slightly curved, parallel rows; the posterior row is
the longer, and the two intermediate eyes, which are somewhat
oval, and nearer to each other than they are to the lateral eyes
of the same row, describe with the intermediate eyes of the an-
terior row, which is situated immediately above the frontal mar-
gin, an oblong-quadrangular figure; each lateral eye of the
posterior row is seated on a minute tubercle, and the inter-
mediate eyes of the anterior row are the largest and darkest-
coloured of the eight. The cephalothorax is large, compressed
before, truncated in front, rounded on the sides, thinly clothed
with hairs, convex, glossy, with a narrow indentation in the
medial line of the posterior region; it is of a yellow-brown co-
lour, the anterior part, which is the darkest, being faintly tinged
with red, and the narrow lateral margins have a brown hue.
The falees are subconical, rather prominent, and armed with a
few teeth on the inner surface; the maxillee are convex at the
base, enlarged at the extremity, which is obliquely truncated on
the inner side, and slightly curved towards the lip, which is
long, and truncated and hollowed at the apex. These parts are
of a dark-brown colour, the maxille and lip, which are the palest,
being tinged with yellow at the extremity. The sternum is
oval, glossy, of a pale dull yellowish colour, and is supplied with
collected in the Cape de Verde Islands. 89
long hairs, which are densest on the narrow dark-brown lateral
margins. The abdomen has an oblong subcylindrical figure,
tapering a little to the spinners; it is slightly convex above,
projects but little over the base of the cephalothorax, and is
clothed with hairs; the upper part is of a brown colour, obscurely
intermixed with yellowish white, and has some long black hairs
at its anterior extremity ; the under part and the spinners, which
are cylindrical and prominent, have a yellowish-white hue, the
latter bemg tinged with brown.
This spider, which was captured in the Island of St. Antonio,
is very closely allied to Drassus lapidicolens, but differs from it
not only in colour, but also in the relative length of its legs and
in the structure of its palpi and falces.
Family Crn1rLonIp&.
Genus Orituyra, Blackw.
Orithyia luteola.
Length of the female 1th of an inch; length of the cephalo-
thorax ;!;, breadth ~;; breadth of the abdomen ;1;; length of
an anterior leg =3,; length of a leg of the third pair ~;.
The eyes, which are unequal in size and seated on brown
spots, are disposed on the anterior part of the cephalothorax in
two transverse curved rows, whose convexity is directed forwards;
the anterior row, which is the less curved, is situated immediately
above the frontal margin, and the two intermediate eyes are
seated on a protuberance; the lateral eyes of both rows are
placed on minute tubercles, and are wide apart, those of the
anterior row being the smallest of the eight. The cephalothorax
is short, broad, convex, somewhat oval, with two furrows on
each side converging towards a shallow indentation in the medial
line of the posterior region ; it is clothed with coarse, pale-yel-
lowish hairs, and is of a yellowish-brown colour; a dark-brown
band passes from the eyes to its base, and a broad one of the
same hue extends along each side. The falces are short, strong,
subconical, and vertical; the maxillz are nearly straight, power-
ful, and greatly enlarged at the extremity, which is obliquely
truncated and protuberant on the inner surface; the lip is tri-
angular; and the sternum is oblong heart-shaped, hairy, and
has eminences on the sides, opposite to the legs. These parts
have a dull brownish-yellow hue. The legs are very unequal in
length, the first pair being much the longest and most robust ;
the fourth pair surpasses the second, and the third pair is the
shortest ; they are provided with hairs, and the metatarsus of
each posterior leg has a calamistrum situated in a curve at its
superior surface; the femora, genua, and tibie of the anterior
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. 7
90 Mr. J. Blackwall on recently discovered Spiders
legs are of a brownish-black hue, the inferior surface being
much the palest; a pale yellowish annulus occurs on the first,
and is situated nearer to its extremity than to its base, which is
the darkest-coloured ; the metatarsi and tarsi are disproportion-
ally slender, and have a brownish-yellow hue ; the second, third,
and fourth pairs of legs are of a pale brownish-yellow colour,
marked with brownish-black annuli. The palpi are short, hairy,
and of a pale brownish-yellow colour, the extremity of the hu-
meral joint, the cubital joint, and the base of the radial joint
having a brown-black hue. The abdomen is oviform, pointed at
the spinners, convex above, with a conical protuberance on each
side of the upper part, near its anterior extremity, and projects
over the base of the cephalothorax; the upper part is of a pale
yellowish colour, a fine black line, which is rather the broadest
near its anterior extremity, extending along the middle; a space
above the spinners, the sides, and under part are strongly tinged
with brown, a spot of a darker hue occurring on each side of the
latter nearly midway between the branchial opercula and the
spinners ; the sexual organs are moderately developed, and have
a brownish-yellow process directed backwards from their anterior
margin: the spinners are eight in number; those constituting
the inferior pair, which are the shortest, consist of a single joint
each, and are united throughout their entire length.
Two adult females of this species were taken in the Island of
St. Nicholas.
Orithyia gnava.
Length of the female 4th of an inch; length of the cephalo-
thorax -!;, breadth 25; breadth of the abdomen +4;; length of
an anterior leg =4-; length of a leg of the third pair 1.
In the relative size and disposition of its eyes, also in the
relative length and proportions of its legs, and in its general
structure, this species is similar to Orithyia luteola. The ce-
phalothorax has a black hue, a short longitudinal streak in the
middle, and a triangular mark at the base, whose vertex is di-
rected forwards, being of a yellowish-white colour. The falces,
maxille, and lip have a pale brownish-yellow hue, the base of
the lip and maxillee, the extremity of the falces, and the sternum
being of a dark-brown colour. The femora, genua, and tibiz
of the anterior legs have a black hue, and that of the slender
metatarsi and tarsi is brownish yellow; the colour of the second,
third, and fourth pairs of legs is brownish black, with a few pale
brownish-yellow annuli. The palpi are of a pale brownish-yellow
colour, the extremity of the humeral joint, the cubital and radial
joints, and the extremity of the digital joint having a brownish-
black tint. The abdomen is soot-coloured, mingled with white,
the sides being the darkest and the under part the lightest-
collected in the Cape de Verde Islands. 91
coloured ; the anterior extremity, contiguous to the cephalothorax,
has a white hue, and comprises a vertical black bar, which is
enlarged at its lower extremity, and is crossed at its upper
extremity by a shorter one of the same hue.
The collection contained two females; but the abdomen of
each was so corrugated that it was not possible to define with
accuracy the design formed by the distribution of its colours ;
the characters given above, however, are sufficient to distinguish
this spider from other known species of the genus. Both speci-
mens were captured in the Island of St. Nicholas.
Family THERrp11p#.
Genus TuEripion, Walck.
Theridion fallax.
Length of the female trd of an inch; length of the cephalo-
thorax +, breadth +; breadth of the abdomen +; length of an
anterior leg 2; length of a leg of the third pair -3,.
The cephalothorax is convex, nearly oval, slightly compressed
before, rounded on the sides, which are marked with furrows
converging towards a large indentation in the medial line, and
is thinly clothed with short, whitish hairs ; the falees are conical
and vertical; the maxille are obliquely truncated at the extre-
mity, on the outer side, and are inclined towards the lip, which
is semicircular ; and the sternum is heart-shaped. These parts
have a very dark-brown hue; the extremity of the falces is
tinged with red, particularly on the inner side, and the extremity
of the maxille, the apex of the lip, and a spot on the sternum
opposite to the base of the lip have a yellowish-brown tint.
The eyes are disposed on the anterior part of the cephalothorax
in two transverse, nearly straight, parallel rows; the four in-
termediate ones form a square, the two anterior ones, which
are seated on a protuberance, being the largest and darkest-
coloured of the eight; the eyes of each lateral pair are placed
on small tubercles, and are separated by a considerable interval,
The legs are long, robust, and provided with hairs, the meta-
tarsi and tarsi being disproportionally slender; they are of a
yellowish-brown colour, the metatarsi and tarsi, which are much
the palest, having a slight tinge of red; and the extremity of
the femora and tibize, and the genua, have a brownish-black hue;
the first pair is the longest, then the fourth, and the third pair
is the shortest; each tarsus is terminated by three claws; the
two superior ones are curved and pectinated, and the inferior
one 1s inflected near its base. The palpi are short, and have a
curved, pectinated claw at their extremity; they resemble the
legs in colour, but are without any brownish-black marks. The
Vie
92 Mv. J. Blackwall on recently discovered Spiders
abdomen is short, broad, oviform, thinly clothed with hairs,
very convex above, and projects ereatly over the base of the
cephalothorax ; it is of a dark-brown colour above, minutely
spotted with white ; in the medial line of the anterior part there
are a few irregular white lines encompassing small dark-brown
spaces; several rather obscure, slightly curved, whitish, trans-
verse lines occur above the spinners, and at the base of those
organs, on each side, there are two minute spots of the same
hue; the sides are of a pale dull yellowish-white colour, re-
ticulated with brown, and are marked with long, oblique,
whitish bands curved downwards and narrowly bordered
with brownish black; these bands, which taper to their lower
extremity, comprise in their broader part a dark-brown space,
and their superior extremity, which is abruptly contracted, is
curved upwards; the under part is paler than the sides, and
has a large white mark in the middle, reticulated with pale and
bordered with dark brown ; the anterior part of this mark is the
broadest, and comprises an oblong-oval, brownish-black spot on
each side of the medial line; and the middle of its posterior
extremity is produced in the form of a small semicircle; the
branchial opercula have a brown hue, and that of the sexual
organs, which are well developed and rather prominent, is dark
red-brown.
One female of this species, which is nearly allied to the spiders
of the genus Latrodectus by the disposition of its eyes, was taken
in the Island of St. Antonio. In consequence of its abdomen
being much disfigured, it was not possible to describe with per-
fect accuracy the desi ign formed upon it by the distribution of
its colours; however, from the characteristics given above, there
can be no difficulty in identifying it.
Theridion quinquenotatum.
mength of the female ths of an inch; length of the cephalo-
thorax -!,, breadth ’,; breadth of the abdomen ts 3 length of
a posterior leg 4+; length of a leg of the third pair +!
The eyes are disposed on the anterior part of the BA
in two transverse rows; the four intermediate ones describe a
square, the two anterior ones, which are rather the smallest and
darkest-coloured of the eight, being seated on a slight protuber-
ance; the eyes of each lateral pair are placed obliquely on a
tubercle, and are contiguous. The cephalothorax is oval, convex,
glossy, with an indentation in the medial line of the posterior
region, and slopes gradually from the anterior extremity to the
base; the falces are conical, vertical, and armed with a few teeth
at the extremity on the inner surface ; the maxille are obliquely
truncated at the extremity, on the outer side, and strongly in-
collected in the Cape de Verde Islands. 93
clined towards the lip, which is semicircular; and the sternum
is heart-shaped. These parts have a red-brown hue, the lateral
margins of the cephalothorax and sternum and the base of the
lip being soot-coloured. The legs are slender, provided with
hairs, and of a red-brown hue, with the exception of the femora,
which are black, and the extremity of the tibize of the first and
fourth pairs, which are soot-coloured; the fourth pair is the
longest, then the first, and the third pair is the shortest ; each
tarsus is terminated by three claws; the two superior ones are
curved and pectinated, and the inferior one is inflected near its
base. The palpi resemble the legs in colour, and have a curved
pectinated claw at their extremity. The abdomen is oviform,
thinly clothed with hairs, convex above, projecting over the base
of the cephalothorax, and is of a brownish-black hue ; an oblique
white spot is situated on each side of its anterior extremity,
another near the middle of the upper part of each side, and an
oblong one immediately above the spinners, in the medial line ;
the branchial opercula have a brownish-yellow hue, and that of
the sexual organs, which are moderately developed, is dark red-
brown.
Two females of this species were taken in the Island of
St. Antonio.
Theridion sagax.
Length of an immature male 2ths of an inch; length of the
cephalothorax ~3;, breadth 1; breadth of the abdomen +; length
of an anterior leg 3; length of a leg of the third pair 44.
The cephalothorax is convex, glossy, and nearly oval, being
slightly compressed before, and rounded on the sides, which are
marked with furrows converging towards an indentation in the
medial line; the falces are short, conical, and vertical; the
maxille are obliquely truncated at the extremity, on the outer
side, and inclined towards the lip, which is semicircular; the
sternum is heart-shaped; the legs are long, and provided with
hairs ; the first pair is the longest, then the fourth, and the third
pair is the shortest ; each tarsus is terminated by three claws ;
the two superior ones are curved and pectinated, and the inferior
one is inflected near its base; the palpi are short; the radial is
larger than the cubital joint, and the tumid digital joint is some-
what oviform; but, as the palpal organs were not developed, it
is evident that the spider was immature. These parts are of a
yellowish-brown colour, the base of the lip and an oblique trans-
verse bar near the extremity of the maxille having a red-brown
hue. The eyes are disposed on the anterior part of the cephalo-
thorax in two transverse rows; the four intermediate ones form
a square, the two anterior ones, which are seated on a protu-
berance, being rather the smallest and darkest-coloured of the
94 Mr. J. Blackwall on recently discovered Spiders
eight; the eyes of each lateral pair are placed obliquely on a
tubercle, and are contiguous. The abdomen is large, glossy,
sparingly clothed with hairs, somewhat oviform, convex above,
and projects greatly over the base of the cephalothorax ; it is of
a black hue, and has a curved yellowish-white band at its ante-
rior part, contiguous to the cephalothorax, which passes nearly
to the middle of the upper part of each side, and whose enlarged
extremities are directed upwards; there is a triangular point in
the middle of the interior of this curve; and a series of triangular
yellowish-white spots, comprising a brown longitudinal streak,
and diminishing in size as they approach the spimners, extends
along the middle; a few obscure slightly curved lines of the
same hue occur above the coccyx, and on each side, near the
enlarged extremity of the anterior curved band, there is an ob-
long yellowish-white spot ; the colour of the under part is yel-
lowish white, irregularly marked with dark brown, and a brown
quadrangular space in the middle has a yellowish-white border.
Captured in the Island of St. Antonio.
Family Epririp2.
Genus Errira, Walck.
Epetra mesta.
Length of the male tth of an inch; length of the cephalo-
thorax =!,, breadth ,; breadth of the abdomen ~;; length of
an anterior leg +; length of a leg of the third pair +>.
The legs are slender, provided with hairs, and have a brown-
black hue, with yellowish-brown annuli on the tibize, metatarsi,
and tarsi; the first pair is the longest, then the second, and the
third pair is the shortest; each tarsus is terminated by claws of
the usual number and structure. The palpi are short, and of a
brown colour, the digital joint being much the darkest; the
radial joint is stronger than the cubital, and has a few long, fine,
curved hairs at its extremity, in front; the digital joint is of an
oblong-oval form, with a process at its base curved outwards ;
it is convex and hairy externally, concave within, and the palpal
organs connected with it are highly developed, prominent, com-
plex in structure, with a fine prominent spine near the base, a
curved one on the under side, and a short bifid one near their
extremity; these organs have a yellow-brown hue; and the
convex sides of the digital joints are directed towards each other.
The cephalothorax is oval, convex, glossy, with a large indenta-
tion in the medial line of the posterior region; it is sparingly
supplied with short white hairs, and is of a brown-black colour,
the anterior margin being the brownest. The eyes are disposed.
on the anterior part of the cephalothorax in two transverse rows
collected in the Cape de Verde Islands. 95
at a moderate elevation above the frontal margin; the four in-
termediate ones nearly form a square, the two anterior ones,
which are placed on a protuberance, and are wider apart than
the two posterior ones, being the largest of the eight ; the eyes
of each lateral pair are the smallest, and are seated obliquely on
a minute tubercle near to each other, but are not in contact.
The falees are conical, and inclined towards the sternum, which
is heart-shaped, with small eminences on the sides, opposite to
the legs; the maxillz are short, powerful, and greatly enlarged
at the extremity, which is produced on the inner side; the lip
is semicircular, and prominent at the apex. These parts have a
black hue tinged with brown, the sternum being the darkest,
and the extremity of the maxillee and apex of the lip the palest.
The abdomen is somewhat oviform, with a small conical protu-
berance on each side of its posterior extremity, which extends
considerably beyond the spinners ; it is moderately convex above,
projects over the base of the cephalothorax, and has a black hue;
a small spot composed of short white hairs occurs at the extre-
mity of each conical protuberance, and a third at the base of the
coccyx.
Captured in the Island of St. Nicholas.
Epeira blanda.
Length of the male {th of an inch; length of the cephalo-
thorax =1,, breadth +4;; breadth of the abdomen ;; length of
an anterior leg 2; length of a leg of the third pair 53,.
The cephalothorax is compressed before, rounded on the sides,
somewhat pointed in front, convex, glossy, and has a large in-
dentation in the medial line of the posterior region; it is of a
brown colour, with a tinge of yellow in the middle and in the
region of the eyes, and the narrow lateral margins are soot-
coloured. ‘The eyes are disposed on the anterior part of the
cephalothorax in two transverse rows; the four intermediate
ones nearly form a square, the two anterior ones, which are the
largest and darkest-coloured of*the eight, and are rather wider
apart than those of the posterior row, being situated imme-
diately above the frontal margin; the- eyes of each lateral pair
are seated obliquely on a minute tubercle, and are near to each
other, but not in contact, the posterior ones being the smallest.
The falces are subconical, slightly divergent at the extremity,
and inclined towards the sternum; they are of a brown colour,
the inner surface and extremity having a pale-yellow hue. The
maxillee are short, straight, powerful, and enlarged and rounded
at the extremity; the lip is semicircular, but somewhat pointed
at the apex; and the sternum is heart-shaped. These parts are
of a yellow-white colour, the base of the maxille and lip and the
96 My. J. Blackwall on recently discovered Spiders
margins of the sternum having a dark-brown hue. The legs
are long, and provided with hairs and spines, the tibial joint of
the second pair having numerous short, black, pointed spines on
its anterior surface; the colour of the femora is pale yellow at
the base and dark brown at the extremity, the dark-brown hue
being most extensive on the anterior pair, and merely forming
an annulus on the third pair; the other joints of these limbs
have a brownish-yellow hue, and are marked with dark-brown
annuli; the first pair is the longest, then the second, and the
third pair is the shortest ; each tarsus is terminated by claws of
the usual number and structure. The palpi are short, and of a
pale-yellow colour, with the exception of the digital joint, which
has a brown hue; the cubital joint projects two long curved
hairs from its anterior extremity, in front; the radial is larger
than the cubital joint, prominent on its external and internal
surfaces, and supplied with long hairs; the digital joint 1s oval,
with a glossy process at its base curved outwards ; it is convex
and hairy externally, concave within, and the palpal organs con-
nected with it are highly developed, prominent, with several
short, curved processes at their extremity, and are of a dark-
brown colour mingled with yellow. The convex sides of the
digital joints are directed downwards. The abdomen is oviform,
convex above, projecting over the base of the cephalothorax, and
is thinly clothed with long dark and light-coloured hairs; on
the upper part there is a large brownish-black, leaf-shaped mark,
which gradually decreases in breadth to the spinners, and whose
sinuous margins have an obscure, white, lateral border, forming
at its anterior extremity a conspicuous spot on each side of the
base of a triangular black spot situated at the anterior part of
the abdomen and having its vertex directed forwards; the leaf-
shaped mark comprises in the middle of its anterior part a fusi-
form mark faintly bordered with white; the sides are of a
brownish-black colour mingled with dull white, and the brownish-
black under part has a curved yellowish-white line on each side,
whose posterior extremity is the falest and most conspicuous.
Captured in the Island of St. Nicholas.
Genus Neruiua, Leach.
Nephila Grayit.
Length of the female 1/4; inch; length of the cephalo-
thorax =, breadth -2,; breadth of the abdomen +; length of
an anterior leg 2; length of a leg of the third pair +.
The legs are long, provided with fine spines and hairs, a dense
brush of the latter occurring on the inferior surface and sides of
the tibize of all the legs except those of the third pair; they are
collected in the Cape de Verde Islands, 97
of a yellow colour, with the exception of the extremity of the
femora and tibie, the genua, and the metatarsi and tarsi, which
are of a black hue, the base of the metatarsi of the first and
second pairs having a tinge of yellow; the first pair is the
longest, then the second, and the third pair is the shortest ; the
tarsi are terminated by claws of the usual number and structure.
The palpi are short, and have a yellow hue, the digital joint,
which has a slightly curved, pectinated claw at its extremity,
being strongly tinged with brown. The eyes are disposed on
the anterior part of the cephalothorax in two transverse rows ;
the four intermediate ones nearly form a square, the two anterior
ones, which are seated on a protuberance, and are rather nearer
to each other than the two posterior ones, being the largest of
the eight ; the eyes of each lateral pair are placed obliquely on
a prominent tubercle; they are the smallest, and are separated
by a considerable interval. The cephalothorax is long, rather
convex above, truncated in front, compressed before, moderately
rounded on the sides, which are marked with furrows converging
towards a transverse pair of indentations in the medial line of the
posterior region, and has two convex, glossy eminences situated
transversely near its middle; it is of a brownish-black colour,
which is almost concealed by a covering of short, adpressed,
white hairs having a silvery lustre. The falces are powerful,
conical, vertical, convex at the base, in front, and armed with
teeth on the inner surface ; the maxille are short, strong, and
greatly enlarged and rounded at the extremity ; the lip is some-
what oval. These parts are of a brownish-black colour, the
falees bemg the darkest, and the extremity of the mavxillz and
the apex of the lip have a dull yellow hue. The sternum is ob-
long heart-shaped, glossy, with eminences on the sides, opposite
to the legs, and is of a yellow colour, with brownish-black lateral
margins. The abdomen is oviform, glossy, sparingly clothed
with short pale hairs, convex above, and projects over the base
of the cephalothorax ; it is of a brown colour, tinged with olive,
and has a large, crescent-shaped, yellow mark near the anterior
extremity of its upper part, followed by four spots of the same
hue in the medial line, which diminish in size as they approach
the spinners ; an obscure, curved, yellowish band passes above
each branchial operculum, and a dull spot of the same hue
occurs on the upper part of each side ; four minute yellow spots
on the under side form a transverse row immediately below the
sexual organs, and a large, irregular, yellow spot is situated
midway between those organs and the spinners; the sexual
organs are moderately developed, and, with the branchial oper-
cula and spinners, are of a dark-brown colour. Some speci-
meus have the yellow spots in the medial line of the upper
98 Mr. J. Blackwall on recently discovered Spiders
part of the abdomen divided, which then form two longitudinal
series.
The name of John Gray, Esq., of Wheatfield House, near
Bolton, is associated with this fine species of Nephila (which
abounds in the Island of St. Antonio, and constructs its web
among the branches of trees), in acknowledgement of the obliga-
tion I am under to him for the interesting spiders described in
this paper, and for numerous specimens of Araneidea collected
in Algeria and Rio de Janeiro.
Genus ArGyoprs, Savigny.
Argyopes Clarkii.
Length of the female 7,ths of an inch; length of the cephalo-
thorax 1, breadth .°-; breadth of the abdomen ;5,; length of an
anterior leg 1; length of a leg of the third pair 3.
The cephalothorax is compressed before, rounded in front
and on the sides, slightly convex, with a broad shallow indenta-
tion in the medial line of the posterior region; it is of a dark-
brown colour, which is concealed by a thick covering of short,
adpressed, white hairs, which have a silvery lustre. The eyes
are disposed on the anterior part of the cephalothorax im two
transverse rows; the four intermediate ones are seated on a
prominence and form a square, the two anterior ones being the
largest of the eight; the eyes of each lateral pair are placed
obliquely on a tubercle, and are near to each other, but not in
contact, the anterior ones being much the smallest. The falces
are powerful, conical, vertical, and armed with teeth on the inner
surface; the maxille are short, strong, and greatly enlarged and
rounded at the extremity; the lip is semicircular, but pointed
at the apex; and the sternum is heart-shaped, supplied with
hoary hairs, and has eminences on the sides, opposite to the legs.
These parts are of a dark-brown colour, the imner surface and
extremity of the falees, the extremity of the maxille, the apex of
the lip, a streak in the medial line of the anterior part of the
sternum, and an oval spot at its extremity having a yellow hue.
The legs are long, provided with hairs and spines, and have a
very dark brown hue, the inferior surface of the coxe being
tinged with yellow ; the first pair is the longest, then the second,
and the third pair is the shortest; the tarsi are terminated by
claws of the usual number and structure. The palpi are rather
paler than the legs, and have a curved, pectinated claw at their
extremity. The abdomen, which is oviform, is more convex on
the under than on the upper part; it projects over the base of
the cephalothorax, and its posterior extremity extends beyond
the spmners; the sides are festooned, four somewhat conical
collected in the Cape de Verde Islands. 99
obtuse prominences occurring on each, and there is a smaller
and more pointed one on each! side of its anterior extremity; the
upper part is densely clothed with adpressed silky hairs, and is
of a pale-yellow colour, with small black spots, many of which
are disposed in transverse rows; there are two short, curved,
black lines near the middle, whose coneave sides are directed
towards each other; and along each side a strongly dentated
black line extends; the anterior surface of the lateral promi-
nences, the branchial opercula, spinners, and under part have a
dark-brown hue; and an irregular spot on each side of the last,
a small one in front of the spinners, and a streak at their base,
on each side, are of a pale yellow colour; the sexual organs,
which are well developed, prominent, and of a very dark brown
colour, have a strong process directed from their posterior
margin obliquely forward and downward, whose extremity is
enlarged, convex, and glossy.
This distinctly marked species of Argyopes, which was cap-
tured in the Island of Brava, is dedicated to the Rev. Hamlet
Clark, who, in conjunction with Mr. Gray, has favoured me with
many exotic spiders of great interest.
Genus TreTraGNnaTHa, Latr.
Tetragnatha maculata.
Laigie of the female 1th of an inch ; length of the cephalo-
thorax ;';, breadth =1;; breadth of the abdomen 4'> length of
an anterior leg }+; length of a leg of the third pair +
The abdomen is short, broad, oviform, sparingly clothed with
hairs, convex above, projects over the base of the cephalothorax,
and is of a brownish-black colour marked with white; the ante-
rior extremity 1s white, and near it, on each side of the medial
line of the upper part, there is a rectangular mark, to which
succeed. six spots disposed in pairs, more or less contiguous, and
dimmishing in size as they approach the spinners; two spots
occur on the posterior half of each side, the anterior one of
which is the larger, and somewhat triangular, and situated lower
on the anterior half there is a slightly “curved line; four spots
are disposed in a square on the under part, and there are two
minute ones on each side of the spimners, at their base; these
marks and spots are white, more or less reticulated with brown,
and reflect a silvery lustre: the sexual organs are moderately
developed, and of a dark red-brown colour. The cephalothorax
is compressed before, rounded in front and on the sides, convex,
glossy, with an indentation in the medial line of the posterior
region, and is of a pale brownish-yellow colour, with narrow,
soot-coloured lateral margins. The eyes are seated on black
100 = Myr.J. Blackwall on recently discovered Spiders.
spots, and are disposed on the anterior part of the cephalothorax
in two transverse rows; the four intermediate ones nearly form
a square ; the two anterior ones are the largest of the eight, and
are placed on a slight protuberance; the eyes of each lateral
pair, which are the smallest, are seated on a minute tubercle,
and are almost in contact. The falces are powerful, conical,
vertical, and armed with teeth on the inner surface; the legs
are long, slender, and provided with hairs; the first pair is the
longest, then the second, and the third pair is the shortest ; the
palpi are slender, and have a slightly curved claw at their ex-
tremity. These parts have a pale brownish-yellow hue. The
maxille are slightly divergent, and increase in breadth from the
base to the extremity, which is somewhat angular on the outer
side; and the lip is semicircular and prominent at the apex.
These organs are of a brown colour tinged with yellow, the base
of the hp being the darkest. The sternum is heart-shaped,
glossy, supplied with some long hairs, and has a black hue,
The male is smaller than the female, but resembles her in
colour. The cubital and radial joints of its palpi are short, the
latter being rather the larger; the digital jot is oval, with a
process at its base curved outwards; it has a yellowish-brown
hue, is convex and hairy externally, and concave within, com-
prising the palpal organs, which are well developed, not very
complex in structure, convex, glossy, and terminate in a point
that extends beyond the extremity of the joint: the colour of
these organs is yellowish brown. The convex sides of the digital
joints are directed towards each other.
One male and three females of this Tetragnatha, which be-
longs to Walckenaer’s second family of the genus, the Coadunate,
were captured in the Island of St. Nicholas,
Tribe Senoculina.
Family Scyropipa.
Genus Scyropns, Latr.
Scytodes pallida.
Length of an immature male 4th of an inch; length of the
cephalothorax +1,, breadth 3's; breadth of the abdomen 51, ;
length of a leg of the second pair +; length of a leg of the third
pa 3°.
The eyes, which are small and disposed in pairs on the ante-
rior part of the eephalothorax, are seated on black spots; those
of each lateral pair are placed obliquely on a shght tubercle, and
are almost in contact, the anterior ones being the largest of the
six; the eyes of the anterior pair are situated transversely, in
advance of the lateral pairs, high above the prominent frontal
Mr. Hf. W. Bates on Longicorn Coleoptera. 101
margin, and are contiguous or nearly so. The cephalothorax is
compressed before, rounded 3 in front and on the sides, moderately
convex, glossy, and has an indentation in the medial line; the
falces are conical, rather prominent, and are armed with a short
curved fang, and a single pointed tooth on the inner side, near
the extremity; the maxille are curved towards the lip, and
touch at their extremity, which is truncated on the inner side ;
the lip is large and somewhat triangular, but rounded at the
apex; the sternum is nearly circular, and glossy ; the legs are
long, slender, provided with hairs, and each tarsus is termi-
nated by two curved, pectinated claws; the second pair is the
longest, the fourth pair rather surpasses the first, and the third
pair is the shortest; the palpi are moderately long; the radial
joint is much longer than the cubital, and the digital joint,
which has an oblong-oviform figure, is tumid, but compact,
proving by its undeveloped state that the specimen had not
arrived at maturity. These parts are of a pale dull yellowish
colour, the falces and lip having a tinge of red, and the anterior
part of the cephalothorax a slight tinge of brown. The abdo-
men is of an oblong-oviform figure; it 1s somewhat convex
above, projects a little over the base of the cephalothorax, and
is clothed with pale soot-coloured hairs, particularly on the
upper part ; it has a brownish-white hue, with a faint brownish
band, which tapers to its posterior extremity, extending froin
the base of the upper part, contiguous to the cephalothorax, a
little beyond the middle.
This spider, which was taken in the Island of St. Iago, belongs
to Walckenaer’s family Depresse of the genus Seytodes, and is
very closely allied to the Scytodes erythrocephala of Koch (Die
Arachniden, Band vy. p. 90, tab. 168. figs. 399, 400), but may
readily be distinguished from it, even when immature, by marked
differences in the structure of the palpi, and especially by the
form of the digital joint.
X.— Contributions to an Insect Fauna of the Amazons Valley.
CoLeorters: Lonaicornes. By H. W. Bares, Esq.
[Continued from vol. xv. p. 394. ]
29. Colobothea nevigera, n. sp.
C. modice elongata, postice regulariter attenuata, nigricans, sericea,
vertice thoraceque supra lineis duabus, elytris maculis paucis
discretis, cinereis ; his truncatis, angulis externis spinosis. Long.
i-7i lm. ¢ 2.
Head black, forehead with three ashy lines, cheeks with a
spot of the same colour, and vertex marked with two ashy lines
102 Mr. H. W. Bates on the Longicorn Coleoptera
diverging on the occiput. Antenne greatly elongated and
robust, black, sixth joint ringed with white, tenth joimt with an
exterior white line (3), in the 9 the eighth and eleventh joints
also streaked with white. Thorax blackish, clothed with an
olivaceous silky pile, the upper surface with two tawny-ashy,
slender, nearly parallel lines; sides each with a single similar
line, besides a broader streak above the coxee. Elytra broad at
the base, with prominent and not markedly oblique shoul-
ders, regularly attenuated thence to the apex, which is truncated
and has the external angles produced into spines; the surface
has a few fine punctures surmounted by acute granulations to-
wards the base, and beset with short black bristles; the colour
is blackish, clothed with silky olivaceous pile, and ornamented
with a small number of scattered and distinct, rounded, tawny-
ashy spots, the extreme apex having an ashy-white border de-
creasing in width from the suture to the external angle. Body
beneath black, thinly clothed with ashy tomentum ; the sides of
the breast have a tawny-ashy streak im continuation of the one
on the prothorax, and the sides of the abdomen are spotted with
the same colour. The legs are blackish, ringed with grey.
g. Terminal ventral segment narrowed to the apex, trun-
cated, with the angles produced into stout spines; dorsal seg-
ment obtuse. Legs stout; anterior tarsi moderately dilated and
frmged. In the smaller males the legs are not perceptibly
thicker than in the females.
?. Terminal abdominal segment projecting considerably be-
yond the apex of the elytra, broad; dorsal segment notched,
ventral truncated, angles not produced.
A common insect at Ega and S. Paulo, Upper Amazons.
30. Colobothea lucaria, n. sp.
C. modice elongata, nigra, vertice lineis duabus divergentibus, tho-
race lineis tenuibus quatuor, elytris maculis paucis hic illic con-
gregatis, griseis; his apice cano marginatis, oblique truncatis,
angulis externis spinosis. Long. 5 lin. ¢.
Head black, forehead with three obscure grey lines, vertex
with two divergent lines of similar colour, and the posterior part
of the orbits Aiko grey. Antenne black, ibe of fourth, eighth,
and tenth joimts grey on one side, Sean jot with a whine
ring. ‘Thorax black, with a silky olivaceous gloss, upper surface
with two slender parallel grey lines, each side also with a similar
line visible in part when the insect is regarded from above ;
there is also a grey line above the coxee. Elytra prominent, and
scarcely oblique at the shoulders, thence eradually attenuated
to the apex, which is on each side obliquely truncated, 2. e. the
sutural portion is more advanced than the lateral angles, which
of the Amazons Valley. 103
.
are produced into spines; the surface is finely punctate-granu-
late, and of the same colour as the thorax; the grey spots are
nearly all of equal size and distinct; but they are collected
partly mto groups, and here and there confluent ; the grey
apical margin is of equal width from the sutural to the external
angle. Body beneath thinly clothed with grey pile; sides of
breast not striped with thicker tomentum. Legs black, ringed
with grey.
¢. Terminal ventral segment truncated, angles produced into
short spines; dorsal segment rounded. Anterior tarsi mode-
rately dilated and fringed.
S. Paulo, Upper Amazons. Very closely related to C. nevi-
gera, differing only in the oblique truncature and somewhat
different arrangement of spots of the elytra.
31. Colobothea crassa, n. sp.
C’. major, robusta, nigra, tomento olivaceo-griseo vestita, vertice tho-
raceque dorso lineis duabus divergentibus, elytris maculis nu-
merosis, minimis, discretis, fulvo-griseis, apice cano marginatis.
Long. 8-10 lin. 3 Q.
Differs from C. nevigera in being of much larger size, in the
spots of the elytra being very much smaller and more numerous,
and in the dorsal lines of the thorax being posteriorly divergent.
In shape and in colour the two species offer no tangible point
of difference. As in C. nevigera, there are only two thoracic
hnes visible from above, although there is a lateral line on each
side and a broader streak above the coxe (yellower in colour
and extending to the abdomen) ; the form of the terminal abdo-
minal segment in both sexes offers also no difference in the two
species. C. crassa is still more closely alhed to a Cayenne spe-
cies, C. lineatocollis* (Dej. Cat.), which is similar to it in size
- and other respects, and differs chiefly in the multitudinous grey
* Colobothea lineatocollis (Dej. Cat. sec. Dom. Chevrolat). Elongata,
antice et postice attenuata, nigra, obscure olivaceo-grisea, sericea,
griseo lineata et maculata. Caput nigrum, griseo lineatum, vertice
lineis griseis duabus postice divergentibus, genis griseo plagiatis,
utennse validee, nigra, articulo sexto albo annulatos Thorax lineis
tenuibus duabus dorsalibus subparallelis, alteris duabus lateralibus,
vittaque utrinque supracoxali, griseis. Elytra postica modice at-
tenuata, humeris parum obliquis, apicibus truncatis, angulis externis
dentiformibus, supra sparse punctata maculis minutis griseis con-
fluentibus, reliquo spatio subapicali immaculato, ipso apice albo mar-
ginato. Corpus subtus nigrum, griseo sparse tomentosum, abdomine
maculato. Pedes nigri, griseo annulati. Maris segmento dorsali ter-
minali truncato, angulis prominulis; ventrali profunde emarginato,
angulis spinosis. Foeminz segmento ultimo dorsali apice lato ; ven-
trali profunde emarginato, angulis productis, Hab. in Cayenna.
104 Mr. H. W. Bates on the Longicorn Coleoptera
specks of the elytra being confluent and forming irregular mar-
bled lines. C. Osculatit of Guérin (Cat. des Ins. Col. recueillis
par Gaetano Osculati, no. 261) appears to be another alhed form
similar in size and colours to C. crassa and C. lineatocollis ; but
the description given of the thoracic markings (“ quatre fines
lignes longitudinales blanches”) leaves us in doubt whether
there are not four lines on the upper surface, which would re-
move the species from the neighbourhood of the two mentioned ;
for, if the lateral lines are to be included, the description ought
to mention six instead of four. The distinctive character of C.
crassa is the minute and equal size, great number, and equidis-
tant position of the grey specks of the elytra.
Common in the neighbourhood of Para. C. Osculatii is pro-
bably a native of the banks of the Napo, where M. Osculati
formed his collection.
32. Colobothea ordinata, n. sp.
C. elongata, postice attenuata, olivaceo-nigra, vertice postice bi-
lineato; thorace supra lineis quatuor crassiusculis vittaque lata
supracoxali fulvo-cinereis ; elytris maculis numerosis subquadratis
fulvo-cinereis in seriebus subordinatis; thorace ante basin utrin-
que breviter tubereulato. Long. 72 lin. ¢.
Head black, forehead with three slender lines, vertex with
two divergent lines, and cheeks with a broad streak, tawny
ashy; there is also a tawny-ashy streak behind each eye. An-
tenn stout, black, sixth joint with a narrow white ring, the
bases of the fourth, eighth, tenth, and eleventh joints with an ashy
streak on one side(¢). Thorax slightly constricted at the base,
and with a small tubercle on each side ; surface black, with four
rather thick tawny-ashy lines; there is also a broad tawny-ashy
vitta above the coxa on each side. Elytra with prominent and
rather acute shoulders, thence gradually attenuated to the apex,
which latter is truncated, the external angles produced each into
a longish spine; surface olivaceous black, marked with a large
number of well-separated and squarish tawny-ashy spots, mostly
arranged in rows, and leaving a distinct belt beyond the middle and
another near the apex unspotted ; apex itself edged with whitish.
Body beneath ochraceous ashy. Legs greyish, varied with
black.
g@. Terminal ventral segment broadly truncated, angles pro-
duced; dorsal segment obtuse, entire.
Ega; rare.
33. Colobothea subtessellata, n. sp.
C. clongata, postice attenuata, olivaceo-nigra, vertice postice bi-
lineato, thorace lineis duabus dorsalibus crassiusculis alteraque
laterali et vitta supracoxali cinereo-ochraceis ; elytris maculis nu-
of the Amazons Valley. 105
merosis cinereo-ochraceis in seriebus subordinatis, spatio lato api-
cali immaculato ; thorace absque tuberculis. Long. 81 lin. 9°.
Head black; forehead with three slender lines, vertex with
two divergent lines, and cheeks with a broad streak tawny ashy.
Antenne stout, black, sixth joint with a broad white ring, tenth
joint with an ashy streak on one side(¢). Thorax not con-
stricted at the base, broadest at its basal angles, and free from
tubercles ; surface black, with two moderately thick tawny lines,
sides each with a similar line, not visible from above, and a
broad tawny vitta above the coxa. Elytra moderately broad at
the shoulders, and narrowed thence to the apex, the latter trun-
cated, with the outer angles spinose; surface olivaceous black,
marked with a large number of tawny spots, which are in some
examples arranged in rows, and in others more or less confused ;
there is a broad immaculate space at the apex, and the apex
itself is broadly margined with white. Body beneath black,
thinly clothed with ashy pile, and having a broad, distinct,
ochreous lateral vitta. Legs blackish, ringed with grey.
?. Terminal abdominal segment elongated and tapering ;
dorsal plate broadly notched; ventral truncated, angles acute.
Banks of River Tapajos; rare.
34. Colobothea octolineata, n. sp.
C. valde elongata, postice attenuata, olivaceo-nigra, vertice linea
unica, genis utrinque lineis duabus cinereis; thorace lineis tenui-
bus cinereis octo, quarum quatuor dorsalibus ; elytris humeris
prominentibus, maculis cinereis discretis irregulariter dispersis.
Long. 73-ll lin. ¢ Q.
Head black, forehead with two greyish lines, vertex with a
single narrow line, and cheeks on each side with two oblique
greyish lmes. Antenne black, sixth joint thickened, with a ring
of dense white hairs in both sexes. Thorax marked with eight
slender, greyish or tawny lines, of which four are on the upper
surface and two on each side, including the supracoxal streak,
which in this species is slender, like the other lines. Elytra
greatly elongated ; shoulders very prominent, thence gradually
narrowing to the apex, the latter truncated, with outer angles
spinose ; surface olivaceous black, marked with a moderate num-
of larger and smaller spots, widely separated from each other,
but very irregularly dispersed ; apex edged with whitish. Body
beneath black, marked with ashy or tawny streaks and spots.
Legs black, ringed with tawny and grey.
¢. Terminal abdominal segment narrowed from the base;
apex of both dorsal and ventral plates emarginate-truncate.
Anterior tarsi very broady dilated and fringed.
Ann. & Mag. N. Hist. Ser.3. Vol. xvi. 8
106 Mr. H. W. Bates on the Longicorn Coleoptera
? . Terminal abdominal segment elongate and tapering ; apex
of both dorsal and ventral plates emarginate-truncate.
Para, also Ega, Upper Amazons; common.
35. Colobothea contaminata, Serville.
Colobothea contaminata, Serv. Encycl. Méth. x. p. 337.
C. valde elongata, angustata, postice vix attenuata, olivaceo-nigra,
vertice linea unica cinereo-fulva, thorace lineis quatuor, quarum
externa utrinque usque ad oculum extensa et vitta supracoxali
supra genas continuata; elytris maculis cinereo-fulvis passim
confluentibus, vel cinereo-fulyis nigro irregulariter maculatis, fas-
cia lata subapicali nigra; antennis utroque sexu nigris, articulo
sexto annulo incrassato albo. Segmento ultimo abdominali maris
attenuato, apice emarginato; foeminee angustato, lamina dorsali
obtusa, ventrali angulis productis ; maris tarsis anticis valde dila-
tatis. Long. 64-10 lin. 3 Q.
Generally distributed and common throughout the Amazons
region ; also found at Cayenne.
36. Colobothea geminata, n. sp.
C’. elongata, postice vix attenuata, olivaceo-nigra, vertice linea unica,
thorace lineis duabus antice et postice conjunctis ; elytris maculis
numerosis in lineis curvatis confluentibus fulvo-griseis. Long.
74-8 lin. SQ.
Head black, forehead with three tawny-grey lines, and vertex
with a single line; cheeks with a tawny-ashy stripe. Antenne
black, sixth joint with a broad white ring. Thorax black, sides
each with two tawny-ashy stripes jomed together near the front
and posterior margins, and continuous with the cheek-stripe.
Elytra elongated, of very nearly the same width from base to
apex in both sexes; external angle of the truncature spinose ;
surface blackish olivaceous, sprinkled with a large number of
tawny-ashy spots, which are mostly confluent, and tend to form
a pattern consisting of three irregular pale rings, on each ely-
tron, enclosing a blackish space; apex edged with tawny whitish.
Body beneath ashy, but tawny towards the sides; ; abdomen
spotted with black. Legs ashy, ringed with black.
g. Terminal abdominal segment short; ventral plate emar-
ginate-truncate, angles produced; dorsal plate obtuse and
notched in the middle. Anterior tarsi not dilated.
?. Terminal abdominal segment tapering; dorsal plate
notched in the middle ; ventral truncate, angles not produced.
Guiana side of the Lower Amazons and banks of the Tapajos ;
also found at Cayenne.
37. Colobothea concreta, n. sp.
C. valde elongata, angustata, olivaceo-nigra, vertice linea unica, tho-
of the Amazons Valley. 107
race vittis quatuor (quarum duabus externis usque ad oculos ex-
tensis) fulvo-cinereis; elytris basi thorace vix latioribus, apice
truncatis, angulis externis spinosis, maculis cinereo-fulvis con-
fluentibus dense vestitis, apice macula magna nigra. Long. 6—9
lin: oO
Head black, forehead streaked with tawny ashy, vertex with a
single line ; occiput on each side with a short line continuous with
the external thoracic stripe, cheeks with a transverse stripe con-
tinuous with the supracoxal vitta. Antennee black, sixth joint with
a broad white ring. Thorax black, surface with four rather thick
tawny-ashy lines, sides having only the supracoxal vitta. Elytra
elongated, scarcely tapering; shoulders very oblique, and not at
all prominent ; apex truncate, external angles spinose; surface
very thickly clothed with tawny or tawny-ashy spots, mostly
confluent, but leaving a broad unspotted space at the apex, the
latter margined with tawny white. Body beneath ashy, sides
streaked with tawny ; abdomen spotted with black. Legs black,
ringed with tawny and grey.
dg. Terminal abdominal segment narrowed from the base ;
dorsal plate deeply notched; ventral plate semicircularly emar-
ginated, with angles much produced. Anterior tarsi widely
dilated and fringed.
?. Terminal abdominal segment elongate and much nar-
rowed; dorsal plate very obtuse, ventral truncated, angles
slightly prominent.
Para, and banks of the Tapajos.
38. Colobothea bilineata, n. sp.
C. valde elongata, postice vix attenuata, nigra, vertice linea unica,
thorace lineis duabus usque ad oculos extensis, griseis; elytris
griseis, nigro dense maculatis, apice macula magna nigra. Long.
7-103 lin. C.
Head black, forehead streaked with ashy, vertex with a single
line, occiput on each side with a short line continuous with the -
thoracic stripe; cheeks crossed by an ashy streak continuous
with the supracoxal vitta. Antennz black, sixth joint with a
broad white ring. Thorax black, surface with only two ashy
stripes, each continuous to the hind margin of the eye. Elytra
elongate and scarcely tapering, very little broader at the base
than the thorax, but shoulders prominent and conical; apex
sinuate-truncate, the sutural angles being prominent and acute,
the outer angles spinose; surface grey, thickly spotted with
black ; some of the spots confluent, and a large spot at the apex
spotless; apex itself edged with white. Body beneath thinly
clothed with grey; abdomen spotted with black. Legs black,
ringed with grey.
Sk
108 Mr. H. W. Bates on the Longicorn Coleoptera
&. Terminal abdominal segment with the ventral plate semi-
circularly emarginated, angles acute; dorsal plate triangularly
emarginated. Anterior tarsi dilated and fringed.
Ega and 8. Paulo, Upper Amazons ; rare.
39. Colobothea lunulata, Lucas.
Colobothea lunulata, Lucas, Voyage de Castelnau, Entomologie, p. 190,
pl. 13. f. 5 (1857).
—— Fryi, Pascoe, Trans. Ent. Soc. vol. i. 41 (1861).
C. elongato-elliptica, nigra; vertice, thorace et elytris albo bivittatis,
vittis longe ante apicem elytrorum convergentibus et annulo albo
utrinque connexis. Long. 73-93 lin. ¢ 2.
This very distinct and handsome species was one of the com-
monest of its genus at Ega, on the trunks of fallen trees in the
forest. The shoulders are extremely oblique and scarcely pro-
minent, so that the insect has the form of an elongated ellipse
truncated at the elytral end. The terminal abdominal segment
in the male has both the dorsal and ventral plates truncated ;
in the female it is elongated, and the angles of the ventral plate
are produced. The anterior male tarsi are widely dilated and
fringed*,
* The following species of Colobothea have not yet been described :—
Colobothea hebraica (Chrevrolat, MS.). Modice elongata, postice at-
tenuata, fusco-nigra, griseo maculata. Caput nigrum, fronte griseo
trilineata, occipite maculis duabus, genis vitta lata, griseis. Antenne
nigra, articulis basi griseis. Thorax basi paulo angustatus, dorso
linea abbreviata, disco utrinque maculis parvis, lateribus vitta latius-
cula cinereo-griseis. Elytra apud humeros lata, deinde usque ad
apices attenuata, truncatura angulis externis spinosis, supra fusco-
nigra maculis cinereo-griseis (partim subagglomeratis) adspersa, apice
haud pallide marginato. Corpus subtus griseum, lateribus cinereis,
nigro maculatis. Pedes nigri, cmereo annulati. Foeminee segmentum
ultimum abdominale attenuatum; lamina dorsali apice rotundata, ven-
trali truncata, angulis productis. Long. 5-7 lin. 2. Had. in Mexico.
Colohothea fusciata. Modice elongata, postice valde attenuata, tomento
brunneo fulvo-maculato vestita; elytris fascia lata nigro-velutina.
Caput nigrum, fulvo-brunneo vestitum, vertice limea unica fulva.
Antenne breviores, nigra, breviter setosz, articulis basi griseis.
Thorax fusco-niger, dorso vittis duabus fulvo-brunneis. Elytra apud
humeros lata, deinde valde attenuata, apice sinuato-truncata, angulis
externis longe spinosis, supra brunnea obscure fulvo maculata, pone
medium fascia nigro-velutma apud dorsum dilatata, apices versus
nigro liturata. Corpus subtus rufescens, medio nigricans. Pedes
nigri. Maris segmentum ultimum ventrale subtumidum, apice obtuse
truncatum ; tarsi antici haud dilatati. Foeminze segmentum ultimum
paulo elongatum, valde attenuatum, lamina ventral: sinuato-truneata,
haud spmosa. Long. 4-6. 3 2. Hab. in Rio Janeiro.
Colobothea lateralis. Elongata, postice valde attenuata; corpore supra
cinereo-ochraceo, rufo variegato, lateribus nigris. Caput nigrum,
fronte fulvescente, vertice et maculis quatuor occipitalibus cinereis
of the Amazons Valley. 109
Subtribe Lamuirm.
Genus TantoTEs, Serv.
Serville, Ann. Soc. Ent. Fr. iv.
This well-known and handsome genus is the only one belong-
ing to the typical Lamiaires found in the Amazonian forests,
the allied genus Ptychodes, common in other parts of Tropical
America, being absent from the low-lying Equatorial region.
The other Tropical American representants of this subtribe, so
rich in forms in the Old World (namely, Plectodera, erm
derus, and Deliathis), seem to be confined to the northern por-
tion of the zone—Central America, Mexico, and thence extend-
ing into the Southern States of North America.
1. Teniotes decoratus, Castelnau.
Teniotes decoratus, Casteln., Animaux articulés, ii. p. 479.
T. nigro-velutinus, capite fascia utrinque infra oculos, vitta laterali
alteraque coronali per thoracem et scutellum continuata, maculis-
que rotundis elytrorum utrinque circa 13 leete flavis ; corpore sub-
tus vitta flava laterali: maris pedibus anticis vix elongatis, tarsis
haud pilosis. Long. 13 lin. ¢ 2.
I met with this fine species only in the neighbourhood of
Para, on felled trees in broad roads through the forest. The
terminal ventral segment in both sexes is broadly truncated,
with a distinct spine at each angle. M. Guérin-Méneville (Icon.
Reégne Animal, p. 243) believes this species to be the same as
the 7. subocellatus of Olivier (Ent. no. 67. pp. 69, 89, pl. 2.
f. 12a, 6), and that the latter is founded on a worn or immature
individual.
2. Teniotes D’ Orbignyi, Guérin.
Teniotes D’ Orbignyi, Guérin-Méneville, Icon. Régne Animal, p. 444.
7’. nigro-velutinus, capite fascia utrinque infra oculos, vitta laterali,
alteraque Perea per thoracem et scutellum continuata, vittaque
elytrorum utrinque medio interrupta et maculiformi leete flavis ;
corpore subtus vitta flava laterali: maris pedibus anticis vix
elongatis, tarsis haud pilosis. Long. 8-l3 lin. ¢ Q.
This le only discovered in the wooded plains of
fulvo ee Antennz grisez, articulis apice nigris. Thorax an-
tice angustatus, dorso cinereo-ochraceus, rufo maculatus, lateribus
nigris. Elytra apud humeros lata, deinde attenuata, apice truncata,
angulis extermis spinosis, supra cinereo-ochracea, rufo maculata, la-
tenbus irregulariter nigris, nigredine ramos tres dentatos in discum
emittente, ine rufo marginatis. Corpus subtus cinereum, medio
nigrum, segmentis primo et ultimo abdominalibus nigris. Foeminz
segmentum ultimum attenuatum, lamina ventrali truncata, dorsali
medio emarginata. Long. 73 lm. 2. Hab, in Brasilia,
110 Mr. H. W. Bates on the Longicorn Coleoptera
Bolivia by M. D’Orbigny, was common on the Upper Amazons
at Ega. The yellow (partially macular) stripe of the elytra
varies a little in the degree in which it is broken up into spots ;
but it never forms a double row of distinct round spots from
base to apex, as shown in 7. decoratus, and can scarcely be con-
sidered a local form of the same stock.
3. Teniotes Amazonum, Thomson.
Teniotes Amazonum, Thoms. Archives Entomologiques, i. p. 172.
T. niger, capite linea curvata frontali, vitta utrinque laterali, altera
coronali per thoracem scutellum et elytros continuata (hic dentata)
pallide flavis; thorace utrinque linea tenuissima grisea; elytris
maculis parvis numerosis, quarum duabus vel tribus discoidalibus
majoribus, flavis; corpore subtus vitta flava laterali: maris pedi-
bus anticis valde elongatis, tibiis curvatis, tarsis haud pilosis.
Long. 9-16 ln. ¢ Q.
A common insect in the forest at Ega, on the Upper Amazons.
It is probably a local form of 7. scalaris, Fabr., but differs much
from the description given by that author. The terminal ventral
plate is formed as in 7’. decoratus.
4. Teniotes farinosus, Linneus.
Cerambyx farinosus, Linn. Syst. Nat. 1. 626. 24; Oliv. Ent. Ixvii. p. 50,
f. 46a.
— pulverulentus, Oliv. Ent. Ixvii. p. 50, f. 46 6.
T. niger, griseo vestitus ; capite thoraceque lineis tenuibus tribus,
elytris maculis numerosis parvis, flavo-griseis, his apice acutis ;
corpore subtus flavo maculato: maris pedibus anticis valde elon-
gatis, tibiis curvatis, tarsis hirsutis. Long. 13 lin. ¢.
This species was a rare one in the Amazons region, and found
only in the dry forests of the Tapajos. The spines of the ter-
minal ventral segment are more elongated than in the other
species.
Subtribe ONcIDERITA.
Group Onciderine.
Genus HypseLomvus, Perty.
Perty, Delectus Anim. Articul. Brasil. p. 95 (1830-34).
Syn. Hypsioma, Serv. Ann. Soc. Ent. Fr. p. 38 (1835).
This genus is distinguished from its allies by its short sub-
trigonal form of body, with projecting and often acute shoulders
of the elytra. The claw-joint of the tarsi is not so much elon-
gated as in Oncideres, or even Clytemnestra. It is very closely
allied to the latter genus, but is distinguishable at once by the
abrupt clavate form of the basal joint of the antenne and the
of the Amazons Valley. 111
curved shape of the third. The males of most species have a
short, slender, curved joint at the tip of the eleventh joint of
the antennz, which is sometimes visible (but much smaller) in
the female.
1. Hypselomus basalis, Thomson.
Hypsioma basalis, Thomson, Classif. des Cérambye. p. 117.
H. modice elongatus, brunneus ; capite, thorace et elytrorum parte
antica rufescenti-ochraceis; summa fronte acute bituberculata ;
antennis nigris, basi rufescenti-ochraceis, articulis ceeteris basi
rufescentibus ; elytris basi utrinque vix elevatis, nigro tuberculatis
humeris, apice nigris; abdomine lateribus rufo vittatis; pedibus
nigricantibus, tibiis compressis, posticis( ¢ ) apice dilatatis. Long.
6-9 lin. ¢ Q.
A common insect throughout the Amazons region, being
found, like the rest of the species, on dead branches, closely
adhering to them, and gnawing the bark and wood all round,
until the bough is sometimes severed. The face and parts
of the mouth are much elongated and directed a little back-
wards between the anterior haunches, so that when the legs are
extended, grasping a branch, the jaws are in a good position to
gnaw effectually. The supplementary joint of the antennz is
very conspicuous in the males of this species.
2. Hypselomus picticornis, n. sp.
H. suboblongus, brunneus, elytris fascia obliqua indistincta_palli-
diore ; antennis brunneis, articulo 2° toto et caeteris basi rufes-
centibus ; elytris basi haud tuberculatis, humeris oblique conicis
modice productis. Long. 7 lin. @.
Head brown, forehead near base of antennz with two very
small conical tubercles. Antenne about the length of the body,
setose beneath ; basal joint strongly and abruptly clavate, third
much bent, dark brown; second joint, basal half of third, and
bases of each remaining joint pallid-reddish. Thorax scarcely
uneven on the surface, uniform dingy brown. Llytra oblong
trigonal; shoulders moderately prominent, and thence gradually
narrowed to the apex, which is broadly rounded; surface con-
vex ; centrobasal ridges not at all prominent, and quite destitute
of tubercles, the basal half of the elytra being simply punctured.
Body beneath rufescent tawny, centre of abdomen black ; legs
brown, claw-joints of tarsi with their basal halves pale reddish.
Ega; rare.
3. Hypselomus Amazonicus, Thomson.
Hypsioma Amazonica, Thomson, Classif. des Céramb. p. 119.
H. convexus, brunneus ; elytris humeris conicis, subuncinatis, pone
medium fascia irregulari pallidiore, deinde ad apices pallide mar-
112 Mr. H.W. Bates on Longicorn Coleoptera.
moratis ; antennis articulis basi rufescentibus: maris tibiis posticis
apice valde dilatato-compressis. Long. 9 lin. 3 2.
Closely allied to H. picticornis, but larger and darker, with
the elytra behind the middle much more variegated with pale
ashy brown, and the conical protuberances of the shoulders
strongly curved anteriorly. The antenne are coloured as in H.
picticornis, the second and basal half of the third, with bases of
the remaining jomts being pale reddish. The underside of the
body is tawny brown, with the centre of the abdomen black.
The elytra are smoothly and strongly convex from base to apex,
without any trace of centrobasal ridge or tubercles.
Ega, Upper Amazons.
4. Hypselomus dimidiatus, n. sp.
H. modice convexus, fuscus, fulvo irroratus; elytris apud medium
ochraceo fasciatis, deinde usque ad apices pallide ochraceo-brun-
neis fusco striatis et maculatis; thorace supra quinquetuberculato,
lateribus acute tuberculatis. Long. 6-7 ln. ¢ @.
Head dingy brown. Antenne dull brown, base of each joint,
from the third, pallid-reddish. Thorax uneven, disk on each
side with two prominent tubercles, and dorsal line elevated be-
hind into a ridge, sides each with an acute tubercle; dingy
brown. Elytra with very prominent shoulders, the anterior side
of the subconical projection oblique; centrobasal ridges slightly
elevated, but not tuberculated; dark brown, sprinkled with ful-
vous ; behind the middle a pale oblique belt or broad triangular
spot darker in the middle, thence to the apex light brown with
darker lines and spots. Body beneath tawny brown, middle of
abdomen black. Legs black, apex of thighs fulvous, claw-joint
red, apex black. Posterior tibize in the male dilated at the apex;
supplementary antennal joint in the same sex very short or
wanting.
Ega. Rather variable in the colour of the posterior part of
the elytra, the pale belt being sometimes extended into a large
triangular patch, and sometimes blended with the pale-brown
shade of the apical half of the wing-cases. The species seems
to be very closely allied to H. subfasciata, Thomson (Classif. des
Céramb. p. 118).
5. Hypselomus rodens, n. sp.
H. oblongus, nigro-fuscus, carneo-fulvo strigatus ; thorace supra haud
tuberculato; elytris humeris apice truncatis, postice uncinatis,
pone medium fascia obliqua pallida. Long. 6 lin. 9.
Head dingy black, crown sprinkled with reddish tawny. An-
tennee black, sprinkled with tawny ; base of each joint, from the
fourth, pale. ‘Thorax convex above, and free from tubercles,
Occurrence of recent Shells in the fossil state near Melbourne. 118
sides with an inconspicuous tubercle. Elytra oblong, shoulders
prominent, but the apex of the cone largely truncated, with the
posterior edge of the truncature projecting ; surface coarsely
punctured, blackish, streaked with reddish tawny, behind the
middle tawny streaked with black, the tawny part separated
from the anterior darker portion by a pale-ochreous fascia.
Body beneath tawny, middle of abdomen black. Legs tawny,
sprinkled with black, base of claw-joint reddish.
Para.
[To be continued. ]
XI.—On the Occurrence of Limopsis Belcheri, Corbula suleata,
and some other recent Shells in the fossil state in Miocene
Tertiary Beds near Melbourne. By Frupuricx M‘Coy, Pro-
fessor of Natural Science in the University of Melbourne, and
Director of the Melbourne National Museum, &e.
Havine occupied myself lately, in my capacity of Paleontologist
to the Geological Survey of Victoria, with the investigation of
the Tertiary fossils collected by the Survey Staff from the strata
of Bird-Rock Bluff, near the mouth of Spring Creek, about
fifteen miles south of Geelong, I was much struck with the
geographical distribution of the very few recent species found
associated with the large majority of extinct species in a rich
fossil fauna unmistakeably of the Lower Miocene age. The
whole facies of the fossil contents of these beds resembles closely
that of the Lower Miocene beds of Doberg (near Biinde, West-
phaha), Malta, and some other European beds of the same age,
as well as the so-called Upper Eocene North-American beds
near Vicksburg on the Mississippi; and many of the genera, as
well as the great majority of the species, are extinct. Amongst
the extinct genera of shells, Adwria amongst the Nautili may be
mentioned as conspicuous; and amongst Fishes, Carcharodon
may be mentioned as an abundant Upper Eocene and Miocene
genus of Sharks, not more than one species of which is found
in our present seas, represented by the two best-known and
most widely distributed HEocene and Miocene species found
abundantly in such strata in England, Germany, and other parts
of continental Europe, and in North America, namely, the
Carcharodon megalodon (Ag.), specimens of which occur in our
Spring-Creek beds (though not very commonly) perfectly iden-
tical with those from Malta or England, or the supposed Hocene
beds of South Carolina, or the Miocene beds of Virginia and
Maryland,—and the Carcharodon angustidens (Ag.), which occurs
abundantly in our Australian beds so perfectly identical with
specimens from the Lower Miocene of Doberg near Biinde, that,
114 Occurrence of recent Shells in the fossil state near Melbourne.
when compared side by side, it is impossible to distinguish them
by the slightest difference; and as this species, according to
Prof. Agassiz’s recently published opinion, includes the Sheppey
London-Clay C. Toliapicus amongst other varieties, I need only
say that the best-marked Kocene and Miocene varieties found in
Europe and America are perfectly represented, on comparison
of specimens, by the different varieties in our Bird-Rock Bluff
beds.
It is, I think, a very curious result of the careful comparison
I have made between the fossil species of the Bird-Rock Bluff
Mollusea and their nearest allies, that I can with certainty an-
nounce one of the commonest of them to be specifically identical
with the Limopsis Belcheri of Adams and Reeve, of which the
few known specimens were brought up alive from a prodigious
depth by Admiral Belcher, off the Cape of Good Hope. The
identification, I should say, does not rest on an examination of
the published figure and description, which would not have been
sufficient for the purpose, but, having been fortunate enough to
procure a living specimen for the National Museum which I
take a pleasure in forming during my residence in Melbourne,
I have perfectly satisfied myself of the complete identity of our
Miocene Tertiary abundant shell with the hitherto very rare
recent one by direct comparison.
The Limopsis aurita (Sassi), perfectly identical with specimens
which I have used for comparison from the Coralline Crag
of Suffolk and many Miocene localities in Germany (which
Mr. Jeffreys has lately dredged from 85 fathoms off Unst, in
Shetland), is also common, though not so abundant in the
Australian beds as the L. Belchert.
The third living species of the Arcide in these beds is the
Pectunculus laticostatus (Quoy & Gaimard) of New Zealand,
occurring just as abundantly as the others.
The last bivalve I shall mention in this communication is an
extremely abundant Corbula, which I can safely pronounce
identical with the C. sulcata now living on the west coast of
Africa. Lest it might be supposed that, judging from figures
or descriptions, I had mistaken the North-east Australian Cor-
bula tunicata or other allied forms for the C. sulcata, I should
state that this is not so, but that, in working out the paleonto-
logy of our Australian deposits, I have thought it my duty to
science to take the precaution of procuring every recent species
I refer to for comparison before assuming an identity.
The commonest Dentalium in these beds I believe to be a
mere variety of the Upper Eocene D. Mississippiensis (Conrad)
from Vicksburg.
Melbourne, May 25, 1865.
Prof. G. Gulliver on Raphides and other Crystals in Plants. 115
XII.— Observations on Raphides and other Crystals in Plants.
By Grorce Guuiiver, F.R.S.
[Continued from vol. xv. p. 458.]
THE object of this paper being to show how the order Vitacez
differs, in the possession of the character of raphis-bearing, from
its alles, they will here follow as enumerated by Prof. Lindley.
Droseracee.—Dried specimens of Drosera rotundifolia and D.
anglica: no raphides.
Fumariacee.—Of these were examined fresh plants of Fumaria
officinalis and another English species, Dielytra spectabilis, and
Corydalis, sp., i none of which could raphides be detected.
Berberidacee.—Leaves and fruit of Berberis vulgaris, B. Japo-
nica, B. Darwin, B. dulcis, B. aquifolia, and leaves and ovaries
of Epimedium alpinum, E. macranthum: no raphides in any of
them ; a few spheraphides in the fruit of Berberis vulgaris.
Vitacee.—Raphides and other crystals in the Grape-vine
have been long known (Edwin Quekett, Lindley’s ‘ Introduction
to Botany’); and | have already- indicated that the character may
pervade the whole order (‘ Annals,’ Dec. 1863 and Jan. 1865).
Lately I have repeated my former observations, and extended them
to more species of this order and its allies. The Vitacez exa-
mined are Cissus discolor, Vitis vinifera, V.odoratissima, V. apii-
folia, Ampelopsis hederacea and two other species, and two species
of Leea. Every one of these plants afforded raphides and sphe-
raphides in more or less abundance. The Leee, though merely
old dried fragments of leaves and flowers, exhibited the raphides
and spheraphides in abundance, the raphides often in bundles,
and still more frequently swimming separately in the water on
the object-plate. All the other Vitaceze were fresh and healthy
plants. In Cissus, the sprigs, tendrils, young leaves, and stipules
all abound in raphides, some within short oval-shaped cells;
there were also other cells, longer, much tougher, and narrower
than the former, pointed or nipple-shaped at the ends, and con-
taining raphis-like objects. Whether these be true raphides
requires further exammation to determine; for they are very
fine and fragile, and (unlike the obvious raphides of this plant)
difficult to separate from each other and from their cells.
They are common, with the regular raphidian cells, in the leaves,
and especially plentiful in the thick base of the stipules.
Pittosporacee.—F resh leaves and twigs of Pittosporum undu-
latum and P. tobira: some spheeraphides in the leaves and meso-
phloeum, but no raphides. Dried fragments of leaves and flowers
of Bursaria spinosa, Marianthus candidus, M. sp., and Chetran-
thera linearis: a few spheraphides in each of these plants, but
no raphides. Fresh leaves of Sollya heterophylla: no raphides,
116 Prof. G. Gulliver on Raphides and other Crystals in Plants.
but many spheraphides. In short, these Pittosporaceze afford
spheeraphides, but are quite devoid of raphides.
Olacacee.—Dried leaves of Olax scandens, O. stricta, Liri-
osma, sp., Heisteria cyanocarpa, Ximenia americana, Icacina sene-
galensis, Aphodytes, sp., Gomphandra axillaris, Pogopetalum acu-
leatum, and Cansjera scandens: all these Olacaceze and Icaci-
nacez devoid of raphides.
Araliacee and Rhamnacea.—Of these orders the following
lants were examined, and none of them afforded any raphides :
Aralia leptophylla, A. nudicaulis, Hedera Helix, Rhamnus Ala-
ternus, Ceanothus azureus, and C. divaricatus. Some of them
abound in spheeraphides, as may be well scen in Aralia (‘Annals,’
April 1864) and Rhamnus. In the last plant they form a beau-
tiful spheraphid tissue, of which there is a plate from Lythrum
in the ‘Annals’ for September 1863, pl. IV. fig. 13. This
tissue occurs in the leaves, liber, and between the medullary
rays and alburnum of Rhamnus.
On the present occasion negative results of searches for ra-
phides are detailed more particularly than has been usual in
these papers, in order that botanists may estimate the observa-
tions on Vitaceee at their true value, and more especially as
Mr. W. H. Baxter has kindly afforded me the means of making
comparative examinations of all the above-named Leez, Pitto-
sporacee, and Olacacez.
Excepting the little order Cyrillaceze, of which I have yet seen
no member, the first six orders in this paper form the whole of
Prof. Lindley’s Berberal Alhance, in which the order Vitacez.
occupies the central place lneally. The affinities of this order
he thus indicates :—
Araliacee.
Berberidaceze.— Virace &.—Pittosporacee.
Rhamnacee.
The result of the present observations is remarkable. No
lant of the central order examined without finding raphides ;
while, on the contrary, these were never found at all in any
examination of its allies and surrounding orders. Thus Vitaceze
must surely be entitled to the character of a raphis-bearing
order. But whether this character will always certainly prove
diagnostic, as now seems probable, can only be decided after a
complete examination of all the orders in question. So novel is
this subject of raphides as natural characters in systematic
botany.
Balsaminacee, Galiacea, Onagracee, Phytolaccaceea, and Nye-
taginacee.—And the same remark applies to these raphis-bearing
Exogens, although my observations in the ‘Annals’ for July
Dr. O. A. L. Morch on the Operculum and its Mantle. 117
1864, and many since made, have convinced me that, so far as
regards the British flora, the raphidian diagnosis is not only
quite true, but very natural. Yet, as formerly noticed, this
character might more easily escape attention in Galhiacez than
in the other orders.
Finally, the propriety of retaming Leea under Vitacez has
been disputed ; and the present observations will tend to support
the conclusion of those botanists who, with Adrien de Jussieu
and Lindley, persist that this genus ought not to be separated
from Vitacez.
Edenbridge, July 17, 1865.
(To be continued. }
XIL1.—On the Operculum and its Mantle (lobus operculigerus,
pomatochlamys). By Dr. O. A. L. Morcu.
Apanson* regarded the operculum of univalve shells as an-
swering to the second valve of the bivalves—an opinion main-
tained by Oken and lately by Dr. Gray} and Prof. Macdonald ft.
In this point of view the lobus operculigerus (Lovén), or “ the
opercular mantle,” would correspond with one moiety of the
mantle of bivalves.
Prof. Lovén regards the bivalve shell as produced by a cloven
or bipartite mantle, and the operculum as homologous with the
byssus.
Prof. Keferstein§ supports Lovén’s opinion, considering the
slit in Emarginula and Tenagodus as a trace of division. The
porous slit of Haliotis, Tenagodus, &c., corresponds with the
notch or channel in canaliferous shells (Hntostomata, Blv.).
There is, however, a more important trace of division in many
univalves—for instance, the dentated furrow in Monoceros,
Pseudoliva, Ancillaria, and some species of Murex (Cerastes), but
chiefly in Carinaria. In this last genus the keel is formed by
the two sides of the shell, which are pressed against each other
in such manner that a piece of paper can be introduced into
the middle of the keel as far as the foetal shell. In Onustus
(Humphr.) the two sides are cemented together, but the union
can be clearly seen, Akera bullata shows something similar in
* Hist. Naturelle du Sénégal.
+ J. E. Gray “On the Operculum of Gasteropodous Mollusca, and an
attempt to prove that it is homologous or identical with the second valve
of Conchifera”’ (Annals and Mag. of Nat. Hist, ser. 2. v. p. 476; and
Phil. Trans. 1833).
t “On the Homologies of the so-called univalve shell and its Opercu-
lum ” (Proe. Linn. Soe. v. 1860).
§ Bronn u. Keferstein, Die Klassen u. Ordnungen des Thierreichs.
118 Dr.O. A. L. Mérch on the Operculum and its Mant’e.
the line of suture. The shell of the young Dentalium is also
split throughout its whole length. The best proof of analogy
would perhaps be the carapace of Limnadia and Estheria, among
Crustacea, which is bivalve, while that of closely allied genera,
as Nebalia and Apus, is univalve.
Pinna saccata, L., has both valves united in the adult state ;
but it has never been observed that the two valves have their
origin in the division of a single shell: on the contrary, the
division is manifest in the larval shell.
Nearly all organs are double in the Acephala: there are thus
two ovaria with distinct external orifices, two kidneys (organs
of Bojanus) with distinct external orifices, two pairs of labial
palpi, two pairs of gills. It seems to me therefore probable that
the Acephala also have two shells originating in the same way
as the other organs above mentioned. This duplicity of the or-
gans is very indistinct among the univalves, as in Dentalium and
Chiton, and it becomes rarer and rarer among the higher
Mollusca.
The larva (Glochidium) of Anodonta has in each shell a distinct
byssus-bundle (“ cordons ombilicaux,” Quatrefages*), and a dis-
tinct intestinal channel with distinct oral orifices+; in other
words, it is a true Diplozoon in the larval state. This curious
fact is perhaps not quite solitary among Mollusca. Thus, ac-
cording to Koren and Danielssen {, several eggs (from 1 to 100)
are, in Buccinum undatum, united to form a single embryo. The
difference is chiefly that in the much lower mollusk, Anodonta,
the amalgamation takes place in a more advanced state of the
embryos, so that some organs, the intestinal channel and the
byssus, are united into one, and the other organs are kept in
their original condition. The animal would then be composed of
two “zonites,” reminding us of the “egg-producing process”
of Hydra, regarded by Prof. Huxley as a reduced individual,
or an organ homologous with an individual §. An Acephalous
mollusk must therefore be considered an individual in the same
sense as a plant or flower composed of individuals (leaves) reduced
to organs. The question is, Does the opercular lobe with its
operculum represent one lobe of the mantle and its shell in the
bivalves, or is it something different ?
The epipodial line of Huxley (‘ manteau inférieur,” Lacaze-
* Ann. des Se. Nat. tom. iv. (1835) p. 283, and tom. v. (1836) p. 321,
&e.
+ Von Siebold, Vergleichende Anatomie, Wirbellose Thiere, p. 294.
+ Bidrag til Pectinibranchiernes Udviklingshistorie ; ‘‘ On the Develop-
ment of Buccinum undatum”’ (Athenzeum, 1852, p. 1066).
§ Lecture upon Animal Individuality, Royal Institution (Ann. Nat.
Hist. ser. 2. vol. ix. p. 505).
Dr.O. A. L. Morch on the Operculum and its Mantle. 119
Duthiers*) often produces posteriorly a shell (operculum) analo-
gous to the shell of the true mantle; daterally it can be deve-
loped into fins, as in Aplysia and Gymnosomata (Pneumodermon,
&c.), analogous to the pallial fins of the Cephalopoda; it can
form a fimbriated or undulated edge, as in Haliotis, Trochide,
Elysiide, and Philine+; it can form a siphon, as in Cephalopoda
or in the American Ampullaria, analogous to the mantle-siphon
of Buccinide ; perhaps it forms anteriorly the tentacula, ommato-
phores{, and intertentacular lobes of Trochide, corresponding to
the mantle-edge of Pectines, Solenes, Galeomma, &c. The epi-
podium is attached to the foot; but it is not quite clear that it
is a part of it. One author considers the foot homologous with
the adductor muscle of the Acephala ; but it must be remembered
that the foot of the Acephala is homologous with that of the
Gasteropoda. Dr. Gray regards the muscles which connect the
columella and the operculum as homologous with the adductor
muscles of bivalves.
Prof. Lovén considers the operculum homologous with the
byssus; but, as this organ is found in several univalves, even in
those with an operculum, this opinion cannot be adopted, as was
pointed out by Prof. Macdonald. It must also be remembered that
it is not known how the byssus of univalves is formed. Swain-
son (Treatise, p. 186, f. 29) represents Cyclostoma suspensum,
Gould, and A. Adams (Voyage of Samarang, p. 44, t. 13. f. 3),
Cerithidea obtusa, Lam., as attached to a branch by a byssus
during the zstivation. According to Macdonald, Planazis, and
to Gray, Rissoa parva, spin a byssus. It is possible, from its re-
semblance to that of Mytilus, that the deep posterior groove in
the footsole of Cerithiopsis tubercularis (Forb. & Hanl. Brit.
Moll.) produces the byssus. The nature of the slimy thread of
Litiopa, too, is very doubtful ; perhaps it only corresponds to the
thread of Limazx filans, Hoy. The “float” of Ianthina, which
attaches the animal to the surface of the water, is probably ho-
mologous with the byssus §, judging from its ventral position ||.
* “ Mémoire sur le Systeme nerveux de |’Haliotide”’ (Ann. des Se. Nat.
Zoolog. 1858, sér. 4. vol. xii. p. 226).
+ Morch, Journal de Conchyliologie, 1863, p. 39.
{ The double-eyed monstrosities of Hmarginula and Patella vulgata,
the latter of which, with a double tentacle, described by Fischer, are not
without importance for this comparison (Journal de Conchyhiol. tom. y.
p- 230, tom. xil. p.89). Lacaze-Duthiers, ‘Sur les Monstres doubles de
la Bullea aperta” (Compt. Rend. Acad. Se. tome xli. 1855, pp. 1247-
1250).
§ Morch, Journal de Conchyliologie, 1860, Juiliet.
Macdonald, ‘*On the Homologies, &c.”? (Proc. Linn. Soe. vol. y.
Nov. 14, 1860, p. 209).
| In the young Cyclas the byssus has, however, a posterior position.
120 Mr. J. Hogg on some Amphibians.
It has a remarkable analogy to the singular vesicular develop-
ment of the cement-tissue of the peduncle of Lepas (Dosima)
fascicularis, Sol. & Ellis*.
The byssus in the Acephala is generally corneous; but in
Anomia it forms a calcareous plate (the plug), possibly corre-
sponding with the opercular valve in Hipponyx and Lithedaphus,
which may be considered a calcareous secretion of the ventral
face of the foot. The epiphragm of the Helices would also
be homologous, if this plate be really a secretion of the foot,
as M. P. Fischer states; but it is probably secreted by the
mantle, like the septa of Vermeti, Runcina decollata, &ce. To this
category belong probably the tubes of Teredo, Gastrochena,
Clavagella, &c., and the accessorial valves of Pholades. The two
pallets in Teredo, which have a striking analogy to the opercula
of some Serpule (Hydroides norvegica, Gunn.), might perhaps be
compared with the posterior supplementary shells of Talona.
The shell of Argonauta, considered by Mr. Adams to be homo-
logous with the egg-cases of Murex, agrees with Nautilus in its
position and the black colour of the carina; but it seems to be
formed by the arms only. Its homology is therefore uncertain.
It appears that all parts of the skin in Mollusca can secrete a
shell. There are likewise found calcareous spicula or grains in
all parts of the body, in the clypeus in Gymnobranchia, the
tentacula of Pleurobranchus, and even in the intestinal channel.
In the Bullide and some Pellibranchiata there are thick calea-
reous plates in the stomach.
Note. The ligament is a thickening of the epidermis, which
is part of the skin of the animal, but not specially of the shell.
This seems evident to me from examining, for instance, a specimen
of Mya truncata in spirit. The connexion of the two valves by
the ligament proves, therefore, not that the valves were originally
one only, but that the bivalve shell is formed in the same manner
as the two lateral mandibles of the olde.
XIV.—WNotes on some Amphibians.
By Joun Hoee, M.A,, F.R.S., F.L.S. &c.
Dr. J. E. Gray, in his paper “ On the Clawed Toads (Dactyle-
thra) of Africa,’ published in the ‘Annals and Magazine of
Natural History’ (vol. xv. p. 334), well observes, that this
kind has “ large webbed hinder feet, some of the toes of which
are armed with very distinct horny black claws—a_ peculiarity
of structure that is quite an exception amongst the Batrachian
animals.”
* Darwin’s ‘ Cirripeds,’ p. 96.
Mr. J. Hogg on some Amphibians. 121
Cuvier, in the second edition of his ‘ Régne Animal’ (1829),
bestowed the generic name of Dactylethra on the only one then
known, which had been discovered in South Africa, and which
is now called D. capensis.
The Greek appellation of the genus, daxTvAjOpa, properly
means a “case” or “sheath for the finger,” 7. e. a thimble ; and
it is clearly a very correct one for the sort of horny case which
covers three of the five toes of this curious animal. Dr. Gray
describes it as a “black horny claw, which covers the last joint
of the three outer toes and the spur of the hind foot.”
The same zoologist further describes this Toad as having its
skin “ scattered with small white lines disposed in a symmetrical
manner, which, when examined by a magnifier of rather high
power, display linear series of close minute perforations or
glandular openings.”
These small perforations or pores are probably of use in
exuding, under a dry and hot atmosphere, a fluid that is service-
able in moistening the naked skin, which, in several species of
Frog, is known to perform the function of breathing. This
cutaneous respiration possessed by some of the Amphibians was,
I believe, first made known by Dr. Edwards, in Paris, more than
a quarter of a century ago; but how far that function may
assist, or be employed in lieu of, pulmonary respiration I have
not been able to learn.
When I wrote my first paper on the ‘“ Classifications of the
Amphibia,” which was published in the ‘ Magazine of Natural
History’ (n.s. vol. i. p. 265, 1839), I kept the genus Dactylethra
apart from the genus Pipa, and took the D. capensis as the type
of a distinct family, which I termed Dactylethride. For so
doing, more than twenty-seven years since, several zoologists,
whose classifications were not in accordance with mine, censured
me; but I am now very happy to find that Dr. Gray has
adopted (p. 340) the family “ Dactylethridz ” as an established
one. Although this distinguished naturalist does not assign the
author to this family, yet by consulting Prof. Agassiz’s ‘ Nomen-
clator Zoologicus,’ it will be seen that I was the originator of it.
The entries in that useful work are as follow :—
In the ‘Index Universalis’ (p. 115), “ Dactylethride, Hogg,
Rept. Ad. 1838.”
Again, in the ‘Addenda’ to ‘Reptilia’ (p. 3), ‘ Dactyle-
thride, Hogg, Ann. Nat. Hist. 1. 1838. Dactylethra. Pipe.”
And should the animal named by Dr. Gray Silurana prove a
distinct genus, and not the larval or tadpole-state of a species
of Dactylethra, it will constitute another interesting genus in
the family Dactylethride.
Ann. & Mag. N. Hist. Ser.3. Vol. xvi. 9
122 Mr. J. Hogg on some Amphibians.
I will now make a few remarks on the Axolotl of Mexico—
named by Cuvier Avolotus Meaicanus, and by myself Siredon
pisciformis (1838).
Dr. Gray, in his very useful ‘ Catalogue of the Specimens of
Amphibia in the British Museum,’ part 2, printed in 1850
(wherein he has done my labours justice), places this remarkable
Amphibian in his suborder II. Gradientia, and family III. Ple-
thodontide ; and he says at p. 49, that it “has only been observed
in its larva state.” He also there cites this passage from Baird
(Journ. A. N.S. Phil. 1849, p. 292) :—It (Siredon) so much
resembles the aa va of Ambystoma punctata in both external form
and internal structure, that I cannot but believe it to be the
larva of some gigantic species of this genus. It differs from all
known Perennibranchiates (the Manentibr anchians, mihi) im
possessing the larval character of the gular or opercular flap,
this being unattached to the subjacent integuments, and free to
the extremity of the chin. The non-discovery of the adult is
no argument against its existence.”
Also Charles Bonaparte, Prince of Musignano, in the same
year (1850), in his Classification of the Amphibia, considered the
Stredon as the mere tadpole of a Salamandra or Batrachian.
Although Dr. Gray, with his usual accuracy, has referred to
“ Home, Phil. Trans. 1824,” yet he seems not to have full
examined that memoir, and to have overlooked the following
passage which I wrote in 1838 respecting it :—
“TLatreille places the Axolotl (Siredon pisciformis) amongst
the Caducibranchious Amphibia; but it had been previously
discovered that its branchi@ are persistent; the details of which
may be learnt from a paper by Sir Everard Home, published in
the ‘Philosophical Transactions’ for the year 1824, p. 419.
One of the accompanying plates accurately represents the ex-
ternal gills as still remaining on a female Axolotl when in the
state of possessing fully developed ovaria, and just before the
ova are shed ; thereby proving her to be a perfect animal. Con-
sequently Latreille should have stationed the Axolotl next to
the Proteus in his second order.” This fact has been fully con-
firmed ; and the permanency of the external gi//s throughout
the life of the animal is now well determined. It is frequent in
the lake near the city of Mexico; and the common people con-
sidering it a fish (as indeed some naturalists are inclined to do),
sell it as such; and, as Hernandez says, “salubre et gratum
preebet alimentum.” There seem to me to be three or four
species which are not yet correctly known or distinguished.
M. Duméril has very recently given an account of the hatch-
ing of the young from the ova of the Mexican species (Szredon
pisciformis) in the menagerie of the Museum of Natural History
Mr. J. Hogg on some Amphibians. 123
in Paris. The ovum is like that of all the Batrachians. The
gills in the tadpole of this species consist of three short appen-
dages, which are cylindric and only shghtly ramified.
For a full description of the interesting development of the
tadpole from the egg of this Amphibian, see the last April
Number (16) of the ‘Comptes Rendus,’ tome Ix. p. 765.
I may, however, note that it appears that the time required
for the hatching of the tadpole of the Axolotl is about one month
—the same as that, in our ordinary springs, for the birth of the
common tadpole.
This genus was placed, in my modified Branchial Classifica-
tion, in 1841, thus :—
Class IV. AMPHIBIA.
Subclass IT. Diplopneumena.
Order IIL. MANENTIBRANCHIA.
Tribe I. RAMIBRANCHIA.
Family II. Proteide.
Genus SIREDON.
And I do not think it necessary for me to alter its position, even
after a period of twenty-four years.
But, before I conclude, I must point out two errors in Prof.
Agassiz’s valuable ‘ Index Universalis,’
The first is in attributing to me the word Cadnabranchia,
which I have never used. The entry at p. 56 stands thus :—
* Cadnabranchia, Hogg, Rept. Ad. 1838” (which he corrects
to “ Caducibranchia).” And he then inserts the following :—
“ Caducibranchia, Bonap. Rept. 1831” (which he corrects to
“V. cadnabranchia”’).
The word Cadnabranchia is, I conclude, only a misprint.
And the second error occurs at p. 310, as follows :—
“ Proteidea, Hogg, Rept. Ad. 1841” (which he corrects to
“ Proteoide’’).
Now the term “ Proteidea,” which is seen at p. 3855, Ann. &
Mag. Nat. Hist.’ (No. 45, July 1841), is not my own, but it is
that of Prof. J. Muller: the original is published in Oken’s
‘Tsis’ (p. 710) for the year 1831; and a translation from the
German, made by myself, is there inserted.
Norton House, Stockton-on-Tees.
July 11, 1865.
Ox
124 Prof..W. King on the Histology of
XV.—Remarks on the Histology of two Specimens of Rhynchopora
Geinitziana, De Verneuil, from near the River Oukhta, Pro-
vince of Archangel, and belonging to the Collection of the Corps
des Mines of St. Petersburg. By Professor Witt1am Kine.
Tne specimens which are treated of in the present communica-
tion have been recently noticed by Dr. Carpenter* in connexion
with a dispute which occurred between us in 1856. They were
kindly submitted to our individual examination, at the time, by
Mr. Davidson, who, on account of the light they seemed to
throw on the histology of the Rhynchonellide, spared no labour
in procuring the loan of them for the purpose stated.
Paleontologists will remember that, in a paper of mine, entitled
“Notes on Permian Fossils,” which appeared in the ‘ Annals
and Magazine of Natural History’ for March and April 1856,
I asserted that the species to which the specimens belong is “as
distinctly and regularly perforated” as any of the family Tere-
bratulide ; and that, on the contrary, Dr. Carpenter, in a letter
published nearly twelve months afterwards, maintained that it
has “only pits upon the inner surface of the shell’’+.
Before proceeding further, I wish it to be understood that I
have no desire to enter on a controversy with Dr. Carpenter in
your pages: my object is solely to place on record, in a scientific
journal like the ‘Annals,’ the observations which I made on the
specimens in question {. Dr. Carpenter, I am fully aware, will
also publish his observations on the same specimens—a proceed-
ing which I shall be perfectly satisfied with, provided he confine
himself to the main point at issue between us§. Should this
be the case, it may be reasonably expected that the discussion
will stimulate others, who have sufficiently favourable opportu-
nities, to endeavour to determine which of us is right.
One of the specimens has a considerable portion of the um-
bone of the large valve broken off: the other is nearly perfect.
* See ‘ Reader’ for July 8, p. 45.
+ See ‘Annals and Magazine of Natural History,’ ser. 2. vol. xix. p. 214.
Dr. Carpenter, however, had previously admitted, in a paper published in
the ‘ Annals,’ vol. xvii. p. 504, that my statement, as regards some German
specimens, was correct, “so far as can be judged by the external appear-
ance”’ of two which he had examined.
{ My observations would have been published at the time they were
made but for certain reasons, which it would be a waste of your valuable
space to be occupied with: they will be stated, however, I expect, in an
immediate number of the ‘ Reader.’
§ Dr. Carpenter will have ample opportunity of adding any other
matters im the ‘ Reader,’ in which he has already introduced a “ personal
discussion.”
two Specimens of Rhynchopora Geinitziana. 125
Neither specimen exhibits any portions of an impression of the
inner surface of the valves.
1. The specimen with the umbone of the large valve broken off.—
The outer or original surface of this specimen seems to have
disappeared, while in some places it is obscured with foreign
matter. In the former condition, the remaining test displays
numerous black dots*. In a few places the specimen has been
a little filed or cut down. The test that remains is, in several
places, of considerable thickness, which is well exhibited on the
fractured portion, where the umbone is broken off. At this part
the black dots are seen (as represented in the surface-figure and
longitudinal section, figs. 1 and 2) not only on the surfaces a
and c, but also on the intermediate por- Fic. 1
tion 6, A broken part on the small Pitas
valve, between the umbone and the an-
terior margin, also shows the test to be
of considerable thickness ; and in certain
lights some of the black dots appeared as
if they formed the terminations of fine
hair-like lines passing down through its Rie 2:
thickness. In no instance could I con-
vince myself that they were “ mere acci-
dental results of infiltration,’ as main-
tained by Dr. Carpenter in one of his
letters to me: their regularity completely forbids the idea.
Black dots are nearly everywhere exhibited on the abraded
surface of this valve: the exceptional cases will be noticed here-
after.
The observations stated completely prove, in my opinion, that
the black dots are not simply “ pits upon the internal surface of
the shell;” or they ought not to be seen on the surface of every
layer of tissue exposed: the whole of the layers, be it understood,
form a series of considerable thickness. On the contrary, seen
under the circumstances named, they cannot but represent
broken transverse sections of tubular perforations, similar to
those characteristic of the Terebratulide, and passing through
the entire thickness of their shell-substance. I am disposed to
regard their dark colour as due to the carbonaceous residuum
of the membrane with which the perforations were originally
occupied. The perforations of fossil Terebratulide are often
filled with a similarly coloured matter: this is remarkably the
case with some specimens before me of Waldheimia ornitho-
cephala, which in this respect offer so striking an analogy to
what is seen in the Permian species under consideration as to
* My observations, made with a Stanhope lens, were most successful
when made in subdued sunlight.
126 Prof. W. King on the Histology of
render it a matter of surprise how its perforated character can
be doubted.
From the surface of the large valve, a little in front, and on
the left of the part where the umbone is broken, some shreds
have been removed, apparently for microscopic examination.
This portion of the specimen appears to be so much altered by
mineralization as to render it, in my opinion, of /ittle or no ser-
vice in settling the question at issue. On those parts from which
the shreds have been removed the black dots are distinctly
visible; and the remaining test on which they are seen is of
such a thickness as to completely preclude the idea of their being
“only pits upon the internal surface of the shell.”
2. The nearly perfect specimen.—This specimen is exceedingly
valuable, inasmuch as it decisively exhibits, on the left side and on
the umbone of the large valve, the original or outer surface of the
shell, All the other parts are Rie. 3.
either obscured with foreign 2
matter, or they show that por-
tions of the shell-substance
have been accidentally worn
down or otherwise removed.
The black dots are distinctly
seen where the abrasion oc-
curs; and in two or three
places on the umbonal region
of the small valve they are
visible on at least three dif-
ferent layers, a, b, c, as repre-
sented in fig. 3. The parts
marked with an asterisk con-
sist of a mere crust of mineral
matter, which appears to have
replaced the original surface-
layer. A portion of the nght
half of the anterior region of
the shell, where the margins of
the valves join, also exhibits
black dots very distinctly, as represented in fig. 4: the test is
evidently of considerable thickness at this part.
With respect to the umbone of the large valve, it does not, to
me, show any black dots, either on the original surface or in
the tissue beneath it. Whether this absence is due to the shell-
substance of the umbone having undergone a molecular change
or metamorphism, or has been produced by any other cause,
are points on which I cannot offer any decided opinion*. I am
* To show the effects of fossilization, I may mention that the type
two Specimens of. Rhynchopora Geinitziana. 127
unwilling, however, to admit that the test is here in its original
condition, because the surface-layer on the left side of the valve,
passing on to its margin, displays a few faint black dots*; while
the markings are distinctly visible on the third rib, where a
small film-like portion of the original surface has been removed.
Figure 5 is a representation of what has just been described,—
a being the original surface, with a few Fig. 5.
faintly-marked black dots; and 4, the
abraded part, with others distinctly vi-
sible. This absence of what may now
be termed tube-apertures on the um-
bone is paralleled in many specimens
I have examined of Terebratulide be-
longing to the Carboniferous and Per-
mian systems. I have also observed it in
Jurassic specimens. In all these cases, mineralization was pro-
bably the obliterating agent. Furthermore the test at the
umbonal portion of the small valve of the first-described speci-
men does not exhibit the black dots so well as the anterior
region: their paucity on an old part of the shell is possibly due
to senility.
On the whole, I cannot but regard these two specimens as
completely confirming the view | took of the histology of the
German fossils noticed under the head of Rhynchonella Gei-
nitziana, in my paper already alluded to. The German speci-
mens, however, exhibit the tubular perforations more decidedly
than those from Russia, which is in favour of the latter having
undergone a greater amount of metamorphism. I have nothing
more to add to my description of the German examples: their
perforated structure, which is correctly represented in the plate
appended to my paper, is identical with that of a species of
Silurian Retzia, specimens of which are now before me. In the
latter the perforations are indicated by black dots, similar to
those in the German specimens.
I therefore repeat, in conclusion, that the species under con-
specimen of the species, belonging to my friend M. de Verneuil, who
kindly lent it me for examination, shows no black dots, nor any appearance
of perforations; but its test has undoubtedly undergone a remarkable
change, which has developed a concentric structure like that of Beekite, and
which,in my opinion, has obliterated all indications of perforations. Another
specimen, collected at Ropsen, near Gera, which I procured from Mr. Damon
of Weymouth, differs remarkably, im the apparent absence of perforations,
from some, procured at the same place, which I had noticed in the “* Notes”
already cited as published in the ‘Annals.’
* T exclude some accidental superficial black specks, which might be
confounded with the ‘ black dots ;” but the latter rather appear to be in
the outer layer of tissue than on it.
128 Royal Society :—
sideration has both valves “as distinctly and regularly perforated
as those of any Terebratulide.’ Hither Dr, Carpenter or I must
be labouring under some serious mistake. If the mistake be
mine, I shall readily bow to correction ; but | may be excused
maintaining my view until the appearances on which it 1s
founded are shown to support a contrary conclusion.
Belmont, near Galway, July 14, 1865.
PROCEEDINGS OF LEARNED SOCIETIES.
ROYAL SOCIETY.
June 15, 1865.—Major-General Sabine, President, in the Chair.
“A Description of some Fossil Plants, showing Structure, found
in the Lower Coal-seams of Lancashire and Vorkehires: By E. W.
Binney, F.R.S.
The author stated that, although great attention has been devoted
to the collection of the fossil remains of plants with which our coal-
fields abound, the specimens are generally in very fragmentary and
distorted conditions as they occur imbedded in the rocks in which
they are entombed ; but when they have been removed, cut into
shape, and trimmed, and are seen in cabinets, they are in a far worse
condition. This is as to their external forms and characters. When
we come to examine their internal structure, and ascertain their true
nature, we find still greater difficulties, from the rarity of specimens
displaying both the external form and the internal structure of the
original plant. It is often very difficult to decide which is the out-
side, different parts of the stem dividing and exposing varied sur-
- faces which have been described as distinct genera of plants.
The specimens described were collected by the author himself,
and taken out of the seams of coal, just as they occurred in the matrix
in which they were found imbedded, by his own hands. ‘This has
enabled him to speak with certainty as to the condition and locality
in which they were met with.
By the ingenuity of the late Mr. Nicol of Edinburgh, we were fur-
nished with a beautiful method of slicing specimens of fossil wood so
as to examine their internal structure. The late Mr. Witham, as-
sisted by Mr. Nicol, first applied this successfully, and his work on
the internal structure of fossil vegetables was published in 1833. In
describing his specimens, he notices one which he designated Ana-
bathra pulcherrima. This did not do much more than afford evi-
dence of the internal vascular cylinder arranged in radiating series,
somewhat similar to that described by Meer Lindley and. “Hutton
as occurring in Stigmaria ficoides, in the third volume of the ‘ Fossil
Flora.’
In 1839 M. Adolphe Brongniart published his truly valuable
memoir, ‘ Observations sur la structure intérieure du Siyillaria ele-
On Fossil Plants from the Coal of Lancashire, &c. 129
-gans comparée a celle des Lepidodendron et des Stigmaria et a celle
des végétaux vivants,’” in the Archives du Muséum d’ Histoire
Naturelle. His specimen of Sigil/aria elegans was in very perfect
preservation, and showed its external characters and internal struc-
ture in every portion except the pith and a broad part of the plant
intervening betwixt the internal and external radiating cylinders.
Up to this time nothing had been seen at all to be compared to
M. Brongniart’s specimen, and no person could have been better
selected to describe and illustrate it. His memoir will always be
considered one of the most valuable ever contributed on the fossil
flora of the Carboniferous period.
In 1849, August Joseph Corda published his ‘ Beitrage zur Flora
der Vorwelt,’ a work of great labour and research. Amongst his
numerous specimens, he describes and illustrates one of Diploxylon
cycadeoideum, which, although not to be compared to M. Bron-
gniart’s specimen, still affords us valuable information, confirming
some of that author’s views rather than affording much more original
information. All these last three specimens M. Brongniart, in his
‘Tableau de végétaux fossiles considérés sous le point de vue de leur
classification botanique et de leur distribution géologique’ (published
in 1847), classes as Dicotylédones gymnospermes, under the family
of Sigillarées—amongst other plants his Sigillaria elegans, Mr.
Witham’s Anabathra, and Corda’s Diploxylon.
In 1862 the author published, in the ‘Quarterly Journal of the
Geological Society’ of that year, an account of specimens which
confirmed the views of the three learned authors above named as
to Sigillaria and Diploxylon being allied plants ; but showed that
their supposed pith or central axis was not composed of cellular
tissue, but of different-sized vessels arranged without order, having
their sides barred by transverse strize like the internal vascular
cylinders of Stgillarta and Lepidodendron. These specimens were
in very perfect preservation, and showed the external as well as the
internal characters of the plants.
All the above specimens were of comparatively small size, with
the exception of that described by M. Corda, which, although it
showed the external characters in a decorticated state, did not
exhibit any outward resemblance to a plant allied to Sigil/aria with
large ribs and deep furrows so commonly met with in our coal-fields,
but rather to plants allied to Sigillaria elegans and Lepidodendron.
In the present communication the author has described some speci-
mens of larger size than those previously alluded to, and endeavoured
to show that the Sigillaria vascularis with rhomboidal scars gradu-
ally passes as it grows older into a ribbed and furrowed Sigil/aria, and
that this singular plant not only possesses two woody cylinders
arranged in radiating series, an internal and an external one divided
by a zone of cellular tissue, both increasing on their outsides at the
same time, but likewise has a central axis composed of hexagonal
vessels, arranged without order, having all their sides marked with
transverse strize. Evidence is also adduced to show that Sigillaria
dichotomizes in its branches something like Lepidodendron, and that,
130 Royal Society :—
like the latter plant, it has a Lepidostrobus for its fructification. The
outer cylinder in large Sigillarie is composed of thick-walled quadran-
gular tubes or utricles arranged in radiating series, and exhibiting
every appearance of the tree having been as hard-wooded as Pinites,
but as yet no disks or strize have been observed on the walls of the
tubes. Stigmaria is now so generally considered to be the root of
Sigillaria, that it is scarcely necessary to bring any further proof of
this proposition ; but specimens are described which prove by simi-
larity of structure that the former is the root of the latter.
The chief specimens described in the memoir are eight in number,
and were found in the lower divisions of the Lancashire and York-
shire coal-measures, imbedded in calcareous nodules occurring in
seams of coal.
No.1, Diploxylon cycadoideum, was from the first-named district,
and the same locality as the T'rigonocarpon, described by Dr. J. D.
Hooker, F.R.S., and the author, in a memoir on the structure of
certain limestone nodules enclosed in seams of bituminous coal, with
a description of some T'rigonocarpons contained therein*; and the
other seven (Sigillaria vascularis) were from the same seam of coal
in the lower coal-measures in which the specimens described in a paper
entitled ‘On some Fossil Plants showing Structure from the Lower
Coal-measures of Lancashire’ +, were met with, but from a different
locality in Yorkshire.
“On the Fossil Mammals of Australia.—Part II. Description of
an almost entire Skull of Thylacoleo carnifex, Ow.” By Professor
Owen, F.R.S. Xe.
In this Part the author gives additional cranial and dental charac-
ters of the extinct marsupial carnivore, Thylacoleo, deduced from
examination of better-preserved fossils, obtained from freshwater
deposits in Darling Downs, Queensland, Australia.
The fore part of the skull, wanting in the first-described specimen
from similar deposits in the province of Victoria, is preserved in
the present specimen, showing the premaxillary bones, which are
relatively larger than in placental felines. Each bone has three
teeth, of which the foremost is developed into a tusk, the second
and third being very small. There is no canine, or no tooth de-
veloped as a laniary in the maxillary bone. In the short extent
of the alveolar border of this bone between the great carnassial
molar and the maxillo-premaxillary suture, there are two approximate
small round sockets, which lodged either one double-rooted tooth or
two small single-rooted teeth. But dental development has mainly
expended itself upon the perfection of a pair of laniary incisor tusks,
in both upper and lower jaws, for piercing, tearing, and holding, and
a pair of carnassials in both jaws for flesh-cutting. These, in the
present specimen, closely agreed with those described in the former
one, but were more worn: they are the largest examples of these
* Philosophical Transactions, 1855, p. 149.
+ Quarterly Journal of the Geological Society of London for May 1862.
Prof. Owen on the Skull of Thylacoleo carnifex. 131
peculiarly modified shear-blade teeth in the mammalian class. Al-
though the tusks are incisors—not, as in placental carnivora, canines
—they possess, through the singular shortness of the facial part of :
the skull in Thylacoleo, the same mechanical advantage, in their
proximity to the biting-power of the enormously developed temporal
muscles, as in Felis. In the lower jaw there is, anterior to the car-
nassial, either a socket for a small double-rooted premolar, or two
approximate sockets for as many single-rooted ones; and, as in the
upper jaw, these cavities do not range in the same longitudinal line
with the carnassial, but extend obliquely inward and forward, from
the inner side of its fore part. There is no other alveolus in the
lower jaw between the premolar one and that of the large lower
tusk. The small ‘tubercular’ molar on the inner side of the hind
end of the upper carnassial, and the two ‘ tuberculars’ behind the
lower carnassial, are indicated by their sockets in the present speci-
men. The author sums up, from acquired data, the dental formula
: 3—3 : 2 aa
of Thylacoleo as follows :—Incisors ;—, Canines ;, Premolars —;
1—)?
2—2 - 1-1 1 o <
=o Carnassials —- Tuberculars —. Of the incisors, the fore-
or =
most above are long and large tusks, like the pair below: of the
other teeth, the carnassials, of unusually large size, are functioned as
flesh-cutters, and the small tuberculars would serve for pounding
gristle or tendon, as in Felis: the premolars indicated by sockets,
and the small upper incisors, represent a remnant of the dental
family type under its extreme adaptive modifications in T’hylacoleo.
In the rest of the skull of the subject of the present Part, many
particulars are yielded in addition to those deduced from the frag-
mentary fossils which indicate the genus. They confirm the deduc-
tions of the marsupial nature of the large extinct Australian carnivore,
determine the alternative expressed in the author’s first communica-
tion as to the homologies of the inferior tusks, and show that the
genus Thylacoleo ranges, not with the series now including Didel-
phys, Dasyurus, and Thylacynus, but with the Diprotodont group,
more eminently characteristic of the Australian continent, and which
is at present represented by, or reduced to, the genera Phascolarctos,
Phalangista with its subgenera, Macropus with its subgenera, and
Phascolomys. The carnassial of Thylacoleo, in its large propor-
tional size, absence of the tubercular part, and indications of subver-
tical groovings of the enamel, most closely resembles that tooth of
the more ancient marsupial carnivore Plagiaulaz, and is associated,
in the lower jaw, as in that genus, with two small posterior tuber-
culars, one or two small premolars, and one large incisive tusk, simi-
larly directed obliquely upward and forward. Few facts in mamma-
lian paleontology are more interesting and suggestive than the
occurrence in our hemisphere, during secondary geological periods, of
Marsupial forms, which find their nearest representatives in existing
or tertiary extinct Marsupialia of the continent of our Antipodes.
The present Part of the author’s series of Papers on Extinct Aus-
tralian Mammals is illustrated with drawings of the entire skull of
the Thylacoleo carnifex.
132
ZOOLOGICAL SOCIETY.
Feb. 28, 1865.—Dr. J. E. Gray, F.R.S., in the Chair.
DESCRIPTION OF A NEW SPECIES OF PoRPOISE IN THE MUSEUM
oF Buenos Ayres. By Dr. H. Burmeister, F.M.Z.S.
PuHocaNna SPINIPINNIS, Sp. nov.
The animal has the general figure of the common European spe-
cies, but differs entirely in the position of the dorsal fin, which is
placed further backwards, and has spines on the upper edge.
The whole body is black, without any other colour, and the sur-
face of the skin is transversely striated with fine excavated lines, like
the inside of the human hand. The upper lip is somewhat shorter
than the under, and the figure of the mouth, on both sides, rather
curved behind; the length of the opening is 84 centim. on each
side. From the hinder corner of the mouth the eye is distant
7 centim., and from the eye to the beginning of the pectoral fin is
16 centim. The opening of the nose has the form of a broad trans-
verse ridge, somewhat curved forwards; it is 3 centim. broad, and
16 centim. distant from the top of the upper lip. The figure of the
whole body is fusiform, but much more elongated behind than be-
fore; it measures from the top of the upper lip to the notch of the
tail-fin 162 centim., and the circumference of the thickest part of the
body, at the middle, is 102 centim.
The distance from the nasal aperture to the beginning of the dor-
sal fin is 84 centim. ; but the elevation of this fin is so gradual, that
it is difficult to say exactly where it begins. The figure of the whole
fin is triangular, somewhat curved forwards near the end, and its
height 14 centim. (see fig. 1). This curving forwards is a peculiar
and very distinguishing character of the species, as is also the
clothing of the anterior margin of the fin with small spmes. These
spines are not different from the skin, but elevations of the skin itself,
like small angles, of an elongated-oval form. I have figured part of
the middle (where the spines are most elevated) as seen from above
(see fig. 2), to show that every spine is surrounded by a ridge of the
skin, and that from the sides of the lateral spines other ridges begin.
Some small spines begin in the middle of the back, at the distance
Dr. H. Burmeister on a new Porpoise. 133
of 25 centim. in front of the fin, as a single line of moderate spines ;
but soon another line begins on each side, so that in the beginning
of the fin there are already three lines of spmes. These three lines
are continued over the whole rounded anterior margin of the fin,
and are augmented on both sides by other small spines irregularly
scattered, so that the whole number of spine-lines in the middle of
the fin is five. Towards the end of the fin they become smaller,
and on the rounded tip of the fin there are no spines at all.
From the hinder margin of the dorsal fin to the notch of the
tail-fin is 54 centim. ‘The tail-fin is 39 centim. broad, and each
fluke 20 centim. long on the anterior margin. This margin is some-
what curved backwards, and the hinder margin sinuated.
The underside of the body is somewhat more curved and extended
than the upper side, and the tail more descending.
The anus is situated under the beginning of the dorsal fin, 70 centim.
distant from the notch of the tail-fin.
The individual seems to be a very young one, because all vestiges
of genital organs are wanting in the exterior. The anus has a dozen
radial folds, of which the largest, 6 centim. long, runs forwards ; all
are very deep, and transversely ridged.
The pectoral fin is faleated, 26 centim. long and 10 broad. At
its proximal end there are many fine ridges in the skin, and in the
middle part are ridges indicating the finger-bones beneath.
The skull proves that the animal is a very young one, and that it
has come perhaps only to half its natural size; because all the bones are
very weak, not perfectly ossified, and the vomer entirely cartilaginous.
It has the general figure of the skull of the European Phocena, dit-
fering principally in the form of the hinder part of the intermaxil-
lary bones, which is more abruptly elevated in this new species than
in the European (see figs. 4 & 5).
Fig. 3. Fig. 4.
Side view of the skull of Phocena spinipinnis, reduced one-third.
134 Zoological Society :—
The upper jaw has sixteen small teeth, and the lower jaw seventeen,
on each side, there being no vestige of an alveolar ridge behind them
The first teeth are smaller and conical, the hinder
in either jaw.
broader and truncated, as seen in figures 3 & 4. ‘This is another cha-
racter distinguishing it from the European species; the skull of a
young individual of the latter, which I examined, had twenty-four
teeth in the upper jaw, and twenty-five in the lower, in both extending
more towards the hinder part of the jaw than in the new species.
The specimen of P. spinipinnis which is preserved in the public
Museum of Buenos Ayres, was captured in the mouth of the River
Fig. 5.
nN,
nV EA EN Bay
y any
Nay
Skull of Phocena spinipinnis, seen from above, reduced one-third.
Plata, and was afterwards exhibited in Buenos Ayres to the public,
some years before I came to this country.
Length of the whole skull, 29 centim.
Breadth between the orbits, 17 centim.
Length of the external margin of the upper jaw, 12 centim. ; of
the lower jaw, 22 centim.
Note.—The tympanic bone is lost ; the figure is therefore defective
in this part.
On THE OsTEoLocy oF MicroGiossa ALECTO.
By W. K. Parxer, F.Z.S.
Having been busy of late with the study of the skull and its
development in the Ostrich tribe, Iam the more sensitive to the
Indeed, but for
peculiar ornithic excellences of the Parrot family.
their Livery, it could hardly have been supposed that these opposite
Mr. W. K. Parker on the Osteology of Microglossa Alecto. 135
creatures belonged to one house: they are the most perfectly anti-
thetical of all the feathered tribes.
Judged by the mere power of flight, the Parrots would not be
accounted worthy to stand in so high a position; but this is only
one, among many, of the talents possessed by birds of noble degree.
Like all those who glory in “ high degree,’’ the Parrots have a
poor relation or two to abate their pride. The Owl-billed Parrot
(Strigops habroptilus) of New Zealand is as lowly as “‘ the younger
son of a younger brother.” If birds were to be classified by the
sternum only, then the Strzgops should be put near the Apteryz,
and the Tinamou attached to the train of the Peacock.
If birds be ranked according to the degree of their intelligence,
then, without controversy, the familiar Crows and Starlings, Finches,
and Singing-birds may take the highest room ; but if power of flight,
mere brute strength, and savage audacity shall be considered most
decent and becoming to a bird, then let the Eagles and Falcons sit
on the throne of the feathered kingdom. But there are qualities,
dear to the morphologist, in which the Parrots have the preemi-
nence, and stand higher, as Birds, than all other birds ; and although,
all things considered, the Crow is the best type and model with
which to compare the whole plumy brotherhood, yet in many things
the Parrot is a bird of birds; he is an ultra-type, and sets bounds to
the class to which he belongs.
But this bird, with the wise and solemn face of an Elephant, has,
like us, its chief and best qualities resident in its head; and if the
skull of an Ostrich be compared with that of the most psittacine of
the Parrots, the difference will appear almost as great as exists be-
tween a larva and an imago.
The type under consideration is one in which the characters of
the Parrot, and indeed the characters of a Bird, as such, are carried
to their highest pitch. I have long been familiar with this highest
kind of Psittacine skull in the genera Plyctolophus and Calypto-
rhynchus (see Cat. Mus. Coll. Surg. vol. 1. pp. 277, 278, nos. 1440
& 1445), and have recently discovered it in the Grass-Parakeet
(Melopsittacus undulatus); but the genus Microglossa carries it
to the fullest degree.
The teleologist might write a fair volume on the fitnesses dis-
played in the skull of this bird; but the adaptive conditions are of
secondary importance to him who would trace the clue of morpho-
logical unity through the mazes of nature’s unutterable variety.
The first thing that strikes the eye of the observer is the cleaving
of a great transverse cleft through the whole face, in front of the
eyes, leaving the enormously developed intermaxillary apparatus, en-
closing the vestibular parts of the olfactory organs, on one hand,
and the skull, maxillary apparatus, and true olfactory region, on the
other. Then we see that not only is the eye bounded beneath by a
blending of the lachrymal with the postfrontal, but the latter is an-
chylosed to the squamosal also ; and thus, with the true zygomatic
arch below, we have three pairs of facial bridges. But the deep,
steep-sided, beautifully arched intermaxillaries, the fair, broad fore-
136 Zoological Society :—
head, the well-roofed eyebrows, the perfectly bony orbit, and a man-
dible such as the eye searches for in vain elsewhere—all these are
outstanding characters in the highest type of Parrots, and, above all,
in the genus Microglossa.
The huge, mobile face is but one bone in the adult, and yet it is
composed of a great variety of parts that have become blended into
one thick mass, ; perfectly void of sutures. The nasals, intermaxil-
laries, prevomers (the vomer is not developed in the Psittacidze), the
nasal septum, the inferior turbinals, and the alee nasi, all these go
to form this large compound bone. ‘There are, therefore, six splint
bones; and the axial bones are four for the septum, two (at least)
for the inferior turbinals, and two for the alee nasi, thus making
eight more, or fourteen bones in all. The highly complex skull
also is completely fused into one bone, and it has in it the separate
parts that form the auditory and olfactory sense-capsules. But the
original attachment of the pieces of the arrested palato-pterygoid
arch is loosened so as to let the ascending (proximal or orbital) pro-
cess of the palatine lie half an inch below its proper foundation, viz.
the pars plana or antorbital. Anteriorly, the palatine is thick and
transversely expanded, and its convex elliptical end fits in a glenoid
cavity in the end of the prevomer of the same side. Further back,
at its proximal plate, it is two-thirds of an inch high, it scarcely
becomes less than half an inch ; and its emarginate hinder end reaches
to behind the ‘‘ membrana tympani,” full a quarter of an inch be-
hind the somewhat slender rod-like pterygoids. The latter bones,
although an inch in length, are thus completely overlapped by the
palatines. The sraall, late-appearing mesopterygoids have early coa-
lesced with each other, and they have united also with the front corner
of the basicranial edge of the left palatine. The malar bone articu-
lates, like its axis, the palatine, with the prevomer. The epiptery-
goid process of the pterygoid is obsolete ; the metapterygoid pro-
cess of the quadrate bone is small, conical, and anteriorly placed, as
in its autogenous counterpart in the non-venomous Serpents. The
hinge- convexity of the quadrate bone is semicircular; the cupped
process for the jugal is large and projecting ; and a well-developed,
outstanding, oval condyle i is received by the cup at the end of the
pterygoid. The heads of the os quadratum—answering to the crura of
our anvil-bone (‘‘incus’’)—are well developed, but do not stand as in
other birds ; for that which is related to the sympletic cartilage of
the stapes is directly ims¢de the outer or prootic head. In birds
generally, this incus-head projects far backwards, overlapping the
opisthotic, and overshadowing the auditory “ fenestra,” to articu-
late with the exoccipital. The splints of the lower jaw, ten in
number, have all become one piece, as unlike as possible to the
simple Meckelian rod on which they were modelled. The sym-
physis is an inch in extent, and the bone is transversely flattened
below, so as to be an inch wide at what should be the intermandi-
bular angle; this is, there, a gently concave transverse margin
having a rounded edge. The greatest height of the mandible is
1; inch; the angular process passes further back than the exocci-
Mr. W. K. Parker on the Ostevlogy of Microglossa Alecto. 137
pital. The occipital condyle is an extremely neat hemisphere. The
scooped occipital plane forms a very obtuse angle with the basis
cranii, which latter region is very small, triangular, and protected by
sharp ridges that meet at the fore angle of the coalesced basitem-
porals, below the small, closely placed Eustachian openings. At first
the “rostrum’’ of the basisphenoid is sharply~carinate, then it
becomes thick, rounded, and covered with articular cartilage, under
which the palatines and anterior ends of the pterygoids g vlide. The
height of the skull is so great that, although the hemispheres of the
brain lie down between the eyes more than in most birds, yet the
compressed rostrum of the basisphenoid and the lower edge of the
perpendicular ethmoid do, together, make a great keel, larger than
the: sternal keel of the Love-bird ( Agapornis ‘puller ia). The ante-
rior pterygoid processes are thrown out of relation to the pterygoids,
which grow no spur to answer to them; they are dull forthstanding
prickles. The exoccipitals are not nearly so much scooped to make
a drum-cavity as in the smaller Parrots; the tympanics, like the
columellie, are lost. ‘The main piece is large i in some of the smaller
kinds. In front of the great cranio- facial hinge, the nasals and
nasal processes of the intermaxillaries are converted into the merest
swollen sponge; behind the hinge, on each side, the lachrymals are
also swollen; but the frontals dip to form a valley between the or-
bits. Then there is a pair of frontal, and another pair of parietal,
smooth, large, rounded swellings, with a shallow, equally smooth
valley between them. The width of the head is nearly two inches at
the point where the postorbital process of the frontal melts into the
postorbital spur of the alisphenoid (post-frontal proper). Below and
behind this point it is more than two inches wide. The junction of
the thick quadrate splint (squamosal) with the post-frontal spur is
so extensive as almost to cover in the small heart-shaped “temporal
fossa.” This bridge of bone is half an inch across. The optic fora-
mina are about one-third of an inch apart; the olfactory fissures are
at the same distance. There is an elegant, small, shell-like middle
turbinal on the front of the self-dev eloped “pars plana,” or antor-
bital, and the simple crus of the ethmoid curls upon itself, so as to
form an upper turbinal. There are evidently full two coils to the in-
ferior turbinals, which are ossified in a fenestrate manner, as in mam-
mals, and which project far beneath the alee nasi. These latter are
ossified separately in the Parrots, and then, in many instances as in
this, acquire an adhesion with the nasals and the inferior turbinals.
The outstanding spurs of the antero-inferior septal bone increase the
complexity of the nasal labyrinth.
The sternum has its fenestree nearly filled up. The sternal keel
is, as in Parrots and many of their nearest allies, coincident with the
upturned, somewhat bifurcate episternal process. This is perfectly
normal ; for the keel, the episternal process, and the coracoid grooves
really belong to the shoulder-girdle ; together they form the true
episternum or manubrium. This might be called “ omo-sternum,”
in contradistinction to the rib-sternum (‘ pleuro-sternum’’), or that
which relates to the tnner cartilaginous belts, which grow directly
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. 10
138 Zoological Society :—
from the centra of the vertebrae *. ‘The furcular bone is only appa-
rently simple, although in this specimen of Microg/ossa no sutures
can be seen. In the Ash-coloured Parrot (Psittacus erythacus),
however, and in the East-Indian Pale@ornis torquatus, the thick,
broad end of each ramus is seen to be a separate piece. This is
also to be seen in the Toucan (Ramphastos Toco) and in the King-
fisher (A/cedo ispida), but is still better developed in all the ‘‘ Rap-
tores”’ and Totipalmatie, in the Baleniceps and Umbretta, and, in
a less degree, in most typical Herons. I have already spoken of
this part (P. Z. S. 1864, p. 339 ef seq.), and may now say that it is
a rudiment of the so-called ‘ clavicle’’ of the Batrachian, Chelonian,
and African Ostrich, and is well seen as a distinct bone in the
shoulder-girdle of the Salmon tribe and some other allied Fishes.
In Birds this rudiment is proximal; in Mammals, generally, it is
distal or sternal ; but I have found such a piece at both ends of the
clavicle in certain Insectivora, e. g. the Mole (Talpa europea), and
in the Shrew (Sorex tetragonurus). In Lizards the counterpart of
this cartilage is the anterior boundary of the coraco-acromial fenes-
tree. The supposed rudiment of the clavicle in certain small Par-
rots, e. g. the Love-Bird ( dgapornis pullaria) and the Grass- Parakeet
(Melopsittacus undulatus), is an ossification of this acromial car-
tilage. In Psephotis multicolor neither this nor the furcular bone is
present.
March 28, 1865.—John Gould, Esq., F.R.S., m the Chair.
Notice or A New Species or Porpotse (PHOCHNA TUBER-
CULIFERA) INHABITING THE Mouru oF THE THAMES. By
Dr. Joun Evwarp Gray, F.R.S., F.L.S., ere.
The fact of a new species of Porpoise being found on our own
shores, at the mouth of the Thames, must be considered a proof of
how little we at present know of the species of Cetacea.
The Zoological Society, who are so anxious to obtain specimens
of these animals that their habits may be studied, procured with
considerable trouble a fine male Porpoise, which had been caught at
Margate. It was carried to the Gardens, and placed in the pond
Formed for these animals; but, though showing no external injury,
it was in so weak a state een it arrived that it sank to the bottom,
and was obliged to be taken out and suspended by bands on the sur-
face of the water so that it might not be choked. After a time it
recovered so as to be able to swim about by its own exertion, but it
only survived the transport a few days.
Messrs. Bartlett and Gerrard, when it was alive, said that it dif-
fered so much in general appearance from the Common Porpoise that
they were induced to believe that it might be a species of Lageno-
rhynchus or Grampus.
* T would remark that, to trace the affinities of the Parrot tribe, we should
take such forms as the Common Grey Parrot (Psi¢tacus erythacus), Nestor,
Psephotis, &e., in which the Psittacine characters are somewhat enfeebled. I
have not found any other “family”? so isolated as this.
Dr. J. E. Gray on a new British Porpoise. 139
The general form of the head, and examination of the teeth after
death, proved at once that it was a species of Phocena, very nearly
allied to, if not identical with, Phocena communis.
Dr. Burmeister’s description of a Phocena from the River de la
Plata (contained in the Museum at Buenos Ayres), which is peculiar
for having some spines on the upper edge of the dorsal fin, naturally
made me careful in examining the edge of the fin of this specimen ;
and to my astonishment I discovered that this species also was pro-
vided with a series of compressed tubercles, giving the fin a sharp,
hard, serrated appearance.
The tubercles or spines on the dorsal fin having been observed in
two specimens from very different localities, I was induced to inquire
if this was a character common to the genus, which had been over-
looked ; but, on examining the stuffed specimen of the Common
English Porpoise in the Museum, it is clear that they are not found
in the common state of the species. It then occurred to me that
it might be a peculiarity of the male sex; but Mr. Flower informs
me that the male specimen which lived for some weeks in the Gar-
dens of the Society, and which he lately dissected, certainly had no
spines on the edge of the dorsal fin: so that cannot be the case.
Under these cireunistances I think I am justified in considering
that the possession of these spinous tubercles is a peculiarity of the
species, and therefore a specific character. The examination of the
skull shows that there are differences in its form which confirm
this opinion.
The species of Phocena may be thus defined :—
a. Back in front of the dorsal fin, and upper edge of the dorsal fin,
smooth, without tubercles or spines. Dorsal fin in the middle
of the back.
1. PHOoCHNA COMMUNIS.
Hab. North Sea and mouths of rivers.
b. Back in front of the dorsal fin smooth; the upper edge of the
dorsal fin with a single series of oblong compressed tubercles,
which are more crowded near the upper end of the fin. Dorsal
fin in the middle of the back.
2. PHOCEHENA TUBERCULIFERA, Sp. Noy.
Hab. Mouth of the Thames, Margate.
c. Back in front of the dorsal fin with a single series, and upper
surface of the dorsal fin with three series, of square-based com-
pressed tubercles or spines. Dorsal fin behind the middle of
the back.
3. PHOCHNA SPINIPINNIS, Burmeister, P. Z.S. 1865, p. 228.
Hab. Rio de la Plata.
10%
140 Zoological Society :—
The new species may be described as follows :—
The specimen was 52 inches, measured along the side from the
end of the nose to the notch in the middle of ibe tail. The front
edge of the dorsal fin is 23 inches from the tip of the nose, measured
over the arch of the back; the hinder edge of the dorsal fin, mea-
sured in the same manner, is 22 inches from the notch in the tail.
The front edge of the base of the pectoral fin is 9 inches from the
end of the nose; and the fin itself is 9 inches long, measured along
its front margin. The tail is 13 inches wide, measured across the
hinder edges ; the lobes are rounded, and rather overlap each other
at the central noteh.
The hinder part of the back, the whole of the dorsal fin, and the
upper and lower surfaces of the pectoral and caudal fins are black ;
the head, the lower lip, the front part of the back, and the sides i
the base of the pectoral fins are greyish black ; the upper parts of
the sides of the body behind the pectoral fins are grey, more or less
mottled with a darker shade; the chin, throat, ches belly, and
under parts of the body ae The upper and lower jaws are of
the same length. The upper lip covers the edge of the lower one,
the covered part being pale-coloured, flattened, and gradually shelv-
ing in towards the upper margin. There are two minute pits (which
may have been the places from which whiskers arose) in the upper
part of the upper lip, situated about where the depression is placed
that separates the beak from the head in those genera which have
the beak marked.
The dorsal fin is scarcely faleate, with a rather broad, rounded
upper margin, which is armed with a single series of dienes com-
pressed paherciede ; the tubercles have an oblong base, with a slightly
raised conical centre, and the surface is Concrcde with irregular radia-
ting wrinkles. Those on the front part of the edge are largest, and
separate from one another ; they diminish in size and become crowded
near the hinder upper part of the fin, forming a ridge which is hard
and serrated to the touch.
The skull is much like that of Phocena communis in size, general
form, and in the number, disposition, form, and size of the teeth; but
it differs from the skull of that species in the beak of the skull being
rather narrower, more tapering in front. The foramen maximum is
narrow, much higher than wide, and the condyles larger; while in
P. communis ane foramen maximum is nearly circular, and the con-
dyles smaller and more oblique. The sy mphysis of the lower jaw is
longer, and the sloping lower edge is more oblique and considerably
longer than in P. communis.
A skeleton is being formed of the bones of this animal; and the
skin has been preserved in spirits, which is certainly one of the best
ways of preserving the specimens of Cetacea, as it allows the outer
surface to be examimed at any future time in a state most nearly
resembling that of living specimens.
Dr. A. Giinther on Australian Pipe-fishes. 141
On THE PIpr-FISHES BELONGING TO THE GENUS PHYLLOPTERYX.
By Ausert Ginruer, M.A., Pu.D., M.D., F.Z.S,
Many Pipe-fishes are provided with short or thin cutaneous ap-
peudages, symmetrically disposed on the different dermal scutes.
These appendages are most developed in the species which may be
referred to the genus Phyllopteryx (Swains.), Kaup. The first of
these extraordinary forms was described and indifferently figured by
Shaw (Zool. v. pl. 180). He named it Syngnathus foliatus, which
name must be preferred to that given in the same year by Laccépede
(Syngnathus teniopterus, Ann. Mus. iv. pl. 58. f. 3), since the
author of a work may be presumed to have named the species at a
much earlier period than the writer of a memoir.
The British Museum possesses, among others, a fine example,
13} inches long, of this Phyllopteryx foliata from Tasmania; and
there is a beautiful coloured figure in the collection of drawings made
by Ferdinand Bauer, Dr. Brown’s companion during Capt. Flinders’s
voyage.
A second species was described by Dr. Gray as Haliichthys
teniophorus in ‘ Proc. Zool. Soc.’ 1859, p. 38, and figured plovins;
it is from Freycinet’s Harbour.
A third species has been lately presented to the British Museum
by Mr. George French Angas, who received it from Port Lincoln,
South Australia. I name it Phyllopteryx eques. Its form is still
more extraordinary than that of the preceding species, the spines,
crest, and cutaneous appendages being much more developed,
and the trunk being dilated into an upper and three lower pro-
minences. ‘The snout is as long as the distance of the front mar-
gin of the orbit from the hind part of the nape; it bears a pair of
small spines behind the middle of its upper edge, a pair of minute
barbels at the chin, and a pair of long appendages in the middle of
its lower part. The forehead bears an erect, broad, subquadrangular
crest, with a shorter single spine behind ; a horizontal spine above
each orbit ; a cluster of spines with narrow appendages on the occi-
put. Nape of the neck with a long spine, dilated at the base into a
crest, and carrying a long bifid appendage.
The trunk is compressed, somewhat dilated, strongly arched on
the back, and with two deep indentations in its lower profile. There
are seventeen bony rings between the pectoral fin and the root of the
tail. The spines are of three kinds: 1. The band-bearing spines are
the strongest, strongly compressed, not flexible, each terminating
in a pair of short points. There are one pair of these spines in the
middle of the back, and one on each of the three prominences: of the
abdominal outline ; the flaps are Jong and bifid. 2. Very long, com-
pressed, and somewhat flexible spines, without appendages; these
oceupy in pairs the uppermost part of the back, and in a single series
the median line of the belly. 3. Small, short, conical spines run in
single series along the median line of the sides, and along the lateral
edges of the belly ; a pair of similar spines in front of the lower part
of the base of the pectoral fin.
142 : Miscellaneous.
Tail quadrangular, with sharp edges, and with five pairs of band-
bearing spines along its upper side ; its end is slightly prehensile.
P. 20. D.37. The dorsal is situated entirely on the tail.
The specimen, being dry, has lost its original colours, which were
probably red during life. The iris is crossed by radiating streaks ;
and several other streaks (of a whitish colour) radiate from the eye
over the opercles and the upper part of the head.
There is no doubt that these fish attach themselves with the pre-
hensile end of their tail to stems of seaweed or other objects; and
when they are in the vicinity of seaweed of a similar colour, their
resemblance to it must be so great that they would easily escape
being observed by their enemies.
MISCELLANEOUS.
The Food of the Aye-Aye.
To Dr. J. E. Gray.
Dear S1r,—The specimen of sugar-cane I sent for your examina-
tion a few days since exhibits in a clear manner the mode of using
its incisor teeth by the Aye-Aye. This animal, as you are aware,
came here in August 1862, and during the period of nearly three
years has been kept in good health and condition, its food being
varied from time to time. It was only recently that I obtained some
fresh green sugar-cane, and placed two or three sticks of the same in
the cage of the Aye-Aye. I soon found the animal was fond of this
kind of food ; and it is interesting to observe how well its teeth are
adapted for obtaining the juice and sugar from each of the joints of
the cane. As will be seen, the long points of the incisors cut deeply
into the cane, the fibre being pulled forward, and the moisture
chewed out. In the observations made by me, and published in the
‘Annals’ for July 1863, p. 72, I stated that the animal feeds freely
on a mixture of milk, honey, eggs, and any thick glutinous fluid;
and, from what I had observed, I was led to think the creature fed
upon the juices of trees; and I am induced to send you this short
notice as an additional proof of the correctness of my statements.
I am, dear Sir,
Yours faithfully,
A. D. BARrtTLettT.
On the Histology of the Acalephee. By Prof. KOLLIKER.
Professor Kolliker has published in the ‘ Wiirzburger naturwis-
senschaftliche Zeitschrift’? some observations made by bim upon the
histology of the Hydrozoa and Ctenophora in the Firth of Clyde.
In these animals he distinguishes three kinds of connective tissue.
One forms the tentacles of the Hydroid Polypes and all the solid
tentacles of the Medusze. It presents the appearance of a series of
cells (muscular cells of Keferstein) occupying the axis of the tenta-
cle. These cells possess no contractility ; at least the tentacles of
Miscellaneous. 143
the Aginide and Trachynemide, which present this structure, are
rigid.
“The contractile tentacles owe their contractility to a muscular layer
situated between the cellular axis and the external epithelium. This
cellular axis is only a dependence of the internal epithelium which
lines the digestive cavity (Hydroids) or the marginal canal (Medusze).
It probably acts as an elastic organ antagonistic to the muscular
layer.
The second kind of connective tissue is a substance destitute of
cells, which forms the umbrella of all the simple Medusze, including
the gelatinous substance of the natatory bells and covering laminge
of the Siphonophora. Sometimes this substance is entirely homo-
geneous; sometimes it is traversed by numerous fibres very like the
elastic fibres. In an quorea these fibres are attached to a mem-
brane capable of isolation, placed beneath the epithelium of the
convex surface of the umbrella.
The third form is the well-known gelatinous substance with dis-
seminated cells of the umbrella of the higher Medusze. Professor
Kolliker agrees with Professor Virchow in denying the existence of
these ale in Cyanea capillata.— Bibl. Univ. May 1865, Bull.
Seient. p. 66.
On a New Type in the Group of Ascidians—Chevreulius callensis.
By M. Lacazr-Duruters.
After describing the general characteristics of the Ascidia, the
author says :—The specimen which forms the subject of the present
memoir exhibits an exceptional and very remarkable arrangement,
which masks the true characters of the group. All the indivi-
duals of the genus Chrevreulius are without stolons or buds which
might lead to their being approximated to the social Ascidia, and
still less to the compound forms. Their form is that of a cylinder,
free at one extremity, adherent by the other, and shghtly flattened
on that side which is in contiguity to the foreign body. The free
superior extremity presents the characters of the genus.
The test, which is of a yellowish colour and cartilaginous, is suffi-
ciently resistant to retain its form after desiccation ; its thickness is
not great, and it resembles a thin lamina of pale horn. When it is
contracted, the orifices are not visible; but as it becomes distended,
more than half of the flat upper .extremity of the cylinder is soon
seen to detach itself towards the circumference, and to rise by moving
as if round a hinge placed on the side of the cylinder which is
flattened.
Beneath the plate which rises thus so as to form a right angle
with its former position, and which represents a valve, there appears
a white transparent tissue—a membrane stretched from one side to
the other of the separated parts, so as to fill up the great fissure
produced by this sort of gaping. Upon this membrane two mamillee
soon rise, at the summit of which open the two orifices characteristic
of the Ascidia. One of these leads into the branchial chamber, and
144 Miscellaneous.
consequently to the mouth; this is the highest one: the other, less
prominent and placed laterally, gives passage to the water which
traverses the branchiz, to the residues of digestion, and to the pro-
ducts of reproduction.
Between these two orifices a small opaque-white nucleus may be
distinguished through the tissue, with delicate filaments issuing from
it: this is the nervous ganglion.
Thus Chevreulius is undoubtedly an Ascidian, but it is a bivalve
Ascidian, of which the test is divided into two parts moveable upon
each other, as in the Acephala; and the Ascidia themselves must be
arranged in two series—one for those in which the external envelope
is a true little leather bottle with two apertures, the other for those
in which the test, divided into two parts by a broad horizontal cleft,
becomes bivalve.
Having met with Chevreu/ius for the first time in the waters of
Calle, I have named it C. callensis. It lives at great depths (60, 80,
or 100 fathoms), and belongs to the fauna of the coralligenous zone.
In conclusion, the author remarks upon the interest attaching to
the discovery of Chevreulius, as an Ascidian with an upper and
lower valve, in connexion with the relation existing between the
Tunicata and Brachiopoda. — Comptes Rendus, June 19, 1865,
p. 1264.
On some singular Organs appended to the Feet of certain Crustacea.
By MM. Crats and Sars.
Professor Claus (Zeitschr. wiss. Zool. xiii. p. 422) and Professor
Sars (Videnskabsselskab. FGrhandl. 1863) have independently in-
vestigated the Schizopod Crustacea of the family Kuphausidee with
regard to the singular organs already alluded to by Dana, Semper,
and Kroyer, and “regarded by Semper as eyes, and by Kréyer as
auditory organs. These are spherical organs, of a reddish colour,
situated at the base of several of the thoracic legs and of the first
four pairs of abdominal appendages. Both the authors above men-
tioned have demonstrated the correctness of Semper’s view, although,
besides these pedal eyes, the animals possess the two large com-
pound eyes common to all Decapoda. Each of the thoracic and
abdominal eyes receives a special nerve from the ventral ganglionic
chain. The organ itself is a spherical bulb, moved by special mus-
cles; and in it may be distinguished a crystalline lens, a vitreous
body, a pigment- -lay er, and a retina of complex structure. The
existence of a cr ystalline lens distinct from the cornea is very striking,
as remarked by M. Sars; for in other Crustacea no true crystalline
exists, its function being performed by the thickened and inflated
cornea. According to M. Claus, the position of the four pairs of
abdominal eyes is very remarkable: the first pair looks forwards,
the last pair backwards, and the two intermediate pairs downwards.
— Bibl. Univ. May 1865, Bull. Scient. p. 63.
THE ANNALS
MAGAZINE OF NATURAL HISTORY.
[THIRD SERIES. ]
No. 93. SEPTEMBER 1865.
XVI.—On a new Lizard, with Ophidian affinities, from the Lower
Chalk (Saurospondylus dissimilis). By Harry Srexey,
F.G.S., of the Woodwardian Museum, Cambridge.
Proressor Owen described the Raphiosaurus subulidens from
the Lower Chalk of Cherry Hinton, and I should perhaps have
been inclined to refer the vertebra here described from that
locality to the same genus, had not a sight of Mr. Carter’s type
specimen shaken my faith in its reptilian character. So far as
external features go, there is nothing to suggest that it is not
the jaw of a fish. Even were it reptilian, it is so dispropor-
tionately large in comparison with this vertebra, that the iden-
tity of the two would still be doubtful. But Professor Owen
appears to know the vertebre of Raphiosaurus; for, in the
‘Paleontology’ (p. 311, 2nd ed.), it is on such evidence that
the species is said to be based.
The Lizards yet known from the Chalk have proccelian ver-
tebree with that simple structure of the zygapophyses in which
the front articulations are turned up and exposed, and the back
pair turned down. This structure, characteristic of most ver-
tebree, would appear to result from the fact that the hmbs sup-
port that part of the skeleton which is in front of them—a func-
tion manifest in the straight or upward tendency of the neck,
where each vertebra rests on its zygapophyses, and the down-
ward direction of the tail, where the vertebree hang without
support under the zygapophyses.
The development and origin of these processes and the form
of the bones depend on the functions of the muscles, though in
a less degree than in the limbs, where bones appear to have
owed their very existence to muscular and functional action.
But in Saurospondylus the vertebra has ten articular facets, as in
Serpents and in the Iguana Lizards, in which the neurapophyses
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. 1]
146 Mr. H. Seeley on a new Lazard
overhang and lock on to the zygapophyses of the vertebra in
front. Such a structure indicates a flexible vertebral column,
for it allows of dorsal vertebrae being supported by the anterior
limbs. In the lumbar region of many mammals, such as the
Armadillo, the Raccoon, and even the wild Cat, where there is
much upward and downward motion, there is a near approach
to a like modification of yoking.
This vertebra, with depressed centrum, obliquely overhanging
and transversely oval cup and ball, zygosphene, and zygantrum,
indicates the lower dorsal region of a small reptile having its
nearest affinities with Jguana. It is $ths of an inch long, not
quite so wide in front where widest. It was found in the lower
Chalk of Cherry Hinton, near Cambridge.
There is no neural spine, and no hypapophysis.
The inferior surface of the centrum is subtriangular. The
length from the base of the cup in front to the base of the ball
behind is equal to the width of the zygapophyses in front.
From these, two strong curved ridges descend and approximate
to the bottom of the sides of the ball. The subtriangular area
so enclosed is a little convex transversely and concave in length.
The vertebra is ;%-ths of an inch high in front, nearly one-
half being the height of the centrum, and the remainder that of
the neural arch, which is higher behind than in front, and may
there have had a slight neural spine. The neural arch on each
side is a smooth cupped surface, with a concave border, and
contracts behind.
The anterior zygapophyses are horizontal square surfaces,
hardly above the border of the cup, from which they are sepa-
rated in front by a perpendicular concavo-convex surface on
each side, about the size of their own articulations.
The zygosphene projects in front of the vertebra, and is just
as wide as the cup. Its superior front margin is concave and
horizontal. Its flat articular surfaces, which look forward and
outward, are very narrow, and entirely between the zygapo-
physes, above and in front of which they project about half their
own length.
The neural arch is very thin in front, thicker behind, where
the processes are less perfectly preserved.
The sides of the vertebra are narrow, concave, wedge-shaped
surfaces bordered by (1) the basal ridges already mentioned,
and (2) the concave ridges at the sides, which from above make
the outline of the neural arch, are laterally parallel with its top
ridge, and connect the anterior with the posterior zygapophyses.
Both ridges meet in front, below the zygapophyses in the
tubercle for the rib, which is broken off on both sides.
The body of the centrum and the neurapophyses appear to be
from the Lower Chalk. 147
cellular. The outline of the posterior end of the vertebra is
pentangular, as high as wide. The height of the ball appears
to be less than the depth of the cup.
Of the vertebral differences of Serpents and Lizards little is
known. This fossil resembles a serpent more than could have
been expected, and yet in other modifications comes near to the
Lizards. A Cretaceous serpent may have been more Lacertian
than any now known, and a lizard of the Chalk may have been
more Ophidian.
The chief Lacertian features which I detect are—
(1.) The absence of an hypapophysis, which all serpents ap-
pear to have, though in some (as in Python) it is very slight.
(2.) The depressed centrum, with transversely oval and over-
hanging cup and oblique ball, are Lizard characters, though the
cup is oblique in Crotalus, Paleryx depressus, &c., and the ball
is transversely elliptical in some other forms.
(3.) The absence of additional diapophyses besides the costal
tubercle is characteristic, though they are not found in all
Ophidia.
(4.) The neural arch is not notched between the zygantra, as
in Serpents, but is prolonged back a little between and over
them, as in Jguana. The zygantra are excavated in the middle
of the sides of the neural canal, as in Jguana, and not at its
summit, as in most serpents, though Naja, Hydrus, Natriz, &c.,
are exceptions.
(5.) The zygosphene projects well over the cup, as in Iguana,
and is not level with it, as in Ophidians.
(6.) Iguana has similar basal ridges, and depressions under
the costal tubercle, like those in the fossil, only more developed.
But neither in lizards nor in serpents does the basal ridge meet
the ridge between the zygapophyses, because the costal tubercle
is always lower down.
The more marked Ophidian characters are—
(1.) The broad quadrate form, which is nearer to Paleophis
than to Iguana, though the anterior vertebre of this and most
lizards are as short. In Scincus there is much the same general
form of the vertebra, and a like absence of neural spine and
hypapophyses ; but the zygosphene can hardly be said to exist,
the zygapophyses are never horizontal, and are well raised above
the tubercle for the ribs.
(2.) The horizontal zygapophyses, level with the top of the
cup, find their parallel in EHrya and most Serpents; but in
Iguana, the nearest to it of the Lizards, they are higher. Zguana
wants the sharp ridge connecting the zygapophyses; it is cha-
racteristic of some serpents, but is also found in Scincus.
The balance of evidence from the few data at my command
11*
148 Dr. J. E. Gray on a new Finner Whale from Formosa.
rather inclines to the conclusion that Sawrospondylus was an
Iguanoid Lizard, hardly separable from the Serpents, than that
it indicates a Cretaceous Ophidian. So classed, it is the type of
a new family.
XVII.—WNotice of a new Finner Whale from Formosa.
By Dr. J. BE. Gray, F.R.S. &e.
Mr. Swrnuocr has kindly sent me some bones of a Finner Whale
which was cast ashore on the coast of Formosa.
The cervical vertebrae show that it is quite distinct from any
Whale the bones of which have previously come under my
examination.
It agrees with the smaller Finner, Balenoptera rostrata of
Europe, in the second and third cervical vertebre being united,
while in all the other true Finners known they are free; and
also in the subcircular form of the front part of the neural canal.
I am therefore inclined to refer it provisionally to the genus
Balenoptera as restricted in my paper (Proc. Zool. Soc. May 24,
1864) ; but I think it probable that, when we know the entire
number of the vertebre and other details of the skeleton, it will
prove to be a distinct form.
The Whale may be named Balenoptera Swinhoei.
The second cervical vertebra with large, broad, truncated,
lateral processes, with a large, oblong, subcentral perforation ;
the lateral processes are each two-thirds of the transverse dia-
meter of the articulating surface of the body of the vertebra.
The third cervical united to the second by the anchylosis of
the neural arches; the body thin, oblong, transverse, broader
than high ; the lateral processes slender, truncated at the end,
not so long as the transverse diameter of the body, curved to-
wards each other at the end, but not forming a ring.
The rest of the cervical vertebre free.
The sixth or seventh cervical with a thin body, and a slender,
nearly straight upper lateral process, and only a very short
tubercle on each side below.
The neural cavity of the second cervical vertebra subcircular,
rather less high than broad, and not quite so wide as half the
diameter of the front side of the body.
The neural cavity of the third cervical vertebra oblong, trans-
verse, rounded above, as wide as half the transverse diameter of
the body, and about one-third broader than high.
The bones are nearly the same size as the similar bones in the
Physalus antiquorum, which is between 60 and 70 feet long when
alive ; they therefore belong to an animal at least three times as
large as the Balenoptera rostrata of Europe.
Dr. A. Krohn on the Male Generative Organs of Phalangium. 149
XVIII.—On the Male Generative Organs of Phalangium.
By Dr. A. Kroun*.
From dissections which I have very recently made, it appears
that the notions of Treviranus and Tulk as to the male sexual
apparatus of Phalangium still generally adopted require to be
essentially modified. The principal question here is as to the
still unexplained signification of a gland-like organ furnished
with two efferent ducts, which is situated in the abdomen upon
the lower wall of the alimentary tube, occurs only in the male,
and appears, as Treviranus asserts, to have some connexion with
the other parts of generation.
My investigations have proved that the above-mentioned organ
is the testis; so that the pair of glands consisting of ramified
lobes or czeca situated in the anterior part of the abdomen, to
which Treviranus and Tulk ascribe the function of preparing
the seminal fluid, have quite a different destination.
When the abdomen is opened from the ventral surface, the
testis falls out, and appears, after the removal of the adherent
fatty body, as a sausage-shaped greatly curved organ t of a dull
white colour, bridged over by the two retractor muscles of the
penis (fig., a). From the extremity of each of its horns (which
The male generative apparatus of the second species of Phalangium men-
tioned in the text, without the accessory glands. Its component parts are
removed from their natural position in order to show their connexion.
a. The Testis. 66. Vasa efferentia. c. Coil of the vas deferens.
d. Dilated portion of the vas deferens. e. Sheath of the penis, with
the penis (f) within it. gg. Retractors of the penis.
are directed forwards, and do not reach to the part where the
two breathing-orifices or stigmata occur on the outer surface
* Translated by W. 8S. Dallas, F.L.S., &e., from Wiegmann’s Archiv,
1865, pp. 41-48.
+ The sigmoid or zigzag form ascribed to this organ by Treviranus and
au is probably only a consequence of injury or displacement during
issection.
150 Dr. A. Krohn on the Male Generative Organs of Phalangium.
of the body) there issues one of the above-mentioned narrow
canals, which may with perfect justice be characterized as vasa
efferentia (bb). In its forward course each of these canals
strikes first of all upon the origin of the tracheal stem of its side,
then bends inwards, and runs to the median line of the abdo-
men, where it meets with the canal of the opposite side, and
both pass into the origin of the vas deferens. Of the nature
of this long duct, which gradually increases in diameter, and is
chiefly rolled together into a close coil (c), Tulk has already
given a satisfactory account. I can completely confirm the
statement of this naturalist that the vas deferens, after passing
through the penis, opens at the extremity of the so-called glans*,
which is armed with a curved spine or hook at its apex, and is
moveably articulated upon the shaft of the penis. I may, how-
ever, remark that the vas deferens, after having become suddenly
and greatly dilated (d) just before its entrance into the penis,
appears so exceedingly narrow during its passage through
the latter, that the transverse section of its lumen bounded by
a chitinous coat only seems very slightly to exceed the diameter
of a single seminal corpusclet.
The testis possesses a bounding membrane which passes into
the outer envelope of the efferent ducts; it is, however, not
hollow, but its mass consists throughout of round cells, fur-
nished with a distinct wall, and closely pressed together, which
contain a great number of small transparent vesicles. As a
nucleus, often surrounded by dark granules or molecules, makes
its appearance in these vesicles on the addition of acidulated
water, I regard them as the formative cells of the semen, whilst
the cells enclosmg them seem to represent their mother cells.
* This so-called glans no doubt functions as an excitant organ during
copulation.
+ The above-mentioned dilated portion of the vas deferens is distin-
guished from the preceding portions of this canal by a very thick chitinized
liming membrane and by a very strong muscular coat. From the great
narrowness of the duct im its passage through the penis, it might be sup-
posed that the dilated portion may act as an organ of propulsion in the
ejaculation of the semen. According to Tulk, the excitant organ (the so-
called glans), which, during repose, is always bent back over the end of
the shaft of the penis, ean be elevated or extended (that is, brought into
the same line with the shaft of the penis) by means of two muscles. I
must deny the existence of these muscles; but, on the other hand, it is
not difficult to detect the presence of a single powerful muscle which is
evidently destined to this purpose. This muscle, which has hitherto been
overlooked, occupies half the length of the interior of the penis from the
base, and is connected with a strong sinew running straight to the excitant
organ, to the base of which it is attached. The mode of action of this
muscle may be easily ascertained by a simple experiment, namely, by laying
bare the sinew, inserting a fine needle into it, and pulling in the direction
of the traction of the muscle,
Dr. A. Krohn on the Male Generative Organs of Phalangium. 151
In support of this view I may refer to the data already extant
as to the development of the semen in some Arachnida (sce
Von Siebold, Vergl. Anatomie, p. 544, note 6; and Leydig,
** Ueber den feineren Bau der Arthropoden,” in Miiller’s Archiv,
1855, p. 470).
Mature semen is usually found in greater or less abundance
in the entire portion of the vas deferens before the dilatation.
The seminal corpuscles are rounded structures, furnished, I
believe, with a disciform nucleus. The oscillating movement
which is observed in them when they are not too closely pressed
together appears to be referable to the phenomenon of the so-
called molecular movement *.
The two accessory glands situated in the anterior portion of
the abdomen immediately above the sheath of the penis are con-
nected by connective tissue and tracheal ramifications with the
coil of the vas deferens, which is placed between them. Their
intimate structure is founded on the same plan that has been
made known to us in the glands of many insects by the ad-
mirable works of H. Meckel, and especially of Leydig. Thus
we may distinguish in them a homogeneous external envelope
(tunica propria), a subjacent, proportionally thick layer of se-
creting cells, and within this a ling membrane (intima). The
lumen of the lobules or ceca appears to be a comparatively
narrow canal, from the circumference of which, throughout the
whole length of the canal, numerous fine tubules pass deeply
into the cellular layer. In the male of P. opilio (P. cornutum)
single tubules are seen at intervals, which are distinguished
from the rest both by their greater size and by their branching
within the cellular layer. The canals of all the sacs, after
uniting to form larger branches, finally combine into a main
duct extending forward through the midst of the gland, and
opening upon the upper wall of the sheath of the penis not far
from the sexual aperture. This duct, however, is never free, as
the layer of secretion-cells is continued upon it and envelopes
it as far as its outlet. The orifices of the two main ducts at the
point just mentioned lie close together on each side of the
median line. On the lining membrane of the main ducts and
their first branches a so-called spiral filament, similar to that
of the trachez, may be detected. Inthe male of P. opilio it may
be pretty clearly distinguished even on the above-mentioned fine
tubules which penetrate the cellular layer.
* T have not been able to give a satisfactory description of the appear-
ance of the seminal corpuscles under a high magnifymg power. Accord-
ing to Leydig’s investigations, they are round, flat structures, with a cen-
tral band-like elevation. Leydig regards their oscillatmg movement as
quite spontaneous, and therefore assumes that they probably possess a fine
capillary appendage (J. c. p. 469, pl. 17. fig. 41 d).
152 Dr. A. Krohn on the Male Generative Organs of Phalangium.
These two glands also occur in the females; but even when
their pregnancy is very far advanced, the size of these glands is
less than in the males. In their structure they differ from those
of the males only in the circumstance that the spiral filament
appears to be entirely wanting in the ramifications, although
present in the main ducts. The spot at which the two main
ducts discharge themselves corresponds exactly with that of the
male. Their orifices are also in the vicinity of the sexual orifice,
upon the upper wall of the sheath embracing the laying-tube*.
As to the use of the secretion of these glands nothing can at
present be stated with certainty. In the male, the secretion is a
clear, tenacious, thickly fluid substance, apparently very similar
to the spinning-material of the Araneida.
In conclusion, I must refer to an exceedingly remarkable
phenomenon which I have observed in the examination of nearly
all males of P. opiliot. This is nothing less than a production
of eggs from the testis, at the same time that the development
of the semen is by no means diminished. The number of ova
produced by the testis may sometimes be so great that, as in
the ovary, they occupy the entire surface; or it may be very
small, and in this case the ova occur only on particular spots of
the testis. In the former case the ova, as on the ovary, present
the most various states of development, from the smallest, with
the vitellus still clear, to those in which the vitellus appears
more or less turbid. These ova, however, appear never to attain
the full size of those produced on the ovary. I have only ob-
served one case in which, among a number of ova, two or three
were remarkable for their preponderant size. These ova agreed
perfectly with the nearly mature ovarian ova, not only in their
size, but also in the nature of the vitellus, which appeared of a
chalky whiteness. In a second species (which, from the form
of the penis, appears to be the one investigated by Treviranus
and Talk), the males of which I was able to procure much more
frequently, I have rarely detected ova upon the testis, and when
present they were always but little developed.
To remove the suspicion that I may perhaps have erred in
* J may remark here, in passing, that the two supposed czecal tubes ex-
tending to the laying-tube, which Tulk has regarded as cement-glands, are
really nerves, as has already been indicated by Gegenbaur (Grundziige der
vergl. Anat. p. 276). I have succeeded im tracing them to their origin in
the thoracic ganglion. They also occur, although of less size, in the male,
in which they accompany the portion of the vas deferens issuing from the
coil to its immersion in the penis. In both sexes they supply the retrac-
tors of the copulatory organs, and also penetrate with a portion of their
branches into the interior of the latter, and then become further divided.
+ The males of this species are exceedingly rare in comparison to the
females.
Mr. J.S. Baly oa new Species of Crioceride. 153
the interpretation of what I have seen, I may refer to the testi-
mony of a celebrated witness, who detected the same pheno-
menon long before me. This is Treviranus, who makes the
following statement :—“ In one of the Phalangia that I examined
I found an ovary filled with eggs, but, mstead of the laying-tube,
a male genital organ. Hermaphroditism, which has often been
observed in the Lepidopter a, appears, therefore, not unfrequently
in the Phalangia” (/.c. p. 38).
This case, in my opinion, agrees exactly with those observed
by me, if we only admit that “the organ described as an ovary
can have been nothing but the testis. And this will be the less
doubtful when we consider that, as already stated, Treviranus
certainly saw the testis, but was not in a position to recognize
it as such.
As regards the ultimate fate of these eggs, there seems to be
no doubt that, after persisting for a longer or shorter time, they
disappear. In favour of this we may quote as an analogous
example the case of some of our indigneous toads (Bufo variabilis,
B. calamita, and especially B. cinereus), the males of which, ac-
cording to the thorough investigations of Wittich, possess,
besides a testis, a more or less rudimentary ovary*. From these
investigations it appears clearly that the ova produced by this
ovary, nee attaining a certain degree of maturity, finally become
aborted and disappear tre
XIX.—Descriptions of new Species of Crioceride.
By J. S. Baty.
Crioceris scabrosa, Baly.
. C. elongata, subcylindrica, plumbea, subnitida, pube aureo-sericea
brevi vestita ; capite rugoso ; antennis dimidio corporis longioribus,
obscure rufo-piceis, basi et apice nigris; thorace sat profunde
transversim strigoso, fere glabro, lateribus vix constrictis ; elytris
crebre rugoso-punctatis, elevato-reticulatis, reticulis fere leevibus,
glabris, disco anteriore dilatatis, et ibi superficiem totam fere am-
plectentibus ; tarsis obscure cyaneis; tibiis intermediis ad apicem
incrassatis, intus curvatis.
Long. 4 lin.
Hab. Mexico.
Elongate, subcylindrical. Head not constricted behind the
eyes ; face subelongate, finely rugose, sparingly clothed on the
* “© Beitrage zur morphol. und histol. Entwickelung der Harn- und
Geschlechtswerkzeuge der nackten Amphibien,” Siebold and Kolliker’s
Zeitschrift, Bd. iv. p. 159.
+ Morphologically it is an interesting fact that the ovary in the above-
mentioned toads occurs as a perfectly independent organ having no further
connexion with the testis, whilst the seminal gland of Phalangium (espe-
cially in P. opilio) has completely the character of an hermaphrodite gland,
154 Mr. J.S. Baly on new Species of Crioceride.
labrum, lower parts of epistome, and round the eyes with short,
adpressed, aureo-sericeous hairs; front impressed with a deep
fovea, on either side of which is a fine but distinct groove, which,
commencing at the apex of the epistome, runs obliquely upwards
on either side to surround the upper portion of the orbit of the
eyes, but at a considerable distance from the eye itself; eyes
not mounted on a raised orbit, rotundate, entire; antenne
slightly thickened towards their apex, moderately robust. Tho-
rax cylindrical, slightly flattened above, subquadrate, its surface
closely covered with deep, irregular, transverse grooves ; just in
front of the base, and again behind the apex, are two indistinct
transverse depressions. Scutellum smooth, triangular, its apex
obtusely truncate. Hlytra much broader than the thorax, pa-
rallel, acutely rounded or almost angulate at their apex; sub-
cylindrical above, slightly flattened along the back, obsoletely
depressed transversely below the basilar space, their apical por-
tion suddenly and obliquely deflexed; closely and somewhat
finely rugose, closely clothed with short, adpressed, aureo-seri-
ceous pubescence ; whole surface (with the exception of the de-
flexed apex) covered with large, broad, raised, glabrous reticula-
tions; on the anterior half of the inner disk these elevations
become much dilated, and coalesce so as to occupy by far the
greater portion of the whole surface. Body beneath clothed with
coarser hairs than the upper surface ; abdomen smooth, impunc-
tate, each segment impressed on either side its centre with a deep
fovea, from the surface of which springs a tuft of pale suberect
hairs; hinder thighs extending rather beyond the apex of the
abdomen.
Crioceris rugata, Baly.
C. elongata, nigra, nitida, thoracis dorso elytrisque fulvis ; antennis
subfiliformibus, vix compressis et perfoliatis ; thorace subquadrato,
lateribus leviter coarctato, dorso rude punctato ; elytris infra basin
obsolete transversim depressis, profunde punctato-striatis, inter-
stitiis basi planis, apicern versus subelevatis ; scutello piceo, glabro.
Long. 33 lin.
Hab. Japan.
Head deeply constricted behind the eyes ; forehead impressed
with a deep groove; face triangular, closely covered with ad-
pressed sericeous hairs ; eyes deeply notched ; antenne half the
length of the body, moderately robust, subfiliform, first, third,
and fourth joints equal in length, fifth to the eleventh each half
as long again as the fourth, equal, somewhat thickened and
compressed, scarcely perfoliate. Thorax subquadrate, subcy-
lindrical, sides moderately constricted in the middle, stained in
front with nigro-piceous; upper surface deeply and coarsely
punctured. Scutellum triangular, its apex truncate. Elytra
much broader than the thorax, parallel, upper surface slightly
Mr. J.S. Baly on new Species of Crioceride. 155
flattened along the suture, indistinctly depressed transversely
below the basilar space, very deeply punctate-striate, interspaces
impunctate, plane at the base, indistinctly rugulose on the
transverse depression, distinctly thickened towards their apex,
being subcostate near the outer margin and suture. Body be-
neath nearly glabrous, the pleura clothed with adpressed seri-
ceous pubescence ; abdomen smooth, impunctate, very sparingly
furnished with subdepressed silky hairs. Hinder thighs not
thickened, not extending beyond the second abdominal segments.
Very closely allied to C. impressa, smaller, much more coarsely
punctured ; antenne more slender.
Crioceris ruficollis, Baly.
C. oblonga, nigra, nitida, collo thoraceque rufis, hoc vix elongatulo,
subcylindrico, lateribus medio modice coarctato, dorso vage sub-
remote punctato; elytris convexis, nigro-ceeruleis, infra basin
leviter transversim impressis, distincte sed tenuiter punctatis, in-
terstitiis planis.
Var. A. Capite antice obscure rufo ; antennis basi piceis.
Long. 3% lin.
Hab. Northern China. Collected by Mr. Fortune.
Face triangular, clothed with aureo-sericeous pubescence ;
antenne moderately robust, slightly compressed, subperfoliate,
more than half the length of the body. Seutellum rufo-piceous.
Elytra broadly oblong, much broader than the thorax. Pubes-
cence on the under surface of the body silvery white, that on
the legs with a golden tint. Abdominal segments each with a
transverse groove, which is clothed with a smgle row of sub-
erect silvery hairs. Hinder thighs very slightly thickened, their
apex not reaching beyond the apex of the third abdominal
segment.
Lema quadriplagiata, Baly.
LZ. oblongo-elongata, fulva, nitida, subtus sparsim fulvo-sericea,
ore supra nigro; antennis modice robustis, corporis longitudine
brevioribus, articulo tertio secundo duplo longiore ; thorace cylin-
drico, subquadrato, lateribus medio valde constrictis, dorso leevi,
pone medium profunde transversim suleato, disci medio obsolete
lineatim punctato; elytris infra basin obsolete transversim im-
pressis, sat fortiter punctato-striatis, striis ad apicem sulcatis, in-
terspatiis postice elevatis, utrinque plagis magnis duabus obscure
violaceis.
Var. A. Antennis (basi excepta) fuseis; corpore subtus fusco
variegata.
Var. B. Elytris totis metallico-ceruleis.
Long. 23 lin.
Hab. Pachybouri. Collected by the late M. Mouhot.
This species may be separated from L. hastrio, Clark, by the
156 Mr. J.S. Baly on new Species of Crioceride.
following character :—In the present insect the third joint of
the antenna is double the length of the second, whilst in L.
histrio it is only from one-third to one-half longer.
Lema Adamsii, Baly.
L. elongata, subcylindrica, pallide fulvo-flava, nitida ; capitis maculis
duabus, thoracis maculis quatuor, quadratim dispositis, elytrorum
maculis quatuor, duabus pone basin parvis, duabusque ante apicem
magnis, pectore, femorum singulorum macula, tibiarum apice,
tarsis antennisque nigris, his subfiliformibus, dimidio corporis
longioribus, articulis duobus basalibus fulvo-flavis ; thorace sub-
quadrato, lateribus sat coarctato, ante basin transversim suleato,
disco levi fere impunctato; elytris fortiter punctato-striatis, infra
basin vix transversim impressis, interspatiis leevibus, antice trans-
versim rugulosis, apice callosis.
Long. 3 lin.
Hab. Chusan. Collected by Mr. A. Adams.
The more robust, shorter, black antenne, with the second
and third joints nearly equal, will distinguish this species from
L. quadriplagiata and L. histrio.
Lema Downesii, Baly.
L. subelongato-parallela, pallide fulva, nitida, antennis (basi excepta)
tarsisque fuscis ; scutello elytrorumque vitta suturali, ante apicem
abbreviata, nigris; thorace elongatulo, basi transversim sulcato,
lateribus vix pone medium modice constrictis, antice subgloboso,
leevi, lateribus et disci medio fortiter subremote punctato ; elytris
infra basin haud transversim impressis, profunde punctato-striatis,
interspatiis prope apicem elevatis, secundo octavoque apice coéun-
tibus callosisque.
Long. 2 lin.
Hab. Bombay ; Bengal.
Thorax smoother and more swollen than in L. suturella;
third joint of antenna one-half longer than the second, equal in
length to the fourth.
Lema suturella, Baly.
L. oblongo-parallela, fulva, nitida, tibiis tarsisque infuscatis, pectore
abdominisque basi nigris ; antennis (basi excepta) fuscis ; thorace
cylindrico, lateribus medio modice constrictis, supra ante basin
transversim suleato, leviter irregulariter punctato ; scutello piceo ;
elytris infra basin vix transversim impressis, fortiter punctato-
striatis, interspatiis prope apicem costatis, linea suturali cyanea,
ante apicem abbreviata, instructis.
Long. 2 lin.
Hab. Bengal.
Thorax less swollen than in L. Downesii, irregularly punctured;
a longitudinal space down the middle of the disk, together with
Mr. J. S. Baly on new Species of Crioceride. 157
the sides in front, more coarsely punctured than the remainder
of the surface; third joint of antenna one-half longer than the
second, distinctly longer than the fourth.
Lema concinnipennis, Baly.
L. elongata, parallela, subcylindrica, nitido-ceerulea ; antennis nigris,
abdominis apice fulvo; thorace subquadrato, subeylindrico, lateri-
bus medio modice constrictis, disco ante basin transversim sulcato,
fortiter punctato, spatio longitudinali centrali levi, fere impunc-
tato; elytris infra basin transversim impressis, intra callum hu-
merale sulcatis, punctato-striatis, punctis ad apicem minus fortiter
impressis, interspatiis planis; antennis corporis dimidio longiori-
bus, modice robustis.
Var. A. Abdomine corpore concolore.
Long. 24 lin.
Hab. Northern China.
Face subelongate ; eyes deeply notched; lower portion of
face closely subrugoso-punctate; forehead impressed with a
short longitudinal groove, very finely punctured. Legs mode-
rately robust.
Lema Psyche, Baly.
L. oblonga, valde convexa, chalybeo-czerulea, nitida ; antennis pedi-
busque nigris; thorace subtransverso, cylindrico, lateribus medio
modice constrictis, disco tenuissime punctato, basi impresso ;
elytris purpureis, robustis, thorace multo latioribus, infra basin
transversim depressis, sat fortiter punctato-striatis, striis integris,
interspatiis ad apicem convexiusculis, obsolete costatis ; antennis
gracilibus, filiformibus, articulis 3° et 4° zequalibus.
Long. 3 lin.
Hab. Northern India.
Face subelongate, vertex coarsely punctured; eyes notched ;
antennze two-thirds the length of the body; transverse depres-
sion of elytra shallow, obsoletely wrinkled. Legs slender ;
thighs scarcely thickened.
The broad elytra and robust form of this insect at once dis-
tinguish it from its congeners.
Lema bipunctata, Baly.
L. subelongata, piceo-fulva, nitida; antennis (articulo basali preeter-
misso), scutello, thoracis vitta laterali, abdominis fasciis, tibiarum
apice tarsisque nigris; elytris singulatim fovea infra basin, linea
suturali alteraque marginali, apice abbreviata, chalybeis.
Long. 2-2 lin.
Hab. Port Natal.
Forehead impressed with a deep oblong fovea; antennz half
the length of the body, subfiliform, second and third joints
each equal in length to the first, obovate. Thorax scarcely
158 Mr. J.S. Baly on new Species of Crioceride.
broader than long, subcylindrical, sides moderately constricted
just behind their middle, anterior angles acute, upper surface
convex, transversely excavated in front of the base, distinctly
punctured down the middle of the disk and along the anterior
border. Elytra broader than the thorax, not transversely de-
pressed below the basilar space, punctate-striate ; imterspaces
towards the apex furcate.
Lema globicollis, Baly.
L. elongata, parallela, subcylindrica, picea, nitida; ore antennisque
nigris; pectore, scutello thoraceque rufis, hoc elongatulo, basi
constricto, antice globoso, fortiter punctato, utrinque spatio longi-
tudinali impunctato ; oculis nigris, prominulis, intus vix sinuatis ;
elytris cyaneis, dorso planiusculis, infra basin haud transversim
impressis, fortiter punctato-striatis, interspatiis prope apicem
elevatis.
Long. 2+ lin.
Hab. India.
Head short; face triangular, concave between the eyes, the
latter very prominent, globular, shining black ; forehead nearly
impunctate ; antennz scarcely equal in length to half the body,
robust, gradually increasing in thickness towards their apex,
basal joints pitchy. Thorax rather longer than broad, deeply
impressed with subremote large punctures ; a longitudinal space
on either side the disk smooth, impunctate.
Lema Dia, Baly.
L. fulva, nitida ; pectoris lateribus pedibusque obscurioribus, femori-
bus (basi excepta) abdominisque utrinque macula basali nigro-
piceis ; antennis fuscis; thorace fere transverso, lateribus medio
valde constrictis, dorso ante basin transversim sulcato, leevi, disci
medio longitudinaliter tenuissime punctato; elytris obscure ceeru-
leis, violaceo micantibus, infra basin transversim depressis, tenuiter
punctato-striatis, punctis vix ante apicem deletis, interspatiis
planis; limbo (basi excepta) fasciaque centrali fulvis.
Long. 33 lin.
Hab. Ega, Upper Amazons.
Head deeply constricted behind the eyes; face triangular ;
antenne two-thirds the length of the body, slender, filiform,
basal joint fulvous, second very short, third nearly twice as
long as the second, fourth one-third longer than the third, the
rest each equal in length to the fourth. Scutellum triangular.
Elytra much broader than the thorax, suboval, their apex broadly
rounded.
Lema ornata, Baly.
L. oblongo-elongata, subcylindrica, nigra, nitida; capite, scutello
Mr. J.S. Baly on new Species of Crioceride. 159
thoraceque rufo-piceis, hoc vix transverso, cylindrico, lateribus
medio modice constrictis, dorso levi, ante basin transversim sul-
cato; pedibus antennisque pallide fulvis, his apice fuscis; elytris
infra basin leviter transversim depressis, subfortiter punctato-
striatis, interspatiis postice subcostatis, disco antico obsolete
transversim crenulatis, flavo-albis, vitta brevi suturali ante me-
dium abbreviata, plaga communi apicali et utrinque vitta lata a
callo humerali longe ultra medium extensa, postice paullo am-
pliata, obscure ceeruleis.
Long. 3 lin.
Hab. Guatemala.
Lema preclara, Baly.
I. subcylindrica, testaceo-fulva, nitida; capite (antennarum apice
albo excepto), thorace basi et apice, tibiis tarsisque nigris; tho-
race subtransverso, lateribus medio sat valde constrictis, dorso
leevi, ante basin transversim sulcato, disci medio tenuiter lineatim
punctato ; elytris lete purpureis, infra basin profunde transversim
depressis, punctato-striatis, stria nona medio obsoleta, punctis
apicem versus minus distinctis, interspatiis planis, ad apicem con-
vexiusculis.
Long. 3 lin.
Hab, Nauta, Upper Amazons.
Lema Pithys, Baly.
L. elongata, subcylindrica, rufo-fulva, nitida, subtus aureo-sericea ;
antennis gracilibus, filiformibus, fusco-fulvis, apice pallidis ; tho-
race transverso, pone medium strangulato; elytris nigris, infra
basin et intra humeros impressis, disco exteriore pone medium
foveolatis, punctato-striatis, striis postice sulcatis, stria nona medio
late interrupta, callosa, interspatiis antice planis, crebre sed tenuis-
sime crenulatis, postice costatis.
Long. 33-4 lin.
Hab. §. Paulo, Upper Amazons.
Antenne nearly equal to the body in length, second and third
joints equal, each more than twice the length of the second,
fourth equal to the two preceding united, fifth and following
joints nearly equal, each somewhat shorter than the fourth.
Thorax broader than long, deeply constricted behind the middle,
smooth, impunctate. Scutellum triangular, its apex truncate.
Elytra longitudinally sulcate within the humeral callus, trans-
versely excavated near the suture below the basilar space, inter-
spaces strongly costate behind the middle of the elytra; ninth
stria from the suture broadly interrupted, the interrupted por-
tion costiform ; on the inner side of this costa, near its apex, is
a broad shallow fovea which extends across several of the adja-
cent striz.
160 Mr. J. 8. Baly on new Species of Crioceride.
Lema Idalia, Baly.
L. elongata, subcylindrica, rufo-fulva, nitida, subtus aureo-sericea ;
antennis elongatis, filiformibus, fulvo-flavis; thorace vix trans-
verso, lateribus medio sat coarctato, pone medium transversim
sulcato, leevi impunctato ; elytris leete violaceis, nitidissimis, infra
basin et intra humeros impressis, tenuiter punctato-striatis, stria
nona medio late interrupta, interspatiis planis, leevibus, postice
obsolete convexiusculis.
Long. 3} lin.
Hab. Upper Amazons.
Antenne rather shorter than in L. Pithys; otherwise in form
and structure very similar. Thorax less transverse, less deeply
strangulated.
Lema pulchra, Baly.
L. subcylindrica, testaceo-fulva, nitida, subtus aureo-sericea, antennis,
genibus, tibiis tarsisque nigris; elytris infra basin transversim
depressis, punctato-striatis, striis ad apicem subsulcatis, inter-
spatiis postice convexis, stria nona medio late interrupta; lete
violaceis, apice pallide fulvis.
Long. 3 lin.
Hab. Nauta, Upper Amazons.
Face elongate-trigonate ; epistome and labrum clothed with
coarse yellow hairs; eyes obliquely notched; antennze slender,
filiform, nearly as long as the body. Thorax cylindrical, sub-
quadrate, sides moderately constricted in the middle; upper
surface smooth, impunctate, impressed before the middle by a
broad transverse suleation. Hinder thighs slightly thickened,
extending nearly to the apex of the abdomen.
Lema Latona, Baly.
L. rufo-fulva, nitida; antennis pedibusque (femoribus basi et infra
exceptis) nigris; thorace levi, transverso, lateribus medio con-
strictis, dorso ante basin transversim sulcato; elytris infra basin
leviter transversim depressis, punctato-striatis, striis ad apicem
distinctis, interspatiis ad latera et ante apicem costatis; leete vio-
laceis, vitta lata laterali pone medium valde angustata rufo-fulva.
Long. 33 lin.
Hab. 8. Paulo, Upper Amazons.
Head deeply constricted behind the eyes; face triangular,
front impressed with an oblong fovea; antennze moderately
slender, filiform, slightly tapering towards their apex.
Mr. A. G. Butler on a new Species of Cetonia. 161
XX.—Deseription of a new Species of Cetonia in the Collection
of the British Museum. By Anrtuur G. Burirr, F.ZS.,
Assistant Zoological Department, British Museum,
Schizorhina Nortoni.
Clypeus black, elongate, emarginate; thorax shining black, with
black margins ; elytra shining black, sulcated, the strie broad and
clothed with short white hair, apices of elytra coarsely pilose.
Clypeus large, black, smooth, elongate, cylmdric, emarginate
in front, densely punctured, with black, opake, elevated lateral
margins. Lyes lateral, pitchy. Antenne black.
Thorax smooth, very finely punctured, densely near the mar-
gins, shining blacks, with black edges ; anterior portion narrower
than the posterior, and somewhat “depressed at the sides so as to
form an obtuse keel; hinder portion a little narrower than the
elytra, with trisimuate hind margin.
Scutellum very large, smooth, nearly triangular, black.
Elytra smooth and shining, nearly covering the abdomen, a
little narrower at the apex than at the base, with four longitu-
dinal smoothly hollowed strize on each elytron, not reaching the
base, and filled with very short white sete, the two outer striz
meeting at the apex, and filled at their apical termimations with
very long golden-yellow hairs, which extend beyond the ab-
domen.
Body shining black beneath; head, thorax, and sides of abdo-
men clothed with coarse golden-yellow hairs ; sternum produced,
compressed, subtriangular.
Legs black ; femora compressed, those of fore and middle legs
densely punctured and sparsely clothed with short yellow sete ;
femur of hind leg sparsely punctured, the sides not clothed with
hair; tibia of fore leg short, compressed, strongly punctured,
outer edge trispinose, inner edge with a marginal line of minute
yellow hairs, and terminated by a long spine; tibia of middle
leg cylindrical, finely and sparsely punctured, a few yellow setze
extending half along its inner side, apex quadrispinose ; posterior
tibia elongate cylindrical, outer edge coarsely punctured and
bluntly unispinose in the middle, the remainder smooth ; imner
edge clothed with long, straight, yellow hairs; apex quadri-
spinose: tarsi five-jointed.
Length 16 lines.
Habitat. Sydney.
Closely allied to S. Philipsii, Schreib. (Trans. Linn. a yor vi.
p. 193, t. 20. fig. 4, Gory, Monog. Ceton. p. 158, pl. 27. fig. 2),
from which it differs in being “altogether longer aad much
larger, and more quadrate ; the clypeus being longer, narrower,
Ann. & Mag. N. Hist. Ser.3. Vol. xvi. 12
162 M. Hesse on new or rare Crustacea
and destitute of sete ; the thorax much more finely and sparsely
punctured, more depressed at the sides in front, much more
deeply trisinuate behind, shiny black, and destitute of sete ; ;
the scutellum almost imperceptibly punctured ; the elytra more
roof-shaped, the strize more irregular, narrower above than be-
low, filled with shorter and white hair, apical portion filled with
long golden-yellow sete, protruding beyond the abdomen.
The underside is not so densely clothed, and the femora are not
so densely punctured and are much more destitute of hair; the
tibiez much more finely punctured, the middle leg with no me-
dian spine, and the hind leg with little beyond an excrescence.
This interesting species was presented to the National Collec-
tion by Mr. H. Norton, and is one of the most beautiful insects
in the genus.
XXI.—Jnwestigations on new or rare Crustacea of the French
Coasts. By M. Hesse*.
§ NoropreRoPHorvs.
M. Hessz has obtained individuals of all ages of the curious
Crustacean described by him (Ann. Sci. Nat. 5¢ sér. tome 1.)
under the name of Notopterophorus papilio, and now gives an
account of the life-history of the species.
The male is one-third of the size of the female (2 mill. in
length), and has the head large and the body short and stout ;
and the thoracic region, which is of uniform width, does not
present an enlargement for the reception of the ova at its base.
The abdominal portion, which is cylindrical, is also shorter
and more robust than in the female, and the dorsal membranous
expansions are but small at the base, whilst the lobes which
accompany them are very long, and gradually diminish to a
point. In number and position these expansions are exactly
similar to those of the female. The colour is a slightly yellowish
white, through which the large intestinal tube, of a yellow
colour, with red and black points, may be perceived. The eye is
black. The males occur in much smaller numbers than the
females, in the interior of Phallusia canina.
In their early states the Notopterophori resemble Cyclops.
The body is eylindrical, and formed of four thoracic segments
(including the cephalic shield), all of which present posteriorly
two triangular acuminate processes ; of these the points, directed
obliquely, project outwards and on the back, and they are evi-
dently destined to become the membranous expansions of the
* Abstract, by W. S. Dallas, F.L.S. &e., from Aunales des Sciences
Naturelles, 5° sér, tom, ili, (1865) pp. 221-242,
of the French Coasts. 163
adults. The abdomen, antenne, and feet exactly resemble those
of the adults.
At its first escaping from the egg the body of the young ani-
mal is perfectly cylindrical. The antennee are broad, flat, and
rounded at the end; their joints are not distinctly marked. The
first legs are very strong, and composed of four joints, terminated
by a stout claw; then follows a pafr, thinner, of footjaws, ter-
minated by bristles or spines, and beneath these is the rostrum,
formed of two pairs of jaws; beneath these is another pair of
short, slender feet, placed laterally, followed by another larger
pair, curved upwards, and furnished with a comb of strong
spines or rigid hairs. The thoracic feet are already biramose.
Notopterophorus Bombyx, Uesse.
This Crustacean, found in the interior of Phallusia intestinaks,
is doubtfully described as a new species by the author. The
male, as in the preceding species, is one-third of the size of the
female. Its head is much larger; the thorax is broad and short,
and diminishes gradually in diameter to the abdomen ; it is very
retractile, the segments forming it have their margins everted
so as to favour their invagination, and the lower extremity of
the last segment, being capable of vertical elevation, forms a sort
of broad, flat surface, which probably serves as a point of sup-
port for propulsion.
The membranous dorsal expansions appear to be much less
extensive than in N. papilio, and that which is implanted upon
the nape presents at its base an occipital protuberance. The
colour is entirely yellowish white.
The female is elongated, especially in the abdominal region ;
the head is also very long, and the ridge which forms the base
of the occipital membranous expansion is much thicker than in
the other species ; the membranous expansions are also smaller
and thicker. The antenne and feet are more slender, and the
last thoracic segment, which contains the eggs, is remarkable
for a peculiar structure which, when seen in profile, presents
some analogy with that of Doropygus. The female is very little
smaller than that of N. papilio; its colour is pale yellow, with a
rusty red streak in the middle. The eye is red. The eggs are
very dark green, with a transparent limb.
In this species the membranous expansions were more lace-
rated than in the other. The Crustaceans are seen constantly
extending and contracting themselves as if endeavouring to re-
move something, or to force a passage through resisting objects;
their movements of propulsion are aided by the strong claws
and spines with which the feet are armed, and the hooks which
terminate the abdominal appendages enable them to move back-
12*
164 M. Hesse on new or rare Crustacea
wards. M. Hesse thinks that the membranous expansions may
also assist in locomotion; they are moved in the manner of the
wings of a butterfly. They are probably employed as a point of
support and traction, by being applied hermetically, and by pro-
jecting, upon surfaces to which the animals wish to adhere, the
lobes by which they are accompanied. There is also a very fre-
quent and active movement of the mandibles, which might lead
to the supposition that these parasites are rather masticators
than suckers.
The young animal, at the second or third transformation, has
a nearly uniformly cylindrical body, diminishing gradually, how-
ever, towards the posterior extremity, by which it acquires the
appearance of a Cyclops. There is as yet no dorsal appendage ;
but the last thoracic segment has commenced the modification
which fits it for the reception of the ova. The other appendages,
although imperfectly formed, resemble those of the adult. The
body is hyaline, with the eye alone red.
Genus PLevRocryrra, gen. noy.
M. Hesse, in September 1864, discovered under the arch of
the branchial cavity of Galatea squamosa a parasitic Crustacean
belonging to the group of the Isopoda, which he regards as
forming the type of a new genus intermediate between Gyges
and Phrywus. He gives the following characters of this para-
site, for which he proposes the name of
Pleurocrypta Galatea.
“ Male. Body elongate-ovate, divided into seven nearly equal
thoracic segments, of which the first is amalgamated with the
head, which is deeply inserted into it; and the last is attached
to the abdomen, which is triangular and of a single piece. Feet
terminated by a strong hooked and denticulated claw.
“ Female. Body ovate, symmetrical, provided above with very
long ineubatory laminz, which entirely cover the thoracic por-
tion ; abdomen divided into six [five ?] segments, furnished with
simple acuminate branchiz of unequal size; feet terminated by
an oblique, ampulliform joint, having a prehensile orifice.
“ Length of the male 0-001, of the female 0-007 m.”
The head of the male is amalgamated with the first thoracic
segment, forming a sort of buckler, rounded in front and hemi-
spherical above ; the six following thoracic segments are of nearly
equal width, but the sixth is a little narrower and soldered to the
abdomen. ‘The latter is of an elongate triangular form, rounded
at the extremity, and presents no trace of segments, unless it be
that the lateral margins are slightly undulated. On each side
of the French Coasts. 165
of the body, in each segment except the first, the white branch-
ing reproductive organs are seen shining through the skin.
On the lower surface of the body the foremost organs are
two pairs of short antenne placed obliquely on the sides of
the head, and scarcely passing its margin. The superior an-
tenna is shorter than the other, and of three joimts; the inferior
antenna has four. The antenne of each side spring from a
common flat basilar piece.
The buccal apparatus is of a somewhat conical form, with the
apex, bearing the buccal orifice, directed forward. The organs
of the mouth consist of two pairs of hard corneous denticulated
jaws, forming a sort of curved nippers, of four other small foot-
5
jaws, each consisting of three jomts and terminated by a crooked
fo)
claw, and of two flat pointed lamine forming a lower hp. The
thoracic legs consist of five joints, of which the femoral and the
apical are the largest; the latter is terminated by a powerful,
curved, and denticulated claw, which, by folding down upon a
protuberance of the lower surface of the inflated apical joint,
becomes a prehensile organ. The body is of a buff colour, with
the abdomen reddish brown. The intestinal canal is brick-red,
with a fine white line upon it, indicating the course of the in-
terganglionic cord. The surface is covered with short, rigid,
and scattered hairs.
The female is much larger than the male, measuring 7 mill.
in length by 3 mill. in breadth; its form is a regular oval.
The head is hemispherical, and embraced by the first pair of
incubatory plates, of which there are in all four pairs, mereasing
in dimensions as they descend towards the abdomen. The last
two plates are longer than the others, and their posterior margin
is turned down almost perpendicularly upon the base of the
abdomen so as to close the incubatory chamber, At their base
these plates present small niche-like cavities for the reception
of the feet.
The epimeric pieces are alternately large and small, so as to
facilitate the movements of the thorax; from the fifth onward,
they become more and more pointed to the extremity of the
abdomen. Tach of the five segments of the abdomen is pro-
yided on each side with a branchial lamina, which is very deli-
cate and contractile ; these laminz, like the segments to which
they are attached, diminish in size posteriorly.
The ventral surface is nearly flat, or very shghtly concave.
The buccal apparatus is placed close to the membranous anterior
margin of the head; the orifice is pierced im the middle of a
large lip, and from it issue two pointed denticulated jaws, form-
ing a pair of pincers. A little above this orifice are situated
the antenne, which, as in the male, are four in number; the
166 M. Hesse on new or rare Crustacea of the French Coasts.
superior antennz are shortest, and composed of three joints ;
the imferior have a large basal joint, and are terminated by a
cylindrical filament of five joints. The eyes are situated on
small rounded protuberances close to the base of the inferior
antenne.
The body is divided into twelve segments, seven thoracic and
five abdominal; and, from the arrangement of the former, the
animal is enabled to roll itself up into a ball like the woodlice.
Each thoracic segment is furnished with a pair of feet, which
are articulated laterally to the epimeric pieces at the base of the
incubatory lamin ; each foot is formed of five nearly equal
joints, and curved in such a manner that its extremity is di-
rected towards the ventral surface. The apical joint, which is
larger than the others, is inflated above and flat beneath, and
on the first two pairs of feet forms a sort of elastic pad; the
apical joints of the other feet are hollow internally and nar
rowed at the apex, where there is a small contractile aperture
with a raised margin. The anterior half-circumference of this
aperture appears to be of a more solid substance, and, furnished
with denticulations, may render the organ more efficient in
fixation.
The head, abdomen, and whole lower surface of the female
are bright yellow, with the two small lateral protuberances of
the head reddish brown; the incubatory plates are light vinous
grey, and the branchial lamine transparent and bright blue.
Beneath, the segments and epimeric pieces are bounded by
white lines.
The young, on escaping from the egg, are minute and very
active, swimming rapidly by jerks. The head is hemispherical,
rounded in front, and as wide as the first thoracic segment in
which it is immersed ; this is followed by seven other segments,
all of the same size, except the last, which has a small rounded
process in the middle. The eyes, situated on the sides of the
head, are large and hemispherical ; the abdomen is formed of a
single piece, as in the male.
The mouth, which is proboscidiform and retractile, is placed
at the lower extremity of an oval protuberance, which projects
in the middle of the first thoracic segment. The superior an-
tenne are short, stout, and formed of three jomts, the last of
which is slender and cylindrical, and truncated at the extremity,
and the second bears some strong pointed sete. The inferior
antenne are much elongated, and composed of five nearly eylin-
drical joints, which diminish in thickness from the second to
the apex.
The legs, as in the adult, are seven on each side, and all
formed of five joints, of which the last is the most developed.
Mr. H. W. Bates on the Longicorns of the Amazons Valley. 167
The first pair, as in the male, is terminated by a powerful claw;
the last joint of the others is hollow, and terminated by a round
orifice.
The abdomen has on each side a broad, lamellar, bifurcate
false leg, and these appendages are preceded by others which
are also lamellar, pointed and denticulated on the margins ; these
are organs of propulsion, and subsequently form the branchiz.
In some individuals the extremity of the abdomen bears, close to
the anus, two small, flat, rounded, margined laminz. The body
of the young animal is of a pale violet-grey colour, as are also
the eggs; and it is to the latter, seen through the transparent
incubatory laminz, that the suprathoracic pouch of the female
owes its peculiar tint.
The females of these Crustaceans reside in tumours produced
by them on the inner surface of the arch of the branchial cavity
in Galatea squamosa; and the males are found adhering to the
abdomen of the females, frequently to the branchial laminze.
In many cases two males are attached to one female.
XXII.— Contributions to an Insect Fauna of the Amazons Valley.
CotnorreRA: Loneicornes. By H. W. Barss, Esq.
[Continued from p. 113.]
6. Hypselomus paganus, Pascoe.
H. sordide fuscus, nigro obscure irroratus ; thorace dorso tuberoso,
lateribus tuberculo acuto ; elytris humeris subconicis, antice cur-
vato angulatis, cristis centrobasalibus prominulis, obtusis. Long.
7-8 lin. o Q.
Head dingy brown. Antennze blackish brown, bases of the
joints (from the fourth) pallid. Thorax with prominent dorsal
ridge and, on each side, two well-marked tubercles, sides each
with a small acute tubercle ; colour dingy tawny brown, speckled
with dusky. LElytra with projecting shoulders, the projection
somewhat conical, but anterior slope curved or angulated, the
apex formed by a thick black tubercle ; centro-basal ridges pro-
nounced, but not crested with tubercles; surface dingy tawny
brown, speckled or irregularly marked with» dusky. Body be-
neath dingy brown; abdomen black in the middle. Legs
blackish, speckled with tawny; base of claw-joimt reddish :
posterior tibiz in the male dilated at apex. Supplementary
antennal joint of male wanting.
Ega and S. Paulo, Upper Amazons.
7. Hypselomus seniculus, n. sp.
H. parvus, fuscus griseo vestitus, summa fronte acute bituberculata ;
168 Mr. HI. W. Bates on the Longicorn Coleoptera
elytris grosse punctatis, humeris modice productis, obtuse trun-
catis, truncaturee angulo postico acuto; maris articulo 12™° an-
tennarum longiusculo, curvato. Long. 43 lin. ¢.
Head clothed with thick tawny-grey pubescence, vertex
spotted with brown; inner side of each antenniferous tubercle
(3) produced into an acute tooth. Antenne towards the base
grey, spotted with dark brown ; apices of third to eleventh joints
dusky, bases of joints from the fourth testaceous. Thorax con-
vex, unarmed, grey, coarsely punctured (especially on the sides)
and spotted with dark brown. Elytra moderately broad at the
shoulders, the latter not conically produced, but obtusely trun-
cated, with the posterior end of the truncature acute ; surface
thinly clothed with grey pile, and coarsely punctured, simply
convex. Body beneath and legs clothed with tawny-g grey pile,
spotted with blackish, base of ilaae -joint testaceous ; apical half
of posterior tibize strongly dilated ( ¢).
ga. :
8. Hypselomus crassipes, n. sp.
HT, robustus, brunneus ; thorace lateribus patlidis; elytris utrinque
macula oblonga transversa eretacea ; pedibus crassis, nigris, tibiis
posticis maris trigonis. Long. 83 lin. ¢.
Head coarsely wrinkled, black; antenniferous tubercles pro-
duced on the inner side into a stout spine (¢). Antennee
scarcely so long as the body, bases of joints, from the fourth,
pale testaceous ; twelfth joint (3) short and twisted. Thorax
convex in the middle, without distinct tubercles, a short obtuse
tubercle on each side; above dark brown, sides asy tawny
white, traversed by an indistinct dusky stripe. Elytra broad
and but slightly convex ; shoulders conically produced, base on
each side obtusely elevated and very coarsely granulate-punctate,
sides under the humeral projections also coarsely punctured,
rest of the surface faintly punctured; dark brown, base dingy
tawny white ; each clytron beyond the middle Grnanmenten with
a distinct oblong, transverse, chalky spot. Body beneath dingy
tawny; abdomen black in the middle. Legs very stout, black ;
tibiee compressed; hind tibize (¢) dilated from the ager and
obliquely truncated at the apex; claw-joint red.
Tapajos. Apparently alhed to H. fasciatus of Thomson ; but
no mention is made by this author of any peculiar formation
in the legs.
9. Hypselomus simplex, n. sp.
H. subelongatus, brunneo-fulvus, unicolor ; elytris modice attenuatis,
humeris conicis ; antennis gracilibus, ar ticulis basi griseis. Long
63-9 lin. SQ.
Rather more elongate than the allied species; but the elytra
rather convex, and the third antennal joint strongly bent. fed
of the Amazons Valley. 169
dusky. Antenne slender, a little longer than the body in the
female, much longer in the male; basal joint strongly clavate ;
dark brown, bases of the Joints, from the fourth, grey. Thorax
bituberculate on each side the central ridge, dingy tawny brown.
Elytra elongated, gradually and slightly tapering from base to
apex; shoulders conical, base en tach obtusely raised, finely
punctured, colour uniform brownish tawny. Body beneath
tawny brown; abdomen black down the middle. Legs simple,
posterior tibize scarcely dilated in the male ; black, thinly clothed
with tawny pile; claw-joints black.
Kga. tia
10. Hypselomus lignicolor, n. sp.
H. subcylindricus, brunneus; thorace et pectore vittis lateralibus
obliquis, elytris sutura vittisque lateralibus abbreviatis curvatis,
nigris pallide marginatis; elytris compressis, sparsim punctatis,
humeris paulo productis haud tuberculatis. Long. 5; lin, Q.
Head tawny, spotted with dark brown. Antenne as long as
the body (¢), moderately stout, brown, unicolorous. Thorax
unarmed and free from tubercles, surface smooth, brown; sides
each with two oblique, blackish vittee, the upper one margined
with dull ochreous; there is also a short dusky central line near
the middle of the hind margin. Scutellum blackish in the
middle. Elytra nearly cylindrical, sides compressed, shoulders
produced each into a slightly elevated ridge not surmounted by
a tubercle; surface sparingly and finely punctured, brown, su-
ture and several curved streaks on each side blackish, the lateral
streaks margined on the upper sides with pallid brown. Body
beneath brown; breast with oblique stripes, dull ochreous and
blackish ; basal half of abdomen dusky. Legs simple, tawny
brown.
Ega. This species is much more elongate and narrow than
the typical forms of the genus ; it consorts, however, much better
with the Hypselomi than with Hesycha or Oncideres (which com-
prehend elongated forms), having antennze approximated on the
forehead instead of widely separated at their bases. It seems to
be nearly allied to Hypselomus egens, Erichson (Consp. Col. Peru.
p. 148).
11. Hypselomus obscurellus, n. sp.
H. subelongatus, nigricans, griseo variegatus ; antennis articulo ba-
sali apice subgloboso ; thorace postice constricto ; elytris elongato-
trigonis, humeris conico-elevatis, obtusis. Long. 53 lin. ¢.
Head dusky, eyes ample; forehead narrow, coarsely punctured ;
antenniferous tubercles unarmed. Antennze black, base of joints
grey, basal joint very abruptly clavate near the apex, subglobose,
third joint very slightly curved. Thorax cylindrical, constricted
behind the middle, surface very uneven, coarsely wrinkled trans-
170 Mr. H. W. Bates on the Longicorn Coleoptera
versely, dark brown. Elytra moderately elongated, wide at the
base, and narrowed thence towards the apex ; shoulders conically
produced, but apex of cone obtuse and not tuberculated ; surface
very roughly punctured near the base, more finely so towards
the middle, colour dark brown or blackish, thinly variegated
with greyish pile. Body beneath tawny brown; abdomen in the
middle glossy blackish, and sides spotted with black. Legs
blackish, varied with tawny ; hind tibize dilated near the apex ( ¢).
Obydos, Lower Amazons. Similar in size and general figure
to H. Syrinz* (Hesycha syrina, D}. Cat. and French collections),
but differmg in the shape of the basal jomt of antennee and in
fo)
the constricted thorax.
Genus JamestA, Jekel.
Jekel, Journal of Entomology, i. p. 259.
This genus is distinguished from Hypselomus by the basal
joint of the antenne being very gradually thickened from the
base to the apex, not abruptly clavate, and by the third joint
being quite straight instead of crooked. The claw-joints of the
tarsi are quite as long as the three remaining joints taken to-
gether. The species have the same heavy ficure and dull colours;
but the elytra are much more elongated, and less trigonal. The
genus is distinguished also by the large volume and subquadrate
form of the eyes.
There seems to be scarcely sufficient difference to warrant the
separation of Jamesia from Clytemnestra (Thoms.)}, the larger
volume of the eyes being the only apparent definite character.
* Hypselomus Syringe. Subelongatus, brunneus vel nigricans, elytris
utrinque vitta obscura obliqua pallidiore. Caput angustum, fronte
impunctata; tuberis antenniferis intus dente armatis. Antenne cor-
pore paulo longiores, articulis basi pallidioribus, articulo basali paulo
mcrassato. Thorax basi latus, antice angustatus, linea dorsali elevata.
Elytra elongata, postice paulo attenuata, menniiiter punctata, br unnea,
linea curvata mediana obscure fulva; humeris prominulis, in carinam
levem curvatam desimentibus. Corpus subtus fuscum. Pedes fusci,
unicolores ; tibiis compressis. Long. 43-53 lin. ¢ 2. Hab. Rio
Jaineiro.
+ Since the early part of the genus Hypselomus in this memoir was in
print, I have found that Perty and Serville happen to have described the
types of two distinct genera under the respective names of Hypsioma and
Hypselomus. The latter genus is equivalent to Clytemnestra of Thomson,
which therefore becomes!a synonym. M. Thomson, in his later work,
‘Systema Cerambycidarum,’ has adopted this change of nomenclature.
The following rectification of synonymy is therefore necessary :—
Gen. 1. Hypselomus, Perty, Delect. An. Art. Bras.
= Clytemnestra, Thomson, Class. des Cérambycides.
=Jamesia, Jekel, Thomson, Systema Cerambycid. (section).
Gen. 2. Hypsioma, Serville, Ann. Soc. Ent. Fr. iv.
=Hypselomus, Thoms. (Class. des Céramb.) Bates (ut supra)
and authors, nec Perty.
of the Amazons Valley. Val
1. Jamesia globifera, Fab.
Lamia globifera, Fabricius, Syst. Eleuth. 11. 284. 15.
Hypselomus variolosus, Pascoe, Trans. Ent. Soc. n. s. v. pt. 1 (1859).
J. subelongata, sordide griseo-brunnea ; thorace transverse ruguloso
et acute tuberculato ; elytris prope basin tuberculis globosis nigris
politis et postice maculis nigris leviter impressis variegatis ; capite
lateribus parallelis, oculis magnis, subquadratis ; antennis brun-
neis, maris corpore multo longioribus ; pedibus simplicibus. Long.
10 lin.
Not uncommon on dead trees throughout the Amazons region;
also found at Cayenne.
2. Jamesia pupillata, Pascoe.
Hypselomus pupillatus, Pascoe, Trans. Ent. Soe. n. s. v. pt. 1.
Jamesia bipunctata, Jekel, Journ. of Entom. i. 260.
J. subelongata, parum convexa, olivaceo-brunnea, nigro punctata ;
elytris medio utrinque ocellatis ; maris capite infra dilatato, corni-
bus frontalibus magnis acutis porrectis; antennis quam corpus
duplo longioribus. Long. 11 lin. ¢ Q.
Differs from J. globifera chiefly by the more depressed form
of the elytra, and the absence of basal elevation with globular
tubercles. It may readily be recognized also by the eye-like
spot on the disk of each elytron, consisting of a rounded, black,
slightly impressed spot, surmounted by a white speck. The
antenne are much more elongated, and the projecting angles of
the antenniferous tubercles in well-developed males are very
large and acute, and are directed horizontally. The base of the
elytron has a few minute granulations with punctures, and the
rest of the surface is sprinkled with rounded, dark-brown,
shghtly impressed spots, as in J. globifera.
Ega; not uncommon.
Genus Hrsycua (Dj. Cat.), Thomson.
Thomson, Archiv. Entom. i. 187 (1857).
Faimaire, Ann. Soc. Ent. Fr. (1859), p. 523.
This genus was first characterized, in few words, by M. Thom-
son in 1857; but the description subsequently published by
M. Fairmaire defined more accurately its points of distinction.
It agrees with Hypselomus in having the first joint of the an-
tenn abruptly clavate, and the third joint curved; the curva-
ture, however, is much less pronounced than in Hypselomus,
and is sometimes very slight. Its other distinguishing charac-
ters are (1) the elongate, parallelogrammical, and depressed
form of body, (2) the more elongated claw-joint of the tarsi,
and (3) the wide separation of the antenne at their origin.
172 Mr. H. W. Bates on the Longicorn Coleoptera
1, Hesycha Nyphonoides, Pascoe.
Hesycha Nyphonoides, Pascoe, Trans. Ent. Soe. n.s. v. pt. 1.
H, parallelogrammica, depressa, obscure fusca cinereo-fulva varie-
gata; elytris medio fascia undulata, obscura, cinereo-fulva. Long.
6i—s8 Soin 3 gf.
Head dull brown; forehead broad, sparingly punctured ; an-
tenniferous tubercles in the male produced on their inner side
into a stout pomted tooth. Antenne in the male nearly twice
the length of the body, with the apical joint greatly elongated ;
in the female about the len igth of the body, apical joint shorter
than the preceding ; colour dull brown ‘or blackish. Thora
uneven above, sides with a short pointed tubercle; dull hha
speckled with black. Klytra shghtly narrowed from base to
apex, shoulders slightly prominent and surmounted by an ob-
tuse shining tubercle; surface even, thickly but finely punc-
tured, dull brown, covered with dingy tawny confluent spots,
and crossed beyond the middle by a zigzag fascia of a little
paler hue. Body beneath and legs ‘blackish or dull brown.
Common on branches of dead trees at Kga. There are two
closely allied species in collections from the interior of French
Guiana*,
* Hesycha jaspidea, u. sp. H. Nyphonoidet simillima, robustior, maris
elytris postice magis angustatis et fronte valde cornuta. Obscure
fusca; thoracis lateribus utringue tuberis duobus obtusis armatis.
Elytra humeris prominentibus, basi rugoso-punctata et inaequalia
fusco-nigra, maculis sordide fulvis sparsis quarum tribus majoribus
medianis in fasciam abbreviatam conjunctis. C orpus subtus fulvo to-
mentosum. Antenne valde elongatie, articulis basi griseis. Long.
Slin. ¢. Had. In Cayenna interiore (Dom. Bar).
Hesycha liturata, n. sp. Minor, brunnea, elytris litura tenui obliqua
albicante. Caput fuscum, fronte pune tata, tuberculis antenniferis
utroque sexu intus acutis. Antenne brunnez, maris corpore paulo
longiores. Thorax quadratus, lateribus tuberculo distineto subacuto,
supra brunneus vittis tribus nigris, lateribus cierascentibus. Elytra
postice paulo angustata, apice oblique breviter truncata, humeris vix
produetis, obtusis; dorso punctata, brunneo et fulvo variegata, infra
humeros (cum prothoracis et pectoris lateribus) nigricantia, “apud me-
dium litura tenui valde obliqua albicante. Corpus subtus et pedes
brunneo tomentosa, Long. 5-6 lin. ¢ 2. Had. In Cayenna (Dom.
Bar).
The following species belongs also to this genus, from its linear sub-
depressed form and the somewhat wide separation of the antenne at their
bases :—
Hesycha xylina, n. ‘Sp. Elongata, sordide brunnea ; ely ie rugoso-pune-
tatis, fusco et griseo strigatis, humeris subuneinatis. Caput fuscum,
fronte grosse sparsim punctata, tubereulis antenniferis intus dente va-
lido eurvato armatis( ¢). Antenne valde elongate, brunnew, apice
pallida, articulis (4 tertio) basi testaceis, articulo 12™0 acuto, curvato.
Thorax supra ineequalis, mermis, brunneus, Elytra valde elongata,
of the Amazons Valley. 17
co
2. Hesycha maculosa, n. sp.
IT. clongata, convexiuscula, fusca, maculis numerosissimis partim
confluentibus fulvis ; vertice nigro trilineato ; thorace nigro macu-
- > D ? o
lato. Long. 8h lin. 3 9.
Head dusky, front channeled down the middle, punctured ;
eyes rather elongated, margined on the inner side narrowly with
tawny ; vertex tawny, marked in the middle with three parallel
black lines; antenniferous tubercles produced into a short acute *
tooth on the i inner side, longer in the male than in the female.
Antenne longer by one half than the body in the male, and the
terminal joint very slender and much longer than the preceding ;
in the female a little longer than the body, with the terminal
joint shorter than the preceding ; basal jomt abruptly clavate,
third joint scarcely perceptibly curved ; colour blackish. Thorax
quadrate, surface uneven, with several impressed curved lines
and raised interspaces, sides behind the middle with an acute
tubercle; colour tawny, marked with two short black lines in
front in the middle and a spot behind them, and four spots on
each side of the disk. Scutellum black. Elytra elongate and
rather convex, slightly tapering; shoulders prominent, and sur-
mounted by a glossy black tubercle; surface quite even and
moderately punctured, dark brown, covered uniformly with a
multitude of tawny specks, mostly confluent. Body beneath
tawny. Legs blackish.
ga.
3. Hesycha cretacea, 0. sp.
H. oblongo-elongata, subdepressa; elytris maculis numerosis fulvis
maculaque magna laterali eretaceo-alba. Long. 8 lin. 9.
Head grey, margins of eyes with tawny lines, front punctured;
eyes elongated; antenniferous tubercles acute on their inner
side, vertex with three short black streaks. Antenne a little
longer than the body, dark brown; basal joint clavate, third
joint very slightly curved. Thorax quadrate, sides each with
two large obtuse tubercles, surface with transverse furrows,
tawny mixed with grey, and spotted with black. Scutellum
black, margined with grey. Elytra oblong, a little dilated be-
yond ‘the middle, slightly convex, shoulders moderately promi-
nent ; with irregular clusters of punctures arranged in lines,
black, covered with pinkish-tawny spots, partly confluent, and
LL ———
parum convexa ; humeris productis, antice curvatis, postice tuberculo
nigro armatis, quasi uncinatis; supra grosse punctata, punctis partim
confluentibus, sordide brunnea, strigis pallidis et fuscis variegata.
Corpus subtus brunneum. Pedes fusci. Long. 43 lin. 3. Hab. Rio
Janeiro, 1 D, Squires capta.
174 Mr. H. W. Bates on the Longicorn Coleoptera
having in the middle on each side a large chalky-white spot.
Body beneath dull chalky white; breasts with pinkish streaks,
and abdomen spotted with black. Legs black, thinly clothed
with grey pile. :
Ega; rare. This handsome species, hke the preceding (H.
maculosa), approaches Oncideres in many of its characters, espe-
cially the elongate eyes, subconvex form of body, and scarcely
curved third antennal joint; but it lacks the massive head,
cylindrical form of body, and short transverse thorax of Oncz-
deres, and therefore must be classed with Hesycha.
Genus TracHysomus, Serville.
Serville, Ann. Soc. Ent. Fr. iv. (1835).
(Char. emend.) Buquet, Ann. Soc. Ent. Fr. 1852, p. 345.
This remarkable group is distinguished from the allied genera
chiefly by the elytra being disfigured by tubercular excrescences,
and by the antennz being composed of short joints reaching
only three-fourths the length of the body. The head is mode-
rately narrow, the eyes oblong (not narrow and elongated as in
Oncideres), the basal joint of the antenne very abruptly clavate,
the third joint very slightly curved, the thorax subcylindrical,
and the claw-joint of the tarsi shorter than the remaining joints
taken together. The species are found closely clinging to thin
woody stems of plants, and strongly resemble portions of the
stems distorted by glandular prominences or galls.
Trachysomus Santarensis, 1. sp.
T. Trachysomo fragifero (Kirbii) valde similis, differt colore ochra-
ceo- vel rufo-fulvo ; thorace supra ochraceo ; elytris juxta scutel-
lum utrinque spinis quatuor acutis, fasciculis singulis pilorum
subapicalibus nigris linea curvata nigra communi connexis.
Long. 7} lin.
This is so closely similar in form of body and _ tubercular
excrescences to the South-Brazihan 7. fragifer, that it can
scarcely be considered more than a local form of the same stock.
It is a little broader and more robust, the thorax is less uneven
on the disk, and is there of a bright yellowish-tawny colour.
The two tubercles on each elytron, near the scutellum, are longer
and more acute. The elytra are of a nearly uniform reddish or
orange-brown hue; the subapical fascicle of hairs is a little
further removed from the apex and margin of the elytra; it is
connected with the corresponding fascicle posteriorly by a curved
black line, and a large portion of the disk behind each basal
—exerescence 1s quite smooth.
Dry woods near Santarem,
of the Amazons Valley. 175
Genus OncrpEREs, Serville.
Serville, Ann. Soe. Ent. Fr. (1835) iv.
The chief characters of this, the typical genus of the group,
are furnished by the elongate-oblong or cylindrical form of body;
the broad head and convex occiput, with consequent wide sepa-
ration of the antennz at their bases; the elongated eyes; the
clavate shape of the basal antennal joint, and straight form of
the third joint; the short transverse thorax ; and, “lastly, the?
great length of the claw-joint of the tarsi, which exceeds that of
the three remaining joints taken together.
The species are all found on the branches of trees, which they
amputate from the living tree by gnawing deeply into the bark
and wood, making a ring-like incision, until the bough breaks
off by its own weight. ‘T have often seen boughs thus severed
from green and living Cajt trees, and hence discovered that the
best means of finding the insects was by examining the ampu-
tated portions lying on the ground in woods or the thinner parts
of the forest. The object of the severance is apparently to
create a supply of dead wood in which to deposit their eggs and
rear the larvee.
1. Oncideres Callidryas, un. sp.
O. minus convexus; thorace griseo-tomentoso ; elytris basi minute
granulatis, medio confertim punctatis, nigris, guttis numerosis-
simis carneo-griseis. Long. 103 lin. 3 Q.
Head much narrower than the middle part of the thorax,
clothed with pinkish-tawny pile; forehead plane, punctured ;
antenniferous tubercles (¢) on each side armed with longish
acute teeth directed forwards ; eyes oblong. Antennz about the
same length as the body in the female, twice the length in the
male, black. Thorax with transverse depressions, sides each
armed with a strong conical tubercle, clothed with hoary-grey
pile. Scutellum and basal margin of elytra hoary grey. Elytra
less cylindrical and convex than im the more typical species ;
shoulders prominent and surmounted by a retrocurved tubercle,
base and shoulders thickly and finely granulated, middle part
simply but thickly punctured, Papel becoming finer poste-
riorly, and disappearing before the apex ; colour black, sprinkled
throughout with small grey or pinkish- -grey spots, some very
minute, others larger; near the middle of each side the spots
are whiter, and tend to aggregation. Body beneath hoary
white. Legs black, thinly clothed with grey pile.
Para, banks of the T apajos, and Ega; one pair taken in co-
puld on a branch of a felled tree at Paré. The elytra are much
more thickly spotted in the Kga examples than in those from
Para and the Lower Amazons.
176 Mr. O. W. Bates on the Longicorn Coleoptera
Oncideres Satyrus, n. sp.
O. cylindricus, fulvo-brunneus; elytris guttis albis paucis sparsis,
basi tuberculis nigris; antennis validis; thorace basi valde con-
stricto. Long. 10-12 lin. ¢ @.
Head in the @ much narrower than the thorax, in the ? as
wide as the widest part of the thorax, with broad plane front,
colour tawny brown, a black stripe below each eye. Antenne
ubout the length of the body in the female, a little longer in the
male, with the apical joint twice the leneth of the preceding ;
they are robust in both sexes, but the four basal joints are
thicker in the g than in the ?; colour black. Thorax with
transverse depressions; a conical tubercle on each side, and
much constricted behind the tubercle ; brownish tawny, with a
fine, black, central, transverse line. Hlytra cylindrical, brownish
tawny, sprinkled with a small number. of minute white spots ;
base and shoulders with a few polished rounded tubercles ; rest
of surface impunctate, smooth. Body beneath and legs thickly
clothed with tawny pile; sides of breast chalky white.
Para. Closely allied to O. vomicosus, Germar (Ins. Nov. 482),
but differing greatly in the maculation of the elytra, the spots
being small, few in number, and all distinct from each other.
3. Oncideres fulvus, n. sp.
O. oblongo- -subeylindricus ; thorace postice haud constricto, guttis
higtis quinque discoidalibus in linea transversa dlispositis, tuber-
culo parvo laterali; elytris modice elongatis, valde convexis, leevi-
bus, guttis parvis ‘albis Sparsis, prope “basin tuberculis utrinque
cirea duodecim nigris. Long. 11 hn. 9°.
Closely resembles O. Satyrus ; but the body is proportionately
shorter and broader in the female than in the corresponding sex
of that species ; the thorax is shorter, and shows no constriction
near the base; the elytra are uniformly convex and impunctate,
and there are very few tubercles near the base, only two con-
spicuous ones on each side of the scutellum, and a small number
under each shoulder. The colour is entirely ochreous tawny,
with the exception of five small spots placed in a transverse row
across the thorax, the black elytral tubercles and a small number
of widely separated, but tolerably uniformly distributed, white
specks over the elytra. The antennz are somewhat darker, and
and there is a very distinct oblong chalky spot on each side of
the breast.
Tapajos.
4. Oncideres Diana, Olivier.
Lamia Diana, Oliv. Ent. 67. p. 107. f. 168.
O. subcylindricus, griseus ; elytris quarta parte basali dense ac mi-
of the Amazons Valley. wir
nute tuberculata, parte apicali lineis tenuissimis fureatis nigris,
medio guttis sparsis nigris; thorace linea transversa nigra: foe-
minze capite lato fulvescente ; maris capite angusto, fusco, inermi.
Long. 8-11 lin. ¢ Q.
This species is distinguished by the basal portion of the elytra
being thickly covered with small glossy-black tubercles, of which
one at the hinder part of the humeral prominence is much
larger than the rest. The tuberculated area ceases abruptly
behind, and the disk of the elytra has only a very few scattered
and slightly elevated black specks, which towards the apex sub-
side into simple spots, not raised at all from the smooth surface.
The general colour is pale ashy grey (white beneath); the apical
part of the elytra has a few fine black lines in the form of a
double or treble fork joined at the base. The male differs
greatly in width of head from the female, but the antenns
scarcely differ in proportionate length or stoutness; they are,
however, more nearly approximated at their bases by one-half
in the male than in the female, which gives to a male insect an
appearance quite foreign to the genus. The male specimen be-
fore me has a finely reticulated black patch across each elytron
at the tips of the forked lines, of which there is only a trace in
one of the female examples.
Para, and at Santarem on the Tapajos.
5. Oncideres crassicornis, n. sp.
O. subcylindricus, postice utroque sexu attenuatus, fulvo-brunneus ;
elytris basi tuberculis diversis sparsis instructis, postice punctis
impressis rufescenti-brunneis in lineis furcatis ordinatis ; maris
antennis basi valde incrassatis, capite bicornuto. Long. 9-10
lm. G2.
Head not much wider in the female than in the male, brownish
tawny, with the usual black stripe below each eye; antenniferous
tubercles in the male dentiform on each side. Antenne dark
brown, simple in the female, one-half longer than the body in
the male, with the basal and third joints much thickened, espe-
cially the latter. Thorax impressed transversely, and furnished
on each side with a tubercle; colour brownish tawny. LElytra
narrowed to the tip in both sexes, tawny brown, inclining to-
wards ashy near the middle ; the basal part raised in the middle,
and studded with a moderate number of scattered tubercles, dif-
fering greatly in size, and all glossy black; from the middle to
the apex there is a number of shallow punctures covered each
with a reddish-brown spot and arranged in forked lines. Body
beneath and legs clothed with tawny-brown tomentum.
Ega, and banks of the Tapajos.
Ann. § Mag. N. Hist. Ser, 3. Vol, xvi. 13
178 Mr. H. W. Bates on the Longicorn Coleoptera
6. Oncideres dignus, un. sp.
O. cylindricus, fuscus; thorace tuberculis quinque in linea trans-
versa ordinatis ; elytris prope basin tuberculis magnis globosis
utrinque sex nigris, postice guttis numerosis albis. Long.10 lin. 3.
Head ( 3) moderately narrow; forehead very narrow, being
encroached upon by the voluminous eyes, which are oblong and
reach very nearly to the extremity of the muzzle; antenniferous
tubercles unarmed ; colour dark brown. Antenne nearly twice
the length of the body, black, basal joint gradually thickened
from base to apex, rest of the antenne tapering to the tip.
Thorax longer and narrower than in the typical species of Onci-
deres ; lateral tubercles small, obtuse, and black; in a line with
them is a row of five similar glossy-black tubercles lying across
the middle of the thorax; colour dark brown. Elytra cylin-
drical, clear dark brown, impunctate; middle of base with six
very prominent glossy-black tubercles, arranged in two rows ;
besides these, there are ten or twelve smaller tubercles on each
side, three of which are on the shoulder: the rest of the elytra
smooth, and ornamented with a number of small clear white
spots, distributed regularly and widely apart over the surface.
Body beneath and legs dark brown.
This handsome species was very rare, at Ega, Upper Amazons.
7. Oncideres pulchellus, n. sp.
O. minor, cylindricus, griseo-brunneus; elytris cinereo maculatis,
dimidio basali tuberculis rotundatis, dimidio apicali maculis im-
pressis, nigro-nitidis. Long. 63 lin. @.
Head and thorax of same breadth; head ashy brown, with a
streak down each side of the front tawny; buccal organs and
cirewt of the mouth red. Antenne a little longer than the
body, dark brown. Thorax ashy brown, with three shining-
black tubercles in a triangle on the disk, and two smaller ones
on each side, the outermost of which is in the position of the
ordinary lateral tubercle. Elytra cylindrical, obtuse behind,
ashy brown, varied with a small number of equal-sized and equi-
distant pale ashy spots, and with a number of scattered shining
round spots, those over the basal half covering large rounded
tubercles of small elevation, and those towards the apex shallow
impressions; the tubercles are not crowded near the base or
shoulders, but are widely dispersed. Body beneath and legs
light brown; sides of breast with an ashy patch.
Kga; rare.
8. Oncideres Cephalotes, n. sp.
O. magnus, robustus, convexus, postice attenuatus, cinereo-brunneus;
elytris prope basin dense, pone basin sparsim tuberculatis, tuber-
of the Amazons Valley. 179
culis ovatis, obliquis et postice elevatis ; thoracis tuberculis latera-
libus elongatis, fronte magna, latissima, nuda, punctulata. Long.
15 lin., lat. capitis 44 lin. 92.
Head brown; front naked, coriaceous, punctured, black ; eyes
moderate, reaching little more than halfway down the forehead;
vertex very convex. Antenne rather shorter than the body ( ? ),
tapering to the apex, basal joint curved; colour brown. Thorax
twice as broad as long, a little narrowed behind the lateral tu-
bercles, which are long and spiniform ; surface dull ashy brown,
with a central transverse black line. Elytra massive, narrowed
to the apex, convex, especially in the middle of the basal part on
each side ; shoulders prominent and oblique, with a conspicuous
tubercle at their hinder angles; colour ashy brown, paler near
the middle, and covered with small, oblong, raised, scale-like
tubercles, which are very crowded and strongly elevated at their
posterior ends near the base, much scattered and very slightly
elevated near the middle, and arranged in rows, simply as spots,
near the apex. Body beneath ashy white; legs ashy brown.
Ega*,
Genus Euprsmvs, Serville.
Serville, Ann. Soc. Ent. Fr. iv. (1835) p. 82.
This well-marked genus resembles Oncideres in its cylindrical
form of body. Its distinguishing character is derived from the
bulbous ovate shape of the third antennal joint in the male.
* The following new species have lately been received from entomological
travellers in South America :—
Oncideres limpidus. Cylindricus, fusco-nitidus ; elytris fulvo-ochraceo
irroratis. Caput (3g) modice angustatum, fronte punctata, ochracea,
vitta infraoculari nigra; tuberculis antenniferis intus prominulis,
acutis. Antenne corpore longiores, nigrz nitide; articulo basali
distincte clavato, articulis tertio et quarto infra dense ciliatis. Thorax
postice angustatus, tuberculis lateralibus modice productis, nigris ;
supra ochraceo-brunneus, linea nigra transversa, ante medium fascia
rufo-fulva. Elytra cylindrica, fusco-nitida maculis numerosissimis
discretis tomentosis ochraceo-fulvis ; juxta basin tuberculis globosis
paucis; deinde leviter granulata, humeris confertim tuberculatis.
Corpus subtus fulvo-tomentosum. Pedes nigricantes, femoribus
fulvo tomentosis. Long. 10 lin, ¢. Hab. in Bahia Brasiliz, a Dom.
Reed lecto.
Oncideres Bouchardii. Cylindricus, cano-griseus; elytris nigro pune-
tatis et maculis majoribus rotundatis fulvis sparsis. Caput latum,
griseum, maris paulo angustius, tuberculis antenniferis intus vix pro-
minulis. Antenne grisee ; articulo basali gradatim incrassato, nigro,
maris valde rugoso. Thorax griseus, linea transversa nigra. Elytra
convexa, vage punctata, cano-grisea, maculis rotundatis carneo-fulyis
conspersa, punctis nigris; prope basin tuberculis numerosis globosis.
Corpus subtus pedesque cano tomentosa. Long. 10-11 lin. ¢ 2.
Hab. in Sta. Martha Noy Granatee, a Dom. Bouchard copiose missus,
13*
180 Mr. H. W. Bates oa the Longicorn Coleoptera
The females of some of the species resemble Oncideres very
closely; and almost the only feature by which their generic
position may be recognized is the peculiar dark patch, streaked
with paler colours, which exists on the apical part of the elytra
of all the species. The head is broad, very little broader in the
females than in the males; but the forehead is not so plane or
so much elongated as in Oncideres. The basal joint of the an-
tenn forms a smooth ovate club ; the thorax is relatively a little
longer than in Oncideres ; the elytra are free from ridges and
tubercles, and are obtusely rounded at the apex; the claw-joint
of the tarsi is moderately clongated, and is about equal in length
to the remaining three.
1. Eudesmus rubefactus, n. sp.
Hf. cylindricus, convexus, rufescens; thorace nigro-lineato ; elytris
dimidio basali grisescente, apice utrinque macula magna ovata
saturatiore strigis nigris et griseis ornata. Long. 73-9 lin. ¢ 2.
Head reddish tawny, vertex streaked with black ; front plane,
coarsely punctured, dingy grey ; eyes oblong, one-half the length
of the front ; antenniferous tubercles in the male acute on their
inner sides. Antenne about the length of the body, reddish
tawny; apices of joints, from the fourth, blackish. Thorax
cylindrical, of same width as the head, very uneven, especially
on the sides, where the inequalities rise to broad, obtuse tuber-
eles; colour pinkish red, centre with two black lines continuous
with those on the vertex, sides each with two or three much-
broken and oblique lines. Scutellum and basal margin of ely-
tra reddish, spotted with black. Elytra cylindrical, convex,
abruptly declivous near the apex ; surface uneven, with faintly
raised lines, thickly punctured, especially towards the base,
basal half occupied by a large, triangular, common, dingy-grey
patch ; on this follows a belt of pale greyish red, which broadens
greatly on the lateral margins; the apical portion of each elytron
is occupied by a dark, neatly limited, oval patch, streaked longi-
tudinally with black, tawny red, and grey. Body beneath and
legs reddish brown; breast ashy in the middle.
ga, clinging to dead boughs of trees; rare.
2. Eudesmus caudalis, n. sp.
£. cylindricus, depressiusculus, cinereo-brunneus ; thorace postice
fusco notato; elytris dimidio basali griseo-fusco, apice utrinque
macula magna ovata nigricante fulvo strigata, medio cinereo fas-
ciata. Long. 53-6 lin. ¢ Q.
Very closely allied to HE. rubefactus, and scarcely differing in
the disposition of the colours and markings of the elytra. The
latter, however, are much more depressed ; and the insect is of a
of the Amazons Valley. 181
dull ashy-brown hue, and much narrower and smaller. The
forehead is uneven, punctured, and of a dull slaty hue; the
third antennal joint m the male is much less swollen than in
Ei. rubefactus, and therefore more elongate, and fusiform rather
than ovate in shape. The thorax is uneven and obtusely tuber-
culated on the sides, but is destitute of longitudinal lines, except
two very short ones near the base. The elytra are of the same
grey leaden hue over their basal halves, and have a pale belt
beyond the middle; but the latter does not expand on the mar-
gin. The dark apical streaked spot has an ashy transverse
streak across the middle.
Also found at Ega.
3. Eudesmus posticalis, Guérin.
Eudesmus posticalis, Guérm-Méneville, Icon. Regne Animal, p. 248.
E. cylindricus, subdepressus, brunneus; thorace dorso valde in-
eequali immaculato, tuberculis lateralibus parvis; elytris medio
fascia obliqua grisea, deinde brunneis griseo et griseo-brunneo
strigatis, ante apicem signatura nigra griseo marginata ; antennis
brunneis, articulis (duobus basalibus exceptis) basi testaceis ;
maris articulo tertio valde inflato, ovato. Long. 65 lin. ¢.
“Tun eris-brunatre couvert @’un duvet trés-court et trés-fin
@une couleur cendrée, surtout en dessous, sur les cdtés du
corselet et au milieu des élytres, ot. ce cendré blanchatre forme
une bande crochue en arriére, terminée en pointe prés de la
suture et précédant une tache arrondie dun brun plus foneé,
en arriére de laquelle on voit une petite tache allongée blanche
et deux ou trois petites lignes noirdtres. Antennes d’un gris
brun, avec la base du troisiéme article et des suivants d’un jaune
roussatre pale, une petite pointe avancée la saillie du front sur
laquelle s’insérent les antennes. Pattes courtes et fortes, un
gris brun dessus, cendrées en dessous. Long. 14, lat. 5 mill.—
Brésil intérieur.” (Guérin-M neville, /. c.)
My example was found at Ega.
4. Hudesmus sexvittatus, n. sp.
£. elongatus, depressus, fulvo-brunneus; thorace supra vittis sex
nigris ; elytris ultra medium dilatatis, plaga laterali infra humeros,
linea basali strigisque ante apicem fuscis, vitta curvata laterali
cinerea; fronte abbreviata, oculis magnis subconvexis. Long.
6Llin. Q.
Head slightly convex on the forehead, with short muzzle ;
eyes very large, broad, and somewhat convex, reaching very
nearly to the edge of the epistome; vertex bright tawny, and
marked with a semicircular figure of a blackish-brown hue,
Antenne rather longer than the body (2) and stout, ochreous
182 Mr. P. M. Duncan on some Fossil Corals
brown, base of joints (from the fourth) pallid. Thorax convex,
but depressed near the hind margin; lateral tubercle small,
conical; colour above bright tawny, with six blackish-brown
vittee; sides ashy, with a broader and paler dusky stripe. Scu-
tellum pale tawny ochreous. LElytra dilated a little behind the
middle, depressed, and thickly punctured (except towards the
apex), rusty tawny, with a few short ashy streaks and a number
of dark-brown strigz a little behind the middle, the innermost
of which runs near the suture to the apex: the basal half of the
suture is broadly margined with dusky, and there is a short
blackish stripe on each side near the scutellum, and a broad
patch of similar hue beneath each shoulder, on the upper edge
of which is an ashy streak, which continues in a curved line to
the lateral margin, and then to the apex. Body beneath ashy ;
sides of breast and abdomen dark brown. Legs reddish ; femora
and tibize each with a blackish ring round the middle.
I met with the female only of this remarkable species, which
differs so much from the other Hudesmz in the shortness of the
muzzle. If the male, when discovered, should be found not to
possess the swollen third antennal joint, the species will have to
be removed from this genus. It was found at Ega.
[To be continued. }
XXIII.—A Description of some Fossil Corals from the South
Austrahan Tertiaries. By P. Martin Duncan, M.B. Lond.,
Sec. Geol. Soc.
[Plate VIII]
THE corals about to be described were derived from the same
Tertiary beds which yielded the species noticed in the ¢ Annals’
for Sept. 1864*, A new genus is represented by three well-
marked species ; the well-known genus Sphenotrochust has two
species in the collection ; and the genus Anéilliat, which attains
so great a development in the Nivaje shale of San Domingo, is
represented by a very interesting new species.
Last of Species.
1. Sphenotrochus australis, Woods & Duncan, sp. nov.
2. emarciatus, sp. NOV.
* The Rev. J. Woods, who collected those formerly described, classes
the various beds of Muddy Creek, Geelong, and the Murray beds as the
“Hamilton ”’ Tertiaries. I have to thank him for the specimens now
determimed and for others which require some further study before their
publication.
+ Edwards and Haime, Hist. Nat. des Coralliaires, vol. ii. p. 65.
{ P. Martin Duncan, Quart. Journ. Geol. Soc. Feb. 1864, p. 28.
from the South Australian Tertiaries. 183
3. Conosmilia elegans, gen. et sp. nov.
4, anomala, gen. et sp. nov.
5. striata, gen. et sp. nov.
6. Antillia lens, sp. nov.
1. Sphenotrochus australis*, n. sp. Pl. VIII. fig. 1 a, 8, ¢, d.
The coral is very compressed, especially inferiorly, where on
either side of the centre of the base a process passes downwards,
giving a “fish-tail” appearance. At the calice the compression
is less; but the great axis is at least twice the length of the
smaller. The coral is longer than its breadth. The costz are
broad, somewhat wavy, and are separated by well-marked lines:
those of the inferior appendages arise from the extremities of
the processes, and pass upwards and inwards; and the lateral
costze, wavy below, become straight above. All are plain. The
wall is much thicker at the ends of the long axis than elsewhere
(in sections). The calice is not shallow, is elliptical, and pre-
sents, deeply seated, a long lamellar columella, which is joined
to the primary and secondary septa by processes. The septa
are well developed and plain; they are not exsert, but pass
straight downwards and inwards towards the columella ; they do
not correspond with the cost, but with the intervals between
the costz, and they number thirty-two. There are three cycles,
with the orders of a fourth, in two systems.
Height 4 inch, breadth + inch; small diameter, halfway,
zp inch.
Hamilton, Victoria, South Australia. Coll. Geol. Soc.
2. Sphenotrochus emarciatus, n. sp. Pl. VILL. fig. 2 a, 6, ¢, d.
The coral is generally much compressed, especially inferiorly,
where two lateral processes give a notched or emarciate appear-
ance to the base. Superiorly the relation of the long to the
short axis is at least 2 to]. The coral is short and broad ; the
base is nearly as wide as the calice is long. The cost are large
and plain, and are separated by well-marked lines : the coste of
the appendices are the largest ; they pass upwards to the calice,
and all are more or less wavy, the central widening out near the
calicular margin. The calice is shallow and elliptical. The
columella is not long, and, from being jomed to the primary
and secondary septa by processes which are rounded above, is
confused in its appearance. The septa are in six systems of
three cycles; they are wider at the wall than elsewhere, are
granular, and those of the third cycle are much smaller ‘than
* Mr. Woods gave the name; but I have not had the opportunity of
seeing his MS.
184 Mr. P. M. Duncan on some Fossil Corals
the others. All the septa correspond to the depressions between
the costee.
Height + inch, breadth 5%, inch.
Hamilton, Victoria, South Australia. Coll. Geol. Soe.
ConosMILIA, noy. gen.
Coral simple, pedicellate, conical. Columella formed of one
or more twisted laminee, which extend from the base upwards.
Kndotheca scantily developed. Septa apparently with simple
margins, and variable in regard to the number of the primary.
3. Conosmilia elegans, n. sp. Pl. VIII. fig. 3a, db, ¢.
The pedicel is large. The coste, equal, sharp, and prominent
at the base, become ‘broad, flat, and granular above, where the
are separated by very faint lines. The columella is formed by
one twisted lamella, and occupies much space. The septa are
in eight systems of three cycles. There are eight primary septa
which reach the columella; the secondary are smaller and reach
midway; and the tertiary are very small. The septa are nearly
plain, are as thick at the columella as at the calicular margin,
and appear to arise between the costa. The calice is nearly
circular.
Height ;8, inch; breadth of calice +5 inch.
Geelong, Victoria, South Australia. Coll. Geol. Soc.
4. Conosmilia anomala, n.sp. Pl. VIII. fig. 4a-e.
The coral is tall in relation to its small pedicellate base. The
costee are not prominent, but are traced by the faint lines which
separate them, and by the fine herring-bone pattern which marks
each of them. The columella is large, strong, and consists of
two twisted riband-shaped lamine. The septa are in eight sys-
tems of three cycles; the laminze are sparely granular, and the
primary are attached to the columella by processes. The se-
condary are smaller than the primary, and their inner edge is
wavy; the tertiary septa are small. The septa arise between
the coste. The endotheca is sparely developed. The wall is
very thin. ‘The calice is slightly elliptical.
Height 5% inch, greatest breadth -%, inch.
Hamilton, Victoria, South Australia. Coll. Geol. Soc.
5. Conosmilia striata, n. sp. Pl. VIII. fig. 5 ac.
The coral has a very narrow base, and does not expand gra-
dually. The costz are very broad, have marked lines between
them, are very flat, and have wavy transverse markings like
those of a pellicular epitheca. Septa in six systems of three
cycles; the primary, which are granular, reach the columella,
Jrom the South Australian Tertiaries. 185
which appears to be formed by one twisted process. The septa
arise anes the coste. The calice is more or less elliptical.
Height = inch, greatest breadth ;2; inch.
Geelong, Victor South Aogielia, Coll. Geol. Soe.
6. Antillia lens, n. sp. Pl. VIII. fig. 6 ae.
Coral in the shape of a cyclolite Fungia. The base is circular
in outline, nearly flat, the concavity being very slight. The
epitheca is pellicular and fait. The costz are seen as radiating
flat elevations, those corresponding with the smallest septa
being the smallest. The margin of the base presents slightly
exsert, equal processes, which are the septa. The upper surface
of the coral is convex and nearly hemispherical, the depression
for a small essential columella, formed by processes from the
base and septal ends, being slight. The septa are in six systems
of four cycles; the primary and secondary septa are equal,
and the tertiary are nearly as large; those of the fourth and
fifth orders are somewhat less: all are very convex superiorly,
and less so and nearly straight externally. The lamine are
thin, and are very strongly marked by sharp ridges, which,
radiating from the basal part of each septum, are more or less
parallel, and give at the free margin a laterally dentate appear-
ance. The appearance is less in the smaller septa. There is
often a paliform process on the larger septa near the columella ;
and the terminations of the ridges give the dentate character to
the free margin of the septa. The endotheca is scanty, stout,
and inclined.
Breadth 53; inch, beight 2, inch.
Hamilton, Victoria, South Australia. Coll. Geol. Soe.
Remarks on the new Genus and Species.
There is much that is very imteresting in these Australian
forms; they are so novel to those who are acquainted with the
coral-fauna of the past m Kurope and America, and moreover
they present structural peculiarities which remove some broad
lines of demarcation between some of the principal families in
our classification.
The new genus Conosmilia possesses the twisted riband-shaped
columella of the subfamily Caryophyllacez, the endotheca and
septal margin of the Trochosmihacez, and the irregular septal
arrangement which was so common in the corals of the Oolitic
age, and which, from its octomeral type, reflected the rugosa of
palzeozoic times.
A simple conical coral with a twisted “ sérialaire” columella,
an endotheca, and an octomeral arrangement of its septal sys-
186 Mr. P. M. Duncan on some Fossil Corals.
tems, is as abnormal as the Hehidna hystria, as far as Huropean
classifications are concerned. ‘The new genus must be placed in
the neighbourhood of Avosmilia; and it connects the families
of the Turbinolides and the Astreeides.
The connexion between the septal and costal arrangements in
the species of the genus is very remarkable. The bases of the
septa and of the coste are not continuous, but the septa appear
to correspond with the line of depression between the coste.
This is common in species of other genera in Australia, but is
very rare indeed in any specimens from any other part of the
world. It was noticed in the ‘Annals’ of September 1864 in
Flabellum Victoria, nobis, and the arrangement is seen in the
two species of Sphenotrochus described in this communication.
It gives a sort of Australian stamp to the corals. The coste
are much broader than the septa; and it will be observed that
in Sphenotrochus emarciatus the line of depression between the
costz is continuous with the line which separates the two laminze
of which the septa are composed. The cost to the left and
right of the depressed line give each a root to the septum. The
species of the new genus are readily distinguished.
The Sphenotrocht are at first sight not unlike well-known
European older Pliocene and recent forms; but the emarciate
base and appendages, with the direction of the plain costa, and
the septal arrangements, distinguish the Australian species, and
prevent their being confounded with the genus Platytrochus.
The cyclolitoid Antilia is a most interesting species. The
genus superseded Montlivaltia during the Miocene (it is a
Montlvaltia with a well-formed columella); and it would appear
that all the various forms of the elder genus are represented in
the more modern. The tall cylindro-turbinate, the shorter, the
forms with oval, elliptical, or circular. calices, those with large
bases and short or tall sides, and those with many or but few
septa, amongst the Montlivaltia, are represented in the Miocene
of San Domingo, Guadeloupe, Jamaica, and Sinde by Antillie of
corresponding shape. In the Hamilton Tertiaries the interesting
cyclolitoid Montlvaltie of the Oolites have a representative in
the Antilla lens.
EXPLANATION OF PLATE VIII.
‘ig. 1. Sphenotrochus australis: a, lower half, natural size; 6, part of
calice and columella, magnified 4 diams.; c, transverse section,
magn. 4 diams.; d, coste, magn. 2 diams.
Fig. 2. Sphenotrochus emarciatus : a, nat.size ; b, side view, magn. 4 diams.;
c, calice, magn. 6 diams.; d, continuation of septa and intercostal
lines, magn. 6 diams.
Fig. 3. Conosmilia elegans: a, nat. size; 6, side view, magn. 3 Heer 3
ce, calice, magn. 6 diams,
Prof. F. M‘Coy on the Australian Species of Arvipis. 187
Fig. 4. Conosmilia anomala: a, nat. size; b, columella, magn. 4 diams. ;
e, costee, magnified 4 diams.; d, transverse section, magn. 4 diams.
(one system is closed below by endotheca) ; e, septum with endo-
theca, magn. 4 diams.
Fig. 5. Conosmilia striata : a, nat. size; 5, costee, magn. 6 diams.; c¢, trans-
verse section, magn. 6 diams.
Fig. 6. Antillia lens: a, nat. size, view from above and side; 0, base,
nat. size; c, side view (part of), magn. 4 diams.; d, base (part
of), magn. 4 diams.; e, septum, magn. 4 diams.
XXIV.—Notes on the Australian Species of Arripis. By Frepe-
rick M‘Coy, Professor of Natural Science in the University
_ of Melbourne, and Director of the National Museum at Mel-
bourne.
I rrnp that nearly all the scales of the Victorian fishes of the
genus Arripis have a more or less distinct fan-like structure of
the base, from the supposed absence of which the genus was
originally named.
Having dissected a great number, I am sure there must be
some mistake (probably a clerical error) in Dr. Giinther’s state-
ment that the pyloric appendages are from seventeen to fifty in
number, as I find them always about one hundred and sixty.
The Australian species to be found in hooks are Centropristes
Georgianus (Cuv.), C. salar (Richardson), C. Tasmanicus
(Homb.), C. trutéaceus (Cuv.), Perca trutta (Cuv.), and probably
Perca marginata (Cuv.). I have perfectly satisfied myself, from
a laborious examination of a great number of fresh specimens,
at different seasons and of all ages, that the whole of these six
supposed species should be reduced to one, and that the more
important characters relied upon by Cuvier, Richardson, and
Giinther are the peculiarities only of different ages of the fish.
The adult form is the Centropristes (Arripis) Georgianus (Cuv.)
and the C. Tasmanicus (Homb.). It reaches nearly 2 feet in
length; and, although abundant in the market, it is eaten with
great hesitation, owing to the many cases (sometimes fatal)
reported of poisonous effects produced on certain persons eating
it, although others at the same table suffered comparatively
little. It is the fish improperly called “Salmon” by the colo-
nists. It is of a nearly uniform pale olive-colour. Probably
from having counted the fin-rays of so large a number of speci-
mens, I am able to announce an extraordinary variation in this
character: thus the pectorals vary from 14 to 16, the soft anals
from 9 to 11, soft dorsals from 16 to 19.
The young, up to about 10 or 11 inches in length, are com-
monly supposed by the colonists to be a different fish, which
they call “Salmon-trout” in the markets; and they are the
188 Prof. F. M‘Coy on the Australian Species of Arripis.
Centropristes or Arripis salar of Richardson and Giinther’s
works. They have the belly silvery, back olive, sides rich green
with vertical darker bands, and four or five longitudinal rows of
round yellow spots, like lacquered brass, on the sides. This
style of colouring, so different: from that of the adult, is most
strongly marked in the young of three or four inches in length;
and I have traced in the most gradual and satisfactory way its
gradual confusion and obliteration as the size approaches 1
foot, beyond which only traces can be seen of any difference
from the nearly uniform dull colouring of the adult. The su-
perior size of the eye, the difference of proportional distance
between the orbits, and the shape of the forehead, relied upon
by authors amongst the characters separating the C. Georgianus
from the others, are more and more exaggerated as the size and
age of the individuals are less aud less.
In small, very young individuals the posterior edge of the
preoperculum is not denticulated ; and this is the great charac-
ter relied on by Cuvier and Giinther for the specific distinction
of the C. tru¢taceus in their works (the fin-rays of the adult
varying to the amount I have shown above) ; but I have clearly
demonstrated the gradual appearance and development of the
serration with increase of size; so that this is certainly (as might
even be seen by observing the relative lengths of the radiating
ridges forming the denticles going to the posterior and inferior
edges of the properculum respectively m an old fish) only a
character of immaturity.
Living specimens of the young fish three or four inches long
have the caudal fin bright yellow, with a broad posterior margin
of rich black; both these colours fade quickly, and totally dis-
appear in spirit or on a dried skin. Now as this particular
colouring, noted by Cuvier on a drawing from life of a fish of
which he had never seen a specimen, was the foundation of the
species Perca marginata in his ‘ Histoire Naturelle des Poissons,’
and all the other characters are those found likewise in the
young of Arripis Georgianus, I have no doubt, from my observa-
tion of these fugitive colours in the living fish, that Perca
marginata should be added to the synonyms of the one Australian
species of Arripis found here—the A, Georgianus. I mean to
publish figures from the life, shortly, in the ‘Decades of the
Prodromus of the Zoology and Paleontology of Victoria,’ which
I am preparing as part of the “ Memoirs of the Melbourne
Museum,” the establishment of which occupies all my leisure so
pleasantly.
Melbourne, June 24, 1865.
Mr. G.S. Brady on undescribed Fossil Entomostraca. 189
XXV.—On undescribed Fossil Entomostraca from the Brick-
earth of the Nar. By Grorce Stewarpson Brapy.
[ Plate IX. ]
For the opportunity of describing the following species of Ostra-
coda I am indebted to the kindness of Professor I’. Rupert Jones,
from whom I received the specimens. An account of the deposit
in which they occurred was given in the ‘ Geological Magazine,’
vol. il. p. 8, to which the reader is referred. The carapaces were
very few in number, and belonged to the four species here
described.
Order OSTRACODA.
Fam. Cypride.
Genus Cyrneripea, Bosquet.
Cytheridea punctillata, n. sp. Pl. UX. figs, 9-11.
Valves oblong, subtriangular, convex. Dorsal margin gently
arched, highest at its anterior third ; ventral margin straight.
Anterior border broad and well rounded; posterior narrower,
and sloping steeply to its lower extremity, which forms a
rounded angle. Seen from above, the carapace is oval in out-
line, and shows scarcely any appearance of pitting. nd view
suborbicular. Surface marked with fine and thickly set
puncta. Length 515th of an inch.
This is nearly allied to Cytheridea pinguis, Jones, and to
Bardia punctatella, Bosquet, but is not strictly referable to
either of these species. It differs from the former in surface-
ornament, as well as in the absence of angulation of the dorsal
border; from the latter, as well as from B. Hebertiana, in its
more triangular shape and finer surface-ornament.
Genus Cyruere, Miller.
Cythere carinata, n. sp.* Pl. IX. figs. 1-4.
Carapace obliquely subtetragonal, convex; margins flexuous.
Dorsal margin arched, gibbous in the middle ; ventral margin
convex, produced anteriorly into a broad, strongly developed
keel. Anterior extremity narrow, bordered partially by the
ventral keel; posterior extremity broad, somewhat truncate.
Dorsal outline broadly oval. End view ovate, tumid. Surface
* This species was noted by Prof. T. R. Jones in the ‘ Geological Maga-
zine,’ vol. il. p. 306, under the name Normania carinata. I have, however,
thought it advisable to abandon the MS. genus Normania, which was
meant to include the “ peach-stone” forms, but which I found incapable
of accurate definition or separation.
190 Mr.G.8. Brady on undescribed Fossil Entomostraca.
covered with conspicuous concentrically arranged pits, which
are well developed towards the margins, but nearly obsolete
at the centre of the valves. Length =4;th of an inch.
This species is either identical with, or very closely related to,
a recent form which is common in deep water on many parts
of the British coast, but which appears hitherto to have escaped
description.
Cythere arborescens, n. sp. Pl. IX. figs. 5-8.
Carapace broadly oval, well rounded in front, about once and a
half as long as broad. The left valve is much larger than the
right, overlapping it considerably on the dorsal and posterior
margins. The dorsal margin is strongly arched, and slopes
somewhat steeply behind towards the ventral margin, the two
being produced at their junction into an obtusely angular
prominence. Ventral margin nearly straight, somewhat in-
curved at its anterior thir d, and sloping gently upwards be-
hind. The dorsal outline is oblong oval, compressed. End
view ovate. Surface of the shell finely punctate, marked at
the extremities and along the ventral margin with an elevated
reticulated pattern, the ramifications of which are gradually
lost on the surface of the valves. Length =1th of an inch.
The recent species Cythere convexa differs from the present
only in the general outline of the valves, which in C. arbores-
cens are more decidedly quadrangular, and in the ornamentation
of the surface. But though there is much diversity in the sculp-
turing of C. convexa, I have never met with any specimens, either
recent or fossil, which show the least trace of the beautiful ar-
borescent ribbing characteristic of the present species. The
surface is also more finely punctate than in C. convexa; but I
should not, on this account alone, have thought it justifiable
to propose for it a distinct specific name.
Cythere aspera, i. sp. PL. IX. figs. 12-19.
Valves oblong, quadrilateral, compressed. Extremities nearly
equal, the anterior obliquely rounded, bordered by an elevated
nodulated ridge, which terminates in a conspicuous tubercle
over the anterior hinge, and is fringed with short blunt spines.
Posterior border produced into a broad flattened lamina, which
bears at the ventral angle three or four strong squamous
spines. Dorsal margin nearly straight ; ventral smuated and
squamous behind. Seen from above, the carapace is com-
pressed, oblongo-ovate, tuberculated, spinous behind. Hnd
view quadrilater al. Surface of the valves marked by three
conspicuous longitudinal ridges, the ventral ridge sharply
Mr. A. E. Verrill on the Classification of Polyps. 191
defined, the others nodulated and less distinct. The valves
are covered, between the ridges, with rounded tubercles. In
young specimens the longitudinal ridges are sharper, the
surface-tubercles are sharp and spinous, and the elevated an-
terior border is absent or indistinct. The young state of this
species is represented in figs. 12-15. Length (of the adult)
s4rd of an inch.
The above description applies to well-marked specimens ; and
much latitude must be allowed as to the amount of spinous and
tubercular development, especially with reference to the squa-
mous spines and lamina of the posterior extremity.
EXPLANATION OF PLATE IX.
Fig. 1. Cythere carinata (Brady), left valve, x 50.
Fig. 2. The same, seen from above, x 50.
Fig. 3. The same, seen from below, x 50.
1
2
3
Fig. 4. The same, end view, x 50.
Fig. 5. Cythere arborescens (Brady), perfect carapace, x 40.
Fig. 6. The same, seen from above, x 40.
Fig. 7. The same, seen from below, x 40.
Fig. 8. The same, end view, x 40.
Fig. 9. Cytheridea punctillata (Brady), left valve. x 40.
Fig. 10. The same, seen from above, x 40.
Fig. 11. The same, end view, x 40.
Fig. 12. Cythere aspera (Brady), right valve (young), x 40.
Fig. 13. The same, seen from above, x 40.
Fig. 14. The same, seen from below, x 40.
Fig. 15. The same, end view, x 40.
Fig. 16. The same, adult right valve, x 40.
Fig. 17. The same, seen from above, x 40.
‘ig. 18. The same, seen from below, x 40.
Fig. 19. The same, end view, x 40.
XXVI. Classification of Polyps. (Extract condensed from a Syn-
opsis of the Polypi of the North Pacific Exploring Expedition
under Captains Ringgold and Rodgers, U.S.N.) By A. E.
VERRILL*.
Tue report upon the collection made by Dr. William Stimpson,
naturalist to the expedition, having been much delayed, the
following tabular view of the classification adopted is here pre-
sented, with the hope that, if imperfect like every other, it may
nevertheless afford some aid in illustrating the natural affinities
of these humble forms.
Although, in a communication read before a zoological club
at Cambridge, January 1862, I attempted to demonstrate the
* From the ‘ Proceedings of the Essex Institute,’ U.S., for 1865.
192 Mr. A. E. Verrill on the Classification of Polyps.
existence of the three natural orders among Polyps, I refrained
from presenting this view in a paper published last year, in
order that I might make further investigations upon the sub-
ject before finally publishing it.
Class CNIDARIA or POLYPI.
Order I. MADREPORARIA.
Polyps simple or compound, with embryonic or rudimentary
basal or abactinal region, which has no special function, unless
for vegetative attachment while young. Actinal area well deve-
loped, form broadly expanded, having a tendency in the higher
groups to become narrowed towards the mouth. Tentacles
simple, conical. Dermal tissues and, usually, the radiating la-
mellie depositing solid coral ; the radiating plates, being between
the lamellze, are therefore ambulacral, and appear to originate
from the surfaces of the lamelle and the connective tissues ex-
tending across the ambulacral chambers and fillmg them from
below. Interambulacral spaces distinct.
Suborder I. Srauracea (Madreporaria rugosa) *.
Coral simple, or compound by budding ; chiefly epidermal and
endothecal; septa apparently in multiples of four, sometimes
wanting. Type embryonic, like a young Astrea or Fungia.
Families: Stauride, Cyathophyllide, Cyathaxonidi, Cysti-
phyllidee.
Suborder IT. Funeacna.
Polyps either simple or compound by marginal or disk-bud-
ding, rarely by fissiparity. Tentacles numerous, in multiples
of six, impertectly developed, scattered on the actinal surface,
usually short and lobe-like. Upper part of polyps scarcely ex-
sert. Coral broad and low, growth mostly centrifugal, tissue
* This group is placed here with considerable hesitation, and principally
on account of the close resemblance in structure to the young of the suc-
ceeding and higher groups when they first begin to form a coral, which
then consists of a ring of epitheca or epidermal deposit, with a few, imper-
fect, rugose septa radiating from the centre. If the number four be a
constant feature of the arrangement of their septa, it is possible that they
may be entitled to rank as a separate order of Polyps. To this opinion
Prof. J. D. Dana inclines. Prof. Agassiz unites the group with Hydroid
Acalephs, on account of their resemblance im some features to the Tabulata.
It seems to me, however, that the absence of transverse plates in Cyathaxo-
nidee and Cystiphyllida, and the perfection of the vertical septa in Stau-
rida, Cyathaxonid, and some of the Cyathophyllide, together with their
general structure, show them to be more closely allied to the Fungacea
and Astreacea, of which they may be considered embryonic types, while at
the same time the group is a synthetic one, having analogies with nearly
all the higher groups of Polyps and also, in some respects, with Hydroids.
Mr. A. E. Verrill on the Classification of Polyps. 193
chiefly septal; walls imperfectly developed, often perforate, sub-
ordinate, usually forming the basal attachment.
Families: Cyclolitide, Lophoseride, Fungide, Merulinide.
Suborder IIT. Astreacea.
Polyps mostly compound, either by fissiparity or various modes
of budding. ‘Tentacles usually well developed, long, subcylin-
drical, limited in number, in multiples of six, encircling the disk.
Coral mural, septal, and endothecal ; growth vertical and centri-
fugal, producing turbinated forms which are often elongated.
Families: Lithophyllhidz, Mzandrinide, Eusmiliide, Caryo-
phyllidz, Stylinide, Astreinz, Oculinide, Stylophoride.
Suborder IV. Mapreproracga (Madreporaria perforata).
Tentacles in definite numbers, twelve or more, well developed,
encircling the narrowed disk, therefore nearer the mouth ; po-
lyps with the upper portion much exsert, flexile ; growth chiefly
vertical ; coral mural and septal, porous. Polyps compound by
budding, sometimes simple.
Families: Eupsammidz, Gemmiporide, Poritide, Madre-
poridee.
Order II. ACTINARIA.
Polyps with well developed, often highly specialized, basal or
abactinal region. Walls well developed; tentacles longer, more
concentrated around the mouth, which is also usually, if not
always, furnished with special tentacular lobes or folds. Ambu-
lacral spaces always open, destitute of connecting tissues and
solid deposits.
Suborder I. ZoaNTHACEA.
Polyps encrusting, adherent, budding from mural expansions ;
tentacles simple, short, at edge of disk.
Families: Zoanthide, Bergide.
Suborder II. ANTIPATHACEA.
Polyps connected by a ccoenenchyma, secreting a solid sclero-
base or coral-axis. Tentacles few, six to twenty-four, simple,
conical.
Families : Antipathide, Gerardide.
Suborder III. Actinacea.
Polyps free, capable of locomotion, with a highly specialized
muscular base or abactinal area. Tentacles well organized,
either simple or branched, varying from ten to many hundreds,
often with accessory organs arising from the same spheromeres,
Ann. & Mag. N. Hist. Ser.3. Vol. xvi. 14
194. Mr. A. E. Verrill on the Classification of Polyps.
such as inner tentacles, verruce, complicated or simple branchial
lobes, cinclide, eye-spherules, suckers, &e. Mouth with special
lobes or folds. Most of the species are simple, a few are com-
pound by fissiparity, many abnormally bud from the wall near
the base, a few secrete from the base a horn-like deposit similar
to the axis of Antipathes.
Families: Actinidze, Thalassianthide, Minyide, Ilyanthide,
Cerianthide.
Order III. ALCYONARIA.
Polyps with well developed actinal, mural, and abactinal re-
gions, compound by budding. Tentacles eight, pinnately lobed,
long, encircling a narrow disk. No interambulacral spaces.
Ambulacral ones open and wide.
Suborder I. ALCYONACEA.
Polyps turbinate at base, budding in various ways, encrusting,
adherent to foreign bodies by the coenenchyma.
Families: Aleyonide, Xenide, Cornularide, Tubiporide.
Suborder II. GorconacEa.
Polyps cylindrical, short, connected by a coenenchyma, se-
creting a central supporting axis.
Families: Gorgonide, Plexaurida, Primnoide, Gorgonellide,
Isidee, Corallide, Briaride.
Suborder II]. PennatTuLacea.
Polyps forming free moving colonies, the composite basal
portion with locomotive functions and special cavities, with or
without a solid free axis.
Families: Pennatulide, Pavonaride, Veretillide, Renillide.
Among the most interesting species in this collection the
following may be mentioned :—
Stephanoseris lamellosa, Verrill.
Coral low, subcylindrical, with a broad base, which completely
covers small univalve shells, with the exception of the opening ;
wall rudimentary ; septa in four cycles, the primaries much the
largest, with subentire rounded tops; columella well developed,
papillose ; costee prominent, unequal.
Loo-Choo Islands. Dr. Wm. Stimpson.
Heterocyathus alternata, Vervill.
A low species with very unequal septa and cost, the primary
Mr. A. E. Verrill on new Species of Polyps. 195
septa very prominent. Encrusts and covers small univalve
shells.
Gaspar Straits. Capt. John Rodgers.
Balanophylia capensis, Vervill.
A species about half an inch high, broadly attached, slightly
turbinated, with an epitheca rising - within a line of the mar gin.
Calicle deep, broadly oval. Septa in four cycles, the principal
ones much exsert, vertical, narrowed at top, those of the fourth
eycle joining the columella in pairs. Colour of the living Polyp
bright orange.
Cape of Good Hope. Dr. Wm. Stimpson.
Eupsammia Stimpsonii, Verrill.
Coral free, elongated, turbinated, blunt at base. Calicle oval,
deep; columella well developed ; septa broad, the principal ones
with entire inner edges, rounded. Length an inch or more ;
breadth of cell °3 in.
Interesting as a living representative of a genus hitherto
known only in the fossil state.
North China Sea. Dr. Wm. Stimpson.
Metridium fimbriatum, Verrill.
A species closely allied to M. marginatum of this coast, but
apparently more elongated, with longer and more slender tenta-
cles, which are almost hair-like. Disk within the tentacles nar-
row. “Colour pale orange, translucent; body punctate with
dark brown; mouth deep orange.”
San Francisco, California. Dr. Wm. Stimpson.
Phellia collaris, Verrill.
Edwardsia collaris, Stimpson, Proc. Philad. Acad. Nat. Science, May and
June 1855.
A species remarkable for its great size compared with pre-
viously known species from Europe.
Hong-Kong, China. Dr. Wm. Stimpson.
Phellia clavata, Vervill.
Edwardsia clavata, Stimpson, /. c. 1855.
A species even larger than the last.
Near Ousima, Japan. Dr. Wm. Stimpson.
AMMONACTIS, Nov. gen.
Column elongated, subcylindrical, with well developed basal
disk, covered, as in Phellia, with a persistent epidermis extend-
14
196 Mr. A. E. Verrill on new Species of Polyps.
ing to near the summit, naked above; but differs in having a
lobe-like tubercle below each tentacle, distinct from the margin.
Tentacles long and numerous.
Ammonactis rubricollum, Verrill.
Edwardsia rubricollum, Stimpson, l. ¢. 1855.
Hong-Kong, China. Dr. Wm. Stimpson.
Halocampa brevicornis, Vervill.
Edwardsia brevicornis, Stimpson, 1. c. 1855.
Hong-Kong, China. Dr. Wm. Stimpson.
Halocampa capensis, Verrill.
Body elongated; tentacles twenty, blunt; ambulacra sub-
papillose. Six tentacles have their inner bases dark brown ;
body pale reddish, with dots and patches of flake white ; inner
side of tentacles flake white.
Cape of Good Hope, 12 fathoms, sand. Dr. Wm. Stimpson.
Cerianthus orientalis, Verrill.
A large species, similar to C. americana, nobis. Body elon-
eated, ina tube of mud. Tentacles long and slender. Colour
of body deep reddish brown ; outer tentacles translucent, yellow-
ish and white, pale brown on their inner sides, greenish at base ;
inner ones purplish brown, or sometimes grass-green.
At low-water mark, Hong-Kong, China. Dr. Wm. Stimpson.
Nephthya thyrsoidea, Verrill.
Polyps forming thyrsiform bunches of closely clustered branch-
lets, 3 inches high and 2 inches broad. Colour wine-yellow or
light brown, with a dark purplish tinge below the tentacles ;
tentacles nearly white ; spicula forming elevated transverse lines
of silvery white on the stalks.
Cape of Good Hope, 20 fathoms, rocks. Dr. Wm. Stimpson.
Telesto ramiculosa, Verrill.
Cornularia aurantiaca, Stimpson, J. c. 1855, non T. aurantiaca, Lamx.
Hong-Hong, 10 fathoms, shelly bottom. Dr. Wm. Stimpson.
Parisis laxa, Verrill.
Coral forming openly reticulate fronds; papillze numerous,
rounded, on all sides of the branches; coenenchyma minutely
villous in alcohol. Calcareous joints shorter, and internodes
longer, than in P. fruticosa, nobis.
Hong-Kong. Dr. Wm. Stimpson.
Royal Society. 197
Acanthogorgia coccinea, Verrill.
Nepthya coccinea, Stimpson, /. ce. 1855.
Hong-Kong, 10 fathoms, on shells. Dr. Wm. Stimpson.
Veretillum Stimpsonii, Verrill.
A large species, 6 or 8 inches long, the upper portion en-
larged, more than half the entire length. Polyps much exsert,
upwards of an inch long; tentacles very long. Axis thick,
short, fusiform, a third of an inch long. Base white, somewhat
striated; body light cream-colour ; polyps transparent, bluish
white at the bases of the tentacles.
Hong-Kong, 6-10 fathoms, mud. Dr. Wm. Stimpson.
Veretillum baculatum, Verrill.
Club-shaped, the base about a third of the length. Polyps
scattered, not numerous. Axis small, fusiform, about half an
inch long in a specimen 3 inches long.
Sea of Ochotsk, off Siberia. L. M. Squires.
Kophobelemnon clavatum, Verrill.
Veretillum clavatum, Stimpson, J. c. 1855.
Polyps more numerous and crowded than in K. Burgeri,
Herkl., which it resembles; body more claviform, naked dorsal
space very narrow.
Hong-Kong, 6 fathoms, mud. Dr. Wm. Stimpson.
PROCEEDINGS OF LEARNED SOCIETIES.
ROYAL SOCIETY.
June 15, 1865.—Major-General Sabine, President, in the Chair.
‘On the Anatomy and Physiology of the Nematoids, parasitic
and free; with observations on their Zoological Position and Affini-
nities to the Echinoderms.’ By Henry Charlton Bastian, M.A.,
M.B. (Lond.), F.L.S.
After commenting upon the many conflicting statements which
have been made concerning the anatomy of these animals, and more
especially with regard to the presence or absence of a nervous
system, and of real organs of circulation, the author alludes to the
increased interest which has lately been thrown over this order by
the discovery of so many new species of the non-parasitic forms,
marine, land, and freshwater.
He has entered fully into the description of the tegumentary
organs, and has recognized a distinct cellulo-granular layer inter-
vening between the great longitudinal muscles and the external chiti-
198 Royal Society :—
nous portion of the integument. This layer is one of great import-
ance in the economy of these animals; the author looks upon it as
the deep formative portion of the integument, from which the chi-
tinous lamellee are successively excreted. It is bounded internally
by a fibrous membrane, which serves as an aponeurosis for the at-
tachment of the four great longitudinal muscles ; and the well-known
lateral and median lines which have so long been a puzzle to ana-
tomists are, he says, in reality nothing more than intermuscular
developments of this layer. In some species each of the lateral lines
contains an axial vessel, though in very many others nothing of this
kind is to be met with. A periodical ecdysis of the chitinous por-
tion of the integument takes place in all Nematoids during the period
of their growth.
The author agrees with Dr. Schneider as to the nature of the
transverse fibres attached to the median lines. They are contractile
prolongations from the longitudinal muscles, and may be considered
extrinsic muscles for the propulsion of the intestinal contents, since
the intestine itself has no muscular tissue in its walls.
Schneider’s description of the nervous system in Ascaris megalo-
cephala has been-confirmed, and a similar arrangement has been re-
cognized by the author in several other Nematoids. It exists as a
nervous ring encircling the commencenient of the cesophagus, in
connexion with many large ganglion-cells. The principal peripheral
branches are given off from the anterior part of the ring, and pro-
ceed to the region of the mouth and cephalic papilla. Although
well developed ocelli exist in many of the free marine species, no
nerve-filaments have yet been detected in connexion with them.
The organs of digestion are mostly simple, the principal variations
being met with in the presence or absence of a pharyngeal cavity,
and in the structure of the cesophagus. In some species its parietes
are distinctly muscular, whilst in others, as in the Trichocephali
and Trichosomata, they are as distinctly cellular. Those possessing
a pharyngeal cavity sometimes have well-marked tooth-like processes
developed from its walls; but the author believes that the chitinous
plates which are sometimes met with in posterior swellings of the
cesophagus are not ‘gastric teeth,’ as they have been hitherto de-
scribed, but rather valvular plates for ensuring greater perfection in
the suctorial process by which these animals | pass their food along
this portion of the alimentary canal.
The water-vascular system may be seen in many Nematoids in its
most elementary condition, as a small tubular gland, with an excretory
orifice in the mid-ventral region of the anterior part of the body. In
other Nematoids no trace of such a system exists; whilst its most
developed condition yet recognized in these animals may be seen
in Ascaris osculata and A. spiculigera, where an intimate plexus of
vessels, still in connexion with an anterior ventral pore, is met with
in a peculiar development from the left lateral band. Intermediate
conditions between these extreme forms may be traced in other spe-
cies; and from the obviously glandular nature of the tubular or
pyriform organ met with so commonly in the free, and also in many
Mr. Bastian on the Anatomy and Physiology of Nematoids. 199
of the parasitic species, he thinks considerable light is thrown upon
the function of the “ water-vascular”’ system. He says, ‘‘ Here we
have undoubtedly to deal with an excretory glandular apparatus.
No one could for a moment regard these structures as at all analo-
gous to vessels destined alternately to receive and discharge an ex-
ternal fluid medium. I believe that in the Trematoda and Teniada
also, where similar though often more developed systems exist, their
function is in like manner one of a purely eliminatory kind; and I
therefore cannot but look upon the name of ‘ water-vascular ’ appa-
ratus as a singularly inappropriate appellation for this system of
vessels.”
Other very peculiar transverse vessels exist in the deep integu-
mental layer of Ascaris megalocephala and A. lumbricvides, mostly
running in pairs from median line to median line, and, strangely
enough, being about twice as numerous on the right as on the left
side of the body.
The author believes that in the Nematoids but little provision
exists for the oxidating portion of the process of respiration, and
thinks that this deficiency may be compensated by a greatly increased
activity of glandular eliminating organs. Considering the conditions
under whose influence so many of the parasitic forms pass their
existence, we can easily imagine that the presence of any organs for
effecting an oxidation of their tissues would not only be useless, but
actually baneful. Glandular organs exist in the greatest abundance
in all Nematoids, and many of these are excretory organs. In those
species in which no modification of the ventral excretory apparatus
is met with, the author has found a very large number of channels
running through the chitinous portion of the integument, so as to
bring its deep cellular layer into communication with the exterior.
These pores are, he believes, complementary respiratory organs, and
their development is always in an inverse proportion to that of the
other excretory organs. Thus amongst the free Nematoids he has
found them most numerous in Dorylaimus stagnalis and Leptoso-
matum figuratum—species in which the ventral excretory apparatus
is entirely absent. The same arrangement is met with in the T7i-
chocephali and Trichosomata, in which these integumental channels
attain their maximum development. The gradually widening longi-
tudinal band long known to exist in the 7'’rchocephali is due to the
presence of thousands of these channels in connexion with a glandular
development of the deep integumental layer beneath.
Many interesting facts are brought forward concerning the “ tena-
city of life’ of some of the free Nematoids, and their power of re-
covery after prolonged periods of desiccation. This has been long
known as one of the characteristics of T'ylenchus tritici*, but the
author has found it common only to the species of four land and
freshwater genera— Tylenchus, Plectus, Aphelenchus, and Cephalo-
bus. The remainder of the free Nematoids are remarkably frail, and
incapable of recovering even after the shortest periods of desiccation.
* Vibrio tritici of older writers.
200 Royal Society :—
In the last section, on ‘The zoological position and affinities of
the Nematoids,” the author enters fully into what he believes to be
the points of resemblance between these animals and the Echino-
derms. The strongest evidence is, he thinks, to be found in the fact
of the very close resemblance between the nervous systems of these
animals, differing notably as they do at the same time from what we
find in the Scolecida or Annelida. Then the integumental pores
which he has now discovered in so many Nematoids can, he thinks,
be paralleled only by the ambulacral and other pores met with in
the Echinoderms. Great similarities in the distribution of these
pores may also be observed in the two groups. The Nematoids
present no trace of segmentation or lateral appendages to their bodies,
but traces of a radiate structure do exist. Their various parts and
organs exhibit a quadrate mixed with a ternate type of development.
He looks upon the order Nematoidea as an aberrant division of the
class E’chinodermata, which at the same time tends to connect this
class in the most interesting manner with the Scolecida—since,
although in the points above mentioned they display their affinities
to the Echinoderms, still, as regards the structure and different
modifications of the ventral excretory apparatus, they agree more
closely with the 7’ematoda or flukes.
“Researches on the Structure, Physiology, and Development of
Antedon (Comatula, Lamk.) rosaceus.” By Dr. W. B. Carpenter,
F.R.S.
The author, after adverting to the special interest attaching to the
study of this typical form, as the only one readily accessible for the
elucidation of the life-history of the CrinorpEa, states it to be his
object to give as complete an account as his prolonged study of it
enables him to offer, of its minute structure, living actions, and
developmental history, taking up the last at the point to which it
has been brought in the memoir of Prof. Wyville Thomson.
He prefaces his memoir with an historical summary of the progress
of our knowledge of the distinctive peculiarities of this genus, and of
its relation to the Crinoidea; and he shows that the first recoguition
of this relationship was most distinctly made by Llhuyd, at the
beginning of the last century, though that recognition has been
passed without notice by most subsequent writers, and is altogether
ignored by MM. de Koninck and le Hon in their recent history.
The author then proceeds to describe the external characters of
Antedon rosaceus; and shows, from its habits as observed in a
vivarium, that although possessed of locomotive power, it makes so
little use of this under ordinary circumstances, that its life in the
adult condition, no less than in its earlier stage, is essentially that of
a pedunculate Crinoid.
He then gives a minute description of the several pieces of the
skeleton—the accounts of these previously given by J. S. Miller
and Prof. Joh. Miller not being in sufficient detail to serve as
standards of comparison to which the parts of fossil Crinoids may be
Dr. Carpenter on Antedon rosaceus. 201
referred. And he directs special attention to the curiously inflected
rosette-like plate, previously unnoticed, which occupies the central
space left within the annulus formed by the adhesion of the first
radials. This plate is in spectal relation to the organ termed by
Joh. Miiller the “heart,” but certainly having no proper claim to
that designation, being a quinquepartite cavity in the central axis,
from the walls of which there pass out not vessels but solid cords
of sarcode, into the rays and arms, and also into the dorsal cirri.
The inflexions of the rosette-like plate serve for the support aud
protection of the large cords passing into the rays, each of which has
a double origin, and a connexion with the adjacent radiating cords
that reminds the anatomist of the ‘circle of Willis.”
The skeleton of the adult differs so widely in the forms and rela-
tions of its parts from that of the early Pentacrinoid larva described
by Prof. Wyville Thomson, that the derivation of the former from
the latter can only be understood by observation of all the inter-
mediate stages. When the calcareous skeleton of the calyx first
shows itself, it consists only of five oral plates arranged conform-
ably upon five dasal plates, as thus :—
OCOs OF 200
B By BBB
At a stage a little more advanced (which has been described by
Prof. Allman, Trans. Roy. Soe. Ed. vo]. xxiii. p. 241), the rudiments
of the first radials are found interposed between the orals and basals,
alternating in position with both, as in the
margin ; and between two of these first radials O O O O O
there appears a single small unsymmetrical a a a a a
plate, which afterwar ‘ds proves to be the anal. B B B B B
The first radials undergo a rapid increase in
size, and soon become Renenouied by second and third radials,
which project between the orals; whilst the orals and basals, under-
going no such increase, are relatively very much smaller; the anal
plate i is still found on the line of the first
radials. But from this time the radials A® A® A*® A® A?
form the principal part of the calyx, A? A? A? A? A?
which opens out widely in conformity OF 02 O72 OO
with the increase of space required for A’ A’anA’ A* A?
the digestive apparatus, the intestinal Bo Be bab ae
canal being now developed around what
was originally a simple stomach with one orifice. The highest joint
of the stem also undergoes a remarkable increase in size, and begins
to acquire the form of a basin with an inflected rim, constituting
what is known in the adult as the centro-dorsal piece. When the
calyx opens out, the five oral plates, which originally formed a circlet
around the mouth, retain that position, and detach themselves en-
tirely from the divergent radials, nothing but the soft perisomatic
membrane filling up the space between them. These oral plates
never increase in size, and towards the end of the Pentacrinoid stage
202 Royal Society.
they begin to undergo absorption. I can still trace their basal
portions in young specimens of the free Antedon; but as the
creature advances towards maturity they are altogether lost sight of.
When the intestinal canal has been sufficiently developed to open on
the surface of the oral disk, the anal plate is lifted out of the posi-
tion it originally occupied, and is at last found on the anal funnel,
far removed from the radials. This, like the oral plates, begins to
undergo absorption towards the end of the crinoidal stage, and
completely disappears in the early part of the life of the free Ante-
don. The radial plates increase not only in size but also in thick-
ness; and channels which are left on their internal surface by
vacuities in the calcareous network, are converted into canals by a
further inward growth of this, which completely covers them in.
It is through these canals that the cords of sareode pass to the arms.
The édasal plates, like the oral, remain stationary in point of size,
and present no change in appearance. or position until after they
have been completely concealed externally by the centro-dorsal piece
(the highest joint of the stem), which rapidly augments, both in
absolute and in proportional size, when the development of the dorsal
cirri is taking place from its convex surface. By the end of the
Pentacrinoid stage, this plate has extended itself so far over the base
of the calyx as completely to conceal the basals; and as the free
Antedon advances towards maturity, it gradually extends itself over
the first radials, which then become adherent to it and to each other.
The basals then undergo a most curious metamorphosis, consisting
in absorption in one part and extension in another, by which they
finally become converted into five peculiarly shaped pieces, the
ultimate union of which forms the single rosette-like plate which
has been already stated to lie within the annulus formed by the first
radials of the adult Antedon. Hence the calyx finally comes to be
thus composed :—
b BR b R b Rb RD RF
BF
CENTRO-DORSAL.
As the orals and the anal have entirely disappeared, no part of
the primordial calyx of the Pentacrinoid larva is traceable in it, until
we separate the adherent pieces which form its base, and search out
the minute and delicate rosette-like plate which is formed by the
metamorphosis of the dasals.
The structure, physiology, and development of the digestive, cir-
culatory, and respiratory apparatus, and of the neryous and muscular
systems, will form the subject of a future memoir.
203
ZOOLOGICAL SOCIETY.
March 28, 1865.—John Gould, Esq., F.R.S., in the Chair.
Notice OF AN APPARENTLY UNDESCRIBED SPECIES OF AMERI-
cAN Porcupine. By Dr. J. E. Gray, F.R.S., F.L.S., erc.
There has been in the British Museum since 1853 a small specimen
of a short-tailed American Porcupine, which was sent from Columbia.
I suspected that the animal might be young ; and I have been wait-
ing, expecting that we might receive another specimen from the
same source, which would enable me to give a more complete account
of the animal; but as no additional materials have come to hand, I
shall now proceed to give a short notice of it, in the hope of draw-
ing the attention of collectors to the animal.
ERETHIZON (ECHINOPROCTA) RUFESCENS.
Pale brown, varied with black ; head white, speckled with black
and pale brown; tail and feet black ; chin, throat, and beneath pale
brown. A short white streak on the centre of the nose, and a few
white spines, forming a slight crest, on the nape; a whitish mark on
the side of the cheek. The bristly spines of the head thin, white,
with a small black subterminal band and yellow tip; the spines of
the back elongate, white, with a black subterminal ring and elon-
gated rufous tips; those of the front part of the back and sides very
slender, bristle-like, gradually becoming thickened, stronger, and
shorter, until on the hinder part of the back, above the tail, they are
well developed, short, thick. Spines with black ends and very small
brown tips. The end of the nose, chin, and underside of the body
covered with uniform pale brown slender bristles. The tail and
feet covered with short black bristles. Whiskers black, slender,
flexible.
Hab. Columbia.
There are a few spines on the top of the head, with one white to
the tip, making a kind of occipital crest; but I am not sure that
this may not be an individual peculiarity.
The soles of the hind feet are bald to the heel. Cutting-teeth
yellow, slender, rounder in front. Unfortunately I have not been
able to see the skull.
If this is a true Hrethizon, the genus may be divided into two
sections :—
1. Erethizon. The back covered with elongated bristles and
short spines. J. dorsatus and E. epixanthus.
2. Echinoprocta. The back covered with one kind of elongated
slender spines, which become shorter, thicker, and more rigid over
the rump. J. rufescens.
Noricre or A Srecies or TupatA FROM BORNEO, IN THE
CoLLECTION oF THE British Museum. By Dr. Joun
Epwarp Gray, F.R.S., F.L.S., erc.
There has been in the British Museum for some years a specimen
204 Zoological Society :—
of a Tupaia in spirits, which was received from Borneo, and also a
stuffed specimen without a habitat, evidently of the same species.
These specimens have the general coloration of Tupaia tana, and
have evidently been regarded as varieties of that species; but they
are most distinct. The head and skull are short and broad, of about
the same form and proportion as those of Tupaia ferruginea; the
fur and tail is of the same bright shining bay as 7. tana, but it is
entirely destitute of the three black streaks between the shoulders,
which are so well marked in that species.
The skull shows that the stuffed specimen is that of an adult
animal not so large as 7’. tana, and more nearly resembling in size
T. ferruginea. It may be known at once from the latter species by
the dark red-brown colour of the tail, with its very red underside.
I propose to call it
TUPAIA SPLENDIDULA.
Fur dark red-brown, blackish-washed. Tail dark red-brown ;
pale red beneath; the shoulder-streak yellow. The head conical,
about twice as long as wide behind.
Hab. Borneo.
The head is large compared with the size of the body; the ears
rounded, with several ridges on the conch, and a well-developed con-
vex tragus, not unlike the human ear. The palm and soles are bald
to the wrist and heel.
I thought at first that this species might be the Tupaia speciosa of
Wagner; but that animal is stated to have a head as long and as
tapering as 7’. tana, and, indeed, seems to be only a slight variety of
that species.
Norice oF A New GENUS AND SPECIES OF THE FamiLy Trio-
NYCHID FROM WESTERN Arrica. By Dr. J. E. Gray,
F.R.S., F.L.S., ec.
The British Museum has just received two specimens of a Trionyx
with covered legs from Western Africa (collected by the late Dr. B.
Baikie, probably on the Niger), which is evidently different im
structure from any we have before received from that country, and
which I am inclined to believe is an entirely new form.
It differs from the other African 7’rionyches with covered feet in
only having two pairs of callosities on the sternum ; while Hepta-
thyra has seven, and Cyclanosteus has nine such hardnesses on the
sternal bones. These callosities differ in disposition and mode of
development, as well as in manner, in the three genera, The skull
is in form like that of the genus Cyclanosteus; that is to say,
the face is moderate, with eyes about halfway between the front of
the zygomatic arch and cavity of the temporal muscle and the end
of the nose ; but it differs from the skull of the latter genus in the
forehead and crown being wider and flatter.
The genus (which I should refer to the tribe Cyelanosteina) may
be defined thus :—
Dr. J. E. Gray on a new Form of Trionychide. 205
TETRATHYRA.
The face of the skull short, convex, arched in front ; orbits lateral,
shelving, about midway between the end of the nose and the front
of the zygomatic arch ; forehead flat, rhombic, broad. The dorsal
shield with flexible margins, without any marginal bones ; front of
dorsal shield warty above and without any odd nuchal bone. Ster-
num flat, with broad rounded lobes covering the feet, and two pairs
of sternal callosities ; the front pair small, rounded, on the front
ends of each of the front pair of sternal bones ; the lateral pairs
are large, oblong, broadly notched out behind, and very rugose.
This genus differs from Oyclanosteus in the want of any odd bone
in front of the dorsal shield, as well as in the number and disposition
of the sternal callosities.
The upper surface of the front of the disk is closely covered with
roundish warts. The sternal callosities are not developed in the
young specimen, the larger lateral pair being first indicated as the
animal increases in size. The dorsal disk of the young specimen is
marked with close grains, or warty, in rather arched longitudinal ridges.
There are some young specimens in spirits from West Africa in the
Museum, which belong to this species ; they differ from the young
of C. senegalensis in being marbled, while that species is marked
with distinct small subcircular black spots.
This second genus of Cyclanosteina may explain the reason why
we have two skulls from West Africa the one with the front and
the other with the whole upper edge of the lower jaw dilated, as
figured in the ‘ Proceedings of the Zoological Society’ for 1864,
fig. 18, p. 95, and fig. 21, p. 96.
TETRATHYRA Baiktl, sp. nov.
Head olive, white-spotted. Back olive, marbled with black above ;
206 Zoological Society :—
the lower surface pale, irregularly black-marbled or spotted. The
front pair of callosities small, oblong.
Younger specimen, the head and dorsal shield pale brown, mar-
bled with large black (often inosculating) streaks ; lower part of head
and sternum black, with large, irregular-sized, pale spots, some of
which are symmetrical.
Hab. West Africa, River Niger?
The largest specimen, which is not full-grown, is 11 inches long;
the dorsal shield 7 inches long and 5 inches wide. .
April 11, 1865.—Prof. T. H. Huxley, F.R.S., V.P., in the Chair.
DescrIPTION OF A New Species or INDIAN PORCUPINE.
By P. L. Scuater, M.A., Pa.D., F.R.S.
About three years ago I received a communication from our excel-
lent Corresponding Member, Colonel Sir William Thomas Denison,
K.C.B., Governor of Madras, inquiring of me whether anything was
known in Europe of a second Indian Porcupine, distinguished from
the common species by having some of its quills of a deep orange-
colour. Upon my replying that this Porcupine appeared to be un-
represented in our collections of animals either living or dead in this
country, and would moreover probably prove new to science, Sir
William promised to do his best to obtain living specimens of it for
the Society’s Menagerie. The first examples of this animal obtained
by Sir William for transmission to this country died, I believe, before
they were shipped. But in the latter part of last year Sir William
was successful in obtaining four other living specimens, which reached
this country in safety on the 22nd of December last. Three of these
Porcupines are still living in the Society’s Menagerie. The fourth
died a few days after its arrival, and was found one morning already
partially devoured by its carnivorous companions. Enough, however,
remained of it to make a tolerably good skin, which, together with
the skull, I now exhibit. Upon these materials I propose to attempt
to give characters to this hitherto undescribed species.
Before doing so, however, I should mention that this species,
although it has never yet been described, and, as far as I can ascer-
tain, has never reached Europe before, alive or dead, has been already
provided with a name, which I do not propose to alter. Mr. Francis
Day, Fellow of this Society, late of H.M. Madras Medical Service,
in his work on the native Indian state of Cochin, called ‘The Land
of the Permauls,’ published at Madras in 1863, has spoken of this
animal as “ The Orange Porcupine, Hystriv malabaricus,” and given
some details respecting it*. Mr. Day has also kindly supplied me
with some further notes respecting it, which I shall give presently.
I commence, however, by characterizing the species, which belongs
to the typical Hystrices, and is very closely allied to H. leucura, as
HystrRix MALABARICA, Sp. nov.
Hl. crist@ setis purpurascenti-nigris, unicoloribus ; rostro pilis
* Land of the Permauls, pp. 446, 447.
Dr. P. L. Sclater on a new Indian Porcupine. 207
minutis obsito: colore corporis antici purpurascenti-rubro,
spinis ad basin aurantiacis, inde ad apicem purpurascenti-
nigris: spinis dorsi elongatis, aliis aurantiaco-rubro et nigro,
aliis, sicut in specie vulgari, albo et nigro annulatis: dorsi
postict linea mediali distincta, e spinis aliis albis, aliis auran-
tiucis composita: cauda longa, spinis aliis albis, aliis auran-
tiaco-rubris.
Long. tota a rostro ad basin caudze 28°0 poll., caudze 8-0.
Heb. India Meridionalis, prov. Cochin.
Obs. Affinis H. leucure, sed spinarum colore, rostro minus
setoso, et cauda longiore distinguenda.
Although the general external appearance of this Porcupine is re-
markably different from that of H. lewcura, so that the living animal
strikes one at the first glance as being undoubtedly distinct, I have
been somewhat disappointed, on comparing the two skins together, to
find how difficult it is to detect any very decided differences in their
structure. The muzzle in the present specimen of H. malabarica
(which is the only individual I have been able to examine) seems to
be decidedly less clothed with hair than in H.leucura. This is one
of the few points in which H. lewcura differs externally from H.
cristata, and in this respect the present specimen seems more like
H. cristata. The whole of the short spines and hairs of the anterior
portion of the body in H. malabarica are dark reddish orange at
their bases, growing into purplish brown at their tips; and the same
is the case with those of the flanks and Jegs. The elongated spines
of the middle of the back are some of them black, annulated with
white, just asin H. /eucura ; others, more especially towards the sides,
where these latter rather predominate, have the white replaced by a
bright orange-red. The medial line of the rump is well defined, as
in H. leucura; but the white spines are mixed with others wholly
orange. This is likewise the case with the spines round the base of
the strong spines which terminate the tail: some of these are wholly
white, and some wholly orange. The strong spines which surround
the tail, and extend beyond its extremity, are mostly wholly white,
with some wholly orange intermixed. In the centre of these are
about twelve of the singular hollow truncated quills mounted on pe-
dicels, just as in ZH. leucura and H. cristata*. About one-fourth
part of these abnormal quills are orange; the others are white.
As the cranial characters of the species of Hystrix are generally
very well marked, and indeed the only test by which the species can
be certainly distinguished, I was in hopes of finding in the cranium
of Hystrix malabarica some more certain evidence of its real distinct-
ness from H. leucura. Ihave therefore carefully compared the skull
of the new species with a fine series of six skulls of H. /ewcura in the
British Museumf, in doing which I have received the valuable as-
* T am not aware whether any explanation has ever been given of the use of
these curious quills. My impression is that they serve to act as a rattle, which is
thus formed, as in the Rattle-Snakes (Crotalus), by a cutaneous development at
the end of the tail.
+ A. cristata and H. leucurus of the ‘ Catalogue of the Bones of Mammalja in
the British Museum ’ (1862), p. 191.
208 Zoological Society :—
sistance of my friend Dr. Peters, who happened to be present on the
oceasion. The skull of Hystriv malabarica, which is that of a very
old animal with the molar teeth worn very low and the cranial sutures
nearly obliterated, agrees in the shape of the nasal and intermaxillary
bones with JZ. /eucura. As in the latter species, so in H. malabarica
the nasal bones have their sides nearly parallel with the hinder mar-
gin, terminating nearly in a line with the anterior edge of the orbit,
and the nasal processes of the intermaxillaries are broad and truncated.
At first I was inclined to think there was some difference in thé pat-
terns of the molar teeth of the two species, those of H. malabarica
being surrounded by a complete cingulum of enamel, and the internal
areas being completely isolated, which is not the case in H. leucura.
But this, I suspect, is only due to the age of the specimen. It would
therefore be desirable to have further specimens of the skull of H.
malabarica for comparison upon this point; but in other respects
there seem to exist differences in the skulls of the two species which
are amply sufficient to confirm their specific separation.
wy i
Skull of Wystrix malabarica.
1. In Hf. /eucura the total length of the molar series is greater
than the distance between the molars and the tympanic bone ; in //,
malabarica it is rather less.
2. In H. malabarica the entopterygoid is more remote from the
tympanic bone, and is of a different form.
3. The facial surface of the lachrymal is very small in H. mala-
Zarica—much smaller than in H. leucura.
4. The rostral part of the cranium is more elongated and more
Dr. P. L. Sclater on a new Indian Porcupine. 209
compressed in H. malabarica, and the foramina incisiva are longer
and narrower.
These and other minor peculiarities will, I think, sufficiently serve
to separate H. maladarica from its nearest ally, although it is of course
desirable that further specimens should be obtained for comparison.
With regard to the habits of H. malabarica, Mr. Day has kindly
furnished me with the following particulars :—
“During my residence at Cochin I was informed by the natives
that a species of orange-coloured Porcupine was found in the neigh-
bouring hills, the flesh of which was more highly esteemed for food
than that of the common variety. It was said to be a smaller species,
and that the two never lived in the same locality. Small families of
them, I subsequently ascertained, are found in various places along
the ghawts of Cochin and Travancore.
** At Trichooe, about forty miles north-east of Cochin, there was a
colony of these animals. They had formed their burrows in the
laterite rock, in a spot from which it was impossible to reach them
by digging. As I was anxious to obtain one of them, the burrows
were stopped and a pitfall dug before two, which were the most fre-
quented; brushwood was then heaped before the other apertures
and set on fire, but the prisoners did not venture out until they had
been smoked three days and nights.
** The native sportsmen declare that the aroma from these burrows
is quite sufficient to distinguish the different species.
«In 1862 I placed a pair, about a third grown, in a cage, and kept
them there nearly two months: although they permitted the dogs
and cats to steal their food, they never became tame or even fr iendly
with those who fed them.
*“They were omnivorous ; and, though quiet all day, as soon as
it became dusk they commenced to gnaw their cage, and continued
to do so until daybreak ; subsequently, when the bars were encased
with tin, they passed the night scratching.
“In captivity they lose much of their orange-colour; and its
vividness greatly decreases when they are ill.
«The natives consider wounds caused by their quills to be venom-
ous, and the effects frequently fatal.”
It may be useful to add to this paper a list of the known species
of Hystriz, and their localities, arranged according to Mr. Water-
house’s excellent system *.
a. Species nucha cristata.
1. H. cristata, Linn. et auct. (Acanthion Cuvieri, Gray); Water-
house, 7. c. p. 448: ex Europa merid. et Africa bor. et occ.
2. H. Arric% AUSTRALIS, Peters, Reise n. Moss. i. p. 170: ex
Africa austr. orient.
3. H. tevcura, Sykes (H. hirsutirostris, Brandt ; Waterhouse,
l.c. p. 454; H. cristata et H. leucurus, Gray): ex Asia occiden-
tali usque ad Indiam extremam.
4. H. mavaparica: ex India merid.
* Nat. Hist. Mamm. vol. ii. p. 446 e¢ seg.
Ann. & Mag. Nat. Hist. Ser. 3. Vol. xvi 15
210 Zoological Society :—
6. Species nucha non cristata.
5. H. Honeson1, Gray ; Waterhouse, /. c. p. 461 : ex India supe-
riore.
6. H. savanica (F. Cuv.) ; Waterhouse, 7. ¢. p. 465: ex Java.
The Society’s collection contains at the present time fine living
specimens of four of these, namely, H. Africe australis, H. leucura,
H. malabarica, and H. javanica.
Nores ON THE WHALES OF THE Care; BY E. L. Layarop,
Esa., or Care-Town, Corr. Mems. Wits Descriprions
or Two New Species; By Dr. J. E. Gray.
Mr. E. Layard, the Keeper of the South African Museum at
Cape-Town, has most kindly sent me descriptions and drawings,
made by Mr. Trimen, of the skulls of the Cetacea contained in that
museum. Amongst these is the drawing of a Porpoise or Grampus
taken in Kalk Bay (Simon’s Bay). Unfortunately the skull of this
animal was placed in the skin during Mr. Layard’s absence from the
Cape ; so that it cannot be got at for description ; but, from what he
saw of the dentition, he believes it is like a Grampus—very like the
figure of the skull of G. Cuvierti in the ‘Catalogue of Cetacea in the
Collection of the British Museum,’ t. 5. f. 1. He says that there is
a separate skull, greatly resembling that figure, in the South African
Museum.
The Grampus (?) prepared with the skull in the skin, mentioned
above, is represented as having a rounded head, without any appear-
ance of a beak. ‘‘ It is entirely deep brown black ; the skin smooth,
with a few wrinkles behind the chin and on the front edge of the
pectoral fin.
‘“‘ The entire length, from the nose to the end of the tail, 8 feet ;
from the nose to the front base of the dorsal fin, along the curve,
3 feet 9 inches; of the dorsal, 10 inches; of the back, from the
hinder edge of the dorsal fin to the end of the tail, 3 feet 10 inches ;
width of the tail, 1 foot 11 inches.
“‘ Length from the front of the mouth to the base of the pectoral,
1 foot 545 inches ; of the upper edge of the pectoral, 1 foot 5 inches.”
In the South African Museum are two smaller skulls from the
coast of the Cape,—one apparently of a Steno, with a teeth; and
the other of a Delphinus, probably the common one of Table Bay,
which has oe teeth.
These are probably new species, to be described.
Mr. Layard observes, ‘‘ These Cetaceans are constantly in the Bay ;
but I cannot get the fishermen, who catch plenty of the Delphinus,
to bring them to the museum. I have offered the market value, be-
sides all the flesh and the blubber ; but they are so prized as food by
the men that they are cut up instantly and sold by auction.”
“Two, if not more, species of Whales come into our bays to calve.
I have never been fortunate enough to see them entire; but, from
the remains, I think them to be the ‘ Right Whale”? (Balena) and
Mr. E. L. Layrad on the Whales of the Cape. 211
Humpback (Megaptera). By the way, do you know the meaning
of Peskop? The Dutch are the dirtiest-minded people I ever met
with: they have heaps of such names for their animals and plants.”
“I have seen off the coast several species of Whale (one near
Agulhas, with an enormous elongated back-fin ; which could it be?)
They are in sight for an hour at least.”
**T send you a drawing, by our friend Mr. Trimen, of the skull of
a Cetacean which I have taken to be a Ziphius, probably a very old
Ziphius sechellensis ; but the figure in your ‘ Catalogue of the Ce-
tacea in the British Museum,’ ack. 2, does not convey any idea
how the curious flattened teeth arch over the upper jaw, as shown
in Mr. Trimen’s drawing. I stood by him all the time, so can answer
for the correctness of the sketch ; and I took the measurements my-
self.’ The drawing shows that it is the skull of an animal more
tf
a, 6. Skull and lower jaw of Ziphius Layardii. c. Teeth of lower jaw, from front.
allied to Ziphius micropterus than to Z. seychellensis. It differs from
Z. seychellensis in the lower jaw being elongate, slender, gradually
tapering in front, like the lower jaw of Z. micropterus. It differs from
the latter species in the tooth on the side of the jaw being elongated,
strap-shaped, with a small process* in the front side of the truncated
apex, and especially in these teeth being arched inwards, forming a
high arch “over the upper jaw,” the crown of the lateral teeth
being short and triangular in Z. micropterus. It is evidently quite
distinct in the form of the rostrum of the skull and the shape of the
teeth from the Ziphius micropterus of the coast of Europe. I
therefore propose to call it Ziphius Layardii.
The entire length of the skull, from condyle to tip of the rostrum,
* The process is not so distinctly shown as it ought to be in the woodcut.
15*
212 Zoological Society :—
3 feet 7 inches ; of the rostrum, from tip to the notch, 2 feet 6 inches ;
the width at the widest part of the brain-case 1 foot 6 inches; the
length in a straight line, from the tip of the rostrum to the crest
over the blower, 2 feet 11 inches; the height of the skull, from the
hinder part of the palate to the crest over the blower, 1 foot 2 inches.
The entire length of the lower jaw 3 feet ; the length from the con-
dyle to the hinder edge of the base of the tooth, 1 foot 114 inches ;
the length of the exposed part of the tooth along the anterior edge,
93 inches; the width below the teeth of the side of the lower jaw,
measured from the inner part of their base, 3 inches.
There is a partial hollow, as if it were the cavity of an old tooth
that had fallen out, on the margin of the inner jaw, behind the base
of the elongated arched tooth.
“In your letter you sent me a sketch of the skull of Ziphius in-
dicus with two teeth in the front of the lower jaw, and a short
stumpy head, totally unlike the skull of ZipAius figured in the ‘ Ca-
talogue of Cetacea.’”’
«There is a skull in the South African Museum which I have got
down as a Globiocephalus. It is the skull of a very old animal,
without teeth; but I think I can trace that it has had two front
teeth in the lower jaw, if not also along (the edge of) the upper and
lower jaw. The animal was taken on our coast.”
The figures of the skull which accompany this note appear to
me to represent the skull of a species of Hyperoodon, which differs
from Hyperoodon of Europe in haying only a low crest on each side
of the maxillary bones. I would propose to designate the species
Hyperoodon capensis.
\
s ‘ AN .
SSE Ehg SSSA EES Swab
2 SW GMM VEN
Skull and lower jaw of Hyperoodon capensis.
The length of the skull, from the end of the rostrum to the occi-
pital condyle, is 3 feet; the height of the skull, from the crest of
the blower to the condyle, 2 feet; the greatest width of the brain-
case 1 foot 7 inches.
Dr. W. Peters on Platacanthomys lasiurus. 213
April 25, 1865.—Dr. J. E. Gray, F.R.S., in the Chair.
Nore on THE SysTEMATIC PosiTION OF PLATACANTHOMYS
Lasiurus. By Dr. W. Perers, For. Memes.
Amongst the many interesting objects which have come under my
observation in the British Museum through the kindness of my
friends Prof. Owen, Dr. Gray, and Dr. Giinther, is a specimen of the
curious Rodent shortly noticed by Mr. Blyth (Journ. A. S. B. xxviii.
p- 289) under the name of Platacanthomys lasiurus. The specimen
in question is that exhibited by Mr. Sclater at a Meeting of this
Society in 1860*, and subsequently presented by him to the British
Museum.
It has always been difficult to me and other workers on the Mam-
mals to understand how a Rodent with only three molars in each
jaw could be referred to the Myowvina ; and I was therefore very
anxious to examine this very interesting form. But the results of my
observations will show that Platacanthomys does not belong to the
Dormice, but appertains strictly to the Murine family of Rodents,
being nearly allied in many respects to Phlwomys and Meriones.
The generic characters of Platacanthomys may stand as follows :—
PLATACANTHOMysS, Blyth.
Habitus myoxinus. Rostrum acutum, rhinario nudo, labro fisso ;
oculi mediocres ; auricule mediocres nude ; vellus molle, setis
dorsalibus latis sulcatis; artus mediocres, palme planteque
pentadactyla, digito primo abbreviato, falculis modicis curva-
tis, acutis ; cauda villosa, versus apicem fere disticha. Dentes
: . . . 33 . .
primores leves, compressi, acuti, molares utrinque =, complicati-
Cranium murinum, sed foraminibus incisivis parvis, coarctatis,
ossibus intermaaillaribus inclusis, palato perforato et processu
coronoideo brevissimo. Ossa antibrachii sejuncta, cruris con-
nata.
The resemblance of this genus to the Dormouse, at first sight,
is very striking, principally on account of the long-haired tail. But
in other respects, in its smaller eyes, very thin ears, and the well-
developed, although very short, thumb of the fore foot, it more ap-
proaches several Murine genera of Tropical India.
The skull is rather broad and flattened behind ; but it is quite
impossible for any one who knows anything about the craniological
characters of the Rodentia not to recognize at first sight the typical
form of the Murine, in the two- rooted zygomatic process of the
upper jaw, together with the peculiar form of the foramen infra-
orbitale, which is very high, narrowed, and widened above, and in
the development of supra-orbital ridges, which form together a lyri-
form figure. As peculiar and deviating from the typical skull of
the Murine, are before all to be noted the small and narrow fora-
mina incisiva formed only by the intermaxillary bones, the imper-
fect perforate palate, and the very short coronoid process of the
lower jaw.
* See P. ZS. 1860, p. 260.
214 Zoological Society :—
The incisors are narrow, compressed, and pointed. The molar
series are distant from and parallel to each other. The first and
second upper molars are nearly of the same size, and much larger
than the third and last. All three are composed of five enamel-
folds or laminze, obliquely directed inwards and hindwards: the first
and second of these are united as well on their inner as on their outer
side ; the third, fourth, and fifth are united on the inner side; but
on the outer side only the first and fifth enamel-folds are united.
The lower molars are of the same size as the corresponding upper
ones; but their enamel-folds are all united on the inner, and sepa-
rate on the outer side, except in the first (which has six enamel-
folds) the three anterior ones, and in the second and third (which
have four enamel-folds) the first and second ones.
PLATACANTHOMYS LAsIuRUS, Blyth.
P. magnitudine Muris ratti, auriculis acuminatis, capitis dimidio
longioribus, vibrissis longissimis ; supra umbrino-fuscus, subtus
albidus, jugulo pectoreque flavescentibus, cauda umbrino-fusea,
apice albido.
Long. a rostri apice ad caudze basin 0138; caudz 0™110.
Hab. India orientalis, prov. Malabar.
The size of this curious little animal is nearly the same as that of
the Black Rat. The head is rounded, rather flattened, with pointed
snout, naked muzzle, extremely long whiskers, eyes of moderate size ;
ears moderate, pointed, and, with the exception of a few scattered
hairs on the outer side, entirely naked. The fur is soft, on the upper
part, from neck to tail, intermixed with flat, longitudinally grooved
bristles. The limbs are proportionate and of moderate length, the
anterior shorter than the posterior ones. The fourth toe is the
longest, but only a little longer than the third ; the second and fifth
toes are much shorter, and nearly of the same length; but the first
is very short, and provided with a well-developed claw. The tail is
nearly of the same length as the body ; it is thickly covered with hair,
which is short on its base, and becomes more lengthened and disti-
chous from its second third.
metre
Total length RR ator ee ar 0°248
Distance from snout to base of tail ............ 0°138
Bengthrofthe headt wo ae mie cst. 6 atest 0:030
OF ENG CATS Ae core Ps etehioes teen eee ete 0°014
of the anterior extremity (from the elbow
to the end of the fourth finger) ............. 0°035
_ of the sole of the hand and fingers ...... 0-014
of the hinder extremity (from knee to the
fourth. toe). as sce ie rote eee eee 0°050
of the sole of the foot and toes.......... 0-025
Total leneth of the tail 05... 21a see ete te osc ee Oe
Length of the tail without hair................ 0°080
The specimen represented was obtained by the Rev. H. Baker, of
Mr. Ei. L. Layard on a new Zebra. 215
Mundakyum, Alipi, in Southern Malabar, who gives the following
note on the species (J. A. S. B. xxviii. p. 289) :—
‘“‘T was ignorant of the existence of this animal till about a year
ago, when rT found it in a range of hills about 3000 feet high. It
ee in the clefts of the rocks awl hollow trees, is said to Hoard ears
of grain and roots, seldom comes into the native huts, and in that par-
ticular neighbourhood the hill-men tell me they are very numerous.
I know they are to be found in the rocky mountains of Travancore ;
but I never met with them in the plains.”
May 9, 1865.—Dr. J. E. Gray, F.R.S., in the Chair.
The following extracts were read from a letter addressed to Dr. J.
E. Gray by Mr. E. L. Layard, of Cape Town, Corr. Memb. :—
“‘T send you herewith figures and descriptions of a new species of
Zebra. You have had a alae sent you* which you rejected as a
‘stray specimen of 1’, montanus, which had got down on the plains
and had been shot by accident’+. Iam sure you will, on perusal
of these notes, alter your opinion; and I shall be obliged to you to
read them at the Zoological Society. I wish to name the animal
Equus Chapmanni, after ‘its discoverer, my friend James Chapman,
who has done so much for African discovery, and who has hitherto
reaped no reward. I send you photographs of a horse and a mare
of this Zebra in different positions to show the markings, which differ
entirely from those of LZ. montanus (vel FE. Zebra) in the union of all
the black stripes with a medial one on the belly ; also on the back,
in wanting the ‘ gridiron’ pattern, as Baines calls it, on the rump.
I also send coloured sketches by Baines to show the colour. This
new animal also differs from the other Zebras in having the callo-
sities on the legs far larger and of a more rounded shape, in having
shorter and more equine ears, measuring only 6} inches instead of
113, and in having a shorter and more equine head and tail. The
hoofs also are flatter than in 2. montanus, and not adapted for
mountain-work. The mane grows several inches down on the fore-
head, and stands up between the ears, so that when seen in full face
it stands far higher than them. Chapman and Baines give the di-
mensions of several individuals; and all who have seen them here,
who are competent to judge from knowing the other species well, at
once detect the differences. I am quite convinced of them myself;
and, if you still doubt, please read this letter and the notes, and ex-
hibit the drawings, to the Zoological Society in my name. They roam
in large herds, and are first met with about 200 miles from the
coast inwards on leaving Walwich Bay, where Hquus montanus (or
rather a variety of that animal) prevails. 1 add some extracts from
* The skin sent me by Mr. Baines arrived in bad condition, with scarcely any
hair on it. It was that of a very young animal, and I could not see any differ-
ence, as far as I could judge in its very bad state, from that of a young Common
Zebra.—J. E. G.
+ I have no recollection of having made such a statement as the latter part of
this quotation—.J. E. G.
216 Zoological Society :—
the journals of Mr. Chapman and Mr. Baines relating to this
Zebra.”
“ Extract from Mr. J. Chapman’s Journal, dated May 21, 1862.
«The Quaggas here, I think, from about Sechellies’, though by
no means new to me, are different to any we see described in books
of natural history. The brush of the tail of one I shot to-day, and
which is rather a young specimen, is a dark grey, while the base is
white. In older specimens the brush is black, with a few white hairs
intermixed. It has a head band traversing the middle of the belly,
from which the transverse bands diverge alternately. The stripes
are of a very deep rich brown, nearly black ; while the ground-colour
is raw sienna on the upper parts (back, rump, sides, &c.), but gra-
dually fading into white on the lower parts. It has an erect mane
of alternate bands of white and black, edged with brown. ‘The ears
are white, with a dark band near the tip and broader band at the
base. The muzzle is grey or lead-coloured, and behind the nostrils
a brown coffee-colour. It has a bare spot on all four fetlocks, with
a brown crescent-shaped spot on either side of it. A bare patch above
the knee, on the inside of each fore leg. The pastern joints are brown,
excepting at the back, where it is divided vertically by a white line
from fetlock to hoof, The ears are decidedly equine. The mane is
6 inches long on the back; commencing from about 4 inches down
the forehead, extends to the length of 24 feet down the back. The
markings of it are continuations of the transverse lines which cross
the back. The white bands on the mane are quite superficial, the
hair underneath being actually black, edged with brown. Length of
ears 6 inches, The head measures 2 feet from the top of the skull
to the point of upper lip, From the root of the mane on the fore-
head and from top of forehead narrow lines of white and black (the
latter sometimes streaked with brown in the middle) diverge in a
triangular manner towards the eyes, where the outside lines, making
an angle, continue down the face, drawing closer towards the extre-
mity of the face (the inside lines being straight), where they blend
and form a dark brown patch behind and above the nostrils, the
muzzle and the lips being grey. Broader bands emanate from this
dark muzzle, and cross the chest in a crescent shape, leaving a white
margin around the eyes, behind which the regularity of the lines is
interrupted by those of the neck ; and the space from below the eye
is filled up with markings of a hieroglyphical character. The stripes
under the chin are light brown. ‘The circumference of the neck is
2 feet. The dorsal line extends to the brush of the tail, which is of
a dark grey ; and on the base of the tail, which is white, it becomes
narrower, and is dotted all the way down on either side with spots of
black, edged with brown. The form of the tail approaches nearer to
that of the Horse in the largeness of the brush than the Zebra or the
Ass ; but it is still not exactly like a Horse’s tail. On the thighs
the stripes are alternately pale brown and deep brown, horizontal, but
curving and forming a right-angled triangle on the flank ; and an
acute and more perfect triangle is formed on the shoulder-blades by
Mr. E. L. Layard on a new Zebra. 217
the junction there of the stripes from the neck and breast with the
transverse stripes. A longitudinal dark band traverses the whole
length of the belly, becoming narrower and deeper on the breast,
around which it winds and continues, forming one of the oblique lines,
to the centre of the shoulder-blades. From out of this ventral line
diverge the transverse lines tending towards the dorsal line, but not
connected therewith. On the legs the stripes gradually assume a
horizontal direction from the top downwards, but continuing the ob-
lique direction longer on the hind legs, and are distinctly, though
sometimes only faintly, visible to the hoofs in this specimen. Others
are more strongly marked. In some cases the transverse lines do
run into the dorsal line; but in no two specimens do the markings
seem to be exactly alike, the lines sometimes branching into two or
three as they approach the dorsal line on the flank and the angle at
the junction of the horizontal or oblique lines, these with the trans-
verse being sometimes filled up with disconnected hieroglyphical
characters.
«The height of a young male shot in June 1862, at the shoulder,
was 43 feet, at the rump 5 feet.”
** Notes of a supposed new variety of Quagga observed on the ele-
vated flats between the Botletle and Zambesi Rivers during the
late journey of J. Chapman and T. Baines. By T. Baines.
«Extract from my diary :—
«© 20th May, 1862.—Chapman had shot a Quagga answering most
nearly to the Bonte Quagga or Burchell’s Zebra, which is striped
over the neck and body, the legs only, from the knees and houghs,
being white; in this, however, faint markings were continued all the
way down, and a peculiar line was run along the centre of the sto-
mach, making me think it must be a new variety. Unfortunately it
is already cut up by Damaras and Bushmen.
“* As nearly as I can remember, Chapman, on returning, remarked,
‘The Quaggas here are not like those of Vaal River ; they have stripes
on their legs ;’ then said, ‘and if they are not Zebras they must be
new, for only two kinds are described—the common one of Kafirland
with no stripes on the rump or legs, and L. Burchellii, the Bonte
Quagga, with no stripes on its legs’ *. Chapman considered they were
not Zebras (as the animal is called here), L. montanus having longer
ears and asinine head and tail, whereas the head and ears of these
were more like those of a Horse, and the tail more bushy. Besides
this, #. montanus is strictly confined to hills and broken ground,
while these live in immense herds on the flat, with no mountains
within many days’ journey. We determined on further investigation.
* Latitude of the camp 20° 5! 55" south.
“ June 26th.— Chapman shot a Quagga strongly marked, like the
former ones, on the parts of the legs that are usually white; he sent
* This passage reads obscurely. Baines means only two kinds of Quagga: the
hunters call &. guagga and E. Burchellii ‘ Quaggas,” while E. montanus they call
“ Zebra.”—E. L. L.
218 Zoological Society :-
to let me know; but John, who has no idea of anything that has
not a market value, had called the Damaras to cut it up.
“‘ 30th.—The head and legs of a Quagga were brought in, the
latter being, as before, strongly marked quite to the hoofs, the re-
currence of this peculiarity showing that it cannot be a mere indi-
vidual accident, such as is seen in difference of colour in domestic
animals.
“July 10th.—Chapman shot a Quagga and Sable Antelope at a
distance from the waggons. I sketched from the skin and horns of
the latter, and the legs and ears of the Quagga. This had been a
smaller animal, but of stouter and more compact build than those
hitherto seen. I have already mentioned those at the Salt-pan with
decided markings on the legs below the knees and houghs, while the
two described species are perfectly white; and now this animal,
besides being stouter and shorter of limb, is more strongly marked,
the colours being distinct and pure black and white, the black spread-
ing almost half over the pastern-joint and fetlock, and having a small
white edging between it and the hoofs; the ears are strongly banded
and slightly tinted with brown. I thought at first it might be a
Zebra; but Chapman considered it a true Quagga, and I am inclined
to think so too.
«This was at Daka (lat. 18° 40'1!"). After coming down off the
elevated plain into the mountainous valley of the Zambesi system,
we were encamped on one of the spruils of the Luisi, the first run-
ning water we had seen since leaving the Botletle river.
« Thursday, 17th, Matietue River.—Chapman had shot a Quagga
mare ; and, hastening to the spot, I found an eager group of natives
with difficulty restrained from rushing at once upon the prey. In
this case we had to omit the measurement ; but I sketched the stripes
carefully, and the camera of course cannot be gainsaid. The general
colour was a yellowish or raw-sienna brown on the upper parts, and
deepest on the rump, fading into white on the neck, belly, and legs ;
the stripes were of the deepest brown or nearly black, and the dif-
ference between this and the known varieties consisted in their being
continued quite down to the hoof on all four legs, slightly fainter on
the inside ; the belly was marked by a broad black band along the
centre, to which all the side stripes were joined ; on the back was a
similar black line, but only the stripes above the shoulder were con-
nected with it; the mane was upright, as usual (the neck-stripes
being continued vertically through it); the ears small and equine,
and a bare spot (rather small) was observable on the inside of the
fore legs only, the Zebra, I believe, having it on all four, as well as
large ears.
“T made two sketches of this, and Chapman two photographs.
There are intermediate brown stripes between the black ones on the
hind legs above the hough.
“Saturday, July 19th.—We proceeded about a mile north-east
by north, when, near the small conical hill on our left, Chapman
brought down a fine young Quagga stallion of the same kind as
the mare previously killed ; but age, I suppose, not having deepened
Mr. E. L, Layard on a new Zebra. 219
the colours, its whole body was of the purest white, marked with
jet-black bands down to every hoof, in the manner of the other, but
slightly fainter on the inside of the legs, and also where the stripes
of the sides joined to the longitudinal line of the belly, some of
those on the flanks having these points so faintly marked that the
junction could not be called complete ; like the other, a central stripe
ran along the back, with which two or three of the shoulder-stripes
(on each side) were connected, the broad stripes of the hinder parts
originating near the central line about the insertion of the tail, and
diverging laterally over the hip, flank, and side till they completely
or nearly reached the ventral line, the longest of them meeting on
their way the ventral stripes of the sides, and forming the most
beautiful possible combination of curves and angles, even the slight
variation of regularity on either side conducing to the effect; the
ears were small, and banded and tipped with black and dark brown ;
the head well shaped, with a little sienna-brown towards the nose ;
and the whole form lighter and more elegant than in the older spe-
cimens.
“Sunday, September 14th.—I shot two, which at first I took
for Mountain-Zebras ; but on comparing notes with Chapman, I came
to the conclusion they were also Quaggas. The stallion fell at a di-
stance, and was cut up while I was sketching and observing the mare.
She was full-striped, somewhat smaller than most of those Chapman
had killed; ears, if anything, shorter and more equine. Callosities
or small bare patches of skin on the inside of the fore legs only, and
not on the hinder legs; striped right down to the hoofs ; inside more
faintly marked than the outer. Dokkie and others thought it like
the Wilde Paard of Ozembengue, and different to the Quacha of the
plains. I believe they would have said anything, so that I would
have done talking and let them begin to cut it up.
“Sunday, December 7th.—Went out from Logu Hill, Zambesi
River ; tracked spoor several hours ; wounded a mare, which was run
down late in the afternoon, and killed with a stone. Fully striped,
as before, down to the hoofs, all four legs, the inside of the forearm
and thighs being more faintly marked; the ears small and tipped
with black ; the stripes on the sides extended from the dorsal line
to the ventral, which last, reaching from between the fore legs to the
hinder, was of not quite so deep a black ; the ground-colour was light-
yellowish brown on neck, back, and sides, passing into white on the
cheeks, throat, and under parts of body; the teats, two in number,
were situated in the after part of the black ventral line. She had
warts or callosities on the inside of the forearms only, and none on
the inside of the thigh.
«I sketched carefully, and took the skin home, attempting to pre-
serve it; but the weather was so damp that, even in a hut with a
fire in it, I could not dry it.
«Tuesday, 14th April, 1863 (after our return to the salt-pan on
the elevated plain between the Zambesi and Botletle rivers).—A
few Quaggas were standing on the further plain, and creeping behind
a point at 300 yards’ range. I shot one through the neck and fore-
220 Zoological Society.
head: it proved to be a well-grown, handsomely marked filly of the
first year; and as the rest retreated, I noticed that the mare hung
back and looked frequently round for her lost little one, returning
when the others were out of sight and gazing wistfully at the spot
where it lay.
««T had no means of measuring the beautiful little creature on the
spot ; and for convenience of carrying I had only my small sketch-
book, so carefully outlined one of the fore legs. I sent Pompey
back for assistance, and in the interval sketched on a small scale, and
stripped off the skin, which is a good size for a small museum, and,
as carriage is a consideration, suits me better than a large one.
“It is perfectly marked after the manner of Quaggas in this loca-
lity, but not so fully as those of Daka and the Zambesi, and is most
certainly an intermediate link between already described varieties
and the Zebra. The chief points worthy of note are that the legs,
instead of being white as in the Bonte Quagga (2. Burchellii) from
the houghs and knees, are marked with transverse bands, not so dark
as those on the body, quite down to the hoofs ; there is a dark stripe,
commencing between the fore legs and extending along the belly to
between the hinder, where it becomes broader and somewhat fainter ;
the first three stripes behind the shoulder are joined to this; the
dark stripes on the rump are alternated with others of a medium
brown, but those on the fore part of the body and neck are of a full
deep black ; there are callosities on the inside of the fore legs only,
and none on the hinder.
“Chapman killed two Quaggas during the day. I believe they
were very faintly marked on the legs ; but the vultures and Damaras
destroyed them. The skins are quite worthless, which is much to be
regretted, as we think it certain they are true Quaggas undescribed
in any work we know of, and, as a new variety, would have been a
handsome gift to any museum.
« Pereira told me subsequently, the Quagga of Damaraland has
legs very nearly white ; there are faint stripes, but not visible till you
come close to them; there are warts on the fore legs only. The
Wilde Paard is darker, the stripes blacker; the head is larger, and
the ears also; they stand up so as to be visible above the mane. The
Wilde Paard goes in the hills, the Quagga on the flats.
“T sent down the skin of the filly to Mr. Logne in Cape Town,
and he forwarded it to the British Museum.”
With reference to this communication, Mr. Sclater remarked that
the female Zebra in the Society’s Gardens (presented to the Menagerie,
May 26th, 1861, by H.E. Sir George Grey), which he had hitherto
referred to Equus Burchellii, appeared to answer the description
above given in every way, and must probably be referred to Equus
Chapmanni if that species were allowed to stand.
221
MISCELLANEOUS.
On the Chilian “Anguilla.” By Dr. R. A. Parviprr.
Dr. Puixrepr has succeeded in obtaining a specimen of the fish
known under the name of ‘ Anguilla” in Chili: it is a new species
of Lamprey, which the author describes under the name of
Petromyzon acutidens.
It is much darker than the three other Chilian freshwater Lam-
preys; above and on the sides blackish grey, with a violet and rusty-
brown lustre, the latter especially on the tail. Each branchial ori-
fice stands in the middle of a whitish spot. The ventral surface is
grey, yellowish beneath the branchial orifices. The caudal fins are
blackish grey ; the two dorsal fins rather light grey. Seen from the
side, the muzzle appears rather acute, the mouth being almost in a
line with the belly ; its hinder end projects somewhat, and is sepa-
rated from the gular region by a transverse fissure, nearly 3 lines
broad, which leads into a sort of shallow pouch. This does not form
a sac, as in Petromyzon? Anwandteri and Velasia chilensis, but is
somewhat inflated. The total length is 14 inches ; the depth at the
last branchial orifice is 9 lines, at the first dorsal 7+ lines, and at the
anus 5 lines. The eye is 12 lines from the apex of the muzzle, and
2 lines in diameter ; the orifice of the mouth is 114 lines long ; tie
first branchial orifice is 19 lines from the tip of the snout, and the
last nearly 3 inches. The first dorsal commences 7 inches from the
tip of the snout, and is 13-14 lines in length and 23 lines in height.
The second dorsal is of the same height, but more than 2 inches
long; the interval between them is 14 inch. The caudal fin is
acutely rhomboidal; its dorsal margin is 14 inch long; its ventral
portion runs, gradually diminishing, nearly to the anus, which is
2 inches 4 lines from the extremity of the tail.
On each side of the head there are three rows of mucus-glands :
one runs from the snout towards the lower margin of the eye, but with-
out attaining the latter; the second forms an oblique line close to the
antero-inferior margin of the eye; and the third commences below
the first, halfway between the apex of the snout and the eye, and is
continued to the throat, where it terminates between the hinder
margin of the mouth and the first branchial orifice.
The mouth forms an ellipse, or, when fully extended, a broad
oval, and has double lips, the outer grey, with a row of small warts,
the inner white, short, and entire at the margin. The teeth are
remarkably acute. In front of the two inner lingual teeth there
stands a transverse row of eight teeth; on the palate there are two
groups, each consisting of three acute teeth; and, lastly, there are
about four concentric series of acute denticles, gradually diminishing
in size from the gullet to the margin of the lips.
In the two groups of three teeth, and the lips destitute of fringes,
the species resembles P.? Anwandteri, which, however, has a row of
large teeth in the external circumference of the mouth, and is further
222 Miscellaneous.
distinguished by a large gular sac (as in Velasia) and by the different
form of the caudal fin. The fish inhabits the brooks of some parts
of Chili, and is thrown away by the fishermen, who regard it as
unwholesome.—Wiegmann’s Archiv, 1864, p. 107.
On the Parasitic Nature of the Mistletoe.
By Joseru Borum.
The author divides plants in general into the two following groups:—
1. Chlorophyll-bearing, which assimilate the inorganic substances
drawn up by the roots from the soil, and thus become the ancestors
of all the rest of living nature.
2. Chlorophyll-free, which either extract the assimilated juices
from other organisms, or nourish themselves from dead organic
matter. The latter plants alone, which live in the manner of ani-
mals, are regarded by the author as parasites.
The Mistletoe has always been regarded as a plant which extracts
the organic juices from the plant on which it grows, and consequently
leads a parasitic existence. Boehm calls attention to the following
circumstances, which are particularly adverse to this view :—
1. The mode of insertion of the roots of the Mistletoe into the
wood of the tree on which it grows.
2. The occurrence of the plant in question upon more than thirty
species of trees, all, however, of indefinite growth (Hndumsprosser).
3. The different results of the analysis of the ashes of the Mis-
tletoe and its supposed nutritive plants.
4. The comparative size of the branches bearing Mistletoe above
and below the insertion of the apparent parasite.
Recent investigations, repeated by Boehm, have placed it beyond
a doubt that, in trees with indefinite growth, the ascent of the crude
nutritive material takes place in the wood, but the assimilated forma-
tive juices descend in the bark. Even Knight was aware that when
annular strips are removed from the branches of these plants, the
latter become thickened only above the annular wound.
This circumstance enabled the author to decide with absolute cer-
tainty that the Mistletoe has precisely the same relation to its nutri-
tive plant as a twig to its parent branch, or the graft to the stock.
From thirty branches bearing Mistletoe (on Acer, Populus, and
Quercus) the terminal twigs above the attachment of the Mistletoe
were cut away and the branches ringed below the Mistletoe. Whilst
in Acer and Quercus the branches thus treated usually died soon,
the Mistletoe plants on the Poplars not only continued their normal
growth, but a thickening of the branch above the annular wound
took place. ‘This can only have occurred at the expense of the
juices assimilated by the Mistletoe.
The fact that the development of the branches adove the insertion
of the Mistletoe is hindered has, in the author’s opinion, nothing to
do with the parasitic nature of that plant. The Mistletoe acts only
in the same way as any branch of the tree of which the development
is in advance of its neighbours. The injurious effect of the presence
Miscellaneous. 223
of Mistletoe upon the growth of the twigs below it is to be ascribed
partly to the aborted condition of the terminal shoots, and partly to
the fact that the juices assimilated by the Mistletoe are chiefly ap-
plied to its own increase, and may be less fitted for the development
of the tree on which it grows.——Bericht der Akad. der Wiss. in Wien,
June 30, 1865, p. 113.
On a Fungus which is developed in Ivory and Bone.
By Proressor WEDL.
In examining some sections of human teeth which had been
macerated for a few days in water, Professor Wedl found that the
cement and the peripheral layers of dentine were furrowed by micro-
scopic channels. He soon recognized in these channels small parasitic
plants, closely resembling those which perforate the shells of Mol-
lusca. A careful examination of the water in which the sections had
been macerated furnished numerous small cells, which might be re-
garded as the spores of the Fungus. Fragments of normal teeth
placed in the same water were soon infested by these little parasites,
the operation of which is, however, confined to the cement and
dentine, and never extends to the enamel. The Fungus also
attacked fragments of bone macerated in the water.
These little Fungi seem to be developed at the expense partly of
the organic and partly of the inorganic matter of the ivory and bone ;
and the conditions of their multiplication doubtless frequently occur
in nature. They do not, however, appear to attack teeth until
after death; so that they have nothing to do with caries. Pro-
fessor Wedl has ascertained that these parasites have been in action
from a high antiquity, many teeth of fossil Fishes and Mammalia
exhibiting unequivocal traces of their action.—Sitzungsber. Akad.
Wiss. in Wien, July 14, 1864; Bibl. Univ. 1865, Bull. Sei.
p- 231.
Note on the Ammobroma Sonore.
This (the literal translation of which is ‘‘sand food of Sonora’’) is
the name of an extraordinary root-parasitic plant, of the region at
the head of the Gulf of California, which Dr. Torrey has just de-
scribed and figured in the eighth volume of the ‘Annals of the
Lyceum of Natural History of New York.’ It has been briefly
noticed before (but never fully characterized) as a new genus allied
to the rare Mexican Corallophyllum of Kunth (or Lennoa, Lexarza),
and still more to the Californian and hardly better known Pholisma
of Nuttal. It hardly throws any new light upon the affinity of
these strange plants, which, though justly thought to be rather
Monotropaceous than Orobanchaceous, are still obscure. This
plant, growing in a forlorn sandy desert, almost covered by the sand
in which it lives, was found by its discoverer, the late Col. A. B. Gray,
to form a considerable part of the sustenance of the Papigos Indians
of the district, and is said to be very luscious when first gathered
and cooked, resembling in taste the sweet potato, only far more de-
licate.—Silliman’s Journal, July 1865.
224 Miscellaneous.
On the Intercellular Matter and the Vessels of the Latex in the
Root of the Dandelion. By Dr. A. VOGEL.
The back of the root of the Dandelion is traversed by a great
number of lactiferous vessels containing a very abundant bluish-
white milk. These vessels form a great number of larger or smaller
bundles disposed in a tolerably regular concentric manner, and
united to each other by numerous ramifications. The ramifications
are always parallel to the surface of the root, so that the bundles
form concentric sheaths independent of each other. The outermost
peridermic layer of the bark, however, is destitute of lactiferous
vessels, which exist chiefly in the inner portion. The cellular paren-
chyma of these two parts of the bark contains a great quantity of
intercellular matters, especially in the vicinity of the lactiferous vessels.
According to the author, the lactiferous vessels originate by the
union of conductive cells (Lettzellen), of which the adjacent septa
become gradually converted into pectose, and finally disappear. He
has detected many intermediate stages, which leave him no doubt as
to this mode of formation of the vessels. According to him, the
lateral walls of these same cells likewise finally become converted
into pectose; so that the fully developed lactiferous vessels are not
formed by a cellulose membrane.
It was by observing the action of iodized liquids, acids, and alka-
lies upon the substance forming the envelope of the lactiferous ves-
sels, that Dr. Vogel was led to regard it as cellulose in progress of
conversion into pectose. He arrived at the same results by setting
the same reagents in action upon the intercellular matter of the
Dandelion-root, which he also regards as pectose produced by the
gradual transformation of the membrane of cells—Bericht Akad.
Wiss. in Wien; Bibl. Univ. 1865, Bull. Sci. p. 239.
On the Structure of the Luminous Organs in the Male of Lampyris
splendidula. By M. Scuu.rze.
The author has found that the numerous branches of the tracheze
in the luminous organs of Lampyris splendidula terminate each in a
small cell of stellate form. Under the action of osmic acid these
cells rapidly acquire a black tinge, whilst the cells of the parenchyma
remain uncolonred. These cells therefore readily reduce the osmic
acid by absorbing its oxygen; and the author attributes to them an
important part in the production of the phenomenon of phosphores-
cence by this insect.—Sitzung der Niederrhein. Ges. fiir Natur- und
Heilkunde zu Bonn, 1864 ; Bibl. Univ. 1865, Bull. Sci. p. 232.
De Jeude’s Collection of Mollusca.
The fine collection of Mollusca formed by the late Prof. Lithe de
Jeude, for many years Professor of Zoology in the University of
Utrecht, has been purchased by Mr. Damon (of Weymouth). The
collection, rich in the rare shells of the Moluccas, was displayed in
140 glass cabinets, and formed one of the chief scientific attractions
in the city of Utrecht.
THE ANNALS
MAGAZINE OF NATURAL HISTORY.
[THIRD SERIES.]
No. 94. OCTOBER 1865.
XXVII.—On Ammonites from the Cambridge Greensand.
By Harry Szetey, F.G.S., of the Woodwardian Museum,
Cambridge.
[Plates X. & XI.]
Ammonites (Scaphites) equals, Sow.
Shell much inflated, convex, with a wide back, and the convo-
lute portion so coiled as not to produce an umbilicus: this part
forms about half the length of the shell, and is always half the
width of the back, or wider. The back is about twice as wide
as the side, and less convex. When the whorl recurves and
forms the mouth, it contracts.
Both spire and hamus are marked with fine elevated ribs,
which are most elevated on the sides, and bifureate before cross-
ing the back. On the spire they are curved slightly away from
the mouth, so that the lines are concave in front ; on the hamus
they pass over straight, and are separated by wider concave
channels.
The symmetrical septa consist of a rather small square dorsal
lobe, with two small notches on each side, and two digitated
terminal branches. The dorsal saddle is enormously wide, eX-
tending to the limit of the back, where the ribs bifurcate ; ‘it is
centrally cleft by a branch half as large as the dorsal lobe. The
superior lateral lobe is as wide as half of the dorsal saddle, has
one small notch on each side, and terminates in two large
branches, which bifurcate, are digitated, and are near together.
The inferior lateral lobe is small, and at the base of the side.
On the ventrum are the ventral lobe and four pairs of acces-
sores.
The Scaphite of the Cambridge Greensand are abundant in
individuals, though few in forms. Authors have generally, and
perhaps rightly, referred similar fossils to the S. equals of
Sowerby.
Nearly all the specimens found are the last chamber, or
Ann. § Mag. N. Hist. Ser. 3. Vol. xvi. 16
226 Mr. H. Seeley on Ammonites
hamus, which may indicate that, after these had become filled
with phosphate of lime, the partitioned part was broken off and
floated away at the surface, much as the recent Spirula is found
drifting. Perfect shells are rare, and young ones never found.
There are three variations from this form: one, which is
ornamented with a row of tubercles on each side, is also found
in the Gosau Chalk.
Ammonites rostratus, Sow.
Ammonites rostratus, Sow. A. symmetricus, Sow.
Numerous citations have been given by Continental writers,
to whom the species seems scarcely known, being regarded as
synonymous with A, inflatus, Sow.
Shell keeled, few-whorled, with a flat back, flat sides, quadrate
mouth, and umbilicus becoming relatively smaller with age, but
never less than the height of the body-whorl; not very deep, it
is angular, though not making a sharp angle with the side,
The back is generally a little narrower than the side is high ;
out of its middle arises the narrow keel, from each side of which
the ribs are directed backward and outward at an angle of 45°,
then descend straight, elevated, perpendicular, and separated by
wide spaces, and terminate, generally separate, im tubercles at
the base. On a whorl there are from 25 to 35 ribs, each
having a tubercle above the middle, and another where they
reach the back. The wide intercostal spaces are smooth. When
full-grown, the last two or three ribs incline frontwards; the
keel and adjacent lateral parts of the back become elevated, and
are directed upward and forward in a curve to form the rostrum,
which is hollow and rhomboid in section; for the last mb but
one, instead of dying away on nearing the keel, as those behind
it do, is continued up its side, becoming less and less distinct.
The whole whorl becomes narrow. This rostral prolongation is
open in front, and in its upper third curved slightly down.
The septa are symmetrical. The dorsal lobe is longer than
wide, extending between two tubercles, and being margined by
the rising of the ribs. It has three notches on each side, the
two terminal of which become with age digitated branches.
The dorsal saddle is relatively very wide, extending down to the
lateral tubercle; it is doubly cleft, the upper accessory being
much the larger, above the middle, and in a line with the dorsal
tubercles. The upper lobe gets relatively wider with age, and is
from a third to more than half the width of the dorsal saddle ;
it has a branch on each side, and two terminal branches digi-
tated with age. The lateral saddle is slightly wider than the
superior lateral lobe. The lower lateral lobe 1s on the ventral
angle, and very small.
from the Cambridge Greensand. 227
Besides this, there are four rare forms varying from the type
chiefly in the degree of development or suppression of the ribs
and tubercles.
This Ammonite is found on every digging along a line of
thirty miles, and is the most abundant. The whorls are always
coiled on the dorsal tubercles or the homologous thickenings of
the ribs; so that the size of the umbilicus depends on the rate
of increase in the height of a whorl. As a general rule, the
more compressed the shell, the closer the ribs.
In a very young state, the whorls are smooth and round, and
continue so for three or four whorls, and then rapidly assume
the typical form; and as this part appears devoid of septa, it
might have been formed in the egg.
Large specimens are very rare, and the largest found are
much inferior in size to those of the Gault at Folkestone or the
Upper Greensand of the Isle of Wight and Devizes, though
there can be no doubt that some Cambridge specimens reached
a diameter quite as great as that of their southern brethren.
Part of a whorl with the rostrum attached is only 13 inch high.
Broken rostra are comparatively common ; and it can only be
supposed that the large shells to which they belonged, having
but weak septa and being filled with phosphate of lime, were
broken up before fossilization. The result was not the production
of the numerous small examples; for, as the last half whorl is
almost invariably devoid of partitions, these died small if not
young, and there is no evidence that a rostrum was ever formed.
It is probably the true A. symmetricus of Sowerby; but that
type is seemingly now lost.
Ammonites pachys. Pl. XI. fig. 4.
Keeled, few-whorled ; whorls flattened on the sides and back,
so as to appear in section nearly quadrate [about half-embracing].
Umbilicus as high as the back is wide, and higher than the side
of the last-formed chamber; it is deep, with the ventrum nearly
at right angles with the side of the whorl, where it is rounded :
this space is marked (at a diameter of 24 inches) with about
twenty-two strong rounded ribs descending from as many large
spinous tubercles which margin the base of the side. From
each tubercle two ribs (rarely three) ascend the side of the
whorl, at first but little elevated, but higher and wider near the
back. The interspaces are never wider than the ribs, and on
the back are narrower; on the sides, owing to a curve in alter-
nate ribs, they are equal. On the dorsal surface the ribs are
curved anteriorly, and become obliterated near the keel. On the
back (and in old specimens on the sides too) the ribs are crossed
transversely by small elevated lines placed rather close together.
16*
228 Mr. Il. Seeley on Ammonites
The back is a little inflated. The keel is round, wide, moderately
elevated, with the smooth spaces on each side slightly depressed.
The mouth is a little wider than high.
Septa very like those of the southern forms of A. rostratus (to
which the form is nearly related), but with the lobes deeper, and
with finer digitations. The dorsal. lobe is as long as the back
is wide, extends in width to the line where the ribs arise, has
two rather narrow branches on each side, and terminates in two
acute branches conspicuous in having no digitations on the
inner side. The lateral lobe is in the middle of the side, not
quite so long as that of the back, with two branches on each
side, and terminating in three, of which the middle one is much
the largest. At a diameter of 13 inch the distance between two
septa where they cross the keel is #ths of an inch. In examples
of the southern form of A. rostratus, at a diameter of 24 inches
the interseptal space on the keel is only half an inch.
Much as it differs, I incline to regard this as a constituent
variety of A. rostratus, with near affinities to A. inflatus.
Ammonites Timotheanus, Pictet, Grés Vert, pl. 3. figs. 1 & 2.
Few quadrate whorls, enlarging rather rapidly, two-thirds
embracing, gibbous ; with a flat back, flattened sides, and flat
ventrum. ‘he sides, very slightly inflated, round into the um-
bilicus and into the back, shghtly converging, so that the back
is narrower than the base. The mouth is nearly a fourth wider
than high. The umbilicus is not shallow, and is half as high as
the mouth is wide. The cast is perfectly smooth, and only
marked with the elegant foliations of complex septa.
The siphuncle is unusually wide, being of the width of the
dorsal lobe. The septa are almost exactly the same as in A.
latidorsatus (Mich.). The dorsal lobe is relatively a little deeper
by the terminal branches being separated for only half as far ;
these and the preceding branch on each side are bifid. The
superior lateral lobe is deeply cleft, making the two bifid ter-
minal branches large. The superior lateral saddle is not deeply
cleft.
[ have seen but two specimens of this form, both collected by
Mr. Carter. So far as figures and descriptions enable me to
judge, it might be classed as a variety of A. latidorsatus, differing
chiefly in the flatness of the back, and perhaps in the rate of
enlargement; but it appears, from Michelin’s figure, that the
young states are sufficiently unlike to justify a distinction. This
form always had a flat back and trapezoidal mouth, &c., while
the other has a round back and lune-shaped mouth, It exactly
corresponds with the Ammonite figured by Prof. Pictet in the
‘Gres Vert’ (pl. 2. fig. 6) as A. Timotheanus; but it does not
from the Cambridge Greensand. 229
correspond with his description and the figure answering for
Mayor’s MS. species. That form is spoken of as “ornée de
cing & six sillons qui partent de Pombilic dans Ja direction
qu’auraient des tangentes au cercle ombilical.” This character
may perhaps not be sufficient to distinguish a species; but it
certainly makes a well-marked form, indicating au animal which
formed periodic varices.
Ammonites latidorsatus, Mich.
A, latidorsatus, D’Orb. Paléont. France; Pictet, Grés Vert, pl. 3. fig. 5.
Few-whorled, much inflated, whorls two-thirds embracing, a
little wider than high, with nearly parallel flat sides, and a
rather depressed round back, which rounds into the sides. The
body-chamber is half as high again as the whorl at the opposite
side of the shell, than which the umbilicus is slightly narrower.
The umbilicus is deep, with the flat and almost horizontal ven-
trum making a sharp angle with the side, though the line of
meeting is just rounded. The cast appears perfectly smooth,
but is marked by shallow and narrow suleations arising in the
umbilicus and passing over the back, where two are separated
by interspaces narrower than the height of the whorl. The
mouth is lunate.
The septa are like those of A. planulatus, the only difference
being that the inferior lateral lobe is not cleft so deeply.
This is a very rare fossil; and the only specimen known to
me was detected in the collection of the Rev. J. F. Blake, who
has presented it to the University Museum. The diameter is
1+ inch, and the last fourth of the outer whorl is devoid of
chambers. It corresponds with the Continental figures and
description, though this sulcated form, which cannot be con-
sidered typical, is a variety distinguishable from Michelin’s
smooth shell.
Ammonites Mayorianus, D’Orb.
A. planulatus, Sow. Min. Con.; Sharpe, Chalk Moll. pl. 12. fig. 4.
A. Mayorianus, D’Orb. Paléont. France, T. C
A. octosulcatus, Sharpe, pl. 19. fig. 3.
A. Griffithsii, Sharpe, pl. 11. fig. 3.
Inflated, with few whorls, more than half embracing ; mouth
as wide as high. Sides a little inflated, and passing impercep-
tibly into the round back. Umbilicus about as high as the
body-whorl, with the horizontal but inflated ventrum rounding
into the side. The depth of the ventrum exposed is about the
same as the width of the unembraced part of the adjacent whorl,
with which it forms a right angle. The cast commonly shows
very fine depressed ribs, arising about the middle of the side,
and passing over the back; they are close together, and on
230 Mr. H. Seeley on Ammonites
nearing the back are curved mouthwards. In the umbilicus
arise four or five more or less deep, wide suleations, which are
mostly flexuous, being first directed a little forward, then
perpendicularly upward, and finally curved forward on the back
to the siphuncle, the two sides meeting there m a broad V-shape
and dying away without impressing it. Between each two sulci
there are commonly from twelve to eighteen ribs.
The septa consist of the dorsal lobe, three lobes on each side,
and one or two in the umbilicus, all except the last much
branched, digitated, and close together. The dorsal lobe is
longer than wide, has three notches on each side, and terminates
in two large branches, which are cleft laterally, the parts which
continue backward being also eleft. The superior lateral lobe
has two or three small notches on each side, and terminates in
three large branches, which are trifureate and digitated. The
dorsal saddle is larger than the lobe; it is centrally cleft, and
has a few notches, besides each half being cleft. The inferior
lateral saddle is much smaller, but similar.
This form, which differs a little from those most common at
Cambridge, has been described first, because one of the types of
it is the identical specimen figured by Mr. Daniel Sharpe as A.
planulatus, and, moreover, it is the most inflated modification
that occurs. The example figured, Palzeont. Monogr. Cret. Mol.
pl. 12. fig. 4, had septa to the end ; and so has one of 4 inches
diameter, the largest I have; but fragments with the shell pre-
served occur, showing the back of a whorl to have been from 4
to 5 inches wide. The small line-like ribs passing over it are in
this large size a quarter of an inch wide, though but little ele-
vated, being separated by narrow shallow grooves. The sulca-
tions go on deepening and widening.
On comparing small examples with corresponding specimens
of A. Mayorianus, D’Orb., from the Upper Greensand of the
Perte du Rhone, I fail to notice any distinction: compare them
as we may, there is no character they have not in common ;
consequently no hesitation is felt in regarding our form as iden-
tical with those of France and Switzerland; but it is not so
absolutely identical with the Chalk fossil known as A. planu-
latus. I know that form from a beautifully distinct cast of the
umbilicus partly formed by an Hxogyra growing attached to the
specimen ; it is from the lower Chalk of Burwell in Cambridge-
shire. The umbilicus measured not less than 4 inches across,
and was shallow. The umbilical part of the whorls is flat, and
the depth of the ventrum exposed only one-fourth the width of
the unembraced part of the side. On the Hwogyra are the
foliations of the septa. And this enables me to state that the
sulci, which were straight on the sides. were not a sixth of the
from the Cambridge Greensand. 231
width or depth of the internal casts of those in A. Mayorianus,
and did not bend into the umbilicus, but terminated at the base
of the side. The ribs on the back are more elevated and wider
apart; the number of whorls fewer. The sulci are arranged
relatively to each other like those figured in ‘ Cret. Mol.’ pl. 12.
fig. 3. It will be readily seen, on referring to that plate, how
much better the above description coincides with the Lewes
Chalk fossil than with the ancestral race figured beside it.
Hence Sowerby’s name, specially used, should be restricted to
the Chalk form to which it was originally given; while D’Or-
bigny’s would with more propriety be preserved for the Conti-
nental fossil, with which ours corresponds.
In the variations from A. Mayorianus, the whorls gradually
become more compressed, with flatter sides rounding into the
umbilicus, which is relatively smaller. There are from five to
eight sulci. These specimens frequently occur with part of a
whorl devoid of septa at a diameter of an inch and a half. In
the young state (diam. ? inch) the sides are flat, and a little
converging to the back; and as the sulcations are scarcely im-
pressed, the shell has the aspect of A. Beudantit.
Some specimens, which died small, exactly correspond with
Mr. D. Sharpe’s figure of A. octosulcatus in the number of sul-
cations, their relative size, and the way they pass over the back :
though the form of the whorl is not quite the same, yet so near
are the two, that I have no doubt of the propriety of regarding
A, octosulcatus as a slight variety of A. Mayorianus.
There are no specimens at Cambridge of A. Griffithsii; but,
judging from Mr. Sharpe’s figure and description, it cannot be
regarded as other than a variation from A. Mayorianus.
The type of shell here discussed is one of the more abundant
of the Greensand Cephalopods, occurring in this neighbourhood
wherever that deposit is worked.
Ammonites Weistii, Sharpe, ‘ Chalk Mollusca,’ pl. 21. fig. 3.
Few-whorled, inflated ; back round; mouth semilunar, much
wider than high; umbilicus moderately open, with its deep
border forming a large angle with the side of the shell.
Ornamented with about twenty-two wide, straight ribs, nearly
all of equal length; every third rib is somewhat thickened on
the sides as it nears the umbilicus, while the ribs between these
thickened ones frequently become obliterated before reaching
the edge of the umbilicus.
Septa simple, consisting on each side of three lobes. Dorsal
lobe marked on each side with two simple digits. The dorsal
saddle, half as wide again as the dorsal lobe, is indicated by a
232 Mr. H. Seeley on Ammonites
small accessory. The superior lateral lobe is about half the
width of the dorsal lobe, and much shorter; the terminal
branches are similar, only less developed.
Only two specimens of this Ammonite have come under my
notice.
The exact affinities of A. Weistii are not quite clear. Speci-
mens of A. navicularis come very near to it, but have not the
constant greater elevation of occasional ribs; rather im this it
approaches 4. peramplus, which, however, has spmes in the
young state at their umbilical termination.
Ammonites navicularis, Mant., var.
The flattened back is slightly inflated, and rounds into the
side; the flattened side, which is also a little swollen, rounds
into the umbilicus. The mouth is higher than wide. The few
whorls are almost entirely embracing, forming a deep and small
umbilicus about half the diameter of the whorl opposite to the
mouth.
Ornamented with about (thirty to) forty wide rounded ribs,
which are straight, strongest where they pass over the back,
and separated by spaces of not more than their own width.
About half the ribs arise in the umbilicus, the remainder near
the middle of the side.
There are three lateral lobes on each side. The dorsal lobe
is wide and square, with three branches on each side, the lower
of which have five digits. Dorsal saddle rather wider than the
lobe. Superior lateral lobe half as wide as the dorsal saddle,
and deeper than the dorsal lobe; it has two branches on each
side and a large terminal one which bifurcates. Lateral saddle
like dorsal.
The few specimens found show considerable variation in the
form of the mouth, which is sometimes as wide as high. They
are more flattened than is usual in examples of A. navicularis
from the Chalk, and differ in never having any tubercles; the
umbilicus is also commonly smaller. The Warminster Upper
Greensand contains similar shells; but they have tubercles on
the back.
It may be necessary to separate the Cambridge shell as a
variety ; for it is intermediate between A. Weistit and A. navicu-
laris, and may be an extremely compressed variety of the former.
But A. Weistit can scarcely claim to be more than a well-marked
variety of A. navicularis, connecting it with A. peramplus. Our
fossils also nearly resemble A. vectensis, Sharpe, and cannot be
distinguished as more than a variety, the only differences being
that the ribs are straight (seemingly more elevated), the back
slightly flatter, and the umbilicus commonly larger. This being
from the Cambridge Greensand. 233
so, confusion may be avoided by marking Cambridge specimens
A. navicularis, var. nothus.
Diam. 2+ inches, with septa to the end.
Ammonites rhamnonotus. Pl. XI. fig. 7.
Few-whorled, flat, with a round back and small umbilicus.
Ornamented with about thirty-six radiating ribs, which con-
tinue uninterruptedly over the back, and are alternately long
and short. The long ribs mostly arise in the umbilicus, and the
shorter ones at a third or half the width of the whorl from the
back; they are nearly straight, elevated, and become tumid
where the side rounds into the back, but are most elevated in
the middle of the side. On the back the ribs are rather less
distinct, bend slightly towards the mouth, and each bears in its
centre a small sharp tubercle. In a younger state there are
also tubercles at the extreme edge of the back, which seem to
disappear with a diameter of twelve lmes. These are, moreover,
the only tubercles in specimens of five lines diam., the back being
till then smooth and rounding.
Mouth twice as high as wide, forming more than half the
diameter of the shell.
Septa complicated, divided on each side into four lobes. The
dorsal, which is wider and shorter than the superior lateral, is
or namented with two branches on each side; the lower of these
has three digits. The saddles are all half as wide again as the
lobes they correspond to, and divided by an accessory lobe into
two unequal parts. The superior lateral lobe, which is long and
narrow, has on each side two branches, of which the lower has
three digits, and in the middle a branch which bifurcates. In
the inferior lateral lobe the terminal branch does not bifurcate,
but terminates in three digits.
Height of shell 1? inch; height of umbilicus less than } inch.
Height of mouth 1 inch, width 4 inch.
It nearly resembles A. sevangulatus; but the much more
numerous and finer ribs, differently arranged, and the absence
of lateral tubercles from. the back, readily ‘distinguish it. The
A. Itierianus, D’Orb., has a very distant resemblance. Essen-
tially the shell is a compressed form of A. Mantelli, with a me-
sial row of dorsal tubercles instead of two lateral rows.
Loc. Cambridge. Coll. University Museum.
Ammonites sexangulatus. Pl. XI. fig. 1.
Few-whorled, discoidal, compressed, with an angular tuber-
culated back and small umbilicus.
Ornamented with about twenty-five wide, rounded, radiating
ribs, which are somewhat wavy, being for the most part bowed
234 Mr. H. Seeley on Ammonites
a little forward on the middle of the side: they are separated
by spaces never narrower than the width of the ribs. Less than
half of the ribs reach the umbilicus, two commonly uniting in
a fork at about a third the width of the whorl from it, aud a
single free one (which dies away at the same distance) sometimes
occurring between pairs of forks.
The sides, which are parallel, slope into the umbilicus, and
make a large angle with each half of the back, by tubercles
being developed on the ribs at the line where the back would
begin to round. The ribs are prolonged, somewhat widening
and curving forwards, to the centre of the back, where they ter-
minate in prominent tubercles. Thus the back is ornamented
with three rows of tubercles.
The mouth is six-sided, with the sides opposite and parallel.
It is half as high again as wide, and nearly half the height of
the shell.
The septa appear to consist of three lobes on each side. The
dorsal is wider and shorter than the superior lateral; they both
have three small branches on each side, and at the end two
larger ones trifurecate. The dorsal saddle, which is about as
wide as the lobe, is cut into by two very small accessory lobes.
In a young state (diameter # inch) the sides are perfectly
parallel ; only one or two of the ribs reach the umbilicus, and
all the others are much shorter than the short ones in the larger
specimen.
Height 13 inch; width of umbilicus } inch. Height of
mouth +2 inch, width 52. inch.
Loe. Cambridge. Coll. University Museum.
This form belongs to the small series with trituberculated
backs, typified by A. papalis. Iam not familiar with any form
which closely resembles it. 4. Itiertanus, D’Orb., has some
likeness to the young form; but the much more numerous ribs,
smaller umbilicus, &c., easily identify the shell described. A.
Brottianus is nearly related.
Ammonites acanthonotus. Pl. XI. fig. 5.
Few-whorled, compressed, with the sides gently inflated ;
back rounded, bearing a mesial row of spines; umbilicus as
high as the whorl at the opposite side of the mouth.
The umbilicus is shallow, with the lower third of the side,
which it includes, gently bevelled down to the preceding whorl ;
it is marked with radiating ribs, each terminating under the
succeeding whorl in a small tubercle.
On the side of a whorl there are about twelve tubercles, from
which its upper two-thirds inclines inwards, rounding gently on
nearing the back. From each eminence diverge two ribs (in
Jrom the Cambridge Greensand. 235
the young state, sometimes three); and there is a free rib not
descending quite so far as these, intermediate between each two
tubercles. The ribs, parted by spaces of about twice their width,
are rather small and obtuse, becoming relatively less elevated
and narrower with age. Ata third of the width of the side from
the back all the ribs are parted by regular distances, but from
about that point two of them converge towards a tubercle on
the back. The intermediate ribs, which do not pass over the
back, terminate where the back and side pass into each other.
The back is about half as wide as the umbilicus is high, and
supports on the last whorl a row of spines one-fourth more nu-
merous than the lateral tubercles. They are short and large,
having for the base the whole width of the back, are directed
forward, and get steadily higher. They are an adult character,
the ribs passing over the back till the specimen gets of more
than nine lines diameter.
The mouth, less than half the shell’s diameter, is about a
fourth higher than wide.
The septa are obliterated; they were simple, with a small
inferior lateral lobe, and below it three small accessory lobes.
A slight inflation extends all round one side of the whorl;
but, from the near resemblance the shell has to Ammonites
glossonotus, I am not inclined to give that weight to the distor-
tion it otherwise would have.
The late Dr. S. P. Woodward, in 1862, regarded this shell as
a monstrosity of Ammonites lautus, Sow.,—a view with which I
cannot agree.
Ammonites glossonotus. Pl. X. fig. 4.
Few-whorled, discoidal, moderately compressed ; back rounded;
umbilicus moderately large.
Around the umbilicus are (about ten) prominent tubercular
spines, from each of which commonly arise two ribs, and an-
other is generally placed in each of the hollow spaces inter-
mediate between the tubercles. These ribs are elevated, narrow,
and not nearly so wide as the spaces between them, which are
of about equal width. Two of the ribs ascend the side of the
shell, nearly parallel to each other, for two-thirds of its height,
when the hinder one bends rather suddenly forward so as to
unite, on the side of the back of the shell, with the front one,
which curves forward slightly ; united, they pass over the back
as a thick, elevated, tongue-like fold extending forward. Rarely
a rib passes over the back singly.
The mouth, about three-fourths as wide as high, is shaped
like an ass-shoe.
The septa are obscure, but appear to be unsymmetrical. The
dorsal lobe is short, nearly square, and has a branch on each
236 Mr. H. Seeley on Ammonites
side, which bifurcates. The dorsal saddle is unequally divided.
The superior lateral lobe has a few digits on each side, and two
terminal trilobed branches.
eth 12 inch; width of umbilicus } inch. Height of
mouth ? inch ; width nearly 2 inch.
This 1 is one of those remarkable Ammonites which undergo a
transformation of ornamentation. The characters described are
only those of the adult state. In a younger condition, the ribs
appear to have been alternately long and short, and to have
each passed over the back without a forward curve.
The example figured is the only one I have seen. There is
no cretaceous shell that can be compared with it; and one of
the few Ammonites having two ribs united to pass over the back
is a species from the Lias of Amberg, described by Miinster as
A. Fischert. That, however, has no umbilical tubercles; the
ribs are wide and obtuse, and fewer, and the aspect more
compressed.
Loc. Ashwell. Coll. University Museum.
Ammonites Woodward. Pl. XI. fig. 3
Few-whorled, inflated, with convex sides bearing spines, and
a round back. Umbilicus mostly higher than the whorl oppo-
site to the mouth, and appearing relatively high from the half-
embracing whorls enlarging but slowly : it is quite smooth and
rather deep ; but the sides are inflated, and round imperceptibly
into the sides of the whorl.
Around the side of the whorl, midway between the back and
the umbilicus, is a row of some ten or eleven rather elevated
spines, from which slight inflations descend to the umbilicus,
and strong round elevated ribs arise to pass over the back. In
each tubercle are collected three ribs, but only two of them pass
over to the corresponding tubercle; for the spines of one side
are placed a little between those of the opposite side, rather
than facing them, so making a zigzag, which does not, however,
strike the eye. The ribs, separated by spaces fully as wide, in
the more compressed forms, where the tubercles are a little be-
low the middle of the whorl, pass straight over the back; but
in inflated forms the coste are noticeably arched forward ;
and in this variety the tubercles are above the middle of the
side ; and consequently the back, which is very much broader,
is much less convex than in the less inflated form. There is no
area that can be named a side, the lateral spines dividing the
umbilicus from the back ; thus defined, the back will be nearly
as wide as three-fourths the height of the umbilicus.
The mouth is wider than high, shaped lke a moon entered
on her fourth quarter.
from the Cambridge Greensand. 237
The septa are simple, the dorsal lobe being square, with two
small terminal branches. There are two lateral lobes, one above
and one below the spines.
Diameter 1} inch, with septa to the end.
It also occurs in the Gault of Folkestone; and is not easily
distinguished from one described by Von Hauer, from the Lias,
as A. spinescens.
The name has for some years been associated with that of the
late Dr. S. P. Woodward, under whose friendly guidance it was
my privilege to gain a knowledge of shells.
Ammonites cvlonotus. Pl. X. figs. 2 & 8.
Few-whorled, much compressed, with nearly flat though
slightly inflated sides; back rounding, with a deep mesial
groove; umbilicus as high as the whorl at the opposite side to
the mouth.
The umbilicus is shallow, but well defined, its narrow hori-
zontal spiral boundary forming right angles with the vertical
sides. Around this umbilical angle are about twenty-five little
eminences—the thickened origin of the ribs. From these points
the ribs ascend towards the back, beg directed forward at a
considerable curve for about one-fourth: of their length. Each
rib then, on the side towards the mouth, gives off a branch,
and these bend back a little, so as to be for about half their
length perpendicular, and then again curve forward in a small
are, passing on to the back, where they continue to be directed
towards the mouth till terminating on the margin of the dorsal
groove at the distance of the fifth rib in front of their own
straight part. The ribs are wide, rounded, and depressed, and
separated by sulcations of about half their width, which taper
gradually both towards the back and umbilicus.
The back is half the width of the umbilicus, with a deep
mesial groove, towards which the sides gently round; the sides
of the sulcation make a sharp angle with the back.
The mouth at its base is two-thirds as wide as high; at its
upper part, where the sides begin to round into the back, it is
half as wide as high.
The septa of this shell are remarkable for the small size of the
dorsal lobe, which is contained in the dorsal groove, and bifur-
cates. There are two lateral lobes: the superior lateral is twice
as wide and half as long as the dorsal lobe; there are two
notches on each side of it, and at its termination three branches,
the central of which has three digits. The dorsal saddle, which
is more than twice as wide as the superior lateral lobe, is divided
into two subequal parts.
This is one of the less common forms; but the few specimens
238 Mr. H. Seeley on Ammonites
I have seen (perhaps twenty-five) show a wide amount of varia-
tion. The fossil figured (PI. X. fig. 2) is one of the most com-
pressed forms; and from it the umbilicus gets higher, the whorls
thicker, the ribs more numerous and less elevated, till at last,
to judge from fragments, the section of a whorl must have
been wider than high. In the form described the dorsal channel
is a third the width of the back, but in the widest form it is
only a ninth.
A variety oecurs in which the whorls are nearly half-embracing
(Pl. X. fig. 8), flattened on the sides, rounding on the back, and
step-like around the umbilicus, ornamented with about thirty-
two rather elevated wide ribs separated by suleations of about
equal width. The ribs are generally alternately long and short,
and terminate in fifteen umbilical tubercles. Aperture rather
higher than wide. Diameter 1} inch minimum.
MM. Pictet and Campiche, in their work on the fossils of Ste.
Croix, pl. 27. f. 2, have referred this type to A. falcatus of Mantell.
But at Cambridge no specimen of A. falcatus has ever occurred,
nor do the ribs vary in the least so as to approach that fossil
more than is seen in the specimen figured. The roundness of
the back and every feature of the ribs are matter for distinction ;
hence, and especially as the distribution is different, the forms
are separated. It is, no doubt, nearly related to A. falcatus,
having a channelled back and ribbed sides.
Diam. 2} inches; septa to the end.
Ammonites splendens, Sow.
A. splendens, Sow. M.C. t.103; Pictet, Grés Vert, pl.6. fig.6; D’Orb.pl.63.
A, Fittoni, D’ Arch.
A. auritus, D’Orb. T. C. vol. 1. pl. 65. figs. 3 & 4.
Shell compressed, with a small umbilicus, high, flattened
sides, and a very narrow flat back. The umbilicus, about as
high as the mouth is wide, and never more than a third the
height of the whorl, is shallow, with the horizontal ventrum,
which rounds into the side, not much deeper than the unem-
braced part of the whorl on which it abuts. The sides of the
whorls are very slightly inflated, and converge, so that the back
is only half as wide as the base. The mouth 1s less than half
the height of the shell. The sides of the cast are smooth,
or marked only with a few broad flexuous ribs scarcely elevated.
The dorsal angles are each crenated, with a row of minute tu-
bercles, which send slight thickenings a short way down the
sides.
Commouly in larger specimens the lower half of the side is
slightly inflated, so that the upper half looks more compressed :
the same peculiarity oceurs rarely in specimens from Folkestone.
from the Cambridge Greensand. 239
Septa commonly unsymmetrical, though the degree of in-
equality varies. The dorsal lobe is square, with two digitated
terminal branches, and two or three small notches on each side.
The dorsal saddle is half as wide again as the lobe, and mesially
cleft, though not deep. The inferior lateral lobe is almost as
large as the dorsal saddle: it terminates in three large branches,
all well digitated, the lateral ones bifurcating in full-grown
forms, but not into equal parts. There are three other lobes,
which are mere notches. The septa are very close together.
I have referred this fossil to the A. splendens of Sowerby
rather than to the A. Fittonz of D’Archiac, because it is quite
identical with typical specimens from Folkestone, though, were
Sowerby’s figure followed, no doubt it should be named A. Fi-
tont. But for the mineralization, it might have been supposed
that ours were southern Gault fossils, the only difference being
that, from the smaller size of the crenule, the back is commonly
a little convex instead of being slightly concave. Nothing ap-
pears to be gained by separating A. Fittoni from A. splendens ;
for it is not a well-marked variety, and our specimens are
slightly intermediate. It is a common fossil, and abundantly
represented in all collections, particularly those of the University
and Mr. Carter.
One variety, for which I am indebted to Mr. C.S. P. Darroch,
of Trinity College, has the septa at first slightly unsymmetrical,
and afterwards symmetrical. The shell is inflated, the mouth
being two-thirds as wide as high. The rather deep umbilicus
is bordered at the ventral angle with sixteen round tubercles.
The dorsal tubercles are larger than those at the umbilicus.
The back is round. The sutures are the same as in A. splendens,
except that there are three small lobes in the umbilicus instead
of two, while the lateral lobes are relatively only half as wide.
There are many variations of A. splendens, through which the
smooth forms pass into others having a sharp and elevated
flexuous rib descending from each small dorsal crenulation to
the base of the side, where two commonly unite to form a slight
thickening; between each two is a free rib, commonly not
descending so far. Occasionally two ribs unite in one dorsal
tubercle.
Some specimens reach as large a size as those from the Gault,
and must have had a diameter of 7 or 8 inches, but are only
found in fragments.
Passing on from these forms, the ribs gradually get less
sharp and wider apart, the umbilical thickenings more elon-
gated, and unite three ribs with intermediate free ones. Two
unite more commonly in each dorsal tubercle, which becomes a
trifle larger. The whole shell gradually thickens, the umbilicus
240 Mr. H. Seeley on Ammonites
enlarges, the ribs strengthen, the umbilical tubercles, as well as
those bordering the back, are more elevated; and thus A.
splendens varies into a new form, which it may be useful to dis-
tinguish as A. cratus.
Ammonites cratus. Pl. XI. fig. 2.
Form inflated, with half-embracing whorls, and a mouth nar-
rower than high, though wider than the umbilicus. The sides
are convex. The flat back is less than half as wide as the
mouth.
Around the umbilicus is a row of twelve large and elevated
spines, separated by spaces wider than their bases: they send
thickenings down the umbilicus, into which the most convex
part of the side rounds, abutting on the embraced whorl. They
also give rise to extremely elevated narrow ribs, separated by
wider, deep, concave channels, curving moderately mouthward. -
Three ribs are collected in each spine, and there is a free one
between each two bundles. About one half reach the back single,
and terminate each in a strong, elevated, tubercular thickening,
which extends obliquely forward into the middle of the back ;
the remainder unite in twos at the dorsal angle, and form similar
tubercles. These tubercles are so arranged as to give a slightly
dendritical aspect to the back.
Septa symmetrical, consisting of the dorsal Jobe and, on each
side, three (? or more) lobes. ‘The dorsal lobe is rather longer
than wide. The dorsal saddle, much wider than the lobe, is
centrally cleft. The superior lateral saddle is as wide as the
dorsal, has a single notch on each side, and terminates in three
large trifurcate branches. The other parts of the suture have
the same structure as those described, but get rapidly smaller.
As the forms depart from the original series, and the whorls
get more inflated, the septa become less and less unsymmetrical.
This extreme form is not common. The largest example in
the University Museum measures 34 inches high, and has septa
to the end.
Another branch of the series now passes on rapidly to A. au-
ritus, Sow., with which should be united A. Guersanti, Pict.
(not D’Orb.); for our specimens are almost identical with fig. 7,
pl. 5 of the ‘Grés Vert,’ differmg only in having rather fewer
tubercles on the back—a character which is the only one to
show that the figure is not copied from D’Orbigny’s A. auritus,
plo67. vol: i. “Terr.Cret?
Ammonites leptus. Pl. X. fig. 5.
Few-whorled, greatly compressed; sides nearly flat; back
from the Cambridge Greensand. 241
very narrow; umbilicus rather more than half the height of the
last whorl.
The umbilicus, which is angular, is bordered on the side
by about fifteen (somewhat elevated, but not very large) tubercles,
separated from each other by fully the width of their bases.
To each of these tubercles converge three or four ribs, two or
three of which generally die away on reaching the eminence.
The ribs are round, very gradually widen, and are separated by
sulcations about equally wide; for the lower two-thirds of their
length they are straight, and then gracefully curve forward,
dying away either separately or uniting in twos in large expanded
dorsal tubercles, which occupy the whole of the back, bend a
little outwards, and (probably) somewhat resembled those of A.
auritus ; they numbered about thirty on each side, and were
neither opposite nor regularly alternate.
The mouth is high and narrow, with the sides converging not
unlike an Egyptian doorway; it is more than twice as high as
wide.
Height 2+ inches; width of umbilicus =°, inch; height of last
whorl +2 inch ; width of base of mouth =, inch.
The specimen described is the only one I have seen. Though
the shell is otherwise well preserved, the dorsal tubercles are all
broken off quite at their bases. A line of the last whorl, broken
away, extends two-thirds round the shell; so that perfect speci-
mens were probably not less than 4 inches in diameter.
The affinities of this form are very near to A. splendens, Sow.,
very evident with A. serratus, Park, and not too distant to
recall the idea of A. auritus, Sow. The close ribbing, large dorsal
tubercles, and compressed aspect sufficiently and severally dis-
tinguish it from all of them.
Loc. Ashwell, University Museum.
Ammonites auritus, Sow.
a. A. auritus, Sow. M.C. vol. ii. pl. 134; D’Orb. T. C. vol.i. pl. 65,
A, Guersanti, Pictet (not D’Orb.), Grés Vert, pl. 5. f. 7.
8. A, Raulinianus, Pictet & Campiche, T. C.; Ste. Croix, pl. 29; D’Orb,
pl.,68. T. C.; Pictet, Grés Vert, pl. 7. fig. 2.
As has already been seen, A. Fittont passes into A. auritus ;
and similarly A. auritus passes into the fossil figured by Pictet
and Campiche as A. Raulinianus, which is only a variety of the
A, Raulinianus of D’Orbigny. The 4A. auritus figured by
Sowerby is a more robust form than that of D’Orbigny, more
intermediate in the series, and consequently a convenient type
for our forms.
The first form is a compressed shell, with the umbilicus half
as high as the mouth, which is nearly two-thirds as wide as
Ann. & Mag. N. Hist, Ser. 3. Vol, xvi. 17
242 Mr. H. Seeley on Ammonites
high. The sides are very slightly inflated, and converge so that
the back is only half as wide as the lower part of the mouth ;
they round into the umbilicus, bordering which are ten slightly
elevated tubercles, forming the boundary for the embracing
whorl. The flat back has on each dorsal angle a row of eighteen
or nineteen tubercles, which alternate, are larger than those of
the umbilicus, and are directed slightly upward, though scarcely
rising above the back. From the tubercles arise flexuous ribs
twice or three times as numerous as the dorsal tubercles, at which
they meet commonly in twos, sometimes in threes, with usually
a free rib between each two tubercles; they are similarly col-
lected at the umbilicus. The degree of elevation of the ribs,
which are sometimes indistinct, varies much, as does the degree
of flexure.
The septa are unsymmetrical, with a square dorsal lobe
having two branches on each side, and two terminal branches,
between which are a row of dorsal tubercles. The dorsal saddle,
wider than the lobe, is cleft mesially by a branch in a line with
the other row of dorsal tubercles. The superior lateral lobe is
longer than the dorsal, has on each side two lateral branches,
and terminates in three branches larger than the others. There
do not appear to be any accessory lobes.
From this, forms diverge having the septa variously sym-
metrical and unsymmetrical, in which the whorl is thicker, while
the umbilical tubercles are elevated into spines, and those of the
back become higher and wider. These are more typical forms of
A. auritus, about intermediate between the figures of Sowerby
and D’Orbigny.
Ammonites Raulinianus, var.
This is quite inseparable as a species from A. auritus.
Shell inflated, with a flat back tuberculate at the sides;
whorls half-embracing. Umbilicus nearly as high as the mouth,
and bordered with spines.
Mouth as wide as high, with the sides slightly converging to
the back; in their lower third they round into the umbilicus,
and on its margin support a row of nine or ten spines, generally
large, but varying. The back has about eighteen tubercular
spines, larger than those of the umbilicus, sometimes directed
upward, sometimes outward. The rows on the two sides are
alternate, so that the ornament of the back is zigzag. The ribs
are commonly strong and obtuse, and slightly curved forward.
Three always diverge from each umbilical spine, and two always
meet in each dorsal tubercle. Occasionally a dorsal tubercle
sends down a free rib,
Septa nearly symmetrical, with a square dorsal lobe, narrower
from the Cambridge Greensand. 243
than the back, with two. notches on each side, and terminating
in two small branches. The dorsal saddle is considerably wider.
The upper lateral lobe is long and narrow, terminating in three
small branches, and having one branch on each side. The lower
lobe is very small, and in a line with the umbilical tubercles.
It is unwillingly that this form has been described separately
from A. auritus, of which it is a badly marked variety, differing
chiefly in a different inflation of the shell. It has been classed
by the Swiss naturalists with A. Raulinianus; but in this series
names nowhere mark real boundaries or breaks.
The young shell so nearly resembles A. Studeri that I am
unable to discover any character not common to the two forms.
This circumstance in no way invalidates the conclusions arrived
at on the affinities of this shell; for all the forms, from A. Fit-
tont to the most extreme variation of A. Studeri, have round
backs in the young state; and if this shell is more inflated than
usual, that is because the adult is one of the most gibbous of the
series.
There are many variations from this form, in one of which
the ribs disappear; in another the dorsal tubercles gradually
become obsolete, while the back gets narrower and the sides
more convex, till at last the back as a flat region becomes obso-
lete, and the alternate ribs almost meet in alternate thickened
terminations along its middle. The ribs curve much toward the
mouth as they near the back. This small shell with a dendrous
back is probably immature.
Ammonites Salteri, Sharpe, Cret. Moll. pl. 28. figs. 5 & 3, is
a variation including those more compressed forms in which the
dorsal spines are reduced to tubercles and the umbilical spines
are small and the ribs slightly elevated.
In a variety which may be named A. ¢etragonus the shell has
flat sides and a flat back, and the mouth nearly quadrate. On
the lower third of the side is a row of ten or eleven small spines,
which are moderately elevated and separated by wide intervals.
At each dorsal angle is a row of about eighteen tubercular spines,
larger than the umbilical row, and, though short, directed late-
rally, which widens the back. These spines are connected by
ribs, which on the last half-whorl are very slightly elevated, and
ultimately become obliterated. Three appear to have always
diverged from each umbilical spinous eminence. The last half-
whorl is devoid of septa. Diameter 2 inches; width of mouth
4 inch.
Another shell (Pl. XI. fig. 6) of the same group scarcely differs
as a variety from A. Renauxianus, P. & C. Ter. Crét. Ste. Croix,
pl. 381. figs. 2-5 ; D’Orb. T. C. vol. i. pl. 27. Shell compressed,
with very few whorls, one-third-embracing, and enlarging very
LZ
244. Mr. H. Seeley on Ammonites
slowly. The flat sides, so slightly converging as to be almost
parallel, are half as high again as wide. The ventrum is hori-
zontal, and rounds into the side. The shallow and open
umbilicus shows that, up to the first, the whorls were smooth.
Here there are fourteen thickenings, which rapidly become
moderately elevated tubercles. A rib given off from each of
these bifurcates a little way up the side, but is very little
elevated. One or both of the branches reach the back, ter-
minating in a tubercle not larger than that at the umbi-
licus. In the last quarter- -whorl the ribs are curved towards
the mouth, and are obliterated in the upper part of the whorl.
The dorsal tubercles in the same space rapidly get smaller, be-
coming oblique thickenings ; they form a row of twenty-five.
The last:half-whorl is devoid of septa. These shells are as un-
symmetrical as any in the young state, but become finally nearly
symmetrical. The square dorsal lobe, more than half the width
of the back, has a notch on each side and two small terminal
branches. ‘The dorsal saddle is of the same width, and mesially
cleft. The upper lateral saddle is rather narrower, with a notch
on each side and three terminal branches. The other parts are
similar, but much smaller. There are two small accessory
lobes.
The dorsal tubercles are alternate, and the back slightly convex.
There are other specimens, one-fourth larger, with the mouth
perfect and as wide as high, with the shell inflated. The dorsal
angle is obtuse and rounded; the side rounds more noticeably
into the ventrum.
I believe these shells to be variations from A. auritus.
Ammonites Vraconensis, Pictet & Campiche, T. C. Ste. Croix,
plod. sie. A.
Inflated, few-whorled; whorls two-thirds-embracing, with
(diam. 2 inches) flat sides and a nearly flat back. The sides
converge, giving the mouth something of the outline of an in-
verted flower-pot. At one-third up the side is a row of eight or
nine large elevated spines, the space interior to which rounds
down to the whorl it embraces, and is smooth. From each
spine commonly arise (three or) four ribs, which extend up the
side to the margin of the back. At the angle where the side
and back meet is a row of tubercles about three times as nu-
merous as the umbilical spines, and in these the ribs terminate,
commonly two, sometimes but one, in each; so that there are
generally one or two free ribs between all the spines. Many of
the ribs are straight ; but the hinder one of the two, meeting in
a dorsal tubercle, necessarily has a bend in its upper third. The
dorsal tubercles of the two sides are not opposite, but alternate,
from the Cambridge Greensand. 245
and, compared with the umbilical spines, small. The umbilicus
is deep, and about as high as or higher than the whorl opposite
the mouth.
Ammonites Studeri, Pictet & Campiche.
A, Studeri, P. & C.; Ste. Croix, pl. 30.
Few-whorled, more than two-thirds-embracing, greatly in-
flated, with a flattened back, and umbilicus as high as the whorl
opposite the mouth, and bordered by a row of large spines.
The mouth is rather wider than high, and nearly twice as
wide as the back. The exposed part of the ventrum is inflated
and nearly horizontal ; but quite on its border, where the side
rounds into it, is placed the row of ten massive tubercles ; these
are much-elevated cones, with bases a fourth the height of the
side; above them the sides are flat, and converge to the back.
From each spine arise two or three ribs, which are slightly
curved mouthward, obtuse, and not much elevated till reaching
the margin of the back, where they terminate each in a thick-
ening which can scarcely be termed a tubercle, and which extends
a short way on to the back. Those of the two sides are alter-
nate, so that the slightly convex back presents a distant approach
to the kind of zigzag ornament which marks the back of A.
Raulinianus.
In the most typical specimens the septa are almost effaced ;
they appear nearly, if not quite, symmetrical. The dorsal saddle
is wide and unequally cleft by a small branch. The superior
lateral lobe is long, with two small notches on each side, and
terminates in three trifid branches. The inferior lobe is small,
and in the line of the tubercles.
Specimens identical with those figured in the ‘ Paléontologie
Suisse’ are rare; but small specimens having all the characters
of ornament the same, and differing only in being relatively
much less inflated, are by no means uncommon. But it is with
doubt that I have cast these in with A. Studert; for the young
state of A. Raulinianus is identical.
The adult shells of this type vary much in the degree of ele-
vation and number of the ribs, as well as the way in which they
are gathered in the tubercles.
I believe the facts given in this paper compel the mnion under
one specific type of every shell it describes after Ammonites colo-
notus. Throughout the series the variations in the septa are
insignificant. Every variety of shape is nothing but an inflated
form of Ammonites splendens modified by the different develop-
ment of crenule and ribs. The four chief species, Ammonites
splendens, A. auritus, A. Raulinianus, and A. Studeri, are in-
separably linked together by intermediate forms; while the
246 Mr. H. Seeley on Greensand Ammonites.
young of A. auritus is the species A. splendens, and small shells
like A. Studeri become with age the species A. Raulinianus.
Yet the other forms all have a value, though for convenience
they may be regarded as varieties of these types, which are but
subspecies of a larger group now named Ammonites permutatus.
Ammonites (Crioceras) occultus. Pl. X. fig. 1.
Moderately compressed, flattened, with few whorls, which
rapidly enlarge, and are so closely coiled that, while the whorls
do not appear to have actually touched, the tubercles of the
back have impressed themselves into the underside of the suc-
ceeding whorl. The transverse outline of the last whorl is four-
sided. The back is flat, and the base is a little concave. The
sides round into the base, and approximate each other with in-
creasing rapidity as they near the back, into which they also
gradually round. The back is half as wide as the base, and one-
third the height of the side. In the earlier whorls the back
appears to have been more round.
The shell is ornamented with a great number of moderately
elevated rounded ribs, which, below the middle of the side, are
slightly inflected forwards, as they are on nearing the back. At
the base of the side the ribs are collected in twos and threes,
forming elongated, elevated, obtuse tubercles; they ascend the
side at about equal distances apart, and so pass over the back ;
but, at the angles which the sides make with the back, every
third or fourth rib developes a large elevated tubercle, the base
of which is at least as wide as the space between the ribs: the
tuberculated ribs are often stouter than the others. On the basal
side, where the ribs are bent forwards, are two impressed lines
marking the width of the back of the preceding whorl; the
space between the lines is rather more than a third of the width
of the base.
The septa are indistinct. The dorsal lobe is twice as long as
wide, and extends over three ribs; it has two large, bifurcating,
many-digited, terminal branches, and two branches on each side,
the lower one being large. The dorsal saddle is as wide as half
the height of the side, divided by one large and many smaller
branches. The superior lateral lobe is about as large as that on
the back, but longer; it terminates in a trifid branch, the cen-
tral ramus of which has three digits. The inferior lateral lobe
is short; the basal lobe minute.
This remarkable Crioceras was obtained by the Rev. Dr. Cook-
son from near Hunstanton. But I suspect that both it and
the Trigonia tormerly named 7. Hunstantonensis have been ob-
tained from the Drift. It has been liberally presented to the
University Museum.
‘Mr. J. 8. Baly on new Genera and Species of Gallerucide. 247
For the drawings which illustrate this paper I am indebted
to the kindness and skill of the accomplished artist, Mr. Robert
Farren.
EXPLANATION OF PLATES X. & XI.
[All the figures are of the natural size. |
PLATE X.
Fig. 1. Crioceras occultus, Seeley : a, lateral view ; 6, ventral view, showing
how the dorsal spines indented the succeeding whorl ; ¢, a section
and septum; d, dorsal view. :
Fig. 2. Ammonites celonotus, Seeley.
Fig. 3. » var.
Fig. 4. glossonotus, Seeley.
Fig. 5. —— leptus, Seeley.
PuaTeE XI.
Fig. 1. Ammonites sexangulatus, Seeley.
Fig. 2. cratus, Seeley.
Fig. 3. —— Woodwardi, Seeley.
Fig. 4. pachys, Seeley. °
Fig. 5. acanthonotus, Seeley.
Fig. 6. Var. of A. Renauxianus, Pictet & Camp.
Fig. 7. A. rhamnonotus, Seeley.
XXVIII.—Descriptions of new Genera and Species of Gallerucide.
By J.S. Baty.
Fam. Gallerucide.
Subfam. HALTICIN”.
Genus SIM@THEA.
Corpus elongatum, parallelum, subeylindricum. Caput exsertum,
fere perpendiculare, pone oculos constrictum ; oculis orbitu circum-
datis, prominentibus, integris; facie inter antennarum insertiones
elevata; encarpis triangularibus, supra fossa transversa profunda
terminatis ; antennis corporis longitudini fere equalibus, filiformi-
bus, articulis cylindricis, primo breviter curvato, a basi ad apicem
paullo incrassato, secundo brevi, obconico, tertio ad primi longitudinem
eequali. Thorax transversus, basi vix transversim sulcatus, disco
convexus, lateribus anguste marginatis, rotundatis, angulis anticis
dente obtuso armatis. /ytra thorace paullo latiora, parallela, apice
subacute rotundata, supra convexa, regulariter punctato-striata,
Pedes robusti; coxis anticis non contiguis, suberectis ; femoribus
paullo, posticis magis incrassatis, his subtus non suleatis; ¢7bis
omnibus apice spina acuta armatis; farsorum posticorum articulo
primo duobus sequentibus paullo breviore; uwnguiculis appendicu-
latis. Prosternum distinctum sed angustissimum.
Type, Simethea Laportei, Baly.
Simethea must be placed in close proximity to Podagrica. In
248 Mr. J.S. Baly oa new Genera and Species of Gallerucide.
addition to its much greater size, it may be at once known by
the absence of the short perpendicular grooves present at the
base of the thorax in Podagrica ; it presents also an abundance of
other distinctive characters.
Simethea Laportei.
S. elongata, parallela, subcylindrica, nitida ; ore, pectore femoribusque
piceis ; tibiis tarsisque fusco-fulvis; antennis flavis, apice paullo
infuscatis ; thorace profunde sed remote punctato ; scutello rufo-
piceo ; elytris sat profunde punctato-striatis, rufis, apice nigris.
Long. 4 lin.
Hab. Tringanee.
Epistome triangular, the lower part of its surface plain, the
apical portion obliquely elevated ; face thickened and elevated
between the insertions of the antenne, but without forming the
usual facial ridge ; encarpz contiguous, large, subquadrate, the
lower and inner angle of each produced downwards ; bounding
the encarpz above is a deep transverse groove, the upper edge
of which is oblique, and gradually lost on the surface of the
front ; running upwards from this groove are a number of short,
nearly perpendicular grooved lines. Thorax rather broader
than long; sides narrowly margined, moderately rounded ;
upper surface convex. Scutellum trigonate, its apex obtuse.
Elytra scarcely broader than the thorax, parallel, each elytron
faintly impressed on the middle of the disk just below the
basilar space; deeply punctured, the punctures somewhat
remotely placed in ten or eleven longitudinal rows ; the black
apex varies greatly in extent, in some specimens being nearly
lost, in others occupying almost a third of the surface.
Genus XUTHEA.
Corpus elongato-ovatum, convexum. Capué exsertum ; facie tri-
angulari, carina lata, modice elevata; encarpis non contiguis; antennis
gracilibus, filiformibus, corporis longitudine brevioribus, articulo
primo paullo curvato, incrassato, secundo illo fere dimidio breviore,
a basi ad apicem modice incrassato, ceeteris gracilibus, simgulatim
primo eequalibus ; oculis orbitu cireumdatis, prominulis, integris.
Thorax transversus, lateribus anguste marginatis, fere parallelis,
angulis anticis tuberculo setifero armatis, dorso ante basin trans-
versim sulcatus, suleco utrinque fossa perpendiculari brevi ad basin
producta terminato. lytra regulariter punctato-striata. Pedes
modice robusti; cows anticis distantibus, prosterno fere zequialtis ;
JSemoribus posticis modice incrassatis, subtus leviter sulcatis ; ¢idiis
omnibus apice spina acuta armatis; ¢arsorum articulo basali ¢ am-
pliato, posticorum articulo basali duobus sequentibus conjunctim
Mr. J.S. Baly on new Genera and Species of Gallerucide. 249
fere aquali, tibiee apici inserto; wnguiculis appendiculatis, Pro-
sternum angustum.
Type, Xuthea orientalis.
This genus resembles Diplaulaca in the form of its thorax,
Crepidodera in the form and punctuation of its elytra; from the
latter it is separated by all the tibize having a short spine at
their apex, from the former by the entirely different shape and
punctuation of the elytra.
Xuthea orientalis.
X. elongato-ovata, convexa, supra viridi-cyanea, nitida, subtus ob-
scure viridi- aut piceo-znea; coxis, tibiis tarsisque piceis, plus
minusve neo tinctis; thorace fortiter subremote punctato ;
elytris nitidis ¢, minute granulosis, subnitidis 9, regulariter punc-
tato-striatis ; antennis fulvis, extrorsum piceis.
Long. 3-33 lin.
Hab. India.
Head exserted, face triangular; mouth pale piceous ; space
between the eyes rugose; carina moderately elevated, not very
broad ; encarpze ill feamed: remote ; face separated from the
front by a deep flexuose groove which runs obliquely upwards
on either side from the apex of the carina; vertex obsoletely
wrinkled in front, very remotely impressed with large punc-
tures; whole face clothed with coarse, depressed, sritiisln hairs.
Thorax one-half as broad as long; sides straight and parallel,
slightly converging and sinuate im front; surface nitidous,
strongly but subremotely punctured, all the angles slightly pro-
duced, acute, the anterior somewhat reflexed. Elytra smooth
and nitidous in the 3, subnitidous and very minutely granulose
in the 2, regularly punctate- -striate, interspaces plane ; ; basilar
space in each elytron bounded by a semicircular depression.
Body beneath clothed with coarse, adpressed, whitish hairs.
Basal joints of all the tarsi in the ¢ dilated, the middle elevated
into a longitudinal ridge.
Subfam. GALLERUCIN 2.
Genus Cynorra.
Corpus elongatum, angustum, parallelum. Caput exsertum;
facie subelongata, plana, subporrecta, inter antennarum insertiones
elevata ; mandibulis sat robustis, antrorsum productis ; antennis graci-
libus, filiformibus, corporis longitudini fere aequalibus, articulo primo
elongato, paullo curvato, a Bai ad apicem incrassato, secundo bre-
vissimo, obovato, czeteris fere eequalibus, singulis primo brevioribus,
quarto tertio paullo longiore; pa/pis maxillaribus apice ovatis ;
oculis prominentibus. Thorax transverso-quadratus, lateribus an-
guste marginatis, fere parallelis; disco modice convexo, profunde
250 Mr.J. 8. Baly on new Genera and Species of Gallerucide.
impresso. Llytra thorace paullo latiora, parallela, crebre punc-
tata, leviter elevato-vittata. Pedes elongati, graciles; cows anticis
contiguis, erectis ; femoribus vix incrassatis, subcompressis; ¢ibiis
omnibus singulatim apice spina acuta armatis; farsorum posticorum
sequentibus longitudine eequali; unguiculis appendiculatis. Pro-
sternum obsoletum.
Type, Cynorta porrecta.
The long, narrow form and produced head will serve to dis-
tinguish this well-marked genus at first sight from its allies.
Cynorta porrecta.
C. elongata, subfiliformis, viridi-zenea, nitida; capite (vertice ex-
cepto), thorace femoribusque fulvis ; thorace granuloso, arcuatim
bisuleato, violaceo-zeneo suffuso; femoribus dorso, tibiis, tarsis
antennisque (his basi exceptis) piceis; elytris granulosis, elevato-
vittatis, interspatiis confuse bifariam punctatis, interstitils inter se
reticulatis.
Long. 34 lin.
Hab. Java.
Face subporrect, vertex granulose, metallic green, apex of
jaws and palpi piceous. Thorax scarcely broader than long ;
sides slightly diverging from their base to beyond their middle,
then slightly converging to the apex; disk moderately convex,
somewhat flattened in the middle, impressed on the centre with
two curved fovez, contiguous at the base, diverging towards
their apices.
Genus NADRANA.
Corpus anguste ovatum, sat convexum. Caput modice exsertum,
perpendiculare ; antennis corporis longitudine, gracillimis, _ fili-
formibus, articulo primo curvato, a basi ad apicem leviter incrassato,
secundo brevi, tertio secundo fere duplo longiore, quarto tertio
dimidio longiore, czeteris articulo quarto singulatim fere aequalibus ;
oculis sat magnis, modice prominulis, integris ; palpis maxillaribus
apice ovatis, acutis. Thorax brevis, transversus, lateribus mar-
ginatis, angulis anticis obliquis, incrassatis; disco leviter transversim
suleato. Elytra ovata, convexa, infra basin non transversim im-
pressa, confuse punctata. Pedes graciles, sat elongati ; cowis anticis
erectis, contiguis ; Semoribus posticis non incrassatis ; ¢¢b7is omnibus
apice spina “acuta armatis ; ¢arsorwm posticorum articulo primo
elongato, duobus sequentibus conjunctim plus duplo longiore ; wn-
guiculis appendiculatis. Prosternum fere obsoletum ; metasternum
utrinque oblique depressum.
Type, Nadrana pallidicornis.
Jadrana is nearly allied to Luperodes ; it is to be distinguished
from it by the grooved thorax and more slender antenne, toge-
ther with the relatively longer third joint of the latter.
Mr. J.S. Baly on new Genera and Species of Gallerucide. 251
Nadrana pallidicornis.
N., elongato-ovata, valde convexa, nigra, nitida; vertice, tibiis quatuor
anticis apice tarsisque piceis; antennis pallide flavis; elytris
tenuiter suberebre punctatis, rufis.
cet
Long. 32 lin.
Hab. Tringanee.
Thorax three times as broad as long; sides rounded, their
outer margin narrowly reflexed ; above transversely convex, very
finely punctured ; middle of disk covered with a broad, shallow,
transverse excavation, which does not quite extend to the outer
border of the thorax. Apical border of elytra narrowly edged
with piceous.
Genus ANTIPHA.
Corpus ovatum, postice ampliatum, convexum. Caput exsertum,
subperpendiculare ; antennis gracillimis, filiformibus, corpore bre-
vioribus, articulis cylindricis, primo curvato, a basi ad apicem incras-
sato, secundo brevi, tertio primi longitudinis aut breviore, quarto
duobus preecedentibus eequali aut longiore, czeteris singulatim quarti
longitudini fere zequalibus, lis prope apicem paullo brevioribus ; oculis
magnis, prominentibus, integris. Thorax transversus, dorso non im-
pressus, lateribus fere rectis, parallelis, angulis anticis incrassatis.
Llytra thorace multo latiora, oblonga, postice paullo ampliata, con-
vexa, infra basin non transversim impressa, confuse punctata. Pedes
graciles, simplices ; cowis anticis suberectis, non contiguis ; femoribus
posticis non incrassatis; ¢/b¢is omnibus apice muticis; farsorum
posticorum articulo basali duobus sequentibus conjunctim longiore ;
unguiculis appendiculatis. Prosternum angustissimum, distinctum.
Type, Antipha picipes.
The glabrous upper surface, smooth thorax, long slender legs
and antennz, unarmed tibie, and appendiculated claws afford
sufficiently good characters (taken conjointly) for the foundation
of the present genus.
Antipha picipes.
LL. ovata, postice ampliata, valde convexa, nitida, fusco-fulva ; oculis
nigris ; epistomate, antennis (basi excepta), thorace infra, abdo-
minis limbo pedibusque piceis, thoracis disco fusco maculato.
Long. 43 lin.
Hab. India.
Face triangular: thorax nearly twice as broad as long; sides
straight, shghtly converging from base to apex, hinder angles
acute, anterior thickened, oblique; smooth on the disk, very finely
punctured on the sides: elytra more coarsely punctured, on the
outer disk is a longitudinal costa which runs parallel to the outer
margin for its middle two-fourths.
252 Mr. J.S. Baly on new Genera and Species of Gallerucide.
Genus Moma.
Corpus elongatum aut subelongatum, convexum. Caput crassum,
valde exsertum ; Jronte lata, declivi ; facie perpendiculari ; mandi-
bulis magnis; antennis corporis longitudini eequalibus aut paullo
brevioribus, filiformibus, ad apicem attenuatis, articulo primo cur-
vato, a basi ad apicem incrassato, secundo primi dimidiz parti eequali
aut paullo breviore, tertio elongato, articulo quarto longiore, ceeteris
singulatim quarto equalibus ; pa/pis maxillaribus crassis, apice acu-
miiinatis. ; ocults vix prominulis, integris. Thorax transversus, lateribus
anguste marginatis, obtuse anculatis, angulis omnibus tuberculo seti-
gero instructis ; disco transversim concavo, medio longitudinaliter ex-
cavato, utrinque transversim sulcato, L£lytra metallica, thorace
latiora, fere parallela, convexa, infra basin transversim depressa, con-
fuse punctata, glabra aut postice pube tenuissima sparse vestita.
Pedes modice robusti, sat elongati; cows anticis erectis, fere conti-
guis ; femoribus posticis non incrassatis ; ¢¢biis omnibus apice muticis ;
tarsorum posticorum articulo basali duobus sequentibus longiore ;
unguiculis bifidis. Prosternum lineariforme.
Type, Momea viridipennis.
The longer third joint of the antenne, together with the
different manner in which the thorax is excavated, will separate
this genus from Nicea and Humea, two nearly allied forms.
There is a slight error in the characters given by me of the latter
genus (Annals, January 1865, p. 37) ; “it ought to read— Pro-
sternum angustissimum aut obsoletum; ¢ibits coxis anticis con-
tiguis.” In both Nicea and Humea all the tibize have their apices
unarmed.
Momea viridipennis.
M. elongata, convexa, fusco-fulva, nitida ; eapite (epistomate excepto)
nigro-piceo ;_ pedibus antennisque nigris; elytris viridi-eneis,
crebre punctatis, sparse fusco-sericeis.
Long. 53 lin.
Hab. Mysol.
Head large; jaws robust, prominent; forehead deflexed, im-
pressed in the middle with a ‘deep longitudinal fovea; sur face shi-
ning, irregularly but not closely punctured : encarpe contiguous,
triangular ; ; lower portion of face, together with the base of the
jaws, obscure fusco-fulvous. Thorax more than twice as broad
as long; sides obtusely angled, diverging and slightly sinuate
from their base to just beyond the middle, then converging
to their apex ; upper surface transversely concave, impressed on
the hinder half of the middle disk with a large longitudinal
fovea; on either side is a deep but ill-defined transverse depres-
sion which extends from the outer border nearly to the medial
line. Elytra subelongate, nearly parallel, convex; their surface
convex, excavated and sinuous below the basilar space, about the
Mr.J.S. Baly on new Genera and Species of Gallerucidie. 253
middle of the disk, and on the sides; on the hinder disk are
several shallow, ill-defined, longitudinal sulcations.
Genus MIMASTRA.
Corpus elongatum, modice convexum, dorso subdepressum. Caput
exsertum ; facie subperpendiculare ; antennis corpore brevioribus,
gracilibus, filiformibus, articulo primo elongato curvato, a basi ad
apicem incrassato, secundo ad tertiam partem primi quali, tertio
secundo duplo longiore, quam articulum quartum paullo breviore,
ceeteris singulatim quarto fere eequalibus; pa/pis maxillaribus lan-
ceolato-ovatis, apice acuminatis. Thorax transversus, lateribus an-
guste marginatis, obsolete angulatis; disco irregulariter excavato.
Llytra thorace latiora, postice paullo dilatata, lateribus anguste ex-
planato-marginata, modice convexa, dorso deplanata, confuse punc-
tata. Pedes graciles, sat eclongati; covxis anticis erectis, contiguis ;
JSemoribus posticis non inerassatis ; ¢/b¢’s omnibus apice muticis; ¢ar-
sorum posticorum articulo basali duobus sequentibus conjunctim
eequali; wnguiculis bifidis. Prosternum obsoletum.
Type, Mimastra arcuata.
The less exserted and smaller head, the shorter third joint of
the antennee, the flattened upper surface, together with the
obsolete prosternum, separate the genus before us from Momea.
Mimastra arcuata.
M. elongata, subnitida, dorso subdepressa, subtus obscure olivacea ;
capite thoraceque fusco-fulvis, illo vertice, hoc disci maculis quinque
metallico-olivaceis ; antennis pedibusque piceis, femoribus infra
tibiisque apice fulvis; scutello nitido, nigro; elytris subelongatis,
postice leviter ampliatis, granulosis, obsolete rugulosis, ferniter
punctatis, metallico-olivaceis, singulatim limbo laterali fasciaque
arcuata prope medium posita, ad marginem adfixa, fulvis.
Long. 43 lin.
Hab. India.
Thorax one-half broader than long; sides distinctly margined,
nearly straight and parallel, moderately dilated, obtusely angled
just before their middle; upper surface covered with about six
large excavations; the green patches are arranged two in the
middle of the disk, and three in front of the basal margin, the
two lateral larger than the others.
Genus SASTRA.
Corpus elongatum, convexum, supra pube brevissima adpressa
plus minusve dense vestitum. Caput exsertum; facie perpendi-
culari, brevi, transversa; mandibulis mediocribus ; antennis corporis
longitudini fere sequalibus, gracilibus aut sat gr Acilibug, filiformibus ;
articulo primo curvato, a basi ad apicem incrassato, secundo brevi,
tertio duobus preecedentibus longitudine eequali aut paullo longiore,
quarto et sequentibus singulatim tertio brevioribus, inter se fere
eequalibus ; oculis magnis, prominentibus, integris ; palpis maxillari-
254 Mr.J.S. Baly on new Genera and Species of Gallerucide.
bus apice anguste ovatis, acuminatis. Thorax transversus, lateribus
medio szepe angulatis; disco subplano, medio longitudinaliter
suleato, utrinque transversim impresso ; angulis omnibus tuberculo
setigero armatis. L/ytra elongata aut subelongata, fere parallela,
postice vix ampliata, anguste explanato-marginata, convexa, dorso
subdeplanata, infra basin non aut vix transversim depressa, pube
sericea plus minusve dense vestita, confuse punctata, seepe disco ex-
teriore longitudinaliter sulcata. Pedes graciles, elongati; covis anticis
contiguis aut fere contiguis ; femoribus posticis non incrassatis ; ¢ébzis
omnibus apice muticis ; farsorum anticorum articulo basali duobus
sequentibus longiore; unguiculis bifidis. Prosternum angustissimum,
saepe fere obsoletum.
Type, Sastra placida.
The smaller head, much shorter and transverse face, together
with the much more pubescent upper surface of the body, will
sufficiently distinguish Sastra from Momea.
Sastra placida.
S. elongata, modice conyexa, sordide flava, nitida; abdomine anten-
nisque fuscis, his basi pallidioribus ; elytris dense punctatis, pube
adpressa fusca vestitis, pallide fusco-violaceis.
Long. 4 lin.
Hab. Mysol.
Head impressed with a longitudinal groove, which extends
from the anterior edge of the epistome to the vertex; encarpse
contiguous, large, pentagonal. Thorax nearly twice as broad as
long ; sides nearly parallel, slightly rounded in front, narrowed
posteriorly ; middle of disk impressed with a longitudinal sulca-
tion, which extends from base to apex, but is interrupted in its
middle; on either side is a broad transverse depression which
occupies nearly a third of the whole surface ; scattered distantly
over the thorax, but rather more crowded on the anterior mar-
gin, are some large deep punctures. Elytra subparallel, lateral
border narrowly dilated, flattened along the suture, closely punc-
tured; densely clothed with adpressed fuscous pubescence ; on
the outer disk are two shallow longitudinal sulcations, separated
from each other by an elevated ridge.
Sastra limbata.
S. elongata, pallide flava, nitida; oculis nigris; elytris metallico-
violaceis, singulatim pallide flavo limbatis.
Long. 35 lin.
Hab. New Guinea.
Face short, transverse; space between the insertions of the
antennee broad, impressed with a longitudinal groove, which runs
downwards across the epistome and upwards to the vertex; ca-
rine large, contiguous, subpyriform; antenne four-fifths the
length of the body, slender; apex of jaws piceous, Thorax
Dr. E. von Martens on Australian Species of Paludina. 255
nearly twice as broad as long; sides subparallel, slightly rounded,
narrowed and sinuate behind their middle; surface smooth, im-
punctate, the longitudinal sulcus interrupted in the middle, less
deeply impressed than the lateral fovez, which are broader and
deeper, but ill defined. Elytra not very closely punctured,
nearly glabrous in the single specimen before me (which is, in all
probability, worn) ; the entire limb, with the exception of a small
space near the scutellum, narrowly edged with flavous ; basilar
space bounded beneath by a shallow depression ; running along
the outer disk, and bounded exteriorly by an indistinct ridge, is a
broad, shallow, longitudinal groove ; on the hinder disk near the
middle are also to be seen the traces of a second, very ill defined.
XXIX.—On the Australian Species of Paludina.
By KE. von Martens, M.D., C.M.Z.8.
Onty one Australian species is mentioned in the list of the
species of this genus given by Frauenfeld in the ‘ Verhandlungen
des zoologisch-botanischen Vereins in Wien,’ 1862, as well as in
Reeve’s ‘ Conchologia Iconica.’ Having enjoyed the advantage
of examining some others in the British Museum and im the
Zoological Museum of Berlin, I shall here give comparative de-
scriptions of them.
1. Paludina australis, Reeve, Conchol. Icon. 1863, no. et
fired:
Probably P. essingtonensis, Shuttleworth, Frauenfeld, 7. ce.
p. 1162.
P. testa conico-globosa, perforata, tenui, confertim spiraliter undu-
lato-striata, virescenti-cornea, fasciis rufo-fuscis 3-5 picta ; spira
gradata; anfr. 5-6 inflati, sutura profunda distincti; apertura
subperpendicularis, circulari-ovata, angulo supero modice acuto ;
peristoma interruptum, album.
Altitudo 38, diameter major 31, minor 25, aperturee altitudo 21,
latitudo 17 mill.
Operculum normale.
Australia; collected by Mr. Gilbert at Port Essington. (B.M.)
Similar in size and form to the European P. vivipara, Mill.,
Lam. (P. Listeri, Forbes), but readily distinguished by its sculp-
ture being similar to that of some species of the Indian Archi-
pelago. The three principal bands occupy the same place as
those of the European species, or as the principal ridges in the
Indo-Chinese (P. angularis, Mill., and P. costata, Q. & G.) ; but
in several specimens there are two additional bands, narrower
and paler, the one above, the other beneath the uppermost of
the three principal ones.
256 Rev. H. Clark on Dejean’s Genus Coelomera.
Paludina affinis.
P. testa conico-ovata, suboblate perforata, solidula, lineis spiralibus
impressis, subtilissimis vel obsoletis, fusco-cornea, fasciis rufo-
fuscis 3-6 picta; spira convexo-conoidea ; anfr. 5, convexi, sutura
mediocriter profunda divisi; apertura paulum obliqua, subcireu-
laris, angulo supero rotundato; peristoma subcontinuum, iterate
nigro limbatum.
Alt. 27, diam. major 23, min. 183, apert. alt. 163, latit. 13} mill.
Operculum normale.
Australia; collected at Fitzroy river and near Port Essington
by Capt. Wickham, R.N.; other specimens by J. R. Elsey, Esq.
(B.M.)
This species stands nearly in the same relation to the pre-
ceding as P. fasciata, Mill. (P. achatina, Drap., Lam.), to P.vivi-
para, Miull., the chief difference being in the outlines of the
whorls ; besides, the spiral sculpture is much less developed in
P. affinis. ‘The bands are almost the same as in P. australis;
but there is a third, secondary band between the second and
third principal ones.
This species varies somewhat in the elevation of the spire,
several smaller specimens found at the same localities having
it more produced, and therefore being of a more oblong form,
and having a relatively smaller aperture. The dimensions of
one of these are as follows :—altit. 194, diam. maj. 15, min, 12,
-apert. alt. 114, lat. 9 mill.
3. Paludina polita.
P. testa globoso-conica, perforata, nitida, lineis spiralibus impressis
subtilissimis haud valde confertis sculpta, corneo-lutea vel pallide
carnea, fasciis nullis; spira conoidea, subgradata; anfractus 5,
inflati, valde convexi, sutura mediocri; apertura paulum obliqua,
ovata, angulo supero modice acuto; peristoma plerumque con-
pum, iterate fusco limbatum.
Alt. 223, diam. maj. 19, min. 17, apert. alt. 13, lat. 11 mill.
Operculum normale, rufum.
South Australia, on the Balonne river, New South Wales ;
found by John Macgillivray. Other regularly decollated speci-
mens, of a brighter red colour, in Lake Alexandria, found by
Mr. Strange. (B.M.)
XXX.—An Examination of the Dejeanian Genus Coelomera (Co-
leoptera Phytophaga) and its Affinities. By the Rev. Hamuirr
Crakk, M.A., F.L.S.
Tue genus Calomera of Dejean’s Catalogue (3rd edition, 1837)
Rev. H. Clark on Dejean’s Genus Coelomera. 257
and of cabinets consists of a variety of forms. As at present
constituted, it represents simply one of several forms of Galleru-
cide, which can neither be referred, on the one hand, to any of
the genera shadowed forth by Adorium, nor, on the other, to
the group of insects more immediately related to Adimonia and
Galleruca proper. In Dejean’s Catalogue it is made to compre-
hend six or seven distinct real genera, the species of which are
found in North America, South America, Africa, and Asia;
while in modern cabinets the limits of the group would appear
to be, if possible, more undefined still; so that the name has be-
come a sort of refuge for everything in the neighbourhood which
is unknown or which has been uncharacterized. An examination
of my material, during my summer holiday, has enabled me to
fix with some certainty the limits of the several genera which it
has comprised. I need not here analyze the Dejeanian species«
they will be found in the following pages in their natural places.
I have been able to trace them all. It may be convenient, how-
ever, to prefix a brief synoptical table of the several genera which
contain the species, and which are dealt with in this paper.
List of Genera.
Antenne robust, short, incrassated; body ovate ... I. CerocHroa.
Antenne incrassated, cylindrical, joints 3, 4, and 5
being equal; body subcylindrical, subovate ...... If. ALeuipia.
Antenne incrassated, joint 3 being longer than 4,
joints 5-11 broadly compressed; body ovate...... III. Cuirena.
Antenne incrassated, joints 1, 3, 4 being subequal,
and 9 and 2 minute and equal; body subparallel IV. HyMrENzst1a.
Antenne incrassated, serrated, joints 3-7 being the
broadest, joints 3-7 equal in length; body ovate V. OrTHOXIA,
Antenne incrassated in g, joints 4~7 dilated and
compressed, 3rd joit shorter than 4th; body
Du MIG) ao bade bapbbadbaer con saricuoL npadcapaancnencslgsoaonerdc VI. Pyesia.
Antenne incrassated in 3, very long, joints gradu-
ally diminishing in thickness from 1 to I1......... VII. Procauus.
Antenne robust, filiform, joints 3, 4, and 5 being
subequal; body robust, subparallel...............0.. VIII. PacuyToma.
Antenne filiform, robust, jomts 1 and 3 equal, and
4, 5, 6 equal and somewhat shorter; body short,
jsiatenl | ll Beeanaiconcoquésoscuncne mcpeecococnic rocouoneceDLtc IX. SPHENORAIA.
Antenne filiform, joints 4 and 5 equal and shorter
than 1 and 3; body parallel; thorax much con-
RURICLCOUAL SERCO IDASE) tcanccsetees sens cance cecnsecredeasir X. DirceMa.
Antenne filiform, moderate in length, joints 4 and 5
subequal; body robust or broadly ovate ......... XI. Monocesta.
Antenne filiform, moderate in length, third joint
very long; body generally broadly ovate ......... XII. Ca:LoMERA.
Antenne filiform, robust, nearly as long as the body,
joints 3 and 5-10 nearly equal; body parallel ... XIII. Corata.
Antenne filiform, slender, nearly as long as the
body; body subparallel...........ecessessscanee sirepare) SL Vig NESTINUS:
Ann. & Mag. N, Hist, Ser.3. Vol, xvi. 18
258 Rey. H. Clark on Dejean’s Genus Ceelomera.
Genus I. Cerocuroa, Gerstiick. Peters, Mossamb. 1862,
Zool. Ins. 341; Bericht Akad. Berl. 1855, p. 3.
Ovata. Caput verticale; palpi mazillares subcylindrici, apice
acuminato. Thoraw elytris multo attenuatior, levis, latere anteriore
paulum emarginato, angulis anticis satis porrectis et acutis, lateribus
paulum rotundatis, haud marginatis, margine postico subtransverso ;
disco levi punctato. Scutellum subtriangulare, apice rotundato.
Elytra thorace latiora, versus medium ampliata, leevia, vix marginata,
crebre punctata. Pedes robusti, unguiculis fortiter appendiculatis.
Metasternum versus apicem vel obsolete, vel in dentem acutum pro-
ductum. Antenne robustee, breves, incrassate, articulis 4°-11™ bre-
vibus et subzequalibus, 1° longiore, 3° primo breviore.
The species on which this genus is based (Cerochroa ruficeps,
Gerst.) is more nearly allied to Adorium than to Celomera. The
genus may be separated from all other neighbouring forms by
its short and robust antennee ; it 1s also conspicuous for the very
narrow breadth of the thorax. From Adorium it may be sepa-
rated by the form of its antennz ; from Rhombopala, by its more
elongate form; from Calomera, by its smooth and even thorax.
Three species of this genus are known :—
1. C. ruficeps, Gerst. Bericht Verh. Akad. Berl. 1855 ;
Peters, Reise Mossamb. Zool. 341.
2. C. brachialis, Suffr. Efver. Vetens. Ak. Forh. 1858. Caffraria.
I think it probable that this species may prove to be a variety
of C. ruficeps.
3. C. maculicollis, Baly, Descript. of Uncharacterized, &c.,
Trans. Ent. Soc. Lond. 1864, p. 282. Old Calabar.
Genus IJ. Aupuipra.
Robusta, subparallela. Caput verticale. Palpi mavzillares art.
penultimo globoso. Thorax declivis, transversus, angulis anticis
depressis, posticis obsoletis; disco levi. Scutellum triangulare,
leve. Elytra robusta, leevia, punctata. Antenne robuste, incras-
satee, art. 4°-11™ gradatim incrassatis, art. 3°, 4° et 5° longitudine
eequalibus. Pedes robusti, unguiculis simplicibus.
The genus A/phidia will stand near to Clitena ; it is separated
from it by its cylindrical (not flattened) incrassation of. the an-
tenne, by the 3rd, 4th, and Sth joints being equal in length,
and by the simple unguiculi.
A. comitata, (Galleruca) Klug, Ins. von Madagascar, p. 124.
A common species, apparently, in Madagascar. In colour
flavous, with elytra bright green or bluish green. Length 43,
breadth 24 lines.
Rey. H. Clark on Dejean’s Genus Coelomera. 259
Genus III. Crrrena, Baly, Descript. of Uncharacterized, &c.,
Trans. Ent. Soc. Lond. 1864, p. 229.
Callopistria, Chev. De}. Cat. p. 402.
Robusta, subparallela. Caput fortiter punctatum. Thorazv trans-
versus, angulis anticis prominulis, margine anteriore circulariter
emarginato, lateribus subrotundatis et marginatis, basi subcirculari
et leviter marginato; disco leevi (vix ut in Ca/lomera transverse de-
presso) punctato. Sceutellum subquadratum, apice circulari. Elytra
thorace latiora, parallela, apice rotundata, ante medium transverse
depressa vel constricta, leevia, punctata, nitida. -dntenne robuste,
satis elongatze, art. 1°, 3° et 4° subzequalibus, art. 2° brevi, art. 5°-11™
latis compressis, art. 6°—11™ brevibus. Pedes robusti, unguiculis bi-
hamatis.
Clitena differs from Celomera and others in the form of its
elytra, which are more robust and antemedially depressed, in the
smooth form of the thorax, which shows no trace of the usual
deep transverse depression, and especially in the peculiar arti-
culation of the antennz, the apical joints of which are flattened,
broad, and short. Two species have been described by Mr. Baly
—C. limbata and C. melancholica, both from Siam.
C. cyanea.
C. Indica, Dej. Cat.
C. robusta, subparallela, cyanea, pedibus et antennis nigris; caput
fortiter punctatum, nigro-cyaneum; thorax leviter punctatus;
scutellum nigrum ; elytra robusta, ante apicem transverse depressa,
leevia, punctata; corpus subtus, antenne, et pedes vel nigra vel
nigro-cyanea.
Long. corp. lin. 5; lat. lin. 23.
Java.
Genus IV. HyMeEnesta.
Parallela, subdepressa. Caput verticale. Palpi mavzillares art.
penultimo triangulari, robusto. Thorax transversus, angulis anticis
brevibus, posticis rotundatis, disco ineequali. Scutellum triangulare,
leve. lytra parallela, confuse punctata. dntenne robuste, arti-
culis presertim apicalibus ineequalibus (subcylindricis, vix depressis),
art. 1°, 3° et 4° subeequalibus, 5°, 6°, 7° et 8° brevioribus et robustiori-
bus, 9° et 2° brevibus, minutis zequalibus 10° et 11° conjunctis. Pedes
robusti, tarsorum art. basali et 2° penitus eequalibus, unguiculis bipec-
tinatis.
I have formed this genus on the species H. Tranquebarica,
Fab.: it is quite distinct, by reason of the very peculiar articu-
lation of the antennee.
H. Tranquebarica, Fab. Syst. El. i. p.479. 8, Suppl. p.93. 6,7,
is punctate and rufous, the antenne, legs, and underside
being black; the elytra are thickly punctate and rufous, the
18*
260 Rev. H. Clark on Dejean’s Genus Coelomera.
apex being marked by a purple transverse band, which broadly
extends, near the margination, for nearly half the elytra. The
length is 44, breadth 24 lines. From the East Indies.
Genus V. OrrHoxta.
Subparallela et satis depressa. Caput verticale. Palpi maxillares
articulis apicali et penultimo globosis. T'horaw transversus, margine
antico transverso, lateribus subrotundatis, margine postico sinuato,
angulis anticis vix prominentibus, disco in typo ineequali. Seutedlum
triangulare. Elytra thorace paulum latiora, parallela. Antenne
robust, serratse, articulis (presertim 3°-7™) apud apices incrassatis,
art. 3°-7™ longitudine subeequalibus. Pedes robusti; unguiculis ad
apices extremos, bihamatis.
This genus is nearly related to Monocesta, but differs from it
in the globularly inflated maxillary palpi, in the minute bifurca-
tion of the unguiculi (those of Monocesta being cleft almost to
the base), and the serrated articulation of the antennee.
O. Boisduvalli, De}.
O. subparallela, satis robusta, crebre rugosa, rufa, elytris, antennis,
capite et pedibus nigris; caput rugosum, nigrum; thorax trans-
versus, angulis anticis subobsoletis, marginibus leviter rotun-
datis, versus apicem obsolete angulatis, basi sinuata, disco
depressione media basali, alteraque utrinque inzequali obliqua
versus latera; thorax crebre punctatus, rufus; scutel/wm rufum ;
elytra thorace paulum latiora, parallela, robusta, creberrime punc-
tata, obsolete tomentosa, nigra; antenne robustee, articulis 3°-6™
sensim ampliatis, nigree : corpus subtus rufum ; pedes nigri.
Long. corp. lin. 4 ; lat. lin. 2.
Java.
Genus VI, Pyrsta.
Differt a genere Monocesta in antennis art. 3° primo breviore, tenui,
art. 4° tertio multum longiore, art. 5° et 6° subsequalibus et tertio
paulum longioribus; antennis in @ simplicibus, filiformibus; in ¢
art. 4°-7™ dilatatis, compressis.
P. laticornis, Germ. Insect. Spec. 589, 843.
Closely allied in pattern to M. consularis, Dej., but a much
smaller insect, and more depressed in form; the elytra are not
finely rugose, but levigate and punctate. The articulation also
of the antennze, both in 9 and g (which is the special character
of the separate genus into which it must be erected), abun-
dantly distinguishes it from this insect.
Brazil.
Genus VII. Procauvs.
Robustus, parallelus, brevis. Caput verticale. Palpi mavillares
elongatuli. Yhorax magnus, transversus, vix elytra amplitudine
Rey. H. Clark on Dejean’s Genus Coelomera. 261
gequans, margine anteriore valde excavato, angulis anticis rectis,
lateribus subrotundatis, marginatis, angulis posticis obsoletis, basi
transversa, disco levi. Scutellum triangulare, leve. Hlytra parallela,
brevia, robusta, thorace paulum latiora, punctata (punctis confusis,
inzequalibus) ; antenne vel filiformes vel incrassatee ; in ¢ penitus
corpus ipsum longitudine eequantes, et ad basin valde robustze, com-
pressze, sensim versus apicem attenuantur; art. 1° valde ad apicem
ampliato, 2° et 3° minoribus, brevioribus, et subzequalibus, 4°, 5° et
usque ad 11™ longitudine subzequalibus, latitudine sensim attenuatiori-
bus; in 2 antenne filiformes ; pedes robusti, satis breves, femoribus
subincrassatis, tibiis versus apicem robustioribus et ad insertionem
tarsorum subtus excavatis; unguiculi ab infra breviter utrinque bi-
dentati.
This diagnosis is based on a Chilian species, Celomera mutans
of cabinets, which is very different from all allied groups, not
only by reason of its shorter and more robust form, but also by
the largely developed basal joints of the antennz in the male.
P. mutans.
P. et colore et maculis incertus, flavo-viridis vel fuscus, thorace et
elytris, vel neutro vel utroque, nigro maculatis ; caput longitudi-
naliter oblique foveolatum, rufo-flavum ; ¢orax rarius punctatus,
vel flavus vel nigro maculatus, maculis quatuor (duabus mediis
et una utrinque postmedia laterali) ; scutellum impunctatum ;
elytra punctis subordinatis minutis, plerumque flava, sed ali-
quando maculis nigris juxta basin transverse ordinatis, maculisque
etiam juxta apicem ; antenne nigre, flavo annulate ; corpus subtus
in $ abdominis apice fortiter ad medium foveolato, fuscescens,
abdomine aliquando rufo ; pedes rufi, tibiis et tarsis nigris.
Long. corp. lin. 3-4; lat. lin. 13-2.
The head-quarters of this species is Chili, where it would
appear to be abundant; it is widely distributed. I have re-
ceived specimens from Brazil and also from Bolivia.
Genus VIII. Pacuyroma.
Robusta, brevis, parallela. Caput breve, verticale. Palpi mazil-
lares graciles, elongatuli. Thorax declivis, elytris paulum at-
tenuatior, angulis anticis prominulis, lateribus paulum curvatis, mar-
gine basali subsinuato ; disco levi, punctato. Scutellum triangulare,
apice truncato. Llytra brevia, robusta, confuse punctata. Antenne
filiformes, robust, art. 1° longissimo, 2° brevi, 3°, 4° et 5° subsequali-
bus. Pedes robusti, unguiculis pectinatis.
1. P. Westermanni, De}. Cat.
P. e majoribus, fusca, antennis, pedibus et corpore subtus nigris ;
caput longitudinaliter foveolatum, punctatum ; ¢horaa crebre et
fortiter punctatus ; scutellum nigrum; e/ytra crebre punctata ;
262 Rey. H. Clark on Dejean’s Genus Coelomera.
corpus subtus et pedes nigra; antenne nigri, articulis 1°-3™ flaves-
centibus.
Long. corp. lin. 6; lat. lin. 33.
Western Africa.
2. P. flava.
P. subovalis, lata, punctata, flava; caput in medio leviter lon-
gitudinaliter foveolatum, punctatum, flavum, basi transverse nigra;
thorax transversus, lateribus et angulis posticis rotundatis, lateri-
bus marginatis, disco erebre punctato; scutellum triangulare,
nigrum ; elytra post medium paulum ampliata, confuse et minute
punctata vel rugosa; antenne nigro-fuscze ; pedes et corpus sub-
fus nigra.
Long. corp. lin. 5, lat. lin. 3.
Natal.
Genus IX. SpHENORATA.
Lata, subdepressa, brevis, parallela. Caput verticale. Palpi
maxillares art. penultimo incrassato, brevi. Thorax transversus,
lateribus marginatis et rotundatis, angulis anticis prominulis, basi
subsinuato. Scwtellum triangulare, leve. Llytra parallela, brevia,
striato-punctata (punctis interdum penitus confusis). Antenne fili-
formes, robustee, art. 1° et 3° eequalibus, et art. 4°, 5° et 6° (articulo
primo paulum minoribus) inter se zequalibus. Pedes tenues, ungui-
culis simplicibus vel leviter appendiculatis.
1. S. flavicollis.
S. rufo-flava, elytris (marginibus et sutura exceptis) nigris; caput
transverse foveolatum, impunctatum ; thorax rufo-flavus, impunc-
tatus (punctis sparsis in medio disco obsoletis) ; seutellum ruto-
flavum, impunctatum; e/ytra parallela, punctata (punctis nec
magnis nec acie instructis, confuse ordinatis), fusco-nigra, sutura et
marginibus tenuiter flavis ; pedes, antenne et corpus subtus flava.
Long. corp. lin. 32; lat. lin. 2.
Northern India.
2. S. nigripennis.
S. flava, elytris, thorace maculis et femoribus apicalibus nigris ;
caput inter oculos transverse foveolatum, leeve, flavum basi nigra ;
thorax flavus, macula utrinque nigra insulata, fortiter sed sparsim
punctata et paulum depressa; scutellum leve, mgrum; elytra
striato-punctata (punctis magnis et profundis), nigra; antenne
nigro-fuscze, art. 1°—4™ flavescentibus; corpus subtus et pedes fusco-
flavescentia.
Long. corp. lin. 4; lat. lin. 23.
Northern India.
Genus X. Dircema.
Parallelum, satis depressum. Caput verticale. Palpi mazillares
elongatuli. Thorax transversus, margine anteriore paulum emar-
ginato, angulis anticis porrectis, lateribus oblique arcuatis, angulis
Rey. H. Clark on Dejean’s Genus Coelomera, 263
posticis obsoletis, disco valde transverse depresso. /ytra parallela,
depressa, subtiliter rugosa. Antenne filiformes, art. 1° et 3° sub-
eequalibus, 2° minuto, 4° et 5° subzequalibus et tertio paulum brevi-
oribus. Pedes graciles, unguiculis utrinque bifidis.
The form of the thorax in this group is peculiar; it is nar-
rowly transverse, the anterior angles are laterally producedcon-
siderably beyond the head ; the sides converge gradually towards
the base, being arcuate or constricted medially ; the basal angles
are slightly prominent ; and the disk of the thorax is distinctly
and deeply transversely depressed. I know of three exponents
of this form, all common species in the tropics of South America.
1. D. nigripenne, Fab. Ent. Syst. 1.11. 14. 9; Syst. El. i. 480.
Notable by its uniformly opaque-black elytra (rugose and to-
mentose) and its clearly coloured rufous or flavous thorax.
Long. corp. lin, 43-53; lat. lin. 2-23.
The species is very common in Cayenne : some examples have
the apical joints of the black antenne testaceous ; and one ex-
ample in my cabinet has the head testaceous instead of black.
2. D. cinctipenne.
D. oblongum, parallelum, opacum, reticulosum, tomentosum, fusco-
flavum, elytris (sutura et marginibus exceptis) et capite nigris vel
viridi-nigris ; caput inter peules longitudinaliter fov Sigh crebre
punctatum, nigrum ; thorax fortitee transverse depressus, punc-
tatus, plus anak tomentosus, flavus, macula utrinque nigra magna
aliquando insulata; scwtel/um flavum ; elytra parallela, nigra vel
nigro-viridia, sutura et marginibus tenuiter flavis ; pedes nigri,
fompribas plus minus flavis ; corpus subtus flavo-fuscum ; antenne
nigree, art. 9°-11™ flavis.
Long. corp. lin. 33-7; lat. lin. 13-
D. cinctipenne varies not only in size, but in the sculpture
and coloration of the thorax, which is either entirely flavous or
flavous with two submedial black markings, one on either side,
these markings sometimes occupying nearly the whole disk.
The species has been found at Para, by Mr. Bates and others.
I have two Columbian representatives of it, which have the ely-
tra opaque green instead of black, and the thorax of which is not
so deeply depressed. I have seen some interesting examples of
this species in Mr. Baly’s cabinet, which, at first sight we are
tempted to declare, represent distinct species: I have little
doubt, however, on more mature examination, that the above
diagnosis 1s the true definition of the species. The insect is as
variable as it is beautiful; and at present, at any rate, we have
not sufficient material to justify us in breaking it up into dis-
tinct races or species.
2
a
2s
264 Rey. H. Clark on Dejean’s Genus Coelomera.
3. D. ruficrus, Chev.
D. parallelum, opacum, tomentosum, rufo-flavum, elytris, antennis
et genibus nigris; caput leviter rugosum, impunctatum, flavum ;
thorax subpubescens, rufo-flavus ; scutellum fuscum ; elytra pa-
rallela, tomentosa, nigra; antenne nigre; corpus subtus rufo-
fuscum ; pedes fusci, femoribus rufo-fuscis.
Long. corp. lin. 433; lat. lin. 14.
Cayenne.
Genus XI. Monocesta.
E majoribus, robusta, plerumque versus apicem dilatata. Caput
verticale, basi longitudinaliter foveolatum. Thorax transversus,
margine anteriore paulum emarginato; angulis anticis sat productis,
lateribus subrotundatis; angulis posticis omnino vel penitus obso-
letis ; discus transverse et fortiter depressus est. Scutel/um trans-
versum, apice rotundatum. /ytra robusta, thorace latiora, post
medium plus minus dilatata, aliquando versus apicem dehiscentia, et
utrinque angulata, marginata, punctata. Antenne vel filiformes
robustee vel subincrassatee, art. 1° apice incrassato, art. 1°, 3° et 4°
subeequalibus, art. 2° minore, art. 5° et 6° subzequalibus, paulum
quarto brevioribus, art. 7°-11™ paulum sexto brevioribus et gradatim
attenuatis. Pedes robusti, art. tarsorum basali penultimo duplo
longiore ; unguiculis fortiter utrinque bifidis.
The genus Monocesta, as thus defined, is very natural: it
represents those species in which the elytra are postmedially
dilated, the thorax is transversely depressed, and the antennze
in the more broadly ovate species filiform, in the more parallel
species subincrassated, the third and fourth jomts being sub-
equal, and the apical joints sufficiently produced and attenuate.
These characters comprehend two distinct subgroups, which will
constitute an excellent genus, well bounded and separated from
the several other forms with which the species representing
them have been, in Dejean’s Catalogue, mixed up. The metro-
polis of the genus is evidently the tropical region of South
America and Mexico. One species (M. cory) is found as far
north as Illinois, where it infests the hazel; and one other (M.
elegantula of this paper) I have received as from Brazil.
Diviston A.
Species of large size; in form (for the most part) postmedially
dilated; the thorax is deeply transversely depressed ; the an-
tenne filiform and sufficiently elongate. Species 1-12.
Section I.
‘lytra for the most part bright blue or bright green, with flavous
markings. Sp. 1-9.
1. M. imperialis.
MM. grandis, apice dilatato, crebre punctata, nigro-cerulea, elytrorum
Rey. H. Clark on Dejean’s Genus Coelomera. 265
dimidio apicali rufo-flavo ; ‘caput fovea longitudinali ad frontem,
impunctatum ; thoraw subtiliter punctatus ; scutellum transverse
quadratum, impunctatum ; elytra thorace latiora, et versus apicem
multum ampliata, marginata, crebre punctata; antenne, corpus
subtus et pedes nigra, femorum apicibus rufo-fuscis.
Long. corp. lin. 93; lat. lin. ad humeros 4, ad ¢ apicem i
Columbia.
2. M. equestris, De}.
M. e majoribus, versus apicem subampliata, crebre punctata, viridis
vel azurea, fascia media in elytris transversa flava; caput longi-
tudinaliter foveolatum, punctatum ; thorax ut in speciebus aliis in
medio valde transverse depressus, punctatus ; scutellum sub-
quadratum, impunctatum ; e/ytra versus apicem ampliata, crebre
et rugose punctata, fascia recta flava transversa ad medium, versus
margines latior, elytra ornat; antennae, corpus subtus et pedes
nigra,
Long. corp. lin. 9-63; lat. lin. 53-23.
M. equestris, the Cayenne species, differs clearly and uniformly
in pattern from the Mexican species, M. ducalis. In the species
before us, the transverse band on the elytra is medial and even,
sometimes, by reason of its greater breadth towards the margins,
being slightly inflected; in M. ducalis the transverse band is
never straight, but arcuate, in each elytron, broader in size and
more irregular in pattern.
Mr. Baly has shown me in his cabinet a species taken by Mr,
Bates on the Amazons, in which the transverse band is a trifle
broader, and rather nearer the scutellum, than in M. ducalis—
M. Batesi, Baly. It is probable that the two are slight modi-
fications of the samc species.
The males of M. equestris have sometimes the apex of the
elytra, at a little distance from the suture, produced into an
angle.
A common species in Cayenne.
3. M. ducalis.
M. elongata, versus apicem subampliata, punctata, nigro-czerulea,
fascia lata et apice flavis; caput foveolatum, punctatum ; thorax
transverse depressus, leviter punctatus; scutellum leve; elytra
crebre punctata vel subrugosa, nigro-ceerulea, fascia lata ineequali,
ad margines versus humeros et apicem tendente, et apice flavis ;
pedes, antenne et corpus subtus nigra.
Long. corp. lin 8-6; lat. lin. 5-3.
I have before me several specimens of this form, which agree
entirely in pattern, and are readily separated from the Cayenne
species, M. equestris: the band of the elytra is broader, more
irregular in outline, and extending along the margination to the
apex, and sometimes to the shoulders. I have an interesting
266 Rev. HH. Clark on Dejean’s Genus Coclomera.
variety in which the transverse band is represented by a medial
spot, the margins being obscurely flavous.
Mexico. A common species.
4. M. splendida.
M. versus apicem leviter ampliata, punctata, leete viridis; caput ob-
solete punctatum ; thorax crebre punctatus, ad medium transverse
depressus ; scutellum subquadratum, leviter punctatum; elytra.
crebre punctata (punctis minutissimis) et leviter rugosa; corpus
subtus fusco-yiride, testaceo pubescens; antenne nigre ; pedes
nigro-virides.
Long. corp. lin. 73, lat. lin. 43.
Conspicuous among its congeners by its uniform brilliant
green colour.
Para. I have in my collection a single example, received
from the Marquis La Ferteé.
5. M. consularis, De}.
M. subparallela, postice paulum dilatata, satis robusta, vix depressa,
tomentose rugosa, vix punctata, rufo-flava, elytris (fascia media
excepta rufo-flava) viridibus ; caput longitudinaliter foveolatum,
rugosum, rufum; ¢horaw rugosus, rufus; scuted/um rufum ; ely-
tra subparallela, apice transverse rotundata et subdilatata, sub-
tiliter rugosa, ceerulea, fascia media transversa eequali rufo-flava ;
antenne fusco-nigre, art. 1° rufo; pedes et corpus subtus ruta.
Long. corp. lin. 63; lat. lin. 33.
This species resembles as to pattern the Pyesia laticornis of
Germar; but, besides being a larger insect, it is more robust,
less depressed, and the surface is not, as in that species, punc-
tate, but thickly rugose and tomentose.
Brazil. One of the examples in my collection is labelled
Mexico.
Section II.
Elytra for the most part flavous, with darker markings. Sp. 6-12.
6. M. illustris.
M. satis lata, subparallela, crebre punctata, pube flava subvestita,
rufo-flava, elytrorum apicibus fuscatis; caput fronte foveolatum,
punctatum ; thorax transverse ad medium depressus, punctatus ;
elytra leviter pubescentia, punctata, rufo-flava, apicibus nigris ;
antenne, pedes et corpus subtus migra.
‘Long. corp. lin. 6; lat. lin. 33.
Cayenne.
7. M. coryli, (Caelomera) Say, Acad. Philad. i. 1824, p. 455;
Complete Writings, vol. u. p. 220.
This is the only species which represents the genus on the
Rey. H. Clark on Dejean’s Genus Coelomera. 267
North American continent: it is found in Illinois, Maryland,
and Virginia; in the latter State it is so abundant that it often
entirely strips the Corylus americanus, on which it feeds.
8. M. depressa.
M. late ovalis, valde depressa, flava, antennis, elytrorum marginibus
et apice necnon corpore subtus nigris ; caput longitudinaliter foveo-
latum, leviter punctatum; ¢horax valde ad medium transverse
depressus, impunctatus; sewtellum flavum ; elytra lata, valde de-
pressa, late marginata subtiliter rugosa, subpubescentia, flava,
apice et marginibus late nigrescentibus ; antenne nigre, art. 1°-3™
flavis ; corpus subtus nigrum ; pedes flavi.
Long. corp. lin. 53; lat. lin. 33.
I am indebted to Mr. Baly for this species, received by him
from the river Magdalina. »
9. M. Balyi.
M. late ovalis, nigra, thorace, capite, elytrorum basi et femoribus
anticis rufo-flavis; caput leviter longitudinaliter foveolatum, im-
punctatum ; thorax late transverse depressus, lateribus ad medium
subangulatis, impunctatus ; scwted/wm impunctatum, rufum ; e/y-
tra ampliata, late marginata, subdepressa, rugosa, opaca, nigra,
basi tenuiter fulvescente ; antenne fusco-nigre, art. 1° et 2° flavis ;
corpus subtus nigrum ; pedes nigri, femoribus anticis rufo-flavis.
Long. corp. lin. 7; lat. lin. 4.
Brazil.
10. M. elegantula.
M. pallide purpurea, thorace maculisque 4 elytrorum flavo-pubes-
centibus ; caput ad basin foveolatum ; thorax omnino flavo
pubescens ; e/ytra macula utrmque magna media, alteraque minore
circulari ad apicem flavo pubescentibus; antenne rufo-fusce ;
pedes et corpus subtus fusco-purpurea.
Long. corp. lin. 5; lat. lin. 3.
Brazil.
11. M. Hopfneri, De}. Cat.
M. subparallela, crebre punctata vel rugosa, testaceo-flava, tibiis,
tarsis et antennis nigris; caput leviter longitudinaliter foveolatum;
thorax transversus, subquadratus, transverse modice depressus,
lateribus rotundatis, crebre et fortiter punctatus; sewtellum sub-
triangulare, subpunctatum ; e/ytra thorace latiora, subrugosa vel
crebre punctata; antenne sat elongate, nigre; corpus subtus
fuscum ; pedes flavo-fusci, tibiis et tarsis nigris.
Long. corp. lin. 5; lat. lin. 25.
M. Hopfneri may be distinguished not only by its flavous
coloration, but by its thorax, which is less deeply transversely
depressed, and coarsely and closely punctate.
Mexico.
268 Dr. W. R. Scott on the Occurrence of
12. M. sanguinicollis.
M. eclongatula, postice subdilatata, crebre punctata, nigra, thorace
rufo, elytris rufo vittatis; capué punctatum, rufum ; ¢horaz in
medio transverse depressus, lateribus rotundatis, punctatus, rufus;
scutellum impunctatum, rufum ; e/yfra versus apicem subdilatata,
satis depressa, crebre punctata, rufa, vittis duabus parallelis (sub-
suturali et laterali) basin sed vix apicem attingentibus, post medium
elytrorum conjunctis ; anfenne nigre, art. basali interdum rufo ;
pedes et corpus subtus vel nigra vel rufo-fusca.
: Renee sal Anion : 3.91
Long. corp. lin. 33-43 ; lat. lin. 13-27.
I have examples of this species from Bolivia, and also from
Brazil. ,
[To be continued. }
XXXI.—On the Occurrence of Orcynus alalonga on the Coast of
Devon. By Dr. W. R. Scorn.
SEVERAL fish of but rare occurrence in British waters have from
time to time been taken on the Cornish and Devon coasts.
The close and accurate observations of Mr. Couch have seldom
allowed any found on the former to pass unnoticed. On the
coast of Devon, however, recorded captures of these rarer spe-
cies are less common, owing probably in some degree to the
want of that zealous watchfulness which has animated the la-
bours of the Cornish ichthyologist. Amongst the rarer species
that pay our coasts an occasional visit are ‘those of the genus
Thynnus ; and amongst the very rarest of these is the Germon,
separated now, however, by Cuvier into a distinet genus, and
which fish he has named Orcynus alalonga, from the length of
its pectoral fin—which constitutes the chief, if not the only, dif-
ference between it and the true Tunnies.
The Oreynus alalonga has been very rarely found in British
seas. One has been recorded as taken at Portland, which was
presented to the British Museum, and it has been twice taken
in Mounts Bay, Cornwall.
I have pleasure, therefore, in now recording another specimen
of this rare British fish, taken in Devonshire. This fish was
captured on the 26th of August last, not really in channel, but
a little way up the river Exe, about three miles from its mouth,
and at about half-tide. The fish had got entangled amongst
some palings which had been driven into the river about a foot
from the edge, where a kind of quay had been made, and which
formed a cul-de-sac. Into this the fish got ; and so violent were
its struggles to get out, that it drew the attention of some work-
men who were at a little distanee, when one of them got his gun
Orcynus alalonga on the an, 269
\
and shot it. They describe its efforts to free itself as shaking
the palings like the strength of two men, which agrees with the
observations of Mr. Couch, who says that a specimen taken in
Mount’s Bay showed “ extraordinary strength when caught with
a line.”
Unfortunately I did not hear of this fish having been captured
till a week after, and when it had become much putrified, and,
indeed, had been buried; so that I cannot give so full and accu-
rate a description of it as might be desirable. Still enough re-
mained to show to what species it belonged; and on showing
the plates of the Tunmies in Mr. Couch’s volume to the person
who killed it (an intelligent foreman of some works near the
river), he declared it to agree with that of the Germon.
The shot by which the animal was killed had destroyed all
the first dorsal fin and part of the second, so that no examina-
tion could be made of these; but the general form was not
much injured, and the other fins remained sufficiently defined
to enable me to mark their position and numbers; and the pec-
toral fin, one of the most important in identifying the species,
was the least injured of all. From this state of the fish I was
enabled to note the following particulars.
The full length of the fish, from the nose to the base of the
caudal fin, was 24 inches, the girth round the pectoral fin 19, and
the girth immediately in front of the second dorsal 154 inches ;
the flesh in this part was very firm and solid.
The head was pointed, the under jaw slightly the longest, the
teeth small and incurved, and the gape about 4 inches; the
nostrils very obscure ; the eye was large, and, when fresh, was
slightly elevated, and placed over the angle of the mouth. The
sections of the gill-covers were well defined ; and from the nose
to the gill-opening was about 7 inches.
The pectoral fin, lodged in a deep depression, was 84 inches
long, and reached to about the middle of the anal fin. I could
not say if the anal and second dorsal were falcated, as the upper
parts of both had perished; but I was informed that both were
so. I could trace that there was a short space between the first
and second dorsal fins, and that the finlets were eight above
and seven below, which I was told had been tinged with yellow,
but not so deeply as are those in Mr. Couch’s plate. The tail
was deeply cleft; but, one-half being gone, I could only judge
of this approximately. The weight of the fish when caught was
twelve pounds, and nothing was found in its stomach.
On comparing this fish with the figure given by Couch, I
thought the latter more slender for its length than the former ;
but this appearance might have arisen from the body having
become depressed, from the treatment to which it had been
270 Prof. H. J. Clark on the Animal Nature
subjected. The length of the pectoral fin was also not so great
as that figured by Couch. The proportions given in the speci-
men killed at Portland are—length of body 30 inches, length
of pectoral fin 114 inches; while in the specimen examined by
me the length of body was 24 inches, and the pectoral fin
84 inches. ‘This latter length, however, comes quite up to that
given by Cuvier for the species. He says the Scomber thynnus
has the pectoral fin one-fifth part of its length, while the Orcynus
has it one-third the length of the body, and that this difference
is the only one between the two fishes. It will be seen, then, that
84 inches is nearly the proportion given by Cuvier, being a
little more than one-third of 24 inches, the full length of the
fish.
It is fortunate that the pectoral fin was sufficiently perfect
to allow of its beimg accurately measured, and thus enable us
to record another instance of this south- of-Europe fish paying
a visit to our northern shores.
It is further worthy of remark that Mr. Couch has reported the
capture of the Short-finned Tunny (a fish never before taken on
our coasts) on the 16th of August last. It would be interest-
ing to know what causes have led these fish so far north on this
occasion.
XXXIT.—Proofs of the Animal Nature of the Cilio-flagellate Infu-
soria, based upon Investigations of the Structure and Physio-
logy of one of the Peridinia (Peridinium ecypripedium, n. sp.).
By Prof. H. James Ciarx, A.B., B.S.*
[Plate XII.]
Wuartver tends to elucidate the doubtful nature of any group
of beings which stands undecided (as it were on the dividing line)
between sentient and non-sentient things has an importance at
the present day which would not have been deemed worthy of
very grave consideration before the theories of Spontaneous
Generation and what is sometimes mistakenly called Darwinism
had been revived. The resurgence of these doctrines has given
a prominence to the discussion of the character of the lowest,
obscure forms of life, simply because, in their extreme simplicity,
they hardly seem to rise above a state of inorganic nature, and
their vitality is exhibited in such a guise as would readily be
mistaken for the operation of exo-endosmotic, inanimate, inor-
ganic forces. Hence the readiness, the eagerness, with which
the physicists of the Materialistic school clutch at these “toys”
* Communicated by the Author, having been read before the American
Academy of Science and Arts, February 14, 1865.
of the Cilio-flagellate Infusoria. 271
of the older microscopists, hoping therein to find an abundance
of argument by which they may prove that rock and flesh do not
incompatibly jostle each other whenever they come in contact.
Claiming, and justly too, that these extremes of the inorganic
and organic bodies are naturally and incontestably related to
each other through their common basis the simple elements of
the chemist, it does not seem possible to the materialist that
their relations should be changed or dissevered by the introduc-
tion of any modes of existence, however varied or elevated. The
carbon, the hydrogen, the nitrogen, and the oxygen once being
established as definite existences, they always remain C, H, N,
and O, no matter under what forms or relations they may be
disguised, the various modes of being not in the least changing
the. fact of their existence. For instance, they say, the transi-
tion from one kind of animate being to another kind is only a
graduated change in the mode of existence, or of the manner of
an outward expression of the relations of the component elements
of the organism ; certainly not an actual metamorphosis of the
nature of these elements. To this assertion there may not pos-
sibly be any objection ; but if the same explanation were urged
for the transition to the Monad from the infinitesimally small,
vibrating, imorganic corpuscle of the “ Brownian motion,” we
have not come to that state of knowledge of the forces of nature
to accept it so readily as in the former case. Still the growing
tendency, among the philosophical chemists, to merge the vital
and the inorganic forces into one would seem to be inevitably
preparing us to regard such a transition as identical in kind
with that which obtains among the undoubtedly organized bo-
dies, whether animals or plants.
In this state of hesitancy to step across the vanishing line of
demarcation between the animate and the inanimate, we can at
least safely venture to give, in general terms, an expression of
the relations of the three forms of existence. We may say that
it is the mode of existence which constitutes the difference be-
tween the morganic and the organic bodies, or between the two
forms of organic life, viz. animals and plants; so that every
fact enunciated in regard to an animal or plant is the record of
a symbol of one of the methods of existence, or of the nature of
the influences which enter into the life of the being.
From this point of view the study of these insignificants rises
to the rank of the highest philosophical inquiry, and the minute
wonders of the microscopist become the agents in the pursuit
after the knowledge of the ways of the Infinite, which one could
hardly have the temerity to smile at.
These thoughts have been suggested by the results of some
investigations into the thus far doubtful animal nature of the
272 Prof. If. J. Clark on the Animal Nature
Cilio-flagellate Infusoria, as the Péridiniens and their congeners
are designated by Claparéde in his, conjointly with Lachmann,
most recent publications upon the Infusoria*.
In order that the various points of the proof that the Péri-
diniens are undoubted animals may be comprehended in sys-
tematic sequence, it seems most desirable to present them under
separate sections, each devoted to some particular vital function.
Habitat and Form.—There is probably no generic difference
between the species in question here and those described and
figured by Allman in the third volume of the ‘Journal of Micro-
scopical Science,’ 1856, and by Claparede in the memoir above
referred to; but in their specific relations no doubt they are
distinct. This (Pl. XII. figs. 1, 2,3) has an oblique pyriform
outline, more than one-third longer than its greatest breadth,
and hollowed on one side by a broad longitudinal depression (d),
extending from the narrower end (p) to a short distance beyond
the broadest part of the body. Not far from the narrower end
the so-called flagellum (jl) is attached, in the middle line of the
broad depression, and is so long as to project beyond the end
near to which it is situated. As the narrower end (P) is always
the posterior, and the broader end (a) the anterior, in the act of
swimming, and the relations of the other parts of the body,
such as the position of the mouth (m) and particularly the trend
of the cesophagus (@), correspond to these, the one which pre-
cedes should be called the anterior, and the other the posterior
end of the body; and as such they will hereafter be designated
in this article.
There are two shallow furrows which encircle the body; one
(pf), rather broad, passes obliquely backwards and around it
just behind its middle, and the other (af), quite narrow, encir-
cles the broader end just in front of the termination of the broad
longitudinal depression above mentioned. The whole of the
body posterior to the narrower transverse furrow is clothed with
vibratile cilia; but the anterior end is devoid of them, and appears
to be covered by a low cap (pe) in the form of a segment of a
sphere. In the young this cap is so shallow as to be readily
overlooked during the motions of the animal. Close to the
posterior end there is a large, clear vesicle (cv), which is quite
conspicuous, even during the rapid motions of the animal. This
is the contractile vesicle, which will be described presently. In
point of sensitiveness this Peridinium exhibits it in almost as
ereat a degree as Plewronema and many other timid Infusoria,
These are the most evident and striking features, such as readily
attract the attention when the body is in motion ; and moreover
* Claparéde and Lachmann, “ Etude sur les Infusoires et les Rhizo-
podes,”’ Mém, de l’Ins. Genévois, tomes y., vi-, vil. (1858-1861),
of the Cilio-flagellate Infusoria. 273
they are the chief and characteristic traits of this species. The
specific name is derived from the resemblance in the form of the
body to the labellum of one of the Orchidaceze, viz. Cypripedium.
It is very common in the freshwater ditches and slow streams
about Cambridge,U.S. ; and in the aquarium congregates in great
numbers around decaying matter. It varies from 3}; to =;
of an inch in length; but occasionally adults were found +; of
an inch long. It is probable, however, that the latter were in a
preparatory state, just before self-division. The colour is a uni-
form light brown, which resides mostly in the derm.
Contractile Vesicle.-—This organ (cv) is so conspicuous, in the
species before us at least, that one is apt to wonder why it has
not been discovered before. The only reasonable excuse for this
seeming delinquency would appear to be, that the animal is so
incessantly active and so rapid in its motions, that a large amount
of patience could hardly compensate for the want of a quiet
subject. Fortunately, at the present day our lenses, even of
moderate power, are constructed with such large angles, broad
fields, and excellent definition, that the difficulty of keeping the
Infusorian in sight, and of getting a clear decided view of its
interior, is almost done away with. By strewing the glass slide
with abundance of indigo, little lagoons are formed here and
there, in which, when the specimens are plentiful, there is no
difficulty in findmg and confining any particular individual,
without the necessity of a thin glass covering. In this way the
motions of the body are reduced to a simple revolution on its
longer axis, with an occasional inversion, end for end. The eye
soon gets accustomed to the rhythmical appearance of any par-
ticular region as it comes round at each revolution, so that, by
a systematic study of each and every feature, a knowledge of the
whole organism may be obtained as readily as in most Infusoria.
The contractile vesicle is invariably situated close to the nar-
rower, or posterior, end of the body, but at a considerable dis-
tance from the ventral, dorsal, or lateral surfaces. At the mo-
ment just before systole it has a perfectly globular form (Pl. XII.
fig. 1 cv), and a very sharp, strongly refracting, conspicuous
contour, and occupies rather more than the middle third of the
transverse diameter of the body at this point. The systole and
diastole are as regular in their recurrence as in any of the
ciliated Infusoria, and as conveniently observed. The systole,
in perfectly fresh specimens, occurs with perfect regularity once
in forty seconds, as numerous and carefully registered observa-
tions prove. As in other Infusoria, between diastole and sys-
tole, the vesicle is more or less irregular in outline, but gradu-
ally approximating to a spherical form. At the moment of sys-
tole it rather quickly changes from a broad spheroidal figure to
Ann. & Mag. N, Hist. Ser.3. Vol. xvi. 19
274 Prof. H. J. Clark on the Animal Nature
one which is globular, and then contracts suddenly and rapidly
until it is nearly invisible. The diastole then follows slowly,
aud during this it passes from a jagged rounded outline (fig. 3 ev)
to a lenticular form (fig. 2), then to a hemispherical shape (fig. 4)
with the flattened side next to the posterior end of the body,
and finally, assuming a spheroidal contour, it remains quiet
awhile, until the time for the next systole. If the water is not
renewed, the specimens become unhealthy, which they exhibit
by changing their form, and swelling up into an oval and finally
a globular mass. In such a condition the systole of the con-
tractile vesicle oftentimes occurs five or six times in a minute,
and will continue at that rate even when the animal is very
much flattened out, and until it bursts or falls to pieces. Time-
ture of opium stops the action of the contractile vesicle almost
immediately, even before the rest of the body is sensibly affected
by it. The effect is to swell the vesicle to an enormous size ;
and then, breaking through the posterior end of the animal, it
expands to a dimension often exceeding that of the whole body,
before it bursts.
The Mouth (m).—That this creature has a mouth might be
premised from the manner in which particles of indigo or car-
mine approach and recede from the body. When the animal is
moored by its flagelliform appendage (fl), and gyrates about it
as if on a pivot, these particles of coloured food may be seen to
pass along the face of the broad longitudinal depression (qd),
and, striking the body just behind its mid-region, glance off in
a backward direction. At the point where the indigo strikes
may be seen an obliquely longitudinal ovate opening (m), which
leads into an elongated funnel-shaped cavity (@): the former is
the mouth, and the latter is the cesophagus. The mouth hes
altogether within the posterior obliquely transverse furrow (pf),
and extends from its anterior to its posterior edge, trending
diagonally across the axial plane of the body, from the right,
backwards, toward the left. Its anterior edge (m!) is bro oad,
and thence it gradually narrows to a sharp angle, which forms
the posterior edge. It is so inconspicuous that in all probability
it is nearly or altogether closed, except when taking in food;
certainly it is not one of the prominent features of the organism,
although one of the most important. When the animal is in a
sickly condition, and swollen up, the mouth is easily deseried ;
but its relations are not readily made out, because in this state
the annular furrows are all obliterated; yet its connexion with
the cesophagus at such a time is clearly seen. There are no
appendages whatever about or belonging to the mouth; not
even the flagelliform body (fl) has anything to do with it, but
is attached to the body at a very sensible distance ( fl’) behind
of the Cilio-flagellate Infusoria. 275
it. It would seem, therefore, to be dependent upon the simple
cilia around it for the transfer of food to its lips. From the
mouth the wsophagus (@) passes obliquely backwards and toward
the dorsal region, at least halfway through the body, and then
terminates rather abruptly just before the contractile vesicle,
but a little to the right side (fig. 8 @) of the axial plane. At
the mouth it is widest antero-posteriorly, but suddenly narrowing
a little, it afterwards gradually lessens its calibre as it extends
into the body, and finally ends as just described. The whole
track of this channel is much more readily seen than the mouth.
The food is taken in such excessively small particles that its
entrance into the mouth cannot be detected with any degree of
satisfaction; and a single digestive vacuole (dv) requires from
twenty minutes to half an hour to form and fill; and although
it may be comparatively quite large, even two- thirds of its fullest
capacity, yet so infinitesimally minute are the particles, that
even indigo or carmine is not readily seen, although it may be
the only Tem of food present. Beyond ne point, however,
these colouring-matters become rapidly visible, so that when a
vacuole is fully formed, the indigo or carmine is as conspicuous
as in any other Infusorian. These vacuoles are very large, in
fact equalling in size the contractile vesicle; and as they form
sometimes pretty far back, they are apt to obscure the latter—
without doubt thus causing this vesicle to be mistaken for one
of them, since they bear a certain resemblance to it. No anus
was detected during these investigations, although the specimens
at times were kept well fed.
The Locomotive Organs.—The most prominent among the cilia
is the so-called flagellum (fl). This, however, is not a single
filament, as has usually been asserted ; but, owing to the manner
im which it is used, it very naturally appears to be so. Most
frequently its compound nature becomes apparent when the
numerous Cilia of which it is composed divide into two groups
(fig. 7 fl, fl), thus simulating a double flagellum*. At other
times, after having divided into two groups, they twist about
each other in such a way as to resemble a sharply pointed screw,
with a long drawn-out double thread. Such is the condition in
which this pseudo-flagellum is most frequently seen, and then,
with the best magnifying -powers, up to five hundred diameters,
its compound nature is not easily recognized. But there are
times when the whole group of cilia spreads out into a distinct
brush, so that each individual cilium may be seen. The base
(fl) of attachment is in the axial plane of the body, a short dis-
* Claparéde (loc. cit.) speaks of frequently noticing that some of the
Ceratiums, &c., appeared to have a double flagellum. Probably they were
a group of cilia divided as here described.
19
276 Prof. H. J. Clark on the Animal Nature
tance posterior to the mouth (m), and distinctly disconnected
from it, as has been already noticed. When not in motion,
which seldom occurs, the brush les along the median furrow
(mf), which trends from the mouth to the posterior end of the
animal; and in this position it projects for nearly half its length
beyond the body. Its most ostensible use would seem to be
that of a sort of rudder when the creature is swimming, and as
a means of attachment when not progressing. The body may
be seen gyrating and at the same time revolving on its longi-
tudinal axis, for long periods, around a point to which the
pseudo-flagellum is attached, and upon which it turns like
a pivot. Most frequently, during this act, a part of the
brush separates from the rest, and performs the office of an
extra propeller. When the animal is darting and spinning
through the water, this appendage projects obliquely from its
point of attachment (as in fig. 1), and always following, with
the narrower end of the body, in the rear, it seems pretty evi-
dently to be the main agent in the various and sudden tackings
to which this Infusorian is addicted, and also the axis upon
which the body revolves; at least the latter whirls, repeatedly
changing as quick as thought from right to left, or vice versa,
upon an imaginary axis, which is oblique to its greatest length,
and which exactly corresponds to the trend of the flagelliform
appendage when operating in this capacity. Under these con-
ditions the animal shoots along with a compound motion, which
might be described as wabbling, or like the action of an excen-
tric wheel. Apparently in confirmation of this view, the annular
obliquely transverse anterior (af) and posterior (pf) furrows
trend almost exactly at right angles to this imagimary axis.
These two furrows seem, at first sight, to be bands of vibrating
cilia; and in fact it is in the line of their trend that these cilia
are most readily detected, simply because they are rather more
crowded along their edges than elsewhere ; but an attentive ex-
amination reveals their presence all over the body, posterior to
the anterior transverse furrow. Between the two furrows (7. e.
from af to pf) they are longer than at the narrower end of
the body, and at both points they have a pretty uniform length,
moderate extension, and are very delicate, so as not to be easily
observed when the body is in motion. At the anterior trans-
verse furrow they appear to be a little longer than elsewhere,
and, acting more or less in concert, they have the semblance of
a wreath disposed along the edge of the low skullcap-like cover-
ing (pe) of the anterior end.
The Cuirass ( pe).—It is pretty evident that in the species be-
fore us. this is a mere dermal specialization, without any trace
of indurated matter which would entitle it to the name of a
of the Cilio-flagellate Infusoria. 277
genuine cuirass. Where vibratile cilia are present, no such
covering can be said to exist; and as the broad anterior end (a)
of the body is devoid of them, its skull-cap covering is the only
portion of the derm where one could expect to find a cuirass.
But this it is only in form, since it participates with the rest of
the body in the general expansion when an individual is dying.
It has, without doubt, a different character from the rest of the
skin; for the style a ornamentation is not of the same kind,
and, curiously enough, too, it is less truly ornamented than
the other regions of the body, amounting to a mere scattered
punctuation ; whereas over the field where the cilia prevail these
punctuations, which are in reality minute, cylindrical, strongly
refracting bodies standing perpendicularly to the surface of the
derm, are arranged in perfectly regular rows, which have a dif-
ferent character in the three regions posterior to the pseudo-
cuirass. In the space (fig. 5 p) between the anterior (af) and
posterior (pf) transverse furrows, the rows trend longitudinally
and transversely ; in the posterior transverse furrow (pf) they
have the same arrangement as the last, but they are more closely
set together; and in the region behind the latter furrow they
trend in decussating lines (e), like those in the carapace of
Arcella vulgaris.
This region is also characterized by being divided longitudi-
nally, on the ventral side, by a furrow (fig. 1, 2, 6 mf) which
trends in a direct line from the end of the body to the mouth,
and gradually widens anteriorly, where it joms the annular
transverse furrow (pf). At this point of juncture the flagellar
appendage arises, and opposite to it the anterior edge of the
transverse furrow just mentioned forms an inequilateral angle at
the broader margin (m') of the mouth, so that the right and left
halves of this furrow are rendered asymmetrical—a character im
perfect accordance with that of many, if not of all, the Peradinia.
The Nucleus (n).—At the period when these observations were
made, viz. early last December, the genital organ invariably lay
transverse to the longitudinal axis, and occupied a very large
portion of the bulk of the posterior end of the body. Most
frequently it had a U-shaped form (fig. 3), and embraced the
contractile vesicle with its two limbs. It was then of a yellowish-
brown colour, and perfectly homogeneous. Occasionally it was
observed to be divided into three or four masses, which extended
toward the region encompassed by the posterior annular furrow.
While in the U-shaped form, the whole semiopaque mass was
enclosed in a transparent envelope (ne). Oftentimes there was
to be seen immediately over and close to the dorsal region of the
nucleus, and directly im the plane of the axis of the body, a mi-
nute, clear, vesicular corpuscle (fig. 3 ¢), which seemed to have
278 Prof. H. J. Clark on the Cilio-flagellate Infusoria.
the character of a “nucleolus” or (as is now becoming the be-
lef, since the investigations of Balbiani and Claparéde)
testicle.
Reproduction from the egg has not been observed, but trans-
verse division occurred in a number of instances. In the latter
case it agrees, in the process, with what Allman (loc, cit.) has
described, excepting that the resultants (fig. 6, 1. 11.; fig. 7) are
quite different in their proportions from the adults (figs. 1, 2,8).
At the moment of separation the young offshoot (fig. 7) is about
two-thirds the size of the adult, and is almost as broad as long,
and bulges strongly on the ventral side (v), in front of the
mouth (m). It has a very flat anterior end, and the pseudo-
cuirass (pe) of this part is represented by an inconspicuous
unguiform body. The anterior transverse furrow, on account of
its narrowness, hardly attracts attention, except along its ventral
edge (af), where it is rendered conspicuous by the strong pro-
jection of the unguiform cuirass. As in the adult, it is broadest
ventrally, but, growing shallower, thins out (af), going dorsally
to almost nothing. The relations and structure of the various
organs, cilia, &e., are the same as in the full-grown individuals;
but with progressing growth the proportions of the different
regions of the body change insensibly, as may be seen by com-
paring figures 7, 4, and 1, which are respectively representations
of the youngest, middle-aged, and adult individuals,
EXPLANATION OF PLATE XII.
In all the figures the same letters refer to corresponding parts. |
8 ] SI
A. The anterior end of the body. pv. The posterior end.
D. The dorsal side. v. The ventral side. Rr. The right side. L. The
left side.
af, anterior transverse or annular furrow; af}, dorsal part of af;
pf, posterior transverse or annular furrow; mf, median or longitudinal
furrow; d, depression on the ventral side; cv, contractile vesicle ;
m, mouth, m', anterior edge of m; @, oesophagus; dv, digestive vacuoles;
Ji, pseudo-flagellum, jl, base of fl; n, nucleus or gener rative or gan; ne, en-
velope of n (this is the reproductive organ, pr operly speaking, ‘and n is the
conteuts or reproductive material, the future eggs); ¢, nucleolus or testes ;
pe, pseudo-cuirass.
I., 11. The two products of self-division. 111. The annular constriction
which finally separates 1. and 11.
Figs. 1 to 7. Peridinium cypripediun, n. sp.
g pry
Fig. 1. Profile of an adult, seen from the left side; magn. 500 diams.
Fig. 2. View of the ventral side of an adult; magn. 500 diams.
‘ig. 3. Posterior view of an adult, the anterior end in the distance ;
magn. 500 diams.
Fig. 4. A young individual ; magn. 300 diams.
Fig. 5. An adult, gradually dried up; dorsal view, to show the arrange-
Bibliographical Notices. 279
ment of the punctiform ornamentation of the derm; magn.
500 diams.
Fig. 6. The process of self-division, just half an hour before separation ;
ventral view; magn. 200 diams.
Fig. 7. Profile of 1. fig. 6, just at the moment of separation; magn.
200 diams.
BIBLIOGRAPHICAL NOTICES.
Travels and Researches in Crete. By Captain T.A.B.Sprart, R.N.,
C.B., F.R.S. &c. In two vols. 8vo. London: Van Voorst,
1865.
In carrying out the Mediterranean Survey, the Island of Crete
came under examination by Capt. Spratt, whose acquaintance with
the requirements of his own profession, with the ancient and mo-
dern history of the Greeks, their early works of art, coins, monu-
ments, and. buildings, with the natural history of land and sea in
the Mediterranean area, and with the geological structure of every
mountain, coast, and islet he visited, render him peculiarly capable
of doing justice to so interesting a region as Crete. The form and
character of,that island, from mountain to plain, the sites of its cities,
its ravines, caves, and water-courses, are so visibly explicable by
their rocky structure, that to shut one’s eyes to their geological is
to misinterpret their topographical relations. Its highlands and
valleys, as well as the coast and the deep sea, are strikingly remarkable
in their natural products. Its old forgotten cities rise up to intelli-
gent research, and the ancient ruins take definite form and their true
place in history, when learning and sagacity unravel the half-true
legends of the place. In Crete are found statuary and coins of the
finest style, and of a school dating from an earlier time than Athenian
art could boast of; for it was the cradle of Greek learning and much
of Greek mythology. Lastly, there still exist genuine Cretan Greeks,
whose ancestors (under the Roman sway) heard Paul preach at Fair
Havens,—under the Byzantines, Saracens, Franks, and Venetians,
played their medizeval part in quarrels, bigotry, and trade, and, well
versed in war, withstood the Turk for more than twenty years,—
and under the Turk have suffered all that brings out the debasing
vices and exceptional virtues of a conquered race.
Vollowing Capt. Spratt in his account of Crete (the eastern part
of which he more particularly treats of, as having been left unde-
scribed by Pashley), we find the natural features of the country, the
remnants of Greek buildings and works of art, medizeval relics, the
peculiarities of the present population—the old highland Sfakiote
breed, hardy, unscrupulous, and cruel, and the lowland Candiotes of
mixed origin—all carefully noted and elucidated by a scientific
acquaintance with nature and by a knowledge of classic literature
and history ; whilst an eye for beauty in anne and art—enthusiasm
in working out the traces of long-past civilization, the early source,
in great part, of our present culture—a warm sympathy with all
280 Bibliographical Notices.
that is human, lowered though it be as the outcast leper of benighted
Crete—and a hearty, honest, common-sense view of men and man-
ners, give a good tone and genuine feeling to all his observations.
In fact, the naturalist, geologist, geographer, antiquary, and general
reader cannot fail to be interested and instructed by this work. Its
illustrations are first-rate : two excellent geological and topographical
maps; a dozen good chromo-lithographs of scenery, with some other
plates; numerous small lithographs on india-paper inserted in the
text, besides several woodcuts, are all well executed, and help the
reader. A delicately tinted lithograph of Cestum Venerts and Beroé
illustrates a long and careful account of these beautiful creatures. A
chapter is devoted to the sponge-divers and their surroundings ; and
a picturesque group of their fishing-boats is shown in a coloured plate.
Appendices on Cretan and modern Greek (by Viscount Strafford);
on Deep-sea Soundings ; on Currents in the Mediterranean ; on the
Salinity of the Black Sea and Mediterranean; on the Geology of
Crete, and its relations with Malta and Africa; on the Birds (by
Col. Drummond-Hay) and the Land-Shells of Crete ; and on the
Greek inscriptions found in Crete (by Dr. Churchill Babington),
carry out more fully some of the researches and favourite topics
of our author.
One of the characteristics of Capt. Spratt is most pleasantly shown
in the honest and genial acknowledgment of the labours of his col-
leagues in the Nautical Survey, of the aid of other friends in his
scientific and literary work, and of the strong and lasting influence
that he believes the genius and philosophy of his lamented friend
Edward Forbes have had in rousing, shaping, and supporting that
activity of research which is so handsomely represented by these
volumes—which is so well known by many circles of his countrymen
and foreigners, and always so modestly referred to by himself.
Handbook of British Water-weeds, or Alga. By Dr. Joan Epwarp
Gray, F.R.S., late President of the Botanical Society of London.
The Diatomacea, by W. Carrutuers, F.L.S. &c. — London:
Hardwicke, 1865.
Tus little work contains an arrangement of all the Algee or Water-
weeds hitherto recorded as found in Great Britain and Ireland, re-
ferred to the most recent genera, and fills up a desideratum that has
for several years been felt by the botanical student.
The black- and red-seeded Algae, which, with very few exceptions,
are all marine, are arranged in the families, genera, and subgenera
used by Professor Jacob George Agardh in his ‘ Species, Genera, et
Ordines Algarum,’ lately published in Sweden, with the alterations
suggested in the system proposed by Professor Harvey, in his ac-
count of the American Alge, published by the Smithsonian Insti-
tution. The species are all accompanied by a short diagnosis and a
reference to the best figure which has been given of them from spe-
cimens in a living state, Harvey’s ‘ Phycologica Britannica’ being the
work almost always referred to.
Bibliographical Notices. 281
The Green Algee (Chlorosperme), which contain both freshwater
and marine species, are arranged according to the system proposed
by the author in his paper on the distribution of those Algze, pub-
lished in the ‘ Annals of Natural History’ for November 1861.
As these plants are very difficult to be distinguished, except in a
living state (the chief character often disappearing when they are
dry, and indeed often shortly after they are gathered), the author
has not attempted to give any diagnosis of the species, but has only
referred to the works in which the species or presumed species are
figured, preferring, where he can, figures that are taken from living
specimens.
Dr. Gray has suggested some improvements in the arrangement
of the Algee. Thus he has proposed to separate the families of
Melanosperme used by Agardh and Harvey into three orders,
according to the structure of the frond; thus—
Order I. Scyropuyces. Frond leathery or membranaceous, formed
of compact cellular substance : containing—l. Fucacee ; 2. La-
minariacee ; 3. Dictyotacee; 4. Sporochnacee.
Order II, Tricuoruyces. Frond subarticulate, with a jointed
axis, and furnished with tufts of pinnate, jointed (deciduous)
threads. 5. Arthrocladiacee.
Order III. AnrHropuyces. Frond formed of jointed filaments,
which are either free or united into a compound body. 6. Chor-
dariacee ; 7. Ectocarpacee.
In the families he has characterized three new genera, viz.,
1. Fasciaria for Laminaria fascia.
2. Spherophorus for Ectocarpus granulosus and its allied species.
3. Hincksia for Ectocarpus Hincksit.
In the Rhodosperms he regards the anomalous genus Hapalidium
as the type of a family. It has been suggested that that genus may
be only the very young state of Melobesia; but this theory wants
further examination, as the glassy texture, the form of the frond,
and cells are very unlike those of any species of the latter genus,
which is always calcareous and opake, and formed of several layers
of cells, even in its thinnest state of development. Again, if it is
the young state of that very common and universally spread genus,
why is it so seldom observed, when the Melodesie are to be seen on
almost every kind of marine body ?
In the Chlorosperms, Dr. Gray has characterized the following
genera as new, V1z.—
Leptocystea tor Cladophora pellucida.
Vagabunda for Cladophora fracta.
Cystothrix for Cladophora Rudolphiana.
Cystophora for Cladophora littorea.
Calonema for Callothrix mirabilis.
The list of the Diatomaceze seems to have been prepared by
Mr. Carruthers with great care; and it will be very useful for the
collectors of that very numerous and intricate class of minute plants.
282 Bibliographical Notices.
Natural-History Transactions of Northumberland and Durham.
Vol. I. Part 1. 8vo. 1865.
The ‘Natural-History Transactions of Northumberlandand Durham,’
of which this is the first part, are to be looked upon as a continuation
of the ‘Transactions of the Tyneside Naturalists’ Field-Club’ under
a different title, being in fact the Proceedings of the ‘ Natural- His-
tory Society of Northumberland, Durham, and Newcastle- on-Tyne,’
incorporated with those of the Tyneside Naturalists’ Field-Club.
To make perfect catalogues, zoological and botanical, for the
natural history, recent anal fossil, of Northumberland and Durham
is a main object of the Tyneside Field-Club and of the naturalists
now associated with them. This aim is well kept in view in the
present volume of their Transactions, which is largely composed of
“Reports of Deep-Sea Dredging on the Coasts of Northumberland
and Durham in 1862-64,” edited by Mr. G. S. Brady. Among the
new or little-known species are especially mentioned :—Echinoderms
—Lehinocardium pennatifidum, Norman, MS., Psolus squamatus,
Echinus pictus, Norman, Antedon rosaceus, Ophiocoma nigra. Mol-
lusea—Chiton albus, L. Several stalk-eyed Crustacea—Adtelecyclus
heterodon, Pagurus Cuanensis, P. Hyndmani, P. ferrugineus, Cran-
gon Allmani, C. spinosus, C. nanus, C. fasciatus. Of Amphipods,
Lysianassa Coste and several otbers. Of Ostracods, six new species
of Cythere were taken; also a new and very interesting Copepod
(Calanus Clausii, Brady) and a new Pycnogon (Nymphon rubrum,
Hodge). Among Polyzoa, Lepralia annulata and Tubulipora lobu-
lata are new to. the coast. Lastly, some rare Hydrozoa were col-
lected ; and five Foraminifera were added to the local list.
Mr. Joshua Alder reports on the Mollusea, Tunicata, and Zoo-
phyta; the Rev. A. M. Norman on the Crustacea; Mr. G. 8. Brady
on the Pelagic Entomostraca; Mr. G. Hodge on the Pyenogoncidea
and the E chinodermata ; ; and Mr. H. B. Brady on the Foraminifera.
These reports are er with tabulated catalogues showing
the species found in 1862, 1863, and 1864 respectiv ely, with notes
as to frequency and other conditions. Eight plates illustrate this
part of the volume.
Mr. Norman, in the next succeeding memoir, describes Cyanea
imporeata (a new Medusa taken off the “Northumberland coast), and
illustrates it with a beautiful chromolithograph (pl. 11) by T. West.
Mr. Alder then describes three new or rare Polyzoa (pl. 8)—Hschara
Landsborovii, Johnston, L. pavonella, Alder, and Scrupocellaria
Delilii, Avdouin, Mr. Kirkby’s paper on some remains of Fishes
and Plants from the Upper Limestone of the Permian series of Dur-
ham succeeds, with Plate 9, illustrating Pal@oniscus altus, P. va-
rians, and P. Abdsii. The next monoid is entitled “ A Catalogue of
the Recent Foraminifera of Northumberland and Durham, by H. B.
Brady,” with Plate 12, in which ten forms figure as new or rare in
the British seas. One point of interest mienboned in this paper is
the occurrence of certain Foraminifera in brackish pools at Hylton
Dene, and near the mouths of the Wansbeck and the Coquet, “such
Zoological Society. 283
as Quinqueloculina agglutinans, Polystomella striato-punctata, No-
nionina depressula, Rotalia Beccarii, Trochammina inflata (abun-
dant), and Globigerina bulloides (one specimen). These often pre-
sent modified shell-structure, and are evidently the remnants of sea-
born families, left to struggle with the adverse influence of fresh
water, herein reminding us of the marine Crustacea found in fresh-
water lakes in Norway, and of other like instances.
Mr. G. 8. Brady supplies a suggestive paper on Naturalists’ Field-
Clubs and their objects, giving some statistics as to half-a-dozen of
the best, comparing some of the different methods of research
adopted, and concluding with a well-urged plea against the destruc-
tion of small birds by the farmer, even for his own sake, and against
the extermination of rare plants by curiosity-hunting botanists, for
science-sake. Very valuable papers and notes on meteorology (Mr.
Atkinson), flowering-time of plants (Mr. G. S. Brady), entomology
(Mr. Bold), &c., complete this rich volume of natural-history facts
collected by the men of Northumberland and Durham.
PROCEEDINGS OF LEARNED SOCIETIES.
ZOOLOGICAL SOCIETY.
May 9, 1865.—Dr. J. E. Gray, F.R.S., in the Chair.
Description or A New Genus or Tricuturoip FIsHES os-
TAINED AT MADEIRA, wIrH REMARKS ON THE GENUS DrI-
crotus, GUNTHER, AND ON SOME ALLIED GENERA OF TRI-
cHIuRID&. By James YATE Jounson, Corr. Mem. Z. 8.
Order ACANTHOPTERYGIL.
Fam. TRICHIURID.
NEALOTUS, gen. nov.
Body elongate, compressed, incompletely clothed with delicate
scales. Cleft of the mouth deep. Small teeth im the jaws and on
the palatine bones ; none on the vomer. First dorsal fin continuous,
extending to the second; finlets behind the second dorsal and anal
fins. Each ventral fin represented by a single small spine. A
dagger-shaped spine behind the vent. No keel on the tail. Caudal
fin well developed. Seven branchiostegal rays.
This genus may be entered in the synopsis of Trichiuroid genera
in the ‘Catalogue of the Collection of Fishes in the British Museum’
thus :—
Each ventral represented by a single spine ; a dagger-shaped spine
behind the vent.
NEALOTUS TRIPES, Sp. n.
Hurst D2 21> “Second D:. 19, .P. 137) Asis. C. 16.
The compressed body is very elongate, and has a few large deci-
duous simple seales of delicate structure scattered here and there on
284 Zoological Society :—
the skin, which is faintly reticulated with oblique grooves or wrinkles,
and has a steel-grey colour with a silvery lustre. ‘The height of the
body, compared with the total length, is as 1 to 93 ; whilst the length
of the head, compared with the total length, is as 1 to4}. The
black compressed head is flattened above, and is concave between
the eyes, where there are four low ridges, the inner pair of which
enclose an elongated diamond-shaped space. The lower jaw is longer
than the upper, and each is armed with a single series of small del-
toid distant teeth. Those of the upper jaw are inserted in the pre-
maxillary. In front there are seven longer teeth, which are conico-
compressed, and curve slightly backwards ; two of them at each side
stand within the outer row of teeth. On the palatine bones there is
a single row of minute teeth; whilst the vomer is unarmed. The
tongue is also without teeth, and is black like the rest of the mouth
and the inside of the gill-covers. A membrane with a tongue-like
lobe stretches across the palate.
The diameter of the round lateral eye is contained in the head
about five times, and is distant from the muzzle 13 of its diameter.
Near the angle of the preopercle are three very small flat teeth. The
opercle terminates in two obtuse projections separated by a notch.
The first dorsal fin commences a little in front of the root of the
pectoral fin. Its height is rather more than half the height of the
body ; and its length is less than half that of the fish. It rises out
of a groove, and is supported by twenty-one slender spines, which
are not tuberculated. The second dorsal fin commences shortly behind
the termination of the first, to which it is not quite equal in point of
height, and it is less than half as long. It is supported by nineteen
rays, of which the first one or two are short; and it is followed by
two longish finlets. The pectoral fin is inserted under the angle of
the opercle ; it contains thirteen rays, and equals in length the second
dorsal fin. ‘The pair of spines representing the ventral fins are in-
serted close together under the hinder part of the roots of the pec-
toral fins. Their length is about a fourth of the height of the body;
and, being longitudinally grooved, each appears to consist of two or
three spines fused together. ‘The vent is a little behind the middle
of the fish. Behind the vent there is a flat dagger-shaped spine,
which is longitudinally grooved. Its length is less than half the
greatest height of the body ; but it is rather longer than the ventral
spines. The anal fin commences about the length of the spine behind
it, and is opposite to, but rather shorter than, the second dorsal fin.
It contains eighteen rays, and is followed by two finlets, the second
of which is elongated. The deeply forked caudal fin contains six-
teen rays, with five or six short exterior rays on each side.
The lateral line falls obliquely from its commencement above the
opercle to the middle of the length of the fish, and is then continued
with a gentler obliquity along the posterior part of the body to the
tail, where it has two-thirds of the height above it.
The single specimen of this fish which has occurred was obtained
in the month of December, and it has been deposited in the British
Museum. ‘The fish bears a close external resemblance to the
Mr. J. Y. Johnson on a new Trichiuroid Fish. 285
“Coelho” of Madeira (Thyrsites Prometheus, Gthr. ; Prometheus
atlanticus, Lowe). From that fish it may be distinguished by the
possession of a dagger-shaped spine in front of the anal fin*, by the
spines of the first dorsal fin being twenty-one in place of eighteen,
by the rays of the second dorsal fin being nineteen in place of twenty-
one, and by the rays of the anal fin being eighteen in place of six-
teen. It may be further noticed that in the present fish the ventral
spines are placed under the posterior angle of the base of the pec-
toral fin, instead of being inserted a little before that fin, and that
the lateral line does not descend rapidly under the anterior part of
the first dorsal fin, as in Prometheus atlanticus. With Nesiarchus
nasutus it cannot be confounded, since the latter has perfect ventral
fins and fleshy and cartilaginous prolongations of the Jaws.
The dimensions of the fish which has afforded materials for this
description are given in the following table :—
inches.
Wotalpleneth ofp fis lis 2,215) <5) oie clevislnvels cee ncroonene 10
TCIM tee ycreya) spats soscosver stac.e Sings Subisiente = © chose paeianee I5'5
(hiekness behind pectoral’ -..2 ..<). 2 =/cesine= -ess step,
lalate eee Fe aes sec oe anol con. cke oop oF 2-4;
live, dtameter, MEATY, cis ore scram siete c/eierseeais Stores 2
Mliectae lenothworarcest) tases eis se siete otet cater ore i
First dorsal, distance from muzzle .............. 2
Birstudorsaleplenptla) ea. te.c)s ers meuete nacre ereann tte 4t
First dorsal, height in front ..... Beak Jods eta aE 3
Necondudorsallemetly io occ. «tvaseie <j. oie oncusisy sues cui Neath;
Secondudorsal, sere ht sie paiete, eke ol satis ieee wares ahs
Pectoral, distance from tip of lower jaw.......... 2555
eetoralysl Girth pacineclesiegor on oicioyereyerioie or rarneiene lyo
Wientralispimes, lomo < cate wyore vexadeset fevalel acy ale sel = met
Ventral spines, distance from tip of lower jaw .... 2;4
Spine in front of anal, length .................-- oy
/ATPEIL A Gina ns aes eee aoe Reet omr ters dstieste cin. slat
Analy height amuront, soe. 2~ sei criers cele sterile | yy
Wandals ene thity pein. 1c cawiowsner a ofa s)<toneie) o iere re oss ls
The family of Trichiuride is composed, according to Dr. Giinther’s
Catalogue, of the genera Aphanopus, Lepidopus, Trichiurus, Epin-
nula, Dicrotus, Thyrsites, and Gempylus. To these have to be added
the recently described genera Nesiarchus and Nealotus. With re-
spect to Dicrotus, Giinther, a genus founded on a small fish only
2+ inches in length, it appears to me that it ought to be abolished,
the fish having been most probably a young individual of some species
of Thyrsites or Gempylus—an opinion which has been entertained by
Dr. Giinther himself for some time. From Thyrsites Prometheus,
for example, it would seem to differ only by the absence of finlets
and the presence of minute teeth on the vomer. But finlets are not
* Aphanopus carbo, Lowe, and Nesiarchus nasutus, a fish described by me in
the Society’s ‘ Proceedings’ for 1862, p. 173, pl. xx1L, have a similar spine be-
tween the vent and the anal fin.
286 Zoological Society :—
developed in very young fishes, such as Dicrotus armatus probably
was ; and teeth are apt to disappear from the vomer when fishes ac-
quire their full growth. It may be mentioned in confirmation of
this view, that I obtained a scaleless fish, not quite six inches in length,
which had its ventrals reduced to single spines, had teeth on both
palatines and the vomer, and had the last four or five rays of the
second dorsal fin distant from, and unconnected by membrane with,
the rest of the fin; whilst the last two or three rays of the anal fin
were separated from the anterior portion. This was therefore a Di-
crotus with imperfectly formed finlets, showing a closer approach to
a fully developed Prometheus atlanticus than D. armatus,
After attentively considering the descriptions of the species placed
by Dr. Giinther under the genus Thyrsifes (Brit. Mus. Cat. ii. 350),
as well as some of the fiahes themselves, it appears to me that a more
satisfactory arrangement would be to distribute the species amongst
are genera, thus :—
Tuyrsites. Fishes having teeth on the palatines, perfect
ae finlets, and a skin naked or furnished with simple scales.
- Atun, C. & V., and T. lepidopoides, C. & V.
Ruverrus. Includes a single very distinct species, remark-
ae for having a keeled abdomen, and the skin ev erywhere furnished
with bony Hodies each bearing several spines—possessing also teeth
on the palatines, perfect ventrals, and finlets.
R. pretiosus, Cocco.
3. Promerueus. Distinguished by having each ventral reduced
to a single spine, as well as by having teeth on the palatines, finlets,
and a skin either naked or furnished with simple scales.
P. atlanticus, Lowe; P. Solandri, C. & V.; P. prometheoides,
Bleek.
The genus Gempy/us is distinguished from all these by the absence
of teeth from the palatines.
To return for a moment to Ruvettus pretiosus (‘ce curieux, ce
précieux poisson,’’—Valenciennes), the ‘ Escolar’? of Madeiran
tishermen, it may be noted that, although one of the characters given
in the ‘ British Museum Catalogue’ is the want of a lateral line, this
line may be made out in fishes fresh from the sea. It commences
on a level with the upper border of the opercle, but at some distance
behind it, and then descends gently until it arrives at the middle of
the height of the fish, which position it keeps on the posterior half
of the body.
May 23, 1865.—John Gould, Esq., F.R.S., in the Chair.
Notice or a New Species or AUSTRALIAN SPERM WHALE
(Caropon Krerrrit) IN THE SYDNEY Museum. By Joun
Epwarpb Gray, Pu.D., F.R.S., V.P.Z.S., F.L.S., ere.
In a letter which I lately received from Mr. Gerrard Krefft, the
intelligent Secretary and Curator of the Australian Museum, he sent
me some photographs (taken like those formerly sent by Mr. Henry
Dr. J. EH. Gray on a new Sperm Whale. 287
Barnes) of a separate atlas vertebra and of the second and other
cervical vertebree united into one mass of a species of Whale, which
are contained in the museum under his charge. The two bones,
though not united, fit one another so exactly that Mr. Krefft has no
doubt of their having belonged to the same animal; and the photo-
graphs sent justify this conclusion. However, should there be any
mistake in this matter, it will not in the least invalidate the conclusion
that I have come to, from the examination of these photographs,
that they indicate the existence of a second species of Sperm Whale
in the Australian Seas, very distinctly characterized by the subcir-
cular form of the atlas vertebra and of the neural canal in it.
The mass formed by the second and other cervical vertebre is
somewhat similar to these bones in the skeleton of the Australian
Catodon lately received by the Royal College of Surgeons, which I
hope will shortly be described by Mr. Flower, the energetic Curator
of their Museum, who, in his late paper on the Balenide, has shown
how well he can describe and determine the species of Whales.
The genus Catodon should be divided into two subgenera, accord-
ing to the form of the atlas, thus :—
I. The atlas oblong, transverse, nearly twice as broad as high; the
central canal subtrigonal, narrow below. Catodon.
1. CaTODON MACROCEPHALUS of the Northern Ocean. A ske-
leton from Scotland, in the British Museum.
2. Caropon austratis, Macleay, of the Southern Ocean. A
skeleton in the Museum of the Royal College of Surgeons, from
Hobart Town.
Il. The atlas subcircular, rather broader than high; the central
canal circular in the middle of the body, widened above. Me-
ganeuron.
Caropon (MreGanEvron) KRerrtii, sp. nov.
The atlas vertebra oblong transverse, about one-third wider than
high ; the lateral processes only a little produced beyond the articular
surface, with an arched edge; the lower edge arched; the neural
arch low, broad, with a slight central prominence on the upper sur-
face; the canal for the spinal marrow very large, circular, rather
contracted on the sides above, and then dilated, becoming oblong and
transverse.
The atlas is thin, high, being only about one-fourth wider than
it is high. The lower and lateral margins are arched, the lower
edge being the most so. The neural arch is low, transverse, with a
nearly straight lower edge. It is thickest in the middle. The
upper surface is shelving on the sides, with an angular central pro-
minence.
The central aperture is very large, nearly circular, and dilated
above into an oblong transverse aperture, which is rather wider than
the widest part of the central circle. The front articulating surface is
horseshoe-shaped, continued to the upper outer angle, and obliquely
288 Zoological Society :—
shelving off on the upper edge to the base of the oblong part of the
aperture. The articulating surface of the hinder side is similar ;
but the articulating surface is shorter at the sides, and transversely
truncated in a line with the middle of the upper, oblong, transverse
opening (figs. 1, 2).
Fig.'2.
Fig. 1. Front of atlas of Catodon Krefftii.
2. Hinder side of ditto (reduced).
The second and other cervical vertebree are all united together into
one mass, anchylosed by their bodies, lateral processes, and neural
arches. ‘The neural arches form a triangular mass, which is strongly
Fig. 3.
Hinder view of cervical vertebre of Catodon Krefftii.
keeled on the central line; and the keel is stronger and produced
into an acute point at the hinder end (figs. 3, 4).
Dr. J. KE. Gray on a new Sperm Whale. 289
The lateral processes of the second, third, and fourth vertebre are
produced and united into a broad, thick, angular process, which is
expanded at the side, giving the united mass a rhombic appearance,
the width of the side being about one-fourth more than the height
of the mass.
Side view of the hinder side of the cervical vertebree of Catodon Kreffttt.
There is a tubercle, which is most probably the end of the lower
lateral process of one of the anterior cervical vertebrae, at the lower
part of the hinder side of the front lateral expansion.
The three hinder vertebrz have no distinct lower lateral processes ;
their place is only marked by three slight ridges on the lower edge
of the hinder side of the mass. The upper lateral processes of the
hinder cervical vertebree are small, slender, forming a strap-like sec-
tion, rather tapering towards and truncated at the tips on the side of
the apertures for the passage of the nerves for the neural canal.
The neural canal is rather large, oblong transverse, the height being
about two-thirds of the width ; it is rather larger and higher behind.
The hinder surface of the body of the last cervical vertebra is ob-
long transverse, about two-thirds of the height of its width at the
widest part; the lower edge is rounded and rather angularly pro-
duced in the centre, and the upper margin transverse, with a slight
central depression; the surface is concave, with a central, linear,
perpendicular, compressed line.
The cervical vertebrae in Catodontide are united into a single mass
by their bodies, the neural arch, and the lateral processes. The
lateral processes of the anterior vertebree are produced, and form a
thick, subconical, triangular prominence on the side of the mass.
The front side is nearly flat, and the lateral processes of the hinder
vertebree are shorter and shorter to the last. The hinder surface
Ann. & Mag. Nat. Hist. Ser. 3. Vol. xvi. 20
290 Zoological Society :—
shelves from before backwards, and is crested with some conical pro-
minences which indicate the lateral processes of the different ver-
tebree of which the mass is formed. The first dorsal vertebra is
sometimes partially anchylosed with the seventh cervical.
The arm-bones are very short.
NoTEs UPON THE CUCKOOS FOUND NEAR SypNEY, New SoutrH
Wates. By Epwarp P. Ramsay.
(1.) The Bronze Cuckoo (Chalcites lucidus) : Gould, B. Austr.
iv. pl. 89.
We have for many years been under the impression that the females
of this species lay two distinct varieties of eggs, which, although in
many instances exactly the same in size, differ widely in colour and
in style of marking.
The most satisfactory way of determining this question was to
procure specimens of each of these different eggs, and to place them
in nests of the Malurus cyaneus, or of various Acanthize (which
had been built sufficiently near our residence to admit of our occa-
sionally visiting them), until they were hatched, and then to com-
pare the young birds so hatched from each of the different eggs.
This we succeeded in doing in more instances than one, and found
that the young birds were in every case alike, and that when they
were sufficiently fledged we had no difficulty in recognizing them to
be the young of the Bronze Cuckoo (Chaleites lucidus).
The first variety of the eggs in question (var. A), usually recog-
nized as the egg of the Bronze Cuckoo, varies in colour from a uni-
form ashy grey to a rich dark olive-brown or bronze, many of the
light ashy-grey specimens having minute dots of deep olive towards
the larger end. In one specimen, in which these dots form a blotch,
they are more inclined to reddish brown.
Var. B has a purely white ground, blushed with pink before the
egg is emptied, and minutely freckled over the whole surface with
dots of light brownish red or dull salmon-colour, running in some
instances into blotches which stretch half across or round the surface,
leaving patches of the white ground without any markings. Occa-
sionally we find a specimen in which the salmon-colour and bronze
seem to be blended, forming a curious brownish-lilac tint.
Both varieties vary much in size: we have specimens of var. A
varying from 8 by 6 lines to 10 by 53 lines ; of var. B, from 8 by 5
lines and 84 by 6 lines to 95 by 6 lines in breadth. The colouring-
matter of both varieties easily rubs off, especially when the eggs are
freshly taken. The Bronze Cuckoo seems to give no preference to
any particular character of country, being found equally numerous
in all parts. In the thick shrubs and low brushwood it finds a secure
place tor depositing its eggs in the nests of Malurus Lamberti and
Acanthiza pusilla. In the half-cleared patches of land and even in
our gardens and shrubberies it seeks for the nests of the Malurus
cyaneus, Acanthiza lineata, A. reguloides, and A. nana.
From a nest of this last-mentioned species (4. nana) I remember
Mr. KE. P. Ramsay on Cuckoos of New South Wales. 291
taking, in the year 1855, no less than six eggs. Among them were
three of the Bronze Cuckoo—two of var. A and one of var. B. In
November last (1864) we took another nest of the same species, con-
taining one of each variety. In this instance one of the eggs, var. A,
was imbedded below the lining of the nest, and had evidently been
laid before the nest was completed, as is not unfrequently the case.
The other egg, which was a specimen of var. B, my brother Percy
placed in a nest of Acanthiza lineata, which he had found on the
previous day and left for such an occasion. On returning to it about
a week afterwards we found that the young Cuckoo had been hatched.
After the lapse of seven days the bronze feathers were just com-
mencing to appear, and in about a week or ten days more the young
bird was nearly able to fly, the bronze on the wings, head, and back
now showing plainly.
All the species of Acanthize that we have met with construct oval
dome-shaped nests, having the entrance near the top, and more or
less covered with a hood. The nests are either suspended (as in the
case of 4. lineata) from the end of some drooping or horizontal
bough, or, like those of the Maluri, placed in some low bush or
cluster of vines, or, as is often the case with 4. reguloides, placed in
the thick forks or loose hanging pieces of bark of the Hucalypti and
white-barked Tea-trees (Melaleuca).
Now, as the apertures of the nests of the dcanthize are exceed-
ingly small, a question naturally arises whether the Bronze Cuckoo
lays its eggs in the nest, or places them there by some other means.
To this I can only answer that the apertures of those nests which
have contained Cuckoos’ eggs are nearly twice as wide as the open-
ings of those nests which we have taken before the Cuckoo’s egg has
heen deposited in them. This is more easily noticed in the nest of
A. lineata, of which the aperture is very small, and neatly covered
over with a hood.
The following are a few extracts from my note-book, showing the
species which are most frequently the foster-parents of the Bronze
Cuckoo :—
|
Eggs |
Nest. ot Eggs of Cuckoo.
owner.
Sept. 29th, 1862 ...|Adcanthiza pusilla| 2 |i of C. lucidus, var. B.
Sept. ]1th, 1863* ... a | 2 {1 of C. lucidus, var. A.
Sept. 12th, 1864 ...|4. lineata ........ 3 {1 of C. lucidus, var. A.
Sept. 12th, 1864 ...|4. reguloides ...... 2 {1 of C. lucidus, var. B.
Sept. 14th, 1864 ...|Malurus cyaneus.| 3 |1 of C. lucidus, var. B.
Sept. 14th, 1864 ...|4. reguloides ...... 3 |1 of C. lweédus, var. A.
Nova S640 ei rccec- see PA MIUATEC Me ceaeeess 4 2 of C. lucidus: 1 of var. A and
| 1 of var. B.
Sept. 16th, 1864 .../J Meliphaga sericea 1 1 of C. lucidus, var. B.
Oct. 2nd, 1864 ...... Meliphaga sericea 1 {lof C. segs var. B.
Mr. Gould tells us that the Bronze Cuckoo is cianeledi over tae
* This nest also contained one of Cuculus cineraceus.
~
292 Zoological Society :—
whole continent of Australia, as well as New Zealand. In the latter
country I have myself met with it at every port I visited, from
Stewart’s Island to Auckland, where it arrives about September, and
leaves during February and March.
(2.) The Unavornep Cuckoo (Cuculus inornatus) : Gould, B.
Austr. iv. pl. 85.
When the eggs of two or more species of Cuckoo are found in the
same locality, and the birds themselves equally plentiful during the
same months, it becomes difficult to determine which is the egg of
each species, except perhaps where there is a great difference in the
size of the birds. Even this, however, must not be depended upon
in too great a degree, as will be seen in the present case. Following
the same plan as in the case of the Bronze Cuckoo (Chalcites lucidus),
we succeeded in procuring two young Cuckoos from eggs which we
had left in the nests of the Yellow-whiskered Honey-eater (Péilotis
auricomis). These, when fledged, we at once recognized to be the
young of Cuculus inornatus.
The young, upon leaving the nest, have the throat, face, and
shoulders black; the rest of the upper and under surface and tail
irregularly marked with dashes and stripes of black, scarcely two fea-
thers, even of wings, being alike. They retain this plumage until
March and April, during which months all the specimens I procured
were commencing to assume the more dusky plumage of the adult.
During these months all the old birds seem to have left us, the young
of the last season alone being found.
The present species arrives early in September, and is usually met
with in pairs, showing a preference for the half-cleared land and belts
of trees skirting the more cultivated parts. They may frequently be
seen perched upon the dead tops of trees, or among the lower open
branches, or often on the posts and fences, from which they pounce
down upon any unhappy grasshopper or cricket that they may have
discovered lurking in the grass.
Their food consists chiefly of Gryllide and Phasmide, various
species of Mantis, and often the beautiful larvee of the Cequosa
triangularis and Antherea eucalypti, which they obtain among the
leafy tops of the Hucalyptus trees. The crops in some specimens,
procured in October last, contained nothing but grasshoppers, which
appear to be their favourite food.
In this neighbourhood they usually deposit their eggs in the nests
of Ptilotis auricomis, but also occasionally in those of Ptilotis chry-
sops, but rarely in those of P#ilotis fusca and Melithreptus lunu-
latus ; in other districts, doubtless, in any nests suitable for the pur-
pose. I have frequently observed that whenever the eggs of Cuckoos
have been deposited in open nests, there is manifested a decided pre-
ference for those of birds which lay eggs similar to their own.
The Cuckoo’s eggs mentioned in my notes upon the Yellow-
whiskered Honey-eater (P. auricomis) in the ‘Ibis’ (vol. vi. 1864,
p- 245) as being found in the nest of that bird, I have now no
doubt belong to Cuculus inornatus, and not, as I then supposed from
their small size, to Cuculus cineraceus.
Mr. E. P. Ramsay on Cuckoos of New South Wales. 293
The eggs of the Brown Cuckoo (C. inornatus) closely resemble the
large and almost spotless variety of the Yellow-whiskered Honey-
eater ; they are, however, somewhat more rounded, and of a much
lighter tint, being of a pale flesh-colour, sprinkled with a few dots
of a deeper hue, but often without any markings at all. In length
they vary from 11 to 123 lines, being from 83 to 9 lines in breadth.
They are usually hatched about the twelfth or fourteenth day,
when the young Cuckoo, a little fat helpless creature, is scarcely
larger than its foster-brethren. However, as it grows more rapidly,
it soon fills up the greater part of the nest, and its unfortunate com-
panions, either smothered by its weight or starved to death through
its greediness, are thrown out by their parents.
On the 30th of October last (1864) we found two unhappy young
birds which had been hatched in company with a Cuckoo in a nest
of Ptilotis auricomis, tossed out and lying upon the ground just
under the nest; they were of course quite dead, and appeared to have
been about three or four days old.
During the months of October and November it is no uncommon
sight to see the smaller birds feeding the young Cuckoos; even the
little Acanthize, which I believe are never the foster-parents, at least
of this species (C. inornatus), join in supplying their wants, which
are easily made known by their continual peevish cry, stopping only
when being fed, or when their appetites are appeased.
While walking towards home, through a half-cleared paddock, on
the 27th of last October, I was not a little surprised, upon hearing
the cries of a young Cuckoo, to see a pair of adult birds of the same
species (C. inornatus) flying after it, settling beside it, and apparently
paying it great attention. Several times they flew away, but returned
to it again; and from their actions I feel convinced that they were
feeding it, although, much to my regret, I was unable to obtain a
view sufficiently close to make sure of the fact.
(3.) The Cinereous Cuckoo (Cuculus cineraceus): Gould, B.
Austr. iv. pl. 86.
This, the third and remaining Cuckoo which annually visits us,
arrives much earlier than either of the former species.
During May I have found it very plentiful, preferring the lonely
and more closely wooded parts, and the sandy scrub-lands studded
with aged Banksie (B. serrata) and widely branching Eucalyptz,
where the undergrowth consists of low, thick, scrubby Lambertie
(L. formosa), Acacias, and dwarf Banksias, &c. Such are the parts
of our neighbourhood frequented by this species for nearly a month
after its arrival. Their clear wailing cry is often heard from the
depths of the bush, giving quite a melancholy tone to the surround-
ing neighbourhood.
June comes, and they leave their lonely haunts for the more open
wooded parts. Here they may be seen, either singly or in pairs,
often frequenting the gardens and orchards, where, among the leafless
fruit-trees, their undulating flight and the peculiar cuckooish upward
jerk of their tails at once render them conspicuous, As spring
294 Zoological Sacrety :—
advances, their melancholy cry assumes a more cheerful tone, but is
less often heard, giving place to a quicker and more harsh note.
The shrill whistle of the Bronze Cuckoos (Chaleites lueidus) is
now more often heard, accompanied by the mellow notes of the
Brown Flycatcher (Micraca macroptera), singing on the topmost
bough of some neighbouring tree; and the twittering of the dcan-
thize as they sport among the leafy branches of the Lucalypti, cling-
ing to the ends of the twigs and leaves in every possible attitude, the
tremulous anxious piping of the Spine-bills (Acanthorhynchus tenut-
rostris), the varied inward note of the Silver-eye (Zosterops dor-
salis), with other species far too many to mention here, keep up a
merry chorus, and, tired of the winter fogs, welcome the bright
spring mornings.
As the birds pair off and the nesting-season commences, this Cuckoo
seems to be less plentiful. Either some of them leave us, or they
scatter over the bush so thinly that we do not observe their numbers.
If some do migrate at this time, still many remain to deposit their
eggs and to avail themselves of the nests of those species most suited
to become the foster-parents of their young, after which they com-
mence to leave us, and, with the exception of a few stragglers and
young, appear to have all departed before the end of December.
Among those species the nests of which are favoured by visits
from this “ parasite” is Acanthiza pusilla, from a nest of which, in
September 1863, we took no less than four eggs—two laid by the
rightful owner of the nest, the other two by Cuckoos. One of these
was a very fine specimen of var. B of Chalcites lucidus, the other
an egg of the present species—Cuculus cineraceus. The entrance
of this nest was greatly enlarged, being in width fully two inches ;
and the hood, which usually conceals the entrances (which are near
the top of the nest, and not generally wider than one inch across),
was pushed back to such an extent that the eggs were rendered
quite visible.
I have now before me ten nests of Acanthize and four of Maluri,
the former comprising Acanthiza lineata, A. nana, A. pusilla, and
what at present I believe to be that of A. regulotdes, the latter
Malurus cyaneus and M. Lamberti.
Now, having compared the greatly enlarged entrances of those
from which we have taken Cuckoos’ eggs with the entrances of those
which did not contain the egg of a Cuckoo, and which we took as
soon as the bird had laid its full number of eggs for a sitting, I
cannot but feel convinced more than ever that the eggs of these pa-
rasites are laid in the nests, and not deposited in any other manner.
The average width of the entrances of the nests of deanthiza lineata
which have not been visited by a Cuckoo is 1 inch, while those
which have contained Cuckoos’ eggs vary from 2 to 23 inches. In
addition to the nests of Acanthiza pusilla, we have known this
Cuckoo (C. cineraceus) deposit its eggs in the nests of 4. reguloides (?)
and Chthonicola minima. Wow great is the difference between the
Cuckoo’s eggs and those of this last bird (Chthonicola minima),
which are of a bright reddish chocolate !
Dr. P. L. Sclater on Animals from Madagascar. 295
The eggs of Cuculus cineraceus are from 10 to 103 lines in length,
by 7 to 74 in breadth. The ground-colour is a delicate white,
spotted and dotted with wood-brown, deep brewnish lilac, and fair
lilac dots, which appear beneath the surface.
Some specimens are faintly sprinkled all over, and the dots have
a washed-out appearance ; others are marked more strongly, and in
these the markings formed are in a distinct zone at the larger end,
which is sometimes broken by a batch of very deep-coloured dots.
I have seldom met with the eggs of this species in collections
(although sometimes I have seen those of Cuculus inornatus), whereas
the eggs of Chalcites lucidus are to be found in almost every col-
lection of eggs made in New South Wales. It is curious that one
variety of the egg of the Chalcites lucidus (var. A) should be so
different from the eggs of the species in the nests of which it is
placed, whereas both the other species here mentioned lay eggs very
similar to those of their foster-parents.
June 13, 1865.—Dr. J. E. Gray, F.R.S., in the Chair.
Repvort oN A COLLECTION OF ANIMALS FROM MADAGASCAR,
TRANSMITTED TO THE SOCIETY BY Mr. J. CALDWELL. By
P, L. Scuarer, M.A., Pa.D., F.RB:S.
Mr. J. Caldwell, of Port Louis, Mauritius, has recently transmitted
to me a small collection of animals in spirits, collected in Madagas-
car, in the vicinity of Antananarivo. The species represented in the
series are two Mammals, five Reptiles, and a Crayfish.
The Mammals, which have been kindly determined for me by my
friend Dr. W. Peters, are of the following species :—
1. Nyctinomus (Mormoprterus*) suGuLARIs, Peters, n. sp.
N. supra fuscus, pilis bast albis, subtus fusco-canus, alis nigris ;
capite depresso, rostro lato ; auriculis triangularibus, sejunctis ;
fovea jugulari magna.
The only specimen of this very interesting species is a male, dis-
tinguished from all other species by a deep transverse fossa imme-
diately before the manubrium sterni.
The head appears more flattened than in any other species, and
terminates with a broad flattened snout. The triangular large ears
are, compared with those of other species, rather thin, not united,
but separated by an interspace of 4 millim.
The fur is soft, of moderate length. The hair of the upper parts
is dark brown, at the base white; that of the underside greyish.
* Mormopterus, nov. subg. In the formula of the teeth (Gs + = + xs"
when younger a -- i + sa) it differs from Nyctinomus with 2 molars, and
approaches more to Molossus. The lips also are not so much plicated as in Nye-
tinomus. It is a species intermediate between Nyctinomus and Molossus, thus
showing another instance of the relationship of the fauna of Madagascar to the
American fauna.
296 Zoological Society :—
The skull is more flattened than in other species, and remarkable
for a strongly developed ante-orbital erista,
metre.
Motallengtlic. vos see ehta a wees 0-089
Length of, the head fk koe verve 0-021
Of the (ear MN fronts: cess «ina 0-014
Breadth of itheears = jac fae tees ence 0-012
engi of phn crus 2 tj ccs he eaypnint er 0-024
ORAOTEATM, tesa eaters Rr 0°037
—- Of thumb car. coins eee 00085
— GRSecond, fren sec tis ays none 0°0335
——w— of third finger .............. 0-067
——— of fourth finger ............ 0°0565
Of fifGhy fn sets itch sim oe ise Uo
————_— of thigh) : 4). .saete'sie avesete. alone a 0-013
——————=~ Of tibia 5 6c hisins eisai ee ee 0-0105
—— of foot with claws .......... 0-008
——— of tail .............. ecwiciciee (OLDS
— of free end of the tail ........ OOll1
2. Mus, sp.?
A very young, indeterminable specimen, with only two molars de-
veloped. Above brown, penicillated with black, with the bases of the
hairs blackish grey ; below white. In its colour and the length of
the ear, this species is allied to the South-African Fieldmice, as Mus
colonus, M. natalensis, &e.
The Reptiles, which Dr. Giinther has named for me, consist of
two Snakes (Dipsas colubrina and Herpetodryas Bernieri), a Cha-
meleon (Chameleon lateralis, Gray), several fine specimens of a
Lizard of the genus Gerrhosaurus (G. lineatus, Cocteau = Cicigna
ornata, Gray), and an example of another Lizard (Liolepisma Belli,
Gray). All these are species already known to the fauna of Mada-
gascar.
The Crayfish I have submitted to Mr. Spence Bate, as our lead-
ing authority on this branch of natural history. Mr. Spence Bate
pronounces it to be a new species of Astacus, which he proposes to
call after its discoverer, with the following characters :—
Astracus CALDWELLI, Spence Bate, sp. nov.
The eyes are planted on short peduncles. The first pair of an-
tennze have the third joint of the peduncle reaching to the extremity
of the rostrum. Both branches of the flagellum are slender; and
the primary branch, which is half as long again as the secondary,
is about half the length of the anterior division of the cephalon.
The second pair of antennz are about three times the length of the
first ; and the flagellum is minutely articulate, each articulus being,
in length, less than half its breadth, and at the basal extremity being
about half the breadth of the last joint of the peduncle. The squa-
Dr. P. L. Sclater on Animals from Madagascar. 297
migerous process of the third joint is rounded and thickened upon the
outside, straight, thin, and ciliated upon the inner, and obtuse at
the apex. The rostrum reaches to the extremity of the penultimate
joint of the peduncle of the external antennze, rounded at the ex-
tremity, dorsally concave, the margins fringed within and above
the actual edge with a rim of short, blunt denticles. The ocular
orbit is deeply excavate, and armed posteriorly near the centre by a
small denticle, and at the infero-lateral extremity by a short, sharp,
curved, and anteriorly directed strong tooth. The lateral walls of
the cephalon are thickly covered with numerous, subequally distant,
short, spinous protuberances, which gradually lessen in importance
towards the dorsal surface of the carapace, which is perfectly smooth,
except for the well-defined fissure that distinguishes the anterior
portion of the carapace from the posterior—the demarcation between
the antennal and mandibular somites. The first or large chelate
pair of pereiopoda are subequal in size, but differ in form from those
of every other species of the genus with which I am acquainted, and
resemble more in general aspect those of the genus Homarus. The
dactylos is curved inwards, and tipped with a sharp unguis; the
dactyloid process of the propodos is similarly formed, and meets the
dactylos only at or near the apex; the approximating edges, how-
ever, are armed with a few small and one large tubercle opposite to
corresponding ones. The inferior and external margin of the pro-
podos, from the extremity of the dactyloid process to the carpal ar-
ticulation, is convex, and longer than that of the intero-superior
margin of the propodos and dactylos together. The carpus is armed
with three blunt and one sharp anteriorly directed teeth upon the
inner edge, and two sharp strong teeth upon the under surface.
The meros is furnished with two rows of teeth, that converge toge-
ther towards the ischium upon the inner surface. The other pereio-
poda have little to attract attention. The second somite of the pleon
has a tuberculous ridge just above the lateral margin. The inner
scale of the posterior pair of pleopoda is furnished with a central
row of short, sharp teeth ; and the telson is armed with similar teeth,
of which there are a few in the median line and others in two late-
ral obsolete rows.
The specimen from which the description is taken is a male. Of
all the species of this genus, this form approximates the nearest to
its marine allies, in the appearance of the great chelate pereiopoda,
of any that we are acquainted with. The generally close resemblance
of the several species of this genus is certainly very remarkable,
when we take into consideration the vast geographical distribution
that it has—larger, perhaps, than that of any genus of Crustacea
that is not of marine habits. Species have been taken in the frozen
waters of North American rivers, in the hot latitudes of Chili, in
temperate Europe and Tasmania, and now from the African island
of Madagascar. We do not know of any having yet been recorded
from the inland rivers of that continent.
298
MISCELLANEOUS.
On Pristolepis marginatus, Jerd.
To the Editors of the Annals and Magazine of Natural History.
GENTLEMEN,—Some time since, Dr. Giinther founded the genus
Catopra on a freshwater fish from Siam ; and he has recently added
another species to the genus, from the west coast of India, under the
name of Catopra malabarica.
In 1849 or 1850 I described that very genus under the name of
Pristolepis, founding it on the identical species lately described by
him from the Malabar coast, which I named Pristolepis marginatus.
It is very possible that Dr. Giinther may not have seen my paper on
the freshwater fishes of Southern India, published in the ‘ Madras
Journal of Literature and Science’; but it is quite as likely that he
has seen and ignored it ; and I therefore beg to call his attention to
it, as well as that of other naturalists who may not be disposed to
treat so slightingly the labours of fellow-workers in natural science,
writing under every disadvantage in a foreign land, It is very pos-
sible that the generic name bestowed by me may have been pre-
viously applied, in which case Dr. Giinther’s name will stand. This
fish, I may remark, is found in rapid rivers im Malabar, and also in
the elevated region of the Toynaad, the waters of which flow into the
Cauvery, on the eastern coast of India. It frequents chiefly rapids ;
and I have taken it with bait. I have not seen it longer than
9 or 10 inches.
Some two or three years ago Dr. Ginther contributed a short
paper to the Zoological Society, remarking on the extension of several
marine genera of fishes to Nepal, apparently on the strength of cer-
tain specimens in Mr. Hodgson’s collection. I have not the paper
now by me to refer to; but among others were Therapon and, I
think, Scatophagus. I need hardly say that the extension of any of
those marine and estuary fishes to Nepal is perfectly mythical ; and
Tam sure that Mr. Hodgson himself would not countenance the idea
for one moment. He probably purchased the fishes at Calcutta.
I intended at the time I saw this paper to have sent you a note on
the subject, and indeed wrote out a short paper; but it was delayed
through some cause or other.
Tam, Gentlemen, yours obediently,
Srinuggur, Cashmere, T. C. JeRvDON,
Aug. 7, 1865. Surgeon-Major.
[In answer to the above statements, we have received the following
from Dr. Ginther.—Kp. Ann. Nar. Hist. ]
1. It is scarcely necessary to say that Mr. Jerdon is in error in
believing that the genus Catopra was founded by me; it was esta-
blished by Dr. Bleeker in 1851.
2. Mr. Jerdon’s paper on Indian Fishes is known to me; the
description of Pristolepis, however, bears so much the stamp of
being written “under every disadvantage in a foreign land,” that
neither I myself nor Dr. Bleeker were able to recognize Catopra in it.
Miscellaneous. 299
3. Since I wrote my paper on Mr. Hodgson’s collection of fishes,
I have ascertained that not only all the Indian species of Therapon
enter fresh waters freely, but that several are exclusively freshwater
fish.—A. GUNTHER.
On the Constitution of the Fruit in the Crucifere.
By M. E. Fournier.
When a horizontal section is made of a bilocular Cruciferous
fruit, especially of a young ovary after the amalgamation of the two
parts of the septum, the latter is seen to be bifurcate at each ex-
tremity, and to embrace in the angle produced by this bifurcation
the elongated column from which the two rows of ovules originate,
described by the author as the placenta. This arrangement pro-
duces a triangular canal extended longitudinally within each placenta,
the horizontal section of which forms a triangle, with its apex at the
point of bifurcation of the two lamellee of the septum, and its base
upon the placenta itself.
The placenta presents, passing inwards, the epidermis, a green
parenchyma, cortical fibres, ligneous fibres, and tracheeze. The epi-
dermis presents projections formed by the cuticle, which are very
common in the Cruciferee. The green parenchyma completely sur-
rounds the placenta in most genera. It is continuous on each side
with the subepidermal parenchyma of the valves, and more internally
with the double origin of the septum, which springs directly from it.
The cortical fibres exist only on the outer side of the placental
column. The woody fibres, which contain chlorophyll at an early
period, form around the trachez a ring which is thicker exteriorly
than interiorly. The trophosperms originate from the placenta,
sometimes within, sometimes outside of, the triangular canal ; in the
former case they perforate one of the lamelle of the septum, to
which they appear to be adnate.
The valves present a double epidermis, the outer one with longi-
tudinally elongate cells, the interior with transverse cells, arranged
in two or three series. Within the outer epidermis there is a paren-
chyma, in which vascular bundles ramify in various ways, according
to the genera and species; this is separated from the inner epidermis
by a remarkable undescribed fibrous layer. It is formed of very
thick fibres, of which the section presents several concentric lines,
and strongly refracts light. The form of the section is circular in
Lunaria biennis and Psychine, elliptical in Sisymbrium. These fibres,
when examined in the middle part of the horizontal section, form a
simple row in Lunaria and Sisymbrium, several rows with parallel
elements in Psychine, two rows with crossed elements in Frdigia
clypeata, Med., and several rows with alternately crossed elements
in Raphanus and Enarthrocarpus. Near the placentas they are
always approximated, in several rows, and form a thicker tissue than
in the middle of the valve. Analogous fibres are met with in many
fruits (Malus, Fraxinus, Nigella, Ervum); but they are never so
frequent in other families as in the Cruciferee. They are absent
from the walls of the ovary in the Resedacee and Capparidacee.
300 Miscellaneous.
Below, these fibres terminate in a point, without attaching’themselves
to any analogous organ; above they are continuous with a fibrous
sheath surrounding the stylar canal; laterally they are in contact
with the annular parenchyma which surrounds the placenta, and
becomes converted into suberous tissue at maturity, causing the
dehiscence of the fruit and the separation of the valves. In the
genus Cardamine, of which the dehiscence is different, the herbaceous
layer of the placenta is interrupted at the level of the line of attach-
ment of the valves.
In certain Cruciferze, which have a free funiculus and a spherical
indehiscent fruit, these fibres do not exist, and the horizontal section
of the fruit only shows tracheze ramifying in a parenchyma.
The anatomical structure of the septum has not yet been thoroughly
investigated. Its two lamellze present at first cells filled with green
matter, which in some rare cases is retained until the fruit is mature.
The form gradually acquired by these cells, the direction of their
elongation, and the thickness of the membranes formed by them
may furnish specific and even generic characters for the Cruciferee.
The Alyssinee may even be divided into two sections according to
the form of the septal network.
Fibres and vessels are frequently developed between the two
lamellee. Sometimes the cells of the septum acquire the character
of fibres upon the median line. In many cases there exists in the
middle of the septum a fibrous bundle, which encloses a dotted duct
in Sisymbrium tanacetifolium. In Matthiola, Maleolmia, and several
Sisymbria, of which the author makes a distinct group under the
name of Maleolmiastrum, there is between the two lamellz an actual
membrane formed of juxtaposed fibres, among which are some
tracheze.
In some genera, especially Farsetia (excl. Fibsgia, Dec.), the fibres
of the septum are pierced with holes, by which they communicate,
forming a very elegant network unconnected with any fibro-vascular
bundle.
In Psychine stylosa the septum, which is very transparent and
formed of polyhedric cells with delicate and inconspicuous walls, pre-
sents long branched tubes of very unequal diameter, with distinct
walls and greenish granular contents before the maturity of the fruit.
These tubes generally ascend nearly parallel to each other; but they
anastomose irregularly, so as to resemble a laticiferous network.
But it is in the triangular canal that these formations, sometimes
closely resembling certain varieties of laticiferous vessels, are especially
met with, containing, however, only granules of chlorophyll, starch,
and fatty matters. They consist of isolated ramose cells, or more
frequently of elongated lateral vessels emitting branches at right
angles to their direction. Among the various elements of this system
we may observe sometimes complete partitions communicating by
the ordinary punctures, sometimes walls perforated like sieves, some-
times open canals, probably produced by the disappearance of former
partitions.
These anatomical observations throw a new light on the constitu-
Miscellaneous. 301
tion of the fruit in the Cruciferee. Upon this point some writers
have put forward singular opinions, in consequence of the difficulty
originating from the position of the stigmata in this family. Now
all those who suppose that the ovules originate from the median
part of a carpellary leaf reduced to the placenta, or joined to its fellow
upon the median line of the valves, are refuted by the fact that the
placenta presents a perfectly peculiar structure. The opinion of
De Candolle, that the septum was formed by the reentering margins
of the carpels, is also invalidated, as the four lines of bilateral origin
of this ergan are situated upon the parenchymatous circumplacentary
ring, and “the structure of the septum is quite different from that of
the valves. The fruit is therefore to be regarded as formed of two
carpels alternate with the placentas, and of tao intervalvar placentas,
from which the septum issues on each side and by a double origin.—
Comptes Rendus, September 4, 1865, p. 404.
Male Generative Organs of Phalangixm.
To the Editors of the Annals of Natural History.
Chislehurst, Kent, 5th Sept. 1865.
GENTLEMEN,—The ‘Annals and Magazine of Natural History’ for
this month contains a translation of Dr. Krohn’s memoir on the Male
Generative Organs of Phalangium, in which he points out certain
mistakes made by Treviranus and Tulk, and explains the true rela-
tion and homologies of those organs.
I had, however, four years ago made the same observations, and
given a figure in all essentials identical with that of Dr. Krohn (‘On
the Generative Organs in the Annulosa,” ‘ Philosophical Transac-
tions,’ 1861, p. 612).
This memoir appears to have escaped the notice of Dr. Krohn.
I am, Gentlemen, your obedient Servant,
JoHNn LuBBOcK.
On the Mode in which the Long-eared Bat captures its Prey.
Botanic Gardens, Regent’s Park,
Sept. 14, 1865.
My pear Si1r,—I have lately noticed a curious way in which
the Long-eared Bat (Plecotus auritus) captures its prey; and al-
though it may be familiar to naturalists, I have not found it men-
tioned by authors.
The peculiar structure of Bats is well known. The highly deve-
loped membrane used as the flying-apparatus or wings is also ex-
tended from the hind legs to the tail, forming a large bag or net
(the interfemoral membrane), not unlike two segments of an um-
brella, the legs and tail being the ribs.
Having caught a lively male specimen of the common Long-
eared Bat, and placed the little fellow in a wire-gauze cage, and
inserted a few large flies, he was soon attracted by their buzz, and,
pricking up his ears (just as a donkey does), he pounced upon his
prey ; but instead of taking it directly into his mouth, he covered it
with his body, and beat it by aid of his arms, &c., into the bag or
302 Miscellaneous.
interfemoral membrane ; he then put his head under his body, with-
drew the fly from the bag, and devoured it at leisure. This appeared
always to be the modus operandi, more or less cleverly performed.
Several times, when the fly happened to be on the flat surface of the
ground, the capture appeared more difficult, and my little friend was
by his exertions thrown on his back ; the tail could then be seen
turned round, with its tip and the margin of the membrane pressed
against the stomach, forming a capital trap, holding the fly, the
captor remaining on his back till he had withdrawn the fly from
the bag.
I had no opportunity of observing the action when the Bat was in
full flight ; but if the insect was captured a few inches from the side
of the cage, the mode was the same. When flying, the interfemoral
membrane is not extended to a flat surface (and appears not capable
of being so stretched), but always preserves a more or less concave
form, highly calculated to serve the purposes of a skim-net to cap-
ture insects on the wing. Occasionally, when the Bat was sleepy,
sitting at the bottom of the cage, nodding his head, a poor silly
** Bluebottle Fly,” no doubt of tender age, and not read in the
natural history of the Vespertilionidee, with the greatest confidence
walked quietly under his friend, passing nose, ears, and eyes without
danger ; but immediately he touched the sensitive membrane of the
bag it was closed upon him, and there was no retreat except by
being helped out of the difficulty by the teeth of the Bat.
On looking through books at hand to see if the above was noticed,
I find that most accurate of observers of nature’s works, Gilbert
White, of Selborne, speaking of a tame Bat, says, “If you gave it
anything to eat, it brought its wings round before its mouth, hover-
ing and hiding its head in the manner of birds of prey when they
feed”’ (a capital description of the action of my little friend, only no
mention is made of the bag). Also, in Bell’s ‘ British Quadrupeds’
is the following :—‘‘ Of Bats, the interfemoral membrane is probably
intended to act as a sort of rudder in rapidly changing the course of
the animal in the pursuit of its insect food.” ‘ In a large group of
foreign Bats which feed on fruits or other vegetable substances, as
well as some of carnivorous habits, but whose prey is of a less active
character, this part is either wholly wanting or much circumscribed
in extent and power.”
May it not also be, that they do not require an entomological
bag-net ? Believe me yours truly,
To W. Francis, F.L.S. Wo. SowERBy.
On the Habits of the Water-Shrew (Crossopus fodiens).
By N. L. Austen, Esq.
I am induced to offer you the following account of the Water-
Shrew, as the animal in question, though tolerably abundant in many
localities, may not have come under the personal observation of some
of my hearers. I have also never seen it mentioned as having been
kept with success in confinement, and therefore will attempt to de-
scribe as accurately as possible the habits of a pair that lived in my
possession for a considerable time, hoping that the details may not
Miscellaneous. 303
prove altogether uninteresting. In form this Shrew closely resembles
the common species, the snout being lengthened in the same manner,
and the fur having the same velvety softness of texture. In size,
however, it is superior, a full-grown male measuring a little more
than 5 inches in total length, whereas the Field-Shrew rarely exceeds
4 inches; the feet and tail are fringed with stiff white hairs, which are
of great assistance to the creature when swimming. The colour on
the head and back is commonly a rich jetty black, on the sides and
underparts pure white; the line of demarcation between the two
colours very distinctly defined, adding much to the beauty of the fur ;
a small tuft of white hairs is also noticeable at the corner of the ear.
The Water-Shrew, as its name implies, is usually found in the
vicinity of pools and rivulets, where it forms in the banks long and
winding burrows, which penetrate for a considerable distance into
the loose soil, and end in a small chamber, furnished with a bed of
moss and dry grass. In this secluded retreat the young are produced
about the middle of May, there being usually from six to ten in the
litter. When first born, they are curious pinky-white little animals,
with round blunt noses and semitransparent bodies, bearing as little
resemblance as possible to their parents. A small colony of these
Shrews frequently inhabit the same spot, and towards the cool of the
evening may be observed searching for food, and sporting with each
other in the water, now hiding behind stones or large leaves to elude
their companions, and then darting out to engage in a general skir-
mishing chase, diving and swimming with the greatest activity, and
occasionally taking a plunge into their holes. By constantly traver-
sing the same ground, in going and returning from their burrows,
they gradually tread down a path among the grass and herbage, by
which their presence may readily be discovered by an experienced
eye. When under water their fur is covered with multitudes of tiny
air-bubbles, that shine like silver, and have a beautiful effect when
seen against the dark surface of the body. Spots where the stream
in some bend of its course forms a little pool are the favourite re-
sorts of this pretty little creature; and, although easily startled by
the slightest noise, their range of vision seems far from extensive, as,
by quietly approaching, I have often succeeded in watching their
gambols without causing alarm among the small community. The
food of the Water-Shrew includes insects, worms, young frogs, and
small fish, which latter it pursues and captures with all the graceful
dexterity of the Otter. I am enabled to speak with certainty as to
this fact, by observing the mode employed by my own pets in seizing
their prey. I obtained them in the following manner :—Having no-
ticed a very fine pair that frequented a small pond, | set several circular
wire mouse-traps, baited with small frogs, in what I supposed to be
their favourite runs, and secured both male and female by the next
morning. I had already had a cage constructed as much as possible
in accordance with what I knew of their mode of life. It was shaped
like an ordinary arched dormouse-cage, but considerably larger than
those used, being 12 inches in height by 18 in length; a zine tank
was also adapted to hook on to the doorway, so that they might
enjoy the comfort of a bath. When first introduced into their new
304. Miscellaneous.
dwelling, the Shrews evinced no symptoms of fear, appearing quite
at home, and feeding freely on worms, raw meat, and insects. A
few days after I procured them, I placed three or four minnows in
the bath attached to the main part of the cage. Directly the Shrews
caught sight of the fish, they both plunged instantly into the water,
anil quickly reappeared, each having secured a victim, which they
proceeded to discuss with great apparent gusto, having first killed
it by a bite through the head. I remarked that while feeding they
held the fish firmly between their fore paws, in the same manner as
the Otter, and, commencing at the head, ate gradually downwards,
by a succession of sharp snapping bites. Their appetites were very
good, as they frequently consumed two or three minnows each in
one day—a very tolerable amount, considering their size. When
running about their cage, these Shrews often uttered a shrill sibilant
chirp, resembling the note of the Grasshopper-Lark. They would
also play in the water, half rearing up and striking with their fore
paws, or rolling over and over each other on the surface. Though
appearing perfectly reconciled to captivity, they manifested no attach-
ment, nor especial tameness, biting viciously when touched. They
lived with me in this way several months in perfect health, till the
cage-door being accidentally left open one day in my absence, the
inmates levanted, as a matter of course, and were never seen or heard
of afterwards. I hope, however, shortly to obtain more, as when
treated properly, and supplied with plenty of water, they thrive, and
might probably be induced to breed in confinement. Besides the
Common Shrew, which is exclusively terrestrial, another species, the
Oared Shrew (Crossopus remifer), is found in Britain. For some
time this animal was confounded with the Water-Shrew, as its habits
are similar, and it frequents the same situations. It differs, how-
ever, in colour, the black on the back and sides being flecked with
white hairs, the throat and abdomen blackish grey tinged with yel-
low. Though scarcer than the two other kinds, the Oared Shrew
is more abundant than is often supposed by naturalists, as I have
several times caught it in different parts of Hertfordshire and Surrey.
I must here remark that the ears of both the Oared and Water-
Shrew are furnished with a peculiar and beautifully contrived appa-
ratus by which the water is excluded from those organs. It consists
of three small valves, which fold together when the animal dives,
effectually preventing the entrance of a single drop of moisture. As
soon, however, as the pressure is = ara, on the Shrew rising to
the surface, they reopen spontaneously. Without this provision of
nature, the animal would constantly be annoyed by the water filling
the cavities and irritating the delicate membranes of the ear.
The dimensions of full-grown individuals of the three species are
as follows :—
Common Shrew. in. lin. Water- Shrew. in. lin. Oared Shrew. in. lin.
Total length 4 1 Total length 5 7 Total length 6 1
lelervole San lr Head 5 Heady... 47
Pailkleenens gg Tale ser 8, UT] Iie ee 2 4
oT —
Hind foot.. 0 3 Hind foot.. 0 Etind! foot. 2.07
Proc. Zool. Soc. June 27, 1865.
THE ANNALS
MAGAZINE OF NATURAL HISTORY.
(THIRD SERIES. ]
No. 95. NOVEMBER 1865.
XXXIII.—On the Microscopic Structure of the Shell of Rhyncho-
nella Geinitziana. By Witxiam B. Carvenrer, M.D., F.RS,,
Beis.) E.G-S.
Ly consequence of my prolonged absence on the Continent, it
has been only within the last few days that [ have seen Prof.
King’s “ Remarks on the Histology of Rhynchopora Geinitziana,”
contained in the Ann. Nat. Hist. for last August (p. 124). These
remarks have led me to subject my preparations of that shell to
a renewed microscopic examination, of which I have now to
state the results. Before doing so, however, I may say that I
have done. my best to dismiss from my mind any prejudice in
favour of that view of its structure which I might be supposed
to derive from the conclusion to which I had been led by my
previous researches—that whilst the perforation of the shell by
canals passing from surface to surface is the family character of
the Terebratulide, the absence of such perforation is the family
character of the Rhynchonellide. The progress of natural-his-
tory inquiry is continually bringing to light examples in which
features essentially characterizing one group appear in particular
types belonging to another. Thus, a paper “On Rose-spored
Mushrooms,” by Mr. Berkeley, now lying before me, commences
as follows :—“ 1 have already pointed out that a single species
with decidedly rose-coloured spores (Agaricus euosmos) occurs in
the white-spored series; but its affinities with the common
Oyster-Mushroom (A. ostreatus) are so intimate that it would be
in direct opposition to nature to separate them.” It would not
in the least surprise me, therefore, to meet with a perforated
Rhynchonellid ; and IT can honestly say that no wish to make
out Rhynchonella Geinitziana imperforate is father to the belief
that, as regards its outer layer, it really is so.
The preparations in my possession consist (1) of transparent
lamellee, scaled off from the exposed surface of German and
Ann. & Mag. N. Hist. Ser.3. Vol. xvi. 21
306 = Dr. W. B. Carpenter on the Microscopic Structure
Russian specimens of the fossil shell in question, and therefore
passing, more or less exactly, in a direction parallel to that sur-
face ; and (2) of a transparent vertical section of a German speci-
men. As these specimens were supplied to me by Mr. Davidson,
there can be no doubt of their authenticity.
Some of the transparent lamellae exhibit distinct and regular
perforations, filled with black matter, of considerable size; and
had these lamellz been my sole materials of judgment, I should
have readily accorded with the description of Prof. King. But,
as I stated when the matter was formerly under discussion (Ann.
Nat. Hist. March 1857, p. 214), this appearance is presented
only by lamelle taken from abraded surfaces, and therefore be-
longing to the infernal layer of the shell. In a transparent
fragment in which the natural surface of the shell is partially
preserved, the large and regular black spots are seen only near
one edge; towards the middle they give place to small black
dots, so irregular in size and form as scarcely to be distinguish-
able from others which are obviously due to infiltrating deposit;
and towards the other side they are wanting altogether. Now
in the whole of this specimen the characteristic structure of
the shell is most perfectly preserved, so that the absence of
the marks of perforation cannot be ascribed to metamorphic
action.
The key to this variety in the appearances presented by pa-
rallel lamellze is afforded by the vertical section. In one part
(a) of this section there
is an obvious deficiency — _2
of the external layer of “—&
the shell, and the per-
forations are seen to pass ~
continuously through Vertical section of a portion of the shell of
the remaining internal Rhynchopora Geinitziana.
layer. But in another part (4c), the external layer is preserved
in great part, if not the whole, of its thickness; and this layer is
plainly seen not to be perforated at all, the passages all stopping
short of it, sometimes ending abruptly in rounded terminations
(4), sometimes more pointedly (cc’). Hence it is obvious that
if the plane of a parallel section pass along the line de, it will
show at a large perforations, at c small perforations, and at bc!
none at all, which is exactly what is seen in the specimen pre-
viously described. And further, as the transparence of the shell
allows the large black spots with which the inner layer is regu-
larly marked to be plainly seen through the outer layer, even
when this is perfectly preserved, it is easy to understand how
readily the conclusion might be drawn from incomplete obser-
vation, that the perforations extend through the whole thickness
of the Shell of Rhynchonella Geinitziana. 307
of the shell,—a conclusion which I have shown to be negatived by
the decisive test of a vertical section.
I feel myself justified, therefore, in reiterating my former state-
ment, that the passages which are visible in the shell of Rhyncho-
nella Geinitziana traverse the internal layer only, and are there-
fore of the nature of pits, having no physiological relationship
with the canals which traverse the whole thickness of the shell of
the typical Terebratulide, and which open out in large trumpet-
shaped orifices on its external surface, although presenting such
a rudimental approximation to that structure—as might almost
be expected in some member of the imperforate series.
The readers of the ‘ Annals’ have now both sides of the case
fully before them, and can form their own judgment whether it
is more likely that Prof. King or that I have fallen into a “ serious
mistake” in this matter. But I must ask them to bear in
mind that Prof. King’s observations upon this shell have been
made, by his own showing, only with a Stanhope lens, upon the
exposed surfaces of his specimens; whilst mine have been made
with a Binocular microscope and a magnifying-power of 120
diameters, upon transparent lamelle and sections. Further, I
would recall to their recollection that it was by surface-obser-
vation with the Stanhope lens that Prof. King was formerly led
to commit himself to the conclusion that all Brachiopod shells
are perforated; from which conclusion, if true, it would necessa-
rily follow that the elaborate drawings and descriptions which I
had given (in the Reports of the British Association for 1844),
of the microscopic structure of the non-perforated forms, had no
prototypes in nature*.
University of London, Burlington House, W.
October 16th, 1865.
* So far from having ever expressed his regret for this grave imputation,
of the fallacy of which he has had ample opportunity of convincing him-
self, Prof. King has recently pursued the very same course, in asserting that
Eozoon Canadense is not a fossil, but is a product of chemical and physical
agencies. For, if this be true, it necessarily follows, either (1) that my de-
scription of its Foraminiferal characters has no foundation in fact, or (2)
that I am incompetent to pronounce upon what I assert to be mdubitable
Foraminiferal structure. As he has not adduced one single fact to justify
either of these charges, I have felt myself called upon to repudiate an toto
his claim to authority in this matter. Whether, under such circumstances,
the charge of “ personality” is to be laid at my door or at Prof. King’s, |
leave it to others to decide. Although he may have used no hard words,
the imputations conveyed by his assertions would, if true, be more
damaging to my personal as well as to my scientific character than any
epithets he could employ.
2A
308 Mr. H. W. Bates on the Longicorn Coleoptera
XXXLV.— Contributions to an Insect Fauna of the Amazons Valley.
CotnorrerRa: Lonercornes. By H. W. Bares, Esq.
[Continued from p. 182. ]
Genus XYLOMIMUS, nov. gen.
Body cylindrical, narrow. Head vertical, or slightly inclined
backwards; muzzle moderately elongated ; sides rounded ; fore-
head very slightly convex ; eyes small, lower lobe nearly circular.
Antenne moderately distant at their bases, with inner side of
antenniferous tubercles prominent and angular; basal joint
dilated almost from the base, and forming a thick, oblong club,
with the lower edge slightly waved ; shard. joint one- thindl longer
than the first, and also thickened nearly from the base, continu-
ing of the same thickness to the apex, furnished on the under-
side with a fringe of long bristles ; fourth joint slender, slightly
thickened in the middle, and about one-half the length of the
third ; fifth to seventh joints each about one-half the length of the
fourth, slender (rest wanting). Thorax cylindrical, longer than
broad, and deeply wrinkled transversely ; lateral tubercles in-
conspicuous. Llytra linear, obtusely rounded at the apex, sur-
face free from excrescences ; pro- and meso-sterna plane. Legs
very short, thighs clavate, tibize broad; claw-joint of the tarsi
as long as the remaining joints taken together.
The species on which this genus is founded presents, from its
shape and style of coloration, a striking resemblance to a frag-
ment of a slender decayed branch.
Xylomimus baculus, n. sp.
X. angustatus, cylindricus, thorace transversim crebre ruguloso ;
elytris stria impressa suturali, apice singulatim obtuse rotundatis ;
corpore supra brunneo, lateribus obscure ochraceo; elytris pone
medium fascia lata flexuosa brunnea ochraceo lineata; antennis
brunneis, articulo quarto flavo. Long. 5j lin. ¢?
Head dingy ochraceous, front uneven, punctured ; vertex and
occiput ample, brown, streaked with rusty ochreous ; antenni-
ferous tubercles slightly prominent on their inner sides, and
leaving a small semicircular notch between them. Antenne
with the first and third joints dark brown varied with ochreous,
bristly, fringe of the third also dark brown, fourth joint yellow,
fifth, sixth, ad seventh rusty brown. Thorax cylindrical, sur-
face covered with numerous, irregular, transverse wrinkles ;
lateral tubercles small, conical, dark brown in the middle, with
three indistinct rusty-brown vitte ; sides each with an ochreous
vitta, below which is a broader brown vitta. Llytra linear,
shoulders not prominent, apex of each obtusely rounded; surface
slightly uneven, plane towards the base and more convex beyond
of the Amazons Valley. 309
the middle, punctured (except near the apex) and marked with
an impressed stria near the suture; colour rusty ochreous, with
a broad common brown vitta over the suture from the base to
beyond the middle, and a broad irregular brown fascia (lineated
with rusty brown) at the termination of the vitta, the space
near the apex having an irregular ochreous spot followed by a
similarly shaped brown. spot. Body beneath light brown; sides
of prothorax and breast with an ochreous-white vitta; abdomen
streaked with ochreous white. Legs clothed with pale tawny-
brown pile.
Found on a slender dead branch of a tree in the forests of the
Tapajos.
Genus Ectua@a, Pascoe.
Pascoe, Trans. Ent. Soc. n. s. iv. p. 244 (1858).
Syn. Talasius, Buquet, Thoms. Arcana Nature, p. 99 (1859).
This remarkable genus is distinguished from the allied groups
by many well-marked features, which have been well described
by the authors above quoted. I myself met with female exam-
ples only, and have not been able to examine the opposite sex,
which bears one of the chief marks of the genus—namely, four
horn-like projections from the forehead. The body is large and
cylindrical; the head very broad, and remarkable (besides the
horned forehead of the male) for the great convexity of the crown,
which rises very much higher than the base of the antenne, and
descends perpendicularly from its front edge towards the tuber-
cles which support those organs. The elytra are broad and
square at the apex, and each one is deeply sinuated in the mid-
dle, so as to form two projections or lobes. The antenne are
rather slender, in the female as long as the body, with the basal
joint tumid on one side at the apex, and the third joint slightly
curved.
My specimens differ in colour from the one figured by M.
Buquet ; but I believe them to be referable to the EL. quadri-
cornis of Olivier. The Trachysomus faunus of Erichson (Consp.
Peru. p. 148) seems to be quite a distinct species of this genus.
Ectheea quadricornis, Olivier.
Cerambyx quadricornis, Oliv. Ent. iv. p. 97, pl. 20. f. 158.
Talasius quadricornis, Buquet, Thoms. Are. Nat. p. 100, pl. 5. f. 6.
The female example now in my collection, and which I found
at Ega, is 94 lines in length, the head being 23 lines in width.
The upper part of the forehead is yellow, brown near the crown,
where it is marked with three black spots; the lower part is of
a blackish olive-colour, the line of demarcation between the two
colours being a transverse carina, from which in the male rise
the two lower frontal horns. The thorax is very uneven on each
310 Mr. H. W. Bates on the Longicorn Coleoptera
side, one of the elevations near the anterior part of the disk on
each side forming an acute tubercle; the colour above is rusty
ochreous, the hind part having two blackish lines, which are
severally continuous with the rounded velvety black spots on the
elytra, on each side of the scutellum. The elytra are of a light
green hue, except on the apical fourth, where there is a large
ashy-ochreous spot, streaked with dark brown, very similar to
the streaked apical spots in the genus Hudesmus. The under-
side of the prothorax and breast is greenish ashy. The legs are
green, varied with greenish ashy. These green and rusty-
ochreous hues, combined with the rugged surface of the insect,
give it very much the appearance of a mossy fragment of wood,
when it is seen clinging close to a dead bough, as is the habit
of the creature.
Genus Treston1a, Buquet.
Buquet, in Thomson’s ‘ Arcana Nature,’ p. 45.
like many other generic groups of Longicorns, the present
one is recognizable rather by a similar general form and colora-
tion than by definite structural characters. The species are
cylindrical or linear and depressed in shape, and exhibit a dark-
brown or black curved mark towards the apex of the elytra,
preceded by a pale-ashy or greenish patch, and succeeded by
fulvous strigee nearer the apex. The possession of this charac-
teristic mark points to a near relationship with Hudesmus and
Ficthea; but some species answer very well to the definition of
the genus Hesycha, as far as structure is concerned. All the
species, however, are more linear in form than the Hesyche, and
the antennee in nearly all are more nearly approximated at their
bases. The head is variable in width, and the forehead is some-
what convex in the middle; the latter is in most species clothed
with pale-coloured tomentum. The antenniferous tubercles, in
the broader-headed species, have prominent and sometimes
cornuted inner angles. The antenne themselves are slender
and setaceous, in the males often twice the length of the body ;
their basal joint is clavate, and the third joint, with few excep-
tions, a little curved. The thorax is cylindrical and uneven,
never short and broad. The elytra are linear, obtusely rounded
at the apex, free from centro-basal elevations and tubercles ; the
shoulders are prominent and acute, and curved anteriorly. The
legs are moderately short, the thighs clavate, the claw-joint ro-
bust, as is universal in the Onciderite, and equal in length to
the three remaining taken together.
The Trestonie, ike the other genera of the present group,
are found on branches of trees, clinging closely and gnawing the
bark and surface-wood.
of the Amazons Valley. 311
1. Trestonia Chevrolati.
Trestonia Chevrolatii, Buquet, Thoms. Are. Nat. p. 46.
T. elongata, subdepressa ; capite lato, tomento flavescente dense ves-
tito, maculis duabus verticis alterisque frontalibus nigris, genis et
gula nigricantibus ; antennis basi distantibus, brunneis, tuberculis
antenniferis intus modice productis acutis (Q); thorace obscure
fusco-grisescente, supra transverse ruguloso sulcis duobus trans-
yersis juxta marginem posticum distinctioribus ; elytris postice
paulo attenuatis, dorso depressis, humeris subconicis, granulatis,
disco bicostato lineaque elevata suturali, punctatis, griseis, ante
apicem utrinque plaga curvata nigra (antice albo marginata), dein
fulvo-brunneis macula subapicali pallida ; corpore subtus pedibus-
que viridi-griseis, abdomine ferrugineo-brunneo, segmentis tribus
posterioribus lateribus ochraceis. Long. 10 line 79%
One example, taken at Ega, and named as above, from the
typical specimen formerly belonging to M. Chevrolat. It would
be impossible to determine the species from the meagre descrip-
tion of M. Buquet.
2. Trestonia ramuli, n. sp.
T. elongato-oblonga, postice ( d ) angustata, subdepressa, fusca, fulvo
variegata ; elytris medio macula magna laterali viridi-cinerea
postice dentata et fusco marginata, intra apicem macula distine-
tiore fulva; tuberculis antenniferis distantibus, intus utroque sexo
prominulis acutis ; antennis corpore paulo longioribus, articulo
tertio curvato. Long. 6-63 lin. ¢ Q.
Head moderately broad, forehead punctured, dingy brown
varied with tawny; antenniferous tubercles with their inner
angles in both sexes prominent, acute, conical, and distant from
each other somewhat widely. Antennz very little longer than
the body in either sex, dark brown; joints paler at the base ;
third joint rather strongly bent in the middle. Thorax sub-
cylindrical, widest in the middle, convex, transversely depressed
near the hind margin, very uneven above, and obtusely tuber-
culose on the sides ; dark brown, varied with rusty tawny. Ely-
tra with prominent conical shoulders, and gradually narrowed
towards the apex (much less so in the female than in the male),
surface scarcely convex, simply punctured (except near the apex),
dark brown, minutely varied with rusty tawny, and having on
each side in the middle a large, oblique, greenish-ashy spot,
widest on the margin: this spot is bordered posteriorly by a
broadish, flexuous, blackish streak ; and close to the apex there
is a tawny spot, larger and clearer in colour than the other
tawny marks. Body beneath and legs clothed with olivaceous-
ashy tomentum.
On dead branches, Kga.
312 Mr. H. W. Bates on the Longicorn Coleoptera
3. Trestonia albilatera, Pascoe.
Hesycha albilatera, Pascoe, Trans. Ent. Soc. n. s. v. pt. 1. 25.
1’. elongato-oblonga, apicem versus paulo attenuata, subdepressa,
fusca, fulvo minute varia ; capite latiusculo, fronte ochracea, tuber-
culis antenniferis intus in lobulos erectos oblongos productis ( ¢) ;
elytris utrinque plaga maxima laterali (fere ad basin extensa) cana,
postice nigro marginata. Long. 63 lin. ¢.
Similar, in its elongate-oblong subdepressed form of body and
general colour, to 7. ramuli, but differs in the elytra being much
less prominent at the shoulders, and not attenuated, except from
very near the apex; the pale lateral spot, too, is much larger
and whiter, extending from behind the middle to the shoulders.
The thorax is cylindrical and very uneven on its surface, as in
T. ramuli; but it has two transverse impressed lines near the
hind margin, and a distinct conical lateral tubercle, much behind
the middle. The forehead is clothed with dense tomentum of a
pale ochreous hue. The underside of the body is ashy, with a
broad rusty-tawny stripe down the middle of the abdomen.
The antennz are very slender and twice the length of the body
in the male; the terminal joints are greatly elongated, and the
third with a scarcely perceptible bend.
Ega, on branches of trees.
4, Trestonia coarctata, n. sp.
Trestonia terminata, Buquet, Thoms. Are. Nat. p. 47, pl. 5. f. 3?
T. cylindrica, cinereo-fusca, fulvo varia, vertice coarctato ; antennis
basi valde approximatis, articulo basali elongato, apice abrupte
clavato; elytris crebre punctatis, apice nigris, fulvo lituratis.
Long. 43-6 lin. 3 Q.
The form of body and situation of the dark apical spot (close
to the apex of the elytra) in this species so closely resemble the
same features in the figure above quoted of 7. terminata, that it
is not unlikely the specimens here treated of belong to that
species. | cannot, however, reconcile the description of the
colours given by M. Buqnet with my insects; and the figure is
as uncertain in this respect as the description. His words are,
“Couleur générale d’un gris-verditre mélangé de blane et par-
fois de jaundtre sur le devant de la téte, sur les bords latéraux
du prothorax et sur la partie inférieure des élytres.” The head
in all my specimens is of a pale ashy hue, with a dark-brown
spot on the upper part of the forehead between the eyes. The
elytra as well as the thorax are dark brown, clothed with thinnish
ashy pile, and sometimes varied with tawny, and becoming of a
paler ashy hue near the dark apical spots. The thorax has a
number of large scattered punctures, and the elytra are thickly
of the Amazons Valley. 313
punctured, except at the extreme apex. The antennz are closely
approximated at the base, the bases of the tubercles being sepa-
rated only by the impressed line on the vertex; the angles of
the tubercles are not produced. ‘The antenne are more than
twice the length of the body in the male, the apical joint being
twice the length of the preceding, and of great tenuity ; in the
female they are but little longer than the body, but the apical
joints are very slender and more elongated than is usual in the
female sex of Longicorn insects; the basal joint is as long as
the third, and clavate at the apex.
Found, rather commonly, on slender branches on the banks
of the Tapajos, and also at Ega.
Genus PERITROX, nov. gen.
Body subcylindrical. Head moderately narrow; face plane,
inclined obliquely backwards ; eyes ample, convex ; antenniferous
tubercles with their inner angles produced. Antenne elongated,
simple; basal joint gradually thickened from the base; third
joint straight, one-fourth longer than the first, fringed beneath
with fine hairs. Thorax subcylindrical, uneven, sides armed
with prominent, acute lateral tubercles. Elytra cylindrical, free
from ridges and tubercles; apex rounded. Legs moderate ;
thighs clavate ; claw-joint of tarsi greatly elongated, longer than
the three remaining joints taken together.
This new genus, founded on one species only, is very closely
allied to Trestonia, differing, in structural characters, chiefly in
the gradually thickened basal joint of the antennz. The cha-
racteristic feature in the coloration of the elytra of Trestonia is
entirely absent, the colours beg dull and uniform. In form
of body and head, the species described below resembles much
Trestonia terminata and T. coarctata.
Peritrox denticolls, n. sp.
P. subcylindrica, paulo convexa, fuliginosa ; elytris maculis tomen-
tosis fulvo-brunneis adspersis; capite inter antennas profunde
impresso ; thorace transverse ruguloso, lateribus acute tubercu-
lntis: Wong. Slime 1%
Head sooty black, coarsely punctured on the forehead and
crown, and deeply grooved between the antenniferous tubercles,
which are closely approximated at their bases, and have their
inner edges produced into short ear-like lobes. Antenne
blackish, shining. Thorax subcylindrical, surface uneven, and
marked with a few sharp transverse wrinkles, besides two
impressed lines parallel to the hind margin; lateral tubercles
conical, acute; colour sooty brown. Elytra cylindrical, narrowed
314 Mr. H. W. Bates on the Longicorns of the Amazons Valley.
only very near the apex, the latter rounded; surface thickly
punctured, except near the apex, sooty brown, sprinkled with
spots formed of dingy-tawny tomentum. Body beneath and
legs pitchy, thinly clothed with ashy pile.
Santarem, on a dead branch: one example.
Genus Pacuypeza, Serville.
Serville, Ann. Soc. Ent. Fr. (1835) iv.
The forehead, muzzle, and eyes in this genus resemble much
the same features in Oncideres; but the crown is narrower and
more depressed between the antenniferous tubercles. The body
is elongate, but narrower than in any species of Oncideres. The
antenn have their joints beneath (including the basal joint)
clothed more or less densely with longish hairs. The thorax is
cylindrical, about as long as broad, and covered above with
transverse wrinkles. The pro- and meso-sterna are extremely
narrow. The legs are short and stout, the femora clavate, the
tibize very short and compressed, and the tarsi have the claw-
joint, although elongated, much less robust and shorter than in
Oncideres.
Pachypeza lanuginosa, n. sp.
P. cylindrica, robusta, fusco-cinerea; capite latiore ; antennis dis-
tantibus, articulis sex basalibus infra pilis tenuibus dense vestitis ;
elytris prope basin confertim et subtiliter granulato-punctatis.
Long. 93-10 lin. ¢ 2.
Head rather broad, forehead between the antenniferous tuber-
cles depressed; eyes large, oblong, ashy tawny. Antennz a
little longer than the body in the male, about the same length
in the female ; terminal joints shorter than the median ones,
last joint short and pointed; basal and five succeeding jomts
densely clothed beneath with very fine hairs ; colour ashy brown.
Thorax scarcely so long as broad, surface closely wrinkled, many
of the wrinkles not continuous; colour ashy brown. Elytra
cylindrical, convex ; shoulders somewhat prominent ; basal fourth
of the surface studded with small, regular granulations, accom-
panied by punctures; finely punctured in the rest of their sur-
face; colour ashy brown, deflexed sides paler. Body beneath
and legs tawny brown; base of abdomen on each side, and hind
legs, sooty brown.
Ega and S. Paulo, Upper Amazons, on slender woody stems.
Rey. H. Clark on Dejean’s Genus Coelomera. 315
XXXV.—An Examination of the Dejeanian Genus Coelomera (Co-
leoptera Phytophaga) and its Affinities. By the Rev. Haunt
Cuaxrk, M.A., F.L.S.
[Continued from p. 268. |
Genus Monocesta.
Division B.
Species of smaller size; in form more robust, short, and parallel ;
elytra not postmedially dilated; thorax with the transverse
depression more obsolete ; antenne shorter, more robust, and
slightly incrassated. Species 13-26,
Section I,
Elytra black, or fuscous black, or dark green. Species 13-19.
13. M. obliquenotata.
M. robusta, lata, satis parallela, rugosa, leviter pubescens, purpureo-
fusca, thorace humeris fascia obliqua et corpore subtus rufo-flavis;
caput leviter punctatum ; thoraa transverse depressus, lateribus
mediis angulatis, punctatus, rufo-flavus, macula utringue insu-
lata magna nigro-fusca ; scutellum subtriangulare, impunctatum,
flavum ; elytra robusta, subrugosa, purpureo-fusca, humeris et
fascia obliqua (a sutura ad margines postmedios tendente) atque
etiam sutura et apice (obsolete) rufo-flavis ; antenne fuscee, art. 1°
2° et 3° infra flavescentibus; corpus subtus rufo-flavum ; pedes
rufo-fusci.
Long. corp. lin. 6; lat. lin. 3.
Amazons. Collected by Mr. Bates.
14. MM. cincta.
M. lata, robusta, subparallela, subtilissime pubescens et rugosa,
nigro-fusca, thorace elytrorum fascia corpore subtus femoribusque
flavo-fuscis ; caput ad frontem foveolatum ; thoraw lateribus ad
medium angulatis, disco depresso, subtiliter punctatus, rufo- flavus,
disco medio nigro ; scute/lum subtriangulare, impunctatum, flavum ;
elytra robusta, subparallela, leviter rugosa, opaca, nigro-fusca,
fascia media recta marginibus et tenuiter sutura flavis; antenne
fuscee, art. 1° et 2° flavo-fuscis; pedes fusci, femoribus flavis ;
corpus subtus rufo-flavum.
Long. corp. lin. 6; lat. line.3:
Amazons. Collected by Mr. Bates.
15. M. spectanda.
M. robusta, parallela, subtilissime rugosa, rufo-flava, elytris (margini-
bus exceptis) viridescentibus ; ceput ad basin foveolatum, minute
punctatum, rufum, macula triangulari basali fusca ; thorax trans-
yerse depressus sparsim pubescens rufo-flavus, disco medio fusco
316 Rev. H. Clark on Dejean’s Genus Coelomera.
adumbrato ; seutellum rufo-flavum ; elytra rugosa, viridescentia,
marginibus flavis exceptis ; corpus subtus et pedes flava ; antenne,
art. 1° et 2° exceptis flavis, in exemplo unico desunt.
Long. corp. lin. 6; lat. lin. 4.
Cayenne. Received from the cabinet of M. Chevrolat.
16. M. flavo-cincta.
M. parallela, lata, subdepressa, rugosa, opace viridis, corpore subtus
thorace anteriore femoribus et elytrorum marginibus flavis ; caput
ad basin foveolatum, impunctatum, rufo-flavum, inter oculos super
antennarum margines nigrum; thorax vix ad discum medium
depressus, sed leevis, planus, subrugosus, rufo-flavus, basi et disco
medio nigris ; seutel/um subquadratum, impunctatum ; elytra lata,
rugosa, opace viridia, marginibus tenuiter rufo-fuscis ; antenne
fuscee, art. 1°-3" rufo-flavis ; pedes fusci, femoribus flavis ; corpus
subtus rufo-flavum.
Long. corp. lin. 43; lat. lin. 23.
Relatively broader, and with the thorax differently formed
from M. circumcincta. The two species are nearly allied.
Amazons. Taken by Mr. Bates.
17. M. cireumeincta, De}.
M. satis depressa, subparallela, rugosa, rufo- vel fusco-flava, an-
tennis nigris ; elytris nigris vel nigro-cyaneis ; capué longitudina-
liter ucaleuinte: flavum ; thorax impunctatus, fortiter transverse
depressus, flavus; scutellum fuscum ; elytra. thorace paulum
latiora et versus apicem subampliata, rugosa, nigro- cyanea, mar-
ginibus ab humeris ad apicem rufo- flavis; AY Ie nigree, art. 1°-3™
interdum subtus flavescentibus ; pedes et corpus subtus vel flava
vel fusco-flava.
Long. corp. lin. 33-5 ; lat. lin. 2-3.
Brazil.
18. M. carbonaria.
M. subparallela, paulum depressa, leviter rugosa, opaca, flava; ely-
tris antennis tibiis tarsisque nigris; caput rugosum, flavum, inter
oculos fuscescens ; thorax ad medium longitudinaliter foveolatus,
leviter punctatus, subpubescens; scutellum nigrum; elytra sub-
parallela, crebre rugosa, opace nigra; antenne filiformes, nigree ;
corpus subtus fusco-flavum, abdomine flavo ; pedes flavi, tibiis et
tarsis nigris.
Long. corp. lin. 4; lat. lin. 2.
Amazons. Taken by Mr. Bates.
19. M. nigriventris.
M. parallela, attenuata, rugosa, flava; antennis elytris abdomine et
tibiis nigris; caput longitudinaliter foveolatum ; thorax leviter
punctatus ; seutellum fascum : ; elytra parallela, rugosa, attenuata,
Rey. H. Clark on Dejean’s Genus Coelomera. 317
opace nigra; antenne filiformes, nigre; corpus subtus rufo-
fuscum, abdomine nigro; pedes rufo-flavi, tibiis et tarsis nigris.
Long. corp. lin. 3}; lat. lin. 14.
Brazil.
Section II.
Elytra flavous for the most part. Species 20-26,
20. M. Klugii, De}.
M. robusta, parallela, rugosa, opaca, subpubescens, fusco-flava, ma-
culis viridibus ornata; caput foveolatum ; thorax transverse de-
pressus, subrugosus ; scutellum rufo-flavum ; elytra robusta, ru-
gosa, fusco-flava, notis quatuor utrinque viridescentibus, 1* scutel-
lari magna, longitudinali, 2* marginali antemedia minuta, 3° mar-
ginali postmedia minuta, 4° suturali postmedia circulari ; pedes
et corpus subtus rufo-flava ; antenne robuste, fusce, art. 1° rufo.
Long. corp. lin. 5-6; lat. lin. 24-3.
Brazil.
21. M. rubiginosa.
M. robusta, parallela, crebre rugosa, rubiginosa, vittis duabus viridi-
nigris a basi ad apicem; caput rugosum, subpubescens, versus
apicem flavescens; thorax transversus, nigro-rubiginosus, lateribus
pallidioribus ; elytra thorace latiora, parallela, rugosa, rubiginosa,
vittis duabus (subsuturali et marginali) indistinctis indeterminatis
nigro-viridibus a basi ad apicem ; hee vittee aliquandofinterruptze vel
post medium omnino evanescunt ; antenne robuste, nigree ; pedes
et corpus subtus rufo-flava, tibiis aliquando nigro-fuscescentibus.
Long. corp. lin. 33-43; lat. lin. 2-23.
Brazil.
22. M. glauca, De).
M. robusta, subparallela, crebre punctata vel subrugosa, testaceo-
pubescens, vel flava vel rufo-flava vel fusca, antennis nigrescen-
tibus ; eapwé leviter rugosum ; thorax paulum transverse depressus,
rugosus ; scu¢ellum triangulare, apice truncato ; elytra thorace
latiora, leviter pubescentia ; antenne satis breves et versus apicem
incrassatee, fuscee, art. basalibus plerumque testaceo annulatis ;
pedes et corpus subtus rufo-fuscescentia.
Long. corp. lin. 43; lat. lin. 23.
New Granada; Bolivia.
23. M. frontals.
M. parallela, crebre punctata, flavo-fusca, capitis fronte nigra ; caput
fortiter longitudinaliter foveolatum, flavum ; thorax leviter punc-
tatus, flavus ; sewtellum fuscum ; elytra parallela, crebre et minute
punctata, subtiliter pubescentia, flavo-fusea ; antenne elongatule,
nigree, art. 1°-6" ab infra preesertim flavescentibus ; pedes et
corpus subtus flava.
Long. corp. lin. 4, lat. lin. 2.
Campeche ; Central America.
318 Rev. H. Clark on Dejean’s Genus Coelomera.
24. M. fuscescens.
M. parallela, crebre punctata, fuscescens, antennis et tibiis nigris ;
caput longitudinaliter foveolatum, leviter rugosum ; ¢horaw trans-
verse evidenter depressus, rugosus ; elytra subtiliter pubescentia ;
antenne nigree ; corpus subtus et pedes flavescentia.
Long. corp. lin. 43; lat. lin. 27.
Bolivia.
25. M. nigricornis.
M. robusta, parallela, crebre punctata vel rugosa, leete pallide rufa,
antennis et tarsis nigris; caput rugosum ; ¢horawx vix transverse
sed evidentius juxta latus utrinque depressus, crebre punctatus ;
scutellum subquadratum ; elytra thorace multum latiora, robusta ;
antenne robust, nigre ; corpus subtus rufo-flavum ; pedes rufo-
fusci, tibiis et tarsis nigris.
Long. corp. lin. 4; lat. lin. 23.
Bogota. Conspicuous by its bright pale-red coloration.
26. M. atricornis.
M. robusta, parallela, crebrius punctata, flava vel flavo-fusea ; caput
longitudinaliter foveolatum, punctatum ; thorax punctatus ; elytra
parallela; antenne robuste, paulum versus apicem incrassate,
nigree, art. 1° rufescente ; corpus subtus rufum; pedes rufi, tibiis et
tarsis interdum fusco-nigris.
Long. corp. lin. 33; lat. lin. 15.
This species may be easily separated from M. nigricornis by
its smaller size, its more depressed and more parallel form, and
its flavous (not rufous) colour.
Jacquelin du Val has described (in the ‘ Histoire de Vile de
Cuba,’ Insectes, p. 304) a “ Calomera” which Iam unable to re-
cognize: the description, unfortunately, omits all notice of the an-
tennee, and otherwise is somewhat imperfect. It is not a Dircema.
Probably it is a species of Monocesta, this genus being the only
one which extends northwards as far as the United States :—
“ Celomera opacipennis.
«C. oblonga, testacea ; elytris brunneis, opacis, pube subtili brevis-
sima sericeis, margine reflexo pectoreque medio nigro-cyaneis ;
thorace valde transverso, medio transversim impresso, basi sinuato,
angulis posticis acutiusculis. Long. 63 millim. Cuba.”
Genus XII. Catomera, Erichs.; Chev. Dej. Cat. ed. 3. 399;
D’Orbign. Dict. Univ. d’Hist. Natur. iv. 75; Erichson,
Conspectus Ins. Peruan. p. 164.
Generi Monocesta, ut hic definito, affinis; differt (ut a Dom.
Erichson indicatum est) in antennis ; art. 1° elongato, 2° brevi, 3° art.
srimo multum longiore, 4° primo eequali, 5°-11™ velut art. secundus
I g
Rev. H. Clark on Dejean’s Genus Coelomera. 319
brevibus: differt etiam in forma corporis; in speciebus plurimis
corpus parallelum, rarius post medium ampliatum : in reliquis genus
Monocestam equat.
The above diagnosis will suffice to point out the peculiarity of
the antenne, which is sufficiently important to constitute a very
natural genus: the third joint is more than usually produced,
and the fifth and subsequent joints are very short. There is also
a general difference in form: while one or two species [and those
from the head quarters of Monocesta, Cayenne and the Amazons],
C. Cayennensis, C. modesta, resemble Monocesta in the post-
medial dilatation of the elytra, the majority of species (which are
from Brazil) are decidedly more parallel, though robust im form ;
and several species from New Granada and Columbia are en-
tirely parallel and subattenuate.
Section I.
Species more or less broadly ovate ; elytra broadly margined.
Species 1-7.
1. C. modesta.
C. depressa, lata, punctata, flavo-fusca, prothorace et antennis flavis ;
caput ad basin longitudinaliter foveolatum, punctatum, rufo-
flavum ; thoraz satis latus, punctatus, rufo-flavus ; scutellum sub-
quadratum, punctatum ; elytra depressa, lata, versus apicem am-
pliata, leviter flavo-pubescentia, rugosa, flavo-fusca ; antenne rufo-
fuscee, art. 1°-3™ flavescentibus ; corpus subtus et pedes rufo-fusca.
Long. corp. lin 77; lat. lin. 5.
This species differs from MM. rufo-fusca in the greater breadth
of its thorax, its differently formed scutellum, its more depressed
form, and its coloration.
Amazons.
2. C. rufo-fusca.
C. lata, subpubescens, leviter rugosa, rufo-fusca; caput longitudina-
liter atque etiam transverse foveolatum, punctatum, rufum, fovea
longitudinali iterumque basi nigris; thorax vix ut in C. modesta
latus, sed transversus, punctatus, rufus, macula utrinque nigra
insulata; seutellum punctatum, rufo-fuscum ; elytra satis lata,
satis robusta, rugosa, subpubescentia, rufo-fusca; antenne art. 1°
—4™ nigris, reliqui desunt; pedes nigri, femoribus fusco-nigris ;
corpus subtus rufum.
Long. corp. lin. 63; lat. lin. 4.
C. rufo-fusca is certainly nearly related to the Amazonian
C. modesta. 1 decide, however, that it must be a distinct spe-
cies : its coloration is different (this of itself has no value in this
group); but, besides coloration, the thorax is relatively much
narrower.
Brazil. A single example from the collection of M. Laferté,
320 Rev. H. Clark on Dejean’s Genus Coelomera.
3. C. bajula, Oliv. Ent. vi. 618. 5, pl. 2. f. 17.
This is a not uncommon Cayenne species ; it is subject to a
considerable variation in the form of the thorax, the sides of
which are sometimes rounded, sometimes rectangular,—and espe-
cially in the form of the elytra; some examples in my cabinet
have the elytra much compressed. Length 63 lines, breadth
44 lines.
4. C. (Galleruca) lanio, Sahlb., Dalm.
C. derasa, Hoffmans. C. Braziliensis, De}. Cat. C. leta, Baly, Trans.
Ent. Soc. (1865) 344.
C. parallela, ¢elytris paulum post medium ampliatis, leviter rugosa,
elytris nigro-cyaneis; caput leviter foveolatum, rufum ; thorax
sparsim punctatus, vel rufus vel flavo-rufus; sewfe//umimpunctatum,
flavum ; e/ytra leviter rugosa, sparsim pubescentia, vel nigra vel
nigro-cyanea; antenne nigree; pedes fusci, femoribus rufis;
corpus subtus rufam.
Long. corp. lin. 54-64; lat. lin. 23-33.
This species is nearly allied to C. Cayennensis, Fab., but is
readily separated from it by its larger size and by the black
basal joints of the antenne.
Brazil. A very common species.
5. C. Cayennensis, Fab. Syst. El. 1.480. 11; Ent. Syst. u. 14.
Oliv. Ent. vi, 617.
C. Cajennensis, Fab. Mant. i. 74. 93, and in S. Nat. Gmel. i. 4. 1669. 85.
Differs from C. lanio of this paper in its smaller size, in the
rugose, almost reticulated (not quite so much punctate) surface
of the elytra, and in the black coloration of the antenne and
underside. One or two examples have the antenne rufo-
flavous.
The species has a very extended range: I have examples not
only from Cayenne, but from Peru, Columbia, and Brazil. It
is probably the C. Columbica of Schénh. C. Peruana, Kvichs.
(Consp. Ins. Peruan. 165) must be referred to it.
6. C.rujicollis, Oliv. Ent. 6.616; Encyel. Ins. 6. 586 (1790).
C. ruficornis, Baly, Trans. Ent. Soe. (1865), 343.
Very nearly related to C. Cayennensis, Fab., of which indeed it
may be but a local variety. Both C. ruficollis, Oliv., and C.
Cayennensis, Fab., are subject to some variation, the species
before us being distinguished by its rufous antennze and some-
what more brightly rufous thorax.
Brazil ; Campos, Espiritu Santo. I have examples also from
Peru, which are identical, with the exception of the colour of
the seutellum.
Rey. H. Clark on Dejean’s Genus Coeelomera. 321
Olivier gives C. nigripennis (Fab. Ent. Syst. Em. 2. 14. 9,
published 1775; Syst. El. i. 480) as a synonym of this species.
C. nigripennis, F., is from Surinam. I cannot quite trace the
reasons of Olivier’s position, but prefer to retain his name with-
out hesitation, as being well established, as well as based on his
excellent authority.
7. C. picta, Baly, Descript. of New, &c., Trans. Ent. Soe.
Lond. 1865, 344.
Length 5-4 lin. ; lat. 24-24 ln.
A common species at Ega, Upper Amazons.
Section II.
Species robust, parallel, abbreviated in form. Species 8-10.
8. C. induta.
C. robusta, satis parallela, pallide et subtiliter pubescens, testaceo-
fusca, thorace medio macula utrinque elytris antemediis et apice
elytrorum late fusco-nigris ; caput subrugosum, obscure testa-
ceum, basi nigra; thorax satis magnus, transversus, lateribus
antemediis subdilatatis, subpubescens, macula media nigro-fusca ;
scutellum rufo-fuscum ; elytra robusta, subrugosa, pubescentia,
testaceo-fusca, macula basali indeterminata insulata (alteraque
minore ad antemedium marginem) et dimidio apicali nigris;
antenne nigre, art. 1°-3™ fulvescentibus ; pedes rufo-fulvi, tibiis
apicalibus et tarsis fulvis ; corpus subtus flavo-rufum.
Long. corp. lin. 53; lat. lin. 3.
Amazons. Taken by Mr. Bates. |
9. C. tibialis, De}.
C. robusta, paulum ampliata, flavo-fusca, tibiis tarsis et antennis
nigro-fuscis ; differt a C. dajula Oliv. in corpore robusto, parum
depresso, vix ampliato, in antennis gracilioribus, fuscis (vix nigris),
in colore quoque corporis subtus, prothoracis et femorum.
Long. corp. lin. 6; lat. lin. 3.
Probably a distinct species from C. bajula, which is very va-
riablen form and size; it may be separated as well by its colour
(rufo-flavous prothorax and underside) as by the greater attenua-
tion of the antenne. .
Cayenne.
10. C. maculicollis.
C. parallela, robusta, subpubescens, fusco-rufa, tibiis, tarsis, antennis,
fronte capitis et maculis thoracis tribus nigris ; caput longitudina-
liter foveolatum, subpunctatum, ad frontem bimaculatum ; thorax
transverse leviter depressus, fortiter punctatus, nigro maculatus,
macula basali media et macula laterali insulatis; sewtellum fusco-
rufum ; elytra thorace latiora, parallela, apice rotundata, subtiliter
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. 22
2 Rev. H. Clark on Dejean’s Genus Coelomera.
pubescentia, crebre punctata ; anfenne robusta, nigra ; corpus
subtus rufo-flavum ; pedes rufo-flavi, tibiis et tarsis nigro-fuscis.
Long. corp. lin. 5; lat. lin. 25.
I have received this very distinct species from M. Chevrolat’s
collection, with a label which I read “ Honduras.”
ll. C. erata.
C. parallela, satis attenuata, punctata, impubescens, nigra, elytris
eeratis ; caput haud foveolatum, punctatum; thorax vix elytra
latitudine sequans, impunctatus, fusco-flavus ; sewtel/um apice ro-
tundatum, levissime punctatum ; e/yéra parallelo-punctata, leevia,
nitida, impubescentia, wrata; wntenne nigre, corpus subtus ni-
grum, abdomine rufo ; pedes nigri, femoribus rufis.
Long. corp. lin, 63 ; lat. lin. 3.
Columbia.
Section Ill.
Species parallel, attenuate in form. Sp. 11-16.
12. C. submetallica.
C. subparallela, leviter pubescens, punctata, rufa, thorace flavo-
rufo, elytris metallice rufo-purpureis ; caput leviter foveolatum,
impunctatum, rufum, basi nigra; thorax thorace C. arate latior,
in medio transverse depressus, impunctatus, flavo-rufus ; sewtellum
leviter punctatum, flavum; e/ytra crebre punctata vel rugosa,
metallice purpurea; antenne nigre ; corpus subtus rufum ; pedes
nigro-fusci, femoribus rufo-flavis.
Long. corp. lin. 63; lat. lin. 33.
Broader both in body and in relative breadth of thorax than
C. erata; the coloration also of the whole body is different.
Columbia.
13. C. parallela.
C. elongata, parallela, leviter rugosa, subtiliter testaceo pubescens,
nigra, thorace et abdomine rutis; caput leviter ad medium foveo-
latum, punctatum ; ¢horaz impunctatus ; scwfel/um impunctatum,
nigrum ; e/ytra parallela, testaceo pubescentia, rugosa ; antenne
nigree ; pedes nigri, femoribus rufo-fuscis ; corpus subtus nigrum,
abdomine rufo.
Long. corp. lin. 63; lat. lin. 63.
C. parallela differs abundantly in form from most of its con-
geners, being attenuate and parallel; it resembles C. violacer-
pennis, but may be separated from it by its fuscous-black, thickly
pubescent, and rugose elytra.
New Granada.
14. C. violaceipennis.
C. elongata, parallela, subcylindrica, vix pubescens, punctata, nigra,
abdomine pedibus et thorace rufis, eclytris nigro-violaceis ; caput
longitudinaliter foveolatum, impunctatum, nigro-fuscum ; thorax
Rev. H. Clark on Dejean’s Genus Coelomera. 323
transversus, lateribus antemediis subrotundatis, in medio trans-
verse valde depressus, impunctatus, flavus ; scwfellum impunctatum,
rufo-fuscum ; elyéra parallela, impubescentia, punctata, nigro-
violacea ; antenne rufo-fusce ; pedes ruto-fusci, femoribus rufis ;
corpus ioe nigrum, podoaine rufo.
- Long. corp. lin. 7; lat. lin. 3.
Nearly allied to C. parallela of this paper, but separable from
it by the blue (not fuscous-black) colour of the elytra, which are
impubescent, nearly glabrous, and distinctly punctate, not
rugose.
Columbia.
15. C. tenuicornis.
C. elongata, subparallela, subpubescens, nigra, femoribus et thorace
rufis 5 caput longitudinaliter ad medium et transverse inter
oculos foveolatum, punctatum, nigrum; ¢horaxv penitus planus,
vix ut in C. parallela fortiter transverse depressus, rufus; scu-
tellum leviter punctatum, nigrum; elytra subparallela, elongata,
rugosa, nigra; antenne subgraciles, nigre ; pedes nigri, femoribus
rufo-fuscis ; corpus subtus nigrum.
Long. corp. lin. 53-63 ; lat. lin. 23-34
Very nearly allied to C. parallela, but differing from it in its
less parallel form, the transverse fovea on thea head, the less
marked transverse depression of the thorax, and the ‘colour of
the underside; the antennee, also, are evidently somewhat more
attenuate.
Bolivia.
16. C. binotata, De}.
C. parallela, subrugosa, rufo-flava, elytris et maculis duabus tho-
racis nigro-cyaneis ; caput leviter punctatum, foveolatum, nigrum;
thorax sparsim punctatus, flavo-rufus, macula utrinque circulari
submedia nigra; scuéellum impunctatum, flavum; e/ytra leviter
rugosa, subpubescentia, nigro-cyanea; antenne nigre; corpus
subtus rufo-fuscum ; pedes nigro-fusci, femoribus rufo-flavescen-
tibus.
Long. corp. lin. 6; lat. lin. 3.
To be distinguished from all its congeners by the two con-
stant black maculations on the thorax; in size not unlike .
Brasiliensis, but differing in coloration, a trifle more parallel
and narrower, and with the thorax not quite so obviously de-
pressed.
Brazil.
Genus XIII. Corata.
Satis elongata, parallela et robusta. Cuput verticale. Palpi mazxil-
lares elongati. Thorax subquadratus, penitus transversus, margine
anteriore recto, angulis anticis distinctis (incurvatis, corpus arcte
amplectentibus), lateribus atque etiam angulis posticis subrotundatis ;
22%
324 Rey. H. Clark on Dejean’s Genus Colomera.
discus plus minus transverse et ineequaliter depressus. Scutel/um
subtriangulare, apice truncato. lytra parallela, pubescentia, punc-
tata, paulum latitudine thoracem superantia. dntenne robustee,
elongatee, filiformes, longitudine corpus penitus eequantes, art. 1° ad
apicem distincte incrassato, art. 3° et 4° subsequalibus et art. primo
brevioribus, art. 2° brevi, art. 5°-10™ eequalibus et penitus art. ter-
tium eequantibus, art. 11° longiore et attenuato. Pedes elongati,
robusti, tarsorum art. 1° et 2° eequalibus, unguiculis utrinque bifidis.
Coraia is a form which, by its almost quadrate thorax, its
long and robust antennee, and its parallel body, reminds us of a
Longicorn. It is allied to Monocesta, but differs from all the
species composing that genus: it is elongate and very parallel,
instead of being (as in many species of Monocesta) broad and
apically dilated; the antenne are relatively considerably longer,
and the form of the thorax is different. The genus Coraia is
related to Celomera and Monocesta by the bipectinated unguiculi
of the foot, by the relative lengths of the joimts of the antenne,
and by the broadly transverse depression on: the disk of the
thorax. Coraia differs from G'alleruca (inter alia) by the greater
relative length of the antenne. It may be separated from Nes-
tinus by its much more robust antenne, the third joint of which
is equal to the fourth (in Nestinus the fourth being the longer),
in its parallel form of body, and in the form of the thorax; the
anterior angles closely embrace the sides, bemg depressed in
position, not subporrect.
C. maculicollis.
C. elongata, parallela, vix depressa, rufo-fusca, humeris, thoracis ma-
culis, tibiis, tarsis, femoribus maculis antennisque nigro-fuscis ;
caput super antennarum bases bituberculatum, rugosum, rufo-
flavum, labro tuberculis et basi late nigris; thorax rufus, macula
media longitudinali et lateribus mediis (late) nigris; scutellum
punctatum, rufo-fuscum ; elytra thorace paulum latiora, parallela,
rugosa, humeris et margine tenui elytrorum antemedio nigris ;
antenne nigre ; corpus subtus rufo-fuscum ; pedes nigri, femori-
bus rufis, ad medium nigro maculatis.
Long. corp. lin. 53; lat. lin. 23.
Not an uncommon species in Mexico. I have received it
under the name of astuta, Chevr.
Genus XIV. NeEstTINUs.
Parallela. Caput subverticale. Thorax transversus, margine an-
teriore subemarginato, angulis anticis modice productis, lateribus at-
que etiam angulis posticis subrotundatis, marginibus undique leviter
marginatis, disco plano vel leviter depresso. Scutellum subtri-
angulare, apice rotundato. Hlytra thorace paulum latiora, crebre
punctata, plerumque submetallescentia. Antenne graciles, elongate,
Rev. H. Clark on Dejean’s Genus Ceelomera. 525
filiformes, corpus longitudine penitus sequantes, art. 1° subgloboso,
2° minore, 3° primum equante, 4° et 5° tertium superantibus et sub-
zequalibus, 6°-11™ subzequalibus. Pedes elongati, tarsorum art.
basali longitudine art. 2" superante, unguiculis utringque bifidis.
Nestinus approaches Coraia in the length of the antennz and
the relative length of the basal joint of the tarsus ; it differs from
that genus in the slender form of the antenne, in the fourth
joint being longer than the third, and also in the form of the
thorax. The known species that represent the genus are three
or four in number, and all from Mexico.
1. N. bimaculatus.
N. parallelus, rugosus, fusco-flavus; thorace et elytrorum apicibus
nigro maculatis ; caput ad basin leviter foveolatum, punctatum,
fusco-flavum, fovea tenuiter nigra; thorax crebre punctatus,
macula media insulata, alteraque etiam utrinque laterali nigris ;
scutellum impunctatum, nigro-fuscum ; e/ytra fortiter et crebre
punctata, fusco-flava, macula utrinque apicali circulari insulata
nigro-cyanea; corpus subtus, pedes et antenne flavo-fusca.
Long. corp. lin. 64; lat. lin. 24.
Guatemala.
2. N. regalis.
N. elongatus, subparallelus, fortiter punctatus vel rugosus, pallide
flavus, elytris vel zeneo vel aureo metallescentibus, antennis et
tibiis nigris; caput leviter foveolatum, punctatum, pallide flavum,
macula basali media nigra; ¢horax punctatus, pallide flavus, ma-
cula media minuta, et altera utrinque media laterali nigris; sev-
tellum impunctatum, rufo-flavum ; e/y¢ra fortiter et crebre punc-
tata vel eneo- vel aureo-metallescentia; antenne nigree; pedes
nigri, femoribus flavo-fuscis ; corpus subéus flavo-fuscum.
Long. corp. lin. 43-53; lat. lin. 2-23.
Mexico. I possess an example from California, which pre-
sents no variation from the Mexican form.
3. N. incertus.
N. subparallelus, punctatus, laevis, flavus; elytris fusco-viridescenti-
bus, thorace fusco-flavo, antennis fuscis ; caput crebre punctatum,
flavum, basi late nigra; thorax transversus, latus, subdepressus,
planus, lzevis, angulis posticis rotundatis, crebre punctatus, fusco-
flavus ; scwtellum leviter punctatum, flavum; e/ytra_parallela,
leevia, crebre punctata ; antenne robuste, nigree ; pedes rufo-flavi,
tarsis fuscis ; corpus subtus rufo-flavum.
Long. corp. lin. 6; lat. lin. 3.
A somewhat aberrant form of Nestinus, but which agrees in
the main with the generic diagnosis. . It is possible that here-
after it may constitute a separate genus.
Brazil.
326 Rey. 8S. Haughton on the Muscular Anatomy
XXXVI.—On the Muscular Anatomy of the Leg of the Crocodile.
By the Rev. Samuen Haveuron, M.D., Fellow of Trinity
College, Dublin.
[Plate XVI.]
Durine the Easter recess of 1864, I had an opportunity of
explaining to Professor Gratiolet*, of Paris, the investigations
I had made with respect to the mechanism of the leg of the
Ostrich, and the theory I had formed to explain it. This dis-
tinguished anatomist did me the honour of approving of my
explanation, and urged me to procure a Crocodile, mm the poste-
rior limb of which he assured me I should find a mechanical
problem exceeding in complexity that presented by the leg of
the Ostrich, and as yet unsolved by anatomists.
During the month of March last I was furnished with a
young Crocodile from Egypt, by Mr. Thomas Moore, of Liver-
pool, to whom I had communicated my earnest desire to have
an opportunity of dissecting such an animal ; and the results of
my examination fully bear out the anticipation of Prof. Gratiolet,
and also furnish a complete confirmation of the principles I
made use of in my theory of the leg of the Ostrich.
The interlacing of tendons in the hind leg of the Crocodile
is very remarkable, and more complex than in the Ostrich,
although in one respect it somewhat resembles it.
* The incaleulable loss that science has sustained, in the early part of
the present year, by the premature death of this distinguished anatomist
is exceeded by the loss experienced by his friends, to whom his genial
social qualities endeared him even more than his brilliant scientific attain-
ments. 1 extract from the ‘ Journal des Débats’ of the 19th of February
1865 the following just tribute to his memory :—
‘“‘Les sciences viennent de faire une perte aussi cruelle qu’imprévue :
M. Gratiolet, professeur de zoologie a la Faculté des Sciences de Paris, a
suecombé hier matin a une attaque d’apoplexie.
“M. Gratiolet n’avait pas cinquante ans; avant hier, encore plein de vie
et de santé, il travaillait & son laboratoire du Muséum d’Histoire Natu-
relle, lorsque, 4 deux heures, frappé d’une congestion subite, il dut étre
ramené & son domicile; quelques heures plus tard, il avait perdu connais-
sance; hier matin a quatre heures, il rendait le dernier soupir.
“* Nous ne saurions peindre l’émotion profonde qu’a causée dans le
monde scientifique annonce de cette mort prématurée. M. Gratiolet
était aimé de tous; son affabilité, la droiture de son caractére, lui avaient
concilié toutes les sympathies. Ses travaux d’anatomie comparée, ses re-
cherches sur le syste¢me nerveux et sur le cervean, ete., Vavaient mis au
nombre des naturalistes les plus distingués de notre pays; son merveilleux
talent d’élocution l’avait placé au premier rang parmi nos professeurs les
plus renommés, et aptitude de son esprit pour les études métaphysiques
avait imprimé A ses ceuvres un caractére d’originalité quappréciaient les
philosophes aussi bien que les savans.
“La mort est venue le frapper au moment ot, aprés de longues années
de lutte, il semblait sur le point de recueillir le fruit de ses laborieux
efforts.”
of the Leg of the Crocodile. 327
On removing the skin and dissecting away the fat, the mus-
cles shown in Pl]. XVI. fig. A are exposed.
1. M. gluteus maximus (0). Origm: from central half of
the ilio-ischiadic line. Insertion: into the fascia outside and
above the knee-joint. This is a broad flat muscle, and straps
down the tendon of the rectus femoris in its passage over the
knee. 0°15 oz.
2. M. rectus femoris (a). Origin: from anterior spine of
ilium, close to the acetabulum. Insertion: as in leg of Ostrich,
into a tendon passing over the knee, outwards, and terminating
in a remarkable muscle* (z) in the calf of the leg, associated
with the gastrocnemius (w), and deriving a second origin from
the agitator caudze (c), as shown in the figure. 0°08 oz.
3. M. agitator caude (c). Origin: from the ischiadie line,
behind the gluteus maximus. Insertion: by a double tendon.
(1) One tendon passes through a pulley on the outside of the
knee, formed by the tendon of the biceps (¢) as it passes to its
fibular insertion, and is inserted in the head of the muscle («)
in the calf of the leg. (2) The second insertion is by means
of a tendon that goes to the top and front of the tibia; this
second tendon also serves to strap down the tendon of the rectus
femoris (a). 0°03 oz.
4. M. biceps femoris (d). Origin: from the tho-ischium,
under and behind the origin of the gluteus maximus. Inser-
tion: partly into the top of the fibula, forming a pulley for the
agitator caudze (c), and an additional strap for the rectus fe-
moris (a), and partly, by means of another tendon, into the
head of the peronzeus longus (y). 0°05 oz.
5. M. semitendinosus (e). Origin: from the posterior point
of the tuber ischii. Insertion: by a remarkable looped tendon
having one end inserted into the back of lower end of femur,
and the other end into the os calcis. 0°18 oz.
6. M. semimembranosus (f). Origin: tuber ischi. Inser-
tion: into the top of the tibia, by a tendon common to this
muscle and the gracilis. 0°11 oz.
The muscles of the calf, shown in the figure, are the follow-
ing :—
7. M. gastrocnemius (w). This muscle, as usual, has an outer
and inner head. 0:14.02. Outer head:—Origin: from the tendon
of the great caudal extensor of the thigh, half an inch from its
insertion into the outer condyle. Insertion: into the under side
of the outer tarsal bone (vide a, fig. B) and into the plantar
fascia. O°ll oz. Inner head:—Origin: from the top of fibula
and inner condyle of femur. Insertion: by a tendon, which
unites with that of the outer head before reaching the os calcis,
* This muscle may be the plantaris.
828 | Rev. 8S. Haughton on the Muscular Anatomy
under which it passes to be inserted into the outer and under
side of the outer tarsal bone. 0:03 oz.
M. plantaris? (z). Origin, double: from rectus femoris
and from agitator caude. Insertion: having become partially
blended with the outer gastrocnemius, it is inserted into the os
ealcis and under surface of the plantar fascia. 0°04: oz.
M. peroneus longus (y). Origin: from the shaft of the
fibula and from the tendon of the biceps femoris (d). Insertion:
into the outer tarsal bone, uniting with the tendon of the
gastrocnemius. 0:03 oz.
Mm. tibialis anticus and exterisor digitorum communis.
Insertion: into the tarsal ends of first, second, and third meta-
tarsal bones. O:11.
The interlacing of muscles in the thigh and leg of the Croco-
dile, just described, is very remarkable, and more complicated
even than that found in the Ostrich; and at first I was dis-
posed to think that it threw some doubt on the explanation I
had given previously of the reason for such an arrangement in
the bird’s leg. In the case of the Ostrich, the necessity for
strict simultaneity of action was made evident by the great force
of the muscles employed, and the great delicacy of the bones on
which they had to act. What could there be, in the case of the
Crocodile, to correspond to such a peculiarity i in the case of the
Ostrich? After some careful dissection, | found the ready
answer to my question in the remarkable muscle which I shall
now describe.
On clearing away the superficial muscles of the thigh and
tail, I found the enormous mass of muscle, figured at J, fig. B,
Pl. XVI., which acts as the chief and powerful extensor of the
thigh :—
11. M. extensor femoris caudalis* (6). Origin: from the
transverse and inferior spinous processes of the caudal vertebree, °
from the third to the fifteenth inclusive. Insertion: into the
back of the upper part of the femur, and into a great round
* This remarkable muscle is noticed and accurately described by Meckel
in tom. 11. p. 152, 153 of his ‘ System der vergleichenden Anatomie,’ Halle,
1828; but it is very strange that he transposes its origin and insertion,
and seems not to have had any idea of its real use. It is regarded from
his point of view as a descriptive anatomist, and without the remotest
reference to its final cause. He says, “ Der zweite, tiefere, weit dickere
Muskel ist von dem ersten [the superficial. muscle of the tail] wie einer
breiten Binde umgeben, entspringt mit zwei ganz getrennten; einer weit
kiirzern, breiten Sehne oben von den hintern Flache des Oberschenkel-
beines; durch eine weit langere, schlanke, unten zwischen den beiden
Gelenkknorren desselben Knochens, und setzt sich an die ganze Seiten-
flache der untern Dornen, so wie der Zwischendornenhaut und die untere
Flache der Wurzeln der Querfortsatze.”’
of the Leg of the Crocodile. 829
tendon, which receives, in particular, the anterior fibres of this
enormous muscle, and, passing down the back of the femur, is
inserted by a strong common aponeurosis into the outer con-
dyle of the femur and into the head of the fibula. This com-
mon aponeurosis also gives a partial origin to the gastrocnemius
(a, fig. B) and to the plantaris (2, fig. A). 1°81 oz.
There are two muscles, accessory to this great caudal extensor
in their action, which are as follows :—
12. M. extensori femoris caudali accessorius. Origin: from
the fascia covering the great caudal extensor, and by a tendinous
head from the quadratus femoris, which is also an accessory to
the great caudal. Insertion: into the looped tendon of the
semimembranosus already described. 0-01 oz.
13. M. quadratus femoris. Origin: posterior, superior, and
inner surface of the pubis, near its symphysis. Insertion: into
the back of the femur, with the action and position of the
quadratus femoris in mammal quadrupeds, and into the tendon
of the great caudal extensor. 0°05 oz.
The effect of the interlacing of the tendons of the various
muscles already described must be to produce simultaneity of
action among them, such as I have already endeavoured to de-
scribe in my account of the leg of the Ostrich ; and in the pre-
sent instance of the Crocodile there seems to be a similar prin-
ciple involved. The Crocodile, resting on mud, progresses
chiefly by using his hind feet as paddles; and in this use of
them the great caudal extensor of the thigh is the most powerful
and important muscle employed. And it seems to me that the
simultaneity of action of all parts of the leg, rendered necessary
by the employment of so powerful a muscle, is fully secured by
the interlacing of the tendons I have described, which renders
it impossible for one set of muscles to act without the others
being also exerted. :
The remaining muscles of the posterior limb are as follows :—
14. M. glutzus medius. Origin: from the central part of
the ilio-ischiadic surface. Insertion: its tendon passes over the
great trochanter, to be inserted into a line down the upper half
of the outside of the femur, between the origins of the two por-
tions of the vastus externus. 0:06 oz.
15. M. gluteus minimus. Origin: from the anterior point
of the ilium. Insertion: into the inner side of the knee, under
the fascia of the rectus femoris. 0:02 oz.
16. Mm. vastus internus, externus, et crureus. The vastus
externus consists of two distinct muscles, as in the Ostrich.
0°22 oz.
17. M. psoas. This large muscle takes an origin as high as
the last rib, and is inserted into the lesser trochanter and the
380 On the Muscular Anatomy of the Leg of the Crocodile.
intertrochanteric line leading to the outer side of the femur. It
lies outside the iliacus. 0°57 oz.
18. M. iliacus. Origin: from the anterior transverse surface
of the ilium, with a slip from the spine. Insertion: altogether
into the lesser trochanter. 0-11 oz.
19. M. sartorius. Origin: behind the origin of the rectus,
on the inner side, at the junction of the ilium and marsupial
bone. Insertion: into the fascia of the inner side of the thigh,
for two-thirds of its length. 0-04 oz.
20. M. gracilis takes an origin from two heads, one at the
posterior point of the pubis and the other on the pectineal line.
Insertion: into the head of the tibia by a tendon common to it
with the semimembranosus. 0:08 oz.
21. M. pectineeus. Origin: between the two heads of gra-
cilis, from the central part of the surface of the pubis and from
the pectineal lme. Insertion: into the top of the linea aspera.
0:06 oz.
22. Mm. adductores. There are three adductor muscles :—
Ist adductor. Origin: anterior pectineal line of pubis. In-
sertion: into the upper half of the linea aspera. 0°13 oz.
Qnd adductor. Origin: from posterior edge of pubis, its middle
third. Insertion: into the middle of the linea aspera. 0:03 oz.
8rd adductor. Origin: from the posterior edge of the pubis,
close to the symphysis. Insertion: into the back of the top of
the fibula, with a fascial union with the tendon of semitendi-
nosus. 0:05 oz.
23. M. obturator externus? Origin: from the tuber ischil,
the posterior edge of the ischium, and the obturator membrane.
Insertion : an oblique line in the back of the femur, below the
insertion of the quadratus femoris. 0°13 oz.
24. M. marsupialis externus. 0°07 oz.
25. M. marsupialis internus. 0:10 oz.
These two muscles take their origin respectively from the
outer surface of the marsupial bone, and from its inner surface
and the last abdominal rib; and they are inserted by a common
tendon into the top of the posterior intertrochanteric line.
Their action is to rotate the femur directly inwards.
26. M. flexor proprius hallucis. Origm: from the outer
condyle of femur. Insertion: into the first, second, and third
toes. 0:02 oz.
27. M. flexor digitorum communis. Origin: from the fibula
and tibia. Insertion: into the first, second, and third toes.
0:05 oz.
28. M. tibialis posticus. This muscle is inserted into the
tarsal ends of the first, second, and third metatarsal bones.
0:06 oz.
Prof. G. Gulliver on Raphides and other Crystals in Plants. 331
29. M. peronzo-caleaneus. Origin: from lower part of
shaft of the fibula. Insertion: ito the upper surface of the
calcaneum. 0:01 oz.
XXX VII.— Observations on Raphides and other Crystals in Plants.
By Grorce Guiuiver, F.R.S.
[Continued from p. 117.]
Vitacee and Araliacee.—In the last communication (‘Annals’
for Aug. 1865) it was stated that raphides abound in all the
plants, therein specified, which I had examined of the order
Vitacese, while every species of the allied or related orders, of
which comparative examinations were made, proved to be devoid
of this raphidian character. I have had an opportunity, through
the courtesy of a botanical friend, of dissecting a dried fragment
of the receptacle-stalk of that most curious plant, Plerisanthes
(Vitis Pterisanthes, Mic., 8. borneensis), a bit of the dried leaf-
blade and fruit-shell of Bersama abassynica, Fresen., and a part
of the dried leaf and flower of Natalia lucens, Hochst. (Rhaganus
lucens, E. Meyer). To the same genus, I have been told, Ber-
sama abassynica is referred by Hooker and Bentham.
Pterisanthes, like Vitis, Cissus, and Leea, abounds in true
raphides and spheraphides. The raphides of Pterisanthes are
about =4,th of an inch long and ;;1,, thick; the average
diameter of the spheraphides is +,55th of an imch. The Ber-
sama and Natalia are destitute of true raphides, but contain
numerous crystal-prisms, about ~1,th of an inch long and
=aszth thick. These may be well seen in the leaf and imner
membrane of the fruit-shell of Bersama, and in the leaf, calyx,
petals, and pedicel of Natalia. The prisms have four equal
faces, and their ends slope off either from angle to angle or
from face to face.
Thus species of all the genera adopted by Lindley under the
order Vitacee—Cissus, Vitis, Pterisanthes, Leca, and Rhaganus
—have now been examined, though too often im imperfect or
unsatisfactory fragments ; and in every one of these plants true
raphides were found, except the Bersama and Natalia (Rhaganus),
in which raphides are replaced by crystal-prisms. It may be
recollected that a like phenomenon occurs in the last order
(Roxburghiacez) of the raphidian class Dictyogenz, as described
in the ‘ Annals’ for June 1865.
Of Araliaceze and Vitacez, the comparative structure in the
leaves and some other parts has already been described (‘Annals ”
for August 1865). I have lately examined fresh leaves and twigs
of Aralia spinosa, and a bit of a dried leaf-blade of A. racemosa,
332 Prof. G, Gulliver on Raphides and other Crystals in Plants.
These plants, like their congeners of the same order, abound in
spheraphides, but are destitute of raphides. Nothing of the
kind can be more beautiful than the spheeraphid tissue (‘Annals,’
Sept. 1863), extending beneath the cuticle of the whole leaf and
in the bark, of A. spinosa. These spheraphides are about
izizzth of an inch in diameter; they are somewhat larger in the
bark, and larger still in the pith.
“Tf,” says Prof. Lindley, “the Vine is compared with Aralia
racemosa, the relationship of Vitaceze to it will be too obvious
to be mistaken. Suppose that Aralia racemosa had an adherent
calyx, erect ovules, with stamens opposite the petals, and it
would be a Vitis.” But now, while recognizing the similarity
or identity of the sphzraphides of Araliaceze and Vitacez, we
perceive that Arala would require also the raphidian character
to be a Vitis. This difference is so remarkable that it may be
very easily and quickly seen by a comparison of the cells in the
leaves and other parts of the species before named of Vitis and
Aralia. Indeed the contrast in this respect between these plants
forms a very pretty microscopic object, and for this purpose the
leaves of Vitis apitfolia and Aralia spinosa answer admirably.
Callee, Orontiea, and Acoree.—We have already seen, under
the head of the raphidian order Aracez (‘ Annals,’ May 1865,
p- 881) how raphides could not be found in Acorus calamus.
In a fragment of a dried leaf of Gymnostachys anceps I found a
few raphides or raphis-like objects; but this scarcely affects the
fact of the deficiency of raphides in Acorez, and their profusion
in the members of the two other tribes of Orontiaceee—a re-
markable difference, which may be well seen by a comparison of
Calla, Monstera, Pothos, and Orontium with Acorus and Gym-
nostachys.
Hemodoree, Conostylee, and Velloziee.—Besides the species
formerly mentioned as affording numerous raphidian cells, my
dissections of Hemodorum planifolium, Anigosanthus rufus, and
A. humilis show a more or less abundance of raphides im all
these plants. But Vellozia is probably devoid of raphides; for
I could not detect them either in the withered leaves or bark of
an old dead trunk of this plant. Hence a comparative examina-
tion of the cellular structure of the Conostyles of New Holland and
the Vellozias, as well as of all the other species now placed by
botanists under Heemodoraceze, appears to be very needful and
likely to increase our know ledge of the natural affinities of this
curious order. After describing some analogy between the
stems of Pandanus and Vellozia, “Prof. Lindley judiciously says,
“ Don proposed to make an order of the Vellozias ; but, till their
structure and that of the Bloodroots shall have been thoroughly
investigated, this step would be premature.”
Prof. F. M‘Coy on the Cretaceous Deposits of Australia. 833
Pandanacee.—This order, like all the rest of Lindley’s Aral
Alliance, abounds in raphides. Besides the plants noticed in
the ‘Annals’ for May 1864, I have lately examined a leaf and
the bark of Pandanus odoratissimus and the root and a leaf of
Freycinetia imbricata. In the leaf of the former plant raphides
swarm, and occur more or less plentifully in the mesophleum,
endophleeum, and alburnum; of the latter plant raphides are
very abundant in the root, but less so in the leaf.
Thus, as far as these observations warrant the inference,
Pterisanthes and Leea have the intimate structure of a Vitis,
while the two species of Rhaganus (Bersama or Natalia) depart
from that structure, but agree well together. Acorus and Gym-
nostachys are deficient in the raphidian character of their allies ;
and the Velloziz differ in like manner from Hemodorez, Cono-
style, Pandanez, and Cyclanthee.
Edenbridge, Oct. 14, 1865.
XXXVITI.—WNote on the Cretaceous Deposits of Australia. By
Freperick M‘Coy, Professor of Natural Science in the Uni-
versity of Melbourne, and Director of the National Museum
of Victoria.
Messrs. D. Carson and J. Sutherland, of Collins Street, Mel-
bourne, recently placed in my hands, for our public Museum,
a series of specimens which they collected on the western bank
of the Flinders River, at the base of Walker’s Table Mountain,
nearly in the middle of the continent, in lat. 21° 13’ and
long. 143° 25’. The examination of these enables me to an-
nounce for the first time with certainty the existence of the
Cretaceous formations in Australia. Mr. Gregory doubtfully
indicated Cretaceous fossils in his last paper to the Geological
Society, but without any generic or specific recognition of fos-
sils of that age ; and his materials, when referred to by the officers
of the Geological Society, were only quoted as Mesozoic. Mr.
Selwyn also alluded formerly to a specimen of an Echinide in
flint, given to him as found in gravel in sinking a well at
Prahran, near Melbourne, having been identified by me as the
European Cretaceous Conulus albogalerus; and I had a flint
Ananchyies ovatus of the same age, given to me as found at
Richmond, near Melbourne also; but both of those specimens
were unsatisfactory, as far as the proof of their having really
belonged to any Australian stratum. I can now, however, re-
cognize the Lower Chalk; and this nearly fills up the great
series of marine Mesozoic formations supposed to be absent in
Australia when I left Europe, but most of which I have recog-
334 Mr. H. L. Smith on a new Growing Slide for the Microscope.
nized from fossil evidence. The most common of the fossils is
a species of Inoceramus with coarsely fibrous shell, nearly + inch
thick, agreeing in size and shape almost exactly with the English
Inoceramus mytiloides (Sow.), from which it differs in having
the hinge-line rather longer, the anterior end more poited, and
the superior posterior angle rather more obtuse. This species
I have named, in a paper read recently before the Royal Society
of Victoria, L. Carsoni (M‘Coy), in honour of one of the donors.
The second most common fossil is a much larger and broader
species of the same genus, which I at the same time named
Inoceramus Sutherlandi (M‘Coy), after the other donor of the spe-
cimens, which were so painfully carried, from the remote point
indicated to the settled districts, on their saddle. This second
species, in form, size, and concentric undulations of the surface,
nearly agrees with the French and English common Cretaceous
I. Cuvieri, but is less curved at the ventral mar gin near the beak.
The next shell is an Ammonite, in size, number and involu-
tion of whorls, shape, markings, and septa, so nearly identical
with the very common A. Beudanti (Br.) of the French Lower
Chalk, that, but for being slightly less compressed, and a slight
difference in some of the septal lobes, it could scarcely be sepa-
rated, even as a variety. I have named it Ammonites Flindersi
(M‘Coy), to call attention to the locality. It may be described
as follows :—
~ Ammonites Flindersi.
Discoid, moderately compressed ; periphery narrow, obtusely
rounded; whorls 44, about one-fourth of the width of each ex-
posed in an obtusely angular-edged, flat-sided umbilicus; sur-
face crossed by obtuse sigmoid striz, some of which are more
prominent than the more numerous intervening ones. ae
6 inches, proportional thickness 5225, width of last whorl 542..
Seven much divided lobes in the septa of each side, two of which
are within the edge of the umbilicus.
With these shells three vertebre of a large Teleosteous fish
occur.
The matrix of these specimens is an olive calcareo-argillaceous
marl.
XXXIX.—On a new Growing Slide for the Microscope.
By H. L. Smiru, Kenyon College, U.S.*
In studying the growth and conjugation of the Diatomacee, I
have felt the want of some means of keeping them alive for a
long time under the microscope, and have devised for this pur-
* From Silliman’s American Journal of Science, September 1865.
Mr. H. Ll. Smith on anew Growing Slide for the Microscope. 335
pose the slide to be described, which appears fully to meet all
requisitions ; and, as it can be readily made by any tolerably
expert microscopist, it will, I am certain, be considered a valu-
able addition to microscopical apparatus.
The whole slide, as I have constructed it, is a trifle more than
ith of an inch in thickness. It consists of two rectangular glass
plates, 3 x 2 inches and about .';th of an inch thick, separated
by thin strips of glass of the same thickness, cemented to the
interior opposed faces, as shown in the figure.
This closed cell, ultimately destined to be filled with water, is
not of such thickness as to prevent the use of the achromatic
condenser—a very important requisite. The glass I use is such
as is employed for the small cheap looking-glasses, and is easily
obtained.
The upper plate has a small hole (a) drilled through it. This
is effected by means of the or-
dinary writing-diamond and the
sharp edge ofa broken steel brooch }
or small rat-tail file. A hole can @
be drilled through glass of this fF
thickness in a few minutes. One ff
corner of the upper glass is re-
moved, as at 5, and a small strip ff
of glass cemented at c serves to !
prevent the thin glass cover placed.
over the object from sliding. Another slip of glass is cemented
on the lower side of the cell at d, but not extending as far as the
removed part at b. The object of this is to prevent the water
in the cell from being removed by capillary attraction, in case
the slide in the neighbourhood of 4 should be a little wetted.
This strip is not, however, absolutely necessary.
To use the slide, fill the space between the two plates with
clean water, introduced at 6 by means of a pipette, and also place
a drop on a to remove the air. The object being put on the top
of the slide and wetted, is now to be covered with a large square
of thin glass, e, at the same time covering the hole a. The
slide can now be placed upright, or in any position, as no water
can escape. It is, im fact, only a new application of the old
principle of the bird-fountain. As the water evaporates from
under the cover, more is supplied through the hole a, and from
time to time an air-bubble enters at b; thus a constant circula-
tion is maintained. A cell of the size named will need replen-
ishing only about once im three days, and this is readily effected
without disturbing the object. I have been enabled to make
observations by means of this slide which it would have been
very difficult, if not impossible, to have made without it.
-
af
336 Mr. J. Blackwall on new Species of Araneidea
XL.—Descriptions of recently discovered Species, and Characters
of a new Genus, of Araneidea from the East of Central Africa.
/ By Joun Bracxwatt, F.L.S.
Tue Spiders described in the following pages were captured in
the region through which the river Shiré flows to its confluence
with the Zambesi, by Mr. Horace Waller, at the particular request
of Mr. Richard Thornton, made shortly before he fell a victim
to the climate of Africa. These specimens were comprised in a
collection of Araneidea forwarded by the relatives of Mr. Thornton
to my friend Mr. Meade, who transmitted it to me for the pur-
pose of having its contents examined and descriptions made of
such species as might appear to be new to arachnologists, being
prevented himself from bestowing the requisite time on the
undertaking, for which he is so well qualified, by his numerous
professional engagements.
The late Mr. R. Thornton accompanied Dr. Livingstone in his
last expedition to South Africa, in the capacity of geologist ; and
Mr. H. Waller held the appointment of lay superintendent of
the mission to the Zambesi.
Tribe Octonoculina.
Family Lycost1p#.
Genus Crenvus, Walck.
Ctenus velox.
Length of the female ;2,ths of an inch; length of the cephalo-
thorax +, breadth +; breadth of the abdomen +; length of a
posterior leg 1,3,; length of a leg of the third pair +3.
The eyes are disposed on the anterior part of the cephalothorax
in three transverse rows; the two anterior ones, with the two
intermediate ones of the four constituting the second row, de-
scribe a trapezoid whose shortest side is before; and each of the
two eyes forming the posterior row, with a lateral one of the
second row, is seated on a tubercle ; the intermediate eyes of the
second row are the largest, and the lateral ones, which are in a
line with them, much the smallest of the eight. The cephalo-
thorax is truncated in front, compressed before, and rounded on
the sides, which are depressed and marked with furrows con-
verging towards a narrow indentation in the medial line of the
posterior region; it is clothed with short, dull-yellowish hairs,
and is of a reddish-brown colour, with narrow, dark-brown
lateral margins, parallel to which a broad, rather obscure, yel-
lowish-brown band, having its superior margin somewhat den-
tated, extends along each side; a narrow, pale red-brown band
passes from between the posterior pair of eyes to its base, and
from the East of Central Africa. 337
comprises in its anterior part a fine dark-brown line that termi-
nates at the medial indentation. The falces are powerful, coni-
cal, vertical, armed with teeth on the inner surface, provided
with long yellowish hairs, and have a dark-brown hue. The
maxille are long, straight, and truncated obliquely at the extre-
mity on the inner side, which is fringed with reddish hairs ;
and the lip is short, broad, and somewhat quadrate, but rounded
on the sides. These parts are of a reddish-brown colour, the
outer side of the maxillee and the base of the lip being much the
darkest. The sternum, which has a broad oval. form, is clothed
with short greyish hairs, interspersed with long upright ones of
a darker hue, and is of a pale red-brown colour. The legs are
long, provided with hairs and sessile spines, and of a reddish-
brown hue; the fourth pair is the longest, then the first, and
the third pair is the shortest ; the metatarsi and tarsi have hair-
hke papille on their inferior surface, and the latter are termi-
nated by two curved claws, pectinated at their base. The palpi
are long, and resemble the legs in colour (with the exception of
the base of the humeral joint, “which has a brighter tinge of red),
and have a short, curved, pectinated claw at their extremity.
The abdomen is oviform, hairy, convex above, and projects a
little over the base of the cephalothorax ; the upper part and
sides are of a brown colour freckled with yellowish grey, and a
broad dentated band of a yellower hue, bordered with black,
extends along the middle of the former, and comprises a longi-
tudinal yellow band in its anterior half; a broad, triangular,
black mark, having its truncated apex directed backwards, and
comprising within its base two oval white spots placed trans-
versely, occurs on the under part; it is bounded on each side
by a bright orange-red band, and these bands converge to the
spinners, where they meet ; the sexual organs, which are highly
developed, have a large process directed backwards from their
anterior margin, and, with the branchial opercula, are of a dark
reddish-brown hue, the latter beg the paler.
The specimen from which the description was made was the
only one of the species in the collection.
Ctenus vividus.
Length of the fouls 4°ths of an inch ; length of the cephalo-
iota 13, breadth 55, ; breadth of the abdomen =3;; length of
an anterior leg 1+; length of a leg of the third pair 1,3,.
The disposition and relative size of the eyes of this species are
similar to those of Ctenus velox. The legs are long, robust,
provided with hairs and sessile spines, and of a yellowish-brown
colour; the first pair is the longest, then the fourth, and the
third pair is the shortest ; the metatarsi and tarsi have hair-like
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. " 28
338 Mr. J. Blackwall on new Species of Araneidea
papillee on their inferior surface, and the latter are termimated
by two curved claws, pectinated at their base. The palpi are
long, and resemble the legs in colour; and the digital joint,
which is strongly tinged with brown, has a short, curved, pecti-
nated claw at its extremity. The cephalothorax is truncated
in front, compressed before, and rounded on the sides, which
are depressed and marked with furrows converging towards a
narrow indentation in the medial line of the posterior region ; it
is clothed with hairs, and of a brown colour, with a broad yel-
lowish-brown band (whose superior margin is somewhat dentated)
extending along each side, and a narrow longitudinal one in the
middle, which comprises a fine brown line in its anterior part.
The falees are powerful, conical, vertical, and densely clothed
with yellow hairs at the base in front; the maxillee are straight,
enlarged at the extremity, which is rounded on the outer side,
and obliquely truncated on the inner side, where it is supplied
with long hairs; and the lip is short, broad, and somewhat
quadrate, but rounded on the sides. These parts have a red-
brown hue, the falees being much the darkest. The sternum
has a broad oval form, with small eminences on the sides, oppo-
site to the legs; it is thinly clothed with long hairs, and of a
dark-brown hue, with a faint tinge of yellow in the medial line.
The abdomen is oviform, hairy, convex above, and projects a
little over the base of the cephalothorax; it is of a dull pale-
yellow colour, streaked and spotted with brown, the under part,
which is the yellowest, being marked with the fewest and smallest
spots; a broad, dentated, dull-yellow band, bordered with brown,
and comprising a longitudinal row of irregular brown spots, ex-
tends along the middle of the upper part, “and on each side of it
there is a series of brownish-black spots; the sexual organs are
moderately developed, with a large process directed backwards
from their anterior margin, whose extremity is dilated; these
organs, with the branchial opercula, are of a brownish-yellow
colour, the latter being the paler.
The collection contained six females of this species, five of
which were immature.
Genus PasttnHea, Blackw.
Pasithea pulchra.
Length of the female +2ths of an inch; length of the cephalo-
thorax - =§;, breadth 3; ; ; breadth of the abdomen 55 length of
an anterior leg 11; length of a leg of the third pair 1+
The eyes are unequal in size, and disposed in three transverse
rows on the anterior part of the cephalothorax, high above its
frontal margin; the four posterior ones form a curved row
from the East of Central Africa. 339
whose convexity is directed backwards, and the other four de-
scribe a trapezoid whose shortest side is before; the posterior
eyes of the trapezoid are seated on tubercles, and are the largest,
and the anterior ones, which are near to each other, are much
the smallest of the eight; the entire group describes a sector of
a circle, whose radii converge towards the frontal margin. The
cephalothorax is convex, glossy, compressed before, vertical in
front, rounded on the sides, which are marked with slight fur-
rows converging towards an indentation in the medial line of
the posterior region, and has a few black bristles behind and
before the eyes; it is of a yellow-brown colour, with a broad,
angular, pale-yellowish mark on the posterior part of the ce-
phalic region, whose vertex extends to the medial indentation,
and has a small red spot on each side, towards its termination ;
this angle is bisected by a faint line of the same hue, which
originates at the intermediate eyes of the posterior row; the
space comprised between the large posterior eyes of the trapezoid
and those of the posterior row is of a dark-brown colour; a red-
brown line extends from each minute anterior eye to the frontal
margin, and there is a parallel one of the same hue on each of
its external angles. The falces are long, powerful, subconical,
and vertical; they are of a dull-yellow colour, with longitudinal
lines of a réd-brown hue, corresponding to those in front of the
cephalothorax, the exterior ones being the shortest. The maxille
are long, convex near the base, which is curved towards the lip,
but straight and pointed at the extremity; the lip is much
shorter than the maxille, somewhat triangular, and hollowed at
the apex; and the sternum is heart-shaped, with a few black
bristles distributed over its surface. These parts are of a pale-
yellowish colour tinged with green, the sternum being the palest.
The legs are very long, slender, provided with hairs and long
spines, and are of a yellow colour, spotted with brownish black,
a longitudinal red line, of greater or less extent, occurring on
the inferior surface of the femora; the first pair is the longest,
then the second, and the third pair is the shortest; each tarsus
is terminated by three claws; the two superior ones are curved
and pectinated, and the inferior one is inflected near its base.
The palpi resemble the legs in colour, but are without any red
line on their inferior surface, and have a curved pectinated claw
at their extremity. The abdomen is of an oblong-oviform figure,
somewhat pointed at the spinners, convex above, and projects a
little over the base of the cephalothorax ; it is sparingly supplied
with hairs, and of a pale-yellow colour, finely reticulated with
dull green ; a yellow band, streaked and freckled with red, and
strongly dentated on its inner margin, extends from the anterior
extremity of the upper part along each side of the medial line
23*
340 Mr. J. Blackwall on new Species of Araneidea
to the spmners, where the two meet; and in the anterior part
of the space comprised between the two bands there is a short
longitudinal one, of a dark dull-greenish colour, which is
rather irregular in outline, ramified at its posterior extremity,
and has a short transverse bar of the same hue near its anterior
extremity ; a somewhat obscure yellow band extends along each |
side of the medial line of the under part, and two nearly parallel,
irregular, fine, black lines pass from the sexual organs halfway
towards the spmners; the sexual organs are highly developed,
with a strong septum in the middle, and are of a very dark
greenish colour, tinged with red, that of the spinners being pale
green.
Two adult females were comprised in the collection, one of
which had deposited its ova.
It is proposed to transfer the Spiders of the genus Pasithea
from the family Thomiside, in which they were originally placed,
to that of the Lycoside, as, by the disposition of their eyes and
the structure of their legs, they evidently possess a relation of
affinity to certain species of the genus Sphasus.
Family THomisip2&.
Genus SeLEenops, Dufour.
Selenops. alacer.
Length of the female #ths of an inch; length of the cephalo-
thorax ;5,, breadth =°,; breadth of the abdomen 1; length of
a leg of the third pair 1,1,; length of a leg of the first pair 1.
The eyes are disposed on the anterior part of the cephalo-
thorax ; four, nearly equal in size, form, immediately above the
frontal margin, a slightly curved transverse row, whose convexity
is directed forwards; the eyes of each lateral pair are seated
obliquely on a tubercle apart, the two anterior ones, which are
oval and the smallest and lightest-coloured, being situated a
little in advance of the lateral eyes of the transverse row ; and
this row, if extended, would include in its curve the two posterior
eyes, which are the largest of the eight. The cephalothorax is
large, depressed, compressed before, slightly rounded in front,
and greatly so on the sides, which are strongly marked with
furrows converging towards an oblong indentation in the medial
line of the posterior region ; it is of a red-brown colour, and is
supplied with hoary hairs having a yellowish tint. The falces
are short, powerful, conical, very convex in front, vertical, and
armed with a few teeth on the inner surface; the maxille are
straight, and increase in breadth towards the extremity, which
is obliquely truncated on the inner side; and the lip is semi-
circular. These parts are of a dark-brown colour, tinged with
from the East of Central Africa. 341
red; the falces are the darkest, and the extremity of the maxille
and apex of the lip have a brownish-yellow hue. The sternum
is nearly circular ; the legs are long, robust, and provided with
hairs and spines ; their relative length could not be ascertained,
as the second pair was missing; but the third pair is a little
longer than the fourth, which somewhat surpasses the first pair;
each tarsus is terminated by two plain, curved claws, and below
them there is a scopula; the palpi are short, and the digital
joint, which is provided with spines, has a curved, minutely
pectinated claw at its extremity. These parts are of a red-brown
colour; the sternum is the palest, and the metatarsal joint of
the legs and the digital joint of the palpi are the darkest-coloured.
The abdomen is of a depressed oviform figure, and the anterior
extremity, which is fringed with fine bristles, and has the ap-
pearance of having been cut directly across, projects very little
over the base of the cephalothorax; it is clothed with heary
hairs having a yellowish tint, and is of a dull brownish-yellow
colour, obscurely freckled with brown, particularly on the upper
part and sides; the sexual organs are moderately developed,
and have a triangular reddish-brown process connected with
each lateral margin, whose vertices nearly meet, a small semi-
circular one of the same hue directed forwards from the posterior
margin, and a larger semicircular process of a reddish-yellow
colour, directed backwards from the anterior margin.
The collection contained two adult females of this species.
Family Turriprip#&.
Genus Latropectus, Walck.
Latrodectus cinctus.
Length of the female 2ths of aninch; length of the cephalo-
thorax 4, breadth =3;; breadth of the abdomen +; length of an
anterior leg 2; length of a leg of the third pair +.
The abdomen is very convex above, thinly clothed with hairs,
and projects over the base of the cephalothorax; it is of a
brownish-black colour, with an oblong mark extending upwards
from the spinners, and three curved transverse bands of a deep
orange-colour on the upper part; the first of these bands is the
shortest, and is situated in front, the second is longer, and the
third, which is much the longest, and curved very obliquely,
increases in breadth towards its extremities, whose pointed ter-
mination is in contact with the posterior extremity of the oblong
orange-coloured mark above the spinners ; the sexual organs are
well developed, and of a dark-brown hue, tinged with red, that
of the branchial opercula being red-brown. Immature females
are of a pale-yellow hue where adults are orange-coloured. The
342 Mr. J. Blackwall on new Species of Araneidea
eyes are disposed on the anterior part of the cephalothorax, high
above the frontal margin, in two transverse, nearly straight
rows ; the four intermediate ones almost form a square, the two
anterior ones, which are seated on a prominence, being rather
nearer to each other than the two posterior ones; the lateral
eyes of both rows are the largest, and each is placed on a tubercle.
The cephalothorax is oval, somewhat convex, hairy, and has a
large indentation in the medial line of the posterior region; the
maxillz are short, obliquely truncated at the extremity, on the
outer side, and inclined towards the lip, which is broad and
semicircular ; the sternum is heart-shaped; the legs are robust
and hairy ; the first pair is the longest, then the fourth, and the
third pair is the shortest; each tarsus is terminated by three
claws ; the two superior ones are curved and pectinated, and the
inferior one is inflected near its base; the palpi are moderately
long, and have a curved, pectinated claw at their extremity.
These parts are of a dark-brown colour, the extremity of the
maxillze having a yellowish-brown tint. The falces are conical,
vertical, and of a red-brown hue.
An adult and an immature female of this Latrodectus were
comprised in the collection,
Family Eeriaipm.
Genus Eprira, Walck.
Epeira vigilans. ;
Length of the female ;2,ths of an inch; length of the cephalo-
thorax 1, breadth +; breadth of the abdomen ?; length of an
anterior leg +3; length of a leg of the third pair =3.
The cephalothorax is convex, compressed before, rounded in
front and on the sides, with an indentation in the medial line of
the posterior region; it is thinly clothed with yellowish-grey
hairs, and is of a red-brown colour, the sides being much the
darkest. The falces are powerful, conical, vertical, and armed
with teeth on the inner surface ; the maxillze are short, straight,
and enlarged and rounded at the extremity ; and the lip is semi-
circular, but somewhat pointed at the apex. These parts are of
a pale red-brown colour, the extremity of the maxillz and the
apex of the lip having a pale-yellow hue. The sternum is heart-
shaped, with small eminences on the sides, opposite to the legs,
and is of a dull-yellow colour, tinged with brown on the lateral
margins. The eyes are disposed on the anterior part of the
cephalothorax in two transverse rows ; the four intermediate ones
are seated on a protuberance, and describe a trapezoid, the two
anterior ones, which are the largest of the eight, being wider
apart than the posterior ones; the eyes of each lateral pair are
from the East of Central Africa. 343
placed obliquely on a small tubercle, and are near to each other,
but not in contact. The legs are long, provided with hairs and
spines, and of a red-brown colour, strongly tinged with brown
at the extremity of the joints; the first pair is the longest, then
the second, and the third pair is the shortest; the tarsi are ter-
minated by claws of the usual number and structure. The palpi
are of a red-brown hue, and have a curved, pectinated claw at
their extremity. The abdomen is of a depressed oviform figure,
and projects over the base of the cephalothorax; it is thinly
clothed with short pale hairs, and of a dull-yellow colour, with
a large oval brown mark on each side of its anterior extremity ;
a triangular brown spot, whose vertex extends between the two
oval marks, occurs in the medial line of the upper part, and
comprises three pairs of short oblique black streaks; this tri-
angular spot, which has a strongly marked streak of the same
hue directed obliquely outwards and backwards from each side,
is followed by a large, pale-brown, leaf-shaped mark that tapers
to the spinners, and has its sinuous margins bordered by a fine
black line whose continuity is much interrupted; the middle of
this mark is occupied by a longitudinal, taper, yellow band,
crossed by lines of the same hue, and comprising some short
black streaks in its anterior part, and several fine, long, brown
lines in its posterior part; two depressed dark-brown spots,
situated on each side of the medial line, nearly form a square,
the two posterior ones being larger and rather wider apart than
the anterior ones; the sides are marked transversely with nu-
merous fine brown lines, most of which meet at their superior
extremities ; the under part has longitudinal brown streaks on
its sides, and a large dark-brown mark in the middle, whose
contracted extremity extends to the spinners; it has an obscure
yellowish hue in the medial line, and is bordered anteriorly and
laterally with yellow, two spots of the same hue occurring on
each side of the brown spinners; the sexual organs, which are
not highly developed, have a long process directed backwards
from their anterior margin, whose surface is concave and extre-
mity rounded; they are of a red-brown colour, that of the
branchial opercula being yellowish-brown.
Three adult females of Hpetra vigilans were included in the
collection.
Genus Nepuiia, Leach.
Nephila Keyserlingit.
Length of the female 1,8, inch; length of the cephalo-
thorax 14, breadth ;; breadth of the abdomen +; length of
an anterior leg 27; length of a leg of the third pair 12.
The eyes are disposed on the anterior part of the cephalo-
344. Mr. J. Blackwall on new Species of Araneidea
thorax in two transverse rows; the four intermediate ones
nearly form a square, the two anterior ones, which are seated
on a protuberance, and are rather nearer to each other than the
two posterior ones, being the largest of the eight; the eyes of
each lateral pair are placed obliquely on a prominent tubercle,
and are separated by a considerable interval. The cephalo-
thorax is long, somewhat convex, particularly in the cephalic
region, truncated in front, compressed before, moderately rounded
on the sides, which are marked with furrows converging towards
a transverse pair of indentations in the medial line of the poste-
rior region, and has two conical glossy eminences placed trans-
versely near its middle; it is of a dark-brown colour, which is
almost concealed by a covering of short, adpressed, white hairs
having a silvery lustre. The falces are powerful, conical, ver-
tical, glossy, convex at the base, in front, and armed with teeth
on the inner surface; the maxille are short, strong, and greatly
enlarged and rounded at the extremity ; and the lip is somewhat
oval. These parts are of a very dark brown colour, the extremity
of the maxille, the apex of the lip, and a line extending along
the middle of the latter having a dull-yellow hue. The sternum
is heart-shaped and glossy, with prominences on the sides, and
a somewhat pointed one opposite to the base of the lip; it is of
a bright yellow colour, with a large, irregular, transverse dark-
brown mark in the middle, and narrow lateral margins of a
similar hue. The legs are very long, slender, provided with fine
spines and hairs, the latter bemg the longest and densest on the
inferior surface and sides of the tibize of all the legs except those
of the third pair; they are of a yellow colour; a broad annulus
near the middle and a narrow one at the extremity of the femora,
the genua, the base, and a broad annulus near the middle of the
tibize of the first and second pairs, the extremity of the femora,
the genua and tibiz of the third pair, and the extremity of the
femora, the genua, and about two-thirds of the tibie from the
base of the fourth pair, with the metatarsi and tarsi of all the
legs, have a dark-brown hue; the first pair is the longest, then
the second, and the third pair is the shortest ; the tarsi are ter-
minated by claws of the usual number and structure. ‘The palpi
are short, and of a yellow colour, the digital jot, which has a
curved, slightly pectinated claw at its extremity, being strongly
tinged with brown. The abdomen is subcylindrical, projecting
over the base of the cephalothorax, and is broader at the anterior
than at the posterior extremity, which is rounded and extends
beyond the spinners ; it is thinly clothed with short hoary hairs
on the upper part and sides, and is of an olive-brown colour, the
under part being the darkest ; a broad, yellow, transverse band
near the anterior extremity passes along the upper part of each
from the East of Central Africa. 345
side to the posterior extremity, its posterior half being irregular
in outline, particularly on its superior margin ; four large yellow
spots are disposed longitudinally in the medial line, and a series
of four small, depressed, brown spots occurs on each side of it ;
several fine brown lines extend along the middle, which are most
conspicuous on the yellow spots ; a yellow band, whose posterior
half is broken into spots and streaks, some of which are covered
with white hairs, passes along the lower part of each side to the
spinners, and above those organs there are six small yellow spots
disposed in pairs; on the under part a yellow, curved, transverse
band passes immediately below the sexual organs, and midway
between those organs and the spinners there is an irregular
transverse band of the same hue; the sexual organs are well
developed and prominent, the anterior margin being oval, and
the posterior one somewhat triangular; these organs, with the
spinners and branchial opercula, have a dark-brown hue, that
of the inner margin of the opercula being pale-yellow.
The collection contained four females of this large and hand-
some species of Nephila, with which I have connected the name
of M. Keyserling, whose researches in this department of arach-
nology have contributed greatly to extend our knowledge of the
Epeiride. :
Nephila venusta.
Length of the female 4ths of an inch; length of the cephalo-
thorax =3,, breadth 4; breadth of the abdomen -3,; length of
an anterior leg 12; length of a leg of the third pair 2.
The legs are long, slender, provided with fine spines and hairs,
the latter being the longest and most abundant on the tibie and
base of the metatarsi of the first, second, and fourth pairs ; they
are of a reddish-brown colour, the metatarsi and tarsi being the
darkest ; the first pair is the longest, then the second, and the
third pair is the shortest; the tarsi are terminated by claws of
the usual number and structure. The palpi are short, and paler
than the legs, the extremity of the radial and the whole of the
digital joint being strongly tinged with brown; the latter is
supplied with long hairs and spines, and has a curved, slightly
pectinated claw at its extremity. The eyes are disposed on the
anterior part of the cephalothorax in two transverse rows; the
four intermediate ones nearly form a square, the two anterior
ones, which are seated on a protuberance, and are rather nearer
to each other than the two posterior ones, being the largest of
the eight ; the eyes of each lateral pair are placed obliquely on
a prominent tubercle, and are separated by a considerable in-
terval. The cephalothorax is long, somewhat convex, particu-
larly in the cephalic region, truncated in front, compressed be-
fore, moderately rounded on the sides, which are marked with
346 Mr. J. Blackwall on new Species of Araneidea
furrows converging towards a transverse pair of indentations in
the medial line of the posterior region, and has two small, coni-
cal, glossy eminences placed transversely near its middle, and
numerous very minute ones on its lateral margins; it is of a
very dark brown colour, which is almost concealed by a covering
of short, adpressed white hairs having a silvery lustre. The falces
are powerful, conical, vertical, glossy, and armed with teeth on
the inner surface; the maxille are short, strong, and greatly
enlarged and rounded at the extremity ; and the lip is somewhat
oval. These parts are of a dark-brown colour, the falces being
much the darkest ; the maxille have a yellow-brown hue at their
extremity, on the inner surface, and the lip is marked with a
longitudinal yellow band, which is contracted in the middle.
The sternum is heart-shaped and glossy, with prominences on
the sides and a pointed one opposite to the base of the lip; it is -
of a bright-yellow colour. The abdomen is subcylindrical, pro-
jecting over the base of the cephalothorax, and its posterior ex-
tremity, which is rounded, extends a little beyond the spinners ;
it is of a yellow-brown colour, reticulated and spotted with pale
dull-yellow on the upper part and sides, the anterior extremity
being the yellowest ; many of the spots, which vary in size and
form, are covered with white hairs having a silvery lustre; a
series of three depressed spots occurs on each side of the medial
line, and on its posterior half four fine lines are disposed longi-
tudinally ; these spots and lines have a brown hue, and two
large irregular spots of a darker brown are situated on the lower
part of each side; the under part is of an olive-brown colour,
with four slightly curved, pale-yellow lines, which describe a
quadrilateral figure in the middle ; the spinners and the branchial
opercula are of a brown colour, the inner margin of the latter
having a dull-yellow hue; the sexual organs are well developed,
with a longitudinal septum in the middle, and the anterior mar-
gin presents two prominent, glossy convexities; their colour is
dark-brown tinged with red.
Two adult females of Nephila venusta were included in the
collection.
Genus Arcyopes, Savigny.
Argyopes caudatus.
Length of the female 1,3, inch ; length of the cephalothorax 2,
breadth }; breadth of the abdomen +%;; length of an anterior
leg 2; length of a leg of the third pair 1,1,
The abdomen is large, and of a depressed oviform figure ; its
anterior extremity, which has the appearance of having been
cut directly across, projects over the base of the cephalothorax,
and its posterior extremity extends greatly beyond the spinners;
from the East of Central Africa. 347
three strong conical prominences project from each side; and it
is terminated by a membranous caudal appendage, with a minute
conical prominence on each side of its base; the upper part is
clothed with hoary hairs, and is of a yellow colour, with a broad,
semilunar, brown mark at its anterior extremity, whose convexity
is directed upwards; to this mark succeed five transverse bands
of a darker hue, whose margins are very irregular, the first and
last having their continuity interrupted in the middle; there is
a series of four depressed brown spots on each side of the medial
line, and four fine lines of a similar hue are disposed longitudi-
nally on the posterior part; the colour of the caudal appendage
is brown, and above it there is an oblong yellow spot ; the sides
and under part have a brown hue; the former are corrugated
and marked with brownish-yellow spots and streaks, which are
almost concealed by hoary hairs, and the yellow lateral tubercles
are marked transversely with brown in the middle; a series of
large, irregular, yellow spots extends along each side of the me-
dial line of the under part, and minute spots of a similar hue
occur in the interval between the two; the sexual organs, which
are moderately developed, have a long, pale reddish-brown pro-
cess in connexion with their anterior margin that is directed
backwards, and their predominant colour, with that of the
branchial opercula, is dark-brown faintly tinged with red, the
latter being the paler. The eyes are disposed on the anterior
part of the cephalothorax in two transverse rows; the four in-
termediate ones nearly form a square, the two anterior ones,
which are seated on a small protuberance, being the largest of
the eight; the eyes of each lateral pair are placed obliquely on
a tubercle, and are near to each other, but not in contact, the
anterior ones being much the smallest. The cephalothorax is
compressed before, rounded in front and on the sides, slightly
convex, with a broad, shallow indentation in the medial line of
the posterior region ; it is of a dark-brown colour, with a broad
pale reddish-brown band extending from the front to the medial
indentation, and brownish-yellow lateral margins ; these colours
are concealed, except at the base, by a dense covering of white
adpressed hairs. The falces are powerful, conical, vertical, armed
with teeth on the inner surface, and of a red-brown colour.
The maxillze are short, strong, and greatly enlarged and rounded
at the extremity ; the lip is semicircular, but somewhat pointed
at the apex; and the sternum is heart-shaped, hairy, and has
eminences on the sides opposite to the legs. These parts are of
a dark-brown colour; the extremity of the maxilla and the
apex of the lip have a brownish-yellow hue ; and a band extend-
ing along the middle of the sternum, from each side of which a
streak is directed obliquely backwards and outwards, and two
318 Mr. J. Blackwall on new Species of Araneidea
minute spots on each side of its anterior part are of a yellow
eolour. The legs are long, provided with hairs and spines, and
are of a dark-brown colour, the inferior surface of the coxee and
of the femora, especially at their base, being strongly tinged
with dull-yellow; the first pair is the longest, then the second,
and the third pair is the shortest; the tarsi are terminated by
claws of the usual number and structure. The palpi are of a
dull-yellow colour, the digital joint (which is supplied with spines
and terminated by a curved, pectinated claw) being tinged with
brown at its extremity.
The collection contained four adult females of this species.
Genus Eurysoma, Koch.
Eurysoma Thorntoni.
Length of the female +$ths of an inch ; length of the cephalo-
thorax .{;, breadth +; breadth of the abdomen ;4; length of a
posterior leg +; length of a leg of the third pair 1.
The eyes are situated near the frontal margin of the cephalo-
thorax ; the four intermediate ones are seated on a protuberance
and form a square, the two anterior ones being the largest and
darkest-coloured of the eight ; the eyes of each lateral pair are
placed a little apart on a tubercle, and are distant from the four
intermediate ones. The cephalothorax is convex, glossy, de-
pressed towards the extremities, compressed before, broadly
truncated in front, rounded on the sides, which are marked with
furrows converging towards an indentation in the medial line of
the posterior region, and is of a yellow-red colour. The falces
are short, powerful, conical, armed with teeth on the inner sur-
face, inclined towards the sternum, and of a brown colour, faintly
tinged with red. The maxille are straight, and enlarged and
rounded at the extremity; the lip is semicircular; and the ster-
num is heart-shaped, with eminences on the sides, opposite to
the legs. These parts are of a yellow colour, the maxillz and
lip having a tinge of brown. The legs are short, provided with
hairs, and are of a dark-brown colour, with the exception of the
coxee and about two-thirds of the femora from the base, which
have a yellow hue; the fourth pair is the longest, then the first,
and the third pair is the shortest; the tarsi are terminated by
claws of the usual number and structure. The palpi, which are
short, resemble the legs in colour, and have a curved, pectinated
claw at their extremity. The abdomen is nearly circular, rather
broader than long, without spines, glossy, moderately convex
above, and projects greatly over the base of the cephalothorax ;
the upper part is of a deep-black hue, marked with numerous
circular depressions, and with twenty-two bright-yellow spots,
from the East of Central Africa. 349
of various shapes and sizes; sixteen occur on the margins, two
in the medial line, and two on each side; the two smallest are
situated on the frontal margin, and between them there are two
others of an oval form; these latter spots are followed by a cir-
cular one in the medial line, to which succeeds a large one of
an oblong heart-shape, veined with pale-brown; the anterior
spot on the side is large and reniform, and the posterior spot is
connected with a marginal one; the under part is much corru-
gated, and of a brownish-black colour, that of the spinners,
which are encircled by a rim, being dark-brown; a large semi-
circular band behind the spinners, and the space surrounding
the pedicle by which the cephalothorax is attached to the abdo-
men, including the branchial opercula, are of a yellow hue, with
transverse, forked, soot-coloured lines; the sexual organs are
moderately developed, and of a dark-brown colour, tinged with
red, that of the anterior margin, which is semicircular and rather
prominent, being brownish-yellow.
This elegantly marked Eurysoma, two specimens of which
were comprised in the collection, is dedicated to the memory of
of the late Richard Thornton, Esq., whose premature death,
deeply deplored by his friends, has removed from his sphere of
usefulness here an intelligent and zealous votary of natural
science.
Eurysoma Walleri.
Length of the female +/ths of an inch ; length of the cephalo-
thorax =3,, breadth -2;; breadth of the abdomen 43; length of
a posterior leg 41; length of a leg of the third pair 4.
The cephalothorax is convex, depressed towards the extremi-
ties, glossy, compressed before, broadly truncated in front,
rounded on the sides, which are marked with slight furrows con-
verging towards an indentation in the medial line of the poste-
rior region, and is of a brownish-yellow colour. The falces are
short, powerful, conical, armed with teeth on the inner surface,
inclined towards the sternum, and of a brown hue, tinged with
yellow on the sides and at the base. The maxille are straight,
and enlarged and rounded at the extremity; the lip is semi-
circular; and the sternum is heart-shaped, with eminences on
the sides, opposite to the legs. These parts have a yellow hue,
the maxillz and hp being tinged with brown. The eyes are
situated near the frontal margin of the cephalothorax; the four
intermediate ones are seated on a protuberance, and form a
square, the two anterior ones being the largest and darkest-
coloured of the eight ; the eyes of each lateral pair are placed
apart on a tubercle, and are distant from the four intermediate
ones. The legs are short, provided with hairs, and are of a
yellow colour, with brownish-black annuli, each femur haying a
350 Mr. J. Blackwall on a new Genus of Araneidea
single annulus at its extremity; the fourth pair is the longest,
then the first, and the third pair is the shortest; the tarsi are
terminated by claws of the usual number and structure. The
palpi are short, somewhat darker-coloured than the legs, and
have a curved, pectinated claw at their extremity. The abdomen
is nearly circular, rather broader than long, without spines,
glossy, moderately convex above, and projects greatly over the
base of the cephalothorax; it is of a yellow colour, with five
depressed brownish-black spots, forming, near the frontal mar-
gin, a transverse curved row whose convexity is directed back-
wards, the two exterior spots being the smallest of the five ;
four similar spots occur on each side, parallel to the margin ;
four smaller ones form a transverse, slightly curved row near
the posterior margin, whose convexity is directed forwards ; and
there are three similar spots on each side of the medial line,
those constituting the intermediate pair being the largest and
widest apart ; the under part is much corrugated, and has a
large, triangular, dark-brown mark in the middle, comprising
within its vertex. the spinners, which are encircled by a rim ;
the whole is encompassed by a broad yellow margin marked
with depressed brownish-black spots, from which lines of the
same hue pass towards the centre; the sexual organs are mo-
derately developed, with a small, oval, brownish-yellow. process
in connexion with their anterior margin, and are of a dark-brown
hue, tinged with red, that of the branchial opercula being
yellow,
I have conferred upon this syecies the name of Horace Waller,
Esq., an ardent naturalist, and the friend and fellow-traveller of
Mr. Thornton,
Eurysoma Walleri bears/a strong resemblance to the Gaste-
racantha hemispherica of /M. Koch (Die Arachniden, Band xi.
p- 49, tab. 375. fig. 874), but differs from it in the number,
distribution, and relative size of the depressed brownish-black
spots on the abdomen, and also in some other particulars. The
generic name Gasteracantha being quite inapplicable to a spider
absolutely devoid of spines, I have placed this species, notwith-
standing the numerous depressed dark-coloured spots with
which its carapace is marked, in the genus Hurysoma, to which,
for the same reason, I think it would be expedient to transfer
Gasteracantha hemispherica.
Genus Pycnacantua, Blackw.
Eyes small, disposed on the anterior part of the cephalo-
thorax ; the four intermediate ones are seated on a narrow, pro-
minent protuberance directed obliquely upwards and forwards,
and nearly describe a square; the two superior ones are placed
from the East of Central Africa. 351
on the summit of the protuberance, and the two inferior ones,
which are rather wider apart and the largest of the eight, are
situated near its middle, in front ; the lateral eyes are the small-
est ; those of each pair are seated obliquely on the outer side of
a slender, elevated, upright tuoercle, a little below its somewhat
pointed extremity, and are separated by a moderately wide
interval.
Mazille short, straight, and enlarged and rounded at the
extremity.
Lip semicireular, but slightly pointed at the apex.
Legs moderately robust ; the first and second pairs are much
longer than the third and fourth pairs, the first pair being the
longest and the third the shortest.
Abdomen subglobose, provided on the upper part and sides
with numerous, close-set, sharp-pointed spines, varying greatly
in their dimensions.
Pycnacantha Meadi.
Length of the female + an inch; length of the cephalo-
thorax +, breadth $; breadth of the abdomen 1; length of an
anterior leg =4;; length of a leg of the third pair 3.
The cephalothorax is compressed before, truncated in front,
with prominent lateral angles, broadly rounded on the sides,
somewhat convex, and has a shallow indentation in the medial
line of the posterior region; it is thinly clothed with short,
whitish hairs, and of a dull-yellow colour, the base bemg much
the palest ; a narrow brown band passes from between the lateral
tubercles to the medial indentation ; a longitudinal one, of a
similar hue, but somewhat paler, whose posterior part is much
the broadest, occurs on each side, and small black prominences,
which are most numerous on the medial band, are distributed
over its surface. The falces are conical, vertical, armed with
teeth on the imner surface, and of a pale-yellow colour in front,
the outer side and extremity having a reddish-brown hue. The
maxillee and lip are of a dark reddish-brown colour, the extremity
of the former and the apex of the latter having a yellowish-white
hue. The sternum is heart-shaped, with small eminences on the
sides, opposite to the legs, and is of a yellow colour, with dark-
brown margins. The legs are provided with hairs and spines,
two parallel rows of the latter occurring on the inferior surface
of the metatarsi and tarsi of the first and second pairs; they
have a pale dull-yellow hue, and the tarsi are terminated by
claws of the usual number and structure. The palpi resemble
the legs im colour, and have a short, curved, pectinated claw at
their extremity. The abdomen, which is clothed with short,
pale hairs, is of a subglobose form, very convex above, projecting
352 Mr. H. Seeley on two new Plestosaurs from the Lias.
over the base of the cephalothorax, and is armed on the upper
part and sides with about fifty sharp-pointed yellow spines;
two, much larger than the rest, are unequally forked, having a‘
short spine on “their outer side, and are of a red-brown colour,
the anterior surface of the prongs and a broad annulus at their
base being much the darkest; these spines -are placed trans-
versely on the most elevated part of the abdomen ; it is of a
pale dull-yellow hue, and has small black prominences distri-
buted over its upper part and sides; the former has a narrow
brown band extending from its anterior extremity to the spin-
ners, near the middle of which there are two short brown lines
curved towards each other, comprising in the interval between
them a pair of spines; and the latter are marked with dark-
brown transverse lines, bordered anteriorly with pale yellow ;
the under part is corrugated, and minute, depressed, brown
spots are disposed in rows in the curved transverse furrows ;
the sexual organs present a narrow transverse orifice, and have
a reddish-brown hue, that of the branchial opercula being yel-
lowish-brown.
I have much pleasure in associating the name of my valued
friend and correspondent, R. H. Meade, Esq., with this very
remarkable Spider, which forms a connecting link between the
species of the genera Epetra and Acrosoma.
XLI.—On two new Plesiosaurs, from the Lias. By Harry
Srey, F.G.S., of the Woodwardian Museum, Cambridge.
[Plates XIV. & XV.]
Tuomas Hawkins, Esquire, to whose zeal English science owes
many exquisitely wrought- out Reptiles of the Liassic seas, pre-
sented to the University of Cambridge a series of Saurians
which grace the walls of the Woodwardian Museum. Two are
Plesiosaurs, both of new species: one of them, already famous
for having the unanchylosed bones described by the late Lucas
Barrett in his paper on the Plesiosaurian atlas and axis, displays
limbs, ribs, all the dorsal vertebrae, and much of the neck; the
other and smaller fragment has a part only of the neck and back,
ribs, episternum, coracoid, s scapula, and clavicle.
The lar ger fossil seems to have been imbedded laying on its
back, with the neck swayed laterally ; but all the vertebrae now
rest on the left side, so that the neck is curved, as though the
head had been drawn back. Except the first four caudal verte-
bre, the tail is wanting, and many vertebre are absent from the
middle of the neck. Mr. Barrett considered the remains to in-
Mr. H. Seeley on two new Plesiosaurs from the Lias, 353
dicate a young animal; but, though probably not aged, there is
no evidence that it was immature.
The species hitherto undetermined, indicated by twenty-four
vertebrae between the last of the neck and the first of the tail,
and by eight carpal bones and five bones in the tarsus, is
different from every other yet characterized. So it will now be
described as
Plesiosaurus eleutheraxon*.
The atlas and axis are contrasted with the same bones in Plesi-
osaurus Etheridgii, and well figured in the ‘Annals of Natural
History’? for November 1858. ‘To these succeed eleven verte-
bra, which with their interspaces occupy 9% inches. The para-
pophyses for the hatchet bones occupy at first all the width be-
tween the epiphyses, but at the thirteenth they are found only
on the hinder half of the side of the vertebre. The neura-
pophyses soon extend to the whole length of the vertebree; the
zygapophyses are long, the neural spine compressed, wide, and
not very high. The sides of these cervical vertebre are flat-
tened, but the under surface between the hatchet bones is exca-
vated.
The third vertebra is ;°, inch long, and # of an inch wide
over the articular end, which is ;°; of an inch high. Subsemi-
circular in outline in front, being flattened above, it is moderately
concave, and, like all the cervical vertebree, has a bevelled border,
which is always of the same width. The articular surface is
oval in the sixth, which is nearly 2 of an inch wide and % high.
The thirteenth is = of an inch long.
Two inches’ interspace here separates the next three vertebrae,
which are noticeably larger, have long narrow venous foramina
between the small ploughshare bones, and extend over 33 inches.
The last of these vertebrae measures 24 inches from base to top
of the neural spe, which is much less high in front than be-
hind, and from front to back nearly as long as the vertebra.
After these there is a vacant imterspace of 11 inches; and
then the remainder of the spinal column is continuous to the
end, the thirty-five vertebree maintaining a nearly uniform size.
In the first seven the circular parapophysis rapidly ascends the
side, becoming elliptical, and at the eighth the articulation for the
rib, which there becomes very much larger, is entirely supported -
on the neural arch. ‘The sides are concave, so that the epiphyses
form a broad rim, and inferiorly they meet in a sharp angle,
which disappears with the first dorsal. The seven vertebre
measure 10 inches. The neural spine is now much longer, and
makes the height of the first of these seven cervicals 34 inches.
* @hevOepos, Lat. liber; dEav, Lat. avis.
Ann. & Mag. N. Hist. Ser.3. Vol. xvi. 24,
354 Mr. H. Seeley on two new Plesiosaurs from the Lias.
Each centrum is much higher than long. Thus there are, with
two scattered on the slab, twenty-seven cervical vertebra pre-
served; and if the interspaces represent the number missing
there must have sheet originally about tLe -six. The hinder
cervical ribs are 2 inches long.
In a former paper on Plesiosaurus macr opterus, I adopted as
the first dorsal that vertebra in which the rib rises entirely on
the neural arch—a view since sanctioned by Professor Owen in
his Monograph of Plesiosaurs.
The length of the dorsal and sacral region in this specimen is
2 feet 5 inches, and it includes twenty- four vertebrae, twenty-one
of which, measuring 2 feet 14 inch, have the ribs_ entirely
supported on diapophyses. The remaining three, with large
articular facets, may be considered pelvic. The first dorsal is
an inch anda quarter long ; but these vertebrae gradually become
a little shorter, and the last dorsal measures only 1;4,inch. The
venous foramina rise to the middle of the side, which is much
more concave than it was in the neck. The diapophysial articu-
lation is vertical, looks behind, is subelliptical, and in the first few
vertebree more than an inch deep; but it soon becomes smaller,
and more circular. The caudal vertebre are shorter than those
of the back, have flatter sides, and are marked by large heema-
pophysial pits. The ribs are very short, and, like those of the
hinder part of the neck, taper to an end in about 2 inches. The
four caudal vertebre with their interspaces measure 4+ inches.
As in the typical specimens of all the species with which this
one presents any affinity, it is the under surface of the limbs which
is displayed: those of the right side are zz sztu; but on the other
side only the femur and humerus remain, the latter showing the
upper surface. The fore limb is slightly the shorter, measuring
23+ inches, while the hind limb is 24 inches long.
The humerus measures 83 inches long, is 32 inches over the
radial end, and 13 inch over the greater part of the shaft. The
proximal end is a little contracted and bent backward, and the
rough condyle is moderately convex. The under surface is flat-
tened and rounded, and displays at an inch below the condyle a
slight oblique rough process, probably for attachment of the
latissimus dorsi. The anterior border is somewhat straight,
- being slightly concave below and slightly convex above; but the
posterior border is more deeply and regularly cupped. The
superior is rather flatter than the inferior surface, except at the
proximal end, which becomes elevated to a trochanteroid thick-
ening for the deltoid, subscapularis, and pectoralis muscles, and
makes the bone concave in length. The ulna is flattened, and
slightly reniform, and a little shorter than the radius, being
3+ inches long and 2+ inches wide, and 25%; inches long in
Mr. H. Seeley on two new Plesiosaurs from the Lias. 355
front ; the radius, more convex than the ulna, is contracted below.
It measures 21 inches over the top, 12 inch in the middle,
12 inch at bottom, and is 84 inches long. There are 8 carpal
bones—5 in the upper row, 3 in the lower row; they add
4 inches to the length of the limb.
Thephalanges and metacarpal bones, which are gracefully curved
backward, are in five rows, and measure 101 inches. There are
3 bones in the first row, 6 in the second, 9 in the third, 8 in the
fourth, and 4 in the fifth row: they have the usual flattened
hour-glass form, and vary much in length, the row usually named
metacarpal being the longest. The metacarpal of the first digit
is subquadrate, and not quite an inch long; the metacarpal of
the fourth digit is 13 inch long. The most massive metacarpal
is that of the fifth digit. The terminal phalange is always nar-
row at the distal end.
The extreme length of the femur is 81 inches: it has a large
hemispherical condyle, 14 inch deep and 12 inch wide. The
smallest diameter of the shaft is 1+ inch, and the diameter of
the distal end of the bone is 31 inches. The anterior side is
longer than the posterior side, but is less deeply cupped. The
under surface is more rounded than the under side of the
humerus, and has a stronger rugose thickening below the con-
dyle for the psoas muscle.
The tibia and fibula are much like the ulna and radius, ex-
cept that the fibula is more reniform.
The fibula is 2 inches wide, 22 inch long, and 2+ inches long
in front. The tibia is 24 inches at top, 14 inch in the middle
and 12 inch at the base, and 2+ inches long. The tarsus adds
2+ inches to the limb. There are 5 bones—d large ones under
the fibula and 2 small ones under the tibia, two large ones
being in the upper row, and the others in the lower row.
The phalanges and metatarsals are straighter than the corre-
sponding bones in the fore limb; they are 11 inches long.
There are 3 bones in the first digit, 7 in the second, 9 in the
third, 8 in the fourth, and 5 in the fifth. As in the wrist of the
fore limb, the metatarsals progressively get longer to the fourth ;
and, as in that limb, the fifth digit articulates with a tarsal of
the first row.
The ilium is near the proximal end of the femur, 4 inches
long, 1+ inch wide at the compressed spathulate end, and 3 of
an inch wide in the shaft. The proximal end is large, and has
two articular facets—one for the femur, the other probably with
the pubes.
Midway between the limbs is a triradiate bone, shaped some-
what like a Greek letter vy, and probably showing the upper
surface. It resembles the ischium, but corresponds exactly with
24%
356 Mr. H. Seeley on two new Plesiosaurs from the Lias.
Professor Owen’s account of the scapula. It is deeply cupped
for the pectoral vacuity, and measures 44 inches along the per-
pendicular and flexuous exterior side, which is elevated to a
ridge as usual, interior to which the inferior side is narrow,
oblique, and concave. Its anterior margin is convex, and
measures 51 inches. This anterior limb is much wider than the
lateral limb, which contracts in the middle. Both terminate in
an articular end, in each measuring 11 inch; the distance over
articulation to articulation is 34 inch. Where the two limbs meet,
they become prolonged into a short spur directed forward and
outward. I cannot regard cither this spur or the broad flat limb
as representing the clavicle. All analogy would indicate that, if
this species had such a bone, it was applied to the exterior flat
side of the scapula. Nor can I believe that the scapula had in
the animal the wonderful position which it has in Prof. Owen’s
restoration, fig. 93, ‘ Paleontology,’ where the left scapula appears
to me to be applied to the right side of the animal. Certainly in
this species the broad limb, which is there free and directed
backward, was directed forward and downward towards the other
scapula. Similar bones are figured in the scapular region of a
Plesiosaur by Mr. Hawkins in pl. 25 of the ‘ Great Sea-Dragons.’
This species nearly resembles P. dolichodeirus (Conyb.), and
is but little smaller than the example recently figured by Prof.
Owen in the Paleeontographical monograph.
The distinctive characters are, (1) that in this species the atlas
and axis are short and separate, instead of being long and united ;
(2) there are 24 vertebrae between the neck and the tail, instead
of 23; (3) the humerus is more slender, being longer; (4) the
bones of the forearm are wider ; (5) there are 8 carpal bones in-
stead of 7; (6) there but 4 bones in the fifth digit, stead of 8.
The more marked differences of the hind limb may be esti-
mated from the figure. From P. Etheridgit it is well distin-
guished by the more numerous cervical vertebrae, the 21 dorsals,
and larger size.
Nor does it approach any other species so near as to need a
minute comparison.
Plesiosaurus cliduchus*.
Professor Owen founded the Plesiosaurus rugosus on a charac-
ter which is by no means limited to that species; for the Plesiosaur
just described shows in the cervical region, only in a less degree,
roughnesses round the epiphyses; and the cervical vertebra of
that which is now to be described have the same character. But
Mr. Wm. Davies, of the British Museum, assures me that in
the typical specimen of Plesiosaurus rugosus the characteristic
* Kreis, Lat. jugulum; and éxa, Lat. habeo.
Mr. H. Seeley on two new Plesiosaurs from the Lias. 357
rugose border is present in the whole series of vertebree ; and
adds that the whole of the limb-bones, including the phalanges,
as well as other parts of the skeleton, are more or less rugose.
Our animal appears to have been mature, though not aged :
what remains of it rests on the right side.
In P. rugosus there are said to be about 35 or 386 cervical
vertebrae : but the only one described is supposed to be the 15th ;
and plate xiv. of the Monograph of Lias Plesiosaurs shows it
to be about two-thirds the size of those at the base of the neck.
Now, as the vertebre from the base of the neck in our specimen
are shorter than that from the middle of the neck in P. rugosus,
and as the lower dorsal vertebrae want the rugose margins, there
is strong probability that the species is new.
It would naturally seem a portion of a vertebral theory that, just
as certain vertebrae between the back and tail are reckoned pelvic
or sacral, so there might be some between the neck and back simi-
larly modified by relations to the fore limbs, which could only be
reckoned pectoral. At present there is no possible means re-
cognized for determining in Plesiosaurus the limits of dorsal
and cervical vertebree. Prof. Owen’s dictum in 18389 is different
from Prof. Huxley’s in 1858; and Prof. Owen, in 1861, lays
down a different law to that given by him in 1865. Indeed
the conditions seem to be different im nearly every species, the
difficulty being to know what to do with those vertebree where
the costal surface is passing from the centrum to the neural
arch, in which the ribs are extremely variable. The confu-
sion which has hitherto marked descriptions of this region
might be easily avoided by reckoning such vertebr pectoral,
and counting them separately, while it would introduce a new
character whereby to distinguish species.
There is the usual difficulty in this species in determining
the first dorsal. As in P. eleutheraxon, the costal surface in
the last few cervicals enlarges, becoming more elliptical, and
then in those vertebre where it is partly formed by the neura-
pophysis becomes smaller, and enlarges again in the back.
The first three vertebre are clearly cervical, and the first of
them measures—
From base of centrum to top of neural spine. . 44 inches.
The height of the centrum is .............. 14 inch.
ithe leneth of theicentrumy is’ i... w--<e se es 55
but it is longer at the top and at the bottom.
Meneth through thercentie sare nce sree oes ty Cas
§ From prezygapophysis to postzygapophysis is 2% inches.
b
The articular surfaces of the centrum are flat at the outer
part, and rather deeply cupped in the centre, where there is a
358 Mr. H. Seeley on two new Plesiosaurs from the Lias,
prominent mamillate eminence, as in P. Neocomiensis. The
sides are flat, and margined by wide, elevated, rugose borders ;
they meet beneath in a sharp mesial ridge. The subqua-
drate neural spine is rounded at the top, and from front
to back measures 1+ inch. The costal surface is circular. The
limit of the neurapophyses on the sides of the centra may be
clearly seen in the figure. The last cervical has the costal sur-
face divided by a horizontal groove.
Then follow three pectoral vertebrae. The costal surface has
now become narrow, elongated, slightly elevated, and marked
by a depression on the centrum, immediately under it: it is
not so near the anterior border of the vertebra. The neural
spine of the last of these is a little broader and higher than m
the succeeding vertebree.
The seventh centrum is unquestionably the first dorsal. The
costal surface is now entirely on the neural arch, is greatly
elevated and enlarged, oblique, and looks backward. ‘The
centrum is 15%, inch long. There are eleven consecutive dorsal
vertebrae, and three more indicated by continuous ribs zn situ.
The rib of the second dorsal is 102 inches long; but towards
the end of the series the ribs rapidly become short, and the four-
teenth is only 4 inches Jong. ‘The tenth and eleventh (which
are the only centra seen) show pinched-in sides, sharp articu-
lar margins, and the outer part of the articular surface flat.
The eleventh is 14 inch long, and about 2 inches high.
The episternum is a large bone with broad wings which
taper to a sharp point behind, and between which, in front,
there is a deep crescentic cup. It is flat and thin, measuring
mesially—from middle of cup in front to end of wedge behind
22 inches ; from horn of crescentic cup to end of wedge 44 inch ;
from middle of cup to end of lateral wing 84 inches.
What remains of the coracoid conforms to the usual type ; but
the scapula and clavicle are remarkable.
Two inches of the coracoid go to form, with 1+ inch of the
scapula, the deep glenoid cavity for the left humerus, measur-
ing 24 inches across.
On its under side the scapula sends down a sharp ridge, which
extends all along the straight, flat, and perpendicular anterior
side. The side interior to the ridge is narrow, oblique, and
concave, and, with the coracoid, forms the pectoral foramen.
What is preserved of it measures 5 inches along the exterior
ridge. From centre of the glenoid cavity to the pectoral foramen
is lt inch. Least distance from ridge to pectoral foramen is
14inch. The pectoral foramen seems to have been elliptical,
23 inches long and 1% inch wide.
But the clavicle, which does not appear to have been detected
Bibliographical Notices. 359
before, resembles nothing so much as the spine of a Mammalian
scapula. It is a V-shaped bone, placed exteriorly on the per-
pendicular side of the scapula, to which it is attached for
nearly 4 inches, widening from nothing in front to 12 inch
where the attachment ends behind. ‘The exterior border is
produced backward till the bone is 64 inches long; but the inner
border contracts in a curve so as to produce a free spine 22 inches
long, which overhangs the glenoid cavity.
The clavicle holds the same relative position that it has in
Ichthyosaurus.
My thanks are due to Mr. W. Farran for the interest taken
in producing the beautiful photographs from which the plates
are taken.
EXPLANATION OF PLATES.
PLATE XIV.
P. eleutheraxon (Seeley): a, ilium; £8, scapula; a’, distal end of left ilium ;
y, analogue of lesser trochanter on the under side of left femur ;
6, great trochanter, on the upper side of left humerus; e«, large
heemapophysial pit of first caudal vertebra; 1. cervical and pec-
toral vertebre ; 11. dorsal vertebree; 111. pelvic; Iv. caudal.
PLATE XV.
P. cliduchus (Seeley): a, episternum ; f, coracoid ; y, scapula; 4, clavi-
cle; ¢, pectoral foramen; ¢, glenoid cavity for humerus; 1. last
three cervical vertebra; 11. pectoral vertebra ; 111. dorsal.
BIBLIOGRAPHICAL NOTICES.
A History of British Ferns. By Enw. Newman. The Fourth or
School Edition. London: Van Voorst.
This does not pretend to be a scientific work, and, in that respect
as well as many others, differs from the author’s former books upon
Ferns; bat why it is called a “school edition”? we cannot under-
stand. A work for schools ought to possess something of an educa-
tional character; but the total omission of scientific arrangement is
not the most fitting mode of teaching. We should rather say that
this book is intended for the purely unscientific collector and culti-
vator of these beautiful and now popular plants. Even for such a
purpose we much doubt if it is desirable to neglect the chance of
conveying some knowledge of science, and transforming the mere
collector into something, however little, of a botanist. Doubtless
many who commence as collectors do really in time learn to desire
some scientific knowledge of the objects in which they take an in-
terest ; and therefore the book placed in their hands should give the
information that they at first require.
Mr. Newman brings forward the difference of opinion that exists
amongst botanists upon the division of Ferns into genera as a suffi-
360 Bibliographical Notices.
cient reason for neglecting all generic names and characters. This
may be convenient for the mere collector, but effectually excludes
him from the knowledge of the structure of the fructification ac-
quired by the study of the generic characters as given by any one of
the botanists to whom he refers. Because A. Gray, Roth, Babington,
and Hooker have been led to call the “same group of species” by
different names is no reason for not pointing out the characteristics
of the group, about which we believe that these authors do not ma-
terially differ.
It may be that the genera as at present accepted are to some ex-
tent artificial,—that is, if British Ferns are alone considered; but are
they quite so artificial if all Ferns are taken into account? And as
they are founded upon structure, is it not better, in a ‘school edi-
tion,” that even they, as the best that our present knowledge sup-
plies, should be placed before the reader? We certainly think so ;
for there is much education for the observing faculties in determining
these genera from a study of structure. The effect of neglecting the
generic names has caused only the specific terms to be used in this
book—a step, as it seems to us, in the wrong direction. Mr. New-
man says that ‘authors plume themselves on the number and length
of the Latin appellations they bestow on each species: no less than
86 Latin names have been assigned to filia foemina and its varieties,
and 47 to Scolopendrium and its varietics.> We think that this is
rather misrepresenting the matter. No botanist does so, although
gardeners do give useless names to an infinite number of forms.
What botanist cares for the 47 forms of Scolopendrium? If we look
at the books published by the above-named authors, we shall proba-
bly find not a single one of these ever-varying forms noticed by
name, but only the collective species characterized. Under the
the filix-feemina some two or three of the more marked and con-
stant forms are usually distinguished. ‘These botanists certainly do
not “plume themselves’? on the length or number of the names
given to the plants.
The author takes credit to himself for goimg back to the very
oldest names in all cases—a very good thing, doubtless, if done with
judgment ; but where is the advantage of hunting-up some obsolete
name when all the best botanists have agreed to accept one uniform
nomenclature? Why should we add to the confusion caused by
synonymy, by using Polypodium myrrhidifolium (Villars) for the
Cystopteris montana (Link), even if Villars really meant that plant
by that name? And why defend it by finding and using another
obsolete name, Polypodium montanum (Vogl), for the Polypodium
(or Aspidium or Lastrea) Oreopteris of all the best botanical au-
thorities of all countries? He says that as ‘‘ the principle of restor-
ing prior names is now universal in zoology, I can only regret it is
so frequently disregarded in the sister science.” We thought that
zoologists were very generally protesting against the attempt to
change the recognized names for the uncertain and ill-defined ones
of old authors. They think, and with much reason, that we are
only confusing the nomenclature, and unnecessarily adding to the
Bibliographical Notices. 361
difficulties of science, by so doing. We believed that it was generally
allowed that the nomenclature of botany was in a better state than
that of zoology. Many changes have been made recently ; but they
have been rendered necessary by an endeavour to bring the nomen-
clature of different countries into harmony, not by an attempt to
rake up and use obsolete terms which may be detected by a careful
search into ancient or obscure writers. Hooker has not acted in
the latter way, neither has Fries or any of the great authorities on
botanical nomenclature.
But enough of this. Under each species the author has given a
fairly good popular description of the plant, and some useful hints
concerning its best mode of cultivation, and also, occasionally, other
remarks of more or less interest. The Cystopteris dentata is appa-
rently combined with the C. fragilis (we say, apparently, for the
name is not mentioned) ; but the C. Dickieana is kept distinct. It
does not seem to us to be more constant or more distinct than several
of the 86 forms of Athyrium filix-fomina. The difference stated to
exist in their spores, however, is undoubtedly a point in Mr. New-
man’s favour. Recurvum is retained as the specific name of the
Lastrea emula or fenisecii of authors. If the oldest name is
adopted, it should be emula, which is much older than Lowe’s
feenisecti or Newman’s recurva. We do not find that Bree ever
called the species by that name; and even if he had done so, the
‘Hortus Kewensis’ is of much earlier date. In this case no name
has been universally adopted, and we are therefore fully justified in
reverting to the oldest. Mudtiflorum and spinosum are still retamed
for the LZ. dilatata and L. spinulosa of nearly all other authors.
Mr. Newman’s own glandulosum is kept distinct. He also sepa-
rates uliginosum from the <Aspidium cristatum of Smith. A.
remotum is likewise retained as a species. Mr. Newman con-
siders the Aspidium (or Polypodium) alpestre “very closely allied to
the common Lady Fern.’ Superficially considered, it seems so;
but we think that they are far from being closely allied in reality.
Asplenium acutum is separated from A. adiantum-nigrum. We give
no opinion upon this, although inclined to consider them forms of
one species. Asplenium Petrarche is stated to be a real native of
Ireland. We hope that it may prove to be so. \ Mr. Newman ad-
heres to the name of Hymenophyllum unilaterale for the H. Wilsont,
although, if we mistake not, it has been shown that they are not the
same plant. Ophioglossum lusitanicum is said to grow near the
Land’s End in Cornwall. Is it not the O. vulgatum B. ambiguum
which is found there, as it certainly is in the Scilly Isles?
Mr. Newman takes credit to himself for having done much towards
causing the present popularity of Ferns. It may be a questionable
point if he has done good or harm thereby. The result is that all
our most interesting Ferns are being uselessly extirpated in all tole-
rably accessible places,—uselessly ; for a very small number of those
pulled up are ever kept alive, or preserved as specimens, or studied
botanically. Those who visit Wales after some few years of interval
cannot fail to notice and deplore the result. The plants which they
362 Bibliographical Notices.
used to look at, not gather, on the mountains are hopelessly eradi-
cated; and they have to go to spots more fitted for the Alpine Club
than the botanist, to see the rarer or more interesting species. Hap-
pily there are a few such spots to which no “tourist” is likely to
attain, where our rare Ferns may perhaps be preserved for the grati-
fication of a future generation, and from which, when the present
fashion has passed away, they may spread to more accessible places
on the hills.
Chart of Fossil Crustacea. By J. W. Satter and H.Woopwarp.
With Descriptive Catalogue. Lowry & Tennant: London, 1865.
This is a large chart (2 feet 2 inches by 2 feet 9 inches) of the
genera of fossil Crustacea, showing the range in time of the several
orders, together with some recent Crustacean types analogous to
the extinct forms. The Chart is divided transversely into fifteen
zones of varying thickness, alternately dark and light, and corre-
sponding to geological stages; and vertically, across these bands, are
represented eight streams, varying in width and length, of Crustacean
forms,—some (as the Trilobites and Eurypterids) beginning early
and dying out in paleeozoic times, others (as the Decapods, Tetra-
decapods, and Xiphosures) beginning either in the Devonian or the
Carboniferous period and still flourishing ; whilst the Brachyurous
Decapods are first found in the Jurassic rocks. Among the lower
groups of Crustacea, the little Bivalved Entomostraca seem to have
had representatives for almost as long as fossiliferous strata take us
back in time; for their ‘“‘stream of history’? ranges upwards from
the chart’s lowest band (“‘ Cambrian”’ or “ Lingula-flags”’). The
Cirripeds also are included in this conspectus of Crustacean life ;
and the Catalogue explains that though in the Chart they range
only from the Rheetic strata upwards, yet good specimens of a trust-
worthy representative (Turrilepas) have of late been recognized by
Mr. H. Woodward among Upper Silurian fossils; and woodcut
figures are given at page 26.
The Chart fully answers the purpose proposed—supplying the
carcinologist with an eye-sketch of the Crustacean types and sub-
types, and enabling the paleeontologist to see the coexistent forms at
any epoch, and to trace at a glance the range of each group, whether
occupying the stage at once in force, as in the case of the Trilobites
and Eurypterids, or beginning with obscure traces or uncertain forms,
and whether giving place to incoming allies or continuing in true
succession to the present day. We need not wonder that the Chart
is good, well-devised, and conscientiously worked out ; for Mr. Salter
is devoted to Trilobites, Paleeocarids, et hoe genus omne, whilst Mr.
H. Woodward is as fond as anybody of Crabs and Lobsters, Prawns,
Shrimps, and “such small deer,” not only in the fresh but in the
fossil state; both also have made a study of the Eurypterids, and
both have command of the goodwill and help of their brother
paleontologists; while Mr. Lowry, the engraver, has long been
known for the successful application of his art to geology and fossils,
prompted by a genuine love of the science in all its branches.
Bibliographical Notices. 363
To show the use of the Chart to the student, we cannot do better
than adopt Mr. Salter’s explanation, as given in the Introduction to
the Catalogue :—
“This Chart is intended to show at a glance the development in
geological time of the different orders of the class Crustacea. Pic-
torial representations of the different groups of fossils, in their geo-
logical order, have been often before attempted, and the ‘Tabular
View of British Fossils,’ compiled and engraved by Mr. Lowry
(published by the Society for Promoting Christian Knowledge), has
been conspicuously useful in this way. But we do not know of any
published Chart in which the relations of the different members of a
family or group are preserved in an unbroken series, so as to show
the course taken by that family or order of animals in the successive
geologic periods, from their first appearance to the present time. It
has been found practicable (due regard being had to size and shape
of the sections of the Chart) to arrange the species nearly in their
right natural-history order. This could not always be done, inas-
much as there are often members of the same genns in two geological
divisions. It has been thought best, therefore, in the Catalogue, to
follow the same order as that of the plate, that the student may the
more easily, in studying each group, follow it from its commence-
ment to its close. The rule followed is to take each group in
ascending order, from its commencement in the lower strata to its
close, or to modern times. The Chart shows through what length
of time any genus existed ; and it will be observed that comparatively
few genera (except among the small Bivalve Crustacea) range through
more than one or two formations. When it is otherwise, the student
has only to look in the Chart, at the next overlying formation, for
the members of the genus he is occupied with (and they are arranged
as nearly together as the space and other circumstances will allow),
and then, turning to the Catalogue, he will be conducted by it over
all the genera peculiarly characteristic of the lower formation before
proceeding to those of a higher one. Thus, for instance, in the group
Lurypterida he will find, in the Lower Silurian, that only foot-
marks of a gigantic species are known; in the Upper Silurian, the
Hemiaspis and Bunodes are confined to that formation, while the
great Pterygotus and Hurypterus are figured as reaching through
two formations. Consulting the Catalogue, he will find that Ptery-
gotus problematicus is an Upper Silurian species, and that P. anglicus,
a very similar form, belongs to the Devonian formation. Again,
Eurypterus tetragonophthalmus is a Lower Devonian form ; but the
gigantic #. Hibberti belongs to the Carboniferous formation. One
or two in the Coal close the series. Here, then, the range of each
genus is extended through the formations to which it belongs. ‘But,
in this particular case only, the size of the animals is so great that,
instead of figuring P. anglicus and P. problematicus one exactly over
the other, a single large figure is made to do duty for both. In all
other cases, as in the genus Limulus on the right hand of the chart,
or the genus Homalonotus or Calymene among the Trilobites, the
successive species are placed one over the other, and the range of the
364 Bibliographical Notices.
whole genus in time is thus made evident. The student is recom-
mended to take each group of animals as indicated by the curved or
vertical lines of separation * by itself, as the object of the Chart is
to show him how each tribe or order has been gradually developed
and perfected, or otherwise, in its course. By taking, then, the
genera and species belonging to the lowest formation first, he will
the more readily see what changes have been introduced among a
particular set of animals; and having made himself thus master of
the separate groups, he will be able afterwards better to see their
mutual relations.’’
Genera Plantarum: auctoribus G. Benruam et J. D. Hooxer.
Vol. i. Pars 2. London, 1865.
We have much pleasure in announcing the publication of another
Part of this admirable work. It consists of 293 pages, and contains
the genera included in the orders Leguminosze, Rosacez, Saxifragez,
Crassulaceze, Droseraceee, Hamamelidez, Bruniaceze, Halorageze,
Rhizophoreze, Combretaceee, and Myrtaces ; and we are informed
that a third Part will complete the Polypetalous orders and the first
volume. Itis much to be desired that no great delay may attend
its publication.
It is scarcely possible to give any idea of the amount of labour
which has been expended upon this work, which must form a neces-
sary part of the library of every botanist. We have looked rather
hastily threugh the present part, and observe very few points re-
quiring notice. In Leguminosie the Genisteze, Trifolieee, and Loteze
are regarded as tribes of the Papilionaceee, and of equal rank with
Vicieee and Hedyracez ; and, amongst the genera, Sarothamnus is
combined with Cytisus, Ar throlobium with "Or nithopus, Ervum is
joined to Vicia, and Orobus to Lathyrus. The order Rosaceze is
retained entire, notwithstanding the apparently epigynous structure
of the Pomeze. Amongst its genera, Potentilla includes Sibbaldia,
Agrimonia includes Aremonia, Poterium includes Sanguisorba,
Pyrus includes Mespilus. The Grossulariaceze are combined with
the Saxifragese, and also the genus Parnassia. The genus Calli-
triche is placed in Halorageze, but Ceratophyllwm is considered to
constitute a Monochlamydeous order.
There are many other alterations made in the usual mode of
grouping, but we do not think it necessary to mention them.
Those enumerated are of the most interest to the British botanist,
as relating to the flora of his own country.
We have only to add that all botanists must feel anxious for the
early continuation of this very useful work, and express our hope
that its sale may be such as to encourage the learned authors to
proceed as rapidly with its publication as they properly can.
* The groups may be made more distinct by colouring the lines by
different paints or crayons.
Zoological Society. 365
PROCEEDINGS OF LEARNED SOCIETIES.
ZOOLOGICAL SOCIETY.
May 23, 1865.—John Gould, Esq., F.R.S., in the Chair.
A Revision oF THE GENERA AND SPECIES OF AMPHISBENIANS,
WITH THE DESCRIPTIONS OF SOME NEW SPECIES NOW IN
THE COLLECTION OF THE British Museum. By Dr.
JoHN Epwarp Gray, F.R.S., F.L.S., V.P.Z.S., etc.
Sir Anprew Smiru having kindly presented to the British Museum,
along with a number of other reptiles which he has described, the
types of his genus Monotrophis, which I had not before seen, and
two Amphisbzenians from Africa having been received from Mr. Wel-
witsch and from the collection of my late excellent and lamented friend
Dr. Balfour Baikie, and from Mr. Bates a species from the Amazons
which I believed had not hitherto been recorded in the Catalogue, I
proceeded to examine them; and, for the purpose of making the
comparison the more complete, I was led on to study all the speci-
mens of this tribe which we have in the Museum.
The natural result’of such an investigation was, that I was dis-
satisfied with the manner in which the species had hitherto been
arranged and described, and, after repeated examination, I have
reduced my observations to the following results :—
The determination of the species themselves, and the means which
a paper resulting from the re-examination and coinparison of all the
species in a large collection afford to a student, are much more im-
portant than any isolated description of the species regarded as new,
however detailed and particular the description may be; and ina
comparative review of the species of a group or order the distinctions
may be stated in a more condensed form.
The Amphisbeenians are very rarely collected; hence few species
are found in museums and noticed in systematic catalogues. This is
explained by their living almost exclusively in the nests of ants, and
being seldom seen by the casual observer. ‘There is reason to believe
that every warm country which has ants has some form of Amphis-
beenians. Until lately they were thought to be confined to Tropical
America, though one was described by Vandeli as occurring in Spain
as long ago as 1780; but his essay and the animal itself were alike
so little known to naturalists, that Professors Hemprich (in 1820)
and Wagler each described Vandeli’s species as new, the latter as a
South-American species. Professor Kaup described a species from
North Africa in 1830, and M. Gervais redescribed it as new in 1835.
MM. Duméril and Bibron have described a specimen in the Leyden
Museum from Guinea, Dr. Andrew Smith one as occurring at the
Cape, and Dr. Peters has added another from the east coast of Africa.
The number of African species is in this essay raised to seven. As
yet none have been received from Asia Proper; but Sir Charies
Fellows brought from Xanthus the same species that is found In
Spain, Portugal, and North Africa.
366
The following table shows the geographical distribution
Zoological Society :—
species here recorded :-—
14.
EASTERN HEMISPHERE.
Fam. Trogonophide.
Trogonophis Wiegmanni. North Africa.
Fam. Amphisbenide.
Blanus cinereus. Spain, North Africa, Asia Minor.
Amphisbena? violacea. East Africa.
Cynisca leucura. Guinea.
Baikia africana. West Africa.
Tribe Cephalopeltine.
. Monotrophis capensis. South Africa.
Dalophia Welwitschii. West Africa.
WESTERN HEMISPHERE.
Fam. Chirotide.
Chirotes lumbricoides. Mexico.
Fam. Amphisbenide.
Amphisbena alba. Brazil.
A. americana. British Guiana.
A. Petrei. Brazil.
A. vermicularis. Brazil.
. A. Darwinii. Monte Video, Buenos Ayres.
Bronia brasiliana. Brazil.
Sarea ceca. West Indies.
Cadea punctata. Cuba.
Anops Kingii. Buenos Ayres.
Fam. Lepidosternide.
Lepidosternon microcephalum. Brazils.
L. Grayit. Tropical America.
L. phocena. Buenos Ayres.
Tribe Cepholopeltine.
Cephalopeltis lepidosterna. Brazils.
of the
The rings of oblong scutella on the skin are in most species inter-
rupted on the sides, and in some species also on the vertebral line ;
these interruptions forma more or less wide depressed groove on the
surface of the body, and are called the lateral and dorsal lines.
The skin at this interruption is usually marked at each transverse
ring with two oblique grooves, which form a cross and divide the
space into four minute triangular shields; in some cases, where the
line is wider and less sunken, the transverse ring of shields is only di-
vided at the sunken line by a single oblique groove caused by the
Dr. J.B. Gray onthe Amphisbenians. 367
tapering end of one of the oblong shields going before the end of the
other. Sometimes this is the case with the dorsal line, and not with
the lateral one. In some species, instead of-only the four triangular
shields in the lateral line, the shield between the cross groove is di-
vided into several minute scale-like shields.
In some of the larger species, as Amphisbena alba, some of the
rings of shields are marked with an oblique groove crossing several
shields, dividing each of them into two parts; but these seem to be
mere individual variations occurring on several parts of the back
of some specimens, and not present in others.
Duméril and Bibron give the number of the teeth as one of the
specific characters. I have not been able to verify their observations ;
they give the following as the number. There seems to be always an
odd number of intermaxillary teeth, the middle one being usually
large.
. 4.5.4 13
' ee ans
Trogonophis Wiegmannt .......+-+++++ Sig 18
: ‘ Sess
Chirotes Caniculatus’ << cere cc sian vine 6.6 —b
i : 5 55 1G,
Amphisbena americana et A. alba,..... Se ie
. Rey 5
J MOHALI ep elo BOT De oPawiatstenrate a 16
abel 4.7.4 15
—— Darwinit ...... 0 crc eee eceee., SSG
5.5.5__15
IST CONCCEED Melee aici icy-Neboi a aietat eds biel me aisle elias ag
ATA 1S
Cadeg PUnelala, ais see sate seks gee Son ee
Anops Kingii ..— 4.7.4 15
Blanus cinereus “es © 8 @ eee oe (Pye. 21.8.8. 8) 658 Chee) 14
Fam. 1. TROGONOPHID.
Head oblong, depressed, rounded below; nostrils lateral, in large
nasal shields ; teeth conical, on the edge of the maxilla. Body
cylindrical, covered with rings of uniform, elongate, oblong, four-
sided shields, without any sternal disk ; lateral line sunken, narrow,
covered with a few minute scales; preanal pores none; tail conical,
acute.
Glyphodermes acrodontes, Dum. et Bibr. Erp. Gén. v. 467.
TroGonorpuis, Kaup, Isis, 1830, p. 880.
Head oblong, depressed; nasal shields large, united by a short
straight edge, behind the large triangular convex rostral ; crown with
two pairs of shields ; temple with many small shields ; upper labial
plate moderate ; lower labial shield larger, with a series of large chin-
shields on each side, and a central gular one. ‘Tail conical, acute ;
preanal pores none.
The skull of this genus has been figured by Dr. Kaup in his paper
in the ‘Isis’ quoted below.
368 Zoological Society :—
TroGonopuis WieGMANnt, Kaup, Isis, 1830, p. 880, t. 861;
Féruss. Bull. Sci. Nat. xxv. 203, 1831; Dum. et Bibr. Erp. Gén. v.
470.
Amphishena elegans, Gervais, Bull. Sci. Nat. de France, 1855,
. 135; Mag. Zool. 1835, class 3. t. 11 (details not good).
Hab. Tangier (Fraser, B.M. 1848); North Africa (B.M. 1846) ;
Algeria (Duméril, B.M.).
This animal was first described by Dr. Kaup, who showed that
the teeth of it were placed on the edge of the jaw, as in the genera
of the family Agamide, which are all confined to the eastern he-
misphere and Australia; while all the other genera of the order
that have been examined have the teeth on the inner side of the jaw,
as in the family Lywanitde, which is restricted to the New World.
It was afterwards described by M. Gervais; and even when Dr.
Kaup had informed him, after inspecting the specimen, that it was
the same as he had previously described, he still regarded it as new,
because he said the skull did not agree with Dr. Kaup’s figure:
but this was a mistake. Dr. Kaup figured the skull of Tr ogonophis
and of an Amphisbena for the sake of showing the difference between
them; and M. Gervais must have compared his animal with the
wrong figure.
Fam. 2. Carrotip#, Gray, Cat. Tortoises, &c., B. M. 74.
Head depressed, rounded on the sides; nostrils on sides ; teeth on
the inner side of the maxille. Body cylindrical, covered with rings
of uniform oblong four-sided shields, and with two short weak front
limbs, provided ath five subequal clawless toes ; lateral line sunken,
covered with scales; preanal pores distinct. Tail cylindrical.
Currores, Duméril; Bimanus, Oppel; Chamesaura, Schneid.
Characters those of the family.
CHIROTES CANICULATUS, Cuvier.
Chirotes lumbricoides, Gray, Cat. Tortoises, &c., B. M. 74.
Hab. Tropical America, Mexico (B. M.).
Professor J. Miiller has figured the skeleton and skull of this
animal.
Fam. 3. AMPHISBHENIDA.
Head oblong, rounded below; nostrils lateral, in nasal shields ;
teeth conical, on the inner edge of the maxillee. Body cylindrical,
covered with rings of uniform, elongate, four-sided shields, without
any sternal disk ; preanal pores distinct ; lateral line linear, sunken,
with a few small scales. Legs none. Tail cylindrical, rounded at
the end.
Tribe 1. Ampaispanina. The head depressed, rounded on the
sides in front; nostrils on the upper part of the sides of the
head.
Dr. J. E. Gray on the Amphisbenians. 369
A. Lateral and dorsal lines distinct, sunken, covered with small tri-
angular scales; nasal shields large, square, lateral, forming
part of the edge of the upper lip, and separated in front by a
broad, square, convex rostral shield.
BLANws.
The rostral square, convex ; the nasal shields large, forming part
of the edge of the upper lip; the crown with a large pentagonal
frontal shield and two pairs of square shields behind it; eye-shield
triangular, between upper edge of the front labial shield and the
frontal. ‘Temples covered with a series of squarish shields ; labial
shields large, the hinder smallest ; the lower shields without any chin-
shield between them and the gular one. Tail rather tapering, blunt ;
preanal pores distinct.
BLANUS CINEREUS, Gray, /. c. 72.
Amphisbena cinerea, Vandeli, Mem. Acad. Lisbon, i. 1780.
A, oxyura, Wagler.
A. rufus, Hempr.
Blanus rufus, Wiegm.
Hab. N. Africa, Tangier (Fraser, B. M.); S.W. Europe, Spain
(Vandel, 1780) ; Oporto (Allen).
M. Gervais (Mag. Zool. 1837, class 3. t. 10) gives a figure of 4.
cinerea; but the details of the head do not perfectly agree with
our specimens; perhaps this may be from want of care in the artist.
The number of pairs of plates on the occiput varies from two to
four.
B. Lateral lines linear, distinct, sunken; dorsal none, or very in-
distinct ; nasal shields not forming part of the upper lip.
a. Nasal plates large, extending across the muzzle, united by a long
straight suture, or united into one cross band ; the rostral trian-
gular, under front edge of nasals ; crown with two pairs of
broad shields.
AMPHISBAENA.
Head depressed, broad, and rounded in front ; frontal plates with
one or two pairs of rather smaller similar plates behind them ; pre-
anal pores eight.
* Head depressed, broad ; occiput covered with square shields, like
the body ; preanal plates numerous.
1. AMPHISBEZNA ALBA, Linn. ; Gray, Cat. Tort., &c., B.M. 70.
Body thick, one-coloured, with only one pair of plates behind the
frontal plates ; occiput shielded like the body.
Hab. Brazil.
Varies in the size and form of the hinder pair of frontal plates ;
preanal pores eight, often seven.
Ann. & Mag. N. Hist. Ser.3. Vol. xvi. 25
370 Zoological Society .—
There is a specimen in the British Museum sent by Mr. Brandt
under the name of 4. Darwinii.
2. AMPHISBENA AMERICANA, Schreb.; Gray, Cat. Tort., &c.,
B. M. 70.
Body rather thick, black, varied ; two or more pairs of plates
behind the frontal plates.
Amphisbena fuliginosa, Linn.
A. vulgaris, Laur.
Hab. Tropical America: British Guiana; Berbice; Demerara
(BM):
The labial shields vary in number and shape ; the shields behind
the frontal vary in number and size, but they are generally in pairs
and subsymmetrical.
Gervais figures the skull of the species (Ann. Sci. Nat. 1854, xx.
t. 14. f. 4).
** Head rounded, narrow ; preanal pores and preanal shields ten
or twelve.
3. AMPHISBZNA PETRI, Dum. et Bib. Erp. Gén. v. 487; Gray,
oe. 80.
Hab. Brazils (Mus. Paris).
**E Head rounded, narrow, rather produced in front; crown-shields
large; occipital shields polygonal. Body slender, one-coloured ;
preanal pores two or four; preanal plates six, middle ones
elongate.
4. AMPHISBANA VERMICULARIS, Dum. et Bibr. Erp. Gén. v.489;
Gray, U.c. 71.
Hab. Brazil (Dr. Gardner, B. M.; Mus. Paris); Porto Bello
(Capt. Austin, R.N., B. M.: head in a very bad state).
5. Ampuisp£na ? Darwin, Dum. et Bib. Erp. Gen. v. 491;
Gray, 7..¢.0711..
Hab. Monte Video (Mr. Darwin; Mus. Paris).
6. AMPHISBHNA ?v10oLAcEA, Peters, Berlin Monatsb. 1854,
p- 620; Wiegmann, Arch. 1855, p. 49.
Hab, Kast Africa, Inhambane (Peters).
This species is unknown to me; it is without a single frontal
shield, and has four preanal pores and visible eyes.
Cynisca, Gray, Cat. Tort., &c., B. M. 71, 1844.
Head flat, narrow; nose conical, four-sided, rounded at the end ;
rostral triangular ; nasal plates very large, soldered together, cover-
ing the front of the head ; crown with a small frontal and a pair of
parietal shields ; eyes distinct ; temples and occiput with large shields.
Body very slender; lateral line distinct. Tail cylindrical, elongate,
truncated ; preanal pores numerous.
Dr. J. KE, Gray on the Amphisbenians. 371
Cynisca LEucuRA, Gray, l. c. 71.
Amphisbena leucura, Dum. et Bibr. Erp. Gén. v. 498.
A. macrura, Schlegel, Mus. Leyden.
Brown ; end of tail white.
Hab. Guinea (Mus. Leyden) (not Guiana, as stated by mistake in
the Catalogue).
b. Nasal shields small, separate above, on the side of a large swollen
rostral shield.
BrRonia.
Head ovate, rather convex ; rostral shield very large, hemisphe-
rical, with the small nasal shields inserted in notches on its hinder
edge, which is placed over the front labial; crown convex, rounded
on the side, covered with two pairs of shields ; the front pair square,
the hinder smaller, triangular, with a small triangular occipital shield
on its outer side; eye-shield triangular; labial shields — the
second upper and front lower large ; gular shield single, square, with
a cross series of shields behind it. Body cylindrical ; lateral line
well marked; the dorsal shields elongate, narrow ; the ventral ones
rather broader, smooth ; preanal pores four ; the preanal shields six or
eight, the central pair largest, the lateral ones very small. Tail blunt.
BRONIA BRASILIANA,
Pale brown; dorsal shields with a dark central spot.
Hah. Tropical America; Santarem, on the Amazons (Bates, B.M.).
Fig. 2. Fig. 1.
Bronia braziliana.
C. Lateral and dorsal lines not defined, or the lateral line only
visible on the hinder part of the body ; rostral shield small ;
nasal shields far apart, small, placed on the side of the high
rostral.
SarEA, Gray, Cat. Tort., &c., B. M. 71, 1844.
Head conical; rostral narrow, higher than broad, rounded in
front, placed behind the triangular nasal ; crown with two pairs of
shields, the front largest, elongate, the hinder trigonal ; eye-shield
triangular ; the labial shields = ; the second upper and lower labial
shields very large, the others smaller; with one large gular plate.
25%
372 Zoological Society :—
Body slender; the dorsal seutella square, as long as broad, with a
dark central dot ; two central longitudinal series of ventral seutella
broader than long, smooth, white ; the lateral line very indistinct,
searcely visible except on the hinder part of the body ; preanal pores
four; preanal shields six, square. The eyes are slightly visible
through the shields.
SAREA C&CA, Gray, Cat. Tort. &c., B. M. 71.
Amphisbena ceca, Cuvier, R. A. 773; Dum. et Bibr. Erp. Gén. v.
492.
Hab. West Indies, St. Thomas’s (4. H. Riise, B. M.).
The specific name is not characteristic, as the eyes are as much
seen through the shield as in many Amphisbeenians.
Capea, Gray, Cat. Tort., &c., B. M. 71, 1844.
Head conical ; rostral narrow, higher than broad, truncated at the
tip, convex in front ; nasals ovate, lateral ; crown with two large, tri-
angular shields ; frontal with a small linear shield on each side of it ;
and two pairs of square occipital shields, the hinder pair smaller ; eye-
shield rhombic; eyes hidden; labial shields = subequal, middle
one in each lip largest ; temples covered with square shields ; gular
plate single, elongate. Body cylindrical; lateral line very indistinct,
scarcely to be distinguished except on the hinder part of the body ;
shields of the back square, of the under surface rather wider, but
scarcely wider than long; preanal pores four; the preanal shields
six, central ones elongate.
Capra punctaTa, Gray, /.c. 71.
Amphisbena punctata, Bell, Zool. Journ. ii. 236, t. 20. f. 2.
A. ceca, La Sagra, Cuba, 195, t. 21.
Pale brown, dotted and varied with deeper brown.
Hab. Cuba (WV. S. Macleay, B. M.): the specimen described by
Mr. Bell.
Tribe 2. ANoprntna. Head compressed, keeled on the sides in
front ; the nostrils lateral, on the under side of the keel.
A. Lateral line distinct and impressed.
Awnoprs, Bell, Proc. Zool. Soc. 1833, p. 99; Zool. Journ. v. 391,
t. 16. f.1; Gray, Cat. Tortoises, &c., B. M. 72, 1844.
Lateral line distinct, sunken; preanal pores ‘‘none”’ (Bed) ;
“four ?? (Dum. § Bibron).
Anors Kineau, Bell, Proc. Zool. Soc. 1833, p. 99; Zool. Journ. y.
391, t..16. £21); Gray, Cat: Tortoises) &e., BAM 72:
Amphisbena Kingii, Dum. et Bibr. Erp. Gén. v. 497.
tab. S. America (P. P. King; Charles Darwin) ; Buenos Ayres
(D’ Orbigny, Mus. Paris).
Dr. J. E. Gray on the Amphisbenians. 373
I have not been able to examine this genus, which was described
by Mr. Bell from a specimen brought from South America by Capt.
P. P. King, R.N.; it is described in more detail by MM. Duméril
and Bibron from specimens obtained by Mr. Charles Darwin.
The existence of the lateral line, which, according to Mr. Bell, “is
more distinct than in Aimphisbena, though less so than in Chir otes,”
and the account of the plates of the head as described by MM.
Duméril and Bibron, show that it must be distinct from the follow-
ing, which comes from Africa.
B. Lateral line none, or only very slightly visible on the hinder
part of the body.
BaAIkKiA.
The head compressed, elevated; rostral plate very large, com-
pressed, forming an arched crest from the mouth to the forehead,
with a groove on the hinder part over the nostrils ; crown with two
pairs of band-like shields behind the upper edge of the rostral, the
front pair narrow; eye-shield very minute ; eye invisible; temples
with two small shields ; the upper labial shields 3—3; the second
upper large, produced, keeled on the side; the hinder, under the
temporal shield, larger, square ; lower lip covered with a single large
shield on each cide! “separated by a square inferior rostral acid nad
by two small gular plates placed one behind the other ; nostrils large,
lateral, under “the edge of the keel of the frontals. Body and chest
covered with rings of similar oblong square shields; preanal pores
2—2, separated by acentral shield. Tail cylindrical, rather conical
at the tip.
In spirits the skin is loose and inclined to form a fin-like fold,
sometimes on one and sometimes on another part of the body, with
a central longitudinal ventral groove, without any appearance of a
lateral line.
BAIKIA AFRICANA.
Hab. West Africa (Dr. Balfour Baikiec). B.M.
Fig. 4. Fig. 3
Baikia africana.
Fam. 4. LepIpOSTERNID®.
Lepidosternide, Gray, Cat. Tortoises, &c., B. M. 73, 1844.
Head oblong, depressed, with a short horizontal keel in front ;
374 Zoological Society :—
nostrils in shields, under the keel of the rostral shield ; teeth conical,
on the inner side of the maxilla. Body cylindrical, covered with
rings of oblong four-sided shields ; the sternum with a disk formed
of differently shaped shields ; preanal pores distinct.
Tribe 1. LeprposrerninA. Head conical, covered with symme-
trical polygonal shields ; the pectoral disk covered with many
polygonal shields placed in oblique lines ; the dorsal and lateral
lines well marked, broad, smooth, formed by the overlapping
of the narrow ends of the sections of the rings. America.
LEPIDOSTERNON.
Head conical, covered with three pairs of symmetrical and a ver-
tebral shield; rostral shield large, broad, rounded in front; the
pectoral shield formed of regular, nearly equal, symmetrical rhombic
or six-sided shields, sometimes united into long shields which are
not symmetrical.
* Sternal plates of central series united into elongated bands.
Lepidosternon.
1. Lep1pOSTERNON MICROCEPHALUM, Wagler, Serp. Bras. 70,
t. 26; Miller, Tiedem. Zeitsch. 1832, iv. t. 22. f.4; Dum. et Bibr.
Erp. Gén. v. 505; Gray, Cat. Tortoises, &c., B. M. 73.
“© T,, macrocephalum, Miller’”’ (fide A. Smith).
Amphisbena punctata, Neuwied, Abh.
Lepidosternon maximilianus, Wiegmann.
Head short, broad; the vertebral plate broader than long, six-
sided ; frontal short, broad, band-like ; parietal small, square ; ocular
higher than broad.
Hab. Brazil, Rio (Dr. Gardner, B. M.).
The specimen in the British Museum has the shields on each side
of the central line of the sternal disk united into an elongate shield,
which is not symmetrical on the two sides, and appears like an acci-
dental peculiarity.
In the Free Museum at Liverpool there are two specimens of this
species, obtained by Mr. Jobert in Brazil. They are similar, but
show that the sternal plates are liable to coalesce and form larger
plates in an unsymmetrical manner.
In the larger specimens the first series of sternal plates on each
side of the central line are united into longitudinal shields, which
are not of equal length. The series of plates on the outer side of
them are separate, but not quite symmetrical.
In the other specimen, which is rather smaller, the first series of
sternal plates on the sides of the central line, and the second series
on the outside of it on the right side, and second and third series
on the left side, are united into longitudinal parallel plates, which are
of unequal length, the two central ones being the longest, and the
two outer plates on the left side much longer than the outer one on
the right side.
Dr. J. E. Gray on the Amphisbenians. 375
The head-shields in both these specimens are exactly alike, the
central crown-shield being much broader than long ; the hinder pair
of frontal shields rather shorter than the front pair, and narrower on
the central edge by the angular front edge of the crown-shield ; the
two pairs of occipital shields are shorter than broad, the hinder pair
being the smaller, and in one instance coalesced on one side with the
plate of the front pair.
They all have a brown spot on the centre of each of the dorsal
sceutella.
** Sternal plates all separate, symmetrical. Sphenocephalus,
2. LeprIposTERNON Grayil, A. Smith, MS. Brit. Mus.
Head rather short, broad; the vertebral plate hexagonal, elon-
gate, as long as broad; the frontal plate very short and broad ;
the parietal shields oblique ; the occipital much longer than broad ;
temporal shields larger on the side of the occipital; plates of the
sternal disk symmetrical, in oblique diverging lines.
Hab. South America? (Brit. Mus.).
Sternal disk formed of four diverging lines of uniform, similar-
sized, symmetrical shields; the shields on the central line smaller,
being divided down the centre by a straight suture.
In 1848, Dr. (now Sir Andrew) Smith sent to the British Museum
this specimen, with the name of Lepidosternon Grayii, informing me
that he had described (or intended to describe) it in the ‘ Proceedings
of the Zoological Society’ under that name. I cannot find that it
has been so described, nor can I find any notice of it in any other
work, though it is very like, but evidently distinct from, the Lepido-
sternon phocena of Duméril and Bibron, figured by M. d’Orbigny
(Voy. Amér. Mérid.).
Fig. 6.
Lepidosternon Grayii.
3. LEPIDOSTERNON PHOCENA, Dum. et Bibr. Erp. Gén. v. 507;
Gray, Cat. Tortoises, &c., B! M. 73; D’Orbigny, Voy. Amér. Mérid.
Rept. t. 6. f. 7-10.
Head broad ; the vertebral plate elongate, small, larger than broad,
acute at each end ; the frontal and occipital plates large, shorter than
broad, the frontal the largest ; the parietal plates short, broad, band-
like; the plates of the sternal disk uniform, symmetrical, oblique.
Hab. Buenos Ayres (Bridges, B. M.: stuffed).
376 Zoological Soctety.
M. d’Orbigny’s figure is very like the preceding species ; but. the
head is represented shorter, the frontal plates are rather larger. The
long occipital shield of that species is here represented by two pairs
of square shields, as if the large plate of the former species were
divided across; it also appears to be a shorter, thicker species.
The stuffed specimen which we received from Mr. Bridges agrees
with the figure in all these particulars ; but the head appears rather
larger, perhaps from its being rather distorted in the preparation.
Both the figure and the specimen belong to a species evidently
very distinct from ZL. Grayii, and much thicker.
Tribe 2. CepHaLopreLtTina. The head depressed, covered above
with a single simple or transversely divided, flat, horny, nail-like
shield ; pectoral disk formed of elongated, symmetrical shields ;
the dorsal and lateral lines very narrow, indistinct, except near
the hinder part of the body.
A. The pectoral disk formed of large, diverging, unequal, polygonal,
symmetrical shields ; crown-shield divided into two by a trans-
verse suture. America.
Creenarore tis, J. Miller, Tied. Zeitsch. fiir Phys. 1831, iv. 269.
Head covered with two large shields, the front one smaller ; the
sternal disk of eight or ten large shields, the two central pairs par-
rallel, one in front of the other; the lateral pairs diverging.
CEPHALOPELTIS SCUTIGERA, Gray, Cat. Tort., &c., B. M. 73.
Cephalopeltis lepidosternon, Miller, J. c. t. 21. f. 6 (skull), t. 22.
f. 5 (head).
Amphisbena scutigera, Hempr. Naturf. Freund. zu Berlin, 1820,
. 127.
Lepidosternon scutigerum, Dum. et Bib. Erp. Gen. v. 509.
Cephalopeltis Cuvieri, Miller (fide Dum.).
Coleopeltis Cuvieri, J. Miller (fide A. Smith).
Hab. Brazils (Miller).
B. The pectoral disk formed of six or eight elongate, longitudinal,
parallel shields; head-shield single. Africa.
Monorropuis, A. Smith.
The head covered with a single nail-like shield, without any slits
on the hinder part of its side edge; the rostral plate between the
nasal plates, transverse, four-sided, broader than high; the shields
of the sternal disk regular, broad, and truncated in front ; the rings
of shields in front of the sternal disk formed of regular square shields,
like those of the rest of the body.
Monorropuis capensis, A. Smith, Zool. S. Africa. Rept. t. 47
(white ; pink when alive).
Hab. S. Africa (B.M.). The type specimen of Sir Andrew Smith.
Miscellaneous. 377
Dr. Peters records Monotrophis capensis as found in Mozambique;
but on comparison it may prove a distinct species. In my notes I
have a reference to Lepidosternon sphenorhynchum, Peters, MS., as
an East-African species, but I cannot find it described or noticed
anywhere. Can it be the name Dr. Peters gave to his Monotrophis
before he discovered it had been described by Sir Andrew Smith?
DALOPHIA.
The head covered with a single nail-like shield, with a linear slit
on the hinder part of its side edges; the rostral plate small, trian-
gular, with the point upwards between the nasal plates; the shields
of the sternal disk rather irregular, but symmetrical, each with an
Fig. 8. Fig. 7.
Dalophia Welwitschti.
acute front edge; the rings of shields in front of the sternal disk
formed of unequal but symmetrical polygonal shields.
Datoenta WELWITSCHII.
Monotrophis capensis, Giinther, MS. B.M. (not A. Smith).
Pale brown.
Hab. Angola; Pungo Andongo (Welwitsch). B.M.
MISCELLANEOUS.
Dr. Sturm’s Collection of Objects of Natural History in Nuremberg.
Tue families of the late brothers Dr. J. H. C. F. and Dr. J. W.
Sturm wish to dispose of the above collection, which consists of the
undermentioned four divisions :—
I. Birds.—Of these there are 1700 species, with 2700 specimens:
1600 of these specimens were stuffed by the master hand of the late
Dr. Fr. Sturm, and set up by him in 557 glass-cases ; the remainder
consists of well-preserved skins.
A specimen of almost every species of bird is to be found, and of
several species there are numerous specimens. It would be hardly
possible to meet with another collection of specimens so well pre-
served or so artistically prepared. The 100 species with 253 speci-
mens of Humming-Birds, as well as the 26 species with 62 specimens of
Rhamphastides, deserve especial notice. The latter served Dr. Sturm
378 Miscellaneous.
as a basis for the monograph in which he has described them. Of
other groups, as, for instance, of Pigeons, Parrots, and Owls, the
specimens are scarcely less numerous.
I]. Birds’ Nests and Eggs.—Seventy-seven nests of European
Birds, with 1597 eggs; seventy-five nests of non-European Birds,
with 769 eggs.
III. Insects.—This division contains nearly 23,000 species, with
70,000 specimens, and is perhaps the largest private collection in
Germany. The order of Beetles, of which there are 16,640 specimens,
is the most richly represented. Besides the Beetles there are also
numerous specimens of the following orders, viz. :
Eliymenoptera cere cites ae ni 2193 in number.
Neuroptera.. .. Sec OO 3
Lepidoptera (non- European) .. 413 a
Lepidoptera (Kuropean)...... 800 6
Diptera cic h-i>: tyes LOSS +
Henintera; sweet ee 1439 .
SPIGeESS fais sesamin. hiaek BOOS ;
COMMONS 19 cymes ci vere Send 68 3
IMiyiapodas oy .ses eine bere: oie ey AO -
IV. Terrestrial, Fluviatile, and Marine Shells.—Land and Fresh-
water Shells, 13,000 specimens; Marine Shells, 2500 specimens.
This division may be also considered to be one of the richest of its
kind, and it contains many original specimens discovered by Say,
Adams, and other scientific men. A large number of wax models of
Land-Snails, prepared from nature by the late Dr. Fr. Sturm, deserves
particular mention, as no similar collection of models is perhaps to
be found elsewhere. Of almost all classes of the animal kingdom
there are specimens enough to form a good nucleus for anyone
wishing to commence a Collection.
This collection has been made use of for a long time in the com-
pilation of several works, and it has enjoyed a high degree of favour
because it contains the new and rare specimens deeenthel and illus-
trated in the works of Dr. Sturm himself. On this account, as well
as on account of its extent, Prof. Burmeister has pronounced it to
be “a very first-rate scientific collection.” Prof. Leiblem in Wurz-
burg, Prof. von Siebold in Munich, and Dr. Will in Erlangen, may
be quoted as University Professors sine have also expressed them-
selves in the very highest terms of the worth of the collection.
Should there be no purchaser for the whole collection, each divi-
sion of the same will be sold separately.
On the Existence of Liquid and Solid Matters in the Trachean
Vessels of Plants. By M. T. Lestrsovupots.
As facts in opposition to the arguments adduced in support of the
opinion that the trachean vessels of plants are aériferous, the author
indicates that vessels may appear empty because the liquids contained
in them are perfectly limpid, or they may lose their fluids by age,
Miscellaneous. 379
and will then emit bubbles of air; finally, the experiments cannot be
regarded as conclusive if made upon branches separated from the
trunk, as then the air would easily find its way into the vessels.
Nothing can be concluded from the circumstance that the walls of
these organs are not thickened by deposits, as many originally suce-
culent organic elements are in the same case.
If liquids circulate in non-vascular plants, this only proves that
the vessels are not the sole organs of circulation; and although
water charged with nutritive substances enters by spongioles which
contain no vessels, this does not prove that it does not penetrate
subsequently into the latter organs.
As regards the course followed by the pollinie granules, no con-
clusion can be drawn from it, as there is not the least relation be-
tween the materials of fecundation and the sap.
Tf we have no demonstration that the trachean vessels are exclu-
sively destined to the transport of gases, we find nothing more con-
clusive in the arguments advanced by Mirbel and Schultz, and other
authors, who hold the opinion that these vessels serve for the circu-
lation of liquids. They say that we see in them bubbles -of air,
which would not be visible if they were not circumscribed by a
liquid—that the absorbed water diffuses itself so rapidly in plants
that it cannot but follow the direct courses presented by these con-
duits—that when the branch of a tree is placed in a coloured liquid,
the latter ascends in the vessels, into which it also penetrates even
when absorbed by the roots—lastly, that as the elaborated sap cir-
culates in the laticiferous vessels, the ascending sap must ascend by
analogous ducts, &c.
But the liquids which surround the air-bubbles may have pene-
trated into the vessels when the observed tissues were cut into thin
sections; those which ascend into the branches may be introduced
by the gaping orifices of section ; the colouring-matters which tinge
the vessels may only impregnate their walls externally ; lastly,
if it be true that the proper juices of certain plants move in true
vessels, it does not follow that there is anything analogous for the
ascending sap, or that it is the office of the trachean vessels to trans-
port it.
Direct observation shows that at the earliest period of the forma-
tion of the tissues the trachean vessels are full of juices, like the
other organic elements, and that they are only deprived of these at
a later period. But even then they may be traversed by liquids of
considerable density.
The wood of certain plants, such as Ulmus campestris, Robinia
pseudo-Acacia, and Quercus Ilex contains large vessels, the interior
of which is occupied by a more or less consistent reticular tissue.
This tissue evidently could not have been produced unless the vas-
cular tubes had been filled with a liquid containing organic materials
in solution.
Another observation proves that trachean vessels may contain
substances which become thickened so as to obstruct their cavity.
In a section of the stem of Calamus Rotang the author found, in
380 Miscellaneous.
most of the fibres, the enormous vessel occupying their centre filled
with a solid white substance, forming continuous or interrupted
cylinders of variable length. This substance, when detached and
put into water, breaks up into granules; and, singularly enough, the
suspended grains were sometimes agitated by a very lively move-
ment, although the cane from which they were derived had long
been dry. This substance was contained in well-marked porous
vessels.
In proof that the solid substances are deposited in the vessels
during the life of the plant, the author cites an example to show
that the solid matter, far from penetrating into the vessels after
section, has a tendency at that time to escape from them. An old
vine-stem, 6 centimetres in diameter, had been cut into pieces from
1 to 2 métres in length. In a short time the cut surfaces were
covered with an abundant, transparent, gummy layer. Having
made new and very smooth sections, the author found on the follow-
ing day that filaments of gummy matter 5 or 6 millimétres in length
had issued from the large vessels. Hence it appears certain that,
even at an advanced age, the trachean vessels may contain, not only
gaseous bodies, but also substances which sometimes acquire a great
density.— Comptes Rendus, Oct. 2, 1865, p. 544.
On the Organization of the Cypridine.
By Professor Ciaus.
During a residence at Messina, Prof. Claus turned his attention to
the little Crustacea which swarm in the waters of the sea. He was
particularly struck by a small Ostracode of the genus Cypridina, in
which he detected, even with a low power of the microscope, an
accessory single eye in addition to the large, paired, compound eye,
and a heart beating with regular pulsations. This latter discovery
naturally surprised him, as in the other two families of Ostracoda
(the Cypride and the Cytheride) the heart is entirely deficient.
A more attentive examination of these Crustacea soon showed, how-
ever, that the Cypridine differ much more from the other Ostracoda
than the Cypride@ and Cytheride from each other. :
The fact that an organ so important as the heart may sometimes
exist and sometimes be deficient in animals so nearly allied to each
other is doubtless surprising, but by no means without precedent.
Thus it has been demonstrated that the Copepoda are in the same
case. M. Claus himself has shown that if the Cyclopide, Harpac-
tide, and Corycaide are always destitute of a heart, the allied
Pontellide and Calanide@ are always furnished with one. Moreover
the author is not the only person who has observed the heart in the
Cypridine, as M. Fritz Miller mentions it in a recent work (Fiir
Darwin, 1864).
The sole visual organs hitherto known in the Cypridine were the
paired eyes, in which M. Lilljeborg has detected a complication of
organization very similar to that of the eyes of the Cladocera, although
the latter are fused into a single mass, forming as it were a median
Miscellaneous. 881
eye. Nevertheless traces of a primitive division into two halves in the
Side, the Lyncei, and the Estherie enable us to establish unhesi-
tatingly the homology of this apparently single eye of the Cladocera
with the paired eyes of the Cypridine. A further homology is
presented when we find in the Cypridine, besides the large com-
pound eyes, a small, simple, median eye, perfectly similar to that
which exists, in addition to the compound eye, in the Daphnia.
The Cypridine present other peculiarities worthy of mention.
As a general rule, the Ostracoda are characterized by the small
number of their appendages, as there exist only two, or at most
three, pairs of locomotive appendages behind the gigantic maxillze.
In fact, the last pair of feet disappears completely, and the others
are converted into organs of manducation. On the other hand, the
mandibles are converted into locomotive appendages. The antenne
also serving for locomotion, we find that throughout their whole life
the Cypridine employ the three anterior pairs of appendages as
locomotive organs. Now this is exactly the case in all Entomostraca
during the Naup/ius-phase, and furnishes a new argument to be
added to those adduced by Fritz Miller in favour of the derivation
of all Crustacea from the Nauplius-form.—Siebold § Kolliker’s
Zeitschrift, 1865, p. 143.
Remarks on the Anatomy of Tridacna elongata.
By M. Léon Vaiuuanr.
Tridacna elongata, Lam., occurs very abundantly in the Bay of
Suez, where it is often employed as food ; the author has accordingly
been able to examine a great number of individuals of this animal.
The retractor muscle of the foot, which is of considerable size in
proportion to the protractor, serves in part for the closure of the
valves ; hence it may be that in those Monomyary Acephala which
have an adductor muscle distinctly divided into two parts, the upper
portion is to be regarded as representing the retractor of the foot
diverted from its normal functions. The byssus of the Tridacna,
already described by Miiller, consists of two parts—one adhering to
the bottom of a cavity of the foot, the other uniting this with ex-
ternal bodies ; each of these is secreted by a distinct organ,—the
former by the bottom of its cavity, the latter by a collection of
racemose glands lining a circular groove in the wall of the cavity.
The large notches of the margins of the shells enabled the author
to ascertain the force which the mollusk is capable of exerting. He
fixed an individual by one of its valves, and suspended a weight to
the other. In this way he found that a specimen 21 centimétres in
length, of which the valves weighed 1-264 kil., could support a
weight of 4°914 kil. ; so that it may be supposed that an individual
weighing 250 kilogrammes (and these are not uncommon) might at
a given moment put out a force of more than 900 kilogrammes.
In the nervous system the branchial ganglia, forming a single mass
with no trace of longitudinal division, exhibit transverse furrows
bounding two false cireumvolutions. A sort of inelastic tendon
382 Miscellaneous.
accompanies the connective extended from the branchial ganglion to
one of the buccal ganglia during its passage through the gastro-
genital mass; the object of this arrangement appears to be the pre-
vention of the dragging (¢iraillement) of the nerve when the organ
is distended with eggs.
Another remarkable arrangement is to be seen in the passage of
the last portion of the intestine through the heart. At the entrance
of the intestine into the ventricle there are muscular bundles starting
from the wall of the latter and inserted perpendicularly into the wall
of the digestive tube; at the moment of contraction these bundles
must, by their shortening, tend to draw apart the walls of the intes-
tine, which would otherwise be compressed during the systole, and
thus the course of the feecal matters will not be interrupted.
In these large mollusks the difference between the arteries and
veins is very easily seen: the former have a very distinct double
epithelial and fibrous wall, whilst the latter are simple sinuses hol-
lowed out in the tissues. All the blood is compelled to traverse an
organ of heematosis (branchize or mantle) before returning to the
heart.
The proper temperature of the animal, compared with that of the
bottom at which it lives, appeared to be rather high. The tem-
perature registered by thermometers sunk at the point inhabited by
the animals was about 63°°5 F. (17°°5 C.); the average temperature
of the Tridacne was 68°°5 F. (20° C.).—Comptes Rendus, Oct. 9,
1865, p. 601.
Remarks on the Protective Sheath and on the Formation of the
Stem of the Root. By M. R. Caspary.
In preceding memoirs M. Caspary has indicated a layer of very
closely approximated cells, placed in a single series in thickness,
which exists in stems, roots, and leaves, the vascular system of which
it envelopes and protects. He has given it the name of the protec-
tive sheath, although in certain cases (in Berberis, for example)
this layer is ruptured during growth, and consequently does not
serve to protect the organs which it envelopes. In describing this
protective sheath, M. Caspary indicated upon the lateral walls of its
cells some darker spots or streaks, which he thought were formed
by very small pores. He now again maintains the existence of
these spots or streaks; but he has ascertained that they are due to
foldings of the walls of the cells, and not to pores. He has observed
these folds in the protective sheath of Ficaria ranunculoides, Roth,
Elodea canadensis, Mich., Brasena peltata, Peret, and Charlwoodia
rubra, Planch. When the cells of the protective sheath become
thickened, the folds gradually disappear. M. Caspary attributes
this change to the elongation of the walls of the cells—an elongation
of which he has convinced himself by direct measurements.
Several authors (especially M. Karsten) have regarded this pro-
tective sheath as a lignified residue of the layer of cambium which
has produced the other parts of the stem; but M. Caspary combats
Miscellaneous. 3583
this opinion. He declares himself also against the opinion which
derives all the parts of the stem from a single layer of cambium
existing in the terminal bud. According to him, the entire terminal
bud is formed of cambium, and already contains the mother cells of
all the kinds of tissue which will subsequently form the various parts
of the stem.—Jahrb. fiir Wiss. Bot. 1864; Bibl. Univ. 1865, Bull.
Sci. p. 87.
Graduation from “ Individual Peculiarities”’ to Species in Insects.
The following are the concluding paragraphs of a paper by Dr. B.
D. Walsh “ On Phytophagic Varieties and Phytophagic Species.”
The name phytophagic is given to those otherwise identical insects
which differ, as varieties or species, according to the species of plant
they feed upon. ‘‘ When certain unimportant characters in the
insect are correlated with the food-plant, while at the same time there
is no sufficient reason to doubt that the two varieties freely inter-
cross,” the forms are called phytophagic varieties. When, from the
lack of intermediate forms, intercrossing may be inferred not to take
place, they are called phytophagic species. Dr. Walsh sums up his
conclusions thus :—
“From the facts referred to above, and those recorded by me else-
where, we may construct the following almost unbroken series, from
the first dawnings of the Phytophagic Variety to the full develop-
ment of the Phytophagic Species :—
“1st. Difference of food, even when the food-plant belongs to
widely distinct botanical families, is accompanied by no difference
whatever either in the larva, pupa, or imago state: dAttacus Cecropia,
Linn. ; Dryocampa imperialis, Drury; Lachnus Carye, Harris
(Proc. Ent. Soc. Phil. vol. i. p. 303) ; and hundreds of other species.
“2nd. Difference of food is accompanied by a marked difference
in the colour of the silk-producing secretions : Bombyx Mori, Linn.,
the common silkworm.
‘3rd. Difference of food is accompanied by a tendency toward the
obliteration of the normal dark markings in the imago: Haltica
alternata, Ilhg.
‘Ath. Difference of food is accompanied by marked, but not per-
fectly constant, colorational differences in the larva, but none what-
ever in the ¢ 2 imago: Datana Ministra, Drury.
“5th. Difference of food is accompanied by a marked and perfectly
constant difference in the size of the imago: Chrysomela scalaris, Lec.
“6th. Difference of food is accompanied by a marked difference
in the chemical properties of gall-producing secretions, the external
characters of the ¢ 2 imago remaining identical: Cynips q. spon-
gifica, O. S., and C. q. inanis, O. S.
‘7th. Difference of food is accompanied by a slight but constant
change in the coloration of the abdomen of the ¢ 2 imago, and by
a very slight change in the chemical properties of the gall-producing
secretions, the galls of the two insects, though typically somewhat
distinct, being connected by intermediate grades in the case of the
latter: Cynips q. punctata, Basset, and C. q. Podagre, Walsh.
384 Miscellaneous.
“8th. Difference of food is accompanied by one marked and per-
fectly constant colorational difference, and others which are not per-
fectly constant, in the larva, but none whatever in the ¢ 2 imago:
Halesidota tessellaris, Sm. Abb., and H. Antiphola, Walsh.
“9th. Difference of food is accompanied by several slight but con-
stant structural differences in the ¢ imago, but none whatever in the
2 imago: Olytus Robinia, Forst., and Cl. pictus, Drury. AG
“10th. Difference of food is accompanied by a slight but constant
structural difference in both d and ?imago: 1. Tingis Tilia, 1. sp.,
and T'. amorphe, n. sp.; 2 (doubtful). Diapheromera femorata,
Say, and D. Velii, n. sp.
* 11th (doubtful). Difference of food is accompanied by very strong
structural and colorational differences in the larva and in all proba-
bility by a constant structural difference of generic value in the @
imago, the ¢ imagos being to all external appearances identical, and
the two insects belonging to different genera: Sphingicampa di-
stigma $ 2, Walsh, and Dryocampa bicolor 3, Harris.
“12th. Difference of food is accompanied by marked and constant
differences, either colorational or structural, or both, in the larva,
pupa, and imago states: Halesidota tesellaris, Sm. Abb., and H,
Carye, Harris, and hundreds of species belonging to the same genus,
and commonly considered as distinct species.
“*The constitution of the human mind is such, that the same evi-
dence carries with it very different degrees of weight when presented
to different intellects. Others will no doubt draw different conclusions
from the facts catalogued above; but for my own part, as on the
most careful consideration I am unable to draw any definite line in
the above series, and to say with certainty that here end the Varieties
and here begin the Species, I am therefore irresistibly led to believe
that the former gradually strengthen and become developed into the
latter, and that the difference between them is merely one of mode
and degree.” —Silliman’s American Journal, September 1865.
Note. on the Cultivation of Eels. By M. L. Souserran.
The author states that for several years past considerable quan-
tities of young eels have been taken at the mouths of the French
rivers and distributed in the inland waters ; but he adds that, from
his own experience, this course is not always judicious, and is fre-
quently unprofitable. He mentions that in 1856 certain landed
proprietors in the neighbourhood of Caen transported great quan-
tities of young eels to the ponds and other waters on their estates,
and after feeding them at great expense obtained nothing but loss from
their undertaking, the produce being only 150 francs against an ex-
penditure of 2220 francs. Besides this, the waters into which the
eels were introduced, and those into which they subsequently pe-
netrated, were entirely depopulated of other species of fish ; so that
the multiplication of eels must be regarded as in every respect
a losing speculation.—Comptes Rendus, 4th September, 1865,
p. 424.
PE ANN ALS
MAGAZINE OF NATURAL HISTORY.
[THIRD SERIES. ]
No. 96. DECEMBER 1865.
XLII.—On the Systematic Value of the Organs which have been
employed as Fundamental Characters in the Classification of
Mollusca. By Dr. O. A. L. Morcu.
WHETHER Classes, orders, and genera are founded in nature, or
are only artificial divisions, is a question rarely disputed. It is,
however, still a matter of discussion whether now-existing spe-
cies are direct descendants of extinct forms from remote geo-
logical periods, which have been gradually changed in the course
of natural selection or in consequence of physical changes of the
globe, or are entirely new creations of any one geological era.
The habits of an animal often cause a considerable modification
of the external form, size, or colour, which are often improperly
considered of specific value; but it must be admitted that such
differences are subject to limits which cannot be passed, and do
not become hereditary. For instance, the corns of the human foot
are, like the nails, a thickening of the epidermis ; but the former
are produced by accidental pressure on the foot, while the latter
originate in the foetal structure of the animal. There is the
same distinction between false and genuine species.
Linné divided all creation into three kingdoms—animal,
vegetable, and mineral. The last-named division is less logical,
because minerals can only be considered as parts of the great
celestial bodies, which may be regarded as organic beings with
involuntary motion impressed upon them, corresponding with
that of the heart or stomach of animals. Geoffroy St.-Hilaire
has more correctly made the division into phanerobiotie and
cryptobiotic kingdoms. <A predestined scheme is shown in the
chronology of our planet as well as in the development of organic
beings. Thus the oceans with their lower types are regularly
in course of time changed to islands and continents with their
Ann. & Mag. N. Hist. Ser.3. Vol. xvi. 26
3886 Dr. O. A. L. Morch on the Systematic Value of the
higher terrestrial types. Even the form of the now-existing
continents seems to be subject to a certain definite plan. The
three main continents (1. Europe and Africa, 2. North and South
America, 3. Asia and Australia) extending from north to south,
nearly forming a continuous range, show a remarkable resem-
blance in their configuration—towards the south a large tri-
angular mass of land without considerable gulfs (Africa, South
America, Australia) united by an isthmus (Suez, Panama, Ma-
lacca and Sunda islands) to a larger land with numerous gulfs
and peninsulas, forming a large median gulf (the Mediterranean
Sea, the Gulf of Mexico, Mindoro Sea) with numerous archi-
pelagos at the entrance (Canarian and Cape de Verde Islands,
West-India Islands, Philippine and Molucca Pacific islands) ;
close to each of the southern parts is a satellite island with a
most remarkable aberrant fauna— Madagascar, New Zealand, and
Galapagos Islands. These three continents would geologically
be considered three different formations. Australia, probably the
youngest continent, representing the Liassic period of the Old
World, is characterized by the lowest Mammalia, viz. the abor-
tive Marsupiala. America has only a few Marsupialia, fre-
quently without pouch (marsupium), but the greatest number
of the lowest Placentalia (Microsthena), the Edentata. Africa
has no Marsupialia, but a few Edentata, and abundance of the
higher Mammalia (Macrosthena). If an order is common to
the Old and New Worlds, there is always a marked difference
according to the continent, the species of the Old World belong-
ing to groups of a superior stamp. Thus the Quadrumana and
Scansores (Parrots, among birds, corresponding with Monkeys)
are divided into those of the Old and those of the New World
—the latter being chiefly long-tailed, a character considered
inferior,
The Ampullarie of the New World are nearly all provided
with a siphon, which is wanting in those of the Old World.
The same is the case with the genus Polymesodon in the Cyrenide,
and Leila among the Unionide. The difference between the
faunze of the three continents diminishes towards the north,
where the faunz are fused into one circumpolar fauna, like the
three continents themselves. The land-faune are limited by
thermal differences or mountain-ranges. Species living in or
or close to water (e. g. Succinea) are very similar all over the
world,
The three continents above mentioned divide the ocean into
three parts geographically, but not zoologically—the Atlantic,
the Pacific, and the Indian oceans. These are limited by the
following coast-lines :—
1. From Behring’s Straits, Norway, Spain, Guinea, Cape.
Organs employed in the Classification of the Mollusca. 387
2. From Behring’s Straits, Greenland, Florida, Brazil, Cape
Horn.
3. a Oregon, California, Peru, Cape Horn,
4. 3 Kamtschatka, Japan, China, Tas-
mania.
5 Cape, Arabia, India, Sumatra, Tasmania.
The three first-named coast-lmes may zoologically be con-
sidered the boundaries of the Atlantic Ocean, in the same way
that America is considered an island in the Atlantic Ocean,
separated from the Pacific Ocean by the western edges of the
polar streams and the great abyss between the Galapagos Islands
and the Sandwich Islands. The coast-lines are divided into
analogous faunz according to the climate: 1. Polar; 2. Sub-
polar (North Sea); 3. Subtropical (Mediterranean) ; 4. Tropical
(Guinea). The southern polar and subpolar are not sufliciently
known (Kerguelen Island, St. Paul’s, &c.).
It would seem easy for most animals, even for Mollusca, the
young of which generally are natatory, to move or to be carried
by currents from one end of these coast-lines to the other; but
in reality this’is not the case. The temperature of the sea
seems to put a nearly insurmountable barrier to the distribution
or migration of species, and even of genera. A general view
of those faunze shows that they are composed of animals of all
subkingdoms, classes, orders, families of the animal kingdom
(Gulls, Seals, &c). The genera are often different, although
allied ; and the species are nearly always different in each fauna,
The question now arises, Are these species and genera originally
different, or changed by various physical causes, such as climate,
light, the saltness of the sea, &c.? The influence of these causes
on the form and size of shells is chiefly seen by comparing
the faunz of the Baltic and Mediterranean with that of the
ocean.
It would be a very slow process to compare the species of all
these faunze, chiefly because no museum possesses a sufficient
collection of animals in spirits for investigation of the teeth,
even of European species. ‘The specific importance of the teeth
is evident, chiefly from the researches of Troschel ; many species
which were considered slight varieties are now proved to be
distinct—e. g., Natica millepunctata and N. maculosa, Natica
clausa and N. consolidata, &c., &c., Viviparus contectus and V,
fasciatus, Bithinia ventricosa and B. tentaculata.
The genus Aporrhais, which was abundant in former geo-
logical periods, from the Oolitic formation upwards, is now
restricted to five or six recent species inhabiting the Atlantic
only. Aporrhais occidentalis is found on the coast of Canada
26*
388 Dr. O. A. L. Morch on the Systematic Value of the
and Greenland. A. pes-pelecani and a variety, perhaps specifi-
cally distinct, are found from the north of Norway to the Medi-
terranean. In Shetland a second species is added, the A. pes-
earbonis, which in the Mediterranean is represented by A. Ser-
resiana, Phil. In Senegal is found A. senegalensis, Gray. The
most northern species is the largest of the genus, as is generally
the case; the outside of the outer lip is smooth, as in all other
arctic univalves. In A. pes-pelecani and the following southern
species the inside lip is plaited, as is generally the case in species
of warmer climes (e. g. Nassa and other univalves). The A.
senegalensis, living in the hottest water, is the smallest. Un-
fortunately it will be very difficult to get the tongues of these
species. The teeth of Aporrhais pes-pelecani as represented by
Lovén, Troschel, and Wilton differ considerably from one an-
other, making it probable that two species are confounded.
A monographic research, chiefly based on the teeth of the
genera Nassa, Fusus, and Buccinum, found on the coast-lines
from the arctic regions to the equator, would probably be suffi-
cient to prove whether species in each fauna are created originally
or are only varieties dependent on different climates, and would
at the same time prove the relations between the species of suc-
ceeding geological periods.
Large suites of specimens, from various depths and localities,
are in most cases sufficient to prove the difference or identity of
so-called species, without a knowledge of the animals; but the
affinity of the genera and families can only be safely understood
by anatomical researches: the anatomical as well as the zoolo-
gical characters are, however, generally either misunderstood or
overvalued. I will therefore endeavour to show the relative
systematic value of the various organs of the Mollusca. Until
these questions are quite cleared up, it will not be possible to
solve the problem in regard to the origin of species among the
Mollusca.
Cuvier founded his primary divisions (classes) of the Mollusca
on the locomotive organs, viz. Cephalopoda, Pteropoda, Gastero-
poda, Acephala (this last changed by Oken to Pelecypoda),
Brachiopoda, and Cirrhopoda. The group Heteropoda, correctly
considered by Cuvier only a family, was by most subsequent
naturalists considered of the same value as the above-mentioned
divisions. ‘The study of the homology of the parts of the Mol-
lusea, commenced by Professors Lovén and Huxley, has shown
that the Pteropoda are true Gasteropoda, and that the funnel of
the Cephalopoda is homologous with the foot of the Gymnosomata.
Such a system seems to me to correspond with that classification,
by the old authors, of the Vertebrata (Quadrupeda, Bipeda,
Pinnata, and Apoda), maintained so pertinaciously by Klein in
Organs employed in the Classification of the Mollusca. 389
opposition to the Linnzean system founded chiefly on the ana-
tomical researches of Ray. The Cetacea, but chiefly the Reptiles
(Anguis, Bipes, &c.), afford striking examples of the insignifi-
cance of the locomotive organs as a base for the primary divi-
sions.
The secondary divisions (orders) of Cuvier were founded on
the respiratory organs. Respiration is indispensable for the life
of all organic beings ; but special organs for this purpose are not
always necessary. Respiration of some importance takes place
through the skin, even in the higher Vertebrata, and can, in the
lower Vertebrata (e. g. Batrachians), temporarily replace that
through the lungs: it is quite sufficient for many Mollusca.
The Vertebrata have two kinds of respiratory organs—lungs,
and gills—which, according to J. Miller, are not homologous,
as they can be found together in the same individual, although
not always exactly performing the same function—for instance,
the Batrachians and the foetus of the higher Vertebrata. In
fishes the lungs are reduced to a swimming-bladder, the walls
of which are provided with some bloodvessels, making it serve
as a secondary, but very imperfect, respiratory organ comparable
to the lung-sac of the Pulmonata. That this organ, even among
the Mollusca, cannot be homologous with the gills, as advanced
by Prof. Agassiz *, is proved in Ampullaria and perhaps Onchis,
each of which has lung-sacs simultaneously with true gills.
Many Mollusca (as Cyclostoma, Neritina, and Littorina), which
are undoubtedly provided with gills, live always or nearly always
in the air, probably having the power of keeping their gills moist,
like the Land-crabs and several fishes (e. g. Anabas scandens).
Whether the vena branchialis and vena pulmonalis are identical
is at least not yet proved. In Mollusca not requiring a hard
covering for their protection, respiration takes place through the
skin; but when the skin is thickened, or a shell developed, a
respiratory organ becomes necessary. ‘The use of the gills is to
produce, by the complication of a part of the skin, a surface
corresponding with the area made impenetrable to the oxygen
of the surrounding medium (air or water). The effect of this
dermal gland is increased by vibratile cilia, producing a swifter
circulation of the oxygeniferous medium. If the branchial sac
is very deep, the circulation of the water is effected by a siphon
acting like a chimney-pipe, often assisted by muscular contrac-
tions (Cephalopoda, Acephala). The larger the shell is in pro-
portion to the uncovered parts of the animal, the more compli-
cated and compressed are the gills.
Several of the internal glands are subject to the same change.
* <QOn the Cireulation of Fluids in Insects,” Proc. Bost. Soc. 1850,
p- 237.
390 = Dr. O. A. L. Mérch on the Systematic Value of the
Thus the liver is strongly arborescent in animals having a dermal
respiration (Pellibranchiata, Gymnobranchiata), and becomes
more and more compressed in proportion as the gills are deve-
loped (Scyllaa, Pleurophyllida) ; but it is most complicated in
Mollusca covered with a shell, The same is the case with the
renal organ, which becomes rounded and spongy, as is to be
seen in the plates of Mr. Hancock’s excellent paper* on the renal
organ of nudibranchiate Mollusca, and in the work of Gegenbaur +
on the Heteropoda and Pteropoda. The compact structure of
this organ in the testaceous Mollusca may be seen in many of
the plates to the ‘ Voyage de |’Astrolabe.’
In the Pellibranchiata the generative glands partake of the
arborescent form of the liver and kidney: thus in Elysia f, &c.,
the albuminous gland, the hermaphrodite gland, and “la glande
en tréfle” of Moquin§ are strongly arborescent.
The dorsal plates of Placobranchus and the arborescent tufts
of Dendronotus and Onchis may be considered the most imper-
fect forms of gills. The hepatic papille of the Molidide are
probably not homologous with the branchial leaves of Gastero-
poda (e.g. Lanthina, Pterotrachea, Doris), as these organs in
Bornella are found simultaneously with arborescent tufts of the
skin, which may be considered homologous with the branchiz
of the Doridide. There appears to be a gradual transition in
the respiratory organs of Tritonia, Heptabranchus, Hexabranchus,
and Lamellidoris. It must, however, be remembered that, ac-
cording to Dr. Hancock, the gills of Doris receive hepatic blood.
The gills in symmetrical animals are generally situated on each
side of the body, e.g. Acephala, Cyclobranchia, Inferobranchia.
In asymmetric spiral testaceous Mollusca the single gill becomes
smaller, and is said to be reduced to a filiform undulated vessel,
as in Vermetus and Onustus ||. I have observed, however, in a
specimen of O. trochiformis, that this vessel opens through a
pore on the outer side of the mantle opposite to the shell ; it
may perhaps be in some relation with the renal organ.
The insignificance of the gills as a systematic character is
evident by comparing the Heteropoda, from the entirely gill-
less Firoloides and Pterotrachea with external gills, to <Af-
lanta exhibiting perfectly internal gills. The same fact is to be
seen in the following series of allied genera :—Stylochilus, No-
tarchus, Aplysia, Bulla, &c., Acteon, Odostomia, and Obeliscus.
* Trans. Linn. Soe. 1864.
+ Untersuchungen iiber die Pteropoden u. Heteropoden, 1854.
{ Journal de Conchyhologie, vol. 1.
§ “Le talon” (St. Simon) = “ petite glande séeréteur” (Souleyet, /. c.
to Guteo))-
|| Moreh, Journal de Conchyliogie, vi. p. 308.
Organs employed in the Classification of the Mollusca. 391
These two series prove not only that the size and form of the
gills stand in connexion with the development of the shell, but
that the divisions Opisthobranchiata and Prosobranchiata, as
originally proposed, are not natural ; this has been indicated by
Messrs. Huxley and Macdonald.
The two kinds of respiratory organs indicate only relative
superiority and inferiority, but not limits of systematic divisions.
Thus, among Vertebrata, lungs and gills meet in a family of the
Batrachians. Among the Mollusca, it is probably in Auricula
and Obeliscus that the two kinds of respiratory organs are found
to meet.
It has been generally understood that neither the locomotive
nor the respiratory organs offer characters for the primary divi-
sions; most authors have therefore divided the Mollusca, ac-
cording to the presence or absence of a distinct head, into
Cephalophora and Acephala. As several Gasteropoda (e.g. The-
cosomata) properly have not a distinct head, Prof. Lovén has
changed the names to Glossophora and Aglossa. Cephaliza-
tion, as explained by Prof. Dana, is, no doubt, of importance as
a character indicating relative superiority and inferiority, but is
not sufficient for natural divisions. Thus, according to this
principle, the Vertebrata would only be divided into two classes,
1. Cephalophora with a distinct head—Mammialia, Birds, and
Reptiles ; 2. Acephala, with the head united to the thorax, con-
taining the Fishes only.
My. study of the Mollusca for about twenty years enables me
to state that the heart and generative organs offer characters of
a much higher systematic value than is generally believed. It
is perhaps “somewhat hazardous to compare the organs of the
lower animals with those of the higher; but it does not seem
probable that organs which have no systematic value in the
higher can have it im the lower animals.
The accompanying synopsis of the Mollusca is chiefly founded
on the intromittent male organ, which seems to me to be the
best indicator of the sensibility of the nervous system, and con-
sequently of the relative systematic rank of the animal. Thus
the lowest class of Vertebrata (the Fishes) wants an intromittent
male organ, although the sexes, with few exceptions, are sepa-
rate; there exists, consequently, no copulation, but impregna-
tion takes place as among plants. In the Plagiostoms the pos-
terior locomotive organs “of the male are e changed into conduits
for the sperm*, Hilees the hectocotylized arm of the Cephalopods.
In the Batrachians the anterior locomotive organs are used as
prehensile organs during the pseudo-copulation; but a true
* Steenstrup, Om agg preter p. 26, Kgl. Danske Videnska-
bernes Selskabs Skrifter, 5. Raekke, 4. Bind.
392 =Dr.O. A. L. Morch on the Systematic Value of the
male intromittent organ is first found among the Reptiles, and
becomes more and more developed among the higher Vertebrata.
That this fact is not in consequence of aquatic habits is evi-
dent by comparing the Cetacea and Sea-serpents, which are
provided with a true male organ.
A similar fact may be observed among the Articulata, of
which the Insects only may have a true male organ.
Among the Mollusca the Androgyna are provided with the
most developed male organ, and which seems to be the most
sensitive. The male organ of the Exophallia is always external,
not retractile, sometimes concealed in a furrow of the right ten-
tacle (e. g. Viviparus). The Pseudophallia have no male organ*.
The copulation is probably effected in the same way as among
Batrachians ; but direct observations are wanting. The male
Acephala disperse their sperm, which is taken up by the females
like the pollen by flowers. The few observations on the copula-
tion of Acephala are very problematical. My synopsis confirms
the rule of Prof. Agassiz that land-animals.are more perfect than
marie; but this rule may be explained in the sense that the
divisions with the largest number of terrestrial forms always are
the superior. Thus, among the Vertebrata, the Thermaliat+ are
higher than the Psychraemia, containing the greatest number of
aquatic forms. The lowest class, Acephala, is entirely aquatic
and chiefly marine. Among the Pseudophallia, Helicine are the
only terrestrial, and Neritine fluviatile. The number of ter-
restrial genera is considerably increased among the Exophallia ;
thus Cyclostomacee are truly terrestrial, Ampullarie, Paludine,
Melanie, and Potamide fluviatile. Among Androgyna are the
greatest number of terrestrial genera.
There is the same concordance with the law of Prof. Owen,
a that the multiplicity of organs indicates inferiority in organiza-
tion.” As was shown in a former paper, the duplicity of the
organs of Acephala descends as the system ascends.
The development of the young is of less systematic value than
is generally believed: this is proved in the Crustaceans, the
marine species having a larval form very different from the
fluviatile species (Astacus fluviatilis and A. marinus). All larvee
of marine Mollusca swim by means of a velum, which in the
marine Acephala, Chiton and Dentalium is changed to a flagel-
lum. In the melicertigene Gasteropoda (Rhachiglossata) the
larva loses the velum before it leaves the egg-case. The larva
* In the ‘ Voyage de l’Uranie’ is represented a Neritina with something
like a short male organ between the mouth and right tentacle, but which
seems not to have been observed in the European species.
+ The difference between hot- and cool-blooded animals seems not to
be sufliciently insisted upon in the modern systems,
Organs employed in the Classification of the Mollusca. 393
of Auricula agrees with that of marine Mollusca, according to
Dr. C. Semper, although the most allied forms (Limnea) are not
subject to this kind of metamorphosis.
All the terrestrial forms leave the egg in a perfect state. The
Cephalopoda may be considered Gasteropoda stopped in the
larval stage, reminding one of Macgillivrayia. The number of
the eggs may also be of some importance for the determination
of superiority and inferiority, as nature seems to compensate
want of intelligence, or power to provide for the offsprmg, by
great fertility: thus plants are more fertile than animals; the
Helicea produce but few eggs compared with the Acephala.
Since the time of Cuvier nearly all naturalists have considered
the Cephalopoda the highest type of Mollusca, chiefly on account
of a presumed affinity with the Fishes, their great size, great
muscular power, as well as the apparent superiority of the
nervous system and organs of circulation. Naturalists are not
yet agreed which families may be considered the highest in
each class, except in the Mammalia. Among the Birds, the
Parrots are considered the highest by the best authors; the
Serpents may perhaps be the highest Reptiles, although such
an authority as Prof. Agassiz considers the Chelonians the
highest, on account of the completeness of the ossification.
According to this principle, the Edentata would be the highest
Mammalia! The largest and strongest Arthropods, the Lob-
sters, have a similar claim to be considered the highest of that
subkingdom. Marine animals are always larger than their kin-
dred on the land, but not the most perfect, as Prof. Agassiz
has proved.
The systematic place of the Cephalopoda may depend on the
structure of the heart and the explanation of the hectocotylized
arm as a male organ. If the branchiocardiae veins of Cephalo-
poda may be considered auricles, as stated by Milne-Edwards,
Kolhker, Huxley *, and Gegenbaur +, the place of the Cephalo-
poda must be between Dentalium and Acephalat—a place not
more strange than that of the Cecilie standing before the Pla-
giostomes, or the Linguatula before the Decapod Crustacea.
The hectocotylized arm of the male indicates a kind of copu-
lation between two individuals, giving the Cephalopoda claim to
a higher place than the Acephala; but it must be remembered
that the manner of copulation of the Pseudophallia is entirely
unknown at present.
If the cardiac auricles only prove to be tumefactions of the
* On Morphology, &e. p. 57—‘ the presumed highest Mollusca.”
+ Vergleichende Anatomie, p. Soe
{ Solen swims, according to Prof. Deshayes, like the Cephalopoda, by
driving water out of the respiratory cavity.
394 Dr.O.A. L. Morch on the Systematic Value of the
branchiocardiac veins, the Cephalopoda must then be removed,
according to the septiserial teeth of the tongue, to the Tzenio-
glossata, the larvee of which (Macgillivrayide) are not unlike the
Cephalopoda. This position would answer to that of Cetacea
being united by the Sirenians to Pachydermata.
The pelagic Cephalopoda are certainly the most powerful and
ferocious of all Mollusca, like the Sharks and Cachalots among
Vertebrata; but in the same divisions are found large animals
living on small animals (e. g. the Whalebone-Whale and the
Squalus glacialis). The same is probably the case with the
largest of all Cephalopoda, Architeuthis dux, Stp. The two man-
dibles of the Cephalopoda are generally very powerful and very
conspicuous ; but their little value as a character for a class is
best seen by the two divisions of mandibulated Echinodermata
and the Helicea. The eyes offer the same differences as in other
parts of the animal kingdom, being small in the diurnal species
and larger in the nocturnal. The size of the lens cannot be
considered a character of perfection of the visual organs, because
the fishes would in such case have the most perfect eyes among
Vertebrata. Nautilus wants a lens, according to Prof. Keferstein.
The exterior form of Cephalopoda is a combination of the form
of the Fishes and Radiata*, depending on their manner of loco-
motion. Cuirroteuthis has thus a striking resemblance to a Me-
dusa. The Cephalopoda seem to me not to have more just
claims to be considered a distinct class in relation to the other
Mollusca than Pteropoda or the Cetacea among Vertebrata.
They exhibit a remarkable analogy with the Plaviostomes in
their habits, in the male organ, the manner of depositing their
eggs, and in the yolk-bearing young.
Subkingdom III. MOLLUSCA.
Series I. MONOTOCARDIA.
The heart with a single auricle. Copulation between two individuals.
Class I. ANDROGYNA (Musivoglossata olim).
All imdividuals alike in respect of the sexual organs, having the two
sexes united. Always provided with a receptaculum seminis (petio-
late bladder). Male organ retractile. Teeth of the tongue generally
multiserial. Predominantly mandibulated.
PULMONATA.
GropuHILa : Phyllovora, Agnathaf.
HyGropuita: Planorbis, Physa, Limnea, Siphonaria, Ancylus,
Auricula.
* Owen, Proceedings of the Zool. Society, 1836, p. 19.
+ Journal de Conchyliologie, July 1865.
Organs employed in the Classification of the Mollusca. 395
TECTIBRANCHIA : Pyramidella, Obeliscus, Odostomia, Chemnitzia,
Acteon, Bulla, Aplysia, Notarchus.
Preropopa: Gymnosomata,—Clione, Pneumodermon (Ianthina?) ;
Thecosomata,— Clio, Hyalea, &e.
GYMNOBRANCHIA.
1. Pygobranchia: Doride, &c.
2. Pleurognatha: Plewrophyllidia, Dendronotus, Tritonia, Bornella,
Molis, Glaucus, &e.
3. Pellibranchia: Tethys, Chionera, Hermea, Elysia, Limapontia,
Pelta, &c.
Class HI. EXOPHALLIA (Arthroglossata olim).
Sexes distinct ; male organ not retractile (often concealed in the bran-
chial cavity or in the tentacle). Mouth predominantly suctorial.
Lingual strap not having more than seven rows of teeth.
TA#NIOGLOsSATA. ‘Tongue with seven rows of teeth, with reflected
edge. Larva of the marine species swimming.
A. Rostrifera. With a short muzzle, not retractile.
#. terrestria. Cyclostomacea (Truncatella).
8. fluviatilia. Ampullaria,-Paludina, Melania, Potamides, Ceri-
thium, Turritella (with nine? rows of teeth), Littorina, Lacuna,
Velutina, Onchidiopsis.
y- parasitica. The eggs hatched in pouches attached to the in-
side of the shell. Vermetus, Crepidula, Hipponyx, Capulus.
6. Pelagica Heteropoda. Firoloides, Pterotrachea, Cardiapoda,
Peltaria*, Carinaria, Helicophlegma, Atlanta, Bellerophon,
Onustus.
e. Strombi.
B. Proboscidifera. Rostrum retractile. Natica, Ovula, Pedicularia,
Trivia, Cyprea, Cassis, Dolium, Pyrula, Triton, Trichotropis,
Aporrhais.
RHACHIGLOSSATA. Tongue not having more than three rows of
teeth, which are not reflected at the edge. Rostrum long, retrac-
tile (without cheek-plates?). Eggs deposited in cartilaginous cap-
sules, which the young leave when the metamorphosis is complete.
Marginella, Voluta (lateral teeth absent), Harpa, Oliva, Ancillaria,
Bullia, Nassa, Buccinum, Fusus, Fasciolaria, Turbinella, Murex,
Purpura (Magilus?), Mitra.
ToxoGLossaTa. Mouth with a suctorial veil. Tongue (?) with
subulate teeth provided with an internal or external veniferous
canal. Conus (Borsonia?), Pleurotoma, Clionella, making a transi-
tion to Terebra, Cancellaria. The egg-cases of this division are
probably like the preceding ; but direct observation is wanting.
Series I]. DIOTOCARDIA.
Heart with two auri¢les. (That this character has no relation to the posi-
tion of the gills, as Prof. Huxley suggested, is evident in the case of
Pleurophyllidia, which, according to Messrs. Hancock and Bergh, have
* Carinaria depressa, Rang, Man. t. 3. f. 1.
396 Dr. O.A.L. Mérch on the Classification of the Mollusca.
a single auricle, although the gills are lateral—and because Turbo mar-
moreus, with a single “gill, has two auricles, according to Quoy and
Gaimard.) Sexes distinct, but without male organ.
Class III. PSEUDOPHALLIA (Aspidobranchia olim).
Tongue with the lateral teeth of two different forms. Male organ
rudimentary ? ? Development only known in Chiton and Dentalium.
RurprpoGLossaTa. Median teeth broad, with reflected edge, ge-
nerally 5.1.5; marginal teeth compressed ; these are very numerous,
with inflected tips.
Terrestria. Helicina. (Eyes sessile.)
Fluviatilia. Neritina. (Kyes petiolate.)
Marina. Nerita, Turbo, Trochus, Haliotis, Fissurella, Emar-
ginula. (Kyes sessile.)
HereroGLossata. (Dochoglossata, Troschel). Tips of the teeth
(always?) having a black pigment. The marginal teeth of the
preceding division are absent.
Cyclobranehia. Patella, Tectura.
Polyplacophora. Chiton, Chitonellus.
Cirrobranchia. Dentalium, Siphonodentalium.
CrPHALOPODA.
1. Dibranchiata.
A. Octopoda.
B. Decapoda.
Oigopside,
Myopsidee.
2. Tetrabranchiata. Nautilus.
Class IV. ACEPHALA (Dithyra).
Dimyaria.
Heteromyaria. Mytilacea et Ostreacea?
Monomyaria, Pecten, Spondylus, Lima, Tridacna.
Observations.
Tanthina has perhaps the same relation to Pneumodermon as
Carinaria to Pterotrachea. Tanthina has a pair of epipodial fins.
Dr. Gould has represented (American Explormg Expedition)
some aciculate bodies, probably homologous with the hooks on
the arms of Pneumodermon. The latter genus has, according to
D’Orbigny, a small hump on the back, probably a rudimentary
mantle ; but intermediate genera are yet unknown.
The Thecosomata differ considerably from the Gymnosomata.
I find it at least very doubtful whether Eurybia makes a true
transition between the two divisions. The*Thecosomata have
3-5 band-like lateral mandibles, and a triseriate lingual dentition,
quite different from that of the Gymnosomata. According to
Souleyet, they have a petiolate bladder, and are consequently
Mr. A. G. Butler on new Species of Butterflies. 397
androgynous ; but they are perhaps more nearly allied to Gas-
teropteron or the Bullide. The systematic position seems to me
not sufficiently clear.
The relation between the pulmoniferous Awricu/a with a marine
larva and the marine branchiferous Pyramidella requires further
observation. QQuoy and Gaimard have pointed out their affinity.
The relation between a rostrum (not retractile) and a proboscis
or haustellum (retractile) is not yet sufficiently understood.
Perhaps there is no other difference than between a long and
short siphon. Strombus appears to make a transition between
the two, and seems to approach most to Cyprea.
XLITI.—Description of four new Species of Butterflies in the
Collection of the British Museum. By A. G. Burier, F.Z.S.,
Assistant, Zoological Department, British Museum.
1. Anthocharis Leo.
Upperside—front wings snowy-white, with a pale orange patch
on the inner margin, bounded above by the third median ner-
vule; base greyish ; apex and a subapical line grey; nervures
black : hind wings snowy white, suffused with orange on the
front margin.
Body grey ; head and prothorax yellowish.
Underside white, tinted with ochreous, darker at the apex of
front wings and front margin of hind wings; a curved line of
small indistinct brown spots crosses the hind wings just below
the middle.
Expanse of wings 1 inch 9 lines.
Hab. White Nile.
2. Danais Mariana.
Upperside—front wings, basal half, an irregular subapical band,
a small spot on the costa just beyond the middle, and two mar-
ginal spots between the median nervules sap-green; apical half
and nervures rich brown, its inner outline angularly and irregu-
larly notched : hind wings, basal half sap-green, tapering to-
wards the termination of the upper disco-cellular nervule, with
an angular and irregular outline ; apical half and inner marginal
nervures rich brown; other nervures reddish.
Body brown; head reddish, spotted with white; antennz
black.
Underside—front wings as above, but with three additional
small marginal green spots and paler-coloured nervures ; hind
wings as above, but with two submarginal rows of pale-green
spots placed in fours between the nervures along the hind mar-
gin, and the nervures margined with brown.
398 Mr. A. G. Butler on new Species of Butterflies.
Body: thorax brown, spotted with green; palpi green; ab-
domen olivaceous brown.
Expanse of wings 1 ch 11 lines.
Hab. New Caledonia.
3. Diadema Mena.
Hestina Mena, Moore, MS.
Upperside—front wings pale greenish ; nervures, end of cell,
and a submarginal band along the hind margin broadly rich
brown ; two indistinct inner submarginal bands of brown scales:
hind wings pale ochreous, nervures rich brown; a submarginal
row of brown lunules between the nervures along the hind
margin.
Body brown above ; head and thorax streaked with ochreous;
antennee black.
Underside—front wings pale greenish; end of cell, base of
nervures, an indistinct band of scales between the nervures just
beyond the cell, and an indistinct submarginal row of spots
along the hind margin brown; termination of nervures fulvous:
hind wings pale ochreous ; nervures brown; a submarginal band
of indistinct brown spots between the nervures along the hind
margin ; front margin darker ochreous.
Body rich brown; tibiz and tarsi of front legs and tarsi of
middle and hind legs alternately brown and white.
Expanse of wings 34 inches.
Hab. North India.
This insect was named by Mr. Frederick Moore, of the Indian
Museum.
4. Heterochroa Lydia.
Upperside rich brown, with a central white band, faintly
tinted with violaceous, extending, entire, from the first discoidal
nervure of the front to the anal angle of the hind wings, con-
tinued above the first discoidal to the subcostal nervure in the
form of two minute and indistinct white spots; a large angular
orange blotch near the apex of the front wings, extending from
the costa to the first discoidal nervure ; a small orange blotch
at the anal angle of hind wings; two submarginal dark-brown
bands, interrupted by the nervures, near the hind margin of
front wings, and three similar bands on the hind wings; base
irrorated with orange scales; cell of front wings crossed by five
equidistant, oblique, black lines, the one nearest the base more
oblique than the others, extending half across the cell; three
similar short black lines below the cell; hind-wing cell only
crossed by two black lines, and closed by a third ; inner margin
pale brown.
Mr. H. J. Carter on Peridinium eypripedium. 399
Body olivaceous ; head fulvous; antenne black, tipped with
fulvous.
Underside—central band extending to the costa, but inter-
rupted near it by the nervures, which are fulvous, otherwise as
above; base silvery, crossed by two pinkish bands margined
with brick-red, the outer one lying close to the central band,
except at the costa of the front wings; base of hind and cell of
front wings crossed by an oblique brick-red dash : front wings
with a subapical patch, creamy above, white and very narrow
below, interrupted by the nervures ; apex fulvous ; the remainder
of the apical half of the front wings pale brown, interrupted by
the nervures, and varied with white lunules between the sub-
costal and second and third median nervules: apical half of
hind wings violaceous outwardly, pale brown inwardly, the
brown portion crossed longitudinally by a brick-red fascia; a
lunulate, submarginal, fulvous line along the hind margin, an
orange spot at the anal angle, and a small black lunule near it.
Body dirty cream-coloured.
Expanse of wings 2 inches.
Hab. Honduras.
Closely allied to Heterochroa Iphicla, Linn., differing from it
above in having the central band produced above the first me-
dian nervule, the subapical orange patch much more angular,
and the submarginal bands more interrupted ; below, the basal
bands are wider apart, the white submarginal lunules are fewer
in number, the submarginal bands less curved, more regular,
and not so much interrupted, the anal orange patch on the hind
wing is much nearer to the margin, and the central band much
more regular in outline.
XLIV.—Remarks on Prof. H/J. Clark’s Peridinium cypripedium,
By H. J. Carrmr, F.R.S. &e.
In the last two Numbers of the ‘ Annals’ (viz. 94 and 95) are
contained the description and illustrations of an animalcule
called by Professor Clark Peridinium cypripedium ; and no mi-
croscopical inquirer into such organisms can have read it, in
connexion with his figures, without admiration and hope of
future contributions of the kind from the same author.
Prof. Clark, however, not unlike those who have preceded him
in such investigations, has confounded two kinds of infusoria,
which, although extremely alike, nevertheless belong, one to the
animal, and the other to the vegetable side of the imaginary
line which divides the two great kingdoms of organized beings.
Nor would this confusion haye been made had the authors of
4.00 Mr. H.J. Carter on Peridinium cypripedium.
that most excellent work to which Prof. Clark alludes, viz. ‘ Les
‘tudes sur les Infusoires, &c.,’ Messrs. Claparede and Lach-
mann, been able to contribute as much to the description of
their second family of Vorticellina as they wished.
This has now been added by Prof. Clark himself so completely
that henceforth no such confusion can exist. But while Prof.
Clark’s mistake serves to show how very like Urocentrum Turbo,
Ehr. (which I believe to be Prof. Clark’s Peridinium cypripedium)
is to Peridinium, it also affords me the opportunity of pointing
out more strongly than has hitherto been done, the striking
resemblance between these two Infusoria, situated on different
sides of the line mentioned, which, from a late examination of
the former, appears to be desirable.
Prof. Clark’s Peridinium cypripedium, if not identical with
Ehrenberg’s Urocentrum Turbo, seems to differ from it so slightly
that it can hardly be termed another species.
Of Urocentrum Turbo Claparede states (op. cit. p. 134), among
other characters, that the mouth or buccal cavity is spiral, and,
on behalf of his lamented coadjutor, the late M. J. Lachmann,
that the anal orifice is posterior; while Ehrenberg asserts, on
the contrary, that the mouth is “not spiral ” (Micrograph. Dict.).
Claparéde also adds that the setaceous or large ciliary appendage
is composed of “long cilia agglomerated into a bundle.”
Urocentrum Turbo has been placed by Ehrenberg among his
Vorticelle ; but if the mouth be not spiral, as he has stated, and
the anal orifice be posterior, as observed by Lachmann—while
Prof. Clark, in his minute examination, although unable to de-
termine the position of the latter, mentions nothing spiral about
the mouth or buccal cavity, and my own observations are of a
like nature—then such testimony is opposed to placing this
organism in the position assigned to it by Ehrenberg.
On this account, probably, the intelligent authors of ‘ Les
Ictudes’ have made a separate family for it, under the head of
“Urocentrina,” which they have placed between their Vorticellina
and Oxytrichina, probably also seeing, among other things, that
the setaceous or large ciliary appendage, by its brush-like form,
allied it more strongly to the Oxytrichina (characterized by
large ciliary feet) than to any other family of Infusoria.
While, then, Urocentrum differs so much from the ciliated
animalcules, on the animal side, as to afford the type of a sepa-
rate family, it so nearly resembles Peridinium, on the vegetable
side, that Prof. Clark has set it down as one of the latter—an
oversight which needs explanation, lest the organization of Uro-
centrum Turbo should be applied to that of the Péridimiens,
By reference to the ‘ Annals’ (1859, vol. i. p. 15) it will be
observed that I have endeavoured to clear up the confusion which
Mr. H. J. Carter on Peridinium eypripedium. 401
Ehrenberg and Dujardin made by mixing up the Trachelius tri-
chophorus of the former, which is the Astasia limpida of the latter,
with the Huglene; and have proposed that under the latter should
only be included Infusoria of the type Huglena, and under that
of Astasia the type only of Astasia limpida,—EHuglena belonging
to the vegetable, while dstasta belongs to the animal side of the
boundary of the two kingdoms ; Astasia being colourless, pre-
senting an oral and an anal orifice, and taking in crude material
for food, while Huglena is for the most part green, presents no
oral or anal orifice, and cannot be seen to incept crude material.
There are other distinctions between these organisms ; but ge-
nerally they are so much alike that Dujardin placed the whole
under the head of Euglene; so that if the organization of Asta-
sia limpida were attributed to Huglena, the same erroneous view
respecting the organization of the latter would arise as that
which would be caused by attributing the organization of Uro-
centrum to Peridinium. Claparéde has also some observa-
tions (p. 346) bearing on the subject; but they were written
unknown to me, if not probably published subsequently to mine.
For my own, 2n extenso, I must refer the reader to the volume
of the ‘Annals’ already mentioned.
Now, Urocentrum Turbo bears a similar resemblance to Peri-
dinium that Astasia linpida (Duj.) bears to the Huglene. Hence
the object of this communication.
As yet, Peridinium must be viewed as closely allied to Huglena
(see my description of Peridinium sanguineum, which imparts ¢
red colour to the sea round the shores of the island of Bombay,
‘ Annals,’ 1858, vol. i. p. 258). It has a large cilium, which
does not appear to be composed of a lash of hairs; a reddish
eye-spot, which may be double or quadruple, according with the
number of divisions which the’ organism may be undergoing
within its lorica, but always connected with a hyaline space,
which in Huglena is seen to be the contracting vesicle; a nu-
cleus, and vesicles of green or otherwise coloured chlorophyll
(see the communication to which I have last referred) ; but no
oral or anal orifice, and no appearance of being supported by
the inception of crude material for food.
On the other hand, Urocentrum Turbo is colourless, has an
oral and probably anal orifice; incepts crude material for food,
as pointed out by Prof. Clark; presents a circular and a median
groove, but with the crown of minute cilia not situated along
the line of the former, as in the cinctum of Peridinium, but in
front of it, and the large cilium, although issuing from the me-
dian groove, composed “of a bundle of hairs, instead of a single
filament. Yet this organ serves to anchor Urocentrum, just as the
simple filament can arrest the progress of Peridinium, while the
Ann. § Mag. N. Hist. Ser. 8. Vol. xvi. 27
402 Mr. J.S. Baly oa new Genera and Species of Gallerucide.
latter swims with its small, and the former, Vorticella-like, with
its large end foremost.
There is no lorica in Urocentrum; and, according to Prof.
Clark, its surface, excepting the crown, is scattered over with
small cilia, in which it further differs from Peridinium—where,
on the other hand, there is a distinct lorica, and cilia only on
the anterior border of the cinctum.
These are some of the chief differences between the two or-
ganisms, of which the most distmguishing are first named.
It is not my object here to go further into the detail of Uro-
centrum Turbo, nor am I prepared to enter the lists with Prof.
Clark respecting its organization.
My observations were for the most part made cursorily, on a
supply of this animalcule which I found here in a freshwater pool
partly filled with decayed leaves, in July last; but, seemg that
the vesicula (contracting vesicle) was particularly active, I
availed myself of the opportunity of watching it carefully with
reference to the question whether it threw back into the body
or discharged externally its contents, and noted down—(1) that
from a simple globular form it became surrounded with a chap-
let of ten (?) small globular sinuses; (2) on these disappearing,
the vesicula became still more enlarged, contracted, and in its
turn disappeared; and (3) that immediately after this, one or
two vesicles became visible, which, breaking into each other,
gave place to the globular form of the vesicula again, to be fol-
lowed by the same series of changes, and so on,—still further
convincing me that the contents of the vesicula do not return
to the body through the sinuses, but are probably ejected from
it through the cuticula direct, or through some excretory chan-
nel—in this instance close to the posterior extremity, where
the vesicula and probably the anal orifice also, are situated.
Urocentrum was also formerly confounded with Cercaria by
Nitzsch and Bory; but the improved quality of our modern
microscopical instruments no longer admits of such a mistake.
Budleigh Salterton, Devon.
Nov. 7, 1865.
XLV.—Descriptions of new Genera and Species of Gallerucide.
By J.S. Baty, F.L.S.
Subfam. Hauricin a”.
Genus Dramputpra, Gerst.
Diamphidia vittatipennis.
D. oblongo-ovata, convexa, sordide fulva, subopaca, antennis, plaga
frontali, oculis, thoracis maculis septem (4.3 dispositis), scutello,
Mr. J.S8. Baly on new Genera and Species of Gallerucide. 403
genibus, tibiis tarsisque nigris; elytris crebre punctatis, singu-
latim puncto apicali vittisque duabus, una submarginali, plerumque
puncto apicali confluente, alteraque subsuturali, ante apicem desi-
nente, basi dilatata, nigris.
Long. 42 lin.
Hab. Damara Land, South Africa.
Head coarsely punctured; apex of jaws and a quadrangu-
lar patch between the eyes black: antenne robust, slightly
compressed, not serrate. Thorax more than twice as broad
as long, coarsely punctured; on the surface are seven black
patches, which resemble in shape and are arranged in a similar
way to those on the thorax of D. flabellicornis, the front row
consisting of four spots, of which the outer two are minute ; the
hinder row is formed of three patches, the outer two of which
are large and irregular, the middle small and oblong.
Diamphidia flexuosa.
D. robusta, convexa, flava, subnitida, antennis, verticis maculis
duabus, thoracis maculis septem (4 . 3 dispositis), scutello, geni-
bus, tibiis, tarsis pleurisque nigris ; elytris postice paullo ampliatis,
crebre punctatis, singulis apice plaga transversa subapicali, fasciis-
que latis flexuosis duabus, una ante, altera vix pone, medium dis-
positis, nigris.
Long. 64 lin.
Hab. Zulu country, Port Natal.
Very closely allied to D. ornata, but larger and more robust
than that insect ; antenne similar in form; sides of the thorax
more regularly rounded.
Genus Popont1A, Dalman.
Podontia evanida.
P. oblongo-ovata, convexa, subtus pallide fulvo-picea, nitida, supra
sordide flava; thorace remote punctato, pallide rufo-piceo varie-
gato, apice utrinque longitudinaliter sulcato; elytris sat profunde
punctato-striatis, punctis pallide piceis, interspatiis ad apicem con-
vexis, puncto humerali nigro, limbo inflexo nigro-tessellato; femori-
bus posticis vaide incrassatis.
Long. 33-4 lin.
Hab. Damara Land, South Africa.
Larger, narrower, and more parallel than P. stolida, with
which insect it otherwise agrees very closely.
Podontia marmorata.
P. ovata, convexa, pallide picea, nitida; antennis extrorsum infus-
catis ; thorace minute granuloso, teuuiter punctato, apice utrinque
longitudinaliter impresso; elytris flavis, rufo-piceo marmoratis,
27*
404 Mr. J.S. Baly on new Genera and Species of Gallerucidee.
profunde punctato-striatis, punctis piceis, interstitiis apicem versus
convexis, limbo inflexo piceo-tessellato ; scutello rufo-piceo; fe-
moribus posticis valde inerassatis.
Var. A. Elytris rufo-piceis, flavo irroratis.
Mas. Gdeagus vix curvatus, apice ovatus, apice ipso subacuminato.
Long. 3-33 ‘lin.
Hab. Port Natal.
More ovate than P. evanida; very similar in form and size to
P. stolida: from the latter species it may be known by the
coarser and rather more distant punctures on the elytra.
Podontia nigrotessellata.
P. ovata, convexa, pallide picea, nitida; thorace subremote punc-
tato, basi biimpresso, antice utrinque longitudinaliter sulcato ;
elytris sordide fulvis, sat profunde punctato-striatis, punctis piceis,
interspatiis remote nigro tessellatis, ad apicem convexiusculis ;
femoribus posticis valde incrassatis.
Mas. Cideagus vix curyatus, lateribus parallelis, apice lanceolato-
ovato.
Long. 23-22 lin.
Hab. ee Natal.
The much smaller size and the different coloration of the
elytra will without difficulty distinguish this species from its
congeners.
Podontia reticulata,
P. anguste oblonga, convexa, pallide piceo- fulva, nitida, antennis
extrorsum infuseatis, ¢ capite scutelloque rufo-piceis ; therace piceo-
marmorato, apice utrinque longitudinaliter impresso ; elytris flavis,
rufo-piceo reticulatis, profunde punctato-striatis, punctis piceis,
limbo inflexo piceo-tessellato ; femoribus posticis valde incrassatis.
Long. 33 lin.
Hab. isan Africa.
Larger and more parallel than P. séolida, more convex than
P. evanida. I only know a single specimen of this species.
Podontia flava.
P. elongata, parallela, convexa, flavo-nitida ; thorace basi biimpresso,
ad latera foveolato, utrinque infra medium ad apicem longitudi-
naliter suleato ; elytris ante medium gibbosis, sat fortiter punctato-
striatis, interspatils ad apicem convexiusculis ; femoribus posticis
modice incrassatis.
Long. 6 lin.
Hab. Sarawak.
This species may be at once known from P. lutea, Oliv., by
the unicolorous body and legs, as well as by the coarser punctua-
tion of the elytra.
Mr. J.S. Baly on new Genera and Species of Gallerucide. 405
Podontia Dalmani.
P. elongata, parallela, postice vix attenuata, pallide picea, nitida;
thorace basi biimpresso, disco medio longitudinaliter sulcato,
utrinque late foveolato, antice utrinque (a margine apicali ad
foveam lateralem) longitudinaliter impresso ; elytris ante medium
gibbosis, fortiter punctato-striatis, uigro-piceis, flavo irroratis,
singulatim plaga basali, fascia lata prope medium alteraque ante
apicem flavis, punctis super partes flavescentes piceis ; femoribus
posticis modice incrassatis.
Long. 6 lin.
Hab. Lombok, Siam.
Podoniia rufo-castanea.
P. subelongata, convexa, subtus cum antennis (his basi excepta)
nigra, nitida, corpore supra et thorace infra rufo-castaneis ; thorace
basi obsolete biimpresso ; disco utrinque profunde foveolato ; ely-
tris aute medium gibbosis, distincte sed tenuiter punctato-striatis ;
femoribus posticis modice incrassatis.
Long. 6 lin.
Hab. India.
This and the preceding insect were originally described by
me as varieties of P. 14-punctata (Journ. Ent. i. p.451). Sub-
sequent investigation, however, has convinced me that they are
entitled to the rank of distinct species.
Podontia congregata.
P. oblonga, convexa, pallide rufo-piceo, nitida, capite elytrisque ful-
vis ; his ante medium gibbosis, punctato-striatis, maculis numerosis
rufo-piceis, preesertim ad latera et apud suturam congregatis, or-
natis; thorace ad latera foveolato, basi breviter bisulcato, ante
medium utrinque longitudinaliter impresso; femoribus posticis
modice incrassatis.
Long. 55 lin.
Hab.
Bs
Subfam. GALLERUCIN &.
Genus Dircema, Clark.
The Rev. H. Clark, in his description of Dircema cinctipenne,
has included five or six closely allied but distinct species: their
diagnoses (together with those of several others entirely new) I
have given below. The males of Dircema may be known from
having the apices of the intermediate tibize armed with a short
tooth or acute process; both sexes have the anal segment of the
abdomen notched ; in the ¢ this notch is large and angular, its
extreme apex being impressed with a distinct fovea; in the 9
the notch is smaller, somewhat variable in shape, although usu-
ally lear, and the apical fovea is obsolete.
406 Mr.J.S. Baly on new Genera and Species of Gallerucide.
Dircema discoidale.
D. elongatum, convexum, dorso subdeplanatum, flavum, nitidum,
vertice, fronte, mandibulis antennisque nigris, his apice sordide
albidis; thorace longitudine plus duplo latiore, lateribus ante me-
dium modice ampliatis, transversim sulcato, utrinque foveolato,
dorso fere glabro, remote punctato, nitido, nigro, marginibus basalt
et laterali flavis; elytris thorace multo latioribus, fere parallelis,
convexis, dorso subdepressis, granulosis, metallico-cyaneis, sub-
nitidis, pube brevissima vestitis, sutura anguste margineque flavis;
femoribus quatuor anticis dorso fusco-lineatis.
Mas. (deagus basi incrassatus, curvatus, lateribus parallelis, apice
ovato-lanceolatus, apice ipso producto, recto, truncato.
Long. 5 lin.
Hab. Nauta, Peru.
Forehead impressed with a large obovate depression, coarsely
punctured, sparingly clothed with coarse hairs; eyes ovate-
rotundate ; four upper joints of antennze pale fulvous. Apical
border of thorax concave, entire, side-margin moderately dilated
in front, nearly straight and parallel behind, hinder angles
acute; disk shining, sparingly clothed with suberect hairs ;
transverse excavation broad, less deeply impressed than in most
of the other species; it is, however, rendered deeper on either -
side by a large roundish fovea, which is placed about halfway
between the middle and the lateral border. Elytra shorter than
in the other species, much broader than the thorax, depressed
above.
My collection contains two male specimens of this species.
Dircema laticolle.
D. elongatum, parallelum, convexum, fulvo-flavum, nitidum, capite
(facie inferiore excepta) nigro, antennarum articulis tribus ultimis
sordide flavis; thorace brevi, longitudine triplo latiore, lateribus
ante medium late explanatis, disco minus crebre, medio subremote
punctato, pube depressa nigra sparsius vestito, profunde trans-
versim excavato, pone sulcum distincte elevato; elytris nigris,
opacis, rude granulosis, pube adpressa fusca aut nigra vestitis,
sutura margineque flavis; pedibus nigris, femoribus basi et infra
fulvis.
Fem. Abdominis segmentum anale apice obtuse angulatum, medio
vix emarginatum.
Long. 33-4 lin.
Hab. Amazons.
Forehead depressed, rugulose, vertex nitidous ; epistome im-
pressed with a deep longitudinal groove, which extends from the
apical border to the encarpze, its upper half very strongly marked;
three upper joints of antennz obscure fusco-fulvous. Thorax
short, quite thrice as broad as long; apical margin concave,
Mr. J.S. Baly on new Genera and Species of Gallerucide. 407
often slightly emarginate in the middle; lateral margin broadly
dilated in front, hinder portion subparallel, hinder angles acute,
slightly produced; upper surface smooth and nitidous, some-
what sparingly clothed with pale hairs; transverse sulcation
deeply impressed, the surface of the thorax behind its posterior
margin thickened and distinctly raised; in the centre of this
raised surface is a more or less distinct longitudinal groove.
Elytra broader than the thorax, nearly subcylindrical in the 3;
rather broader and more depressed in the ? .
Dircema cinctipenne, Clark.
Ann. & Mag. Nat. Hist. Oct. 1865, p. 263.
D. elongatum, parallelum, convexum, flavum, nitidum, capite (facie
inferiore excepta) nigro, antennis apice sordide flavis aut albidis ;
thorace brevi, longitudine fere triplo latiore, lateribus ante medium
late explanatis, disco ruguloso, pube adpressa sat dense vestito,
medio longitudinaliter canaliculato, profunde transversim excavato,
pone sulcum distincte elevato, utrinque plaga magna nigra ornato ;
elytris opacis, granulosis, pube depressa fusca aut nigra dense ves-
titis, nigro-viridibus aut obscure viridi-metallicis, sutura margine-
que flavis ; pedibus fusco-nigris, rarius obscure flavis, fusco
lineatis.
Mas. Cideagus basi incrassatus, curvatus, anguste lanceolatus, apice
producto, subacuto, recurvato.
Fem. Abdominis segmentum anale apice modice incisum.
Long. 4—43 lin.
Hab. Amazons.
Head short ; front clothed with adpressed hairs, with a more or
less distinctly defined ovate depression, which is more or less finely
rugulose ; vertex nitidous ; face grooved as in D. Jaticolle ; three
or four upper joints of antennee yellowish white. Thorax three
times as broad as long; apical margin concave, sometimes
slightly notched in the middle; side margins broadly dilated in
front, nearly straight and slightly diverging posteriorly, hinder
angles acute ; disk impressed in a similar manner to that of D.
laticolle, but rugose-punctate, more closely covered with pu-
bescence. Elytra similar in form to those of D. laticolle, dark
metallic green, subopaque.
Dircema pulchrum.
D. elongatum, pallide flavum, nitidum; fronte, vertice antennisque
nigris ; his apice albis, articulo basali flavo ; thorace brevi, longi-
tudine triplo latiore, lateribus late ampliatis, disco profunde trans-
versim excavato, minus crebre punctato, sparse pubescente, utrin-
que plaga nigra ornato ; elytris pube adpressa brevi vestitis, sub-
opacis, rugosis, metallico-viridibus, margine suturaque flavis ;
femoribus dorso nigro-piccis ; tibiis ad apicem tarsisque fuscis.
408 Mr. J. 8. Baly on new Genera and Species of Gallerucidee.
Mus. Gdeagus apice lanceolato-ovatus, lateribus ad apicem bi-
sinuatis, apice ipso vix producto, truncato, recurvato.
Fem. Abdominis segmentum anale utrinque foyeolatum, apice angu-
lato sat profunde inciso ; thorace toto flavo ; antennis totis, femori-
bus dorso, tibiis tarsisque nigris.
Var. A. (fem.). Elytris obscure nigro-purpureis.
Long. 5-6 lin.
Hab. Nauta, Upper Amazons.
This species, although of larger size and brighter coloration,
very closely resembles D. daticolle, more especially in the form
and punctuation of the thorax; the ? may be known by the
linear notch at the apex of the abdomen, the ¢ by the paler legs
and the brighter colour of the elytra. Front with a large semi-
ovate rugulose depression ; vertex nitidous.
I possess three specimens of this species, one ¢, two 9.
Dire2ma columbicum.
D. clongatum, parallelum, sordide flavum, nitidum, vertice, fronte
(hoc maculam flavam includente), antennis, femoribus dorso, tibiis
tarsisque nigris; thorace longitudine duplo latiore, lateribus ante
medium modice ampliatis, disco modice sed late transversim ex-
cavato, crebre punctato, pube adpressa dense vestito, utrinque
macula ovata nigra ornato; elytris metallico-viridibus, subopacis,
pube subdepressa brevi vestitis, margine suturaque flavis.
Tom. Abdominis segmentum anale apice obtuse angulatum, breviter
incisum.
Long. 6 lin.
Hab. Bogota.
I only know a single specimen of the present species (a ? ).
It may be easily separated from all its congeners by the peculiar
form of the thorax ; this part is less transverse, beg only about
twice as broad as long; its sides in front are less dilated, and
the upper surface much less deeply excavated ; the hinder angles
are produced into an obtuse tooth. The elytra are much broader
than the thorax, somewhat depressed above. Face long, clothed
with pale hairs ; vertex and front black, the latter nearly plane,
marked in the centre with a transverse yellow patch.
Dircema letum.
D. elongatum, parallelum, convexum, dorso subdepressum, sordide
fusco-fiavum, nitidum, vertice antennisque nigris, illo rugoso, his
basi et apice flavis ; thorace longitudine vix duplo latiore, lateribus
ante medium modice ampliatis, disco late transversim excavato,
crebre rugoso, pube brevi adpressa dense vestito, sordide flavo,
utrinque plaga magna nigra ornato; elytris rugoso-granulosis,
pube adpressa brevi fusco-fulva dense vestitis, viridi-zeneis, sutura
Mr. J.8. Baly on new Genera and Species of Gallerucidee. 409
marginibusque anguste flavis; pedibus nigris, femoribus basi et
infra sordide flavis.
Mas. Gideagus basi incrassatus, curvatus, parallelus, ad apicern acute
angulatus, apice producto, truncato, recurvato.
Fem. Abdominis segmentum anale apice obtuse angulatum, sat
profunde incisum.
Long. 43-5 lin.
Hab. Banks of the Napo, Ecuador.
Thorax rather less broadly dilated than in D. cinetipenne, its
upper surface much less deeply excavated. Elytra pale, almost
sea-green, fusco-sericcous, broader and more depressed than in
D. cinctipenne; more nearly approaching in habit to D. colum-
bicum, but from that species the smaller size and differently
formed thorax will separate it; three upper joints of antennz
yellowish white, the remainder more or less fulvous beneath.
I obtained three specimens of this pretty species (two ¢ and
one ?) from the late Mr. H. Cuming, who received them from
one of his correspondents in Keuador.
Dircema sordidum.
D. elongatum, parallelum, convexwm, fulvo-fuscum aut sordide ful-
vum, nitidum, femoribus dorso, tibiis tarsisque piceis, vertice
fronteque viridi-nigris ; antennis nigro-piceis, basi externa et apice
fulvis ; thorace longitudine triplo latiore, lateribus ante medium
late explanatis ; disco rugoso, profunde transversim excavato,
pone sulcum elevato, pube adpressa dense obsito, nigro, vitta lata
centrali et margine apicali dilatato flavis; elytris rude granulosis,
obscure metallico-viridibus, pube adpressa fusca dense vestitis,
sutura margineque obsolete flavo-marginatis.
Fem. Abdominis segmentum anale apice obtuse angulatum, semi-
circulariter incisum,
Long. 4 lin.
Hab. Kga and Nauta, Upper Amazons.
Very similar in form, both of thorax and elytra, to D. cincti-
penne ; easily distinguished from that insect by the coarser and
closer pubescence of the upper surface, and by the extremely
narrow and almost obsolete fulvous margination of the elytra ;
the notch on the apex of the anal segment of the abdomen in the
? is semicircular, instead of lmear, as in D. cinctipenne.
I know two specimens of the species, both 9. The specimen
from Ega has the scutellum dark fuscous.
Direema modestum.
D. elongatum, parallelum, convexum, dorso subdeplanatum, flavum,
nitidum, verticis macula bilobata antennisque nigris, his apice
flavo-albis ; thorace longitudine fere triplo latiore, lateribus ante
410 Mr.J.8. Baly on new Genera and Species of Gallerucide.
medium sat late ampliatis, disco profunde transversim sulcato,
pube adpressa sparse obsito, nitido, subremote punctato; elytris
convexis, dorso subdepressis, rude granulosis, subremote punctatis,
fusco-sericeis, murinis vel pallide viridibus, opacis ; genibus,
tibiis, tarsisque nigris.
Mas. (deagus basi incrassatus, curvatus, parallelus, ad apicem ovato-
lanceolatus, apice ipso producto, capitato.
Fem. Abdominis segmentum anale apice obtuse angulato, breviter
angulariter inciso.
Long. 5 lin.
Hab. Magdalena River, Columbia.
Broader and flatter above than D. cinctipenne, closely resem-
bling D. nigripenne, Fab., in form and unicolorous elytra. From
this latter species it may be at once known by the bilobate black
patch on the vertex, as well as by the paler elytra; the males of
the two species differ also in the form of the cedeagus ; the apex
of this organ, instead of being capitate as in the insect before us,
is produced in D. nigripenne into a broad slightly recurved pro-
cess, the extremity of which is truncate and slightly notched,
the lobes of the notch being also reflexed ; the females may be
separated by the difference in the apical notch of the last abdo-
minal segment: in D. nigripenne this notch is linear, a longi-
tudinal groove extending from its apex to the base of the seg-
ment; in D. modestum the notch is short and slightly angular,
the longitudinal groove being entirely obsolete. The head of
D. modestum is rather longer than in any of the other species
described in the present paper; the front is impressed with the
usual semiovate space.
Genus HyPpHa@NIA.
Corpus angustatum, parallelum, modice convexum. Caput ex-
sertum ; facie perpendiculari ; antennis filiformibus, 3 seepe cor pore
longioribus, gracilibus ; articulis primo curvato, a basi ad apicem in-
crassato, secundo brevi, czeteris cylindricis, subtus in ¢ seepe setis erectis
vestitis, singulis longitudine primo fére eequalibus ; secundo, tertio
quartoque interdum subtus compressis et dilatatis; 9? corpore
paullo brevioribus, articulis nunquam setiferis; ocwlis magnis, pro-
minulis ; pa/pis maxillaribus ovatis, apice acutis. Thorax transverso-
quadratus, lateribus rectis, angulis singulatim tuberculo setifero
armatis; dorso subdeplanato, medio transversim sulcato. Elytra
thorace paullo latiora, elongata, parallela, confuse punctata. Pedes
graciles, subelongati; covis anticis erectis, contiguis ; femoribus pos-
ticis non incrassatis; ¢éb¢is omnibus apice muticis; ¢arsorum posti-
corum articulo primo duobus sequentibus conjunctis eequali; un-
guiculis appendiculatis. Prosternum fere obsoletum.
Type Hyphenia (Luperus) pilicornis, Motsch. Birmah.
Dr. Mérch on the Limits of the Subkingdom Mollusca. 411
XLVI.—On the Limits of the Subkingdom Mollusca.
By Dr. O. A. L. Morcu.
Tue Vertebrata form the only subkingdom with uncontroverted
limits ; yet the late Prince Bonaparte considered the Aphanozoa
(Sagitta, &c.) the lowest Vertebrata.
Since the time of Cuvier, the Tunicata have been considered
by zoologists true Mollusca. Professor Huxley has shown that the
Bryozoa are inseparable from the Tunicata. The Brachiopoda
are considered allied to the former, thus forming the subclass
Molluscoidea of several modern authors. Dr. Macdonald* has
lately added the Ctenophora + to the Brachiopodat.
Prof. Lovén first raised a doubt as to the affinity of the Tuni-
cata to the Mollusca, chiefly on account of the form of the larva
of Ascidians. Dr. Macdonald founded the same doubt chiefly
on the reproduction by budding, which is foreign to the nature
of Mollusca. This reason alone cannot be considered sufficient,
as there is found in nearly every class a division (subclass, mi-
crosthenic division) differimg in having an abortive progeny, e. g.
Marsupialia, Amphibia, Orthoptera among Mandibulata, and He-
miptera and Homoptera among Haustellata, Entomostraca, &c.
The systematic position of the Tunicata seems chiefly to be
based on a supposed resemblance between the two apertures of
Ascidians with the siphons of Acephala, and a still more super-
ficial resemblance between “the shells.”
Another division, contaiing gelatinous and arborescent forms
(the Acalephze), seems to have a much greater affinity to the
Acephala, with which they agree in the four labial palpi and the
fringed oculiferous edge of the mantle. This resemblance is
very striking if the Acaleph be compared with a swimming
Lima, as mentioned by Sars. The radiated form indicates the
same relation to Acephala as Cephalopoda bears to Gasteropoda.
The group Radiata as defined by Cuvier do not really exist.
Cnidz and phlebenterism are found in the Gymnobranchia to a
great extent, bemg dependent on the want of a shell§. ‘The An-
thozoa, considered inseparable from the Acalephe, look no
stranger among the Mollusca than the Bryozoa.
There are, however, some other Coelenterata which may be
referred to the Mollusca with still more precision. I have
always thought it strange that the Mollusca, although inferior to
* Transactions of the Royal Academy of Edinburgh, 1864.
+ Morch, Fortegnelse over Gronlands Bloddyr, 1857; Rink, Grén-
land, p. 97.
t Malacozoologische Blatter, vi. p. 104.
§ In Cyanea is found a brown membrane reminding us of the Loligo-
shell.
412 Dr. O. A. L. Moreh on the Limits
the Arthropoda, contained only a single true intestinal species
(the Hndoconcha of John Miiller), whilst the “Entozoa” have
contributed considerably to the Arthropoda, viz. Lernea, Pelto-
gaster, Clistosaccus, Linguatula.
The Helmintha are generally divided into two sections,—viz.
Nematodes with distinct sexes, and Platyhelmia with the sexes
united. Mr. Bastian* has, at the last meeting of the Royal
Society, on the 15th of June, pomted out that the Nematodes,
parasitic and free, are closely allied to the Echinodermata.
The Platyhelmia, contaming the Trematoda, Cestoda, and Tur-
bellaria, Seem to me not to differ materially from the androgy-
nous Mollusca.
The Turbellarians are united to the Mollusca by Mr. Ch.
Girard}, on account of the resemblance of the embryonic state
of Planocera elliptica with that of Hlysia. It must, however, be
remembered that the ‘ chrysalis” state is not known among the
Mollusea, and that all larvee of Gasteropoda are provided with a
nautiloid shell, except the larva of Chiton. The Turbellarians
differ chiefly from the Gasteropoda in the presence of numerous
eyes; but this difference is perhaps not of great importance, as
Dr. Bergh has discovered four eyes in a species of Midiana from
St. Thomas, and Agassiz{ indicates that Margarita 1s provided
with eyes, at the base of the pedal filaments, corresponding to
those in the mantle-edge of Pecten and other Acephala. Dr.
Bergh, however, has not found any lens in the “eyes” of the
pedal filaments in Margarita. According to Prof. Hensen, the
lens is absent in Nautilus.
The nervous system shows no important difference from that
of Gasteropoda. The want of a trne foot is of no consequence.
In Bulla§, Akera, &e., the locomotion is partly, and in Pelh-
branchiata almost entirely, effected by the hind part of the body,
the true foot being reduced to a crescent-lke disk, not unlike
the sucker in many Entozoa. The arborescent form of the in-
testinal canal, generative organs, and kidneys is common to
all animals wanting special respiratory organs.
Salivary glands are present, but the liver is entirely absent, as
is usually the case in animals (e. g. perfect Insects) living upon
food not requiring preparation. ‘The Zenie want even a ‘mouth
and intestinal canal, living im a prepared nutrimental juice,
2) .
which the parasite absorbs through the surface of its skin; a
* The Atheneum, June 1865, p. 850. [Annals for September,
Foal (i
+ Researches upon Nemertians and Planarians. Philadelphia, 1854.
+ Lectures on Embryology, p 86, 8vo.
§ Mérch, “Contributions & la Faune malacologique des Antilles
danoises,”’ Journ. de Conch. 1863, p. 19.
of the Subkingdom Mollusca. 418
circulatory system, therefore, would be of no use. The Platy-
helmia seem to me not to differ more from the Mollusca than
Peltogaster and Clistosaccus from the Crustacea. In the ento-
zoic Crustacea some of the harder oral parts are generally pre-
served, in order to serve the animal for attachment. It is there-
fore likely that the hooks of Cestoda are homologous with simi-
lar hard organs of Mollusca. Gegenbaur* compares in this
way the four uncigerous retractile organs of Tetrarhynchus with
the cheek-hooks (harpagz) of the Pteropoda gymnosomata.
Perhaps a closer study of the genus Homoderma of Van Beneden +
(outwardly resembling a Distoma, but anatomically agreeing
with Pneumodermon) may throw some light on this question.
The hooks of the Cestoda may also be compared with the
lingual teeth of the Pellibranchiata, and chiefly with that part
called by Messrs. Alder and Hancock { the “prehensile collar ”
in Limapontia nigra. These hooks probably assume a circular
arrangement. ‘the lingual teeth are frequently absent among
Gasteropoda, e. g. Doridopsis (Hancock), Cirroteuthis and Tethys
(according to Bergh).
A secretory organ (kidney) is never absent in any animal
taking food ; and it therefore exists in all Entozoa, agreeing in
form with the renal organ of Elysia as represented by Souleyet.
The generative organs of the Platyhelmia agree exactly with
those of the androgynous Mollusca in general, but chiefly with
those of the genera § Chalidis and Pelta; at least I cannot dis-
cover any notable difference on comparing the anatomy of the
two latter genera with that of Amphistomum subclavatum as
represented by Dr. Walier||.. The Huirudines and Lumbrici,
which are considered by Prof. Quatrefages distinct classes, are
also androgynous, with reciprocal copulation; but the gene-
rative organs, like the nervous system, seem to be formed
according to quite another plan, although they may have some
distant resemblance to those of H/ysia. It may be questioned if
androgyny is of sufficient importance to waite animals differing
so greatly in their exterior form, especially as hermaphroditism
is only of specific value among Fishes (Serranus scriba) and
Acephala (Pecten opercularis). I, however, believe that andro-
* Vergleichende Anatomie, p. 329.
+ Exercices zootomiques, 1839, p. 54, t. 3. f. 4-6 (Mém. de l’Acad.
Roy. des Sciences de Bruxelles, tome xi.)
{ Ann. Nat. Hist. ser. 2. vol.i. p. 208, pl. 20. f. 2. Quite different from
“the prehensile collar” of the same authors in their ‘ British Nudibran-
chiata.’ ;
§ Quatrefages, Mémoire sur les Gastéropodes phlébentérés (Ann. Se.
Nat. 3 sér. 1844, tome i. p. 155, t. 4. f. 4); Morch, ‘‘ Contributions a la
Faune malacologique des Antilles danoises,” d. e. p. 24.
|| Archiv fir Naturgeschichte, xxiv. p. 269, t. 11. f. 5.
414, Prof. T. R. Jones and Dr. H. B. Holl on
gyny is of the highest importance in connexion with the other
characters mentioned above, as opposed to the vermicular form
and white eolour—characters common to most burrowing ani-
mals and plants.
If this proves correct, there will only remain among the
Coelenterata Lumbrici, Hirudines, Annelida, Echinodermata*,
Nematoidea, Tunicata, Bryozoa, Brachiopoda, Ctenophora, Rhizo-
poda, Spongiaria. It seems very doubtful whether these divisions
together would correspond to the three other subkingdoms.
The limits of the second subkingdom, Arthropoda, are much
litigated, as the lowest Crustacean does not show the same
degradation of the copulatory organs as the Fishes among Ver-
tebrata, and the Acephala among the Mollusca. I therefore
consider it probable that the subkingdom Articulata, chiefly
on account of the nervous system, ought to be reestablished
as it was defined by Cuvier.
XLVII.—Notes on the Paleozoic Bivalved Entomostraca. No. V1.
Some Silurian Species (Primitia). By Professor T. Ruprrr
Jonrs, F.G.S., and Dr. H. B. Hott, F.G.S
[Plate XIII.]
In the ‘ Annals of Natural History’ for August 1855, September
1855, and April 1857 were published descriptions of some
Sihimienn Bivalved Entomostraca, comprising, among others,
Beyrichie of three types,— simplices,” “ corrugates,” and
“jugose.” The first of these groups, the simple or unisulcate,
seems to us now to be deserving of generic distinction, since,
among a still larger number of forms, we find a persistent oc-
currence of the chief features, with a passage towards Leperditia,
by the complete loss of the furrow, rather than towards the two-
furrowed or real Beyrichie.
We do not presume, however, that we hereby do more than
somewhat improve our classification of these necessarily obscure
Silurian Entomostraca, represented only by carapace-valves,
always minute and often variable in form and ornament, besides
being subject to alterations by pressure and by chemical change,
and rarely to be cleared of their matrix on all sides. In some
cases, too, we have had to be content with what we could make
out of casts and imprints.
There remain, therefore, several difficulties in classifymg these
little Bivaive Entomostraca—and especially since with the total
disappearance of the dorsal sulcus we do not seem necessarily to
* According to Mr. Williamson, closely allied to the Annelida (British
Association Report, 1857).
the Paleozoic Bivalved Entomostraca. 415
enter on a distinct genus, judging from the general shape and
character of some of these minute valves associated with others
that are furrowed and, from similarity of style and structure,
apparently congeneric with them. We allude to the dwarf
forms of Leperditia, on one hand, where relative size is often the
only available basis of distinction (for the eye-spot may be want-
ing, the muscle-spot invisible, and the flanged edges may be
hidden), and, on the other hand, to those true Beyrichie with a
single sulcus, and in which the second sulcus is reduced to a
minimum or altogether wanting (B. arcuata, &e.). In this case
also relative size is distinctive, as well as, perhaps, the difference
of geological horizon.
Another difficulty is found in defining the probable alliances
of some minute Silurian Entomostraca which have been grouped
under “ Cytheropsis,” and which in outline agree with Primitia,
but want the sulcus altogether. It is possible, however, that
the sulcus is not an essential character zoologically, and that the
merely slight impression (as in P. Beyrichiana and P. obsoleta)
leads us to altogether non-sulcate forms : and here the balance of
probabilities, judged of by the general aspect of the specimens,
must be our guide ; and we must still be content with imperfect
classification, if we wish to make our present knowledge of these
little Palzeozoic fossils available. If, therefore, they are to be
catalogued and brought into relationship with their larger con-
temporary allies and their modern representatives, we must ac-
cept and make the most of such features as are apparent, and
give credit for probable divergency in the unpreserved soft parts
when the valves show differences of contour, foldings, and sculp-
ture. Hence we have been induced to value more highly than
formerly the differences in recorded varieties of the so-called
Beyrichia strangulata (Annals Nat. Hist. Sept. 1855), and we
shall offer diagnoses for them accordingly.
PRIMITIA, gen. nov.
Carapace minute, bivalved, either equivalved or nearly so,
convex, more or less oblong, often approaching Leperditia in
shape, by the sloping of the dorsal angles; hinge-line straight,
sometimes nearly as long as the valve. Surface of each valve
usually impressed on the dorsal region, either medially or to-
wards the anterior extremity, with a vertical sulcus, variable in
size, sometimes barely visible, sometimes passing into, or even
merely represented by, a navel-like pit; and sometimes the
sides of the sulcus are swollen, and even raised up into tubercles*,
* With some additional details, the diagnosis for “ Beyrichiz simplices,’
given in Ann. Nat. Hist. ser, 2. vol. xvi. p. 85, serves for Primitia.
416
Prof. T. R. Jones and Dr. H. B. Holl on
Table of the Distribution of Primitia.
Name.
1. P. strangulata, Salter, sp....
i EAVALS Co sncsaenesee ks Sener
2. P. Salteriana, nov. .........
3. P. semicordata, nov. ......
lee SIMPIOK, GIGMES-1.ve cess ee
o. P. Logant, Jones. .25..00:.00-
5*, ——., var. reniformis
one, , var. leperditioides.
6. P. matutina, nov......... Ace
7. P. Seminulum, Jones ......
OP PSigtllatay J OWES acinateea ae
PRA vanrolatas Ove scesses ces
oh , var. paucipunctata .
10. P. rugulifera, Jones
HV SPS renmlimas 200. cc..0.c00 <4
12. P. mundula, Jones
13. P. nana, nov. .
14. P. bicornis, Jones.........
15. P. umbilieata, nov.
16. P. eristata, nov.
LJ PPS terse. 200. ckiscceos
Iitekew] 2
Oe E
20. P. Roemeriana, nov..........
. obsoleta, nov.
. oblonga, nov.
. Ovata, nov.
semicircularis, n0v.
sPUsilla OD. cs.sesacececs'
trigonalis, nov. .......
Beyrichiana, nov... ..
Cs
Cr
eeetee
. coneinna, Jones ..
- muta, nov.
seen
isuleus and,
.. pit (famt)
_ very faint |
Having
a sulcus
33
no suleus
a sulcus
39
33
33
39
suleus and
tubercles
(faint)
suleus and
tubercles
tubercles
(strong)
pit
>
sulcus
3)
3”
no sulcus
33
33
small sule.)
no suleus
33
Formation.
Lower Silurian
3)
9
Upper Silurian
3)
39
3)
Be
33
ide
} Lower Silur.
ee
Upper Silurian
33
339
Lower Silurian
Upper Silurian
3)
{
Locality.
Lancashire.
Pembrokeshire.
33
Salop.
Portugal and Salop.
Canada
33
59
Salop.
Montgomery.
Beechey Island.
near Malvern.
33
Beechey Island.
near Malvern.
Sweden and near Mal-
vern.
Salop.
near Malvern.
+B)
2)
bP)
Sweden.
near Malvern.
Sweden.
”)
near Malvern,
Canada. [tia.
Gothland; Nova Sco-
Beechey Island.
1. Primitia pias
Beyrichia strangulata, Aun. Nat. Hist. ser.
Salter, sp.
2. vol. xvi. p. 17], pl. 6. fig. 18.
This is one of the largest forms belonging to Primitia, and it
has a larger marginal rim than any other.
Found in ealeareous
schists (Lower Silurian) at Coniston Waterhead, Lancashire.
This species was mentioned, under the provisional name of
“ Cytherina levigata,” by J Mr. Salter in 1845 (Quart. Journ,
Geol. Soe. vol. i. p. 445), as occurring abundantly in the Conis-
the Paleozoic Bivalved Entomostraca. 417
ton Limestone ; but in 1852 (Brit. Pal. Foss. Camb. Mus. Ap-
pendix A, p. ii.) he decided to call it B. strangulata.
Schrenk recognizes this species in the brown bituminous marl
(Brandschiefer) of the Lower Silurian of the Baltic Provinces
(Untersuch. p. 195).
1*, Primitia strangulata, Salter, sp., var. a.
Beyrichia strangulata, var. x, ibid. p. 172, pl. 6. fig. 19.
Differing from the foregoing chiefly by the furrow being faint
and extending far across the valve. In fossiliferous schist (Lower
Silurian) at Robeston Wathen, Pembrokeshire.
2. Primitia Salteriana, nov.
Beyrichia strangulata, var. 8, ibid. fig. 20.
This differs from P. strangulata in the absence of the raised
rim, in the more acute anterior extremity, and in the punctation
of the surface. In fossiliferous schist (Lower Silurian), Sholes
Hook, Haverfordwest.
Schrenk speaks of this variety (but with papille instead of
pits) as occurring in the Lower Silurian Brandschiefer and
Borkholm beds of the Baltic Provinces (Untersuch. p. 196).
He also refers to another form (“ var. crenulata’’), with a broad
and notched border, from the Lower Silurian beds at Paggar
and Borkholm (Untersuch. p. 196).
3. Primitia semicordata, nov.
Beyrichia strangulata, var. 8, young, ibid. fig. 21.
Besides being smaller than any of the above, this is relatively
short and broad, nearly semicircular, and smooth. Accompany-
ing the last mentioned.
4. Primitia simplex, Jones.
Beyrichia simplex, ibid. figs. 25, 26, 27.
This small, smooth, subovate species was first observed in the
Lower Silurian schist of Busaco, near Coimbra, Portugal ; and
the very slightly different forms from a Lower Silurian schist
at Harnage, near Shrewsbury, do not appear to be separable
from it.
5. Primitia Logani, Jones ; et varr.
Beyrichia Logani et vary. reniformis et leperditioides, Ann. Nat. Hist.
ser. 3. vol. i. p. 244, pl. 9. figs. 6-10.
A large number of this gregarious species comprise so many
variations of form, from oblong to reniform, on one hand, and
from oblong to subovate, on the other, with or without puncta-
tion of surface, but always sulcated, that it seems impossible to
draw any lines of distinction, excepting such as may limit the
Ann. & Mag. N. Hist. Ser.3. Vol. xvi. 28
418 Prof. T. R. Jones and Dr. H. B. Holl ox
oblong forms to P. Logani proper (whether long-oblong or
ovate- ‘oblong),— —dividing off the kidney-shaped valves as var.
reniformis, and those that have the most sloping dorsal angles
as var. leperditioides. They all come out of a Lower Silurian
limestone (upper portion of the Calciferous Sandrock) at Gren-
ville and Hawkesbury, Canada.
6. Primitia matutina, noy. Pl. XIII. figs. 7a, 7 &.
Length 5-22,, height +23, inch.
This small, smooth, convex, Leperditia-shaped, non-sulcated
Primitia might be catalog ‘ued as a variety of B. Logani, had we
ever seen a “specimen of the latter without a sulcus, or with an
inclination to lose its furrow. Under existmg circumstances,
however, we give the benefit of the doubt, and the value of
geographical distance, such as it is, to the probability of this
little Lower Silurian form being distinct from its Canadian ally.
The Trinucleus-shale (belonging to the upper part of the Bala
or Caradoc formation) in the river Onny, near Cheney-Longville,
Shropshire*, abounds with such small Entomostraca as this
Primitia ; but we have been able only to pick out this form in
a well-preserved state.
7. Primitia Seminulum, Jones.
Beyrichia Seminulum, Ann. Nat. Hist. ser. 2. vol. xvi. p. 173, pl. 6. fig. 24.
A neat little semicircular form, from the Upper Silurian
schists of Montgomery, where it occurs with Beyrichia Kleedent.
8. Primitia sigillata, Jones.
Beyrichia sigillata, Ann. Nat. Hist. ser. 3. vol. i. p. 242, pl. 9. fig. 5
More oblong than P. Seminulum, less deeply furrowed, and
somewhat more coarsely punctate. From the Upper Silurian
Limestone of Beechey Island, with P. rugulifera, P. muta, and
Leperditia gibbera. P. variolata is the British representative of
P. sigillata.
9. Primitia variolata, nov. PI. XIII. figs. 6 a, 6 4.
Length ;22,, height 32, (as 8 to 2).
Carapace ier convex, varying between suboval and
subquadrate ; rather more obtuse behind than before. Hinge-
line straight, but somewhat overhung by the dorsal part of the
valves. Valves somewhat flattened, impressed with a distinct
dorsal sulcus, which ends, towards the middle of the valve, in a
* An account of this Trinucleus-shale is given im a paper by Messrs.
Salter and Aveline, Quart. Journ. Geol. Soc. vol. x. p. 62, &e. See also
*Siluria,’ 2nd edit. p. 72, &e. The Caradoc and Bala beds are regarded as
belonging to a higher horizon than the Caleiferous Sandrock of Canada,
in which P. Logani abounds.
the Paleozoic Bivalved Entomostraca. 419
subcircular umbilical pit. The surface of the valves is dotted
with coarse shallow pits, and sometimes slopes gradually to the
margins, but is generally deflected abruptly. There is a slight
rim at the border. P. variolata is not far removed from P. sigil-
lata and P. Seminulum ; indeed, it may be said to be the British
representative of the former. From the Woolhope Limestone,
west of the Wych, Malvern.
9*, Primitia variolata, var. paucipunctata, figs. 6c, 6d.
Length +235, height +347 inch.
This bears fewer pock-marks, has a less well defined border-
rim, and a rather smaller sulcus. From the Woolhope Lime-
stone, west of the Wych.
10. Primitia rugulifera, Jones.
Beyrichia rugulifera, Aun. Nat. Hist. ser. 3. vol. i. p. 242, pl. 9. fig. 4.
Oblong, ornamented with minute transverse wrinkles, and
impressed with a broad and deep sulcus on the anterior half.
From the Upper Silurian Limestone of Beechey Island, together
with other small Bivalved Entomostraca.
11. Primitia renulina, nov. Pl. XIII. figs. 5 a, 5b.
Length =, height 1; (as 3 to 2), thickness ;1; inch.
Carapace convex, ovate-oblong, straight on the back, boldly
curved on the other margins, especially backwards; dorsal
angles distinct ; anterior region compressed. Valves smooth,
bilobed, impressed with a deep well-defined dorsal sulcus, and
bordered all round with a narrow depressed rim, which runs
into the sulcus at the back or upper part of the valve. From
the Wenlock Limestone at the Crofts, Malvern.
12. Primitia mundula, Jones.
Beyrichia mundula, Aun. Nat. Hist. ser. 2. vol. xvi. pp. 90 & 174, pl. 5.
fig. 23, & pl. 6. figs. 28-31.
In addition to the diagnosis of this neat little species given
at p. 174, op. cit. we have to allude to the swelling of the sides
of the sulcus in some well-grown individuals, which, however,
are not specifically distinct from others with less developed
sulcus,—also to the beautifully delicate reticulated sculpture of
the surface, which sometimes appears as excessively fine longi-
tudinal wrinklings, with inosculating meshes, and sometimes as
a minute pitting.
Small individuals (long =, high 3; inch) are more oblong
than others, having rounded ends, parallel upper and lower
borders, and a faint sulcus. The normal form approaches that
of Leperditia; and, with a short hinge-line, angular ends, and
convex belly, the valves become almost aval. P. mundula has
28%
420 Prof. T. R. Jones and Dr. H. B. Holl on
some of the characteristics of the Lower Silurian P. dzcornis,
but in a much less degree.
From the Upper Silurian (Scandinavian) Limestone found as
drifted blocks in North Germany. We have seen a small speci-
men in a limestone of the Wenlock Series from near Malvern.
13. Primitia nana, nov.
Beychia strangulata, var. y, ibid. p. 173, pl. 6. fig. 22.
Very small, nearly oblong, but proportionally longer than P.
strangulata. "Edges of anlar swollen into two minute unequal
tubercles. In soft fossiliferous schist (Lower Silurian) at Har-
nage, near Shrewsbury, in company with P. simplex and P. bi-
cornis. It is possibly the young of the latter.
14. Primitia bicornis, Jones.
Beyrichia bicornis, ibid. fig. 23.
Very small, but readily distinguished by its two subcylindrical
tubercles bordering the sulcus on each valve, and by its crested
marginal rim. From the Lower Silurian beds at Harnage, near
Shrewsbury.
15. Primitia unliaat: noy. Pl. XIII: figs. 2a, b7e 7d,
Length 54°,, height ;22,, thickness ;2%., inch.
Carapace convex, rounded-oblong; dorsal angles sharp ;
hinge-line straight, and sunken in an elliptical or acute-ovate
depression formed by the convexity of the dorsal borders of the
two valves; this flattened area is broadest posteriorly, and ter-
minates, near the antero-dorsal angle, by a shght notch. The
hinge itself is formed by the meeting of simple edges. The
ends of the valves are boldly rounded ; “the ventral line is gently
convex; surface of the valves convex, rather compressed in
front, usually smooth, but sometimes ornamented with minute,
close-set, undulating lines. Each valve is marked in the middle
with a short longitudinal furrow, more or less pronounced in
different individuals, and widened at its centre into a navel-like
pit. A well-defined rim extends from one dorsal angle to the
other, round the ventral border of each valve, nearly parallel to
the edge, from which it is separated by a shallow groove. The
edge itself 3 is sculptured with minute transverse pits.
This is the characteristic Primitia of the Aymestry Limestone
of Chances Pitch, near Malvern.
16. Primitia cristata, nov. Pl. XIII. figs. 1a, d, e.
Length ;38,, height ~24,, thickness ;18, inch.
Carapace convex, but most tumid posteriorly ; suborbicular
or rounded-oblong, and about one-third longer than high,
the Paleozoic Bivalved Entomostraca. 421
Dorsal margin straight or slightly depressed; dorsal angles
marked ; extremities broadly rounded, the posterior rather Tess
obtuse than the other; ventral margin boldly convex. Valve-
surface compressed forwards, giving a wedge-like outline to the
carapace seen edgeways; it is also turned inwards at the mar-
gins, thus being flattened at the margins, particularly on the
ventral aspect, where the angle of deflection is marked by a
slight ridge; a delicate rim also accompanies the terminal and
ventral edges. Between the marginal angle above mentioned
and the border itself, there is, at the posterior extremity of each
valve, a sharp crest, directed upwards and inwards to meet its
fellow at the postero-dorsal angle, giving a notched appearance
to the dorsal aspect of the carapace. The surface is smooth,
but bears a deep umbilical pit on the middle, rather towards the
dorsal line.
In the Wenlock Limestone at Crofts Quarry, near West
Malvern.
Symptoms of the marginal angle of deflection, so strong in
P. cristata, are seen in several Primitia, and particularly in P.
umbilicata and P. tersa. These three forms are nearly allied ;
and P. umbilicata is of medium development, and may stand as
the type of the subgroup they represent.
17. Primitia tersa, nov. Pl. XIII. figs. 3 a—-e.
Length ;33,, height 7374, thickness =34, inch.
Carapace tumid, most convex at the hinder third, rounded-
oblong, one-fourth longer than high; dorsal border straight,
angles pointed ; ventral border very convex ; ends nearly equally
rounded. Surface of valves smooth, bearing a pit or umbilical
depression, with irregular outline, in the dorsal region, and
having a slight marginal ridge, which, commencing at the
antero-dorsal angle, runs round the anterior extremity parallel
to the border, and becomes lost, or very faint, at the middle of
the ventral border. In the Wenlock Limestone at Crofts
(Quarry, near West Malvern.
18. Primitia trigonalis, nov. Pl. XIII. figs. 4a, bd.
Length ;42, = 71, height 35, = 74, thickness 33, =, inch
(about) — Pgh 3.6:
Carapace convex, compressed towards the margins, trigonal,
very much like the right valve of Leperditia arctica in shape,
having sloping dorsal angles and protruding ventral border ;
but it has no eye-spot, or radiating muscle-spot, and the valves
are symmetrical and almost equal; indeed the right valve is
slightly smaller than the left, though of similar outline, instead
of the left bemg much the smaller, of different shape, and
422 Prof. T. R. Jones and Dr. H. B. Holl on
strongly overlapped on its ventral border. A slight impression
on the middle of the valves, and the nearly equivalved condition,
characterize this as a Primitia, m spite of its Leperditia-like
outline,—the last being a feature taken on by other Primitia, as
we have already shown. The most convex portion of the surface
is faintly and irregularly corrugated by broadish shallow pits,
which enlarge towards the middle, and there merge into the
feeble sulens: From the Wenlock Limestone of Crofts Quarry,
near West Malvern.
19. Primitia Beyrichiana, nov. Pl. XIII. fig. 9.
3 (as 3 to 2).
Carapace-valve suboblong, straight along the back, boldly
curved behind, gently convex ventrally, obliquely truncate with
a gentle curve in front. Surface convex, and marked with a
broad, subtriangular, faint impression in the middle of the dorsal
region ; and, excepting on the dorsal edge, margined with a neat
flattened rim, rather narrower at the anterior edge than elsewhere,
and uniformly sculptured with minute elongate pits, perpendi-
cular to the peripheral curve of the valve, and thus forming a
radiate ornament. Such a style of margin is present also in P.
Roemeriana, and is still more developed in Beyrichia Maccoyiana ;
and a simply pitted rim is found in B. Salteriana, Isochilina
gracilis, &e.
P. Beyrichianais rare, and is one of the small Bivalve Entomo-
straca referred to (under the terms Cytheres, and Cytheropses) in
Ann. Nat. Hist. ser. 2. vol. xvi. p. 84, and ser. 3. vol. i. p. 249,
as abounding in the drifted Scandinavian blocks of Upper
Silurian Limestone found in North Germany, with some of which
material Prof. E. Beyrich, the eminent palzeontologist at Berlin,
long ago supplied us.
20. Primitia Roemeriana, noy. PI. ee figs. 8 a, b.
Length ;+7,, height ~25,, thickness +23, inch.
Carapace ovate, Back Ime less convex than the ventral border.
Valves convex, bearing a very faint subcentral impression, and
marked with numerous, small, roundish, shallow pits, the inter-
vening surface being a smooth, nearly regular meshwork (like
the pattern on the side of a thimble). The valves are also bor-
dered with a flat marginal rim, broadest ventrally, thinning away
at the dorsal slopes, and neatly ornamented with a row of minute
subquadrate pits, forming a radiate fringe, as in P. Beyrichiana.
The ventral profile of the united valves is acutely ovate.
This ornamented species, differmg markedly from the pretty
radiated P. Beyrichiana of Sweden, is from the Wenlock Limestone
of the Crofts near Malvern; and we name it after our friend
the Paleozoic Bivalved Entomostraca. 423
Professor Dr. Ferdinand Roemer, of Breslau, who has already
enlarged our knowledge of palozoic fossils from the Scandina-
vian Limestones.
21. Primitia obsoleta, nov. Pl. XIII. figs. 12 a, b,c.
Length 513; (+), height =4,, thickness ;$, inch.
5
300 0
Carapace-valves obtusely subovate, straight at the back, con-
vex ventrally ; one end broadly and obliquely curved, with a
dorsal angle, the other semicireular; a slightly raised smooth
rim runs along the ventral border, continues along the broad
end, widens at the dorsal angle, and dies out on the hinge-line.
Surface convex, slightly flattened towards the narrower end,
smooth, and presenting, at the middle of the dorsal region, a
faintly indicated impression, fainter even than the shallow sulcus
in its companion P. Beyrichiana.
In Scandinavian limestone (Upper Silurian), drifted blocks,
North Germany.
22. Primitia oblonga, nov. Pl. XIII. figs. 14a, b,c.
Length +,, height ~,, thickness 35 inch (84: 2:1).
Carapace-valves convex, sloping gently and nearly equally to-
wards the margins, oblong, with rounded, almost symmetrical
ends; the dorsal line slightly sinuous, and the ventral line very
faintly convex. Surface smooth. The slight sinuosity of the
dorsal line is probably a trace of the sulcus, as in P. radiata and
other associated forms. The ventral margin is very slightly
thickened by being suddenly inturned. Excepting this last
feature, the absence of positive characters in this relatively large
form might have made us doubtful whether it be really a Pri-
mitia as defined above, or whether we ought to have classed it
as a Cythere, with some fossil and recent carapaces of which it
might almost equally well be compared.
In the Scandinavian limestone blocks (Upper Silurian) from
Northern Germany.
23. Primitia ovata, nov. Pl. XIII. figs. 13 a, d, c.
Length ;3., height ;2,, thickness ¢4 inch.
679
Carapace-valves nearly ovate, excepting that the dorsal margin
is not so convex as the ventral ; extremities round, but one much
narrower than the other; surface smooth, convex, sloping off
gently towards the ends ; ventral border thickened by a sudden
inturn of its edge, with a low rounded ridge at the angle of
deflection. In this last feature it has an alliance with P..odso-
leta; but its outline, want of postero-dorsal angle, and smaller
convexity distinguish it.
Drifted Scandinavian limestone (Upper Silurian), with P. ob-
soleta, &e.
424 On the Paleozoic Bivalved Entomostraca.
24. Primitia semicireularis, nov. Pl. XIIT. figs. 10 a, d, ¢.
Length 4, height 24, thickness 4; inch (8: 6: 4).
Rea valves moderately convex, See sometimes sub-
orbicular, with a straight back, and with one end more or less
acute at the dorsal angle, and the other well rounded and form-
ing a bold semicircular curve with the ventral line. The ventral
margin is thickened, as in P. ovata. There is no doubt that
P. semicircularis, P. ovata, P. obsoleta, and P. oblonga are all
closely allied forms of the simplest of Primitie ; still, presuming
that their soft parts may have had distinctive characters (as 1s
likely) we keep them apart for convenience. They are grega-
rious, and, together with P. mundula, form a considerable pro-
portion of some parts of the Upper Silurian Limestone of
Sweden.
In the Scandinavian limestone (drifted), North Germany.
P, pusilla seems to represent P. semicircularis in Britain.
25. Primitia pusilla, nov. Pl. XIII. figs. 11 a, 11 6.
Length ;27,, height +21, inch.
This little smooth subovate form mi :ay possibly be a variety of
P. semicircularis ; but its less convexity and more rounded out-
line distinguish it, to say nothing of its faint suleus—a feature,
however, which may possibly be present in some specimens of
P. semicircularis. From the Wenlock Limestone, near West
Malvern.
26. Primitia concinna, Jones.
Cytheropsis concinna, Ann. Nat. Hist. ser. 3. vol. 1. p. 249, pl. 10.
figs. 3, 4.
This neat subeylindrical carapace probably belongs to a furrow-
less Primitia; there is, indeed, a faint dorsal hollow in the
original, which may stand for the suleus. From the Lower
Silurian Limestone on the Ottawa River, Canada.
The same form is not uncommon among the little Entomo-
straca of the Upper Silurian Limestone of Gothland (from
Dr. Lindstrém) ; and we have seen one like it also in the Upper
Silurian Limestone of Arisaig, Nova Scotia (from Dr. Honeyman).
It is not impossible that Eichwald’s Leperditia minuta may prove
to be the same species; for though the published figures* are
not quite similar, yet some specimens with which M. E. d’Eich-
wald has favoured us can scarcely be distinct ; the state of pre-
servation, however, hinders exact comparison.
* Bullet. Soe, Nat. Imp. Moscou, 1854, part 1. p. 99, pl. 2. fig. 6; and
Lethza Rossica, livr. 7 (1860), p. 1335, pl. 52. fig. 2. Schrenk finds L.
minuta in the Lower Sinan of the Baltic Provinces (Untersuch. p- 195).
Dr. J. E. Gray on the Names of the Genus Mystomys. 425
27. Primitia muta, nov.
Cytheropsis concinna?, ibid. p. 254, pl. 9. fig. 3.
Oblong-ovate, nearly Leperditia-shaped ; back straight, ends
rounded but unequal; ventral edge convex; surface smooth,
Though resembling P. concinna at first sight, it is less cylin-
drical, broader at the obliquely rounded end, and its ventral
dge is more symmetrically curved : it 1s near to P. rugulifera in
shape. From the Upper Silurian Limestone of Beechey Island,
with P. rugulifera, P. sigillata, Beyrichia clathrata, B. plagosa,
and Leperditia gibbera.
EXPLANATION OF PLATE XIII.
[The figures are magnified about 20 diameters. |
>
=
. Primitia cristata, J. & H.: a, carapace, view of right valve ;
b, posterior view; ¢, dorsal view.
. P. umbilicata, J. & H.: a, left side of carapace; 5, dorsal view ;
c, ventral view; d, inside of valve (filled with matrix).
Fig. 3. P. tersa, J. & H.: a, carapace, side view; 6, dorsal view; c, ven-
tral view.
es
eS
bo
Fig. 4. P. trigonalis, J. & H.: a, carapace, view of left valve; 6, dorsal
view.
Fig. 5. P. renulina, J. & H.: a, carapace, side view; &, dorsal view.
S
~~
. 6. P. variolata, J. & H.: a, left valve; 6, its edge-view; c, right
valve of var. paucipunctata; d, its edge view.
Fig. 7. P. matutina, J.& H.: a, right valve; 6, its edge view.
8. P. Roemeriana, J. & T.: a, carapace, side view (left valve) ; 6, pro-
file.
‘ig. 9. P. Beyrichiana, J. & H., right valve.
Fig. 10. P. semicircularis, J. & H.: a, left valve; 0, its end view ; ¢, ven-
tral edge.
Fig. 11. P. pusilla, J. & H.: a, right valve; 6, its edge view.
‘tg. 12. P. obsoleta, J. & H.: a, right valve; 6, its end view; c, edge
view (ventral).
Fig. 13. P. ovata, J. & H.: a, right valve; 4, its end view; c, edge view
(ventral).
Fig. 14. P. oblonga, J. & H.: a, right (?) valve; 6, its end view; ec, edge
view (ventral).
XLVIII.— On the Names of the Genus Mystomys. (In a Letter
to Professor Allman.) By Dr. J. E. Gray, F.R.S., V.P.Z.S.,
F.L.S. &e.
British Museum, Noy. 15, 1865.
Dear Proressor ALLMAN,
As I have been informed that, in your paper on the animal
that Du Chaillu noticed as Cynogale velox, you persist in retain-
ing the generic name of Potamogale, I venture to send you the
following observations, in the hope that I may induce you to re-
consider the question, and avoid adding another to the several
useless names which the animal has already received.
426 Dr. J. EH. Gray on the Names of the Genus Mystomys.
I did hope that I had clearly explained why I rejected that
name in my paper in the ‘Annals and Magazine of Natural
History’ for July 1861, vol. vin. p. 62.
M. du Chaillu’s description of the Cynogale velox is so incor-
rect that, if the skin had not fortunately come into the posses-
sion of the British Museum, the animal must have remained,
like the genus of Bats proposed by Bowdich because his speci-
men had a large Acarus affixed mside of the ears, one of the
puzzles of zoologists.
M. Du Chaillu observes :—‘ Cynogale velox. This resembles
the Asiatie Cynogale Bennettii, Gray. I have now nothing but
the skin of the animal, the skull having been destroyed by fire.
The teeth resemble those of the above genus of Gray, as well as
the general appearance; but the size of the animal, the length
and character of the tail, and the habitat indicate a distinet
species.” Then follow the description and some observations
on its habits, which are succeeded by the followmg remarks :—
“Only a single species of Cynogale being described, and that a
native of Asia, I thought the different shape and proportion of
the tail, with its African habitat, were sufficient to make this
the representative of a different genus, for which I proposed the
name of Potamogale, preferring, however, to wait until I can
procure the skull and skeleton. I have placed it with the genus
Cynogale, to which it certainly bears a close resemblance.”
This is all that M.du Chaillu says upon the question. Is
such a general observation sufficient to establish a genus, more
especially when the animal described has not the slightest re-
semblance, either in external form, character of feet and claws,
or in dental character, to the animal with which it is compared ?
I need not say that the teeth have not the slightest resemblance
to those of Cynogale, though he says he had the skull, but it
was destroyed—and that the extremities can scarcely be called
“small, the first joimt enclosed within the skin of the body,” and
the fore claws ave not “very slightly if at all webbed,” nor are
the “ hind claws partially webbed.” = With such a description you
had every excuse for believing that your animal was an “ entirely
new genus,” as you did when you first spoke to me about it, be-
fore you were shown Du Chaillu’s skin of Cynogale velox in the
British Museum.
I can only repeat what I said in the paper before referred to,
“ As M.duChaillu has not characterized his genus Potamogale,
and as he has given such an erroneous description of the feet of
the specimen that no one could recognize it, I do not think that
his name has any reason to be retained,” more especially as in
one place he gives the same reason for considering it a species
of Cynogale which he gives in another for thinking that it may
be a different genus.
Dr. J. E. Gray on the Names of the Genus Mystomys. 427
If the name Potamogale is to be used, it must by every just
naturalist be quoted as Potamogale, Allman, as you have charac-
terized the genus, and Du Chaillu has not done so. This would
be all well, if the genus had not already been characterized, as
far as the materials at command would allow, and in a manner
which can leave no doubt of the identity of the animal, as it
is the only one which has the characters assigned to it.
If the rules of nomenclature are rigidly adhered to, you are
in an untoward dilemma yourself; but I cannot believe that
can have any influence on you on this occasion. The paper in
which you established and characterized the genus Potamogale
was read several months ago; but there was no abstract of the
paper including the character of the genus printed in the ‘ Pro-
ceedings,’ and the account of the genus which is to appear in
the ‘Transactions’ of the Society has not been published yet,
and may not appear probably until the end of the year; so in
fact your character of the genus has not been published even yet.
In the meantime Dr. Barboza du Bocage has read a paper in
which he has described and figured the teeth, and established for
it a genus under the name Bayonia. This paper was published
in the second part of the ‘Proceedings’ for the year 1865,
p- 401, and therefore it must have priority. The synonyma of
the genus will stand thus:
1. Cynogale, part, Du Chaillu, 1860.
2. Mystomys, Gray, Ann. & Mag. Nat. Hist. 1861, p. 63.
3. Bayonia, Dr. Barboza du Bocage, Proc. Zool. Soc. 1865.
4. Potamogale, Allman, Trans. Zool. Soc. med. 1865 or 1866.
From the form of the feet and tail, and also from the simi-
larity of the fur to that of Castor and Fiber, I observed that
“T suspect that it is a Glirine animal, much more nearly allied
to Fiber, Hydromys, and Castor than to any ferine genus.” ‘This
has turned out to be an unfortunate suspicion, the fur being as
like that of the aquatic Insectivore Galemys, which did not at the
instant occur to me, as it is to that of the aquatic Glires.
From this observation I have been accused of referring the
genus to Glires*. Perhaps the name | adopted may have had
something to do with this mistake; I only said “TI suspect”
it might be one. But I used the Greek for Mouse as we use it
in English: thus we call a Bat Fluttermouse, a Marsupial
Opossum Mouse, and, more bearing on the question, several
Insectivores Shrewmouse, Elephant Mouse, Hopping-mouse, and
Musk-rat; and no one that I am aware of has objected to
the names of Hylomys, Temm. (a Mole), Echinomys, Licht.
(the Elephant Shrew), and especially Myogale or Myogalea or
* «M. Gray pour un Rongeur,” Proce. Zool. Soe. 1865, p. 402.
428 Dr. E. von Martens on new Species of Shells.
Galemys for the Musk-rat, a genus very nearly allied, as is now
proved, to Mystomys; and the animal is quite as much allied to
a Mouse or Rat as it is to a Weasel, which the name Potamogale
implies. Both the names, if strictly interpreted, have the objec-
tion that Fabricius so forcibly put against the use of generic
names having a signification, which has so frequently induced
me to use names which it is the fashion of some to call bar-
barous, though they appear to me much less barbarous than
many of the sesquipedalian Greek names which some of these
purists have given to the genera they have described.
I see in M. Bocage’s paper that he quotes “ Potamogale velox,
Du Chaillu, Journ. N. H. Soe. of Boston, 1860, p. 361.” But
no such combination of words is to be found in that work at the
page quoted, or in any other that I can find, not even in the
index.
Therefore your adoption of this name is only adding another
to the superabundant names that have been applied to this
animal.
I can only hope that you will reconsider the question. No
one is more desirous than I am that every one should have his
due claim for priority of description properly considered; but I
cannot but believe that in the interest of science one is called on
to resist the adoption of names given, as Potamogale was, with-
out any character, and with particulars that were only fitted to
mislead the student.
I am, my dear Professor,
Yours sincerely,
Joun Epwarp Gray.
XLIX.—Descriptions of new Species of Shells.
By KE. von Martens, M.D.
1. Paludina purpurea.
P. testa conico-globosa, obtecte perforata, solidula, lineis spiralibus
elevatis subtilibus numerosis sculpta, rufo-fusca, non fasciata ;
spira convexe conoidea ; anfr. 5, convexi, sutura mediocriter pro-
funda divisi; apertura vix obliqua, subcircularis, swperne rotun-
data, non angulata, intus purpurea ; peristoma interruptum,
rectum.
Alt. 25, diameter major 22, minor 17, aperturze alt. 15, lat. 123 mill.
Australia, Murray River. The specimens in the Berlin Zoo-
logical Museum were received from Mr. Krefft.
In young specimens a narrow umbilicus is to be seen, which
is shnt up in the full-grown by the inner lip; the upper two
whorls are worn off in the last. I am not aware of any species
closely resembling it.
Dr. E. von Martens on new Species of Shells. 4:29
2. Cyclophorus cruentus.
C. testa anguste umbilicata, turbinata, oblique striatula, striis spira-
libus rarioribus et costis obtusis supra suturam 3, infra 5 sculpta,
subunicolori, rufo-fusca ; spira turbinata, apice obtusa; anfr. 43,
convexi, supremi distinctius oblique striati, carinis in tertio demum
incipientibus, ultimus non descendens ; apertura parum obliqua,
circularis, intus intensive rubra; peristoma multiplex, externum
leviter expansum, ad umbilicum vix arcuato-productum, internum
rectum, continuum.
Diam. maj. 24, min. 19, alt. 19; apert. incluso peristomate 13, ex-
cluso 10 mill.
Loquilocun, island of Samar, Philippines. Collected by
Mr. F. Jagor.
Allied £6 C. tigrinus, Sow., but of smaller size, the last whorl
more flattened, the imner lip | scarcely dilated, never covering the
umbilicus. On the upper whorls a flammulated pattern is
sometimes found, as in C. tigrinus.
3. Tellina semilevis.
T. testa elongata, tenui, convexa, alba, intus flavescente, verticibus
coccineis ; valva dextra sulcis concentricis confertis scabra, sinis-
tra levis, nitida; margo ventralis antice arcuatus, postice ascen-
dens, dorsalis antice vix declivis, convexiusculus, postice descen-
dens, primum concavus, dein convexus; pars antica longior, pos-
tica rostrata, biangulata, angulo superiore magis obtuso ; costz
umbonales_ valvze aes dae subzequales, rotundatee ; Giessice
distincta ; dens cardinalis (compressus, prominens) unus in valya
sinistra, iden in valva dextra; ligamentum breve, immersum ;
dentes laterales nulli.
Long. 48, alt. 24, crass. 10 mill; vertices in 54, longitudinis.
Querimba Islands, near Mossambique (Peters); Red Sea
(Ehrenberg and Dr. Schweinfurt).
Near 7. Pharaonis, Hanley ; distinguished by its form, which
is like that of 7. perna, Spengl.
4. Tellina depauperata.
T’. testa transversim oblonga, tenui, pellucida, concentrice striata,
nitida, lactea, antice rotundata, postice subrostrata, margine ven-
trali vix ascendente, dorsali postice subrectilineo ; flexura indis-
tincta ; dentes. cardinales parvi, in utraque valva ‘singulo, valvze
sinistree simplex, compressus, valvee dextree latior ; laterales nulli;
sinus palliaris magnus, per totam longitudinem Iinens palliari ad-
natus, musculo antico appropinquans, etiam in facie externa test
conspicuus.
Long. 26, alt. 15, crass. 6 mill.; vertices in 7; longitudinis.
Cavite, Bay of Manila, Philippines.
Near T. iilium, Hanl., and 7. cygnus, Hanl.
430 Dr. E. von Martens on new Species of Shells.
5. Tellina Moluccensis.
7’. testa transversim oblongo-ovata, inzequilatera, inzequivalvi (valva
dextra magis convexa), tenui, concentrice leviter striata, alba, ad
margines epidermide brunnea vestita, antice rotundata, postice
breviore, angustata, extremitate subperpendiculariter truncata ;
flexura distincta, carinula nulla; dentes cardinales in utraque
valva bini, anterior valvee dextree parvus; ligamentum infossum ;
sinus palliaris latus, ascendens, magna parte a linea palliari solutus.
Long. 22, alt. 16, crass. 7 mill.; vertices in 2 longitudinis.
Island of Batjan, Moluccas.
Allied to 7. angulata, Chemn., but smaller and without keel.
6. Tellina pretexta.
7’. testa transverse ovata, compressa, ineequilatera, subineequivalvi,
concentrice tenuiter striata, nitida, rosea, zonis pallidioribus con-
centricis picta, margine albo, antice rotundata, postice breviore,
subrostrata, margine ventrali paulum ascendente, dorsali recti-
lineo, valde declivi; flexura distincta ; dentes cardinales in utraque
valva bini, laterales nulli; area ligamenti excavata, carinula cincta;
sinus palliaris modicus, transversus, superne angulatus, antice
acutus, a musculo antico longe remotus, per 5 longitudinis suze
cum linea palliari concretus.
Long. 31, alt. 20, crass. 8 mill. ; vertices in 3—;% longitudinis.
Yokohama, Japan.
At first sight nearly allied to 7. ¢enuis, Maton; on closer
inspection, more nearly to 7. nitida, Poli.
7. Tellina dissimilis.
T’. testa trigono-oblonga, convexiuscula, solidula, striis concentricis
subtilibus sculpta, opaca, alba, vertice et zonis nonnullis ochraceis,
ad margines epidermide fusco-grisea vestita, antice rotundata,
postice acuminata; margo ventralis medio subrectilineus ; flexura
distincta ; ligamentum superficiale, dimidium fere marginis dorsalis
postici occupans ; carinula in utraque valva a vertice prope mar-
ginem superum usque ad extremitatem posticam excurrens. Dentes
cardinales valvee dextree duo, anterior minor simplex, posterior
validus bifidus ;- valvee sinistree unicus, validus, bifidus, interdum
adjecto altero posteriore minore simplici; dentes laterales nulli.
Sinus palliaris magnus, primum versus verticem ascendens, dein
angulatim deflexus, lineam palliarem petens et cum hae aut angulo
recto conjunctus, aut antea paulum recurrens, 7m valva sinistra
musculo antico approximatus, in deatra inde sat remotus.
Long. 49, alt. 335, crass. 143 mill.; vertices ad 3 longitudinis,
Yeddo Bay, Japan.
Contours like those of 7. daceridens, Hanl.
4
8. Tellina incongrua.
T'. tes ‘| i d eX solidul bsol 1
. testa trigono-rotundata, convexa, solidula, ODSO ete concentrice
_
Dr. E. von Martens on new Species of Shells. 431
striata, opaca, albida, prope vertices subfasciata et facie interna
media ochracea; antice rotundata, postice subrostrata, margine
ventrali postice ascendente ; flexura distincta ; ligamentum super-
ficiale, dimidium marginis dorsalis postici occupans ; dens cardinalis
valvee sinistree unicus crassiusculus, valvee dextree duo, posterior
bifidus; margo cardinalis incrassatus ; sinus palliaris magnus, per
totam longitudinem cum linea palliari concretus, 7n valva sinistra
museulum anticum pie es in dextra ab oe sat remotus.
Long. 33, alt. 27, crass. 15 mill. ; vertices ad 12 longitudinis.
Yokohama, Japan.
Similar to 7. frigida, Hanl.; but the posterior part shorter,
more sloping, and distinctly bent to one side.
9. Tellina iridella.
T. testa transversim oblonga, sat compressa, ineequilatera, tenul, con-
centrice striata, nitida, roseo-albida, iridescente ; antice rotundata,
postice breviore, subrostrata, margine ventrali subrecto, dorsali
postico valde declivi; flexura parum distincta; dentes cardinales
in utraque valva bini, parvi, in valva dextra subzequales, in sinistra
anterior major ; sinus palliaris magnus, musculo antico appropin-
quatus, per totam longitudinem lineze pe adnatus.
Long. 19, alt. 10, crass. Pak mill. ; vertices in +4 longitudinis.
Japan.
10. Gastrana Japonica.
G. testa transversim pyriformi-ovata, ventricosa, valde inzequilatera,
concentrice striata et striis radiantibus elevatis obtusis superfi-
clalibus confertis decussata, albida, unicolori, antice breviore, ro-
tundata, postice producta, subperpendiculariter truncata, carinula
nulla, margine ventrali medio recte utrinque ascendente. Dens
cardinalis valvee sinistree anterior porrectus, triangularis, leviter
bifidus, posterior parvus, postice sensim in marginem cardinalem
abiens ; sinus palliaris ascendens, rotundatus, major parte a linea
palliari solutus, dimidiam longitudinem teste attingens.
Long. 33, alt. 235, crass. 17 mill. ; vertices in 4, longitudinis.
Yokohama, Japan.
11. Psammobia (Psammotea) connectens.
P. testa ovato-oblonga, subzequilatera, postice paulo elatiore, utrinque
rotundata, leviter concentrice striata, violacea, epidermide intense
violaceo-fusca, nitida vestita; margo ventralis rectus; dens cardi-
nalis valvee dextree unicus, compressus, valvze sinistree duo, mi-
nores, anterior compressus, posterior depressus, nymphis accli-
natus ; sinus palliaris primum transversus, dein oblique descendens,
lineze palliari totus fere adnatus.
Long. 28, alt. 155, crass. 85 mill. ; vertices in + longitudinis.
Banka Island, east of Sumatra.
4.32 M. Serres on Glyptodon ornatus.
12. Solen brevissimus.
S. testa parva, recta, brevi, concolori, cornea; margo anticus per-
pendicularis, extus sulco notatus, intus incrassatus ; cardo in utra-
que valva unidentatus, pone sulcum situs.
Long. 37, alt. 9, crass. 84 mill.
Singapore.
The relatively shortest species of Solen I know—distinguished
by the perpendicular anterior edge from Ceylonensis, Leach, =
intermedius, Phil., and from S. brevis, Gray, =abbreviatus, Phil.,
= truncatus, Reeve, Conch. Syst. i. 25. 1, and both from these
and from S. versicolor, Phil., by the position of the hinge be-
hind the anterior marginal furrow.
L.—On Glyptodon ornatus. By M. Serres*,
Recent investigations have made us tolerably well acquainted
with the peculiarities of the organization of the Glyptodons ;
but as regards the number and characteristics of the species
little has been done since Nodot, in 1856, summed up the state
of science on this point. Unfortunately Dr. Burmeister was
unable to consult this work, and consequently his memoiry,
although more recent, and notwithstanding the new facts its
author was enabled to observe at Buenos Ayres, does not con-
tribute much to the history of the different species the remains
of which have hitherto been discovered.
The lateral margins of the carapace which forms the subject
of this note are alone deficient ; but it is easy to see that upon
these margins the osteites of the dermal skeleton, instead of
acquiring a tubercular form, as in other species of this group,
tended, on the contrary, to become diminished, as in the Arma-
dillo.
The carapace has a semiovoid form, emarginate at the two
extremities. Its dimensions are as follows :—
Distance from the centre of the anterior notch to that of the
posterior one 1:14 metre.
Perimeter of the carapace from one of these points to the
other 1°33 m.
Greatest distance from one lateral margin to the other 0°68 m.
Corresponding perimeter 1°23 m.
Aperture of the caudal and cephalic notches, about 0°35 m.
When this earapace is looked at in profile, one is immediately
struck by the complete difference of its aspect from that of the
* Translated from the ‘Comptes Rendus de l’Académie des Sciences,’
Oct. 2 & 23, 1865, by W.S. Dallas, F.L.S.
+ Annals, vol. xiv. p. 81.
M. Serres on Glyptodon ornatus. 433
carapaces figured by Owen and Nodot. The latter have a very
distinct convexity from one extremity to the other. Ours, on
on the contrary, presents a well-marked sort of constriction at
about the second third of the length of the body. Hence a rule
tangent to the line of the back may touch it at two points, and
subtend an are (which, however, is very flat) between the scapular
and iliac regions. Behind this constriction the carapace, through-
out the iliac region, presents the form of the segment of a
sphere.
All the osteites forming it have a common characteristic type,
which, however, is more or less altered in certain regions—
namely, in front, behind, and on the margins. These modifica-
tions, on which I cannot dwell here, are nevertheless useful to
know. They might lead to error in specific determinations made
from too small fragments. This common type evidently corre-
sponds with the fragment of a carapace figured by Owen under
the name of Glyptodon ornatus, and reproduced less successfully
by Nodot.
Here and there we perceive little shapeless plates fillmg the
vacant spaces left by the regular design of their neighbours,
which does not accommodate itself well to the spherical surface
of the carapace. But this aberration is a normal fact,—the
dermal bones of the Edentata belonging, like the squamose
bones of the cranium, to that variety of the primary osseous
organs which originate without preexistent cartilages, and which,
in consequence of their spontaneous mode of origin and develop-
ment, always present great variations of number and configura-
tion.
The inner face of the plates generally presents a remarkable
uniformity. At the points where they were in connexion with
the bones of the muscular skeleton the union was effected by
fibrous tissue, and not, as Burmeister gives us to understand,
by cartilaginous tissue. Indeed in the Armadillo it is likewise
fibrous tissue that unites the extremities of the ischia to the
processes of the posterior buckler to which they apply them-
selves—a very dense fibrous tissue, with fibres of variable but
generally considerable diameter, attaining as much as 0-007 mill.
These fibres are arranged in slightly undulated and imperfectly
limited bundles.
Throughout the anterior region of the carapace the mner face
of the plates is perfectly smooth. They all uniformly present a
nutritive foramen in their centre, analogous to that which is
seen on the inner face of the plates of the Armadillo, into which
an arterial branch of considerable volume always penetrates. It
is evident that in this region the carapace reposed upon a thick
cushion of adipose tissue.
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. 29
A434. M. Serres on Glyptodon ornatus.
Behind, in the vicinity of the posterior emargination, the
inner face of the plates is, on the contrary, irregular, rugose,
and uneven; nor do we find the same regularity in the distri-
bution of the vessels, each plate having often several nutritive
foramina. To all appearance this region gave attachment to
muscles analogous to the elevators of the tail in the Armadillo,
muscles which are seen to attach themselves, on the one hand,
to the inner face of the posterior buckler, and, on the other, to
the spinous apophyses of the first caudal vertebree.
‘he margins by which the plates are articulated together
present the asperities usual on such organs. They were united
by fibrous tissue. Perhaps, at the perfectly adult age to which
our specimen had attained, some of these articulations were ossi-
fied in the dorsal region; but certainly this was not the case
throughout. It may be asserted that the carapace no more
formed a shell homogeneous and solid in all its parts, than the
thoracic and iliac bucklers of the Armadillo are solid and in-
flexible. It must have become flattened or arched a little in
accordance with the efforts of the animal; im other words, it
must have changed its form, certainly within narrow limits, but
still appreciably.
The caudal emargination is formed by a single row of plates
arranged in an are, like the stones of a bridge.
The cephalic emargination is much more complicated in its
structure. It is not simply circular, but shows on the median
line a projection in the form of a hood, shghtly promment, but
massive, with its margins rounded at the expense of the outer
face of the carapace, sharp at the level of the inner face.
Three different rows of plates assist im the formation of this
anterior emargination, and furnish in turn its marginal pieces.
The series which borders the most lateral part of the emargina-
tion extends, gradually diminishing, over one-third of its com-
pass. Here it disappears. The row which was the second on
the sides thus becomes marginal, and disappears in its turn in
the same manner. There remains the third, which thus becomes
marginal in the middle of the notch. In other words, the row
which bounds the emargination in the centre alone constitutes
a complete arch, resting on each side upon two other arches,
which become less and less complete.
All the plates in the vicinity of the anterior emargination
bear on their surface excavations measuring about 4 millimetres
in all directions ; these are rounded, with two or three Haversian
canals opening in their bottom. These cavities lodged the bulbs
of large hairs, directed forwards, which fringed the anterior
margin of the carapace of the animal.
I now come to that which constitutes the great interest of this
M. Serres on Glyptodon ornatus. 435
carapace—namely, the persistence of its natural relations with
the pelvie ring.
Besides the pelvis, a fragment of the dorsal column was found
in its interior, but not in place. This fragment will be the subjcet
of a second note (p. 438), in which I propose to examine the other
bones which appear to me to be referable to Glyptodon ornatus,
and which present remarkable differences from the same organs
in G. clavipes.
Anatomists are not yet agreed as to the number of vertebrze
which must be regarded as constituting the sacrum in the
Glyptodons. With Huxley, I refer to this bone the two verte-
bre with large transverse apophyses, united by their extremities
to each other and to the ischium, which limit the pelvic cavity
behind. Huxley calls them coccygeal vertebre of the sacrum.
There is no doubt that the first should be referred to the mass
of the sacrum; as to the second, it has with the former such
connexions and such a community of relations (especially in a
gigantic species which I shall hereafter have occasion to bring
before the Academy), that it seems difficult to separate it from
the same group. In Glyptodon ornatus these two vertebre ap-
pear to me to be the tenth and eleventh, counting as the first
the vertebra of which the lateral expansion forms the sacral
crest.
In our specimen the iliac crests and bones, as also the cotyloid
eavity, present nothing particularly remarkable.
The rami of the pubis have not been preserved.
Behind the cotyloid cavities originate the ischia, which are
quite different here from the oblique and fan-like ischia of G.
clavipes. These ischia are directed straight backwards, only
diverging a little. With the sacral crests in front and the two
coccygeal vertebra of the sacrum behind, they bound a nearly
quadrangular cavity, over which the narrow region of the sacrum
is thrown like a high-arched bridge.
The ischium has the form of a triangular lamina placed in a
vertical plane. The apex of the triangle is at the cotyloid cavity
in front, the base at the carapace behind. This base, or posterior
margin of the ischium, is free below. Above it is cut oif some-
what obliquely, to adapt it to the convexity of the carapace,
which rests upon it. The upper margin is thick, and regularly
concave ; the lower margin is more irregular. Near the cotyloid
cavity it presents a concavity, a portion of the orifice of the
obturator foramen. Beyond this it was united with the ramus
of the pubis, now destroyed. Further on it forms a lozenge-
shaped synarthrodial surface, bristling with highly developed
osseous rugosities, which give attachment, during life, to a
powerful interarticular ligament. Behind this rugose surface,
29%
~
4.36 M. Serres on Glyptodon ornatus.
and quite at the extremity of the lower margin of the ischium,
there is a small, smooth articular surface, about 1 centimetre in
diameter.
The outer surface of the ischium is convex from above down-
wards, and slightly concave from before backwards. The inner
surface presents the opposite arrangement.
At the lower posterior part, in the immediate vicinity of the
lozenge-shaped surface of the lower margin of the ischium, the
imner face also presents a large synarthrodial surface, of a some-
what pyriform shape, 2 centimetres broad in front, narrower
behind, and about 6 centimetres in length.
The ischium, regarded as I have just described it, measures
25 centimetres in length from the margin of the cotyloid cavity
to the carapace. But it is easy to see that it is really composed
of two different osseous pieces. The bad condition of the parts
has prevented my investigating this curious point of osteogeny
so completely as I could have wished. It is necessary to take
into consideration the difficulties inherent in such observations
on broken and reconstructed bones. Moreover the individual
itself was by no means favourable for investigations of this
nature, age having already produced in it certain synostoses
which were evidently accidental.
Nevertheless I think it very probable that we may regard the
posterior region of the ischium as independent of the region
united to the sacrum, as a sort of epiphysis of which the anchy-
losis is very late, if indeed it ever takes place. I may, moreover,
remind the reader that in the young Armadillo the ischium
likewise presents at its extremity a distinct osseous piece, the
nature of which must be the same.
In our specimen, at about three-fourths of the length of ie
ischium, we see very distinctly a solution of continuity parallel
to the base of this bone, and consequently nearly vertical, which
separates it into two regions. This solution of continuity is an
articulation. This may be very well seen towards the upper
margin of the ischium, where the articular surfaces of the two
pieces become considerably widened in consequence of the thick-
ness of that margin. They are there nearly 2 centimetres in
diameter, and present pretty nearly the aspect of the epiphysary
surfaces of young bones.
This curious synarthrosis is very distinctly visible on the two
sides. Below, it pretty nearly cuts im half the lozenge-shaped
and pyriform articular surfaces of which I have already spoken as
occurring on the inner face and inferior margin of the ischium.
From this arrangement it follows that, in Glyptodon ornatus,
the sacro-iliac region of the pelvis was not united in a perfectly
rigid manner to the carapace by the ischium. It is evident that
M. Serres on Glyptodon ornatus. 437
at this point there existed certain conditions of mobility, which
were doubtless of but little extent, but of which the multiplica-
tion and complication of all these synarthroses accumulated at
the same point furnish a certain indication. The whole of this
organic construction, moreover, is in accordance with what I
have said above as to the partial flexibility of the carapace.
The two coccygeal vertebree of the sacrum present modifica-
tions corresponding with those of the ischium, with which they
are articulated. The first of these vertebre, which is intimately
united to that which precedes it, is placed nearly in a vertical
plane, slightly inclined from above downwards, and from before
backwards. The total height of the vertebra is 8 centimetres ;
its apex is about 3 centimetres distant from the carapace. The
spinous lamina which surmounts its arch presents posteriorly
two diarthrodial facets (articular apophyses), corresponding with
two facets of the following vertebra.
The transverse apophysis, which is lamellar as in G. clavipes,
is placed at first in the same plane as the body of the vertebra,
but undergoes a slight torsion and becomes horizontal. It con-
ceals the transverse apophysis of the following vertebra, and
articulates therewith much more completely than in G. clavipes.
By its squarely truncated extremity it articulates with the pyri-
form synarthrodial surface which occurs on the inner face of the
ischium.
The second coccygeal vertebra of the sacrum is placed in a
much more oblique plane than the preceding one. Hence the
body of this vertebra makes with that of the preceding one a
salient angle about equal to the sacral angle in man. In its
form the body of this vertebra exactly resembles that of the
caudal vertebre. The rhachidian canal becomes suddenly much
less in diameter on reaching its level. The spinous apophysis
is no longer a lamina, but is rounded and obtuse. At its base
in front are articular apophyses corresponding to the two facets
borne by the arch of the preceding vertebra.
It was of interest, with regard to the mechanism of the tail in
Glyptodon, to ascertain exactly the position of this vertebra with
relation to the caudal emargination of the carapace. A straight
line drawn from the centre of this emargination to that of the
cephalic notch passes through the apex of the salient angle which
this vertebra assists to form. The distance from the posterior
face of the body of the vertebra to the centre of the caudal
emargination is 0°125 metre. The distance from the apex of
its spious apophysis to the same point is 0:080 m.
The transverse apophysis passes entirely beneath that of the
preceding vertebra. But, contrary to what occurs in G. clavipes,
it passes this, and articulates by its superior face, which is
438 M. Serres on Glyptodon ornatus.
roughened with rugosities, with the lozenge-shaped synarthro-
dial surface of the lower margin of the ischium. A small diar-
throdial facet also corresponds with that which terminates this
margin.
At the level of its articulation with the ischium, the transverse
apophysis bends suddenly downwards nearly at a right angle.
It becomes vertical, and continues the outer face of the ischium
downwards for some centimetres. I may remark, in conclusion,
that this arrangement of the second coccygeal vertebra of the
sacrum, which is thus suddenly bent downwards in G. ornatus,
occurs also upon all the middle caudal vertebra of the different
species of Glyptodon.
When we take up the study of fossils we are mevitably led
towards the highest questions of zoology. These ancient spoils
of an extinct world bear imprints which the genius of Cuvier
taught us to read—imprints by the aid of which we can approxi-
mate the action of the forces at such remote periods to the
action of the forces which are still exerted upon the develop-
ment of organized beings.
In organized beings there exist constant differences of orga-
nization, upon which is founded their arrangement in classes,
families, genera, and species. From these differences, and from
them alone, results the variety in the animal and vegetable
kingdoms ; but is the cause or principle of this variety, or of
these differences, inherent in the vegetable or animal organism?
or is it in great part exterior to it ?
Let us remark,in the first place, that whatever may be the oppo-
sition of these two methods of looking at the differences of
plants and animals, there is one idea and one word which neither
of them has been able to eliminate—the idea and the word
type. For it is certainly necessary to recognize differences be-
tween organisms, diverse imprints, and to give them a name.
But while some believe that every specific difference corresponds
to an immutable type, others think that there exists only one
type indefinitely modifiable by the action of the media in which
it is developed.
This latter hypothesis, to which Mr. Darwin’s work on the
origin of species (which, at the time of its appearance, produced
so deep a sensation among naturalists) referred, appears to me to be
equally irreconcileable, on the one hand, with what logic compels
us to admit philosophically, and, on the other, with the facts of
experience. In fact, this unique type, not being immutably de-
termined, would not be so essentially, which destroys the very
idea of type, and substitutes in its place that of the indetermi-
nate substance. Notwithstanding the development given to this
M. Serres on Glyptodon ornatus. 439
hypothesis, the question of variation, or of the differences of
animals and plants, has not made a single step; it remains
entire, inasmuch as there always remains to be explained the
first origin of form or of determination, which evidently has a
cause, a principle anterior to what are called media.
As I have said elsewhere, in the numerous differences which
organized beings may be subject to, they never pass the limits
of their class to acquire the forms of a higher class: a Fish
will never ascend to the form of a Reptile ; the latter will never
attain that of a Bird, or a Bird that of a Mammal. Even in
monstrosities, a monster may repeat itself—it may present two
heads, two tails, six or eight extremities ; but it will always re-
main circumscribed within the limits of its class. This surpri-
sing phenomenon is doubtless connected with the general har-
mony of creation. What can be its cause? Of this we are
ignorant. But it results from this, nevertheless, that everything
is not primitively in the materials—that evidently we must
conceive a principle exterior to them, which determines their
employment and presides in their arrangement.
Media, however, exert a powerful influence in the production
and development of organized creatures. They do not create
the types; but, in the physical world, they are an indispensable
condition of their evolution. If they created the types, only a
single type could exist in the same medium, there would be onl
a single determination for the bemgs which would be produced
there. Now, on‘the contrary, the same media present multitudes
of creatures differmg in nature or in form. Moreover, if the
type which specializes each of these creatures were only the
effect, the term of action of the media, it would be in itself only
a simple modality, something purely passive, whilst facts show
evidently that, in each type, there resides an active energy, a
distinctly determinate organizatory power.
Specific differences therefore cannot have their cause in the
media in which the animals are developed, since they affect ani-
mals developed in the same medium, and persist immutably in
these same animals when transported into another medium.
The influence of these, bemg powerless to change their specific
organization, only introduces into it slight and superficial modi-
fications. Consequently, this specific organization, bemg fun-
damentally unalterable, depends on an internal cause which
is itself unalterable ; from, which it follows that, both im ani-
mals and plants, there exist true species, not by a single type,
but by infinitely numerous types, which become involved and
linked together in proportion as the organisms, increasing in
perfection, become more complex.
To prove the development of the diversity of types in the
440 M. Serres on Glyptodon ornatus.
same medium, I shall select the Fauna of the Tourtia, so well
appreciated by M. d’Archiac.
“What strikes us at once,” says that illustrious geologist*,
“in the examination of this Fauna, which is still so imperfectly
known, the elements in our possession having only been col-
lected at three or four points, is the prodigious development
and the almost infinite variety of the type of the Terebratule.
“‘ Of these we have determined and described forty-eight spe-
cies. Of this number thirty-four, or nearly three-fourths, are
new; and there are, besides, more than twenty equally well-
characterized varieties. This genus of itself includes nearly
one-fourth of all the species that we know in the Tourtia; and
when we consider the small thickness of this bed, and the re-
stricted space in which it is deposited, we cannot but be asto-
nished that a single organic type, after probably a very short
lapse of time, should present in the combination of its forms,
or of its dimensions, so manifest a proof of the admirable fe-
cundity of nature, that it might lead us to doubt of the reality
of the species considered in itself.”
Among fossil Vertebrata, the Glyptodons, which flourished at
the period of formation of the Subapennine stage of the Pampas
of Buenos Ayres, in which the remains of these animals are
found exclusively, reproduce the same fact, although in a much
less degree ; and hence the necessity of well-characterizing the
species, however difficult this may be. In fact, if anatomical
investigation applied to creatures which no longer exist presents
im some respects an attractive interest, it also sometimes exposes
us momentarily to certain zoological confusions, which cannot
always be avoided.
The fragments of several extinct and unknown animals, of the
same group and the same size, reach us, collected together with-
out order, in the same layer of soil. If each skeleton were
complete, nothing would be easier than to bring together these
disjointed organs, and to reestablish the same natural relations
which formerly united them into a single living whole. The
difficulty commences when each of them is represented only by
a bone, a different organ. Then the naturalist cannot escape
the danger of describing as belonging to the same species organs
really pertaining to several, except by fallimg into another error,
that of describing a single species under several names. In this
alternation of two causes of error, both difficult to shun, I
found myself in the comparisons which I propose now to bring
before the Academy.
In my first note on the Carapace of Glyptodon ornatus 1
* «Rapport sur les fossiles du Tourtia,’’ Mém. de la Soc. Geol. de France,
2° série, tome 1. p. 291.
M. Serres on Glyptodon ornatus. 44)
mentioned that a fragment of the dorsal column had been found
adherent to its inner surface, but not in its proper place (see
ante, p. 45). This fragment commences at the articulation of
the third dorsal vertebra, and includes twelve vertebrz soldered
together ; except the number of vertebrze thus united, it presents
nothing in its general configuration that has not already been
described by Huxley, and especially by Burmeister in Glyptodon
clavipes.
The number of vertebre is easily counted by the number
of apertures seen in the bottom of the two vertebral channels.
These apertures gave passage to the posterior branches of the
spinal nerves, and also to venous canals. The median crest in
front is cut off very obliquely, thin and trenchant; it was no-
where articulated or even in direct contact with the carapace.
Each lamina of the twelfth vertebra of the bone is produced in
front into the bottom of each vertebral channel above the la-
mina of the eleventh, by an apophysis (at least ?) three centime-
tres in length. On each side, this apophysis is held in a sort of
mortice, formed by two vertical surfaces, depending from the
arch of the eleventh vertebra. There is synostosis; but this
must no doubt be ascribed to age.
The laminz of the twelfth vertebra present analogous mor-
tices behind; these, no doubt, received the apophyses of the
following vertebra in the same manner.
Between this point and the vertebre soldered together to form
the sacrum, there exists a gap, which is the more to be regretted,
because, the hinder part of the skeleton having, in this indivi-
dual, remained in its normal relations to the carapace, the inte-
grity of the vertebral column would have enabled us to appreci-
ate the true relations of the neck and head with the margin of
the cephalic emargination.
In seeking for the bones of the neck, I met with one which,
although not belonging to the same individual, appeared never-
theless to be referable to the same species. The articular sur-
faces seemed constructed to coincide ; the posterior part of the
voluminous apophysis, which surmounts the vertebral gingly-
mus, was hollowed by a narrow furrow exactly in relation to the
anterior oblique and trenchant extremity of the median crest of
Glyptodon ornatus.
But the skeletal constitution of the neck was not the same
here as in Glyptodon clavipes. In place of the trivertebral and
pentavertebral bones, named by Huxley and myself, I found
two bones, each consisting of four vertebree. The articulation
separating them, the mechanism of which I have already had
the opportunity of describing in G. clavipes, instead of existing
between the fifth and sixth cervical vertebre, occurs between the
442 M. Serres on Glyptodon ornatus.
fourth and fifth. The names of trivertebral and pentaverte-
bral bones, evidently cannot serve to designate these two tetra-
vertebral bones ; for the sake of clearness and brevity I prefer to
adopt the denominations mesocervical bone and metacervical bone.
The metacervical bone presents the general lineaments of the
trivertebral bone of G. clavipes. It is only a little longer in
proportion to its breadth. The two costal impressions are more
oblique. Of the four apertures of conjugation, the last three
present the same general arrangement as in G. clavipes, and the
first is immediately in front of the second.
The articulation which unites the metacervical to the meso-
cervical bone is closer than in G. clavipes. Below, in particular,
it leaves no hiatus, even in its extreme movements. The juxta-
posed vertebral bodies, instead of being reduced to the condition
of a trenchant lamina, measure about five millimetres in diameter.
The line which separates them describes a curve with its con-
cavity in front. It is here the fifth cervical pair of nerves that
traverses the articulation.
The mesocervical bone corresponds with the pentavertebral
bone ; it has all its general features, except that it includes only
four vertebree. The ala of the bone, resulting from the coa-
lescence of the transverse apophyses, is comparatively much
slenderer than in G. clavipes. On the outside it terminates in
a narrow, nearly triangular surface. The posterior face of the
bone does not exhibit any special aperture for the passage of
the vertebral artery.
We have here, therefore, an anatomical structure very similar
to that of G. clavipes, but to a certain extent displaced, thrown
back one vertebra. In this arrangement we cannot, however,
see a teratological anomaly: the existence, in the collection of
of the Museum, of two identical specimens, does away with any
supposition of this kind.
Does the bony constitution of the neck just described pertain
to Glyptodon ornatus? Everything leads me to suppose so, but
the future alone can settle the question.
I hoped to get some light from another dorsal column
derived from an imdividual similar in size to that of which we
have the carapace. But this dorsal column was accompanied
by the metacervical bone, of which it had been the continuation,
and this was a true trivertebral bone analogous to that of G.
clavipes. But on examining it more closely some differences
were found between this dorsal column and that of G. ornatus.
Thus the anterior extremity of the spinous crest instead of beng
oblique, thin, and trenchant, is five or six millimetres im thick-
ness, and rises vertically to a great height. The lateral chan-
nels are also narrower and deeper.
Bibliographical Notice. 443
I consequently found myself in the presence of several (at
least two, perhaps three) species of animals of the same size,
nearly allied in their organization, but very distinct in the ver-
tebral arrangement of the neck, and all developed in the same
medium. Anatomy was enriched by a new fact, and at the same
time created a new difficulty in the classification of the fossil
cuirassed Mammalia, which is already rather confused.
BIBLIOGRAPHICAL NOTICE.
British Conchology, Vol. Il. Marine Shells, comprising the re-
maining Conchifera, the Solenoconchia, and Gasteropoda as far
as Littorina. By Joun Gwyn JeErrreys, F.R.S., F.G.S., &e.
London: Van Voorst. 1865.
Eacu succeeding year brings to us another volume of Mr. Jeffreys’s
new work on the British Mollusca, which is coming out with praise-
worthy regularity. At the present rate of progress two more vo-
lumes will be required to complete the subject. We propose to
continue the course which was adopted in noticing the preceding
volumes, and compare the genera and species here described with
those contained in Forbes and Hanley’s ‘ History of the British
Mollusca,’ which, up to the present time, has been the standard
authority on this portion of the marine zoology of our islands.
The following are the generic changes. Sphenia of Turton is
merged in Mya, and Cochlodesmain Thracia. Panopea Norvegica
is regarded by our author as a Sawzicava and not a Panopea; but
the latter genus is not excluded from our lists, since the little shell
figured by Forbes and Hanley (pl. 6. figs. 1-3), but not named
(vide vol. iv. p. 248), appears here with the name of Panopea pli-
cata, Montagu. The genus Patella is divided, and P. pellucida is
placed in the genus Helcion of De Montfort: on the other hand,
Pilidium fulvum falls into Teetura of Cuvier, a name adopted
by Mr. Jeffreys instead of Acmea of Eschscholtz. Capulus, De
Montfort, supersedes Pileopsis, Lamarck ; and, lastly, the genus
Cyclostrema of Marryat is adopted in the family of the Trochidse
for the reception of three little shells,—C. Culterianum, Clark
(=Skenea Culteriana, F. & H.); C. nitens, Philippi (=Trochus
pusillus, F. & H.); and C. serpuloides, Montagu (=Skenea divisa,
Eis E.):
The aie species, which were admitted in the ‘ History of the
British Mollusca,’ are excluded in vol. iii. of ‘British Conchology :’—
Corbula ovata, Teredo palmulata, and Trochus conulus, as not being
indigenous ; Shkenea costulata, as occurring only in a fossil state ;
Pandora obtusa, Thracia villosiuscula, Corbula rosea, Saxicava
arctica, Skenea levis, and Patella athletica, assigned respectively as
varieties to Pandora inequivalvis (P. rostrata, F. & H.); Thracia
papyracea, Poli (T. phaseolina, F. & H.); Corbula gibba (C.
nucleus, F. & H.); Saxicava rugosa ; Cyclostrema nitens ; and
Ad Bibliographical Notice.
Patella vulgata. In the genus Littorina, L. fabalis and L. palliata
are referred to Littorina obtusata (L. littoralis, F. & H.), and Lit-
torina tenebrosa, L. saxatilis, and L. patula are grouped, as Forbes
and Hanley suggested they probably should be, under ZL. rudis.
Several Mollusca of much interest are added to the British fauna
in the present volume.
Neera rostrata, Spengler, a form which ranges from the Augean
and Mediterranean seas to the coasts of Norway and Sweden, is ad-
mitted on account of a single right valve, dredged in seventy-six
fathoms forty miles from the east coast of Shetland.
Teredo pedicellata, Quatrefages, inhabits fir and oak used in sub-
marine and fixed woodwork in the Channel Islands.
Lepeta ceca, Miller. A single specimen has been dredged in the
Shetland seas, and a second off the west coast of Scotland.
Trochus amabilis, Jeffreys. A very beautiful new species, diseo-
vered among ‘fine sand, mixed with gravel, in 85 to 95 fathoms,
about twenty-five miles N.N.W. of Burra Firth Lighthouse, Unst.
The area in which it occurs appears to be limited to a few square
miles. I discovered this new and beautiful species in 1861, while in
company with my friend Mr. Waller; and we obtained specimens
again in 1864 by dredging on the same ground. Living together
with it were Limopsis aurita, Cylichna alba, Buccinopsis Dalei var.
eburnea, and other treasures. I do not know any other place, at
home or abroad, where it has been found.’’
Trochus Duminyi, Requien. Found among shell-sand at Bun-
doran, in Donegal Bay. This is a Mediterranean species, and was
first described as a fossil by Philippi, under the name of Valvata?
striata. It was recorded as British by Mr. Jeffreys, in his ‘‘Addi-
tional Gleanings in British Conchology”’ in the ‘Annals’ of Sep-
tember 1859, as Skenea striata.
Although not prepared to acquiesce in all the changes which have
now been enumerated, yet with the greater part of them we entirely
agree, and shall only here call attention to two of the points in
which we venture to entertain a different opinion from that which
is maintained in the book before us.
We must enter a strong protest against the uniting of Pandora
obtusa with its ally P. inequivalvis. Mr. Jeffreys gives the fol-
lowing reasons for the course which he has adopted in treating
the former as a variety of the latter species :—‘ The difference be-
tween the typical shell and the variety obtusa apparently arises from
the nature of their respective habitats—the one being sublittoral,
and the other belonging to deeper water. An intermediate form has
been taken by Cailliaud on the coast of Brittany, and by M‘Andrew
at Corunna. On a superficial view, indeed, it would seem as if a
valid distinction existed in the length from the beak to the front mar-
gin being always greater in P. inequivalvis (or rostrata), and on
the posterior side in P. obtusa; but this only shows that varieties,
as well as species, have some character of their own. Such may
be expected when the conditions of life vary. The extension of
the posterior side in the typical form may be caused by the differ-
Bibliographical Notice. 4.45,
ence of locality. When the Littoral zone is sandy, the surface is
apt to be disturbed by waves and occasional storms, so that the
stratum may be of a greater or less thickness at one time than at
another: now it is covered by a deposit of material thrown up by
the sea ; in a few days this cover may be stripped off. In order to
prevent its tubes being choked by an accumulation of the imported
material, the Pandora living between tide-marks gradually lengthens
that end of its shell. The variety which inhabits deeper water is
not exposed to fluctuations of this kind; it therefore does not re-
quire any such provision, aud lies undisturbed in its level bed.
This may explain the variation in the proportion of length and
breadth which is exhibited by the two forms. The difference of
thickness in the shells of P. inequivalvis and its varieties also de-
pends on habitation. I am also inclined to think that, with regard
to every species living both in the Littoral and Coralline zones, the
shell is thicker in the former than in the latter. Examples to illus-
trate this proposition occur in Venus gallina and its varieties stria-
tula and laminosa, Mactra solida and its variety elliptica, Trochus
ziziphinus and its small conical variety, Buccinum undatum and its
variety zet/andica, and in many other species. Experiments made
by Dr. Davy, Forchhammer, and Bischoff have proved that the
quantity of carbonate of lime held in solution by sea-water, and
from which shells are produced, is greater on the coast than in the
ocean: it is derived from the land, and brought down to the sea by
rivers and streams, the washings of rain, and the action of waves.
This fact ought not to be lost sight in discriminating species from
varieties, of which the comparative solidity and size are the sole or
chief criteria.”
Now, surely, there is a great deal assumed here. That conve-
nient little word “‘may”’ holds a very important place in this argu-
ment ; and Mr. Jeffreys supposes an instance of wonderfully rapid con-
formity to requirement when he urges that the posterior extremity
of the shell may be lengthened in the course of a few days to pre-
vent the tubes being choked by an accumulation of the shifting ma-
terial of the sea-shore. We would suggest that the animal might
crawl upwards out of the deepening sand somewhat more rapidly
than the shell would be likely to grow, and, moreover, that it must not
be forgotten that the size and form of the shell depend upon the size
and form of the animal contained within it. Again, it is undoubted]
true that a species which has a considerable range in depth, will be
frequently found to have its shell much more strongly developed in
the Littoral and Laminarian zones than it is in greater depths of
water ; but this fact does not prove that a thin-shelled, deep-water
member of a genus is specifically identical with a stronger-shelled
form which invariably inhabits shallower water, and differs from it
not only in respect to the substance of the shell, but in many other
particulars of structure also; moreover we are not aware that the
comparative tenuity of P. obtusa has ever been relied upon as of
weight in maintaining its specific character. Forbes and Hanley
clearly point out the characteristic differences of the two allied
4.46 Bibliographical Notice.
species of the genus; and it appears to us that Mr. Jeffreys yields
everything when he acknowledges ‘it would seem as if a valid dis-
tinction existed in the length from the beak to the front margin
being always greater in P. ¢nequivalvis, and on the posterior side in
P. obtusa ;” for the point of difference here alluded to will be found
to be constant and all-important. In P. obtusa, the greatest dia-
meter from the dorsal to the ventral margin is always situated
towards the posterior extremity of the shell, whereas in P. ineequi-
valvis it ts anterior, or in a line drawn from the beaks. We have
examined more than 600 specimens of P. inequivalvis from Jersey,
for the express purpose of finding intermediate gradations between
that species and P. obtusa, but without meeting with any such
Specimens ; and the more experience we have had of the two shells,
the more firmly have we become convinced that differing, as they
do, 1st, invariably in form, 2udly, invariably in habits, and, 3rdly, in
geographical distribution, Pandora inequivalvis and P. obtusa can-
not but be regarded as valid and distinct species.
By far the most interesting portion of the present volume is the
graphic and full account of the ship-worm (Teredo), to the history
of which the author devotes no less than forty-five pages. The details
respecting this genus have evidently been drawn up with especial
care, and must be read in their entirety to be appreciated. To
attempt to give here partial extracts would be doing the author an
injustice. Mr. Jeffreys describes only six species of the genus,
whereas, in his “ List of the British Species of Teredo,”’ published in
the ‘Annals’ of August 1860, no less than fifteen were admitted.
On examining the reasons for this wide discrepancy, we find that
Teredo nana and T. subericola are now regarded as varieties of
T. megotara, and that T. bipinnata of Turton is the same as 7’
pennatifera of Blainville—the former author having assigned in his
description the valves of one species (7'. malleolus) to the pallets of
another (7. pennatifera). Again, T. bipartita, T. fusticulus, T.
spatha, and T. fimbriata (T. palmulata, F. and H.) undoubtedly
have no claim to be admitted into our fauna. The valves and
pallets of these species have indeed been found in pieces of wood
floating in our seas ; but those woods have always been of extra-
European origin—West-India cedar and mahogany—and the con-
tained Teredines have never been met with alive, the animals
having apparently perished in the cold waters of our seas.
We are, however, at a loss to understand upon what principle 7.
malleolus and T. bipinnata (T. pennatifera, Blainville) are admitted,
while 7’. exeavata and T’. cucullata are excluded. These four species
occur under precisely similar circumstances, and are frequently found
associated together and living in the same pieces of European woods
drifted to our shores. We cannot acquiesce in the statement that
‘the habits of the Teredo are littoral,’ and that “when they are
met with far from land, the piece of wood which contains them has
been accidentally detached and carried out to sea by some marine
current.’ On the contrary, the four species just mentioned, though
frequently met with in floating balks of timber, have never as yet,
we believe, been found in fixed woodwork. They appear, like the
Miscellaneous. 44.7
Lepadide, with which they so constantly live associated, to be
essentially pelagic. The position which they invariably occupy in
drift-wood proves that they have attacked the wood while it has
been in a floating position; for as the wood floats in the water it will
be found that the entrances to the tubes of the shipworms are always
on the upper portion, where they are frequently brought into direct
contact with the atmosphere, while the more deeply immersed angle
of the log is hung with vast masses of barnacles. If the grounds
on which those Teredines frequently found living in floating timber
drifted to our shores by westerly winds are excluded from our fauna
be valid and just, then, all we have to say is that Mr. Jeffreys has
established a precedent which, if followed out in other branches of
science, would lead us to refuse to admit into our fauna all occasional
ornithological visitants, the Tanthine, Salpa, many oceanic Crusta-
cea, Physalia, Velella, Diphyes, &c., and all the Lepadidze except
Scalpellum vulgare. We commend to Mr. Jeffreys’s notice the
following observations of Mr. Darwin upon the genus Lepas, which
presents us with an exact parallelism to Teredo:—‘'The species
abound over the Arctic, temperate and tropical parts of the Atlantic,
Indian, and Pacific Oceans, and are always, or nearly always, attached
to floating objects, dead or alive. The same species have enormous
ranges; in proof of which I may mention that, of the six known
species, five are found nearly all over the world, including the British
coast, and the one not found on our shores (the Z. australis) appa-
rently inhabits the whole circumference of the Southern Ocean.”
In conclusion, we will only add that the more we see of ‘ British
Conchology,’ the more do we recognize the value of the work both
to conchologists and Tertiary paleeontologists, and the more confi-
dently are we able to commend it to our readers. And now, Mr.
Jeffreys, we shall for the present wish you good-bye, looking forward
with especial interest to the appearance of your next volume, and
anxious to learn what you are going to do with those horribly tor-
menting Odostomia, over the study of which we poor conchologists
have so often strained our eyes, and racked our brains, and scratched
our heads in the agonies of perplexed doubt! We will not say,
“Woodman, spare that tree,” but rather, ‘ Don’t be afraid of the
pruning-knife.”
MISCELLANEOUS.
Capture of Muscicapa parva at Scilly.
To Dr. J. E. Gray, F.RS. $e.
Sir,—It may be interesting to you to know that another example
of Muscicapa parva, very nearly in the same state of plumage as its
predecessor at Scilly, was captured on Sunday week, at Trescoe Isle,
Scilly. The variation in its plumage consists in the scapularies and
wing-coverts being more decidedly bordered with rufous. This, I
think, shows it to be a bird of the year. I expect it breeds in
Britain. Yours obediently,
Penzance, Noy. 14, 1865. Epwarp Herarue Ropp.
448 Miscellaneous.
On the Canine Teeth of Thylacoleo carnifex (Ow.).
By Prof. M‘Coy.
To the Editors of the Annals and Magazine of Natural History.
GENTLEMEN,—I beg to enclose you a rough pen-and-ink sketch
of the exact natural size of one of two canine teeth of Thylacoleo
found, with part of the jaw and teeth of Moto-
therium Mitchellii (on which it had probably
been feeding), on the surface of Mr. Bell’s
station, Murchill, not far from Geelong, in
this colony of Victoria. As hitherto only the
molars and back part of the skull of Thyla-
coleo have been known, the discovery of the
great canine is of much interest. The trans-
verse section of the crown is rotundato-oblong,
having two long nearly parallel lateral bounda-
ries, and the anterior and posterior faces ob-
tusely rounded, the anterior a little larger than
the posterior ; the great fusiform bony root is
very coarsely marked with short, irregular, in-
terrupted longitudinal sulci, and narrow ridges
about a line in thickness. The specimen is
nearly 5 inches long and 1 inch 5 lines wide
at about the middle of the fusiform, com-
pressed, gently incurved root. The crown is
worn down obliquely almost to the base, only
about an inch of it remaining.
The specimens have been presented, through
Dr. Greene, to the Melbourne National Mu-
seum; and [ shall shortly figure them in the
Decades I am preparing of the recent zoology and paleontology of
Victoria.
I remain, Gentlemen, yours, &c.,
Melbourne, Aug. 24, 1865. Freperick M‘Coy.
Ageon Alfordi.
To the Editors of the Annals and Magazine of Natural History.
GENTLEMEN,—A second specimen of Ayeon Alfordi was found
here on August 24th by Mr. Mundie, 93 Richmond Road, Dalston.
Amongst other favourite haunts, I showed him a ledge of rocks stretch-
ing into the Roads under the Garrison Hill, and here, with many of
the more common species, he found one Anemone which was new to
him. He examined this treasure carefully with a lens, and found it to
be exactly like Aigeon Alfordi in form, and also in colour on base,
column, and disk ; but the tentacles, instead of being “satiny green
throughout with a faint line of grey on the outer edge,” were of a lus-
trous satiny green on the back, whilst the front was marked as in those
of Bunodes Ballii, with bars and dots of opaque white on a ground of
neutral tint, this tint brightening into a lovely rose-colour at the tip.
I described this specimen to Mr. Gosse, and he judged it to be avariety
of A’geon Alfordi, and advised me to write you an account of it. This
Miscellaneous. 449
specimen seems at least to make another link between the Antheade
and Bunodide. The Anemone still lives in the possession of Mr.
Mundie.
IT have had to change my opinion about the quantities of Aiptasie
around these islands. Their very great number soon made me think
that I had been mistaken. I sent some specimens to Mr. Gosse, and he
imagined them to be Atptasie, but they did not recover the journey
sufficiently for him to be sure aboutit. I have had no accommodation
for them to disport themselves fully, and consequently I was misled
by a general resemblance. I must conclude now, that though there
are probably Aiptasie to be found here, yet they certainly do not
exist in the numbers [ imagined, but I have mistaken the grey
Anthee for them. I hope you will enter this in your ‘Annals’ as
soon as possible, lest Anemone-fanciers should be tempted to our
distant retreat by false hopes.
St. Mary’s Parsonage, Isles of Scilly, Sincerely yours,
November 4, 1865. D. P. Aurorp.
On the Nest of the Ten-spined Stickleback.
By W. H. Ransom, M.D.
Although it has long been known that the three-spined and fif-
teen-spined Sticklebacks build nests—the former on the ground, and
the latter among weeds—no one has yet described, so far as I am
aware*, the nest of the ten-spined Stickleback, or Gasterosteus pun-
gitius, so common in our ditches, and well known by the name of
Tinker, from the black nuptial livery of the male. I was fortunate
enough, in 1854, to have a nest built by a little Tinker in my aqua-
rium, and I found that its position resembled that of the fifteen-
spined species. Last spring I was able more carefully to witness
this interesting fact. On May Ist, 1864, a fine black male G. pun-
gitius was put into a well-established aquarium of moderate size, in
which he soon became at home, but did not build any nest for three
days. I then supplied him with two ripe females of the same spe-
cies. Their presence at once roused him to activity, and he soon
began to build a nest of bits of dirt and dead fibre and of growing con-
fervoid filaments, upon a jutting point of rock among some interla-
cing branches of Myriophyllum spicatum—all the time, however,
frequently interrupting his labours to pay his addresses to the females.
This was done in most vigorous fashion, the male fish swimming, by
a series of rapid little jerks, near and about the female, and even
pushing against her with open mouth, but usually not biting. After
alittle coquetting, if she be ripe, she responds and follows him, swim-
ming just above him as he leads the way to the nest. When there,
the tables are turned, the gentleman now coquets, he seems not to
* Since my paper was read, I have discovered that Mr. Charles Strange
described the nest of the Tinker in ‘Once a Week,’ vol. i. p. 145. There
is also a figure accompanying the communication, but that appears to be
the nest of the three-spimed Stickleback. Mr. Couch, im his ‘ History of
British Fishes,’ now in course of publication, remarks that he has never
seen the nest of the Tinker.
Ann. & Mag. N. Hist. Ser. 3. Vol. xvi. 30
450 Miscellaneous.
know where it is, will not swim to the right spot, and the female,
after a few ineffectual attempts to find the proper passage into it,
turns tail to swim away, viciously chased this time by the male.
When he first courts the female, if she, not being ready, does not
soon respond, he seems quickly to lose temper, and, attacking her
with great apparent fury, drives her to seek shelter in some crevice
or dark corner. The coquetting of the male near the nest, which
seems due to the fact that he really has not quite finished it, at
length terminates by his pushing his head well into the entrance of
the nest, while the female closely follows him, placing herself above
him, and apparently much excited. As he withdraws she passes
into the nest, and pushes quite through it, after a very brief delay,
during which she deposits her ova. While she is in the nest,
the male, in some impatience, pushes his snout against her tail,
and as she passes through he also enters, fertilizes the ova, and then
makes his exit at the opposite side. At once he proceeds seriously
to attack the female and to drive her to a safe distance; then, after
patting down his disturbed nest, away he goes in search of another
female. In this case both the females spawned in his nest, and it
is quite likely that more would have done so had I supplied them.
The nest was built, and the ova deposited, in about twenty-four hours.
My Tinker continued to watch day and night; and during the
light hours he also continued to add to his nest, which at first
was a rather hurried and imperfect structure. After eight days I
put a ripe female three-spined Stickleback into the tank with him.
At once he paid court to her, and solicited her most palpably to
accompany him to the nest; this, however, she obstinately refused,
and he drove her away in real anger. During the night following
she deposited her ova in another part of the tank among the weeds.
This unwillingness of the female seems to be the sole cause of our
finding no hybrids naturally, as artificially it is not difficult to get a
cross between G. leturus and G. pungitius. As the time approached
for the eggs within the nest to hatch, the Tinker was even more con-
stant in his attention to it. About the tenth day, before I could see
any young Sticklebacks, he began, by mistake, to catch some young
Perch, which just then were hatched in the same tank, and to project
them into his nest. In order to be quite sure, I took some young
Perch from another batch ina dish with a pipette, and put them into
the tank near his nest. Isaw him at once seize them, put them into
his nest, and fan them afterwards with great apparent affection ; this
attention was, however, not appreciated; for the fry of the Perch
are very erratic and wandering in their habits, and they swam out
immediately with a skipping sort of motion. About the twelfth
day young Sticklebacks were visible, but they seemed quite still,
resting here and there upon the fibres of which the nest was built ;
none of them, as yet, seemed to have any errant desires, and not
one of them wandered or gave their anxious parent any real
trouble. About this time the Tinker was occasionally seen to visit
and to inspect rather frequently another spot in the tank; and
on the following day (the 14th) he began there to build another
uest, this time free from any rockwork, and just in the middle of the
Miscellaneous. 451
tank, where a few branches of Myriophyllum, Chara, and Anacharis
crossed each other; these he connected by an accumulation of inter-
laced, decayed, and growing fibres of confervee, which he broke off er
collected from neighbouring parts. When the fibres were added to
the nest, he generally pushed thein into the nest with his snout, but
did not, so far as [ could see, agglutinate them with a viscid secre-
tion, such as I have seen the three-spined Stickleback do. When he
brought bits of dirt, he was content to project them into or upon the
mass of fibres which formed the nest. By dint of great industry, in
about forty-eight hours he built a large well-formed nest, about the
size and shape of a full-sized walnut, with a central aperture passing
almost but at first not quite through it, in a direction nearly hori-
zontal. During all this time he never interrupted his labours for,
and care of, the young fry in his first nest, but continued both to
catch the young Perch, and to watch and fan the nest; and after a
few days he had to catch and bring back his own young ones when
they began to wander. About this time his labours were incessant
during the light hours ; and although he seemed to take no food, he
looked well. I then (the fifteenth day) put two ripe female ten-
spined Sticklebacks into the tank; he courted them with great vigour;
but as they did not respond, he drove them into the most distant and
obscure corners with great ferocity: he then set to work with greater
vigour than ever to enlarge and beautify his nest, and in the course
of the following day the females deposited their ova in it so that
they were easily seen, from the favourable position of the nest with
regard to light. For three days more he continued his care of both
nests, and caught indiscriminately young Perch and young Stickle-
backs and put them indifferently into eithernest. By the eighteenth
day from the deposition of the ova in the first nest, and the sixth
after hatching, the Stickleback fry were independent, and roamed in
search of food; and from this date the parent took no more notice of
them. I neglected to note whether the ova deposited in the second
nest hatched safely; but I have reason to think that they did, as
about a fortnight later I saw some_very young Sticklebacks in the
tank. This instance of the failure of instinct is remarkable, as the
Tinker was almost starving in the midst of plenty. I am not aware
that anyone has hitherto noticed the simultaneous building and tend-
ing of two nests by any of the Gasterostez; it would be interesting
to ascertain if this be an exceptional fact, or limited to the ten-spined
species. One would hardly expect it among the three-spined Stickle-
backs, as they can and do enlarge their nests very considerably, so as
in the nest of one male to receive, fertilize, and protect the spawn of
several females. Itis well known that the female Sticklebacks spawn
more than once in a season; but it is not known, so far as I have
read, how many times they spawn, nor how many times the males
build; there will probably be a relation between the two. The ob-
servation here given shows at least two consecutive generations.
This repeated ripening of ova in one season is doubtless connected
with their large comparative size, and their consequent small number,
as compared with those of other osseous fishes.--Transactions of
the Midland Scientific Association.
4
2
INDEX to VOL. XVI.
Acacta mAcntrica, on the gland
of the phyllodium of, 12.
Acalephie, on the histology of the,142.
Aigeon, description of the new genus,
41.
/Egeon Alfordi, note on, 448.
Agassiz, Prof., on the metamorphoses
“of certain Fishes, 69.
Alford, Rev. D. P., on Ageon Al-
fordi, 448.
Alphidia, characters of the genus,
258.
Ammobroma Sonor, note on
220:
Ammonactis, characters of the
genus, 195.
Ammonites, descriptions of new, 225,
334.
Amphibians, notes on some, 120.
Amphiprora, new species of, 5.
Amphisbeenians, on the genera and
species of, 365.
Anabathra pulcherrima, description
of, 128.
Antedon rosaceus, on the structure,
physiology, and development of,
200.
Anthocharis, new species of, 397.
Antillia, new species of, 185.
Antipha, description of the new ge-
Mus, Lolli.
Argyopes, new species of, 98, 346.
Arripis, on the Australian species of,
187.
Artamus, new species of, 60.
Ascidians, on a new type in the group
of, 143.
Astacus, new species of, 296.
Asterionella, new species of, 4.
Austen, N. L., on the habits of the
Water-shrew, 302.
Australia, on the cretaceous deposits
of, 333.
Axolotl of the Mexicans, on the, 122.
Aye-Aye, on the food of the, 142.
Baikia, description of the new genus,
Soe
Baird, Dr. W., on a new species of
Entozoon, 52.
Baleenoptera, new species of, 54, 148.
Balanophylhia, new species of, 195.
Baly, J.S., on new species of Crio-
the,
new
ceride, 153; on new genera and
species of Gallerucidee, 247, 402.
Bartlett, A. D., on the food of the
Aye-Aye, 142.
Bastian, H. C., on the anatomy and
physiology of the Nematoids, 197.
Bat, on the mode in which the Long-
eared, captures its prey, 301.
Bate, C.S., on a new species of As-
tacus, 296.
Bates, H. W., on the Longicorn Co-
leoptera of the Amazons V: alley,
101, 167, 308.
Bence Dr., on the habits of. the
Lyre-bird, 53,
Binney, E. W., on some fossil plants
from the lower coal-seams of Lan-
cashire, 128.
Blackwall, J., on new Spiders from
the Cape de Verde Islands, 80;
from Central Africa, 336.
Boehm, J., on the parasitic nature of
the Mistletoe, 222.
Books, new :—Spratt’s Travels and
Researches in Crete, 279; Gray’s
Handbook of British Water-weeds,
280; Natural-History Transactions
of Northumberland and Durham,
282; Newman’s British Ferns,
359 ; Salter and Woodward’s Chart
of Fossil Crustacea, 562; Bentham
and Hooker’s Genera Plantarum,
Pars 2, 364; Jeffreys’s British
Conchology, vol. 11., 443.
Bothridium, new species of, 52.
Brachiolaria, on a new form of, 70.
Brady, G. 8., on undescribed fossil
Entomostraca from the brick-earth
of the Nar, 189.
Brady, H. B., on the nomenclature
of the Foraminifera, 15.
Bronia, characters of the new genus,
Siile
Burmeister, Dr. H., on a new species
of Whale, 54; on a new species of
Porpoise, 135.
Butler, A. G., on a new species of
Cetonia, 161 ; on four new species
of Butterflies, 39].
Butterflies, new species of, 397.
Carpenter, Dr. W. B., on the strue-
ture, physiology, and development
IND EX.
of Antedon rosaceus, 200; on the
microscopic structure of the shell
of Rhynchonella Geinitziana, 305.
Carter, H. J., on Peridinium eypri-
pedium, 399.
Caspary, R., on the protective sheath
and on the formation of the stem
of the root, 382.
Catodon, new species of, 286.
Cephalopus, new species of, 63.
Cerianthus, new species of, 196.
Cerochroa, characters of the genus,
258.
Cetonia, new species of, 161.
Chevreulius callensis, description of,
143.
Cinclus aquaticus, on the anatomy
and habits of, 49.
Clark, Rev. H., on the Dejeanian
genus Ceelomera and its affinities,
Do, Silt.
Clark, Prof. H. J., on the animal na-
ture of the cilio-flagellate Infusoria,
270.
Claus, M., on some singular organs
appended to the feet of certain
Crustacea, 144; on the organiza-
tion of the Cypridine, 380.
Clitena, characters of the genus, 259.
Ceelomera, on the Dejeanian genus,
256, 315; new species of, 319.
Colobothea, new species of, 101.
Conosmilia, characters of the new ge-
nus, 184,
Coraia, new species of, 323.
Corals, descriptions of fossil, 182.
Corbula suleata, on the occurreace of,
in a fossil state, 113.
Corvina, new species of, 48.
Creswellia, new species of, 4.
Crioceris, new species of, 153.
Crisp, Dr. E., on the anatomy and
habits of the Water-ousel, 49.
Crocodile, on the muscular anatomy
of the leg of the, 326.
Crossopus fodiens, on the habits of,
302.
Crucifere, on the constitution of the
fruit in the, 299.
Crustacea, on some singular organs
appended to the feet of certain,
144; on new or rare, of the French
coasts, 162.
Ctenus, new species of, 336,
Cuckoos of New South Wales, notes
on the, 290.
453
Cyclophorus, new species of, 429.
Cynorta, description of the new ge-
nus, 249.
Cypridinee, on the organization of the,
380.
Cythere, new species of, 189.
Dactylethra, on the, 120.
Dallas, W.S., on the feathers of Di-
nornis robustus, 66.
Dalophia, characters of the genus,377.
Danais, new species of, 397.
Dandelion, on the intercellular mat-
ter and the vessels of the latex in
the root of the, 224.
Diadema, new species of, 398.
Diamphidia, new species of, 402.
Diatoms, descriptions of new genera
and species of, 1.
Dinornis robustus, on the feathers of,
66.
Dircema, new species of, 262, 405.
Dodo, on some bones of the largest
known species of, 61.
Drassus, new species of, 86.
Duncan, Dr. P. M., on some fossil
Corals from the South Australian
Tertiaries, 182.
Eels, on the cultivation of, 384.
Entomostraca, on undescribed fossil,
189; on the paleeozoic bivalved,414.
Entozoon, on a new species of, 52.
Epeira, new species of, 94, 342.
Ephebe, new British species of, 10.
Erethizon, new species of, 203.
Eudesmus, new species of, 180.
Eupsammia, new species of, 195.
Eurysoma, new species of, 348.
Fishes, on the metamorphoses of
certain, 69; on the structure of the
encephalon of, 70.
Foraminifera, on the nomenclature
of the, 15.
Fournier, E., on the constitution of
the fruit in the Cruciferae, 299.
Fungus, on a, developed in ivory and
bone, 223.
Gallerucidze, new genera and species
of, 247, 402.
Gasterosteus pungitius, on the nest
of, 449.
Gastrana, new species of, 431.
Glyptodon ornatus, observations on,
432.
Gonionema, new British species of, 8.
Gosse, P.H., on a new British Sea-
Anemone, 41,
454
Gould, J., on two new Australian
birds, 60; on the egg of Parra
gallinacea, 70.
Gray, Dr. J. E., on a new species of
Cephalopus, 63; on a new British
species of Porpoise, 133; on a new
Finner Whale, 148 ; on a new spe-
cies of American Porcupine, 203 ;
on a new species of Tupaia, 203 ;
on a new genus of Trionychide,
204 ; on two new species of Whales,
210; on a new species of Catodon,
286; on the genera and species of
Amphisbeenians, 365 ; on the names
of the genus Mystomys, 425.
Greville, Dr. R. K., on new genera
and species of Diatoms, 1.
Giinther, Dr. A., on a new species of
Corvina, 48 ; on the British Salmo-
noids, 59; on the Pipe-fishes of
Australia, 141.
Gulliver, Prof. G., on the pollen-
grains of Ranunculus arvensis, 65;
on the occurrence of raphides and
other crystals in plants, 115, 331.
Halocampa, new species of, 196,
Haughton, Rev. S., on the muscular
anatomy of the leg of the Croco-
dile, 326.
Hemiaulus, new species of, 5.
Hersilia, new species of, 81.
Hesse, M., on new or rare Crustacea
of the French coasts, 162.
Hesycha, new species of, 172.
Heterochroa, new species of, 398.
Heterocyathus, new species of, 194.
Hogg, J., on some Amphibians, 120.
Holl, Dr. H. B., on the paleeozoie bi-
valved Entomostraca, 414.
Hollard, M., on the structure of the
encephalon of Fishes, 70.
Hymenesia, description of the new
genus, 209.
Hyperoodon, new species of, 212.
Hypheenia, characters of the new ge-
nus, 410.
Hypselomus, new species of, 111,167.
Hystrix, new species of, 206.
Infusoria, on the animal nature of
the cilio-flagellate, 270.
Insects, on the graduation from “ in-
dividual’ peculiarities to species in,
383.
Jamesia, on some species of, 170.
Jerdon, T. C., on Pristolepis mar-
ginatus, 298.
INDEX.
Jeude’s, De, collection of Mollusca,
note on, 224.
Johnson, J.G., on a new genus of
Trichiuroid tishes, 283.
Jones, Prof. T. K., on the nomencla-
ture of the Foraminifera, 15; on
the palsozoic bivalved Entomo-
straca, 414.
King, Prof. W., on the histology of
Rhynchopora Geinitziana, 124.
Kolliker, Prof., on the histology of
the Acephala, 142.
Kreift, G., on a ey species of Rock-
Kangaroo, 72
Krohn, Dr. ree on the male genera-
tive organs of Phalangium, 149,
Lacaze-Duthiers, M., on a new type
in the group of Ascidians, 143.
Lampyris splendidula, on the strue-
ture of the luminous organs in the
male of, 224
Latrodectus, new species of, 341.
Layard, KE. L., on the Whales of the
Cape, 210; on a new Zebra, 215.
Leighton, Rev. W. A., on British Li-
chens, 8; on the gland of the phyl-
lodium of Acacia magnifica, 12.
Lema, new species of, 155.
Lestiboudois, T., on the existence of
liquid and solid matters in the tra-
chean vessels of plants, 378.
Lichens, notes on British, 8.
Limopsis Belcheri, on the occurrence
of, in a fossil state, 113.
Lizard, on a new, from the Lower
Chalk, 145.
Longicorns of the Amazons Valley,
on the, 101, 167, 308.
Lubbock, Sir J., on the male genera-
tive organs of Phalangium, 301.
Lycosa, new species of, 80.
Macalister, A., on the presence of cer-
tain secreting organs in Nematoi-
dea, 45.
M‘Coy, Prof. F., on the occurrence of
Limopsis Belcheri and other recent
shells in a fossil state near Mel-
bourne, 118; on the Australian
species of Arripis, 187; on the
cretaceous deposits of Australia,
333; on the canine teeth of Thyla-
coleo carnifex, 448.
Malurus, new species of, 60,
Martens, Dr. E. von, ou the Australian
species of Paludina, 255; on new
species of shells, 428.
UN'D EX,
Metridium, new species of, 195.
Microglossa Alecto, on the osteology
of, 134.
Microscope, on a new growing slide
for the, 334.
Mimastra, description of the new ge-
nus, 253.
Mistletoe, on the parasitic nature of
the, 222.
Moreh, Dr. O.A.L., on the homology
of the buccal parts of the Mollusca,
73; on the operculum and _ its
mantle, 117; on the systematic
value of the organs employed as
fundamental characters in the clas-
sification of Mollusca, 385; onthe
limits of the subkingdom Mollusca,
411.
Mollusca, on the homology of the
buceal parts of the, 73; on the
systematic value of the organs em-
ployed as fundamental characters
in the classification of, 385 ; on the
limits of the subkingdom, 411.
Momeea, description of the new ge-
nus, 252.
Monocesta, new species of, 262, 315.
Muscicapa parva, note on the capture
of, at Sciliy, 447.
Mystomys, on the names of the ge-
nus, 425.
Nadrana, description of the new ge-
nus, 250.
Nealotus, description of the new ge-
nus, 283.
Nematoidea, on the presence of cer-
tain secreting organs in, 45; on the
anatomy and physiology ofthe, 197.
Nephila, new species of, 96, 343.
Nephthya, new species of, 196.
Nestinus, new species of, 325,
Newton, A., on some bones of the
Didus nazarenus, 61.
Notopterophorus, on the early stages
of, 162.
Nyctinomus, new species of, 295.
Oncideres, new species of, 175.
Operculum and its mantle, on the,
Orcynus alalonga, on the occurrence
of, on the coast of Devon, 268.
Orithyia, new species of, 89.
Orthoxia, characters of the genus,
260.
Owen, Prof., on the skull of Thyla-
coleo carnifex, 130.
455
Pachypeza, new species of, 314.
Pachytoma, characters of the genus,
261.
Palmeria, description of the new ge-
nus, |
Paludina, new species of, 255, 428.
Parisis, new species of, 196.
Parker, W. K., on the nomenclature
of the Foraminifera, 15; on the
osteology of Microglossa Alecto,
34,
Parra gallinacea, on the egg of, 70.
Pasithea, new species of, 338.
Peridiniun cypripedium, observations
on, 270, 399,
Peritrox, description of the new ge-
nus, 313.
Peters, Dr.W., on the systematic po-
sition of Platacanthomys lasiurus,
PAIS
Petromyzon, new species of, 221,
Phalangium, on the male generative
organs of, 149, 301.
Philippi, Dr. R. A., on the Chilian
* Anguilla,” 221.
Phoezena, new species of, 132, 138.
Phyllopteryx, on the Australian spe-
cies of, 141.
Plants, on the occurrence of raphides
and other crystals in, 115, 331;
description of some fossil, 128;
on the existence of liquid and solid
matters in the trachean vessels of,
378.
Platacanthomys lasiurus, on the sys-
tematic position of, 213.
Plecotus auritus, note on, 301.
Plesiosaurs, on two new, from the
Lias, 352.
Pleurocrypta, characters of the new
genus, 164.
Podontia, new species of, 403.
Polyps, on the classification of, 191.
Primitia, description of the new ge-
nus, 415.
Pristolepis marginatus, note on, 298.
Procalus, new species of, 261.
Psammobia, new species of, 431.
Pterogale, new species of, 72.
Pyenacantha, description of the new
genus, 350.
Pyesia, characters of the genus, 260.
Ramsay, E. P., on the Cuckoos of
New South Wales, 290.
Ransom, Dr. W. H., on the nest of
the ten-spined Stickleback, 449.
456
Ranunculus arvensis, on the pollen-
grains of, 65.
Raphides, on the occurrence of, in
plants, 115, 331.
Rhynchopora Geinitziana, on the his-
tology of, 124, 305.
Rodd, E. H., on the capture of Musci-
capa parva at Scilly, 447.
Royal Society, proceedings of the,
128,.197.
Salticus, new species of, 82.
Sars, M., on a new form of Brachio-
laria, 70; on some singular organs
appended to the feet of certain
Crustacea, 144.
Sastra, description of the new genus,
253.
Saurospondylus dissimilis, descrip-
tion of, 145.
Schultze, M., on the structure of the
luminous organs in the male of
Lampyris splendidula, 224.
Sclater, Dr. P. L., on a new species
of Indian Porcupine, 206 ; on some
animals from Madagascar, 295.
Seott, Dr. W. R., on the occurrence
of Oreynus alalonga on the coast
of Devon, 268.
Seytodes, new species of, 100.
Sea-anemone, on a new British, 41.
Seeley, H., on a new Lizard from the
Lower Chalk, 145; on Ammonites
from the Cambridge Greensand,
925 on two new Plesiosaurs from
the Lias, 352.
Selenops, new species of, 340.
Serres, M.,on Glyptodon ornatus, 432.
Sigillaria, on some species of, 129.
Simeethea, description of the new
genus, DATE
Smith, H.L., on a new growing slide
for the microscope, 334.
Soubeiran, L., on the cultivation of
Eels, 384.
Sowerby, W., on the mode in which
the long-eared Bat captures its
prey, 301.
INDEX.
Sphenoraia, new species of, 262.
Sphenotrochus, new species of, 183.
Spiders, new, from the Cape de Verde
Islands, 80; from Central Africa,
336.
Spilonema, new British species of, 9.
Stephanoceris, new species of, 194.
Stickleback, on the nest of the ten-
spined, 449.
Surirella, new species of, 4.
Tzeniotes, on the species of, 109.
Teeth, on a fungus developed in, 223.
Tellina, new species of, 429.
Tetragnatha, new species of, 99.
Tetrathyra, description of the new
genus, 204.
Theridion, new species of, 91.
Thomisus, new species of, 85.
Thylacoleo carnifex, on the skull of,
130; on the canine teeth of, 448.
Trachysomus, new species of, 174.
Trestonia, new species of, 311.
Trichiuridz, on some allied genera of,
283.
Tridacna elongata, on the anatomy
of, 581.
Tupaia, new species of, 203.
Vaillant, L., on the anatomy of Tri-
dacna elongata, 381.
Veretillum, new species of, 197.
Verrill, A. E., on the classification of
Polyps, 191.
Vogel, Dr. A., on the intercellular
matter and the vessels of the latex
in the root of the Dandelion, 224.
Walsh, Dr. B. D., on phytographic
varieties and species, 383.
Wedl, Prof., on a fungus which is
developed in ivory and bone, 223.
Whale, new species of, 54, 148.
Xuthea, description of the new genus,
248.
Xylomimus, description of the new
genus, 308.
Zebra, on a new species of, 215.
Zoological Society, proceedings of
the, 49, 132, 203, 283, 365.
END OF THE SIXTEENTH VOLUME,
PRINTED BY TAYLOR AND FRANCIS,
RED LION COURT, FLEET STREET.
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