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THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY,
INCLUBING
ZOOLOGY, BOTANY, ann GEOLO
( BEING A CONTINUATION OF TIE ‘ANNALS’ COMBINED WITIL LOUDON AND
CHARLESWORTH’S ‘ MAGAZINE OF NATURAL HISTORY.’)
CONDUCTED BY
ALBERT C. L. G. GUNTHER, M.A., M.D., Ph.D., F.RB.S.,
WILLIAM CARRUTHERS, F.B.S., F.L.S., F.G.S.,
AND
WILLIAM FRANCIS, sun., F.US.
.
eee
LONDON:
PRINTED AND PUBLISHED BY TAYLOR AND FRANCIS.
SOLD BY SIMPKIN, MARSHALL, HAMILTON, KENT, AND CoO., LD.;
WHITTAKER AND CO.: BAILLINRE, PARIS:
MACLACHLAN AND STEWART, EDINBURGH ?
HODGES, FIGGIS, AND CO., DUBLIN: AND ASHER, BERLIN,
1900.
“Omnes res create sunt divinse sapientiz et potentix testes, divitiz felicitatis
human :—ex harum usu onitas Creatoris; ex pulchritudine sapientia Domini ;
ex ceconomia in conseryatione, proportione, renoyatione, potentia majestatis
elucet. Earum itaque indagatio ab hominibus sibi relictis semper estimata ;
a veré erudigs et sapientibus semper exculta; malé doctis et barbaris semper
inimica fuit — -Linnaus.
“Quel que soit le principe de la vie animale, il ne faut qu’ouvrir les yeux pour
voir qu’elle est le chef-d’ceuvre de la Toute-puissance, et le but auquel se rappor-
tent toutes ses opérations.”—Bruckner, Théorie du Systéme Animal, Leyden,
1767.
eee ee ee eo es. Lhesylvan powers
Obey our summons; from their deepest dells
The Dryads come, and throw their garlands wild
And odorous branches at our feet; the Nymphs
That press with nimble step the mountain-thyme
And purple heath-flower come not empty-handed,
But scatter round ten thousand forms minute
Of velvet moss or lichen, torn from rock
Or rifted oak or cavern deep: the Naiads too
Quit their loved native stream, from whose smooth face
They crop the lily, and each sedge and rush
That drinks the rippling tide: the frozen poles,
Where peril waits the bold adventurer’s tread,
The burning sands of Borneo and Cayenne,
AUl, all to us unlock their secret stores
And pay their cheerful tribute.
J. Taytor, Norwich, 1818,
AUERE J rpAMMAM,
» ~ SS
|
Sy 5 2
CONTENTS OF VOL, VI.
[SEVENTH SERIES. ]
NUMBER XXXI.
Page
I. On some Fish-remains from the Parana Formation, Argentine
Republic. By A. Smirn Woopwarp, LL.D. F.L.S. (Plate L)..
II. The New Mexico Bees of the Genus Megachile and a new
Andrena. By T.D, A. CockExE.L, Professor of Entomology, New
Blexico-Aenicultural College... Meas tied oe on su cunccscvesaeusde.
II. On the West-Indian Species of Madrepora. By J. W.
GreGory, D.Sc., F.G.S., Professor of Geology in the University of
PRL OEIO oor cece) rac ee vs Soey tice le stele Riasaid, ni ialol eo sina eha oy eaeM aes
IV. British Amphipoda.—lV. Families Stegocephalide and Cidi-
ceride (part.). By Canon Norman, M.A., D.C.L., LL.D., F.R.S.,
eee abe N DEL 2 ters) daar a sean betatehtec i were ee hw ad Sole Sek ee oe
V. Description of a new Sea-Horse (Hippocampus) from Muscat.
Pyphraits MOTION LE Seth amelie aes ak'scs 0 Ste AIR eke wae ©
VI. A List of the Fishes collected by Mr. Rupert Vallentin in
the Falkland Islands. By G. A. BouLenesr, F.R.S. With Notes
ema tiess © le hOT rn 2 aes cis cave Way tarehe ac ol beepic = ico RTH Ns choke abe nee
VII. Rhynchotal Notes—V. Heteroptera: Asopine and Tes-
Hae tant OILUGOLC RTE Tory NY Ae Arie PIER RONG! © Pate c veg sclape Sis oah aro ue en, « chee ak
VIII. Descriptions of some new Species of Heterocera from Tro-
pical South America. By Hrrsertr Drucr, F.LS. &c........6..
IX. Report on the Neuroptera Odonata collected by Mr. E. E.
Austen at Sierra Leone during August and September 1899. By
Week) Krnpy,, W035.) FbaS. | (Plate TE.)) v5. ad alee Cas vale «
X. Notes on the Forficularia.—V. Descriptions of new Species
and anew Genus. By Matcotm Burr, F.Z.5., F.E.S. (Plate IV.
Ue hy gl Oey as win coche ah oso Be ae Marea eta iel seein int bic phe oie ad oases 8 >
XI. Notes on the Forficularia.—VI. On a Collection of Forficu-
laria from Sarawak. By Matcoitm Burr, F.ZS8., F.E.S. (Plate
Ae eI) Oc nn Wig ater onc v are Nats a(n sy o@iweNe. 4 ass
XII. The Lower Paleozoic Crinoids of Bohemia. By F. A.
EW AMMAR NN oR RE. kay 'g cots aN we gc TIO Mee AO Ee kto De
XIII. Asiatic Tortricide. By the Rt. Hon. Lorp WatsincHaM,
VPM Oro Re PE adn: 7a)'5.9 Sais al naavs 9 wieke Sy is eis nry se eg Pier
XIV. On Giant Squirrels from the Amazonian Region. By OLp-
MEE A LOMAS OS crea ats aks cielo 9p h.a,84shh tals doe UR etd sv Hlele wala aie Se a
ik
20
32
iv CONTENTS.
Page
XV. Plankton Studies.—II. On Pleodorina ilinoisensis, 1 new
Species from the Plankton of the Illinois River. By C. A. Kororp,
Ph.D. (Plates V8 VIL) os coe. as see apes ieee eee 139
New Book :—Catalogue of the Lepidoptera Phaleenze in the British
Museum. Volume II. Catalogue of the Arctiade (Noline,
Lithosiane) in the Collection of the British Museum. By Sir
Grong FE: ELAMPSON, JBarty ance doicticrecen ears cc oe suseres oer 157
On the ‘Ankiindung eines systematischen Werkes yon dem Schmet-
terlingen der Wienergegend’ of Schiffermiiller and Denis, by
Louis B. Prout, F.E.S.; Abundance of the Greater Siil-Smelt
(Argentina silus, Ascan) on the Market of Boston, Lincoln-
shire, by2G. A. poulenger HOR Sn aac s jes oicn oe eae ene 158, 160
NUMBER XXXII.
XVI. Note on Diptera from South Africa (Tabanide and Asilide).
By Miss GERTRUDE RicaRpo
nha dhe want Goede eae en aeaete aie asa tk nee 161
XVII. Note on a Hermaphrodite Frog. By R. C. Punnett,
BAG CE BEOER A), hoc awa. poe shen eee treme ERNE 179
XVIII. Descriptions of new Batrachians and Reptiles collected by
Mr. P. O. Simons in Peru. By G. A. Bouteneer, F.R.S......... 18h
XIX, Descriptions of new Batrachiins and Reptiles from the Larut
Hills. Perak By .G.A. BovimNnenn, FURS). ota. teehee 186
XX. Description of a new Lizard from Jamaica, By G. A.
IBOULENGER, HEuiSe! Gross ate v's «ies Fh ae eee e atiaiiogs oeata Mere 193
XXIJ. Description of a new Lizard from British East Africa. By
GA. BOpUENGER, PURIS. “ity. sists ts sient es ue aepeperes Ocoee 194
XXII. On a new Species of the Genus Alepas (A. Lankestert),
from the Collection of the British Museum. By A. Gruven, Chargé
de Cours a la Faculté des Sciences de Bordeaux. (Plate VIII.) .. 195
XXIII. Two Spiders new to the British Fauna. By Groner H.
CarPEntTER, B.Sc, Lond., of the Science and Art Museum, Dublin. 199
XXIV. An undescribed Type of Rusine Deer. By R. Lyprxxrr. 204
XXV. On Two English Millipedes (Zuius londinensis, Leach, and
dulus' teutonicus, sp. 0.) 5. by, be. es ROCOCK exit a. tcp oie ae 206
XXVI. Descriptions of Two Species of Cyprea, both of the Sub-
genus Trivia, Gray. By James Cosmo Metviut, M.A.,F.L.S. .. 207
XXVIT. Description of a new Species of Papilio from Bwool,
North Celebes. By H. Grosr-Smita, F.ES., F.Z.S., &e. ......., 210
XXVHI. Notes on the Collection of African Phasgonuride formed
by Mr. W. LL. Distant in the ‘lransvaal &c., with Descriptions of
Two new Species.. By W. F. Kinpy, F.LS., FES. &. ...0.... 210
CONTENTS.
Vv
I.
Page
XXIX. Note on a Variation in the Number of the Genital Pouches
in Thalussema neptuni, Gaertner. By F. Hl. Stewart, M.A.,
Gatty Marine Laboratory, St. Andrews
XXX. Description of a new Species of Buprestide. By Cuas. O.
DEPP STISIURITO UHee Wek on ne atau ints, ayeu ike a ofan dyes Ghapaiaaine wins, eats oS
XXXII. Rhynchotal Notes.— VI. Heteroptera: Dinidorine,
Phyllocephalineg, Urolabidine, and <Acanthosomine. By W. L.
Distant
XXXII. Asiatic Tortricidae. By the Rt. Hon. Lorp Watsrne-
ER UaB Vc MPa ETM OR huron erry che foe seve eee eigen sin ong xs co araqnane ts
XXXII. Note on Variation in the Weasel and Hedgehog. By
G. E. H. Barrerr-Hamtton
New Books :—Zoological Results based on Material from New
Britain, New Guinea, Loyalty Islands, and elsewhere, collected
during the Years 1895, 1896, and 1897, by Arthur Willey,
DSe. “Lond., Hon. M.A.Cantab. Part IV.—A Monograph of the
Coleopterous Families Corylophide and Spherude. By the Rev.
pA a IVIEAUINTUED RY WIS oye Vl essyersy oy 20 oo 4 evrevsb eusy ce sey esol. wk sraite ah sy on haaatione 246,
Priority or Usage, by F. Jeffrey Bell, M.A. &e.
Cele tei OOo Ls CECE OEO RS
NUMBER XXXIII.
XXXIV. On some Malacostracous Crustaceans from Malaysia in
the Collection of the Sarawak Museum. By W. I. Lancuesrer,
M.A., King’s College, Cambridge. (Plate XII.)
XXXY, On new Species of Histeride and Notices of others. By
G. Lewis, F.L.S. (Plate X.)
XXXVI. Descriptions of Five new Species of Pangonine from
South America. By Miss GERTRUDE Ricarpo
S)ieern e's eet 5a, oilaciar iw! @) 8)
i) ajN) ime’ iel \q)p ol ce! (6 Re) (ej tet oi.m) 6) mh. aM 8) ie rel a) le Le: el teuie? ie!
AOD 0. 00) Sr ONC i
XXXVII. Descriptions of new Rodents from Western South
America. By OLDFIELD THOMAS .
XXAVIII. On the “ Tohi,” the East-African Reedbuck currently
known as Cervicapra bohor. By OLDFIELD THOMAS
XXXIX. New Species of Eastern and Australian Moths. By
Colonel C.iSwinnon, MUA. F.GS, Se. sk Sas ihe
XL. On anew Theriodont Reptile (Jefidosuchus primevus) from
the Karoo Beds, South Africa. By R. Broom, M.D., B.Sc. ...... :
XLI. Some new Arachnida from Cape Colony. By R. I. Pocock.
XLII. Asiatic Tortricide. By the Rt. Hon. Lorp WaALSsINGHAM,
M.A., LL.D., F.RS.
New Books :—A Treatise on Zoology. Edited by E. Ray LanKESTER,
M.A., LL.D., F.R.S. Part II. The Echinoderma. By F. as
BATHER, M. mw , assisted by J. W. Grecory, D.Sc., and EK.
GoopricH, M. a Monograph of Christmas Island. P hy “ical
Features and Geclogy by CHarLes W. Anprews, B.A. ; with
Descriptions of the Fauna and Flora by numerous Contri-
UNEOES 5 2 Cae rpms 82! ae) Hriy'sy 5 65) A nc Go auce Oe Mega o41,
Leo SAIN Len ORDR CRC EC. UBS OAC OM tet Sa C ery CC LACS Oy EN Gy WeCeeL Yer Ve 5
218
219
247
248
vi CONTENTS.
NUMBER XXXIV.
Page
XLUI. Description of Sponges from Funafuti. By R. Kinx-
PATRICK, Assistant, Natural History Museum. (Plates XIII-XV.) 345
XLIV. The Scorpions of the Genus Heterometrus. By R. L
POCOGKE Nan ches <ileueenatn ie Phage ac hiels Ahhe B RID Pee Oe eremne 362
XLV. Rhynchotal Notes.—VII. Heteroptera: Fam. Coreide. By
Ws Ts. STANT 7 c/o cer itearain ne ee etcve re alate tiateeate cree ier lca. ceiehe ne Paced 366
XLVI. Notes on some Insects from the Yang-tse-Kiang. By
Sig SS ary sp oe PS PM he Die ers eee ee eos gers Si 380
XLVII. Descriptions of new Rodeuts from Western South
Anierica, By OLDPIELDDHOMAS 2 huhinscis aeieee fe ee Heese Pea 383
XLVIII. A new Dassie from North Nyasaland. By OLpFrELD
PROM AS j- Was se ap a.e cia) ste Yb opi oka he AES Boa os et ony OnE 387
XLIX. Description of a new Species of Ornithoptera. By H.
GROSE-OMMrHA eb ARRIGO ZiS: Ose. 0k i. eee ce at «css Sains 388
L. Descriptions of Brazilian Coccide. By ApotpH HeMPEL,
Rare UO, bream Nd SS, Beret att tag eps i Sima eons ER catcher nee 389
LI. Descriptions of new Species of Japanese Laa:d-Shells. By
$i. GU, BLS «sass Sie ose, hens gee ne ee ee ee 398
LU. Asiatic Tortricide. By the Rt. Hon. Lonp Watstneuam,
ILA ea DD SERS? 5 chai Gis Glave dleaeie bods: coset eee eve CM Le 401
LIL. Deseription of a new Snake of the Genus <Ablabes from
Bunnie. By GA. BouLENGER, BARS... Stews Se5.po te ee 409
LIV. Descriptions of new Genera and Species of Hymenoptera.
By BOA MEROR a Rissitetes cote setae ae eet atari eta tere ete . 410
LY. On the Presence of Echis coloratus, Giinther, in Africa. By
Jonun AnpERson, MAD; ELD. iS. 2 aise cae ice: oo > tee 419
New Books:—In Bird-land with Field-glass and Camera. By
Oniver G, Prxr.—The Birds of Eastern North America. Parts
I., 11. By Cuarres Cory.—A Natural History of the British
Lepidoptera. A Textbook for Students and Collectors. By
J. W; Torr; FOES. Wola. cies vie pee ee 426, 426
NUMBER XXXV.
LVI. Asiatic Tortricide. By the Rt. Hon, Lorp WatsineHam,
NSA LD ERS. ies... pics ae een ese eet he peer kc i neice 429
LVII. Descriptions of some .Species of Cvccide collected by
Mr. James Lidgett in Victoria, Australia, By EK. Ernest GREEN,
F.E.S., Government Entomologist, Ceylon. (Plate XI.).......... 448
LVIII. Descriptions of new S;ecies of Japanese Land-Shells. By
AeA UDG, ZI. « pro a-hole hatene abet ebayeta stat atlo’a Sela c's Ste ea deeds ar 453
CONTENTS. Vil
LIX. Notes on the Neuropterous Family Nemopterrde. By
Metre, FLS:, Fite ties sete ews rads lee eta wee ene ss 456
LX. On Eyuus Penricei, a Representative of the Mountain Zebra
(Equus zebra, L.) discovered by Mr. W. Penrice in Angola. By
Pe CID "TOMEAGS | seat atretelscet ead «nyu d dais sw bln wy ole oa se bt aun letphe. aie 465
LXI. New Peruvian Species of Conepatus, Phyllotis, and Akodon.
Aig PE By MEE VEY PL EEOBEA 5 eyafe st <n) aces; wn)'0 0-57 oie o-e « dae wre eB men oasis ais 466
LXII. A new Free-tailed Bat from Central America. By Grrrit
SAN EREAAIT ASE” CN Pets oh, rae oh pe TCS Ake iy ob ols ae da Werner weve 470
LXIII. Natural History Notes from H.M. Royal Indian Marine
Survey Ship ‘ Investigator,’ Commander T. H. Heming, R.N., com-
manding.—Series III., No. 4. Some Results of the Dredging Season
1899-1900. By A. F. McAro te, B A., M.B., Capt. I.M.S., Surgeon-
Naturalist to the Survey ............ BC ere See ne eo 471
LXIV. Diagnoses of new Fishes discovered by Mr. J. E. S. Moore
in Lake Tanganyika. By G. A. Boutenarr, F.RS.,..........- 478
LXY. Note on Diatoms from Chincha Guano. By C. MEREscH-
TP Sia Oe ad OG | eae ae 481
LXVI. Some new African Theraphosoid Spiders in the British
Mrapemims sy bee LP OCOGK . 55 soa 5 « Geers etapa ryerated 66 elas ce 0 slols ofa 489
LXVII. Descriptions of new Genera and Species of Hymenoptera.
ERE AMEE MONG ica lout Save de ete sk eeae a eelnero RA EMU, vcttieis 495
New Books:—Catalogue of Eastern and Australian Lepidoptera
Heterocera in the Collection of the Oxford University Museum.
Part I. Noctuina, Geometrina, and Pyraldina. By Col. C.
SwinHog, M.A., F.LS., F.ZS. FES. Pterophoride and
Tineine by the Right Hon. Lorp Watsineuam, M.A., LL.D.,
F.RS., &e., High Steward of the University of Cambridge, and
Joun HartLtEy Durrant, F.E.S., Memb. Soc. Ent. France.—
Sexual Dimorphism in the Animal Kingdom. By J. T. Cun-
NINGHAM, M.A.—Colour in Nature. By Marion I. New-
UR errs eerenicler tes osc fet: laa ae wna late Te gos we 506, 507
NUMBER XXXVI.
LXVIII. A suggested Origin of the Segmented Worms, and the
Problem of Metamerism. By H. M. Bernarp, M.A. Cantab. .... 509
LXIX. On some little-known African Silurid Fishes of the Sub-
family Doradine. By G. A. Boutenesrr, F.R.S. .............. 520
LXX. Descriptions of new Genera and Species of Hymenoptera.
Pee ONION aks Sirs oie Sct ay Se NP MOTE AO ae eee ass Hee a ee has 530
LXXIL. Plankton Studies.—IiL. On Platydorina, a new Genus of
the Family Volvocide from the Plankton of the Ilinois River. By
eA. Roromprine. (Plate VIL) ogc. ace ns chads ce Se ee 541
LXXII. On the Luminous Organs of Selachian Fishes. By Rup.
Burcxuarnt, Nat. Hist. Mus, 8S. Kensington
Vill CONTENTS.
Page
LXXITM. Mormopterus Whitleyi—a new Species of Bat from
AY; Aitica:* By R.-F) Scxaerr, PhDs Be: See 569
LXXIV. A new Bat from Peru. By Gerrit S. Mrnurr, Jr. .. 570
LXXV. Reptiles and Batrachians collected in German New
Guinea by the late Dr. Erik Nyman. By Dr. Ervar LONNBERG .. 574
New Book :—The Story of Bird-life. By W. P. Pycrarr........ 582
Proceedings of the Geological Society 2.0.2... cece eee eens ... 583
MOOR Goyongtr vs Os: cig Pola be ots Soh A ete bb ok Vem Se Pk eee Ack: 584
PLATES IN VOL. VI.
Prater I. Teeth of Elasmobranch Fishes from the Parana formation.
II. Neuroptera from Sierra Leone.
III. British Amphipoda.
IV. New species of Forficularia.
we Pleodorina illinoisensis.
VI. §
Vil. Platydorina caudata.
VIII. Alepas Lankesteri.
IX. Anatomy of an hermaphrodite frog,
X. Species of Histeridee.
XI. New Australian Coccidie.
XII, Crustaceans from Malaysia.
XSLT.
XIV. > New Sponges from Funafuti.
XY.
XVI. Diatoms from Chincha guano.
ERRATUM.
Page 71, lines 18 and 27, for sectors of the arculus
read sectors of the triangle.
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES. ]
Cpeeoccrerecnacce ... per litora spargite muscum,
Naiades, et circiim vitreos considite fontes:
Pollice virgineo teneros hic carpite flores:
Floribus et pictum, dives, replete canistrum.
At vos, o Nymphe Craterides, ite sub undas ;
Ite, recurvato variata corallia trunco
Vellite muscosis e rupibus, et mihi conchas
Ferte, Dew pelagi, et pingui conchylia succo.
N. Parthenii Giannettasi, Bel.
No. 31. JULY 1900.
I.—On some Fish-remains from the Parana Formation,
Argentine Republic. By A. Smita Woopwarp, LL.D.,
BPS.
[ Plate I.]
THERE is an interesting marine deposit exposed in the banks
of the River Parana near the city of Parana, in the province
of Entrerios, Argentine Republic. It has been known since
the explorations of d’Orbigny and Darwin, and is of special
importance not only as containing the remains of land-
mammals itself, but also as being intimately associated with
other deposits which yield abundant evidence of extinct
mammalian faunas. Since marine fossils are of much more
value in determining the age of a formation than the remains
of land animals, this small Parana deposit may therefore be
expected to afford a clue to the geological age of some of the
South-American terrestrial faunas, concerning which various
opinions have been expressed. Its interest in this connexion
has already been recognized by several observers, including
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. wy:
2 _ Dr. A. S. Woodward on some
Bravard *, Doering +, Ameghino}, Burmeister §, and Stelz-
ner ||, of whom the first three consider that its marine fossils
indicate an Eocene or Oligocene age, while the two latter
place the formation in the Upper Tertiary. The latest
examination of its fish-remains is supposed to have proved
its equivalence or homotaxy with the Eocene formations of
the northern hemisphere §.
Fortunately a considerable number of the Parana fish-
remains are the teeth of sharks, which may be supposed to
have had a very wide distribution in the oceans at the time
when they flourished. ‘The evidence of geological age which
they afford may thus be regarded as of considerable value.
I therefore propose to record a few observations on the Hlas-
mobranch fossils from the Parana collections in the National
Museum of Buenos Aires, the La“ Plata Museum, and the
San Paulo Museum, which have been kindly lent to me by
Dr. Carlos Berg, Dr. F. P. Moreno, and Dr. H. von Jhering.
The loan of these fossils has also enabled me to identify a
small series of Parana fish-remains labelled by Bravard in
the British Museum.
1. Raja Agassiz’, Larrazet.
1886. Raja Agassiz, Larrazet, Bull. Soc, Géol. France, [3] vol. xiv.
p- 259, pl. xiii. figs. 1-6.
The large dermal tubercles of an extinct skate thus
described are still known only from the Parana formation.
One typical specimen in the La Plata Museum measures no
less than 0°088 m. by 0°075 m. across, by 0°03 m. in
maximum thickness.
2. Dynatobatis paranensis, Larrazet.
1886. Dynatobatis paranensis, Larrazet, loc. cit. p. 268, pl. xiv. figs. 1-4.
These dermal tubercles evidently belong to a member of
* A. Bravard, ‘ Monografia de los Terrenos Marinos Terciarios de las
Cercanias del Parana’ (Parana, 1858).
+ A. Doering, in Roca, ‘Informe Oficial Com. Cient. Exped. al Rio
Negro,’ pt. 111, (1882) p. 429.
{ F. Ameghino, ‘Contribucion al Conocimiento de los Mamiferos
Fosiles de la Republica Argentina’ (1889) p. 14.
§ H. Burmeister, ‘ Description Physique de la République Argentina,’
vol. ii. (1876) p. 219.
|| A. Stelzner, ‘ Beitrage zur Geologie und Paleontologie der Argentin-
ischen Republik,’ pt. i. (1885) p. 189.
q G. de Alessandri, “ Ricerche sui Pesci Fossili di Parana (Repubblica
Argentina),” Atti R. Accad. Sci. Torino, yol. xxxi. (1896) pp. 715-750,
with plate.
Fish-remains from the Parana Formation. 3
the family Trygonide, as already observed by Jackel *.
There are two typical examples in the National Museum,
Buenos Aires.
3. Myliobatis americanus, Bravard.
1858. Myliobates americanus, A. Bravard, Monogr. Terr. Marinos Tere.
Parana, p. 53.
1896. Mylhobates americanus, G. de Alessandri, Atti R. Accad. Sci.
Torino, vol. xxxi. p. 724, pl. i. fig. 6.
The best-known specimen is described and figured by
Alessandri, but this is too imperfect for specific determination.
It seems probable that some of the detached teeth labelled as
belonging to this species are truly referable to RAc‘noptera.
4. Cestracion paranensis (Alessandri).
1896. Acrodus paranensis, G. de Alessandzri, loc. cit. p. 723, pl.i. fig. 5.
There are two teeth of this species in the San Paulo
Museum. On careful comparison with Cestracion they are
found to agree exactly with the larger grinding-teeth in this
existing genus. There is therefore no justification for
referring them to the extinct Mesozoic genus Acrodus.
5. Odontaspis elegans (Agassiz). (PI. I. figs. 1-5.)
1843. Lamna elegans, LL. Agassiz, Poiss. Foss. vol. iii. p. 3869, pl. xl. 8,
fic. 24 (non p. 289, pl. xxxv. figs. 1-7, pl. xxvii. a, figs. 58, 59),
1858. Lamna elegans, A. Bravard, op, eit. p. 52.
1858. Lamna unicuspidens, A. Bravard, op. cit. p. 52.
1885, Lamna eleyans, F. Noetling, Abh. Geol. Specialk. Preussen u.
Thiiring. Staaten, vol. vi. pt. 3, p. 61, pl. iv.
1896. Odontaspis elegans, G. de Alessandri, loc. cit. p. 720, pl. i. fig. 1.
1896, Odontaspis Hopet, G. de Alessandri, loc. cit. p. 720, pl. i. fig. 2.
1899. Odontaspis elegans, A. S. Woodward, Proc. Geol. Assoc. vol. xvi.
p. 8, pl. i. figs. 15-18.
All the teeth from Parana referred to this species are rather
small, and five well-preserved specimens are represented in
Pl. I. figs. 1-5. It seems impossible to distinguish them
from the typical teeth of O. elegans from the European Eocene,
but the striation of their inner face is perhaps more feebly
marked than in the latter. Most of the specimens are much
rolled and abraded. When their inner face is worn quite
smooth they are of the form referred to O. Hopet, Ag., by
Alessandri; when they have lost their lateral denticles they
are the Lamna unicuspidens of Bravard, as shown by his
label in the British Museum.
* O. Jaekel, ‘ Die Mocaenen Selachier yom Monte Bolea ’ (1894), p. 140,
1*
4 Dr. A. S. Woodward on some
6. Oxyrhina hastalis, Agassiz. (PI. I. figs. 6-8.)
1838-43. Oxyrhina hastalis, L. Agassiz, op. cit. vol. iii. p. 277, pl. xxxiv.
figs. 3-18, 15-17.
(?) 1858. Squalus eocenus, A. Bravard, op. cit. p. 51.
1881. Oxyrhina Agassizi, R. Lawley, Studi Comp. Pesci foss. coi viv.
generi Carcharodon, Oxyrhina, e Galeocerdo, p. 93, pls. v.-ix. (Oxy-
rhina). {The species renamed by Lawley.
1889. Oxyrhina hastalis, A. S. Woodward, Catal. Foss. Fishes B. M.
pt. i. p. 385.
Several teeth in the National Museum, Buenos Aires,
agree both in size and shape with the European Miocene and
Pliocene teeth ascribed to this species by Lawley. A well-
preserved anterior mandibular tooth is shown from the outer
face and in side view in Pl. I. figs. 6, 6a, while two
characteristic upper teeth, the first with imperfect base, are
seen, outer aspect, in figs. 7, 8.
7. Carcharodon megalodon, Agassiz. (PI. I. fig. 9.)
1885-43. Carcharodon megalodon, L. Agassiz, op. cit, vol. ili. p. 247,
1. xxix,
188. Carcharodon gigas, R. A. Philippi, Zeitschr. f. gesammt. Naturw,
vol, li. p. 685, pl. xix.
1881. Carcharodon megalodon, R. Lawley, op. cit. p. 35, pls. vi—xi.
(Carcharodon).
1889. Carcharodon meyalodon, A. 8. Woodward, Catal. Foss, Fishes
B. M. pt. i. p. 416.
There are two upper teeth of this species from Parana in
the National Museum, Buenos Aires, one being in the Lelong
collection, the other presented by Sefior Don Marcos Sastre.
The first is considerably the larger, and is shown of two
thirds the natural size in Pl. 1. fig. 9. It evidently
belongs to the middle of the side of the upper jaw, and is
quite unabraded. Another upper lateral tooth, from a Tertiary
formation at Coquimbo, Chili, has been described by Philippi
under the name of C. gigas, and teeth either of the same or
a closely related species occur in the Patagonian formation
of Chubut. The Chilian specimen was found in a deposit
which has yielded teeth indistinguishable from those of the
existing Carcharodon Rondeleti (A. S. Woodward, loc. cit.
1889, p. 421).
8. Carcharias (Prionodon) obliquidens (Bravard).
1858, Squalus obliquidens, A. Bravard, op. cit. p. 51,
1858. Lamna amplibasidens, A. Bravard, op. cit. p. 53.
1858, Lamna serridens, A, Bravard, op. ert. p. 68,
Fish-remains from the Parana Formation. 5
1896. Curcharias (Aprionodon) Gibbesii, G. de Alessandri, loc, ett.
p. 721, pl. i. fig. 3.
1896. Coraz: aff. falcatus, G. de Alessandri, loc. cit. p. 722, pl. i. fig. 4.
The meaning of Bravard’s specific names quoted above is
indicated by his labels in the British Museum and in the
National Museum, Buenos Aires. Two teeth in the British
Museum marked Lamna obliquidens are upper lateral teeth
of Carcharias (Prionodon); numerous specimens in the
National Museum labelled Zamna serridens are obviously
upper teeth of the same species, while other specimens labelled
Lamna amplibasidens in both museums are for the most part,
if not all, lower teeth of the same fish. ‘he majority of the
lower teeth are distinctly serrated, like the specimen erro-
neously referred to C. (Aprionodon) Gibbesi by Alessandri ;
but many of the smaller specimens exhibit quite smooth
edges—possibly a character of immaturity *. Some of the
upper teeth have been referred to Corax by Alessandri; but
I have made sections of several examples identical in shape
with that represented in his fig. 4, and found all to have
a central cavity. I therefore regard his determination of
Corax from the Parana formation as a mistake,
9. Galeocerdo aduncus, Agassiz. (Pl. I. figs. 10, L0 a.)
1835-43. Galeocerdo aduncus, L. Agassiz, op. cit. vol. iii. p. 231,
pl. xxvi. figs. 24-28,
1889. Galeocerdo aduncus, A. 8. Woodward, Catal. Foss. Fishes B. M.
pt. i. p. 444.
Like those of Carcharias the species of Galeocerdo are
difficult to determine, but the tooth from Parana represented
in Pl. I. figs. 10, 10a, seems to belong to the side of the
jaw of the European and North-American Miocene species
G. aduncus. It is readily distinguished from the Kocene
G. latidens by its deeper crown and finer serrations.
10. Hemipristis serra, Agassiz. (PI. I. figs. 11, lla.)
1835-43. Hemipristis serra, L. Agassiz, op. eit. vol, iii. p, 237, pl. xxvii.
figs, 18-30, 3
1889, Hemipristis serra, A.S. Woodward, Catal. Foss, Fishes B, M,
pt. i. p. 449.
This species is rare in the Parana formation, but a few
well-preserved upper teeth are unmistakable. ‘The best
specimen, not in the least abraded, is shown in Pl. I,
figs. 11, 11a. It is in all respects a typical upper tooth.
* A, Giinther, Catal. Fishes B, M. vol. viii. (1870) p. 357,
6 On some Fish-remains from the Parana Formation.
11. Undetermined Stluroids cc.
Remains of Siluroid fishes are very abundant in the Parana
formation, but all the known specimens are too fragmentary
for exact determination. They evidently represent Arius,
Pimelodus, Platystoma, and other genera which still live in
the fresh waters of South America.
Fragments of Characinoids also occur and are easily recog-
nized; but other remains, such as those which have been
referred to Sparoids and Labroids by Bravard and Alessandri,
are not satisfactorily determinable. The so-called tooth of
Lepidosteus described by Alessandri (loc. cit. p. 726, pl. i.
fig. 8) is not sufficient to indicate the presence of this genus.
If the foregoing determinations of the fish-remains from the
Parana formation be accepted, it is evident that Alessandri’s
argument for the Hocene age of this deposit has no founda-
tion in fact. The so-called teeth of Acrodus and OCorax
(which are typically Mesozoic genera) become, on renewed
examination, evidence of Cestracion and Carcharias, which
range throughout the Tertiary formations and survive at the
present day. It is by no means certain that the teeth deter-
mined as Odontaspis elegans in the Parana collection belong
to the same fish as those originally thus named in European
Kocene collections; and even if Carcharias (Aprionodon)
Gibbest were correctly identified, the Phosphates of South
Carolina, from which the type specimens of that species were
obtained, seem to include fossils of all Tertiary ages from the
Eocene to the Pleistocene. In fact, the only species in the
Parana collection which seem to be of real importance for
stratigraphical purposes are Oayrhina hastalis, Carcharodon
megalodon, and Hemipristis serra. All these in Europe are
exclusively Miocene and Pliocene fossils, while the only teeth
from the undoubted Hocene of North America (Alabama)
commonly referred to the same species are comparatively
small, not of typical size like those from Parana. Moreover,
it is to be noted that several teeth of Oayrhina hastalis and
Carcharodon megalodon were dredged from the bed of the
South Pacific Ocean by the ‘Challenger’ expedition *, this
discovery probably implying that these great sharks did not
become extinct until quite the latest geological times. I
therefore conclude, with Burmeister and Stelzner, that the
* J. Muwray and A. Renard, ‘‘ Deep-sea Deposits” (‘Challenger’
teports, 1891), pls. v., vi.
On Bees from New Me.xico. 7
Parana formation is truly of late Tertiary age, and may
probably be correlated with the Pliocene of the northern
hemisphere.
EXPLANATION OF PLATE I.
Teeth of Elasmobranch fishes from the Parana formation, Province
of Entrerios, Argentine Republic.
Figs. 1, 1a. Odontaspis elegans, Ag. ; outer and lateral aspects.
Fig. 2. Ditto; inner face.
Figs. 3, 3a. Ditto; inner and lateral aspects.
Figs. 4, 5. Ditto ; inner face.
Figs. 6, 6a. Oxyrhina hastalis, Ag.; outer and lateral aspects.
Figs. 7, 8. Ditto; outer face.
Fig. 9. Carcharodon megalodon, Ag.; outer face, two thirds nat. size.
Figs. 10, 10 a. Galeocerdo aduncus, Ag. ; outer and inner faces.
Figs. 11, lla. Hemipristis serra, Ag.; outer and inner faces.
The original specimens are in the National Museum, Buenos Aires, and
all the figures except no. 9 are of the natural size.
Il.—The New Mexico Bees of the Genus Megachile and a
new Andrena. By T. D. A. CocKkERELL, Professor of
Entomology, New Mexico Agricultural College.
Megachile Wootoni, Ckll., 1898.
The type was a male. I have before me two females from
the Rio Ruidoso, about 6900 feet, at flowers of Verdascum
thapsus, July 23 (C. H. T. Townsend). They differ at once
trom the female of WM. calogaster by having little or no black
hair on the vertex and mesothorax; one has the orange
scopa as in calogaster, but the other has the scopa orange
in the middle and black at the sides, thus approaching
M. melanophea.
This belongs to the subgenus Megachile s. str., as restricted
by Friese.
Megachile sapellonis, sp. n.
? .—Length 17-22 millim., the shorter examples having
the abdominal segments retracted.
Black, with rather thin pubescence, white on sides of
face, cheeks, pleura, metathorax, femora, sides of first seg-
ment of abdomen, and hind margins of second and following
segments, more or less interrupted in the middle, at least on
8 Prof. T. D. A. Cockerell on
the second segment ; vertex, mesothorax, and scutellum with
erect black hair; basal portions of second and following
abdominal segments with short black hair ; labrum with
erect pale orange hair; hair on inner side of tarsi more or
less orange or reddish, on hind tarsi deep orange-ferruginous ;
ventral scopa entirely very pale yellowish to yellowish white,
never quite a pure white ; spurs bright ferruginous ; antenne
wholly black, reaching about to tegule; clypeus shining,
with strong punctures, sparse in the middle, its anterior
margin with four short teeth; head large, subquadrate ;
cheeks simple ; vertex broad, with strong punctures, dense in
the middle, where they are of two sizes; mandibles broad,
4-dentate, not counting the inner angle; mesothorax densely
punctured at the sides, more sparsely in the middle ; tegule
black, microscopically sculptured, punctured anteriorly ;
wings smoky ; claws with a ferruginous denticle at the base.
Allied to M. bucephala, Smith, but considerably larger.
Hab. Beulah, Sapello Cafion, N. M., July 26,2 2, one
at flowers of thistle (W. Porter); Beulah, Aug. 18, 2 9
(Ckll.) ; hill west of Beulah, Aug. 23, 1 9 (W. Porter) ;
Haivey’s Ranch, 9600 feet, Aug. 22,4 9 (Porter & Chili.) ;
South Fork, Hagle Creek, White Mts., about 8100 feet, at
flowers of Senecio Bigelovii, Aug. 18,1 2 (Townsend).
Greene, ‘ Pittonia,’ iv. p. 118, announces that the plant of
the White Mts., distributed as Senecio Rusbyt, is really
S. Bigelovit. It is visited also by Bombus improbus and
Andrena apacheorum.
Megachile pugnata, Say, 1837.
Santa Fé,5 g,2 9; Las Vegas, July 22, at flowers of
Rtudbeckia laciniata,1 9 (W. Porter); Beulah, July 26,3 3
(W. Porter) ; hill near Beulah, Aug. 19,2 9 (W. Porter).
The female is easily distinguished from the last by its
smaller size (not over 15 millim.) and the large tooth at the
lower hind angles of the cheeks. The abdominal bands are
entire.
The Santa Fé males were at flowers of Lactuca pulchella
and udbeckia, the females at Lactuca pulchella and
Grindelia.
Megachile fidelis, Cress., 1878.
Las Vegas Hot Springs, at flowers of Senecio Douglasit,
Aug. 10,3 9 (W. Forter) ; Burnt Cafton, Aug. 14, at flowers
of Cleome,1 9 (Sarah L. Mize); Las Vegas, July 6 and
Bees from New Mexico. 9
19, at flowers of Verbena, 2 g (Porter & Chil.) ; July 7, at
flowers of Senecio Douglasti, 1 9 (W. H. Rishel) ; Aug. 10,
at flowers of Grindelia squarrosa, 1 9 (S. L. Mize) ; San
Ignacio, Sept. 1, 1 2 (Lorter & Ckil.); Beulah, July 26,
1 9 (W. Porter); west fork of Gila River, July 16, g
(Townsend); Rio Ruidoso, about 6500 feet, July 31 and
Aug. 1, at flowers of Heliopsis scabra, 2 3 (Townsend).
The female differs at once from that of pugnata by the
yellow abdominal bands and the large tooth or lamina on
each side of the anterior margin of the clypeus.
Megachile exilis, Cress., 1872.
The male has the first three joints of the anterior tarsi
flattened at the side. The female resembles the male; the
ventral scopa is white, sometimes with a few black hairs at
the extreme tip ; in one specimen from the Rio Ruidoso the
scopa on the last three segments is greyish brown; the ante-
rior margin of the clypeus has two little teeth in the middle
and a broad tooth or lamina at each side.
West fork of Gila River, July 12 to 16, many males
(Townsend) ; Santa Fé, at flowers of Pentstemon Torrey? in
a garden, crawling on the outside of the flower, July 11,1 ¢
(Ckll.) ; Las Vegas Hot Springs, July 11, 1 g; Las Vegas,
July 18, at flowers of Lyctum vulgare, 1 9 (W. Porter) ;
Gallinas River at La Cueva, at flowers of Psoralea tenuiflora,
2 2 (Porter & Ckil.) ; Rio Ruidoso, about 6700 feet, at
flowers of Vicia near pulchella, July 29, 2 2, 6 @
( Townsend).
This species differs from most of its genus in not being
addicted to the Compositee.
Megachile occidentalis, Fox, 1894.
A narrow species, much like the last, but rather larger, and
both sexes with two spots of white hair on the anterior part
of the mesothorax. The female has the anterior margin of
the clypeus with a broad but shallow median excavation, in
the middle of which is a tooth ; the ventral scopa is white,
wholly black on the last segment.
Riley’s Ranch, at base of Organ Mts., Aug. 26, 1 2
(Ckll.) ; Las Cruces, June 8,1 ¢ (Ckll.) ; Mesilla, May 10,
at flowers of Prosopis glandulosa, 1 $ (Jessie Casad) ;
April 22, at flowers of Phacelia, 1 g (Chll.).
This is a species of the Middle and doubtless Lower
Sonoran, whereas M. exilis belongs to the Upper Sonoran
and transition zones,
10 Prof. T. D. A. Cockerell on
Megachile latimanus, Say, 1823.
The female is easily distinguished by its rather large size
(13-15 millim.), shovel-shaped abdomen, ventral scopa whitish
at base, becoming ferruginous at apex, broad face, and the
greyish-white pubescence of the vertex and mesothorax
hardly at all mixed with black.
Santa Fé, July and August (Ckil.); Las Vegas, June 1,
3, June 9, ? (&. Devine); July 6, at flowers of Verbena
Macdougali, 1 2 (W. Porter) ; July 11, at flowers of Cleome
serrulata, 2 9 (M. Winters & Chil.) ; June 19, at flowers of
Medicago sativa, 1 9 (Ckll.) ; Aug. 11, at flowers of Petalo-
stemon candidus, 1 2? (W. Porter); Aug. 9, at flowers of
Grindelia squarrosa, 1 9 (W. Porter); San Ignacio, Sept. 1,
9 (Porter. & Ckll.).
This is a species of the transition zone, and it is remarkable
that it was neither found at Beulah nor in the White
Mountains.
Megachile fortis, Cress., 1872.
This species in the male exhibits a dichroism like that of
Anthophora occidentalis; the pale torm is the M. comata,
Cress., 1872, but in a long series it becomes impossible to
recognize it as aseparate species. The female has never been
described as such, but from analogy with M. latémanus and
the facts of distribution I feel certain that it is the insect
described by Cresson in 1878 as M. texana. The male
assigned to tevana by Cresson is a different species.
This species inhabits higher altitudes than M. latimanus,
though it also occurs in the transition zone. Las Vegas Hot
Springs, Aug. 10, at flowers of Senecio Douglasti and Verbena
Macdougali, 3 8 (W. Porter) ; Gallinas River, at La Cueva,
Aug. 6, 1 ¢ (W. Porter); San Ignacio, Sept. 1, g 9
(Porter & Ckll.); Beulah, end of August, 1 ¢ (Céil.) ; Mesca-
lero Indian Reservation below the Agency, at flowers of
Bigelovia graveolens, var. glabrata, Oct. 1 and 2,7 3 (Ckil.) ;
Rociada, Aug. 20,1 2? (W. Porter) ; South Fork of Eagle
Creek, 8000-8200 feet, Aug. 18-20, g at flowers of Heliopsis
scabra, at flowers of Erigeron macranthus and Senecio
Bigelovit (Townsend) ; Rio Ruidoso, July 8, at flowers of
Pentstemon, ¢ (Wooton); July 21—Aug. 3, 6500-7600 feet,
males at flowers of Verbena Macdougali (some of these var.
comata), Vicia near pulchella, Monarda stricta, Heliopsis
scabra, Astragalus humistratus, and Potentilla Thurber, but
only single specimens on the last two; females at flowers of
Bees from New Mexico. 11
Verbascum thapsus (very many), Rhus glabra, and Verbena
Macdougali.
Megachile relativa, Cress., 1878.
This and the next form a little group distinguished in the
female by the ferruginous scopa, punctures of mesothorax
relatively sparse in the middle, abdomen with hair-bands
mostly interrupted, and the clypeus normal. MM. relativa is
rather small (about 11 millim.) and has the hair-bands of the
abdomen white, overlapped with orange hairs, so as usually
to appear yellowish or orange. In monardarum there are no
overlapping hairs, unless it may be some black ones.
Rio Ruidoso, July 30, 1 9 (C. M. Barber); Rociada,
Aug. 20, 1 9 (W. Porter); Harvey’s Ranch, Aug. 22
(W. Porter); Beulah, July 26, Aug. 18, Aug. 25, many
females, one at flowers of Polemonium filicinum (W. Porter).
The species evidently extends right across the northern
part of the continent, as I have a female from Olympia,
Washington State, at flowers of Potentilla, June 30 (T.
Kincaid).
Megachile monardarum, sp. n.
? .—Length 13-16 millim.
Similar to redat’va, but differing in its larger size; the
abdominal hair-bands (very broadly interrupted on the second
and third segments, narrowly interrupted or entire on the
fourth and fifth) white instead of orange; the black hairs on
the dark parts of the abdomen longer, conspicuous at the
sides when the abdomen is viewed from above; the vertex
more sparsely punctured at the sides, and the two apical teeth
of the 4-dentate mandibles perhaps not so long.
Hab. Hill near Beulah, Aug. 19 (W. Porter); Rio
Ruidoso, 6500-6700 feet, at flowers of Monarda stricta and
Astragalus humistratus, July 27-31 (Townsend).
Megachile pruina, Smith, 1853.
Our insect is M. facunda, Cress., now considered to be
identical with pruzna. The following records are based on
the male, which is distinguished from the allied males flying
in the same region by the simple anterior tarsi and the large
teeth at the apex of the abdominal venter. It is a larger
insect than M. Townsendiana, though variable in size
(11-14 millim.), and does not have the emarginate apical
12 Prof. T. D. A. Cockerell on
portion of the abdomen curved downwards as in that species ;
but the two are closely allied.
Las Vegas Hot Springs, Aug. 10, at flowers of Senecio
Douglast’, two (W. Porter); Las Vegas, July 22, at flowers
of Rudbeckia laciniata (W. Porter); Aug. 9, at flowers of
Melilotus alba (W. Porter); Aug. 12 (A. Garlick).
Megachile Townsendiana, Ckll., 1898.
Las Cruces, Aug. 23, 4 ¢ (Chil. & Townsend) ; Rincon,
July 6, at flowers of Actenella, 2 g (Ckll.).
A species of the Middle Sonoran zone.
Megachile pollicaris, Say, var. pereximia nov.
6 —Length 13 millim.
Anterior and middle femora and tibie ferruginous ; hind
femora and tibie black; middle tibie with a black basal
streak behind; anterior tibiz yellow in front at apex; second
joint of anterior tarsus white, with a long linear apical
process; apex of flagellum somewhat dilated.
Hab. Beulah, May 30, at flowers of wild plum (W.
Porter).
Megachile Casade, Ckll., 1898.
Found as yet only in the Mesilla Valley.
The female has the scopa entirely white; clypeus and
mandibles normal; punctures of mesothorax large, the
intervals between them shining, and presenting some ex-
tremely minute punctures.
Megachile soledadensis, sp. n.
o.—Length 10 millim.
Agreeing with the description of M. legalis, Cress., but
possessing the following peculiarities:—Hyes pale green ;
apical joint of flagellum broad and flattened ; anterior coxa
with a bright ferruginous hair-patch, their spines short,
oblique, pointed, and slightly curved ; anterior femora with
the basal two fifths above pale ferruginous, beneath with a
large yellowish keel, subtriangular in shape, like that of a
racing-yacht ; anterior tibie three-sided, ferruginous, except
on the outer side ; anterior tarsi yellow, moderately dilated,
with a long white fringe, which is brownish within, second
joint with a conspicuous black spot within ; hind tibiz stout,
their tarsi broadened ; hair of vertex and thoracic dorsum
Bees from New Mexico, le
slightly greyish, not at all mixed with black; tegule clear
testaceous ; wings nearly clear, costal nervure ferruginous ;
abdomen rather broad, but parallel-sided, clothed with
greyish-white hair, hind margins of the segments with con-
spicuous entire white hair-bands ; apex rounded, irregularly
serrulate, not emarginate; apex of venter with a rather
large tooth on each side and a small one in the middle.
Has the general aspect of 1. Townsendiana, from which its
front legs and abdominal structures at once separate it. The
non-emarginate apex of abdomen allies it with M. manifesta,
but it has not the long median ventral apical tooth of that
insect.
?.—Similar to the male, except for the usual sexual
differences. Hind border of scutellum and sides of meso-
thorax conspicuously bounded by white pubescence; apical
teeth of mandibles very little developed ; ventral scopa white,
black on the last segment.
Hab. Soledad Cafion, Organ Mts., Aug. 15, 2 g, 1 9
(C. H. T. Townsend); Mesilla Valley, at flowers of Baileya
multiradiata, 1 g (Townsend).
Megachile manifesta, Cress., 1878.
3 .—The abdominal bands, described as white by Cresson,
are often ochraceous.
?.—Similar to the male, except in the usual sexual
characters, the broader more shovel-shaped abdomen, and the
vertex and disk of mesothorax (but not the scutellum) being
clothed with black hair. ‘The abdominal hair-bands are entire
and very distinct, and the second and following segments
have their dark portions clothed with black hair. Ventral
scopa white, black on the apical segment and apex of penulti-
mate one, white, however, at extreme base of apical segment.
Las Vegas Hot Springs, Aug. 10, at flowers of Senecio
Douglasi, 1 § (W. Porter); Las Vegas, Aug. 9-14, at
flowers of Grindelia squarrosa, 14 g, 3 2? (S. L. Mize &
W. Porter); Aug. 11, at flowers of Petalostemon candidus,
1 ¢ (W. Porter). Apparently absent from the White
Mountain region.
Megachile cleomis, sp. n.
? .—Length 11-13 millim.
Pubescence mostly dull white, on vertex black, on meso-
thorax and scutellum thin, greyish white at the sides, black
in the middle; on the middle of the second and following
14 Prof. T. D. A. Cockerell on
abdominal segments black, very conspicuous at the sides
when the abdomen is viewed from above; white pubescence
dense on sides of face, pleura, tubercles, sides of metathorax,
sides of first abdominal segment, and on hind margins of
segments 1 to 5, forming conspicuous entire white bands;
ventral scopa white or with a yellowish tinge, black on last
segment; pubescence on inner side of tarsi dull ferruginous ;
clypeus ordinary, with strong punctures, well separated in
the middle; antenne short; vertex with large punctures;
mesothorax microscopically tessellate, with large punctures,
well separated on the disk ; abdomen inversely mitre-shaped,
strongly punctured ; tegule black, punctured all over; wings
dusky.
This agrees almost exactly with Cresson’s description of
the female M. grandis, but that is the female of M. pollicaris,
which our insect certainly is not. Among the females found
in New Mexico it is known by the white scopa, black on
the last segment, normal clypeus, rather broad form, and the
absence of any spots of white pubescence on the mesothorax
or white band between the mesothorax and scutellum. The
absence of the last-mentioned marks at once separates it from
the superficially similar M. sidalcee.
3 .—Length 11 millim.
Abdomen rather parallel-sided; antenne long, last joint
not modified; punctures of mesothorax closer; mesothorax
and scutellum with scarcely any dark hairs; face densely
covered with silky white hair; black hair on abdomen incon-
spicuous ; cheeks simple; anterior coxee armed with black
spines of moderate length; anterior femora ferruginous
beneath and with a ferruginous patch above; anterior tarsi
simple, but fringed with white hair ; margins of tegulee more
or less ferruginous ; apex of abdomen emarginate, irregularly
denticulate on each side of the emargination ; apex of venter
with three very short teeth.
This appears to be the male which Cresson supposed (erro-
neously, as I hold) to belong to his M. terana. It differs
from that of M. rufimanus by the armed anterior coxe.
Hab. Santa Fé, July 5-25, many males, one at flowers of
Cleome serrulata, 1 ? (Ckll.); Las Vegas, June 28-July 20,
both sexes numerous at flowers of Cleome serrulata, 1 9? at
flowers of Medicago sativa (alfalfa), one of each sex at flowers
of Verbena Macdougali (M. Holzman, N. Stern, H. K. Rishel,
A. Garlick, W. Porter, M. Winters, Ckil.) ; Albuquerque.
1 ¢, June 30 (CkIl.).
Bees from New Mexico. 15
Megachile cleomis, var. lippice nov.
@ .—Always has two transverse white hair-marks at the
front of the scutellum, but lacks the two marks at the front
of the mesothorax, which are present in M. sidalceew ; scopa
yellowish white, black on apical segment; mesothorax
between the punctures dull; tegulze more or less edged with
ferruginous; flagellum sometimes a little ferruginous beneath ;
abdomen with less black hair at the sides.
6 .—Usually with more black hair on the thorax, espe-
cially on the scutellum.
This is the Middle Sonoran race of the species. La Cueva,
Organ Mts., about 5300 feet, at flowers of Lippia Wrightir,
Sept. 3-5, 2 ¢,2 9 (C0. H. T. Townsend) ; West Fork of
Gila River, July 12-16, ? (Townsend); Las Cruces, ?,
Sept. 7 (Ck/l.), and males as follows:—June 16, at flowers
of Aster spinosus (Ckll.); Aug. 11 (Townsend); Sept. 4, at
flowers of Solidago canadensis, var. arizonica (Ckll.).
The following males have no black hair on the scutellum,
and offer no satisfactory differences from those of typical
cleomis :—Las Cruces, Aug. 24, at flowers of Solidago cana-
densis, var. arizonica; Mesilla, July 4 (C. M. Barber) ;
Mesilla Park, April 24, at flowers of Onobrychis (Jessie
- Casad); Fillmore Cafion, Organ Mts., Aug. 29 (Ckll.) ;
Lone Mountain, July 7 (Ckl/.).
The species is recognized in the male by the ferruginous
patch on the upperside of the anterior femora.
Megachile sidalcew, Ckll., 1897.
Described originally from the male. The subjoined records
are based on the female, which is very similar to that of
M. cleomis and is distinguished by the following combination
of characters:—Ventral scopa pure white, black at extreme
tip; a conspicuous white hair-band in the scutello-meso-
thoracic suture; two white hair-marks, converging poste-
riorly, on mesothorax in front; mesothorax microscopically
tessellate between the punctures ; borders of tegule more or
less ferruginous ; pubescence of face white, of vertex often
mixed with black; clypeus and mandibles normal, anterior
edge of clypeus smooth, often with a small median tubercle ;
abdomen mitriform.
Mesilla, May 13, two at flowers of Prosopis glandulosa
(Jessie Casad) ; June 24, one at flowers of Helianthus ciliaris
(Ckll.) ; Mesilla Park, two at flowers of Lsocoma Wrightii,
Sept. 11 (Ckll. & Porter) ; Las Cruces, Aug. 23 (Ckll.) ;
~
16 Prof, T. D. A. Cockerell on
east of Las Cruces, about 4000 feet, two at flowers of Senecio
filifolius, Sept. 10 (Townsend) ; Soledad Cafion, Organ Mts.,
one at flowers of Pectis papposa, Aug. 15 (Townsend).
The abdomen, viewed from above, does not show the
black hair at the sides, which is so conspicuous in MV. cleomis,
except on the last two segments. This separates it at a
glance from the typical female cleomis, but cleomis var. lippice
has conspicuous lateral black hair only on the last two
segments, as in sidalcee. It would be easy to regard lippie
and sidalcee in the female as forms of one species, but the
males are quite different in the structure of their anterior
legs. There is a bare possibility that the female here assigned
to stdalcee does not belong to it, but is really a form of lippie,
in which case one of the next two species should probably be
mated with s¢dalcee ; but in the absence of absolute proof I
have followed the apparent probabilities.
Megachile prosopidis, sp. n.
@ .—Length 124-154 millim.
Resembling 1. stdalcee in the pubescence, but abdomen
subcylindrical and parallel-sided ; pubescence white, not
hiding the surface except in particular places; the vertex and
mesothorax have some very scanty and short dark hair,
scarcely visible; pubescence dense, forming conspicuous
white marks at sides of face, below tegule, two white marks
on mesothorax in front converging posteriorly, a spot at
lateral hind angles of mesothorax and on hind margins of
abdominal segments, forming narrow entire bands; pubes-
cence also dense on pleura and sides of mesothorax, but no
well-defined band at the scutello-mesothoracic suture; band
on first abdominal segment produced at each side into a
triangular patch ; scopa white, black on last segment; legs
black ; margins of tegule dull ferruginous ; wings hyaline,
slightly smoky on margins; antenne black, flagellum
faintly ferruginous beneath ; mesothorax with large very close
punctures, not at all sparse on disk; clypeus deeply and
broadly emarginate, the sides of the emargination directed
inwards, the middle occupied by a large lobe, so that the
opening, if closed by an imaginary line connecting the oppo-
site lower corners, would have a reniform outline; labrum
with the apical lateral angles pointed, the apical middle
thickened but not produced, bearing a brush of brown hairs
directed somewhat backwards; mandibles broad, 5-dentate
counting the inner angle, only the apical tooth large.
Bees from New Mexico. 17
Allied to M. rufimanus.
Male unknown.
Hab. Mesilla, May 10, two at flowers of Prosopis glandu-
losa (Jessie Casad) ; Las Cruces, one (Townsend) ; Rincon,
July 5, at flowers of Chilopsis linearis, one (Ckll.).
The maxillary palpi are covered with bristles.
Megachile chilopsidis, sp. n.
? .—Length 14 millim.
In form, size, and pubescence just like MW. prosopidis, so
that, without looking at the face, one would take it for the
same species, but clypeus and mandibles entirely different.
Clypeus extremely short, produced in the middle into a very
broad truncate lobe, more than twice as broad as long, pro-
jecting, shining and punctured above ; labrum broadly trun-
cate and thickened at end, the lateral corners pointed, the
middle with a narrow brush of brown hairs directed obliquely
backwards; mandibles long and narrow, somewhat broadened
at apex, with three small teeth; a large open space between
the mandibles and the clypeus. First joint of labial palpi
shorter than in M. prosopidis, being not greatly longer than
the second.
Male unknown.
Allied to M. rufimanus, which resembles this species in
the mandibles, and the last more in the clypeus.
Hab. Mesilla, one at flowers of Prosopis glandulosa,
May 13 (Jessie Casad) ; Rincon, one at flowers of Chilopsis
linearis, July 5 (Ckil.).
Megachile popult, sp. n.
? .—Length about 13 millim.
Black, with white pubescence ; black on disk of mesothorax,
middle of scutellum, and vertex, but in these places thin,
wholly exposing the surface; hair of cheeks and sides of face
long, dense, and pure white; antenne short, hardly reaching
to tegule, flagellum faintly ferruginous beneath; vertex
strongly punctured, but the punctures separate; clypeus
shining, strongly punctured at the sides, the punctures evanes-
cent on the disk, anterior edge straight and normal; man-
dibles normal, 4-dentate counting the inner angle; first joint
of labial palpi very broad, shorter than the second; meso-
thorax shining, though microscopically tessellate, densely
punctured at the sides, sparsely in the middle; tegule
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 2
18 Prof. T. D. A. Cockerell on
piceous, punctured all over; wings hyaline, faintly smoky ;
tarsi with ferruginous hair on the inner side; abdomen long-
mitriform, the segments with entire white hair-bands ; ventral
scopa white, black on the last segment and sometimes the |
hind margin of the penultimate one.
There are no white hair-marks on the front of the meso-
thorax nor at the scutello-mesothoracic suture.
Superficially resembles I. cleomis, var. lippic, but easily
distinguished by the character of the mesothoracic punctuation
and the much longer second joint of the labial palpus.
Hab. Mesilla Park, campus of New Mexico Agricultural
College, April 16, cutting the leaves of Populus, to use in
preparing their nests.
Megachile vallorum, sp. n.
? .—Length 14-15 millim.
Much like MZ. populi, with the same kind of pubescence
and entire white abdominal bands, but differing in many
details. Black pubescence arranged on vertex, mesothorax,
and scutellum as in MZ. populi’; flagellum wholly dark or
faintly reddish beneath; mandibles with prominent sharp
teeth ; second joint of labial palpus a little longer than first ;
clypeus with large punctures, dense all over; anterior border
of clypeus with a broad shallow emargination, beneath which
is a conspicuous fringe of orange hair; mesothorax densely
punctured all over, the areas between the punctures wholly
dull; no white hair-marks on anterior part of mesothorax
nor at scutello-mesothoracic suture ; tegule dark, punctured
all over; wings hyaline, broadly smoky on outer margin, and
especially at apex; basal joint of hind tarsi long and flat,
considerably longer than the other joints together, covered on
inner side with orange-ferruginous hair; spines at apices of
first four tibize quite long, dark ferruginous ; abdomen, viewed
from above, showing abundant black hair at the sides of the
last two segments, but not the others; ventral scopa white,
varying to quite yellowish, black on last segment and apex
of penultimate one.
By the clypeal structure allied to M. montivaga, but other-
wise different.
Hab. Las Cruces (Agnes Williams); Mesilla, June 24,
entering burrow in an adobe wall (Ckl/.) ; Socorro, June 29,
two at flowers of a species of Composite (Ckil.).
Megachile heterodonta, sp. n.
? .—Superficially, in size, form, and colour, even to the
Bees from New Mexico, 19
tint of the abdominal bands and dark shading of the wings,
exactly like I. pugnata, but differing as follows :—-Cheeks
not toothed; head not so large behind the eyes; middle of
vertex with the punctures large and well-separated, the space
between them shining, though microscopically tessellate ;
first joint of flagellum shorter; clypeus ordinary, densely
punctured all over, with very large punctures, its anterior
margin inconspicuously tridentate and fringed beneath with
long orange hairs; mandibles stout, shorter than in pugnata,
4-dentate, the apical tooth long and pointed, the next tran-
cate, the next quadrate, broader than long, with its inner
corner somewhat produced, the innermost tooth short and
pointed ; mesothorax dull and as densely punctured as possible
all over; ventral scopa white, black on last segment. The
second joint of the labial palpus is conspicuously longer than
the first. The abdominal bands are very distinct.
Hab. Las Vegas, Aug. 1 (Porter & Ckll.) ; Fillmore Cajion,
Organ Mts., about 5700 feet, Sept. 1 (C. H. 7. Townsend).
Megachile brevis, Say, 1837.
Hab. Gallinas River, at La Cueva, Aug. 6,1 2 (Porter
& Ckil.).
This agrees with I. brevis, received from Mr. Charles
Robertson, and is the only genuine brevis in the New Mexico
collections. The insect is recognized by its rather small
size (11-12 millim.), wholly white ventral scopa, mesothorax
dull and densely punctured, abdomen mitriform, mandibles
_curved at the apex, the two apical teeth close together.
I have the species also from Baldwin, Kansas, July (J. C.
Bridwell).
Andrena sapellonis, sp. n.
9 —Length 94-11 millim.; g about 8 millim.
With a wholly dark face and very long antenna, the
flagellum entirely black. In Robertson’s table in Trans.
Amer. Ent. Soc. xviii. p. 50, this runs to A. salicis, but it is
a little larger, and differs from the description of the female
by having the basal process of labrum narrow but quite large,
produced, and rounded at the end instead of truncate; the
well-developed hair-bands on the second, third, and fourth
abdominal segments are white instead of fulvous. ‘The
clypeus, as in salicts, has a well-developed median impunc-
tate ridge, and the long rather dense hair of the thoracic
dorsum is ochreous, though that of the cheeks and pleura is
white. The mesothorax is dull, microscopically tessellate,
2
20 Prof. J. W. Gregory on the
with sparse punctures; the abdomen is impunctate. The
anal fimbria is greyish brown. ‘The enclosure of the meta-
thorax is granular and ill-defined. Tarsi dark in both sexes,
I had thought it possible that this might be Andrena tri-
zonata (Ashmead, as Ciulissa), so I sent a specimen to
Mr. Ashmead, who kindly compared it with his type, and
reported as follows:—‘'The Andrena sent is not my ¢ri-
zonata, although it superficially resembles it. Your specimen
is slightly larger, differently sculptured, and has quite a
different pygidial plate. The hind legs and tarsi are also
differently coloured. It is quite a different insect.”
These remarks relate to the female; the male of trizonata
is said to have a banded abdomen; that of sapellonis @ is
shining, without bands, though the first segment, lateral hind
margins of the two following, and whole hind margins of the
rest are clothed with rather pale brown hair, which is only
conspicuous under a lens.
The female sapellonis must resemble Robertson’s recently
described A. salicacea, but it differs from the description of
the latter as follows :—Pubescence of thorax above ochraceous ;
facial grooves white, their width about as great as length of
first flagellar joint; enclosure and sides of metathorax
rugose-reticulate, but sculptured alike; anal fimbria pale
brown. A. sapellonis agrees with salicacea in the process of
labrum, proportions of the first three flagellar joints, fuscous
pubescence on tibiz, and third submarginal cell at least twice
as long as the second. ‘The two doubtless are closely allied.
Hab. Beulah, 4 9,1 ¢ at flowers of Salix, 2 ? at flowers
of wild plum, May 380, 1899 (W. Porter).
Mesilla Park, New Mexico, U.S.A.,
March 28, 1900.
Il].—On the West-Indian Species of Madrepora*. By
J. W. Grecory, D.Sc., F.G.S., Professor of Geology -in
the University of Melbourne.
Tue term muricatum was first applied to West-Indian corals
* [This paper was read before one of the London Societies in June last
year after a visit to the West Indies to study, amongst other questions,
the Madrepore of that region, The paper was withdrawn by request of
the Society.
I delayed publication in order to reconsider the matter after a few
months’ interval. A recent letter from Mr, J. HK, Duerden, of Jamaica,
West-Indian Species of Madrepora. 21
by Sloane * in 1707. Sloane used the word in describing
the three forms of Madrepora which are now generally known
as M. palmata, M. cervicornis, and M. prolifera. Linneus
accepted the term as the name of a species which he called
Millepora muricata in 1754 and Madrepora muricata in 1767 f.
Linneus founded the species to include all the Madrepore
with an arborescent branching corallum. It was adopted in
the sense of either Sloane or Linneus by subsequent zoologists
until 1816, when Lamarck { broke up the IZ. muricata, L. et
auct., into five species and abandoned Linneeus’s specific name.
For the West-Indian Madrepore he founded the species
M. palmata, M. cervicornis, and M. prolifera. Lamarck’s
course of action was adopted by all students of corals until
1890 ; in that year Prof. Heilprin suggested that the West-
Indian branched and palmate forms of Madrepora are members
of the same species. “I feel doubtful,” says Prof. Heilprin §,
“if the palmate form of the corallum, as seen in IZ. palmata,
M. flabellum, and M. alces (Kast Indies), is in itself a character
sufficient to distinguish the species from those forms, agreeing
with the palmate types in other respects, in which the corallum
is strictly digitate. My associate, Mr. J. E. Ives, has called
my attention to the tendency in the direction of digitation
which many individuals of the palmate species exhibit. This
is carried so far in some of the specimens contained in the col-
lections of the Academy of Natural Sciences that it becomes
difficult, if not really impossible, to class the individuals.”
By the digitate types Prof. Heilprin presumably means
M. cervicornis, for M. prolifera he kept quite distinct. The
same conclusion was reached and extended in 1893 by Brook ||,
who is making a detailed study of the polypes of the West-Indian corals,
shows that, like myself, he had been misled by deference to Brook’s
opinion, It therefore seems to me advisable to publish the paper, and
it is issued exactly as written last June. My. Duerden says :—“ Relying
upon Brook’s statement that he had met with intermediate specimens of
cervicornis and palmata, 1 was inclined to regard them all {including alc7-
formis| asone species. I have examined acres of Madrepora growth with
the object of finding such intermediate forms, but without any success,
although such would be expected considering that prolifera and palmata
grow together.”’} :
* Hans Sloane, ‘A Voyage to the Islands Madera... Jamaica,’ vol. i.
(1707) pp. 51-58, pl. xviii. figs. 3, 4, pl. xvii. figs. 2, 3.
+ Linneus, Syst. Nat. ed. x. p. 792, ed. xii. p. 1279.
t Lamarck, Hist. Nat. Anim. s. Vert. vol. ii. 278, 281.
§ A. Heilprin, “The Corals and Coral-reefs of the Western Waters of
the Gulf of Mexico,” Proc. Acad. Nat. Sci. Phil. 1890, p. 304.
|| Geo. Brook, ‘The Genus Madrepora,’ Cat. Madrep. Brit. Museum,
yol. i. 1893, pp. 25-380.
22 Prof. J. W. Gregory on the
whose opinion was no doubt formed independently, as he does
not refer to Prof. Heilprin’s paper. In Brook’s great mono-
graph of the genus Madrepora he not only merged Lamarck’s
three West-Indian species, but adopted for them Linneus’s
name of muricata.
During a recent visit to the West Indies I have had the
opportunity of studying the three forms of Madrepora on the
reefs, and have been led to revert to the Lamarckian arrange-
ment. As in 1895* I accepted Brook’s proposals, it may be
advisable to state the reasons for my change of opinion.
It will be convenient first to consider whether the West-
Indian Madrepore are all to be included in a single species.
Brook supported this idea by two lines of evidence :—Ist, the
distribution of the corals on the reefs ; 2nd, the existence of a
series of specimens having characters intermediate between
those of Lamarck’s species.
HABIT AND ENVIRONMENT.
The first argument was based on statements that Brook
attributed to Pourtalés. Thus he sayst that Pourtalés has
“hinted that the three species of Lamarck may prove to be
variations of one species, dependent on environment for their
precise habit.” But this is not quite a correct account of
Pourtalés’s opinion. The only reference to Pourtalés which
Brook includes in his synonymy is to the memoir on the
* Deep-sea Corals.” Therein Pourtalés } does hint that possibly
M. cervicornis and M. prolifera may be specifically identical ;
but he makes no suggestion that Jl. palmata should be
united with them. He even comes finally to the conclusion
that MM. cervicornis and M. prolifera may be conveniently
kept apart. The passage referred to is as follows :—‘‘ Some
specimens partake so much of the characters of both this
fi. e. MW. prolifera] and the preceding species [ JZ. cervicornis|
as to shake the belief in their specific difference. Still the
greater number of specimens examined are readily distin-
guished, more perhaps by their habitus than by the more
minute characters of the calicles.”
Pourtalés’s conclusion seems to me sound. Specimens of
M. prolifera and M. cervicornis are distinguishable without
* J. W. Gregory, “ Contributions to the Paleontology and Physical
Geography of the West Indies,” Quart, Journ. Geol. Soc. vol. li. (1895)
p. 282.
t+ Brook, op. cit. p. 18.
} L. F. de Pourtalés, “ Deep-sea Corals,” Hl. Cat. Mus. Comp, Zool.
no. iv. 1871, p. 84.
West-Indian Species of Madrepora. 23
difficulty, though fragments may sometimes be indeterminable.
But the two forms are sufficiently allied for their separation
to be a mere matter of convenience.
The difference between M. palmata and M. cervicornis is,
however, far more definite, and Pourtalés had no hesitation
in keeping them distinct. Brook *, however, unites them ou
the ground that “ Pourtalés has pointed out, with regard to
the West-Indian specimens of palmata, cervicornis, and pro-
lifera, that the proper habit and robustness of each form is
associated -with a different position on the reef. MM. palmata
grows in situations exposed to the force of the sea; J. cervi-
cornis in less exposed localities ; while for its full develop-
ment I/. prolifera appears to require sheltered spots on the
inner side of the reef.’ This passage involves another
unfortunate misrepresentation of Pourtalés, for, according to
that author, the species which ‘requires a rather sheltered
position for its full development ” f 1s MZ. cervicornis, and not
M. prolifera. In respect to the position of growth of the
latter, Pourtalés gives no information ; and, so far as my own
observations go, M. prolifera does not flourish in sheltered
spots inside the reef, but in deeper water than M. cervicornis,
and often outside the main reef. Thus the typical species in
the quiet coves of Parham Sound, Antigua, are M. palmata
and WM. cervicornis. The best specimens of MW. prolifera that
I obtained from Antigua came from the depth of 3 fathoms
from an exposed position on the slopes of Sandy Island.
Pourtalés did say that M/. palmata is characteristic of the
exposed positions on the outside reefs, a statement, however,
which is only true with one important limitation. The
particular form of JZ, palmata known as “‘ the car of Neptune,”
which has a massive corallum formed of thick lamellar ex-
pansions, is no doubt the typical form of Madrepora found in
exposed positions in the West-Indian reefs. The fragile
‘branched coralla of M. cervicornis and M. prolifera would be
shattered if struck by the full force of a breaker ; they accord-
ingly grow in protected situations or at a depth below the
limit of the surf.
It is possible that it was the “‘ Neptune’s car” form of palmata
which Pourtalés had in mind when writing the previously
quoted remark. His statement, so far as my observations go, is
not correct for M. palmata as a whole. ‘The alciform variety
of palmata, which both Pourtalés and Brook include in that
form, grows under identically the same conditions as AZ, cervi-
cornis. For example, I collected specimens of both forms
* Brook, op. cit. p. 28.
+ Pourtalés, op. cit. p. 84.
24 Prof. J. W. Gregory on the
which were growing side by side, not 18 inches apart, on
precisely the same sea-floor, rising to exactly the same level,
and equally exposed to wave and current. The position was
sheltered in the extreme, for it was on the shore of a small
land-locked bay in Parham Sound on the lee side of Bird
Island; and the bay was further protected by a shoal across
its mouth. In an adjacent patch of reef MM. palmata and
M. cervicornis were growing interlocked, but each species was
perfectly distinct. In another bay in deeper water there were
circular patches of MZ. palmata and M. cervicornis, forming
flat-topped tabular masses from 10 to 15 feet in diameter.
They were growing under identical conditions.
The statements therefore that M. palmata and M. cervi-
cornis are dimorphic forms of one species and that they have
acquired different habits owing to their occurrence at different
situations on a reef are not in accordance with their distri-
bution on the coasts of Antigua. Indeed, the fact that where
M. palmata grows in association with MM. cervicornis the
former is represented by a digitate or branched variety is fatal
to the assumption of their specific identity ; for the J. alces of
Dana, and not the J. cervicornis of Lamarck, is the branched
variety of I. palmata.
THE EVIDENCE OF INTERMEDIATE SPECIMENS.
Brook supported his argument by the existence of corals
intermediate between I/. palmata and M. cervicornis. I care-
fully looked out for such in all the reefs I had the chance of
examining, but the search was unsuccessful. Brook stated
that the ‘ intermediate forms occur chiefly in the collection of
the British Museum.” He enumerates them on p. 29 of his
monograph. ‘The specimens are four in number, and, thanks
to the kindness of Prof. Bell, I have had the opportunity of
examining them.
The first specimen was collected by the ‘ Challenger’ expe-
dition at St. Thomas. Its registration number is 86. 12. 9. 274.
The specimen is 200 millim. long, and consists of a central
stem which gives off a series of cylindrical branches. On one
side there is, a little above the base, an imperfectly separated
branch which subdivides into two and shows the proximal
ends of six cylindrical branches. On the other side there are
eighteen branches or branchlets. I fail to see any approach
to M. palmata in this specimen. If the specimen were
palmate we should expect it to be so at the base. But at the
bottom the central stem measures 30 millim. wide and
20 millim. thick. There is nothing palmate in that. The
West-Indian Species of Madrepora. 25
specimen appears to me only a M. cervicornis in which the
branches are numerous and mainly in one plane.
In regard to the three other intermediate forms, my diffi-
culty is to understand why Brook assigned them to the
palmata-cervicornis group. I had the privilege of examining
them in conjunction with Mr. H. M. Bernard, who agreed
as to the improbability of their specific identification. There
is no evidence that the corals came from the West Indies,
and from the characters of the specimens this source seems
unlikely.
The second specimen is 93. 4. 7. 22, and its locality is
unknown. It is apparently the young basal portion of a
corallum, and is, perhaps, too immature for specific identifica-
tion. It is 120 millim. long by 100 millim. wide, and its
surface bears three rows of subconical elevations. On the
middle row one process has grown upwards into a branch
30 millim. wide by 25 millim. thick, rising 55 millim. from
the bottom of the furrow between the rows and rising 35 millim.
above the slit which separates the branch from the adjacent
subconical elevation. In one of the outside rows there is a
lower branch, 45 millim. long by 30 millim. thick by 32 millim.
high. In these characters I fail to see anything to ally the
specimen to either J/. palmata or M. cervicornis. The coral-
lum agrees more nearly with Brook’s description of that of
M. conigera*.
The third specimen is no. 93. 4. 7. 23, and its locality is
also unknown. It is divided almost to the bottom into
branches which if broken into fragments would be indistin-
guishable from those of JZ. cervicornis, as they would be from
several Pacific species. But the corallum is reticular and its
general aspect is not that of cervicornis, much less of palmata.
It appears to me to be more like MV. dbrevicollis t, though I do
not care to venture an attempt at a specific identification of
any Indo-Pacific Madrepore.
The fourth specimen (93. 4. 7. 85, locality unknown) is
labelled in Mr. Brook’s handwriting “ I, muricata?”’ The
note of interrogation seems amply justified, unless that species
be accepted in its original Linnean sense for Indian Ocean
muricated Madrepore. ‘The specimen consists of thick,
flat, basal lobes, whence arise short thick branches, which
divide into a crowded and irregular series of branchlets. If I
had to give the specimen a name I should feel tempted to call
it a short-branched form of the corals which Brook has iden-
tified as MW. Hhrenbergi t.
* Brook, op. cit. p. 34.
+ Brook, op. ct. p. 159, pl. xxvii. figs. A, B. *
i eens & Haime, Hist. nat. Cor. vol. iii, p. 143; Brook, op. cit.
p- .
26 Prof. J. W. Gregory on the
Hence the specimens which Mr. Brook quoted as linking
M. palmata and M. cervicornis do not seem to me to give any
support to the belief in the specific unity of these corals. To
dismiss such differences in the form of the corallum as not
worthy ‘of specific value appears to me inconsistent with
Mr. Brook’s practice in later pages of his monograph. Thus
he founded a species, J. attenuata, for a form which appears
to be based on a series of fragments of slender branches of
M. cervicornis, and he accepted Dana’s M. cyclopea, which
appears to be only an alciform variety of MZ. palmata. In
the case of A/. attenuata it may be objected that the terminal
axial corallites are shorter than in J/. cervicornis; but they
are not shorter than in Agassiz’s * figures of that species,
which show that the character is inconstant. A more serious
inconsistency is that Brook divided his subgenus Conocyathus,
Brk., non d’Orb.f, into four sections, characterized solely by
the form of the corallum. The following are his diagnoses of
those sections :—
A. Corallum corymbose, with or without confluent branches.
If the central branches are long the habit is bushy.
(2.161,
B. Corallum forming a subcomplanate reticulum, with
short twigs on the upper surface. (P. 166.)
C. Corallum cespitose. (P. 166.)
D. Corallum subarborescent or bushy, usually with nume-
rous short proliferations. (P. 169.)
If the difference between a cespitose ¢ corallum and a
bushy corallum is of more than specific value in “ Cono-
cyathus,”” why is the well-marked difference between the
palmate and arborescent coralla of less than specific value in
Eumadrepora ?
* L. Agassiz, “ Report on the Florida Reefs,” Mem. Mus. Comp. Zool.
vol. vii. no. 1, 1880, pl. xviii. figs. 1, 4, & 8,
+ This name was preoccupied for a genus of corals which has living
Australian representatives. Another of Brook’s subgeneric names, Odonto-
cyathus, is preoccupied for a deep-sea coral dredged by the ‘Challenger’
and described by Moseley.
{ The difference that Brook intended to suggest between a czspitose
and a bushy corallum is not very easy to realize, According to Murray’s
new English Dictionary, czespitose means “growing in thick tufts or
clumps.” But Brook places M. Forskali, in which he describes the
“corallum [as] forming dense and much branched clumps,” among the
bushy and not among the czespitose section; and MW. Rousseawi, in which
he describes the corallum as “consisting of tufts,” is also excluded from
the cxespitose section. Both Ogilvie and Worcester’s dictionaries define
ceespitose as ‘ growing in tufts.”
West-Indian Species of Madrepora. 27
THE Species “ wvrroara.”
Hence, in the absence of corals intermediate between
M. palmata and M. cervicornis, and in view of the fact that
the differences in form between them are not due to growth
under different conditions, it appears advisable to return to
Lamarck’s arrangement of the species. That decision raises
the question whether Linnzus’s name ought not to be retained.
But if we follow Brook, and unite the three species, MZ. pal-
mata, M. cervicornis, and M. prolifera, and take the first as
the typical form—for Brook accepts it as forma A—then the
name muricata is both inappropriate and inapplicable. It is
inappropriate, since the name muricata was probably suggested
by Linnzus from the resemblance of the branchlets of many
species to the varices of Murex *. And M. palmata is not
a muricated species in this sense.
But the name is inapplicable, since, although Linnzus
used it to cover all the ramose Madrepore that he knew, he
carefully excluded the palmate variety from J/. muricata.
He excluded it in three ways. In the first place, both in his
own diagnosis and in his additional remarks, he describes the
species as a ramose form—‘ Madrepora ramosa composita ”’ ;
“rami albi’”’; ‘ corallium seepe format pulcherrime ramis suis
corymbum rosaceum.” lLinneus makes no reference to
palmate or alciform varieties. He also quotes from earlier
authors a series of descriptive phrases in which references
to the ramose condition continually recur. In the second
place, Linneeus carefully excluded the palmate form by
omitting reference to the figures of that coral in the list of
literature on his muricata. ‘Thus Sloane figured an excelient
example of the alciform variety ; Linneus accepts Sloane’s
figures of the cervicornis and prolifera types, but not of the
palmata}t. Seba§ also figured all three torms, the prolifera
on pl. evil. fig. 6, the cervicornis on pl. exiv. fig. 1, and a
typical palmata on pl. exii. Linneeus again accepted the two
first, but excluded the last. In the third place, the inclusion
of M. palmata in M. muricata is rendered unsatisfactory by
the geographical evidence. When Linneus founded the
latter species in 1754 he gave as its habitat “ Pelago Asiatico.”
* Muricata, as Prof. Bell has remarked to me, means spiny, with sharp
points. r ;
+ Sloane, ‘ Voyage... . Jamaica,’ vol. i. pl. xvii. fig. 3.
{ Le, he accepts Sloane, dézd. vol. i. pl. xviii. fig. 3, pl. xvii. fig. 4;
but not pl. xvii. fig. 3.
§ Seba, ‘ Loc, Rerum Natur. Thesauri,’ vol. iii. 1758.
28 Prof. J. W. Gregory on the
And M. palmata is typically, if not exclusively, West Indian,
not East Indian.
It is therefore undesirable to take as the type form of
Linneus’s species the one Madrepora known to Linneus,
which he excluded from it. It seems to me advisable to drop
the name muricata altogether, on the ground that Linneus
used that name for all the ramose Madrepores he knew, as
well as for ramose corals which belong to other families,
Thus Linneeus included in muricata the coral figured by Seba
on his pl. exvi. fig. 5, which is not a Madrepera at all. He
included the three corals figured by Morris * as ‘ anomalous
submarine plants”; they are equally anomalous as specimens
of Madrepora muricata as defined by Brook. To take one of
the many corals included by Linneus in J/. muricata would
be an arbitrary proceeding ; but if it is to be done the name
ought to be applied to an Indo-Pacific species, both since
Linneus assigned it to that area and as the best figures he
quotes are those in ‘ Rumphius Herbarium Amboinense’ f.
Not one of the three species J. palmata, M. cervicornis, or
M. prolifera has been recorded from Amboyna.
THe RANGE OF THE WEST-INDIAN Maprerorz.
The argument from the geographical distribution raises the
question as to the range of M. palmata and M. cervicornis.
J refer to this question with reluctance, and only at the strong
suggestion of Prof. Bell.
According to most authorities the three forms or species of
Madrepore tound in the West Indies and the western tropical
Atlantic are contined to that region. According to Mr, Brook
they also occur in the Pacific and Indian Oceans, ranging
from Tahiti to the Red Sea. As Prof. Bell pointed out to
me, the distribution of these forms as accepted by Brook is
very remarkable; for all three forms are very abundant in
the West Indies, and they all occur very widely but very
sparsely distributed in the Western Pacific and Indian Oceans,
The Indo-Pacific specimens referred by Brook to WM. muricata
are eight in number. I examined some of them in 1895, but
did not see any one character by which they could all be
separated from the West-Indies species, though demurring to
the idea that they were all members of one phylogenetic
species. After a more careful examination of the specimens,
the doubts then expressed are strengthened. The specimens
* Morris, ‘Plantarum Hist. Oxon, Univ.’ pt. iii, 1699, sect. 15, pl. x,
fiys. 3, 9, & 10.
+ 1750, pl. Ixxxvi. figs. 1 & 2.
West-Indian Species of Madrepora. 29
in the zoological collection of the Museum appear to me
insufficient to justify the attribution of an Indo-Pacific range
to M. palmata, M. cervicornis, and M. prolifera.
It will be advisable to consider the specimens separately in
the order in which Brook catalogued them.
A. palmata.
1. No. 92. 6. 8. 213. Port Darwin. Saville Kent coll.
This specimen isa fragment showing no signs of the base. It
is a thin flat lamellum, interrupted by lacune ; it is compara-
tively level on one face, but has a series of muricate branchlets
on the other face.
The specimen differs from typical forms of palmata by the
presence of the numerous varices, of the small lacune, and
by the thinness of the lamellum, hese differences are perhaps
unimportant, but the specimen is such a fragment that its
evidence also is unimportant.
2. No. 93.4. 7.24. Singapore. This specimen is a fine
palmate vasiform corallum; the growth is irregular, and
lacune pierce the lamellae. The upper surface is covered with
numerous small flat-topped branchlets, at the end of which is
a deep pit. The walls are dense.
What specific name should be given to this coral I do not
propose to enquire. It is sufficient to point out that it differs
from palmata by the presence of the numerous branchlets on
the upper surface, and that some of its characters necessitate
its transference to a different division of Madrepora from that
to which M. palmata belongs.
Brook divided Medrepora into four divisions. The first divi-
sion he characterized as follows :—“Madrepore with cylindrical
axial corallites, which project to a greater or less extent at the
apex of each division of the corallum; wall usually very porous,
margin plane, exterior more or less distinctly striateor rugose.”
Now in the Singapore specimen each branchlet does not end
in a projecting axial corallite ; on the contrary, the branchlets
are flat-topped and the axial corallites are not exsert. The
specimen must therefore be transferred from the division con-
taining ELumadrepora to the division comprising the two
subgenera Isopora and T'ylopora. It cannot, however, rest in
either of those genera as they were defined by Brook ; for
according to the characters of the corallum it would be an
dsopora, and according to those of the branchlets it would be
a L'ylopora.
30 Prof. J. W. Gregory on the
B. prolifera.
3. No. 46. 7. 30. 8. Wreck Bay, Great Barrier Reef,
N.E. Australia.
This specimen is the only one in the collection which is
catalogued as a Pacific form of prolifera, It is a small
fragment, 2 inches long, and its evidence is insufficient.
Form intermediate between prolifera and cervicornis.
4, No. 93.4.7. 43. Tahiti. (MM. regalis, Ehr.)
Milne-Edwards & Haime * described M. regalis as “ trés-
voisine du WM. prolifera, mais ayant les branches plus grosses.”
That definition accurately describes the habit of this specimen.
The thickness of the branches agrees with that of JZ. cervi-
cornis. ‘The specimen differs from WM. cervicornis by having
very short (1-2 millim. exsert) terminal corallites, in which
the primary septa are very unequal. According to Brook the
axial corallites of cervicornis have the terminal corallite
6-8 millim. exsert and the primary septa subequal.
The corallites differ from those of M. prolifera by having
well-developed septa, whereas, according to Brook, in that
form ‘the directive septa are moderately developed, but the
remaining members of the primary cycle are more or less
rudimentary.”
C. cervicornis.
5-7. Nos. 92.6.8. 210-212. Port Darwin. Saville Kent
call.
These three specimens, though differing somewhat in the
relative closeness of the branches probably belong to the same
species. The habit is cervicorn; but the most striking feature
of the coral is that the terminal axial corallites are broad,
short, and thick-walled ; the wall, in fact, is equal in thick-
ness to the transverse diameter of the calice. In the youngest
corallites there are 6 septa; in older corallites the septa
number from 12-16; in the largest and best-preserved coral-
lites (e. g. in one marked with an ink-dot on specimen
92. 6. 8. 210) there are 3 complete cycles of septa.
Now, according to Brook, in the subgenus Kumadrepora the
axial corallites have a “ relatively thin wall and 12 septa.”
Hence the Port Darwin corals are not typical members of
the same subgenus as JZ, cervicornis.
* Hist. nat. Cor. vol. ii. p. 159.
9
West-Indian Species of Madrepora. al
8. No. 92. 6. 8. 214. Thursday Island. Saville Kent
collection.
This is the last specimen in Brook’s list, and it is that
which most closely resembles M. cervicornis. It agrees with
that form in (1) the radial corallites being nariform below
and tubo-nariform in the distal parts, (2) in the thin walls of
the terminal radial corallite, and (3) in the striate or echinu-
late ornamentation of the walls of the corallites,
These three characters are common to most of the arbo-
rescent Madrepores; the first and third characters occur in
nearly all, as, e. g., in MM. entermedia. The most important
point of resemblance between this specimen and J, cervi-
cornis is the length of the terminal corallite. This structure
is shown, however, only on one branch of the Thursday Island
specimen.
There are not wanting differences between this coral anl
Brook’s description of J. cervicornis. Thus he states that
the primary septa are subequal, whereas in this specimen they
are very unequal, while the one terminal corallite, though of
the same length as in J. cervicornis, is narrower, being
3 willim. instead of 4—5 millim. in diameter.
Why this specimen, with its long narrow terminal corallite,
was regarded as the same species and variety as the Port
Darwin specimens, with their short, broad, thick-walled,
terminal corallites, is not obvious.
Hence I am driven to the conclusion that the evidence of
the eight Madrepores which Brook catalogued as Indo-Pacific
representatives of M. palmata, cervicornis, and prolifera is
insufficient to prove the occurrence of those species in the
Indo-Pacific Ocean.
SUMMARY OF CONCLUSIONS.
1. M. palmata, Lam., may be conveniently kept distinct
from JM. cervicornis, since (a) the two forms live under
identical conditions, their differences are not due to
environment, and (0d) the evidence of the intermediate
forms is inconclusive.
2. M. palmata, Lam., should not be treated as the typical
form of MJ. muricata, L., from which Linneus ex-
cluded it.
3. If the name JZ. muricata be retained, which seems un-
desirable, it should be used for an Indo-Pacific species.
4, The evidence of the range of M. palmata, M. cervicornis,
and J. prolifera into the Indo-Pacific is inadequate.
32 Canon A. M. Norman on British Amphipoda.
IV.—British Amphipoda.—IV. Families Stegocephalide to
CHdiceride (part.). By Canon Norman, M.A., D.C.L.,
LED:, F:R-S.,.ce*
[Plate IIL.]
Fam. VI. Stegocephalide.
Genus 1. STEGOCEPHALOIDES, G. O. Sars.
81. Stegocephaloides christianiensis, Boeck.
1869. Stegocephalus ampulla, Norman, “ Last Report Dredging Shet-
land Isles,” Brit. Assoc. Rep. for 1868, p. 276 (nec Cancer ampulla,
Phipps).
1870. iia christianiensis, Boeck, (137) p. 48.
1876. Steyocephalus christianiensis, Boeck, (188) p. 424, pl. vill. fig. 4,
pl. ix. fig. 1.
1892. Stegocephaloides christianiensis, G. O. Sars, (142) p. 202, pl. Ixx.
fio. 2.
Hab. St. Magnus Bay, Shetland, 50 fathoms, 1867
(A. M. N.); ‘Porcupine,’ 1869, Stat. 18, to.the west of
Galway, 183 fathoms: Mus. Nor. Loch Fyne, 40-70 fathoms
(D. R.) ; S.W. of Ireland, 750 fathoms, 1888, and west coast
of Ireland off the Skilligs, 52-62 fathoms, 1890 (specimens
in Dublin Museum, fide A. O. W.).
Distrib. Trondhjem Fiord, Norway, 40-300 fathoms
(A. M. N.); West Norway (G. 0. Sars): Mus. Nor. Sars
states that it is found as far north as the Lofoten Islands.
Bohusliin (Malm) ; Skagarak (Metnert); Belle Ile, France
(Chevreus).
[Genus 2. STEGOCEPHALUS, Kroyer.
[ Stegocephalus inflatus, Kroyer.
1842, Stegocephalus inflatus, Kroyer, Naturhist. Tidsskr. vol. iv. p. 150;
Voyage en Scandinavie &c. pl. xx. fig. 2.
1876. Stegocephalus ampulla, Boeck, (188) p. 421 (but not Cancer
ampulla, Phipps).
1892. Stegocephalus inflatus, G. O. Sars, (142) pl. Lxix.
Off Holsteinborg, Greenland, in 57 fathoms, ‘ Valorous’
Exped., 1875, Stat. 5.
Trondhjem Fiord, 200-300 fathoms ; Hardanger Fiord,
between the islands Valoddin and Hidle, 110 fathoms (in both
these cases I found it among Lophohelia prolifera) ; also
* See for preceding paper ‘ Annals,’ vol. v. 1900, p, 326.
Canon A. M. Norman on British Amphipoda, 33
Klosterelv Fiord, Kast Finmark (A, M@. N.) ; West Norway
(G. O. Sars) ; ‘ Triton,’ 1882, Faroe Channel, lat. 60° 9! N.,
long. 7° 16° W., in 466 fathoms; Spetsbergen (Lovén) ;
mera’ Eixped., lat.) 67° 7’ N’, long. 173° 24" W. (i.e.
Behring’s Strait), 24 fathoms (from Stockholm Mus.) ; ‘ Wil-
lem-Barents,’ Barents Sea (Stebbing) ; U.S. Fish. Comm.,
Gulf of Maine, 51 fathoms (8S. Z. Smith): Mus. Nor. It
will thus be seen that this is a true Arctic species with
circumpolar distribution. ]
Genus 3. ANDANIA, Boeck,
82. Andania abyssi, Boeck.
1870. Andania abysst, Boeck, (137) p. 49.
1876, Andania abyssi, Boeck, (138) p. 426, pl. ix. fig. 2.
1892. Andania abyssr, G. O. Sars, (142) p. 207, pl. lxxi, fig. 2, and
pl. lxxii. fig. 1.
Hab. ‘ Porcupine,’ 1869, Stat. 24, south of Rockall, lat.
56° 26! N., long. 14° 28’ W., 109 fathoms: Mus. Nor.
Distrib. ‘Trondhjem Fiord, 200-300 fathoms (A. MZ. N.) ;
West Norway (G. O. Sars): Mus. Nor. Sars speaks of it
as a true deep-water form, only occurring in greater depths
from 200-400 fathoms; it has occurred from Southern Norway
northwards to the Lofoten Islands.
Fam. VII. Amphilochide.
Genus 1. AMPHILOCHUS, Bate.
83. Amphilochus manudens, Bate.
1862. Amphilochus manudens, Bate & Westwood, (1) vol. i. p. 180, ¢.
1876. Amphilochus concinnus, Stebbing, ‘ Some new and little-known
Amphip. Crust.,” Ann. & Mag. Nat. Hist. ser. iv. vol. xviii. p. 443,
pl. xix. figs. 1 a,b, 9.
1876. Callimerus acutidigitata, id. ibid. p. 445, pl. xx. figs. 3 a, b.
1890. Amphilochus Boeckii, Meinert, (71) p, 160.
1892. Amphilochus manudens, G, O. Sars, (142) p. 217, pl. lxxiv,
Flab. Off Farland Point, Cumbrae, 20 fathoms (A. JZ, N.) :
Mus. Nor. Mull (G. Brook, fide T. S.) ; Upper Loch Fyne
and Granton Harbour, Firth of Clyde (7. S.); St. Andrews
(M‘Intosh) ; off North Wales and Isle of Man (A, O. W.) ;
‘Torbay (Stebbing) ; Jersey (Sinel & Hornell).
Distrib, Drobak, Christiania Fiord, and Trondhjem Fiord,
200 fathoms (A. M. N.) : Mus. Nor. On Norwegian and
Finmarkian coasts, not rare from Christiania to Vadsé (G4. 0.
Sars) ; Greenland (/ansen) ; Kattegat (Mednert) ; west coast
of France (Chevreuz).
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 3
34 Canon A. M. Norman on British Amphipoda.
84. Amphilochus tenuimanus, Boeck.
1870, Amphilochus tenuimanus, Boeck, (187) p. 51.
1876. Amphilochus tenuimanus, Boeck, (138) p. 487, pl. ix. fig. 7.
1892. Amphilochus tenuimanus, G. O. Sars, (142) p. 218, pl. Ixxv. fig. 1.
Hab., Cumbrae (DD. #.); Firth of Clyde (728) 5 0m
Spurm Head, Yorkshire (7. S.).
Distrib. Sars says that it is a deep-water form living in
100-200 fathoms, especially among deep-sea corals, and that
it is found in several places and West Norway.
85. Amphilochus neapolitanus, Della Valle. (PI. III. fig. 1.)
1893. Te amie neapolitanus, Della Valle, (139) p. 595, pl. xxix
figs. 16, 17.
1895. Amphilochus melanops, A. O. Walker, “ Revision Amphipoda of
Liv. M. B. C. District,” Trans. Liv. Biol. Assoc. vol. ix. p. 298,
pl. xviii. fig. 12, pl. xix. figs. 13-15.
Hab. Off Little Ormes Head, Denbighshire, 5-7 fathoms
(A. O. W.); Falmouth Harbour; Jersey, tide-marks
(A. M.N.): Mus. Nor. Also Menai Strait, near Beaumaris,
5-10 fathoms (A. O. W.).
Distrib. Naples (Della Valle): Mus. Nor. West France
(Chevreuz).
Having compared typical specimens of A. melanops kindly
given me by Mr. Walker, and other British examples taken
by myself in the localities mentioned above, with a specimen
ot A. neapolitanus received from Della Valle, I am satisfied
as to their identity. A. neapolitanus and A. brunneus, both
described by Della Valle, are very closely allied. OF the
latter I took several specimens at Naples in 1887, and thus
have had the opportunity of comparing the two forms. The
one point in which they are described by Della Valle as
differing consists in the character of the gnathopods: those
of the second pair are very broad, triangular, and the carpal
process reaches to the extremity of the posterior margin of the
hand; the armature of the palm and dactylus is exactly as
represented by Della Valle in the allied species A. brunneus
(Della Valle, pl. xxix. fig. 14), the palm being bounded by
two spines and its edge excessively finely denticulated, and
within the denticulations a row of small spinules.
Mr. Walker’s fig. 14 gives the carpal process shorter than
in any specimen I have seen; but in A. brunneus it only
extends about half the length of the hand. ‘lhe very close
resemblance of the species might raise a question in the mind
whether they are really distinct. In favour of their distine-
tion is the fact that all the specimens I took at Naples agreed
Canon A. M. Norman on British Amphipoda. 35
with A. drunneus and all the British examples which I have
examined agree with A. neapolitanus. Of this last species
Della Valle wrote that he had only seen a single specimen ;
but he must have obtained others subsequently, since imme-
diately after the publication of his work he kindly sent mean
example. A. tenuimanus, G. O. Sars, is distinguished from
the present species by the character of the gnathopods, and
especially by the telson exceeding in length the peduncle of
the last uropods, whereas in A. neapolitanus the telson is
much shorter and scarcely equals half the length of the
peduncle of the much produced last uropods.
Genus 2. AMPHILOCHOIDES, G. O. Sars.
86. Amphilochoides serratipes (Norman).
1869. Probolium serratipes, Norman, “ Last Report Dredging Shet-
land Isles,” Brit. Assoc. Rep. for 1868, p. 273.
1892. Amphilochvides odontonyx, G. O. Sars, (142) p. 221, pl. Ixxv.
fig. 2 (nec A. odontonyx, Boeck).
1895. Amphilochoides Boeckit, G. O. Sars, (142) p. 690.
Hab. St. Magnus Bay, Shetland, in 50 fathoms; Isle of
Cumbrae, 5 fathoms (A. JZ. N.): Mus. Nor. Blackwater-
foot, Arran, N.B., 20 fathoms (D. &.); 8 miles off Flesh-
wick Bay, Isle of Man, 33 fathoms (A. O. W.).
Distrib. West Norway (G. O. Sars): Mus. Nor. On the
Norwegian coast here and there from Christiania to the
Trondhjem Fiord, in 50-100 fathoms (G@. O. Sars); Con-
carneau (Chevreux).
87. Amphilochoides odontonyx, Boeck.
1870. Amphilochus odontonyx, Boeck, (137) p. 51.
1876. Amphilochus odontonyx, Boeck, (158) p. 434, pl. xi. fig. 3.
1892. Amphilochordes pusillus, G. O. Sars, (142) p. 222, pl. lxxvi. fig. 1.
1895. Amphilochoides odontonyx, id. ibid. p. 690.
1896. Amphilochoides odontonyx, T. Scott, Fourteenth Annual Rep.
Scotch Fish. Board, p. 159, pl. iv. figs. 4-6.
Ilab. Plymouth, 8 fathoms, 1887 (A. M. N.): Mus. Nor.
Near Bass Rock, Firth of Forth, 22-23 fathoms; St. Andrews
Bay ; off Spurm Head, Yorkshire; several places in the
Clyde district (7. S.).
Distrib. Christiania Fiord, 20-40 fathoms, and Vadsd,
E. Finmark, 30-60 fathoms (G. O. Sars); Denmark
(Mevnert).
OF
36 Canon A. M. Norman on British Amphipoda.
88. Amphilochoides intermedius, 'T. Scott.
1896. Amphilochoides intermedius, T. Scott, Fourteenth Annual Report
Scotch Fish. Board, p. 159, pl. iv. figs, 1-3.
Hab. Various parts of the Firth of Forth (7. S.).
Genus 3. Giranopsis, G. O. Sars.
89. Gitanopsis bispinosa (Boeck).
1870. Amphilochus bispinosus, Boeck, (137) p. 51.
1876, Amphilochus bispinosus, Boeck, (188) p. 435, pl. x. fig. 1.
1892. Gitanopsis bispinosa, G. O. Sars, (142) p. 224, pl. Ixxvi. fig. 2.
flab. “ Dredged off Blackwater-foot, Arran, N.B., in
20 fathoms, bottom sand and mud” (D. &.).
Distrib. Lofoten Islands, Norway (G. O. Sars): Mus.
Nor. South and West Norway as far north as Lofoten
Islands, but nowhere in abundance, generally in 50-100
fathoms (G. O. Sars) ; Greenland (Hansen); west coast of
France (Chevreuz).
90. Gitanopsis inermis, G. O. Sars.
1882. Gitanopsis inermis, G. O. Sars, (102) p. 51.
1892. Gitanopsis inermis, G. O. Sars, (142) p. 224, pl. Ixxvi. fig. 2.
Hab. Specimens taken by Professor G. 8. Brady * off
Cullercoats, Northumberland, were identified by Mr. Stebbing
as this species (Nat. Hist. Trans. Northumb., Durham, and
Newcastle-upon-Tyne, vol. xiii. pt. 3, 1900, p. 442).
Distrib. Professor Sars has met with this species only in a
single locality, and that a very northern one—Vads6—where
it was found rather sparingly in 20-50 fathoms.
Genus 4. GiTana, Boeck.
91. Gitana Sarsii, Boeck.
1870. Gitana Sarst, Boeck, (137) p. 52.
1876. Gitana Sarst, Boeck, (188) p. 439, pl. xi. fig. 2.
1878. Amphilochus Sabrine, Stebbing, ‘Two new Species of Amphi-
podous Crustacea,” Ann. & Mag. Nat. Hist. ser. 5, vol. ii. p. 365,
pl. xv. figs. 1 a-g.
1892. Gitana Sarsit, G. O. Sars, (142) p. 228, pl. Ixxviii. fig. 1.
1893. Gitana Sarsii, Della Valle, (159) p. 590, pl. xxix. figs. 18-32.
* It may be desirable to mention that Professor Brady, who has done
so much good work among the Entomostraca, does not study the Amphi-
poda, and that when his name is added to a locality, if it be this present
locality the specimens have been determined by Mr. Stebbing, and from
all other localities they have been identified by myself.
Canon A. M. Norman on British Amphipoda, 37
Hab. Off Little Orme, Denbighshire, 2-7 fathoms
(A. O. W.): Mus. Nor. Tenby (Stebding); Kilchattan
Bay, Firth of Clyde, 5 fathoms, muddy sand (D. &.); Inch-
keith, Firth of Forth (7. S.) ; 8 miles off Fleshwick Bay,
Isle of Man, 33 fathoms, and Valentia, Ireland (4. O. W.).
Distrib. West Norway (G. O. Sars); Baie de Houat,
France (Chevreux): Mus. Nor. Among alge in shallow
water all along coasts of Norway and Finmark (G. O. Sars) ;
Spetsbergen (Norw. North Atlant. Exped.); Franz-Josef
Land, Jackson-Harmsworth Exped. (7. 8S.) ; Kattegat
(Meinert); Naples (Della Valle).
Genus 5. Cyproip1A (Haswell ?), Stebbing.
=? Peltocova, Catta.
It is at this time impossible to say in what genus the
Cyproidia damnoniensis, Stebbing, should be located, and I
think it best therefore to leave it where he placed it. As
both Della Valle and myself, however, have found this
species in the Mediterranean, probability is given to the
assumption of Della Valle that Cyproidia is a synonym of
Peltocoxa, Catta, described in 1875* from specimens taken
at Marseilles, but that author’s description of the genus is so
inadequate that it requires much to be assumed to employ it.
Haswell, in his amended description of Cyprotdia (‘ Cat.
Australian Stalk- and Sessile-eyed Crustacea,’ Sydney, 1882),
states that the “ antenne are subequal, superior without an
appendage.” If that is really so, Stegoplax of G. O. Sars is
asynonym. But Stebbing appears to have concluded that
the secondary appendage, which is extremely minute, had
been overlooked by Haswell, and consequently placed in
Haswell’s genus his Cyproidia damnoniensis, which has such
an appendage.
Haswell put two species in his genus, Cyproidia lineata
and C. ornata. Quite recently Mr. Stebbing +, having
received specimens of the first of these species from
Mr. Haswell, has instituted, apparently on very slight
grounds, another genus (Paracyprovdia) for its reception, but
gives us no information as to the antenne either of his new
genus or of Cyproidia ornata, which he leaves as the type of
Haswell’s Cyprotdia.
* Peltocova Marioni, Catta, Revue des Sci. Nat. vol. iv. 1875, p. 161.
+ Stebbing, “ Revision of Amphipoda,” Ann, & Mag. Nat. Hist, ser. 7,
vol. iv. 1899, p. 207.
38 Canon A. M. Norman on British Amphipoda.
Still more recently Dr. Charles Chilton * has described a
species from New Zealand under the name Cyproidia otakensis,
which accords in all generic characters with C’. damnoniensis,
Stebbing.
Two other species have been characterized under the name
Cyproidia: one from New Zealand, doubtfully referred by its
author to Cyprotdia, and since affirmed by him to be quite
distinct ; for this Cyprotdia? crassa, Chilton t, Mr. Stebbing f
has instituted a genus Tetradeion.
The other, Cyproidia brevirostris of T. & A. Scott I here
place in the genus Peltocova, G. O. Sars, because the upper
antenne have no secondary appendage.
92. Cyproidia damnoniensis, Stebbing.
1885. Cyprotdia damnoniensis, Stebbing, “ Descr. of a new English
Amphipodous Crustacean,” Ann. & Mag. Nat. Hist. ser. 5, vol. xv.
p- 59, pl. ii.
1893. Peltocoxa damnoniensis, Della Valle, (139) p. 648, pl. xxx.
figs. 19-382, pl. lx. figs. 9, 10.
Hab, Starcross, Devon (C. W. Parker): Mus. Nor,
Blackwater-foot, Arran, N.B., 20 fathoms, muddy gravel,
and off Fairland Point, Cumbrae, 19 fathoms, gravel (V. &.).
Distrib. Naples (Della Valle & A.M. N.): Mus. Nor. Le
Croisic, France (Chevreux).
Genus 6. SteGOPLAX, G. O. Sars.
93. Stegoplax brevirostris (T. & A. Scott).
1893. Cyproidia brevirostris, T. & A. Scott, “ On some new and rare
Crustacea from Scotland,” Ann, & Mag. Nat. Hist. ser. 6, vol. xii.
p. 244, pl. xiii. figs. 1-11.
Hab. Loch Fyne (7. S.): Mus. Nor. Moray Firth,
washed from Filograna implexa from a depth over 40 fathoms
(7. S.), 8 miles west of Fleshwick Bay, Isle of Man, in
33 fathoms, and Valentia, Ireland (4. O. W.).
* Chilton, “A New-Zealand Species of the Amphipodous Genus
Cyproidia,” Ann. & Mag. Nat. Hist. ser. 7, vol. v. 1900, p. 242.
+ Trans. New Zealand Institute, vol. xv. 1882, p. 80, pl. iii. fig, 1.
¢{ Ann. & Mag. Nat. Hist. ser. 7, vol. iv. 1899, p. 207.
Canon A. M. Norman on British Amphipoda. 3y
Fam. VIII. Stenothoide.
Genus 1. STENOTHOE, Dana.
94. Stenothoe marina (Bate).
1860, Stenothoe Danai, A. Boeck, Férh. ved de Skand. Naturf. 8de
Mide, p. 655.
1861. Montagua marina, Bate & Westwood, (1) vol. i. p. 58,
1892, Stenothoe marina, G. O Sars, (142) p. 236, pl. Ixxx,
Hab. Shetland; Skye; Isle of Cumbrae;~ Cullercoats,
Northumberland ; Durham coast ; Plymouth; Donegal Bay,
Ireland (A. M.N.); ‘ Porcupine,’ 1869, Stat. 6, off S.W. Ire-
land, 90 fathoms; Firth of Forth (7. 8.) : Mus. Nor. Moray
Firth and off Spurm Head (7. 8.) ; St. Andrews (Ad‘Intosh) ;
Loch Fyne (Murray) ; North Wales ; Isle of Man; near
Bray Head, W. lveland (A. O. W.); Jersey and Sark
(Kehler).
95, Stenothoe crassicornis, A. O. Walker.
1897. Stenothoe crassicornis, Walker, Rep. Brit. Assoc. for 1896, p. 420.
1897. Sienothoe crassicornis, Walker, “ New Species of Edriophthalma
6
from the Irish Sea,” Journ. Linn. Soc., Zool. vol. xxvi. p. 229,
pl. xviii. figs. 3-3 e.
Hab. “Three males taken in the bottom tow-net 6 miles
W.S.W. of Calf of Man, in 23 fathoms” (A. O. W.).
96. Stenothoe monoculoides (Montagu).
1861. Montayua monoculoides, Bate & Westwood, (1) vol. i. p. 54.
1892, Stenothoe monoculoides, G. O. Sars, (142) p. 240, pl. lxxxii. fig. 1.
Hab. Apparently to be found all round our coasts. Shet-
land; Firth of Clyde ; Plymouth ; Falmouth ; Guernsey ;
Jersey ; Roundstone, Ireland (A. MW. WV.) ; Cullercoats, North-
umberland (J. Alder) ; Starcross, Devon (C. Parker) : Mus.
Nor. Farne Islands, Northumberland (A. M. N.) ; Ardbear
Bay, Ireland (Brady & Robertson) ; North Wales and Isle of
Man (A. O. W.); St. Andrews (M‘Intosh) ; Firth of Forth
(7. 8.) ; Loch Fyne (Murray) ; Valentia, Ireland (A. O. W.).
Distrib. South and West Norway, as far north as Trond-
hjem Fiord (@. 0. Sars); Denmark (Meinert) ; West France
(Chevreux) ; Azores (Barrois).
97. Stenothoe setosa, sp.n. (Pl. ILI. figs. 2-4.)
Coxal plates of second pereopods resembling those of
S. marinain form. Superior antenne with basal joint longer
40 Canon A. M. Norman on British Amphipoda.
than the second; flagellum of twenty-four articulations.
Lower antennz with the last joint of the peduncle one fourth
shorter than the penultimate ; flagellum of twenty-one articu-
lations. First gnathopods (fig. 2) having the lobe of the
meros well rounded at the extremity; hand slightly longer
than the wrist, widening distally ; palm defined, occupying
the slightly oblique extremity. The hand of the second
gnathopods (fig. 3) elongated ovate, widest in the middle,
front margin evenly and regularly arcuate, fringed throughout
with long sete, the length of which almost equals the breadth
of the hand, wholly without spines, tubercles, or crenation,
and the palm undefined ; finger broad, extending to only half
the length of the hand. The last pereeopods (fig. 4) have the
meral and propodal joints subequal in length, the carpal
joint somewhat shorter, the nail half as long as the propodos ;
the posterior lobe of the meros is well arched, but not greatly
developed, extending to half the length of the carpus.
Colour white, tinted with rose-colour; eye red.
Hab. A single specimen taken at Plymouth, August 1889
(A. UM. N.): Mus. Nor.
My dissection does not show the mandible, but from the
general character of the species it would seem to belong to
this genus. 4S. setosa has its nearest known ally apparently
in S. Dollfust, Chevreux *. It resembles that species in the
form of the first gnathopods and, judging from the drawing,
in the proportionate length of the joints of the last pereeopods ;
but it differs in the form of the second gnathopods and the
absence of all teething of the palm of its propodos.
Genus 2. Mreropa, Boeck.
98. Metopa Alderi (Bate).
1861. Montagua Alderi, Bate & Westwood, (1) vol. i. p.
p. 61 22
1868. Metopa norvegica, Bate & Westwood (nec Lilljeborg), (1) vol. ii.
p. 500, ¢.
1876. Metopa clypeata, var., Boeck, (138) p. 451, pl. xviii. fig. 5, ¢.
1892. Metopa Alderi, G. O. Sars, (142) p. 250, pl. xxxvi.
Hab. Aberdeenshire (R. Dawson): Mus, Nor. Northum-
berland coast (A.M. N.) ; Firth of Forth (7. S.); Firth of
Clyde and Mull of Kintyre (D. &.); North Wales (A. O.W.) ;
Torbay (Stebbing).
Distrib. Norway, from the south northwards to Tromsé
(G. O. Sars); Spetsbergen and Sweden (Goés) ; Murman
* Chevreux, “ Quatriéme Campagne de ‘1’Hirondelle,’ 1888: Hyale
Grimaldi et Stenothoe Dollfusi,” Bull. de Soc. Zool. de France, 1891.
Canon A. M. Norman on British Amphipoda. Al
coast (Stuxberg); Iceland (Sars); Denmark (Mernert) ;
Holland (Hoek); Boulogne (J. Bonnier); La Manche
(Chevreux).
99. Metopa norvegica (Lilljeborg).
1850. Leucothoe norvegica, Lilljeborg, “ Bidr. till Norra Rysslands och
Norrige fauna &c.,” K. Vet.-Akad. Hand. vol. ii. p. 335, pl. xx. fig. 4.
1855. Montagua pollexiana, Bate, Brit. Assoc. Rep. p. 57.
1861. Montagua pollexiana, Bate & Westwood, (1) vol. i. p. 64.
1887. Metopa pollexiana, H. J. Hansen, (141) p. 92, pl. iii. figs. 5, 5 a.
1892. Metopa pollexiana, G. O. Sars, (142) p. 269, pl. exv.
The Metopa which Bate and Westwood figured as Leuco-
thoe norvegica, Lilljeborg, was the male of MW. Alder’, and to
that species Sars has referred Lilljeborg’s species; but from
his reference to Bate and Westwood’s figure it would almost
seem that he had not consulted Lilljeborg’s papers. That
author’s Leucothoe norvegica is, it appears to me, undoubtedly
the present species. The following description applies to it,
and not to M. Alderi :—“ L. clypeate, Kroyer, sat affinis.
Antenne superiores inferioribus longiores, flagello pedunculo
longiore, articulo primo secundum superante, et articulo tertio
minimo ; flagellum antenni inferi ultimo pedunculi articulo
brevius vel equale; manus pedum secundi paris maxima,
dilatate, apice vero acuminato, aculeoque marginis posterioris
medii validissimo et ungut cequali ; epimera quarti annuli
thoracici maxima, latitudine vero altitudine parum majore.”
The italics are Lilljeborg’s, but for comparison with M. Alder
the statement as to comparative lengths of the antenne should
also be italicized. ‘The figure represents a second gnathopod
with the palm projected forwards and the lateral process
reaching beyond the palm (cf Sars’s figure p.’ 3), so that
the finger impinges upon it; and in no other species is this
the case.
Hab. Shetland; near Holy Island, Northumberland;
Durham coast (A. M. N.); Aberdeen (&. Dawson): Mus.
Nor. St. Andrews (M‘Intosh); Firth of Forth (7. &.).
There is thus no record of it as yet from the west side of our
islands except St. Ives, whence Spence Bate received
specimens from Mr. George Barlee.
Distrib. Bejan, at entrance of ‘Trondhjem Fiord, Norway,
in 80-50 fathoms (G. O. Sars) ; Tromsé (Schneider) ; Green-
land (H. J. Hansen) ; Murman coast (Jarzinski, fide Sars).
100. Metopa rubrovittata, G. O. Sars. (Pl. III. fig. 5.)
21876, Probolkiwm Spence-Batei, Stebbing, Ann. & Mag. Nat. Hist.
ser. 4, vol. xvii. p. 344, pl. xix, fig. 4,
42. Canon A. M. Norman on British Amphipoda.
1882, Metopa rubrovittata, G. O. Sars, (102) p. 90, pl. iv. figs. 2, 2 a.
1892. Metopa rubrovittata, G. O. Sars, (142) p. 255, pl. Ixxxix, fig. 2.
Hab. Cullercoats, Northumberland, Oct. 5, 1854. Fresh
specimens, mounted in Dean’s medium at that date, have now
their beautiful crimson markings perfectly preserved, exactly
corresponding to Sars’s coloured drawing. Isle of Cumbrae ;
Lerwick Bay, Shetland (A. 4. N.): Mus. Nor. Off north
side of Little Cumbrae, 10-15 fathoms (D. 2.) ; North Wales
(AL OAWS.
Distrib. Christiansund, Norway, and Vadsé, East Finmark
(G. O. Sars) ; Kattegat (Meinert) ; Holland (Hoek); West
France (Chevreuz).
101. Metopa abscisa*, sp.n. (Pl. III. figs. 6-10.)
1869. Montayua clypeata, Bate & Westwood, (1) vol. ii. p. 499 (nec
Leucothoe clypeata, Kroyer).
Upper antennee with two first joints of peduncle subequal,
third not quite half length of second; flagellum about two
and a half times as long as the peduncle, consisting of about
17 articulations. Lower antenne shorter than the upper, its
flagellum shorter than the peduncle ; last joint of peduncle
slightly longer than the penultimate.
First gnathopods (figs. 6 and 7) with the hand a little
shorter than the carpus, oblong, upper and lower margins
parallel for the first two thirds of the length, from which
point the hinder margin curves upwards to the base of the
nail; nail short, not one third the length of the hand; hand
with a few sete but no spines.
Second gnathopods (figs. 8 and 9) with the hand oblong,
slightly widening distally, breadth equalling from one half to
three fourths of the length ; palm absolutely transverse, some-
what shorter than the margin; not toothed and scarcely
waved, bounded by a tooth-formed process, near to which the
palmar margin is extremely minutely serrated (but the serra-
tions are so minute that they are only visible under a high
magnifying-power) ; finger very stout and strong.
Last pereeopods (fig. 10) with the meros and carpus sub-
equal in length, the hinder lobe of the meral joint small, not
reaching beyond half the length of the carpus; propodos
much longer than the preceding joints; nail as long as the
carpus, the tip turned inwards, with a little seta at the base of
the tip.
Telson devoid of dorsal spines.
* Abscisa, “cut off,” in reference to the truncate extremity of the
hand of the second gnathopods.
Canon A. M. Norman on British Amphipoda. 43
Hab. The specimens I have seen of this species are all
from the eastern side of our island. That figured is one of
those from Cullercoats, Northumberland, which Bate and
Westwood recorded under the name Montagua clypeata.
Other specimens in my collection are from St, Andrews
(Professor M‘Intosh) and Aberdeen (Mr, R. Dawson).
I have thought it best to figure a Cullercoats specimen,
since it was one of these which was recorded by Bate and
Westwood. My St. Andrews example is, however, more
mature, and, agreeing in all other respects, differs somewhat
in the form of the hand of the second gnathopod, which is
somewhat longer in proportion to the breadth, nearly twice
as long as the greatest width; the palm &c. is as in the
Cullercoats example.
This species is nearest allied to M. rubrovittata, from which
it may be easily distinguished by the totally different form of
the first gnathopod (fig. 5, first gnathopod of M. rubrovittata),
by the absolutely transverse palm of the second gnathopods,
and by differences in the last pereeopods.
My friend the late Dr. D. Robertson, in the appendix to
his first report (p. 91, separate copy), recorded “ Mefopa
celypeata” from Cumbrae. It is probable that the Amphipod
found by him was Metopa rubrovittata, since I have on two
occasions, in 1854 and in 1885, taken that species at Cumbrae.
Dr. Hoek has described from Holland a Metopa nearly
allied to M. rubrovittata under the name Metopa Normani
(Hoek, ‘‘ Crustacea Neerlandica, II.,”’ Tijdsch. der Nederl.
Dierkundige Vereen. ser. 2, vol. ii. p. 21 (separate copy),
pl. vil. figs. 5 and 5’); but it appears to be more closely
allied to M. rubrovittata than the species which I have here
described.
102. Metopa borealis, G. O. Sars.
1882. Metopa borealis, G, O. Sars, (102) p. 91, pl. iv. fig. 4 (but not
fio. 4 a).
1899, ae borealis, G. O. Sars, (142) p, 254, pl. Ixxxix. fig. 1.
Hab. Isle of Cumbrae, 1885 (A. M. N.); South Bay,
Firth of Forth, 1888 (7. S.); Menai Strait (A. O. W.):
Mus. Nor. Guernsey (A. O. W.).
Iiistrib. Tromsé (J. 8. Schneider): Mus. Nor. Norway
(G. O. Sars) ; Greenland (Hansen).
103. Metopa propinqua, G. O. Sars.
1892, Metopa propinqua, G. O. Sars, (142) p. 264, pl. xciii. fig. 1.
Hab. The only record of this species occurriug in our fauna
44 Canon A. M. Norman on British Amphipoda.
is one by Mr. T. Scott, who found it off Crail in the Firth of
Forth in 1892 (Twelfth Ann. Rep. Fishery Board of Scot-
land, 1894, p. 263).
Distrib. The only specimens known to Sars were taken by
him in about 40 fathoms in company with Cressa dubia in
two places in the Trondhjem Fiord.
104, Metopa pusilla, G. O. Sars.
1892. Metopa pusilla, G. O. Sars, (142) p. 256, pl. xe. fig. 1.
Hab. Off Fairland Point, Isle of Cumbrae, 20-24 fathoms
(A. M. N.); Menai Strait, 7-12 fathoms (A. O. W.): Mus.
Nor. Rhos Bay, on north coast of Wales, just below tide-
marks (A. O. W.).
Distrib. South and West Norway as far north as the
Trondhjem Fiord, in comparatively shallow water among
Alge and Hydroids (G@. O. Sars); Franz-Josef Land, Jack-
son-Harmsworth Expedition (Thomas Scott, Journ. Linn.
Soc., Zool. vol. xxvii. 1899, p. 72).
105. Metopa tenuimana, G. O. Sars.
1892, Metopa tenuimana, G. O. Sars, (142) p. 259, pl. xci. fig. 1.
Hab. A single specimen taken at Shetland in 1867
(A. M. N.): Mus. Nor.
Distrib. West Norway (A. M. N.): Mus. Nor. Where
Sars’s type specimens were also found.
106. Metopa Bruzelit, Goés.
1865. Montagua Bruzelii, “Crust. Ampbip. Maris Spetsberg. &c.,”
Cefy. af K. Vet.-Akad. Forhand. (p. 522, fide Sars), p. 6 separate
copy, pl. xxxviil. fig. 10.
1887. Metopa Bruzelu, H. J. Hansen, (141) partim (?), p. 97, pl. iv.
figs. 2 c-g (? figs. 2 a, 6).
1892. Metopa Bruzelii, G. O. Sars, (142) p. 261, pl. exii. fig. 1.
1900. Proboloides Bruzelii, Stebbing, Ann, & Mag. Nat. Hist. ser. 7,
vol. v. p. 15.
(Nec Metopa Bruzelit, Boeck, = M. Boeckii, G. O. Sars.)
Hab. Port Erin, Isle of Man, 24 fathoms (A. O. W.):
Mus. Nor. Colwyn Bay and Little Orme, North Wales
(A. O. W.); Firth of Forth (T. S.).
Distrib. Widely distributed in West Norway and Fin-
mark (G. O. Sars); Spetsbergen (Goés) ; Greenland
(Hansen).
Canon A. M. Norman on British Amphipoda. 45
Genus 3. METOPELLA, G. O. Sars.
First gnathopods not subchelate. Basos of last three
pereopods not expanded behind and almost or quite linear in
form, and the meros only very slightly produced behind.
107. Metopella nasuta, Boeck.
1870. Metopa nasuta, Boeck, (137) p. 65.
1876, Metopa nasuta, Boeck, (138) p. 465, pl. xviii. fig. 6.
1892. Metopa nasuta, Sars, (142) p. 276, pl. xeviii. fig. 1.
Hab. Taken incompany with my late friend Dr. D. Robert-
son in 1888 off Fairland Point, Isle of Cumbrae, in 20 fathoms:
Mus. Nor. Deep water off May Island, in the Firth of Forth,
and in the Moray Firth (7. 8.).
Distrib. West Norway (G. O. Sars) : Mus. Nor. Hansen
records it with doubt from Greenland.
Genus 4. METOPINA, gen. nov.
I institute this genus to receive three aberrant forms of
Metopa, in which the first gnathopods are not subchelate, and
have the hand long and very slender, and the terminal joint
very minute, flattened, not nail-formed. The second gnatho-
pods are very robust; the last pereeopods have the posterior
projection of the meros well and often enormously developed.
The type species is Metopa palmata, Sars; the others are
M. clypeata, Kroyer, and M. robusta, Sars.
108. Metopina robusta, G. O. Sars.
1892. Metopa robusta, G. O. Sars, (142) p. 270, pl. xevi. fig. 1.
1894. Metopa robusta, Thomas Scott, ‘‘ Some new and rare Crustacea
from Scotland,” Ann. & Mag, Nat. Hist. ser. 6, vol, xiii, p, 148.
Hab. Firth of Forth, 1884 (Dr. J. R. Henderson): Mus.
Nor. In this same locality Mr. Scott subsequently found
three specimens, and first recorded the species as British.
He has since also found it in the Moray. Firth.
Distrib. Varanger Fiord, Kast Finmark, 125-150 fathoms,
1890 (A. M. N.): Mus. Nor. Sars’s specimens were from
Bejan, in the outer part of the Trondhjem Fiord, and
Hammerfest, West Finmark ; in both places in 30-50 fathoms
among Hydroids, Tromsé (Schneider),
46 Canon A. M. Norman on British Amphipoda.
Genus 5. Cressa, Boeck.
=Danaia, Bate (nec M.-Edw. & Haime).
109. Cressa dubia (Bate).
1861. Danaia dubia, Bate & Westwood, (1) vol. i. p. 68.
1870. Cressa Schiddter, Boeck, (137) p. 65.
1876. Cressa Schiédtez, Boeck, (138) p. 467, pl. xviii. fig. 8.
1876, Danaia dubia, Stebbing, “ New and little-known Amphipodous
ae Ann. & Mag. Nat. Hist. ser. 4, vol. xviii. p. 444, pl. xix.
os, 2, 2a-c.
1890. Cressa dubia, J. Bonnier, “ Les Amphipodes du Boulonnais,”
Bull. Sci. de France et Belgique, vol. xxii. p. 500, pl. x. figs. 1-11.
1892. Cressa dubia, T. Scott, Tenth Ann. Rep. Fishery Board of
Scotland, p. 262, pl. viil. fig. 15 (the mandible).
ae gs dubia, G, O. Sars, (142) p. 278, pl. xeviii. fig. 2, & pl. exix.
od.
Hab. Off the Isle of Cumbrae (A. I. N.); Torquay
(Stebhing): Mus. Nor. Firth of Forth; Moray Furth; off
Tarbert, Loch Fyne (7. S.); off Cullercoats, Northum-
berland, 20 fathoms (G. 8. Brady & fide Stebbing) ; North
Wales; Isle of Man (A. O. W.) ; Isle of Mull (G. Brook).
Distrib. South and West Norway as far north as the
Trondhjem Fiord (G. O. Sars). Boulogne (Bonnier); 8.
Croisic (Chevreux).
Fam. IX. Leucothoide.
Genus 1. Leucornor, Leach.
110. Leucothoe spinicarpa (Abildgaard).
17-9. Gammarus spinicarpus, Abildgaard, Zool. Dan. vol. iii. p. 66,
pl. cxix. figs. 1-4.
1804. Cancer (Gammarus) articulosus, Montagu, “ Desc. several new
Animals found on South Coast of Devonshire,” Trans. Linn. Soc.
vol. vii. p. 70, pl. vi. fig. 6.
1862. Leucothoe articulosa, Bate & Westwood, (1) vol. i. p. 271.
1892. Leucothoe spinicarpa, G, O. Sars, (142) p. 283, pls. ¢. & ci. fig. 1.
1893. Leucothoe spinicarpa, Della Valle, (159) p. 652, pl. vi. fig. 4,
& pl. xix. figs. 1-20.
Hab. Shetland in Ascidians ; Skye; tlhe Minch; Cumbrae;
Loch Fyne in Ascidians; Lulworth, Dorset; Plymouth ;
Jersey ; Roundstone,;Ireland, in Ascidians (d. Mf, W.);
Polperro, Cornwall (Laughrin): Mus. Nor. St. Andrews
(M‘Intosh) ; Firth ot Forth (2. Scott); Anglesea; Isle of
Man; Valentia, Ireland (A. O. W.).
Distrib. Hardanger and ‘Trondhjem Fiords, Norway
(A, M. N.); Adriatic (from Heller and Claus as L. denti-
Canon A. M. Norman on British Amphipoda. 47
culata, A. Costa) ; Naples (Della Valle): Mus. Nor. Katte-
gat (Metnert); Greenland (Hansen) ; W. France (Chevreuz) ;
Azores (Barrois).
111. Leucothoe Lilljeborgit, Boeck.
1855. Leucothoe articulosa, Lilljeborg, Céfvers. af K, Vet.-Akad.
Forhand. p. 126.
1860. Leucothoe Lilljeborgvt, Boeck, Forhand. Skand. Naturf. 8de
Made, p. 653.
1888. Leuccthoe furina, Chevreux, Bull. Soc. d’Etudes Scient. de Paris,
11° année, 1 semestre, p. 9 (separate copy) (nec LZ. furina, Savigny)
1889. Leucothoe imparicornis, Norman, “ Notes on British Amphipoda,”
Ann. & Mag. Nat. Hist. ser. 6, vol. iv. p. 114, pl. x. figs. 1-4.
1892. Leucothoe incisa, D. Robertson, “ Second Contrib. Cat. Amphip.
and Isop. Firth of Clyde, &c.,” Trans. Nat. Hist. Soc. Glasgow,
vol. iii. (p. 25, separate copy).
1892. Leucothoe Lilljeborgii, G. O. Sars, (142) p. 284, pl. ci. fio, 2,
1897. Leucothoe incisa, Stebbing, “ Amphipoda from Copenhagen
Museum and other Sources,” Trans. Linn. Soc., 2nd ser. Zool. vol. vii.
Pi 505; pl..x.
.
The Leucothoe described by Sars under the name of
L. Lilljeborgit, Boeck, is certainly the same as my L. impari-
cornis. My description was taken from a mounted and only
partially dissected specimen, and thus the recurved process
of the last segment of the metasome could not be seen. It
appears to me that the characters by which Mr. Stebbing
would separate L. incisa, Robertson, from this species are
insufficient ; the gnathopods, especially those of the first pair,
show some differences dependent upon age.
This species has been named at former periods “ Leucothoe
furina” by Sars, Chevreux, and myself.
I am also inclined to think that Lewcothoe serratipalma,
Della Valle, is the same species. The chief difference, and
it is a very unimportant one, appears to be that there is a
slight serration at the commencement of the palm of the
second gnathopods, where the point of the finger impinges on
the palm.
Hab. Shetland, 1863 (A. MW. NW.) ; Banff (7. Edward) ;
Ballock Bay and off Fairland Point, Isle of Cumbrae,
20 fathoms (J. #.); near the Bass Rock and in Upper
Loch Fyne (7. S.) ; Mull (@. Brook, fide T. 8.) ; Pool Krin,
Isle of Man, and Guernsey (A. O. W.).
Distrib. Kopervig, west coast of Norway, 40-60 fathoms
(G. OU. Sars) ; Kullaberg, Sweden (Liljeborg) ; West France
(Chevreux) ; Naples (?) (Della Valle).
48 Canon A. M. Norman on British Amphipoda.
Fam. X. Cdiceride.
[Genus 1. Gipiceros, Kroyer.
[ Gidiceros saginatus, Kroyer.
1865. Cediceros saginatus, Goés, “ Crust. Amphip. Maris Spetsberg.
&e.,” Cifvers. af K. Vet.-Akad. Forhand. p. 10 (separate copy),
pl. Xxxix. fig. 18,
1876. Cediceros saginatus, Boeck, (138) p. 257, pl. xiii. fig. 3.
1892. Cdiceros saginatus, G. O. ‘Sars, (142) p. 288, pl. cii.
Greenland, lat. 67° 50! N., long. 55° 17' W., 20 fathoms,
‘ Valorous,’ 1875.
Vadsé, East Finmark, 1890 (A. M. N.): Mus. Nor.
Iceland (Torell) ; Spetsbergen (Goés) ; Murman coast (Jar-
zynski); Siberian Polar Sea (Stwxberg). A truly Arctic
species, the most southern locality from which it has been
recorded is Christiansund (V. Diiben, fide Sars).]
[| Hdiceros boreaits, Boeck.
1870. Gidiceros borealis, Boeck, (137) p.
1876. CGdiceros borealis, Boeck, (138) p. 61, pl. xiv. fig. 1.
1892, Gdiceros borealis, G. O. Sars, (142) p. 290, pl. cil. fig. 1
Holsteinborg Harbour, Greenland, in 7-35 fathoms,
‘ Valorous,’ 1875.
Finmark (Boeck) ; Kara Sea (Stuxberg).
The Gdiceros borealis of Buchholz (‘ Die zweite deutsche
Nordpolarfahrt,’ 1874, vol. 11. p. 325, pl. v.) cannot be
this species, nor, indeed, a member of this genus. It is
perhaps Monoculodes simplex, H. J. Hansen, but the telson
does not seem to agree. |
[Genus 2. Parapiceros, G, O. Sars,
[ Pareediceros lynceus (M. Sars).
1858. CGediceros lynceus, M. Sars, “ Oversigt over de i den Norsk-
Arctiske Region forekommende Krebsdyr,” Christ. Vid.-Selsk.
Forhand. p, 143.
1876. Ctdiceros lynceus, Boeck, (188) p. 259, pl. xiii, fig. 4.
1883. Ctdiceros lynceus, J.8. Schneier o Norges kyster forekommende
Arter af familien Cidiceride,” Tromso Museums Aarshelter, vi.
p- 14, pl. ii. fig. 12.
1892. Parediceros lynceus, G. O, Sars, (142) p. 292, pl. ciii. fig. 2
& pl. civ. fig, 1.
Holsteinborg Harbour, Greenland, 7-35 fathoms, ‘ Valorous,’
1875.
Sydvaranger and Varanger Fiord, Kast Finmark ; Svolver,
Lofoten Islands, 5-10 fathoms, 1890 (4. MZ. N.) ; Tromsé
Canon A. M. Norman on British Amphipoda. 49
(J. S. Schneider); Spetsbergen (Lovén); Gulf of Maine,
N.E. America, 90 fathoms (Prof. 8S. I. Smith): Mus. Nor.
Barents Sea (Hoek); Iceland (TZorel/?) ; Murman coast
(Jarzynski) ; Siberian Polar Sea (Stuaberg); Franz-Josef
Land, Jackson-Harmsworth Exped. (7. 8.) ; Iceland (Tore//);
East Greenland (Buchholz) ; Barents Sea (Hoek) ; Murman
coast, Kara Sea, and Jugor Schar (Stuxberg).|
Genus 3. MONOCULODES, Stimpson.
112, Monoculodes carinatus, Bate.
1855. Westwoodia carinata, Brit. Assoc. Rep. p. 58.
1859. Gidiceros affinis, Bruzelius, Skand. Amphip. Gammariden, p. 93,
pl. iv. fig. 15, g junior.
1862. Monoculodes carinatus, Bate & Westwood, (1) vol. i. p. 165.
1862. Monoculodes Stimpsoni, iid. ibid. p. 160.
1876. Monoculodes affinis, Boeck, (188) p. 265, pl. xiv. fig. 6, 2.
1883. Monoculodes carinatus, Schneider, Af Norges kyster forekom.
Art. af Gidiceride, p. 19, pl. i. fig. 4.
1889. Monoculodes carinatus, Norman, ‘‘ Notes on British Amphipoda,”
Ann. & Mag. Nat. Hist. ser. 6, vol. iii. p. 447, pl. xix. figs. 1-5.
1892. Monoculodes carinatus, G. O. Sars, (142) p. 295, pl. ev.
1893, Cidiceros affinis, Della Valle, (189) p. 548, pl. iv. fig. 3 &
pl. xxxiii. figs. 27-31. %
For notes on the above synonymy see my paper in the
‘Annals.’ It is not Gdiceros affinis, Goés.
Fab. Shetland; Isle of Cumbrae; off the Northumberland
coast; Plymouth (A. M. N.); Moray Firth (7. Edward) ;
off May Island, Firth of Forth, 35 fathoms (Sir J. Murray) :
Mus. Nor. Loch Fyne (Sir J. Murray); Isle of Man,
Jersey, and Valentia, Ireland (A. O. W.).
Distrib. Floré, Norway (A. M. N.); Croisic, France
(Chevreux) ; west coast of Sweden (Bruzelius); Kattegat
(Meinert) ; Naples (A. MM. N.).
113. Monoculodes subnudus, Norman.
1889. Monoculodes subnudus, Norman, ‘‘ Notes on British Amphipoda,”
Ann. & Mag. Nat. Hist. ser. 6, vol. il, p. 450, pl. xvi. fig. 11,
& pl. xix. figs. 6-10.
1892. Monoculodes falcatus, G. O. Sars, (142) p. 802, pl. evil. fig. 2.
Hab. Shetland; Sleat Sound, Isle of Skye (4. M. N.):
Mus. Nor.
Distrib. Kors Fiord, near Bergen ; Rédberg, in the Trond-
hjem Fiord, in 150 fathoms, and Klosterelv Fiord, East
Finmark (A. M. N.): Mus. Nor. Sars has found it at several
places in West Norway.
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 4
50 Canon A. M. Norman on British Amphipoda.
114. Monoculodes borealis, Boeck.
1865. Gdiceros affinis, Goés (nec Bruzelius), ‘Crust. Amphip. Maris
Spetsb. &c.,”’ Gifv. K. Vet.-Ak. Férh. p. 11 (sep. cop.) (partim),
pl. xxxix. fig. 21’ (nec 21).
1870. Monoculodes borealis, Boeck, (137) p. 88.
1876. Monoculodes borealis, Boeck, (138) p. 278, pl. xv. fig. 4.
1883. Monoculodes borealis, Schneider, J. ¢. p. 22, pl. i. fig. 3.
1892. Monoculodes borealis, G. O. Sars, (142) p. 298, pl. evi. fig. 2.
Hab. Dredged off Blackwater-foot, Arran, N.B., in 20
fathoms, muddy sand (D. f.).
Distrib. Roédberg, in the Trondhjem Fiord, in 40-150
fathoms; Bog Fiord, EK. Finmark, 3-5 fathoms (A. M. N.) ;
Tromsé (Schneider): Mus. Nor. The former is the most
southern locality in which it has been found by Sars;
Spetsbergen (Goés) ; Greenland (Hansen) ; Kara Sea (Stua-
berg) ; Franz-Josef Land, Jackson-Harmsworth Exped.
CEOS)
115. Monoculodes tuberculatus, Boeck.
1865. Gdiceros affinis, Goés (nec Bruzelius), “ Crust. Amphip. Maris
Spetsb. &c.,” Cifv. K. Vet-Ak. Forh. p. 11 (sep. cop.) (partim),
pl. xxxix. fig. 21 (nec 21’).
1870. Monoculodes tuberculatus, Boeck, (137) p. 87.
1876. Monoculodes tubercalatus, Boeck, (138) p. 277, pl. xv. fig. 2.
1883. Monoculodes tuberculatus, Schneider, J. ¢. p. 29, pl. i. fig. 8.
1892. Monoculodes tuberculatus, G. O. Sars, (142) p. 308, pl. evii. fig. 3.
Hab. A single specimen in tow-net gathering a few miles
east of the Island of Arran, N.B. (7. Scott)—No doubt the
tow-net here meant was a net attached near the dredge, and
not at surface.
Distrib. West coast of Norway ; Trondhjem Fiord; Va-
ranger Fiord, East Finmark (G. O. Sars); Tromsé
(Schneider) ; Spetsbergen (G'oés) ; Greenland (Hansen).
116. Monoculodes Packardi, Boeck.
1870. Monoculodes Packardi, Boeck, (137) p. 86.
1876. Monoculodes Packurdi, Boeck, (138) p. 274, pl. xiv. fig. 3.
1883. Monoculodes Packardi, Schneider, /. c. p. 27, pl. i. fig. 6.
1892. Monoculodes Packard. G. O. Sars, (142) p. 807, pl. cix. fig. 1.
Hab. Loch Striven, Firth of Clyde, 40 fathoms (D. 2.) :
Mus. Nor. Upper Loch Fyne (T. S.).
Distrib. Varanger Fiord, E. Finmark, 125-150 fathoms
(A. UM. N.); Tromsé (Schneider): Mus. Nor. On whole
coast of Norway from Christiania to Vadsé in 10-100 fathoms
(G. O. Sars).
On a new Sea-Horse from Muscat. aL
Genus 4. PerrocuLopes, G. O. Sars.
117. Perioculodes longimanus (Bate).
1869. Monoculodes longimanus, Bate & Westwood, (1) vol. ii. p. 507.
1870. Monoculodes Grubet, Boeck, (137), p. 85.
1876. Monoculodes Grubet, Boeck, (138) p. 269, pl. xvi. fig. 1.
1883. Monoculodes Grubei, Schneider, J. ¢. p. 24.
1887. Monoculodes longimanus, Chevreux, Cat. Crust. Amphip. du
Sud-ouest de la Bretagne, p. 13, pl. v. figs. 1, 2.
1888. Monoculodes equimanus (Norman, MS.), Robertson, “ Contrib.
Cat. Amphip. and Isop. of Firth of Clyde,” Trans. Nat. Hist. Soc.
Glasgow, vol. ii. B 26 (separate copy), 3.
1889. Monoculodes longimanus, Norman, “ Notes on British Amphip.,”
Ann. & Mag. Nat. Hist. ser. 6, vol. iii. p. 451, pl. xx. figs. 6-9.
1892. Perioculodes longimanus, G. O. Sars, (142) p. 313, pl. ex. fig. 2,
lexi, fie. ¥:
1893. Celiceros longimanus, Della Valle, (139) p. 547, pl. iv. fig. 9,
pl. xxxill. figs. 32-36.
Hab. Kames Bay, Isle of Cumbrae, 1-4 fathoms ; Oban;
off Marsden, Co. Durham, 10 fathoms ; Starcross, Devon, 2-3
fathoms; Clew Bay, Co. Mayo (A. M. N.); Banff (T.
Edward) ; Firth of Forth (7. 8.) : Mus. Nor. Upper Loch
Fyne and off Spurm Head (T. S.); Isle of Man; North
Wales ; Jersey ; Guernsey; Valentia, Ireland (A. O. W.).
Distrib. Svolver, Lofoten Islands; Trondhjem Fiord,
5 fathoms (A. M. NV.) ; South Norway (G. O. Sars) ; Naples
(A. M. N. & Della Valle): Mus. Nor. Kattegat (Meinert);
West France (Chevreux) ; Spezia (G. O. Sars).
EXPLANATION OF PLATE IIL.
Fig. 1. Amphilochus neapolitanus, Della Valle. Second gnathopod.
Fig. 2. Stenothoe setosa, sp.n. First gnathopod.
Fig. 3. Ditto. Second gnathopod.
Fig. 4. Ditto. Last peraeopod.
Fig. 5. Metopa rubrovittata, G. O. Sars. First gnathopod.
5
6. Metopa abscisa, sp. n. First gnathopod.
7. Ditto. Terminal joints of first gnathopod, more magnified.
Fig. 8. Ditto. Second gnathopod.
9. Ditto. Palm of second gnathopod, more magnified.
0. Ditto. Last perzeopod.
V.—Description of a new Sea-Horse (Hippocampus) from
Muscat. By G. A. BOULENGER, F.R.S.
Hippocampus Jayakari.
11 segments on the body. All the tubercles produced into
long slender spines; supraorbital spine longer than the
diameter of the orbit, longer than the azygous spine in front
4%
j
52 - Mr. G. A. Boulenger on Fishes
of it; two spines below the orbit ; a very long spine, flanked
by a similar one on each side, in front of the coronet, which
is rather elevated and terminates in five spines. Snout three
times as long as the eye, half the length of the head. Dorsal
with 18 rays, inserted on four segments, two of the body, two
of the tail. Yellow laterally and ventrally, brown dorsally,
with dark brown rings; most of the spines yellow at the tip.
A single specimen, measuring 85 millim. when stretched
out, was picked up at Muscat by Surgeon-Lt.-Col. A. 8. G.
Jayakar, who, on kindly sending it as a present to the
British Museum, pointed out to me its close affinity to
H. hystrix, Kaup. The latter differs, however, in the longer
snout and the shorter and more acutely pointed spines on the
head, also in the coloration.
Dr. Jayakar is, I regret to hear, leaving Muscat. It
gives me great pleasure to connect his name with this
latest discovery of his at the station where for so many
years he has been collecting fishes for the British Museum,
with the result of enriching ichthyology by many new species,
which have been described in the ‘ Proceedings of the Zoolo-
gical Society ’ and in these § Annals.’
VI.—A List of the Fishes collected by Mr. Rupert Vallentin
in the Falkland Islands. By G. A. BouLENGER, F.R.S.
With Notes by the Collector.
Trachinide.
1. Eleginus maclovinus, C. & V.
“ ¢ Mullet,’ as they are called, abounded in the sea, especially
from the Falkland Islands. 53
on the sands near the lighthouse at Cape Pembroke. The
fish from this locality were large, from 380-36 inches in
length, and were mostly meshed ina trammel. The specimens
brought homewere caught in asmall seine in Stanley Harbour.”’
2. Notothenia sima, Richards.
Stanley Harbour. Very common.
3. Notothenia macrocephalus, Gthr.
Stanley Harbour.
4, Harpagifer bispinis, Forst.
“ The little Cottus-like fish occurring so abundantly along
the shores of Stanley Harbour in shallow pools during low
water were subject to great variations in colour; indeed one
never found two specimens marked exactly the same. The
ground-colour of these fish was usually grey, with longitu-
dinal streaks of darker tint. Occasionally one would find a
specimen with brilliant carmine patches on the head, with
bands of the same colour running along the dorsal surface
posteriorly above the pectoral fins. The ventral surface was
invariably of a uniform light grey. I was unable to keep
any of these brilliantly coloured specimens in captivity so as
to find out if these bright colours were permanent or only
transitory. These variations of colour had nothing to do
with local surroundings, there being no red sea-weeds in
Stanley Harbour—at least, I could find none—although
there were plenty in the open ocean.”
Lycodide.
5. Lycodes latitans, Jen.
“ Fairly common along the shore and also in the deep
water in Stanley Harbour.”
6. Lycodes flavus, sp. n.
Depth of body equal to length of head, 8 times in total
length. Head as deep as broad, once and a half as long as
broad; snout short, rounded, projecting beyond the mouth,
which extends to below the posterior border of the eye; the
diameter of the latter equals interocular width and is con-
tained once and one third in length of snout, five and a half
times in length of head; nostril tubular, near the end of the
54 On Fishes from the Falkland Islands.
snout ; large pores on the snout and on both lips; a single
series of moderately large conical teeth in the jaws and on
the vomer; gill-opening very narrow. Body naked, com-
pressed ; lateral line indistinct; the vent more than twice as
distant from the end of the tail as from the ventrals. Dorsal
with about 80 rays, originating just behind the occiput; anal
with about 60 rays. Pectoral about two thirds length of
head; ventral nearly one third length of pectoral.
Dark or light saffron-colour; a dark brown stripe along
each side of the head, passing through the eye, separated
from the dark brown upper surface of the head by a narrower
yellowish-white stripe; lower surface of head and body
yellowish white.
Total length 113 millim.
This species differs strikingly from L. latitans, apart from
the coloration, in the shorter head and the larger eye.
“The specimens secured were detected in the hollow tangled
roots of Macrocystis, whilst the specimens of the blackish-
brown L, latitans were found under stones resting on mud.”
Galaxiide.
7. Galaxias attenuatus, Jen.
“ Fish known to the inhabitants as ‘smelts’ were fairly
common, and occurred in shoals in the shallow water along
the shore. The specimens brought home were dipped from
the sea with a large hand-net while being pursued by a
penguin (Hudyptes chrysolophus).”
8. Galaxias maculatus, Jen.
“Freshwater fish. The Falkland Island ‘trout’ is very
abundant in certain small brooks and streams in the Falkland
Archipelago. The specimens sent were secured in a small
brook called Wier Creek, a small stream in direct communi-
cation with the higher portion of Port William.
‘* Owing to the abundance of peat and the heavy rainfall one
would naturally expect these fish to be black in colour, like
moorland-trout. Great was my astonishment to find these
Falkland Island trout almost transparent, the principal blood-
vessels being plainly visible in the living fish.”
On Asopine and Tessaratomine. 53
VII.—Rhynchotal Notes.—V. Heteroptera: Asopine and
Tessaratomine. By W. L. DIsTant.
THE present communication calls for little comment. ‘The
Asopine still require more generic revision to bring them
into line with the main division of the Pentatomide, to which
they belong. The cause is not difficult to see, and is an
adherence to supposed authority. Stal treated the Asopinze
in the first part of his ‘numeratio Hemipterorum,’ and
there used subgenera for what in the later parts of his work
he would distinctly have regarded as genera.
This paper refers to Walker’s genera and species belonging
to the above two subfamilies, as contained in vols. 1.11. of
his ‘ Catalogue of Hemiptera-Heteroptera.’
ASOPINZE.
Genus HETEROSCELIS.
Heteroscelis bimaculata.
Platynopus bimaculatus, Walk. Cat. Het. i. p. 124, n. 6 (1867).
Genus MINEUS.
Mineus triangularis.
Strachia triangularis, Walk. Cat. Het. ii. p. 323. n. 42 (1867).
Genus CERMATULUS.
Cermatulus nasalis.
ZBlia nasalis, Westw. in Hope Cat. i. p. 32 (18387).
Rhaphigaster pentatomoides, Walk. Cat. Het. ii. p. 370. n. 81 (1867).
Asopus binotatus, Walk. Cat. Het. i. p. 144 (1867).
Walker gives Brazil as a habitat for his A. dinotatus,
which is clearly an error, there being not the slightest doubt
that it is a synonym of the common Australian species.
Genus DorRyYCORIS.
Dorycoris fuscosus.
Asopus fuscosus, Germ. in Silb. Rev. p. 187 (1837).
Rhaphigaster perornatus, Walk. Cat. Het. iii. p. 567 (1868).
56 Mr. W. L. Distant on
Genus EALDA.
Ealda, Walk. Cat. Het. ii. p. 409 (1867).
Abdomen with the lateral segmental angles moderately
spinous, the two penultimate angles strongly spinous, the
apical angles longly spinous.
I have placed the genus near Hoploawys, Dall.
Ealda minaz.
Ealda minax, Walk. Cat. Het, ii. p. 409, n. 1 (1867).
Hab. New Caledonia.
GORDONERIUS, gen. nov.
Head moderately long and broad, subtruncate in front,
where it is slightly notched centrally; antenne with the
second joint a little longer than the third; rostrum reaching
the intermediate cox, third joint a little shorter than second
or fourth joints, which are subequal. Pronotum with the
lateral margins very slightly sinuate, much broader at base
than long, posterior angles subprominent. Scutellum nar-
rowed at about centre, apex rounded. _Abdomen with a short
basal spine; anterior femora spined before apex; anterior
tibie very slightly dilated.
In general shape and appearance allied to Dorycoris and
Zicrona, from both of which it is at once distinct by the
character of the spined anterior femora.
Gordonerius lineatus.
Oplomus lineatus, Walk. Cat. Het. i. p. 122. n. 20 (1867).
Hab. Abyssinia (Brit. Mus.).
Genus PLATYNOPUS.
Platynopus rostratus.
Cimex rostratus, Drury, Ill. Nat. Hist. iii. p. 59, pl. xliii. fig. 3 (1782).
Platynopus badius, Walk. Cat. Het. i. p. 126. n. 12 (1867).
Var. Platynopus trijunctus, Walk. loc. cit. n. 11.
Var. Platynopus optabilis, Walk. loc, cit. p. 126. n. 13.
Platynopus parvus, sp. n.
Allied to P. rostratus, Dru., but much smaller; colour
paler, pale castaneous, luteous markings similar but paler ;
Asopine and 'Tessaratominz. 57
antenne with the basal joint and the bases of the fourth and
fifth joints ochraceous; second and third joints subequal in
length. Other characters as in P. rostratus.
Long. 9-10 millim.; exp. pronot. angl. 54-6 millim.
Hab. Sierra Leone (Brit. Mus.) ; Congo (Coll. Dist.).
Platynopus scutellatus, sp. n.
Above bright shining ochraceous. Head dark olivaceous
green, with a central lineate ochraceous spot near base; an-
tenne fuscous. Pronotum sparingly, coarsely, and darkly
punctate ; a distinct submarginal lateral linear fascia of dark
punctures, two blackish ringlets on anterior portion of disk,
and the pronotal spines shining black; the space behind the
pronotal angles and the posterior margin impunctate. Scu-
tellum sparingly, coarsely, and darkly punctate, the basal
angular areas impunctate, and a small dark spot in each
basal angle. Corium sparingly, coarsely, and darkly punc-
tate. Membrane cupreous. Body beneath and legs ochra-
ceous ; sternum coarsely punctate, a large impunctate spot on
each lateral area of pro-, meso-, and metasternum; a trans-
verse fascia at junction of pro- and mesosternum, a much
waved sublateral fascia, large subapical spot and anal ap-
pendage to abdomen, anterior and intermediate femora (ex-
cluding base and a subapical spot), outer surface and base
and apex of inner surface of anterior tibie, apices of posterior
femora, bases and apices of intermediate and posterior tibia,
and the tarsi castaneous.
Scutellum with the apex obtusely angulate. Anterior
tibie strongly dilated, Abdominal basal spine long, passing
the posterior coxe.
Long. 13 millim. ; exp. pronot. angl. 8 millim.
Hab. Upper Congo, Bopoto (Coll. Dist.).
A species to be known by its angulated apex to scutellum
and distinct coloration.
Platynopus melanoleucus.
Elia melanoleuca, Westw. in Hope Cat. i. p. 33 (1837).
Platynopus polygraphus, Walk, Cat. Het. i. p. 126. n. 17 (1867).
Var. Platynopus melanoleucus, Dall. (nec Westw.), List Hem. i. p. 87.
n. 1 (1851).
The British Museum possesses specimens from Celebes and
Ceram which agree with Westwood’s type from Java. All
the Philippine specimens I have seen are of a varietal form in
which the pronotal fascie are more strongly marked and
continuous.
58 Mr. W. L. Distant on
Platynopus leetus.
Platynopus letus, Walk. Cat. Het. i. p. 127. n, 19 (1867).
Platynopus dotatus, Walk. loc. cit. p. 128. n. 20.
Platynopus semiscitus, Walk. loc. cit. p. 129. n. 21.
Canthecona decorata, Voll. Faun. Ind. Neerl. iii. p. 9, pl. ii. fig. 2 (1868).
Platynopus melacanthus.
Pentatoma melacantha, Boisd. Voy. Astrol., Ent. p. 628, pl. ii. fig. 7
(1835).
Platynopus tenellus, Walk. Cat. Het. i. p. 127. n. 18 (1867).
Platynopus splendidulus.
Cimex splendidulus, Fabr. Syst. Rhyng. p. 163. n. 40 (1803).
Platynopus splendidulus, Stal, Hem. Fabr. i. p. 16. n. 1 (1868),
Oplomus elongatus, Dall, Trans. Ent. Soc. Lond. 1852, p. 6, pl. i. fig. 1.
Dallas gave the habitat of his species as “ in Brasilia?”
Specimens exactly agreeing with his description, which repre-
sents the form described by Fallou as P. metallicus (Rev.
d’Ent. x. p. 5, 1891), are in the British Museum from Sierra
Leone and in my own collection from West Africa and the
banks of the River Aruwini (Central Africa).
Genus CANTHECONA.
Canthecona cognata.
Canthecona cognata, Dist. Ent. Month. Mag. xix. p. 157 (1882).
Canthecona insularis, Kirby, Journ. Linn. Soc., Zool. xxiv. p. 79,
pl. iv. fig. 4 (1891).
Genus PICROMERUS.
Picromerus obtusus.
Picromerus obtusus, Walk. Cat. Het. i. p. 183. n. 6 (1867).
Picromerus nigrivitta, Walk. loc. cit. n. 7.
Genus Popisus.
Podisus modestus.
Arma modesta, Dall. List Hem. i. p. 101. n. 18 (1851).
Rhaphigaster aggressor, Walk. Cat. Het. ii. p. 359. n. 19 (1867).
Podisus semialbatus.
Mormidea semialba, Walk. Cat. Het. iii. p. 553 (1868).
A species allied to P. falcatus, Dist.
Asopine and Tessaratomine, 59
Podisus turbidus.
Arma turbida, Walk. Cat. Het. i. p. 140. n. 43 (1867).
This species is still represented only by the unlocalized
type specimen in the British Museum. It is allied to
P. fuscescens, Dall.
Rostrum dark castaneous, the second joint black.
Podisus cynicus.
Pentatoma cynica, Say, New Harm. Ind., Dec. 1831 ; Compl. Writ. i.
p. 312. n. 1 (1859).
Arma grandis, Dall. List Hem. i. p. 96. n. 3 (1851).
Podisus cynicus, Leth. & Sev. Cat. Gén. Hém. t. i. p. 217 (1898).
Podisus grandis, Leth. & Sey. loc. cit. p. 218.
Uhler (Check-list Hem. Het. N. America, 1886, p. 4)
correctly sank Dallas’s species as a synonym of the above ;
but as the authors of our best catalogue have kept them as
distinct, it is necessary to again draw attention to their being
conspecific.
Genus TYLOSPILUS.
Tylospilus megaspilus.
Hoploxys megaspilus, Walk. Cat. Het. i. p. 141, n. 2 (1867).
Genus ANASIDA.
Anasida funebris, sp. n.
Pitchy black ; eyes and a spot at base of head ferruginous.
Head thickly and finely punctate, with a small impunctate
space before each eye; antenne with the second and third
joints subequal in length, remaining joints mutilated. Pro-
notum coarsely punctate and rugulose, the sublateral carinz
near posterior angles very distinct and prominent. Scutellum
coarsely punctate and rugulose. Corium more opaque,
smooth, somewhat more sparingly punctate. Membrane
shining bronzy.
Long. 16 millim.; exp. pronot. angl. 7 millim.; abd.
74 millim.
Hab. Natal (Gueinztus: Brit. Mus.).
Allied to the West-African type of the genus, A. tenebrio,
Karsch, but differing by the scutellum being more elongate
and distinctly and angularly narrowed at about two thirds
from base; abdomen narrower, with the connexivum much
les prominent, &c.
60 Mr. W. L. Distant on
Genus ASOPUS.
Asopus erythromelas.
Strachia erythromela, Walk. Cat. Het. ii, p. 339, n. 81 (1867).
Strachia pyrophila, Walk. loc. cit. p. 340. n. 88.
Var. Strachia precipua, Walk. loc. cit. p. 339. n. 82.
These described forms were reported by me (‘ Annals,’
May 1900, p. 434) as not to be found under the genus and
subfamily in which they were described. I have since
discovered them under the genus Asopus, where I find Walker
subsequently mentions them (Cat. Het. i. pp. 533, 534,
1868).
Genus CHCHALIA.
Cchalia consocialis.
Pentatoma consociale, Boisd. Voy. Astr., Entom. ii. p. 630. n. 3, pl. ii.
fig. 9 (1835). ;
Rhaphigaster perfectus, Walk. Cat. Het. ii. p. 371. n, 88 (1867).
Mormidea decora, Walk. Cat. Het. ii. p. 256. n. 25 (1867).
This species clearly represents an undescribed genus in
Asopine; but as the type and only specimen is without
antenne, it must remain for the present a generic nondescript.
Abdomen unarmed; rostrum reaching the posterior coxe ;
head rounded in front, the central lobe longer than the
lateral lobes ; posterior pronotal angles produced into mode-
rately long, slender, acute spines; scutellum somewhat long,
subtriangular, its apex extending a little beyond base of
membrane.
T'BSSARATOMINAE.
Genus PIEZOSTERNUM.
Piezosternum subulatum.
Cimex subulatus, Thunb. Nov. Ins. Sp. ii. p. 41, pl. i. fig. 55 (1788).
Piezosternum retractum, Walk. Cat. Het. iii. p. 467. n. 4 (1868).
Salica excellens, Walk. loc, cit. p. 469. n. 1.
Genus TESSARATOMA.
Tessaratoma papillosa.
Cimex papillosus, Drury, Ill. Nat. Hist. 1. p. 96, pl. xliii. fig. 2 (1770).
Tesseratoma striata, Walk. Cat. Het. ii. p. 463. n. 16 (1868).
Tesseratoma furcifera, Walk. loc. cit. n. 17.
Tesseratoma timorensis, Walk. (Voll. ?) loc. cit. p. 464. n. 18.
Asopine and 'Tessaratomine. 61
Genus HYPENCHA.
Hypencha apicalis.
Tessaratoma apicalis, St.-Farg. et Serv. Enc. Méth. x. p. 591. n. 3
(1825).
Tesseratoma semicuprea, Walk. Cat. Het. iii. p. 465. n. 21 (1868).
HHypencha opposita.
Tesseratoma opposita, Walk. Cat. Het. iii. p. 466. n. 22 (1868).
Walker’s type specimen was unlocalized. The Museum
now possesses a second specimen from Johore.
Genus PYGOPLATYs.
Pygoplatys cribatus.
Prezosternum cribatum, Walk. Cat. Het. iii. p. 458. n. 6 (1868).
Pygoplatys trucidus, Walk. loc. cit. p. 461. n. 8.
Genus HUSTHENES.
Eusthenes thoracicus, sp. n.
Head, pronotum, scutellum, and legs piceous; corium and
body beneath very dark castaneous; apex of scutellum and
eyes reddish ochraceous. Antenne black; second joint a
little longer than the third, remainder mutilated. Pronotum
with the lateral margins distinctly reflexed, rounded ante-
riorly, and a little concavely sinuate before the posterior
angles, which are subprominent ; before the rounded anterior
lateral margins is a broad sublateral rugosity, which is
strongly transversely striate; remainder of disk somewhat
faintly striate and moderately punctate. Posterior femora in
male strongly incrassated, beneath with a long robust curved
spine near base, and with a series of small obtuse spines near
apex. Other characters as in E. robustus, Lepell. et Serv.,
but differing from that and all other species of the genus by
the structure of the pronotum.
Long. 36 millim.; exp. pronot. angl. 17 millim.; max.
Jat. abd. 20 millim.
Hab, N. India, Assam (Atkinson Coll., Brit. Mus.).
Genus DALCANTHA,
Dalcantha dilatata. |
Dalcantha dilatata, Amy. & Serv. Hem. p. 171. n. 1 (1848),
Dalcantha regia, Walk. Cat. Het, iii. p. 474, n. 3 (1868).
62 Mr. W. L. Distant on
Stal (En. Hem. i. p. 76, 1870) treated Walker’s species
as a synonym of D. Stali, Voll. Walker’s descriptions were
made from specimens identified by Dallas as D. dilatata,
which Stal had previously seen under that name when visiting
the British Museum.
Summarized Disposition of Walker’s Genera and Species.
Asopine and Tessaratomine.
Genera considered valid.
Blachia, Walk. Cat. Het. i. p. 117 (1867).
Cecyrina, Walk. loc. cit. p. 118.
Ealda, Walk. loc. cit. ii. p. 409.
Muscanda, Walk. loc. cit. iii. p. 576 (1868).
Genera treated as synonymic.
Bodetria, Walk. Cat. Het. i. p. 119 (1867), =Gen. Heteroscelis, Latr.
Gilva, Walk. loc. cit. p. 141, = Gen. Coryzorhaphis, Spin.
Salica, Walk. loc, cit. ili. p. 469 (1868), = Gen. Piezosternum, Amy. &
[Sery.
Species considered valid and described under correct Genera.
Blachia ducalis, Walk. Cat. Het. i. p. 117. n. 1 (1867).
Cazira internexa, Walk, loc. cit. p. 118. n. 4.
Cecyrina platyrhinoides, Walk. loc. ct. p. 119. n. 1.
Platynopus letus, Walk. loc. cit. p. 127. n. 19.
purpurascens, Walk. loc. cit. iii. p. 5380 (1868).
Canthecona concinna, Walk. loc. cit. 1, p. 181. n. 9 (1867).
Glypsus truculentus, Walk. loc. cit. p. 132. n. 4.
Picromerus obtusus, Walk. loc. cit. p. 183. n. 6.
Macrorhaphis infuscata, Walk. loc. cit. iii. p. 581 (1868).
spurcuta, Walk, loc. cit.
Ealda minax, Walk. loc. cit. ii. p. 409. n. 1 (1867),
Pygoplatys lancifer, Walk. loc. cit. iii. p. 460. n. 7 (1868).
Amissus nitidus, Walk. loc. cit. p. 466, n. 2.
Stphnus dilatatus, Walk. loc. cit. p. 467. n. 3.
Lyramorpha ramifera, Walk, loc. cit. p. 476. n. 4.
Muscanda testacea, Walk. loc. cit. p. 577.
Species considered valid, but requiring generic revision.
Oplomus lineatus, Walk. Cat. Het. i. p. 122. n. 20 (1867), belongs to gen.
Gordonerius, g. n.
Platynopus bimaculatus, Walk. loc. cit. p. 124. n. 6, belongs to gen. Hete-
roscelis, de Cast.
Arma eli Walk. loc. cit. p. 135. n. 15, belongs to gen. Podisus, Herr.-
chaff.
—— ampla, Walk. loc. cit. p. 138. n. 34, belongs to gen. Podisus, Herr.-
Schaff.
caliginosa, Walk. loc. cit, n. 35, belongs to gen. Podisus, Herr.-
Schaff.
Asopine and Tessaratomine. 63
Arma submarginata, Walk. loc. cit. p. 189. n. 87, belongs to gen. Podisus,
Herr.-Schaff.
turbida, Walk. loc. cit. p. 140. n. 43, belongs to gen. Podisus, Herr.-
Schaff.
Hoploxys megaspilus, Walk. loc. cit. p. 141. n. 2, belongs to gen. Tylospilus,
Stal.
Piezosternum firmatum, Walk. loc. cit. iii. p. 458. n. 5 (1868), belongs to
gen. Pygoplatys, Dall.
eribratum, Walk. loc. cit. n. 6, belongs to gen. Pygoplatys, Dall.
ingenum, Walk. loc. cit. p. 459. n. 7, belongs to gen. Pygoplatys, Dall.
Tesseratoma forticornis, Walk. loc. cit, p. 465, n. 23, belongs to gen. Pygo-
platys, Dall.
opposita, Walk. loc. cit. p. 466. n. 22, belongs to gen. Hypencha,
Amy. & Serv.
Pycanum stabile, Walk. loc. cit. p. 472. n. 12, belongs to gen. Carpona,
Dohrn.
Species treated as synonymic.
Bodetria brenthoides, Walk, Cat. Het. i. p. 119. n. 1 (1867), = Heteroscelis
Servillet, de Cast.
chrysochlora, Walk. loc. cit. iii, p. 528 (1868), = Heteroscelis Servillet,
de Cast.
indecora, Walk. loc. cit.,= Heteroscelis Servillei, de Cast.
—— scutellaris, Walk. loc. cit. p. 529, = Heteroscelis lepida, Stal.
Oplomus biarcuatus, Walk. loc. cit. p. 121. n. 17,= Oplomus rutilus, Dall.
chrysomelas, Walk. loc. cit. n. 18,= Oplomus tripustulatus, Fabr.
basalis, Walk. loc. crt. p. 122. n. 19,= Oplomus tripustulatus, Fabr.
Platynopus conspersus, Walk. loc. cit. p. 128. n. 38,= Oplomus proteus,
Stal.
trijunctus, Walk. loc. cit. p. 125. n. 11,= Platynopus rostratus, Dru.,
var.
badius, Walk. loc. cit. n. 12,= Platynopus rostratus, Dru.
optabilis, Walk. loc. cit, p. 126, n. 13,= Platynopus rostratus, Dru.,
var.
polygraphus, Walk. loc. cit. n. 17,= Platynopus melanoleucus, Westw.
tenellus, Walk. loc. cit. p. 127. n. 18,=Platynopus melacanthus,
Boisd.
dotatus, Walk. loc. cit. p. 128. n. 20,= Platynopus letus, Walk.
semiscitus, Walk. loc. cit. p. 129. n. 21, =Platynopus letus, Walk.
Picromerus nigrivitta, Walk. loc. cit. p. 133. n. 7,= Picromerus obtusus,
Walk.
Arma monospila, Walk. loc. cit. p. 186. n. 16,= Podisus sagitta, Fabr.
colorata, Walk. loc. cit. nn. 17,= Euthyrhynchus floridanus, Linn.
lateralis, Walk. loc. cit. p. 188. n, 36, = Tynacantha cincticeps, Stal.
Gilva varipes, Walk. loc. cit.1. p. 142. n. 1 (1867), = Coryzorhaphis spinole,
Sign.
Asopus binotatus, Walk. Joc. cit. p. 144. n. 2, = Cermatulus nasalis,
Westw.
Piezosternum retractum, Walk. Joc. cit. iii. p. 457 (1868),= Prezosternum
subulatum, Thunb.
eyscpiaty trucidus, Walk. loc. cit. p. 461. n. 8,= Pygoplutys cribatus,
alk.
Tesseratoma striata, Walk. loc. cit. p. 463, n. 16,= Tessaratoma papillosa,
Dru.
— furcifera, Walk. loc. cit. n, 17,= Tessaratoma paptllosa, Dru.
64 Mr. H. Druce on some
Tesseratoma timorensis, Walk. loc. cit. p. 464. n. 18,= Tessaratoma
papillosa, Dru.
clara, Walk. loc. cit. n. 19,= Tessaratoma papillosa, Dru.
senucuprea, Walk. loc. cit. p. 465. n. 21,= Hypencha apicalis, Lep. &
Serv.
Salica excellens, Walk. loc. cit. p. 469. n. 1,=Piexosternum subulatum,
Thunb.
Pycanum rubidum, Walk. loc. cit. p. 471. n. 11,=Pycanum pretiosum,
Stal.
ea laos Walk. Joc. cit. p. 472. n. 18,=Carpona angulata,
tal.
pallipes, Walk. loc. cit. p. 473. n. 17,= Mattiphus oblongus, Dall.
Daleantha regia, Walk, loc. cit. p. 474. n. 3,= Dalcantha dilatata, Amy. &
Serv.
To be treated as non-existent.
Species the types of which are not now to be found in the British Museum.
Arma velata, Walk. Cat. Het. iii. p. 5382 (1868).
Species incorrectly placed in the Asopine.
Genus CARBULA (Pentatomine).
Carbula humerigera.
Pentatoma humerigera, Uhler, Proc. Ac. Philad. 1860, p. 223. n. 4.
Arma japonica, Walk. Cat. Het. iii. p. 583 (1868).
Genus Tyoma (Pentatomine).
Tyoma cryptorhyncha.
Cimex cryptorhynchus, Germ. in Silberm. Rey. y. p. 169 (1837).
Arma nanula, Walk. Cat. Het. i. p. 139. n. 38 (1867).
VIII.—Descriptions of some new Species of Heterocera from
Tropical South America. By Herpert Druce, F.L.S.
&e.
Fam. Syntomide.
Dycladia lydia, sp. n.
Female.—Head, palpi, antennz, thorax, abdomen, and legs
black ; collar and base of the abdomen chrome-yellow; the
anal segments of the abdomen dark blue ; a fine yellow line on
each side of the abdomen extending from the base almost to
the anus. Primaries black, crossed beyond the middle from
new Species of [Heterocera. é 65
the costal to the outer margin bya curved semiliyaline yellow
band: secondaries black, with a fine hyaline line from the
base to the middle of the outer margin.
Expanse | inch.
Hab. South Brazil, Porto Real (Mus. Druce).
Hucereon giganteum, sp. n.
Male.—HUead, palpi, and antenne black, collar bright
carmine ; tegule and thorax black, streaked with yellowish
brown ; a large yellow spot at the base of thorax; abdomen
black, banded with bright carmine, the anal segments bright
carmine above, black on the underside; the legs black.
Primaries brownish black, with a brownish-white spot
at. the end of the cell; a band very similar in colour be-
yond the cell; a submarginal pale brownish line extending
from the apex to the anal angle; the veins all pale yellowish
brown: secondaries white, broadly bordered with black, the
veins black.
Hxpanse 24 inches.
Hab. Colombia (Mus. Druce).
Eucereon testaceum, sp. n.
Male.—Wead, palpi, collar, tegule, thorax, abdomen, and
legs grey; antenne black. Primaries semihyaline white
beyond the cell, the apex, outer and inner margin clouded
with grey: secondaries semihyaline white.—The femule
almost identical with the male, excepting that the base of the
thorax is clothed with white hairs.
Exxpanse, ¢ 14, 2 12 inch.
Hab. Venezuela (Mus. Druce).
Eucereon (?) fanum, sp. n.
Male,-—Head, palpi, antenne, tegule, and thorax black,
the head and base of the tegulez spotted with white ; abdomen
bluish black, with a row of white spots on each side; legs
black above, white on the underside; the underside of the
thorax yellow. Primaries black, with a large greyish patch
round the end of the cell, the costal and inner margin
slightly streaked with grey: secondaries black, the central
part of the wing from the base to the end of the cell semi-
hyaline white, the veins black.
Expanse 2 inches.
Hab. Venezuela (Mus. Druce).
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 5
66 On some new Species of Heterocera.
Fam. Arctiidae.
Automolis melea, sp. u.
Male.—Front of head blue; palpi black above, chrome-
yellow on the underside; antenne black; collar and tegule
chrome-yellow ; thorax and basal half of the abdomen black,
with a large yellow spot on each side close to the base; the
anal segments of the abdomen spotted with metallic blue ; the
underside black, banded with chrome-yellow; legs black.
Primaries black, the costal margin from the base to the end
of the cell edged with chrome-yellow ; a wide chrome-yellow
band extends from the base of the wing to the outer margin,
which it does not quite reach; the fringe black: secondaries
black, the costal margin from the base to the apex chrome-
yellow ; the fringe black.—Female very similar to the male,
but larger and without the chrome-yellow spots near the base
of the abdomen.
Expanse, ¢ 13, 2 2 inches.
Hab. Venezuela (Mus. Druce).
Automolis Pratti, sp. n.
Male.—Head, palpi, and antenne black, head spotted with
metallic blue in front; tegule yellow, edged with black on
the outer side; thorax black, metallic blue at the base, with
a yellow spot on each side; abdomen black, the anal segments
spotted with metallic blue; legs black. Primaries dark
yellow, the costal margin, apex, outer and inner margin edged
with black: secondaries black, the costal margin from the
base to the apex broadly yellow.
Expanse 1° inch.
Hab. Colombia (Mus. Druce).
Amaia theon, sp. n.
Male.—Head and collar yellow; antenne brown; palpi
brown, white on the underside; tegule brown, edged with
yellow; thorax yellow; abdomen dark brown above, white on
the underside, the anal segment pale yellow. Primaries semi-
hyaline pale yellow, spotted and banded with brown, very
similar to those of Amawxia pardalis, Walker, but with more
yellow at the base of the wing: secondaries semihyaline
white, shaded with pale brown at the anal angle and round
the outer margin.
Expanse 1,°5 inch.
Hab. Venezuela (Mus. Druce).
Mr. W. F. Kirby on Odonata from Sterra Leone. 67
Idalus pythia, sp. n.
Male.—Head, collar, and tegule pale greyish fawn-colout ;
antenne fawn-colour, the tips white; abdomen yellow, the
underside white, the anal segment pale greyish fawun-colout ;
legs white. Primaries semihyaline yellowish white, the
basal half of the wing pale greyish fawn-colour, with several
yellowish-brown spots along the inner margin ; a large, round,
pale greyish fawn-coloured spot with an indistinct darker
brown centre close to the apex: secondaries semihyaline
yellowish white, dark yellow alony the inner margin.
Expanse 14 inch.
Hab. Bolivia, Songo (Mus. Druce).
Opharus amata, sp. n.
Female.—Head, antenne, collar, tegule, thorax, abdomen,
and legs black; abdomen banded with yellow. Primaries
olive-brown, palest at the base and along the costal margin,
the veins black; a rather large indistinct blackish spot at the
end of the cell: secondaries semihyaline brownish black, the
fringe dark brown.
Expanse 2 inches.
Hab. Venezuela (Mus. Druce).
IX.—Report on the Neuroptera Odonata collected by Mr. E. E.
Austen at Sterra Leone during August and September 1899,
By W. F. Kirsy, F.L.8., F.E.S.
[Plate II.]
Durine Mr. Austen’s visit to Sierra Leone, as a member of
Major Ross’s expedition to inquire into the connexion between
mosquitos and malaria, he collected a considerable number of
dragonflies; but as he captured long series whenever he was
able, the number of distinct species obtained amounted only
toeighteen. However, I am able to describe seven new species
in the present paper, two of which I have made the types of
new genera. Ishould also mention that specimens of most of
these have been in the Museum for some time, but have not
previously been described.
In place of quoting full synonymy in the present paper, I
generally confine myself to quoting references to ae original
*
68 Mr. W. F. Kirby on Odonata from Sierra Leone.
author of a species and to my ‘Catalogue of Neuroptera
Odonata.’
The only other Neuropt
specimens of Myrmeleon tri
1899, and one or two sma
era in the collection were four
stis, Walker, taken on Sept. 5,
ll species of Ephemeride and
Termitide not at present determinable.
The following species were obtained by Mr. Austen =
Libellulide.
LIBELLULINZ.
Pantala, Hag.
flavescens, Fabr.
Rhyothemis, Hag.
notata, Fabr.
Palpopleura, Ramb.
Lucia, Drury.
Portia, Drury.
Trithemis, Brauer.
Kalula, Kirb. (sp. n.).
serva, Kirb. (sp. n.).
Helothemis, Karsch.
dorsalis, Ramb.
Beblecia, Kirb. (g. 0.).
adolescens, Kirb. (sp. N.).
Crocothemis, Brauer.
erythrea, Brullé.
Thermorthemis, Kirb.
Austeni, Kirb. (sp. n.).
leonina, Karsch.
Apeleutherus, Kirb. (g. n.).
Strachant, Kirb. (sp. n.).
Orthetrum, Newm.
chrysostigma, Burm.
Julia, Kirb. (sp. n.).
CorDULIIN”.
Macromia, Ramb.
Selysi, Kirb. (sp. n.).
JEschnide.
/ESCHNINA.
Anacieschna, De Selys.
triangulifera, McLachl,
Agrionide.
AGRIONINE:,
Sapho, De Selys.
ciliata, Fabr.
Libellago, De Selys.
dispar, Beauv.
Pantala flavescens.
Libellula flavescens, Fabr. Ent. Syst. Suppl. p. 285 (1798).
Pantala flavescens, Kirb, Cat. Neur, Odon. p. 1. n. 1 (1890).
9 specimens, Aug. 25, Sept. 18, 21, 27 (1899).
Rhyothemis notata.
Libellula notata, Fabr. Spee. Ins. i. p. 390. n. 12 (1781).
Rhyothemis notata, Kirb. Cat. Neur, Odon. p. 6, n, 21 (1890).
3 specimens, Aug. 26, Sept. 2 (1899).
Palpopleura Lucia.
Libellula Lucia, Drury, Il. Exot. Ent. ii. pl. xlv. fig. 1 (1778).
Palpopleura Lucia, Kirb. Cat. Neur. Odon. p. 9. n. 1 (1890),
35 specimens, Aug. 21 to Sept. 22 (1899).
One of the commonest and most widely distributed African
dragonflies.
Mr. W. F. Kirby on Odonata from Sierra Leone. 69
Palpopleura Portia.
Libellula Portia, Drury, Ill. Exot. Ent, ii. pl. xlvii. fig. 3 (1778).
Palpopleura Portia, Kirb. Cat. Neur. Odon. p. 9. n. 4 (1890).
9D specimens, Aug. 26 to Sept. 21 (1899).
Although long known and widely distributed, this species
does not seem to be nearly so abundant as P. Lucia, Drury.
Trithemis Kalula, sp.n. (PI. II. figs. 2, 2 a.)
Long. corp. 82-34 millim. ; exp. al. 53-55 millim.; long.
pter. 2 millim.
Male.—Rufo-testaceous, with hyaline wings. Very similar
to T. arteriosa, Burm., from which it differs as follows :—
The segments of the abdomen are distinctly shorter and
broader, and instead of a row of lateral black spots succeeded
by a continuous band, there is a row of lateral black spots
enlarging and expanding behind, which are more or less
expanded on the last three segments, sometimes so much as
to cover most of these segments, except a reddish spot on each
side in front. The middle appendages of the same segment
consist of a pair of long slender recurved hooks. The neura-
tion is brown, reddish only sometimes in the costal and _post-
costal areas. The yellow shade at the base of the hind wings is
darker than in 7’, arteriosa and the cells in it are usually centred
with pale brown. In adult specimens the vertex is violet, and
the thorax and abdomen are slightly pruinose, but more of a
reddish plum-colour than blue. An immature male (and pre-
sumably the female) has a broad brown median band on the
thorax, and a narrower one on each side, the latter connected
below with a longitudinal black band on the pleura, which
emits three oblique brown bands downward, the hindermost
connected with a black space, filled up with two pale spots in
front and a large transverse one behind on the metapectus.
At the base of the abdomen there are also three brown stripes
above the level of the lateral markings already described. In
the more mature specimens these markings are more or less
obscured. Legs black. Pterostigma brown, more or less
centred with yellow.
4 specimens, Aug. 26, Sept. 13, 21 (1899).
Trithemis serva, sp.n. (Woodeut, fig. 1.)
Long. corp. 34 millim.; exp. al. 57 millim.; long. pter.
23 millim.
Male.—Dull black, more or less varied with testaceous in
immature specimens, especially on the face; in the most
70 =Mr. W. F. Kirby on Odonata from Sierra Leone.
mature specimens the vertex is violet and the thorax pruinose
blue. The sides of the mentum are sometimes yellow.
Wings hyaline, with black nervures; fore wings with 11-13
antenodal and 7-10 postnodal ecross-nervures ; pterostigma
dark brown, traversed by a narrow yellow line. Triangle
traversed, followed by three rows of cells, increasing; sub-
triangular space consisting of 3 cells: hind wings with 7-8
shin Have Bali: aL Wine se
a seg
g5 Shi
Trithemis serva.
antenodal and 9-11 postnodal cross-nervures ; membranule
white, stained with blackish; a small yellow cloud beyond,
sometimes obsolete, with its outer border straight, and above
the lower basal cell, only visible at the extreme base. Ap-
pendages of the second segment very prominent, the two
hindermost converging like a pair of pineers.
Described from six specimens, two of them taken by
Mr. Austen on Sept. 5 and 21 (1899).
Allied to the Kast-Indian 7. fest/va, Ramb., but easily
distinguished by the different shape of the markings at the
base of the hind wings and the form of the appendages of the
second segment of the abdomen. It must also be allied to
1’. dichroa, Karsch, which seems to be a still darker coloured
species in the male.
Helothemis dorsalis.
Libellula dorsalis, Ramb. Ins. Névr. p. 89 (1842).
Trithemis dorsalis, Kirb. Cat. Neur. Odon, p. 19. n. 16 (1890),
Helothemis dorsalis, Karsch, Berl. ent. Zeitschr, xxxili. p. 878 (1890).
10 specimens, Aug. 19, 26, Sept. 8, 9, 138, 19, 20, 21 (1899).
I am not satisfied that this genus is sufficiently distinct
from Trithemis.
Mr. W. F. Kirby on Odonata from Sierra Leone. 71
BEBLECIA, gen. nov.
Eyes connected by a short space; frontal tubercle rounded
above, and slightly concave in front; abdomen with segments
2 and 3 carinated. Wings with only one nervure in the
lower basal cell and no supratriangular nervures: fore wings
with 12-13 antenodal cross-nervures, the last not continuous,
and 10-11 postnodals; pterostigma long and thick, nodal
sector slightly waved beyond the middle, and with a double
row of cells between it and the subnodal sector for the last
third or fourth of their length; arculus rising between the
first and second antenodal cross-neryures; sectors of the
arculus stalked; triangle traversed by 1 or 2 nervures and
followed by a row of 4 cells, then by 2 or 3 more or less
irregularly, before rising again to 4 increasing, or by seven
rows of 3 only; subtriangular space consisting of 3 or 4 cells:
hind wings with 10 antenodal and 11 postnodal cross-
nervures ; triangle traversed by a nervure or (rarely) free ;
sectors of the arculus widely separated, the upper one rising
considerably above the lower angle of the triangle, where the
lower sector rises ; base of triangle nearly corresponding with
the arculus. Claws dentated beyond the middle. Appendages
of the second segment in the male rather large; upper
terminal appendages rather long and slender, fully as long as
the ninth segment ; lower appendage nearly as long as the
upper ones, truncated at the extremity.
This genus appears to be nearly allied to Trdthemis, but in
typical Trithemis the sectors of the arculus are scarcely
separated in the hind wings and the triangle of the hind wings
is never divided.
It is to be regretted that Mr. Austen obtained only three
specimens of this interesting species, all in bad condition.
Beblecia adolescens, sp. n. (Woodcut, fig. 2.)
Exp. al. 60 millim.; long. pter. 5 millim.
Lighter or darker ochraceous; pleura, pectus, and face
paler; vertex darker, sometimes slightly greenish; antenne,
ocelli, spines on the legs, and central and part of the lateral
caring on the abdomen black; a pale brown shoulder-stripe
and a similar stripe cn each side of the upper part of the
abdomen: wings hyaline, with black neuration; costal
nervure and pterostigma yellow, the latter between black
nervures ; membranule of hind wings whitish, beyond it the
wing is very slightly stained with yellow.
72 My. W. F. Kirby on Odonata from Sierra Leone.
Described from three specimens (2 ¢,1 ¢), all more or
less damaged and perhaps not quite mature, captured on
Sept. 2, 9, and 13 (1899).
Beblecia adolescens.
Crocothemis erythraa.
Libelluda erythrea, Brullé, Expéd. de la Morée, iii. (1) p. 102, pl. xxxii,
fic. 4 (1832).
Crocothemis erythrea, Kirb. Cat. Neur. Odon. p. 21. n, 1 (1890).
19 specimens, Aug. 25 to Sept. 22 (1899).
Specimens from West Africa are rather smaller and
slenderer than typical ones from South Kurope.
Thermorthemis Austent, sp.n. (Pl. IL. figs. 1, 1 @.)
Long. corp. 54-56 millim.; exp. al. 94-98 millim.; long.
ter. 4 millim.
Male.—Rufo-testaceous, thorax with a slight brown line on
each shoulder and the borders of the patagia black ; abdomen
in the male pulverulent blue. Legs black in the adult male,
except at the base of the femora; otherwise rufo-testaceous,
with black tarsi; spines strongly developed. Wings hyaline,
with black nervures, very slightly tinged with yellowish
brown at the base; pterostigma yellow, between black ner-
vures; tips very slightly browned. Fore wings with 20-23
antenodal (the Jast continuous) and 13-14 postnodal cross-
nervures; triangle divided by two nervures; two supra-
triangular nervures ; triangle followed by four rows of cells,
increasing subtriangular space consisting of 5-8 cells : hind
wings with 13-15 antenodal and 13-16 postnodal cross-
nervures; triangle traversed and with the base generally
Mr. W. F. Kirby on Odonata from Sierra Leone. 73
placed a little nearer the base of the wings than the arculus ;
one supratriangular nervure; sectors of the arculus slightly
separated, the lower onecoinciding with the baseof the triangle.
Abdomen with strong black median and lateral serrated carine ;
terminal appendages of male as long as the ninth segment,
the lower appendage broad, half as long as the others.
Female.—Uniform rufo-testaceous, mouth-parts inclining
to yellowish, dark shoulder-stripe slightly indicated, carina of
abdomen black, eighth segment perfoliate.
This species agrees with YZ. madagascariensis, Ramb., in
general neuration, the bifid frontal tubercle, the Orthetrum-
like frontal depression, &c., but differs in the fewer and
stronger spines on the hind tibiee.
Described from five male specimens, one taken by
Mr. Austen at Sierra Leone on Sept. 21, 1899, and the others
from West Africa, without special locality.
The single female is from Sierra Leone (Morgan).
I append the description of an allied species from Angola* :—
Female.—Dark brown; head testaceous; under surface and
terminal segments inclining to ferruginous ; eighth segment
perfoliate, with the edges black. Wings yellowish hyaline,
with black nervures: fore wings with 19 antenodal and 12
postnodal cross-nervures; neuration otherwise asin 7. Austen’,
One specimen from Angola, collected by Mr. and Mrs,
Monteiro.
Thermorthemis leonina.
Orthetrum leoninum, Karsch, Ent. Nachr. xvii. p. 59 (1891).
Two specimens (¢ ¢ ) from Sierra Leone, the male taken
by Mr. Austen on Sept. 2, 1899, and the female taken by
Dr. W. G. Clements.
The insect is a true T’hermorthemis, except that the body is
more slender, and the hind tibie have fewer and stronger
spines than in 7’. madagascariensis; but it shares the latter
character with 7. Austenz. In the male the triangle of the
fore wings is followed by four rows of cells increasing, and in
the female by five, shrinking to four and then again increasing.
APELEUTHERUS, gen. noy.
Male.—Frontal tubercle strongly bifid; front large and
broad, projecting beneath it to about half the width of the
eye, almost quadrangular, with a conspicuous median depres-
sion. Kyes connected for a short space. Legs rather long,
* Thermorthemis Monteirot, sp. n.
74 Mr. W. F. Kirby on Odonata from Sierra Leone.
spines scarcely more than sete on the femora, numerous and
rather short and strong on the tibie, but with a few long
slender ones at their base. Abdomen distinctly shorter than
the hind wings, not inflated at base; second and third
segments transversely carinated; appendages of second
segment prominent, consisting of a small lobe, and an ap-
pendage in front directed obliquely backward; upper anal
appendages rather longer than the ninth segment, slender,
hardly longer than the broad lower appendage. Wings with
only one cross-nervure in the lower basal cell and no supra-
{riangular nervures; nodal sector slightly waved; arculus
rising between antenodals 1 and 2, its sectors with a long
stalk. Fore wings with 13-14 antenodal cross-nervures, the
last not continuous, and 8—9 postnodal cross-nervures ; triangle
crossed by 2 nervures (rarely 1) and followed by one or two
rows of 4 cells (rarely 3) and then several rows of 3, in-
creasing; subtriangular space consisting of 3 cells (occasionally
2 or 4 on one side). Hind wings with 8-10 antenodal and
9-11 postnodal cross-nervures; triangle traversed by one
nervure, its base lying rather nearer the base of the wings
than the arculus; sectors of the triangle subcontiguous ;
triangle (above the sectors) followed by one row of 3 (rarely 2)
cells, and then by several rows of 2, increasing.
Female.—Eighth segment not expanded; vulvar scale
distinct, about one third as long as the ninth segment.
Allied to Thermorthemis, Hadrothemis, &c. The presence
of the vulvar scale in the female allies it to Crocothemis.
Apeleutherus Strachani, sp.n. (PI. I. figs. 4, 4 a.)
Long. corp. 38-40 millim. ; exp. al. 71-73 millim.; long.
pter. 4 millim.
Head blackish, face more or less testaceous, especially the
labrum and mentum. Thorax brown or blackish; a large
oval spot on each shoulder, a longitudinal stripe on each side
of the central carina in front, a transverse one behind, in
front of the patagia, the inside of the patagia, 3 spots on the
upper part of the pleura and others below, and some slighter
marks on the interalary spaces pale yellow, or more rarely dull
reddish; abdomen greyish brown above (rarely red in the
male), but usually reddish below. In the female the base
and sides are marked with irregular pale markings and with
large pale irregularly-shaped spots, more or less distinct
above, on each side of most of the segments, near the base.
In the male these markings are usually obsolete. Legs
testaceous, more or less blackish above, and with black spines.
Mr. W. F. Kirby on Odonata from Sterra Leone. 75
Wings hyaline, with black nervures; pterostigma rather
long and broad, blackish, rather paler in the middle and
towards its lower edge. Membranule of hind wings whitish
in the middle and bordered with brown: adjacent part of
wing slightly stained with yellow in the reddest males;
otherwise this is wholly wanting. Anal appendages yellowish
or reddish.
Described from sixteen specimens (seven males and nine
females) collected by Dr. Strachan at Lagos. Only one of
the males has the abdomen red and the pale abdominal
markings obliterated, but the markings of the thorax are of
the usual pale yellowish colour seen in the other specimens.
Mr. Austen collected two males at Sierra Leone on Aug. 24
and Sept. 4; in both of these the abdomen is red, but in one
the pale abdominal spots are indistinctly visible. In both
the spots on the thorax are dull red, which is not the case in
any of Dr. Strachan’s specimens. ‘There is also an unusually
well-marked female, with only one cross-nervure in the
triangles of the fore wings, from Abyssinia, which appears to
belong to the present species.
Among other interesting species obtained by Dr. Strachan
at Lagos is a male belonging to the genus Dicranopyga,
Karsch, which is allied to the Hast-African D. mundula,
Karsch. I believe that Dr. Karsch is correct in referring my
Aithrimanta rezia from Madagascar to Dicranopyga, although
the triangle of the fore wings is traversed by a triangle on
the right side in the unique type. Dr. Strachan also obtained
a single damaged specimen of a new species of Nympheutria,
Karsch, which I deter describing for the present.
Orthetrum chrysostigma.
Libellula chrysostigma, Burm. Handb. Ent. ii. p. 857, n. 58 (1839).
Libellula barbara, De Selys, Lucas, Expl. Alg. iii. p. 117, pl. 1. figs. 2,
2a, b (1849).
Orthetrum chrysostigma et barbarum, Kirb, Cat. Neur. Odon. p. 36,
nn. 3, 4 (1890).
45 specimens, August and September (1899).
The most adult specimens are pulverulent blue in both
sexes, and the least mature are of a light red; but there is no
median thoracic band, as in VU. Julia, and the cross-nervures
in the lower costal space are nearly always yellow, instead of
black like those in the upper space.
Orthetrum Julia, sp.n. (Pl. IL. fig. 3.)
Long. corp. 89-48 millim.; exp. al. 60-74 millim. ; long.
pter. 3-4 millim,
76 Mr. W. F. Kirby on Odonata from Sierra Leone.
Male.—Head with the vertex, the strongly bifid frontal
tubercle, the occipital triangle, extremity of labrum, and a
broad central band on the labrum black ; hinder orbits yellow,
with one or two brown transverse spots; frontal shield
greenish, strongly punctured ; the rest of the face (except fo
the black markings already described) yellowish. Prothorax
yellow, with a brown transverse band, narrowed in the middle.
Mesonotum yellow, with brown and blackish markings; the
front transversely black, with a transverse yellow mark in the
middle ; a median dark central band, with two yellow lines
near the centre and a dark shoulder-stripe, not reaching the
hinder lappets, which are yellow, enclosed by black carine.
Interalary spaces black, spotted with yellow in the middle, and
with a yellow spot at the base of each wing. On the yellow
pleura, below the shoulder-stripe, are five more black or
brown bands on or between the sutures, the first double above,
The metapectus is enclosed by almost a square of black
earine, the hindermost detached, and slightly curved. Legs
black; femora more or less testaceous. Abdomen black or
pruinose blue, with all the carine black, the first three
segments with yellow markings, separated by the carinze and
by dark hind borders, and a lateral stripe; the three following
segments with a broad subbifid spot on each side in the
middle, the rest black. On the under surface these pale
lateral markings are more extended. Anal appendages yellow,
upper ones slender, as long as the ninth segment, the extreme
tips black; lower appendage truncated at the extremity,
nearly as long as the upper ones, and with a black carina
beneath on each side. Appendages of the second segment
large. Wings hyaline, with black nervures; fore wings with
13-15 antenodal and 11-12 postnodal cross-nervures. Ptero-
stigma black, with a yellow transverse central line. Mem-
branule blackish, with a white spot at the base: hind wings
smoky yellow at the extreme base, the sectors of the arculus
subcontiguous at the base.
Female similar, with the lines on the pleura darker and
broader. Abdomen with the testaceous spot on the under
surface of the seventh segment expanded to the sides far
enough to be visible above; eighth segment pertoliate.
Anal appendages yellow, longer than the tenth segment, which
isalso yellow. Wings clouded hyaline, with black nervures ;
14-17 antenodal and 11 postnodal cross-nervures.
Abdomen similar in both sexes, much inflated at base,
slightly constricted behind the third segment; the rest of the
segments rather broad; the fourth about three times as long
as broad, widening at the extremity ; the rest subparallel,
Mr. W. F. Kirby on Odonata from Sierra Leone. 77
but successively shorter. Costal nervure black, with a
barely perceptible trace of an intersecting yellow line at the
base.
In the most adult males, which I was at first inclined to
regard as a distinct species, the body is almost entirely
pruinose blue, through which a row of long tawny spots may
sometimes be seen extending as far as the eighth segment of
the abdomen. There is usually a black mark in the middle
of the labrum. ‘I'he appendages of the second seement are
apparently better defined, and the upper anal appendages are
black, though the lower appendage generally remains tawny.
The abdomen is apparently less inflated at the base, less
constricted behind the third segment, and somewhat more
slender. Yet I find it impossible to venture to separate these
specimens specifically.
The species is described from four specimens from Sierra
Leone, two of which were collected by Mr. Austen on Aug. 26
and Sept. 11 (1899).
Of the adult form nine males were collected by Mr. Austen
between Aug. 26 and Sept. 15. There are also specimens of
both forms in the Museum from Accra and Lagos.
This is a species of great interest. The semiadult form,
which I have taken as typical, closely resembles O. Sabina,
Drury, except in the shorter and broader abdomen. In fact,
O. Julia stands in almost the same relation to O. Sabina in
the Old World as that occupied by Mesothemis attala, De Selys,
to M. verbenata, Hagen, in the New. I have not seen the
true O. Sabina from Africa, though De Selys indicates a var.
africana from the Camaroons (Ann. Soc. Hnt. Belg. xxxi.
p- 22, 1887). The insect is also very near O. trinacria,
De Selys, and is, I imagine, regarded by some authors as
this species, or as O. brachiale, Beauv. But I have not been
able to determine O. brachiale and O. africanum, Beauv.,
satisfactorily. According to Prof. Calvert, who has examined
the types, the former should have the base of the wings
entirely unclouded, and A. annulatum is described as having
the thorax uniform brown. As regards O. trinacria, it is a
long-bodied insect much resembling O. Sabina.
Macromia Selysi, sp. n.
Long. corp. 67 millim.; exp. al. 88 millim.; long. pter.
24 millim.
Male.—Head: frontal tubercle bifid and, as well as the
frontal lobes below, with strong green and violet reflections ;
face lighter or darker ferruginous brown, with orange mottlings.
78 = =Mr. W. F. Kirby on Odonata from Sierra Leone.
Thorax and pleura shining metallic green; thorax and base
of abdomen clothed with grey hairs. Pleural sutures and
interalary spaces with slight pale markings; abdomen black,
with narrow pale belts in the middle of the second, third, and
fourth segments, and near the front of the second and sixth
segments, the last the broadest.
Wings rather long, narrow, and pointed, clear hyaline, with
black nervures. Fore wings with 15-16 antenodal and 9
postnodal cross-nervures; the nodal and subnodal nervures
nearly straight and parallel for most of their length; triangles
small, free, followed by a cell divided by a curved line
running from the triangle, and then by several single or
irregularly divided cells, followed by several rows of two,
increasing. Only 3 supratriangular nervures; dcross-nervures
in the lower basal cell. Pterostigma deep black ; membranule
dark brown, intersected by a pale line. Hind wings with 11
antenodal and 11-12 cross-nervures ; 2 supratriangular ner-
vures and cross-nervures, 4 in the lower basal cell; no lower
triangle ; upper triangle free, followed by a triangular cell,
then by 3 single oblong ones, and then by two rows of
cells, increasing. Membranule black, with a white speck at
the base and tip; anal triangle divided low down, the lower
part forming a long narrow isosceles triangle (in J. Sophia,
De Selys, it is almost equilateral), and the nervure bounding
the anal angle is more distinctly marked with red. Anal ap-
pendages nearly as in M. Sophia.
One male taken by Mr. Austen on Sept. 16 (1899).
This species is closely allied to M. Sophia, De Selys, but
differs in the clear hyaline wings and many little details of
neuration, especially by the commencing single row of post-
triangular cells on all the wings.
Anacieschna triangulifera.
Anacieschna triangulifera, McLachlan, Ann. & Mag. Nat. Hist. (6)
xvii. p. 407 (1896).
Sept. 9 (1899).
A single damaged specimen, which I refer to this Hast-
African species (Delagoa Bay and Natal) with some un-
certainty.
Sapho ciliata.
Agrion ciliata, Fabricius, Spec. Ins. i. p. 528. n. 3 (1781).
Sapho ciliata, Kirb, Cat. Neur. Odon. p. 100. n. 1 (1890).
34 specimens, Freetown, Sierra Leone, Sept. (1899).
A very interesting series. ‘The immature specimens of
Mr. M. Burr—WNotes on the Forficularia. To
both sexes are of a clear iridescent hyaline, darkening to
yellowish hyaline in the female, and in the male passing on
through shades of brown to the deep purple, strongly suffused
on the hind wings with coppery red, which characterizes the
adult male. Fabricius’s type was an adult female.
Libellago dispar.
Agrion dispar, Beauv. Ins. Afr, Amér. p. 85, Neur. pl. vii. fig. 2 (1805 ?).
Libellago dispar, Kirb. Cat. Neur. Odon. p. 112. n. 1 (1890).
Two specimens, Sept. 13 and 21 (1899).
EXPLANATION OF PLATE II.
Fy. 1. Thermorthemis Austeni, 9 , p. 72.
Fig. 1a. Ditto, 3, appendages of second segment.
Fg. 2. Trithemis Kalula, 3, p. 69.
Fig. 2a. Ditto, 3, appendages of second segment.
Fig. 3. Orthetrum Julia, 3, p. 75.
Fig. 4. Apeleutherus Strachani, 2, p. 74.
Fig. 4a. Ditto, 3, appendages of second segment.
X.—WNotes on the Forficulariaa—V. Descriptions of new
Species and a new Genus. By Matcoim Borr, F.Z.S.,
¥.E.S.
[Plate IV. figs. 3, 5, 7.]
Anechura ahrimanes, sp.n. (PI. IV. fig. 5.)
Statura majore; colore nigro, rubro-variegato ; elytra et ale per-
fecte explicate ; abdomen medio paullo dilatatum, ovale ; forcipis
brachia elongata, gracilia, subsinuata, intus fortiter bidentata. ¢.
© ignota.
Long. corporis...... 12-145 mm.
ORC UBIE ers 2.9 « 9 =
Head deep red; eyes black; sutures indistinct; antenne
dark red (seven segments remain).
Pronotum black, broader than the head ; anterior margin
straight, posterior margin rounded; the sides parallel.
Elytra broad, finely punctulated, deep red, shaded with
black towards the apex and the margins.
Wings prominent, black, with a large yellow spot near the
base.
80 Mr. M. Burr—WNotes on the Forficularia.
Feet deep red, the knees and tibie darker or black ; tarsi
darker.
Abdomen deep dark red, somewhat dilated in the middle,
narrowed towards the extremity ; lateral tubercles black, very
distinct; the whole abdomen is finely punctulated; last
segment transverse, hinder margin straight, a faint median
depression, a reddish blunt elevation over the insertion of the
forceps, and a small tubercle on the outside angles ; penulti-
mate ventral segment ample, rounded.
Pygidium strongly depressed, triangular; two small pale
obtuse lobes are visible below the apex of the pygidium,
projecting beyond the border of the produced penultimate
segment, visible from below.
Forceps & with the branches slender, long, remote at the
base and tricarinate there, slightly diverging at first, then
subsinuate until the apices meet and cross; there is at the end
of the first and second third of the total length a strong trian-
gular depressed tooth: the forceps are red at the base, darker
towards the apex; the inner margin of the basal third is
finely crenulated.
Patria. North India, Sikkim, 2 ¢.
Type in my collection.
This is a very handsome and distinct species. It appears
to be most closely allied to A. Hugeli, Dohrn, from Luzon,
but is coloured very differently.
STRONGYLOPSALIS *, gen. nov.
Corpus convexum : antenn segmentis 1° et 3° longis, ceteris brevi-
oribus, 4° et sequentibus conicis: pronotum quadratum: elytra
perfecte explicata; ale nulle: abdomen medio modice dilatatum,
apicem versus ¢ minus, 9 maxime attenuatum; segmentis 2° et
3° dorsalibus tuberculis pliciformibus distinctis instructis: tar-
sorum segmentum secundum simplex, cylindricum: forcipis
brachia ¢ basi remota, gracilia, basi recta, deinde valde incurva,
asymmetrica ; 9? contigua, recta, apice decussata.
This genus stands nearest to Carcinophora, Scudd., from
which it may be distinguished by the slender forceps, quite
remote at the base, and not stout and subcontinuous as in
that genus. It has the appearance of certain species of
Chelidura, Latr., but is easily separated by the simple second
tarsal segment. From Anzsolabis, Fieb., it may be distin-
guished by the fully developed elytra and the presence of
tubercles on the second and third abdominal segments.
* grpoyyvros, circle; Wadis, forceps.
Mr. M. Burr— Notes on the Forficularia. 81
Strongylopsalis inca, sp. n.
Statura minore; colore fusco ; antennw ?-segmentate (segmenta 11
restant), segmento primo et tertio elongatis, secundo brevi, quarto
et sequentibus brevibus, conicis; pronotum quadratum, quam
caput tam latum, pallido-marginatum ; elytra perfecte explicata,
marginata, postice oblique truncata, metanotum liberantia; ale
nulle ; pedes fusco-testacei: abdomen convexum, medio modice
dilatatum, apicem versus, ¢ modice, 9 maxime attenuatum;
segmentum ultimum dorsale g magnum, medio sulcatum, utrinque
tuberculatum, margine postico rectum; 2 parvum, medio sul-
catum, vix tuberculatum: pygidium ¢ quadratum, apice trunca-
tum: forcipis brachia, ¢ basi valde remota, gracilia, inermia,
basi ipso recta, dimidia parte apicali valde incurva, mucronibus
haud attingentibus, brachio dextro intus, magis, brachio sinistro
extus, minus incurvis; 2 recta, contigua, apice decussata,
CIF -
Long. corporis.... 85-9°5 mm. & mae
ee TORCIDIS vis 2 Fe Lie 2
Head reddish brown, the eyes small and black ; the mouth-
parts somewhat paler. The antenne are darkish testaceous ;
eleven segments remain, these are all small and conical,
except the first and third, which are long, and the second,
which is very short and cylindrical.
Pronotum square, as broad as the head, testaceous, the
margins lateral, slightly raised, and paler in colour.
Elytra tully developed, testaceous, the lateral margins
slightly turned up and paler in colour; hinder margin
obliquely truncated.
Wings absent.
Feet dark testaceous.
Abdomen dark blackish brown, the centre of the segments
reddish; broadest in the middle, strongly attenuated at the
apex in the ?, less soin the ¢; tubercles of the second and
third segment present, but not very distinct. Last segment
g transverse, with a central longitudinal furrow and a
tubercle on each side thereof, the hinder border straight ; in
the ¢ the furrow is noticeable, but the tubercles scarcely so.
Forceps: in the g the branches are remote at the base,
slender, thicker at the basal part than in the apical part,
reddish in colour, straight at first, then strongly arched in-
wards, the right branch being most strongly bowed and arched
within the left; in the ? the branches are subcontiguous,
straight, the apices crossing.
Patria. Peru, 2 6,1 2, 2 nymphs.
Type in my collection.
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 6
82 Mr. M. Burr—Notes on the Forficularia.
This is a very curious species. At first glance it might be
mistaken for Chelidura, but the simple second tarsal segment
at once shows that it belongs to a different group of genera.
It might be provisionally ranged in Caretnophora, but, owing
to the very different form of the forceps, I have preferred to
erect a new and very distinct genus.
Forficula orientalis, sp.n. (PL. IV. fig. 7.)
Statura majore. JF. auriculurie, atque FP. luride vicina, a quibus
differt forma forcipis. @ ignota.
3 forcipis brachia basi valde deplanata et dilatata, hac parte mar-
gine interno crenulato, tota inermia; dehine valde divergentia,
attenuata ad duas tertias partes longitudinis; tune subito re-
curva, valde attenuata, apice fere attingentia.
Long.corporis cous teweko 15 mm.
Sap POTGL DURE Fh. kicpete sis eye A pits
A, ss NAYAK inne aes 4:5
Head reddish; antenna testaceous, 12-segmentate, the
third segment small and round, the others elongate.
Pronotum dark brown, the lateral margins clear testaceous.
Elytra and wings fully developed, of the same form and
colour as in F, auricularia.
Feet pale testaceous.
Abdomen dark reddish, the lateral tubercles very distinct ;
the segments finely granulated. The last segment is square,
the posterior angles rectangular and depressed ; the hinder
border is straight, slightly tuberculated and broken; in the
middle of the segment there are three faint depressions, arranged
in a line horigontally ; the abdomen is broadest about seg-
ments 5-6, and then slightly narrower towards the last
segment. ‘The penultimate ventral segment nearly covers
the ultimate segment and is rounded. No pygidium visible.
Forceps: the branches are very strongly dilated in the
basal third, where the sides are at first parallel ; from the end
of the dilated part the branches diverge and are narrowed ;
in the apical third they are strongly arched inwards and
attenuated, the apices almost meeting; the space thus en-
closed between the branches is a transverse oval; the
branches are entirely unarmed, except that the inner margin
of the basal third, the dilated part, is faintly crenulated. The
colour of the forceps is black, except the centre of the dilated
part, which is testaceous.
Patria. Constantinople (ex coll. Seeldrayers), 1 g and
1 nymph.
Type in my collection.
Mr. M. Burr—WNotes on the Forficularia. 83
This species, for which I am indebted to my friend M. Seel-
drayers, of Brussels, in size and appearance resembles
Ff. auricularia and F. decipiens. The unarmed and much more
strongly dilated forceps distinguish it from F. aurteularia ;
the presence of wings and the very strongly bowed forceps
distinguish it from £. dectpiens, to which perhaps it stands
nearest ; its larger size and strongly bowed forceps separate
it from £. lurvda. ‘The general appearance of the forceps
recalls that of 2. circinata, Fin., but they are more strongly
arched and the insect is larger.
Apterygida Mackinderi, sp. n. (Pl. LV. figs. 3, 3 a.)
Statura majore ; elongata; oculis parvis, nigris ; (antenne desunt) ;
pronotum capite paullo angustius, antice rectum, postice rotun-
datum ; elytra et ale perfecte explicatie, illa latiora, apice trun-
cata; he parum prominentes: pedes breves, femoribus in-
crassatis ; tarsorum segmento secundo lobato: abdomen deplana-
tum, apicem versus paullo dilatatum ; segmentis 2° et 3° utrinque
tuberculo distincto instructis, primo parvo, secundo maximo;
segmentum ultimum dorsale magnum, quadratum, utrinque supra
insertionem forcipis tuberculatum ; segmentum penultimum ven-
trale maximum, segmentum ultimum totum obtegens, angulis
ipsis posticis exceptis, margine postico rotundato, medio late sed
haud profunde emarginato.
Forcipis crura elongata, gracilia, basi valde remota, et triquetra,
basi ipso paullo divergentia, tum sensim appropinquantia, apice
decussata, margine interno basi dente parva obtuso, denteque
valido medio armata, margine interno usque ad dentem medianum
crenulato ; pygidium magnum, breve, latissimum, inerme, margine
postico recto.
Colore fusco-testaceo, fusco-variegato. og. @Q ignota.
Long; COLPOLIS. sens 5,0 145 mm,
fOrapines 4... ae Uy GG abe
Colour generally dark brown, varied with darker; legs
aler.
2 Head darker anteriorly than posteriorly ; the eyes small
and dark. (Inthe type, only the first segment of the left an-
tenne is left; this is long, cylindrical, and testaceous.)
Pronotum reddish testaceous, slightly narrower than the
head, the front border straight, the hinder border rounded,
Elytra well developed, dark reddish testaceous, broader
than the elytra at the shoulders, smooth, truncated at the
apex.
Wings of the same colour as the elytra, smooth, not pro-
jecting far beyond the elytra. ~
84 Mr. M. Burr—WNotes on the Forficularia.
Feet light testaceous, short, the femora rather strongly
compressed.
Abdomen depressed, slightly dilated posteriorly. The
segments, except the last, are all finely granulated, each
segment being also slightly broader than the previous one ;
the tubercle on the second segment is very distinct, black,
the tubercle on the third segment is considerably larger and
black ; these two segments are much darker than the re-
mainder, and the part round the tubercles also is black; the
last dorsal segment is very large, square, light testaceous,
smooth and shining; at each angle on the posterior margin
above the insertion of the forceps is a large round tubercle,
slightly furrowed in the middle, giving the appearance of a
double tubercle. The underside of the abdomen is much
paler than the dorsal side and is entirely granulated. The
penultimate segment is very large, only leaving the ultimate
segment visible at the extreme posterior corners. The poste-
rior margin of the last dorsal segment has an impression in
the middle.
Forceps: these are very long and slender, almost as long
as the body. At the base they slightly diverge at first, then
gradually point inwards, to meet and cross at the apex. At
the base they are triquetric and armed with a small conical
tooth on the inner margin, just at the pygidium; there is a
second, much stouter tooth halfway down on the inner margin ;
the part between these teeth is finely crenulated. The colour
of the forceps is clear testaceous, on the underside they are
flattened.
Pygidium short, rectangular, very broad, not toothed or
emarginate, the hinder border straight and simple ; the angles
are sharp; in the middle on the upperside it is slightly
bi-impressed.
Patria. British East Africa, Nairobi, 5500 feet; Kikuyu
Country, July 1899 (HZ. J. Mackinder & CO. B. Hausburg).
Type in Mus. Hope, Oxford.
In the form of the forceps, lengthening of the body, and
largeness of the abdominal tubercles this species recalls certain
species of Foreipula; but the lobed second tarsal segment
definitely shows that it cannot be related to that genus.
I have great pleasure in dedicating it to Mr. H. S. Mackinder,
the eminent geographer, who discovered it during the recent
expedition to Mount Kenia.
For the drawings of this species I am indebted to my
friend Mr. EH. H. J. Schuster, F.Z.S.
Mr. M. Burr—Notes on the Forficularia. 85
Opisthocosmia oannes, sp. n.
Statura majore; colore nigro, elytris alisque rufescentibus ; antennz
13-segmentate, rufescentes ; pronotum capite angustius, nigrum,
pallide marginatum, rotundatum ; pedes nigri, tarsis pallidioribus :
abdomen nigrum, medio subdilatatum; segmentum ultimum
dorsale angustius, quadratum, margine postico subbituberculatum,
angulis acutis: forcipis brachia ¢ basi remota, rotundata, recta,
gracilia, apicem versus incurva, mucronibus decussatis, ante
medium dente magno obtuso supra sursum spectanti armata,
paullo ante apicem margine interno dente parvo acuto armata ;
pygidium haud prominulum. 6.
Tongs COrporis’ "i... vss «ss 14 mm.
pe GRCADIS yr ct 3's ops cs Gh
Head black; antenne 13-segmentate, the first segment
black, the remainder reddish.
Pronotum slightly narrower than the head, the anterior
border straight, the hinder border rounded, all angles rounded ;
black in the middle, the sides pale.
Llytra smooth, considerably broader than the pronotum at
the shoulders, black, shaded with dark red.
Wings prominent, black, with a red spot in the middle.
Feet black, the tibie towards the apex and the tarsi paler,
Abdomen black, very slightly dilated about the middle, the
lateral tubercles very distinct, the sides of segments 5-6
slightly produced backwards; last dorsal segment narrower,
square, reddish, the hinder border straight, with two faint
obtuse tubercles above the insertion of the forceps.
Pygidium not apparent.
Forceps 8 with the branches remote at the base, straight,
black, cylindrical, incurved at the apex and strongly de-
cussating there; armed near the base above with a strong
blunt conical tooth, pointing upwards, and a smaller acute
tooth on the inner margin near the apex.
Patria. Assam (Linden, 1894, ex coll. Seeldrayers).
‘Type in my collection.
For this fine novelty I am indebted to my friend M. Seel-
drayers.
It is not closely allied to any known species, but approaches
rather to an as yet undescribed species brought from Siam
by Mr. Annandale; it differs, however, very markedly in
colour.
Ancistrogaster inopinata, sp. i.
Statura parva; colore nigro, abdomine zeneo-nitenti, pedibus testa-
ceis; antenne 13-segmentate, fusce, apice pallescentes; pro-
86 Mr. M. Burr—WNotes on the Forficularia.
notum capite paullo angustius, nigrum ; elytra punctulata, nigro-
castanea, nitentia, apice truncata; ale parum prominentes, nigre,
apice macula parvula pallide ornate: abdomen medio dilatatum,
nigro-rufescens, tuberculis lateralibus distinctis, segmentis 4
ultimis Jateribus paullo retrorsum productis ¢ ; segmentum
ultimum dorsale apice angustatum, margine postico recto, sub-
tuberculato, Q distincte tuberculato: pedes testacei, tibiis necnon
infuscatis: forcipis brachia g basi subcontigua, plus minus
dilatata et deplanata, medio divergentia, tunc incurva, apice attin-
gentibus, prope basin supra dente magno trigonali sursum spec-
tanti armata, margine interno in parte media crenulata; ¢ recta,
contigua, inermia, apice decussata. of Q.
mm. 62°5 ‘ane
29 2
Long, corporis... .. 5
ath AONCINIS Ne xeye pe 2°75 a
Head black; autenne 13-segmentate, fuscous, paler at the
apex.
Fronotum slightly narrower than the head, black, indis-
tinctly marginate with reddish ; anterior border straight, with
the angles rounded ; posterior border rounded.
Evytra short, very finely punctulated, shining, very dark
reddish castaneous, truncate at the apex.
Wings not very prominent, of the same colour as the elytra,
with a very small indistinct pale spot on the inner margin at
the apex of the suture.
Feet testaceous, pubescent, the tibiee and tarsi somewhat
darker.
Abdomen dilated in the middle, dark castaneous, shining,
with a slightly metallic reddish sheen, the sides darker ;
lateral tubercles very distinct ; in the ¢ the sides of segments
€-9 are slightly produced backwards, but this is not very
distinct; last dorsal segment attenuated, the hinder margin
nearly straight, broken with indistinct obtuse tubercles.
Pygidium not apparent.
Forceps: 3 with the branches subcontiguous at the base
and somewhat depressed and dilated there, then diverging,
and attenuate and incurved towards the apex, where the
points meet ; near the base there is a very strong triangular
sharp tooth pointing straight upwards; the middle third,
where the dilated part is fading out, is crenulated on the
inner margin; @? with the branches straight, simple, un-
aimed, the apices crossing ; colour dark reddish black,
Patria.’ Costa Rica, 3 9 (Linden, 1894, ex coll. Seel-
drayers).
Type in my collection.
Mr. M. Burr—WNotes on the Forficularia. 87
For this interesting species I am also indebted to my friend
M. E. Seeldrayers, of Brussels.
It is a distinct species, coming next, perhaps, to A. varie-
gatus, Dohrn. It has the appearance of an Opisthocosmia.
Ancistrigaster intermedia, sp. 0.
Castanea ; antenne fusco-rubrescentes, segmentis elongatis, 2° parvo,
duobus ultimis pallidis ; pronotum antice rectum, postice rotunda-
tum ; elytra perfecte explicata, latiora, levia ; ale perfecte expli-
cate, valde prominentes: abdomen medio dilatatum, segmentis
2° et 3° valde tuberculatis, segmentis 4° et 5° obsolete, vix, vel
haud tuberculatis; segmentum ultimum dorsale ¢ quadratum,
rectangulum, inerme, supra insertionem forcipis subtuberculatum ;
© angustum, attenuatum, declive, inerme: pedes graciles: for-
cipis brachia ¢ basi distantia, intus basi ipso dente conico
armata, dehine paullo divergentia, medio unidentata, apicem
versus valde incurya, mucronibus decussatis; @ simplicia, recta,
gracilia, inermia, apice mucronata. ¢ Q.
3. an
oie, CORPOLIS: 6) <r yere)s yy 8 mm. 8 mm.
Si, LORCAPISN «Y= 3 bs Cae mes as 2 for 2
Head flattened, clear red; antenne reddish, the last two
segments paler, all elongate, except the second, which is
short ; eyes jet-black.
FPronotum about as broad as the head, straight on the fore
border, rounded posteriorly, dark testaceous or dark red, the
margins clear luteous.
Kilytra ample, broad, smooth, shining, fusco-testaceous,
the hinder border truncate.
Wings ample, projecting well beyond the elytra, fuscous,
sometimes with a large pale spot in the middle at the base
and a smaller one at the apex.
Feet long and slender, pale testaceous.
Abdomen dilated in the middle, fusco-testaceous or dark
brown. Segments 2 and 3 with lateral tubercles dark or
black, very distinct ; in the male segments 4 and 5 have very
faint, barely visible, obsolete tubercles. In the ¢ the last
segment narrow, quadrate, rectangular, the hinder border
straight, with a small tubercle above the base of each branch
of the forceps; in the ¢ the last dorsal segment is narrowed,
square at the hinder border, and unarmed.
Pygidium not visible.
Forceps: g with the branches distant at the base, armed
there on the inner margin with a strong conical tooth, flat-
tened, slightly diverging, with a sharp tooth in the middle of
88 Mr. M. Burr—WNotes on the Forficularia.
the inner margin ; towards the apex the branches converge
strongly, and are very strongly dilated immediately before
the apex, where they are pointed and hooked; in the ¢ the
branches are simple, straight, smooth, unarmed, and incurved
slightly at the apex.
The feet, abdomen, and forceps are clothed with a pale
short yellowish pubescence.
Patria. Peru, 1 §,3 9 (ex coll. Staudinger).
Type in my collection.
This species stands nearest to A. Championi, Borm., from
which it may be distinguished by the square ultimate dorsal
segment, which has not the angles sharply produced as in
A, Championi, by the abdominal tubercles being distinct only
on segments 2-3, and not on segments 2-7, and by the forceps,
which have the branches less strongly diverging, with fewer
teeth.
Dormans Park, East Grinstead,
April, 1900.
EXPLANATION OF PLATE IV. Fas. 3, 5, 7.
Fig. 3. Apterygida Mackindert, sp.n. Somali. .
Fig. 3a, Ditto, Segmentum ultimum ventrale, from below.
Fig. 5. Anechura ahrimanes, sp.n. Sikkim. 6.
Fig. 7. Forficula orientalis, sp. n. Constantinople. ¢. Last segment
and forceps.
XJ.—Notes on the Forficulariaa—VI. On a Collection of
Forficularia from Sarawak. By Maucotm Burr, F.Z.S.,
[Plate IV. figs. 1, 2, 4, 6.]
Mr. R. SHELFORD, of Sarawak, has very kindly handed to
me for determination a small collection of earwigs from that
region, The collection includes twenty-five species, of which
nine are new, and there are some very curious forms.
The localities from which they come are thus explained by
Mr. Shelford :—
Kuching.—The capita] of Sarawak.
Matang.—A mountain 7 miles from Kuching, 2800 feet.
Penrissen.—A mountain 50 miles from the sea, at the
head of the left branch of the Sarawak River, reaching
My. M. Burr—Wotes on the Forficularia. 89
an elevation of 4800 feet; the specimens were taken
between the levels of 3300 feet and 4000 feet.
Pankalan Ampat.—At the foot of Penrissen.
Batu Song.—A mountain in the Baram district.
For the drawings I am indebted to Mr. H. C. Philips.
The collection works out as follows :—
1. Apachys chartaceus, de Haan.
Kuching, Feb. 2, 1899, 2 g; June 13th, 1899, 1 9;
Dec. 12th, 1899, I 3.
This species has been recorded by de Haan from Borneo
and Sumatra and by Dubrony from Sarawak. According to
de Haan, it lives under the bark of dead trees.
2. Pygidicrana marmoricrura, Serv.
Batu Song, 1 2 (C. Hose, Esg.). There are also two
females in bad condition which I refer to this species, from
Penrissen, Feb. 24th and May 1899.
The species is common in the Hast Indies.
3. ? Pygidicrana modesta, Borm.
Matang, August 1899. One fragmentary female, which I
refer with some doubt to this species.
Recorded from Burmah.
4. Echinosoma sumatranum, de Haan.
Kuching, Sept. 13th, 1899, and Nov. 27th, 1899, 2 @;
Sept. 25th, 1899, 1 3; Oct. 6th, 1899, 1 ¢.
De Haan remarks of this species, “ Vivit sociatim in ligno
putrido, Motus alacris.”
5. Psalis borneensis, Kirb.
Kuching, Feb. 1899, Dec. 26th, 1899, 2 ¢; Dec. 15th,
13995 1:2
As Kirby suggests, this is probably a melanic form of ©
P. indica, Hag.
6. Spongiphora rubriceps, sp. n.
Statura mediocri; caput rufum; antenne 15-segmentate, fusce,
nigro-annulate ; pronotum postice rotundatum, nigrum, pallide
marginatum ; elytra aleeque fusce ; pedes testacei, fusco-variegati :
90 Mr. M. Burr—WNotes on the Forficularia.
abdomen cylindricum, fuscum; segmentum ultimum magnum,
quadratum, margine postico tuberculato: forcipis brachia ¢
basi remota, recta, apice incurva, mucronibus attingentibus, rubra,
margine interno crenulata, haud dentata; 9 recta, deplanata,
minute crenulata.
Long.corporis .<<.- J 11 mm. 10°25 mm.
Pr) forcipis oweeeve 4 39 3d 9
Head flattened, clear brick-red; eyes small, black: an-
tenne 15-segmentate; first segment long, testaceous, dark at
the apex; the other segments are black, except segments
10-11, which are clear white ; from the second, the segments
are short, gradually lengthening.
Pronotum as broad as the head, chocolate-brown, the
margins whitish or testaceous ; the anterior border is slightly
round-convex ; the sides are parallel; the hinder border is
rounded.
Elytra perfectly developed ; chocolate-brown, dull, a little
broader than the pronotum at the shoulders, hinder borders
obliquely truncate, slightly sinuate.
Wings ample, projecting well beyond the elytra, of the same
colour, with a pale spot at the apex on the inner margin (this
spot is sometimes very faint).
Feet dark testaceous, varied with paler.
Abdomen slightly depressed, the sides parallel; rich brown
in the centre, darker at the sides; abdominal tubercles
distinct, black. Last dorsal segment of the g simple, not
sulcate, with a row of small tubercles on the hinder margin,
which is straight. In the ? the last dorsal segment slightly
narrower, with a faint sulculus near the hinder margin, the
tubercles smaller and fewer in number. The penultimate
ventral segment ¢ is ample, almost entirely covering the last
segment, nearly rectangular, the hinder margin rounded.
Pygidium not visible.
forceps: & branches remote at the base and three-keeled
there, then simple, nearly straight, gradually converging to
meet at the apex; in the apical half on the inner margin there
are three small teeth; the forceps are bent noticeably down-
wards from the base. @ branches simple, straight, remote
at the base, more or less flattened, the apices meeting; the
inner margin near the base is very slightly dilated and crenu-
late ; otherwise the branches are unarmed.
Patria. Sarawak, Kuching, Dec. 3rd, 1898; July 15th,
1899, 2 g; March 1899, 2 ¢ ; Oct. 23rd, 1899, 1 9 ;
July 14th, 1899, 1 nymph.
‘his species is most closely allied to S. n¢tzdipennis, Borm,
Mr. M. Burr—WNotes on the Forficularia. 91
7. Spongiphora semiflava, Borm.
Kuching, Jan. 2nd, 1900, 1 9; July 7th, 1899, 1 ¢@.
This species, which closely resembles Labia mucronata,
Stal, in appearance, has hitherto been recorded only from
Burmah.
8. Spongiphora sphinz, sp. n.
Statura mediocri, gracili; colore testaceo vel fusco; antennex 18-
segmentate ; caput, antenne, pronotum, elytra, ale, pedes testa-
cea vel rubro-testacea; abdomen cylindricum, testaceum vel
rubrum ; forceps gracilis, brachiis basi remotis, rectis, apice
mucronatis, attingentibus, margine interno minute denticulatis ;
abdomen tuberculis distinctis instructum. o¢ 9.
Long. corporis .... 11°5 mm. 10 mm.
pe TOLEIDIEG Gore) Ey Be
lead depressed, clear brick-red: antenne 18-segmentate ;
the first segment long, testaceous, the remainder darker;
second segment very small, the third nearly as long as the
second and fourth united, the fourth, fifth, and sixth
gradually lengthening, the apical segments long and slender.
Pronotum slightly narrower than the head, long and
narrow, slightly broader posteriorly than anteriorly, rectan-
gular, the hinder angles rounded, posterior border nearly
straight ; clear testaceous, reddish anteriorly, the margins
paler; in the ? shaded with fuscous on the disk posteriorly.
Elytra long and narrow, clear testaceous, the outer and
inner margins darker.
Wings conspicuous, clear testaceous.
feet clear testaceous, clothed with a few fine pale hairs.
Abdomen cylindrical, reddish testaceous in the ¢, reddish
black in the $3 lateral tubercles very distinct, all the sey-
ments very finely granulated ; last dorsal segment ¢ square,
hinder margin straight, slightly tuberculated above the inser-
tion of the forceps on each side; similar in form in the 9;
penultimate ventral segment ¢ large, rounded posteriorly,
not covering the apical part of the last segment, which has
the posterior border straight, very deeply emarginate in the
middle, divided into two rounded lobes, the outer angles
folded, forming a bicarinate tubercle on each side. In the ?
the penultimate ventral segment is rounded, larger, covering
the ultimate segment entirely except for the outer angles,
Pygidium 8 ? quadrate, the side slightly emarginate near
the apex.
92 Mr. M. Burr—WNotes on the Forficularia.
Forceps: & with the branches remote at the base, slender,
almost straight, slightly depressed, the apices meeting, and
strongly mucronate ; at two thirds their length on the inner
margin there is a small tooth ; from the base to this tooth the
inner margin is very finely denticulated. In the 9 the
branches are straight, remote at the base, less depressed, the
apices meeting, the inner margin finely denticulated to about
two thirds the length, where there is an obsolete tooth barely
distinguishable. ‘lhe forceps are clear red in the male,
reddish black in the female.
Patria. Sarawak, Kuching, August 1897 (“Z.”), 1 3;
Oct, 12th, 1899, 41 39 ..
This curious species is very close to Labidura? decipiens,
Kirb., which must almost certainly be moved, at least pro-
visionally, to this genus. It differs in the form and armature
of the forceps. It has the appearance of certain species of
Apterygida, as A. erythrocephala, Oliv., A. suturalis, Serv.,
and A. bipartita, Kirb. It may be easily mistaken for the
latter at first glance; but an examination of the second tarsal
segment shows that it is simple and cylindrical; the distinct
lateral tubercles prevent its being placed in Labidura. The
form of the penultimate ventral segment of the male distin-
guishes it from typical Spongiphora, and, as Mr. W. F.
Kirby suggests, a new genus must be erected tor his Labidura ?
decipiens and for this species when better known.
9. Spongtphora nitidipennis, Borm.
Kuching, July 1899, 1 ?.
Hitherto recorded from Burmah and Sumatra.
10. Sparatta setulosa, sp. n.
Colore sordide testaceo; pronotum valde elongatum, parallelum ;
pygidium haud perspicuum ; forcipis brachia ¢ basi remota,
basi ipso intus laminata, dehine recta, sensim incurva, tertia
parte apicali dente inermo armata, mucronibus attingentibus, valde
pubescentia. d.
rofte
TONES COTPOTISs a se ele sare 1 ani
a MOrcipis yess ree 3
Head dirty yellow; eyes small, black; antenne with
13 segments remaining, fuscous, the segments elongated and
slender, numbers 2-4 enlarged at the apex.
Pronotum twice as long as the head, very narrow, the sides
Mr. M. Burr—WNotes on the Forficularia. 93
parallel, anterior border produced to a neck, posterior border
rounded, tawny yellow.
Elytra narrow, obliquely truncated posteriorly.
Wings prominent, black, the dark colour showing through
the clear elytra.
Feet tawny.
Abdomen slightly broader at the apex than at the base ;
tawny, slightly darker in the middle ; all the segments very
finely punctulated except the last, which is smovth, large,
and quadrate, the posterior margin straight, with a row of
small tubercles.
Pygidium barely visible, quadrate.
Forceps slender, remote at the base; on the inner margin
at the base itself there is a strong depressed laminated tooth ;
the branches are then very faintly curved outwards, then
gradually inwards, the apices meeting ; near the apex on the
inner margin is a strong but small oblique tooth; the forceps
are dirty yellow at the base, darker towards the apex, clothed
with long fine bristles,
The feet and abdomen are also clothed with a few fine
long bristles.
@ unknown.
Patria. Sarawak, Matang, August 1899, 1 ¢.
This species is not closely allied to any described form
except S. Horsfieldi, Kirb., trom which, however, it is very
distinct,
11. Sparatta Brunneri, Borm. ?
Penrissen, May 1899, 2 ?.
J refer two females with some doubt to this North-Australian
species.
12. Chetospunia parvula, sp. n.
Statura minore; colore nigro, rubro-variegato; caput, pronotum,
et elytra nigra ; ale deficientes ; abdomen segmentis 1°—7™ nigris,
ceteris rubris; forcipis brachia ¢ basi distantia, valida, basi
dente gracili instructa, margine interno denticulata, incurva,
mucronibus decussatis. ¢. Q ignota.
Dongs COMPOris: ...°..« «<< oe 5 mm,
eget (0; CCl Ost pee Renae ee 1°23 5;
Head and eyes jet shining black; antenne with only one
or two segments remaining, paler.
Pronotum jet-black, as broad as the head, slightly broader
posteriorly than anteriorly, the angles rounded; anterior
border rounded, posterior border straight.
94 Mr. M. Burr—WNoées on the Forficularia.
Elytra black, rounded at the apex.
Wings absent.
Feet clear testaceous.
Abdomen slightly broadest in the middle, shining black,
except segments 8-9, which are red, and the last segment,
which is red, quadrate, slightly narrower posteriorly than
anteriorly ; the hinder border straight, smooth, black.
Pygidium not visible.
Forceps § with the branches stout, three-keeled at the
base, straight, then gradually incurved towards the apex,
where the points decussate; at the base itself, on the inner
margin there is a fine oblique tooth, and the inner margin is
denticulated for the first two thirds of the length; the forceps
are black near the base, red at the apex.
The abdomen is clothed with a few fine pale hairs.
Patria, Sarawak, Kuching, Dec. 15th, 1898, 1 ¢.
This species is nearest to Ch. fee, Borm., from which it
differs entirely in the shape of the forceps.
13. Chetospania Jupiter, sp. n.
Statura majore ; colore fusco-testaceo ; antennie 15-segmentate, seg-
mento 2° parvo, ceteris elongatis, cylindricis ; pronotum quadratum,
marginibus anteriori et posteriori rotundatis, lateribus parallelis,
angulis posticis rotundatis; elytra setulosa, apice truncata; alee
setulosee ; abdomen paullo dilatatum, tuberculis vix prominulis,
segmento ultimo margine postico recto, integro: pygidium elon-
gatum, apice attenuatum, emarginatum; Q brevius: forcipis
brachia ¢ elongata, basi distantia, triquetra, depressa, basi ipso
dente valido armata, carina superiori margine interno ante
medium dentata, carina inferiori in parte apicali bidentata, dehine
crenulata, mucronibus attingentibus, decussatis ; brachia 9 illis
maris similia, rectiora, apice fortius decussata. dQ.
G. :
Long. corporis 12 mm. 10-10°75 mm.
35, MORCIDIEs/nntewt » O-OlO 1, o'20-6 ,,
Head reddish fuscous, sutures well marked; eyes not
prominent; antenne fuscous, with 15 segments, the second
small, the remainder long.
Pronotum longer than broad, reddish fuscous, as broad as
the head, hinder border and anterior border rounded, the
angles rounded, the sides parallel ; posterior part of the disk
clothed with fine yellow hairs, directed backwards.
Elytra fuscous, truncated at the apex, pubescent like the
pronotum.
Mr. M. Burr— Notes on the Forficularia. 95
Wings prominent, coloured and pubescent like the elytra,
the margins very narrowly paler.
Feet dark testaceous, pubescent.
Abdomen reddish brown, very finely granulated, very
faintly broader towards the apex; lateral tubercles barely, if
at all, distinct. Last segment smooth, deep red; hinder
border unarmed, straight in the g,sinuate in the ?. Penul-
timate ventral segment large, rounded, almost entirely
covering the ultimate segment.
Pygidium prominent, attenuated towards the apex, conical,
emarginate at the apex in the @, in the ? shorter and
thicker or barely distinguishable.
Forceps: the branches are remote at the base, tricarinate,
with a strong triangular depressed tooth on the inner margin
at the base itself in both sexes; upper keel on the inner
margin before the middle with a strong tooth (¢ ?); lower
margin with two teeth in the apical half, and then denticu-
lated (g¢ 2); the whole of the inner margin, lower keel, is
more or less denticulated in both sexes, this armature
varying in individuals. The branches ¢ are nearly straight,
very gradually incurved, with the points meeting and barely
decussating at the apex; in the ¢? the branches are less
curved, straighter, but pointing more inwards, meet sooner,
and decussate more strongly at the apex ; in both sexes they
are clothed with fine pale hairs; they are testaceous in the
6, darker at the apex, much darker throughout their length
in the ?.
The @ is smaller and generally darker in colour than
the “Gu
Fatria. Sarawak, Penrissen, May 1899, 2 $,2 2.
This is the largest species of the genus. It stands
nearest to Ch. inornata, Karsch, from Madagascar, but is
larger, and the forceps are different.
14. Chelisoches Ritseme, Borm.
Kuching, Sept. 13th, 1899, 1 ¢ ; Dec. 15th 1898, 4 ¢,
ee
This species has been taken in Sumatra and in Burmah.
15. Chelisoches Dorie, Borm.
Kuching, Oct. 28th, 1898, 1 $; Dec. 29th, 1898, 4 ¢;
May 20th, 1899, 1 g; Dec. 1898, 2 ?; August 11th, 1899,
Reo!
This splendid species has been recorded from Sumatra
and Sarawak, where it does not appear to be rare.
96 Mr. M. Burr—WNotes on the Forficularia.
Mr. Shelford’s specimens from Sarawak all have the
branches of the forceps ¢ strongly toothed and crenulate
near the base, and then smooth and unarmed almost to the
apex, just short of which on the inner margin is a strong
tooth. The specimens vary considerably in size, from
15-23 millim. in length of body, excluding the forceps, which
are of a more uniform length.
16. Chelisoches pulchripennis, Borm.
Kuching, Oct. 17, 1898, 3.4; Oct. 28th, 1698, cee
Oct. 17-18, 1898, 2 9.
Recorded from Burmah and ‘ Indes orientales.”’
17. Chelisoches Shelford, sp.n. (PI. LV. fig. 4.)
Statura maxima. Habitus generis Opisthocosmic. Fusco-castanea,
metallescens; pronotum ovale, capite paullo angustius; elytra
granulata, latiora, apice truncata; ale granulate: abdomen
cylindricum, tuberculis pliciformibus distinctis; segmentum
ultimum transversum: forcipis brachia basi remota, recta, elon-
gata, sinuata, margine interno utrinque dentibus duobus vyalidis
armata, apice mucronata, attingentia. J. Q ignota
Vong. Gotporis. 6a <.<1- 15°5 mim.
ay) LORGIDIS |<. wares ote bs Rae
Head and eyes reddish castaneous; antenne with 18
segments, fuscous, the last four dirty white.
Pronotum slightly narrower than the head, oval, all the
margins rounded.
Elytra ample, broad reddish castaneous, with a metallic
sheen, finely granulated, truncated at the apex.
Wings of the same colour as the elytra, granulated.
Feet tuscous, the tarsi and apices of the tibie dirty testa-
ceous; the lobe of the second tarsal segment very conspicuous.
Abdomen cylindrical, reddish castaneous, granulated; the
last segment is transverse, slightly narrower than the pre-
ceding, with a row of tubercles on the hinder border. Penulti-
mate ventral segment large, rounded, entirely covering the
ultimate segment.
Forceps & with the branches widely remote and _ tri-
carinate at the base, curved at first slightly outwards, the
inner keels on this part denticulated; the branches are then
nearly straight, subsinuate, each faint sinuation marked
on the inner margin with a long oblique sharp tooth; these
teeth are two in number on each side; after the second tooth
the branches curve in, the apices hooked and meeting. Seen
Mr. M. Burr— Notes on the Forficularia. 97
from the side, the forceps are sinuate, curving slightly
upwards.
The legs and forceps are covered with a thick yellowish
pubescence.
Patria. Sarawak, Pankalan Ampat, May 1899, 1 ¢@.
I have very great pleasure in dedicating this very fine
species to Mr. Shelford, to whom I am indebted for the
opportunity to work out this interesting collection.
It is a very distinct species and not closely allied to any
known form. It comes closest perhaps to Ch. superbus. It
has somewhat the appearance of a large Anechura, but the
lobe of the second tarsal segment is very distinct.
18. Chelisoches Hercules, sp.n. (PI. IV. figs. 2, 2a.)
Statura robusta ; colore nigro, elytris alisque ceruleis, metallescenti-
bus ; pronotum ovatum, cum elytris alisque granulatum: abdomen
cylindricum, nitidum, nigrum, testaceo-pubescens, tuberculis
lateralibus distinctis; segmentum ultimum dorsale magnum,
margine postico utrinque tuberculo obtuso magno, apice trun-
eato, valido obliquo armatum: forcipis brachia ¢ valida, basi
depressa et deplanata, hac parte margine interno valde denticu-
lata, dehine valde semicirculariter incurva, intus denticulata,
apice haud attingentia ; forceps et pedes valde pubescentes. <.
@ ignota.
Long. corporis ..... . 15°5-20 mm.
sip SRORCUDIS? Wah ata. 4°5 a)
Latit. max. forcipis.. 5 i
Head and eyes black ; antenne with 15 segments, typical,
black, the eleventh whitish ; sometimes the apical half of the
antenne dirty whitish.
Pronotum as broad as the head or very slightly narrower,
round, black, granulated; with a very faint median carinula,
on each side of which anteriorly there is a round elevation,
with a slight depression in the middle; the margins are
turned slightly upwards.
Hlytra considerably broader than the pronotum at the
shoulders, granulated, deep blue, with a metallic sheen;
they are truncated at the apex.
Wings of the same colour as the elytra, granulated, very
narrowly edged with pale. ‘The elytra and wings are clothed
with a short thick pubescence.
Feet stout, fuscous, the tibie clothed with a thick testa-
ceous pubescence, the tarsi with a reddish pubescence.
Abdomen cylindrical, finely punctulated, shining black, the
lateral tubercles very distinct and prominent; the last dorsal
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 7
98 Mr. M. Burr—Notes on the Forficularia.
segment is ample, the hinder border rounded ; at the hinder
border in the middle there are two very stout, long, elevated
tubercles, pointing upwards and obliquely outwards, truncated
at the apex. Penultimate ventral segment large, rounded,
covering the ultimate segment except at the corners.
Pygidium visible from beneath, short, conical, emarginate
at the apex.
Forceps with the branches g short, stout, strongly de-
pressed and dilated at the base, the inner margin of this
dilated part is strongly toothed; at half their length the
branches are cylindrical and curved inwards in a semicircle,
the apices are pointed but do not meet; the inner margin of
this curved part is denticulated.
The whole of the abdomen, tubercles, and forceps are
thickly coated with a short dense testaceous pubescence.
Patria, Sarawak, Kuching, Dec. 19th, 1899, 1 ¢; Pan-
kalan Ampat, May 1899, 1 ¢@.
This remarkable earwig is very distinct from any form
familiar to me. ‘The male from Pankalan Ampat presents a
curious form of aberration. The right branch of the forceps
is scarcely developed, very short, almost straight, bent slightly
downwards and inwards, with no sign of teeth or serrulation ;
the left branch is not fully developed, the denticulation of the
curved part being wanting. ‘This arrest of development is
well known to occur in Forficularia, and is often called
hermaphroditism or gynandromorphism ; but it is extremely
doubttul whether this is ever really the case. In the specimens
which I have examined the normal number of segments of
the male—that is, nime—are visible ; in the female only seven
segments of the abdomen are visible. I have in my collec-
tion two specimens of Chelisoches morio, Fabr., in which the
lett branch is that of the male in the form and the right is
that of the female, and these I have recorded elsewhere * as
heimaphrodites ; but it is more probable that the phenomenon
is due to airested development, owing to want of nourish-
ment or some accident. I have a male of Pygidicrana
marmoricrura in which both branches of the forceps are
abnormal, being nearly straight and unusually pubescent ;
this is certainly an aberration, and not a regular dimorphic
form. The specimen of Ch. Hercules in which the right
branch is malformed is noticeably smaller than the other
tully developed male, and it may well be due to defective
nourishment,
* Ent. Month, Mag, 1897, p. 147,
Mr. M. Burr—WNotes on the Forficularia. 99
19. Anechura coriacca, Borm.
Penrissen, May 1899, 1 g, 1 9; Matang, Dec. 1898,
1 ¢.
Recorded from Burmah,
20. Anechura scabriuscula, Serv.
Matang, Dec. 1898, 1 ¢.
Common in the Oriental Region.
21. Anechura Hermes, sp.n. (PI. IV. figs. 1, 1a.)
Colore atro ; pronotum antice rectum, postice rotundatum ; elytra
pronoto latiora, pubescentia; ale prominentes: abdomen con-
vexum, medio valde dilatatum, apice attenuatum; segmentum
ultimum dorsale angustum, margine postico medio bicarinatum :
forcipis brachia ¢ valida, triquetra, basi subcontigua, dehine
valde divergentia, subito fortiter incurva, mucronibus haud
attingentibus, prope basin supra dente maximo obtuso perpen-
diculari armata. ¢. @ ignota.
NSE COLONIE! i 5s 6 ois wfole cig oarersa Lees 11°5 mm
RA MLORETDUS, Sr taietaa hic sis es Gas awh Oe
Minti, WA LOLCIDIS 2c dws ee ere es ve ts 4,
Altitudo max. forcipis (tuberculo basali
‘LEE ECS(T)) R ARS BS age 7 a tae
Head jet-black ; antenne ?-segmentate (eleven segments
remain, all black, except the eleventh, which is whitish),
second segment small, the rest longer, conical.
Pronotum very slightly narrower than the head, straight in
front, rounded behind, all black.
Elytra jet-black, very finely punctulated, truncated at the
apex.
Wings the same colour as the elytra.
Feet black, with a few fine short bristles.
Abdomen dark reddish brown, strongly dilated in the
middle, attenuated towards the apex ; lateral tubercles very
distinct; edges of the fifth to seventh segments slightly
produced; last dorsal segment narrow and small, hinder
border straight, with a small tubercle on each side above the
insertion ot the forceps, produced backwards into a faint
carina; penultimate ventral segment large, rounded.
Pygidium not visible.
Forceps with the branches g short, stout, almost con-
tiguous at the base, cylindrical, not keeled, strongly diverging
at first, then the points suddenly and sharply turned inwards
7*
100 Mr. M. Burr— Notes on the Forficularia.
eta right angle, then obtuse, not meeting ; near the base on
the upperside on each branch there is a long and strong
conical obtuse tooth, pointing upwards and obliquely outwards
backwards ; forceps all black, entirely unarmed except for
these large tubercular teeth,
Patria, Sarawak, Penrissen, May 1899, 2 ¢.
This curious form recalls Ancistrogaster in the form of the
abdomen, but the antenne are different. It is very distinct
from any known form of Anechura, and in appearance
resembles certain species of Chelisoches, except for the dilated
abdomen, which is oval in shape.
22. Apterygida borneensis, Dubr.
Kuching, Dec. 14th, 1898, Dec. 19th, 1899, July 14th,
1899, 3 @; May 1897,1 92.
This species is represented by a variety in which the pro-
notum has no pale border, the head is reddish. The antenne
have 21 segments, of which the tenth to twelfth and the last
two are pale whitish.
23. Forficula? ares, sp.n. (Pl. IV. fig. 6.)
Niger ; pronotum quadratum, postice rotundatum ; elytra latiora,
minutissime punctulata ; alse nigre, apice margine interno minute
testaceo-maculate ; pedes nigri: abdomen apicem versus paullo
dilatatum, apice attenuatum, tuberculis lateralibus distinctis ;
segmentum ultimum dorsale attenuatum, medio profunde rotun-
dato-impressum ; segmentum penultimum ventrale magnum,
rotundatum ; pygidium haud perspicuum: forcipis brachia
eracilia, basi plus minus dilatata et deplanata, hac parte denticu-
lata dehine inermia, rotundato-incurva, apice mucronibus attin-
gentibus, haud decussatis. ¢. 2 ignota.
Long. corporis ........ 10°75 mm,
g5. MOUGIDIS) Petts. stoi: AS Wan
Head jet-black ; antennee with nine segments remaining, of
which the second is very small, the remainder gradually
lengthening, conical.
Pronotum black, as broad as the head, straight in front,
the posterior margin rounded.
Elytra short, broad, truncated at the apex, black, very
finely punctulated.
Wings of the same colour as the elytra.
Feet black, with a thin pubescence.
Abdomen black; lateral tubercles very distinct; broadest
Mr. M. Burr—Notes on the Forficularia. 101
near the apex, but attenuated at the apex itself; last dorsal
segment very small, with a very distinct deep round depres-
sion in the middle. Penultimate ventral segment large and
rounded, completely covering the ultimate ventral segment.
Pygidium not visible.
Forceps & with the branches slender, more or less dilated
and depressed at the base, this dilated part crenulated on the
inner margin, ending with a small, conical, nearly obsolete
tooth ; beyond this tooth the branches are very slender, curved
outwards and then inwards, enclosing a circular space, the
apices almost meeting but not decussating; beyond the
obsolete tooth the branches are quite smooth and unarmed.
Patria. Sarawak, Penrissen, May 1899.
This species recalls in appearance Chelisoches ? picticornis,
Kirb., but it is not so long and is more thick-set. It appears
to be allied to the little-known Forficula lobophoroides of
Dohrn, of which C. picticornis may be the male, which is
not definitely known. J. ares is allied to Anechura hermes
and may have to be moved to that genus. Its uniform black
colour and the form of the forceps distinguish it.
24, Opisthocosmia centurio, Dohrn.
Kuching, Dec. 10-14, 1898.
Previously recorded from Borneo.
25. Opisthocosmia lugens, Borm.
Penrissen, May 1899,2 ¢,4 9.
Previously recorded from Burmah.
Dormans Park, East Grinstead,
April 1900.
EXPLANATION OF PLATE TV. Fies. 1, 2,4 6.
Fig. 1. Anechura Hermes, sp.nu. o. Forceps.
Fig. 1a. Ditto. From the side.
hig. 2. Chelisoches Hercules, sp.n. . Forceps,
Fig. 2a, Ditto. From the side.
Fig. 4. Chelisoches Shelfordi, sp.n. . Forceps.
Fig. 6. Forficula? ares, sp.n. 6. Forceps.
‘102 Mr. F. A. Bather on the
X1.—The Lower Paleozoic Crinoids of Bohemia.
By F, A] BAtuer, MAL, Gs:
THE latest volume of ‘The Silurian System of Bohemia’ *
is welcome indeed to students of Paleozoic geology and pale-
ontology, and more than welcome to the worker on Crinoids.
The account of the rich and varied Cystid fauna, published
13 years ago, had given us hope that the unwearied efforts of
Barrande might have garnered up a fairly rich harvest of
Crinoids, while the known differences between this province
and that of North-west Europe led us to expect considerable
novelty. The eminence and experience of the senior author
_and the intelligent zeal of his collaborator were further war-
rant for pleasurable anticipation. We knew that no pains
were being spared in looking up the literature, while some
valuable papers by Dr. Jahn indicated that the statements of
horizon and locality were being placed on a sure basis. If,
after all, disappointment be our dole, this is due to many
causes, for most of which the authors are in no way re-
sponsible. In the first place, the material is neither rich nor
well preserved: the many-plated calyces are often crushed
and their plates disarranged; they are covered to a deplor-
able extent by “an extremely hard, irremovable limestone
matrix ’’; several specimens that might have been interesting
are represented by mere ochreous powder. Then the vast
majority of forms come from one stratum alone, namely reef-
like bands of limestone that form a transition between the
beds e1 and e2 (about equivalent to our Woolhope Lime-
stone): the first fauna (Cambrian) furnishes no crinoid
remains; the second fauna (Ordovician) includes a few
indeterminable columns chiefly from a4 and 45, and two
species of a new genus, Calerdocrinus, from a4 (about equal to
Middle Bala) ; the third fauna (Silurian s. str.) yields a very
few stem-fragments from e1, rather more from e2, along with
recognizable specimens of Scyphocrinus and of the new genera
Bohemicocrinus, Carolicrinus, and Laubeocrinus, all which,
except the last, occur more abundantly in the underlying
transition beds; f1 contains a few doubtful remains, and f2
yields Beyrichocrinus humilis, gen. et sp. nov., as well as a
* ‘Systéme Silurien du centre de la Bohéme,’ par Joachim Barrande,
lére partie: Recherches Paléontologiques. Continuation éditée par
le Musée Bohéme. Vol. vii. Classe des [chinodermes. 2° partie:
Famille des Crinoides. Par le Prof. Dr. W. Waagen et le Dr, J,
Jahn, Traduit par A. 8S. Oudin. vi and 216 pp., 40 pls. numbered x].
Ixxix. Prague (Dec. 1899).
Lower Paleozoic Crinoids of Bohemia. 103
doubtful Zchthyocrinus and a few undetermined fragments ;
G and H are barren of crinoids, except for rare fragments in
g1,g2,andhi. From this it is clear that the crinoids of
Bohemia could not be made by anyone to throw much light
on the general evolution of crinoids or on the succession of
faunas in the Bohemian basin.
The next obstacle with which the authors have had to
contend is “‘piété pour la mémoire de Barrande.” This praise-
worthy sentiment has caused them to follow with marvellous
success the monumental style of the great ‘ Systeme Silurien,’
and the arrangement which necessitates numerous chapters
after the model of the notorious one on Snakes in Iceland. It
is no doubt the same piety that has induced them to publish
25 plates by a pitiable person called Langhans. Concernin
these plates the authors themselves write: “ most of the
drawings are more or less defective ...many are so ill
executed that they give not the faintest idea of the original ;
&e. &e.” Worse than this, the majority of the specimens
misrepresented on these plates are themselves obscure frag-
ments, to which Barrande, doubtless for his own convenience,
had casually attached manuscript names “au point de vue
scientifique .... nullement justifiés.” And yet “ respect
for his memory ”’ is supposed to be shown by the retention
and publication of all these provisional names. It is useless
for the authors to say, as they do on p. 1389: “ nous déclinons
toute responsibilité sur la question de leur admissibilité.”
Here, unfortunately, are the names, and they have to be
reckoned with. As our knowledge of the Bohemian erinoids
increases, what dissensions will there not be among sys-
tematists as to the value of these names! What hideous
confusion, wrangling, and waste of time! For all of it those
who have published the names will be responsible.
In this article the drawings by Langhans will be ignored ;
difficulties enough will confront us in reconciling the artistic
representations by Swoboda with the outlines thoughtfully
annexed on the thin covering-papers and with the diagrams in
the text. For example, the first question that arises is as to
the position of the small basal in Beyrichocrinus. Plate 59,
fig. 4 shows no basal smaller than the two others; the
covering outline places it in the left posterior interradius ; the
diagram on p. 12 (our Fig. 1) shows it in the left anterior inter-
radius; fig. 1, on p. 18, has it in a position that is different
but not more precisely determinable. A point of equal
importance, especially considering the authors’ doubt as to
the monocyclic nature of this same base, is the orientation of
the lobes of the axial canal as seen in the base of the cup.
104 Mr. F. A. Bather on the
The lobation is doubtless peculiar, but in every figure it is
represented differently, so that one can only enquire whether
the lobes are radial or interradial: they are clearly radial in
one drawing, clearly interradial in another, and not clearly
anything in the rest. The text throws no light on the
matter, unless the following sentences may suggest a solu-
tion of the riddle: ‘Ce genre n’est représenté que par un
fragment de la couronne” (p. 12) ; ‘sous cette dénomination,
nous comprenons le calice et les bras, abstraction faite de la
tige”’ (p. 3); “la base est... . presque entigrement cachée
par la tige” (p. 13); “les exemplaires [note plural!] du
genre Beyrichocrinus étaient munis d’une tige” (p. 15).
But the apparent contradiction is probably due to an ambi-
guity introduced by the translator—another obstacle in the
path of the authors. Whether due to the imperfection of the
specimens or the carelessness of the artists, such discrepancies
are only too numerous.
The Crinoidea of the Lower Paleozoic rocks of Bohemia,
as presented in this volume, may be divided into three
classes: (i.) Incerte sedis; (ii.) Scyphocrinus species and
varieties ; (iii.) eight new species, distributed among seven
new genera. If this statement can be substantiated, then
the authors are to be congratulated on having at least
fulfilled our dream of novelty. But their caution and candour
are so manifest that they will be the first to admit the rather
unsatisfactory foundations for some of these genera, Let us
examine them in order, the order being first by geological
periods and then alphabetical—a final Barrandean touch.
The absence of a strict systematic arrangement and the
doubt that surrounés many of the genera have prevented our
authors from drawing up diagnoses. In attempting to
abstract the salient features, I hope to be pardoned if I use
the terminology elsewhere employed by me*.
* The old-fashioned terminology retained by Messrs. Waagen and
Jahn for the arm-structures obscures their homologies in Camerate
Crinoids. The term pieces brachiales is due to L. de Koninck, who
applied it solely to the fixed arm-ossicles, while he called the free arm-
ossicles articles brachiaux. Since the present authors restrict the term
brachialia to the free ossicles, they should not translate it by pieces
brachiales (p. 5). A similar error as to articles brachiaux occurs on p. 17
of Wachsmuth and Springer’s ‘Crinoidea Camerata.” The term voute
(vault) connotes a structure now admitted by all to be imaginary ; while
the terms ¢rompe (proboscis) and canal nourricier (alimentary canal) are
equally misleading. The well-known term cu’r?, though introduced on
p. 9, is replaced in practice by the term vriles (tendriis), which is no
gain, Again, the term calyx is defined on p. 3 in the sense in which
it has been used by Wachsmuth and Springer and others: “ the employ-
ment of the word calyx to designate that part which is only the dorsal
Lower Paleozoic Crinoids of Bohemia. 105
Beyrichocrinus humilis, the sole species of the genus,
comes from the limestone f2 of Konéprus*. Only the cup
is preserved (Fig. 1). It consists of the following primary
Fig. 1.—Analysis of the cup of Beyrichocrinus humilis, outlines copied
from Waagen and Jahn, and about twice nat. size. Inter-
brachials (i[Br) and anals are shaded from right to left;
intersecundibrachs (iIIBr) from left to right; intertertibrachs
vertically,
elements: 3 basals, 2 large and 1 small; 5 radials, which
appear to abut laterally by short sides, except in the posterior
interradius; 2 fixed primibrachs in each ray, the first being
hexagonal except in the right and left posterior rays; 2 fixed
secundibrachs in each half-ray; 2-4 fixed tertibrachs, of
which the last is axillary and bore the free quartibrachs.
The secondary elements of the cup are: 5-10 interbrachials
arranged in 4-6 horizontal zones, the proximal interbrachial
resting on the steep shoulders of two adjacent radials and
coming very near to the basals; in the wider anal inter-
radius are about 14 interbrachials (anals), of which the
cup is an abuse.” It is therefore perplexing to find the authors per-
sistently using the term in the sense they here deny toit. The fact is
that the word calyx has so often been used for the dorsal cup, that our
attempts to define it otherwise seem bound to fail. The adoption of
the term theca, as used by Haeckel, for the whole case or test con-
taining the central organs of the animal, seems the most rational escape
from the difficulty.
The symbols and terminology of the present paper differ but slightly
from those explained in the Ann. & Mag. Nat. Hist. ser. 6, vol. ix. p. 57,
Jan. 1892. The nomenclature and classification follow Part II1., The
Echinoderma, of ‘ A Treatise on Zoology,’ edited by H. Ray Lankester :
London, 1900.
* Oddly enough, the authors have adopted for the Bohemian names a
spelling different from that hitherto employed in the ‘ Systéme Silurien.’
Their reasons, though not given, are probably good enough to warrant
me following them in this article.
106 Mr. F. A. Bather on the
proximal rests, though by a very short side, on the base,
while it supports 3 plates above ; the intersecundibrachs are
at least 2, but are small and obscure; the existence of inter-
tertibrachs is doubtful. The authors profess not to know
whether the base was monocyclic or dicyclic. But, since no
camerate crinoid has yet been described in which infrabasals
are associated with fused basals, we are justified in placing
Beyrichocrinus among Monocyclica Camerata. Therein it
falls apparently into the Suborder Batocrinoidea, and the
family Periechocrinide, although we do not know whether
it possessed the biserial free arms characteristic of the latter.
Many obscure genera have been referred to this family,
mostly as supposed synonyms of Perdechocrinus. Whether
Beyrichocrinus may be identical with any of these, is a
question for the future. It appears allied to Periechocrinus,
but the cup is rounded as in Megistocrinus ; its most dis-
tinctive characters are the wedge shape of the proximal anal,
and the absence of a distinct median vertical row of anals.
Bohemicocrinus (why not Bohemicrinus?) is established
for two imperfect cups, both assigned to one new species,
B. pulvereus. The trivial name, taken from a manuscript
Fig. 2.—Analysis of the cup of Bohemicocrinus pulvereus, outlines copied
from Waagen and Jahn, a little over nat. size. Shading as in
fig. 1. The small plates at the angles of some of the inter-
brachials are described by the authors as accessory plates; they
do not occur in the only other specimen known.
label by Barrande, is consistently spelled pulverens—a mani-
fest misprint of familiar nature. One specimen comes from
the white limestone, e2 of Kosor, the other from black
limestone transitional between e1 and e2, near Dvorce. The
Lower Paleozoic Crinoids of Bohemia. 107
authors refer their genus to the family Actinocrinide as
defined by Zittel in 1879. ‘There is, however, nothing in
the part preserved (Fig. 2) to prevent its assignment to the
family Carpocrinide founded by the same authority ; and, to
speak even more precisely, there is no particular reason why
these cups should not belong to a Carpocrinus or Desmido-
erinus. ‘The authors indeed say that ‘‘ Bohemicocrinus is
distinguished from all other genera of crinoids described
down to this day, by the characteristic conformation of the
two radii on each side of the anal interradius, as well as by
the structure of the latter and of the other interradii.”’
When this was written the authors were doubtless unac-
quainted with the structure of the cup in Barrandeocrinus,
since that was first published by Wachmuth and Springer
in May 1897. The same peculiarities are emphasized in that
genus; and yet the structure of the cup alone would scarcely
justify the removal of Barrandeocrinus trom Carpocrinus.
It is highly improbable that Bohemicocrinus possessed the
remarkable arms of Barrandeocrinus ; therefore, though the
name Bohemicocrinus may be a convenient way of expres-
sing the imperfect nature of the known specimens, there
seems reason to think that it may ultimately prove a syno-
nym of Carpocrinus or Desmidocrinus. HEucalyptocrinus, Co-
rymbocrinus (= Clonocrinus, Quenst.), and Dolatocrinus have
no anal resting on the base, and it is not clear why they
should have been dragged into comparison.
Carolicrinus is based on a crown and an arm-fragment
from the black limestone (e 1-2) of Karlstein (whence the
name), and another arm-fragment from the white limestone,
e2, near Lochkov. These are all placed in a new species,
C. Barrandei. This species is a many-plated camerate
crinoid not unlike Scyphocrinus, with which it is associated
in the rocks ; but the biserial arrangement of the brachials
would alone justify its severance therefrom. Unfortunately
the authors, impressed by the general resemblance to Scypho-
crinus, have not thought it necessary to compare Carolicrinus
with genera foreign to Bohemia. The imperfection of the
base also is held to excuse them from discussing the sys-
tematic position of the genus. ‘The authors believe that
there were three basals, two large and one small, the latter
being in the posterior interradius (strange position!) ; and
they will not admit that the proximal anal rested on the
base. At the same time they “cannot quite guarantee the
correctness of the explanation just given.” ‘I'he published
figures do not help one even to the bare facts of the case, so
that an attempt at any other interpretation is out of the
108 Mr. F. A. Bather on the
question. But such facts as are undisputed all indicate that
this is nothing but a large species of Abacocrinus, although
that genus has 4 basals and a proximal anal resting on
the base.
Laubeocrinus Barrandet is a new genus and species based
on an incomplete cup provided with 3 millimetres of stem,
from the white limestone, e 2, of Lochkov (Fig. 3). Noattempt
Fig. 3.—Analysis of the cup of Laubeocrinus Barrandei, outlines copied
from Waagen and Jahn, nat, size. Shading as in fig. 1.
is made by the authors to determine the systematic position
of this genus. The peculiarities of the specimen are, indeed,
such that they might be considered individual abnormalities ;
but if that had been the opinion of Messrs. Waagen and
Jahn, they would scarcely have based a new genus on them.
“It is,” they say, “impossible to determine if the base is
monocyclic or dicyclic ””; but since the basals are 4, not 5,
we have every reason to believe that it is what it appears to
be, namely, one of the Monocyclica Camerata. The fused
basals are the right posterior and right anterior. The proximal
anal rests on the base; but instead of the posterior basal
being widened and truncated to support it, the left posterior
basal is widened to occupy the whole of the left posterior
radius, and so to come into contact with the anal. Both of
these features are without parallel in accepted genera. Con-
sidered by itself, the base is like that of Compsocrinus or
Abacocrinus (members of the family Xenocrinide), but in
them the larger basals are posterior and anterior. ‘The re-
maining portions of the cup might belong to any typical
Actinocrinoid. The anal area differs from that of the Xeno-
crinide, and, indeed, of all known Batocrinoidea, in the
presence of only two plates in the second row. PAillipso-
crinus, M‘Coy, is believed by Wachsmuth and Springer to
have been based on an abnormal specimen of Actinocrinus,
and it is noteworthy that the peculiarity is similar to that of
Laubeocrinus, though still more abnormal.
Lower Paleozoic Crinoids of Bohemia. 109
Vletavicrinus, a name derived from the ancient appellation
of the river Moldau, on which lies Dvorce, is based on a
unique specimen from the black limestone (e1-2) of that
locality. The specimen is an almost perfect crown, to which
47 millim. of stem remain attached. It forms the type of a
new species, V. Hauer’, The crinoid (Fig. 4) is a Camerate,
AK) >”
Fig. 4.—Analysis of the type of Vletavicrinus Hauer, outlines copied
from Waagen and Jahn, nat. size. Shading as in fig. 1. The
portions of arms that surmount the anals are intended as repe-
titions of the arm-branches drawn also in the adjoining rays ;
the object doubtless was to show the relations of the pinnules,
but the chief result is to prove the inaccuracy of the diagram.
but whether monocyclic or dicyclic is uncertain, since the
base is almost entirely hidden by the stem. Hach ray con-
tains 2 fixed primibrachs, and the fixed secundibrachs in each
half-ray are said to be 7 or 8 (a number which each of the
five figures chooses its own way of contradicting), the last of
these being axillary. ‘The left posterior interradius, as cleaned
from matrix by the authors, exhibits a proximal interbrachial
resting on the shoulders of the radials, and supporting two
parallel rows of interbrachials, which gradually become
smaller and more irregular and pass into the tegmen. Fifteen
plates in all are exposed in this interradius. ‘The posterior
interradius shows over 30 plates, which seem to have passed
up into an anal tube. ‘The proximal anal rests on the base,
and is followed by a single plate which supports two parallel
rows of irregular plates, 5 in each row, followed by smaller
plates. Rather large intersecundibrachs are visible in the
right and left posterior rays, arranged as in the diagram, and
succeeded by smaller and less regular plates, The arms,
110 Mr. F. A. Bather on the
which are free from the secundaxil, may branch at least
twice more, 7. e. up to quintibrachs, and are composed of low
uniserial brachials bearing closely set pinnules. Unfortu-
nately the authors attach so little importance to the mode of
arm-branching that they have passed over without comment
the very different appearances drawn by Mr. Swoboda in two
identical views of the single specimen. One gathers only
that the branching was not a regular dichotomy. It is also
regrettable that the relation of the pinnules to the brachials is
to be learned from neither description nor figures, the latter
being discrepant in this important point also. We are told
that immediately after each bifurcation the fine pinnules form’
a solid pavement, but we are not told whether any of these
apparently united pinnules form part of the cup-wall; nor,
what is far more important, is it stated if more than one
pinnule is borne by each brachial. Whenever the whole base
is not exposed as in a diagram, Messrs. Waagen and Jahn
seem to think that they are absolved trom any attempt to
relegate the specimen to its place in the system.. Had they
likewise refrained from weighting such a specimen with a new
generic name, their avoidance of responsibility would have
been more readily excused. When, however, they go so far,
we may demand to have such details as it is obvious could
be learned from the specimen, without being forced “ to wait
for more abundant material.” The facts vouchsafed to us do,
all the same, permit some inference. ‘he families of Di-
cyclica Camerata, as at present known, afford no place for
Vletavicrinus. Its reference to Monocyclica Camerata is
more than consistent with the small amount of base that is
visible. If placed in this Order, the relations of its proximal
anal prevent admission to the Melocrinoidea, while the
number of secundibrachs, if ‘nothing else, keeps it out of
Actinocrinoidea. The disposition of the anals, though not
that typical of the Batocrinoidea, is not absolutely discordant
therewith, and it is in that Suborder that it seems to find its
nearest allies. If the brachials do bear more than one pinnule
apiece, if, in other words, they are compound structures, one
would place the genus near Carpocrinus. In any case it
might provisionally be placed in the Carpocrinide. Our
conclusion, then, is similar to that reached in the case of
Bohemicocrinus, from the same horizon and locality ; and we
now remember that the anals of that genus have an arrange-
ment almost identical with that of the present specimen.
Comparison of the descriptions reveals many minor resem-
blances, so many, indeed, that one is impelled to ask why
Lower Paleozoic Crinoids of Bohemia. LiL
Vietavicrinus is not the same as Bohemicocrinus, even if
V. Haueri be not a synonym of B, pulvereus itself.
The last of the alleged new genera from the black lime-
stone, e 1-2, is based on an ill-preserved cup from Dvorce,
and is now called Zenkericrinus melocrinoides. It is the
Xenocrinus mentioned by name only on p. 416 of Dr. Jahn’s
“ Beitriige zur Stratigraphie und Tektonik der mittelbéhm-
ischen Silurformation” *. ‘he authors refer it to the Melo-
crinide, to which family, even in its latest and most restricted
sense, it certainly belongs. ‘hey say that “it is nearest to
Melocrinus, from which, however, it differs in many essential
features.” This opinion also is well founded ; but the authors
seem unaware that the genus Mardacrinus, Hall, is distin-
guished from Melocrinus by those same features. There is
nothing to prevent the relegation of the preseat specimen to
Marvacrinus.
Fig. 5,—Caleidocrinus multiramus. Proximal portions of stem and
crown, seen from alleged right posterior radius; the supple-
mentary plates of the alleged posterior interradius are in two
vertical rows, those of the other are in a single row. This is
fig, 28. @ of Waagen and Jahn, and is said to be “ grossi”; but
not a single measurement is given throughout the account of
this genus. The block has been kindly lent by Dr. Anton
Fritsch, on behalf of the Barrande Committee.
In the second fauna the only representative of the Crinoidea
is a new genus, Calecdocrinus, based on impressions in the
“schistes de grauwacke,’ a4, of Haj Hill, near Zahoran.
* Jahrb. d. k. k. geol. Reichsanst, Wien, xlii. pp. 897-462, Feb. 1893
(not 1892),
112 Mr. F. A. Bather on the
Several specimens were distributed by Barrande under the
manuscript name, Kchinoencrinites multiramus. 'The present
authors place it in or near the Taxocrinide of Angelin,
which is as much as to say that it belongs to the Flexibilia
Impinnata. It is of interest as being older than any Flexible
genus hitherto known, and the interest is enhanced when we
see how its structure accords with its age in the eyes of the
evolutionist (Fig. 5). The Flexibilia Impinnata of the
Silurian rocks fall into four main groups, which may pro-
visionally be regarded as families and named Ichthyocrinide,
Taxocrinide, Calpiocrinide, and Sagenocrinide. The two
latter are clearly more specialized than the two former and
have a larger proportion of representatives in later rocks.
Both Ichthyocrinide and Taxocrinidee have isotomous arms,
which may abut, and in some Ichthyocrinide even interlock,
by their sides. Taxocrinide have a few interbrachials, of
which the proximal is the largest; their anals form a well-
defined vertical series resting on the posterior basal. The
Ichthyocrinide have no interbrachials, and their simplest
genus, Ichthyocrinus, has no anals. Now Caleddocrinus
resembles Ichthyocrinus in the absence of anals from the radial
circlet and in the isotomy of its arms, which are, as in that
genus, sometimes inrolled at their distal ends; but it re-
sembles the Taxocrinide in the presence of interbrachials
with occasional intersecundibrachs, which, however, are all
very small and irregular. The authors believe themselves
able to distinguish an anal interradius by the presence of
3 vertical rows of interbrachials instead of 2. But since
these interbrachials lay in a flexible integument, a greater or
less expansion of the arms would of itself expose more or
fewer interbrachials. However this may be, we have in
Caleidocrinus a genus that approaches the common ancestor
of Ichthyocrinus and Tagxocrinus, although an important point
of divergence, and one by no means primitive, lies in the
minuteness of the basals. All the specimens except one are
assigned to C. multiramus, which has two primibrachs. The
remaining specimen, having 38 primibrachs, is made another
species, C. Barrandet. The authors’ argument is not without
force, but ‘‘ the possibility that more abundant material of the
two species may disclose yet other distinctive characters”
seems to me far from the probability.
We return to the consideration of those members of the
third fauna which are referred either with doubt or certainty
to genera previously known. ‘The first is called Calpioerinus?
bohemicus, or sometimes Calpiocrinus ??? bohemicus (Fig. 6).
The latter mode of expression is preferable, for, as the authors
Lower Palcozote Crinotds of Bohemia. 113
fully recognize, their own interpretation of the fragment does
not agree with Angelin’s diagnosis of his genus. ‘Their
interpretation is that the fragment represents the lower portion
of a cup, consisting of 3 basals, 5 radials, each followed by
about two thirds of a broken primibrach, and 5 large inter-
radials, all but one reaching from the basals to the same level
a b Cc
Fig. 6.—The so-called Calpiocrinus? ? ? bohemicus, after Waagen and
Jahn, pl. lix. figs. 7,8,9; x 4diam. a, “ calice incomplet, vu
d’en bas,” but from above if regarded asa root. 6, from the
side, in what W. & J. consider the normal position, but upside
down if it be a root. ec, “vu d’en haut,” but showing the
surface of attachment to the sea-floor if a root.
as the broken top of the first primibrachs. It is admitted
that all the plates are extremely irregular. This point,
however, need not be laboured, for a glance at Mr. Swoboda’s
drawing of the alleged upper surface (pl. lix. fig. 9; our
Fig. 6) is enough to suggest—nay, to force upon one—a very
different interpretation. The published figures having failed
us so constantly, it may seem the height of rashness to take
their evidence in opposition to the opinion of two eminent
naturalists who have devoted so many years to the study of
these fossils. But no human being could have drawn that
figure 9 without having either before him or in his mind’s
eye the root of some stalked Echinoderm. Here is faithfully
represented the flat surface of adhesion to the sea-floor, with
grooves, like those of JLichenocrinus and other genera,
radiating from the central cavity. Within that cavity are
shown what the authors themselves describe as “ écailles
nombreuses, imbriquées, ornées de cétes fines longitudinales,
et traversées par un grand nombre de petits canaux radiaires.”
This is no description of the thecal cavity of any fossil
crinoid, but is perfectly intelligible in the light of certain roots
well known to geologists in North America. As for an
appearance of pentamerous symmetry in the outer plating,
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 8
114 Mr. F. A. Bather on the
that also is so far from unusual that it has recently led an
American paleontologist to make the remark (extraordinarily
exaggerated no doubt) that “ the root of crinoids appears to
be the counterpart of the calyx in type of symmetry.”
Should this interpretation commend itself to Dr. Jahn, he
will be able to discuss it in that future volume which is to
deal with Loboliths and other crinoid roots.
An imperfect crown from the white limestone, £2, near
Koneprus was labelled by Barrande “ Lecanocrinus bohemi-
cus,” but Messrs. Waagen and Jahn incline to place it in
Ichthyocrinus, to which it certainly presents a closer resem-
blance.
To the long-known genus Scyphocrinus are devoted no less
than sixty-three pages and nearly twenty-two plates, not to
mention those on which are described and figured the nume-
rous fragments probably belonging to it. ‘The more one
appreciates the thoroughness of this piece of work, the more
one regrets that its wonderful array of detail is unprovided
with a summary in the form of diagnoses of the genus and
its species. There is a key to the latter, but it has proved
incomplete and inapplicable in practice. For the generic
diagnosis the reader is referred to the description by Zenker
(1883), which, he is told, “forme une diagnose générique
fort exacte.’ Even had Zenker given a diagnosis, it would
have been framed in accord with the knowledge of his time,
and would have required revision after the discovery of
hundreds of genera unknown to him, some of them closely
allied to Scyphocrinus. Merely to learn that the present
authors refer Scyphocrinus to the Melocrinide (which they
nowhere define) one has to hunt back from p. 73 to p. 11.
But such is the vast plan to which they will be faithful at all
costs !
The Melocrinide belong to that division of Monocyclica
Camerata which has the radials in contact all round. ‘They
are distinguished by having 4 basals; in each half-ray 2-5,
occasionally more, fixed secundibrachs, which support 2 or 4
main rami, giving off pinnules or pinnulate ramuli; nume-
rous and usually definite interbrachials, intersecundibrachs,
and anals; a tegmen of numerous, small, and irregular
plates ; a stem circular in section. The known genera are
Scyphocrinus, Mariacrinus, Melocrinus, and Ctenocrinus ;
and of these the oldest and, in some respects, least specialized
is Scyphocrinus.
Scyphocrinus may be diagnosed as a Melocrinid with a
large number of interbracliials, anals, and intersecundibrachs,
varying both in number and arrangement within a single
Lower Paleozoic Crinotds of Bohemia. 115
species or even a single individual; with the anal area
distinguished only, and that slightly, by its greater width
and greater number of plates; with arms bifurcating fairly
regularly from 3 to 5 times (possibly more), and composed of
simple brachials, which in the free portions are pinnulate and
either uniserial (especially in the proximal region) or alterna-
ting (especially in the distal region), but not biserial in any
authentic specimen; with both the primibrachs, all the
secundibrachs, which are numerous, and the more proximal
tertibrachs loosely joined to those of adjacent rays by a pave-
ment or network of plates.
A very careful study of a large number of specimens
belonging to this variable genus has led the authors to
separate them, according to the ornament of the cup-plates
and especially the interbrachial network, into the following
species and varieties, which form a closely connected series :
Scyphocrinus subornatus; S, excavatus, var. Zenonis, var.
Schlotheimt, var. typica, var. Schroetert; S. decoratus. Except
S. excavatus, all these names are new ; the name S. subornatus
is indeed ascribed to Barrande, but apparently only on manu-
script evidence. One misses the familiar S. elegans, Zenker ;
but this yields to the name Pentacrinites excavatus, which
Schlotheim in 1820 applied to pl. iv. fig. 2 of J. S. Schroe-
ter’s ‘ Vollstiindige HKinleitung &c.,’ 1778, in which work
the honour of original discovery is ascribed to F. Zeno,
Professor at Prague University. The case is, however, no
clearer than usual with these old names. It is admitted that
the details given by Schroeter and Schlotheim do not indi-
cate definitely any one even of the species, much less one of
the varieties, distinguished by the present authors. ‘he
name excavatus has by no previous writer been resuscitated
or applied to Scyphocrinus. One would therefore be inclined
to let it rest and to adopt S. elegans, Zenker, did not Messrs.
Waagen and Jahn assert their inability to find in either
figures or description of Zenker the characters absolutely
necessary for any decision as to which of their species may
claim the name eleyans. Since, moreover, not one of the
numerous writers on this genus has attempted to give pre-
cision to the specific name, the way seems clear for the
present authors to exercise their free choice. ‘Therefore they
make the loosely defined elegans a synonym of the loosely
defined excavatus, and adopt the older term for their largest
species. ‘The chief objection to this proceeding is that
one can after all determine the species to which Zenker
applied the name S. elegans. In the very words of the
volume before us: “l’espéce de Zenker différe notablement
ge
116 Mr. F. A. Bather on the
des Crinoides du Silurien de la Bohéme, désignés par Barrande
sous le nom de Scyph. subornatus, et par nous, sous le noms
de Scyph. decoratus et Scyph. excavatus, var. Zenonis.
Cependant la description de Scyphoer, elegans Zenker con-
vient parfaitement 4 la plupart des spécimens que nous
rangeons dans trois variétés nouvelles: Scyph. excavatus var.
typ., var. Schlotheimi, et var. Schréteri; il y a méme une
concordance remarquable entre les spécimens et les figures
données par Zenker.” Briefly then: Scyphocrinus eacavatus
Waagen and Jahn, non Schlotheim, is identical, even in its
var. typtca, with S. elegans, Zenker. It seems a pity that
our authors did not draw the obvious conclusion, and so
preserve a name in universal use. Had they chosen, “ par
respect pour la mémoire de” Schlotheim, to give the name
excavatus to their new species S. decoratus, none could have
said them nay. However, for the sake of concord, let us
accept the action which the authors themselves have seen fit
to base on their profound research.
To the features of great morphological interest presented
by Scyphocrinus we can here do little more than allude. It
has been supposed that the peculiar network or pavement
which unites the proximal portions of the arms is formed of
modified ramuli; but to this conception the authors do not
refer. The resemblance to fixed ramuli or pinnules, such as
are known in other genera (e.g. Uintacrinus), is more
obvious in the figures of S. ewcavatus varr. typ. et Schroe-
teri than in those of the other varieties and species, some of
which represent only a solid and continuous pavement.
‘There is no geological evidence to show which is the more
primitive type, since, as the authors are careful to point out,
all the varieties are contemporaneous and associated. We
must turn to anatomical and comparative evidence. The
arms of Scyphocrinus are simpler than those of other Melo-
crinide, and have not the ramuli of Mardacrinus and Melo-
crinus. Therefore the structures in question are not so
likely to be modified ramuli as modified pinnules. Two
statements made by the authors suggest possible arguments.
They say (p. 72) “les brachiales de premier ordre [distal
secundibrachs] sont en forme de croissant ou de fer 4 cheval.
L’échancrure .... correspond au sillon ambulacral du trone
des bras, lequel sillon passe dans la voute du calice.” If
branches were given off from this ventral groove to the lines
of plates forming the interbrachial network, this would con-
firm the view that the latter were fixed pinnules. But
specimens examined by me, notably a fine one apparentl
referable to Scyphocrinus excavatus var. Schlotheimi (Brit.
Lower Paleozoic Crinotds of Bohemia. 117
Mus. 15417), show that the supposed ventral groove cuts
into the actual brachials either not at all or only to the
smallest extent, but that the markedly crescentic appearance
is formed by the incurving of the adjoining interbrachials or
fixed pinnules (Fig. 7). This helps to explain the otherwise
Fig. 7.—Scyphocrinus excavatus, var. Schlotheimi, from British Museum
specimen 15417, xX 3 diam. Drawn by Gilbert C. Chubb.
«. Tertibrachs (IIT Br) seen from the inner side of the ray, showing their
relations to the supposed fixed pinnulars (pn) that occupy the
intersecundibrach area. The subhexagonal outlines of the latter
plates are much clearer in such a diagram, where only the sutures
are accurately represented; in a shaded drawing the linear
arrangement of the plates would stand out more clearly. The
latter effect is suggested here by thickening the lines between
the rows.
». The distal surface of the same set of tertibrachs, showing how the
ventral groove (v.g.) is formed by the supposed fixed pinnulars
(pn). Some of these, being broken, do not show the striated
sutural surface.
unintelligible, and doubtless incorrect, fig. 25 on p. 90, pur-
porting to show some arms of Scyphocrinus decoratus. The
other statement alluded to is that the components of the net-
work are joined to the arms and to one another by articular
surfaces. ‘This expression, if used in its strict morphological
sense, would indicate, not merely that the structures were
of brachial origin, but that they had not long become incor-
porated in the cup. Unfortunately for the argument, the
union, at least in the specimens at my disposal, is by loose
suture, with a crenelated edge and probably a striated joint-
surface, just like that between the primary elements of the
cup. On the other hand, since this is also the mode of union
between the fixed brachials, the pinnular origin of the net-
work remains undisproved. The above-quoted statement,
118 Mr. F. A. Bather on the
that the ventral groove passes into the vault (i.e. tegmen),
implies that the distal secundibrachs are still above the level
of the tegmen. If this were so, the plates uniting them could
not be ordinary interbrachials, and their pinnular origin
would be as good as proved. But, since our authors profess
themselves unacquainted with so much as a fragment of a
teemen referable to Scyphocrinus, their rather inexplicable
remark may be dismissed as a vague and inaccurate gene-
rality. The tegmen, if discovered, would certainly throw
light on the question, since, if it could be proved to pass
into the covering plates of the arms and pinnules, and if no
rigid line could be drawn between free pinnules and those
which were partially or wholly united, then the arrangement
would resemble that of Uintacrinus. But if the network
were found to pass gradually into the strictly interambulacral
area of the tegmen, then its plates would have to be regarded
as more or less modified interbrachials. Had this problem
been present to the minds of Messrs. Waagen and Jahn,
they could probably have solved it from their abundant
material. With the help of Mr. Gilbert C. Chubb in pre-
paring the specimen of S. excavatus above alluded to, I have
been able to see that the tertibrachs and proximal quarti-
brachs (beyond which the arms are broken away) are all
fixed by small plates arranged in definite rows; that the
brachials in question are wedge-shaped and alternating, just
as the free brachials of any pinnuliferous arm; and that the
thickened end of each brachial corresponds with a single one
of the rows of plates just mentioned. In other words, the
relation of the rows to the fixed tertibrachs and quartibrachs
is precisely the same as that of pinnules to free brachials.
It is not so easy to trace this relation in the case of the distal
secundibrachs, partly because the plates are here folded and
less regular; partly because the secundibrachs are not low
and wedge-shaped, but are more than twice the height of the
tertibrachs, and are possibly compound ossicles. In this
particular specimen there appear to be 138 secundibrachs:
the lower four are flat cup-plates; the fifth and sixth are
irregular in shape, and the line of the ramus is hard to trace;
from the seventh onward they assume the form of free
brachials. It seems probable that the plates uniting the lower
secundibrachs are true interbrachials, and that their simu-
lation of ramuli or pinnules is due to their axial folding.
The true resemblance to pinnules increases in the more
distal region, although in this specimen, as in many others,
the plates there form a flat pavement. Iam convinced that
the gradual passage of these lines of plates into free pinnules
Lower Paleozoic Crinoids of Bohemia. 119
‘could be traced in specimens that have the arms better
preserved.
It is curious that, in describing some undetermined tegmina
which may possibly belong to Scyphocrinus, the authors
should twice observe that they are unable to see “ l’ouverture
buccale sur le fragment,” especially as they most definitely
refer these specimens to the Camerata of Wachsmuth and
Springer or the equivalent Hypascocrina of Neumayr,
which are characterized by the total absence of a mouth-
opening. Possibly these remarkable sentences are due to the
translator.
Of all the specimens of Scyphoerinus only four retain any
of the stem, and in the most perfect of these it consists of but
7 columnals. Nevertheless there is reason to believe that the
stem often attained a length of many metres. Various roots
are associated with the remains of Scyphocrinus, but the
authors do not feel justified in referring any one of these to
the genus. Some of these roots sprang from the curious
hollow and chambered spheroids known as Loboliths. But
if these cautious paleontologists ever intend to hint that those
bodies may be a part of Scyphocrinus, they have reserved
their remarks for a future volume on roots.
Realizing the futility of discussing, on the evidence of
professedly incorrect figures, those remains which the authors
themselves decline to determine, we may summarize in a table
(p. 120) what this analysis has brought out concerning the
Paleozoic crinoid fauna of Bohemia.
It is probable that all the species are new, but they should
-be compared afresh with known species of the genera to which
they are here referred. The genera, however, are not so
strange as they seemed at first. Bohemicocrinus and Caleido-
crinus may be accepted without hesitation ; Beyrichocrinus
and Laubeocrinus are open to slight question. But if half of
the genera are new, even that is a large proportion ; and since
the authors believe that all records of Scyphocrinus outside
Bohemia are insufficiently supported, five out of the eight
genera may possibly be peculiar to that province.
The authors have nowhere ventured to arrange their genera
in systematic order. The attempt here made, if near the truth,
is rather startling. ‘The extraordinarily large proportion of
Monocyclica and the truly remarkable absence of Inadunata
afford much food for reflection. But considering the rarity of
crinoid-bearing beds in Bohemia, perhaps one should not lay
great stress on negative evidence. Some of the doubtful
remains are of rather Inadunate appearance. ‘The absence of
Monocyclic Adunata is less noteworthy, since this order did
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Lord Walsingham on Asiatic Tortricide. 121
uot originate till Silurian times and was nowhere represented
in large numbers. Flexibilia, too, have not hitherto been
known earlier than the Wenlock Age, so that their rarity in
the lower beds and the appearance of an Jchthyocrinus in f 2
are just what one might expect. The importance of Caletdo-
erinus has already been dwelt on. The Monocyclica Came-
rata, which form the dominant assemblage, present no genus
at all puzzling except Laubeocrinus. The others belong to
widely spread families, and even the new genera among them
are not so distinct as to indicate any great isolation of their
life-province. Indeed, the vertical distribution of the Crinoids
within the Bohemian basin, and such a curious detail as the
fact that all the stem-fragments attacked by parasites (Myzo-
stomidee, the authors say) come from the one band f2, seem
to show that the nature of the genera found was dependent
rather on their selection by a local and temporary environ-
ment than on any wider conditions governing the migration
of faunas or the evolution of orders.
Since the foregoing pages were written the senior author
has passed beyond reach of criticism. I have not, for that
reason, thought it necessary to modify any of my remarks,
since they were never intended to depreciate the labours of
those to whom we are indebted for this exhaustive account of
the Lower Paleozoic Crinoids of Bohemia. It is clear that
the conditions imposed on them rendered their task one of
peculiar difficulty; but the volume they have produced is
none the less worthy of its place in the magnificent work of
Barrande.
Natural History Museum,
London, 8.W.
XIIL.—Astiatic Torrrrerp2.
By the Rt. Hon. Lorp Watsineuam, M.A., LL.D., F.R.S.
(Continued from vol. v. p. 490.]
Carposina, H.-S.
912 (3). Carposina niponensis, sp. n.
Antenne (3) biciliate (3); dirty brownish white. Palp?t
whitish, the median joint shaded externally with fuscous, the
terminal broadly annulate with fuscous at its base. Head
122 Lord Walsingham on Asiatic Tortricidae,
and thorax dirty brownish white. ore wings dirty brownish
white ; with a short triangular basal patch, much widened on:
the costa, pale brownish fuscous, with a darker fuscous spot
of raised scales near the lower extremity of its outer edge ;
four or five ill-defined pale brownish fuscous shade-spots
along the costa, of which one only is before the middle, from
this a diffused shade of scattered fuscous scaling slopes out-
ward to the lower angle of the cell, where there is a small
raised spot, thence a similar diffused shade, much raised at
the upper angle of the cell, is continued to the two outer costal
spots; a broken shade-line is indicated around the apex and
termen; cilia brownish grey. Hap. al.16 mm. Hind wings
‘grey; cilia brownish grey. Abdomen brownish grey. Legs
whitish, with hind tarsal shade-rings.
Type, 3 (70601) Mus. WlIsm.
Hab. JAPAN (Pryer, 1886). Unique.
The colouring of this species is very obscure and ill-defined,
and although it is obviously distinct I should scarcely have
thought it worthy of description had it not been desirable to
indicate this interesting geographical extension of the genus.
PROPEDESIS, gen. nov.
(pomndnois=a springing forward.)
Type Propedesis eacisa, Wlsm.
Antenne (2?) simple. Labial palpi projecting nearly
twice the length of the head beyond it; terminal joint short,
median joint somewhat triangular owing to upstanding scales
above at the middle and depressed scales below at its apex.
Head densely clothed above. Thorax smooth. Fore wings
narrow, elongate ; costa slightly arched near the base, straight
beyond; termen very oblique, sinuate. Newratzon, 12 veins ;
2 from very near angle of cell; 3 and 4 closely approximate
at angle of cell; 7 to termen; 8 and 9 stalked, 8 to a little
above termen. Hind wings with the costa straight, apex
produced, termen sinuate, flexus rounded, median vein with
long loose hairsabove. Neuwration, 6 veins; 3 and 4 stalked.
Abdomen smooth. Legs with slight projecting scales at hinder
extremity of tibia.
The genus is closely allied to Carposina, H.-8., from which
it is distinguishable only by the stalking of veins 8 and 9 of
the fore wings and the somewhat more produced apex of the
hind wings. The male will probably have biciliate antenne
and short palpi.
Lord Walsingham on Asiatic Tortricide. 123
912 (4). Propedests eacisa, sp. n.
Antenne dirty white, annulate with fuscous towards the
base. Palpi white, black beneath and at the base. Head
and thorax white. Fore wings with the termen very oblique,
strongly sinuate beneath the produced apex; white, dusted
with greyish fuscous, the markings dark greyish fuscous; a
triangular basal patch, wider on the costa than on the dorsum,
is followed by a costal spot before the middle, succeeded by
three smaller ones beyond the middle; an oblique patch at
the outer end of the cell is parallel to the outer edge of the
basal patch, it reaches down to the fold and upward nearly to
the costal spot, and is accompanied by tufts of raised scales,
as is also the outer edge of the basal patch itself; there is a
series of spots along the termen, with a parallel shade-line
faintly indicated before them; cilia brownish cinereous.
Ezp.al.20 mm. Hind wings with strongly produced apex,
the termen more deeply excised than in the fore wings;
brownish grey ; cilia brownish cinereous. Abdomen brownish
grey. Legs dirty whitish.
' Type, 2 (70602) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886). Unique.
912 (5). Propedesis japonica, sp. n.
Antenne dirty white, very faintly annulate. Palp?
whitish, shaded externally with greyish fuscous. Head and
thorae whitish, tinged with greenish grey. ore wings
greenish white, sparsely dusted with olive-grey, with a trian-
gular patch of pale olivaceous grey at the base, wider on the
costa than on the dorsum, its outer edge consisting of raised
scales; a slight olive-grey shade-spot on the costa before the
middle, followed by three others beyond it; an oblique
fuscous streak at the end of the cell, consisting of raised scales,
and preceded by other tufts of raised scales (white dusted with
fuscous), one at the upper edge of the cell, one resting on the
middle of the fold, another preceding it below the fold; the
termen shows a faint indication of greyish marginal spots
preceded by a parallel greyish shade; cilia pale cinereous.
Exp. al.20 mm. Hind wings pale rosy grey ; cilia greyish
cinereous. Abdomen cinereous. Legs whitish.
Type, 2 (70603) Mus. Wlsm.
Hab, JAPAN (Pryer, 1886). Unique.
This species differs from e#cisa in the different outline of
the wing as well as in the markings; both in the fore and
hind wings the apex is less produced and less acute, the
termen being only very slightly sinuate.
124 Lord Walsingham on Asiatic Tortricide.
RuyYAcioniA, Hb.
Type Zortrix buoliana, Schiff. (Stph. 1834).
915. Rhyactonta duplana, Hb.
Retinia duplana, Stgr. & Wk. Cat. Lp. Eur. 246. No. 915 (1871) *;
Fern. Tr. Am. Ent. Soc. X. 27. No. 158 (1882) °.
Hab. Europe}. JAPAn (Pryer, 1886)—Howpo—Yoko-
hama (Manley, 1888). Unirep States—Oregon *.
919. Rhyacionia turionana, Hb.
Retinia turionana, Stgr. & Wk. Cat. Lp. Eur. 246, No, 919 (1871)*;
Fem, Tr. Am. Ent. Soc. X. 27. No. 157 (1882) ?.
Hab. Evrorr’. Japan (Pryer, 1886). UNITED STATES
—Nevada’.
921. Rhyacionia buoliana, Schiff.
Retinia buoliana, Steger. & Wk. Cat. Lp. Eur. 246, No, 921 (1871) 1.
Hab. Europe’. SrpertA'. CoreaA—Gensan, VI. 1886
(Leech).
923 (1). Rhyacionia retiferana, Wk.
Retinia *margarotana, Hein. Schm. Deutsch. Tortr. 95-6. No. 168
(1863). Retinia retiferana, Wk. Zts, Ent. Bresl. (n. s.) VII. 73
(1879). Retinta *margarotana, Brt. Ent. Mo. Mag. XXVI. 49
(1890); Hdgn, Ent. XXIII. 119 (1890). Retina retiferana, Brt.
Ent. Mo. Mag. XXIX. 113-4 (1893); Rag. Ann. Soc. Ent. Fr.
LXIII, 200. No. 923 bis (1894).
Hab. Evrore. JAPAN (Pryer, 1886).
EXARTEMA, Clem.
924 (1). Exartema castaneanum, sp. n.
Antenne brownish grey. Lalpi whitish ochreous, with
two fuscous spots on their outer sides. Head and thorax
brownish grey, the tegule and patagia paler. Fore wings
rich chestnut, with shining leaden grey markings narrowly
outlined with pale ochreous and some brownish fuscous
shades; a basal patch extending to one-third is mottled with
brownish fuscous, leaden grey, and chestnut, its outer edge
straight, but with a slight outward angle below the costa, and
bounded by a leaden grey band narrowly outlined with pale
ochreous; a leaden grey patch at the tornus sends two
branches inwards from its upper extremity, the upper one
Lord Walsingham on Asiatic Tortricide. 125
moderately straight, the lower one angulated downward at its
middle, all narrowly outlined with ochreous ; there are five
pairs of outwardly oblique pale ochreous costal streaks beyond
the middle, alternating with brownish fuscous, the first pair
emitting a sinuate leaden grey line which reaches the termen
above the middle; cilia brownish ochreous, with a fuscous
spot at the apex and another on the middle of the termen.
Eap. al. 15-17 mm. Hind wings greyish brown; cilia pale
ochreous, a dark dividing line running through them near
their base. Abdomen greyish brown. Legs ochreous, shaded
and banded with greyish brown.
Type, & (60792, Kiusiu) ; ¢ (60793) Mus. Wlsm.
Hab. CorrEA—Gensan, VII.-IX. 1887 (Ito). JAPANn—
Krusrv (Leech, 1890). Five specimens.
Allied to the American Hvartema ferrugineanum, Riley,
but the only specimen I have of this differs decidedly in the
position of the first geminated. costal streak beyond the
middle, the leaden line extending to the termen being much
shorter and coming from the second instead of from the first
pair,
924 (2). Hwartema velutinum, sp. n.
Antenne fuscous. Palpi dirty whitish ochreous. Head
and thorax fuscous. ore wings dark brownish fuscous,
bright reddish ferruginous towards the apex and below the
middle of the termen, with a pale brownish olivaceous
ochreous patch on the middle of the dorsum, mottled on the
basal patch and on the rather oblique median band with steel-
blue; with seven pairs of pale whitish ochreous costal
streaklets, of which two only are before the middle, the first
pair beyond the middle giving out a sinuous steel-blue line to
above the middle of the termen; a steel-blue patch at the
tornus sends inward two branches from its upper extremity,
forming the blue mottling on the dark band; cilia ochreous,
tessellated with fuscous, a fuscous line along their base.
Eep. al. 15 mm. Hind wings rather deeply emarginate
below the lobe in @; greyish brown; cilia pale cinereous, a
dark shade-line near their base. Abdomen greyish brown.
Legs greyish, banded with pale cinereous.
Type, & (60459, Mupin); ¢ (60796) Mus. Wlsm.
Hab. W. Cuina—Mupin, 21 V., 28 V. 1890 (Leech).
CoreA—Gensan, VI. 1886 (Leech), VII.-IX. 1887 (Ito).
JAPAN (Pryer, 1886)—Krusrv (Leech, 1890)—Satsuma, V
1886 (Leech). Hight specimens,
126 Lord Walsingham on Asiatic Tortricidae.
924 (3). Exartema transversanum, Chr.
Penthina transversana, Chr. Bull. Soc. Imp. Nat. Mose. LVI. 75-7.
No, 108 (1881) ': sep. 163-5 (1882) '.
Hab. AMur—Pompejefka, VII.’ ; Wladiwostok', 15 VIL.,
3-4 VIII. 1877 (Christoph). CoreA—Gensan, VI. 1886
(Leech). CHiInA — Chang Yang, 4000-6000 feet (Pratt,
1886). JAPAN (Pryer, 1886)—Yxsso—Hakodate, VIII.
1886 (Leech) ; Howpo—Tsuruga, VII. 1886 (Leech).
924 (4). Hxartema japonicum, sp. n.
Antenne cinereous. falpi whitish cinereous. Head cine-
reous. Thorax mixed whitish cinereous and olive-brown.
Fore wings shining whitish cinereous, with olive-brown
streaks and blotches; the first series bounding a much reticu-
lated basal patch, wider on the dorsum than on the costa, and
containing an admixture of steel- grey scales ; the second series
forming a moderately straight transverse fascia in the middle
of the wing, wider on the dorsum than on the costa and
bulging outwards in the middle in two large irregular ex-
crescences, broken up with dark fuscous and shining steel-
erey scaling ; an olive-brown spot before the tornus is more
or less connected by a pale olivaceous shade with an elongate
oblique olivaceous patch above it, which again is connected
with some oblique streaks of the same colour from the costa,
alternating with creamy white geminated streaks, each divided
by steel-grey lines which run together to the middle of the
termen ; the apex is olive-brown ; cilia pale whitish ochreous,
with an olive-brown line along their base; the whole of the
paler portions of the wing are rendered more or less lustrous
by the admixture of steel-grey scaling. Hp. al. 20 mm.
Hind wings dark brown; cilia shining whitish cinereous, with
a dark line through them near their base. Abdomen greyish
brown, anal tuft cinereous. Legs cinereous.
Type, & (70537); ¢ (60392) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886)—Howpo—Tsuruga, VII. 1886
(Leech). Four specimens.
924 (5). Hxartema Pryeranum, sp. n.
Antenne brownish cinereous. Palpi pale brownish cine-
reous, with some fuscous scales. Head and thorax brownish
fuscous, mixed with brownish cinereous. ore wings
brownish cinereous, much shaded across the middle and on
the basal patch, and sprinkled toward the apex with brownish
fuscous; a reduplicated narrow leaden grey band at one-
Lord Walsingham on Asiatic Tortricide. 127
third, narrowly outlined with brownish cinereous, ends in two
pairs of brownish cinereous costal streaklets, and from a little
beyond the middle a similar band tends obliquely outward
from costa to tornus, sometimes broken about its middle,
where it sends a branch obliquely inward to the dorsum ;
the shape of this band seems to be somewhat variable, but at
its upper-extremity is a pair of costal streaklets, and these are
succeeded by three other pairs before the apex, which tend to
converge upon a sinuate leaden grey line running toward the
middle of the termen ; cilia pale brownish cinereous, shaded
along their base and on their outer half with fuscous. up.
al. 19 mm. Hind wings dark greyish brown; cilia pale
brownish cinereous, with a shade-line near their base. Adbdo-
men greyish brown. Legs pale cinereous, with slightly
spotted tarsi.
Type, 3 (70542); ¢ (70546) Mus. Wlsm.
Hab. Jaran (Pryer,1886)—//onpo —Oiwake ( Pryer, 1885).
Four specimens.
924 (6). Hxartema fasciatanum, Clem.
Exartema fasciatana, Clem. Pr. Ac. Nat. Se. Phil. XII. 357 (1860) ’.
Servaphila decisana, Wky. Cat. Lp. Ins. B. M. XX VIL. 340-1 (1863) ?.
Evartema fasciatana, Why. Cat. Lp. Ins. B.M. XXX. 994 (1864) *.
Sericoris fasciatana, Clem. Pr. Ent. Soc. Phil. V. 134 (1865) *. Zv-
artema albofasciatum, Z. Verh. ZB. Ges. Wien, XXV. (1875) Abh.
272-3 (1875) °. Exartema fasciatana, Z. Verh. ZB. Ges. Wien, XXV
(1875) Abh. 273 (1875) °. Eccopsis fasciatana, Fern, Tr. Am. Ent.
Soc. X, 80. No. 177 (1882)7; Mfft. Can. Ent. XIX. 88 (1887) °.
Hab. JAPAN (Pryer, 1886) — owpo— Yokohama (Manley,
1888). CaNaDA—Ontario®, UNITED Stares**— Maine’,
Massachusetts *, New York", Pennsylvania’, Maryland (Mus.
de Joannis), Ohio’, North Carolina (Morrison), IUlinois’.
924 (7). Hwxartema semicremanum, Chir.
}Penthinea semicremana, Chr. Bull. Soc, Imp. Nat. Mose. LVL. 77-8.
No. 109 (1881) !: sep, 165-6 (1882) *. Penthina semicremana, Rag.
Ann. Soe. Ent, Fr, LXIII. 203, No, 949 dcs (1894) *.
Hab. AMUR—Pompejefka, VII.', 13 VIL. 1876 (Christoph).
JAPAN— Yzsso (Pryer, 1882).
OLETHREUTES.
924 (8). Olethreutes arcuella, L.
Penthina arcuella, Stgr. & Wk. Cat. Lp. Eur. 249, No. 980 (1871);
Stgr. Hor. Soc. Ent. Ross. XV. 250 (1879) *. Eucosma arcuella,
Meyr. HB. Br. Lp. 465 (1895) ’.
Hab, Europe’, Astatic TURKEY — Auvpavenpix14ar—
a
128 Lord Walsingham on Asiatic Tortricide.
Brussa, VI2 Srperta*. CorEA—Gensan, VI. 1886
(Leech). JAPAN *—Iowpo—Oiwake, VI. 1885, VI-VII.
1887 (Pryer).
The specimens of this species from Corea and Japan differ
from our English form in their larger size and paler colouring
on the underside, but precisely correspond in their markings.
The males all have a small hanging appendage near the base
of the limbus in the hind wings, a character hitherto confined
to the genera Evrartema, Clem., and Cymolomia, Ld. The
discovery of this character in Japanese specimens caused me
to re-examine carefully a series of British specimens of
arcuella, which has been placed by different Kuropean authors
in Penthina, Ty., and Sericoris, Tr. It was at once apparent
that all the males in the series possessed a very small rudi-
mentary appendage, scarcely noticeable unless special atten-
tion had been called to it. The stronger development of this
character in Japanese specimens is scarcely sufficient to justify
their description as a distinct species, especially as I find
German specimens of intermediate size and decidedly paler
on the underside than our British examples.
Meyrick, whose attention had been called to the existence
of the lobe, wrote (HB. Br. Lp. 465) that this was so rudi-
mentary that it appeared insufficient for generic separation.
In this I am unable to agree with him, for it is certainly a
structural character, corresponding with that which was relied
upon by Clemens when describing the genus Hvartema.
The question is one of degree, but it may fairly be argued
that the possession of a structural character rather than the
mere degree of its development is the true test of generic
yank ; and Meyrick has recognized this principle very forcibly
in including Péycholoma, Stph., and Lozotenia, Stph., in the
genus Cacecia, Hb., the only cause of separation by the
older authors having apparently been the inferior development
of the costal fold.
If Meyrick is justified in placing arcuella in the same
genus as many other species which possess no limbal fold or
lobe, it follows that the Japanese form (specially inseparable,
as I think, from arcwella) would fall into a different genus
through the slightly greater development of the limbal lobe,
unless he is prepared to include with it Hxartema, Eccopsis, &e.
Clemens, in describing Hwartema, clearly refers to the
hind margin being deeply and sharply excised opposite to the
median nervules. He described five species: the first two
only, viz. nitidana and permundana, possess this character ;
the other three species, versicolorana, inornatana, and fascia-
tana, have the dorsum evenly rounded beyond the flexus,
Lord Walsingham on As/atic Tortricide. 129
although there is an excavation on the limbus as well as the
tubular lobe at the base. In the outline of the wing these
three species agree with Cymolomia, Ld., rather than with
Olethreutes, which possesses the lobe without any excavation
of the limbus, but the American species differ from Cymo-
fomia in the tubular structure of the lobe. Thus we have in
this group four published genera, which can be distinguished
by structural differences :-—
1. Eccorsis, Z.—Limbus rolled at base, without lobe, with
strongly angular limbal excavation and deep dorsal
excavation beyond the flexus.
2. ExarTEMA, Clem.—With partially free tubular lobe from
base of limbus, moderately excised limbal margin, and
excised dorsum beyond the flexus.
3. CymotomiA, Ld.—Limbus rolled at the base, without free
lobe ; limbus excised, dorsum not noticeably excised.
4, OLETHREUTES, Hb.—Limbus rolled, with slightly deve-
loped lobe ; limbus and dorsum not excised.
924 (9). Olethreutes notata, sp. n.
Antenne cinereous, basal joint yellow-ochreous. Pa/pi
pale whitish ochreous, shaded externally with fuscous. Head
yellow-ochreous, shaded with fuscous. Zhorazx bright yellow-
ochreous, mottled with fuscous. Fore wings bright yellow-
ochreous, reticulated on the basal half and deeply suffused
on the outer half with blackish fuscous, and with numerous
spots and short cross-bars of bright shining steel-blue; the
spots for the most part being evenly distributed over the
basal half, and with three bars and some spots on the outer
half, one below the termen at the commencement of the
apical third, one above the tornus, and one within and below
the apex, the two outer moderately straight, the inner slightly
oblique, except where the deep outer shade touches the costa
immediately beyond the middle; the edges of the wing are
uniformly bright yellow-ochreous, mottled and spotted with
dark fuscous; cilia greyish fuscous, with a broad pale whitish
ochreous interruption below the apex, a black line along their
base. Hap. al. 19-20 mm. Hind wings dark brown; cilia
pale whitish ochreous, a dark line running through them
near their base. Abdomen brownish. Legs grey.
Type, 3d (70014); 2 (70016) Mus. Wlsm.,
Hab, JAPAN—Yesso (Pryer, 1882). Six specimens.
This species has a very rudimentary excrescence near the
base of the limbus of the hind wings, scarcely more than a
thickening of the cuticle on the margin itself, in this respect
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 9
130 Lord Walsingham on Asiatic Tortricidae.
agreeing with the European form of arcuella rather than with
Eccopsis, in which this character is much more strongly
developed.
PHZCADOPHORA, gen. nov.
(patxads = a woolly shoe; gopeiv = to carry.)
Type, o 3, Phecadophora fimbriata, W\sm.
Antenne (6) slightly ciliate. Palpt (¢) short, projecting
only the length of the head beyond it; terminal joint very
short, almost concealed in the strong tuft above and below
the median joint: ¢ longer than the ¢, more porrect, ter-
minal joint more apparent and slightly depressed. Head
with a strong erect crest. Jhoraxv with a slightly erect tuft
posteriorly. Fore wings narrow, elongate, costa gently
arched, ¢ without a costal fold; apex acute or abruptly
angulated; termen straight or slightly impressed, not convex ;
tornus rounded, dorsum straight. Newration, 12 veins, all
separate ; 2 from middle third of cell; 7 to termen; 10 not
nearer to 11 than 9. Hind wings much broader than the
fore wings, more or less trapezoidal, apex rounded, termen
slightly indented below it; ¢ with a strong pencil of hairs
near the limbus on the upperside anda fringe of long hairs
on the underside between the branches of vein 1. Neuration,
8 veins; 3 and 4 connate; 5 bent over to near origin of 4;
6 and 7 separate but parallel and closely approximate at
their kase. Legs, hind tibiz and tarsi strongly clothed above
with long tufts of woolly scales.
This genus differs from Phecasiophora, Grt., in its nar-
rower fore wings and in the hind tarsal joints as well as the
tibiae being clothed with thick hair-tufts, as well as in the
more strongly developed fringes of hairs above and below
the hind wings near the limbus.
924 (10). Phecadophora fimbriata, sp. n.
Antenne and palpi tawny cinereous. Head and thorax
dark tawny vinous red; tegule pale cinereous, shaded with
reddish grey, with a strong raised tuft of vinous red scales
on the thorax posteriorly. Fore wings whitish stone-colour,
thickly and slenderly streaked throughout with rosy pink
and olive-brown ; costa with oblique striz throughout its
length, a pair before the apex turning downwards and meet-
ing the end of the oblique streak which precedes them; at
the end of the cell are a few blackish scales, and along the
dorsum a conspicuous dark band of tawny brownish, bluish
fuscous on the extreme margin, with a blackish brown line
Lord Walsingham on Asiatic Tortricide. 131
running along its middle and bent obliquely upwards at two-
thirds from the base, where it is bounded on each side by a
narrow margin of the pale ground-colour of the wing, running
to a point at the lower angle of the cell, the upper edge of
this dorsal band is somewhat waved, a slight bulge occurring
before the middle of the wing, its upper edge carrying a
reduplication of the blackish brown central line; about the
tornus the dorsal band is continued after a slight interruption,
and here also it contains some blackish brown lines; an
elongate olive-brown shade lies below the outer half of the
discal cell, with some lines of the pale ground-colour running
through it; above the tornus is a small shining whitish
ocelloid spot, tinged with rosy pink scales immediately above
the angle; cilia beautifully mottled with olive-brown, silvery
whitish, greyish fuscous, and rosy pink, the greyish fuscous
being at their tips on the middle of the termen, also at the
extreme apex and at the tornus, the rosy pink coming above
and below the middle and also on the outer half of the cilia;
the reddish brown is situated on the basal half of the cilia
on the upper half of the margin, and the silvery whitish is
immediately below it in a small spot also en the basal half
of the cilia. zp. al. 20mm. Hind wings dark brewn, on
the upper surface of vein 2 is a distinct fringe of brownish
gvey hairs; cilia shining whitish, with a dark line running
through them near their base. Abdomen brownish grey.
Legs greyish white, clouded with greyish fuscous.
Type, 3 (70257, Japan) ; 2 (70249) Mus. Wlsm.
Hab. Japan (Pryer, 1886)—Krusiw (Leech, 1890). As-
saAmM—Margherita (Doherty, 1889). E. Peau—Kuarew Hizrs
—Thandsung, 4000 ft. (Doherty, 1890). ‘Twenty-four
specimens.
This description is taken from a male in beautiful condition,
which exhibits some differences from others in the series. |
find that in one variety (40124) the longitudinal streaks are
more distinctly carried through the wing and there is more
olive-brown alternating with longitudinal lines of fuscous,
the rich rosy tint being chiefly confined to the space below
the costa, especially in a long streak beneath the outer half;
the black scales at the end of the cell are almost obsolete, and
the cilia, although they exhibit somewhat the same pattern,
are not so rich in colouring. It is evidently a variable
species.
924 (11). Phecadophora (?) acutana, sp. n.
Antenne pale stone-grey, faintly spotted above. Palpi
long, porrect, extending twice the length of the head beyond
Ox*
132 Lord Walsingham on Asiatic Tortricide.
it, thickly clothed to the base of the exposed terminal joint,
above and beneath; light stone-grey. Head light stone-
grey. Zhorax pale stone-grey; tegule with two brownish
spots. Fore wings elongate, apex slightly produced, termen
slightly concave, scarcely oblique, angulated at the extre-
mity of vein 3, thence very oblique to the dorsum, which is
straight; stone-white, streaked throughout with pale olive-
brown; a series of short oblique streaklets along the costa,
olive-brown mixcd with biownish fuscous scales; an olive-
brown spot at the apex is followed by a greyish fuscous spot at
the base of the cilia immediately below it ; a small patch of
greyish fuscous at the base of the dorsum, followed by some
scattered specks of the same colour, especially along the
lower half of the wing to the tornus, and a small streak of
brownish fuscous scales at the angle of the termen on vein 3 ;
cilia shining stove-white, inclining to olivaceous about the
tornus. Lap. al. 21mm. Hind wings brownish fuscous,
costa whitish; cilia greyish fuscous, a dark line running
through them near their base. Aldomen greyish fuscous.
Legs stone-white, yellowish white on their underside; hind
tarsal joints somewhat densely clothed, but not so thickly as
in the male of the allied species fimbriata.
Type, 2 (60191) Mus. Wlsmn.
Hab. Japan — Kivsrv — Satsuma, V. 1886 (Leech).
Unique.
I should have regarded this as a variety of jimbriata had
it nct been for its much longer palpi and the absence of the
dark dorsal band. The male is at present unknown, but
the generic determination is almost certainly correct.
924 (12). Pheecadophora (?) divisa, sp. n.
Antenne, palpi, and head cinereous. Thorax mixed
greyish cinereous and dark reddish fuscous. Fore wings
with the dorsal half pale cinereous, slightly mottled with
greyish fuscous, the costal half dark reddish fuscous; the
colour on the costal half extends to the fold at the base, is
slightly indented before the middle, and diminishes obliquely
from about the lower angle of the cell to the apex; along
the extreme costa is a series of very small greyish cinereous
geminated dots, with a slight tawny gloss, connected with
the three pairs which precede the pair nearest to the apex;
on the pale half of the wing is an oblique dash from the
dorsum before the tornus, pale chestnut-brown, containing
five or six short blackish tuscous transverse dashes; at its
upper end this streak is connected with a brownish fuscous
Lord Walsingham on Asiatic Tortricide. 133
spot, about the middle of the termen, which sends a dark
shade through the whitish ochreous cilia; below this spot
the space within the tornus inclines to whitish grey, and
above it the triangular space between the dark costal shade
and the upper half of the termen is white, a narrow dark line
running around the termen at the base of the cilia; on the
underside the geminated streaks are distinctly visible on the
outer half of the costa. Hxp. al. 22mm. Hind wings dark
greyish fuscous; cilia tipped with greyish. Abdomen dark
greyish fuscous. Legs cinereous, clouded with fuscous on
the hind tibiz and tarsi.
Type, 2 (40228) Mus. Wlsm.
Hab. AssAM—Nae@a Hirrs—Gologhat (Doherty, 1890).
Unique.
The termen of the fore wings in this species differs from
that of acutana in not being more distinctly impressed below
the apex than in the typical species. The ¢ is unknown,
but it will probably be found to be congeneric with fimbriata.
924 (13). Phecadophora (?) ochracea, sp. n.
Antenne cinereous. Palpi whitish ochreous, the fringes
at the end of the median joint and the terminal joint purplish
fuscous. Head fuscous. Thorax cinereous. Fore wings
with the termen slightly impressed beneath the apex; pale
ochreous on the dorsal half, dark tawny red on the costal
half, a dark purplish fuscous shade overspreads the basal
and lower portions of the costal half, and on the costa is a
series of pale ochreous geininated streaks terminating (espe-
cially those towards the apex) in very oblique pale steel-grey
lines; the pale ochreous dorsal colouring extends to the costa
at the extreme base and separates the costal shade from the
told for about one-fourth the wing-length; it also encroaches
upon it by a small angular excrescence at about one-third,
from a little beyond the middle ot the wing it extends obliquely
across the fold in a straight line to the extreme apex; a
slight brownish ochreous shade passes upwards trom the
tornus direct, not obliquely, and contains three or four dark
fuscous dots, this shade extends to the upper half of the
termen; there is also a shade of a similar colour below the
middle of the fold, with a small patch of fuscous scales at the
extreme base of the dorsum; cilia pale ochreous, shaded with
brownish ochreous at their middle, and with greyish fuscous
for a small space above the middle of the termen; under-
side with the costal spots showing distinctly throughout.
Exp. al. 20 mm, Hind wings dark brownish fuscous ; cilia
134 Lord Walsingham on Asiatic Tortricidae.
greyish. Abdomen dark greyish fuscous ; anal tuft ochreous.
Legs whitish ochreous, shaded with greyish ochreous, hind
tarsal joints spotted.
Type, 2? (60421) Mus. Wlsm.
Hab. CorrEA—Gensan, VII.-IX. 1887 (Ito). Unique.
Dupua, Wkr.
Dupva, Wkr. Cat. Lep. Ins. B. M. XXX. 1000 (1864).
Dudua hesperialis, Wkv.
Dudua hesperialis, Wkr. Cat. Lp. Ins, B. M. XXX. 1000 (1864) }.
Antenne dark brownish fuscous. Palpi short, porrect,
teiminal joint almost concealed; bluish grey. Head very
dark brownish fuscous. Thorax with an erect crest poste-
riorly ; very dark brownish fuscous; tegule spotted with
steel-blue. Fore wings dark brownish fuscous, freely spotted
with shining steel-blue ; before the middle of the costa are
three pairs of steel-blue streaks, touched with whitish at their
upper ends; on the extreme costa beyond the middle are
four pairs of oblique geminated whitish streaklets, with a
single one, not oblique, before the apex; of these the first
pair, and the third pair (counting from the middle of the
wing), send out oblique lines of steel-blue towards the termen ;
a steel-blue band from the tornus, running upwards for more
than half the breadth of the wing, is preceded and followed
by some patches of cinereous scales, which also extend up-
wards along the termen; the remainder of the lower half of
the wing, with the exception of a space above the middle of
the fold, is freely sprinkled with steel-blue spots; these are
more thickly grouped beyond the middle than before it,
except perhaps at the extreme base ; cilia shining steel-blue,
tipped with fuscous, at the apex dark fuscous, ap. al.
19 mm. Hind wings greyish brown; cilia pale cinereous,
a dark shade running through them near their base. Abdo-
men greyish brown. Legs pale cinereous; posterior tarsal
joints shaded and banded with greyish fuscous.
Type, & (Saunders Coll.), Mus. Br.—acquired 1894.
Hab. Japan (Pryer, 1886)—Kivsrv—Satsuma, V. 1886
(Leech). Cutna—Chang Yang, 4000-6000 tt. (Pratt, 1886).
BorRNEO— Sarawak |.
The above description, taken from a female from Satsuma
(60012), was made betore the rediscovery of the missing
types ‘(in Mr. Saunders’s Collection.” I have only females
and therefore cannot determine whether the genus Dudua is
Lord Walsingham on Asvatic Tortricidae. 130
distinct. I noted that the type, which is unset, had broadly
fringed hind tibiz, and Mr. Durrant, who drew the neuration,
thought that the flexus of the hind wings was probably some-
what bulged as in Phecadophora. At present it cannot be
determined whether Dudua should be referred to Phecasio-
phora, Grt., or to Phaecadophora, Wism., or whether it is
distinct from both.
PHACASIOPHORA, Grt.
924 (14). Phecasiophora Fernaldana, sp. n.
Antenne reddish fuscous. Palpt reddish brown at the
sides, paler towards the apex. Head reddish brown. Thorax
rosy fawn, mottled with umber-brown. Fore wings rosy
fawn, with a short basal patch and three irregular fasciaform
markings all confined to the costal side of the fold; these
markings are all broken up and jagged at the edges and
consist of greyish fuscous and olive-brown, margined and re-
ticulated with dark umber-brown ; the short basal patch is
narrow on the costa, dilated and angulated on the upper edge
of the cell, its lower margin not reaching the fold and ill-
defined; the first fasciaform band is, at one-third the wing-
length, broken into two patches, the first adjacent to the
costa, irregularly quadrate, the other beneath it, adjacent to
the fold, somewhat reniform; the second fascia, beginning
scarcely beyond the costa, is wider and more conspicuous
than the other two, and sends out three excrescences on its
outer side and two slight projections on its inner side, the
pale ground-colour between the outer excrescences gives the
appearance of a spot (or reduplicated spot) about the upper
angle of the cell; the third fascia does not reach the costa,
but is curved from near its outer fourth, first outwards towards
the apex, then downwards in the direction of the tornus,
sending out two projections on its outer side towards the
termen; between this fascia and the apex of the wing is a
slight grey shade, followed by a slender dark umber-brown
line and a small triangular apical spot; the extreme costa is
narrowly pale fawn-colour, with numerous short striae, some
oblique, some almost straight; the rosy fawn portions of the
wing between the tasciz, as well as the dorsal fourth of the
wing-width, are clouded and reticulated with grey and greyish
fuscous and with a small proportion of umber-brown; cilia
pale fawn, with a narrow brownish line near their base and
three or four greyish spots running through them at the
apex and on the termen, Hap, al. 22-24 mm. Hind wings
136 Lord Walsingham on Aszatic Tortricide.
brownish fuscous, inclining to whitish on the costa and
limbus; cilia pale cinereous, a dark line near their base.
Abdomen greyish fuscous, anal tuft cinereous fawn. Legs:
¢ with the posterior pair densely and widely tufted above
with long white hair-seales ; posterior tarsal joints lightly
spotted with greyish fuscons above.
Type, 3 (70279, Japan); 2 (70275) Mus. Wlsm.
Hab. CorFA—Gensan, VII.-IX. 1887 (Jio). Japan
(Pryer, 1886). Sixteen specimens.
‘This species is larger than the American confizana, Wkr., .
which it greatly resembles, although the actual form of the
markings is somewhat different, the pattern appearing fairly
constant through a considerable series of specimens.
924 (15). Phacasiophora Pryert, sp. n.
Antenne pale brownish fawn. Pa/pi whitish fawn, touched
with brownish externally. Head brownish fawn. Thorax
evenly mottled with whitish fawn and brown. Fore wings
whitish fawn, divided into innumerable streaks and reticu-
lations by brown spots, lines, and patches; a basal pateh
oceupying about one-third of the wing is but faintly indi-
cated by three brownish spots in a fasciaform series, each
margined with umber-brown, and is much reticulated through-
out; from the middle of the costa is a large brown bloteh
shaded with umber-brown, and another below it, not crossing
the fold, and partly connected with the first, at its upper
edge this patch is also clouded with dark umber-brown and
sends out two exerescences on its outer side, these are out-
lined by whitish fawn streaks enclosing shining bluish grey
scales and forming an elongate whitish fawn spot between
the excrescences, extended outwards and downwards; beyond
them is a curved patch of fawn-brown, through whieh run
six or more transverse lines of dark umber-brown; this patch
is narrowly connected with the termen about its middle, and
on its upperside is bounded by a long eurved whitish fawn
line, reaching from the dark costal pateh to the middle of
the termen, having some shining grey scales along its upper
edge; above it are five pairs of pale costal geminations,
separated by brown spaces, the apex itself is brown, a pale
waved line running from the last costal gemination nearly to
the middle of the termen; there is also a rather incon-
spicuous brown patch on the dorsum before the tornus ; cilia
reddish fawn, a reddish brown shade running throughout’
them near their base, and some greyish spots about the apex
and termen. ap.al. 20mm. Hind wings dark greyish
On Giant Squirrels from the Amazonian Region. 137
brown; cilia whitish cinereous, with a greyish brown line
near their base. Abdomen greyish brown, anal tuft inclining
to ochreous. Legs: 3, posterior pair, densely and widely
tufted above with long whitish cinereous hatr-scales ; pos-
terior tarsal joints faintly spotted with pale brownish,
Type, & (40270) ; 2 (70269) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886)—Horvo—Foochau, IV. 1886
(Leech). Nine specimens.
X1V.—On Giant Squirrels from the Amazonian Region.
By OLDFIELD THOMAS.
TuE British Museum has received during the last two or
three years a number of Giant Squirrels belonging to the
Sciurus Langsdorfi group from several different localities in
South America. On determining these one proves to need
description, while additional localities may be recorded for
others.
During the progress of this work Dr. Kiikenthal, of Bres-
lau, has been good enough to render me valuable assistance
by the loan of the original type of S, tricolor, Péppig, from
Maynas, Peru, without which it would have been impossible
to determine to what form that name should be applied.
Scturus igniventris, Wagn.
Nericagua and Munduapo, Upper Orinoco (G. K. Cherrie) ;
Bogota (G@. D. Child).
‘The type locality of this species is Marabitanas, on the
Upper Rio Negro, not far from the latter river’s junction with
the Upper Orinoco. The further extension of the species
westwards to Bogota follows the line of the Rio Meta, which
falls into the Orinoco in the same district that Mr. Cherrie
collected in. South-eastwards the species ranges along the
Rio Negro to its mouth.
Sciurus tricolor, Péppig.
Mouth of Rio Coca, Upper Rio Negro. Collected and
presented by Mr. W. Goodfellow.
The type locality of Péppig’s species was Maynas, in the
angle between the Lower Huallaga and the Marajion; but as
Mr. Goodfellow obtained two species on the Coca to the
northwards, and a third had been found by Signor Balzan to
the southwards, I was in great difficulty as to which was the
real S. tricolor. Happily Dr. Kiikenthal’s kindness has
1388 On Giant Squirrels from the Amazonian Region.
enabled me to fix S. tricolor as one of the two species found
by Mr. Goodfellow on the Coca. Its differences from its
ally are enumerated below. The type is an old female, and
though it was said by Prof. Gravenhorst * to have only
6 mamme, careful search shows that it has 8, the usual
number in this group.
The skull of S. tricolor is less bowed than in the other
species and has an unusually long narrow muzzle.
The melanoid S. fumigatus, Gray, from the ‘ Upper
Amazon,” may possibly be referable to S. trécolor, but its
determination must always remain somewhat doubtful. The
same author’s S. brunneoniger is also a member of the present
group, but is not like any properly localized specimen that I
have seen.
Sciurus cocalis, sp. n.
Not dissimilar at first sight to S. tricolor, found in the same
place, and it is only after the examination of three skins of
one form and seven of the other that I have satisfied myself
of their essential difference. General colour above grizzled
yellowish anteriorly, becoming chestnut-rufous posteriorly.
Crown finely grizzled rufous, not blackish. Backs of ears
rich rufous, a prominent spot behind their posterior bases
bright fulvous. Under surface clear buffy or orange- buffy,
becoming more rufous on the throat and inner sides of the
limbs; line of demarcation on sides sharply defined and in
most specimens with a distinct black line running along the
lower edge of the upper colour, and an indication of a rufous
line along the pale colour just below it. Fore limbs bright
orange-rufous on their outer surfaces from the elbows down-
wards. Hind limbs chestnut-rufous externally as far down
as the ankles, then the feet are again orange-rufous like the
fore limbs. ‘Tail long, bushy, black for its basal 3 inches,
then broadly washed with bright orange-rufous, and so far
like that of S. tricolor, but below the black which succeeds the
rufous on the hairs there is a broad band of yellowish or
orange, which may either extend downwards to the bases of
the hairs or be succeeded again by another dull ring.
Skull shorter, more bowed, and with a more normal-shaped
muzzle than in S. tricolor.
Dimensions of the type (an adult male, measured in
skin) :—
Head and body 315 millim.; tail 283, with terminal
hairs 345; hind foot (wet), s. u. 63, c. u. 663 ear (wet) 32.
* Tschudi, Faun. Peruana, Mami. p. 157 (1845),
I (
Dr. C. A. Kofoid on Pleodorina illinoisensis. 139
Skull: basilar length (of a second specimen) 47°5; nasals
(of type) 18:3* x85; interorbital breadth 20°5; inter-
temporal breadth 19-2; diastema17°8; length of upper molar
series 10-4.
Hab. Mouth of Coca River, Upper Rio Napo.
Type. Male. B.M. no. 0. 6. 3.4. Collected in June 1899
and presented by Mr. Walter Goodfellow.
It is probable that just as S. tricolor ranges northwards
from Maynas to the Coca River, so S. cocalis will be found to
range up that river westwards and northwards, and that the
two species only overlap just at this point. As the two
species are found together it may be well to emphasize their
differences by repeating that S. cocalis, as represented by
seven specimens, differs from 8. tricolor (1) by its grizzled
instead of blackish crown, (2) by the light patches behind the
ears, (3) by its bright orange instead of chestnut fore limbs
and hind feet, (4) its black-lined sides, (5) its basally ringed
caudal hairs, and (6) its shorter muzzled skull.
Sciurus pyrrhonotus, Wagn.
Yungas and Missiones, on the Upper Mamoré, Bolivia
(LZ. Balzan, in Museo Civico, Genoa).
The type locality of S. pyrrhonotus is Borba, near the mouth
of the Madeira, so that its occurrence on the Mamoré, on the
far upper waters of that stream, affords another instance of
the manner in which the ranges of these squirrels follow the
lines of the great Amazonian rivers.
XV.— Plankton Studies.—I1. On Pleodorina illincisensis, a
new Species from the Plankton of the Illinois River. By
C. A. Koroip, Ph.D.T
[Plates V. & VL]
THE genus Plevdorina was discovered in 1893 by Shaw
(1894) at Palo Alto, California, and in May of the following
year the species Pleodorina californica, upon which the genus
was founded, was detected by Mottier (1894) in water from a
shallow stagnant pool near Bloomington, Indiana. During
* Diagonally.
+ From the ‘ Bulletin of the Illinois State Laboratory of Natural His-
tory,’ vol. v. pp. 273-298. From a separate impression communicated by
the Author.
110 Dr. C. A. Kofoid on Pleodorina illinoisensis.
the same summer the form also occurred in the Illinois River
and its adjacent waters (Clinton, 1894), and it has been found
in the plankton of these situations in succeeding years from
June to September. ‘The distribution of the species in this
continent is thus quite extended, and it is not at all im-
probable that continued investigation of freshwater plankton
will demonstrate that this genus has a cosmopolitan dis-
tribution similar to that of some other genera of the family
Volvocinez to which it belongs.
On June 16, 1898, a form which may be referred to the
genus Pleodurina was found in the Illinois River in water
entering the stream in large part from Cook’s Slough and
Quiver Lake. Owing to high water (10 feet above low-water
mark) prevailing at the time, a considerable portion of the
habitat of the form in question consisted of submerged terri-
tory, with shallow warm water abounding in growing aquatic
and semi-aquatic vegetation.
This Pleodorina could not be found in Quiver Lake collec-
tions made on the 7th of June, but on the 16th it was present
in the river in small numbers, increasing until the 20th, when
a maximum was reached. From this time the numbers
decreased until the 27th, when, following a rise in the river,
the species seemingly disappeared entirely from the plankton.
It was also found sparingly in Thompson’s Lake during this
period, a large area of slightly submerged territory being at
this time tributary to the lake.
Associated with this species in great abundance was Hudo-
rina elegans in all stages of asexual reproduction, and Pando-
rina morum was also present in smaller numbers and in like
condition. Volvox, Huglena, Phacus, Lepocinelis, Trachelo-
monas, Dinobryon, Synura, Mallomonas, Uroglena, Melosira,
and Lragillaria occurred in varying frequency, but only a
single specimen of Pleodorina californica was found in collec-
tions containing the species described in this paper. The
animal plankton was represented in the main by rotifers, Poly-
arthra being most abundant, while Syncheta, Euchlanis,
Pterodina, Brachionus, and Anurea were also present.
Diffiugia, Codonella, Bosmina, Cyclops, and nauplii complete
the list of the more common associates of this P/leodorina in
the plankton.
Pleodorina illinoisensis, sp. n.
The species here described consists of an ellipsoidal cceno-
bium or colony of 82, rarely 16 and still more rarely 64,
Dr. C. A. Kofoid on Pleodorina illinoisensis. 141
biflagellate cells. The shape is quite constant, occurring in
the youngest colony and continuing throughout the asexual
cycle until the daughter colonies abandon the gelatinous
matrix of the maternal organism. Among the large number
examined only a few specimens were seen which approached
a spherical form. Measurements of twelve seemingly full-
grown colonies from material freshly killed in 2 per cent.
formalin showed a range of 101 to 137 yw in long diameter and
an average of 113 w. ‘The transverse diameter ranged from
84 to 102 w, and averaged 94 w. Individuals in which the
gonidia have begun to divide show a considerable swelling of
the hyaline gelatinous envelope. One specimen containing
2- and 4-cell stages measured 178x155 pw, and when the
young colonies are ready to escape the parent may measure as
much as 200X175 pw. At the time of escape the young
colonies measure 46X38 yw. The measurements of the
colonies approach very closely those given by Biitschli (1880-
1889, p. 840) for Hudorina, viz. 100-150 yw; and the colonies
of this genus found in association with the form here described
exhibit dimensions almost, if not quite, identical with those
above recorded for the Pleodorina.
The colony (PI. V. fig. 1) contains, as a rule, 32 cells,
arranged, as Henfrey (1856) first noted for Hudorina, in five
circles, two of which are polar and contain four cells each,
while eight cells are found in each of the remaining three
circles, one of which is equatorial, and the other two lie be-
tween the latter and the polar circles. The cells resemble
those of Hudorina in that they are situated in the periphery
of the hyaline gelatinous matrix and are not closely crowded
together, the degree of separation depending upon the age of
the colony, and varying considerably in different cases. Their
inner ends do not approach the centre of the colony, as is the
case in Pandorina. No trace of any protoplasmic connexion
between the cells of a colony could be detected in the living
organisms, nor in material killed in formalin or in chromo-
acetic acid and afterwards stained in fuchsin, hematoxylin, or
Bismark brown. Specimens treated by Zograt’s method
(1 per cent. osmic acid followed by 4 per cent. crude pyro-
ligneous acid), or by 1 per cent. osmic acid followed by picro-
carmine, showed no connexion between the cells.
The colony is surrounded by a common gelatinous sheath
(sh.) increasing in thickness (3°5 to 12 w) with the age of the
organism. ‘lhis membrane or sheath is of equal thickness in
all regions, and consists of two parts—an outer, thin, denser,
more highly refractive layer (0./.), and an inner homogeneous
142 Dr. C. A. Kofoid on Pleodorina illinoisensis.
one (7./.), which shows no traces of the concentric structure
found in Pandorina. It is within this latter layer that the
increase in thickness takes place in the older colonies. It is
limited centrally by a thinner and less highly refractive
layer (m.m.), which encloses the common matrix (m.) in which
the cells of the colony lie. Frequently among the older
organisms there occur upon the posterior end of the colonies
blunt pseudopodia-like protuberances (Pl. V. fig. 4) of the
sheath, of irregular form and of no constant number. ‘Their
position and the fact that they are often, though not always,
found in old colonies from which some of the daughter colonies
have already apparently escaped, suggest that they may mark
the place of exit of the young individuals from the parent.
Similar protuberances were observed upon Hudorina and
Pandorina, under similar conditions, in the collections in which
the Pleodorina under discussion was found. Wills (1880)
found that the daughter colonies of Volvox globator escaped
through a rift in the posterior hemisphere of the parent, and
Klein (1889) observed the same phenomenon in Volvox aureus.
The escape of the daughter colonies in Pleodorina has not
been observed by me. .
The sheath stains deeply in an aqueous solution of methylen
blue, more deeply, in fact, than the enclosed matrix, the outer
layer taking the deeper stain. It also-shrinks to about one
fourth its former thickness. ‘This shrinkage, together with
that of the central matrix, causes the sheath to wrinkle along
lines which bound hexagonal areas from whose centres the
cells now project, thus giving the appearance of a division of
the surface of the colony into regular polygons. The sheath
shows no trace of the layer of radial rod-like structures
found by Klebs (1886) in Pandorina, but iodine or methylen
blue demonstrates a finely granular condition like that
described for Hudorina. 'The sheath is traversed by the pairs
of flagella which arise from the outer ends of each of the cells.
The matrix (m.) is a gelatinous substance of some con-
sistency, filling the colony inside of the inner membrane, In
the living colonies, in those which were killed in the various
reagents mentioned above and afterwards stained, and in
disintegrating material, no traces of any divisions can be
detected in this substance that are not due to wrinkling caused
by shrinkage. Methylen blue or iodine causes the matrix to
show a faintly reticulated or vacuolated appearance due to
different densities of staining. ‘That the substance of the
matrix has considerable consistency, even in the swollen con-
dition found in the maternal colonies, is shown by the fact
that the flagella of the young forms, before rotation begins, \
Dr. C. A. Kofoid on Pleodorina illinoisensis. 143
can be seen to penetrate the matrix of the parent very slowly.
Their ends are often blunted, or even knob-like, and their
lateral motion is very limited. The movement of the young
colonies through the matrix is a very slow and gradual one,
showing the gelatinous consistency of the substance in which
they are imprisoned.
The striking feature in the structure of this species, as in
the case of P. californica (Shaw, 1894), is the presence of
two distinct types of cells in the colony (Pl. V. fig. 1), the
vegetative (v.c.) and the gonidial (g.c.) cells. The presence
of these two types of cells at once places this new species in
the genus Pleodorina rather than in Hudorina, which it other-
wise closely resembles.
The vegetative cells (v.c.) are four in number and consti-
tute the anterior polar circle, being always directed forward
in locomotion, as in the other species of the genus. Their
number remains the same in the smaller colonies of sixteen
cells and in the larger ones of sixty-four. ‘The diameter of
these cells ranges trom 9-5 to 16°8 w, twelve cells averaging
12°25 w. The size of these cells varies even in the matured
colonies, measurements at this stage ranging from 9°6 to
15°6 w. At birth the cells of the young colonies vary in
diameter from 3°5 to 5 w indifferent parents. In the daughter
colonies, while still in the maternal matrix, no distinction in
size between the vegetative and gonidial cells can be detected,
nor can this distinction be made in the younger free-swimming
colonies, it being thus impossible at this stage to distinguish
the young P.zldnotsensis from the similar stages of Eudorina
elegans with which they were associated. When the young
colonies have attained dimensions of 46 x 38 wu the vegetative
cells measure 4 w and the gonidia 4°8 uw. A like similarity
between the two kinds of cells in the young colonies exists,
according to Shaw (1894), in P. californica.
In structure the vegetative cells (Pl. V. fig. 2) are in most
particulars similar to the gonidia, described below. They
sometimes appear to be a trifle lighter green in colour—a
difference which may be due to their smaller size. The
principal differences lie in the smaller number of pyrenoids
and the larger size, both absolute and relative, of the stigma
or eye-spot.
As to the fate of the vegetative cells, the evidence at hand
is insufficient and conflicting. In three colonies in which the
daughters were moving about in the maternal matrix, some
having already escaped, the vegetative cells showed very
evident signs of degeneration, the contents being shrunken
and irregular. In the larger number of instances of this stage
144 Dr. GC. A. Kofoid on Pleodorina illinoisensis.
under observation the cells appeared normal, showing no trace
of degeneration or division. In one instance only have I
found a specimen in which the vegetative cells had divided
beyond question. This was a colony in which the gonidia
had completed their division but had not escaped. Three of
the vegetative cells were in the two-cell stage and one was
undivided. In two instances matured colonies have been
found in which four smaller daughter colonies (of eight and
sixteen cells respectively) were present at one pole.
The gonidial cells (Pl. V. fig. 3) constitute the remainder
of the colony. They usually number 28, rarely 12 or 60,
and occupy the parts behind the anterior polar circle of vege-
tative cells. These cells in most instances can be easily
distinguished by their larger size. In form they are spherical,
though some specimens in preserved material are slightly
flattened on their inner ends. In diameter they range in
seemingly full-grown colonies from 15 to 25 w, averaging in
twelve specimens 19'2 w. Their dimensions just before their
division, that is, in colonies in which division has begun, also
show the extreme range quoted above, the smaller diameter
having been found in a sixteen-cell colony. As a rule the
gonidia are all of the same size, but occasionally specimens
have been found in which one er more dwarf cells occur
among them. ‘These are irregular in their distribution and
can be distinguished at once from the vegetative cells by
their position. Similar dwarf cells were found in both Pan-
dorina and Eudorina. In matured colonies gonidial cells are
frequently found which fail to divide. The gonidia are of a
light green colour, a trifle darker than the vegetative,¢ells.
Their colour in general is similar to that in Hudorina, and is
somewhat lighter than that in Volvow and Pandorina, with
which they are associated.
A distinct cell-membrane (¢.m.) is found about each of the
cells. In the living condition and in the material preserved
in formalin it forms a highly refractive hyaline layer, about
1 » in thickness, outside of the green contents of the cell. It
stains very faintly in hematoxylin and assumes a deep brown
tint with long-continued action of iodine and sulphuric acid.
In the case of diseased colonies hereafter mentioned the cell-
membranes persist, often retaining their original form and
shape, after the entire disappearance of the contents.
‘he greater part of the cell-contents consists of what seems
to be one large chromatophore (Pl. V. fig. 2, chr.), which
occupies all of the cell except the centrally placed nucleus
with its enveloping protoplasm, and a slender column (p.c.)
passing from this region to the anterior end of the cell. In
Dr. C. A. Kofoid on Pleodorina illinoisensis. 145
many cells a faintly marked notch or furrow (/u., fig. 1) is to
be detected on one side of the chromatophore at the anterior
end of the cell. ‘This seems to mark the line of contact of
the sides of the chromatophore which has surrounded the
nucleus. In the 2- and 4-cell stages of the gonidial cells
the nucleus and the protoplasmic mass are plainly seen
to occupy one side of the cell (Pl. VI. figs. 7, 8), but in
the cells of the young colony it again occupies a central
position. The chromatophore is uniformly of a bright chloro-
phyll-green, and shows a finely granular structure under high
magnification. In the youngest colonies each cell contains
but a single spherical pyrenoid (pr.), which occupies a lateral
position in the chromatophore, in the inner hemisphere of the
cell. Inthe older colonies the number of pyrenoids increases,
as many as twelve having been found. ‘They are scattered
irregularly through the chromatophore, and may occur in any
art of it. A similar increase of pyrenoids is reported by
Shaw (1894) for P. californica, In the vegetative cells the
number of pyrenoids is often but 2-4, and is, as a rule, less
than that of the gonidial cells. In a very few instances as
many as eight have been found, and in one ‘old colony the
vegetative cells seemed to be packed full of pyrenoids. In the
young colonies the pyrenoids have a diameter of about 1p,
and in the older colonies of 2°5 w,
The nucleus (n.) lies in about the centre of the cell in the
midst of a mass of protoplasm enclosed by the chromatophore.
In mature gonidial cells before division it has a diameter of
7-8 w, and contains a sub-central nucleolus (ne/.) whose
diameter is 3m. ‘The nucleolus stains deeply with picro-
carmine, and is by this means easily distinguished from the
pyrenoids, which it resembles in appearance and size. The
nuclear membrane is detected with difficulty. It encloses a
faintly stained nuclear reticulum (7.). In the younger cells
the nucleus is much smaller (4-5 ~), the nucleolus is rela-
tively larger, and the reticulum is not evident. In the living
cell the nucleolus alone can be seen in the midst of the
greyish protoplasmic mass at the centre of the cell. The
rotoplasm is continued from this central region peripherally,
in the axis of the cell as a slender column (p.c.), to the
anterior end, where it includes the stigma and bases of the
two flagella. A protoplasmic mantle enclosing the chromato-
phore was not demonstrated.
The stigma or eye-spot (s.) lies at the anterior end of the
cell, near its axis, and is often so placed that an equilateral
triangle may be drawn with it and the bases of the two flagella
as apices. It is of a bright reddish-brown colour, though in
Ann. & Mag. N. List, Ser, 7.’ Vol. vi. 10
146 Dr. G. A. Kofoid on Pleodorina illinoisensis.
some of the posterior cells the colour is often very faint, giving
the stigma the appearance of a slightly tinged oil-globule.
It is of an elongated hemispherical shape when seen from the
side, and has a circular outline when seen from above. Its
upper end often projects slightly so as to elevate the cell-
membrane. The application of killing agents and alcohol
soon removes its colour, and even in formalin this fades out in
the course of a few days, leaving merely a colourless, highly
refractive structure. ‘The larger stigmata have a diameter of
2°5 w and a depth of 2°8 w, and are to be found in the cells
in the anterior part of the colony, especially in the four vege-
tative cells of the anterior polar circle. Posteriorly the stig-
mata are less prominent, and are often not to be found at all
as brightly coloured spots but merely as pale globules whose
position alone affords a clue to their real character.
This specialization of the stigmata in the anterior end of
the colony occurs also in Hudorina, Pandorina, and Volvoz,
and Shaw (1894) states that in P. californica the stigmata,
which are present in the posterior part of the young colonies
(in gonidial cells), become less conspicuous and disappear as
the colony enlarges and the differentiation of the cells pro-
ceeds. ‘This prominence of the stigmata in the anterior end,
together with the facts that this end is always directed fore-
most in locomotion and that the species showing this differ-
entiation are positively phototactic in the vegetative condition
when the differentiation is prominent, all point toward the
participation of the stigmata in the function of light percep-
tion. An interesting phenomenon occurs at the time of the
division of the gonidia, for the stigma of the mother cell
persists and is passed on through the five successive cell-
divisions to the outer end of one of the cells of the daughter
colony, situated in the margin of the cup which arises from
the plate of cells and closes to form the ellipsoidal daughter
colony. Inasmuch as this cup always closes from the inside
out, that is, with the opening directed outward, it is evident
that the stigma must traverse the distance between the outer
end of the mother cell and its inner end, which corresponds
to the outer ends of the cells of the daughter colony. New
stigmata arise in the cells of the daughter colony, but being at
first very small are thus quickly distinguishable from the
persisting stigma. ‘I'he ultimate fate of this persisting stigma
has not been traced. .
No contractile vacuole was observed in the living cells, and
careful search with a Zeiss j4-inch oil-immersion lens for
this structure in preserved and stained material has led to no
positive identification of a vacuole, ‘The bleached stigma
Dr. C. A. Kofoid on Pleodorina illinoisensis. 147
and what seem to be the enlarged bases of the flagella are the
only areas discernible in the anterior end of the cell which at
all resemble a contractile vacuole. Shaw (189+) finds in
picronigrosin material a single vacuole in the anterior end of
the young cells of P. californica.
The flagella ({) are two in number for each cell, and unite
with the cell at the anterior end adjacent to the stigma. The
two flagella have the same proportions, and in adult colonies
they measure 40 in length. In the young colonies they
are relatively longer. They are visible on the young colonies
shortly after the cup closes, and persist upon the maternal
colony during the early divisions of the gonidia.
The locomotion of the colonies of Pleodorina illinotsensis
is of the type prevailing among other spherical or ellip-
soidal genera of the Volvocinee; viz., rotation about the
principal or long axis of the colony, either from right
over to left or the reverse, frequently with one direction
predominating, progression being usually along the line
of the axis, the same end of the colony always leading.
In P. illinoisensis the vegetative pole always leads in loco-
motion in horizontal, oblique, and vertical movements, and
is therefore the anterior pole of the colony. Under normal
conditions, when under observation, this species ts rarely quiet
during the period of growth. While still within the matrix of
the mother colony the ceaseless rotation with its frequent rever-
sals begins. Colonies in the life cell, while favourable con-
ditions prevail, can be seen in active movement, jostling one
another and their neighbours in their seemingly aimless wan-
derings. When an object is met which does not yield to their
persistent rotation, their movements may slacken for a time
to be resumed shortly in some line of less resistance. The
rotation of this species is prevailingly from right over to left,
as the following tables show, which indicate the number and
direction of the reversals of rotation in ten individuals in one
minute.
Pirection: ..< es 6. < i AO Gee ele S| 9» | 10: |\Tatal,
Right over to left...) 2 2 1 2 1 ON |e /4: Bgl Oe
Left over toright..| 2 Bee eee aren reek ieee eine EEG
A few days later a second set of observations was made
with the following result -—
10*
148 Dr. C. A. Kofoid on Pleodorina illinoisensis.
Direction ........ 1 | 2) Billa a | Gy A Sao 10 ‘Total.
Right over toleft ..|) 3-4 | 5+ | 3+
Left over to right... 2— 5+) 2+
In the majority of instances where the direction of rotation
was observed it was from right over to left, the ratios being
22 to 16, and 35 to 80. These tables give some idea of the
frequency of change in direction and its variation in different
individuals, but do not show the duration of the directions of
rotations. This is indicated in a general way in the second
table by the plus and minus signs, which show the direction
in which the rotation was of longer and shorter duration.
Tn conclusion it may be said that both directions of rotation
occur, though that from right over to left is more frequently
met with, or, in other words, is of longer duration.
With regard to locomotion in P. californica, Shaw (1894)
says that “the movement of the plant in the water was fol-
lowed in the case of a few individuals bearing well-developed
gonidia. In swimming through the water the vegetative pole
is directed forward and the plant revolves to the right (in
observed cases) on the axis connecting the vegetative and
reproductive poles. The path is parallel to this axis in up-
ward vertical as well as in horizontal movement.” The
polarity of this genus thus expressed physiologically in the
movements of the colony is accompanied by a corresponding
structural differentiation of the cells composing the organism,
In Gonium, according to Fresenius (1856), the motion of
the colony resembles that of a wheel, progression taking place
in the line of the axis of rotation. According to Biitschli
(1883-1887, p. 858), locomotion is accomplished by the
rotation of the plate-like colony around its shorter axis, the
direction of rotation being to the right in some individuals
and to the left in others. Pfeffer (1884), on the other hand,
describes the rotation during the forward movement as alter-
nately from the right and the left. Migula (1890) calls
attention to the wavering, often backward, and irregular
movements of this genus, and also notes its rotation about
an axis through the middle of the colony. ‘This rotation is
either to the right or to the left, no predominance being men-
tioned. Polarity is thus marked in the activity of the Gontwm
colony, though not expressly marked in its structure except
as it appertains to the individual cells.
Dr. C. A. Kofoid on Pleodorina illinoisensis. 149
In Stephanosphera the polarity in structure is but slightly
marked in the colony, being indicated in some colonies by
the asymmetrical position of the cells, but there is a physio-
logical differentiation in that one pole of the colony leads in
locomotion. In this genus also, according to Cohn (1852),
‘the rotation is in either of the two directions and is subject
to frequent change. No predominant direction was noted
by him.
In Pandorina the only structural expression of polarity is
found in the greater development of the stigmata in the cells
in the anterior end of the colony. In other particulars the
poles are not differentiated. Braun (1851) maintains that in
this genus the rotation is constantly around the long axis of
the colony in the direction of the hands of a clock, when the
motion is toward the observer. Niigeli (fide Biitschli, 1883—
1887, p. 858), on the other hand, observed rotation in both
directions. My own observations upon Pandorina morum
show beyond question that the direction of rotation is not
constant, as the following table demonstrates :—
Piraction) *.-0....<% Pelee en Sees We Ge tar eo hon h kO! Metal,
Right over to left ..)6+ |12-+) 54+ |1—|24+ |8+ |74)14/|1+/84| 41
Left over to right . .| 6— |12—| 4+ | 24+ )2+)4+/64+)2—|1-|38+)]| 42
We
The table gives the direction and number of changes in
direction in rotation of ten colonies, each observed for one
minute. ‘The plus and minus signs indicate the estimated
predominance in duration, According to the table the in-
stances of direction observed are approximately equal for the
two directions, though that from right over to left showed
the greater duration, The younger and smaller colonies
showed much the greatest activity and exhibited more
frequent changes in direction than the older colonies. In all
observed cases the same end continues to lead in locomotion,
physiological polarity being thus fully developed in this genus.
In the case of Hudorina the structural polarity of the
vegetative colonies is no more marked than it isin Pandorina,
though according to Carter (1858) there is in the moncecious
sexual colony a differentiation, in that the four cells at one
pole divide to form spermatozoa, while the remaining twenty-
eight become egg-cells. It should be noted in this connection
that no such colonies were observed by Goroschankin (1875)
in the sexual generation, The literature at hand presents no
150 Dr. G. A. Kofoid on Pleodorina illinoisensis.
precise statement as to locomotion in this genus. As ob-
served by me, it closely resembles that described above for
Pleodorina illinoisensis ; viz., rotation around the long axis
of the colony, the same pole constantly leading in progression.
The direction of rotation is frequently reversed, though it was
predominantly from right over to left in the cases observed.
A functional polarity thus exists in this genus.
In Volvow, according to Klein (1890), there is a polar differ-
entiation as regards the stigmata that is even more marked
than it is in the genera previously mentioned. He finds that
the cells of the pole directed forward in locomotion each
possess a stigma which is especially large and intensely
coloured; that the colour fades out and the stigmata become
smaller and paler as the equator is approached; and that
beyond this they are usually represented merely by a colour-
less oil-drop, which in some cases may even disappear. ‘The
posterior hemisphere is also marked by the development
there of the gonidia, as was first shown by Cohn (1856), and
occasionally ellipsoidal colonies are found whose long axis
connects the anterior and posterior poles. Locomotion in
Volvoz is accomplished, as elsewhere in the family, by the
rotation of the colony about its principal axis. Wills (1880)
observed the predominance of the rotation to the right and
its occasional brief reversal. Klein (1889) states that this
preference is found in V. globator, but that it is not shown by
V. aureus. In this latter species the changes are frequent and
are often separated by a brief pause. Backward motion is
rarely seen and lasts but a short time. In the case of Volvow
the axis of rotation is slightly oblique, the centre of the
colony remaining in the line of progress, but the axis of
rotation being inclined from above the line at the anterior
pole to below it at the posterior one.
We thus find that Pleodorina dllinoisensis, which exhibits
both a structural and physiological polarity, shares with
most, if not all, of the genera of the family to which it
belongs the physiological differentiation which is expressed
in locomotion, and also, in observed cases, exemplifies the
extreme form of a predominance of rotation in one direction.
We also find that the structural differentiation shown in the
decadence of its posterior stigma obtains in varying degrees
in the other spherical and ellipsoidal genera of the family—
least in Pandorina, most in Volvoew. ‘The genus Pleodorina
agrees with Volvox in having astructural polarity based upon
the division of the colony into vegetative and gonidial regions,
but the differentiation is simpler. Of the two species of Pleo-
dorina, the one here described exhibits the simplest possible
Dr. C. A. Kofoid on Pleodorina illinoisensis. 151
differentiation of the colony consistent with the symmetry
of the organism; viz., the differentiation of the anterior
polar circle of four cells as vegetative members of the colony.
Of the two species of the genus it thus stands nearer Hudo-
rina, while its sister species, P. californica, approaches more
closely to Volvow both in the number of cells and in the
extent of the differentiation.
The discovery of this additional species of the genus Pleo-
durina thus supports the opinion expressed by Shaw (1894),
who founded the genus, that it was intermediate between
Eudorina and Volvow but nearer the former. Judging merely
from the asexual stage, P. illénoisensis affords additional evi-
dence of the close relationship of Pleodorina and Hudorina.
Throughout the preparation of this paper the writer has
had constantly in mind the possibility that the form here
described is merely a stage in the life cycle of Eudorina. A
number of facts lend support to this hypothesis: (1) the
occurrence of Pleodorina illinoisensis with Hudorina elegans ;
(2) their marked similarity, aside from the four vegetative
cells, in structure and measurements; (8) the impossibility
of separating the youngest free-swimming colonies of the
two forms; (4) a considerable variation in the size of the
vegetative cells in Pleodorina, grading toward the condition
in Hudorina; (5) some evidence that in certain cases at
least the vegetative cells may divide, one case of a 2-cell
stage having been seen in the hundreds, if not thousands,
of specimens examined, and one instance noted in which
a maternal colony containing thirty-two daughter colonies
had at one pole four colonies which were slightly smaller
than the remaining twenty-eight; and (6) the occurrence
of pleomorphism in the family Volvocinex, Klein (1889 and
1890) citing no less than twenty-four “ combinations ”’ in the
case of Volvow aureus. It may then be that the form here
described as Pleodorina illinoisensis is only a ‘ Pleodorina
stage” of Hudorina.
The abrupt disappearance of this supposed new species
from the plankton prevented the carrying out of breeding
experiments designed to test its validity, and it seems that
the matter must remain undecided for the present. In the
absence of satisfactory proof that the form here described is
but a phase of the life cycle of Hudorina it has seemed best
to the writer to make the above suggestion and to take the only
course open in publication, namely, the description of the form
as a new species, inviting the criticism of subsequent investiga-
tion. The dilemma here presented is by no means an isolated
one in plankton work, nor is it new to the family Volvocinea :
152 Dr. C. A. Kofoid on Pleodorina illinoisensis.
witness the long confusion which existed over the two species
of Volvoxv, aureus and globator, which has been at last cleared
up by the excellent work of Klein (1889, 1889 a, and 1890)
and Overton (1889). Another instance is often presented
when Pandorina and Eudorina both occur in the same
collections and the plankton statistician must decide to which
genus each specimen observed must be referred. ‘Typical
specimens of each can be found, but all individuals do not
conform to the type, or they may present conditions in
which the conformation is obscured by some phase of the
life cycle.
The asexual reproduction of Pleodorina illinotsensis (P1. VI.)
resembles that of other species of the genus in that it is
accomplished by the repeated division of the gonidial
cells, resulting in the formation of daughter colonies in the
maternal matrix. ‘These escape later from the parent organ-
ism, and by growth attain the adult condition with the differ-
entiation of the four vegetative cells. Five successive cell
divisions, pervading all the cells of the parent organism
except the vegetative cells, are necessary for the completion
of the process, and result in the 2-, 4-, 8-, 16-, and 32-cell
stages of the forming colonies. The first two of these divi-
sions result in the formation of a quadrangular plate of cells
—a form which is retained through the two succeeding
divisions, which produce the 8- and 16-cell stages. The
cupping of this plate, which results in the formation of an
ellipsoidal colony, is apparent as early as the 4-cell stage
(Pl. VI. fig. 9) and continues through the later stages
(figs. 11, 13), so that by the time the 16-cell plate is formed
it has almost the curvature of a saucer. With the formation
of thirty-two cells the closure of the cup proceeds and is soon
completed. The orifice of the cup is directed outward in all
cases, and thus the ends of the cells of the daughter colony
which are formed from the outer end of the maternal gonidial
cell come to lie in the inner side of the cup, and are the inner
ends of the cells of the daughter colony. In the matured
colonies the young usually lie with their long axes parallel
to the surface of the parent. I have not, however, been able
to identify the point of closure of this cup with this region
or positively with any other.
‘Lhe sequence and position of cleavage planes which produce
the quadrangular plate of the 16-cell stage is, in the main,
similar to that described by Goroschankin (1875) and Braun
(1875) for Hudorina and Volvoz. Beyond this stage there is
some doubt as to the agreement. A full discussion of the
subject is beyond the scope of the present paper, for which the
Dr. GC. A. Kofoid on Pleodorina illinoisensis. 153
following brief description must suffice. The first cleavage
plane (I) divides the gonidial cell into two hemispheres along
the axis of the cell, and the daughter nuclei, with the sur-
rounding protoplasm, are placed close together in the centre
of the opposing faces of the new cells (Pl. VI. fig. 7).
The second plane (II) is at right angles (figs. 8, 9) to the first
and aiso passes through the regions representing the axis
of the ancestral cell. In this instance also the nuclei are
gathered near the centre of the young colony, which exhibits
to an appreciable extent the curving indicative of the later
formation of the cup. The 8-cell stage results from the
divisions of each of the quadrants of the 4-cell stage by a
plane (III) which is parallel to one of the previous planes
and perpendicular to the other, meeting the latter at a point
about midway between the centre and the circumference. By
a subsequent adjustment of the cells the four more centrally
placed ones come to form a sort of Greek cross whose angles
are filled by the other four (Pl. VI. figs. 10,11). The 16-
cell stage is formed by four additional planes, each of which
divides one of the cross-cells and its corner neighbour. The
location of these planes may be described in the same terms
as the last, excepting that they meet the radial planes, I and
II, at about one fourth the distance from the circumference
toward the centre. The cupping of the plate soon advances
to such an extent that it consists of a square of four centrally
placed cells, upon each of the four sides of which there over-
hangs a row of three cells, of three grades of elevation (PI. VI.
figs. 12, 13). ‘The succeeding division and the completion of
the cup (PI. VI. fig. 14) result in the young colony’s
assuming the ancestral form. Throughout these divisions the
number of pyrenoids in the daughter cells grows steadily less.
But one can be found in each cell in the 32-cell stage, while
in the 16-cell stage two are readily recognizable in each cell.
In the earlier stages and before division the number often
varies, and the pyrenoids are frequently so crowded that
enumeration is difficult, if not impossible. It seems not im-
probable that these structures also must undergo some division
during the process of cell-multiplication. During the pro-
cesses of division the nuclei continue to occupy a position
near the inner ends of the cells (in the new colony), and it is
only after the divisions have been completed that they come
to occupy their usual positions at the centre of the cells—
perhaps as a result of the growth of the chromatophore.
No stage of sexual reproduction has been positively identi-
fied for this species.
A peculiar condition of the colonies of this species, also
154 Dr. GC. A. Kofoid on Pleodorina illinoisensis.
occurring in Pandorina and Eudorina, deserves passing
notice. It occurred with considerable frequency in all three
genera, and resulted in each case in the destruction of the
entire colony affected. The early stages of the disease, if it
be such, are indicated by the homogeneous condition of the
cells and the fading out of the colour, together with a flat-
tening of the cell-contents into a disk- or lozenge-shaped
mass (Pl. V. fig. 4). In the subsequent stages this mass
assumes a yellow and then a brownish colour, takes on an
irregular shape (Pl. V. fig. 5), and disintegrates, leaving the
empty cell-walls occupying the matrix. In spite of the
suggestion in the above description there was never any trace
of the formation of spermatozoa in the colonies presenting
these phenomena, neither was there any indication of encyst-
ment. There was no indication of either a fungous or an
algal parasite, and it seems not improbable that the occur-
rence of these diseased forms may have been due to some
unfavourable local condition in the water tributary to the
habitat of the genera affected.
The following brief synopsis of the prominent characters of
this genus and its two species will serve as a convenient
diagnosis for their determination :—
PLEODORINA, Shaw.
Colony consists of a spherical or elliptical coenobium of
greenish biflagellate cells of two types, vegetative and
gonidial, in the anterior and posterior parts of the colony
respectively, which lie in the periphery of a hyaline gelatinous
matrix and are surrounded by a common hyaline envelope.
Cells each with one reddish stigma, which is more prominent
in the anterior part of the colony. No connecting filaments
between the cells. Non-sexual reproduction by gonidia,
which are formed by increase in size of a part of the cells of
the colony. Daughters escape from parent as small colonies
of biflagellate cells, which at this stage are all similar,
Sexual reproduction not known.
Pleodorina californica, Shaw.
Number of cells in colony 64 or 128. Maximum diameter
of colony 175-340 . Vegetative cells constituting approxi-
mately one half the colony. Gonidial cells 2-3 times diameter
of vegetative cells. Known habitat: ponds, ditches, and
streams in California, Indiana, and Illinois,
Dr. C. A. Kofoid on Pleodorina illinoisensis. 155
Pleodorina illinoisensis, sp. 0.
Number of cells in colony usually 32, rarely 16 or 65.
Dimensions of colony range from 46x 38 & to 200 x 175 pw.
Vegetative cells always 4in number. Gonidial cells approxi-
mately 1, 1—2 times diameter of vegetative cells. Known
habitat : submerged lands along the Illinois River. Types
deposited in collections of Illinois State Laboratory of Natural
History and United States National Museum.
Illinois Biological Station,
July 25, 1898.
Bibliography.
Braty, A. (1851.)—‘ Betrachtungen iiber die Erscheinung der Ver-
jiingung in der Natur, insbesondere in der Lebens- und Bildungs-
geschichte der Pflanze.’ xvi+864 pp., 38 Taf. Leipzig.
——. (1875.)—“ Ueber einige Volvocineen,” Ber. d. Gesellsch. natur-
forsch. Freunde zu Berlin, 1875, pp. 9-18.
Birscuut, O. (1880-89,)—* Protozoa,” Bronn’s Klassen und Ord-
nungen des Thierreichs, Bd.i. Th. iii. 2035 pp., 79 Taf. Leipzig
und Heidelberg.
Carter, H. J. (1858.)— On Fecundation in Eudorina elegans and
Cryptoglena,” Ann. & Mag. Nat. Hist. ser. 8, vol. 11. pp. 287-2538,
1. vill.
Corin G.P. (1894.)— Pleodorina in Illinois,” Botan. Gazette,
vol. xix. p. 585.
Conn, F. (1852.)—‘‘ Ueber eine neue Gattung aus der Familie der
Volvocineen,” Zeitschr. f. wiss. Zool. Bd. iv. pp. 77-116, Taf. vi.
——. (1856.)—“ Observations sur les Volvocinées,” Ann. d. Sci. Nat.,
Botan. iv. sér. t. v. pp. 323-332; also, with some changes, in
34 Jahresber. d Schles. Gesellsch. f, vaterl. Cultur f. 1856, pp. 39,
40, 77-83.
Fresenius, G. (1856.)—“ Ueber die Algengattungen Pandorina,
Gonium, und Raphidium,” Abhandl. der Senckenberg, naturforsch.
Gesellsch. Bd. ii. pp. 187-200, Taf. viii.
GoROSCHANKIN, J. (1875,)—‘ Genesis im Typus der palmellenartigen
Algen. Versuch einer vergleichenden Morphologie der Familie der
Volvocinea’’ (German title of Russian article), Mittheil. der kais.
Gesellsch. naturforsch. Freunde in Moskau, Bd. xvi. 40 pp., 2 Taf.
Henrrey, A. (1856.)—“Notes on some Freshwater Confervoid
Algve new to Britain,” Trans. Micr. Soc. London, n, s., vol. iv.
pp. 49-54, pl. iv.
Kurss, G. (1886.)—“ Ueber die Organisation der Gallerte bei einigen
Algen und Flagellaten,” Untersuch. :a. d. botan. Inst. zu Tiibingen,
Ba. ii. pp. 853-418, Taf. iii., iv.
Kuxrtn, L. (1889.)—‘‘ Morphologische und biologische Studien tuber
die Gattung Volvox,’ Pringsheim’s Jahrb. f. wiss. Botan. Bd. xx,
. 183-211, Taf. x.—xii.
——. (1889 a.)—“ Neue Beitriige zur Kenntniss der Gattung Volvox,”’
Ber. d. deutsch. botan. Gesellsch., Jahrg. 1889, Bd. vii. pp. 42-53,
Taf. ii.
—. (1890.)—‘ Vergleichende Untersuchungen iiber Morphologie
und Biologie der Fortpflanzung bei der Gattung Volvox,” Ber. d.
naturf. Gesellsch. zu Freiburg i. B., Bd. v. pp. 29-120, Taf. ii.-vi.
156 Dr. GC. A. Kofoid on Pleodorina illinoisensis.
Mieuta, W. (1890.)—“ Beitrige zur Kenntniss des Goniwm pec-
torale,” Botan. Centralbl. Bd. xliv. pp. 72-76, 103-107, 143-146,
Taf. ii.
Mortrer, D. M. (1894.)—“‘ Pleodorina in Indiana,” Botan. Gazette,
vol. xix. p. 383.
Overton, E, (1889.)—“ Beitrag zur Kenntniss der Gattung Volvox,”
Botan. Centralbl. Bd. xxxix. pp. 65-72, 118-118, 145-150, 177-
182, 209-214, 241-246, 273-279, Taf. i.-ii.
PFEFFER, W. (1884.) — “ Locomotorische Richtungsbewegungen
durch chemische Reize,” Untersuch. a. d. botan. Inst. zu Tiibingen,
Bd. i. pp. 363-482.
Suaw, W. R. (1894.)—“ Pleodorina, a new Genus of the Volvocineze,”
Botan. Gazette, vol. xix. pp. 279-283, pl. xxvii.
Wits, A. W. (1880.)—*On the Structure and Life-History of
Volvox globator,’ Midland Naturalist, vol. iii. pp. 209-214, 233-
237, pls. vil., vill.
EXPLANATION OF THE PLATES *.
Abbreviations.
A, Anterior pole. nel, Nucleolus.
chr. Chromatophore. o.l. Outer layer of sheath.
cm. Getlnent rane: P. Posterior pole.
f. Flagellum. p.c. Protoplasmic column.
Ju. Furrow. pr. Pyrenoid.
g.c. Gonidial cell. r. Reticulum.
2.1, Inner layer of sheath. s. Stigma.
m. Matrix. sh, Sheath.
mm. Matrix membrane. v.c. Vegetative cell,
n. Nucleus. I, I, II. Cleavage planes.
PLATE V.
. Pleodorina illinoisensis, lateral view of colony. X 500.
. Lateral view of vegetative cell. x 1500.
. Lateral view of gonidial cell. x 1500.
Lateral view of matured colony, showing posterior lobes. X 185.
. Diseased cell, early stage. XX 1250.
. Diseased cell, later stage. x 1250.
S
> Or CO BD
PLATE VI.+
7. Pleodorina illinoisensis, top view of 2-cell stage.
Fig. 8. Top view of 4-cell stage.
Fig. 9. Lateral view of 4-cell stage.
Fig. 10. Top view of 8-cell stage.
Fig. 11. Lateral view of 8-cell stage.
Fig. 12. Top view of 16-cell stage.
Fig. 13. Lateral view of 16-cell stage.
Fig. 14. Top view of 32-cell stage.
* Figures drawn by C. A. Kofoid and inked by Miss L. M. Hart.
+ Figs. 7-14 magnified 1000 diameters,
Bibliographical Notice. 157
BIBLIOGRAPHICAL NOTICE.
Catalogue of the Lepidoptera Phalene in the British Museum.
Volume II. Catalogue of the Arctiade (Noline, Lithosianse) in the
Collection of the British Museum. By Sir Grores F. Hampson,
Bart. 8vo. Pp.xx,589. Plates xvili-xxxv. London: printed
by order of the Trustees, 1900,
Tue first Catalogue of Moths (Lepidoptera Heterocera as they used
to be called, or Lepidoptera Phaleenz as some recent authors prefer
to call them) was published in thirty-five small volumes, without
plates, between 1854 and 1866, and was edited by the late Francis
Walker. Notwithstanding the numerous.errors which have fairly or
unfairly been charged against this work, its publication lent an
enormous impetus to the study, for it enumerated upwards of 20,000
species, most of which, except in a portion of the Micro-Lepidoptera,
were described, with full synonymy in the case of known species.
But Walker’s Catalogue is now quite out of date, and the earlier
volumes are entirely out of print, and therefore the Trustees of the
British Museum have projected a new Catalogue covering the same
ground, but profusely illustrated with plates and figures (rendered
all the more necessary by the stringency of the official regulations,
which do not admit of specimeus once registered ever being lent out
of the building), and have entrusted the commencement of the work
to Sir George F, Hampson. We say the commencement, for though
Sir George is not an old man, and his energy and rapidity of work
are well known, it will take two or three men’s official lifetimes at
least to complete the work on the same plan, unless several men are
employed to work at different families at the same time.
The two volumes which have already appeared (in 1898 and
1900) include descriptions of 2377 species, and yet the ground
which they cover only corresponds to a comparatively small portion
of the first two volumes of Walker’s Catalogue, with the corre-
sponding supplements. This will be enough to show the enormous
increase in our knowledge of the subject within the last fifty years.
Sir George is working undcr very favourable circumstances, for
the entomological section of the reference library at South Ken-
sington is to all intents and purposes practically complete ; and the
collection of Moths has been so largely increased lately, that it is
now, beyond dispute, one of the very best in the world, if not incon-
testably the best. Moreover, in the earlier famities he has the
advantage of the assistance of Mr. Kirby’s approximately complete
Catalogue of Sphinges and Bombyces, published in 1892, and for
later years the ‘ Zoological Record’ &c.; so that the chances of
anything important being overlocked are reduced to a minimum.
Sir George Hampson’s methods of work are so similar in all his
publications that those who have seen one volume of his work on
the ‘ Moths of India’ or one volume of the present series, may form
a fair judgment of the whole. All the species are briefly described
(the bulk of the book forbids very lengthy descriptions, which are,
moreover, less necessary in the case of recognizably figured species),
158 Miscellaneous.
and the numerous tables of genera and species will be very useful.
We are glad tosee, too, that transformations are described in the case
of the comparatively few species in which they are known. We
may mention that the excellent (and, in the present volume,
crowded) coloured plates are by Mr. Horace Knight.
MISCELLANEOUS.
On the *‘ Ankiindung eines systematischen Werkes von den Schmetter-
lingen der Wienergegend’ of Schiffermiiller and Denis, By Lovts
B. Prout, F.E.S.
Tur well-known ‘Systematisches Verzeichniss der Schmetterlinge
der Wienergegend’ by the above-named authors is invariably cited
by modern writers as dating from 1776, and I am not aware that
that date has ever been challenged. I have, however, for some
years past, been acquainted with a copy of the same work under a
different title, bearing date 1775, and have been somewhat puzzled
by it; but as my studies have not hitherto necessitated a critical
investigation of the work, I have not until quite recently taken up
the question seriously. In doing so, however, I find sufficient
evidence to be worthy of publication, and by the advice of
Mr. John Hartley Durrant, with whom I have been in corre-
spondence on the subject, I have prepared the following notes.
In the library of the British Museum (Bloomsbury) is the copy
which first arrested my attention, coming from the library of
Sir Joseph Banks, and quite accurately catalogued by Dryander in
‘ Bibl. Banks.’ i. p. 254. The title reads “ Ankitindung | eines |
systematischen Werkes | von den Schmetterlingen | der Wiener-
gegend | herausgegeben | von einigen Lehrern | am k. k. Theresi-
anum. | Wien, | verlegts Augustin Bernardi Buchhiindler. 1775.”
Another copy, identical with this, was acquired by Smith in 1784
from Linné’s library, and is, of course, preserved by the Linnean
Society ; this latter is of considerable interest, as it is still in the
original boards, while that at Bloomsbury has been re-bound, and
lettered on the back “ Ankiindigung eines Werkes von den Schmet-
terlingen der Wienergegend. Wien. 1775.” With the exception of
the titlepage and the frontispiece, this early issue is identical with
the well-known ‘ Systematisches Verzeichniss’ of 1776, so that, as
Mr. Sherborn says (i Uitt., 13th Feb. 1900), * there was only one
printing of the body of the work—the same broken letters occur in
every copy.” Mr. Sherborn further writes :—‘ Note that the 1775
T.P. isa ‘ woodblock,’ not type set, and the 1776 T.P. is engraved on
copper.’ The frontispiece and plates are coloured, but the former is
arranged as in the wncoloured 1776 copies. A third issue (1776 also)
has coloured plates, but the frontispiece is somewhat differently
designed. I have seen no copy of the ‘ Ankiindung’ excepting the
two above mentioned, but the following bibliographical references
deal with it.
Mr. Durrant called my attention to the fact that Hiselt, ‘ Ges,
Syst. Lit. Ins.’ 203 (1836), and Percheron, ii, 89-40 (1837), both
Miscellaneous. 159
cite ed. 1775—the latter, however, under the title ‘ Systematisches
Verzeichniss’ &c. This latter, perhaps, accounts for Hagen’s slip
(i. 167), where, under Denis, he gives nearly the 1776 title, with
the 1775 date. Engelmann (505) falls into just the opposite error,
citing the 1775 title with date 1776.
Hagen (ii. 122), under Schiffermiiller, gives “i. Ankiindigung
.... 1775,” but only at second hand, as he had not seen it.
Percheron (J. c.) quotes among the reviews of the work ‘“ Jena
gel. Zeit. 1775, p. 825.” As there is no copy of the ‘Jenaische
Zeitungen von Gelehrten Sachen’ in this country, 1 wrote to
Dr. O. Taschenberg, in Halle, who very obligingly sent for a copy
from Jena and wrote me out the review verbatim. It appeared in
“ rxxxxviiI Stiick. Freytags den 8 December 1775,” pp. 825-826,
and described the work as ‘ Ankiindigung .... 1775. 322 S. in er.
4, nebst 8 Kupfertafeln,” quite accurately, except that I suppose 8
to be a typographical error for “3,”’—an error, however, which has
been followed by Schréter and by ‘“ L'Esprit des Journaux.” The
reviewer says, ‘‘ Dieses hier angekiindigte fiirtrefliche Werk
haben wir vorziiglich den Herren Professoren Schifermiiller [sic]
und Denis zu danken,” and goes on to discuss appreciatively the
authors’ plan. ‘The title ‘« Ankiindung” was of course indicative of
the fact (well-known to lepidopterists) that even this great catalogue
of 322 pp. was only intended as the precursor of a much greater
work which never appeared. But the title in question has given
rise to much confusion, and it is highly probable, as Dr. Taschenberg
suggests, that it was changed in 1776 to ‘‘ Systematisches Verzeich-
niss” &c. because its scope had bcen misunderstood. It is quite
obvious, as Mr. Durrant points out, that there cannot ‘“ have been
such a demand for this work in 1775 as to necessitate a second
edition in 1776” (in litt., 9 Feb. 1900); moreover, Mr. Sherborn
has shown from internal evidence that there is only one printing in
question. Mr, Durrant inclines to regard the 1775 copies as
‘advertisement advance-copies with a temporary title.”
Other contemporary references to the ‘“‘ Ankiindung” may be
briefly noted as follows—I give nothing which I have not personally
verified :—
Schroter, ‘“‘Abhandlungen iiber verschiedene Gegenstiinde der
Naturgeschichte,” erster Theil, 1776, ‘‘ Vorrede” (dedication is
dated 28 Feb. 1776): ‘* Mein Manuscript war bereits in den Hiinden
meines Herrn Verlegers, als mir noch eine Arbeit zu Gesichte kam
die ich nicht tibergehen kan.” He then gives the title ‘* Ankiindig-
ung” &c., 1775, and remarks that the book might appear to many
too detailed for a mere *‘ Ankiindigung,” &c., &c.
‘ L’Esprit des Journaux’ for April 1776, p. 400, gives “* Ankundig-
ung [ste] einer systematischen werks von der sehmerligen” [sic!},
&c., 1775, very probably merely after the account in the ‘Jena.
Zeit.’
Schulze, in ‘ Neue Mannigfaltigkeiten,’ iv. pp. 26 (bis—recte 28)
et seq. (Berlin, 1st June, 1776), reviews the ‘* Ankiindigung....
1775” very fully.
Berlinische Sammlungen,’ ix. p, 219 (recte 308), 1777, refers to
160 Miscellaneous.
both issues, apparently assuming that of 1775 to be a true An-
kindigung.
Cobres, ‘Delicize Cobresiane,’ p. 387 (178%), gives “ Syst.
Verzeichniss .... 1776,” but adds, ‘ Anfiinglich kam es unter dem
Titel Ankiindigung eines systematischen Verzeichnisses [sic]... .
heraus.”
Prange, ‘ Verz. Schrift. Naturgesch.’ p. 207, copies Cobres.
The earliest review which I have seen of the issue entitled
‘Systematisches Verzeichniss’ &c., is Erxleben’s, in his ‘ Physikal-
ische Bibliothek,’ iv. Stiick 2, pp. 190-207 (1777); but others soon
folluw. In Denis’s own work cn ‘ Die Merkwiirdigkeiten der k. k.
garellischen 6ffentl, Bibliothek am Theresiano’ (Wien, Bernardi,
1780), of which the preface is dated 3rd Nov., 1778, we find
(pp. 16-32) a chronclogical list of works in the library which
appeared from 1747 to 1778. Under 1775 there is no mention of
the ‘ Anktindung, but under 1776 the Syst. Verz. is duly regis-
tered, with the comment ‘‘ durch des ersten [7. ¢. Schiffermiiller’s |
Beforderung zum Directorate des so genannten nordischen Colle-
giums in Linz ist die Fortsetzung dieses Werkes wenigstens auf
einige Jahre gehemmet worden.”
Mr. Durrant calls my attention toa possible further complication.
Engelmann, 514, and Hagen, 11. 122, quote the 1776 edition as
“Wien (Beck)”; but all the copies which I have seen or which
are noticed in the early writers were published by Bernardi; and
Dr. Taschenberg explains to me that the fact that Engelmann places
** Beck” in brackets indicates that that name will not be found on
the titlepage, but that ‘‘ das ist entweder der Commissions verlag
oder der Eigentiimer der Firma Bernardi” (in ltt., 8 Mar., 1900).
I have at present quite an open mind as to whether the date 1775
should be accepted for this work or not; but in the meanwhile I
venture to make a practical suggestion that for “ priority ” rank it
should be placed after the other 1775 literature (Fabricius, Syst.
Ent.; Naturforscher vi., vii., &c.), but before that of 1776 (Sulzer,
Abgek. Geschichte; Miiller, Zool. Dan. Prodr.; Naturforscher viii.
&e.). I also gladly follow Mr. Sherborn’s suggestion (in litt.), and
recommend that henceforth it be cited as ‘ Schmett. Wien,’ as this
does not cause any confusion in the title.
246 Richmond Road, N.E.,
16th March, 1900.
Abundance of the Greater Sul-Smelt (Argentina silus, Ascan.) on the
Market of Boston, Lincolnshire. By G. A. Boutenenr, F.R.S.
The Natural History Museum has received, through the kindness
of Mr. W. H. Shrubsole, F.G.S., specimens of this rare fish, measur-
ing up to 14 inches, which had been sent to him by Mr. F. Kime,
with the information that great numbers had been brought on the
25th June to Boston and sold in every direction. The only record
of the capture in English waters of this deep-sea fish well known
from the coast of Norway is of a quantity taken on the 15th June,
1898, off the south coast of Ireland (cf. Holt, Journ. Mar. Biol.
Assoe. v. 1898, p. 341). Mr. Kime writes to me that these fish,
which excited great curiosity on the market, were caught in deep
water somewhere about the coast of Norway.
THE ANNALS
MAGAZINE OF NATURAL HISTORY,
(SEVENTH SERIES. ]
~~
No. 32. AUGUST 1900.
XVI.—Notes on Diptera from South Africa (Tabanide and
Asilidz). by Miss Gerrrupe Ricarpo.
THE Diptera treated of in this paper are those collected by
Mr. Distant, chiefly from the Transvaal, but a few come trom
Angola and Nyasaland. The new species will be figured
later in his work on the insect fauna of the Transvaal. The
other families represented in his collection will be described on
a future occasion.
Tabanida.
P4ANGONINE.
Cadicera nigrescens, 3, sp. n.
One female from Zomba, British Central Africa (Rendua/i).
This species, rather resembling C. rubra-marginata, Macq.,
differs from it in the following particulars :—T'he antennz are
imperfect in this specimen. ‘The red colouring on the abdo-
men is not so widespread, there is xo central dorsal red stripe
on the first three segments, the red at the sides is less marked
and does not begin till the third segmenr, on the ventral side
it extends as a band on the last segment. There is a greyish
band on the forehead behind the antenne, extending to the
sides, plainly visible when viewed from behind. The palpi
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. ll
162 Miss G. Ricardo on
are not quite so thick. The wings are rather darker and the
insect is smaller.
Length 15 millim.
Type (female), Zomba, British Central Africa (Rendall).
PanGontia, Rond.
Pangonia biclausa, 2, Loew, Dipt. Siidafrik. p. 19 (1860).
One male from Pretoria (W. Z. D.).
This agrees with the description of the female given by
Loew ; it is one of the species with the fourth posterior cell
closed (see Ricardo, Ann. & Mag. Nat. Hist. (7) v. p. 109,
and remarks on P. Br evis, Loew, which will es apply to this
species).
Pangonia chrysopila, Macq., Hist. Nat. Dipt. i. p. 194 (1884) ;
Walker, List Dipt. pt. v. Suppl. 1, p. 137 (1854).
One female from Pretoria (Rendall).
Probably belongs to this species. Macquart’s description ts
very meagre; if correct, it belongs to Pangonia, Rondani,
the eyes being bare.
Pangonia angulatu, Fabr., Syst. Antl. p. 91. 5 (1805);
Wiedem., Auss. zweifl. Ins. 1. p. 97 (1828) ; Macq., Ann.
Soc. Ent. Fr. vi. p. 429, tab. xv. fig. 4 (1837) ; Loew,
Dipt. Stidafrik. p. 20 (1860).
Panyonia obesa, Walker, List Dipt. pt. v. Suppl. 1, p. 185 (1854) ;
Schiner, Reise der Nov ara, p. 99 (1866).
One female from Stellenbosch, 1888.
Corizoneura varicolor, Wiedem., Auss. zweifl. Ins. i, p. 98
(1828); Walker, List Dipt. pt. i. p. 134 (1848), pt. v.
Suppl. 1, p. 137 (1854) ; Loew, Dipt. Siidafrik. p. 17
(1860).
Gne temale from Pretoria (W. L. D.); two males and two
females trom Namaqualand (Cochrane); two males from
Barberton (Rendall); one female from Kowie, Jan. 1895
(F. Pym) ; one male from Pretoria (W. L. D.), Oct. 1895.
The female from Kowie has the first posterior cell closed
and pedunculated, similar to a female in the British Museum
collection ; there is no doubt these belong to the above species,
which evidently must be added to those other species of Pan-
gonine which vary in this particular in different individuals.
The males have the prolongations on the fore tarsi (see
Diptera from South Africa. 163
Ann. & Mag. Nat. Hist. (7) v. p. 110); one male is very
dark in colouring, and all the males have the thorax darker
than in the females.
Corizoneura suavis, 2, Loew, Dipt. Siidatrik. p. 17 (1860).
One female from Pretoria (W. LZ. D.).
Corizoneura lateralis, Fabr., Syst. Antl. p. 9L. 4 (1805) ;
Wiedem., Auss. zweifl. Ins. 1. p. 101 (1828); Walker,
List Dipt. pt. i. p. 1384 (1848), pt. v. Suppl. 1, p. 1385
(1854) ; Loew, Dipt. Siidatrik. p. 17 (1860).
One male from Cape Town.
The stripes on the thorax are not distinct, nor is the white
spot on the wing, but the eyes not quite meeting agree with
Loew’s description.
Rhinomyza denticornis, Wiedem , Auss. zweifl. Ins. 1. p. ILL
(1828); Walker, List Dipt. 1. p. 192 (1848); Loew, Dipt.
Siidatrik. p. 21 (1850).
Stlvius denticornis, Wiedem., J. c.
Dichelacera binotata, Macq., Dipt. Exet. i. p. 113.
One female from Barberton (Rexdull) ; two females from
Pretoria, one male trom Rustenburg (IW. L. /).
Chrysops stigmaticalis, Loew, Dipt. Sidafrik. p. 29, tab. 1.
fig. 18 (1860).
‘Three females, all from Pretoria.
1d
ABANINZ.
Tabanus biguttatus, Wiedem., Auss. zweill. Ins. u. p. 623
(1828); Walker, List Dipt. pt. v. Suppl. 1, p. 231
(1854); Loew, Dipt. Siidafrik. p. 37 (1860); Karsch,
Berlin. ent. Zeit. xxxi. p. 370 (1887).
Tabanus cilipes, Macq., Dipt. Exot. i. p. 120 (1838); Walker, 7. ¢.
p. 286.
Tabanus cerberus, Walker, /. c. pt. i. p. 48 (1848).
Tabanus tripunctifer, Walker, Zool. viii. Appendix 95 (1850).
Tabanus noctis, Walker, Dipt. Saund. pt. i. p. 42 (1850).
One female from Graham’s ‘Town, June 1893 (Rev. Wal-
lace) ; one temale trom Fort Johnston, Nyasaland (Rendall) ;
one female from Namaqualand (Cochrane).
i
164 Miss G. Ricardo on
Tabanus sagittarius, Macq., 1. c. p. 123; Walker, List Dipt.
pt. v. Suppl. 1, p. 228 (1850).
Tabanus socius, Walker, l. c. pt. i. p. 160.
Tabanus serratus, Loew, 1. c. p. 39, tab. i. fig. 21.
Nine females from Pretoria (W. ZL. D.); eight females
from Fort Johnston, Nyasaland (Rendall); one female from
Uganda (Ansorge).
Tabanus latipes, Macq., 1. c. i. p. 119; Loew, 7. c. p. 36;
Peters, Reise nach Mossambique, Zool. v. 2 (1862).
Tabanus latipes, Walker, List Dipt. pt. i. p. 236, pt. v. Suppl. J, p. 828
(1854).
Fabanus fenestratus, Walker, Zoologist, viii. Appendix 67 (1850) ;
List Dipt. v. Suppl. 1, p. 219 (1854),
Tabanus africanus, Gray.
One female from Zomba, British Central Africa; three
females from Angola (Montero).
Tabanus nyase, 2, sp. n.
One female (type) from Fort Johnston, Nyasaland (Ren-
dall).
Belongs to division B A638 of Loew. Eyes naked. No
appendix. Fore tibie not thickened. Wings clear.
Brown. Face and palpi greyish, the latter with short
black jubescence on the upperside. Forehead yellowish
grey, with a shining reddish oblong spot above antenne, a
stripe ot the same colour about half as long again proceeding
from it towards the veitex. Antenne with the first joint on
its upper half thickly covered with black hairs, the second
reddish, the third on first annulation greyish. Thorax with
giey tomentum and five grey stripes. Abdomen with trian-
gular greyish spots on segments 2-5 forming an irregular
stripe; on the side of each segment a greyish oblong spot, not
reaching ile anterior margin; the posterior margins of seg-
ments aie narrowly grey; the extreme lateral margins of
fifth and sixth segments are fulvous ; the dorsum of abdomen
covered with short black pubescence. The ventral part of
abdomen is reddish, with an indistinct black stripe in the
centie. legs with Jorg grey and some black hairs on anterior
femoraand tibiae; the posterior tibie have shorter pubescence,
chiefly black. Wings hyaline, the fore border and base
hrownish yellow, veins brown. Halteres brown, with yellowish
club.
Length 153 millim. ; width of head 5 millim.
Diptera from South Africa. 165
Tabanus nigrohirtus, ?, sp. n.
One female in British Museum Ooll., labelled ‘‘ Bonny,
July 1872, 73. 66”; two females in Mr. Distant’s Coll.
labelled “ Bonny, July 1872.”
Belongs to division BAb2 of Loew. Eyes naked. No
appendix. Fore tibiz not thickened. Wings shaded.
Reddish brown. Face red, with white pubescence ; palpi
yellow, with black pubescence. Beard white. Forehead
greyish, with a long red-brown stripe in the centre. Antenne
red, the apex of the joints black; pubescence on the first two
joints black, on the third ereyish. Thorax with three in-
distinct black stripes, the middle one divided by a narrow
median line; covered with hoary tomentum and short black
pubescence, with longer black hairs on the sides; breast red,
with grey pubescence and tomentum. Scutellum red, with
some hoary tomentum. Abdomen dark red, with a narrow
dorsal greyish line extending from the first to the fifth seg-
ment; the whole surface of the abdomen is covered with
short black pubescence, which is usually thickest on the
fourth and following segments; on the extreme lateral
margins there are some white hairs; in one specim3n the
median line is indistinct: the underside of abdomen ts lighter
red, with less black pubescence and some white hairs; the
posterior margins of the segments are lighter in colour and
only the extreme apex dark.
Legs reddish yellow, the coxe with grey pubescznce;
the anterior femora, the apical half of the tibize, and all the
tarsi are blackish, and sometimes the posterior femora; the
hairs on the outer edge of the femora are white, and there is
some white pubescence on the tibiz, elsewhere it is black.
Wings grey, on fore border yellowish, veins brown; no
appendix on fork of third longitudinal vein.
Length 16 millim.; width of head 3 millim.
Type (female), Bonny, July 1872, 73. 65, in British Mu-
seum Coll.
Aitylotus nigromaculatus, 2, sp. n.
Thirteen females; five from Pretoria, the rest from Fort
Johnston, Nyasaland.
Resembles Tabanus bipunctatus, Wulp (‘ Notes Leyden
Museum,’ vii. p. 72, 1885), from West and South Africa, in
having two black spots on the forehead ; but the eyes have a
band which Wulp expressly mentions as non-existent in his
species, and the abdomen does not agree with his description.
I theretore venture to make it a new species.
166 Miss G. Ricardo on
Dark grey. Face and palpi whitish, with white hairs ;
some black pubescence on the upperside of the latter, more
marked in some of the series than in the type. Beard
scanty and whitish. On the forehead are two distinct black
shining spots, the upper one near the antenne lheart-shaped;
with a longitudinal groove, the other oval. ‘The pubescence
on the eyes is very slight, hardly perceptible ; there is a cross-
band extending to opposite the cordiform spot. The antenne
are light yellow, the first joint greyish, with some black hairs ;
the apex of the third joint is more orange in colour, Thorax
with greyish or light yellow short pubescence. Abdomen
with a distinct central grey stripe extending from the first
to the sixth segment; this is not so distinct in those from
Nyasaland ; on each side is another stripe more yellow in
colour, not extending beyond the fifth segment. The ventral
side has no stripes, clothed with short light yellowish pubes-
cence. Legs reddish yellow, the coxe with light grey pubes-
cence, the anterior tibiae at their apex and the tarsi dark
fuscous; on the posterior legs only the last four joints of the
tarsi are darker.
Wings hyaline, the fore nerves yellow; a short appendix
on the fork of the third vein, not always present.
Length 124 millim.; width of head 4 millim.
Type (female), Pretoria (W. L. D.).
Asilida.
Laparus albicinetus, 2 , sp. n.
One female from Barberton (P. Rendall).
This species will belong to Loew’s division A } (see Dipt.
Siidafrik.). Fore tibize having an end spur and four to six
bristles on the anterior margin of the mouth.
Black. Face brown, shining, with testaceous tomentum at
the sides, bordering the eyes, and round the mouth. The
four large bristles near the mouth are black, with a few black
hairs below. Antenne red, the first joint and the base of the
second blackish, a large bristle on the latter. Forehead
black, with some greyish-red tomentum; a short central
longitudinal furrow extends from the antenne to the ocelli-
eerous tubercle. The bristles and hairs on the hind part of
head are black. Proboscis and palpi black, the latter with
black hairs.
The thorax is black, with a whitish band on each side
extending the whole length, and three faint white stripes on
the dorsum, only noticeable on the anterior border. Scutellum
Diptera from South Africa. 167
bordered with grey. ‘The breast-sides shining black, with
very faint reddish-grey tomentum. Abdomen blue-black,
shining. Halteres brown. Legs dark red, the coxe and last
joints of tarsi, the fore femora, and the base of the posterior
femora black; there are no bristles on the fore femora; all
the bristles and pubescence on the legs black.
Wings light brown, lighter at the apex, the opening of the
fourth posterior cell is a little narrowed.
Length 14 millim.
Type (female), Barberton (P. Rendall).
Microstylum dispar, Loew, Dipt. Siidafrik. p. 78 (1860).
Four males from Rustenburg, two males from Pretoria.
Microstylum nigribarbatum, Bigot, Ann, Soc. Ent. Fr. (5)
vil. p. 408 (1878).
One male from Pretoria probably belongs to this species,
though the third joint of antenne is reddish, not black, and
the hind tibia have white hairs on the underside, extending
nearly halfway, of which no mention is made by Bizot.
Microstylum acutirostre, Loew, Ber. d. Akad. d. Wissench.
zu Berlin, p. 658 (1852) ; Dipt. Siidafrik. p. 79 (1860) ;
Peters’s Reise, p. 7, tab. 1. figs. 5, 6 (1862).
One male from Fort Johnston, Nyasaland.
The third joint of palpi is red on its apical half, not wholly
black; the last segment of abdomen has an orange-yellow
border; the double black stripe on the thorax is not wider
anteriorly, nor reddish ; and the wings are not so dark as in
Loew’s description.
Microstylum rufinevrum, Macq., Dipt. Exot. Suppl. v. p. 49
(1850).
One female from Barberton (Rendall).
The pubescence on the first joint of tarsi is orange-red
instead of white; the wings are hardly brown as in Mac-
quart’s description, but greyish. Macquart’s type came from
Gaboon,
Microstylum spurinus, Walker, List Dipt. pt. ii. p. 308
(1849).
Dasypogon spurinus, Walker, J. ¢.
One female from Pretoria.
168 Miss G. Ricardo on
This differs from the type in having stiff black bristles on
the thorax, instead of long white hairs, otherwise it seems
identical with Walker’s type.
Microstylum glabrum, 2, sp. n.
One female from Pretoria (W.L.D.); one female from
Zoutpansberg (Kessner).
This species closely agrees with Loew’s description of
M. gulosum, g, but is probably a distinct species; it differs
in the following particulars from M. gulosum :—
The abdomen is dull black on the first four segments,
shining black on the last four; there is no pubescence on the
dorsum of abdomen, and only some on the first segment of
the underside. Legs black; the pubescence on the femora
and the posterior tibie white, on the anterior tibize black.
The first posterior cell of the wings is not narrowed at the
apex; the 4 spines at the apex of the tibie are very stout.
Length 26 millim.
Type (female), Pretoria (W. L. D.).
Microstylum elegans, § & 2, sp. n.
Three males and two females from Pienaars River (W.
L. D.); two females from Pretoria (W. Z. D.).
Belongs to division Ba of Loew.—Moustache confined to
the lower part of the face, thick, with no stout bristles inter-
mixed. ;
Black. Face with yellowish-white tomentum ; the mous-
tache composed of long white hairs ; palpi black, covered
with long white hairs; beard white; antenne black, with
black hairs on the first two joints. Forehead with two
whitish spots reaching from the eyes towards the centre,
the remaining portion black; at the sides are long white
hairs reaching to the antenng; on the posterior half they
are black; on the hind part of head and on the collar
white. Thorax brown, witha broad grey stripe on each side,
clothed with long white hairs and bristles, which extend to
the posterior part of the thorax; seutellum grey, with long
white pubescence ; breast-sides brown, with white tomentum
and a few white hairs.
Abdomen black, with very distinct grey-white bands on
the posterior border of each segment, narrowest on the last
one, extending up the side of each segment; they are bor-
dered by a slight fulvous edge, more noticeable when viewed
from behind; the whitish pulescence on the abdomen is short
and spare, except at the sides of the first two segments, where
Diptera from South Africa, 169
it is long; the underside of abdomen is grey, with some
scattered white hairs. Legs black; the coxe grey, with
long white hairs; the femora and tibie with short white
pubescence; all the bristles on legs are white, with the ex-
ception of some on the last four tarsal joints. Halteres brown.
Wings clear, grey shading on the fore border as far as the
second longitudinal vein, extending into the submarginal cells,
where it gradually ends in a point; it also fills the centre of
the first and fourth posterior cells, and there is a small irre-
gular patch of it in the basal half of the second posterior
cell; veins yellow ; the first posterior cell is nearly closed
in some specimens, in others hardly narrowed.
Length 27 millim.
Type (female), Pienaars River (W. LZ. D ).
The female is very similar, but the last three segments of
the abdomen are wholly black, though on the side of the sixth
the grey stripe is present ; the abdomen is more shining and
less pubescent. The pubescence on the legs is a little
slighter. The first posterior cell of the wings varies as
above.
Length 30 millim.
Type (female), Pienaars River (W. LZ. D.).
‘These fine flies were found flying near the ground, on
bare open veldt. One female was caught attacking a grass-
hopper.
Microstylum nigrescens, § & 3, sp.n.
Two males and two females from Angola, in British Museum
Coll., 76.55 and 73.663; one male and two females from
Angola (Monteiro), in Mr. Distant’s Coll.
Belongs to division B 6 of Loew.—Moustache confined to
the lower part of the face, thin, with very stout bristles inter-
mixed.
3. Black. Face brown, with yellowish or whitish tomen-
tum; the moustache consists of four large black bristles,
intermixed with a very few black hairs; palpi black, the
tips reddish, clothed with black hairs. Beard black. An-
tenne black, the first and second joints more or less red with
black hairs and bristles. Forehead with an oblong shining
black spot on the vertex, continued as a narrow black line
to the anteune, sides greyish with black hairs and bristles,
‘he hind part of head grey, with black hairs. ‘The collar
covered with greyish tomentum, ieddish on the posterior
half in the centre. ‘Lhorax black ; the shoulders, sides, and
posterior halt reddish, with greyish tomentum, forming one
ceitral bruad stripe divided in the middle, and one on each
170 Miss G. Ricardo on
side, not attaining the fore border; all the bristles and the
short hairs on the dorsum are black. Scutellum red, with
grey tomentum.
Abdomen dull black, some dull reddish colour on the
second, third, and fourth segments, becoming more or less
distinct bands on the posterior margins; some very short
black pubescence on the sides; genital organ red, with black
pubescence. Halteres red. Legs bright red, with black
pubescence and bristles; the coxe are brown, with greyish
tomentum. Wings light brown, veins yellowish red,
Length 20 millim.
Type (male), Angola, 73. 66 in British Museum Coll.
?. The greyish stripes on the thorax are not so distinct.
The last three segments of abdomen are shining black, the
extreme edge of the last one reddish. Wings in the type
are brownish, but one female has almost clear wings; the
veins are yellowish red in all, those on the fore border deepest
in colour.
Length 26 millim.
Type (female), Angola 73. 66 in British Museum Coll.
Scylaticus rufescens, 3, Sp. Nn.
One female from Barberton (P. Rendall).
Allied to S. costalis, Wiedem., but distinguished from it
by the orange-red moustache, wholly red antenne, and the
presence of grey spots on the second segment of the abdomen.
Black. Face with white tomentum. The moustache,
which extends nearly halfway up the face, consists of bright
orange-red hairs. Palpi and proboscis black, with reddish
pubescence. Beard reddish. Antenne red, the first two
joints with red hairs. The forehead black with reddish hairs.
The hind part of head bordered with orange-red hairs.
Collar and thorax black, the latter with the shoulders and
posterior margin red, covered with yellowish pubescence.
Scutellum red, with some long reddish bristles. Abdomen
with the first segment black, the anterior border of the
second black, with an oblong grey spot on each side,
the space between and behind the spots is red, and the
posterior border of the segment consists of a light yellow
continuous band; the third, fourth, and fifth segments are
orange-red, bordered with a zigzag black band, and on their
lateral margins the light yellow band appears; the sixth and
seventh are almost wholly orange-red; the pubescence on
the abdomen consists of sparse light reddish hairs, longer
and lighter in colour at the sides: on the underside the
Diptera from South Africa. 171
yellow bands are continuous on the second, third, and fourth
segments. Legs red; the coxe black with grey tomentum,
the hairs and bristles on the legs are whitish yellow. Halteres
red. Wings hyaline, on the fore border brown, reaching to
the discal cell, but not extending to the base of the wing;
the fourth posterior cell is slightly narrowed at the opening.
Length 133 millin.
Type (female), Barberton (P. Rendall).
Damalis speciosa, 3, Loew, Dipt. Siidafrik. p. 108 (1860).
One female from Barberton (P. Rendall). Is apparently
the female of this species.
The third joint of the antenne, which was wanting in
Loew’s type, 1s black, with the long bristle black, and white
towards the tip. ‘The pubescence on the dorsum of the
abdomen is sparse, so that the ground-colour is seen, dark
reddish with transverse black bands.
Lamyra gulo, Loew, Bemerk. iiber Fam. Asiliden, p. 19
(Berlin, 1853) ; id. Dipt. Stidafrik. p. 113 (1860).
One male from Lydenburg District (Zutrzenka); one
male from Waterberg District (Wi/de); two males from
Angola (Monteiro).
Vhe above specimens have three white bands on the
abdomen, the first segment having a rather narrower one
than the other two, leaving the anterior border free, but
almost reaching the posterior border; it is not visible on the
underside. In ‘ Bemerk. ittber Fam. Asiliden,’ Loew men-
tions the three bands, but in his ‘ Dipt. Siidafrik.’ he only
mentions two bands. ‘he third joint of the antenne is
somewhat grey. In every other particular these specimens
agree with the description. They vary from 20-32 millim.
in length. ‘The two from Angola are both injured and have
lost the third joint of antenne.
Laphria aureoptilosa, 8, sp. n.
One male from Durban (W. Z. D.).
Blue-black, shining. Face covered with yellowish to-
mentum, the lower part is occupied by a large black tubercle ;
below the antennez are black bristles and hairs, with a few
long golden-yellow hairs, which extend down each side of
the face. The moustache is composed of black bristles and
some light yellowish ones on the lower half. Beard yel-
lowish. Palpi small, black. Antenne black, the first two
172 Miss G. Ricardo on
joints with thick black pubescence. Forehead black, with
black bristles and hairs. Thorax covered with short dense
yellow hairs. Breast-sides have some grey tomentum and
black pubescence. Scutellum and abdomen have similar
pubescence to that of the thorax; genital organ black,
shining, with black pubescence. Underside of abdomen
black, with yellow hairs. Legs black; the anterior and
middle coxe covered with grey tomentum; the pubescence on
the legs consists of long light yellowish hairs, except on the
apex of tibia and first joint of tarsi, which are clothed under-
neath with dense fulvous pubescence, and the last four joints
of all the tarsi, which have black pubescence; hind tibize
much thickened, red at the extreme base, no bristles on the
underside. Halteres yellow. Wings dark brown, clear on
the basal half, veins dark brown.
Length 13 millim,
Type (male), Durban (W. Z. D.).
Heplistomera nolilis, Loew, Dipt. Siidafrik. p. 120 (1:60).
‘Two males and two females from Fort Johnston, Nyasa-
land (P. Rendall); one male from Lydenburg District
(Zutrzenka).
Lawenecera zonata, Loew, Dipt. Siidafrik. p. 123 (1860).
One female trom Pretoria (W. L. D.).
Laxenccera albicinta, Loew, Dipt. Siidafrik. p. 122 (1860).
One female from Pretoria (W. L. D.).
Dasythrix brachyptera, Loew, Bemerk. tiber. Fam. Asiliden,
p. 21 (Berlin, 1853) ; id. Dipt. Siidafrik. p. 126 (1860).
D. brachyptera, 3, Loew, and DP, stenura, 3, Loew (Dipt.
Siidafrik. p. 125), are probably the male and female of one
species, as Loew himself suggests, and Gerstaécker, in Decken’s
Reisen in Ost-Afrik., concurs; if thisis the case, brachyptera
having priority must be adopted.
One male from Pretoria (W. LZ. D.); two females from
Pretoria (W. L. D.).
These three specimens from the same place seem to con-
firm the above supposition, being very similar in general
appearance,
Promachus fulvipes, Macq., Dipt. Exot. 1. (2) p. 98 (1888) ;
Loew, Dipt. Siidafrik. p. 182 (1860),
Oue male from Pretoria (W. L. D.).
Diptera from South Afric c. 173
Promachus albicinctus, 6 & 2, sp. n.
One male and three females from Pretoria (W. L. D.).
Allied to fu/vipes, but easily distinguished from it by the
wholly black antenna.
g and 2. Grey. Face brown, covered with dense yellow
tomentum. The moustache consists of yellow bristles, from
which there extends up each side of the face a row of long
yellow hairs reaching to the antenne, and ending with a few
black bristles and hairs. Palpi black, with long yellow
hairs. Beard white. Forehead with black bristles and
hairs on the anterior half, the posterior with yellow ones.
Antenne black, the first two joints with yellow pubescence.
Hind part of the head in the centre with yellow bristles, at
the sides yellow and black intermixed, with long white hairs.
Thorax brown, with the central stripe rather lighter; the
shoulders, sides, and anterior half covered with dense reddish-
brown tomentum, which borders the side-stripes and runs
between their several divisions as a narrow line; the pubes-
cence is black on the dorsum, with a fringe of longer white
hairs on the fore border, some scattered ones on the sides
and on the posterior part. Scutellum brown, with grey
tomentum, black bristles, and white hairs. Breast-sides
brown, with reddish-brown tomentum and sparsely scattered
black and white hairs; the tuft over the halteres is red.
Abdomen grey, with transverse black bands, which take up
more than the anterior half of each segment, not reaching
the side border, with rounded corners; the pubescence
whitish yellow, thicker on the first three segments; the
underside grey, with white pubescence; the genital organ
shining black. Legs chestnut-red; the coxz black, with grey
tomentum and long white hairs; the anterior and middle
femora with a black stripe on the upperside; knees black ;
the pubescence on the legs is white, and all bristles black.
Wings clear, yellowish at the extreme base; veins brown,
those on the fore border red.
Length 20 millim.
Types (male), Pretoria (W. LZ. D.); (female), Pretoria
(WES D:)
Promachus bicolor, 3 , sp. n.
One female from Pretoria (W. L. D.).
Black. Face red on the lower half, with yellowish to-
mentum, above black. Moustache extending halfway up
the face consists of yellow bristles below -nd black bristles
174 Miss G. Ricardo on .
above, from it extends on each side of the face to the antenne
a line of short yellow and black bristles. Palpi black, with
white hairs. Forehead black, with black hairs on the ante-
rior half, a few white hairs on the vertex. Antenne black ;
the pubescence on the first two joints is black above, white
below. Beard white. Hind part of head clothed with black
bristles. ‘Thorax black, the shoulders and posterior curners
bright red; some fulvous tomentum on the shoulders and
sides of thorax ; pubescence black, white hairs on the sides
and posterior part. Scutellum black, with black bristles
and white hairs. Breast-sides black, with stripes of grey
tomentum and some sparse white hairs, those over the hal-
teres fulvous. Abdomen iron-grey, with black bands on the
segments nearly reaching to the posterior border, not at-
taining the sides, with rounded corners; the pubescence on
the bands is black, with a fringe of longer white hairs on the
posterior borders of each segment, and some grey tomentum
on the sides of abdomen; viewed from above the abdomen
appears black, with narrow bands of white hairs; the under-
side is black, with white pubescence. Legs red; coxe, knees,
apex of tibia, and tarsi black; a faint indication of a black
stripe on the underside of the anterior and middle femora;
pubescence on the legs white, longest on the anterior coxa ;
all bristles black. Wings clear; veins brown, those on the
fore border and at base red. Halteres red.
Length 22 millim.
Type (female), Pretoria (W. LZ. D.).
Alcimus rub/ginosus, Gerst., Decken’s Reisen in Ost-Afrik.
p. 387 (1873), tab. xvi. fig. 5.
Two females from Zomba, British Central Africa (P.
Rendall); two females from Uganda (Ansurge) ; two males
from Zomba, British Central Africa (P. Rendall).
Gersticker described the female. In the above specimens
the thorax varies from dark red-brown with black stripes to
fawn-colour with the central stripes a little darker and only
the side-stripes black; the former and Gersticker’s type have
probably become denuded of tomentum, leaving the ground-
colour (reddish brown) prominent. ‘The only other differences
between these females and the description of the type are
the following :—The pubescence on the anterior part of the
scutellum is white; the last segment of the abdomen is black,
shining. The fere femora have black bristles on the under-
side, with a few white ones intermixed and have no fringe
of long white hairs; there are white bristles on the tarsi of
Diptera from South Africa. 175
the two anterior pairs of legs, besides tho-e at the tip of the
tibie.
In the two male specimens there are no bristles on the fore
femora, but a few long yellowish hairs, there are some black
bristles on the side of abdomen, the male organ is blackish
with short white pubescence, and the anal appendage bright
red. |
Alcimus longurio, Loew, Dipt. Siidafrik. p. 137 (1860).
Two females trom Durban (W. LZ. D.) are evidently the
female of the above. Loew described the male only. ‘They
differ from his description thus :— ;
The thorax in one of the specimens has the median black
stripes very narrow, posteriorly merged in one broad black
stripe; in the other it is divided by only a narrow umber-
brown line. The fore tibize and the apical half of the posterior
ones, and the first joint of the fore and posterior tarsi, clothed
with short fulvous pubescence on the inner side; the basal
half of the tarsal joints is dark red. ‘lhe moustache extends
to the antenne, with a few black bristles intermixed with some
long yellow hairs. The last segment of abdomen is wholly
black, shining.
Alcimus tristrigatus, 3, Loew, Dipt. Sitdafrik. p. 134, tab. i.
fig. 51 (1860).
One male from Pretoria (W. L. D.).
The third joint of antenne, which was wanting in the
type, is black, and the bristle the same.
Two females from Pretoria (W. LZ. D.).
Though these vary somewhat from the male, I feel con-
vinced they are the same species, their general appearance
being identical. They differ from the description of the male
in the following particulars :—
Antenne wholly black, the moustache continued to the
antennee as a tew scattered white hairs ; cheeks black ; palpi
black, with yellow hairs. The middle stripe of thorax is
darker, the two narrow black lines on the side-stripes of the
male here become respectively a triangular and an oblong
spot; the bristles on the posterior half are black instead of
yellow ; the hairs on the scutellum are all yellow. On the
abdomen the dark spots become a broad band on each segment,
not reaching to the posterior margin; the last sezment is
wholly black, shining; the pubescence on the bands is
yellowish, giving them a fulvous appearance, elsewhere
whitish. Legs black, the tibia on basal half reddish; the
176 Miss G. Ricardo on
pubescence white, becoming fulvous on the basal inner edge
of fore tibia and of the first joint of fore tarsi and on the
inner and under side of the posterior tibia and of the first
joint of tarsi.
Wings with grey shading at the apex, extending to the
first posterior cell, and continuing in a narrow line to the base
of the first submarginal cell.
Alcimus cinerascens, § & 3, sp. n.
Three males and four females from Fort Johnston, Nyasa-
land (P. Rendall).
g.—Grey. Face brown, with whitish tomentum; palpi black,
with white hairs; the moustache consists of a row of black
bristles above the mouth, then of long white hairs, which from
halfway up the face dwindle into a few short ones reaching
to the antenne. Forehead with a few black hairs at the sides.
Antenne red; the third joint black, with grey tomentum, the
bristle black ; the first two joints with black hairs. The hinder
part of head bordered with black bristles intermixed with
white ones, the hairs on sides of head and the beard white,
Thorax and scutellum brown, with grey tomentum; the median’
stripe of the former blackish brown, divided on its whole
length by a narrow brown stripe ; the side-stripes brown, with
an ill-defined black oblong spot on the last three divisions ;
the shoulders reddish ; the bristles on the posterior part of
thorax are black, the pubescence on the scutellum white; the
breast-sides are brown, with grey tomentum and scanty white
pubescence ; the bristles near the halteres white.
Abdomen reddish brown, with darker bands on the seg-
ments, bordered posteriorly by narrow bands of white tomen-
tum, and a stripe of white tomentum on each side; the
pubescence on the dark bands is black, elsewhere white; the
bristles on the sides of the first four segments are white, on
the remaining ones black; the underside cinereous, with grey
tomentum,
Legs red, the fore coxe with grey tomentum and long
white hairs; the posterior coxe partly black, with black
bristles on the outer edges ; the inner side of the femora and
of the anterior and middle tibize, the posterior tibia: except
at the extreme base, and all the tarsi black ; the pubescence
on the legs white ; the fore femora on the basal half and the
fore tibia with long white hairs on the underside ; the poste-
rior tibie and the first joint of the posterior tarsi clothed with
dense fulvous pubescence on the underside; bristles on the
legs black, with some white ones intermixed,
Diptera from South Africa. 17%
Wings clear, a grey shadow in the extreme apex extending
to the third longitudinal vein; the veins brown.
Length 18 millim.
Type (male), Fort Johnston, Nyasaland (P. Rendall).
¢ .—The black bristles of the moustache are only at the sides
of the mouth, those on the back of the head are mostly white.
The last segment of the abdomen is wholly black, shining ;
the bristles on the side of abdomen are wanting, those on the
posterior coxe and the greater number on the legs are white.
Length 22 millim.
Type (female), Fort Johnston, Nyasaland (P. Rendall).
Ommatius fuscovitiatus, g, sp. n.
One male from Pretoria (W. L. D.).
In general appearance this resembles O. jaculator, Walker,
but is distinguished by the first two joints of the antenne
being red.
Brownish grey. Face clothed with silvery- white tomentum ;
moustache composed of white hairs, with two black bristles on
each side ; afew white hairs on the middle of the face, reaching
to the antenne. Palpi black, with white hairs. Beard white.
Antennee red, the last joint and the bristle black ; the hairs
on the first two joints are black. Forehead grey, with
yellowish tomentum, the hairs on the vertex and at the sides
black. The hind part of tne head with a row of black bristles.
Thorax covered with silvery-grey tomentum ; the stripes very
distinct, brown; the median stripe appears widened at its
anterior border and is divided in the centre by a greyish line ;
the short pubescence and the bristles black. Scutellum
brown, with two black bristles. Breast-sides covered with
silvery-grey tomentum, slightly fulvous in the centre; the
scattered hairs are white.
Abdomen brown, with cinereous segmentations ; the pubes-
cence white, the small bristles on the sides of the last
segments black ; the genital organ shining black, the under-
side brown. Legs light yellow; the coxe the same colour as
the breast-sides, with white pubescence ; the knees, the apex
of the first four tarsal joints, and the fifth joint black; the
fore and middle femora with white hairs on the underside,
the middle and posterior femora armed with short black
bristles ; the short pubescence on the legs is black; the
bristles on the outer edge of the anterior tarsi and on the first
joint of the middle tarsi are white, elsewhere black.
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 12
178 On Diptera from South Africa.
Wings clear, veins brown; yellowish at the base and the
whole of the first longitudinal vein.
Length 103 millim.
Type (male), Pretoria (W. L. D.).
Lophonotus albofasciatus, 3, sp. n.
Two males from Pretoria (W. ZL. D.).
Belongs to division I. 2. a of Loew. The abdomen with
bristles before the segmentations, the underside with bristles,
the thorax with long hairs or bristles reaching to the anterior
border.
Brown. Face pale yellow; moustache black, reaching to
the antenne, a few white hairs on the upper half. Forehead
brown anteriorly, fawn-coloured posteriorly, with black bristles
and a few white hairs near the antenne. Beard white. An-
tenne black. Palpi black, with black hairs. Hind part of
head bordered with black bristles, then white hairs. Thorax
bronze-coloured, with some white tomentum on the sides,
stripes indistinct ; the mane composed for two thirds of its
length of short black hairs, the last third of longer white
hairs, on its whole length bordered by black bristles, thicker
at each end; some short white hairs are scattered on the sides
and posterior border. Scutellum the same colour as the
thorax, clothed with long white hairs and a double row of
long black bristles. Abdomen with a black central stripe and
irregular black spots, and reddish-brown tomentum on the
sides ; the pubescence on the first segment consists of long
white hairs in the centre and black ones at the sides; else-
where the pubescence is black, except at the sides of the
apical half, where are some white hairs; the bristles are
black; the genital organ brown above, red below, with black
and white pubescence, longer on the underside ; the underside
of abdomen brown, with some greyish tomentum.
Legs black, metallic, the basal halt of the tibie chestnut-
brown; the bristles black, with the exception of some on the
apex of the tibiz and on the first three joints of the tarsi,
which are white ; the pubescence of legs white, thickest on
the tibia and tarsi. Halteres brown.
Wings clear ; veins brown, red on the fore border.
Length 15} millim.
Type (male), Pretoria (W. LZ. )).).
One specimen is labelled ‘‘ Caught while attacking common
Geometer, Sept. 1895.”
On a Hermaphrodite Frog. 179
XVII.—WNote on a Hermaphrodite Frog.
By R. C. Punnett, B.A.
[Plate IX.]
Last February my attention was called to an anomalous
condition of the genital organs in a frog which was being
dissected by a member of the practical class in this University
(St. Andrews). The specimen was preserved for future
examination, the results of which are now given,
Right Side.
The Miillerian duct was well developed and considerably
convoluted. It possessed a small uterine dilatation, an
opening into the body-cavity, and was slightly pigmented
here and there. The ureter expanded to form a vesicula
seminalis, though this was hardly so well developed as in the
male frog at this period. ‘The ureter and Miillerian duct
were bound together near the cloaca by some connective tissue,
but possessed separate openings into the cloaca. ‘The genital
gland was a well-developed testis. On the side furthest from
the vena cava and about one third of the distance from the
antelior end a small pigmented patch was seen, containing
two white specks (Pl. 1X. fig. 1, *). Sections through the
gland showed that this pigmented portion contained a single
normal ovum (PI. [X. fig. 2, 0v.), whilst the rest of the gland
was purely male and showed no traces of ova either normal or
degenerate. Vasa efferentia were present, and some of them
entered into close relation with the small pigmented ovari:n
patch (PI. IX. fig. 2, ef). The fat-body was well developed.
Left side.
The Miillerian duct was developed as in the normal female
and was slightly larger than that on the opposite side. It
possessed a well-marked uterine dilatation (PI. 1X. fig. 1, wé.).
The ureter showed a small vesicula seminalis and bore the same
relations to the Miillerian duct as its fellow on the opposite
side. The genital gland from external appearance was an
unmistakable ovary, though in size it was slightly smaller
than in the normal females which were dissected at the same
time. At about the middle of the gland and on the side
nearest the inferior vena cava was present a small yellowish
patch, resembling the testis im its external appearance.
Sections showed that this small patch consisted of testis tissue,
containing, like the testis on the opposite side, nearly ripe
12*
180 On a Hermaphrodite Frog.
spermatozoa. The patch was almost isolated from the rest of
the gland, and at one spot (Pl. IX. fig. 3, ov.) contained a well-
developed ovum. It was also devoid of pigment except near
its attachment to the main gland. The vasa efferentia entered
into functional relation with it. With the exception of this
small patch the gland consisted entirely of ovarian tissue
containing ova up to ‘7 millim. in diameter. Vasa efferentia
passed to it as well as to the small testis portion. ‘The fat-
body was well developed. Lastly, it should be mentioned
that the male glandular enlargement on the fore foot was
well marked on both sides.
One of the chief points of interest with regard to this
specimen lies in the completeness of the hermaphroditic con-
dition in the ducts, associated with a sharp distinction be-
tween the sexual characters of the glands. ‘This is the more
remarkable, as at the time the animal was killed (the middle
of February) the breeding-season was not far distant, and
yet neither gland appears to show any very marked prepon-
derance over the other. Though on the whole the testis
seems more developed than the ovary, it is possible that the
peculiar sexual manifestation of activity in this specimen
would, had life continued, have been determined by chance
in the shape of the sex of the first frog which happened to
cross its path.
A full bibliography on this subject is given by F. J. Cole
in the Anat. Anzeiger, Bd. xi. (1895).
Gatty Marine Laboratory,
St. Andrews,
EXPLANATION OF PLATE IX.
Fig. 1. Dissection of the generative organs, slightly enlarged. The upper
portions of the kidneys are not shown. The Miillerian ducts
have shrunk in the spirit and are not so thick as in the fresh
state,
fb. Fat-body. t. Testis.
we. Inferior vena cava. ve. Vasa efferentia.
k. Kidney. vs. Vesicula seminalis.
o. Ovary. ut. Uterine dilatation.
ovd. Oviduct. * Ovarian patch on testis.
ovdo. Opening of oviduct. T Testicular patch on ovary.
r. Rectum.
Fig. 2. Section through the uv 11 patch on the testis. X 30,
bv. Blood-vessel. pg. Pigment.
do. Degenerate ovum (?). st. Seminal tubules.
ov. Ovum. ve, Vasa etferentia.
Fig. 3. Section through the testicular patch on the ovary. x 30, Letters
as in preceding figure.
On new Batiachiars wid Reptiles from Peru. 181
XVIII.—Deseriptions of 2.9 Batvachians and Reptiles col-
lected by Mr. P. O. Simons in Peru. By G. A. BOULENGER,
F.R.S.
Nototrema peruanum.
Tongue subcircular, slightly nicked and free behind. Vo-
merine teeth in two short, straight, transverse series between
the choane. Head moderate, broader than long; snout
rounded, as long as the diameter of the orbit; loreal region
slightly concave ; canthus rostralis distinct ; interorbital space
narrower, or at most not broader, than the upper eyelid;
tympanum distinct, two thirds the diameter of the eye.
Fingers with a very slight rudiment of web; toes one-third
webbed; disks of fingers and toes smaller than the tym-
panum ; subarticular tubercles moderate; a fold along the
inner edge of the tarsus. The tibio-tarsal articulation reaches
the tympanum or the eye. Upper parts covered with smooth
warts of unequal size, the largest of which are parotoid-like
or may be confluent into longitudinal folds; lower parts
granulate. Greenish above, with insuliform black-edged
dark spots, most of which correspond to the larger glandular
tubercles ; limbs with dark transverse bars ; whitish beneath.
Male with a subgular vocal sac.
From snout to vent 43 millim.
Several specimens from Carao, 7000 feet altitude.
Very closely allied to NV. marsupiatum, D. & B. Distin-
guished by the narrower interorbital region and the extra-
ordinary development of the dorsal glands.
Bufo cophotis.
Crown without bony ridges; snout short, blunt; inter-
orbital space as broad as the upper eyelid; tympanum quite
hidden, but eustachian tubes perfectly developed. Fingers
and toes short, flattened, blunt, with double subarticular
tubercles ; first tinger extending a little beyond second ; toes
one-third webbed; two moderate, feebly prominent meta-
tarsal tubercles; no tarsal fold. The tarso-metatarsal articu-
lation reaches the eye. Upper parts covered with smooth
tubercles of different sizes, pierced with large pores, many of
which are comparable to so-called parotoid glands ; such a
large gland on the middle of the upper surface of the leg or
crus. Dark olive; upper parts speckled with black, lower
parts marbled with greyish white. Male without vocal sacs,
182 Mr. G. A. Boulenger on new
with blackish nuptial asperities on the upper and inner sides
of the three inner fingers.
From snout to vent 55 millim.
A single male specimen from Paramo, Cajamarca, 9000 feet.
Several smaller specimens from Carao, 7000 feet.
Nearest allied to B. vartegatus, Gthr. Distinguished by
the well-developed eustachian tubes and the somewhat longer
inner finger.
Paludicola Simonsiz.
Tongue oval, entire. Vomerine teeth none. Snout
rounded, as long as the diameter of the orbit; canthus ros-
tralis distinet ; nostril a little nearer the tip of the snout than
the eye; interorbital space as broad as the upper eyelid; no
tympanum; no eustachian tubes. Fingers and toes moderate,
slightly swollen at the end; first finger not extending as far
as second; toes free; subarticular tubercles moderate; two
rather large, feebly prominent metatarsal tubercles. The
tibio-tarsal articulation reaches the angle of the jaws.
Upper parts with porous smooth warts, some of which are
confluent into longitudinal folds on the body ; throat, belly,
and lower surface of thighs coarsely granulate. Olive-brown
above, whitish beneath ; a dark brown canthal and temporal
streak ; a blackish band in the groin, another on the inner
side of the femoro-tibial articulation, and a third on the outer
side of the tibio-tarsal articulation ; a few blackish spots on
the sides.
From snout to vent 30 millim.
T'wo specimens from Paramo, Cajamarca, 9000 feet.
Allied to P. marmorata, D. & B., but tympanum absent
and first finger shorter than second.
Stenocercus melanopygus.
Pterygoid teeth. Anterior border of ear denticulated.
Upper head-scales very feebly keeled, without transversely
enlarged supraoculars; occipital not enlarged; temporal
scales very teebly keeled. Side of neck with a very short
curved antehumeral fold. Body a littledepressed. No dorsal
crest or denticulation. Dorsal scales large, strongly imbri-
cate, sharply keeled, sharply pointed; the keels forming
continuous lines, which are slightly oblique on the posterior
part of the back; lateral scales smooth or feebly keeled,
passing gradually into the ventrals, which are rounded,
smooth, and considerably smaller than the dorsals; 46 to 50
seales round the middle of the body. The adpressed hind
Batrachians and Reptiles from Peru. 183
limb reaches the ear or not quite so far; fifth toe not ex-
tending as far as second. ‘Tail nearly twice as long as head
and body, tapering, scarcely compressed, scaled like the body,
the scales forming rings. Dark olive-brown above, with
more or less numerous small yellow spots; young with a light
dorso-lateral streak ; lower parts whitish, tinged with green
or blue; anal region, lower surface of thighs, and of base of
tail black in the males.
millim,
Moral; Fen et ieap es a alehaccact er vest oaks 5ifora 187
IESG US iy OS aan & rae ne cere een eee 7
WhUsia lala gout Lavereye bdr other eeeeR a Core 12
ERGY roe pil Oe NOR OE mabey oe eee ace 50
HORE IAM oe. othehc habeas Perel. 26
La bbarel Joie caked Pee ia Ak ry caret CR eR ae 43
Rall pp a) ea IR eR cE ER Fie 98 cp ch on cova 120
A larger specimen, with reproduced tail, measures 85
millim. from snout to vent.
Several specimens from Bafios, Cajamarca, 9000 feet.
Stenocercus Simonsti, Blgr., described from the Andes of
Ecuador, occurs in the same locality.
Stenocercus chrysopygus.
Pterygoid teeth. Anterior border of ear denticulated,
Upper head-scales smooth or feebly keeled, some of the
‘upraoculars feebly enlarged transversely ; occipital not en-
arged ; temporal scales feebly keeled. Side of neck with
olds enclosing shallow pockets covered with granular scales ;
antehumeral fold much stronger than in the preceding
species, with a serrated edge on its lower half, asin S. cupreus,
Blgr. Body depressed. No dorsal crest or denticulation.
Dorsal scales rather large, strongly imbricate, sharply keeled,
sharply pointed; the keels forming continuous lines, which
are parallel or slightly oblique on the posterior part of the
back; lateral scales passing gradually into the smaller
smooth ventrals; 54 to 60 scales round the middle of the
body. ‘The adpressed hind limb reaches the humeral fold or
between it and the ear; fifth toe not extending as far as
second. ‘Tail about twice as long as head and body, tapering,
scarcely compressed, scaled like the body, the scales forming
rings. Bronzy or greyish brown above, with darker or
lighter spots, which may be confluent into longitudinal
streaks; a more or less distinct light dorso-lateral streak
constantly present; lower parts whitish, sometimes bluish
184 Mr. G. A. Boulenger on new
grey, the throat usually with a dark grey network; anal region
and lower surface of hind limbs bright yellow in the males.
millim.
Totalletipth.. fect. s eee eee 238
TER e ads.) 2 sronbeakach aie canes eoikers tiers eye 20
Width tofvhieady fe ptr teeter weit are 14
OGY). 5. Soi oe AM a iene oe ee ee ee 63
Moves ss.1c/Sucies ec ek ce eee 30
Hand glimb tyre hire eco Gece. 51
PN ar, cok cicare woh et okcrae rs icaeta mend eatenee ace 155
Numerous specimens from Carao, 8000 feet, Huaras,
10,000 feet, and Recuay, 11,000 feet.
Distinguished readily from the preceding species by the
granular patches on the sides of the neck.
Tropidurus Thomast.
Upper head-scales smooth ; a series of four to seven trans-
versely enlarged supraoculars ; occipital very large, broader
than long, at least as broad as the supraocular region; nostril
above the canthus rostralis; ear-opening with a fringe of
long pointed scales; temple granulate. A strong curved
antehumeral fold, nearly meeting its fellow on the throat;
latter with more or less. distinet cross folds; sides of neck
minutely granulate, strongly plicate. Body depressed, with
a more or less distinct fold along the side, and a vertebral
series of enlarged tectiform scales, torming a low crest on the
nape; dorsal scales very small, juxtaposed, feebly keeled,
smaller still, granular, and smooth on the sides; ventrals
larger, imbricate, smooth. The hind limb reaches the ear or
between it and the antehumeral fold. Tail about once and a
half as long as head and body, rounded or feebly compressed,
with a low serrated dorsal ridge ; caudal scales much larger
than dorsals, keeled and shortly mucronate. Grey above,
dotted with blackish and yellowish white, the dots having a
tendency to forming transverse series; antehumeral fold
black ; throat and breast black in the males.
millim
Dotalwonoth . -. ciisie nites e an se 6 165
cad rick. s waarmee Boeri ek 15
Waidthsofshesd? nf chy honk nee seers i153}
Body: .S¥ee Ticak Wi Mob chen eer LUes 51
Fore limb tnhe hescsknnctereeeecnes St
Hind dlimb 245... Feces cee ech Ree 45
Tibia gece eee ties aka Pe Ne evateroke 99
Several specimens from Eten, coast of Peru.
Batrachians and Reptiles from Peru. 185
This species, named in honour of my colleague Mr. Oldfield
Thomas, is allied to 7’. peruvianus, Wiegm., from which it is
well distinguished by the shorter hind limbs and the stronger
auricular fringe.
Proctoporus ventrimaculatus.
Body elongate, limbs weak. Fronto-nasal much longer
than broad, much larger than the frontal; fronto-parietals
forming a long suture ; interparietal narrower and a little
shorter than the parietals, widening posteriorly ; three sub-
equal occipitals; three supraoculars; no loreal; a series of
very small infraorbitals ; temple with large irregular shields ;
chin-shields, one anterior and three pairs; gular scales sub-
quadrangular, in 9 transverse series between the chin-shields
and the collar; 10 collar-shields. Dorsal scales elongate-
quadrangular, smooth, juxtaposed; 31 series between the
occiput and the base of the tail. Ventral plates quadrangular,
in 10 longitudimal and 22 transverse series. ‘Two large
preanal shields in the first row, four in the second. Limbs
with smooth shields. Three femoral pores on each side.
Tail thick, scaled like the body. Uniform brown above ;
white beneath, with large black spots.
millim.
JEL TE aro Bit tko De DOC DICer Toes
Wwiruheotheadee seta vere. «occa attic: 5
From end of snout to fore limb........ 12
From end of snout to vent............ 34
ores lime: epee has eats oh Soe aes 9
Elinidilimibier tester, 6 scree: occa losin 28 10
A single specimen from Cajamarca, 10,000 feet.
Philodryas Simonsit.
Hye three fifths length of snout. Rostral broader than
deep, just visible from above; internasals shorter than the
preefrontals ; frontal once and two thirds as long as broad,
longer than its distance from the end of the snout, slightly
shorter than the parietals ; loreal longer than deep ; one pra-
ocular, not reaching the frontal; two postoculars ; temporals
2+3; eight upper labials, fourth and fifth entering the eye ;
four lower labials in contact with the anterior chin-shields,
which are nearly as long as the posterior. Scales smooth,
with single apical pits, in 19 rows. Ventrals rounded, 182 ;
anal divided; subcaudals 105. Greenish yellow above, with
three olive longitudinal stripes and a dark brown vertebral
186 Mr. G. A. Boulenger on new |
line; the lateral stripe extending forwards to the nostril,
passing through the eye; upper lip yellowish white; yellowish
white beneath, speckled with olive.
Total length 780 millim. ; tail 240.
A single specimen from Cajamarca, 9000 feet.
Closely allied to P. elegans, Tsch. Distinguished by the
shorter snout and the shorter frontal shield.
XIX.— Descriptions of new Batrachians and Reptiles from
the Larut Hills, Perak. By G. A. BOULENGER, F.R.S.
I AM indebted to Mr. A. L. Butler, Curator of the Selangor
Museum, for an opportunity of examining a number of
batrachians and reptiles collected by him in the Larut Hills,
among which I was pleased to find examples of several unde-
scribed species. Unless otherwise stated, types of these new
species are preserved both in the British Museum and in the
Selangor Museum at Kuala Lumpur.
Leptobrachium heteropus.
Tongue large, pyriform, feebly notched behind. Vomerine
teeth none. Head moderate, as long as broad; snout short,
truncate at the end; canthus rostralis strong; loreal region
concave; interorbital space as broad as the upper eyelid;
tympanum distinct, half the diameter of the eye. Fingers
moderate, blunt, first and second equal; toes moderate, blunt,
webbed at the base only, the web continued as a slight fringe
along each side of the toes; a strong dermal ridge or keel,
formed by a modification of the subarticular tubercles, runs
along the lower surface of the third and fourth toes, which
thus appear to be compressed; a small oval inner metatarsal
tubercle. The tibio-tarsal articulation reaches the centre of
the eye. Skin smooth, with small tubercles on the upper
eyelids. Grey above, with darker light-edged symmetrical
markings, the largest occupying the middle of the back; a
Jlack lumbar spot; a black canthal and temporal streak ;
black spots on the sides; dark cross-bars on the limbs;
lower parts grey, speckled with black ; a round wlutish spot
on each side of the breast, at the base of the arm, another on
the back of each thigh.
From snout to vent 33 millim.
Batrachians and Reptiles from Perak. 187
A single specimen was obtained in the Larut Hills at an
altitude of 3500 feet. Selangor Museum.
This species is closely allied to L. pelodytoides, Blgr., from
which it may be distinguished by the lesser web and the
extraordinary dermal ridges under the toes, a point of struc-
ture which is only foreshadowed in the types of L. pelodytoides.
Ixvalus larutensis.
Snout rounded or obtusely pointed, as long as the diameter
of the orbit ; canthus rostralis distinct ; loreal region concave ;
nostril a little nearer the end of the snout than the eye; inter-
orbital space as broad as the upper eyelid; tympanum mode-
rately distinct, half the diameter of the eye. Fingers free;
toes half-webbed ; disks of fingers as large as the tympanum ;
subarticular tubercles moderate ; a small inner metatarsal
tubercle. ‘The tibio-tarsal articulation reaches between the
eye and the tip of the snout. Upper parts smooth or with
small flat warts; throat, belly, and lower surface of thighs
granulate. Grey-brown or reddish brown above, with dark
brown symmetrical markings, a cross-band between the eyes
being constant ; usually a )( or )-( shaped marking on the
anterior part of the body ; sides of body and of thighs with
white spots between a brown network; limbs with dark
cross-bands; lower parts white, spotted or speckled with
brown.
From snout to vent 35 millim.
Several specimens from the Larut Hills at 4000 to 4500
feet.
Iealus vermiculatus.
Head large, broader than long; snout rounded, as long as
the diameter of the orbit ; canthus rostralis distinct ; loreal
region concave ; nostril a little nearer the end of the snout
than the eye; interorbital space as broad as the upper eyelid ;
tympanum distinct, two fifths the diameter of the eye.
Fingers with a rudiment of web; toes half-webbed; disks of
fingers as large as the tympanum; snbarticular tubercles
moderate ; a small inner metatarsal tubercle. The tibio-tarsal
articulation reaches between the eye and the tip of the snout.
Upper parts smooth; throat, belly, and lower surface of
thighs granulate. Olive-green above, closely vermiculate
with black; upper surface of thighs with a series of large
black blotches; sides of thighs and anal region orange-
yellow ; white beneath. Male with a large gular vocal sac.
From snout to vent 33 millim.
Three specimens from the Larut Hills at 4000 feet.
188 Mr. G. A. Boulenger on new
Microhyla Butleri.
Habit slender. Snout rounded, as long as the orbit ; inter-
orbital space broader than the upper eyelid. Fingers and
toes rather slender, the tips dilated into small but well deve-
loped disks; first finger much shorter than second; toes
webbed at the base; subarticular tubercles small; two very
small metatarsal tubercles. The tibio-tarsal articulation
reaches the eye. Skin smooth. Grey on the back, pale
reddish on the sides and limbs, with symmetrical dark brown
markings forming bars on the limbs; some small scarlet
spots on the sides; a whitish oblique streak from the eye to
the base of the fore limb and a whitish spot on the end of the
snout; whitish beneath, throat and breast speckled with dark
brown.
From snout to vent 21 millim.
A single specimen from the Larut Hills at 4000 feet. Se-
langor Museum.
Closely allied to M. achatina. Distinguished by the
shorter limbs.
Microhyla annectens.
Habit slender. Snout rounded, as long as the orbit ; inter-
orbital space broader than the upper eyelid. Fingers and
toes moderately slender, the tips dilated into rather large
disks; first finger much shorter than second; toes half-
webbed ; subarticular tubercles feebly prominent; a very
small inner metatarsal tubercle. Hind limb remarkably long,
the tibio-tarsal articulation reaching far beyond the tip of the
snout; tibia two thirds length of head and body. Skin
smooth. Brown above, with symmetrical blackish lght-
edged markings, a large one on the back being produced to
between the eyes, where it expands into a transverse bar ;
side, from the shoulder to the lumbar region, black, with
sharply defined upper outline; an oblique whitish streak from
below the eye to the base of the fore limb; a blackish spot at
the knee, a dark cross-bar on the thigh, another on the femur,
and a third on the tarsus; lower parts closely marbled with
dark brown.
From snout to vent 15 millim.
Several specimens from the Larut Hills at 4000 feet.
This species connects M. achatina with M. Berdmorit.
Gehyra larutensis.
Body and limbs moderately elongate. Head oviform ;
snout a little longer than the distance between the eye and
Batrachians and Reptiles from Perak. 189
the ear-opening, which is small and round; head covered
with finely granular scales, which are larger on the snout ;
rostral twice as broad as deep, with a short median cleft
above; nostril pierced between the rostral, the first upper
labial, and three nasals, the upper of which is the largest and
separated from its fellow behind the rostral by several minute
granules ; 9 upper and 9 lower labials ; symphysial truncate
behind; a series of small chin-shields, the median pair
largest, scarcely longer than the symphysial. Scales uni-
formly granulate on the back, limbs, and throat, larger, flat,
and imbricate on the belly. Digits free, strongly dilated,
with entire lamella, the Jargest of which are angulated or
chevron-shaped ; these chevron-shaped lamella number one
under the hallux, three under the third toe, four under the
fourth. Tail subcylindrical, covered with small imbricate
scales above and beneath. Male with a long continuous
series of 42 femoral and preanal pores. Grey-brown above
and beneath, tail yellowish, with small darker spots on the
back and a vertebral series of small blackish spots widely
separated from each other; a dark line on each side of the
head, passing through the eye.
millim.
TOCA 129d a ee ere eae re 69
1 BIGING! saci aalerc eo ates Rear ae Ee 10
Wide of beadter ss Slate. te Sate ras 6:5
MSO Uhpep de cheat te ee tiettstoee dec Fs ne ote Ue 27
HOVER ad eater. «ach koeiaron: 10
JSlnAGlAihinloy wpe eee ee Ake ee Eo ener ae 13
EWTN) SSOP g Se Wiehe ICE APOE OO EES OR EE 32
A single specimen from the Larut Hills, under a house, at
3500 feet altitude. Selangor Museum.
Draco punctatus.
Head moderate; snout as long as the diameter of the
orbit ; nostril lateral, directed outwards ; tympanum naked,
nearly as large as the eye-opening. Upper head-scales very
unequal, keeled; two subtriangular, compressed, enlarged,
erect scales on the posterior part of the supraciliary region ;
10 or 11 upper labials. The male’s gular appendage a little
shorter than the head. Male with a very distinct nuchal
crest. Dorsal scales unequal, keeled, not larger than ventrals ;
a lateral series of enlarged distant scales. The fore limb
stretched forwards extends a little beyond the tip of the
snout; the hind limb reaches the axilla or between the latter
and the elbow of the adpressed fore limb. T'ail with a dorsal
190 Mr. G. A. Boulenger on new
crest of distinct large, pointed, compressed scales. Dark grey
above, with a paler, reddish, vertebral stripe ; back, and head
above and beneath, with large black dots; wing-membranes
black above, with interrupted whitish streaks, colourless
beneath; throat and belly pale blue; inner side of neck-
lappets and extremity of gular appendage lemon-yellow.
millim
DiptaltenGth oa con on ois sae ate ve one 247
TIGA Tee GG Ee oe feasts siete eek clctals 18
Wadthtotshead’, A.iastt. tims ose kis ecto e 12
WOU Pa kiry ba coms osels sakes ot mei peat 73
Moreslimb tee... Picl anchors wc usheteerme eee 37
FAs rig MAT ssa, 6 eca-5 nuoteclnwikr ipia ak Bis « 45
MAEM ee emacs chs tgs inte is ace ane RG ee 156
A single male specimen obtained in the Larut Hills by
Mr. A. L. Butler at an altitude of 3000 feet. I have also
examined a male from Sarawak, collected by the late Mr. A.
Everett, which I had referred to D. cristatellus.
Draco formosus.
Head small; snout slightly longer than the diameter of the
orbit; nostril directed upwards, perfectly vertical ; tympanum
naked, smaller than the eye-opening; upper head-scales
unequal, keeled ; a prominent tubercle at the posterior corner
of the orbit; nine or ten upper labials. The male’s gular
appendage as long as the head, very thin, translucid, covered
with very large scales. No nuchal fold. Dorsal scales
equal, very feebly keeled, not larger than ventrals; a few
widely separated enlarged scales on the side of the back.
The fore limb stretched forwards extends considerably
beyond the tip of the snout ; the adpressed hind limb reaches
the axil. Brown above (in life), head greyer, with a few
dark spots ; wing-membranes olive above, edged with maroon
or crimson, with five more or less regular black transverse
bands, uncoloured beneath ; throat of male, under the lappets
and right to the base of the gular appendage, maroon or
crimson, of female dark green.
millim. _millim.
Total length .........+54.. 272 P
Hlead) devescn coats a ees Fae ye 19 23
Wadthtotihead! eo grr acter 12 15
Ody fe: cs iscis oh oe bee apes 78 92
orem bilo. mntereitte acres 42 48
ind limb) osha s ieee sicker 53 60
Darl 40j9--crccqebhe ater auc eieiets 175 ?*
* Reproduced.
Batrachians and Reptiles from Perak. 191
Several specimens were obtained in the Larut Hills by
Mr. A. L. Butler at elevations varying between 1500 and
3000 feet.
Intermediate between D. Blanfordii, Blgr., and D. tenio-
pterus, Gthr. D, Blanfordii occurs also in the Larut Hills.
Lygosoma presigne.
Section Hinulia. WUabit lacertiform ; the distance between
the end of the snout and the fore limb is contained once and
a half in the distance between axilla and groin. Snout
short, obtusely pointed. Lower eyelid scaly. Nostril pierced
in a single nasal; no supranasal ; rostral forming a straight
transverse suture with the frontonasal, which is broader than
long; prefrontals forming a median suture; frontal very
narrow behind, as long as frontoparietals and interparietal
together, in contact with the three anterior supraoculars ;
four supraoculars, followed by avery small fifth ; nine or ten
supraciliaries, first and last largest ; frontoparietals and inter-
parietal distinct, latter longer than former ; parietals forming
a very short suture behind the interparietal; three pairs of
nuchals ; fifth upper labial below the centre of the eye. ar-
opening oval, nearly as large as the eye-opening ; no auri-
cular lobules. 28 smooth scales round the middle of the
body; dorsals largest, especially those of the two median
series, which are more than twice as broad as long. A pair
of enlarged preanals. The hind limb stretched forwards
reaches the elbow of the adpressed fore limb. Digits rather
long, compressed ; subdigital lamelle smooth, 20 under the
fourth toe. Reddish brown above, with scattered black dots,
grey on the sides, spotted with black and white; a series of
large roundish black spots on each side of the neck on ante-
rior part of body ; lips spotted with black; tail black above
and on the sides, with irregular annuli of whitish scales;
lower parts white.
millim
MOtAWlEHAG A TOY veut bak Rees vee aes 240
Cer eyed edene hoes. cies sialepeh hepa ahs, ear oe 25
RNa ofsheaad -y.),0%.d)sjoele cee. ss. § Tf,
Core ttt ei teeta, Aono eer 05d) eR Hal gies epe 2 85
rer MUI ees 5 210) vues atses aA s.2 ard ae 0 a0 34
[EPIC aS Artie tia Lactate as-ae. 3 47
Bail, (ROpPOdWCOd) 2 iL ¥ ain chle.e vied claws 130
A single specimen from the Larut Hills at 4000 feet.
Selangor Museum.
192. On new Batrachians and Reptiles from Perak.
Lygosoma stellatum.
Section Henulia. Habit lacertiform; the distance be-
tween the end of the snout and the fore limb is contained
once and one fourth to once and one third in the distance
between axilla and groin. Snout moderate, obtusely pointed.
Lower eyelid scaly. Nostril pierced in a single nasal;
rostral forming a straight transverse suture with the fronto-
nasal, which is a little broader than long; _ prefrontals
uniting or forming a short median suture (in one specimen
separated by a small azygous shield); frontal as long as
frontoparietals and interparietal together, in contact with
the two or three anterior supraoculars; four supraoculars,
followed by a very small fifth ; eight supraciliaries, first and
last largest ; frontoparietals and interparietal distinct, sub-
equal; parietals forming a suture behind the interparietal ;
two or three pairs of nuchals; fifth upper labial below the
centre of the eye. Ear-opening oval, a little smaller than
the eye-opening; no auricular lobules. 24 smooth scales
round the middle of the body, dorsals largest, those of the
two median series more than twice as broad as long. A pair
of enlarged preanals. ‘The hind limb stretched forwards
reaches the wrist or the elbow of the adpressed fore limb.
Digits rather long, compressed ; subdigital lamellae smooth,
22 to 25 under the fourth toe. Bronze-colour above, spotted
all over with black and white, the black spots preceding the
white ones and more crowded on the sides ; lips spotted with
black ; the spots disposed in transverse series on the tail ;
lower parts bluish or greenish white.
millim.
Total lente « Yiswele cit hater asetene 173
Head hel ik- eG eaictctcnh ere eer as pally
Wadthiof head. tic. creer acters 10
Body ski swan woe cake tes Cee eee 63
Hore ambi: S20. SR ee 26
Find dimb’ cow. fine SRE ore ie ee 36
Pag rea oh. ceaie Sales ate een Crees 93
A specimen, badly preserved unfortunately, was brought
home by Mr. 8. 8. Flower, who obtained it in the Larut
Hills, at an altitude of 4400 feet, in Apri! 1898. I have
since examined two younger specimens from the same hills,
forming part of Mr. A. L. Butler’s collection. One was
found in a rotteu tree at 3500 feet altitude, the other in a
house at 4000 feet.
Mr. Butler’s collection also contains an example of Lygo-
soma Bampfyldii, Ki. Bartlett (Journ. Str. Br. As. Soe.
On a new Lizard from Jamaica. 193
no. 28, 1895, p. 96), a species described from the Upper Rejang
River, Sarawak, one of the types of which is now in the
British Museum. ‘This form is remarkable in establishing a
connecting-link between the sections Riopa, Gray, and Lygo-
soma, Gray. It agrees with the former in the presence of
supranasals, forming a suture behind the rostral, with the
latter in the frontal shield being much broader than the supra-
ocular region. In the Sarawak specimen the fifth upper
labial is broken up into several shields, there are 38 scales
round the body, and the upper parts are yellowish brown
with a dark brown band across the frontal region and another
across the occipital. In the Larut specimen the fifth labial is
as large as the fourth and borders the eye, the scales number
40, and the dark brown of the occiput extends along the
dorsal surface of the body and tail, the sides of which are
reddish.
Lycodon Butlert.
Closely allied to Z. fasciatus, Anderson, but with a larger
eye and more strongly angulate ventral and subcaudal shields.
Body slightly compressed. Rostral twice as broad as deep,
hardly visible from above ; internasals three fifths the length
of the preefrontals ; frontal a little longer than broad, as long
as its distance from the end of the snout, shorter than the
parietals; loreal more than twice as long as deep, bordering
the eye below the single preeocular; two postoculars; tem-
porals 2+2; eight upper labials, third, fourth, and fifth
entering the eye; five or six lower labials in contact with the
anterior chin-shields, which are as long as the posterior.
Scales in 17 rows ; dorsals very feebly keeled. Ventrals 224-
228, strongly angulate laterally ; anal entire; subcaudals
88-92 pairs. Blackish brown above and beneath, with 43 or
45 rather irregular annuli of whitish spots or edges to the
scales.
Total length 540 millim. ; tail 115.
‘T'wo female specimens from the Larut Hills at altitudes of
4000 and 5000 feet.
XX.— Description of a new Lizard from Jamaica.
By G. A. BouLencer, F.R.S.
Diploglossus Bakeri.
Lateral teeth obtusely tricuspid. Head small, not distinct
from neck; snout short, with obtuse canthus; ear-opening
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 13
194 On a new Lizard from British East Africa.
moderately large, smaller than the eye-opening; a large
azygous prefrontal, broader than and in contact with the
entire anterior border of the frontal, in contact with the
loreal and separated from the rostral by two pairs of shields ;
frontal once and one third as long as broad; parietal on each
side separated from the frontal and supraoculars by three
shields ; occipital much smaller than the interparietal ; inter-
nasal in contact with the first labial; a single postnasal ;
two consecutive loreals ; rostral twice as broad as the sym-
physial ; the suture between the fifth and sixth upper labials
falls below the centre of the eye; four chin-shields on each
side, the first three in contact with the lower labials. Body
elongate, cylindrical. 39 scales round the middle of the
body, dorsals strongly striated. Limbs very short; the fore
limb stretched forwards extends scarcely beyond the ear; the
hind limb measures one third the distance between axilla and
groin. ‘Tail cylindrical; only the basal scales striated.
Bronzy brown above, with small black spots, which are more
crowded on the sides and limbs; a black dorso-lateral line,
light-edged above, widening on the neck, and extending,
through the eye, to the end of the snout ; belly whitish, throat
speckled with black.
millim
AUG TAL MOM OGM hdetate’s.ctaieca'e stem eter a sare 15]
Flea eas Tis cee Ses Pde ae eis 11
IWirdithyofihieads. sm bes carsteqeie eis his fe wie sate 8
Bod V's ict at sek aki le oem enne ees 57
Fore limb......... Die et sawets Reautovton 10
Mandelimibr tence ee os Fee eke 13
Tal Bee SEAS EAS oe er eet esc 83
A single specimen, a gravid female, was sent from Jamaica
by Mr. C. H. Baker to the Corporation Museum of Leicester,
through the kindness of whose curator, Mr. Montagu Browne,
it has been presented to the British Museum.
XXI1.— Description of a new Lizard from British East
Africa. By G. A. BOULENGER, F.R.S.
Lygosoma clathrotis.
Section Liolepisma. Body elongate, limbs short. The
distance between the end of the snout aud the fore limb is
contained twice in the distance between axilla and groin.
Snout short, rounded. Lower eyelid with an undivided trans-
parent disk. Nostril pierced in the nasal; no supranasal ;
On a new Species of the Genus Alepas. 195
frontonasal broader than long, forming a broad suture with
the rostral and with the frontal; prefrontals small; four
supraoculars ; seven supraciliaries ; frontoparietals distinct,
much larger than the interparietal, behind which the parietals
form a suture; a pair of nuchals; third and fourth upper
labials under the eye. Har-opening small, with six inter-
locking pointed lobules within the meatus, three pointing
downwards and three pointing upwards. 22 subequal smooth
scales round the body. A pair of feebly enlarged preanals.
Fore limb, stretched forwards, reaching the ear; hind limb
half as long as the distance between axilla and groin. Digits
short ; 12 smooth lamella unter the fourth toe. Tail rather
thick, once and two thirds the length of head and body.
Brown above, yellowish beneath ; sides dotted with black.
millim.
SE oar OENR: 5 a creer ac afo) ont + Stn) Smid s) ap alta 167
Header aee as mek ot eae ain ore eran eeay ete Meas ay 11
Wiidthuotneddey + 56) oncta ns roaches attra 7
BS Ye as 1s eke Beco eae. pases ait foltes AID tress F 49
(Hore linn bik pictrerd Ware Bie ete ny share estate nS 10
Drv arrays «8 fer vice vosoty ode os ickos suerte NG 16
Msi eRe eee en coe eka tojeres srtoshe te shee eres fe 1
A single specimen from the foot of Mount Kenia. Pre-
sented to the British Museum by Lord Delamere.
XXII.—On a new Species of the Genus Alepas (A. Lankesteri),
from the Collection of the British Museum. By A.
GRUVEL, Chargé de Cours 4 la Faculté des Sciences de
Bordeaux.
[Plate VIII.]
Draevosis.—Capitulum swollen laterally, covered with a
thick transparent cuticle, without plates. Orifice slightly
tubular and projecting. No crests on the median dorsal
line, but a slight continuous ridge extending from the orifice
to the peduncle.
The peduncle is nearly as long as the capitulum, without
visible ornamentation, with the exception of irregular trans-
verse folds.
Mandibles with four teeth. Inner branch of the 5th and
6th pairs of cirri atrophied and unequal.
Distribution.—West Indies, Mona Channel, 814 fathoms.
Collected by Captain Cole. British Museum Collection,
13*
196 M. A. Gruvel on a new
Capitulum.—The capitulum, entirely devoid of plates, is
covered with a thick chitinous envelope, extremely trans-
parent even after prolonged immersion in alcohol, and orna-
mented with transverse folds which are especially numerous
near the orifice. It is swollen towards the median and
lateral regions, and then strongly retracts to form a kind
of short, tubular duct in which the cirri are set. The
capitulum has no dorsal crests, but simply a slight continuous
ridge which follows the median dorsal line from the opening
of the capitulum to the commencement of the peduncle.
Seen in profile, the general form of the eapitulum is that
of a semicircle, of which the anterior part, forming the base,
is straight, and the posterior part regularly curved.
The dimensions of the capitulum of the largest specimen
are as follows:—Height 20 mm., breadth 18°50 mm., thick-
ness 8°5 mm.
Peduncle.—The peduncle continues the capitulum without
a break. At first broad, it contracts and is almost cylin-
drical towards its middle part; then it broadens again to its
base to form its surface of attachment. The cuticle is the
direct prolongation of that of the capitulum.
This cuticle of the capitulum and peduncle presents some
interesting features. It is composed of chitinous processes
separated by somewhat irregular spaces. Some are wide at
their base, and their summit terminates in three or four
pointed branches which are recurved and divergent, forming
hooks (Pl. VIII. fig. 2); others are shorter and simply conical
(fig. 3). These processes have nearly the same height in
the same zone, and this height varies from 4°8 uw to 24 p.
Near the middle of each of these zones is a sensory bristle
receiving at its base a nerve-filament which is very distinct,
long, slender, and terminates in a fine point. The average
length is from about 95 to 100 wu (fig. 4).
In general each zone of many-pointed hooks is surrounded
by a zone of conical spines, and it is usually also in the zone
of hooks that the sensory bristle is placed.
The opening of the capitulum is heart-shaped, presenting
on the median dorsal line a circular protuberance delimitating
a depression and not a true canal.
Mantle.—The mantle which clothes the chitinous cuticle
on the interior is composed of the usual two epithelial layers,
including between them two muscular layers, the one longi-
tudinal, the other oblique, crossing at various angles and
together forming a kind of very elegant tessellation, these
bundles being separated one trom another by a distance of
about 70 ww. There is, moreover, quite a system of branching
Speeies of the Genus Alepas. 197
elastic fibres, obliquely binding the epithelial surfaces one to
the other.
The chitinous cuticle which clothes the inner part of the
mantle is thin, transparent, and interspersed with comb-
shaped chitinous ornaments with a greater or less number
of teeth, sometimes one only, irregularly placed and serving
to retain the ovigerous sac in the intrapallial cavity.
Mouth.—The labrum has on its free margin fourteen. short,
strong, and blunt chitinous teeth, separated into two series
of seven by a smooth space; the lateral parts are furnished
with stiff and short but fine bristles.
The palpi are flattened and provided with long barbed
bristles over about half their length (fig. 5),
The mandibles have four strong teeth on their free margin.
The distance between the first and second slightly exceeds
that between the points of the second and fourth (fig. 6).
The median dorsal region is covered with short, stiff, but
rather fine bristles ; the lateral faces of the teeth are fur-
nished with strong, short, and pointed bristles, especially well
developed in the vicinity of the last three teeth.
The maxille have the free margin divided into two un-
equal parts by a deep notch. ‘The upper part is furnished
with a very strong chitinous tooth, which continues the dorsal
margin, and a smaller one. The lower part is scalariform
and bears three short and thick projections, between which are
finer bristles, uniformly not barbed, which likewise cover the
lateral parts of the maxilla (fig. 7).
The labial palpi of the lower lip are broad, nearly square
in shape; the anterior free margin is divided by a notch
into two unequal parts, the upper part about three times
as broad as the lower. ‘The upper dorsal region of the
palp is covered with long fine bristles collected in a thick
tuft. These bristles become shorter and stiffer on the lower
parts. Finally, on the posterior part are irregular bristles,
bent and few in number (fig. 8).
Cirrt.—The cirri are generally very long, covered with
long fine bristles, especially long towards the free extremity
of the rami. The first pair is somewhat sharply separated
from the others.
1st pair.—This is much the shortest. The two rami are
unequal. The anterior ramus, formed of 13 rather short
joints (the basal joint being almost equal to four ordinary
joints), bears numerous long, fine, and barbed bristles on its
posterior border; these being much fewer on the anterior
border. ‘The posterior ramus is formed of 21 joints (the
first being nearly equal to four ordinary joints). The hairs
198 On a new Species of the Genus Alepas.
are longer than those of the anterior ramus and the terminal
joints are thinner, while keeping about the same length.
2nd pair.—The rami are nearly equal to each other and
about twice as long as the posterior ramus of the first pair ;
that is to say they attain 12 millim. with 50 to 55 joints, the
joints at the base not being very distinct. These segments
are short, straight, and each bears some long anterior bristles
and some shorter and finer posterior bristles only at the border
of the segments.
The 3rd and 4th pairs are almost exactly similar to the 2nd.
The 5th par has the inner ramus atrophied. The length
of this ramus does not exceed 3°5 to 4 millim. ; the number of
joints is 19, long and slender towards the extremity, bearing
some long fine bristles on the anterior margin and two or
three very short bristles on the posterior margin, at the
border of the segments. The normal ramus is similar to the
preceding.
Finally, the 6th pair of cirri have also the inner ramus
atrophied, similar to the preceding, however, but shorter (16
joints) (fig. 10).
Caudal appendages.—These are long, slender, and cylin-
drical (1°5 millim. in length, with 10 joints). The basal
joints are very broad compared with the superior (fig. 9).
The last joint is furnished with fine bristles longer than itself ;
but these diminish rapidly in length as they approach the
base until they become simple spines.
Filamentary appendages.—A single very short pair at the
base of the first pair of cirri.
Penis.—The penis, which is about 4°5 millim. in length,
is neaily regularly cylindrical, terminating at its extremity
in a blunt point (fig. 11). Itis covered with a thin transparent
cuticle having annular parallel folds in the depressions, be-
tween which are placed, quite irregularly, long fine bristles
more or less bent, which form at the apex a somewhat irregular
tuft.
‘This new species, to which I propose to attach the name
of the Director of the British Museum, approaches in some
of its characters A. cornuta, Darw., and A. japonica, Auri-
villius, but it has most affinity with the first-mentioned
species.
Bordeaux,
7 May, 1900.
EXPLANATION OF PLATE VIII.
Fig. 1. Alepas Lankesteri, sp. n.
Fiy. 2. Hook-shaped processes on the cuticle of the capitulum.
Fig. 3. Conical processes on ditto.
On Two Spiders new to the British Fauna. 199
Fig. 4. Sensory bristles situated in the midst of the preceding structures,
Fig. 5. Left palp.
wg. 6. Right mandible.
Fig. 7. Left maxilla.
Fg. 8. Right labial palp.
Fig. 9. Caudal appendages.
Fg. 10. Cirrus of the 6th pair of cirri.
Fig. 11. Penis (free extremity).
XXIII.—Two Spiders new to the British Fauna. By Grorae
H. CARPENTER, B.Sc. Lond., of the Science and Art
Museum, Dublin.
DurinG the past few years I have received consignments of
spiders for identification from Mr. A. Randell-Jackson, of
Southport, collected by him in various parts of Northern
England and the Isle of Man. Many of his captures are of
considerable faunistic interest, and these will be duly recorded
in local lists. ‘Two species, however, are of particular im-
portance, being undoubtedly additions to the British fauna,
and one of them seems to be new to science. Both were
obtained in the neighbourhood of Southport.
Family Agelenide.
Genus AGELENA, Walck.
Agelena longipes, sp. n. (Figs. 1-5.)
Female.—Length 12°5 millim.; length of carapace 5:5
millim. ; breadth 4 millim. ; length of legs i., il, ii, iv. re-
spectively 20, 18°5, 18, 24 millim.
Eyes of hind row equal to each other; centrals a diameter
apart, each lateral a diameter and a half from its neigh-
bouring central. Front lateral eyes of same size as hind
laterals, which they almost touch, each a diameter from
neighbouring front central. Front centrals larger than the
other eyes, half a diameter from each other, a diameter from
the hind centrals (figs. 2, 3).
Epigyne forming a simple deep rounded depression, broader
than long, with thickened edges; a dark horseshoe-shaped
area in front of it (fig. 4).
Upper spinnerets with the terminal segment flattened and
pointed and fully twice as long as the proximal (fig. 5).
Carapace yellowish brown, with a fine black marginal
line and a broad brown band with black markings on either
200 Mr. G. H. Carpenter on Two
side. Sternum reddish brown, with a black horseshoe-shaped
marking, open in front, drawn out behind, and reaching to
the apex.
Legs remarkably long, the hindmost being nearly twice as
long as the body ; reddish brown, with a few dark markings
on the thighs.
Abdomen dark brown above, with a central clear yellowish
longitudinal band, bounded by a waved black line, and
slightly expanded in front, where it encloses a lance-shaped
red marking with serrate edges (fig. 1). Abdomen beneath
yellowish white, with central black longitudinal band. Spin-
nerets reddish brown, the terminal segment in the upper pair
darker.
Agelena longipes, sp. n.
Fig. 1.—Female (without appendages), dorsal view, x 2.
Fig. 2.— Head-region and eyes, dorsal view, X 7.
Fig, 3.—Eyes, face view, X 7.
Fig. 4.—Epigyne, x 7.
Fig. 5.—Spinnerets from side, x 7.
Locality. Lancashire (Southport).
The specimen described above was taken on a bunch of
flowers which had been brought indoors from a garden. It
is possible therefore that the species may prove to be an
introduced exotic ; but I have failed to find a description of
an Agelena from any part of the world that agrees with this
form.
The genus Agelena has been hitherto represented in our
fauna only by the well-known A. labyrinthica (Clerck) ; it is
by no means impossible that A. longipes will turn out to be a
truly indigenous species.
A, longipes may be distinguished at a glance from A. laby-
rinthica by its smaller size, brighter coloration, and relatively
Spiders new to the British Fauna. 201
Jonger legs and spinnerets, as well as by the altogether diffe-
rent form of the epigyne. The European species to which
A. longipes comes nearest is A. agelenoides (Walck.) *, a
South European spider showing a very similar abdominal
pattern to that of the present species. But the female of
A, agelenoides also has much shorter legs and spinnerets than
A, longipes and a very differently shaped epigyne.
In the last-named character A. opulenta, L. Koch +, from
Japan, shows considerable likeness to A. longipes, but this
species again has relatively short legs and apparently a
unicolorous yellowish-brown abdomen.
On the whole A. longipes seems most nearly related to the
North American A. nevia, Walck. {, which has the legs rela-
tively longer than in the European Agelene. But though
the general structure of the epigyne is similar, none of its
forms as figured by Emerton agrees with that of our spider,
and the American species has no lance-shaped marking on
the abdomen, while the terminal segments of its upper spin-
nerets are relatively short and cylindrical. I have to thank
the Rev. O. P. Cambridge for kindly sending me Canadian
specimens of A, nevia for comparison.
Family Argiopide.
Subfamily Hereonr#.
Genus CNEPHALOCOTES, Simon §.
Cnephalocotes silus (Camb.). (Figs. 6-15.)
Eriyone sila, Camb. Proc. Zool. Soc. 1872, p. 758, pl. Ixy. fig. 7.
Cnephalocotes pusillus, Simon, Arachn. France, v, (1884) pp. 706-7
(nec Microneta pusilla, Menge, Preuss. Spin.).
Cnephalocotes silus, Chyzer and Kulezynski, Aran, Hungar. ii. (1894)
pp. 118-9, pl. iv. fig. 41,
Several specimens of this interesting addition to the British
spider-fauna have been taken on the coast sandhills at South-
port by Mr. Randell-Jackson during the early months of this
year. Full descriptions of the species are given by the
authors referred to in the above synonymy. A few structural
figures may assist British students of spiders to discover
fresh localities for it. Aspects of the male palp somewhat
* Simon, Arachn, France, ii. pp. 115-6.
+ Verh. zool.-bot. Gesell. Wien, xxvii. (1877) pp. 757-9.
{ Emerton, Trans. Conn. Acad. vill. (1890) pp. 197-200, pl. viii.
figs. l-ln. A. californica, Banks (Journ, N. Y. Ent, Soe. iv. (1896)
pp. 89-90), seems a nearly allied form.
§ E. Simon, Arachn, de France, vy, (1884) p. 699; Hist. Nat, Araignées,
2° éd. (1892) tome i. p. 650,
202 Mr. G. H. Carpenter on Two
different from those drawn by former observers are given
(figs. 9-12). The female, fully described by M. Simon, is,
I believe, now figured for the first. time. The epigyne
(fig. 15) is very characteristic, consisting of a simple semi-
circular cavity, with a truncate tongue-like process within its
forward region and a rounded tubercle on either side.
Cnephalocotes silus (Cb.).
Fig. 6.—Carapace of male, with left palp, x 20.
Fig. 7.—Front end of carapace, showing eyes, x 60.
Fig. 8.—Side view of male without appendages, x 20.
Fig. 9.—Left palp of male from side-front, x 40.
Fig. 10.—Genua and tibia of left palp from above, x 40,
Fig. 11.—Left palp from side (spines removed), x 40.
Fig. 12.—Left palp from below, x 40.
Fig. 13.—Outline of female, x 20.
Fig. 14.—Eyes of female from above, x 60.
Fig. 15.—Epigyne, x 40. %
The type of this species (with which Mr. Cambridge has
very kindly compared one of Mr. Randell-Jackson’s speci-
mens, confirming my identification) came from Nuremberg,
Spiders new to the British Fauna. 203
Bavaria. M. Simon records the spider from both the north
and south of France and from Corsica, while. MM. Chyzer
and Kulezynski found it on the Croatian shore of the Adriatic
(Fiume). It seems therefore to belong to a southern distri-
butional type.
This is the fourth species of Cnephalocotes which has been
added to the British list within recent years. ‘The genus was
represented in our fauna only by C. odscurus (Bl) * until
1888, when Mr. Cambridge described as a new species
C. interjectus t from Hertfordshire; this spider, lately
recorded from the Edinburgh district {, is now believed to be
identical with C. lesus, L. Koch §, from Central Siberia. In
1894 two more species of Cnephalocotes—C. curtus, Simon,
and C. elegans, Camb.—were added to the British list ||, the
former occurring on the shores of the Firth of Forth, the
latter in Inverness-shire. C. curtus has since been found on
the west coasts of Scotland and Ireland {]. C. si/us makes,
therefore, the fifth species of the genus known to inhabit our
islands.
The species of Cnephalocotes are small dark-coloured spiders
with strongly chitinized skin and short blunt carapace; eyes
small, those of the hinder row moderately procurved, the
centrals nearer to each other than to the laterals **. The very
wide sternum is produced between the hindmost haunches in
a broad, blunt, rounded process. The legs are short and stout,
the front tarsi being fusiform (especially in the male) and
nearly as long as the metatarsi. ‘The tibia of the male palp
is usually broad and truncate, with one or two short processes ;
the tarsus is always large and the bulb prominent, with a free-
ended, coiled, thread-like spine.
The males of our British species may be tabulated thus :—
I. Head-region more or less elevated, distinct impres-
sions running backward from lateral eyes.
1. Tibia of palp above with an internal blunt and
* Blackwall, Spid. Gt. Brit. Irel. (1864) pp. 297-8, pl. xx. fig. 212.
+ O. P. Cambridge, Trans. Herts. Nat. Hist. Soc. v. (1888) p. 18; Proc.
Dorset Field-Club, x. (1889) pp. 121-2, pl. A. fig. 6.
{ O. P. Cambridge, Proc. Dorset Field-Club, xvii. (1896) p. 60.
§ L. Koch, Kongl. Svensk. Vetensk.-Akad. Handl. xvi. (1878) no. 5,
p- 67, pl. ii. fig. 19. See W. Kulezynski, ‘Fauna Aranearum Austriz
inferioris’’ (Cracow, 1898), pp. 63-4.
|| O. P, Cambridge, Proc. Dorset Field-Club, xv. (1894) p. 112, fig. 4;
G. H. Carpenter and W. Evans, Proc. R. Phys. Soc. Edinb. xii, (1894)
pp. 572-3 ; Ann, Scot. Nat. Hist. 1894, p. 232.
4, G. H. Carpenter, Proc. R. Irish Acad. (8) v. (1898) p. 162.
** In the large and closely allied genus Lophocarenum, Menge, the eyes
of the hind row are equidistant and the row greatly procurved ; also the
skin in Lophocarenum is more strongly coriaceous.
204 Mr. R. Lydekker on an
a central tooth-like process. Cephahe lobe
Gistinet 2. cups ves piteleedeede A ieiedeee AE obscurus (Bl.).
2. Tibia of palp above with a single, short, sharp
process pointing outward. Cephalic lobe
GIStIRCE 2s Shae aes me rene ieee elegans (Cb.).
3. Tibia of palp above with a rather long, sinuous,
sharp-pointed process. Head-region only slightly
Pleveted.: 2. caw ce os epee eaaaee nse apae lesus (L. K.),=
interjectus, Ch.
II. Head-region not elevated ; no impressions behind
lateral eyes.
1. Clypeus strongly conical. 23.00. ..5.4. ede stlus (Cb.).
2. Clypeus vertical; tibia of palp above with a very
short, straight, sharp-pointed process ........ curtus (Simon).
XXIV.—An undescribed Type of Rusine Deer.
By R. LypDeKkKer.
In my work ‘The Deer of All Lands’ a brief notice *, to-
gether with a photograph, was given of three peculiar male
Rusine deer at that time living in the collection of the Duke
of Bedford at Woburn Abbey. They were of small size—a
little larger than a hog-deer—and agreed in general character
with the members of the Sambar group, although differing
from all named forms by the complexity of the antlers. No
name was given to these deer, on account of the possibility of
their proving to be abnormalities or hybrids, or even the
adult of Cervus culionensis. No definite information is
available with regard to their place of origin, although it is
very probable that they came from the Philippines.
One of the three specimens has since died and been presented
by the Duke and Duchess of Bedford to the British Museum,
where its skin is now mounted. A closer examination is now
practicable than was the case during life, and as the result of
this I feel justified in describing the mounted specimen as the
type of a new species of Rusine deer, since it appears different
from any named form, and there seems little probability that
its peculiar characters are due either to abnormality or to
hybridism.
As mounted, the specimen stands 30 inches in height at the
withers. In general form, and especially in the large size of
the face-glands, it agrees with the Rusine group (subgenus
Rusa). From all the various races of the sambar (Cervus
* Page 171, fig. 45.
undescribed Type of Rusine Deer. 205
unicolor) it differs by the hairs being completely annulated
with black and yellow, as it also does by the form of the
antlers and their comparative slight degree of rugosity.
These appendages are primarily of the three-branched Rusine
type, with the inner or hinder tine of the terminal fork
forming the continuation of the beam, and longer than the
outer or front tine. ‘The tail, too, is thinner and less bushy
than in the sambar.
In all these three respects—namely, the annulated hair, the
general form and slight rugosity of the antlers, and the rela-
tively thin tail—the specimen agrees with the rusa (C. Aippe-
laphus). It is, however, very considerably smaller than either
of the three local races of the latter, from which it also differs
by the complexity of the antlers. And here it may be men-
tioned that at the time of its death the animal was apparently
bearing its third pair of antlers, those with which it was
figured in ‘ The Deer of All Lands’ having been shed and
replaced ; it is therefore approximately adult.
As regards their special characters, the antlers are more or
less flattened throughout and display a marked tendency to
palmation. This brow-tine is much flattened, with a sharp
posterior edge, and on the right side is distinctly bifurcate,
although only imperfectly so on the left. The outer tine of
the terminal fork is likewise much flattened, sharp-edged
behind, and trifureate, but the inner tine on the right side is
conical and simple, although showing a tendency to branch
on the left side. ‘The number of points on each antler is thus
SIX.
The species may be shortly defined as allied to QO. hippe-
laphus, but much smaller (30 inches at the shoulder), with
flattened and somewhat palmated antlers, which, when fully
developed, show at least six points on each side. This species
I propose to call Cervus (Rusa) tavistocki, the mounted
example in the British Museum being the type. In giving
this name I must take the risk of the Woburn deer being
identical with one of the numerous forms from the Philip-
pines which have been described by Heude as species ¥.
I may take this opportunity of mentioning that specimens
now at Woburn seem to indicate the identity of Cervus
Luedorfi, Bolau (1880), with C. xanthopygus, Milne-Edwards
(1867). When this year’s antlers are shed the point can be
definitely decided.
* See ‘ Deer of All Lands,’ pp. 186, 187.
206 On Two English Millipedes.
XXV.—On Two English Millipedes (Lulus londinensis, Leach,
and [ulus teutonicus, sp. n.). By R. I. Pocock.
Icrvs tonprvensts was originally described in Trans. Linn.
Soc. Lond. xi. p. 378 (1815), and redescribed and figured in
Zool. Mise. iil. p. 33, fig. 135 (1817). The type and two
other specimens are in the British Museum.
Leach speaks of this species as occurring very commonly
amongst moss in woods near London, but unfortunately does
not say exactly where his specimens were actually collected —
unfortunately, because the species has never, to my knowledge,
been discovered since Leach’s time either near London or in
any other locality at home or abroad. It is true that there is
an allied species, common in some parts of the south of
England and of Western Europe, which passes as londinensis
and has been more than once described under that name by
students of European Millipedes. English specimens of this
species taken in the vicinity of London have been compared
by Dr. Carl Verhoeff with continental examples, and pro-
nounced to be specifically identical with them. A comparison,
however, between examples of this species and Leach’s
| original examples of londinensts shows that the former has
been wrongly determined. It therefore requires a fresh name.
I propose to call it Iwdus tewtonicus, and to select as the type
an example taken by myself at Sevenoaks in Kent.
Careful reading of Leach’s description, brief as it is, of
I. londinensis shows that this species differs from J. teutonzcus
in two important particulars. It is, in the first place, very
much larger, and, in the second place, has the caudal process
submucronate, the caudal process of J. tewtontcus being in no
sense describable as mucronate. This discrepancy was
detected by Verhoeff (Berl. ent. Zeitschr. xxxvi. p. 137, 1891),
who, however, passes it over as due to an error on Leach’s
part. Asa matter of fact, Leach was correct. |
Again, as to size. Leach states that his specimens were
21 inches (that is to say, 58 millim.) long. Meinert (Nat.
Tidssk. v. p. 8, 1868), on the contrary, gives 34 millim. and
Verhoeff 38 millim. as the maximum size of the species they
identified as J. londinensis, neither of them paying heed to
the dimensions given by Leach. In this case, however, Leach
seems to have exaggerated considerably, since all of his
specimens in the British Museum fall short of 2 inches long,
and this is about the length of the specimen represented in
the drawing in the ‘ Zoological Miscellany,’ which purports
to have been taken from life. It is of course possible that
On Two Species of Cyprea. 207
Leach saw larger specimens than those that he placed in his
cabinet. However that may be, there is no question that the
true J. londinensis, judging from the only examples of it that
are known, is a much larger species than the one that has
been mistaken for it on the Continent. Add to this that the
tergal striae are much more numerous and close-set in I. lon
dinensis than in teutonicus, and no one can doubt that the two
are perfectly distinct species. It is safe, moreover, to
prophesy that when fresh examples of I. londinensis come to
hand for examination further differences will be found in the
structure of its copulatory organs.
The differences between the two may be tabulated as
follows :—
a. Total length from about 38 to 48 mm., width 4; tergal
striz yery numerous, fine, and close-set, the intervening
spaces rarely exceeding and generally less than the dia-
meter of the porous area; caudal process short, subcylin-
drical, blunt-pointed or obsolete (submucronate) ...... londinensis.
b. Total length from about 25 to 35 mm., width 2°5; tergal
striz much less numerous and further apart, the inter-
vening spaces generally much exceeding the diameter of
the porous area; caudal process obtusely angular, not
eyem srbmucrOnAter Lh G.I, A. ae SU ieee. oe 5 teutonicus.
I, teutonicus occurs in Scandinavia, Denmark, Western
Germany, the north of France, and the south of England.
The British Museum has specimens from Kent, Middlesex,
Surrey, Hampshire, Oxford, and Warwickshire, but none
from South Wales, Gloucestershire, Somerset, Devon, or
Cornwall, although the Millipedes of these counties have been
fairly well worked.
XXVI.— Descriptions of Two Species of Cyprea, both of the
Subgenus Trivia, Gray. By James Cosmo MELVILL,
MEAs? Lis.
For the opportunity of examining the two cowries now
thought worthy of description I am under much obligation to
Mr. Frederick L. Button, of Oakland, California, a most
enthusiastic cypreologist who has devoted especial attention
to the 7rivie. With much liberality he has from time to
time forwarded me series of species, inhabitants of the Western
American seas, including fusca, californica, and sanguinea,
all of Gray, all three exhibiting much variation, with several
208 Mr. J. C. Melvill on
doubtful forms. Recently the authorities of Stanford Uni-
versity, California, organized a scientific expedition to the
Galapagos Isles, and Mr. Snodgrass collected there one of
the two following, a remarkable shell, on which I have
obtained the opinion of Mr. Sowerby, Mr. Edgar Smith,
Mr. HK. R. Sykes, and others, they all confirming my own and
Mr. Button’s views that it could hardly be referred to any
known species,
Cyprea (Trivia) galapayensis, sp. n.
_ C. (Trivia) testa ovato-rotunda, parva, nigrescenti-purpurea, lateri-
bus paullum dilatatis, extremitatibus vix productis, obtusis, costis
numerosis, crassis, levibus, pallide cinereis, dorsaliter apud
medium superficie callosa omnino obtecta, versus extremitatem
utramque callositatis, velut ocello, albo-maculata, nitidissima, basi
convexiuscula ; apertura arcuato-recta ; columella haud yaricosa.
Long. 8, lat. 5°75 mm. (spec. maj.).
Py ia fama iat Sy », (spec. min.).
Hab. Insula “ Albemarle,” e grege Galapagensi.
This particularly interesting and select form, of which I
have seen three specimens, as just stated, collected for the
Stanford University of California by Mr. Snodgrass, differs
from sanguinea, Gray, in its much smaller size, distinct
coloration, being blackish purple instead of madder-brown,
and more numerous ribs, these being flatter and thicker in
Cyprea galapagensis.
proportion. But the chief peculiarity consists in the shining.
enamelled callosity over the whole centre of the dorsal region,
completely obliterating the sulcus, if any exists, which I.
Two Species of Cyprea. 209
doubt, and further rendered conspicuous by two white spots,
one towards either extremity, at the point of junction of the
ribs with the callosity, and both quite covered by it.
Affinity also exists between galapagensis and pulla, Gask.,
and likewise subrostrata, Gray, both inhabitants also of the
Galapagos group. ‘These small species are much of the same
size, the latter possessing a decided sulcus, and with beaked
extremities—hence its trivial name; whilst pudla is less
globose, having fine ribs, with hardly any definition even of
a sulcus.
No trace of any dorsal callosity has, so far as I am aware,
ever been found in any other 7rivéa such as exists in the
species before us.
Cyprea (Trivia) Buttont, sp. n.
C. (Trivia) testa parva, ovato-globosa, pallide straminea, apud
latera pauilum dilatata, costis ad quatuordecim, fortibus, con-
tinuis, hie illic interruptis vel subdivaricatis, ad dorsum precipue
latis, levibus, albis, sulco nullo, apnd extremitates paullum pro-
ducta, basi subconvexa, dentibus labialibus ad octodecim.
Long. 5°50, lat. 4 mm.
Hab. 2? (Ff. L. Button, Esq.).
A small, globular, straw-coloured Trivia, few ribbed, these
being continuous, occasionally interrupted or subdivaricate,
broad, especially dorsally ; there is no sulcus present; the
shell is slightly produced at the extremities; labial teeth
eighteen in number.
Cyprea Buttoni.
But few species are very comparable with this: acutidentata,
Gask., may be akin, but the type is lost, and I have never
seen anything but the original insufficient description ; pauc?=
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 14
210 On a new Species of Papilio.
lirata, Sowb., possesses a well-defined sulcus, and the ribs
seem more acute than are those of Buttont; candidula, Gask.,
is larger and whiter, with more frequent coste ; producta,
Gask., as its name implies, is produced at its extremities, and
the ribs are of a different character, very acute and thin.
There is no sulcus, however, in this species. The ribs, like-
wise, of pellucitdula, Gask., are far more numerous, and the
substance more delicate than in our shell, which it affords me
sincere pleasure to be able to dedicate to Mr. F’. L. Button,
its discoverer.
XX VII.—Desecription of a new Species of Papilio from Bwool,
North Celebes. By H. Grosu-Smitu, F.E.S., F.Z.8., &c.
Papilio Dixont.
2 .—Upperside. Anterior wings rather pale fuliginous
brown, darker at the base and the costal and outer margins ;
the dark areas more restricted than in the same sex of
P. Kiihni, Honrath, the veins and rays in the cell and be-
tween the veins also dark fuliginous brown. Posterior wings
pale fuliginous brown, with the base and outer third darker ;
the crimson markings on the underside showing indistinctly
through the wings.
Underside, Anterior wings as above. Posterior wings
brownish black, with a pale crimson irregular band a
little beyond the cell, extending obliquely from near the
abdominal margin to the upper median nervule, much nar-
rower than the crimson band in a similar position on the
underside of the posterior wings of P. Kiihni; outside this
band crossing the disk, between the veins, is a row of four
large pale crimson lunules, of which the first and fourth are
the narrowest.
Expanse of wings 44 inches.
In the collection of Mr. Grose-Smith.
Nearest to the female of P. Kiihni, which it resembles in
shape. This butterfly was captured by Mr. Frank Dixon
about 80 miles inland at an elevation of 800 feet.
Mr. W. F. Kirby on African Phasgonuride. 211
XX VIII.—Notes on the Collection of African Phasgonuride
formed by Mr. W. L. Distant in the Transvaal &e., with
Descriptions of Two new Species. By W. EF. Kirsy,
F.L.S., F.E.S., &e.
Tuis collection (exclusive of the families Stenopelmatide,
Gryllacridz, and Hetrodide, which were discussed in the
‘Annals’ for June 1899, pp. 475-480) includes the following
twenty species, belonging to various families. ‘Two species
of the genus T'ylopsds are described as new.
ORTHOPTERA-PHASGONURIDA.
Decticide.
GAMPSOCLEINE.
Arytropteris, Herm.
basalis, Walk.
Sagide.
Clonia, Stal.
vittata, Thunb.
Wahlbergi, Stal.
Conocephalide.
CoNOCEPHALIN2,
Pseudorhynchus, Serv.
pungens, Schaum.
Conocephalus, Serv.
consobrinus, Walk.
tuberculatus, Rossi.
XIPHIDIINZ,
Aiphidium, Serv.
aris, Serv.
Pseudophyllide.
PsEUDOPHYLLINA,
Zabalius, Bol.
orientalis, Karsch,
Bocaget, Bol.
CYMATOMERINZ.
Cymatomera, Schaum.
denticollis, Schaum.
spulophera, Walk.
Mecopodidz.
Anedepoda, Karsch.
latipennis, Burm.
Phaneropteride.
ACROMETOPINZE,
Rhegmatopoda, Brunn.
Brunneri, n. a.
PHANEROPTERINA.
Phaneroptera, Serv.
nana, Charp.
TYLOPSIN#,
Tylopsis, Fieb.
continua, Walk.
marginata, Brunn,
punctulata, sp. n.
ruUbrescens, SP. 1.
AMBLYCORYPHINE,
Amblycorypha, Stal.
cereris, Stal.
proserpineé, Brunn.
14*
212 Mr. W. F. Kirby on African Phasgonuride.
Decticide.
G AMPSOCLEINZ.
Arytropteris basalis.
Thyrconotus basalis, Walk. Cat. Derm. Salt. ii. p. 247. n. 6 (1869).
Arvtropteris angulosa, Herman, Verh. zool.-bot. Ges, Wien, xliv. p. 204
(1874).
1, Zomba (P. Rendall).
Natal (B. M.); Zululand (Herman).
Thorancistus, Pictet, appears to me to be a different genus.
Sagide.
Clonia vittata.
Locusta vittata, Thunb. Nov. Ins. Spec. v. p. 102 (1789) ; Mém. Acad.
Pétersb. v. p. 280 (1815).
Clonia vittata, Stal, Rec. Orth. p. 119 (1874).
Saga maculosa, Walk. Cat. Derm. Salt. ii. p. 294. n. 16 (1869).
1, Pretoria (Distant).
Between Olyfants River and Slang River (Thunberg).
Clonia Wahlbergi.
Clonia Wahlberg’, St81, Ree. Orth. 11. p. 119 (1874); Dist. Nat. in
Transvaal, p. 83, cum fig. (1892).
1, Transvaal, Waterberg (W. Z. D.); 2, Zomba (P. Ren-
dall) ; 2, Barberton (P. Rendall).
Natal (Stal).
Conocephalida.
ConocEPHALIN#
Pseudorhynchus pungens.
Conocephalus pungens, Schaum, Monatsb. Akad. Berl. 1853, p. 778;
Peters, Reise nach Mossamb. v. p. 126, pl. vii. tig. 12 (1862).
Pseudorhynchus pungens, Redt. Verh. zool.-bot. Ges. Wien, xli. p. 365,
pl. iii. tig. 18 (1891).
2, Fort Johnsion (P. Rendall).
Zanzibar, Mozambique (Ledtenbacher).
Conocephalus consobrinus.
Conocephalus consobrinus, Walk. Cat. Derm. Salt. ii. p. 315. n. 4 (1869),
9, Pretoria (W. L. D.); 1, Barberton (P. Rendall),
Natal (B. M.).
Mr. W. F. Kirby on African Phasgonuride. 213
Conocephalus tuberculatus.
Locusta tuberenlata, Rossi, Faun. Ftv. i. p. 269 (1790).
Ineusta mandibularis, Charp. Hor, Ent. p. 106 (1825).
Conocephalus mandibularis, Brann, Prodr, Eur. Orth. p. 304 (1882) ;
Redt, Verh. zvol.-bot. Ges. Wien, xii. p. 427. n. 101 (1891).
1, Fort Johnston (P. Rendall).
Comnion in South Europe and almost throughout Africa,
A IPHIDIINE:,
Xiphidium iris.
Aiphidion iris, Serv. Ins. Orth. p. 506 (1839).
Xiphidium iris, Redt, Verh. zool.-bot. Ges. Wien, xli. p. 515 (1891).
Xiphidium punctipenne, Walk. Cat. Derm. Salt. ii. p. 272. n, 14 (1869).
Aiphidium tenue, Walk. d. c. n. 15 (1891).
1, Pretoria (W. ZL. D.); 1, Fort Johnston, Nyasaland
(2 Ttendali).
Mauritius, Rodriguez, Madagascar, Gaboon, Zanzibar
(Redt.); S. Africa (B. M.).
Pseudophyllida.
PsEv DOPHYLLINE.
Zabalius orientalis.
Mateus orientalis, Karsch, Berl. ent. Zeitschr. xxxvi. p. 85 (1890);
Brunn. Mon. Pseud. p. 30 (1895).
1, Barberton (P. Rendalf).
Usambara, Tanganyika (Berlin) Museum); Zanzibar
(Dohrn) ; Bihe (?) (Bolivar).
The descriptions of this species are very poor, Mr. Dis-
tant’s specimen has broad oblique yellow lines, with slight
pseudopodiform projections, and narrowly bordered on the
outside with black. In fact it much resembles Z. Boerger,
but the thoracic granules are less numerous, and are yellow in
front, and reddish behind instead of black.
Zabalius Bocaget.
Mustius Bocaget, Bol. Jorn. Sci. Lisb. (2) i. p. 221 (1899),
1, Angola (Monteiro).
The Natural History Museum possesses a discoloured
specimen from the Congo which agrees with Bolivar’s deserip-
tion in having the inside of the hind femora blood-red.
Mr. Distant’s specimen is only slightly faded on the head &e.,
and is pale green, with the thoracic granules black, ‘The
214 Mr, W. F. Kirby on African Phasgonuride. |
radial nervure of the tegmina is orange at the base, as well
as the front border of the pronotum; the transverse nervures
of the tegmina are broadly yellow, narrowly edged behind
with black, and the inner marginal area is brownish yellow
except at the base, but may be slightly discoloured. ‘The
hind femora are green, with a row of reddish-brown spots on
the inner side, and a double row of short stout yellow spines
beneath, most numerous on the outer carina. The tips of the
femora and the whole of the tibie and tarsi are reddish. In
the faded Museum specimen the pale transverse Jines of the
tegmina are bordered outside with pink.
Until a larger series of fresh specimens is received, I
should not be justified in separating Mr, Distant’s specimen
as a distinct species.
CYMATOMERINA.
Cymatomera denticollis.
Cymatomera denticollis, Schaum, Monatsh, Akad. Berl. 1853, p. 778;
Peters, Reise nach Mossamb. v. p. 123, pl. vii. fig. 9 (1862) ; Karsch,
Berl. ent. Zeitschr. xxxvi, p. 97 (1891); Brunn. Mon. Pseud. p. 83
(1895), :
Cymatomera Schaum, Stal, ify. Vet.-Akad. Forh, xiii. p. 170 (1856).
2, Barberton (P. Rendall).
Mozambique (Schaum); Delagoa Bay, Tanganyika
(Karsch) ; Zambesi (Brunner) ; Natal and Nyasa (B. M.).
Cymatomera spilophora.
Cymatomera spilophora, Walk. Cat. Derm, Salt. iii. p. 455. n. 3 (1870).
Cymatomera Brancsikt, Brunn. Mon. Pseud. p. 86, pl. iv. fig. 34 (1895).
1, Barberton, Zomba (P. Rendall).
Zambesi (runner); Bangani, German East Africa
(Berlin Museum) ; East Africa (type) ; Mombasa, Nyasa
(B.. Mas
Mecopodide.
|needopoda latipennis.
Mecopoda latipennis, Burm. Handb. Ent. ii. p. 686, n. 2 (1839).
1, Zomba (P. Rendall).
Natal, Nyasa, Uganda, Sierra Leoue (B. M.),
Mr. W. F. Kirby on African Phasgonuride. 215
Phaneropterid2.
ACROMETOPINZ.
Ithegmatopoda Brunneri, n. n.
|| Zoratosphaga leptocerca, Brunn. Mon. Phan, p, 89. n. 38, pl. i. figs. 9 a, b
(1878).
Rhegmatopoda leptocerca, Brunn, Verh. zool.-bot. Ges, Wien, xli. p. 45
(i891).
1, Barberton (P. Rendall).
This cannot be Horatosphaga leptocerca, Stal (CErv. Vet.-
Akad. Férh. xxxiii. p. 59, 1876), in which the subgenital
plate of the male is described as narrow.
PHANEROPTERINE.
Phaneroptera nana.
Phaneroptera nana, Charp. Fieber, Lotos, ii. p. 49 (1853); Brunn.
Mon. Phan. p. 215 (1878).
||Phaneroptera bilineolata, pt., Walk. Cat. Derm. Salt. ii. p. 337. n. 16
(1869).
Phaneroptera sparsa, Stal, difv. Vet.-Akad. Forh. xiii. p. 170 (1858).
Phaneroptera conspersa, Stal, Rec. Orth. ii. p. 29 (1874).
Phaneroptera tetrasticta, Gerst. Arch. f. Nat. xxxv. p. 215 (1869) ;
Von der Decken’s Reisen, iii. (2) p. 832 (1873).
1, Pretoria (W. L. D.); 4, Barberton (P. Rendall); 12,
Fort Johnston, Nyasaland,
Portugal, Fernando Po, Cape, Uru, Zanzibar, Rio Janeiro
(Brunner) ; Natal (B. M.).
TYLOPSINe.
Tylopsis continua.
Phaneroptera continua, Walk. Cat. Derm. Salt. ii. p. 837. n, 20 (1869).
Phaneroptera vicaria, Walk. l. c. p. 388. n, 22 (1869).
|| Phaneroptera bilineolata, pt., Walk. 7. c. p. 837. n. 16 (1869).
Tylopsis longipennis, Stal, Gify. Vet.-Akad. Forh. xxxili. (5) p. 58
(1876).
Tylopsis vittata, Brunn. Mon. Phan. p. 229 (1878).
Tylopsis inhamata, Karsch, Berl. ent. Zeitschr. xxxii. p. 455 (1888).
1, Fort Johnston, Nyasaland (P. Rendall) ; 2, Masil Nek ;
i Pretoria CW. 2...) )
Damaraland (Sta/) ; Cape (Berlin Museum) ; Delagoa Bay
(Karsch); Natal, Zululand (B. M.).
I consider all the above names to refer to a single species.
216 Mr. W. F. Kirby on African Phasgonuride.
Tylopsis marginata.
Tylonsis marginata, Bruyn. Verh. zool.-bot. Ges. Wien, sli. p. 113
(1891).
1, Pretoria (W. LZ. D.); 2, Barberton (P. Rendall); 1,
Zomba (P. Rendall) ; Natal (Brunner).
Apparently a rather scarce species,
Tylopsis punctulata, sp. n.
3.—Long. corp. 18 millim., cum app. 21 millim. ; long.
teom. 31 millim. ; long. al. 38 millim.
@ .—Long. corp. 20 millim., cum ovip. 24 millim.; long.
teem. 32 millim. ; long. al. 38 millim.; exp. tegm. 65 millim. ;
exp. al. 69 millim.
Brownish testaceous, probably green when living; an-
tenne reddish brown; a broad brown band runs from the
occiput over the pronotum and the inner margin of the teg-
mina, extending over the basal area of the latter, for about one
third of their length, where it narrows to a border for the rest
of their length. Pronotum with a narrow yellow lateral line
on each side of the brown band; the deflexed lobes are pale, a
little longer than high, with both the lower angles rather ob-
tusely rounded off; they are also marked with a rather large
brownish blotch in the middle. ‘Tegmina with rows of sub-
obsolete small brown spots between the longitudinal and
oblique nervures. Abdomen not denticulated on the median
line; the last two segments with black spots in the male;
subgenital lamina broad, upeurved, bifid; cerci curved round
it, inwards and upwards, and waved towards the tips, which
are attenuated and pointed. Female with the ovipositor
short, broad, rugose-punctate, suddenly upcurved, and with a
short rather abrupt point at the tip. Legs (especially the
hind legs) very long and slender; femora unarmed ; front
and middle tibize sulcated, finely spinulose above and below ;
hind tibie closely and thickly set with short black-tipped
spines on both carinz above ; below there is a row of spines
placed more widely apart on the inner carina only.
3, Zomba (P. Rendall).
Distinguished from every other species by the dark spots on
the tegmina.
Tylopsis rubrescens, sp. n.
Long. corp. 16 millim., cum app. 20 millim.; long. tegm.
29 millim.; long. al. 35 millim,
Mr. W. F. Kirby on African Phaszgonuride. 217
Male.—Very similar to the last species, evidently green
when alive, but except on the tegmina the colour in the only
specimen before me has faded to yellowish. Antenna an
legs red. A broad reddish-brown band runs from the occiput
over the pronotum and inner margin of the tegmina, as in
the last species ; on the pronotum it is darkest on the sides,
the middle being marked by an obsolete pale line. ‘The
deflexed lobes are pale, bordered with a pale line (probably
yellow in life) above and below. ‘The lower mouth-parts
are black. ‘The dark edging of the inner margin of the
tegmina is broadly bordered with reddish, shading into
purplish brown beyond the middle, the colonr being con-
tinued on the exposed part of the wings. There are a few
small dark spots between the oblique nervures, but not more
than one or two in each interspace. Front tibia apparently
unarmed above ; middle tibie with only a single spine visible
above. Abdomen red above, with a central carina, but appa-
rently not denticulated; slightly dusted with red on the
sides. Cerci red, rather shorter than in 7. punctu/ata.
1, Zomba (P. Rendall).
Apparently intermediate between 7. marginata, Brunner,
and 7’, punctulata.
A MBLYCORYPHINE.
Eurycorypha cereris.
Phylloptera cereris, Stal, GEfv. Vet.-Akad. Forh. xiii. p. 170 (1856).
Eurycorypha cereris, Stal, Ree. Orth. ii. p. 39 (1874); Brunner, Mon.
Pian. p. 273 (1878).
|| Phyllopterst proteifolia, Walk. (nec Burm.) Cat. Derm. Salt. ii. p. 378.
n. 11 (1869).
1, Figtree Creek, Barberton (P. Rendail).
Caffraria (Sta/) ; Grahamstown (Brunner) ; Natal (B. M.).
Eurycorypha proserpine.
Eurycorypha proserpine, Brunn. Mon. Phan. p. 274, pl. vi. fig. 83
(1878).
|| Orophus gramineus (?), Walk. (nec Serv.) Cat. Derm. Salt. iii. p. 434
(1870).
2, Barberton (P. Rendall).
Natal (B. M.).
218 On the Genital Pouches in Thalassema neptuni.
XXIX.—Noteona Variation in the Number of Genital Pouches
in Thalassema neptuni, Gaertner. By F. H. Stewart,
M.A., Gatty Marine Laboratory, St. Andrews.
WHILE dissecting a specimenof Thalassema neptun?, Gaertner,
I observed that it varied from the normal in possessing five
genital pouches instead of four, one being placed in the
middle dorsal line, the remaining four lying, as usual, two on
each side. But on examination by means of sections the
pouch which appeared to be median proved to belong to the
right side and to correspond to the anterior pouch on the left.
Behind these on the right side came an unpaired nephridium,
followed by a second normal pair.
The question then arose, In what relations did the five
nephridia of this specimen stand to the normal four? Was
the unpaired pouch interpolated between the normal two
pairs, or did it represent the second normal pair, the fourth
and fifth pouches being an additional pair? It was obviously
impossible to obtain any direct proof on this point; but in
order to obtain some indication I measured the distances
between the successive nephridiopores in the abnormal and
in several normal specimens, at the same time taking the
distance between the genital hooks and the first pair of
nephridiopores as a standard of comparison. In the abnormal
specimen the distances were as tollows:—(a) Genital hooks
to first pair of pores ‘5 millim. (8) First pair to unpaired
*25 millim. (c) First pair to second pair ‘7 millim. The
relation of distance (a) to distance (0) is thus 2: 1, while that
of (a) to (c) is 5:7.
In the normal specimens the relation of the averages of the
distances were :—Distance of genital hooks from first pair of
pores to distance between two pairs as 2 to 3, 7. e. '5 to *75.
"25
x arto 0
Abnormal.
x 0
SS - —/\———~- I
5 Tf
x 0
Normal (mean),
x 0
5 15
(xX =genital hooks; 0=nephridiopores.)
On a new Species of Buprestide. 219
This point is illustrated by the accompanying diagram.
It would appear from this comparison that the unpaired
genital pouch is interpolated between the two normal pairs ;
and if this be granted it would have to be assumed either that
a segment which does not normally bear pouches intervenes
between the two which normally do, or that a partial dupli-
cation of the nephridia of one segment has oceurred—a pheno-
menon which we find in its complete form in the family of
the Capitellidee.
Mr. Punnett, Assistant Professor of Natural History in the
University of St. Andrews, who kindly gave me the abnormal
and the various normal specimens, informs me that they were
all obtained near Plymouth in the same locality. There can
thus be no doubt that this case is an actual variation, not a
local variety.
XXX.—Description of a new Species of Buprestide.
By Cuas. O. WatTEeRHOUSE, V.P.E.S.
SPECIMENS of the species described below have been in the
Museum collection for many years, separated as distinct from
Psiloptera quadrioculata, A fresh specimen just brought
from Upper Egypt by Mr. D. A. MacAlister shows that the
differences between this and P. guadritoculata are constant,
and I| therefore venture to give it a name, and I propose to
eall it P. MacAlister? after the donor.
Psiloptera Mac Alistert, sp. n.
P. quadrioculate valde affinis ; vitta obliqua thoracis, elytrorumque
vitta laterali cupreo-rubris; corpore subtus tomento griseo-albo
induto, utrinque maculis majoribus circularibus ornato.
Long. 20-27 mill.
Very similar to P. guadrtoculata, but perhaps a little less
convex. ‘lhe general blackish-wneous colour is the same,
but the impressions on the thorax and lateral stripe of the
elytra are coppery red. ‘The front of the head is clothed with
pale yellow pile. ‘he thorax is moderately closely and
strongly punctured, with a slight raised median line, and
four rotundate, smooth, black spots as in P. quadrioculata,
but the rugose surface surrounding them is coppery red and
forms a distinct oblique band from the anterior spot to the
base, leaving a triangular space at the posterior angle slightly
220 Mr. W. L. Distant on Dinidorine,
raised, and smoother than in P. guadrioculata. The sculp-
ture of the elytra is nearly the same, but the stria are a little
more strongly impressed, and, being filled (in fresh examples)
with a sandy-white pile, are more conspicuous. ‘The inter-
stices are broken up into irregular quadrate spaces by rugose
punctures; the smooth raised parts are more convex than in
PP, quadriocufata. The underside of the body is clothed
(except along the median line) with whitish (or sandy
coloured) pile; the first to fourth seements have each a
round smooth spot on each side, and the terminal segment
has an oblique smooth line.
Hab. Upper Egypt, Northern Etbai (D. A. MacAlister) ;
White Nile (Consul Petherick); Nubia; Suez.
The specimens from Nubia and Suez have the pubescence
on the underside of the body sandy coloured.
XX XI.—Rhynchotal Notes—VI. Heteroptera: Dinidorine,
Phyllocephaline, Urolabidine, and Acanthosomine. By
W. L. Distant.
THIS contribution completes the examination of the family
Pentatomida, as contained in the British Museum, including
the genera and species described by Walker in vols. 1.—i1t
of his ‘Catalogue of Hemiptera-lHeteroptera’ (1867-8).
Some recent acquisitions to the Museum and my own collec-
tion are also described.
Drx1porivz.
Genus CYCLOPELTA.
Cyclopelta dorsalis.
Cyclopelia dorsalis, Walk. Cat. Tet. iii. p. 478. n. 5 (1868).
Allied to C. funebris, Fabr., but not separable by the
differential character given by Walker—‘‘ somewhat broader
than C. funebris”—but by having the lateral margins of the
pronotum much more oblique and less rounded than in the
Fabrician species.
Cyclopelta parva, sp. n.
Closely allied to C. obscura, Lep. & Serv., but differing by
its much smaller size and by the antenne, which are shorter
and broader than in that species.
Long. 10-12 millim.
Phyllocephaline, Urolabidinz, and Acanthosomine. 221
Hab. China, Kiukiang (Pratt, Brit. Mus.) ; Shantung
(Coll. Dist.) ; Rangoon and Pegu (Atkins. Coll., Brit. Mus.).
I have long possessed a single specimen from Shantung ;
and as the British Museum now contains six other specimens
from Kiukiang, I have not hesitated to separate them, as
constituting a distinet species. All the general characters are
those of C. obscura. ‘The connexivum is generally spotted
with ochraceous and there is usually a small central basal
ochraceous spot to the scutellum. ‘The antenna are shorter
and the joints broader and more spatulate than in C. obscura.
Cyclopelta? vilis.
Cyclopelta vilis, Walk. Cat. Het. iii. p. 478. n. 6 (1868).
The typical and only specimen of this species which I
have seen possesses merely the first, second, and third joints
of one antenna. It may probably prove to be a species of the
genus Aspongopus.
Genus ASPONGOPUS.
Aspongopus fuscus.
Asponyopus fuscus, Westw. in Hope Cat. i. p. 26 (1887).
Aspongopus margimalis, Dallas, List Hem. i. p. 350. n. 9 (1851).
A spongopus cuprifer.
Aspongopus cuprifer, Westw. in Hope Cat. i. p. 25 (1837).
Asponyopus sepuichralis, Stal, Hem. Afr. i. p. 214. n. 5 (1864).
Aspongopus solitus, Walk. Cat. Het. iii. p. 484. n. 30 (1868).
Aspongopus patruelis.
Cyclopelta patruelis, Stal, CEfv. Vet.-Ak. Foérh. 1853, p. 233. n. 1.
Cyclopelia dotata, Walk. Cat. Het. iii. p. 479. n. 7 (1868),
Aspongopus binotatus, sp. n.
Above dark castaneous ; apical joint of antennez and two
small spots at apex of scutellum luteous; base of lateral
margins of corium and body beneath ochraceous ; legs dark
casfaneous.
Second joint of antennae minute, third much longer than
fourth. Pronotum obscurely transversely rugulose ; scutellam
distinctly wrinkled; corium very obscurely tomentose ; femora
with their inner margins spinous.
Long. 17-19 millim. ; max, abd. lat. 11-12 millim.
Hab. Nyasaland (Sir H. Johnston, Brit. Mus.).
By the minute second joint of the antennz this species is
allied to A. patruelis, Stal. |
222 Mr. W. L. Distant on Dinidorine,
Aspongopus figlinus, sp. n.
Testaceous, somewhat paler beneath ; apical joint of the
antenne luteous.
Antenne with the second joint minute, third not quite twice
the length of fourth and subequal to fifth, third and fourth
joints flattened and sulcate. Head with the lateral lobes
convex at their apices, between which the anterior margin is
distinctly cleft ; the lateral margins sinuate. Pronotum with
the lateral margins convex and slightly reflexed, its surface
finely punctate and obscurely rugulose, especially near base.
Scutellum transversely wrinkled and finely punctate. Corium
obscurely wrinkled and finely punctate. Membrane dark
ochraceous. Coxe and disk of abdomen ochraceous.
Long. 17-18 millim.; exp. pronot. angl. 10-11 millim. ;
max. abd. lat. 12-124 millim.
Hab. EK. Africa: Livingstonia (Simons, Coll. Dist.) ;
Nyasaland ; Fort Johnston (P. Rendall, Coll. Dist.).
Allied by structure to the group of species having the
second joint of the antenne minute, viz. A. patruelis, Stal,
A. cyclopeltus, Dist., and A. binotatus, Dist.
Aspongopus singhalanus, sp. n.
Bronzy brown ; antenne, eyes, rostrum, and legs piceous ;
apical joint of antenne, base of rostrum, and the tarsi
ochraceous.
Antenne thick ; the second, third, and fourth joints deeply
suleate; second and third joints subequal in length or third
a little longer than second ; fourth and fifth joints subequal,
fifth joint narrowest and cylindrical. Body above finely
rugulose and punctate; abdomen above reddish. Rostrum
reaching about halfway between the anterior and intermediate
COX.
Long. 18 millim.; exp. pronot. ang]. 10 millim.
Hab. Ceylon (Green, Brit. Mus. and Coll. Kirkaldy ; Lewis,
Coll. Dist.).
Allied to A. brunneus, Thunb., and A. obscurus, Fabr.,
from both of which it differs by the thick and deeply sulcate
antenne.
Genus MEGYMENUM.
Megymenum dentatum.
Megymenum dentatum, Boisd. Voy. Astr., Ins, ii. p. 632, pl. xi. fig. 11
1835).
Merion instructum, Walk. Cat. Het. iii. p. 502, n. 14 (1868).
Walker had confused JZ. dentatum, Boisd., with MZ. semé-
vestitum, Voll., and then redescribed Boisduval’s species.
Phyllocephaline, Urolabidine, and Acanthosomine. 223
PHYLLOCEPHALINA.
Genus DALSIRA.
Dalsira humeralis,
Phyllocephala humeralis, Walk. Cat. Het. iii, p. 490. n, 22 (1868).
Basicryptus? humeralis, Leth. & Sev. Cat. Gén. Hém. t. i. p. 242
(1893).
Dalsira vicina.
Phyllocephala vicina, Sign. Rey. et Mag. Zool. 1851, p. 446. n. 13.
Phyllocephala funesta, Walk. Cat. Het. iii. p. 490. n. 23 (1868).
Genus BASICRYPTUS.
Basicryptus distinctus.
Phyllocephala distincta, Sign. Rey. et Mag. Zool. 1851, p. 446.
Phyllocephala impressa, Walk. Cat. Het. ii. p. 489. n. 21 (1868).
Walker’s type is a small specimen of Signoret’s species.
The series now before me varies in length from 17 to 22 millim.
Basicryptus diversus, sp. n.
Brownish ochraceous; inner lateral margins, a central
anterior spot and a central transverse line to pronotum, a
broad central longitudinal fascia to scutellum, inner margins,
and a submarginal fascia to corium black. The divergent
lateral lobes of head shorter than in preceding species; a
distinct spine in front of eyes; antenne mutilated. Pro-
notum moderately convex, the lateral angles subprominent,
the lateral margins acutely dentate ; two transverse levigate
lines on disk, posterior half rugose and punctate. Scutellum
with the basal and lateral margins and margins of the central
black fascia raised and levigate; centrally thickly punctate,
outwardly sparsely so. Corium sparsely punctate, membrane
ochraceous. Sternum, rostrum, and legs ochraceous; abdo-
men beneath brownish ochraceous; stigmatal spots and a
prominent spot on lateral areas of prosternum piceous.
Long. 15 millim.; exp. pronot. angl. 84 millim.
Hab. Australia; Swan River (Coll. Dist.).
Allied to B. rugicollis, Westw., and Bb. Frenchi, Bergr.,
but differing from both by the strongly serrate lateral margin
to the pronotum, the non- or subprominent pronotal lateral
angles, &c.
224 Mr. W. L. Distant on Dinidorine,
Basicryptus negus, sp. n.
Ochraceous, unicolorous; membrane greyish, with the
nervures fuscous.
Antenne pale Juteous; second, third, and fourth joints
almost subequal in length, fitth longest. Head long, the
lateral lobes a little divergent at apices, their margins up-
wardly reflexed. Pronotum rugose, sparingly punctate,
convex, without any central ridge, the lateral angles sub-
acutely rounded, the anterior lateral margins strongly serrate.
Scutellum rugose, scarcely so at apex, darkly and sparingly
punctate. Corium sparingly punctate, the punctures in some
places black and in small clusters. Body beneath and legs
sparingly punctate.
Long. 16-20 millim.; max. Jat. 10-12 millim.
Hab, Arabia; Hadramaut (Dent Exped., Brit. Mus.) ;
Abyssinia (Coll. Dist.).
Apparently most nearly allied to the West-African species
B. rugosus, Fabr., from which it differs by the non-ridged
pronotum &e.
Two Abyssinian specimens in my own collection reach the
maximum in size.
Genus GONOPSIS.
Gonopsis, Amyot & Serv. Hem. p, 180 (1848).
Bessida, Walk. Cat. Het. ili. p. 577 (1868).
Lethierry and Severin in their Catalogue (t. 1. p. 226)
placed this proposed genus of Walker in the Tessaratominee,
misled doubtless by the erroneous description of the head—
‘rounded in front.’
Gonopsis coccinea.
Macrina coccinea, Walk. Cat. Het. iii. p. 497. n. 7 (1868).
Bessida scutellaris, Walk. loc. cit. p. 578.
Gonopsis angularis.
Macrina angularis, Dall. List Hem. i. p. 860 (1851).
Gonopsis mantis.
Macrina mantis, Sial, QEfv. Vet.-Ak. Férh. 1853, p. 225. n. 2.
Lichelurhinus indicator, Walk. Cat. Het. iii. p. 499. n. 4 (1868),
Gonopsis bantu, sp. n.
Pur lish black ; head, anterior half of pronotum, a sub-
Phyllocephalinz, Urolabidine, and Acanthosomine. 225
marginal crenulate line on each side of the scutellum (ex-
tending for about two thirds its length), and lateral margin
of corium (excluding apex) pale ochraceous. Body beneath
reddish ochraceous, a pale oblique lateral sternal fascia on
each side extending through the pro- and mesosterna, on outer
side of which are two piceous spots, a submarginal elongate
black spot on each side of metasternum, and two submarginal
black fascize on each side of abdomen.
Antenne ochraceous, the second joint considerably longer
than the third, which is a little shorter than the fourth ;
fifth joint mutilated. Head coarsely punctate. Pronotum
rugulose, with a distinct central carination between the
lateral angles, which are subacutely prominent. Scutellum
strongly rugulose. Corium somewhat thickly punctate.
Long. 14 millim.; exp. pronot. angl. 8 millim.
Hab. Brit. E. Africa, Lake Kibibi (Gregory, Brit. Mus.). -
Genus DIPLORHINUS.
Diplorhinus furcatus.
Atelocerus? furcatus, Westw. in Hope Cat. i. p. 20 (1837).
Diplorhinus sinensis, Walk. Cat. Het. iii. p. 494, n. 2 (1868),
Diplorhinus quadricornis, Stal.
Diplorhinus quadricornis, Stil, En. Hem. v. p. 122 (1876).
Diplorhinus furcatus, Dall. (nec Westw.) List Hem. i. p. 359, n. i
(1851); Walk. Cat. Het. iii. p. 494. n. 1 (1868),
Genus TETRODA.
Tetroda histeroides.
Acanthia histeroides, Faby. Ent. Syst., Suppl. p. 526. n, 24 (1798).
Tetroda bilineata, Walk. Cat. Het. ii. p. 494, n. 11 (1868).
Tetroda obtusa.
Tetroda obtusa, Dall. List Hem. i. p. 357. n. 6 (1851).
Gellia (?) obtusa, Stal, En. Hem. v. p. 124 (1876).
Gellia obtusa, Atkins. Notes Ind, Rhynch. y. p. 106 (1888).
FRISIMELICA, gen. nov.
Body elongate. Head broad, lateral lobes somewhat re-
curved upwardly, meeting in front of central lobe at about
two thirds from base, which is moderately gibbous; antenne
with the second and fifth joints subequal in length, third
shortest. Pronotum convexly gibbous, the lateral margins
Ann. & Mag. N. Hist, Ser. 7. Vol. vi. 15
226 Mr. W. L. Distant on Dinidorine,
moderately laminate; posterior margin not wider than the
scutellum and inwardly angulated at centre; posterior angles
broadly rounded, from thence slightly sinuate to posterior
margin. Scutellum a little longer than half the length of
abdomen, moderately deflected on each side, narrowed at
about two thirds from base. Corium slightly and convexly
rounded from near base; membrane with longitudinal veins
extending slightly beyond apex of abdomen. Rostrum
reaching anterior coxee; second joint shortest, third joint a
little longer than fourth. Mesosternum gibbous and with a
central longitudinal levigate carination.
A genus to be placed near Gellia and Megarhynchus.
Frisimelica signata.
Cimex signatus, Fabr. Ent. Syst. p. 712. n. 76 (1775).
Phyllocephala signata, Dall. List Hem. i. p. 355. n, 10 (1851).
Hab. West Africa; Sierra Leone (Banksian Coll., Brit.
Mus.) ; Gambia (Brit. Mus.).
URroLABIDINZ.
Genus UROLABIDA.
Urolabida histrionica.
Urostylis histrionica, Westw. in Hoye Cat. i. p. 46 (1887).
Urolabida binotata, Walk. Cat. Het. ii. p. 415, n, 4 (1867).
Genus UROCHELA.
Urochela quadripunctata.
Urochela quadripunctata, Dall, Trans. Ent. Soc. Lond. 1850, p, 3, pl. ii.
tig. 1.
Urostylis lopoides, Walk. Cat. Het. ii. p. 414. n. 12 (1867).
Urochela distincta, sp. n.
Brownish ochraceous; lateral margins to pronotum and
basal lateral margins of corium luteous; a spot on lateral
margins of corium and a subbasal spot to lateral margins of
corium piceous; two discal rounded piceous spots on each
corium situate one above the other, the lower one near the
apical margin; connexivum alternately luteous and black ;
membrane cupreous, its apex paler; antenna piceous, the
basal area of fourth and fifth joints luteous. Body beneath
and legs brownish ochraceous, the spots on lateral margins of
pronotum and corium as above.
Phyllocephalinz, Urolabidine, and Acanthosomine. 227
Antenne with the first and second joints subequal in
length, third shortest, fourth longer than fifth ; body above
sparingly, coarsely, and darkly punctate ; lateral margins of
the pronotum oblique.
Long. 10-11 millim.; max. lat. 5 millim.
Hab. China; Kiukiang (Brit. Mus.).
Species wrongly included in the Urolabidine.
Genus Norius (Pentatomine).
Notius, Dall. List Hem. i. p. 155 (1851).
Ebora, Walk. Cat. Het. ii. p. 415 (1867).
In his description of the proposed genus Hbora Walker
describes the rostrum as “ extending a little beyond the fore
coxe,” whereas the intermediate core should have _ been
written. Misled by Walker’s differential comparison with
the genus Urochela, Lethierry and Severin placed Hbora in
the Urolabidine.
Notius depressus.
Notius depressus, Dall. List Hem, i. p. 155. n. 1, t. iv. fig. 1 (1851).
Ebora circumdata, Walk. Cat. Het. ii. p. 416, n. 1 (1867).
ACANTHOSOMIN.
Genus ABULITES.
Abulites sparsus.
Cimex sparsus, Germ. Silb. Rev. Ent. v. p. 174. n. 111 (1837).
Rhaphagaster fusco-wroratus, Walk, (nec Stal) Cat. Het. ii. p. 362. n, 48
(1867).
Genus ANDRISCUS.
Andriscus armatus.
Rhaphigaster? armatus, Dall. List Hem. i. p. 291, n, 48 (1851),
Genus ACANTHOSOMA.
Acanthosoma vittata.
Cimex vittatus, Fabr. Ent. Syst. iv. p. 104. n. 96 (1794); Syst. Rhyng,
p. 165. n, 52 (1803).
Acanthosoma vittatum, Dall. List Hem. i. p, 307, n. 13 (1851).
Anubis? vittatus, Leth. & Sev. Cat. Gén, Hém, t, i. p. 250 (1893),
15*
228 Mr, W. L. Distant on Dinidorine,
Genus SASTRAGALA.
Sastragala firmata.
Cuspicona firmata, Walk, Cat. Het. iii. p. 569 (1868).
Sastragala murreeana, sp. n.
Ochraceous, coarsely black punctate ; lateral pronotal spines
red, blackly punctate, long, directed forwards and upwards.
Body beneath and legs ochraceous, abdomen with reddish
apical spots.
Antennz with the second joint much longer than the third,
apical half of third piceous; remaining joints mutilated.
Pronotum and corium somewhat thickly and coarsely punc-
tate, the scutellum more sparingly so.
Long. 14-15 millim. ; exp. pronot. angl. 84-10 millim.
Hab. N. India, Murree (Atkins. Coll., Brit. Mus,, and
Coll. Dist.).
Sastragala heterospila,
Acanthosoma heterospila, Walk. Cat. Het. ii. p. 394. n. 15 (1867).
Sastragala affnis, Atkins. Journ. Asiat. Soc. Beng. vol. lvii. p. 344
(1889).
Sastragala edessoides, sp. n.
Luteous, coarsely and darkly punctate. Antenne luteous,
apical half of third joint and fourth and fifth joints piceous.
Second joint of antenne a little longer than the third;
pronotum and scutellum somewhat sparingly punctate, the
corium much more thickly so; pronotal angles long, robust,
slightly ascending ; abdomen above pale sanguineous, mar-
gins of connexivum ochraceous. Body beneath and legs
ochraceous, small stigmatal black spots and two similar spots
at posterior margin of apical segment.
Long. 14-15 millim. ; exp. pronot. ang]. 93-103 millim.
Hab. Sikkim (Atkins. Coll., Brit. Mus.); Naga Hills
(Doherty, Coll. Dist.).
The long robust pronotal angles render this species distinct,
and it has a striking structural resemblance to some species
of the American genus Edessa.
Sastragala elongata.
Acanthosoma elongatum, Dall, List. Hem, i. p. 809. n. 17 (1851).
Acanthosoma elongata, Atkins, Notes Ind. Rhynch, Het. y. p. 24 (1888),
Phyllocephaline, Urolabidine, and Acanthosomine. 229
Sastragala Hampson, sp. n.
Olivaceous, coarsely and darkly punctate; pronotal angles
acute, directed outwardly ; anal appendage provided with
two long red forceps; antennx olivaceous, apical half of
third and the whole of fourth joint piceous; fifth joint
mutilated.
Third joint of antenne slightly longer than second ; head
impunctate, the pronotum, scutellum, and corium about
equally coarsely and sparingly punctate; body beneath and
legs very pale yellowish green.
Long. 13 millim.; exp. pronot. angl. 9 millim.
Hab. Nilgiri Hills (Sir G. F. Hampson, Coll. Dist.).
Genus ANAXANDRA.
Anaxandra alaticornis.
Acanthosoma alaticornis, Walk. Cat. Het. iii. p. 573 (1868).
Anaxandra levicornis.
Acanthosoma levicorne, Dall. List Hem. i. p. 311. n, 24 (1851).
Acanthosoma levicornis, Notes Ind. Rhynch. Het. vy. p. 23 (1888) ;
Leth. & Sev. Cat. Gén. Hém. t. i. p. 254 (1893).
Anaxandra nigricornis.
Acanthosoma nigricornis, Walk. Cat. Het. iii. p. 574 (1868).
Anazandra nigrocornuta, Reut. Berl. ent. Zeitschr. xxv. p. 77 (1881).
Anaxandra bovilla, sp. n.
Brownish ochraceous, thickly and coarsely punctate ; pro-
notal angles largely developed, slightly recurved, and pointed
posteriorly at apices, which are a little paler in hue,
Antenne with the second joint longer than the third ;
posterior area of the pronotum from between the lateral
angles much more coarsely punctate ; scutellum very coarsely
punctate, the apical margins somewhat raised and levigate, a
central levigate line traversing the pronotum and scutellum ;
corium more thickly and finely punctate, with a discal levi-
gate spot, which in some specimens is very indistinct. Mem-
brane pale brownish. Body beneath and legs brownish
ochraceous.
Long. 10 millim. ; exp. pronot. angl. 8 millim.
Hab. Assam (Atkins. Coll., Brit. Mus.); Naga Hills
(Doherty, Coll. Dist.).
230 Mr. W. L. Distant on Dinidorine,
Genus STICTOCARENUS.
Stictocarenus placidus.
Acanthosoma placida, Walk. Cat. Het. ii. p. 397. n. 26 (1867).
Stictocarenus chlorophilus.
Acanthosoma chlorophila, Walk. Cat. Het. ii. p. 398. n. 27 (1867).
Stictocarenus suffusus, sp. n.
Pale ochraceous ; a basal fascia to pronotum, internal and
external areas (excluding bases) and apical area (broadest at
apex) of corium pale sanguineous. Body beneath and legs
pale greenish ochraceous ; tarsi testaceous ; abdomen with a
discal, longitudinal, pale greyish linear callus on each side.
Antenne brownish ochraceous ; second joint slightly longer
than the third, fourth and fifth joints subequal in length and
darker in hue. Pronotal angles distinctly subprominent,
pronotal lateral margins moderately concavely sinuate; san-
guineous fascia at base blackly punctate, remaining area
(excluding that of callosities) coarsely and concolorously
punctate. Scutellum coarsely punctate, but much less so
at basal area. Corium thickly and coarsely punctate, the
sanguineous internal area darkly punctate.
Long. 9 millim.; exp. pronot. angl. 44 millim.
Hab. King Island, South Seas (Brit. Mus.).
Apparently nearest allied to S. nigropunctatus, Reut., but
differing by the absence of black punctures to the scutellum
and also by the presence of the discal linear callosity to the
abdomen.
Genus ELASMOSTETHUS.
Elasmostethus lineatus.
Acanthosoma (Sastragala) lineata, Dall, Trans, Ent. Soc. v. p. 194 (1849).
Sastragala lineata, Atkins. Notes Ind. Rhynch. Het. v. p. 28 (1888).
Acanthosoma binotata, Walk. Cat. Het. ii. p, 395. n. 16 (1867).
Elasmostethus truncatulus.
Acanthosoma truncatula, Walk. Cat. Het. ii. p. 896, n, 18 (1867)
Elasmostethus asperus.
Acanthosoma aspera, Walk. Cat. Het. ii. p. 395, n. 17 (1867),
Phyllocephalinew, Urolabidine, and Acanthosomine. 231
EHlasmostethus nebulosus, sp. n.
Ochraceous, with coarse brown punctures; basal spot to
head, two anterior marginal spots to pronotum, and marginal
spots to connexivum black. Pronotum with brownish suffu-
sions principally on posterior area and at lateral angles ;
scutellum much suffused with dark brownish at base, disk,
and at each apical margin; a small pale levigate spot in each
basal angle; corium with a transverse central and a broad
apical brown suffusion ; abdomen above reddish brown, with
the extreme apex piceous.
Antenne with the two basal joints ochraceous, the re-
maining joints fuscous, second joint subequal to or slightly
shorter than the third. Body beneath and legs ochraceous ;
prosternum and femora darkly punctate; sternal spots near
coxe, stigmatal spots, and outer marginal spots at segmental
incisures black.
Long. 10 millim. ; exp. pronot. angl. 54 millim.
_ Hab. N. India, Naga Hills (Doherty, Coll. Dist.).
Elasmostethus nilgirensis, sp. n.
Ochraceous ; basal areas of pronotum and scutellum, claval
and apical marginal areas of corium, castaneous or reddish
castaneous ; pronotal angles, a basal submarginal line to
corium, and apical angle of corium black ; membrane hyaline,
fuscous at base and apex ; abdomen above reddish, with the
apical area black ; lateral margins of the pronotum, a small
spot in each basal angle of the scutellum, and a faint longi-
tudinal central line traversing the pronotum and scutellum
levigate pale ochraceous.
Antenne brownish ochraceous, second joint distinctly
longer than the third, apical joint somewhat infuscated ; pro-
notum, scutellum, and corium coarsely and sparingly punc-
tate, the central marginal area of corium much less punctate.
Body beneath and legs pale luteous ; the odoriferous apertures
and two small subapical abdominal spots black; apex of
abdomen reddish.
Long. 9 millim.; exp. pronot. angl. 44 millim.
Hab. Nilgiri Hills (Sir G. F. Hampson, Coll. Dist.) ; Uta-
kamand (Brit. Mus.).
Allied to the Chinese H. nubilus, Dall., but differing
structurally by the much greater length of the second joint of
the antenne.
232 Mr. W. L. Distant on Dinidorina,
Elasmostethus Lewitst, sp. n.
Ochraceous; pronotum, scutellum, and corium coarsely
and darkly punctate ; scutellum with a central cordate levi-
gate spot surrounded by castaneous shading, in which is a
short central dark lineate spot above and beneath; extreme
apices of pronotal angles, apical margins or only angles of
corium, a small spot at base of membrane, and the apical
abdominal segmental angle black ; abdomen above reddish,
with its Jateral margins ochraceous; membrane hyaline,
slightly brownish on inner and outer margins.
Antenne ochraceous, second joint distinctly longer than
the third, apical joint somewhat infuscated ; head transversely
wrinkled ; pronotum with two transverse levigate callosities
on anterior area; corium with the whole marginal area very
finely and concolorously punctate.
Long. 8 millim.; exp. pronot. angl. 43 millim.
Hab. Ceylon; Punduloya (Atkins. Coll, Brit. Mus. ;
G. Lewis, Coll. Dist.); Nilgiri Hills; Utakamand (Brit.
Mus.).
Elasmostethus delicatulus.
Acanthosoma delicatula, Walk. Cat. Het. ii. p. 397. n. 24 (1867),
Mormidea erythrospila, Walk. loc. cit. 111. p. 555 (1868),
Elasmostethus lineus.
Acanthosoma linea, Dall. List Hem. i. p. 308. n. 15 (1851).
Elasmostethus emeritus.
Cimex emeritus, Faby. Syst. Ent. p. 705 (1775).
In Banksian Coll. (Brit. Mus.).
Summarized Disposition of Walker's Genera and Species.
Dinidorinz, Phyllocephaline, Urolabidine, and
Acanthosomine.
Genera considered valid.
Enada, Walk. Cat. Het. ili. p. 485 (1868) (Tessaratomine).
Thalma, Walk. loc. cit. p. 503.
Urusa, Walk. loc. cit. p. 504.
Genera treated as synonymic.
Erga, Walk. Cat. Het. iii. p. 485 (1868),=Gen. Avona, Stal.
Ucia, Walk. loc. eit. 11. p. 407 (1867), =Gen. Panetius, Stal.
Ebora, Walk. loc, cit. iii, p. 415 (1868), =Gen. Notius, Dall. (Pentato-
; ; [mine.
Bessida, Walk. loc. cit. p. 577, =Gen. Gonopsis, Amy. & Serv.
Phyllocephalinz, Urolabidinez, and Acanthosomine. 233
Species considered valid and described wnder correct Genera.
‘Cyclopelta dorsalis, Walk. Cat. Het. iii. p. 478. n. 5 (1868),
? vilis, Walk. doc. cit. n. 6.
Aspongopus circumeinctus, Walk. loc, cit. p. 433, n. 28,
-—— e@neus, Walk. loc, cit. p. 484. n. 29,
Enada rosea, Walk. loc. cit. p. 485. n. 1 ( Tessaratomine).
Megymenum basale, Walk. loc. cit. p. 502. n. 13.
Thalma biguttata, Walk. loc. cit. p. 503. n. 1.
Urusa crassa, Walk. loc. cit. p, 504. n. 1.
Vrolabida octomaculata, Walk. loc. cit. p. 576,
Urochela discrepans, Walk. loc. cit. ii. p. 411. n. 4 (1867),
Urostylis lateralis, Walk. loc. cit. p. 412. n. 7.
sinensis, Walk. loc. cit. n. 8.
— lygoides, Walk. loc. cit. n. 9.
— fumigata, Walk. loc. cit. p. 413. 1. 10.
—— philoides, Walk. loc, cit. n, 11.
Species considered valid, but requiring generic revision.
Phyllocephala humerals, Walk. Cat. Het. iii. p. 490, n. 22 (1868), belongs
to gen. Dalsira, Amy. & Serv.
—— albidicosta, Walk. loc. cit. p. 491. n. 24, belongs to gen, Basicryptus,
Herr,-Schaff.
subtruncata, Walk. loc. cit. n, 25, belongs to gen. Basicryptus, Herr.-
Schaff.
Sehtemadiops ensignis, Walk, loc. cit. p. 495. n. 2, belongs to gen. Schizops,
Opin.
Bie eee chiens Walk. loc. cit. p. 497. n. 7, belongs to gen. Gonopsis,
Amy. & Serv.
Megarhynchus diversus, Walk. loc. cit. p. 498. n. 4, belongs to gen. Go-
nopsis, Amy. & Serv.
Cuspicona firmata, Walk. loc. cit. p. 569, belongs to gen. Sastragala,
Amy. & Serv.
Acanthosoma truncatula, Walk. loc, cit. ii. p. 396. n. 18 (1867), belongs
to gen. Elasmostethus, Fieb.
—— subducta, Walk. loc. cit. n. 23, belongs to gen. Sastragalu, Amy. &
Serv.
delicatula, Walk. loc, cit. p. 397. n, 24, belongs to gen. Elasmo-
stethus, Fieb.
—— placida, Walk. loc. cit. n. 26, belongs to gen. Stictocarenus, Stal.
— eeeenin, Walk. loc. cit. p. 898. n. 27, belongs to gen. Stictocarenus,
tal.
—— alaticornis, Walk. loc. cit. iii. p. 573 (1868), belongs to gen. Anaz-
andra, Stal.
oe pecorn Walk. doc. cit. p. 574, belongs to gen. Anaxandra,
Stal.
Spectes treated as synonymic.
Cyclopelta dotata, Walk. Cat. Het. iii. p. 479. n. 7 (1868), = Aspongopus
patruelis, Stal.
Aspongopus solitus, Walk. Joc. cit. p, 484. n, 30,= Aspongopus cuprifer,
Westw.
234 Lord Walsingham on Asiatic Tortricidae.
Erga roseoflua, Walk. loc. cit. p. 486. n. 1,= Axona longitudinalis, Westw.
(Tessaratomine).
Phyllocephala impressa, Walk. loc. cit. p. 489. n. 21,= Basicryptus dis-
tenctus, Sign.
Funesta, Walk. loc. cit. p. 490. n. 28,= Dalsira vicina, Sign.
Tetroda bilineata, Walk. loc. cit. p. 494. n. 11,= Tetroda histeroides, Fabr.
Diplorhinus sinensis, Walk. loc. crt. n. 2,= Diplorhinus furcatus, Westw.
Macrina seita, Walk. loc. cit. p. 496. n. 6,= Macrina juvenca, Burm.
vacillans, Walk. loc, cit. p. 497. n. 8,= Gonopsis affinis, Uhler.
Dichelorhinus indicator, Walk. loc. cit. p. 499. n. 4,=Gonopsis mantis,
Stal.
Megymenum instructum, Walk. loc. cit. p. 502. n. 14,= Megymenum
dentatum, Boisd.
Vera mutilata, Walk. loc. cit. ii. p. 408, n. 1 (1867), = Paneetius lobulatus,
Stal.
Urostylis lopoides, Walk, loc. cit. p. 414. n. 12,= Urochela quadrpunctata,
Dall.
Urolabida semicircularis, Walk. loc. cit. nu. 2,=Urolabida histrionica,
Westw.
——- binotata, Walk. loc. cit. p. 415. n. 4,= Urolabida histrionica, Westw.
Ebora cireumdata, Walk. loc. cit. p. 416. n. 1,= Notius depressus, Dall.
(Pentatomine).
Bessida scutellaris, Walk. loc, cit. iii. p. 578 (1868),= Gonopsis coccinea,
Walk.
Species the types of which are supposed to be in Australia.
Acanthosoma immunda, Walk. Cat. Het. iii. p. 573 (1868). National
Museum, Melbourne.
Ebora postica, Walk. loc. cit. ii, p.416. n.2 (1867). Nat. Mus., Melbourne.
? plana, Walk. loc. cit. n. 8. + a
——? patula, Walk. loc. cit. p. 417. n. 4. - -
XXXII.—Ast1atic Torrrrerp2.
By the Rt. Hon. Lorpv WatstneuaM, M.A., LL.D., F.R.S.
[Continued from p. 137.]
ARGYROPLOCE, Hb.
925. Argyroploce profundana, F.
Penthina profundana, Stgr. & Wk. Cat. Lp. Eur. 247. No. +625 [$925]
(1871)!; Chr, Hor. Soc. Ent. Ross. XIL. 225 (1876) 2
Hab. Europe’, Transcaucasta — Lenkoran’, 24 VI.
1874 (Christoph). JAPAN (Pryer, 1886).
Lord Walsingham on Asiatic Tortricide. 235
927 (1). Argyroploce vicinana, Rag.
Penthina vicinana, Rag. Ann. Soc. Ent. Fr. LXIII. (1894) 200-1.
No. 927 bis (1894) ?.
Hab. AMuR*. JAPAN—Yesso (Pryer, 1882).
926. Argyroploce dimidiana, C).
n. syn.=Schreberiana, L.
[Phalena (Tortrix)| dimidiana, Cl. Ic. Ins. I. Pl. VIII. 10 (1759).
Phalena (Tortrix) Schreberiana, L. Fn. Suec. (2 edn.) 348. No. 1338
(1761). {[Phalena (Tortrix)| Screberana, Cl. Ic. Ins. LI. Register,
p. (3) (1764). Penthina Schreberiana, Stgr. & Wk. Cat. Lp. Eur.
247, No. 926 (1871).
Hab. Eurorr. JAPAN—Howpo—Oiwake, VI.-VII. 1887
(Pryer).
Staudinger and Wocke quote this species thus :—“ Schreberi-
ana (Cl. Ic. 8. 10. Figura sine nom. typogr.) L. F.S. p. 348.”
In the Berlin, Tring, and Merton copies of Clerck’s ‘ [cones,’
Pl. VIII. f. 10 has the ENGRAVED name “ dimidiana.”
In the following copies “ dimidiana” has been erased,
“ Screberiana”’ being substituted in MS.: Mus. Br. (NH),
Herrich-Schiiffer, and Freyer (teste Zeller), Lin. Soc. Lond.
(Linneus’s own copy), Oxtord Mus. (Radeliffe Library). In
two copies, Mus. Br. (Banksian Library) and Mr. Godman’s,
“ Schreberiana”’ occurs in MS., and in the Zool. Soc. Lond.
copy the name is “ Scebertana”’ in MS.
This figure is indexed as “ Screberiana, 8. 10,” in the
register to the ‘ Icones,’ II. p. (3).
In the ‘ Fauna Suecica’ citations from Clerck’s ‘ Icones ’
occur (e. g. gelatella, Cl. VIII. 3) ; precedence must there-
fore be given to Clerck’s names, and since at least three
copies are still extant with dimidiana engraved on the plate,
this name must be accepted as valid. Linneus omits to
cite Clerck’s figure, and Clerck probably adopted Linneus’s
name when he issued his second part in 1764, but neither he
nor Linneus (in 1761) had the power to change a name
which had become established in 1759.
931. Argyroploce capreana, Hb.
Penthina capretina, Stgr. & Wk. Cat. Lp. Eur. 247. No. 931 (1871)';
Fern. Tr. Am, Ent. Soc. X. 31. No. 189 (1882) ”.
Hab. Eurore*. JApan (Pryer, 1886). LABRADOR’.
236 Lord Walsingham on Asiatic Tortricide.
931 (1). Argyroploce basipunctana, sp. n.
Antenne brownish grey. Palpi white. Head and thorax
brownish grey, the latter with a raised chestnut-brown tuft
posteriorly. Hore wings mottled with bluish grey and dark
fuscous, some obscure chestnut-brown patches interspersed
across the middle, followed by a rather shining white tornal
patch produced upward nearly to the costa, but not reaching
the apex, which is occupied by a triangular chestnut-brown
patch; there is some greyish fuscous mottling on the inner
and outer sides of the white patch ; on the costa are five pairs
of oblique geminated white streaks, two before and three
beyond the middle, the latter followed by two single ones
before the apex ; at the base is a conspicuous white spot on
the cell; cilia pale cinereous, mottled with brown and fuscous,
a dark brown line along the termen at their base, preceded by
a narrow white streak along the upper half of the termen.
Exp. al. 22 mm. Hind wings greyish brown; cilia pale
cinereous, a greyish brown line running through them near
their base. Abdomen and legs brownish grey.
Type, 2 (701385) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886). Two specimens.
931 (2). Argyroploce lactetfacies, sp. n.
Antenne brownish grey. Palpi short, porrect, median
joint rather thickly clothed, terminal joint small and partly
concealed ; white, shaded with greyish brown externally at
their base. Head greyish brown above, face white. Thorax
greyish brown, sprinkled with white, with a moderately deve-
loped tuft posteriorly. Fore wings milky white, strongly
shaded with bluish grey and greyish fuscous along the costa
and dorsum to two-thirds the wing-length, the costal shade
narrower at the base and widening outwards, the dorsal shade
wider at the base and narrowing outwards, both sprinkled
with whitish scales and with some whitish geminations on
the costa; there is a slight olivaceous tint at the outer por-
tion of these costal and dorsal shades passing across the pale
space between them, and below the middle of the wing
beyond it is a mixed blue-gr ey and greyish fuscous spot ; the
apical third white, with a slight olivaceous hue, an olivaceous
rey shade curving inwards and upwards from below the
middle of the termen, with three olivaceous grey costal streaks
above it; cilia greyish, sprinkled with white, a greyish
fuscous line along their base. Hap. al. 17 mm. Hind wings
Lord Walsingham on Asiatic Tortricide. 237
greyish brown; cilia pale whitish cinereous, with a slender
greyish brown shade running through them near their base.
Abdomen greyish brown. Legs whitish, hind tarsal joints
faintly spotted.
Type, & (60059) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886)—Howpo—Yokohama (Man-
ley, 1888). Four specimens.
This species appears to be allied to corticana, Hb., but the
pale ground-colour of the wing almost obliterates the central
fascia below the middle; the apical markings also are some-
what different and the palpi are white instead of dusky.
933 (1). Argyroploce auricristana, sp. n.
Antenne subochreous, basal joint white. Palpi white.
Head hoary grey above, white in front. Thorax olive
ochreous mixed with white, with a conspicuous golden olive
crest posteriorly, ore wings olive ochreous, shading to
olive-brown, especially along the costa; apical third of the
wing snow-white, the apex slightly tinged with brownish
grey; the olive ochreous colour is irrorated with lustrous
chalybeous scales which extend as shining silvery streaks
along the inner margin of the white space and around the
termen; cilia shining whitish, slightly clouded with grey.
Exp. al, 19-20 mm. Hind wings greyish brown; cilia
shining pale cinereous. Abdomen greyish brown. Legs
shining cinereous, hind tarsal joints spotted with brownish.
Type, & (701387); 2 (70138) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886)—Kzuszv (Leech, 1890). ‘len
specimens.
934 (1). Argyroploce geminata, sp. n.
Antenne (3) simple; greyish. Palpi porrect, with the
tuft on the underside of the median joint considerably deve-
loped; whitish, shaded with greyish fuscous externally.
Head whitish ochreous. Thorax with the tuft scarcely deve-
loped; whitish, mixed with greyish fuscous and olivaceous,
Fore wings white, much mottled, streaked, and clouded with
greyish fuscous, grey, and olivaceous; an irregular curved
band of mixed greyish fuscous and olivaceous within the
basal third, further trom the base on the dorsum than on the
costa, encloses a speckled and striated basal ‘patch, in which
is a tuft of olivaceous ochreous scales on the dorsum; the
238 Lord Walsingham on Astatic Tortricide.
space beyond this basal patch corresponds in colour with the
extreme base, and is followed by another oblique band,
greyish fuscous towards the costa, containing a few blackish
scales about its middle, with an olivaceous spot near the end
of the cell and a larger one below the fold, its outer edge
biangulated about the middle; beyond this the apical third
of the wing is white shaded with greyish, the apex and
termen mottled with olivaceous in a triangular form, with three
small elongate spots of black scales at its inner edge on veins
6, 7, and 8; the costa with a series of white geminated
streaks throughout, more plainly visible on the outer than on
the basal half of the wing and separated and divided by
greyish fuscous and olivaceous spots and streaklets; cilia
greyish fuscous sprinkled with white, at the tornus white,
tipped with olivaceous immediately above the angle. Lup.
al. 16 mm. Hind wings brownish grey ; cilia slightly paler,
with a darker shade running through them near their base.
Abdomen brownish grey. Legs whitish.
Type, & (70084); 2 (70144) Mus. Wlsm.
Hab. Japan (Pryer, 1886) —Kziusrv (Leech, 1890). Five
specimens.
This species greatly resembles sororculana, Zett., but differs
in the more streaky appearance of the basal two-thirds of the
fore wings, in the less developed thoracic tuft, and the longer
clothing of the median joint of the palpi; the antenne also
are somewhat stouter and the costal geminations are more
numerous and more clearly defined.
937. Argyroploce variegana, Hb.
Penthina variegana, Stgr. & Wk. Cat. Lp. Eur. 247, No. 937 (1871)';
Stgr. Hor. Soc. Ent. Ross. XV. 249 (1879) *.
Hab. Europe’. Asiatic TURKEY—Kauparenpix1ar—
Brussa, VI.-VII.’; Azozw—Smyrna’”; Hazes—Shar De-
vesy (Native Coll. 1890) ; Aruznra’.
938. Argyroploce pruniana, Hb.
8. pruniana, Hb.+pruneticolana, Z.
Penthina pruniana, Hb., var. pruneticolana, Stgr. & Wk. Cat. Lp. Eur.
248. No. 938 a (1871)"; Chr. Hor. Soc. Ent. Ross. XII, 225 (1876) *;
Stgr. Hor. Soc, Ent. Ross. XV. 249 (1879) °.
Hab, Kurore*. Asiatic TuRKEY—Brussa, V.*; Taurus’;
Lord Walsingham on Asiatic Tortricide. 239
Amasia, VI.*; Goelkutschuk*; Aintab, 10 V. 1892 (Native
Coll.). WN. Persta—Asterabad, V.’
940. Argyroploce atropunctana, Zett.
=§ dimidiana, Sdf.
Penthina dimidiana, Stgr. & Wk. Cat. Lp. Eur. 248. No, 940 (1871) ';
Ster. Hor. Soc. Ent. Ross. XV. 249 (1879)?; Fern. Tr. Am. Ent.
Soe. X. 31-2. No. 190 (1882) *.
Hab. Evrore*. Astatic TurRKEY—Szvas—Amasia, V.—
VII. Unirep States—Missouri’.
The adoption of dimidiana, Cl., in lieu of Schreberiana, L.,
renders dimidiana, Sdf., a homonym, and the name afro-
punctana, Zett., should be used for this species.
943. Argyroploce oblongana, Hw.
Penthina oblongana, Steger. & Wk. Cat. Lp. Eur. 248. No. 948 (1871).
Hab. Kvrorr. Asiatic TurkKeY—Hazez—Shar Devesy
(Native Coll, 1893).
945. Argyroploce gentianana, Hb.
Penthina gentiana, Stgr. & Wk. Cat. Lp. Eur. 248. No, 945 (1871) ';
Ster. Hor. Soc. Ent. Ross. XV. 249 (1879) *.
Hab. Europe’. Astatic TurKEY—Kavpavenpiki4r—
Brussa, VIII.’ JAPAN (Pryer, 1886).
946 (1). Argyroploce (?) catapittoma, sp. n.
Antenne smoky black, paler beneath; basal joint with a
white spot at its apex. Pal/pi black; the terminal joint and
the apex of the median joint white. Head black; face white.
Thorax black, with a white transverse band anteriorly and a
white patch posteriorly. Fore wings smoky black for two-
thirds their length, the outer third white, a large white patch
on the dorsum at one-fourth, and some small white spots along
the costa before the middle; there is a white spot at the end
of the cell near the edge of the black suffusion, and for a little
space beyond and below it, between the black and white
portions, occurs a patch of brownish purple accompanied by a
tew black scales; cilia white, with two costal, one apical,
and three terminal black streaks running through them, some
black scales are also scattered along the margin at their base ;
240 Lord Walsingham on Asiatic Tortricidae.
underside dark fuscous, with a slight purplish reflection (on
both wings), the tessellated white cilia showing very clearly.
Exp. al. 22 mm. Hind wings cupreous; cilia brownish
grey. Abdomen cupreous. Legs: posterior pair coppery
brownish, the tarsi blackish, with three or four white annula-
tions.
Type, 2 (61150) Mus. Wlsm.
Hab, CHINA — Pan-tse-Fang, VI.-VII. 1892 (Leech).
Unique.
A very distinct species, not closely allied to any known
Kuropean form. ‘The male is unknown, but it will probably
be found to belong to the genus Argyroploce.
963. Argyroploce Branderiana, L.
Penthina Branderiana, Stgr. & Wk. Cat. Lp. Eur. 249, No. 968 (1871).
Hab, Europe. JAPAN— Yusso (Pryer, 1882).
963 (1). Argyroploce inornata, sp. n.
Antenne dull cinereous. Palpt appressed to the face,
terminal joint short, concealed; pale ochreous at the sides,
touched with fuscous above. Head fuscous. Thoraz crested ;
greyish fuscous, slightly mottled with subochreous. Fore
wings greyish fuscous, indistinctly mottled with grey, sub-
ochreous and olive-brown to two-thirds the wing-length, the
apical third subochreous, shaded with olive-brown ; on.the
costa are five or six pairs of pale ochreous geminated streaks,
separated and divided by greyish fuscous, a slender slaty grey
line running from the pair nearest the middle of the costa to
the termen; below this line is a series of inconspicuous
narrow fuscous lines crossing the paler portion of the wing
towards the termen, but interrupted below the middle by an
elongate upright slaty grey patch, forming the outer edge of
the ocelloid spot; cilia pale ochreous, much mottled and
speckled with brownish fuscous, and with a brownish fuscous
line along their base. Hap. al. 24-25 mm. Hind wings
brownish fuscous ; cilia very pale ochreous, with a brownish
fuscous line running through them near their base. Abdo-
men brownish fuscous. Legs subochreous, hind tarsi barred
with dark fuscous.
Type, 2 (70161); g (70162) Mus, Wlsm.
Hab, Japan (Pryer, 1886). Six specimens,
Lord Walsingham on Asdtatic Tortricidae. 241
966 (1). Argyroploce major, sp, u.
Antenne and palpi cinereous. Head brownish ochreous.
Thorax dark fuscous, tegule brownish ochreous. Fore wings
brownish ochreous, tending to olive-brown across the middle,
much shaded with dark fuscous to two-thirds the wing-length
and crossed by numerous irregular waved pale steel-blue
bands, or chains of connected spots, beyond the base; the
fuscous shading occurs for the most part along the dorsum,
obliquely across the middle of the wing to near the tornus, at
the base of the extreme costa and below it at a little distance
from it, the apical third of the wing showing scarcely any
fuscous scales; the pale shining steel-blue bands are distri-
buted from the costa before the middle to the dorsum beyond
it, and profusely also on the pale apical third, where they
follow the outer margin of the oblique dark shade which
precedes it, and are carried round the costa and termen, but
a little within them; cilia light brownish ochreous along their
basal half, their outer half pale ochreous. zp. al. 20 mm.
Hind wings dark brownish fuscous ; cilia whitish cinereous,
tinged with ochreous at the apex. Abdomen dark brownish
fuscous. Legs whitish cinereous.
Type, 2 (70232); g (70565) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886)— Yusso (Pryer, 1882). Four
specimens.
In the hind wing veins 3 and 4 are normally connate, but
sometimes stalked.
970. Argyroploce stibtana, Gn.
Penthina stibiana, Stgr. & Wk. Cat. Lp. Eur. 249, No. 970 (1871)? ;
Stgr. Hor. Suc. Ent. Ross. XV. 250 (1879) ?.
Hub. Europe’. Astatic ‘TurKey—Szvas—Ak-Dagh,
13 VIL; Harezs—Shar Devesy, 15-18 VI. 1890 (Native
Coill.).
984. Argyroploce lacunana, Schiff.
Penthina lacunana, Stgr. & Wk. Cat. Lp. Eur. 250. No. 984 (1871) ';
Stgr. Hor. Soc. Ent. Ross. XV. 251 (1879) ?.
Hab. Europe’. Asiatic ‘luRKEY — Brussa, V.-Vi?
Cui1na—Chang Yang, 4000-6000 feet (Pratt, 1886); Mupin,
21 V. 1890 (Leech).
989 (2). Argyroploce obovata, sp. n.
Antenne brownish grey. Palpi whitish ochreous. Head
rust-brown. Thorax brownish grey, mottled with rust-brown
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 16
242 Lord Walsingham on Asiatic Tortricide.
and pale cinereous. Fore wings rather short and stumpy,
with rounded costa, obtuse apex, and rather obliquely convex
termen ; pale cinereous, mottled with grey, based and banded
with rust-brown, shaded with grey; the basal patch ill-
defined, its outer edge dilated a little inward on the fold;
the median band wide, attenuated inward to the middle of
the costa, beyond which are five oblique pale ochreous
geminated costal streaklets, a sinuous steel-grey line from the
first pair meeting a line from the apex a little above the
middle of the termen ; cilia pale ochreous, a rust-brown line
along their base. Hzp. al. 14mm. Hind wings brownish
grey ; cilia pale cinereous, a shade-line near their base. Ad-
dumen brownish grey. Legs pale cinereous.
Type, & (60386) Mus. Wlsm.
Hub. JAPAN (Pryer, 1886)—Howpvo—Foochau, IV. 1886
(Leech). Four specimens.
998 (2). Argyroploce humeralis, sp. n.
Antenne fuscous. Lalpi pale cinereous. Head and thorax
fuscous. ore wings with a basal patch extending to one-
fourth, bent outward about the middle, fuscous mottled with
leaden grey ; the space beyond it is pale ochreous, a slender
fuscous line running through it parallel to the outer edge of
the basal patch; the outer portion of the pale space is diffused
and strongly tinged with ferruginous ; along the costa a series
-of fuscous spots alternate with oblique geminated pale ochreous
streaks; from the third spot, about the middle, arises a
fuscous shade, dilated outward and downward to the dorsum
before the tornus and much mottled with leaden grey, a larger
patch ot which occurs at its outer edge above the tornus; the
outer portion of the wing is bright ferruginous, a slender
leaden grey line running from the fifth costal spot to the
termen below the apex ; cilia pale ochreous, based (and tipped
along the middle of the termen) with greyish fuscous. ap.
al. 17 wm. Hend wings dark greyish brown; cilia pale
cincreous, with a dark line running through them near their
base. Abdomen greyish brown. Legs pale cinereous, with
four blackish spots on the hind tarsi.
Type, & (70532) Mus. Wism,
Hab. Japan (Pryer, 1886). Unique.
1003 (1). Argyroploce (?) acharis, Btl.
Penthina acharis, Btl. Ill. Typ. Lp. Het. B. M. III. 80, Pl. LX. 9
(1879) |.
Hab. CoreEA—Gensan, VI. 1886 (Leech). JAPAN (Pryer,
On Variation in the Weasel and Hedgehog. 243
1886)— Yesso—Hakodate, VILL. 1886 (Leech). Hoxpo—
Yokohama’. Krusru (Leech, 1890)—Satsuma, V. 1886
(Leech).
This is not a true Argyroploce, as it does not possess a
thoracic tuit; it cannot be referred to Enarmonia, as at
present constituted, since veins 3 and 4 of the hind wings are
connate, not stalked.
[To be continued. }
XXXIIT.—WNote on Variation in the Weasel and Hedgehog.
By G. E. H. Barrerr-HAmMILTon.
I HAVE read with much interest the two notes by Dr. Kinar
Loénnberg which appeared in the ‘ Annals’ for May and
June 1900. In these days of much writing and little atten-
tion it is pleasant to find that one’s work attracts notice,
even if that notice be critical or condemnatory. It is doubly
valuable to receive criticism from one who resides in Scandi-
navia, a country whose climate and configuration lends itself
in a very peculiar manner to the formation of local subspecies
of mammals, or, in other words, to variation.
Before noticing Dr. Lounberg’s remarks I may say, by
way of preface, that in my studies of Kuropean mammals
my main object has been, first to record, and secondly to
throw light, however dimly, upon the origin of the numerous
variations which occur. ‘lhe making of species or subspecies
is therefore to me of quite subsidiary impurtance, and I| care
not a jot whether the torms upon which I find it necessary
to bestow technical names be styled species, subspecies,
races, forms, or phases. On the whole I incline to the latter
word; but the use of the term subspecies is now so general
and, I had thought, so well understood that 1 have found it
convenient. No one who works for any little time at such
matters can fail to meet with numberless ditficulties or to
notice the inequality between the various subspecies. That,
however, is the fault of the system, or, if you like, of the
animals themselves, which refuse to accommodate themselves
to any scheme which man can invent, and which conse-
quently excite the frequent protests of those who fail to see
the troubles which must beset any system of minute inquiry.
Even, however, if | were to find that I had made numerons
244 Mr. G. E. H. Barrett-Hamilton on
bad subspecies, I would vastly prefer to be on the side of
those who attempt to unravel the mysteries of variation (it
may bea task heavier than the cleaning of the Augean stables)
rather than to cultivate the icy scepticism of the modern school
of “Jumpers,” to whom the many phases of animal variation
are like the ripples of the ocean to the sailor—things to be
detested in proportion as their magnitude makes them trouble-
some. By such a school no real progress can now or ever be
made.
Let us look at Dr. Lénnberg’s criticisms. Admitting that
the meaning of a subspecies varies somewhat with different
authors, he supposes ‘that even in a subspecies the distin-
guishing characteristics (although they are of less import-
ance than specific ones) must be constant to a certain degree
and inherited from one generation to another; in the oppo-
site case it is only an individual variety. Such independent
individual varieties must not be called subspecies, in my
opinion, even if they are numerous and dominate in some
region.”
Reading my introductory remarks, it is easy to understand
where Dr. Lénnberg and I differ and where we agree.
Finding that the weasels of the far north turn white in
winter, while those of the south do not, 1 apply to each of
these, in their extremes highly distinguishable, phases of the
same animal a different trinomial name, This I find to be
the best method at my disposal of calling attention to such
differences. Dr, Liénnberg, on the contrary, prefers to
minimize the importance of these really important colour
changes by refusing to accord them the hall-mark of nomen-
clatural distinction.
But it is not this which puzzles and annoys Dr. Linnberg
so much as the existence in Scandinavia of weasels belonging
to both of these forms. Well, why not take things as they
are, and admit the difficulty, with the impossibility of ever
completely surmounting it? Is it altogether preposterous
that, while we have the regularly white-turning Putorius
nivalis typicus in North and Middle Sweden, and the always
brown P. n. vulgaris in Scania (connected, as we know they
are, by various intermediates), we should similarly find the
P.n. typicus on a mountain-top and the P. x. vulgaris in
the valley of the same parish? Is not all this due to the
same laws of climatic variation, and need it deter us from
further investigation to find that such variation is in the
highest degree perplexing? Shall we not rather do well to
Variation in the Weasel and Hedgehog. 245
accept the situation, pregnant as it is with interest, and to
welcome each step in the road to its explanation ?
It is, in fact, only to be expected that a mountainous
country like Scandinavia, with one flank fully exposed to the
damp blasts of the Atlantic, the other chilled by the near
proximity of a vast continent, should present us with at
least two or even more phases of each common mammal.
This is certainly the case, as Dr. Lénnberg points out, with
the polar hare, Lepus canescens, and | have shown that it is so
also with the squirrel and also probably with the long-tailed
field-mouse, Mus sylvaticus. Why, then, should Dr. Lénn-
berg be at such pains to demonstrate the occurrence of the
“vulgaris” form together with the “ ¢ypicus,” an incident
which was not only probable but necessary for the proper
appreciation of their réle as subspecies? And why should
Dr. Lénnberg regard such intergradation both of colour and
size as rendering the distinguishing characteristics derived
from them unimportant, when in the very same paragraph
he shows his thorough agreement with me that “ the variation
of the weasel certainly does not lack significance, because
intermediate stages occur which unite the extreme forms ”’ ?
Of Dr. Lénnberg’s second paper—‘ Note on the Indi-
vidual Variation of the Common Hedgehog ’’—I have less to
say. I had found what I thought to be solid points of
differentiation between the skulls of hedgehogs from England
and Scandinavia, whereby all the examples included in
a fair series contained in the British Museum of Natural
History were readily distinguishable. Relying on the rule,
soon learnt in working at mammals, that such differences,
even if slight, are usually not meaningless, I assumed that
they would be borne out by a larger series of specimens
than I at that time had before me. Dr. Lénnberg finds that
this is not so; and I can only say that, while I am sorry that
my opinion seems to have been erroneous, I am only too glad
to find here in the hedgehog another check to those who,
while refusing to recognize colour differences, pin their faith
with an inconsistent fidelity on what are frequently the
shadowy characters of the cranium. Such characters of the
cranium, as every year only scems to teach us, may be full
of value or worthless just according to the individual idio-
synerasy of the animal in which they occur. They are in
many cases not one bit more reliable than those presented
by colour, proportions, or size. In fact, in regard to some of
the more important cranial characters, such as those of the
246 Bibliographical Notices.
dentition of the vole *, 25 per cent. of specimens examined
may be abnormal, a fact which, when still larger numbers
are available, may yet prove the saving of my subdivision
of the hedgehogs.
In conclusion, I must thank my critic for the exceedingly
temperate and forbearing way in which his remarks are
couched. Criticisms thus conscientiously formed and fairly
expressed cannot surely fail to advance our science.
BIBLIOGRAPHICAL NOTICES.
Zoological Results based on Muterial from New Britain, New Guinea,
Loyalty Islands, and elsewhere, collected during the Years 1895,
1896, and 1897, by Arthur Willey, D.Se. Lond., Hon. M.A.
Cantab. Part IV. Cambridge, May 1900.
‘Tun long-delayed fourth part of Dr. Willey’s ‘ Zoological Results ’
is now before us and proves fully equal, both in interest and in the
general excellence of its contents, to its predecessors. It contains ten
memoirs, the majority of which are devoted to reports on the collec-
tions made by Dr. Willey in various groups of the animal kingdom.
Three, however, are on subjects of morphological interest. The first
of these is the opening paper of the volume by Mr. J. Stanley
Gardiner, ‘‘On the Anatomy of a supposed new Species of Ceno-
psanmic from Lifu.” Mr. Gardiner divides his subject into four
heads, dealing respectively with the general anatomy of the skeleton
and specific description, the general anatomy of the polyps, minute
anatomy, and some conclusions relating to the body-layers in the
Actinozoa. He comes to the conclusion that the whole filament of
the primary and secondary, and probably also that of the tertiary,
mesenteries is ectodermic in origin, and that the whole of the
digestion of the animal is performed by these filaments, and draws
the important deduction that the stomodeum of Actinozoa is not
comparable with that of the Triploblastica, but is rather, with
the mesenterial filaments, the homologue of the whole gut. The
so-called endoderm is homologous with the mesoderm of Triplo-
plastica, and the Actinozoan polyp ought to be regarded as a Triplo-
blastic form.
The second of the morphological papers is by Mr. J. J. Lister on
Astrosclera Willeyana, the type of a new family of sponges. This is
a very remarkable organism, with a massive calcareous skeleton of
polyhedral elements united to form a rigid skeleton and excluding
the soft parts, an arrangement which is only approached among
living sponges in the genus Petrostoma. Among several points in
which Astrosclera differs from the rest of the Porifera may be men-
tioned the absence of a central atrial space, the minute size of the
flagellated chambers, and the peculiar form of the flagellated cells,
* As shown by Mr. G, 8S. Miller, Jun.
Bibliographical Notices. 247
which appear to be without “collars” and to taper gradually into
the flagellum.
The skeleton of Astrosclera in many ways recalls that of the
extinct Pharetrones, and Mr. Lister devotes a considerable space to
the consideration of the possible relationship of the two groups,
but comes to the conclusion that there is no clear affinity between
them.
The paper on Asérosclera is followed in the volume by a memoir
by Mr. W. P. Pycraft, ‘On the Pterylography of the Megapodii,”
dealing with the pterylosis of the adult, the nestling, and the
embryo. In the course of some remarks on the nature of the nest-
ling plumage, Mr. Pycraft advances the view that this does not,
in Megapodius, consist of true neossoptiles, but of a growth of
feathers intermediate between the latter and the definitive contour
feathers of the adult. Similar feathers are found in the nesiling
owl. There are no preplumule and the true prepennsx are shed
during embryonic life.
The remuining papers in the volume are respectively by Dr. D.
Sharp on the Insects from New Britain, by Mr. L. A. Borradaile
on the Stomatopoda and Macrura, by Mr. Walter Collinge on the
Slugs, Miss Philipps on the Polyzoa, Miss Thornely on the Hydroids,
Professor Hickson and Miss Hiles on the Stolonitera and Aley-
onacea, and Dr. Ashworth on the Xeniidx, the whole series torming
a valuable addition to our knowledge of the zoology of the regions
visited by Dr. Willey.
A Monograph of the Coleopterous Families Corylophide and
Spheerlide. By the Rev. A. Marrurws, M.A. Edited by Parzip
B, Mason, M.R.CS., F.LS., F.Z.S., &c. London: 0. E. Janson,
1899.
Tais work is published in the same form as the author’s previous
monograph on the Trichopterygide. It consists of 220 pages of
letterpress, and is illustrated with nine plates in outline, with
details of structure. One hundred and sixty-nine species are
described. ‘he author concludes that the affinity of the Cory lophides
is with the Silphide and Leptinide, and places the families thus :—
Leptinide, Corylophide, Pheenocephalide, Silphide.
The Pheenocephalide consists of a single genus, Pheenocephalus
separated from the Corylophidz on account of the maxillie having
two lobes, &ce. Aphanocephalus, originally described as a member
of the Corylophide, is separated as a distinct family, Pseudo-
corylophide, chiefly on account of the maxille having “ three lobes,”
somewhat as in the Trichopterygide. The family is placed between
the Corylophide and Phzenocephalide in the work, but the author
would “retain it in the position now occupied by the whole family
in the vicinity of the Coccinellide.” This is a most interesting
point, for the Coccinellide, formerly placed at the end of the
Coleoptera entirely away from the Clavicornia, are now by some
eminent Coleopterists located with them.
The various genera included in the Corylophide are so diverse
“248 Miscellaneous.
that the author divides them, and we think rightly, into five tribes.
This diversity gives the study of this family great interest, as light
may be thrown on some of the difficult problems connected with the
classification of the Clavicornia, and may determine the proper
position of the Coccinellid, which is still somewhat doubtful.
Those who are acquainted with the author's work on the
Trichopterygidze will not, we feel sure, be disappointed with the
present volume, which shows everywhere the great care that has
been taken in its preparation. The small size of these insects
(apparently nearly all considerably less than two millimetres in
length) has deterred many from examining them; but this work
has rendered the study of the group possible even to an outsider, as
the figures of details are very good. The plates are admirably
executed, but we cannot help regretting that only half-figures are
given: these do not satisfactorily convey to the eye the form of the
insect, and there is room on the plates for complete figures.
The editor and publisher are both to be congratulated on the
publication of this work; they seem to have spared no pains to
make it worthy of the author. We presume the wing of T'richo-
pteryx on the cover, as it does not in any way indicate the contents
of the book, is to be regarded as the author’s entomological coat
of arms.
MISCELLANEOUS.
Priority or Usage.
To the Editors of the ‘ Annals and Magazine of Natural History.’
GentLeMEN,—Many who, like myself, had “the advantage of a
classical education” must remember the joy, alike of master and
scholar, when the “ note” said that a given word was an drat
Aeyopevor. It was as water in a dry land, as the word Mesopotamia
to the historical old woman.
And now to think of the joy of it! dos (see Sharpe, Preface to
‘ Hand-list of Birds,’ il. p. vi) is found to bea Greek word meaning
‘“‘ weariness.”” Whether Euripides ever used this word is very
doubtful ; but it seems certain that he never used it twice, and that
no other known Greek author ever used it at all. It is a true
dak Neyopevor, a gem of the purest water !
But see to what shifts the priority-purist who wants to apply
Apus to the swift (see the preceding volume of these ‘ Annals,’
p. 480) is reduced! At first sight it is astounding ; on reflection it
is seen to be appropriate; for, after all, what is the priority-purist
but a searcher after dag \eyéueva—names used once and decently
buried in some dusty magazine, which he drags to the light and
uses to eject world-known terms. Let these purists beware lest,
when the history of zoology be written, they find themselves drag
Aeyopevor, and not too politely at that!
Your faithful Servant,
14th July, 1900. F. Jerrruy Bec.
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY,
[SEVENTH SERIES. ]
No. 33. SEPTEMBER 1900.
XXXIV.—On some Malacostracous Crustaceans from Ma-
Taysia in the Collection of the Sarawak Museum. By
W. F. Lancuester, M.A., King’s College, Cambridge.
[Plate XII.]
THE Crustaceans with which I deal in this paper were sent
to me by Mr. R. Shelford, B.A., Curator of the Sarawak
Museum, when I was at Singapore in the early part of 1899,
and I am greatly indebted to him for giving me the oppor-
tunity of identifying them. He has, further, been kind
enough to let me retain all but two of the specimens, and
those that may be desiderata to the Natural History Museum
at South Kensington will be deposited there. It is a small
collection of 38 species (27 genera), and only one of these is
new ; but some, in particular the Potamons, of which there
are five species, have only been recently described. This
Potamon group, which seems to be especially well represented
in Borneo, and which ranges there from sea-level to 6000 feet,
numbers many species of more or less definiteness, and
greater additions to our collections will no doubt enable us
to reduce the number of these somewhat and at the same
time to get a clearer idea of their mutual affinities. There is
also a specimen of a Callianassa, unfortunately somewhat
damaged, but almost certainly identical with OC. Martensi,
Ann, & Mag. N. Hist. Ser. 7. Vol. vi. re
250 Mr. W. F. Lanchester on some
first described from Mauritius by Mr. Miers and later from
Amboina by Dr. de Man, which, if my diagnosis be correct,
would point to a disappearance of the eyes in this species ;
for the corne, which in Mr. Miers’s younger example are of
normal size, are in this one small and out of all proportion to
their peduncles, which are flattened and produced, so as to
overlap them on all sides.
Since my return from the Straits I have been occupied in
describing the Brachyura collected there by Mr. Bedford and
myself: the paper dealing with these was read to the Zoolo-
gical Society in June, and will appear in the ‘ Proceedings ’ of
that Society later in the year. This being so, I have contented
myself with giving references to that paper in cases where, in
this, species are mentioned which I have already described.
As regards what few measurements I have taken, I should
say that, except where otherwise stated, I have measured the
breadth between the bases of the last antero-lateral spines and
the length between the posterior margin and a line joining the
bases of the internal supraocular spines ; my reasons for this
I have given in my earlier paper.
It may perhaps be advisable to add that the numbers
given in Roman figures with each species correspond to
labels atfixed by Mr. Shelford, and are inserted for his better
convenience in identifying the corresponding exaimples in his
inuseum.
BRACHYURA.
IT. Genus ATERGATIS, de Haan.
1. Atergatis integerrimus, Lam.
Cancer integerrimus, Lam. Hist. Anim. sans Vert. t. v. p. 278 (1818).
Atergatis integerrimus, de Haan, Crust. Jap. p. 46, pl. xiv. fig. 1
(1839) ; A. M.-Edw. Nouv. Arch. Mus. t. 1, p. 285 (1865).
Distr. Philippines, Malaysia, Siam, Java, Tuticorin.
No. cctxxul. Hab. Singapore.
A female.
Dim.: Length 34°5 millim., breadth (greatest) 55:5.
2. Atergatis foridus, Rumph.
Cancer floridus, Rumphius, d’Amboinsche Rariteit-Kammer, p. 16,
pl. viii. fig. 5 (1705). :
Cancer ocyroé, Herbst, Naturgesch. d. Krab. pl. liv. fig. 2 (1798).
Atergatis floridus, A. M.-Edw. Nouy. Arch. Mus, t. i. p. 243 (1865).
Distr. Ceylon, Malay Archipelago, Duke-of-York Island,
Samoa, Clairemont.
Malacostracous Crustaceans from Malaysia. 251
No. cocx. Hab. Pulau Satang.
A female.
Dim.: Length 30 millim., breadth (before last tooth) 43:5.
II. Genus CarPiLopEs, Dana.
3. Carpilodes socius, Lanchester.
Carpilodes socius, Lanchester, P. Z. 8. Lond. (1900).
INXstr. Singapore.
No. coct. Hab. Singapore.
A fine male specimen, quite agreeing with my original
description. Colour deep red above, a lighter brown-red
below ; fingers brown, with dirty-white tips.
Dim.: Length 17:25 millim., breadth 29°25.
ILI. Genus Actma, de Haan.
4. Acta areolata, Dana.
Actea areolata, Dana, U.S. Expl. Exp., Crust. vol. i. p. 162, pl. vill.
fig. 1 (1852); A. M.-Edw. Nouy. Arch. Mus. t. i. p. 264 (1865).
Distr. Port Molle, Mergui Is., Sooloo Sea.
No. crxxit. //ub. Singapore.
A male, in which, as with other specimens from Singapore,
the internal lobule of the protogastric lobe is no broader than
the mesogastric ; front only moderately prominent.
Dim.: Length 9 millim., breadth 15:5,
5. Actea pulchella, var. modesta, de Man.
Acteodes modestus, de Man, Arch. f. Naturgesch. Jahrg. liii, Bd. i.
p. 257, pl. ix. fig. 2 (1887).
Actea pulchella, Lanchester, P. Z. S. Lond. (1900).
Distr. Singapore, Amboina.
No. cccxcr. lab. Singapore.
A female. I cannot quite convince myself of the specific
distinctness of Actwodes modestus and Actwa pulchella. The
genus at least cannot be sustained, based as it is practically
on the one character of the greater or less emargination of the
finger-tips. But it is, again, more particularly on this cha-
racter that Dr. de Man has formed his species— Ks scheint
mir nun, dass sich die pulchella von unserer Art gleichfalls
durch die Scheerenfinger unterscheidet.” These differences
are :—in A. pulchella the fingers meet along their length, are
only slightly emarginate, and the dactyl slants obliquely ; in
A. modestus the fingers gape a little, the tips are strongly
1
252 Mr. W. F. Lanchester on some
emarginate, and the dactyl curves downwards. Now this
example and two males which I have already described from
Singapore (¢. ¢c.) show the modestus arrangement pretty
clearly, though even in this small number one may see a
degree of gradation, though a slight one, to the pulchella
form; while a fourth example, a female, rather smaller
(already described, ¢. ¢.), shows the pulchella arrangement,
not perfectly, but distinctly. There are no other noticeable
differences between the two forms (sonstige Unterschiede
giebt es kaum,” de Man, ¢. c.), and I do not think the slight
variations in the hand-structure are as yet more than varia-
tions. It is true that three distinctions are noticed, but these
three are really only one variation, the curvature of the
dactyl and the gape of the fingers being implied by the
emargination of the finger-tips. And the amount of this
emargination, even in my four examples, is not at all con-
stant, the two smaller examples having sharper finger-tips in
proportion. It seems to me, therefore, better to retain
Dr. de Man’s specific name in a varietal sense for those forms
of A. pulchella in which the spoon-shaped arrangement is
well marked, while leaving it open to decision as to whether
the difference is truly varietal, or only one of age, sex, or
both.
Dim.: Length 11 millim., breadth 16°25.
IV. Genus Errtsopgs, Dana.
6. Htisodes anaglyptus, M.-Edw.
Etisus anaglyptus, M.-Edw. Hist. Nat. Crust. t. i. p. 411 (1835),
Etisodes anaglyptus, Lanchester, P. Z. S. Lond. (1900).
Distr. Philippines, Clairemont, Torres Strait.
No. cott. Hab. Pulau Satang.
A female. I have already (¢. c. supra) given a fuller
description of this species, as a supplement to M.-Edwards’s
short diagnosis.
Dim.: Length 35 millim., breadth 52.
V. Genus Pitumnus, Leach.
7. Pilumnus vespertilio, Fabr.
Cancer vespertilio, Faby. Suppl. Entom. p. 838 (1798).
Pilumnus vespertilio, M.-Edw. Hist. Nat. Crust. t. i. p. 418 (1885) ;
Haswell, Catal. Austr. Crust. p. 65 (1882).
Malacostracous Crustaceans from Malaysia. 253
Distr. Seychelles, Tuticorin, Malay Archipelago, N.W.
Australia, New Zealand, Samoa.
No. ccctxv. Hab. Singapore.
A female.
Dim.: Length 11 millim., breadth 14:5.
VI. Genus Errputa, Latr.
8. Eriphia levimana, var. Smithi’, McLeay.
Lriphia Smith, McL. Annul, in Smith’s Zool. 8. Africa, p. 60 (1838).
Eriphia levimana, v. Smithii, Miers, Ann, & Mag. Nat. Hist. ser. 5,
vol. v. p. 287 (1880).
Distr, Natal, Kurrachee, Singapore, New Guinea, off
Madagascar.
No. cotvi. Hab. Natunas.
A female.
Pim.: Length 39°5 millim., greatest breadth 54.
VII. Scytua, de Haan.
9. Scylla serrata, Forskal.
Cancer olivaceus, Herbst, Naturg. d. Krab. ii. p. 157, pl. xxxviil. fig. 3
(1796).
Lupea tranquebarica, M.-Edw. Hist. Nat. Crust. t. i. p. 448 (1835).
Scylla serrata, de Haan, Faun. Japon. Crust. p. 44 (1839) ; Haswell,
Cat. Austr. Crust. p. 79 (1882).
Distr. Cape of Good Hope, Natal, India, Malay Archi-
pelago, China, New Caledonia.
No. Lxxx1x. Hab. Moratabas.
A young female.
Dim.: Length 48°5 millim., breadth 73.
VIII. Genus Neprunus, de Haan.
10. Neptunus pelagicus, Linn.
Cancer pelagicus, Linn. Syst. Nat. (ed. xii.) p. 1042 (1766),
Lupea pelagica, M.-Edw. Hist. Nat. Crust. t. i. p. 450 (1835).
Neptunus pelagicus, Hasw. Cat. Austr. Crust. p. 77 (1882).
Distr. Muscat, Persian Gulf, India, Malay Archipelago,
Port Curtis, Sydney, Shanghai, Zanzibar.
No. ccctxvi. Hab. Buntal.
A female.
Dim.: Length 28 millim., breadth 52°5.
= .
254 Mr. W. F. Lanchester on some
No. ccoxxx. Hab. Santubong.
A male.
Dim.: Length 16°25 millim., breadth 28.
1 R.
Length of last lateral teeth in 9 =5 mm. 9°5 mm.
” ” ” g=45 ,, 45,
IX. Genus Gontosoma, A. M.-Edw.
11. Goniosoma affine, Dana.
Charybdis affinis, Dana, U.S. Expl. Exp., Crust. vol. i. p. 286, pl. xvii.
fie, 2 (1852).
Gonrosoma affine, A. M.-Edw. Arch. Mus. t. x. p. 384 (1861) ; de Man,
Mergui Crust. p. 80, pl. v. fig. 2 (1888).
Distr. India, Mergui Is., Singapore.
No. ccocxxxt. Hab. Buntal.
A female, in which the two median lobes of the front are
broken off. I can detect no signs of teeth on the posterior
margins of the penultimate joints of the natatory (last) legs.
Dim.: Length 23°5 millim., breadth 37.
12. Goniosoma roseum, Jacq. et Lucas. (Pl. XII. fig. 1.)
Thalamita rosea, Jacq. et Luc. t. iii. p. 5, pl. v. fig. 11 (=Hombron
et Jacq. Voy. au Péle Sud, Crust. 1853).
Goniosoma roseum et rostratum, A. M.-Edw. Arch. Mus. t. x.
pp. 378-9, pl. xxxv. fig. 2 (1861).
Distr. Mouths of Ganges, Calcutta, Gulf of Martaban, New
Guinea.
No. 1x. Hab. Buntal.
A male, in which there is a small distinct tooth on the
base, posteriorly, of each first antero-lateral tooth, and a slight
pubescence on the branchial regions. G'. rostratum differs
only in being less orbicular, in having the two median teeth
of the front more advanced, and in the possession of only two
spines on the hand, and I do not think it is specifically
distinct.
Dim.: Length 17°5 millim., breadth 24°5.
X. Genus THALAMITA, Latr.
13. Thalamita Dane, var. Stimpsonit, A. M.-Edw.
Thalamita Stimpsoni, A. M.-Edw. Arch. Mus. t. x. p. 862, pl. xxxv.
fig. 4 (1861).
Thalamita Dane, de Man, Mergui Crust. p. 78, pl. iv. figs. 8, 9 (1888).
Malacostracous Crustaceans from Malaysia. 255
Distr. Torres Strait, Sandwich Islands, W. Australia,
Malay Archipelago.
No. cccititr. Hab. Singapore.
A young female.
Dim,: Length 8 millim., breadth 12°35.
XI. Genus Poramon, Sav.
14. Potamon (Perithelphusa) Biittikofert, de Man.
Potamon (Perithelphusa) Biittikofert, de Man, Notes from the Leyden
Museum, vol. xxi. pts. i—iii. pp. 80-86, pl. vi. fig. 6 (1899).
Distr. Sintang.
No. coxcit. Hab. Mt. Matang, 3000 feet.
A male. Dr. de Man describes the “ ischial line” on the
posterior maxillipedes as being parallel to the inner margin of
the ischial joint; in this example it is oblique, slanting away
from it from before backwards.
Dim.: Greatest length 28°5 millim., greatest breadth 35.
15. Potamon (Perithelphusa) borneense, var. hilare, de Man.
Potamon borneensis, var. hilaris, de Man, Notes Leyden Mus. vol. xxi.
pts. iii. p. 71, pl. v. fig. 4 (1899).
Distr. Sintang.
No. xcvu. Hab. Kuching: freshwater.
A male.
Dim.: Greatest length 18°5 millim., greatest breadth 23.
16. Potamon (Parathelphusa) tridentatum,
var. incertum, Lanchester. (PI. XII. fig. 2.)
Potamon (Parathelphusa) tridentatum, var. incertum, Lanchester,
P. Z.S. Lond. (1900).
Distr. Singapore (Botan. Gardens).
No.cxxxvi. Hab. Kuching: presumably freshwater, but
I have no note to that effect.
A male, quite agreeing with the forms I have already
described from Singapore; though in this specimen the
lobulation of the extraorbital tooth is even more striking,
giving it all the appearance of having four antero-lateral teeth,
such that the second is truncated, while the rest are sharp.
Dim.: Length (including rostrum) 30°5 millim., breadth
between last teeth 37:5.
256 Mr. W. F. Lanchester on some
17. Potamon (Geothelphusa) Birgeri, de Man.
Potamon (Geothelphusa) Biirgeri, de Man, Notes Leyd. Mus. vol. xx1.
pts. i-ili. pp. 121-127, pls. xi., xii. fig. 14 (1899).
Distr. Mount Liang Koeboeng.
No. xcv. Hab. Kuching.
A female example, quite agreeing with Dr. de Man’s
description of a male, except in regard to the following
points :—The left chelipede is the larger, and not the right ;
the fingers, moreover, which on the right side meet all along
their length, are somewhat widely separate on the left. The
mesogastric groove is continued faintly on to the front, and
shows a tendency to bifurcation behind; there is also a
faint oblique depression between the gastric and branchial
regions on each side.
Dim. : Greatest length 15°5 millim., greatest breadth 21.
18. Potamon (Thelphusa) bidiense, sp. n.
(Pl. EOE tie: 13.)
No. ccctxxvu. Hab. With this specimen I have the
following note from Mr. Shelford :—“ Caves at Bidi in pools ;
the caves were absolutely dark. Body pale brown, legs
white.”
A male. This form is closely related to one obtained by
Dr. de Man from the Dutch Expedition to Central Borneo, and
named by him P. Melanippe, which is itself allied to P. Aus-
tentanum, a form described by Wood-Mason from Assam.
Like them, this specimen is most noticeable for the length and
slenderness of its legs ; but the penultimate pair are a little more
than three times as long as the carapace, and their meri are also
longer than the carapace by nearly a fifth of their length. In
this, too, they differ from P. Melanzppe in that the meri of the
last four legs are armed with a small blunt spine anteriorly,
quite close to the distal end. The external maxillipeds are
precisely similar to those of P. Melanippe; the chelipedes, too,
are essentially similar, except that the fingers do not quite
meet along their length and their tips decussate. The shape
of the male abdomen is, however, quite different: the stxth
segment is indeed quite similar to that of P. Melanippe; but
the -fifth, instead of being narrower at its base than at its
extremity, is, if anything, ever so little broader, with its
sides straight and practically parallel; as a consequence of
this, the transition from the narrower terminal to the broader
basal segments is much less marked than in Dr. de Man’s
form.
Malacostracous Crustaceans from Malaysia. 257
Epigastric lobes very prominent; no crest parallel to the
anterior margin of the front, general surface of the carapace
rather thickly punctate than granular. Upper surface of
meri of ambulatory legs with two granular eminences. ‘There
is a minute epibranchial tooth as in P. Melanippe, behind
which the sides are obliquely rugose. ,
In regard to the habitat from which it was taken, there is
no evidence to show that this species is essentially a cave-
dwelling one; though its occurrence in a dark cave is very
interesting, and shows the possibility of such being the case.
The eyes in this individual are normal; its colour is perhaps
paler than in the Thelphuside generally (but ef. Potamon
bicristatum, de Man: “ carapace lead-coloured, legs yellowish
marmorate’”’), But a loss of colour of this nature is not
necessarily a permanent feature, and proves very little as to
the ways of the species as such. The possibility must always
be borne in mind that a particular habitat, such as this, may
be one temporarily acquired by certain, possibly locally limited,
members of a species; should this residence become _per-
manent, a local race may be formed. But in this case, at
least, the evidence is very little, and not much can be deduced
from it in either direction.
Dim.: Greatest length 12°25 millim., greatest breadth 14;
length of penultimate pair of legs 40, length of merus of
penult. pair 15.
XII. Genus SESARMA, Say.
19. Sesarma calypso, de Man.
Sesarma (Parasesarma) calypso, de Man, Zoolog. Jahrb. Syst. Bd. ix.
pp. 185-9, fig. 34 (1896).
Sesarma calypso, Lanchester, P. Z. S. Lond. (1900),
Distr. Atjeh, Singapore.
No. xxxviul. //ab. Buntal.
A female. Superior border of hand raised to a strong crest on
its inner side, against which the four peclinated ridges abut.
Fingers with seven or eight of the characteristic ‘“ 'Treppen-
formig ” tubercles.
Dim.; Greatest length 16 millim., breadth (including ex-
ternal orbital angles) 20°5.
20. Sesarma quadrata, Fabr.
Cancer quadratus, Fabr. Ent. Syst. Suppl. p. 841 (1798)
Grapsus (Pachysoma) quadratus, de Haan, Crust. Jap. p. 222, pl. viii.
fig. 3 (1859).
Distr. Malay Archipelago, Japan.
258 Mr. W. F. Lanchester on some
No. Lxxx1u. Hab. Santubong.
A small female.
‘Two obscure pectinated ridges, and 8-10 low tubercles on
the dactyl.
Dim.: Greatest length 9°25 millim., breadth (including
external orbital angles) 12.
XIII. Genus Ocypope, Fabr.
21. Ocypode ceratophthalma, Pallas.
Cancer ceratophthalmus, Pall. Spic. Zool. ix. p. 83, pl. v. fig. 17 (1772).
Ocypode ceratophthalma, Fabr. Suppl. Ent. Syst. p. 347 (1798) ; Miers,
Ann. & Mag. Nat. Hist. ser. 5, vol. x. p. 379, pl. xvii. fig. 1 (1882).
Distr. Very wide; from EK. Africa to the Pacific.
Nos. LXXXVI, CCCLXI., ccxc. Hab. Santubong, Buntal.
A male, with styles 11°5 millim. long (breadth of carapace
at epibranchial angles 36); a female, with styles :75 long
(breadth of carapace 28) ; and a small male, with no ocular
styles (breadth of carapace 18°5).
gal o 2.
millim. millim. millim.
Dim.: Greatest length of carap. 32 15:5 24°5
Greatest breadth of carap. 36 18°5 28
XIV. Genus Uca, Leach (= Gelasimus, Latr.).
22. Uca annulipes, M.-Hdw.
Gelasimus annulipes, M.-Edw. Hist. Nat. Crust. t. ii. p. 55 (1837) ;
de Man, Mergui Crust. p. 118, pl. viii. figs. 5-7 (1888), ubi syn.
Distr. Zambesi, Durban, India, Malay Archipelago, Samoa,
Lu-chu (China).
No. xxv. Hab. Santubong.
A male, left-handed.
Dim.: Greatest length 7°5 millim., breadth (with external
orbital angles) 13°5.
XV. Genus MACROPHTHALMUS, Latr.
23. Macrophthalmus carinimanus, M.-Edw.
Macrophthalmus carinimanus, M.-Edw. Hist. Nat. Crust. t. ii. p. 65
(1837); Miers, Ann. & Mag. Nat. Hist. ser. 5, vol. v. p. 8306 (1880) ;
Haswell, Cat. Austr. Crust. p. 88 (1882).
Distr. Malaysia.
Malucostraccus Crustaceans from Malaysia. 259
No. ccctv1. Hab. Santubong.
A male.
Dim.: Greatest length of carapace 9°5 millim., breadth
(across ext. orb. angles} 21.
24. Macrophthalmus depressus, Riippell.
Macrophthalmus depressus, Riipp. Beschreib. 24 Art. kurzschw. Krab.
p- 19, pl. iv. fig. 6 (1830).
Re aes affinis, Guérin, Crust. ‘ Favorite,’ p. 172, pl. 1. fig. 2
Air op hihi depressus, Hend. Trans. Linn. Soc., 2nd ser. Zool. v.
p. 889 (1893) ; de Man, Zoolog. Jahrb. Syst. Bd. viii. p. 578 (1895).
Distr. Pamban, Rameswaram, Bombay, Red Sea, North
Australia.
No. xiv. Hab. Buntal.
A male. The granulations on the carapace are very fine,
distinctly visible under the lens; and there is a well-marked
mesial groove on the front, extending back to the level
of the external orbital angles. In a smaller female specimen
Dr. de Man found no spine on the merus of the last leg;
there is one present in this example, which is a little larger.
Prof. Henderson (¢. ¢.) says: ‘The ambulatory legs are
pubescent, with a single tooth near the anterior distal end of
merus,’ —making no exception as regards the last pair.
I note, in addition, in my specimen the presence of a thick
hairy tuft on the anterior surface of the merus of the cheli-
pedes, and another at the base, internally, of the fingers.
Dim.: Greatest length of carapace 11°25 millim., breadth
(across orbital angles) 14.
Prof. Henderson gives the length of a male as 11 millim.,
and its breadth 17; Dr. de Man’s specimen is given as
8°25 millim. long and 12-2 broad.
XVI. Genus CALApPPA, Fabr.
25. Calappa hepatica, Linn.
Cancer hepaticus, Linn, Syst. Nat. p. 1048 (1766).
Calappa tuberculata, Fabr. Ent. Suppl. p. 346 (1798) ; Herbst, Naturg.
d. Krab, i. pl. xiii. fig. 78 (1796).
Calappa hepatica, Hasw. Cat. Austr. Crust. p. 186 (1882).
Distr. Red Sea, E. Africa, Natal, India, China, Sandwich
Is., Australia.
No. cccxxxvil. Hab. Natunas.
A male.
Dim.: Length 35 millim., greatest breadth 55,
260 Mr. W. F. Lanchester on some
XVII. Genus Marura, Fabr.
26. Matuta victrix, Fabr.
Cancer victor, Fabr. Ent. Syst. ii. p. 449 (1793).
Matuta victor, id. Ent. Suppl. p. 369 (1798).
Matuta victrix, Miers, Trans. Linn. Soc., 2nd ser. Zool. vol. i. p, 243,
pl. xxxix. figs. 1-3 (1877).
Distr. Red Sea, EK. Africa, Natal, Madras, Japan, Port
Jackson, Fiji Is.
No. vi. Had. Santubong.
A young male.
Dim.: Length 26 millim., breadth 28°5.
ANOMURA.
XVIII. Genus Ca@nosita, Latr.
27. Cenobita spinosus, M.-Edw., var. olivieri, Owen.
Cenobita oliviert, Ow. Crust. ‘ Blossom,’ p. 84; Dana, U.S, Expl. Exp.
Crust. vol. i. p. 470 (1852) ; Hasw. Cat. Austr. Crust. p. 160 (1882).
Cenobita spinosa, var. olivierit, Ortmann, Zool. Jahrb. Syst. Bd. vi.
p. 318, pl. xii. fig, 24 (1892). :
Distr. Madras, Nicobar, Tahiti, N.W. Australia.
No. cccx1x. Hab. Santubong.
A male. Fore part of carapace much less tumid than in
the species; outer surface of penultimate joint of the left
third leg with a rounded crest in its distal half. It is possible
that this variety is the young stage of the species. A speci-
men in the Museum collection at 8. Kensington, labelled
“CO, olivier’, Pelew I.,” should, I think, be referred to C. per-
lata. Length of carapace 27°5 millim.
XIX. Genus PETROLISTHES, Stimpson.
28. Petrolisthes hastatus, Stm.
Petrolisthes hastatus, Stm. Proc. Ac. Nat. Sci. Philad. p. 241 (1858).
Porcellana inermis (Heller), de Man, Merg. Crust. p. 212 (1888).
Petrolisthes hastatus, Ortmann, t. c. p. 260 (1892).
Distr. Mergui, Nicobar, Noordwachte I. (Java), Japan.
No. cctxiv. Hab. Pulau Satang.
A female. I have compared this with Dr. de Man’s
slightly smaller specimens, and find the following differences :—
The antero-lateral margin in this female is grooved in its
anterior third, this groove passing up and obliquely backwards
Malacostracous Crusticeans from Malaysia. 261
‘on the carapace. his groove is less conspicuous in Dr. de
Man’s examples according to their smaller size. In the Sara-
wak form also the tooth at the end of the posterior margin,
and the three teeth on the anterior margin of the carpus of
the chelipedes, are reduced in size; in fact, the foremost
tooth on the anterior margin, and the one just behind it, are
only just visible. These distinctions [I consider to be, with-
out doubt, due to age only.
Length 8°5 millim., breadth 8:25.
MACRURA.
XX. Genus CAuLiANassa, Leach.
29. Callianassa Martenst, Miers. (Pl. XII. figs. 4, 4 a.)
Callianassa Martensi, Miers, P. Z. 8. Lond. (1884) p. 13, pl. i. fig. 1;
de Man, Arch. f. Naturg. Jahrg. 53, Bd. i. p. 482, pl. xxi. fig. 1
(1887).
Distr. Mauritius, Amboina.
No. yxvi. Jab. Buntal.
Amale. This specimen, though rather damaged—having,
in particular, lost .both chelipedes—I believe to be a repre-
sentative of Mr. Miers’s species. 1t exhibits certain differences
on a comparison with the type specimen, but these seem to be
explained by the greater size of this example, 7. e., they are
differences of age. The most striking difference is in the
frontal region; the rostrum is shorter than in the type, in
which latter it reaches to the middle of the cornee of the
eyes, while in this it barely reaches halfway to the cornee.
These cornez are themselves much reduced in size in propor-
tion to their peduncles, while the latter have become very
acutely angulated at their antero-internal angles, now ¢n front
of the cornee, so that the two peduncles form in front a short
blunt tooth in the median line. From this angle to their
antero-external angles the border is concave, there being
another tooth, smaller than the median one, at the latter
angle. ‘The whole peduncle is very flattened and the cornea
appears only as a small black spot just in front of the centre
of its upper surface. Besides this, the ischium and merus of
the third maxillipedes form a broad opercular plate, and the
dactyl is bent back on the greatly enlarged propodos, so as
to make this appendage subchelate. Both these points are
more prominently developed in the larger animals than in
Mr. Miers’s smaller one. In other respects these two forms
agree.
Pim.: Length from tip of rostrum to tip of telson about
81 millim.
262 Mr. W. F. Lanchester on some
XXI. Genus Senex, Pfeffer (= Panulirus, Gray).
30. Senea ornatus, Fabr.
Palinurus ornatus, Fabr. Suppl. Ent. Syst. p. 400 (1798); Hasw. Cat.
Austr. Crust. p. 171 (1882).
Senex ornatus, Ortmann, ¢. c. p. 34 (1892), ubi syn.
Distr. Kast Africa, Ceylon, Malay Archipelago, N. Aus-
tralia, Hongkong, Japan, Samoa.
No. xc. Hab. Pulau Satang.
A male of this very variable species. On the antennal
segment are four forwardly-directed teeth arranged in a
square, the two anterior being larger; between these latter,
but nearer to the left one than the right, is a small ‘ Neben-
dorn.” Ina larger specimen in the Museum collection, from
Natunas, there are two ‘‘Nebendornen” halfway between
each of the anterior and posterior teeth. The second and
third abdominal segments are marked with an interrupted
transverse hirsute furrow; the specimen from Natunas is
without these, but has a deep furrow on the first abdominal
segment, without hairs, as in this specimen.
Dim.: Base of eyes to posterior margin of carap. 37 millim.
XXII. Genus Atya, Leach.
31. Atya armata, A. M.-Edw.
Ee A. M.-Edw. Ann. Soc. Entom. France, ser: 4, t. iv, p. 149
4).
? yrs aes de Haan, Crust. Jap. p. 186, pl. O (1889).
Age ee de Man, Weber’s Zool. Ergebn. p. 357, pl. xxi. fig. 20
( :
Distr. Malay Archipelago, Samoa, New Caledonia.
No. uxxx. Hab. Simanggang.
A male, which is certainly identical with A. armata.
Whether it is the same as de Haan’s species, as Mr. Miers
seems to think (Ann. & Mag. N. H. (5) v. p. 382), is rather
more difficult to determine, for de Haan’s description is too
brief. I have compared the mouth-parts of this specimen
with de Haan’s figures (vide pl. O), but the latter are too
diagrammatic for an accurate comparison; as far as I can
determine, however, the two are essentially similar.
Dim. : Length from orbital margin to tip of telson
50 millim.; length from orbital margin to posterior margin
of carap. 15°5.
XXIII. Genus ALPHEus, Fabr.
32, Alpheus gracilipes ?, Stimpson.
Alpheus gracilipes, Stm. Proc. Ac. Nat. Sci. Philad. p. 100 (1860) ;
Sp. Bate, ‘ Challenger ’ Macrura, p. 561, pl. ci. fig. 3 (1888) ; Ortmann,
op. cit. Vv. p. 488 (1891).
Malacostracous Crustaceans from Malaysia. 263
Distr. Zanzibar, S.W. Japan, Tahiti, Samoa, Bass Strait.
No. coxxu. Hab. Singapore.
A male. The chelipedes are lost, so that I cannot be quite
certain of its identity with Stimpson’s species.
Dim.: Base of rostrum to posterior margin of carap. 5°25
millim. ; base of rostrum to tip of telson 13°5.
XXIV. Genus PALz&moN, Fabr.
33. Palemon carcinus, var. Lamarrei, M.-Edw.
Palemon Lamarret, M.-Edw. Hist. Nat. Crust. t.1i. p. 397 (1837); de
Haan, Crust. Jap. p. 171 (1839); Ortmann, ¢. c. p. 701, pl. xlvii.
fig. 2 (1891).
Palemon carcinus, Hend, Trans. Linn. Soc., 2nd ser. Zool. vol. v. p. 441
(1893).
Distr. India, Malay Archipelago, Ecuador, Amazon.
No. xxviul. Hab. Kuching, freshwater.
Dental formula 42, and there is a wide gap between teeth
8-9 above. The rostrum exceeds the scaphocerite by one
fourth of its own length, while the second pair of pereiopods
exceeds the scaphocerite by half the carpus and the hand.
The carpus and hand are covered with minute teeth at
moderately wide intervals.
Dim.: Orbital margin to posterior margin of carap. 36°5
millim. ; length of second pair of legs 105, of merus 18, of
carpus 25, of palm and fingers 42.
XXV. Genus Penzus, Fabr.
34. Peneeus velutinus, Dana.
Peneus velutinus, Dana, U.S, Expl. Exp., Crust. p. 604, pl. xl. fig. 4
(1852); Sp. Bate. Chall. Macr. p. 258, pl. xxxiii. fig. 1 (1888) ;
Ortmann, ¢. c. p. 452, pl. xxxvi. fig. 6 (1891); Henderson, Trans.
Linn. Soc., 2nd ser. Zool. vol. v. p. 449 (1893).
Distr. Red Sea, Mauritius, Malay Archipelago, India,
Japan, N. & W. Australia, Sandwich Is.
No. cxxx. Hab. Singapore.
A male. The tip of the telson is broken off in this speci-
men, but there is no doubt that it belongs to this species;
the form of the male appendage is very characteristic.
Dental formula $; the peduncle of the antennules just
reaches the tip of the scaphocerite.
Dim.: Length from orbital margin to posterior margin of
carap. 9°5 millim,
35. Penaeus indicus, M.-Edw.
Peneus indicus, M.-Edw. Hist. Nat. Crust, t. ii. p. 415 (1837) ;
264 On Malacostracous Crustaceans from Malaysia.
Sp. Bate, Chall. Macr. p. 248, pl. xxxiii. fig. 2 (1888) ; Hend. Trans.
Linn. Soce., 2nd ser. Zool. vol. v. p. 447 (1395).
Peneus merguiensis, de Man, Merg. Crust. p. 287, pl. xviii. fig. 8
(1888).
Distr. India and Malay Archipelago.
No. uxxy. Hab. Buntal.
A female. Dental formula 8; rostrum just overreaches
the scaphocerite. The first tooth is situated behind the level
of the hepatic spine; antennular peduncle shorter than
scaphocerite.
This specimen is certainly identical with Dr. de Man’s
species ; but I do not separate it from P. indicus because, on
a comparison of some examples of the latter in the Museum
collection with two of Dr. de Man’s specimens and my own,
I find them essentially similar in all respects but two—the
proximal half of the rostrum is raised into a very prominent
crest and its first tooth is situate behind the hepatic spine.
The first of these distinctions is certainly a very obvious one
and very striking ; the second less so. It is hard to decide
whether these characters are of specific importance; for
myself, should a larger series demonstrate their constancy, I
should still be inclined to consider P. merquiense as a variety
only, though a well-marked one, of P. indicus.
Dim.: Orbital margin to posterior margin of carap. 17
millim.
STOMATOPODA.
XXVI. Genus SqQuiLua, Fabr.
36. Squilla scorpio, Latr.
Squilla scorpio, Latr. Encyel. Méth. x. p. 472 (1825); Miers, Ann.
& Mag. Nat. Hist. ser. 5, vol. v. p. 18, pl. ii. fig. 7 (1880) ; Hend.
Trans. Linn. Soe., 2nd ser. Zool. vol. v. p. 453 (1893).
Distr. Madras, Singapore, N. Australia, Shanghai.
No. vir. Hab. Buntal.
A young female. Length about 24 inches.
37. Squitla raphidea, Fabr.
Squilla raphidea, Faby. Ent. Syst. Suppl. p. 416 (1798) ; Miers, Ann.
& Mag. Nat. Hist. ser. 5, vol. v. p. 27 (1880).
Squilla harpax, de Haan, Crust. Jap. p. 222, pl. li. fig. 1 (1839).
Distr. Zanzibar, Madras, Indian Ocean, Mergui Is., Singa-
pore, Borneo, China, Japan.
No. xu. Hab. Moratabas.
A female. Length about 4? inches.
On new Species of Histeride. 205
ISOPODA.
XX VII. Genus NErociLA, Leach.
38. Nerocila depressa, M.-Edw. (Pl. XII. fig. 5.)
Nerocila depressa, M.-Edw. Hist. Nat. Crust. t. iii. p, 254, pl. xxxi,
fig. 17 (1839); Schiddte & Meinert, Naturhist. Tidsskrift, Kréyer,
Raek. iii, Bd. 13, p. 15, pl. i. figs. 10-11,
Distr. Penang, Zamboango, Amoy.
No. xxx. Had. Buntal.
A female.
The figure given by Schiddte and Meinert exaggerates, I
think, the relative proportions of the telson, which appears to
be too short for its breadth (vide text, p. 16, where the
proportions are given as 8:7). The exterior branch of the
uropods, which is long and styliform, is pigmented with black,
and this pigmentation is continued up along the sides of the
abdomen, though it fades away on the sides of the thorax.
Dim. : Greatest length 17°5 millim., greatest breadth 11 ;
breadth of telson 3°5, length of telson 4°25.
EXPLANATION OF PLATE XII.
Fig. 1. Goniosoma roseum. Antero-lateral border showing reduplicated
first tooth.
Fig. 2. Potamon tridentatum, var. incertum. Antero-lateral border.
Fig. 3. Potamon bidiense.
Fig. 4. Callianassa Martensi. Frontal region and eyes. 4a, Third
maxillipede.
Fig. 5. Nerocila depressa, Abdomen and telson.
XXXV.—On new Species of Histeridee and Notices of others.
By G. Lewis, F.L.S.
{Plate X. ]
TuIs is the eighteenth paper of a series on the //ister‘de
published in this Magazine, and in these various memoirs I
have described about 360 new species, and in other publica-
tions, during the same period, I have noticed 150 more. In
and between the years 1884 and 1897 Herr J. Schmidt
described about 230 species, and after the year 1868, the date
of the Munich Catalogue, Marseul described about 185,
Mr. L. Casey 48, and other authors about 157 species. These
figures represent in all 1180 species, which, with those of the
Munich Catalogue, 1151, bring the present number of species
recognized up to 2281. I have a new Catalogue in MS.
which I hope to publish shortly after a few points of synonymy
Ann. & Mag. N. Hist, Ser, 7, Vol. vi. 18
266 Mr. G. Lewis on
are settled and the lineal arrangement of the genera somewhat
adjusted.
The genus Probolosternus founded here is a remarkable
one, and as three species of it have been recently discovered
(one in Guinea and two others in the French Congo), it is
highly probable that many more exist in Central Africa, and
the species also vary so much in size, which increases the
probability that intermediate forms exist. It is also likely,
from the torm of the sterna, that the species have the same
habits as those of Pachycrerus, which are known to follow
the xylophagous beetles, feeding on the euphorbize and other
African shrubs and trees.
There are two species of the genus ister noticed here
(H. Marshalli and H. augoniensis) which have the inner
subhumeral striae complete, and a reference to them will show
how essential it is at times to recognize the importance of the
basal oblique humeral stria of the elytra. Sometimes, although
rarely, the inner humeral stria is so similar to the first dorsal
stria that it is impossible to distinguish the one from the other
without noting their position in regard to it.
List of Species.
Hololepta dyak. *Phelister festivus, Lew.
syntexis. *Notolister sulcicollis, Lew.
*Placodes opacus, Lew. Hister monbasanus.
Apobletes parallelus. zambesius.
—— cavifrons. augoniensis.
macilentus. *—— Marshalli, Lew.
platessee. brahminius.
Liopygus chalcis. frontalis.
*Platylister mirabilis, Lew. *Coproxenus Marshalli, Lew.
* extrarius, Lew. Stictostix parra, Mars.
cathayi. Notoccelis satur.
enodis. Sitalia Severini, Lev.
Platysoma assamense. Homalopygus remex.
disparile. Terapus muricatus.
Pachycreerus chlorites.
Probolosternus africanus,
permundus.
—— minor.
*Baconia loricata, Lew.
Epitoxus coryceus.
Saprinus punctisternus,
—— castanipes, Curtis.
leesus.
Teretrius Braganzz.
Epiechinus Marseuli.
Hololepta dyak, sp. n.
Oblongo-ovalis, depressa, nigra, nitida; fronte impressa, mento
haud carinato; thorace lateribus sparse punctulato; elytris
striis dorsalibus brevissimis, i* appendice curvata; propygidio
* Species with an asterisk are not described in this paper, but are
figured in the Plate.
new Species of Histeride. 267
extus sparse punctato ; pygidio dense ocellato-punctato ; prosterno
lato; tibiis anticis 4-dentatis.
L. 10 mill. (absque mandibulis).
Oblong-oval, depressed, black and shining; the head
smooth, impressed between the mandibles; mentum in male
concave and smooth, not carinate: the thorax, lateral stria
complete and similar to that of 7. Baulny?, Mars.; there is
a band of fine, not conspicuous, punctures scattered along the
sides ; the anterior angle has a small emargination and close
to the emargination there is a deep oval, somewhat oblique,
fossette : the elytra, lateral furrow widest in the middle and
abbreviated a little before and behind; striee, first dorsal is
one fourth of the elytral length and has a short bent appendage,
second shorter, not quite half the length of the first : the pro-
pygidium is sparsely punctured on the outer edges, but not
at the base ; the pygidium is densely and ocellately punctate,
with the posterior border narrowly smooth; the prosternal
keel is broad and scarcely narrowed at or before the coxe.
This insect is very similar to H. Baulnyi, Mars., but it
differs from it in the thoracic auriculiform fovea being deeper,
more circular in outline, aud in its position being nearer the
angular emargination, the apical elytral stria is bent, not
straight, and the pygidium is distinctly ocellately punctured.
Hab. Borneo.
Hololepta syntevis, sp. n.
Oblongo-oyata, subdepressa, nigra, nitida; fronte inconspicue bi-
striata; pronoto impunctato, stria laterali in angulo antico desi-
nente ; elytris striis 1-2 brevibus, 3 vix conspicua; propygidio
apice bifoveolato, punctis sparsis late cincto.
L. 8 mill. (absque mandibulis).
Oblong-oval, depressed, black and shining; the forehead
smooth, with two very short, almost obsolete, transverse
strie, g mentum with a short but distinct carina; the
thorax is impunctate, marginal stria very fine, lateral stria
well marked, especially at the anterior angles, both strie
terminate behind the angle, male has no thoracic emargination
or fovea; the elytra, lateral fossette reaches the base, but it is
shortened before the apex, 1 dorsal stria is about a fourth of
the elytral length, 2 half the length of the first, 3 is just
visible as a rudiment, there is no appendage ; the propygidium
is bifoveolate at the apex, with a border of scattered punctures,
except between the foveee, where the surface is smooth; the
pygidium is rather closely punctured, the punctures varying
somewhat in size and leaving the posterior edge narrowly
16*
268 Mr. G. Lewis on
smooth ; the prosternum is not very wide between the coxe,
but at the base it widens out considerably and is somewhat
triangular (in this character it agrees with H. striatidera,
Mars.) ; the mesosternum has a short curved sulcus at each
anterior angle; the anterior tibiew are 4-dentate, the two apical
teeth are robust and near together.
Hab. 8. Thomas I, Gulf of Guinea (Mocquerys).
Apobletes parallelus, sp. n.
Oblongus, planatus, parallelus, brunneus, nitidus ; fronte punctulata,
stria transversa utrinque interrupta; pronoto, stria marginali
antice late interrupta ; elytris striis 1-2 integris, 3 late inter-
rupta, humerali externa completa, interna in medio separata ;
prosterno lobo antico immarginato.
L. 33 mill.
Oblong, flat, parallel at the sides, brown and shining; the
head slightly impressed anteriorly, punctulate, also the epi-
stoma, striate over the eyes, transverse stria rather wide and
shallow and confined to the frontal impression; the thorax,
lateral stria continues round the anterior angle to a point
behind the eye and it is equally strong there as at the sides,
at the base on either side opposite the second dorsal stria is a
shallow, somewhat transverse fovea, in lieu of a scutellar
fovea there is a minute tubercle, along the thoracic border,
but well away from the edge, is a longitudinal cluster of
punctures; the elytra, strie, outer humeral complete, inner
humeral separated in the middle, each part extending to a
point transversely opposite to each other, oblique basal rather
long and very fine, dorsal 1-2 complete, 3 widely interrupted
in the middle, the apical edge is narrowly rugosely punctured ;
the propygidium is punctured throughout, in the centre and
along the posterior edge the punctures are fine, but otherwise
the punctures are shallow, irregular, and imbricate, in no part
are they dense; the pygidium is flat, surface more densely
punctate than the propygidium, and the punctures appear
ocellate under the microscope ; the prosternum, anterior lobe
immarginate anteriorly, but behind the middle it is striate on
either side, the surface of it and the keel is finely punctulate ;
the mesosternum is very distinctly bisinuous, so that the
middle of the anterior edge is slightly acuminate, on either
side is a curved marginal sulcus, but there is no marginal
stiia; the anterior tibie are 3—4-dentate.
This species is more parallel than any other known species
and its specific characters generally do not correspond with
any other.
Hal. Sumatra.
new Species of Histeride. 269
Apobletes cavifrons.
Ovalis, planatus, niger, nitidus; fronte excavata, stria transversa
nulla, mandibulis distincte punctatis ; pronoto lateribus rugose
punctato, stria marginali post caput interrupta; elytris striis
1-3 integris, 4 apicali; pygidio dense et fortiter punctato.
L. 33-3? mill.
Oval, flat, black and shining; the head, forehead excavated
and evenly, not closely, punctured, the stria over the eyes is
strong and bent, there is no transverse stria, but the suture
dividing the forehead and the epistoma is straight and clear ;
the labrum is punctured like the forehead and is minutely
strigose at the base; the mandibles are densely and rather
rugosely punctured; the thorax, marginal stria strong laterally,
but fine behind the head, and it does not meet in the middle,
lateral borders are rugosely and densely punctured, especially
behind the anterior angles; the elytra, the outer humeral
stria is wide, basal, and dimidiate, inner humeral is wanting,
oblique basal fine but clear, 1-3 dorsal complete, 4 apical
and very short. ‘he elytra, striz are almost similar to those
of A. Schaumz, Mars., and also closely similar to those of
A. tener, Mars., but in the last the third dorsal stria is inter-
rupted in the middle. The propygidium is somewhat impressed
and punctate on either side, with the middle and _ posterior
border smooth; the pygidium is deeply punctate, punctures
clear and of equal size, set close together, the posterior margin
is slightly elevated; the prosternum, the anterior lobe is
marginate, and in this and other respects it is similar to that
in A. tener; the mesosternum also resembles that of A. tener,
but the anterior border is more distinctly bisinuous ; the
anterior tibize are 4—5-dentate.
This is a very distinct species, but resembles the species
named superficially.
Hab. Patkai Mountains, Assam (Doherty).
Apobletes macilentus, sp. n.
Oblongo-ovatus, parallelus, depressus ; fronte sat dense punctulata ;
pronoto lateribus punctulato, stria integra; elytris striis 1-3
integris, 4 apicali ; pygidio subconvexo, dense punctato.
L, 22-3 mill.
A. macilentus differs from A. nirvana, Lew., in being
smaller, more depressed and more parallel laterally, the head
and thoracic borders are much less distinctly punctured, the
elytral strize are finer and nearer to each other, the fourth stria
is short and apical and the fifth is either wholly wanting or
270 Mr. G. Lewis on
represented by a very rudimentary line or one or two pune-
tures. The pygidium is less convex and somewhat less
coarsely punctured. Beneath the punctuation in A. macilentus
is more fine, especially as it is visible on the anterior lobe of
the prosternum, and the posterior lobe of the prosternum is
wider than in A. nirvana.
This is evidently the species referred to by Herr J. Schmidt
as A. nirvana, Lew., in the Ann. Mus. Genova, ser. 2, xvii.
p- 286 (1897).
Hab, Si-Rambe, Sumatra (Modigliani). I possess six
examples with the label of the Genoa Museum attached to
them.
Apobletes platesse, sp. n.
Oblongus, parallelus, planatus, piceo-niger, nitidus ; fronte punctu-
lata, stria transversa utrinque interrupta ; pronoto stria marginali
antice interrupta; elytris striis 1-3 integris, 4 basi abbreviata ;
propygidio utrinque imbricato-punctato ; pygidio parum dense et
subocellato punctato.
L. 23 mill.
Oblong, parallel at the sides, flat, pitchy black and shining ;
the head impressed anteriorly, distinctly but not densely
punctulate, lateral stria strong and does not pass beyond the
eye, transverse stria very feebly sinuous and on either side
rather widely separated from the lateral stria; the thorax,
lateral stria well marked, but fine anteriorly and ceasing
behind the eyes, lateral border with a few small scattered
punctures, base with a small scutellar fovea ; the elytra, outer
humeral stria complete, inner wanting, oblique basal fine and
relatively rather long, 1 dorsal complete but not quite so
long apically as the second, 2-3 complete, 4 shortened by
about one fifth at the base, all the striae are nearly parallel
one to another, 5 obsolete or consisting of 2 or 3 punctures,
sutural absent ; the propygidium has a lateral cluster of rather
large, shallow, imbricate punctures, with a few very fine
points in the centre; the pygidium is somewhat densely
punctured, punctures somewhat ocellate and each one clearly
separate from another; the prosternum widens out broadly
before the coxee and the anterior lobe is margined at the sides
only by a fine stria; the mesosternum is anteriorly bisinuous,
but the marginal stria neither meets in front nor does it con-
tinue along the sides; the anterior tibiee are 3—4-dentate.
Hab. Nguela, Usambara.
LInopygus chalcis, sp. n.
Ovatus, depressus, piceo-brunneus, nitidus ; fronte punctulata, stria
new Species of Histeride. 271
transversa utrinque interrupta; pronoto lateribus inconspicue
punctulato, stria integra; elytris striis 1-3 integris, 4-5 api-
calibus ; propygidio transversim punctato ; prosterno haud striato ;
mesosterno lato, distincte marginato ; tibiis anticis 4-5-dentatis.
L. 3 mill.
Oval, depressed, pitchy brown and shining; the head
punctulate, but not closely, slightly impressed behind the
transverse stria, which is straight with the ends on either side
turned backwards, and it is not connected with the lateral
strie, which are bent and do not extend beyond the eye; the
thorax, surface microscopically punctured, punctures more
visible along the lateral borders, marginal stria strong later-
ally and continued behind the head, the stria leaves the
margin at the angle, cutting off a triangular space, it then
passes along the margin until it passes the eye, when it again
leaves the margin and behind the neck leaves a fairly wide
border between it and the edge, there is no scutellar fovea.
The elytra, there are no humeral striz except the oblique
basal, which is very fine, dorsal stria, 1 a little abbreviated
at the apex, 2 complete and turning a little away from the
first at the base, 3 complete but extremely fine at the base,
the interstice between it and the second is of the same width
throughout, 4 apical and one third of the elytral length, 5
shorter with a wider interstice, sutural wanting. ‘The pro-
pygidium has rather large shallow punctures; the pygidium
rather deeply bifoveolate and smooth; the prosternum, the
anterior lobe is broad, with a marginal stria, which is, however,
interrupted at the suture before the keel, keel without striz ;
the mesosternum is wide, very feebly sinuous anteriorly, and
clearly marginate; the metasternum has a shallow median
furrow ; the anterior tibie are 4—5-dentate.
This species is somewhat similar to L. Gestror, Lew., but
the latter is less oval, the thorax more distinctly punctured
laterally, the apical margin of the elytra narrowly punctulate,
the second dorsal stria widens out at the base from the first
in a similar manner, and the second and third are similarly
parallel to each other, but the two apical striz are shorter and
the fifth is more distant from the suture. In ZL. chalcis the
fovee in the pygidium are nearly round, in L. Gestrod they
are transverse.
Hab. Sipora, Mentawei Islands, Sereinu, 5th June, 1894
(Modigliani). ‘Two examples, and I have a third labelled
“Perak” by Herr J. Schmidt.
Platylister cathayt, sp. n.
Oblongo-ovatus, subdepressus, niger, nitidus ; fronte impressa, stria
o42 Mr. G. Lewis on
fere recta; pronoto stria integra; elytris striis 1-3 integris,
4 valde abbreviata, 5 dimidiata, suturali brevissima vel obsoleta ;
pygidio margine anguste levi haud conspicue elevato.
L, 5-54 mill.
Oblong-oval, rather depressed, black and shining; the
head impressed anteriorly, transverse stria feebly sinuous,
labrum transverse and narrow ; the thorax widest at the base,
anterior angles moderately depressed, marginal stria com-
plete; the elytra, strie 1-3 complete, 4 apical, short, and
sometimes broken, 5 apical and dimidiate, sutural short,
much abbreviated apically, and not reaching the middle of
the elytron ; the propygidium punctured, the punctures vary in
size and are not closely set; the pygidium, the punctures are
larger, nearly equal in size, and much more closely and
evenly set, the outer margin is smooth but scarcely elevated ;
the prosternum, anterior lobe striate, stria arched and only
marginal anteriorly; the mesosternum, anteriorly rather
widely but not deeply emarginate, stria complete ; the
auterior tibie rather strongly 4-dentate.
Differs chiefly from P. suturalis, Lew., in its labrum being
much more transverse, in its shorter and less defined sutural
stria, and in the posterior margin of the pygidium being
smooth and scarcely elevated.
Hab. Kuatun, China. Three examples.
Platylister enodis, sp. n.
P. abrupti persimilis, sed differt pygidio fere impunctato et lobo
prosterni marginato.
L. 53 mill.
This species is extremely like P. abruptus, Er., the elytral
strie are exactly similar, viz. 1 and 2 complete, 3 interrupted
near the middle, but the thorax is rather more transverse, the
lateral outline rather less parallel, and there is a conspicuous
fovea before the scutellum. The propygidium is smooth, with
a shallow impression on either side near the posterior edge, in
which there are a few small punctures, and the pygidium on
either side near its base has a small cluster of punctures, but
it is otherwise smooth and flat. In P. enodts the stria of the
prosternal anterior lobe is marginal; in P. abruptus it is
an independent arched stria not following the margin; in
P. enodis the mesosternum has a slightly wider emargination
and the marginal stria is more marked than in P. abruptus.
Hab. Mailu, British New Guinea (Anthony, July 1897).
new Species of Histeride. 273
Platysoma assamense, sp. n.
Oblongo-ovatum, depressum, nigrum, nitidum ; fronte concava,
punctata, stria integra; pronoto lateribus dense punctato ; elytris
striis 1-3 integris, 4 dimidiata, 5 apicali, subhumerali externa
brevi, interna punctiformis ; pygidio convexo, grosse et dense
punctato,
L. 33 mill.
Oblong-oval, rather flat, black and shining, legs somewhat
reddish; the head, forehead and epistoma concave, clearly
and evenly, not densely, punctate, stria complete, transverse
part nearly straight; the thorax, marginal stria complete,
narrowly canaliculate at the sides, fine behind the head, and
behind the neck it deviates from the edge, the disc is micro-
scopically and sparsely punctured, but the lateral borders are
widely and conspicuously punctate and many of the punctures
are much larger than those on the head. ‘The elytra, lateral
margins clearly but finely punctulate, striz, external humeral
short and relatively deep and is seen just below the shoulder,
the inner humeral is punctiform and evanescent at both ends,
1-3 dorsal complete, 4 dimidiate or faintly traceable beyond
the middle, 5 apical and nearly one third of the elytral length,
apical margin punctured and very minutely strigose. The
propygidium and pygidium are punctate, punctures round and
deep, closely set together on the pygidium, less regularly on
the propygidium, the pygidium is convex ; the prosternum,
anterior lobe is marginate and clearly punctured over the
whole of its surface, ‘at the base beyond where the marginal
stria terminates are some large shallow punctures, and on
either side of the keel before the coxee are similar punctures ;
the mesosternum anteriorly is widely sinuous, the sinuosity
extending almost from angle to angle, anteriorly the mar-
ginal stria is fine and close to the edge, the under surface
generally is finely punctulate; the anterior tibie are 4-
dentate.
P. novum, Lew., and rimarium, Mars., resemble this
species ; all have a convex pygidium and the thorax similarly
punctured.
Hab. Khasia Hills, Assam.
Platysoma disparile, sp. n.
Oblongo-ovale, subdepressum, piceum, nitidum ; fronte leviter
impressa, stria transversa utrinque interrupta; pronoto impunc-
tato; elytris striis 1-2 integris, 3 late interrupta; pygidio
punctato; tibiis anticis obtuse dentatis.
L, 3% mill.
274 Mr. G. Lewis on
Oblong-oval, rather depressed, piceous, shining ; the head
faintly impressed anteriorly, transverse stria not well-marked
and interrupted on either side; the thorax gradually narrowed
from the base to the anterior angles, marginal stria extremely
fine and continued round the angle to a point behind the eye,
lateral stria almost reaches the marginal behind the anterior
angles, surface smooth, with a very small anti-scutellar punc-
ture; the elytra, humeral stria wanting, save the oblique
basal which is very fine, dorsal 1-2 complete and parallel one
to another, each turning in a little at the base, 3 widely
interrupted in the middle and extremely fine, only seen in
certain lights, at the base ; the propygidium is transversely and
irregularly punctured ; the pygidium is more regularly punc-
tate, but the basal edge is narrowly smooth, the sides are a
little raised; the prosternum, anterior lobe marginate, keel
rather wide; the mesosternum with a well-defined marginal
stria and anteriorly widely sinuous; the anterior tibia have
three large blunt teeth, resembling those of Hololepta ob-
tusipes, Mars., and a rudiment of a fourth at the base.
There is no known species of Platysoma to which this can
well be compared, but it may be placed near P. sincerum,
Sch.
Hab. Lombok. “ Sapit, 2000 feet, April 1896” (H. Fruh-
storfer) .
Pachycrerus chlorites, sp. n.
Oblongo-ovatus, subconyexus, viridi-cyaneus, nitens; fronte im-
pressa, stria interrupta; pronoto stria marginali integra; elytris
striis 1-3 integris, ceteris nullis; pygidio punctato; prosterno
bistriato ; mesosterno valde prominulo, stria marginali antice
interrupta ; tibiis anticis 5-dentatis.
L. 54 mill.
Oblong-oval, rather convex, greenish blue, shining, beneath
black ; the head rather feebly impressed anteriorly, lateral
stria narrowly interrupted over the eyes, oblique anteriorly
and not joining in front, surface with some very small
scattered irregular punctures; the thorax, marginal stria
complete but fine behind the head and close to the margin,
surface with a few punctures chiefly at or just behind the
anterior angles; the elytra, strie, outer humeral complete and
very distinct, inner basal and dimidiate, oblique basal very
fine, dorsal 1-2 complete and not very deeply impressed,
3 finer and although almost complete has a tendency to be
evanescent behind the middle, 4, 5, and sutural are wanting ;
the propygidium is faintly impressed on either side of the
posterior edge, surface distinctly but not closely punctured ;
new Species of Histeride. 275
the pygidium similarly but rather more evenly punctured ;
the prosternum is bistriate, strie rather fine, scarcely divergent
anteriorly, but widen out a little behind, keel, anterior lobe
smooth, a few fine punctures only are seen under the micro-
scope ; the mesosternum is markedly prominent anteriorly and
has a lateral well-defined marginal stria, but it is not con-
tinued along the projection; the anterior tibiz are 5-dentate.
This species should be placed close to P. eleguns, Lew.,
both having a very prominent mesosternum, but superficially
it resembles P. chalybeus in colour and outline, being in the
first character somewhat bluer and in the second somewhat
more oval. It is remarkable in having only three dorsal
striae.
Hab. 8. Thomas I., Gulf of Guinea (Mocquerys, 1900).
One example.
PROBOLOSTERNUS, gen. nov.
Body oval, rather convex ; head retractile, forehead without
a transverse stria, labrum transverse, mandibles acute at the
tips ; antenne inserted below the frontal margin, club oval,
and when at rest is received into a fovea under the thoracic
angle like that in the genus Pachycrerus; the thorax is
transverse, arched at the sides, gradually narrowed to the
anterior angle, anterior angles somewhat acute, marginal
stria extremely fine and not passing behind the head, lateral
stria complete close to the edge, leaving only a narrow lateral
rim; the elytra, epipleure finely striate, the interstice be-
tween the inner humeral and first dorsal stria is wide at the
base; the propygidium is very large, hexagonal, and convex ;
the pygidium is also rather large and semicircular in outline
behind, but being deflected it is invisible from above; the
prosternum, the keel is narrow, bistriate, and incised at the
base; the mesosternum is very prominent anteriorly, and
from its base is arched at its sides and is sharply pointed
anteriorly where it fits into the base of the narrow prosternal
keel, the marginal stria is more or less fine but entire; the
tibie are all dilated, the outer edge of the anterior pair is
bowed and multidentate ; the tarsi are short with two claws
and the tarsal grooves are nearly straight. The curious form
of the mesosternum is shown in fig. 1 (p. 277), which is
drawn from an example of P. permundus.
Probolosternus africanus, sp. n.
Ovalis, parum conyexus, niger, nitidus, supra leviter punctulatus ;
elytris striis dorsalibus 1-3 integris, 4 antice interrupta, 5 apicali,
276 Mr. G. Lewis on
suturali basi abbreviata; prosterno bistriato; mesosterno stria
marginali antice tenuissime impressa; tibiis anticis multiden-
ticulatis.
L. 7 mill.
Oval, rather convex, black and shining; the head rather
finely and rather closely punctulate, epistoma transverse and
flat, lateral stria fine and not passing behind the epistoma ;
the thorax, lateral stria close to the margin and continued
behind the head, surface clearly punctulate, but the punctures
are rather less dense than those of the head. The elytra,
surface punctulate like that of the thorax ; striz, outer humeral
fine and complete, inner also complete with its outer edge at
the base a little raised, 1-3 dorsal complete with the outer
edges somewhat raised, especially those of the first and second
strie, 4 fine and shortened before the base, 5 apical eva-
nescent and punctiform, sutural fine and evanescent before
the base and in the middle it bends a little away from the
suture. The propygidium and pygidium are much more dis-
tinctly punctured than the head; the prosternum is bistriate,
strie are fine and gradually diverge posteriorly along the
narrow keel; the mesosternum is acutely pointed in front
fitting into the incision in the prosternum ; there is a very
fine marginal stria only to be seen in front in certain lights ;
the metasternum has a well-marked lateral stria and the
suture is shown by a straight stria ; the tibie are all dilated,
anterior pair bowed on the outer edge, along which are small,
evenly set denticulations (about 16 or 17 in number) ; inter-
mediate and posterior tibiz are spinose, the spines are some-
what irregular and not closely set together.
The size and the incomplete sutural stria will distinguish
this species from the two following.
Hab. W. Guinea.
Probolosternus permundus, sp.n. (Pl. X. fig. 4.)
Ovalis, parum convexus, niger, nitidus; fronte punctulata, antice
leviter impressa ; elytris striis 1-4 suturalique integris, 4 cum
suturali arcuatim connexa, 5 antice abbreviata; prosterno bi-
striato, striis basi divergentibus; mesosterno stria marginali
integra.
L. 423 mill.
Oval, rather convex, black and shining; the head lightly
impressed behind the epistoma, surface punctulate, lateral
stria fine; the thorax, lateral stria continued behind the head,
surface punctulate. The elytra sparsely punctulate especially
in the apical region; striez, outer humeral very fine and
new Species of Histeride. 277
complete, inner humeral and 1-3 dorsal strong, complete, and
similar in having the outer edge somewhat elevated, the third
dorsal is rather less strong than the second, 4 dorsal and the
sutural are fine and almost complete, being joined anteriorly,
but they are slightly shortened apically, 5 dorsal is apical,
Fig. 1.
iw)
Probolosternus permundus, Lew.
fine, and reaches just beyond the middle. ‘The propygidium
and pygidium are clearly punctulate; the prosternum is bi-
striate, strie divergent posteriorly ; the mesosternum is like
that of the last species, except that the fine marginal stria is
conspicuously complete anteriorly ; the metasternum is striate
laterally and the suture is marked by a fine transverse stria ;
the legs are similar to those of P. africanus.
Hab. Kuilu, French Congo (Mocquerys, 1892).
Probolosternus minor, sp. n.
Ovalis, parum convexus, niger, nitidus; fronte punctulata, stria
laterali perspicue angulata ; pronoto stria laterali integra; elytris
striis 1-4 suturalique integris, 4 cum suturali arcuatim connexa,
5 basi abbreviata; prosterno bistriato, striis utrinque divergenti-
bus ; mesosterno stria marginali antice integra,
L. 3 mill.
Oval, rather convex, black and shining; the head punctu-
late, punctures varying in size, with very minute points
intermixed, lateral stria rather strong and very distinctly
angulate in the middle, area behind the epistoma is very
faintly impressed ; the thorax punctured like the head, except
along the base, where the punctures are larger, lateral stria
entire and feebly crenulate behind the neck; the elytra, the
striz resemble those of P. permundus, except that the fourth
and sutural are not shortened at the apex, the surface punctu-
ation is only seen under the microscope and consists of
extremely fine points ; the propygidium is margined laterally
with a very fine stria, surface clearly not closely punctate,
with very fine points intermixed ; the pygidium is curiously
278 Mr. G. Lewis on
punctured, the punctures are very irregular, some crescent-
shaped, some semicircular but not close together, and these
again are mixed with fine points, the apex is margined by a
fine stria; the prosternum is bistriate, the striz are divergent
before and behind; the mesosternum is outwardly margined
by a rather fine stria complete anteriorly, the suture between
it and the metasternum is marked by a straight crenulate
stria, the latter segment has a few imbricate punctures along
its base, the punctures being most numerous at the sides ; the
legs are similar to those of the last species.
The small size, prosternal striae divergent at both ends,
and the crenulate metasternal stria are characters which easily
separate this species from the last. ‘The stria on the pygidium
1s not very easy to see.
Hab. Kuilu, French Congo (Mocquerys, 1892).
HPITOXUS, gen. nov.
Body oval or suborbicular, convex; the head retractile,
frontal stria complete; the thorax with an impression before
the scutellum more or less distinctly arcuate or biareuate ;
the elytra with six dorsal striz, the fourth and sutural some-
times joining at the base; the propygidium is transverse, the
pygidium semicircular and declined, the prosternum bistriate,
and the anterior tibie multispinose. ‘The other characters
agree with those of Phelister. Type Phelister circultfrons,
Mars.
There are five species known which should be placed in
this genus: three have the sutural dorsal strize shortened,
viz. Lhelister circulifrons, Mars., hilarulus, Lew., and the
species described below; while P. breviusculus, Fahr., and
nitidus, Lew., have the fourth and sutural dorsal striae com-
plete and joined at the base. Of P. circulifrons and bre-
viusculus there are excellent figures in Marsuel’s monograph.
Epitoxvus coryceus, sp. n.
Ovalis, parum convexus, Diger, nitidus ; fronte stria integra, antice
fere recta; pronoto lateribus punctato, ante scutellum_bi-
arcuatim impresso; elytris striis 1-3 integris, 4-5 brevibus,
suturali paululum obliqua, dimidiata; pygidio punctulato ; pro-
sterno bistriato ; mesosterno marginato.
L. 43 mill.
Oval, rather convex, black and shining; the head, fore-
head smooth, stria complete, not strong, feebly sinuous behind
the mandibles, almost straight anteriorly; the thorax, mar-
ginal stria complete, lateral border with a band of punctures,
new Species of Histeride. 279
punctures most conspicuous behind the anterior angles, fine
at the base, on the basal edge before the scutellum are two
arched impressions free of punctures ; the elytra, outer humeral
stria apical and scarcely dimidiate, inner wanting, oblique
basal very fine and close to the first dorsal, dorsal 1-3 com-
plete, 4 apical but reaching the middle and it has a short
basal appendage, 5 apical very short and somewhat puncti-
form, sutural apical but reaching quite to the middle and
from the apex the striz diverge from the suture, its base is in-
dicated by a puncture; the propygidium is distinctly punctured
at the sides and minute punctures are intermixed with the
larger ones, the minute punctures alone extend over the disc ;
the pygidium is similarly but less conspicuously punctured,
and on either side near the middle of the margin is a fovea;
the prosternum is bistriate, the striz do not meet anteriorly
and they diverge posteriorly from the coxe to the base; the
mesosternum is bisinuous anteriorly and narrowly transverse,
it posterior limit is marked by a bent not crenulate stria.
Differs from circulifrons, Mars., in size, more oval outline,
less convex form above, frontal stria less strong and not
semicircular, and the thoracic scutellar impression is distinctly
biarcuate and free of punctures.
Hab. Kuilu, French Congo (Mocquerys, 1892).
flister mombasanus, sp. n.
Oblongo-ovalis, convexus, niger, nitidus ; fronte punctata, stria
integra antice recta; pronoto lateribus bistriato; elytris striis
1-3 integris, 4 apicali dimidiata, 5 brevissima, suturali utrinque
abbreviata ; propygidio pygidioque dense punctatis; mesosterno
antice marginato ; tibiis antice 3-dentatis.
L. 7 mill.
Oblong-oval, convex, black and shining ; the head punctu-
late at the base, rather densely and rugosely punctate behind
the stria, stria complete and straight anteriorly, epistoma
and mandibles also densely and rugosely punctured, the last
are not marginate; the thorax, lateral marginal stria ex-
tremely fine, external lateral ceasing behind the eye, interstice
narrow, internal, feebly sinuous laterally, complete behind
the head, and distinctly sinuous behind the eyes, interstice
rather wide and punctulate, with some irregular strize or
rugosities at and behind the anterior angle, the sides are
microscopically punctulate; the elytra, strie, external
humeral wanting, internal apical and dimidiate, oblique basal
extends outwardly beyond the inner humeral, dorsal striz
1-3 complete and rather wide, best marked on the edge,
280 Mr, G. Lewis on
interstices distinctly punctulate, 4 much less in width, apical
and dimidiate, 5 represented by a short apical line or puncture,
sutural shortened a little before the apex, extending just
beyond the middle and slightly turning from the suture at
either end; the propygidium and pygidium are wholly and
coarsely punctured, but without impressions ; the prosternum,
the lobe is partly punctured and has a bordering stria, which
is close to the edge at the apex; the mesosternum is very
feebly sinuous anteriorly and the marginal stria is complete ;
the anterior tibiz are punctured and 3-dentate.
This species belongs to the HH. longicollis, Mars., group.
Hab. Makeré, Mombasa.
Hister zambesius, sp. n.
Ovalis, convexus, niger, nitidus; fronte impunctata, stria integra;
pronoto lateribus bistriato ; elytris striis 1-4 integris, 5 basi
abbreviata, suturali vix abbreviata; propygidio pygidioque dense
punctatis ; tibiis anticis 3-dentatis.
L. 7-74 mill.
Oval, convex, black and shining ; the head impunctate, stria
strong and complete, anteriorly nearly straight, labrum trans-
verse, mandibles impunctate; the thorax, marginal stria
very fine, outer lateral stria terminates after passing the
anterior angle, feebly sinuous in the middle, outer interstice
impunctate, inner lateral straight and continued behind the
head, interstice roughly punctate behind the anterior angle
and there is a small cluster of punctures anteriorly just within
the inner thoracic stria, the scutellar fovea is fine and linear ;
the elytra, striz, outer humeral is represented by a very short
median sulcus, inner humeral is complete, oblique basal
inconspicuous, 1-4 dorsal complete, somewhat wide, with all
the interstices more or less irregularly punctured, especially
near the apices, 5 stria is straight and shortened before the
base, sutural is bent and a little longer; the propygidium
and pygidium are evenly and closely punctured, both are
without impressions ; the sterna resemble those in the last
species ; the anterior tibiz are 3-dentate.
This, like the last species, also belongs to the H. longi-
collis group.
Hab. N. Batoka, Zambesi: December 1895.
Hister angoniensis, sp. n.
Suborbiculatus, convexus, niger, nitidus ; fronte biimpressa, punctu-
lata, stria carinata antice recta ; pronoto striis 2 lateralibus
new Species of Histerides. 281
validis, integris; elytris striis 1-3 et subhumerali interna
similiter integris; mesosterno in medio foveolato ; tibiis anticis
valde 3-dentatis.
L. 52 mill.
Suborbicular, convex, black and shining ; the head, surface
uneven and punctulate, with two impressions behind the
frontal stria, frontal stria cariniform and straight anteriorly,
epistoma transversely concave, labrum robustly carinate,
mandibles concave above with the rims elevated; the thorax
gradually narrowed from base to anterior angles, angles
depressed and somewhat acute, marginal stria fine, somewhat
carinate, especially behind the anterior angles, and ceasing
anteriorly behind the eyes, there are two lateral stria parallel
to one another, both strong, with the interstices a little raised,
the outer lateral ceases after passing the anterior angle, the
inner continues as far behind the head as the marginal
stria, and where the two ends overlap they enclose a narrow
space in which there is a small but well-marked puncture.
The elytra are wider than the thorax and have each four
strong and complete strie with crenulate edges; the outer
of these striz is evidently the inner humeral stria, because
the short oblique basal stria is seen between it and the next
stria, the four striz all widen out a little at the base, stria 4
is indicated by a short apical line only, 5 wanting, sutural
faint, not reaching the base but extending to the middle. The
propygidium and pygidium are rather densely punctured, but
the apex of the latter is narrowly smooth; the prosternum,
lobe punctured, the lateral strie meet at the apex; the
mesosternum is truncate, the marginal stria complete and in
the middle there is a conspicuous oval fovea; the anterior
tibiz are 3-dentate, the apical tooth being very broad and
prominent.
This is a remarkable species, both from its general form
and also for the inner subhumeral stria being similar to the
first dorsal stria. Its position in the genus should be near
HI. ignavus, Mars., and H. Marshalli, Lew. The latter
figured here (Pl. X. fig. 8) also has an inner subhumeral
stria similarly formed to the first dorsal.
Hab. Zomba, Upper Shire River, Angoni, 3000 feet:
10th Dec., 1895 (Dr. P. Rendell).
Hister brahminitus, sp. n.
H. pteromalo simillimus, sed major, elytris striis subhumerali
interna vix dimidiata, 1-2 dorsalibus integris, 3 postice abbre-
viata, et mesosterno antice distincte emarginato,
L, 10 mill.
Ann. d Mag. N. Hist. Ser. 7. Vol. vi. 19
282 Mr. G. Lewis on
This species very closely resembles H. pteromalus, Mars.,
but it is considerably larger, the external thoracic lateral
stria is longer, the inner subhumeral elytral stria is rather
fine but distinct, it does not quite reach the apex and ante-
riorly it does not quite reach the middle, the second dorsal
stria is complete, and the mesosternum anteriorly is very
distinctly emarginate. It also very closely resembles H. di-
visifrons, Sch., but it again differs as regards size ; it agrees
with the latter in having rather a longer external thoracic
stria behind the anterior angle, and in having an internal
subhumeral elytral stria. In H. divis¢frons the subhumeral
stria is longer and much deeper. The dorsal striz in
Schmidt’s species are much stronger, and the third dorsal is
widely interrupted in the middle ; in H. brahminius the third
dorsal is only basal, that is, there is no appendage. The
mesosternum is anteriorly sinuous in H. pteromalus, slightly
emarginate in H. divisifrons, and distinctly emarginate in
H. brahminius.
Hab. Ceylon (Andrew Murray).
Hister “nanus, Mars.,” Reitt. Deutsche ent. Zeit. p. 209
(1879). E. Siberia.
This species is, I have no doubt, the same as H. niponicus,
Lew., 1895. Reitter’s name of nanus is a misprint for
navus, Mars.
Hister frontalis, sp. n.
Ovalis, parum convexus, niger, nitidus ; fronte biloba, stria valida
antice recta; pronoto stria marginali antice interrupta, interna
sinuata basi adunca; elytris striis 1 et 3 integris, 2 et 4 basi
haud attingentibus, 5 apicali, suturali dimidiata; mesosterno
emarginato, stria marginali integra; tibiis anticis 3-dentatis.
L. 54 mill.
Oval, rather convex, black and shining; the head is feebly
divided into two lobes by a median impression, which widens
out triangularly behind the stria, stria strong and straight
anteriorly, surface inclosed by the stria is punctulate; the
thorax, marginal stria fine and interrupted in front behind the
head, outer stria very short, little irregular, and confined to
the region behind the anterior angle, inner stria complete,
rather strong, and sinuous laterally, and at the base it turns
away from the posterior angle, the scutellar point is small and
on the basal edge; the elytra, striae, outer humeral wanting,
oblique basal fine and nearly reaching the middle, a line of
new Species of Histeride. 283
rather obscure punctures represents the inner humeral, 1 and
3 dorsal complete, the first turning inwards at the base, 2 and
4 nearly complete, being a very little shortened at the base, 5
nearly dimidiate and apical, sutural nearly twice the length of
the fourth ; the propygidium is impressed on either side, clearly
but not densely punctate; the pygidium, the punctures are
similar but are arranged chiefly along the base; the pro-
sternum, the anterior lobe at the apex is margined with a
stria, but the stria leaves the edge laterally and_ passes
obliquely down the sides ; the mesosternum is rather widely
emarginate and the marginal stria is complete; the meta-
sternum is longitudinally sulcate in the middle; the anterior
tibie are 3-dentate.
Hister bifrons, Mars., agrees with this species in the form
of the head, but in HZ. frontalis the thorax is not bisinuous,
there is a short stria at the angle, the outer subhumeral stria
is wanting, the pygidium is not so densely punctured, and
the mesosternum is emarginate not straight.
Hab. Khasia Hills, Assam.
Stictost’x parra, Mars. Ann. Soc. Ent. Belg. xii. p. 92
(1870).
As there are now four species assigned to the genus Stvcto-
stig from Australia and two from North America, I have
thought it desirable to give a figure of the type of the genus,
S. parra, Mars. (fig. 2). All the species are highly sculp-
tured, but nothing as yet has been recorded of their habits.
Stictostia parra, Mars.
NOTOCGLIS, gen. nov.
Body oblong, gibbous in the dorsal region; head retractile
and declivous, frontal stria carinate ; the thorax contracted at
the base, with the disk almost wholly excavated; the elytra
19*
284 Mr. G. Lewis on
costate; propygidium very large and hexagonal; prosternum
narrow and bistriate; mesosternum anteriorly truncate, stria
complete on the margin; the tibizw are all widely dilated.
‘This genus may be placed near to Discoscelis. In establishing
a genus on a single species it is impossible to give many
generic characters.
Notoceelis satur, sp.n. (Pl. X. fig. 9.)
Oblongus, postice gibbus, rufo-brunneus ; fronte striata, stria margi-
nali carinata; pronoto disco perample excavato ; elytris 4-costatis ;
propygidio longitudinaliter striato; pygidio punctulato ; tibiis
valde dilatatis.
L. 23 mill.
Oblong, gibbous in the dorsal region, dark reddish brown,
somewhat shining ; the head declivous and not visible from
above, forehead transverse and flat, anterior edge sinuous
before the eyes, frontal stria complete and carinate anteriorly,
surface with fine, irregular, transverse stria and some obscure
shallow punctures chiefly visible at the sides; the thorax,
when viewed from the front a stria is seen behind the neck
shaped like a widened-out V, and on either side of it are two
longitudinal striz, the thorax viewed from above is seen to be
rather spread out and thickened at the sides, marginal stria
complete, behind the anterior angles the thorax is longitudi-
nally impressed, and inside the impression is a lateral stria,
the disk of the thorax is wholly and deeply excavated except
in front of the scutelium, where there is a protuberance, feebly
convex on its vertex and ridged and inclined anteriorly, the
anterior rim of the excavation is carinate, the excavation is
nearly smooth ; the scutellum is somewhat elongate, with its
base narrowly transverse ; the elytra, striz, outer humeral
fine and complete, dorsal are represented by four thick costa
which are punctured along their ridges, the first and second
cost are conspicuously sinuous, the third less so and the
sutural costa is only feebly turned from the suture at either
end, there is a fine stria along the sutural margin and the
apices are rounded off at the suture; the propygidium is large
and hexagonal, about half its surface can be seen from above,
and it is conspicuously and longitudinally striated ; the
pygidium is clearly punctulate, but not striate; the pro-
sternum, the keel is narrow and bistriate, the striz are parallel
at the sides and joined at both ends, the anterior lobe is mar-
ginate and in front of the coxe are two semicircular strie
stretching outwards ; the mesosternum is straight anteriorly
and the marginal stria is strong and continues unbroken along
new Species of Histeride. 285
the base of the metasternum, in the centre of which it is
angulate, the suture of the mesosternum is invisible ; the legs,
anterior tibize dilated, external edge nearly semicircular,
intermediate and posterior tibie externally triangularly dilated,
tarsi short with two claws.
Hab. Paraguay (Dr. J. Bohls). ‘Two examples found on
the 5th October, 1892, while it was running over a mound
made by termites.
SITALIA, gen. nov.
Corpus oblongum, subparallelum, convexum, brunneum ; caput inter
oculos planum, antennis funiculo brevi sensim incrassato, clava
ovali haud compressa; pronotum supra convexum ; elytris cari-
natis(fig.3 d); propygidium postice acuminatum ; pygidium declive;
prosternum basi incisum, bistriatum ; mesosternum in medio
minute productum ; tibiz late, fossa tarsali parum lata, subrecta,
leviter exarata.
While establishing the above genus for the reception of
Paratropus Severint, Lew. (Ann. Soc. Ent. Belg. xxxvi.
p. 142, 1892), I have availed myself of M. G. Severin’s kind
Fig. 3.
Sitalia Severin, Lew.
loan of the blocks to reproduce the woodcuts made of the
species from his original drawings. Only one example of
this curious insect is known and no dissections have been
made of it.
Homalopygus remex, sp.n. (PI. X. fig. 10.)
Oblongus, subdepressus, piceus, nitidus ; fronte abrupta, tenuissime
punctulata; pronoto stria marginali basi abbreviata, laterali
obliqua ; elytris striis 1 integra, 2-4 inqualiter abbreviatis,
suturali geminata; pygidio punctulato; prosterno bistriato ;
mesosterno antice biemarginato, stria marginali integra; tibiis
valde dilatatis.
L. 23 mill.
286 Mr. G. Lewis on
Oblong, rather depressed, piceous, shining ; the head very
finely punctulate, abruptly declivous anteriorly ; the thorax,
sides impressed, especially behind the anterior angles, mar-
ginal stria very fine, shortened before the base and continued
round the anterior angle, behind the neck there is a fine
detached stria, not quite close to the edge (this stria appears
to be an appendage to the lateral stria), the lateral stria is also
fine, commencing at the base a little within the angle and
passes obliquely inwards for about three fourths of the thoracic
Jength, on the inside of this stria is another very fine and
very short stria, slightly bent ; the elytra, striee, outer humeral
is shortened apically after passing the middle, inner humeral
is complete, both are very fine, dorsal 1 complete, 2-4 are
shortened apically, 4 is rather longer than the third, 3 longer
than the second, 5 rudimentary or more usually wanting,
sutural is geminate and resembles those figured for H. latipes,
Bohem. (Mars. Mon. 1861, fig. 1); the pygidia are finely
and evenly punctulate; the prosternum, keel bistriate, striee
divergent anteriorly, less divergent at the base, anterior lobe
has a fine marginal stria and a transverse stria marks the
suture between the lobe and the keel; the mesosternum is
biemarginate, the marginal stria is fine and complete, but it
is only near the edge behind the emarginations, at the sides
it joins the lateral metasternal stria; the legs are similar to
those of H. latipes, Bohem.
Marseul’s figure of H. latipes suggests the idea that the
elytral striz are costate, whereas they are very fine, and it is
evident that Marseul mistook the inner part of the sutural
stria, which is geminate or double, for the fifth stria. In
some of my specimens of H. remex the fifth stria is repre-
sented by a rudiment midway between the sutural and fourth
stria. Both species are similar to each other, especially in
the form of the mesosternum and legs.
Hab, Paraguay. I have eleven examples, taken by Dr. J.
Bohls on the 5th Oct., 1892. ‘The beetles were running
about on the mounds made by termites, some were in
copul@ and were fairly numerous. Afterwards Dr. Bohls
searched several hundreds of termite-mounds, but failed to find
other examples either on their surfaces or in the interiors.”
Notocelis satur occurred at the same time.
Terapus muricatus, sp. np. (PI. X. fig. 12.)
Oblongus, cinereo-niger, undique muricatus ; fronte concaya ; pro-
noto antico ad angulos explanato ; elytris leviter costatis ; pygidio
in medio conspicue muricato.
L. 4 mill.
new Species of Histeride. 287
Oblong, ashy black, body and legs densely muricate,
antenne obscurely red; the head, forehead concave, concavity
somewhat smooth, with a lateral stria on either side which
becomes carinate anteriorly and terminates in a tubercle, the
mandibles are microscopically striate ; the thorax is cut out
behind the head in a semicircular form, and the anterior angles
are explanate, the marginate stria is visible on the anterior
edges of the angles, but it is obliterated laterally by the
muricate sculpture; the elytra are costate, the first costa
represents the inner humeral stria, the three next the 1-3
dorsal, the fourth is faint, the fifth and sutural are almost
obliterated ; the propygidium is sculptured like the thorax
and the pygidium also, except in the middle where there is
a bunch of longer sharp-pointed cones and behind them two
lines of small tubercles ; the prosternum, the keel is bordered
laterally by two rough ridges, which widen out a little from
the coxe to the base, on either side of the keel there is a
scooped-out lobe-shaped space which is free of tubercles or
muricate sculpture and it is bordered outwardly by a carina,
the anterior lobe is rough and tuberculate, the keel has an
arch-like incision extending across its base ; the mesosternum
is bisinuous and bordered by a rough muricate ridge; the
legs, anterior tibiz are straight on the inner edge, somewhat
sinuous on the outer, which is armed with 6 or 7 irregular
denticulations ; the intermediate tibia are angulate in the
middle of the outer edge, to which point the tarsal grooves
extend, outer edge with 11 or 12 irregular teeth, inner edge
with small, evenly set, denticulations, posterior tibize also
angular in the middle and denticulated, the thighs on their
upper surface are conspicuously muricate.
This curious species differs much from Terapus Mniszechi,
Mars., but I do not think it necessary to establish another
genus for it. The chief important differences are in the
sculpture, in the thorax having explanateangles and not being
bisinuous behind the head, in the intermediate and posterior
legs being angulate in the middle, and as these angulations
are ab a point where the tarsal grooves end, the tarsi are
relatively longer than in Marseul’s species, where the angles
are nearer to the tarsal end. In 7’. bicarinatus, Lew., the
thorax is not bisinuous and the tarsal grooves are much
shorter than those of 7. Mniszechi, but there is no doubt these
last two species are congeneric.
Hab, Paraguay (Dr. J. Bohls). One example only.
Saprinus punctisternus, sp. n.
Obscure yiridi-eneus, nitidus ; fronte transyersa, utrinque punctata,
288 Mr. G. Lewis on
stria integra; pronoto lateribus basique punctato ; elytris striis 1-4
dorsalibus in medio postice abbreviatis, suturali basi abbreviata ;
prosterno striis integris, subparallelis ; mesosterno marginato,
parce et grosse punctato.
L. 9 mill.
Oval, convex, with a slight greenish brassy tinge; the
head and epistoma transverse (like that of S. semipunctatus, F.),
frontal stria well-marked and complete, vertex punctured,
punctures particularly conspicuous near the eyes ; the thorax,
marginal stria complete, along the sides is a broad band of
rather coarse punctures, which terminate behind the eyes but
continue more narrowly along the base nearly to the scutellum,
there is a small antiscutellar puncture ; the elytra, dorsal strize
are rather oblique and punctured and in form otherwise closely
resemble those of S. splendens, Er., figured in Marseul’s
monograph, fig. 22, but the second and third interstices are
smooth, the basal half of each elytron is rather coarsely and
closely punctured; the propygidium is chiefly punctured at
the sides ; the pygidium has a fine median raised line and is
closely punctured, except at the apex which is nearly smooth ;
the prosternum, keel is parallel and formed like that of S. semz-
punctatus, F’., lateral striee are complete and feebly widen out
at the base ; the mesosternum is marginate and sparingly but
coarsely punctured; the metasternum has a band of points
along its base.
The species belongs to the semdpunctatus-group, but the
punctuation of its mesosternum resembles that of S. optabilis,
Mars. In most of the large Saprini the mesosterna are
smooth. ‘The type specimen of this species is the largest
Saprinus I have seen.
Hab. 'Tai-yuen-fu, Shansi: 5th June, 1897. Collector
unknown to me.
Saprinus castanipes, Curtis, Trans. Linn. Soc. xix. p. 442
(1845),
~An examination of the type specimen in the British Museum
proves this species to be the same as S. fulvopterus, Mars.
Curtis’s name has the priority of ten years. Two other
species described by Curtis at the same time are S. Matthews?
= 8S. bistgnatus, Er., 1834, and S. furcatus = S. connectens,
Payk., 1811.
Saprinus lesus, sp. n.
Ovalis, supra seneus, nitidus ; fronte dense punctata; pronoto haud
ciliato ; elytris macula humerali rufa, stria suturali apice
new Species of Histeride. 289
abbreviata antice hamata; mesosterno stria marginali haud
interrupta.
L. 33 mill.
Oval, convex, aneous above and shining, black beneath ;
the head is closely not coarsely punctured and without frontal
strie ; the thorax, marginal stria entire, punctured on the
lateral borders, with a small impression behind the eye, much
narrowed anteriorly and not ciliate at the sides. The elytra,
the humeral area is red, the dorsal striz, except the sutural,
are very short, or wanting (as the fifth), the sutural is shortened
before the apex, otherwise it is similar to the sutural stria
figured by Marseul for S. béstgnatus, Kr. (Mon. 1855, fig. 13);
on either side of the sutural stria, just behind the middle, is
a small fovea, the-elytral apices are moderately, not densely
punctured, the epipleural stria is continued round the apex of
each elytron, but it does not quite reach the suture. The pro-
sternum, the striz on the keel are strong and anteriorly diverge
(similarly to those of S. bistgnatus) ; the mesosternum is
faintly and sparingly punctulate, the marginal stria is complete
and widely sinuous, but it is not quite close on the anterior
edge; the anterior tibize are 7-8-dentate.
This species is easily distinguished from its congeners,
S. bisignatus, Er., S. lepidus, Ky., and others, by its red
shoulder and complete mesosternal stria.
Hab. Ecuador.
Teretrius Braganze, sp. n.
Cylindricus, niger, nitidus, undique punctatus ; pronoto stria mar-
ginali antice interrupta; prosterno valide bistriato, striis antice
sensim divaricatis, lobo antice marginato; mesosterno antice
obtuse producto et immarginato ; mesosterno metasternoque in
medio canaliculatis.
L. 2 mill.
Cylindrical, black and shining, legs and antenne reddish,
surface above rather densely and strongly punctured, many of
the punctures appear under the microscope minutely ocellate;
the head is rather convex; the thorax, lateral stria well-marked,
but it is not continued behind the neck, on the edge before
the scutellum there isa cluster of dense punctures ; the elytra,
the smooth humeral space is very distinct ; the prosternum,
there is a transverse marginal stria along the apical margin
of the anterior lobe, but down the sides there are only a few
marginal punctures, the keel and lobe have a very few but
conspicuous and irregular punctures, the striz on the keel
are very conspicuous and posteriorly do not quite reach the
290 On new Species of Histeride.
base, from the base they are gradually divergent until they
cease near the edge of the lobe, the base of the keel is impressed
triangularly ; the mesosternum is strongly and obtusely pro-
duced in front, the projection is immarginate, but at the sides
there is an oblique stria which is not, however, joined to the
metasternal lateral stria, there is a median canaliculation
common to both the meso- and meta-sterna and there is no
visible suture between these two segments, the punctures on
the sternal plates are large, sparse, and very irregular; the
anterior tibie are 6-dentate.
This is a very distinct species, but in size and form it
resembles 7. Walkeri, Lew. The chief distinguishing specific
characters of all the Teretrdé are in the sternal plates. Ihave
compared it with the type of 7’. corticalis, Woll.; the iast is
smaller and of a lighter colour.
Hab. 8. Thomas I., Gulf of Guinea (Mocquerys, 1900).
Eptechinus Marseuli.
This name is given to EL. hispidus, Mars., it being clearly
evident that the species described as hispidus, Payk., in the
‘ Abeille,’ i. p. 70 (1864), cannot be the same as Paykull’s
species. ‘The latter is described (Mon. p. 98) as having five
frontal longitudinal carine ; the first has the frontal lateral
borders raised and one median carina. In my collection there
are six Asian species of this genus; Marseul apparently only
knew one. «
EXPLANATION OF PLATE X.
Fig. 1. Placodes opacus, Lew.
Fig. 2. Platylister mirabilis, Lew.
Fig. 3 extrarius, Lew.
1g. 4, Probolosternus permundus, sp. n.
Fg. 5. Baconia loricata, Lew.
Fig. 6. Phelister festivus, Lew.
Fig. 7. Notolister sulcicollis, Lew.
Fig. 8. Hister Marshaili, Lew.
9
Fig. 9. Notocels satur, sp. n.
Fig. 10, Homalopygus remex, sp. n.
Fg. 11. Coproxenus Marshalli, Lew.
Fig. 12. Terapus muricatus, sp. 0.
Except the new species, those figured in the Plate are described in
former numbers of the Ann, & Mag. Nat. Hist.
On Five new Species of Pangonine. 291
XXX VI.—Description of Five new Species of Pangonine from
South America. By Miss GERTRUDE RICARDO.
THE following species were among some Tabanide sent me
by Dr. K. Kertész, of the Budapest Museum, for comparison
with the Walker types in the British Museum collection. He
has kindly allowed me to describe these Pangonine before
returning them to him.
Subfamily Paveoyrvz.
Scione fuscus, 2 , sp. n.
Brown. Face reddish brown, shining and bare. Proboscis
more than half as long as the body. Antenne red, with a
green tinge on the first two joints and apex of third; a few
short black hairs on the first two joints, chiefly on the under-
side. Forehead brown, with greyish-red tomentum, and a
dark brown central stripe ; pubescence short and brown; the
ocelli are very distinct. Palpi yellow, the second joint
brownish; the two joints about equal in length, the second
curved and tapering. Beard white. Eyes hairy. Thorax
reddish brown, with two narrow red stripes, at the sides
lighter in colour; pubescence brown, longer on the sides.
Breast-sides light yellow, with white pubescence and a few
brown hairs in the centre. Scutellum reddish brown. Ab-
domen brown, the first two segments light yellow, with a few
irregular black spots, the third yellow on its anterior margin,
its posterior half and the remaining ones brown, with irregular
black markings; the posterior half of the last segment is
yellow; the dorsum of abdomen covered with short black
hairs, at the sides a few silvery-white hairs are intermixed
with the black ones, beginning from the third segment; the
underside of abdomen is light yellow, with a few small, round,
black spots. Legs yellow, apex of tarsal joints brown.
Wings smoky brown, fore border yellow; there is a clear
band across the middle of wing tinged with yellow, com-
prising the basal half of the marginal and submarginal cells,
three quarters of the first posterior cell, and the whole of the
discoidal cell; veins brown. MHalteres yellow, somewhat
brown on the head of club.
Length 11 millim.
Type (2) from Bolivia (Songo).
292 Miss G. Ricardo on Five new
Scione claripennis, 2, sp. n.
This species is allied to S. maculipennis, Schiner (Dichia
maculipennis), but differs in having clear wings, cross-veins
not shadowed, and no spots.
Brown. Face brown, greyish on the lower part near
mouth and at the sides, with brown pubescence. Antenna
brown, the basal half of the third joint bright red, the first
two joints with long brown hairs. Forehead fawn-coloured,
with a large, almost square, dark brown spot, not reaching
the sides ; a central dark line extends from it to the ocelli-
gerous tubercle, which is also brown, as is the pubescence on
the forehead. Palpired, the two joints about equal in length,
the second curved and tapering. Beard white. Eyes
hairy. Thorax with three grey stripes ; the side ones do not
reach the border, but join the central stripe; at about half
their length a branch stripe begins, which is extended to the
shoulders ; pubescence on dorsum and sides brown, yellowish-
white tufts at base of wings; breast-sides brown, with tufts
of white and brown hairs. Scutellum brown, grey in the
middle and on the margin, with white hairs in the centre and
brown ones on the margin. Abdomen brown, with tufts of
white hairs on the posterior borders of segments in centre,
forming an irregular stripe; the first three segments are
yellowish, with dark markings, the posterior borders of the
following dark brown segments are lighter in colour; the
pubescence consists of black hairs, except on the posterior
border of second segment, where they are white; the sides of
segments 2, 3, 4, 5 have white hairs; underside of abdomen
yellow, with dark markings and bands of yellow on the
posterior borders, the Jast segments almost wholly black.
Legs black, knees yellow, the basal half of the anterior and
middle tibie brown, coxe and femora clothed with long
black hairs. Wings hyaline; at the base and on the fore
border, extending also to the cross-veins of the basal cells, is
a slight yellowish tinge.
Length 103 millim.
Type (2), Peru (Callanga).
PanGontA, Latr.
Erephrosis niger, 3, sp. n.
Black. Antenne black, the third joint dull red, with black
segmentations, first and second joints with black hairs.
Palpi and face black, the latter with the second joint longer
Species of Pangoninze from South America. 293
than the first, tapering to a point. Proboscis nearly as
long as body. Beard brown above, grey below. ‘Thorax
and scutellum dull brown, with spare black pubescence and a
white tuft of hairs at the base of wings; the pleure fringed
with white hairs. Abdomen black and shining; a white
tuft of hairs on the lateral margins of the last two segments,
the other segments bordered at the sides with black hairs.
Legs black, the fore tibize and tarsi dull brown, with fulvous
pubescence on the underside of the first tarsal joint; the last
tarsal joint of posterior legs is reddish. Wings grey, at base
and on fore border yellowish; a white oblong spot at the
extreme base of the wing, another at the root of the first basal
cell, and a short white line between the first and second
longitudinal veins, starting from the root of the latter ; veins
yellow, no appendix.
Length 14 millim. ; proboscis 11 millim.
Type (¢) from Surinam, 8. America.
Erephrosis rufopilosis, 2, sp.n.
Dark brown. Antenne and face black, the latter with a
band of ashy-grey tomentum below the antenne. Palpi
black, long, extending far beyond the face; the second joint
nearly twice as long as the first, broad at the base, with a
deep furrow on the outer side, tapering to a point, edged with
black pubescence. Forehead brown, covered with rather long
black pubescence. Beard black. Thorax with long brown
hairs on the anterior half and on the sides ; scutellum reddish
brown; a thick tuft of black hairs at base of wings. Abdo-
men dark brown, the first three segments somewhat reddish ;
with bright orange-red pubescence on the sides of the seg-
ments, widest on the second segment, becoming a transverse
fringe on the posterior margins of the fourth and fifth seg-
ments, and appearing as scattered hairs on the dorsum of the
posterior segments ; underside black and shining, the orange
hairs appearing on the extreme lateral margins only. Legs
black, the anterior coxe with greyish pubescence ; the femora
and tibie fringed with long black hairs, thickest on the ante-
rior ones, some fulvous pubescence on the underside of the
tarsi. Wings brown, yellow on fore border and at base, with
lighter centres to the cells on the posterior border and two
white spots at the base of the wing, besides a white streak
between the first and second longitudinal vein; an appendix
on the branch of the third longitudinal vein.
Length 16 millim. ; proboscis 8 millim.
Type (?) from Bolivia (Songo).
294 Mr. O. Thomas on new
This species resembles Hrephrosis rufo-hirta, Walker, and
other Brazilian species in the shape of the abdomen and
general appearance (see Ann. & Mag. Nat. Hist. (7) v. p. 175,
1900).
Erephrosis rufescens, 2, Sp. n.
Two females from Peru (Callanga) and Bolivia (Songo).
This species should belong to Scione, Walker (Diclisa,
Schiner), having the fourth posterior cell closed; but as it
does not agree with the description of the genus in any other
particulars and has all the appearance of a true Pangonia
(Latr.), it seems best to include it for the present under
Pangonia (Latr.) (see Ann, & Mag. Nat. Hist. (7) v. p. 104,
for similar cases).
Reddish brown. Antenne dull red, lighter at the apex,
with black segmentations ; the first joint stout, nearly three
times as long as the second, both with black hairs at the
sides. Face and palpi brown, with fawn-coloured tomentum,
the latter curved; the second joint longer than the first,
broad, ending in a point bordered with black pubescence.
Forehead reddish, fulvous at the sides bordering the eyes,
with short black pubescence. Beard black, thin, with a few
red hairs. Thorax brown; scutellum red: both with short
fulvous pubescence. Abdomen red, shining, with irre-
gular black markings and orange-red pubescence on the lateral
margins, becoming thicker on the apex. . Legs red; the coxe
brown, with black pubescence; the femora reddish brown,
with short black pubescence ; the anterior and posterior tibie
and tarsi with some reddish pubescence. Wings hyaline,
tinged with yellow on the fore border; no appendix ; fourth
posterior cell closed.
Length 17 millim.; proboscis 7 millim.
Type (2) from Peru (Callanga).
XXXVII.—Descriptions of new Rodents from Western South
America. By OLDFIELD THOMAS.
Sciurus (Microsciurus) Stmonsi, sp. n.
Size as usual in this group. General colour above grizzled
olivaceous, about as in S. Alfari, though slightly darker.
Under surface also very much as in that species, the hairs
tipped with buffy, not ferruginous. Sides of nose yellowish.
Rodents from Western South America. 295
Eyes with marked yellowish rings round them. Lars well-
haired, without lighter spots: behind them, dark rusty red.
Feet grizzled yellowish. Tail fairly long-haired, the hairs
ringed black and reddish basally, with black subterminal and
dull yellow terminal rings.
Skull slightly larger than in S. A/fari; nasals decidedly
shorter, their posterior edge directly transverse, some dis-
tance in front of the termination of the premaxillary processes.
Molars larger than in the allied species.
Dimensions of the type (measured in the flesh) :—
Head and body 138 millim.; tail 112; hind foot, s. u. 35,
c. u. 38; ear 16.
Skull: greatest length 38°8; basilar length 29; greatest
breadth 23°5; nasals 9°4 x5; interorbital breadth 14; tip to
tip of postorbital processes 20; palate length from henselion
16 ; diastema (to front of p.*) 9; length of tooth-series (ex-
cluding the minute p.’) 6°3.
Hab. Porvenir, near Zaparal, Province of Bolivar, Hcuador.
Altitude 1500 m.
Type. Female. B.M. no. 99. 9.9.12. Original number
261. Collected 16th March, 1899, by Mr. Perry O. Simons.
Of the seven species and subspecies of the subgenus Micro-
sciurus represented in the British Museum, S. Simons? is
undoubtedly nearest to the Costa-Rican S. Alfar?, Allen, but
differs by its rusty red ears, more prominent eye-rings, shorter
nasals, larger molars, and widely different locality.
Scturus (Microsciurus) perwanus napi, subsp. n.
Quite like S. peruanus of N.W. Peru in the general oliva-
ceous colour, absence of eye-rings, white patches on and
behind ears, and other characters, but distinguished by the
coloration of the belly, which, instead of being fairly defined
yellowish rufous, is only faintly and indistinctly suffused
with the same colour, the dark bases to the hairs giving the
general tone to the under surface.
Skull as in true S. perwanus, but the nasals apparently
rather shorter and more oval.
Dimensions of the type (measured in skin) :—
Head and body 157 millim.; tail (broken at tip); hind
foot s. u. 84; ear 12.
Skull: greatest breadth 21; nasals 9x5; interorbital
breadth 13; diastema 8; length of upper tooth-series (ex-
cluding p.*) 6.
Hab. Mouth of Coca River, Upper Rio Napo.
Type. Female, slightly immature. B.M. no. 0. 6. 3. 6.
296 Mr. O. Thomas on new
Collected June 1899 and presented by Walter Goodfellow,
Esq. A second specimen, labelled as from the Rio Napo,
has been in the Museum since 1875.
Phyllotis amicus and its subspecies.
After obtaining the specimens of this species—at Eten,
Reque, and Tolon—mentioned in the original descrip-
tion *, Mr. Simons collected a number further inland and
higher up at Uramarca, on the River Ushpe, not far from
Pallasca. The three localities (for Eten and Reque may be
treated as one) are all approximately at the same latitude ;
but the first is on the coast, in the desert region, at an alti-
tude of only 20-50 metres, the second, Tolon, about 60 miles
inland at 100 metres, and the last is at about 1200 metres on
the Andean chain.
On comparing the three sets it proves that the coast
specimens are smaller and paler in colour than the middle,
typical, set from Tolon, while, on the other hand, those from
Uramarca are larger, with longer tails and, especially, much
larger ears. Further south, on the Shigray River, ‘Tambo
(1600 metres), and at Marca (2000 metres), the same large
long-eared form occurs, while at Chosica, near Lima, at
850 m., the local representative of Ph. amicus is again much
like that found at Tolon.
It would therefore seem that along the higher level (1200-
2000 m.) from about lat. 7° to 12° 8., the long-eared form
occurs, that the intermediate typical one ranges along the
middle altitudes (100-850 m.), while the Eten one may also
hereafter be found to occur further southwards along the
coast, perhaps to Callao.
The subspecies may be briefly distinguished as follows :—
Phyllotis amicus maritimus, subsp. n.
Size small (head and body averaging 78°5 millim. in six
specimens) ; tail short (96°6) ; ears short (22°25).
Colour paler and more sandy than in the typical form,
especially posteriorly ; approximating to the darker specimens
of the desert species Ph. gerbillus.
Dimensions of the type (measured by collector in the
flesh) :—
Head and body 80 millim.; tail 100; hind foot 223 ear
22:5.
Skull: basilar length 17:1.
* Ann, & Mag. Nat. Hist. (7) v. p. 355 (1900).
Rodents from Western South America. 297
Hab, Kten, coast of N.W. Peru. Altitude 20 m.
Type. Male. B.M. no. 0. 3.1.62. Original number 589.
Collected 16th September, 1899, by Mr. P. O. Simons.
Phyllotis amicus (typical).
Average measurements of five specimens from the type
locality, Tolon, altitude 100 m. :—
Head and body 85:2 millim.; tail 100; hind foot 22°6;
ear 22°6.
Colour rather darker than in either of the other subspecies.
Phyllotis amicus montanus, subsp. n.
Size comparatively large (average of four specimens:
head and body 90 millim.; hind foot 24); tail long
(average 117°5) ; ears very large (average 24°75).
Colour about as in the typical subspecies or slightly paler.
Dimensions of the type (measured in the flesh) :—
Head and body 93 millim.; tail 122; hind foot 24;
ear 26.
Skull: basilar length 19:3.
Hab. Uramarca, near Pallasca, N.W. Peru. Altitude
1200 in.
Type. Female. B.M. no. 0. 6. 6. 28. Original number
760. Collected 7th December, 1899, by Mr. P. O. Simons.
The four Tambo and Marca specimens of PA. a. montanus
have the following average measurements :—
Head and body 90 millim.; tail 111; hind foot 22°75;
ear 24°75.
The two from Chosica, near Lima, which I refer to the
typical Ph. amicus :—
Head and body 85 millim.; tail 107°5; hind foot 23;
ear 22°.
Eligmodontia sorella, sp. n.
A medium-tailed fawn-coloured species with long soft hair.
Size rather less than in Mus musculus. Fur long, smooth,
very soft and fine, the ordinary fur about 9 millim. long on
the back and 6-7 millim. on the belly, and profusely mixed
with longer hairs about 16 millim. in length. General
colour sandy fawn, darker and closely lined with black on
the back, clearer on the sides, where there is a well-defined
fawn-coloured line edging the white of the belly. Tips of
Aun, & Mag. N. Hist. Ser. 7. Vol. vi. 20
298 Mr. O. Thomas on new
the longer dorsal hairs shining silvery. Under surface
whitish, the hairs visibly slaty grey basally, white terminally ;
line of demarcation on sides well marked. Face slightly
greyer than back. ars of medium length, their visible
surface when folded brownish fawn; a marked white patch
behind their posterior bases. Upper surface of hands and
feet uniformly pure white; proximal third of soles thinly
hairy ; fifth hind toe reaching to the middle of the basal
phalanx of the fourth. Tail rather shorter than head and
body, uniformly closely haired, so as to hide the scales, not
pencilled ; brownish fawn above, darkening nearly to black
at the tips, white below and on the sides.
Skull delicate and papery; nasals and premaxillary pro-
cesses of equal extension behind ; interorbital region flat, its
edges square but not beaded; palatal foramina long, reaching
backwards to the middle of m.*; bulle small.
Dimensions of the type (which is adult, but not old),
measured by collector in the flesh :—
Head and body 72 millim.; tail 62; hind foot 18 (ce. u.
19°5) ; ear 17.
Skull: greatest length 22-7; basilar length 17:2; zygo-
matic breadth 10°7 ; nasals 8°8 x 3; interorbital breadth 3:7;
breadth of brain-case 11:0; interparietal 2°2 x 8°5; diastema
57; palatine foramina 5°5x2; length of upper molar
series 3°5.
Hab. Hight miles south of Huamachuca, N.W. Peru.
Altitude 3500 m.
Type. Female. B.M. no. 0. 6.6.29. Original number
741. Collected 28th November, 1899, by Mr. Perry O.
Simons.
This pretty little mouse has no near relationship to the
E. lepida * of Central Peru, which is one of the very short-
tailed species, but is most closely allied to the Argentine and
Paraguayan ZH. lancha, Desm., from which it differs by its
longer and softer fur, more fawny colour, longer and slen-
derer feet, and other details,
Oxymycterus inca, sp. Nn.
Size rather large, not quite equal to the large Brazilian
species, but greater than in OU. rufus of Paraguay or La
Plata. Fur coarse, rather short ; hairs on back 11-12 millim.
in length. General colour rufous chestnut, more grizzled
into black along the centre of the face and dorsal area, more
* Hesperomys bimaculatus, var. lepidus, Thos, P. Z. 8, 1884, p. 454,
pl. xlii. fig. 2.
Rodents from Western South America. 299
uniform reddish on the sides. Ears well haired, blackish
brown. Chin prominently contrasted white ; whole re-
mainder of under surface rich ochraceous rufous, not defined
laterally from the red of the sides; bases of the hairs indis-
tinctly plumbeous. Upper surface of hands and feet blackish
brown. Tail well haired throughout, blackish brown, slightly
lighter along the median line below.
Skull long, the muzzle of the peculiar trumpet-shape
characteristic of the large Brazilian species of the O. hispidus
group. Nasals long, broadest anteriorly, reaching backwards
to the level of the front edge of the anteorbital bridge ; supra-
orbital edges as usual, not beaded or ridged, but there is a
slight angular projection in the position of the postorbital
processes, whence parietal ridges run nearly directly backwards,
giving the brain-case an oblong parallel-sided appearance ;
interparietal well developed ; anterior zygoma-root much
slanted backwards; palatal foramina large, widely open,
extending backward to the middle of m.’.
Dimensions of the type (measured in flesh by the col-
lector) :—
Head and body 135 millim.; tail 105; hind foot, s. u.
80, c. u. 33; ear 21.
Skull: greatest length 38; basilar length 30; zygomatic
breadth 18; nasals 14:3 x 5; interorbital breadth 6°7; tip
to tip of rudimentary postorbital processes 11*7 ; interparietal
2°6 x 6; diastema 9; palatal foramina 8 xX 3°2; length ot
upper molar series 5°7.
Hab. Perené, Ucayali watershed, Department of Junin,
KE. Peru. Altitude 800 m.
Type. Male. B.M. no. 0. 7.7.45. Original number
925. Collected 10th April, 1900, by Mr. P.O. Simons. Four
specimens examined,
This is the first Peruvian Owymycterus described. _ Its
size and cranial characters will readily distinguish it from
any known species.
In the same collection, Mr. Simons secured at Galéra, the
highest point of the Oroya railway, altitude 4800 m., an
example of the remarkable Neotomys ebriosus, 'Thos., described
in 1894 * trom a specimen collected by Mr. J. Kalinowski in
the Valley of Vitoc. Mr. Simons’s skin shows that the under-
side of the tail is not white but dull buffy, and that the
chest is dirty brownish, a colour which runs backward a short
distance along the centre of the belly.
* Ann, & Mag, Nat. Hist. (6) xiv. p. 348,
20*
300 Mr. O. Thomas on new
Proechimys Simonst, sp. n.
Size rather smaller than in P. chryscolus, its nearest ally.
Rump practically spineless; nape and sides also much less
spinous than the centre of the back. Spines short, only
about 15 millim in length, and narrow (0°6-0°8 millim. in
breadth). General colour above very like that of P. rosa,
with the same grizzled mixture of black and fulvous. Head
rather darker and less fulvous. Under surface and inner
sides of limbs pure sharply defined white, much more sharply
defined than in the allied species. Upper surface of feet dull
brown, lighter along the inner margins. Tail of about the
same length as in P. rosa and chryscolus, but much less
hairy, the hairs not hiding the scales; scales unusually large,
the rings running about 8 to the centimetre ; its colour promi-
nently bicolor, black above, white below.
Skull remarkably like that of P. brevicauda, Giinth.,
although in external characters the species has little re-
semblance to that animal. In size it is smaller than in any
of the species mentioned. Nasals long, narrow, rounded
behind, extending about 2°5 millim. beyond the premaxille
posteriorly. Interorbital region narrow. Parietal ridges
almost obsolete, a faint indication of them in the positions
described in P. chryseolus, not continuous as in P. rosa.
Palatal foramina broad and rounded, almost as broad as long,
the ridges on the palate just behind them nearly obsolete.
Palate emarginate to the back of m.”. Pterygoid processes
narrow, curved, not spatulate. Molars small and rounded.
Dimensions of the type (measured by Mr. Simons in the
flesh) :—
Head and body 210 millim.; tail 165; hind foot, s. u.
47,c. u. 49; ear 26.
Skull: greatest length 53:2; basilar length 36°5; zygo-
matic breadth 25°3; nasals 20 X 5:5; interorbital breadth
11-3; greatest breadth on parietal ridges 20; interparietal
75 x 12:2; diastema 10°4; palatal foramina 4°3 x 3:5;
length of molar series 7-5.
Hab. Perené River, Junin Province, Peru. Altitude
800 m.
Type. Adult male. B.M. no. 0. 7. 7. 50. Original
number 942. Collected 14th April, 1900, by Mr. Perry O.
Simons. “ Haten by natives. Found in deep woods.
Native name ‘ Pericota.’”’
‘This species presents the curious anomaly of being closely
similar externally to P. chryseolus and rosa, while its skull is
scarcely distinguishable from that of the outwardly very
Rodents from Western South America. 301
different P. brevicauda, Giinth. Its most obvious diagnostic
characters are its thinly haired, large-scaled tail, its sharply
defined white belly, its small skull, suppressed parietal ridges,
and short open palatal foramina.
In his description of P. brevicauda * Dr. Giinther speaks
of “an adult male in spirit and the skin of an adult female
brought by Mr. E. Bartlett from Chamicuros, Huallaga
River”; but, as a matter of fact, it is only the skin
(received in 1869) that is labelled as from Chamicuros, and
the spirit-specimen is merely recorded from the Upper
Amazon, and was received by the Museum three years
before, in 1866. It is therefore probable that they are from
different localities, and there is unquestionably great doubt as
to their specific agreement. Under these circumstances it
would be advisable to treat the skin by itself, accurately
labelled and with good skull, as the type, and to ignore the
second specimen, which has no exact locality and whose skull
is broken to pieces.
Ctenomys tucumanus, sp. n.
Size small, about as in C. talarwm. General colour above
brownish fawn, with a faint reddish suffusion ; middle line of
face blackish. Cheeks like back, a faint lighter patch below
ear. Under surface pale buffy, the hairs plumbeous basally ;
large white axillary and inguinal patches present, the former
almost extending across the chest (but this may be due to the
great age of the specimen, which is also becoming hoary on the
back). Upper surface of hands well haired, whitish, of feet
nearly naked, the few hairs also white. ‘Tail practically
naked, a few whitish hairs forming a slight terminal crest.
This nakedness of hind feet and tail is possibly due to age
and wear.
Skull broad and flattened, much more so than in C. mendo-
cinus, and still more different to the lumpy rounded skull
of C. Perrenst. Nasals evenly tapering backwards, their
posterior end behind the level of the anteorbital bridge.
Interorbital region flat, shorter and broader than in C. talarum,
as also is the brain-case. No interparietal discernible. Malar
less heavily ridged than in C. talarum. Posterior nares
broad and open, the pterygoid ridges much further apart than
in the allied species.
Dimensions of the type (measured in the flesh) :—
Head and body 172 millim.; tail 71; hind foot, s. u. 27,
c. u. 30°55 ear 6.
* PZ, S. 1876, p. 749.
302 On new Rodents from Western South America.
Skull: greatest length in middle line 43°3; basilar
length 38; zygomatic breadth 27:7; nasals 16 x 6-4; length
of frontal suture 10-2; interorbital breadth 9°1; breadth
across postorbital processes 11'1; least posterior breadth
across brain-case 17; greatest posterior breadth on meatus
27°5; palate length from benselion 20°2; diastema 12;
length of upper molar series (crowns) 9°5; greatest diameter
of anterior tooth (p.*) 4.
Hab. Tucuman. Altitude 450 m.
Type. Old male. B.M. no. 0. 7. 9. 14. Original number
133. Collected 25th September, 1899, by Signor Luis
Dinelli.
Although geographically nearest to C. mendocinus, Phil.,
and C. Perrenst, Vhos., this Tuco-Tuco is probably most nearly
allied to C. talarum, with which it shares the small size and
flattened shape of the skull. It differs, however, from that
species by its broader skull, especially the broader interorbital
region, more open choane, and much paler coloration.
Signor Dinelli also obtained at Tucuman an example of the
rare armadillo Dasypus vellerosus, Gray, and of the Chilian
opossum Marmosa elegans, Waterh., not hitherto known
from Argentina.
Cyclopes* didactylus ida, subsp. n.
Geneial colour of the more greyish type characteristic of
the typical C. didactylus from Guiana, not yellowish or
golden as in the Mexican and Central-American C. d. dor-
salis, Gray. Rump, legs, and tail grey, not yellow. Dorsal
stripe indistinet, irregular, almost lost in the coarse marblings
of the back. Under surface uniform dull yellowish or buffy,
without any trace of the dark sternal line present in the other
subspecies.
Skull very like that of the Guianan form; edges of inter-
orbital region similarly evenly diverging backwards.
Dimensions of the type, in skin :—
Head and body (c.) 180 millim. ; tail (c.) 198.
Skull: greatest length 49; greatest breadth of brain-case
23°5; length of nasals (laterally) 13°5; interorbital breadth
9°6; length of frontal suture 23:6, of parietal suture 8°7 ;
length of lower jaw 38.
Hab. Oriente of Ecuador. Type from Sarayacu, Upper
Pastasa River; another specimen from the Rio Napo.
Type. Female. B.M. no. 80. 5. 6. 69. Collected by
Mr. Clarence Buckley. Four specimens from Sarayacu
examined and one from the Napo.
* Gray, 1821. Cyclothurus, Gray, 1825, auctorum.
On the East-African Reedbuch. 303
XXXVIII.—On the “ Tohi,” the East-African Reedbuck cur-
rently known as Cervicapra bohor. By OLDFIELD THOMAS.
MEssrs. ROWLAND WArD have recently put into my hands
three Reedbuck skulls which, collected some twenty or
thirty years ago, had become scattered into different collec-
tions, but which, noticing their peculiarity, Messrs. Ward had
kept in touch with and have now brought together again in
order that I might examine and report upon them.
They were obtained in some part of the Upper Nile, and,
according to one account, at Kassala. ‘Their collector was a
Herr Essler, by whom a number of other large mammals
now in the British Museum were precured at the same time.
Their northern locality has still more recently been con-
firmed by Dr. Donaldson Smith, who obtained several
specimens of the same form during his recent journey from
Lake Rudolf to the Nile, in about 5° N. latitude.
Now these specimens all agree among themselves, and
differ from all the species recognized in the ‘ Book of Ante-
lopes,’ by the peculiar graceful curvature of their horns,
which, while first sloping backwards and then outwards
somewhat as in C. arundinum, are distinctly (though not
abruptly) recurved forwards and inwards terminally. As a
result their back view is not altogether unlike the more
distant figure of C. arundinum in the ‘ Book of Antelopes,’
pl. xlii., though the tips approach each other terminally
much more, while their side view is similar both to those of
the East-African antelope currently known, since Dr. Giinther’s
paper on the subject *, as C. bohor, and also to Riippell’s
figure of ‘“ Antilope redunca” +, afterwards the type of his
C. bohor.
The side view of the horn-curvature being therefore the
same, Dr. Giinther, in the absence of Abyssinian material,
not unnaturally assigned the Kast-African animal, the “ Tohi”’
of Mr. Jackson in ‘Big Game Shooting,’ to Riippell’s
species; but it is now quite clear, both by locality and b
some details about the type kindly sent me by Dr. Kobelt,
that Messrs. Ward’s specimens are the true Bohor, being the
first examples of it that have come to this country. By their
aid we see that it is a peculiar northern species, most nearly
allied to C. arundinum, to which it approximates in size, but is
distinguished by its terminally incurved and recurved horns.
* P. Z. 8. 1890, p. 604.
t+ N. Wirb. Abyss. pl. vii. fig. 1 (1835).
304 On the East-African Reedbuck.
A second name applicable to it is “* Cervicapra? odrob,”
Heuglin*, based on an antelope observed by him on the
River Setit.
But if this antelope, with its widely expanded semilyrate
horns, is the true C. bohor, the question arises as to what is
the proper appellation for the very different Reedbuck from
East Africa which has of late been known by that name.
That animal, as is shown in the ‘ Book of Anutelopes’ f, is
closely allied to the West- African Nagor (C. redunca, Pall.),
being distinguished from it merely by its greater size, the
difference there quoted being 9 inches as compared to 8 in
the basal length of the skull.
A renewed comparison not only confirms the alliance of
the two forms, but shows that the difference is even less than
we then supposed; for Messrs. Ward, who had already helped
so materially in this matter, have lent me the skull of an
unusually fine Nagor which had been shot by Lieut. A. A. E,
Ellison, R.N., 160 miles up the River Gambia, and this skull
measured no less than 8°65 inches (220 millim.) in basal
length, nearly as much as in ordinary EHast-African Reed-
bucks. The fur, at least of the bead, is closer and finer than
that of Kast-African specimens, but is quite similar in colour,
These being the facts, it would appear better to treat the
Tohi as a subspecies of C. redunca, and in recognition of the
help received from Messrs. Ward I would suggest the name
Cervicapra redunca Wardi, subsp. n.
Size rather larger than in the typical subspecies, basal
length of skull 9 inches or more. Hair coarser and shaggier.
Colour similar.
Skull of the type (an old male) :—
Basal length 234 millim. (9:2 inches) ; greatest breadth
114 (4:48) ; muzzle to orbit 138 (5°45). Horns: length round
curve in front 235 (9°25), circumference at base 146 (5°75),
breadth across most distant points externally 190 (7:5), tip
to tip 143 (5°6).
Hab. Kast Africa. Type from the Mau Plateau.
Type. Male (skin and skull). B.M. no. 0. 8.15.1. Col-
lected by Mr. F. J. Jackson. Presented by Mr. Rowland
Ward.
* Reise N.O.-Afr. ii. p. 109 (1877).
+ Vol. ii. p. 155, synopsis,
On new Species of Hastern and Australian Moths. 305
XXXIX.—New Species of Eastern and Australian Moths.
By Colonel C. Swinuog, M.A., F.L.S., &c.
Family Syntomide.
Eressa catoria, nov.
3. Fore wings as in LZ. confinis, Walker: hind wings all
black, with one small circular hyaline spot beyond the cell,
with indications of a smaller spot below it ; in many speci-
mens both these small spots are indistinct. Antenne white
at the tips; a large orange spot on the prothorax, orange
streak on metathorax ; abdomen with a line of orange spots
on vertex, on the sides, and also underneath.
Expanse of wings 1 inch.
Jaintia Hills. Many examples, all males.
Allied to £. musa, Swinh., and L£. conjfinis, Walker.
Differs from the former in having white tips to the antenne,
and from the latter in the orange streak on metathorax, and
from both in having the hind wings all black.
Family Chalcosiide.
Pidorus leno, nov.
g ?. Antenne, body, and wings deep black, with an
olive tinge, collar crimson ; face, body below, and the under-
side of fore legs white. Fore wings with a wide oblique
yellowish-white band from the costa beyond the middle to
the hinder angle, touching neither: hind wings with a
yellowish-white costal band extending round the apex,
showing a whitish apical narrow patch: underside of both
wings paler, with the basal portions greyish.
Expanse of wings 1,/5 inch.
Jaintia Hills. ‘Three males and one female.
Superficially like a very small P. geminus, Walker, but
the white costal band of the hind wings above and the different
colouring below are very striking, the underside of the wings
in P. geminus, Walker, being streaked and shot with blue,
Isbarta curtosa, nov.
3 ¢. Antenne, head, thorax, and abdomen black, slightly
metallic ; antenne white at the base, a white spot behind the
head, and a white spot on each tegula; face white, body and
legs also white on the underside. Wings dull black, with
306 Col. C. Swinhoe on new Species of
yellowish-white spots and streaks in the interspaces, the
female with the spots and streaks somewhat as in J. imitans,
Butler ; the spots, however, are shorter, and the three long
streaks below the cell are disjointed outwardly: the hind
wing is also somewhat similarly marked, but the basal
streaks fill up the whole of the interspaces, leaving no black
except on the veins, and the colour is rather deep yellow.
The male differs from the female in being much smaller; the
wings are almost entirely black, there being little more than
indications of the spots and streaks.
Expanse of wings, ¢ 27, 2 2;%5 inches.
Malang, Java. One pair.
Family Drepanulide.
Deroca hidda, nov.
3 9. Shaft of antenne, head, thorax, and abdomen
white, branches of antenne dark grey. Wings semihyaline,
two sinuous grey bands close together before the middle on
the fore wings, three similar bands close together before the
outer margin of both wings, marginal space grey, or it might
be described as having three sinuous white bands before the
outer margin on a grey ground; the hind wing has also
indications of a band before the middle, marginal line
black, veins grey and more or less prominent.
Expanse of wings 179-17 inch.
Jaintia Hills. T'wo males and three females.
Much smaller than D. hyalina, Walker; wings better
clothed, no marginal spots.
Family Lymantriide.
Gazalina intermizta, nov.
¢@. Head white, collar and thorax pale rufous; abdomen
white, with black segmental thin bands. Wings pure white ;
fore wings with two medial, oblique, thick, black transverse
lines, the outer one outwardly elbowed beyond the cell; a
subbasal black line; three or four short black streaks on the
veins between the outer line and the margin: hind wings
without markings. Underside pure white, both wings with
black central, transverse, thin bands.
Expanse of wings 1} inch.
Jaintia Hills. Two examples.
Allied to G. chrysolopha, Kollar; differs in its white
Eastern and Australian Moths. 307
abdomen and in the absence of the black streak along the
median vein of fore wings, and the black streaking of all
the veins outside the outer transverse line.
Lelia adalia, nov.
3 2. Shaft of antenne pale pink, branches dark grey ;
male with the head, thorax, and fore wings dark red-brown,
in the female paler and more red; a black lunule at the
upper end of cell, a pale streak through the lower half of
cell; the median vein and its veinlets lined with black, and a
black streak opposite the lunule: abdomen and hind wings
red-grey, paler and more red in the female ; without markings
in both sexes. Underside paler; both wings uniformly
coloured ; fore wings with all but the borders suffused with
brown.
Expanse of wings, ¢ 14%, 92 245 inches.
Jaintia Hills. One male and six females.
The male somewhat like ZL. atestacea, Hmpsn., but
differently marked ; the female of L. atestacea is, however,
of a uniform pale yellowish white, with internervular grey
streaks,
Family Epiplemide.
Epiplema kohistaria, nov.
&. Hind wings with slight tails at veins 4 and 7, with
lesser productions on all the other veins; long stiff hairs on
the basal half of costa beneath. Upperside dark greyish
purple ; abdomen with a broad purple band near the base :
fore wings with a purple line edged with ochreous on both
sides, from costa one sixth from apex to hinder angle, the
upper half curved to an angle near the outer margin at the
middle, then sharply bent inwards, then again straight to the
hinder angle: hind wings with a somewhat similar line, but
farther away from the outer margin, marked with black on
the inside and with some black streaks on the outside; there
are also several black patches inside the wing somewhat in
the shape of an antemedial band; both wings with black
marginal lunules and ochreous cilia. Underside nearly
white, suffused in places with purple.
Expanse of wings | inch.
Port Blair, Andamans. ‘Two examples.
308 Col. C. Swinhoe on new Species of
Family Boarmiide.
Subfamily Bosrurwz.
Boarmia delatina, nov.
2. Of a uniform pale greyish fawn-colour: fore wings
with three outwardly curved indistinct grey lines, antemedial,
medial, and discal, the last two meeting rather close together
on the hinder margin; all three lines marked with black
spots on the costa and with some black points on the veins:
hind wings with two grey similar lines, one central, the other
from three fourths of costa, much curved, terminating on
abdominal margin rather closer to the central line than it is
on the costa. Underside greyish white, with two lines across
both wings, broad marginal grey borders, with two large
white spots on outer margin of both wings, apical and medial.
EXxpanse of wings 1,°5 inch.
Koni, Shan States (Manders). One example.
Subfamily Lwwourrwz.
Anaxa kaluga, nov.
9. Pale yellow; wings sparsely irrorated with pinkish
atoms: fore wings with a thin pinkish band from costa, quite
close to apex, to hinder margin, just beyond the middle, the
space beyond being filled in with pinkish except at the hinder
angle: hind wings with the thin pinkish band across its
centre, but obsolete towards the costa. Underside as on
upperside, but slightly paler.
Expanse of wings 1,5 inch.
Jaintia Hills. Four examples, all females.
Closely allied to A. cesadaria, Walker,=A. sulphurea,
Butler, of which I have a fine series of both sexes ; the apex
of fore wings is more produced and the band is straight and
very nearly touches the apex instead of curving inwards
in a sinuous form one fifth before apex, as in A. cesadaria,
Walker.
Hypephyra etawa, nov.
g. Above olive-brown, with a decided pinkish tinge: fore
wings with the apical portion paler, containing three or four
brown spots: hind wings with a broad marginal even band
of pale pinkish ; both wings crossed by ante- and postmedial
Eastern and Australian Moths. 309
zigzag pale pinkish thin bands; cilia brown, with pinkish
tips. Underside uniform pale pinkish grey, without markings.
Expanse of wings 17%5 inch.
Jaintia Hills. One example.
Allied to H. subangulata, Warren, = Stegania micans,
Hmpsn.
Genus PRISTOPERA, nov.
Fore wing with costa nearly straight, apex minutely acute,
outer margin curved and very strongly crenulate : hind wing
with outer margin also strongly crenulate, the apex truncate;
palpi upturned, second joint slightly hairy, third joint short.
Antenne bipectinate with fascicles of cilia; the branches
rather short ; tongue and frenulum present. Fore wings with
fovea, cell more than half the length of the wing, disco-
cellular concave ; vein 2 at three fourths, 3 close before end
of cell, 4 from the end, 5 a little below centre of discocellulars,
6 from upper angle of cell, 7, 8, and 9 stalked from shortly
before end of cell, 10 anastomosing with 11, which is given
off from 12: hind wing with vein 3 from before end of cell,
4 from end, 6 from upper end, 7 from betore end, 8 anasto-
mosing with 7 till near middle of cell.
Type P. hepaticata, nov.
Pristopera hepaticata, nov.
3. Of a uniform dark red-brown colour; costa of hind
wings whitish. Fore wings with pale dots on the costa and
crossed by two central oblique pale lines, the inner one bent
inwards on the costa: hind wing with one central pale
transverse line, in continuation of the outer line of the fore
wing. On the underside the colour is also uniform and
more red, with two black transverse central lines across both
wings, obsolete on the hinder margin of fore wings, which is
whitish.
Expanse of wings 1,3, inch.
Central China. One example.
The shape of the wings is somewhat similar to Ocaelophora
bastpuncta, Moore, from Sikkim.
Subfamily Macarrya.
Calletera schistacea, nov.
¢@. Of a uniform dark shining lilac-grey ; costa of fore
wings with many ochreous specks; a brown spot at end ot
310 Col. C. Swinhoe on new Species of
each cell, lines across both wings brown, dentated—first ante-
medial, second discal, third submarginal, the two latter
running evenly and rather close together, marginal line
black ; cilia ochreous, with brown spots opposite the angles
of the crenulated border. Underside pure dark grey ; costa
of fore wings marked with black and ochreous ; cell-spots
and cilia as above; one transverse dentated line a little
beyond the middle.
Expanse of wings 1,%) inch.
Sarawak, Borneo. ‘I'wo examples.
Family Larentiide.
Cryptoloba etaina, nov.
g. Dull cinereous grey: hind wings as dark as the fore
wings, the latter with white specks on the costa and two
transverse nearly upright brown lines, angled outwards above
their centre and sinvous—the first from the costa before the
middle to the hinder margin at the middle, the second from
the costa at two thirds to the hinder angle—the space between
the first line and the base ochreous in all three examples ;
cilia ochreous, marked with grey.
Expanse of wings 3% inch.
Khasia Hills. ‘lhree examples.
Allied to C. cinerea, Butler ; lines somewhat similar, but
the wing-colour is very different and the insect is very much
smaller.
Family Sterrhide.
Perizera maculifera, nov.
?. Uniform ochreous grey, sparsely irrorated with brown ;
a large round brown spot at the end of each cell; a large
brown spot midway between the cell-spot and the outer
margin on the hind wings, the irrorations collected and
formed into small clusters on various parts of both wings,
and black marginal dots in the interspaces.
Expanse of wings 1,°5 inch.
Dawson, Queensland. One example.
Placed provisionally in the genus Pertxera until a male is
procured to determine it properly; it is, however, such a
curiously marked insect it deserves a description,
Eastern and Australian Moths. alt
Family Quadrifide.
Subfamily Pozypzsuiy2.
Melioptis ankara, nov.
9. Head, thorax, and fore wings olive-grey, the latter
with a large black spot touching the costa a little before the
apex; a small spot in the middle below it and another near
hinder angle; a grey suffused band across the wing, its outer
edge touching the three spots; a grey lunule at the end of
cell and two outwardly curved sinuous grey lines—ante- and
postmedial—each terminating in a black spot on the costa ;
marginal lunules grey: hind wings grey ; a medial brown
thin band corresponding to the outer line of the fore wings ;
a broad blackish marginal band ; the space between the bands
white; cilia of both wings grey, with a pale basal line.
Underside nearly white ; fore wing with a brown spot at end
of cell; both wings with a central, outwardly curved, thin
brown band and broad brown marginal borders and white
cilia.
Expanse of wings 1} inch.
Quetta, one example ; Kandahar, one example.
Family Hypenide.
Subfamily Dezrorprvz.
Catada charalis, nov.
g. Of a uniform pale purplish-brown colour ; palpi with
a white band and end of second joint and a thin white collar:
fore wings with three rather prominent, upright, dark brown
bands—first antemedial, second a little beyond the middle,
third submarginal; the first irregularly outwardly curved, the
second and third with two blunt outward angles and out-
wardly edged with white ; a brown spot at the end of cell:
hind wings without markings. Underside much paler; fore
wings with a white streak at costa representing second band,
submarginal white dots and white dots on cilia; hind wings
with discal and submarginal sinuous brown bands, outwardly
edged with white, and white dots on cilia; both wings with
white lunules at the end of each cell.
Expanse of wings 1 inch.
Coomoo, Queensland. Five examples.
Allied to Catada vagalis, Walker.
312 Col. C. Swinhoe on new Species of
Subfamily Hrrrwrvz.
Genus ABRIESA, nov.
Fore wing with costa nearly straight, apex minutely
pointed, outer margin acutely angled at vein 4, concave
between the angle and apex, nearly straight to hinder angle,
which is well angled; hind wing with the outer border
crenulate ; fore wing with veins 3, 4, and 5 from lower angle
of cell, 6 and 7 from upper angle, 8 and 9 anastomosing to
form the areole, 10 from 9 near apex, 11 from centre of cell,
12 free; hind wing with 3 and 4 from lower angle of cell, 6
and 7 from upper angle, 8 free, straight; palpi upturned far
above head, second joint well clothed, with long hair above,
third joint as long as second, with short stiff hairs below and
very long tufts of hairs towards extremity above ; hind tibiz
with a pair of stout terminal spurs, one short, the other very
long ; legs well clothed.
Type A. derna, nov.
Abriesa derna, nov.
@. Pale pinkish fawn-colour above; palpi with brown
bands, the tufts of hairs white: fore wings thickly striated
with grey, collected together in places, forming several in-
complete transverse bands, and having three oblique white
transverse streaks towards the base; a brown straight line
inwardly edged with white near the outer margin from the
costa close to the apex to the hinder angle: hind wings
with a brown spot at end of cell suffused with reddish grey ;
a broad discal brown band and marginal black lunules on
both wings. Underside nearly white; fore wings suffused
with pink; hind wings striated with brown; both wings
with two indistinct brownish bands, central and discal ; fore
wings with a white band from near apex to hinder margin.
Expanse of wings 13 inch.
Dawson, Queensland. One example.
Family Thyridide.
Rhodoneura melilialis, nov.
3d. Costa straight, outer margin oblique, evenly curved,
shape as in 2. bastialis, Walker. Pure white; antenne red,
as is also the top of head, fore part of thorax, and a broad
costal stripe on fore wings, the latter, however, becomes
obsolescent on the apical third; both wings striated with
ochreous; on the fore wings the striations are very uniform ;
there is another more indistinct red stripe below the costal
Eastern and Australian Moths. 313
stripe, terminating in a red spot at end of cell; on the hind
wings the striations form a rather prominent and_ broad
central band, and on both wings are submarginal red dots,
Underside as above, but the striations are darker, more red,
and more diffuse.
Expanse of wings 75 inch.
Dawson, Queensland. One example.
Family Schenobiide.
Cirrhochrista rauma, nov.
3. Antenne and palpi chestnut-brown ; a stripe on palpi
above and top of head white; a large chestnut-brown spot
behind the head; body and wings shining white: fore wings
with a broad costal chestnut-brown stripe, which becomes
thin towards the apex; a large wedge-shaped spot of the
same colour attached to the costal stripe, deseending to the
end of the cell, a stripe of same colour on the outer margin,
and a chestnut cilia: hind wings with some chestnut-red On
a small portion of the outer margin and cilia below the
middle. Underside and legs pure white, without markings.
Expanse of wings 14 inch.
Brisbane. ‘Two examples.
Allied to C. etherialis, Led.
Family Pyraustide.
Subfamily DicxocrocriwZ.
Hedylepta vildersalis, nov.
g 9. Of a uniform olive-grey, glossy and nearly semi-
hyaline: fore wings with a brown dot in the cell outside the
first line, a brown lunule at the end: fore wings crossed by
two brown lines, the outer one crenulated, deeply bent
inwards underneath the cell-lunule, then straight to hinder
margin beyond the middle ; the inner line at one third, out-
wardly curved: hind wings with the inner line from costal
third straight to anal angle; the outer line from the costa
beyond the middle straight down to vein 3; both wings with
a brown marginal line and brown cilia, interlined with white.
Expanse of wings 1 inch.
Cherra Punji. ‘Twenty-two examples.
The lines are disposed as in H. cuprealis, Moore, from tle
Andamans, of which I have many examples, but the lines are
crenulated and not smooth as in that species, and the colora-
tion is altogether different.
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. yal
314 Dr. R. Broom on a new Theriodont Reptile
XL.—On a new Theriodont Reptile (Ictidosuchus primevus)
from the Karoo Beds, South Africa. By R. Broom, M.D.,
B.Sc.
In the lower beds of the Karoo formation near Pearston I
have been fortunate in recently discovering the remains of a
small Theriodont reptile of a remarkably generalized type.
The remains were met with in the beds of indurated shale,
and though, unfortunately, much of the skeleton has been
weathered away and irretrievably lost, sufficient has been
left in the rock to give a very good idea of the type. The
parts discovered consist of a moderately complete lower jaw,
a considerable portion of the maxilla with teeth, and a number
of other portions of the skull; the almost perfect scapula,
coracoid, and precoracoid ; the humerus and radius; a femur
and a tibia and fibula ; two or three imperfect vertebra and
ribs; and a number of other fragmentary remains. Unfor-
tunately the various bones are mingled together in almost
inextricable confusion—the tibia and fibula lying across the
scapula and the femur right across the humerus—so that it is
a matter of great difficulty to develop the one bone without
injuring the other. As the form is of much interest, and the
thorough examination of the remains will necessarily take
considerable time, 1 have thought it advisable to give a short
preliminary account of the animal.
Lower jaw of Ictidosuchus primevus, nat. s1ze.
The lower jaw resembles much more the Theriodont jaws
from the Upper Karoo beds than any of those hitherto found
from the Karoo Beds, South Africa, 315
in the Lower. In its general form and proportions the
resemblance to the jaw of Tribolodon from Lady Frere (Phil.
Trans, 1894, B, pl. Ixxxviii. fig. 6) is very marked. The
horizontal ramus of the jaw is long and rather slender and
moderately uniform in depth. The coronoid process is very
well developed, long and fairly thick, and makes an angle of
about 120° with the ramus. Almost the whole lower jaw
appears to be formed by the dentary, the splenial being a
feeble splint and the articular, which is lost, probably not
large.
Perhaps the most remarkable feature of the form is the
simple structure of the teeth. Seven teeth still remain in
the jaw, and though it is possible that one or two others are
lost, those that remain show that there is no marked distinc-
tion between incisors, canines, and molars, all the teeth being
modifications of the simple pointed Saurian type. All the
teeth are feebly ribbed, and the anterior teeth differ from the
posterior only in being longer and moderately sharp, while
the latter have rounded apices.
The fragment of the maxilla shows the upper teeth to be
very similar to those of the lower jaw.
The scapular arch resembles that of Rhopalodon (Phil.
Trans. 1894, B, p. 703) more closely than that of Dicynodon.
The scapula, while expanded and flat above, is narrow in the
middle. There is no very distinct acromion. Inferiorly the
scapula is much expanded and forms a large articulation with
the precoracoid. ‘The precoracoid is considerably larger than
the coracoid and differs from that in Dicynodon in not
entering into the glenoid cavity and in completely surrounding
the precoracoid foramen. ‘I'he coracoid closely resembles
that in Dicynodon.
The humerus is very mammal-like and bears considerable
resemblance to that of Gomphognathus (Phil. T'rans. 1895, B,
p- 29), though less robust. here is a large epicondylar
foramen, and the whole lower half of the bone so far as is
preserved is much like that of the Phalangers. Above, there
is a sharp delto-pectoral crest, somewhat less prominent than
that in Gomphognathus, because less directed outwards from
the bone.
The femur is slightly longer than the humerus, and, like it,
shows much resemblance to the mammalian types. There is
a well-marked great trochanter, which forms a prominent
trochanteric ridge more resembling the condition in some of
the “Edentates and Marsupials than that of either of the
Monotremes. On the whole, however, the affinities of the
bone are mainly with the type seen in Hehidna; and if the
21*
316 © Mr. R. I. Pocock on some
femur of this Monotreme were a rounded instead of a flattened
bone, the general resemblances between it and that of the
fossil form would be very marked. ‘The lower end of the
femur especially is flattened and quite Monotreme-like.
The tibia and fibula are both long simple bones, con-
siderably longer than the radius. ‘The fibula is only about
half the thickness of the tibia.
As soon as a thorough examination has been made of the
more fragmentary remains, and the more perfect elements
more thoroughly cleared of matrix, an endeavour will be made
to describe in detail, with figures, the various remains of this
primitive type, for which I propose the name Ictidosuchus
primevus, gen. et sp. n.
The following are some of the principal measurements of
the bones :—
millim.
Lower jaw : symphysis to coronoid process ........ 94+
Depth of jaw at largest tooth (canine P)............ 10
Heipht.ot largest toothy. 24 aks, < cto spew. tieoera ate 5
Antero-posterior diameter of largest tooth .......... 2:3
Imengthsonscapel a coc ny earns ws cee archery 73+
Width of upper part of-scapula, cas... 225 2 tee 19
Width in narrow middle region ..........000ee eee 9
Wiadtheat basoi.: smi zi -y absentee nats es bi geeteo rn 28
Length of humerus (72 millim.+) ........ probably 82
Width of humerus near middle ............2.0006 9
Length of femur (86 millim.+) .......... probably 90
Width of femur near middle. ..225% AcistoGaten ee stercee 8
Length of dubia: «Gh <dncc xe tnjeciauie ere Altes ost 94
Width of tibiatat lower third (...<):scs0e os sage ok 9x5
Pearston, 8. Africa.
XLI.—Some new Arachnida from Cape Colony.
By R. I. Pocock.
Order SOLIFUGA.
Genus Sotpuea, Licht.
Solpuga Schénlandi, sp. n.
3 .—Colour a uniform pale yellow, as in &. venator;
width of head less than tibia of palp and than patella or tibia
of fourth leg. Form of mandible recalling that of S. Dar-
Lingit, but with upper jaw armed with only one minor tooth
in front of and remote from the two large distal teeth; the
new Arachnida from Cape Colony. 517
terminal fang shorter, directed more upwards, and less curved
at the tip, its upperside more strongly hollowed at base ;_ basal
portion of flagellum high, conical, with rounded summit and
straight anterior border; distal portion of flagellum rising
above the proximal terminal tooth, short, scarcely surpassing
the basal portion when reclining backwards, lightly curved
or narrowed distally, with apex truncate and furnished with
a minute process; on the jaw in front of the base of the
flagellum there is a small upstanding tooth.
Total length 35 millim.; width of head 9°5; length of
patella of palp and also of tibia and tarsus 14, patella and
tibia of fourth leg 12.
Loc. Grahamstown (Dr. Schénland).
Genus Daista, C. Koch,
Desia Bernardi, sp. n.
? .—Prevailing colour yellow, clouded with greyish black
laterally ; tubercle black ; mandibles with faint fuscous lines ;
palp infuscate, with the basal three fourths of the femur
yellow ; apex of femur and base of patella of third and fourth
legs lightly infuscate.
Abdomen with three narrow dorsal stripes.
Mandible with upper jaw furnished with two minor teeth,
the distal major teeth subequal; lower jaw with one minor
tooth close to base of posterior major tooth.
Palp with tibia lightly fusiform, armed as in D. Letpolde?,
Pure. ; tarsus rather more than one third the length of the
protarsus ; patella armed below with two stout seta inside
and three thinner sete outside.
Carapace as long as patella of palp, a little shorter than its
tibia and tarsus and than patella of fourth leg.
Abdomen: third segment furnished beneath on each side of
the median sternal plate close to the posterior border with a
transverse series of four slender but strong and sharp, curved,
claw-like spines raised on low elevations of the integument.
Immediately adjacent to these on the anterior portion of the
underside of the fourth abdominal segment there is a trans-
verse row of six or seven smooth, shining, subcylindrical
ridges, separated by deep intervening grooves,
Total length 13 millim.; width of head 3; length of
palp 10.
Loc. Hex River Valley, Cape Colony (H. MZ. Bernard).
No structure on the abdomen resembling that described
above has been mentioned in the diagnoses of any of the
previously described South-African species of Desia.
318 Mr. R. I. Pocock on some
Order ARANE ZK.
Family Theraphoside.
Genus PTERINOCHILUS, Poc.
Pterinochilus Lugardi, sp. n.
&$ .—Carapace distinctly longer than patella, tibia, and tarsus
of palp, shorter than patella and tibia of first leg, subequal to
those of fourth and only just exceeding protarsus of fourth.
Ocular tubercle and clypeus practically as in P. voraz,
Foe:
Front leg with tibia not inflated, the spur projecting in-
ternally in such a way that a large space is left between the
spur and the base of the protarsus when the latter is flexed
on the tibia.
Palpal organ much like that of P. voraz, but with the
base of the spine less stout.
Protarsus of leg lightly sinuous, not nearly so strongly as
in P. voraz.
Total length 24 millim.; carapace 12, first leg 41, second
37, third 34, fourth 43.
Loc. Kwebe Hills, near Lake Ngami (Capt. E. J. Lugard,
Ds.0:),
Pterinochilus Schénland, sp. n.
6 .—Carapace about as long as patella, tibia, and tarsus
of palp, shorter than patella and tibia of fourth leg, just
equal to those of second, not or scarcely exceeding protarsus
of fourth, equal to protarsus and half the tarsus of the third.
Legs with patella and tibia of fourth scarcely longer than
protarsus and tarsus of first; tibia of first not inflated; the
spur directed forwards in such a way that it is almost in
contact with the inner side of the base of the protarsus when
the latter is flexed ; protarsus not sinuous.
Spine of palpal organ stouter than in the preceding species,
compressed, distally sinuous, and suddenly narrowed to a
point just before apex.
Total length 18 millim.; carapace 9, first leg 30, second
leg 29, third leg 25, fourth leg 33.
Loc. Grahamstown (Dr. Schénland).
Longer in the leg than P.nigrofulvus, Poc., and with different
palpal organ.
new Arachnida from Cape Colony. 319
Family Ctenizide.
Genus STASIMOPUS, Simon.
Stasimopus Schénlandt, sp. n.
Nearly allied to §. oculatus, Poc., in disposition of eyes &c.,
but recognizable by the absence of the apical band of spines
on the lower side of the protarsus of third leg and by the
restriction of the spinules on the lower side of the protarsus
of fourth leg to a small posterior tuft.
Total length up to 35 millim.
Loc. Grahamstown (Dr. Schénland).
Genus HERMACHASTES, nov.
Allied to Hermacha and Nemesia.
Fovea large, transverse, or a little recurved.
Rastellum consisting of numerous close-set stout bristles
and spines.
Labium armed with a row of cusps; coxa of palp with
about 20-25 strong short scattered cusps; inferior claw of all
the legs long and distinct; superior claws with two rows of
strong teeth.
Posterior sternal s¢gi//a submarginal.
Apical segment of posterior spinners about half as long as
the second.
Male with longer legs than female; labium and maxille
unarmed. ‘Tibia of first leg armed on the inner side with
two low prominences, each bearing a stout curved spine.
Tarsus of palp short, truncate.
Hermachastes collinus, sp. n.
? .—Colour olive-yellow or brown on carapace and limbs ;
femora darker than the rest of the leg-segments; abdomen
ashy black, variegated above with pale patches and spots,
paler below.
Carapace longer than patella, tibia, and tarsus of palp, and
than patella and tibia of first or fourth leg. Palp with tibia
and tarsus spined. Patella and tibia of first and second legs
weakly, protarsus more strongly spined ; patella, tibia, and
protarsus of third spined; patella of fourth unspined.
Tarsi of legs unspined.
3 .— Carazace as long as patella and tibia of first leg,
shorter than those of fourth.
320 Mr. R. I. Pocock on some
Falpi unspined; tibia swollen, bristly below; bulb sub-
globular, spines short, stout, lightly curved, pointed at apex ;
protarsus of first leg lightly sinuous at the base.
Measurements in millimetres.— 2 . Total length 18; cara-
pace 7, first leg 15, second leg 14, third leg 12, fourth leg 17°5.
; 3. Total length 12; carapace 6:5, first leg 19, fourth
eg 23.
Loc. Table Mountain (J. Hull).
Genus BESSsIA, nov.
Carapace smooth, fovea deep, procurved; ocular area
transversely oblong, more than twice as wide as long, parallel-
sided ; lateral eyes narrowly separated; eyes of anterior line
procurved, medians a little smaller, with their anterior edges
on a line with the centres of the laterals.
Rastellum consisting of many long slender spines over-
hanging the base of the fang.
Mandible armed below with a single row of about 15-16
teeth, a few smaller cusps at the posterior end of the row;
fang long and slender.
Labium twice as wide as long, armed, like the base of the
maxilla, with many close-set cusps; maxille lightly im-
pressed.
Sternum with posterior sigilla of medium size, elongate,
about their own length from the margin of the sternum and
twice that distance apart.
Legs not scopulate, anterior pairs weaker than posterior,
with protarsi much longer than tarsi; first leg scarcely
spined ; protarsus of second with some strong spines below ;
third leg with a few spines on patella and tibia above, about
6 in two rows on protarsus; fourth with protarsus nume-
rously spined; claws armed with 3-4 basal teeth. Posterior
spinner with apical segment small, much smaller than
second, .
According to Simon’s tabulation of the genera of “ Cyrtau-
cheniex,” this new genus seems allied to Amblyocarenum
and Aptostichus, especially to the latter.
Bessia fossoria, sp. n.
Colour of carapace and legs pale mahogany-red.
Measurements in millimetres.—Total length 12; carapace 6,
palpus 7, first leg 10, second leg 9, third leg 11, fourth leg 14.
Loc. Port Elizabeth (Dr. Broom).
new Arachnida from Cape Colony. 321
Family Scytodide.
Genus Sicarrus, Walck.
Sicarius spatulatus, sp. n.
3 .—Integument of carapace and sternum castaneous ;
legs yellower; abdomen testaceous; in nature the whole
body is covered with particles of sand &c. and presents a
uniform greyish-black tint.
Width of carapace about equal to length of patella and
tibia of third leg, a little longer than tibia of first ; median
eyes on a distinct tubercle, longitudinally elliptical, space
between them less than their transverse diameter ; lateral
angles of head prominent, interval between the lateral eyes
less than their diameter.
Legs strong, first more than three times as long as width
of carapace, third less; femur of first narrowed at base,
swollen in middle.
Falpi short ; tibia globular; tarsus short ; spine of palpal
organ stout, curved, distally expanded, and triangularly
spatulate, with truncate extremity.
9? .—Like male, but with legs shorter, first less than three
times the width of the carapace, which is about equal to the
patella and tibia of the second.
Total length (¢) 9 millim., width of carapace 4, length
of first leg 14.
Loc. Port Elizabeth (H. A. Spencer, Dr. Broom).
Differs entirely from S. Hahnit, Karsch (sec. Simon), in
the form of the palpal organ.
Family Caponiide.
Genus CAPonrA, Sim.
Caponia secunda, sp. n.
9 .—Resembling C. natalensis (Cambr.) in colour and
most structural features.
Carapace rugulose, the anterior median and anterior lateral
eyes forming a line which is distinctly recurved, the posterior
edge of the medians being on a level with the centres of the
laterals ; the three eyes which form the curved lateral line
very unequal in size, the anterior lateral being the largest
and the posterior lateral the smallest, the latter being only
‘about half the area of the former,
According to Cambridge, in C. natalensis the anterior
medians and anterior laterals form a straight transverse line.
322 Mr. R. I. Pocock on some
Simon represents this line as slightly procurved. Moreover,
the latter author declares the lateral eyes to be equal, the
former says “ nearly equal.”
Total length 8 millim.
Loc. Grahamstown (Dr. Schénland).
Family Ereside.
Genus ERESUS.
Eresus Spenceri, sp. n.
Much smaller than H. fumosus and differently coloured.
Hairy clothing olive-grey; integument of carapace and
mandibles deep brown, of legs deep reddish, of abdomen
olive-yellow ; narrow white rings round the dorsal sigilla.
Carapace as long as tibia, protarsus, and tarsus of first
leg ; width of head about equal to patella and tibia of first
leg ; posterior median eyes large, about two diameters apart.
Vulva with large subcircular lateral pits, which are at least
equal to the width of the median septum. -
Total length 13 millim.
Loc. Port Elizabeth (H. A. Spencer).
Genus DRESSERUS, Sim.
Dresserus Darlingi, sp. n.
Allied to D. obscurus, Poc., but with shorter legs, the
width of the head being distinctly greater than the patella
and tibia and than tibia and protarsus of first leg ; head also
noticeably higher above the line of eyes; median septum of
vulva with its lateral edges more converging posteriorly.
Total length 14 millim.; carapace 6°3.
Loc. Mashonaland: Enkeldoorn and Mazoe (J. ff. Dar-
ling).
Dresserus olivaceus, sp. n.
Allied to D. Darlingi, but much darker in colour above
and below, being of a silky greenish black. Head less flat
above, more evenly convexly rounded from before backwards ;
the sides of the median sclerite of vulva more parallel and
the lateral impressions narrower.
Total length 14 millim.
Loc. Grahamstown (Dr. Schénland).
new Arachnida from Cape Colony. 323
Dresserus collinus, sp. n.
Differing from D. obscurus, olivaceus, and Darlingi in
having the posterior median eyes very large and separated
from the anterior laterals by a space which is only about
equal to twice the diameter of the medians; width of head
slightly exceeding length of patella and tibia of first leg.
Median area of cribellum subequal to the external area.
Total length 11 millim.
Loe, Table Mountain (H. A. Spencer and J. Hull).
Family Amaurobiide,
Genus AUXIMUS, Sim.
Auximus capensis, sp. n.
?.—Colour. Carapace and limbs pale castaneous, the
latter infuscate distally, hairy clothing pale olive; mandibles
black; abdomen symmetrically spotted above; legs palely
castaneous, clothed with pale olive hairs, the anterior pairs
distally infuscate.
Eyes of posterior line slightly procurved, medians smaller
than laterals and slightly nearer each other than to the
laterals; eyes of anterior line slightly procurved, medians
smaller, about a radius apart and a diameter from the laterals.
Mandibles with posterior border of fang-groove armed with
5-8 teeth, those in the middle of the row largest.
Legs 1, 4, 2, 3.
Vulva consisting of a large, shallow, transversely ellip-
tical pit, the rim of which is semicircularly incurved on each
side behind.
Total length 13 millim.
Loc, Port Elizabeth (H. A. Spencer); Cape Town (H. A.
Spencer) ; Table Mountain (J. Hull).
Auzximus hottentottus, sp. n.
Smaller than the preceding; the head darker posteriorly
and at the sides than in the middle; sternum darker than
coxe ; legs indistinctly annulate below.
Eyes a little more widely separated, the whole ocular area
thus wider ; eyes of anterior line subequal.
Vulva consisting of a large transverse lightly convex plate,
marked on each side at the margin with a circular pit and in
the middle by a longitudinal groove, bordered behind by an
anteriorly emarginate crest.
Total length 10 millim.
Loc. Little Namaqualand, Garies (Dr. Broom).
Mr. R. I. Pocock on some
(sh)
bt
ne
Family Argiopide.
Genus GASTERACANTHA, Sund.
Gasteracantha Spencer?, sp. n.
Colour. Carapace and mandibles deep red ; sternum black,
with anterior yellow spot; legs with coxe, trochanters, and
femora red, the remaining segments black ; abdomen black
below, with yellow spots; uniformly yellow above, with
dark sigilla, and slightly clouded with fuscous at the base of
the anterior spine and posteriorly at base of median spine ;
spines reddish, with black tips.
Abdomen not twice as wide as long; posterior spines as
large as in G. ensifera, Thor. (?=G. versicolor, Walck.) ;
anterior and median spines much shorter than in that
species ; the medians scarcely as long as the posteriors, straight
and strong, and about four times as long as the anteriors.
Loc. East London (H. A. Spencer) ; also a closely allied
form from Grahamstown (Schénland).
Genus ARANEUS, L.
Araneus mensamontis, sp. n.
? .— Colour much as in A. Rumpfi or A. nauticus ; cara-
pace yellowish or red, darker at the sides; legs banded,
femora of anterior pairs reddish or black; abdomen black
below, with a pair of large yellow spots in front of spinners.
Eyes of anterior line straight, medians about three times as
far from the laterals as from each other.
Tibia and protarsi of anterior /egs armed with many spines ;
tarsi also spined.
Vulva somewhat as in A. haploscapus, but the svape
shorter and not bent at right angles, but meeting the basal
portion at an obtuse angle.
Total length up to 15 millim.
Loc. Table Mountain (J. Hull); Port Elizabeth (Dr.
Broom).
Araneus Graemit, sp. te
? .— Colour. Carapace yellow, black on thoracic portion
and on middle of head; legs yellow, banded with black ;
patella and tibia of anterior legs black below ; abdomen
olive, varied with black and white above, black below, with
a broad yellow band on each side of the middle.
Eyes of anterior line strongly procurved. Carapace about
new Arachnida from Cape Colony. 325
as long as patella and tibia of first leg; tibia of first armed
with about 6 spines in front below, protarsus with about 8.
Vulva with its basal portion very short and projecting
distally on each side of the base of the scape as a distinct
horny process; scape rather short, lightly constricted laterally
near its base.
Total length 10 millim.
Loc. Grahamstown (Dr. Schénland).
Family Zodariide.
Genus CYDRELICHUS, nov.
Allied to Cydrela and Cesetius, with the eyes of the ante-
rior line approaching those of Cesetius, the laterals being
from three to four diameters apart; those of the posterior
line recurved to about the same extent as in Cydrela.
Type C. Spencert.
Cydrelichus Spencer, sp. n.
? .—Colour. Carapace deep castaneous; legs clearer,
scantily clothed with whitish and blackish hairs; abdomen
ashy black, covered above and below with a scanty clothing
of whitish and darker hairs and ornamented beneath with
four pale lines.
Palpi strongly spined distally.
Anterior two pairs of legs weakly spined, posterior two
pairs very strongly spined.
Vulva (? subadult) consisting of a small transversely oblong
plate, impressed with a pair of deep irregularly oval pits,
separated by a partition and defined externally by a dark
rim.
Total length 10 millim.; carapace 4, first leg 8, fourth 10.
Loc. Port Hlizabeth (Hl. A. Spencer).
Genus CHARIOBAS, Simon,
Chariobas lineatus, sp. n.
Colour. Carapace bright reddish yellow, with a median
longitudinal black band and a much narrower black marginal
band; sternum black at the sides, with a median pale stripe ;
abdomen chalky grey, with a median longitudinal dorsal
black band extending to the spinners and continuing the
black band on the carapace; in the ventral median line a
similar stripe which gradually expands posteriorly ; spinners
326 Mr. RB. I. Pocock on some
black ; legs orange-yellow, darker apically, and with black
tarsal and protarsal scopule.
Carapace about as long as tibia and protarsus of first leg.
Measurements in millimetres.—Total length 11°53; length
of carapace 4, width 1°6; length of abdomen 7°5, width 2 ;
length of first leg 7°5, of second 6°5, of third 5, of fourth 7.
Loc. King Williamstown (Stenning).
Family Agelenide.
Genus AGELENA.
Agelena ocellata, sp. n.
?.—Colour. Integument ochre-brown; carapace with
indistinctly defined broad median and lateral white bands ;
legs with white hairs, the femora distinctly banded ; sternum
with pale median stripe; abdomen rubbed; the dorsal in-
tegument ornamented with a pair of bright yellow spots,
some black spots arrange in two longitudinal lines, and
some small dark spots laterally.
Anterior median eyes a little longer than laterals, their
inferior borders on a level with the superior borders of the
laterals.
Vulva marked with a pair of obliquely oval pits separated
by a broad posteriorly clavate partition.
Total length 7 millim.; carapace 3:2; first leg 9.
g.—With much longer legs than female. Palp with
patella short, armed externally with a pair of subequal short
apophyses, the upper of which is more acute than the under ;
tibia shorter than patella, but higher, armed below externally
with a slender tooth-like process; tarsus much longer than
patella and tibia taken together.
Total length 6 millim.; carapace 3; first leg 13.
Loc. Table Mountain (J. Hull).
Genus Roruus, Sim.
Rothus auratus, sp. n.
@ —Much paler than £2. vittatus, Sim. Integument testa-
ceous ; carapace covered with yellowish hairs, with a paler
median brown-bordered band; legs covered with yellowish
hairs ; abdomen similarly covered, without bands.
Head \ess constricted and frontal line of eyes less promi-
nent than in &. vittatus.
new Arachnida from Cape Colony. 327
Vulva very similar to that of #. vittatus, but the median
sclerite marked with a deep median longitudinal impression.
Total length 14 millim.; carapace 5°5; first leg 20°5.
Loc. Little Namaqualand, Garies (Dr. Broom).
Genus Lycosa, Latr.
Lycosa hectoria, sp. n.
? .—Colouring apparently as in Z. capensis, Simon, but
with tibia of fourth leg strongly banded below with black at
base and apex; tibia of third leg similarly but much less
distinctly banded; tibia of second and first legs yellowish
brown below, with pale base; sternum, coxe, and lower side
of abdomen clothed with yellowish-grey hairs, the abdomen
with faintly defined median band behind the vulva.
Structurally also apparently as in ZL. capensis, except that
the teeth of the posterior border of the fang-groove are sub-
equal in size.
Vulva wider than long, the median keel about half the
length of the transverse bar, the pits subcircular and defined
by a ridge curved like a ram’s horn.
Total length 16-19 millim.; carapace 10; first leg 23,
fourth leg 30. ;
Loc. Table Mountain (J. Hull).
Lycosa subvittata, sp. n.
?.—Smaller than L. hectoria, but very similar in colour
on the dorsal side, though perhaps on the whole darker; the
two yellow stripes on the abdomen strongly defined; ventral
surface of abdomen with a median longitudinal black stripe,
broader in front behind the vulva and narrowing posteriorly
and defined on each side by a broad yellow stripe, which is
itself defined by a darker stripe composed of blackish spots ;
coxe infuscate like the sternum; legs infuscate below,
scantily clothed with pale hairs, the tibie not distinctly
banded.
Structurally as in L. hectoria, but with vulva longer than
broad, the median keel broad in front, narrowed behind, and
as long as the transverse bar, the depressions longitudinally
ovate, three times as long as wide.
6 .—Like female in coloration, except that the black on
the ventral surface of the abdomen spreads laterally behind
the epigastric fold so as to cover the underside, the epigastric
area being black in the middle, pale at the sides; coxze much
paler than sternum, as in young female.
328 Mr. R. I. Pocock on some
? .—Total length 14 millim.; carapace 7; first leg 19,
fourth 16.
3S .—Total length 11 millim.; carapace 6°5; first leg 19,
fourth 23.
Loc. Port Elizabeth (H. A. Spencer) ; Table Mountain
(J. Hull).
Lycosa Schénlandi, sp. n.
? .—General colour reddish yellow, carapace with two dark
stripes; abdomen mottled black above and at sides, greyish
below, with median and lateral blackish stripes, much as in
L. Spencert, Poc.; coxe and legs reddish, clothed with pale
hairs ; legs spotted and irregularly banded with black ; base of
mandible covered with yellowish hairs of the same colour as
those on the face.
Eyes of anterior line lightly procurved, their inferior edges
in a straight line, medians a little longer than the laterals,
posterior median separated by a space which is much less
than their diameter.
Vulva with very narrow median keel and stout, posteriorly
convex, and projecting cross-bars, on each side of the median
keel there is a narrow outwardly curving crest, forming the
inner border of the shallow oval longitudinal impression.
Measurements in millimetres.—Total length 15; carapace
6:5; first leg 16, fourth leg 21.
Loc. Grahamstown (Dr. Schénland).
Lycosa promontorit, sp. n.
Allied to Z. Schénlandi, but darker in colour, carapace
with narrower whitish submarginal stripe; abdomen dark
reddish grey above, mottled with black and marked with a
short paler median stripe in front; entirely black below,
coxa and sternum deep blackish brown; legs mottled with
darker and lighter spots; base of mandible scantily clothed
with yellow hairs.
Eyes of anterior line straight by their lower borders,
medians noticeably the larger; posterior medians scarcely
wider than anterior line, separated by considerably less than
their diameter ; distal tooth on posterior border of fang-groove
very much smaller than the others. .
Vulva with rim thick and curved like a horseshoe, the
median bar very broad in front, narrow behind, about half
the length of the transverse bar.
Measurements in millimetres.—Total length 13 ; carapace 6;
first leg 14°5, fourth leg 17°5.
Loc. Wynberg (H, A. Spencer) ; Table Mountain (J,
Hull).
new Arachnida from Cape Colony. 329
Lycosa algoensis, sp. n.
Colour. Carapace normally coloured, but the lateral stripes
not so distinct as in L. Schénlandi; upperside of abdomen
black in the middle, with a yellow stripe formed of two pale
patches, recalling the markings on Ocyale atalanta, on each
side ; ventral surface banded as in L. Schénlandi; coxe and
sternum yellowish red; legs pale, obscurely mottled ; man-
dibles covered with greyish hairs at base.
Posterior median eyes very large, scarcely a radius apart ;
anterior medians much larger than anterior laterals.
Vulva very abnormal, consisting of a narrow median keel,
which posteriorly runs out into a strong conical process and is
flanked on each side by asubspherical black prominence, which
is bordered externally and below by a sinuous ridge continuous
in the middle line with the anterior end of the median keel.
Measurements in millimetres.— Yotal length 13; carapace 7 ;
first leg 15°5, fourth leg 19.
Loe. Port Elizabeth (H. A. Spencer).
Lycosa bessiana, sp. n.
gd. Resembling Z. algoensis, but with the whole of the
ventral surface of the abdomen olive-black, the sternum also
infuscate with a central darker line; the legs uniformly
yellowish red, not banded or mottled with dark spots ; upper-
side of abdomen marked with two very distinct, broad, yellowish
bands uniting in front, separated by a dark ventral stripe,
and bordered externally by a darker ill-defined stripe ; inferior
portion of lateral surface white ; dark bands on carapace
broader than pale bands.
Measurements in millimetres.—Total length 9; carapace 5;
first leg 14, fourth leg 17.
Loc. Port Elizabeth (H. A. Spencer).
Family Oxyopide.
Genus PeucetiA, Thor.
Peucetia maculifera, sp. n.
9 —Oolour. Carapace greenish, with short radiating dark
stripes ; clypeus and mandibles without bands ; an abbreviated
and interrupted dark stripe on each side of the head above
the basal mandibular spot; legs yellow, femora and cox
minutely and rather thickly spotted at the bases of the spines
and hairs; tibie and protarsi with larger and fewer spots at
the base of the black spines, a faint blackish ring round distal
end of patella and basal and distal end of tibia; sternum
Ann. & Mag. N. Hist. Ser. 7. Vol, vi. 22
330 Mr. R. I. Pocock on some
green, minutely spotted at base of hairs; abdomen suffused
with rosy pink above, yellower in front, greenish yellow
below, paler in middle line.
Carapace as long as patella and tibia of third leg and three
fourths of the protarsus of the first; clypeus almost vertical.
Abdomen oval, not twice as long as broad.
Legs moderately long, first about five times, fourth nearly
four times as long as carapace.
Vulva consisting of a pit crescentic in front and filled
posteriorly by a pair of black oval sclerites, separated by a
median crest and each marked behind by a deep pit.
Total length 13 millim.; carapace 5°3; first leg 26, fourth
leg 20.
ere King Williamstown (Stenning).
Differs in colouring, form of vulva, &c. from the rest of the
S. African species.
Family Heteropodide.
Genus SPARASSUS.
Sparassus Schénlandi, sp. n.
Colour of integument yellowish, clothed with whitish hairs ;
mandibles a little darker.
Carapace longer than wide, width of head as in S. Bates?,
Poc. ; eyes of posterior line slightly procurved; carapace as
long as tibia of second leg.
Legs 2,1, 4,3; second exceeding first by its tarsus and one
fifth of the protarsus ; third leg scarcely reaching tip of tibia of
second; patellee unarmed, tibize with two pairs of inferior spines.
Vulva consisting of a large transversely oblong plate, the
posterior border of which is semicircularly excised, the
posterior angles of the excision produced inwards into a dark
horny prominence.
Total length 12 millim. ; length of carapace 6:5, of first
leg 22, second 25, third 16, fourth 18-5.
Loc. Grahamstown (Dr. Schénland).
Genus Patysres, L. Koch.
Palystes lycosinus, sp. n.
=Palystes megacephalus, Pocock, Ann. & Mag. Nat. Hist. (6) xvii.
p. 63, 1896 (? P. megacephalus, C. Koch).
3 .—Apparently resembling P. megacephalus, C. Koch, in
colour and most structural features.
new Arachnida from Cape Colony. B51
Tibia of palp armed at its distal end above with a single,
slightly sinuous, basally stout, apically pointed spur directed
forwards and outwards over the base of the tarsus.
¢ —Like male in colour.
Vulva bordered behind by a straight transverse crest, in
front of which there is on each side a short thicker crest ;
the middle of the fore part of the abdomen bilobate.
Total length 21 millim.
Loc. Port Elizabeth (H. A. Spencer).
Palystes cultrifer, sp. n.
3 .—Resembling the preceding, but with two tibial spurs
on the palp—a distal superior slender and upstanding, and a
proximal external, stout, curved, blade-like with a sharp
oint.
; Total length 17 millim.
Loc. Grahamstown (Dr. Schénland).
P. megacephalus, C. Koch, may be identical with either or
neither of these species.
Palystes perornatus, sp. n.
Allied to P. lunatus, Poc., but with the sternum and coxa
a rich golden yellow; the femora of the legs palely olive-
yellow below and mottled with silvery white spots.
Vulva with median horny process much shorter than in
P. lunatus.
Total length 25 millim.
Loc. Queenstown, Cape Colony (Capt. C. K. Bushe).
Genus SELENOPS, Latr.
Selenops Broom, sp. n.
@ .—Allied to S. Krausii, Poc., in curvature and relative
size of eyes of ocular quadrangle, but differing in having five
instead of six pairs of inferior spines on tibice of first and
second legs *, and in the form of the vulva, the lateral lobes of
which are in contact in the middle line and circumscribe a
more transversely cordate pale median sclerite.
Total length 11 millim. ; carapace 5°5; first leg 19, fourth
leg 21.
Loe. Little Namaqualand, Garies (Dr. Broom).
* In the type of S. Arausw the anterior leg on the right side has been
reproduced and shows only five anterior spines beneath.
22*
332 On some new Arachnida from Cape Colony.
Selenops parvulus, sp. n.
9 .—Very small, dark-coloured, thickly mottled with black,
white, and bronze-yellow. ;
Eyes of quadrangle not so strongly recurved as in S. ato-
marius; the posteriors larger than anteriors and with their
lower rims just below the level of the upper rim of the latter,
which are nearly two diameters apart and a little more than
half their diameter from the posterior medians.
Tibi of anterior legs with four pairs of inferior spines,
protarsi with three pairs.
Vulva consisting of a very large plate, representing the
two lobes, impressed by a median sutural line, a small trans-
verse pale area just in front of the plate.
Total length 5 millim.; carapace 2°5; first leg 7, fourth
leg 8.
Loc. Port Elizabeth (Dr. Broom).
Family Thomiside.
Genus Tuomtsus, Walck.
Thomisus Stenningi, sp. n.
? .—Prevailing colour yellow ; carapace with a sharply
defined yellow stripe on each side, starting at the sides of the
head and running upwards to a point on each side of the
fovea; face white, orange-yellow round the anterior median
eyes ; mandibles white, with darker basal spot; a brown trans-
verse stripe in the middle of the femur and tibia of first and
second legs, also some brown near the tip of the protarsus
and tarsus, the dark patches set off with white markings.
Carapace with horns much higher and longer than in
T. albus; the summit of the head between these more strongly
concave, with deeper notch adjacent to posterior median eyes ;
eyes of anterior line more strongly recurved. Protarsal spines
5 in front, 4 behind.
Vulva consisting of a swollen, hairy, indistinctly bilobed
area, in front of which there is a shining plate marked with
a pair of parallel narrow crests, which meet in front and
posteriorly end in a kind of loop, darkened with pigment.
Total length 5°5 millim.
Loe. Pirie Bush, King Williamstown (Stenning).
? Genus Camanicus, Thor.
Camaricus marmoratus, sp. n.
Colour. Carapace deep red, with narrow black inferior
Lord Walsingham on Asiatic Tortricide. 333
border on clypeus and large black patch on each side of head,
involving anterior and posterior lateral and posterior median
eyes; mandible reddish, sternum and mouth-parts pale; coxe
and trochanters of legs black, rest of segments yellow, lined
with black ; palpi with femur and trochanter black, the other
segments yellow and lined with black ; upperside of abdomen
yellow, marked with six transverse black stripes, the anterior
three of which are mesially interrupted; sides of abdomen
with a broad inferior black band, ventral surface pale.
Carapace as broad as long, high, strongly convex, not
narrowed in front. yes of posterior line very slightly
recurved and slightly wider than those of anterior line, the
medians nearly twice as far from each other as either is from
the lateral; eyes of anterior line recurved ; clypeus low, not
much more than one fourth of the distance between the
anterior median eyes.
Mandibles with inferior band of hair.
Legs weak, tibie of first and second with three pairs of
slender spines below, protarsi of first and second with two
pairs of spines below.
Abdomen subglobular, a little longer than wide; vulva
marked with a deep suboblong or heart-shaped pit, which is
broader in front than behind.
Total length 10 millim.
Loc. Grahamstown (Schdnland).
XLII.—Astatic Vorrerrer2.
By the Rt. Hon. Lorp Watsrncuam, M.A., LL.D., F.R.S.
[Continued from p. 243.]
Bactra, Stph.
1006. Bactra lanceolana, Hb.
Aphelia lanceolana, Stgr. & Wk. Cat. Lp. Eur. 251. No. 1006 (1871) ';
Ster. Hor. Soc. Ent. Ross. XV. 252 (1879) *.
Hab. Europe. ArFricaA. AUSTRALIA. New ZEALAND,
S. America. UNITED STATES. ASIA.
Asiatic TurKEY—Brussa, 1V.-V.’; Rhodes Island’; Shar
Devesy (Native Coll. 1893); Palestine (Tristram) ; Keras-
dere, 30 V.’; Jenikeui-Hochebene, 23 V.* TRANSCAUCASIA
—Lenkoran, 18 VI. 1874 (Christoph). PErsta—Asterabad,
15 V. 1873 (Christoph). ‘lTURKESTAN—Krasnowodsk, 5 VI.
334 Lord Walsingham on Asiatic Tortricide.
1872 (Christoph). CorEA—Fusan, 9 VI. 1886 (Leech).
Inp1A—Berhampore, III. 1882 (Minchin) ; Hyderabad, IV.
1886 (Swinhoe) ; Karachi (Swinhoe, 1885). CryLton—Na-
walapitiya (Pole, 1890). SumATRA—Padang Rengas, low
country (Doherty, 1891). CELEBES—Macassar, 500 feet
(Doherty, 1891).
1009 (1). Bactra roseana, sp. n.
Antenne pale cinereous. Palpi slightly recurved, terminal
joint short, not concealed; pale cinereous. Head pale cine-
reous. Thorax smooth, pale cinereous, tegule touched with
greyish fuscous. ore wings elongate, costa evenly rounded,
apex somewhat angular, termen not oblique, tornus much
rounded; pale greyish cinereous, speckled, streaked, and
shaded with pale greyish fuscous, the dorsum shaded through-
out with greyish fuscous; the apical part of the wing to the
middle of the costa and to below the middle of the termen
suffused with rosy carmine, through which run about nine
short longitudinal blackish streaks, crossed obliquely near
their base and again beyond their outer extremities by slender
steel-grey lines coming from a series of costal geminations,
alternately black and whitish ochreous; at the extreme apex
one of these whitish ochreous streaks, longer than the others,
descends to the termen, where it interrupts a narrow black
line which extends along the extreme margin from the apex
for two-thirds of its length in the direction of the tornus ; cilia
bright fawn-ochreous, tipped with greyish fuscous at the apex.
Exp. al. 21 mm. Hind wings brownish fuscous; cilia
whitish cinereous, a slender greyish fuscous line running
through them near their base. Abdomen brownish fuscous,
anal tuft subochreous. Legs [missing].
Type, S (70258) Mus. Wlsm.
Hab. JAPAN — Howpvo — Gifu, 1V.-Y. 1886 (Pryer),
Unique.
Potycurosis, Rag.
1022. Polychrosis porrectana, Z.
Eudemis porrectana, Stgr. & Wk. Cat. Lp. Eur. 251, No, 1022 (1871).
Hab. Kvrore. Asiatic TuRKEY—Hazzz—Shar Devesy
(Native Coll. 1893).
Nortocexia, Hb.
1004. Notocelia Uddmanniana, L.
Aspis Uddmanniana, Stgr. & Wk. Cat. Lp, Eur. 250, No. 1004 (1871).
Hab. Kuropr. Asiatic TuRKEY—Hazez—Shar Devesy
(Native Coll. 1893); Pazzsrrve (Tristram, 1883).
Lord Walsingham on Asiatic Tortricidae. 335
PELATEA, Gn.
1030 (1). Pelatea bicolor, sp. n.
Antenne fuscous. Palpi moderate, porrect, median joint
closely scaled, more thickly above than below, terminal joint
short, exposed ; fuscous, terminal joint tipped with ochreous.
Head and thorax fuscous. Fore wings with a basal patch
occupying one-fourth, its outer margin straight, blackish
fuscous ; a broad yellow ochreous band across the middle, its
inner and outer edges both straight, whitish on its inner side
and with a narrow whitish line along its outer margin, with
three minute fuscous costal dots and two or three also on the
dorsum ; the apical third of the wing dark brownish fuscous,
sprinkled with shining greyish fuscous, especially about its
margins; cilia shining greyish fuscous, a slender pale line
along their base. Hap. al. 125 mm. Hind wings greyish
brown; cilia slightly paler. Abdomen greyish brown. Legs
cinereous.
Type, 8 (70167, Japan); 9 (70168) Mus. Wlsm.
Hab, JAPAN (Pryer, 1886). Assam— Kohima (Naga
Hills), 4700 feet, VI. 1889 (Doherty). Six specimens.
HYSTRICHOSCELUS, gen. nov.
(vorpeE=a porcupine; oKedAos=a lee.)
Type, ¢ 2, Hystrichoscelus spathanum, Wlsm.
Antenne (@ ) slightly ciliate. Palpi short, recurved, ap-
pressed to the face; median joint somewhat coarsely scaled
beneath, terminal joint short, porrect, exposed. Head rough
above. Thorax with a moderately erect tuft posteriorly.
Fore wings of moderate width, costa slightly convex, ¢ with
a weak costal fold; termen slightly oblique, scarcely im-
pressed, tornus rounded. Neuratzon: 12 veins, all separate ;
7 to termen; 3 somewhat recurved upwards. Hind wings
broader than the fore wings; termen and dorsum evenly
rounded. Neuration: 8 veins; 3 and 4 stalked; 5 mode-
rately straight, not closely approximate to 4 at base; 6 and
7 separate, but closely approximate at base, sometimes almost
coincident. Abdomen (@) with a rather long anal tuft.
Legs: 8, hind tibiz clothed with long hairs above, stretching
to the middle of the tarsal joints, but not so densely packed
as in Phecasiophora, Grt.: ¢, hind tibie slightly clothed
above.
This genus is apparently allied to Heldctophanes, Meyr.,
from which it differs in emitting vein 7 to the termen and
also probably in the more extended clothing of the hind
tibiee.
336 Lord Walsingham on Asiatic Tortricide.
1030 (2). Hystrichoscelus spathanum, sp. n.
Antenne tawny grey. Palpiand headdark tawny. Thorax
grey, mottled with tawny brown. Fore wings leaden grey,
mixed with some whitish scales and transversely streaked
and banded with umber-brown to two-thirds their length,
the apical third white, slightly shaded with grey, and having
numerous short black lines running longitudinally through it ;
the brown transverse streaking is somewhat plentiful on the
basal third, at the outer edge of which a thicker band, much
sinuated outwardly, forms the oblique margin of a basal
patch in which the grey ground-colour of the wing is much
mixed; scarcely beyond the middle is a broader band of
brown, much widened beneath the costa, but again narrowed
towards the dorsum, this is bounded on each side by leaden
grey; beyond it there are five pairs of geminated oblique
white streaklets along the costa, the space about their ends
being tinged with chestnut-brown, through which one or two
leaden grey lines are visible; a minute black dot lies within
the chestnut-brown apex; the white outer third of the wing
possesses a row of short black dashes towards its inner edge,
and a curved row of similar dashes above it reaching to the
middle of the termen, and some shining silvery grey scales,
set in a sprinkling of grey and brown scaling, lie on the
white space above the tornus ; cilia white at the tornus, with
a slender greyish line running through them above to the
apex and a dark line along their base on the upper half of
the termen. Hap. al. g 15, 2? 17-18 mm. Hind wings:
3 whitish, shaded with greyish brown, especially towards
the apex; cilia white: 9 greyish brown; cilia pale cine-
reous, with a greyish brown shade running through them
near the base; the flexal cilia contain a number of Jong
spatulate dentate scales, these arise from the margin, not
from the wing-surface, they are more strongly developed in
the female than in the male. Abdomen brownish grey.
Legs: & whitish, the long hairs above the hind tibize snow-
white, hind tarsi spotted with greyish fuscous: 2 whitish
cinereous; hind tibiz slightly clothed above, hind tarsi
barred with brownish grey.
Type, & (70154); ¢@ (70156) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886). Four specimens.
Evucosma, Hb.
1041 (1). Eucosma conformana, Mn,
Grapholitha conformana, Mn. Verh. ZB, Ges. Wien, XXII. (1872)
Abh, 86-7 (1872)1; Stgr. Hor. Soc. Ent. Ross, XV. 259 (1879) 2,
Lord Walsingham on Asiatic Tortricidae. 337
Hab. Evrope—Daruarra—Ragusa’, Spalato’. Croarra
—Josefsthal’. Huweaasary — Ofen*. Asratic TURKEY —
Kuvupaveyprxrark — Brussa, V1.1; Pacesrrve (Tristram,
1883),
1049 (1). Hucosma medullana, Steger.
Grapholitha medullana, Stgr. Hor. Soc. Ent. Ross. XV. 254-5 (1879) 3,
Hab. Astatic Turkey — Kavpavenprerar — Brussa!;
Arpin — Smyrna’; Karamavra —'Taurus!; Hares — Shar
Devesy, 1 VII. 1890 (Native Coll.).
1057. EHucosma hepaticana, Tr.
Grapholitha (Pedisca) hepaticana, Stgr. & Wk. Cat. Lp. Eur. 253,
No. 1057 (1871).
Hab, Kurope. Astaric TurKEyY—Hazzez—Shar Devesy
(Native Coll. 1893).
1061. Hucosma graphana, Tr.
Grapholitha (Pedisca) graphana, Stgr. & Wk. Cat. Lp. Eur. 253,
No. 1061 (1871).
Hab. Kuropre. Astatic TurKEY—/Hazzes—Shar Devesy
(Native Coll. 1893).
1069 (1). Eucosma pica, sp. n.
Antenne greyish, faintly annulate, basal joint white.
Palpi white, with a black spot externally at the base of the
median joint. Head white, touched with black at the sides
behind the antenne. Thorax white, with a black band
across in front and some black scales behind the middle,
Fore wings: costa gently arched, ¢ with a narrow fold at the
base, apex slightly rounded, termen impressed, slightly
oblique; white, streaked and mottled with black on the costa
and dorsum and on the fold to two-thirds the wing-length; a
large black eostal patch about the middle is carried obliquely
downwards and outwards nearly to the fold, where it is bent
upwards towards the apex and somewhat narrowed, joining
an irregular black patch below the apex, which runs to the
middle of the termen ; beneath it before the tornus is a trian-
gular black patch containing some white dots on the dorsum,
and beyond its outer edge, which runs at right angles from
the margin, is a pale steel-grey ocelloid patch mixed with
white ; on the costa beyond the median patch are three black
spots, sending out black streaks to the subapical shade, slender
338 Lord Walsingham on As?atic Tortricidee.
black lines lying between them on the white ground-colour,
the apex is black ; cilia white below the middle of the termen,
grey above it, with a black line along their base reaching
only to the middle of the margin. zp. al. 17-20 mm.
Hind wings slightly impressed below the apex ; pale brown;
cilia pale cinereous, with a dark shade running through them
near their base. Abdomen pale brown. Legs whitish, shaded
and spotted with grey and black.
Type, 2? (60023); g (70075) Mus. Wlsm.
Hab. Japan (Pryer, 1886)—Howno— Yokohama (Manley,
1888). Six specimens.
1091. Hucosma tripunetana, Schiff.
' Grapholitha (Pedisca) tripunctana, Stgr. & Wk. Cat. Lp. Eur, 255,
No. 1091 (1871)'; Stgr. Hor. Soc. Ent. Ross. XV. 257 (1879) *.
Hab. EKvrore*. Asiatic TurKEY—Kuopavenprerar—
Brussa, [V.-V.?; Szvas—Amasia, VI.?; Awarozra*; HaeB
—Shar Devesy, 24 VI. 1890 (Native Coll.).
1095. Hucosma Pflugiana, Hw.
Grapholitha (Pedisca) Pflugiana, Stgr. & Wk. Cat. Lp. Eur. 255,
No. 1095 (1871) 1; Stgr. Hor. Soc. Ent. Ross. XV. 257 (1879) *.
Hab. Europe. Astatic ‘URKEY—KuavpavenpDre14r—
Brussa, V.°; | Agrmenra’. CuHinA—Ta-chien-lu, V.—VI.
(Leech, 1891).
1095 (1). Eucosma Pryerana, sp. n.
Antenne greyish fuscous. Pali short, scarcely pro-
jecting beyond the head; dark brownish cinereous. Head
dark brownish cinereous. Yhorax dark purplish fuscous.
Fore wings blackish brown, thickly sprinkled with shining
dark blue-grey streaks and blotches; a conspicuous white
patch on the middle of the dorsum reaching to the fold, its
apex sometimes overlapping the fold; three or four pairs of
whitish geminated streaks on the outer half of the costa;
four black spots on the ocelloid patch le between bands of
shining dark blue-grey ; the apical portion of the wing is
tinged with ferruginous, through which run blue-grey streaks
from the costal geminations; cilia greyish, tinged with ferru-
ginous at the apex. wp. al. 22-25 mm. Hind wings dark
brownish fuscous; cilia creamy whitish, touched with
brownish at the apex, a dark brownish fuscous line along
their base. Abdomen dark brownish fuscous. Legs greyish
brown, hind tarsal joints spotted with whitish ochreous.
Lord Walsingham on Asiatic Tortricide. 339
Type, (70069) Mus. Wlsm.
Hab. JarpaN—Honvo—Oiwake, VII. 1887 (Pryer). Six
specimens.
This species is nearly allied to Pfluytiana, Hw., but differs
in the more pure whitish cilia of the hind wings and in the
more clearly defined white dorsal patch. I have seen none
of the mottled varieties that occur in the European species.
1095 (3). Hucosma (?) macrorris, sp. n.
Antenne brownish cinereous. Papi rather long, median
joint with a long projecting tuft beneath, terminal joint con-
cealed; brownish cinereous, shaded with fuscous. Head
brownish cinereous. Thoraxdark umber-brown. Fore wings
broad, costa arched, termen slightly oblique, not impressed,
tornus rounded ; dark umber-brown mixed with ferruginous ;
acream-white upright dorsal patch before the middle is divided
by a single dark umber streak on its lower half; above and
before it, at about one-third from the base, is a smaller
creamy white costal spot, also divided through its middle by
dark umber seales; three or four pairs of very short creamy
white geminations occur on the outer half of the costa; near
the base beneath the fold, and also about the region of the
ocelloid patch, which contains some indistinct patches of
leaden grey, the ferruginous scaling is more conspicuous than
on the other parts of the wing, it extends also upwards along
the termen ; cilia creamy white on their upper half, touched
with fuscous at the apex, leaden grey on their lower half, with
a fuscous line along their base. EHzp. al. 25mm. Hind
wings dark brown; cilia shining greyish, a brown line
running through them near their base. Abdomen dark brown.
Legs brownish, hind tarsal joints spotted with fuscous and
whitish ochreous.
Type, 2 (70067) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886). Unique.
This species has much the appearance of Hucosma ex-
pressana, Chr. (=contrasignata, Chr., 2), but differs in its
longer palpiand more rounded costa. It is probably correctly
referred to the genus Hucosma, but the male is at present
unknown.
1095 (4). Hucosma contrariana, Chr.
Grapholitha contrariana, Chr. Bull. Soc. Imp. Nat. Mose. LVI. 424-6,
No, 126 (1882)?: sep. 187-8 (1882)?. Grapholitha (Pedisca) con-
trarvana, Snell. Tijd. v. Ent. XXVI, 218-9, Pl. XIII. 3, 34
(1883) *,
340 Lord Walsingham on Asvatic Tortricide.
Hab. E. Srserta—Irkutsk, 30 VI.-30 VII’; Pompe-
jefka*; Wladiwostok, VII; Amur, 21 VII CxHma—
Chang Yang, 4000-6000 feet (Pratt, 1886).
1106 (1). Hucosma inconspicua, sp. n.
Antenne greyish fuscous. Palpi with a rather strong pro-
jecting tuft of scales beneath the median joint, terminal joint
small, exposed; greyish fuscous. Head greyish fuscous.
Thorax dark brown. Fore wings moderately broad, costa
slightly convex, ¢ with a narrow fold at the base reaching
to beyond the basal third, termen slightly oblique ; greyish
fuscous, mixed with dark umber-brown and ferruginous, with
leaden grey lines about the apical third, which is especially
tinged with ferruginous ; a very faintly indicated triangular
ereyish patch on the middle of the dorsum contains a dark
streak at its middle; a curved steel-grey band, from a little
beyond the middle of the costa, runs obliquely outwards in
the form of two parallel streaks, tipped with whitish on the
extreme costa and joining at their lower end the steel-grey
band which precedes the ocelloid spot; beyond this are three
pairs of whitish geminated costal streaks, throwing out steel-
grey lines to the termen, and beneath these the outer edge of
the ocelloid spot, which contains three black transverse
streaks set in ferruginous, is also steel-grey, the termen and
apex are also ferruginous ; cilia dark bluish grey. zp. al,
19-22 mm. Hind wings dark brown; cilia pale cinereous,
with a darker shade running through them near their base.
Abdomen dark brown. Legs greyish fuscous, hind tarsal
joints spotted with whitish ochreous.
Type, 3 (60122); ¢ (60125) Mus. Wlsm.
Hab. Japan (Pryer, 1886)—Honpo—Tsuruga, VII. 1886
(Leech). Eleven specimens.
1107. Eucosma fanella, L.
Grapholitha (Pedisca) feenella, Ster. & Wk. Cat. Lp. Eur, 255, No. 1107
(1874).
Hab. Europe’. ArMmenrIA!. CoreEA—Gensan, VI. 1836
(Leech), VIL—-IX. 1887 (Ito). Cui1na—Chang Yang, 4000-
6000 feet (Pratt, 1886). Japan (Pryer, 1886) — Kivsrv
(Leech, 1890)—Satsuma, V. 1886 (Leech). InvIA—Punsaz
—Dharmsala (Hocking).
The series is extremely variable.
Bibliographical Notices. B41
1110 (1). Eucosma fessana, Mn.
Grapholitha fessana, Mn. Verh. ZB. Ges. Wien, XXIII. (1875) Abh.
573 (1873) !; Stgr. Hor. Soc. Ent. Ross. XV. 257-8 (1879) *.
Hab. Asiatic TurKEY—Kuaramanra—Kiilek* (=Giilek ’) ;
S1vas—Jenikeui-Hochebene, 15 VI.*; Maidan 11, V.*; Az-
menr4a—Manglis*; Hazres—Shar Devesy, 25 VII. 1890
(Native Coll.).
Larva in stems of Salvia candelabrum ’.
A small specimen from Shar Devesy with rather darker
hind wings cannot be described as distinct.
[To be continued. |
BIBLIOGRAPHICAL NOTICES.
A Treatise on Zoology. Edited by E. Ray Lanxustrr, M.A., LL.D.,
F.R.S.—Part IIL. The Echinoderma. By F. A. Barner, M.A.,
assisted by J. W. Grecory, D.Sc., and E, 8. Goopricn, M.A.
London: Adam & Charles Black, 1900.
Tue present volume, on the Echinoderma, is the first published,
but third in order of a Comprehensive Treatise on Zoology, which
has been for some time past in preparation under the guidance of
Prof. E. Ray Lankester.
Rather more than half of this volume has been written by Mr. F.
A. Bather. Mr. E. 8. Goodrich is responsible for the section on the
Holothuroidea, and Dr. J. W. Gregory for that on the Stelleroidea
and Echinoidea.
This is essentially a student’s book. Its aim is to be a syste-
matic rather than an anatomical work, hence facts that are of
purely anatomical interest find no place here. This is well, for
they have recently been dealt with in Prof. Lang’s excellent
compendium, and would only crowd out matter more germane to
the purpose. Already this book has been much compressed and
any further condensation would seriously threaten the usefulness
of the whole.
This work is reared upon the foundation of phylogeny and
ontogeny ; and if it be objected that this is a somewhat insecure
foundation, it must at least be admitted it is the ideal aim of every
post-Darwinian taxonomist. It may be claimed for this book that
it is unique, in that fossil and living forms are regarded as common
material for the building thereof. The former are not regarded as
merely decorative elements. So well has the piecing together of
these fragments been done, that their true relationships to the living
forms can be grasped with something more like certitude than ever
342 Bibliographical Notices.
before. Asa result, we have for the first time a real insight into
the inter-relationships of this most difficult group.
The calycinal system is no longer the governing factor in our
reckoning of the morphological level of the Echinoderma, as in the
systems of Lovén, Carpenter, and Sladen. The plates taking part
in the apical system of the Echinoidea and Stelleroidea cannot,
Mr. Bather shows, now be regarded as merely homogenetic with
those of the Crinoidea. The evidence of the fossils is fatal to this
conclusion, inasmuch as the Eleutherozoa “if they arose from
stalked forms at all, indubitably did so ages before the calycinal
system had been evolved.”
The most primitive Echinoderms which we know at present
appear to be the Cystidea Amphoridea. From this stock probably
branched the Cystidea Rhombifera, Cystidea Diploporita, Blastoidea,
and Crinoidea on the one hand, and the Edrioasteroidea and Eleu-
therozoa on the other, these last being derivable possibly from the
Edrioasteroidea.
The account of the larval forms, which is absolutely indispensable,
is lucidly, if briefly, sketched. Herein the complex coilings of the gut,
the changes in the development of the ccelom, following the changes
from a free-moving to a fixed habit, and the gradual evolution into
the characteristic radial symmetry are made as clear as such a
difficult matter can possibly be made. Most of the figures
illustrating this section are new. That showing the change from
the Pentactea to the Eleutherozoic Stelleroid type is very in-
structive.
Mr. Bather supports the view held by many that the simplicity
of structure of the Synaptide is a secondary feature, and that
therefore this form cannot be regarded as the simplest and most
ancestral of the Echinoderms. The class he regards as a probable
early offshoot from the Edrioasteroidea.
Mr. E. §. Goodrich’s summary of the Holothuroidea is admirable
and well illustrated. We venture to think he missed an opportunity
in not directing attention to the fact that Ludwig’s classification
does not agree with his phylogenetic tree ; and that whilst the latter
may be taken as a more or less probable expression of the relation-
ships of the forms included, one to another, the former is artificial,
and savours rather too much of a “ Key.”
Dr. Gregory has certainly sustained the high standard which
characterizes this work. He insists on the close relationship of the
Stelleroidea and Ophiuroidea, wherein most will agree with him,
It is ineorrect, however, to state that Ophioteresis agrees with the
Stelleroidea in having an ambulacral furrow. This is an important.
point, for Dr. Gregory uses it as an instance showing the unreli-
ability of this character asa taxonomic factor for the division of the
Ophiuroidea from the Stelleroidea. Again, the supposed pore-plates
of Bothriocidaris are really tubercles. But these are to be regarded
as slips which will creep into every work in spite of the most zealous
precautions.
Prof. Lankester’s choice of authors for this work has been in
Bibliographical Notices. 343
every sense justified, and both Editor and Authors have placed
zoological students under a great obligation by bringing within easy
reach, and with marvellous completeness, all the essential facts
concerning a group which has always ranked as one of the most
difficult of comprehension. This book, we may safely say, as yet
knows no rival.
It is beautifully and profusely illustrated and remarkably free
from misprints. The only one which we have detected, so far, is on
page 9, where constructed appears for constricted.
W. P. Pyorart.
A Monograph of Christmas Island. Physical Features and Geology
by Cuartes W. Anprews, B.A.; with Descriptions of the Fauna
and Flora by numerous Contributors. Printed for the Trustees
of the British Museum. London, 1900.
Tus is a book of remarkable interest, and one of more than ordinary
scientific value. It is an embodiment of the results of a ten-months’
stay on Christmas Island by Mr. Andrews during 1897-98 ; and the
Trustees of the British Museum, in publishing these results, have
conferred a great and lasting benefit upon students of natural science.
Our thanks, however, are not alone due to the Trustees. “It
seemed highly desirable,” writes Sir John Murray in an Introductory
Preface, “ that this interesting island—which was evidently an up-
raised coral atoll—should be carefully examined and described by a
competent naturalist and geologist before being opened up by Euro-
peans for agricultural and commercial purposes. Accordingly it was
arranged with the Trustees of the British Museum that Mr. C. W.
Andrews, B.Sc., F.G.8., of the Geological Department, should be
granted leave to carry out this exploration. I undertook to pay all
the expenses and to present a complete set of all specimens procured
to the National Collection.”
The physical features and geology have been written by Mr. An-
drews. The zoology has been worked out by various specialists,
most of whom are members of the Museum staff. Their work has
been well done. Comparisons are odious, so we refrain from
comment in this direction. Field-notes by Mr. Andrews are often
appended to the descriptions of species, and some of these notes are
of extreme interest. Perhaps one of the most vivid of these descrip-
tions is that of the frigate-bird. “ About the beginning of January,”
he writes, “ the adult males begin to acquire the remarkable pouch
of scarlet skin beneath their throat. This they can inflate till it is
nearly as large as the rest of the body, and a dozen or more of these
birds sitting in a tree with outstretched drooping wings and this
great scarlet bladder under their heads are a most remarkable sight.
When a hen bird approaches the tree the males utter a peculiar cry,
a sort of ‘ wow-wow-wow-wow, and clatter their beaks like casta-
nets, at the same time shaking the wings. When they take to flight
j44 Bibliographical Notices.
the air is allowed to escape from the pouch, but occasionally they
might be seen flying wfth it partly inflated.”
There is a point concerning this pouch of very great interest. It
is a secondary sexual character, developed during the breeding-
season, but so far we have no information concerning its nature or
the mechanism by which it is inflated and deflated. From an ex-
amination of spirit-specimens we believe that it will be found to be
nothing more than a widening of the cesophagus, filled through the
anterior nares, and held inflated by occlusion of the esophagus by
means of a sphincter at the back of the mouth.
One of the commonest mammals of the island is Mus Maclearii.
“Tt occurred in swarms. During the day nothing is to be seen of
it, but soon after sunset numbers may be seen running about in all
directions, and the whole forest is filled with its peculiar querulous
squeaking and the noise of frequent fights. ..... As may be
imagined, they are a great nuisance, entering the tents or shelters,
running over the sleepers, and upsetting everything in their search
for food. They seem to eat anything, and destroy any boots or skins
incautiously left within their reach.”
“ One of the chief objects of the expedition to Christmas Island
was to find out whether its structure would throw any light upon
the vexed question of the nature of the foundation of atolls.”
This exceedingly difficult question is most ably and lucidly handled
in the concluding part of this volume.
The occurrence in an oceanic island of thick beds of lower and
middle Tertiary limestone is a point of extreme interest and import-
ance, and the demonstration of this fact forms one of the many
important results obtained by Mr. Andrews. ‘“ Perhaps the most
remarkable of the rocks of Christmas Island are the thick deposits
of nearly pure phosphate of lime which cap several of the higher
hills. This substance is probably derived from ancient (? Pliocene)
guano-beds formed on the low islets which existed before the first
elevation of the island, and is the insoluble residue of beds of lime-
stone altered by this guano.”
Space forbids that we should extend this notice further; enough,
it is hoped, has been said to show haw admirably this work has been
carried out and how fruitful have been its results.
We entirely agree with Sir John Murray, who, in his Introductory
Note, says, “It will, I think, be admitted that in the present
elaborate report we have the best account of a true oceanic island
that has ever been published.”
The illustrations, which are numerous, are, on the whole, very
good. The text-figures, however, and one or two of the plates are
about as badly reproduced as could well be. W, P. Presser
THE ANNALS
AWD
MAGAZINE OF NATURAL HISTORY.
(SEVENTH SERIES. ]
No. 34. OCTOBER 1900.
XLIII.—Deseription of Sponges from Funafuti.
By R. Kirkpatrick, Assistant, Natural History Museum.
{Plates XIII-XV.]
THE sponges described in this paper were obtained by the second
Boring Expedition to Funafuti, and were entrusted to me by
Prof. Judd for description. ‘The specimens, which are mostly
very small, incrust or are attached to fragments of corals and
corallines dredged up from depths of from 30 to 145 fathoms.
The fragments were obtained only with great difficulty, since
they had to be detached from the bottom by means of chisels.
The collection includes the representatives of twenty-one
species, of which eleven are new.
In 1897 Mr. Whitelegge described (30, p. 323) from the
same locality a small collection obtained from the surface of
the reefs. Of the sixteen species recorded by him only one
occurs in the present collection.
Altogether thirty-six species have been recorded from this
locality. The special features of interest are the occurrence
(1) of Astrosclera Willeyana, Lister, which represents a new
family, or, possibly, order of calcareous sponges, (2) of a
recent species of the Lithonine genus Plectroninia, hitherto
found only in the Eocene, and (3) of a new genus of Clionide.
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 23
346 Mr. R. Kirkpatrick on
The following is a list of the species :—
Subclass CALCAREA. Fam. Spirastrellide.
1. Astrosclera Willeyana, Lister. 12. Latrunculia clavigera, sp. n.
2. Plectroninia Hindei, sp. 0.
8. Clathrina depressa (Dendy).
Suborder HALICHONDRINA.
Subclass DEMOSPONGIDA. Fam. Peoeciloscleride.
Order CARNOSA. 13. Agelas gracilis, Whitelegge.
Suborder OLIGOSILICINA. 14. Tedania levis, sp. 0.
4, Chondrilla mixta, Schulze.
Fam. Haploscleride.
Suborder MIcROSCLERO-
15. Chondropsis ceratosus, sp. N.
Misa 16. Pachychalina fibrosa, Ridley &
5. Curticium candelabrum, O. Dendy.
Schmidt.
6. Placinolopha spinosa, sp. 0.
7. Placinastrella clathrata, sp. n. Order MONOCERATINA
Order TETRACTINELLIDA. 17. Luffariella variabilis (Polé-
jaeff).
geometrica, sp. 0.
19, Psammopemma purpureum
Order MONAXONIDA. (Carter).
20. Stelospongus cavernosus (Pallas),
var. pyriformis, Lendenfeld.
Section CLAVULIDA, 21. Polyfibrospongia Sweeti, sp. n.
Fam. Clionidz.
8. Erylus monticularis, sp. 0. 18.
Suborder HADROMERINA.
9. Cliona mucronata, Sollas.
10. Schmidti (Ridley).
11, Dyscliona Davidi, gen. et sp. 0.
Astrosclera Willeyana, Lister.
1900. Astroselera Willeyana, Lister (16, p. 459, fig. A, 1-5).
The single large specimen of this species from Funafuti
was sent to Mr. J. J. Lister, of Cambridge, who was at the
time engaged in working out the structure of four small
specimens of the same species from Lifu. Mr. Lister’s
description of the external features of the Funafuti specimen
is as follows :—
«The specimen from Funafuti (fig. A, 1-5) has grown
attached by a short stalk about 6 millim. wide at the base,
which expands into a broad nearly citcular disc, convex
above, and resembling the pileus of a mushroom in shape.
The diameter of the disc is about 20 millim., and the distance
Sponges from Funafuti. 347
from the broken end of the stalk to the centre of the upper
surface of the disc is 16 mm.
“A smooth imperforate cortical layer covers the outer
surface of the stalk and under surface of the disc... . Tha
upper surface of the disc is in part perforatel by pores.
“There appears to have been an interruption of the growth
of the specimen after the formation of the disc, and the later
expansion has taken place not uniformly, but only from parts
of its upper surface. At these parts rounded bosses of diffe-
rent sizes have been formed, and they, like the original disc,
are perforated by pores on their convex upper surfaces and
covered on the sides by an imperforate cortical layer... . the
greater mass of the new growth appears to have been formed
by the fusion of three originally distinct bosses.
“The upper surface, whether of the original disc or of the
later formed bosses, is perforated by closely-set pores. These
are sometimes isolated, but often they open into curving and
branching grooves, recalling those of the coral Meandrina.
“At seven places on the surface of the later growth, and
at one on the original growth, the lines of pores are seen to
be disposed in a radiating manner about so many centres.
The pores along these lines are large, being at least twice the
diameter of those distributed elsewhere.
“The fully formed skeleton is built up of a solid mass of
polyhedral elements (40 to 150 w in diameter), whose surfaces
are united closely together to the complete exclusion of the
soft parts.”
For a full account of the structure of the skeleton and soft
parts of this sponge reference must be made to Mr. Lister’s
memoir (16). ‘The skeletal structure of Astrosclera differs so
entirely trom that of other sponges that it will probably have
to be placed in a third order of Calcarea, the other two being
Lithonina and Dialytina; but Mr. Lister would defer the
establishment of such a division until more material becomes
available for investigation.
PLECTRONINIA, Hinde (12, p. 51).
Plectroninia Hindei, sp. n. (Pl. XIII. fig. 1, a-r.)
The specimen, which forms a minute oval crust on a coral-
line, is 43 millim. in area and 1 millim. in thickness at the
centre, but thinning away to the edges. ‘The surface appears
vitreous, with a faint yellow tinge, the interior being erystal-
line vitreous. The crust, which was easily flaked off from
the cc line, showed a granular crystalline basal layer; on
23%
348 Mr. R. Kirkpatrick on
vertical section a second granular “ basal” layer, parallel
with the first, occurs in the middle of the section.
Under magnification, the polygonal openings (105 @ in
diameter) of the excurrent canals are seen occupying the
whole upper surface.
The thin edges of the specimen allow of no room for a
dermal layer, but in one small recess there was a little nest of
loose oxeas and styles, possibly the constituents of such a
layer.
The skeleton is mainly built up of stout quadriradiates,
usually smooth, with fused facial rays and with the sharp
apical ray more or less free and pointing almost vertically to
the surface; occasionally the surface of the rays is slightly
spined or tuberculated.
The basal layers are composed of densely packed but
separate small quadriradiates, varying considerably in shape
and size, with rays frequently spined and terminating in flat
expansions.
Spicules—Skeletal network : length of beams formed by
facial rays 70-100 mw, and 40 yw in thickness; apical rays
85-200 w in length by 15-45 w in thickness.
The rays of the smaller basal quadriradiates vary from
18 to 35 w in length.
Triradiates: (1) Sagittal, rays sharp-pointed, basal ray
120x7 yp, laterals 60x 7 p.
(2) Irregular, with slender pointed unequal rays, resting
on the apices (tripod type).
(3) Tuning-forks: (a) with prongs curved away from the
plane of the handle, with equal prongs each ending in trifid
oints.
; (b) Prongs and handle in one plane, prongs of unequal
length and spined; there are several variations in the shape
of form (0).
Lance-shaped subtylotes and styles, finely spined with
disk- or crook-shaped head, 200-500 x 6 yw, head 5 w, neck 3 ph.
Finely spined oxeotes, 200-400 x 7 p.
Plectroninia Hindei constitutes the second recent Lithonine
sponge that has been discovered, the first being Petrostroma
Schulzet, Léderlein, from Japan (10, p. 15, pls. ii.—vi.).
The new species is near to Plectroninia Halli, Hinde,
from Eocene beds at Geelong and Flinders. So far as can
be made out from crushed fragments and a broken surface of
the minute specimen, the skeletal framework is fairly regu-
larly reticulate, without presenting any radiating main beams,
and the facial rays of the quadriradiates are always expanded,
and not tapering. The chief specific differences from
Sponges from Funafutt. 349
P. Halli lie in the smaller size of the beams and meshes of
the network and in the characters of the loose spicules. The
type of the genus is turbinate in form, but it cannot be said
what would be the ultimate shape of the probably young
specimen of P. Hindei.
Locality. Beacon Island, Funafuti, 50 fath.
Clathrina depressa (Dendy).
1891. Leucosolenia depressa, Dendy (8, p. 65, pl. iii. figs. 4, 4a, pl. viii.
fig. 8, pl. xi. fig, 4).
The specimen forms a small whitish patch about 6 millim.
in area on the surface of Luffariella variabilis. ‘The dermal
triradiates are slightly larger than in specimens from Port
Phillip, being 240 x25 yw, as against 200 x28 w from the
latter locality ; the quadriradiates are few in number and
with the apical ray almost aborted.
Locality. Funamara, Falefatu, 25-45 fath.
Distribution. Port Phillip, Victoria; Funafuti Atoll.
Chondrilla mixta, F. E. Schulze.
1877. Chondrilla mixta, F. FE. Schulze (23, p. 116).
The one specimen forms a small bluish-black incrustation
onan Agaricia. The surface of the sponge is finely granular
to the naked eye. The spherasters are 25-30 w in diameter,
with about 20 pyramidal prickles, and the oxyasters 24-28 pw
in diameter, with a centrum 7 yw in diameter.
The spherasters just below the surface are united into little
heaps, which give rise to the granular appearance of the
surface ; the oxyasters are only very rarely present in the
ectosome.
Locality. Funafuti Islet.
Distribution. Java; Amirante Is.; Red Sea; Funafuti
Atoll.
Corticum candelabrum, O. Schmidt.
1862. Corticiwm candelabrum, O. Schmidt (21, p. 42, pl. iii. fig. 25),
1881. Corticium candelabrum, F. E. Schulze (26, p. 410).
The specimen forms a small greenish-black disk, 3 millim.
in area by 1 millim. in thickness, attached to a piece of
coralline.
Locality. Fuafatu, 50-70 fath.
Distribution. Adriatic; Zebu; Funafuti Atoll.
350 Mr. R. Kirkpatrick on
Placinolopha spinosa, sp.n. (Pl. XIII. fig. 2, a—m.)
The specimen forms a whitish-brown crust about 10 millim.
in area and 1-2 millim. thick, creeping over a nodule of
coralline. Several oval oscules, ‘5 x ‘25 millim. in diameter,
with membranous edges level with the surface, are present.
The chcanosome forms more or less vertical folds beneath
the ectosome. The skeleton is composed of micro-calthrops
of various sizes densely scattered in the ectosome and walls
of the choanosomal folds, and of lophodiactines arranged
tangentially in the ectosome and in the walls of the folds.
_ Sptcules.— Microcalthrops in graduated sizes, from very
slender forms with rays 24x 2 yw, smooth or slightly spined,
with simple or bifid ends, up to stout forms with rays 42 x 9 p,
much spined and di- or trichotomously branched at the ends.
Lophodiactines 240 x12 yw, with simple or branched spines
and once or twice branched at the ends.
The lophodiactines of P. spinosa resemble in form a spicule
figured by Sollas (26, pl. xxxv. fig. 24), who found it asso-
ciated with spicules of the Lithistid Corallistes Thomasi from
the Ki Islands. Sollas, who was doubtful whether to regard
the spicule as Tetractinellid or Monaxonid, assigned it to a
new genus and species— Orthorhachis problematica. It seems
very probable that Sollas’s species comes under Placino-
lopha. The spicule figured by him is 450x40 p, nearly
twice the size of the largest lophodiactine of P. spinosa.
Including that of Sollas, there are three species belonging
to this genus :—
P. Bedoti, Topsent.—Philippines (28, p. 429).
P. spinosa, sp. n.—Funafuti.
P. problematica, Sollas.—Ki Islands.
Locality. Ei. end of Fuafatu, Funafuti Atoll, 50-70 fath.
Placinasirella clathrata, sp.n. (Pl. XIII. fig. 3, a-o.)
The specimen, which forms a small, rounded, soft nodule
8x 5x5 millim. in size and of dirty white colour in spirit,
appears to have been cut off from a coral.
The surface is smooth, but when highly magnified shows
an extremely fine pile formed by the points of a palisade of
cortical diactines. An oscule in the form of an irregular
fissure about 1 millim. in length, and level with the surface,
occurs on the summit of the specimen. The pores occupy
irregular cribriform areas with well-defined margins; the
sieve-meshes are 70 mw in diameter and the pores about 30 yu
(about two or three to a mesh). The pores lead by short
Sponges from Funafute. 351
canaliculi into a system of subdermal spaces, which open into
incurrent channels and spaces in the interstices of a close
network of tubes, the lumen of the tubes constituting the
excurrent canal system ; this arrangement recalls that found
in the Ascon genus Clathrina.
Skeleton.—The ectosome is supported by a palisade of
slender diactines, but in the meshes of the pore-areas the
diactines are tangential. The walls of the choanosomal tubes
are supported by tangentially arranged diactines, triods, and
microcalthrops of various sizes.
Spicules.—Diactines attaining 140 x 5p, straight or slightly
curved, with aslight kink or irregular thickening in the centre,
and often abruptly bent near each end. HKctosomal diactines
36 x 1:5 w, with a slight kink in the centre.
Triods regular or irregular, with two rays (together
120 x 4 w) in a line and a shorter third ray (36 w) forming an
angle.
Calthrops, larger forms with rays 36 x5 yw, smaller with
rays 18X3 wm.
The only other species of this genus Placinastrella c piosa,
Schulze (24, p. 483, pl. xxi. figs. 17-21), is found at Naples.
A cortical palisade of diactines is present in both; but the
large definitely orientated calthrops occurring in P, copiosa
are wanting in P. clathrata.
The tubular reticulum appears to be merely an elaboration
of the system of simple folds found in Placina simplex, and
the proper position for Placinastrella is probably in the
family Placinide, where Schulze placed it (24, p. 449), rather
than in the Pachastrellide, to which the genus was assigned
by Sollas (26, p. 103).
Locality. Funafuti Islet.
Erylus monticularis, sp.n. (Pl. XIV. fig. 3, a-A.)
The sponge forms an extremely thin pale brown lamella
incrusting an Agaricia. Several minute conical oscules,
‘07 millim. in height and ‘175 millim. broad at the base, are
scattered over the flat surface, the walls of the cones being
supported by a sloping palisade of small oxea 80 x 3 yu.
Skeleton.—Beneath the surface-layer of disciform sterrasters
is a layer of scattered orthotrianes, with reduced rhabdomes
directed vertically downwards.
a et ee NEBR oxea (few) slightly curved, 210 x
[bs
Orthotriznes: rays of cladome each 186x3 yp, slender,
straight, sharp-pointed, rhabdome 6 pw.
Microscleres: sterrasters oval, with diameters 150 x 114 y,
352 Mr, R. Kirkpatrick on
both surfaces granulose, with radiating strie, edges finely
serrated,
Microxea (medium) 80x3 yw, slightly curved. Very
slender microxea 40 x 1 p.
Small chiasters 10 w in diameter, with small centrum and
about 12 slightly tylote rays.
Oxyasters 18-30 mw in diameter, with about 6 rays.
The new species is near H. placenta, Thiele (27, p. 5, pl. 1.
fig. 1, pl. vi. figs. 1, a-h), but differs from it chiefly in the
proportions of the oxea and orthotrienes. In Thiele’s species
the oxea attain a size of 800 w, and the microxea are centro-
tylote. Further, in the new species there are no orthotrieenes
with long rhabdomes 500 yw in length, but only aborted
forms.
Locality. Funafuti Islet.
Cliona Schmidti (Ridley).
1870. Vioa Johnstont, var., O. Schmidt (22, p. 5, pl. vi. fig. 18).
1881, Vioa Schmidti, Ridley (18, p. 130).
1884. Vioa Schmidti, Ridley (19, p. 622).
1898, Vioa Schmidtu, Lendenfeld (14, p. 72, pl. iii. fig. 31, pl. vi.
fig. 58, pl. vii. fig. 74, pl. x. figs. 185-189).
The specimens of this species ramify in fragments of
Echinopora along with Dyscliona Davidi, and in a lamina of
Agaricia along with Cliona mucronata, Sollas. Colour
purple.
One of the two kinds of spinispirulas is 54 x 1°5 p, slender,
with eight or more curves and with separate short blunt
spines, the other kind being much stouter, 48x 6 mw, with
about six bends and with sharp pyramidal spines.
The tylostyles are 8360 x 4 w, straight or slightly curved,
with oval or spherical heads about 7 mw in diameter, but
showing a certain amount of variation in shape and size.
Locality. Funamara, 25-45 fath., and Funafuti Islet.
Mistribution, Amirante Is.; Funafuti Atoll; Adriatic.
DYSCLIONA, gen. nov.
Clionide with diactinal (usually strongylote) megascleres
and spined diactinal microscleres.
Dyscliona Davidi, sp.n. (Pl. XIV. fig. 1, a-g.)
Boring sponges of orange-yellow colour. Oscular papille
(and poral papille ?) projecting slightly above the surface,
cylindrical, patent, and about 1 millim. in diameter. Mem-
branous poral areas (?) irregular in form and level with the
Sponges from Funafuti. 353
general surface. Lobes (mainly following the shape of the
meshes of the madrepore in the present specimens) usually
elongated and quadrangular, attaining a size of 35x 1°5
millim., but varying in shape and size ; lobes connected with
each other by short tubular channels varying in diameter ;
the interior of the lobes occupied by a labyrinth of thin-
walled channels and spaces. Flagellated chambers oval,
eurypylous, 30 x 24 mw.
Skeleton formed of scattered strongyles occasionally forming
an irregular reticulum, and mostly lying tangentially in the
walls of the lobes.
Spicules.—Megascleres: strongyles varying from 126 to
246 w in length and from 3 to 5°5 mw in thickness, curved at
the centre, smooth, rarely slightly roughened at the ends.
Microscleres: microstrongyles (rather rare) 90x3 yp,
slightly curved, with truncate ends, and with spines arranged
in a regular closely-whorled spiral.
Surface of the “ galleries,” and especially of the connecting
channels, finely shagreened, but frequently smooth,
The specimens consist of three fragments of the coral
Echinopora excavated by the sponge. ‘Two of the pieces are
dried, the sponge being of a dark brown colour; the third
piece is in formol, the sponge here being orange-yellow when
the coral is freshly fractured, but gradually becoming brown
on exposure to light. The fragments are also excavated by
the purple boring-sponge Cliona Schmidt, Ridley. The
new species is nearly related to the Cliona purpurea of
Hancock (11, p. 343, pl. xii. fig. 6), the latter species now
coming under the new genus,
Topsent (28, pp. 576-7) was of opinion that Hancock’s
species was not a genuine boring-sponge, but possibly a
Desmacidine which had grown into excavations made by
some other organism. In answer to my request for informa-
tion about Hancock’s collection, Mr. R. Howse, Curator of
the Newcastle Museum, very kindly forwarded to me the
type specimen of Chona purpurea. In view of the close
affinity of this species to D, Davidt, a brief description of the
former is given here for comparison.
The type specimen consists of several small fragments of
Tridacna gigas, with holes for the oscular and poral papille
on the inner and outer surfaces. The broken edges show a
labyrinth of small oval cavities with finely shagreened surface
connected by tubes or foramina, and lined with a dull red
fluffy membrane. ‘The megascleres are amphi-subtylote and
measure 246 x 5°25 yw, the ends being 6 wu in breadth, slightly
spined, and sometimes with a terminal tuft of spines. ‘lhe
354 Mr. R. Kirkpatrick on
microscleres are strongylote or substylote, slightly curved,
finely and irregularly spined, and 90-144 w in length by
3-5 pw in breadth.
After a careful examination of Hancock’s specimen I |
consider Dyscliona purpurea to be a genuine boring-sponge.
To return to the Funafuti species: the Hchinopora has
been bored by Cliona Schmidti and by the filamentous alga
Achlya, and precautions were necessary to ensure that the
Dyscliona was obtained free from admixture with the Cliona.
Fragments of the coral with the cavities lined solely with the
yellow tissues of Dyscliona were separated, none of the tylote
and spinispirular spicules of C. Schmedti keing found. Hence
I consider that these excavations were formed by the
Dyscliona. Further, the existence of two closely allied
species, D. purpurea and D. Davidi, ramifying deep down in
the substance of shell and coral, tends to confirm the hypo-
thesis that we have here genuine boring-sponges.
If this supposition be correct, the true position of these
species would seem to be in the Clavulid family Clionide,
despite the fact that only diactinal megascleres are present in
them and that true spinispirulas do not occur. The un-
doubted Clionid sponges, Cliona nodosa and C. labyrinthica,
have only diactinal megascleres; and, further, the spirally-
spined microscleres of Dyscliona Davidi may be regarded as
modified spinispirulas, the modification having proceeded still
further in D. purpurea.
If D. Davidi and D. purpurea are not Clionids we have to
assume that the faculty of boring belongs to Halichondrine
as well as to certain Clavulid and Tetractinellid sponges.
To have placed these two species in the genus Cliona
would have been to unduly disturb what Topsent has termed
the “incomparable homogénéité ” of that genus; accordingly
they have been put into a new genus, Dyscliona. The new
species is named after Prof. Edgeworth David, the leader of
the Australian Boring Expedition.
Locality. Funamara and Funafuti Islet, 25-45 fath.
Latrunculia clavigera, sp.n. (Pl. XIV. fig. 2, a-e.)
Sponge forming a very thin greyish-white crust (on the
stem of a Gorgonia).
Skeleton formed of a surtace-layer of discasters, and, beneath
this, of scattered tylotes arranged horizontally ; scattered small
nail-shaped tylotes with points upwards and heads on basal
surface.
Spicules.—Megascleres: tylotes, average size 850x9 yw,
Sponges from Funafuti. 355
slightly curved; head faintly trilobed, 12 » in breadth (one
large tylote 1295 x 15 w was present on the microscopic slide).
Microscleres : discasters, 54 mw in length, straight or
slightly curved, shaft spinose, tylote; disks concave above,
convex below, with thick spinose rims; upper disk 19 p,
lower 26 yw in diameter.
Small nail-shaped tylotes, 120 x 6 yw, straight, ending in a
very sharp point, with double head 11 yu in breadth.
Carter gives a figure of a discaster (2, p. 358, pl. xxix.
fig. 20) obtained by him from the debris of the root-tuft of
Euplectella cucumer from the Comoro Islands, which closely
resembles the discaster of the new species.
Locality. W. of Tutanga, Funafuti, 86 fath.
Tedania levis, sp.n. (Pl. XIV. fig. 4, a-e.)
Sponge forming a pale slate-coloured crust about 10 millim.
in area and °5 millim. in thickness, on a nodule of coralline.
Surface smooth ; no oscules or pores visible; consistence
firm.
Skeleton.— Dermal: a felt-like network of tangentially
arranged, contorted, spined strongyles. Choanosomal: a
rather dense mass of contorted spined strongyles, occasionally
with a tendency to unite into bundles; also scattered smooth
styli (rather rare) and scattered smooth strongyles (rather
rare). Trichodragmata and rhaphides abundant, the former
being cylindrical and frayed at the ends.
Spicules.—Spined strongyles 250 to 300 w in length and
3 to. 12 w in breadth, the dermal spicules being the more
slender. Styles smooth, straight, or slightly curved, 245 to
1380 p in length and 4 to 12 w in breadth. Smooth stron-
gyles straight, averaging about 1085 by 12 p.
Rhaphides 150 to 200 » in length and about 1 w in thick-
ness ; trichodragmata 150 to 200 w in length and about 6 to
12 w in thickness.
The new species, which is only placed in Tedania pro-
visionally, should probably come under a new genus near
Tedania, but differing from it in having spined diactinal
megascleres in the choanosome. ‘The contorted megascleres
call to mind those of certain Axinellid sponges.
Locality. Fuafatu, 50-70 fms.
CuHonpDROPSIS, Carter (6, p. 122).
Chondropsis ceratosus, sp. un. (Pl. XV. fig. 3, a-c.)
Two specimens. Sponge incrusting. Colour reddish
brown; texture soft and elastic.
356 Mr. R. Kirkpatrick on
Oscules 3 millim. in diameter, level with surface, with thin
margins. Dermal membrane smooth, supported by a reticu-
lum of primary and secondary fibres.
Skeleton formed of a square-meshed reticulum of horny
fibres, which latter may be clear or filled with sand-grains.
Width of meshes about 245 w; width of fibres from 24 to
48 p.
Spicules.—Megascleres : oxea (rare and possibly foreign)
slender, straight, sharp-pointed, 120 x2 wp.
Microscleres: sigmata abundant, 30 to 50 pu in length.
Locality. W. of Tutanga, 30 fath.
The specimens, which are of irregular shape, are growing
on fragments of Sertatopora, on branches of which they form
a thin semitransparent coating, but attaining some thickness
between the branches.
The new species differs considerably from other species of
Chondropsis in its comparative freedom from sand and other
foreign bodies ; but I can find no other position for it than
in the genus Chondropsis, Carter, as emended by Dendy
(9, p. 250).
Pachychalina fibrosa, Ridley and Dendy.
1887. Pachychalina fibrosa, Ridley and Dendy (20, p. 21, pl. iv.
figs. 3, 4).
18¢8. Pibleiial Jibrosa, Lindgren (15, p. 293, pl. xix figs. 6 a-e).
The specimen is in the form of a small dried fragment
40 millim. in height, finger-shaped and expanded at the upper
end. ‘The colour is pale yellow and the spinules 2 to 3 millim.
in height.
The spicules are strongyles 114 x 3°5 y, slightly curved.
Had it not been for Lindgren’s description of transition
forms in the spicules, which, in his specimens, include oxea,
tornota, and strongyla, I would have hesitated before placing
the specimen in the above species. Doubtless Puchychalina
sptnosissima, Dendy, and Chalina spinifera, Carter, are very
nearly allied to the present species, but it is doubtful whether
they are identical.
Locality. Funafuti Atoll.
Distribution. Off Bahia; Philippines; Funafuti Atoll ;
Cochin China; Java; Christmas I. (?) ; Mergui (?).
Luffariella variabilis (Poléjaeff ).
1884. Luffaria variabilis, Poléjaeff (17, p. 69, pl. ix. figs. 1-6).
1889. Luffaria variabilis, Lendenfeld (18, p. 387, pl. xxxiv. fig. 5).
1899. Luffariella variabilis, Thiele (27, p. 25).
Sponges from Funafuti. 3957
Several small, massive, and digitate specimens of this
sponge occur from depths of 25-86 fathoms, either as dried
washed-out skeletons or preserved in formalin ; the branches
of one of the digitate varieties anastomose. The dermal
membrane is dark slate or black and the interior rich yellow.
Localities. Funamara, 25-45 fath.; Falefatu, 80 fath.;
and Funafuti Islet.
Distribution. Api, New Hebrides, 60-70 fath.; Tahiti,
reefs; Funafuti Atoll.
Luffariella geometrica, sp. n.
(Pl. XV. fig. 1, ac.)
Sponge forming two subspherical masses attached to a
branch of Serdatopora, the larger being 20 millim. in diameter.
Colour reddish brown ; consistence soft and elastic.
The dermal membrane sandy in parts, but in other parts
transparent, the whole sponge being more or less transparent
and allowing one or more concentric spheres (indicating
periods of growth) to be visible in the interior. Oscules level
with the surface and 6 millim. in diameter.
Surface conuli 210 mw, formed by projecting main fibres.
Skeleton forming a regular rectangular network composed
of radiating primary main fibres, joined by secondary hori-
zontal fibres in such a manner as to form triangular shafts ;
tertiary fibres also present, more or less irregularly distributed.
Dermal skeleton a regular reticulum in which usually six
secondary fibres radiate out from each nodal vertical primary
fibre, the joined bases of the resulting triangles forming more
or less regular hexagons; the primary meshes filled in with
a network of more slender fibres.
Fibres.— Primary main fibres 175-210 yw, secondary 110 yp,
tertiary 16 p.
Width of meshes about 700-1500 yw in width and 400-650 wu
in height.
Flagellated chambers subspherical, 32 mw in diameter.
The present form constitutes the fifth described species of
the genus. JL. geometrica is more nearly allied to L. calya,
Lendenfeld, from the Indian Ocean, than to the other species,
but differs from it in being devoid of gyriform protuberances
on the surface.
The pith of the fibres occupies about one third of their
diameter and is clearly differentiated by staining with borax
carmine.
Locality. W. of Tutanga, 30 fath.
355 Mr. R. Kirkpatrick on
Psammopemma purpureum (Carter).
1880. Aplysina purpurea, Carter (3, p. 36).
1881. Aplysina purpurea, Carter (4, p. 103, pl. ix. figs. 1, 2).
1885. Pseudoceratina durissima, Carter (5, p. 204).
1885. Holopsamma fuliginosa, Carter (5, p. 213).
1889. Aplysina purpurea, Dendy (7, p. 97).
1889. Psammopemma fuliginosum, Lendenfeld (13, p. 636).
The sponge, which forms a dark purple incrustation on
Halimeda, has a few cactiform protuberances on its surface
and fragments of the coralline imbedded in its substance.
The ground-substance contains fine fibrillz 2 w in thickness.
After examining the type specimens of Pseudoceratina
durissima, Carter, and H. fuliginosa, I agree with Lenden-
feld in considering these species as synonyms of A. purpurea ;
but I can see no reason for rejecting the oldest species name
purpureum.
Locality. Tutanga, 50 fath.
Distribution. Port Phillip, Victoria; Funafuti Atoll; Gulf
of Manaar.
Stelospongus cavernosus (Pallas), var. pyriformis,
Lendenfeld.
1889. Stelospongus cavernosus, var. pyriformis, Lendenfeld (13, p. 509).
A small specimen, 70 X40 millim. in area by 20 millim. in
thickness, consisting only of the hard brown skeleton of
flattened-pyriform shape.
The type specimen of the variety is in the British Museum
(registered, year 1838. 4. 16. 10), the specimen having been
purchased from a dealer, and it is important to note that the
locality is unknown, and probably not “ West Indies,” as
recorded by Lendenfeld. In fact, the type specimen may
have come from the Pacific, since various specimens presented
by Sir E. Home, who brought home collections from the
Pacific, are registered on the same page. It is not unlikely
that the dealer obtained his specimen from some fellow-
voyager of Sir E. Home.
On the under surface of both specimens an oval patch of
ale brown dermal membrane incrusted with foreign bodies
still persists, and probably coincides with the area of attach-
ment of the sponge.
Locality. Funafuti Atoll.
PoLYFIBROSPONGIA, Bowerbank (1, p. 459).
1889. Hircinia, subgenus Polyfibrospongia, Lendenfeld (18, p. 587).
1884. Stelospongus, Ridley (19, p. 383).
Stelospongine with a loose skeleton-net, the trabecule of
which are formed of fascicles of parallel slender fibres fre-
quently anastomosing.
Sponges from Funafuti. 359
Polyfibrospongia Sweet’, sp.n. (Pl. XV. fig. 2, a-c.)
Sponge forming an irregular incrusting mass, 4 centim. in
length, 2°5 centim. in width, and 1 centim. in thickness,
nearly encircling the stem of a Gorgonia; with long mem-
branous yellow oscular tubes about 2 centim. in height,
occasionally branched. Dermal membrane smooth, perga-
mentaceous, yellowish white.
Skeleton.—Kctosomal: composed of stout primary fibres
30 to 60 w in thickness, partly cored with foreign bodies,
the meshes being filled in by thinner clear fibres and crowded
with foreign bodies; the oscular tubes consisting of a con-
tinuation of the dermal layer, but with the cored fibres forming
a regular network with square meshes.
Choanosomal skeleton composed of an oval-meshed network
of loose bundles of slightly branched anastomosing fibrils, the
diameter of the bundles being 210 w and of the fibrils 9 w;
also of sparsely scattered slightly branched fibres 50 wu thick,
and passing from the base to the surface, composed of foreign
bodies cemented by spongin.
Filaments absent.
This remarkable species is nearly related to Polyfibro-
spongia flabellifera, Bowerbank, a thin flabellate species from
New Guinea, but differs in shape and in having the oscular
tubes. The type specimen of Bowerbank’s species is in the
Dresden Museum, but the Bowerbank collection contains two
slides prepared from the type, and quite sufficient to show the
close similarity in the skeletal structure of the two species.
Lendenfeld puts Bowerbank’s species in a subgenus Poly-
jibrospongia under the genus Hircinia. After a careful search
I have been unable to find filaments in either of the above
two species, though these mysterious objects are abundant in
a specimen from Port Jackson named by Lendenfeld Hircinia
( Polyfibrospongia) gigantea.
To include HH. gigantea in the same genus with Polyfibro-
spongia Sweetiand flabellifera would be tantamount to regarding
the filaments as unessential elements from the systematic point
of view. I havenot at presenta definite opinionon this difficult
controversial question of the systematic value of the filaments,
a question on which equally eminent spongologists take
diametrically opposite views.
For the present I shall retain Bowerbank’s genus with
the two species Sweeti and flabellifera, placing it next to
Stelospongus, and shall leave H. gigantea and fasciculata
under Hircinia. The new species is named after Mr. G.
Sweet, whose munificent donations rendered possible the
success of the Australian Boring Expedition.
Locality. W. of Tutanga, 86 fath., Funafuti Atoll.
=
Mr. R. Kirkpatrick on
. BowrrBank, J. 8. “ Descriptions of Five new Species of Sponges
discovered by A. B. Meyer on the Philippine Islands and New
Guinea,’ Proc. Zool. Soc. 1877, p. 456.
. Carter, H. J. “ Contributions to our Knowledge of the Spongida,’
Ann. & Mag. Nat. Hist. 1879, (6) vol. ix.
. ‘Report on Sponges dredged up from the Gulf of Manaar,”
Ann. & Mag. Nat. Hist. 1880, (5) vol. v.
“ Contributions to our Knowledge of the Spongida,” Ann.
& Mag. Nat. Hist. 1881, (5) vol. viii.
——. “Descriptions of Sponges from Port Phillip Heads,” Ann.
& Mag. Nat. Hist. 1885, (5) vol. xv.
—. “Sponges from South Australia,” Ann. & Mag. Nat. Hist.
1886, (5) vol. xvil.
. Denpy, A. “ Report on Sponges from the Gulf of Manaar,” Ann.
& Mag. Nat. Hist. 1889, (6) vol. iii.
“ A Monograph of the Victorian Sponges, Part I.,” Trans.
Roy. Soc. Vict. 1891.
—. “Catalogue of Non-Calcareous Sponges... . collected from
.... Port Phillip Heads, Part I.,” Proc. Roy. Soc. Vict. 1891,
vol. vii. n. s. 1895.
. DoperLE, L. “Ueber die Lithonina, eine neue Gruppe von
Kalkschwimmen,” Zool. Jahrb. 1897, vol. x. Systematik.
. Hancock, A. “On the Excavating Powers of certain Sponges
belonging to the Genus Cliona,” Ann. & Mag. Nat. Hist. 1849,
(2) vol. iii.
. Hrypz, G. J. “On some remarkable Calcisponges from the
Eocene of Victoria,” Quart. Journ. Geol. Soc. 1900, vol. lvi. p. 50.
. LENDENFELD, R. v. Monograph of the Horny Sponges, 1889.
.—. “Die Clavulina der Adria,’ Nova Acta, 1898, vol. lxix.
. Linpe@ren, N.G. “ Beitrag zur Kentniss der Spongienfauna des
malayischen Archipels und der chinesischen Meere,” Zool. Jahrb.
1898, vol. xi. Systematik, p. 288.
. Lister, J. J. “ Astrosclera Willeyana, the Type of a new Family
of Sponges,” Willey’s Zoological Results, part iv. 1900.
. PotésaEFF, N. Report on the Keratosa collected by H.M.S.
‘ Challenger,’ 1884.
. Riviey, 8. O. “Spongida collected during the Expedition of
H.M.S. ‘Alert’ in the Straits of Magellan and on the Coast of
Patagonia,” Proc. Zool. Soc. 1881.
Report on the Zoological Collections made in the Indo-
Pacific Ocean during the Voyage of H.M.S. ‘ Alert’: Spongiida,
1884.
. Riptey, 8S. O., and Denpy, A. Report on the Monaxonida
collected by H.M.S. ‘ Challenger.’
. Scumipt, 0. Die Spongien des adriatischen Meeres, 1862.
. Grundziige einer Spongien-Fauna des atlantischen Ge-
bietes, 1870.
. ScnuuzE, F. E. “ Untersuchungen tiber den Bau und die Ent-
wicklung der Spongien: III. Die Familie der Chondroside,”
Zeitschr. wiss. Zool. 1877, vol. xxix. p. 87.
“Die Plakiniden,” Zeitschr. wiss. Zool. 1880, vol. xxxiv.
p- 407.
. —. Corticium candelabrum, O. Schmidt,” Zeitschr. wiss. Zool.
1881, vol. xxxv. p. 410.
Sottas, W. J. Report on the ‘Challenger’ Tetractinellida,
1888.
. Turecs, J.“ Studien uber pazifische Spongien,” Zoologica, Stutt-
gart, 1898, Heft 24 (ii.) part 2, p. 1.
Sponges from Funafute. 361
28. Topsent, E. “ Deuxiéms Contribution 4 l'étude des Clionides,’’
Arch, Zool. Expér. 1891, (2) vol. ix. p. 555.
. “Spongiaires de la Baie d’Amboine,” Revue Suisse de
Zoologie (Bedot), 1897, vol. iv. p. 421.
30. Warren. EGGE, T. “ The Sponges of Funafuti,” Australian Mu-
seum, Memoir III. 1897, p. 823.
29.
EXPLANATION OF THE PLATES.
Pirate XIII.
Fig. 1. Plectroninia Hindei, sp. n.
a, Specimen, natural size.
6, c. Large fused quadriradiates, forming skeleton mesh, x 200.
d, e. Smaller quadriradiates of basal layer, X x 200.
i g, h. Triradiates, x 200.
?-l, Tuning-fork ticles, x 400.
m-o. Tylotes and subtylotes, x 400.
p-r. Style and oxea, x 400.
Fig. 2. Placinolopha spinosa, sp. n.
a, Natural size.
b-z. Microcalthrops in graduated sizes, X 200.
k-m. Lophodiactines, x 200.
Fig. 3. Placinastrella clathrata, sp. u.
a. Surface, showing a pore area, X 6.
6. Section, x 10.
e. Small calthrops, x 200.
d. Larger calthrops, X 200.
e-g. Triods, x 200.
h. Dermal diactine, x 200.
zo. Larger (choanosomal) diactines, x 200.
PuaTE XIV.
Fig. 1, a. Fragment of Echinopora bored by Dyscliona Davidi, sp. n.
b. Portion of the sponge separated by decalcification, showing
the lobes, x 10.
ec. Strongyle, x 200.
d. Spirally spined microstrongyle, x 200.
e. The same, X 600.
fg. Spicules of D. purpurea, Hancock, copied from Hancock’s
figures.
Fig. 2. ips Sey clavigera.
a, Specimen incrusting Gorgonia, natural size.
6. Tylote, x 125.
ec, d. Discasters, x 400.
e. Small tylote, x 400.
Fig. 3. Erylus monticularis, sp. n.
a, Surface, x 50. e. Smallest microxea, x 250.
b. Sterraster, x 400. Jf. Orthotrizne, x 100.
ce. Large oxeum, X 200. g. Chiaster, x 400.
d. Medium-sized oxeum, x 250. | A. Oxyasters, x 400,
Fig. 4. Tedania levis, sp. n.
a, Spined strongyle, x 200.
b, Tylote, x 200.
ce. Smooth strongyle, x 200,
d, Trichodragma, x 200.
e. The same with rhaphides compact, x 200.
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 24
362 Mr. R. I. Pocock on the
PuatTE XV.
Fig. 1. Luffariella geometrica, sp. n.
a, Natural size.
b. Section, x 10.
ce. Derma] skeleton, X 25.
Fig. 2. Polyfibrospongia Sweeti, sp. n.
a. Natural size.
b. Section, K 25.
c. Wall of oscular tube, x 15.
d. Bundle of fibres teased out, x 260.
Fig. 3. Chondropsis ceratosus, sp. n.
a. Skeletal network, x 25.
6. Sigmata, x 1265.
c. Oxeum, X 125.
XLIV.—The Scorpions of the Genus Heterometrus.
By R. I. Pocock.
Pror. KRAEPELIN (Das Tierr., Scorp. p. 124, 1899) re-
cognizes, under the name Scorpio *, two species of the genus
Heterometrus, namely maurus and Boehmi, a form referred on
the authority of Birula to testaceus of C. Koch, from Syria,
being regarded as doubtfully distinct. To the synonymy of
maurus is added palmatus of Ehrenberg and propinquus of
Simon, the latter being qualified with a mark of interrogation.
An examination of the material at my disposal has led me
to conclusions by no means in accord with those contained in
‘Yierreich.” A. Boehmi and H. propinquus are unknown
to me; but including the latter 1 recognize the following
four species as occurring in the area inhabited, according to
Prof. Kraepelin, by the one form maurus :—
(1) Heterometrus maurus (Linn.).
Scorpio maurus, Linn, Syst. Nat. ed. 10, p. 624 (1758).
Buthus testaceus, C. Koch, Die Arachn. vy. p. 3, fig. 259 (1839).
Scorpio maurus, Kraepelin, Das Tierr., Scorp. etc. p. 124 (1899) (in
part.).
* The system, if any, of determining the type species of genera, which
is adopted by Prof. Kraepelin, and presumably sanctioned by the Tier-
reich-Committee, is most puzzling. By elimination maurus is not the
type species of Scorpio, since it was removed from that genus under the
name palmatus by Hemprich and Ehrenberg and fixed on to Heterometrus
by Thorell (Ann. & Mag. Nat. Hist. (4) xvii. p. 2,1876). Ifit be regarded
as the type of Scorpio on the strength of standing first under that heading
in the 10th edition of the ‘Systema,’ it must for the same reason be
regarded as the type of Heterometrus. In that case Heterometrus must
be a synonym of Scorpio. Prof. Kraepelin, however, retains by elimina-
tion the name Heterometrus for the second species it originally contained,
namely spinifer.
Scorpions of the Genus Heterometrus. 363
Loc. Maroceo and Algeria. The British Museum has
many examples from Tangier (B. B. Woodward, d&c.), Cape
Spartel (Z'. Annandale), Tunis, and Algiers.
(2) Heterometrus palmatus, Hempr. & Ehrenb.
Heterometrus palmatus, Hempr. & Ehrenb. Symb. Phys., Scorpiones,
no. 1 (1829).
Heterometrus palmatus flavus and ? rufus, iid. ibid.
Scorpio maurus, Kraepelin, op. cit. (in part.).
Loc. Egypt: Cairo (Dr. Anderson).
Hemprich and Ehrenberg recognized two varieties of the
Egyptian species—one from Alexandria, which was named
flavus; the other from Sinai, named rufus. I have seen no
examples from Sinai, but a single male specimen from Cairo
(Dr. Anderson), which is doubtless referable to H. palmatus
flavus, belongs to quite a distinct species from the Algerian
A, maurus.
(3) Heterometrus fuscus, Hempr. & Ehrenb.
Heterometrus palmatus fuscus, Hempr. & Ehrenb. Symb. Phys., Scorp.
no. 1 (1829) ; Simon, Ann. Soc, Ent. Fr. (5) ii. p. 258 (1872).
Heterometrus palmatus, var. minor, Simon, ibid.
Heterometrus testaceus, Birula, Hore Soc, Ent. Ross. xxxiii. p. 138
(1898) ; Kraepelin, op. cit. (nec testaceus, C. Koch).
Loc. Syria: Jerusalem (Herr Rolle), Tiberias (A. Smith
Woodward), &c.
Birula seems to have recognized the distinctness of the
Syrian species, but erroneously used the name ¢estaceus for
it: testaceus was applied by C. Koch to a specimen from
Algeria co-specific with maurus of Linn.
(4) Heterometrus propinquus, Simon.
Heterometrus propinquus, Simon, Ann. Soc, Ent. Fr. (5) ii. p. 259
(1872).
Loc. Syria: Damascus and Nablous.
This species is said to differ from the preceding (/. fuscus)
in having the median eyes larger, the vesicle more globular,
and 14 pectinal teeth.
Simon, unfortunately, did not determine the sex of his
specimens; but since the type of H. propinquus is said to be
smoother than specimens of H. fuscus, it is safe to assume that
the former species is based upon the female sex.
To the above-mentioned I have to add the following new
forms :—
(5) Heterometrus arabicus, sp. n.
Colour a nearly uniform yellowish brown, the legs and tail
24*
364 Mr. R. I. Pocock on the
entirely pale ochre, except for the normal brown spot at the
extremity of the femur and patella in front. Carapace and
sternavery weakly granular laterally. Median eyes larger than
in HZ. fuscus and maurus. Tail narrow, second segment about
as long as wide; intercarinal spaces almost smooth; keels
normal, but the inferiors on the third and fourth segments
scarcely denticulated, merely roughened with setal pores;
vesicle larger, much wider than the fifth and as wide as the
second caudal segment. Hand of chela ornamented above with
a reticulated pattern of low ridges, passing into tubercles
towards the base of the fingers and becoming obsolete on the
inner edge of the hand ; two very strong finger-keels on hand,
the external the stronger. Pectines with 12 teeth; genital
operculum a little wider than long. Spine-armature of fourth
tarsus 10 or 9-7.
Length 10 millim. ; carapace 8; tail 27; width of hand 6:7.
Loc. Arabia (Mrs. Burton).
Differs from H. fuscus in being much paler, much smoother,
and in having the inferior keels on the third and fourth caudal
segments not denticulated, the vesicle large, the hand very
strongly everted, and the median eyes much larger.
In size of vesicle and of median eyes 7. arabicus approaches
Hi. propinquus, Simon (Ann. Soc. Ent. Fr. (5) u. p. 259,
1872), from Damascus and Nablous ; but since Simon noticed
no difference in the carination of the inferior surface of the
tail and of the hand between H. propinguus and fuscus, I
cannot do otherwise than conclude that the two are alike in
these particulars.
(6) Heterometrus Townsendt, sp. n.
? .—Resembling the preceding in colour, granulation, &e.,
but with the hand almost entirely smooth above and much
less strongly crested, the genital operculum as long as wide,
the vesicle only slightly wider than the fifth and much narrower
than the second caudal segment.
Total Jength 59 millim.; carapace 9:2; tail 28; width of
hand 7°5.
Loc. Fort Reshire (near Bushire) and Bushire on the
Persian Gulf (F. W. Townsend).
(7) Heterometrus Boehm, Kraepelin.
Heterometrus Boehmi, Kraepelin, MT. Mus. Hamb, xiii. p. 187 (1896).
Scorpio Boehmi, id. Das Tierreich, Scorp. ete. p. 125 (1899).
Loc. Lake Tanganyika.
Evidently very distinct from the foregoing species by the
Scorpions of the Genus Heterometrus. 365
smoothness of the inferior keels on the last abdominal and first
caudal segments, the spine-armature of the tarsi, &e.
The known species may be tabulated as follows :—
a. Keels of last abdominal sternum and of underside of
first caudal segment smooth (2) ....... Soyente gas Boehm,
6. Keels of last abdominal sternum and of underside of
first caudal segment coarsely granular (¢ 2 ).
a‘, Genital operculum divided ; terga and sterna finely
granular or shagreened (<¢ ).
a’, Hand in adult wider than length of carapace,
distinctly crested above ; immovable finger short,
triangular; a crest on underside of brachium
behind; tail thick, fifth segment less than
twice as long as wide, third as wide as or wider
CLAS Oca SON CI groin sok ere miei MAUrUS.
6, Hand narrower than length of carapace, not
visibly crested above; immovable finger long;
no crest on underside of brachium ; tail narrow,
third caudal segment longer than wide ...... palmatus.
6‘. Genital operculum undivided ; terga and sterna not
shagreened (2).
a*, Third and fourth caudal segments with inferior
keels strong and granular.
a‘, Median eyes small, vesicle not inflated.
a’. Genital operculum much wider than long,
transversely elliptical; lower side of bra-
chium with distinct smooth crest behind ;
hand more strongly granular and keeled, &e. mauwrus.
6°. Genital operculum heart-shaped, as long or
nearly as long as wide ; lower side of bra-
chium evenly rounded behind; hand less
coarsely granular and crested .......... Suscus.
6', Median eyes larze ; vesicle expanded........ propinguus,
8, Inferior keels of third and fourth caudal segments
smooth, only uneven with setal pores; median
eyes large.
a°, Hand with a pair of strong smooth finger-keels,
and ornamented above with a distinct net-
work. of smooch ridges cc cmc e wens ene e ahs arabicus.
6°. Hand almost smooth above, scarcely visibly
sculptured ; its crests obsolete ............ Townsendi.
N.B.—Attention must be drawn to the omission from the
‘Tierreich ’ of the following names which have been applied
to various forms of the genus Heterometrus: flavus, rufus,
fuscus, and minor. Apart from the possibility that these
forms may in future take rank as distinct species, the re-
cording of such names is necessary to prevent their applica-
tion to new species by students who have no opportunity of
consulting all the literature of the subject and look to a work
of the aspirations of the ‘Tierreich’ to supply at least a
complete list of the names that have been employed in each
genus.
366 Mr. W. L. Distant on Coreide.
XLV.—Rhynchotal Notes.—V II. Heteroptera: Fam. Coreide.
By W. L. Distant.
THE following notes and descriptions refer to the subfamilies
Merocorinz, Mictine, Amorbine, Petasceline, and Dala-
derine. Vol. iv. of Walker’s Catalogue refers to the Coreide,
and the species and genera therein described are here reviewed
so far as they belong to the subfamilies named above. A
number of new species contained in the British Museum are
described and also a few from my own collection,
Merocortnz.
MENARDUS, gen. nov.
Head beneath armed with a prominent spine or tubercle.
Scutellum about the length of the clavus, entire near base.
Mesosternum with two small tubercles at base. Posterior
femora armed beneath with two rows of strong spines or
tubercles. Other characters as in Flavius, to which it is
primarily allied by the prominent tubercles on the under
surface of the head, and from which it may be at once sepa-
rated by the much shorter scutellum. .
Menardus notatus.
Meropachys notatus, Walk. Cat. Het. iv. p. 70. n. 6 (1871).
Hab. N. Brazil, Central America.
Genus PERANTHUS.
Peranthus tinctus.
Mirileus tinctus, Walk, Cat. Het. iy. p. 72, n. 5 (1871).
Micrinz.
Genus DEREPTERYX.
Derepteryx Hardwicki.
Cerbus (Derepteryx) Hardwickii, White, Ann. & Mag. Nat. Hist. (2)
iii, p. 542 (1839). °
Mictis amplectens, Walk. Cat. Het. iv. p. 25. n. 58 (1871).
Genus PRIONOLOMIA.
Prionolomia porrigens.
Trematocoris porrigens, Walk. Cat. Het. iv. p. 85, n. 11 (1871).
Mr. W. L. Distant on Coreide. 367 ©
Prionolomia fulvicornis, Fabr., var. ?
The unique type is an unlocalized male specimen erro-
neously described by Walker as a female. Fourth joint of
the antenne subequal in length to first joint, not “ shorter
than the first.”
Very closely allied to P. fulvicornis, Fabr.; posterior tibiee
more slender, posterior femora a little more granulated.
Prionolomia mandarina, sp. n.
Fuliginous-brown; body beneath, legs, and antenne
piceous (in some specimens the colour is fuliginous brown
throughout) ; apical joint of antenne ochraceous. Abdomen
above red, connexivum and apex fuliginous.
Antenne with the first and fourth joints subequal in length,
second a little longer than the third; pronotal angles widely,
broadly, and spatulately extended, their apices subtruncate
and slightly directed upwards, their anterior margins strongly
dentate, their posterior margins undulated ; in some specimens
the pronotal basal margin is somewhat deeply notched in
front of scutellum, but this is not a constant character; con-
nexivum distinctly extending in front of the posterior half of
the corium.
6. Anterior and intermediate tibia moderately outwardly
dilated at a little beyond base; anterior and intermediate
femora with a distinct spine beneath near apices; posterior
femora incrassated, strongly tuberculated in a longitudinal
series above and with a much more obsolete series beneath,
and with two strong acute spines beneath, one near centre,
the other a little before apex; apex angulated and finely
dentate beneath; posterior tibie dilated on each side, in-
wardly broadly angulated at about centre, and outwardly
convexly widened in the same position.
9. Posteriur femora moderately incrassated, tubercles.
above more obsolete than in male; posterior tibiee convexly
dilated on each side near base, not inwardly angulated as in
male.
Long., ¢ 2, 22-26 millim.; exp. pronot. angl. 11-12
millim.
Hab. China, Kiukiang (Pratt, Brit. Mus.).
Genus PHYLLOGONIA.
Phyllogonia limosa.
Sulpicia limosa, Walk. Cat. Het. iy, p. 39. n. 5 (1871),
368 Mr. W.L. Distant on Coreide.
Genus EKLASMOPODA.
Elasmopoda affinis, sp. 0.
3. Dark pitchy brown; corium shortly ochraceously
pilose; antenne, eyes, rostrum, anterior and intermediate
legs, and posterior tarsi ochraceous ; anterior and intermediate
coxw and apices of femora piceous; body beneath reddish
ochraceous, pro- and mesosternum somewhat paler, abdominal
spines piceous ; disk of apical segment and a marginal spot
on apical and penultimate segments piceous, the marginal
spots preceded by ochraceous shading.
Pronotum somewhat narrow, anterior lateral margins entire,
not dentate, lateral angles produced in foliaceous subacute
spines, their margins dentate.
Long. 21 millim.; exp. pronot. angl. 8 millim.
Hab. British East Africa, Samburu (C. S. Betton, Brit.
Mus.).
Closely allied in structure, colour, and general appearance
to HL. undata, Dall., but differing by the narrower pronotum,
with its non-dentate anterior lateral margins and its narrower
and more acute lateral angles.
Genus HOLOPTERNA.
flolopterna Rothi.
Mictis Rothii, Dall. List Hem. 1. p. 395. n. 28 (1852).
Cypia rubra, Lethierry, Ann, Mus. Civ. Genova, xvi. p. 286 (1881).
Cipia rubra, Leth. & Sev. Cat. Gén. Hém. t. ii. p. 8 (1894).
This species varies in the colour of the anterior and inter-
mediate femora, which are either red or black ; the corium is
either unicolorous or with the apical half darker. Dallas’s
specimens ot HH. Rethii exhibit the variation shown in
Lethierry’s species.
Holopterna Elliott, sp. n.
Castaneous; head, scutellum, inner claval margin, lateral
areas of corium (excluding apex), membrane, rostrum, coxe
and trochanters, anterior and intermediate femora, base of
posterior femora, posterior tibia, a broad central mesosternal
patch, abdominal tubercles, and apex of abdomen piceous.
Antenne with the third joint shortest, second and fourth
joints subequal, first joint a little longest and thickest. Head
very distinctly cleft at apex, eyes brownish. Pronotum
finely rugulose, lateral margins distinctly pilose, lateral
Mr. W. L. Distant on Coreide. 369
angles subacutely produced, slightly directed upwards. Scu-
tellum transversely rugulose. Corium thickly and somewhat
coarsely punctate. Membrane opaque. First and second
abdominal segments each with two robust conical tubercles.
Long., ¢ 22, 2 25 millim.
Hab. East Africa; Ruwenzori, 7000-8000 feet (Scoté
Elliot, Brit. Mus.).
EVAGRIUS, gen. nov.
Allied to Holopterna, Stal, but differing in the following
particulars :—
g. First joint of the antenne longer than the fourth joint,
but considerably shorter than the “third and fourth joints
together ; third joint moderately dilated and grooved, a little
shorter than fourth joint. Second, third, and fourth abdo-
minal segments armed with a prominent tubercle on each side
of disk.
The pronotal angles are very strongly, laminately, and
arcuately produced, and between their area and the head the
pronotum is suddenly and profoundly deflected.
Evagrtus gladius, sp. n.
Pale castaneous, femora and antenne darker in hue, the
last with the apical joint pale stramineous; membrane
piceous ; abdominal tubercles black ; meso- and metasternum
with a broad oblique ochraceous fascia on each side. An-
tennz with the bases and apices of the second and third joints
very narrowly ochraceous, third joint moderately dilated and
distinctly grooved, fourth joint the most slender and cylin-
drical. Pronotum with the depressed space between the area
of the angles and head ochraceously pilose, with a narrow
obscure central fascia extending therefrom to base; the angles
extremely prominent, laminate, somewhat arcuately directed
forwards and upwards, their apices slightly recurved and
subacute, their margins moderately dentate, posterior femora
with a prominent tooth and some smaller teeth near apex ;
posterior tibizs moderately dilated, with an inner prominent
spine near apex.
Long. 23 millim.; max. abd. lat. 6 millim.; exp. pronot.
ang]. 11 millim.
Hab. Brit. East Africa; Maziwa Mitatu and Maungu
(C. S. Betton, Brit. Mus.).
370 Mr. W. L. Distant on Coreides.
Genus PTERNISTRIA.
Pternistria octolineata.
Melucha octolineata, Walk. Cat. Het. iv. p. 56, n. 10 (1871).
Genus MYGDONIA.
Mygdonia tuberculosa.
Mictis tuberculosa, Sign. Rev. et Mag. Zool. 1851, p. 448, pl. xv. fig. 6.
Melucha atra, Walk. Cat. Het. iv. p. 55. n. 7 (1871).
Genus OCHROCHIRA.
Ochrochira lunata, sp. n.
Fuliginous-brown ; apical joint of the antennz fulvous ;
abdomen above red, connexivum and apex fuliginous.
Antenne with the first joint a little longer than the fourth,
second a little longer than the third; pronotum with the
lateral angles widely, broadly, and somewhat lunately pro-
duced, their margins dentate anteriorly, profoundly crenulate
posteriorly, their apices subacute and slightly directed up-
wards. ‘The body (excluding membrane) is finely ochra-
ceously pilose.
3g. Anterior and intermediate femora with a distinct spine
beneath near apex, the intermediate femora also with a series
of short spines beneath; posterior femora incrassated and
with two or three distinct tuberculous spines near apex
beneath, and with a few spines on apical half of inner margin ;
posterior tibiae moderately dilated, spinously produced at
about two thirds from base, and thence narrowed and in-
wardly dentate to apex.
@. As in male, but posterior femora much less incrassated
and only spined at apex; posterior tibie simple, longitudi-
nally sulcated.
Long., ¢ ?, 25-26 millim. ; exp. pronot. ang]. 6-7 millim.
Hab. China; Kiukiang (Pratt, Brit. Mus.).
This species finds its nearest ally in O. fuliginosa, Uhler,
from Japan, from which it is separated by the much more
produced and lunate pronotal angles.
Genus Mictis.
Mictis lateralis.
Mictis lateralis, Walk. Cat. Het. iv. p. 29. n. 69 (1871).
Allied to M. albovittata, Stal, from which it differs by the
Mr. W. L. Distant on Coreide. 371
absence of the pale longitudinal fasciz beneath and by the
lateral angles of the pronotum being directed outwardly as in
Stal’s species, but not recurved posteriorly at their apices.
Mictis longicornis.
Myctis longicornis, Westw. in Hope Cat. ii. p. 11 (1842).
Mietis javana, Walk. Cat. Het. iv. p. 30. n. 70 (1871).
Mictis filicornis.
Mictis filicornis, Walk. Cat. Het. iv. p. 27. n. 64 (1871).
Mictis ambotinensis.
Mictis amboinensis, Walk. Cat. Het. iv. p. 28. n. 66 (1871).
Mictis profana.
Lygeus profanus, Fabr. Syst. Rhyng. p. 211. n. 33 (1803).
Mictis symbolica, Dall. List Hem. ii. p. 404. n. 52 (1852),
Mictis crux, Dall. loc. cit. p. 405, n. 53,
Mictis limbativentris.
Mictis limbativentris, Stal, Trans. Ent. Soc. Lond. 1863, p. 603.
This species was described by Stal from a female specimen
only, and was afterwards placed by him (En. Hem. 111. p. 51,
1873) in “ Species Mictariorum tncerti generis.” ‘The British
Museum has since acquired two other specimens (one a
male), of which the following are the principal structural
characters :—
é. Posterior tibiz moderately dilated on each side, with a
strong spine on inner surface at about one third from apex ;
second abdominal segment with a large raised conical tubercle
on each side parallel with the posterior coxe.
Long., ¢, 30 millim.
Hab. New Guinea, Dorey ( Wallace, Brit. Mus.).
Mictis farinulenta.
Mictis farinulenta, Bredd. Mitt. nat. Mus. Hamb. xvi. p. 168 (1899), ?.
3. Abdomen beneath at junction of second and third
segments broadly and tuberculously elevated ; posterior tibiz
gradually dilated from base on inner side to a somewhat
broad tooth about centre, from thence concavely narrowed
to apex, and in structure somewhat resembling the femora of
M. tenebrosa, Fabr.
Long., ¢, 21 millim.
372 Mr. W. L. Distant on Coreide.
The British Museum possesses both sexes from Lombok
and a female specimen from Savu, Philippines, all collected
by the late Mr. Everett.
Mictis oceanensis, sp. n.
Castaneous, obscurely and finely ochraceously pilose ; apical
joint of antenna, apices of tibie, tarsi, and sternal segmental
margins ochraceous; abdomen above ochraceous, lateral
margins, a central longitudinal fascia, and the apex black ;
tarsal claws piceous.
Antenne with the first and second joints subequal in
length, third a little longer than fourth ; pronotum with the
lateral margins very strongly and coarsely serrated, the serra-
tion piceous, the lateral angles prominent and broadly sub-
acute, its surface reticulately rugulose and finely but obscurely
punctate; scutellum transversely rugose ; corium obscurely
and finely punctate; rostrum about reaching the intermediate
coxe, its apex black.
3. Abdomen at junction of second and third abdominal
segments broadly and_ tuberculously elevated; posterior
femora moderately incrassated, very finely and obscurely
tuberculate, with a lineate carina on outer edge and with
two short blunt teeth at apex; posterior tibiz sulcate,
gradually dilated interiorly into a somewhat broad tooth
about centre, thence concavely narrowed to apex.
@. Abdomen unarmed; posterior tibie simple, sulcaie, —
and slightly dilated near base.
Long., ¢ ?, 25 millim. ; exp. pronot. angl. 9 millim.
Hab. New Hebrides (Dr. D. McNabb, Brit. Mus.).
A species to be primarily recognized by the coarsely
serrated pronotal margins and by the distinct colour of the
upper surface of the abdomen.
Genus ANOPLOCNEMIS.
Anoplocnemis tristator.
Lygeus tristator, Fabr. Syst. Rhyng. p. 206. n. 18 (1803).
Mictis luteitarsis, Walk. Cat. Het. iv. p. 19, u. 84 (1871).
Anoplocnemis mosta.
Mictis mesta, Dall. List Hem. ii. p. 400. n. 41 (1852).
Posterior tibize in male angulated as in A. pagana, Dall.
Anoplocnemis Dallasiana, Leth. & Sev.
Mictis scutellaris, Dall. (nom. proce.) List Hem.1i. p. 390. n. 17 (1862).
Mr, W. L. Distant on Coreide. 373
The type of this species is no longer to be found; there
are, however, a long series of specimens in the British
Museum which undoubtedly belong to it, and which have
been received from §. Africa, Angola, Zomba, and the
‘Transvaal.
Anoplocnemis gracilicornis.
Mictis gracilicornis, Stal, Hem. Afr. ii. p. 42. n. 22 (1865),
Melucha aurulenta, Walk. Cat. Het. iv. p. 55, n. 8 (1871).
Anoplocnemis phasianus.
Lygeus phasianus, Fabr. Spec. ii. p. 861 (1781).
Mictis dubia, Dall. loc. cit. p. 389. n.13: 2.
Mictis castanea, Dall. List Hem. ii. p. 389. n. 14 (1852): 2.
Mietis lata, Dall. loc. cit. p. 390. n. 15: 9.
Mictis ferrifera, Walk. Cat. Het. iv. p. 24. n. 57 (1871).
Physomerus mictiformis, Walk. loc. cit. p. 61.n. 8: Q.
Anoplocnemis signata, sp. n.
3. Very dark castaneous, finely ochraceously pilose, espe-
cially on the pronotum; apical third of the scutellum ochra-
ceous ; antenne with the first, second, and third joints casta-
neous, fourth joint ochraceous, with its apical half infuscated ;
body beneath and legs a little darker in hue, anterior and
intermediate tibiz and all the tarsi castaneous; sternum
densely ochraceously pilose, more or less exhibiting distinct
Jateral ochraceous lines. Abdomen above reddish, with the
margins and apex pitchy.
@. Slightly paler in hue; abdomen more ochraceously
pilose beneath.
Antenne pilose, with the first and fourth joints subequal
in length, second a little longer than the third; apex of the
second segment of the abdomen in the male produced at
centre in a flat oblong process extending about halfway across
the third segment. Femora in male strongly incrassated,
curved, and distinctly angulated beneath about one third from
apex ; tibie in both sexes compressed, moderately and evenly
dilated.
Long., ¢ 16 millim., 2? 17-18 millim.
Hab. East Africa, Ruwenzori, 6000-8000 feet (Scote
Lilliot, Brit. Mus.).
Apparently allied to A. castaneicornis, Stal, from which it
differs by the colour of the upper surface of the abdomen,
marking of the scutellum, smaller size, &c.
374 Mr. W. L. Distant on Coreida.
Anoplocnemis Whytet, sp. n.
Castaneous; corium dull stramineous, its lateral margins
irregularly castaneous ; posterior tarsi dull ochraceous ; abdo-
men above black, its lateral margins and a central lineate
spot on anterior margin of fourth and fifth segments ochra-
ceous ; connexivum and apex castaneous ; membrane bronzy.
Body and legs finely ochraceously pilose; antennae with the
basal joint a little darkest, first, second, and fourth joints
subequal in length, third shortest ; pronotum with the lateral
angles subprominent and rounded ; body beneath somewhat
paler ; sternum thickly ochraceously pilose, with an indistinct
ochraceous fascia near the coxe.
g. Abdomen beneath broadly and tuberculously gibbous
at the junction of the second and third abdominal segments;
posterior tibize inwardly angulated at less than midway from
base, and thence finely dentate to apex.
?. Posterior tibiz simple, but finely dentate along inner
margin.
Long., g 22 millim., 9 25 millim.
Hab. Nyasaland, Nyika Mts., 6000-7000 feet (A. Whyte,
Brit. Mus.).
Genus MELUCHA.
Melucha Biolleyt, sp. n.
@. Fulvous; first and second joints of antenna, lateral
margins and apices of lateral angles to pronotum, about basal
half of lateral margins to corium, lateral margins and apex of
scutellum, and a large basal spot to membrane black ; body
beneath more rufous than above, intermediate and posterior
femora luteous, posterior femora with the apex and apical
spines black, posterior tibize with about apical third luteous,
basal rufous portion very narrowly edged with black ; con-
nexivum fuliginous ; membrane cupreous.
Antennz with the first joint longer than the second, their
bases narrowly rufous ; remaining joints mutilated ; pronotum
coarsely punctate and rugulose, its lateral margins finely
crenulate, the lateral angles acutely and straightly produced,
with their posterior margins crenulate; scutellum coarsely
punctate, basal margin levigate; corium finely punctate ;
posterior femora armed with about five short teeth ; posterior
tibie outwardly dilated, convex, not toothed.
Long., ?, 22 millim. ; exp. pronot. ang]. 9 millim.
Hab. Costa Rica; Turrialba, Atlantic slopes (P. Biolley,
Coll. Dist.).
Mr. W. L. Distant on Coreide. 375
Genus MozENaA.
Mozena alata, sp. n.
In coloration and general appearance closely allied to
M., lunata, Burm., but differing from that species by the more
produced pronotal angles, which have their apices more elon-
gately acute, less directed forward, and slightly recurved at
their tips.
Long., ¢ ¢, 21-22 millim.; exp. pronot. angl. 12 millim.
Hab. Costa Rica; Tuis, Atlantic slopes (P. Biolley, Coll.
Dist.).
Genus NEMATOPUS.
Nematopus fasciatus.
Nematopus fasciatus, Westw. in Hope Cat. ii. p. 14 (1842).
Nematopus decoratus, Walk. Cat. Het. iv. p. 78. n. 14 (1871),
Genus ACANTHOCERUS.
Acanthocerus clavipes.
Coreus clavipes, Fabr. Syst. Rhyng. p. 196 (1803).
Camptischium tenebrosum, Walk. Cat. Het. iv. p. 114, n. 4 (1871),
Camptischium verrucosum, Walk. loc. cit. p. 115. n. 6.
Camptischium subvarium, Walk. loc, cit. p. 116. n. 7.
Acanthocerus sublevis.
Camptischium subleve, Walk. Cat. Het. iv. p. 115. n. 5 (1871).
Acanthocerus lobatus.
Crinocerus lobatus, Burm, Handb. 11. 1, p. 318. n. 2 (1835),
Acanthocerus lobatus, Uhler, Proc. Zool. Soc. Lond. 1894, p. 178.
Acanthocerus tuberculatus, Uhler (nec Herr.-Schaff.), op. cit. 1898,
p. 705.
Prof. Uhler has returned specimens collected by Mr. H. H.
Smith on the island of St. Vincent as belonging to A. tuber.
culatus, H.-S.; this identification was probably a slip of the
pen, as they cannot be separated from other specimens from
the island of Grenada and also identified by Mr. Uhler as
A. lobatus, Burm. It is therefore necessary to erase the name
of A, tuberculatus, Herr.-Sch., from the fauna of these islands,
at least so far as the collections made by Mr. Smith enable us
to form an opinion.
376 Mr. W. L. Distant on Coreidee.
AMoRBINE.
Genus AMORBUS.
Amorbus alternatus.
Amorbus alternatus, Dall. List Hem. i. p. 408. n. 1 (1852).
Amorbus planus, Walk. Cat. Het. iv. p. 42. n. 11 (1871).
PETASCELINE.
Genus PETILLIA.
Petillia calcar.
Mictis calear, Dall. List Hem. ii. p. 397. n. 33 (1852).
Trematocoris subvittata, Walk. Cat. Het. iv. p. 34. n. 9 (1871).
Trematocoris vittata, Walk. loc. cit. p. 36. n. 12.
Melucha notatipes, Walk. loc. cit. p. 56. n. 9.
The Melucha notatipes, Walk., represents a rudimentary
form of P. calcar.
Petillia biserrata.
Mictis biserrata, Walk. Cat. Het. iv. p. 29. n. 68 (1871).
DaALADERINZ.
Genus DALADER.
Dalader Horsfieldi, sp. n.
Pale brownish ochraceous; antenne, femora, and mem-
brane fuscous ; basal and apical joints of antenna, tibia, and
tarsi brownish ochraceous, mottled with fuscous.
Antenne moderately hirsute, first joint slightly longer than
the second, third dilated and spatulate; head and pronotum
with a faint central pale levigate line; pronotum with the
anterior margins finely dentate, the lateral angles broadly and
angularly produced, posterior margin concave before the base
of the scutellum; abdomen above reddish ochraceous; the
segmental margins ochraceous ; connexivum brownish ochra-
ceous, with subquadrate pale fuscous spots ; femora somewhat
thickly spined in longitudinal series beneath.
Long., dg, 20 millim. ; exp. pronot. angl. 83 millim.
Hab. Java (Horsfield Coll., Brit. Mus.).
This small species is allied to D. rubiginosus, Westw.,
from which it differs by the much more produced pronotal
angles.
Mr. W. L. Distant on Coreide. or
Genus HORMAMBOGASTER.
Hormambogaster, Karsch, Ent. Nachr. 1892, p. 131 (May).
Ovengua, Dist. Ent. Month. Mag. 1892, p. 285 (November).
Hormambogaster expansus.
Hormambogaster expansus, Karsch, Ent. Nachr. 1892, p. 181 (May).
Ovengua aperta, Dist. Ent. Month. Mag. 1892, p. 285 (November).
Summarized Disposition of Walker's Genera and Species.
Merocorinz, Mictine, Amorbinz, Petasceline, and
Daladerine.
Genus treated as synonymic.
Mictoides, Walk. Cat. Het. iv. p. 88 (1871),=Gen. Curtius, Stal.
Species considered valid and described under correct Genera.
Derepteryx truncata, Walk. Cat. Het. iv. p. 11. n. 4 (1871). (The type
is an immature form of the species.)
Mictis filicornis, Walk. loc. cit. p. 27. n. 64.
ambotnensis, Walk. loc. cit. p. 23. n. 66.
lateralis, Walk. loc. cit. p. 29. n. 69.
Archimerus tndecorus, Walk. loc. cit. p. 64. n. 17.
Phidippus asper, Walk. loc, cit. p. 71. n. 1.
Nematopus ferrinus, Walk. loc. cit. p. 77. n. 13.
varius, Walk. loc. cit. p. 78. n. 15.
Species considered valid, but requiring generic revision.
Mictis biplagiata, Walk. Cat, Het. iv. p. 22. n. 51 (1871), belongs to gen.
Ochrochwra,
insolita, Walk. loc. cit. p. 27. n. 65, belongs to gen. Liaspis.
biserrata, Walk. loc. cit. p. 29. n. 68, belongs to gen. Petzllia.
Trematocoris pardalipes, Walk. loc. cit. p. 33. n. 3, belongs to gen. Petillia.
notatipes, Walk. loc. cit. p. 34. n. 5, belongs to gen. Petillia.
bicoloripes, Walk. loc. cit. p. 35. n. 10, belongs to gen. Petillia.
porrigens, Walk. loc, ett. n. 11, belongs to gen. Prionolomia.
elegans, Walk. loc. cit. p. 37. n. 14, belongs to gen. Petzliia.
patulicollis, Walk. loc. cit. n. 15, belongs to gen. Petillia,
Sulpicia limosa, Walk. loc. cit. p. 39. n. 5, belongs to gen Phyllogonia.
Melucha octolineata, Walk. loc. cit. p. 56. n. 10, belongs to gen. Pternistria.
Physomerus nigrorufus, Walk. loc. cit. p. 60. n. 7, belongs to gen. Ochro-
chira.
Meropachys notatus, Walk. loc. cit. p. 70. n. 6, belongs to gen. Menardius,
gen, Noy. ’
Hirileus tinctus, Walk. loc. cit. p. 72. n. 5, belongs to gen. Peranthue.
Camptischium subleve, Walk. loc. cit. p. 115. n. 5, belongs to gen. Acan-
thocerus, Pal. Beauv.
Ann, & Mag. N. Hist. Ser. 7. Vol. vi. 25
378 Mr. W. L. Distant on Coreide.
Species treated as synonymic.
Mictis luteitarsis, Walk. Cat. Het. iv. p. 19. n. 34 (1871),= Anoploene-
mis tristator, Fabr.
ee laces Walk. doc. cit. p. 28. n. 58, = Ochrochira fulginosa,
er.
ferrifera, Walk. loc. cit. p. 24. n.57,= Anoploenemis phasianus, Fabr.
— reer, Walk. loc. cit. p. 25. n. 58,=Derepteryx Hardwicki,
hite.
insularis, Walk. loc. cit. p. 26. n. 63,= Pternistria macromera, Guér.
—— javana, Walk. loc. cit. p. 30, n. 70,= Mictis longicornis, Westw.
Trematocoris subvittata, Walk. loc. cit. p. 34. n. 9,= Petilla calcar, Dall.
vittata, Walk. loc. cit. p. 36. n. 12,= Petillia calear, Dall.
Amorbus planus, Walk. loc. cit. p. 42. n. 11,=Amorbus alternatus, Dall.
Melucha atra, Walk. loc. cit. p. 55. nu. 7,= Mygdonia tuberculosa, Sign.
aurulenta, Walk. loc. cit. n. 8,= Anoplocnemis gracilicornis, Stal.
notatipes, Walk. loc. cit. p. 56. n. 9,= Petillia calcar, Dall.
Physomerus mictiformis, Walk, loc. cit. p. 61. n. 8,= Anoploenemis phasi-
anus, Fabr,
Bs muticus, var., Walk. loc. cit. p. 63. n. 10,= Capaneus tetricus,
Stal.
—- acutiusculus, Walk. loc. cit. p. 64. n. 16,=Lycambes varicolor, Stal.
guttiventris, Walk. loc. cit. p. 65, n. 18,=Lycambes varicolor, Stal.
maculifer, Walk. loc. cit. p. 65. n. 19,= Archimerus scutellaris, Stal.
Archimerus dolosus, Walk. loc. cit. p. 66. n. 20,= Capaneus odiosus, Stal.
Hirileus collaris, Walk. loc. cit. p. 73. n. 6,= Lycambes varicolor, Stal.
Nematopus decoratus, Walk. loc. cit. p. 78. n. 14,=Nematopus fasciatus,
Westw.
Camptischium tenebrosum, Walk. loc. cit. p. 114. n. 4,=Acanthocerus
clavipes, Fabr.
a Walk. loc. cit. p. 115, n. 6,=Acanthocerus clavipes,
Fabr.
subvarium, Walk. loc. cit. p. 116. n. 7,= Acanthocerus clavipes, Fabr.
SYNONYMICAL NOTES.
Fam. Pentatomide.
TESSARATOMINE.
Lyramorpha picta.
Lyramorpha picta, Dist. Ann. & Mag. Nat. Hist. (6) xi. p. 480 (1893).
Lyramorpha Vollenhovii, Horvath (part.), Term. Fiiz. xxiii, p. 351
(1900), tab. ix. fig. 7, tab. x. fig. 7.
Dr. Horvath has included my ZL. picta asa synonym of
L. Vollenhovii, Stal, a species he has evidently not seen, and
apparently misled by the spotted corium— a speciebus reli-
quis hujus subgeneris corio maculato facillime distinguenda.”
The following characters, partly derived from Dr. Horvath’s
own description and figures, will serve to differentiate the
Species :——
Mr. W. L. Distant bi Coreidee.
L. Vollenhovit, Stl (Horv.).
3. Posterior angles of sixth
abdominal segment broadly and
obtusely angulated.
379
L. picta, Dist.
3d. Posterior angles of sixth ab-
dominal segment acutely and more
elongately produced; inner mar-
gin gradually narrowing to apex.
Antenne brownish ochraceous;
apex of the first joint, upper sur-
face of the second, apex of the
third, and the whole of the fourth
and fifth joints (excluding their
bases) fuscous.
Long. 21 millim.
Antenne “fusco-ferrugineis, arti-
culo primo subtus dilute roseo.”
Long, 24-25 millim.
Lyramorpha diluta.
Lyramorpha diluta, Stl, Trans. Ent. Soc. Lond. 1863, p. 598; Bredd.
Ent. Nachr. xxvi. p. 35, n. 2, 3, fig. 4 (1900); Horv. Term. Fiiz.
Xxiil. p. 348. n. 6, tab. ix. fig. 11, 2 (1900), nec fig. 3, ¢.
Stal’s type (a female specimen) is in the collection of the
British Museum. Breddin has correctly figured the anal
characters of the male, but Horvath’s figure (tab. ix. fig. 3, )
does not apply.
Genus TAMOLIA.
(Horvath, Term, Fiiz. xxiii. p. 8365 (1900).)
Tamolia Horvathi.
Tamolia ramifera, Horv. (nec Walk.) Term. Fiiz. xxiii. p. 8365 (1900).
Walker’s unique type of Lyramorpha ramifera (Cat. Het.
ili. p. 476. n. 4, 1868) is in imperfect condition, possessing
only the first, second, and third joints of the antenne, and
although somewhat divergent from Lyramorpha by the shape
of the body, I considered it best under the circumstances to
leave the species in that genus (cf. Ann. & Mag. Nat. Hist.
ser. 7, vol. vi. p. 62). Dr. Horvath has, however, founded a
genus (Zamolia) on a specimen, “uti videtur, nonnihil imma-
turum,” which he takes to be Walker’s species, but which is
clearly proved to be not so by the character he describes—
‘spina ventrali antrorsum ante coxas anticas producta.” In
L. ramifer the ventral spine does not reach the anterior
coxe. Walker’s species cannot therefore be considered the
type of Horvath’s genus.
Plisthenes dilatatus.
Tessaratoma dilatatum, Monty. Ann. Sci. Phys. (2) vii. 1, p. 100 (1855).
Plisthenes dilatatus, Dist. Trans. Ent, Soe. Lond. 1880, p. 151; Ann, &
25*
380 Mr. W. F. Kirby on some
ee Nat. Hist. (6) iii. p. 272 (1889) ; Horv. Term. Fiiz. xxiii. p. 363
900).
Plisthens ventralis, Horv. Term. Fiiz. xxiii. p. 864 (1900).
Dr. Horvath, who has examined the type of Montrouzier’s
species, is able to give a distinct character as “ articulo quarto
antennarum articulo tertio distincte breviore.”’ Following
this up he has specifically described a very closely allied
species as P. ventralds in which the third joint of the antennz
is compared with the fourth “ subbreviore.”’ As a synonym
of this new species he adds P. dilatatus, Dist. (cf. supra). I
cannot quite understand this course, as Dr. Horvath has not
seen the specimen thus identified, nor did I in the structural
differentiation between the species of Fabricius and Mon-
trouzier allude to these characters, as most of my specimens
have unfortunately reached me minus the apical antennal
joint. Inow, however, possess a specimen from the island of
Bouro with complete antenne which agrees with Horvath’s
character and in all other respects with the specimens I pre-
viously identified as P. dilatatus, Montr. Another character
relied upon by Dr. Horvath, viz. the intensity of the fasciz
to the underside of the abdomen, cannot, with the material
before me, be maintained, and is clearly a variable character.
Plisthenes ventralis, Horv., may be a distinct species, but
certainly not if the specimens I identified as P. dilatatus are
synonymic with it, as stated by Dr. Horvath.
XLVI.—Notes on some Insects from the Yang-tse-Kiang.
By W..’F. Kirey, F.0.8., FS. cc.
DuRING a journey on the Yang-tse- Kiang a small collection
of insects was formed by Capt. A. W. 8S. Wingate, and
presented by him to the Natural History Museum in the
course of last year. The Diptera were represented by a
single species of Hristalis, not in sufficiently good condition
for determination ; and the Homoptera were represented by
some larvee belonging to the family Flatide. The dragonflies
and locusts were, however, of more interest, and of these a
list is given below, including the description of a locust
which appears to be new.
NEUROPTERA.
ODONATA.
Libellulida.
Neurothemis fulvia, Drury.
Insects from the Yang-tse-Kiang. 381
Sympetrum sanguineum, Miiller.
Crocothemis soror, Rambur.
Deielia phaon, De Selys.
Orthetrum brunneum, Fonscolombe.
Aischnide.
Hemianax ephippigerus, Burmeister.
Agrionide.
Neurobasis chinensis, Linné.
ORTHOPTERA.
Locustide.
Aularches miliaris, Linné. ‘Two rather small and brightly
coloured specimens.
Cyrtacanthacris succincta, Linné,
Cyrtacanthacris Wingatei, sp. n. (infra).
Trilophidia annulata, Vhunberg.
Cyrtacanthacris Wingatet, Kirby, sp. n.
Exp. al. 82 millim. ; long. corp. 42 millim.
Male.—¥ront of head and pronotum dark red; vertex,
cheeks, lower part of the face, and sides of the thorax much
paler. The space between the facial carine and the thorax
thickly rugose-punctate ; back of the head smooth, with two
rather indistinct rows of black spots running backwards from
between the eyes, and diverging behind. Thorax irregularly
mottled with black on the back and sides. Abdomen
shining chestnut; interalary space paler. Legs brownish
testaceous, four front femora spotted with black above.
Hind femora with all the caring set with small black serra-
tions, and with three black bands on the upper surface, the
third narrower than the others, and followed by three pairs of
black spots, the two hinder pairs more or less connected in
the middle. Hind tibiz red, with yellow spines tipped with
black. Tegmina testaceous brown, darker towards the base,
and mottled with black towards the base, and more sparingly
with brown beyond. Wings smoky hyaline, darker towards
the tips, yellowish brown along the costa nearly to the ex-
tremity, and broadly suffused with rosy at the base.
Allied to C. succincta, Linné.
One specimen.
382 On some Insects from the Yang-tse-Kiang.
I append a list of the butterflies drawn up by Dr. A. G.
Butler.
List of the Butterflies collected by Capt. Wingate at Hunan.
By A. G. BuTuer, Ph.D.
Fam. Nymphalide.
Subfam. EvpL@in.
Salatura plevippus, Linneeus. One example.
* Parantica aglea, Cramer. One example.
Trepsichrois mulciber, Cramer. One example.
Subfam. NYMPHALINZ.
Precis almana, Linn. Two examples.
enone, Linn. Five examples,
# orithyia, Linn. One female.
® laomedia, Linn. One example.
—— iphitu, Cramer. One example.
Pyrameis cardui, Linn. Two examples.
calliroé, Hiibn. One example.
Cethosia biblis, Drury. One example.
Neptis eurynome, Westwood. One example.
Subfam. SATYRINZ#.
*Tansima verma, Kollar. One example.
Lethe confusa, Aurivillius. One example.
rohria, Fabricius. One example.
Fam. Lycenidez.
Polyommatus beticus, Linn, One example.
Everes filicaudis, Pryer. Three examples.
Fam. Papilionide.
Subfam. Prerinz.
Delias aglaia, Linn. Two males, one female.
Terias anemone, Felder. One male.
Catopsilia pomona, Fabr. Two males, one female.
Ganoris melete, var. dulcinea, Butler. One male.
Subfam. PAPILIONINZ.
Papilio cloanthus, Westwood. One example.
asiaticus, Ménétriés. One example.
xuthulus, Bremer. One example.
Nevilli, Wood-Mason. One example.
chaon, Westwood. One example.
*.
K
Six examples (species marked above by an asterisk) have been retained
for the Museum collection.
On new Rodents from Western South America. 383
XLVII.—Descriptions of new Rodents from Western South
America. By OLDFIELD THOMAS.
Ctenomys opimus, Wagu.
Ctenomys opimus, Wagn. Arch. f. Nat. 1848, p. 75.
There can be little doubt that the specimen on which this
species was founded was one of the “ large number” obtained
by Mr. Bridges in Potosi and other parts of Bolivia, and
referred by Waterhouse to C. braziliensis *.
The Museum contains several specimens from the same
collector, and in addition a series of four from Tetiri, S. Peru,
others from Sahama, N.W. Bolivia, and an isolated example
from Jujuy; and these seem to represent three subspecies, of
which the Sahama one, as being like some of Bridges’s
specimens, and the most like Wagner’s description, may be
taken as the typical subspecies. The other two are as
follows :—
Ctenomys opimus nigriceps, subsp. n.
Similar in size to the typical form, or perhaps rather larger,
and the tail rather longer.
Fur similarly soft and fine. General colour greyish fawn,
more greyish and less sandy than in true C. opimus. Whole
centre of face as far back as the nape prominently deep black ;
on the neck the black becomes much more grizzled with
fawn, but is traceable as an ill-defined line down the back to
the loins. On each side there is also a black band passing
through the eye and across the ear, behind which it forms a
distinct patch, separated from the central black of the crown
by a projection forward of the body-colour. The sides of
the muzzle and the chin are also black, behind which latter
there is a contrasted transverse light mark corresponding in
position with the rami of the mandible. Behind this again
the centre of throat is prominently blackish. Upper surface
of hands, feet, and whole of tail dark chocolate-brown ; the
comb-like fingers of the feet and the longer hairs at the end
of the tail are, however, yellowish.
Skull as in the typical form.
Dimensions of the type (measured in the flesh by col-
lector) :—
Head and body 230 millim.; tail 100; hind foot (s. u.) 38,
(c. u.) 43.
* N. H. Mamm. ii. p. 273.
384 Mr. O. Thomas on new
Skull: greatest length in middle line 56; basilar length
47°8; zygomatic breadth 37; nasals 19x 8°55 interorbital
breadth 12°5; least breadth across brain-case 21; greatest
breadth on meatus 35; diastema 16; length of tooth-row
(alveoli) 11°6.
Hab. Tetiri, about 40 miles W. of Puno, on the Puno-
Moquegua road. Altitude 16,000 feet.
Type. Male. B.M. no. 97.10.8.42. Collected 5th July,
1896, by J. Kalinowski. Four specimens examined.
Besides Mr. Kalinowski’s four specimens, one of those
collected by Mr. Bridges, though much faded, shows some-
thing of the same marking and may, perhaps, be referable to
the same form. His other specimens agree better with
Herr Garlepp’s Sahaman examples of what may be considered
as the typical C. opimus.
Ctenomys opimus luteolus, subsp. n.
Varying from true C. opimus in exactly the opposite
direction to C. 0. nigriceps, owing to the suppression of all
the darker markings. General colour bright sandy buff all
over, the centre of the face, if anything, rather lighter than
the rest instead of being darker, the nasal region almost
white. No darker markings round eyes, at back of ears, or
on the chin. Under surface similar to upper, but rather
brighter, almost ochraceous buff; no white markings any-
where. Hands, feet, and tail buffy, the last-named rather
darker terminally.
Skull apparently as in the typical form.
Dimensions of the type (measured in skin, therefore only
approximate) :—
Head and body 220 millim.; tail 70; hind foot (s. u.) 36°5,
(c. u.) 41.
Skull: greatest length in middle line 51;. zygomatic
breadth 32°7; nasals 19-5 x 8:6; interorbital breadth 12°7 ;
least breadth across brain-case 19°5; diastema 15; length of
tooth-series (alveoli) 11:2.
Hab. Cordilleras of Jujuy, Argentine Republic.
Type. Male. B.M. no. 99, 2. 22.17. Presented by the
La Plata Museum.
This tuco-tuco may be readily distinguished from its allies
by its light-coloured muzzle, this part being brown or black
in all the more northern specimens.
Rodents from Western South America. 385
Ctenomys dorsalis, sp. n.
Size about as in Ct. talarum. Coloration very much as in
the large Ct. boliviensts, Waterh. Fur soft and_ fine.
General colour above shining buffy fawn, a marked black
dorsal line running from the tip of the nose backward on to
the rump; on the head this line is sharply defined and about
half an inch broad, but broadens out and is less defined on
the back. No dark lateral face-markings round eye or ear.
Cheeks and chin like body, but the usual light collar behind
them is well marked, running up on each side to the ear.
Throat, chest, outer edges of belly, and a narrow line down
its centre pale buffy, the hairs slaty basally; rest of belly
white, the hairs white to their bases. Upper surface of
hands and feet dirty whitish. Tail-hairs mixed black and
white.
Incisors about the breadth of those of Ct. talarum, orange
above and below.
Dimensions of the type (measured in skin, approximate) :—
Head and body 156 millim.; tail46; hind foot (s. u.) 27°2,
(c. u.) 30°2.
Breadth of two upper incisors 4°3.
Hab. Northern Chaco of Paraguay.
Type. Female. Original number 255. Collected 7th May,
1900.
7 Native name Sumkum.’”’ Presented by J. Graham Kerr,
sq.
The typical skin is unfortunately without a skull, but its
striking coloration will readily distinguish this handsome
little tuco-tuco from every species hitherto known except the
enormously larger Ct. boliviensis.
Sciurus versicolor, sp. n.
Closely allied to S. variabilis, Geoff., but with the belly
rich rufous.
Colour very variable, that of the specimen selected as the
type as follows:—Head and posterior back dark grizzled
olivaceous, the central area of the back darkened to black.
Nape, shoulders, and outer side of arms rich rufous, grizzled
along the centre line of the neck, deeper and purer on the
arms. A small patch of yellowish at the posterior base of
each ear. Outer side of hind limbs grizzled rufous. Whole
under surface and the inner sides of arms and legs deep vivid
orange-rufous. ‘Tail grizzled blackish olivaceous for its basal
fourth, then broadly washed with orange-yellow (more com-
monly bright rufous), its tip black.
386 On new Rodents from Western South America.
In another specimen from the same locality the whole
body is dull blackish rufous, nearly as rufous on the sides of
the rump as on the shoulders.
Skull not appreciably different from that of S. variabilis
from Santa Marta. Premolars }.
Dimensions of the type (measured in the flesh) :—
Head and body 230 millim.; tail 225; hind foot (s. u.) 58,
(c. u.) 63; ear 27.
Skull: greatest length 57; basilar length 43°3 ; greatest
breadth 33°5 ; nasals (diagonally) 18 x 8:8; palate length 25;
length of upper tooth-row 9°5.
Hab. (of type). Cachabi, Prov. Esmeraldas, N. Ecuador.
Alt. 160 m.
Type. Male. B.M.no.97.11.7.32. Original number 25.
Collected 3rd Jan., 1897, by Mr. W. R. Rosenberg.
The British Museum contains a large number of squirrels
from the north-western corner of South America which have
been referred to S. variabilis, Geoff. ; but now that Mr. Bangs
has shown that the true S. vartabilis is constantly white-
bellied (as is also the type of Gray’s S. Gerrardi), it is
evident that the red-bellied forms require a special name.
As is the case with S. variabilis, this species is a very
variable one, and each of the several series of itin the Museum
presents some striking variation among its members. The
general tone varies in depth and degree of grizzling: the
centre of the back varies from deep shining black to hardly
darker than the sides, the ear-patches are occasionally
suppressed, the middle part of the tail may be either yellow,
orange, or red, the end of the tail may be either red or black
(more frequently the latter), and even the distinctive red
belly may be picked out with white markings in the axille,
middle line, and groins. It is possible therefore that other
variations will lead into S. variabilis, in which case it will
have to be considered as a subspecies of that form.
Other localities from which the Museum has specimens of
S. versicolor are—in Ecuador, Paramba (Rosenberg, Miketta) ;
in Colombia— Valdivia, Lower Cauca (Pratt), Medellin
(Salmon), San Pablo (Hopke), and Bogota (Child).
A rather similar coloration to that of S. versicolor is found
in the true S. griseogena, Gray, of Venezuela; but that is a
smaller form, as shown by its skull, and has a more uniformly
grizzled back, less blackened behind and less reddened in
front.
Mr. Nelson’s Sciurus caucensis is not a member of this
group, but is allied to—indeed, I do not see how it is diffe-
rent from—Gray’s S. medellinensis, also from the Valley of
On a new Dasste from North Nyasaland. 387
the Cauca. Perhaps this again will prove to be a synonym
of S. Pucherani, Fitz. Neither species is referred to by
Mr. Nelson.
XLVITII.—A new Dassie from North Nyasaland.
By OLDFIELD THOMAS.
Procavia mima, sp. n.
General appearance, colour, and length of hair almost
exactly as in P. (Dendrohyrax) arborea, from which the
number of the mamme (1—2=6) widely separates it, and
allies it to the more typical members of the genus. Fur very
long, soft, and sleek, the hairs without the crinkling present
in arborea. General colour above pale sandy fawn, heavily
grizzled and lined with black, especially along the middle
line ; each hair dark brown for its basal 25-27 millim., then
sandy for about 5 millim., and its terminal 5 millim. shining
black. Crown and rump particularly black, owing to the very
numerous shining black ends to the longer hairs. Upper lip
and an inconspicuous spot over eye whitish. Ears rounded,
well-haired, greyish white. Dorsal spot pale buffy white,
the hairs this colour to their bases; peripheral hairs of spot
not deeper buffy, as is often the case in arborea. Under
surface white, with a faint buffy tinge, the bases of the hairs
blackish. Upper surface of hands and feet grizzled black
and pale buffy.
Dimensions of body apparently about the same as in P. ar-
borea. Hind foot 57 millim.
Hab. Nyasa-Tanganyika Plateau, Northern Nyasaland.
Type. Female. Skin without skull. B.M.no.99. 6. 28. 16.
Presented in 1898 by Col. Manning, then Acting Com-
missioner, British Central Africa.
This Dassie is so strikingly like P. arborea that it has
hitherto been regarded as belonging, or at least allied, to that
species. But now that I find that it has 1—2=6 mamma,
the formula of Procavia (s. s.) and Heterohyrax, as against
the 0—1=2 of Dendrohyrax, there can no longer be any
doubt that it represents a distinct species. In colour and
length of hair it is widely different from any known species
with the same mammary formula.
388 On a new Species of Ornithoptera.
XLIX.—Description of a new Species of Ornithoptera.
By H. Grose-Smiru, B.A., F.E.S., F.Z.S., &e.
Ornithoptera titan, sp. n.
3 .—Upperside. Anterior wings black, with a broad
yellowish-green band extending along the costa and partly
invading the cell, narrowest at the base, but becoming wider
in the middle and tapering towards the apex, which it does
not reach; on the disk from the base and thence below the
median nervure to the inner margin is a yellowish-green
triangular area, the apex of which is a little above the lowest
discoidal nervule ; this green area, between the median ner-
vules, does not extend to the median nervure, nor does it
reach the outer margin, which is broadly black; there is no
stigmatic brand asin the O. priamus group. Posterior wings
golden yellow, with a black outer border, which is narrow
along the costa and becomes wider at the apex ; the outer
edge of the yellow area is golden green, which colour extends
inwardly along the veins; beyond the yellow area, which
extends rather beyond the lowest median nervule, to the inner
margin the wings are jet-black, the abdominal fold, which is
bordered basally by golden scales, being very wide; on the
disk towards the apex in the yellow area between the veins
are three large oval golden-green patches, in the upper part of
each of which is a black spot, which represent the spots on
the underside.
Underside. Anterior wings golden green, the costal, outer,
and inner margins broadly black, the veins black; on the
disk between the veins is a row of four rather narrow black
lunules. -Posterior wings golden yellow, with a golden green
submarginal and inner-marginal area, the latter very wide ;
three large oval black spots on the disk, the uppermost sub-
apical, the two others on either side of the lowest subcostal
nervule ; outer margin narrowly black; abdominal fold pale
buff, with long hairs of the same colour.
In shape the anterior wings are very broad, less produced
at the apex than in the O. priamus group; posterior wings
convex and rather truncate at the anal angle. Antenne,
thorax, and legs black; abdomen yellow, with small black
spots on either side ; orbits white.
Expanse of wings 7? inches.
Hab. New Guinea.
In my own collection.
This magnificent butterfly was sent to me in a collection
Mr. A. Hempel on Brazilian Coccide. 389
stated to be from New Guinea (collector unknown to me).
The rest of the collection is in very bad condition. I con-
jecture, however, from the series of species contained in it
that the insects may have been captured towards the southern
end of British New Guinea. I hope to obtain further parti-
culars later on.
The absence of the stigmatic brand on the anterior wings
separates it from the group of O. priamus, and the shape
and neuration of the wings from the Schoénbergia group.
I have compared it with the drawing of the neuration of
Schoénbergia (?) goliath, Honrath, as figured by Mr. Rippon,
and find little, if any, difference; if the true locality of
S. goliath is New Guinea (Mr. Rothschild gives its locality
as Waigeu?, Dorey ?), it is very probable that my butterfly
is the male of that species.
A new genus should be created for this species.
L.— Descriptions of Brazilian Coccide.
By ApvoteH HemPe., 8. Paulo, Brazil.
THE writer has published, in the ‘ Revista do Museu Paulista,’
vol. iv., a paper on the Coccide of Brazil. One hundred and
thirty-one species are included in that paper, which is printed
in the Portuguese ]anguage; and under the circumstances it
was thought advisable to publish the descriptions of the new
species in the English language as well.
The measurements of the scales, body, antennz, and long
hairs are in millimetres ; those of the joints of the antenne
and legs, and of short hairs, spines, and glands, are in micro~
millimetres.
Coccide.
Subfamily Monopuresivz.
Genus Icery4, Signoret.
Icerya brasiliensis, Hempel.
Adult female elliptical, pink; antenne and legs dark
brown, entirely covered with white secretion ; consisting of
one long caudal tuft, one cephalic tuft, a lateral and a sub-
lateral row of nine tufts on each side, and a central longitu-
dinal mass. A tuft on each side of the caudal and cephalic
tufts is longer than the other marginal tufts. Ovisac large,
390 Mr. A. Hempel on Brazilian Coccide.
white, sometimes showing a creamy tinge, distal end curved
up. Beneath it is convex and slightly striated longitudinally.
Dorsum and side longitudinally fluted, with 14 or 15 longi-
tudinal furrows. In the largest individual examined the
anal tuft was 20°5 millim. long. The caudal and cephalic
tufts are usually fluted with four longitudinal ribs. The
ovisac has one or two longitudinal slits on the medial dorsal
line, through which the young escape. Forty-four eggs
were found in one sac.
Antenne usually 11-jointed: joint 5 is the shortest, and
sometimes unites with joint 4, making the antenne only
10-jointed ; joints 2, 4, 6-10 are subequal in length ; joint
11 equals or slightly exceeds joints 9 and 10 in length.
Length of antenne variable; the longest observed was
1:1 millim. long. Joints 1-10 each bear a whorl of about 6
hairs, while joint 11 has a terminal band of 15 or 16 hairs.
Legs ordinary ; tarsus curved near the distal end ; digitules
absent. Digitules of claw fine, hair-like, short.
Rostrum large, situated just between the first pair of legs.
Mentum with about 28 hairs. Rostral loop reaching beyond
the insertion of the second pair of legs. Numerous hairs are
scattered over both surfaces and around the margin; while
the body is terminated by two terminal tufts of 5 long hairs.
The entire dorsal surface is crowded with glands; these are
round and appear to be composed of 6 to 9 parts placed in a
circle, with a long glassy filament in the centre.
Length of insect and ovisac, excluding the tufts, 10°5
millim.
Larva (just hatched).—Elliptical, deep red with a pinkish
tinge. On the dorsum there are four tufts of yellowish wax,
forming a diamond-shaped patch, with the long diameter
placed cephalo-caudad. Antenne 6-jointed : joints 2-5 sub-
cylindrical and subequal in length, ‘066 millim. long ; joint 1
convex on the inner side; joint 6 club-shaped, *164 millim.
long; joints 2-5 have each one long slender hair and several
shorter ones ; joint 6 with 6 very long hairs (longer than the
antenne), ‘64 millim. long, and about 12 short ones.
The dorsum bears many slender hairs, placed in about ten
irregular longitudinal rows. On the head there are four
bristles between the eyes; the two middle ones are very long,
reaching nearly to the tip of the antenne. ‘There are 6 anal
bristles about 1°35 millim. long, equalling nearly twice the
length of the body. There are also 6 shorter bristles on each
side of the abdomen not one third the length of the anal
bristles. The lateral margin of thorax and head also bears
some short bristles. Dorsum with many round secretory
Mr. A. Hempel on Brazilian Coccide. 391
pores, placed more or less in transverse rows. Hyes 2, small,
conical, dark brown, nearly black. Antenne and legs also
dark brown. ‘Tibia of second and third pair of legs +20 millim.
long, tarsus and claw a little shorter; claw long, slender,
slightly curved, and notched at the end. Digitules of claw
slender, buttoned, a trifle longer than claw. No tarsal
digitules. Length °73 millim.
Female (third stage)—Body ovate; secretion or wax
usually light yellow, arranged in two lateral rows each with
about 10 tufts, two sublateral rows of 8 tufts each, one ter-
minal tuft at each end, and one median longitudinal row of
5 tufts. Antenne 9-jointed, joint 9 the longest. Legs
shorter than in the larve. Rostral loop reaching to the in-
sertion of the third pair of legs. Mentum with about a dozen
short hairs. Both surfaces of the body are covered with
hairs, those on the dorsum fewer and longer. The dorsal
surface also contains a large quantity of round secretory pores,
each situated above a group of five or six cells. These pores
have the same construction as those in the adult, and are
most abundant on the head and the margins of the body.
Hab. Sent from I[guape by Mr. E. Young, where it occurs
in such numbers on Codieum sp.? as to kill the plant. Also
found in Ypirauga and Sao Paulo on Ficus sp., rose, and
other cultivated plants.
It has killed a number of shade-trees in So Paulo, and is
apt to cause considerable damage to the parks. The indi-
viduals usually cluster on the undersides of the twigs and
branches in great numbers. Also occurs in large numbers on
Liriodendron tulipifera, L., Laurus camphora, L., and on a
species of palm. Many Hymenopterous parasites have been
bred from this species; but the parasites do little harm to the
insect, as the eggs are not affected, and hatch, although the
adult is full of parasites. A species of Coccinellid larva has
also been observed feeding on the growing insects.
Icerya Schrottky?, Hempel.
Adult female, massed together and all covered with a dense
white secretion, so that it is hard to distinguish individual
characters. Each insect, however, is covered with a dense
mass of long white filaments of secretion, which seem to
proceed from glands placed in two concentric rings on the
dorsum; all the filaments pointing backwards, and some
attaining a length of 30 millim. On the abdomen there are
two small patches of white secretion. The ovisac is secreted
under the abdomen, and consists of a dense mass of white
392 Mr. A. Hempel on Brazilian Coccide.
woolly secretion, very sticky, adhering to everything it
touches. Denuded of wax orange-yellow in colour; legs and
antenne dark brown. Body ovate, wider posteriorly than
anteriorly. The dorsum has two concentric rings of pits or
glands extending around it, dividing it into three areas. The
abdomen is transversely furrowed.
Length 7°50 millim. ; width 5 millim. ; height 3 millim.
Boiled in asolution of KOH it gives the solution a yellow
muddy appearance. The derm is thin and transparent.
Antenne variable, of 10 or 11 joints; 11, however, seems
to be the typical number of joints, of which the last is the
longest. Joints 1-10 each bear a whorl of 7-9 hairs ; joint
11 has a brush of many hairs. Length about 1:10 millim.
Approximate formula: 11 2 13 (789) 10 6 (45).
Length of joints: (1) 110, (2) 123, (3) 97, (4) 66, (5) 66,
(6) 75, (7) 93, (8) 93, (9) 93, (10) 84, (11) 173.
Legs long and hairy. Length of joints of first pair of
legs: coxa 191, trochanter and femur 594, tibia 604, tarsus
252, claw 66. Tarsal digitules wanting. Digitules of claw
short, hair-like. Eyes close to the base of the antenna,
small, conical, dark brown. Rostrum lar~s, situated between
the first pair of legs. Rostral loop extending to the second
pair of legs. Mentum with about 20 hairs. The dorsal and
ventral surface of the body is crowded with hairs and large
round glands; the hairs on the ventral surface, however, are
smaller than those on the dorsal surface.
Larva (just hatched).—Orange-red, elliptical, ‘812 millim.
long and ‘400 millim. wide. A very little white secretion on
the back. Antenne about *555 millim. long; of 6 joints,
the terminal joint longest and club-shaped. Length of joints:
(1) 57, (2) 70, (3) 79, (4) 79, (5) 79, (6) 191. All the joints
bear hairs ; joint 6 bears six very long hairs and many shorter
ones; joint 5 bears also one very long hair. LKyes small,
conical, dark brown. The six central caudal bristles are
very long, attaining a length of 1:46 millim. Besides these
there are six shorter bristles on each side, but these are
scarcely one fifth as long as the others. The margin of the
body and the derm also bears numerous hairs, none of them
very long. Many round glands are also present in the derm.
The legs are long and thin, with many hairs. Length of
joints of first pair of legs: coxa 79, trochanter and femur
222, tibia 244, tarsus 164, claw 40. Digitules of claw long,
slender, with slightly expanded ends. ‘Tarsal digitules
wanting. Claw slightly notched. Rostral loop short, ex-
tending a little beyond the third pair of legs.
Mr, A. Hempel on Brazilian Coccide. 393
Hab. Jundiahy, State of Sio Paulo, on the bark of an
indigenous tree. Collected by Mr. C. Schrottky.
Several hundred small Hymenopterous parasites were bred
from this species. As in J. brasiliensis, the parasites are
present in the adult, and do not prevent the eggs from
hatching, and consequently are but a slight check to this
species.
Subfamily Coccrraz.
Genus Errococcus, Targ.
Eriococcus brasiliensis, Ckll.
Adult female reddish brown, oval in outline. Anal ring
with 6 long hairs. Antenne variable. In some specimens
joint 3 is the longest, whilé in others joint 4 is the longest,
being 44 long; joint 1 is 22 long. All joints except joint 3
bear one or more hairs.
Male sacs of same consistency and colour as those of the
female, but a little smaller. The adult male is dark brown
in colour. Antenne variable, usually of 10 joints, but
sometimes with only’ or 9 joints: joints 2-9 are dilated
at the distal ends; joint 2 is very thick, being twice the
diameter of joint 3. Approximate formula; 10 2 (9 3) 87
(456) 1. All the joints bear many small hairs, while in
addition to these joints 8 and 9 each bear one, and joint 10
bears five large thick hairs. Thorax large; abdomen wide,
with several hairs on the margin of each segment. Genital
spike short and acuminate. Wings ordinary, the pocket, for
the insertion of the balancers, being large. Balancers or
halteres long; the last joint long and slender, with a large
hook at the distal end. Claws toothed as in the female.
Length °95 millim. ; extent 1°87 millim.
Hab. Ypirauga. Usually crowded on the ends of th
twigs of Baccharis dracunculifolia, DC.
The insect is active until just before gestation, when it
constructs a closely felted sac, occupying three or four days
for the work.
Eriococcus perplexus, Hempel.
Largest female sacs 11 millim. long, 3°5 millim. wide, and
1°75 millim. high; spindle-shaped, widest caudad of the
middle. Snowy white, closely felted, pointed, and with a
small opening at the posterior end. The dorsal surface is apt
to be slightly flattened and shows traces of transverse furrows.
? .—Orange-yellow, with a brown longitudinal median
Ann. d& Mag. N. Hist. Ser. 7. Vol. vi. 26
394 Mr, A. Hempel on Brazilian Coccide.
stripe. After boiling in a solution of KOH it is 4°5 millim.
long and 2°75 millim. wide. It colours the liquid light
yellow. Antenne variable, 7-jointed, about °30 millim. long ;
joints 1, 3, and 4 nearly equal in length; formula: 1 (8 4)
27 (5 6), varying to 3 (1 4) 27(65). The antenne are
large and are little reduced in size in the first four joints.
All joints except joint 3 bear hairs. Legs short and stout ;
cox with two hairs and three or four short spines; tro-
chanter with two terminal hairs and one spine; femur twice
as long as wide; tibia and tarsus equal in length, about
seven tenths as long as femur; claw long, curved; all the
digitules slender, with expanded ends. Anal ring with six
hairs. Mentum is situated in front of the insertion of the
first pair of legs ; rostral loop short, extending halfway to the
middle pair of legs. The entire surface of the body is
covered with straight and curved spines and minute round
glands. The abdomen ends in a pair of small tubercles.
Larva (just hatched).—Orange, pyriform; the abdomen ends
in a pair of tubercles, each terminated by a long bristle.
Between the tubercles there are two long and four shorter
hairs. The surface of the dorsum bears six longitudinal rows
of large sharp spines and numerous small tubercles. An-
tenn 6-jointed, joint 3 the longest. Legs small, claw long
and slender and slightly curved; digitules slender. Anal
ring with six hairs. ‘There are two conspicuous hairs on the
anterior margin between the antenne. yes small, spherical,
inconspicuous. Rostral loop long, reaching nearly to the
anal ring.
Hab. Ypirauga, State of SH Paulo, on the underside of
leaves of a plant of the order Myrtacee ; and State of Minas
Geraes, where it occurs on the bark of Eugenia jaboticaba.
Eriococcus armatus, Hempel.
Female sacs oval, flattened, with a large elliptical aperture
in the caudal end ; composed of a thick closely felted material.
White, with a creamy tinge. 3°25 millim. long and 2°25
millim. wide. Adult temale oval in outline; reddish brown ;
abdomen transversely wrinkled. Boiled in a solution of
KOH it colours the liquid a light red.
Antenne placed very close together, of seven joints, joint 7
the longest, variable, about *320 millim. long. Approximate
formula: 7 (1 2)463 5. All the joints bear hairs. Length
of segments: (1) 44, (2) 44, (8) 36, (4) 40, (5) 31, (6) 38,
(7) 89. Legs short; tibia and tarsus nearly equal to femur
and trochanter. Tarsal digitules slender, with knobbed ends,
Mr. A. Hempel on Brazilian Coccide. 395
extending to the tip of claw. Digitules of claw larger, with
expanded ends. Rostrum small, situated between the an-
tenne and first pair of legs. Mentum large, dimerous.
Rostral loop long. Eyes small, oval. Anal ring with six
hairs. Anal tubercles present, each terminating in a long
seta, and bearing several hairs and four or five short thick
spines. ‘The last five or six segments of the abdomen bear,
on the lateral margins and dorsum, several groups of these
short, thick, spear-shaped spines, each group consisting of
four or five spines. Scattered over both surfaces of the body
are small round spinnerets, short spear-shaped hairs, and
many short cylindrical glands. ‘These glands are especially
numerous on the lateral and caudal margins of the abdomen.
Length 2°70 millim. :
Larva.— Length *44 millim., oval. Antenne of six joints,
joint 6 the longest. Legs short, thick; digitules slender.
Anal ring with six hairs. Anal tubercles not conspicuous,
each ending in a seta, and bearing two short sharp spines.
The dorsum bears about sixteen transverse rows of short hairs.
Hab. Ypirauga, on Baccharis sp. The individuals are
clustered around the stem, close to the ground, on the roots,
and also on the ends of the branches.
Genus DactyLopius, Costa.
_ Dactylopius grandis, Hempel.
Adult female oval in outline; dorsum convex, rounded,
dark orange in colour. The dorsum is covered with a white
powdery secretion, arranged in one submedian and one sub-
lateral longitudinal row on each side. Around the lateral
margin there is a fringe of short white tufts. ‘There are also
two long acuminate anal tufts. Sometimes the secretion has
a yellowish tinge. ‘The adult rests upon a cushion of white
cottony matter that contains the young; this cotton readily
adheres to anything it touches. ‘The largest specimens are
7:50 millim. long, 5 millim. wide, and 3 millim. high.
Antenne of eight joints, joint 8 being the longest ; joint 1
thick, nearly twice the diameter of joint 2. Length of joints
variable, joints 3, 6, 5, and 7 being subequal and joints 1 and
2 being subequal; sometimes 1, sometimes 2, is the longer.
Approximate formula: 8 215 (367) 4. Average length
of antenna ‘48 millim. All the joints bear hairs. Length of
the joints: (1) 67, (2) 71, (8) 49, (4) 36, (5) 53, (6) 47,
(7) 49, (8) 98. Eyessmall, conical. Legs short, stout, with
few hairs; coxa wider than long; tarsus and tibia about
26*
396 Mr. A. Hempel on Brazilian Coccida.
equalling femur in length; claw small, with the digitules
short and slender, with buttoned ends. ‘Tarsal digitules
slender, scarcely reaching to the tip of the claw. Rostral
loop very short. Anal ring with six hairs. The two anal
tubercles are inconspicuous, but each one bears several hairs
and a number of small triangular glands and about 15 short,
thick, sharp spines. On the dorsal surface of the body near
the lateral margin there are about 32 groups of glands and
spines, each group consisting of from 8 to 12 small trian-
gular glands or spinnerets and from 5 to 8 short sharp spines.
The lateral margin is also fringed by a number of short
hairs. Dorsal surface of body bears many small triangular
glands and short sharp spines, placed singly, apparently in
transverse rows. ‘The ventral surface of body bears glands
and many small hairs.
Young (just hatched).—Elliptical, yellow; eyes small,
conical, dark brown. Antennee 6-jointed, joint 6 being the
longest, equalling joints 3, 4, and 5. Rostral loop long,
nearly extending to the anal ring. Legs long, claw slender,
digitules of claw and tarsus long and fine, buttoned. Anal
tubercles inconspicuous, each bearing a terminal seta.
Around the margin of the body there are short sharp spines,
while each of the last two abdominal segments bears two
spines on each side. Length 45 millim.
Hab. Ypivauga. On leaves and twigs of a plant of the
fam. Myrtacee.
Dactylopius setosus, Wempel.
Adult female elliptical, flat, orange-red in colour; legs and
antenne yellowish. Thorax and abdomen transversely
wrinkled; the abdomen ends in two stout sharp filaments of
white secretion, while both surfaces of the body are dusted
with a white powder. On the dorsum there is one median
row and on each side a sublateral and a marginal row of
long glassy filaments, which stand out in all directions and
"give the insect a bristly appearance. Largest specimens
5 millim. long and 2°75 millim. wide.
Antenne usually 8-jointed, although sometimes joints 3
and 4 are united into one; slender, joints 4~7 slightly ex-
panded at the distal end. All of the joints bear hairs. Joint
8 the longest. Approximate formula: 8 3 (2 1) 54 (67).
Antennz varying in length from ‘60 millim. to °70 millim.
Average length of antennal joints: (1) 89, (2) 89, (3) 102,
(4) 64, (5) 84, (6) 62, (7) 62, (8) 1383.
Legs long, slender, with many hairs. Coxa short and
Mr. A. Hempel on Brazilian Coccide. 397
wide. The joints of first pair of legs measure in w :—femur
333 long; tibia 312; tarsus and claw 125. Digitules of
tarsus slender, with small buttons at tip, reaching to tip of
claw, Digitules of claw large, with widely expanded tip.
Eyes small, conical. Lostral loop very short. Anal ring
_ with 6 hairs. Anal tubercles present, each ending in a long
seta, and bearing two short sharp spines and a number of small
hairs, and minute triangular glands. Grouped around the
anal orifice and arranged singly near the lateral margin of
the dorsal surface are some characteristic cylindrical glands,
each one 35 w long and 9 w wide. Three to five short hairs
are arranged around the external openings of these glands.
Lhe dorsal surface also bears many minute triangular spin-
nerets, and in the cephalic region many hairs. The ventral
surface also has hairs and glands scattered over it.
Hab. Sio Paulo. On the twigs of a species of Ficus
planted as shade-trees in some of the streets of the city.
Dactylopius secretus, Hempel.
Female active ; body ovate, transversely furrowed; very
light yellow; the dorsum dusted with a fine white powdery
secretion. ‘he lateral margin bears a fringe of small tufts
of white wax. A pair of these tufts, at the caudal extremity,
are longer than the others, and between them there is another
pair of fine hair-like tufts. The largest specimen was 2°25
millim. long and 1:25 millim. wide, but it was probably im-
mature. It inhabits globular or cylindrical galls. These
galls are formed by having part of the leaf thickened and
folded upon itself, with its long axis parallel to the long axis
of the leaf. The gall is on the underside of the leaf, with
the opening on the upperside, and reaches a length of
12 millim.
Antenne short, thick, of eight joints; each joint with
several thick hairs; joint 8 the longest. Length about
‘42 millim. Approximate formula: 8 2 1 3 (5 7) (4 6).
Length of the segments of an antenna: (1) 57, (2) 62, (3) 43,
(4) 85, (5) 40, (6) 35, (7) 40, (8) 98. Legs short. Joints
of first pair of legs: femur 191, tibia 182, tarsus and claw
102. Digitules of tarsus fine, slender, with ends slightly
expanded, not extending beyond the tip of claw. Digitules
of claw thick, end enlarged, extending beyond the tip.
Rostral loop long, reaching about halfway between the
second and third pair of legs. Eyes very small, oval. Anal
ring with six large hairs. Anal tubercles not conspicuous,
each ending in a large seta, and bearing two small sharp
398 Mr. G. K. Gude on new Species of
spines, smaller hairs, and small triangular and larger round
glands. The surfaces of the body bear hairs and scattered
spines and numerous small and large glands.
Adult male light yellow ; eyes black. Length, including
style, ‘85 millim., extent of wings 2°25 millim. Antenne
10-jointed ; joint 10 the longest; joints 3-9 subequal.
Halteres short, expanded in the middle, bristle fine, with
a large hook at the end. Legs long, slender, with numerous
hairs. ‘Tibia twice the length of tarsus. Claw very long
and slender, one third length of tarsus. Digitules hair-like,
short. Style very short, acuminate. The last segment of
the body bears on each side of the style one long hair and
several shorter ones. The other abdominal segments also
bear several short hairs on the lateral margins.
Hab, Ypirauga. In galls on leaves of a plant of the
family Solenacez.
But few of the galls contain insects, and it is probable that
they are made by other insects and appropriated by this
Dactylopius. This species is accompanied by an ant
( Cremastogaster ?),
[To be continued. |
LI.— Descriptions of new Species of Japanese Land-
Shells. By G. K. Gung, F.Z.8.
Amoneé a number of Helicoid land-shells received from
Mr. Hirase the following appear to be undeseribed :—
Arnouldia ceratodes, sp. n.
Shell imperforate, trochoid, smooth, shining, dark corneous ;
spire depressed, apex obtuse, suture impressed. Whorls 54,
increasing slowly, the last not descending in front, at first
keeled at the periphery, becoming rounded towards the
mouth, Aperture oblique, lunate; peristome thin, straight,
acute; margins distant, upper descending slightly, colu-
mellar margin slightly expanded above, the umbilical region
sunk.
Diam. maj. 5°5, minor 4°75; alt. 4 millim.
Hab. Kashima, prov. Harima. ‘Type in my collection.
The new species is allied to A. stenogyra, A. Ad., but it is
darker and considerably Jarger and more deeply sunk around
the umbilical region. Six specimens were received.
Japanese Land-Shells. 399
Arnouldia nanodes, sp. n.
Shell imperforate, depressed conoid ; under the microscope
seen to be very finely striated and decussated with close faint
spiral lines, the latter a little more distinct and wider apart
below ; thin, dull above, a little shining below, pellucid,
pale corneous. Spire depressed, apex obtuse, suture im-
pressed. Whorls 4, increasing slowly, the last angulated at
the periphery, tumid below near the mouth. Aperture
oblique, securiform ; peristome thin, straight, acute ; margins
approaching, the upper forming an obtuse angle with the
outer margin at the periphery; columellar margin vertical,
slightly expanded above.
Diam. 2°25, alt. 1°5 millim.
Hab. Kioto. Type in my collection.
The nearest ally of this form appears to be A. obtusangula,
Reinh., but that species is perforated and is composed of 54
whorls. Six specimens.
Crystallus sulcatus, sp. n.
Shell perforate, discoid, smooth above, very finely striated
below ; shining, pellucid, corneous. Spire depressed, suture
channelled. Whorls 4, rounded, increasing slowly at first,
the last widening rapidly, not descending in front, sunk round
the umbilicus. Aperture nearly vertical, roundly lunate ;
peristome thin, straight, acute; margins distant, upper
arcuate, columellar margin slightly dilated over the narrow
umbilicus.
Diam. maj. 4, minor 3°75; alt. 2 miflim.
Hab. Kioto. Type in my collection.
Two specimens only, one of which is imperfect. This
species is allied to C. microdiscus, Reinh.
Crystallus velatus, sp. n.
Shell narrowly perforate, discoid; under the microscope
seen to be faintly striated and decussated with excessively
fine spiral lines; shining, pellucid, corneous. Spire de-
pressed, suture impressed. Whorls 4, a little rounded, in-
creasing slowly, the last not descending in front, sunk round
the umbilicus. Aperture slightly oblique, narrowly lunate ;
peristome thin, straight, acute; margins distant, upper
arcuate, columellar margin slightly expanded over the narrow
umbilicus.
Diam, 3°5, alt. 1°5 millim.
Hab. Kioto. ‘Type in my collection.
400 On new Species of Japanese Land- Shells.
Allied to the preceding, but that shell is more elevated in
the spire, the aperture is more widened laterally, the umbilicus
is a little wider, and it is devoid of spiral sculpture.
Microcystis Hiraset, sp. n.
Shell perforate, depressed-conoid, very finely striated, de-
cussated with microscopic spiral lines; thin, shining, pel-
lucid, dark corneous, whitish below. Spire depressed, suture
linear, faintly margined. Whorls 43, a little rounded above
and below, increasing rapidly, the last twice as wide as the
penultimate, not descending in front. Aperture a_ little
oblique, lunate; peristome thin, acute; margins distant,
upper depressed, columellar margin a little dilated over the
narrow umbilicus.
Diam. maj. 9°75, minor 8; alt. 4°75 millim.
Hab. Kashima, prov. Harima. Type in my collection.
Three specimens received. This new species is interme-
diate between J/. rejecta, Pfr., and M. Doenitze’, Reinh.
Three other specimens (apparently immature) I also refer to
this form ; they measure barely 6 millim. in diameter.
Trishoplita cretacea, sp. n.
Shell deeply umbilicated, conoid, irregularly striated, in-
distinctly and minutely granulated; chalky white under a
deciduous thin yellowish-white cuticle. Spire elevated, apex
obtuse, suture impressed. Whorls 63, rounded, increasing
slowly, the last descending very shortly in front, obsoletely
angulated at the periphery, becoming rounded and a little
widened at the mouth. Aperture oblique, elongate-rotundate ;
peristome scarcely thickened, expanded all round, the margins
approaching, columellar margin broadly reflected over the
deep but rather narrow umbilicus.
Diam. maj. 15°5-17°5, minor 13°5-15°5; alt. 13 millim.
Hab. Inga, prov. Hoki. Type in my collection.
The nearest ally is 7. mesogonia, Pils., but that species is
much smaller and possesses only 5} whorls. The twelve
specimens received are all decorticated, but some have patches
of the cuticle adhering.
Plectotropis conica, sp. n.
Shell deeply and perspectively umbilicated, conical, finely
striated, the striz decussated by very fine wrinkled spiral
lines; from pale fuscous to reddish brown, paler around the
umbilicus; the thin deciduous cuticle is raised transversely
Lord Walsingham on Asiatic Tortricida. 401
into membranous lamelle, which are rather long and not
much interrupted at the peripheral region, but becoming
shorter above near the suture and below near the umbilicus.
At the periphery the cuticle is produced into conspicuous
triangular membranous scales. Spire elevated conoidal,
apex prominent, acute, suture impressed. Whorls 64-7,
increasing regularly, the earlier rounded, the later ones more
or less flattened above, the last rather tumid below near the
mouth, descending very shortly and slightly in front, acutely
keeled. Aperture oblique, subcircular; peristome thickened
and reflexed, pale rufous; margins approaching, the colu-
mellar a little reflected over the umbilicus, which is deep and
perspectively widened, exhibiting the larger portion of the
penultimate whorl.
Diam. maj. 18-19, minor 17-17°5; alt. 11-11°5 millim.
Hab. Izuhara, Tsu-shima. ‘Type in my collection.
The only known species to be compared with this form is
P. trochula, A. Ad.; but the latter is much smaller and more
depressed, the umbilicus is narrower, the cuticular lamelle
are much more interrupted, the spiral sculpture is much finer,
and the aperture less circular.
LII.—Asiatic Torrercrpz.
By the Rt. Hon. Lorp Watsinauam, M.A., LL.D., F.R.S.
[Continued from p. 341.]
HENDECANEURA, gen. nov.
(evdexa=eleven ; vedpov=a nerve.)
Type (¢ 2?) Hendecaneura impar, W|sm.
Antenne simple, or scarcely pubescent beneath. Palpi
slightly recurved, median joint roughly clothed above and
beneath, terroinal joint short, exposed; projecting to scarcely
more than the length of the head beyond it. Head slightly
tufted anteriorly. Thorax smooth. Fore wings, costa slightly
arched, ¢ with a narrow costal fold at the base, termen
slightly impressed below the rounded apex, tornus rounded,
dorsum very slightly convex. Neuration, g 11 veins (9
absent, coincident with 10 or 8): 9 12 veins all separate; 2
from middle third of cell; 3, 4, 5, and 6 more or less
assembled towards the termen; 7 to termen. Hind wings
402 Lord Walsingham on Astatic Tortricide.
broader than the fore wings, somewhat triangular, apex
slightly rounded, termen slightly indented above vein 5.
Neuration, 8 veins; 3 and 4 stalked; 5 moderately straight,
remote from 4; 6 and 7 nearly coincident along their base,
diverging at about one-third their length. Legs smooth,
hind tibia with a slight hair-pencil above.
The absence of one of the radial veins, presumably 9 by
coincidence with either 10 or 8, and the possession of a
costal fold enables this genus to be readily identified from
the male.
(1). Hendecaneura impar, sp. n.
Antenne ochreous. FPulpi ochreous, touched with fuscous
externally at the end of the median joint. Head ochreous,
touched with fuscous above the short, square, ochreous frontal
tuft, face whitish. Thorax ochreous, mixed with greyish
fuscous. Fore wings dark purplish fuscous, apex ferruginous ;
with an ochreous spot at the extreme base and a conspicuous
shining white streak on the middle of the dorsum, much
widened at its upper end beneath the fold ; near the base is a
reduplicated shining steel-blue patch on either side of the
fold, not reaching to the margins, a similar patch beneath the
costa a little before the middle, with two whitish ochreous
streaks above it on the costa; beyond the middle the costa
has seven oblique whitish ochreous streaks, the spaces be-
tween them being purplish fuscous; from the first pair of
these streaks an outwardly oblique shining steel-blue line is
angulated at the upper angle of the cell, and much widened
towards the tornus, but turns outwards towards the termen
before reaching it; the fourth of the costal streaks is also
tipped with shining steel-blue, and the fifth sends out a
shining steel-blue streak which nearly reaches the incision
below the apex ; below this is another slightly oblique steel-
blue streak running nearly to the termen at one-third above
the tornus, and enclosing between it and the wider steel-blue
band behind it an ocelloid spot or space, crossed by six or
seven transverse black lines, this and the whole apical space
above it is rich ferruginous, and some ferruginous spots are
scattered about the tornus and above the outer third of the
fold; cilia steel-grey, with a dark line along their base.
Exp. al. 15-18 mm. Hind wings dark brownish fuscous ;
cilia cream-white, shading to greyish fuscous at the flexus,
with a brownish fuscous line running through them near
Lord Walsingham on Asiatic Tortricide. 403
their base. Abdomen brownish fuscous. Legs whitish
ochreous, hind tarsal joints spotted with brownish fuscous.
Type, 3 (70237); ¢ (70236) Mus. Wlsm.
Hab, JAPAN (Pryer, 1886). Four specimens.
(2). Hendecaneura (?) cervinum, sp. n.
Antenne, palpi, and head pale fawn-brown. Thorax
blackish; tegule pale fawn-brown. ore wings blackish ;
a short pale fawn-brown basal patch, with some blackish
scales on the dorsum, is bounded outwardly by a trans-
verse bright steel-blue band, which is slightly indented upon
the fold; at one-third from the base is another bright
steel-blue band with an excrescence on its outer side above
the dorsum ; at its upper extremity are two whitish costal
dots; on the middle of the costa commences a series of
oblique whitish costal streaks (seven or eight), from the first
two a slightly interrupted oblique steel-blue band passes to
the tornus, widened towards its lower extremity; the space
beyond this band is brownish ochreous or olive-brown, trans-
versely and slenderly striated with black and containing a
bright steel-blue spot opposite the middle of the termen;
two smaller spots above it, placed obliquely backwards,
sometimes combine, forming a slender streak to the costal
streak fifth from the apex; cilia shining steel-grey, tipped
with fuscous near their apex and with a slender dark line
near their base. Hep. al. 14 mm. Hind wings dark
brownish fuscous; cilia pale greyish, with a dark line
running through them near their base. Abdomen dark
brownish fuscous. Legs greyish fuscous, hind tarsal joints
spotted with pale ochreous.
Type, 2 (70023) Mus. Wlsm.
Hab. JAPAN— Yzsso (Pryer, 1882). Three specimens.
The male is at present unknown, but this species so greatly
resembles ¢mpar, Wlsm., that there can be little doubt it is
correctly referred to the genus Hendecaneura.
(3). Hendecaneura apicipictum, sp. n.
Antenne pale cinereous. Palpt and head fawn-ochreous,
tinged with cinereous. Thorax greyish fuscous. Fore wings
dark reddish fuscous, mottled and streaked with bright
shining steel-blue; on a not otherwise defined short basal
patch are two or three bright steel-blue spots, a pair of
whitish costal streaks above the upper one ; before the middle
404 Lord Walsingham on Asiatic Tortricidae.
of the wing is an oblique elongate steel-blue spot beneath the
costa, with two white costal streaks at its upper edge, opposite
to which is a rounded steel-blue spot between the fold and
the dorsum ; this is followed by a smaller spot at the lower
edge of the fold halfway to the tornus ; from a little beyond
the middle of the costa a slightly outward-curved bright steel-
blue band runs to a little above the tornus, where it is
diverted towards the termen; at its costal extremity are two
oblique white costal streaks, these are followed by three pairs
of similar costal streaks before the apex, the third from the
apex sending out an oblique steel-blue line to the termen;
some steel-blue spots follow the outline of the termen below
it, and the space preceding these latter markings is streaked
with olive-brown; cilia pale steel-grey, a slender whitish
ochreous line along their base followed by a narrow fuscous
line; the extreme apex of the wing around the last two
costal streaks is tinged with olive-brown and the cilia at the
extreme apex are whitish. Exp. al. 18-14 mm. Hind
wings dark brownish fuscous; cilia white, tipped with
greyish before the apex, a slender whitish ochreous line along
their base, followed by a narrow brownish fuscous line.
Abdomen brownish fuscous.. Legs brownish grey; hind
tarsal joints spotted with whitish ochreous.
Type, 2 (70020); 3g (70019) Mus. Wlsm.
Hab. JAPAN—Yasso (Pryer, 1882). Four specimens.
Tutop1A, Hb.
1110 (2). Thiodia infessana, W\|sm.
Antenne pale fawn-ochreous. Palpi projecting more than
the length of the head beyond it; pale ochreous, the end of
the tuft beneath the median joint fawn-brownish. Head and
thorax pale ochreous, the latter with a brownish shade.
Fore wings fawn-brown, mixed with minute brownish fuscous
scales, profusely mottled and transversely strigulate with
shining pale ochreous, and with two short black lines in the
obscure ocelloid patch above the tornus ; cilia pale ochreous,
with brownish mottling. zp. al., ¢ 20, 2 22mm. Hind
wings brownish grey; cilia pale brownish cinereous, with a
slender pale ochreous line along their base. Abdomen and
legs brownish grey.
Type, & (60827); ¢? (60828) Mus. Wlsm.
Hab. Asiatic ‘luRKEY—H/azren—Shar Devesy, 15 VI,
1890 (Native Coll); Parustinz (Tristram, 1883). Six
specimens.
Lord Walsingham on Aséatic ‘Tortricide. 405
This species much resembles Zucosma fessana, Mn., from
which it differs in the absence of the costal fold, in its much
larger size, and in the colour being rather rich fawn-brown
instead of brown-grey, and in the numerous transverse
strigule ochreous instead of white.
1112. Thiodia citrana, Hb.
Grapholitha (Semasia) citrana, Star. & Wk. Cat. Lp. Eur. 256.
No. 1112 (1871) +; Stgr. Hor. Soc. Ent. Ross. XIII. 258 (1879) ?.
Hab. Kurope’. Asiatic TurKEY—Hazes—Shar De-
vesy, 18 VI. 1890 (Native Coll.) ; Kuvpavenpix1ar —
Brussa, VI.*; Szvyas—Amasia’.
1118. Thiodia Metzneriana, Tr.
Grapholitha (Semasia) Metzneriana, Stgr. & Wk. Cat. Lp. Eur. 256.
No. 1118 (1871) ?.
Hab. Europe’. N. PerstA—Asterabad (Lederer), Zell.
Coll. Srperta*. AMuUR— Pompejefka, 16 VII. 1876
(Christoph). CoreA—Gensan, VI. 1886 (Leech). CuHina
—Chang Yang, 4000-6000 feet (Pratt). Japan (Pryer,
1886 ; Leech, 1892)—Yusso (Pryer, 1882).
1121 (2). Thiodia intacta, sp. n.
Antenne yellowish white, annulated with fuscous. Palpi
porrect, brush-like, terminal joint concealed ; whitish, shaded
at the sides externally with brownish grey. Head yellowish
white. Thorax pale brownish ochreous. ore wings
yellowish white, with a slight shade of pale brownish ochreous
near the base, and with some very indistinct mottlings of the
same colour about the dorsum; a small group of black scales
lies at the upper edge of the ocelloid patch, which is shining
white; cilia yellowish white. ap. al.19mm. Hind wings
brown ; cilia white, with a pale brown shade along their
base. Abdomen brownish, anal tuft white. Legs whitish,
hind tarsal joints faintly shaded with brown.
Type, & (70172) Mus. Wlsm.
Hab. Japan (Pryer, 1886). Unique.
1123 (1). Thiodia rigidana, Snell.
Grapholitha (Semasia) rigidana, Snell. Tijd. vy. Ent. XXVI. 203-5,
Pl. XIL 4, 4a (1883) *.
Hab. E. SipertA—Askold Id." Corza—Gensan, VI.
1886 (Leech). JAPAN — Yusso (Pryer, 1882); Kivsrv
(Leech).
406 Lord Walsingham on Asiatic Tortricide.
1123 (2). Thiodia glebana, Snell.
Grapholitha (Semasia) glebana, Snell. Tijd. v. Ent. XXVI. 206-8,
Pl. XII. 6, 6a (1883) }.
Hab, EK. S1per14—Pokrowsk, 29 V.'; Baitonowa,6-9 VI.';
Tschernajewa, 11 VI.'; StarikowoR., 23 VI. 1876 (Christoph).
1123 (8). Thiodia sinensis, sp. n.
Antenne, palpi, and head dusky cinereous. Thorax
brownish olivaceous. Fore wings olive-brown, paler towards
the costa, along which is a series of short, slightly oblique,
reduplicate whitish streaklets alternating with dark brown ;
these are of about equal length throughout beyond the
middle, but shorter before it where they commence at about
one-fourth from the base; a small white ocelloid patch con-
taining two short black streaks lies opposite to the lower
half of the termen, with a dark brown shade-patch above and
below it; cilia dusky cinereous, with fuscous speckling.
Exp.al.18mm. Hind wings dark greyish brown ; cilia pale
cinereous, with a shade-line near their base. Abdomen
greyish brown. Legs cinereous, tarsi fuscous, annulate with
whitish,
Type, & (60151) Mus. Wlsm.
Hab. Cuina—Cuz-x1avea—Ningpo, IV. 1886 (Leech).
Unique.
Allied to asptdiscana, Hb., glebana, Snell., and corculana,
Z., but distinguished by the absence of any lengthening of
the costal streaks before the apex.
1219 (2). Thiodia niveicaput, sp. n.
Antenne simple in the male; yellowish white. Palpz very
thickly clothed, with a long hairy brush beneath completely
concealing the terminal joint; white. Head and thorax white,
the latter shaded with brown posteriorly. Fore wings white,
obliquely blotched and striated with brown along the dorsal
half of the wing and with a series of oblique brown costal
streaklets of varying lengths; these and the dorsal patches
contain also some blackish scales ; the first dorsal patch is at
about one-third the wing-length, and reaches a little way
across the fold, the second and larger dorsal patch is at two-
thirds, the space between them more or less occupied by
short waved streaks of the same colour; beyond the outer
patch is a white ocelloid spot, margined at its imner and
Lord Walsingham on Aszatic Tortricidee. 407
outer edges with shining silvery white, and containing one or
two transverse black streaks; above it the apical portion of
the wing is shaded with brown and greyish, and there are
some silvery lines between the costal strie ; cilia white, shaded
with brown and greyish towards the apex. [In some varieties
the brown patches are considerably shaded with greyish or
greyish fuscous, and the costal streaks are also rather fuscous
than brown.] ap. al. 14-15 mm. Hind wings pale
brownish grey; cilia shining white. Abdomen brownish
rey. Legs whitish.
Type, 8 (70097); ¢ (70098) Mus. Wlsm.
Hab. Japan (Pryer, 1886). ‘Ten specimens.
HuceE.is, Hb.
1124 (3). Hucelis (?) falcana, sp. n.
Antenne fawn-grey. alpi triangular, much clothed
above, porrect ; fawn-grey. Headfawn-grey. Thorax fawn-
brown. Fore wings with the costa slightly arched, the apex
somewhat falcate; fawn-brown, very slightly darker along
the basal half of the dorsum, and becoming somewhat
_ suffused with fawn-grey toward the termen, before which a
series of small fuscous spots is very indistinctly visible ;
along the costa also are some very indistinct, short, darker
fawn-brown streaks ; cilia chestnut-brown along their base,
pale fawn brownish beyond it. Hap. al. 18 mm. Hind
wings rather dark grey ; cilia slightly paler, with a slender
fawn-brown line running along their base. Abdomen
brownish grey. Legs pale greyish cinereous.
Type, 2 (70560) Mus. Wlsm.
Hab. Japan (Pryer, 1886). Two specimens.
A second female (70593) tends to show slender dark lines
along the veins, especially beyond the cell. Probably
correctly referred to Hucelis, Hb., but the male is at present
unknown.
1124 (4). Eucelis ochreocervina, sp. n.
Antenne cinereous fawn, faintly annulated. Palpi trian-
gular, much extended, projecting twice the length of the head
beyond it, terminal joint almost concealed; greyish fuscous.
Head greyish fuscous. Thorax cinereous fawn. Fore wings
elongate, rather narrow at the base, widened outwardly, costa
gently arched, male without a fold; apex acute, termen
408 Lord Walsingham on Asiatic Tortricide.
slightly impressed below the apex, tornus rounded; pale
fawn-ochreous, shaded with greyish fuscous and fawn-brown,
reticulated with whitish grey along the extreme costa, the
intermediate spaces being greyish fuscous; the markings
consist of an ill-defined basal patch, much wider below the
fold than above it, mixed greyish fuscous and whitish grey ;
a subcostal shade beyond the middle of the same colour, the
space below it tinged with fawn-brown, and containing at the
outer end of the cell a greyish fuscous spot, preceded by some
greyish white scales; near the termen are a few groups of
black scales; cilia fawn-brown, a slightly darker line along
their base, touched with greyish fuscous at the tornus. Hap.
al. 20mm. Hind wings pale greyish fuscous; cilia pale
cinereous, a slightly darker shade near their base. Abdomen
pale greyish fuscous, anal tuft fawn-ochreous. Legs greyish,
posterior tarsal joints pale spotted.
Type, 3 (60137, Goorais) Mus. Wlsm.
Hab. KasuMir—Goorais Valley, 7000 feet, VIII. 1887
(Leech). JAPAN (Pryer, 1886). Five specimens.
In some of the Japanese specimens the costal shade is
almost absent and the basal patch is reduced to a deep shade
along the dorsum.
1125 (1). Hucelis (?) divisa, sp. n.
Antenne ferruginous at the base, shading to brownish
grey beyond. Falpt projecting the length of the head
beyond it, median joint thickly clothed above, terminal joint
short, depressed, scarcely exposed; pale ferruginous. Head
pale ferruginous, somewhat roughly clothed above. Thorax
dark ferruginous. ore wings narrow at the base, widened
outwardly, costa very slightly arched, termen slightly oblique,
scarcely impressed, tornus rounded ; dark terruginous to two-
thirds from the base, the outer edge of the dark portion
slightly oblique, further from the base on the dorsum than on
the costa; beyond it the apical portion of the wing is
brownish fawn-colour, transversely striated with shining
steel-grey, with a few lines of ferruginous scales, especially
along the termen; on the costa the brownish fawn-colour
contains four pairs of pale ochreous geminated streaks, sepa-_
rated and divided by dark ferruginous; a dark ferruginous
spot lies within the extreme apex; cilia shining fawn-grey,
with a dark line running through them near their base. xp.
al. 18mm. Hind wings brownish fuscous ; cilia pale cine-
reous, with a dark line running through them near their
base. Abdomen [missing]. Legs whitish cinereous, shaded
On a new Snake from Burma, 409
externally with greyish fuscous, hind tarsal joints faintly
spotted. ~
Type, 2 (70095) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886). Unique.
A small and very distinct species, of which, unfortunately,
I have only a single female.
(To be continued. |
LIIL.—Description of a new Snake of the Genus Ablabes
from Burma. By G. A. BouLencer, F.R.S.
Ablabes Hamptoni.
Snout short, convex, profile curved from the frontal region
to the lip; eye three fourths the length of the snout; rostral
once and a half as broad as deep, just visible from above ;
nasal divided ; internasals a little longer than broad, a little
shorter than the prefrontals; frontal once and two thirds as
long as broad, longer than its distance from the end of the
snout, shorter than the parietals; loreal small, longer than
deep; a large preocular, with a second very small shield
below it; two postoculars; temporals 1+2; eight upper
labials, fourth and fifth entering the eye; four lower labials
in contact with the anterior chin-shields, which are much
longer than the posterior. Scales smooth, in15 rows. Ven-
trals 194; anal undivided ; subcaudals 76. Uniform greenish
grey above (in spirit), white beneath and on the upper lip.
Total length 1050 millim. ; tail 220.
This species, which is closely allied to A. Dorie, Bler.,
differing in the broader rostral shield, was discovered by
Mr. Herbert Hampton at Magok, on the Irawaddi, about
12 miles north of Mandalay. ‘Together with the snake
which I have the pleasure of naming, Mr. Hampton has
presented to the British Museum examples of the following
reptiles obtained by him at the same place, which are inter-
esting from the point of view of the geographical distri-
bution :—Gymnodactylus khasiensis, Jerdon; Trirhinopholis
nuchalis, Blgr.; Lycodon fasciatus, Aud, ; Simotes violaceus,
var. multifasciatus, Jan; Nata tripudians, var. ceca, Gmel,
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 27
410 Mr. P. Cameron on new
LIV.—Descriptions of new Genera and Species of
Hymenoptera. By P. CAMERON.
A. New Species of Vespide from India and Japan.
Polistes Rothney?, sp. n.
Ferrugineus, nigro maculatus; facie, clypeo apicequetarsorum flayis ;
alis hyalinis, nervis stigmateque flavis. ¢.
Long. 20 mm.
Hab. Barrackpore, Bengal (Rothney).
Antenne rufous, the scape yellow beneath; the basal six
joints black above; the third joint fully longer than the
succeeding two united ; the terminal joint black except at the
base, compressed, rounded and slightly dilated at the apex.
Head distinctly broader than the mesothorax ; in front densely
covered with silvery pubescence, granular; the lower part of
the front, the face, and clypeus yellow; the occiput and a
broad band across the vertex, extending to the eyes, black.
Clypeus broadly roundly projecting in the middle, the sides
waved. Mandibles rufous, tinged with yellow; the three
teeth are black, the middle is slightly smaller. Thorax
rufous, the lower parts of the pleurae with a yellowish tinge ;
mesonotum black, the middle, especially at the base, with two
broad irregular rufous marks. Scutellum and postscutellum
rufous, the two separated by a black line. Metanotum
black, the sides and two somewhat pyriform marks in the
middle rufous; it is not very strongly or closely transversely
striated. Mesopleure sparsely and indistinctly punctured ;
the base narrowly, the furrow, a broad irregular splash be-
hind on the upper half, and with a small rufous mark in the
middle, the apex more broadly than the base; the base, the
lower side, the part along the furrow and the apex, except for
a large mark in the middle, black—or, in other words, the
mesopleure are black, with an upper and two lower rufous
marks, the posterior being much the larger. Wings hyaline,
slightly suffused with yellow along the eyes. Legs rufous,
the four anterior yellowish in front, the four hinder coxe,
femora, and tibie lined with black above; the basal two thirds
of the metatarsus black. ‘The base of the first and second
abdominal segments and the apices of the others, except the
last, broadly black.
A distinct species. The form of the terminal joint of the
antenne is as in P. schach; that species differs from it in
Genera and Species of Hymenoptera. 411
having the eyes “ very small, separated from the base of the
mandibles by more than two thirds of their own length,”
they being here separated by one third only or by the length
of the three mandibular teeth ; in schach there are only two
teeth, and it bears no black markings. Saussure (‘ Vespides,’
i. p. 104) says that the mandibles of the male are “ trés fortes,
longues et arquées,.... leur base trés épaisse, armée d’une
forte dent dirigée en dedans,” which is certainly not the case
in our species, in which they are broad at the base, slightly
obliquely narrowed towards the apex. ‘The clypeus is quite
flat, not convex ; the sides at the top are broadly rounded ;
the centre is transverse; the occiput is sharply margined.
Polistes gallicus, Fabr.
This common European species may be added to the
Indian fauna. I have seen an example from Gilgit. It
belongs to the European form, not to the Eastern Asian var.
chinensis, Fabr.
Polistes rufolineatus, sp. n.
Ferrugineus; vertice, meso- metathoraceque nigris, lineis 2 mesonoti,
scutello postscutelloque rufis; pedibus rufis, coxis, trochanter-
ibus, femorum anterioribus basi, femoribus tibiisque posticis
nigris ; tarsis flavis, basi nigra; alis fulvo-hyalinis.
Long., 2 23, ¢ 15 mm.
Hab. Khasia Hills. Coll. Rothney.
Head ferruginous; the ocellar region from shortly behind
the posterior pair, from eye to eye, and extending along the
upper part of the eye-incision, black; the front and vertex
punctured, but not very strongly or closely ; the antennal keel
stout, roundly dilated behind and carinate in the middle.
Clypeus broader than long, roundly convex, the top trans-
verse, the angles oblique; the apex with the sides oblique ;
the punctures are sparse and moderately large. Mandibles
rufous, sparsely punctured; the teeth black, the middle one
smaller than the others. Antenne rufous; the five basal
joints of the flagellum black above. Thorax black ; the pro-
thorax, two lines on the centre of the mesonotum, a much
shorter one opposite the tegule, a mark, narrowed below,
under the tubercles, the scutellum and postscutellum, rufous.
Mesonotum almost impunctate ; the scutellum with shallow
scattered punctures. Metanotum, except at the base, irregu-
larly and rather widely transversely striated. Pleure bearing
shallow punctures ; the apical half of the metapleure obliquely
27*
412 Mr. P. Cameron on new
striated. Legs black; the greater part of the four anterior
femora, the anterior tibiz, the apical third of the middle, and
the apex of the third rufous ; the four posterior tarsi yellowish ;
the basal joints black, except at the apex. Wings fulvous,
with a violaceous tinge; the stigma rufo-testaceous; the
second cubital cellule at the top is as long as the space
bounded by the first transverse cubital and the first recurrent
nervures; the second and third transverse cubital nervures are
parallel and roundly curved outwardly from shortly above the
middle. Abdomen dark rufous. The base of the petiole
and a narrow line on the base of the second segment black.
The male has the face, clypeus, and mandibles paler, of a
more yellowish hue; the clypeusis longer than broad, almost
impunctate ; the apex is bluntly rounded, the lateral angles
above are more rounded ; the antennal tubercle is narrowly
furrowed in the middle above ; the mesopleure rufous, with
three irregular marks above and in the middle behind, the
latter being the larger and narrowed in the middle; the four
anterior coxe are only black at the apices, the posterior are
irregularly marked with yellow behind; the black on the
vertex is more extended and is united in the middle by a
narrow line to the black on the occiput.
Polistes rugifrons, sp. n.
Ferrugineus; meso- metathoraceque nigris; pedibus rufis ; coxis,
trochanteribus, femoribus tibiisque posticis nigris; alis fulvo-
fumatis. @.
Long. 25 mm.
Hab. Khasia Hills. Coll. Rothney.
Antenne rufous, the third and following joints black above.
Head rufous, the ocellar region and the occiput black; the
inner orbits of a paler colour; the vertex, the eye-incision,
and the centre of the front strongly and distinctly punctured ;
the space between the antenne bluntly tuberculate. Hyes
large, separated by, if anything, slightly more than the length
of the last joint of the antenne from the base of the mandibles.
Clypeus roundly convex, strongly but not very closely punc-
tured. The base of the mandibles with scattered punctures,
their three apical teeth are of equal size. Thorax black,
except the prothorax and the sides of the scutellum, which are
ferruginous; the pronotum is lined with yellow on the top.
Mesonotum closely and strongly rugosely punctured, the
punctures running into reticulations in the middle behind.
‘The scutellum and postscutellum are more rugosely punctured
than the mesonotum. The median segment is transversely
Genera and Species of Hymenoptera. 413
but not closely striated, and has a wide, moderately deep
furrow down the middle. Pro- and mesopleure closely
rugosely punctured; the basal half of the metapleure is
covered sparsely with shallow round punctures ; the lower
half has them much more widely separated. Legs rufous ;
all the cox and trochanters, the hinder femora, and the
hinder tibie except at the apex are black. Wings dark
fuscous-fulvous, iridescent; the stigma testaceous; the ner-
vures blackish ; the second cubital cellule is equal in length
to the space bounded by the first transverse cubital and the
second recurrent nervures. Abdomen dark ferruginous, the
base of the segments darker, the apex of the first and second
with a broad dull yellowish band.
Polistes khasianus, sp. un.
Ferrugineus, lineis pronoti abdominisque flavis ; mesosterno femori-
busque posticis subtus nigris; tarsis posticis flavis; alis fulvo
fumatis, stigmate fulvo. 9.
Long. 14 mm.
Hab. Khasia Hills. Coll. Rothney.
Antenne rufous, the flagellum darker above; the third
joint is slightly but distinctly longer than the following two
united. Head smooth, the front and vertex covered with a
silky down, the clypeus sparsely with black hairs and with a
few punctures ; above it is distinctly incised, roundly in the
middle ; the apex is triangularly produced, the middle point
rounded. Mandibles rufous, sparsely punctured in the middle;
the three teeth are sharply pointed and of almost equal size.
Thorax smooth, the mesopleure closely punctured, the punc-
tures deep and distinct; metapleure obscurely striated, the
base indistinctly crenulated. The mesosternum has the apex
and the middle at the base black, as are also the edges of
the divisions and the central furrow on the metanotum.
Legs coloured like the thorax, the hinder femora lined with
black on the outer half above; the apical four joints of the
metatarsus yellow. Abdomen smooth and shining, silky:
the petiole is black at the base, it and the other segments are
narrowly lined at the apex with pale yellow.
A distinct species. Characteristic is the form of the clypeus,
which is longer than usual compared to its breadth, it being
much longer than broad, and is more largely developed below
the eyes than usual.
Polistes nigritarsis, sp. n.
Ferrugineus ; abdomine flavo-lineato, dimidio apicali nigro; metanoto
414 Mr. P. Cameron on new
flayo bilineato; tarsis posticis nigris; alis hyalinis, apice fu-
mato. o.
Long. 15 mm.
Hab. Barrackpore, Bengal (Rothney).
Antenne rufous, darker towards the apex, the last joint
one half longer than the penultimate. Head densely covered
with pubescence, which is very dense and silvery on the
clypeus, darker and longer on the top; the occiput is black,
the outer orbits and the eye-incision narrowly lined with
yellow. The clypeus is twice longer than broad, is slightly
narrowed towards the top, where it is slightly and roundly
incised; the antennal tubercle is not very convex, broad,
slightly and gradually dilated towards the apex, which is
rounded. The eyes are separated from the base of the man-
dibles by the length of the second antennal joint. The lower
and upper teeth of the mandibles are large and sharply
pointed, the middle one shorter and narrower. Thorax
densely covered with a silky fulvous pubescence; the base of
the mesonotum—the mark dilated in the middle behind—a
transverse narrow line on the apex, the extreme base of the
median segment, the central furrow (narrow at the top,
broader below), and a mark (narrowed gradually behind on
either side at the apex) black ; a narrow line on the pronotum,
a narrow line on the base of the scutellums, spots at their
sides, and two large marks on the median segment (straight
on the inner side, rounded on the outer, and narrowed gradually
towards the apex) yellow. The pro- and mesothorax are
granular, the sides more coarsely than the top; the median
segment obscurely transversely striated; the metapleure
obliquely striated above, below the furrow sparsely punctured.
The hinder tarsi are entirely black, the middle black, white
at the base, the anterior yellowish white; calcaria white.
Wings hyaline, the stigma fulvous, the nervures darker;
there is a fuscous cloud occupying the greater part of the
radial cellule and the upper part of the apical cubital cellules ;
the apex of the costal cellule is brownish. Abdomen black,
the two basal segments rufous; the sides broadly and apex of
the basal segment and the apices of the second, third, and
fourth segments yellow; the petiole below, except for a
narrow line in the middle, and the apices of the second and
third segments are yellow.
A distinct species allied to P. maculipennis and P. stigma ;
easily known from them by the much longer and narrower
clypeus ; it differs also in having a distinctly defined area
above the clypeus, bounded by distinct furrows, and the apex
of the clypeus is quite differently formed, it making a semi-
circle, while in sé¢gma its sides are straight and oblique.
Genera and Species of Hymenoptera. 415
Polistes ephippium, sp. n.
Niger ; clypeo, orbitis oculorum, mandibulis, prothorace, scutello,
postscutello, lineis duabus metanoti petioloque rufis; pedibus
rufis, apice tarsorum nigris ; alis fuscis, costa stigmateque rufo-
testaceis. 9.
Long. 16-17 mm.
Hab. Khasia Hills. Coll. Rothney.
Antenne rufous, the flagellum darker above. Head black,
the outer orbits, the inner from the inside of the lower part of
the incision, the space between the antenne and the clypeus
rufous. The front and vertex rugose, thickly covered with
short fuscous hair. Clypeus with scattered punctures and
with short fuscous hair; its edges narrowly black. Man-
dibles rufous; the teeth black. Prothorax rufous, except
the basal slope. Mesothorax black; the scutellum, post-
scutellum, the sides near the tegule, the tubercles, and an
oblique mark (narrowed gradually beneath) below them
rufous. Pro- and mesonotum closely rugosely punctured, as
are also the scutellum and postscutellum, thickly covered with
fuscous pubescence. The striz on the metanotum are not
very close together; on each side is a large red mark, broad
at the top, gradually narrowed towards the apex. Legs
rufous ; the greater part of the coxe below, more or less of
the trochanters, the base of the femora, and the apical three
joints of the four hinder tarsi, black. Abdomen black, the
petiole and the apical segment rufous.
A stout easily recognized species.
Polistes khasianus, sp. n.
Niger ; clypeo, pronoto late, apice petioli segmentoque 4° rufis ; alis
fuscis, stigmate ochraceo. &.
Long. 12 mm.
Hab. Khasia Hills. Coll. Rothney.
Antenne black, thickly covered with pale pubescence.
Head black, the clypeus (except narrowly round the eyes)
red; there is a narrow red band on the upper outer orbits.
Vertex closely and distinctly punctured, as are also the eye-
incisions and the front broadly in the middle; they are
covered thickly with longish fuscous hair. Clypeus sparsely
punctured and with a few blackish hairs. Thorax black,
closely punctured and thickly covered with short pale hairs;
the base of the prothorax to near the coxe and the pleure
broadly above brick-red ; the red on the pleuree broad and
416 Mr. P. Cameron on new
rounded behind. Scutellum sparsely, postscutellum more
thickly rugosely punctured, behind almost impunctate. The
apex of the median segment has an oblique slope; on either
side on the upper part are three curved distinctly separated
keels. Pro- and mesopleure closely and distinctly punc-
tured; the base of the metapleuree smooth, except for a
narrow row of striz at the base; the apex and the lower part
behind the oblique furrow with shallow round punctures and
irregularly striated. Wings dark smoky, lighter behind and
below towards the apex ; the stigma ochraceous; the nervures
black. Abdomen black, the apical third of the petiole and
the fourth segment almost all round brick-red; the red band
on the fourth segment is incised at the base on either side.
A distinct species. There is no known Indian species
with which it can be confounded. The apex of the meta-
notum is not quite so closely striated as in most of the
species.
Polistes Wattit, sp. n.
Flavus ; tibiis posticis supra nigris ; alis fulvo-hyalinis, nervis testa-
ceis, 8.
Long. 13 mm.
Hab. Bengal (Dr. George Watt, F.L.S.).
The third joint of the antenne is nearly as long as the
following three united, the apical slightly longer than the
penultimate. Front and vertex finely granular, of a darker
shade than the face. Clypeus broader than long, the middle
at the top transverse, the sides oblique, the apex with the
sides oblique, the middle rounded. Mandibles yellow; the
teeth black, of nearly equal size, bluntly triangular. Pro-
and mesothorax smooth; the scutellum has a narrow longi-
tudinal furrow in the centre at the base. Median segment
closely transversely striated, the furrow deep. Wings fulvo-
hyaline, suffused with fuscous ; the second cubital cellule at
the top is slightly less than the distance bounded by the first
transverse cubital and the first recurrent nervures. Legs
coloured like the thorax, except that the hinder tibie are
black behind, this being also the case with the base of the
metatarsus. The base of the petiole, the greater part of the
second segment, and to a less extent the base of the third, are
infuscated, the darkened bands being dilated at the sides at
the apex.
This species has no near ally, and is readily known by the
unicolorous body. ‘The clypeus is broader than usual; the
antennal keel is broad, forming a large tubercle with a tovea
Genera and Species of Hymenoptera. 417
on the top; above it is a short narrow furrow; on the front,
below the ocelli, is a narrow, not very distinct furrow.
Polistes japonicus, sp. n.
Long., 6 2, 22 mm.
Hab. Japan (George Lewis).
This species so closely resembles the exceedingly variable
P. hebreus, especially the dark varieties, that it easily might
be passed over for it. The difference in the form of the
clypeus, however, enables them to be readily separated. In
hebreus ¢ the clypeus above has its sides straight and oblique;
in japonicus the top forms a semicircle, the sides not being
straight. The males are easily separated by the fact that in
hebreus the bordering furrow of the clypeus touches the eyes,
while in japonicus it 1s separated from them by a clear space;
in the latter, too, the top is bordered by a distinct furrow,
which is incised in the middle; in the common species this
upper bordering furrow is absent or scarcely noticeable and
it does not project downwards in the middle. In japonicus
the curved transverse lines on the abdomen are united to a
broad black band at the base of the segments, so that enclosed
lateral marks are formed. Head testaceous ; the occiput, the
vertex from shortly behind the middle to the upper half of the
eye-incision, the part between the antenne, and the sides and
top of the clypeus narrowly black. Clypeus sparsely punc-
tured on the lower half, the apex with the sides oblique, the
centre rounded. Front and vertex distinctly, but not very
closely, punctured, the front with a narrow but distinct furrow.
Mandibles testaceous, the three teeth of nearly equal size.
Thorax black, the edge of the pronotum behind, the upper
third of the sides, their base more narrowly, leaving in the
centre a triangular black space, two large lines in the middle
of the mesonotum, a curved spot near the tegule, the scutel-
lum, postscutellum, two lines (narrowed gradually below) on
the slope of the metanotum, a mark (narrowed below) near
the tubercles on the mesopleure, and a more irregular one
(narrowed gradually towards the middle) on the apex lower
down, and a mark in the centre of the metapleure: behind the
furrow, yellow. Mesonotum obscurely punctured on the sides ;
the metanotum transversely striated, its central furrow much
wider than in Aebreus. Pleure obscurely punctured, the lower
part of the mesopleuree smooth at the base. Legs testaceous ;
the coxa, trochanters, the base of the front femora, the basal
half of the middle, the hinder (except at the apex), the four
anterior tibie and tarsi, and the hinder tarsi (except at the
418 On new Genera and Species of Hymenoptera.
base) rufo-testaceous. Wings hyaline, with a smoky viola-
ceous tinge; the costa and stigma rufo-testaceous. Abdomen
testaceous, the basal two thirds of the petiole (except for two
irregular curved marks), the base of the second segment
broadly (with the middle dilated), a curved line running from
its centre to the sides, where it becomes united to the base,
thus forming a large enclosed space, and similar curved lines
on the third, fourth, and fifth segments, but with the basal
lines much narrower, black; the ventral segments are some-
what similarly marked, the basal two are almost entirely
black.
The male is similarly coloured, but with more black on
the vertex behind; the clypeus is clearly bounded by a deep
furrow at the sides and above and distinctly separated from
the eyes ; the middle is turned down on the top; the sides of
the epipygium are broadly and roundly raised, in the centre
behind are two stout teeth, rounded behind.
Polistes erythrocerus, sp. n.
Niger; prothorace late, lineis 2 mesonoti, scutello, postscutello
lineisque 2 metanoti flavis; abdomine late flavo balteato ; alis
fulvo-fumatis, stigmate nervisque fulvis. 9.
Long. 22 mm.
flab. Kamakura, Japan (George Lewis).
Antennze rufous, the scape lined with black above; the
third joint is slightly longer than the fourth and fifth united,
the last two equal in length. Head fulvous yellow ; a broad
band on the ocellar region, extending to the eyes and round
the upper part of the eye-incision, a line above and below
the base of the antenne, the occiput, a line above it (united
to the eyes by a small oblique line), black; the part between
the antenne is distinctly tuberculate, the centre of the upper
part keeled. Front and vertex distinctly but not very closely
punctured. Clypeus very sparsely punctured, longer than
broad ; the sides of the apex oblique, the top narrowed, its
sides rounded, the middle roundly incised. Mandibles
coloured like the head, except that the teeth are black.
Thorax black; the prothorax (except for a triangular black
mark near the middle behind), two lines in the middle of the
mesonotum, two short ones opposite the tegule, the scutel-
lum (except in the middle), its keels, the postscutellum, two
large lines (narrowed below) on the metanotum, and a small
irregular mark under the tubercles, yellow. Pro- and meso-
notum closely and distinctly but not deeply punctured, the
punctures running into reticulations ; the middle impunctate
On the Presence of Echis coloratus in Africa. 419
at the base. The scutellum is, if anything, more rugosely
punctured than the mesonotum; the postscutellum is more
sparsely punctured. Median segment transversely striated
as usual. Propleuree punctured, more closely above than in
the middle. Mesopleure more closely and strongly punctured ;
metapleuree much less strongly punctured, the middle almost
smooth. Coxe, trochanters, and femora black; the fore
cox below, the fore femora above and below, and the hinder
at both sides below yellow ; the fore tibie rufous, tinged with
yellow ; the middle tibiz rufous, lined with black at the base
behind ; the posterior entirely black behind ; the tarsi rufous,
the joints black at the apex. Wings fulvous, tinged with
violaceous, the hinder more hyaline in tint. Abdomen
black ; the apex of the petiole narrowly and the apical half
of the other segments testaceous.
One of the examples bears a large species of Stylopide.
[To be continued. |
LV.—On the Presence of Kchis coloratus, Géinther, in
Africa. By JoHN ANDERSON, M.D., LL.D., F.R.S.
THIS viper, along with four other species of reptiles, was
found by Mr. D. MacAlister in the neighbourhood of the
Emerald Mines on the coast of the Red Sea, in nearly the
same latitude as Assuan, and was presented by him to the
British Museum. I am indebted to the courtesy of Mr. G. A.
Boulenger, F.R.S., for his permission to direct attention to
the occurrence of this snake in the Eastern Desert of Egypt,
as the species is not included in my work on the Reptiles and
Batrachians of Egypt, this being the first occasion on which
it has been met with on the African continent.
Mr. MacAlister informs me that the five species of reptiles
collected by him in the afore-mentioned locality were obtained
in a tract of country not more than 9 miles in breadth at any
point, lying between the Wadis Rousbaid, Sakét, and Nugrus,
which open one into another and ultimately join the Wadi
Gemal. ‘The first two of these Wadis are dry and stony, but
they are covered more or less with “low scrub, Gash (Bas-
silla) and Sayal (Mimosa), and occasionally with small
pumpkins.” ‘lhe Wadi Nugrus is sandy and with rather
more scrub than in the two preceding Wadis, and the Wadi
Gemal partakes of the same character, but it is more open.
420 Dr. J. Anderson on the Presence of
The snakes were caught in the Wadis Nugrus and Sakét,
and the lizards in the latter and in the Wadi Rousbaid.
Mr. MacAlister says, “ the Reptiles apparently live in the
cracks in the gneiss.”
Echis coloratus, Gunther.
Mr. MacAlister captured two females, the measurements &c.
of which are recorded in the following table along with those
of the type specimen :—
Echis coloratus, Gthr.
: 2. Q.
Type.
Midian, Eastern Desert Eastern Desert
Arabia. of Egypt. of Egypt.
Snout to vent........ 660 485 378
ail orth caves tne oda 80 67 54
Wenttals scsi oases fon 208 188 194
NAD Cr oct as ee eae 1 it 1
Caudals 2.2.50) wos 46 51 51
Senles eft cueremis 35 35 35
The type of E. coloratus, Giinther, was found by the late
Sir Richard Burton * at Jebel Sharr, behind El Mewaylah,
in Midian, to the south of the entrance to the Gulf of Akabah,
at an elevation of 4500 feet above the level of the sea.
The scales on the upper surface of the head, from the snout
to between the eyes, anteriorly, are almost smooth in &. colora-
tus, whereas in EZ. carinatus they are strongly keeled ; but on
each scale can be detected a central eminence, the equivalent
of the elongated and generally well-developed keel which
occurs on the scales further back. ‘There are 13 rows of
scales across the head between the eyes, but no defined supra-
ocular is present. The number of scales round the eyes varies,
as on the right side there are 19, and on the left side only 16.
There are four rows of scales between the upper labials and
the eyes. Twelve to thirteen upper labials, the numbers
varying on the opposite sides of the head. Rostral twice as
high as broad. Nasal shield variable, not wholly divided on
the right side, but completely so on the left, invariably sepa-
rated from the rostral by two almost shield-like scales, the
outermost lying on the first labial, with the innermost, the
larger of the two, placed above it and separated from its fellow
of the opposite side by two small scales.
* «Gold Mines of Midian,’ 1878.
Echis coloratus, Giinther, in Africa. 421
Gitinther described the colour of the type as “¢ greyish, with
large pinkish spots on the upperside; they are rounded on
the front part of the body, but more irregular and broken up
on the posterior. Lower parts whitish, speckled and powdered
with greyish.”
Mr. MacAlister’s two specimens present slight variations
from the type, and these are brought out in the two accom-
panying figures of the front of the snout of (a) the type and (6)
of one of the individuals from the Eastern Desert.
Echis coloratus.
In the type of Z. coloratus the rostral is a dome-shaped
shield, whereas in the Hgyptian specimens it is more or less
quadrangular. The form of this shield, however, is doubtless
subject to much variation, depending on the number and
disposition of the shields around it. In the Midian viper one
infranasal intervenes between the nasal and the rostral, and
behind it lie two other shields, one above the other, the upper-
most in contact with the nasal and the one below it resting on
the rostral and on the first labial. The internal infranasals
are separated from each other in the mesial line by two small
scales resting on the rostral. The supranasals are broadly
excluded from the rostral by the foregoing two scales and by
the first infranasal. In fig. 6 only one infranasal rests on
the rostral, but behind it another infranasal exists between
the nasal and front labial without being in any way in contact
with the rostral. In this individual the equivalent of the
second suprarostral of the type seems to have fused with the
rostral and with the front labial, and in so doing to have
given greater transverse breadth to the former shield and
greater height to the latter. ‘These variations in no way
connect the vipers presenting them with LE. carinatus.
In the type of £. coloratus the supranasals are more or less
irregular in form, widely separated from the rostral by the
internal infranasals and by the median pair of scales which
lie between the latter shields; but in fig. 6 (Hastern Desert
viper) the right supranasal is prolonged down to the rostral on
422 Dr. J. Anderson on the Presence of
the right side, whilst the supranasal of the opposite side is
excluded from the rostral by asmall scale. In the other viper
from the Emerald Mines both supranasals touch the rostral,
but in the two individuals from that locality the supranasals
in each case are separated in the mesial line by a single scale.
The downward projection of the supranasals in these
examples of H. coloratus, feeble though it be, is a variation in
the direction of £. carinatus, in which these shields are
markedly developed, broadly in contact with each other in
the mesial line and in a varying degree with the rostral. In
a specimen of H. coloratus from the Dead Sea the supranasals
are large and directly in contact with the rostral, but in
another from the same locality they are excluded by the infra-
nasals. In both of these vipers the nasals are separated from
the rostral by infranasals. The same variation occurs in
vipers from the Hadramut, but, as in . car¢natus, the supra-
nasals are in contact mesially. In a specimen in the British
Museum said to be from Socotra the rostral is rounded like
the type and the supranasals are excluded from contact with
it by the infranasals, of which there are two, the second
resting on the first upper labial.
The essential feature of “. coloratus is the broad separation
of the nasal shields from the rostral and from the first upper
labial, whereas in £. carinatus the nasals rest directly on the
rostral and first upper labial. The accompanying figures
(c and d) represent the front view of the snout of £. carinatus
from Sind and from Jebba on the Upper Niger respectively ;
and if they are compared with the two modifications of E. colo-
ratus (figs. a and b) the differences between the two species
become apparent.
Echis carinatus.
As the scales around and between the eyes in the two
species vary very nearly within the same limits as to
numbers, they are practically by themselves of little use as
guides to the distinction of the species; and the same may
be said of the scales between the eyes and upper labials, as
Echis coloratus, Giinther, in Africa. 423
examples of £. coloratus are met with having three rows of
such scales, a number occurring occasionally in #. carinatus, in
which, however, these scales are usually in two rows, whereas
in L. coloratus they are either in three or four. The enlarged
supraoculars generally present in the former species are
usually absent in the latter, and when present they are not
well developed.
In the specimens of /. coloratus here tabulated the ventrals
range from 188 to 203, and the subcaudals from 46 to 51;
whereas in /. carinatus these shields vary, over the vast area
of its distribution, the ventrals from 132 to 194, and the
subcaudals from 23 to 48. So few examples of LZ. coloratus
are known, it is quite possible that the numbers of its ventrals
and consequently of its trunk-vertebre may rise much higher
than the maximum number given above and also fall much
below the minimum of 188. The type of Z. carinatus had
only 150 trunk-vertebree, whereas now, after more than a
century of research, individuals have been found with as few
as 132 and others with as many as 194—that is to say, that
there may be as great a variation as 62 vertebrae. This
result is obtained by the consideration of many individuals
from Asia and Africa brought together from between Madras
and West Africa; but the vertebrae in question may vary as
much as 89 among individuals from the same locality
(Deccan, India), whereas in others from Sind and from West
Africa the number of trunk-vertebree may be practically the
same, viz. 143 and 145. ‘The existence of this enormous
range of variation in the number of the vertebra of the indi-
viduals of this genus, even although it takes place in structures
which are mere repetitions the one of the other, is very
remarkable, because associated with it there is of necessity
a corresponding modification of the nervous and vascular
systems. With such variations occurring in the form of an
animal the possible modification of the epidermal structures
would seem to be almost limitless. The modus operandi by
which Nature effects these unstable variations of the skeleton
and soft tissues associated with it is an enigma unsolved by
any hypothesis as yet formulated.
The coloration of these vipers from the Eastern Desert is
much the same as in the type, and as they have been quite
recently killed, its general character is better made out.
There are from 43 to 46 more or less transverse, narrow, pale
greyish areas, becoming almost light grey spots in the younger
specimen on the hinder part of the body. In both these pale
areas are margined with dark finely speckled grey, the inter-
424 On the Presence of Echis coloratus in Africa.
vening spaces being reddish buff. The sides of the body on
the region of the serrated scales are marked with finely
speckled dusky spots, more or less connected with the dorsal
reticulations. There are also some dusky spots on the angles
of the ventrals. The ventral surface is white, but with faint
dusky spots here and there. This type of coloration corre-
sponds in its broad outlines with that of H. cartnatus. In
two females of the latter species from Sind in the British
Museum there are 36 and 40 spots respectively much more
clearly and vividly defined than in £. coloratus, whilst on
the tail the spots are practically absent. A specimen from
Muskat in the British Museum has the same general type of
coloration, but the dark and light markings are less defined,
whilst in another young specimen from the same locality the
light spots are absent and the dark markings are reduced to
two very narrow parallel dorsal lines, largish brown spots
occurring on the lateral serrated scales and a small brown
spot on the angle of nearly every third ventral. In the
specimen supposed to have come from Socotra the general
colour above is dark slaty, with brown spots more or less
obscurely present along the mesial line of the back, and irre-
gularly shaped brown spots along the sides. No pale dorsal
spots are present, but the ventral surface is obscurely spotted
posteriorly. The specimens from the Dead Sea are pale
yellowish, with the dark and pale markings not clearly defined
and the ventrals immaculate.
In no viper with infranasals has the head ever been found
to have its upper surface bearing the symmetrical dark brown
markings present in a greater or less degree in vipers with
their nasals resting directly on the rostral and on the front
upper labial.
£. coloratus is as yet known only from Arabia, Southern
Syria (Dead Sea), the Eastern Desert of Egypt, and from
the island of Socotra.
Ptyodactylus Hasselquistii, Donndorft.
This is an example of the pallid typical form with the
nostril but little, if at all, tubular. This is the most southern
point it has yet been recorded from the Eastern Desert, but
it is common as far south as Wadi Halfa.
Agama spinosa, Gray.
Two specimens, differing in no respect from the examples
from Suakin. As already pointed out by me, the type of
Bibliographical Notices. 425
this species, presented very many years ago to the British
Museum by James Burton, the distinguished Egyptologist,
came, in all likelihood, from the Hastern Desert about the
latitude of Keneh.
Eremias guttulata, Licht.
Three specimens. This species is distributed all over
Egypt, from Suez to Suakin.
Psammophis Schokart, Forskal.
This specimen, in its almost pale fawn-colour and the
presence of two darker-coloured lines along the back and the
obscure dotting of the ventrals, especially on the sides, and
distinct head-markings, resembles the individuals of this
species found at Durrur to the south. The number of its
ventrals (190) largely exceeds the number found there and at
“Suakin, and in this respect it conforms to snakes of this
species found at Assuan. It has 118 caudals.
BIBLIOGRAPHICAL NOTICES.
Bird Books.
In Bird-land, with Field-glass and Camera. By Otiver G. Prxe.
T. Fisher Unwin & Co. Pp. 1-280, with 83 photographs from
Nature. 1900. Price 6s.
The Birds of Eastern North America. By Cuartrs Cory. Printed
for the Field Columbian Museum, Chicago, Ill. 1899. Parts
| a
Tue strides which photography has made among us of late years
we regard with a jealous eye. Not satisfied with ousting the
beautiful wood and steel engravings which adorned the pages of
our older books, it has gone now so far as to displace literature
itself, so that many of the ‘“‘ books” which are thrust upon a long-
suffering and defrauded public to-day are practically little more
than collections of bad pictures served up with a sprinkling of worse
text—added for appearance’ sake.
To every rule, however, there are exceptions ; and there are
occasions when we feel real gratitude towards the enthusiastic
photographer.
Ornithologists are without doubt indebted to the Kearton brothers
for the work which they have done and recorded in this field. The
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 28
426 Bibliographical Notices.
present little book ‘In Bird-land,’ by Mr. Pike, may well rank
beside the best in this direction. The birds and their ways are for
the most part feelingly described, and the author shows himself to
be not only an observer but also a lover of nature. We are at one
with him in his condemnation of the gamekeeper. The fauna of
these Islands has suffered great and irreparable losses at his
ignorant hands. But if the keeper is to be subjected to the lash of
our displeasure, how much more 0 shall his employer, who signs
the death-warrants which he executes ?
Many of the illustrations of this book are excellent. Some are
bad. ‘The frontispiece forces a grumble from us. It is really
beautiful, and it seems a pity that, since such work is possible,
some of the inferior or less interesting pictures were not suppressed,
and the remainder reproduced in the same way as the frontispiece.
What was lost in quantity would be repaid a hundredfold in
quality.
There is some excuse for the use of photography in reproducing
actual outdoor scenes, or objects of natural history taken at first
hand, but to call in its aid to reproduce bad drawings is indeed a
sin. ‘The illustrations to the ‘ Birds of Eastern North America’ is
acase in point. The figures in this work are for the most part
grotesque ; they could scarcely be worse.
As a “key” the book is doubtless useful. The terms, however,
employed in describing the topography of a bird are often faulty,
sometimes very much so. We must protest against the use of the
word ‘‘tertials ” to indicate the long inner secondaries of the wing.
On page 3 “tertials” are defined as “the few remaining remiges
which grow from the humerus.” ‘The feathers called “ tertials ” in
the plates are only long inner secondaries: it is very doubtful
whether the parapteron and hypopteron can legitimately be regarded
as remiges.
With a little pruning and revising this book could be made worthy
of its author.
A Natural History of the British Lepidoptera. A Textbook for
Students and Collectors. By J. W. Tort, F.E.S. Vol. I.
London: Swan Sonnenschein & Co., May 1900. Pp. viii, 584 ;
plates vii.
Tue first volume of Mr. Tutt’s great work on British Lepidoptera
appeared in January 1899, and already the second volume is
lying before us. We are glad to find that the author has
received so much encouragement that he is enabled to proceed
with the book without delay and in the most elaborate manner.
The second volume is thicker than the first by no less than 24
pages, and is similarly divided into two parts. The first part
Bibliographical Notices. 427
is divided into five chapters, of which the first two deal with
metamorphosis in Lepidoptera, and the others with the external
morphology, internal structure, and phylogeny of the Lepidopterous
pupa. The second part includes the Psychides (divided into
Micropsychina and Macropsychina), a catalogue of the Palearctic
Psychides, the first portion of the Lachneides, and Index. The
author has not only epitomized a large part of the extensive
literature relating to the various subjects of which he treats, but
has added a very large amount of entirely new and original matter,
derived from the observations of himself and his correspondents.
No less than 334 pages of the second volume are devoted to the
interesting, but extremely difficult, group of the Psychides, which is
one of the most remarkable among the Lepidoptera. The females
are almost always apterous, and in some species are almost destitute
of legs and antenne as well, being thus reduced to the condition of
mere helpless egg-bags. ‘The larvee form cases for themselves on
the plants on which they feed, somewhat resembling those formed
by the larvee of caddis-flies (Trichoptera), to which some entomo-
logists have considered the Psychide to be allied. Here the pupa
is formed, and the more helpless females never quit it, but deposit
their eggs within it. Another peculiarity is that parthenogenesis
is So common in some of the species, especially in the genus Solenobza,
that you may go on breeding from the larva-like female for genera-
tion after generation without ever seeing a male, which greatly
adds to the difficulty of satisfactorily separating and defining the
species.
Mr. Tutt has thoroughly reviewed this difficult group. His
Catalogue of the Palearctic Psychides includes no less than 11
families, 20 subfamilies, 36 genera (of which 8 are new), and 143
species, besides varieties, &¢. Among the Psychide, Mr. Tutt places
several genera which many previous authors have included in the
Tineides, such as Diplodoma, Lypusa, Melasina, Solenobia, Taleporia,
&c. But if we exclude these, we find that the Psychides proper,
which a few years ago used to form a single family, of three genera
at most, and which were often included in one, has now expanded
to four families, comprising twelve sections, and twenty-seven
genera! ‘This will appear to old-fashioned entomologists a terrible
and unnecessary amount of subdivision, but in most similar cases
the foresight of the innovator is, sooner or later, largely justified
by the judgment of his successors.
The natural history of each species is also worked out as
exhaustively as before; thus the account of Pachythelia villosella,
Ochs., occupies more than eighteen closely printed pages.
The reprint of the original description of each genus and species,
whether short or long, is a great assistance, especially as the original
types of the genera are clearly indicated. Had this always been
done, we should have been spared a tremendous amount of con-
fusion, though few cases are quite so glaring as that of the genus
428 Bibliographical Notices.
(Ecophora, to which we may here allude, though it does not belong
to the families that have yet been discussed by Mr. Tutt. Latreille
established the genus Geophora for @. sulphurella, Linné, which
remained the type for twenty years, Curtis actually figuring it as
such, and Stephens sinking his own genus Dusycera as a synonym.
But Zeller gave the genus a wider extension, dividing it into three
sections, for the last of which he adopted Stephens’s discarded name
of Dasycera. Then Stainton adopted Geophora for Zeller’s first
section, separating sulphurella (the inalienable type of @eophora) as
Dasycera. Then Mr. Meyrick established two families, (icophoride
and Dasyceridee, thus turning the very type of Gcophora out of
the family Cicophoride itself! This has since been rectified, but it
is only an illustration of many similar cases which will occur to any
entomologist who troubles himself about synonymy.
The remainder of the present volume is devoted to a portion of
the Lachneides (or Lasiocampides), and the classifications of Hubner,
Aurivillius, Dyar, and others are quoted in full. Ouly five species,
however, are dealt with in the present volume : Pocilocampa populi,
Trichiura crategi, Lachneis lanestris, and Malacosoma castrensis and
M. neustria. Mr, Tutt estimates that the remaining five genera
and six species of the superfamily Lachneides will occupy 200
pages of the next volume. In his remarks on the phylogeny of
the Lachneides, we are pleased to see that while freely expressing
his own views, and criticizing those of his predecessors, he puts them
forward tentatively, and quite avoids the dogmatic tone assumed by
certain writers on what must necessarily long remain one of the
most difficult and uncertain problems in entomology—all the more
so, because, in Lepidoptera at least, we have nothing but the barest
fragments of any Geological Record to help us to verify any of our
conclusions ; and without this we are necessarily only groping in
the dark.
Five of the seven plates in the present volume are devoted to
Psychides—phylogeny, neuration, spurs, antenne; imagines and
cases of Whittleta retiella; and the transformations of the American
Thyridoptery« ephemereformis. Plate 1 is devoted to the wings,
wing-scales, &c. of Lepidoptera, and plate 7 to Dyar’s phylogeny of
the Lachneides.
We can fully sympathize with what Mr. Tutt says in his preface
about the difficulty of getting more matter into each volume, but
yet we should like to suggest that it would be very useful to include
in the Contents a list of the British genera and species discussed
in each volume. As the number of these is very limited, this would
require very little space, and would probably not involve the
sacrifice of more than a single page. W. oge
THE ANNALS
AND
MAGAZINE OF NATURAL HISTORY.
[SEVENTH SERIES.]
No. 35. NOVEMBER 1900.
LVI.—Astatic Torrrrorp2.
By the Rt. Hon, Lorp Watsineuam, M.A., LL.D., F.R.S.
[Concluded from p. 409.]
LASPEYRESIA, Hb.
1127 (1). Laspeyresia tridescens, sp. n.
Antenne greyish fuscous. LPalpi short, recurved, slightly
erect, with appressed scales ; very pale ochreous. Head small
and prominent; greyish. Thorax brownish fuscous, sprinkled
with whitish ochreous scales. ore wings with the termen
slightly indented below the apex ; brownish fuscous, minutely
and profusely speckled with whitish ochreous (with a strong
greenish iridescence in a bright light if held at certain angles) ;
on the outer half of the costa are some short whitish ochreous
oblique streaks, the spaces between them being greyish
fuscous (darker than the ground-colour) ; from these streaks
run one or two very inconspicuous slaty grey lines, and the
ocelloid patch, which contains three blackish spots, is bounded
outwardly by a similar line; cilia brownish grey, with a
slender dark line along their base; underside strongly irides-
cent, costal markings not visible. Hap. al. 17-19 mm.
Hind wings dark brown, slightly paler towards the base ;
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 29
430 Lord Walsingham on Asiatic Tortricide.
cilia yellowish white, at the flexus greyish, with a dark line
running throughout them near their base. Abdomen greyish
fuscous. Legs greyish, hind tarsal joints pale spotted.
Type, 2 (60138); g (60141) Mus. Wlsm.
Hab. CorrA—Gensan, VI. 1886 (Leech) ; Fusan, 9 VI.
1886 (Leech). JAPAN (Pryer, 1886)—Kzrvsrv—Satsuma,
V. 1886 (Leech). Five specimens.
This species appears to the naked eye dull and unicolorous
with the exception of the costal streaks; it differs from nigri-
cana, F., in the absence of any visible markings on the under-
side, as well as in its larger size and shorter costal streaks.
1128 (1). Laspeyresia quadrocellana, sp. n.
Antenne brownish fuscous. Palpi ochreous. Head and
thorax brownish fuscous. ore wings brownish fuscous, with
a beautiful greenish and purplish iridescence, especially on
the basal half; on the costa are five pairs of oblique pale
ochreous geminated streaks, the first two pairs indistinct
before the middle; a single streak lies between the third and
fourth pairs, and these both terminate in outwardly oblique
steel-blue lines, scarcely disconnected from two similar lines
beneath them which form the margins of an upright ocelloid
patch opposite to the middle of the termen, containing four
black spots set in pale ochreous rings ; two black spots occur
between the ocellus and the termen; on the dorsum is an
obscure pale ochreous patch, consisting of four parallel lines,
all curved outward and attenuated to their termination at the
lower margin of the outer third of the cell; cilia shining
grey, with a dark line along their base, interrupted by a
slight incision below the apex. Hap.al.l14mm. Hind wings
dark brown; cilia whitish cinereous, with a dark line along
their base. Abdomen and legs greyish brown.
Type, & (60797) Mus. Wlsm.
Hab, JAPAN—Krwusiv (Leech, 1890). Unique.
Allied to nebritana, Tr.
1128 (2). Laspeyresia pavonana, sp. n.
Antenne greyish fuscous, paler on their underside. Palpi
somewhat recurved, slightly erect; ochreous. Head more
retracted than in zrédescens, Wlsm.; greyish fuscous. Thorax
greyish fuscous, with a slight greenish iridescence. Sore
wings, 3 without a costal fold, termen indented below the
apex; greyish fuscous, strongly iridescent, with green and
purplish reflections; from the middle of the dorsum four
waved parallel subochreous lines extend obliquely outwards
Lord Walsingham on Asvatic Tortricidae. 431
towards the outer end of the cell, reaching halfway across the
wing ; on the costa isa series of oblique pale ochreous streaks,
two before the middle, each geminated, short, and incon-
spicuous, six beyond the middle, of which the two nearest the
apex are longer and less oblique than those which precede
them; the spaces between these streaks are dark umber-
brown ; from the first and fourth streaks beyond the middle
dark steel-blue lines extend outwards, the second reaching to
the apex of the outer pair of ochreous streaks; beneath it is
an olivaceous brown ocelloid patch containing some transverse
black lines and margined by steel-grey streaks, the inner one
of which reaches nearly to the tornus, where it is preceded by
an umber-brown shade; beyond the ocelloid patch the termen
is shaded with umber-brown, containing a series of about five
black dots; a slender dark line runs along the extreme
margin at the base of the brownish grey cilia; on the under-
side, which is also iridescent in certain lights, the ochreous
costal streaks are plainly visible. zp. al. 16mm. Hind
wings dark brown ; cilia pale greyish, with a dark line along
their base; underside iridescent. Abdomen greyish fuscous.
Legs cinereous, inclining to ochreous.
Type, 2 & Mus. Wlsm.
Hab. JAPAN—Howpo—Shimonoseki, VII. 1886 (Leech).
Two specimens,
Very similar to guadrocellana, Wlsm., from which it may
be distinguished by the black markings in the ocellus and
preceding the termen. In quadrocellana the series of four
ocelloid black dots set in pale ochreous rings slant outwards
to the tornus, and only two black dots occur between the
ocellus and the termen. In pavonana the upper three black
streaks point inward, the two lower ones forming, with three
or more black dots between the ocellus and termen, a sub-
terminal series parallel with the margin of the wing.
1135 (1). Laspeyresia adenocarpi, Rag.
Grapholitha adenocarpi, Rag. Bull. Soc. Ent. Fr, 1875, lxxiii; Ann.
Soc. Ent. Fr. (5s.) VI. 406-8, Pl. VL 4 (1876).
Hab. Europe. Asiatic TuRKEY—Hazzez—Shar Devesy
(Native Coll. 1893).
1136. Laspeyresia cecana, Schig.
Grapholitha (Semasia) cecana, Stgr. & Wk. Cat. Lp. Eur. 256.
No, 1136 (1871)*; Stgr. Hor. Soc. Ent. Ross. XV. 258 (1879) 2.
Hab. Europe’. Asiatic TuRKEY — Kuaupaveyprerar—
Brussa, V.?; Szvas—Amasia, IV.?; //ares—Shar Devesy
(Native Coll. 1890).
29*
432 Lord Walsingham on Asiatic Tortricide.
1148 (2). Laspeyresia quadristriana, sp. n.
Antenne brownish grey. Palpi short, porrect, median
joint brush-like beneath, terminal joint smooth exposed ;
whitish. Head brownish grey, face whitish. Thorax
brownish grey. ore wings with the termen indented below
the apex; brownish grey at the base, shading to brownish
beyond the middle; on the middle of the dorsum is a patch
of four whitish ochreous lines running at right angles to the
margin for one-third of their length, thence bent obliquely
outwards and reaching nearly halfway across the wing; on
the costa is a series of nine oblique whitish ochreous streaks,
the outer one somewhat inverted, of these the third and fourth
and the fifth and sixth combine in sending obliquely forward
from each pair a short bright blue streak, and the last streak
before the apex is also tipped with bright blue; the extreme
apex is black, the ocelloid spot above the tornus is indicated
only by a narrow upright shining cupreous patch; cilia
shining rosy grey; underside shining iridescent green-grey,
the pale costal streaks distinctly visible. Hp. al. 11°5 mm.
Hind wings brownish; cilia pale cinereous; underside
strongly iridescent. Abdomen greyish, whitish beneath.
Legs whitish ochreous.
Type, & (70112); ¢ (70113) Mus. Wlsm.
Hab, JAPAN (Pryer, 1886). W. Cutna—Foochow, IV.
1886 (Leech). Hight specimens.
1155 (2). Laspeyresia gradana, Chr.
Grapholitha gradana, Chr. Bull. Soc. Imp. Nat. Mose. LVI. 419-20.
No. 122 (1882)+: sep. 182-8 (1882) 3.
Hab. Amur, VII.'—Starikowo R., 8 VII. 1876 (Christoph);
Wladiwostok, 26-29 VII. 1877 (Christoph). Japan (Pryer,
1886).
1165. Laspeyresia duplicana, Zett.
Grapholitha (Semasia) duplicana, Stgr. & Wk. Cat. Lp. Eur. 258.
No. 1165 (1871).
Hub. Europe. Asiatic TurKEY—Hares—Shar Devesy
(Native Coll, 1893).
1167 (3). Laspeyresia fimana, Snell.
Grapholitha fimana, Snell. Tijd. v. Ent. XX VI. 225-6, Pl. XII. 7, 7 a
(1883) ?.
Heb. KE. Srperta—Askold Id.’; Wladiwostok, 25 V.
1877 (Christoph). JAPAN (Pryer, 1886).
Lord Walsingham on Asiatic Tortricide. 433
1168. Laspeyresia pallifrontana, Z.
Grapholitha (Semasia) pallifrontana, Steger. & Wk. Cat. Lp. Eur. 258.
No. 1168 (1871); Stgr. Hor. Soc. Ent. Ross, XV. 260 (1879) *.
Hab. Europe’. Asrtatic TurKEY—KuavpavenpickrAar—
Brussa®. Szvas—Kerasdere*, Maidan, 10-21 V*. Hazep
—Shar Devesy (Native Coll. 1893).
1168 (1). Laspeyresia dificilana, sp. n.
Grapholitha difficilana, Bang-Haas, Stgr. List, XLII. 25 (1888-9) MS.
Antenne blackish. Pu/pi somewhat erect; white. Head
blackish above; face white. ore wings bronzy blackish
brown, with a leaden grey tinge at the base; four pairs of
short silvery white costal streaks, each pair terminating in
shining steel-blue, all except the last pair pointing obliquely
outward ; on the middle of the dorsum a longer pair of evenly
curved silvery white streaks, the first terminating in a small
blue spot on the cell; arising from the tornus is a slightly
inverted bright iridescent steel-blue band, not quite reaching
the apex of the second pair of costal spots ; cilia whitish grey,
a dark fuscous line running along their base, but interrupted
below the apex by a white dot; underside whitish, with the
exception of the dark costal streaks on the outer half of the
wing, termen shaded with fuscous. vp. al. 11mm. Hind
wings: f whitish at the base, apex and lower margin
broadly cupreous brown; cilia snow-white, a dark line along
their base, around the apex only; underside whitish, with
one or two dark costal streaks near the apex, termen shaded
with fuscous: $ somewhat more darkened than in the ¢.
Abdomen blackish, banded with white beneath. Legs, hind
pair silvery whitish, much shaded with greyish fuscous ex-
ternally, terminal joint of the tarsi fuscous, spurs white.
Type, & (61611); 2 (7277) Mus. Wlsm.
Hab. Astatic ‘TuRKEY—Hazrzes—Shar Devesy (Native
Coll. 1893) ; Zeitun (Stgr.). Five specimens.
This species is allied to pallifrontana, Z , and interstinctana,
Clem. ; from the first it may be at once distinguished by its
more erect and whiter palpi and by the hind wings in both
sexes having a pale space at the base; the latter distinction
also separates it from interstinctana, but interstinctana need
not be confused with pallifrontana, since that species is easily
recognized by the much wider separation of the first and
second pairs of costal streaks.
434 Lord Walsingham on Aséatic Tortricide.
1168 (2). Laspeyresia geministriata, sp. n.
Antenne greyish fuscous. Palpi recurved, slender, smoothly
clothed, terminal joint about one third the length of the
median ; pale ochreous. Head greyish. Thorax brownish
grey. ore wings brownish grey, inclining to brownish
fuscous beyond the basal third; a pair of upright pale
ochreous streaks beyond the middle of the dorsum are
widened in the middle and narrowed towards their apex, the
first somewhat bent outwards, the second erect ; on the costa
are from five to seven very pale ochreous streaks, somewhat
ureguiar and varying in size, but all rather wide and con-
spicuous (from the difference in the number of these streaks
on the right and left wings they have evidently a tendency
to run together); beneath and connected with these costal
streaks are three short oblique rosy metallic dashes, and the
ocelloid patch is indicated by a rosy cupreous upright patch,
somewhat widened at its upper end; cilia shining rosy grey ;
underside strongly iridescent, the costal streaks being visible ;
there is also a pale spot on the termen below the apex which
does not appear on the upperside. xp. a/.12mm. Hind
wings brownish fuscous, slightly paler at the base; cilia
greyish cinereous; underside iridescent. Abdomen greyish
fuscous. Legs pale cinereous.
Type, 2 (60142) Mus. Wlsm.
Hab. W. Cutna— Foochow, IV. 1886 (Leech).
1169. Laspeyresia fissana, Fro).
Grapholitha (Semasia) fissana, Stgr. & Wk. Cat. Lp. Eur. 258. No. 1169
(1871).
Hab. Europe. Asiatic TuRKEY—L/arzesn—Shar Devesy
(Native Coll. 1893).
1173. Laspeyresia dorsana, F.
Grapholitha (Semasia) dorsana, Stgr. & Wk. Cat. Lp. Eur. 258. No. 1173
(1871).
Hab. Evrore. Asiatic TurKeY—fazrzs—Shar Devesy,
15 VI. 1890 (Native Coll.).
1173 (2). Laspeyresia junctistrigana, sp. n.
Antenne dark greyish fuscous. Pa/pi and head smoky
brown-grey. Thorax smoky blackish. Fore wings smoky
brownish black, shading to dark tawny brownish about the
apex ; with five pairs of short silvery white costal streaks
Lord Walsingham on Asiatic Tortricidae. 435
terminating in sinuate lines of steel-grey, one of such lines
serving for the third and fourth pair and apparently emanating
from the outer streak of the third; an ocelloid patch, con-
taining three or four short black streaks, is margined on both
sides by shining metallic lilac; a snow-white dorsal streak,
nearly erect, but slightly outcurved at its outer end above the
fold, is of equal width throughout, a few dark scales in its
middle at each extremity indicating reduplication ; cilia dark
brownish fuscous, with a slight cinereous admixture. Hap.
al.16 mm. Hind wings dark brown, whitish along the costa
nearly to the apex; cilia dirty whitish, a dark shade-line
running through their base. Abdomen dark fuscous. Legs
whitish cinereous, shaded externally and on the hind tarsi
with greyish fuscous.
Type, 2 (61610) Mus. Wlsm.
Hab. Astatic TurKey—Hazres—Shar Devesy (Native
Coll. 1893).
Cypra, Hb.
1181. Cydia pomonella, L.
Carpocapsa pomonella, Stgr. & Wk. Cat. Lp. Eur. 258. No, 1181
(1871).
Hab. Europe. ArricaA. Asta. N. & S. AMERICA.
AustraLiA. New Zeauanp. Astatic TuRKEY—Hares—
Aintab, 10 V. 1892 (Native Coll.). KaAsHmtr—Dras Ladak,
7000 feet, 20 VI. 1887 (Leech).
1182. Cydia grossana, Uw.
Carpocapsa grossana, Ster. & Wk. Cat. Lp. Eur. 258. No. 1182
(1871).
Hab. Evrorve. Asiatic TuRKEY—Hazrezs —Shar Devesay
(Native Coll. 1893). JAPAN (Pryer, 1886).
PaMMENE, Hb.
1186 (1). Pammene nannodes, sp. n.
Antenne pale cinereous. Palpt, head, and thorax whitish
cinereous, dusted with fuscous. ore wings whitish cinereous,
with a minutely speckled or transversely strigulated appear-
ance; the markings are greyish fuscous and consist of a
series of small costal spots, more conspicuous beyond than
before the middle ; an oblique dorsal streak before the middle
and an upright dorsal patch before the tornus, with a terminal
436 Lord Walsingham on Asiatic Tortricide.
patch, the lower portion of which is enclosed by silvery
streaks tending to converge downward; cilia whitish. cine-
reous, with a dark fuscous line running through them near
their base. Hp. al. barely 10mm. Hind wings white, with
a brown shade across the apical third and around the termen
and dorsum ; cilia white, with a brownish shade along their
base. Abdomen greyish. Legs whitish.
Type, & (60147) Mus. Wlsm.
Hab. SyriA—Beyrout, III. 1886 (Pratt). PALESTINE
(Tristram). Four specimens.
A small but distinct species having the appearance of a
dwarfed Laspeyresta succedana, Schiff.
1192. Pammene amygdalana, Dp.
Phthoroblastis costipunctana, Hw.+-amygdalana, Stgr. & Wk. Cat. Lp.
Eur. 259. No. 1191 a (1871). Grapholitha (Semasia) Lobarzewski,
Ster. & Wk. Cat. Lp. Eur. 259, 425. No. 1154 (1871). Phthoro-
blastis Lobarzewskit, Steger. & Wk. Cat. Lp. Eur. 259. No. 1192
(1871).
Hab. Evrore. Asiatic TURKEY—Hazrez—Shar Devesy,
10 VII. 1890 (Native Coll.).
1194. Pammene Juliana, Crt.
Phthoroblastis Juliana, Stgr. & Wk. Cat. Lp. Eur. 259, No. 1194
(1871).
Hab, Evrorr. Asiatic TuRKEY—/azes—Shar Devesy
(Native Coll. 1890).
1194 (2). Pammene (?) griseana, sp. n.
Antenne, palpi, head, and thorax greyish fuscous. Fore
wings grey, mottled with greyish fuscous; with an oblique
whitish patch on the middle of the dorsum striated with
greyish fuscous ; a whitish ocelloid patch above the tornus,
outwardly bounded with shining grey and containing some
transverse black streaks; on the costa are five or six pairs of
slender white geminations, the outer pair extending nearly
to a slender small white spot on the termen below the apex ;
a narrow dark fuscous line runs along the termen; cilia
greyish fuscous, a dark line along their base, and a small
whitish patch on the slight indentation below the apex. Ezp.
al. 18mm. Hind wings greyish brown; cilia pale whitish
cinereous, with a greyish brown line along their base. Ab-
domen greyish brown. Legs pale brownish cinereous, hind
tarsi banded with greyish fuscous.
Lord Walsingham on Asiatic Tortricidae. 437
Type, & (70177) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886). One specimen.
This species, almost certainly a Pammene, differs from
nimbana, H.-S8., in the more oblique and more mottled dorsal
patch, and in the more distinct blackish lines crossing the
ocelloid patch, which is less clearly margined on both sides
with lustrous steel-grey lines than in némbana; in other
respects the markings of the wing are similar and the shape
corresponds.
1207. Pammene nitidana, F.
Grapholitha (Semasia) nitidana, Steger. & Wk. Cat. Lp. Eur. 257.
No. 1160 (1871). Phthorodlastis fexana, Stgr. & Wk. Cat. Lp. Eur.
260. No. 1207 (1871)?. Stigmonota nitidana, Brt. Ent. Mo. Mag.
XII. 7-8 (1875) °. Phthoroblastis flexana, Stgr. Hor. Soc. Ent. Ross.
XV. 261 (1879)*. Pammene nitidana, Rag. Ann. Soc. Ent. Fr.
LXIII, 217. No. 1160, 220. No, 1207 (1894)*; Meyr. HB. Br. Lp.
506 (1895) °.
Hab. Europe. Asiatic TurRKEY—Kavpavewprcrar—
Brussa*. JAPAN (Pryer, 1886).
Barrett * and Meyrick*® adopted the synonymy flexana, Z.
=wetrana, Dgl., while Ragonot’ wrote nitedana, F.=flew-
ana, Z.
Zeller, Stett. ent. Ztg. X. 281-2 (1849), described flewana,
Z. in litt. (rotundana, Koll. in litt.), giving its habitat as
Danzig; Sweden (Bohemann) ; Riesengebirge (Standfuss) ;
and Pisa lV.-V. (Mann). The two latter specimens, labelled
respectively “ Hlexana, Z. 1 Ex. v. Bohemann, 1847. 21”
and ‘ rotundana, 'Tosc. FR. 638,” are the only types still in
the Zeller Collection. These are both nitidana, F., and
therefore confirm Ragonot’s determination of flerana, Z.
TmMeETOocERA, Ld.
1210. Tmetocera ocellana, Schiff.
Tmetocera ocellana, Stgr. & Wk. Cat. Lp. Eur. 260. No. 1210 (1871) ';
Fern. Tr. Am. Ent. Soc. X. 48. No. 349 (1882) 2.
Hab. Europe’. JApan (Pryer, 1886). UNnitTEep STATES’.
CANADA.
1210 (2). ZLmcetocera prognathana, Snell.
Grapholitha (Tmetocera) prognathana, Snell, Tijd. v. Ent. XXVI.
227-8, Pl. XIIL 8, 8a, 8d (1883) '.
Hab. Amur, 19 VII. 1877 (Hedemann)—Chingan Mts.,
20-22 VII. 1877°; Pompejefka, 9 VII. 1876 (Christoph) ;
Raddefka, 25 VII. 1876 (Christoph). JAPAN (Pryer, 1886).
438 Lord Walsingham on Asiatic Tortricide.
Gypsonoma, Meyr.
1212. Gypsonoma dealbana, Fri).
Steganoptycha incarnana, Stgr. & Wk. Cat. Lp. Eur. 260, No. 1212
(1871). Steganoptycha alnetana, Stgr. & Wk. Cat. Lp. Eur. 260.
No. 1214 (1871)*. Steganoptycha reconditana, Ster, & Wk. Cat. Lp.
Eur. 260, No. 1215 (1871) °. Steganoptycha incarnana, Fern. Tr.
Am. Ent. Soc. X. 46. No. 337 (1882) +; Rag. Ann. Soe. Ent. Fr,
LXIIT. 221, Nos. 1214-15 (1894)°. Gypsonoma dealbana, Meyr.
HB. Br. Lp. 481-2 (1895) ®.
Hab. Evrore’*. Cutna—Chang Yang, 4000-6000 feet
(Pratt, 1886). UNirep STATEs*.
Haworth, Lp. Br. 485 (1812), described Tortrix incarnana,
thinking it probably the same as “ Tortrix incarnana, Hiib.
Schmet. Tort. 30. 191?” He had no intention to bestow a
new name on his insect, but misquoted Hiibner’s name
“ incarnatana,”’ Pl. XXX. 191. Haworth’s name can only
be regarded as a homonym erroneous in adoption (and quota-
tion), and dealbana, Frél., should be used for this species.
1224. Hnarmonia diniana, Gn.
Sphaleroptera diniana, Stgr. & Wk. Cat. Lp. Eur. 241. No. 788
(1871) '. Steganoptycha pinicolana, Stgr. & Wk. Cat. Lp. Eur. 261.
No. 1224 (1871) 2; Fern. Tr. Am. Ent. Soc. X. 47. No. 846 (1882) *.
Steganoptycha diniana, Rag. Ann.Soe. Ent. Fr, LXIII. 185. No. (+) 787,
({788) 221, No, 1224 (1894) *.
Hab. Evrore®. SIBertA*. JAPAN — Howpo — Oiwake
(Pryer, 1885). Unirep STAaTEs*.
1225 (2). Enarmonia infausta, sp. n.
Antenne simple; brownish fuscous. Palp¢ brownish
fuscous. Head and thorax dark brown, with minute pale
brownish cinereous speckling. Fore wings dark brown,
minutely speckled with pale brownish cinereous, and mottled
with dark brownish fuscous, which is specially noticeable in a
costal patch a little beyond the middle, in another before the
apex, and in a third at the apex, the first of which is pre-
ceded and followed by two or more smaller ones; all these
dark spots are narrowly separated on the costa by whitish
ochreous; cilia brownish, tipped with whitish ochreous.
Exp. al. 15 mm. Hind wings dark tawny brown; cilia
greyish brown, tipped with whitish ochreous. Abdomen and
legs dark brown.
Type, & (60759) Mus. Wism.
Hab. JAPAN— Yusso—Hakodate, VI-VII. 1887 (Native
Coll.). Unique.
An obscure but distinct species.
Lord Walsingham on Asiatic Tortricide. 439
1227 (1). Enarmonia cristata, sp. n.
Antenne cinereous, faintly annulated. Palpz ferruginous.
Head thickly crested above; bright ferruginous. Thorax
fulvous, touched with silvery grey posteriorly. Fore wings
narrow, elongate, termen oblique, not sinuate, costa gently
arched; silvery grey, with a basal patch extending to one-
third, fulvous, with a slight silvery-grey sheen; beyond the
middle is a ferruginous patch narrowly touching the costa on
its inner side and extending to the dorsum beneath, but
reaching outwards above its middle and almost connected
with two spots of the same colour, one within and one at the
apex ; costa mottled with ferruginous and fulvous scales
beyond the middle, and a few ferruginous and fulvous scales
are scattered about the termen; cilia silvery grey, speckled
with pale grey. Exp. al, 13:5 mm. Hind wings pale
brownish ; cilia brownish grey. Abdomen [missing]. Legs
greyish cinereous.
Type, (70174) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886). ‘Two specimens.
ANCYLIS, Hb.
1250 (1). Ancylis partitana, Chr.
Phoxopteryx partitana, Chr. Bull. Soc. Imp. Nat. Mose. LVI. 430.
No. 130 (1882)?: sep. 193 (1882) *.
Hab. AMur*—Raddefka, 17 V. 1876 (Christoph) ; Wladi-
wostok, 30 V. 1877 (Christoph). Japan (Pryer, 1886)—
Yusso—Hakodate, VI-VII. 1877 (Native Coll.).
1250 (2). Ancylis pulchra, Btl.
Phoxopteryx pulchra, Btl, Il. Typ. Lp. Het. B. M. III. 79-80, Pl. LX.
7 (1879) }.
Hab. JAPAN (Pryer, 1886)—Hownpo— Yokohama’.
1256. Ancylis biarcuana, Stph.
Phoxopteryx biarcuana, Stgr. & Wk. Cat. Lp. Eur. 262. No. 1256
(1871)'. Phoxopterts biarcuana, Fern. Tr. Am. Ent. Soc, X. 51.
No. 373 (1882) *.
Hab. Evrore’. AmMur—Raddetka, 17 V. 1876 (Christoph).
JAPAN (Pryer, 1886). Unirep States—California ’*.
1258 (1). Ancylis latipennis, sp. n.
Antenne greyish fuscous. JPalpi thickly and roughly
clothed below, terminal joint short; greyish fuscous on their
440 Lord Walsingham on Aszatic Tortricidae.
outer sides, the end of the median joint and the terminal
joint whitish cinereous. Head greyish fuscous above, face
whitish cinereous. Thorax brownish grey. Fore wings
rather wide, with the costa very slightly arched, the apex but
slightly faleate; shining pale brownish grey, slightly striated
and blotched with umber-brown and brownish fuscous ; across
the basal half of the wing are some very indistinct dark
brownish strize; a small brownish fuscous patch on the dorsum
at one-third and some brownish fuscous dots along the costa ;
at about the middle of the costa is an oblique fasciaform
umber-brown band, somewhat widened on reaching the discal
cell and becoming less distinct and narrower towards the
dorsum, which it attains before the tornus; at the end of the
cel] this band sends out obliquely upwards towards the apex
a dark brownish fuscous pointed projection ; on the outer half
of the costa are five pairs of whitish geminated streaks, sepa-
rated and divided by umber-brown lines; the apex is umber-
brown, and a slender umber-brown line runs along the
termen to below its middle; cilia greyish, shaded with brown
and touched with brownish fuscous at the apex. zp. al.
155 mm. Hind wings brown; cilia cinereous, shaded with
brown. Abdomen brown, anal tuft slightly ochreous. Legs
cinereous, shaded with brown.
Type, 2 (70486) Mus. Wlsm.
Hab. JAPAN (Pryer, 1886). Unique.
In this species the wings are somewhat broader and shorter
than is usual in this genus; it perhaps somewhat approaches
unguicella, L., in the form of the markings, but the basal
patch is confined to a small dorsal spot.
1264 (2). Ancylis mandarinana, sp. n.
Antenne cinereous. Palpi, head, and thorax white. Fore
wings white, shading to grey beyond the middle; a tawny
chestnut-brown patch on the dorsum widens outward from
the base, forming an obtuse angle above the fold and sloping
thence with slightly convex outer margin to a little beyond
the middle of the dorsum, its outer edge clearly defined by a
narrow, rather shining, white line ; beyond and above it is a
large triangular costal patch of the same colour, extending
from the middle of the costa to the apex, containing one long
and four short outwardly oblique shining grey costal streaks,
the long one forming an acute angle with a similar line
bounding the outer side of the triangle and enclosing slender
black lines; the upper part of the triangular patch fades to a
greyish tinge before the anteapical costal streak; a whitish
Lord Walsingham on Asiatic Tortricida. 441
line along the termen precedes the greyish cinereous cilia.
Exp. al. 14mm. Hind wings and cilia pale brownish cine-
reous. Abdomen brownish grey. Legs white, with blackish
annulations on the hind tarsi.
Type, & (60352) Mus. W1sm.
Hab, K. CutnA—Ningpoo, LV. 1886 (Leech). AmMur—
Anossowo, 15 VI. 1877 (Hedemann). JAPAN—Yeazsso (Pryer,
1882). Three specimens.
This is distinguished from burgessana, Z., by its paler hind
wings, the paler dorsal patch, and by the pale space above the
tornus, the costal triangle not extending to the tornus. It is
nearly allied to daciniana, Z., and lundana, F.
Ruoposota, Ld.
1268. Rhopobota nevana, Hb.
Rhopobota nevana, Stgr. & Wk, Cat. Lp. Eur. 263. No. 1268 (1871) '.
Hab. EKurore'’, JApan (Pryer, 1886). InptA—Dharm-
sala (Hocking) ; Ootacamund, I. 1884 (Minchin).
CROCIDOSEMA, Z.
1269. Croctdosema plebeiana, Z.
Crocidosema plebetana, Steyr. & Wk. Cat. Lp. Eur. 263. No. 1269 (1871) ';
Wlsm, Pr. Z. Soc. Lond. 1897, 127. No. 174 (1897) ?.
Hab. Europe’. PALESTINE (Tristram). HAWAIIA
(Blackburn). AUSTRALIA*, West Inpies*. BraziL’.
PERU*®. ARGENTINA’.
1269 (1). Crocidosema demutata, sp. n.
Antenne (3) thickened, not ciliate; shining white, basal
joint dull white. Palpi white, touched with greyish at the
middle of the median joint externally. Headwhite. Thorax
evenly mottled with white and grey. Fore wings (3) with
a costal fold from the base; basal patch whitish ochreous,
costa and dorsum mottled with grey, and with some olive-
brown on its lower half; beyond the basal patch is a strong
grey shade, faintly reticulated with white and fuscous, and
continued from the costa to a triangular patch, on the dorsum,
consisting of a number of more or less connected white spots,
varied with grey on the extreme margin; beyond this dorsal
patch the outer part of the wing is stained with olive-green ;
there is a large circular patch of fuscous white below the
commencement of the outer third of the costa, into which two
pairs of geminated white streaks run from the margin above
442 Lord Walsingham on Asiatic Tortricidae.
it; these are preceded and followed on each side by a pair of
similar costal streaks, the last pair before the apex sending
out a leaden-grey line running within the termen nearly to
the tornus, and thence reverting towards the white patch ;
the olive-green on the apical half of the wing is mottled with -
dark fuscous, especially around the lower margins of the white
subcostal patch ; cilia greyish fuscous, with pale parallel lines
running through them along the margin and with two whitish
interruptions above the middle. zp. al. 18 mm. Hind
wings pale brown; with a tuft of long, loose, hair-like scales
arising from the median vein and from the space above and
below it at its basal half (not upstanding as in plebedana, Z.,
but combed backwards towards the limbus) ; cilia very pale
cinereous, a dark line running through them near their base.
Abdomen pale greyish brown. Legs pale greyish, hind tarsal
joints slightly spotted.
Type, 3 (70068) Mus. Wlsm.
Hab. JAPAN—Yesso (Pryer, 1882). Unique.
The fore wings are somewhat less elongate than in plebeiana,
Z., but the form of the antenne and the tuft on the hind
wings, although not quite brush-like, as in that species, sepa-
rate it from all other described genera.
LivoptycHa, Id.
1299. Lipoptycha plumbana, Se.
Dichrorampha (Iipoptycha) plumbana, Stgr. & Wk. Cat. Lp. Eur.
264. No. 1299 (1871)*. Diehrorampha plumbana, Fern. Tr. Am.
Ent. Soc. X. 55. No. 503 (1882) ?.
Hab. Europe’. UNITED StaTes—Cazzornr4*. ASIATIC
TurKeEY—Hazrzes—Shar Devesy (Native Coll. 1893).
ADDENDA.
Arcuips, Hb.
692 (4). Archips tnornatus, sp. n.
Antenne brownish cinereous. Palp? not extending beyond
the head; pale brownish ochreous. Head fawn-ochreous.
Thorax reddish fawn-ochreous. Fore wings reddish fawn-
ochreous, with a faint costal indication of a slightly darker
band of the same colour diffused obliquely outward toward
the tornus, and followed by another slight costal shade before
Lord Walsingham on Asiatic Tortricide. 443
the apex ; cilia pale ochreous; the costal fold is not closely
appressed. Hap. al. 283mm. Hind wings brownish grey ;
cilia paler grey. Abdomen and legs brownish grey.
Type, 8 (60073) Mus. Wlsm.
Hab. Cor—EA—Gensan, VI. 1886 (Leech). Unique.
This species appears to be allied to Archips Lafauryanus,
Rag., but is of a richer ochreous colour and rather larger.
692 (4). Archips luticostanus, Chr.
n. syn.=gigantana ( Bang-Haas MS.), Kennel.
Tortrix luticostana, Chr. Hor. Soc. Ent. Ross. XXII. 311 (1888) '.
Tortrix (Heterognomon) gigantana, Kennel, Ent. Zts. Iris, XL. 6-7,
No. 4, Pl. I. 4 (1899) *.
Hub. E,. StpER1A— Wladiwostok, VII.*; Amur’.
Tortrix, L.
725 (2). Tortrix trigonana, sp. n.
Antenne brownish grey. Palpi, head, and thorax buff-
brown. Fore wings with the costa evenly arched, apex obtuse,
termen oblique, slightly convex; buff-brown, with some
greyish fuscous shading toward the termen and a series of
obscure greyish fuscous spots along the costa and along the
base of the brownish ochreous terminal and apical cilia; a
triangular dark fuscous spot lies a little beyond the middle
of the fold; above it is a small reduplicated spot of the same
colour at the end of the cell, between this and the termen,
to which they are parallel, a line of dark fuscous dots margined
with ferruginous brown (in some specimens the markings
fade almost to obliteration). Hp. al. 20 mm. Hind wings
and cilia brownish, the latter fading to brownish cinereous
outwardly, and with a slender pale line along their base.
Abdomen and legs brownish grey.
Type, & (70551) Mus. Wlsm.
Hab. JapAN—Howpo—Gity (Pryer, 1886). Two speci-
mens.
PuHarmacis, Hb.
854 (1). Pharmacis magister, sp. n.
Antenne greyish. Palpi dark grey, tinged with ferrugi-
nous externally. Head and thorax brownish ochreous. Fore
wings rich brownish ochreous, with ferruginous markings and
shining leaden-grey cross-streaks and wavy lines; after a
faint ferruginous shade, accompanied above and beyond by
444 Lord Walsingham on Asiatic Tortricide.
leaden grey on the middle of the base, is a reduplicated patch
on the middle of the dorsum, outlined before and behind with
leaden grey, the inner line produced backward obliquely to
the costa; before the tornus is a similar ferruginous dorsal
patch, also outlined with leaden grey, the inner line furcate
from above the fold and enclosing between its two branches
which reach the costa a triangular ferruginous medio-costal
patch, the outer line also produced upward to the costa, where
it forms the inner margin of an anteapical ferruginous costal
patch; the apical and terminal area is somewhat speckled
with leaden grey, a reduplicated slender ferruginous marginal
line running at the base of the shining coppery ochreous cilia
(the first costal and the second dorsal patches are sometimes
obliquely and narrowly confluent, but they send out no out-
ward projection toward the apex, as in aleella, Schulze).
Exp. al. 22 mm. Hind wings tawny brownish grey, with a
shade-line near the base of the whitish cilia. Abdomen
brownish grey. Legs whitish.
Type, & (60856) Mus. Wlsm.
Hab. Asiatic TurKEY—J/azes—Shar Devesy, 18 VI.
1890 (Native Coll.). ‘Three specimens.
This species has the appearance of a very large form of
aleella, Schulze, with which it agrees in neuration, but which
is placed by Meyrick, wrongly as I think, in the genus
Phalonia, Hb.
Loxopera, Stph.
}Lozoprra, Stph. }fLOxopERA, Scudder.
862 (3). Lowopera ferruginea, sp. n.
Antenne cinereous. Palpi [broken]. Head and thorax
ochreous. ore wings pale ochreous, profusely sprinkled with
bright orange-ochreous, and with two oblique bright ferrugi-
nous transverse fasciz, much diffused at their edges; the
first runs from the middle of the costa to the basal half of
the dorsum, somewhat dilated at its extremities, especially
toward the costa, and attenuated on the fold; the second
arallel with the first and attenuated at its middle beyond the
end of the cell; there is a narrow ferruginous shade along the
costa from the base to the first fascia; cilia pale ochreous.
Exp. al. 21 mm. Hind wings rather dark grey ; cilia paler.
Abdomen dark grey. Legs pale cinereous. |
Type, 2 (61588) Mus. Wlsm.
Hab. Asiatic ‘TurRKEY—Hazzs—Shar Devesy (Native
Coll. 1893). Unique.
Lord Walsingham on Asvatic Tortricide. 445
This species has much the appearance of sangudnana, Tr.,
but the markings are more blurred and veins 7 and 8 of the
fore wings are stalked.
PuatoniA, Hb.
866 (1). Phalonia eestiva, sp. n.
Antenne white. Palpi white, the median joint shaded
with brown externally, the terminal joint fuscous. Head and
thorax white, the latter shaded with light brownish ochreous.
Fore wings white, with light brownish ochreous markings ;
three equidistant costal spots with ferruginous dots between
them, one before, one at, and one beyond the middle; an
oblique medio-dorsal blotch pointing outward and reaching to
the middle of the cell, below which it is margined on both
sides with silvery grey and some fuscous scales; a fainter
blotch before the tornus, less distinctly margined, and some
pale ochreous spots on the terminal and apical area accom-
panied by silvery scaling; cilia whitish ochreous, dusted with
fuscous. Hep. al.l8mm. Hind wings brownish grey; cilia
whitish, with a shade-line running through them. Addomen
brownish grey. Legs white, the tarsi dusted with fuscous.
Type, ¢ (60865); & (60866) Mus. Wlsm.
Hab. Astatic ‘TurKEY—Jl/4zzn—Shar Devesy (Native
Coll. 1890). Three specimens.
872. Phalonia Richteriana, F. R.
Cochylis Richteriana, Stgr. & Wk. Cat. Lp. Eur. 245. No. 872 (1871).
Hab. Evrore. JAPAN—Yzsso (Pryer, 1882).
HyYsTerosiA, Stph.
= TIprocraputs, Ld.
Stephens, who had originally placed cnopiana, Hw., in the
Phaloniad genus Xanthosetia, Stph., seems to have recognized
its divergence from hamana, L., and its allies in proposing
the genus Hysterosia for its reception. Wilkinson, finding a
costal fold in the male, placed it in the genus Halonota, Stph.
(Olethreutine). In the same year (1859) Lederer charac-
terized Idiographis, of which he made dnopiana the type,
placing it as a subgenus under Tortriz, Tr.; he was followed
by von Heinemann and Staudinger & Wocke. Walsingham,
in 1879, placed the genus Jdiographis between Retinia, Gn.,
and Conchylis, Tr., recognizing the structural character
“vein 2 of the fore wings arising from the outer third of the
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 30
446 Lord Walsingham on Asiatic Tortricide.
cell,”” but relying on the costal fold for its separation from
both. Fernald, who in 1882 instituted the subfamily Con-
chyline, started with [diographis preceding Conchylis.
The genus Phtheochroa, Stph. (type rugosana, Hb.), has
been rightly recognized as belonging to this subfamily, but to
this genus have been added two species which differ from it
in the possession of a costal fold—Duponcheliana, Dp., and
amandana, H.-S. I am unable to refer them to any other
genus than Hysterosia. Meyrick (HB. Br. Lp. 554-5) placed
the smaller species maculosana, Hw., by itself in the genus
Eupecilia as having a costal fold and thoracic tuft, vein 7
of the fore wings running to the apex. I find that in macu-
losana vein 7 distinctly runs to the termen, which justifies its
inclusion in the genus Hysterosia, the type of which, ¢nopiana,
Hw., does possess a posterior thoracic tuft, although Meyrick
unaccountably failed to observe it and described the thorax as
smooth. In any case the name Kupecilia, Stph., is inad-
missible for maculosana, Hw., Westwood in 1840 having
cited angustana, Hb., as its exponent, thereby rendering
Eupecilia a synonym of Huxanthis, Hb., Meyr.
909 (1). Hysterosia syriaca, sp. n.
Antenne pale brownish, whiter towards the base. Palpi
porrect, extending twice the length of the head beyond it;
white. Head and thorax white, witha brownish tinge. Fore
wings white, with a few silvery scales scattered around the
edges of the darker markings; these are of two shades of
brown and consist of a streak along the costa from the base,
covering the narrow costal fold and joining the upper extre-
mity of a broad transverse brown band, slightly indented at
the middle on its inner side and adorned with four or five
tufts of raised brown scales; three costal spots beyond the
middle, the outer two merging in a transverse narrow brown
band reaching to the tornus and scarcely separated from a
dorsal blotch before the tornus ascending to the upper angle
of the cell; two black spots mark the end of the cell and
along the termen is a chestnut-brown shade, broken into a
series of small marginal patches; cilia white, mottled with
brownish grey, a darker shade running along their base. zp.
al. 22-28 mm. Sind wings greyish brown; cilia whitish,
with a dark line near their base. Abdomen greyish brown,
with white cross-bands. Legs dirty whitish, peppered with
brown scales.
Type, 3 (60832) Mus. Wlsm.
Hab. Astatic TuRKEY—Hazrzes—Shar Devesy, 23-27 V.
1890 (Native Coll.). Five specimens.
Lord Walsingham on Asiatic Tortricide. 447
This species is closely allied to Hysterosia Duponceliana,
Dp., but is separable through its larger size and the broader
patch of white before the apex and much paler cilia.
909 (2). Hysterosia decipiens, sp. n.
Antenne pale brownish grey. Palpi projecting more than
twice the length of the head beyond it; white. Head and
face white. Zhorax brownish grey, with some black scales.
Fore wings: 2 stone-white, mottled with brownish grey,
the outlines of the patches narrowly and irregularly marked
with black scaling ; a patch of black scales surrounding a
ferruginous spot rests on the middle of the fold, extending
upward to the anterior margin of the cell and downward
nearly to the dorsum; the brownish grey mottling is specially
noticed in a basal patch, a small costal patch about the middle,
and in a patch above the tornus, followed by an outwardly
curved transverse band before the apex, ending below the
middle of the termen ; cilia stone-white, mottled with brownish
grey, and a few blackish scales: @ with a narrow darkened
costal fold. Kap. al. 16mm. Hind wings pale brownish
grey ; cilia pale cinereous. Abdomen brownish grey. Legs
whitish.
Type, 2 (61601); ¢ (60874) Mus. Wlsm.
Hab. Asiatic ‘TurKEY—fazzen—Shar Devesy (Native
Coll. 1890, 1893). Four specimens.
910 (1). Hysterosia coreana, sp. n.
Antenne white, with a brownish tinge. Palpi white, the
median joint shaded with brown externally nearly to the apex.
Head and face white. Thorax black. Fore wings white,
with some silvery scales surrounding pale olivaceous grey
mottling on the outer half and a black transverse fascia a
little before the middle; a short black basal patch, not ex-
tending to the dorsum, sends forth a streak of black mottling
along the costa, including the narrow costal fold, and.reaching
the black fascia; this contains some steel-blue scaling and is
slightly bowed outward externally ; it is followed by a small
black triangular costal spot, with which it is almost con-
nected ; a slight costal shade before the apex is followed by a
black terminal band including the apex and upper two-thirds
of the termen and mixed with ferruginous scales; cilia
whitish, with brown and black mottling on their outer half,
a blackish line running through them near their base. ep.
al. 18mm. Hind wings brownish grey; cilia whitish, with
30*
448 Mr. E. E. Green on some
a reduplicated shade-line running through them. Abdomen
brownish grey, white posteriorly. Legs whitish, with some
black bands.
Type, & (60364) Mus. Wlsm.
Hab. Corva—Gensan, VI. 1886 (Leech). Unique.
In appearance this species is intermediate between /Hyste-
rosia amandana, H.-S., which has a white thorax and a costal
fold, and Phtheochroa pulvillana, H.-S., which has a black
thorax and no costal fold; except by its black thorax, it
is also undistinguishable from the Japanese albiscutellum,
Wism., of which unfortunately the male is not known, but
this is a much larger species than amandana.
LVII.—Descriptions of some Species of Coccidee collected by
Mr. James Lidgett in Victoria, Australia. By K. ERnust
GREEN, F.E.S., Government Entomologist, Ceylon.
[Plate XI.]
Mr. EK. E. Green has sent to the British Museum some
specimens of Coccide new to the collection, and with them
the following descriptions and figures, which I have much
pleasure in sending to the ‘ Annals’ for publication.
Cuas. O. WATERHOUSE, V.P.E.S.
Mytilaspis indentata, sp.n. (Pl. XI. fig. 1.)
Female puparium brownish straw-colour, the sides and
hinder part darker, with a very narrow whitish margin.
First pellicle very pale yellow. Second pellicle almost
. colourless. Strongly convex above. LHlongate, rather nar-
row, but widening slightly behind the exuvie ; usually much
curved or contorted, as in J. intermedia, Mask. (Trans. N. Z.
Inst. 1890, p. 7). Ventral scale moderately developed.
Length about 2 millim.
Male puparium not observed.
Adult female of usual elongate form, broadest across the
basal abdominal segments. Antenna consisting of the usual
small tubercle and curved bristle. Mouth-parts large and
conspicuous. Parastigmatic glands absent or represented
only by a single pore at the anterior spiracles. Abdominal
segments with groups of oval pores and short cylindrical ducts
on the lateral margins. Pygidium (Pl. XI. fig. 1) deltoid,
Coccide from Australia. 449
the margin deeply indented at the junctions of the suppressed
segments, these indentations made more conspicuous by a
thick chitinous rim. Median lobes minute and incon-
spicuous. First lateral lobes undivided, large and broad but
not very prominent, the free margin evenly curved, the sides
straight. Other lobes obsolete. Margin on each side
strongly chitinous and serrated. There are no squames and
only two small spines are noticeable on each side imme-
diately laterad of the large second lobes. No circumgenital
glands. Anal at about same level as genital aperture, near
centre of pygidium. Dorsal pores few, but strongly marked
by thickened chitinous rims.
Total length of insect averaging (50 millim.
Hab. On an unidentified plant collected at Werribee
Gorge, Bacchus Marsh, Victoria, Australia. (Lidgett Coll.
no. 47.)
A very distinct species, easily recognizable by the minute
median lobes followed by the large simple lateral lobes, and
by the absence of circumgenital glands. The only other
Australian species of Myétlaspis with this character are
M. conveva, Mask., and M. drymidis, Mask. In convewa the
puparium is greyish white and the second pellicle is so much
raised that its posterior edge forms a conspicuous ridge over
the secreted portion. The pygidium is also said to have
broad median lobes, though (from an examination of the
figure) it seems possible that these may really represent the
lateral lobes as in indentata. M. drymidis is easily distin-
guished by the remarkable fringed tubular processes on the
margin.
Ctenochiton (?) araucarie, sp.n. (PI. XI. figs. 2, 2 a.)
Adult female oblong-oval; strongly convex above; naked,
or with a very imperfect coating of irregular waxy plates
which appear to be brittle and easily detachable. Colour of
dried examples dark chestnut-brown; the dorsal surface
strongly rugose, usually with a well-defined median longi-
tudinal ridge. Antenna (fig. 2) with eight joints, third
longest, 6, 7, and 8 very short, subequal. The terminal
joint bears several stout spines. Antennal formula—
3, 4, (2, 5), 1, (6, 7, 8). Legs stout; tarsus equal to about
two thirds length of tibia. Foot with four digitules, the
ungual broadly spatulate, the tarsals stoutish knobbed hairs.
Anal scales usually widely divergent; base shorter than
outer edge, which is rounded and bears a few truncate spines.
Anal ring with six stout hairs; the invaginated tube which
surrounds them strongly rugose. Margin closely set with
450 Mr. E. E. Green on some
moderately long stout spines, their extremities truncate.
Stigmatic cleft shallow, with three or four stout pointed
stigmatic spines, one or more of which are longer and curved
(fig. 2 a). Derm with small and rather obscure circular and
oval pores, more conspicuous towards the margin.. Nume-
rous small circular pores (circumgenital glands) surrounding
the genital aperture, and a group of similar pores (para-
stigmatic glands) below the stigmatic cleft.
Length 5 millim.; breadth 3-33 millim.
On the twigs bearing the adult females are some very
imperfect empty and crushed tests, which may represent either
the second stage of the female or the male puparia. They
are composed of granules or small irregular plates of trans-
parent waxy matter.
Hab, On Araucaria sp., Victoria, Australia. (Lidgett
Coll. no. 50.)
I have considerable difficulty in determining the proper
genus in which to place this species. The structural cha-
racters of the insect itself suggest Hrvochiton ; but the female
forms no felted or cottony covering—in fact, the adult female,
as submitted to me, is practically naked, bearing only scat-
tered fragmentary patches of glassy wax. The complete
marginal series of stout spines distinguishes it from Lecantum.
Although I can find no vestige “of a marginal fringe of
flattened waxy plates, I have provisionally attached this insect
to Maskell’s genus Ctenochiton, to which in other respects it
conforms fairly well. In Cten. depressus, Mask., the fringe
is said to be ‘ inconspicuous or sometimes absent” in the
adult female.
BIRCHIPPIA, gen. nov.
Test of adult female completely enclosing the insect, with
a small aperture at posterior extremity.
Male puparium with a well-defined circular valve at hinder
end, through which the adult insect makes its escape.
Adult female shrivelling to anterior extremity after gesta-
tion, the hinder part of the test being filled with the eggs.
Legs absent. Antenne more or less rudimentary. Mentum
monomerous. Stigmatic spines present. Posterior extremity
cleft. Anal aperture surrounded by chitinous plates, which
do not meet to form a valve as in typical Lecaniine. Anal
ring with ten hairs.
Adult male unknown.
Larva with 6-jointed antenne; the terminal joint long.
Legs normal. Anal plates as in adult; anal ring with six
Coccide from Australia. 451
hairs. Posterior extremity of body with a pair of long
caudal sete, not springing from the anal plates.
Having only a single species with which to deal, it is
difficult and inadvisable to define the generic characters very
minutely.
I have still further difficulty in determining the proper
systematic position of the new genus. It has some characters
(stigmatic spines, anal cleft, &c.) which associate it with the
Lecaniine, others which suggest Dactylopiine, but not such
*as would warrant its inclusion in the intermediate family
Hemicoccine, where the larve are Dactylopiinid and the
adults Lecaniinid. I think that the Lecanoid characters
have here the predominance, the principal differences being
the non-valvular aual plates and the marginal position of the
candal sete.
The name of the genus is taken from that of the locality
in which the specimens were collected.
Birchippia anomala, sp.n. (Pl. XI. figs. 3-38 d.)
Test of adult female corneous, semitransparent, fulvous,
more or less obscured by a fragmentary coating of brownish
waxy matter. The dark body of the dead insect can be
distinguished through the test at the anterior extremity.
Broadly oval to oblong-oval according to position. Strongly
convex above, smooth. A small circular opening at the
posterior extremity.
Length 3-4 millim.; breadth 2 millim.
Male puparium whitish, opaque. Rather broadly oval,
with a circular valve-like opening at posterior extremity.
Moderately convex above, more so above the abdominal area.
Surface marked by numerous transverse depressed lines and
usually four longitudinal furrows ; the area between the two
median furrows standing up asa rounded ridge. The general
character of the puparium 1s strongly suggestive of that of a
male Lecaniodiaspid.
Length 1:25 millim.; breadth 0°75 millim.
Adult female (fig. 3) shrivelling to anterior part of test,
subcircular (after maceration). Posterior extremity cleft.
Mentum monomerous. Antenna (fig. 3a) atrophied, varying
in development in different individuals. In some examples a
distinct basal and a compound terminal joint only can be
distinguished. In others at least three distinct joints are
present, the median as long as the other two combined. The
terminal joint has two or more imperfect divisions and bears
six or seven stout bristles at its extremity. On the margin,
452 On some Coccide from Australia.
opposite the stigmatic areas, are some stout dilated blunt
spines, flattened towards the ends (fig. 3), two (rarely three)
at anterior point, one (rarely two) at the other point, placed
on dorsal surface. On the ventral surface is an extended
group of ceriferous pores, directed towards the opening of the
spiracle. Derm with a large number of bituberculate spin-
nerets scattered over the surface and forming a more or less
continuous series on the margin. They are connected with
some remarkable tubular ducts, each terminating in a lateral
finger-like process. The connexion between the external
spinneret and the tubular duct is very indistinct; but it can
be made out by careful illumination (see figs. 3b and 3¢).
Anal aperture at base of cleft, surrounded dorsally by four
more or less confluent triangular chitinous plates (fig. 3c),
the lower pair largest, with two stout spines near the apex of
each; these plates do not meet to form a valve, as in typical
Lecaniine. Anal ring with ten stout flattened hairs pro-
jecting into the cleft. Margin of body with a scattered
series of short spines, which increase in size near the anal
cleft.
Diameter 1:50-1:75 millim.
Newly hatched larva oval. Antenna 6-jointed, sixth
longest ; extremity abruptly narrowed and truncate, bearing
a longish fine hair. Anterior stigmatic area with two, poste-
rior with one stout spine. Anal aperture (fig. 3 d) surrounded
by chitinous plates as in adult. A long stout seta on each
side of the cleft, not attached to the chitinous plates, but
springing from a minute tubercle on the margin of the body.
Hab. “ Qnasmall Leguminous shrub, collected at Birchip,
Victoria.” (Lidgett Coll. no. 49.)
The scales are thickly clustered on the twigs of the plant.
Peradeniya, Ceylon,
24th May, 1900,
EXPLANATION OF PLATE XI.
Tig. 1. Mytilaspis indentata. Pygidium.
Fig. 2. Ctenochiton araucarie, Antenna of female.
Fig. 2a. Ditto. Stigmatic area of adult female.
lig. 3. Birchippia anomala. Adult fema’e, ventral view.
Fig. 8a. Ditto. Antenna of adult female.
fig. 3b, Ditto. Anterior stigmatic area of adult female.
Fig. 38¢. Ditto. Extremity of abdomen, dorsal aspect.
Fig. 3d. Ditto. Posterior extremity of young larva.
On new Species of Japanese Land- Shells. 453
LVIII.—Descriptions of new Species of Japanese
Lend-Shells. By G. K. Guo, F.Z.8.
A FURTHER consignment of Helicoid land-shells received from
Mr. Hirase contains seven undescribed species.
Kaliella elata, sp. n.
Shell subperforate, elongate-conoid, thin, fragile, sub-
pellucid, corneous, under the microscope seen to be covered
with about ten spiral lire above the periphery, below minutely
spirally striated, shining. Spire elevated, apex obtuse, suture
slightly impressed. Whorls 6, compressed, increasing slowly ;
the last not descending in front, angulated at the periphery,
a little convex below, narrowly impressed around the axis,
Aperture a little oblique, subquadrate; peristome thin,
straight, acute; margins distant, the columellar margin
vertical, slightly reflected over the narrow perforation.
Four specimens.
Diam. 1°5, alt. 2 millim.
Hab, Kashima, prov. Harima, Type in my collection.
Kaliella crenulata, sp. n.
Shell imperforate, conoid, thin, fragile, subpellucid, pale
corneous, minutely ribbed above, shining and minutely striated
below. Spire conoid, apex obtuse, suture linear. Whorls 6,
flattened, increasing slowly, the last not descending; peri-
phery with a crenulate keel, convex below, narrowly im-
pressed round the axis. Aperture subvertical, subquadrate ;
peristome thin, straight, acute; margins distant, columellar
margin subvertical, slightly reflected above.
One specimen.
Diam. 2°75, alt. 3 millim.
Hab. Kashima, prov. Harima. Type in my collection.
Differs from its nearest ally K. acutangula, A. Ad., by the
flatter whorls, less convex base, and crenulated keel. It
is also nearly double the size, although having but half a
whorl in excess of its congener. Five specimens from
Kioto, previously referred to K. acutangula, prove to pertain
to this new species.
Kaliella pagoduloides, sp. n.
Shell imperforate, ovoid-conoid, thin, subpellucid, ochreous-
corneous ; under the microscope seen to be finely ribbed
454 Mr. G. K. Gude on new Species of
above, spirally wrinkled below. Spire convex-conoid, apex
obtuse, suture impressed. Whorls 53, rather tumid, in-
creasing slowly, the last not descending in front, convex
below, narrowly impressed round the axis. Aperture ver-
tical, truncate-lunate; peristome thin, straight; margins
distant, columellar margin vertical, slightly reflected above.
Two specimens.
Diam. 2, alt. 1°75 millim.
flab. Kashima, prov. Harima. Type in my collection.
Allied to K. stenogyra, but it has fewer whorls than that
species and is rounded at the periphery. In shape it much
resembles a very minute Ganesella pagodula, Ehrm.
Pyramidula (s. 8.) pretiosa, sp. n.
Shell umbilicate, discoid, corneous under an_olivaceous
cuticle, distantly costulate. Spire depressed, apex obtuse,
suture rather deep. Whorls 34, a little rounded, increasing
rather rapidly ; the last scarcely descending in front, angu-
lated at the periphery. Aperture roundly ovate; peristome
thin, straight, acute ; margins approaching, columellar margin
a little reflected over the narrow deep umbilicus.
Ten specimens.
Diam. 14, alt. 1 millim.
Hab, Fukura, Awaji Island. Type in my collection.
Allied to P. amblygona, Reinh., but much smaller.
Trishoplita Dacoste, sp. nu.
Shell deeply umbilicated, conoid, finely striated, obsoletely
decussated with microscopic spiral lines, perceptible only in
the depressions between the striz, corneous, pellucid, rather
solid, shining below, duller above. Spire depressed, apex
obtuse, suture deeply impressed. Whorls 5-53, rounded
above, a little flattened below, increasing rather rapidly ; the
last shortly deflected in front, rounded at the periphery,
widened towards the mouth and sloping into the umbilicus
behind the peristome. Aperture oblique, subrotundate ;
peristome scarcely thickened, slightly expanded all round,
the margins approaching, columellar margin a little reflected
over the deep but moderately wide umbilicus, which exhibits
the anterior half of the penultimate whorl.
T'wo specimens.
Diam. maj. 10°5-11'5, minor 9-9°5; alt. 7-8 millim.
Hab. Kagashima, prov. Satsuma. ‘Type in my collection.
Allied to 7. conospira, Mts., but more solid and with wider
Japanese Land- Shells. 455
umbilicus. The spiral sculpture is scarcely perceptible under
a strong lens. A specimen in the collection of Mr. 8. J.
Da Costa, and labelled by the late Dr. Hungerford “ Satsuma,
sp., Ikao, Nippon,” I also refer to this new species. It is a
little more shining and smaller than the type, measuring :—
Diam. maj. 9°5, minor 8; alt. 6 millim.
Ganesella tosana, sp. n.
Shell narrowly umbilicated, depressed- conoid, finely stri-
ated, decussated with fine spiral lines, thin, pellucid, a little
shining below, silky above, with a broad milky-white band
under the suture, the umbilical region more or less clouded
with milky white. Spire depressed, apex prominent, suture
impressed. Whorls 54-6, rounded, increasing rather rapidly,
the last widened towards the mouth, descending shortly in
front, obsoletely angulated at the periphery ; aperture oblique,
ovate-rotundate ; peristome thin, acute, margins convergent,
the lower expanded, the columellar margin reflected over the
narrow deep umbilicus.
Three specimens.
Diam. maj. 14, minor 12; alt. 9 millim.
Pee toe 8. Lk os) alt; O°: online
Hab. Ushirogawa, Tosa Island. Type in my collection.
The present species is allied to G. Hilgendorfi, Kob., but
it is thinner, more depressed, and with narrower umbilicus.
Eulota (Huhadra) grata, sp. n.
Shell sinistral, deeply umbilicated, conical; embryonal
whorls smooth, the others finely regularly ribbed, the inter-
stices minutely spirally striated; thin, glossy, subpellucid,
the earlier whorls pale corneous, the later ones straw-yellow.
Spire elevated, apex obtuse, suture margined. Whorls 64,
rounded, increasing somewhat rapidly, the last shortly de-
flected in front, angulated at the periphery, becoming rounded
and widened at the mouth, a little flattened below. Aperture
oblique, broadly lunate; peristome rather thin, white ; mar-
gins approaching, upper straight, acute, outer and lower
broadly expanded and reflected, columellar reflected over the
narrow deep umbilicus.
‘'wo specimens.
Diam. maj. 27, minor 24; alt. 21°5 millim.
Hab. Nishigo, prov. Uzen. ‘Type in my collection.
456 Mr. W. F. Kirby on the
Var. zonata, nov.
Ornamented with a wide, ill-defined, and more or less
interrupted ochreous-brown zone, reaching from near the
suture to below the periphery.
Hab. Same as the type.
This handsome new species belongs to the group of
E. quesita, Desh. It is, however, much smaller, much more
elevated in the spire, thinner in texture, more shining, and
has a narrower umbilicus. The dark zone of the variety is
evidently the result of the diffused colouring-matter of three
bands—one peripheral, one supra- and one infraperipheral—
which are faintly visible on holding the shell to the light.
LIX.—Notes on the Neuropterous Family Nemopteride.
By W. F. Kresy, F.L.S., F.E.S.
THIRTY-THREE species of this small but very interesting
family are enumerated in the present paper, of which the
Natural History Museum possesses fourteen, which T have
distinguished by an asterisk (*). They are remarkable for the
great length of the hind wings, which take the form of long
slender tails.
I have given the synonymy with tolerable completeness,
as the references are very scattered, and many of them occur
in old books not always easy of access.
It is somewhat remarkable that, as in the Ascalaphide,
the brightly coloured species are nearly all Mediterranean,
while those with hyaline fore wings mostly occur elsewhere.
Genus J. NEMOPTERA.
Nemoptera, Latr. Hist. Nat. Crust. Ins. iii. p. 296 (1802), xiii. p. 20
(1805) ; Olivier, Enc. Méth. viii. p. 173 (1811) ; Klug, Abhandl. Akad.
Berlin, 1836, p.92; Walker, List Neur. Ins. il. p. 470 (1858) ; Hagen,
Stett. ent. Zeit. xxvii. pp. 874, 451 (1866) ; Proc. Bost. Soc. Nat. Hist.
xxiii. p. 250 (1886). uel
Nemopteryx, Leach, Zool. Mise. ii. p. 74 (1815).
Physapus, Leach, Edinb. Encyel. ix. p. 187 (1815).
Nematoptera, Burm, Handb. Ent. ii. p. 984 (1840) ; Westwood, P. Z.S.
1841, p. 9; Ann. & Mag. Nat. Hist. viii. p. 376 (1842).
The types of this genus are Nemoptera coa, L., as figured
by Coquebert and Latreille. The latter figure has been
referred to N. bipennis, but seems to possess the distinctive
characters of NV. coa.
Neuropterous Family Nemopteride. 457
*1. Nemoptera sinuata.
Nemoptera sinuata, Oliv. Enc. Méth. viii. p. 178. n. 2 (1811); Ramb.
Ins. Névr. p. 885 (1842); Hagen, Proc, Boston Soc. Nat. Hist. xxiii.
p. 250 (1886),
|| Punorpa coa, Borkhausen, Scriba, Beitr. Ins. ii. p. 155, pl. xi. fig. 1
(1791).
Nemoptera coa, pt., Latr. Hist. Nat. Crust. Ins. xiii. p. 20 (1802) ;
pt., Kleg, Abh, Akad. Berl. 1836, p, 92. n. 1.
|| Nematoptera coa, pt., Burm. Handb. Ent. ii. p. 987. n. 9 (1840).
Nematoptera petiveri, Westw. P. Z.S. 1841, p. 9. n. 1; Ann, & Mag,
Nat. Hist. viii. p. 376. n. 1 (1842),
Hab. Turkey, Asia Minor, Syria.
This species differs from the others in the wings being
much less mottled.
*2, Nemoptera bipennis.
Panorpa bipennis, Mliger, Ahrens, Faun. Eur, fase. i. fig. 16 (1812).
Nematoptera bipennis, Westwood, Proc. Zool. Soc. Lond. 1841, p. 10.
n. 3; Ann, & Mag, Nat. Hist. viii. p. 377. n. 3 (1842).
Nemoptera bipennis, Hagen, Proc. Bost. Soc. Nat. Hist. xxiii. p. 253.
n. 4 (1888).
Nemopteryx lusitanica, Leach, Zool. Mise. i. p. 74, pl. Ixxxv. fig. 1
1815).
‘ie lusttanica, Klug, Abh. Akad. Berl. 1836, p. 93. n. 3;
Rambur, Faune Andal. ii. pl. ix. fig. 1 (1842); M‘Lachl. Proc. Ent.
Soc. Lond. 1886, p. lviil.
|| Nematoptera lusitanica, Burmeister, Handb. Ent. ii. p, 907. n. 8 (1840).
Hab. Spain, Portugal.
] have not seen Ahrens’s figure.
*3, Nemoptera egyptiaca.
, Savigny, Descr. Ins. Neur. pl. 11. fig. 15.
Nemoptera egyptiaca, Rambur, Ins. Névr. p. 834 (1842) ; [fagen, Proc.
Bost. Soc. Nat. Hist. xxiii. p. 252 (1886).
Nemoptera hebraica, Westwood, Thes. Ent. Oxon. p. 178, pl. xxiii. fig. 5
(1874).
Hab. Egypt, Syria, Palestine.
*4, Nemoptera coa.
Ephemera coa, Hasselquist, Iter Paleest. p. 423. n. 110 (1757).
Panorpa coa, Linné, Syst. Nat. (ed. x.) i. p. 552. n. 3 (1758); Coque-
bert, Icones, p. 15, pl. 111. fig. 3 (1799).
Nemoptera coa, Oliv. Enc. Méth., viii. p. 178. n.1 (1799) ; Walker, List
Neur. Ins. B. M. ii. p. 470. n. 1 (1858); Ramb. Ins. Névr. p. 333,
pl. iii. fig. 3 (1842) ; Hagen, Proc. Bost. Soc. Nat. Hist. xxiii. p. 253.
n. 3 (1880).
Nematoptera coa, pt., Burm, Abhandl. Akad, Berl. 1836, p. 92. n. 1.
Nematoptera Petiveri, pt., Westwood, Proc. Zool. Soc. Lond. 1841,
p. 9.n.1; Ann, & Mag. Nat. Hist. viii. p. 376. n. 1.
Nematoptera Coqueberti, Westwood, Ul. ce. p. 9. n. 2. p. 876. n. 2.
Hab. 8. Europe, N. Africa.
458 Mr. W. F. Kirby on the
Genus II. HALTER.
Halter, Ramb. Ins. Névr. p. 8385 (1842).
The time has hardly yet arrived for subdividing this genus,
to which we may provisionally refer all the transparent-winged
species with long beaks. The species are divisible into several
very distinct groups, most of which will probably hereafter
rank as genera.
Section 1.
Hind wings linear, with a long dark band before the extremity.
*5. Ilalter africana.
Nemopteryx africana, Leach, Zool. Mise. ii. p. 74, pl. Ixxxv. fig. 1
(1815).
Nemoptera africana, Gray, Griffith’s Anim. Kingd. xv. p. 324, pl. ev.
tig. 4; Westwood, Proc. Ent. Soc. Lond. i. p. xxv (1836); Walker,
List Neur. Ins. ii. p. 475. n. 14 (1853); Hagen, Proc. Bost. Soc. Nat.
Hist. xxiii. p. 262. n. 10 (1888).
Nematoptera africana, Westwood, Proc. Zool. Soc. Lond. 1841, p. 12.
n.9; Ann. & Mag. Nat. Hist. viii. p. 379. n. 9 (1842).
Nemoptera bacillaris, Klug, Abhandl. Akad. Berl. 1836, p. 95, fig. 2.
Nematoptera bacillaris, Westwood, ll. cc. p. 12. n. 10, p. 879. n. 10.
Nemoptera halterata, pt., Klug, l. ¢. p. 94. n. 8 (1886).
Nematoptera latipennis, Burm. Handb. Ent. 11. p, 986. n. 5 (1840).
Nematoptera latipennis, Westwood, JU. ce. p. 12. n. 11, p. 879. n. 11.
Hab. S. Africa.
The type is in the British Museum.
6. Halter remifera.
Nemoptera remifera, Westwood, Thes. Ent. Oxon. p. 179, pl. xxxiii.
fig. 9 (1874) ; Hagen, Proc. Boston Soc. Nat. Hist. xxiii. p. 265, n. 11
(1888).
Tab. Cape.
7. Halter costalis.
Nemoptera costalis, Westwood, Proc. Ent. Soc. Lond. i. p. Ixxv (1886) ;
Thes. Ent. Oxon. p. 179, pl. xxxiii. fig. 6 (1874).
Nematoptera costalis, Westw. Proc. Zool, Soc. Lond. 1841, p. 12. n.13;
Ann. & Mag. Nat. Hist. viii. p. 379. n. 15 (1842),
3. Nemoptera angulata, Westwood, Proc. Ent. Soc. Lond. i. p, Ixxv
(1836); Duncan, Jardine’s Nat. Libr., Intr. Ent. p. 293, pl. xxvii. fig. 3
(1840); Hagen, Proc. Bost. Soc. Nat. Hist. xxiii. p. 259. n. 9 (1888).
Nematoptera angulata, Westwood, Proc. Zool. Soc. Lond. 1841, p. 12.
n. 12; Ann. & Mag. Nat. Hist. viii. p. 879. n. 12 (1842).
Nemoptera angula, Walk. List Neur. Ins. ii. p. 475, n. 16 (1858).
Hab, Cape.
Neuropterous Family Nemopteridee. 459
As the name costalis stands before angulata in the Proc.
Ent. Soc., the former must stand for the species if the two
names are correctly placed together as sexes, as appears
probable.
8. [alter biremis.
Nemoptera biremis, Kolbe, SB. Ges. naturf, Fr. Berlin, 1900, p. 16.
Hab. Cape.
Section 2.
Hind wings more or less expanded towards the extremity (once, twice, or
thrice), and with a broad dark band towards the base of the expanded
part of the wings.
9. Halter albostigma.
Nemoptera albostigma, Westwood, Thes. Mnt. Oxon. p. 179, pl. xxxiil.
fig. 7 (1874).
Hab. Gululand.
*10. Halter halterata.
Panorpa halterata, Forskal, Descr. Anim, p. 97 (1775) ; Icones, pl. xxv.
fie. EK (1776).
Nemoptera halterata, De Selys, Ann. Soc. Ent. Belg. x. pp. 254, 255,
pl. ii. figs. 5, 6 (1887); Hagen, Proc. Boston Soc, Nat, Hist. xxiii.
p. 257. n. 8 (1887).
Nemoptera pallida, Olivier, Enc. Méth., viii. p. 179. n. 5 (1811) ; Ram-
bur, Ins, Névr. p. 385, pl. vii. fig. 4 (1842).
Nematoptera pallida, Westwood, Proc. Zool. Soc. Lond. 1841, p. 11.
n.8; Ann. & Mag. Nat. Hist. vi. p. 378. n. 8.
Nematoptera Forskalli, Westwood, Ji. cc. p. 11. n. 6, p. 378. n. 6.
Hab. Arabia, Baghdad.
This is the type of the genus, N. pallida, Olivier, being
the first of four species referred by Rambur to J/alter, and,
moreover, being figured. The specimen in the British
Museum is from Mount Sinai. Several other species have
been wrongly called N. halterata by various authors, whose
references | have not always quoted.
*11. Halter tipularia.
Nemoptera tipularia, Westwood, Thes. Ent, Oxon. p. 179, pl. xxxiii.
fic. 10 (1874).
Hab. Damaraland.
460 Mr. W. F. Kirby on the
*12. LHalter Bettont, sp. n.
Dimensions.—
3. 2.
Loneicorp.? aCe eee on 15 mm.
Exp Salant Sele ee ones AG? 5
Long. al//past: \. “se (eurea)o0o5 of
Body reddish brown, beak (except at the tip) and sides and
under surface of thorax testaceous ; abdomen slender, slightly
hairy towards the extremity. Fore wings hyaline, with
brown nervures, the costal nervure and the second median
nervure spotted with paler in the spaces between the trans-
verse nervures. Subcostal nervure slightly reddish. Towards
the extremity of the wing is a dark pterostigma, filling up
the narrow oblong space between two approximate nervures,
but not extending quite to the costa. Hind wings very long
and slender, white, with a yellowish central line, from which
numerous small blackish T-shaped markings diverge asym-
metrically on each side. ‘The extremity is broadly triflabel-
late, with a reddish-brown band extending over the first ex-
pansion, except at the base, and the basal portion of the second ;
the terminal part of the wing snow-white.
Hab. Maragoja Fundi, ‘Tara Desert, British Hast Africa ;
a pair collected by Mr. C. 8. Betton on March 3, 1897.
A very distinct species, somewhat allied to the West-
African A. imperatriz, Westwood.
The costal nervures vary in number from 33 to 37 before
the pterostigma on the fore wings, and are not uniform on
the two sides, showing that this is a variable character.
*13. Llalter tmperatriv.
Nemoptera imperatrix, Westwood, Trans. Ent. Soc. Lond. (3) v. p. 507
(1867); Thes. Ent. Oxon. p. 178, pl. xxxiil. fig. 8 (1874).
Hab. W. Africa.
14. Halter remipennis.
Nemoptera remipennis, Kolbe, SB. Ges. naturf. Fr. Berlin, 1900, p. 13.
Hab. Nyassa, Usambara.
15. [Halter toyonica.
Nemoptera togonica, Kolbe, SB. Ges. naturf. Fr, Berlin, 1900, p. 15.
Hab. Togo, Upper Guinea.
Neuropterous Family N emopteride. 461
Section 3.
Hind wings long, more or less expanded two or three times towards the
tips, and with two blackish bands on the expanded portion.
The British Museum possesses no species belonging to this
section, to which I refer H. graczl’s with a little doubt.
16. Halter gracilis.
Nemoptera gracilis, Hagen, Proc. Boston Soc. Nat. Hist. xxiii. p. 255
(1886).
Hab. Cape.
17. Halter barbara.
Nemoptera barbara, Klug, Abhandl. Akad. Berl. 1836, p. 94. n. 5;
Selys, Ann. Soc. Ent. Belg. x. p. 254, pl. ii. figs. 3, 4 (1887); Hagen,
Proc. Boston Soc. Nat. Hist. xxiii. p. 255. n. 6 (1886).
Nematoptera barbara, Westw. Proc. Zool. Soc. Lond. 1841, p. 11. n. 7;
Ann. & Mag. Nat. Hist. viii. p. 578. n. 7.
|| Panorpa halterata, Fabricius, Ent. Syst., Suppl. p. 208. n. 8 (1798).
Nemoptera algirica, Rambur, Ins. Névr. p. 3386 (1842); Luc. Expl.
Alg. iii. p. 139, pl. ili. fig. 3 (1849).
Hab. Algeria.
18. Halter Lederert.
Nemoptera Lederert, Selys, Ann. Soc, Ent. Belg. x. p. 254, pl. ii.
figs, 1, 2 (1887) ; Hagen, Proc. Boston Soc. Nat. Hist. xxiii, p. 254.
n. 5 (1886).
Hab. Asia Minor.
Section 4,
Hind wings less than twice as long as the fore wings, with two large
black dilatations and a short white terminal process.
19. Halter dilatata.
Nemoptera dilatata, Klug, Abhandl, Akad. Berl. 1836, p. 94. n. 6, fig. 1.
Nematoptera dilatata, Westw. Proc. Zool. Soc. Lond, 1841, p. 11. n. 5;
Ann. & Mag. Nat. Hist. viii. p. 378. n. 5 (1842),
Hab. S. Africa.
Section 5.
Hind wings less than twice as long as the fore wings, with a wide dark
dilatation towards the extremity, which is formed by a broad lobe,
rounded at the extremity, white at the base and dark beyond.
*20. Halter extensa.
Nemoptera extensa, Oliv. Ene. Méth, viii. p. 178, n. 4 (1811); Guér.
Ann. & Mag. N. Hist. Ser. 7. Vol. vie oe
462 Mr. W. F. Kirby on the
Icon. R. Anim., Ins. p. 384, pl. Ixi. fig. 1 (1829-1844) ; Klug, Abhandl.
Akad. Berl. 1836, p. 93. n. 4; Ramb. Ins, Névr. p. 336 (1842).
Nematoptera extensa, Westwood, Proc. Zool. Soc. Lond. 1841, p. At
n.4; Ann. & Mag. Nat. Hist. viii. p. 397. n. 4 (1842).
Hab. Baghdad.
Genus III. CHASMOPTERA, gen. nov.
Antenne very thick ; beak prominent; hind wings only one
and a half times as long as the fore wings ; fore wings hyaline,
except a slightly marked pterostigma; hind wings wholly
dark except at the extremity and on the median line in the
middle of the principal lobe, which expands very broadly on
each side, occupying the third quarter of the wing; at the
base it expands gradually, and at the extremity it is trun-
cated, the angles, however, being rounded off; from’ the
central line a number of bifid nervures diverge on each side.
This is followed by a second smaller lobe on each side, and a
broad terminal projection, white on the apical half.
*21. Chasmoptera Hutiit.
Nemoptera Huttii, Westwood, Proc. Ent. Soc. Lond. v. p. 27, pl. viii.
fig. 1 (1847) ; Hagen, Proc. Boston Soc. Nat. Hist. xxiii. p. 265.
n. 12 (1888).
Hab. Australia (Perth, Swan River).
Genus IV. SAVIGNIELLA, n. n.
\|Brachystoma, Rambur (nec Meigen), Ins. Névr. p. 337 (1842).
Mouth not produced into a beak ; hind wings Jong, linear,
scarcely enlarged towards the extremity.
22. Savigniella costata.
, Savigny, Descr. Egypte, Neur. pl. ii. fig. 14 (details).
|| Nemoptera halterata, Oliv. (nec Forsk.) Encycl. Méth. viii. p. 178
1811).
weaeene costata, Klug, Abhandl. Akad. Berl. 1836, p. 94. n. 7.
Nemoptera Olivieri, Rambur, Ins. Névr. p. 337 (1842),
Nematoptera Oliviert, Westwood, Proc. Zool. Soc. Lond. 1841, p. 13,
n. 14; Ann. & Mag. Nat. Hist. viii. p. 379. n. 14 (1842).
Hab. Alexandria.
Hagen considers Savigny’s fig. 13, usually referred to this
species, to represent Halter Ledereri, Selys.
Neuropterous Family Nemopteride. 463
Genus V. STENORRHACHUS.
Stenorrhachus, M‘Lachlan, Proc. Ent. Soc. Lond. 1886, p. viii.
|| Stenoteenia, M‘Lachlan (nec Gervais), Trans. Ent. Soc. Lond. 1885,
p. 376.
23. Stenorrhachus Walker?.
Stenotenia Walkeri, M‘Lachl. Trans. Ent. Soc. Lond. 1885, p. 377.
Hab. Chili.
The only described American representative of this family.
Genus VI. Croce.
Croce, M‘Lachlan, Trans. Ent. Soc. Lond. 1885, p. 378.
*24. Croce filipennis.
Nematoptera filipennis, Westwood, Proc, Zool. Soc. Lond. 1841, p. 13,
n. 19; Ann. & Mag. Nat. Hist. viii. p. 380. n. 19 (1842).
Nemoptera filipennis, Hagen, Proc. Boston Soc. Nat. Hist. xxiii. p. 267.
n, 13 (1888).
Hab. N. India.
The Natural History Museum possesses three specimens of
what may possibly be a second Indian species, the hind wings
being apparently much narrower than in the typical form ;
but this may perhaps be due merely to the pubescence being
more closely appressed to the shaft of the wing.
*25. Croce aristata.
Nemoptera aristata, Klug, Abhandl. Akad. Berl. 1856, p. 96, n. 13.
Nematoptera aristata, Westwood, Proc. Zool. Soc. Lond. 1841, p. 13.
n. 17; Ann. & Mag. Nat. Hist. viii. p. 380. n. 17 (1842),
Hab. Ambukohl (Klug); Thebes (B. M., presented by
A. Carter, Esq.).
26. Croce (?) alba.
Nemoptera alba, Oliv. Enc. Méth. viii. p. 179. n. 6 (1811); Klug,
Abhandl, Akad. Berl. 1836, p. 96. n. 12; Rambur, Ins. Névr. p. 336.
n. 6 (1842).
Nematoptera alba, Westw. Proc. Zool. Soc. Lond. 1841, p, 13, n. 18;
Ann. & Mag, Nat. Hist. vill. p. 380. n, 18,
Hab. Baghdad.
Found in abundance in houses in the evening towards the
end of May by Olivier, and described by him ; but no further
information about the species has been given by any author,
and it was apparently unknown in nature even to Rambur.
al*
464 On the Neuropterous Family Nemopteride.
27. Croce pusilla,
Nematoptera pusilla, Tascheuberg, Zeitschr. f. Naturw. lvi. p. 183
(1884). s
Nemoptera pusilla, Hagen, Proc. Boston Soc. Nat. Hist. xxiii. p. 268.
n. 14 (1888).
Hab. Socotra.
*28. Croce setacea.
Nemoptera setacea, Klug, Abhandl. Akad. Berl. 1836, p. 95. n. 10, fig. 3.
Nematoptera setacea, Westwood, Proc. Zool. Soc. Lond. 1841, p. 13.
n.15; Ann. & Mag. Nat. Hist. viii. p. 379. n. 18 (1842).
Hab. S. Africa.
29. Croce capillaris.
Nemoptera capillaris, Klug, Abhandl. Akad. Berl. 1836, p. 96, fig. 4.
Nematoptera capilaris, Westwood, Proc. Zool. Soc. Lond. 1841, p. 18.
n. 16; Ann, & Mag, Nat. Hist. viii. p. 379. n. 16 (1842).
Hab. Arabia Felix.
30. Croce ephemera.
Nemoptera (Croce) ephemera, Gerstaecker, Mitth. Ver. Vorpomm, xxy,
p. 152 (1893).
Hab. Mesopotamia.
31. Croce Baudii.
Nemoptera (Croce) Baudii, Gritfini, Boll. Mus. Torino, x. no. 214, p. 2
(1895).
Hab. Cyprus.
32. Croce Chobautt.
Croce Chobaut?, M‘Lachlan, Bull. Soc. Ent. France, 1898, p. 169.
Hab. Algeria.
38. Croce damare.
Croce damare, M‘Lachlan, Bull. Soc. Ent. France, 1898, p. 170.
JTab. Damaraland.
Mr. O. Thomas on Equus Penricei. 465
LX.— On Equus Penricei, a Representative of the Mountain
Zebra (Equus zebra, L.) discovered by Mr. W. Penrice in
Angola. By OLDFIELD THOMAS.
Durin@ his last expedition to Angola Mr. W. Penrice
discovered an animal which he rightly recognized as a repre-
sentative of the Mountain Zebra, hitherto only known * from
South Africa, and now, alas, nearly exterminated there.
Of this most interesting animal he brought home a flat
skin, which on examination proves to possess the deeper and
more essential characters of EH. zebra, such as the forward
slope of the median dorsal hairs, the presence of a “ eridiron-
pattern” on the rump, &c., but to differ from it so much in
other details that it clearly cannot be assigned to the typical
form of that species. The only question is as to whether it
should be considered a species or subspecies ; but since it is
now isolated geographically, and its general appearance is so
strikingly different from that of #. zebra, it would seem
necessary to call it a species until at least any evidence is
brought forward that intermediate specimens occur. ‘T’o no
other species than JZ. zebra is it at all allied, although the
equal striping of the body, the short close fur, and the buffy
tone of the light stripes give it a superficial resemblance to
£. Grevyt, which it may be said to rival in the extreme
handsomeness of its markings.
Equus Penricet, sp. n.
Spinal hair-slope, character of rump-stripes, gridiron-pattern
of loins, and complete striping of limbs as in &. zebra. Size
apparently rather larger than in that animal, but exact
measurements are not at present available. Fur short, close,
and glossy, the hairs of the back only aboutd millim. in length,
and therefore very different from the comparatively long and
shaggy hairs of #. zebra. Light ground-colour not white, as
in £. zebra, but buffy or creamy dun, as in L. Grevyi,
lightening, however, nearly to white on the belly. Dark
bands glossy brownish black. Striping throughout modified
from that of E. zebra by the broadening of the light and the
narrowing of the dark bands, the light bands being everywhere
(with the exception of those on the sides of the neck) as broad
as or broader than the dark ones, the latter being far the
* In Prof. Bocage’s “ List of the Mammals of Angola” (J. Sci. Lisb.
(2) v. p. 28, 1890) “ Equus zebra” is included, on the strength of observers
who had seen zebras in Southern Angola. Although Z. Burchelliis more
common there, Bocage’s name may in part refer to Penrice’s zebra.
466 Mr, O. Thomas on new Peruvian Species of
broader on all parts of the allied species. This difference is
most noticeable along the back, where, instead of dark stripes
about 14 inch wide alternating with white ones of half an
inch, both light and dark are equally about 1 inch wide.
Similarly on the face all the fine light stripes are broader
than the dark ones, which latter are, as usual, reddish instead
of black. On the ears again the white is increased at the
expense of the black. The transverse dorsal markings on
the whole are rather more uniform and less irregular than in
E. zebra, and they are throughout (except in two cases)
divided from each other by a well-defined narrow median
black line which runs from the withers right through on to
the tail. The gridiron-pattern of the loins is essentially
similar to that of HL. zebra, but owing to the alteration in the
proportions of the colours would rather be called black
markings on a buff ground instead of white ones on a black
ground. Upper light rump-stripe nearly twice the breadth of
that of H. zebra.
Length of the tanned skin, from tip of nose to base of tail,
8 feet 4 inches.
Hab. Mossamedes, Southern Angola. Type from Provi-
dencia, near the River Moninho, about 70 kilometres N.E. of
Mossamedes. Altitude 300 m.
Type. Male. B.M. no. 0. 9.12.1. Shot in May 1900
by Mr. G. W. Penrice:
Mr. Penrice informs me that the country where this zebra
is found consists of sand-flats dotted all over with stone kopjes
in a state of disintegration, the flats in many cases being
thickly strewn with boulders. He had heard of zebras being
occasionally found on the top of the Chella range, 2100 metres,
but was unable to say to which species they belonged.
He had first met the magnificent animal with which I have
associated his name on the Coroca River, 50 kilometres east
of Port Alexander, and at an elevation of only 180 m.,
and was also told that a zebra of this species had been shot
some two years ago within a few miles of the sea.
LXI.—New Peruvian Species of Conepatus, Phyllotis,
and Akodon. By OLDFIELD ‘THOMAS.
Conepatus arequipe, sp. n.
Closely allied to the Ecuadorean C. quitensis, with which
it agrees in the great breadth of the white stripes anteriorly.
Size rather less than in that animal. Fur of body long and
Conepatus, Phyllotis, and Akodon. 467
thick, as befits the high altitude ; that of the nape reversed
forwards from the withers to the crown. White stripes
ending on the loins, not continued on to the root of the tail,
as they are in C. quitensts and chinga. ‘Tail long, thick,
very bushy, its basal half black, its terminal half mixed
black and white, the hairs of the two colours about equally
numerous.
Skull broad and stout, shorter and broader than that of
C. quitensis, larger than that of C. rex and chinga. Upper
molar comparatively small.
Dimensions of the type (measured by collector in the
flesh) :—
Head and body 370 millim, ; tail 200; hind foot (s. u.) 70,
(c. u.) 80; ear 28.
Skull: basal length 70; greatest breadth 48; length of
nasals (in middle line) 13; interorbital breadth 25; greatest
diameter of m.* 9°5.
Another skull, that of an aged male, measures :—
Greatest length 87; basal length 73 ; zygomatic breadth
56; intertemporal breadth 20; mastoid breadth 47; palate
length 36:5.
Hab. Sumbay, Province of Arequipa, S. Peru. Alt. 4000 m.
Type. Male (young adult), B.M. no. 0.10.1. 2. Ori-
ginal number 1032. Collected 6th June, 1900, by Mr. Perry
O. Simons.
This skunk is an intermediate link between C. quitens?s and
C. rex. It differs from the former by its smaller size, by
the more definite reversal of the nape-hairs, and by the white
stripes ending on the loins and not passing backwards to the
root of the tail. From C. rex, on the other hand, it differs
by its rather larger size and by the white stripes being
throughout separated from each other by the median black
line.
Phyllotis sublimis, sp. n.
A vole-like species of medium size, with short tail and very
long fur. SOON),
Size medium; proportions rather Microtine, intermediate
between those of the medium-tailed Ph. boliviensis and the
remarkable short-tailed PA. Garlepp?. Fur long, extremely
soft and fine; hairs of back about 14 millim. in length.
General colour dull pale greyish fawn, greyer anteriorly,
more distinctly fawn on the sides and posteriorly on the
rump. Centre of back browner. Head like back, no darker
markings round eyes, Ears of medium length, not long as
468 Mr. O. Thomas on new Peruvian Species of
in the more typical Phyllotes, their anterior halves externally
brown, rather darker than the general head-colour; hairs
on their inner surface yellowish, an ill-defined patch of
yellowish fawn at their bases. Under surface white, not very
sharply defined, the bases of -the hairs slate. Upper surface
of hands and feet silvery white. Tail only about half the
length of the head and body, uniformly finely haired, not
pencilled ; a narrow line along the top ill-defined fawn, the
remainder white.
Skull more normal in general shape than the widely
expanded skull of Ph. boliviensis or the very peculiar one of
Ph. Garleppi. Nasals narrow, evenly tapering backwards,
considerably surpassing the premaxillary processes posteriorly.
Interorbital region flat, its edges square but not ridged,
Interparietal narrow, strap-like, its anterior edge concave
forwards. Anterior zygoma-root slanted backwards. Palatal
foramina long; choane narrow; bulle small, much smaller
than in Ph. boliviensis. Molars normal. Incisors narrow,
though not so much so as in Ph. Garleppi; pale yellow
above and below.
Dimensions of the type (measured in the flesh) :—
Head and body 105 millim.; tail 54; hind foot (s. u.) 21°5,
(c. u.) 23; ear 23.
Skull: greatest length 25:3; basilar length 20:2; greatest
breadth 14:5; nasals 10x3°6; interorbital breadth 3°7;
breadth of brain-case 12°5 ; palate length 12°1; diastema 6°9 ;
palatal foramina 6; length of upper molar series 4:7.
Hab. Rinconado Malo pass, above Caylloma, on the Sumbay
road, Peru. Altitude 5500 metres (nearly 18,000 feet).
Type. Female. B.M. no. 0. 10. 1. 60. Original number
1104. Collected 18th June, 1900, by Mr. P. O. Simons.
Nine specimens examined, all dug out of one burrow.
This most interesting little mouse lives at the highest alti-
tude from which mammalian life has been recorded in the
New World, and in the Old is only surpassed in this respect
by a few of the Himalayan species. Zoologically it is also
of interest, owing to its forming a link between the peculiar —
short-tailed Phyllotis Garleppi* and the other members of
the genus, so that it tends to resolve the doubt with which I
assigned that species to Phyllotis.
Akodon amenus, sp. n.
A medium-sized species with close fur and sharp-edged
supraorbital region.
* Aun, & Mag. Nat. Hist, (7) i. p. 279 (1898).
Conepatus, Phyllotis, and Akodon. 469
Probably most nearly allied to, and of about the same size
as, A. punctulatus, Thos.* Fur rather short, close and
almost crisp, the ordinary hairs about 8 millim. in length,
though these are mixed on the back with much longer fine
hairs 12-13 millim. long. General colour above dull yel-
lowish, rather, but not conspicuonsly, darker in the middle of
the back. Muzzle, cheeks, rings round eyes, and base of ears
clearer and deeper buffy yellow. Longer hairs of back brown,
tipped with white. Under surface dull whitish, fairly well
defined, the bases of the hairs plumbeous except on the chin,
where they are white to the roots. Ears rather short, well-
haired, of about the same colour as the head. Upper surface
of arms and legs, as far down as the metapodials, orange-
yellow; fingers and toes white ; fifth hind toe (without claw)
barely reaching to the base of the fourth. Tail about as long
as the body without the head, well-haired, its upper surface
dull orange, its lower white.
Skull very different from that of the more typical Akodons,
narrow, strongly bowed, its upper outline very convex and
its interorbital region sharply square-edged, though not
ridged. Nasals short and rather broad. Interparietal very
small. Palatal foramina long. Molars of normal Akodont
structure.
Dimensions of the type (measured in the flesh) :—
Head and body 100 millim.; tail 75; hind foot (s. u.) 19°5,
(e.7n-)21:5)3\ ear 16.
Skull (much damaged): extreme length 26°5; nasals
8°5x 3°73; interorbital breadth 4:9; breadth of brain-case
11:4; length of upper molar series 4:2.
Hab. Calalla, Rio Colea, near Sumbay, Peru. Altitude
3500 metres.
Type. Male. B.M. no. 0. 10. 1. 77. Original number
1109. Collected 19th June, 1900, by P. O. Simons.
This is an isolated species, apparently only allied to 4. pune-
tulatus, from which it is distinguishable by its very different
colour.
Besides these three species, Mr. Simons’s Caylloma and
Arequipa collection contains examples of the remarkable
Chinchillula sahame, of Akodon pulcherrimus, Phyllotis boli-
viensis, and Reithrodon pictus.
* Ann. & Mag. Nat. Hist. (6) xiv. p. 861 (1894),
470 Ona new Free-tailed Bat from Central America.
LXII.—A new Free-tailed Bat from Central America.
By Gerrit 8. MILxer, Jr.
A SPECIES of free-tailed bat represented by two skins from
Chiriqui, submitted to me for determination by Mr. Oldfield
Thomas, is strikingly distinct from any hitherto known. In
size and external appearance it suggests a small “ Nyctinomus”’
of the gracilis-europs group; but its skull and teeth show it
to be a Promops related, notwithstanding its small size, to
P. glaucinus.
Promops nanus, sp. n.
Type (in British Museum *),—Adult male (skin and skull),
no. 56 collection of H. J. Watson, taken at Bogava,
Chiriqui, Panama (alt. 250 m.), October 7, 1898. “ Caught
under the roof of a house.”
Character.—Essentially a miniature of Promops glaucinus ;
forearm about 39 millim. instead of 56 millim., greatest length
of skull 16°5 millim. instead of 22 millim.
Muzzle and lips—So far as can be determined from the
dried specimens, after relaxation of the parts, the lips are
slightly less wrinkled than in P. glauctnus, and the pointed
horny projections on the ridge above and between nostrils are
even smaller and less distinct.
EFars.—The ears are of the same form as in P. glaucinus ;
laid forward they extend about 2 millim. beyond tip of
muzzle; their inner borders arise together on forehead, about
5 millim. behind tip of muzzle. In outline the conch is
irregularly squarish, the anterior and superior borders about
equal in length, the posterior much shorter, the corners
bluntly rounded. Antitragus about two thirds as high as
wide, its anterior and posterior borders converging regularly
toward the bluntly rounded tip, the width of which is a little
less than half that of base. Keel well developed, its edge
very noticeably thickened. ‘Tragus minute, scarcely more
than 1 millim. in length, its highest point at extremity of
posterior margin.
Feet and membranes.—The feet and membranes present no
characters of importance. Wing attached at distal extremity
of tibie.
Fur and colour.—The fur is not peculiar either in quality or
distribution. On middle of back it is about 5°5 millim. in
length. General colour above dark bistre, very faintly washed
* [Now registered as no. 0, 7, 11. 99,—0, T.]
Some Results of Indian Dredging. 471
with ecru-drab. The drab is only visible in certain lights.
Underparts broccoli-brown, strongly washed with ecru-drab.
Throughout the body the hairs are whitish smoke-grey at
base.
Skull.—Notwithstanding its size, less than that of Nycti-
nomus europs or NV. brasiliensis, the skull of Promops nanus
closely resembles that of P. glaucinus. The brain-case is
relatively larger and less angular, and the basisphenoid pits
are actually larger. Bony palate entire anteriorly.
Teeth.—The teeth throughout are similar to those of Pro-
mops glaucinus, except that the upper incisors do not quite
come in contact with the canines and the posterior molar,
both above and below, is relatively larger and better deve-
loped. The third upper molar contains a small but distinct
metacone, a cusp that is lacking in the corresponding tooth of
Promops glaucinus. Similarly, the third lower molar is
provided with an entaconid of considerably greater relative
size than that of the corresponding tooth in the larger species.
Hypocone of first and second upper molars as in Promops
glaucinus.
Measurements. — Exxternal measurements of type: head
and body 41 mm.*; tail 84*; tail (free) 15; tibie 11; foot
78 (6°6); forearm 38; thumb 5°5; second digit 38; third
digit 75; fourth digit 58 ; fifth digit 37. The other specimen
(a female) is slightly larger; forearm 39-5.
Cranial measurements of type: greatest length 16-4 mm. ;
basal length 15; basilar length 138; median palatal length 6 ;
zygomatic breadth 9:8; least interorbital breadth 3:6; mas-
toid breadth 9; greatest breadth of brain-case above roots of
zygomata 8 ; lachrymal breadth 5; mandible 11:6; maxillary
tooth-row (exclusive of incisor) 6°4; mandibular tooth-row
(exclusive of incisors) 6°8.
LXIUI.—Natural History Notes from the Royal Indian Marine
Survey Ship ‘Investigator,’ Commander T. H. Heming,
R.N., commanding.—Series III., No. 4. Some Results of
the Dredging Season 1899-1900. By A. F. McArpte,
B.A., M.B., Capt. I.M.S., Surgeon-Naturalist to the
Survey.
Durine the season 1899-1900 the R.I.M.S. ‘ Investigator’
was for the greater part of the time without the services of a
Surgeon-Naturalist. At the beginning of April 1900 I joined
* Collector s measurement.
472 Capt. A. F. McArdle—Some Results
the ship at Colombo, and remained in her till the 4th May,
when she went into dock at Bombay. Altogether there were
twelve trawls made in over 100 fathoms, yielding a fair
number of Crustacea, Fishes, Mollusca, &c. Of these the
Fishes and Crustacea have been identified, while the Worms
and Mollusks remain still to be described by the specialists to
whose hands they have been entrusted.
Of the most interesting finds made during the year may be
mentioned :—
1. A very large specimen (an adult male), 12 inches long,
of Bathynomus giganteus, A. M.-Edw., dredged off the north-
west of Ceylon in 594 fathoms.
2. A specimen of that strange crab T’richopeltarium ovale,
from a depth of 445 fathoms.
3. Five specimens of the Atlantic fish Hoplostethus atlan-
ticum.
4, A trawl of twenty-eight specimens of Glyphocrangon
investigatoris.
5. ‘lwo fine specimens of Pennatula from 487 fathoms, in
the branches of each of which was found a small polycheete
worm of one and the same species.
6. Two specimens of Pleurotoma symbiotes, from 771 and
464 fathoms. As on the two previous occasions on which
this interesting mollusk was dredged, the same species of
Epizoanthus was found encrusted on the living shell.
7. A large collection (forty-five specimens) of Turbo indicus,
from 595 fathoms. The living shells were generally covered
on the spire by Scalpellum and sponges.
8. ‘I'wo fine specimens of a species of Conus were found at
the very unusual depth of 487 fathoms.
Most of the Crustacea obtained had been previously known,
but the following three species are new :—
BRACHYURA.
OXYRHYNCHA.
Fam. Maiide.
Subfam. Jwacurva.
CyrTomalA, Miers.
Cyrtomaia Goodridge, sp. n.
Description of an adult male.—Carapace bluntly triangular,
broader than long, vertically depressed anteriorly, broadly
rounded over the branchial regions, and with the gastric an |
of Indian Dredging. 473
cardiac regions greatly elevated. It is finely granular and
has numerous spines arranged as follows:—In the gastric
region one very long spine on either side, with a very short
one in the median line between them, a fourth gastric spine
intermediate in size being situate in the median line posterior
to the other three. In the cardiac region are two strong
spines on well-marked tubercles close to and on either side of
the median line. There is a well-marked hepatic spine. In
the branchial region there is a very long and strong spine
directed forwards, and laterally there are numerous small
spines, mostly arranged in two horizontal lines running
through the branchial and pterygostomial regions. There isa
prominent postocular, but no preocular nor supraocular spines.
A very small intestinal spine. The rostrum appears, when
looked at from in front or from above, to be trifid, owing to
the presence of a strong interantennulary spine, which is as
Jong as either of the proper rostral spines. The latter are
two in number, triangular, and run forward horizontally and
almost parallel to one another. They are bent downwards at
the tips. The abdomen consists of seven distinct segments ;
the first six carinate and spinous, the spine on the sixth
segment being the longest, the seventh segment granular and
rounded at its distal extremity. The sternum is spiny, both
anteriorly, where the spines are situated more or less trans-
versely, and laterally, where they are placed in two rows, one
at the bases of the chelipeds and ambulatory legs, and the
other row midway between them and the margin of the abdo-
men. ‘The eyes are short, robust, with a small spinule at
their distal extremities. The basal joint of the antenne has
two spinules on its outer border, one of which—that nearer
the base—points downward, the other pointing forward.
Arising between these two, but from the inferior surface of
the basal joint, there is a third spinule, pointing downwards.
The second and third joints are flattened from above down,
have each two smallspinules on their outer margins, and the
third joint reaches to the tip of the rostral spines. The
flagellum is long, slender, and setose. The third maxillipeds
have the ischium and merus strongly spinous, and the antero-
external angle of the latter is produced to a spiny tubercle.
The exognath is as long as the ischium and merus together,
and is somewhat narrowed anteriorly, The chelipeds are long
and slender, over two and a half times as long as the carapace
(77 : 28), the merus slightly shorter than the palm and fingers
combined (32:35), fingers regularly toothed and not fitting
closely. ‘Ihe first pair of ambulato1y legs are between twice
and one and a half times as long as the chelipeds [122: 77],
474 Capt. A. F. McArdle—Some Results
the second pair about two thirds as long as the first. The
first two pairs are strongly spinose, the spines being arranged
in longitudinal rows. ‘The last two pairs are shorter and
much more slender, and have no spines, with the exception of
one at the distal extremity of the merus. Dactyli nearly
straight and hirsute.
Colour in spirit dull ivory -white.
Adult ¢.
millim.
Length of carapace to base of rostrum..........+06- 28
Greatest breadth...) . sine emu rims alive mint 32
ene th Of POSEN etna elsiamtaqeieke Sree iets ontay eats stata 53
Jj) ABASETIC pile WRG ant ile le taeterdt Tid Seiden 13
. branchial spine |... irvestcash! «mateo ey haste 8
i chelipeds ..... PSA oO E ESO Ye oc. 14
A. first ambulatory legs ss s:< iss oe atric ii 122
A perfect adult male, dredged at Sta. 267 in about 500
fathoms, in lat. 7° 02! 30" N., long. 79° 36', off the west coast
of Ceylon.
Of the species heretofore described it is most like C. Smithit,
but differs in the shape and character of the rostrum, in
having no supraorbital spine, in possessing a large branchial
spine, and in the description of the basal joint of the antenne.
From C. Murrayi it can be easily distinguished by its
rostrum, by the absence of spines on its third and fourth
ambulatory legs, by having no preorbital spine, and by its
shorter and more robust eyes.
From C. Suhmi it differs in its shorter and more robust
eyes, in its general appearance, the regions of the carapace
being much better defined, in the general arrangement of the
spines, and by the rostrum.
A figure will be given in an early issue of the “ Illustrations
of the Zoology of the R.I.M.S. ‘ Investigator.’ ”
OXYSTOMA.
Fam. Dorippide.
Subfam. Dorrerivz.
Eruusa, Roux.
Ethusa hirsuta, sp. n.
Carapace subquadrilateral, flat, with its length equal to its
extreme breadth. It is expanded, but not greatly so, over
the branchial regions, and the lateral convergent borders are
of Indian Dredging. 475
almost straight. The whole of the carapace and the sternum
are covered with hairs, which are particularly long and strong
over the anterior and lateral borders. he cardiac region is
not depressed, and the branchio-cardiac lines, which are well
marked, do not meet in front. The front of the carapace is
four-toothed, the teeth equal in length, but the median being
more widely separated from one another than they are from
the lateral. The widest and deepest space is that between
the lateral teeth and the external orbital spine, which is long,
slender, acutely triangular, and extended forwards beyond the
tips of the rostral spines, and is directed slightly outwards.
The eyes are small, well pigmented, situated on moderately
sized, freely movable eyestalks, which extend well beyond
the angles of the orbital sinuses.
There is no distinct epistome. Antennules folded obliquely.
The bases of the antenne moderately developed. Fla-
gellum short. The outer maxillipeds have the ischium pro-
duced at its antero-internal angle; the merus articulates with
the next joint at its antero-external angle and has its antero-
internal angle rounded. ‘I'he exognath is long, slender, and
narrowed anteriorly.
The chelipeds are equal, more than three fourths the length
of the first pair of ambulatory legs. Palm and fingers taken
together are twice the length of the carpus. Fingers not as
long as the palm. The chelz fit well and there is no space
at the base.
The first pair of ambulatory legs are longer than the cheli-
peds (24: 193) and the second than the first, but the dispro-
portion is much less than in some allied forms. The last
two pair of legs are small, covered with hair, and have small
recurved dactyli. ‘The abdomen in the female is 7-jointed,
in the male 5-jointed, owing to the coalescence of the third,
fourth, and fifth segments.
Colour (in spirit) yellowish white.
ie
millim.
ere tihi Gl CHAE. i)dc ni eles a ee slchiolsles io s@aldea 15
WES LHCOUCATAMECE (os 2) tails 5 aidin.w ie area sre 18 tiga ed 15
iC enG he Gi CHOM BOOS O's «(5 's.6 aa 6 a vale cies a's ay eines Sua 194
- first, ambulatory legs: svi ck vee es 24°
+ second ambulatory legs ....cscesncvace’ 284
A female and a young male dredged at Stations 267 and
268 in between 500 and 600 fathoms, lat. 7° 02’ 30” N. and
7° 36’ 00” N., and long. 79° 36’ 00” E. and 78° 05' 00” KE
respectively.
476 Capt. A. F. McArdle—Some Results
The species is nearest HE. indica of Alcock and JZ. lata of
M. Rathbun (£. pubescens, Faxon). It differs from the former
in having a flatter carapace with less swollen branchial
regions, a cardiac region not depressed, in being covered with
hair, in having longer eyestalks, and in having the external
orbital spines directed forwards instead of obliquely outwards.
From £. lata by its general hairy covering, by the longer
eyestalks which extend beyond the angle of the orbit, the
external orbital spine projecting beyond the rostral spines,
and the chelipeds being much longer in proportion to the
ambulatory legs.
In the “Key to the Indian Species of Hthusa,” as pub-
lished by Major Alcock in the J. A. S. B. vol. Ixv. part ii.
no. 2 (1896), the species will be placed with H. andamanica in
part 2 of the first division, being divided from it by the dis-
tinction that the external orbital spines are long, acute, and
project beyond the level of the frontal spines, and the body
being hairy.
The species will be figured in the “ Illustrations to the
Zoology of the R.I.M.S. ‘ Investigator.’ ”
MACRURA.
Fam. Thalassinide.
CaALocaRris, Berl.
Calocaris Alcock, sp. n.
Description of an adult female.—Carapace laterally com-
pressed and smooth. Rostrum long, narrow, and curved
upwards, deeply grooved on its upper surface, the margins
of the groove being produced backwards as well-defined ridges
over the anterior part of the carapace, and giving rise on
either side tu a small spine directed upwards and forwards,
The rostrum itself has two small spinules on its left margin,
one on its right. All the regions of the carapace are smooth
and devoid of hair. A slight median carina runs the whole
length of the carapace. ‘I'he abdomen is longer than the
carapace, smooth, non-carinated.
Eyestalks short, eyes rudimentary; no traces of pigment
or cornea.
The second antenne have their first or basal joint unarmed,
the second joint has two fixed spines. A short internal spine
and an external, which is longer, but which does not reach
the next joint. From the second joint there also arises an
articulatory spine or scaphocerite, which projects but for a
of Indian Dredging. 477
very short distance over the third joint. The third joint is
slender and very long, longer than the other three segments
taken together; both it and the fourth joint, which is short,
are unarmed.
Flagellum long. Epistome well developed.
The chelipeds are long and slightly asymmetrical; coxa
unarmed; the ischium has one small spine above at its
proximal end, two small spines below at its distal. Merus
very long, flattened laterally, and curved on itself; at the upper
border of its distal end is a prominent spine. Carpus un-
armed. The palm has a large spine distally on its upper
border. The chele are narrow and long (but not so long as
the palm) ; on the right side the margins are sharp, finely
dentate, with curved crossed tips. On the left side the chele
are imperfect.
Second pair of legs slender, with small perfect chelee ;
remaining Jegs slender, with short curved dactyli.
With the exception of a few hairs on the last and penulti-
mate joints all the legs are naked.
The female genital openings are easily seen, and there is a
tubercle in the position of the male openings.
Eggs large, blue in life.
The outer plate of the large swimmeret has a diagonal
suture running obliquely from its outer to its posterior border.
There is a spinule at the outer end of this, but none along
the course of the suture. No spinules on the inner plate.
The telson is longer than broad, with a rounded end.
Colour in life reddish, in spirit yellowish white.
Adult 3.
millim.
Hueneth Of TOSEUME 0.5 Nese sn elea nts vce sas she Were s 5
-- carapace to base of rostrum.............. 163
* abdomen’ with telson 5. cn. owecesencees « 32
chelipeds :
ESC HATIMAN Petes cle share fees erate sect e's = soa 4
IUGHITE, saetteieets «ala rchivee sir nie eon nce wie. 6! ele 16
WORT ESS ares clei s ladle’ “Gawrles were eek hue rf
GEO latech aver wibetoteite, sastehene spermcainye tae aia se ll
GG CTS oot elation eres acini ale #5 oettte age vale 10
One adult female dredged off the north-east of Ceylon in
Sta. 266, in 542 fathoms, lat. 87,36 15" N., by long.
81° 20! 30" E. ali 4 i
I have placed this new species 1n the genus Calocaris of
Bell, in spite of its having a styloid scaphocerite on the
peduncle of its second antenne, a point which Faxon has
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 32
478 Mr. G. A. Boulenger on
thought of sufficient importance to found a new genus (Cata-
stacus) on. Moreover, Bell, in his description of the genus
Calocaris, says that the second antenne have “a large trian-
gular scale reaching to the end of the first joint,” and gives a
figure of it at the head of his description of the only species.
The length of this scaphocerite seems to be only a matter of
degree; it is very short in the species which has just been
described, it is longer in Calastacus investigatoris, And., and
Calastacus felix, and longest of all in Calastacus stilirostris,
Faxon.
C. Alcock? is easily distinguishable from C. Macandree by
the general naked appearance of the body and legs and by
the marked differences in the rostrum and chelipeds.
From Calastacus stilirostris by the short external spine
and scaphocerite on the second antenne, the rostrum being
grooved, the gastric area being smooth, and by differences in
the chelipeds and the telson.
From C. investiyatoris and C. felix by the carapace having
no hairs and not being granular, in having shorter spines on
the second antenne, no denticle at the end of the carina, and
the abdominal terga smooth, and by differences in the rostrum,
telson, and chelipeds.
The species will be illustrated in an early issue of the
* Tilustrations of the Zoology of the R.I.M.S. ‘ Investigator.’ ”
LXIV.—Diagnoses of new Fishes discovered by Mr, J. E. 8S.
Moore in Lake Tanganyika. By G. A. BouLENGER,
F.R.S.
I. Cyprinide, Silurida.
Capoeta tanganice.
D120). A, dilat tee Oeatell ina
14—15°
Depth of body 3? to 4 times in total length, length of head
5. Diameter of eye 34 in length of head, 2 in interorbital
width; a very small barbel. Third ray of dorsal very strong,
ossified. Caudal peduncle twice as long as deep. Olive
above, each scale darker at the base, silvery white beneath.
Total length 320 millim.
North end of Lake Tanganyika.
new Fishes from Lake Tanganyika, 479
Barbus platyrhinus.
D.11, A.8. L. lat. 40. L. tr. 35%,
Depth of body 3} times in total length, length of head 4.
Snout broad and rounded, twice as long as eye, which is 54
times in Jength of head, 24 in interorbital width ; barbels two
pairs, subequal, as long as eye. No bony dorsal ray. Ven-
tral below middle of dorsal. Caudal peduncle 12 as long as
deep. 384 scales between lateral line and root of ventral.
Olive-brown above, golden yellow beneath.
Total length 390 millim,
South of Usambura.
Barbus altianalis,
Dez. oA. oe doe lat.go.. Lh. tr. ze
Depth of body equal to or slightly greater than length of
head, 4 to 44 times in total length. Snout moderately broad
and rounded, 14 to 13 as long as eye, which is 5 to 54 times
in length of head, 2 in interorbital width ; barbels two pairs,
subequal, as long as ora little longer than eye. Third dorsal
ray very strong, bony, not serrated. Longest anal ray 3
length of head, nearly reaching caudal when folded. First
ventral ray corresponding to origin of dorsal. Caudal pe-
duncle twice as long as deep. 3 scales between lateral line and
root of ventral. Olive-brown, very dark above.
Total length 350 millim.
Lake Kivu and Rusisi River, N.E. of Tanganyika.
Barbus serrifer.
D; 10." AL 8. The lat. 28-30, E: tr. a,
Depth of body 8 to 3} times in total length, length of head
4 to 44. Snout rounded, as long as or a little longer than
eye, which is 4 to 43 in length of head and 14 to 14 in inter-
orbital width ; barbels two pairs, posterior longer, twice as
long as eye. ‘Third dorsal ray very strong, bony, strongly
serrated. Last ventral ray below first of dorsal. Caudal
peduncle 1} to 13 as long as deep. 3 scales between lateral
line and root of ventral. Olive-brown above, silvery white
below; a greyish stripe above lateral line; a small blackish
spot at base of caudal.
Total length 120 millim.
North end of Lake Tanganyika.
32*
480 On new Fishes from Lake Tanganyika.
Barilius Moorit.
D. 12. A, 16-17. 1, lat. 56-60. L. tr. “=.
Depth of body equal to length of head, 4 times in total
length. Mouth extending to below anterior third or centre of
eye. Anal originating below middle of dorsal. 3 scales
between lateral line and root of ventral. Silvery, brownish
on the back; about 10 dark vertical bars on the body ; dorsal
blackish at the end.
Total length 115 millim.
North end of Lake Tanganyika.
Barilius tanganice.
D138,.- +A, 20 2. lat82. eat: P
Depth of body equal to length of head, 44 times in total
length. Mouth extending to below posterior border of eye.
Anal originating below posterior third of dorsal. 4 scales
between lateral line and root of ventral. Silvery, olive on
the back ; 16 or 17 dark vertical bars on the body.
Total length 260 millim.
North end of Lake Tanganyika.
Chrysichthys brachynema.
D.16. A. 12-13.
Head little longer than broad, smooth above ; snout twice
as broad as long; diameter of eye 5 to 6 times in length of
head; maxillary barbel 4 or 3 length of head, outer man-
dibular 4 or $; nasal barbel not or but scarcely longer than
diameter of eye; teeth on the palate forming a broad crescentic
or horseshoe-shaped band on the vomer and _ pterygoids.
Dorsal spine 4 or ? length of head. Adipose fin measuring 4
or 4 its distance from the rayed dorsal. Pectoral spine very
strongly serrated. Caudal deeply notched, with obtusely
pointed lobes. Olive above, white beneath.
Total length 400 millim.
Several specimens from Kalambo and Usambura.
Synodontis granulosus.
LD Ho Dar jem ea: Bs
Head granular above; eyes supero-lateral; maxillary —
barbel simple, as long as or a little longer than the head;
On Diatoms from Chincha Guano. 481
mandibular barbels with short simple branches; anterior
mandibular teeth very short, 40 to 42. Adipose dorsal 4
times as long as deep, 24 to 34 times as long as its distance
from the rayed dorsal. Humeral process narrow, keeled,
sharply pointed. Body covered with granular papilla. Olive
above, yellowish beneath ; dorsal, anal, and paired fins black
in front, orange behind ; caudal black, edged with orange.
Total length 230 millim.
North end of Lake Tanganyika.
LXV.—Note on Diatoms from Chincha Guano.
By C. MERESCHKOWSKY.
[Plate XVI]
] HAVE, through the kindness of Mr. E. Thum, of Leipzig,
been provided with a very interesting slide containing a great
number of Diatoms from Chincha guano (in Peru). Having
carefully studied the various forms which it contains and
determined the species so far as possible with the aid of
the few books at my disposal, I give in the present note a
list of forty-one forms accompanied by a few remarks con-
cerning several of them and by the description of some new
species and varieties.
The majority of Diatoms of which the Chincha guano
material is composed belong to the group Anaraphidiez or
Cryptoraphidiee. Different kinds of Beddulphia, small species
of Coscinodiscus and Chetoceros, represented by a great
number of species, form the greatest bulk of this material.
There are only a few representatives of the group Raphidiez
and still fewer belonging to the group Pseudoraphidiew, or
Bacilloideze, as I propose to call this group *.
1. Diploneis vacillans, var. delicatula, Cl. Very rare.
2. Navicula Hennedyi, var. subrostrata, nov. var. (Pl. XVI.
fig. 14.) Very rare.
Size small, length 0:044 mm., breadth 0°025 mm.: valve
elliptic, with slightly rostrate apices ; lateral areas moderately
* In a note which will soon be published I have separated the Nitzschi-
oidez and Surirelloidez from the rest of the Pseudoraphidian Diatoms,
and given to the latter the name Bacilloidez, while the former have been
united in a new group called Carinate.
482 Mr. C. Mereschkowsky on
large, smooth, with almost parallel internal margins ; marginal
striae 11 in 0-‘01 mm., axial strie 13-14in 0-01 mm. Puncta
indistinct.
Among the numerous varieties of this species enumerated
by Cleve * there is not a single one which is described as
having valves with rostrate ends. I think therefore that the
above-described form belongs to a new variety. From its
small size it might be referred to the varietas minuta, Cl.
(size, according to Cleve, 0:027-0°05 mm.), or the varietas
tahitensis, Cl. (size 0°C4-0:045 mm.); but as Cleve does
not give in his ‘Synopsis of the Naviculoid Diatoms’ a suffi-
cient description of these two varieties and I do not possess
his work ‘On New and Rare Diatoms,’ where they are
figured, it is impossible for me to ascertain whether it is so
or not. It seems, however, not very probable that this
form belongs to the first variety on account of the striz, which
are here 11 in 0°01 mm., while in the varietas minuta they are
8-10 in 0°01-mm. ‘The same reason makes it even more
improbable that the above-described variety belongs to the
varietas tahitensis, which has 13-14 marginal strie. From
Tahiti I know another variety of Navicula Hennedy?, the
valves of which have also subrostrated apices; but this latter
(Pl. XVI. fig. 15) differs from the varietas subrostrata in the
lateral areas, which are of a somewhat different shape, their
interior margins being divergent; the ends of the valve are
also more or less elevated. I will give the description of the
variety from Tahiti in a future note on Polynesian Diatoms ;
I give here the figure of the latter in order te make the com-
parison of both forms easier.
3. Trachyneis aspera, Khr. Rather rare.
4, Pleurosigma nicobaricum, Grun. Rare.
Cleve, in his ‘Synopsis of the Naviculoid Diatoms’ f, attri-
butes to this species a length of 0°14 mm. I have seen a
specimen attaining 0°267 mm. in length, with a breadth of
0:036 mm., not differing in other respects from the description
which Cleve gives of this species.
5, Cocconeis costata, var. pacifica, Grun. Very rare.
6. Cocconeis scutellum, Khr. Mather rare.
* Cleve, ‘Synopsis of the Naviculoid Diatoms,’ part 11. p. 58,
t+ L.c. part i. p. 36,
Diatoms from Chincha Guano, 483°
7. Grammatophora angulosa, Ehr. Very rare.
There exist at least two different forms similar in their
general aspect, but differing in the strie, which are usually
confounded. In one of them the striw are distinctly seen
under a moderately high magnifying-power (12-14 strie in
0-01 mm.), while in the other they are invisible under such
conditions (about 20 strie in 0°01 mm.). The Chincha form
belongs to the latter, which I consider to be the genuine
G. angulosa, the other belonging to a quite different species.
8. Grammatophora maxima, Grun. Rare.
This species has been found hitherto, so far as I know,
only in a fossil state in Californian deposits (Redondo, Santa
Monica, San Pedro, Monterey, &c.). I have seen several
beautiful specimens in the Chincha guano, and it is also
sometimes to be found in a living state in California.
9. Sceptroneis caduceus, Ehr. Rare, and always in a broken
condition.
Attains a large size, over 0°1 mm.
10. Chetoceros bicoronata, sp.n. Very rare.
Of this species I have seen only a single endocyst, which
in its general outlines, as well as in the disposition of the
awns, resembles greatly the endocysts of C. sedracanthus,
Gran, recently described by H. Gran*. It differs essen-
tially from the latter in having a corona of spines not only on
one of the margins of the median part of the endocyst, but on
both ; the spines are also much stronger than represented in
the figures of Gran. Unfortunately I neglected to make a
drawing of this species at the time 1 found it, and afterwards
I did not meet with it again.
11, Chetoceros borealis, Bail. Rare.
12. Chetoceros chinche, sp. n. (Pl. XVI. figs. 3-7.) Not
very rare.
Of this species I know only the endocysts, which have a
very peculiar appearance. ‘heir form is elongated elliptical,
with rounded ends; the margins are provided with short,
more or less irregular spines, which become shorter towards
* H. Gran, in ‘ Norske Nordhays-Expedition,’ plate iii. figs. 40, 41.
484 — Mr. C. Mereschkowsky on
the ends of the valve; they do not form any regular row, as
in other species, but are disposed at rather variable intervals
one from another, and their apices are usually not very acute,
often bent in a hook, sometimes bifid (fig. 6). The surface
of the valve shows very peculiar, coarse, irregularly contorted
lines or markings, resembling somewhat Arabic inscriptions
or shorthand writing signs; these markings are sometimes
distributed all over the surface of the valve, and in this case
the diatom has a striking resemblance to Liradiscus ovalis,
Grev., the more so as the general outlines of both are nearly
the same *; but usually the lines or markings are not so nume-
rous, sometimes even more scantily represented than in the
figure 4, and sometimes there are only a few of them, as
shown in the figure 5. As will be seen from fig. 7, these
lines or markings are irregular membranaceous ribs elevated
over the surface of the valve.
Length: 0-024 0:0245 0:0265 0:027 0:031 0:042 0:042 0:049
Breadth: 0:0138 0-014 x 00146 x 0014 x O018
13. Chetoceros diadema (Ehr.), Gran. Common.
A very characteristic species, of which I have seen not
only the endocysts, but the frustules also. Filaments of this
species with endocysts enclosed in their cellules have been
recently observed and figured by H. Gran f.
14. Chetoceros didymus, Ehr. Very common.
15. Chetoceros incurvus, Bail. Rather rare; known only
by its endocysts. (PI. XVI. figs. 1, 2.)
This very peculiar species is easily recognized by the
disposition of the four awns with which the endocysts are
provided. The figures given by Brightwell are correct as to
their general outlines, but this author has failed to represent
the granulation of the surface of the valves; the granules
are small and irregularly scattered over the whole surface of
the body of the endocysts (figs. 1, 2). In the above-cited
work of H. Gran we find the description of a new species of
Chetoceros—the Cheetoceros cinctus, Gran, which has endo-
cysts very similar to those of C. incurvus, Bail.f, differing
* It would not be surprising if Zzradiscus should prove to be the endo-
cyst of some unknown species of Chetocer'os.
+ H. Gran, in ‘Norske Nordhavs-Expedition,’ plate i. fig. 18.
+ H. Gran, /. c. plate ii. figs. 23-27.
Diatoms from Chincha Guano. 485
only in the greater relative length of the awns and the size,
which seems to be larger ; the surface of the endocysts is also
granulated. It might represent only a variety of C. incurvus.
I give here some measurements of the latter :—
General length : 0:023 0:028 0:029 0:027 x
Length of the valve: 0°014 0:019 0:019 0:0195 0-021-
Breadth of the valve: x 0:010 0:010 0-009 0:0115
16. Chetoceros Lorenzianus, Grun. Very common.
17. Chetoceros peruvianus, Brightw. Rare.
There are a few more species of this genus, which, however,
I was unable to determine.
18. Rhizosolenia styliformis, Brightw. Very common.
19. Skeletonema costatum (Grev.), Cl. Very common.
20. Skeletonema costatum, var. spiralis, var.nov. (Pl. XVI.
fig. 8.) Not rare.
Differs from the type only in the oblique disposition of the
spines, which unite the frustules together ; the degree of in-
clination of the spines is variable, and there are always some
frustules in the same filament whose spines are straight, as in
the type species.
Although Skeletonema costatum is known to me from very
different localities all over the world, I have never before
seen such a spiral disposition of the spines; it seems therefore
advisable to regard the present form as a separate variety.
21. Eucampia zodiacus, Ehr. Rather rare.
22. Biddulphia (Triceratium) alternans, Bailey. Very
common,
23. Biddulphia (Triceratium) alternans, var. variabilis,
Brightw. Rare.
These two forms are generally regarded as belonging to
two distinct species—the second being characterized by the
irregular form of the valve, its concave margins and the acute
apices of the three ends; the first having a regular form,
straight margins, and broadly rounded apices. The exam-
ination of both of these species from Chincha guano, in which
486 Mr. C. Mereschkowsky on
the first (B. (7.) alternans) is very frequent, has shown me
without any doubt that they cannot be considered two
different species, being connected one with the other by all
possible intermediate forms. Usually the B. (7.) alternans
has regularly triangular valves, with the three apices of equal
length; but there occur forms in which, while one or two
apices are broadly rounded, as in the type species, the other
one (or sometimes two) is acute and has the peculiar irregular
appearance which characterizes the varietas variabilis.
In figs. 9,10, I have represented some of these interme-
diate forms. As to the structure of the valve, it is identical in
both, and their size is also the same,
B. (7.) alternans being the more common form, I regard
it as the type species and the other as its variety.
24, Cyclotella striata (Kiitz.), Grun. Rare.
Diameter from 0°0185 to 0-031 mm.
25. Auliscus sculptus, var. celata, Bail. are.
The only specimen observed agreed pretty well with the
figure of Bailey. ;
26. Actinoptychus undulatus, Ehr. Not very rare.
Often with a small process placed on the submarginal
median portion of each alternate compartment, sometimes of
all compartments.
27. Actinoptychus areolatus, Brightw. Rather rare.
Seems to be only a variety of the preceding species.
28. Actinoptychus splendens (Schadb.), Ralfs. Rare.
29. Asteromphalus flabellatus (Bréb.), Grev. Not very rare;
a widely distributed species.
30. Asteromphalus heptactis (Bréb.), Ralfs (Pritchard, Inf.
pl. viii. fig. 21; Spatangidium Ralfsianum, Norm. Micr.
Journ, vil. (1859), pl. vii. figs. 7,8; Asteromphalus Ralfs-
tanus, A. Schmidt, Atlas, pl. xxxviii. figs. 5-8) *.
Rather rare.
* The above-cited synonyms are given after Cleve, “ Report on the
Phyto-Plankton collected on the Expedition of H.M.S. ‘ Research,’ 1896,”
Fifteenth Annual Report of the Fishery Board for Scotland, part i.
p. 297.
Diatoms from Chincha Guano. 487
31, Asteromphalus malleus (or malleiformis), Wall., var. paci-
fica, var.nov. (Pl. XVI. figs. 11,12.) Not very rare.
Valve almost circular or slightly ovoid, with the upper
part (containing the median ray) a little narrowed. The
median ray is placed more or less asymmetrically, forming an
acute angle with the longitudinal axis; its central part is
conical ; 8 rows of puncta in 0°01 mm., the puncta bordering
the rays being a little larger than the others.
Length : 0:043 0:046 0:051 0:057
Breadth: 0-041 0:0455 0:050 0-057’
This diatom greatly resembles the type species from the
Indian Ocean which has been described by Wallich, differing
from it in the central part of the median ray. In the type
species this central part is quadrangular, with a deep con-
striction in the middle, dividing it into two parts of equal size
and shape; in the var. pacijica this part has never such an
appearance, being of a more or less conical form. In order
to show more clearly this difference I have reproduced in the
fig. 13 the type species as given by Wallich.
Although this difference is very constant, occurring in all
specimens which I have observed, and there is another differ-
ence consisting in the asymmetry of the median ray, still it
seems to me that these characteristics are too trifling to be
of any specific value. I think it therefore more reasonable
to consider the Pacific form simply a variety of the Indian
species.
32. Asteromphalus variabilis, Grev. Very rare.
Resembles exactly figure 7 of Greville, but has distinct
puncta, which are, I suppose, omitted in Greville’s figures.
33. Coscinodiscus radiatus, Ehr. Rare.
34, Coscinodiscus radiatus, var. asteromphala, Ehr. Very
rare.
The large central alveoles surrounding a small hyaline
circular spore are provided with elongated puncta along their
periphery.
35. Coscinodiscus excentricus, Khr. Very common.
36. Coscinodiscus gigas, Ehr. Rather common.
488 On Diatoms from Chincha Guano.
37. Coscinodiscus perforatus, var. cellulosa, A. S. Rare.
38. Coscinodiscus polyacanthus, Grun. Not rare.
Resembles C. sudtdl’s, but has numerous marginal spines.
39. Coscinodiscus subconcavus, Grun. Not very rare.
40, Coscinodiscus subtilis (Ehr.?), Grun. Rare.
Without marginal spines.
41. Spermatogonia antiqua, Leud. & Fortmorel. (Pl. XVI.
figs. 16-21.) Not rare.
This curious form, as to the nature of which there still exist
some doubts, was originally found in Java; but it seems to be
a widely distributed species. Besides the Chincha guano, I
have observed it in New Guinea (Tami Islands) and very
often among diatoms from the Adriatic Sea (Triest), and
Cleve has recently found it in the Plankton of the Atlantic
Ocean *,
The inflation at the lower end is not triangular, as described
by Leudiger and Fortmorel, but has exactly the form of the
head of a human spermatozoon. The frustule is usually not
straight, but more or less sigmoidally bent. The upper part
is enlarged and terminates in a point; one margin of the
enlarged portion is more straight, the other is always convex,
sometimes angular (fig. 17). ‘The striz are not capitate, as
described by the authors, this appearance being due to the
fact of their crossing the double outlines of the frustule. It
might be that the strie are superficial ribs, which give to the
margins a somewhat indented appearance of the same kind
as in Thalassiothriz, although in a much lesser degree. ‘The
number of striz is 8 in 0°01 mm.; they gradually disappear
near the lower end. Figure 21 represents a small and
straight individual with an obscure longitudinal line which
might possibly be a kind of keel.
Length: 0:096 0118 0:126 0:152 0161 x
Breadth :; 0-003 0:0024 0:0035 0:004 0:004 0:005°
The Chincha guano contains also a form known as Acti-
niscus sirius, Ehr., which, however, is not a diatom, but
* Cleve, ‘A Treatise on the Phytoplankton of the Atlantic and its
Tributaries’ (Upsala, 1897), p. 26.
On some new African Theraphosoid Spiders. 489
rather belongs to the so-called Silico-Flagellate. I have
found this same form in California and in the Mediterranean.
EXPLANATION OF PLATE XVI,
Figs. 1, 2. Chetoceros incurvus, Bail. Endocysts.
Figs. 3-7. Chetoceros chinche, Mer. Endocysts, (Fig. 6 in a somewhat
oblique position ; fig. 7 in profile.)
Fig. 8. Skeletonema costatum, var. spiralis, Mer.
Figs. 9, 10. Intermediate forms between Biddulphia (Triceratium) alter-
nans, Bail., and varietas variabilis, Brightw. The puncta are not
represented.
Figs. 11, 12. Asteromphalus malleus (malleiformis), var. pacifica, Mer.
900
es
Fig. 13. Asteromphalus malleus (malleiformis), Wallich, reproducing the
figure of Wallich. “
Fig. 14. Navicula Hennedyi, var. subrostrata, Mer.
Fig. 15. Another variety of N. Hennedy? from Tahiti.
Frgs. 16-21. Spermatogonia antigua, Leud. & Fortm.
LXVI.—Some new African Theraphosoid Spiders in the
British Museum. By R. I. Pocock.
Family Theraphoside.
Subfamily HvuzvorHorr.s, Poc.
Genus HYSTEROCRATES, Sim.
A few weeks ago the Trustees of the British Museum
acquired by purchase seven large Theraphosoid spiders which
were captured on the island of St. Thomas in the Gulf of
Guinea, and were offered to the British Museum under the
name Selenocosmia CGrreeffi of Karsch. Who may be respon-
sible for the identification I know not, but examination
showed the specimens to be referable to three perfectly distinct
species, neither of which is identical with the species described
by Karsch, if any reliance is to be placed on the description of
the latter.
The species may be described as follows :—
Hysterocrates didymus, sp. n.
? Phoneyusa Greeffi (Karsch), Simon, Hist. Nat. Araignées, i, p. 153
(1892).
¢.—Colour. Integument deep blackish brown, hairy
clothing a rich ruddy olive-brown, with paler tips to the
490 Mr. R. I. Pocock on some
segments of the legs; legs without distinct pale lines; cara-
pace, abdomen, and femora more sooty than the remaining
segments of the legs.
Carapace normal in form, shorter than patella and tibia of
fourth or of first leg, and than protarsus and tarsus of fourth,
and than patella, tibia, and tarsus of palp; its width sub-
equal to or a little less than distance between ocular tubercle
and posterior emargination, equal to or slightly exceeding
femur of fourth leg, less than femur and patella of third leg ;
no perceptible transverse groove in front of fovea.
Mandibles tubercular in tront.
Legs 4,1, 2, 3 in length, fourth exceeding the first by the
length of its tarsus; fourth leg not in any sense thickened,
the height of the femur not quite a third of its length
(7: 23°5); patella and tibia narrower than femur, patella a
little thicker and higher than tibia; tibia cylindrical, about
three times as long as high or wide (18: 5°5); length of
upperside of patella more than twice its height or width.
Palp with tarsus tumid at base, being proximally higher
and wider than tibia (woodcut, fig. b, p. 493).
Measurements in millimetres.— Total length 58; carapace
28, its width 25; length of first leg 73, of second 62, of
third 60, of fourth 84; patella and tibia of first 29, of fourth
30; protarsus and tarsus of fourth 31, of first 22.
Loc. Island of St. Thomas in the Gulf of Guinea (Moc-
querys coll.).
This species is probably the same as that determined by
Simon as Phoneyusa Greeff, Karsch, which, according to
Simon, has the tarsus of the palp tumid above at the base.
Karsch’s description contains nothing to justify this statement,
no special modification of this segment being mentioned,
Since, moreover, the length of the legs in P. Greeff is,
according to the description, different from that of the legs
in the form here described, I am compelled to regard the two
as specifically distinct.
HHysterocrates scepticus, sp. n.
9 .—In appearance and structure closely resembling the
preceding, but with the tarsus of the palp normal and the
legs shorter as compared with the carapace, the latter being
longer than the patella and tibia of first or fourth legs.
Most nearly allied to H. hercules *, Poc., from Jebba, up
* In the description of this species the carapace stands as 34 millim.
long. Including the posterior epimeral piece (which, consistently, should
be included), the length is 35°5,
new African Theraphosoid Spiders. 491
the Niger (P. Z. S. 1899, p. 844), and like that species in
presenting a shallow transverse groove in front of the fovea,
but certainly differing in the much greater inequality in
length between the first and fourth legs. In H. hercules the
first leg falls short of the fourth by nearly half the length of
the tarsus of the fourth, whereas in H. scepticus it falls short
by more than the length of the tarsus; the fourth leg in
H. hercules is exceptionally short, its patella and tibia being
less than those of the first.
Measurements in miliimetres.—Total length 63; length of
carapace 32, its width 27 ; first leg 74°5, second 63, third 60,
fourth 86; patella and tibia of first leg 30, of fourth 31;
tarsus and protarsus of fourth 31; femur of fourth 24°5, its
height 7; tibia 17°5, its width 5.
Loc. Island of St. Thomas in the Gulf of Guinea (Moc-
querys coll.).
Hysterocrates apostolicus, sp. n.
?.—Resembling H. scepticus in having the tarsus of the
palp normal, but distinguishable by the much greater length
and strength of the fourth leg.
Fourth leg strong, rather thicker than the first, the femur
three times as long as high (22: 7), the tibia a little less than
three times as long as wide (16:5°5); height of tibia equal
to that of patella; the patella and tibia or protarsus and
tarsus are very noticeably longer than the carapace, the whole
limb surpassing the first leg in length by its tarsus and one
third of its protarsus.
Carapace about equal to patella and tibia of first leg and
to patella, tibia, and tarsus of palp; no transverse groove in
front of the fovea.
Measurements in millimetres. —Total length 51; carapace
25:5, its width 22; length of first leg 62°5, of second 53, of
third 52, of fourth 79; patella and tibia of first 25, of fourth
28; protarsus and tarsus of fourth 28-5.
Loc. Island of St. Thomas in the Gulf of Guinea (Moc-
querys coll.),
Of the species that occur on the mainland, H. apostolicus
approaches H. gigas, Poc. (P. Z. 8. 1897, p. 762, and 1899,
p- 845), from the Cameroons, and H. laticeps, Poc. (op. cit.),
trom Old Calabar. From the formerit may be recognized by
its broader carapace and by having the carapace shorter
instead of longer than the patella and tibia or protarsus and
tarsus of the fourth leg; from the latter by having the patella
and tibia of the fourth greater than of the first leg instead of
492 Mr. R. I. Pocock on some
equal, and by the fourth leg exceeding the first by more than
its tarsus in length instead of by less than its tarsus.
The only species of this group previously recorded from
St. Thomas is that described by Karsch as Selenocosmia
Greeffi, the generic position of which is still a little doubtful.
In his original description (SB. Ges. Naturwiss. Marburg,
1884, p. 60) Karsch writes, when describing the legs, ‘ Die
Metatarsen der beiden Hinterpaare an der Spitze unterhalb
mit... Scopula’”’; whereas subsequently (Berl. ent. Zeit.
1886, p. 83) he says “ Scopula an Metatarsus iv. fast hart bis
zur Basis des Gliedes reichend.’”’ ‘These statements clearly
involve either an error of identification, the author con-
founding two distinct genera as one species, or of observation.
Simon has assumed the latter to be the case, and, following
him, I have on the strength of the assumption referred
S. Greeffi to the genus Hysterocrates. If, however, as is
possible, the specimens that Karsch first described as
S. Greeffi have the scopula on the fourth protarsus confined
to the apex of the segment the species will fall no doubt into
Phoneyusa.
Assuming that it falls into Hysterocrates, the known species
from the island of St. Thomas may be tabulated as follows :—
a. Legs short; first leg just exceeding twice the length of
the carapace (62:30); fourth leg only twice and one
sixth the length of the carapace (according to Karsch). Greeff.
b. Legs longer; first legy* not less than twice and one
third the length of the carapace; fourth leg not less
than twice and two thirds the length of the carapace.
a, Tarsus of palp swollen at base, wider and higher
HAMID oe i orien Peleiehemiele cle tet: wheter eles didymus.
b'. Tarsus of palp normally cylindrical, not wider than
tibia.
a’, Fourth leg not thickened ; carapace a little longer
than its patella and tibia or protarsus and tarsus.. scepticus.
b?, Fourth leg a little thickened; carapace con-
siderably shorter than its patella and tibia and
protarsug aud tarsus Cyan, ss «acts slot omens wee» apostolicus.
CITHARISCHIUS, gen. nov.
Resembling Phoneyusa, Karsch (with which, pending a
re-examination of the types, I provisionally include Harpaxo-
theria, Sim., and Pelinobius, Karsch), in the restriction of the
* The legs are here measured from the base of the femur. Karsch’s
measurements may contain a greater number of segments, but cannot
contain a smaller. I have assumed that they contain the same. If they
contain a greater number of segments, the shortness of the legs in Greeff
is even greater than here represented.
new African Theraphosoid Spiders. 493
scopula of the fourth protarsus to the apical third of the
segment, but distinguishable from all the known genera of
the subfamily by the size and peculiar shape of the labium,
which is as long as half the length of the sternum, has the
form of a truncated cone, being so much narrowed anteriorly
that its apical width is only half the basal width, the apex
itself being rounded and not transversely truncated (wood-
cut, fig. a). Moreover the third and fourth legs are much
stouter than the first and second.
Type Citharischius Crawshay?, sp. n.
a.—Citharischius Crawshay?, gen. et sp. n. Labium and sternum.
b.—Hysterocrates didymus, sp.n. Extremity of palpus, 2.
Citharischius Crawshay?, sp. n.
2 .—Colour. Integument reddish brown, hairy clothing
rusty red, shining with silky lustre, and brighter in tint on
the distal segments of the legs than on the femora, carapace,
and abdomen.
Carapace much longer than wide, the width less than the
distance between the ocular tubercle and the posterior median
emargination ; length equal to that of patella, tibia, and half
the protarsus of first leg, exceeding patella, tibia, and tarsus
of palp, just exceeding patella and tibia of fourth leg, but
barely equal to protarsus and tarsus of fourth.
Legs 4,1, 2, 3 in length: third and fourth much more
robust than first and second, first a little more than twice as
long as the carapace ; second and third subequal, less than
Ann, & Mag. N. Hist. Ser. 7. Vol. vi. 33
494. On some new African Theraphosoid Spiders.
twice as long as the carapace, fourth more than twice and
a half as long; first leg falling short of fourth by the
tarsus and one fourth of the protarsus of the latter ; no spines
on legs except at apices of protarsi beneath ; femur of fourth
considerably less than three times as long as high (23:9);
tibia about as wide as high, less than three times as long as
high (163: 6); patella rather more than twice as long as
high (15: 63) ; the inner side of the patella, tibia, protarsus,
and tarsus pseudoscopulate—that is to say, the hairs are
short, erect, close-set, and equal in length like those of a
mole’s skin; on the protarsus this peculiar modification of
the hairy clothing extends well on to the upper and under-
side of the segment.
Measurements in millimetres.—Total length 52; length of
carapace 33, its width 25°5; length of palpus 47, of first
leg 69, of second leg 64, of third leg 63, of fourth leg 85 ;
patella and tibia of first 27, of fourth 31:5.
Loc. Kinani, British East Africa.
A single female specimen, collected and presented, together
with a male of Phoneyusa Bettont, Pocock, by Mr. Richard
Crawshay.
The distinctive features of the undoubted genera of Humeno-
phorine, a subfamily of Theraphoside characterized by the
presence of aremarkable stridulating-organ between the basal
segments of the palpus and first leg (an organ described in
P. Z. 8. 1897, p. 772, and figured in the ‘ Zoologist,’ 1898
P ’ 8 gist, ’
pp. 14-21), may be tabulated as below :—
a. Scopula on fourth protarsus extending almost to
base of segment.
a’, Fovea of carapace large and wide ............ Loxomphalia.
bl, Fovea of carapace small, crescentic .......... Hysterocrates.
b. Scopula on fourth protarsus confined to distal third
of segment.
a. Tarsa] scopula of fourth leg divided by a broad
band oF sete i). .':1. 2 {23 . Satnel. wth. Senor ates hale Lumenophorus.
b?, Tarsal scopula of fourth leg undivided.
a®, Labium very lovg, conical, half the length of
the sternum ; fourth leg much thicker than
GIS Spee ces esctay spicier cic alleae eyanacerate i oles tas Rech Citharischius.
b*. Labium less than half the length of sternum ;
fourth leg not thicker than first.
a‘, Anterior tibia of male with spine-tipped
SPUN aid wit ee tenth eee eRe wee rahcy Naa Monocentropus.
4, Anterior tibia of male unarmed.
a’. Fovea lerge,'straight (. \o.t. sss pele nse Anoploscelus.
6°, Fovea small, crescentically procurved .. Phoneyusa.
On new Genera and Species of Hymenoptera. 495
LXVII.—Deseriptions of new Genera and Species of
Hymenoptera. By P. CAMERON.
[Continued from p. 419. ]
The Males of Icaria.
Bingham, in his ‘ Hymenoptera of India,’ does not describe
the male sex of any species of Jcarta. Saussure, in his
‘ Htude sur la Fam. d. Vespides,’ only describes the male of
one species, from which it would appear that the males must
be rarer than the females or workers.
The male of J. ferruginea is of the same size as the female,
and agrees with it in general coloration. ‘The underside of
the scape, the eye-incision, the face, clypeus, and mandibles
are pale yellow ; the clypeus is strongly punctured, except at
the apex ; the clypeusis broader than long, is slightly convex,
and is roundly and broadly incised in the middle, the sides
being oblique; the fifth and following joints of the antenna
are roundly incised, the incision becoming deeper towards the
apex ; the apical joint is deeply incised, almost hook-shaped,
somewhat as in Humenes.
The male of variegata has the middle of the front, the eye-
incision, the clypeus, and mandibles lemon-yellow; the clypeus
is smooth, the incision on its top is somewhat horseshoe-
shaped, deep and black ; the lateral keel does not reach to it;
the flagellum becomes gradually thicker to the penultimate
joint, the joints are not incised, the last is about three times
Jonger than the preceding and is broadly curved on the lower
side; on its inner side beneath it is broadly hollowed, the
hollow bordered by a keel, which is obliquely curved. The
pro- and mesopleure are for the greater part yellow, streaked
with rufous; the mesosternum 1s also yellow; the mesonotum
blackish ; the scutellum, postscutellum, and metanotum are
also yellow ; the yellow band on the apex is one fourth of the
length of the segment and has a black band at its base; the
other segments are for the greater part yellowish; the petiole
is shorter than the second segment by about one fourth, and
it is distinctly longer than it is in the %. In length it is
12 millim.
Tam not certain, but this may be the male of artifex, with
which, in some points, it agrees better than it does with the
female of vartegata. ‘Lhe petiole appears to be intermediate
between the two, being larger than in variegata, but not so
long compared with the second segment as in artifex. The last
joint of the antenne is longer compared with the preceding
than it is in ferruginea and is differently formed; the joints
preceding it, too, are simple, not incised, as in Sorrugines.
33
496 Mr. P. Cameron on new
Tcaria quadrimaculata, sp. n.
Brunnea, flavo-maculata; petiolosegmentoque 2° flavo-bimaculatis ;
alis hyalinis, nervis stigmateque nigris. o.
Long. 10 mm.
Hab. Bengal, probably Barrackpore (Rothney) .
Antenne blackish, the scape yellow below; the flagellum
below and at the apex above brownish; the last joint is
shortly but distinctly longer than the preceding. Head
brownish ; the eye-incision, the face, clypeus, and outer orbits
narrowly yellow ; the vertex and front are thickly covered
with silvery pubescence ; the frontal keel is stout and extends
to the top of the clypeus, which is roundly incised in the
middle; the sides are oblique. Mandibles yellow, like the
clypeus, with the teeth black. Thorax brownish; a narrow
line on the pronotum, the base of the propleure narrowly, a
conical mark below the tegule in front, a smaller more oval
one below it behind the furrow, a large irregular mark on the
mesopleure on the lower side behind, a mark below the hind
wings, the base of the scutellum (more broadly laterally), the
postscutellum, and two marks on the metanotum, lemon-
yellow. Legs fuscous, the anterior lighter in tint; the tarsi
black ; the apex of the hinder tibize and of the hinder femora
yellowish. Wings hyaline; the radial cellule infuscated.
Abdomen brownish black: two spots on the node of the
petiole near the middle, two in the centre of the second
segment, and lines on the sides of the others, two large
marks on the second segment beneath, broad lines, narrowed
in the middle, on the base of the third to sixth, and the last
broadly, lemon-yellow. ‘The petiole is long and slender and
not nodose at the apex, it becoming only gradually and slightly
wider towards the apex from the middle.
This species is not, I am sure, the unknown male of
I. fuscipennis ; it appears to be too small; the markings
on the thorax and abdomen are smaller and of a distinct
Jemon-yellow colour, not pallid; the head and abdomen are
darker, more distinctly black in colour,
Icaria ornaticeps, sp. W.
Nigra, late flavo-maculata, fronte mesonotoque flavo-lineatis ; pedibus
posticis nigris, coxis apiceque femorum flavis ; alis hyalinis, apice
fumatis, nervis stigmateque nigris. %.
Long. 8 mm.
Hab, Khasia Hills. Coll, Rothney.
Genera and Species of Hymenoptera. 497
Antennz black; the scape yellow; the flagellum rufous
beneath, densely covered with pale pubescence. Head black,
the outer orbits entirely below, more narrowly above, the
inner orbits from shortly above the top of the eye-incision
(broadly below the incision, more narrowly above), a large
irregular mark on the middle of the front extending trom the
ocelli to the base of the antenne, the clypeus (except round
the edges), and an irregular line in the centre above, a curved
line on the sides behind the ocelli, and the mandibles, lemon-
ellow. The mandibular teeth are black at the apex,
Thorax black ; the prothorax at the base, on the sides broadly
(especially on the upper half, where it extends to the middle
of the pleure), two lines on the centre of the mesonotum at
the base, the sides of the scutellum widely, the base of the
postscutellum widely, the median segment (except in the
middle), a large irregular mark on the base of the mesopleurze
under the wings, an irregular mark on the lower side at the
apex, its base prolonged backwards below, and an elongated
mark above the hinder coxe, lemon-yellow. The thorax is
thickly covered with a pale pile and is impunctate ; the short
furrow on the metapleure black. Legs black; the fore
coxe, the four posterior (except broadly at the base below), a
line on the fore femora in the middle in front, the apex
broadly below, the apical half of the middle, the apex of
the posterior, with a streak in the centre, the fore tibice
behind, and a mark on the middle pair, lemon-yellow. Wings
hyaline, the costa, stigma, and nervures black; the radial
cellule with a broad black cloud in front. Abdomen black ;
a pear-shaped mark on the side of the petiole, two large
irregular marks on the base of the second segment, its base
below broadly, its apex all round narrowly, the sides of the
third broadly, lemon-yellow. The petiole is about two thirds
the length of the second segment and becomes gradually
wider towards the middle; the apical half is of uniform
width ; the second segment cup-shaped, widely rounded at
the base; the second cubital cellule is much narrowed above ;
both the recurrent nervures are received behind the middle of
the cellule.
Easily known by the yellow marks on the head and meso-
notum,
Icaria rufocollaris, sp. n.
Nigra ; capite, prothorace, scutello, dimidio apicali petioli apiceque
segmenti secundi rufis; apice clypei, basi postscutelli medioque
metanoti flavis ; alis fuscis, nervis nigris, stigmate testaceo, 9.
Long. 13 mm,
Hab, Khasia Hills. Coll. Rothney.
498 Mr. P. Cameron on new
Antenne black, the scape and the greater part of the
second and third joints rufous; the flagellum of a duller
rufous colour beneath. Head rufous; the vertex, occiput,
the edges of the clypeus, and a large mark in its centre
black ; the apex of the clypeus pallid yellow. Mandibles
dull rufous, their base pallid yellow, the teeth black. Front
and vertex distinctly and closely punctured; the clypeus with
some scattered punctures, its apical tooth sharp, distinct ;
below each antenna is a short, deep, oblique furrow. Thorax
black ; the pronotum, a large oblique mark on the meso-
pleuree, and the greater part of the scutellum rufous ; the base
of the postscutellum and the middle of the median segment
pale lemon-yellow. Pro- and mesonotum with the scutellum
closely punctured and thickly covered with pale pubescence ;
the postscutellum punctured at the base only; the median
segment finely transversely striated. Pro- and mesopleure
closely punctured, the punctures almost forming reticulations ;
the lower part of the metapleure obscurely punctured. Legs
black, the four anterior coxe yellow below; the femora and
tibiee rufous, black above; the tarsi black. Wings fuscous,
the front and apex smoky; the nervures and costa black ;
the stigma testaceous; the second cubital cellule is much
narrowed at the top; the first recurrent nervure is received
near the base of the apical third of the cellule. Abdomen
black ; the apical dilated half of the petiole, the sides of the
second segment narrowly at the junction between the dorsal
and ventral parts, and a narrow line before the apex, rufous ;
the apex itself is yellow.
The male has the face and clypens pallid yellow, this being
also the case with the lower part of the eye-incision; the
front and vertex are entirely black; there are two large
yellow maiks on the sternum between the four front legs;
the neck of the second segment and a streak along the sides
at the base are yellow; otherwise as in &.
Comes near J. speciosa and ungulata; easily known from
them by the rufous prothorax. Instead of there being a
detached spot on the base of the second segment there is a
short yellow line laterally at the extreme base.
Icaria nigroplagiata, sp. n.
Ferruginea ; metathorace nigro, medio flavo ; abdominis segmento 2°
basi biflavo-maculato, apice flavo; alis fere hyalinis, apice fu-
mato, stigmate flavo, nervis nigris. - 2.
Long. 11 mm,
Hab. Khasia Hills. Coll, Rothney.
Genera and Species of Hymenoptera. 499
Antenne dark rufous; the flagellum blackish above.
Head thickly covered with short pale pubescence; the front
and vertex closely covered with round, not very deep punc-
tures ; the ocellar region black. Clypeus yellow, its edges
and a broad band down the middle black, the black edged
with rufous, sparsely punctured and thickly covered with
silvery pubescence, its top broadly roundly incised in the
middle; the lower half of the outer orbits is black. Thorax
rufous ; the postscutellum and a large mark in the middle of
the metanotum (incised at the top and bottom) lemon-yellow.
Pro- and mesothorax with the scutellum closely and distinctly
punctured ; postscutellum smooth. Mediansegment obscurely
striated; the yellow mark on it does not extend far below the
middle. The punctuation on the metapleure is obscure.
Legs dark rufous, the coxe, trochanters, femora, and hinder
tibie darker coloured; the fore coxe largely, the middle
slightly, marked with pale yellow; they are thickly covered
with white hair. Wings hyaline; the radial cellule and the
upper part of the apical cubital cellules smoky ; the top of
the second cubital cellule is as long as the space bounded by
the second recurrent and the second transverse cubital ner-
vures. ‘The narrowed basal half of the petiole is black ; the
node is longish ; the petiole distinctly longer than the second
segment; the second and following three segments are closely
punctured and thickly covered with pale hair; the two yellow
maiks on the base of the second are irregular in shape; the
apical band is narrow.
This species is closely related to ZL. artifex; it may be
known from it by the metathorax being entirely black, except
for the yellow mark on the middle of the metanotum, by the
scutellum not being furrowed down the middle, by the yellow
mark on the metanotum being smaller, it not extending much
beyond the middle, the pleure entirely without yellow, and
the head is more broadly developed behind the eyes, especially
above.
Icaria carinata, sp. n.
Ferruginea; abdomine nigro, flavo-maculato, basi ferruginea ; alis
fusco-hyalinis, stigmate nervisque nigris. 6.
Long. 15 mm,
Hab. Khasia Hills. Coll. Rothney.
Comes into Bingham’s Section Aa (/. ¢. p. 286), in which
it will form a new division, distinguished by having the second
and following segments of the abdomen black. Antenne
ferru ginous, the flagellum darker above. Head ferruginous
500 Mr. P. Cameron on new
darker above ; the face thickly covered with silvery pubes-
cence; the antennal tubercles longish, extending equally
below and above the antenna, and sharply keeled down the
middle ; the front and vertex closely punctured, closely trans-
versely above the antenne ; the sides narrowly and the apex
of the clypeus broadly pale yellow; the apex of the clypeus
broadly rounded, not projecting in the middle. Mandibles
pale yellow, thickly covered with silvery pubescence; the
teeth black ; the palpi pale yellow. ‘Thorax dark ferruginous ;
a narrow line on the base of the postscutellam and two large
marks (longer than broad and oblique at the base and apex)
on the apex of the median segment lemon-yellow. The
scutellum is darker coloured than the mesonotum and is
widely furrowed down the centre on the apical half; the post-
scutellum is finely longitudinally striated in the centre. The
median segment is narrowly furrowed at the base, the rest
with the furrow much wider; the segment is thickly covered
with short fuscous hair and is obscurely transversely striated ;
the furrow is blackish on the apical two thirds. The pro-
pleuiz below, the mesopleure above, and the base and apex
of the metapleure more or less blackish. Legs dark rufous ;
the four hinder femora infuscated above; the tibie black,
except in front, the tarsi for the greater part black. Wings
hyaline, with a fuscous tinge ; the radial cellule and the apex
ot the costal cellule smoky. Petiole dark rufous, blackish at
the base and apex ; the rest black; there is a broad yellow
band on either side of the second segment at the base; two
smaller ones on the base of the third; a mark on the sides of
the fifth near the middle; the last segment is broadly obscure
yellow on the base. The apex of the petiole is yellowish
and sharply bordered laterally ; the basal three fourths of
the second and third segments, the basal half of the fourth,
the base of the fifth narrowly, and a longer semicircular mark
on the base of the sixth, yellow.
A very distinct species, intermediate between Bingham’s
sections “A. Reddish or ferruginous brown, with yellow
markings,” and “ B. Black, with red or yellow markings.”
Icaria Wroughtoni, sp. n.
Long. 15 mm.
Hab. Poona (Wroughton).
Comes into Bingham’s section “ A. Reddish or ferruginous
brown, with yellow markings,” and “a. Second abdominal
segment with no transverse yellow band on its apical margin,”
which now stands :—
Genera and Species of Hymenoptera. 501
Abdomen with no yellow markings; the scutellum
MUO Retin Livd nse aiols sy do e'e xin ued acoihe guttatipennis.
Abdomen with large yellow markings; the scutellum
deeply and widely furrowed behind.......... Wroughtont.
Antenne rufous, darker towards the apex. Head rufous;
the antennal tubercles and the apex and sides of the clypeus
broadly lemon-yellow; the antennal tubercle punctured,
narrowly keeled in the middle; the clypeus with scattered
shallow punctures, its sides oblique, ending in the middle ina
black blunt tooth; the apex of the clypeus with scattered
punctures; the face bears a white pile ; the clypeus is sparsely
covered with long fuscous hair, black along the apex. Man-
dibles yellow ; the teeth black. Thorax rufous; the pleure
more or less and the sides of the median segment yellow.
Scutellum broadly furrowed down the middle. Median seg-
ment broadly furrowed down the middle, the top, sides, and
furrow blackish, the centre yellowish, the furrow finely
transversely striated. Propleurze broadly lemon-yellow round
the edges, except behind; the lower part sharply keeled at
the base, the lower part behind the coxe raised, hollowed
above the raised part; there is a bifurcated furrow on the
mesopleurz, from the centre of which a furrow runs obliquely
to the fore coxe. On the metapleura below the wings is a
curved furrow ; behind this is an obliquely striated space ;
the base of the pleurez more coarsely striated, the striated
part becoming wider on the lower part. Metapleure finely
obliquely striated. Legs rufo-testaceous, the coxe yellowish.
Wings hyaline; the radial cellule and the upper part of the
costal smoky; the costa and stigma rufo-testaceous; the
nervures slightly paler, especially towards the apex. Petiole
rufous above, the sides yellowish ; the other segments ferra-
ginous, the second to the fifth with two large transverse
yellow marks; the second ventral segment is for the greater
part yellow ; the others broadly yellow at the base.
Icaria fuscipennis, sp. n.
Long. 13-14 mm. 9°.
Hab. Khasia Hills. Coll. Rothney.
Comes into Bingham’s section A and is closely related to
I. carinuta, with which it agrees in having the abdomen
black and the wings fuscous-hyaline, but may be known by
the shorter petiole, by the pleura and sternum being marked
with yellow, by the mesonotum having two yellow lines, by
the four hinder legs being largely marked with black and
502 Mr. P. Cameron on new
yellow, and by the second abdominal segment being marked
with yellow at the base. In the presence of the two yellow
marks on the base of the second abdominal segment it agrees
with artifex and variegata, from which it is otherwise very
distinct.
Antenne dark rufous, darker above; the scape yellow
below. Head dark rufous; the clypeus, the inner orbits to
the top of the eye-incision, the upper half of the antennal
tubercle, the outer orbits (narrow above, broader below) from
near the top to the base of the mandibles, the palpi, the man-
dibles (except at the apex), a transverse line on the pronotum
above, one on its sides, the postscutellum, two large marks on
the metanotum, an oblique squarish mark under the tegule,
a larger oblique one (narrowed at the base) on the lower side
on the apex of the mesopleure, a mark on the top of the
metapleure, triangularly dilated below, on the sides of the
mesosternum is a large mark turned outwardly at the base
and largely dilated at the apex, on the apex is a larger
oblique mark, at the tubercles a smaller one, and on the
lower half of the metapleure is a large mark, incised in the
middle above, and the base of the propleure, all clear yellow.
Scutellum narrowly furrowed in the middle from the base to
the apex; the median segment widely but not deeply furrowed
down the middle. Wings fuscous-hyaline; the apex of the
costal cellule and the radial cellule smoky, especially above.
The four front legs yellowish, the trochanters, the femora in
front (except at the apex), the basal half of the tibie, and
the basal joint of the tarsi fuscous; the middle coxe broadly
on the inner side below, the basal half of the femora, the
basal two thirds of the tibiz and the tarsi black ; the hinder
legs similarly coloured, but with the black more extended.
Petiole dark rufous; the sides (except at the apex) and a
mark on the sides of the dilated part, two large marks on
the sides of the second segment at the base, two large marks
on the base of the third, two smaller ones on the base of the
third and two large marks on the last segment, the sides of
the petiole beneath, a large mark (narrowed at the base) on
the base of the second, a large transverse line (dilated at the
sides) on the base of the third, and a narrowerone on the base of
the fourth segment, lemon-yellow. Thesides of the clypeusare
oblique, its middle ending in a short blunt tooth ; the antennal
tubercle is large ; the edge of the pronotum is sharply keeled
above; the thorax above thickly covered with short white
pubescence.
Genera and Species of Hymenoptera. 503
Icarta tinctipennis, sp. n.
Ferruginea ; abdomine, coxis femoribusque posticis nigris; alis
fusco-hyalinis, nervis stigmateque nigris. 9.
Long. 21 mm.
Hab. Khasia Hills. Coll. Rothney.
Antenne rufous; the basal joints of the flagellum darker
above. Front and vertex shagreened, thickly covered with
fulvous pubescence ; the part between the antenn projecting,
keeled in the middle. Clypeus shining, sparsely punctured,
thickly covered with fuscous hair; the apex in the middle
forming a distinct tooth. Mandibles pilose, sparsely and
distinctly punctured; the teeth deep black. Thorax rufous ;
the edges of the mesonotum, a line down the centre, the apex
of the mesopleure, the part along the sutures, the breast, the
base of the metapleure, the central furrow of the metanotum
and its sides black. The entire thorax is thickly covered
with short pubescence, which has a fulvous tint on the meso-
notum ; the pleuree are shagreened ; the metanotum obscurely
transversely striated. The nervures are paler, more testa-
ceous in tint than the costa and stigma; the second cubital
cellule above is distinctly wider than the space bounded by
the two recurrent nervures ; the space bounded by the second
and by the third transverse cubital nervures is slightly but
distinctly wider than the space enclosed by them. Legs
coloured like the thorax ;- the coxz, the hinder trochanters,
and the hinder femora blackish ; the four hinder tarsi are
infuscated. Petiole longer than the second segment, the node
becoming slightly and gradually narrowed towards the apex ;
the third segment is one third longer than the fourth, which
is equal in length to the fifth.
The differences between this species and I. fulvinerva may
be best shown in synoptical form :—
Petiole rufous; wings yellowish; the fourth abdo-
minal segment distinctly longer than the fifth .. fulvinerva.
Petiole black ; wings not yellowish ; the fourth abdo-
minal segment equal in length to the fifth...... tinetipennis,
The petiole is stouter and more distinctly dilated towards
the apex, the scutellum is not furrowed down the middle, and
the propleure not striated as in fulvipennis. It is one of the
largest of the Indian species.
504 Mr. P. Cameron on new
Icaria fulvinerva, sp. n.
Ferruginea ; abdomine nigro, petiolo ferrugineo ; tarsis posteriori-
bus nigris; alis flavo-hyalinis, nervis stigmateque rufis. 2.
Long. 17-18 mm.
Hab. Khasia Hills. Coll. Rothney.
Antenne ferruginous ; darker above, the scape brownish.
Front and vertex alutaceous; the incision and the inner orbits
black; the front with a narrow furrow in the middle; the
part between the antennz projecting, its sides oblique, the
centre keeled. Clypeus shining, of a brighter colour than
the front, sparsely punctured, its apex thickly fringed with
fulvous hair. Mandibles dark ferruginous, sparsely punc-
tured, the teeth black. Thorax thickly covered with a silky
pile. Mesonotum and scutellum shagreened, the latter with
a shallow furrow down the middle. ‘The striz on the meta-
notum not prominent and absent from the furrow, which is
shallow and black on the apical half. The lower half of the pro-
pleure striated; the furrow on the mesopleure black, deep, its
basal fork not continued to the base and not much longer than
the apical. Legs coloured like the body; the four hinder
tarsi black above and for the greater part below; the calcaria
pale. Wings hyaline, with a distinct yellowish tint, espe-
cially in front; the second cubital cellule is fully more than
half the length of the third above and slightly less than the
space bounded by the first transverse cubital and the first
recurrent nervures ; the latter is received shortly in front of
the middle. Petiole rufous, longish, its node not much
dilated ; it is nearly as long as the second and third segments
united; all the segment very smooth and covered with a
silky pile.
A distinct species, easily known by the red head, thorax,
and petiole, black abdomen, and fulvous wings.
Odynerus hindostanus, sp. n.
Long. 9mm. Q.
Hab, Barrackpore (Lothney).
Agrees closely with QO. sibilans; differs from it in the
yellow mark on the front being separated from the yellow
lateral marks, in the clypeus being broader compared to its
length, it being broader than long, its sides are not so straight
and its apex in the middle is more distinctly incised, and the
second ventral segment bears two large yellow marks.
Antenne black, the scape yellow, the flagellum rufous
beneath. Head black; the clypeus, a somewhat crown-
Genera and Species of Hymenoptera. 505
shaped mark on the front, the eye-incision, and the posterior
orbits yellow. Vertex closely, the front somewhat more
widely punctured. Clypeus wider than long, its sides
roundly curved; the apex with a shallow incision. Mun-
dibles yellow, the apex broadly rufous. Thorax black; a
broad mark on the sides of the pronotum (the two almost
touching), the tegule, a large mark un ler the tegule, the
scutellums, and the sides of the median segment broadly,
yellow. Pro- and mesothorax closely and strongly punctured,
the upper basal slope of the pronotum, the base and apex of
the mesopleure, the base of the metapleurz, and the middle
of the metanotum smooth; the punctuation on the median
segment is slight, its centre is broadly and slightly hollowed
and is keeled down the middle. Legs yellow; the anterior
coxe entirely, the four posterior behind, the anterior femora
at the base behind and the posterior entirely (except at the
apex), black; the four posterior tibiee are marked with black
behind. Wings fuscous-hyaline, the apex violaceous; the
second cubital cellule much narrowed above, scarcely one
fifth the length of the third. Abdomen black; the apical
half of the petiole broadly at the nodes and the apex entirely,
a large irregular mark (wider than long) on the sides of the
second segment, its apex broadly, widest in the middle, the
apex of the third to fifth, and the sixth broadly in the middle,
yellow. On the second segment beneath are two yellow
marks ; its apex and that of the third broadly yellow.
The male is similarly coloured; there is less black on the
legs ; the yellow mark on the front is prolonged between the
antenne as a short streak; the flagellum is black, except
at the apex; the yellow on the scutellum is narrowed and
incised in the middle.
Odynerus Antoni, sp. n.
Niger, dense albo-pilosus ; basi clypei, linea pronoti lineaque abdo-
minis segmentis 1° et 2° rufis; alis fusco-violaceis. ¢.
Long. 7 mm.
Hab. Barrackpore (Rothney).
Antenne entirely black, except the apical claw, which is
brownish ; it is thick and reaches to the base of the eleventh
joint. On the head, the base of the clypeus broadly, a mark
between the antenne, and a small mark behind the eyes are
brick-red ; the front and vertex are closely and_ strongly
punctured and covered with long white hair ; the keel between
the antennz is stout. Clypeus roundly convex, sparsely
punctured, thickly covered with white, somewhat depressed
pubescence ; its apex depressed and slightly roundly incised
506 Bibliographical Notices.
in the middle. Pro- and mesothorax strongly rugosely punc-
tured, covered thickly with white hair; the pronotum is
transverse at the base, its sides triangularly projecting ; on
the posterior half of the mesonotum are two shallow furrows.
Apex of postscutellum rounded. The apex of the median
segment has an oblique slope and is transversely aciculated ;
it is bounded above and at the sides with a stout keel, which
is bent down towards the centre above; from this a narrow
keel runs down the middle. Pro- and mesopleura strongly
punctured; the punctures large and distinct ; there is a some-
what triangular red mark at the tubercles. Metapleure
finely and closely longitudinally aciculate ; the upper half
irregularly longitudinally striated. Legs black ; the anterior
tibie entirely in front and the middle pair at the base testa-
ceous. Wings fuscous-violaceous, the posterior pair paler;
the second cubital cellule above not quite one half the length
of the third. Abdomen black ; the first and second segments
lined with black on the apex; the petiole has an oblique
slope on the base and is sharply keeled above; it is some-
what strongly but not closely punctured ; the other segments
are obsoletely punctured ; the base of the second is crenulated.
Allied to UO. stkhimensis.
Note—At p. 415 antea, for Polistes khasianus read
P, lepcha.
[To be continued. }
BIBLIOGRAPHICAL NOTICKS.
Catalogue of Eastern and Australian Lepidoptera Heterocera in the
Collection of the Oxford University Museum.—Part IL. Noctuina,
Geometrina, and Pyralidina. By Colonel C. Swryuox, M.A.,
F.LS., F.Z.8., FES. Pterophoride and Tineina by the Right
Hon. Lord Watstnenam, M.A., LL.D., F.R.S., &c., High Steward
of the University of Cambridge, and Joan Harriey Durrant,
F.E.S., Memb. Soc. Ent. France. With eight Plates. (Oxford,
Clarendon Press, 1900.) Pp. vii, 630.
Tur importance of public collections of insects depends partly on
the number of species accurately named, and partly on the possession
of types, by which the descriptions of the original describers of a
species can be checked or verified. The original specimens on
which species are founded should always find their way eventually
into public collections ; for private collections are not always easy
of access, and their contents are almost sure, sooner or later, to be
lost or dispersed. Great Britain is peculiarly rich in public col-
lections, and may boast of a considerable number of the types of
Linné, preserved by the Linnean Society ; though Queen Ulrica’s
collection, from which Linné likewise described many species,
Bibliographical Notices. 507
remains at Stockholm. Fabricius, the pupil of Linné, and his
worthy successor in entomology, was much in England, and
described the insects in the collections of Sir Joseph Banks,
Dr. Hunter, and others; and though some of these types are
missing, a large proportion are still preserved in the British Museum
(Natural History) and also in Glasgow.
Oxford University Museum contains no types of the last century ;
but the nucleus of its entomological collection consists of the
united collections of Hope and Westwood, to which large additions
have been made from other sources, especially from the collection
of Wilson Saunders, which was specially rich in the types of
moths described by Francis Walker from the collections formed
by Dr. A. R. Wallace in the Malay Archipelago. As Walker’s
descriptions are frequently short and unsatisfactory, it was a matter
of considerable scientific importance to verify them as far as possible
from the original types; and hence the present work was under-
taken by Col. Swinhoe, and the first volume, containing Sphinges
and Bombyces, and illustrated by eight plates, was published in
1892. The second volume, just issued, and twice the thickness of
the first, completes the subject. It is a full synonymie catalogue
of the Eastern and Australian moths in the Oxford Museum, and
special attention has been paid to the elucidation of Walker’s types,
a considerable number of which are figured, as well as many new
species which are now described and figured by Col. Swinhoe and
his coadjutors for the first time.
Books like the present are of great use to all entomologists
who are working at exotic moths, and we cordially recommend
Col. Swinhoe’s work to their special attention.
Sexual Dimorphism in the Animal Kingdom. By J.T, Cunnrnewam,
M.A. London: Adam and Charles Black, 1900.
Colour in Nature. By Marton J. Newstery. London: John
Murray, 1898.
Iy the production of the first-mentioned book, the nucleus of which
appeared in the pages of the now unhappily extinct ‘ Natural
Science,’ Mr. Cunningham has, without doubt, spared neither time
nor pains. As a result, he has brought together a considerable
number of facts of real and lasting value. Whether, however, his
interpretation of these facts will find favour with students of this
subject is another matter: we shall be surprised if he succeeds in
making a single convert.
Mr. Cunningham is Lamarckian in principles. His object has
been, he tells us, not to ‘* attempt to prove that acquired characters
are inherited,” but “‘ merely to point out how remarkably the multi-
tudinous facts all agree with the hypothesis that secondary sexual
characters are due to the inheritance of acquired characters.” This
very cautious statement of his case looks somewhat as though
Mr. Cunningham were a little afraid of the ghost which he has
conjured up.
The beard of man, and especially of the Caucasian races, it is
suggested, owes its conspicuous development to the stimulation of
508 Bibliographical Notices.
the growth of the hair by tugging of the teeth or hands in the
combats of mature males. But, as Mr. Cunningham remarks, “ It
is unlikely that men should fight in the way suggested after they
had reached even the stage of evolution known to us in the lowest
existing savages, for they seem always to fight with weapons.”
Thus, from the dawn of man’s evolution the stimulation which
Mr. Cunningham invokes has ceased to act, at least with any degree
of periodicity ; therefore by the tenets of his faith these secondary
sexual characters should have disappeared, from disuse. After all,
as Mr. Cunningham remarks, ‘“ what is wanted is evidence con-
cerning the influence of mechanical irritation of the hair-follicles in
the growth of the hair.”
The common fowl, we are reminded, in fighting erects the long
neck-hackles, “ which accounts for their elongation.” The blows of
the beak upon the head received during these contests had similarly
been made to account for the comb and wattle. The crest of the
peacock has been produced, we are solemnly informed, like the
comb of the common cock, by strokes of the beak, which, being less
violent, have plucked at the feathers instead of injuring the skin,
and so causing an outgrowth from it.
Are we to account in the same way for the two remarkable
feathers which depend, one on either side of the head, in the King
of Saxony’s Bird of Paradise? or the wonderful balls of down which
invest the legs of the humming-birds of the genus Spathura?
The brilliantly coloured bare skin of the cassowary is the result
of the irritation, we are asked to believe, ‘“‘ produced by the blows of
the beaks of the birds when fighting ... . the habit of fighting with
the beak also explains the presence of the bony crest on the skull.”
As a matter of fact the cassowary fights not with the beak, but
with the legs, which by powerful down-strokes can inflict serious
injuries. No one who knows anything of the structure of the
helmet of the cassowary would suggest that its existence was due to
blows of the beak.
We venture to think that the above instances are indicative that
Mr. Cunningham has failed to seriously injure the case for sexual
selection, which, as yet, it must be admitted, holds the field, even
though it may fail to satisfy us as an explanation of all the problems
which it is called upon to solve.
As a whole we may say of this book that it contains many new
things and many good things; but the new things are not good and
the good things are not new. Nevertheless it will doubtless be
widely read and will find a place on the bookshelves even of those
who differ from Mr. Cunningham on this very important subject.
It is well printed, well illustrated, and nicely got up.
In the little work on ‘Colour in Nature’ we have an undoubtedly
important contribution which will enable us to test the value of
much that has hitherto been of a purely speculative nature. The
book has probably by this time become tolerably well known. To
those who have not yet made its acquaintance we would recommend
it as an extremely interesting and helpful work.
THE ANNALS
MAGAZINE OF NATURAL HISTORY,
[SEVENTH SERIES.]
No. 86. DECEMBER, 1900.
LXVIII.—A suggested Origin of the Segmented Worms, and
the Problem of Metamerism. By H. M. Brrnarp, M.A.
Cantab.
Durine the last five years in which I have been working at
the stony corals, the Coelenterata, as an animal group, have
been more or less continuously in the foreground of my
imagination. I do not pretend to have seriously worked at
any other than the Madreporaria; yet the fact just stated
will, I trust, be accepted as some excuse for the following
article. In it I propose to show how the Ccelenterates
may have given rise to the segmented Worms. The idea
forced itself upon me by its simplicity, and, although not
altogether new, it has never, so far as I am aware, been
stated as here proposed. I may add that I am still further
impelled to trespass beyond the region of my own serious
studies by perceiving that, if there is any truth in the sugges.
tion, it offers a possible solution to that vexed problem “ What
was the origin of the mesoderm ?”’
In the geological record metamerism first appears in the
Trilobites, which are constructed of a number of segments
jointed together. With the exception of a head-piece and, in
most cases, a tail-piece, which differ from one another and
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. a4
510 Mr. H. M. Bernard on the
from the other joints, all the segments in the typical Trilo-
bite are alike in form; they diminish in size, however, pro-
gressively from front to back. Further, with regard to the
head-piece, it is not difficult to see that this has been built up
of a certain number of fused and thus modified segments.
The same is true of the tail-piece, as we gather not only from
its markings but also from the fact that some of the very
oldest Trilobites are found without any such specialized fusion
of segments. Characteristic of the Trilobite segment is the
pair of immense pleural folds, which can only be regarded as
secondary specializations. We conclude, then, that the
Trilobites must have been descended from some form made
up of simpler segments, 7. e. of segments without pleure, and
that these segments, with the exception of the first, containing
the mouth, and the last, containing the anus, were in all
respects similar in form, but diminished progressively from
the oral to the anal end.
We look elsewhere, then, for more primitive segmentation
than that which we find in these ancient fossils. We have
not far to look, for there exists a group in which it attains its
greatest development, the primitive segments being apparently
but little altered by secondary specializations. I refer to the
segmented Worms. In typical forms we have here a long
string of segments, the first and the last alone being peculiar ;
all the rest, no matter how many, are structurally exact
repetitions of one another. Not only are the appendages and
markings of the segments faithfully repeated from segment to
segment, but if cut open the internal machinery of each
segment, excluding the reproductive, is found simply to
repeat the internal machinery of that which precedes it;
muscles, nerves, blood-vessels, excretory organs, are all more
or less exactly repeated.
‘That metamerism reaches its highest development in the
Worms there can then be no doubt; hence it is to animals
below the Worms in structural complexity that we must look
for its origin. Where segmentation or vestiges of it are
found in animals above the Worms—for instance, we have
traces of it in ourselves—we are justified, in the absence of
clear evidence to the contrary, in assuming that it has been
derived from Annelidan ancestors. It is to my mind almost
outside the bounds of probability that such a remarkable ©
repetition of parts could have been developed twice on quite
independent lines.
Now the probfem is, How did such an animal come into
being? Several interesting suggestions have been made,
Origin of the Segmented Worms. 511
but zoologists have not been able to agree as to the process
which best satisfies the conditions. But, on the other hand,
zoologists are all, I think, agreed that the form which comes
with the greatest probability next below the typical segmented
Worm is the Cceelenterate ; for as yet no satisfactory position
has been found for the other groups of vermiform animals,
and though we usually place them near one another, we travel
outwards from the typical Annelid towards these groups with
uncertain steps. We only reach solid ground again when we
come to the Ceelenterates. Morphologists have, indeed, long
recognized that the Ccelenterate body marks a well-defined
stage in the evolution of the Metazoa. The mechanism is
simple, yet very perfect, for the two primary functions of
animal life, the vegetative and the reproductive. Its success
may be gathered from the extraordinary wealth of the Ccelen-
terates still peopling the waters of our planet both in numbers
and in forms. But we are justified in assuming that this
success has attended them from their first appearance in early
geological periods, bearing in mind that, as we go back to
simpler conditions of environment—simpler in the fact that
many animal forms preying and to be preyed upon were not
then evolved,—the variations in form would be less and less
numerous, until we reach a time when the simplest of all
conceivable types of Ccelenterate was the only representative
of the race.
It is to the Celenterates therefore that we naturally look
first for the ancestors of the segmented Worms, and more
than one attempt has been made to sketch the form-changes
which would be necessary. So far as I remember, the earliest
and, to my mind, the best suggestion refers the segmented
Worms to the serial budding which is known to occur among
the Coelenterates, e. g. in the “ strobila.” This is, however,
so far only a suggestion; I know of no attempt to work it
out in detail; one apparently insuperable difficulty blocks the
way, viz. the presence, in the Annelids, of the ccelomic
cavities, which are unknown in the Ceelenterates. Prof,
Sedgwick * proposes to deduce the ccelomic cavities of the
Annelids from the mesenterial chambers (‘‘ gastric pouches ”’)
of a Ccelenterate-like ancestor. This would not explain the
addition of new segments, progressively diminishing in size, at
the posterior end. ‘The method of the increase in the number
of the mesenteries which takes place in the higher Ccelen-
terates is quite different. Itseems to me that the reference of
* Quart. Journ. Micr. Sci. xxiv. (1884),
34*
512 Mr. H. M. Bernard on the
segmentation to serial budding is singularly apt. No group
of animals before or since, excluding perhaps the Protozoa,
have shown such marvellous developments of this method of
reproduction. The number of successful colony-formations
among the Coelenterates, both stationary and free-swimming,
is perfectly bewildering. We have, then, in this power of
increase by budding a wealth of possibilities upon which we
can safely draw without putting any strain whatever upon
our sense of what is probable. I go even further, and think
it possible to show that free-swimming strings of buds must
at one time have existed as a necessary interaction between
an organism like the Ccelenterate, capable of budding any-
where, and its environment.
I prefer, then, to go lower down among the Ccelenterates
than Prof. Sedgwick, lower down either phylogenetically or
ontogenetically, for the origin of the Annelids—v. e. to forms
either primitive or larval, and before the mesenteries (which,
I think, developed as muscular bands drawing and holding in
the primitive mouth in order to form an cesophageal infolding*)
appeared.
The chief assumption which we have to make, then, is
that some free-swimming ciliated Coelenterate of the very
simplest type, instead of becoming early attached, continued
to be free-swimming long enough to put out buds. There is
nothing improbable in this. Dritted away by currents,
thousands must have missed finding anything to attach them-
selves to. In times before the seas swarmed with carnivorous
fish or Crustacea—both, I believe, descendants of the Worms
whose origin we are discussing—there would be abundance of
opportunity for some of these unattached forms to lead a
free-swimming life with safety, feeding on still simpler forms,
animal and vegetable.
* This method of deducing the Scyphozoa with ectodermal gullet from
the Hydrozoa without ectodermal gullet is to my mind preferable to the
belief, entertained by Sedewick and others, that the mesenteries were
developed primarily as folds for the increase of the digestive surface,
The differentiation of the muscles round the primitive mouth into
radiating bundles could hardly fail to result in the formation of internally
projecting folds, the body being but a flexible sac frequently distended by
fluid. The more strongly these ridges developed the more permanent
would be the tucking of the primitive oral aperture into the body, the
ectoderm around it becoming the lining of an cesophageal tube. The
advantages to the organism not only of this more perfect mouth, with its
stronger sphincter and radial muscles, but also of the increase in the
digestive surface afforded by the mesenterial folds, would lead to its
further development.
Origin of the Segmented Worms. 513
We merely assume, then, that a certain number of forms
remained free-swimming long enough to feed and to grow and
to put out buds. Now, without going so far as to say that
there is only one place where such a free-swimming form
could bud and yet continue to pursue its active life, viz. at its
hinder end, it is certainly clear that the most likely place for
a bud to appear would be at the posterior end. Not only
would any other place make forward locomotion impossible,
but at the hinder end the conditions seem to be more favour-
able than anywhere else. The external pressure would here,
in the wake of the animal, be less than anywhere else in the
body—indeed, during rapid forward swimming there would
be something like suction at this spot. Now if this is true
for the first bud, it is surely more true for the second, and it
seems clear that we get all the conditions for rapid serial
budding: for the addition of the first bud, by increasing the
number of available cilia, would propel the animal faster
through the water ; this greater speed would enable more food
to be swallowed, part of which would find its way into the
bud, while at the same time it would still further diminish
the external pressure of the water at the extreme hinder end,
and we have still more favourable conditions for the forma-
tion of a second bud behind that already formed, and so on.
If there is any probability in this argument at all, it would
seem that the more buds there were the faster would buds be
produced, until some limit, presently to be discovered, was
reached.
But apart from this argument, and merely bearing in mind
the wealth of colony formation developed among the Ccelen-
terates, is there not full justification for believing that round
the shores of the ancient seas, soon after the primitive forms
appeared, there would be free-swimming individuals trailing
behind them longer or shorter strings of buds diminishing in
size progressively backwards? Of these, some, we may fairly
assume, would sooner or later again give up more or less com-
pletely the laborious free-swimming life and become creeping
forms. It is in this direction that I should be inclined to
look for the ancestors of some of the other members of the
ancient class ‘‘ Vermes”’*.
* The Turbellarian Microstomum lineare, Ehr., still swims about trailing
short strings of buds after it. I was once fortunate enough to get a
glimpse of it living, under a high power, and saw very clearly what
appeared to be typical nematocysts scattered here and there in its skin.
The derivation of the Platihelminthes from primitive Coelenterates, which
gave up a free-swimming for a creeping manner of life, has much to be
514 Mr. H. M. Bernard on the
But confining our attention solely to the forms which we
assume persisted in their more active free-swimming habits,
a vista of possibilities opens out with regard to them which
seems to lead straight to the typical Annelid.
It appears at first sight as if there would be no limit to the
possible addition of fresh buds, so that, excluding accidents,
each original (or parent) animal would soon trail after it a
string of buds of indefinite length. Mechanical difficulties,
however, would most certainly sooner or later arise. The
most important difficulty would, it seems to me, be the
following :—The first-formed buds would have considerable
difficulty in getting rid of the indigestible remains of their
food. The parent animal, or first segment, was probably able
to get rid of feecal remains through the mouth; but this would
be very difficult for the buds, and, indeed, progressively
difficult the further back they were. The aperture leading
from sac to sac we may justly presume to have been, at least
primitively, very small, and it may perhaps have remained so
in the immediate ancestors of the segmented Worms. Before
long, therefore, particles of feecal matter would be mixed with
the food which found its way back into the last-formed bud,
until in time the fecal matter from the progressively increasing
number of digesting buds would make the arrival of food-
particles from the mouth impossible. Some nutriment might
for a while be extracted from this faecal matter, but the poste-
rior segments would eventually be so poorly nourished that
further budding would cease. This difficulty was hardly
likely to last long. Apertures can occur almost anywhere in
the Ceelenterate wall, as we know from the apertures in the
walls of the Sea-anemones for the discharge of acontia.
Hence a posterior opening might early be developed in order
to free the terminal buds from their useless and deleterious
burden *, The muscular fibres of the walls would provide the
said for it. The tentacles of Temnocephala may be secondary adaptations
to its sessile manner of life; but the fact discovered by Professor Haswell
(Quart. Journ. Micr. Sci. xxvili., 1888), that the young of the Australian
forms invariably have six, while the adult has only tive tentacles, is signi-
ficant rather of the opposite view. My esteemed friend and teacher
Prof. Arnold Lang (Text-book of Comp. Anat. pt. i.) also suggests an
origin from free-swimming Coelenterates which have secondarily adopted
a creeping manner of life, but along an entirely different line, viz. through
the Ctenophora, certain specialized forms of which furnishing almost ideal
intermediates.
* Cf. Prof. Sedgwick’s account (J. c.) of the possible origin of nephri-
dial apertures out of the constricted gastric pouches, which, according to
his theory, gave rise to the ccelomic chambers.
Origin of the Segmented Worms. 515
sphincters. Such an anal aperture, acquired of necessity by
the lengthening chain of digesting buds, would be earlier and
earlier developed by that force in living matter called heredity,
which causes it to repeat itself so long as the habits of life
demand.
This specialized aperture, however, would now be in the
exact position from which budding had formerly taken place.
Although division of highly organized cells is known in the
Protozoa, it is certainly commoner in cells not differentiated
for special functions. Hence we may suppose that the cells
which were specialized as epithelial, muscle-, or nerve-cells
to line and regulate the new opening would not divide, at
least actively, but the less differentiated zone of cells imme-
diately round these would, and by so doing push the specialized
anus out in front of them, so that it would always be at the
posterior end of the chain.
At first this anal aperture would have been very simple and
small, but as it became more highly specialized the budding
zone would retreat from the extreme tip until there appeared
a regular “anal segment,”’ the budding zone being always
between it and the last-formed bud. ‘This is, in fact, still
the law in all animals that develop metamerically—the new
segments are added between the anal “segment” and the one
in front of it.
The mechanical difficulty due to the accumulation of feces
being thus simply got over, the possibility of further advance
is instantly furnished ; more food would now be passed along,
and the posterior segments being better nourished, the budding
process at the end could be renewed. ‘The number of buds
would, however, still be limited, as, even though fecal matter
could now be got rid of, yet as the length increased it is
obvious that more fecal matter than food would reach the
posterior end of the body, which would thus be badly
nourished and cease to grow. This limit could only again be
surmounted by the development of a vascular system.
So far, then, we have arrived at long free-swimming strings
of Ceelenterate buds with a mouth at one end and an anus at
the other, and with a tendency to produce fresh buds up to a
limit between an anal prominence or segment and the last-
formed bud. ‘This brings us so near to the typical Annelids
that it is worth seeing whether the structural differences
which still separate them are insurmountable. Here I might
parenthetically remark that it is just such active free-
swimming organisms as those we have sketched which attain
to higher levels of organization. ‘They, least of all, would
516 Mr. H. M. Bernard on the
stand still in the midst of the developing life of the sea. We
have a right to expect that, if in the age of the Ccelenterates
any such forms existed, profound structural modifications for
the perfecting of their activities would take place in them.
The difficulty, and the only great difficulty, which lies in
the way of the deduction of the typical Annelid from the
hypothetical organism described above, lies in the fact, already
mentioned, that the former have paired ccelomic cavities.
The rise of these in the animal kingdom is still one of the
great riddles of zoology. Professor Sedgwick’s suggested
solution, which has been deservedly widely accepted, would,
I think, have satisfied me entirely had division of existing
segments been the rule for their multiplication. As it is,
their known method of production has always seemed to me
to point so unmistakably to serial budding, that nothing which
has been said on the subject has been able to deter me from
watching for clues in that direction. On the other hand, his
argument based upon the blastopore is very complete, and, so
far as I can see, can only be attacked as coming too much
within the field of purely developmental adaptations. I repeat,
then, that this paper will have some value if it suggest a simple
and natural line of development for these cavities—that is, a
line of development without resort to any hypothetical ‘change
of function,” which is one of the convenient stocks-in-trade of
the puzzled morphologist ;—if it show that in the further
development of strings of Ccelenterate buds as free-swimming
organisms some such mesodermal structures would inevitably
arise, fulfilling essentially the same function from their earliest
rudiments to their latest specialization.
We are safe in assuming that the body-cavities arose, as
all new structures arise in the animal kingdom, in response
to some physiological need. In the present instance we do
not require to look far for that need. In the struggle for
existence between such chains of Ccelenterates it is not likely
that the muscles in the walls would be long kept out from
assisting in locomotion. A waving of the body is a far more
efficient method of propulsion than the rowing of cilia. Hence
we should expect the wall-muscles in each bud to be gradually
specialized into systems in order to bring about definite
serpentine movements of the whole string, and, increasing in
size and strength, gradually to take on the whole function of
propulsion. But muscles require not only to be fed and
oxygenated, but also to be relieved of the waste products of
their activities. ‘The more they are used the more elaborate
must be the nutritive and excretory apparatus to enable them ~
Origin of the Segmented Worms. 517
to maintain their efficiency. Further, this growth and ever
increasing concentration of the muscular systems of the indi-
vidual buds would necessarily be accompanied by an ever
increasing development of their nervous systems. These also
would require to be well nourished and cleansed. The
feeding and the carrying away of the waste products of this
developing neuro-muscular sheath could hardly be carried on
in the original Ccelenterate wall. Structural modifications,
increasing in specialization as the systems they subserved
became more and more perfect, were bound to take place;
these, I think, we may safely outline. In the simpler stages
scattered cells with fluid spaces would appear in the mesen-
chymal layer; their increase would result in the development
of a parenchymatous tissue, with still larger fluid spaces, for
a compact tissue would be a hindrance to contractility.
These spaces would gradually separate the ectoderm from the
endoderm—. ¢., on the one hand, the ectodermal layer and
locomotory muscles from the digesting endoderm, the only
source of nourishment; and, on the other hand, the digesting
endoderm from the ectoderm, the only breathing surface.
Further advance along these lines would necessitate definite
streams, and ultimately a vascular system, collecting nutri-
ment from the digesting layer and oxygen from the ecto-
dermal. In this way, then, the endoderm and the ectoderm
would be gradually divorced from each other by an increasing
development of intervening tissue. The former (endoderm)
would be ultimately confined to a central hollow strand, the
alimentary canal, suspended and held in place by membranous
tissue and wrapped round by muscles and cells split off from
the early mesenchymatous or parenchymatous layer already
described. In passing we may note that though the deep
columnar epithelium characteristic of the alimentary canal of
the Annelids could doubtless be developed at any time, still
it is just possible that it may be an expression of the shrinkage
of the surface due to the detachment of the endoderm from
the outer wall and to its becoming the inner lining of a
narrowing axial tube. The suggestion arises naturally from
the remarkable difference seen between the endoderm of an
expanded and that of a contracted polyp.
Here, then, we have a very simple and natural origin for
the mesoderm in the Metazoa, the activities of our string of
buds supplying us with an adequate reason for its develop-
ment. ‘The case is, I think, a strong one, for the muscles in
each bud necessary to enable it to take its share in giving
to the whole string a serpentine movement would have to be
518 Mr. H. M. Bernard on the
far stronger than any which the bud itself would require had
it existed as a single animal. Similarly the nervous system
necessary to coordinate the movements of these muscles
within each bud and with those of the buds before and behind it
would have to be very much more highly developed than would
have been necessary in single animals the size of the buds or
segments. We need not be surprised, then, at the enormous
development of the mesodermal chambers. It is significant
here to note that the Turbellarians, which I am inclined to
regard as the nearest existing kindred of our early strings of
buds, having, with a few exceptions, lost the power of forming
buds, have their muscles and nerves simply embedded in a
parenchymatous tissue not unlike that which we imagine to
have been a stage in the development of the mesodermal
cavities of the Annelids. Neither muscles nor nerves in
these single animals required the same high specialization as
was required by the individuals forming the separate links
in a long chain which had to move as one, comparatively
speaking, immense composite organism. We thus have, in
the specially pronounced development of the neuro-muscular
sheath of each segment of our string of buds—a far more deve-
loped sheath, I repeat, than any single animal could possibly
require,—an efficient cause to account for so highly specialized
a nutritive and excretory apparatus as the mesoderm of the
Annelids, with its cavities, vascular system, and nephridial
funnels.
On turning to the embryological records of the mesoderm
we find them confused, and, for want of a clue, frequently
contradictory. We have the ccelomic cavities arising in
masses of parenchymatous tissue, or forming early out of very
few cells, or, again, as definite invaginations. All these fall
into their places in the light of the origin above assigned to
the mesoderm. ‘They are fragments and abbreviations of the
true phylogenetic history. The invaginations may be either
for the purpose of rapidly supplying cells or else as shortened
methods of forming the coelomic cavities. They obviously
serve both purposes.
We have here an excellent illustration of the difficulty
of discovering phylogeny from embryology. The parenchy-
matous mesoderm suggests little or nothing, whereas meso-
dermal invaginations seem so clearly to indicate some once
useful structures which, by “change of function,” might
have supplied the mesoderm, that they immediately acquire
a greater importance than really belongs to them. One
primitive function to which they have been attributed—besides
Origin of the Segmented Worms. O19
that suggested by Professor Sedgwick, that they were at one
time due to foldings of the digestive layer—is the reproductive.
But I cannot help remarking that the view here put forward,
which regards the mesoderm as arising by gradual structural
modifications in response to the simplest and most frequently
recurring of life’s activities, viz. to locomotion in pursuit of
food—that is, to locomotion which lasts from the beginning
to the end of life,—supplies us with a better motive for its
production than do the needs of the reproductive function,
which are periodic and confined to the adult. Further, I
need hardly again emphasize the fact that the suggested
origin put forward in this paper avoids all appeals to “ change
of function’’; from its earliest rudiments to the highest
specialization the mesoderm, if developed along the lines here
sketched, had essentially the same function.
With the perfection of the powerful neuro-muscular sheath
and the necessary mesodermal apparatus for its nutrition and
excretion we have brought our strings of Ccelenterate buds
near enough to the typical Annelid to show that, whether
this was the true origin of the Annelids or not, it could easily
have been so. It is admittedly only an outline sketch. A
much closer comparative study of kindred animals and of
their homologous structures would be necessary in order to
fill in details. One somewhat prominent point may, how-
ever, be noted.
For the purpose of bringing about the strong lateral con-
tractions of each segment necessary to give the whole complex
its serpentine motion, the longitudinal muscles would tend to
stretch from end to end of each segment. Their points of
insertion would consequently be more and more concentrated
at the ends of the buds, leaving the middle regions free. As
already noted, the ectoderm was the only available respiratory
surface. What more natural than that this middle region
should become specialized for respiration, the greatly developed
neuro-muscular system demanding for its efficiency ever-
increasing supplies of oxygen. ‘The necessary mechanism
for obtaining this extra supply was well within reach and the
structures developed would be from the first functional. I
refer to the lateral skin-folds which would arise in the middle
regions above mentioned every time the longitudinal muscles
contracted. From this point of view the parapodia, whatever
specialization and differentiation they may since have under-
gone, were originally nothing more than gill-folds, and they
arose concomitantly with the rise and concentration of the
powerful longitudinal muscles as these latter required more
and more oxygen for their work.
520 Mr. G. A. Boulenger on some
We can thus trace our rows of Ccelenterate buds along a
line of development, each step in which, so far as I can see,
would be a necessary result of their locomotory activities,
not as so many separate buds, but acting together as a single
complex organism. This line has led us, without strain or
difficulty, to the typical Annelid with its coelomic cavities and
its parapodia.
On page 2 I stated that I considered the metameric seg-
mentation so remarkable a phenomenon that wherever traces
of it can be discovered we are justified in assuming descent
from Annelidan ancestors. ‘The segmentation found so
widely spread in the vegetable kingdom is a parallel pheno-
menon, being due to periodical buddings. In the Vertebrata
we have organisms which show very distinct traces of seg-
mentation, but complicated by the fact that their most
characteristic structure, the notochord, shows no traces of
having been primitively segmented. The jointing which
later appears in the vertebral column has apparently been
impressed upon it as a mechanical necessity by the segmented
organism in which it developed. For an attempt to show
how the vertebrate organization with its primitively unseg-
mented notochord can be deduced along what appears to be
a similarly straightforward line of mechanical adaptations
from an annulate ancestor, see ‘‘ A new Reading in the Annu-
late Ancestry of the Vertebrata,” Natural Science, vol. xii.
1898, p. 17.
[Notre.—This paper was written eighteen months ago, and
was finally corrected for press before the appearance of
Prof. Lankester’s critical résumé of the subject in the recent
instalment of his ‘ T'ext-book of Zoology.’]
LXIX.—On some little-known African Silurid Fishes of the
Subfamily Doradine. By G. A. BouLEeNnGER, F.R.S.
Among the least-known freshwater fishes of Africa there is a
group of small or very small Silurids, more or less closely
allied to Synodontis and falling under Giinther’s division
“© Stenobranchie,” which have been described in a more or less
satisfactory manner by various authors under the generic
names of Mochocus, Rhinoglanis, Chiloglanis, Doumea, Atopo-
chilus, and Peltura. Having lately had the privilege of
examining the few specimens of these fishes preserved in the
little-known African Silurid Fishes. 521
Paris, Hamburg, and Genoa Museums, and having received
examples of new or rare species from the Nile and Congo, I
have attempted to meet a long-felt requirement by drawing
up original comparative descriptions of all the genera and
species.
Chiloglanis Pers. Mochocus, Joannis.
Deckenii, Ptrs. niloticus, Joannis,
niloticus, Blgr. Doumea, Sawv.
Euchilichthys, Bigr. typica, Sawv.
Guentheri, Schilth. Phractura, Bigr.
Dybowskii, Vail. Bovei, Perugia.
Atopochilus, Sawo. scaphirhynchura, Vail.
Savorgnani, Saue. Andersonia, Blgr.
leptura, Bigr.
1. CHILOGLANIS.
Chiloglanis, Peters, Mon. Berl. Ak. 1868, p. 599.
Body moderately elongate, feebly depressed. Anterior
dorsal entirely in advance of ventrals, formed of 1 spine and
5 branched rays ; an adipose fin opposed to the anal; pectoral
with a spine ; ventral with 7 rays. A cephalo-nuchal bony
shield. Mouth inferior, surrounded by a very large, circular,
papillose lip; conical teeth on the premaxillaries and on the
vomer * ; movable, slender, curved teeth in the mandible; a
maxillary and two mandibular barbels on each side ; nostrils
rather remote from each other, both with a valve, the ante-
rior tubular; eyes small, without free border. Gill-clefts
short, confined to the sides, Air-bladder small, partially
enclosed on each side between the enlarged processes of the
anterior vertebra.
1. Chiloglanis Deckenti.
Chiloglanis Deckenti, Peters, Mon. Berl. Ak. 1868, p. 599, pl. ii.;
Pfeffer, Fische O. Afr. p. 58, fig. (1896).
Synodontis eurystomus, Pfeffer, Jahrb. Hamb. Wiss. Anst. vi. 1889,
p. 14, and x. 1893, p. 31, pl. 1. fig. 5.
Body slightly depressed, its depth 5} to 64 times in total
length. Head strongly depressed, a little longer than broad,
its length 3 to 34 times in total length. Eye directed up-
wards, in the second half of the head, its diameter about 5
times in length of head and 1} in interorbital width, which
equals the distance between the eye and the nostril; two
widely separated oval groups of pramaxillary teeth, forming
3 or 4 transverse series ; a series of 6 to 8 short mandibular
* “ Premaxillary teeth” of Peters.
522 Mr. G. A. Boulenger on some
teeth ; maxillary barbel about 4 length of head, a little longer
than the lower labials. Dorsal I 5; spine not distinctly
serrated, about § length of head. Adipose fin 3 to 4 its
distance from the rayed dorsal. Anal III 6. Pectoral spine
a little longer than the dorsal. Ventral extending to beyond
origin of anal. Caudal forked. Caudal peduncle nearly
twice as long as deep. Olive-brown, with irregular darker
cross-bands; a dark bar at the base of the caudal and another
across each lobe of the fin.
Total length 70 millim.
German Hast Africa (Korogwe, on the Rufu River).
Type in Berlin Museum.
Thanks to the kindness of Prof. Pfeffer, one of the speci-
mens obtained by Stuhlmann at Korogwe is now in the
British Museum.
2. Chiloglanis niloticus, sp. n.
Distinguished from C. Deckenit in the following cha-
racters :—
Depth of body 4 to 43 times in total length. Diameter of
eye 6 times in length of head; interorbital width much
greater than the distance between the eye and the nostril.
Dorsal I 6, the spine about ? length of head. Pectoral spine
3 length of head. Anal III 8. Pale greyish olive above,
yellowish beneath; four broad blackish cross-bands on the
body ; a horizontal blackish bar on each lobe of the caudal.
Total length 45 millim.
Several specimens were obtained in May last by Mr. W. L.8.
Loat whilst on the island of Arko, Soudan.
2, EUCHILICHTHYS, gen. nov.
Body moderately elongate, not or but feebly compressed.
Anterior dorsal entirely in advance of ventrals, formed of 1
spine and 6 branched rays; an adipose fin opposed to the
anal; pectoral with aspine; ventral with 7 rays. A cephalo-
nuchal bony shield. Mouth large, inferior, surrounded by a
much-developed, circular, papillose lip; a broad band of
pointed, curved, movable teeth in the preemaxillaries, this
band entirely in advance of the narrower band of mandibular
teeth, the base of which is long and slender, the crown trun-
cate or notched and bent up. hook-like; conical teeth on the
vomer; a barbel on each side, and a pair of very short ones
on each side of the lower section of the circular lip; nostrils
little-known African Silurid Fishes. 523
rather close together, the posterior with a valve; eyes with
free border. Gill-clefts short, confined to the sides, Air-
bladder small, partially enclosed on each side between the
enlarged processes of the anterior vertebree.
Distinguished from Chiloglanis by the angularly bent,
truncate or bicuspid mandibular teeth, which form a broad
band ; from Atopochilus by the pointed premaxillary teeth
so different from those on the mandible.
Two species.
1. Euchilichthys Guentheri.
A topochilus Guenther, Schilthuis, Tijdschr. Nederl. Dierk. Ver. (2)
ill. 1891, p. 86, pl. vi. fig. 2.
Depth of body 3 its greatest width, 7 times in total length.
Head large, much depressed, covered with soft skin, a little
longer than broad, its length 34 times in total length; snout
broadly rounded, nearly twice as long as postocular part of
head ; nostrils nearer end of snout than eye, the diameter of
which is 5 times in interocular width; buccal cleft straight,
nearly half length of head; lateral barbel + length of head,
posterior barbels about half diameter of eye. D. I 6, origi-
nating immediately behind pectorals; spine strong, covered
with skin, measuring + length of head. Adipose fin very
short. A. 10 (38 raysrudimentary). Pectoral spine 4 length
of head, its inner border with strong retrorse spines. Ven-
trals not reaching anal. Caudal forked. Caudal peduncle
14. as long as deep. Skin villose. Dark brown, with small
round black spots on the anterior parts.
Total length 190 millim.
The single known specimen, from the Stanley Pool, Congo,
is now in the British Museum.
2, Huchilichthys Dybowskii.
Chiloglanis Dybowskz?, Vaillant, Bull. Soc. Philom, 1892, no. 16, p. 2.
Body a little deeper than broad, its depth 44 times in total
length. Head large, moderately depressed, covered with a
very thin skin, not quite 1} as long as broad, its length 23
times in total length; snout rounded, nearly twice as long as
postocular part of head; nostrils nearer end of snout than
eye, the diameter of which is 5 to 54 times in length of head,
twice in interocular width ; buccal cleft straight, ? length of
head ; lateral barbel nearly } length of head, posterior shorter
still. D.16; spine strong and smooth, 2 length of head.
524 Mr. G. A. Boulenger on some
Adipose fin much longer than deep, nearly as long as its
distance from the first dorsal. A. 10 (8 rays rudimentary).
Pectoral spine 2 length of head, its inner border with 4 or 5
strong retrorse spines. Ventrals extending a little beyond
origin of anal. Caudal deeply emarginate. Caudal peduncle
as long as deep. Skin villose on the sides of the body.
Pale brown; a blackish band across the dorsal, another across
the pectorals ; ventrals and anal blackish.
Total length 39 millim.
The two type specimens, from the Ubanghi, French Congo,
are preserved in the Paris Museum.
3. ATOPOCHILUS.
Atopochilus, Sauvage, Bull. Soc. Philom. (7) iii, 1878, p. 96, and N,
Arch. Mus. (2) iii. 1880, p. 42.
Body moderately elongate, feebly depressed. Anterior
dorsal entirely in advance of ventrals, formed of 1 spine and
6 branched rays; an adipose fin opposed to the anal, which
is short ; pectoral with a strong spine; ventral with 7 rays.
A cephalo-nuchal bony shield. Mouth large, inferior, sur-
rounded by a much developed, circular, papillose lip; teeth
fine, slender, movable, truncate, bent at an obtuse angle and
forming a very broad band in the premaxillaries, this band
entirely in advance of the narrower band of mandibular teeth,
which are similar to the premaxillaries, but closer together ;
conical teeth on the vomer; a barbel on each side, and another
pair on the posterior border of the lip; nostrils close together,
the posterior with a valve; eyes with free border. Gill-
clefts short, confined to the sides.
A single species.
1. Atopochilus Savorgnant.
Atopochilus Savorgnani, Sauvage, ll, cc. pl. iii. fig. 3.
Depth of body a little less than its greatest width, 52 times
in total length. Head depressed, 14 as long as deep, a little
longer than broad, its length 32 tines in total length ; upper
surface of head a little rugose; snout broadly rounded, its
length 24 postocular part of head; nostrils nearer end of
snout than eye, the diameter of which is 43 times in length
of head, 13 in interorbital width ; buccal cleft straight, ? length
of head; band of mandibular teeth not half as broad as that
of premaxillaries ; lateral barbel + length of head, more than
twice as long as posterior barbel. Occipito-nuchal shield as
long as broad, ending in two points. A striated, acutely
little-known African Silurid Fishes. 525
pointed humeral process. D.I 6; spine striated, 3 length of
head. Adipose fin 3 times as long as deep, 3 its distance
from rayed dorsal. A.11 (2 rays rudimentary). Pectoral
with a strong striated spine bearing 12 retrorse teeth on its
inner border, its length ? that of head. Ventral nearly
reaching origin of anal. Caudal peduncle 13 as long as
deep *.
Total length 90 millim.
The single specimen, preserved in the Paris Museum, is
from the falls of Doumé, Upper Ogowe.
4, Mocuocus.
Mochokus, Joannis, Mag. Zool. 1835, Poiss.; Giinth. Cat. Fish. vy:
p. 217 (1864). .
Rhinoglanis, Giinth. t. ¢. p. 216.
Body moderately elongate, slightly compressed. Anterior
dorsal ending over the ventrals, formed of a strong spine and
7 or 8 branched rays ; a second dorsal, formed of soft rays,
above the short anal; pectoral with a strong spine; ventral
with 7 rays. A cephalo-nuchal bony shield. Mouth sub-
inferior, without labial folds; small conical teeth in both
jaws; a maxillary and two mandibular barbels on each side;
nostrils rather remote from each other, both with a valve, the
posterior very large; eyes moderate, without free border.
Gill-clefts short, confined to the sides. Air-bladder large,
free.
1. Mochocus niloticus.
Mochokus niloticus, Joannis, 7. ¢. pl. viii.
Rhinoglanis typus, Giinth. J. c. p. 216, fig., and in Petherick's Trav. ii.
p. 287 (1869).
Rhinoglanis Vannutelliz, Vinciguerra, Ann. Mus. Genova, (2) xix. 1898,
p. 254.
Depth of body 53 to 5§ times in total length, length of
head 4 to 44. Head depressed, as long as broad, rugose
above; a large frontal fontanelle ; eye directed upwards, 3 to
34 times in length of head, not much narrower than inter-
orbital width ; barbels long and slender, the maxillary reach-
ing end of pectoral spine or beyond. Anterior dorsal I 7-8 ;
spine nearly as long as head, with ascending serre on anterior
border ; second dorsal 10-15, about as long as its distance
from the first. Anal 9-10. Pectoral spine a little longer
* The caudal fin is injured in the type specimen; it was probably
notched or forked, not truncate, as figured by Sauvage,
Ann. & Mag. N. list. Ser. 7. Vol. vi. ao
526 Mr. G. A. Boulenger on some
than dorsal, serrated like it on outer border, with very strong
retrorse teeth on inner border. A long, sharp, humeral pro-
cess. Ventral not reaching anal. Caudal forked. Caudal
peduncle nearly twice as long as deep. Pale greyish olive
above, speckled and marbled with blackish ; two more or less
distinct, irregular, dark bands across the back, one under each
of the dorsal fins ; belly white; dorsals and caudal with
blackish dots.
Total length 55 millim.
The type specimen is not to be found in the Paris
Museum.
The figures given by Joannis are very incorrect, as may be
inferred from the discrepancies between them. A comparison
of specimens obtained by Mr, Loat at Assuan with the type
of Rhinoglanis typus, from Gondokoro, and those of R. Van-
nutellii, trom Lake Rudolf, has convinced me that these
supposed species cannot be maintained as distinct from
Mochocus niloticus.
5. DouMEA.
Doumea, Sauvage, Bull. Soc. Philom. (7) iii. 1878, p. 96, and N. Arch.
du Mus. (2) iii. 1880, p. 41.
Body elongate, depressed, with extremely slender caudal
peduncle. ‘Two short dorsal fins, the anterior entirely in
advance of the ventrals and formed of one simple, flexible,
and 6 branched rays, the second opposed to the anal and
formed of fine soft rays partly concealed in adipose tissue ;
pectoral and ventral fins large, horizontally expanded, with-
out spine, but with thickened, simple outer ray; 12 rays to
the pectoral, 6 to the ventral, No cephalo-nuchal shield.
Mouth small, inferior, surrounded with large papillose he
no mandibular teeth ; small conical premaxillary teeth * ; a
maxillary and two mandibular barbels on each side; nostrils
rather remote from each other, both with a valve; eyes
small, without free border. Gill-clefts narrowly interrupted
at the isthmus. ;
A single species.
Sauvage referred this genus to the ‘ Pimelodine”’ of
Giinther. Vaillant (Bull. Mus. Paris, 1897, p. 82), on the
contrary, referred it to the vicinity of Synodontis, a view in
which I entirely concur.
* The ‘“ vyomerine ” teeth of Sauvage.
little-known African Silurid Fishes. 527
1. Doumea typica.
Doumea typica, Sauv. ll. ce. pl. iii. fig. 1.
Depth of body slightly less than its width, 8 times in total
length. Head strongly depressed, with slightly rugose skull
covered with a thin skin, 1} as long as broad, 5} times in
total length ; snout pointed, projecting a little beyond the
mouth, 12 as long as postorbital part of head; internarial
space equally distant from end of snout and eye; latter
directed upwards, its diameter 64 times in length of head,
1} in interorbital width; maxillary barbel 4 length of head,
mandibular barbels shorter stil!. Occipital process narrow,
3 times as long as broad, not reaching interneural shield.
Dorsal I 6, first ray a little longer than the head. Anal I 7.
Pectorals reaching the ventrals, which extend far beyond
origin of anal. Caudal with crescentic notch. Caudal
peduncle a little depressed, 3 total length. Uniform brownish
in spirit.
Total length 90 millim.
The unique specimen in the Paris Museum is from the
falls of Doumé, Upper Ogowe.
6. PHRACTURA, n. n.
Peltura (non M.-Edwards), Perugia, Ann. Mus. Genova, (2) x. 1892»
p-. 972
Body elongate, depressed, with extremely slender caudal —
peduncle; a series of imbricate scutes along each side of the
back and belly from the dorsal and ventral fins to the caudal
peduncle, which is entirely surrounded by bony scutes. Two
short dorsal fins, the anterior entirely in advance of the
ventrals and formed of one simple, flexible, and 6 or 7
branched rays, the second opposed to the anal and formed of
fine soft rays; pectoral and ventral fins large, horizontally
expanded, without spine, but with thickened, simple outer
ray; 10 rays to the pectoral, 6 to the ventral. No cephalo-
nuchal shield. Mouth small, inferior, surrounded with large
papillose lips; no mandibular teeth; small conical pre-
maxillary teeth; a maxillary and two mandibular barbels
on each side; nostrils rather remote from each other, both
with a valve; eyes small, without free border. Gill-clefts
narrowly interrupted at the isthmus.
Two species.
35*
528 Mr. G. A. Boulenger on some
1. Phractura Bovei.
Peltura Bovei, Perugia, /. c. fig.
Depth of body 9 times in total length, length of head 6
times. Head 13 as long as broad; skull a little rugose,
covered with thin skin; snout a little longer than the post-
ocular region of the head, pointed, projecting beyond the
mouth; the space between the nostrils equally distant from
the end of the snout and the eye, which is directed upwards ;
the diameter of the eye 10 times in the length of the head
and 24 in the interocular width; maxillary barbels 3 length
of head, nearly reaching the gill-cleft, mandibular barbels
much shorter. Occipital process 3 times as long as broad,
not reaching interneural shield. Dorsal 17; first ray longest,
+ longer than the head; second dorsal very small. Anal 17.
Pectoral a little longer than the head, as long as the ventral ;
latter nearly reaching the anal. Caudal with crescentic notch.
Caudal peduncle depressed, 3 total length. 23 dorsal and 19
ventral scutes, the last 5 on the caudal peduncle. Uniform
yellowish in spirit.
Total length 110 millim.
Lower Congo. Type in Genoa Civic Museum.
2. Phractura scaphirhynchura.
see ee
D.ffers in the following points from the preceding species :—
Length of head 5 times in total length, 14 its width; snout
twice as long as postocular part of head; diameter of eye
7 times in length of head, 1} in interocular width; maxillary
barbel 7 length of head, 3 the distance separating its root
from the gill-cleft. Occipital process 24 as long as broad,
reaching interneural shield. Dorsal I 6, first ray as long as
head. Pectoral not longer than head. 22 dorsal and 18
ventral scutes.
Total length 120 millim.
Diélé, on Alima R., French Congo. Type in Paris
Museum.
7. ANDERSONIA, gen. nov.
Body elongate, depressed, with extremely slender caudal
peduncle; a series of imbricate scales along each side of the
back and belly from the dorsal and ventral fins to the caudal
peduncle, which is entirely surrounded by bony scutes. Two
short dorsal fins, the anterior above the ventrals and formed
little-known African Silurid Fishes. 529
of one simple ossified ray or spine and 6 branched rays, the .
second opposed to the anal and formed of fine soft rays pre-
ceded by a short spine ; pectoral and ventral fins moderately
large, the former with a spine; 8 rays to the pectoral, 6 to
the ventral. No cephalo-nuchal shield. Mouth small, in-
ferior, toothless ; lips not much developed; a maxillary and
a mandibular barbel on each side; nostrils remote from each
other, both with a valve; eyes small, without free border.
Gill-clefts narrowly interrupted at the isthmus.
Nearest allied to Phractura, from which it is well distin-
guished by the position of the anterior dorsal, the presence
of a spine in front of each of the dorsals, the presence of a
single mandibular barbel, and the total absence of teeth.
Named in memory of the late Dr. John Anderson, to whose
exertions during the latter years of his lite Science is indebted
for much progress in the zoology of the Nile region, and to
whose initiative we owe the organization of a survey of the
Nile Fishes which is now being carried on by the Keyptian
Government.
1. Andersonia leptura, sp. n.
Depth of body 9 times in total length, length of head
5 times. Head 1} as long as broad; skull rugose; snout as
long as the postocular part of the head, thrice as long as the
diameter of the eye, which is half the interorbital width ;
barbels slender, as long as the snout. Occipital process
keeled, 24 as long as broad, widely separated from the inter-
neural shield ; a shorter process on each side of the occipital
process. Anterior dorsal I 6; first ray longest, nearly as
long asthe head. Anal I 9. Pectoral § length of head, as
long as the ventral, widely separated from the anal. Caudal
with crescentic notch. Caudal peduncle depressed, 4 total
length. 25 dorsal and 23 ventral scutes, the last 9 on the
caudal peduncle. Greyish above, marbled with blackish,
white beneath.
Total length 50 millim.
A single specimen was found by Mr. Loat ina pond near
Koshek, Soudan, in March 1900.
530 Mr. P. Cameron on new
LXX.—Deseriptions of new Genera and Species of
Hymenoptera. By P. CAMERON.
[Concluded from p. 506. }
Rthynchium khasianum, sp. n.
Nigrum ; clypeo, scapo antennarum subtus, pronoto late pedibusque
ferrugineis ; tarsis posterioribus nigris; alis violaceis. 92.
Long. 17-18 mm.
Hab. Khasia Hills. Coll. Rothney.
Antenne black, the scape rufons, spotted with black on the
top. Head black, rugosely punctured; the clypeus rufous.
On the vertex behind the ocelli are two large, deep, smooth
fover, the space between them being smooth. Antennal
keel smooth and shining, dilated at the top, and with a small
yellow mark on either side of the top. Clypeus rufous, longi-
tudinally punctured ; its apex wide, not much narrower than
the base; the extreme apex depressed and with a broad slight
curve. Mandibles rufous, black on the upperside. Palpi
black, sparsely covered with stiff hairs. ‘The base of the
pronotum is smooth and shining; the upper half of the sides
rufous, as are also the tubercles, a spot on either side of
the metanotum, a line on the mesonotum near the tegule, and
the tegula. Mesonotum closely and strongly punctured; the
extreme base smooth and distinctly separated from the pro-
notum. There is a bordered furrow on the basal half of the
scutellum in the middle. Median segment rugosely punc-
tured, the middle smooth, alutaceous; the basal half furrowed
in the middle, the apical keeled ; the top and middle of the
mesopleure closely rugosely punctured ; there is a wide deep
furrow on the top of the smooth basal part, with a keel down
its middle. Metapleuree deeply rugosely reticulated. Legs
ferruginous, the tarsi blackish; the fore tarsi rufous at the
base ; the fore coxe for the greater part black. The basal
slope of the petiole smooth; the rest of it and the second
segment closely punctured ; the third segment is less strongly
punctured, smoother at the base; the fourth and fifth pune-
tured, smooth at the apex.
Belongs to Bingham’s group “ A. Basal segment of abdo-
men slightly constricted, the apex distinctly narrower than
the base ot the second segment.” What appears fo be a
variety has the rufous band on the pronotum obliterated, and
there is a yellow mark on either side of the clypeus at the top.
Genera and Species of Hymenoptera. 531
Rhynchium elypeatum, sp. n.
Nigrum, dense argenteo-pilosum, basi medioque clypei albo ; alis
violaceis. <o.
Long. 14 mm.
Hab. Barrackpore, Bengal (Rothney).
Antenne black, the flagellum slightly brownish beneath ;
the claw brownish, stout. Head strongly and closely punc-
tured, thickly (especially on the face and outer lower orbits)
covered with silvery pubescence. Clypeus large, its greatest
width as long as its length, its base broadly rounded, its apex
roundly incised, the sides forming triangular teeth ; its upper
third, and the lower half broadly in the centre, yellow, the
apical yellow mark being separated from the basal yellow
part ; the edges of the black separation are irregular; the
Inner orbits opposite the antenne are yellow. ‘Thorax ru-
gosely punctured, covered with a silvery pile; the lower part
of the propleure bear four stout clearly separated keels. ‘The
base of the mesopleure is smooth; in the middle is a longi-
tudinal furrow ; above this the punctures are somewhat more
widely separated; the lower part of the apex is smooth,
The upper part of the metapleure above the hollow is longi-
tudinally striated; the lower part is smooth and impunctate ;
the upper part of the apex is strongly but not closely punc-
tured; the lower part is smooth. ‘The sides of the median
segment are broadly rounded above and have no teeth.
Wings violaceous, the anterior pair deeper in tint than the
posterior ; the second cubital cellule is much narrowed at the
top, about one sixth of the length of the third and about half
the length of the space bounded by the first transverse cubital
and the recurrent nervures. Abdomen thickly covered with
silvery pubescence, sparsely punctured; the ventral surface
with the punctures stronger and closer.
Comes nearest to R.argentatum, Fabr. (sec. Saussure, Stett.
ent. Zeit. xxiii. p. 187,=metaliicum, Sauss. Htudes, i. 114),
but is larger, has the clypeus broader compared to its length,
more rounded on the top, and yellow in the middle at the
apex ; the thorax is less strongly punctured ; the sides of the
median segment bulge out more and the apex is more exca-
vated therefore in the middle; the third cubital cellule is of
nearly equal width at the top and bottom, while in 2. argen-
tatum itis distinctly wider at the top than at the bottom, and
the hind wings are not so clearly hyaline compared with the
anterior.
Rk. argentatum, Bingham (‘ Fauna of India,’ Hym. i.
532 Mr. P. Cameron on new
p. 858), is a different species from the argentatum of Saussure’s
Revision of the East Indian Odynerides (Stett. ent. Zeit. xxiu.
pp- 129 and 207). It is probably a form of 2. bengalense,
Sauss., which=W. argentatum, Saussure, Etudes, i. p. 115,
non Fabr., sec. Sauss. Stett. ent. Zeit. .c. p. 191. Col. Bing-
ham does not appear to have been acquainted with the
above-mentioned Revision. 2. carbonarium, Sauss., from
Tranguebar, mentioned by Bingham shortly in a footnote
(2. c. p. 858), is described in full by Saussure (Stett. ent, Zeit.
i,c. p, 183). It is a Pararhynchium.
Rhynchium basimacula, Cam.
This is a good and distinct species, and not a variety of
R. flavomarginatum, as stated by Bingham (* Fauna of India,’
Hym. i. p. 859). It is easily known from flavomarginatum
by the head and thorax not being “ coarsely and very closely ”
punctured, the clypeus in particular having only some small
scattered punctures; 1t is much broader at the apex, almost
transverse, the sides not projecting into teeth; the abdomen
is impunctate, very smooth, and shining. The form of the
clypeus is different from what it is in most of the species, it
being not roundly convex, but flat, except at the sides, which
have an oblique slope, ‘The entire insect is thickly pruinose,
which gives it a whitish appearance, ‘he only species with
such a broad and flat clypeus is 2. dugubrinum, which, how-
ever, differs from it in having the head, thorax, and base of
abdomen closely rugosely punctured.
Rhynchium lugubrinum, sp. n.
Nigrum ; abdominis segmentis basalibus duobus flavo-lineatis ; alis
fulvo-violaceis ; tegulis rufis. @Q.
Long. 14-15 mm.
Hab. Khasia Hills. Coll. Rothney.
Antenne and head entirely black. Front and vertex
rugosely punctured, thickly covered with longish fuscous
hair. Antennal keel stout. Clypeus slightly obliquely
narrowed at the apex; the sides slightly obliquely depressed,
closely rugosely punctured, the apex with a rounded incision,
closely punctured and thickly covered with silvery pubes-
cence ; the extreme apex depressed and punctured. Pro- and
mesothorax closely and almost uniformly punctured; the
mesopleuree above with the punctures running into reticula-
tions; the basal half of the metapleures smooth, the apical
reticulated ; the upper part of the median segment reticulated,
Genera and Species of Hymenoptera. 533
the lower in the middle smooth except for some transverse
strie, the centre carinate. Tlie whole thorax is thickly
covered with pale pubescence. ‘The second cubital cellule is
much narrowed at the top, being there in length less than the
distance bounded by the second transverse cubital and the
second recurrent nervure. Legs entirely black, thickly
covered with pale pubescence. ‘Tegule rufous, Abdomen
black, the basal two segments banded with yellow, the bands
interrupted in the middle; the basal two segments closely
punctured, the third and fourth more closely, the apical
alutaceous.
The apex of the clypeus is broader than usual and does not
project into teeth; it is flat, with the sides oblique; the
insect is more densely pilose than usual; the apex of the
median segment is slightly and roundly concave; the upper
part at the sides broad, not forming teeth; the central keel is
stout ; the second cubital cellule is much narrowed at the top,
being there only one half the length of the space bounded by
the second transverse cubital and the second recurrent nervure ;
the second abdominal segment is not much narrowed at the
base.
This species comes nearest to R. flavomarginatum, with
which it agrees in coloration ; it may be known from it by
the body being more densely pilose, and is readily separated
zat it by the clypeus not being deeply grooved in the
middle.
Rhynchium rugolatum, sp. n.
Long. 12-14 mm.
Hab. Barrackpore (fothney), Poona (Wroughton), Khasia
Hills.
This species closely resembles &. brunneum. The differ-
ences between the two are best expressed in synoptical form :—
@.—Mesonotum shining, sparsely and slightly punc-
tured ; the scutellum impunctate ; the sides of the
median segment with sharp teeth .............. brunneum.
Mesonctum and scutellum closely rugosely punec-
tured; the sides of the median segment without
SC Men errata fee al Aenea fos dics catia tcs © Ae Se a eons rugolatum.
¢ .—The top of the clypeus transverse ; the oblique lower
part shorter than the upper part; the middle femora
not narrowed and twisted at the base .......... rugolatum,
The top of the clypeus rounded, its narrowed lower
part longer than the upper; the middle femora
narrowed and twisted at the base .............. brunneum,
The head rufous; the ocellar region broadly (the mark
slightly narrowed and rounded behind), the part between the
534 Mr. P. Cameron on new
antenne, a broad, short, oblique line over each, and the ccciput,
black. Front and vertex coarsely closely punctured, thickly
covered with short white hair. Clypeus pyriform, broadly
rounded above, sparsely and somewhat strongly punctured ;
the apex in the middle depressed and with a shallow incision ;
the sides forming oblique stout teeth. Thorax rufous, coarsely
and closely punctured except at the base and apex ; the lower
part of the pronotum, the edges of the mesonotum (narrowly
at the base and sides, broader at the apex), the base broadly
in the middle, the apex of the scutellum, the sides of the post-
scutellum, the middle of the metanotum broadly, its sides, the
base of the pronotum narrowly, the base and apex of the
mesopleure, a large irregular mark, a line along the furrow,
the base and lower half of the metapleure, and the breast
black. The scutellum is more closely punctured than the
mesonotum ; the postscutellum has behind and above the
oblique apical slope a stout somewhat irregular ridge; the
apex of the segment has an oblique slope ; it is smooth and
shining, the centre hollowed, the sides oblique ; in the middle
is a keel bordered by fine striae. Pro- and mesopleura
coarsely and closely punctured, except at the base; meta-
pleuree smooth, sparsely punctured; there are no teeth on the
sides; the apex roundly dilated. Legs rufous; the four
anterior coxee behind, the hinder almost entirely, the fore
femora narrowly at the base behind, the middle narrowly at
the base in front and broadly behind, and the posterior except
at theapex black. Wings yellowish, fuscous along the edge ;
the apex with a violaceous cloud; the costa and stigma
rufous, the nervures black ; the hinder wings hyaline. Ab-
domen rufous; the basal slope of the petiole, an irregular
line on the base of the second segment, and the bases of the
third and fourth segments black ; the basal ventral segments
are more or less marked with black.
In the male the black on the vertex is broader and extends
to the eyes; the part below it is yellowish, tinged with
rufous; the oblique marks above the antenne are as in the
female, but there is no black between the antenne; the
clypeus is as wide as its longest length, is smooth, except for
some punctures; the oblique lower part is slightly shorter
than the upper; the incision is shallower and wider than in
the female, and the part above it is not depressed; the
markings as in the female.
In general coloration it more resembles carnaticum, with
which it also agrees in the form of the male middle femora ;
but in the form of the clypeus &c. it differs, as it does also
from brunneum.
Genera and Species of Hymenoptera. 535
Montezuma bisulcata, sp. n.
Nigra, sparse albo-maculata ; alis fusco-violaceis ; scapo antennarum
basique mandibularum albo-maculatis. ©.
Long. 14-15 mm.
Hab. Khasia Hills. Coll. Rothney.
Scape of antenne sparsely and minutely punctured, yel-
lowish beneath. Front and vertex rugosely punctured, the
vertex more sparsely than the front; the part between the
antenne broadly carinate; foveate above, a yellow mark on
either side of the fovea. Clypeus closely rugose, the apex
with scattered punctures ; depressed and roundly and distinctly
incised at the point. Mandibles with a small yellow mark in
the middle near the base, its lower side obliquely widened at
base and apex, the apical part being the longer. Thorax
black, except for an interrupted line on the pronotum and a
mark on either side of the apex of the median segment.
Pro- and mesonotum closely rugosely punctured; the scu-
tellum, if anything, more closely, almost rugosely, punctured,
and with a shining line, gradually widened towards the apex
in the middle; the postscutellum has the punctures, if any-
thing, more widely separated than on the scutellum; its
centre is smooth and has an elongated shallow depression in
the middle. Pro- and mesopleure coarsely rugosely punc-
tured, the punctures on the latter running into reticulations ;
the apex is alutaceous, impunctate. Metapleuree alutaceous,
the apex obscurely obliquely striated; on the base are five
fovex, separated by stout keels, and having at the top a much
larger more elongated depression. Wings fuscous-violaceous,
the apical half of the posterior more hyaline. Legs black, a
line on the basal two thirds of the anterior tibize in front and a
line on the base of the posterior behind yellow; the front
femora are similarly marked on the apical two thirds in
front. Abdomen black ; a line, interrupted in the middle, on
the apex of the petiole, and three lines on the apex of the
second segment, yellow. Petiole above closely rugosely
punctured ; a shallow indistinct furrow on the apex in the
middle above; the base sparsely covered with long white
hair; the lower side is regularly and stoutly transversely
striated. The second segment is closely but not very deeply
punctured; the third segment is slightly depressed and
punctured on the apex.
The insect is covered with a pale pile. On the median
segment the central furrow is large and deep and is divided
into two parts by a transverse partition—a shorter basal,
536 , Mr. P. Cameron on new
rounded and narrowed at the base and apex, and a longer,
wider apical one, with a stout keel in the middle. The
tegule are brownish on the outer side.
Allied to WM. burmanica, Bingh.
Labus armatus, sp. n.
Niger; clypeo, scapo antennarum subtus, basi mandibularum, linea
pronoti, maculisque 2 scutelli flavis ; alis fusco-hyalinis, stigmate
nervisque nigris.
Long. 9mm. 6.
flab, Khasia Hills. Coll. Rothney.
Antenne covered with a pale down; black, the scape
yellow below; the terminal hook rufous. Head black; the
elypeus (except narrowly round the edges) and the base of
the mandibles to near the second tooth lemon-yellow. Front
and vertex closely and somewhat strongly and uniformly
punctured all over; covered with silvery pubescence; the
lower part of the front and the eye-incision covered with
silvery pubescence. Clypeus sparsely punctured; the apex
obliquely narrowed from the eyes to the middle, where there
are two minute black teeth. On the mandibles are one large
apical tooth, followed by two blunter, more rounded ones, of
half the length of the apical; the third has a much shorter
tooth or tubercle near its base. Palpi black. Thorax black ;
a broad line on the pronotum with its teeth, two marks on
the base of the scutellum, rounded and slightly narrowed on
the apex, from the outer side to the inner, yellow. Pro- and
mesonotum closely, uniformly, and rather strongly punctured.
Scutellum more sparsely punctured than the mesonotum,
The tooth on the postscutellum large, sharply pointed towards
the apex. Median segment sparsely punctured; the central
furrow becoming gradually wider towards the apex. Pro-
pleure strongly punctured above, below smooth, aciculated ;
mesopleure similarly punctured, except at the base beneath ;
metapleuree more sparsely punctured than the meso-, espe-
cially towards the base. Legs black, the four anterior tibiz
in front and the basal joint of the four front tarsi yellow.
Abdomen black; the apex of the petiole above and of the
second segment all round yellow; the petiole has the basal
half coarsely punctured ; the punctures are large and longer
than broad; below, the punctures extend nearer to the apex.
‘The whole insect is covered with a pale pile. Wings fuscous-
hyaline; the stigma and nervures black; the apex of the
radius is bluntly rounded, it is distinctly curved upwards at
the second cubital cellule; the tegule are yellow, broadly
black in the middle.
Genera and Species of Hymenoptera. 537
This does not appear to me to be LZ. Humbertianus, a
Ceylon species recorded by Bingham from Burma and Tenas-
serim. ‘The metanotal furrow is certainly not ‘supra bi-
spinosa,’ but the apical teeth are distinct enough and luteous
in colour. The scutellum in our species can hardly be said to
be divided in the middle by a furrow, except, perhaps, be-
tween the yellow marks, which have the appearance of being
slightly raised.
Bingham (‘ Fauna of India,’ Hym. i. p. 348), it may be as
well to point out, omits from his generic and specific descrip-
tions one of the most characteristic features of this genus,
namely, that the postscutellum is armed with a stout erect
spine. It is possible that the Burmese examples described
by Bingham as Humbertianus may be different from Saussure’s
species, which was from Ceylon.
B. Descriptions of Two new Genera and Species of Braconidse
from Britain and Two Species of Pompilus from India.
HARKERIA, gen. nov.
Radial nervure abbreviated, not reaching to the apex of the
stigma, the radial cellule therefore confluent with the cubital ;
the cubital nervure almost entirely obliterated, only indicated
by a stump beyond the first transverse cubital, which is rather
faint, but not bullated. The first cubital cellule is confluent
with the first discoidal, through the obliteration of the cubitus
at the base; the transverse median discoidal nervure is not
interstitial with the marginal discoidal, being received shortly
beyond it; the second discoidal cellule is completely enclosed,
it and the costal and the median cellnles are the only complete
cellules. Stigma linear, elongate, narrow; the pterostigma is
distinct. In the hind wings there is only one nervure, which
is probably the subcostal, which ends in what may be a
stigma. Antenne filiform, 19-jointed. Occiput margined.
Mandibles ending in a longish sharp tooth. Eyes prominent.
Parapsidal furrows distinct at the base. Scutellum large,
roundly convex. Median segment large; it has a gradual
rounded slope to the apex, which has a large distinctly defined
area, rounded at the top on either side, and between them in
the middle a narrower one, which is not very distinctly defined
on the top. Abdomen with a distinct petiole, which is about
four times longer than broad; its apex is triangular and ends
in a short projecting ovipositor.
This genus belongs to the Huphorides. It comes nearest
perhaps to Microtonus, with which it agrees in the paucity of
538 Mr. P. Cameron on new
the alar nervures ; but it may be easily known from it and
from all the other genera in having the radial cellule com-
pletely open at the apex, the radius being very short.
Harkeria rufa, sp. n.
Rufa ; vertice mesonotoque nigris; antennis longiore quam corpore,
flagello nigro; alis hyalinis, stigmate pallido, nervis pallide
fuscis. Q.
Long. fere 4 mm.
Antennee blackish, the scape and the base of the third joint
rufo-testaceous. Head, pro- and mesothorax smooth and
shining ; the face and clypeus of a much paler colour than
the vertex; the ocellar region deep black; on either side of
the front ocellus is a fuscons splash which extends to and
behind the eyes. The mesonotum, except in front at the
sides, black ; the scutellum and postscutellum are of a darker
rufous colour; the median segment is obscurely shagreened
at the base. The petiole closely shagreened ; the rest of the
abdomen is smooth and shining.
Taken near Gloucester in June many years ago, probably
on the banks of the Severn, when in the company of the late
Prof. Allen Harker.
LAMADATHA, gen. nov.
Wings abbreviated and narrowed; the stigma and meta-
carpus greatly thickened, broad, projecting beyond the costa ;
the stigma not separated from the metacarpus; the radius
irregular, thickened, about half the thickness of the stigma ;
it is united to the stigma shortly behind the middle by a short
branch, near which there is on it a small cellule, which is
longer than broad; the transverse preebrachial and the pre-
brachial nervures are normal; in the hind wings the costa is
thick ; the subcostal area is distinctly defined; both wings
are ciliated with long hairs. Antenne filiform, longer than
the body, 24-jointed. Head large, broader than long, broadly
projecting behind the eyes ; the occiput not margined, rounded
inwardly. Middle lobe of the mesonotum distinctly separated ;
the scutellum roundly raised; the postscutellum convex,
distinctly separated from it. Metathorax rugosely punctured,
it has a rounded slope from the base to the apex. Petiole
distinct, thick, longer than broad, rugose, the other segments
flat, smooth, and shining, its apex bluntly rounded. Eyes
bare, distant from the mandibles.
The face is rngose, the clypeus roundly convex, smooth,
Genera and Species of Hymenoptera. 539
shining, and is clearly separated from the face by a suture ;
the suture on the mesosternum is wide, deep, and transversely
striated ; the prosternum is long and furrowed down the
centre. The mandibles are small; the apical tooth is long,
curved, sickle-shaped; in the middle is a small triangular
tooth : above them, on either side of the clypeus, is a large
deep depression.
Belongs by the form of the mouth and mandibles to the
Exodontes, and probably to the Dacnuside, from the fact of
there being apparently only two cubital cellules, counting the
small cellule at the base of the radius as the second cubital.
The thickened radius also allies it to Dacnusa. The form of
the mandibles is different from what we find either in the
Dacnuside or the Alysioide, they being much more slender
and sharper, particularly at the apex. The abdominal petiole
is also more slender and distinctly separated.
Lamadatha testaceipes, sp. n.
Nigra ; basi antennarum pedibusque testaceis ; alis hyalinis.
Long.2mm. 4.
Antenne longer than the body, filiform; the basal half
clear, the apical dark testaceous. Head smooth and shining,
sparsely covered with longish hair; the eyes strongly facetted.
Pro- and mesonotum smooth and shining, sparsely covered
with longish black hairs: in the middle of the hinder half of
the mesonotum is a broad, punctured, longitudinal furrow ;
the sides of the raised basal lobe are punctured ; in its centre
is a narrow, smooth, longitudinal furrow. Scutellum and
postscutellum smooth; the depression at the base of the
former is smooth. ‘The lower part of the propleure is closely
rugosely punctured; the part of the mesopleure above the
furrow is smooth and shining ; the metapleure and the meta- |
notum closely rugosely punctured. Wings hyaline, the basal
nervures testaceous; the apical nervures, the stigma, and meta-
carpus blackish, the nervures in the hinder wings testaceous.
Legs testaceous, longish. Abdomen smooth and shining; the
petiole closely rugosely punctured; behind the middle, on
either side, is a small projecting tubercle.
Found early in June in Cadder Wilderness, near Glasgow,
in a fungus much frequented by dipterous larve.
Pompilus perturbans, sp. n.
Niger, pruinosus; alis fusco-violaceis, cellula cubitali 2* duplo
longiore quam 3%, 2°,
Long. 13 mm.
Hab. Barrackpore (Rothney).
540 On new Genera and Species of Hymenoptera.
Head densely covered with a grey pile, the front and vertex
more sparsely with longish blackish hair. Eyes parallel,
slightly converging on the top; the ocelli are in a curve, and
the hinder are separated from each other by a greater distance
than they are from the eyes. Apex of clypeus broadly
rounded. Mandibles broadly rufous in the middle, their base
thickly covered with depressed pubescence. Pronotum in the
middle not quite so long as the head; its apex is broadly
rounded. The median seement has a gradually rounded
slope; looked at from above the apex is transverse ; the sides
at the apex bituberculate, the upper tubercle being larger
and more rounded, the lower smaller, more distinctly defined,
more projecting, and more triangular in form. Wings fusco-
violaceous; the radial and second and third cubital cellules
are lighter in tint, but not hyaline; the second cubital cellule
is about three times longer than the third; the first and
second transverse cubital nervures are parallel, the first has a
more rounded slope, the third is obliquely bent near the top ;
the first recurrent nervure has an oblique slope and is re-
ceived near the transverse cubital nervure as in Salzus, the
second at the apex of the basal fourth of the cellule; the anal
nervure in the hind wings is interstitial. Legs densely
pruinose ; the tibial and tarsal spines are long and stout ;
the underside of the tarsi is covered densely with short
spines ; the claws have one long, curved, and a short, thick,
bluntly pointed tooth. Abdomen black; the apical halves of
the segments with lighter-coloured bands of a dark bluish
tinge,
Comes near to P. parenthope, Cam.: that is a larger and
stouter insect, it has the second cubital cellule longer com-
pared with the third, the second recurrent nervure is received
in the middle of the cellule, and the wings are more uniformly
coloured and have a deeper more violaceous tint ; the head is
much more largely developed behind the eyes, it being there
nearly as long as the width of the eyes, while in the present
species they are not half the length, it being also there more
oblique, less rounded.
P. simillimus appears to be a close ally of this species.
Smith’s description is not detailed enough for satisfactory
determination ; Bingham (Hymen. of India, p. 166) says that
the second and third cubital cellules are equal in length, but
this statement is contradicted in the synoptical table on p. 150,
where the second it said to be wider than the third.
Pompilus implacabilis, sp. n.
Niger, dense argenteo-pruinosus ; alis brevis, flavis, apice fumatis. 2.
Long. 10-11 mm.
Hab. Bengal (Rothney).
Dr. C, A. Kofoid on Platydorina. 541
’ Antenne brownish towards the apex, stout, and placed
close to the top of the clypeus. ead rather long and flat
and wider than the thorax ; the eyes are almost parallel; the
ocelli in a curve (*. *) and separated from the eyes by about
the same distance they are from each other. Clypeus short,
its apex transverse. Mandibles rufous, black at the apex ;
the base thickly covered with depressed silvery pubescence.
Pronotum large, but not quite so long as the head; it is
depressed at the base, transverse at the apex. ‘The entire
thorax is thickly covered with depressed silvery pubescence ;
its apex has an oblique slope. Wings short, not reaching to
the apex of the second abdominal segment, yellowish hyaline,
the apex near the middle of the radial cellule infuscated ; the
radius is roundly curved from base to apex; the second and
third cubital cellules are equal in length above and beneath ;
the first transverse cubital nervure has an oblique slope
on the upper half; the first recurrent nervure is received
shortly behind the second, beyond the middle of the cellule;
the transverse median nervure is received at a distance—by
about its own length—beyond the transverse basal; the anal
nervure in the hind wing is appendiculated. Legs densely
pruinose, the tibiae bare, the tarsi bearing thick spines, the
claws bifid. Abdomen with the segments bordered with
silvery pubescence ; it is about as long as the head and thorax
united; the second ventral segment has a transverse furrow.
The presence of a transverse furrow on the second (or third
according to one method of counting, the first segment forming
part of the median segment) brings this species into Bingham’s
section “ B, Ferreola Group.” It has certainly no near
relationship beyond the abdominal furrow to such species as
P. Cameronii, Bingh. (fenestrata, Bingh., nec Smith), The
present species has some resemblance to the Pompilus vivax
group, but is easily known from them by the abdominal
furrow, by the transverse basal nervure being widely remote
from the transverse median, and by the short wings.
LXXI.—Plankton Studies.—III. On Platydorina, a new
Genus of the Family Volvocide from the Plankton of the
Illinois River. By C. A. Koroip, Ph.D.*
(Plate VIL]
THE family Volvocide is well represented in the plankton of
freshwater ponds and streams. Indeed, with the possible
* From the ‘ Bulletin of the Illinois State Laboratory of Natural His-
tory,’ vol. v. pp. 419-440. From a separate impression communicated by
the Author,
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 36
542 Dr. C. A. Kofoid on Platydorina.
exception of Stephanosphera, all of tle colonial forms in-
cluded in the subfamily Volvocinee—Spondylomorum, Gonium,
Stephanosphera, Pandorina, Pleodorina, and Volvox—are
pelagic in habit and are found only in the freshwater
environment. For the past four years, during the summer
and autumn months, a colonial form belonging to this sub-
family has occurred in plankton collections from the [linois
River and its adjacent waters, to which I have given the
name of Platydorina caudata. It appears as early as June 15,
and becomes abundant in the months of August and September,
diminishing in numbers in October, and disappearing in
November. It thus attains its greatest development toward
the close of the maximum period of summer heat, when the
temperature of the water in which it is found often reaches
36° C. This species has occurred in all the waters examined
in the course of the operations of the Illinois Biological
Station, viz. in the Illinois River, in Thompson’s, Quiver,
Flag, Matanzas, and Phelps Lakes, at Havana, Ill., and in
the Illinois River and Meredosia Lake at Meredosia, III.
During the summer and fall of 1899 it was also found in
abundance near Urbana, III., in Salt Fork, a small stream
tributary to the Vermilion River—a confluent of the Wabash.
It was not equally plentiful in all these localities, but showed
a decided preference for shallow water free from vegetation,
reaching its maximum development when the turbid water
was but a few feet, or even less than a foot, deep. In such
situations the shallowness of the water and the absence of
vegetation conduce to a maintenance of the high temperatures
which seem to favour its multiplication. The bottom of the
lakes in question is usually composed of soft mud, rich in
decaying organic matter, and often covered by a mat of
Oscillaria, but otherwise quite free from vegetation. At
Havana we have found Platydorina in greatest numbers in
Phelps Lake, which in 1896, 1897, and 1898 afforded the
conditions above described. It was likewise abundant in
Thompson’s Lake in the late summer and early fall of 1897
and 1898, when the lake was at a low level and contained
little vegetation. In the shallow open waters of Matanzas
Lake it was much more abundant than in Quiver Lake, where
there was usually a large amount of vegetation. At the
time of the maximum abundance of Platydorina in Salt Fork
in September the stream was reduced by drought to a series
of stagnant pools with no vegetation. In the early part of
August it was full of alge and other aquatic vegetation, and
Platydorina was then present in considerable numbers,
although not so abundant as it was in the following month,
Dr. C. A. Kofoid on Platydorina. 543
On August 2, 1888, Professor H. Garman, while con-
ducting a biological survey of the aquatic life in the vicinity
of Quincy, Ill., in the bottoms of the Mississippi River (see
Garman, 1890), found a specimen of this interesting species
in the waters of Libby Lake. He records and sketches it in
notes now on file at this Laboratory, but published nothing
concerning it,
The occurrence of this new genus in the waters of the
Wabash, Illinois, and Mississippi River systems, and _ its
recurrence in our collections for several successive years
indicate its wide distribution and firm establishment in the
Mississippi Valley in waters of some permanency. It has
not yet been noted in temporary pools.
The associates of Platydorina in the plankton have varied
with the season, the locality, and the year. It may be said,
in a general way, that the plankton in which it occurs is
characterized by an abundance of flagellates, of rotifers—
especially Brachionide—and of immature Copepoda. <A
water-bloom, composed largely of Huglena, Trachelomonas,
Carteria, and other green flagellates, often appears at the
suiface of waters where Platydvrina is abundant. Gontum
is frequently associated with it in large numbers, as are also
Pandorina, Eudorina, and Pleodorina, though these three
genera may also be plentiful in the early summer, when
Platydorina may be absent or rare. Pleodorina californica
was extremely abundant in Salt Fork in August, but had
almost entirely disappeared by the time that Platydorina had
reached its maximum. A few specimens of Volvox, which,
in this locality, is common in the spring months, were also
noted, while perhaps the most interesting associate in Salt
Fork was Ceratium kumaonense, discovered by Carter (1871)
in Hindostan. Other chlorophyll-bearing associates tre-
quently seen are Pediastrum, Scenedesmus, Actinastrum, and
Olosterium. Among the diatoms, Melosira, F'ragillaria, and
Surirella were to be seen; and among the Peridinide Peri-
dinium tabulatum was almost always represented.
The zodplankton associated with Platydorina is not less
varied than the phytoplankton. The Protozoa were usually
represented by Arcella, Difflugia, and occasionally by pelagic
Ameba; by Synura, Mallomonas, Dinobryon, and Uroglena ;
and by Codonella and Coleps. Among the Rotifera the order
Ploima was well represented; Z?iarthra and the Brachionidee
—notably Brachionus militaris, 2, angularis, B. punctatus,
and B, Bakeri and its varieties—were most common during
the summer months, while the Syncheetide increase in numbers
in the early autumn. Tolyarthra was frequently abundant,
30*
544 Dr. C. A. Kofoid on Platydorina.
and Rotifer, Philodina, Asplanchna, Euchlanis, Cathypna,
Distyla, Monostyla, and Pterodina were often represented by
one or more species. The paucity of Entomostraca stands in
sharp contrast with the abundance of rotifers, the former
group being represented by relatively few species and few
adult individuals. The nauplii of Cyclops were, however, as
a rule, abundant, but only occasional specimens of adult
Cyclops, Diaptomus, Bosmina, Chydorus, Certodaphnia,
Daphnia, and Cypridopsis were to be seen.
Platydorina caudata, gen. et sp. n.
The species here described consists of a horseshoe-shaped
ceenobium or colony (Pl. VII. fig. 1) of 16 or 32 biflagellate
cells, the anterior end corresponding to the toe of the horse-
shoe and the truncate posterior end to the heel, the latter
carrying 3 or 5 prolongations or tails formed by the gelatinous
substance of the coenobium. The colony is plate-like and
flat, except that the plate is slightly twisted in a left spiral.
This spiral is scarcely noticeable in a face view (fig. 1) except
by focussing with a high-power objective, but it can be easily
detected in a side view (fig. 3). It varies from one eighth to
one thirty-second of a turn of the spiral, and in twenty-five
colonies especially examined on this point it was invariably a
left spiral, with the location of the twisting always in a defi-
nite relation to the colony, the right anterior and the left
posterior regions of the colony in face view being high, while
the left anterior and the right posterior are low. Repeated
examinations of specimens, both living and preserved, indi-
cate that this spiral form is a constant feature of structure,
that it is not reversed in direction, and that it is subject to
but slight variation in the degree of the torsion. No move-
ment within the colony which would produce or vary the spiral
was noted in living individuals. ‘he form of Platydorina
seems to be as constant and as characteristic as that of other
genera of the family.
The size of the colony varies with the age, with the number
of cells present, and also perhaps with the locality and the
season. Colonies of 32 cells in which the first division
leading to the formation of daughter colonies is taking place
average about 150 w in length, 130 w in width, and 20 w in
thickness. ‘he largest colonies are about 165 x 145 x 25 p,
and the smallest free-swimming ones about 25x21 x4 b
Colonies of 16 cells are smaller than those of 32 cells and
are also narrower in proportion to their length, measuring
about 70 x 43 x 16 w.
Dr. C, A. Kofoid on Platydorina. 545
The colours of Platydorina are quite as striking as those of
related forms. The cells, which are imbedded in the trans-
parent gelatinous matrix of the colony, are a bright chloro-
phyll-green, and each has, as a rule, a red stigma or eye-spot
of unusual brilliancy.
The substance in which the cells of the colony are imbedded
is similar in appearance to that in Hudorina. It is a trans-
parent colourless substance of considerable consistency, showing
in the living condition, as a rule, no trace of differentiation.
The gelatinous nature of the substance is shown by the great
numbers of bacteria which swarm within it in moribund
specimens. Colonies killed in formalin and stained in Dela-
field’s hematoxylin exhibit a difference in the intensity of
coloration, indicating the presence of a denser peripheral
layer or sheath 3-4 w in thickness (Pl. VII. figs. 1, 4, p.sh.).
This is apparent along the edges of the colony and presumably
extends over its faces. In several living colonies a granular
differentiation of this outer layer was noted about the margin.
This sheath is similar to that of Hudorina and Pleodorina,
but shows no trace of the concentric layers so prominent in
Pandorina.
One of the most characteristic features of the colony is the
presence upon the posterior border of 3 or 5 projections or
tails, which are merely extensions of the sheath. Colonies of
16 cells have but three tails, while those of 32 cells have
uniformly five. ‘These projections are bluntish finger-like
processes without structural differentiation, tapering some-
what to a rounded or pointed end. Occasionally the outer-
most pair, and more rarely the inner one, are slightly divergent.
In the 16-cell colony there are two latero-posterior tails and
one median one (fig. 2), the former being better developed
and measuring 15 to 20 w in length. The median tail is
variable in length, being sometimes a mere rudiment appearing
on a slight elevation on the margin. Its average length is
about one third that of the adjacent pair, though it occasionally
attains two thirds their length. The latero-posterior tails are
upon each side of the colony directly behind the marginal
row of cells, while the median tail is midway between the
central rows. In the 32-cell colony (fig. 1) there is, in
addition to the three tails above noted, another pair which
may be designated as the lateral pair. ‘hese tails are slightly
divergent, arising at the outer posterior angles of the marginal
row of cells, between the last transverse quartet and the last
sextet of cells. They are from 10 to 15 w in length and are
often of the same size as the median tail of the colony. The
other three tails occupy the same position with respect to the
546 Dr. C. A. Kofoid on Platydorina.
posterior quartet of cells that they do in the 16-cell colony,
but are, as a rule, much larger, the postero-lateral pair
measuring from 20 to 30 mw in length, while the median one
reaches only a length of 15 to 18 w. The five tails do not
lie in one plane, but share in the spiral of the colony, at
times, indeed, exceeding it in the degree of the twisting.
These structures are all subject to considerable variations and
nregularities of development (fig. 5), such as suppression,
inequality of members of pairs, differences in size and relative
development, in attenuation, and in degree of divergence.
These irregularities are often correlated with the loss of cells
in the colony due to parasites or other causes. The tails
nevertheless exhibit such a constancy of position and so much
of symmetry and regularity of development that they cannot
for a moment be considered as ephemeral features of little
structural importance. In their position they recall the pro-
tuberances noted by me (1898) at the posterior end of Pleo-
dorina illinoisensis. In Pleodorina, however, these struc-
tures are apparent only in disintegrating maternal colonies,
and it may be that they also indicate the point at which the
embryonic cup closes, On the other hand, in Platydorina
caudata these tails are present upon the colonies at the time
of their escape from the maternal matrix, and persist through-
out the life of the adult, being permanent structures, charac-
teristic of the species.
Within the outer sheath is a homogeneous gelatinous matrix
(fig. 1, m), which in Delafield’s hematoxylin stains less
readily than the sheath. In the living colony no differentia-
tion of this matrix is to be seen, but, after staining, a delicate
sheath showing deeper colour is demonstrated about each of
the cells. In most places a considerable space intervenes
between this secondary sheath (fig. 4, s.sh.) and the inclosed
cell, so that the sheaths crowd upon each other and appear
to divide the field of the matrix into irregular polygonal areas.
These areas, as a rule, fill the greater part of the plate, leaving
unoccupied only a few corners, principally about the second
transverse row. The two poles of this swollen secondary
envelope are not of equal size, the inner being somewhat the
smaller, and slightly overlapped by those of the contiguous
cells. This is due to the intercalation of the cells of the two
sides of the plate and to the fact that the outer ends of the
cells are slightly nearer the surface of the plate than are the
inner ones. ‘The gelatinous substance within the secondary
sheaths does not differ in structure or stainibility from that
of the surrounding matrix. As a result of the form of the
Dr. C. A. Kofoid on Platydorina, 547
colony the amount of the matrix substance is much less in
Platydorina than in related forms such as Hudorina.
The cells of the colony are all of one type, alike in structure,
and approximately similar in size. Each is biflagellate and
has a central body of protoplasm with a nucleus, two contrac-
tile vacuoles, one stigma, and one chromatophore with a single
pyrenoid.
The number of cells in the colony is either 16 or 32; at
least no normal colony with cells of any other number has
been detected among the hundreds, if not thousands, of
colonies examined. Colonies are frequently seen which, by
reason of parasites or from other causes, have lost one or more
cells—indeed, in some cases all but one or two; but the form
of these colonies is usually preserved, and the empty secondary
sheaths frequently remain as evidence of the original comple-
ment of cells. ‘lhe 16-cell colonies are not mere stages in
the development of the 32-cell form, for division of the cells
of this type in observed cases leads to the development of new
colonies and not to the formation of the 32-cell stage. As in
other nearly related genera of the family—for example, Hudo-
rina, Pandorina, and Pleodorina—the number of cells in the
colony varies, within narrow limits, in the ratio of geometrical
progression. In Platydorina, however, this pleomorphism is
manifested not only by this difference in the number of cells
in the colony, but also by a structural distinction—the presence
of three tails in the 16-cell and five tails in the 32-cell colony.
Inasmuch as the two types always occur together, and since
this pleomorphism is in some respects similar to that of
related genera, it does not seem justifiable to regard the two
as distinct species of the genus. ‘They are, I believe, two
forms of one species.
The arrangement of the cells is characteristic and is
strikingly different from that of any other genus of the family.
The gelatinous matrix and sheath conform to the horseshoe-
shaped plate of cells, and even the caudal appendages bear a
fixed relation to the plan of cell-arrangement. The 32-cell
colony is composed of a marginal U-shaped row of 12 cells
about three sides of a 20-celled somewhat rectangular plate,
which, in turn, consists of an outer row of 12 cells on three
sides of a row of four pairs of cells. The colony might also
be regarded as made up of three U-shaped rows of 12, 12, and
8 cells respectively, nested in such a fashion that the inner
two project one cell beyond the outermost. ‘The cells also
fall into six quite irregular transverse rows of 4, 6, 6, 6, 6,
and 4 cells respectively, and into the same number of corre-
548 Dr. C. A. Kofoid on Platydorina.
sponding longitudinal ones. As before stated, the lateral
tails are posterior to the marginal row, while the postero-
laterals are behind the first row within the marginal, and the
median one midway between the innermost rows. In the
colony of 16 cells (fig. 2) the marginal row has but 10 cells
and the central plate but 6. The cells fall into four some-
what irregular transverse rows, and there are the same
number of longitudinal ones of 4 cells each. The horseshoe-
shape, however, masks somewhat this simple Gontum-like
arrangement. The plate-like form of the colony and the
arrangement of the cells, especially in the 16-cell form, give
this new genus a superficial resemblance to Gontum. It is,
however, fundamentally different, for in Platydorina the two
faces of the plate are exactly alike, while in Gontum the face
anterior in locomotion bears all the flagella, and the other
face presents only the bases of the cells. This similarity of
the two faces in Platydorina, neither of which is anterior or
posterior, is brought about by the fact that every other cell
upon either face presents to that face the pole which bears
the stigma and the flagella, while the intervening cells present
the opposite pole, with its pyrenoid. This alternation of stigma
and pyrenoid is constant, and can be followed in any row of
cells except the diagonal ones (cf. fig. L). The cells of the
marginal row, in both the 16- and 32-cell colonies, point
obliquely outward, the direction alternating, however, in
conformity with the arrangement of cells in the central area,
as may be seen in a view of the edge of the colony (fig. 3).
The alternation of the cells in the colony as a whole is the
same whichever face is presented, the right-hand cell of the
posterior row of four cells always presenting the basal end
uppermost. Fyrom this as a starting-point the regular alter-
nation of stigma and pyrenoid can be traced from cell to cell
throughout the whole colony. An examination of twenty-five
colonies showed that all conformed to the same plan of alter-
nation, there being no case of reversal. In the arrangement
of the cells in the colony Platydorina is thus more like
Eudorina than like Gonium, being not a simple plate like the
latter genus, but, in reality, a flattened ellipsoid so much
compressed that the cells of the two faces intercalate
regularly, and thus give to the colony its superficial resem-
blance to Gontum.
The individual cells are all substantially alike in size and
structure. ‘They have the form of an oblate spheroid, slightly
larger in the outer hemisphere*. Some cells, especially the
* As in the case of Eudorina and Pleodorina the terms “ outer” and
Dr. C. A. Kofoid on Platydorina. 549
marginal ones, often exhibit a slight flattening or even a
depression at the outer pole. In the full-grown colony the
cells have an equatorial diameter of 15-20 w and a polar one
of 15-18 p. The cells of young colonies still within the
maternal matrix do not exceed 4-6 w in diameter. I do not
find that the cells of the 16-cell colonies are appreciably
larger than those of the 32-cell.
The protoplasm is small in amount, consisting of a very
thin pellicle (fig. 4, p.) on the surface of the cell on the out-
side of the chromatophore, and an axially-placed knob-
shaped mass (pr.) located somewhat nearer the outer pole
than the inner one. Near the centre of this mass lies the
spherical nucleus (fig. 4, n.), containing a single spherical
nucleolus (ncl.). Within this protoplasmic mass lie the two
contractile vacuoles (c.v.) and the stigma (s¢.), while from
the outer end of the cell arise the two flagella (/.).
There is but a single cup-shaped chromatophore (fig. 4,
chr.), which is inclosed within the pellicle above noted, and
itself contains the knob-shaped protoplasmic mass. It is of a
brilliant chlorophyll-green colour, and contains numerous
small granules of irregular and somewhat angular outline.
Towards the inner end of the cell, imbedded in the thickest
part of the chromatophore, there is a single spherical py-
renoid, having a diameter of 4-6 uw.
The stigma or eye-spot, seen from above, is circular in
outline, but in lateral view has the form shown in fig. 4, s.
The slightly convex outer surface appears to project some-
what beyond the rounded contour of the cell. The colour is
usually a bright reddish brown, often brightest in the anterior
and marginal cells and rarely entirely faded in the posterior
ones. ‘lhe stigma is a homogeneous body, showing no trace
of structure beyond the well-defined contour-line, which is
best seen in fading and moribund cells. It is normally
present in all cells of the colony, and may readily be demon-
strated by full illumination. The position of the stigmata in
the cells is somewhat unusual, and is significant of the pro-
nounced polarity of the organism. The customary position
in other genera is adjacent to the bases of the flagella. In
Platydorina, however, the location of the stigma is not con-
stant with respect to the flagella, but seems rather to bear a
definite relation to the form of the colony, since it lies towards
“inner” are used to designate respectively the ends which bear the
stigma and the pyrenoid,
550 Dr. C. A. Kofoid on Platydorina.
the peripheral and posterior region of the cell (fig. 1), while
the flagella are centrally located and project outward in the
usual manner. ‘This relation appears not only in the mar-
ginal regions, but also in the central. The physiological
significance of this arrangement is not apparent, but it seems
to be correlated with the pronounced polarity of the organism.
Platydorina is positively phototactic. A miscellaneous plank-
ton collection placed in a window with southern exposure in
an aquarium 6 inches in diameter was, after ten minutes,
quite barren of Platydorina except along the margin towards
the window. On the other hand, this species avoids bright
light. This was very evident in collections fresh from the
ficld when examined under low power (75 diameters), a very
slight increase above a moderate illumination causing them
to leave the field of view with considerable rapidity. In one
case, where twenty-eight colonies were in the field when
placed under the microscope, only one of them remained after
an exposure of twenty-five seconds. <A very slight decrease
in the amount of light would invariably insure their return to
the field with almost equal rapidity, the number increasing
as the intensity of the illumination was decreased. It may
be that the asymmetrical position and the somewhat unusual
arrangement of the stigmata are connected with the pro-
nounced phototaxis of this organism. At least the asym-
metrical position has a tendency to place the long axis of the
stigma parallel to the main axis of the colony, with the outer
end directed towards the source of light in negative, and away
from it in positive, phototaxis.
The flagella are uniformly two in number for each cell, are
similar in the same cell and in different parts of the colony,
and are in the adult colony 20-25 win length. From the
outer pole of the cell they pass through the matrix, leaving
the appearance of a tube-like structure in the gelatinous sub-
stance (fig. 4). When not in activity the flagella project
beyond the sheath in a position perpendicular to the surface
of the colony at the place of exit. As in other genera of this
family, the flagella persist after the division of the cell to
form the daughter colony, and even after the divisions are
completed still provide locomotion for the maternal organism.
In some instances the flagella could be seen passing through
the matrix toward that cell of the daughter colony which
bears the largest eye-spot.
The contractile vacuoles (fig. 4, c.v.) are two in number,
and are located in the peripheral layer of protoplasm near
the outer end of the cell. ‘They lie in the outer part of the
knob-shaped mass of protoplasm upon either side of the place
Dr. C, A. Kofoid on Platydorina. 551
of origin of the flagella, being somewhat widely separated.
At diastole the vacuoles of an adult colony have a diameter of
15-2 w. The contraction is rhythmical and the two vacuoles
usually alternate at regular and equal intervals. At a tempe-
rature in the laboratory of 20° C. each vacuole contracted at
intervals of forty-tive to fifty seconds. In rare instances the
contractions of the two vacuoles were separated by unequal
intervals, being almost coincident in one case observed.
The method of locomotion in Platydorina is similar in
many respects to that of other genera of the family. The
lashing of the flagella produces a forward movement of the
colony, and causes its rotation about the major axis either
from left over to right or from right over to left. ‘The rounded
end of the colony is uniformly directed forward in locomotion ;
at least, no instance in which the caudal end led was noticed.
The forward movement is, as a rule, accompanied by the
rotation of the colony, though the amount of rotation varies
somewhat with the individual, the freedom of movement, and
the speed of locomotion. When locomotion is blocked by
obstructions the rotation continues, as in Pleodorina, with
frequent reversals in direction. In fact, obstruction to pro-
gress seems frequently, though not uniformly, to act as a
stimulus to the reversal of the direction of rotation.
The two directions of rotation are not equally prevalent,
that from right over to left having a marked predominance.
Thus, of twenty-five colonies observed in motion twenty were
rotating from right over to left and but five from left over to
right. In another twenty-five the corresponding numbers
were nineteen and six respectively. Keeping a single colony
under observation for some time, it is found to rotate from
right over to left about four fifths of the period and to turn
in the opposite direction the balance of the time, this pro-
portion representing the totals of the periods of rotation,
while the individual periods vary greatly in length, that from
left over to right lasting at times but a few seconds.
This predominance of one direction in locomotion is doubt-
less correlated with the torsion of the colony, whose shape is
such that the rotation would necessarily be from right over
to left in forward locomotion, as a result of the resistance of
the water, unless, of course, there should be some disturbing
factor. ‘lhe immediate and most potent cause of the direc-
tion of rotation is doubtless the coordinated action of the
flagella, since reversal of direction does not seem to be
accompanied by any change in the direction of the torsion of
the colony. ‘lhe evidence upon this point is not conclusive,
but repeated efforts have failed to detect any change in the
552 Dr. C. A. Kofoid on Platydorina.
form of the plate when the direction of rotation is reversed
in the living and moving colony ; and, again, colonies when
killed suddenly have always the usual form of spiral, though
some of them were moving in the reverse direction. When
the usual direction of rotation is reversed, the forward motion
still continues in spite of the fact that then the form of the
plate favours a backward movement; the form of the colony,
therefore, does not control the direction of rotation, though it
is correlated with the direction which predominates. The
fact that the rotation from right over to left predominates
also in Pleodorina illinoisensis and Eudorina elegans, where
there are no structural features favouring such a predomi-
nance, suggests the possibility that the form of the colony in
Platydorina is the result and not the cause of this predomi-
nance, and that the function of turning from right over to
left predominantly preceded the structure which favours it.
The organization of Platydorina suggests a descent from a
Eudorina-like form, in which event the’systematic series and
the phylogenetic series alike afford evidence of a function
arising in an organism before the structure with which it is
correlated appears.
In another connexion (1898) the subject of locomotion and
polarity in the different genera of the Volvocine was reviewed
and discussed. It will suffice, therefore, for the present to
give a brief résumé of the facts. In the lower genera of the
family, Stephanosphera and Gonium, as also in Pandorina,
the rotation seems to be indifferently to the right or left, while
in Hudorina, and especially in Pleodorina illinoisensis, it is
oftenest to the left, rotations to the right in observed cases of
the latter species being to those to the left as 100 to 117-138.
With respect to Volvow, there are no data at hand. In Platy-
dorina we find by far the most pronounced predominance of
one direction of rotation, the ratio in observed cases being
100 rotations from left over to right to 355 from right over
to left. In this respect, then, so far as there is evidence,
Platydorina is the most highly differentiated genus of the
family.
The polarity of the lower genera, Stephanosphera and
Gonium, is likewise of the simplest form, being merely
physiological, the same pole or face of the colony always
leading in locomotion. In Pandorina, Eudorina, and Volvox,
however, there is the added feature of the greater brightness
of the anterior stigmata, and in Pleodorina the two poles are
differentiated by the two types of cells, as well as by the
characters found in the genera just mentioned; but Platy-
dorina is the only genus of the family in which polarity is
Dr. C. A. Kofoid on Platydorina. 553
expressed by the arrangement of the cells and by structural
features of the envelope. In regard to polarity, also, the
new genus is thus the most highly specialized member of the
Volvocine.
The reproduction of Platydorina has been observed by me
repeatedly in the past five years, but only the asexual phase
has thus far been discovered. All of the cells of the organism
are gonidial, each dividing to form a daughter colony. The
sequence of the divisions and the position of the successive
planes are of the type found in Hudorina and Pleodorina,
the resemblance being so close that the figures illustrating _
the asexual development of Pleodorina illinoisensis (1898,
pl. xxxvil.) might almost be used for cleavage in Platydorina.
‘There is one difference, however, for in Platydorina the
curved plate of cells, which becomes first cup-shaped and
then ellipsoidal, subsequently flattens, the cells of the two
faces intercalating during the process. The daughter colony
acquires the adult form, including the tails and the torsion of
the plate, before it escapes from the maternal matrix, the
young colonies moving about for some time in the disinte-
grating matrix before making their escape through the
ruptured outer sheath. The secoudary sheath surrounding
the gonidial cell becomes the outer or primary sheath of the
new colony. No stages of sexual reproduction have been
seen, though the collections examined represent a considerable
range of season and locality. It may be that these are to be
sought upon the bottom rather than in the superjacent strata
of water where plankton collections are usually made.
Aquaria about to dry up were also searched in vain for sexual
stages of Platydorina.
The mode of development of Platydorina is significant of
its systematic position and its relationships. ‘The number
and the original arrangement of the cells, the type of deve-
lopment, and the character of the envelope, all indicate that
Platydorina is a more highly specialized form descended from
some Hudorina-like ancestor, and that it is more closely
allied to Audorina than to any other existing genus.
‘Throughout this paper the customary term ‘colony ”’ has
been used to designate the organism herein described and
others related toit. ‘The wide use of the term in the literature
of the subject is doubtless due to the fact that, as a rule, the
organisms are composed of similar cells arranged in symme-
trical form with no pronounced axial differentiation, without
contact or protoplasmic connexion, separated from each other
by a non-living gelatinous matrix, and each capable of per-
forming all the functions necessary for its own life and the
5d4 Dr. C. A. Kofoid on Platydorina.
continuance of the species. T'urthermore, the destruction of
individual cells does not impair the life of the other cells of
the organism, for so long as a single cell remains it continues
its customary activity. “he useof the term colony is, however,
objectionable. A number of facts militate against this con-
ception of the organism, and the discovery of the new genus
here described adds to the array. (1) The cells are not
always similar, for in all forms with poles physiologically
or otherwise differentiated the anterior stigmata are brighter
than the posterior, and in Pleodorina there are two kinds of
cells, the vegetative and the gonidial, the former distinctly
smaller than the latter. (2) There is in all of the higher
genera a well-defined physiological polarity accompanied by
the difference in the anterior and posterior stigmata, and
also, in Platydorina, by a differentiation of the poles by the
arrangement of the cells and the structure of the envelope,
and by the further differentiation of a transverse axis.
(3) In Landorina the cells are almost in contact, and in
Volvox they are actually connected by protoplasmic processes.
(4) The beginnings of histological differentiation are also
evident in the cells composing the so-called colony. In
Eudorina, according to Carter (1858), the cells are differen-
tiated into male and female in definite regions, the male cells
developing from the anterior quartet and the remainder
becoming female ; in Volvoa sexual and asexual reproduction
alike are limited to a few of the cells; and in Pleodorina the
asexual process is confined to the posterior hemisphere. The
cells of the organism are thus histologically and functionally
differentiated in this particular in these higher genera.
Although the degree of differentiation is slight, it is never-
theless appreciable. (5) In the matter of locomotion the
activities of the individual cells of the organism are not
independent of each other, but are correlated, the flagella
acting together to produce rotation, its reversal, or its cessa-
tion. The piedominance of the direction in the higher genera
plainly exhibits the phenomenon of correlated locomotor
activities of the constituent cells.
The facts above cited emphasize the desirability of regarding
each of these so-called colonial flagellates of the subfamily
Volvocine as a unit, with an organization of its own, and not
as a colony, that is, an aggregation of independent and similar
cells associated merely as a result of descent from a common
parental cell, the form being a matter of chance or circum-
stance. ‘lhe group of cells as a whole, and not each of the
constituent cells, is the unit of descent, of form, and of func-
Dr. C. A. Kofoid on Platydorina. 555
tion, and the word colony can be applied to it only by the
license of usage and as a matter of convenience.
Reference has been made frequently in the preceding pages
to the prevalence of colonies whose symmetry has been dis-
turbed by loss of cells. In most instances only the empty
secondary sheath remains, giving no clue to the cause of the
loss of its contents. In collections made in Phelps Lake,
Havana, IIl., in August 1896, however, colonies were often
found which were parasitized by one of the Sporochytriacez,
which upon examination proves to be Dangeardia mamillata,
described by Schréder (1898) as a parasite of Pandorina
morum. As these infested colonies frequently showed a loss
of one cell or more and exhibited all stages in the destruction
of the cell, it seems probable that the loss was due to the
parasite. Hudorina elegans and Pandorina morum occurred
in the same collection and were similarly infested. Two
additional genera, Platydorina and Eudorina, are thus to be
added to the list of hosts of Dangeardia.
For the convenience of systematists a brief statement of
the generic and specific characters of the form herein described
is now given, followed by a key to the genera and species of
the Volvocine for the assistance of students of this inter-
esting and not uncommon group of freshwater organisms.
Species not as yet reported to my knowledge from Illinois are
indicated by an asterisk when found elsewhere in this con-
tinent and by a dagger when not as yet reported from it. It
is not at all improbable that all the species here listed will
yet be found within this State.
PLATYDORINA, gen. nov.
Colony flattened, the two faces compressed so that the cells
of the two sides intercalate; flagella upon both faces on
alternate cells. Anterior and posterior poles of major axis
differentiated by the arrangement of the cells and by the
structure of the envelope. Long and short transverse axes
differentiated by the flattening of the colony. Cells similar,
biflagellate, each with stigma, chromatophore, and pyre:oid.
Asexual reproduction by repeated divisions of all of the cells,
each forming a daughter colony.
Platydorina caudata, sp. n.
Colony flattened, horseshoe-shaped, twisted about one
eighth of a turn from right over to left; cells 16 or 32,
556 Dr. C, A. Kofoid on Platydorina.
arranged in a marginal row of 10 or 12 and a central area
of 6 or 20; posterior end with 3 or 5 prolongations or tails
formed by extension of the common outer sheath.
Known habitat, lakes and streams in Central Illinois.
Types in the collections of the Illinois State Laboratory of
Natural History and deposited in the United States National
Museum,
Keys to the Genera and Species of the Subfamily Volvocine.
Cells arranged in ccenobia of definite forms varying with the species,
biflagellate, with stigma and one or more chromatophores; surrounded
by a gelatinous envelope, whose development separates them to a greater
or less degree; number not uniform, varying in the different species,
often, but not always, definite. Asexual reproduction by repeated divi-
sions of gonidial cells, which constitute the whole or only a part of the
parental organism, to form daughter organisms ; sexual reproduction in
some species (in others unknown) by the conjugation of male and female
gametes, resulting in the formation of a resting stage, which later develops
into a uew organism.
Genera.
( Cells arranged in form of plate, with flagella upon
J one free only Sie. : MOR een Sele 2.
14 Cells arranged in spherical, ellipsoidal, or flat-
| tened colonies ; flagella not confined to one
te PUCR tsa se apis Sing os Eemtas ett onl te ia eee ree 3.
(Cells in a squarish plate; envelope closely ad-
sy MGrent ls CULT Le ete ae ae oe Ree Gonium,
“\ Cells in a rounded plate ; envelope swollen, oval,
L ormpherteall tag: ete ese ek crab phat oe eee Stephanosphera,
(Colony ellipsoidal or spherical; cells crowded
| together, conical, reaching towards centre,
34 outer membrane of concentric layers ...... Pandorina.
| Cells not crowded together nor reaching towards
contre of colony) yicead4 eee arn apts shapes 4,
( Colonies ellipsoidal or flattened; cells uniform in
| SIZE aa ur roe ste nasa las outer ste SLs ee oP 5.
42 Colonies spherical or spheroidal, or, if ellipsoidal,
| with small vegetative and large gonidial
ie Cells vise% Seven ete re aoe cane on eee 6.
( Colony ellipsoidal or spherical ; poles not differen-
| tiated by arrangement or size of cells or by
5 structure or envelope’ = 21.7. fete te sen ee Eudorina,
\ Colony flattened, horseshoe-shaped, with poles
| differentiated by arrangement of cells; poste-
L nor .6nd ~withstaile>\: vores hae ote bear ete Platydorina.,
( Cells not connected by protoplasmic processes, of
| two sizes, smaller vegetative at anterior pole
: and larger gonidial at posterior............ Pleodorina.
| Cells connected by protoplasmic processes, not
i markedly different in size ........... . is att) Volpe.
6
Dr. C. A. Kofoid on Platydorina. 557
Species.
GONIUM.
MEME Se diver need ei sa hile spare ane sociale (Duj.).t
| Calls AGS ss% Sete ttey as ia snk 83 pectorale, Mill,
STEPHANOSPHERA.
Represented by a single species, characterized as
follows :—Cells 4 or 8, ovoid or spindle-shaped,
with numerous processes ..........02000005 pluvialis, Cohn.*
PANDORINA.
Represented by asingle species, characterized as
follows :—Cells 16 or 32, crowded, each with
a single chromatophore and pyrenoid........ morum, Bory.
EuporINa.
Represented by a single species, characterized as
follows:—Cells 32, 16, or 64, similar, not
crowded together; common outer membrane
without marked concentric structure........ elegans, EKhrb.
PLATYDORINA.
Represented by a single species, characterized as
follows :—Cells 16 or 32, arranged in a horse-
shoe-shaped plate, those of the two faces inter-
calated. Posterior end with 3 or 5tails .... caudata, Kofoid.
PLEODORINA.
{ Cells 64 or 128; gonidial cells about two to three
times the diameter of vegetative cells, which
constitute about one half the total number
4 and lie in anterior hemisphere............ californica, Shaw.
Cells 32, rarely 16 or 64; gonidial cells not more
than twice the diameter of the vegetative
{ cells, which constitute the anterior quartet, . ¢//inoisensts, Kofoid.
VOLVOXx,
(Cells about 10,000 (minimum 1500, maximum
22,000), angular, with stout connecting
protoplasmic processes into which the chro-
matophore may enter. Diameter of colony
about 700 » (minimum 400, maximum
J 1200) ; diameter of cell-body 3-5 ...... globator, Li.
Cells 500-1000 (minimum 200, maximum 4400),
rounded, with slender connecting protoplas-
mic processes into which the chromatophore
does not enter. Diameter of colony 170-
Ke 850 4»; diameter of cell-body 5-8 yw ...... aureus, Ehrb.
University of Illinois,
Urbana, IIl., U.S.A.,
Dec. 5, 1899.
Ann. & May. N. Hist. Ser. 7. Vol. vi. 37
558 Prof. R. Burckhardt on the
Literature cited.
Carter, H. J. (1858.)— On Fecundation in Eudorina elegans and
Cryptoglena,” Ann. & Mag. Nat. Hist. ser. 3, vol. ii. pp. 237-253,
1. Vili.
Pe (1871.)—* Note on a Freshwater Species of Ceratiwm from the
Lake of Nynee (Naini) Tal in Kumaon,” Ann. & Mag. Nat. Hist.
ser. 4, vol. vii. pp. 229, 230.
Garman, H. (1890.)—“ A Preliminary Report on the Animals of the
Mississippi Bottoms near Quincy, Illinois, in August 1888,” Part L.,
Bull. Ill. State Lab. Nat. Hist. vol. iii. pp. 123-184.
Kororp,C. A. (1898.)—“ Plankton Studies, II. On Pleodorina illinois-
ensis, a new Species from the Plankton of the Illinois River,” Bull.
Ill. State Lab. Nat. Hist. vol. v. pp. 273-293, pls. xxxvi., xxxvii.
ScuroveEr, B. (1898.)—“ Dangeardia, ein neues Chytridineen Genus
auf Pandorina morum, Bory,” Ber. d. deutsch. Bot. Gesellsch.
Jahrg. 1898, Bd. xvi. pp. 314-321, Taf. xx.
EXPLANATION OF PLATE VIL*
Abbreviations.
A, Anterior pole. P. Posterior pole.
e.v. Contractile vacuole. p. Outer pellicle of protoplasm.
f. Flagellum. pr. Knob-shaped mass of proto-
m. Matrix, plasm,
n. Nucleus. pyr. Pyrenoid.
nel. Nucleolus. s.sh, Secondary sheath.
p.sk, Outer or primary sheath. st. Stigma.
Fig. 1. Platydorina caudata, face view of 32-cell colony, x 550.
Fig. 2. Face view of 16-cell colony, x 628.
lag. 3. Edge view of 32-cell colony, x 350.
Fig. 4. Lateral view of one of the marginal cells, x 1400.
Fig. 5, a-e. Outline of the posterior ends of several deformed colonies,
x 280.
LXXII.—On the Luminous Organs of Selachian Fishes. By
Rup. BurckuHarpt, Nat. Hist. Mus., 8. Kensington.
In the ‘ Zeitschrift ftir wissenschaftliche Zoologie,’ vol. Ixvi.
1899, Johann, at the instigation and under the guidance of
Prof. Blochmann, of Tiibingen, published a paper entitled
‘““Ueber eigenthiimliche epitheliale Gebilde (Leuchtorgane)
bei Spinax niger,” in which are given the results of some
researches he made in this subject.
In this paper he describes some very minute organs with
* Figures drawn by C. A. Kofoid and inked by Miss L. M. Hart.
Luminous Organs of Selachian Fishes. 55)
which the skin of Sp/naa niger is covered in great numbers,
more particularly so on its black-coloured parts. In addition
to describing their distribution upon the surface of the body
and their histological structure, which in the main points
corresponds with those of Phyllirhoé and the Pennatulide, he
also compares them with other laminous organs.
He moreover refers to a previous observation on phospho-
rescence made in another Spinacide species, Jsistius bra-
siliensis, by Bennett as early as 1840 during a whale voyage,
in which he states expressly that, with the exception of a
ring on the throat remaining dark, the whole of the ventral
surface of this fish was aglow for about three hours. It
struck Johann as being remarkable, when he discovered on
the throat in Sp¢naz two regions extending towards the median
line which were devoid of these structures. From this fact
alone, and quite apart from purely structural characters, he
was right in his surmise, that in all likelihood they would
have to be regarded as organs of luminosity.
The accuracy of his notions as regards their functions was
moreover fully borne out in a note from Th. Beer, dated
Jan. 31, 1899, which reached Johann in time to find a
place at the end of his paper.
In his note Beer says that, whilst engaged in ophthalmo-
scopic observations on Spinax in a room darkened for this
purpose, its phosphorescence was vivid enough to enable
him to see it at a distance of from 3-4 metres. ‘¢ The whole
of the ventral surface of the animal,” he continues, ‘ from
the snout to the root of the tail, was glowing with a feebly
shining greenish lustre, as if it were impregnated with phos-
phorus or had been coated with a luminous paint, with this
difference, however, that luminosity appeared and disappeared
at short intervals, but invariably increased in intensity just
before its disappearance.”
In this wise Johann’s interpretation was indisputably
confirmed. He further, too, more fully entered into thuse
questions of their physiology which are so closely connected
with these remarkable structures ; and he finally endeavoured
to supply proof of their presence in other Selachians which
he examined for this purpose, without, however, being able to
do so.
The genera investigated by him with the aforesaid object
were :— Stegostoma, Carcharias, Scylium, Pristiurus, Muste-
lus, Crossorhinus, Zygena, Chimera, Centrophorus (granu-
losus), Lamargus (borealis), and Scymnus.
At the time when Johann published his work I had already
noticed some peculiar structures in Lemargus rostratus, a
37*
560 Prof. R. Burckhardt on the
rare shark from the Mediterranean. These I had put aside
for the purpose of future microscopical use, pending the
completion of other anatomical rescarch; I then concerned
myself with this shark. Unquestionably they were organs of
luminosity, of the correctness of which opinion I became
more firmly convinced, since a splendid specimen of Spinaw
niger afforded me an opportunity for confirming Johann’s
observations.
When, in the spring of 1898, I was working at the Zoolo-
gical Station in Naples, Dr. Salvatore Lobianco, with his
wonted kindness, seized upon the first opportunity that pre-
sented itself to procure for me half adozen living specimens
of this species, which, however, arriving one evening, died
the next morning. | was greatly struck at the time by the
splendour of the spectral colours which these fishes exhibited,
and of which, so far as I am aware, no mention anywhere in
literature seems to have been made. This latter circumstance
induced me therefore to prepare a coloured sketch of this
phenomenon from these fishes.
A later scrutiny of this sketch convinced me of the fact
that I had been able to observe the phosphorescence of these
organs by daylight, so strong was their luminous power *.
{t is not my intention here to enter into details relating to
the purely histological modifications of these organs amongst
other Selachians, feeling sure that they all share more or less
the type of form which Johann has ascribed to them for
Spinag.
A tew figures, however, which I made of Lemargus ros-
tratus may well find a suitable place here—firstly, because
they will prove that the organs observed by myself are really
identical with those of Johann; and, secondly, they go to
show, as before stated, that the organs pertain to that form
whose merpholegical structure is so much affected by the
scales, as is the case with many of the ‘leleostean fishes.
‘The transverse section made through the luminous organ
of Lewargus rostratus (fig. 1) shows an epidermal swelling,
projecting crater-like above the ordinary level of the epidermis.
Beneath a few layers of epidermal cells of normal texture are
some which contain a prismatic corpuscle. The actual limit
* It should be stated here that with regard to its outer appearance
Spinax has repeatedly been the victim of misrepresentation, f. 2. :—
1. In the figure which Rafinesque gives of it, by having three gill-
openings only.
2. Its natural colours were quite unknown to all iconographers of
fishes, from Bonaparte to Fries. .
3. In being represented with its abdomen turned upwards in
Moreau’s drawing, without any reasons given for so doing.
Luminous Organs of Selachian Fishes, 561
between the epidermis and the cutis could not be established
with precision on material which had been preserved in a
solution of formalin, Certain it is that here, too, an incur-
sion of large pigment-cells into this prominence took place by
means of the strand leading to it, and which itself is enveloped
Fig. Y.
Microscopical section of the luminous organ of Z. rostratus.
by them. This strand seems to consist of a fibrous tissue
admitting a nerve of considerable size. On comparing it with
the luminous organ described by Johann, it appears to me in
process of atrophy, and may possibly be met with only in a
state of perfect development in the fully gestated embryo,
afterwards becoming resorbed.
Fig. 2 shows the luminous organ in correlation with the
surrounding scales. From a bulbous centre, which latter
represents the swelling seen in fig. 1, extend on two sides
some epithelial strands which are covered by pigment.
These scales differ from the normal ones in the following
points :—
(1) By having a more abundant pigmentation of their
pulpa, thus making the parts which these organs occupy, and
which are already prominent under ordinary macroscopical
observation, more conspicuous.
562 Prof. R. Burckhardt on the
(2) By having their apices blunter than is the case with
the normally constituted scales, in particular those rows of
scales which are situated orally with respect to the luminous
organs.
(3) By their base not terminating in a simple rhomboidal
edge, but possessing finger-shaped projections which secure
for these organs a firmer hold.
Fig. 2.
iit ma
pine ee
agree
ZZ
A=
Scale-bases and pigment of the luminous organ of Z. rostratus.
In Lemargus borealis the luminous organ appears to have
arrived at a stage of further reduction because its histological
structure is more vestigial still than is the case with that
described by Johann.
Besides this, the scales which surround the luminous organs
of this species differ from the normal ones in their stronger
pigmentation only and in being more crowded, but not in
variety of form.
Luminous Organs of Selachian Fishes. 563
I will now proceed to the description of the topographical
distribution of these photogenic organs, concerning which it
should be said that it differs in every species.
In the genus Spinax these organs were discernible, besides
in S, niger, in S. pusillus and S. granulosus, of which latter the
type is preserved in the British Museum. In puszllus they are
distributed in an almost identical manner to those in S. niyer,
2. é. the whole of the black under surface is densely beset with
them, whereas on the upperside they are very widely dispersed
or isolated.
On examining several specimens I[ was greatly im-
pressed with the fact that these organs appear extremely
varied in point of distinctness, variations which are clearly
attributable to the amount of pigmentation consequent upon
the more or less fresh condition the specimens were in at the
time they were preserved.
Up to the present time no phosphorescent organs were
known to exist in the genus Lemargus, although L. borealis
has been known for many years and is a common inhabitant
of the Northern Seas.
The distribution of these organs in Lamargus rostratus is
as tollows:—On both sides of the head and immediately
in front of the opercles are two bands which seem to agree
with the posterior crura of a circlet consisting of these organs,
as described by Johann to exist on the corresponding portion
of the head in Spinax. The remaining bands are distri-
buted in irregular swarms, resembling small stripes, which
are placed transversely. Accumulations of this kind are
situated on the occipital region and also on the base of the
anterior dorsal fin orally. Moreover, an irregular row of
these organs runs along the body, beginning at the back of
the head and ending at the root of the caudal fin a short
distance from and above the lateral line. Lastly, there is a
short row, composed of seven of these organs, situated below
the hindmost gill opening, which converges, together with
the one on the other side, towards the ventral median line.
Excepting these latter, 1 have not discovered any more of
these structures below the lateral line.
Their distribution in Lemargus borealis is similar; but
whereas 125 of these structures can be counted on one side
of L. rostratus, in L. borealis their number is 95 only. On
comparing their distribution in the two species the following
are the principal differences in borealis :—
The number of occipital organs is fewer. ‘The preedorsal
Prof. R. Buseck havik on the
564
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'sypa..og snhuvuaT
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‘9nqp.igso.t snbipuvny
1
ive)
iD
‘eZIs ‘YOU | "SULTLO SNOUTLUNT Jo WOT|NLUSIp ay} Furmoyg ‘zpuogury snwouojpo.dngy
Luminous Organs of Selachian Fishes.
¢ “Bry
ie
566 Prof. R. Burckhardt on the
cluster is slightly removed towards the snout and more
reduced than that of L. rostratus. I met with some scattered
organs on the anterior dorsal fin itself and also above the
foremost gill-opening, while the supralateral row does not
extend beyond the posterior dorsal.
With regard to these organs in Lemargus brevipennis, I
was able, from a specimen in the Paris collection, to prove
their presence in this species, and to state the fact that they
have a general resemblance to those of Z. borealis. It is
quite probable that luminous organs are functional in the
majority of the Lamargide, but possibly so in the young only
of the two larger species, in which they most likely have
escaped observation.
It is to the authentication in Jsistius brasiliensis of these
organs, however, that I attach particular value.
Here, as in Spinax pusillus, similar conditions obtained
for the non-uniformity of these structures in the different
specimens which I consulted. For instance, while they
could be seen plainly enough in some of them, in others,
whose state of perfection previous to their preservation
was probably impaired, absolutely no traces of them were
perceptible.
I found their expansion in perfect accord with Bennett’s
statements concerning this species, who had studied its
sphere of luminosity. In place of the non-luminous portion
referred to by this author a yellowish-coloured zone was even
yet visible on the old spirit-specimen, having its origin above
the gill-openings in a breadth of about | centim., and widening
out ventrally to one of about 2 centim. The zone lying in
our figure between the dotted lines contained not a single one
of these organs, whilst just behind it the whole of the
ventral surface is densely beset with them, though they are
more widely dispersed towards the back.
Someisolated organs can nevertheless also be found scattered
along the back. In addition to those mentioned above, the
further occur on both dorsal fins. On the top of the head they
are equally as scarce as on the back of the body, reappearing,
however, in somewhat greater numbers dorsally of the poste-
rior angle of the eyes, below the eyes, on the spiraculum, in
the angle of the mouth, on the cheeks, and also on the rostrum,
In regard to number these organs stand to the scales in the
ratio of about four of the former to one of the latter on the
abdomen, as against one to every ten scales on the upperside.
Euprotomicrus Labordii is only slightly longer than
Isistius, but considerably darker in colour. Its luminous
organs are more strongly pigmented and recall to mind rather
those of Spinaw—though present only on the ventral side,
where they are densely massed and quite equally distributed,
Luminous Organs of Selachian Fishes. 567
without any interruption on the throat at all. There are
no fewer than twelve to every scale, and in some places
they accumulate behind each scale in angle-shaped groups.
4 nat. size.
Bigs te
Centroscyllium granulosum, Giinth.
Luminous organs of Paracentroscyllium ornatum, Alcock, n. gen. Nat. size.
Dorsally they scarcely reach to the lateral line, which they
certainly do not cross.
Centroscyllium granulosum was first described by Dr. Gtin-
ther. Qn the type specimen I found the luminous organs
568 On the Luminous Organs of Selachian Fishes.
rather evenly distributed over the entire specimen, though
somewhat sparser perhaps when compared with forms pre-
viously mentioned.
In fig. 7 I was obliged to have recourse to a restoration in
part after Centroscyllium Fabricii for the first dorsal and the
caudal fin, as well as for the distribution of the photogenic
organs, the skin of the specimen of C. granulosum having
sustained some damage. ‘These organs are also present in
C. Fabrict’, but unfortunately the specimen at my disposal
was likewise defective, a circumstance which entirely
hindered the study of the distribution of these structures in
this species.
An oceanic Spinacid, Paracentroscyllium ornatum, which
has been described by Alcock, seems to be more closely
allied to Spinawx than to Centroscyllium, to judge from the
description of its colours by this author. Only one of the
three specimens belonging to this species is in the British
Museum, and on this I found the organs of luminosity
present.
They are relatively large in size, and are confined merely
to a few places—for instance, on the muscular portions of the
pectoral fins, then again on the two ventrals, and on both
dorsal ones. They further occur on the inferior lobe of the
caudal, consisting of a single row only, in smaller groups and
more widely diffused over and below theeyes. Dorsally they
make their appearance between the spiraculum and a little
beyond the ventral fin, over the whole of the gill-region, and
towards the anus on the ventral surface.
It may here be remarked that I have searched in vain for
these photogenic organs in Centrophorus granulosus, C. cal-
ceus, UC. squamosus, Scymmodon ringens, Scymnus lichia, Cen-
trina, Notidanus, Echinorhinus, and Chlamydoselache.
In conclusion, it will be seen that organs of phosphorescence
exist in eleven species of Selachians, inclusive of Jsistius
brasiliensis (Bennett) and of Spinax niger (Johann).
In structural characters they are very similar, and exhibit
a primitive degree of development *.
The distribution of luminous bodies is characteristic of the
respective genera and species. All Selachians which possess
them are pelagic, and they belong to the family Spinacide
(Giinther) = Spinacide + Lemargide of authors.
* Luminous organs of a similar form have been described in a Teleo-
stean fish (The Phosphorescent Organs in the Toad-fish, Porichthys
notatus, Girard,” by Ch. Wilson Green, Journ. of Morphol. vol. xv.
1899), With reference to this I may mention that I have also noticed
their existence in the occipital region of the haddock.
On a new Species of Bat from West Africa. 569
LXAXIIT.—Mormopterus Whitleyi—a new Species of Bat
from W, Africa. By R. F. Scuarrr, Ph.D., B.Sc.
THREE specimens of a bat, which I was unable to identify
with hitherto published descriptions, were sent to me by
Dr. J. C. Whitley from Benin City in West Africa. Before
describing them I forwarded one of them to Mr, Oldfield
Thomas, of the British Museum, who was good enough to
confirm my opinion that it belonged to a new species. Ihave
great pleasure therefore in naming the bat after its discoverer,
Sc
Head of Mormopterus Whitleyi, Schartf.
Dr. Whitley, and herewith give a description of its most
salient external features,
Kars shorter than the head and approaching one another
on the forehead to within 2 millim. Both the ears and their
short tragus are rounded off above. The antitragus is well
developed and separated posteriorly by a distinct notch.
The extremity of the muzzle projects very much beyond the
mandible, and the end of the nose is distinct from the upper
lip. The nostrils open almost laterally. ‘The upper lip is
only very indistinctly folded and covered with thick bristles
and fine hairs. On the lower lip the bristles are confined to
the angle of the mouth, but the hairs extend all over it.
The gular sac is small.
The dentition being } 43%, M@. Whitley? differs from all
other species of the genus in the possession of four lower
incisors instead of the usual six.
The fur is dark brown above, the bases of the hairs being
pale. The underside is of a light reddish-yellow colour.
Nearly half the tail projects beyond the wing-membranes,
which reach to the ends of the tibie.
The dimensions of the type, now in the British Museum
(adult g in formalin), are:—Total length 81 millim. ;
head 22; body 36; tail 23; length of forearm 36, of lower
leg 27.
The height of the ear is 15 millim., the width 10.
570 Mr. G. 8. Miller, Jr., on a
Three other species of Mormopterus are known to science,
viz., M. albiventer, M. acetabulosus, and M. setiger. The
first of these is confined to Madagascar, the second has a
wide range from South-east Africa to Madagascar and
Mauritius, while the last has been taken in German Kast
Africa.
M. Whitleyi differs from M. albiventer in being somewhat
smaller, in having a well-defined antitragus in the ear-conch,
and from all the three in having only four lower incisors
instead of six.
M. Whitley? differs from M. acetabulosus and M. setiyer in
having the ears close together on the forehead, in the more
regularly triangulate shape of the ears, and in the squareness
of the tragus.
Finally, 42. Whitleyi differs from M. setiger in being
smaller and in having the tragus rounded off above.
LXXIV.—A new Bat from Peru.
By Gerrit 8. MILuer, Jr.
AMONG some Peruvian bats submitted for determination by
Mr. Oldfield Thomas there are three specimens whose super-
ficial appearance is much like that of Pépistrellus hesperus
(H. Allen), but whose structural characters show relationship
with the members of the genus Rhogeéssa. The peculiarities
of the ear and teeth, however, are so great that the species
cannot be placed in any of the known genera of its family.
TomoPEASs, gen. nov. ( Vespertilionide).
Type Tomopeas ravus, sp. nov.
Characters.—Similar to Rhogeéssa, H. Allen, except that
there are only four incisors in the mandible and the structure
of the ear is essentially like that of the small-eared species of
Nyctinomus. Har-conch provided with a distinct though
rudimentary keel occupying the same position as in the
members of the family Molosside. Anterior border of ear
terminating simply, and without trace of basal lobe or “ hem.”
Tragus low and rounded, no larger than in some of the
species of Nyctinomus, and barely visible in the dried skin.
Nostrils distinctly short-tubular. Upper lip broad and
spreading.
Dental formula: 7. 5, ¢. =3) pm. sag, Ms = 28.
new Bat from Peru. O71
Tomopeas ravus, sp. nov.
Type.—Adult male (skin and skull), no. 689, collection of
P. O. Simons, Yayan, Cajamarca, Peru (alt. 1000 metres),
November 4, 1899.
Characters.—Slightly larger than Rhogeéssa tumida, H.
Allen (total length about 75 millim., forearm about 33) ;
ear broader, its anterior margin concave below tip, its posterior
Margin convex to base of antitragus; colour yellowish grey,
almost exactly like that of Pipistrellus hesperus from the
desert regions of the western United States, and more pallid
than in any known species of Rhogeéssa; skull slender, the
rostrum and brain-case depressed ; teeth much as in Rhoge-
éssa tumida, but first and second upper molars with more
distinct hypocone,
a, Tomopeas ravus; 6. Rhogeéssa tumida.
(Somewhat enlarged.)
Muzzle and lips.—Muzzle produced in conformity with the
broad lips, its extremity 3°5 millim. in front of incisors.
Front of muzzle naked except for a sprinkling of fine short
hairs at middle on upper surface. Nostrils with rim of aper-
ture produced into a distinct though very short tube. The
rim is slightly more produced at upper outer edge of nostril,
forming there a distinct blunt point. Upper lips thin and
widely spreading, the outer surface irregularly wrinkled and
little swollen, the inner surface distinctly and finely trans-
verse-wrinkled. Edge of lip conspicuously fringed, the
hairs longest about midway between nostril and angle of
mouth, but the fringe continuous across lower surface of
muzzle. ‘The lower lip is unusually tumid in front, but
otherwise it shows no peculiarities.
Ears.—The ear is moderately long ; laid forward it extends
to nostril. The anterior margin arises simply and without
trace of lobe or “hem” about 1 millim. above inner canthus
of eye. It is at first abruptly convex, the convexity ex-
tending through lower three fourths, though less distinct
above middle. At termination of main convexity the outline
becomes abruptly concave. Beyond this it is straight to
572 Mr. G.S, Miller, Jr., on a
narrow rounded tip. Posterior margin convex from tip to
antifragus, though nearly straight through upper third. Anti-
tragus small but well developed, and sharply defined from
rest of ear. It is roughly rounded-triangular in outline, the
anterior border faintly convex, the posterior slightly concave.
The anterior border is continued forward to terminate at a
distinct wart on lower lip at angle of mouth. Inner surface
of conch nearly smooth, though inconspicuously sprinkled
with very fine hairs. About halfway between meatus and
tip of ear two or three faint cross-ridges may be detected.
From anterior base of tragus there extends forward and
upward a distinct keel about 3°5 millim. in length and nearly
as well developed as in Nyctinomus minutus. ‘Tragus scarcely
rising above level of eye. In form it is much like that of a
specimen of Nyctinomus Kalinowskii, though in size it is
actually, as well as relatively, somewhat larger. Anterior
border faintly concave immediately above base, then strongly
convex, the convexity passing directly into that of tip, and
this again into that of posterior border. Posterior border
expanded near base into a narrow upright plate whose surface
is vertical to that of tragus. From tip of tragus spring a few
hairs, the length of which is about equal to height of tragus
from anterior base.
Membranes.—The membranes are thin, delicate, and in no
way peculiar in structure. Wing from ankle, its attachment
on outer and lower side. Uropatagium ample, enclosing tail
to base or middle of penultimate vertebra.
Feet.—The foot is small, about one third as long as tibia.
Its structure calls for no comment. Calcar about as long
as tibia, its tip forming a small but distinct lobe. Keel
narrow and inconspicuous.
Fur.—The fur is soft and dense, its length over entire
dorsal surface about 8 millim. Beneath it is shorter. It is
very closely confined to body, barely extending on extreme
base of membranes. ‘The uropatagium both above and below
is sprinkled with fine hairs. Basal half of ear and entire face
densely furred.
Colour.—Dorsal surface pale wood-brown, faintly washed
with ecru-drab across shoulders. Underparts dull buff,
fading to whitish cream-buff posteriorly. Face, ears, and
membranes blackish. Basal half of fur everywhere dull
slaty grey.
Skull.—The skull of Tomopeas ravus resembles in a general
way that of Rhogeéssa tumida, but is less robustly formed,
and the brain-case is less elevated. Zygomata very slightly
flaring, much less prominent than im Rhkogeéssa tumida.
new Bat from Peru. 573
Viewed from the side, the rostrum is seen to be much flat-
tened as compared with that of Rhogeéssa. The flattening is
visible in dorsal view also, though less conspicuous. Ant-
orbital foramen separated from rim of orbit by a wide space,
in the middle of which the relatively large lachrymal foramen
opens directly outward. Bony palate as in Rhogeéssa tumida,
except that the backward extension behind plane of molars is
shorter and without median spine.
Teeth.— Except in their uniformly smaller size and in the
absence of the outer incisors the mandibular teeth do not
differ appreciably from those of Rhogeéssa tumida. ‘The
maxillary molars, on the other hand, are immediately distin-
guishable by their greater development. ‘The first and second
are each provided with a distinct hypocone separated from
the commissure of the protocone by a deep notch. In Rhoge-
éssa the hypocone is so merged with posterior commissure
of protocone as to have practically lost its identity. Third
molar much broader than in Rhogeéssa tumida, owing to the
presence of a well-developed metacone.
Measurements.—External measurements of type (skin):
head and body 42 millim.*; tail 32*; tibia l0; toot 5 (42);
calcar 11; forearm 33:4; thumb 4°6; second digit 27;
third digit 56; fourth digit 48; fifth digit 38.
External measurements of an adult female from Chosica,
Peru (in alcohol) : head and body 39*; tail 34*; tibia 10;
foot 56; calcar 12; forearm 32°6.
Measurements of an adult female from Tolon, Peru (in
alcohol) : head and body 36°4; tail 33; tibia 11; foot 5;
forearm (broken) 31; ear from meatus 12; ear from crown
10°4; width of ear 11°6; height of tragus from anterior
base 1°8.
Cranial measurements of type: greatest length 12°6;
basal length 11:4; basilar length 9; zygomatic breadth 7;
Jachrymal breadth 4°8 ; least interorbital breadth 3; greatest
breadth of brain-case above roots of zygomata 6°6; depth of
brain-case 4°4; mandible 8°6; maxillary tooth row (exclu-
sive of incisors) 4°4; mandibular tooth-row (exclusive of
incisors) 5.
Remarks.—While Tomopeas ravus is so readily distinguish-
able from other bats as to require no special comparisons, its
relationships are much involved. ‘The general external struc-
ture, so far as wings, membranes, feet, and tail are concerned,
shows no departure from a strictly Vespertilionine type. The
somewhat tubular nostrils and the broad upper lip show, on
* Collector's measurement.
Ann. & Mag. N. Hist. Ser. 7. Vol. vi. 38
574 Dr. E. Linnberg on Reptiles and
the other hand, a distinct tendency toward Molossine cha-
racters; while the ear, in all the essentials of structure, is
practically identical with that of some of the species of Nycti-
nomus. Probably the questions suggested by this combination
of peculiarities can only be answered after detailed study of
the skeleton.
The type specimen will be presented by Mr. ‘Thomas to
the British Museum * and the spirit-specimen from Chosica
to the United States National Museum.
LXXV.—Reptiles and Batrachians collected in German New
Guinea by the late Dr. Erik Nyman. By Dr. EINar
LONNBERG.
During his visit to New Guinea last year my late friend, the
Swedish botanist Dr. Erik Nyman, made a collection con-
taining the following reptiles and batrachians, which were
sent home to Upsala. It is not very large, which can easily
be understood, as Dr. Nyman laid most stress on making
large botanical collections, in which he succeeded. Never-
theless the zoological collection treated of in these pages is of
great value, as it includes several forms new to science and
also adds to the zoogeographical knowledge. All the speci-
mens were collected in German New Guinea. ‘The following
geographical names are found on the labels :—
Stephansort.
Simbang.—In a bay at the outlet of a river, a missionary
station on a hill 50 metres above the sea-level.
Sattelberg.—A missionary station about 800 metres above
the sea.
Friedrich Withelms Hafen.
Most of the specimens are from Sattelberg. Some of the
labels had, however, fallen off during transit. “ Sattelberg ”’
was written on all except one, on which “ Fr, Wilhelmus
Hafen ” was inscribed. Consequently one of the specimens
recorded in this paper, for which no exact locality is given, is
from Friedrich Wilhelms Hafen, but all the others are from
Sattelberg. In those instances where the locality is indi-
cated it is quite certain, and quoted from an original label
* (B.M. no. 0. 3. 1. 101.—O. T.]
Batrachians from German New Guinea. 575
tied to the specimen. ‘The fauna of Sattelberg is particu-
larly interesting, because it is to be regarded as most typical
for the interior of German New Guinea, and from that place
the best specimens of the collection have been obtained,
REPTILIA.
LACERTILIA.
Gymnodactylus pelagicus, Gir.
One example, Stephansort.
Gymnodactylus louisiudensis, De Vis.
Description.—Head large; snout longer than the orbit, the
diameter of which is nearly equal to its distance from the
nostril, but a little shorter than its distance from the ear-
opening ; forehead concave, loreal region also concave, espe-
cially anteriorly ; ear-opening roundish, about one third of
the eye. Body and limbs rather elongate. Digits somewhat
depressed at the base, strongly compressed distally. Head
granular, with small tubercles on the temporal, occipital, and
posterior part of parietal regions. Rostral subquadrangular,
its height being about three fourths of its breadth, with a
deep median cleft above ; a large supranasal, separated from
its fellow by a small azygous shield ; nostril pierced between
the rostral, the supranasal, the first upper labial, and three or
four small granules ; twelve upper and ten to twelve lower
labials ; symphysial triangular; two pairs of chin-shields,
median largest and forming a long suture behind the sym-
physial; throat granular. Upper surface of arm above the
elbow covered by subimbricate scale-like granules; with
this exception, body and limbs covered above with small
granules intermixed with small, round, feebly keeled tubercles ;
scales covering a fold extending on each side of the body,
from axilla to groin, enlarged to similar tubercles. Ventral
region covered by scales, larger and more imbricate towards
the middle of the body, where their diameter is considerably
greater than that of the dorsal tubercles. Male with a long
uninterrupted series of femoral and preeanal pores, about 30
on each side, forming aright angle in the middle, but (unlike
the condition in G. Lorie) this series is not preceded by any
additional preeanal pores. <A distinct preanal groove. Tail
cylindrical, tapering, covered by a heterogeneous lepidosis.
The proximal third bears the same kind of tubercles as on
the back, but the granules between them are larger, scale-like,
and subimbricate. Distally the tubercles disappear and the
35*
576 Dr. E. Lénnberg on Reptiles and
scales become transversely arranged, more or less. On the
lower side there is a median series of transversely enlarged
shields. Reproduced tails have a less regular lepidosis.
Greyish brown above, a broad, dark brown, light-edged band
from eye to eye over the nape; a saddle-band of the same
kind over the shoulders, and two similar ones behind each
other on the back; the male with a third similar band
across the loins. Tail with broad dark brown or blackish
rings, which are less regular on the lower side. Lower parts
of body and limbs uniformly brownish white (in spirit). Two
smaller specimens, probably young of the same kind, are
brown above, with narrow undulated bands of a darker shade
across the back ; tail as in the large specimens.
“ 3.
millim. millim.
Total lengths. <heiewine =e 185 203
Le (=)¥0 Vie Beni Ac Buekok nee 27 33
Width of head’. <., .caakan 19 23
GAY hehe von aiie eee 63 84
DRAIN TM Yapett ates who each tees 95 86 *
One male, two female, and two young specimens from
Sattelberg.
Gonyocephalus dilophus, Dum. & Bibr.
T'wo specimens, Sattelberg.
Gonyocephalus modestus, Meyer.
One specimen.
Varanus indicus, Daud.
A young specimen.
Lygosoma elegantulum, Ptrs. & Dor.
Three specimens.
Lygosoma jobiense, Meyer.
One specimen,
Lygosoma Baudint’, Dum. & Bibr.
One specimen.
* Distally reproduced.
Batrachians from German New Guinea. 577
Lygosoma rufescens, Shaw.
One specimen.
OPHIDIA.
Chondropython viridis, Schleg.
“Yamomong.” ‘Two specimens from Sattelberg. ‘The
larger is dark bluish black (in spirit), with small, widely
scattered, yellow spots, which usually only occupy one scale.
These spots are not systematically arranged, and are found as
well on the back as on the sides. ‘he scales in three or four
of the outer rows are edged with yellow, and those in the
series next to the ventrals are completely yellow, like the
latter. The young specimen is entirely different in colour,
being yellow with red markings. ‘These are partly regular.
A streak running from the temporal region through each eye
and joining that from the other side just above the rostral; a
short transverse streak between the eyes; a \ f-shaped
band on the parietal region; a round spot on the nose and
two elongate rings in front of the parietal band; a longitu-
dinal median streak beginning on the nape and extending to
the tail; in connexion with the latter small subtriangular
rings, often alternating with each other and with the tips
pointing downwards; on the flanks numerous small rings
and round spots of a paler colour than the other markings.
The tip of the tail is provided with dusky spots. All
these markings make the specimen deserve the epithet
“ beautiful’ written by the collector on the label; but it
seems peculiar that the young animals should have such a
striking coloration, when that of the full-grown animal might
be termed protective. The body of this young snake is
extremely compressed. At the middle it is fully 2 centim.
deep, but hardly 1 centim. broad. The back is compressed
to an edge.
Enygrus asper, Gthr.
Two specimens from Stephansort.
Tropidonotus Mairi, Gray.
Two specimens, both with the frontal once and two thirds
as long as broad. ‘The number of ventrals resp. 145 and 154,
and the number of caudals resp. 66 and 88.
578 Dr. E. Lonnberg on Reptiles and
Stegonotus modestus, Schleg.
Two specimens.
Dendrophis calligaster, Gthr.
Two specimens, one resembling var. D in Boulenger’s
‘ Catalogue of Snakes,’ the other var. G. (¢bid.). Both are
me) ’
from Simbang.
Dipsadomorphus irregularis, Merr.
‘T'wo adult and one young specimen from Simbang.
PSEUDAPISTOCALAMUS, gen. nov.
This genus is allied to Apistocalamus, Blgr., and Pseud-
elaps (D. & B.), Blgr.
Maxillary extending forwards fully as far as the palatines,
with two grooved teeth, anteriorly followed by four grooved
teeth, gradually, but feebly, decreasing in length. Head
small, depressed, hardly distinct from neck. Eye very small,
with round pupil. Nostril pierced in a single nasal, but the
margin of the latter bordering the first labial and the inter-
nasal comparatively narrow; a large preocular, in contact
with the single nasal. Body cylindrical; scales smooth, in
15 rows; ventrals rounded. ‘Tail short ; subcaudals in two
rows.
Pseudapistocalamus Nymani *, sp. n.
Snout short, bluntly rounded. Rostral broader than deep,
visible from above, but the visible portion hardly one fifth of
its distance from the frontal ; internasals rather small, about
half as long as the prefrontals. Preefrontals not quite so
long as the frontal. Frontal almost as broad as long, almost
as long as its distance from the rostral, much shorter than
the parietals ; preeocular single, longer than deep, forming a
suture with the nasal. Normally two postoculars, upper
much larger than lower, sometimes both fused to one. Tem-
porals 1+2; six (or seven) upper labials, third and fourth
entering the eye, sixth largest, sometimes divided; five lower
labials, three in contact with the anterior chin-shields; poste-
rior chin-shields smaller, more or less separated by a scale,
which sometimes reaches the anterior chin-shields, sometimes
* In memory of the late Dr. Erik Nyman:
Batrachians from German New Guinea. 579
extends only to the middle of the posterior ones. Scales
smooth, in 15 rows. Ventrals 196-205; anal divided; sub-
caudals 26-29. Dark bronzy brown (almost blackish in the
largest specimen), somewhat iridescent above, lighter on the
flanks; ventrals and subcaudals blackish brown, edged with
light grey (in the youngest specimen the outer rows ‘of scales
coloured in a similar manner). Upper lip and a spot on each
side of the nape yellowish, as is also a transverse band across
the prefrontals from lip to lip, althongh the latter is less
pronounced.
The largest specimen ‘measures about 445 millim. in total
length; tail about 47 millim. ‘This specimen is a female
and contains eggs measuring about 13 millim, in length.
Three specimens from Sattelberg.
BATRACHIA.
ECAUDATA.
Rana papua, Less.
Three large and five half-grown specimens from Sattelberz.
The interorbital breadth compared with that of the upper
eyelid seems to be variable, and to be greater in old specimens.
Cornufer corrugatus, A. Dum.
Several specimens from Sattelberg.
Mantophryne robusta, Bley.
A fine “specimen from Sattelberg. This species was
described in 1898 from specimens collected on Bt. Aignan
Island, British New Guinea.
HHyla eucnemts, sp. n.
Tongue broadly heart-shaped, nicked and free behind.
Vomerine teeth in two large but confluent groups, behind the
level of the choane, which are large, but not quite so large
as the patches of vomerine teeth. Head large, about as
broad as long. Snout subtriangular, the tip truncate, longer
than the diameter of the orbit; canthus rostralis very distinct,
curved ; loreal region concave. Interorbital space broader
than the upper eyelid. ‘l'ympanum distinct, about two thirds
the diameter of the eye. A glandular fold extending from
the eye backwards above the tympanum. Three outer fingers
extensively webbed, the membrane reaching the disks of the
580 Dr. Ei. Linnberg on Reptiles and
secoud and fourth; disks about the same size as_ the
tympanum ; toes entirely webbed ; subarticular tubercles well
developed. The hind limb being carried forward, the tibio-
tarsal articulation reaches just beyond the eye. Upper
surfaces of body and head very minutely granulate; belly
and lower surfaces of thighs coarsely granulate. Throat
covered with scattered warts; a series of similar warts along
the margin of the lower jaw. A well-developed denticulated
fringe along the outer side of the forearm and the outer finger ;
a similar fringe along the outer side of the tarsus and the outer
toe. Heel with a triangular dermal appendage ; some smaller
dermal flaps near the vent. Colour (in spirit) dark plum
above, uniform, with the exception of some very indistinct
cross-bands on the hind legs; light below. The largest
specimen measures 67 millim. from snout to vent.
Two specimens from Sattelberg.
With regard to the great development of the web between
the fingers, this species resembles Hyla gracilenta, Ptrs., but
differs from it in so many characteristics that there can
hardly be any close relationship between them.
Hyla dolichopsts, Cope.
A large specimen from Sattelberg, and a young from
which the following notes are taken :—
The vomerine teeth being wanting, this little frog might
be classed as belonging to the genus Hylella.
Tongue almost round, nicked and free behind. Head
longer than broad. Snout longer than the eye; canthus ros-
tralis indistinct, rounded; loreal region concave. Interorbital
space much broader than the upper eyelid. ‘'ympanum dis-
tinct, two thirds the diameter of the eye. Fingers webbed at
the base; toes nearly entirely webbed. Disks smaller than
the tympanum. ‘The hind limb being carried forward, the
tibio-tarsal articulation reaches the tip of snout. Upper sur-
face of head granulate; back and upper surface of fore legs
also granulate, but less coarsely ; a fold over the tympanum.
Belly coarsely granulate; lower surface of thighs less so.
Colour (in spirit) dark bluish brown, light below; a white
streak along the margin of the lower jaw to the axilla; a less
distinct similar streak along the outer side of the tarsus.
Hyla obsoleta, sp. n.
The most striking characteristic of this form is that the
tympanum is completely covered by the granular skin, so that
no trace of it can be seen outwardly.
Batrachians from German New Guinea. 581
Tongue broadly rounded anteriorly, nicked and free be-
hind. Vomerine teeth in two small well-separated groups
between the choane. Snout subtriangular, longer than the
diameter of eye, truncate at the tip; canthus rostralis distinct ;
loreal region concave ; interorbital space broader than the
upper eyelid. Fingers with a short web at the base; toes
nearly entirely webbed ; disks about half the diameter of the
eye. A slight cutaneous fold with a row of tubercles along
the outer side of the forearm; a less pronounced row of
tubercles along the outer side of the tarsus ; a short dermal ap-
pendage at the heel. The hind limb being carried forward,
the tibio-tarsal articulation reaches beyond the snout. Upper
surfaces minutely granulate ; a fold from the eye to the axilla ;
another fold across the throat. Belly and lower side of thizhs
granulate. Colour (in spirit) dark brown above, light below,
without markings. From snout to vent 34 millim.
A single specimen from Simbang.
Hyla impura, Peters & Doria.
Habit slender. Tongue subelliptic, slightly nicked and
free behind. Vomerine teeth in two obliquely transverse series
between the choane. Head longer than broad ; snout longer
than the diameter of the eye, rather acuminate; canthus ros-
tralis angular, loreal region concave ; interorbital space broader
than the upper eyelid. Tympanum distinct, but small, about
half the diameter of theeye. Fingers nearly free, a short web
at the base of the outer fingers. Second finger longer than the
first, which has a brown wart on its inner side in the male ;
disks of fingers somewhat smaller than the tympanum. ‘Toes
broadly webbed, the web reaching the disks of the fifth and
third toes; disks of the toes smaller than those of the fingers ;
subarticular tubercles well developed. A large inner meta-
tarsal tubercle connected by a short web with the hallux, thus
producing the impression of arudimentary toe; another small
metatarsal tubercle at the base of the third toe. The hind limb
being carried forward along the body, the tibio-tarsal articu-
lation reaches between the eye and the snout. A cutaneous
fold along the side from above the tympanum, another fold
across the throat. Skin minutely granular above, coarsely
granular on the belly and under the thighs; scattered granules
on the throat and a row of similar granules along the forearm.
Greyish brown (in spirit), with some irregular darker brown
markings; there is, however, no anal blotch as described in
H. impura.
This species is represented by a single female specimen
582 Bibliographical Notice.
from Sattelberg, which agrees well with Hy/la impura as
described by Mehely (Termeszetr. Fiizetek. 1898).
The herpetology indicates, as other branches of zoology,
the mixed origin of the fauna of Papuasia. There are ancient
forms, such as Geckonide, ‘l'yphlopide, and Boide, the
members of which have had time to become widely distributed,
cosmopolitan so far as the climatical conditions allow. it.
Others originating from Asia reach their extreme limit, in a
south-easterly direction, in Papuasia (and Australia), as, for
instance, Dendrophis, Dipsadomorphus, Tropidonotus*, and
Ranide. Another group has reached New Guinea from Aus-
tralia, and there become checked from further distribution :
Hylidae is typical in this respect. ‘l'his family, being chiefly T
Australian and South-American, has extended from the latter
to North America, and from there to Hurasia, as its distri-
bution clearly shows. This family is thus a typical repre-
sentative of Australia in the fauna of Papuasia. But the great
number of endemic genera and species in the herpetological
fauna of Papuasia indicates its long isolation.
BIBLIOGRAPHICAL NOTICK.
The Story of Bird-life. By W. P. Pycrart,
London: George Newnes, Ltd., 1900. Pp. 244.
Tus recent addition to the ‘ Library of Useful Stories” is by a
rising ornithologist whose name is well known to specialists by
the admirable work which he is doing in the physiology and com-
parative anatomy of birds. We believe that this is about his first
appearance before the general public, and we hope it will not be the
last, for he has succeeded in compressing a large amount of useful
elementary information respecting birds, much of which is not
easily accessible in a convenient form elsewhere, into a very
moderate compass. The structure of birds, especially of the wings
and feathers, their courtship, nests, migration, geographical distri-
bution, pedigrees, &c. are discussed in an easy and attractive style
in twelve chapters. There is also a series of text-illustrations,
among others a restoration of Archwopteryx, and two full-page
illustrations, ‘‘ The summer home of the Ruff and Reeve ” and ‘* The
love-display, or ‘ showing off, of the Great Bustard, Otis tarda.”
Those interested in the latter subject will find a beautiful group of
Great Bustards in the Ornithological Gallery of the Natural History
Museum at South Kensington. We congratulate Mr. Pyeraft on
the production of this interesting little book, for it is not every
specialist who is able to write successfully and popularly on his
subject, as well as scientifically.
* The two former perhaps on driftwood or by the agency of man, the
last by swimming.
+ Like Cystignathide.
Geological Society. 583
PROCEEDINGS OF LEARNED SOCIETIES.
GEOLOGICAL SOCIETY.
February 7th, 1900.—W. Whitaker, B.A., F.R.S.,
President, in the Chair.
The following communication was read :—
‘Foraminifera from an Upper Cambrian Horizon in the Malverns,’
By Frederick Chapman, Esq., A.L.S., F.R.M.S.
The foraminifera described in this paper were found in a shaly
limestone which Prof. Groom obtained ‘from the débris of a
small ridge composed of black shales, with intercalated basalts,
which forms a spur on the north-west side of Chase End Hill.
The rock belongs to the well-known and widely-spread zone of
Spherophthalmus, Peltura, and Ctenopyge, which in Britain forms
the upper half of the Dolgelly Beds or Upper Lingula-Flags.’ The
specimens have been sliced, and have yielded a few forms other than
Spirilina; sections of echinoderm-spines, ostracod-tests, and occa-
sionally sponge-spicules (?) are also to be seen in the slides. The
tests of the foraminifera are infilled with a crystalline substance.
The following species are mentioned :—Lagena levis, Montagu,
L. apiculata, Reuss, LZ. ovum, Ehrenberg, Nodosaria pygmea (?)
Terquem, NV. abnormis(?) Reuss, Marginula soluta(?) Reuss,
Cristellaria acutauricularis (?) (Fichtel & Moll); and a new species
of Spirillina is described. The Spirillinew are in a good state of
preservation ; and the valves of mollusca, cut through in section,
are sometimes seen to be quite filled with the tests of this genus.
A record is given of some of the earliest known foraminifera,
including the somewhat doubtful forms described by Cayeux from
pre-Cambrian rocks in Britanny, those by Ehrenberg from the so-
called ‘Silurian Clay’ near St. Petersburg, by W. D. & G. F.
Matthew from the rocks bearing the Protolenus-fauna in New
Brunswick, by Keeping from the shales above the Bala Limestone
near Welshpool, by Blake from the Llandovery of Cwm Symlog,
by Brady from the Woolhope Limestone of the Malverns, and by
Terquem from the Upper Silurian rocks of Indiana. The Author
has also frequently met with Zagena in the Wenlock Limestone of
Shropshire.
584
INDEX to VOL. VI.
ABLABES, new species of, 409.
Abriesa, characters of the new genus,
312.
Actea pulchella, note on, 251.
Agelena, new species of, 199, 326.
Akodon, new species of, 468.
Alcimus, new species of, 176.
Alepas, new species of, 195,
Amaxia, new species of, 66.
Amphipoda, revision of the British,
32.
Anasida, new species of, 59.
Anaxa, new species of, 308.
Anaxandra, new species of, 229,
Ancistrogaster, new species of, 85.
Ancylis, new species of, 439.
Anderson, Dr, John, on the presence
of Echis coloratus in Africa, 419.
Andersonia, characters of the new
genus, 528.
Andrena, new species of, 19.
Anechura, new species of, 79, 99.
Anoplocnemis, new species of, 373.
Apeleutherus, characters of the new
genus, 73.
Apobletes, new species of, 268.
Apterygida, new species of, 83.
Apus, on the name, 248.
Arachnida, new, 199, 316, 363, 489.
Araneus, new species of, 324.
Archips, new species of, 442.
Argentina silus, on the abundance of,
on the market of Boston (Lincoln-
shire), 160.
Argyroploce, new species of, 236.
Arnouldia, new species of, 398.
Asopine, revision of the, 55,
Aspongopus, new species of, 221.
Asteromphalus malleus, new variety
of, 487.
Atylotus, new species of, 165.
Automolis, new species of, 66.
Auxinus, new species of, 323.
Bactra, new species of, 334.
Barbus, new species of, 479.
Barilius, new species of, 480.
Barrett-Hamilton, G. E. H., on varia-
tion in the common hedgehog and
weazel, 243.
Basicryptus, new species of, 223.
Bather, F. A., on the Lower Paleozoic
Crinoids of Bohemia, 102.
Batrachians, new, 181, 186, 579; on,
from German New Guinea, 574.
Beblecia, characters of the new
genus, 71.
Bell, Prof. F. J., on the name of the
swift, 248.
Bernard, H. M., on the origin of the
segmented worms, 509.
Bessia, characters of the new genus,
320.
Birchippia, characters of the new
genus, 450.
Boarmia, new species of, 308.
Books, new:—Systéme Silurien du
centre de la Bohéme, vol. vii. pt. 2,
102; Hampson’s Catalogue of the
Arctiadée in the Collection of the
British Museum, 157; Willey’s
Zoological Results, pt. iv., 246;
Matthews’s Monograph of the Co-
leopterous Families Corylophidee
and Spheriide, 247; Lankester’s
Treatise on Zoology, pt. ii., 341;
Andrews’s Monograph of Christ-
mas Island, 348; Pike’s In Bird-
land with Field-glass and Camera,
425; Cory’s Birds of Eastern North
America, 426; Tutt’s Natural
History of the British Lepidoptera,
426; Catalogue of Eastern and
Australian Lepidoptera Heterocera
in the Oxford University Museum,
pt. u., 506; Cunningham’s Sexual
Dimorphism in the Animal King-
dom, 507; Newbigin’s Colour in
Nature, 507; Pycraft’s The Story
of Bird-life, 582.
Boulenger, G. A., on a new sea-horse
from Muscat, 51; on fishes from
the Falkland Islands, 52; on the
abundance of the greater siil-smelt
on the market of Boston (Lincoln-
shire), 160; on new batrachians
and reptiles from Peru, 181; from
Perak, 186; on new lizards from
Jamaica and British East Africa,
INDEX.
193, 194; on a new Species of
Ablabes, 409 ; on new fishes from
Lake Tanganyika, 478 ; on African
Silurid fishes, 520.
Broom, Dr. R., on a new theriodont
reptile from 8S. Africa, 314.
Bufo, new species of, 181.
Burckhardt, Prof. R., on the luminous
organs of Selachian fishes, 558.
Burr, M., on new species and a new
genus of Forficularia, 79; on For-
ficularia from Sarawak, 88.
Butler, Dr. A. G., on butterflies from
Hunan, 382.
Cadicera, new species of, 161.
Calleteera, new species of, 309.
Callianassa Martensi, remarks on, 261.
Calocaris, new species of, 476.
Camaricus, new species of, 522.
Cameron, P., on new genera and
species of Hymenoptera, 410, 495,
530,
Capoeta, new species of, 478.
Caponia, new species of, 321.
Carpenter, G. H., on two new British
spiders, 199,
Carposina, new species of, 121.
Catada, new species of, 311.
Cervicapra bohor, observations on,
303.
redunca, new subspecies of, 304.
Cervus, new species of, 204.
Cheetoceros, new species of, 483.
Cheetospania, new species of, 93.
Chapman, F., on foraminifera from
the Malverns, 583.
Chariobas, new species of, 325.
Chasmoptera, characters of the new
genus, 462.
Chelisoches, new species of, 96,
Chiloglanis, new species of, 522.
Chondropsis, new species of, 355.
Chondropython viridis, note on, 577.
Chrysichthys, new species of, 480.
Cirrhochrista, new species of, 313.
Cnephalocotes silus, on the occur-
rence in Britain of, 201.
Coccide, new, 389, 448.
Cockerell, Prof. T. D. A., on the
New Mexico species of Megachile
and a new Andrena, 7.
Coleoptera, new, 219, 265.
Conepatus, new species of, 466.
Crinoids, on the Lower Paleozoic,
of Bohemia, 102.
Crocidosema, new species of, 441.
Crustacea, new, 39, 195, 256, 472 ;
585
on Malacostracous, from Malaysia,
249,
Cryptoloba, new species of, 310.
Crystallus, new species of, 399.
Ctenochiton, new species of, 449,
Ctenomys, new species of, 301, 388.
Cyclopelta, new species of, 220,
Cyclopes didactylus, new subspecies
of, 502.
Cydrelichus, characters of the new
genus, 525,
Cypreea, new species of, 207.
Cyrtacanthacris, new species of, 881.
Cyrtomaia, new species of, 472.
Cytharischius, characters of the new
genus, 492,
Dactylopius, new species of, 395.
Deesia, new species of, 317.
Dalader, new species of, 376.
Deroca, new species of, 306.
Diatoms from Chincha guano, on, 481.
Diploglossus, new species of, 193.
Diptera, new, 161, 291.
Distant, W. L., revision of the Aso-
pine and Tessaratomine, 55; of
the Dinidorinz, Phyllocephalina,
Urolabidinze, and Acanthosomine,
220; of the Coreide, 366.
Draco, new species of, 189.
Dresserus, new species of, 322.
Druce, H., on new species of Hetero-
cera from South America, 64,
Dudua hesperialis, note on, 134.
Dycladia, new species of, 64.
Dyscliona, characters of the new
genus, 352.
Echis coloratus, on the presence of,
in Africa, 419.
Elasmopoda, new species of, 368.
Elasmostethus, new species of, 231.
Eligmodontia, new species of, 297.
Enarmonia, new species of, 438.
{piechinus, new species of, 290.
Epiplema, new species of, 307.
Epitoxus, characters of the new
genus, 278.
Equus, new species of, 465,
Erephrosis, new species of, 292,
Eressa, new species of, 305.
Eresus, new species of, 322.
Eriococeus, new species of, 393,
Erylus, new species of, 351.
Ethusa, new species of, 474.
Eucelis, new species of, 407.
Eucereon, new species of, 65,
Euchilichthys, characters of the new
genus, 522,
586
Eucosma, new species of, 337.
Eulota, new species of, 455.
Eumenophorine, synopsis of genera
of, 494.
Eusthenes, new species of, 61.
Evagrius, characters of the new
genus, 369.
Exartema, new species of, 124.
Fishes, new, 51, 55, 478, 522; on
remains of, from the Argentine
Republic, 1; on the luminous
organs of Selachian, 558.
Foraminifera from the Malverns,
on, 985.
Forficula, new species of, 82, 100.
Forficularia, new species of, 79; on,
from Sarawak, 88.
Frisimelica, characters of the new
genus, 225.
Frog, note on an hermaphrodite, 179.
Ganesella, new species of, 455.
Gasteracantha, new species of, 324.
Gazalina, new species of, 306.
Gehyra, new species of, 188.
Geological Society, proceedings of
the, 583.
Gonopsis, new species of, 224.
Gordonerius, characters of the new
genus, 56.
Green, E. E., on new Australian
Coccidee, 448.
Gregory, Prof. J. W., on the West-
Indian species of Madrepora, 20.
Grose-Smith, H., on a new species
of Papilio, 210; on a new species
of Ornithoptera, 388.
Gruvel, A., on a new species of
Alepas, 195.
Gude, G. K., on new Japanese land-
shells, 598, 453.
Gymnodactylus louisiadensis, note
on, 970.
Halter, new species of, 460.
Harkeria, characters of the new
genus, 597.
Hedgehog, on variation in the
common, 243.
Hedylepta, new species of, 313.
Hempel, A., on Brazilian Coccide,
389,
Hendecaneura, characters of the new
genus, 401.
Hermachastes, characters of the new
genus, 319.
Hermaphrodite frog, note on an, 179.
Heterometrus, new species cf, 863.
Heteroptera, new, 56, 221, 366.
INDEX.
Hippocampus, new species of, 51.
Hister, new species of, 279.
Histeridee, new, 265,
Hololepta, new species of, 266.
Holopterna, new species of, 368.
Homalopygus, new species of, 285.
Homoptera, new, 389, 448.
Hyla, new species of, 579.
Hymenoptera, new, 7, 410, 495,
530.
Hypephyra, new species of, 308.
Hysterocrates, new species of, 489,
Hysterosia, new species of, 446.
Hystrichoscelus, characters of the
new genus, 335.
Icaria, on the males of, 495; new
species of, 496.
Icerya, new species of, 389.
Ictidosuchus, characters of the new
genus, 314.
Idalus, new species of, 67.
Isbarta, new species of, 305.
Isistius brasiliensis, on the luminous
organs of, 566,
Tulus, new species of, 206.
Ixalus, new species of, 187.
Kaliella, new species of, 453.
Kirby, W. F., on Odonata from
Sierra Leone, 67; on African
Phasgonuride, 211; on insects
from the Yang-tse-Kiang, 380;
ou the Neuropterous family Nemo-
pteridee, 456.
Kirkpatrick, R., on sponges from
Funafuti, 345.
Kofoid, Dr. C. A., on Pleodorina
illinoisensis, 139 ; on Platydorina,
a new genus of Volvocide, 541.
Labus, new species of, 536,
Leelia, new species of, 307.
Lemargus, on the luminous organs
in species of, 558.
Lamadatha, characters of the new
genus, 538,
Lanchester, W. F., on Malacostra-
cous crustaceans from Malaysia,
249,
Laparus, new species of, 166.
Laphria, new species of, 171.
Laspeyresia, new species of, 429.
Latrunculia, new species of, 354.
Lepidoptera, new, 64, 121, 210, 234,
305, 334, 388, 401, 429.
Leptobrachium, new species of, 186.
Lewis, G., on new species of His-
teridze, 265.
Liopygus, new species of, 270,
‘INDEX. 587
Loénnberg, Dr. E., on reptiles and
batrachians from German New
Guinea, 574.
Lophonotus, new species of, 178.
Loxopera, new species of, 444.
Luffariella, new species of, 357.
Lycodes, new species of, 53.
Lycodon, new species of, 193.
Lycosa, new species of, 327.
_ Lydekker, R., on an undescribed
type of Rusine deer, 204.
Lygosoma, new species of, 191, 194.
McArdle, Capt. A. F., on some re-
sults of Indian dredging, 471.
Macromia, new species of, 77.
Madrepora, on the West-Indian
species of, 20.
Mammals, new, 138, 204, 294, 383,
387, 465, 466, 470, 569, 570.
Megachile, new species of, 7.
Melioptis, new species of, 311.
Melucha, new species of, 374.
Melvill, J. C., on new species of
Cypreea, 207.
Menardus, characters of the new
genus, 366.
Mereschkowsky, C., on diatoms
from Chincha guano, 481.
Metopa, new species of, 42.
Metopina, characters of the new
genus, 45,
Microcystis, new species of, 400.
Microhyla, new species of, 188.
Microstylum, new species of, 168.
Mictis, new species of, 372.
Miller, G. S., jr., on a new bat from
Central America, 470; on a new
bat from Peru, 570.
Mollusca, new, 207, 398, 453.
Montezuma, new species of, 535.
Mormopterus, new species of, 569.
Mozena, new species of, 375.
Myriopoda, new, 206.
Mytilaspis, new species of, 448.
Navyicula Hennedyi, new variety of,
481].
Nemopteride, notes on the, 456.
Neuroptera, new, 69, 460.
Norman, Canon A. M., revision of the
British Amphipoda, 82.
Notoccelis, characters of the new
genus, 283.
Nototrema, new species of, 181.
Ochrochira, new species of, 370,
Odonata, new, 69.
Odynerus, new species of, 504.
Olethreutes, new species of, 129.
Ommatius, new species of, 177.
Opharus, new species of, 67.
Opisthocosmia, new species of, 85.
Ornithoptera, new species of, 388.
Orthetrum, new species of, 75.
Orthoptera, new, 79, 89, 216, 381.
Oxymycterus, new species of, 298.
Pachycrerus, new species of, 274.
Paludicola, new species of, 182.
Palystes, new species of, 330.
Pammene, new species of, 435.
Papilio, new species of, 210.
Parana formation, on the age of the,
6.
Pelatea, new species of, 335.
Perixera, new species of, 310.
Peucetia, new species of, 329.
Pheecadophora, characters of the new
genus, 130.
Pheecasiophora, new species of, 135,
Phalonia, new species of, 445,
Pharmacis, new species of, 443,
Phasgonuride from the Transvaal,
on, 211.
Philodryas, new species of, 185,
Phractura, definition of the new
generic name, 527,
Phyllotis, new species of, 296, 467.
Pidorus, new species of, 805.
Placinastrella, new species of, 350.
Placinolopha, new species of, 350.
Platydorina, characters of the new
genus, 555.
Platylister, new species of, 271.
Platynopus, new species of, 56.
Platysoma, new species of, 273.
Plectotropis, new species of, 400.
Plectroninia, new species of, 347.
Pleodoriua, new species of, 139.
Pocock, R.I., on two English milli-
pedes, 206; on new Arachnida
from Cape Colony, 316; on the
scorpions of the genus Hetero-
metrus, 862; on new African
spiders, 489.
Polistes, new species of, 410.
Polyfibrospongia, new species of, 359.
Pompilus, new species of, 539.
Potamon, new species of, 256.
Prionolomia, new species of, 367,
Pristopera, characters of the new
genus, 309.
Probolosternus, characters of the new
genus, 276,
Procavia, new species of, 387.
Proctoporus, new species of, 185,
Proechimys, new species of, 300.
Promachus, new species of, 173.
Promops, new species of, 470,
588
Propedesis, characters of the new
genus, 122,
Prout, L. B., on the Ankiindung
eines systematischen Werkes von
den Schmetterlingen der Wiener-
gegend of Schiffermiiller and Denis,
158.
Pseudapistocalamus, characters of
the new genus, 578.
Psiloptera, new species of, 219.
Pterinochilus, new species of, 318.
Punnett, R. C., on an hermaphrodite
frog, 179.
Pyramidula, new species of, 454.
Reptiles, new, 182, 188, 198, 194,
314, 409, 578; on, from German
New Guinea, 574,
Rhegmatopoda leptocera, new name
for, 215,
Rhodoneura, new species of, 312.
Rhynchium, new species of, 530.
Ricardo, Miss G., on Diptera from
South Africa, 161 ; on new Pan-
gonine from 8. America, 291.
Rothus, new species of, 326.
Saprinus, new species of, 287.
Sastragala, new species of, 228.
Scharti, Dr. R. F., on a new bat
from West Africa, 569,
Schiffermiiller & Denis’s Ankiindung
eines systematischen Werkes von
den Schmetterlingen der Wiener-
gegend, on the date of, 158.
Scione, new species of, 2U1,
Sciurus, new species of, 138, 294,
385.
Scylaticus, new species of, 170.
Selachian fishes, on the luminous
organs of, 558.
Selenops, new species of, 331.
Sicarius, new species of, 321.
Sitalia, characters of the new genus,
285.
Skeletonema costatum, new variety
of, 485,
Solpuga, new species of, 316.
Sparassus, new species of, 550.
Sparatta, new species of, 92.
Spermatogonia antiqua, note on, 488,
Sponges from Funatuti, on, 345.
Spongiphora, new species of, 89.
Stasimopus, new species of, 319.
Stenocercus, new species of, 182.
IND EX.
Stenothoe, new species of, 39.
Stewart, F. H., on a variation in the
number of genital pouches in
Thalassema neptuni, 218.
Stictocarenus, new species of, 230.
Strongylopsalis, characters of the
new genus, 80.
Swinhoe, Col. C., on new eastern
and Australian moths, 305,
Synodontis, new species of, 480.
Tabanus, new species of, 164.
Tedania, new species of, 355.
Terapus, new species of, 286.
Teretrius, new species of, 289.
Tessarotomine, revision of the, 55.
Thalassema neptuni, on a variation
in the number of genital pouches
in, 218.
Thermorthemis, new species of, 72.
Thiodia, new species of, 405,
Thomas, O., on giant squirrels from
the Amazonian region, 137; on
new rodents trom Western 53.
America, 294, 3838; on the E.
African reedbuck currently known
as Cervicapra bohor, 303; on a
new dassie from Nyasaland, 387 ;
on Equus Penricei, 465; on new
Peruvian species of Conepatus,
Phyllotis, and Akodon, 466.
Tuomisus, new species of, 322.
Tohi, on the, 3035.
Tomopeas, characters of the new
genus, 670,
Tortricidae, new Asiatic, 121, 234,
333, 401, 429.
Tortrix, new species of, 443.
Trishoplita, new species of, 400, 454.
Trithemis, new species of, 69,
Tropidurus, new species of, 184.
Tylopsis, new species of, 216.
Urochela, new species of, 226.
Volvocine, key to the genera and
species of, 556,
Walsingham, Lord, on Asiatie Tor-
tricide, 121, 254, 333, 401, 429.
Waterhouse, C. O., on a new species
of Buprestidee, 219.
Weasel, on variation in the, 243.
Woodward, Dr. A.S., on fish-remains
from the Argentine Republic, 1.
Worms, on the origin of the seg-
mented, 509.
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CONTENTS OF NUMBER 31.—Seventh Series.
[. On some Fish-remains from the Parana Formation, Argentine
Republic. By A. Surrx Woopwarp, LL.D., F.L.S. (PlateL) ....
Il. The New Mexico Bees of the Genus Megachile and a new
Page
Andrena, By T. D. A. Cocxerext, Professor of Entomology, New |
Mexico Agricultural College
esevure eee et eee eeeeeee sere eee eee re
III. On the West-Indian Species of Madrepora. By J. W. ay
Greeory, D.Sec., F.G.8., Professor of Geology in the University of
Melbourne
ee ero eer eee eee eee eee eee eee rm ere eeereseeeerer eres
IV. British Amphipoda.—IV. Families Stegocephalide to Cdi-
st
ceride (part.). By Canon Norman, M.A., D.C.L., LL.D., F.RS., &e.
(Tate UT) cho cay eas Oho eae eee ae ee hea
V. Description of a new Sea-Horse (Hippocampus) from Muscat, :
By. G: A) Bouuenerr, ORB. 2015 ok seine wee sys ete Onn Ree
VI. A List of the Fishes collected by Mr. Rupert Vallentin in the
Falkland Islands, By G. A. Boutpnerr, F.R.S. With Notes by the
Collector
VII. Rhynchotal Notes.—V. Heteroptera: Asopine and Tessara-
LONE, BY West Li. DIBDANT : '5\0 cis vps, » Sn cete stale e ounaie: ate er Ra
VIII. Descriptions of some new Species of Heterocera from Tropical
South America. By Hurserr Drucr, F.LS. &e. 22.2... eee
IX. Report on the Neuroptera Odonata collected by Mr. E. E.
Austen at Sierra Leone during August and September 1899. ‘By
W. F. Kinsy, F.L.S., F.E.S. (Plate IL.) SHER Dipl athe abe Dee
X. Notes on the Fon rficularia.— V. Pete ae of new Species
ereer er ecer er er ee ee eer eer ee eee eee ewe ee eee ee wh ew et
and. anew Genus. By Matcorm Burr, F.Z.8., F.E.S. (Plate eh .
figs ABUT) ss ass A MURR SI
XI. Notes on the Forficularia.—VI. On a Collection: of Ponfiew
laria from Sarawak. By Maxcorm Burr, F.Z.S., F.E.S, _ (Plate TV.
Fn LO) ales CN eh a pce ee RRR Ghee bree
XII. The Lower Paleozoic Crinoids of Bohemia. By F. A.
Bare MAL EG Bre icp. oe aie oo oN Mars neces ole ee en a ie aia
XIII. Asiatic Tortricide. By the Rt. Hon, Lorp Warstvenant,
MEAs EL Dis, WRB ie ae o aiethlern olate whe teey bie Ste, ae tore RE ae an
XIV. On Giant Squirrels from the Amazonian Region, uy Oxp-
FIELD THomas
© 8.6) ole fee fe iene .@ © 00 © © 0 6 0 ee 2 wie @, 6 e Ose, ere eden 010, 6 ible Ib eis la
XY. Plankton Studies.—II. On Pleodorina illinoisensis, a new
Species from the Plankton of the Illinois River. By C. A. Kororp,
Ph.D. (Plates V. & VI.)
BIBLIOGRAPHICAL NOTICE.
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Hampson, Bart.
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CONTENTS OF NUMBER 82.—Seventh Series.
Page
XVI. Notes on Diptera from South Africa (Palenihe and — c
Asilide). By Miss GertrupE RIcaARDO..........00.0 00 ee cece 161
XVII. Note on a Hermaphrodite Frog. By R. C. Punnezrt, BA.
(Plate ER yi cc iie de lade mane egunie sede a Aiea salle wae ne atta 179
— XVII. Descriptions of new Batrachians and Reptiles collected by
Mr. P. O. Simons in Peru. By G. A. Boutenerr, F.R.S. ..... 181
XIX. Descriptions of new Batrachians and Reptiles from the Larut Aa
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XX. Description of a new Lizard from Jamaica. By Goa.
BovLenGen, Pek Fie Ces Pave a etaneete map atnasie seer ae Matas Aes 193
XXI. Description of a new Lizard from British East Africa. By —
GLA BoriENncER, BeBe igs Slee ec Ae ee a ea 194
XXII. On a new Species of the Genus Alepas (A. Lankesteri),
from the Collection of the British Museum. By A. Grover, Chargé
de Cours & la Faculté des Sciences de Bordeaux. (Plate VIII.).... 195
XXIII. Two Spiders new to the British Fauna. By Gzorez H.
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XXVII. Description of a new Species of Papilio from Bwool, North
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XXVIII. Notes on the Collection of African Phasgonuride formed
by Mr. W. L. Distant in the Transvaal &c., with Descriptionsof Two
new Species. By W. F. Kirsy, F.L.S., FES. j MGa os peta eipinioteea ee 211
XXIX. Note on-a Variation in the Number of theGenital Pouches __
in Thalassema neptum, Gaertner. By F, H. Srewarr, M.A., Gatty
Marine Laboratory,’ St: Andrews (48050 hs Pics P cdsce spas wha ds 218
XXX. Description of a new Species of Buprestide. By Cuas. O.
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XXXI. Rhynchotal Notes.—VI. Heteroptera : Dinidorine, Phyllo-
cephaline, Urolabidine, and Acanthosomine. By W.L. Distant .. 220
XXXII. Asiatic Tortricide. By the Rt. Hon. Lorn WatsineHaM,
MOA TLD. BRB SS UR ee 234
XXXIII. Note on Variation im the Weasel and Hedgehog.
By Gi EH, Baneerrblamiivon es eh tag ei nietane eee) ae ace 243
BIBLIOGRAPHICAL NOTICES.
Zoological Results based on Material from New Britain, New Guinea,
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1896, and 1897, by Arthur Willey, D.Sc. Lond., Hon. M.A.
Cantab. Part LY Coe eae a ee 246
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a MISCELLANEOUS. ae
Priority or Usage. By F. Jurrrey Brrr, M.A. &............... 248
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XXXIV. On some Malacostracous Crustaceans from Miliysia in
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CONTENTS OF NUMBER 35.—Seventh Series. ae
Pigs “
LVI. Asiatic Tortricide. By the Rt. Hon. Lorp Wasi
M.A., LL.D., F.B.S.
LYVII. Descriptions of some Species of Coccide eatleated by Ne
Mr. James Lidgett in Victoria, Australia. By E. Ernest oe “oe ‘ile
F.E.S8., Government Entomologist, Ceylon,” (Plate, Mb yal yl) eeiaaaes ae aye
LVIII. Descriptions of new th of Japanese Tand-Shelle. By
G RAGHEB ZR) or si ee nah: i
LIX. Notes on the Neuropterous Family roi gia? By Tae
WB. Karey FB BBS, ea ot wie + ere plars. nie rie we lyiaiie eit he
LX. On ELquus Penricei, a Representative of the Mountain Zebra
(Equus zebra, L.) discovered by Mr. W. Penrice in Angola. cs wi
Qi brn Pome eS ee ki Se aN ous ee le lee a 465
LXI. New Peruvian Species of Conepatus, Phy oe and Akodon.
By OMReLD PROWAB 0 Loa. eine here eee ie. area tS 466
LXII. A new Free-tailed Bat nt Central Aineciaes nae Gunert
SU) GL itil SF eee CaCO am WARE Naa YOLEN Pama TURN TENA \ ( AT70
LXIII. Natural History Notes from H.M. Royal Indian Mesias
Survey Ship ‘Investigator,’ Commander T. H. Heming, R.N., com-
manding.—Series IIT., No. 4. Some Results of the Dredging Spat
1899-1900. By A. F. McAnpte, B.A., M.B., Capt. L.MLS., igi :
Naturalist to, the Surveye jonas Boe Oaks et 471
LXIY. Diagnoses of new Fishes discovered by Mr. Ae E. s iysees YATRA 231
in Lake Tanganyika. By G. A. Bounenerr, FBS... 0... 478
LXV. Note on Diatoms from Chincha Guano. By C. somo ae
KOWSE Ys. (Plate; AVE) 58 tee ae aie olone be tetah > Ube tumles Kata cane 481
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Museum. By R. I. Pocock
LXVII. Descriptions of new Genera and Species of ieee
By P. CAMERON 0.06 ecu eee ee ne nee oles teen een ees B05
BIBLIOGRAPHICAL NOTICES.
Catalogue of Eastern and Australian Lepidoptera Heterocera in the _
Collection of the Oxford University Museum.—Part II. Woc-
tuina, Geometrina, and Pyralidina, By Col. C. Swinnor, M.A.,
E.LS., F.Z.8., FES. Pterophoride and Tineine by the Right —
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Ham, M.A
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CONTENTS OF NUMBER 36.—Seventh Series,
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LXVIII. A suggested Origin of the Segmented Worms, and the
Problem of Metamerism. By H. M. Bernaxp, M.A. Cantab. ..... . 509 :
LXIX. On some little-known African Silurid Fishes of the Sub-
family Doradine. By G. A. Bounenerr, F.RS. ..............4. 520
LXX. Descriptions of new Genera and Species of Hymenoptera.
By P. CamEeron
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the Family Volvocide from the Plankton of the Illinois River. By
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LXXII. On the Luminous Organs of Selachian Fishes. By
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LXXIIL. Mormopterus Whitleyi—a new Species of Bat from
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LX XIV. A new Bat from Peru. By Gerrit 8. Mitimr, Jr. .... 570
LXXV. Reptiles and Batrachians collected in German New Guinea
by the late Dr. Erik Nyman. By Dr. Ernar L6nnsere
BIBLIOGRAPHICAL NOTICKH.
The Story of Bird-life. By W. P. Pycrarr 58
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