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ALERE @ FLAMMAM,
CONTENTS OF VOL, XIX.
(EIGHTH SERIES, }
NUMBER 109,
I. Corylophide [Coleoptera] from the Seychelles and Rangoon.
By Huexu Scorr, M.A., F.L.S., Curator in Entomology in the
University of Cambridge. (Plates I.-V.) ...........ccceceeeees
If. Notes on Exotic Chloropide. By C.G. Lamp, M.A., B.Sc.,
2) Se O° a Ce i eT
IIT. Some Systematic Notes on Melolonthine Coleoptera. By
Bae Pat, ee IRONY 85, os 58: 5 .vi's ahgiyrs. nisrecelae etm dale satan mie « Fa gee wre et
IV. Descriptions of New Tyralide of the Subfamilies Epi-
paschiane, Chrysaugine,. Endotrichine, and Pyraline. By Sir
GrorceE F. Hampson, Bart., F.Z.S., &e. 2.1... PREPS Gee aoe
V. The Homoptera of Indo-China. By W.L. Distant........
VI. Notes on Fossorial Hymenoptera.—XXV. On new Sphecoidea
in the British Museum. By Row.anp E. Turner, F.Z8., FES.
VIJ. On the External Characters of the Felide. By R. I.
Pococs, F.R.S., Superintendent of the Zoological Society’s Gardens.
VIII. On some new Mites of the Suborder Prostigmata living on
eaeen ee CAML EAR ST tos ne we ned cn giene wed tenwass
IX. Cassidine and Bruchide [Coleoptera] from the Seychelles
Islands and Aldabra. By 8, Mauurx, B.A. (Cantab.)...........+
Page
144
iv CONTENTS.
Page
X. Notes on Fossorial Hymenoptera.—XXVI. On the Genus
Homonotus, Dablb. By Rowtanp E. Turner, F.Z.S., FES. .... 147
XI. Notes on the Species of the Genus Cavia. By OLDFIELD
ROMANS Ceci a ais aee valet ont + sss tinue mies ae Loe 7 - 152
New Book :—African Freshwater Fishes ........2+ceeeeeeeeres 160
a
NUMBER 110. .
XII. Coleoptera, Heteromera (excluding Tenebrionide) from the
Seychelles Islands arid Aldabra. By GroreE CHARLES CHAMPION,
Mei SAME AVE) anc oc, a Beaks « 9% wlemincssi luis «ib aip o.02h ge viele rae eee 161
XIII. On new Species of Indian Cureulionide.—Part III. By
LGA. Sh MA BBHAES » DOG. oo. Gs ca aly ne es +, 2 nist eeele 188
XIV. A Revision of the Clupeid Fishes of the Genus Pellonula
and of Related Genera in the Rivers of Africa. By C. Tate
EBGAN, WEA oe UN fects oe 8 Pea ee EReES > se wee ee he ei eine 198
XV. New Species of Tabanide from Australia and the Fiji
Islands.” By GueTRUDE RICARDOT: wears. oh ewan ae baunp 207
XVI. New Species of Hematopota from India. By GERTRUDE
PERRIN iets Ge Seo wien meet es Micon ate tet oe G Sree Re iets EARS 225
XVII. The Fishes of the Genus Clupea. By C. Tarr Reaan,
IT aA See Stet ces oe oa teroRebs fanars a tel cade canes nv we Rm ee emne ee Mean 226
XVIII. Barnacles from the Hull of the ‘Terra Nova’: a Note.
By 1; A. BORRADAILE «094%. 2 bee ween genes ae hse eee en han 229
Proceedings of the Geological Society...........0.cecaeeeee 230, 231
NUMBER 11}.
XIX. Notes from the Gatty Marine Laboratory, St. Andrews.—
No. XL. By Prof. M‘Inrosu, M.D., LL.D., F.R.S., &c., Gatty
Marine Laboratory, University, St. Andrews. (Plates VII.-XII.) . 233
XX. Notes on Exotic Helomyzide, Sciomyzide, and Psilide. By
C. G. Lams, M.A., B.Sc., Clare College, Cambridge ............ 266
‘CONTENTS. Vv
Page
XXI. Further Notes on the New Zealand Ampbipod Hyale
grenfelli, Chilton, By Cuas, CHILTON, M.A., D.Sc., M.B., C.M.,
LL.D., O.M.Z.S., Professor of Biology, Canterbury College, New
Zealand 273
XXII. Descriptions of new Lizards of the Family Lacertide. By
GovAc BouLenGur, PURIS So. i. ccc ive ee teteaied idee eee es 277
XXIII. A new Bat of the Genus Scotecus. By OLpFIELD
Be MIN ee Gh or ear eb olcke ato Siegert ffeleeoe Mania ope Not eae erage aber Kier 280
XXIV. A new Species of Aconemys from Southern Chili.
By OLDFIELD THOMAS 20... 000s cere eres eee t neice er enaenes 281
XXV. Descriptions and Records of Bees. —LXXIV. By T. D. A.
CockERELL, University of Colorado ......+ seer eee ee reer eens 282
New Book :—Catalogue of the Lepidoptera Phalene. Supplement,
Pca Mies a orci car hat sss eyes 3 ce Sw ead dime hohe nue: rao. ar aan 8a SA 291
Proceedings of the Geological Society .....+.+.+eseeeeeees 291—295
NUMBER 112.
XXVI. A Revision of the Clupeoid Fishes of the Genera Promo-
lobus, Brevoortia and Dorosoma, and their Allies. By ©. TatE
Rs eA sie eae ns Adel wes ye Whale vine ew ese HHH ao He's 297
XXVII. Notes on Fossorial Hymenoptera—XXVII. On new
Species in the British Museum, By RowLanp E, Turner, F.Z.8.,
Re oot Aca Sides naga Cai ate Roi ANA en el halGe ee we ws 317
XXVIII A new Tuberculate Terrestrial Isopod from New
Zealand. By Cuas. Cuinton, M.A., D.Sc. MBs, CM, LL.D;
C.M.Z.S., Professor of Biology, Canterbury College, New Zealand.
PT ee open gc rida aleh nape sien crete Mo Heald eso wo ere 027
XXIX. South-African Talitride. By the Rey. Tuomas R. R.
SUEBBING IG Rae ER in cee eens ec etle sso c essen nner eewentnes 330
vi CONTENTS.
Page
XXX. New Species of Indo-Malayan Lepidoptera. By Colonel
C.SWINHOM, MGA.) BAGS. 12 5. 2. ss ce boneeen kb pee Beta deve 331
XXXIJ. The Lemurs of the Hapalemur Group. By R.I. Pocock,
PEE 5 spo aye sai «o'er « i a's <'o. 2 Say ee eee eae 343
XXXII. Some Notes on Three-toed Sloths. By Oxprrexp
POMS 01% vila Wiese 25s tele 5 Se See Ale see tan dee 352
NUMBER 118.
XXXII. Descriptions of New Pyralide of the Subfamilies
Hydrocampine, Scopariane, &e. By Sir Groner F, Hampson,
Bart; RZ. 8. 2... 25s eee bc nag oe ae cs eee 561
XXXIV. A Revision of the Clupeid Fishes of the Genera Sardi-
nella, Harengula, &e. By CO. Tate Ruean, M.A. .............. 377
XXXYV. On new Weevils of the Genus Mecysmoderes from India.
By Goy A. K. Mans smepaieemamcer oi. eee ue. e ee as ae 395
XXXVI. Occurrence of a Holothurian new to the Fauna of
Bermudas. | By W. JAG nonin tc tence wwescds sper on teneesees 405
XXXVII. Descriptions of a new Lizard and two new Frogs
discovered in West Africa by Dr. H. G. F. Spurrell. By G. A.
BOULENGER, EURis.setis cos see tk sO un eee ete Chee aan ec 407
XXXVIII. New Species of Indo-Malayan Lepidoptera. By
Colonel C) Swinton sea ALS iy iis sins orcas sce kot eee cats 409
XXXIX. Notes on Myriapoda.—V. On Cylindroiulus (Leucotulus)
nitidus (Verhoeff), By Hitpa K. Brapsz,; M.Sc. L.R.C.P.,
M.R,C.S5., and the Rev. S. GRaHam Breks, M.Sc. ........00000 417
—
“
CONTENTS. ; Vil
NUMBER 114.
Page
XL. Notes on Collembola.—Part 4. The Classification of the
Coilembola; with a List of Genera known to occur in the British
Isles, By Joun W. Suorsoruam, N.D.A., Berkhampsted, Herts. 425
XLI. Notes on Fossorial Hymenoptera. — XXVIII. On new
Ethiopian Species of Bembex in the British Museum. By RowLanp
HE; TURNER, F°Z.5., FES. 2.0.6.6 s008 SOR es ee iy er 436
XLII. On Fabricius’s Types of Odonata in the British Museum
(Natural History). By Harpert Campion
XLIII. A new Vole from Palestine. By Otprretp THomas .. 450
XLIV. On the small Hamsters that have been referred to Cri-
cetulus pheus and campbelli. By OLDFIELD THOMAS............ 452
XLV. Descriptions of New Pyralida of the Subfamilies Hydro-
camping, Scopariane, &e. By Sir Groner F. Hampson, Bart.,
Sere Me Ral ase om Ulla a 5 Kuga oy PAL RH Sade an ote daa bag 457
XLVI. Descriptions and Records of Bees —LXXV. ByT.D. A.
Cocnunur, University of Colorado”... 0.600... es cee sccesceees 473
XLVI. The Khapra Beetle (Trogoderma khapra, sp: n.), an
Indian Grain-pest. By GILBERT J. ARROW ..........000eeeee 481
Proceedings of the Geological Society ........ccccccecccecuses 483
Notice to the Zoological Profession of a Possible Suspension of the
International Rules of Zoological Nomenclature in the Cases
of Musca, Linneus, 1758, and Calliphora, Desvoidy, 1830 .... 484
Index
PLATES IN VOL. XIX.
PLatE I
II
iG WE
IV
]
|
(vorylapand beetles.
<
VI. Heteromerous Coleoptera from the Seychelles and Aldabra.
VALT)
VIL.
<a >Nervous and muscular ayqreme of Owenia and Myriochele.
XI.
XII. J
XIII. Cubaris milleri, sp. n.
ERRATUM.
Page 330, line 14, for 1913. Taltriator, Matthews, P. Z.S. Lond. p, 109,
read Lalitriator, Methuen, P. Z. 8. Lond. p. 109.
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MAGAZINE OF NATURAL HISTORY.
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No. 109. JANUARY 1917.
1.—Corylophide [Coleoptera] from_the Seychelles and Ran-
goon. By Huau Scort, M.A., F.L.S., Curator in Ento-
mology in the University of Cambridge.
[Plates I.-V.] °
Tue main purpose of this paper is to give an account of the
Corylophid beetles obtained by the Percy Sladen Trust Expe-
dition of 1905 and 1908-9 in the Seychelles and other islands
of the Western Indian Ocean. But I have also included
certain forms taken at Rangoon in 1911. ‘The actual sources
of these two sets of material may be considered separately,
as follows :-—
(A) Rancoon.—The specimens were collected from a nest
of Munia striata, a bird belonging to the Ploceide or weaver-
birds, on Oct. 9th, 1911, by Dr. H. H. Marshall, M.O.H., and
sent by him in alcohol to Professor G. H. F. Nuttall at the
Quick Laboratory, Cambridge. Professor Nuttall kindly
handed over the Coleoptera to me. They consist of three
species of Corylophide — namely, Arthrolips flavicollis,
Matthews, Orthoperus muni@, sp. 0., and Orthoperus sp.
indet., as well as a single example of an undetermined
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. L
2 Mr. H. Scott on Corylophide from the
Cuenjid which seems closely allied to Silvanus longicornis,
Grouvelle, a form known from Singapore. In addition to
Coleoptera, the tube contained some Lepidopterous larvee, a
spider (Scytodes sp.), and some Gammasid mites, all from the
same bird’s-nest. I do not know of other recorded cases of
Corylophidz being found in birds’-nests, but I have myself
taken a specimen of Orthoperus from a blackbird’s or thrush’s
nest of the preceding year at Henley-on-Thames, 25. iii. 1910.
(B) Srycuettes Istanps.—It was intended that all
results of the Perey Sladen Trust Expedition should appear
together in one publication, but circumstances have rendered
this impossible. The work in question consists of certain
special volumes of Trans. Linn. Soc, London, five of which
are already complete (ser. 2, Zool., vols. XIl.—-xvi.), while a
sixth is in progress: these contain, inter alia, a number of
reports on insects. In the present paper much the same
plan is followed as in my two previous articles on certain
groups of Seychelles Coleoptera (op. cit. vol. xv, p- 215, 1912 ;
vol. xvi. p. 193, 1913).
No Corylophide have been recorded from these islands
before, ‘Those dealt with here amount to twelve species, ten
of which are described as new (see below, under “ determina
tion of species”), while one is undetermined and one is
referred to a previously described species, They belong to
eight genera, one of which is described as new. The series
may be briefly analyzed thus:—Suctum, 4 spp. n.; Arthro-=
liys, 1 sp. n., 1 sp. indet.; Meioderus, 1 sp. n. ; Sericoderus
(Anisomeristes), 1 sp. n. ; Daubania, g. n., 1 sp.n.; Lew-
istum, 1 sp. n.; Rhypobius, 1 sp. n, ; Orthoperus, 1 sp.
(previously known),
Listribution.—One species, Rhypobius aquilinus, was found
only on a coralline island of the Amirantes Group. The
other eleven were all taken by the writer in the mountainous
granitic islands of the Seychelles proper. Six of these were
found exclusively in the island of Silhouette, which was
visited only during the drier months of August and September ;
one was only obtained in Long Island, a small cultivated
islet near Mahé, in July, also one of the drier months ; the
remaining four, including the new genus Daubania, were
taken in two or more of the larger islands, and in both
the drier and wetter seasons.
Two species are represented by single specimens, two
(Sericoderus and Lewisium) by big series of over 50 and of
neatly 200 respectively, the remainder by series of from 3 to
15 examples. ‘They were all preserved dry.
Seven kinds were obtained only at high elevations, in the
Seychelles and Rangoon, 3
endemic forests; one (Arthrolips insule-longe), as stated
above, only onacultivated islet. Of the remainder, Daubania
gen, nov.) occurred in the high forests and at more moderate
elevations, while the two most abundant forms (Sericoderis
and Lewtstum) seemed generally distributed from the culti-
vated country up into the endemic woods at high altitudes.
Most of the material was collected somewhat promiscuously,
by general sweeping and beating of vegetation, but in some
cases I have exact records of the manner in which specimens
were taken. Thus some of the Sericoderus and of the
Lewisium were swept from long grass, and most of the
Sactum picaultianum were beaten from dead palm-leaves, a
very fruitful source of beetle-life. Two individuals of the
Lewisium were found in-a fallen branch containing an ant’s
nest (see p. 24), though whether their presence was acci-
dental or intentional I cannot say.
A ffinties.—The world-fauna of creatures so minute as
Corylophidze must be at present but very imperfectly known,
therefore it is not profitable to discuss at length the affinities
of the Seychelles series. Moreover, having regard to the
highly peculiar nature of the endemic vegetation, and to the
large number of peculiar insects and other animals existing
there, it is probable that some at least of the species herein
described will prove to be absolutely confined to these islands.
But such indications of affinities as exist may be briefly con-
sidered for what they are worth.
The only form referred to a previously described species
is Orthoperus minutissimus, Matth., hitherto recorded from
S. America and W. Indies. The new genus Daubania is
allied to Oligarthrum, known only from §. America, and to
Corylophus, widely distributed in Europe and Asia. Meio-
derus was previously reeorded only from Japan, Lewis/um
from Ceylon and Japan. The other genera are known from
all parts of the world.
The Corylophid fauna of Madagascar appears to be very
little known. ‘he only species included in Alluaud’s * Liste
des Insectes Coléoptéres de la Révion Malgache’* (p. 105)
is Sactum monstrosum (Schaufuss) ¢, which, from its descrip-
tion, seems’ quite unlike any of the Seychelles forms.
Matthews describes his Sacium bifasciatum (Mon. p. 54)
from Madagascar, and this is a little like my Saciwm p:caulti-
anum. J have found no further records of Corylophidze
* Vol xxi. of Grandidier’s ‘ Histoire Physique, Naturelle, et Politique
de Madagascar,’ Paris, 1900.
+ = Clypeaster monstrosus, Schaufuss, Tijdschr, Ent. xxxiv. 1891, p. 2;
Matthews, Mon. Corylophide, p. 217. 18
4 Mr. H. Scott on Corylophide from the
from Madagascar in tle subsequent literature. Rvitter’s
(1908) descriptions of E. African species have been studied,
but without seeing specimens it is hard to pronounce on ‘fran
relationships nauk those of the Seychelles. In comparing
the latter with forms in Matthews’s collection, I have several
times found that the nearest to the Sevchelles species are
Oriental forms, from Ceylon, Japan, &c. (¢f. the distribution
of the genera Meioderusand Lewisium, mentioned above), but
the resemblance. is’ not generally very close. However, if
these apparent indications of Oriental affinities should prove
genuine, this would only tally with what has been found
so strongly marked in certain other groups of Seychelles
insects.
On the whole, the Seychellean forms are very minute, even
jor Corylophidee. In comparing, I have been repeatedly
struck with their small size in relation to their congeners,
STRUCTURE.—Various anatomical points are dealt with
under the headings of particular genera and species. Thus
secondary sexual characters have come to light in Rhypobius
and Orthoperus, and differential specific characters in the
form of antenne and mouth-parts in certain species of Sacium,
Sericoderus, and Lewistum—in Saeium also in the form of the
prosternum. Attention is called to the presence of diverging
metasternal strize in Orthoperus.
The condition of the hind wings is stated, so far as it has
been examined, in the case of each particular species. I
follow Matthews in using the term “ample” to denote that
the wings are not reduced, vestigial, or absent, but much
longer than the elytra, under which they are folded. It
appears that they are ample i in ten out of the fifteen species
dealt with below, the remaining five being :—Arthrolips
sp. indet., wings present but could not be examined ; Arthro-
lips flavicollis, Matth., Orthoperus minutissimus, Matth., and
Orthoperus sp. indet., wings not examined ; Rhypobius aqui-
linus, sp. n., wings present and longer than the elytra in the
6, but seemingly quite absent in the 9. This last case is
interesting, exhibiting a sexual difference in the wing-
development. The genus Rhypobius (= Moronillus) is said
by Ganglbauer (Kit. Mitteleur. ili, pp. 273, 283-4) to have
the hind wings quite absent. Matthews makes the less
general statement (Mon. p pp. 172-3) that these organs are
absent in the “ genotype,” 2. marinus, Leconte, but says
nothing of their condition in the other species. Ina pair of
the European R. rujicollis (Duval) which I have examined IL
find no trace of hind wings in either sex. I have not investi-
gated their condition in other species of the genus.
>
Seychelles and Rangcon. 5
Matthews also states (Mon. pp. 109, 115) that the hind
wings are either absent or small and narrow in Svricoderus
and Andsomer/stes, but in those specimens of S. (A.) seychel-
lensis, sp. n., which I have dissected they are much longer
than the elytra. For the rest Matthews describes them as
“ample” in his diagnoses of all the other genera except six,
in which he either states that he had not examined them or
does not mention them at all. But in the case of some
genera examination of larger numbers both of species and
individuals is probably required.
TECHNIQUE.—Jn fixing the generic position of species I
have never relied on general appearance alone, but have in
all cases made balsam-preparations of antennze and mouth-
parts for examination under the compound microscope.
These preparations are mounted between two cover-slips, one
of which is attached to a cardboard framework ; the thinness
of the glass then allows of both sides of the object being
viewed through a high-power objective, while the cardboard
framework admits of the preparations being pinned beside
the insects. Balsam-preparations appear almost essential in
dealing with Corylophide, and are indispensable in describing
aly new genus.
Measurements of leneth have been made with a yr ola
micrometer-eyepiece. Drawings made with the aid of
drawing-apparatus,
For comparison I have used the British Museum Collection,
which, including Matthews’s Collection and his balsam-
‘preparations, is fairly complete up to the date of his ‘ Mono-
graph’ (1899). Descriptions of older forms not included in
the Monograph, and of all species and genera described since,
have been consulted.
LiTERATURE.—Matthews’s ‘ Monograph of Corylophide:
and Spheertidze ’ appeared in 1899, after its author’s decease.
A number of species unknown to ‘him were not included in
his manuscript, but the editor of the Monograph refers to
these on pp. 19-21 and p. 217. The Monograph may there-
fore be taken as a fairly complete enumeration of the species
up to and including 1899.
The following is a list of the subsequent literature, compiled
from the ‘ Zoological Record,’ the nature of each work being
bricfly indicated. Though a catalogue of the family has
receutly appeared, this list may also be of some use :—
1900. Casry. Journ. New York Ent. Soc. viii. pp. 60-75, review of
N. Amer. forms, describing several new genera and species:
Bathona, g. u., Gronevus, g.n., Eutrilia, g.u, near Orihoperus,
M olamba, ¢ g. n. near Sacrum.
6 Mr. H. Scott on Corylophidee from the
1900. Doprro. Ann. Mus. Genova, xl. p. 565, records Saciwm for-
mosum, Matth., from Burmah,
Reirrer. Wien. ent. Zeit. xix. p. 182, synonymic notes ; Deutsch.
ent. Zeitschr. p.82, describes Sericoderus chobuuti, sp. n., from
S. France [see 1908].
1901. Reirrer. Deutsch, ent. Zeitschr. p. 70, Orthoperus acariformis,
sp. n., from West ‘Turkestan.
1902. Reirrer. Wien. ent. Zeit. xxi. p. 187, Orthoperus schneidert,
sp. n., from Corsica.
1903. Fauver. Rev. Ent. Franc. (Caen), xxii. pp. 289-291, three new
species of Arthrolips and one of Corylophus from New Cale-
donia *,
Morritt. Ent. News (Philadelphia), xiv. pp. 1385-138, pl. vi.,
metamorphosis of Corylophodes marginicollis.
1908. Reirrer. Wien. ent. Zeit. xxvii. pp. 59-63, describes a num-
ber of forms from E. Africa (Homogrypinus, g. n. near Serico-
derus, and new species of Sactwm, Arthrolips, Sericoderus, Cory-
lophus, and Orthoperus); t. c. p. 198, synonymic notes, and
a Sericoderus chobauti, Reitt. (1900), as a var. of S. reveldiert,
eltt.
Scorr. ‘ Fauna Hawaiiensis,’ iii. pp. 415-8, includes description
of Sacitum angusticolle, sp. n. [omitted by Csiki from his Cata-
logue, 1910).
1909. Reirrer. Bull. Soc. ent. Egypte, i. (1908) p. 40, deser. Serico-
derus ( Anisomeristes) pecirkanus, sp. n., from Egypt.
1910. Buatrcuiry. Bull. Indiana Dept. Geol. i. pp. 501-506, describes
Indiana species.
Csixt. Rovart. Lapok. xvii. p. 28, synonymic notes and new
names ; Coleopt. Catalog. (Junk & Schenkling), part 18, pp. 5—
28, catalogue of the family.
1912. Suarp and Murr. Tr. Ent. Soc. London, p. 507, ¢ genital arma-
ture.
1913. Hetscuxo. Wien, ent. Zeit. xxxil. p. 181, Matthewstella, nom.
nov. for Microwm.
ReitteEr. Deutsche ent. Zeitschr. pp. 653-4, Sericoderistes, gen.
noy. near Sericoderus, with a new species, from Turkestan.
SauLBErG. Ofv. Finsk. Vetensk.-Soc. Férh. (Helsingfors),
vol. lv. 1912-18, Afd. A, no. 8, p. 12, Catoptyx levantinus, sp.n.,
Lebanon.
1914. Broun. New Zealand Institute, Bull. 1, part 3, p. 173, Sacina
curtula, sp. 0., New Zealand.
In the following portion of this paper dates in brackets
after authors’ names refer to the above list.
* In this paper Fauvel also records (p. 289) Arthrolips souverbiet
(Montr.) from New Caledonia. This species was described by Mon-
trousier from that country as one of the Heteromera, being made the
type of a new genus Apelta (Ann. Soc. Linn. Lyon, vol. xi. 1864, p, 124),
aud as such it is included in the Munich Catalogue (vol. vii. p. 1972)
under Tenebrionidee. But Fauvel, as stated above, records it as a Cory-
lophid, giving Apelta as a synonym of Arthrolips. The name souverbeet
does not, however, appear to be mentioned in Csiki's ‘Catalogue of
Oorylophide ’ (1910) either as a yalid species or as a synonym.
Seychelles and Rangoon. 7
Types.—A first set of the material, including the types of
the new genus and of all new species, will be placed in the
British Museum ; a second set will be retained in the Cam-
bridge University Museum.
Sacrum, Leconte.
(PI. I. figs. 1-9.)
Sacium, Leconte, Proc. Ac. Philad. vi. 1852, p. 142.
The material includes four species from the Seychelles, all
quite distinct from each other and from anything in Matthews’s
coliection ; neither do the descriptions of the few species
which I have not seen correspond at all with any of the
Seychelles forms. Reitter (1908) has described five species
from Hast Africa; but after careful study of his descriptions
I conclude that none of my species is identical with any of
his.
Structural Characters.—In examining the Seychelles
collection I have noticed certain structural differences between
the species, of a kind which does not seem to have been
hitherto employed. Thus, among these four species there are
two distinct types of prosternum: (i.) of appreciable length
in front of the ae and furnished with an elevated median
keel (fig. 6) ; (ii.) exceedingly short in front of the coxe and
with no keel ae 3); further details are given in the
specific descriptions. Matthews (Mon. p. 41) writes “ pro-
sterno parvo, inter coxas elevato...,” but makes no state-
ment as to specific differences in its form.
Another category of characters is exhibited by the mouth-
parts. A balsam-preparation was made in order to fix with
certainty the generic position of each species. These prepara-
tions exhibit slight differences in the form and relative pro-
portions of such “parts as the mentum and joints of the palpi,
differences which are briefly mentioned in each description
(Ones, 2.0), 0, oO).
Characters such as these are not necessary for separating
the Seychelles species, which are amply distinct in other
ways. But they are indicated in case they should prove
useful in further studies of this large genus of minute
creatures.
1. Sacium picaultianum, sp. n.
(PI. L. figs. 1-3.)
Oblongo-ovale, supra nitidissimum, fere glabrum; piceo-nigrum,
thoracis margine anteriore testaceo, elytris maculis 4 (in utroque
8 Mr. H. Scott on Corylophide from the
elytro 2) rufo-flavis, corpore subtus rufo-piceo, pedibus rufo-
testaceis, antennarum clavis infuscatis ; supra tote fortiter dense
punctatum, thoracis basi plus minusve regulariter seriatim punc-
tata; metasterno et segmento 1° abdominis subtiliter dense
punctatis.
Long. corp. 1:05-1:25 mm.
Oblong-oval, with elytra nearly parallel-sided, not very
much broader than the thorax at their widest point; upper
surface very shining, with the punctures bearing such exces-
sively short minute hairs (only visible under a compound
microscope) that it may almost be called glabrous. Colour:
thorax pitechy black, with the anterior explanate margin
translucent and testaceous, the testaceous colour extending
back a little on to the dise in two places, one on either side
of the middle line in front ; scutellum black; elytra pitchy
black, with two reddish-yellow marks on each, the front pair
of marks extending from the base to 4 the length or more,
fairly widely separated from the outer margins and at the
suture ; the hind pair only very narrowly separated at the
suture, sometimes quite confluent across it, fairly widely
separated from the apex of the elytron, each mark extending
obliquely forwards from the suture nearly to the outer margin.
In a few examples the spots of the front pair also are nearly
coufluent across the suture ; and insome (possibly immature)
the whole elytra are much paler, almost uniform pitchy
reddish or even testaceous. Underside reddish pitchy, apex
of the abdomen rufescent. Legs reddish testaceous. Clubs
of antenne dark. Thorax and elytra closely and strongly
punctured, the punctures separated by from once to twice
their own diameter, the thorax with a basal series of more
closely placed punctures (very distinct in the figured speci-
men, but less regular in others) ; edytra with lateral margins
reflexed and visible from directly above throughout the
greater part of their length ; sutural stria present, obsolete in
about the anterior 4. Wings dissected out and’ found to be
ample. Metasternum and abdomen finely and closely punc-
tured and finely pubescent; the punctuation more sparse on
the postero-median part of the metasternum.
Prosternum (Pl. I. fig. 3) extremely short, forming in
front of each coxa a bridge so narrow that it can scarcely
be seen in looking directly down on to the under surface ;
there is consequently no room for a median elevated keel
in front of the coxze (contrast Sacium grossinianum, fig. 6).
A balsam-preparation of the mouth-parts shows that the
Seychelles and Ringoon. 9
mentum (fig. 2) is narrow, pointed in front, and the terminal
joints of the labial palpi slightly longer than the second.
Sacium picaultianum approaches three species which I
have seen—S. bifasciatum, Matth. (Madagascar), S. quadri-
maculatum, Matth. (Ceylon), and S. flaviventre, Matth.
(Ceylon), Mon. pp. 53, 54. — S. bifasciatum is slightly
longer in proportion, more tapering behind, much more finely
punctured, with the basal thoracic series much less distinct,
and the light marks on the elytra less sharply defined and
differently arranged. S. quadrimaculatum and S. flaviventre
are both larger and differently shaped in outline, having the
elytra less parallel-sided and broadening out rather more
behind the shoulders ; both have the dise of the thorax dark
red instead of pitchy black and the marks on the elytra much
smaller; moreover, the upper surface is entirely glabrous,
the punctures being devoid of even sueh minute hairs as are
present in S. picaultianum. The latter is quite distinct from
any of the three.
Toe. Seychelles: Silhouette Island, 1908.
Fifteen specimens. Nine were beaten from dead palm-
leaves on the Mare aux Cochons plateau, over 1000 feet,
25. ix. 1908 ; five others are from the same locality, though
how obtained is not recorded ; and one is from the other side
of the island, near Mont Pot-a-eau.
Named after Captain Lazare Picault, who commanded one
of the earliest expe litions to the Seychelles, in 1742 *.
2. Sacium grossinianum, sp. n.
(PI. I. figs. 4-6.)
Oblongo-ovale, supra sat nitidum, subtiliter dense punctatum atque
pubescens, piceo-nigrum, thoracis margine anteriore late testaceo,
elytris fasciis 2 transversis rufo-flavis, in sutura interdum an-
guste interruptis; metasterno et segmento 1° abdominis nigris,
thorace subtus et segmentis posterioribus rufescentibus, pedibus
antennisque rufo-flavis, harum clavis haud nigricantibus.
Long. corp. 1°15 mm.
Oblong-oval, with thorax rather long, its anterior margin
forming a curve that narrows considerably in front, and
with elytra nearly parallel-sided, but considerably broader
than the thorax at their widest point ; upper surface fairly
shining, covered with fine, short, decumbent, pale pubescence.
* For this and other historical particulars, see J. Stanley Gardiner,
“The Seychelles Archipelago,” Geographical Journal, Feb. 1907,
pp. 148-174.
10 Mr. H. Scott on Corylophidee from the
Colour: thorax pitchy black, with front margin rather
broadly reddish testaceous; elytra pitchy black, with two
broad transverse reddish-yellow fasciz, the anterior or both
of which may be narrowly interrupted by darker colour at
the suture, thus almost forming four separate marks; in one
example the scutellum is included in the anterior pale fascia,
in another it is darker; metasternum and first abdominal
segment pitchy black, posterior segments paler ; underside of
thorax, legs, and antennee reddish yellow, clubs of the an-
tenne not black. Zhorawx and elytra closely and very finely
punctured, the punctures twice their own diameter, or rather
more, apart; the thorax has no distinct basal series, but an
impressed line immediately before the base ; sutural stria
present but vanishing in nearly the anterior 4; lateral mar-
gins of elytra reflexed through about 2 the length from the
shoulder, visible from directly above. Wings apparently
ample, but not dissected out. Metasternum and abdomen
finely and closely punctured and pubescent.
Prosternum (PI. 1. fig. 6) much longer than in Sactum
picaultianum, forming in front of each coxa a bridge about
half as broad in an antero-posterior direction as the dimen-
sions of the coxe in the same direction, and having a
sharply elevated median longitudinal keel. A balsam-
preparation of the mouth-parts shows that the mentum
(Pl. I. fig. 5) is broader, not pointed in front, more like
Matthews’s figure (pl. i. D 6).
Several species resemble this in general scheme of colour,
but its pubescent surface distinguishes it in many cases, and
I have seen none very closely similar to it. Among thie
other Seychelles species it is abundantly distinct from
S. picaultianum by its pubescence, its narrower form, finer
punctuation, longer prosternum, by the confluence of the
light marks on the elytra to form transverse fasciz, Kc.
Loc. Seychelles: Silhouette Island.
‘Three examples, fiom the same place as most of the pre-
ceding species, the Mare aux Cochons plateau or near by,
ix. 1908.
Named in memory of Captain Grossin, a member of
Picault’s expedition to the Seychelles in 1742.
3. Sacium roslanianum, sp. n.
(Pl. I. figs. 7 & 8.)
Late ovale, supra nitidissimum, tote glabrum, modice sat dense
punctatum ; piceo-nigrum, margine anteriore thoracis late pallide
testaceo, elytro utroque macula singula media rufo-flava, corpore
Seychelles and Rangoon. 11
subtus piceo, pedibus piceis vel fusco-testaceis, antennarum clayis
fuscis.
Long. corp. 1:0 mm.
Rather shortly and broadly oval, with thorax forming
almost a perfect semicircle (not a narrowing curve), and
elytra considerably wider than the thorax, reaching their
widest point a little before the middle ; shining and entirely
glabrous above. Colour: pitchy black ; front margin of the
thorax broadly pale testaceous and translucent ; each elytron
has a median pale spet, narrowly separated from its neigh-
bour at the suture, more widely separated from the outer
margin ; in one specimen the spots are clear yellow, in others
darker, reddish, and suffused; the black ground-colour is
slightly diluted at the apices of the elytra ; underside pitehy ;
legs pitchy or fusco-testaceous, with paler tibiae; head and
clubs of antenne dark. Thorax. and elytra moderately
strongly and closely punctured ; lateral margins of elytra
reflexed throughout the greater part of their length, visible
from directly above ; sutural stria present, vanishing in the
anterior portion. Wings apparently ample, but not dissected
out. Metasternum and abdomen with remote punctures
bearing fine short hairs, the former nearly impunctate in the
middle.
Prosternum in front of each coxa forming a bridge of con-
siderable breadth in an antero-posterior direction, and having
an elevated median longitudinal keel, 7. e, approaching the
condition found in Sacium grossinianum (cf. fig. 6). The
balsam-preparation of the mouth-parts shows that the mentum
(fig. 8) is rather narrow and bluntly pointed in front, the
apical joints of the /abial palyt shorter than the second (con-
trast S. picaultianum), and the penultimate (third) joints
of the maxillary palpt proportionately longer than in some
other species.
Sacium concinnum, Matth. (Ceylon), S. formosum, Matth.
(Ceylon), and 8. politum, Matth. (Japan) [Mon. pp. 52, 56,
57], all have the same general scheme of colour—each elytron
with a single. pale mark on a dark ground. S. roslanianum
is, however, quite distinct from them all. S. concinnum is
differently shaped, having the elytra very little wider than
the thorax, its punctuation is much closer, and the ligit
marks on the elytra are more longitudinal in direction and
much more widely separated from the outer margins and
from one another. SS. formosum is larger, longer, and
narrower, with thorax forming a longer narrowing curve ;
also its thorax is reddish instead of black, and the pale marks
lie farther back on the elytra and are much more widely
12 Mr. H. Scott on Corylophidee from the
separated at the suture; the punctuation also is finer.
S. politum is much larger, proportionately longer, and nar-
rower, with red thorax; its pale marks are much shorter in
an antero- -posterior direction—2. e., they form a narrow trans-
verse fascia on the elytra.
Loc. Seychelles: Silhouette and Mahé, 1908-9.
Five specimens, from the high forests. In Silhouette two
were found, near Mont Pot-d-eau (ca. 1500 feet), and at
Mare aux Cochons ; in Mahé three, from Cascade Estate at
about 1000 feet, and from the Mare aux Cochons district
at about 1500 feet.
Named after Monsieur du Roslan, under whom an early
expedition visited the Seychelles in 1769.
4. Sacium rochonianum, Sp. ne
(PL. I. fig. 9.)
Minutum, ovale, supra nitidissimum, glaberrimum, omnino impune-
tatum ; thorace rufo-flavo ; elytris piceo-nigris, vel unicoloribus,
vel fascia pallida transversa suffusa, plus minusve distincta,
munitis ; metasterno piceo-nigro, abdomine rufescente, ore an-
tennis pedibus flavis, antennarum clayis haud nigricantibus.
Long. corp. 0°9-1:0 mm.
“Minute, oval, the front margin of the thorax forming an
elliptical curve narrowing slightly in front, sides of the elytra
gradually curved, reaching their widest point a little before
the middle; very shining, absolutely impunctate, and
glabrous above. Colour: thorax unicolorous reddish yellow,
rather paler at the front margin ; scutellum in most examples
reddish yellow, in some darker; elytra pitchy black, diluter
at the apices ; in some specimens practically unicolorous, but
in most there is near the suture just behind the middle a
paler area, which, though very indistinct in some, in other
cases forms a suffused transverse pale fascia; metasternum
pitchy black, underside of thorax yellowish testaceous, of
abdomen reddish; head, antenne, and legs yellow, clubs
of the antenne not darkened. Hlytra with lateral margins
narrowly reflexed throughout most of their length from the
shoulder, these margins visible from above immediately
behind the shoulder and again in the posterior half, but
scarcely visible (or invisible) for a short space just before
the middle; sutural stria present, extending forwards a little
beyond the middle. Wings apparently ample, but not
dissected out. Aletasternum quite smooth, glabrous, and
impunetate in the middle, with scanty very short pubescence
at the sides; abdomen with longer yellowish pubescence.
Seychelles and Rangoon. 13
Prosternum formed rather as in Sacium picaultianum, very
short, forming only a very narrow bridge in front of each
coxa, and sloping steeply upwards (2. e., dorsalwards) in the
middle in front, not forming a median keel. The balsam-
preparation of the mouth-parts shows that the mentum (fig. 9)
is broader than long (contrast S. picaulttanum).
No species which I have seen is closely like this. The
Hawaiian S. angusticolle, Scott (1908, p. 416), resembles it
in its minute size and general colour-scheme—red thorax and
black elytra. But S. angusticolle is distinctly though finely
punctured and pubescent above, and is proportionately longer
and narrower, less oval in outline, and with elytra less
broadened about the middle.
Loc. Seychelles: Silhouette, 1908.
Fifteen examples, all from the high endemic forest above
Mare aux Cochons, well over 1000 feet.
Named after the Abbé Rochon, a member of du Roslan’s
expedition in 1769; he left a written record, and his name
has been given to a river in Mahé.
ARTHROLIPS, Wollaston.
The material includes three species—A. flavicollis, Matth.,
hitherto known from Java, an example of which is now
recorded from Rangoon; A. insule-longe, sp. n., from the
Seychelles ; and an undetermined species from the Seychelles.
Since the appearance of Matthews’s Monograph, Fauvel
(1903) has described three new species from New Caledonia,
and Reitter (1908) two new species from Hast Africa. But
those before me do not appear to be identical with any of
these.
5. Arthrolips flavicollis, Matthews.
Arthrolips flavicollis, Matthews, Ann. & Mag. Nat, Hist. (5) vol. xix.
1887, p. 107 ; Mon. Corylophide, p. 92.
One example, agreeing closely with the type.
Loc. Rangoon; from nest of Munia striuta, 9, x. 1911
(Dr. H. H. Marshall). Previously recorded from Java.
6. Arthrolips insule-longe, sp. n.
CPIAT, fos 10.- BL.)
Sat breviter ovalis, convexus, nitidus, castaneus, fere unicolor, sed
elytris ad latera et antice ad suturam indistincte infuscatis, pedi-
bus antennisque castaneis, harum clavis haud_ nigricantibus ;
corpore supra subtusque dense punctato, pallide pubescente.
Long. corp. 1:15-1°25 mm,
14 Mr. H. Scott on Corylophide from the
Rather shortly oval, more convex than several of its con-
geners, shining, castaneous, almost unicolorous above and
beneath, but with the front margin of the thorax paler and a
dark mark on its dise where the head shows through the
chitin, and with indistinct dark areas along the sides of the
elytra and near the front part of the suture, the latter forming
a median dark mark common to the two elytra; legs and
antennee castaneous, the latter with the clubs not darker ;
body above and beneath covered with fine pale yellowish
pubescence. Thorax with base almost straight, only very
slightly sinuate on either side of the scutellum, with surface
finely punctured, the punctures about twice their own diameter
apart. Seutellum finely punctured. /ytra about as long as
their combined breadth, with sutural stria indistinet (not indi-
eated in fig. 10 and in some positions hardly visible) and
obsolete in the anterior 4, more strongly punctured than the
thorax, punctures about twice their own diameter apart;
reflexion of lateral margins very slight, scarcely noticeable
from above. Wings apparently ample, but not dissected out.
Ventral surface closely punctured, except the middle of the
metasternum, which is almost impunetate.
It is not easy to describe the differences separating this
form from others. It is not identical with any species I have
seen. The following four are selected from Matthews’s
collection for comparison, as they seem nearest to it. A. testu-
dinalis, Woll, (Madeira), is larger, less convex, more parallel-
sided, with the dark areas at the sides and suture of the
elytra contrasting much more strongly with the paler areas
between, and the elytral punctures very much closer.
A. croceus, Matth. (Siam), is narrower, much less convex,
more parallel-sided, and muck paler and yellower; in punctua-
tion it is not unlike A. ¢nsule-longe. ‘The same remarks
apply very nearly to A. senegalensis, Matth. A. westwood,
Matth. (Ceylon), is larger, proportionately longer, less convex,
and generally lighter in colour, though with the darker areas
on the elytra much the same as those of A. ¢nsule-longe@ ; its
antenne are much lighter coloured, being bright yellow; in
punctuation it is not far removed from <A. insule-longe.
The Jatter differs from all these four species in its shorter,
more convex, less parallel-sided form, as well as in the other
ways mentioned in each separate case,
Reitter (1908, p. 61) has described a species—A. centri-
maculatus, from East Africa—which seems to resemble
A. insule-longe in many respects; but without seeing a
specimen it is hard to say exactly how the two forms are
related. A, centrimaculatus is described as “ breviter ovalis,”’
Seychelles and Rangoon. 15
but as “levissime convexus”? and “dilute fulvus,” whereas
A. insule-longe is more convex than several of its congeners
and dark castaneous. The dark areas on the elytra of
A. centrimaculatus appear to be in the same positions as those
of A. insule-longe.
Loc. Seychelles: Long Island, a small coconut-planted
islet close to Mahé, vii. 1908. Hight specimens, obtained by
beating (probably from coconut-trees).
7. Arthrolips sp.
A single specimen, in bad condition, with one elytron
broken. So far as can be seen, the form is rather depressed
and suboblong—that is, more nearly parallel-sided than in
some allied species. Shining, thorax and elytra pitchy black,
the thorax paler (dirty ferruginous) in front, and the apices
of the elytra, where the light shows through, appearing
pitchy ferruginous. Underside of thorax ferruginous, meta-
sternum and first abdominal segment pitchy, hind margins
of abdominal segments testaceous. Legs ferruginous ; clubs
of antenne not black. Body above covered with fine pale
pubescence, much worn in the unique example. Thorax
very finely and subobsoletely punctured. Scutellum and
elytra with stronger, larger punctures, about their own
diameter apart; sutural stria very fine and close to the
-suture, but distinguishable through almost the whole length
of the elytron excepting right at the base. Wings present,
but not examined. Ventra//y, metasternum and first abdo-
minal segment finely and rather closely punctured, the punc-
tuation reticulate towards the sides; the pale pubescence is
rather dense, especially towards the sides of the sternum and
hind margins of the abdominal segments,
Length 1:0 mm.
As the specimen is unique and in bad condition, I have
not named it, though it is not identical with any exaniples
J have seen. A, odlongus, Matth. (Japan), has the same
shape and colour, but is much larger and differently punc-
tured, its thoracic punctures being stronger, while conversely
the elytral punctures are finer and more remote.
Loc. Seychelles: Silhouette, from Mare aux Cochons or
the forest near by, over 1000 feet, 1908.
| Merroperus, Matthews.
Meioderus, Matthews, Mon. Corylophide, p. 102.
This genus was erected to include a single species—J/. niti-
dus, Matth., from Japan,—till now its only known repre-
16 Mr. H. Scott on Corylophide from the
sentative. The new form described below agrees closely with
M. nitidus in generic characters—in general shape, form of
antenne, mouth-parts, sterna, tarsi, &c.,—but is quite distinct
in specific characters.
8. Meioderus quinssyanus, sp. n.
(PI. IL. fig. 12.)
Sat late ovalis, supra fortiter nitidus, omnino glaber ; prothorace
unicolore, rufo, scutello elytrisque unicoloribus, piceis, corpore
subtus fusco-testaceo, pedibus antennisque testaceis, harum clavis -
haud nigricantibus; prothorace fere impunctato, elytris subtiliter
remote punctatis, sine stria suturali.
Long. corp. ca. 1:1 mm.
Rather broadly oval, moderately convex, very shining,
and quite glabrous above. Colour: prothorax unicolorous
reddish, the colour broadly diluted at the translucent front
margin, scutellum and elytra unicolorous pitchy, underside
brownish testaceous, legs and antennz testaceous, clubs of
antenne not blackened. Thorax rather short, its front mar-
gin forming a wide curve; for ordinary purposes it may be
called impunctate, though under a very high power a few
very remote and exceedingly fine punctures are visible, as
indicated in fig. 12. Elytra gradually curved, with lateral
margin narrowly reflexed, though when viewed from directly
above this is generally visible only in front, as shown in
fig. 12 ; punctures fine, remote, shallow, slightly elongate ;
sutural stria entirely absent. Wings ample (mounted in
balsam). Metasternum and first abdominal segment glabrous,
the former impunctate in the middle, finely and remotely
punctured at the sides, the latter finely and remotely punc-
tured.
M. nitidus, Matth., is larger, more elongate-ovate in out-
line, with thorax much darker; the elytra are much deeper
black, their punctuation is, if anything, a little stronger, and
a sutural stria is discernible in the posterior part; the ventral
surface is much blacker and the metasternum more closely
punctured at thesides. When the ventral surfaces of M. nitz-
dus and M. quinssyanus are viewed side by side the greater
relative breadth ot M. guinssyanus is apparent, and the cox
of its middle and posterior pairs of legs look even more
widely distant, inter se, in spite of its smaller actual size.
Loc. Seychelles: Silhouette, viii.—ix. 1908.
Four examples, one from near Mont Pot-a-eau, at about
1500 feet, three from Mare aux Coclhions, about 1000 feet.
This species is named after Monsieur Le Queau de Quinssy,
Seychelles and Rangoon. i
last of the French Governors of the Seychelles, who served
the Monarchy, the Republic, the Empire, and, finally, the
British Government.
SERICODERUS, Stephens.
(PL. Ti. tiem, 13—F7.)
Subgenus ANISOMERISTES (Matthews),
Anisomeristes, Matthews, Ent. Mo, Mag. xxii, 1886, p. 225; Mon.
Corylophide, p. 108.
Sericoderus, pars, Reitter,
Anisomeristes, treated by Reitter, and here, as a subgenus
of Sertcoderus, is separated from true Sericoderus by having
11-jointed instead of 10-jointed antenne. Otherwise the
species of the two subgenera are closely alike, and it is’
impossible without examination of the antennze to decide in
which of them any particular form should be placed.
The difference is caused by the fusion of two joints—joint 3
and the succeeding one—in Sericoderus, s. str. But in some
species at least of this subgenus there is a fine transverse
line on the third joint, showing where the division would be
if it were present. Fig. 17, made from a balsam-preparation,
shows the antenna of a British specimen in the Crotch Col-
lection placed as S. lateralis; fig. 17a shows the elongated
third joint more highly magnified, and it is clear, both from
the shape of the joint and the presence of the transverse line,
that it is made up of two joints fused. Figs. 16, 16a illus-
trate the antenna of S. (A.) pubipennis, Sharp (Hawaiian
Islands), and figs. 15, 15a give that of S. (A.) seychellensis,
sp.n. In pubspennis the separation of the joints is complete,
but not so marked as in seychellensis ; in pulbipennis the two
joints fit together very closely, while in seychellensis the
distal one is distinctly narrowed at its base. ‘he condition in
S. (A.) pubipennis, therefore, seems to be transitional be-
tween that in S. (A.) seychellensis and that in S. (s. str.) late-
ralis. The antenne also exhibit other differences in length
and in the proportions of the joints inter se. But appear-
ances are sometimes deceptive, and much depends on the
exact position in which the antenna is lying in the balsam.
In many descriptions of Sericoderus spp. no mention is made
of the antenne, and the subgeneric position of some species
is not satisfactorily established. Owing to this inadequacy
of descriptions, it is hard to say exactly how certain described
species are related to the Seychelles form. I have named the
latter S. (A.) seychellensis, though it may possibly prove to
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 2
18 Mr. H. Scott on Gorylophide from the
be identical with some described species which I have not
seen,
Condition of hind wings: see ante (p. 4) and below.
9. Sericoderus (Anisomerstes) seychellensis, sp. n.
(PI. IL. figs. 13-15.)
Obconicus, nitidus, unicolor flavo-testaceus, pedibus antennisque
? . 2 . . . Y . P
flavescentibus, harum clavis haud nigricautibus, sat longe aureo-
. 7.* 5 - 2, 2 *
‘pilosus ; prothorace “stbtiliter punctato, inter punctos levi;
elytris fortius punctatis, inter punctos parum asperatis ; antennis
curtis.
Long. corp. 0°75+1:0 mm.
Obconic, of the form characteristic of Svricoderus—that 1s,
with thorax broader than elytra and produced at the hind
angles, and with. elytra narrowing gradually from the base
backwards, subtruncate at the apices, and with sides straight,
not curved. 8, (A.) seychellensis is narrower in proportion
than some of its congeners. Itis shining, unicolorous yellow-
testaceous, with legs and antenne yellowish, the clubs of the
latter not (or only very slightly) darkened. Body covered
above and below with golden pubescence, rather coarser and
longer than, and not quite so close as, in some species. Thorax
smooth, very finely punctured ; elytra rougher, with coarser
punctuation, which extends right to the base. Wings con-
siderably longer than elytra (mounted in balsam).
Of all the forms which I have seen, the Hawaiian S. (A.)
pubipennis, Sharp*, is nearest to S. (A.) seychellensis, but it
is larger and has the pubescence and punctuation ¢ denser.
Tt also differs in the form of the antennal joints (figs..15, 15 a).
In seychellensis the antenne are short, less than 14 times the
breadth of the head, while in pubipennis they measure over
14 times the breadth of the head. In seychellensis joint 2
is short and conspicuously broad in proportion, 3 and 4 are
short and transverse, and the division between them is con-
spicuous, 4 being narrowed at its base, 5 is very little broader
than long, 6 much more PRD EN OT, 7 conspicuously larger
than 8, and the club-joints are short, 9 and 10 both being
broader than long. In pubipennis (figs. 16, 16 a) joint 2 is
proportionately much longer, 4 is differently shaped and
much less narrowed at its base, 9 and 10 are longer, being
about as long as broad. Perhaps characters of a more defi-
nite nature than some of those hitherto used may be found in
* Ty. Dublin Soe. iii. 1885, p. 128; Matthews, Mon. Conylopbide,
p- 121; Scott, ‘Fauna Lawaiiensis,’ iii. p. 417 (1908).
Seychelles and Rangoon. 19
the antenne to distinguish a number of forms superficially
much alike.
Among species which I have not seen, S. etchelbaumi,
Reitter (1908, p. 62, H. Africa), seems to resemble S, (A.)
seychellensis in some respects, but to differ (as, according to
Reitter, 1. ¢., does also the Australian S. pallidulus, Reitter)
in having the punctuation of the elytra obsolete towards the
base; also eichvlbaumi and pallidulus presumably belong to
the subgenus Sericoderus, s. str., though this is not actually
stated. Certain forms have been described from Australta
by Lea* and from New Zealand by Brount, but it is
impossible to say exactly how they are related to 8. (A,)
seychellensis. S. (A.) pecirkanus, Reitter (1908), from
Egypt, is, according to the description, different in shape,
colour, and nature of the pubescence.
Loc. Seychelles: Silhouette and Mahé, 1903-9.
Over fifty specimens, varying considerably in size. In
Silhouette several were swept from a grassy clearing at over
1000 feet, 30. vii. 1908, and a large number were beaten
all together from one place on the edge of the forest at Mare
aux Cochons, over 1000 feet, in the late afternoon of 18. ix.
1908; others were found in various localities both in the
high forests and at lower elevations, In Mahé examples
were taken in the high forest of Morne Blanc, on Cascade
Estate, &c.
DAUBANIA, gen, nov.
(Pl. 1. fig. 18; PI. IIL. figs. 19, 21-24.)
Antenne (ut in Oligarthro) 8-articulatie, sed ab eis Oligarthri in
forma articulorum differentes. Caput sub pronoto omnino ob-
tectum. Genus in forma mandibulorum, maxillarum, labu,
Corylopho affinis, sed ab hoc genere in numero articulorum
-antennarum differt.
Form (fig. 18) oval, narrowed behind, moderately convex,
glabrous above. Head entirely concealed beneath pronotum.
Antenne (fig. 19) 8-jointed ; joint 1 long, thickened, curved
towards base; 2 pyriform, over twice as long as broad ;
8 slender at base, a little longer than broad ; 4 small, a little
broader than long; 5 may be reckoned as part of the club, it
and 6 are about as long as broad; 7 is rather broader than
long; 8 is longer than broad and tapers to a blunt apex.
Labrum (fig. 21) transversely oblong, anterior angles
rounded, anterior margin slightly bisinuate. Mandibles
* Proc. Linn. Soc. New South Wales, vol. x. p. 309 (1895).
+ Man. New Zealand Cgl. part 5, p. 1072 (1893).
20 Mr. H. Scott on Corylophidee from the
(fig. 22) armed on the inner margin with a comb of long fine
teeth, becoming gradually shorter towards the base. Maaille
(fig. 23) with the lobe finely setose; mazillary palp with
joint 2 large, obliquely truncate at apex, greatly produced
and rounded at the outer apical angle, which bears six long
slender laminate processes (cf. Corylophus, Matth. Mon.
pl. iv. fig. D5), each of which becomes gradually broader
from the base outwards, then tapers to a sharp apex ; the
outermost one is much the longest and is curved, the others
become gradually shorter inwards, the innermost ones being
almost the same length; joint 3 very short, transverse ;
joint 4 a little broader than long, produced at the inner apical
angle, rounded off at the outer angle, bearing short hairs on
the almost truncate apex. Labium (fig. 24), so far as can be
discerned, shaped like a spear-head; ligula very large and
broadly spatulate, truncate at apex, narrowed at base; lalial
palpi short and broad ; joints 2 and 3 both broader than long,
2 shaped Jike an asymmetrical cup produced on the outer
side, 3 a little narrower at its base than the apex of 2, its
truncate apex shortly setose. Prothorax semicircular, ante-
rior margin explanate, base bisinuate, hind angles produced.
Scutellum triangular, broader than long, apex blunt. Elytra
very slightly broader behind the shoulders than the base of
the thorax, gradually narrowing behind ; outer margins not
much curved, explanate for about 2 of their length, the
explanate margin disappearing in the posterior } ; posterior
outer angles broadly rounded off, posterior inner angles
slightly rounded ; a fine sutural stria is present, but vanishes
in the anterior 4 of the elytra. Wangs ample (mounted in
balsam). Pygidium rounded, projecting a little beyond the
elytra. Middle cove moderately, hind cove widely, distant.
Type of the genus, Daubania seychellarum, sp. n.
The only other known genus of Corylophide with 8-jointed
antenne is Oligarthrum, Matthews (Mon. p. 127, pl. iv.
fig. C), established for a single species, O. waterhouse/,
Matth., described from a unique example from Chili. In
Oligarthrum, however, antennal joints 2-5 differ absolutely
in actual form and relative proportions from the corresponding
ones In Vaubania, as will be seen by comparing figs. 19 and
20, the latter of which is copied from Matthews’s Monograph ;
so that, unless many intermediate gradations come to light,
the two insects can hardly be classed in one genus.
The mouth-parts of Ol/garthrum have not been dissected,
but Matthews states that, so far as he could see, they re-
sembled those of Corylophus. This resemblance is also
maiked in Daubania, as will be seen by comparing my
Seychelles and Rangoon. 21
figures of the latter with Matthews’s illustrations of Cory-
lophus.
Daubania is dedicated to Monsieur and Madame Edouard
Dauban, owners of the island of Silhouette, Seychelles.
10. Daubania seychellarum, sp. n.
Cel. 1. fe 13: Pin iin. figs, 19, 21-24. )
Nitida, supra glabra, prothorace rufo, elytris piceo-nigris ad apicem
parum dilutioribus, ore antennis pedibus rufo-testaceis, anten-
narum clavis infuscatis; prothorace subtiliter obsolete punctato,
elytris sat dense strigoso-punctatis.
Long. corp. 0°38 mm.
With the characters of the genus. Colour: thorax red,
anterior margin paler, translucent; scutellum and elytra
pitehy black, the latter diluter towards the apex ; underside
reddish brown; legs, mouth, and antenna reddish testaceous,
the antenne with clubs infuscate. Sculpture &c.: disc of
thorax finely and obsoletely punctured; elytra closely punc-
tured, punctures separated by more than their own diameter,
produced into channels or striz, the general direction of
which is longitudinal, though near the suture they become
oblique ; though quite distinct, these stria are not very deep,
aud under high lights sometimes only the actual punctures
are visible. Pygidium finely pubescent. Metasternum with
surface finely alutaceous at the sides, smooth in the middle,
and with punctuation and pubescence very scanty ; in one
specimen examined it is bare of pubescence in the middle,
the first abdominal segment is also nearly bare and has a
median longitudinal depression; in another example this
depression is absent, the segiwent is more pubescent, and the
metasternum has some scanty pubescence in the middle in
front. These differences possibly may be in part sexual
(cf. Rhypobius, p. 26). The other ventral segments are finely
punctured and pubescent.
This species is quite distinct in general appearance fiom all
other Seychelles Corylophide by its minute size, strigosely
punctured elytra, &. No species of any genus in Matthews’s
collection superficially resembles it. Oligarthrum water-
house is quite different, being larger, unicolorous blackish,
with hind angles of thorax less produced and elytral punc-
tures not drawn out into striole.
Loe. Seychelles: Silhouette, Mahé, Praslin Islands, 1508-9.
Fourteen examples: in Silhouette, collected at Mare aux
Cochons plateau or from the forest near by, over 1000 feet ;
22 Mr. H. Scott on Corylophide from the
in Mahé, from country above Port Glaud, 500-1000 feet,
and from the forest on Cascade Estate, between 800 and
2000 feet ; Praslin, Cétes d’Or Estate.
._ Lewistum, Matthews.
(Pl. ILI. fies. 25-28, 30; Pl. IV. figs. 31, 32, 34, 35.)
Lewisium, Matthews, Mon. Corylophide, 1899, p. 164, pl. v. fig. A.
Lewisium was established for two species—L. ceylonicum,
Matth. (op. cit. p. 166), and L. japonicum, Matth. (op. cit.
p. 167), and no further representative of the genus has since
been described. My material contains a long series of a
species’from the Seychelles, which is referred “to Lewisium
on account of its very close general resemblance to L. cey-
lonicum, but which in the form of its antenne and mouth-
parts differs from that species and in some ways more closely
resembles Catoptyx bowringt, Matth. (Java), the type of
the genus Catoptyx*. The Seychelles form Ch seychellea-
num, sp. n.) thus seems in some respects intermediate between
the types of Lewistwm and Catoptyx, and an examination of
the actual parts in L. ceylonicum and L. seychelleanum, and
comparison with Matthews’s figures of Catoptyx renders one
rather doubtful whether the differences between Lewistum and
Catoptya are more than specific. But one of the chief
diagnostic characters of Catoptyx is that it has the anterior
angles of the pronotum abruptly inflexed and closely fitted to
the sides of the head, and of this there is no trace in L. sey-
chelleanum. Therefore I do not propose to sink Lewistum as
a synonym of the earlier name Catoptye.
Antenne, mouth-paris, &e.—The antenna of L. seychelle-
anum (figs. 25, 25a) has the basal joint much thicker, the
third joint proportionately much longer, than that of L. cey-
lonicum (figs. 26, 26a). This forms a ready means of
distinction in balsam-preparations. The labrum of Ly. sey-
chelleanum (fig. 27) is intermediate between that of Lewisewm
and that of -Catoptyx bowringi as figured by Matthews
(copied in figs. 28, 29), being considerably more tapering
than the former but much less acuminate than the latter.
The mandibles of L. seychelleanum are bifid at the distal
extremity, each of the two apices being armed with two or
three hooks (figs. 30, 80 a}—~. e., rather more complex than
those of Catoptyx bowring?, which, according to Matthews
(pl. vi. fig. B4), have only a single hook at each apex, but
* Catoptyx, Matthews, Ann. & Mag. Nat. Hist. (5) vol. xix, 1887,
p- 111; Mon. Corylophidee, p- 167, pl. vi,-fig, B 1-7,
Seychelles and Rangoon, 23
without the serrations that extend some way down the
mandibles of L. ceylonicum (cf. Matthews, pl. v. fig. A 4).
Maaillary palpi ot L. seychelleanum (fig. 31) with joint 2
much less curved and inflated outwardly, and the apical joint
shorter and blunter, than those of ZL. ceylonteum (fig. 32) ;
mawillary lobes of L. seychelleanum slender, sharply pointed,
with inner edge serrate near the apex [Matthews heures the
lobes in Lewistum as unarmed ; but a balsam-preparation of
the maxilla of L, ceylonicum (fis. 32) shows about six minute
teeth near the apex, though these are scattered on the suiface,
not arranged in a serrate edge as in L. seychelleanum].
Fig. 33, copied from Matthews, shows the maxilla of Cafo-
ptyax bowrengi for comparison. ’ Labial palpi of L. seychelle-
anum (tig. 34) lying Hearty contiguous, not spread apart as
in L. ceylonicum (fig. 33) *; fig. 36, copied from Matthews,
shows the parts in Catoptyw bowringi. Therefore in the
maxillz and labium L. seychelleanwm seems in several points
to resemble Catoptyx bowringi more closely. Tarsi of all
three pairs in L. seychelleanum broadly dilated and bilobed,
the lobes pubescent.
11. Lewisium seychelleanum, sp. n.
(Pl. IIL. figs. 25, 27, 30; PI. IV. figs. 31, 34.)
Late ovale, postice perparum angustatum, valde convexum, niti-
dissimum, supra glabrum; piceo-nigrum, prothoracis margine
antico pallide testaceo et pellucido, disco prothoracis ante scu-
tellum, scutello ipso, elytrorum sutura et marginibus exterioribus
(his anguste) piceo-rufis, antennis pedibusque rufo-testaceis,
antennarum clavis haud nigricantibus; prothorace fere impunc-
tato, elytris dense sat fortiter confuse punctatis. Lewisio ceylonico
simile, sed statura minus, et differt in forma antennarum, man-
dibulorum, &c., que vide supra.
Long. corp. 1:05-1'1 mm.
Broadly oval, slightly narrowed behind, very convex, very
shining, elabrous. Pitchy black, with anterior margin of
the thorax pale testaceous and pellucid, and the middle of the
dise of the thorax before the base, together with the scutellum
and suture of the elytra, lighter—z. e., pitchy reddish ; outer
margins of the elytra also narrowly reddish -[in a few speci-
mens the reddish colour is more extended and the whole body
is a little lighter] ; underside pitchy reddish, centre of meta-
sternum and first abdominal segment darker; legs, mouth,
* Too much reliance must not be placed on this differ rence, Which may
be partly due to greater pressure of the coverslip in one preparation than
in the other.
24 Mr. H. Scott on Corylophidee from the
and antenne reddish tastaceous, clubs of the antenne not
blackened. Thorax and scutellum under a powerful hand-
lens appearing impunctate, but under a compound microscope
the thorax is seen to bear numerous very fine subobsolete
punctures. . Hlytra closely and strongly punctured, punctures
separated by once to twice their own diameter ; sutural stria
not distinguishable. Wings dissected out and found to be
ample. Jdetasternum rather closely and strongly punctured
towards the sides, but with the elevated central part almost
impunctate. Aldumen ventrally clothed with fairly close,
fine, short hairs.
In general appearance closely resembling L. ceylonicum,
Matth., which is, however, distinctly larger. The example
of L. ceylonicum betore me appears a very little less convex,
has scarcely any reddish colour along the suture of the
elytra, the elytra even more strongly punctured, and the
metasternum almost impunctate at the sides as well as slightly
less elevated in the middle. But differences of a more
definite character lie in the form of antenne and mouth-parts,
as stated above.
L. seychelleanum is quite distinct in size and general
appearance from the other previously described species of the
genus—1z. e., L. gaponicum, Matth., and also from Catoptyx
bowringt, Matth. A second species of Catoptyx has been
described recently by Sahlberg (1913)— C. levantinus, from
the Lebanon ; but this is said to have the elytra ‘‘ obsolete
punctata” and the third joint of the antenna as long as broad,
and must be quite different from L. seychelleanum.
Loc. Seychelles: Silhouette, Mahé, Long, Praslin, and
Félicité Islands, 1908-9. Found much more abundantly
than any other species, over 190 specimens being taken ; the
distribution seems fairly general, from sea-level and the
cultivated country up into the endemic forests. In Silhouette
many examples were collected from near Mont Pot-a-eau,
ca. 1500 feet, and from Mare aux Cochons; a number were
swept from long grass; one is recorded as beaten from dead
palm-leaves ; two were found in fallen dry branches con-
taining nests of the ant Pheidole punctulata, Mayr (A. Forel
det.), on the coast near Pointe Htienne, 17. ix. 1908. In
Mahé, generally distributed from the cultivated country up
to elevations of over 1000 feet. In Long Island, a cultivated
islet near Mahé, a specimen was taken from the beach just
above high-water mark.
Seychelles and Rangoon. 25
Ruypostius, Leconte.
Rhypobius, Leconte, Proc. Ac, Philad. vi. 1852, p. 141. :
Moronillus, Jacqu-Duval, Ann. Soc. Ent. France, 1854, Bull. p. 38 ;
Gen. Col. Eur. vol. ii. 1857-59, p. 254.
Nec Gleosoma, Wollaston, Ins. Mader. 1854, p. 480, pl. x. fig. 7.
Rhypobius, founded on the North-American FR. marinus,
Leconte, was originally (but erroneously) described as having
Y-jointed antem.a. AMoroni/lus was erected to contain the
European JW, ruficolis, Duval, and was correctly described
as having the antenne of eleven joints, In 1883 Leconte
and Horn [Classif. Col. N. Amer. (Smithson, Mise. Coll.
xxvi.) p. 113] asserted that Rhypobius and Moronillus are
really the same, and admitted that Leconte had wrongly
stated the number of antennal joints in his original descrip-
tion of Rhypobius. Matthews also followed these writers in
regarding Merenillus as a synonym of Rhypobius (Mon.
Coryloph. p. 173). Ganglbauer, however (Kiaf. Mitteleur.
ili. 1899, p. 283, footnote), was not satisfied that the number
of antennal joints is really the same in the two cases, and
therefore employed the name Moronillus as distinct from
Rhypobius. I have made a balsam-preparation of the an-
tenna of a specimen of L. marinus, Leconte, from Matthews’s
Collection. Itis undoubtedly 11-jointed, and closely resembles
that of 2. aquilinus, sp. n. (fig. 38). - Leconte and Horn and
Matthews were therefore right in regarding the number of
joints as the same in the type-species of Rhypobctus and
Moronillus. The character separating the two disappears,
and Moronillus must be treated as a synonym of Rhypobius.
A preparation of the antenna of the West-Indian 2. brevi-
cornis, Matth., also shows eleven joints.
These remarks, however, do not apply to Gleosoma,
Wollaston. This genus was founded tor Glwosoma velox,
Woll., which was described from a unique example found in
Madeira, but of which other examples, subsequently taken
in Nortli Africa, are also to be seen in the British Museum.
Wollaston described and figured the genus as having 10-
jointed antennze (an assertion which I am glad to be able to
confirm, below). But Duval, in his Gen. Col. Europe, sank
Glaosoma asasynonym of his genus Moronillus. To this
Wollaston replied in his ‘ Coleoptera Atlantidum’ (1865,
pp- 93-5, and footnotes), saying that he had carefully
re-examined the type of G. velox, and was convinced that his
original figure and description were correct, that the antennee
were really 10-jointed, and that the joints differed in form
26 Mr. H. Scott on Corylophide from the
inter se from those of Aforonillus. Nevertheless, Leconte and
Horn and Matthews regarded Glevsoma (like Moronillus) as
a synonym of Fhypobius; but Ganglbauer (/. c.) was not
convinced, and Casey (1900, p. 65) wrote that Gleosoma is
altogether distinct trom Rhypobius. J have examined the
type of G. velox under the highest power applicable to a
carded specimen, and found that the antenna appeared almost
certainly 10-jointed_; but being still not satisfied, I mounted in
balsam the antenna of one of the WNombewean: specimens,
which seem absolutely identical with the type. This antenna
(fig. 39) is 10-jointed, having between the second and the
next large joint one small joint less than in Fhypobius, and,
as stated by Wollaston, the form and proportions of the joints
differ from those of Lthypobius. The three joints (5, 6, 7)
preceding the three club-joints are all much longer in propor-
tion than the corresponding three (6, 7, 8) in RAypobcus, and
the large middle one of the three especially is of a different
shape.
If the number of antennal joints be used as the criterion
for separating the gencra, the matter may be summarized
thus :—
Rhypobius (= Moronillus), antennee 11-jointed.
Gleosoma, autenne 10-jointed.
Secondary Sexual Characters—I do not know of any
reference to these in Rhypobius. But the material before me
includes three specimens of a species, apparently new, two of
which have a marked impression on the metasternuin, while
in the third this is quite absent. In comparing certain other
species with mine, it was seen that some examples have
impressions on the metasternum and sometimes on the first
abdominal segment as well. Having before me two speci-
mens of Lhypobius ruficollis (Duval), one of which has the
sternum impressed while the other has not, I dissected these
aud found that the insect with impressed sternum is g@, while
the otheris 9. I therefore infer that the ventral impressions
are a @ character, though further study is needed to prove
whether they are present in all or only in some species.
Those = which they have so far been observed are :—
(i.) R. rujficollis (Duval), go: a rather faint and narrow
een al impression on the posterior 3 of the metasternum,
and a long narrow impression down the middle of the first
abdominal segment.
(uae, brevicornis, Matth., g: a deep and rather broader
longitudinal impression on the metasternum ; on the first
abdominal segment a very broad and deep impression,
extending the whole length of the segment aud nearly the
-
Seychelles and Rangoon. 27.
whole distance between the hind coxe ; on either side of the
impression the segment is raised into a ridge which bears
rather long pubescence.
(iii.) hk. aquilinus, sp. n., ¢@: a marked longitudinal im-
pression, broadening behind, along the posterior 3% of the
metasternum, the pubescence in the impression being much
closer than on either side of it ; first abdominal segment with
no impression, but with a little median group of hairs.
Condition of hind wings: see ante (p. £), and below.
12. Rhypobius aquilinus, sp. n.
CEL ING Bes 377; Pl, Vi. figs 38.)
Ovalis, postice haud fortiter attenuatus, supra subtusque subtilissime
alutaceus, thorace rufo-flavo, elytris castaneo-brunneis postice ad
suturam interdum rufescentibus, pedibus antennisque flavescenti-
bus, harum clayis haud nigricantibus; thorace impunctato ;
elytris punctis duplicibus sat confertim munitis; metasterno ¢
in medio longitudinaliter valde impresso, segmento 1° abdominis
baud impresso.
Long. corp. 0:85 mm.
Outline shown in fig. 37; the thorax appears a little
shorter than it actually is, owing to its being bent down ;
length of the elytra very nearly equal to their combined
breadth, which is greatest a little before the middle. Body
above shining, glabrous ; ; finely and closely alutaceous above
and beneath. Colour: thorax reddish yellow, elytra dark
castaneous brown, in the type-specimen lighter and more
reddish in the posterior half near the suture ; ventral surface
castaneous brown, antennze and legs yellowish, clubs of the
antenne not blackened. Antenne (fig. 38) a little longer
than the width of the head from eye to eye. Zhorawx nar-
rowly margined at the sides, with base. very shallowly
sinuate on either side of the middle, and hind angles (seen
from the side) slightly less than right angles; surface im-
punetate. Scutellum rounded. £/ytra with lateral margins
narrowly reflexed, but in viewing a specimen from vertically
above the margins are only visible behind the shoulder and
again for a short space behind the middle; sutural stria
quite absent ; surface with fine double punctures, each con-
sisting of two slightly elongated punctures lying close side
by side*; in a transverse direction the double punctures are
about their own diameter apart, but in a longitudinal direc-
tion about twice this distance. Wangs: uo trace of these
* The alutaceous surface and double punctures are characteristic of a
number of other members of the genus.
28 Mr. H. Scott on Corylophide from the
organs can be seen under the partly opened elytra of the
single 9, but actual dissection and search for minute
vestigial wings is prevented by the necessity of preserving
the specimen intact; the two ¢ have ample wings, folded
under the elytra; one of these organs is mounted in balsam,
but I have failed to unfold it completely, so cannot state its
proportions to the elytron accurately ; it is, however, con-
siderably longer than the elytron (see p. 4). Metasternum
3 with a marked median longitudinal impression broadening
behind, on the posterior ? of its length; surface of the meta-
sternum almost impunctate, with pale short hairs, closer in
the impression, very scanty at the sides ; in the 2? the meta-
sternum is convex and glabrous in the middle. Sirst abdo-
minal segment: @, with no impression, but with a median
group of a few short hairs, on either side of which it is bare,
but has a few other hairs near the lateral margins ; 2, no
median group of hairs. ‘The other segments bear scanty pale
pubescence.
This species is quite distinct from any I have seen. The
form most closely resembling it superficially is R.brevicornis,
Matth. (West Indies), A ¢ of this, now before me, is the
same size, but more attenuated behind ; the reticulation of its
thorax is slightly less marked, while its elytral punctures are
a little stronger ; and it differs decidedly in the nature of its
3 ventral impressions (vide supra, p. 26).
Loc. Amirantes Islands. ‘I'hree specimens from Eagle
Island, 1905 (H.M.S. ‘ Sealark’ Expedition).
Named ‘“aquilinus” in allusion to the island of its
discovery.
ORTHOPERUS, Stepliens.
(PI. IV. figs. 40, 41; Pl. V. figs. 42-44.)
The material includes at least two, possibly three, species
of this genus: a new and very distinct form from Rangoon ;
a single g from the Seychelles, referred to a species known
from 8. America and W. Indies ; anda single indeterminable
specimen from Rangoon, possibly the ? of the preceding,
possibly distinct.
Diverging Strie on Metasternum.—I have found in the
literature no mention of diverging striz or lines on the meta-
sternum, curving round behind the middle coxe (fig. 41, 2.) ;
yet they are present in a number of species. ‘They recall
the diverging striw found in a similar position in Acritus
and other Histeridz, but in these there is a second pair of
diverging striz behind the hind coxe on the first abdominal
Seychelles and Rangoon. 29
segment, while in the Ovthoperi there is only the pair on the
metasternum. The species in which I have seen them are :-—
equalis, Sharp, atumarius, Heer, brunnipes, Gyll., coriaceus,
Rey, crotchi, Matth., kluki, Wank., muni, sp. n., ovatus,
Matth. IJ have not examined the other species of the genus
as to whether these striz are present or not.
Secondury Sexual Characters——More than one writer has
noted that the front tibia of some Orthoperus are long and
incurved at the apex. Thus Matthews, in his description of
the genus (Mon. p. 182), ‘* [anterior] tibiz often very long
and much incurved, abruptly incurved at the apex”; and
again, in his descriptions of some of the species, ‘anterior
tibie very long and strongly incurved,” or, contrariwise,
“anterior tibiee nearly straight ”’ (see also his figure, pl. vii.
fig. A 1). But it does not seem to have been stated that this
difference in tle form of the tibig is, in some species at least,
sexual, Thus, in O. muni, sp. n., the front tibiae of some
specimens, which I infer to be ¢, are more incurved towards
the apex, and have a sharp heel or spur at the inner apical
angle (fig. 42) ; while those of other examples, presumably
2, which in all other external characters appear identical
with the preceding, are straighter and have no such heel
(fig. 43). In this case the curvature of the ¢ tibia is not
very marked, but it is much greater in O. minutissimus,
Matth. (fig. 44). Dr. Sharp has pointed out to me the same
kind of sexual difference in the form of the front tibia: in
some of our British Orthoperus. The divergence of the sexes
in this respect is sometimes quite sufficient to be seen with a
hand-lens.
Casey (1908, p. 65) describes for certain North-American
forms a new genus Mutrilza, one of the principal characters
of which is that it has the front tibia more flattened and less
incurved at the apex thanin Orthoperus. It will be necessary
to discriminate between sexual and other differences before
the limits of the two genera are made quite clear.
13. Orthoperus munie, sp. n.
(PI. LV. figs. 40, 41; Pl. V. figs. 42, 43.)
Ovatus, valde convexns, nitidissimus, glaber, piceo-fuscus, pedibus
antennisque testaceis, harum clavis infuscatis; thorace serie
basali punctorum fortium ad latera haud attingente, in medio a
basi magis distante, munito, disco subtilissime ac subobsolete
punctato; elytris sat dense sed subtilissime ac subobsolete punc-
tatis; ¢ tibiis anterioribus ad apicem parum incurvatis, angulo
apicali interiore producto.
Long. corp. 0°7 mm.
30 Mr. H. Scott on Gorylophidee from the
Ovate, very convex, shining, smooth (not at all aluta-
ceous), and quite elabrous above; body above and beneath
and head pitchy fuscous; legs, palpi, and antenne testaceous,
clubs of the latter infuscate. Head impunctate. Thora
with its base sinuate on either side and produced backwards
in the middle, with lateral margins (seen from the side)
slightly sinuate in the middle, hind angles nearly right
angles; with a strong basal series of rather elongate punc-
tures, becoming osolete at the sides, further removed from
the actual base in the middle than at the ends of the series
[it recalls the basal series of some species of Acritus| ; diss
bearing a number of very fine subobsolete punctures, bat in
some lights and positions these are scarcely visible. Hi/ytrau
of nearly the same length as their combined breadth, con-
siderably larger than the ab: lomen, the outline of which is
shown in fig. °40 appearing through the elytra as a dotted
line (perhaps some allowance must be made for shrinkage
of the abdomen) ; lateral margins not visible from directly
above; the elytra have no trace of a sutural stria, and are
finely and rather closely punctate; the punctures under a
high power appear as fine elongate dashes, closer at the
base and suture, and almost obsolete towards the apex (like
those on the thorax, the punctures in some lights and
aspects are difficult to see owing to their shallowness).
Wings ample. Metasternum (fig. 41) very convex, impunc-
tate in the middle, finely punctured at the sides, the diver ging
striae behind the middle cox are punctured and run in a
continuous curve from the anterior to the lateral margins of
the metasternum. Abdomen in several specimens tapering
to a blunt point, first segment almost impunctate, each seg-
ment with a series of very fine short hairs, rather wide apart.
Front tibie of $ (fig. 42) slightly incurved towards the apex,
with the inner apical angle produced into a sharp heel ; in
both g and ¢ (for the latter sex, see fig. 43) the excavation
of the outer margin towards the apex is conspicuous. No
other external sexual distinction is visible.
No species in Matthews’s Collection resembles this at all
closely, and those described since his time seem quite diffe-
rent. O. japonicus, Matth., has a basal thoracic series of
punctures, but they are much finer; it is much larger than
O. munie, has a minutely reticulate surface, and much closer
elytral and thoracic punctuation.
“Loc. Rangoon. Six examples, found in nest of Munia
striata, 9. X. “1911 (Dr, H. H, Marshall),
Seychelles and Rangoon. 31
14. Orthoperus minutissimus, Matthews (?).
(Pl. V. fig. 44.)
Orthoperus minutissimus, Matthews, Mon. Corylophide, 1899, p. 196.
A single g, in bad preservation. Pitchy fuscous, legs
and antenne lighter, shining and quite glabrous above.
Thorax not (or scarcely) punctured, Hlytra finely and sub-
obsoletely punctured, the punctures more than their own
diameter apart. Ventrally the metasternum is impunctate
in the middle, bat its sides and the first abdominal segment
have very fine punctures several times their own diameter
apart. Wings not examined,
So far as can be seen in its bad ,condition, the specimen
agrees in size, colour, and punctuation with an example in
Matthews’s Collection from Grenada, West Indies, placed as
VO. minutissimus *, 'The two agree particularly in the form
of the front tibize, which are sharply incurved at the apex,
the inner apical angle forming a sharp heel. Fig. 44 shows
the right-hand front tibia in the West-Indian specimen.
Loe. Seychelles: Silhouette, from Mare aux Cochons,
1000 feet or more, ix. 1908. O. minutissimus, Matth., is
recorded from South America and West Indies.
15. Orthoperus sp.
Among the material from Rangoon is a single specimen,
pethaps not fully mature, of a very minute species, quite
distinct from O. munie by the absence of the basal thoracic
series of punctures. In size and punctuation of the upper
surface it isnot unlike the example from Silhouette described
above: and referred to O. minutissimus. It is just possible
that it is a 9 of that species, since it probably belongs to the
% sex, the front tibia not being incurved and having no sharp
heel. The metasternum appears quite impunctate, even at
the sides; diverging stria perfectly distinct but not punc-
tured. Determination or further description of this form is
impossible in the absence of more material, Wings not
examined,
. * The name and description of O. minutisstmus are published in square
brackets in Matthews’s Monograph, from his own MS. notes, by P. B.
Mason, editor of the Monograph. Mason gives reasons for thinking that
Matthews probably intended to sink this name as a synonym of O, per-
pusillus, Matth. I have, however, provisionally retained the name
minutissimus, since time has not admitted of an examinationof Matthews’s
material sufficiently close to decide whether minutissimus and perpusillus
are identical or not.
32
On Corylophides from the Seychelles and Rangoon.
Length about 0°7 mm.
Loc. Rangoon: from nest of Munia striata, 9. x. 1911
(Dr. H. H. Marshall).
EXPLANATION OF THE PLATES.
Norse.—The figures of whole insects are approximately, but not exactly,
to scale: they are magnified between 47 and 57 diameters, in most cases
50-53 diameters.
Fig.
Fig.
Fig.
Fug.
Fig.
Fig.
Fig.
Fug.
Fig.
Fg.
Fig.
Fig.
ane
=
KH SOMNM AE wr
18.
ig. 19.
. 20.
wok,
ae
. 23.
. 24,
. 25,
. 26.
ads
2s,
. 29.
. 80.
PLaTe I.
Sacium picaultianum, sp. n.
. Ditto. Mentum.
. Ditto. Underside of prothorax and anterior coxa.
. Sacium grossinianum, sp. 0.
. Ditto. Mentum.
Ditto. Underside of prothorax and anterior coxe.
. Sacium roslanianum, sp. 1.
Ditto. Mentum.
. Sactum rochoniunum, sp.n. Mentum.
. Arthrolips insule-longe, sp. n. Outline.
. Ditto. Punetuation and pubescence of thorax and elytra, to
larger scale.
Daubania seychellarum, gen. et sp. n.
Puate II,
. Metoderus quinssuanus, sp. n.
3. Sericoderus ( Anisomeristes) seychellensis, sp. n. Outline.
. Ditto. Sculpture and pubescence of thorax and elytra, to
larger scile.
. Ditto. Antenna. 15a, joints 5 and 4 more highly magnified.
. Sericoderus (Anixomeristes) pubipennis, Sharp. Antenna, 16a,
cage
joints 3 and 4 more highly magnified.
Sericoderus (s. str.) laterals, Gyll. Antenna. 17 a, joint 3 more
highly magnitied, showing transverse line.
Puate II.
Daubania seychellarum, gen. et sp.n. Antenna.
Oiigarthrum waterhouset, Matthews. Antenna (from Matthews,
Mon. Coryloph. pl. iv. fig. C7).
Daubania seychellarum. Labrum,
Ditto. Mandible.
Ditto. Maxilla.
Ditto. Labium.
Lewisium seychelleanum, sp.n. Antenna. 26 a, joints 3-6 more
highly magnified.
Lewisium ceylonicum, Matthews. Antenna. 26a, joints 3-6
more highly magnified.
Lewisium seychelleanum. Labrum.
Lewisium sp. Labrum (from Matthews, pl. v. fig. A 3).
Catoptyx bowringt, Matthews. Labrum (from Matthews, pl. vi.
fig. B 3).
cae seychelleanum. Mandible. 30a, apex of another
specimen from a different point of view.
Mr. ©. G. Lamb on Exotic Chloropide. 33
Prats LV.
Fig. 31. Lewisium seychelleanum. Maxilla.
Fig. 32. Lewisium ceylonicum. Maxilla.
Fig. 33. Catoptyx bowringt. Maxilla (from Matthews, pl. vi. fig. B 5).
Fig. 34. Lewisium seychelleanum. Labium.
Fig. 35. Lewisium ceylonicum. Labium.
Fig. 36. Catoptyx bowring?,. Labium (from Matthews, pl. vi. fig. B 6).
Fig. 37. Rhypobius aquilinus, sp. u. Outline.
Fig. 40. Orthoperus munie, sp. n.
Fig. 41. Ditto. Metasternum and first abdominal segment, middle and
posterior coxal cavities shaded ; /., diverging metasternal line
or stria.
PLATE V.
Fig. 38. Rhypobius aquilinus, sp.n. Antenna.
ug. 39. Gleosoma velox, Wollaston. Antenna.
Fig. 42. Orthoperus munie, sp. nu. Anterior tibia and tarsus, ¢.
Fig. 43. Ditto. Ditto, 9. ;
Fig. 44. Orthoperus minitissimus, Matthews. ~ Anterior tibia and
tarsus, ¢.
JI.—WNotes on Exotic Chloropide. By C. G. Lams,
M.A., B.Sc., Clare College, Cambridge.
Tue following notes are based on material from two sources.
The larger portion is the collection of Diptera in the Zoolo-
gical Department of Cambridge University, and will be
referred to as ‘‘Cam. Coll.” In 1904 Mr. F. Muir presented
a very large collection of Diptera from Africa to the Cam-
bridge Museum, and his specimens will be marked “ F. M.”
In addition, the Museum was indebted to Dr. G. A. K.
Marshall for many other specimens from the same region,
and there have been various other small accessory collections
incorporated from time to time. The other portion consists
of specimens kindly submitted to the author by Dr. G. A. K.
Marshall—they are part of the extensive collection being
formed by the Imperial Bureau of Entomology ; this will
be referred to as “ Bur. Coll.”
All the insects listed and described in the paper will be
deposited in the British Museum, and hence no indication
of the situation of the type-specimens will be given after
the descriptions; they will all be in the British Museum.
The task of dealing with this family is enormously
lightened and simplified by the valuable and complete
monographs of Th. Becker, which bring the information
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 3
34 Mr. C. G. Lamb on Exotic Chloropide.
available up to the dates of publication of the same, and
hence save much labour in searching out old records.
These monographs are :—
I. Theil. Palearctic Region.
‘ Archivum Zoologicum,’ i, 1910.
Il. Theil. Ethiopic Region.
Ann. Mus. Nat. Hung. vii. 1910.
TII. Theil. Indo-Australian Region.
Ann. Mus. Nat. Hung. ix. 1911.
IV. Theil. Nearctic Region, Neotropical Region, and Addendum.
Ann, Mus. Nat. Hung. x. 1912.
The last brings the list of known species in all the regions
up to date ; it also contains a discussion of Enderlein’s new
genera (Sitz. d. Gesell. Nat. Freunde, 1911), and clears up
many points in that paper which at one time seemed likely
to throw the classification into confusion.
These monographs will be shortly referred to by the
numbers I., II., I/I., IV. after Becker’s name.
As is so often the case, a considerable number of single
specimens occur in both collections. Where the characters
are quite unmistakable and striking, these single specimens
have been described as the types of new species. When the
specimen agrees with fair accuracy with any published
description, it has been thought best to place the insect
under the existing name; but in general it will be found
that this fact is referred to, and any differences recorded.
The Chloropide form a very protean family and include
ereat numbers of genera that run fairly closely into one
another. It might be said that almost every positive
character which limits the family may be separately absent
in some genera—in fact, the allocation of an insect to the
family is in many cases practically due to a “trained eye,”
and cannot be logically justified by the limits of the defini-
tions of the family. This is possibly more true of this
family than of any of the other Acalyptrate groups.
It naturally results that the generic limitations follow the
same tendency, and that the original limits of a genus, as
set by its founder, have to be transgressed, so that finally
the “genus” sometimes bears little resemblance to the
limited form originally prescribed. A good example of this
is to compare Gaurax as founded by Leéw with Becker’s
latest concept of the species forming that assemblage.
Mr. C. G. Lamb on Exotic Chloropide. 35
Such a sequence of events is, from the nature of the case,
inevitable, though it leads to much difficulty both in tracing
species and in assigning genera. The fact is that in some
groups of the Oscininee there is no natural line or lines of
demarcation ; even the known forms merge into one another
and share characters that should belong to different genera
as originally defined; and when the world forms are “really
adequately studied there can be no doubt but that this
tendency will be increased.
It will follow that it is quite possible that the author may
have assigned species to definite genera which, in the opinion
of more experienced students, should be placed elsewhere.
For this reason the descriptions are often made a little
fuller than would be necessary to enable one to discriminate
between the species of a sharply bounded genus ‘such as
Chyliza. For the same reason it will be found that he has
been compelled to place species in genera whose specification
does not exactly meet the case. A good example is Lagaro-
ceras anomalum; if the head were removed it would he
impossible to distinguish this species from one of the
described forms of Becker’s L. megalops ; but the antenna
and vertical triangle are both considerably at variance with
the forms described as characterizing the genus. Wide
interpretation of generic limits is unavoidable in this family,
for if definite and fixed generic characters were to be adhered
to, the family would mainly consist of monotypic genera.
The author hopes to be able to continue with some other
of the Acalyptrate families if time and opportunity permit.
Note.—To save space certain abbreviations will be used.
When describing the head the word “ triangle” will refer to
the fronto-vertical macula usually found there, though its
shape varies greatly. The antennal joints will be referred
to by number only—thus “38rd” will mean third joint of
the antenna. In the case of the thorax the word “callus”
will refer to the front thoracic callus unless qualitied. Inthe
wings the veins will be referred to by the old system of
numbers as being more convenient and simple in this case ;
thus, “ 2nd” will mean the second long vein. Similarly, the
pra segments will be referred to by the numbers ee the
long veins that end there—thus “2 to 8” means costal
distance between ends of second and third long veius
measured on costa.
36 Mr. C. G. Lamb on Exotic Chloropide.
CHLOROPINA.
PacuyLopnus, Loéw.
The following species were in the collections :—
P. lugens, Loéw. Cam. Coll., Durban (fF. M.).
P. splendidus, Ad. Cam. Coll., Durban (Ff. M.).
P. proximus, Ad. Cam. Coll., Durban (7. M.).
P. fossulatus, Ad. Cam. Coll., Durban (Ff. .).
P. varipes, Ad.; a very pale-legged form. Cam. Coll.,
Durban (/. M.).
In addition, there were many specimens included in the
ordinary black-vertexed section. To this section belongs
Fig. 1.
to)
Si oe
Fic. 2, Fig. 3.
ete
Fig. 1.— Wing of large form of P. frontalis.
Fig. 2.— Wing of small form of P. frontalis.
Fig. 3.—Wing of P. frontellinus (?).
Loéw’s species P. frontalis ; Becker, in his monograph, only
recognizes as valid that single species in the section with
black triangle, prominent head, and pale femora. Hesinks
as synonyms both Bezzi’s P. tellinii and Speiser’s P. fron-
tellinus. As regards the first, he makes out what is appa-
rently a good case, but gives practically no reasons for the
second. If one studies the fairly long series in the Cam.
Coll., it can be seen at once that there are two quite definite
venations present, shown in figs. 2 and 3. The species with
the cross-veins fairly apart has a dullish frons, with the
triangle little marked, but a raised shining black central
Mr. C. G. Lamb on Exotic Chloropide. . 37
line, and this form agrees quite well with the description of
frontalis. The species with the more approximate veins is a
little smaller and its vertical “triangle” is more shining ;
the raised central line tends to be multiple—in fact, it agrees
very closely indeed with Speiser’s description of frontellinus
(Kilimandjaro Meru Exp., Diptera, x. p. 198). In the Bur.
Coll. are specimens still more robust than the first species,
with venation as in fig. 1 and with rather more glassy wings,
but otherwise practically identical with the first series.
It appears best to consider the first and last sets of speci-
mens as belonging to frontalis, and the other to frontellinus ;
in none of the published descriptions is any figure of the
venation given,
P. frontalis. The large form: Bur. Coll., Durban (LZ. Bevis).
‘The smaller form: Cam. Coll., Durban (/. M.).
P. frontellinus. Cam. Coll., Durban (PF. M.).
Three species in the collection belong to the smaller
section of the genus with a pale vertical triangle. ' Hitherto
there are described but two species in this section (Becker,
II., p. 388). Neither of these species accords with the
specimens in the collection, and hence they must be con-
sidered new.
Pachylophus lituraticrus, sp. n.
This species belongs to Becker’s section with pale legs
and triangle (II. p. 387).
Head (top view) :—Vertex nearly as broad as twice the
visible eye-breadth, somewhat transversely concave on the
front, bright dullish yellow; the triangle is shining orange
without furrows &c., extending not quite to the frons, with
a rounded darkened tip, very slightly concave sides, and with
the vertical base just less than the eye-distance ; the ocellar
spot black. In profile the angle between the vertical and
facial tangent-planes is a little less than 90°, the frons being
just visibly prominent. The pale yellow cheeks are about 3
the depth of the 3rd joint; the latter is quite normal, yellow
except for the part below aristal insertion; the latter is
black and of usual form. Palpiyellow. Hind head orange;
a darkening behind the ocellar spot, and a pale spot each
side just at the top angle of the eye, on which the tiny
vertical bristles stand.
Thorax: dorsum elegantly striped; along the middle
is a broad stripe of reddish brown bordered by pale narrow
.
38 Mr. C. G. Lamb on Exotic Chioropide.
stripes, which are less covered with the tiny bristles clothing
the rest of the dorsum—hence these pale stripes are
made more conspicuous ; beyond these down to the noto-
pleural suture the dorsum is again reddish brown, though
less intense; callus with a black central spot surrounded
by yellow; pleura orange, shiming. Scutellum with its
middle third oceupied by a bright yellow longitudinal stripe,
bounded by black, though the extreme hind angles are pale ;
terminal crossed bristles just at the end of the black
stripes ; the surface is very faintly and sparsely striate.
The venation is exceptional; the cross-veins are much
approximated, being separated by a distance rather less than
13 times the length of the hind cross-vein. The discal cross-
vein is a little beyond the costal ending of the 1st ; wings
clear, with pale orange veins. Halteres white.
Legs all yellow except for an infuscation on the front
tibia and tarsus and on the Jast joints of the other tarsi. A
very striking and constant character is a darkened ‘ brand ”
ou the back of the hind tibia; this brand occupies about 4
the length of the tibia. Similar structures can be just seen
on the legs of other species, but are not coloured in them ;
they also occur in other Chloropid genera.
The abdomen is the same colour as the thorax, the margins
narrowly paler, and has a well-marked interrupted darkened
middle line.
The intensity of the reddish colour of the insect varies to
a fair extent, .
Length (excluding antennz) nearly 3 mm.
A long series in Cam. Coll., Durban (Ff. M.).
Note.—It is just possible that this species is the same as
Becker’s P. contractus (II. p. 893) ; but it is unlikely that
so careful an observer should not mention the “ brand” or
the relation of the cross-veins.
Var.—There is a single specimen with a slightly paler
triangle and no visible dark brand. This is m the Bur,
Coil., Manganallur, Tangore. Although the localities are
so far apart, the insect is not specifically separable,
STeLeocervs, Beck.
S. lepidopus, Beck. Cam. Coll,, from Chirinda Forest
(G44 i. 1.,),
Mr. C. G. Lamb on Eebtie Chloropidee. 39
Steleocerus nigricornis, sp. 0.
This species is next to Becker’s S. longicollis (II. p. 401),
but differs as follows :—
The halteres are grey, not white; the jowls are larger,
about 4 the depth of the third antennal joint ; the tongue is
pale, not black; the antennz are all deep black, not red ;
legs a little paler; wings more normal, with rounded anal
angle.
Size 55 mm.
Cam. Coll., Durban (F. M.).
S. ensifer (?), Thoms.
A single specimen agrees fairly with Thomson’s descrip-
tion (Bug. Resa, p. 605), but it is possible that we have
another species here. The legs are quite pale; the frontal
triangle is not all yellow, but is very much suffused with
shining brown, which does not, however, entirely cover the
triangle, but occupies the base and shades off forward.
One cannot be sure of the identification from this single
specimen.
Bur. Coll., Mysore.
There is an immature specimen in Bur. Coll. from Coim-
batore, Madras, which is near S. formosus, Beck.
Stelevcerus quadrivitiatus, sp. n.
From the Chirinda Forest, S. Africa, we have a few
specimens of a species of the ensifer-tenellus group.
Head (top view):—Frons yellow and dull, the triangle
equilateral, with nearly straight sides; basally it practically
touches the eyes and extends by a sharp point right to the
front ; itis suffused with shining brown, which leaves narrow
yellow side-lines and broader boundaries on the hind head,
where the black part of the triangle’s base extends as a
broad stripe down the hind head. Side view: outline fairly
circular, the frontal and facial tangent-planes making about
90°, and the hind jowls large ; eyes oval, oblique forwards,
with narrow lower jowls less than half the width of 3rd ; all
the side is whitish yellow. Antenne rather large, the 3rd
projecting backwards a little, so as to be a longish oval with
axis parallel to body-axis ; it is yellow, but darkened dorsally ;
2nd yellow, arista normal. Face, palpi, &c., all pale yellow.
Though there are but few specimens, the thorax varies
somewhat in amount of darkening ; dorsum with a broad
40 Mr. C. G. Lamb on Exotic Chloropide.
black central stripe, sometimes getting browner behind, and
extending forward right on to the prothorax; each side is a
grey poliinated line of ochreous tone, which is moderately
distinct till just before the scutellum, where it suddenly
becomes very marked and forms an elongate spot at each
side of scutellar base; similarly in front, just at level of
calli, it again forms bright long spots; beyond these grey
lines the dorsum is as the mid-line, but is more darkened in
front of the cross-suture, in one case quite black there.
Calli shining, rather orange. Pleura all somewhat shining
orange, with brownish boundaries to the sclerites ; the black
spot over middle coxa may or may not be present. Scu-
tellum darkened orange, in one case paler in centre ; terminal
bristles long and crossed, and a few smaller marginal ones.
Wings clear with brown veins, 2nd ending about 2 down
costa between 1 and 3. Halteres white, with orange stalks.
Legs orange, with front tibia and tarsus a very little
darkened.
Abdomen yellowish at sides, the dorsum forming a broad
darkened continuous stripe.
Size (ex. antenne) just under 2 mm.
Cam. Coll., Chirinda Forest, 8.A. (G. A. K. W.).
The second species is represented by but a single specimen,
but it is very distinct from all the others.
Steleocerus flavipes, sp. 0. -
Head (top view) :—Abont 14 times as long as broad ;
from vertex to the sightly prominent frons it is all bright
yellow ; the triangle is very shining, especially along its
concave side boundaries ; these are slightly depressed, and
the hair-lines on them are exceptionally well marked ; the ~
sharp-pointed apex projects between the somewhat swollen
antennal pits; basally it does not quite occupy the whole
vertical breadth ; eye-margins narrowly silvery, especially in
front; ocellar circle black ; hind head absolutely pale except
for two excessively narrow dark lines from vertex. Side
view :—The angle between frontal and facial planes is about
75°; antennz all quite pale yellow, except that the 3rd joint
is orange just,at the insertion of the black arista, which is
more elegantly and regularly haired than in the other
species, Jowls pale yeliow; palpi pale. In front the
mouth-margin is seen to be very narrowly darkened.
‘Thorax reddish orange, with the following greyish polli-
nated stripes :—Ceutrally a short narrow one extending but
Mr. C. G. Lamb on Exotic Chloropide. 41
a little distance on the disc, each side of this another which
runs to the outer angles of ‘the scutellum, diverging as they
go; beyond these the orange is very slightly suffused with
grey; calli shiny orange, “pleura the same. Scutellum
blackened orange, slightly punctate.
Wings yellowish, with orange veins; small cross-vein
just perceptibly beyond the costal ending of the Ist; hind
one about 3 times its length from the former, and with its
distance from the lower end to the 5th vein-ending about
15 times the distance apart of the cross-veins.
Halteres quite white, with yellowish stalks.
Legs entirely yellow, with no darkening at all.
Abdomen all darkened orange, with very narrow palish
hind margins.
Size nearly 3 mm.
Cam. Coll., Mozambique (F. M.).
Sleleocerus latiseta, sp. n.
There is a single specimen of a very distinct species.
Head (top view) :—Frons dull orange, lighter over an-
tenn, the triangle is remarkable in form ; basally it just
does not touch the eyes ; the margins are formed by raised
straight ridges; about halfway a sudden diminution in
breadth occurs, so that the side boundaries at that point are
suddenly shifted inwards; the space between these forward
parts of the ridges is necessarily a little depressed, but down
the centre runs a very fine raised ridge, which goes to the
front of the head to just behind where the side-ridges meet
in a slightly rounded curve ; the whole is shining bluish
black except the extreme tip, over which the frontal orange
runs; ocelli bright chestnut. Back of head entirely black.
Side-view :—LHyes very large, only leaving very narrow lower
and fairly narrow hind orange. jowls ; the orange frons is
just visible, the 3rd is orbicular and all orange ; the arista
is a little broader than usual and tapers to a fine point.
Thorax : dorsum dullish black, pollinated more and more
strongly with brown pollen towards the scutellum ; the pre-
scutellar depression is present, but is not sharply demarcated
from the rest. Scutellum as thorax, with divergent bristles.
‘The calli and an area below all orange, the pleura shining
dark brown, with a few lighter areas interspersed.
Wings clear, with brown veins, but with a faint smoky
cloud between 3 and 4 extending nearly to level of costal
euding of 2; the second vein long, as in lepidopus.
Halteres with white knobs.
42 Mr. C. G. Lamb on Exotte Chloropide.
Legs entirely yellow, except that the last two joints of the
very slightly dilated front tarsi are suffused.
Abdomen all rather shining brownish black.
Size (ex. antennz) 2 mm.
Cam. Coll., Chirinda Forest, S.A. (G. A. K. M.).
Meromyza, Meig.
M. capensis, Loéw.
There is a long series in Cam. Coll. which shows the very
considerable variation in abdominal and other infuscation
that occurs in this species.
Cam. Coll., Durban (F. M.).
Cam. Coll., Salisbury, Mashonaland (G. A. K. M.).
Bur. Coll., Zomba, Nyasaland (H. Stamus). .
Evurina, Meig.
Eurina oculata, sp. n.
There are two females in the Bur. Coll. which belong to
this genus, but do not fit with any of the hitherto described
species. ‘The eyes are larger than in most, being more of
the proportion shown by Meigen in §.B. vi. tab. lv. fig. 10,
though the frons is less protuberant.
Fig. 4.
Eurina oculata, X 40.
Head (top view):—Breadth nearly twice the distance
from vertex to tip of frons; vertex concave ; eyes promi-
nent and practically bare ; frons ending in a rounded cap
containing the antennal bases. The colour of frons &e. is
pale brown, dull; the rather darker triangle is nearly equi-
lateral, more shiny, with base about % vertical cross-breadth ;
Mr. C. G. Lamb on Exotic Chloropidee. 43
it ends in a very sharp raised ridge extending to the an-
tennal cap; the boundary is formed by two darkened
furrows bordered interiorly by two or three smaller parallel
furrows; the frons itself (along eye-margins exteriorly to
the triangle) has two very deep and broad furrows extending
along the sides of the triangle from the vertex to end of the
triangle ; the minute pairs of vertical bristles stand at the
beginning of these furrows. Ocelli brown, with tiny ocellars
each side of the front ocellus. The hind head is darkened
behind the ocellar triangle except along the actual vertex,
which is yellow in two long confluent spots; the rest of
hind head is brownish yellow.
Side-view :—The eyes are largerthan usual, nearly circular,
though slightly longer horizontally than vertically; the
frons is hence less prominent than usual, the distance from
antennal base to eye-margin being about 0°53 of the hori-
zontal eye-breadth ; the profile is less triangular than
usual, there being well-marked horizontal jowls of about +
the eye-depth running into the frons by a concave face-line.
The side is all brown-yellow except that the frons is there
infuscate. Antenne black, small 2nd joint, 3rd elliptical ;
arista pale, but brown on the swollen basal joint. The pale
face has a slight central swelling below antenne. Palpi
orange.
Thorax discally grey ; two mid-stripes darker, extending
from front to back, just separated till towards the scutellum,
where they meet; alongside these stripes is an interrupted
dark stripe forming a spot about the position of the cross-
suture, and a longer continuing stripe extending to the
scutellum ; just above the side-suture is a similarly broken
indistinct line ; ; callus and pleura dull orange-brown. Scu-
tellum orange, swollen, rounded in profile, ‘hairy, especially
on the margins, though no true bristles are present ; it is
darkened discally, with the orange showing through as a
narrow stripe.
Wings normal, with thick veins ; the thinning out of the
4th vein occurs suddenly at about } of its length; cross-
vein rather oblique. Halteres orange, with dark stalk.
Legs all orange-brown like the pleura, but slightly in-
fuscate dorsally on all the femora (less so on the front pair),
on the tibiz, and hind tarsi.
Abdomen somewhat flattened, dark brown, with sharp
narrow whitish margins ; ventrally all pale.
Size 4 mm.
Bur. Coll., Hagari, Madras Presidency.
Note :—‘“‘ Feeding on leaf-parenchyma of grass.”
44 Mr. 0. G. Lamb on Exotic Chloropide.
Lacaroceras, Becker.
There appear to be five species that fall within the limits
of this genus, of which at least three appear to be un-
described. They all agree with Becker’s diagnosis very
fairly, except that in two of them the 3rd antennal joint is
broader than he figures for his type-species, and is more
simply a long oval ; there appears, however, to be no good
reason for not placing them in his genus.
LI. megalops, Beck.
There is a single specimen which agrees very fairly with
Becker’s description and figure; the dorsum is quite
blackened all over, so that the three stripes are here
confluent.
Cam. Coll., Mozambique (F. .).
Lagaroceras anomalum, sp. 0.
There are several specimens of a species which very closely
resembles the above specimen in thoracic and abdominal
colour, in the legs and general facies, but is a little larger.
The differences are, however, marked and constant; they are
(1) the triangle, which has a different form; it is not truly
leaf-shaped, as in the generic diagnosis, but is practically
triangular, with concave (not convex) sides; it extends to
the front of head: (2) the antennz are relatively shorter,
though still nearly as long as the face ; the third joint is not
quite twice as long as the second and is about 1} times as
long as broad ; it is oval in form, with the upper tip sharply
rounded.
Size (excluding antenne) 3 mm.
Cam. Coll., Durban (Ff. M.).
Lagaroceras pulchellum, sp. 0.
This is a fine handsome species of the megalops group.
Head (top view) :—As broad as thorax ; frons black except
anteriorly just above the antennz, where it is orange; the
surface is dusted with greyish pollen and has many small
hairs; the triangle is highly polished black and is of a
pointed leaf-shape, the sharp stalk extending right to the
base of the antennz ; the base is rounded and occupies a
little over 2 of the vertical breadth. The head-bristles are
well marked ; the back of the head is black. Side-view :—
Profile a little more trapezoidal than normal, the line from
—s ee
Mr. C. G. Lamb on Ezotic Chloropide. 45
antennal base to mouth-edge nearly straight ; jowls about
equal in breadth to that of the 3rd antennal joint—they are
silvery, as are the hind jowls. The antenne are inserted
just on the edge of the orange frons and are just about as
Fig. 5. Fig. 6.
ya ;
Fig. 7
A. B
Fig. 8
\
aa Z
wef,
ee ae
. /
aT
Fig. 5.—Lagaroceras anomalum, X 50.
Fig. 6.—Lagaroceras longicorne (?), x 50.
Fig, 7.—A. Lagaroceras megalops, X 30. 3B. Lagaroceras anomalun,
x 3d.
Fig. 8.—Lagaroceras pulchellum, X 35.
long as the face is deep; they are all darkish orange, except
that the upper half and the tip, of the 3rd joint are black-
ened ; this joint is about 14 times as long as the 2nd, which
46 Mr. C. G. Lamb on Exotic Chloropide.
is itself rather unusually long; arista white and closely
pubescent as usual, with the smooth basal joint yellow.
The face is shining black, with silvery lines from the an-
tennal bases to the mouth and with silvery lower eye-
margins ; palpi black.
Thorax: the dorsum is finely granulated; the general
ground-colour is dark greyish, with the following black
marks :—a broad median line vanishing about halfway, a
_ very fine black line running down the centre of each of the -
rather obsolescent furrows; beyond this a broadish line
from just in front of the callus to the side of the scutellum,
interrupted by the grey ground where the cross-suture
should be; last d. c. well developed; pleura shining black,
with oblique row of three pale yellow spots—a triangular
one just behind the humeral callus, a sloping one on the
mesopleura, a horizontal one above the mid-coxa. Scu-
tellum pale orange, nearly flat, hairless, with long crossed
terminal bristles and small accessory ones beside the main
ones and close to them.
Wings clear, with brownish veins ; the venation is not
quite as given by Becker for megalops (I., tab. iii. fig. 47) ;
the 2nd vein is quite parallel to the 3rd all the way, and the
distances between the ends of 2 to 3and 3 to 4 are nearly
equal.
Legs orange ; femora progressively more infuscate from
front to hind ; last tarsal jomts dark..
Abdomen entirely deep dull black, smooth.
Size (ex. antenne) 3+ mm.
Cam. Coll., Durban (F. M.).
To the section with dark and rather rugose scutellum
belong two species. One of these will pass for L. longicorne
of Thomson (Hug. Resa, p. 604). It agrees quite well with
what would be a dark form of the species, though not so
well with Becker’s description (I., p.108). The remarkable
antenne are even longer than may be inferred from the
description ; the thorax can be described as black, rather
rugose, with three narrow, smoother, grey stripes; the
scutellum like thorax, centrally black, with the sides orange.
It is possible that we have a new species here, but as there
is but the single specimen, it is left provisionally in this
species. Thomson’s species was from China, this is from
S. India.
Size (ex. antennze) nearly 3 mm.
Bur. Coll., Coimbatore, Madras Presidency.
+0
Mr, C. G. Lamb on Ezotic Chloropide. 47
The following is a second species of this section ; it occurs
also as a single specimen which, like LZ. anomalum, has a rela-
tively stout antenna—in fact, the antenna is practically as
shown in fig. 5 (p. 45).
Lagaroceras infuscatum, sp. nu.
Head (top view) :—Frons all palish ochreous brown, dull
and black-haired, the triangle shining dark orange-brown
and of peculiar shape; the basal part is about 2 as broad as
the vertical cross-breadth ; it continues normally along the
frons, but about midway is suddenly constricted, and then
continues like a narrow spear-head to the antennal base ;
each side of the constricted point is a yellow raised spot on
the triangle ; the surface is somewhat variegated in striz
and the middle area is rather darker than the rest; just at
the hind eye-angles occur the usual pale spots carrying the
vertical bristles. The f. 0. b. small, but distinct. Hind head
all black. Side-view :—Frons a little prominent, covering
the antennal base, brown; face-outline nearly linear, if
anything slightly concave, from antenne to mouth. The
total length of the antenne is about equal to the face; the
2nd joint about half as long as 3rd, which is a little lese
than twice as long as broad ; rounded oval in outline, all
darkened except for a tiny spot of orange on the base of
8rd joint below ; arista normal, white and pubescent, with
smooth pale yellow base. Jowls, lower and hind, palish
yellow, the former about half the depth of 3rd joint. Palpi
black. The face is darkened with a narrow emarginate
paler mouth-margin ; the eyes have short silvery margins.
Thorax: dorsum black and finely punctate, with three
very narrow grey lines. Scutellum flattish, similar to thorax,
with a dark orange median line, two terminal and one adja-
cent smaller bristle each side. Pleura very shining brownish
black, except for a yellow stripe just below the mesopleura.
Wings with venation similar to pulchellum, 2nd and 3rd
quite parallel, but the cross-vein is slightly sloped backwards.
Halteres white, with a brown stalk.
Legs orange, the femora progressively more infuscate
from fore to hind pairs, the last tarsal joint darkened.
Size (ex. antennz) 24 mm.
Cam. Coll., Durban (/. M.).
Hapuieeis, Loéw.
Haplegis nitens, sp. n.
A small form, considerably more shining than H. tarsata.
48 Mr. C. G. Lamb on Exotic Chloropidee.
Head (top view) :—Entirely black and somewhat shiuing,
even on eye-borders; the triangle fairly close to eye on
vertex, extending with the usual straight sidés to a sharp
point over antennz, excessively shining, with the usual
shallow but sharp depressed middle trough; the bordering
hair-rows very indistinct. Hind head all black. Side-
view :— Lower jowls dull orange and very narrow ; antennz
of normal form, 3rd black and round, 2nd bright orange ;
arista black and finely pubescent, the longish basal joints
more orange. Face fairly silvery; tongue and palpi
blackened.
Thorax all entirely shining black, including the scutellum,
which has two longish end-bristles ; the whole dorsum has
a regular clothing of very fine brown hairs; the pleura is
faintly orange in some parts.
Wings clear, normal in venation, brown veins. Halteres
with almost white head.
Legs entirely clear orange, including front coxa and all
the tarsi.
Abdomen shining black.
Size 2 mm.
» Cam. Coll., Durban (F. ¥.).
Exacuierereicus, Beck.
HX. bistriatus, Beck.
Specimens from Durban, Cam. Coll. (F. W.).
Camarora, Meig.
(Modo, Oscinis, Latr.)
C. angustifrons, Bezzi.
Specimens from Durban (F. M.) agreeing well with Begzi’s
description.
Merarostiema, Beck.
M., sauteri.
Specimens in Bur. Coll. from Coimbatore, Madras.
Cuatcipomyia, de Meijere.
This genus was described in Tijd. v. Ent. (vol. liii. p. 156)
as a Drosophilid, the error being due to the insect possessing
a remarkable bipectinate arista. Becker redescribed it in its
Mr. ©. G. Lamb on Exotic Chloropide. 49
proper family as Hemispherisoma (II1I., p. 47). The syno-
hymv was given by de Meijere in Tijd. v. Ent. (vol. lvi.
p- 571). In both cases the specific name selected for the
type-species had been politus, but for some reason de Meijere
changed it to beckeri, though both types were the same
species.
C. polita, de Meij.
Specimens in Bur. Coll. from Taliparamba, Malabar,
with the note :—‘ In ginger-stems attacked by Dichocrocis.”
Curomatorrerum, Beck.
Chromatopterum lacteiventre, sp. n.
This species has the pubescent arista of the Iudian species
C. pubescens, Becker (II1., p.82), but its facies is that of the
African C, delicatum, Becker (LI., p. 413).
Head (top view) :—Frons almost entirely covered by the
brilliant shining black ‘‘ triangle,’ which has its sides con-
tiguous with the eyes and a rounded frout margin reaching
to the antennal base ; its sides converge slightly to the
front; the only part of the frons left uncovered by it are
two small, dull orange, triangular patches each side in front ;
the surface is broadly and shallowly depressed ; the ocellar
hump is slightly raised and carries chestnut-coloured ocelli,
Side-view :—T'he semicircular eyes cover the whole, pro-
jecting beyond the face and leaving practically no lower
jowls and only a small hind eye-border, which is shining
black, as is the whole hind head. ‘lhe antennal 3rd joint is
almost orbicular, just a little Jonger than deep, orauge on
lower half, blackened on top; arista inserted basally, hair-
like except for the small pale basal joint, finely pubescent ;
2nd jomt yellow. Face darkened orange ; palpi black.
Thorax (including scutellum and pleura) all shining
black, the dorsum just before the scutellum aud the scutellum
itself very lightly dusted with orange pollen; the rounded
and slightly swollen scutellum with moderately long slightly
divergent end-bristles and a few accessory side-hairs,
Wings with venation as figured by Becker (1I., tab. xii.
fig. 10), but the blackening is different ; the frout blackeuing
is confined to the first part of first vein, the space between
it and where the auxiliary vein would be (ikea long stigma),
and the thickened black costa itself, from which a faint
suffusion runs on to the neighbouring cell ; the end spot is
smaller and discrete, it touches the costa midway between
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 4
50 Mr. C. G. Lamb on Exotic Chloropide.
the ends of 2 and 3, but does not extend to the end of wing
or down to vein 3.
Halteres with ivory-white heads.
Legs mainly orange, all the coxe black and all femora
broadly ringed with black.
Abdomen: dorsum flattened, a little longer than broad,
and tapering in outline from the base; it is of a quite unique
colour, being all suffused with a dense milky-blue glaze ;
the last segment is considerably longer than the others;
beneath, the abdomen is orange ; the last segment, which is
bent under, is all shining black.
Size about 1? mm.
Cam. Coll., Peradeniya, Ceylon (J. C. F. Fryer).
Ops, Becker.
O. madagascariensis, End.
A specimen in Cam. Coll., Durban (F. M.), differs from
the ordinary form only in the femora being somewhat
darkened.
O. callichroma, Loéw.
There are two specimens of this species—the one in Bur,
Coll. from Nyasaland, in which the abdominal cross-bands
are rather weak and indefinite. The other is a very bright
and shining form, which might be taken as a subspecies.
It is a little larger, aud the “triangle” covering nearly all
the frons is very deep excessively shining black instead of
being shining brown. ‘he abdominal markings are also
very clear aud distinct ; they consist of the following on the
yellow background :—Ist segment with very shoit central
bar ; 2ud arched bar with the springings situated basally ;
8rd broad, only leaving narrow hind margin yellow; 4th
niedian, of half total breadth of segment; the pointed Sth
has a narrow basal band.
A specimen in Cam. Coll., Durban (. .).
Ops nigra, sp. n. 5
The whole of head and thorax entirely shining black,
except for the orange antenne and bright yellow scutellum.
The vertical triangle does not cover the whole of the trons,
but leaves eye-margins narrowly widening right from the
vertex.
Wings quite normal, clear.
Mr. C. G. Lamb on Exotic Chloropida. 51
Legs orange, with coxa black ; femur very dusky except
at tip. Knobs of halteres whitish orange.
Abdomen orange, with dark bands somewhat similar in
form to last species, but all of them broader in proportion
and less well demarcated.
Size 2 mm.
Cam. Coll., Mozambique (F. .).
Cuuororpisca, Loéw.
There are two single-specimen species—one resembling
obscurella, but with a rounder head, the other like a true
Chlorops, but with somewhat flattened scutellum. It is not
advisable to describe from these single specimens.
Curorors, Meig.
C. contribula, Loéw.
Cam. Coll., Durban (F. M.).
C. levigata, Beck.
Cam. Coll., Durban (Ff. ™.).
Chlorops zeylanica, sp. n.
There is one species which will not accord with any of
Becker’s species in the Indian fauna. It has a somewhat
exceptionally prominent head (see fig. 9) and belongs to the
section with fine white arista.
Fig. 9.
Chlorops zeylunica, X 80.
Head (top view) :—Frons (fig. 10) dull pale ochreous
yellow covered with black hairs; the triangle very large,
with its boundary well defined nearly up to the vertex, but
there less so ; it extends to the extreme front, with slightly
4%
52 Mr. C. G. Lamb on Exotic Chloropide.
concave sides bordered with hair-lines, and it is the same
colour as the frons but shining and suffused across the
middle with pale brown, as shown by the dotted boundary-
line in the figure; the ocellar spot is black; a very distinct
but narrow furrow runs from front ocellus right to edge of
frons. Hind head broadly black, with pale yellow bordering
stripes starting from the vertical bristles. Side-view as in
fig. 9; all yellow, the haired frons more orange, rest quite
bare except for a few oral hairs. Antenne with yellow
basal joints, deep black orbicular third ; arista white, basal
joints a little suffuse, pubescence very fine. Face all pale
yellow; palpi pale, but just perceptibly infuscate outside at
tip ; tongue yellow.
Thorax: dorsum moderately shining yellow, with black
hairs ; three broad black stripes, the middle one beginning
on neck and extending to scutellum, the side ones abbre-
viated in front but meeting the middle one behind, so that
they form an almost uninterrupted band on hind dorsum ;
small side-lines above the wings run into the main pair ;
humeri pale yellow. Pleura pale yellow; a shining oval
black spot on the lower front angle of the mesopleura, the
usual black triangle over the middle coxa, and a smallish
black oval spot over the hind coxa.
Wings normal, clear, thick-veined; in one of the two
specimens both the hind cross-veins are broken in the
middle. Halteres bright whitish yellow, with darkened stalk.
Legs entirely yellow except that the front tarsus and last
joints of the others are very faintly infuscate. _
Abdomen: dorsum all brown-black, slightly shining, the
hind margins of all segments but last very narrowly yellow,
the last broadly so; venter paler.
Size (ex. antenne) 35 mm.
Cam. Coll., Peradeniya, Ceylon (J. C. F. Fryer).
PaRECTECEPHALA, Beck.
Parectecephala varifrons, sp. n.
A species in the Cam. Coll. is best assigned to this genus;
the triangle is rather longer than normal according to the
descriptions of the known species.
Head (top view, fig. 12) :—Frons about 24 times as broad
as one eye and about 1} times longer than broad, projecting
about 4 its length beyond a line touching the eyes in front ;
eye-margins parallel, the projecting forehead narrowing a
little and ending in a broad pointed tooth overhanging the
7 Mr. C. G. Lamb on Exotic Chloropide. 53
antennal pits; frons bright dull orange, with small scattered
black hairs ; the triangle has its base about § the breadth at
vertex ; the bounding lines are nearly straight, a little raised,
very narrowly yellow, and meet just beyond the level of
the eyes; they continue nearly to the front in a shining
yellow stalk; inside these narrow lines the triangle is
mainly shining chestnut, but is variegated by lighter colours,
Fig. 11.
Parectecephala varifrons, sp. n., X 50,
Fig. 18,
Parectecephala varifrons, sp. n., X 40,
so that the most prominently visible chestnut part is a
rhombus extending from the triangle’s tip to the front
ocellus ; this area is also very shallowly hollowed out ; the’
lighter parts consist of (1) a pair of oval dull brightish yellow
spots each side of the ocellar area, and extending thence-
right to the sides of the triangle; (2) two more orange and:
54 Mr. C. G. Lamb on Ezotic Chioropide.
more shining spots extending from hind ocelius to the outer
angles of the triangle. ‘The hind head is orange except fora
large black patch extending from the base of the triangle.
Side-view (fig. 13) :—All pale whitish yellow, the side of
frons orange ; eyes very nearly circular ; jowls about depth
of 3rd ; face somewhat concave ; antenne as fig. 11, but the
arista for its last 3 is very faintly white pubescent, not bare
as figured, 3rd joint orange with blackened tip ; tongue and
palpi yellow. Face whitish, unkeeled, but depressed, the
actual facial ridges being marked with a narrow pale grey
line ; antennal pits well marked with dark shining chitinous
edges.
Thorax: dorsum dull palish orange ; a broad black central
stripe from neck to end of scutellum—this is very intense
up to about the middle of the dorsum, then gets much
fainter, till it is very faint on the scutellum ; each side is
another uniformly black line, rounded and abbreviated front
and back, and diminishing that way in breadth; below is
another very thin blackened line extending forward from
just above the wing for about ? the pleural length. Scu-
tellum (as above) suffused centrally, sides orange, not
flittened, a little hairy, pair of terminal bristles ; meta-
notum darkened ; the pleura all rather shining pale yellow,
with a small elongate spot.
Wings normal, clear, with brown-orange veins, the
distance between cross-veins about equal to the last part of
5th; halteres with whitish knobs.
Legs long, all yellow except for the last two darkened
tarsal joints.
Abdomen shining brownish orange, with very narrow
pale segmental margins ; venter paler.
Size 44 mm.
Cam. Coll., Durban (F. ™.).
PEMPHIGONOTUS, gen. nov.
In the Bur. Coll. are three specimens (1 ¢,2 2) of a
remarkable insect from Melville Island which exhibits
marked sex-dimorphism.
Characters common to both sexes :—Texture horny,
macrochetes quite absent, though bedy hairy; scutellum
very large, swollen, standing in profile well above the
thoracic level (see fig. 14), with a flattened area of different
texture; abdomen oval and flattened ; wings with very long
discal cell. (see fig. 14), the auxiliary and anal veins just
visible as “ shadows” of veins; legs long ; antenna like that
Mr. C. G. Lamb on Exotic Chloropide. 55
of a true Chlorops; the triangle narrow and ill-defined,
with a better-defined narrow central line, only differen-
tiated by shine from the rest of frons.
The male has a remarkable arch in the first part of the
costa, which carries a fringe of very long hairs ; the mid-
femur and tibia are also clothed with abundant long tangled
hairs,
Fig. 14.
Pemphigonotus mirabilis, x 12.
Fig. 15.
Pemphigonotus mirabilis, x 22.
Pemphigonotus mirabilis, sp. n.
The insect is all red-orange, slightly darkened in various
places except where otherwise mentioned.
56 Mr. C. G. Lamb on Evotic Chloropide.
3 .—Head (top view, fig. 15) :—Frons bare, dull except
for the very narrow redder mid-line extending from ocellus
to forehead and the narrow ill-bounded main triangle; no
eve-mar ins ; hind head hairy at upper corners behind eyes.
Side-view (fig. 14):—Antenne and palpi clear yellow ;
arista hair-like, pale ; tongue fleshy and hooked at tip. In
front the face is wide, with no keel except a tiny bar be-
tween antenne; margin of mouth arched.
Thorax bare on dorsum, which is flattened and dull except
for a central shining line extending to the shining base ; the
sides above the notopleural suture and all the pleura are
abundantly clothed with long pale hairs. Scutellum enor-
mously swollen both sideways and upwards, smooth except
for an extraordinary flattened area on the disc, which
is slightly dimpled ; the base towards thorax has two large
blackened areas with a pale line between; it is hairy, with
pale hairs, which are longest and regular on the margin ;
notopleura smooth,
Wings as fig. 14, the costal elevation from base to Ist
vein with a row of long, dark, silky hairs; the whole surface
much suffused except a rather narrow lower margin from
axillary angle to near the end of the 5th, and again from
beyond that end to just across the 4th.
Halteres practically white.
Legs long, hairy, all pale orange except for a slight suffu-
sion on the front tibia and the darkened tarsi ; all the tarsi
somewhat swollen. Tie middle femur and tibia with abun-
dant long pale hairs.
Abdomen flattened, long-oval, the maximum breadth
being about twice the thoracie breadth.
The 2 differs as follows:—Thorax not so dull and not
flattened ; wings with no costal elevation, the whole costa
being very gently curved in a continuous manner; no long
hairs on costa; no long hairs on middle legs.
Size about 5 mm.
Bur. Coll., Melville Island, N. Australia (G. F. Hill).
BatTHYPARIA, gen. nov.
Becker describes a genus Euryparia (III., p. 84) which
occurs in Formosa; it has very deep jowls, quadrate 3rd
antennal joint, and is covefed with white hairs. Among
the Durban species there are several specimens of a very
handsome small Chloropid that have the above characters,
Mr. C. G. Lamb on Evotic Chloropide. 57
especially the bright silvery clothing, and even a faint
central wing-cloud, in common with Becker’s species; but
they differ greatly in that the eyes are long-oval and the
antenne are smaller. The thorax im the species represented
is black and not striped, and scutellar bristles are present.
They must form the African equivalent of the Asiatic genus.
Head (see figs. 16 & 17) :—The facial and frontal tangent-
planes meet at about 120°; eyes long-oval, with axis nearly
upright; jowls very deep, about half the depth of the long
eyes ; antenne nearly as long as face, with a practically
rectangular 8rd joint about twice as long as broad, and a
very fine, slender, bare arista, thickened basally. Frons
parallel-sided, with a long rather narrow triangle from
vertex to front only just differentiated by its extra shininess
from the rest of frons. Wing-venation as fig. 18, the 3rd
and 4th veins just not reaching the edge.
The whole insect is covered with brilliant shining white
hairs even on the frons; these are very stout and bright on
the thorax and head, but less so on the abdomen. Unlike
Euryparia, there is a pair of scutellar bristles inserted in
the same manner as in Ops.
The palpi are quite peculiar, being rather stout, long, and
spoon-shaped.
Bathyparia preclara, sp. n.
Head (top view) :—Chestnut-brown, the triangle more
shining ; the silvery hairs along the triangle’s border beud
across it ; eye-margins broad and very silvery ; the verticals
and ocellars white ; hind head all black except just on vertex
behind ocelli, where is a long yellowish stripe along the
vertical ridge. Side-view:—Similar in colour, the broad
hind eye-margin very silvery, as is the hind jowl. Antenna
slightly darkened brown; arista pale at base. The palpi
are long, spoon-shaped, and silvery grey ; tongue dark.
Face the same brown colour, side-ridges well developed ; no
median keel, so that the antennz nearly touch basally.
Thorax : dorsum, meso- and sternopleura all shining black
and punctate, covered with bright silvery-white hairs arising
from the punctures ; the rest of pleura bare. Scutellum
bright yellow, with approximated pale terminal bristles and
silvery hairs like thorax. Notopleura black and dull from
very faint shagreen.
Wings (see fig. 18) clear, with brown veins; in several
specimens the central part is very faintly tinged with brown,
Halteres pale yellow.
58 Mr. C. G. Lamb on Exotic Chloropida.
Legs long, with slightly dilated tarsal joints, all covered
with the fine white silky hairs; front pair all bluck except
for orange trochanter and knees ; middle with orange tro-
chanter, black femur, the rest nearly white ; hind with dark
orange trochanter, femur black with pale knee, tibia pale
Fig. 16. Fig. 17.
Fig. 18.
Bathyparia preclara, X 40.
and more or less darkened about the middle, tarsi nearly
white.
Abdomen smooth, shining black, the silky hairs evident
but sparse and fine, a little longer at upper angles.
Size 24 mm.
Cam. Coll., Back Beach, Durban (Ff. .).
Mr. G. J. Arrow on Melolonthine Coleoptera, 59
III.—Some Systematic Notes on Melolonthine Coleoptera.
By GiLpert J. ARRow.
(Published by permission of the Trustees of the British Museum.)
Mr. L. PéRINGUEY, in his “ Catalogue of the Coleoptera of
S. Africa” (Trans. 8. Afr. Phil. Soe. xiii. 1904, p. 174), puts
at the head of the genus Sparrmannia a species which he
calls vertumuus, Pall. (with the names alopex, F., and
brunnipennis, Oast., as synonyms), mentioning a typical
form with pale testaceous colour as inhabiting the Karroo
region, and a form with “light” (apparently meaning dark)
chestnut elytra in Namaqualand, Bushmanland, and Damara-
land. The recent Catalogue of Dalla Torre adopts this
synonymy, but separates as a variety the dark form brunni-
pennis.
Dr. H. Brauns has lately sent a series of this dark form,
which he has found in abundance in the Uniondale district of
Cape Colony, while the light form is equally abundant in the
Willowmore district, only 42 miles to the south, but separated
by the range of the Zwaartberg running from west to east
across the continent. Examination has proved that the two
forms are quite distinct, and Fabricius’s description shows
that itis the dark form which is the true S. alopex. It was
Fabricius himself who, in his Syst. Eleut. ii. p. 163, identified
this insect with the Scarabeus vertumnus otf Pallas, but with
strange carelessness, for the latter is a Russian species,
apparently belonging to the genus Rhizotrogus.
The light-coloured Sparrmannia, described at length by
Péringuey, is therefore without a name, and I propose to
call it
Sparrmannia flava, sp. n.
In addition to the. pale-coloured elytra, this species differs
from 8. alopex in their more distinct and regular punctura-
tion, in the longer tarsi of both sexes, and especially in the
longer middle tarsi and more dilated hind tibize of the male.
The xdeagus is figured by Péringuey. That of S. alopes is
much shorter and blunter. Dr. Brauns states that, while
S. flava occurs together with S. alopex north of the dividing
range, he has never seen the latter south of the mountains,
aud that no specimens of intermediate coloration are found,
S. flava generally appears at Willowmore towards Christmas
time, while S. alopew is later, generally appearing in January
60 Mr. G. J. Arrow on Melolonthine Coleoptera.
and February. Both are nocturnal, and hide in loose soil
during the day.
There is another closely similar species, of which specimens
are probably included amongst those enumerated by Périn-
guey, and which I have wrongly determined as 8S. ver-
tumnus in Denkschr. Med. Nat. Gesellsch. xiii, 1908, p. 438.
I now call it
Sparrmannia similis, sp. n.
Pallide flava, capite, pronoto, scutello, pectore abdomineque longe
et densissime lanatis. S. flave valde similis, sed clypeo paulo
minus profunde exciso, elytris crebrius sed minus distincte
punctatis tarsisque paulo minus elongatis.
Long. 22 mm.; lat. 11°5 mm.
Hab. 8.W. Arrica: Hereroland.
This has an extremely close resemblance to S. flava, but
the elytra are finely and confusedly, instead of strongly and
sparingly, punctured, the clypeus is acutely, but less deeply,
notched in the middle, and its sides a little less rounded, and
the tarsi, or, at least, the middle ones of the male, are not
quite so long. The edeagus of the male is drawn out intoa
tube just behind the orifice.
Upon p. 287 of his Catalogue already referred to, Mr.
Péringuey recognizes two South-African species only of the
genus Asthenopholis—subfasciatus, Blanch., and erassus,
Arrow ; but the species to which he has wrongly applied the
Jatter name is evidently the true A. adspersus, Boh. (=trans-
vaalensis, Brenske), and in <A. subfasciatus he has included
the quite distinct A. mznor, Brenske, These four species
may be distinguished as follows :—
I. Scutellum well punctured; hind tibia little
dilated at the end.
a. Scales of the upper surface long and hair-like. swbfasciatus, Bl.
b. Scales of the upper surface short and broad .. minor, Brenske.
II. Scutellum smooth or almost smooth ; hind tibia
strongly dilated at the end.
ec. Pronotum moderately covered with long sete. adspersus, Bohem,
d. Pronotum closely covered with oval scales .. crassus, Arrow.
A. subfasciatus seems to be confined to Cape Colony,
A. minor to Natal, A. adspersus to Natal and the Transvaal,
whilst A. crassus is known only from British East Africa,
Brenske’s species were determined for me by himself, and
Mr. Péringuey has certainly determined them wrongly,
Mr. G. J. Arrow on Melolonthine Coleoptera. 61
although he has had the assistance of type-specimens. The
genitalia of the males are quite different in the three species
he has united, notwithstanding his statement.
Mr. Péringuey has founded a genus Euronycha, but has
not included in his Catalogue the genus T’riodonta, of which
many African species are known; and as the sole character
by which he differentiates Zuronycha (a feature of the male
alone) is found in Triodonta, they must be considered the
same,
The type of Heterochelus gonager, F., in the British
Museum is the species called by Burmeister HH. longipes, as
Mr, Péringuey has recorded upon my authority (‘lrans. 8.
Afr, Phil. Soc. xiii. 1908, p. 698) ; but the quite different
species to which the name gonager was applied both by
Burmeister and by himself in vol. xii. of the above work
remaius without any available name. 1 propose to call it
Heterochclus melanopygus, sp. 0.
The two following species of South-African Hopliini were
described several years ago at Professor Poulton’s request,
but the descriptions have remained unpublished. The insects
were amongst those collected more than a century ago by
the African traveller Burchell, and now in the British and
Oxford Museums. The data are taken from Burchell’s note-
books in Professor Poulton’s possession.
Gouna burchelli, sp. n.
Rather large, broad, sooty black, naked above, beneath
thinly clothed with black hairs and a few white scales at the
sides ; head broad, rather convex and rugose above, clypeus
short, not angulate but bilobed ; prothorax rather broader
than long, strongly contracted in front, front angles acute,
hind angles very obtuse, surface finely punctate, with a faint
longitudinal channel ; scutellum small, almost semicircular ;
elytra broad, faintly costate, irregularly and inconspicuously
punctured ; pygidium (male) large, inturned, transversely
punctate-rugose ; legs (male) rather long, hind ones slightly
* thickened, unarmed, front tibie tridentate, the innermost
tooth rather small and distant, all the claws single and
minutely cleft, but those of the hind legs hardly visibly.
Length 9 mm. ; greatest breadth 5 mm.
Locality. Burchell’s two specimens (nos. 318 and 319)
62 Mr. G. J. Arrow on Melolonthine Coleoptera.
were captured on the morning of Nov. 3, 1814, at Duyker
River, in the south of Cape Colony, a little to the west of
Mossel Bay.
The type is one of three specimens in the British Museum
derived from the Pascoe Collection, There are also four from
the Fry Collection and one from the Reiche Cellection. A!I
these, as well as the two brought by Burchell, are males, and the
other sex remains still unknown. ‘The species was wrongly
identified with AMonochelus spinipes, FE -, by Reiche, and has a
general resemblance to that insect, but its structural characters
are not those of Monochelus. They negree with those formulated
by Mr. Péringuey for his genus Gouna, one of those created
by the dismemberment of the old Gymnoloma. This dis-
memberment is very unsatisfactory, since by a process of
elimination the original genus is left without tangible differ-
ential features at all. The present form, howls ry is nearly
related to Gymnoloma lineolata, the type of Gouna, although
much larger and broader, Its comparatively large size and
sooty-black surface render it easily recognizable. I at first
suspected that the absence of scales from the upper surface
might be due to age; but the specimens are in general well
preserved, and, as all agree in being smooth on the upper
surface, they are evidently in their natural condition,
Dicranocnemus burchelli, sp. n.
Fuscous, with the elytra and legs reddish. Rather elon-
gate, the thorax distinctly longer than wide and not gibbous,
Clypeus parabolical, the front margin very slightly reflexed
and with scarcely visible angles. Upper surface of the head
uniformly finely rugose and pubescent. Prothorax mode-
rately convex, the sides regularly rounded and converging
forwards. Front angles acute, hind angles obsolete.
3. Prothorax finely rugose and densely clothed with
rather short tawny pubescence, which changes into scales at
the posterior margin, ‘The median sulcus is deep behind, but
vanishes beyond the middle. The scutellum is clothed with
elongate whitish scales and the elytra with round scales
varying in colour from chocolate to pale yellow, the light
ones forming a median longitudinal stripe which is broadest
near the shoulder, a sutural stripe broadest at the apical end,
and a quadrate ‘pateh between these. The pygidium and
propygidium are densely covered with orange scales, with a
darker band at the base of the former. The claws of the
middle feet are without a basal appendage
Length 5°5 mm.
Mr. G. J. Arrow on Melolonthine Coleoptera. 63
2. The prothorax is without a median sulcus. It is not
finely rugose, but strongly punctured, and clothed with
ereyish hair, longer but less dense than that of the male.
There are no scales. The elytra are more thinly clothed with
decumbent sete of an almost uniform tawny colour.
Length 4°5 mm.
Hab. Burcliell’s eight specimens, all of which are accounted
for, were captured in flowers, five of them at Uitenhage
(Nov. 28 and Dec. 1, 1813), and two between Kra Ka Kamma
and Van Stade’s River (Feb. 7, 1814), near (S.W. of)
Uitenhage. ‘lwo from each locality are in the British
Museum, but there is no means of associating these specimens
witli their precise data,
Types (g and ? ) in British Museum.
The description is based upon nos. 1303 and 1305 in the
British Museum. ‘I'he specimen numbered 1308 is rather
‘smaller and shorter, and may possibly prove to be distinct ;
but it is most likely only an aberrant individual of the same
form.
From the description, this species must be very nearly
related to D. hypocrita, Péringuey, which has on each
elytron two discoidal bands of pale scales coalescing at thie
middle, whereas only one is present in our form. In the
female no pattern is traceable. A male and female of the
species were compared by Mr. Guy Marshall and Mr. Pévin-
guey with the Péringuey type at Cape Town and the ? (293)
named L/eterochelus longipes, Burm., the g (294) Dicrano-
cnemus squamosus, Burm. Both, however, show the form of
front tibia distinctive of Dicranocnemus, while D. squamosus
is characterized by a peculiar formation of the middle claws
of the g which is absent here. WD. burchelii is one of the
very numerous species of this group of which the sexes are
quite dissimilar, so that, in the absence of sufficient evidence,
they are frequently associated wrongly. The question has
been settled for us in the present instance by Burchell.
Four males and four females were taken by him, and of these
one of each given to the British Museum were placed on the
same pin, showing his conclusion that they belonged to a
single species. It will be seen in the above description that,
in addition toa difference of shape, the elytra of the male are
decorated with orange scales, with a paler sutural patch and
longitudinal stripe upon each, while the female is uniformly
clothed with grey hair. Hence it is not surprising that, in
the absence of direct evidence, they should have been assigned
to different species, and even different genera,
64 Mr. G. J. Arrow on Melolonthine Coleoptera.
Both generic and specific names of Blackburn’s Neolepid/ota
obscura are redundant, the insect being a common Indian
species, Holotrichia serrata, F., of which an old specimen in
bad condition and of unknown origin unfortunately fell into
Blackburn’s hands. It is now in the British Museum.
I believe Lepidiota bovilli, Blackb., to be identical with
L. rothei, Blackb. In spite of a careful comparison of his
types, I am quite unable to detect the differences mentioned
by him.
NEMATOSERICA, gen. nov.
Corpus nonnihil elongatum. Mesosternum haud productum, sat
latum. Pedes graciles, tibia antica lata, bidentata, postica modice
angusta, fortiter spinosa, tarsorum posticorum articulo primo
quam secundum daplo longiori, Ungues profunde fissi, parte
interna brevi et lata. Antenne 10-articulate, clava(¢) quadri-
articulata, longissima, lamellis eequalibus; (2) triarticulata.
Clypeus .vix angustatus, margine antico reflexo, subtiliter
sinuato, superficie anteriori haud lato. Oculi haud magni,
remoti. Prothoracis latera postice sinuati, angulis posticis acutis,
Elytrorum margines postici arcuati, ad suturam depressi.
Nematoserica cwrulea, sp. n.
Cerulea vel viridi-czrulea, sericea, clypeo tibiis tarsisque nitidis,
antennis nigris ; modice elongata, convexa, capite, corpore subtus
pygidioque pallide setosis, elytroram Jateribus fortiter nigro-
setosis, clypeo parce punctato, margine valde reflexo, antice sub-
tiliter sinuato; pronoto parcissime punctato, lateribus bisinuatis,
angulis posticis acutis, paulo productis, basi utrinque late
impresso; elytris fortiter sulcatis, sulcis sat vage punctatis,
apicibus separatim arcuatis, parte postica ad suturam depressa,
corpore subtus opaco, grosse setoso ; pygidio sat fortiter punctato.
Long. 5-5°5 mm.; lat. max. 3-3°5 mm.
Hab. BoRNEO (Sarawak): Puak (G. EH. Bryant, April,
May).
Type in the British Museum.
This beautiful little insect is chiefly remarkable for the
length of the 4-jointed antennal club of the male, which is
relatively longer than in any other species of Sericineas known
tome. The tour lameilee are of equal length—at least five
times as long as the foot-stalk—and little shorter than the
elytra. The bright blue or greenish-blue colour is also, so
far as I know, unique. The upper surface is silky and sub-
opaque, with the clypeus alone shining, the margin of the
On new Pyralidee of the Subfamily Pyraline. 65
latter broadly reflexed, the front margin very gently excised,
and a row of stiff bristles traversing the middle from side to side.
Lhe eyes are rather small and far apart. The lateral margins
of the prothorax are distinctly sinuated in their posterior half,
the hind angles a little produced and acute and the base
impressed on each side. The elytra are sulcate and the sulci
contain rather coarse but shallow punctures.
The genus is apparently related to ZYeraserica, which I
do not know, and which has been described from the male
alone, the antenna of which has the last four joints rather
long but much less elongate than in the present insect. This
has not the forehead narrow and the eyes very large and
prominent, as in Teraserica, The strongly bisinuated sides
of the prothorax and acutely produced hind angles are very
characteristic, and another peculiarity which, so far as I know,
is not found elsewhere is in the shape of the elytra. hese
are separately rounded at their hinder margins, with the
sutural angles extremely blunt, so that a wide angle is formed
and a considerable part of the abdomen exposed. ‘he
peculiar appearance, however, is chiefly due to the fact that
this part of the elytra is strongly depressed along the suture.
IV.—Descriptions of New Pyralide of the Subfamilies
Epipaschiane, Chrysauginze, Endotrichine, and Pyraline.
By Sir Georcs F. Hampson, Bart., F.Z.S., &c.
[Concluded from vol. xviii. p. 373.]
(26) Pyralis nigricilialis, sp. n.
3. Head and thorax creamy white tinged with purplish red,
especially the tegule; antennz purplish red; abdomen creamy
white mixed with purplish red and dorsally banded with black
except at base and extremity. Fore wing creamy white mixed
with purplish red, the basal area suifused with black except at
base of inner margin, the costa black,:rather diffused towards
apex; antemedial line defining the black area, creamy white
slightly defined on outer side by purplish-red and black scales,
excurved below costa; the medial part of costa with three white
points ; a round white spot defined by purplish red at upper angle
of cell, another below the lower angle conjoined to a patch of
confluent annuli beyond the lower angle, and another annulus
on vein 1; postmedial line white defined on each side by purplish
red, oblique to discal fold, then slightly waved; cilia black mixed
Ann. & Mag. N. Hist. Ser. 8. Vol, xix, 5
66 Sir G. F. Hampson on new
with some purplish red. Hind wing creamy white mixed with
purplish red, the basal area suffused with black; an oblique
slightly sinuous white antemedial line defined on outer side by
purplish pink and some black scales; a white patch defined by
purplish red and with purplish-red point in centre at lower angle
of cell; postmedial line white defined on each side by purplish
rel and some black scales, waved; an interrupted purplish-red
line with some black scales on it before termen; cilia black
mixed with some purplish red. Underside purplish red; fore
wing with the costa black with white points on it to beyonl
middle ; hind wing with waved white postmedial line defined by
deeper purplish red.
Hab. Br. EB. Arrica, Nairobi (Anderson), 13; Br. C. Arrica,
Mt. Mlanje (Neave). 1 ¢ type. Hap. 16 mm.
(2c) Pyralis trifolialis, sp. n.
3S. Head, thorax, and abdomen white mixed with purplish red,
the antenne and tegule purplish red, the abdomen irrorated with
some blackish scales; sides of frons and palpi blackish, the latter
with the extremities of 2nd and 8rd joints white; fore legs
blackish, the tarsi ringed with white; pectus, mid and hind: legs,
and ventral surface of abdomen white tinged with red-brown.
Fore wing creamy white mixed with purplish red and irrorated
with a few black scales, the terminal area more suffused with
purplish red; antemedial line white defined on each side. by
purplish red, slightly sinuous, a small white spot defined by
purplish red on its outer side at vein 1; the medial part of
costa black with four white points on it; a small white spot
with purplish-red annulus at upper angle of cell, others below
the lower angle of cell and on vein 1, and a trifoliate patch
beyond lower angle of cell; postmedial line white defined on
each side by purplish red, expanding at costa, excurved to near
termen at middle, and ending at tornus; cilia fuscous black with
a fine white line at base. Hind wing with the basal area white
mixed with black and some purplish red, the medial area purplish
red irrorated with black especially towards inner margin, the
terminal area purplish red mixed with whitish and black; a
slightly sinuous white antemedial line defined on each side by
blackish ; a figure-of-eight-shaped white discoidal spot defined
by blackish and with black points in its upper and lower parts ;
postmedial line white defined on each side by blackish, waved,
excurved between veins 6 and 3; cilia fuscous blaek with a
fine white line at base. Underside whitish suffused with red-
brown ; fore wing with the costa black with white points on it
to beyond middle; hind wing with the postmedial line whitish
and indistinct.
Hab. Goup Coast, Kumasi (Sanders), 1 ¢ type. Exp.
9
12 mm, ‘
Pyralidee of the Subfamily Pyraline. G7
.
(2d) Pyralis atrisparsalis, sp. n.
2. Head whitish suffused with purplish pink; thorax and
abdomen purplish pink mixed with some whitish and strongly
irorated with black, the pectus, legs, and ventral surface of
abdomen less strongly irrorated. Fore wing purplish pink mixed
with some whitish and strongly irrorated with black, especially
on basal area except towards costa; antemedial line strong,
whitish defined on each side by black, oblique to submedian
fold, where it is angled outwards, angled inwards at vein 1; a
small blackish discoidal spot; a whitish patch on costal area
towards apex, the whitish subterminal line arising from it,
excurved to vein 3, then incurved; the termen purplish pink.
Hind wing whitish tinged with purplish pink, the terminal half
suffused with fuscous and irrorated with black towards tornus;
a whitish postmedial line, excurved at middle and angled out-
wards at veins 3 and 2, then oblique to tornus; the termen
purplish pink; cilia whitish, mixed with pink and black at tips.
Underside whitish mixed with pink and fuscous; fore wing with
the subterminal line indistinct, except the patch on costal area ;
hind wing with whitish postmedial line excurved at middle.
Hab. N. Nicerta, Zungeru (Macfie), 1 9 type. Exp. 18 mm.
(8a) Pyralis costinotalis, sp. n.
3. Head and thorax pale rufous; abdomen whitish suffused
with red-brown; antenne brownish; palpi and legs whitish suf-
fused with red-brown. Fore wing rufous tinged with purplish
red; antemedial line white defined on outer side by black,
expanding into a wedge-shaped mark at costa, to which it is
slightly incurved; the medial part of costa with alternating
black and white points; a slight blackish discoidal spot; post-
medial line white defined on inner side by blackish, expanding
into a wedge-shaped mark at costa, then excurved and very
slightly waved; a faint maculate brownish terminal band; a fine
whitish line at base of cilia. Hind wing whitish suffused with
rufous to the postmedial line except on costal area, the terminal
area irrorated with brown; an oblique sinuous white antemedial
line, joined at inner margin by the white postmedial line, which
is excurved at middle, then slightly waved; a terminal series of
small brown spots; cilia with a brown line near tips. Underside
whitish suffused with reddish brown; fore wing with series of
whitish and dark brown points on costa to the postmedial line;
both wings with slight blackish discoidal spot and slightly waved
whitish postmedial line defined on inner side by brown and
excurved at middle.
(13 ¢@) Pyralis rufibasalis, sp. n.
¢. Head and thorax red-brown ; abdomen white suffused with
pale olive; legs red-brown; pectus and ventral surface of abdomen
5*
68 Sir G. F. Hampson on new
whitish tinged with red-brown. Fore wing with the basal area
rufous irrorated with red-brown, the rest of wing white tinged
with olive; a small black discoidal spot; the curved postmedial
line indicated by a faint olive shade on its inner side. Hind
wing white tinged with olive; a diffused black patch on basal
area; a small black discoidal spot; a diffused curved olive post-
medial line. Underside white thickly irrorated with black-brown ;
hind wing with small black discoidal spot and diffused curved dark
postmedial line.
Hab. Gotp Coast, Kumasi (Sanders), 13; S. Nigeria,
Tlesha (Humfrey), 1 3 type. xp. 14-16 mm.
(136) Pyralis rosettincta, sp. n.
3. Head, thorax, and abdomen white tinged with pale red-
brown. Fore wing white suffused with pale red-brown except
towards the costa and termen; a curved white antemedial line
with a patch tinged with rose-pink before it except at costa and
inner margin; a slight red-brown discoidal spot; a sinuous white
postmedial line with a rose-pink shade beyond it. Hind wing
white suffused with pale red-brown except at termen; a curved
white antemedial line; a white postmedial line excurved at middle
and above inner margin and with rose-pink shade beyond it.
Underside white tinged with rufous; hind wing with curved white
postmedial line. -
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 1 ¢ type. Hap.
14 mm.
(16d) Pyralis tyrialis, sp. n.
@. Head, thorax, and abdomen brownish ochreous, the head,
thorax, and two basal segments of abdomen suffused with purplish
crimson; palpi, legs, and ventral surface of abdomen brownish
ochreous. Fore wing brownish ochreous suffused with purplish
crimson and slightly irrorated with dark brown; a fine curved
white antemedial line; a small black discoidal spot; a white post-
medial line, excurved at middle; cilia white and blackish. Hind
wing brownish ochreous strongly suffused with purplish crimson
and irrorated with black ; indistinct curved white ante- and post-
medial lines defined by black seales ; a blackish terminal line; cilia
blackish. Underside ochreous suffused and irrorated with brown.
Hab. Gotv Coast, Bibianaha (Spurrell), 1 9 type. Exp.
18 mm.
(16e) Pyralis phenicealis, sp. n.
@. Head, thorax, and abdomen ochreous brown with a crimson
band on 2nd segment of abdomen; palpi with some dark brown at’
sides; fore legs suffused with dark brown. Fore wing silky
ochreous brown ; the costal area irrorated with some dark scales,
the medial part of costa with series of black points and the
terminal part of costa glossy black; two indistinct crimson sub-
Pyralidee of the Subfanily Pyraline. 69
basal lines; a crimson and blackish point in middle of cell and
small discoidal spot; a crimson point at middle of submedian fold
and bar at inner margin; a postmedial crimson point at discal
fold and bar from submediaie fold to inner margin; a curved
diffused crimson subterminal line and a terminal band except on
the black costal area; cilia deep crimson. Hind wing glossy
ochreous brown; a crimson subbasal patch from cell to inner
margin; a discoidal patch with oblique line from it to inner
margin ; a strong postmedial line somewhat excurved at middle ;
a Sean einai band expanding into a patch at costa, and a narrow
band before the ochreous terminal line ; cilia deepcrimson. Under-
side ochreous suffused with fuscous brown: fore wing with some
pale points on medial part of costa and both wings with pale curved
postmedial line.
Hab. Goww Coast, Bibianaha (Spurrell) 1 2 type. Exp.
18 mm.
(19a) Pyralis excumbralis, sp. n.
3. Head, thorax, and abdomen brownish ochreous. Fore wing
ochreous; a rather diffused fuscous patch below the cell; a small
black discoidal spot; a fuscous subterminal shade, not reaching
the costa and narrowing to tornus. Hind wing ochreous; a sub-
basal patch of black irroration, the rest of wing urorated with
fuscous; a curved whitish postmedial line. Warteride ochreous
irrorated with fuscous; fore wing. with blackish discoidal point
and both wings with whitish postmedial line.
Hab. Gord Coast, Bibianaha (Spurrell) 1 S$ type. xp.
16 mm.
(10) Pyralis flavirubralis, sp. n.
3. Head, thorax, and abdomen purplish red mixed with some
yellowish. Fore wing purplish red slightly nrorated with brownish,
the medial area yellow irrorated with red and more suffused with
red towards inner margin ; antemedial line whitish, slightly sinuous
below the cell; a blackish discoidal point; postmedial line whitish,
incurved below discal fold ; cilia yellowish tinged with red. Hind
wing purplish red thickly irrorated with fuscous ; an indistinct
oblique slightly sinuous whitish antemedial line and curved slightly
waved postmedial line; cilia purplish red with a fine white line at
base. Underside ochreous white irrorated with red; both wings
with small blackish discoidal spot.
_ Hab. Transvaat, White R. (Cooke), 1 35 type. Hap. 18mm.
(ld) Pyralis perpulverea, sp. n.
@. Head and thorax whitish tinged with rufous and irrorated
with dark brown; abdomen whitish tinged with rufous; palpi,
pectus, legs, and ventral surface of abdomen rufous, the tarsi dark
brown ringed with white. Fore ‘wing rufous mixed with some
whitish, especially towards inner margin and thickly irrcrated with
70 Sir G. F. Hampson on new
fuscous; a slight blackish discoidal striga; cilia black mixed with
some grey, a pale reddish line at base and some reddish scales at
tips. Hind wing whitish tinged with rufous; cilia rufous with a
fine whitish line at base and dark line near tip except towards
tornus. Underside of fore wing pale fuscous brown, the costal and
terminal areas rufous; hind wing whitish tinged with red-brown,
the apical area rufous.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 1 9 type; Port.
E. Arrica, Mt. Chiperone (Neave),1 9. Hap. 22 mm.
(le) TLegulifera trroralis, sp. n.
@. Head, thorax, and abdomen brownish grey irrorated with
black; the anal tuft rufous; antennze whitish ringed with black.
Fore wing brownish grey tinged with rufous especially towards
costa and irrorated with black; a series of whitish points on costa
with some blackish between them except towards base ; a terminal
series of black bars; cilia fuscous with a fine Avhitish line at base
and blackish line near tips. Hind wing brownish grey tinged with
rufous and irrorated with black; an indistinct pale curved post-
medial line defined on inner side by blackish; a terminal series of
black striz; cilia fuscous mixed with grey, a fine whitish line near
base and blackish line near tips. Underside of fore wing rufous
irrorated with blackish, the inner area whitish; a series of white
points on costa with black between them, a blackish discoidal
striga, a pale subterminal line defined on inner side by blackish,
angled outwards to termen at vein 3; hind wing pale rufous
irrorated with blackish, a black discoidal point and postmedial line
defined on outer side by whitish and excurved at middle.
Hab. W. Arrica (Dudgeon), 1 2 type; S. Nieeria, Lagos
(Sir G. Carter),1 9. Exp. 16 mm.
(26) Tegulifera purpurascens, sp. n.°
3. Head and thorax purplish red with a few fuscous scales;
abdomen ochreous suffused with purplish red and irrorated with
black, the extremity clear ochreous; palpi black at tips; pectus,
legs, and ventral surface of abdomen purplish red irrorated with
black. Fore wing purplish red irrorated with black ; a rather irre-
gularly waved almost medial black line defined on inner side by
diffused ochreous; the medial part of costa with some whitish
points with black between them; postmedial line black defined
on outer side by diffused ochreous, waved, exeurved between
veins 5 and 2 and incurved at submedian fold; a terminal series of
black points. Hind wing ochreous suffused with purplish red and
irrorated with fuscous ; traces of a curved blackish antemedial line
and a distinct curved postmedial line; a terminal series of black
points. Underside ochreous tinged with purplish red ; both wings
with small black discoidal spot and curved postmedial line.
Hab. 8. Nigeria (Sampson), 1 3 type. Exp. 22 mm.
Pyralidee of the Subfamily Pyraline. 71
(2d) Tequlifera eleomesa, sp. n.
$. Head and thorax pale olive-brown, the vertex of head and
tips of patagia tinged with purplish red ; abdomen pale olive-brown
suffused with purplish red towards base and irrorated with some
black scales towards extremity; pectus, legs, and ventral surface
of abdomen whitish suffused with purplish red and irrorated with
black. Fore wing pale olive-brown, the terminal area purplish
red irrorated with black; a curved whitish antemedial line with
a purplish-red patch irrorated with black before it from cell to
inner margin; the medial part of costa with some white points
with black between them; a black discoidal point; a slightly in-
curved white postmedial line with some black irroration before it
below the cell; a fine white line at base of cilia. Hind wing
purplish red irrorated with black ; two curved whitish medial lines,
the area between them suffused with blackish ; a fine white line
at base of cilia. Underside purplish red irrorated with black and
mixed with whitish towards base; both wings with obscure black
discoidal spots; fore wing with the postmedial line indistinct ;
hind wing with slightly waved, white, medial line defined on inner
side by rather diffused black.
. Hab. Goun Coast, Aburi (Johnston), 1 3, Bibianaha (Spur-
rell), 2 $type. Hap. 14 mm. -
(29) Tegulifera obovalis, sp. n.
Head and thorax pale red with a few fuscous scales ; abdomen
pale reddish, the base purplish red; subdorsal black fascize except
at base, connected dorsally on 2nd segment and on two terminal
segments ; pectus, legs, and ventral sur face of abdomen pale reddish.
Fore wing pale rdious slightly irrorated with fuscous, the ovate
terminal area chocolate-brown ; antemedial line whitish defined on
outer side by blackish, rather oblique ; a small black discoidal spot;
the medial part of costa with whitish points with some black
between them; postmedial line whitish defined on inner side by
blackish, ineurved; a fine white line at base of cilia. Hind wing
red-brown with a slight purplish-red tinge ; a curved whitish ante-
medial line defined on outer side by dark brown; a small blackish
spot at upper angle of cell; postmedial line whitish defined on
inner side by dark brown, rather obliquely curved; a fine white
line at base of cilia. Underside whitish tinged with rufous espe-
cially in and beyond the cell of fore wing and on terminal areas of
both wings, the costal areas with some black irroration ; fore wing
with the whitish and black points on costa extending to base, the
postmedial line very slightly waved; hind wing with dark ante-
medial line from cell to inner margin, small discoidal spot and
oblique slightly waved postmedial line.
Hab. Gotp Coast, Kumasi (Whiteside), 1 g, 1 Q type;
Narat, Durban (Leigh), 1 9. Hap. 24-28 mm.
.
72 Sir G. F. Hampson on new
(2h) Tegulifera semicircularis, sp. n.
@. Head, thorax, and abdomen whitish suffused with pale red-
brown. Fore wing whitish suffused with pale olive-brown and
slightly irrorated with black, the semicircular terminal area deep
chocolate-red and defined on inner side by an incurved white shade ;
antemedial line white defined on inner side by brown, rather oblique;
a black discoidal spot; cilia white, tinged eh reddish brown
except at base. Hind wing whitish suffused with pale red-brown ;
two oblique dark medial lines defined by white, the inner line on
inner side, the outer on outer side, the area between them rather
whiter; a fine red-brown terminal line; cilia pale reddish, white at
base and with some dark scales at tips. Underside whitish suffused
with rufous; fore wing with black points on costa to beyond
middle, a black discoidal point, the terminal area purplish red
defined on inner side by an incurved white line; hind wing with
oblique very slightly waved reddish-brown postmedial line.
Hab. Gown Coast, Bibianaha (Spurrell), 1 9 type. Exp.
28 mm.
(27) Tegulifera tripartita, sp. n.
¢. Head whitish tinged with red-brown; thorax red-brown
tinged with grey ; abdomen whitish tinged with red-brown ; legs
dark brown, the tarsi ringed with white. _Fore. wing with the
basal and terminal areas dark red-brown with a orey a gloss, the
medial area pale grey slightly tinged with red-brown and irrorated
with dark brown ; aricenenlel hae white slightly defined on outer
side by brown, exeurved to submedian fold, then incurved; the
media] part of costa with a series of white points with dark brown
between them; a small dark brown discoidal spot; postmedial line
white slightly defined on inner side by brown, slightly incurved
below vein 3; cilia pale red-brown with a fine white line at base
defined on outer side by a dark line. Hind wing greyish with dark
red-brown irroration along vein 1 and on terminal half; an oblique
brown line from upper angle of cell to inner margin ‘at the post-
medial line which is pale ‘defined on each side by ‘brow n, curved ;
cilia pale red-brown with a fine white line at base ‘defined on outer
side by a dark line. Underside of fore wing reddish ochreous
irrorated with brown, the terminal area suffused with red-brown,
the inner area white, the basal area darker to submedian fold, the
costa black-brown with series of prominent white points to the
postmedial line, which is dark defined on outer side by white
forming a small spot at costa, a blackish discoidal spot; hind wing
whitish tinged with rufous and irrorated with red-brown except on
inner area, a dark discoidal spot and curved postmedial line defined
on outer side by whitish.
Hab. Assam, Khasis (Nissary), 3 3 type. Exp. 26 mm.
(36) Tegulifera oehrimesalis, sp. n.
2. Head, thorax, and abdomen yellow, the tegule tinged with
Pyralidze of the Subfamily Pyraline. 73
purplish pink, the abdomen suffused with purplish pink and irro-
rated with black except at extremity; pectus, legs, and ventral
surface of abdomen ochreous yellow. Fore wing ochreous yellow
tinged with purplish pink and slightly irrorated with dark scales,
the medial area and termen almost clear ochreous ; antemedial line
yellow slightly defined on outer side by brownish, curved; a blackish
point at upper angle of cell; postmedial line yellow slightly defined
on inner side by brownish, slightly excurved at middle and incurved
at submedian fold. Hind wing yellowish suffused with purplish
pink and irrorated with blackish ; waved whitish medial and post-
medial lines; a terminal series of small blackish spots except
towards tornus. Underside ochreous tinged with brown; both
wings with indistinct pale sinuous ante- and postmedial lines defined
by brownish ; fore wing with slight dark discoidal spot.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 1 2 type. Hyp.
20 mm.
(38d) Tegulifera pallidalis, sp. n.
3. Head and thorax ochreous tinged with reddish; abdomen
ochreous faintly tinged with purplish red and slightiy irrorated
with brown; fore and mid legs suffused with red-brown. Fore
wing pale ochreous slightly irrorated with brown, the terminal area
tinged with purplish pink ; a series of slight dark points on costa;
a slight dark discoidal spot; a straight pale ochreous postmedial
line defined on inner side by brown and on outer by the pink
terminal area; a terminal series of dark points; cilia brownish
ochreous. Hind wing ochreous white; a rather punctiform brown-
ish terminal line except towards tornus; cilia ochreous, tinged with
brown towards apex. Underside ochreous white; fore wing with
the costal and terminal areas tinged with pinkish, some pale points
on costa towards base, a small brown discoidal spot, the postmedia]
line indistinct, whitish ; hind wing with the costa deeper ochreous,
an oblique brown postmedial line from costa to discal fold.
Hab. Uaanpa, Gondotroro (Reynes-Cole), 1 3 type. Exp.
20 mm.
(4a) Tegulifera bostralis, sp. n.
©. Head and thorax ochreous suffused with red-brown ; abdo-
men ochreous irorated with black-brown; legs. suffused with
red-brown; ventral surface of abdomen tinged with reddish. Fore
wing with the basal and terminal areas red-brown, the medial area
ochreous slightly irrorated with brown, more thickly towards costa;
a pale antemedial line defining the basal area; some pale points
on medial part of costa; a small black discoidal spot; postmedial
line pale, oblique, slightly excurved at middle, then incurved; a
terminal series of small blackish spots and a pale line at base of
cilia. Hind wing ochreous tinged with red-brown, the terminal
half suffused with pale red-brown; an indistinct sinuous dark
medial line defined on outer side by ochreous ; a terminal series of
blackish bars ; cilia red-brown with a pale line at base. Underside
74 Sir G. F. Hampson on new
of fore wing ochreous suffused with purplish red except on inner
area, a series of whitish points on costa with dark brown between
them to the oblique pale postmedial line ; hind wing ochreous, the
costal area and terminal half tinged with purplish red, a small
blackish spot at upper angle of cell and oblique sinuous brown
medial line.
Hab. Br. E. Arrica, Kakumega Forest, Yala R. (Weave),
1 9 type; Transvaat, White R. (Cooke),19. Hap. 26-28 mm.
(Ge) Tegulifera metasarcistis, sp. n.
3. Head and thorax ochreous mixed with dark brown; abdomen
ochreous; antennz brownish; palpi dark brown irrorated with
whitish; legs suffused with dark brown, the tarsi black-brown
ringed with whitish. Fore wing ochreous thickly irrorated with
purplish red and some black except on basal inner area; a pale
postmedial line, exeurved below discal fold; a terminal series of
blackish bars; cilia ochreous tinged with red. Hind wing flesh-
pink ; a small brown subterminal spot at submedian fold ; a black-
ish terminal line except towards tornus. Underside ochreous tinged
with red; hind wing with obliquely curved red postmedial line.
Hab. Gourd Coast, Bibianaha’ (Spurrell), 1 3 type. Lup.
18 mm.
(7a) Tegulifera flavicarnea, sp. n.
3. Head, thorax, and abdomen yellow tinged with reddish, the
palpi, pectus, legs, and ventral surface of abdomen more strongly
tinged with red-brown. Fore wing yellow tinged with purplish
pink, especially on terminal half, and slightly irrorated with brown;
the costa with series of white points with dark brown between
them except towards base; a pale subterminal line, excurved from
below costa to vein 2; a fine white line at base of cilia defined on
its outer side by a black line. Hind wing golden yellow with a
black line at base of cilia. Underside yellow, the costal and
terminal areas tinged with purplish red and the former irrorated °
with dark brown ; fore wing with series of white points on costa
with black between them, a curved white subterminal line defined
on inner side by blackish, a terminal series of small blackish spots ;
hind wing with curved white subterminal line defined on inner side
by blackish.
Hab. Borneo, Sandakan (Pryer), 1 5 type. Hap. 22 mm.
(7b) Tegulifera flaveola, sp. n.
2. Head, thorax, and abdomen yellowish suffused with purplish
red ; palpi dull purplish red. Fore wing yellowish suffused with
dull purplish red and irrorated with blackish scales, the area from
middle of wing to the postmedial line more strongly suffused ; a
faint dark discoidal spot; the postmedial line indistinct, excurved
at middle and ineurved below vein 2; a terminal series of small
blackish spots. Hind wing yellowish, suffused and irrorated with
Pyralidee of the Subfamily Pyraline. 75
dark brown to the indistinct curved postmedial line, the terminal
area very slightly irrorated; a terminal series of small dark brown
spots; cilia with a dark brown line through them. Underside
yellow; fore wing tinged and irrorated with brown to the post-
medial line, the terminal area slightly irrorated, more strongly
towards costa; hind wing irrorated with brown to the indistinct
irregular postmedial line, the terminal area sparsely irrorated from
costa to vein 2. bi
Ab. 1. Wings uniformly suffused with red and irrorated with
blackish; fore wing with the postmedial line hardly traceable ;
hind wing with it indistinct; the underside uniformly suffused
with red and irrorated with black, both wings with curved slightly
waved blackish postmedial line.
Hab. Cameroons, Ja R., Bitje (Sates), 3 2 type. Hap. 20-
24 mm.
(7c) Teqgulifera chromalis, sp. n.
6. Head, thorax, and abdomen golden yellow suffused with
purplish red; palpi yellow tinged with purplish red in front towards
base ; fore coxze and mid femora towards base deep purple, the
fore and mid tibiz black-brown, the tarsi black-brown ringed with
whitish. Fore wing golden yellow, the basal area to just below
the cell purplish red, the apical area from middle of costa to termen
at vein | suffused and irrorated with purplish red leaving a conical
almost clear yellow patch from postmedial part of costa to below
vein 5, the inner area irrorated with a few red scales; some yellow
points on medial part of costa; the antemedial line represented by
a yellow bar from costa to median nervure; the postmedial line
faint and excurved from vein 6 to 2, then incurved; cilia glossy
black-brown. Hind wing golden yellow irrorated with purplish
red to the postmedial line and on terminal area from apex to
vein 4; an oblique curved red antemedial line joined at inner
margin by the curved slightly waved postmedial line ; cilia glossy
black-brown except towards tornus. Underside yellow ; fore wing
more evenly irrorated with red, the costa deep purplish red with
pale points on it to the indistinct curved yellow postmedial line ;
hind wing with the costal area irrorated with red, a faint curved
postmedial line formed by red scales.
Hab. Cameroons, Ja R., Bitje (Bates), 1 S type. Lap.
30 mm.
(7d) Tequlifera ochrealis, sp. n.
Q. Orange-yellow. Fore wing with faint traces of curved post-
medial line. Hind wing rather paler.
Hab. MasnonatanD (Dobbie), 1 2 type. Exp. 20 mm,
(9a) Tegulifera conisalis, sp. un.
3. Head, thorax, and abdomen greyish suffused with mid
reddish brown ; fore tarsi dark brown ringed with whitish; dull
76 Sir G. F. Hampson on new
and hind tarsi whitish. Fore wing greyish tinged with red-brown
and thickly irrorated with dark red-brown; an oblique whitish
antemedial line defined on outer side by diffused dark brown; some
whitish points on medial part of costa with dark brown between
them ; a small dark brown discoidal spot; postmedial line whitish
defined on inner side by dark brown, slightly waved and curved to
vein 2 and incurved at submedian fold; a terminal series of small
dark brown spots and whitish line at base of cilia. Hind wing
whitish tinged and irrorated with brown ; a terminal series of snia i
dark brown spots. Underside whitish tinged and irrorated with
purplish brown, the inner areas paler ; both wings with small dark
discoidal spot and curved postmedial line; fore wing with the
costa dark brown with white points on it to the eles: line.
Hab. Germ. E. Arrica, Dar-es-Salaam, 1 ¢ type. zp.
16 mm.
(2) Elealis metachalecistis, sp. n.
3. Head, thorax, and abdomen dark red-brown; antennz whitish
ringed with brown; fore tarsi ringed with whitish, the mid and
hind tarsi whitish tmged with red-brown. Fore wing dark red-
brown with a cupreous gloss ; a series of whitish points ¢ on costa to
beyond middle and a postmedial whitish spot tinged with reddish.
Hind wing golden cupreous irrorated with dark red- brown, the
costal and. terminal areas dark red-brown, the latter narrowing to
tornus; the underside reddish ochreous, the basal part of costal area
and cell mottled with reddish ochreous defining a dark brown
discoidal spot, the terminal area dark brown narrowing to tornus.
@. Fore wing with narrow whitish postmedial band tinged with
reddish.
Hiab. Br. E. Arrica, Kikuyu Escarpment, Ibea (Doherty), 2 3,
1 9 type. Exp. 18-20 mm.
(la) Stemmatophora albiceps, sp. n.
Antenne of male with the basal joint very long.
Head ochreous white, the antenne dark brown except the basal
joint, the palpi with dark brown spot at side of 2nd joint, the
8rd dark brown with white tips; tegule ochreous white irrorated
with some dark brown scales and dark brown at sides; thorax
whitish, the patagia dark brown at sides; abdomen reddish brown
tinged with grey, the anal tuft ochreous; pectus, legs, and ventral
surface of abdomen dark brown tinged. with grey. Fore wing
dark reddish brown; triangular white ante- and postmedial patches
on costa with faint slightly curved whitish lines from them to
inner margin and two white points between them on costa; cilia
with a fine white line at base and some whitish at tips. Hind wing
dark reddish brown with curved whitish ante- and postmedial lines;
cilia with a slight whitish line at base. Underside fuscous brown ;
fore wing with the inner area whitish, a series of ochreous-white
points on costa to an ochreous-white postmedial patch with slight
Pyralids ofthe Subfamily Pyraline. 7
line from it to inner margin; hind wing with oblique whitish
postmedial line defined on inner side by darker brown.
Hab. N. Nicerts, Minna (Macfie), 1 $ type, Zungeru (Macfie),
id, . oo Bap; 6 14, 9 16 mm,
(2a) Stemmatophora oleoalbalis, sp. n.
3. Head and thorax white with a faint brownish tinge; abdo-
men whitish tinged with red-brown and irrorated with dark brown
scales; antennz ringed with brown; pectus and legs suffused with
red-brown, the tarsi dark brown ringed with white; ventral surface
of abdomen dark brown towards extremity. Fore wing white
tinged with pale olive and irrorated with a few black scales; a
slight black mark at base of costa; the medial area black with
white points on costa and defined by the diffused white ante- and
postmedial lines, the former nearly straight, the latter strongly
excurved at middle, then incurved, a wedge-shaped rufous patch
beyond it on costa; a terminal series of faint black points. Hind
wing white with a faint brownish tinge; a faint curved dark post-
mediai line; a terminal series of black points except towards
tornus. Underside whitish tinged with rufous; fore wing with
whitish points with black between them to the postmedial line,
the medial area suffused with blackish; hind wing with slight
dark point at upper angle of cell and rather diffused black post-
medial line defined on outer side by white and excurved at middle.
Hab. Br. HE. Arnica, Nairobi (Anderson), 1 & type. Exp.
20 mm.
(26) Stemmatophora chloralis, sp. n.
Stemmatophora chloralis, Longstaff, Butterfly Hunting in Many Lands,
pl. ii. fig. 9.
2. Head white; antennz with the extreme base of shaft black ;
thorax white tinged with very pale blue-green; pectus, legs, and
abdomen white irrorated with a few black scales, the tarsi slightly
ringed with black. Fore wing white very finely pencilled with
pale blue-green ; a black striga trom base of costa; a black point
on middle of costa ; an oblique black band, defined on each side by
rather diffused white from costa just beyond middle to inner
margin, with some white points on it at costa, expanding into a
large elliptical black patch in and beyond the cell, then narrowing
and again slightly expanding to inner margin; a terminal series of
slight black points with a more prominent point above tornus.
Hind wing white, the terminal area slightly irrorated with black
scales, extending on costa to middle and narrowing to tornus; a
slight fuscous mark at lower angle of cell; a terminal ‘series of
small black spots from apex to submedian fold. Underside of fore
wing with prominent series of black strize on costa from base to
the postmedial band which is obsolescent.
Hab. Zampest, Victoria Falls (Longstaf’), 1 2 type, ¢ in
Coll. Longstaff. Kap. 24 mm.
,
78 Sir G. F. Hampsoii on new
(6c) Stemmatophora cupricolor, sp. n.
; 3. Head, thorax, and abdomen pale red with a whitish tinge,
the pectus, legs, and ‘ventral surface of abdomen whiter. Fore wing
cupreous red ‘slightly irrorated with black; a whitish postmedial
line, oblique to vein 6, then sinuous; a fusenus terminal line ; cilia
fuscous mixed with whitish and with black line near base. Hind
wing fiery red irrorated with blackish; a blackish antemedial line,
oblique to submedian fold, where there is a white patch before it,
then sinuous and defined on inner side by whitish ; a curved white
postmedial line slightly defined on inner side by blackish; cilia
fuscous mixed with whitish and with black line near base. Under-
side of fore wing grey-brown, a slight dark postmedial line defined
on outer side by whitish, oblique to vein 5, then slightly incurved ;
hind wing brownish white slightly irrorated with brown, a slight
curved brown postmedial line.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 1 3 type.- Exp.
26 mm.
(6d) Stemmatophora olivotincta, sp. n.
3. Head, thorax, and abdomen whitish tinged with olive-brown;
pectus, legs, and ventral surface of abdomen tinged with crimson.
Fore wing whitish suffused with pale olive-brown; a slightly
curved white antemedial line ; the medial part of costa with series
of white points with dark brown between them; a straight erect
white postmedial line; a slight crimson terminal line ; cilia bright
crimson with a fine white line at base. Hind wing whitish suffused
with pale olive-brown ; traces of a curved white antemedial line
and a more distinct postmedial line ; a slight crimson terminal line;
cilia bright crimson with a fine white line at base. Underside
white tinged with rufous ; fore wing with large patch of crimson
suffusion in and beyond the cell and below lower end of cell with
slight crimson suffusion beyond it between veins 5 and 2; hind
wing with crimson antemedial line oblique to median nervure, the
medial part of costa and a spot at upper angle of cell crimson, a
strong rather diffused crimson postmedial line, the terminal area
tinged with crimson towards apex and in submedian interspace.
flab. Cameroons, Ja R., Bitje (Bates), 1 9 type. Hap.
24 mm.
(6f) Stemmatophora hemicyclalis, sp. n.
Head and thorax ochreous tinged with rufous and irrorated with
a few dark brown scales; abdomen ochreous white irrorated with
dark brown scales forming diffused dorsal bands except towards
base ; palpi and fore legs dark reddish brown, the latter with the
tarsi ringed with white. Fore wing ochreous whitish irrorated
with dark brown, the basal area suffused with red-brown, the costa
dark brown with slight whitish points to beyond middle ; a minute
black discoidal spot; postmedial line, white, incurved from costa -
towards apex to tornus, the semicircular ter minal area suffused with
Pyralide of the Subfamily Pyraline. 79
dark brown shading to red-brown at termen; a terminal series of
black points; cilia greyish suffused with brown. Hind wing white
irrorated with brown except on terminal area from apex to vein 3
which is faintly tinged with rufous; an indistinct obliquely curved
dark postmedial line ; a terminal series of small black spots ; cilia
tinged with brown and with a brownish line near base. Underside
whitish ; fore wing irrorated with brown especially on basal half,
an indistinct erect dark postmedial line; hind wing with the costal
area irrorated with brown, an indistinct obliquely curved dark
postmedial line.
Hab. Transvaat, White R. (Cooke), 13, 12 type. up.,
6 20, 9 22 mm,
(11 a) Stemmatophora perrubralis, sp. n.
2. Head and thorax fiery rufous; abdomen yellowish tinged
with rufous, the ventral surface deeper rufous. Fore wing fiery
rufous slightly irrorated with dark brown; antemedial line pale
slightly defined on outer side by black scales, rather oblique; a
small black discoidal spot ; postmedial line whitish slightly defined
on inner side by dark scales, almost straight and erect; a slight
dark terminal line and whitish line at base of cilia which are brown
and whitish at tips. Hind wing yellowish suffused with fiery red ;
a curved whitish postmedial line slightly defined on inner side by
red; a fine whitish line at base of cilia. Underside yellowish
suffused with fiery red; both wings with faint red postmedial line
defined on outer side by whitish.
Hab. Lourengo Maregues, Shilouvane (Junod), 2 2 type.
Hep. 28 rom.
(12 a) Stemmatophora minimalis, sp. n.
Head, thorax, and abdomen whitish suffused with pale red-
brown; pectus, legs, and ventral surface of abdomen whitish
irrorated with dark brown. Fore wing whitish suffused with pale
red-brown and irrorated with blackish; the costa with slight
whitish points with blackish between them to the postmedial line ;
a curved whitish antemedial line; a faint dark medial line, slightly
excurved to submedian fold, then incurved; postmedial line whitish
slightly defined on inner side by dark brown and slightly curved ;
cilia brown with pure white tips. Hind wing whitish suffused
with pale red-brown and irrorated with blackish ; an oblique whitish
antemedial line curved inwards to costa; a straight white post-
medial line; cilia brown, pure white at tips. Underside whitish
suffused with reddish and irorated with brown; both wings with
curved white postmedial line.
Hab. Crynton, Trincomali (Green), 1 5, 19 type. Hap.,
6 12, 2 14mm.
80 pir Ge Hampson on new
(12,f) Stemmatophora excurvalis, sp. n.
Q@. Head, thorax, and abdomen whitish suffused with pale red-
brown and irrorated with dark brown, the last with blackish dorsal
bands on two medial segments. Fore wing whitish suffused with
pale red-brown and irrorated with dark brown; a narrow inwardly
oblique whitish antemedial band; the medial part of costa with
slight whitish points with dark brown between them; a faint
blackish discoidal spot; postmedial line whitish slightly defined on
inner side by fuscous, slightly incurved to discal fold, then strongly
excurved to vein 2, then incurved; cilia with a white line at base,
the tips fuscous and white. Hind wing whitish tinged with pale
red-brown and irrorated with brown, the apical area more suffused
with brown ; a diffused curved whitish postmedial line ; a terminal
series of slight dark spots; cilia white with dark lines near base
and tips. Underside white tinged with red-brown and irrorated
with dark brown; fore wing thickly irrorated except on inner
area, the costa with whitish points with dark brown between them
to the postmedial line ; a small blackish discoidal spot; hind wing
with small black discoidal spot and rather diffused blackish post-
medial line defined on outer side by whitish and excurved at
middle.
Hab. Br. E. Arnica, Nairobi (Anderson), 1 2 type. Hap.
20mm. .
(129) Stemmatophora postaurantia, sp. n.
Head, thorax, and abdomen whitish tinged with rufous, the last
irrorated with black on terminal half; antennz slightly ringed with
black. Fore wing whitish tinged with pale rufous and irrorated
with black; a patch of black irroration at base of costal area;
- antemedial line black, diffused, slightly excurved at submedian
fold; the medial part of costa black with white points on it; a
black discoidal spot; postmedial line black, diffused, slightly in-
curved at discal fold and angled inwards at submedian fold; a
patch of black irroration on costal area towards apex; a terminal
series of small black spots. Hind wing reddish orange with a
terminal series of small black spots. Underside of fore wing
orange-red, the costal and inner areas whitish tinged with olive-
brown, the costa with series of whitish points with some black
between them to the diffused black postmedial line, slightly
incurved at discal and submedian folds, a black discoidal spot;
hind wing orange-red, the costal and terminal areas irrorated with
a few blackish scales, an indistinct oblique postmedial line formed
by blackish scales.
Hab. Br. C. Arnica, Mt. Mlanje (Weave), 1 3, 1 & type.
Hxp.20 mn.
(18 a) Stemmatophora erebalis, sp. n.
Head, thorax, and abdomen red-brown largely mixed with black ;
tarsi pale. Fore wing reddish brown very thickly irrorated with
as
Pyralide of the Subfamily Pyraline. 81
black; a pale waved antemedial line; the medial area with series
of black and pale points on costa; a black discoidal spot; a pale
minutely waved postmedial line defined by black on inner side and
excurved at middle ; a terminal series of small black spots; cilia
reddish with dark lines at middle and tips. Hind wing fuscous
with indistinct blackish discoidal spot; a pale curved postmedial
line; a fine black terminal line; cilia pale with diffused dark line
through them.
Hab. Goup Coast, Ajinak (Dudgeon) 2 3,1 2 type; N. Nr-
GERIA, Minna (Macfie), 3 ¢,1 2, Zungeru (Macfie), Fe Oe.
Bida (Mactie),3 2 ; MasHonaLanD (Dobbie), EY. Rep} & 20;
© 26 mm.
(156) Stemmatophora fusilinealis, sp. n.
Head, thorax, and abdomen ochreous suffused with rufous, the
terminal half of abdomen with black strongly mixed; fore legs
black-brown, the tarsi ringed with white; mid legs suffused with
black-brown, the tarsi whitish ringed with black. Fore wing
ochreous suffused with cupreous red and slightly irrorated with
black ; a diffused curved black antemedial line ; the medial part of
costa with series of white points with black between them; a small
black discoidal spot; a diffused black postmedial line, angled in-
wards at discal and submedian folds and with its outer edge
minutely dentate; traces of a waved subterminal line formed by
black scales; a terminal series of black striz; cilia with blackish
lines near base and tips. Hind wing ochreous suffused with
cupreous red ; an indistinct curved slightly waved dark postmedial
line; a terminal series of black striz ; cilia with blackish lines near
base and tips. Underside ochreous suffused with cupreous red ;
fore wing with series of whitish points with blackish between them
to the postmedial line, the other markings as above.
Hab. Br. C, Arrica, Mt. Mlanje (Veave), 3 bd, 1 Q type.
Exp. 22 mm.
(5a) Herculia roseotincta, sp. n.
Head, thorax, and abdomen white tinged with pinkish brown;
palpi and legs suffused with red-brown. Fore wing whitish suf-
fused with brownish pink and faintly irrorated with brownish, the
costal edge whiter; antemedial line white, angled outwards below
costa, then oblique; a slight brownish discoidal spot; postmedial
line white, oblique; a whitish line at base of cilia. Hind wing
white, the terminal half tinged with pink except towards tornus; a
faint white postmedial line slightly defined on inner side by pink;
a terminal series of pink points to submedian fold; the cilia tinged
with pink and with whitish line at base to submedian fold. Under-
side of fore wing rose-red, the inner area white, the costa with
series of white points with brown between them to the obliquely
curved white postmedial line; hind wing white, the costal area
suffused and irrorated with pink, a slight pinkish diseoidal point, a
Ann, & Mag. N. Hist. Ser. 8. Vol. xix. 6
82 Sir G. F. Hampson on new
eurved white postmedial line slightly defined on inner side by pink
from costa to submedian fold.
Hab. Transvaat, White R. (Cooke), 1 2, Pretoria (Distant,
Janse),2 5,49 type. Hap. 22-24 mm.
(15a) Herculia plumbeoprunalis, sp. n.
Head and thorax yellowish suffused with purplish red; thorax
and abdomen greyish suffused with purplish red; pectus, legs, and
ventral surface of abdomen purplish red. Fore wing reddish brown
with a leaden-grey gloss, the costal area yellow suffused with
purplish red; a slightly curved brown antemedial line defined on
inner side by yellowish; a series of yellow points on medial part
of costa with dark red-brown between them; a slight brown
discoidal striga; the postmedial line almost subterminal, with a
yellow bar from costa to vein 6, then a slight pale line excurved to
vein 2; cilia yellow, deep red at base and apex. Hind wing
reddish brown glossed with leaden grey; a faint pale curved post-
medial line; cilia yellow, deep red at base. Underside whitish
tinged with brown ; fore wing with the costal area yellow tinged
with red and with yellow points*on costa with brown between
them to the postmedial line; hind wing with curved brown post-
medial line.
Hab. W. Cotompta, R. Jiminez, 1 9, R. Dagua, 1 9 ; VENE-
ZUELA, Esteban Valley, Las Quiguas,2 9 ; Ecuapor, R. Pastaza,
Alpayacu (Palmer),1 3. Hap. 22-30 mm.
(226) Herculia perpulverea, sp. n.
@. Head, thorax, and abdomen brownish ochreous irrorated with
dark brown; pectus, legs, and ventral surface of abdomen pale
red-brown. Fore wing brownish ochreous thickly irrorated with
dark brown; traces of a curved brownish antemedial line defined
on inner side by diffused ochreous; postmedial line indistinct,
brown defined on outer side by ochreous, strongly excurved; a fine
pale line at base of cilia. Hind wing brownish ochreous thickly
irrorated with dark brown ; a faint curved dark postmedial line; a
fine pale line at base of cilia. Underside ochreous whitish tinged
and irrorated with red-brown; both wings with faint curved brown
postmedial line.
Hab. Goup Coast, Kumasi (Sanders), 1 2 type. Hap.
24 mm.
(246) Herculia griseobrunnea, sp. n.
3. Head, thorax, and abdomen purplish brown mixed with grey ;
antenne whitish tinged with brown; mid tarsi and hind legs
whitish. Fore wing purplish brown irrorated with grey; ante-
medial line white, excurved to submedian fold; the medial part
of costa with white points with dark brown between them ; post-
medial line white, expanding at costa, then slightly waved and
Pyralide of the Subfamily Pyraline. 83
excurved at middle ; a fine white line at base of cilia followed by a
brown line. Hind wing whitish suffused and irrorated with purple-
brown; a curved white postmedial line; a fine white line at base
of cilia followed by a brown line. Underside white thickly irro-
rated with brown; fore wing with white points with dark brown
between them on costa to the postmedial line.
Hab. Transvaat, Groenvlei (Janse), 1 3, Merwe (Janse), 1 ¢
type, Pretoria (Janse), 1 3. Exp. 18 mn.
(25 a) Herculia purpureorufa, sp. n.
2. Head, thorax, and abdomen greyish suffused with purple-red ;
hind legs whitish. Fore wing deep purplish red; a curved white
antemedial line expanding into a patch on costa; two white points
on medial part of costa; a white postmedial bar from costa to
vein 6, then a fine line excurved at middle and above inner margin
and incurved at submedian fold; a white line at base of cilia.
Hind wing deep purple-red irrorated with a few dark brown scales ;
an obliquely curved white antemedial line joined above inner margin
by a similar postmedial line and both slightly exeurved just below
submedian fold; a fine white line at base of cilia followed by a dark
brown line. Underside purple-red thickly irrorated with dark
brown; fore wing with white points on costa to the postmedial
line; hind wing with curved white postmedial line.
Hab. Mapras, Belgaum (Watson), 1 2 type. Exp. 20 mm.
(266) Herculia pyrerythra, sp. n.
3. Head, thorax, and abdomen purplish red, the anal tuft
yellowish; legs irrorated with black. Fore wing deep purple-red
urorated with black, the medial area more thickly irrorated except
towards costa; a curved whitish antemedial line; the medial part
of costal area paler with slight white points with black between
them on the costa; postmedial line whitish, expanding at costa,
excurved at middle and above inner margin; cilia fuscous brown
with fine white line at base and diffused whitish line at middle.
Hind wing deep purple-red irrorated with black, the medial area
more thickly irrorated; an oblique white antemedial line and white -
postmedial line excurved at middle and above inner margin; cilia
fuscous brown with a fine white line at base and more diffused line
at middle. Underside purple-red thickly irrorated with black ;
fore wing with white points on costa to the postmedial line; hind
wing with the postmedial line defined on inner side by black.
Hab. N. Nigeria, Zungeru (Simpson), 1 3 type. Exp.
16 mm.
(26 c) Herculia lacteocilia, sp. n.
Head, thorax, and abdomen greyish suffused with pale purplish
red, the last irrorated with a few dark scales; fore legs dark brown,
the tarsi ringed with whitish. Fore wing greyish tinged with
6*
84 Sir G. F. Hampson on new
purplish red and irrorated with blackish, the terminal area suffused
with fuscous brown; traces of a pale curved antemedial line; the
medial part of costa with whitish points with black-brown between
them, and with the costal area whiter; postmedial line whitish,
oblique ; cilia black-brown at base, pale yellow at tips. Hind wing
purplish red irrorated with blackish; oblique slightly sinuous
whitish ante- and postmedial lines approximated at inner margin ;
cilia black-brown at base, pale yellow at tips. Underside whitish
suffused with purplish red and irrorated with black; fore wing
with white points on costa to the postmedial line; hind wing with
oblique white postmedial line.
Ab. 1. Hind wing brighter purplish red; cilia of both wings
pure white at tips.
Hab. Ucanpa, Toro, Mpanga Forest (Neave),1 3,1 2 type;
Br. C. Arrica, Mt. Mlanje (Weave),1d. Exp.,d 18, 2 20mm.
(82 6) Hereulia perrubralis, sp. n.
©. Head, thorax, and abdomen purplish red; mid and hind
tarsi whitish. Fore wing deep purple-red; antemedial line black
defined ou inner side by whitish, rather oblique; some pale points
on medial part of costa with black between them; a small black
discoidal spot; postmedial line black defined on outer side by
whitish, rather oblique; a terminal series of blackish points and
fine whitish line at base of cilia. Hind wing deep purple-red; an
oblique blackish antemedial line and similar postmedial line defined
on outer side by whitish; a terminal series of slight dark points
and fine whitish line at base of cilia. Underside purplish red;
both wings with small black discoidal spots and oblique postmedial
line ; fore wing with whitish points on costa to the postmedial line ;
hind wing with the costal area irrorated with blackish.
Hab. 8S. Nigeria, Itu (Furquahar), 19 type. Hap. 28 mm.
(84a) Herculia castaneorufa, sp. n.
Head, thorax, and abdomen chestnut-red;° antenne whitish
tinged with red. Fore wing chestnut-red; traces of a whitish
antemedial line; the medial part of costa with yellow points with
dark brown between them; postmedial line yellow, slightly curved,
dilated at costa; cilia glossy fuscous brown. Hind wing chestnut-
red with a curved yellowish postmedial line; cilia glossy fuscous
brown. Underside yellowish tinged with red and irrorated with
brown ; fore wing with yellowish points with dark brown between
them on costa to beyond middle; hind wing with oblique dark
brown medial line.
Hab. Cameroons, Ja R., Bitje (Bates),2 6,12 type. EHuzp.,
‘go 24, 9 28 mm.
(346) Hereulia flavirufalis, sp. n.
Head, thorax, and abdomen yellowish tinged with red. Fore
“1
i
Pyralidze of the Subfamily Pyralince. 85
wing yellow tinged with fiery red and slightly irrorated with
brown; traces of a yellow antemedial line; the medial part of
costa with yellow points with black-brown between them; a yellow
postimedial line faintly defined on inner side by brown, slightly
excurved at middle; a terminal series of slight brown points; cilia
glossy fuscous brown with a fine yellow line at base. Hind wing
yellow suffused with fiery red; indistinct curved dark ante- and
postmedial lines defined by whitish, the former on inner side, the
latter on outer; cilia glossy fuscous brown. Underside yellow
tinged with red; fore wing with dark discoidal point and yellowish
points on costa with blackish between them to the dark postmedial
line defined on outer side by whitish; hind wing with faint curved
dark postmedial line.
Gee CameEnroons, Ja R., Bitje (Bates), 15,1 2 type. Hap.
22 min.
(84¢) Herculia ecrhodalis, sp. n.
6. Head and thorax pale purplish red; abdomen whitish irro-
rated with purplish red; antennz whitish; pectus and legs red-
brown; abdomen whitish tinged with red-brown. Fore wing
whitish tinged with red and irrorated with purplish red, the
terminal area suffused with purplish red; traces of a whitsh ante-
medial line; the medial part of costa with whitish points with dark
brown between them; postmedial line whitish, slightly excurved at
middle. Hind wing whitish suffused with purple-red; a curved
whitish postmedial line; cilia with a whitish line at base. Under-
side whitish suffused with red-brown; fore wing with whitish
points with dark brown between them on costa to the faint pale
postmedial line; hind wing with curved whitish postmedial line.
Hab. Cameroons, Ja R., Bitje (Bates), 2 ¢ type. Hap.
18 mm.
(84d) Herculia echrunnealis, sp. n.
@. Head, thorax, and abdomen greyish tinged with red-brown,
the pectus and ventral surface of abdomen redder, the legs brownish
with the tarsi ringed with whitish. Fore wing greyish tinged
with olive-brown, the terminal area browner; traces of a whitish
antemedial line; the medial part of costa with whitish points with
dark brown between them; a small dark brown discoidal spot;
postmedial line indistinct, dark brown defined on outer side by
whitish, excurved at middle, a terminal series of black-brown points
and tine whitish line at base of cilia, @hich are grey-brown. Hind
wing greyish tinged with olive-brown, the terminal area browner ;
a curved brown postmedial line defined on outer side by whitish ;
a brown terminal line and fine white line at base of cilia, which
are grey-brown. Underside ochreous suifused with rufous and
irrorated with brown; fore wing with small blackish discoidal
spot and whitish points with blackish between them on costa to
the postmedial line; hind wing with dark discoidal point and
curved postmedial line.
Bore. Sir G. F. Hampson on new
Hab. Cameroons, Ja R., Bitje (Bates), 1 2 type: Exp.
18 nm.
(11) Triphassa trichotibialis, n. n.
Triphassa bilinea, Hmpsn. Moths Ind. iv. p. 166 (nec Moore).
Hab. CErYLon.
(1a) Saecada papuana, sp. nu.
3. Head, thorax, and abdomen grey mixed with chocolate-
brown, the tegule mostly chocolate-brown; pectus in front and
the fringes of hair on fore legs more chocolate-red. Fore wing
grey irrorated with chocolate-brown, the basal area from costa to
vein 1 chocolate-brown with some fiery rufous in submedian inter-
space; antemedial line grey, oblique to submedian fold, then
inwardly oblique; a reddish-brown discoidal spot defined by grey ;
postmedial line grey, oblique below vein 4, a broad chocolate-brown
shade beyond it; cilia dark brown mixed with grey. Hind wing
purplish grey suffused with brown. Underside of fore wing
purplish red, the inner area grey, the postmedial line indistinct,
whitish ; hind wing purplish grey, the costal area suffused with red,
an indistinct curved whitish postmedial line ending at tornus.
Hab. Br. N. Guryea, Dinawa: (Pratt), 1 d, Ekeikei (Pratt),
1g type. Hap. 44-48 mm.
(3¢) Sacada erythropis, sp. n.
@. Head, thorax, and abdomen purplish pink mixed with red-
brown; pectus, legs, and ventral surface of abdomen chestnut-red.
Fore wing purplish pink slightly irrorated with brown; a large
elliptical crimson-red patch from below costa to above inner
margin before the strongly curved fuscous antemedial line; a
discoidal bar formed by fiery red and black-brown scales with a
_ pale striga in centre; a diffused obliquely curved rufous line beyond
the cell; postmedial line fuscous slightly defined on outer side by
whitish, rather oblique to vein 5, then inwardly oblique, a fiery
rufous shade beyond it and a chocolate-brown patch between veins
7 and 4; cilia black-brown mixed with red and with a fine whitish
line at base. Hind wing dark reddish brown to the indistinct
curved postmedial line, then purplish red irrorated with brown; a
fine whitish line at base of cilia. Underside of both wings dark
brown to the curved black-brown postmedial line defined on outer
side by white towards costa of fore wing, the terminal areas purple-
red.
Hab. S. NicErta, Dorin (Macfie), 1 2 type. Exp. 30 mm.
(5b) Sacada albioculals, sp. n.
3. Head and thorax greyish mixed with red-brown, the patagia
dark red-brown except at base; abdomen greyish suffused with
red-brown ; antennz red-brown ; pectus, legs, and ventral surface
Pyralidee of the Subfamily Pyraline. 87
of abdomen bright red-brown. Fore wing red-brown mixed with
greyish, the basal part of inner area and the medial area darker
greyish brown; a large fiery-red patch below the cell before the
antemedial line, which is white and strongly excurved from discal
fold to inner margin ; a white discoidal bar with its lower extremity
slightly angled outwards and a small black-brown spot on its lower
part; postmedial line white, rather oblique to vein 5, then inwardly
oblique and sinuous to inner margin, where it is approximated to
the antemedial line, some fiery red suffusion beyondit. Hind wing
whitish suffused with pale reddish ; a faint curved dark postmedial
line. Underside whitish suffused with pale reddish; fore wing
with faint dark postmedial line oblique and sinuous below vein 5;
hind wing with faint curved dark postmedial line.
Ab. 1. Fore wing with the ante- and postmedial lines confluent
at vein 1 and not reaching inner margin.
Hab. Dutcu N. Guinea, Fak-fak (Pratt), 2 5 type. Ezp.
30 min.
(9a) Sacada nyasana, sp. n.
@. Head, thorax, and abdomen purple-brown; the hind tarsi
with the Ist joint whitish towards base and the other joints ringed
with whitish. Fore wing purple; a strongly curved fuscous ante-
medial line with a broad chocolate-brown shade before it; a deep
chocolate-brown discoidal spot with a whitish bar in centre, a
chocolate-brown shade beyond the cell, oblique below vein 4; post-
medial line fuscous slightly defined on outer side by grey, rather
oblique to vein 4, then inwardly oblique, the apical area beyond it
chocolate-brown, its lower edge running obliquely to termen at
vein 4, and a chocolate-brown shade beyond the postmedial line
from vein 4 to inner margin; cilia chocolate-brown. Hind wing
pale purple-brown. Underside purple; fore wing with fuscous
postmedial line defined on outer side by white towards costa and
oblique below vein 4; hind wing with fuscous postmedial line
excurved at middle.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 1 2 type. Hap.
42 mm.
(13) Sacada viridalis, sp. n.
@. Head and thorax dull apple-green ; abdomen grey suffused
with brown; antenne grey-brown; pectus, legs, and ventral
surface of abdomen white tinged with brown. Fore wing dull
apple-green, the costal edge red-brown with a series of white points
on medial area; antemedial line red-brown, sinuous to median
nervure, where it is angled outwards, then oblique ; postmedial line
red-brown, incurved below costa, excurved to vein 4, then incurved
and very ‘slightly waved; cilia pale purplish brown with a fine
white line at base. Hind wing dull apple-green, the costal area
purple; an oblique purple-brown antemedial line and slightly
sinuous postmedial line; cilia pale purplish with a fine white line
88 Sir G. F. Hampson on new
at base followed by a dark line. Underside whitish suffused with
purple except on inner area; fore wing with the costa red-brown
with white points on it to the dark postmedial line, which is ex-
curved at middle, a dark discoidal point; hind wing with dark
discoidal point and curved postmedial line slightly waved to
vein 2.
Hab. Cameroons, Ja R., Bitje (Bates), 1 2 type. Eup.
30 min.
(2a) Paractenia pallidirubra, sp. n.
3g. Head and thorax whitish suffused with pale purplish red;
abdomen white. Fore wing whitish suffused with pale purplish
red, the costa rather darker ; a small blackish discoidal spot; post-
medial line indistinct, fone rather diffused on inner side and
minutely dentate on outer, slightly excurved to vein 2 and in-
eurved at submedian fold; a Serine series of slight blackish
spots. Hind wing white faintly tinged with red- brown ; ; a rather
punctiform red- brown terminal line. Underside white, the fore
wing and costal area of hind wing faintly tinged with red; fore
wing with slight dark discoidal spot and obliquely curved ‘post-
medial line; hind wing with faint discoidal point and postmedial
striga from costa.
Hab. BompBay, Deesa (Nurse), 1 3 type. Hxp. 24 mm.
(8a) Paractenia viridicostalis, sp. n.
Head yellowish white tinged with red; thorax olive-brown
mixed with black-brown Sie some whiti ish abdomen yellowish
white tinged with red, irrorated with black and dorsally banded
with black; antenne ringed with black; palpi red-brown, ochreous
in front; pectus ochreous i in front ; legs ochreous tinged with red
and irrorated with blackish. Fore w ing with the costal and
terminal areas olive-green slightly irrorated with blackish ; some
dark reddish-brown suffusion at base in and below the cell; a.broad,
obliquely curved, diffused dark reddish-brown fascia from near
termen below apex to inner margin near base, a dark brown dis-
coidal spot ; a rather lunulate white mark from costa before apex,
and broad oblique white postmedial band from vein 5 to inner
margin with a dentate brown subterminal line slightly defined on
outer side by white on its outer edge with a reddish shade beyond
it; a terminal series of small blackish spots; a fine whitish line at
base of cilia followed by small dark spots. Hind wing whitish
suffused with purplish red and slightly irrorated with brown; some
brown suffusion at base; a curved slightly sinuous brown post-
medial line defined on outer side by white except towards costa; a
terminal series of dark bars separated by white points from apex to
submedian fold ; cilia with a white line at base followed by a dark
line. Under side whitish suffused with red and irrorated with
brown ; fore wing with broad dark reddish-brown shade on costal
area extending to inner margin before middle, a lunulate whitish
= We
Pyralide of the Subfami’y Pyraline. 89
patch from costa towards apex and oblique waved brown sub-
terminal line defined on outer side by whitish; hind wing with
rather diffused curved slightly v waved dark eden postmedial line
defined on outer side by whitish.
Hab. Durcu N. Gurve EA, Snow Mts., Setakwa R. (Meek),
3d type. Exp. 26-28 mm.
(4a) Paractenia sanguitincta, sp. n.
@. Head, thorax, and abdomen ochreous mixed with some
purplish red ; pectus, legs, and ventral surface of abdomen tinged
with red and irrorated with black. Fore wing yellowish irrorated
with purplish red, more thickly on basal costal area; an oblique
diffused purplish-red postmedial line, somewhat angled inwards at
submedian fold; a maculate deep red terminal line with some
blackish scales on it; cilia white at base, blackish mixed with
whitish at tips. Hind wing yellowish irrorated with purplish red
and blackish; a faint diffused curved reddish postmedial line; a
fine blackish terminal line; cilia white at base, blackish mixed
with whitish at tips. Underside yellow tinged with purplish red
and irrorated with black; both wings with indistinct diffused dark
postmedial line.
Hab. Gotp Coast, Kumasi (Sanders), 1 2 type. Exp.
22 mm.
(5a) Paractenia phanerostola, sp. n.
@. Head, thorax, and abdomen pale glossy red-brown, the palpi
and fore legs rather deeper red-brown. raore wing pale glossy red-
brown, the costa rather deeper red-brown; a faint rather diffused
curved brown antemedial line; a faint bis postmedial line, ex-
curved to vein 4, then oblique. Hind wing pale glossy red-brown ;
a faint diffused oblique brown antemedial line and rather more
distinct curved postmedial line. Underside of fore wing whitish
suffused with red-brown; hind wing whitish tinged with red-
brown ; both wings with curved brown postmedial line.
Hab. Ecvapvor, R. Bobonaza, Canelos (Palmer), 1 Q type.
Exp. 34 mm.
(56) Paractenia castaneonigra, sp. n.
Head, thorax, and abdomen chestnut-red, the last dorsally
suffused with black; legs suffused with black, the tarsi black
ringed with chestnut. Fore wing chestnut-red, the base and the
whole wing beyond the antemedial line suffused with black-brown ;
antemedial line blackish, oblique to submedian fold, then angled
inwards at vein 1; the medial part of costa with reddish-ochreous
points ; postmedial line indistinct, ochreous, very slightly waved,
excurved at middle and angled inwards at submedian fold; cilia
blackish at base and with some ochreous scales at middle. Hind
wing glossy black-brown with a chestnut-red tinge; cilia black
at base, bright yellow at tips. Underside black-brown ; fore wing
90 Sir G. F. Hampson on new
with the costal area chestnut-red to the postmedial line, the costal
edge black-brown with ochreous points on it, the postmedial line
yellow on costal area, then indistinct; hind wing with the costal
area and cell chestnut-red to the postmedial line, the inner area
whitish irrorated with black-brown, a yellowish postmedial line
slightly incurved in submedian interspace.
Hab, Cameroons, Ja R., Bitje (Bates), 13,29 type. Euxp.,
3 28, 2 36 mm.
(6a) Paractenia sichimensis, sp. n.
Head and thorax yellow mixed with brick-red ; abdomen yellow
tinged with red and irrorated with brown; legs yellow irrorated
with dark brown. Fore wing yellow suffused with red and irro-
rated with brown, the medial area yellower with a nearly clear
yellow conical patch from costa before the postmedial line and
another patch below end of cell; antemedial line brown defined on
outer side by yellow, excurved to median nervure, then oblique ;
the medial part of costa dark brown with yellow points on it;
a small brown discoidal spot; postmedial line brown, diffused on
outer side, inwardly oblique and somewhat incurved below vein 5;
a terminal series of blackish points; a fine pale line at base of cilia
followed by a dark line. Hind wing yellowish white, the termen
slightly tinged with red and irrorated with brown; a terminal
series of blackish points and fine pale line at base of cilia followed
by a dark line. Underside yellow tinged with red and irrorated
with dark brown; fore wing with the costal area suffused with
brown, a small dark discoidal spot, yellower patch from costa .
beyond the cell, and diffused oblique dark postmedial line; hind
wing with slight brown discoidal spot and curved postmedial line.
Hab. Sixurm (Pilcher, Moller), 8 3,5 Q type. xp. 20-
24 mm. ;
(7 a) Bostra purpurealis, sp. n.
Head and thorax bright purple-red; abdomen pale purple-red
irrorated with brown; antenne white above; fore tibize white on
outer side, the tarsi white. Fore wing bright purple-red irrorated
with brown, the medial area tinged with fuscous, the costal edge
white except towards base; antemedial line fuscous, oblique to
submedian fold, then slightly incurved; pustmedial line fuscous
slightly defined on outer side by white and slightly meurved in
submedian interspace. Hind wing whitish suffused with pale
purple-red and fuscous brown ; a curved fuscous postmedial line
defined on outer side by whitish from costa to vein 1; cilia bright
‘purple-red. Underside grey with pale purplish-red streaks along
the veins; both wings with the costal area purple-red to beyond
middle and with fuscous postmedial line; fore wing with white
scales on the costa.
Hab. Br. C. Arrica, Mt. Mlanje (Neave), 3 5, 1 @ type;
PortucuEsE E. Arrica; Mt. Chiperone (Neave), 1 3. ap.
30-32 mm.
Pyralide of the Subfamily Pyraline. 91
(7 b) Bostra cenochroa, sp. n.
Head, thorax, and abdomen pale purplish red thickly irrorated
with fuscous ; antennz of male pale red. Fore wing pale purplish
red thickly irrorated with fuscous, the costal edge redder; ante-
medial line rather diffused, black, excurved below costa then
slightly incurved ; a small black discoidal spot; postmedial line
rather diffused, black faintly defined on outer side by whitish in
male, and slightly excurved. Hind wing uniform glossy grey-
brown with a purplish-red tinge. Underside greyish brown, the
costa of fore wing purplish red irrorated with dark brown; hind
wing with faint curved dark postmedial line.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 1 d, 2 G type;
TransvaaL, White R. (Cooke), 2 3. Exp. 26-80 mm.
(7h) Bostra pallidifrons, sp. n.
3S. Head white tinged with pale rufous; thorax and abdomen
grey-brown mixed with whitish; tibiz and tarsi whitish tinged
with brown. Fore wing dark brown mixed with grey-white ;
antemedial line diffused, whitish, oblique to median nervure and
slightly incurved in submedian interspace; the medial part of
costa with white points with black-brown between them; a small
blackish discoidal spot ; postmedial line diffused, whitish, incurved
below vein 4. Hind wing uniform grey-brown. Underside grey
suffused and irrorated with brown; fore wing with whitish points
on costa with black-brown between them except at base and apex.
Hab. Masnonatand, Enkeldoorn Distr. (Jliss LE. 8. Younge),
1 ¢ type. Kup. 14 mm.
(77) Bostra lignealis, sp. n.
Head and thorax pale brownish red; abdomen whitish tinged
with red and irrorated with blackish; legs whitish irrorated with
dark brown. Fore wing pale red-brown irrorated with blackish ;
antemedial line whitish defined on outer side by blackish, almost
straight; the medial part of costa with white points with some
black between them; a small black discoidal spot ; postmedial line
whitish defined on inner side by blackish, slightly excurved at
discal fold and below submedian fold; cilia with a white line
at base. Hind wing pale red-brown irrorated with blackish, an
oblique whitish antemedial line defined on outer side by blackish
and joined at inner margin by a slightly sinuous whitish post-
medial line defined on inner side by blackish; cilia with a white
line at base. Underside whitish suffused with red-brown and
irrorated with blackish; both wings with sinuous white postmedial
line; fore wing with blackish discoidal spot.
Hab. Br. KE. Arrica, Nairobi (Anderson), 1 56, 1 Q type.
Eep. 20 mm.
92 , Sir G. F. Hampson on new
(9a) Bostra rusinalis, sp. n.
3. Head and thorax rufous with a few dark brown scales;
abdomen whitish tinged with rufous. Fore wing rufous sparsely
irrorated with dark brown, the basal costal area suffused with
blackish; traces of a waved dark antemedial line; the medial part
of costa with whitish points with dark brown between them; a
small dark discoidal spot ; postmedial line indistinct, dark, slightly
excurved to vein 4 then slightly incurved. Hind wing whitish
tinged with rufous, the termen and bases of cilia rather deeper
rufous; the underside with the costal area suffused with rufous
and irrorated with dark brown.
Hab. Gotp Coast, Appan, 2 ¢ type. Hap. 22 mm,
(10a) Bostra pallidicolor, sp. n.
$. Head and thorax whitish tinged with pale red and irrorated
with a few dark brown scales; abdomen white faintly tinged with
red and slightly irrorated with brown; palpi and pectus in front
redder. Fore wing whitish tinged with purplish red and irrorated
with brown, the red rather deeper on basal costal area and forming
diffused streaks in discal and submedian folds; a very faint diffused
dark antemedial line angled outwards at suhmedian fold; a black
discoidal spot; postmedial line indistinct, dark, diffused, oblique
below vein 4; a terminal series of minute blackish spots. Hind
wing ochreous white ; a terminal series of slight brown points trom
apex to vein 2. Underside white; fore wing with the costal area
bright rufous with the costal edge white, then tinged with ochreous
and irrorated with brown except on inner area ; hind wing with the
costal area tinged with rufous and irrorated with brown, a blackish
spot at upper angle of cell.
Q@. Head and thorax greyish tinged with purplish red ; abdomen
whitish thickly irrorated with brown ; fore wing greyish uniformly
tinged with purplish red; hind wing suffused with pale reddish
brown; underside of fore wing suffused with brown except on
costal area, the hind wing tinged with brown and with curved
brown postmedial line from costa to submedian fold.
Hab. Transvaat, Pretoria (Distant, Janse), 1 3,1 Q type.
Exp., 5 30, 9 26 mm.
(10 c) Bostra dentilinealis, sp. n.
Head and thorax whitish tinged with red-brown and irrorated
with a few black-brown scales; abdomen suffused with fiery red
and irrorated with black. Fore wing whitish suffused with pale
red-brown, sometimes tinged with olive-green and irrorated with
black-brown ; an indistinct dentate blackish postmedial line some-
what ineurved at submedian fold; a terminal series of black striz ;
cilia with reddish-brown lines near base and at tips. Hind wing
whitish suffused with fiery red and slightly irrorated with brown ;
a curved dentate blackish postmedial line, rather diffused on inner
Pyralide of the Subfamily Pyraline. SP
side; a terminal series of black-brown bars; cilia with a red line
near base. Underside whitish suffused with red and irrorated with
dark brown; both wings with slight dark discoidal strie and
diffused curved dentate postmedial line.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 1 S, 3 Q type.
Exp. 24 mm.
(lla) Bostra pyrochroalis, sp. n.
6. Head, thorax, and abdomen bright fiery red; the antennz
blackish except at base; the frons and palpi whitish, the latter
black at tips. Fore wing bright fiery red slightly irrorated with
blackish, especially before the antemedial line and on terminal area;
antemedial line whitish, excurved below costa, then slightly
incurved ; postmedial line whitish, slightly excurved beyond the
cell, then slightly incurved, both the lines faintly defined by
fuscous. Hind wing bright fiery red slightly irrorated with
blackish, especially on apical area; faint oblique whitish ante- and
postmedial lines slightly defined by fuscous and somewhat approxi-
mated towards inner margin. Underside of both wings with
blackish discoidal point and curved white postmedial line; fore
wing with series of white points on costa to the postmedial line.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 1 ¢ type. Hap.
14 mm.
(10 b) Bostra flavalis, sp. n.
3. Head, thorax, and abdomen yellow with a faint olive tinge ;
palpi fuscous below; tibize with the spurs fuscous at base. Fore ~
wing yellow with a faint olive tinge ; antemedial hne formed by
slight black striz defined on inner side by white, slightly sinuous ;
the medial part of costa with black and white points; a slight
blackish discoidal striga ; postmedial line formed by black scales
defined on outer side by white, slightly excurved below costa and
at middle and incurved at discal fold and below vein 3; a terminal
series of black striz; cilia with some blackish at tips. Hind wing
yellowish white tinged with fuscous brown ; a curved fuscous post-
medial line slightly defined on outer side by white; a terminal
series of blackish strize ; cilia with a slight brownish line near base.
Underside of fore wing suffused with brown except on inner area
- to the postmedial line ; hind wing irrorated with brown; a curved
dark postmedial line.
Hab. Formosa, Kanshirei (Wileman),2 $ type. Hxp.16 mm.
(19 ¢) Bostra phenicocraspis, sp. n.
Head, thorax, and abdomen olive-yellow, the last tinged with
crimson at extremity ; antenne whitish tinged with brown; fore
and mid tibiz and the hind tibie at extremity crimson, the fore
legs and ventral surface of abdomen except towards base irrorated
with brown. Fore wing olive-yellow irrorated with a few red-
brown scales ; a faint red-brown antemedial line, oblique to: sub-
94 Sir G. F. Hampson on Gan
median fold; the medial part of costa with whitish points with
blackish between them ; a slight brown discoidal striga ; postmedial
line slight, brown, somewhat oblique to vein 4 and slightly ex-
curved above inner margin; a terminal series of black bars; cilia
deep crimson at base, then paler crimson with the tips blackish to
vein 8. Hind wing olive-yellow; an obscure line formed by brown
- scales from lower angle of cell] to inner margin and a similar
slightly curved postmedial line; the terminal area irrorated with
a few brown scales; a terminal series of black strie; cilia deep
crimson at base, paler crimson at tips. Underside of fore wing
crimson irrorated with black except on costal area to the post-
medial line and below vein 3 to termen, the costa with series of
quadrate black spots to the postmedial line which is black, ex-
curved from below costa to vein 3, then erect, a black discoidal
point; hind wing suffused with crimson and irrorated with black
to the postmedial line and on apical area, the termen then narrowly
crimson, the postmedial line blackish, obliquely curved to vein 2,
then sinuous.
Hab, CamEroons, Ja R., Bitje (Bates), 1 3,2 2 type. Exp.
22 mm.
(19e) Bostra phenicoxantha, sp. n.
6. Head, thorax, and abdomen whitish suffused with dull
purplish red; antenne ringed with brown; fore legs crimson;
pectus, mid and hind legs, and ventral surface of abdomen. ochreous.
Fore wing brilliant crimson with a yellow medial band exeept on
costal area, defined on inner side by the faint curved crimson ante-
medial line and on outer by the similar postmedial line excurved at
middle; a blackish terminal line except towards tornus. Hind
wing brilliant crimson, with a broad yellow medial band defined
on inner side by the slight curved crimson antemedial line and on
outer by the similar postmedial line excurved at middle; a black
terminal line except towards tornus; cilia whitish at tips. Under-
side with the crimson paler and duller.
Hab. Br. C. Arnica, Mt. Mlanje (Neave), 2d type. Bap.
20 mm.
(21a) Bostra maculilinea, sp. n.
Head, thorax, and abdomen white mixed with cupreous red and
irrorated with black, the abdomen suffused with black towards
extremity; antenne white ringed with black; fore tibie and the
tarsi black ringed with white. Fore wing cupreous red thickly
irrorated with black ; some white at base of inner margin; ante-
medial line white defined on outer side by rather diffused black,
slightly waved; the medial part of costa with white points with
blackish between them; postmedial line white defined on inner
side by diffused black and forming a small white spot at discal
fold and larger spot in submedian interspace, slightly waved, ex-
curved at middle and incurved in submedian interspace; a black
terminal line; cilia white with blackish lines at base and middle
Pyralidee of the Subfamily Pyraline. 95
and at tips. Hind wing white tinged with red-brown and irro-
rated with fuscous; a curved slightly sinuous white postmedial
line; a black terminal line; cilia white mixed with fuscous and
with black line near base. Underside of fore wing whitish suffused
with fuscous, the costal area chestnut-red to the white postmedial
line, the costal edge fuscous with white points ; hind wing white
tinged with red and irrorated with fuscous, a white postmedial line
defined on inner side by black, slightly incurved at discal fold, then
excurved.
Ab. 1. Fore wing with the antemedial line obsolescent towards
costa, the postmedial line obsolescent at middle.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 8 3, 4 Q type.
Exp. 14-18 mm.
(216) Bostra ignirubralis, sp. n.
3. Head, thorax, and abdomen fiery red, the last slightly
nrorated with blackish; antenne whitish tinged with red; palpi
and legs deeper red with some blackish mixed, the mid and hind
tarsi whitish tinged with red. Fore wing fiery red; antemedial
line brown defined on inner side by whitish, very slightly curved ;
a slight blackish discoidal spot; the medial part of costa with
slight white points with black between them; postmedial line
fuscous brown defined on outer side ‘by whftish, very slightly
curved and slightly incurved at submedian fold; cilia with some
fuscous mixed. Hind wing fiery red; an oblique dark antemedial
line defined on inner side by whitish, met above inner margin by a
curved dark postmedial line defined on outer side by whitish ; cilia
with some fuscous mixed. Underside paler red irrorated with
fuscous brown; fore wing with the inner area whitish, the costal
edge dark brown with white points to the indistinct dark post-
medial line, a slight dark discoidal spot; hind wing with faint
eurved dark postmedial line. ss
Hab. Cayton (Mackwood), 1 ¢ type. Exp. 32 mm.
(5) Zitha fulviceps, sp. n.
Head fulvous yellow, the antenne dark brown ringed with
whitish, the palpi suffused with brown except above; thorax dark
brown ; abdomen, pectus, and legs yellowish suffused with brown.
Fore wing glossy fuscous brown; an indistinct blackish discoidal
spot; postmedial line indistinct, blackish, exeurved below costa
and slightly incurved below vein 4. Hind wing glossy fuscous.
brown. Underside fuscous brown; both wings with indistinct
curved dark postmedial line.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 2 3,1 9 type ;
Portuguese E. Arrica, Mt. Chiperone (Neave), 1 3. Exp.
16 mm.
Genus PHRYGANOMIMA, nov.
Type, Phryganodes noctifer, Dogn.
Proboscis fully developed ; palpi upturned, the 2nd joint reaching
96 Sir G. F. Hampson on new
to middle of frons, moderately scaled, the 38rd moderate; maxillary
palpi triangularly dilated with scales; antenne of male laminate.
Fore wing with the median nervure strongly downcurved towards
end of cell, the discocellulars strongly excurved; vein 3 from
before angle of cell; 5 from above angle; 6 from upper angle;
7, 8, 9 stalked from before angle; 10,11 from cell. Hind wing
with the cell short; veins 3 and 5 from near angle; 6, 7 from
upper angle, 7 not anastomosing with 8.
Genus LoryMODES, nov.
Type, Pyralis diagonalis, Hmpsn.
Proboscis fully developed ; palpi upturned, the 2nd joint tufted
with scales produced to a point in front at extremity, the 3rd long
and acuminate; maxillary palpi slightly dilated with scales, antenne
of male with fasciculate cilia, the basal joint long. Fore wing
with vein 3 from close to angle of cell; 4, 5 stalked; 6 from upper
angle; 7, 8, 9 stalked; 10,11 from cell. Hind wing with vein 3
from close to angle of cell; 4, 5 stalked; 6, 7 from upper angle,
7 not anastomosing with 8.
(2) Lorymodes stenopteralis, sp. n.
Head, thorax, and abdomen white tinged with red-brown ;
antennz fuscous except towards base. Fore wing very narrow;
whitish suffused with red-brown; a slight brown antemedial mark
in submedian fold; the medial part of costa with slight brown
points; antemedial line slight, black defined on inner side by
whitish, very oblique from costa to the postmedial line at vein 2,
then inwardly oblique to inner margin, defining on inner side the
postmedial line, which is rather diffused black-brown defined on
outer side by whitish, very oblique and slightly curved from costa
near apex to gabmedian fold, then still more oblique to middle of
inner margin; a fine brown terminal line; cilia whitish. Hind
wing silvery white. Underside of fore wing white, tinged with
brown on costal half.
Hab. Gambia, 1 do; N. Nigeria, Zungeru (Macfie), 1 2 type.
Hep. 16 mm.
Genus DATTINIA.
Type.
Dattinia, Rag. Bull. Soc. Ent. Fr. 1887, p. cxxxvil ................46 syrtalis.
Constantia, Rag. Bull. Soc. Ent. Fr. 1887, p. exxxvii (nec Adams,
IMFO IIE BIB GO) marie etree ean aerate nica? he co aint nee aene leonalis.
Buliana, Navas, Bol. Soc. Aragon, x. p. 64 (1913) .................. leonalis,
(4e) Dattinia eumictalis, sp. n.
@. Head and thorax yellowish white mixed with rufous, red-
brown, and dark brown; abdomen white suffused with red-brown ;
antennze white ringed with dark brown; legs white irrorated with
brown. Fore wing yellowish white very thickly irrorated with
rufous and red-brown and with a few dark brown scales; a whitish
Pyralide of the Subfamily Pyraline. 97
subbasal spot below the cell; antemedial line strong, white, rather
oblique to submedian fold and incurved at vein 1; a white spot in
end of cell before the blackish-brown discoidal bar; postmedial
line black-brown defined on outer side by whitish, incurved below
costa, angled outwards at veins 6, 5, 4, then retracted and almost
obsolete to below end of cell, then erect, sinuous and more dis-
tinctly detined on outer side by white; a terminal series of faint
brown spots; cilia chequered red-brown and white. Hind wing
white tinged and irrorated with red-brown; a terminal series of
slight brown strie; cilia whiter. Underside silvery white, the
fore wing except on inner area and the costal area of hind wing
urorated with red-brown.
Hab. Supan, Port Sudan (Waterfield), 1 Q type. Exp.
32 mm. ; 5
Genus ANOBOSTRA, nov.
Type, A. discimacula.
Proboscis present; palpi with the 2nd joint porrect, typically
projecting about the length of head, the scaling at extremity below
produced to a point, the 8rd obliquely upturned, moderate ;
maxillary palpi dilated with scales; antenne of male thickened
and with long fasciculate cilia, the basal joint with tuft of scales.
Fore wing with vein 3 from angle of cell; 4, 5 typically stalked ;
6 from upper angle; 7, 8, 9 stalked ; 10, 11 from cell. Hind wing
with vein 3 from angle of cell; 4, 5 typically stalked; 6,7 from
upper angle, 7 not anastomosing with 8.
(1) Anobostra discimacula, sp. n.
Palpi with the 2nd joint projecting about the length of head ;
both wings with veins 4, 5 stalked.
Head and thorax purplish red; abdomen grey suffused with
brown ; antennze dark brown; pectus, legs, and ventral surface of
abdomen pale red irrorated with brown. Fore wing purplish red,
irrorated with blackish, the medial area slightly irrorated ; ante-
medial line blackish defined on inner side by pale red, oblique to
submedian fold, where it is angled outwards; a series of small
blackish spots on medial part of costa ; a large black discoidal spot ;
postmedial line rather diffused, black defined on outer side by pale
red, excurved below vein 7 and at middle and incurved at discal
fold and below vein 3, a series of slight blackish streaks beyond it
on the veins; a terminal series of minute black spots. Hind wing
erey suffused with brown; a whitish line at middle of cilia.
Underside grey suffused with brown; fore wing with the costa
pale reddish with numerous brown striz.
Hab. Br. C. Arrica, Mt. Mlanje (Neave), 2 5 type; Porrv-
GuESE EK. Arrica, Mt. Chiperone (Weave), 1 9; Masnonananp,
Umtali (Marshall), 1 9. Kzxp., 3 18-20, 2 22-26 mm.
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. Z
98 Sir G. Ff. Uampson on new
(3) Anobostra albilinealis, sp. n.
Palpi with the 2nd joint projecting about twice the length of
head; both wings with veins 4, 5 from the cell.
2. Head, thorax, and abdomen pale red-brown. Fore wing
chocolate red-brown with the basal area pale red-brown except
towards costa; antemedial line white, rather oblique to median
nervure ; a series of minute white spots on medial part of costa ;
postmedial line white, excurved to vein 3, then incurved; cilia
greyish brown with a fine whitish line at base. Hind wing pale
red-brown mixed with some whitish, the base whiter; a curved
whitish postmedial line; cilia greyer brown with a fine pale line at
base. Underside red-brown mixed with whitish; fore wing with
series of small white spots on costa to the postmedial line, the
postmedial line of both wings slightly waved.
Hab. Asysstnta, Dirre Dawa (Drake-Brockman), 1 2 type.
Exp. 30 mm.
(4) Anobostra punctilinealis, sp. n.
@. Head and thorax deep purple-red mixed with black ;
abdomen greyish tinged with purple-red and thickly irrorated with
black; pectus, legs, and ventral surface of abdomen purplish red
mixed with black, the fore tibiz and the tarsi brownish white.
Fore wing deep purple-red irrorated with black ; antemedial line
indistinct, blackish, oblique to just below the cell, where it is
angled outwards, then inwardly oblique and defined on inner side
by white; the medial part of costa with series of white points ;
postmedial line indistinct, blackish defined on outer side by white
scales and by small spots at discal and submedian folds, minutely
dentate, incurved below vein 4; cilia with a fine whitish line at
base. Hind wing grey tinged with purplish red and suffused and
irrorated with brown, the termen and cilia redder; a faint curved
dark postmedial line. Underside dull purplish red irrorated with
fuscous brown; fore wing suffused with fuscous brown to the post-
medial line except on costal area, a series of white points on basal
half of costa; the postmedial line blackish with a slight white mark
at costa; hind wing with rather diffused minutely dentate blackish
postmedial line.
Hab. Br. C. Arrica, Mt. Mlanje (Weave), 3 2 type. LHuxp.
28 mm.
(5) Anobostra radialis, sp. n.
Q. Head and thorax red-brown mixed with ochreous. Fore
wing greyish suffused with red-brown, especially on basal half and
beyond lower angle of cell ; antemedial line whitish defined by dark
brown on outer side, angled outwards below costa, then obliquely
curved and with some black scales before and beyond it at inner
margin; some dark rufous suffusion in and below middle of cell ; °
veins 2 to 6 streaked with whitish and defined on each side by
Pyralide of the Subfamily Pyralinz. 99
slight dark brown streaks to the postmedial line which is very near
termen, white defined by dark brown on inner side, excurved between
veins 6 and 3, then oblique and slightly sinuous; a fine brown
terminal line; cilia whitish with grey medial band and line near
tips. Hind wing yellowish white slightly tinged with rufous
especially towards termen; an indistinct curved whitish sub-
terminal line incurved at vein 2; cilia whitish with a slight brown
line near base; the underside with the subterminal line more
distinct and defined by brown on inner side.
Hab. Br. E. Arrica, Taveta (K. St. A. Rogers), 2 2 type;
Uaanpa, Gondokoro (Reynes-Coles), 19. Lup. 30-34 mm.
(1b) Tyndis medio-pallens, sp. n.
Head and tegulw pale reddish, the thorax pale reddish mixed
with dark brown; abdomen grey suffused with black-brown, the
base whitish, the anal tuft rufous; antennz and palpi irrorated
with dark brown; pectus, legs, and ventral surface of abdomen
whitish tinged with red-brown and irrorated with dark brown. Fore
wing whitish tinged with red-brown and irrorated with dark brown,
the medial area much paler except at costa, the basal costal area
and a broad shade beyond the postmedial line darker; antemedial
line rather diffused “dark brown, slightly curved; the medial part
of costa with minute greyish spots; a black striga on upper disco-
cellular ; postmedial line dark brown slightly defined on outer side
by whitish towards costa, very oblique and straight; a terminal
series of blackish points except towards costa and fine whitish line
at base of cilia. Hind wing creamy white slightly irrorated with
brown, the terminal area more thickly irrorated; traces of a brown
antemedial line from cell to inner margin; a curved brown posts
medial line; a brown terminal line and white line at base of cilia.
Underside whitish tinged with red-brown and thickly irrorated
with dark brown; both wings with small dark discoidal spot and
oblique postmedial line.
Hab. Br. E. Arrica, Nairobi (Betton), 1 g, Kikayu, Fort
Smith (Crawshay), 1 3 type, Eb Urru (Betton), 1 3, Nakuru
(Bodeker), 1 3,1 9; Germ. EH. Arrica, Kilimandjaro (Sjéstedt), -
1d. xp. 26 mm,
(le) Tyndis pallidirufa, sp. n,
Head and thorax whitish suffused with rufous; abdomen
whitish tinged with rufous and irrorated with black, a blackish
band on third segment; pectus, legs, and ventral surface of
abdomen rufous irrorated with brown. Fore wing whitish suffused
with rufous and irrorated with dark brown except on medial area ;
a rather diffused erect black almost medial line from subcostal
nervure to inner margin and a similar oblique postmedial line from
below costa to inner margin; a terminal series of minute black
spots; cilia with blackish lines near base and tips. Hind wing
100 Mr. W. L. Distant on the
whitish suffused with reddish brown; an oblique dark postmedial
shade; a dark terminal line; cilia whitish with dark lines near
base and tips. Underside creamy white tinged with rufous and
irrorated with brown; fore wing with dark discoidal point and
suffused oblique postmedial line from below costa to inner margin ;
hind wing with diffused oblique black postmedial line; both wings
with terminal series of minute black ue
Hab. Sierra Leone (Clements), 13,1 9 type. Exp., gd 20,
Q 22 mm.
(6) Tyndis pyrrhoxantha, sp. n.
2. Head and thorax yellow suffused with brilliant fiery red ;
abdomen yellowish tinged with fiery red and irrorated with fuscous;
pectus, legs, and ventral surface of abdomen yellow tinged aie
red. Hons wing orange-yellow suffused with fiery red a 1rro-
rated with black ; traces of a yellow antemedial line, oblique to
submedian fold and incurved at vein 1; postmedial line rather
diffused yellow, excurved at middle and hpennved at submedian
fold. Hind wing yellow, the inner and terminal areas faintly
tinged with red and the latter irrorated with blackish. Underside
orange-yellow ; fore wing with minute dark discoidal point, the
anal area tinged aia red and irrorated with black; hind wing
with the apex tinged with red and irrorated with black.
Hab. Gown Coast, Bibianaha (Spwrrell), 1 2 type. Hap.
22 mm.
V.—The Homoptera of Indo-China.
By W. L. Distant.
Fam. Cicadide.
For some time Mon. R. Vitalis de Salvaza has sent me
collections of Homoptera from this very interesting region,
and I believe he intends at some future date to publish an
illustrated work on the insect fauna of Indo-China. The
Homoptera already received are from the frontier of Laos,
East Annam, and from Lao Kay and Chapa in Tovkin. In
this contribution I give a rough list of the species already
received, which number fifty-five, belonging to the family
Cicadidz alone. Examples of all these, including types and
uniques, are placed in the collection of the British Museum,
which, as regards this family, is now by far the largest and
most complete i in the world.
T also add the descriptions of six new species.
Homoptera of Indo-China.
101
List of Species already received.
nigrosignata, Dist.
Tosena melanoptera, White.
Rihana bimaculata, Oliv.
Cryptotympana mandarina, Dist.
holsti, Dist.
Inthaxvara rex, Dist.
Salvazana mirabilis, Dist.
Leptopsaltria phra, Vist.
Dundubia mannifera, Linn,
var. terpsichore, Walk.
Cosmopsaltria fratercula, Dist.
vopaga, Dist-
andersont, Dist.
—— tonkiniana, Jac.
Haphsa nana, Dist.
Platylomia nagarasingna, Dist.
operculata, Dist.
radha, Dist.
Meimiuna nicrodon, Walk.
subviridissima, Dist.
—— rava, Dist.
Pomponia intermedia, Dist.
Fusca, Oliv.
lactea, Dist.
Aola scitula, Dist.
bindusara, Dist.
Terpnosia crowfooti, Dist.
Platypleura badia, Dist. |
Terpnosia posidonia, Jac.
madhava, Dist.
chapana, Dist.
ransonneti, Dist.
rustica, Dist.
mesonotalis, Dist,
clio, Walk.
maw, Dist.
Caleagninus sulvazanus, Dist.
Geana vitalist, Dist.
maculata, Dru.
annamensis, Dist.
sultana, Dist.
pavict, Noualh,
Balinta pulchella, Dist.
delinenda, Vist.
Talainga binghami, Dist.
distantt, Jac.
Mogannia cyanea, Walk.
hebes, Walk.
cesar, Jac,
conica, Germ.
indigotea, Dist.
Huechys sanguinea, De Geer.
tonkinensis, list.
Scieroptera splendidula, Fabr.
Lemuriana apicalis, Germ,
HT |
Terpnosia rustica, sp. n.
Head, pro- and mesonotum pale olivaceous green ;
head
with two spots at apex of front and a lateral spot near
base of antenne, two curved central lines on vertex, and a
spot above margins of eyes aud two small spots between
central fasciz and eyes black ; pronotum with two central
longitudinal fascia which are angulated anteriorly and
posteriorly, the furrow behind eyes, and the lateral margins
black ; mesonotum with central obconical lines which are
centrally, posteriorly prolonged, a sublateral line on each
side, and a spot near each anterior angle of the basal
Peleg elevation black; abdomen above and beneath
ochraceous, with the basal margin and apical area black,
the basal segments above are also centrally spotted with
black ; legs and opercula pale olivaceous-green, the latter
with black lateral margins, the tarsi ochraceous and apically
black ; tegmina and wings subhyaline, the first with the
veins ‘blackish, the transverse veins at the bases of first,
second, and third apical areas with pale brown suffusions
aud some obscure spots of the same colour on the longi-
102 Mr. W. L. Distant on the
tudinal veins to apical areas, costal and post-costal mem-
branes ochraceous ; wing-venation blackish; opercula in
female short and. transverse, subconically oblique, not
extending beyond base of abdomen ; face conically produced
and somewhat strongly laterally striate; base of head at
region of ocelli suleate ; pronotum centrally longitudinally
sulcate ; tympanal coverings both narrower and shorter than
tympanal cavities.
Long., excl. tegm., @, 15 mm.; exp. tegm. 49 mm.
Hab. Tonkin; Chapa (R. Vitalis de Salvaza).
This species may be placed near 7. ransonneti, Dist.
Terpnosia chapana, sp. n.
3d. Head, pronotum, mesonotum, face, sternum, legs,
and opercula olivaceous green ; abdomen above and beneath
pale testaceous, the abdominal margins a little darker, and
the apical abdominal area black; lineate markings to
anterior margin of front, a transverse spot near insertion
of autenne, and a suffusion at the region of the ocelli black ;
pronotum with two central longitudinal linear fascie, nar-
rowed: and united at base, the furrows, two spots on each
lateral area, and the extreme basal margin black ; meso-
notum with the margins of two anterior obconical spots,
followed on each side of anterior margin by a small angulate
spot and again by a submarginal longitudinal fascia, a
central longitudinal spot reaching middle of cruciform
elevation and a spot before each anterior angle of same,
black ; tegmina and wings subhyaline, the venation black
or blackish, tegmina with the costal membrane and post-
costal area ochraceous, the transverse veins at the bases of
the second, third, and fifth apical areas, and the apices of the
longitudinal veins to the apical areas spotted with fuscous ;
tympanal coverings well developed, but shorter and narrower
than tympanal cavities ; face centrally sulcate and strongly
transversely striate, vertex between the ocelli sulcate ;
opereula subtruncately oblique, scarcely passing the base of
abdomen ; greatest width of tegmina about one-third
of length.
Long., excl. tegm., ¢, 18 mm.; exp. tegm. 55 mm.
Hab. Tonkin, Chapa (R. Vitalis de. Salwaza).
The nearest allied species is 7. postdonia, Jac.
Terpnosia mesonotalis, sp. 0.
3. Head, pronotum, abdomen above and_ beneath,
sternum and legs ochraceous, the upper surface of the
Homoptera of Indo-China. 103
abdomen moderately rufescent ; mesonotum uniformly pale
ochraceous; pronotum with the sublateral furrow marked
with black ; the mesonotal cruciform elevation dark tes-
taceous ; opereula pale ochraceous ; tegmina and wings
subhyaline, the veins fuscous; tegmina with the costal
membrane and post-costal area dull ochraceous with dark
linear markings, extreme basal angle of upper ulnar area
dark fuscous; vertex of head sulcate between the ocelli;
sublateral furrows to pronotum very profound; face with
very strong transverse ridges; tympanal coverings small,
very much shorter and a little narrower than tympanal
cavities ; opercula short and broad, not quite reaching base
of abdomen. .
Long., excl. tegm., as 17 mm.; exp. tegm. 45 mm.
Hab. Toukin; Chapa (R. Vitalis de Salvaza).
To be placed near T. madhava, Dist.
Calcagninus salvazanus, sp. v.
3. Body and legs ochraceous, mesonotum a little paler,
sometimes blackish markings at base of abdomen beneath ;
tegmina and wings subhyaline, the venation fuscous brown,
tegmina with the whole of the costal and subcostal areas
ochraceous ; tympanal coverings imperfect ; abdomen tuber-
culate beneath on second and third abdominal segments ;
head about as Jong as half the width between eyes ; wings
with six apical areas; mesonotum sometimes with indi-
cations of two dark lateral longitudinal fasciz ; opercula
wide apart, transverse, not passing basal abdominal segment,
apical margins roundly truncate; face with the lateral
striations distinct, but not profound; vertex of head
narrowly longitudinally suleate between the ocelli.
Long., excl. tegm., ¢, 15 mm. ; exp. tegm. 45 mm.
Hab. Tonkin ; “Chapa (Rk. Vitalis de Salvaz 2a).
Mogannia indigotea, sp. 0.
Body and legs very dark indigo-blue ; tegmina and wings
hyaline, the venation dark pcleacedusls ; tegmina with about
basal half flavescent, outwardly margined ‘with a transverse
fuscous fascia and an oblique macular fascia directed
inwardly and the basal cell of the same colour, costal
membrane dark ochraceous; base of wings narrowly dark
ochraceous. Front of head longly hirsute and longly
depressed, between the ocelli longitudinally sulcate ; pro-
notum with the furrows profound ; rostriim reaching the
intermediate coxe.
104 Mr. R. E. Turner on Fossorial Hymenoptera.
Long., excl. tegm., 14-17 mm. ; exp. tegm. 31-40 mm.
Hab. Tonkin; Cliapa (R. Vitalis de Salvaza). N. China
(Brit. Mus.). Philippine Is.; Malinao, Tayabas (C. T.
Baker).
A somewhat variable species, allied to M. effecta, Dist.
In some specimens the basal cell of tegmina is ochraceous,
in others the inner and outer dark fasciz of the basal area
are fused.
Huechys tonkinensis, sp. n.
Head, pronotum, and mesonotum black ; vertex of head
with almost anterior half, the ocelli and av angulated spot
behind them, pronotum with a central, broad, longitudinal
fascia which is strongly, medially, angularly compressed,
mesonotum with the lateral margins and a medial, longi-
tudinal, anteriorly strongly attenuated fascia sanguineous ;
face black, apically sanguineous; sternum, opercula in
male, body beneath and above sanguineous; legs black ;
tegmina dark brownish, the venation darker; wings sub-
hyaline, the venation dark brownish; head with the face
strongly, centrally, longitudinally sulcate for about two-thirds
its length, the transverse lateral striations very coarse ;
head (including eyes) about as wide as base of mesonotum ;
head about as long as pronotum; mesonotum (including
cruciform elevation) longer than pronotum; tegmina with
eight apical areas; opercula in male not passing base of
abdomen, well separated, but inwardly obliquely directed ;
their apices roundly truncate.
Long., excl. tegm., ¢,18 mm.; exp. tegm. 40 mm.
Hab, Tonkin; Chapa (R. Vitalis de Salvaza).
Allied to H. chryselectra, Dist., from Borneo.
VI.— Notes on Fossorial Hymenoptera.— XXV. On new
Sphecoidea in the British Museum. By Row uanp E,
*'TurNER, F,Z.S., F.E.S,
Subfamily Parcayrurs.
Cerceris armigera, 8p, 0.
©. Nigra; clypeo, mandibulis basi, scapo, fronte sub antennis,
segmento dorsali secundo macula basali utrinque, segmentisque
quarto quintoque fascia apicali emarginata flavis; vertice macula
utrinque pone oculos, pronoto macula utrinque, tegulis, femoribus
s
Mr. R. E. Turner on Fossorial Hymenoptera. 105
anticis, femoribus intermediis apice, tibiis tarsisque anticis inter-
mediisque brunneo-ferrugineis; segmentis dorsalibus secundo,
quarto, quintoque omnino, tertioque apice ferrugineis; alis
hyalinis, venis nigris clypeo brevissimo, subporrecto, apice
latissime emarginato, raat apicalibus dente valido armato;
mesopleuris haud dentatis ; segmento mediano area basali sub-
opaca, delicatissime punctata, segmento ventrali secundo area
basali elevata nulla.
Long. 8 mm.
¢. Coarsely and closely punctured ; head very broad, the
eyes distinctly divergent towards the clypeus; cheeks nearly
as broad as the eyes. Antennee inserted rather low down,
nearly three times as far from the anterior ocellus as from
the base of the clypeus, second joint of the flagellum a little
longer than the third. First abdominal segment distinctly
broader than long ; pygidial area coarsely but rather sparsely
punctured, elongate-ovate, rather broadly truncate at the apex.
Abdominal segments very strongly constricted, the ventral
surface almost smooth. First recurrent nervure received a
little before the middle of the second cubital cell, second
close to the base of the third cubital cell.
Hab. 5S. Queensland, Darra near Brisbane (Hacker) ; De-
cember.
The shape of the clypeus is remarkable and quite different
from any other Australian species.
Cerceris unispinosa, sp. 0.
@. Nigra; mandibulis basi, clypeo, fronte usque ad antennarum
basin, seapo, genis late, vertice macula obliqua utrinque, pronoto
fascia utrinque, tegulis, scutello macula transversa utrinque,
postscutello, segmento dorsali secundo macula transversa basali,
tertio fasaa apicali antice late emarginata, quarto fascia an-
eusta apicali, quinto fere omnino, segmentis ventralibus tertio
quintoque lateribus, femoribus anticis intermediisque subtus,
tibiis tarsisque anticis intermediisque flavis; segmento dorsali
secundo dimidio basali, segmentis ventralibus secundo fere toto,
tertio in medio, femoribus anticis intermediisque supra, pedibus-
que posticis ferrugineis; alis hyalinis, apice et cellula radiali
infumatis, venis nigris ; cly peo plano, apice subemarginato, mar-
gine apicali in medio dente nigro parvo armato ; mesopleuris
haud tuberculatis ; segmento mediano area basali subopaea, i im-
punctata ; segmento ventrali secundo area basali elevata nulla.
Long. 9 mm.
?. Strongly and closely punctured; head very broad,
eyes distinctly divergent towards the clypeus, antennee
inserted about twice as far from the anterior ocellus as from
106 Mr. R. E. Turner on Fossorial Hymenoptera.
the base of the clypeus; cheeks very broad, much broader
than the eyes ; first abdominal segment a little longer than
the greatest breadth ; pygidial area rugulose, elongate-ovate,
rather narrowly truncate at the apex ; second ventral segment
almost smooth, the other ventral segments sparsely and
shallowly punctured.
Hab. S. Queensland, Darra near Brisbane (Hacker) ; De-
cem ber,
Not very near to any other Australian species.
Subfamily Nyssonzvz.
Nysson (Acanthostethus) brisbanensis, Turn.
Nysson (Acanthostethus) brisbanensis, Turn. Ann. & Mag. Nat. Hist. (8)
xy. p. 81 (1915). 9.
3g. The male has the ventral segments bare, without a
fringe of hairs; ventral segments 3-6 with a small but
distinct spine on each side at the apical angles; seventh
dorsal segment widely and rather shallowly emarginate at
the apex, the angles produced into short blunt spines.
Hab, Brisbane (Hacker) ; February.
Differs from nudiventris, Turn., to which species the female
is most nearly allied, in the shape of the seventh dorsal seg-
ment, which only has two spines (one at each apical angle),
also in the presence of a short spine at the apical angles of
the sixth ventral segment. The only specimen sent is very
small, measuring only 4 mm. in length.
Subfamily Crazronrv2z.
Encopognathus brownet, sp. n.
9. Nigra, ubique dense rugose punctata; scapo, callis humerali-
bus, postscutello, femoribus apice extremo, tibiis tarsisque pallide
flavis; tibiis intermediis posticisque infra fuscis; alis hyalinis,
iridescentibus, venis fuscis, stigmate testaceo.
Long. 5 mm.
2. Mandibles excised on the outer margin, acute at the
apex. Clypeus subcarinate longitudinally, produced into
two porrect teeth at the apex, with a smaller tooth on each
side near the apical angles. Eyes not hairy, the facets in
front larger than elsewhere, separated from each other at the
base of the clypeus by a distance equal to about half the
length of the scape; frontal groove smooth and shining.
Posterior ocelli a little nearer to the eyes than to each other,
the ocellar region and the vertex coarsely punctured-rugose,
Mr. R. E. Turner on Fossorial Hymenoptera. 107
an oblique groove from the eyes to the posterior ocelli ; the
hind margin of the head slightly raised, subcarinate. Pro-
notum transverse, the anterior margin raised and sharply
pointed at the angles ; thorax very coarsely punctured, the
mesopleuree coarsely rugose. Median segment short ; witha
distinct enclosed basal area, which is very finely rugulose,
with five strong longitudinal carine ; the posterior slope
rather indistinctly transversely striated; the sides of the
segment very finely and closely punctured. Abdomen
smooth and shining beneath, the three basal dorsal segments
coarsely punctured, the three apical segments closely and
finely punctured. Recurrent nervure received before the
middle of the cubital cell; transverse cubital nervure received
just beyond the middle of the radial cell.
Hab. British East Africa, T'ana River, 3000 ft., near
Mt. Kenia (G. Orde Browne) ; November.
This is distinct both in colouring and in structural details
from E. brauer?, Kohil., also in the sculpture of the abdomen.
Rhopalum tuberculicorne, sp. n.
6. Niger; scapo, tuberculis humeralibus, pedibus anticis, pedibus
intermediis tarsis infuscatis, coxisque posticis apice flavis ; fla-
gello fusco subtus ferrugineo ; segmentis dorsalibus 5-7, ventrali-
bus 2-7, tibiis posticis tegulisque pallide ferrugineis; alis
hyalinis, iridescentibus, venis nigris, cellula radiali infuscata,
Long. 4 mm.
3g. Clypeus without a carina, clothed with silver pubes-
cence, the apical margin almost transverse. Mandibles
blunt at the apex, not bidentate. Second joint of the fla-
gellum longer than the third, emarginate at the base beneath
and produced into a stout tubercle at the apex beneath.
Head smooth and shining; the eyes separated at the base
of the clypeus by a distance slightly exceeding the length of
the scape, strongly divergent towards the vertex ; posterior
ocelli as far from the eyes as from each other, and also as far
from the hind margin of the head as from each other, a short
longitudinal sulcus between them. Pronotum short, a minute
spine at the anterior angles. Thorax shining, microscopically
punctured ; median segment smooth and shining. First
abdominal segment scarcely longer than the second, mode-
rately swollen at the apex, second segment broadened from
the base, third segment broader than long. Hind tibiee
broad, with a few small spines on the outer margin. Re-
current nervure received just before two-thirds from the base
of the cubital cell; radial cell broadly truncate at the apex,
108 Mr. R. E. Turner on Fossortial Hymenoptera.
the costal margin shorter than the stigma, the transverse
cubital nervure received a little beyond one-quarter from the
base of the cell.
Hab. 8S. Queensland, Caloundra (Hacker) ; January.
Nearly related to A. tenuiventris, Turn., but the abdomen
is more slender in that species, the third segment being wuch
longer than broad; in tenuéventris g the scape has a small
spine at the apex, the second joint of the flagellum is rather
more strongly emarginate beneath, and the third joint is also
strongly emarginate beneath and subtuberculate at the apex.
The epicnemial area is defined in both species. In most
Australian species of Rhopalum the male antennee are not
strongly differentiated, but in L. alicia, Vurn., and 2. lepto-
spermi, ‘Lurn., the third jolt of the ‘flagellum is strongly
emarginate beneath and subtuberculate at the apex.
Rhopalum testaceum, sp. n.
Q. Testacea; capite mesonotoque nigris; mandibulis, apice ex-
cepto, clypeoque flavis; antennis testaceis ; alis hyalinis, irides-
centibus, venis fuscis.
Long. 4 mm.
¢. Mandibles acute at the apex, not bidentate ; clypeus
broadly rounded at the apex, with four minute teeth on the
apical margin, without a carina. Head smooth and shining ;
the eyes separated at the base of the clypeus by a distance
equal to rather more than two-thirds of the length of the
scape ; posterior ocelli as far from the eyes as from each
other and about the same distance from the hind margin of
the head ; a curved groove from the inner margin of the
eye, extending towards, but not reaching, the posterior ovelli.
Pronotum depressed below the mesonotum, almost vertical.
Thorax closely and minutely punctured, a transverse groove
at the base of the scutellum. Median segment smooth and
shining, with a distinct median sulcus. First abdominal
segment a little shorter than the second, moderately swollen
at the apex, not very slender; second segment longer than
the apical breadth ; third segment much broader than long.
Hind tibie very feebly serrate near the apex. Recurrent
nervure received a little before two-thirds from the base of
the cubital cell ; radial cell oblique at the apex, the costal
margin as long as the stigma, the transverse cubital nervure
received less beyond one-quarter from the base of the cell.
Hab. N. Queensland, Kuranda (PF. P. Dodd).
Easily distimguished by the remarkable colouring. The
first abdominal segment is shorter than in other Australian
Mr. R. E. Turner on Fossortal Hymenoptera. 109
species except frenchi and macrocephalus, and the hind tibize
are much less swollen than is usual in the genus.
Subfamily Tryroxyroniv a.
Pison deperditum, sp. n.
@. Nigra; mandibulis, palpis, antennis, abdomine pedibusque
rufo-ferrugineis ; tegulis testaceis; alis hyalinis, venis fuscis ;
segmenuto mediano crasse rugoso, sulco mediali lato, transverse
striato.
Long. 7 mm.
9. Clypeus broadly rounded at the apex, clothed with
whitish pubescence, which extends on to the front. Second
joint of the flagellum about equal to the third, nearly twice
as long as the first. Hyes separated at the base of the clypeus
by a distance equal to nearly twice the length of the scape,
and by about the same distance on the vertex ; posterior
ocelli ‘a little further from each other than from the eyes,
separated from each other by a distance equal to the diameter
of one of them, with a shallowly impressed transverse line
behind them. Front with a low carina from the base of the
antennee reaching halfway to the anterior ocellus. Pronotum
transverse, with a narrow depression along the hind margin;
thorax smooth, opaque. Median segment very coarsely
rugose-reticulate, with short oblique striz at the base, and a
very wide transversely striated longitudinal sulcus, the poste-
rior slope irregularly transversely striated. Abdomen micro-
scopically punctured, the segments broadly but very shallowly
depressed on the apical margin. First recurrent nervure
received jast before the apex of the first cubital cell, second
received close to the middle of the second cubital cell ; third
cubital cell on the radius very short, shorter than the petiole
of the second cubital cell,
Hab. Port Darwin, Northern Territory (G. F. Hill).
This is very closely allied to P. ruficornis, Sm., from
which, however, it is easily distinguished by the very different
and much coarser sculpture of the median segment. ‘The
neuration in both species is that of the section Pisonitus,
Shuck.
Pison multistrigatum, sp. n.
Q.. Nigra; palpis testaceis; calcaribus unguiculisque ferrugineis ;
alis hyalinis, apice leviter infumatis, venis fuscis; segmento
mediano fortiter longitudinaliter striato.
Long. 9 mm.
110 Mr. R. E. Turner on Fossorial Hymenoptera.
2. Clypeus broadly truncate at the apex, clothed with
silver pubescence. Head opaque, finely punctured; eyes
separated at the base of the clypeus by a distance about equal
to three times the length of the scape, but by only about half
that distance on the vertex ; second joint of the flagellum
distinctly longer than the third and about twice as long as
the first ; posterior ocelli about.twice as far from each other
as from the eyes. Thorax subopaque, finely and closely
punctured ; the pronotum transverse, a little depressed on
the posterior margin. Median segment very coarsely longi-
tudinally striated; the sides of the segment finely hori-
zontally striated, with fine punctures between the strie ;
posterior slope transversely striated, with a deep median
sulcus. Abdomen shining, very finely punctured, the seg-
ments rather feebly depressed at the apex; second ventral
segment more sparsely punctured in the middle than on the
sides; the apical angles of the dorsal segments with a little
white pubescence. First recurrent nervure received close to
the apex of the first cubital cell, second at the apex of the
second cubital cell, almost interstitial with the second trans-
verse cubital nervure. Third cubital cell shorter on the
radius than the petiole of the second cubital cell.
flab, Nyasaland, Mlanje (8. A. Neave) ; February.
Differs from all other species known to me by the very
strong longitudinal striation of the median segment.
Pison strigulosum, sp. n.
©. Nigra; fronte argenteo-pubescente; mandibulis, femoribus
apice, tibiis tarsisque ferrugineis ; tegulis testaceis; alis hyalinis,
iridescentibus, venis nigris, segmento mediano basi oblique, apice
transverse striato.
Long. 8 mm.
@. Clypeus without a carina, broad, the apical margin
slightly oblique on the sides and forming a distinct angle in
the middle. Head opaque, a distinct frontal suleus reaching
the anterior ocellus. Front broad, the eyes at the base of
the clypeus more than half as far again from each other as
on the vertex. Posterior ocelli nearer to the eyes than to each
other ; second joint of the flagellum distinctly longer than
the third. Thorax minutely and closely punctured, the
pronotum and mesopleure clothed with short silver pubes-
cence. Median segment obliquely striated at the base, the
strize becoming rather finer and more transverse towards the
apex, the apical slope coarsely transversely striated ; at the
base of the segment is a very small triangular space enclosed
Mr. R. E. Turner on Fossorial Hymenoptera. 11g
by sulci; from the apex of the triangle a longitudinal trans-
versely striated groove runs to the apex of the segment, and
is continued after a narrow interruption on the apical slope.
Abdomen finely punctured, somewhat pubescent, the three
basal segments shallowly depressed on the apical margin ;
second ventral segment microscopically punctured, more
finely than the third; second to fourth ventral and third to
fifth dorsal segments very narrowly pale testaceous at the
apex. ‘Third cubital cell as long on the radius as the petiole
of the second cubital cell; recurrent nervures received just
before the first and second transverse cubital nervures.
Hab. Gold Coast, Tamale (Dr. C. E. S. Watson).
This belongs to the group of P. wanthopus, Brullé, but
may be distinguished by the less oblique striation of the
median segment, the colour of the pubescence on the front,
and the shape of the clypeus.
Pison carinatum, sp. n.
@. Nigra; mandibulis in medio fusco-ferrugineis; calcaribus
pallide testaceis ; alis hyalinis, margine apicali leviter infuscatis ;
fronte argenteo-sericeo, abdomine segmentis dorsalibus 1-3
margine apicali albido pubescentibus.
$. Femine similis; tarsis rufescentibus ; segmentis abdominalibus
4-7 rufis; segmento dorsali septimo lato, deflexo, apice sub-
truncato.
Long., 2 7, ¢ 6 mm.
9. Clypeus with a low longitudinal carina on the basal
half, broadly subtruncate at the apex. Head opaque, with a
delicate longitudinal sulcus on the front reaching to the
anterior ocellus. Eyes more than half as far again from each
other at the base of the clypeus as on the vertex ; posterior
ocelli a little nearer to each other than to the anterior ocellus,
further from each other than from the eyes; second joint of
the flagellum a little shoiter than the third. Thorax closely
and minutely punctured, more strongly on the mesopleuree
than on the mesonotum ; median segment finely obliquely
striated, depressed longitudinally in the middle, with a
distinct longitudinal carina, the apical slope transversely
striated, the sides of the segment finely and closely punctured.
Abdomen on both surfaces closely and microscopically punc-
tured; sixth dorsal segment broadly triangular, convex,
subcarinate longitudinally in the middle. The position of the
recurrent nervures and also the length of the third cubital
cell on the radius show much variation in this species.
The male has the clypeus more preduced in the middle
112.) - Mr. R. E. Turner on Possorial Hymenoptera.
than in the female, but has the carina at the base; the eyes
are a little further apart on the vertex, the second joint of
the flagellum is fully as long as the third. The broad form
of the seventh dorsal segment is remarkable.
flab. Ashanti, Obuasi (W. M. Graham), April, Feb-
ruary ; Uganda, Hntebbe (C. G. Gowdey) (type), September
and March; Egypt, Meadi (Egyptian Department of Agri-
culture), July; Sierra Leone (Morgan).
I had identified this species as xanthopus, Brullé, in my
recent paper on Pison (Proe. Zool. Soc. 1916), but since
then have found other specimens answering much better to
Brullé’s description.. The present species may possibly be
obscurus, Shuck., but the type of that species is lost and
the description gives the pubescence of the front as golden, as
in wanthopus.
Pison vanthopus, Brulle.
eae zvanthopus, Brullé, Ann. Soc. Ent. France, ii. p. 408 (1883).
Four females in the National Collection answer well to the
description. They differ from cartnatum and strigulosum in
the bright golden pubescence of the front and in the red
colour of the two or three apical abdominal segments. There
is no basal carina on the clypeus as in carinatus, which it
resembles in the sculpture of the median segment, and differs
in the latter point trom strigulosum. 'The clypeus is more
distinctly truncate at the apex than in either of the other
species. P. clypeatus, Cam., seems to belong to the same
group. I do not think that Shuckard’s description of
obscurus can be meant for the present species.
Hab. N. Nigeria, Kateri (J. J. Simpson), December; Gold
Coast, Aburi (L. Armstrong), April.
Pison flavolimbatum, sp. n.
@. Nigra; segmentis dorsalibus tribus basalibus fascia apicali
flavidula ; scapo tegulisque brunneo-testaceis; fronte, pronoto
segmentogue mediano lateribus pallide aureo-pubescentibus ;
alis subhyalinis, costa late infuscata, venis nigris.
Long. 10 mm.
9. Clypeus convex, broadly truncate at the apex, without
a carina ; lead opaque, a distinct frontal sulcus reaching to
the anterior ocellus; eyes a little further apart at the base
of the clypeus than on the vertex, posterior ocelli as near to
each other as to the eyes; an undulating, low, transverse
ridge separating the anterior from the posterior ocellar
On the External Characters of the Felide. 113
region. Second joint of the flagellum slightly longer than
the third. Thorax opaque, minutely and very closely punc-
tured ; median segment similarly punctured, with a median
longitudinal sulcus, shallow and narrow on the dorsal surface,
deep and broader on the posterior slope, the extreme apex
with a few transverse strize. Abdomen closely and minutely
punctured, rather more strongly on the ventral than on the
dorsal segments ; sixth dorsal segment triangular, convex.
Both recurrent nervures received by the second cubital cell,
the first near one-quarter from the base, the second very near
the apex ; radial margin of the third cubital cell variable in
length, but longer than the petiole of the second cubital cell.
Hab. British Guiana, Issororo (C. B. Williams) ; July.
Three females,
The development of the yellow abdominal fascize, which
are chitinous, varies considerably, being rather obscured in
one specimen. This is quite distinct from P. paraense, Spin.,
which also has yellowish abdominal fascize, but is much
smaller and is without the broad fuscous costal margin of the
fore wing, and differs in other details of colour, also in the
position of the first recurrent nervure.
VII.— On the External Characters of the Felide. By
R. I. Pocock, F.R.S., Superintendent of the Zoological
Society’s Gardens.
Tue facts recorded in this paper are based upon an exami-
nation, extending over many years, of specimens that have
died in the Zoological Society’s Gardens. Although un-
avoidably incomplete, the observations probably, I think,
cover the range of variation in the characters discussed
within the limits of the family.
The Ears.
The ears of the Felidz are very constant in their general
features, so far as my observations have carried me, and do
not differ essentially from those of the typical Viverride.
The bursa is always present and large. Its posterior flap
rises behind the rim of the pinna above, and the anterior
flap is always deeply notched. These features are observable
even in newly-born kittens. The main cartilages also differ
but little from species to species ; but neither in the structure
of the bursa nor of the cartilages have I been able to establish
Ann. & Mag. N. Hist. Ser. 8, Vol. xix. 8
114 Mr. R. I. Pocock on the
any features of systematic value. A more detailed comparison
than I have made may, however, show that such differences
Fig. 1.
ul
Hy
M) lif Hee
4 iN {/ jf ly
ee Wi
A. Left ear of Felis ruffus esquinape, nat. size.
B 4 » jaguarondi, sy hs
Cy
3» salinarum, %
exist. For instance, in /. eyra the excrescence on the antero-
internal ridge overlapping the anterior end of the supratragus
(plica principalis) is rather unusually well developed.
External Characters of the Felide. 115
Usually the ears are rounded at the summit, but in
the species of the genus Velis (s.s.), e. g. F. sylvestris,
ocreata, chaus, and their allies, and also in the lynxes,
¥. lynx, ruffus, caracal, they are more angular and
pointed. In all the lynxes, moreover, the tip is provided
with a pencil of hairs, which are especially well developed in
fF’, caracal and smallest in F. ruffus. In the latter they are
sometimes temporarily absent during the moult; but they
are never absent in F. caracal. That these ear-tufts cannot
be regarded as a generic feature is shown by the frequent
presence of similar but smaller tufts in /’. ocreata, F. chaus,
and F. ornata.
Of all the species known to me, /. jaguarondi* and
F. manul have relatively the smallest aud least conspicuous
ears. Inthe former their smallness, coupled with the general
shape of the narrow head, imparts a decidedly musteline phy-
siognomy to the species. In /. manul the width of the head
and the height of the forehead make the ears appear to be set
very low behind the cheek, and they certainly are never raised
above the summit of the head when pricked +. F. serval
presents the greatest possible contrast to F. manul in this
particular, the ears being large and capable of being closely
juxtaposed on the top of the head when pricked. In no
other species is this power developed to the same extent.
The ear of Acinonyx conforms in shape and structure to
that of other round-eared members of the family Felidz
(Aun. & Mag. Nat. Hist, (8) xviii, pp. 422-423, fig. 2.4, 1916).
The Rhinarium.
The muzzle of the Felidz differs from that of the Viverridz,
Cryptoproctid, Mungotidz, and their allies in being bluntly
truncated, the nose, that is to say, projects to a comparatively
slight extent beyond the lower jaw. ‘This feature, coupled
with the shortness of the jaws, gives a very characteristic
appearance to the face of the Felidae as compared with that
of other Ailuroids in which the muzzle noticeably recedes
from the anterior edge of the prominent rhinarium to the
sloping chin. There is, however, a certain amount of variation
in the Felidae with respect to this character. In all species,
it may be added, the upper lip is cleft by a laterally distensible
and mesially grooved strip of naked skin, confluent above with
* If F. braccata, Cope, as stated, has pointed ears, it is probably not
related to F. jaguarondi as claimed by its describer,
+ So far as my memory serves, the ears in F. manul are rounded and
not angular, as one would expect from the affinity of this species to
typical Felis and to the lynxes, :
w
116 Mr. R. I. Pocock on the
the rhinarium and extending inferiorly to the edge of the lip;
and the median groove impressing this strip passes up the
anterior surface of the rhinarium approximately or actually
as high as the upper rim of the nostrils. The infranarial
& j
A. Rhinarium of Felssylvestris, from the front.
B. 7 a ocreata, ns 5
c. i ‘, 3 from the side.
io} es e serval, from the front.
E. 9 - », from above.
real. es és , trom the side.
G. i + salinarum, from_the front.
iY. <i ye i from above.
if. i Pa i from the side.
portion of the rhinarium, so pronounced in Mungotide and
most of the “lower” Ailuroidea, is either suppressed or
developed to only a small extent.
External Characters of the Felide. 117
In the genus Felis (s.s.), as exemplified by F. sylvestris and
F. ocreata, the rhinarium is comparatively small, exhibiting
from the dorsal view a very narrow naked area beyond the
hair of the summit of the muzzle. Viewed from the front,
Fig. 3.
A. Rhinarium of Felis pardalis, from the front.
B. * a »» from above.
C. i 7 » from the side.
D. i 3» Jaguarondi, from the front.
E. , % :; from above.
¥: 5 i is from the side.
its upper edge is horizontal with a slight median depression
and rounded angles. The median portion below the level of
the nostrils, which are moderately far apart, is acutely angled
inferiorly, and there is no definite strip extending laterally
beneath the nostrils. The rhinarium of an example of
118 Mr: R. I. Pocock on the
F. ocreata from Somaliland differs from that of an example
of F. sylvestris from Scotland in having the area between
the nostrils and the angular portion immediately below it
rather narrower (fig. 2, A, B, C).
The lynxes (f. caracal, F. ruffus esqunape, and F. lynx
isabellinus) have the rhinarium relatively larger and more
prominent than F. sylvestris and F. ocreata, the naked portion
seen from above being less overgrown by the hair of the
muzzle. Otherwise there is no great difference between
them. Inanexample of F. lynx isabellinus the upper margin
seen from the front is more convex than in F. caracal and in
F. ruffus esquinape, and the nostrils are somewhat larger,
possibly in adaptation to a life at high altitudes, where the
atmosphere is more rarified (fig. 4, C).
In the smaller tropical and subtropical Felide of America
the rhinarium is large as compared with that of /. ocreata
and sylvestris, as may be seen by comparing the drawings of
this organ in a specimen of F’. ocreata from Somaliland and
of F. salinarum from Cordova in the Argentine, the cats
themselves being approximately equal in size. In the case
of F. salinarum* the rhinarium exhibits a naked area of
considerable size when seen from above, the nostrils are
wider apart than in F. ocreata, and the infranarial portion
is wider transversely and much less acutely angled inferiorly
(hie! Qs Wee Gada oa).
In F. wiedit the rhinarium is very similar to that of
F, salinarum.
In an example of F. pardalis (fig. 3, A, B, C) from Maiaos
the rhinarium is rather more prominent than in examples of
F. wiedii examined, and has the internarial septum wider, the
edge more convex in profile view, and the upper edge also
more convex when viewed from the front. Nevertheless, the
general similarity between them is unmistakable.
In an adult F’. jaguarondi (fig. 8, D, H, F) from Cordova,
in the Argentine, the muzzle projects, and the hairs on its
summit form a high crest, which anteriorly encroaches in
the middle line on the rhinarium, forming an angular pro-
jection over the middle of its upper side. In profile view
the margin is convex. From the front view the upper edge
is mesially notched by the hairy crest, the intervarial septum
is broad, and the portion below the nostrils deep. But in a
young specimen of the typically-coloured form of this species
* This form, described by Mr. Thomas (Ann. & Mag. Nat. Hist. (7)
xii. p. 239, 1903), is closely related to the hetter-known F. geoffroyt.
Possibly it should be regarded merely as a subspecies,
External Characters of the Felide. 119
these characteristics of the rhinarium observed in the adult
eyra-coloured specimen from Cordova are less marked *.
I have only examined the rhinarium in two of the tropical
Asiatic species, namely F. viverrina and FI’. nebulosa. In the
former the rhinarium is tolerably similar to that of F. pardalis,
but is relatively smaller and less prominent. In F. nebulosa,
on the other hand, it differs but little from the rhinarium of
Panthera t described below.
In the matter of prominence and the great size of the
the naked area, when viewed from above, the rhinarium of
F. serval (fig. 2, D, E, F) surpasses that of all other species
of Felide. From the dorsal aspect it is broadly cordate. In
profile view its margin is rather strongly convex and projects
well beyond the lips. From the front its superior edge is
transverse, with rounded angles; the portion above the
nostrils is high, and the area below them wide, comparatively
deep, and not acutely angled inferiorly. The rhinarium,
indeed, is an exaggeration of the type seen in F. pardalis.
The differences between it and the rhinarium of /. ocreata
and sylvestris are particularly worthy of note.
In an example of F. concolor, three months old, the
rhinarium seen from above exhibits a tolerably extensive
naked tract, although not so large as that of /. pardalis.
Nor is the rhinarium so convex and prominent in profile
view as in that species. Moreover, from the front aspect
the area above the nostrils is deeper, that below them is:
narrower, and the nostrils themselves are closer together.
The rhinarium, it may be noted, is not like that of Panthera,
but in its general features approaches the rhinarium of the
smaller members of the Felidee.
In Panthera leo (fig. 4, A, B) the short hair of the muzzle
spreads over the summit of the rhinarium practically to its
anterior margin, so that there is no naked area, or at most
a very narrow naked area, visible in front of the hair from
the dorsal view. The rhinarium itself is tolerably flat, the
median area below the level of the nostrils is narrow and
acutely angled inferiorly, and there is no definite lateral
infranarial extension, the naked skin forming the lower
margin of the nostril being quite smooth and moist like the
inside of that orifice, which is large and expanded.
So far as my observations go, the rhinarium of P. tigris,
* In the description of F. braccata, Cope records the angulation of the
muzzle-hairs in F. jaguarondi (Amer. Nat. xxiii. p. 144, 1889).
+ For the recognition and definition of this genus, see Ann. & Mag.
Nat. Hist. (8) xviii. pp. 221-229 & 306-316 (1916),
120 Mr. R. I. Pocock on the
Fig. 4.
Ae ee
ate ‘ Vavers ¥ aes
Mit ca Pecans
1 REVS vey
it Viv my fe
2,
/,
7
set
B
A. Rhinarium of Panthera leo, from the front.
B. 4 = » from the side,
C. a Felis lyna, from the front.
Dz. iy » 3) from the side.
External Characters of the Felide. 121
onca, and pardus agrees with that of P. leo. The rhinarium
of Uncia uncia I do not know.
The foregoing account of the range of variation in the
structure of the rhinarium in the Felidee, and a comparison
between that organ in the Felidze and in genera referred
to the Viverridz, show that there is practically a complete
gradation between the two.
In Civettictis civetta*, for example, the rhinarium, which
is of the same type as the rhinarium of Paradoxurus and of
Mungos, is very large and prominent, with the infranarial
portions deep and extending laterally beyond the nostrils.
But in Genetia the infranarial portions are reduced in size ;
and in Linsang ¢ they are so much reduced as to be only a
little larger than in some of the Felide—e. g., F. pardalis,
F. eyra, and F. serval, which also have the rhinarium
tolerably prominent and naked above. The difference,
indeed, between the rhinaria of Linsang and of Civettictis
is greater than between the rhinaria of Linsang and F. par-
dalis ; and from the prominent rhinarium, with its naked
upper side, of /’. pardalis, gradations may be traced within
the Felidze to the wide, comparatively flat rhinarium, with
hairy upper side and suppressed infranarial areas, of Panthera
leo, the species which, with its allies, has the highest type of
rhinarium met with in the A®luroidea.
The Facial Vibrisse.
Amongst the Ailuroid Carnivores, as I have already shown,
the Felidz are exceptional for the complete absence of the
interramal tuft of tactile vibrisse. At all events, I have
never found a trace of this tuft in any specimen of the
many species that have passed through my hands. For the
rest, there is nothing particularly noteworthy about the facial
vibrissee. The mystacial and superciliary tufts are always
well developed. The two genal tufts occupy the normal
position on each cheek, the lower being placed in a line
with the corner of the mouth, and the upper a little higher
up and a little farther back than the lower. In species
with short hair on the cheeks each tuft consists usually of
two or three long vibrisse and is very conspicuous, e. g.
Panthera pardus and F. caracal. But sometimes there is
a reduction in the number. Of two specimens of F. wiedii,
* P.Z. Si 1915, py 306.
ft Ann. & Mag. Nat. Hist. (8) xvi. p. 341, pl. xii. fig. 5 (1915).
422 Mr. R. I. Pocock on the
for instance, one had three bristles to each tuft, the other
only one—a difference I suspect to be due to moulting.
On the other hand, in species with long hair or copious
whiskers on the cheek, like Panthera tigris and Felis lyna,
these vibrissz are not always easy to find. In an example
of the Tibetan lynx, F. lynx isabellina, for instance, each of
the genal tufts was represented by a single bristle mixed up
with the fringe on the cheek. Similariy, in examples of
F. sylvestris and of F. ocreata, recently examined, each
the genal tufts was represented by a single bristle.
The Feet.
In the ‘Annals and Magazine of Natural History’ (8)
Xvll. pp. 419-429 (1916), in a paper dealing with the
external characters of the hunting leopard or cheetah
(Acinonyx jubatus), I described the feet of that Feline, and
compared them with those of the common leopard (Panthera
pardus) to show the differences between them and to illus-
trate the range of structural variation in the feet within
the limits of the family Feld. I stated that the feet
of Acinonyx are distinguished from those of all the other
members of that family by the complete absence of cuta-
neous sheaths for guarding the claws; but added that the
feet of the typical Felide by no méans always conform to
the Pantherine type in the degree of development of these
sheaths. In the following pages I have described and
figured the feet * of several species from the Old World
and the New to show how they differ from each other.
Since the selection is tolerably wide, it does not appear to
me probable that any species of cat exists with feet differing
in any important respect from all of those here discussed ;
but a few interesting species, like F. manul, pajeros, and
planiceps, still remain to be done.
Since in their main characters tne feet here described
agree with those of Panthera pardus, it is needless to repeat
what was said on that head in the paper above quoted +.
* The drawings have been taken from measured feet with the hairs
surrounding the pads cut short, and the feet’are represented as naked
with the digits spread, the axes of digits 2 and 5 being approximately at
right angles.
+ Apart from the forms assigned to Panthera, a genus which I have
elsewhere defined, all the species are provisionally referred to the genus
Felis,
External Characters of the Felide. 123
Genus Fruis, Linn.
Feet of some European, African, and Asiatic Species.
Felis sylvestris *.—he feet are comparatively narrow for
Fig. 5.
A. Left fore foot of Felis sylvestris. x 3.
B. ,, hind foot of _,, Be
C. Right fore foot of Felis serval. #7
Dy ig) o) Mind foot of .:,, 3 7
their Icngth, with smallish pads. In the fore foot the
* It is appropriate to begin with this species, because it is closely
related to, and probably one of the agriotypes of, the domestic cat, Felis
catus, the type of the genus Felis. The feet of the two are similar. The
examples of F. sylvestris examined came from Scotland.
124 Mr. R. 1. Pocock on the
second and third digits are provided with inner lobes to
the claw-sheaths, that of the third being larger than that
of the second. There is, however, no distinct inner lobe to
the claw-sheaths of the fourth and fifth digits. The webs
are moderately well developed. In the hind foot the digits
are without inner lobes to the claw-sheaths, or, at all events,
these lobes are so small as to be negligible (fig. 5, A, B).
F. ocreata has feet almost precisely like those of
F. sylvestris.
Felis serval_—The feet in a general way resemble tolerably
closely those of F. sylvestris, except that the sheaths of the
claws are relatively a little larger, the mner lobe of the third
digit in the specimen examined being exceptionally well
developed and larger than the outer lobe. The carpal pad
also is relatively larger (fig. 5, C, D).
Felis caracal.—The fore foot is tolerably similar to that
of F. sylvestris, but the digits are more separable, the claw-
sheaths somewhat larger, and the webs, particularly those
-joining the second and third and the fourth and fifth digits,
shallower and more emarginate. In the hind foot the third
digit carries a well-developed inner lobe to the claw-sheath,
the plantar pad is longer as compared with its width than in
F. sylvestris, and the webs are much shallower, especially
that connecting the third and fourth digits. The digital
pads, also both of the fore and hind foot, are more pointed
distally than in F. sylvestris and F. serval (fig. 6, A, B).
As I have already remarked (Ann. & Mag. Nat. Hist. (8)
XVili. p. 429, 1916), the hind feet of F. caracal recall those
of Acinonyx jubatus in the emargination of the webs.
Felis lynx isabellinus (fig. 7, C, D).—The feet differ in
some interesting particulars from those of F. caracal. In
the fore foot the plantar pad, owing to the encroachment
of the surrounding hair, is shorter as compared with its
width ; the webs are deeper and hardly differ in develop-
ment from those of F. sylvestris and F. serval; the claw-
sheaths are exceedingly well developed on the second and
third digits, the inner lobe of the third being approximately
as large, relatively, as in F. serval, and there is a distinct
inner lobe on the fourth and fifth digits, that of the fourth
being large, that of the fifth smaller but distinct. In the
hind foot the plantar pad is longer for its width than in
the fore foot, but not so long as in F. caracal. The webs
are rather more emarginate than in Ff. sylvestris and
F. serval, but not nearly so shallow as in F. caracal. As in
the fore foot the claw-sheaths are complete on all the digits,
External Characters of the Felide. 125
the second, fourth, and fifth carrying an inner lobe as well
as the third, that of the fifth being the smallest.
A. Right fore foot of Felis caracal (young). x 2,
B. ,, hind foot of 8) ;
7 ” ”?
Felis viverrina.—The feet are shorter and broader than in
the previously-described species, and in the example examined
126 Mr. R. I. Pocock on the
the carpal pad was exceedingly small. The claw-sheaths are
moderately well developed, but the inner lobe of the second
Fig. 7.
D.
A. Left fore foot of Felis ruffus esqguinape. X 3.
B. ,, hind foot of =: ay s
C. Plantar pad of right fore foot of Felis lynx isabellinus. xX 4
D.
a + hind foot of
” ”? 3?
of the fore foot is comparatively large, and there is a small
inner lobe on the fourth and fifth. Similarly, in the hind
External Characters of the Felide. 127
foot there is an inner lobe, but a very small one on the
second, third, and fourth digits. The webs are developed
to approximately the same extent as in F. sylvestris and
F, serval. They do not conceal the tips of the claws, which
(Unclipped.)
1
x A.
d
”
+ hind foot of _,,
Fig, 8.
C. Left fore foot of Felis viverrina.
” D.
x2
”
”
if "Nees y)
1} “i YR
Mie aa] i v
; l
A. Left fore foot of Felis viverrina.
rary
+) |! °
; 2)
Oo aS)
-—— ~ ~
‘ 3
7 421
z | 5
os
if 2
+4 /
gins
J FE
/ jaa)
project to a certain extent, even beyond the hairs of the
toes, especially on the hind feet, as shown by the sketche
of the unclipped feet (fig..8, A, B, C, D).
128 Mr. R. I. Pocock on the
Felis nebulosa.—The feet are very short and broad with
large pads. In the fore foot the carpal pad is very large
Fig. 9.
A. Right fore foot of Felis nebulosa. X ¥.
B
3 bind ddotar © ,, a 3
and rounded at the apex, the webs are deep and extend
approximately up to the distal ends of the digital pads, and
External Characters of the Felide. 129
on all the digits the claw-sheaths are perfected by the
development of inner lobes. In the hind foot the plantar
pad is very broad, the webs are very nearly as deep as in the
fore foot, and all the digits, as in the fore foot, have well-
developed inner lobes (fig. 9, A, B).
The feet of this species differ from those of F. viverrina
in the larger size of the pads, the much deeper webs, and the
much better developed claw-sheaths. In all these respects
they more resemble the feet of Panthera described below.
The Feet of some American Species.
The feet of an American lynx, probably F. rujjus
esquinape (fig. 7, A, B), from Tampico, resemble those of
F. lynx isabellinus in web-development, but the inner lobes
of the sheaths of the claws are relatively smaller, both on
the fore and hind feet, and the plantar pads are differently
shaped, being markedly longer as compared with their
width. Thus the median length of these plantar pads is
about three-quarters their total width. They are less over-
grown by hair than in F. lynex isabellinus, and recall in
their shape and proportions the posterior plantar pad of
F. caracal.
It may be recalled that Bangs has already pointed out
(Proc. Biol. Soc. Wash. xi. pp. 48, 49, 1897) that the
plantar pads of the lynxes (Ff. ruffus fasciatus, etc.) of
the more southern portions of North America are larger
than those of the Canadian species (F. canadensis). Hence
it may be inferred, I think, that the pads of F. canadensis
probably resemble those of F. lyna isabellinus. The point,
however, to be noticed here is that the three species of
lynxes, namely, F. caracal, F. lynx isabellinus, and FP. rufus
esquinape have feet of the same general form, aud that those
of the Mexican animal are approximately intermediate in
character between the feet of F. caracal and of F. lynx
isabellinus *.
F. geoffroyi—The feet are more robust than those of
F, sylvestris, but are otherwise tolerably similar to them
in the size of the pads and the development of the webs and
of the claw-sheaths. The claw-sheaths are small. In the
fore foot the inner lobe is negligible on the fourth and fifth
digits, small upon the second and larger, but still small,
* The Tibetan lynx has been referred to the subgenus which at
present carries the inadmissible name Eucervaria. That is a mistake,
the skull characters being those of the typical forms, F. lyna and
F’, canadensis.
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 9
139 Mr. R. 1. Pocock on the
upon the third; and in the hind foot the inner lobes are
negligible upon all the digits *.
F. jaquarondi.—In an adult example of the F. eyra-
~ mutation from Cordova, in the Argentine, the fore foot is
Fig. 10.
is
A. Right fore foot of Felis wiedit. x 3.
B.. .,,.. bind foot of »,, , ;
C. ,, fore foot of Felis salinarum. ,,
1}, -«,, - bind foot af -1,, rs 5
shorter than in the example of F. geoffroyi examined, has
the pads relatively larger, and the claw-sheaths better
* Tn all cats the edge of the skin upon the admedian or inner side of
the claw is detached from the claw to a greater or less extent. Hence
the rudiment of the inner portion of the sheath is always present, eyen
in cases where it is stated to be negligible or absent in this paper.
External Characters of the Felide. 131
developed, the inner lobes of the second and third digits
being larger and a small one is present on the fourth. The
webs, however, are developed to approximately the same
extent. The hind feet of the two species are also approxi-
mately alike, except that in F. jaguarondi the claw-sheaths
are a little larger and the third digit shows a small inner
lobe. These differences, however, are less marked in a
aay
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hind foot of ,,
fore foot of ,,
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A. Left fore foot of Felis concolor.
B
OF
kitten of the same type from Cordova and in one of the
dark-coloured forms of which the locality is unknown.
In F, salinurum (fig. 10, C, D) the feet closely resemble
those of F. geoffroyi and F. jaquarondi.
F, wiedit (macroura) (fig. 10, A, B).—The feet are broad
and short, with moderately well-developed pads. In the
fore foot the webs are very deep and extend approximately
Q#
132 Mr. R. I. Pocock on the
up to the distal ends of the digital pads, as in F. nebulosa.
The claw-sheaths also are well developed, with distinct inner
lobe upon the second, third, and fourth digits; but this lobe
is sufficiently small to be negligible upon the fifth digit.
In the hind foot the webs are also well developed, although
shallower than on the fore foot. The inner lobe of the claw-
sheath is negligible upon the second and fifth, but well
developed on the third and distinct though small upon the
fourth digit.
F. pardalis has feet similar to those of F. wiedii (Ann. &
Mag. Nat. Hist. (6) xviil. p. 428, fig. 5, A, 1916).
It is interesting to notice that the feet of F. wiedit and
F. pardalis differ rather markedly from those of F. geoffroyi,
F. salinarum, and F. jaquarondi, and also from those of
F. concolor, though to a slightly less extent, in the develop-
ment of the webs and of the claw-sheaths.
F. concolor (fig. 11, A, B, C).—In an example three
months old the feet are shorter and broader than in
F. geoffroyi, and provided with larger pads and better
developed claw-sheaths. Nevertheless, the webs are de-
veloped to approximately the same extent, and the inner
lobe of the claw-sheaths is small upon the second and
third digits and negligible upon the fourth and fifth of the
fore foot, and also negligible upon the second, fourth, and
fifth of the hind foot. In neither foot do the sheaths encase
the claws almost to the tip, and, in the extent to which the
claws when retracted are exposed, the feet approximately
resemble those of I. geoffroyi, jaguarondi, viverrina, and
sylvestris, and do not conform to the type of foot of
Panthera (fig. 12).
Genus Pantuera, Oken.
In the species of the genus Panthera (fig. 12) examined,
namely, P. pardus, onca, tigris, and leo, the feet are very
much alike. They are short, broad, compact, and difficult
to spread. The plantar and digital pads are large and the
sheaths are well developed, both upon the outer and
the inner side of the claw, and almost conceal the tips
of the claws when retracted. In the fore foot the carpal pad
usually has a widely rounded apex, and the webs extend almost
up to the tips of the digital pads, at least on the admedian
side, and show only a shallow emargination when the digits
are stretched. In the hind foot the webs are less extensive
and more emarginate.
External Characters of the Felide. 133
Fig. 12 depicts the feet of a specimen of P. tigris, three
months old. The feet of P. pardus I have already figured
(Ann. & Mag. Nat. Hist. (8) xvii. p. 424, 1916).
Similar, however, as are the feet of this genus, they do
not differ in important points from those of F. nedulusa, a
Three months old.
(Clipped.)
’
”
x2.
gris.
?
”
Fig. 12,
’
”
hind foot of
fore foot of
”
»
A. Right fore foot of Panthera tt
B.
C.
species which has a more Pantherine skull than any species
of Felis in the sense in which the latter term is used in this
paper.
The facts above described show an interesting series of
134 Mr. R. I. Pocock on the
gradations in the specialization of the feet of Felis in the
following particulars :—
Claw-sheaths —In their simplest form, as exemplified
in Felis sylvestris or F. geoffroyi, these structures differ in
no essential respects from those of some genera of Viverrine,
like Viverra and Genetta.
In their most elaborate form, as exemplified in Felis lynz,
Puanthera tigris, aud others, the inner lobes on all the digits
are well developed and form claw-sheaths, complete extern-
ally and internally, but the extent to which they protect
the tips of the retracted claws depends upon the degree of
retraction of the claw-bearing phalanx by the elastic liga-
ments and upon the length of the sheaths themselves.
Between these two extremes every gradation in the
development of the sheaths may be traced.
Webs.—The webs may, exceptionally, be very shallow, as
in the hind feet of F. caracal, but in aimost all cases they
reach up to the proximal end of the digital pads, at least on
their admedian side; but in other cases they extend beyond
that point, and may, in the case of the front foot, reach
practically to the distal end of those pads on the admedian
side aud exhibit only a slight emargination of the edge, as
in F. tigrina, for example. In the hind feet the webs are
always shorter than in the fore feet, but they exhibit a
similar. progressive series in development from species to
species. In almost all cases well-developed wehs are asso-
ciated with well-developed claw-sheaths. A striking excep-
tion to this, however, is shown by the hind feet of F. /ynz,
where short and deeply emarginate webs accompany claw-
sheaths, which are complete both externally and internally.
It is needless to compare the feet of the Felide with those
of Hyena, Mungos, Galidia, Eupleres, Cryptoprocta, Nan-
dinia, Paradoaurus, and their allies. But a few genera of
fluroids, formerly included in the heterogeneous family
Viverridz, approach the Felidz tolerably closely in tlie
structure of the feet, and, at all events, in the develop-
ment of claw-sheaths, have more “ feline” feet than has
Acinonyx. ‘there is scarcely any difference, for example,
between Genetta* and many species of Felis in the extent
to which the claws are retracted and guarded by cutaneous
sheaths. The same may be said of the feet of Linsang and
Poiana t. But in the structure of the plantar and carpal
pads, the low-set pollex, and the presence of the hallux,
* Proc. Zool. Soc. 1915, p. 186, fig. 3.
+ Ann. & Mag. Nat, Hist. (8) xvi. pp. 342 & S45, pl. xii. (1915),
External Characters of the Felide. 135
Geneita, Poiana, and Linsang have much more primitive
feet. On the other hand, Viverricula has a single cordate
carpal pad, a simpie trilobed plantar pad, and a small pollex
set almost as high as in many Felide. The hind foot, how-
ever, retains a small hallux; and it seems that the mvariable
presence of this digit is the only character that can be
definitely affirmed as distinctive of the feet of the Viverrinz
(Viverra, Viverricula, Civetiictis, and Genetia) when com-
pared with those of the Felide *.
The Anus and External Genitalia,
The auus and the external genitalia, both in the male and
the female, of the Felide present very little variation in
structure. The anus itself opens in the centre of a circular
area of naked skin, and in the female the skin immediately
surrounding the vulva is naked or sparsely hairy; the
perineal region between the two is short, hairy, and un-
modified, aud the clitoris is minute. In the male the
perineal region is also hairy aud unmodified, and the pre-
puce is situated close to the scrotum. The glans penis is
short, subconical, usually armed with backwardly directed
spiny papille, is boneless, or, at most, fortified with a smail
bone, and the urethra opens close to the tip.
In its short unmodified perineum, the shortness of the
glans penis, and the closeness of the prepuce to the scrotum
the ano-genital area of Felidze resembles that of Nandinia
and the Mungotide, and, so far as lam aware, of Hupleres and
Linsang. My acquaintance with the area in Lupleres
and Linsang is, however, restricted to the female, and I
do not know whether the prepuce is close to the scrotum or
not. Fossa is another genus about which very little seems
to be actually known with respect to this region, except that
the perineum is unmodified and that the prepuce, judging
from dried skins, is situated far in front of the scrotum,
a character which must be regarded as primitive in the
Carnivora,
So far as this area is concerned, the Felidz may be dis-
tinguished from Nandinia by the absence of the large scent-
gland situated- in front of the prepuce and vulva in that
genus, from the Mungotidz by the absence of the circum-
aval glandular sac and the situation of the small urethral
orifice at the tip of the glans penis instead of beneath it.
Of the remaining genera of A®luroids, the Viverride
* Very exceptionally the hallux is present in the Felide., I have seen
it in a lioness. ;
136 ‘Mr. S. Hirst on some
(Viverra, Paradoxurus, Cynogale, and their allies) have the
perineal area provided with scent-glands, situated in the
male between the ‘scrotum and the prepuce, which are
widely separated. In Galidictis and Galidia a similar
gland is present at least in the female, the position of the
prepuce being unknown. In the Hyzenas there is a large
sac, receiving the secretion of the anal glands, above the
anus, the prepuce is far in advance of the scrotum, and
the glans penis is long.
Finally, Cryptoprocta, which has even been referred to the
same family as the Felidz on account of the misleading
character of its dentition, has widely different external
genitalia and the anus opening into a large sac.
Thus, if we set aside Linsang, Eupleres, and Fossa, about
which our knowledge is defective, it may ke seen that the
genito-anal area of the Felidz possesses a combination of
characters distinctive of this family of Ailuroidea.
VIIIl.—On some new Mites of the Suborder Prostigmata
living on Lizards. By Stan.iuy Hirst.
(Published by permission of the Trustees of the British Museum.)
Tur Acari described below are forms living on lizards, and,
with the probable exception of Pimeliaphilus tenuipes, they
are all true blood-sucking parasites. The species dealt with
in this prelimimary note will be figured and described in
detail in a later paper on parasitic mites.
Genus Prerycosoma, Peters.
Pterygosoma persicus, sp. n.
? —Body much wider than long. On each side of the
anterior end of the dorsum there are two patches of very
short plumose hairs ; these patches being almost continuous
with one another, the inner one consists of about 5-8 hairs,
the outer of 8-16 hairs. Hairs on rest of dorsum very few
in number ; some distance behind the anterior patches of
hairs there is a transverse row of four plain hairs, which are
short and widely separated from one another ; there are also
two or three rather long fine hairs on each side near the
margin ; posteriorly there are two more pairs of short plain
new Mites living on Lizards. 137
hairs (one pair placed behind the other) ; finally, two pairs
of short plumose hairs situated near the posterior margin,
those of the inner pair being placed on either side of the
genital aperture. On each side of the posterior margin
there is a fringe of about nineteen or twenty very long
hairs, which are quite fine, not being feathered or modified
in any way. Venter with only four pairs of fine plain hairs,
which are of moderate length. There is also a tuft of
eleven very long fine hairs on each side of the genital
aperture. Legs slender and of moderate length; coxe
unarmed, being furnished with long fine hairs.
Length of body ‘6 mm.; its width 1:15 mm.
Hab. Sixty miles north-west of Kermanshaw, Persia ;
a few specimens found under the scales of the tail of
Agamu nupta.
Pterygosoma melanus, sp. n.
? .—This species is not so wide as compared with its length
as is usually the case in the genus. On each side of the
anterior end of the dorsum there is a band of short slender
plumose hairs. Hairs on the rest of the dorsum very few
in number and widely separated from one another ; some
distance behind the anterior margin there are two pairs of
racket-shaped hairs arranged so as to practically form a
transverse row, and a little further back there is another
pair of similar hairs. Posterior margin furnished with a
fringe of about 18-22 hairs, which are rather short and
paddle-shaped, the basal portion being short aud cylindrical,
but the rest of the hair flattened so as to form a rather wide
blade-like expansion, which is striated. Hairs on venter
very few in number, a pair of short plain hairs being
situated immediately behind the mouth-parts, and another
pair of similar hairs in the middle of the body ; posteriorly
there are two more pairs of hairs, which have the distal end
plumose. Legs slender and rather short; coxe unarmed,
being furnished with quite fine plain hairs.
Colour (spirit-specimens) usually black, but sometimes
paler.
Length of body *72 mm.; its width 1-1 mm.
Hab. Deelfontein, Cape Colony ; several specimens found
under axillz and ventral folds of neck of Agama atra.
Klipfontein, Damaraland; two specimens found: on the
same host.
138 Mr. S. Hirst on some
Pterygosoma neumanni, Berl.
Hab. Specimens from Agama colonerum, Gooli Mountains,
Somaliland.
It is probaole that this mite is only a variety of P. agame,
Peters.
GECKOBIELLA, gen. nov.
Body long-oval, being much longer than wide, instead of
wider than long as is the case in the genus Pterygosoma.
Scutum absent. Numerous short plumose hairs are present
on the dorsum and sides. Venter only furnished with very
few hairs. Free portion of peritreme rather long and
directed forwards. Coxe not nearly so much fused to-
gether as is the case of Geckobia and Pterygosoma, and only
furnished with fine hairs.
This new genus is founded for Geckebia texana, Banks ;
as will be seen from the details given above, it is mcre
closely allied to Pterygosoma than to Geckobia, but differs
from the former in the shape of the body, which is longer
than wide, instead of the reverse, and in the structure of the
coxee, which are only slightly fused with one another.
Geckobiella texana, Banks.
Hab. Duval County, Texas; two adult specimens and
numerous larve found on Scelopurus spinosus, var. clarkii
(=S. floridanus).
Genus Grcxosia, Mégnin.
Geckolia latasti, Mégn.
Hab. We have specimens of this mite from Castelfusano,
Ostia, and also from Lisbon and Seville. These examples
were taken from between the toes of Zarentola mauritanica.
Geckcbia clelandi, sp. n.
? .— Body about as wide as Jong. Dorsal seutum well
developed and much wider than long ; it reaches its greatest
width just before the posterior margin, being angular and °
salient at this pomt. The scutum is furnished with ten
hairs, arranged in two transverse rows, an anterior row
composed of four hairs (two being placed close together
on each side) and a posterior row of six (three on each
side of the scutum) ; these hairs are similar in structure
to those on the rest of the dorsal surface, and are fairly
new Mites living on Lizards. 139
long. A short distance in front of the outermost hair
of the hinder row there is a minute rounded structure,
which possibly is an obsolete eye. Hairs on rest of dorsal
surface fairly numerous, but not placed close together ;
they are mostly of moderate length and are club-shaped,
the distal end being enlarged and plumose. Hairs on
venter numerous, bat not placed close together ; most of
them are much smaller than those on the dorsal surface,
and have the distal end plumose but uot distinctly en-
larged; hairs at the sides and hinder end large and club-
shaped, however. On each side of the vulva the integument
forms a large conical process. Plumose hair on second
segment of palp, slender, curved, and not very long,
Legs. Hinder legs not swollen, but they are longer than
the front ones. First coxa furnished with two long fine
hairs, which are not plumose. Coxe 2-4 each with two
short hairs, which are plumose distally (sometimes there
are three on the last coxa). There is a conspicuous club-
shaped hair on the dorsal surface of the femora of the legs,
and a similar but much smaller hair is present on the
anterior surface of the first femur.
Length of body *64 mm.; its width ‘61 mm.
Colour red wheu alive (in spirit yellowish).
Hab. Sydney (ii.16) and Narabeen, New South Wales
(14. xi. 15); specimens from Gymnodactylus platurus forming
part of Dr. J. Burton Cleland’s collection.
Geckobia indica, sp. un.
? — Body much wider than long. Sevtum transversely
elongated, being very much wider than long; its posterior
margin is divided into two rounded lobes by a distinct
indentation in the middle. A minute eye is present on
each side near the anterior margin. There are about
84-46 plumose hairs on the scutum, all of them being
quite short, especially the posterior ones. Similar hzirs
are present in the middle of the dorsum. Hairs at sides
_and posterior eud of moderate length, slender and blunt ;
apparently they are not plumose. Hairs of posterior tufts
of moderate length. Anterior hairs on venter very short
and indistinctly plumose. Hairs on rest of lower surface
long, slender and pointed. Hairs on second segment of the
palp quite slender and plumose. Legs. Posterior legs
longer than the anterior ones, but not much_ stouter.
Spurs on coxz well developed, being large and stout;
there is also a plumose seta on the trochanter and femur
140 Mr. S. Hirst on some
of the fourth leg, but these sete are much more slender
than the coxal spurs.
Length of body *24 mm. ; its width °375 mm.
Hab. Several specimens found under ventral scales of a
gecko (Hemidactylus gleadowi) from Upper Sind.
Geckobia papuana, sp. n.
2? .—Body much wider than long. Dorsum furnished with
numerous hairs. At the anterior end there are two groups,
each consisting of six stout plumose hairs, which are not very
long. Behind them there are numerous very short, pointed,
plumose hairs. Hairs at sides and hinder end of body of
moderate length, slender, and not distinctly plumose. Hairs
of posterior tuft long. Eyes present, but very minute and
inconspicuous. Hairs on venter numerous. Anteriorly
there is a number of very short plumose hairs or spinules.
The rest of the lower surface is densely furnished with
hairs, which are shaped rather like long narrow spear-heads,
being flattened dorso-ventrally and having the point long
and narrow. Last pair of /egs greatly swollen, the anterior
pairs comparatively slender. Coxze armed with stout spurs,
which are curved and plumose ; two spurs are present on
the second coxa, two on the third, and three on the fourth.
There is also a spur on the trochanter and femur of the
fourth leg, that on the femur being placed on a large
protuberance.
Length of body °34 mm. ; its width ‘5 mm.
Hab. Specimens found under ventral scales of a gecko
(Gymnodactylus louisiadensis) from German New Guinea.
Geckobia malayana, sp. un.
? .—Closely allied to G. papuana, sp. n. Body much
wider than long. Dorsum furnished with numerous hairs.
Two groups, each consisting of five rather stout plumose
hairs, which are not long, are situated at the anterior end of
the body, and they are followed posteriorly at a short interval
by a pair of similar hairs. Numerous very short plumose
hairs, which are pointed, are present in the middle area
of the dorsal surface. Hairs at sides and posterior end
long, slender, and apparently not plumose. There is a
minute but distinct eye on the outer side of the group
of stout plumose hairs. Hairs of the posterior tuft very
long and slender. Venter with numerous hairs. Imme-
diately behind the coxe there is a band of very short
new Mites living on Lizards. 141
pointed hairs or spinules; hairs on the remainder of the
ventral surface long and very slender. Legs of fourth pair
much larger and stouter than the others. There is the
usual number of spurs on the legs ; the one on the femur of
the last leg is not situated on a protuberance.
Length of body °28 mm. ; its width ‘49 mm.
Hab. Several specimens found on geckoes (Gymnodactylus
pulchellus) from the Jalor Caves, Malay Peninsula.
Geckobia boulengeri, sp. n.
? .—Body longer than wide and attaining its greatest
width some distance behind the middle. Scutum distinct
and almost triangular (wedge-shaped) ; the anterior margin
is slightly concave and strongly salient laterally. ‘Ten
plumose hairs are present on the scutum, all of them being
very short and stout ; six of these hairs are situated close
behind the anterior margin (almost forming a transverse
line), three being placed on each side. Posteriorly there
are two lateral hairs on each side, one being situated imme-
diately behind the other on the margin of the scutum. A
minute eye is present on each side on : the salient portion of-
the anterior margin. Numerous short plumose hairs are
present on the rest of thg dorsal surface, the anterior ones
being usually rather stout, blunt, and very short ; the others
are more elongated, however. Hairs at the sides and hinder
end of the body slender, fairly long, and blunt; apparently
most of them are not feathered. Hairs of the posterior
tuft long. Venter with very numerous contiguous hairs,
the anterior ones being short and plumose, the others of
moderate length, fine, and not feathered. Legs. Anterior
legs slender, those of the third pair considerably longer
and stouter ; whilst the fourth pair are also long and are
greatly swollen. Short stout spurs similar to thuse present
in G. papuana ete. are present on the proximal segments of
the legs.
Length of body ‘47 mm. ; its width °43 mm.
Hab. A number of examples found on a gecko (Gehyra
yunnanensis) from Yunnan Fu, China.
Geckobia socotrensis, sp. n.
2 .—Body wider than long. Scutum absent. Anteriorly
the dorsum is furnished with numerous very short plumose
hairs, which are slender, pointed, and subequal in length,
none of the anterior ones being enlarged. MHairs at sides
and posterior end of body only of moderate length and
142 Mr. S. Hirst on some
often sinuous ; apparently they are not plumose, Hairs on
venter flattened and scale-like as in G. loricata, Berl., but
much narrower and more elongated (spindle-shaped), and
sharply pointed posteriorly. Distal hair on second segment
of palp short, fairly stout, and plumose. Legs. Cox
furnished with the usual spurs, but they are biunt and
not nearly so strong as in G. loricata; trochanters also
with a short but rather stout seta. All the legs are of
approximately the same thickness, the posterior ones being
the longest.
Length of body *3 mm. ; its width -37 mm.
Hab. A few specimens found under axille of a gecko
(Pristurus rupestris) from Jena-Agahan, Socuotra.
Geckohia loricata, Ber).
Hab. I have examined specimens of this species found
under the ventral scales of specimens of Tarentola maure-
tanica from Lisbon and also from the Riviera.
Geckobia australis, sp. n.
2 — Body wider than long. Scutwm absent. Hairs on
the anterior two-thirds of the dorsum much more uniform
both in size and distribution than in G. loricata, Ber!.,
none of the front ones being enlarged, all being very short.
Posterior hairs on dorsum of moderate length and sometimes
plumose, but the feathering is rather difficult to see. Wentral
hairs flattened and scale-like, most of them being spindle-
shaped and pointed posteriorly ; the posterior ones are more
elongated, however. The hair on the dorsal surface of the
palp is stout and plumose. Posterior /egs longer and stouter
than the anterior pairs, those of the fourth pair being con-
siderably swollen. Coxal spurs large and curved ; there is
also a plumose seta on the posterior trochanters and on the
femur of the fourth leg.
Length of body *36 mm. ; its width *425 mm.
Hab. Several specimens found under ventral scales of a
gecko (Lygodactylus capensis), from Beira, Portuguese East
Africa.
Genus PimeviaPuitus, Tragardh.
Pimeliaphilus tenuipes, sp. n.
? —Body oval, being much longer. than wide. Scutum
triangular, the anterior margin almost straight, being very
slightly concave in the middle, the posterior end bluntly
new Mites living on Lizards, 143
pointed ; the scutum is furnished with six plumose hairs, a
transverse row of four hairs being situated on the anterior
margin ; the other two a little behind the middle of its
length ; these hairs are quite long, being slightly longer than
the scutum. Arrangement of hairs on dorsum the same
as in P. podapolipophagus, Trigardh, and P. insignis, Berl.
First of all, there is an outer hair on each side situated
on the same platelet as the eye, the latter being placed
in front of the hair. There are also four longitudinal rows
of slender plumose hairs, the outer rows each consisting of
two long hairs and a shorter posterior hair, the inner rows
each of three long hairs. On each side of the genital
opening there are two hairs of moderate length and also an
inner border of three short hairs. All these hairs on the
dorsal surface are slender and plumose, and their sockets
are not enlarged. Hairs on venter few in number; there is
a pair of short fine hairs between the last cox, followed
posteriorly by three pairs of plumose hairs. Integument
marked with a sculpturing of very fine wavy lines as in
P. podapolipophagus. Projecting portion of peritreme short.
Chelicera shaped very like that of the species of Geckobia,
the basal part being short, compact, well defined, and
strongly convex dorsally; the rest of the chelicera forming
a long slender style, which, apparently, is not bifid at the
end as in the two known species of Pimeliaphilus, but euds
in a single minute tooth or claw, which is slightly curved.
Palp short: the basal segment is salient laterally and has
a sharp prominent transverse ridge on its dorsal surface ;
second segment dorsally with a long slender plumose hair ;
the next two segments each with a shorter hair, which is
very fine and apparently not plumose. Legs long and
slender, and furnished with numerous fine plumose hairs.
With the exception of the last, each of the coxe has a pair
of short hairs, the inner hair being fine and not plumose,
the outer stouter and apparently plumose. There is also a
forwardly directed plumose hair on the anterior surface of
the third coxa.
Length of body °274 mm.; its width -22 mm.
Colour (in spirit). Body red, but whitish anteriorly and
marked with a pale central line both above and below.
appendages pale. :
Hab. A single example found on a gecko (Gonatodes
albogularis), from Honda, Magdalene River, Colombia.
144 Mr. 8. Maulik on Cassidinee and Bruchide from
IX.—Cassidine and Bruchids [Coleoptera] from the Sey-
chelles Islands and Aldabra. By S. Mavuix, B.A.
(Cantab.).
Tuts paper deals with the material of these groups obtained
by the Percy Sladen Trust Expedition, in 1908-9, in the
Seychelles Islands and Aldabra. Many of the results of
this expedition have been published in special volumes
of the Linnean Society’s ‘Trausactions’ (ser. 2, Zool.
vols. xii.-xvil.), in which series the writer of the present
paper has already reported on the Hispinz of the Seychelles
(vol. xvi. pp. 237-242, 1913).
Chrysomelide.
CASSIDIN &.
This subfamily is represented by two species—Hoplionota
lila, sp. n., and Aspidomorpha apicalis, Klug. The former
is allied to certain Madagascar species, the latter is known.
from Madagascar and Africa. The only member of the
group previously recorded from the Seychelles was Copto-
cycla leopardina, Boheman, known also from Madagascar
aud the Comoro Islands; but this was not obtained by the
Percy Sladen Trust Expedition.
Hoexrronota, Hope.
1. Hoplionota lila*, sp. n.
Quadrate, slightly narrowed behind; as seen in profile
very convex behind the middle, from the highest point
of the convexity a gentle slope towards the head and a
sudden decline towards the posterior extremity ; subnitid.
Head, antenne, prothorax with its lateral expansions, scu-
tellum, the elytral expansions, and the underside orange-red.
Eyes black. Basal half of elytra green, without coste,
apical half dark red. Elytra without spines or tubercles,
Length 5 mm. ; greatest breadth 4.5 mm.
Head not completely concealed under the pronotum,
dorsal surface slightly depressed between the eyes ; viewed
dorsally the vertex is bilobed and slightly projecting; the
antenne are situated under the lobes. Eyes oblong-ovate,
Antenne: joint 1 elongate and distally thickened, joint 2
* A Sanskrit word, used with reference to the green colour of the
elytra.
the Seychelles Islands and Aldabra, 145
small and rounded, joints 3-6 elongate and more slender,
joints 7-1] form a dilated club which is covered with
brownish pubescence. Pronotum twice as broad as long,
front margin more or less serrated, lateral margins rounded ;
surface of disc uneven, impunctate ; the lateral expansions
with large and deep punctures, the centres of which are
more or less hyaline. Scutel/um triangular ; apex rounded,
Elytra: basal portion green, deeply and closely punctate,
centres of punctures red; the green portion of each elytron
is separated from the apical red portion by an oblique costa,
one end of which terminates in a swelling at the middle
of the lateral expansion, the other end joining with an
irregularly-branched costa on the apical red portion of the
elytron ; costz shining ; the apical red portion of the elytra
Hoplionota lila, sp. n.
has the suture raised and is deeply and closely punctate ;
elytral expansions sparsely and deeply punctate, centres of
the punctures more or less hyaline.
Loc. Seychelles: Mahé; Cascade Estate, ca. 800 ft., 1909
(H. P. Thomasset).
Type in the British Museum : described from one example.
H. lila is related to H. thiemi, Weise, H. guerini, Weise,
and H. marginata, Boh., from Madagascar. All of these
are without elytral spines or tubercles, and also have the
basal portion of the elytra without any pronounced costa.
Hi. lila differs from all the others by (1) the orange-red
colour of the prothorax, scutellum, &c., (2) its larger size,
(3) the proportionately greater length of the antennz, (4) the
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 10
146 Mr. S. Maulik on Cassidinsee and Bruchide.
greater sloping of the elytra from the highest convex point,
(5) the more pronounced character of the cost on the apical
portion of the elytra.
AspripomorpHa, Hope.
2. Aspidomorpha apicalis (Klug).
Cassida apicalis, Klug, Ins. Madag. 1833, p. 122; Boheman, Mon,
Cassid. 11. 1854, p. 257. -
Cussida decolorata, Boheman, Cat. Brit. Mus. ix. 1856, p. 144; id.
Mon. Cassid. iv. 1862, p. 547.
Cassida subeurop@a, Thomson; Weise, Deutsche Ent. Zeitschr. 1896,
p- 19; Kolbe, Abh. Senckenb. Naturf. Ges. xxvi. 1902, p. 584.
Var. lutea, Fairm., Bull. Soc. Ent. France, 1896, p. 223; Weise, in
Voeltzkow, Reise in Ost- Afrika, ii. 1910, p. 504.
Loc. Seychelles: Mahé; Cascade Estate, ca. 1000 ft.,
i.-ii.1909. Aldabra (teste Fairmaire and Kolbe). Known
also from Madagascar and widely spread in Africa.
The specimens from Mahé are all from cultivated land.
Several were found, together with a larva and a pupa, on
the leaves of sweet potatoes ([pomea batatas) in Jan. 1909.
After death most dried examples fade from green to a
uniform light yellow.
Cortocycia, Chevrolat.
8. Coptocycla leopardina, Boheman.
Coptocycla leopardina, Boheman, Mon. Cassid. iii, 1855, p. 255; id.
Cat. Brit. Mus. 1856, p. 175; Fairmaire, Ann. Soc. Ent. Belg. xxxvii.
1893, p. 525; Alluand, Cat. Col. Région Malgache, 1900, p. 333.
Not obtained by the Percy Sladen Trust Expedition.
Toc. Seychelles (teste Fairmaire, /. c.). Madagascar,
Comoros.
Bruchide.
Apparently no member of this family has been recorded
hitherto from any of the islands under review. ‘Two species
were collected by the Expedition—one in Seychelles, the
other in Aldabra. According to Pic’s ‘Catalogue of Bru-
chide’ (1913), both are Oriental.
Pacuymervs, Thunberg.
Caryoborus, Schénherr.
4. Pachymerus gonager (Fabr.).
Loc. Seychelles: Mahé, two specimens from Port Vic-
toria, xii. 1908. East Indies (Pic, Catalogue, p. 7). The
Mr, R. E. Turner on Fossorial Hymenoptera. 147
British Museum contains specimens from Bombay, South
India, Ceylon, and Java. Lefroy (‘Indian Insect-Life,’
p. 351) states that this insect is common in India, the larva
living in the seeds, and the adult eating the leaves, of the
tamarind: he refers also to the description of the life-
history by Elditt (1860), who reared the beetle from pods
of Cassia.
SpseRmopuacus, Schénherr.
5. Spermophagus convolvuli (Thunberg).
Loc. Aldabra, xi. 1908 (Fryer), sixteen specimens, seven of
which are stated to have been bred from fruits of Mvolvulus
alsinoides, Linn. Pic’s Catalogue (p.59) records the species
from Ceylon, South Russia (introduced), aud doubtfully
from South Africa.
X.— Notes on Fossorial Hymenoptera.— XXVI. On the
Genus Homonotus, Dahlb. By Rowianp E. Turner,
E25. is.
Family Psammocharide.
Genus Homonorus, Dahlb.
Homonotus, Dahlb. Hymen. Europ. i. p. 35 (1843) (nec p, 441, 1845).
Wesmaelinius, Costa, Prosp. men. Ital. ii. p. 46 (1887).
Hemisalius, Saussure, Grandidier, Hist. Madagascar, xx. p. 315 (1892),
This genus is poor in species, but has a wide range in the
Old World, though apparently absent from America. It
may be distinguished by the convex head, strongly hollowed
behind ; the clypeus prolonged and covering the mandibles ;
the long and somewhat flattened median segment, emarginate
at the apex and with the apical angles produced into stout
spine-like processes ; by the bifid tarsal ungues ; and by the
cubitus of the hind wing originating beyond the transverse
median nervure. Second and third joints of the flagellum
subequal, short. The neuration of the fore wing in the
genus is variable, both in the proportion of the second and
third cubital cells and in the length of the submedian cell,
but the first recurrent nervure is received before the middle
of the second cubital cell. As in many genera of the family
there is a group of identical structure with only two cubital
cells, the second transverse cubital nervure being abseit.
The species I have not seen are marked *, 5
10’
148 Mr. R. E. Turner on Fossortal Hymenoptera.
Homonotus sanguinolentus, Fabr.
Sphex sanguinolenta, Fabr. Entom. Syst. ii. p. 211 (1793).
Salius dorsalis, Sm. Ann. & Mag. Nat. Hist. (4) xii. p. 255 (1878). 9.
This is the type of the genus and occurs throughout
Europe, also ranging as far as Kastern Siberia. Though
the thorax and median segment are usually red in the female,
much variation exists in this respect, the female sometiines
having the thorax and median segment wholly black.
Homonotus ariadne, Cam.
Pompilus (Ferreola) ariadne, Cam, Mem. Manchester Lit. & Phil. Soc.
(4) iv. p. 462 (1891).
Hab. N.K. India; 8. India; Ceylon; Tenasserim.
* Homonotus albistylus, Sauss.
Hemiswius albistylus, Saussure, Grandidier, Hist. Madagascar, xx.
p. 315 (1892). 9.
Hab. Madagascar.
Evidently very closely allied to ariadne, having the same
nervure at the base of the first cubital cell.
FHomonotus exulans, Turn.
Pedinaspis exulans, Tarn. Proc. Zool. Soc. London, p. 338 (1910). 2.
I doubt if this is more than a geographical race of the
Indian Homonctus ariadne, Cam., but the spines at the apical
angles of the median segment are distinctly longer and more
acute in Australian specimens.
flab. Mackay and Kuranda, Queensland; February to
June.
Homonotus nudiventris, Turn.
Pedinaspis nudiventris, Turn, Proc. Zool. Soc. London, p. 389 (1910). 9.
This differs from exulans in the colour of the wings and
nervures, and in the much shorter and blunter spines at the
apical angles of the median segment.
Both species and also H. ariadne, Cam., have the submedian
cell of the fore wing as long as the median, not a little
shorter as in the Kuropean 4. sangutnolentus, Fabr. The
first cubital cell is also pointed at the base, projecting towards
the base of the wing a little beyond the basal nervure, in this
also differing from sanguinolentus.
Hab, Mackay, Queensland ; October.
This may prove to be a seasonal form of evulans,
Mr. R. E. Turner on Fossorial Hymenoptera. 149
Hemonotus egyptiacus, Rad,
Wesmaelinius egyptiacus, Rad, Bull. Soc. Natural. Moscou, p. 478
(1888). 3.
A male in the British Museum from Uganda answers well
to the description, but has the greater part of the femora, as
well as the tibiee and tarsi, ferruginous; the apex of the
abdomen is red from the middle of the third segment. With
this I associate a female from North Riodesia in which the
legs are black, the calcaria whitish, and the abdomen xed
from the base of the third segment. The submedian cell in
this species is slightly longer than the median.
Hab. Mt. Kokanjero, S.W. of Elgon, 6000 ft., Uganda
(S. A. Neave), August; 70 miles west of Kariba Gorge,
N. Rhodesia (Silverlock), June.
I assume that this is the species described by Radosz-
kowski, being the only species of the genus with similar
colouring known to me. I have, however, seen a species
more nearly allied to Planiceps in which the three apical
segments of the abdomen are red in the female ; but this has
a short clypeus and the tarsal ungues are bidentate near the
base, and I do not think it can have been mistaken for a
Flomonotus. It was taken at Harar.
Fomonotus nurse, sp. n.
@. Nigra; mandibulis fusco-ferrugineis; antennis fuscis, subtus
fusco-testaceis ; clypeo apice, tegulisque testaceis ; pedibus fuscis ;
tarsis pallide ferrugineis, articulo basali basi, calcaribusque
albidis ; alis hyalinis, venis basi testaceis, apice fuscis.
Long. 4-6 mm.
?. Clypeus produced over the mandibles, very broadly
rounded at the apex ; second and third joints of the flagellum
subequal. Front strongly convex, temples very narrow, the
eyes nearly reaching the hind margin of the head. — Posterior
ocelli very far apart, about four times as far from each other
as from the eyes. Pronotum scarcely longer than the meso-
notum, much broader than long, narrowed anteriorly. Median
segment emarginate posteriorly, the apical angles produced
into stout and rather blunt spines. First and second ab.do-
minal segments about equal in length, the basal half of the
second dorsal segment rather thinly covered with very short
grey pubescence. The longest calcar of the hind and inter-
niediate tibie a little longer than the basal joint of the tarsi.
First cubital cell narrowly rounded at the base; submedian
cell a little shorter than the median; third abscissa of the
radius longer than the second; first recurreut nervure
150 Mr. R. E. Turner on Fossortal Hymenoptera.
received at about one-third from the base of the second
cubital cell, second just before the middle of the third cubital
cell. Cubitus of the hind wing originating beyond the traus-
verse median nervure.
flab. Deesa, W. India (Nurse) ; April.
This is a smaller species than albocalcaratus, Rad., and has
the third cubital cell longer than the second, not shorter as
in that species ; the colour of the antennz and tarsi is also
different.
Homonotus albocalcaratus, Rad.
Wesmaelinius albocalear atus, Rad. Bull. Soc. Natural. Moscou, p, 472
(1888). 9g.
A single male in the British Museum from Karachi
(Comber) corresponds fairly well with the description, but
the wings are hyaline, not infuscate, and the clypeus is
broadly rounded at the apex, not subemarginate ; but I am
inclined to look on the latter as a sexual difference,
Hab. Orenburg ; Caucasus ; Siberia.
* Homonotus caucasicus, Rad.
Wesmaelinius caucasicus, Rad. Bull. Soc. Natural. Moscou, p.472 (1888),
= es
Hab. Caucasus.
* Flomonotus transcaspicus, Rad.
Wesmaelinius transcaspicus, Rad. Hore Soc. Ent. Ross. xxvii. p. 60
(1893). 9.
Hah. Merv.
* Homonotus steini, Schulz.
Homonotus afinis, Stein, Berlin. ent. Zeit. iii. p, 68 (1869) (nec Pome
puus affins, Ky.= H, sangwnolentus, Fab.).
Pompilus steini, Schulz, Spolia Hymen, p, 168 (1906),
Hab. §.K. Hungary,
Doubtfully distinct trom sanguinolentus.
* Homonotus coste, Tourn.
Wesmaelinus coste, Tourn, Entom. Genev. i. p. 156 (1889). 9 ¢.
Pompilus wettsteint, D, T., Cat. Hym. viii. p. 336 (1897).
Hab. Sicily.
Subgenus GILBERTELLA, nov.
Differs from L/omonotus in having only two cubital cells,
the second transverse cubital nervure being absent.
Type of the subgenus, Planiceps umbraticus, Turn,
Mr. R. E. Turner on Fossorial Hymenoptera. 151
Homonotus ( Gilbertella) umbraticus, Turn.
Planiceps umbraticus, Turn. Proc, Zool. Soc. London, p. 337 (1910). Q.
The second cubital cell is very long, receiving the re-
current nervures near the base and near the apex. As in
other Australian species of Homonotus, the base of the first
cubital cell emits the stump of a nervure into the median cell.
The submedian cell is a little shorter than the median.
Calcaria of the intermediate and hind tibiz very long, slightly
exceeding in length the basal joint of the tarsi.
Hab. Mackay, Queensland; January and February.
Homonotus (Gilbertella) disparilis, sp. n.
3. Niger; antennis subtus, tibiis anticis intermediisque subtus,
tarsisque fusco-ferrugineis; calcaribus albidis; alis fusco-
hyalinis, venis nigris.
Long. 5 mm,
&. Clypeus very broadly rounded at the apex, covering
the mandibles ; second and third joints of the flagellum sub-
equal; front shining, moderately convex. Posterior ocelli
about twice as far from each other as from the eyes ; temples
very narrow. Pronotum narrowed anteriorly, scarcely as
long as the mesonotum ; median segment a little longer than
broad, emarginate at the apex, the apical angles produced
into long stout spines. Second abdominal segment a little
longer than the first; the two apical ventral segments
strongly compressed laterally. The longest calcar of the hind
and intermediate tibie not quite as long as the basal joint of
the tarsus ; hind tibie moderately spinose ; tarsal ungues
rather feebly bifid near the apex. ‘Two cubital cells; the
second abscissa of the radius twice as long as the first;
the recurrent nervures received at one-quarter from the base
and at one-quarter from the apex of the second cubital cell ;
second transverse cubital nervure received just before the
middle of the radius. Submedian cell distinctly longer than
the median; cubitus of the hind wing originating far beyond
the transverse cubital nervure.
Hab. Mlianje, Nyasaland (8. A. Neave) ; May.
The second cubital cell is much shorter than in umbraticus
and the recurrent nervures are received much nearer together.
In umbraticus the second transverse cubital nervure is
received just before two-thirds from the base of the radius,
The spines at the apical angles of the median segment are
longer in this species than in any other known to me.
152 Mr. O. Thomas on
XI.—Notes on the Species of the Genus Cavia.
By OLDFIELD THOMAS.
(Published by permission of the Trustees of the British Museum.)
THE genus Cavia ranges from Venezuela and Guiana in the
north of South America to the pampas of Buenos Ayres
in the south, and extends across the whole breadth of the
continent, from Peru to Pernambuco.
Any examination of the species that exist in this area, and
their correct names, has been rendered very difficult by the
occurrence of such widely different specimens in the same
areas, on which account I have long hesitated to attempt to
work out this puzzling group. Definite cranial characters
seemed almost non-existent, and one appeared to be reduced
to distinguishing the local forms purely by average differ-
ences of size and shades of colour in a group where there is
not a great range in either.
On taking up the subject afresh, however, I find that one
character, observed by Lund in 1838, but overlooked ever
since, detinitely and sharply separates the smaller Brazilian
species from the larger ; and then, these smaller forms being
laid on one side, the whole problem immediately becomes
simplified.
This character is the possession by Cavia fulgida, the
smaller Brazilian cavy, of a deep outer re-entrant angle or
notch at the front end of the posterior lobe of 3 *, this angle
being quite shallow in the larger forms. This notch is so
deep and well defined that there is practically never any case
where one is doubtful as to the allocation of an individual
skull.
When writing about the group in 1901 T, I recognized
Cavia fulgida (under the name of rufescens) by its smaller
size, but, not knowing of the tooth-character, I erroneously
made the small Argentine “ quiso ” a subspecies of it. Now,
however, it is evident that there is no special relationship
between the two.
Taking first the ordinary species without the extra molar
notch, and going from north to south on the Eastern non-
Andean part of the continent, we have in Guiana
* Figured by Lund, K. Dansk. Vid. Selsk. viii. pl. xxv. fig. 15,
+ Ann. & Mag. Nat. Hist. (7) vill. pp. 532-584 (1901).
the Species of the Genus Cavia. 153
Cavia guiane, Thos.
C. porcellus yuiane, Thos. Ann. & Mag. Nat. Hist. (7) viii. p. 152
(1901) ;
and in Venezuela
“ Cavia porceilus venezuel@,” All. Bull. Am. Mus. xxx. p. 250°(1911),
whose distinction from guiane appears most doubtful.
As regards guiune, the original statements about its
characters were largely influenced by the fact that at that
date the few available Brazilian specimens «included both
aperea and what we now know to be the quite distinct
species fulgida, Asa matter of fact, gudane has practically
the same colour as the real aperea, but is distinguished by
its smaller size, the largest of three full-grown. shells onl
measuring 63 mm. in length *, with length of bulla +
11°38: mm. and upper tooth-row 14, The skull is pro-
portionately rather broadly built, with unusually developed
postorbital projecting ledges.
Specimens are in the British Museum from the Kanuku
Mountains, Berbice, and the Moon Mountains, all in British
Guiana.
Putting aside the Cav/a porcellus of Linneus, based on
the Cavia coboya of Marcerave, the domesticated guinea.
pig, to which the name should be restricted, we next have
Cavia aperea, Erxl.
Cavia aperea, Erx]. Mamm. p. 348 (1777) (based on the “ Aperea’’ of
Marcgrave, Bras. p. 223, 1648).
Anema hilaria, Geotf. N. H. Mamm. (fol.) ii. text to pl. 282 (1820).
Cavia leucopyga, Brandt, Mem, Ac. Petersb. 1835, p- 436, pl. xvi.
Size largest of the genus. General colour grizzled
brownish grey, not the clearer or more olivaceous grey of
the Argentine forms. Below dull whitish or drabby whitish,
a clear white spot generally present on the middle line of the
chest just behind the brown collar.
The largest of the available skulls measures no less than
73 mm. in total length, while the average of half a dozen
from Minas Geraes and Sao Paulo is 68°7 mm. in total length,
bulla 11-9, tooth-row 15°5. ‘lhe hind foot in adults varies
from 45 to 50 mm.
* The skull-length is here always taken from the tip of the nasals, and
may sometimes be slightly exceeded by a slanting length from occipital
to gnathion.
+ Measured from-the notch in front of the paroccipital process directly
forwards, parallel with the axis of the skull, not to the antero-internal
angle, which ends in an irregular point,
154 Mr. O. Thomas on
Range from Pernambuco to S80 Paulo; inland to Minas
Geraes. Specimens in Museum from Bahia (Zoological
Society) ; Rio Jordao, Minas Geraes (Robert); Alambary
and Ypanema, Sao Paulo (Robert) ; and Victoria, So Paulo
(Hempel). Recorded by Lund from Lagoa Santa.
The Paraguayan cavy is so similar to C. aperea that I
should probably not have distinguished it, but, as it has a
name, it may provisionally stand as
Cavia aperea azare.
Cavia azare, Wagn., Schr. Siiug., Supp. iv. p. 63, footnote (1843).
Colour, as represented by fresh skins, very much as in
true aperea or rather more olivaceous; size averaging
slightly less, though individual specimens overlap. Averages
of four skulls in greatest length 65°38 mm.; bulla 12:4;
tooth-row 14°9. The bulle would, therefore, appear to be
rather larger, but the number of specimens is not enough to
indicate this with certainty.
Hab. Paraguay. Several specimens from Sapucay (W.
Foster).
Next southwards from C. aperea, in the province of
Parana, a special form was discovered by M. Robert, which
may be described as follows :—
Cavia rosida, sp. n.
Size less than in C. aperea, greatest length of skull about
62 mm. General colour saturate, comparatively dark, nearly
as much so as in C, fulvida. Upper surface grizzled
‘‘ mummy-brown ” ; median area of back heavily blackened
with long blackish piles, especially posteriorly, the middle of
the lumbar region being nearly black. ‘The blackening is,
however, variable and occasionally almost absent. Under
surface dull cinnamon-buff, the hairs pale grey basally ;
usual throat-markings scarcely distinguishable, the inter-
ramia buffy, the usual dark collar overlaid with dull buffy,
and the white chest-patch either absent or reduced to a small
spot. Inner side of limbs like belly.
Skull, as compared with that of C. aperea, smaller and
with conspicuously shorter and slenderer muzzle—in fact, the
skull, apart from the muzzle, is scarcely or not smaller than
that of aperea, the difference in the whole length being
almost entirely due to the reduction of the rostrum. Post-
orbital projections not heavily developed. Bullee fairly large.
Dimensions of the type :—
Head and body 395 mm. ; hind foot 46; ear 20.
the Species of the Genus Cavia. 155
Skull: greatest upper length 62; condylo-incisive length
58 ; zygomatic breadth 35; nasals 19°7 x 8°5 ; interorbital
breadth 12°6; breadth of parietals across brain-case 24°5 ;
diastema 17:4; bulla 12-2 x9°3; upper tooth-series 14°6.
Hab. Serra do Mar, Eastern Parana. Type from Roga
Nova. Alt. 1000 m.
Type. Adult female. B.M. no. 3.7.1.96. Original
number 831. Collected 6th September, 1901, by Alphonse
Robert. Six specimens.
This cavy of the Serra do Mar is readily distinguishable
from C. aperea by its dark colour, blackish back, buffy belly,
reduced chest-markings, aud by the short and_ slender
muzzle of its skull. In the lowlands of the same region, at
Morretes (10 m.), M. Robert found a representative of the
C. fu'gida group.
Next comes the well-known quiso of the Argentine and
Uruguay :—
Cavia pamparum, Thos.
Cavia rufescens pamparum, Thos. Ann, & Mag. Nat. Hist. (7) viii.
p. 538 (1901).
Allied to C. aperea, but smaller; the skull usually about
62-63 mm. in length when adult. Colour as in aperea, but
distinctly more greyish or olivaceous, less brown. Under
surface whitish or slightly drabby, the chest-pattern well
marked.
Skull shaped as in aperea, but smaller ; the muzzle of the
same general proportion, not reduced asin C. rosida.
Range from Corrientes and Uruguay southwards to
Southern Buenos Ayres. Specimens in Museum from “ 20
miles north of Corrientes” (Zurner Henderson); Goya,
Corrientes (2. Perrens) ; Maldonado (Darwin); La Plata
(Thomas) ; Los Yngleses, Ajé, Buenos Ayres (EZ. Gibson)
and Bonifacio, 8.W. Buenos Ayres (2. Kemp).
All the specimens from the above considerable range agree
very closely with each other in size and colcur, no geogra-
phical variation being observable. ‘lwo of Mr. Gibson’s Ajé
specimens, however, out of seven are abnormally larger than
the others, with decidedly larger skulls; but these appear
more or less diseased, and it is possible that they represent an
‘infusion of domestic guinea-pig blood, although there is no
colour indication of this, The other specimens of the same
lot are quite like the ordinary quiso. The size of the bulla
is a little variable, two of the Bonifacio series having this
12:1 aud 11 mm. in length, that of the type being 11°7.
2
156 Mr. O. Thomas on
Passing now to the cavies of the Andean countries, Peru
and Bolivia, we have first to identify Cavza cutler/, Bennett,
the earliest name connected with that region.
The type-specimen, with imperfect skull, is in the British
Museum—no. 53. 8. 29. 2,—and I have carefully examined
it and compared it with the other material in the collection.
It is a melano, and on this account its colour.has never been
able to be used for purposes of identification, while, although
called a “ Peruvian ecavy,” its original locality has always
been doubtful.
The conclusion I come to is that it isa domesticated guinea-
pig— Cavia porcellus, L.,—its skull being too large for any
Peruvian wild species, while it is closely matched by examples
of C. porce/lus, among which, of course, black specimens are
by no means infrequent.
With this troublesome name removed, the ordinary
Peruvian cavy should bear the name of
Cavia tschudii, Fitz.
Cavia cutleri, Tschudi, Fauna Peruana, p. 195 (1845).
Cavia tschudit, Fitzinger, SB. Ak. Wien, lvi. pt. i. p. 154 (98 in sepa-
rates) (1867).
with type-locality Ica, on the coast, where Tschudi saw the
specimens he described.
‘he species is comparatively small, the skull about 58 to
62 mm. in length, and with small bullae. In colour it is
coarsely grizzled cinnamon, buffy or greyish, and the under-
side varies from strongly buffy to nearly white.
These variations appear to indicate four subspecies, as
follows :—
Cavia tschudii atahualpe, Osgood.
Cavia atahualpe, Osgood, Field Mus. Publ. x. p. 98 (1913).
Size fairly large, the bullz larger than in the more southern
forms. Colour dark, “evenly grizzled cinnamon and
blackish, the bases of the hairs dark drab followed by two or
more annulations of cinnamon and blackish”; back, and
especially rump, with numerous longer black hairs ; under
surface more or less cinnamon or buffy. Length of type-
skull 60 mm.
Hab. N. Peru: Cajamarca.
No Peruvian cavies that I have seen have more than one
light annulation on the hairs; but, even if there is no mistake
in the observation, I should not consider it sufficient reason
the Species of the Genus Cavia. 157
to distinguish the North-Peruvian cavy specifically from
C. tschudii, in view of its general agreement in size and other
characters.
Cavia tschudit umbrata, subsp. n.
Size as in atahualpe. Colour greyer throughout, the light
rings on the hairs whitish instead of cinnamon or buffy.
Median area of back blackish, the darkening being effected
not by overlaying with long black hairs, as in atahualpe and
rosida, but by the reduction of the light rings on the hairs,
these being often barely 1 mm. in length, while those in the
other subspecies are about 2-4 mm. as is usual. Bases of
hairs pale slaty. Under surface soiled drabby, the belly and
submaxillary lines of this colour ; collar and middle line of
chin greyish brown. Hands and feet pale brown, lighter
on digits.
Skull of average proportions, the bulle longer than in the
two following subspecies.
Dimensions of the type :—
Hind foot 42 mm. ; ear 20.
Skull: greatest length 60; condylo-incisive length 58 ;
greatest breadth 57:7; nasals 20°58; diastema 18:2 ;
bulla 11°8x 95 upper tooth-series 13.
Hab, Junin, Central Peru. Type from Incapirca, Zezioro.
Type. Adult female. B.M. no. 94. 8.6. 23. Collected
20th June, 1890, by J. Kalinowski.
This Junin subspecies agrees with atahualpe by its
darkened back, rather larger size, and larger bullee, as com-
pared with the two more southern forms that follow. From
atahualpe it differs in general colour very much as pam-
parum differs from aperea, and also in the details of the dorsal
dai kening.
Cavia tschudii tschudii, Fitz.
General colour fairly dark, strongly grizzled, the light
rings on the hairs buffy or ciunamon. Under surface more or
less strongly buffy. Median area of back not darkened.
Skull-length about 59-61 mm.; bull rather smaller than
in the previous subspecies, 10°1-10°9 mm. in length.
Range. Middle Peru, from Iga to Cuzco.
The type-locality is Ica, and a specimen from Tambo, on
the coast opposite Arequipa, agrees so precisely with the
description as to be undoubtedly the same form. Four
specimens from Urubamba, Cuzco, collected by O. Garlepp,
agree absolutely with that from Tambo, while three from
Lia Raya Pass, collected recently by K. Heller, are rather
158 Mr. O. Thomas on
greyer and more or less intermediate between this subspecies
and the next; they are, however, all immature.
Cavia tschudit pallidior, subsp. n.
Similar in general characters to tschudii, but colour much
lighter, the pale rings on the hairs a paler buffy, and the
under surface a pale creamy buff approaching whitish.
Collar a paler grey. Hands and feet buffy whitish, a little
browner proximally.
Skull as in tschudit.
Dimensions of the type (measured in flesh) :—
Head and body 242 mm. ; hind foot 24; ear 29.
Skull: greatest length 59:5 ; condylo-incisive length 54;
zygomatic breadth 33°5; nasals 20°3 x 8:7 ; diastema 16°6 ;
bulla 10°2 x 8; upper tooth-row 14.
Hab. Arequipa. Type from 2500 m.
Type. Adult male. B.M. no.0. 10. 1. 85. Original num-
ber 1023. Collected 31st May, 1900, by P. O. Simons.
Presented by Oldfield Thomas. Six specimens.
Distinguished from tschudit by its lighter coloration
throughout.
Cavia nana, sp. n.
A pigmy cavy, conspicuously smaller than any other
species of the group.
Size very small, skull-length only about 52 mm. Colour
about as in C. tschudit pallidior, the light rings on the hairs
buffy or pale cinnamon ; no darkening along the median
area of the back. Under surface creamy whitish, the grey
collar well marked. Hands and feet pale brown.
Skull about as in C. tschudii, but conspicuously smaller.
Bulle especially small.
Dimensions of the type (measured in flesh) :—
Head and body 215 mm.; hind foot 38 ; ear 23.
Skull: greatest upper length 52; condylo-incisive length
47; greatest breadth 29°5 ; nasals 17°5x 7:5; interorbital
breadth 10°5; breadth of brain-case 22; diastema 13:7;
bulla 9°5 x 7°53 upper molar series, crowns 11°8, alveoli 12-5.
Hab. Highlands of Bolivia. Type from Chulumani,
Yungas, 2000 m. Another specimen from tle Desaguadero
River (J. B. Pentland).
Type. Adult female. B.M.no.1.6.7.59. Original num-
ber 1363. Collected 16th February, 1901, by Perry O.
Simons. Presented by Oldfield Thomas. Four specimens
in all.
the Species of the Genus Cavia, 159
This remarkable little cavy furnishes a good example of
the difficulty of distinguishing young specimens from old
in this group ; for, in spite of the fairly close survey of the
collection always kept up, no one has previously noted that
Mr. Pentland’s specimen, received sixty-six years ago, is
fully adult, and it is only on the general examination of the
group now made that I have found this out, and am able to
give Mr. Pentland the credit for a very interesting discovery.
‘The first scientific explorer of the Titicaca plateau, he sent
home quite a number of interesting specimens, but, of course,
had not been instructed as to the proper preservation of data.
On this account I have chosen one of Mr. Simons’s three
specimens as the type. These were erroneously called
C. aperea in my accouut of the latter’s Bolivian collection.
In proof that the specimens are full grown, I may note
that the type has its basilar suture closed, while Mr. Pent-
land’s specimen has already the tell-tale sagittal crest charac-
teristic of old individuals.
Lastly, we have the Brazilian species with the deep notch
on the outer side of m3 already referred to. ‘There appears
to be only one species of this group, whose name and
characters are as follows :—
Cavia fulgida, Wagler.
Cavia fulgida, Wagler, Isis, xxiv. p. 512 (1831) ; Wagn., Schr. Siug.,
Supp. iv. p. 59 (1843) (redescription of type).
Cavia rufescens, Lund, K. Dansk. Vid. Selsk. viii. p. 282 (1841).
Cavia nigricans and “ Kerodon obscurus, Licht.,” Wagn., Schr. Saug.,
Supp. iv. p. 64 (1843),
Size comparatively small, greatest skull-length rarely
attaining 60 mm. Colour rich dark grizzled brown ; under
surface deep buffy or ochraceous, dulled by the greyish
bases of the hairs showing through to a variable extent.
Last upper molar with a deep indentation on its outer side
at the anterior end of the posterior lobe.
Range from Lagoa Santa, Minas Geraes, to Santa Cathe-
rina ; type said to have been obtained on the “ Amazonian”
journey of Spix *, but the species is not known to occur on
the Amazon.
* Spix’s other explorations were mostly in the region inhabited by the
species I now call C. fulgida, and some error probably crept in as to the
particular trip on which it was collected. Or, with the loose geography
of the time, all his Brazilian journeyings may have been spoken of as
“ Amazonian.” Wagner expressed certainty as to the identity of fulgida
and rufescens, and there appears to me no doubt about it.
160 Bibliographical Notice.
Specimens in Museum from Minas Geraes (Zool. Soc.
Museum); Engenheiro Reeve, Espiritu Santo (A. Robert) ;
Rio Janeiro (Capt. Milner and L. Hardy du Drenenf) ;
Cruzeiro and Piqneté, Sao Paulo (Robert) ; Moretes, Parana
(Robert) ; Humboldt (Larhardt) and Joinville (Behr), Santa
Catherina.
A very distinct species, readily recognizable by its peculiar
m*®, In colour it is not unlike Cavia rostda, but has uot the
special darkening on the back.
BIBLIOGRAPHICAL NOTICE.
Africin Freshwater Fishes,
Wiru the completion of vol. iy. of the ‘Catalogue of Freshwater
Fishes of Africa’ (London, the Trustees of the British Museum,
1916) Mr. G. A. Boulenger has earned the gratitude, not merely of
students of African fishes or of ichthyologists in general, but of all
who are concerned with the problems of geographical distribution.
In these four volumes Mr. Boulenger has described the largest
collection of freshwater fishes ever brought together from one area
in any part of the world, comprising as it does over 15,000 specimens
now in the British Museum and an almost equal number in the
museums of the Nile Survey, the Congo (Tervueren), 8. Africa,
Paris, and Luxemburg.
How immensely our knowledge of the freshwater fishes of Africa
has grown during the last thirty years or so may be gathered from
the fact that in 1880 only 255 species were known. Ten years
ago this number had increased to 974. In the present catalogue
no less than 1425 species are described, and this increase is largely
due to the zeal and enthusiasm of the author of this catalogue, of
which he may well be proud.
Though it would materially have increased the bulk of these
volumes, we venture to think that their value would have been
immensely increased by the addition of internal anatomical
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contributed by the collectors. But there were probably good
reasons for reducing the work to the smallest possible dimensions.
Happily it is well illustrated and has a good index.
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I. Corylophide [Coleoptera] from the Seychelles and Rangoon,
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sity of Cambridge, (Plates I-V.) ....., +3 Ae Gee Ree ©
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EYs Desoriplions of New P yralide of the ‘Subfomnilies Epi- —
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VI. Notes on Fossorial Hymenoptera.—X XV. On new Sphecoidea
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VII. On the External Characters of the Felidae. By R. I. Pococx, 3 ee:
F.R.S., Superintendent of the Zoological Society’s Gardens........ 113
VIII. On some new Mites of the Suborder Prostigmata living on
Lizards, “By Srannux Hagen ss is) Stay seas se tings te ced cee s Se
IX. Cassidine and Bruchide [Coleoptera] from the Seychelles. eee
Islands and Aldabra. By 8. Maurie, B.A. (Cantab.) ........0.4- 144
X. Notes on Fossorial Hymenoptera.—XXVI. On the Genus =
Homonotus, Dahlb. By Rowzanp KE. Turnur, F.Z Su F.E.S. ae ee
XI. Notes on the Species of the Genus Cavia. cs Onprieup
THOMAS —
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No. 110. FEBRUARY 1917.
XII. — Coleoptera, Heteromera (excluding Tenebrionidz)
Jrom the Seychelles Islands and Aldabra*, By Guorce
Cuartes Cuampion, F.Z.S.
‘ [Plate VI.]
Tue material reported upon in the present paper forms part
of the collections made by the Percy Sladen Trust Expedi-
tions of 1905 and 1908-9 in the Seychelles and other islands
of the Western Indian Ocean*. The twenty-six species of
Heteromerous Coleoptera enumerated belong to eight
families, the Monommide, Cistelide (= Alleculid), Melan-
dryidee, (idemeridz, Anthicide, Pedilidz, Xylophilide, and
Mordellide. The Tenebrionide (at present in the hands
of Herr Hans Gebien) are not included. The collections
examined illustrate the abundance of certain Cdemerids,
Xylophilids, and Mordellids (Mordellistena) in the- islands
and the presence of two peculiar Melandryid genera in
the Seychelles. A first set of the material, including the
types of all new forms, will be placed in the British
Museum.
* Many results of this Expedition have been published in a special
series of volumes of the Linnean Society’s ‘ Transactions’ (ser. 2, Zool.
vols, xii—xyii.).
Ann. & Mag. N. Mist. Ser. 8. Vol. xix. > Bh
nian ingéis,
ans? mel,
162 Mr. G. G. Champion on Coleoptera from
List of Species.
Fam. Moncmmide. Fam, Anthicide.
1.. Monomma pruinosum, sp. 0. 11. Anthicus oceanicus, Laf. ©
Fam. Pedilide.
Fam, Cistelide (=ALLECU- on
LID#). 12. Eurygenius fragilicornis, sp. B.
13. 3 eonvericollis, sp. 0.
2. Cacoplesia viriditincta, sp. 1.
3. 4 couilipes, Sp. 1. Fam. Xylophilidz.
14. Xylophilus torticornts, sp. 2.
Fam. Melandryide. 14. a6 clavicornis, 8p. 0.
fc Pa ei G, ; seychellarum, sp. N.
A, Stictodrya (gen. nov.) longi- : y pe
_ _pennis, sp. 0. : Fam. Mordellide.
5. Mycteromimus (gen. nov.) t- a ie
sularis, sp. D. 17. *Mordella brauert, Kolbe.
18. 4 peregrinator, sp. M.
19. M6 disparilis, sp. 0.
Fam, C&demeride. 20. Mordellistena mahena, Kolbe.
21. 5 degressa, sp. N.
6. Oavacis grisescens, Fairm. 22. 5 partilis, sp. 1.
7. 5, lineola, Fairm. 23) = colee, sp. n.
8. Ananca aldabrana, sp. N. 24, e septemcarinata,
9, 5 seabripennis, sp. Nn. sp. D.
10. » submaryinata, sp. 1. | 25. a3 dirempta, sp. 2.
| 26, 5 argutula, Sp. Ws
Fam. Monommide.
MonomMa.
Monommea, Castelnau, Hist. Nat. Ins. ii. p. 215 (1840).
1. Monomma pruinosum, sp. n.
Elliptic, rather broad, feebly shining, nigro-piceous or
black, at most obsoletely rufo-variegate, the reddish colora-
tion sometimes becoming more distinct along the lateral
and apical margins of the prothorax, towards the sides of
the elytra before the apex, and on the humeri beneath, the
autennal club, the palpi, and the legs in part also rufescent ;
somewhat thickly clothed above with minute, fulvous,
adpressed, squamiform hairs, which are condensed into a
small patch on each elytron at the base. Head densely
punctate. Prothorax rounded at the sides anteriorly, closely,
finely punctate, the anterior angles arcuately produced, the
hind angles subrectangular. Elytra transversely gibbous
* Not represented in the collections made by the Expedition.
the Seychelles Islands and Alabra. 163
‘before the middle, with rows of somewhat closely placed,
rather coarse, shallow punctures placed in fine shallow striae,
the punctures becoming coarser and less approximate and
the striz obsolete on the gibbous portion of the disc, the
interstices minutely punctate throughout, moderately convex
towards the sides and apex, and almost flat on the disc.
Beneath closely, finely punctate, the punctures on the
anterior and lateral portions of the metasternum coarse and
scattered ; prosternal process moderately broad, the marginal
earine parallel; fifth ventral segment with a very deep,
bisinuate, transverse sulcus extending across the middle
from the outer margin, interrupted in the centre by a
dentiform backward prolongation of the anterior portion of
the segment.
Length 51-6}, breadth 3!-34 mm.
Loc, Aldabra: Takamaka, x.-xi. 1908 (Fryer).
Kleven specimens, almost certainly including the two
sexes. Near M. irroratum, Kiug, from Madagascar, but
smaller; the vestiture finer and more scattered, not con-
densed into two well-defined densely punctate spots on the
disc of the prothorax (well shown in Klug’s figure) ; the
prothorax more finely punctate; the elytra less dilated at
the sides below the humeri, the humeri more aeute, the
seriate punctures smaller and shallower, the dentiform back-
ward prolongation of the basal portion of the fifth ventral
segment narrower and extending to very near the apex,
Fam. Cistelide (=Alleculide),
CACoPLESIA.
Cacoplesia, Fairmaire, Ann. Soc. Ent. Belg. xlii. p. 237 (1898).
The two species provisionally referred to this genus have
the penultimate joint of the tarsi lobed and the tibial spurs
small, as defined by Fairmaire.
2. Cacoplesia viriditincta, sp.n. (Pl. VI. fig. 1, ¢.)
Oblong-oval, moderately elongate, subopaque, subglabrous,
fusco-testaceous, more or less suffused with green or golden-
green, the head and prothorax being almost entirely of this
colour, the elytra more dilute, the antenne and legs testa-
ceous or obscure testaceous. Head closely, finely punctate,
the epistoma separated from the front by a shallow
groove; eyes large, prominent, slightly smailer in ?,
separated by less than the width of one of os as seen
ES
164 Mr. G. C. Champion on Coleoptera from
from above; last joint of maxillary palpi rather narrow,
elongate-triangular; antenne slender, long in g, shorter
in ?, joints 3-11 subequal in length, 3-10 feebly subserrate.
Prothorax transverse, rather small, arcuately narrowing from
a little behind the middle to the apex, the base broadly sub-
truncate, the hind angles obtuse; the punctuation fine and
sparser than on the head, the interspaces alutaceous. Elytra
much wider than the prothorax, moderately elongate, sub-
parallel in their basal half in ¢@, a little widened posteriorly
in @; deeply crenato-striate. the punctures closely placed,
the interstices convex throughout and very sparsely, finely,
irregularly punctate. Aldeagus of ¢ long, tapering, abruptly
bent at some distance before the slender tip.
Length 74-84, breadth 3-35 mm. (3 ?.)
Loc. Aldabra: Esprit Island, xii. 1908 (Fryer).
Five specimens, the two males having the aedeagus pro-
truding. Allied forms occur in Madagascar, and it is just
possible that the present species may be referable to one of
them. It has the upper surface obviously less shining than
in C. micans, Kiug, cerulans and ceruleovirens, Fairm., &c.,
to judge from the descriptions of those insects.
3. Cacoplesia annulipes, sp.n. (Pl. VI. fig. 2, 3.)
Oblong-ovate, convex, the head and prothorax opaque,
the rest of the surface shining; head, prothorax, and coxe,
and the basal joint of the antenne in immature examples,
obscure ferruginous, the rest of the antennz black, the
elytra brown, the legs testaceous, with the knees, and some-
times the apices of the tibiz also, black ; very finely pubes-
cent, the elytra almost glabrous. Head small, densely,
rugosely punctate, the epistoma confused with the front ;
eyes small, strongly transverse, rather prominent, somewhat
distant from the base of the head; last joint of maxillary
palpi small, subtriangular; antenne extending to beyond
the middle of the elytra, joint 2 small, 3 and 5 equal, 4
slightly longer, 5-10 gradually becoming shorter and stouter,
11 ovate, shorter than 10. Prothorax transversely convex,
short, nearly twice as wide as the head, rounded at the sides,
a little more narrowed in front than behind, subtruncate
at the base, the hind angles obtuse; densely, rugulosely
punctate. Elytra convex, moderately long, about one-half
wider than the prothorax, narrowing from the middle, the
humeri rounded; crenato-striate, the punctures closely placed,
the interstices convex, faintly punctulate. Beneath closely,
the Seychelles Islands and Aldabra. 165
finely punctate. Fifth ventral segment with a shallow trans-
verse depression before the apex.
Length 33-4, breadth 14-12 mm. (¢.)
Loc. Seychelles: Mahé.
Four specimens, apparently all males, from the damp
endemic mountain-forests above Cascade Estate and in the
Mare aux Cochons district (between 1000 and 2000 feet).
This species has the legs coloured as in Plesia geniculata,
Klug, from Madagascar. It will doubtless have to be
removed from Cacoplesia, on account of the small head and
eyes, the small apical joint of the antenne, the convex
general shape, &c. It can hardly be referred to Allecula,
a genus at present including many heterogeneous forms.
Allied insects inhabit Borneo.
Fam. Melandryida.
STICTODRYA, gen. nov.
Head short, small, obliquely narrowed immediately before
the very large prominent eyes and parallel-sided behind
them, the epistoma not separated from the front; labrum
strongly transverse; mandibles small; maxillary palpi
rather stout, the apical joint triangular; antenne very
short, slender, subserrate towards the tip; prothorax trans-
verse, without trace of marginal carina, the base feebly
bisinuate, with distinct foveee; scutellum small; elytra
elongate, much wider than the prothorax, subparallel,
without trace of striz, the punctuation uniform, the epi-
pleura not reaching the apex; anterior cox contiguous,
the cavities open behind ; intermediate coxe well separated;
intercoxal process of abdomen narrow, triangular; ventral
segments rather long, the sutures almost straight ; tibial
spurs minute; tarsi with penultimate joint and the one
preceding it lobed beneath, the former broad, the claws
feebly developed and appendiculate ; body elongate, de-
pressed, the integument rather soft, variegated with lighter
and darker. pubescence.
Type, S. longipennis.
This genus seems to be best placed near Thisias and
various Other forms provisionally referred by me to Melan-
dryide.
The structure of the mandibles cannot be seen in the
unique example obtained. The narrow, immarginate, basally
foveate prothorax, elongate, subparallel, uniformly punctate,
166 Mr. G. C. Champion on Coleoptera from
fasciate elytra, smaller eyes, &c., separate Stictodrya from
Mycteromimus. Fairmaire’s Melandryid-genus Diegoa, from
Madagascar, is compared with Marolia, and it cannot, there-
fore, be very nearly allied to the Seychelles insect.
4. Stictodrya longipennis, sp.n. (Pt. VI. fig. 3, J.)
3. Moderately shining, piceous, with a faint zneous lustre,
the antenne and legs testaceous ; variegated above with
very fine, adpressed brownish and flavo-cinereous pubescence,
the latter condensed into dense patches on the prothorax
and elytra, forming irregular interrupted fascize on the
latter; the head, prothorax, and scutellum densely, finely
punctate, the puncturing of the elytra a little more diffuse.
Head convex, transversely depressed in front, the post-
ocular portion about one-third the length of the eyes
antenne reaching the base of the prothorax, very slender,
joints 8-10 gradually decreasing in length, 2 short, stout,
3 about twice as long as 2,9 and 10 triangular, 11 short-
ovate. DProthorax transverse, a little wider “than the head
with the eyes, the sides rounded anteriorly and parallel
behind, the dise obliquely depressed on each side, the small
basal foveee polished. LElytra about four times the length
of the prothorax, the humeri somewhat oblique in front.
Ventral segments ]—5 closely, finely punctate, simple.
Length 53, breadth 2 mm,
Loc. Seychelles: Mahé.
One specimen, beaten from dense forest-vegetation of
“Capucin ”-trees (Northea), Roscheria-palms, &c., on the
summit of Morne Pilot, over 2000 feet, xi. 1908.
MycrrRromimus, gen. nov.
Head short, obliquely narrowed before the eyes, the
epistoma not separated from the front; eyes very large,
reaching the anterior margin of the prothorax ; labrum
strongly transverse; anteunz short, slender, the outer
joints subserrate, 11 short-ovate, simple ; mentum strongly
transverse ; apical joint of maxillary palpi stout, elongate-—
triangular; mandibles acute at tip, toothed towards the apex
beneath ; prothorax short, closely applied to the elytra, bi-
sinuate at the base, distinctly margined to near the apex
at the sides beneath ; scutelium small; elytra much wider
than the prothorax, oblong, without trace of striz, the
sculpture consisting of intermixed minute and larger punc-
tures, the epipleura not reaching the apex; anterior coxe
the Seychelles Islands and Aldabra. _ 167
small, contiguous, the cavities open behind and closed by
the mesosternum ; intermediate coxz narrowly separated ;
intercoxal process of the abdomen rather narrow, triangular ;
ventral segment 5 simple in both sexes, 2 with a pubescent
tubercle in ¢ ; tibial spurs minute; tarsisparsely pubescent
on their lower surface, penultimate joint broad and lobed
beneath, the claws appendiculate ; body oblong-oval, densely
pubescent.
Type, M. insularis.
The single species from which the above characters are
taken is nearly related to the holarctic genus Mycterus,
some of the members of which have a non-rostrate head; but
it differs from these latter in the still shorter head, the greatly
developed eyes, the elongate-triangular apical joint of the
maxillary palpi, the small scutellum, &c. An unnamed
insect from Madagascar (represented by two broken examples
in the British Museum) is still more closely allied to the
Seychelles insect.
5. Mycteromimus insularis, sp. n. (PI. VI. fig. 4, 2.)
Moderately convex, shining, zeneo-piceous, the legs, mouth-
parts, and antennz (except the intermediate joints in mature
examples) testaceous ; densely clothed with pale brownish
or brownish cinereous pubescence (which almost hides the
sculpture), that on the prothorax transversely arranged ;
the entire surface densely, minutely punctate, with scattered,
irregularly placed, slightly coarser punctures intermixed,
these latter giving an asperate appearance to the elytra when
the vestiture is removed. Antenne with joint 3 about
twice as long as 2, 3-10 gradually becoming shorter and
subserrate, 9 and 10 subtriangular, 11 short-ovate. Pro-
thorax strongly transverse, the sides rounded anteriorly and
parallel at the base. LElytra about four times the length of
the prothorax, the humeri rounded.
3. Ventral segment 2 somewhat gibbous in the middle
anteriorly and at this place bearing a small, fulvo-pubescent
tubercle. :
Length 4-64, breadth 2-24 mm. (¢ 2.)
Loc. Seychelles : Silhouette, Mahé.
Twelve specimens, only obtained by beating from the
growing leaves of one species of endemic palm (Stevensonia
sechellarum) in the mountain-forests: Silhouette, above
Mare aux Cochons, over 1000 feet, ix. 1908; Mahé, near
Morne Blanc, and above Cascade Estate, in both cases at
about 1000 feet. The insect is probably pulverulent in Jite
168 Mr. G. C. Champion on Coleoptera from
like its Palearctic allies. The European Mycterus curcu-
lionoides F., has a similar tuft of hairs on the second ventral
segment in the male.
Fam. Edemerida.
One species of this family is quoted by Kolbe as having
been recorded from the Seychelles by Fairmaire in 1893,
but no name was given ; the insect in question is doubtless
one of those subsequently described by the French author *.
OXxAcIs.
Oxvacis, Leconte, New Species Coleopt. p. 165 (1866); Leconte &
Horn, Class. Coleopt. N. Am. p. 405 (1883); Champion, Biol.
Saad Coleopt. iv. 2, p. 149, and Trans. Ent. Soc. Lond. 1896,
p. 39.
The insects placed under this genus have the mandibles
uncleft at the apex, the right one, at most, with a short
tooth before the tip. Amongst the ten species of Ananca
recorded by Fairmaire from Madagascar or the Seychelles
two, at least, A. grisescens and A. lineola, belong to Oxacis
as here understood, and Lagria livida, F. (selected by
Semenow as the type of Sessinia, Pasc.), from Tahiti, is
congeneric with it. Fairmaire notes the extreme rarity of
the males of some of these Gidemerids.
6. Oxacis grisescens. (Text-fig. 1, g genital armature.)
Ananca grisescens, Fairm. Ann. Soc. Ent. Belg. xli. p. 119 (1897).
Elongate, robust, pale testaceous, the eyes and the tips of
the mandibles black, subopaque, the anterior portion of the
head shining, thickly clothed with very fine pallid pubes-
cence. Head above and between the eyes densely, finely
punctate, the punctuation becoming coarser and diffuse on
the auterior half, the epistoma rather long; eyes very large;
left mandible simple, right mandible toothed before the tip ;
antennee nearly as long as the body in @, a little shorter
in ?, joint 3 distinctly longer than 4, 11 shorter than 10
and feebly constricted at the middle. Prothorax oblong-
subcordate, densely, finely punctate, obsoletely, interruptedly
canaliculate down the middle, the shallow groove terminating
in a deeper, transverse, foveiform depression before the base,
the dise transversely flattened or depressed towards the apex,
* Fairmaire also mentions (Bull. Soc. Ent. Fr. 1893, p. xcix) a Can-
tharid and a Rhipiphorid from the Seychelles, but no names are given.
the Seychelles Islands and Aldabra. 169
without definite foveee. Elytra elongate, subparallel in their
basal half, closely, extremely finely punctate, obsoletely
bicostate on the disc from the base to beyond the middle.
6. Sixth (hidden) ventral segment divided into two long,
inwardly curved, sinuous, concave, forcipiform lobes, the
small seventh segment very deeply emarginate, a long, slender,
pilose rod extruding from the emargination ; edeagus ex-
tremely elongate, slender, thickened at the tip, lateral lobes
long, ciliate, arising from a common stem, which is abruptly
bifurcate from a little beyond the middle. (Text-fig. 1.)
Oxacis (A nanca) grisescens, Fairmaire, §. Gen. armature.
Length 10-12 mm. (4 ?.)
Loc. Seychelles : Mahé Silhouette, Praslin, Félicité,
Bird Island (1905 and 1908-9); Round Island (Mus. Brit.).
The specimens were all found at or near the coast, never in
the endemic forests of the mountains.
Fifteen examples seen, including a ¢ from Round Island,
received by the British Museum in 1870. The very fine
close puncturing of the uppersurface, the long third antennal
joint, the form of the mandibles, and the generally robust
body, distinguish O. grisescens from the allied insects occur-
ring in the Seychelles, whence Fairmaire’s type was obtained.
It is the only one to which his brief description applies.
7. Oxacis lineola.
Ananca lineola, Fairm. Ann. Soc. Ent. Belg. xxxix. p. 453 (1895).
Elongate, shining, finely pubescent ; pale testaceous, the
170 Mr. G. C. Champion on Coleoptera from
eyes and the tips of the mandibles black, the prothorax
with a narrow median vitta and an oblong spot on each side
(the vitta sometimes interrupted and the spot wanting), and
the head in some specimens with a spot between the eyes,
fuscous, the elytra fuscous, with the suture, three uarrow
lines on the dise (the outer one fainter and abbreviated
anteriorly), and the Jateral margin more broadly, pale testa-
ceous, the ventral surface and metasternum in part infuscate.
Hiead moderately produced auteriorly, very finely punctate ;
eyes large, separated by more than the width of one of them
as seen from above; mandibles uncleft at the tip ; antennz
long, joint 3 longer than 4, 4-10 decreasing in length, 11
longer than 10 and feebly constricted at the middle. Pro-
thorax longer than broad, moderately constricted behind the
middle, transversely depressed anteriorly and also hollowed
in the centre before the base; the surface polished, very
finely punctate, with an indication of a smooth median line.
Elytra much wider than the prothorax, somewhat attenuate
posteriorly, closely, very finely punctate, without definite
coste.
¢@. Fifth ventral segment excavate down the middle before
the apex.
Length 7-10 mm. (¢ 2.)
Loc. Aldabra (1908, Fryer). Madagascar (Mus. Brit.).
Four specimens, one only of which (a ? ) is from Aldabra,
apparently referable to A. lineola, Fairm., the type of which
was from Madagascar.
ANANCA.
Sessinia, Pascoe, Journ. Ent. ii. pp. 45, 488 (1863) (nomen nudum),
Ananca, Fairmaire et Germain, Ann, Soc, Ent. Fr. 1863, p. 267.
Copidita, Leconte, New Species Coleopt. p. 164 (1866) ; Champion,
Biol. Centr-Am., Coleopt. iv. 2, p. 144, and Trans. Ent. Soc. Lond.
1896, p. 40.
This genus differs from Owacis in having both mandibles
cleft at the tip. No type was given by Pascoe for Sessznda
and his name cannot be accepted. The five species referred
to Ananca by Fairmaire and Germain were all from Chile ;
the first of these, Nacerdes pallens, Sol., which must be taken
as the type, proves to have bifid mandibles, and the name
Ananca, therefore, must be adopted in place of Cojidita,
used by me elsewhere.
8. Ananca aldabrana, sp. n.
Elongate, luteo- or fulvo-testaceous, the eyes and the tips
the Seychelles Islands and Aldabra. 171
of the mandibles black, subopaque, the head shining, thickly
clothed with rather coarse pallid pubescence. Head mode-
rately produced in front, the epistoma rather long, coarsely,
closely punctate; eyes large; mandibles each bifid at the
tip ; antennz not reaching the apex of the elytra, joints 3
and 4 subequal in length, 11 feebly constricted at the middle.
Prothorax considerably longer than broad, subcordate,
narrow, densely, coarsely, subconfluently punctate, broadly
depr essed and subfoveate on each side of the disc anteriorly.
Elytra long, nearly twice as wide as the prothorax, somewhat
convex, closely, finely, scabroso-punctate, each with two
distinct coste on the disc and another near the outer margin,
all three extending from the base to near the apex.
¢. Sixth (hidden) ventral segment divided into two
narrow, curved, concave, forcipiform lobes, the corresponding
dorsal segment similarly shaped; sdeagus long, rather
stout, gradually widened towards the tip, the latter rounded,
lateral lobes very long, slender, and feebly curved.
Leng h 8-12 mm. (¢ 2.)
Loc. Aldabra (1908-9, Fryer). Seychelles: Round
Island (Mus. Brit.).
Found in abundance at Aldabra, in several parts of the
atoll. In the British Museum there are also two females
aud a male of the same species from Round Island. Very
few males are contained in the long series before me; three,
however (including the one from Round I.) have been
identified, and their genital armature examined. ‘The bifid
mandibles, the densely, rather coarsely punctured, dull, sub-
hifoveate, narrow prothorax, and the finely punctate, more
distinctly costate elytra, readily separate A. aldabrana from
Oxacis (Ananca) grisescens, Fairm. A. (Sessinia) andrewst,
Arrow, from Christmas Island, under which two species
were confused by the author, has the terminal joint of the
antennz almost divided into two, the head much smoother,
the prothorax non-foveate, and the elytra sharply bicostate
on the dise, with the rest of their surface very finely, closely
punctate.
9. Ananca scabripennis, sp. n.
(Text-fig. 2, d genital armature.)
Sessmia andrewst, Arrow, Monogr. Christmas Isl, p. 107 (1900)
(25 neces):
Elongate, testaceous or obscure testaceous, the eyes and
the tips of the mandibles black, subopaque, the head and
prothorax shining, finely pubescent. Head moderately
172 Mr. G. C. Cliaampion on Coleoptera from
produced in front, sparsely, rather coarsely punctate, the
punctures becoming more crowded towards the base ; eyes
large, separated by about the width of one of them as seen
from above ; mandibles each bifid at the tip ; antennz nearly
as long as the bodyin g,, shorter in ? , joints 8-10 decreasing
very slightly in length, 11 slightly longer than 10 and feebly
constricted at the middle. Prothorax longer than broad,
narrow, subcordate, rather sparsely, moderately coarsely
punctate, the disc excavate on each side of the middle
anteriorly and also in the centre before the base, appearing
trifoveate. Elytra long, closely and rather coarsely scabroso-
punctate, each with two faint coste on the disc and another
near the outer margin.
6. Sixth (hidden) ventral segment divided into two,
curved, concave, comparatively short lobes ; edeagus mode-
rately long, gradually narrowed at the apex, the long
narrow tegmen divided into two slender, acuminate pro-
cesses (lateral lobes) from about the middle. (Text-fig. 2.)
Fig. 2.
Ananca scabripennis, Champion, g. Gen. armature.
Length 8-10 mm. (¢ 2.)
Loc. Seychelles: Mahé, Silhouette, Praslin (1905,
1908-9). Christmas Island (Mus. Brit.). ‘The examples
from the Seychelles were all taken near the coast, not iu
the endemic mountain-forests.
‘'welve specimens, apparently all females but one. Various
female examples from Christmas Island placed by Arrow
under his Sessinia andrewsi doubtless belong to this species.
They differ from his type (¢,not ¢ as stated) in having the
the Seychelles Islands and Aldabra. 173
elytra roughly sculptured and obsoletely costate, the pro-
thorax subtrifoveate, the apical joint of the antennz feebly
constricted, &e.
10. Ananca submaryinata, sp. n.
6. Moderately elongate, narrow, depressed, shining,
finely pubescent ; pale testaceous, the tips of the mandibles,
the eyes, an oblong spot on each side of the prothorax, and
an evanescent submarginal stripe on each elytron (extending
from the humeral callus to beyond the middle), black or
piceous ; the entire upper surface closely, very finely punc-
tate. Head slightly produced anteriorly ; mandibles each
bifid at the tip ; eyes large, separated by considerably more
than the width of one of them as seen from above ; antennz
slender, extending to a little beyond the middle of the elytra,
joints 3-5 subequal, 6-11 distinctly shorter, 11 feebly con-
stricted. Prothorax longer than broad, subcordate, slightly
hollowed on each side of the disc anteriorly and also in the
middle towards the base. LElytra comparatively broad,
moderately elongate, subparallel, faintly bicostate on the
disc for about three-fourths of their length, the punctuation
a little finer and more diffuse than that on the prothorax,
ideagus (as seen completely everted) long, bisagittate at
the apex, the outer portion of the sheath also sagittate and
divided at the tip into two slender acute processes ; lateral
lobes widely separated from the base, extremely elongate,
slender, and ciliate.
Length 7 mm.
Loc. Aldabra : Takamaka, xi. 1908 (Fryer).
One male. A rather slender form, with a spot on each
side of the prothorax and a submarginal streak on each
elytron infuscate, the eyes widely separated, the upper
surface shining and finely punctate, the mandibles bifid at
the tip.
Fam. Anthicide.
ANTHICUS.
Anthicus, Paykull, Fauna Suecica, i. p. 253 (1798).
A cosmopolitan genus represented in all parts of the
world.
174 Mr. G. C. Champion on Coleoptera from
1]. Anthicus oceanicus.
Anthicus oceanicus, Laferté, Monogr. Anthic. p. 170; Fairm., Rev. et
Mag. Zool. 1849, p. 452; Pic, Ann. Soc, Ent. France, 1894, p. 6C8 ;
Alluaud, Hist. Madag., Coleopt. p. 487 ; Kolbe, Mitteil. Zool. Mus.
Berlin, v. p. 27.
Loc. Seychelles: Bird Island, vi. 1908 (Fryer). Mar-
quesas ; ‘Tahiti; Polynesia.
Three specimens are before me from Bird Island ; this is
one of two small coral-islands situated on the north of the
Seychelles Bank, but which have neither the peculiar flora
nor any of the physical features characteristic of the other
islands of the group. Recorded by Pic as having been found
in numbers by M. Ch. Alluaud in the Seychelles in April,
1892, beneath seaweed on the coast.
Fam. Pedilide.
Lurygenius, Laferté, Monogr. Anthic. p. 1 (1846).
The known species of this genus are mostly from North
or Central America ; two from Madagascar, however, have
been described by Fairmaire, one from Japan by Lewis, one
from E. Africa and another from Bengal by Pic, one from
the Nilgiri Hills by myself, and one from E. Africa by
Kolbe.
12. Eurygenius fragilicornis, sp.n. (Pl. VI. fig. 5, 2.)
Elongate, narrow, somewhat shining, zneo-piceous or
piceous, the anterior portion of the head rufous, the basal
joint of the antennz, the mouth-parts (the tips of the man-
dibles excepted), the humeri, femora, and the tibiz in part
or entirely, testaceous ; somewhat thickly clothed, the legs
and antenne included, with rather long, semierect, pallid
pubescence. Head densely, rugulosely punctate, obliquely
narrowed behind the eyes, the latter extremely large,
rounded, very feebly emarginate in front, coarsely facetted ;
mandibles entire; maxillary palpi with terminal joint stout,
securiform, the two preceding joints angulate within ;
antennz about half the length of the body, a little shorter
in 2, very slender, joints 3-10 elongate-obconic, subequal
in length, 2 shorter than 3, 11 slightly longer than 10 and
constricted beyond the middle. Prothorax narrower than
tlie head (with the eyes), transversely orbicular, the narrow
neck-like anterior portion rather long, the entire surface
densely, rugulosely punctate. LElytra elongate, much wider
the Seychelles Islands and Aldubra. . We
than the prothorax, parallel in 2? , broader at base and some-
what attenuate in ¢ ; closely set with subseriately arranged,
coarse, oblong, foveiform punctures, the narrow interspaces
minutely punctate and here and there transversely confluent.
6. Femora stouter than in 9, the hind tibize more
curved ; fifth ventral segment unimpressed, simply truncate
at tip.
Length 6, breadth 14 mm. (¢ ?.)
Loc. Seychelles: Mahé.
One pair, taken (4. 11. 1909) on the precipitous slopes of
the peak of Morne Seychellois at an elevation of about
2000 feet ; the specimens were obtained by sweeping a dense
low growth of native ferns and shrubs (Melastoma, Rubus,
Senecio sechellensis, &c.), among which were occasional
Roscheria-palms and other small trees. The coarsely and
closely foveato-punctate elytra separates the present species
from the Madagascar forms. ‘The single representative
from the adjaceut island of Silhouette cannot be treated as a
variety of it. .
13. Eurygenius convericollis, sp. n.
g. Elongate, rather narrow, opaque, the elytra and under
surface somewhat shining ; nigro-piceous, the epistoma and
scutellar region rufescent, the two basal joints of the antenne,
the mouth-parts (the tips of the mandibles excepted), femora,
aud tarsi in part, testaceous ; somewhat thickly clothed, the
antennee and legs included, with rather long, pallid, coarse,
semierect pubescence. Head, palpi, and eyesas in Z. fragili-
cornis, the antenne a little shorter and stouter, about as
long as in ? of that species. Prothorax distinctly broader
than the head (with the eyes), much narrowed behind.
Elytra broader than in E. fragilicornis, narrowing from the
base, the narrow interspaces between the subseriately
arranged foveiform punctures more rugose, giving a dull
appearance to the surface. Legs stout, the posterior tibize
feebly curved. Fifth ventral segment broadly hollowed
down the middle, truncate at the apex.
Length 6, breadth 13 mm.
Loe. Seychelles: Silhouette, viii. 1908.
One male example.
Fam. Xylophilide.
XYLOPHILUS.
NXylophilus, Latreille, Fam. Nat. Régne Anim. p. 383 (1825).
A genus almost cosmopolitan in its distribution. Three
176 Mr. G. C. Champion on Coleoptera from
species are represented in the Seychelles collection, all of
them apparently being fairly common insects in the Islands.
Mr. Scott notes that “ many of them were swept from grass
and other low-growing vegetation.”’ So far as known, they
are wood-feeders in their earlier stages. Owing to their
extreme fragility, very few of the specimens obtained are
in good condition, Upwards of a dozen species have been
described from Madagascar, three from Mauritius, one from
Bourbon, &e.
14. Xylophilus torticornis, sp. n.
(Pl. VI. figs. 6 ¢, 7 ¢ antenna.)
Rather short, moderately shining, very finely cinereo-
pubescent, black, the tips of the tarsi, and sometimes that
of the eleventh antennal joint also, reddish. Head short,
together with the eyes broader than the prothorax, finely
punctate; eyes moderately large, occupying nearly the whole
of the sides of the head, feebly emarginate, distant ; antennze
(fig. 9) moderately long, closely setose, somewhat twisted,
stout, joints 2 and 3 short, 4-10 broad, perfoliate, very
strongly transverse, 6-8 wider than the rest, 11 stout, ovate,
about as long as 9 and 10 united. Prothorax trausverse,
convex, rounded at the sides, closely, rather finely punctate,
bi- impressed on the dise posteriorly. Elytra much wider
than the prothorax, subparallel in their basal half, closely, -
rather coarsely punctate, obliquely depressed on the disc
below the base. Legs short; posterior femora moderately
thickened, obsoletely sulcate beneath ; basal joint of pos-
terior tarsi slightly curved.
Var. a. Duller, the punctuation denser and coarser,
Var. 8. Shining, the punctuation more scattered than in
the type.
Length 1-1} mm. (¢ 2.)
Loc. Seychelles: Silhouette [type]; Mahé [var. a] ;
Praslin [var. 8].
Kighteen specimens—twelve of the form from Silhouette
selected as type, two of the var. a from Mahé, and four of
the var. 8 from Praslin, three of these much ‘smaller than
the rest. The examples from Silhouette were found in the
forest near the Mare aux Cochons plateau, ix. 1908; the
two from Mahé are from high elevations in the forests of
Morne Blane and the Mare aux Cochons district; those
from Praslin were collected on Cétes d’Or Estate, xi. 1908.
The females appear to have the antenne a little less widened
and the eyes rather smaller than in the males. A species
the Seychelles Islands and Aldabra. Lite
recognizable by the somewhat twisted antenne, due to the
joints 6-8 being more dilated than those preceding or
following.
15. Xylophilus clavicornis, sp. n.
(Pl. VintiesGe)
Rather short, feebly shining, piceous or nigro-piceous, the
tarsi, the bases of the tibiz, and the antennal joints 2-9 and
the tip of 11 testaceous ; the prothorax and elytra in fresh
specimens variegated with sharply defined, irregular patches
of very fine grey pubescence (tending to form an interrupted
median and subapical fascia on the elytra), the rest of the
vestiture brown. Head short, together with the eyes broader
than the prothorax, densely, finely punctate; eyes large,
occupying nearly the whole of the sides of the head, distant,
almost entire; antenne rather short, sparsely setose, joints
2-8 each longer than broad, 2 nearly as stout as 1, 3 more
slender, 4—8 scarcely stouter, 9-11 wider than those pre-
ceding, 9 transversely subtriangular, 10 broader, strongly
transverse, 11 stout, acuminate-ovate, about as long as 9
and 10 united. Prothorax transverse, convex, somewhat
rounded at the sides, densely punctate, and with an inter-
rupted arcuate depression on the disc before the base.
Elytra much wider than the prothorax, slightly rounded at
the sides, densely, rather coarsely punctate, feebly depressed
on the disc below the base. Legs short ; posterior femora
moderately thickened, obsoletely suleate beneath; basal
joint of posterior tarsi feebly curved.
Length 1-1} mm. mea)
Loc. Seychelles: Silhouette, Mahé, Praslin.
Found in profusion in Silhouette and Mahé, sparingly on
Praslin. Most, if not all, of the specimens are from the
mountain-forests, from a number of different places and
elevations : one was taken from a rotten and fungus-grown
fallen trunk of the endemic ‘‘ Bois Rouge”? (Wormia ferru-
ginea). Many of these examples are now in bad condition,
very few having the cinereous markings intact. Recogniz-
able by the slender, nigro-clavate antenne, with stout second
joint, the variegate legs, and the densely punctured, cinereo-
maculate surface. The antenne seem to be a little shorter
in the females. The variegate vestiture of the elytra is
common to many species of the genus. The beautiful ex-
ample figured was accidentally injured by the artist after
the drawing was completed and finally corrected.
Ann, & Mag. N. Hist. Ser. 8. Vol. xix. 12
178 Mr. G. C. Champion on Coleoptera from
16. Xylophilus seychellarum, sp.n. (Pl. VI. fig. 8, 3.)
- Rather short, moderately shining, nigro-piceous or piceous,
the base and tip of the antenne, the palpi, the base of the
prothorax in the middle in some examples, the humeri or
base of the elytra, and the legs testaceous, clothed with a
fine sericeous pubescence. Head skort, together with the
eyes much wider than the prothorax, finely punctate; eyes
large, occupying almost the whole of the sides of the head,
separated anteriorly by about half the width of one of them
in 6, more distant in ?, feebly emarginate; antenne (2 )
finely pubescent, long, slender , joint 2 short, 8-10 moderately
elongate, becoming gradually shorter and wider, 9 and 10
subtriangular, 11 stouter, obliquely acuminate, nearly as
long as 9 and 10 united, (2) similar, but much shorter.
Prothorax convex, broader than long, parallel-sided at the
base, closely, finely punctate, with au interrupted arcuate
depression on the disc behind. Elytra convex, rather short,
at the middle about twice as wide as the protiorax, slightly
rounded at the sides; closely, moderately coarsely punctate,
obliquely depressed on the disc below the base. Legs rather
long, slender; posterior femora moderately incrassate,
simple in both. sexes ; basal joint of posterior tarsi feebly
curved.
Length 14-1} mm. (4 2.
Loe. Seychelles : Silhouette, Mahé.
Most of the examples were collected in the Fomretes but
in Silhouette at least one was taken in the low country.
Highteen specimens, varying a little in colour, immature
examples having the elytra paler. In this species the
antenne are moderately elongate in ¢,shorter in 2 , slender
and very gradually widened outwards to the stouter apical
joint in both sexes. The type of coloration is common to
many members of the genus, some of which have peculiarly
formed posterior femora in @.
Fam. Mordellide.
Morbet.a.
Mordeila, Linneeus, Syst. Nat. 10th ed. 1. p. 420 (1758).
17. Mordella braueri.
Mordella braueri, Kolbe, Mitteil. Zool. Mus. Berlin, y. p. 27.
Loc. Seychelles: Mahé (Brauer).
This insect is described as deep black and albo-maculate ;
the Seychelles Islands and Aldabra. 179
the prothorax with six spots—two, semilunate, on the ante-
rior portion and four, confluent, near the base; the elytra
with four spots—one sub-basal, median, one submarginal,
posthumeral, one subsutural, a little before the middle, and
one anteapical. “It is compared with the Asiatic M. com-
posita, Walk., and the African M. elegans, Mak.
The unique example known measures 14 mm. in length.
18. Mordella peregrinator, sp. n.
(PEV Ts fie, es)
Robust, elongate, cuneiform, deep black, the anterior coxe,
the anterior femora in part, and the palpi testaceous, the an-
tenn often fuscous, with the base testaceous ; clothed with
black and whitish or cinereous pubescence, which is con-
densed into the following sharply defined markings :—'The
head with a large patch on the middle of the vertex, and the
prothorax with an interrupted median vitta and a large tri-
angular mark on each side of it, infuscate or black, for the rest
whitish, cinereous, or flavo-cinereous; the elytra with three
strongly angulate narrrow fascize—one sub-basal, irregularly
branching forwards (enclosing an oblique oval spot on the
disc, an oblique humeral streak, a common postscutellar
patch, and sometimes a small spot on each side of it), and
one a little before aud another just beyond the middle, these
two connected along the suture—and a broader, simply
arcuate, transverse fascia just before the apex, whitish or
cinereous, for the rest black ; the ventral segments at the
base or laterally, the side-pieces of the metasternum, and
the pygidium in great part above, also white. Antenne
slender, moderately long in g, shorter in ¢, joint 2 shorter
and stouter than 3, 3-11 nearly equal in length, 5-10
serrate ; last joint of maxillary palpi greatly developed, very
broadly securiform, nearly as wide as the inter-antennal
portion of the head in both sexes. Prothorax broader than
the head and elytra, deeply bisinuate at the base, rounded
at the sides. Elytra long, narrowing from the base. Py-
gidium very long, compressed, acute at tip. Ventral segment
5 holiowed down the middle posteriorly in ¢. Auterior
femora and tibiz simply pubescent in both sexes.
Length (excl. head) 64-83, to tip of pygidium 83-11};
breadth (prothorax) 24-3i mm. (¢ 2.)
Loc. Seychelles: Silhouette; Round I. Java; Borneo;
Singapore ; Philippines; Malacca; Ceylon, &c.
One 2 specimen from Silhouette (Mare aux Cvochons,
ix. 1908). This is apparently a common species in. Borneo,
12s
180 Mr. G. C. Champion on Coleoptera from
Ceylon, &c., but it cannot be identified from any of the
published descriptions. There is a long series of it in the
British Museum from many differeut localities ; and Mr.
Bryant has recently captured numerous examples in Borneo.
M. miata, F., from New Guinea &c., is an allied form, and
an unnamed insect from the Andaman Is. in the Museum
collection is, perhaps, a variety of the present species.
M. composita, Walk., has very different elytral markings.
19. Mordella disparils, sp. n.
3. Moderately elongate, rather narrow ; black, the head
(except a large transverse patch on the vertex, which is
sometimes wanting), mouth-parts, joimts 1-3 of the antenne,
the sides of the prothorax broadly, the elytra each with an
oblique stripe extending from the shoulder to near the
suture and a curved or oblique fascia just beyond extending
narrowly backwards along the suture to near the tip (the
latter sometimes nearly or quite obsolete, or represented by
yellowish pubescence), the anterior cox, femora, and tibie,
the intermediate tibiz, the extreme base of the posterior
tibiz, and the calcaria testaceous or rufo-testaceous ; varie-
gated with cinereous, flavo-cinereous, and fuscous pubescence,
the flavo-cinereous hairs mostly placed on the fasciate
portions of the surface, the vestiture of the under surface
almost wholly cinereous. Antenne moderately long, slender,
joints 2 and 3 very short, equal in length, 4-11 much longer
than broad, subserrate. Terminal joint of maxillary palpi
rather stout, subtriangular. Prothorax transverse, a little
broader than the elytra, rounded at the sides. Elytra nar-
rowing from the base. Pygidium about as long as the
posterior tarsi. Anterior femora beneath, and anterior tibie
at the base within, fusco-ciliate.
?.: Similar to g, but with the head, prothorax, and
intermediate femora infuscate, the antennz a little shorter,
the anterior femora and tibiz without longer hairs.
Var. The oblique elytral fasciz connected along the
middle of the disc.
Length (incl. pygid.) 32-41 mm. (Jd 2.)
Loc. Seychelles: Silhouette, Mahé.
Eleven examples; the six from Silhouette were taken in
the high forest above Mare aux Cochons and in the low
coconut-planted country near the coast at Pointe Etienne,
ix. 1908; the five from Mahé were collected in the forest at
the summit of Morne Pilot, over 2000 feet, in the Mare aux
the Seychelles Islands and Aldabra. 181
Cochons district at about 1500 feet, and in the forest above
Cascade Estate.
The eleven specimens vary in the development of the
oblique testaceous elytral fasciz, the posterior one being
sometimes obsolete and in one example (?) united to the
anterior one. The dissimilarly coloured sexes were obtained
in each island. M. biformis, from Central America, and the
European Mordellistena abdominalis are parallel cases of
sexual dimorphism. MM. disparilis (2) seems to be related
to M. homochroa, Fairm., from Diego Suarez, but without
comparison of types it would be unsafe to identify it with
that insect. The variety with confluent fasciz was found in
a burrow in a stick in the jungle at Silhouette.
MorDELLISTENA.
Mordellistena, Costa, Kaun. Regn. Napol., Mordellid. pp. 16, 31 (1854),
A genus of world-wide distribution and abundantly repre-
sented within the tropics. One species from the Seychelles
has been described by Kolbe and three from Madagascar by
Fairmaire. Mordella castanea, Boh., {rom Guam, and various
others from the adjacent regions referred to Mordella by the
older authors may belong here. Mr. Scott’s collections
include about two hundred specimens, belonging to seven
species. One of these insects was bred from larve found
in the wood of Colea pedunculata. Vhe appended table will
help in the identification of these closely allied forms. The
sexes have been identified in nearly every case by an exami-
nation of the genitalia of one or more examples of each
species. The antennal structure is completely ignored by
nearly all authors, presumably owing to difficulties of
manipulation :—
Tibial and tarsal formula—4 or 5, 3 or 4, 2, 2;
body uniformly coloured.
Fourth antennal joint as lorg as fifth ...... mahena, Kolbe,
Fourth antennal joint much shorter than fifth. degressa, sp. n.
Tibial and tarsal formula—4, 5, 2, 1; head in
3S 2, and prothorax also in ¢, testaceous or
rufo-testaceous; elytra cinereo-bifasciate .. partilis, sp. n.
Tibial and tarsal formula— 3, 3, 2,0; fourth an-
- tennal joint short ; body uniformly coloured,
fusco-castaneous, robust .........-...... cole, sp. D.
Tibial and tarsal formula—3, 2, 2,0; fourth an-
tennal joint short.
Body uniformly coloured, fusco-castaneous,
ferruginous, or testaceous above.
Antennal joints 5-11 elongate .......... septemcarinata, sp. n.
“182 Mr. G. C. Champion on Coleoptera from
Antennal joints 5-11 not much longer than
road 5... 2.252% ssshone eee ee dirempta, sp. 0.
Body black, griseo-pubescent; head partly
tostAccoUs IN O's) <u = sheen e s Tee argutula, sp. n.
20. Mordellistena mahena.
(Text-fig. 8, posterior leg.)
Mordellistena mahena, Kolbe, Mitteil. Zool. Mus. Berlin, v. p. 28 (1910).
Moderately elongate, cuneiform, rather narrow ; casta-
neous or fusco-castaneous, thickly clothed with greyish-
brown pubescence. Antenne filiform, very long in 6,
shorter in 9, joints 1 and 2 shorter than 3, 3 in & about
one-third, and in 2 one-half, the length of 4, 4-11 equal in
length. Apical joint of maxillary palpi stout, securiform.
Fig. 3.
Mordellistena mahena, Kolbe. Posterior leg.
Pygidium long, acuminate, as long as hind tarsus. Poste-
rior tibiz with 4 or 5, first joint of posterior tarsi with 3 or
4, and the second and third joints each with 2, oblique
ridges.
Length (incl. pygid.) 833-45 mm. (¢ ?.
Loc. Seychelles: Mahé, Long Island, Round Island.
Félicité, Marie Anne.
This species was originally taken in Mahé by Brauer;
the Seychelles Islands and Aldubra. 183
several examples were obtained by Mr. Scott in the same
island in x. and xi. 1908 near Morne Blanc, not in the highest
forests, but between 500 and 1000 feet. A few were collected
at Long avd Round Islands, small cultivated islets off Port
Victoria, Mahé, vii. 1908. Several were also found in
Félicité and Marie Anne Islands, xii. 1908, in a rather dry
type of forest near sea-level.
Kolbe’s description was made from a single example, and,
as the antennal structure is not mentioned by him, it is not
quite certain whether the name should be applied to this or
the following species. The first ridge on the posterior tibiz
and first tarsal joint is at most feebly developed and often
wanting. Amongst the series examined there are at least
two of each sex with the genital ergans extruded, so that
there can be no mistake as to their identification.
21. Mordellistena degressa, sp. u.
Extremely like M. mahena, but differing from it in having
the antenne less elongate in both sexes (in ¢ about as long
asin 2 of M. mahenu), comparatively short and subserrate
in 2, joints 8 and 4 small and equal in length in the two
sexes, 5 twice as long as 4, 5-11 moderately elongate in ¢.
Posterior tibize and tarsi as in M. mahena.
Length (incl. pygid.) 8{-43 mm. (¢ ¢.)
Loc. Seychelles: Mahé, Silhouette. ;
Nine specimens, eight of which are from Mahé. All are
from the mountain-forests. These appear at first sight to
be females of M. mahena ; but as there is no corresponding
variation in the development of the antenne in the long
series of the allied forms from the Seychelles, the examples
with a short fourth joint must be separated from the rest.
M. degressa, therefore, is based upon examples with seven,
and M. mahena with eight, elongated antennal joints.
22. Mordellistena partilis, sp. n.
g@. Rather short, narrow, convex ; black, the head, an-
tenn, mouth-parts, prothorax, anterior and intermediate
legs, and the posterior tibiee and tarsi in part, testaceous ;
the elytra variegated with cinereous and black pubescence,
the latter condensed into a very large transverse patch at
the base (not quite reaching the suture) and a common,
broad, postmedian fascia (leaving a sharply defined sub-
median and apical fascia cinereous), the vestiture of the
pygidium and under surface cinereous, that of the head and
184 Mr. G. C. Champion on Coleoptera from
prothorax flavo-cinereous. Antenne moderately long, slen-
der, joints 1 and 2 rather stout, 2 shorter than 1, 3 and 4
small, subequal in length, 5-10 longer than broad, sub-
serrate. Apical joint of maxillary palpi stout, securiform.
Prothorax transverse, rounded at the sides anteriorly, not
wider than the elytra. Elytra relatively short, subparallel
in their basal half. Pygidium long, acute, as long as poste-
rior tarsus. Posterior tibiz with 4, first joint of posterior
tarsi with 38, and the second joint with 2, short oblique
ridges, the third joint also with an indication of a single
ridge.
2. Similar to g, but with the prothorax infuscate or
black, the basal margin at most testaceous, the antenne a
little shorter.
Length (incl. pygid.) 23-3 mm. (¢ ?.)
Loc. Seychelles: Mahé, Silhouette.
One male and four females—both sexes from Mahé, a
female only from Silhouette. The Silhouette specimen is
from near Mont Pot-a-eau, about 1500 feet ; those from
Mahé were all found in the forests above Cascade Estate
at about 1000 feet. Easily distinguished from the other
Seychelles forms by the sharply defined elytral markings,
the single male with both the head and prothorax testaceous.
The prothorax is more transverse thanin M. argutula. The
anterior femora are not ciliate in ¢.
23. Mordellistena colee, sp. un.
(Text-fig. 4, posterior leg.)
Moderately elongate, cuneiform, robust; fusco-casta-
neous or castaneous, thickly clothed with greyish-brown
pubescence. Antennze with joints 3 and 4 short, equal, 5 at
least twice as long as 4, 5-11 rather broad, compressed,
elongated, and 11] longer than 10, in ¢, 5-11 shorter and
subserrate in 2. Last joint of maxillary palpi rather nar-
row, elongate-triangular. Pygidium elongate, about as long
as hind tarsus. Anterior tibie slender, sinuous within.
Posterior tibiz with 3 ridges—two very long and oblique,
and a shorter apical one—and with from 8-5 stiff erect
setee along their lower edge in both sexes. First joint of
posterior tarsi with 3, and the second joint with 2, oblique
ridges, the third unarmed.
Length (incl. pygid.) 834-53 mm. (¢ 2.)
Loc. Seychelles: Mahé, Silhouette.
Numerous examples from each island. Extremely like
M, mahena and M., degressa, but averaging larger in size, the
the Seychelles Islands and Aldabra. 185
terminal joint of the maxillary palpi narrower and more
elongate, the antennz distinctly wider, with short fourth
joint as in M. degressa, the posterior tibize strongly, sparsely
se tose along their lower edge, the third joint of the posterior
tarsi without definite ridges. Three of the Silhouette
8 pecimens are labelled as having been bred from larvee found
Fig. 4,
Mordellistena colee, Champion. Posterior leg.
in the wood of an endemic tree, the “ Bilimbi marron,”
Colea pedunculata ; another from the same locality is marked
“bred from a pupa” found in same tree. All the examples
are from various places in the mountain-forests, at altitudes
ranging from about 1000 to 2000 feet.»
24. Mordellistena septemcarinata, sp. nu.
Moderately elongate, narrow, convex, shining ; ferrugi-
nous or castaneous, the eyes black, the under surface in
part and the base of the pygidium black or piceous ; thickly
clothed with greyish-brown pubescence. Antenne long in
6, shghtly shorter in 9, slender, filiform, joints 8 and 4
short, 4 a little longer and wider than 3, subtriangular,
5—11 elongate, subequal, 5 about twice as long as 4. Apical
joimt of maxillary palpi moderately stout, subtriangular.
Prothorax broader than long, not wider than the elytra.
Klytra subparallel in their basal third, gradually narrowing
from the middle, somewhat coarsely and not very densely
186 Mr. G. C. Champion on Coleoptera from
punctate, the interspaces shining. Pygidium long, becoming
very slender beyond the middle, as long as the hind tarsus.
Anterior tibiz slender, sinuous within. Posterior tibize with
3, and the first two joints of the posterior tarsi each with 2,
oblique ridges.
Length (inel. pygid.) 83-34 mm. (¢ 2.)
Loc. Seychelles: Mahé, Silhonette, Praslin.
Mahé; forest above Cascade Estate, and in the Mare aux
Cochons district, in both cases over 1000 feet, also one
specimen recorded from the low country ; Silhouette, various
places in the mountain-forests above 1000 feet ; Praslin, one
specimen from Cotes d’Or Estate.
A long series, mostly in very bad condition. Closely
resembling M. colee, but smaller, narrower, and less cunei-
form ; the elytra subparallel and somewhat coarsely sculp-
tured ; the first joint of the posterior tarsi with two ridges
ouly ; the pygidium very slender.
25. Mordellistena dirempta, sp. n.
Narrow, convex, shining; testaceous or ferruginous, the
elytra often darker towards the sides and apex, the eyes
and under surface black or piceous; thickly pubescent.
Antenne comparatively short, joint 3 very small, 4 slightly
longer and wider, 5-10 broader, subtriangular, not much
longer than broad, subequal, 1] longer than 10. Prothorax,
elytra, and pygidium much asin MW. septemcarinata. Anterior
tibize slender, sinuous within, thickened towards the base (¢ ).
Posterior tibie with 3 (the subapical one short), and the
first two joints of posterior tarsi each with 2, oblique ridges.
Length (incl. pygid.) 23-3 mm. (¢ ¢.)
Loc. Seychelles: Silhouette, Mahé, Praslin.
Silhouette, Mare aux Cochons and forest above, over
1000 feet ; Mahé, high forest of Morne Blanc, and a speci-
men also from the low country ; Praslin, Cotes d’Or Estate.
Described from about a dozen examples, other damaged
_individuals probably belonging here. These were at first
supposed to be diminutive females of M. septemcarinata,
coming as they do from each of the islands quoted; but, as
both sexes appear to be represented in each series, this
cannot be the case. M. dirempta, therefore, may be described
as a rather short, small form of M. septemcarinata, with the
antennal jomts 5-10 subserrate and but little longer than
broad. Bs
26. Mordellistena argutula, sp. n.
¢. Narrow, moderately elongate, convex ; black, the head
(a large transverse patch of variable extent on the vertex
the Seychelles Islands and Aldabra. 187
excepted), mouth-parts, basal joints of the antenne, anterior
coxe, femora, and tibiz, and calcaria testaceous or rufo-
testaceous ; closely, uniformly fusco-cinereo-pubescent.
Head very convex ; antenne slender, moderately long, joints
1 and 2 rather stout, 2 a little shorter than 1, 3 small, short,
4 subtriangular, 5-10 longer than broad, feebly subserrate ;
Jast jot of maxillary palpi stout, securiform. ~Prothorax
moderately transverse, not wider than the elytra, rounded
at the sides anteriorly. Elytra subparalle! in their basal
half. Pygidium long, acute, about as long as the posterior
tarsi. Anterior femora beneath and anterior tibiz at base
fusco-ciliate, the latter sinuous within. Posterior tibiz with
two very long oblique ridges, in addition to the short sub-
apical one. “Posterior tarsi with joints 1 and 2 each with
two oblique ridges.
?. Similar to ¢, but with the head and the anterior legs
in great part or entirely infuscate, the antenne a little
shorter, the anterior femora and tibiz not ciliate.
Length (incl. pygid.) 24-4 mm. (¢ ¢.)
Loc. Seychelles: Mahé, Round Island, Anonyme Island,
Silhouette, Félicité.
A long series, almost all from low elevations. In Sil-
houette a large number of examples was swept from grass in
the low coconut- planted country near the coast at Pointe
Etienne, 17. ix. 1908 ; a few were also taken near Mare aux
Cochons. In Mahé specimens were collected near Morne
Blane, 500-1000 feet; on the marshy coastal plains of
Anse aux Pins and Anse Royale; and a few from over
1000 feet in the Mare aux Cochons district. Examples
were also found in two cultivated islets, Anonyme and
Round Island. Feélicité: six specimens from a rather dry
type of forest near sea-level.
A small, narrow, obscure form, with the head and anterior
legs partly rufo-testaceous in ee the vestiture uniform, the
prothorax rather Jong and not wider than the elytra, the
pygidium long and acute.
EXPLANATION OF PLATE VI.
ty. 1. Cacoplesia viriditincta, Champion, 3.
Fig. 2. Cacop esia annulipes, Champion, 3.
Fig. 3. Stictodrya longipennis, Champion, 3.
Fig. 4. Mycteromimus insularis, Champion, 2.
Fig. 5. Eurygenius fragiicornis, Champion, &.
Tig. ©. Xylophilus torticornis, Champion, ¢.
Fig. 7. Ditto. Antenna.
Tu; 8. Xylophilus seychellarum, Champion, ¢.
. Aylophilus clavicornis, Champion, @.
wi) ie 10, Mordella peregrinator, Champien, 2.
188 Dr. G. A. K. Marshall on new
XIIT.—On new Species of Indian Curculionide.—Part III.
By Guy A. K. Marsuatt, D.Sc.
Subfamily Hreunr.
Genus PELTOTRACHELUS, Nov.
Head continuous with the rostrum, the eyes comparatively
small and widely separated. Rostrum about as long as the
prothorax, its sides sloping outwards from the carine
bounding the median area, the genz more or less dilated,
the apical emargination deep and triangular ; the scrobes
apical and short, visible from above ; the lateral areas im-
pressed and with two furrows—one running just below the
dorso-lateral carina, the other passing from the lower corner
of the scrobe to the lower margin of the eye ; the buccal
aperture extremely oblique and much longer than the lower
surface of the rostrum, the mentum bearing only two setze.
Antenne with the scape but little curved, subeylindrical,
slightly thickened towards the apex, and reaching beyond
the front margin of the thorax ; the funicle variable ; the
club narrowly spindle-shaped. Prothorax transverse, the
base deeply bisinuate and broader than the apex, the ocular
lobes developed or not, but vibrissz always present. Scu-
zellum small. Elytra with the shoulders obliquely rounded
and not prominent, the dorsal outline flat or only slightly
convex, the declivity steep, the apices separately rounded,
the striz partly hidden by the dense scaling. Legs with the
front cox nearer the anterior margin of the prosternum ;
the femora moderately clavate and with a small tooth; thie
tibiz simple, the corbels of the hind pair quite open; the
claws small and free.
Type, Platytrachelus pubes, Fst.
The species included in this genus were erroneously
attributed by Faust to Platytrachelus, Schh., owing to his
having wrongly identified the genotype, P. pistacinus, Boh.
- Some years ago, through the kindness of Dr. Taschenberg,
I was able to examine the type of that species, which is in
Germay’s collection in Halle ; it proved to be identical with
Amblyrrhinus viridanus, Fst. (Stett. ent. Zeit. 1890, p. 74).
Platytrachelus differs from Peltotrachelus principally in the
structure of the rostrum, which has the median area broad
and the sides vertical; the scrobes are therefore quite
invisible from above and extend backwards for more than
half the length of the rostrum. The only other species of
Species of Indian Curculionide., 189
true Platytrachelus known to me are Amblyrrhinus psitta-
cinus, Fst., and Corigetus paviei, Auriv., both of which occur
in Indo-China.
The other described species of Pellotrachelus are Platy-
trachelus propinquus, Fst., P. ovis, Hell., Cyphicerus juvencus,
Fst. (= Myllocerus acacie, Stebb.), Acanthotrachelus albus,
Pasce., and Myllocerus isabellinus, Boh.
Peltotrachelus cognatus, sp. n.
3 ¢. Colour black or piceous, with dense grey scaling
and with the following denuded areas on the elytra: —A
transverse patch just behind the scutellum, extending to
about the fourth stria; a dentate transverse band before the
middle, which is only broken at the suture; a similar but
complete and more curved band behind the middle; these
patches often partly obscured by whitish or yellowish
powdering.
Head with the eyes lateral and almost flat ; the forehead
with a central fovea. Rostrum longer than its basal width,
slightly narrowed from the base to beyond the middle, and
dilated at the apex; the dorsal area broadly and rather
deeply impressed ; the submentum with a projecting tooth.
Antenne with the second funicular jomt much longer than
the first, the others longer than broad. Prothorawx with the
sides slightly rounded and shallowly constricted at the apex,
the postocular lobes prominent, the dorsal anterior margin
rounded; the upper surface with rather coarse confluent
punctation, and with a very shallow transverse impression
before and a small rounded one behind the middle on each
side. Elytra nearly parallel-sided (¢) or dilated behind
the middle (?), the intervals distinctly broader than the
shallow strize and with short curved irregular sete.
Length 6-7, breadth 24-3 mm.
Manpras: Yercaad, 4500 ft., Shevaroy Hills (7. Bainbrigge
Fletcher).
Very closely allied to P. pubes, Fst., but differing in its
colouring and its larger and less convex eyes; the rostrum
is longer and more deeply impressed, the prothorax is more
narrowed in front, the shoulders of the elytra are less promi-
nent, and the sides more dilated behind in the female.
Peltotrachelus rugipennis, sp. 0.
Colour black, with rather thin pale green scaling, which
is often more or less abraded.
Head with the eyes lateral, elongate, and only slightly
190 Dr. G. A. K. Marshall on new
convex, the forehead thinly pubescent and without green
sealing. Rostrum much longer than its width at the base,
almost parallel-sided in the basal half, and strongly dilated
anteriorly, the basal area broadly and rather deeply im-
pressed, the median part of the submentum elevated into a
sharp conical process with the point directed backwards.
Antenne with the scape gently curved and gradually thick-
ened; the funicle with joint 2 longer than 1, and 3 to 7
longer than broad. Prothoraz with the sides subparallel
in the basal half and narrowed in front, the dorsal anterior
edge very slightly rounded, the ocular lobes strongly pro-
duced; the upper surface with close confluent punctation
which is not very distinct through the scaling, with a faint
transverse impression before the middle and a deeper
rounded impression on each side behind. /ytra with rows
of large fovee, the intervals very narrow and irregular; in
the vicinity of the suture the spaces between the fovez are
slightly raised, so that the surface appears transversely
rugose ; the setz extremely short, dense, and suberect.
Length 53-7, breadth 23-3 mm.
Manpras: Anaimalai Hills (H. ZL. Andrewes).
Peltotrachelus illobatus, sp. n.
Biack, with dense pale green or greenish-grey scaling
throughout, the head and prothorax with a yellowish tinge.
Head with the eyes rather prominent and lateral ; fore-
head with a short central stria. Rostrum longer than br oad,
very gradually dilated from the middle to “the apex, the
dorsal carine more elevated than usual and continued on
to the forehead. Antenne with the scape distinctly curved ;
the funicle with joint 2 longer than 1, the latter longer than
3 and 4 together, 5-7 much longer than broad. Prothorar
with the sides almost straight and strongly narrowed from
base to apex, the dorsal anterior margin straight, the ocular
lobes absent, being replaced by a tuft of yellow vibrisse, the
upper surface rather rugosely punctate, especially towards
the sides. Hiytra broadest behind the middle (2), with
rather deep and coarsely punctate striz, aie however,
appear very narrow and finely punctate ‘when the scaling
is intact; the setee mostly very short and depressed, but
scattered among them a number of comparatively long erect
sete.
Length 7, breadth 3} mm.
Burma: Taung-ngu (G. Q. Corbett). Camponia (Mouhot).
Species of Indian Curculionide. TSt
Peltotrachelus smaragdus, sp. n.
Black, with dense bright green scaling, the head usually
with pinkish scales ; sometimes the insect is covered with a
more or less dense whitish coating over the green scaling.
Head with the eyes very small, prominent and lateral ;
forehead with a central fovea. Rostrum longer than broad,
only slightly dilated at the apex, the dorsal area broadly
impressed, the under surface normal and with no projection.
Anienne with the scape aimost straight; the funicle with
jvint 2 nearly twice as long as 1, 1 hardly longer than 3, and
3 to 7 much longer than broad. Prothorax with the sides
scarcely curved, only slightly narrower at the apex than
at the b: ise, the dorsal anterior margin straight, the ocular
lobes absent, being replaced by a tuft of golden-yellow
‘vibrissze ; the upper surface closely punctate and with a
shallow fovea on each side behind the middle. Klytra with
fine distinctly punctate striz and broad intervals where
the scaling is intact, the striae being a good deal broader
when the scaling is removed, but even then distinctly nar-
rower than the intervals ; the setae extremely short, dense,
and suberect.
Length 4-6, breadth 2-3 mm.
Mapras: Nilgiri Hills (Sir G. Hampson, H. L. Andrewes).
A very distinct species. Apart from the absence of the
ocular lobes, the facies is that of a typical Peltotrachelus.
Genus Merronops, nov.
_ Head separated from the rostrum by a very shallow trans-
verse impression; the eyes widely separated, comparatively
small, and almost circular. Rostrum rather broad and stout,
longer than its basal width, the buccal aperture oblique, the
apical emargination angular but rather shallow; the true
scrobe apical and very short, the space from the serobe to
the eye broadly impressed. Antenne elongate and compara-
tively slender; the scape cylindrical, abruptly clavate, and
curved only towards the apex ; the funicle withjoint 1 much
longer than 2, 3 to 7 subequal, and the club narrowly spindle-
shaped. Prothorax simple, strongly transverse, the sides
rounded, its greatest width almost or quite equal to that of
the elytra, the apex narrower than the base, the latter trun-
cate or faintly bisinuate, the ocular lobes not very prominent,
broadly rounded and with short vibrissee, the. front cox
placed in the centre of the prosternum. Seutellum small.
Elytra with the base vertically truncate, its margin being
192 Dr. G. A. K. Marshall on new
slightly raised, the shoulders feeble and obliquely rounded,
punctato-striate, the intervals smooth and even. Legs with
the femora strongly clavate and having a rather large tooth,
all the tibiz sinuate internally near the base, the corbels
of the hind pair entirely open, the tarsal claws free.
Type, M. aspersus, sp. n.
Allied to Phytoscaphus, Schh., but differing from it in the
very broad and rounded prothorax, the vertical basal margin
of the elytra, and the small and widely separated eyes.
Meionops aspersus, sp. n.
Colour piceous, with chocolate-brown scaling and pale
markings; the head fawn-coloured; the prothorax with a
broad dorsal and narrower lateral stripe of yellowish-creamy
scales ; the elytra with a similarly coloured, broad, irregular,
and broken lateral stripe, and with small pale spots on the
disk, which often coalesce along the suture.
Rostrum only slightly widened at the apex, the dorsal area
almost plane and with a fine central earina, the lateral area
with a deep narrow furrow running towards the upper edge
of the eye and a broader one beneath the scrobe. Antenne
with joints 3 to 7 of the funicle about as long as broad.
Prothorax not quite as broad as the elytra at the shoulders,
the base slightly bismuate, the upper surface with shallow
punctures and slightly granulate, the sculpture being almost
hidden by the scaling. lytra jointly sinuate at the base,
the apices jointly rounded, slightly broader behind the
middle, the striz shallow, with very large subquadrate
punctures (somewhat hidden by scaling), the intervals
almost Bee and smooth, with minute subdepressed sete.
Length 53-6, breadth 21-3 mm.
ASSAM.
Meionops glaucinus, sp. n.
Colour black, with dark greenish-grey scaling throughout.
Rostrum with the dorsal area almost plane and without a
carina. Antenne with joints 3 to 7 of the funicle evidently
longer than broad. Prothoraz as broad as the elytra at the
shoulders, the base truncate, the apical portion shallowly
constricted. ilytra truncate at the base, the sides parallel
to beyond the middle, the punctures smaller.
In other respects agrees with M. aspersus, Mshl,
Length 5, breadth 2; mm.
Wy Benga: Chota ‘Nagpur (Cardon).
Species of Indian Curculionidee. 193
Subfamily 4 wrzowomrvs.
Genus OnycHOCNEMIS, nov.
Head exserted, subconical; the eyes lateral, small, almost
eircular. Rostrum broad, fattened dorso-ventrally, longer
than the head or the front tibia, almost straight, deflected,
forming a coutinuous line with the head, and with the apical
margin entire; the scrobes narrow and deep, beginning at
about one-third from the apex and continued obliquely to
beneath the base of the rostrum; the mentum small, sub-
quadrate, about as long as its supporting peduncle, convex,
impunctate, and very shining. <Anienne short, geniculate ;
the scape almost straight, clavate, reaching the middle of
the eye ; the funicle 7-jointed, joint 1 swollen and longer
than any of the others, 2 subconical and as long as broad,
the remainder strong! y transverse, very closely packed, and
rapidly widening outwardly, joint 7 being closely annexed
to the club, which is broadly ovate and 3-jointed. Pro-
thorax without postocular lobes, and with the base bisinuate.
Scutellum distinct, circular. Hlytra oblong, broader than
the prothorax, entirely covering the pygidium, with distinct
shoulders and ten strie. Legs short and stout; the hind
cox ovate, not reaching the edge of elytra; the femora
moderately clavate and not toothed, the hind pair not nearly
reaching the apex of the elytra ; the tibize almost straight,
slightly compressed, strongly uncinate at the apex, and also
with a short sharp mucro projecting perpendicularly from
the inner angle; the tarsi broad, joint 2 transverse, 3
broadly lobate, 4 short, the lower suriace clothed with fine
pale pubescence, which is sparse on the two basal joints,
the claws very small and connate at the base. Sternum:
the prosternum very short, with the front margin shallowly
sinuate, the coxze in the middle and narrowly separated ;
the mesosternum with the epimera not ascending and broadly
separating the episterna from the elytra, the intercoxal pro-
cess broadly truncate at the apex; the metasternum between
the coxee about as long as the middle cox, the episterna
comparatively broad. Venter with the intercoxal process
broadly rounded, the two basal segments fused together and
delimited only by an almost straight shallow stria, the inter-
mediate segments not. angulated externally, 2 almost as lorg
as 3 +4 in the- middle, and 5 but little longer than 4.
Type, Onychocnemis carey@, sp. 0.
Allied to the European Bradybatus, Germ., and the South-
African Thamnobius, Sclih. The former genus differs in its
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 13
194 Dr. G. A. K. Marshall on new
much longer, more slender, and cylindrical rostrum, its
simply uncinate tibiz, free and bifid tarsal claws, and dentate
front femora. In Thamnobius the rostrum is also more
cylindrical, the scape does not exceed the front margin of
the eye, the tibiz are merely uncinate (not mucronate, as
stated by Lacordaire), and the tarsal claws are free and
appendiculate.
Onychocnemis carey@, sp. Nn.
o 9. Colour red-brown, shining and sparsely clothed
with short recumbent white sete; the head darker; the
elytra with the entire suture blackish brown, as well as a
large common patch extending from the base to beyond the
middle and laterally as far as the fifth stria, its outline
being very similar to that of the elytra ; the mesosternum,
metasternum, tarsal claws, and the two apical hooks of the
tibia also dark brown or blackish.
Head rugosely punctate, the forehead a little narrower
than the base of the rostrum and broader than the eye.
tostrum very gradually widened from base to apex, rugosely
punctate above from the base to the end of the scrobe, the
apical area more lightly punctate (g) or impunctate ( ).
Prothorax subconical, almost as long as its width at the
base, gradually narrowed from there to the apex, the sides
gently rounded, without any anterior constriction, the basal
angles nearly right angles, the base angularly produced in
the middle, the apical margin very shallowly sinuate dorsally
and oblique at the sides ; the upper surface with coarse
reticulate punctation throughout and with a median stripe
of deuser pale recumbent sete. /ytra almost parallel-sided
from the shoulders to well behind the middle, broadly
rounded behind, the apices continuous, the basal margin
slightly raised and almost straight from the second stria to
the shoulders ; the strize broad, containing deep closely-set
punctures, which diminish behind, the intervals scarcely
broader than the striz, almost flat and finely aciculate, the
posterior callus nearly obsolete; the dorsal outline flat
from the base to the middle, then gradually declivous. Legs
coarsely punctate and clothed with curved white sete.
Length 2-2}, breadth 1-1} mm.
Mysore: Madhavgiri (H. H. Mann, Pusa Coll.).
This species was found on the leaves of the jak-fruit tree
(Careya arborea).
Species of Indian Curculionide. 195
Subfamily Omoruorry.
Genus TELUROPUS, NOV.
Head globose, with the eyes lateral. Rostrum stout, about
as long as the front tibia, somewhat depressed, the apical
margin very shallowly sinuate ; the scrobes invisible trom
above, beginning at some distance from the apex, curving
rapidly downwards behind the antennee, and ending in a flat-
tened punctate area almost on the lower surface of the base
of the rostrum; mandibles stout, tridentate; mentum small
and square, about equal in length to the peduncle of the
submentum, and not broader than the lateral space on each
side of it. Prothorax with the basal margin deeply bi-
sinuate ; the anterior margin oblique at the sides and without
any postocular lobes. Scuéellum distinct, almost circular.
Elytra short and broad, with ten complete strize ; the in-
flexed lateral margin unusually narrow, involving only the
tenth stria, and without true epipleure. Wings fully deve-
loped. Legs short and stout; the femora only slightly
clavate and each with a small tooth, the hind pair scarcely
reaching the apex of the elytra; the tibize with the external
apical angle strongly uncinate, and the inner angle with a
sharp mucro as well; the tarsi short and broad, the second
joint twice as broad as long, the fourth projecting only a
short distance beyond the third, the claws simple and stout.
Sternum with the front coxe very widely separated and placed
behind the middle, the space between them quite flat; the
median coxe still further apart, the side-pieces of the meso-
sternum fused together but divided by a stria, the suture
between the mesosternum and episternum entirely oblite-
rated, and the intercoxal piece broadly truncate; the length
of the metasternum between the cox not greater than that
of the median coxe, the episternaas broad as the base of the
mid-femora and fused with the metasternum, but the line of
junction quite distinct, the epimera imperceptible ; the hind
coxve as widely separated as the middle pair. Venter short,
with segment 2 nearly as long as 3+4 and separated, from
1 by a deep straight incision, the intercoxal process very
short and broad, with an angular projection in the middle,
and segments 2 and 3 angulate externally.
Type, Teluropus subcostaius, sp. n.
In general form the only known species bears considerable
resemblance to the African genus Omophorus, Schh.*, and
* In most collections this genus stands under the later name Meta-
tyges, Pasc. M. turritus, Pasc., is a synonym of O. stomachosus, Boh.,
while M. parvus, Fst., is identical with O. zndispositus, Boh., the type of
which is now in the Oxford University Museum (Sommer’s collection).
13*
196 Dr. G. A. K. Marshall on new
the Fijian Physarchus, Pasc., except that the shoulders of
the elytra are much less prominent. But both these genera
differ, inter alia, in the absence of the inner apical mucro
on the tibiz, and in having the front coxz contiguous and
the hind pair much closer together than the middle pair.
Teluropus baliardi, sp.
g. Dark red-browa, fairly closely clothed with short,
curved, golden-brown sete.
Head rugosely punctured throughout, the forehead almost
as broad as the base of the rostrum, transversely flattened
and with a central fovea; the eyes almost circular, their
greatest depth at about one-fourth from the hind margin,
Rostrum stout, parallel-sided, and porrect from the base to a
little beyond the middle, thence slightly widened and curved
downwards, somewhat flattened longitudinally at the sides in
the basal half, and rugosely punctured throughout right up
to the apex. Antenne short; the scape stout, slightly com-
pressed, strongly clavate, and coarsely punctate ; the funicle
with joint 1 rather longer than 2, and joints 2 to 7 of about
equal width and widening regularly outwards, 7 being closely
fitted to the club, which has three distinet joints, Prothorax
broader than long, broadest near the base and rapidly
narrowing in front, with a broad apical constriction, the
auterlor margin straight, the base with a large median lobe,
which is emarginate at its apex; the upper surface convex,
coarsely and confluently punctate throughout, and with two
low broad elevations in the middle of the disk ; these merge
and slope gradually away behind, but in front they are
abruptly narrowed where they cross the apical constriction
and enclose a large rounded depression between them ;
below these prominences on each side is another much lower
rounded elevation. Hlytra together nearly as broad as long,
parallel-sided from the shoulders to beyond the middle, and
very broadly rounded behind ; each elytron strungly lobate
at the base, the greatest depth of the lobe being at the third
interval, which bears a slight basal callus; the juxta-basal
area slopes steeply forwards from a transverse postbasal
ridge, which bears a rounded prominence on intervals 3 and
5; a little behind this is a short costate elevation on in-
terval 3 and a less distinct one on 5 ; the strie are deep and
strongly punctate and the intervals are rugose, the alternate
ones being slightly more convex. Legs rugosely punctate,
the femora with scattered granules, set with curved suberect
Species of Indian Curculionidae. 197
sete ; the anterior pairs of tibia rather sharply angulate on
the lower surface not far from the base.
Length 45-5, breadth 23-3 mm.
Mapras: Coimbatore (. Ballard, type). Mysore:
Madhavgiri, on leaves of jak-fruit, Careya arborea (H. H.
Mann, Pusa Coll.).
Subfamily /sorruywcurvz.
Phenomerus angulicollis, sp. n.
?. Colour black, sparsely clothed with rather stout,
transversely recumbent, pale yellowish, hair-like scales,
having the following patches apparently bare, but really
clothed with similar black hairs which are not very con-
spicuous :—A large transverse patch on the anterior half of
the prothorax, and two irregular patches on each elytron,
one before and the other behind the middle.
Head with scattered punctures, the eyes a little more
widely separated than in P, sundewalli. Rostrum red-brown,
the thickened basal portion forming about one-fourth of the
whole, and not sulcate, but with two posteriorly convergent
rows of fine punctures on the disk; the scrobes continued
to beyond the middle as a shallow punctate furrow, and with
a fine stria just above them. Prothoraz at least two and a
half times as long as its basal width, the sides obtusely
angulated in front of the middle, the dorsal margin bounded
by a fine carina in the basal half, the upper surface with
reticulate punctures which are longitudinally subcoalescent,
leaving a distinct smooth central carina. Llytra narrowly
cylindrical, about as broad as the prothorax at its angu-
lation, with shallow striz containing closely set transverse
punctures, the intervals narrowly carinate, except the four
outer ones, which are broader and almost flat. Legs similar
to those of P. sundewalli, except that the large tooth on the
elongate hind femora is more deeply sinuate at the base of
its posterior edge, so that this edge is distinctly angulated
in the middle.
Length 34, breadth ? mm.
BrneaL: Sanderbans, 18. 11. 1915 (C. F. C. Beeson).
Very similar superficially to P. sundewalli, Boh., but
distinguished by its obviously narrower build, more widely
separated eyes, the lateral angulation and basal lateral
carina of the prothorax, and the shape of the tooth on the
hind femora.
Found in burrows in sundri-trees (Heritiera littoralis),
198. MCT, Regan on the Clupeid Fishes
Phaenomerus brevirostris, sp. n.
3 @. Colouring similar to that of P. angulicollis, but the
pale scales more generally distributed, so that the dark patch
on the prothorax is indistinct and those on the elytra are
very much redu-ed,
Head with close shallow punctures, the forehead broad,
about twice the breadth of the funicle. Rostrum unusually
short and stout, the thickened basal portion forming half
(2) or more than half (¢) its length, and bearmg two or
four shallow furrows, the apical area smooth and sparsely
punctate ; the serobe continued to well beyond the middle
in both sexes, but without any distinct furrow above it.
Antenne short, all the joimts of the funicle except the first .
very strongly transverse, the club shorter and more obtuse
than in P. sundewalli. Prothorax about twice as long as
broad, parallel-sided from the base to beyond the middle,
thence narrowing gradtially to the apex, the upper surface
simply reticulate, the punctures not coalescing longitu-
dinally, with an indistinet central costa. E/lytra cylindrical,
very slightly broader than the prothorax, with coarsely
punctate striz, the dorsal intervals narrow, subcarinate, and
crenulate. Legs as in P. sundewalli, except that the hind
femora are much shorter, extending only a short distance
bevond the apex of the elytra, the basal stem is more rapidly
widened, and the large tooth is more deeply sinuate at the
base of its posterior edge, so that the edge is distinct'y angu-
lated in the middle.
Length 3, breadth ? mm.
Unitep Provinces: Khairabenda, Khash Forest, 29. xi.
1913 (C. F. C. Beeson).
This species can be readily distinguished from both
P. sundewalli and P. angulicollis by its short rostrum and
hind femora, and its broad forehead. Found in burrows in
dead sal-tree (Shorea robusta). Mr. Beeson informs me that
all the three species of Phenomerus mentioued here occurred
in burrows of Scolytide &c., and he is of opinion that they
are predaceous upon those beetles.
X1LV.—A Leevision of the Clupeid Fishes of the Genus Pello-
nula and of Related Genera in the Ltivers of Africa. By
C. Tate Reaan, M.A.
(Published by permission of the Trustees of the British Museum.)
Perrroxrra and its allies are distinguished from Clupea,
Sardinella, &c., by the stronger dentition and by the absence
of the anterior supramaxillary bone.
of the Genus Pellonula dc. 199
Synopsis of the Genera.
I. Abdominal scutes sharply keeled ; preemaxillary teeth rather strong:
A. Lower jaw not or but little’ projecting ; anterior mandibulary
teeth enlarged, but no strong canines in either jaw.
1. D., 16-18, “above or just behind pelvics. A. 16-21. Scales
about 45/11- 15. Vertebree 42-43. Supramaxillary large.
1, Pellonula.
2. D. 12-14, above interspace between pelvics and anal.
A. 20-21. Scales 33/8. Vertebre 48 .......... 2. Pecilothrissa.
A. 21-25. Scales 88-42/10-12. Vertebrae 89-40... 3. Microthrissu.
3. D. 13-14, above pelvics. A. 17-18, far behind dorsal. Scales
40-44/10. Vertebree 42. Supramaxillary small.
4. Potamothrissa.
B. Lower jaw strongly projecting.
Premaxillaries with an inner series of 2 or 3 strong
canine-like teeth on each side ; anterior teeth
OF lower jaw Onlatred. cc... cee cee es wo. 58. Cynothrissa.
Premaxillary teeth uniserial, with a canine on each
side; lower jaw with a pair of strong anterior
sa a thats. aeah hat atte bt 6. Odaxothrissa.
Il. Abdominal seutes in front of pelvic fins feebly keeled; pra-
maxillary teeth small.
Maxillary narrow proximally and expanded distally ;
. tongue and palate toothless ............+... 7. Stolothrissa,
Maxillary broad throughout its length; a patch of
teeth on each palatine and a strip on tongue .. 8. Limnothrissa.
1. PELLONULA, Giinth. 1868.
Cat. Fish. vii. p. 452. .
Form elongate, compressed; abdomen sharp-edged. Mouth
moderate, terminal, with the lower jaw a little projecting ;
upper jaw without median notch; maxillary of a narrow
proximal and an expanded distal part ; in front of the narrow
part a ligament runs from the end of the premaxillary to the
broad part of the maxillary ; a single well+developed supra-
maxillary. Teeth in jaws uniserial, conical, acute; pree-
maxillary teeth rather strong, unequal, without well-marked
canines; mandibulary teeth enlarged anteriorly ; maxillary
teeth minute ; ; a patch of conical teeth on each palatine ; an
elongate patch of small teeth on tongue. 6 brancliostegals.
Scales with entire edges, moderate, in a longitudinal series
one to each myotome; ventral scutes sharply keeled and
acutely pointed, commencing on or behind the thoracic keel
formed by the hypocoracoids. Dorsal of 16-18 rays, anal of
16-21. Pelvics 8-rayed, below or a little in advance of
dorsal. Vertebree 42 or 43. A bluish-silvery lateral band.
200 Mr. C. T. Regan on the Clupetd Fishes
Pellonula modesta, Fischer (Jahrb. Hamb. Wiss. Anst. il.
1885, p. 75), from. Eloby, is placed by Boulenger in the
synonymy of P. vorax, but unless the description is quite
incorrect (teeth in jaws minute, tongue and palate toothless,
no lateral band, &c.) it is not a Pellonula at all, and is most
likely a Sardinella.
A species from the coast of Brazil, described by Stein-
dachner as Pellonula bahiensis(Sitzungsb, Akad. Wien, Ixxx.
1, 1880, p. 181, pl. i. fig. 2), of which Sardinel/a pernam-
bucana, Schreiner & Ribeiro (Arch. Mus. Rio Janeiro, xii.
1903, p. 72), appears to be a synonym, evidently belongs to
the genus L/eringia, Fowler, 1911 (Rhinosardinia, Bigen-
mann, 1912), the type of which is H. amazonted, Steind.,
from the Amazon and Guiana.
Synopsis of the Species.
26-30 gill-rakers on lower part of anterior arch. Depth
3 to 4 in the length; caudal peduncle as long as
HEED OGM. wears cen hat bene e es OREM RES Ns ie Meets l. vorax.
27 gill-rakers on lower part of anterior arch. Depth 5
in the length; caudal sats a little longer than
MRED) caret si ack a AIG hsteteiets eR Iee wai pie ima eee 2. leonensis.
33 gill-rakers on lower part of anterior ‘arch. Depth 4 :
in the length ; caudal peduncle 13 as long as deep . 3. stanleyana.
1. Pellonula voraz.
Pellonula vorax, Giinth. Cat. Fish, vii. p. 452 etal
Pellonula vorax (part.), Bouleng. Cat. Afr. Fish. i. p. 156, fig. 124
(1909),
Depth of body 3 to 4in the length, length of head 82 to 44,
Snout neaily equal to diameter of eye, “which is 3 to 32 in
Jength of head; maxillary extending to below aiterine
margin or antenion part of eye ; lower jaw a little projecting,
26 to 30 gill-rakers on lower part of anterior arch, About
45 scales in a longitudinal series, 14 in a transverse series ;
ventral seutes 13- 1548+ 10. Dorsal 16-18 ; origin equi-
distant fiom end of snout and base of caudal, or nearer snout.
Anal 18-21. Pelvics varying somewhat in position, rarely
entirely below the dorsal, sometimes entirely in advance of
it. Caudal peduncle as long as deep. Vertebree 42.
West Africa, from the Senegal to Angola.
Numerous examples, measuring up to 140 mm, in total
length.
This species may occur in the Lower Congo, but none of
the specimens from the Congo enumerated by Boulenger
belongs to it; I refer specimens 18 and 19 to Cynothrissa
of the Genus Pellonula ce. 201
ansorgi?, 20-23 and 24-25 to Mierothrissa parra, 26 to
Potamothrissa acutirostris, 27 and 28 to Peeilothrissa con-
gica, and 29 and 30 to Pellonula stanleyana.
2. Pellonula leonensis.
Pellonula leonensis, Bouleng. Cat. Afr, Fish. iv. p. 172, fig. 111 (1916).
Depti of body 5 to 5) in the length, length of head 32 to 4.
Snout nearly as long as diameter of, eye, which is 3 in length
of head; maxillary extending to below anterior 4 of eye ;
lower jaw a little projecting. 27* gill-rakers on lower part
of anterior arch. 45 scales in a longitudinal series, 11 or 12
in a transverse series ; ventral scutes 13-14+8-9. Dorsal
16; origin equidistant from end of snout and base of caudal.
Anal 16-17. Pelvics below origin or anterior rays of dorsal.
Caudal pedunele a little longer than deep,
Sierra Leone.
Two specimens, 57 mm. long, from the North Sherbo
District.
3. Pellonula stanleyana, sp. n.
+ TLellonula vorax (part.), Bouleng. Cat. Afr. Fish. i. p. 156 (1909).
Depth of body 4 in the length, length of head 43. Snout
as long as diameter of eye, which is 3} in Jength of head ;
maxillary extending to below anterior margin of eye ; lower
jaw a little projecting. 33 gill-rakers on lower part of ante-
rior arch. 45 scales in a longitudinal series, 14 in a trans-
verse series ; ventral scutes 14+9. Dorsal 16-17; origin
nearer to end of snout than to base of caudal, Anal 18.
Pelvics below anterior rays of dorsal. Caudal peduncle 1$
as long as deep. Vertebre 43,
Stanley Falls.
Two specimens, 110 mm. in total length ; one of these is
a skeleton, but I have been able to count the gill-rakers.
2. P@CILOTHRISSA, gen. nov.
Closely related to Pellonula, differing in that the scales in
a longitudinal series are less numerous than the myotomes,
the supramaxillary bone is rather small, and the tongue is
toothless. Dorsal fin of 13 rays, above the interspace be-
tween pelvics and anal, the last with 20-21 rays. Scales
33/8. Vertebree 43.
* | find 27 gill-rakers on the lower part of the anterior arch in each
of the type-specimens, which I have examined under a binocular
microscope,
202 Mr. C. T. Regan on the Clupeid Fishes
Peecilothriss1 congica, sp. n.
Pellonula vorax (part.), Bouleng. Cat. Afr, Fish. i. p. 156 (1909).
Depth of body 4 in the length, length of head 4}. Snout
a little shorter than diameter of eye, which is 2? in length of
head ; jaws equal anteriorly ; maxillary extending to vertical
from anterior edge of eye; maxillary teeth quite distinct ;
19 gill-rakers on lower part of anterior arch. 33 scales ina
longitudinal series, 8 in @ transverse series ; ventral scutes
11-12+9-10. Dorsal 13; origin equidistant from end of
snout and base of caudal. Anal 20-21. Pelvics well in
advance of dorsal. Caudal peduncle longer than deep.
Lateral band narrow, present only on posterior half of fish.
Vertebree 43.
Congo.
T'wo specimens, 55 and 60 mm. in total length, from
Coquilhatville and from Monusembe.
3. Microturissa, Bouleng. 1902.
Ann. Mus. Congo, Zool. 11. p. 26.
Scarcely generically distinct from Pellonula, but tongue
toothless, dorsal fin of 12-14 rays and anal of 21 to 25, and
pelvics in advance of dorsal. Scales 38-42/10-12. Ver-
tebree 89-40. ‘
1. Microthrissa parva, sp. n.
Pellonula vorax (part.), Bouleng. Cat. Afr. Fish. i. p. 156 (1909).
Depth of body 4 to 5 in the length, length of head about 4.
Snout a little shorter than diameter of eye, which is rather
more than 4 the length of head; maxillary extending to
vertical from anterior edge of eye; lower jaw a little pro-
jecting. 26 or 27 gill-rakers on lower part of anterior arch.
AO to 42 scales in a Jongitudinal series, 10 or 11 im a trans-
verse series; ventral scutes 12-13+7-8. Dorsal 12-14;
origin’ nearly equidistant from end of snout and base of
caudal. Anal 21-23, not extending forward to below dorsal,
Pelvies in advance of dorsal. Caudal peduncle longer than
deep. 40 vertebree.
Upper Congo.
Two specimens of 40 mm, from Coquilhatville and three
of 30 mm. from the Tumba Lake.
of the Genus Pellonula §e. 203
2. Microthrissa royauxi.
Microthrissa royauri, Bouleng, Ann, Mus, Congo, ii, 1902, p. 26, and
Cat. Afr. Fish. i. p. 161, tig. 129 (1909).
Depth of body 8-34 in the length, length of head 4.
Snout shorter than diameter of eye, which is 3 in length of
head ; jaws equal anteriorly; maxillary with minute teeth,
extending to below anterior margin of eye; 14 gill-rakers on
lower part of anterior arch. 38 to 40 scales in a longitudinal
series, 12 in a transverse series ; ventral scutes 12-13 + 6-7.
Dorsal 13; origin equidistant from end of snout and
base of caudal, behind the 8-rayed pelvics. Anal 23 (-25),
extending forward nearly to below end of dorsal. Caudal
pedunele a little deeper than long. 39 vertebree,
Ubanghi R.
One of the types, 55 mm. in total length, from Banzyville.
4, POTAMOTHRISSA, gen. nov.
Closely related to Pellonula, but maxillary narrow, supia-
maxillary bone quite small, and no teeth on tongue or on
maxillary. Dorsal fin of 13 or 14 rays, placed well forward,
its origin much nearer to end of snout than to base of caudal,
above or in advance of first ray of prlvies; anal of 17 or 18
rays, far b. hind dorsal. Scales 40-44/10. Vertebree 42.
1. Potamothrissa obtusirostris, (Fig. 1, 2.)
Pellonula obtusirostris, Bouleng. Cat. Afr. Fish. i, p. 158, fig. 126
(1909).
Jaws equal anteriorly. 16 gill-rakers on lower part of
anterior arch. Ventral scutes 9-10+ 9-10.
Aruwimi River, Congo.
Two specimens, 72 mm. in total length.
2. Potamothrissa acutirostris.
Pellonula acutirostris, Bouleng. Cat. Afr. Fish. i. p. 159, fig. 127 (1909).
Lower jaw shorter than upper. 19 gill-rakers on lower
part of anterior arch. Ventral scutes 12-134 10-12.
Upper Congo.
Five specimens, up to 75 mm. in total length.
5. CYNOTHRISSA, gen. nov.
Differs from Pellonula in the very prominent lower jaw
and in the presence of an inner series of pramaxillary teeth,
204 Mr. C. T. Regan on the Clupeid Fishes
comprising 2 or 3 strongly enlarged teeth on each side.
Vertebree 42.
Heads of 1. Cynothrissa mento, 2. Potamothrissa obtusirostris (X 23).
1. Cynothrissa mento, sp.n. (Fig. 1, 7.)
Pellonula vorax (part.), Bouleng. Cat. Afr. Fish. i. p. 156.
Depth of body 4 in the length, length of head (without
lower jaw) 32. Snout longer than diameter of eye, which is
33 in length of head; maxillary extending to below anterior 4
of eye. 19 gill-rakers on lower part of anterior arch. About
45 scales in a longitudinal series, 14 in a transverse series ;
ventral scutes 15+9. Dorsal 17; origin above base of
pelvics, equidistant from end of snout and base of caudal.
Anal 21. Caudal peduncle longer than deep.
Nigeria.
A single specimen, 130 mm. in total length, from Agberi,
Southern Nigeria.
This new species is distinguished from C. ansorgit by the
more slendcr form, fewer gill-rakers, and more numerous
anal rays.
2. Cynothrissa ansorgit.
Pellonula vorax (part.), Bouleng. Cat. Afy. Fish. i. p. 156 (1909).
Odaxothrissa ansurgii, Boulenger, op. cit. iv. p. 172, tig. 112 (1916).
Depth of body 3 to 8} in the length, length of head
(without lower jaw) 32 to 4. Snout as long as or longer
than diameter of eye, which is 3 to 4 in length of head;
maxillary extending to below anterior 4 or middle of eye.
23 to 25 gill-iakers on lower part of anterior arch. 42 to 45
scales in a longitudinal series, 14 or 15 in a transverse
series; ventral scutes 13-16+9-11. Dorsal 15-17; origin
‘of the Genus Pellonula &c. 205
above or immediately behind base of pelvics, nearly equi-
distant from base of caudal and end of snout. Anal 17-19.
Caudal peduncle as long as deep. Vertebre 42.
Lower Congo and Angola.
Nine specimens, 110 to 160 mm. long, including the types
from Angola and two from Boma and from Vivi, Lower
Congo.
6. Opaxorurissa, Bouleng. 1899.
Differs from Pellonula in the very prominent lower jaw,
with a pair of strong canines anteriorly ; preemaxillary teeth
uniserial, with a canine on each side.
1. Odaxothrissa vittata, sp. n.
Odaxothrissa losera (part.), Bouleng. Cat. Afr. Fish, i. p. 160 (1909).
Depth of body 43 in the length, length of head 82.
Snout longer than Gtateeer of eye which is 4 in length of
head ; maxillary extending to hala middle of eye ; canines
very strong. Gill-rakers ‘shorter than gill-filaments, 22 or 23
on lower aaa of anterior arch. 45 scales in a longitudinal
series, 14 in a transverse series; ventral scutes 1249.
Dorsal 16; origin a little nearer base of caudal than end
of snout, immediately behind base of pelvics. Anal 21.
Caudal peduncle 14 as long as deep. A well-detined silvery
lateral band.
Ubanghi River.
A single specimen, 110 mm. long, from Banzyville.
2. Odaxothrissa losera.
Odaxothrissa losera, Bouleng. Ann. Mus, Congo, Zool. i. 1899, p. 64,
pl. xxxi. fig. 1
Depth of body equal to or a little less than length of head,
which is 84 in the length of fish. Snout a little longer than
diameter of eye, which is 32 to 43 in length of head; max-
illary extending to below ‘aiberbn 2 or middle of eye. 27
gill-rakers on lower part of anterior arch, 44 to 46 scales in
a longitudinal, 14 in a transverse series ; ventral scutes
11 +9. Dorsal 16-17 ; origin equidistant from end of snout
and base of caudal, immediately behind base of pelvics.
Anal 21-22. Caudal peduncle as long as deep. Lateralh
band vestigial.
Upper Congo,
206 Clupetd Fishes of the Genus Pellonula dc.
The above description is based on one of the types, a speci-
men of 85 mm. from Coquilhatville, and on the figure of the
larger type-specimen, 160 mm. long. In the smaller fish the
gill-rakers are rather longer than the gill-filaments, but in the
larger they are said to be much shorter and to number only
18 on the lower part of the anterior arch ; this may be a
m sprint for 28, or possibly in the adult the anterior gill-
rakers may be vestigial.
7. STOLOTHRISSA, gen. nov.
Mouwh formed as in Pellonula, but teeth in jaws quite
small and no teeth on palate or tongue. Abdomen in front
of pelvic fins rounded, with the scutes but weakly keeled ;
scutes behind pelvic fins strongly keeled and acutely pointed.
Vertebra: 44.
Stolothrissa tanganice, sp.n. (Fig. 2, 2.)
Pellonula miodon (part.), Bouleng. Cat. Afr. Fish, i. p. 157 (1909).
Depth of body about 6 in the length, length of head 3} to
4, Snout rather longer than diameter of eye, which is 34
in length of head; maxillary not or barely reaching vertical
from anterior margin of eye; lower jaw slightly projecting.
4) to 42 gill-rakers on lower part of anterior arch. About
45 scales in a longitudinal and 13 in a transverse series ;
ventral scutes 13-14+8-9. Dorsal 15; origin nearly equi-
distant from end of snout and base of caudal, Anal 17.
Pelvics below or a little in advance of middle of dorsal.
Lateral band broad.
Tanganyika,
Four specimens, 80 to 95 mm. in total length, from Vua
and Ndanvie.
Heads of 1. Limnothrissa miodon, 2. Stolothrissa tanganice (X 2).
On Tabanide from Australia &c. 207
8. LIMNOTHRISSA, gen. nov.
Maxillary broad throughout its length, its dentigerous
margin extending right up to the preemaxillary ; latter with
quite small teeth, but dentition of lower jaw, palatines, and
tongue as in Pellonula. Abdomen in front of pelvic fins
rounded, with scutes but weakly keeled; sentes behind
pelvic fins strongly keeled and acutely pointed. Vertebree 44.
Limnothrissa miodon. (Fig. 2, 7.)
Pellonula miodon (part.), Bouleng. Cat. Afr. Fish. i. p. 157, fig. 126
(1909).
Depth of body 43 in the length, length of head 32 to 4.
Snout as long as diameter of eye, which is 34 in length of
head ; maxillary extending to below anterior } of eye ; lower
jaw slightly projecting. 31 to 33 gill-rakers on lower part
of anterior arch. About 45 scales in a longitudinal and
14 in a transverse series; ventral scutes 13-14+ 10-11.
Dorsal 15 ; origin equidistant from end of snout and base
of caudal. Anal 16-18. Pelvies below middle or anterior
part of dorsal. Lateral band broad.
‘l'anganyika.
Four specimens, 100 to 140 mm. in total Jength, from
Ndanvie, Tembwi, and Kuasakalewa; also some young
examples not included in tle description.
XV.—New Species of Tabanidee from Australia and the
Fiji Islands. By Gerrrupe Ricarpo.
Tue identification of species and descriptions of new species
contained in this paper are from specimens forwarded to
Mr. Marshall by Dr. E. W. Ferguson and Dr. J. Burton
Cleland.
One new species from the Fiji Islands, the type being in
the British Museum Coll., is included. The types of all the
new species will be presented to the British Museum by
the Imperial Institute of Keonomic Entomology, with the
exception of a few species belonging to the South Australian
Museum and the National Museum, Victoria; in these
cases, paratypes are kept tor the British Museum. ;
208 Miss G. Ricardo on Tabanidee from
PANGONINZ.
Diatomineura ruficornis, Macquart, Dipt. Exot., Supp!. i.
p. 125. (1846) ; Walker, List (Dipti pt. w.,* Supyl,
1, p. 142 (1854); Ricardo, Ann. & Mag. Nat. Hist.
(7) v. p. 118 (1900).
Females and males from the summit of Mt. Wellington,
Tasmania.
I believe these specimens are Macquart’s species, the type
of which was examined by me at Lille in 1906, and the
following note made of it:—“ Palpi with the first joint
short, the second long, flattened at base, broad, ending in
a fine point. Subcallus fulvous, forehead darker. Thorax
blackish, with yellowish tomentum ; this describes it better
than Macquart’s remarks, the stripes he speaks of consist
of pubescence, sides with yellow pubescence ; the other
details of his description are correct.”
This species is very probably identical with Diatomineura
constans, Walker, as suggested by Mr. White.
Diatomineura constans, Walker, Dipt. Saund, 1. p. 15 (1850);
Ricardo, Ann. & Mag. Nat. Hist. (7) v. pp. 1138, 118
(1900) ; White, Royal Soc. Tasmania, 1915, pt. ii,
Diptera Brachycera of Tasmania, p. 20.
Nine females from Waratah, Tasmania (Lea).
One female from Hobart (Lea).
Diatomineura auriflua, Donovan, Gen. Illustr. Ent. Hym. et
Dipt. (1805) [Tabanus}; Wied. Ausszweifl. Ins. 1,
p. 194 (1848) (Pangonia) ; Ricardo, Ann. & Mag. Nat.
Hist. (7) v. pp. 112, 119 (1900) ; id. (8) xvi. p. 27
(1915) ; White, Royal Soc. Tasmania (1915), pt. i,
Diptera Brachycera ‘Tasmania, p. 19.
[ Pangonia solida, W1k. |
[ Pangonia dives, Macq. |
One male and one female from Mt. Washington, Tasmania
(Lea).
Two males and one female from Devonport, Tasmania
(Lea).
Diatomineura brevirostris, Macq. Dipt. Exot., Suppl. iv.
p. 826 (1842); Ricardo, Ann. & Mag. Nat. Hist. (7) v.
p- 118 (1900) ; id. (8) xvi. p. 29 (1915).
Two females from Dorrigo, New South Wales (W. Heron).
Australia and the Fiji Islands. 209
Six females from Clarence River, New South Wales
(A. & FL R. Zietz).
Diatomineura testacea, Macquart, Dipt. Exot. i. p. 103
(1838) ; Walker, List Dipt. pt. v., Suppl. i. p. 145
(1854) ; Ricardo, Aun. & Mag. Nat. Hist. (8) xvi. p. 31
(1915).
One female from Cairns District, Queensland (Dodd).
Diatomineura abdominalis, 2 , sp. n.
Type (female) and another from Laurieton, New South
Wales, 19.10.15.
A species nearly allied to Diatomineura brevirostris, but
distinguished from it by the absence of any white or yellow
haired spots on abdomen and by the first posterior cell
being considerably narrowed at the border, only half open.
Length 15 mm,
Face, palpi, and antenne as in D. brevirostris. Forehead
parallel, also the same; the frontal callus when not denuded
appears to be small, pear-shaped. Thorax dresden-brown .
with short black pubescence, no stripes are visible; shoulders
with chiefly yellowish-white hairs continued on sides of
thorax as far as the scutellum, but they do not quite reach
the apex of scutellum; a faint white spot is visible on each
side of thorax near the suture, on the dorsum. <Addomen
amber-brown, mottled with blackish markings, smooth,
shining, with very short black pubescence; sides with
yellowish-white hairs ; apex and sides of posterior segments
paler in colour; underside paler with black spots. Legs
reddish-yellow ; the tarsi brownish, pubescence chiefly black.
Wings clear, stigma yellowish, veins brown, no appendix;
first posterior cell narrow at border, but open.
Erephopsis maculipennis, Macquart, Dipt. Exot., Suppl. iv.
p- 20 (1849) ; Schiner, Reise Novara, Dipt. p. 99 (1868)
[ Pangonia] ; Ricardo, Ann. & Mag. Nat. Hist. (7) v.
p- 106 (1900); id. (8) xvi. p. 28 (1915).
A series of specimens from South Australia appear to be
this species, Judging from Macquart’s description, though
there are a few discrepancies ; his type came from the East
Coast of New South Wales.
The wings have an appendix and two brown spots, which,
however, amount to little more than dark shading on the
transverse veins. Abdomen black, at base testaceous, with
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 14
he oe
210 Miss G. Ricardo on Tabanidze from
median black spots on the first two segments. Antenne
red, palpi with the second joint a little longer than the first,
which is blackish; the second one reddish, black ateborders,
concave and broad, ending in an obtuse point.
Erephopsis lasiophthalma, Boisduval, Voyage ‘ Astrolabe,’
Zool. 11. p. 666 (1332) [ Pangonia|; Macquart, Suites
& Buffon, i. p. 193 (1834); Walker, List Dipt. Brit.
Mus. v., Suppl.i. p. 1389 (1854) ; Ricardo, Ann. & Mag.
Nat. Hist. (7) v. p. 154 (1900).
The type was described from Cape Jervis, 8. Australia.
Erephopsis contigua, W1k., is not identical as Walker stated.
Pangonia fuliginense, Boisduval, is from New Guinea;
whether it is the same as Lrephopsis lasiophthalma is
doubtful.
Three females from Mt. Kosciusko in N.E. Victoria,
and two females from Moonbar, New South Wales (Mar-
grave) (1915), in Brit. Mus. Coll., belong, I believe, to this
species, answering to the description by Boisduval. Whether
the specimens Macquart placed under this species are
“identical is doubtful.
The wings have one dark band crossing the base of the
discal cell and the apices of the basal cells, and the trans-
verse veins at fork of third vein and apex of discal cell are
shaded ; there is the rudiment of an appendix present; the
first posterior cell is narrowed at border, but open. <Abdo-
men reddish with a black median spot on the first three
segments, then usually darker at the apex. Antenne reddish
yellow. Palpi same colour, very short; the second joint not
much longer than the first joimt, very concave. Forehead
twice as broad anteriorly as it is at the vertex, with dark
furrows above, continued to the antennz. Walker’s species
has two very distinct dark bands on the wings. LE. maculi-
pennis differs in the wings, which are only shaded, and the
first joint of palpi is dark.
Evrephopsis guttata, Donovan, Lllust. Ent, i. Hym. et Dipt.
(1806) | Tabanus}.
One female from Queensland.
Erephopsis gibbula, Walker, List Dipt. i, p. 140 (1848);
Ricardo, Ann. & Mag. Nat. Hist. (7) v. pp. 112, 117
(1900) ; id. (8) xvi. p. 22 (1915).
Five females from Warren River, West Australia.
Australia and the Fiji Islands. 211
Erephopsis aureohirta, Ricardo, Aun. & May. Nat. Hist. (7)
v. pp. 112, 116, pl. i. fig. 10 (1900); id. (8) xvi.
p. 23 (1915).
Two females from Queensland.
Erephopsis doddi, 2 , sp. n.
Type (female) and others from Warren River, West
Australia (W. D. Dodd). The type is in the South
Australian Museum.
A species with shaded transverse veins on the wings.
Abdomen testaceous at base with median black spots and
blackish at apex and greyish or reddish segmentations.
Antenne blackish. Palpi very small, short, and concave.
Legs testaceous and blackish.
Length 15 mm.
Face reddish, with grey tomentum and long black hairs,
some white ones intermixed. Beard yellowish white.
Palpi testaceous, the first joint with long black hairs, the
second one conical and concave with curved upper border,
and a few short black hairs at apex. Antenne black, the
first two joints with long black hairs. Forehead reddish
brown with erey tomentose sides and with black thick
pubescence, broadest anteriorly, being quite a third broader
than at vertex; ocelli distinct. Thorax blackish with. two
narrow, grey, tomentose stripes on anterior half of dorsum
only ; pubescence as in EL. gemina, Walker. Abdomen very
similar to this last species, the third segment with a black
spot similar to the one on the second segment; hairs
on sides chiefly white, black on the third, fourth and fifth
segments; underside bright testaceous with a few white
hairs. Legs testaceous, but blackish on the upper sides of
- femora; pubescence black. Wings with tlie first posterior
cell closed or slightly open.
Pelecorrhynchus eristaloides, Walker, List Dipt. Brit. Mus. 1.
p- 193 (1848) [Silvius] ; Ricardo, Ann. & Mag. Nat.
Hist. (7) v. p. 102 (1900) ; White, Royal Soc. Tasmania
(1915), pt. 1., Diptera Brachycera, p.22.
One male from Huron River, Tasmania (Lea),
Genus Sitvivs. 3
Mr. Taylor has lately forwarded me a copy of his paper
(Proceedings Linnean Soc. of New 8S. Wales, 1915, vol. xl.
EAS
912 Miss G. Ricardo on Tabanidee from
pt. 4, Nov. 24th), containing eight new species of this
genus, which, with the four new species named by me in the
Ann. & Mag. Nat. Hist. (8) xvr., Oct. 1915, brings the known
species occurring in Australia to twenty. Seven new species
are now added, whieh are all distinct from those described
by Mr. Taylor, judging from his descriptions.
Silvius grandis, 2, sp. n.
Type (female) and another from Fortescue River, Ham-
mersley Range, North-West Australia, and another in the
Brit.. Mus. Coll. from N. Australia (J. R. Elsey), 57.13.4—
this latter in bad preservation. ‘The type is in the South
Australian Museum.
A lareze dark brown species; antennz, palpi, and legs
blackish. Abdomen with white-haired segmentations.
Length 15 mm.
Face \lackish, covered with grey tomentum and with
some white hairs. Palpi similar to those of a species of
Tabanus; large, black, stout at base, ending in a point.
Beard scanty, white. Antenne black, the third joint wide
at its base, the first two joints with black hairs. Hyes bare.
Forehead arrow, and narrower anteriorly than at vertex;
frontal callus narrow, pear-shaped, with long lineal exten-
sion; ocelli distinct; traces of golden-coloured pubescence on
forehead. Thorax (denuded) blackish brown, some white
hairs at sides and on shoulders. Abdomen blackish brown,
the segmentations white-haired, the other female has the
first and second segments partly yellowish brown; traces of
grey tomentum appear on segments ; pubescence on abdo-
men black ; underside blackish, with white-haired segmen-
tations. Legs blackish brown with black pubescence, the
tibia more reddish brown. Wings greyish, slightly tinged
yellowish browu; veins and stigma yellowish; no appendix.
Silvius fuscipennis, ? , sp. 0.
Type (male) from Claudie River, Queensland (7. A. R.).
Type (female) from Cape York, N. Queensland (J/ac-
gillivray).
Three females from Claudie River and one female from
Cape York. .
Tne types are in the Nat. Museum, Victoria.
A species apparently belonging to this genus, to be
recognized at once by the brown wings with one irregular
clear band and two clear spots.
Length 138-16 mm.
Australia and the Fiji Islands, 213
Female.—Face covered with yellowish-grey tomentum and
with a few white hairs. Beard white. Palpi fairly stout,
reddish yellow or darker in colour, Antenne blackish, the
first two joints rather large with black hairs, the third broad
at its base. Eyes bare. Forehead narrow, but a little wider
anteriorly, the frontal callus black, club-shaped, with an
extension not reaching the vertex ; forehead blackish with
yellowish-grey tomentum. Thorax and scutellum blackish
brown. Abdomen broad, blackish with narrow reddish-
yellow bands; underside the same, but more reddish brown.
Legs blackish. Wings sepia, the pale irregular band crosses
the bases of the first submarginal, the first posterior, the
discal, the fourth and ‘fifth posterior cells ; there is a small
clear spot at the base of the fork of the third longitudinal
vein, and a larger one on the apex of the discal cell,
embracing the base of the second and third posterior cells ;
stigma dark brown ; wing at base somewhat pallid.
In the female from Claudie River the palpi are blackish,
the abdomen on the first four segments is yellowish with
a black spot in the centre of each segment, the remaining
segments blackish with yellow segmentations ; underside
yellowish with broad whitish segmentations, darker at apex.
Male somewhat different from the female in the colouring
of the abdomen, which has black spots in the centre of each
of the first three segments, and the apical segments are
eutirely black ; pubescence on the yellow parts yellow, black
elsewhere; underside identical. Sides of thorax with bight
reddish-yellow hairs, dorsum deep black. Scutelium the
same, with black hairs. Wings paler brown than in the
female.
Silvius nigripennis, 2 , sp. 1.
Type (female) and another female from Claudie River,
North Queensland (J. A. Kershaw), 11/12 and 2/13. One
female from same place, 28.1. 14.
The type is in the National Museum, Victoria.
A species rather Tabanus-like mm appearance, and very
similar to Silvius grandis, sp. n.,1n all particulars, except
the wings; easily distinguished by its almost wholly brown
wings and black abdomen with white bands.
Length, type, 165 mm., the others 14 and 17 mm.
Face flat in centre, covered with grey tomentum and with
white pubescence. Palpi black, rather stout on their whole
length, ending in a short point. Antenne black, broad at
base of third joint, Tabanus-like in shape. Forehead slightly
214 Miss G. Ricardo on Tabanidee from
broader anteriorly, about four times as long as it is broad,
covered with grey tomentum; frontal callus brown, pear-
shaped, with “short ‘lineal extension’; pubescence scanty,
white ; ocelli distinct. Thorax blackish, covered with grey
tomentum ; pubescence black, with some appressed white
hairs. Scufellum same colour. Abdomen black with black
pubescence ; bands of white hairs on posterior borders of the
first four segmeuts, not reaching the middle of segment ;
underside black with three complete white bands. Leys
black with black pubescence. Wings dark brown, pale at
the extreme apex, with a clear streak below the stigma,
and the extreme edge of the posterior border of wing
al:o pale.
Silvius fergusoni, 9, sp. 0.
Type (female) from Nelson Island, Hawkesbury River,
New South Wales, 26. 2.16.
This handsome species, named after the donor, is nearly
allied to Silvius nigripennis, sp. n., from the northern part of
Australia ; but is distinguished by the pale base of wing.
These two. species, together with Si/vius grandis, sp. n., form
a group unlike the typical forms by their larger size and
dark abdomens marked with paler segmentatious,.and by
their coloured wings.
This species measures 17 mm.
Face covered with hght ashy-grey tomentum and with
some silvery-white hairs in centre of face, and longer, more
numerous ones in the furrows between the face and the
cheeks, joining the beard of same colour. Palpi blackish,
with some ashy-grey tomentum, and short dark hairs ; they
are large and stout with an obtuse point. Antenne black,
the first two joints with black hairs, the third large and
wide at the base with a distinct angle. Subcallus same
colour as face. Forehead same .colour, narrow, parallel,
about six times as long as it is broad anteriorly ; the trontal
callus black, shining, not reaching the eyes, pear-shaped,
with a long stout lineal extension reaching the ocelli.
Thorax blackish brown, somewhat shining, with two grey
tomentose stripes and grey at sides; pubescence chiefly
black ; some white hairs on the stripes; pubescence on
shoulders long, black, with tufts of white hairs at base of
wings. Scutellum the same colour. Abdomen blackish
brown, the first two segments with broad grey tomentose
bands, which are repregeuted on the remaining segments
only at the sides; these bands have white liars, “thick kest at
Australia and the Fiji Islands. 915
the sides, pubescence elsewhere black ; underside is iden-
tical, but the grey bands are present on every segment for
their whole width. Legs black, the fore coxz with whitish
tomentum and pubescence. Wings blackish brown, darkest
on the fore border in the submarginal cells, becoming paler
at apex and on posterior border, and almost clear in the
basal, anal, and axillary cells; stigma blackish ; veins brown.
It is distinguished from Silvius nigripennis by the pale
base of wing, and by the shape of the frontal callus and
forehead.
Silvius niger, 2, sp. v.
Type (female) from Helensburgh, New South Wales,
9.3.15. Was resting on branch of low shrub; when dis-
turbed it circled round my head, evidently seeking to bite
(EL. WF).
A small robust black species, with a broad abdomen and
clear wings. Eyes slightly pubescent. Antennze and palpi
dull reddish yellow. Legs blackish.
Length 1] mm.
Face covered with brownish tomentum, paler on cheeks,
and with some dark hairs in the centre; on the cheeks they
are much thicker and long, increasing in length as they
attain the bases of cheeks. Beard same colour. Palpi
reddish yellow, curved on their upper edges, which are
clothed with a distinct fringe of black hairs, the lower
edges have a similar fringe; they are somewhat stout at
base, ending in an obtuse point. Antenne rather a darker
shade than the palpi, the first two joints with black hairs,
the third joint with a very few at the base and on the
extreme tip; the tooth represented by an obtuse angle.
Forehead same colour as face, with black pubescence,
parallel ; the frontal callus dark brown, very narrow, keel-
like. Ocelli very distinct. Thorax sepia, with two broad
grey tomentose stripes and grey sides; a few scattered white
hairs on dorsum ; long black hairs on the grey tomentose
shoulders. Scutellum a little darker. Abdomen blackish,
grey tomentum on the first segment; on the second, third,
and fourth segments appear short white hairs on the sides
and in the middle, not, however, joining each other to form
a band; pubescence elsewhere black ; segmentations with
traces of grey tomentum ; underside black with very narrow
white-haired segmentations. Legs black; knees reddish
yellow, the tibiz obscurely so; pubescence black. Wings
clear, veins yellowish brown, stigma yellowish.
216 Miss G. Ricardo on Tabanidee from
Silvius montanus, 2? , sp. 0.
Type (female) and another from Mt. Tambourine,
Queensland (4. M. Lea).
The type is in the South Australian Museum.
A blackish-brown species with yellowish segmentations
on abdomen and traces of median yellow spots. Antenne
reddish yellow. Legs blackish brown, some of the tibiz
yellowish.
Length 10 mm.
Face covered with greyish tomentum and with yellowish
tomentum near the eyes; pubescence consists of long blackish
hairs. Beard whitish. Palpi stout, covered with grey to-
mentum and with black pubescence, yellowish at extreme
base. Antenne with the first jomt greyish, covered with
black hairs, the second yellow with black hairs, the third
reddish brown, apex black. Forehead broader anteriorly,
covered with yellowish-brown tomentum, brown in the
centre ; the frontal callus elongate, apparently broader near
the vertex; forehead covered with black hairs; ocelli distinct.
Eyes bare. Thorax reddish brown with two grey stripes,
most distinct anteriorly, covered with greyish-yellow hairs
and with black pubescence intermixed, and longer black
hairs at sides. Scutellum brown, with black hairs. Addo-
men brown; the segmentations widely pale yellow, with
white hairs, which form median spots ; pubescence black ;
hairs at sides chiefly white ; underside blackish, with white-
haired bands. Legs blackish, the fore and mid tibiz ob-
scurely yellowish, the hind tibiz reddish brown ; pubescence
on legs black. Wings greyish, the transverse veins shaded, .
the other ones faintly shaded; no appendix ; stigma yel-
lowish ; veins brown.
Silvius insularis, ¢ , sp. 0,
Type (female) and another from Bathurst Island, N,
Territory.
The type is in the South Australian Museum.
A small narrow blackish-brown species, with a greyish
tomentose stripe on abdomen ; legs pale yellow, the femora
darker. Palpi long, narrow, with truncated tips.
Length 10 mm.
Face reddish, covered with grey tomentum, and with some
short white hairs. Palpi yellowish, with black hairs. An-
tenne yellowish, the first two joints with black hairs, the
third wide at its base. Forehead almost parallel; frontal
Australia and the Fiji Islands. 20%
callus blackish, quadrate, reaching the eyes, with a lineal
extension, covered with grey tomentum. Ocelli distinct.
Thorax and scutellum reddish brown, with white short
pubescence and some grey tomentum anteriorly on thorax.
Abdomen blackish brown, with a broad median stripe of
grey tomentum and grey segmentations, and scattered
white hairs on stripe and segmentations ; underside dark,
with grey segmentations. Legs with some brown colour
on the femora and on apices of tibize and tarsi ; pubescence
white, with some black on tibie and tarsi. Wings clear,
veins yellowish.
Silvius indistinctus, Ricardo, Aun. & Mag. Nat. Hist. (8) xvi.
p- 262 (1915).
Two females from Melville Island, N. Australia (W. D.
Dodd).
Five females from Bathurst Island, Northern Territory.
One female from Coen River, Cape York Peninsula,
N. Australia (W. D. Dodd).
Ectenopsis australis, @ 3, sp. u.
Male (type) from Milson Island, Hawkesbury River.
This type is not in the Brit. Mus. Coll.
Female (type) from Sydney (C. Gilbons), 16.12.14.
A species nearly allied to Kctenopsis vulpecula, Wied., but
distinguished {rom it by the very distinct stripes on thorax
and by the darker, not uniform, colour of abdomen and of
the face. The legs, which, however, appear to be variable
in colouring in the typical species, are here reddish yellow ;
the tarsi dusky.
Female.— Face and forehead chamois-coloured, with some
grey tomentum. dnitenne with the first two joints pale
reddish yellow with black hairs, the third joint with the
first division raw-sienna in colour, the next two divisions
dusky with grey tomeutum, and the remainder blackish.
Thorax covered with yellowish-grey tomentum, with a
narrow median brown mahogany stripe, becoming broader
beyond the suture, and a broad one of the same colour
at each side. Scutellwm uniform brown mahogany-colour.
Abdomen the same colour, with paler grey tomentose seg-
mentations, developing on the second, third, and fourth
segments into triangular median spots; underside almost
a uniforin paler shade. Appendix ef wing lone.
The male is similar, but the triangular median pale spots
218 Miss G. Ricardo on Tabanide from
of abdomen are absent. Legs duskier. The following note
on the eyes of the male is furnished by the collector :—
“Eyes brilliant green, with reddish-brown lines across at
lower third; lines sharply defined below, fading above ;
similar line round whole eye. Eyes appear reddish brown
in some lights.”
TABANUS.
Group LV.
Tabanus angusticallus, 3, sp. n.
Type (female) and another from Melville Island, N. Aus-
tralia (WV. D. Dodd). The type is in the South Australian
Museum.
A small greyish-brown species, which may be included
in Group IV., as the frontal callus is only represented
by a very small black line. Antenne and legs reddish
yellow.
Length 74 mm. (type) ; the other female 94 mm.
Face and forehead cov ered with yellowish-grey tomentum ;
pubescence on face consists of a few white hairs and longer
brown hairs below and on cheeks. Palpi long and narrow,
hardly incrassate at base, yellow, with short black puhes-
cence. Beard. consists of white hairs below and brown
above. - Antenne bright red-yellow, the two first joints
pale yellow with black hairs, the third joint very broad at
base, with a few black hairs at angle. Forehead parallel,
about five times as long as it is broad, with some black
pubescence. Eyes bare. Thorax, scutellum, and abdomen
the same colour, blackish brown, with grey tomentum, and
with short black pubescence ; a few yellowish hairs at base
of thorax, segmentations of abdomen narrowly pale; under-
side with wider yellow segmentations and white pubescence.
Legs yellow, with black pubescence. Wings clear, grey ;
veins and stigma very pale yellow; long appendix present.
Tabanus nemotuberculatus, Ricardo, Ann. & Mag. Nat.
Hist. (8) xiv. p. 388 (1915).
Three females from Cape York.
Oue female from Claudie River, -N. Queensland.
Tabanus nemopunctatus, Ricardo, Ann. & Mag. Nat, Hist.
(8) xiv. p. 888 (1915).
A series of females from Yeelanna, S. Australia, appear
Australia and the Fyi Islands. 219
to belong to this species, though the type came from
Queensland. These females have a broader forehead, wider
anteriorly, whereas in the type it is parallel.
Group VII.
Tabanus stranymanni, g, Ricardo, Ann. & Mag. Nat. Hist.
(8) xiv. p. 898 (1914).
One male from Cairns District (W. D. Dodd).
Tabanus rufinotatus, Bigot, Mém. Soc. Zool. de France, v.
p. 673 (1892) (Atylotus) ; Ricardo, Ann. & Mag. Nat.
Hist. (8) xiv. p. 892 (1914).
A series of females from Melville island, N. Territory.
Tabanus pseudoardens, Taylor, Austr. Inst. Trop. Med. 1911,
p- 66, pl. xiv. fig. 18 (1913) ; Austen, Ann. & Mag.
Nat. Hist. (8) xiii. p. 265 (1914) ; Ricardo, ibid. (3)
xiv. p. 272 (1915).
A series of females from Cairns District, Queensland.
Group VIII.
Tabanus victoriensis, Ricardo, Aun. & Mag. Nat. Hist. (8)
RIV p.2/0 (1915):
Females from Mt. Tambourine, S. Queensland.
Group IX.
Tabanus macquarti, Ricardo, Aun. & Mag. Nat. Hist. (8)
xiv. p. 277 (1915). [ Zabanus limbatinervis, Macq.
Dipt. Exot., Suppl. iv. p. 333 (1852). |
A series of females from Cairns District, Queensland.
Tabanus neogermanicus, Ricardo, Aun. & Mag. Nat. Hist. (8)
x1y. p. 283 (1915).
A series of females from Melville Island, S. Queensland.
Tabanus clavicallosus, 9 , sp. n.
Type (female) from Milson Island, New South Wales,
10. 1.15; other females from the same locality, and one
female from Sydney. (‘ Eyes of a dull brown colour when
alive’; note by collector. )
220 Miss G. Ricardo on Tabanidee from
The type is in the South Australian Museum.
A small species allied to Tabanus neogermanicus, Ricardo,
but differing in the darker colour of the abdomen, and in
the shape of the frontal callus which is rather distinctive
for this species.
Length 11-12 mm.
Face covered with yellowish-grey tomentum, almost devoid
of pubescence. Beard composed of sparse white hairs.
Palpi long and slender, pale yellow, a little stouter on the
basal half, ending im a long point; pubescence on the first
long, white, on the second joint short and black. Antenne
bright reddish yellow, dusky at the tips, the first two joints
with black pubescence, the third not very wide at its base,
with a slight tooth. Forehead and subcallus darker than
the face, the latter often appearing reddish through the
tomentum; the forehead with black short hairs beyond the
frontal callus, which is blackish brown, not reaching the
eyes; large and club-shaped, ending in a very short point,
which is, however, drawn out on some of the specimens ;
the whole callus is long, olten attaining half the length of
the forehead, which is about four times as long as it is
broad and is the same width throughout. Thorax blackish,
covered with yellowish-grey tomentum and with appressed
pale fulvous hairs; pubescence on the reddish shoulders
black, some white hairs at sides at base of wings. Scutel/um
identical. Abdomen blackish brown; posterior halves of the
segments with greyish tomentose bands, extending in the
middle as indistinct median spots, most discernible on
the second to the fifth segments; some very short white
pubescence is present on the segmentations and is longer
at the sides. Legs blackish; the femora with some grey
tomentum, and at their apices reddish yellow, or almost
wholly so; the tibize reddish yellow, dusky at their apices.
Wings clear, stigma yellowish brown, veins brown; a short
appendix present.
Dr. Ferguson states that this is a common sp¢cies.
Tabanus milsonis, 2 , sp. 0.
Type (female) and another from Milson Island, New
South Wales. (‘‘ Occurs on the cattle, but is rare and hard
to catch’: note by collector.)
A medium-sized species, blackish brown, with yellow
palpi and blackish antennz. Legs dusky in colouring.
Abdomen with grey tomentose bands and spots. Wings
with an appendix.
Australia and the Fiji Islands. 221
Length 16 mm.
Face covered with pale tomentum and some white short
hairs. Beard white. Palpi pale yellow, swollen on their
basal half, ending in a point which is about equal in length
to the basal half, some grey tomentum on this latter; pubes-
cence black, white below. Antenne blackish, the first two
joints reddish, the third joint broad at its base. Forehead
parallel, about five times as long as it is broad ; the frontal
callus chestnut-coloured, oblong, not reaching the eyes, with
a lineal extension; forehead darker than face, with black
hairs. Yhorax blackish brown, with two narrow grey tomen-
tose stripes and grey at the sides. Scutellum ideutical.
Abdomen blackish brown with broad grey tomentose bands,
extending into blunt triangular spots in the middle; the
ground-colour under the tomentum often appears reddish,
and the sides are reddish yellow ; pubescence on segmenta-
tions white, and on sides, elsewhere black mixed with the
white at sides, and rather long and abundant at sides;
underside reddish brown with grey tomentum. Legs dusky
in appearance, the femora with grey tomentum, the tibiz
obscurely reddish; pubescence chiefly black, some white
hairs on the femora and long ones on their under sides.
Wings large, clear; veins blackish brown, with an appendix ;
stigma yellowish.
Group X.
Tabanus nigritarsis, Taylor, Report Austr. Inst. Tropical
Medicine, 1911, p. 18 (1913); Ricardo, Aun. & Mag.
Nat. Hist. (8) xiv. p. 288 (1915).
Two females from Northern Territory.
Tabanus sanguinarius, Bigot, Mém. Soc. Zool. de France, v.
p. 675 (1892) [ Atylotus]; Ricardo, Aun. & Mag. Nat.
Hist. (8) xiv. p. 287 (1915).
Series of females from Mt. Tambourine, S. Queensland.
Tabanus kershawi, 2 , sp. n.
Type (female) and another from Claudie River, N. Queens-
land (J. A. Kershaw), 11/12 and 13. 2. 14.
The type is in the National Museum, Victoria.
A small black species, at once distinguished by the pale
tibiz and the reddish-yellow antenne.
Length 10 mm.
222 Miss G. Ricardo on Tabanides from
Face black, with some brown tomentum and black hairs.
Palpi blackish, very stout, ending in a very short point ;
pubescence black. Beard black. Antenne reddish yellow,
with a very small angle on the third joint ; some black hairs
on the first two joints. Subcallus shining black. Forehead
a third narrower anteriorly, about eight times as long
as it is broad anteriorly, covered with grey and brown
tomentum ; frontal callus small, oblong, reaching the eyes,
with a lineal extension. yes bare, with traces of three
stripes. Thorax, scutellum, and abdomen blackish with some
few appressed grey hairs on thorax and scutellum, and on
segmentations of abdomen; pubescence at sides black. Legs
black, the fore tibiz whitish, darker at extreme apex, the
middle tibize and the hind pair wholly whitish; fore tarsi
black, the others almost wholly whitish. Wings clear,
veins and stigma yellow; no appendix.
Group XI.
Subgenus THERIOPLECTES.
Species with pubescence on the eyes.
Tabanus regis georgii, Macquart, Dipt. Exot. i. p. 132
(1838); Ricardo, Ann. & Mag. Nat. Hist. (8) xvi.
p- 276 (1915).
One female from S. Australia (Rev. A. P. Burgess).
(“ Eyes in this species are dull-coloured’’: note by the
collector.)
Tabanus circumdatus, Walker, List Dipt. 1. p. 185 (1848) ;
Ricardo, Ann. & Mag. Nat. Hist. (8) xiv. p. 280
(1915); White, Royal Soc. Tasmania, 1915, pt. u.
p. 14.
(Tabanus nepos, W\k.; abstersus, Wik. ; brevidentatus,
Macq. ; and hebes, WIk., are all synonyms of this species. )
A series of females from Yeelanna, 8. Australia, and from
Swansea, Tasmania.
Tabanus cirrus, 2? , 8p. D.
Female (type) from Milson Island, Hawkesbury River,
New South Wales.
A stout, medium-sized, black species, distinguished by the
Australia and the Fiji Islands. : 923
tufts of white hairs on the thorax at base of wing and by
the white-haired fringe of scutellum.
Length 15 mm.
Face covered with ashy-grey tomentum and in the centre
with long, fairly dense, white hairs. Beard white: Palpi
yellowish with grey tomentum, stout at base, ending in a
rather short point; the pubescence scanty, chiefly pale.
Antenne dusky, the first two joints with black hairs. yes
very distinctly pubescent. Forehead broad, about three
times as long as it is broad anteriorly, where it is distinctly
wider than at vertex; forehead and subcallus a Itttle darker
than face ; frontal callus bare, protuberant, almost reaching
the eyes, pear-shaped, with a short lineal extension, pitchy-
brown in colour; pubescence on forehead black. Thorax
blackish brown, with two grey tomentose stripes and another
on each side below the suture ; shoulders reddish with black
hairs; a tuft of white hairs below base of wings, and ancther
continued to the scutellum; pubescence on dorsum black.
Scutellum blackish brown, fringed with white hairs on its
posterior border. Abdomen blackish brown, with broad
greyish-white tomentose segmentations and traces of white
hairs on them, which are distinct on the side edges of the
segments ; pubescence elsewhere on dorsum black. Legs
black, the fore cox covered with ashy-grey tomentum and
with white hairs; the tibiz-reddish yellow, black at their
apices; femora and tibize with chiefly white pubescence.
Wings clear, veins blackish; an appendix present; stigma
brownish.
Tubanus neocirrus, 2 , sp. n.
Type (female) from Swansea, Tasmania (Lea), and another
female from S. Australia.
The type is in the South Australian Museum.
A black species with whitish spots and segmentations om
the abdomen ; smaller than Tabanus cirrus, sp. n.
Length, type, 12 mm.; the other female 10 mm.
It differs from Tabanus cirrus in the following par-
ticulars :—Palpi slender, the long point nearly as long as
the slightly incrassate basal part, covered with white
pubescence. Third joint of antenne broad at base. Pubes-
cence on thorax black with many white hairs, which are
_long anteriorly and shorter posteriorly. Abdomen with a
distinct stripe composed of white-haired median spots: ;
segmentations also white-haired, thickest at the sides.
224 On Tabanidw from Australia &e.
Tabanus postponens, Walker, List Dipt. i. p. 179 (1848);
Ricardo, Ann. & Mag. Nat. Hist. (8) xvi. p. 282
(1915).
Males and females from South Australia.
The femora are sometimes dark.
Tabanus pacificus, 2, sp. n.
Type (female) and another female from Suva, Fiji [slands,
30. vi. 1910 and xii.1910 (Dr. P. H. Bahr). Presented by
London School Tropical Medicine.
For list of species from these islands and surrounding
region, see Ricardo, Ann. & Mag. Nat. Hist. (8) xiii. p. 476
(1914).
A narrow-bodied dull-coloured species; palpi slender,
blackish. Antenne dull reddish yellow. Forehead narrow.
Legs dull yellowish and brown.
Length 12 mm.
Face covered with tawny tomentum. Beard scanty,
brownish. Palpi long and slender, almost the same width
throughout; blackish with some greyish tomentum and a
few black hairs. Antenne tawny, the first two joints yel-
lowish with black hairs, the third joint with a small but
distinct tooth on its first division at base, clothed at tip
with some black hairs; the next three divisions are equal
in length, yellowish, the last one dusky and nearly as long
as the three preceding ones together. Forehead narrow,
shghtly narrower anteriorly, about six times as long as it
is broad anteriorly ; frontal callus shining dark chestnut-
brown, oblong, reaching the eyes, with a long, stout, lineal
extension reaching nearly to the vertex. Thorax muimmy-
brown, with some traces of grey tomentum and scme
yellowish short hairs, but chiefly black ones, on the dorsum.
Scutellum the same colour with black pubescence. Addomen
mummy-brown, appearing darker by reason of the rather
thick, short, black pubescence; there are small, whitish,
yellow-haired, median spots on each segment except the
last two, and indistinct, narrow, grey segmentations, not
visible on the apical segments, and almost absent on the
other female; underside with white-haired segmentations
and no spots. Leys mummy-brown, the femora yellowish,
pubescence on legs black. Wings grey, faintly tinged brown
on fore-border and on cross-veins; stigma and veins brown;-
appendix present, but very short.
On new Species of Hee.natopota from India. 225
XVI.—New Species of Hematopota from India.
By Gerrrupe Ricarpo.
THeEse species were handed to me for identification by the
Imperial Bureau of Economic Entomology, the types to be
given to the British Museum Collection.
Hematopota montanus, 2, sp. 0.
Type (female) and another from Bababuddin Hills,
Mysore, 4700 ft., vi. 1915 (Ramakreshna Coll.).
A small species allied to Hematopota latifascia, Ricardo
(‘ Records Indian Museum,’ iv. p. 355, 1911), having one
broad band extending across the apex of wing, but dis-
tinguished from it by the absence of a black band on the
face.
Length of type 8 mm.; the other female 10 mm.
Face covered with grey tomentum and with small dark
brown spots on the upper part, in the other female there is
a trace of a black band on the lower edge; there is also a
dark spot between the antennz; hairs on face scanty, white.
Palpi pale yellow with black hairs. Antenne long and
slender; the first joint as long as the first annulation of the
third joint, only slightly incrassate, yellow with black
pubescence ; the second one very small, blackish; the third
yellowish at base, then dusky ; slender throughout. Fore-
head same colour as face; the frontal callus blackish brown,
reaching the eyes, straight on both borders; the paired spots
the same colour, touching the eyes, but not the band; some
black pubescence on forehead. Thorax yellowish brown
with appressed yellow hairs and some black ones ; scuéelium
same colour with black hairs. Abdomen mummy-brown,
with pale yellow segmentations and an obscure pale median
stripe; pubescence black, some yellow hairs at apex ; under-
side paler. Legs yellowish with blackish-brown rings ; fore
femora rather dusky and the others dusky at their apices;
tibize at base and apex blackish and a black ring in middle,
the fore tibize white at base, fore tarsi wholly biack, on the
other tarsi the basal joint is pale yellow; pubescence on
dark parts black, on the pale parts white; fore tibiz in-
crassate. Wings with the usual rosettes; the apical band
starts from the junction of the first vein with the border
and attains the posterior border, it is rather sinuous on both
borders; on the posterior border pale spots are present in
every cell; veins and stigma brown. -
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 15
226 Mr. C. IT. Regan on the
Hematopota hindostani, 2, sp. n.
Type (female) and two other females from DBababuddin
Hills, Mysore, 4700 ft., vi. 1915 (Ramakrishna Coll.).
A species in the same group as H. montanus, sp. n., but
rather allied to H. assamensis in the wing having a single
band at apex, not reaching the border; face with a black
band; palpi rather stout and short, pubescence on them and
on face rather thick.
Length 8 mm.
Face covered with grey tomentum and with some white
hairs, and a black band on upper part of face. Palpi
covered with grey tomentum and with thick black pubes-
cence ; some white hairs on the first joint below. Antenne
dull reddish, the third joint dusky at apex, the first joint
rather stout, shining, with black hairs, the second one very
small, same colour, the third a little broader at base, the
first joint longer than the first annulation of third joint.
Forehead black, covered with brownish tomentum. Frontal
callus black, shining, reaching the eyes, with a straight
border and a black spot between the antenne. Thorax
muumy-brown, with three pale stripes anteriorly, the side
ones ending in a pale spot at the suture and there is another
pale stripe posteriorly at the sides; pubescence chiefly
consists of pale appressed hairs; scutellum same as thorax.
Abdomen same colour as thorax, with pale yellow segmen-
tations aud a pale median stripe; pubescence almost nil;
underside yellowish. Legs yellowish with darker rings;
apices of femora and the tarsi dark. Wings with the usual
rosettes, the apical band short, not reaching far beyond the
fork of third vein; sometimes a pale spot is visible on
border, but never joining the band; veins and stigma -
brown.
XVII.—The Fishes of the Genus Clupea.
By OC. Tate Keaan, M.A.
(Published by permission of the Trustees of the British Museum.)
In a preliminary arrangement of the Clupeoid fishes in the
collection of the Natural History Museum, [ had put together
examples of Clupea arcuata, Jenyns, aid specimens received
from New Zealand as Clupea antipodum, Hector, as be-
longing to a genus distinct from Clupea; on going through
Fishes of the Genus Clupea. 227
the material a second time I find that such a genus cannot
be maintained, and that Clupea must be enlarged by the
addition of C. arcuata and the closely related C. melanostoma
as well as by the species generally known as C. antipodum,
but here called C. muedleri, Klunz., since reference to the
original description shows that the name C. antipodum should
be given to the species recently described by meas C. holodon.
The synopsis given in my former paper (Ann. & Mag. Nat.
Hist. (8) xviii. 1916, p. 3) may be modified to include the
three additional species.
I. Pelvic fins 9- (rarely 8- or 10-) rayed; vomer toothed. D. 17-20.
A. 14-20. Vertebrae 50-59. 40 to 51 gill-rakers on lower part
of anterior arch. (Northern species.)
Ventral scutes: all keeled i7:.2.45 308 Aah clo ok ates 1. harengus,
Ventral scutes in front of pelvic fins not keeled .. 2. pallasi.
II. Pelvic fins 8-rayed. D. 15-i9. <A. 17-21. Vertebre 42-51.
35 to 40 gill-rakers on lower part of anterior arch, (Southern
species.)
A. Vomer toothed.
Depth of body 4} in length. Scales 48/13-14...., 3. antipodum.
Depth of body3 to33in length, Scales45-44/10-1]1. 4. mwelleri,
B. Vomer toothless,
Vertebree 49-51 ..... Sabo ceitenicaonc Bors: axdher Aten 5. fuegensis,
Mental: 4 1 Ua: gel ds 7 Palate tants os 8h ee a'ee 6. bassensis,
LUI. Pelvic fins 7-rayed ; vomer toothless.
A. 34 to 40 gill-rakers on lower part of anterior arch.
A 17-21. Scales 44-50/12-15............2...0-- 7. sprattus,
B. 25 to 380 gill-rakers on lower part of anterior arch.
A 22-23. Scales 42/15. Depth 3 to 33 in length. 8. arcuata.
A 17-20. _ Scales 42/11-12. Depth 83 to 4 in
Peni h For tn da aeldcaces > psoas Lhamisegericat a = 9. melanostoma.
Descriptions of C. antipodum, C. muelleri, C. arcuata, and
C. melanostoma follow.
Clupea antipodum.
Clupea sprattus, var. antipodum, Hector, Edible Fish. N. Zealand,
p. 133 (1872).
Clupea holodon, Regan, Ann. & Mag. Nat. Hist. (8) xviii. 1916, p. 5.
An elongate-ovate patch of teeth on vomer, a broad-ovate
patch on tongue. Depth of body 44 in the length, length of
head 4 to 43. Diameter of eye 4 in length of head ;
maxillary extending to below anterior part of eye; 36 gill-
rakers on lower part of anterior arch. Numerous radiating
grooves at free’ margin of scales ; 48 scales in a longitudinal
series ; 13 or 14 ina transverse series ; ventral scutes keeled
and pointed, 21+12. Dorsal 16-17 ; origin a little nearer
to base of caudal than to end of snout. Anal 16-18.
228 On the Fishes of the Genus Clupea.
Pelvics 8-rayed, inserted below origin of dorsal. Caudal
peduncle longer than deep, Vertebree probably not fewer
than 46.
Stewart Island.
The type, from the Foveaux Straits, was 150 mm. long ;
the specimen in the British Museum measures 122 mm, in
total length.
Clupea muelleri.
Clupea muelleri, Klunzinger, Sitzungsb, Akad. Wien, Ixxx. 1880,
p. 416,
An elongate patch of teeth on vomer, an ovate patch
on tongue, Depth of body 3 to 34 in the length, length of
head 33 to 32. Diameter of eye 34 in length of ‘head ;
maxillary extending to below anterior part or middle of eye; 36
to 39 gill-rakers on lower part of anterior arch. No radiating
grooves at free margin of scales; 43 or 44 scales in a
Jongitudinal series, 10 or 11 in a transverse series ; ventral
scutes strongly keeled and acutely pointed, 19-204 9-11.
Dorsal 15-16; origin nearer to base of caudal than to end
of snout, Anal 16-18. Pelvics 8-rayed, inserted below or
a little in advance of origin of dorsal. Caudal peduncle
deeper than long. Vertebree 42.
New Zealand.
Five specimens, 90 to 100 mm. in total length, from Otago
and Canterbury, received from the Otago and Canterbury
Museums as examples of C. antipodum,
Clupca arcuata.
Clupea_arcuata, Jenyns, Zool. ‘ Beagle,’ Fish. p. 184 (1842) ; Giinth,
Cat, Fish. vii. p. 442,
A narrow strip of teeth on tongue; palate toothless,
Depth of body 3 to 3} in the length, length of lead 4 to 42.
Diameter of eye 3 to 32 in length of head ; maxillary
extending to below atehint 4 a eye; 28 gill-rakers on
lower part of anterior arch. No grooves at free margin
of scales; about 42 scales in a longitudinal and 15 in a
transverse series ; ventral scutes strongly keeled and acutely
pointed, 18-19+9-10. Dorsal 16-18; origin nearer to
base of caudal than to end of snout. Anal 22-23. Pelvics
7-rayed, inserted below or a little in advance of origin of
dorsal.
Uruguay to Northern Palagonia,
Barnacles from the Hull 0) the ‘ Terra Nova.’ 229
Five examples, 60 to 90 mm. long, three from Montevideo,
and two, from Bahia Blanca, types of the species, kindly
lent to me for examination by Mr. C. Forster Cooper.
Clupea melanostoma.
Pomotobus melanostomus, Eigenmann, Proc. Washington Acad. viii.
1907, p. 452, pl. xxiii. tig, 6,
Mouth toothless. Depth of body 34 to 4 in the length,
length of head 44 to 5. Diameter of eye 3 to 34 in length
of head; maxillary extending to below anterior margin or
anterior + of eye; 25 gill-rakers on lower part of anterior
arch. No grooves at free margin of scales; about 42 scales
in a longitudinal and LL or 12 in a transverse series ; ventral
scutes strongly keeled and acutely pointed, 17-20+9-10.
Dorsal 15-16 ; origin nearer to base cf caudal than to end
of snout. Anal 17-20. Pelvics 7-rayed, inserted in advance
of origin of dorsal. Vertebree 43.
Rio de la Plata.
Eleven specimens, 65 to 80 mm, in total length.
XVIII.—Barnacles from the Hull of the‘ Terra Nova’ :
a Note. By L. A. BorRADAILeE.
WHEN the ‘Terra Nova,’ with the British Antarctic Expe-
dition on board, was at Lyttelton, New Zealand, in 1910,
barnacles weie removed from her bottom, Some of these
were included in the collection of Cirripedes taken by the
Expedition, upon which I have recently reported (Brit.
Antarct. (‘‘l’erra Nova’) Exped. 1910, Zool. iii. p. 127, 1916).
Others came into the possession of the Otago Museum, and
are mentioned by Mr. Jennings in an article on the Pedun-
culate Cirripedia of New Zealand, published in 1915 in the
‘Transactions of the New Zealand Institute’ (xlvii. p. 285).
Unfortunately, at the time of writing my report I had not
Mr. Jennings’s work before me, and there are consequently
between our papers certain discrepancies. ‘I'he object of
the present note is to call attention to and explain these, as
follows :—
1. My Lepas affinis is Mr. Jennings’s L. anatifera, var. c.
If I had seen Mr. Jennings’s description of this torm, I
should still have thought it advisable to name it as I did,
because in my view it is as nearly related to ZL. hilld as
to L. anatifera, and all three forms are of the same rank,
23) Geological Society.
whether species or varieties. L. affints is undoubtedly a link
between ZL. anatifera and L. hilli, but in the circumstances I
have not proposed to reduce J. Ailli to the rank of a variety,
preferring to leave the discussion of its status till the genus
is next revised as a whole.
2. In the material at Mr. Jennings’s disposal were speci-
mens of Conchoderma virgatum and C. auritum from the
hull of the ‘Terra Nova.’ In the collection placed in my
hands neither of these species was represented from that
source. C, auritum, which was taken upon whales in New
Zealand waters by the ‘Terra Nova,’ is also reported by
Mr. Jennings from whales in the same neighbourhood.
PROCEEDINGS OF LEARNED SOCIETIES.
GEOLOGICAL SOCIETY.
November 8th, 1916.—Dr. Alfred Harker, F.R.S., President,
in the Chair.
The following communication was read :—
‘Aulina rotiformis, gen. et sp. nov., Phillipsastrea hennahi
(Lonsdale), and the Genus Orionastrea. By Stanley Smith,
GA. De, FGS.
The primary object of the present communication is a descrip-
tion of a new and interesting coral genus of colonial habit, Aulina,
obtained from the highest limestone that can be associated with
the Lower Carboniferous—the Fell Top Limestone of North-
umberland and its equivalent horizon in Teesdale, the Botany
Beds.
Since this form has been confounded with another Carboniferous
species, well known under the name of ‘ Phillipsastrea radiata
(S. Woodward),’ it has been found advisable, in fact necessary,
to extend the original scope of the paper so as to include a
revision of the genus Phillipsastrea and a description of ‘Ph.
radiata’ and its allies, which I have grouped together under a
new generic name, Ordonastrea. Several type-specimens, including
that of Phillipsastrea hennahi (the genotype of Phillipsastrea),
are described and figured.
The new genus from the Fell Top Limestone is a very distinctive
form, on account of the remarkable annular wall developed within
the theca, and may prove of considerable value as a zonal
index.
Geological Society. 231
The corallum in this gents, as also in Phillipsastrea and in
Orionastrea, represents a stage in colonial development in which
the epitheca of the individual corallites has entirely disappeared,
and these are consequently united by their dissepimental tissue—
a type of colony to which the term ‘ Astreiform’ may be applied.
Diagnoses.
Aulina rotiformis.—The corallum is massive, and the corallites
are united by their extrathecal tissue ; all the septa dilate at the
theca, and those of the major cycle again dilate at their axial edges,
in such a manner as to fuse together, and so build a cylindrical wall
or tube within the theca. The structure of the form is in most
respects similar to that of Phillipsastrea, but it appears to carry
forward the septal characters peculiar to that genus to a further
stage of development.
Phillipsastrea.—the corallum is composite and massive; the
corallites are united by their dissepiments, or are only separated by
a thin epitheca; in the former case, the septa are often confluent.
Major and minor septa dilate at the theca; the latter terminate
there, and the major septa attenuate and advance into the intra-
thecal region, and there often dilate again at the axial edge. ‘The
central part of the corallite is occupied solely by tabulz.
Orionastrea.—The characters of this genus are essentially those
of Lithostrotion, but of a modified form. The corallum is compo-
site and massive, and the corallites are either defined by a thin
epitheca, or, in the more typical instances, by no epitheca at all ; in
this latter case the corallites are united by their dissepiments and
the septa are confluent.
The distinguishing characters of the three species recognized and
described are as follows :—
(1) O. ensifer (Edwards & Haime). Septa not confluent. Columella present.
(2) O. phillipsi (McCoy) ........... Septa confluent. Columella present.
(3) O. placenta (McCoy) ............ Septa confluent. Columella absent.
November 22nd, 1916.—Dr. Alfred Harker, F.R.S., President,
in the Chair.
The following communication was read:—
‘Characeze from the Lower Headon Beds.’ By Clement Reid,
F.R.S., F.L.8., F.G.8., and James Groves, F.L.S.
The investigations here recorded have been made at Hordle Cliffs
(Hampshire), where the strata, below the superficial gravel, belong
232 Geological Society.
entirely to the Lower Headon Beds, and consist of freshwater
and brackish-water (more .or less calcareous) deposits, laid down
apparently in wide shallow lakes and lagoons. Such habitats are
the most favourable to the growth of Characez, and several of the
beds have yielded numerous remains of these plants.
There is a great diversity in the fruits of Chara found, repre-
senting evidently a number of species, belonging to several different
sections or genera. With the exception of a few, which are possibly
abnormal variations, the fruits can be roughly grouped under the
following eight types :—
I, Tuberculate series. (Type of C. twberculata Lyell= Kosmogyra Stache,
emend.)
(a) Spherical.
(b) Obovoid or pyriform, with distinctly prolonged base.
II. Non-tuberculate series.
(c) Large spherical, diam. c. 1 mm. (type of C. medicaginula Brongn.),
(d) Large ellipsoidal (type of C. helicteres Brongn.).
(e) Medium-sized, subglobose, tapering more or less at both ends.
(f) Cylindric-ellipsoidal, showing more numerous striz.
(g) More or less pyriform: that is, definitely tapering towards the
base.
(h) Minute, subglobose-ovoid (long.=c. 350 to 500 p).
It is difficult to determine the exact number of species found,
on account of the extreme variability of some of the forms, but
the Authors consider that at least twelve may, for the present, be
conveniently treated as distinct.
The vegetative remains are comparatively few, consisting of
minute portions of stems and branchlets of different diameters,
and these it is impossible at present to connect with any particular
types of fruit.
Though investigations of some earlier formations have shown
that there are extinct forms of Characeze exhibiting important
points of difference from their living representatives, the remark-
ably distinct and characteristic oogonium of five elongated spirally-
twisted cells has remained constant certainly as far back as the
Inferior Oolite, and it is only in earlier formations that any doubt
arises as to whether bodies are or are not Chara fruits.
Characeze are found in still fresh or brackish water all over the
world, under widely different conditions as regards heat, etc., and
may therefore be expected to occur in almost all freshwater
formations. For these reasons it is suggested that the fruits of this
group of plants, when more widely collected, may prove of consider-
able value as zonal fossils for the correlation of lacustrine deposits
lying in isolated basins. Doubtless, on account of their small size,
the Characez have in the past often been overlooked.
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CONTENTS OF NUMBER 110.—Kighth Series,
he oF Ks ‘a Ss at
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XII. Coleoptera, Hateedhia Goialading Tenebrionide) from the
Seychelles Islands and Aldabra.. UF: GEORGE Cmarres Cnamerox,
PLS. (Pita VES eee ey ee Cae mG eee Ae ae
XIII. On new Species of Indian Cureulionide. —Part WL. 2
Gur°A. K, Maninier, DRGs eG oa isa men et ‘188
XIV. A Revision of the Clupeid Fishes of the Genus Pdlonula
and of Related Genera in the Rivers of Africa. By C. Tarn Regan, — eet
BOA oe So Ss ES Soy et arene gt pee aren ae ROB
XV. New Species of Tabanide from Australia and ‘the Fiji a ea
Islands. By Ggrrrupp Ricarpo ............ bee ay ae PU pe
XVI. New Species of Hematopota from India. By GerrrupE
Backs vgs eae i oe Bewethes Te wets eae ey eae re a2.)
Maa ea BARES oe ee ee joe eT TN ney 226
XVIII. Barnacles from the Hull of the ‘Terra Nova’: a Note. eg
By-L. As BOMB AER 2 eis os SEER es wanes oh eae
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No. 111. MARCH 1917.
XIX.—WNotes from the Gatty Marine Laboratory, St. An-
drews.—No. XL. By Prof. M‘Inrosu, M.D., LL.D.,
F.R.S., &c., Gatty Marine Laboratory, University,
St. Andrews.
(Plates VII.-XIL. ]
On the Nervous System and other Points in the Structure of
Owenia and Myriochele.
Since the remarks on Owenia and Myriochele were made in
the volume on the British Marine Annelids lately issued by
the Ray Society, a few observations on both types were
carried out, though, unfortunately, no living forms could be
obtained ; yet Owenia formerly was cast on the beach at
St. Andrews in hundreds, whilst Myriochele is not uncommon
on the west coast of Ireland, and in certain foreign localities
it occurs in swarms. Such blanks, which may stretch over
many years in the British area, are in the case of the fishes
often regarded as evidence of serious diminution ; but, so
far as observed during a long period of years, neither in the
case of the fishes nor in the case of the invertebrates is there
much of a basis for this supposition. The two forms above
mentioned are of interest especially as regards their nervous
system, which differs from that in the majority of the
Polychets in having the cephalic system as well as the
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 16
234 Prof. M‘Intosh’s Notes from the
nerve-trunks wholly hypodermal. The latter arrangement
of the ventral cords is that characteristic of most Polychets
—yet about ten families have their great ventral nerve-
trunks enclosed by the muscular tissues of the body-wall,
besides the basement-layer, hypoderm, and cuticle, showing
how uncertain any single factor is in the classification of this
group. In the Archiannelid Profodrilus the nervous system
agrees with that in Owenia and Myriochele in being mainly
hypodermic, and in Saccocirrus and Sternaspis (though this
is not a Polychzet) the cerebral ganglion is similarly situated
and the ventral nerve-cord is not segmented into ganglia.
They contrast thus with the Nemerteans, in which the
cephalic ganglia are internal and the longitudinal cords either
enveloped in the muscular walls of the body or entirely
within them. It is further interesting, in comparing
the Nemerteans with the Polychets, that no Polychzt
possesses the proportionally large nerve-supply to any organ.
—a supply, moreover, more bulky in its distribution than in
its origin, and which undergoes remarkable changes of form,
both in contraction and dilatation—as that of the Nemertean
proboscis. Hence its lattice-like arraigement gave rise
to the term “ elastic layer”’ in the early memoirs. ‘This
feature is as noteworthy as the passage of the proboscis
between the dorsal and ventral commissures of the cephalic
ganglia. Some consider that this arrangement of the nerves
causes it to be an organof sensation ; but it is often thrown
off when brought into contact with foreign bodies, and,
though renewed, its functions for the interval are in
abeyance. In the Ammocharide under consideration what
appears closely allied to nervous matter is distributed as
a continuous layer beneath the hypoderm of the gullet—a
condition much more primitive than the elaborate system of
the Nemertean proboscis or than the proboscis of a typical
Polychet such as Nereis.
In glancing at the literature of the subject, it is found that
the acute and accomplished Claparéde, familiar as he was
with the ordinary nervous system of the Polycheta, failed to
find the central nervous system in Owenia “qu il m’a été
parfaitement impossible d’en trouver la moindre trace chez
POwenia fusiformis sur les coupes d’individus conservés ” *,
and he even had difficulty in discriminating the ventral cord
in the fresh animal. Yet he had described and figured a
similar condition to that of Owenia in Telepsavus costarum
one of the Chetopteride, in which the central nervous
* Annél. Sédent. p. 129.
Gatty Marine Laboratory, St. Andrews. 235
system forms asubhypodermic band resting on the basement-
tissue, and which Claparéde interpreted as a transverse
commissure between the ganglia*, yet to be considered as
the representative of the cerebral ganglia, and the two eyes
in Telepsarus rest on it. The ventral cords, moreover, show
no ganglia and are wide apart. He does not allude to the
minute structure of the nerve-tissue.
In 1885 von Drasche + gave a careful account, with figures,
of the structure of Owenia “filiformis”’ as it occurred at
Trieste, dealing in the first instance with the external
characters, and especially the “ Lippenorgan” at the oral
aperture, the hypoderm and nervous system, the muscu-
lature, alimentary canal, coelomic cavity, and long mucous
glands. So far as he goes, the structure of these organs is
correctly described, with accompanying figures. He could
not satisfy himself as to the “ nephridia”’ “and the mode of
exit of the genital products. He observed no nerve-cells in
the minute structure of the central nervous system or in the
ventral cord, only a fibrillar structure and Leydig’s punctate
substance. The ventral cord showed no ganglionic enlarge-
ments. Below the epithelium of the alimentary canal a
strand similar in structure to the ceutral system is briefly
mentioned, but nothing definite is recorded concerning
the nerve- supply of the internal organs nor concerning the
nephridia.
In looking around for analogous relations of the central
ganglia, it is found that in Phoronis Caldwell t observed that
the central nervous system remained in the epidermis, and
therefore represented the primitive condition. In the adult
the central system is in the form of a post-oral ring, the anus
lying outside it. In Phoronis buski of the ‘Challenger’ §
the nerve-centre rests on a broad plate of basement- tissue,
with the hypoderm externally extending from the nephridia
forward to the centre of the whorl of tentacles on each side,
and it agrees precisely in minute structure with that in
Cephalodiscus and Owenia,
The central nervous system in Cephalodiscus dodeca-
lophus || occupies an area of considerable proportional size at
* Annél, Sédent. p. 127.
ae ees zur feineren ee der Polychaeten,’ Zweites Heft,
Woe pp. 1 22, 2 plates.
if Proc. Roy. Soc. vol. xxxiv. _p. 372,
§ Zoology, vol. xxvii. part 75, pp. 18-21, pl. ii, figs. 1 & 2, pl. iii,
figs. 1-5
| ‘Challenger,’ Zoology, vol. xx. part 62, p. 23, pl. vi. fig. 3, and pl. vii.
fig. 3.
16*
236 Prof. M‘Intosh’s Notes from the
the base of the plumes external to the median space, and ts
bounded externally by the thick coat of hypoderm and
internally by the basement-layer. It extends laterally in
the hypoderm along the basal region of the plumes and for
some distance along the dorsal side of the buccal shield. In
microscopic structure it is minutely cellular and granular,
intermingled with fibres, and corresponds generally in
position with that in Owenia and Myriochele.
Benham * (1896) described the central nervous system of
the “ Archiannelida”’ as in contact\with the epidermis, and
pointed out that in some Polycheta it holds a similar position
in the epidermis ; but he does not mention this condition
in the Ammocharide, which he associates with his Spioni-
formia, nor in the Chetopteride included by Levinsen and
himself under the same group or suborder.
Gilson (1897-98) devoted much attention to the structure
and function of the various parts of Owenia. Besides a
careful account of the remarkable secreting glands +, which
form such prominent organs, he has furnished an extended
description of the “valves septales” ¢t and of “ cellules
musculo-glandulaires” in the body-wall§. Perhaps the most
important contribution is that connected with the ‘ valves
septales,” wherein he gives a systematic description of each
septum, with its functions, one of the most striking being
the second septum (7. e., between the fourth and fifth seg-
ments), his sphincter muscle being in the position of the
ordinary oblique muscle at its insertion over the nerve-cord.
The muscular arrangements of this septum are specially
connected with the ccelomic fluid and the branchie. After
describing the special apertures in the septa and their
mechanism, he shows that apertures at several of these
connect the celom with the exterior ; that in the sixth seg-
ment two zigzag cutaneous canals springing from funnels at —
the septum between the sixth and seventh segments perform
the function of genital ducts, since nephridia are absent—a
feature of a peculiar character in a Polychet. In his paper
on the musculo-glandular cells he states that a peritoneal
membrane or ccelomic coat proper is absent in Owenia, thus
resembling such forms as the Nematodes, Acanthocephali,
many Annelids and Archiannelids. The body-wall is formed
by a combined musculo-glandylar coat—that is, it cannot be
separated into a muscular and a glandular layer. The inner
* Camb. Nat. Hist., Worms, Rotifers, and Polyzoa, pp. 248, 255, & 325.
+ ‘La Cellule,’ t. x. fasc. 2. ~
{ Ibid, t. xii. fase. 2. § bid. t. xiv. fase. 2.
Gatty Marine Laboratory, St. Andrews. 237
region of this coat secretes albuminoid substances, fat, and
urinary products. As will be shown subsequently, such is
a misapprehension of the structure of the peritoneal surface,
probably owing to the condition of the accomplished Belgian
author’s material.
In connection with Gilson’s opinion, for it is nothing
more, that the hypodermic canals in the sixth segment are
genital ducts, it is noteworthy that Arnold Watson observed
the reproductive elements in Owenia issuing from two pores,
to the right and left of the anus, a portion of the posterior
end of the body projecting from the anterior aperture of the
tube. Thus Gilson’s theory of the advantages of the ante-
rior opening of the hypodermic tubes (his genital ducts)
lapses, were it only by the thrusting out of the much more
delicate tail anteriorly.
Ogneff* (1899), working at the Naples Station, took up
the subject of Gilson’s ‘ cellules musculo-glandulaires”’ in
Owenia. In his preparations he found within the muscular
layer of the body-wall a protoplasmic and cellular layer
which lined the celom. In the muscle-fibres of the longi-
tudinal coat themselves were spindle-shaped cells with nuclei,
as Schwalbe first described in the muscles of worms and
Jamellibranchiate mollusks, and also on the surface of the
muscles in a protoplasmic layer. Over these, however, is a
layer of peritoneal cells, which are cup-shaped, with rounded
inner or deeper surfaces and flattened surfaces toward the
celom, with an oval nucleus, fat, and granules like the white
of egg in the protoplasm. A fine protoplasmic network
stretches from these amongst the muscle-cells. He thought
there were as many as fifteen to twenty peritoneal cells to
one muscle-cell. He did not consider that Gilson’s “‘ muscle-
gland-cells”’ existed in Owenia, the misapprehension being
due to the less elaborate methods of preparation and section-
making.
In 1900 a very interesting paper on Owenia fusiformis
was communicated to the Linnean Society by Mr. Arnold
Watson +, whose observations on the living animals are
noteworthy. His description of the lip-organ and its func-
tions, the occurrence of a prostomial pore, the discovery of
the emission of the sexual elements through twoccelomo-ducts
(anal pores), the structure and repair of the tubes, and the
rearing of the ova to the Mitraria-stage are the chief features
of this contribution to the life-history of the species.
* Biol. Centralblatt. Bd, xix. p. 156.
+ Journ. Linn. Soe. vol. xxviil. p. 259, pl. xxii.
238 Prof. M‘Intosh’s Notes from the
,
One of the latest contributions is a histological paper on
Owenia by Ziircher,*, who enters into the minute structure
of the muscles, showing that the long spindle-shaped muscle-
cells have a spiral character (formerly noted by Ogneff),
which in transverse section give them a barred aspect. A
circular muscular coat occurs only at the second dissepiment
and forward to the branchial lobes. This also has spindle-
shaped cells with nuclei. He combats Gilson’s view, as
Ogneff had previously done, that there is no line of demar-
cation between the muscular coat and the peritoneum, that
the nuclei are rare in the muscular fibres and by-and-by
vanish, and that it is impossible to distinguish the nuclei of.
the muscles and those of the gland- cells. He points out
that the peritoneal nuclei are generally rounded, whereas the
nuclei of the muscles are oval and flattened, with the long
axis in the line of the muscle-cell. He goes somewhat fully
into the histology of the circulatory system (his hemoceel),
the main trunk consisting of a dorsal vessel carrying the
blood forward and a ventral trunk carrying it backward.
The dorsal forms a blood-sinus round the: gut to the second
septum, then breaks up into a network over the canal, the
trunks fusing at the first septum and sending forward a
scries of vessels to the branchie, the returning vessels
uniting to form the ventral trunk below the gut. The walls
of the vessels have a fine epithelial coat and a delicate circular
muscular Jayer with minute nuclei in their spindle-shaped
cells. The author also objects to Gilson’s statement that
no special constrictor to the alimentary canal occurs at the
septa, and points out that at the third septum an efficient
constrictor apparatus exists both for the canal and the blood-
sinus, the muscular apparatus showing the large muscle-cells
at the outer ends of the fibres. Posteriorly also the
alimentary canal is moniliform from its constrictions. He
is inclined to think that Drasche’s bladder-like tissue on the
ventral mesentery is part of the reproductive apparatus.
The ampullz on the ventral blood-vessel, which Drasche
observed to be rythmically contractile, are confined to the
genital region of the body, and bear the reproductive
elements on their outer surfaces, and, though they have
muscular walls, Ziircher would attach more importance to
their nutritive capacity. The red blood contains rounded
or lenticular corpuscles with nuclei, and some corpuscles
undergo mitosis. He found them in the ampulle and less
* Jenaische Zeitschr, fiir Naturwiss. Bd. xlv. pp. 181-220, pls, xv.-xx,
(1909).
Gatty Marine Laboratory, St. Andrews. 239
frequently in the vessels and sinuses. There is a mixture of
venous aud arterial blood in the branchiz, The author
does not touch on the structure of the nervous system of
Owenia, the topography of the alimentary canal, the hypo-
dermic canals of the sixth segment, and other features subse-
quently to be described, and his main points are histological.
The illustrations are chiefly in outline.
No trace of a central nervous system is observed in
Owenia fusiformis till the folds of the mouth are cut in the
transverse sections*, and the first definite appearance of a
layer similar to nerve-tissue is the presence of a pale band
external to the stained basement-layer of the inner border
of a lateral flap of the mouth. It resembles a differentiated
stripe of hypoderm from which cells and pigment are absent,
hut the fine striz are continued tlirough it to the basement-
tissue, the whole being minutely fibrillar like the nerve-
tissue, and generally dotted with minute granules. Ti fades
away before reaching the free or ventral edge of the lateral
flap, and disappears similarly at the dorsal edge of the fold.
Then. (for it is difficult to cut exactly on the same level) a
corresponding band appears on the opposite labial fold.
This pale belt is considered by some, e. g., Atircher, to be
basement-tissue, but it seems to be somewhat different... In
any case, the contrast between it and the condition, for
instance, in the proboscis of the armed Nemertean with its
large strands of nerves and their reticulationus is marked, yet
* Whilst many advantages are gained by the use of paraffin, cellu-
loidine, and other substauces for imbedding, the old plan of fine sections
made directly from careiully prepared spirit-specimens is not without
value in checking the proportional thickness of the muscular layers and
other parts. Thus, in the case of Owenia the great thickness of the
longitudinal muscles of the body-wall can only be appreciated in this
way, and so with the proportional size of the mucous glands and the
tough nature of the basement-layer. In such preparations more than
forty years old the delicacy of the hypodermic! layer has caused most of
it to be removed in the manipulations before and after preservation, but
in every case the nerve-cord firmly adheres to the basement-tissue in the
mid-ventral line, thus demonstrating its comparatively tough nature in
contrast with the hypoderm. In such sections the gut fills the entire
area, with the exception of the mucous glands, though, of course, in life
the coelomic space was larger. The term hypoderm in the structure of
the Polycheta refers to the glandular and granular layer, often areolated,
beneath the cuticle. It is an ectodermic structure.
1 T am indebted to Mr. EK. W. Shann, B.Sc., now Captain in the
Northumberland Fusiliers, Mr. J. W. Pryde, M-A., now Lieutenant in
the Black Watch, and to Miss Harvey, of Ndinburgh, for aid in making
the various sections,
240 Prof. M‘Intosh’s Notes from the
the functions of both are equally well performed. The
Nemertean brain, asin many Polycheets, is distinctly isolated
from the tissues outside it, and the same may be said of the
main trunks in that group. Here, in what is considered to
be a higher series, the opposite condition prevails, the nerve-
centre and main trunks being hypodermal, as are the cords
in the majority of the Polychets. In Owenia this belt
agrees in minute structure with that surrounding the central
system, and occupies a corresponding position.
With the disappearance of a central fillet in the dorsal
arch of the body-wall a slightly pale band is noticeable in
the hypoderm of the region, yet that layer passes to the
basement-tissue (which stains) uninterruptedly, a series of |
the ends of severed fibres being grasped in spaces bounded
by reticulations between the basement-tissue and the ad-
joiming circular muscular fibres. Then the pallor of the
inner portion of the hypoderm becomes more pronounced,
and in the next section or two (PI. VII. fig. 1) a distinct
nervous layer, as in Cephalodiscus, stretches along the mid-
dorsal arch. It shows both fine transverse and vertical fibres
or striz, and minute granules occur next the basement-tissue
(Pl. VII. fig. 2). It fades on each side into the ordinary
cells and areole of the hypoderm, which likewise continues
to the surface externally without evident break. The nerve-
tissue, in short, is marked by no hard-and-fast line from the
hypoderm, but is traversed by its fibres, and the neuropile,
nemoglia, and neurilemma of the ordinary Polychet ganglia
are not distinguishable. From end to end in section the
tissue has a uniform structure, and where, for instance, it is
separated from the basement-layer only projecting vertical
fibres and granules appear. Certain granules occur at its
outer border next the deeply stained cells and granules of
the hypoderm, but these could not be associated with the
nerve-band, the finely fibrillar edge of which coursed evenly
along. Insucceeding sections this great nerve-band stretches
downward at the sides, becomes more distinctly differentiated
from the hypoderm externally and the basement-tissue in-
ternally, and then a slight narrowing of the mid-dorsal arch
takes place, the lateral extensions being thicker. The mouth
is still divided ventrally in these sections, and the nervous
expansion extends over the entire arch of the body-wall with
the exception of a comparatively short region of the ventral
edge of the lateral lip, the thickest layer being lateral, for
the dorsal is now diminishing. A narrow layer, apparently
of basement-tissue, occurs, as indicated, simultaneously in
the sections external to the hypoderm Jining the mouth, and
Gatty Marine Laboratory, St. Andrews. 241
at this level all round, though it is thickest ventro-laterally,
and it was this layer which was first encountered in front.
The structure of this lateral band of so-called basement-
tissue closely resembles that of the central system, and it is
possible that it may perform functions of a sensory kind in
connection with the lateral flaps of the vestibule. The flaps
gradually unite to form the lower half of the vestibule, the
thicker band of pale tissue still being retained ventrally
with a thin connecting-band dorsally. Proceeding a little
backward the main nervous band disappears from the dorsum
and is confined to the lateral regions of the body-wall, from
which it gradually thins off dorsally. Finally, when the
lip-organ appears in the section over the oral gap, a large
nerve-cord alone is left at the lower limit of the former nerve-
band (Pl. VII. fig. 3, me.), all the hypodermic layer dorsad of
it having assumed the usual condition. In sections of Sacco-
cirrus near the mouth Dr. Goedrich * found the lateral cords
(his “ cesophageal commissures”’) ina similar position. This
limited nerve-area presents in section pale, finely granular,
transverse striation, through which delicate fibrils from the
hypoderm external to it pass to the basement-tissue. When
the circle of the body-wall is complete—that is, immediately
behind the oral gap (Pl. VII. fig. 3),—the large nerve-cords
are situated a little below the middle line of the body-wall,
and have a blood-vessel in the muscle to their inner side.
The central region is still lined by hypoderm, and the thick
pale band of the inner layer is infero-laterally conspicuous.
Then the hypodermic layer of the vestibule passes into the
gullet, and sections of the lip-organ (/p.) appear, whilst the
hypodermic inner lining of the dorsal region is shut off by a
deep fold with a narrow layer of hypoderm from the vesti-
bule, the rest of the large arch above having a thick coat of
the same tissue. Externally, again, a change has occurred
in the mid-ventral line, for the thick lateral coat of hypo-
derm in which the nerve-cords lie has thinned off ventrally,
leaving a considerable area with just a trace of it; but this
appears to occur only for a short distance. With the
termination of the vestibule lined by hypoderm, and the
increase of the lip-organ in section, the ventral hypoderm of
the body-wall again gradually thickens from the middle line
outward. Moreover, the narrow pocket formed by the
first septum les on each side of the lip-organ (Pl. VII.
fig. 4), and then is quite shut off from the upper cavity
(vestibule) lined by hypoderm, and which represents the
* Quart, Journ. Mier. Sci. n. 8, vol. xliv.
242 Prof. M‘Intosh’s Notes from the
gullet proper (which may have complex functions), sur-
rounded by a tough (muscular) investment, from which
various strands radiate to the body-wall amongst the blood-
vessels of the region. ‘The body-wall at this part has a
thinner coat of hypoderm both mid-dorsally and mid-
ventrally, its thick layer being lateral. The longitudinal
muscles form a somewhat thin layer of fasciculi ail round,
and the lip-organ shows a thick mass of modified vertical
cells with nuclei, each mass probably rubbing against the
other. The massive lip-organ then forms a thick-walled
tube in section, with a central cavity (Pl. VII. fig. 5) and
an external muscular investment, whilst the cesophagus has
a thick mucous layer, continuous with the hypoderm, to
subserve its special functions, the radiating strands and
numerous blood-vessels still continuing. ‘These radiating
fibres show that the movements of this thick-walled region
(Pl. VIII. fig. 18) must be more or less restricted, yet the
longitudinal bands, especially on its dorsal wall, would point
to protrusion and retraction. Externally the hypoderm— .
thickest laterally at and above the nerve-cords, which are
descending—has increased in depth dorsally, but is thin
ventrally. Behind the foregoing the lip-organ.loses its
central cavity (a fold) and diminishes in size, but its complex
muscular coat is proportionally thicker, and in thesurrounding
area the blood-vessels are larger. Finally, the muscles of
the lip-organ alone are visible, and then disappear, showing
that it is, in short, attached by a muscular stalk, first hollow
and then solid, though the sections would indicate that the
muscular fibres (retractor) are fixed to the body-wall close
behind and for some distance backward. Moreover, a fan-
shaped arrangement (Pl. XII. fig. 17) occurs anteriorly
where the fibres spread into the lip-organ. Besides, various
oblique and transverse fibres act on the folds and give
complexity to the movements (Pl. XI. fig. 16). A double
layer of muscular fibres, further, Jies beneath the basement-
tissue bounding the gland-cells—the one the reverse of the
other,—so that in sagittal section the cut ends of one series
abut on the thick inner (i. e. toward the ccelom) belt.
Gland-cells also occupy considerable areas internally at the
edge of the organ. ‘The whole structure of this organ there-
fore differs from mere labial folds of the vestibule, as more
clearly seen in vertical sections (Pl. VII. fig. 6), the densest
part of the cellular layer being toward the middle of the
ventral fold and thinning off dorsally and laterally. The
lip-organ, in short, is a highly differentiated apparatus, both
secretory and manipulative, for the tube-formation and other
Gaity Marine Laboratory, St. Andrews. 243
functions. It is interesting that Dr. Goodrich * found a
similar organ (his “ muscular pad’’) in Saccocirrus and
Protodrilus. The body-wall at this level again presents a
change in its hypoderm (which throughout has a firm
exterior film or cuticle), since, though somewhat diminished
dorsally, it is now of a considerable thickness mid-ventrally,
its densest part being at the nerve-cords, which have moved
downward, so that they are separated by about a sixth of the
circumference of the body (P]. VIII. fig. 7). The cesophagus
has special fasciculi of muscles laterally and dorsally, besides
the radiating fibres.
The next important change is the merging of the cesopha-
geal region of the canal, with its boldly arranged coat con-
tinuous with the external hypoderm, into the stomachal
region at the third septum, with its granular glandular
surface (Pl. VIII. fig. 7) and its external muscular coat, the
whole internal surface of the stomach being by-and-by thus
transformed.
At the commencement of this region of the gut at the
third septum a complex muscular sheath connected with
the lip-organ occurs ventrally, with thickened muscular
pillars at each side—abutting on a membranous space to
the exterior and just over the nerve-trunks, certain of the
fibres, moreover, a little further back being attached to the
basement-tissue over the outer part of the nerve-cords.
Blood-vessels occur in the large space which is thus soon
formed below the alimentary canal, and the vessel in the
median mesentery (which is attached to the upper border of
the mass of muscle stretching from side to side over the area
above the nerve-cords—a little behind the section figured in
Pl. VIII. fig. 7) is distinct, the special mesenteric area still
being visible externally, though much reduced in size.
Between the basement-tissue of the body-wall and this trans-
verse muscular mass he the ventral longitudinal muscles
(vm.). The great cavity appears to contain coelomic fluid
and corpuscles, and is shut off by a shelf of septal tissue
(Pl. VIII. fig. 7, bt.) continuous at each side with that of
the body-wall, whilst the upper area on each side of the
alimentary canal is occupied by elastic connective-tissue
strands and by the muscular fasciculi along the dorsal wall
of the canal. The body-wall at this region has dorsally the
longitudinal muscles (dm.), which may be held to cease at
the junction of the transverse platform of septal tissue a
little below the middle, of longitudinal muscular fibres
* Quart. Journ, Micr, Sci, n. s., vol. xliv. p. 415, sections 18 & 20.
214 Prof. M‘Intosh’s Notes from the
ending at the junction of the ventral transverse band, and,
lastly, of the median (ventral) longitudinal fibres (vm.)
beneath the latter (Pl. VIII. fig. 7). The nerve-cords at
this part are separated by fully a sixth of the circumference
of the body-wall. The transverse septal plate above the
ventral longitudinal muscles has a central structureless part
—apparently of a homogeneous nature (pale and elastic),
the muscular fasciculi fraying out especially at the dorsal
surface and ends (Pl. VIII. fig. 74). In its progress back-
ward a change in the diminishing area between the nerve-
trunks is inaugurated, the homogeneous central region of
the transverse band, the anterior part of which is indicated
in Pl. VIII. fig. 7, being shortened transversely and increased
vertically, so that it pushes as a lozenge-shaped and then
wedge-shaped area into the centre of tlhe ventral muscular
mass, whilst the upper muscular fibres externally become
defined as more or less independent masses, bounded exter-
nally by sloping muscular fibres which simulate the oblique
in certain sections—at least, at their insertion. The lozenge-
shaped area of the homogeneous (for it can scarcely be called
“tendinous ”) tissue thins off on each side to a plate, to
the upper edge of which fasciculi of muscular fibres are
attached, whilst ventrally processes pass into the median
ventral longitudinal muscles. The whole thus forms a com-
plex muscular apparatus attached to the central tough tissue,
which gradually in its progress backward shrinks, leaving
the fused muscular fasciculi to form the massive ventral
longitudinal muscles as shown in Pl. VIII. fig. 8, vm. By
the gradual diminution of the tough central area of the
before-mentioned transverse band to which the median
mesentery from the gut is attached, and by the grouping of
the several longitudinal ventral muscles into a mass on each
side, the typical ventral longitudinal muscles are formed,
and at this part they exceed in bulk the dorsal muscles.
This evolution of these continuous ventral fasciculi out of
the elements in front is probably connected with a change
in the function of the contents as well as in the body-wall
itself,
The disappearance of the longitudinal muscular fibres and
the radiating strands from the dotsal wall of the gut leaves
the two halves of the upper division of the coelomic space for
ceelomic fluid only, and it is separated from the two much
larger spaces inferiorly by strong muscular bands at each
side of the transversely enlarged alimentary canal; yet the
appearance of the canal beneath—to which the median
mesentery is attached ventrally—apparently leaves a gap by
Gatty Marine Laboratory, St. Andrews. 245
which the two cavities communicate superiorly under the
transversely enlarged canal.
The anterior end of the stomach is a narrow tube as seen
in Pl. XI. fig. 23, and in the various transverse sections. It
further presents a bifid border ventrally, a narrow process of
the cavity ending in a dilated rim on each side below, the
ventral blood-vessel and the mesentery occupying the gap,
whilst a spacious sinus surrounds the stomach. This bifid
condition gradually disappears, the organ assuming the
outline shown in P}. X. fig. 30.
A mesentery with the dorsal blood-vessel in the centre
passes from the upper arch of the gut to the dorsal wall,
and another mesentery, with the ventral blood-vessel, goes
to the mid-ventral tissues, the coelom being thus divided
into halves. Then a process from the wall of the stomach
above the rugose and somewhat triangular ventral arch
appears, and a little behind is tacked to the ventral portion,
and thus cuts it off as a separate canal with folded mucous
membrane internally, the longer upper chamber having its
inner surface smooth and symmetrically folded. The in-
ferior and somewhat pear-shaped chamber (PI. VILL. fig. 8,
st., stomach) is surrounded by blood-vessels, which form a
vascular plexus around it on their way to the branchial
region, and from its apex inferiorly a mesentery passes
to join a mid-ventral homogeneous (pale tough) area arching
‘over a special muscular region which terminates on each
side over the outer edge of the nerve-cord, now approaching
that of the opposite side.
At this level the body-wall has thin hypoderm in the
mid-dorsal line, then it imereases in depth laterally, again
becomes thinner, and then swells out ventrally at the nerve-
cords. Within are the basement-tissue and circular fibres,
then the dorsal longitudinal muscles (dm.), which end below
the attachment of the upper canal on each side, and the
ventral longitudinal (vm.), which are more massive, and
have the differentiated region with the arched fibres in the
middle line, such, indeed, forming the only separation
between them. This differentiated region is probably in
connection with the movements of the alimentary canal.
The nerve-cords in section show a granular and fibrillar
aspect, and they are much better differentiated than in front.
The occurrence of bristle-tufts makes the separation between
the dorsal and the ventral longitudinal muscles more pro-
nounced, and below the tuft is a well-defined pore of the
mucous gland with large nuclei in its cellular wall (PI. VIII.
fig. 9, mp.), one side abutting on the hypoderm, the other
246 Prof. M‘Intosh’s Notes from the
having muscular fibres from the bristle-tuft attached to it;
and the hypoderm is thinned at the tuft and has an incurva-
tion at its upper edge, whilst it rapidly thickens above it.
Moreover, a distinct muscular slip (mec.) occurs in the mid-
ventral line, the remnant of the complex condition in front.
The next change is the infolding of the stomachal wall
(Pl. VIII. fig. 10, s¢.), the loss of its lateral connections, and
the termination of its cavity; whilst the intestine enlarges,
its folds become more prominent and alter their character,
resembling, indeed, the cesophageal hypodermic lining. The
intestine still shows a plexus of vessels, about seven, for
instance, being cut on each side, and they resemble buds
from the investment of the gut, though they are only sections
of longitudinal trunks with their internal and external
investments. ‘The dorsal mesentery and its enclosed vessel
now pass upward from the gut-wall, and inferiorly are the
ventral mesentery and its vessel, the membrane trending to
a fissure between the more massive ventral longitudinal
muscles, since the special median muscular area and its fibres
(shown in Pl, VIII. fig. 7a) have disappeared. The nerve-
cords are separated only by their own breadth from each
other, and they are, perhaps, more distinctly granular than
before. ‘The mucous glands, with their secretion rendered
fibroid by preparation, are now prominent, each placed
above the ventral muscle of its side. ‘The coelomic spaces
(Pl. IX. fig. 11, ¢.), reduced to one on each side, have a
translucent coagulum with granules.
When the nerve-cords touch and fuse (PI. IX. fig. 11, ne.)
it is seen that the glandular tubes in the ccelom approach
each side of the ventral vessel, and slope outward as they go
forward to the excretory duct below the bristle-tuft. The
gut has become pear-shaped, the narrow end being below
with its mesentery, whilst two mesenteries pass from the
dorsal arch and join before reaching the dorsal blood-vessel.
This arrangement makes an additional supra-intestinal
chamber.
‘The hypoderm still presents a symmetrical enlargement
just above the bristle-tuft on each side, this thickened region
being differentiated by the narrow layer immediately above
it, for it gradually deepens dorsally and again becomes
narrow as it reaches the mid-dorsal line. Trom the lower
edge of the bristle-tuft it gently increases to the nerve-cords
in the mid-ventral line. The dorsal longitudinal muscles
are thinner than the massive ventral, but they extend over
a larger area of the body-wall.
A little further back (Pl. IX. fig. 12) the gut increases in
Gatty Marine Laboratory, St. Andrews. 247
vertical diameter, forming a long flattened organ extending
from the dorsal to the ventral region in section, and the
cellular lining is thrown intoruge. Both dorsal and ventral
short mesenteries are double, and the wall of the dorsal
vessel is more muscular. The gut has the same vascular
investment as just meutioned, and the vessels lie outside the
mesentery just alluded to. At the next bristle-bundle a
pore similar to the first gives issue to the secretion of the
second pair of glandular tubes. The ventral longitudinal
muscles retain their more massive outline,
After an interval the body (Pl. IX. fig. 13) increases in bulk
proportionally, but the hypoderm becomes thinner all round,
the thickest part being that situated ventrally on each side of
the nerve-cords. The ventral longitudinal muscles pass far
upward, and encroach on the dorsal, which occupy only the
upper arch of the body, and each pair has a distinct median
notch into which the mesentery fits, the ventral mesentery
having the ovaries or spermaries attached to it laterally,
aud the contents of which are shed into the celom. Two
mucous glands in section occupy the upper half of the
coelom on each side, and the nuclei of the cells forming their
walls are regularly arranged. The gut stretches nearly from
dorsal to ventral arch, held in position by the median
mesenteries and also by the septa at intervals. The double
attachment superiorly forms a blood-channel, which com-
municates with a sinus (séz.) surrounding the gut, so that
here, instead of the isolated though reticulated vessels, there
is a continuous blood-channel—a development in all proba-
bility attained only in the adult or nearly adult condition.
The nerve-cord still has numerous hypodermic fibres passing
from the outer to the inner (ventral to dorsal) or vice versa,
and, in addition, fine reticulations and granules, some of
which are probably nuclei. Usually a slight ventral furrow
and a median peak dorsally indicate the double nature of the
area. The second pair of glandular tubes is situated to the
exterior of the first pair in transverse sections.
The sixth segment is distinguished by the presence of
Gilson’s epidermal tubes, which stretch from the septum
between the sixth and seventh segments to that in front.
‘They are readily recognized ‘by their position in transverse
sections, viz., dorsad of the groove (Gilson’s “ gouttiére de
la soie’”’) which runs along the dorso-lateral region. They
are canals of considerable size, and are separated from the
basement-layer by a stratum of cells, the cavity in section
being also bounded externally by an arch of hypodermic
cells, Gilson supposed that these hypodermic canals served
248 Prof. M‘Intosl’s Notes from the
for the transmission of the reproductive elements, and
possibly also for an interchange between the coelomic cavity
and the exterior as in certain Oligochets. True nephridia,
at any rate, are absent in Owenia.’ Whatever the function of
these canals may be, Arnold Watson has shown that sperms
and ova escape by different channels.
In front of the tail the hypodermic coating of the surface
is of moderate thickness. ‘The massive muscular investment
is conspicuous, and it is difficult to distinguish where the
dorsal longitudinal muscle ends and the ventral begins,
though a fold above the mucous gland seems to indicate the
separation. The nerve-cords have shrunk to a small lenti-
cular area, which in minute structure has the same fibrillar
and granular character as in front. The intestine, held in
position by a dorsal and a ventral mesentery, is considerably
less, but it has large vessels or sinuses on each side, the
ampullee from the ventral vessel passing into the large gonad
below the gut. Two mucous glands are still in evidence
under the dorsal wall, and they have the same character as
in front. Moreover, their ducts open above the long line
ef hooks in the space between these and the bristle-tuft,
which is now dorsal in position, and so leave the entire
Jateral wall to the hooks; thus the restricted area occupied
by the dorsal longitudinal muscles is defined. The whole
lateral and ventral regions are covered by the ventral longi-
tudinal muscles, which, however, are much thinner than the
dorsal, the reverse of the Eatidiaen in front. The con-
spicuous development of the gonads in this region and the
ampulle of the ventral vessel are noteworthy. The mucous
glands have now ceased (PI, IX. fig. 14).
One of the interesting features toward the tail is the
occurrence of the septa (P1. IX. fig. 15). Their first appear-
ance is indicated by the envelopment of the intestine and
its blood-sinuses by a sheath which springs from each side
of the vertical mesentery under the dorsal blood-vessel, and
stretches to the mid-ventral mesentery considerably below
the ventral blood-vessel. In such a view it might be sup-
posed that the middle of the septum has been sliced, leaving
the upper and lower attachments ; but such will not explain
all the outlines of these posterior septa.
Reproductive elements occur in the spaces outside the
septum as well as within it and its areas. Then the upper
and lower arches separate, each having a zigzag outline as
it passes to the body-wall. The ceelom is thus divided into
six areas—two dorsal, two ventral, and the lateral with the
gonads inferiorly on each side of the gut.
Gatty Marine Laboratory, St. Andrews. 249
The first appearance of the septum in the sections is
heralded by a tuft of muscular fibres attached to the exterior
of the gut-wall and the ventral septum; then the muscular
ring (the ventral septum and the gut being free) loosely en-
velops the gut and its vessels, besides the ventral blood-vessel
and its mesentery, almost to the ventral longitudinal muscles,
but leaves the dorsal blood-vessel for the most part free.
The septum, indeed, springs on each side from the lower
wall of the vessel, and encloses that part of the mesentery
between the vessel and the sinus around the gut, whilst the
distal part of the mesentery passes freely to the gap between
the dorsal longitudinal muscles. This muscular sheath or
tubular chamber by-and-by swells out into a large area, its
upper arch or roof being attached on each side to the body-
wall between the dorsal and ventral longitudinal muscles,
and its floor stretches from the median ventral mesentery
to the wall of the body a considerable distance below the
attachment of the roof. This chamber encloses thealimentary
apparatus and the gonads, but the more advanced sperms
lie in the two ceelomie chambers outside its roof—that is,
between the latter and the body-wall (Pl. IX. fig. 15). It
is difficult to explain the exact nature of these septa—
whether they are modifications of the ordinary septa, which
extend far backward in the caudal region, or only the
ordinary septa sliced so as to present these characteristic
appearances,—for it is unlikely that two septa would fall
into the line of section. These septa seem to differ in
disposition and aspect from those in frout, and are probably
associated with the special functions of the caudal region—
respiratory, purely intestinal, or otherwise.
On viewing the animal externally from the dorsum, a
broad fillet passes from each side of the collar anteriorly and
slopes obliquely inward and backward on the dorsum to the
constriction behind the third bristle-tuft, then bends a little
outward, and is continued along the dorso-lateral region
posteriorly. A groove exists at the collar just below the
anterior end, and which apparently is functional also for the
median ventral ridge and groove, so that, if ciliated, it may
send a current outward and forward to it. These ridges
apparently are those which show the remarkable pennate
arrangements in the hypoderm in the preparations (PI. IX.
fig. 20)..
In certain longitudinal sections (Pl. VIII. fig. 18) the dark
pigment stretches as a broad band behind the collar, a gap
intervening between it and the edge of the fold behind, such
probably representing a sensory groove, and its borders have
Ann. & Mag. N. Hist. Ser. 8. Vol. xix, 17
230 Prof. M‘Intosh’s Notes from the
the specially modified cells. Von Drasche* figures’ the
pigment only behind the collar, and his “ ganglion ”’ is small
and considerably in front of the collar, thus diverging from
the condition described here. Ziircher does not deal with
this region.
In longitudinal sections a marked feature is the pennate
condition of the hypoderm of the anterior region—that is,
between the first and second dissepiments, as well as a little
in front of the former in certain cases ; and it appears to be
specially developed on the dorsum. The hypoderm is there
thrown into a series of ridges, which in section present a
streaked granular basal region terminating externally in
a pennate and symmetrical series of small granular cells,
after the manner of the barbs of a feather (Pl. IX. fig. 20),
the breadth of the ridge varying, whilst the processes (in
section) increase in size from behind forward, culminating
in the collar with the deep groove in front (Pl. XL fig. 16, and
Pl. VILL. fig. 18). Some of the ridges, springing from the
continuous base, are narrow distally, so that the lateral rows
of cells are close on the midrib and a few are more or less
conical. The transition from the hypoderm of the succeeding
segment is by a gradual modification in the arrangement
of the vertical cells, which by-and-by are fan-like and then
pennate. This pennate condition in the preparations of
the hypoderm is apparently limited in distribution, since it is
absent in most sections both dorsally and ventrally, and Von
Drasche neither mentions nor figures it. Moreover, in trans-
verse sections, so far as observed, it is not seen, and therefore
may be due to the arrangement of the cells in a vertical plane
after preparation. An-approach to this condition of the
hypodermic cells is observed in some longitudinal sections
of Myzicola, but it is less distinct than in Owenia, and is
probably due likewise to the effect of the preservative fluid
acting on a thick glandular hypoderm. ‘The inner edge of
the collar has a series of minute cells along its anterior
border, and a fan-like series of strands and cells posteriorly,
whilst the tip is symmetrically pennate. The anterior curve
of the furrow is furnished with a special series of granular
pigmented cells, oc., closely arranged at the surface, and
which probably have the functions of eyes. They extend
the whole length of the collar from side to side on the
dorsal surface, and are partly protected by that fold (Pl. VIII.
fig. 18, and Pl. XI. fig. 16).
The mucous glands (mg.) present either a characteristic
* Op. eit. Taf. i. fig. 3.
Gatty Marine Laboratory, St. Andrews. 258
pennate aspect in longitudinal section or a series of straight
or curved transverse bars, according as the long tube is cut
in a median or lateral plane (PI. X. fig. 19). In the former
case a central axis of the secretion is flanked on either side
by a series of plates, often slanting distally, and containing
an occasional nucleated cell or a series of granules in the
plate. Such a condition may be due to the action of the
preservative spirit, or to the method of secretion, but it is
worthy of note. The slender posterior ends of these glands
are curved forward and outward. The secretion forms a
lining to the tube, and attaches foreign structures such as
sand-particles and foraminifera to it externally ; and in
many cases, so firmly do the annelids adhere to it after
preservation, that rupture of the tissues accompanies their
removal.
The hypoderm covering the mouth and buccal region,
including the “lip-organ,” differs from that on the surface
of the body and branchie. It is bounded bya uniform and
definite investment, and has a finely-granular and fibrillar
structure, so that it forms a tougher, more massive, and
more consistent layer, which, however, at certain parts
diminishes in thickness as it approaches the branchie. It
rests on a basement-layer having beneath it a complex series
of muscular fibres. The same kind of hypoderm dips down
and envelops the lip-organ, though it is more translucent
in section, from the paucity of granules which stain more
deeply. Then the organ forms a deep furrow (Pl. VII. fig. 6)
with massive pale walls, whilst a double fold which now
appears to the inner side, as well as the folds dorsad of the
mouth, stain distinctly, as also do all the folds of the mouth
and pharynx. ‘The pale region thus lies in the figure between
a and a in the centre of the organ, but it thins off on each
side—that is to say, the middle region of the fold has thickest
walls. The buccal mucous membrane is like that first men-
tioned in the lateral area of the cephalic region, viz., closely
fibrous and granular, and it continues to the second dia-
phragm. It rests on a basement-membrane and a firm
outer layer of both circular and longitudinal muscular fibres,
the anterior or buccal region having numerous trabecule
fixing it to the body-wall ; and this is specially marked at
the thick pale folds (lip-organ, /p., in the various figures).
The anterior buccal region is probably capable of partial
protrusion. In front of the second diaphragm the folds of
the canal have thick muscular walls, so that a certain amount
of differentiation exists—either as proventriculus or stomach.
Behind the second dissepiment the walls of the canal are
Liz
252 Prof. M'Intosh’s Notes from the
apparently uniform, and contain mud rich in organic
remains.
The former or stomachal region enters the following or
intestinal region by an aperture which is thrust backward as
acone. Moreover, the walls of the organ undergo a struc-
tural differentiation, for a short distance before reaching the
aperture (text-fig.) they become finely reticulated and dotted
as if formed of muscular or erectile tissue (ec.), whilst the
lining of the tube consists of the same mucous membrane
as in front. Further, the adjoining circular fold of the
diaphragm (sp¢.3) is provided with a similar, though
thinner, layer of the same tissue (ec.), which likewise
Longitudinal section of the alimentary apparatus at the third septum,
spt. 3, showing traces of the special muscular layer, ec., enveloping
the posterior wall of the stomach and its sinuses, and continued
over the valvular region, val.; hp., hypoderm; cel.c., coelomic
corpuscles ; mg., mucous glands.
occurs in two of the folds of the organ in front of the
foregoing. This tissue is apparently muscular, and its
minute structure is interesting as showing the peculiar
muscle-cells with their granular contents and nuclei which
stud the free border of the muscle. Zircher * has given a
good description and figures of the structure of this tissue.
It evidently controls not only the wall of the canal but the
blood-sinus on its outer surface, and thus may have con-
siderable effect on the main trunks proceeding forward
to the branchiw. The ccelomic corpuscles often form a
_ * Op. eit. p, 208, and figs. 33 & 35.
Gatty Marine Laboratory, St. Andrews. 253
translucent mass (cwl.c.) behind the diaphragm and on
each side of thejgut.
Myriochele, the second genus of the Ammocharide, differs
from Owenia in the simplicity of its anterior end, for it is
broadly truncated, with a smooth margin, a deep pit or
vestibule leading to the mouth, and a ventral fissure; yet it
is found that, though the branchiz are absent, the nervous
system is formed much on the same plan as that of Owenia.
Myriochele is even more broadly truncate than Owenia, and
as a transparent object its large blood-vessels in front
and the great longitudinal mucous glands which follow
give the body a striped aspect.
The cilia covering the large funnel-shaped oral cavity of
Myriochele are long and powerful, so that the currents they
cause are probably considerable. Moreover, the oral aperture
is oblique, the rim dipping backward to the notch at the
ventral border, thus somewhat resembling the condition in
the young of Owenia, though the oral gap is larger.
In the early sections of the snout of Myriochele the
hypoderm presents more distinct elements thau in Owenia,
and it appears to be somewhat thicker, its minute cells and
‘ granules in the gelatinous matrix being conspicuous. The
sections have a horseshoe-shaped appearance, the wide oral
gap beneath forming the heel of the shoe, which, however,
is tapered at the tip, the wall thinning off at each side. The
exterior of the shoe has cuticle, glandular hypoderm, and
basement-tissue resting on a gelatinoid layer which has
numerous minute nucleated cells along both outer and
inner borders. Basement-tissue, again, bounds the vesti-
bular hypoderm on the inner border, which differs from that
of the outer wall in having a distinct inner coat, from which
cilia probably spring. The basement-layer in both cases
is apparently elastic. The surface-layer of hypoderm
often presents clear spaces or vacuoles—probably from
rupture and extrusion of the glandular tissue. Moreover,
its external surface forms a more definite cuticle, whilst its
inner border rests on the basement-layer, no nervous belt
appearing in the first sections; but circular fibres occur
within the basement-layer, and then a well-developed longi-
tudinal coat of muscle which stretches downward to the
oral edge, from which the epithelium of the mouth passes
inward (PI. XI. fig. 22) as a thick layer of cylindrical cells
with nuclei, bounded internally by a thin sheet of circular
fibres and a few longitudinal strands. The space (ccelom)
between the body-wall and the oral wall shows many
254 Prof. M‘Intosh’s Notes from the
granular cells with fine connective-tissue fibres at certain
parts, besides blood-vessels. Then a narrow pale belt
becomes distinct within the circular fibres and basement-
tissue of the oral wall, apparently corresponding to the
pale sensory layer of Owenia in the same region. The
nerve-centre appears as a narrow pale granular band in
section at the inner border of the hypoderm of the body-
wall, and stretching downward from the dorsum (PI. XI.
fig. 22) as it passes by-and-by into the trunks connecting
it with the ventral cord. This region therefore represents
the prostomium, though devoid of any external indication.
The minuteness of the nerve-centre in comparison with that
of Owenia renders its finer details obscure, and it is more
transparent. No fine strands from the hypoderm could be
made out, the slightly prominent cells and interstitial tissue
alone appearing at the edge, whilst its inner border rested
on a smooth basement-tissue. Its position and extent
agree with that in Owenia.
In horizontal (longitudinal) sections the central nervous
system appears as an area at the inner border of the hypo-
derm about the point of the V-shaped oral funnel (Pl. XI.
fig. 23), and its transverse breadth is shown by its appearing
on each side in these sections. So far as can be ascertained
in the preparations, no special sensory apparatus is present
either in the form of a groove or deposit of pigment in the
body-wall, but the pigment may have been removed by
long preservation in spirit. Therein it differs from Owenia
with its pigmented cells and its groove.
Then, the hypoderm, again, extends over the whole depth
to the basement-layer dorsally, and the nerve-cords are
differentiated laterally—at first high up, nearly on a level
with the dorsal arch of the mouth (Pl. XI. fig. 25), and
then gradually descending as in Owenia. Very soon
between the mid-dorsal and the oral walls a blood-vessel
appears, and one in each lateral space, the counective-tissue
strands and cells which connect the walls apparently
keeping them more or less in position, the vessels being
proportionally large for the size of the annelid and perhaps
subserving respiration (P]. XI. fig. 25 and Pl. XII. fig. 24,
bv.). When the body-wall becomes continuous—that is,
just behind the ventral (oral) slit—the cords have reached
the commencement of the lower third of the body-wall, and
the median arch dorsally and the mid-lateral regions of the
gullet present the thickest layer of cells, the upper angles
and the lower edges being thinner. Moreover, a section of
Gatty Marine Laboratory, St. Andrews. 255
the lip-organ (Jp.) appears. A pale band indicates a differ-
entiation outside the cellular layer dorsally and another
laterally. The chamber by-and-by assumes a figure-of-8
outline, the section of the lip-organ occurring in the lower
division; and this shows a dorsal lenticular.region of firm
pale nucleated cells, somewhat symmetrically arranged, the
lower part still having its cavity surrounded by the softer
and more deeply-stained nucleated cells of the vestibule
(Pl. XI. fig. 26). The figure-of-8 outline of the chamber
is now complicated by a median process on each side and by
the appearance of a diverticulum (gullet) dorsally, whilst the
increase in its size diminishes the space between it and the
body-wall laterally and superiorly, though from the first it
clings to that wall ventrally. The pale streaked dorsal region
of the lip-organ is gradually increasing in size as the sections
pass backward ; the diverticulum joins the upper region of
the canal, which is soon separated from the lower by the
junction of the median processes or isthmuses, thus con-
fining the lip-organ to a special chamber (PI. XII. fig. 27).
Theeupper chamber is lined by the soft cellular mucous
coat; the lower has a thin lining of epithelium, with longi-
tudinal and a few circular muscular fibres externally, the
whole becoming continuous with the upper edge of the lip-
organ on each side, the remains of the ventral wall with its
mucous lining at first linking it to the lip-organ and theu
disappearing, the mid-ventral region being occupied by
strong muscular fibres, probably the protractor of the
organ. The nerve-cords are on each side and widely
separated, and the ventral wall of the body is very thin.
A blood-vessel lies on each side at the upper edge of the
lower chamber; a section of a succeeding part of the canal
appears at the upper border of the wall of the lower chamber,
aud soon stretches across the entire region. A change
is also taking place in the upper chamber, the lower region
of which is thickened and its cells rendered paler. In the
roof of the lower chamber the cells are assuming the chordoid
condition of those in the lower region, so that very soon both
halves make an efficient organ for compression or manipu-
lation. The upper chamber becomes also smaller, and the
space between the two larger. Strong muscular fascicules
appear both dorsally and ventrally over the lingual organ in
the lower chamber; and the nuclei in the elongated cells of
the modified organ form a row nearer the outer than the
inner border. In the interval between the upper and lower
chambers another diverticulum of the upper chamber has
256 Prof. M‘Intosh’s Notes from the
been intruding, and is easily recognized by the numerous
nuclei. The halves of the lip-organ in the lower chamber
are becoming continuous and, fusing, form a dense cylinder
with enly a chink in the centre, the nuclei of the cells being
situated near the external border. The alimentary canal in
the upper region occupies more than half the area within the
body-wall. At this level the hypoderm is thickest laterally
and has increased ventrally with the downward progress of
the nerve-cords, the lip-organ is now solid in section and
smaller, whilst the canal above has increased in size, and
a central chamber of different cellular structure makes its
appearance, whilst by-and-by only muscular fibres occupy
the place of the lip-organ inferiorly, and a cselomic space
occurs at each side. The nerve-cords touch and soon fuse.
The alimentary canal now takes a median position in
the body-cavity, with a dorsal and a ventral mesentery,
and it occupies a large space. The thinning of the dorsal
longitudinal muscles in the mid-lateral region indicates a
differentiation, whilst a considerable mass over the ventral
nerve-cord and a thinner layer on each side represent the
longitudina] ventral muscles. The mucous glands now
appear toward the lower region of the ceelom, and they
seem to have the same structure (Pl. XII. fig. 28) as in
Owenia, and to open by similar wide ducts. The dorsal
and ventral blood-vessels in the respective mesenteries are
large. The hypoderm at this level is thick all round, espe-
cially ventrally, where the nerve-cords are in juxtaposition,
and the dorsal and ventral longitudinal muscles are thicker
and better differentiated (Pl. XII. fig. 29). The alimentary
canal next presents various wrinkles; and a pale band passes
from each side of the nerve-cords (which are proportionally
large) outside the basement-layer, as if extension were
indicated. The alimentary canal is thrown into deep folds,
as if a stomachal or gizzard-like part existed in the lateral
regions, whilst the dorsal and ventral arches have the
ordinary mucous structure; then considerable vertical con-
striction occurs, the dorsal and ventral arches disappear,
anda vascular sinus is established laterally. Thereafter the
gut is pointed above and is split into two lobes ventrally,
with muscular bundles in the gap. The latter (gap) by-and-by
disappears, the canal enlarges, the lumen is much filled up
with the dense coating of cells, and the vascular sinus around
it is continuous in the sections (Pl. X. fig. 80). Externally
at this level the hypoderm is massive ventrally, thins off
laterally, and again becomes thin dorsally. The longitudinal
muscular coat is thinner, and fibres radiate from a little
Gatty Marine Laboratory, St. Andrews. 257
above the duct of a mucous gland, impinging on the wall of
the blood-sinus on one side, and apparently attached to the
depression in the hypoderm and indicative of an opening in
the mid-lateral region. ‘The epithelium in the food-canal
differs quite from that in front, being almost fibro-granular
in aspect from the elongation of the cells, the nuclei of which
lie toward the outer border, and at first the surface presents
reticulations. The alimentary canal enlarges behind the
foregoing, but the character of the mucous lining remains
the same, and the ventral blood-vessel lies in the mesentery
over the nerve-cord, which is large, with a median peak
dorsally and a specially thickened hypodermic layer at each
side. On the dorso-lateral regions of the body the hypoderm
appears to be thicker than on the mid-dorsal and mid-lateral
parts. No specialized dorsal blood-vessel appears in this
region, for it has fused with the sinus.
In horizontal longitudinal section (Pl. XI. fig. 23) the
alimentary apparatus has a similar appearance at the second
septum to that shown in Owenia, though the details are
slightly different and the scale is much less. The mucous
lining is thickened as it approaches the septum, and a
centro-lateral fold bulges forward into the stomachal or
gizzard-like division, whilst the central opening is narrowed
and enters the succeeding part of the alimentary canal as a
prominent papiila. The narrow termination of this region
stains-more deeply than the rest; indeed, it is coloured like
the muscular septum, so that it is probable that it has
specially developed muscular fibres at this part so as to
enable it to perform sphincter-like functions, the food being
retained in this chamber for some time and then permitted
to pass backward by relaxation of the muscular guard.
In vertical sections of the middle region the gut is at first
flask-shaped, the wide part, with the contents, being dorsal,
the narrow part ventral ; and the mucous lining has again
altered its character, the cells being larger, their nuclei
larger, and the inner edge smooth. The ccelomic space in
the female is distended on both sides with large ova having
a clear nucleus, an opaque nucleolus, and granular contents,
and they spring from the epithelium of the mesentery below
the ventral blood-vessel, the smaller ova being inferior, the
larger superior. A noteworthy change is the disappearance
of the blood-sinus around the gut and the presence of a
dorsal vessel in the upper mesentery, the ventral trunk
remaining as before. The thick layer of hypoderm ventrally
has a furrow over the median nerve-cord, and this coat is
comparatively thin laterally and dorsally. The dorsal and
258 Prof. M‘Intosh’s Notes from the
ventral mesenteries separate the muscles of the sides, but
there is little to distinguish between the areas of the dorsal
and the ventral muscles respectively.
Proceeding backward the epithelium of the gut becomes
somewhat finer, the longitudinal muscles form a uniform
layer all round without evident differentiation other than
the attachment of the median mesenteries, and the hypoderm
still remains thicker ventrally, whilst the blood-vessels and
ova show no change. A gelatinoid (protoplasmic) layer
envelops the longitudinal muscles internally, the repre-
sentative of the coelomic epithelial layer.
Then the intestine in section shows a keel ventrally, and
septal strands, apparently muscular, pass from it to the
body-wall, making membranous (and partly muscular) septa
to the latter, viz., one on each side of the gut. These septa,
however, soon reach the dorsal region and become attached
to a process from the dorsal wall of the gut on each side,
the dilated (globular) region of the canal being filled with
food below these, the ventral portion forming a solid apex.
The septa tend to mount upward on the interior of the
body-wall, leaving two great areas ventrally on each side of
the canal and a narrow dorsal chamber above the septa.
Finally, the septum disappears, leaving only a small vessel
at the dorsal mesentery. Then a sinus again forms on the
upper wall of the gut, the ova continue as in front, and
the section of the nerve-cord is more or less circular.
Behind the former region the body-wall becomes somewhat
thinner, the thickest region of the areolated hypoderm being
the ventral. The nerve-area is comparatively large and ovoid.
The basement-layer and, it may be, fine circular fibres occur
internally, whilst the longitudinal muscular fibres are only
differentiated by the median mesenteries dorsally and ven-
trally. The gut is large, with a firm external wall and a
single layer of cylindrical epithelium, the nuclei being
symmetrically arranged in the middle. No dorsal vessel
is visible, but the frilled external wall of a sinus occurs
laterally on the intestine. The ventral blood-vessel is large
and the mesentery leading to the ventral wall is loaded on
each side with developing ova, the larger forms distending
the coelomic cavity on each side (PI. IX. fig. 31).
The tip of the tail is bilobed, with, in addition, a ventral
median semicircular lobe, and is richly ciliated (Pl. X. fig. 32),
a short terminal portion of the intestine being straight, the
next (in front) being indicated by a slight constriction, whilst
the third is almost elliptical, from marked constrictions
in front aud rear. In most cases, when removed from the
Gatty Marine Laboratory, St. Andrews. 259
tubes after preservation, the caudal region is thrown into
various zig-zags or spirals, and the tufts of bristles are
more conspicuous than infront. One or two of the terminal
bristles are single on each side, those preceding being in
groups of two, three, four, and five or more. In certain
examples, in lateral view, the ventral process at the tip
of the tail projects more than the dorsal, though, perhaps,
irregularity of the dorsal lobes occasionally occurs. The
arrangement of the septa is apparently on a similar plan
to that in Owenia, where they are very distinct. The
septa in Myriochele probably cause the constrictions, and
the tips of the lozenge-shaped sections of the intestine are
fixed by membranous attachments to the body-wall. A
pinnate aspect is apparently due to the blood collecting at
the septa, where it was darkened by the stain (hematoxylin).
The reproductive elements appear to be lodged on each side
posteriorly, but their mode of exit has not been demon-
strated. So far as could be made out, no pores were present
posteriorly.
In sagittal sections of the tail (Pl. X. fig. 33) the
constriction present a short distance from the tip appears
to be normal, the gut being narrowed at this point and
furnished with a valvular process projecting forward (val.p.).
The terminal region of the body thus marked off is divided
dorsally into five compartments by short transverse septa,
on the anterior faces of which layers of blood occur, probably
from extravasation, as no walls other than the septa are
visible. The continuation of comparatively large intestinal
sinuses almost to the tip of the tail, in addition to the ventral
trunk, indicates their importance. in the economy of the
annelid, probably in connection with a respiratory function.
The transverse sections of the extreme tip of the tail show
a ring of hypoderm with a ventral gap (Pl. X. fig. 84), on
each side of which the wall is thickened, so that it is lobate.
Moreover, a differentiation occurs in this lobate part, as if an
aperture existed; but such may be due simply to the more
vacuolated condition of the hypoderm of this region. The
cells are larger than on the dorsal surface, and after the
completion of the posterior aperture they form a distinct
pale area on each side of the middle line (Pl. X. fig. 35).
The gut shows, almost before its closure, traces of the vascular
sinus (Pl. X. fig. 35, vs.) on each side, the blood in which had
been rendered of a deep purplish-black hue by the action of
heematoxylin ; and soon the ventral vessel appears, the lateral
sinuses greatly increase in size, whilst ova are present between
the ventral vessel and the body-wall, the nuclei in these being
260 Prof. M‘Intosh’s Notes from the
deeply stained. The lateral sinuses leave only a small portion
of the gut bare above the ventral vessel in front of the fore-
going sections, and the ova occupy the lateral regions, though
their position is variable, for they by-and-by appear, as the
body enlarges, below the alimentary canal and the vessels.
The fine sand-tubes of this form abound in such regions as
the Gulf of St. Lawrence, and occasionally one end (the
caudal) is terminated by a long tapering filament of the
secretion covered with sand.
The main feature in forms like the present is the partial
differentiation of the nerve-tissue from the hypoderm with
which it is in continuity at its centre. Nosheath is evident
anywhere, even in the more distinctly outlined nerve-cords
posteriorly. Yet the position of the cephalic centre and its
connection by two trunks with the ventral nerve-cord agree
with the general type. The innervation of the alimentary
canal seems to be carried out on a similar plan to that of the
main system, viz., by contact with a sensitive layer rather
than by special twigs, since the latter have not been met with
in sections. The whole nervous apparatus, indeed, is m an
elementary condition, and in marked contrast, for instance,
with that of such highly differentiated types as Bispira and
Branchionma, where a chordoid skeleton protects the central
ganglia and the neuroglia is much developed, the whole
central system being shielded by the tissues around it, and
so in the brain of Glycera as described by Gravier*, in
various types by Eisig, and in the brain of Lagis as shown
by Nilsson +. lu Owenia and Myriochele the trunks from
the ceutral system are not cesophageal, but run externally
in the hypoderm to join the ventral cord. Both Owenia and
Myriochele appear to have certain larval characters, as seen
in the young of various polycheets, for instance, in Kleinen-
berg’s t Lopadorhynchus, in which, amongst other features,
the nervous system of the gullet may approach that of the
enigmatical pale layer in the vestibule of the present species.
The structure of Saccocirrus, as given by Goodrich §, also
presents certain analogous conditions.
The alimentary canal of both QOwenta and Myriochele
shows certain valvular complexities, doubtless associated
with the nature of their food—viz., mad or sandy mud con-
taining organic particles of various kinds. Carried into the
* Bull. Se. France et Belg. t. xxxi. p. 159,
+ ‘ Beitrige der Kennt. des Nervensystems der Polycheeten,’ Upsala,
1912.
t ‘Die Enstechung des Annel. aus der Larva von Lopadorhynchus,’
1886.
§ Op. cit.
Gatty Marine Laboratory, St. Andrews. 261
vestibule by ciliary currents, it would in the first instance
be subjected to the action of the oesophagus, then passed in
certain quantities into the stomach with its mobile glandular
walls, and subsequently sent through the funnel-like muscular
valve at the third septum into the intestine. The occurrence,
moreover, of the valve-like folds towards the posterior end
of the intestine in Myriochele, with the adjacent vascular
apparatus, would seem to indicate special functions there,
both in regard to the contents of the gut and respiration.
The central nervous system in Owenia and Myriochele
does not conform to the typical three regions of the able
investigator Racovitza—viz., the paired “ région palpaire”
giving branches to the palpi, the unpaired “ sincipital”
giving branches to the eyes and tentacles, and the paired
“nuchal” to the ciliated sensory- grooves; or to those
of other authors of more recent date, the elementary con-
dition, perhaps, being associated with the feebly developed
and much modified prostomium, especially in Myriochele.
Further, the contrast between the typical form with its
circumecesophageal commissures is noteworthy, since the
homologues of these are as much hypodermal as the central
mass, for the nervous layer beneath the hypoderm of the
vestibule essentially differs.
Another feature of moment is the absence of distinct
nephridia in both Owenia and Myriochele, the only repre-
sentative of a tube communicating with the coelom and the
exterior being Gilson’s long hypodermic tube in the sixth
segment, and which apparently is indicated in Delle Chiaje’s*
original figure as two zig-zag tubes bet ween two bristle-tufts;
and the author was also acquainted with the long mucous
glands and the general arrangement of the branchiz and
their blood-supply. The addition of two eyes to one of the
figures (2), with a pair of pinnate branchie, is, however,
more or less imaginary.
Again, the general structure of Owenia and Myriochele, as
representing thefamily Ammocharidz, gives small grounds for
their association under the same suborder, as Prof. Benham
in his earlier classifications+ seemed to think, with the
Spionide and Chetopteride in his group Spionoformia,
which really comprehends these only, since his Polydoridz
and Magelonide, of which separate families are made, can
without undue laxity be placed under the Spionide. Le-
vinsen, indeed, had previously made a separate group for
* Diseriz. e Nat. degli Anim. Invert. pl. 175. figs. 1-5 (1841),
+ ‘The Cambridge Natural History,’ vol. ii. p. 258 (1896).
262 Prof. M‘Intosh’s Notes from the
his ‘‘Ammochariformia,” keeping under his “ Syllidiformia
Spionina” the Spionide, Chetopteride, Cirratulide, Ari-
ciide, Chlorcemide (?), and Opheliide, an assemblage even
more complex than that of Prof. Benham.
Explanation of the letters used in the figures.
bv. Blood-vessels.
e. Central nervous system.
em. Circular muscular coat.
ce, Coelom.
cel.c. Coelomic corpuscles.
dm. Dorsal longitudinal muscles.
dv. Dorsal blood-vessel.
ec. Special layer of gut and diaphragm,
g. Gonads.
Ap. Hypoderm.
Ape. Thickened layer of hypoderm.
mt. Intestine.
lp. Lip-organ.
m. Mouth.
mg. Mucous glands.
mp. Pore of mucous gland.
ne. Nerve-cord.
es, CEsophagus.
ov. Ova.
spt. Septum.
st. Stomach.
val.p. Valvular process of alimentary canal.
vm. Ventral longitudinal muscles,
vs. Vascular sinus.
vv. Blood-vessels.
EXPLANATION OF THE PLATES *.
Puate VII.
Fig. 1. Transverse section of the anterior end of Owenta fusiformis, Della
Chiaje, to show the central nervous system (¢.). m., mouth,
the lining of hypoderm also having a pale band beneath it at
mc. The ventral gap in the body-wallisstill open. x Zeiss
oc. 2, obj. A.
Fig. 2. More highly magnified view of the central nervous system, c., as
part of the hypoderm, hp., em. Circular muscular coat.
x oc. 4, obj. D.
Fig. 38. Transverse section after the completion ofthe body-wall in front
and the appearance of the lip-organ jp. e., nerve-cord.
X oc. 2, obj. A.
Fig. 4. Section behind the foregoing. The vestibule is now contracting,
and the lip-organ, /p., is in full development, with its inner
ply region and more massive external part. xX oc. 2,
obj. A.
* T am indebted to the Carnegie Trust for part of these figures.
Gatty Marine Laboratory, St. Andrews. 263
Fig. 5. The wall of the esophagus, es., as the sections proceed backward,
is completed, and various blood-vessels, vv., including the
dorsal trunk, dv., are prominent. The lip-organ, /p., presents
a basal mass with a central chink, and a regular arrangement
of its tissue. X oc. 2, obj. A.
Fig. 6. Vertical section of Owenia fusiformis. a., the vestibule ; ¢., cen-
tral nervous system ; dp., lip-organ ; mg., mucous gland ; spt.,
septum. The section is imperfect at 2. X about 40 diam.
PuaTeE VIII.
Fig. 7. Transverse section behind the folds of the lip-organ and at the
point where the tendinous transverse band, 4¢., and its lateral
connections occur in the ventral region. x oc. 2, obj. A.
Fig.7 4. Transverse section indicating the condition of the parts of the
lozenge-shaped tendinous region ventrally, as its upper and
outer edges differentiate into muscular fibres and strands
pass into the fasciculi beneath. x oc. 2, obj. A.
Fig. 8. Transverse section behind the foregoing, showing the diminishing,
pale, elastic band, d¢., and thecomplete condition of the ventral
longitudinal muscles, vm. The stomachal region is surrounded
by blood-vessels, bv. xX oc. 2, obj. A.
Fig. 9. Section in the line of the bristle-tufts with the opening of the
mucous gland, mp. ‘The stomach is still surrounded by the
various vessels and their mesenteries. A trace of the elastic
system is seen in the muscular differentiation, mc., of the
mid-ventral area. X oc. 2, obj. A.
Fig. 10. The thickening of the dorsal and ventral walls of the stomach
is conspicuous in this section (behind fig.9). Onthe left is
a bristle-tuft, 67. The median dorsal and ventral blood-
vessels and their mesenteries, and the ducts of the mucous
glands are seen on the way to the anterior. X oc, 2, obj. A.
Fig. 18. Vertical section of the region of the collar, col., and the central
nervous system, c., with the adjoining body-wall. x oc. 4,
obj. D, with 2-inch draw-tube.
Pirate IX,
Fig. 11, The body-wall has attained its general arrangement with the
exception of the thickened lateral portions of hypoderm, hype.
The coelomic space is large, and the median dorsal mesen-
tery is split inferiorly. The nerve-cords have now fused.
X oc. 2, obj. A.
Fig. 12. In this section, which is posterior to the preceding, the vertical
elongation of the alimentary canal is noteworthy, and the
accompanying vessels are still separate, though in some fusion
is indicated. The ccelom has its corpuscles, and the mucous
glands and their ducts are distinct. x oc. 2, obj. A.
Fig. 13, Transverse section after a considerable interval backward from
the foregoing, showing the great extent of the ventral longi-
tudinal muscles, vm., and the commencement of the male’
gonads, g., below the gut. The large area occupied by the
mgr and its enveloping sinus is noteworthy. x oe. 2,
obj. A.
264
Fig. 14.
Fig. 15.
Fig. 20.
Fig. 81.
Fig. 19.
Fig. 80.
Fig. 32.
Fig. 33.
Fig. 34.
Fg. 35.
Prof. M‘Intosh’s Notes from the
Section in front of the tail in a ripe male. The gut is still
enveloped in the spacious sinus, vs., with its dorsal and ventral
mesenteries, and the coelomic cavity is loaded with sperms.
The gonads, g., are large, as also is their blood-supply, 6v.,
whilst the nerve-cord is smaller than in front. The arrange-
ment of the muscles of the body-wall diverges from that in
front, since the dorsal longitudinal muscles, dm., are thick-
ened on each side of the middle line, as are also the ventral
above the nerve-cord ; but in the figure the parts have been
separated by thé methods of technique. The hooks, w., occur
in numbers along the whole lateral region on each side, the
bristle-tufts being shifted to the dorsal aspect. X oc. 2,
obj. A.
Section through the characteristic arrangement of the caudal
septa, spt., some distance behind fig. 14. The gonads, g., in
this region increase in size from before backward, and the
free sperms lie in the chamber above the upper septa. A
transverse septum is seen below the gonads. The dorsal
longitudinal muscles are thickest, whilst the ventral cover
two-thirds of the body-wall. x oe. 2, obj. A.
Peculiar pennate arrangement of the gelatinous tissue and
glands of the hypoderm anteriorly. x oc. 4, obj. D, with
full draw-tube.
Transverse section of the posterior region in a female with well-
developed ova, ov., which arise from the vascular ovigerous
tissue vetitrally, as in the male; some are cut and others
altered by compression. X 100 diam.
PLATE X.
Lonzitudinal section showing the arrangement of the mucus in
the mucous glands. X oc. 2, obj. 4.
Section through the stomach after the blood-sinus, vs., around
itis established. mg., mucous gland; bv., blood-vessels. The
median ventral furrow, making the organ bifid in section, has
now disappeared. X 100 diam.
Tip of the tail of Myriochele protruding from a tube. Two
dorsal papilla and a slightly more prominent ventral papilla
oceur posteriorly. The zig-zag condition of the gut in this
region is indicated. x 60 diam.
Section of the tip of the tail of Myriochele, showing the valvular
apparatus, val.p., at the constriction of the body-wall, the
septa, spt. The blood has been rendered opaque blackish
by the hematoxylin used in the technique, and apparently
has accumulated at the septa.
Transverse section of the extreme tip of the tail of a female with
the modified areolez of the hypoderm, a larger area on each
side being evident. xX oc. 2, obj. D, with 1 inch of draw-
tube.
Section of the tail a little in front of the foregoing, showing the
large areole of the hypoderm and the blood in the sinus, vs, |
around the gut. % oc. 2, obj. D, with 2-inch draw-tube.
Fug.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fug.
Fig.
16.
ive
24,
27.
28.
29.
Cr
Gatty Marine Laboratory, St. Andrews. 26
PuatE XI,
Vertical section of one half of the anterior end of Owenta fusi-
formis cutting the nerve-centre, ¢., across, and showing its
relation to the sensory groove with its pigment-corpuscles
on the anterior wall, in close relation to the central nervous
system. <A fold of the vestibule is seen at vf. xX oc. 2,
obj. A, with l-inch draw-tube.
2. Transverse section a Jittle behind the tip of the snout of Myrio-
chele on the appearance of the pale central nervous system, c.,
which becomes continuous with the nerve-cord on each side.
The mottled condition of the hypoderm anteriorly is charac-
teristic of this form. xX about 100 diam,
. Horizontal section through the vestibule and three regions of
the alimentary canal, viz., ws., cesophagus, s¢., stomach, and
int., intestine. spt., septa; val., valvular apparatus ; ¢., central
nervous system; bv., blood-vesseis, in some cases the dark
blood (coloured by hematoxylin) has been pushed beyond
the line of the vessel. x Zeiss oc. 4, obj. A, with 13-inch
draw-tube.
. In this section the cords are descending, being a little below the
middle line, and the sides of the vestibule are now slightly
united. Dorsal and lateral blood-vessels are in a similar
position to those in the foregoing figure. x 100 diam.
. The body-wall is completed, as also is the vestibule. The
nerve-cords, zc., are descending, and a section of the lip-organ,
Ip., appears. X 100 diam.
PLATE XII,
Sagittal section of the anterior end, to illustrate the complex
interlacing of the muscular fibres. The collar, col., and
central nervous system, c., are seen on the left. x oc. 2,
obj. A, with l-inch draw-tube. ;
Transverse section before the completion of the body-wall. The
nerve-centre, c., is well shown, and probably the lower ends
represent the commencement of the nerve-cords. Blood-
vessels, bv., occur both dorsally and laterally. x 100 diam.
In this section the outline of the body is slightly altered, the
vertical exceeding the transverse diameter. @s., cesophagus
with its cellular lining; Zp., lip-organ and its radiate arrange-
ment of cells; bv., blood-vessels. The dorsal and ventral
portions of the walls are much attenuated, partly from
stretching. x 100 diam.
The diminution of the dorsal hypoderm and the increase of the
ventral hypoderm are indicated in this figure, together with
the great size of the cesophagus. The nerve-cords are
approaching each other. x 100 diam.
Transverse section after the union of the nerve-cords, ne. The
succeeding region of the canal is joining the oesophagus on
the right, and two mucous glands, mg., are cut obliquely.
x 100 diam,
Ann, & Mag. N. Hist. Ser, 8. Vol. xix. 18
266 Mr. C. G. Lamb on Exotic Helomyzide, ce.
XX. — Notes on Exotic Helomyzide, Sciomyzide, and
Psilide. By C. G. Lams, M.A., B.Sec., Clare College,
Cambridge.
Helomyzidz.
He.omyza, Fall.
In the Wiener Ent. Zeit. for 1904 Czerny critically
examined all the species of this genus up to that date ; since
then only about half a dozen species have been added, and
hence the task of working out the specimens in the collections
was much simplified. ‘There was one well-known species
and three new ones, one of the latter being very interesting
as departing from the almost universal character of having
extra costal bristles.
Helomyza picta, Wied.
S. Ruopesta: Chirinda Forest (G. A. K. Marshall,
Camb. Coll.).
There was a fair series of this bandsome insect. It
exhibits considerable sexual dimorphism. ‘The sex described
is the male, and it has the dorsum elegantly variegated in
ochre and dark ochreous grey; its femora are beautifully
and regularly spindle-shaped, the mid pair less so, and they
bear long dense hairs below. The female has a quite dark
dull brown dorsum and scutellum, which exhibit only faint
signs of the male marks ; the femora are normal, less haired,
and the front ones have an anterior spine row—in fact, the
sexual differences in the legs are like those of some WScato-
phagas. ‘This type of Helomyza is devoid of the upper
patches carrying the orbital bristles, and also of the small
ocellar triangle joined to these, which are usual in the Euro-
pean forms; it has also pictured wings and swollenand hairy
male femora; this form seems to be typically African.
Speiser, in his Kilimandjaro-Meru Expedition paper, describes
two males of the same facies—//. acroleuca and 1, lactnata—
and the next species also belongs to this section.
Helomyza ingens, sp. n.
A single male of the pzcfa group was present ; it is larger
and more stoutly built than that species.
Head (top view) :—Frons and antenne entirely yellow to
orange, in front a little darker, with microscopic hairs and
Mr. C. G. Lamb on Exotic Helomyzide, &e. 267
irregular reddish patches, but with no sign of dark or black-
ened spots except the usual hairy neck-spot, an excessively
faint large spot behind the vibrissa, and the brownish-red
ocellar spot; the antenna has a darkish flagellum with very
long hairs, and the third joint has its absolute ridge red.
Side-view :—Jowls, palpi, tongue (also the face) all yellow ;
hind head and lower side bristly as usual; vibrissa very
strong ; eyes rather rectangularly oval, with long axis
vertical ; jowls about as deep as breadth of third joint of
antenna,
Thorax: dorsum dull dark coffee-brown, except the much
lighter callus and front of dorsum; the light part quickly
ple into the dark just as the true dorsum begins ; two
dark lines start near the neck, and continue right over the
dorsum, but are necessarily very faint on the dark part ;
between them, and also along the d.c. lines, are two faint
narrow lines showing up more ochreous than the rest. Scu-
tellum flat, slightly shining , quite bare, with a few tiny ridge-
airs between end- and side-bristles. All the macrocheetes
are Jong, but slender for the size of the insect; side above the
notopleural suture from the callus rather brown and shining,
below the pleura is as dark as dorsum, though a little more
shining ; mesopleura absolutely bare ; metanotum dull, more
grey ; “all bristles normal,
Wings darkened, the costal bristles stout and about fifteen
in number from end of vein 1, ceasing about level of hind
cross-vein. The general colour of the wing is brown; from
the end of 1 to the tip a darker brown covers the wing up to
about the middle of the cubital cell; the distal part of this
extra darkening extends across the wing; the basal parts of
veins 3 and 4 are included in another dar kened area, the
small cross-vein and neighbouring parts of 3 and 4 in
another, and the hind cross-vein and near parts of 3 in yet
another ; there are clear ‘‘ windows’ between the veins at
the base, and the usual “ window” just distal of the anal
cell shows very brightly ; the absolute costa between ends of
and 4 is pale ochreous, but otherwise all the veins are
brown, Halteres clear white.
Legs: femora all elegantly spindled, mid less so than others,
with long profuse blackish hairs at sides and below. Colour:
all coxee and trochanters orange, more or less darkened, all
femora shining black, all tibize orange, front with black tip,
hind suffused; all with long hairs below, which are ex-
cessively long on distal half of middle tibia; all tarsi orange,
the last joints daik and first joint of middle one with ver
long hairs. Bristles :—front femur a superior row of about 7,
13*
268 Mr. C. G. Lamb on Eaxotic Helomyzide, &e.
tibia with a distal anterior row of 3; mid tibia with crown
of about 4, the inner very long; hind femur with irregular
rows anteriorly totalling 10 to 12 bristles ; all tibize with usual
preapicals, hind ones with small inner spur.
Abdomen pitchy black, except at the scutellar angles of
the first segment ; hypopygium not large, very hairy.
Size 8} mm.
Britisu E. Arrica: Kenia Forest (7. J. Anderson, Imp.
Bur. Ent.).
The following species are of the normal European form,
with well-marked upper vertical patches and ocellar triangle,
and with simple legs. Both belong to the section with long-
haired arista and quite bare mesopleura.
Helomyza balteata, sp. n.
=o
Head (top view) :—Frons dullish orange and hairy, brighter
and bare in narrow lines on each side of the ocellar triangle
and along a mid line to the front; the upper vertical patches
and the ocellar triangle sharply bounded, grey, the former
with a pointed tip and only touching eyes just on vertex ;
hind head orange, with well-marked trapezoidal spot from
neck to vertex; all bristles normal. Face smooth, orange.
Side view :—Eyes rather elongate-oval, with the long axis
in the line joining outer vertical to the protuberant mouth-
angle; the latter is covered with a large dark patch, the rest
of face &c. being orange; antenna orange, arista black, with
long and strong pectination, stout slightly orange basal joints ;
long vibrissa with a small companion below; depth of jowl
about equal to breadth of third joint ; hind head orange and
bristly. Palpi orange, with slightly infuscate tip; tongue
orange.
Thorax: dorsum almost uniformly dull ochreous brown,
the tiny black bristles looking like a close regular punctation ;
a very faint pair of median lines between the d. c¢. bristles,
which stand on brown spots ; callus grey ; scutellum as thorax,
but a little paler centrally on disc and on the absolute tip,
quite bare and flat, with a few tiny hairs between main
bristles ; pleura orange above, merging to yellow below,
dull; mesopleura quite bare; metanotum dark, somewhat
shining orange.
Wing with about nine stout spines from end of vein 1 to
about level of hind cross-vein ; suffused, the darkening being
more intense from costa to just over second vein; both cross-
Mr. C. G. Lamb on Exotic Helomyzide, de. 269
veins well and broadly suffused; veins brown. Halteres
orange.
Legs: colour all orange, except that the hind knees and
all the last tarsal joints are brown, and the tibize have tips
browned. Bristles:—front femur with usual upper row and
inferior hairs ; mid femur with anterior row of 3 on distal
third ; hind femur with 5 bristles, three form an anterior
superior row, the last of these and two others, one above the
other, form a triangle ; usual tibial preapicals.
Abdomen orange, each segment with a black band based
on distal margin, narrow at side and broadening to middle,
with rather indistinct boundary there, so that the appearance
is like an indistinct mid-line with distinct side-teeth ; genital
segment orange, except extreme tip, which is black.
Size 7 mm.
S. Ruopesia: Chirinda Forest (G. A. K. Afarshall,
Camb. Coll.).
Helomyza aspinosa, sp. n.
This large species is quite aberrant, inasmuch as the
characteristic costal spines are not to be seen; they are
apparently so short as to merge in the general costal bristle-
border, which is more strongly developed than usual. ‘The
venation is quite normal, as is the complete cheetotaxy in
every respect. In broad facies the insect is not quite typical
of the genus, but looks rather like a Dryomyzid, The
absence of costal bristles is even more marked than is appa-
rently the case in the aberrant genus Thyreophorella, if one
may judge by the figure of the same.
Head (top view) :—All dull orange, darkened a little in
front, black-haired, especially in front ; the usual short hind
eye-borders and conjoined ocellar triangle are somewhat
paler, as is the hind head; the neck-spot is more orange and
a little silvery. Jace orange, a little darker over lip. Side-
view :—All orange, including the antennal third joint (which
is, however, brown on its absolute edge), arista (with very
long hairs), the hairy palpi, and the tongue; a long vibrissa,
with a few short neighbours below.
Thorax: dorsum dull reddish brown, a little darker in
front and lighter on side from d.c. lines to pleura; two
indistinct blackened lines run along the d.c. rows, with a
similar indistinct line outside from about the level of cross-
suture. Scutellum similar on the disc, which is quite flat
and bare, with a few tiny hairs between the main bristles
along the edge. Pleura (including callus and notopleura)
270 Mr. C. G. Lamb on Exotic Helomyzide, cc.
brightish orange, as are the sides of the scutellum and the
metanotum ; a darkened stripe runs from the end of the
callus to the wing-base ; the mesopleura is absolutely bare.
Wings quite normal, except as described above, somewhat
suffused, especially costally ; veins brown, more orange at
base.
Legs normal. Colour: all yellow, except for tiny side-
spots on the tips of middle and hind femora; front and hind
tarsi with all joints suffused, middle with last joint black.
Bristles : front femur with dorsal row of 6 and hairy below,
middle femur with anterior.row of 3 or 4, shaggy below,
tibia with a crown and shaggy below; hind femur with
superior row of 4 on distal half, rather haired below; all
tibiz with usual preapical.
Abdomen pitchy black, with basal segment more orange,
sides of all segments (except the last) and the venter orange,
genital segment black; all segments with border-bristles,
very long on the sides.
Size 91, wing 84 mm.
S. Ruopesra: Chirinda Forest (G. A. K. Marshall, Camb.
Coll.).
Sciomyzida.
SEPEDON, Latr.
In the Ann. Mus. Nat. Hong. vol. ix. (1911) p. 266,
Hendel published a paper on this genus, bringing the species
from Asia and Africa up to date ; since then only one species
has been added. The paper is conservative in treatment, and
clears up the confusion existent in the species as well as may
be. No references are given here, as the above paper is
used in what follows.
Sepedon violaceus, Hend.
Inp1A: Coimbatore.
Sepedon lobiferus, Hend.
A nice series of this interesting form was in the Cam. Coll.
under the name javanicus, R. D. The insect was hitherto
only known from Formosa, and its occurrence in the Hima-
layan district is of interest.
Inp1A: Showali, Kumaon.
Mr. C. G. Lamb on Evotic Helomyzide, de. 271
There are two species in the Cam. Coll. from Africa.
Speiser, in his paper on the Kilimandjaro-Meru Expedition
(x. 5, pp. 168, 169), describes the species ¢trichooscelis and
argyrostethus ; Hendel considers that the former is a syno-
nym of ornatifrons, Adams, the latter most probably of sene-
galensis, Macq. ; the author had independently come to the
same conclusion. Both of Speiser’s species were founded on
single damaged specimens, and in the case of this genus that
procedure is especially hazardous,
Sepedon ornatifrons, Adams.
A fair series isin the Camb. Coll., showing quite perceptible -
degrees of variation.
S. Ruopesta: Chirinda Forest (G..A. K. Marshall, Camb.
Coll.).
Sepedon senegalensis, Macq.
Natat: Durban (Ff. Muir, Camb. Coll.).
Psilide.
Cuyuiza, Fall.
There are two specimens from Ceylon which do not agree
with any of the known Oriental species ; in general appear-
ance they are very like C. leptogaster.
Chyliza pallidipes, sp. n.
Head (top view) :—The eye-borders black and somewhat
shining, about 4 of total frontal width, extending from
vertex nearly to front, but narrowing sharply in front; the
wider part of the frons lying between these anterior narrowed
parts is bright but dull yellow, the rest is brown ochreous,
but the long ocellar triangle is a little shining and its base
and the absolute vertex are rather shining orange ; the whole
head is covered with tiny golden pubescence; the bristles as
in C. leptogaster, but stouter in proportion, Face yellow, but
the lower half of the antennal pit is shining black ; the
narrow lower eye-margins slightly silvery. Side view :—
Antenna with deep black basal joints and orange third,
which is just perceptibly suffused on its edge ; arista pale
brown, with widely bipectinate flagellum, the total breadth of
the pectination being about equal to the breadth of third;
palpi deep black, tongue yellow. Hind head entirely shining
272 Mr. C. G. Lamb on Exotic Helomyzide, ce.
black above, black below, but widely bordered with yellow, so
that hind jowls and mouth-margin are all that colour; eyes
like C. leptogaster, with sinuate hind margin,
Thorax : dorsum subshining black, with uniform shallow
minute shagreening; entirely covered with elegant pale
yellow hairs, except for two lines, confluent in front and
abbreviated about halfway, which are bare (these are best
seen with oblique light); the dorsum is much like lepto-
gaster, Pleura like the dorsum in front, but with silvery
white hairs, longest below ; the sclerites over the hind leg
are all smooth, hairless, and very shining; just over the
callus is a rather bright orange narrow bar, and a dark orange
one is just visible on the top of the sternopleura. Scutellum
all rather shining orange, bristled as in leptogaster.
Wings slightly smoky, especially broadly so at tip between
costa and vein 4; the latter is parallel to 5 up to about its
distal fifth, when it makes a sudden slight bend upwards ;
veins brown, extreme base of wing orange. Haltere with
snow-white head-and slightly brownish stalk.
Legs all pale straw-coloured, a little whiter proximally on
the femora, with no sign of any rings or darkening.
Abdomen like the thorax in colour and punctation, but the
hairs are brownish ; the shape is more wasp-like than in
leptogaster.
Size 5 mm.
CryLon: Peradeniya (A. Rutherford, Imp. Bur. Ent.).
Loxocera, Mg.
In the Kilimandjaro- Meru Expedition Reports (x. 5, p. 193)
Speiser gives a table of the known African members of this
genus. Of these, L. dispar, Bezzi, is apparently quite
distinct, having a black triangle, sternopleura, and front
femora. He separates the others on the presence or absence
of thoracic stripes and their position: thus, L. rufa, Loéw,
is given as stripeless, L. lateralis, Lw., and L. macrogramma,
Speiser, are striped in different ways.
In the Camb, Coll. are eleven specimens of a red Lowocera
of the latter group, They are evidently closely related, and,
apart from thoracic marks, differ only in the degree of undu-
lation of the fourth vein between the cross-veins, and the angle
between the last cross-vein and the fifth; in such cases
where the veins are wavy or bent (as is the cross-vein here
concerned) the angle in question and the amount of curvature
of the veins is always a little uncertain. Apart from this
and the colour of the thorax, neither of which are correlated
On Hyale grenfelli, Chilton. 273
with one another nor with the sex, no structural difference is
apparent. Now the thoracic dorsum varies greatly ; of the
seven males, one is clear red (typical rufa), one has side-lines
just in front of the scutellum, one has side-lines complete but
no middle one (this is typical macrogramma by comparison
with Speiser’s full description), one has side-lines and a faint
middle one in front, one has all these lines well marked, one
has all the lines broad and even confluent at the middle of
the dise, and another has the lines reddish but a little dark
behind. Of the four females, one is quite immaculate, one
has only the faintest trace of the lines in red, one has the
lines all present but faint, the last has all present and very
strong. The author is quite sure that Speiser’s macro-
gramma is a dark-lined form of rufa. As regards the two
species of Loéw (rufa and lateralis), some doubt may arise.
Loéw evidently had single specimens only (rufa is described
from a 2, lateralis from a @; see B. E. Z. 1874, xliv.
p- 194). The main difference appears to be dark flecks in
the antennal pits in the latter species and (possibly) less
hairy arista. It is impossible to be sure of the true distinet-
ness of these three species, and hence the author considers all
the red Lowoceras with entirely black third joint to be L. rufa,
Loéw.
S. Ruopesia: Salisbury and Chirinda Forest (G. A. K.
Marshall, Camb. Coll.). Natau: Durban (F. Muzr, Camb.
Coll.).
XXI.—Further Notes on the New Zealand Amphipod Hyale
grenfelli, Chilton. By Cuas. Cuitton, M.A., D.Sc.,
M.B., C.M., LL.D., C.M.Z.8., Professor of Biology,
Canterbury College, New Zealand.
In May 1916 * I described a new species of Amphipod from
New Zealand, naming it yale grenfellt. The type-specimen,
which was the only specimen at that time known, was a
male, and was characterized by the peculiar shape of the
second gnathopod and by the great dilatation and setose
character of the terminal joints of the maxillipeds. I pointed
out that it was quite likely that this peculiar development of
* Ann, & Mag. Nat, Hist. ser. 8, vol. xvii. p. 362.
274 ' Prof. C. Chilton on
the maxillipeds would be found in the male only, and was
probably to be looked upon as a secondary sexual character.
On December 12th, 1916, I received from Mr. C. R. Gow,
of the Moko Hinou Lighthouse, a small collection of Crus-
tacea which had been taken between tide-marks on Moko
Hinou, a group of islands off the east coast of Auckland,
situated about 50 miles from Cuvier Island, where the type-
specimen was obtained. Among these Crustacea there are
fortunately a few specimens of yale grenfelli. Most of
these are males, showing the peculiar characters in the
maxillipeds and the second gnathopod as described. One
Fig. 1.
Hyale grenfelli, 9. Maxillipeds.
specimen is a female, 5 mm. long, bearing five large eggs in
the brood-pouch, and I am therefore now able to describe the
characters of the female. In it the maxillipeds (fig. 1),
though, perhaps, a little larger than in the majority of the
species of LHyale, have the terminal joints only slightly
enlarged, and not showing the special form nor the numerous
long setee characteristic of the male; the carpus bears one
long seta at its outer distal angle and a few on the inner
margin near the distal angle, but there are none on the
surface of the joint itself; in the propod the inner margin
Hyale grenfelli, Chilton. 275
bears a regular row of long sete and there are two rows on
the surface near the outer distal end, lying more or less
parallel to the distal border, and proximal to these there is
one short row of four or five setee and a single seta situated
still more proximally ; the dactyl is small, very much nar-
rower than the propod, and bears at the end a long, stout,
curved seta which is proportionately much more prominent
than the corresponding seta on the dactyl of the male. It
will be seen that the maxilliped in the female presents the
ordinary characters common to allied species of Hyale, and
that its terminal joints show none of the numerous transverse
Hyale grenfelh, 2. First gnathopod.
rows of long fine setee on the surface that are so charac-
teristic of the male. ;
In the gnathopoda the first pair (fig. 2) are, on the whole,
similar to those of the male, but more slender ; the side-plate
is large, produced a little anteriorly, so that it is widest
below ; the carpus bears a fringe of sete on its posterior
margin, as in the male, the propod is much more slender
than the corresponding joint in the male, only widening very
slightly distally, and its posterior border bears only a small
tuft or short row of sete near the centre instead of having
nearly the whole of the margin fringed with a row of setx as
276 On Hyale grenfelli, Chilton.
in the male. The second gnathopod (fig. 3) is almost exactly
the same as the first, but is very slightly larger, and the
side-plate differs in being regularly rectangular and not
widening below ; the other joints of the appendage show no
differences from the first worthy of notice.
In other characters the female closely resembles the male.
Nearly all the males are apparently fairly well developed,
and show the characters of the second gnathopod and the
maxillipeds nearly as originally described. In one which is
Hyale grenfelli, 2. Second gnathopod.
about 5 mm. in length the terminal joints of the maxilliped
are rather less expanded and not quite so setose, and in the
second gnathopod the propod is not so wide, the palm is more
oblique and much less concave, being nearly straight or only
slightly concave, and its outer and inner borders are less
widely separated ; the dactyl, however, is short and fairly
stout, almost as in the typical male. Doubtless in still
younger specimens of the male these appendages would show
the characters of the male to a still less extent and be more
like those of the female.
On new Lizards of the Family Lacertide. 277
XXIT.—Descriptions of new Lizards of the Family Lacertide.
By G. A. Boutencer, F.R.S.
(Published by permission of the Trustees of the British Museum.)
Lacerta viridis, var. woosnami.
Head comparatively short, its width 12 to 1} times in its
length. Occipital } to 2 the length of the interparietal, not
or but slightly broader than the latter; 2 to 8 granules
between the supraoculars and the superciliaries ; temple with
12 to 20 shields, with a large or very large masseteric, which
may extend from the upper temporal to the upper labials,
Fig. 1.
Upper and side views of head.
the tympanic well developed and usually in contact with the
upper temporal. Dorsal scales rhombic and strongly keeled,
considerably larger than the laterals; 38 to 43 scales across
the middle of the body. Ventral plates in 6 longitudinal and
24 to 28 transverse series. 17 to 20 femoral pores on each
Fig. 2.
Lepidosis of middle of body.
278 Mr. G. A. Boulenger on new
side. 25 to 27 lamellar scales under the fourth toe. Green
or olive-grey above, uniform or with small black spots,
sparsely scattered on the back, more crowded on the sides,
sometimes (in a single female) forming a regular vertebral
series ; upper surface of head uniform green, olive, or brown ;
lower parts yellow, greenish, but not blue, on the throat and
on the sides of the belly. Young brown or olive, with three
white longitudinal streaks on the back, traces of which may
be preserved in the adult; black spots may be present
between them; a white streak on each side of the neck,
from the tympanum, continued on each side of the body
or breaking up into two series of round spots; another white
line along each side of the belly.
From snout to vent, ¢ 102 mm., 2? 95; tail, ¢ 190,
9 207.
This form connects the var. strigata with the typical
L. viridis, and especially the oriental specimens on which
the name var. vaillanti, Bedr., has been bestowed, agreeing
with the latter in the temporal scutellation and the reduction
in the number of superciliary granules, with the former in
the presence of a light vertebral streak in the young; it
differs from both in the lepidosis of the body, in respect
to which it approaches L. princeps.
This variety is described from eight specimens obtained
by the late Mr. R. B. Woosnam on the South Coast of the
Caspian Sea, and from one young obtained by Mr. R. T.
Giinther at Bash Nurashin, N.W. Persia, which I had
referred to the var. strigata (Journ, Linn. Soc. xxvu. 1899,
p. 378).
Ichnotropis tanganicana.
Form and lepidosis as in J. capensis, Smith, but upper
head-shields rather feebly striated and the four super-
ciliaries in contact with the four supraoculars, only 3 or 4
small granules intervening between the second and third
superciliaries and the supraoculars, and lower nasal but
narrowly in contact with the rostral. 36 scales and plates
round the middle of the body; ventral plates in 8 longi-
tudinal and 25 transverse series. 11 or 12 femoral pores on
each side. 19 lamellar scales under the fourth toe. Bronzy
olive above, with a few small transverse blackish spots in
three longitudinal series on the nape and two on the body;
a black streak from the nostril to the eye, and another on
the edge of the mouth; a white, black-edged streak from
Lizards of the Family Lacertide. 279
below the eye, through the ear, to above the axil; white,
black-edged ocellar spots on the posterior part of the back,
on the hind limbs, and on the tail ; lower parts white.
From snout to vent 38 mm.
This species, which I regard as the most primitive of the
geuus, as it is also the northernmost in its habitat, is based
on a single male specimen, probably half-grown, from. the
East Coast of Lake Tanganyika, presented to the British
Museum by Mr. W. H. Nutt in 1896.
Eremias adramitana.
Head and body strongly depressed, limbs very slender ;
head 14 times as long as broad; snout pointed, with the
nasal shields rather strongly swollen, as long as broad, as
long as the postocular part of the head ; hind limb reaching
between the collar and the ear in males, the shoulder or
the collar in females ; foot 14 to 14 times as long as the
head ; toes slender, feebly compressed ; tail 12 to 24 times
as long as head and body. Lower eyelid with a semi-
transparent disk divided into 5 to 8 scales. Lepidosis as in
EF. guttulata, but occipital minute or absent, the parietals
meeting in the middle, and ventral plates in 10 regular
longitudinal series, mostly as long as broad or a little
broader than long, the outer longer than broad. 31 to 40
scales across the middle of the body. 11 to 15 femoral
pores on each side. Subdigital lamellee tricarinate, 20 to 23
under the fourth toe. J'awn-coloured or pale grey above,
with or without small brown spots, which may be irregular
or disposed in two longitudinal series on the back, with or
without small whitish spots; a dark brown lateral band,
often bearing white spots, from behind the eye to the tail,
bordered below by a white or yellowish lateral streak passing
through the tympanum; upper surface of limbs marbled
with brown, or with white spots ; lower parts white.
From snout to vent 44 mm.
This species has been confounded with £. brevirostris,
Blanf., of which the Syrian , bernoulli, Schenkel, is a
synonym, by Anderson, ‘Herpetology of Arabia,’ p. 43
(1896). It differs in the more depressed head, longer in
proportion to its width, the more slender limbs, and the ventral
plates constantly in ten longitudinal series. Itis only known
from the Hadramut, South Arabia, whilst LZ. brevirostris is
on record from Kalabagh in the Punjab, Bushire in Persia,
Tumb Island in the Persian Gulf, and Syria,
280 On a new Bat of the Genus Scoteecus.
XXIU.—A new Bat of the Genus Scotecus.
By OLDFIELD ‘T'HOMAS.
(Published by permission of the Trustees of the British Museum.)
AMONG a series of well-prepared skins from Nyasaland
presented to the National Museum by Mr. Rodney C. Wood,
there occur examples of several rare bats, notably Myotis
welwitschii and bocage’, Glauconycteris papilio, Eptesicus
megalurus, and a Scotecus which appears to be new. The
last may be called
Scotaecus woodt, sp. n.
Near S. albofiiscus of the Gambia, but smaller.
Size about the smallest of the genus. General colour
above dark brown (near mummy-brown), the tips of the
hairs paler brown; under surface little paler, near Prout’s
brown. Wings coloured as in S. albofuscus, the forearms,
digits, hind limbs, and tail blackish, the membranes internal
to a line from elbow to knee, and the interfemoral dark
brown, those external to forearms dull whitish, rather darker
terminally. Hars short, with large external basal lobe ;
tragus short and broad, its inner margin slightly concave.
Skull short and stumpy, of the characteristic broad shape
usual in the genus, the lacrymal breadth even greater than
in S. albofuscus. Nasal notch very deep. Median part of
zygoma absent in type.
Incisors slender, their bases not touching the canines.
Canines broadened transversely, their basal area broader
than long, and flattened behind, close and parallel to the
front edge of the large premolar ; no small premolar or place
for it present.
Dimensions of the type (the italicized measurements taken
in the flesh) :—
Forearm 28:5 mm.
Head and body 56 mm. ; tail 27 ; ear 12.
Third finger, metacarpus 28, first phalanx 10, second
phalanx 8 ; lower leg and hind foot (c. u.) 17:5.
Skull: greatest length 13°2; median upper length 11;
basi-sinual length 9°8; greatest breadth 10°3; lacrymal
breadth 6:7 ; mastoid breadth 9°1 ; palato-sinual length 4:5;
front of canine to back of m’* 4:9.
Hab. Southern Nyasaland. Type from Chiromo ; alt.
200’.
On anew Species of Aconemys. 281
Type. Adult male. B.M. no. 17. 2.1.1. Original num-
ber 173. Collected 2nd October, 1916, and presented by
Rodney C. Wood, Esq.
This species may be distinguished from its only close,
though geographically very distant, ally S. albofuscus by its
smaller size, proportionally even broader skull, and the
different shape of the base of its canines. The other members
of the genus all have uniformly brown wing-membranes.
I may note that of twelve skulls of Scotewcus, including
examples of all the described species, only two have com-
plete zygomata, although all have been prepared by that
most skilful skull-cleaner Mr. W. Sherrin. Imperfection or,
at least, excessive tenuity of the zygoma would therefore
appear to be an additional character of the genus Scotcecus.
Of forty skulls of Scotecnus similarly prepared by Mr. Sherrin,
nearly all have perfect, although very slender, zygomata.
XXIV.—A new Species of Aconzeemys from Southern Chil.
By OLDFIELD ‘'l’HoMAS.
(Published by permission of the Trustees of the British Museum.)
THE British Museum has recently received from Mr. J. A.
Wolffsohn a specimen of the rare genus Aconemys (Schizodon,
Waterh.) which had been presented to him by the well- known
naturalist Don Carlos E. Porter. The species proving to
be new, I propose to name it in honour of the latter, to whom
the Museum has been indebted for help in various ways.
Aconemys porteri, sp. n.
Fur more woolly than in A. fuscus; tail more completely
bicolor; incisors stouter.
Size about as in A. fuscus or rather smaller. Fur soft,
more woolly, less straight than in A. fuscus, the general
texture and the colour both suggesting that of a Huropean
water-vole (Arvicola amphibius). General colour deep rich
brown, near “ auburn ” of Ridgway, the subterminal rings on
the hairs dull cinnamon. Under surface similar but rather
warmer in tone, the ends of the hairs rich cinnamon. Hands:
and feet greyish white, the middle part of the metatarsus
rather darker. Tail rather longer than in A. fuscus and
completely bicolor, black above and creamy whitish below for
Ann, & Mag. N. Hist. Ser. 8. Vol, xix. 19
282 Mr. T, D. A. Cockerell—Deseriptions and
its whole length; in A. fuscus the terminal part of the
under surface is brown.
Skull apparently somewhat smaller than in A. fuseus, but
the age of the type is not very certainly determinable, An-
terior. part narrower, the breadth between the outer corners
of the anteorbital foramina and the interorbital breadth both
distinctly less.
Incisors very stout and heavy, decidedly thicker than in
specimens of A. fuscus of apparently similar age.
Dimensions of the type :—
Tail (vertebra in skin) 64; hind foot (dry) 28.
Skull: tip of nasals to back of frontals 28°5 ; greatest
breadth 23; nasals 15 x 6°3 ; interorbital breadth 7° 5; breadth
between outer corners of anteorbital foramina 17°6 ; ‘palatilar
length 16°2; front of incisors to back of m* 21:5; upper
tooth-row (crowns) 84; combined breadth of upper in-
cisors 4°7.
Hab. Osorno, 8. Chili.
Type. Adult. B.M. no. 16.11.14.4. Presented by Don
Carlos E. Porter to Mr. J. A. Wolffsohn. °
The British Museum contains eleven specimens of Aco-
nemys fuscus, received at different dates from Mr. T. Bridges,
but whether all were from the ‘* Valle de Las Cuevas, on the
east side of the Andes, near the Voleano of Peteroa, altitude
6000’,” where Mr. Bridges discovered the species, there is,
unfortunately, no evidence to show. But all agree in the
characters used above in separating the southern form, which
is probably an inhabitant of the high slopes on the Volcano
of Osorno, some little distance from the town of the same
name.
Since the time of Mr. Bridges no examples of this genus
have come to the British Museum, nor has our indefatigable
correspondent Mr. Wolffsolin been able to see or hear of any.
Consequently this additional specimen, representing a second
and more southern species of the genus, is an extremely
welcome accession.
XXV.— Descriptions and Records of Bees—LXXIV.
By 'T. D. A. Cockeret, University of Colorado.
ALL the bees recorded in the present part are in the U.S
National Museum.
Andrena lugubrescens, n. n.
Andrena lugubris, Lepeletier, 1841 (not Erichson, 1840).
3 .—Belvidere, Tunis, May 10, 1899 (P. ALagretti).
Records of Bees. 283
This is like the male of A. albopunctata, Rossi, but has the
abdomen shining, irregularly wrinkled, and with scattered
minute piliferous punctures, so it is doubtless the male of
A, lugubris, described by Lepeletier from the female only.
Length about 12 mm.
Process of labrum prominent, shining, truncate, slightly
emarginate ; first recurrent nervure joining second sub-
marginal cell distinctly before the middle.
The name /ugubris is preoccupied, so lugubrescens is pro-
posed as a substitute. -
Andrena cussariensis, Morawitz.
9 .—Kohat, N.W. Provinces, India, March 1906 (Frank
Benton).
Superficially this looks like A. morio, which Bingham
records from the Simla hills; but it is certainly distinct from
morio, and, as far as can be gathered from Morawitz’s quite
full description, agrees well with cussartensis. The abdomen
has very fine punctures, and the process of labrum is much
narrower than in morio. ‘The species is more closely allied
to A. ephippium.
Andrena cussariensis kohatensis, var. nov.
? .—Length about 14°5 mm.
Scutellum and broad bands at sides of mesothorax terra-
cotta red.
Hab. Kohat, India, March 1906 (Frank Benton).
This variety suggests comparison with A. ephippium,
Spin., to which it is closely allied. It differs from ephippium
by the narrower thorax, the scutellum much narrower, and
less closely punctured on disc; the flagellum only very
obscurely reddish beneath, the shorter fourth antennal joint,
the less strongly sculptured area of metathorax, and the
broad hind margin of first abdominal segment excessively
finely punctured, abruptly contrasting with the rest of the
segment. The hair of hind legs is entirely black.
Should comparison of specimens indicate that this species
is to be separated from A, cussariensis, it may be known as
A. kohatensis.
Andrena chionospila, sp. n.
9? .—Superficially exactly like A. albcopunctata, Rossi
(specimen from Ras-el-Ma, Algeria, compared), but differing
thus :—Antennz shorter ; process of labrum, although very
13°
284 Mr. T. D. A. Cockerell— Descriptions and
broad, not so broad ; area of metathorax smaller, less rugose ;
punctures of abdomen conspicuously more feeble and less
dense; white hair-patches at sides of abdomen larger.
&.—Very like the female, except in the usual sexual
characters ; head very broad ; region of mouth, sides of face,
and region of antenna with long black hair, but face otherwise
with long white hair, which is dull, not clear white as in 9 ;
cheeks broad, with black hair; abdomen less distinctly
punctured. :
Hab. Menserah, N.W. Provinces, India, March 1906
(Fra vk Benton).
Perhaps a subspecies of A. a/bopunctata.
Andrena subspinigera, sp. Nn.
9 .—Lenegth about 11 mm.
Head, thorax, and legs black ; abdomen with the first three
segments clear ferruginous (the first with a broad transverse
black band, the third with an interrupted suffused dark band
beyond the middle), the others black, the third and fourth
with heavy fringes of pure white hair, the second with a
thin inconspicuous fringe, the caudal fimbria brownish black.
Hair of head and thorax white, with a slight creamy tint on
thorax above ; facial fovese rather narrow, seen from above
shining white, with the upper end brown; facial quadrangle
broader than long ; process of labrum broad and obtuse, with
sloping sides; clypeus dull except at sides, with sparse
punctures ; flagellum bright ferruginous beneath except at
base; third antennal joint almost as long as next three
together; mesothorax and scutellum dull, without well-defined
punctures, the long hair not concealing the surface; area of
metathorax dull, minutely granular, scarcely defined ; tegulee
pale yellowish testaceous. Wings strongly reddened, stigma
and nervures rufo-fuscous; b. n. meeting t.-m. ; second s.m.
large, receiving first r.n. considerably beyond middle. Scopa
of hind tibiee compact, fuscous behind (above), white in
front; basitarsi broad and flat. Abdomen dull, minutely
granular, without any evident punctures.
Hab. Menserah, N.W. Provinces, India, March 1906
(Frank Benton). On some labels the locality is written
“’ Manserah,” on others “ Menserah.”
This species is very like A. sp’n’gera, Kirby, from Quetta,
buat differs by the flagelluin red beneath, the dusky reddish
wings, the dull abdomen, &c.
Records of Bees. 285
Andrena quettensis, sp. n.
3g .—Length about 9 mm.
Black, the hind tarsi, apical half of middle tarsi, broad
apical band on first abdominal segment, and second segment
except a spot on each side and a dusky cloud in middle (or
only the apical margin and a broad semilunar area on each
side basally) all ferruginous red. Head broad, facial
quadrangle much broader than long; mandibles rather short,
red at end; process of labrum broadly. emarginate ; no light
face-marks ; clypeus dull and granular ; face and front
covered with long sooty hair, paler and reddish about middle
of face, becoming black around margins ; occiput and lower
part of cheeks with long pale fulvous hair; cheeks broad,
but rounded behind ; antenne long, reaching metathorax ;
flagellum thick, crenulate, entirely dark ; mesothorax and
scutellum dull; area of metathorax triangular, coarsely
wrinkled, poorly defined ; hair of thorax long and fulvous ;
tegule fuscous, the outer margin paler. Wings long,
reddish hyaline, stigma and nervures amber-colour ; second
s.m. receiving first r.n. well beyond middle. Legs with
pale hair, golden on inner side of tarsi. Abdomen shining,
the dark segments beyond the middle with a very slight,
hardly observable, greenish tint; segments with very thin
bands of long pale hair ; apical plate broadly emarginate.
Hab. Quetta, India, March 1906 (Frank Benton).
This does not agree with any of the species reported by
Nurse from Quetta ; the nearest is A. balucha, Nurse, which
has more red on the abdomen and much paler hair on head.
It is just possible that A. quedéensis represents an extreine
colour-variation of A. balucha, but it seems to be quite
distinct. In Apidee Europes A. quetéensis runs to A. cingu-
lata and’ A. laticeps, but differs at once by the colour of hair
on head. A. balucha, which I have examined in U.S.
National Museum, has the area of metathorax of the Trach-
andrena type.
Andrena bentoni, sp. n.
? .—Length about 9 mm,
Black, including legs and abdomen ; hair of head and
thorax abundant, erect, but not hiding surface, very pale
greyish ochreous, black on vertex ; facial quadrangle con-
siderably broader than long ; clypeus shining, strongly and
closely punctured, without any distinct smooth line; man-
dibles red apically; process of labrum broadly truncate ;
286 Mr. IT’. D. A. Cockerell—Descriptions and
facial foveee reddish brown, separated from eye by a distinct
punctured band; flagellum obscure brownish beneath except
at base; third antennal joint about as long as next three
together ; fourth and fitth short and about equal, sixth
longer ; mesothorax dull, closely and distinctly punctured ;
scutellum shining ; area of metathorax granular, minutely
plicate at extreme base; tegule fuscous, posteriorly ferru-
ginous. Wings strongly reddened, stigma and nervures
ferruginous; second s.m. receiving first r.n. in middle. Legs
with pale hair, scopa of hind tibize dense, entirely pale golden
fulvous. Abdomen broad and flattish, glistening, very finely
and closely punctured, second segment depressed hardly one-
fourth; hind margins of segments 2 to 5 with rather weakly
developed white hair-bands ; apical fimbria dark chocolate.
Hab. Menserah, N.W. Provinces, India, March 1906
(Frank Benton).
In Apide Europese A. bentont appears to fall nearest to
A. propinqua and A. separanda, but the hair of thorax is
quite differently coloured. ‘There is no close resemblance
to any of the Indian species.
Andrena precocella, sp. n.
6 —Length 7°5-8°5 mm.
Black, with long black and white hair. Very close to
A.-precor, Scop., but differing thus:—Hind margins of
second and third abdominal segments more or less brown or
red; mandibles with no basal tooth beneath; head equally
broad, but longer ; upper part of cheeks punctured; light
hair of thorax above white (not yellowish) ; fourth and fifth
abdominal segments with thin white hair-bands; apical
plate of abdomen emarginate, shaped like a fish-tail.
Compared with the Japanese A. pracociformis, Ckll., it
differs by the large amount of black hair at sides of face, the
cheeks strongly angled behind, the black hair on meta-
thorax, &e.
Hab. Quetta, India, March 1906, 5 g (Frank Benton).
The females of this group are very unlike the males, so I
thought it possible that Nurse might have described the
species from Quetta in the female sex. ‘There is, however, no
description which seems possibly applicable. In A. preco-
cel/a the fourth antennal joint is about 256 microns long,
the fifth 8320. The mandibles are long and falciform,
Apis florea nasicana, Cockerell.
Kohat, N.W, Provinces, India, March 1906 (Frank
Benton).
Records of Bees. 287
Tetralonia pomona (Nurse).
Both sexes ; Quetta, India, March 1906 (Frank Benton).
Tetralonia kohatensis, sp. n.
3 —Length 85-10 mm. ; antenne about 6°5 mm.
Black, with the small joints of tarsi ferruginous ; clypeus,
labrum, and basal half of mandibles clear sulphur-yellow ;
mandibles red in middle and black apically ; antenne long
and slender, bright ferruginous beyond the third joint, the
upper side dusky ; third antennal joint much longer than its
apical width, dark fuscous, abruptly contrasting with fourth ;
eyes green; maxillary palpi rather short, bat six-jointed ;
liead and thorax above, as well as front and upper part of
face, with long pale fulvous hair, cheeks and underside of
thorax with white hair ; dise of mesothorax shining ; tegule
light reddish fulvous. Wings clear, faintly brownish in
apical field ; stigma and nervures reddish fuscous ; first r. n.
meeting second t.-c. or falling a little short of it; marginal
cell obliquely truncate. Outer side of tibize with dense white
hair; tarsi with ferruginous hair on inner side; spurs
creamy white. Abdomen shining, with piliferous punctures ;
apical margin of segments broadly pallid, covered with dense
bands of pale ochreous tomentum, of equal width right across,
the band on first segment narrow ; no definable basal bands ;
lateral margins of sixth segment briefly dentate.
Hab, Kohat, N.W.Provinces, India, 4 ¢,. March 1906
(frank Benton).
Related to 7’. erythrocera, Cam., but easily separated by
the fulvous hair. Superficially the insect is exactly like
Tetraloniella aliena, Ckll.
Anthophora connewxiformis, sp. 0.
6 -—Length about 14 mm.
Robust ; black, including legs and antenne (except a very
small cream-coloured line on scape), with a short linear
creamy mark on each orbital margin below level of antenna,
aud a large cream-coloured area on clypeus, broad below,
narrowed to a band above (inverse goblet-shaped), but
labrum and mandibles wholly black ; eyes bright ochreous;
facial quadrangle much longer than broad; mandibles with
a large rounded tooth on inner side ; malar space well deve-
loped ; third antennal joint fully as long as next three
united, the fourth very short; clypeus, labrum, cheeks
(except upper part anteriorly), and occiput densely covered
288 Mr. T. D. A. Cockerell—Deseriptions and
with very long pure white hair ; front, vertex, sides of face,
and upper part of cheeks anteriorly with black hair ; thorax
with very long hair, mixed grey and white, dark on scutellum,
shining white on mesopleura; tegule black, very hairy.
Wings hyaline. Legs slender, with Jong black and white
hair, dark chocolate on inner side of tarsi; apical joint of
middle tarsi with no noticeable fringe ; hind basitarsus long
and broadened. Abdomen not banded or spotted, but with a
profusion of long erect hair, which is mostly greyish white,
but black on discs of fourth and following segments, though
white and very long at sides.
Hab. Quetta, India, March 1906 (Frank Benton).
Closely allied to A. connexa (Nurse), also from Quetta;
but according to Nurse’s description connexa has the clypeus
all yellow, the apical tarsal joints more or less rufo-testaceous,
the blackish hair of abdomen confined to the apical two
segments, and the front with white hair. It thus seems
probable that our insect is a distinct species, though it may
be only a variety. ‘There is a pencil of white hair on each
side of front, a little above level of antenna. The general
appearance of the insect is very like that of Yetralona
pomona.
Anthophora (Micranthophora) albopicta, sp. n.
9? .—Length about 11 mm., anterior wing 8 mm.
Black, including the legs and antenne, but mandibles
ferruginous with the lower basal corner broadly black ;
labrum black, with a very broad white band down the
middle ; clypeus with a large apical white triangle, attenuated
above, this on a light ferruginous field, which extends as a
band to upper margin, but the upper half of clypeus black
except in middle; eyes greyish ochreous, converging below,
the front very broad; flagellum very obscurely reddish
beneath; third antennal joint about 640 microns long,
the next three together about 735; maxillary palpi with
stout bristles, except on the last two joints; third joint of
Jabial palpi 560 microns from base to origin of fourth joint ;
pubescence very pale ochreous, nearly white, long on head
and thorax; on head and thorax above with black hairs
intermixed ; mesothorax extremely densely punctured ;
teoule piceous. Wings hyaline, with a very taint brownish
tint. Legs with creamy white hair, rusty black on inner
side of hind tibize and tarsi, anterior and middle tibize with a
small patch of ferrugiuous hair at apex. Abdomen broad,
.
Records of Bees. 289
hind margins of segments whitish hyaline ; whole surface of
dorsal segments rather thinly covered with appressed pale
hair, not forming bands ; apex with a patch of black hair ;
apical plate very long and narrow.
Hab. Kotal Malul, 8. Persia, Feb. 1906 (Frank Benton).
A typical Micranthophora, looking just like the Californian
A, anstrutheri, Ckll., though differing in the face-markings
and many other details. It is also related to the Indian
A. candida, Sm., but the pubescence of the abdomen in that
species is much more dense, the face-markings are different,
and the flagellum is red beneath,
Anthophora cincta (Fabricius).
Axim, Gold Coast, Africa (C. R. Mengel).
Anthophora antimena, Saussure.
Mahanoro, Madagascar, May 5, 1895 (W. L. Abbott).
Anthophora acraensis (Fabricius),
Luebo, Congo (D. W. Snyder).
Anihophora flavicollis, Gerstaecker.
Axim, Gold Coast, Africa (C. R. Menge’).
Anthophora leucorhina, sp. n.
3 (type).—Length about 15 mm.
Black, including flagellum and legs, except the reddish
apical joint of tarsi; face-marks creamy white, including
clypeus, labrum (except large black spot at each basal corner,
and black apical margin), elongate spot on base of mandibles,
narrow stripe along each anterior orbit (beginning at about
level of antenne, but not reaching lower corner of face), and
anterior surface of scape ; clypeus prominent, convex ; third
antennal joint about as long as next three combined ; face
and cheeks with long pure white hair, occiput with yellowish,
vertex and front with black hair, but some white on each
side of antenne, and some long black hairs at sides of face ;
malar space well developed; thorax with abundant long
hair, pale greyish-yellow above and on upper part of sides,
290 Descriptions and Records of Bees.
black on anterior part of scutellum, and white on lower part
of pleura; mesothorax duil, slightly shining on dise ; tegule
piceous. Wings hyaline, very faintly brownish apically.
Legs with long white hair; middle tarsi not modified, nor
with any black fringe on last joint; hind basitarsi_ not
toothed. Abdomen shining, with piliferous punctures, the
surface covered with long hair, pale greyish-yellow on first
two segments, black on the others, but hind margins of
segments 2 to 4 with loose bands of white hair ; venter with
long white hair,
¢ .—Length about 16 mm.
Tongue very long ; no pale face-marks, but a red tubercle
on each side of base of labrum, and malar space red ;_ hair of
front pale ; disc of mesothorax and anterior part of scutellum
with some dark hair, not conspicuous ; tegule rufo-testa-
ceous ; patches of fulvous hair at apices of anterior and
middle tibize and on hind knees; hair on inner side of hind
tibiee (except base) black, on inner side of hind tarsi largely
red, in some lights appearing rich fox-red with black margin ;
abdomen with broad pale hair-bands on segments 2 to 4;
apex with black hair ; apical plate long and narrow ; venter
with white fringe on segments 2 to 4, but dense black hair
on apex of 5.
Hib. Kotal Malul, 8. Persia, Feb. 1906 (frank Benton),
a ok ee
Htesembles A. cinerea (Friese), from Sarepta, but is con-
siderably larger, There is a general resemblance to A. cri-
nipes, Sim., but the middle tarsi are not modified as in that
species, and erinipes has a linear malar space. By the white
face-marks and prominent clypeus the male resembles
A. dives, Dours, of which I have a specimen marked
“cotype”’ from Gribodo, but the legs are entirely different.
Friese makes dives a synonym of A, dufourii, Lep., but it is
possibly separable, the male (at least) having no metallic
colour on abdomen, the middle tarsal joints of middle leg
longer and slenderer than in Friese’s figure, and the brush
on last joint wider. They agree, however, in the remarkable
hind basitarsi.
Osmia (Ceratosmia) balucha, Nurse.
Quetta, India, March 1906 (Benton).
The male has the middle femora strongly produced and
angulate beneath, but the hind basitarsi are not dentate.
Bibliographical Notice— Geological Society. 291
BIBLIOGRAPHICAL NOTICE.
Catalogue of the Lepidoptera Phalene. Supplement, Vol. I.
London: the Trustees of the British Museum. 1914-15.
Since the publication of the first two volumes of the ‘ Catalogue of
Moths’ a formidable number of species in the families therein
included have since been described. Hence it became necessary
to prepare a supplement in order that the subject-matter of these
volumes might be brought up to date. The present volume, with a
smaller containing the plates, represents the first instalment of
that supplement.
Some idea of the number of species which have been added to
the lists may be gathered from Dr. Gahan’s Preface to Sir George
Hampson’s work. Thus, the family Amatide in vol. i. contained
169 genera and 1184 species, to which are now added 16 genera
and 945 species. The family Noline in vol. ii. had 13 genera and
162 species, to which are added 1 genus and 116 species; while
the Lithosiane in vol, ii. had 244 genera and 1055 species, to
which are added in this Supplement 73 genera and 880 species!
A supplementary volume to vol. ili. is in progress.
Whether all the species recognized in this Catalogue are really
“‘ eood species” is evidently a matter for debate, since the author,
in this Supplement, frequently admits of this or that new species
that it is ‘very possibly ”’ the male or female, or even a “ variety,”
of some other specific form.
PROCEEDINGS OF LEARNED SOCIETIES.
GEOLOGICAL SOCIETY.
December 6th, 1916.—Dr. Alfred Harker, F.R.S., President,
in the Chair.
Mr. G. C. Crick, A.R.S.M., F.G.S., gave an account of some
recent researches on the belemnite animal. He stated
that it was not his intention to deal that evening with the
homologies of the belemnite shell or with the phylogeny of the
belemnite group, but to confine himself to the restoration of a
typical belemnite animal and its shell, as shown particularly by
examples in the British-Museum collection.
He first demonstrated, by means of a rough model, the construc-
tion of the belemnite shell, including the guard or rostrum, the
phragmocone with its ventrally-situated siphuncle, and its thin
envelope the conotheca, with its forward prolongation and expan-
sion (on the dorsal side) known as the pro-ostracum. He then
292 Geological Society.
exhibited photographic slides of examples in the British-Museum
collection showing these various characters, and noted the abrupt
termination of the chambered cone on the lower part of the pro-
ostracum, of which the dorsal surface may have been partly or
almost completely covered by a thin forward extension of the
guard. To illustrate what was known of the complete body of
the animal as found associated with the guard, he then showed
photographic slides of two of the examples figured by Huxley
in his ‘ Memoir on the Structure of the Belemnitide ’ published in
1864. Each of these exhibited the guard associated with portions
of the pro-ostracum, the ink-bag, and the hooklets of the arms.
The form of the hooklets with their thickened bases was discussed,
this feature in a great measure justifying the attribution to the
belemnite of certain cephalopod remains (found practically at
about the same geological horizon) that included uncinated arms
associated with an ink-bag, and frequently also with nacreous
portions of (presumably) the pro-ostracum.
Of the remains of uncinated armed cephalopods from the Lias,
each exhibiting the same form of hooklets as those figured by
Huxley, he said that the British-Museum collection contained
seventeen examples, all from the neighbourhood of Lyme Regis and
of Charmouth, in Dorset. Each specimen exhibits a number of
uncinated arms associated usually with an ink-bag, sometimes also
with nacreous matter, and in two instances also with the guard or
rostrum. ‘These two examples were those to which he had already
referred as having been figured by Huxley, and unfortunately
the arms are not well preserved in either of these specimens ; in
one (B. bruguierianus, from the Lower Lias near Charmouth)
there are only a few scattered hooklets, while the arms of the
other (B. elongatus, from the Lower Lias of Charmouth) are
represented only by a confused mass of hooklets. Of the other
fifteen examples, in one there are a few solitary hooklets; in
another the number of the arms is very indistinct; in two the
remains of only two arms are preserved; in one there are traces
of three arms; in two there are indications of three, or possibly
four, arms; and in one there is a confused mass of possibly four
arms; and in one there are the remains of four, or possibly of five,
arms. In each of the remaining six specimens six arms can
be more or less clearly made out, while there is not a single
example in which more than six uncinated arms are displayed.
Of the six examples that exhibit six uncinated arms four are
stated to be from the Lias of Lyme Regis; one is from the Lias of
Charmouth ; and one was obtained from the Lower Liassic shales
between Charmouth and Lyme Regis. From a consideration of
these specimens, the speaker concluded that the cephalopod repre-
sented by these uncinated arms is the animal known as the
belemnite, and that the six uncinated arms were arranged in three
pairs of unequal length, of which the longest pair was lateral, the
medium-sized pair probably dorsal, and the shortest pair probably:
Geological Society. 293
ventral. He considered the presence of tentacular arms to be
doubtful. These observations were in accord with those of
Huxley, who, in his ‘Memoir’ already cited, stated that he had
‘not been able to make out more than six or seven arms in
any specimen, nor has any exhibited traces of elongated tentacula,
though the shortness of the arms which have been preserved would
have led one to suspect their existence.’
The speaker regarded certain markings sometimes to be seen on
the guard as indicating that during the life of the animal the
guard was almost, if not entirely, covered by the mantle, in which
case it was highly improbable that the guard was pushed into the
soft mud of the sea-bottom in order to act as an anchor.
He considered the animal to have been a free swimmer, swimming
forward ordinarily, but when desirable, capable also of sudden and
rapid propulsion backwards.
short discussion followed, and the thanks of the Fellows
present were accorded to Mr. Crick for his lecture.
December 20th, 1916.—Dr. Alfred Harker, F.R.S., President,
in the Chair.
Marte C. Sropgs, D.Sc., Ph.D., gave an account of some
recent researches on Mesozoic ‘Cycads’ (Bennetti-
tales), dealing particularly with recently-discovered petrified
remains which reveal their cellular tissues in microscopic prepara-
tions. ‘To make the significance of the various fossil forms clear,
Dr. Stopes first showed some lantern-slides of living Cycads, and
then pointed out that it was in their external features and in their
vegetative anatomy only that the fossil ‘Cycads’ were like the
living forms ; the most importa nt features, the reproductive organs,
differ profoundly in the two groups, and the fossils were funda-
mentally distinct, not only from the living Cyeads, but from all
other living or fossil famihes.
The fossils representing the group that are most frequently
found are (a) trunks, generally more or less imperfect casts or
partial petrifactions, and sometimes excellent petrifactions pre-
serving anatomical details and cell-tissues ; (0) 1 impressions of the
foliage. Not infrequent are the detached impressions of incomplete
‘flowers’ or cones, of one cohort (the Williamsonee), while
petrified fructifications are numerous in some of the well-petrified
trunks of the Bennettitez. The described species of the group
run into hundreds, but probably many of these duplicate real
species, because the foliage, trunks, pith-casts, various portions of .
the fructifications, etc., have often been separately found and
named. In very few cases have the different parts been correlated.
The species of the foliage are the most generally known, as they
are the most readily recognized with the naked eye; they have
been described under a variety of generic names.
294, Geological Society.
%
The following table gives the proved, or probable, associated
parts of some members of the group :—
Foliage. Trunk, Fructifications.
Zamites spp. Bennettites spp. Bennettites spp.
Zamites gigas. Attached, no separate Williamsonia gigas.
name.
Ofozamites'sp. ~ =. au when an eee Williamsonia spectabilis.
Ptilophyliwm pectinoides. ss esas Williamsonia whitbiensis.
Anomozamites minor. (Only slender branches Wielandiella angustifolia.
known, no name.)
Teniopterts wiibata. ~ 9 te anaes Williamsoniella coronata.
Dr. Stopes exhibited slides of microphotographs of the stem
and leaf-base anatomy of the group, including some unpublished de-
tails of Bennettites maximus. The roots of the group have hitherto
been entirely unknown, and a slide was exhibited for the first time
showing rootlets penetrating the leaf-bases of a petrified specimen
(represented by a section in the Geological Department of the
British Museum—Natural History). These roots probably belong
to B. saxbyanus: they are covered with wonderfully petrified
root-hairs, running uncollapsed through the silica matrix. They
raise interesting questions concerning the possible chemical con-
ditions of the infiltration of the silica. Illustrations were also
exhibited of the famous complex ‘ flower’ and cone-structures, and
of Wieland’s brilliant restorations of the same. Microphotographic
slides were exhibited of the seed-cone of an interesting unpublished
new species from the British Gault. This is beautifully petrified,
and adds to our knowledge of the finer anatomy of the seeds and
associated structures. Itisalso the largest cone of the Bennettitales
yet known, though it occurs in the Gault, by which time the group
appears to have begun rapidly to die out.
The following table indicates the distribution of a few of
the most interesting representatives of the Bennetti-
tales (including the cohorts Bennettitee and William-
sone) :—
Upper CRETACEOUS. Very fragmentary and uncertain records; apparently
the group is nearly or quite extinct.
Mipp.Le Cretacrous; The new large-sized seed-cone.
Gault. B. morierei 9 (? described originally from the Jurassic).
Lower Crevracrous; Well-petrified trunks with fructifications.
Lower Greensand, B. gibsonianus (type-species of the Bennettitez),
B. maximus. \ Throughout
these periods
Potton Sands. Trunks, e.g. Colymbetes edwardsi. | :
in America,
Wealden. Trunks (casts and petrifactions), | trunk-remains
foliage, very abun-
B. saxbyanus. dant, often
k : petrified and
Jurassic; Purbeck. Trunks (casts and semi-petrifac- | with fructifica-
tions). Se \ tions, parti-
Buckland’s original Cycadeoidea | eularly from the
Spp- Black Hills,
C. gigantea, South Dakota,
Geological Society. 295
Oolites. Trunks, pith-casts, ete. Much | and Maryland.
foliage of various types. Wil- C. Jenneyana,
liamsonia gigas and other fruit- | C. ingens,
impressions. ) GC. wielandi, ete.
W. scotica. Rich ; : ms
Wi Pail ronata. ich impressions i
. piss iamsoniella een a Wiese. Gk Witkin.
Lias, Foliage and _Williamsonia sonia and many
fruits (India). foliage genera.
Rhetie, Wielandiella angustifolia and foliage.
The group is by far the most characteristic of all the plants of
the Jurassic and Lower Cretaceous, during which periods its
distribution was almost world-wide. It was locally, if not univer-
sally, dominant, and was the most highly evolved plant-group of
the epoch of which we are cognizant.
Three chief points of interest are to be noted in the geological
distribution of these plants: (a) that the most numerous highly-
specialized trunks reach their maximum in the J urassic and Lower
Cretaceous Periods, when their distribution was practically world-
wide; () that the oldest and therefore presumably the most
primitive type, W 2elandiella, is externally less like the living eycads
than the commoner later forms, while these latter are utterly unlike
the living genera in their fructifications ; (ec) that the geologically
youngest cone is the largest yet discovered, occurring in the Gault
when the extinction of the group appears already to have set in.
Contrary to what might have been anticipated from their
external likeness to the living Cycads, coupled with their great
geological age, the fossil ‘Cycads’* are much more complex and on
a higher level of evolution than the living group: * It seems to the
Author to be extremely unlikely that the fossil and the living forms
have any direct phylogenetic connexion nearer than a remote,
unknown, common ancestor. The mooted connexion between the
fossil ‘ Cycads’ and the Angiosperms is highly suggestive, but lacks
data for its establishment.
A short discussion followed, and the thanks of the Fellows
present were accorded to Dr. Stopes for her lecture.
January 10th, 1917.—Dr. Alfred Harker, F.R.S., President,
in the Chair.
The following communication was read :-—
‘Balston Expedition to Peru: Report on Graptolites collected
by Capt. J. A. Douglas, R.E., F.G.S.’ By Charles Lapworth,
LL.D., M:Se., F.R.S., F.G.S.
The specimens of graptolites were collected from the rocks of
the Inambari district in Peru by Capt. Douglas, under whose name
296 Geological Soctety.
the collection has been placed in the Geological Department of
the University Museum, Oxford. These fossils were forwarded
by Prof. W. J. Sollas to Prof. C. Lapworth, who embodied the
results of his study in a Report, of which the following is a brief
abstract.
The specimens are recorded as all occurring in the same locality,
but it is not known whether they were obtained froma single zone.
The majority of the rock-specimens in which the graptolites oceur
are black and somewhat pyritous carbonaceous shales, usually well
bedded and uncleaved, and the graptolites are in general well
preserved. The lithology of the containing rocks and the mode of
preservation of the graptolites are similar to those obtaining in
the richest graptolite-bearing strata of Britain, Europe, and North
America.
The forms apparently represented in the collection are Logano-
graptus logant Hall, a new species of Gontograptus (7), Didymo-
graptus stabilis Elles & Wood and D. bifidus Hall, Phyllograptus
angustifolius Hall, Glossograptus acanthus Elles & Wood, Crypto-
graptus tricornis Hall, var., Amplexograptus confertus Lapworth,
and A. cwlatus Lapworth.
Taken as a whole, this graptolite fauna may best be compared
with that of the Upper Arenig formation of Britain and its North-
American equivalents, answering to the Lower Llanvirnian of
Hicks & Marr and the Didymograptus-bifidus Zone of Elles &
Wood and H.M. Geological Survey.
The assemblage of graptolites discovered in Bolivia a few years
ago by Dr. J. W. Evans corresponds very closely with this Peruvian
fauna, and was probably derived from the southward continuation
of the same Andean graptolite-band. The Peruvian forms in the
Douglas collection, like those from Bolivia, admit almost as close a
parallelism with those of the Arenig-Llandeilo graptolite-beds of
Australia and New Zealand as with their representatives in the
Northern Hemisphere.
Not only is the Douglas Collection of Peruvian graptolites
instructive and valuable from the paleontological point of view,
owing to the number and the good state of preservation of the
species represented, but it is of especial interest from the palzo-
graphical aspect, as affording additional proof of the identity
(in general facies) of the graptolite fauna of the sea-waters of
Lower Ordovician times in those regions of the globe which are
now oceupied by some of the dry lands of Britain, Eastern North
America, Peru, Bolivia, Victoria, and New Zealand. Thus it greatly
‘strengthens the inference that in Arenig-Llandeilo times there
was open-sea communication admitting of the circulation of sea-
currents along some as yet undetermined line or lines, connecting
the above-mentioned regions, which must have extended across the
Equator and apparently throughout a length nearly equal to that
of half the cireumference of the globe.
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XXVI.—A Revision of the Clupesid Fishes of the Genera
Pomolobus, Brevoortia and Dorosoma, and their Allies.
By C. Tate Reagan, M.A.
(Published by permission of the Trustees of the British Museum.)
THE genera dealt with in this revision are the Clupeinz with
a distinct notch in the middle of the upper jaw ; these have
usually been placed in two distinct groups—those with
terminal mouth and the last dorsal ray not prolonged being
associated with Clupea, and the others, with inferior mouth
or last dorsal ray prolonged, forming a group apart (Chato-
essine of Giinther, Dorosomatide “of modern authors) ; in
my judgment this is quite an artificial arrangement. All
these fishes appear either to be ean y entering rivers to
spawn in fresh or brackish water*, or are permanently
fluviatile (e. g., Gudusia, Signalosa).
Synopsis of the Genera.
I. Gill-rakers of epibranchial of first arch folding downwards, those
near the angle overlapping the gill-rakers of the ceratobranchial.
A. Seales with edges entire or feebly serrated ; normal scales from
occiput to dorsal fin; pelvic fins 9-ray ed; ; operculum with
radiating grooves.
Teeth on vomer and palatines (rarely deciduous
in adultsye sae 2 anes, < eet he Cre pEpOor Oe « IL. Caspialosa.
* The breeding-habits of Ethmidium and Ethmalosa are unknown
to me.
Ann, & Mag. N. Hist, Ser. 8. Vol. xix. 20
298 Mr. C. T. Regan on the Clupeoid Fishes
Palate toothless; lower jaw not prominent, its
dapancluded ......'.. ...24s= ee ee ee eee 2. Alosa,
Palate toothless; lower jaw projecting ........ 3. Pomolobus.
B. Scales with edges serrated in young, pectinated in adults ; pelvic
fins 7-rayed.
A well-defined series of pectinated scales on each
side of middle line from occiput to dorsal
fin; operculum striated or nearly smooth .. 4. Brevoortia.
A median series of scutes from occiput to dorsal
fin ; operculum smooth or very feeblystriated. 5. Ethmidiwm.
IT. Gill-rakers of epibranchial of first arch not folding downwards over
those of ceratobranchial ; pelvic fins 8-rayed; operculum smooth.
A. Edge of dentary not reflected outwards in front of maxillary.
1, Last dorsal ray not prolonged.
Upper gill-rakers of first and second arches and
all of succeeding arches bent or expanded,
T-shaped or triangular in section .......... 6. Ethmalosa,
Gill-rakers normal: scales large, 40-50/13-20 .. 7. Hilsa.
Gill-rakers normal ; scales small, 75-100/27-34 . 8. Gudusia.
2. Last dorsal ray prolonged into a filament.
Mouth terminal or subterminal; maxillary nor-
mal, with one supramaxillary ............ 9. Clupanodon.
Mouth terminal; maxillary normal, with two
SULPPAMlexTNAPiOn s..%lnw. pix sie 0's wie ahete wine» 10. Signalosa.
Mouth subterminal or inferior; maxillary slender,
with one supramaxillary ..............5> 11. Dorosoma.
B. Edge of dentary reflected outwards in front of extremity of
maxillary ; mouth toothless, subterminal or inferior, transverse,
its cleft forming an angle: one supramaxillary.
Maxillary slender, distally slightly expanded and
curved downwards ; last dorsal ray produced
into.a filaments si6 |. ie sic say. is abe Sea gs bones 12. Nematalosa.
Maxillary slender, distally slightly expanded and
curved downwards ; last dorsal ray not pro-
Guna 8 Sie FE rch LS cin cisseeus eee hire sae eee 13. Gonialosa.
Maxillary a straight, thin, transversely expanded
lamina, tapering distally ; last dorsal ray not
produced 2. dias a ynktsis SAniahaen et eee 14. Anodontostoma.
1. CaspraLosa, Berg, 1915.
Clupeonella (non Kessler), Berg, Ann. & Mag. Nat. Hist. (8) xi. 1913,
p. 472.
Caspialosa, Berg, Poiss. de l’eau douce de la Russie, p. 22 (1916).
Differs from Alosa in having three patches of teeth on the
palate, borne by the vomer and palatine bones; but in large
examples of C. caspia I find that the palate is toothless.
Black and Caspian Seas.
Berg recognizes thirteen species of this genus.
of the Genera Pomolobus, Brevoortia, cc. 299
2. ALosa, Cuv. 1829.
Régne Animal, ed. 2, ii. p. 319; Regan, Ann. & Mag. Nat. Hist. (8)
xviii. 1916, p. 6.
North Atlantic and Mediterranean.
In my revision five species and six subspecies were
recognized,
3. Pomonosus, Rafin. 1820.
Ichth. Ohiensis, p. 38; Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1896, p. 424.
Distinguished from Alosa by the prominent lower jaw,
with its tip not included.
Western North Atlantic.
Synopsis of the Species.
I, Lower jaw strongly projecting ; 20 to 25 gill-rakers on lower part
of anterior arch.
Anterior teeth persistent; maxillary extending to
below posterior part of eye; caudal peduncle
TOMTSP UMP RECON 5.2 eet oath apie. ciao Phe 1. chrysochloris.
Jaws’ toothless; maxillary extending to below
middle of eye; caudal peduncle as long as deep. 2. mediocris.
II. Lower jaw a little projecting ; 40 to 50 gill-rakers on lower part of
anterior arch. j
Depth 33, head 42 in the length; eye 4: to 5in
head (in specimens of 220-260 mm.) ....... . 9d. @estivalis.
Depth 3, head 4 to 43 in the length; eye 33 to 4 in
head (in specimens of 220-260 mm.) ..... vee 4, pseudoharengus.
1. Pomolobus chrysochloris.
Pomolobus chrysochloris (Rafin. 1820), Jord. & Everm. Bull, U.S. Nat.
Mus. xlvii. 1896, p. 425, and 1900, fig. 187, sd
Depth of body 32? in the length, length of head 4. Snout
longer than diameter of eye, which is 6 in length of head;
maxillary extending to below posterior part of eye; lower
jaw strongly projecting; small conical teeth persistent in
premaxillaries and anterior part of lower jaw; 23 gill-rakers
on lower part of anterior arch. 56 scales in a longitudinal
series, 17 in a transverse series; ventral scutes 21415.
Dorsal 18. Anal 18. Pelvics a little in advance of middle
of dorsal. Caudal peduncle longer than deep. Silvery;
back darker.
Mississippi and southern coast of U.S.A.
A single specimen, 280 mm. long, from Pensacola.
20*
300 Mr. C. T. Regan on the Clupeoid Fishes
2. Pomolobus mediocris.
Pomolobus medioeris (Mitchill, 1815), Jord. & Everm. Bull. U.S. Nat.
Mus. xlvii. 1896, p. 425, and 1900, fig. 188.
Depth of body 3} to 3? in the length, length of head 4 to
41, Snout longer than diameter of eye, which is 5 in length
of head ; maxillary extending to below middle of eye or a
little beyond ; lower jaw strongly projecting ; jaws toothless ;
21 or 22 gill-rakers on lower part of anterior arch. 56 scales
in a longitudinal series, 17 in a transverse series ; ventral
scutes 21-22+16. Dorsal 16-18. Anal 20-22. Pelvics
in advance of middle of dorsal. Caudal peduncle as long
as deep. Silvery; back darker ; each scale on sides with a
dark spot.
Atlantic coast of U.S.A.
Three specimens, 280 to 800 mm. long, from the Potomac
and Woods Hole.
3. Pomolobus estivalis.
Pomolobus estivalis (Mitchill, 1815), Jord. & Everm. Bull. U.S. Nat.
Mus. xlvii. 1896, p. 426, and 1900, fig. 190.
Depth of body 33 in the length, length of head 42. Snout
a little longer than diameter of eye, which is 43 to 5 in the
length of head ; maxillary extending to below anterior part
or middle of eye; lower jaw a little projecting ; jaws tooth-
less ; 44 to 47 gill-rakers on lower part of anterior arch. 52
to 55 scales in a longitudinal series, 15 or 16 in a transverse
series ; ventral scutes20+14. Dorsal 17-18. Anal 18-20.
Pelvics below anterior half of dorsal. Caudal peduncle
longer than deep. Silvery ; back darker.
, Atlantic coast of U.S.A.
Two specimens, 220 and 260 mm. in total length.
4. Pomolobus pseudoharengus.
Pomolobus pseudoharengus (Wilson, c. 1811), Jord. & Evyerm. Bug
U.S. Nat. Mus. xlvii. 1896, p. 426, and 1900, fig. 189.
Depth of body 3 in the length, length of head 4 to 4}.
Snout as long as or shorter than diameter of eye, which is 3}
to 4 in the length of head; maxillary extending to below
middle of eye or a little beyond; lower jaw a little pro-
jecting ; jaws toothless ; 40 to 42 gill-rakers on lower part of
anterior arch. 52 to 56 scales in a longitudinal series, 15
to 17 in a transverse series; ventral scutes 20-21+ 12-14.
3
of the Genera Pomolobus, Brevoortia, de. 301
Dorsal 16-18. Anal 18-22. Pelvics below anterior half
of dorsal. Caudal peduncle as long as deep, or deeper than
long. Silvery ; back darker.
Atlantic coast of U.S.A.
Seven specimens, 220 to 260 mm. in total length.
4, Brevoorti, Gill, 1861.
Proc, Ac. Philadelphia, p. 37; Jord. & Everm. Bull. U.S, Nat. Mus.
xlvii. 1896, p. 483.
This genus is distinguished from Alosa by the pectinated
scales, ‘Vhe gill-rakers are very numerous, long and slender.
1. Brevoortia tyrannus.
Clupea tyrannus, Latrobe, Trans. Amer, Phil. Soc. v. 1872, p. 77, pl. i.
Clupanodon aureus, Agassiz, Spix, Pisc. Brasil. p. 52, pl. xxi. (1828).
Clupea menhaden (Mitchill, 1815), Gunth. Cat. Fish. vii. p. 436 (1878).
Clupea aurea, Giinth, t. c. p. 487.
Brevoortia tyrannus, Goode, Rep. U.S. Fish. Comm. 1877, p, 19,
pls. 1., 1. (1879); Jord, & Everm. Bull. U.S. Nat. Mus, xlvii. 1896,
p. 4933.
Brevoortia patronus, Goode, t. c. p. 26, pl. v.
Depth of body 2} to 34 in the length, length of head 2% to
3}. About 70 gill-rakers (in the adult) on ceratobranchial
of first arch. About 50 scales in a longitudinal and 25 in a
transverse series ; in adults scales very deep, two often
meeting across one of the intermediate longitudinal series,
thus increasing the number of transverse rows ; ventral scutes
18-20+11-12. Dorsal 18-21. Anal 19-24. Pelvics
below or in advance of anterior rays of dorsal. Vertebre 48.
Nova Scotia to Gulf of Mexico and Brazil.
Several examples up to 350 mm. in total length ; one of
these, from Alabama, and therefore B. patronus, is exactly
similar to the specimen 8 inches long, from Woods Hole,
figured by Goode. Giinther’s example named Clupea aurea
is without locality, and may be North American ; the distri-
bution ot this species on the coast of South America has yet
to be made out.
2. Brevoortia pectinata.
Alosa pectinata, Jenyns, Zool, ‘ Beagle,’ Fish. p. 135, pl. xxv. (1842).
Clupea pectinata, Gunth. Cat. Fish. vil. p. 457 (1868).
Brevoortia pectinata, Goode, Rep. U.S. Fish, Comm. 1877, p. 80, pl. vi.
(1879).
Depth of body 23 to 22 in the length; length of head 3 to
34. Gill-rakers more numerous than in B. tyrannus, about
302 Mr. ©. T. Regan on the Clupeoid Fishes
90 on ceratobranchial of first arch. Scales more regularly
arranged and not so deep as in B. tyrannus; about 48 in a
longitudinal series, 20 to 25 in a transverse series ; ventral
scutes 18-20+10-12. Dorsal 17-19. Anal 18-22, Pelvics
below or in advance of origin of dorsal. Wertebree 44.
Northern Patagonia to Southern Brazil.
Here described from the types, skins, 180 and 260 mm.
in length (the larger kindly sent to me for examination by
C. Forster Cooper, Esq.), and from four examples of 220 mm.
from Rio Grande do Sul.
5. Erumrpium, Thompson, 1916.
Proc. U.S. Nat. Mus. 1. p. 458.
Closely related to Brevoortia, but with a median series of
scutes from occiput to dorsal fin.
Ethmidium maculatum.
Alausa maculata, Cuv. & Val. Hist. Nat. Poiss. xx. p. 480 (1847).
Alausa cerulea, Cuv. & Val. t. c. p. 482.
Clupea notacanthus, Giinth. Cat. Fish, vii. p. 443 (1868),
Clupea maculata, Giinth. 1. e.
Clupea (Alosa) notacanthoides, Steind. Sitzungsb. Akad. Wien, 1x.
1869, p. 809, pl. vii.
gen notacanthoides, Thompson, Proc. U.S. Nat. Mus. 1. 1916,
. 458.
Ebimidium cerulea, Thompson, t. c. p. 460.
Depth of body in the adult equal to length of head, 3 in
length of fish ; in the young head relatively shorter and body
deeper. Diameter of eye 4 to 7 in length of head ; maxillary
extending to below posterior part of eye or beyond. 80 (young)
to 160 (adult) gill-iakers on lower part of anterior arch. 24
to 28 scutes from occiput to dorsal fin; about 50 scales in a
longitudinal, 17 to 20 in a transverse series; ventral scutes
18-20+15-17. Dorsal 19-22. Anal 14-18. Pelvics
below anterior $ of dorsal. Vertebize 50. Silvery ; back
bluish ; sometimes a lateral series of dark spots.
Peru and Chile.
Four specimens, 100-280 mm. long, from Callao, Val-
paraiso, and Helladura Bay.
; 6. ETHMALOSA, gen. nov.
Form rather deep and strongly compressed. Upper jaw
with median notch ; lower jaw included ; teeth minute or
absent. Adipose eyelid well developed ; cheek moderately
deep, with a naked area below the suborbitals. Operculum
of the Genera Pomolobus, Brevoortia, Sc. 303
smooth except for a groove parallel to its anterior edge ; sub-
operculum tapering upwards; opercular margin rounded ;
6 branchiostegals. Lower gill-rakers of first and second
arches long, slender, and numerous, those of ceratobranchial
folding over those of epibranchial, which are curiously
expanded, T-shaped or triangular in section, appearing angu-
larly bent on the lower side, but not on the upper; gill-rakers
of third and fourth arches similarly expanded or recurved,
the series fitting closely to form a sieve. About 45 scales in
a longitudinal and 16 to 19 in a transverse series ; edges of
scales crenulated in the young, pectinated in the adult ;
transverse grooves paired, not meeting in the middle of the
scale, only the most posterior groove extending right across ;
a well-defined mid-dorsal double row of scales, commencing
with a large postoccipital pair, extends backwards to the
dorsal fin ; ventral scutes with sharp-pointed keels. Dorsal
fin of 16 to 19 rays ; a very low basal sheath. Anal of 20
to 23 rays. Pelvics 8-rayed, inserted below anterior 4 of
dorsal. Caudal with alar scales.
Ethmalosa dorsalis.
Meletta senegalensis, Cuy. & Val. Hist. Nat. Poiss, xx. p. 870 (1847).
Alausa dorsalis, Cuv. & Val. t. ¢. p. 418.
Alosa platycephalus, Bleek. Verh, Holl. Maatsch. Haarlem, 1862,
Guinée, p. 123, pl. xxvi. tig. 2.
Clupea dorsalis, Gunth. Cat. Fish. vii. p. 458 (1868),
Clupea setusa, Steind, Sitzungsb. Akad. Wien, lx, 1869, p, 311, pl. vi.
Depth of body 24 to 3 in the length, length of head 3 to 32.
Diameter of eye 43 to 6 in length of head. Maxillary ex-
tending to below middle or posterior part of eye. About 45
‘scales in a longitudinal, 16 to 19 in a transverse series ;
ventral scutes 16-19+4+11-13. Dorsal 16-19. Anal 20-23.
Pelvics below anterior 4 of dorsal. Silvery ; back darker ;
tip of dorsal fin blackish. ;
West Africa.
Numerous examples up to 300 mm. in total length.
7. HILSA, gen. nov.
Paralosa (non Bleek.), Regan, Ann. Durban Mus, i. 1916, p. 167.
Distinguished from A/osa by the smooth operculum and
the different arrangement of the gill-rakers of the anterior
arch, from Hthmalosa by the normal structure of the gill-
rakers, and from both by the absence of alar scales on the
caudal fin.
304 Mr. C. T. Regan on the Clupeotd Fishes
Coasts and rivers from Natal to China.
In the young the body is deeper and the head smaller than
in the adults, the greater length of the head in the latter
being mainly due to the size of the operculum.
Synopsis of the Species.
I, Parietal ridges expanded and striated.
A. Depth 23 to 3 in the length.
Head. 3 to 32 Ain the length) ety a Bae ee Ae Eerie 6 1. kanagurta.
Head 32-to 32 in the length ...0. 00s cves ce eee ree. 2, durbunensis.
B, Depth; 2} in the lengthy. ot), les y bee eels 3. brachysoma.
II. Parietal ridges narrow, covered by smooth skin.
A. Maxillary extending to below middle of eye (young) or beyond.
1. Caudal lobes as long as head.
Opereulum 3 to 2 as broad asdeep; scales 45-48/17-20. 4. alisha.
Operculum 2 to 2 as broad as deep; scales 42-45/16-17. 5. reevesi.
2. Caudal lobes longer than head.
Operculum 3 to 2 as broad as deep; scales 40/14-15 .. 6. toli.
B. Maxillary not reaching middle of eye; caudal lobes much longer
than head; scales 45/14-15............006. 7. macrura,
1. Tlilsa kanagurta.
Alosa kanagurta, Bleek. Verh. Bat. Gen. xxiv. 1852, Haringaeht.
p- 84; Atl. Ichth. vi. p. 114, Clup. pl. vii. fig. 6 (1872).
Alosa malayana, Bleek. Ned. Tijdschr. Dierk. iii. 1866, p. 294; Atl.
Ichth. vi. p. 114, Clup. pl. vii. fig. 4.
Clupea tlisha, Giinth. Cat. Fish. vii. p. 445 (1868).
Clupea kanagurta, Day, Fish. India, p. 640, pl. clxii. fig. 4; Weber &
Beaufort, Fish. Indo-Austral. Arch. 1. p. 67 (1918).
Depth of body 23 to 3 in the length, length of head 3 to
32. Parietal ridges expanded and striated. Snout nearly as
long as or a little longer than diameter of eye, which is 32
to 44 in length of head ; maxillary extending to below middle
or posterior part of eye ; width of operculum 4% or less than $
its depth ; 100 to 150 gill-rakers on lower part of anterior
arch. 42 to 45 scales in a longitudinal series, 13 or 14 ina
transverse series; ventral scutes 16-18+11-13. Dorsal
17-20. Anal 19-22. Pelvics below anterior half of dorsal.
Caudal about as long as head. A dark humeral spot, in the
young followed by a series.
Zanzibar to Malay Archipelago.
Fifteen examples, up to 220 mm. in total length.
of the Genera Pomolobus, Brevoortia, &c. 305
2. Hilsa durbanensis.
Clupea durbanensis, Regan, Ann. Natal Govt. Mus. i. 1906, p. 4,
pl. iv.; Gilchrist, S. Afr. Mar. Biol. Rep, i. 1918, p. 59.
Depth of body 23 to 3 in the length, length of head 32 to
32. Parietal ridges expanded and striated. Snout as long
as or slightly longer than diameter of eye, which is 4 to 44
in length of head; maxillary extending to below middle or
posterior part of eye; width of operculum ? its depth; 150
gill-rakers on lower part. of auterior arch. 42 to 44 scales in
a longitudinal, 13 or 14 in a transverse series; ventral scutes
16-17+12-13. Dorsal 17-18. Anal 19-21. Pelvics
below anterior half of dorsal. Caudal fin about as long as
head. A dark humeral spot. Upper edge of dorsal and
posterior edge of caudal blackish.
Natal.
Three specimens from Durban, 140 to 200 mm. in total
length.
Gilchrist has examined a large example, 240 mm. long to
base of caudal fin ; in this the head is 32 in the length, and
there are 200 gill-rakers on the lower part of the anterior
arch.
3. LHilsa brachysoma.
? Alusa brevis, Bleek. J. Ind. Arch. ii. no. 9, 1848, p. 638; Atl. Ichth.
vi. p. 116 (1872).
Alosa brachysoma, Bleek. Nat. Tijdschr. Ned. Ind. v. 1853, p. 527;
Atl. Ichth. vi. p. 115, Clup. pl. iv. tig. 5 (1872).
Clupea platygaster, Giiuth. Cat. Fish. vii. p. 448 (1868) ; Weber &
Beautort, Mish. Indo-Austral. Arch. ii. p. 66, fig. 24 (1918).
Depth of body 2} in the length, length of head 32.
Parietal ridges expanded and striated. Snout as long as
diameter of eye, which is 4 in length of head; maxillary
extending beyond middle of eye ; width of operculum 2 its
depth; 100 gill-rakers on lower part of anterior arch. 42
scules in a longitudinal, 15 in a transverse series; ventral
scutes 17412. Dorsal 17-18. Anal 20-21. “Pelvics below
middle of dorsal. Caudal about as long as head. A dark
humeral spot ; dorsal and caudal dark-edyed.
Sumatra.
Here described from Bleeker’s type and only specimen,
which has the head and body deeper and the lower jaw longer
ete. than in A. kanagurta of this size (120 mm.) ; also the
first mid-dorsal post-cephalic scale is a striated bony plate.
Weber and Beaufort’s figure is of a fish that agrees with
Bleeker’s in the appearance of the head, but is more elongate
in form, the depth about 23 in the length.
306 Mr. C. T. Regan oa the Clupeoid Fishes
4. Hilsa ilisha.
Clupanodon tlisha, Ham. Buch, Fish. Ganges, p. 243, pl. xix. fig. 75.
Alosa palasah, Cuv. & Val. Hist. Nat. Poiss, xx. p. 482 (1847).
Clupea palasah, Giinth. Cat. Fish. vii. p. 445 (1868).
Clupea ilisha, Day, Fish. India, p. 640, pl. clxxii. fig. 3 (1878).
Depth of body 23 to 3 in the length, length of head 33 to
33. Parietal ridges narrow, covered with smooth skin in the
adult fish. Snout as long as or longer than diameter of eye,
which is 42 to 7 in the length of head ; maxillary extending
to below posterior part of eye or beyond ; width of operculum
from a little more than } to 4 of its depth ; 120 (young) to
220 gill-rakers on lower part of anterior arch. 45 to 48
scales in a longitudinal and 17 to 20 in a transverse series ;
ventral scutes 17-19+13-14. Dorsal 18-20. Anal 18-21.
Pelvic fins below anterior part of dorsal. Caudal fin about
as long as the head. Vertebre 47.
Persian Gulf to Burma.
Several specimens, 130 to 350 mm. in total length.
5. Hilsa reevestt.
Alosa reevesii, Richards, Ichth. China, p. 305 (1846).
Alosa palasah, Richards, ¢. c. p. 306.
Clupea reevesti, Giinth. Cat. Fish. vii. p. 446 (1868).
Depth of body 3 to 34 in the length, length of head 3 to
31. Upper surface of head covered with skin ; no striated
bones exposed, except in the young. Snout longer than
diameter of eye, which is 5 to 9 in length of head; maxillary
extending to below posterior part or edge of eye, or a little
beyond ; width of operculum 2 or more than 2 of its depth ;
gill-rakers long and slender. 150 (young) to 250 on lower
part of anterior arch. 42 to 45 scales in a longitudinal, 16 or
17 in a transverse series; ventral scutes 18+13-14. Dorsal
17-18. Anal 18-19. Pelvics below anterior half of dorsal.
Caudal fin about as long as the head.
China.
Seven examples, 150 to 500 mm. long, from Shanghai and
Kiu Kiang.
6. Hilsa toli.
Alosa toli, Cuv. & Val. Hist. Nat. Poiss. xx. p. 485 (1847); Bleek.
Atl. Ichth. vi. p. 118, Clup. pl. vili. fig. 4 (1872).
Alosa ctenolepis, Bleek. Verh. Bat. Gen. xxiv. 1852, Haringacht. p. 32.
Clupea toli, Giinth. Cat. Fish. vii. p. 447 (1868); Day, Fish. India,
p- 641, pl. clxii. fig. 2 (1878); Weber & Beaufort, Fish. Indo-
Austral. Arch. ii. p. 64 (1913).
Clupea chapra, Giinth. l. ¢.
oj the Genera Pomolobus, Brevoortia, &e. 307
Depth of body 22 to 33 in the length, length of head 33 to
4, Parietal ridges narrow, covered with smooth skin in the
adult fish. Snout as long as or longer than diameter of eye,
which is 42 to 74 in the length of head ; maxillary extending
to below posterior part of eye or beyond; width of operculum
from 4 to nearly 2 of its depth ; 70 to 95 gill-rakers on lower
part of anterior arch. About 40 scales in a longitudinal and
14 or 15 in a transverse series ; ventral scutes 17-184 11-13.
Dorsal 17-19. Anal 18~21. Pelvics below middle of dorsal.
Caudal lobes, in the adult fish, nearly 14 as long as head.
India, Malay Peninsula and Archipelago.
Several examples, 120 to 450 mm. in total length.
7. Hilsa macrura.
Alosa macrurus, Bleek. Verh. Bat. Gen. xxiv. 1852, Haringacht. p. 31;
Atl. Ichth. vi. p. 118, Clup. pl. vi. fig. 4 (1872),
Clupea macrura, Giinth. Cat. Fish. vii. p. 448 (1868) ; Weber & Beau-
fort, Fish, Indo-Austyral. Arch. ii. p. 65 (1918).
Depth of body 22 to 32 in the length, length of head 4 to
42, Parietal ridges narrow, covered with smooth skin in the
adult fish. Snout not longer than diameter of eye, which is
4 to 5 in the length of head; maxillary extending to below
anterior 4 of eye ;.width of operculum 4 its depth; 60 to 80
gill-rakers on lower part of anterior arch. About 45 scales
in a longitudinal and 14 or 15 in a transverse series ; ventral
scutes 16-18+11-15.. Dorsal 17-20. Anal 18-21. Pelvics
below middle or anterior part of dorsal. Caudal lobes, in the
adult fish, nearly twice as long as head,
Sunda Islands. :
Three examples, 160 to 350 mm. in total length.
8. GubusiA, Fowler, 1911.
Proce. Acad. Philadelphia, lxiii. p. 207.
Distinguished from H/sa by the smaller scales.
‘T'wo species from the rivers of India and Burma.
1. Gudusia chapra.
Clupanodon chapra, Ham. Buch, Fish, Ganges, p. 248.
Climea indica, Gray, Ill. Ind. Zool.; Giinth. Cat. Fish. vii. p. 444
(1868).
Alausa microlepis, Cuv. & Val. Hist. Nat. Poiss. xx. p. 439 (1847).
Clupea chapra, Day, Fish. India, p. 639, pl. clxi. fig. 1 (1878).
Chepeas are Chaudhuri, Ree. Ind. Mus. vii. 1912, p. 439, pl. xxviii
euls
Depth of body 22 to 3} in the length, length of head 3 to
33- Snout shorter than diameter of eye, which is 4in length
308 Mr. C. T. Regan on the Clupeoid Fishes
of head; maxillary extending to below anterior part or
middle of eye; 200 or more gill-rakers on lower part of ante-
rior arch. 75 to 100 scales in a longitudinal and 27 to 34 in
a transverse series; ventral scutes 18-20+8-10. Dorsal
14-16. Anal 20-24. Pectoral 13-14. Pelvics nearly
below origin of dorsal. Usually a dark humeral spot, some-
times followed by a series.
Northern India, from Sind to Assam.
Several examples, 120 to 140 mm. long.
2. Gudusia variegata.
Clupea variegata, Day, Proc. Zool. Soc. 1869, p. 263; Fish. India,
p- 689, pl. clxi. fig. 4.
Depth of body 23 in the length, length of head 33.
Snout a little shorter than diameter of eye, which is 43 in
Jength of head ; maxi!lary extending to below middle of eye;
more than 200 gill-rakers on lower part of anterior arch.
90 scales in a longitudinal and 32 in a transverse series ;
ventral scutes 19+10. Dorsal 16. Anal 23. Pectoral 14.
Pelvics nearly below origin of dorsal. Back with several
vertically expanded dark spots.
Burma.
A single specimen, 166 mm. in total length.
According to Day, this species differs from G. chapra in
the deeper body, the coloration, and 24 to 29 instead of 21
to 24 anal rays.
9. CLuPANODON, Lacep. 1803 *.
Hist. Nat. Poiss. v. p. 468; Bleek. Atl. Ichth. vi. p. 112 (1872).
Konosirus, Jord. & Snyder, Proc. U.S. Nat. Mus, xxiii. 1900, p. 349.
Mouth toothless, terminal or subterminal, with lateral
* Of the six species placed by Lacepéde in Clupancodon, Jordan (in
collaboration) has at different times regarded as the genotype: 1. jussteut
(by designation), 2. pilchardus (by elimination), and 8. thrissa (the first
species). But before this, in 1872, Bleeker (Atl. Ichth. vi. p. 112) stated
that Clupanodon thrissa was the type of Lacepéde’s genus. Reference to
Lacepéde’s work leaves no doubt that his species was the true Clupea
thrissa of Linneus, who took the name and the diagnosis respectively
from Osbeck and from Langerstrom. Lacepéde took the specitic name
from Linnzus, and gave China as the first locality; his description of
the pointed lower jaw and the notched upper jaw, and his statement
that the fish spawns in fresh water, apply quite well to the Chinese
species, but not to the Antillean species (Opisthonema oglinum), which
so many of the older writers believed to be the same fish. Consequently
I regard the Clupea thrissa of Osbeck, Linneus, and Lacepéde, and not
the Clupea thrissa of Bloch and of Giinther, as the type of Clupanodon.
of the Genera Pomolobus, Brevoortia, cc. 309
cleft ; maxillary normally formed, extending to below ante-
rior part or middle of eye ; anterior supramaxillary absent.
Gill-rakers slender, very numerous. Dorsal 15-18 ; last ray
produced into a filament. Anal 20-28. Pelvics 8-rayed,
below anterior part of dorsal. 48 to 58 scales in a longi-
tudinal series, 20-23 in a transverse series. Vertebre 51
(in C. punctatus).
Coasts and rivers of China and Japan.
1. Clupanodon thrissa.
Clupea thrissa, Osheck, Iter Chinensis, p. 257 (1757); Linn. Syst. Nat.
ed, 10, p. 318 (1758).
Clupanodon thrissa, Lacep. Hist. Nat. Poiss. v. p. 468 (1803).
Chatoessus maculatus, Richards, Ichth. China, p. 308 (1846) ; Giinth.
Cat. Fish. vii. p. 409 (1868).
Chatoessus osbeckt, Cuy. & Val. Hist, Nat. Poiss. xxi. p. 106 (1848).
Depth of body 22 to 3 in the length, length of head 34.
Diameter of eye 44 to 5 in length of head. Mouth terminal ;
maxillary extending to below anterior part or middle of eye.
48 scales in a longitudinal, 20 in a transverse series ; ventral
scutes 18-20+4+10-12. Dorsal 15-16. Anal 22-27. Pelvics
below anterior 4 of dorsal. A dark humeral spot, sometimes
followed by a series of spots.
China; Formosa.
‘Three specimens of 150-200 mm. from Formosa ; two from
China, 60 and 90 mm., are not included except for counts of
fin-rays etc.
2 Clupanodon punctatus.
Chatoessus punctatus, Schlegel, Faun. Japon., Poiss. p. 240, pl. cix.
fig. 1 (1846); Cuv. & Val. Hist. Nat. Poiss. xxi. p. 107 (1848) ;
Giinth. Cat. Fish. vii. p. 408 (1868).
Chatoessus aquosus, Richards, Ichth. China, p. 307 (1846); Cuv. & Val.
Hist. Nat. Poiss. xxi. p. 109 (1848).
es punctatus, Jord. & Herre, Proc. U.S. Nat. Mus, xxxi. 1906,
p. 624,
Depth of body 3 to 33 in the length, length of head 32 to
43. Diameter of eye 43 to 5 in length of head. Mouth
subterminal ; maxillary extending to below anterior part or
nearly to middle of eye. 53 to 58 scales in a longitudinal,
20 to 23 ina transverse series ; ventral scutes 18-21 + 14-17.
Dorsal 16-18. Anal 20-25. Pelvics below anterior 4 of
dorsal. A dark humeral spot ; a dark spot on each scale of
upper half of body. Vertebre 51.
China; Japan.
Kleven specimens, 150-200 mm. in total length.
‘310 Mr.C.T. Regan on the Clupeoid Fishes
10. Sranatosa, Everm. & Kendall, 1898.
Bull. U.S. Fish. Comm. 1897, p. 127.
Month toothless, terminal, with lateral cleft; maxillary
normally formed, extending to below anterior edge of eye or
a little beyond; two supramaxillaries. Giill-rakers slender,
very numerous. Dorsal 13-16; last ray produced into a
filament. Anal 21-27. Pelvies 8-rayed, below or a little in
advance of origin of dorsal. About 40 scales in a longitudinal
series. Vertebre 41. :
Rivers from Southern U.S.A. to Central America.
1. Signalosa mexicana.
Chatoessus mexicanus, Giinth. Cat. Fish. vii. p. 409 (1868).
Dorosoma mexicanum, Jord. & Everm, Bull. U.S. Nat. Mus. xlvii.
1896, p. 416.
Signalosa atchafalaye, Jord. & Everm. ¢. e. 1898, p. 2809, fig. 184.
Signalosa mexicana, Meek, Zool. Publ. Columbian Mus. vy. 1904, p. 94.
Depth of body 22 to 3 in the length, length of head 3 to
33. Diameter of eye 33 to 4 in length of head; maxillary
extending to below anterior edge ort of eye. 200 gill-rakers
on lower part of anterior arch. About 40 scales in a longi-
tudinal series ; ventral scutes 16-18+8-10. Dorsal 13-15;
origin equidistant from end of snout and base of caudal, or
nearer former. Anal 23-27. Pelvics inserted a little in
advance of origin of dorsal. A dark humeral spot.
Louisiania to Central America, in rivers emptying into the
Gulf of Mexico.
Nine specimens, 70 to 100 mm. in total length.
2. Signalosa petenensis.
Meletta petenensis, Giinth. Proc. Zool. Soc. 1866, p. 603.
Chatoessus petenensis, Giinth, Cat. Fish. vii. p. 408 (1868).
Depth of body 3 in the length, length of head 3} to 33.
Diameter of eye 34 to4 in length of head; maxillary ex-
tending to below anterior ¢ or edge of eye. 160 gill-rakers
on lower part of anterior arch. About 40 scales in a longi-
tudinal series; ventyal scutes 14-164+10-12. Dorsal 15-16;
origin nearer to end of snout than to base of caudal. Anal
21-24. Pelvics inserted below origin of dorsal. A dark
humeral spot.
Lake Peten.
Four specimens, 70 to 90 mm. in total length,
of the Genera Pomolobus, Brevoortia, cc. Sin
11. Dorosoma, Rafin. 1820.
Ichth. Ohiensis, p. 39.
Chatoessus (part.), Cuv. Régne Anim. ed. 2, ii. p. 320 (1829); Gtinth.
Cat. Fish. vii. p. 406 (1868).
Chatoessus, Cuy. & Val. Hist. Nat. Poiss, xxi. p. 94 (1848).
Mouth subterminal or inferior ; maxillary narrowed dis-
tally ; anterior supramaxillary absent. Gill-rakers slender,
very numerous. Dorsal 12-15; last ray produced into a
filament. Anal 26-38. Pelvics 8-rayed, in advance of
dorsal. 55 to 80 scalesin a longitudinal series. Vertebree 50.
Atlantic coast and rivers of North and Central America.
1. Dorosoma cepedianum.
Megalops cepediana, Le Sueur, Journ. Acad. Philadelphia, i. 1818,
. 361,
ee heterura, Rafinesque, Amer. Monthly Mag. 1818, p. 354.
Dorosoma notata, Ratinesque, Ichth. Ohiensis, p. 39 (1820).*
Chatoessus ellipticus, Kirtland, Rep. Zool. Ohio, p. 169 (1839).
Dorosoma insverabilis, Abbott, Proc. Acad. Philadelphia, 1860, p. 865.
Chatoessus cepedianus, Giinth, Cat. Fish. vii. p. 409 (1868),
Dorosoma cepedianum, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1896, p. 416, and 1900, fig. 183.
Dorosoma cepedianum exile, Jord. & Everm. 1. ec.
Dorosoma exile, Meek, Zool. Publ. Columbian Mus. y. 1904, p. 94.
Mouth small, subterminal or inferior ; maxillary extending
to below anterior edge of eye. Depth of body 2 to 3 in the
length, length of head 32 to 43. Diameter of eye 4 to 5 in
length of head. 55 to 65 scales in a longitudinal series, 21 to
29 in a transverse series ; ventral scutes 17-19+10-13.
Dorsal 13-15. Anal 30-34. Pelvics inserted in advance of
origin of dorsal. A dark humeral spot, most prominent in
the young.
Cape Cod to Mexico, entering rivers.
Here described from five specimens, 180 to 260 mm. long,
from Virginia, Illinois, and Texas. In these the body is
deeper (depth 2 to 2} in the length) in the examples from
Virginia ‘than in those from Illinois and Texas (depth 22 to
3 in the length); but in young specimens this difference is
not apparent, the depth being about 3 of the length in both
forms.
2. Dorosoma anale.
Dorosoma anale, Meek, Zool. Publ. Columbian Mus. y. 1904, p. 98, fig.
Depth of body 22 to 3 in the length, About 70 scales in
312 Mr. ©. T. Regan on the Clupeoid Fishes
a longitudinal series. Dorsal 13-14. Anal 35-38. ‘In
other respects like D. cepedianum.
Atlantic coast streams of Mexico south of Vera Cruz.
T'wo examples, 120 to 160 mm. long, from Perez (Meck).
3. Dorosoma chavest.
Dorosoma chavesi, Meek, Zool. Publ. Columbian Mus. vii. 1907, p. 112.
Mouth rather large, with the jaws nearly equal anteriorly,
the mandible nearly $ the length of head and the slender
maxillary extending to below the middle of the eye. Depth
of body 24 in the length, length of head 23 to 3. Diameter
of eye 3 to 34 in length of head. 74 to 78 scales in a longi-
tudinal series ; ventral scutes 17-19+9-10. Dorsal 12-15.
Anal 26-30. - A dark humeral spot.
Total length 47 to 210 mm.
Lakes Managua and Nicaragua.
12. NEMATALOSA, gen. nov.
Mouth toothless, subterminal or inferior, transverse, its
cleft forming an angle; maxillary slender, distally slightly
expanded and curved downwards ; edge of dentary reflected
outwards in front of extremity of maxillary; one supra-
maxillary. Gill-rakers slender, very numerous. — Dorsal
13-18 ; last ray prolonged into a filament ; a scaly sheath at
base. Anal 18-24. Pelvics 8-rayed, below or a little in
advance of dorsal. Scales 44-50 in a longitudinal series,
14-21 in a transverse series. Vertebree 43 (in JV. erelz).
Coasts and rivers of Asia and Australia from Arabia to
Japan and New South Wales.
Synopsis of the Species.
I. Second suborbital covering cheek, its anterior edge vertical and its
lower edge horizontal and in contact with lower limb of pre-
Opercy lun fee ke eo Mep alessio s heb bias widen er aie 1. nasus.
II. Second suborbital with oblique antero-inferior edge; a naked area
above lower limb of przeoperculum.
A. Dorsal 16-18; pelvics below anterior part or middle of dorsal.
Anal 21—-93.; depth Siim lense in cea. ieee eile Beg 2. japonica.
Anal 19; depth 22 in length .............000. aiavats een 3. arabica.
Anal 20-22; depth 2 to 23 in length ...... aie eee Te 4. come.
B. Dorsal rays 15-16. Anal 18-22.
Depth 2 to 23 in length ; eye 83 (young) to 5 (very large
fish) in length of head; pelvics below or immediately
in advance of crigin Ofdorsal. 4.3 217.5 supe eat ee 5. erebi,
Depth 23 to 2 in length; eye 4 (young) to 5 (adult) in
length of head; pelvics below anterior 3 of dorsal .. 6. horni.
of the Genera Pomolous, Brevoortia, de. 313
1. Nematalosa nasus.
Clupea nasus, Bloch, Ausl. Fische, ix. p. 116, pl. eecexxix, fig. 1(1795).
Chatoessus aitus, Gray, Ill. Ind. Zool. pl. xci. fig. 2 (1835).
Chatoessus nasus, Cuv. & Val. Hist. Nat. Poiss. xxi. p- LOL (1848) ;
Day, Fish. Tndis a, p. 634, pl. elx. fig. 4 (1878).
Chatoessus chanpole, Giinth. Cat. Fish. vii. p. 410 (1868).
Depth of body 22 to 24 in the length, length of head 33 to
4. Snout as long as or shorter than diameter of eye, which
is 34 to 4 in lenethi of head ; maxillary extending to below
anterior + of eye; second suborbital covering cheek, with
vertical anterior edge and horizontal inferior “edge attached
to lower limb of praoperculum. 45 to 50 scales in a longi-
tudinal series, 15 to 19 in a transverse series ; ventral scutes
16-19 + 10-12. Dorsal 15-17. Anal 21-24. Pelvics
below origin or anterior 4 of dorsal. Dark longitudinal
streaks along upper series of scales; often a dark “humeral
spot.
India.
Several examples, 100 to 200 mm. long, from Sind,
Bombay, Canara, Madras, Calicut, and Burma.
2. Nematalosa japonica, sp. n.
Depth of body 3 in the length, length of head 43. Snout
as long as diameter of eye, which is = in length of head ;
maxillary extending to below anterior 4 of eye; second sub-
orbital with oblique lower edge. 48 to 50 seales in a longi-
tudinal series, 19 or 20 ina transverse series ; ; ventral scutes
19-20+12-14. Dorsal 16-18. Anal 21-25. — Pelvics
below middle or anterior part of dorsal. A dark humeral
spot.
Inland Sea of Japan.
Three specimens, 200 mm, in total length.
3. Nematalosa arabica, sp. n.
Depth of body 22 in the length, length of head 382. Snont
as long as diameter of eye, which is 4 in length of head ;
maxillary extending to below anterior 1 of eye; second sub-
orbital with ‘oblique lower edge. 50 scales in a longitudinal
series, 19 in a transverse series; ventral scutes "18413.
Dorsal 17. Anal 19, Pelvics a little in advance of middle
of dorsal. Dark longitudinal streaks along series of scales on
upper part of body.
Muscat.
- A single specimen, 150 mm. in total length.
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 21
314 Mr. C. T. Regan on the Clupeoid Fishes
4. Nematalosa come.
Chatoessus come, Qichards, ‘Erebus’ and ‘Terror’ Fish. p. 62,
pl. xxxviii. figs. 7-10 (1846).
Chatoessus nasus, Giinth, Cat. Fish. vii. p. 407 (1868).
Dorosoma nasus, Bleek. Atl. Ichth. vi. p. 142, Clup. pl. ii. fig. 4 (1872) ;
Weber & Beaufort, Fish. Indo-Austral. Arch. ii. p. 24 (1918).
Depth of body 2 to 24 in the length, length of head 33 to
4, Snout nearly as long as or shorter than diameter of eye,
which is 3 to 4in the length of head; maxillary extending
to below anterior 4 of eye ; lower edge of second suborbital
oblique. 46 to 50 scales in a longitudinal series, 17 to 20 in
a transverse series ; ventral scutes 18-20+10-12. Dorsal
16-18. Anal 20-22. Pelvics below anterior part or middle
of dorsal. Dark longitudinal streaks along upper series of
scales ; a blackish humeral spot.
Indo-Australian Archipelago.
Several examples up to 200 mm. in total length, including
one that I believe to be the type of the species (C. nasus,
specimen & of Giinther).
5. Nematalosa erebi.
Chatoessus erebi, Giinth. Cat, Fish. vii. p. 407 (1868).
Depth of body 2 to 2} in the length, length of head 34 to
41, Snout as long as or shorter than diameter of eye, which
is 3} to 5 in length of head ; maxillary extending to below
anterior } of eye; second suborbital with oblique lower edge.
46 to 50 scales in a longitudinal series, 17 to 21 in a trans-
verse series ; ventral scutes 18-19+10-12. Dorsal 13-16 ;
origin above or immediately behind base of pelvics. Anal
18-22.
Hast coast of Australia.
Several examples, 100 to 350 mm. in total length, from
Cape York, Burnett R., Mary R., and New South Wales.
6. Nematalosa horn,
Chatoessus horni, Zietz, Rep. Horn. Exped. ii. p. 180, pl. xvi. fig. 6
(1896).
Depth of body 24 to 24 in the length, length of head 3} to
4, Snout as long as or shorter than diameter of eye, which
is 4 to 5 in the length of head ; maxillary extending to below
anterior + of eye; lower edge of second suborbital oblique.
44 to 46 scales in a longitudinal series, 15 to 18 in a trans-
verse series; ventral scutes 16-18+9-11. Dorsal 13-16.
Anal 18-22. Pelvics below anterior 4 of dorsal.
of the Genera Pomolobus, Brevoortia, &c. 815
Australia.
Vive of the types, 100-170 thm. long, from Red Bank
Creek, McDonnell Range ; numerous examples from the
Bulloo Creek, interior of Queensland (‘Challenger’) and
some from the Borwan R., interior of New South Wales
(Stead).
e
13. GONIALOSA, gen. nov,
Mouth formed as in Nematalosa. Dorsal 14-17; a scaly
sheath at base ; last ray not prolonged. Anal 22-28. Pelvics
8-rayed, below or in advance of origin of dorsal. Seales
45-75 in a longitudinal series, 16-25 in a transverse series.
Vertebra 44-46,
Rivers of India and Burma,
1. Gonialosa modesta.
Chatoessus modestus, Day, Proc. Zool. Soc, 1869, p. 622, and Fish.
India, p. 633, pl. clx. fig. 1 (1878).
Depth of body 2 to 2} in the length, length of head 3 to
4, Snout shorter than diameter of eye, which is 3 to 34 in
the length of head ; maxillary extending to below anterior
edge of eye. 45 to 47 scales ina longitudinal series, 16 to 18
in a transverse series; ventral scutes 17-19+9-12. Dorsal
14-17. Anal 24-28. Pelvics below or in advance of origin
of dorsal. Vertebree 44. Usually a dark humeral spot.
Burma.
Seven specimens up to 100 mm. in total length,
2. Gonialosa manmina.
Clupanodon manmina, Ham, Buch, Fish. Ganges, p. 247 (1822).
? Clupanodon cortius, Ham. Buch, ¢. ¢. p. 249,
Chatoessus manmina, Cuv. & Val. Hist. Nat. Poiss. xxi, p, 114 (1848) ;
Day, Fish. India, p. 633, pl. elx. fig. 2 (1878),
Chatoessus cortius, Gunth, Cat. Fish. vii. p, 410 (1868),
Depth of body 22 to 31 in the length, length of head 33 to
43. Snout shorter than diameter of eye, which is 3 to 34 in
length of head ; maxillary not or barely reaching eye. 55 to
65 scales in a longitudinal series, 21 to 25 in a transverse
series. Ventral scutes 16-19+4+10-13. Dorsal 14-17,
Anal 22-26. Pelvics below or in advance of dorsal. Ver-
tebre 46. Sometimes a dark humeral spot.
Northern India and Assam.
Several specimens, to 130 mm, in length,
21*
316 On the Genera Pomolobus, Brevoortia, &e.
14. AnoponTostomA, Bleek. 1849.
Verh. Batav. Genootsch. xxil., Madura, p. 16.
Differs from Gondalosa in that the maxillary is a straight,
thin, transversely expanded lamina, tapering distally, whilst
the supramaxillary is very slender. Dorsal 17-19, with a
well-developed scaly sheath extending to tip of last ray.
Anal 18-21, depressible in a scaly sheath. Pelvics 8-rayed,
below middle or anterior half of dorsal. Scales 40-42 in a
longitudinal series, 12-17 in a transverse series. Ver-
tebree 42.
Coasts and rivers of India and Indo-Australian Archipelago.
1. Anodontostoma chacunda.
Clupanodon chacunda, Ham. Buch. Fish. Ganges, p. 246 (1822).
Chatoessus chacunda, Cuv. & Val. Hist. Nat. Poiss. xxi. p. 111 (1848) ;
Ginth. Cat. Fish. vii. p. 411 (1868); Day, Fish. India, p. 682,
pl. clx. fig. 3 (1878).
Anodontostoma husseltii, Bleek. Verh. Batay. Genootsch. xxii. 1849,
Madura, p. 15.
Chatoessus selangkat, Bleek. Verh. Batavy. Genootsch. xxiy. 1852,
Haringacht. p. 47.
Dorosoma chacunda, Bleek. Atl. Ichth. vi. p. 143, Clup. pl. ili. figs. 5,
(1872); Weber & Beaufort, Fish. Indo-Austral. Arch. ii. p. 2
fiz, 14 (1918).
Depth of body 2 to 23 in the length, length of head 3} to 4.
Snout shorter than diameter of eye, which is 3 to 32 in the
length of head; maxillary extending to helow anterior } of
eye. 40 to 42 scales in a longitudinal series, 12 to15 ina
transverse. series; ventral scutes 16-18+10-11. Dorsal
17--19. Anal 18-21. Pelvics below middle or anterior part
of dorsal. Dark longitudinal streaks along upper series of
scales; a dark humeral spot.
India and Indo-Australian Archipelago.
Numerous examples, up to 160 mm. in total length.
2. Anodontostoma breviceps.
Chatoessus breviceps, Peters, Monatsb. Akad. Berlin, 1876, p. 848.
Depth of body 22 in the length, length of head nearly 4.
Snout 3 as long as eye; maxillary extending to below middle
of eye. 42 scales in a longitudinal series, 17 in a transverse
series. Dorsal 19. Anal 19. Pelvics below middle of
dorsal. Longitudinal dark stripes along upper series of
scales.
Total length 230 mm.
New Hanover.
ma
Mr. R. E. Turner on Possorial Hymenoptera. 317
XXVII.—Notes on Fossorial Hymenopteraa—XXVIL. On
new Species in the British Museum. By Row ann LE.
Turner, F.Z.S., F.E.S.
Family Scoliida.
Subfamily Ezz.
Eilis bodkini, sp. n.
@. Nigra; clypeo lateribus, orbitis internis externisque anguste,
fascia transversa inter antennas, pronoto margine posteriore et
margine anteriore late interrupto, mesonoto macula quadrata
postice maculaque parva utrinque angulis posticis, postscutello
fascia, segmento mediano fascia longitudinali utrinque, meso-
pleuris fascia verticali sub alis, segmento dorsali primo macula
magna utrinque fasciaque angusta interrupta mediana, segmentis
tertio, quarto quintoque fascia basali, sexto macula transversa
basali, segmentis ventralibus 2-4 fascia lata emarginata, quinto
fascia mediana, angusta, interrupta, femoribusque intermediis
anticisque macula apicali flavis; alis subhyalinis, area radiali
late infuscata, venis fuscis ; mandibulis ferrugineis.
Long. 13 mm,
?. Clypeus finely punctured, subcarinate longitudinally in
the middle ; front and vertex coarsely punctured, with sparse
pale fulvous hairs ; frontal prominence subtuberculate on each
side on the inner side of the scape. Thorax closely and
rather coarsely punctured, more finely and closely on the
pronotum than elsewhere ; median segment subcarinate in
the middle at the base, a triangular space at the base much
more finely punctured than the rest ot the dorsal surface, the
sides of the segment shallowly obliquely striated. Abdomen
shining, finely and closely punctured, more strongly and
sparsely on the ventral surface ; sixth dorsal segment closely
and finely longitudinally striated. Second cubital cell very
Jong, the second abscissa of the radius nearly half as long
again as the third; first recurrent nervure received at three-
fifths from the base of the second cubital cell, second at two-
fifihs from the base of the third cubital cell.
Hab. River Mazaruni, British Guiana (G. ZL. Bodkin) ;
November 1916.
This is a smaller species than flavopicta, Sm., and has the
vertex much more closely punctured ; the puncturation of the
thorax is much closer and finer, the markings are somewhat
different, there is no blue gloss on the abdomen, and the
318 Mr. R. E. Turner on Fossorial Hymenoptera.
second abscissa of the radius is much longer. In the latter
character it resembles the Central-American ZH. pulchrina,
Cam., and £. bicincta, Sm., but differs from both in markings
and in the finer and closer puncturation.
Family Sapygide.
Sapyga furtiva, sp. n.
@. Nigra; antennis ferrugineis, apice infuscatis; clypeo lateribus,
fascia arcuata inter antennas, orbitis internis anguste, orbitis
externis supra, pronoto margine antico late interrupto, meso-
pleuris macula sub alis, postscutello fascia transversa interrupta,
segmento mediano macula magna apicali utrinque, segmentis
abdominalibus 2-5 fascia lata transversa, sextoque dorsali
macula magna ante apicem flavis ; femoribus subtus, tibiis, tarsis,
segmento dorsali primo fascia mediana, ventralique secundo basi
ferrugineis ; alis hyalinis, venis nigris, area radiali infumata.
Long. 11 mm.
?. Mandibles very broad, tridentate at the apex; clypeus
subrectangular, broader than long, the apical angles pro-
duced, longitudinally rugose. Head closely punctured-
rugose ; antenne thickened towards the apex, much stouter
at the base than in clavicornis; posterior ocelli at least half
as far again from the eyes as from each other. Thorax very
closely punctured ; abdomen shining, minutely and closely
punctured. Second abscissa of the radius half as long as the
third, more than twice as long as the first
flab, Simla Hills, 6300 ft.
Nearly allied to ecdavicornis, but differs in colouring, in the
larger second cubital cell, and in the stouter basal joints of
the flagellum.
Family Crabronide.
Subfamily Larrwz.
Dimorpha ruficaudata, sp. n.
@. Nigra; flagello fusco; scapo, mandibulis, pedibusque ferru-
gineis ; tegulis fuscis, apice testaceis; segmentis abdominalibus
4-6 rufis; alis hyalinis, venis fuscis.
Long. 8 mm.
9. Head sparsely punctured, the clypeus and front clothed
with long cinereous hairs, clypeus very short, transverse at
the apex, finely punctured and subcarinate in the middle ;
second joint of the flagellum distinctly longer than the third ;
Mr. R. E. Turner on Fossortal Hymenoptera. 319
posterior ocelli twice as far from each other as from the eyes.
Mesonotum shining, with scattered punctures, the anterior
third closely punctured and clothed with cinereous hairs,
which extend on to the pronotum, and more sparsely on to
the mesopleure. Scutellum smooth and shining. Median
segment strongly longitudinally striated, the space between
the striz more finely transversely striated, forming reticu-
lations. . Abdomen shining, microscopically punctured.
Radial cell very short, on the costal margin about twice as
Jong as the third abscissa of the radius, and not more than
half as long again as the apical margin of the cell; third
abscissa of the radius half as long again as the second, but
only one-third of the length of the second transverse cubital
nervure.
Hab. Nyasaland, Zomba (H. S. Stannus).
The colouring of the abdomen is unusual in the genus.
I use Jurine’s name Dimorpha for the genus instead of
Astatus, Latr., as to which there is some confusion.
Notogonia nigricans, Walk.
Notogonia nigricans, Walk. List of Hymen. in Egypt. p. 21 (1871). 9.
Notoyonia sculpturata, Kohl, Ann. Naturh, Hofmus. Wien, vii. p. 221
(1892). ¢.
There is a co-type of Walker’s species in the British
Museum.
Hab. Egypt; Port Soudan ; Albania; Gibraltar ; St. Vin-
cet, Cape Verde Islands.
Notogonia palumbula, Koll.
Notogonia palumbula, Kohl, Ann. Naturh. Hofmus. Wien, ix. p. 804
(1894).
Notoyonia punctipleura, Cam., Sjéstedt, Kilimandjaro-Meru Exp. ii.
p. 285 (1910). .
This is merely the tropical subspecies of nigricans ; the
pygidial area of the female is narrower,
Hab. Cameroons; Kilimandjaro; Lake Nyasa; N.E.
Rhodesia.
Notogonia reticulata, Cam.
Leptolarra reticulata, Cam, Ann. & Mag. Nat. Hist. (7) v. p. 81 (1900).
This is the Indian subspecies of nigricans, differing from
the typical form in the rather finer punctures of the meso-
notum,
320 Mr. R. EB. Turner on Fossorial Hymenoptera.
Aub. Barrackpore, N.E. India; Matheran, W. India;
Chapra, Bengal.
Very closely allied to these three forms -of nzgricans is the
Australian WV. retiaria, Turn., in which the punctures of the
mesonotum are almost obsolete and the eyes nearer to each
other on the vertex. I do not consider that small differences
in the comparative length of the abscissa of the radius are to
be relied on in this genus; there seems to be a slight indi-
vidual variation in this respect. WV. mahensis, Cam., from
the Seychelles, differs from retéculata in the longer and rather
narrower radial cell.
Notogonta irrorata, Sm.
Larrada trrorata, Sm. Cat. Hym. B.M. iv. p. 284 (1856). 9. :
Larva (Notogonia) fraudulenta, Kohl, Ann. Naturh. Hofmus, Wien, ix.
p. 303 (1894). @.
Hab. Senegal; Sierra Leone ; Ashanti ; Uganda.
Notogonia cresus, Sm.
Larrada cresus, Sm. Cat. Hym. B.M. iv. p. 284 (1856). 9.
Notogonia cresus, Kohl, Ann. Naturh. Hofmus. Wien, ix. p. 800
(1894). 9S.
Motes lirioides, Turn, Trans. Ent. Soc. London, p. 753 (1912). 9.
Although the tarsal ungues are toothed in this species in
the female, the very different form of the pygidial area shows
that it is not closely related to A/otes.
Hab. Kast Africa from Mashonaland to Witu; West
Atrica, Gambia and Gold Coast.
Doubtless this species ranges through the whole of tropical
Africa.
Notogonia deceptor, Turn.
Motes deceptor, Turn. Ann. & Mag. Nat. Hist. (8) xvii. p. 253 (1916).
oe
This is closely related to WV. cr@sus, and is not a Motes.
It may possibly prove to be a colour-variety of cr@sus, the
siructural differences being very slight.
Tachysphex excelsus, sp. n.
Q. Nigra; segmentis abdominalibus primo secundoque, tertioque
dimidio basali rufis; alis subhyalinis, leviter infuscatis.
Long. 12 mm,
¢. Clypeus broadly triangularly deflexed towards the
Mr. R. EE. Turner on Fossorial Hymenoptera. 321
apex, the triangular surface shining, with large scattered
punctures, the apical margin transverse. IHyes separated on
the vertex by a distance equal to twice the length of the
second joint of the flagellum; the third joint of the flagellum
about as long as the first.and second combined. Head and
thorax very finely and closely punctured; median segment
opaque, very finely granulate, the sides not striated ; the
posterior slope transversely striolate towards the apex, ail a
deep depression at the base. Abdomen highly polished ;
pygidial area elongate, rather sparsely and finely punctured.
The long spur of the hind tibia distinctly shorter than the
basal joint of the hind tarsus; spines of the fore tarsus
forming a comb, slender and fairly long. Radial cell longer
and natrower than in pectinipes ; the second abscissa of
the radius scarcely shorter than the third,
Hah. Tibet, Gyangtse, 13,000 ft. (1. J. Walton); June.
The sculpture of sthe median segment resembles 7. ee
Jrons, Kohl, from which it differs in other details. The eye
are further ajart on the vertex than in TZ. pectinipes, to
which it is allied in the form of the pygidial area.
Tachysphex filicornis, Koht.
Tachysphex filicornis, Kohl, Deutsch. Ent. Zeitsch. xxvii. p. 169.
This Mediterianean species occurs at Harar (G. Kvristen-
sen). A subspecies occurs at Salisbury, Mashonaland
(G. A. K. Marshall), in which the sculpture of the median
segment 1s much coarser, there being very distinct divergent
strize at the base, whereas the strise, as far as they are deve-
loped in fiicornis, are parallel. For this I suggest the name
Tachysphex filicornis eacerptus, subsp. n.
I do not regard the sculpture of the median segment as a
very reliable Sausete in this genus, considering that it is
liable to considerable variation in some species.
Tachysphex auropilosus, sp. n,
Q. Nigra; callis humeralibus, tegulis, abdomine, pedibusque rufo-
testaceis; segmentis ventralibus nigro intaminatis ; clypeo,
fronte, thor ace, segmento mediano, segmentisque dorsalibus mar-
gine apicali preecipue aureo-sericeo- pubescentibus ; ; alis pallide
-Havo-hyalinis, apice pallidissime infuscatis, venis testaceis,
Long. 14 mm,
¢. Clypeus broadly rounded at the apex; eyes separated
on the vertex by a distance not quite equal to the length of
822 Mr. R. E. Turner on Fossorial Hymenoptera.
the second joint of the flagellum. Thorax and median seg-
ment very closely and minutely punctured, rather thinly
covered with very short, delicate, golden pubescence ; the
posterior slope of the median segment finely transversely
striated, with a deep median sulcus. Pygidial area elongate-
triangular, shining, sparsely and rather strongly punctured,
very narrowly truncate at the apex. Comb of the fore tarsi
long ; tibize with short golden pubescence. Jadial cell
rounded at the apex, not truncate, third abscissa of the radius
longer than the second, which is equal to the space between
the recurrent nervures on the cubitus. Tarsal ungues very
long, as in the genus Wetogonia.
Hiab. British East Africa, Simba, 3350 ft. (S. A. Weave),
April; Makindu, 3300 ft. (S. A. Neave), April.
This seems to belong to the group of ZY. quadricolor,
Gerst., but is a smaller and less robust species, and the eyes
are nearer together on the vertex; the colour of the wings is
also different. The elongate ungues are very remarkable.
Tachysphea depilosellus, sp. n.
Q@. Nigra; mandibulis basi, clypeo dimidio apicali, scapo, flagello
articulo primo, articuloque secundo basi, callis humeralibus,
abdomine, pedibusque rufo-ferrugineis ; alis anticis fuscis, posticis
pallide fusco-hyalinis ; venis nigris.
Long. 12 mm.
9. Clypeus very broadly rounded at the apex, somewhat
deflexed from the middle, the apical half shining, with a few
large scattered punctures; the base of the clypeus and the
front clothed with very short, sericeous, silver pubescence.
Thorax closely microscopically punctured, the mesonotum
and scutellum bare; dorsal surface of the median segment
opaque, very closely and microscopically punctured, the sides
ot the segment obliquely, the apex transversely striated.
Abdomen slender; pygidial area elongate-triangular, sparsely
punctured. No pubescent fascize on the dorsal segments,
'l'arsal comb long, the basal joint of the fore tarsus with eight
spines. Radial cell broadly rounded at the apex ; second
abscissa of the radius longer than the third, which is longer
than the space between the recurrent nervures on the cubitus.
Hab. N. Rhodesia, Pakasa (O. Silverlock) ; January.
A very slender species, easily distinguished by the fuscous
wings from any other Ethiopian species with the abdomen red.
Mr. R. E. Turner on Fossorial Hymenoptera. 323
Tachysphex brinckere, sp. n.
©. Nigra; mandibulis basi, tibiis anticis basi et subtus, tarsis
anticis, tarsisque intermediis posticisque apice fusco-ferrugineis ;
segmentis abdominalibus primo secundoque rufo-ferrugineis ;
tegulis testaceis; alis flavo-hyalinis, apice pallidis; venis testaceis.
Long. 138 mm.
9. Clypeus broadly truncate at the apex, finely and closely
punctured on the basal half; the apical half deflexed, shining,
with large scattered punctures. Vertex very closely micro-
scopically punctured; the eyes separated on the vertex by
a distance slightly exceeding the length of the second joint
of the flagellum ; front and the base of the clypeus clothed
with very short silvery pubescence, which is only visible in
certain lights. Thorax minutely and closely punctured ;
median segment granulate, as Jong as the mesonotum, the
sides and apex ot the segment striated, Second and third
dorsal segments with a little short silver pubescence at the
apical angles; pygidial area elongate-triangular, sparsely
punctured. Comb of fore tarsus long, basal joint of the fore
tarsi with eight spines. Radial cell broadly rounded at the
apex ; second abscissa of the radius longer than the third,
nearly twice as long as the space between the recurrent ne1-
vures on the cubitus,
Hab. Transvaal, Pretoria (Miss J. Brincker).
Tachysphex punctata, Sm.
Larrada punctata, Sm. Cat. Hym. B.M. iv. p. 282 (1856). ¢ (as 2),
Larra punctata, Kohl, Verh. zool.-bot. Ges. Wien, xxxiv. p. 247 (1884).
The type is a male, not a female, and is a true Tachysphea.
The wings are of a darker fuscous than in any other small
black Ethiopian species of the genus known tome. L[yes
separated on the vertex by a distance equal to about twice
the length of the second joint of the flagellum.
Tachysphex subfuscatus, sp. 0.
Q..Nigra; segmentis abdominalibus duobus basalibus fusco-rufis ;
tarsis fusco-ferrugineis ; alis subhyalinis, venis fuscis ; clypeo,
fronte, segmentisque dorsalibus tribus basalibus fascia apicali
argenteo-pubescentibus ; thorace rugose punctato; segmento
mediano longitudinaliter striato-reticulato.
Long. 9 mm.
9. Clypeus broadly truncate at the apex, the apical
324 Mr. R. KE. Turner on Fossorial Hymenoptera.
margin armed with a number of ill-defined teeth. Head
closely and rather finely punctured; antennze short and stout ;
eyes separated on the vertex by a distance slightly exceeding
‘the length of the two basal joints of the flagellum. Thorax
coarsely punctured- rugose; scutellum punctured ; median
segment irregularly longitudinally striate, with finer trans-
verse striz, giving a reticulate appearance, which is more
strongly developed on the more coarsely sculptured sides of
the segment. Pygidial area smooth, rather broadly trian-
gular. Legs slender ; comb of the fore tarsi long ; spur of
the hind tibia much shorter than the basal joint of the hind
tarsi, Radial cell broadly obliquely truncate at the apex ;
second and third abscissee of the radius pchias
Lfab, Nyasaland, Mlanje, 2300 ft. (S. A. Neave) ; October.
Tachysphex strigatus, sp. n.
©. Nigra; tarsis articulis apicalibus brunneo-ferrugineis ; fronte
clypeoque argenteo-pubescentibus; mesonoto crasse punctato,
cinereo-piloso; segmentis dorsalibus tribus basalibus fascia
interrupta apicali argenteo-pubescente; segmento mediano
fortiter longitudinaliter striato; alis hyalinis, venis brunneo-
ferrugineis; tegulis testaceis.
Long. 9-11 mm.
2. Clypeus broadly subtruncate at the apex, the apical mar-
gin somewhat reflexed and with two blunt teeth on each side,
closely and not very finely punctured. Front opaque, finely
punctured-rugulose, the vertex closely and not very finely punc-
tured, yes ‘separated on the vertex by a distance half as great
again as the length of the second joint of the flagellum. Meso-
notum and mesopleurze coarsely punctured-rugose ; scutellum
strongly but not very closely punctured. Median segment
coarsely longitudinally striated, with finer irregular, transverse
strie between ; the sides thre segment coarsely rugose-reti-
culate. Abdomen shining ; pygidial area triangular, shining,
with a few small scattered punciures. Radial cell rather broadly
obliquely truncate at the apex ; second abscissa of the radius
a little longer than the third, equal to the distance between
the recurrent nervures on the cubitus.
Hab. N.1. Rhodesia, between Fort Jameson and Lundazi,
4000 ft. (S. A. Neave), June ; Central Angoniland, Lilongwe
District, 4000-5000 ft. (S. A. Meave), May; Nyasaland,
Mombera District, 4000 ft. (S. A. Neave), June; Nyasaland,
Kotakota (Dr. J. L. S. Old).
Easily distinguished by the coarse sculpture of the thorax
and median segment.
Mr. R. E. Turner on Fossorial Hymenoptera. 325
Tachysphex vulneratus, sp. n.
©. Nigra; mandibulis basi, tegulis, tibiis tarsisque anticis, femori-
bus apice, tibiis intermediis et posticis subtus, tarsisque inter-
mediis et posticis basi nigro intaminatis, brunneo-testaceis ;
segmentis abdominalibus quinto sextoque omnino, quartoque
apice rufis; alis hyalinis, venis fuscis; fronte, clypeo, segmentis-
que dorsalibus tribus basalibus fascia apicaliinterrupta argenteo-
pubescentibus ; segmento mediano longitudinaliter striato.
6. Femine similis, tibiis tarsisque intermediis et posticis brunneo-
testaceis ; segmentis dorsalibus sexto septimoque omnino, quinto-
que apice rufis ; fronte clypeoque aureo-pubescentibus.
Long., 2 10-11, ¢ 8-9 mm.
2. Clypeus truncate at the apex, rather broadly depressed
on the apical margin, with two minute teeth on each side.
Head finely and ciosely punctured ; eyes separated on the
vertex by a distance not quite equal to the length of the
second joint of the flagellum ; antenne slender ‘and rather
long, ‘Thorax closely and not very finely punctured ;
median segment strongly longitudinally striated, the sides of
the seoment more finely obliquely striated, the surface of the
posterior truncation finely transversely neds Pygidial
area triangular, not elongate, shining, with a few scattered
punctures. Comb of the fore tarsus long and slender ; the
Jong spur of the hind tibia almost as long as the basal joint
of the hind tarsus. Radial cell long, rather narrowly rounded
at the apex; third cubital cell about as long as the second,
both on the cubitus and on the radius.
6. Seventh dorsal segment broadly rounded at the apex ;
eighth ventral segment shallowly emarginate, the angles
produced into distinct teeth.
Hab. N.E. Rhodesia, Niamadzi. River, near Nawalia,
2000 ft. (S. A. Meave), August; Mid Luangwa Valley,
2000 ft. (S. A. Weave), July ; "Upper Luangwa Valley
(S. A. Neave), August.
This differs from strigatus in colour, in the much finer
sculpture of the thorax, in the lesser distance between the
eyes, and in the long and slender antenne.
Prosopigastra neavei, sp. n.
©. Nigra; mandibulis in medio, abdomine segmentis tribus basali-
bus, calcaribus, tarsisque articulis apicalibus ferrugineis; tegulis
testaceis ; alis hyalinis, iridescentibus, venis nigris,
do. Femine similis; segmentis abdominalibus 5 ‘apicalibus nigris ;
tarsis ferrugineis ; tibiis basi albido-maculatis.
Long., 9 7-8, d 6 mm.
326 Mr. R. H. Turner on Fossorial Hymenoptera.
9. Clypeus very widely arcuately deflexed towards the
apex, the deflexed portion smooth and shining, the apical
margin subtruncate. HWyes separated on the vertex by a
distance equal to about four times the length of the second
joint of the flagellum; head very distinctly but not very
closely punctured ; a smooth convex area between the ante-
rior ocellus and the base of the antenns. Thorax rather
more strongly punctured than the head, the individual pune-
tures large and clearly separated. Median segment scarcely
more than half as long as the mesonotum ; the dorsal surface
margined by caring at the sides and apex, irregularly and
coarsely striate-reticulate ; the sides of the segment longitu-
dinally striated, the posterior slope rugose. Abdomen closely
and finely punctured ; pygidial area shining, sparsely punc-
tured, very narrowly truncate at the apex. Comb of thie
fore tarsi long and slender. Radial cell short, very broadly
obliquely truncate at the apex; second and third abscissee
of the radius subequal, each at least half as long again as the
fourth.
3d. Seventh dorsal sezment broadly rounded at the apex ;
eighth ventral segment emarginate, testaceous, the apical
anvles produced into short spines. Hyes separated on the
vertex by a distance not exceeding half the length of the
second joint of the flagellum.
Hab. N.E. Rhodesia, Mid Luangwa Valley, about
2000 ft. (S. A. Neave), July and August; Nyasaland, be-
tween I*t. Jameson and Dowa, 4000 ft. (S. A. eave),
October.
The sexual divergence in the distance between the eyes
on the vertex is greater than in any Mediterranean species
of the genus.
Subfamily Trrpoxrroyivaz.
Pison papuanum, Schulz.
Pison papuanum, Schulz, Berlin. Ent. Zeit. xlix. p. 217 (1904).
Pison morosus, Sm. Journ. Linn. Soc., Zool. viii. p. 85 (1864). 9 (nec
Sm. 1856).
Pison constrictum, Turn. Ann. & Mag. Nat. Hist. (8) ix. p, 201 (1912).
3.
Pison constrictum, Turn. Proc. Zool. Soc. London, p. 627 (1916).
I had overlooked the name papuanum in my recent paper
on Pison.
Ona new Terrestrial Isopod from New Zealand. 327
XXVIII.—A new Tuberculate Terrestrial Isopod from New
Zealand. By Cuas. Cuitton, M.A., D.Sc., M.B., C.M.,
LL.D., C.M.Z.S., Professor of Biology, Canterbury
College, New Zealand.
[Plate XIII.]
In 1915 * I described a tuberculate species of Cubaris from
New Zealand under the name C. sutert, Of this species I
had only the one specimen, and I stated that of a second
tuberculate species, CO. hamiltoni, only a single specitnen was
known, these facts showing that our knowledge of the
terrestrial Isopoda of New Zealand was still very incomplete.
I suggested also that a careful survey, especially in the
forests of the North Island, might bring to light other inter-
esting species. This has already proved to be the case, and
I have recently received from Mr. David Miller, of the New
Zealand Agricultural Department, several specimens of
another tuberculate Cubaris found under the bark of fallen
logs in the bush at Levin, Wellington. Of this species
Mr. Miller was fortunate enough to find eight specimens.
In general appearance, colour, markings, etc., they are very
similar to Cubaris suter’, and I at first thought that they
might perhaps be specimens of this species with the tubercles
on the dorsal surface better developed than in the type-
specimen. This, however, proves not to be the case, as the
tubercles, or, rather, ridges, are arranged differently, and I
am therefore describing the specimens as a new species, which
I have much pleasure in naming after their discoverer.
Cubaris millert, sp. n. (Pl. XIII. figs. 1-6.)
Specific description.—Oblong-oval, breadth about half the
length. MKpimeral portions fairly well developed, especially
in the first segment of the pereon ; central portion of each
segment very convex and marked off from the lateral portions
by a longitudinal ridge or flange on each segment (figs. 1 & 2).
Head with the anterior margin produced upwards into a well-
defined ridge projecting slightly above the dorsal surface and
having the upper margin regularly convex and without any
notch : the posterior surface of the head is produced dorsally
into a distinct transverse flange rising high above the general
surface and showing in front view much higher than the
* Journ. Linn, Soc. vol. xxxii. p. 425, pl. xxxvii. figs, 24-28,
328 Prof. C. Chilton on a new Tuberculate
anterior margin; the flange has a slight depression in the
centre, so that. its upper margin is concave (fig. 3). Each
seement of the perzeon bears a pair of longitudinal tubercles
or ridges, which are low anteriorly but become higher towards
the posterior part of the segment; these ridges increase in
size and distinctness on the posterior segments until, in the
seventh segment, the ridge is much higher than the segment
itself and projects backwards over the pleon. In dorsal view
these ridges form an almost continuous row, separating the
central part of the body from the lateral portions. In each
segment there are a few small tubercles or irregularities both
on the lateral portions below the ridge and also on the central
part between the ridges. Inferior margin of first segment of
person deeply cleft posteriorly, the cleft extending nearly
halfway along the whole margin; inferior margin of the
second segment with a distinct tubercle on its inner surface
enclosing a wide notch for the reception of the succeeding
seoment when the animal is rolled up into a ball (fig. 4),
The pleon bears no ridges and shows the usual characters ;
the posterior segment has the hind margin either straight or
very slightly concave (fig. 5).
Antenne (fig. 3) of normal shape, the second and third
segments of peduncle subequal, the fourth a little longer and
the fifth nearly twice as long as the fourth; flagellum a little
shorter than the fifth joint of peduncle, its first joint about
one-third the length of the terminal joint.
The mouth-parts show the usual structure common to the
genus, and do not appear to present any distinctive characters.
The legs are all short and of the usual form. In the single
male dissected the anterior pairs do not show any special
modification; but as the specimen is small and the legs
imperfect the evidence on this point is not quite conclusive.
The pleopoda of the male do not appear to differ in any
important points from those of other species of the genus.
The uropoda (figs. 5 & 6) have the endopod very short,
almost knob-shaped, extending only ashort distance from the
base ; its extremity bears two or three minute sete. The
exopod is also very small, reaching only about halfway from
its attachment to the posterior end of the peduncle ; it bears a
rather long seta, which reaches nearly as far posteriorly as
the pedunele.
Under a high power the whole integument shows minute
seale-like markings.
Colour. Pale reddish brown, with marblings of a darker
brown,
Terrestrial Isopod from New Zealand. 329
Length of largest specimen about 7 mm.
Loc. Under the bark of fallen logs in the bush, Levin,
Wellington, N.Z.
This species appears to be closely related to Cubaris sutert,
Chilton, the structure of the lateral margin of the first and
second segments of the perwon and of the uropoda being
closely similar in the two species. In C. sutert, however,
the ridges are transverse and mainly confined to the posterior
border of the peraon segments, while in the present species
the ridges are longitudinal, extending along nearly the whole
of the length of each segment, and they are much better
developed and consequently more prominent. Another
tuberculate species, C. hamiltont (Chilton) *, probably also
comes near to these two species; but the dorsal surface is much
more profusely supplied with flanges or ridges and with
pointed tubercles. C. hamiltoni is known only from the
single ty pe-specimen which was obtained in the neighbourhood
of Petane, near Napier, in New Zealand, and this specimen is
unfortunately somewhat imperfect, so that our knowledge of
the species is far from complete. The only other tuberculate
species known from New Zealand is C. macmahoni (Chilton),
originally described from Kenepuru in Marlborough, though
I have since had specimens sent to me from one or two
localities in the North Island. C. spinosus (Dana) is a spiny
species, “ the body bristied throughout with subacute spines ” ;
but itis only known from Dana’s brief description and figures,
no specimen having been’since collected. It was found by
Dana near the Bay of Islands.
I am much indebted to my assistant, Miss E. M. Herriott,
M.A., for preparing the drawings to illustrate this paper.
EXPLANATION OF PLATE XIII.
(All the figures refer to Cubaris millert, sp. 0.)
Fig. 1. Dorsal view of whole animal.
Fig. 2. Side view of animal (antennz and legs not shown).
Fig. 3. Front view of head with antenne etc., the ange arising from
the posterior border of the head showing behind the anterior
margin.
Fig. 4. Lateral margins of perzeon segments 1, 2, and 3, from below.
Fug. 5. Terminal portion of pleon, from above.
Fig. 6. Uropoda and terminal segment, from below.
* See Trans. Linn. Soc., Zool. vol. viii. pp. 99-152, pls. xi.-xvi., and
Trans. N.Z. Inst, vol. xlii. pp. 286-291.
Ann. & Mag. N. Hist. Ser. 8. Vol, xix. 22
330 On South-African Talitridee.
XXIX.— South-African Talitride.
By the Rev. Tuomas R. R. Stepsine, M.A., F.R.S.
On Christmas Eve in1916 Mr. H. W. Bell-Marley, of Durban,
Natal, obtained some specimens of Talitridze at Eshowe Bush,
1800 feet above sea-level. Of those which he has kindly
forwarded to me most are females, but one or two males
among them, though less in size than many of the other sex,
will, I think, settle a question which has long been obscure.
The species is clearly that which Spence Bate in 1862 named
Tulorchestia? africana. Clearly, also, it may now be referred
to the genus named Talitriator by Methuen in 1913, and more
fully defined by Barnard-in 1916.
Genus TALITRIATOR, Methuen.
1913. Talitriator, Matthews, P. Z. 8. Lond. p. 109.
1916. Talitriator, Barnard, Ann. 8. African Mus. vol. xv. pt. 3, p.222
ae
Related to Talitrus by feeble minutely chelate second
gnathopods in both sexes; distinguished from it by the first
antennee nearly as long as the peduncle of the second ;
maxillipeds with small fourth joint to the palp; first gnatho-
pod shorter than second; fifth side-plates of perzeon more
‘unequally bilobed ; telson longer than broad.
Of these characters, the last two seem to be scarcely of
generic importance. Spence Bate considers the fifth side-
plate to be equally bilobed in Zelitrus; but neither his own
figures nor the facts support this statement.
Talitriator africanus (Bate).
1862. Talorchestia ? africana, Bate, Amphipodous Crust. Brit. Mus.
p. 15, pl. ii. figs. 6, 62, 64.
1906. Talorchestia ? africana, Stebbing, ‘Das Tierreich, Lief. 21,
. 564.
1910. Talorchestia ? africana, Stebbing, Ann. S. Afr. Mus, vol. vi.
pt. 4, p. 459.
1912. Talitrus ? africanus (Bate), Calman, Ann, & Mag. Nat. Hist.
ser. 8, vol. x. p. 185 (1912).
1918. Taltriator eastwoode, Methuen, P. Z. 8, Lond. p. 110, pls. x., xi.
1916. Talitriator eastwoode, Barnard, Ann. 8. Afr. Mus. vol. xv. pt. 3,
p. 223 (and Talorchestia ? africana, p. 215).
In the male specimen the flagellum of the first antenne has
ten joints, in the female eleven, in hoth sexes the third joint
of the peduncle is the longest, in the second antenne the
flagellum of the male has 22, the larger female 23 joints.
‘The palp of the first maxilla is minute. Of the customary
On new Species of Indo-Malayan Lepidoptera. 331
three teeth on the inner plate of the maxillipeds two are very
conspicuous, but the innermost small, as shown by Methuen.
For the second gnathopod Methuen gives ‘ coxal plate
excavate behind with conical projection.” Barnard mentions
this as having specific value in the genus and as excluding
the typical species from Tu/itrus. It is, however, found in
T. alluaud’, Chevreux, 1896. Methuen states that the first
pereeopod is not quite as long as the second. ‘This, surely, is
an accidental reversing of the true relation. For the grext
size of the anterior lobe of the fifth side-plate there is a
parallel in 7. alluaudi. Our specimens show four pairs of
setules on the telson, while Methuen’s figure shows only two
pairs ; but Barnard supposes that Methuen’s specimens were
probably not quite mature. A fine red colour was retained
by Mr. Bell-Marley’s specimens as received nearly two
months after capture. As this is probably a terrestrial
species, it is desirable to point out that in Metluen’s notes on
distribution the word “ depths ”” has by some mischance been
substituted for “heights” in the quotation from ‘ Das
Tierreich,” .
XXX.—New Species of Indo-Malayan Lepidoptera.
By Colonel C. Swinuosg, M.A., F.L.S.
Danainin.
Salatura pleaippus adnana, nov.
3 ?. A local race of plewippus, uniformly smaller; all the
black vein-markings narrower ; the black apical portion of
the fore wing broader, consequently the bronzy-red interspace
between veins 2 and 3 much shorter; no indication of the
‘small similarly coloured space always present in plew/ppis in
the next upper interspace, just outside the cell-end, aud the
series of subapical bars all much shorter,
Expanse of wings, g 238, 2 -3 inches.
Hab. Luzon.
Staudinger refers to this local race in ‘ Iris,’ 1889, p. 28.
Evrrermy 2.
Tsamia eclecta, nov.
d. Upperside dark blackish brown: fore wing paler on
the outer third, the inner two-thirds with a slight blue-black
Beis 09%
332 Colonel C. Swinhoe on new
gloss; a minute blue-grey spot at the lower end of the cell,
another outside it in the interspace above vein 4, and another
above the upper end of the cell close to the costal margin :
hind wing with the costal space whitish, descending a little
into the cell; a very faint series of blue-grey dots close to
the outer margin; no other markings on either wing. Under-
side fairly uniform blackish brown, paler than it is above ;
fore wing with the hinder marginal space whitish; spots
larger and more prominent, one at the lower end of the cell,
another beyond it ; a rather long oval spot in the interspace
above vein 2, a small spot outside it, and three small spots
close to the margin above the hinder angle, and two small
spots at the base of the wing: hind wing with three basal
small spots, one at the end of the cell, five in a line in the
interspaces 2, 3, 4,5, and 6, a submarginal spot in inter-
space 2, two close together in interspace 3, and one in inter-
space 4; a series of somewhat larger spots close to the
margin in the interspaces up to interspace 4; cilia with white
dots in the interspaces both above and below. Head with
three white spots on each side; thorax above with a white
central line; below, palpi with a white spot between them
and one on each side, thorax covered with white spots, and
the abdomen with a central row of larger spots.
Expanse of wings, ¢, 4 inches.
Hab. Palone, Burma, June 1887.
Prerinz.
Hebomoia solomonensis, nov.
-$. Fore wings with the orange apical portion occupying
more than one-third of the wing, extending well into the cell,
filling up very nearly the whole of interspace 3 and the outer
and upper halt of cell 2; the costal band very narrow, blackish
grey powdered with ochreous, thickens a little at the apex,
runs down the outer margin very narrowly, and ends in a
blackish suffused small patch just above the hinder angle;
the interior blackish band which usually limits the orange
portion entirely absent ; the submarginal blackish spots in
the orange patch spear-shaped and very pale: hind wings
without any marginal band. Head and body powdered with
ochreous.
Expanse of wings, ¢, 375 inches.
Hab. Solomons.
Madais vi.
Teracolus vi, Swinhoe, P. Z. 8. 1884, p. 487, pl. xxxix. figs. 6, 7.
Teracolus immaculata, Rober, Seitz, Macro. Lep. i. p. 86,
Species of Indo- Malayan Lepidoptera. San
My type came frem the vicinity of Aden in Arabia, Rober’s
type from Syria ; [I have both in my museum, and there can
be no doubt they are identical. Teracolus vi is not mentioned
in Seitz.
Family Aganaide.
Asota lara.
Hypsa lara, Swinhoe, Ann. & Mag. Nat. Hist. (6) xii. p. 215 (1893).
Aganais intacta, var., Snellen, Tijd. voor Ent. xxxi. p. 138, pl. ii. fig. 4
(1888).
Hab. Java.
It is a good species, quite different to intacta, Walker,
having a broad, central, longitudinal stripe on the fore wing ;
it is apparently quite common in Java; I have received
several examples from Mt. Gedé and Buitenzorg.
‘ Family Drepanida.
Sewa orbiferata.
Abracas orbiferata, Walker, xxiv. 1126 (1862).
Argyris insignata, Moore, P. Z, 8, 1867, p. 645.
Platypteryx cilicoides, Snellen, 1, c, xxxii, p. 9, pl. i. fig. 3 (1889).
flab. Sarawak, Borneo (type in B. M.). ;
The type of ¢nségnata in the B. M. is marked “ Bengal.”
Snellen’s type is from Java. I have it from Mone, Shan
States (Adanders), and from Kina Balu, Borneo (£verete).
‘They are all very similar.
Ticila argentilinea.
Ticilia argentilinea, Walker, xxxii. 394 (1865); Swinhoe, Cat. Het.
Mus. Oxon. i. p. 244, pl. vii. fig. 18, ¢ (1892).
Platypteryx argentilinea, Snellen, /. ¢. p. 8, pl. 1. fig. 2, 2 (1889).
Hab. Singapore (type 2? in Mus. Oxon).
It is also from Sula in Mus. Oxon. (a ¢). Snellen also
described his type trom Java as argentilinea.
Family Lasiocampide.
Sitina cinyra, nov.
?. Palpi black, with some white hairs on its upperside ;
head and thorax covered with long ochreous-while (nearly
pure white) hairs; abdomen black; anal tuft white: tore
wing black, irrorated with very minute white atoms ; a large
round black spot with a white line through it at the end of
334 Colonel C. Swinhoe on new
the cell; an antemedial, sinuous, transverse white line; a
postmedial diffuse white band, narrowing hindwards, and
through it a black dentate line, curved outwards below the
costa, its points outwards; a series of black lunules with
white outer edges close to the margin ; cilia with some white
spots: hind wing paler, uniform in colour, without irrorations,
a white waved band across its middle from the middle of the
costa to the abdominal margin near the anal angle; cilia
white, with pale blackish spots. Underside uniformly
coloured like the upperside of the hind wing ; a rather broad
white band across both wings, evenly outwardly curved, -
postmedial on fore wing, medial on hind wing; the marginal
matks on both wings as on the upperside. Body and legs
black, with white hairs ; abdomen with white lateral bands.
Expanse of wings 14 inch.
Hab. N. Gippsland, Victoria (H. W. Davey).
It is unnamed in the B. M.
Sitina epipasta, nov.
@. Palpi ochreous brown ; head and shoulders covered
with white hairs; thorax black, with ochreous-grey hairs ;
abdomen black, with some ochreous-grey hairs on the first
two segments, small tufts of white hairs on the middle of the
last two segments and on each segment at the sides ; anal
tuft white: fore wing grey, darkest on the middle of the
costa, blackish on the basal half of the hinder margin, the
wing eovered with minute white irrorations, dense at the
base and on the lower half of the middle; below the cell a
white, sinuous, transverse, antemedial line; a small white
Junular mark at the end of the cell; a postmedial band of
grey Junules outwardly edged with white; a submarginal
row of black lunules, outwardly edged with white; veins
brown, finely marked with white: hind wing uniformly
grey, with a nearly straight white band from the middle of
the costa to the abdominal margin above the anal angle ;
cilia of both wings grey. Underside coloured uniformly
erey as on the upperside of the hind wing; a transverse
medial white band on both wings, nearly straight on fore
wing, outwardly and evenly curved on hind wing. Bod
concolorous with the wings ; abdomen with the lateral white
spots continued into segmental bands, its anal segment white ;
legs with white hairs.
Expanse of wings 1,%5 inch.
Hab. Yackandandah, Victoria (7. W. Davey).
Species of Indo-Ma’ayan Lepidoptera. 335
Family Deilemereida.
Deilemera luzonica, nov.
?. Belongs to the evergista group, nearest to eres, Boisd.,
and gerra, Swinhoe (Trans. Ent. Soc. 1903, p. 63, pl.iv. fig. 1);
fore wing of the same blackish-brown colour ; a longitudinal
white streak near the base, shorter than in gerra, with a small
white spot below it near the base; a very large white patch
with waved outer side, commencing at the upper end of the
cell in a narrow rounded form, broadening hindwards to the
internal vein ; its inner side is slightly excavated at the lower
margin of the cell, and then runs inwards below the outer end
of the basal streak, occupying a large portion of the central
space of the wing; two large white, rounded, submarginal
spots as in gerra, but much larger: hind wing with a narrow
* costal blackish-brown band and an even outer marginal band,
as in gerra, with a submarginal white spot in it, a little
below the apex. Head and body yellow ; collar with two
biack spots ; thorax covered with short green scales ; abdo-
men with broad black segmental bands.
Eixpanse of wings 1,8; inch.
Hab. Luzon, Philippines.
Deilemera purata, nov.
?. Milk-white ; palpi white, the last joint black; top of
head with a black spot, two on the collar; thorax with a
black medial line, and another thinner line on each side of it;
abdomen with a dorsal row of pale blackish spots; legs white,
without markings: fore wings with the veins grey, a darker
grey blotch or patch at the lower end of the cell: hind wing
with dark grey streaks at the vein-ends, decreasing in size
hindwards. Underside with all the vein-ends grey and a
large space on the fore wing blackish from the base to the
end of the cell, extending upwards to the costa, the veins
through this black space white.
Lixpanse of wings 1;%) inch.
Hab. Luzon, Philippines.
Figured by Semper as a female aberration * of Deilemera
sonticum, Swinhoe, also from the Philippines ; but I have in
my museum both sexes of sonticum trom Mindanao and
Luzon. ‘lhe sexes of that species are alike and are widely
different from this form, though the palpi, head, and body
are similarly marked.
* Phil. Schmett. pl. Iviii. fig. 7 (1899).
336 Colonel C. Swinhoe on new
Family Lymantriide.
Euproctis servilis.
Euproctis servilis, Walker, xxxii. 850 (1865). g.
Darala prima, Walker, xxxv. 1917 (1866). ¢.
Euproctis incompta, Snellen, Tijd. voor Ent. xx. p. 9, pl. i. fig, 2
(1879). ¢.
‘Euproctis flwvipennis, Snellen, 1. ec, xxii. p. 107, pl. ix. fig. 1 (1879). 9.
Euproctis cinerea, Heylearts, Ann. Soc. Ent. Belg. xxxvi. p. 10 (1892).
Euproctis nurma, Druce, Ann. & Mag. Nat. Hist. (7) iil. p. 469 (1899).
Type ¢, Celebes, in Mus. Oxon.
Type & prima, Celebes, in Mus. Oxon.
Types ¢ incompta, Java, in coll. Snellen.
Type ? flavipennis, Makassar, Celebes, in coll. Snellen.
Type cinerea, Java.
Type nurma, Timor, in coll. Joicey.
As stated in my monograph of this family in Trans. Ent.
Soc. 1903, p. 420, the colour of the fore wings varies much,
from pale yellow to olive-brown, and the hind wings from
yellow to white. I have the two extremes from the same
locality ; I have received it from Celebes, Java, Talaut, and
Kina Balu, Borneo: the markings are all identical.
Family Hadenide.
Cirphis philippensis, nov.
3 2. Palpi, head, body, and fore wings brownish ochreous,
much as in the common C, doreyi, Dup.: fore wing with a
narrow white streak along the median vein to the end of the
cell, with some blackish scales below its basal half ; narrower
white streaks on all the other veins, and still narrower (very
fine) streaks in all the interspaces ; some blackish scaling on
the basal half of the hinder margin ; a small black mark at
the lower end of the cell, at the end of the white streak first
mentioned, a black dot in the interspace below the middle of
the cell, and another in the same interspace more than half the
distance between it and the outer margin, some black points
on the outer margin ; cilia brownish ochreous, variegated by
the white streaks running into it: hind wings pure white,
without any markings, Underside with the fore wing paler,
with the white streaks less distinct and a black spot one-sixth
before the apex, close to the costa.
Expanse of wings, ¢ 2, 1,%) inch.
Hab, Luzon, Philippines,
Species of Indo-Malayan Lepidoptera. 337
Family Catocalide.
Attatha flavata, nov.
?. Head and body yellow; collar black; a broad black
band across the middle of the thorax ; a square black patch
at the base of the abdomen: fore wings bright yellow ; ashort
black streak from the base; a long black streak on the
hinder margin, not reaching the base nor the hinderangle; a
black band from the middle of the costa, narrowing hindwards
to near the hinder angle, and a triangular black patch from
the costa near the apex, much as in A, regal’s, Moore, from
India ; four black spots on the lower portion of the outer
margin: hind wing paler yellow, with a marginal series of
small black spots. Underside dull yellow, quite uniform in
colour ; a rather large, quadrate, blackish patch at the end of
the cell ; small black marginal spots on the hind wing.
Expanse of wings, ?, 1,8) inch.
fab. Manilla ; two examples received from Herr Semper
as A. flavata, Semper ined., but has never been published.
Attatha coccinea, nov.
?. A larger insect than flavata ; head and thorax yellow ;
frous black ; collar, middle band across thorax, and patch at
base of abdomen black as in flavata ; abdomen scarlet: fore
wing bright yellow, the bands and streaks as in flavata; the
apical patch not excavated on its outer side as in regalis, the
central band narrower: hind wing scarlet, marginal spots
small and black. Underside: both wings and body and
Jegs uniform scarlet ; fore wing with a dark black patch at
end of cell as in flavata, but blacker, no black patch in the
middle of the outer margin as in regalis, one black spot at
the end of vein 3, and a series of black spots on the outer
margin of the hind wing ; the subterminal large black spot
in regalis near the anal angle on the upperside not present.
Expanse of wings, 2, 1% inch.
Hab. Luzon (Semper).
Family Stictopterida.
Stictoptera poliata, nov.
?. Head, body, and fore wings dark grey, covered with
blackish irrorations, which are uniformly distributed throughout
the fore wings except in the middle of the wing, through which
there is a prominent black thick line, uniform, and evenly
outwardly curved, marginal points black: hind wings with
338 Colonel (. Swinhoe on new
broad and even black border, which occupies nearly half of
the outer portion of the wing; a black cell-spot and black
veins ; cilia grey. Underside paler grey, with very broad
black borders to both wings: fore wings with a black dis-
coidal spot and another above it close to the costa: hind wings
with a large black discoidal spot.
Jixpanse of wings 1} inch.
Hab. Singapore.
Received with several examples of S&. plagifera, Walker,
Journ. Linn. Soe. vii. p. 187 (1864), and described by Walker
asa Thirmesia; type in Mus. Oxon., and apparently over-
looked and omitted in Phal, xi.
Stictoptera wetterensis, nov.
6. Fore wing narrow and long, the outer margin very
oblique and but slightly convex; head, body, and fore wing
dark pinkish grey, thickly irrorated with black atoms: fore
wing with a short black linear mark below the cell-end, a
shorter one at the end, a transverse similar mark near the
hinder angle, a longer similar mark parallel with the costa at
the apex, and an obscure blackish mark near the hinder
margin one-third from the base: hind wing dull white, the
veins black; a fairly broad even black band on the outer
margin. Underside dirty white, all the veins black: fore
wing nearly all blackish, the whitish part confined to the
space below the cell: hind wing with a discoidal black line
and blackish borders as on the upperside.
Expanse of wings 14 inch.
Hab. Wetter Island, South-west Islands, Amboina.
‘Lhe Amboina examples are almost identical with that from
Wetter ; when more material comes to hand the genitalia
must be examined to determine its exact position.
Stictoptera tongloana, nov.
3d. Head, body, and fore wings greyish brown with a
slight pinkish tinge: fore wings with a number of indistinct,
transverse, blackish, waved lines; a black spot inwardly
white-edged at the end of the cell, a small black mark below
the cell beyond its middle, another rather larger beyond it,
with a small one above it eontinucd upward ina waved linear
form to near the costa, a similar submarginal disjointed row
of black marks, and two black round spots at the apex; all
the other marks more or less lunular and encircled by a paler
ground than that of the rest of the wing; a row of pale
blackish lunules, inwardly pale-edged, close to the outer
Species of Indo-Malayan Lepidoptera. 339
margin, and small dark black lunules, inwardly pale-edged,
on the margin: hind wings smoky white, veins black ; a very
broad, even, black marginal band, occupying one-third of the
wing-space ; cilia w hite. Underside much as in wetterensis.
Hixpanse of wings 14 inch.
fab. 'Tonglo, Solomon Islands.
Stictoptera dispar, nov.
3 ¢?. Palpi, head, body, and fore wings dark chocolate-
brown, nearly black; palpi grey in front: fore w ing with the
base and outer portions slightly paler, markings very in-
distinct ; a transverse, somewhat oblique, and very indistinct
band, postmedial, parallel with the outer margin, and beyond
the reniform, apaler band adjoining its outer side somewhat
reddish-tinged, with some obscure black spots init, and black
Junular marks. on the margin: hind wings smoky white, the
veins black ; outer margin broadly and evenly black, occu-
pying more than one-third of the wing. Underside of the
usual pattern, but the fore wing has four rather prominent
white spots on the costa before the apex, and the hind wing a
prominent discal lunular bar which runs up to the costa.
Expanse of wings 1,5 inch.
Hab. Mt. Kebea, Brit. N. Guinea, 6000'.
I have four examples which I received as S. macromma,
Snellen (fiom Celebes), but they do not correspond with
Snellen’s figure or description, or with Hampson’s description
in Phal, xi. p. 162.
Stictoptera commutata, nov.
¢. Fore wings much as in dispar, but there is a very
large round white spot below the cell at the base of vein 3,
which slightly enters the cell and also slightly crosses vein 2 ;
at the base of the wing there are some dull ochreous scales
and dull ochreous hairs covering the upper sides of the thorax,
two spots behind, and some on the first two segments of the
abdomen : hind wings and underside as in dispar.
Expanse of wings 1,95 inch.
Hab. Mt. Kebea, Brit. N. Guinea, 6000'.
Two examples.
Family Epiplemide.
Epiplema rhacina, nov.
3. Upperside of a uniform olive-brown colour: fore wing
with the basal half of the costa irrorated with dark brown; a
340 Colonel C. Swinhoe on new
double ring-shaped mark somewhat like a figure of 8 in the
cell, another at its end, and a third below the end, the last
two more or less connected ;. the basal half of the fore wing
is finely striated with brown thin striations ; there are indica-
tions of an outwardly curved brown antemedial line on the
fore wing, and a sinuous outwardly curved middle line con-
nected. with the two outer ring-marks; both wings with a
postmedial line, sinuous in the fore wing, its upper half very
deeply outwardly curved, double on the hind wing, waved
and very slightly outwardly curved ; a row of submarginal
brown spots on both wings; outer margin of the fore wing
somewhat excavated below the apex as in . moza, Butler *,
but not so deeply, and the two tails of the hind wing blunt
aud very short. Underside pale pinkish grey, both wings
with discoidal marks, double brown transverse lines rather
close together, and minute submarginal spots.
Expanse of wings 1,%, inch.
Hab. Khasia Hills.
Family Pyralide.
Crithote horridipes.
Crithote horridipes, Walker, Journ. Linn. Soc., Zool. vii. p. 183 (1864).
Selenis crinipes, Suellen, Tijd. voor Ent. xxiii. p. 109, pl. viii. figs. 4, 4a
(1880).
Type, Sarawak, Borneo, in Mus, Oxon.
Type ertnipes, Bonthain, Celebes, in coll. Snellen.
Apparently a very widely distributed species. I have it
from Gilolo, the Khasia Hills, and from N. Kanara, 8. India ;
there is no appreciable difference in any of them.
Avitta subsignans.
Aritta subsignans, Walker, xv. 1675 (1858).
Oroba surrigens, Walker, Journ. Linn. Soc., Zool. vii. p. 81 (1864).
Epizeuxis inductalis, Snellen, Tijd. voor Ent, xxiii. p. 180 (1680), and
xxiv. p. 68, pl. vi. fig. 8 (1881).
Avitta fasciosa, Moore, Descr. Ind. Lep. Atk. p. 194, pl. vi. fig. 26 (1882).
Type, Kanara, 8. India, in B. M. —
Type surrigens, Sarawak, Borneo, in Mus. Oxon.
‘Type inductalis, Makassar, Celebes, in coll. Snellen.
Type fasciosa, Khasia Hills, in coll. Staudinger.
Another widely-spread species. Snellen records it from
Java, and I have received it from Sumba Island, Java,
Goping, Perak, Coomoo (Queensland), the Andaman Islands,
* Ann. & Mag. Nat. Hist. (5) i. p. 402 (1878).
Species of Indo-Malayan Lepidoptera. 341
Bombay, Nilgiri Hills, and the Khasia Hills—all apparently
identical ; whether the examination of the genitalia will bear
this out remains to be proved.
Osericana albistella.
Osericana albistella, Walker, xxxiv. 1214 (1865).
Pinacia pupilialis, Snellen, Tijd. voor Ent. xxviii. p. 7, pl. i. fig. 7
(1885).
Hab. Sumatra.
Both types are from Sumatra. I have also a pair from
Nias. The fore wing of the male is much paler than that of
the female, the hind wing of both sexes very pale in colour.
Osericana albistella trypheropa, nov.
3 ¢. Both wings of a uniform purplish grey, the hind
wing perhaps a shade lighter in colour than the fore wing,
the pectinations of the long antennz more robust than in
albistella, the markings similar,
Expanse of wings, ¢ ?, 1,%5 inch.
Hab, Palawan, Philippines; 1 ¢,3 2.
Osericana albistella syntypistis, nov.
36 ¢. Uniformly smaller than either of the foregoing
forms ; the colour of the hind wing about the same as in
trypheropa, the colour of the fore wing very much darker ;
the abdomen with more greyish suffusion, the yellow anal
tuft entirely black on the upperside ; in the other two forms
there are only a few blackish hairs.
Expanse of wings, ¢@ 17g, 2 1)5%-1)% inch.
Hab. Lawang, EB. Java; 1 o,4 2.
Simplicia schaldusalis.
Bocana schaldusalis, Walker, xvi. 180 (1858).
Culicula himarginata, Walker, Journ. Linn. Soe. vii. p. 178 (1865).
Simplicia infausta, Felder, Reise Nov., Lep. pl. exx. tig. 45 (1873).
Nabartha marginata, Moore, Lep. Ceylon, iii. p. 234, pl. clxxvii. fig. 2
(1885).
Simplicta griseolimbalis, Snellen, Tijd. voor Ent. xxix. p. 47, pl. ii. fig. 4
(1886).
Hab. Walker’s and Felder’s types are from Sarawak,
Borneo, Moore’s from Ceylon, and Snellen’s from Sumatra.
It appears to be a very widely spread form; I have it also
from the Solomons and from Obi Island in the Moluccas,
and without examining the genitalia [ can find no difference
between them.
342. On new Specivs of Indc-Mulayan Lepidoptera.
Family Pyraustide.
MarGAroniin a.
Margaronia alboscapulalis, nov.
Glyphodes alboscapulatis, Kenrick, MS.
¢ 2. Upperside: head and shoulders black ; a white spot
on the collar; body black, a short tuft of white hairs on each
side from the base of the thorax: fore wings black ; a white
spot below the costa a little before its middle ; a large, oval,
discal white patch as in MW. doleschali, Lederer: hind wings
white, with a broad black band, narrowest on the costa, very
broad at the apex, narrowing somewhat hindwards to the
anal angle. Underside: palpi and body white ; abdomen of
the male with some black marks, anal tuft black, of the
female with the lower half black ; legs white.
Expanse of wings, 3 2, 1;4-1,4 inch.
Hab, Ekeiki, Mt. Kebea, Brit. Cent. N. Guinea.
A fine series of both sexes, allied to M. doleschal’, Lederer,
but is easily distinguishable by its white hind wings, dole-
schali having black hind wings, with a very large, almost
round, white spot.
SYLEPTINZ.
Sylepta zarialis, nov.
gd. Cream-coloured, almost pure white, but not shining ;
palpi chocolate-brown above: fore wing with the costa pale
chocolate, outer marginal fine line, and a little apical suffusion
of the same colour very pale: hind wing with the outer
marginal line very faintly touched with the same tint of
colour; otherwise the head, body, wings above and below,
and the legs without any markings.
Expanse of wings, g, 1 inch.
Hab. Dinawa, 4000', Brit. N. Guinea.
PYRAUSTINZ.
Aphytoceros subflavalis, nov.
3. Pale yellow ; head and body without markings ; abdo-
men with the anal tuft black ; a small brush of yellow hairs
in its middle. Wings above uniform pale yellow, markings
pale chocolate-brown : fore wing with two outwardly oblique
sinuous lines, two more antemedial, more close together ; a dot
in the cell, two short lines from the costa across the end of the
cell, curved towards each other, with a darker line between
On the Lemurs of the Hapalemur Group. 343
them; two more or less sinuous lines from the inner end of
the above, straight to the hinder margin, slightly more sepa-
rated hindwards ; two postmedial lines from the costa to a
little below vein 2, somewhat separated from each other at
the costa, connected near its end by a thin sinuous line with
the lower end of the discoidal lines, and four small rings
attached to the outer half of these two lines ; some marginal
spots and a little suffusion below the middle of the space
between: hind wing with a dark lunular line at the end of
the cell ; two lines from the middle of the costa extending
hindwards towards the ‘anal angle, the lines anastomosing
halfway down ; a sinuous line from the costa near the « apex
to the anal angle, where it somewhat thickens ; some suffusion
at the apex of ihe wing; both wings with dak marginal line
and yellow cilia, interlined by a pale brown line. Underside
pale glistening yellowish white, the markings of the upperside
more or less indicated. Body and legs yellow without any
markings.
Expanse of wings, ¢, 1,5 inch.
Hab. Arfak Mts. 4 6000), 'N. New Guinea.
Not unlike a very large "Margaronia ceesalis, Walker.
XXXI.—The ania ar the Hapalemur Group.
By ht. I. Pocock, F.R.S.
On Hapalemur and Prolemur.
Tw addition to the skeleton and skull of the Hapalemur simus
described by Beddard (P. Z. 8. 1901, pp. 121-129), the
Zoological NSociety’s eolation contains the following ma-
terials. of Flapalemur, all the specimens being labelled
H. griseus, Madagascar, without nearer locality :—
1. The skull of an adult but small specimen, without
history of any kind.
2. The skin of a small, probably immature, male specimen
which was received in Nov. 1837, and has never been
described.
3. The skin of the adult male described by Beddard
(P. Z. 8S. 1884, pp. 391-399), and later by Bland
Sutton (P. Z. 8. 1887, p. 369).
4. The skin and skull of an adult male dated June 9th,
1903, to Sept. 17th, 1904, which like no, 2, has never
been described.
344 Mr. R. I. Pocock on the
This paper is based primarily upon an examination of these
examples.
The two species above named have quite an extensive
literature. Skulls assigned to H. griseus have been figured
on several occasions. ‘l'o these and to the specimens in the
Society’s collection I shall revert later.
Good figures of the skull of HZ. stmus have been published,
notably by Gray (P. Z. 8. 1870, pp. 829-830, figs. 1-4), by
Jentink (Notes Leyd. Mus. vil. 1885, p. 33), by Milne-
Edwards and Grandidier (Hist. Nat. Madag. Mamm., Atlas
ii. pls. exxii. G & H, 1890-1896), and by Elliot (Mon.
Primates, i. pl. xvii., 1912); and it may be noted that these
figures attest no structural variations of moment, suggesting
that more than one form has been described under the name
simus. As I shall presently attempt to show, this is not the
case with skulls ascribed to ZZ. griseus.
The generic name Hapalemur, proposed in 1851 by
I. Geoffroy for the species then known as Lemur griseus,
met with universal and unchallenged acceptance until 1912,
when Eliiot, misled by a superficial inspection of the text,
substituted Jdioxicebus—emended to Myowicebus—on the
alleged, but entirely erroneous, grounds that Lesson in 1840
had given the latter title to the type-species of Hapalemur.
It is quite true that the first species cited under Mowxicebus
was named griseus ; but it is equally and obviously true that
the diagnoses, both generic and specific, of Mioaicebus griseus
have no applicability to Hapalemur griseus. On the contrary,
they fit tolerably closely the species for which they were
intended, namely, Chircgaleus major, then known as mili.
It is possible that Lesson had at the time a specimen of a
different but closely allied species of Chirogaleus before him ;
but until evidence on that head is forthcoming M/voaicebus
griseus must stand as a synonym of Chirogaleus major.
Hapalemur consequently resumes its former place in litera-
ture.
* Another unnecessary change introduced by Elliot into the nomen-
clature of lemurs is the substitution of the new name Altiéidemur for
Opolemur on the alleged grounds that Gray applied the latter generic
term to Chirogaleus milii. That is an incorrect interpretation of the
facts. Opolemur (P. Z. S. 1870, pp. 853-854) was proposed by Gray for
a species represented in the British Museum by specimens which he
wrongly identified as Chirogaleus milii. That his identification was
erroneous is shown by the diagnosis and figures. The characters, stated
and illustrated, of his Opolemur do not fit Chirogaleus; hence the former
cannot be a synonym of the latter, as Elliot asserted, and Opolemur must
be restored to use, if the genus it designates is maintainable, with Altii-
lemur as its synonym.
Lemurs. of the Hapalemur Group. 345.
So far as I am aware, the only other name which can come
generically into the little group of lemuroid species exemplified.
by griseus of I. Geoffroy is Prolemur, which was used by
Gray first in a subgeneric, then in a generic, sense for the
species he described as simus. It appears to me that full
generic rank should be assigned to this form. The characters
upon which this opinion is based have been either figured or
described by previous authors—notably by Gray, Beddard,
Milne-Edwards, Grandidier, and Elliot,;—who, however, did
not attach so much importance as I do to the differences
between griseus and simus where they were appreciated *,
These differences appear to me to be of considerably greater
systematic value than those which distinguish such genera
as Chirogaleus and Microcebus, for instance.
To our knowledge of Prolemur simus I have nothing to
add. In the subjoined comparative diagnoses of Hapalemur
and Prolemur IT have merely made use of characters in Pro-
lemur which have been stated by others or are apparent in
their published figures.
Hapalemur, Geoff.
Type, griseus, I. Geoffr.
Gland on forearm present in
both sexes,
Nasals long, extending back be-
yond lacrymal foramina.
Interorbital constriction not ex-
ceeding half the width of the post-
orbital constriction.
Mesoptery goid fossa much longer
than its greatest width in front.
Width across paroccipital pro-
cesses at most a little greater than
leneth of nasals. ;
Malar orifice large, set back be-
hind middle of orbit.
Symphysis of mandible strongly
eurved, chin rounded,
Ramus of mandible
everted behind dental line.
Upper pm? much lower than
canine, a little higher than pm? ;
pm and pm? unlike in size and
slightly
Prolemur, Gray.
Type, stmus, Gray.
Gland on forearm present in
neither sex.
Nasals short, not extending back
to level of lacrymal foramina.
Interorbital constriction con-
siderably more than half the width
of the postorbital constriction,
Mesopterygoid fossa shorter than
its greatest width in front.
Width across paroccipital pro-
cesses much greater than length of
nasals,
Malar orifice small, set forwards
nearly in line with middle of orbit.
Symphysis of mandible not
strongly curved, chin flattish.
Ramus of mandible strongly
everted behind dental line.
Upper pm' slightly lower than
canine, much higher than pm’;
pm? and pm? approximately alike
* Guay’s opinion, for example, that the species described by Schlegel
as Hapalemur griseus was the same as his 1. simus attests failure in this
respect on his part ; and Beddard, when he suggested that Mivart had
identified stmus as griseus, must haye overlooked that author’s description
of the teeth.
23
Ann. & Mag. N, Hist. Ser. 8. Vol. xix.
346
Hapalemur, Geoffr.
form ; pm® molariform, with quad-
rate inner lobe; m and m? with
simple cingulum, without accessory
cusp; no trace of groove on the
inner cusp of these teeth behind,
Legs shorter; skull about six-
sevenths the length of the femur *.
Mr. R. I. Pocock on dpe
Prolemur, Gray.
in size and form; pm® not molari-
form, with rounded inner lobe;
m' aud m? with bilobate cingulum,
the posterior lobe cuspidate; the
main inner cusp of these teeth
grooved posteriorly.
Legs longer; skull about two-
thirds the length of the femur.
On the Species of Hapalemur.
When Hapalemur was instituted two species were assigned
to it by Geoffroy—namely, griseus and olivaceus. The latter
was said to differ from the former in colour and in the shape
of the lower jaw. Most subsequent authors have concurred
in the specific identity of the two, and Milne-Edwards and
Grandidier, who had access to Geoffroy’s specimens, called
olivaceus a variety of griseus; and their coloured figures show
that griseus is lighter in tint than olivaceus. Elliot, however,
admitted the two species because of the difference in colour
and the larger size of the skull in o/ivaceus. The inference to
be drawn from the literature, whether rightly or wrongly, is
that the two forms may represent distinct subspecies, or
possibly species, but that in any case they are closely related
and exhibit few, if any, constant cranial differences except of
size.
The three skins in the Zoological Society’s collection are
decidedly dark in tint, and may be described as dusky brown,
the hairs being dark bluish grey annulated with rusty brown
towards the tips. On the crown of the head the rusty brown
is more in evidence, but round the eyes and on the cheeks
it is less obvious. ‘The underside is lighter than the upper.
In the small specimen, received in 1887, the belly and thighs
inside are bright buff, the throat grey. In the two others
the throat is darker and the belly dark grey washed with
brown. Provisionally I regard these skins as belonging to
one and the same species and race, and the colouring enforces
* Judging from M.-Edwards’s figures of the skeletons of H. griseus
and P. simus. For instance, in H. griseus the skull measures 73 mm.
and the femur 90; in P. stmus the skull is 81 mm. and the femur 120.
In the Zoological Society’s specimen of P. simus the femur is actually
a little longer, being 122 mm. to the head, whereas the skull is shorter,
namely, 75mm. The skeleton, however, is that of an immature speci-
men, with the last molar teeth still buried in the bone, as Beddard’s
figure indicates. Probably the skull would have increased in length
proportionately much more than the femur.
Unfortunately the leg-measurements of H. schlegeli are unknown.
Hence the character above stated can only be used provisionally in a
generic sense.
Lemurs of the Hapalemur Group. 347
the conclusion that they are the olivaceus-form of griseus, and
not typical grzseus. This conclnsionis borne out by the skull
of the example received on 9. 6. 03, which is a little larger
than the skull of griseus figured by Milne-Edwards and
Grandidier. It also has the muzzle less steeply inclined, the
posterior half of the zygomatic arch a little more arcuate, and
the glenoid a little lower with reference to the dental line.
I have not sufficient material to judge of the systematic value
of these differences. Otherwise the two skulls are very much
alike ; and it is possible that M.-Edwards’s illustration, as
suggested below, was taken from an example of what he
called the olivaceus variety of griseus.
I stated above that skulls of specimens assigned to JHapa-
demur griseus have been figured on several occasions; and
the figures indicate confusion of more than one form under
that name. For instance, if the figure of the skull published
by Schlegel be compared with that published by Milne-
Edwards and Grandidier, it will be seen that the differences
between them fall quite outside the limit of individual varia-
tion exemplified by Prolemur simus or by any single species
of the Lemuride known to me. Gray, indeed, declared that
Schlegel had drawn the skull of an example of Prolemur simus
in mistake for Hapalemur griseus. With this opinion
Beddard was disposed to agree, and Jentink tried to account
for the error of this view by explaining that Schlegel’s
illustration was inaccurate, apparently because it did not
agree with the skulls that he possessed. Doubtless it did
not; but in my opinion Schilegel’s figure was exact in all
essential points, seeing that it agrees singularly closely with
the adult skull in the Society’s collection mentioned first on
my list in the opening paragraph of this paper.
Similarly, the skull of the specimen that lived in the Gardens
from June 1903 to Sept. 1904 agrees in the main, though not
so closely as in the other case, with the skull of LZ. griseus
figured by Milne-Edwards and Grandidier. Since these
French authors had access to Geoftroy’s type of griseus, it
must be assumed that the example they identified as griseus
belonged to that form or to olivaceus, which was regarded as
the same, and that Schlegel’s example was wrongly referred
to griseus. Confirmation of this conclusion was supplied by
Elliot, who also saw the specimens in the Paris Museum, and
remarked in connection with Schlegel’s illustration :-—* This
figure is badly drawn, or does not represent the skull of
H. griseus. It is altogether too broad, especially the muzzle.”
From this passage it seems that Llliot was not prepared
altogether to accept Jentink’s verdict as to the inaccuracy of
Z3*
348 Mr. R. I. Pocock on ‘the
Schlegel’s figure, and that the possibility of another species
being concerned dawned upon him. Nevertheless, the short-
ness of the muzzle misled him apparently in the matter of its
apparent superior width.
Both the literature, therefore, and the skulls in my
possession attest the existence of two well-marked species of
/lapalemur—one exemplified by the small skull above referred
to, which probably belongs to the form Schlegel identified as
H, griseus, the other being the true griseus of Geoffroy, which
has been well figured by Milne-Hdwards and Grandidier.
‘The former species I propose to describe as new, taking the
skull in the Zoological Society’s collection as the type. Since
the only other skull I have at hand is that of the specimen
determined, for reasons already stated, as olivaceus, I have
diagnosed the new species with special reference to olivaceus
rather than to griseus, although the differences between it and
Milne-Edwards’s figure of the skull of griseus are almost as
well marked. .
Hlapalemur schlegeli, sp. n.
¢ Hapalemur griseus, Schlegel, in Pollen & Van Dam, Rech. Faune de
Madag., Mamm. et Ois. p. 6, pl. vii. figs. 4 a-d (skull). Nec H. gri-
seus, 1s, Geoff.
Skull (type) considerably shorter but relatively broader,
higher, and more arched antero-posteriorly along its upper
profile, and less hollowed between the postorbital processes,
than in #., olivaceus, the orbits relatively larger, with the
inferior edge much more salient, giving a strongly sinuous
curvature to the outline of the malar arch, and causing a
deeper groove along the outer surface of its suborbital
portion ; the upper surface of the muzzle more depressed and
curved, the upper portion of the maxilla compressed along
the nasal suture, the lateral edge of the anterior nares emar-
ginate in profile view, this orifice slightly higher than wide,
compressed above. In H. olivaceus the muzzle and anterior
nares are not compressed above and the latter orifice is slightly
wider than high. The zygomatic arch and postorbital bar
are relatively stouter than in olivaceus ; the mastoid is inflated,
reducing the paroccipital process, and the upper edge of the
zygoma is not continued as a crest back to the occiput as it
is in H, olivaceus, where the mastoid is not inflated but flat,
leaving the paroccipital processes salient. The basicranial
axis is more steeply inclined, so that the bull and occipital
condyles are set considerably lower with reference to the
alveolar border of the maxilla than in olivaceus *.
* This difference is not so marked between the skulls of H. schlegeli
and H, griseus, judging from M,-Edwards’s figure of the latter.
Lemurs of the Hapalemur Group. 349
Teeth of H. schlegeli shorter and narrower.
The typical skull of A. schlegeli has fully erupted and
complete dentition and the sutures nearly obliterated. The
obliteration, however, has not extended to quite the same
extent as in the skull referred to . ol’vaceus. Nor is there
in the skull of A. schlegeld a median sagittal ridge on the
parietal region. ‘The low temporal crests are merely con-
fluent near the middle of the parietals. ‘The difference in this
respect may be due to difference of age ; but this is uncertain.
* When the two skulls are placed side by side on a flat surface
they are practically the same height, despite the considerable
disparity in length.
The differences in the shape and the dimensions of various
parts of the skulls may be appreciated from the subjoined
table of measurements of the type of schlegeli and of my skull
referred to olivaceus. In the third column are given the
dimensions taken from the figures of the skull named griseus
by M.-Edwards :—
| schlegelt. | olivaceus.| griseus. |
| | mm. mm. mm,
pS HOHL Aer GRg OYyiat i tidhe wane tian cia hy veka + OR 64 | 60
Length of palate along middle line ....) 25 Bl 29
Length from post. edge of postorb. bar
EOP AOE MME rn hora aise ws egal Hes | 36 a mats yf
Length from post. edge of orbit to
Facrymeal TOr@Met |. /0% hes als & oss 19 19 19
PSA etPL OE OF DUG stets les yd «54s we was 14 14 14
Height from alveolar border to lower
RTE CE IEE alata sere arwratie y batt Coss | 9 12 10°5 |
PNVsid cle ORCraniG nn 5: ban a's es te mae 0 bl 33 Ce
Width of postorbital constriction ...... 23 19 LSet
Width of interorbital constriction...... 8 10 10
Width across zygomata (postorbital) .. 44 A bh ee
[> WV deltas AeNOSS ORBUGA cei e's Sta 218 a6 fs 39 BO ce hs AG
| Width of muzzle above canines........ 14 17 LGAs 4
Length of mandible from condyle.......; 41 . 62 48
| Width of upper pm®.........5.. rsh Fae 4-5 55 Os}
Skulls assigned to H. griseus have also been figured by
Jentink (Notes Leyden Mus. vii. pls. 1. & 11. figs. 3-4, 1885)
and by van der Hoeven (Tijds. Nat. Geschied. 1844, pl. i.
fig. 1); but in both cases there are discrepancies in the
dimensions of the superior and lateral views which make it
impossible to tabulate the measurements. Tor instance, in
the case of Jentink’s specimen the superior view of the skull is
65.mm., the lateral view 62°5, whereas the lateral view of the
mandible from the condyle is 44 and the superior view 40,
350 Mr. R. I. Pocock on the
In Hoeven’s figure the superior view of the cranium is 59,
the lateral viewis 63. It may be noted that in M.-Edwards’s
fivure of the skull of griseus the measurements coincide, as
should be the case, both from the lateral and superior aspects.
Turning to Jentink’s text, we find it stated that sixteen
adult skulls measured 61 mm. in total length and 42 in width
across the zygomata. They are thus considerably smaller
than the skull of griseus on by M.-Edwards, which is
73 mm. long and 48 broad, while my olivaceus is 76 mm.
long and 49 broad. Clearly, therefore, Jentink’s skulls were
considerably smaller than the one depicted by Milne-Edwards
and than the one I have described as oltvaceus. ‘This suggests
the possibility of Milne-Edwards having described a skull of
olivaceus as griseus, a course he might very well have adopted,
seeing that he reg: arded ulivaceus merely as a variety of griseus.
Avain, if the specimen figured by Jentink be a true ‘sample
of the sixteen he had for examination, they all differ from my
olivaceus and Milne-Hdwards’s griseus in having a very much
thinner postorbital bar. "Chis, however, like the smaller size,
may be a matter of age. Moreover, it will be noticed that
the temporal crests are subparallel, showing scarcely any sign
of convergence as far back even as the interparietal region,
whereas in my ol’vaceus and M.-Edwards’s griseus these ridges.
coalesce and form a fairly strong sagittal crest over the
middle line of the parietal region.
But, whether Jentink’s skulls represent a form distinct
from M.-Edwards’s griseus, or are merely less well-developed
individuals of the same species, it is quite clear they are not
referable to the same form as the one f have named schlegel/.
They are too long and narrow, have very slender postorbital
bars, and the frontal bones are depressed as in my skull of
olivaceus.
There is no occasion to publish a figure of the type-skull
of H. schlegeli, since it is in almost punetilious agreement
with Schlegel’s illustration, which shows the inflation of the
mastoid, the sinuous curvature and suborbital salienee of the
malar arch, the thickness of the postorbital bar, the large
orbits, the cranial width, the curvature of the upper profile,
the shortness of the muzzle, ete. One rather marked difference
in the tip of the muzzle may be explained, I suspect, by the
cutting away of this part of the skull in Schlegel’s example
when it was removed trom the skin. The incisor teeth are
missing, as others have remarked, and this defect suggests
that a portion of the premaxilla may have been cut away.
If so, the ends of the nasals may have been truncated at the
same time. ‘This, however, is merely a suggestion. In the
Lemurs of the Hapalemur Group. 351
type-specimen also the angle of the mandible is less rounded
and the upper end of the coronoid is thinner, longer, and less
curved than shown in Schlegel’s figure.
The specimen described by Schlegel as H. griseus was
discovered by Pollen at Ambassuana, three days’ journey
from the north-west coast of Madagascar. If, as I suspect,
the type of H. schlegeli belongs to the same species, it
probably came from the north-west coast of Madagascar,
possibly also from Ambassuana.
The Arm-glands of Hapalemur.
The presence of glands on the forearm in Hapalemur
griseus—or, rather, olivaceus, for such one of the specimens
proves to be—was first pointed out by Beddard, who also
ascertained, from Jentink and Milne-Edwards, that no such
glands are developed in Prolemur stmus. ‘This character
alone is sufficient, in my opinion, for generic separation of
the two species.
In the two male specimens of LZ. olivaceus* in which he
described the glands, he pointed out that the naked tract of
skin above the wrist was covered with long and coarse
papille ; but, judging from his figures, the papillae were much
better developed in the first specimen examined than in the
second.
In the two other skins in the Society’s collection, which
Beddard did not see—namely, the small one received Nov. 10,
1887, and the adult received June 9th, 1903,—the gland
differs in that the tract of integument is comparatively
smooth, being merely roughened, so far as can be judged on
the dried skin, with fine granular papilla.
With regard to the glands on the upper arm, regarded by
Beddard (but, I think, wrongly) as mammee, I can find no trace
of them in the small and presumably immature skin ; and in
the adult skin with the glandular tract of the forearm nearly
smooth they are less well developed than in the specimen in
which-they were first detected—namely, the one with the
glandular area of the forearm exceedingly coarsely papillate.
I do not think any special importance should be attached
to these differences, because in Lemur catta, which possesses
similar glands, the spur on the glandular tract of the forearm,
which may be compared to the papilla in Hapalemur, varies
* T have the dried skin of the example described by Beddard in 1884.
Of the second specimen described in P. Z. S. 1891, p. 449, and 1902,
p- 159, no history was given, and the skin was not preserved. Probably
it was olivaceus,
359 Mr) 0. Thomas—2 Some eles
,
in development with age, being larger in older specimens,
and the gland of the upper arm is, I believe, subject to
seasonal clianges in size.
The occurrence of similar glands in two such widely
different species as Lemur catta aud Hapalemur griseus, and
their complete absence in the other species referred to Lemur
and in Prolemur simus, is a remarkable fact.
XXXII.—Some Notes on Three-toed Sloths.
By OLpFIELD ‘THOMAS.
(Published by permission of the Trustees of the British Museum.)
To those whose interest it is to compare zoological characters
in their relation to geographical distribution no group of
Mammals is so unattractive as the sloths, on account (1) of
their variability, especially in the skull, in specimens from
the same place, (2) the slight and intangible characters that
distinguish specimens from the most distant localities, and
(3) the great state of confusion that has resulted from the
descriptive efforts of Wagler, Gray, and Fitzinger. Early
descriptions, without statements of locality, have been made
the basis of various names, and it is a matter of the greatest
difficulty to disentangle the confusion.
The present notes make no pretence of being complete, and
are purposely worded somewhat vaguely, as such are the
difficulties of the case that there is hardly a statement J can
make which may not prove liable to modification as fuller
series from all localities are studied.
Firstly, with regard to genera, I am disposed to recognize
the collared sloth, Bradypus torquatus, D.smarest, 1817 (not
Llliger, as usually quoted, for the latter author’s two refer-
ences are both nomina nuda), as forming a special genus,
which may be distinguished by the inflated pterygoids, better
developed premaxille, the median spout-like projection on
the mandible, and the absence of a dorsal gland or “ speculum ”
in the male*, all these characters being asin Cholapus. The
5
generic name of Sewopus, Peters, is available for it.
* T can by no means subscribe to Dr. Allen’s conclusion (Bull. Am,
Mus. xx. p. 339, 1904) that “the presence or absence of this highly
differentiated patch is not sexual ’’—a conclusion based on what I must
cousider the incorrect sexing of certain “ females” by one of his collectors,
Not only has it long been generally accepted that the speculum is charac-
teristic of the male, but I find that in every specimen without speculum
in our collection mamme are to be found, while in no example with
speculum is there any trace of them. With so large a collection, in-
cluding so many different forms, this evidence appears to me conclusive.
on Three-toed Sloths. 353
Synonyms of Scwopus torquatus are crinitus, Gray, 1849
(ex Browne), and affinis, Gray, 1849. ‘The species is con-
fined to South-western Brazil, but -the limits of its range are
not known. Mr. A. Robert sent a nice series of it to the
British Museum from Engenheiro Reeve, Hspirito Santo, in
1903.
Passing now to true Bradypus, we find the question of the
identification of its type-species, tridaetylus, Linn., a question
so productive of confusion among the early writers, has now
been settled by the fixation as the typical locality of the latter
as Surinam *—so that the Guianan species should bear the
name tridactylus.
B. tridactylus is the best-marked of the species of the
genus, being readily recognizable by the extension of the
yellow colour of the face down the front of the neck—a
character to which no approach is shown by any other species.
Its skull is rather small and has generally a pair of peculiar
fosse or perforations in the floor (or, more strictly, the roof)
of the anterior part of the mesopterygoid fossa ; but, although
so striking in well-marked cases that a person might excusably
think it a character of generie value, this modification varies
in different skulls, and is sometimes practically absent. ‘The
teeth are of average proportional size, the pseudo-canine T
well differentiated, and the pseudo-incisor T small, usually
about a quarter the size in section of one of the molars.
Wagler (1831) was the first properly to distinguish this
species, to which he gave its current name of #. cuculliger,
while other synonyms of it are Acheus ai, Less.t (1827) ;
guianensis, Blainv. (1839), a name doubtfully valid techni-
cally ; gu/aris, Riipp. (1845); and eristatus, Temm., Fitz-
inger, 1871.
B. tridactylus ranges over the whole of Guiana—French,
Dutch, and British,—and our collection contains a good series
of it, mostly presented by Mr. F. V. McConnell.
* See Thomas, P. Z. 8. 1911, p. 182.
+ These names are used respectively for the second and first upper
teeth, which, really corresponding to the anterior premolars of ordinary
Mammals, take on in the three-toed sloths something of the relative
proportions of a canine and an incisor. The three remaining teeth on
each side, acting as a premolar—-molar series, are subequal, smaller than
the pseudo-canine, larger than the pseudo-incisor.
¢ This name was given to the “ Bradypus tridactylus, L.,” of Des-
marest, which included all the forms of true Bradypus then known. It
seems best placed as a synonym of ¢77dactylus, especially as the animal is
said by Desmarest to be very common in Cayenne. For the Brazilian Ai
of Maregrav it would be the earliest name, but there would be difficulty
in justifying its use for that animal on techuical grounds,
354 Mr. O. Thomas—Some Notes
South of Guiana, from Para to Rio, and westwards up the
Amazon, there occur a very uniform series of forms which
may or may not be divisible into two or more species, but
which it is impossible at present to clear up without many
more sets of specimens. One locality only—Para—is well
represented, as M. Robert got a series there in 1904.
Throughout this area the sloths are rather larger than
B. tridactylus; the speculum is of normal size, as in that
species, and of a rather less dark yellow colour, the general
colour is brown mottled to a very variable extent with white,
and the band of yellow velvety hair which passes across the
forehead is usually about half an inch in breadth—say, 10-
15mm. The skull averages rather larger than that of tri-
dactylus, and has generally a much inflated frontal region.
The floor of the mesopterygoid fossa is usually flat, with or
without median septum, and without special perforations.
The teeth, although variable, are usually of what may be
called normal proportions, the pseudo-incisor smaller—gene-
rally much smaller—than the molars, and the pseudo-canine
decidedly larger than the latter.
‘he names for these sloths may be put in two groups—the
Amazonian and the 8.E. Brazilian—not that as yet I can see
any valid reason for distinguishing the two specifically, but
merely to help later workers.
(1) Amazonian: B. infuscatus, Wag]. 1831 (Brazil near
Peruvian boundary) ; brachydactylus, Wagn. 1855 (syn.
speculiger, Fitz. 1871) (Borba, Lower Madeira) ; proble-
maticus, Gray, 1849 (Para) ; unicolor, Fitz. 1871 (Para) ;
smithi?, Gray, 1869 (Para). .B. marmoratus, Gray, 1849
( Brazil’), seems also to belong here, judging by the type,
and, if the Upper Amazon tnfuscatus proves distinguishable,
would be the first name for the Lower Amazon form.
(2) S.E. Brazilian: B. ai, Wagl. 1831, nec Less. 1827
(R. Matheus, Espirito Santo) ; drasiliensis, Blainv.* 1839
(Rio Janeiro) ; pallidus, Wagn. 1843 (Rio Janeiro) ; blain-
villet, Gray, 1549 (Brazil) ; dorsalis, Fitz. 1871 (Pernam-
buco—based on Marcgrav).
Series from many localities are needed before these Bra-
zilian sloths can be properly worked out, the available speci-
imens—apart from M. Robert’s set from Para already mentioned
—consisting of isolated examples, often without any exact
locality at all.
Bradypus boliviensis, Gray, 1871 (type B.M. no. 46.7.28.24),
seems to be a valid species. Its speculum is of medium size,
* Put in valid form on p. 64 of the article on Bradypua,
on Three-teed Sloths. 355
rather dark-coloured. The hairs of its throat are tipped with
white, a point not seen in any other species, though probably
not of great constancy. Its skull is large and solidly built,
and the teeth are distinguished by the unusual character that
the pseudo-incisor is very large—as large as or even larger
than the pseudo-canine, which in turn is small, not exceeding
the posterior molars in transverse section. ‘These tooth-
characters are, however, not to be seen in Gray’s figure,
which seems to have been taken from a wrong specimen.
But that 46. 7.28.24 is the proper type there can be no
doubt, as its osteological number, 921. a, is quoted by Gray.
In Central America we have 2. castaneiceps, Gray, 1871,
of Nicaragua, and B. griseus, Gray, 1871, of Veragua, of
which we have no further material, and I ean add nothing to
Alston’s account of them except to note that Mr. Goldman
considers them really distinct, and adds to them a third
species—B. ignavus, from Panama and the Atrato River.
It is, however, certain that griseus and tgnavus at least are
very closely allied to B. flaccidus. It is also to be observed
that Dr. Allen has identified a sloth from Rio San Jorge,
Bolivar, Colombia, with Philippi’s B. ephippiger; and as an
example from Condoto, Choco, Colombia, presented by
Dr. Spurrell, agrees with Philippi’s figure in the great size
of the speculum, the breadth of the frontal band, the distri-
bution of the light and dark face-markings, and the size of
the teeth, I am disposed to accept Dr. Allen’s identification,
and put down ephippiger, which was described without exact
locality, as a native of N.W. Colombia, therefore in the same
region as “ B, ignavus” came from.
In any case, however, the relations to each other of
griseus, ignavus, ephippiger, and flaccidus clearly need much
further investigation.
B. flaccidus, Gray, 1849, has as type-locality Venezuela
(probably the region opposite Trinidad), and has as synonyms
dysont, Gray, 1869, and columbicus, Fitz., 1871.
But by what characters it can be positively distinguished
from the Brazilian forms I have not sufficient good material
to be certain.
From Ecuador the Museum contains, firstly, a set of five
adults and two young from Sarayacu on the Upper Pastasa
River, and, secondly, an adult from the Balzar Mts., Guayas
district, W. Ecuador. ‘These appear to me to represent twa
turms tor which no names are available. The first may be
called
356 Some Notes on Three-toed Sloths.
Bradypus macrodon, sp. n.
Most nearly allied to B. infuscatus, but the teeth heavier.
General colour pale brown, the white marbling of the
posterior back generally extensive, with a well-marked brown
median line, but in one specimen there is scarcely any white,
and in another but little. Face and chin brown, the light
frontal band narrow, about half an inch (say, 10-15 mm.) in
breadth, whitish rather than yellow in most of the specimens.
Hairs on crown overhanging the frontal band rather darker
than those on the body generally. Speculum of medium
size, 24 inches long in the type, paler yellow than in most
other species, its median black band broad. Limbs freely
marbled with white.
Skull much as in B. infuscatus, the forehead rather less
convex than usual. Nasals, as usual, quite variable in shape
and length, some convex and some concave anteriorly. Inter-
parietal also extremely variable, its antero-posterior diameter
in the type 15 mm., and in another specimen 7 mm. Meso-
pterygoid fossa widely open, its Hoor smooth and flat, or with
a slightly raised median ridge, but without deep pits or exca-
vations.
Teeth very large, much heavier than in ¢nfuscatus.
Pseudo-incisor very large, oval in transverse section, the
longest diameter pretty well equalling that of the molars,
4-3 mm. in the type, 5 mm. in another specimen. Pseudo-
canine similarly very large and heavy, its greatest diameter
about 6°2 mm., far exceeding the molars in bulk. Molars
rarely less than about 5 mm. in greatest diameter.
Skull-dimensions of type :—
Naso-occipital length 78°5 mm. ; condylo-basal length 78°3 ;
greatest breadth (on squamosal) 56 ; nasals, length 12:5,
least breadth 12°3; interorbital breadth 25°5; palate length
from gnathion 29; postpalatal length 43; tooth-series 29 ;
breadth between outer sides of pseudo-canines 22°6.
Hab. Sarayacu, Upper Pastasa River, Oriente of Ecuador.
Type. Adult male. B.M. no. 80.5.6.56*. Collected by
Clarence Buckley. Seven specimens examined,
This species is no doubt most nearly allied, as is natural, to
B. infuscatus, but differs by the materially larger size of the
teeth. Even in this character, however, one specimen fails ,
(skull no. 80. 5. 6.59); but among sloths such variations
must be expected.
* The skulls were not individually allocated to the skins when they
came, but I think there is little doubt as to the reference to each other
of the type-skin and skull.
Geological Society. 357
Bradypus violeta, sp. n.
Aliied to B. macrodon, but distinguished from that, as from
every other sloth, by the great extent of the yellowish velvety
hair of the face, which extends upwards on the crown to the
level of the ears nearly 60mm. fromits anterior commencement ;
the hairs of the cheeks as far as the ears also of the same
colour and quality. Sides of throat likewise inclining to
yellowish, in continuity with the cheeks, but the chin dark
brown as usual, and the dark eye-streak also present. Nape,
shoulders, and middle line at withers brown, the rest of the
back nearly wholly white, as are also the hind limbs. Fore
limbs brown, marbled proximally with white. Under surface
dark brown anteriorly, whitish brown posteriorly. Speculum
uot known, the only specimen being a female.
~ Skull like that of B. macrodon, the forehead a little more
swollen. Teeth of similar size and proportions.
Skull-measurements of type :—
Naso-occipital length 72 mm.; greatest breadth (on juga!)
49°5; nasals, length 14, least breadth 12°3; interorbital
breadth 24; palate length from gnathion 28-5 ; tooth-series
27:2; breadth between outer corners of pseudo-canines 20 ;
greatest diameter of pseudo-incisor 4, pseudo-canine 6, second
molar 4°8.
‘Hab. Balzar Mountains, Guayas, W. Ecuador.
Type. Adult female. B.M. no. 80.5.6. 83. Collected by
Mr. Illingworth.
Characterized by its nearly wholly velvet-clothed face.
ee
PROCEEDINGS OF LEARNED SOCIETIES.
GEOLOGICAL SOCIETY.
January 24th, 1917.—Dr. Alfred Harker, F.R.S., President,
in the Chair.
Mr. Scoresby Rovurieper, M.A., gave an account of Easter
Island. He-said that the Expedition, that he had had the
honour to command, was organized with the object of carrying
out a long-standing wish of various bodies interested in anthro-
pology. This wish was that Easter Island, and other islands most
near to it, though far distant from it, should be thoroughly
examined, and that all information and material thereon found
should be carefully considered on the spot, or, if possible, be brought
back for comparative study.
This programme necessitated a vessel being specially designed,
built, and equipped for the purpose. A schooner with auxiliary
motor power, the ‘Mana,’ of 90 tons gross register, 78 feet
Lal
358 Geological Society.
on the water-line, 20 feet beam, and drawing 10°5 feet aft,
was accordingly completed by the end of 1912, and she sailed
from Southampton in February 1918 with a company of twelve
all told, of whom four formed the scientific staff. After the
longest voyage ever made by a yacht under canvas, she sailed
into Southampton again in June 1916, without having experienced
accident to man or material.
The course taken was through the Magellan Straits, and thence
through the labyrinth of Andean waterways that stretch north
therefrom, and are known as the Patagonian Channels.
On reaching Juan Fernandez Island, the ‘Mana’ had to put
back to Valparaiso because the geologist of the Expedition, the
late Mr. F. L. Corry, had contracted typhoid fever on the Chilean
coast. Mr. Corry never recovered sufficiently to allow him to
rejoin the Expedition. Hence no formal geological report on
the island could be submitted to the Meeting. It was thought
best, therefore, to endeavour to convey the conditions existent on
Easter Island by means of a series of panoramic and other photo-
graphs, specially taken to illustrate geological features. As these
very largely consist of coast-sections, the opportunity was taken to
show, and explain, other pictures that were closely associated with
them. Such were the ruins of the village called Orongo, consisting
of peculiar canoe-shaped houses built of imbricated slabs of shale,
with the roof convex, both longitudinally and transversely, on its
exterior aspect, and covered with earth. They are romantically
situated on the rim of the voleano of Rano Kao, with an almost
sheer drop of 900 feet into the sea, or of 600 feet into the crater-
lake. At Orongo, too, are found certain large rocks, carved with
the symbol of a bird-headed man, holding in its hand an egg. A
cult, based on annually obtaining the first-laid egg of a certain
migratory sea-bird, was thus gradually brought to light, and appears
to be a unique form. A brief outline only could be given of
some of the knowledge obtained concerning the peculiar routine
associated with seeking, and taking, the sacred egg, and of the part
which it occupied in the former religious life of the island.
Proceeding along the coast, typical examples of the great terraces,
and their giant stone figures, were shown, and their leading cha-
racteristics discussed. A submarine freshwater spring, near the
great image-terrace of Tongariki, and opposite certain typical
lava-formed caves, gave occasion to the lecturer to explain how
had arisen the longstanding, and world-wide spread report, that
man and beast on Easter Island habitually drink sea-water, in the
place of fresh.
The old voleano of Rano Raraku, the centre of the former
religious life of the island, was then described. A series of
panoramic pictures, preceded by an accurate survey made by
Lieut. R. D. Ritchie, R.N., the Cartographer of the Expedition,
showed a crater-lake surrounded by a rim of tuff which rises to a
height of 540 feet above the surrounding plain. The plain is
undulating in surface, formed superficially of hard, dense, but
nevertheless vesicular, lava, and it rests on compact non-columnar
Geological Society. 359
basalt. One section of this crater wall, some 600 yards long,
on both its interior and exterior aspects, was seen to be quarried
right up to the highest point. On the mountain-face, both inside
and out, large numbers of statues, in every state of completion,
were to be seen. The largest of these measured 68 feet in length..
Some of those excavated by the Expedition exhibited fine details,
such as the finger-nails, in perfect condition.
In conclusion, Easter Island might be described as a plateau of
basalt raised from 50 to 100 feet above the sea. Superimposed on
this were numerous cones ranging up to nearly 2000 feet. The
plateau was covered but sparsely with soil, and could only be
crossed with difficulty in any direct line. The cones, on the other
hand, were generally smooth of surface, with a good depth of soil.
Nevertheless the island is practically without trees, bushes, or
shrubs.
February 7th, 1917.—Dr. Alfred Harker, F.R.S.; President,
in the Chair.
The following communications were read :—
1. ‘The Trias of New Zealand.’ By Charles Taylor Trechmann,
M.Sc., F.G.S.
The fossiliferous Triassic rocks of New Zealand have been wholly
or in part at different times attributed by the geologists of that
Dominion to a Devonian, Permian, Permo-Carboniferous, Lower,
Middle or Upper Triassic, or Trias-Jura age. A review of the
previous research on these rocks and of their correlation and nomen-
clature is given. They are quite distinct from the Matai rocks,
which contain a Permo-Carboniferous fauna.
Triassic beds appear at intervals from Kawhia on the western
coast of the North Island to Nugget Point on the south-eastern
coast of the South Island—a distance of 620 miles. Except in two
localities, they are everywhere very steeply inclined, and where they
approach the Alpine Chain of the South Island pass into semi-
metamorphic greywackes or completely metamorphic phyllites and
schists. They are of great thickness. A short description of the
special faunal, lithic, and tectonic features of each of the more
important localities is given, all of which but one occur in the
South Island. In the North Island only the Noric and Rhetic
horizons have been recognized. Wherever the sequence is preserved,
the Trias passes conformably up into Jurassic deposits.
The lowest fossiliferous horizon of the Trias occurs near the top
of a great thickness of greywackes and conglomerates called the
Kaihiku Series, and is separated by several hundred feet from
the next fossiliferous beds above it. The Kaihiku fossils are
scanty in species, and no cephalopods occur. Among those re-
stricted to this horizon is Daonella indica Bittner, which occurs
in Ladino-Carnic deposits in the Himalayas and in the Malay
Archipelago. Members or survivors of a Muschelkalk fauna
occur in the form of Spiriferine of the group of Spiriferina
fragilis Schlothem. It is concluded that the Kaihiku fossil
360 Geological Society.
horizon is either late Middle or early Upper Trias, and the great
unfossiliferous series below it represents the Middle and possibly
Lower Trias.
The most highly fossiliferous division is the Carnic—the Oreti
and Wairoa Series of New Zealand. geologists. Several am-
“monites occur, among which Discophyllites cf. ebneri Mojsisovics
is found in the Carnie and Lower Noric of the Himalayas.
The Halobie include H zitteli Lindstrém, a Spitsbergen fossil,
together with . hochstetteri Mojsisovics and H. austriaca
Mojsisovies. Several of the Carnie fossils show affinities with
European Alpine forms, and can be used for purposes of
correlation.
The Noric horizon, the Otapiri Series in part, is represented by
felspathic sandstones containing immense quantities of Pseudo-
monotis, a genus which characterizes the Noric in all the Cireum-
Pacific Trias. Ps. righmondiana Zittel is known only from New
Zealand and New (MR aonia: but the Author found the Asiatic,
Siberian, and Japanese form, Ps. ochotica Teller, in allits varieties,
in very high Noric beds near Nelson.
The Rhetic, the upper part of the Otapiri Series of local
geologists, comprises a great thickness of sandy and pebbly beds.
Its fossils include an extremely-alate Spiriferina and a group of
specialized bisuleate Spirigerids. An Arcestid of Rhetie aspect
was collected high up in these beds at Kawhia.
Forty-seven genera and species of molluses and brachiopods are
recorded in the present paper, of which three genera and forty-one
species are regarded as new.
The brachiopods are of considerable interest, and exhibit phylo-
gerontic tendencies in several of the groups as they approach
extinction.
The affinities of the New Zealand Trias with that of the Malay
Archipelago, and especially of New Caledonia, is discussed ; and it
is shown that the faunal transgression which occurred over those
regions, at or shortly before the commencement of Upper Triassic
times, extended also to the area now occupied by New Zealand.
2. ‘The Triassic Crinoids from New Zealand. collected by Mr. C.
T. Trechmann.’ By Francis Arthur Bather, M.A., D.Sc., F.R.S.,
F.GS.
The specimens are all from the Kaihiku Series, and comprise :—
(1) an Lntrochus from near Nelson, with a broadly waved suture ;
(2) axrock-fragment from the Hokanui hills, containing imprints
of columnals and brachials representing two genera : namely, (@) an
Entrochus with ridges of the joint-face arranged in pairs separated
by shorter ridges; (6) an Isocrinus of the group of I. dubius
(Goldfuss). Comparison of the three new species based on all these
remains with the Triassic crinoids described from Europe and
especially with those from North America, leads to the conclusion
that they are of Upper Triassic age. They bear, however, no
resemblance to the Upper Triassic crinoids from Timor, which the
Author has in hand for description. ~ ,
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XXXIII.—Descriptions of New Pyralide of the Subfamilies
Hydrocampine, Scopariane, §c. By Sir Georce F.
Hampson, Bart., F.Z.S., &c.
Tur numbers attached to the species in the following paper refer
to my classification of Hydrocampine and Scoparianz in the Trans.
Ent. Soc. 1897, pp. 127-240, and to subsequent supplementary
papers in the Ann. & Mag. Nat. Hist.
E)PIPASCHIAN 2.
Genus PLUTOPASCHIA, nov.
Type, Stericta sinapis, Roths.
Proboscis fully developed; palpi upturned, the 2nd joint reaching E
to vertex of head, the 3rd rather long. Antenne of female mi-
nutely ciliated; tibiae smoothly scaled. Fore wing narrow, the
apex rounded, the termen evenly curved; vein 3 from before angle
of cell; 4, 5 stalked; 6 from upper angle; 7, 8, 9, 10 coincident ;
11 from cell. Hind wing with veins 3 and 5 shortly stalked, 4
absent; 6, 7 from upper angle of cell, 7 anastomosing with 8.
Genus GEROPASCHTIA, 0. n.
Arzopaschia, Hmpsn, A. M. N. H. (8) xviii. p. 1384 (1916), nec Hmpsn,
1906.
Type, G. grisealis.
Ann. & Mag. N. Hist. Ser, 8. Vol. xix, 24
oo
oO
Ww
Sir G. F. Hampson on new
CHRYSAUGINZ.
Murgisca mesozonalis, sp. n.
Q. Head, thorax, and abdomen whitish ; patagia and abdomen
above tinged with rufous. Fore wing pale greenish yellow; a
silvery-white fascia on base of inner area bounded above by a rufous
line; a medial silvery-white band from upper angle of cell to inner
margin bounded by rufous and constricted at lower angle of cell
and vein 1. Hind wing ochreous yellow; cilia of both wings
silvery white.
Hab. W. Avustratta, Sherlock R. (Clements), 2 Q type, Roe-
bourne. xp. 22 mm.
PYRALINE.
Pyralis costinotalis, Hmpsn. A. M. N. H. (8) xix. p. 67 (1917).
Hab. Formosa, Arizan (Wileman), 2 5 type. Hap. 24 mm.
HTYDROCAMPINE.
(La) Gargela niphostola, sp. n.
@. Head, thorax, and abdomen silvery white, the last dorsally
suffused with red-brown except on two basal segments ; palpi with
the 2nd joint golden brown behind ; fore femora and tibiz suffused
with golden brown, the tarsi ringed with brown towards extremity ;
ventral surface of abdomen tinged with golden yellow. Fore wing
silvery white; a faint curved brownish postmedial line; a blackish
point on termen below apex. Hind wing uniform silvery white.
Underside silvery white; the fore wing with a black point on
termen below apex.
Hab. D’Enrrecasteavx Is., Fergusson I. (Meek), 1 9 type.
Exp. 18 mm.
(6) Gargela obliquivitta, sp. n. :
Mid tibia of male dilated with a fold and tuft of long hair and
fringed with hair above.
Head, thorax, and abdomen silvery white; antenne brown except
above; palpi behind, except the 3rd. joint, and base of maxillary palpi
dark brown; fore legs tinged with ochreous, the femora at extre-
mities, tibiz near extremities and tarsi banded with black-brown.
Fore wing silvery white ; a golden-orange medial line, oblique to
upper angle of cell, then inwardly oblique and faint; a golden-
yellow tinge from end of cell to medial part of termen ; an oblique
golden-orange line from costa well beyond middle, where itis dilated
into a small wedge-shaped mark to discal fold near termen, an
oblique golden-orange wedge-shaped mark beyond it from costa to
discal fold just before termen ; a cupreous-brown termiual line from
apex to discal fold, and a double cupreous-brown striga before termen
Pyralides of the Subfamily Hydrocampine. 363
above vein 3; cilia white, metallic golden at tips, with cupreous-
brown line at base from vein 4 to submedian fold, and wholly tinged
with gold towards tornus. Hind wing uniform silvery white.
Hab. Mouvecas, Batchian (Doherty), 1 3,12 type. Hzp.
16 mm.
(56) Argyractis leucostola, sp. n.
Fore wing of male with small fovea below the costa beyond
middle and vein 11 distorted.
Head, thorax, and abdomen silvery white; fore tibiz black at
ase and extremity. Fore wing silvery white, the costa suffused
with blackish brown to end of cell; a reddish-brown spot in end
of cell and oblique black-brown line from median nervure near end
of cell to inner margin, diffused on inner side; a brown discoidal
striga and oblique line from it to inner margin; postmedial line
double, black-brown filled in with yellow and oblique to vein 3,
then single, retracted with an upwards curve to lower angle of cell, |
then strongly excurved to inner margin, its sinus filled in with
yellow; a terminal yellow band, arising just below apex, its inner
side defined by a black-brown line to above vein 2 and its outer by
black points, a silvery spot defined by brown before termen below
vein 2. Hind wing silvery white; a sinuous black-brown line
from middle of costa’to tornus, with a yellow patch before it below
the cell; a double slightly sinuous black-brown line from costa just
before apex to submedian fold, excurved below costa and filled in
with yellow below vein 3; a yellow terminal band from apex to
submedian fold, defined on inner side by a waved black line except
at apex ; cilia with a brown line at base to vein 2.
Hab. Br. C. Arrica, Mt. Mlanje (Neave), 25, 5 2 type.
Exp., 3 18, 9 22 mm.
(18a) Argyractis flavivittalis, sp. n.
do. Head, thorax, and abdomen whitish mixed with some red-
brown; antennz red-brown; fore tarsi with brown bands towards
extremities. Fore wing silvery white, the costal area suffused with
brown ; a brown antemedial line defined on outer side by white, ”
incurved from below costa; a medial brown line defined on outer
side by white, incurved below the cell; a small brown discoidal
spot; postmedial line silvery white, defined on inner side by brown
from costa to vein 4, then angled inwards to median nervure before
end of ceil, then oblique and defined on inner side by brown to
tornus, with a wedge-shaped yellow patch above it from lower angle
of cell to below its angle at vein 4; a silvery-white subterminal
line defined on each side by brown from costa to vein 2 and a small
brown spot with silvery-white mark before it below vein 2; cilia
brown at base, whitish at tips. Hind wing white, the end of cell and
the area beyond it from costa to vein 3 yellow ; a silvery discoidal bar
defined on inner side by a brown striga ; postmedial line silvery white
and oblique from -costa to vein 3; a terminal black patch #rom below
24:
364 Sir G. F. Hampson on new
apex to vein 2, with four ocelli on it with black centres defined ‘on
inner side by white and on outer by golden yellow; cilia white
with brown line near base from apex to vein 2.
Hab. Perv, San Domingo (Ockenden), 1 3 type. Hap. 14mm.
(20a) Argyractis cuprescens, sp. n.
2. Head, thorax, and abdomen cupreous red-brown mixed with
some white, the abdomen banded with white; frons white at sides.
Fore wing cupreous red-brown; a broad oblique silvery-white band
from median nervure before middle to inner margin near base; a
white medial line defined on each side by darker brown, excurved
below costa, then erect; an orange-yellow discoidal bar defined on
outer side bya brown line; postmedial line represented by a silvery-
white bar from costa connected with an oblique wedge-shaped patch
from beyond the cell to above vein 2 and an oblique band from
lower angle of cell to inner margin with an orange-yellow patch
between it and the wedge-shaped patch; a silvery-white band before
termen ending in a point at vein 2; an orange-yellow terminal band
defined at sides by brown lines, ending below submedian fold where
it is bent inwards; cilia brown at base, white at tips. Hind wing
silvery white ; some brown at base; a brown medial band, incurved
below the cell, with an orange-yellow discoidal pateh on it and some
orange-yellow below the cell; postmedial line cupreous brown,
oblique to beyond lower angle of cell, then incurved; an irregularly
reniform white subterminal patch defined by cupreous brown from
costa to vein 4; a black terminal band from apex to vein 2 with
five iridescent silvery annuli on it, defined on inner side by a white
line with a waved black-brown line before it and with some fulvous
yellow from it to tornus; cilia white, red-brown at base except
towards tornus.
Hab, Ecuapor, Rio Verde (Palmer), 1 9 type. Hxp. 18 mm.
(24a) Argyractis productalis, sp. n.
@. Head, thorax, and abdomen red-brown mixed with whitish,
the abdomen with whitish bands on basal and 8rd segments; an-
tenn red-brown ; frons whitish ; ventral surface of abdomen white.
Fore wing red-brown mixed with white, the subterminal triangular
patch from costa to vein 2 deep red-brown ; a rather diffused white
antemedial band from below costa to inner margin; a white medial
band defined on outer side by brown, sharply bent inwards to inner
margin ; a white discoidal bar defined by red-brown ; a white post-
medial band, its outer edge oblique to vein 3, then retracted to near
lower angle of cell, then oblique to inner margin near tornus with
its inner edge defined by brown; a silvery-white subterminal band
from costa to below vein 2 where it ends in a point and is some- -
what bent inwards, a fulvous-yellow band beyond it defined at sides
by brown lines; cilia white, brown at base except towards tornus.
Hind wing white, the terminal area irrorated with black to vein 3;
Pyralidee of the Subfamily Hydrocampine. 365
a faint rather diffused yellow-brown medial line, oblique to below
the cell, then incurved; postmedial line indistinct, yellow-brown,
oblique to below vein 4, then inwardly oblique and sinuous; a black
point just below apex and four black spots defined on outer side by
silver before termen at veins 6 to 2 with a slight waved black line
before them and some brownish beyond them before the blackish
terminal line.
Hab. Peru, Carabaya, Oconeque (Ockenden), 1 2 type. Exp.
32 mm.
(24¢) Argyractis argyrophora, sp. n.
2. Head and thorax silvery white, the vertex of head and
dorsum of thorax with some rufous; abdomen silvery white irro-
rated with brown, the anal tuft rufous; palpi rufous, the extremity
of 2nd joint and the 8rd joint white; pectus, legs, and ventral
surface of abdomen white, the fore legs suffused with rufous and
with black band at the extremity of the tibiew. Fore wing silvery
white, the costal area suffused with rufous; a rufous antemedial
line, interrupted in submedian interspace ; a silvery-white discoidal
bar defined at sides by rufous; postmedial line rufous, oblique to
vein 4, then almost obsolete and retracted to median nervure before
end of cell, then strong and incurved, a silvery-white band beyond
it extending to costa; a wedge-shaped rufous patch from costa to
vein 4 before a white terminal band suffused with metallic silver
defined by fine blackish lines and forming a wedge-shaped mark at
tornus, the area above this mark suffused with yellow; cilia white.
Hind wing white; a fulvous-yellow discoidal bar defined on outer
side by a metallic silvery bar; an oblique postmedial metallic
silvery band, interrupted at vein 4 and ending on termen above
tornus, defined on inner side by a yellow band and with the area
beyond it yellow ; five partly confluent black spots on. termen with
some metallic silvery between them from below apex to vein 2 and
with some silver on termen from them to tornus ; cilia white with
a fine brown line near base from apex to vein 2.
Hab. CotomeBts, Choko Prov., Condoto (Spurrell), 5 2 type,
R. San Juan, Juntas de R. Tamana,1 9. Hep. 20 mm.
(25a) Argyractis brunneosuffusa, sp. n.
@. Head and thorax red-brown mixed with some whitish;
abdomen fulvous brown with whitish segmental lines, the base with
some whitish ; palpi with the 8rd joint white; ventral surface of
abdomen red-brown with white segmental lines. Fore wing
whitish suffused with red-brown especially on costal area; a sub-
basal shade formed by black-brown scales; a curved whitish ante-
medial line defined by shades formed by dark brown scales in the
interspaces ; a slight white discoidal lunule defined by diffused
dark brown ; postmedial line white defined on inner side by rather
diffused dark brown, oblique to vein 6, incurved at discal fold, at
366 Sir G. F. Hampson on new
vein 3 retracted to just below angle of cell and excurved below
vein 2; a white subterminal band narrowing to points at apex and
vein 2, defined on inner side by diffused dark brown and on outer
by a yellow terminal band defined on each side by dark lines.
Hind wing white, the inner area tinged with red-brown ; an oblique
white band bey ond the cell from below costa to submedian fold,
defined on each side by red-brown shades; postmedial line white,
oblique to vein 2 where it is excurved to near termen, incurved at
submedian fold then bent outwards to termen above tornus, defined
on inner side by red-brown and with an oblique white bar before it
near tornus, the area beyond it suffused with brown from costa to
vein 2 witha whitish pateh on it; a black band just before termen
from below apex to vein 2, with four metallic silvery annuli on it ;
the apex yellow followed bya yellow line beyond the ooeflette band ;
cilia white with a brown line at base to vein 2.
Hab. Ecvuapor, R. Pastaza, Banos (Palmer), 1 9 ty pe. Exp.
32 mm.
(316) Argyractis mimicalis, sp. 0.
Head and thorax whitish mixed with pale red-brown ; abdomen
white tinged with pale red-brown ; pectus, legs, and ventral suiface
of abdomen white with a faint rufous tinge. Fore wing white
irrorated with blackish; an inwardly oblique antemedial series of
black striz ; a yellow patch in and above end of cell; medial line
black, erect to just beyond lower angle of cell, then oblique, sinuous
and with another faint line before it towards inner margin ; an
indistinct black postmedial line from vein 3 to inner margin,
angled outwards above vein 1; the apical area yellow, broadly, at
costa and narrowing to a point at submedian fold; a subterminal
white spot below costa before a blackish and silvery line from below
the costa to vein 3 towards which it is rather diffused; a small
black spot at apex, then a terminal series of black points to sub-
median fold where there is a black and silvery mark before it; cilia
white mixed with fuscous. Hind wing white thickly irrorated with
black except on basal and terminal areas ; a diffused oblique black
subbasal band from discal fold to inner margin; a waved white
medial line slightly defined on each side by black; subterminal
line white defined on each side by black, excurved below costa, then
sinuous ; terminal area yellow; a black bar with some silvery-blue
scales on it at apex, then three black ocelli each with two silvery-
blue points on them and a black spot with some silvery blue on it
below submedian fold ; cilia white with a black line near base to
discal fold, then black bars beyond the ocelli,
Hab. Sierra LEonE (Clements), 1 d; N. Nicerta, Zungeru
(Macfie), 1 2 type, Borgu, Yelwa L. (dkigeod), 1 2. Hezp.
14 mm,
Pyralids of the Subfamily Hydrocampine. 367
_ Ble) Argyractis nyasalis, sp. n
¢@. Head and thorax pale yellow mixed with black-brown and
some white; abdomen pale red-brown with white segmental lines
and a blackish patch before extremity ; antenne dark brown ringed
with white ; palpi pale rufous; pectus and legs white tinged with
rufous, the fore tibize with black band near extremity; ventral
_ surface of abdomen white. Fore wing white irrorated with black ;
a blackish patch at base of costa ; a diffused inwardly oblique black
subbasal line ; a golden-yellow antemedial patch defined by black
scales from cell to inner margin; a double inwardly oblique sinuous
black medial line; postmedial line double, black filled in with
white, oblique to vein 3, then retracted with an upwards curve to
lower angle of cell, then strongly excurved above inner margin,
the area beyond it yellow irrorated with black ; a white subterminal
band defined by black lines from costa to vein 4 with some black
below it and an oblique black mark above tornus; a terminal
yellow band to vein 2; cilia white mixed with black. Hind wing
white, the postmedial area to vein 2 and the terminal area irrorated
with black ; a small black spot on inner margin near base ; a broad
black antemedial band from below costa to inner margin with a
small yellow spot on it in lower angle of cell, defined on outer side
by a white line followed by a curved black line; a double black
subterminal line filled in with white, excurved below costa, then
sinuous, the area beyond it yellow ; four rather diffused black ocelli
with silvery-blue points on them on termen between vein 7 and the
submedian fold, with a waved black line before them ; cilia blackish
at base, white at tips.
Hab. Br. C. Arrica, Blantyre (Davey), 1 35 type. LHxp.
16 mm.
(45a) Argyractis melanograpta, sp. 2.
3. Silvery white ; head with some black-brown behind; abdo-
men with black-brown band on 2nd segment and slight bar near
extremity, the anal tuft brown at extremity; palpi with the 3rd
joint black ; fore femora and fore and mid_tibie blaek-brown in
front, the tarsi banded with black-brown. Fore wing with oblique
black-brown subbasal band from costa to median nervure; medial
line black-brown, double and oblique towards costa, then single,
inwardly oblique, rather diffused and bent inwards to inner margin ;
subterminal line black-brown, double towards costa and inner mar-
gin, bent outwards to costa where the inner line has a short streak
on its inner side at costa, excurved to near termen at middle and
with some black-brown suffusion beyond it at tornus; a black-brown
terminal line from below apex to vein 4; cilia tinged with yellow.
Hind wing with some black-brown in end of cell; an obliquely
curved black-brown medial line, arising below costa and diffused on
outer side towards inner margin; a rather interrupted sinuous
black-brown subterminal line, diffused below discal fold; cilia
368 . Sir G. F. Hampson on new
yellow at base and with some black-brown scales at middle at apex,
brown at base from discal fold to tornus. ;
Hab. Br. Guiana, Demerara (Rodway), 1 g type. Eup.
‘10 mm.
(46a) Argyractis pheopastalis, sp. n.
@. Head and tegule white ; thorax and abdomen white suffused
with brown; antenne brown, yellowish white towards base; palpi
white; pectus, legs, and ventral surface of abdomen white, the fore
and mid legs tinged with yellow. Fore wing red-brown; a slight
whitish medial line, angled outwards below costa and excurved
below the ,cell; an obliquely curved white postmedial line from
costa to vein 4 where it is met by an oblique white mark on its
inner side from vein 6; a silvery-white subterminal band from
below costa, where it is bent outwards to vein 4, an orange-yellow
band beyond it on termen extending to vein 2 and defined by
brown lines, some silvery white below it above tornus; cilia white.
Hind wing red-brown; an indistinct sinuous whitish medial line
defined on inner side by darker brown ; four minute ocelli just
before termen between veins 7 and 2 with black centres and metallic
silvery annuli; cilia white, yellow at base and with dark brown line
at middle fr om apex to vein 2.
Hab. CotomsBia, Choko, Prov. Condoto (Spurrell), 1 9 type.
Exp. 10 mm.
(le) Eristena tenebrifera, sp. n.
Antenne of male thickened with scales above towards base ;
hind femora with fringe of short hair behind towards base, the hind
tibize with large tuft of long hair on inner side; fore wing with
veins 8, 4 stalked.
3. Head, thorax, and abdomen white tinged with ochreous
brown’and mixed with some fuscous ; antenne fuscous ; hind legs
with the fringe of hair on femora whitish, the tuft of hair on tibiz
white at base, black at tips. Fore wing white tinged with ochreous
brown ; some black suffusion on basal area; medial area with some
black suffusion, its inner edge rather oblique, its outer incurved
below the cell and excurved above inner margin ; some blackish
suffusion on postmedial costal area and two rather diffused blackish
subterminal lines, somewhat excurved below costa, then oblique; a
slight dark terminal line with some ochreous brown before it.
Hind wing white, the terminal area suffused with ochreous brown.
Hab. Durcu N. Guryea, Mimika R. (Wollaston), 1 3 type.
Exp, 14 mm.
@l SF) Evistena tetralitha, sp. n.
@. Head, thorax, and abdomen grey suffused with brown and
some fuscous ; pectus and ventral surface of abdomen whitish ;
fore tibiz with black stripe. Fore wing whitish suffused with
Pyralidee of the Subfamily Hydrocampine. 369
brown and irrorated with black; an oblique diffused black ante-
medial shade ; the cell whiter towards extremity ; postmedial line
rather diffused, black, obliquely curved to vein 3, then bent upwards
to upper angle of cell, then inwards and oblique to inner margin,
the area above its sinus whitish except towards costa and the area
in its sinus at end of cell and the whole area beyond it rufous; an
obliquely curved rather maculate silvery subterminal line defined on
each side by fuscous ; a terminal series of black points; cilia white
mixed with brown. Hind wing whitish; the basal area tinged
with brown and suffused with black; a rather diffused curved
black postmedial line, incurved below vein 3; the terminal area
suffused with rufous; four small black ocelli before termen between
discal and submedian folds, the two or three upper ocelli with white
points in centre, and the upper one with a black point above it;
cilia whitish tinged with brown and with a brown line near base.
Hab. Durcen N. Guinea, Mimika R. (Wollaston), 5 Q type,
Wataikwa R. (Wollaston), 4 2, Snow Mts., Setakwa R. (Meek),
19; Br. N. Guryea, Kumusi R. (Meek), 3 9. Hap. 14-20 mm.
(16a) Nymphula manilensis, gp. n.
6. Head, thorax, and abdomen yellow mixed with some white
and fuscous, the last white at base and extremity and with black
segmental lines; antennz yellow ringed with black ; pectus, legs,
and ventral surface of abdomen white slightly irrorated with brown,
Fore wing yellow irrorated with dark brown and mixed with some
white, especially on outer half of medial area except towards costa ;
antemedial line white defined by some black scales, oblique below
the cell; a black point in middle of cell and some black suffusion
in end of cell; medial line white defined on inner side by black
scales, oblique to middle of discocellulars, then inwardly oblique, an.
oblique white mark beyond it below costa; postmedial line double,
brown filled in with white, arising below the costa where there is a
white mark before it, oblique and sinuous ; subterminal line formed
by somewhat dentate white marks defined on each side by blackish,
excurved at middle. Hind wing yellow irrorated with black, the
base white; an oblique black mark above inner margin near base ;
medial line double, black filled in with white, sinuous ; medial area
white with a yellow patch defined by black from below costa to.
vein 2; postmedial line double, black filled in with white, obliquely
curved, slightly sinuous towards costa; a rather maculate white
subterminal band defined on each side by black, sinuous; a fine
black terminal line; cilia white.
Hab. Purireprnes, Manila (Ledyard),1 3 type. Exp.14mm.
(19a) Nymphula leucoplagalis, sp. n.
Bocchoris zoilusalis, Druce, Biol. Centr.-Am., Het. ii. p. 558 (part.), nee
Wik.
3. Head, thorax, and abdomen fulvous yellow mixed with fuscous
370 Sir G. F. Hampson on new
and some white; antennz black; pectus, hind legs, and ventral
surface of abdomen towards base white; fore tibiz fuscous, white
at extremities. Fore wing fulvous yellow irrorated with dark red-
brown; a subbasal red-brown shade; a medial red-brown shade
with some white scales on it in end of cell, a white patch beyond it
on costal area, defined on outer side by a red-brown shade joining
the medial shade at lower angle of cell, then with three small white
spots on its outer edge, a small white spot beyond it on costa; the
terminal area suffused with dark red-brown. Hind wing fulvous
yellow, the basal half and terminal area suffused with dark red-
brown. ;
Hab. Mexico, Morelos, Cuernavaca (H. H. Smith), 1 3 type,
Godman-Salvin Coll. Hap. 14 mi.
(306) Nymphula plumbefusalis, sp. n.
2. Head and thorax leaden grey tinged with fuscous ; abdomen
white, basally suffused with reddish brown; antennze dark brown ;
frons white; palpi with the 8rd joint white; pectus, legs, and
ventral surface of abdomen white, the fore and mid legs tinged
with brown, the fore tibie blackish on inner side, white on outer
side except towards base. Fore wing leaden grey tinged with
brown ; an antemedial black spot below the cell and slight rufous
and dark shade towards inner margin; a black discoidal spot; an
indistinct diffused rufous postmedial shade, incurved below vein 4
to below end of cell, arising below the costa and interrupted below
the cell; a faint dark subterminal shade ; a faint punctiform brown
line before termen and terminal series of slight dark spots. Hind
wing pure white; a faint reddish-brown line from lower angle of
cell to vein 1; a reddish-brown postmedial bar at discal fold,
diffused spot at vein 2 and small spot at inner margin; a puncti-
form line before termen and a terminal line; cilia white mixed
with reddish brown and chequered with darker brown at base.
Hab. Supan, Blue Nile (flower), 6 ¢ type. Harp. 20 mm.
(526) Nymphula metastictalis, sp. n.
©. Head, thorax, and abdomen white, the metathorax with
brownish bar, the abdomen tinged with ochreous brown except at
base; fore tibize fuscous brown at extremities. Fore wing silvery
white slightly tinged with ochreous brown except on terminal area ;
small black-brown spots at base of costa and cell; a black-brown
antemedial spot on costa; an inwardly oblique brown media] line
with black spot at costa; a small white discoidal spot indistinctly
defined by brown; postmedial line brown with a black spot at costa
where it arises towards apex, excurved to vein 2, then incurved to
inner margin below end of cell; a curved brown subterminal line
from vein 7 to submedian fold ; a fine black terminal line before a
yellow terminal band, punctiform towards apex ; cilia white suffused
with brown. Hind wing white suffused with reddish brown except
Pyralide of the Subfamily Hy drocampine. 371
at base and on inner area; diffused brown ante- and postmedial
lines and a faint subterminal shade; two small black spots just
before termen below apex and eight between discal and submedian
folds, all defined on inner side by white ; the termen yellow; cilia
brown at base, white at tips.
Hab. Goodenough I, (Meek), 1 2 type. Exp. 20 mm.
(536) Nymphula flavicostalis, sp. n.
Fore wing of male without fovea below the cell.
3. Head and thorax white mixed with black-brown and some
yellowish ; abdomen white irrorated with black forming diffused
dorsal bands except at base ; antenne yellow ringed with brown ;
palpi yellow mixed with red-brown; pectus and ventral surface of
abdomen white irrorated with dark brown; legs white, the tibiz
and tarsi banded with black. Fore wing white tinged in parts
with brown and irrorated with black-brown, the costal area to end
of cell and towards apex orange-yellow; antemedial line white
defined on each side by blackish, curved, obsolete at inner margin ;
an elongate white spot in the cell before the medial line which is
white defined on each side by blackish, slightly waved, oblique
below the cell, a white patch beyond it on costal area: an irregular
yellowish discoidal spot defined by black scales ; postmedial line
white defined on each side by dark brown, ‘maculate and slightly
curved to vein 3, then bent inwards and obsolescent to lower angle
of cell, then erect and sinuous, some alternating black and white
marks before it on costa; a white subapical point, then a series of
white subterminal lunules, larger and extending to near the post~
medial line above and below ..vein 6, with diffused black marks
beyond them to vein 3; the termen narrowly yellow; cilia black
at base, then white with black marks towards apex and at middle.
Hind wing white; rather diffused brownish medial and post-
medial lines; the termen suffused with brownish, its inner edge
slightly waved ; some slight dark marks on termen towards apex ;
cilia with some dark scales at tips; yellowish at base and with a
blackish line at middle between veins 3 and 1,
Hab, Purv, Carabaya, Oconeque (Ockenden), 1 3 type. Exp,
22 mm.
(53) Nymphula graphicalis, sp. n.
2. Head, thorax, and abdomen white, the last with faint dorsal
ochreous-brown bands except at base. Fore wing silvery white; a
subbasal black point below vein 1; a slight antemedial spot formed
by black scales below costa; medial line with an oblique black
striga from costa, then double and pale ochreous brown, incurved
in the cell and obliquely excurved below it; a narrow discoidal
lunule defined by black and a patch of pale ochreous-brown suffu-
sion below end of cell; postmedial line double, brown with a black
point on the inner line at costa, excurved below costa, then oblique,
372 Sir G. F. Hampson on new
slightly waved and paler below vein 4, a pale ochreous-brown tinge
before its costal half and beyond it below vein 5, pale ochreous-
brown streaks beyond it on costa and vein 7 with a black streak
between them from termen; cilia tinged with ochreous brown.
Hind wing silvery white; a double waved black-brown antemedial
line; a discoidal bar defined by black-brown; postmedial line
double, ochreous brown and somewhat dentate, slight ochreous-
brown suffusion before it except towards costa and inner margin
and slight streaks beyond it on the veins of costal half, then brown
suffusion to tornus; cilia with pale ochreous brown mixed.
Hab. Perv, Carabaya, Oconeque (Ockenden), 1 Q type. Exp.
20 mm.
(16) Margarosticha gaudialis, sp. n.
3. Head and thorax greyish mixed with brown, the tegule and
patagia with some fulvous; abdomen fulvous with brown dorsal
bands; antenne whitish tinged with brown; palpi banded with
dark brown ; pectus, legs, and ventral surface of abdomen whitish
_ tinged with brown, the fore legs with the femora black above, the
tibiz black at extremities, the tarsi with black marks at the joints.
Fore wing fulvous orange, the base tinged with brown ; an oblique
silvery band defined on each side by brown from middle of cell to
inner margin; the fovea in end of cell surrounded by brown suffu-
sion; a triangular silvery-white mark beyond the cell defined by
rather diffused brown which extends to the costa; a silvery-white
subterminal band from costa to vein 4 where it ends in a point,
defined on inner side by rather diffused brown and on outer by a
black line extending to below vein 3; a terminal series of small
black spots to vein 2 and a short leaden-brown fascia above tornus
ending in a silvery point; cilia silvery: Hind wing fulvous orange,
the base pale ; a slightly sinuous silvery-white medial band defined
on each side by brown; the postmedial area brown, ending in a
point above tornus; four ocellate black spots on termen between
yein 7 and submedian fold, defined on inner side by a waved white
line and with fulvous orange between them, a small triangular
- black spot above the uppermost ocellus ; cilia silvery.
Hab. Avuiraury Is. (Meek), 3 5 type. Hxvp. 18 mm.
(8a) Margarosticha euprepialis, sp. n.
g. Head and thorax silvery white, the shoulders and some hairs
at tips of patagia yellow-brown ; abdomen silvery white at base
with a yellow-brown band, then pale yellow with slight yellow-brown
dorsal spots to beyond middle and silvery-white segmental lines ;
antennz pale fulvous; palpi white, the 2nd joint banded with pale
red-brown ; pectus, legs, and ventral surface of abdomen silvery
white, the fore tibie tinged with yellow above. Fore wing pale
yellow; a red-brown tinge at base of costa and a triangular white
patch on basal inner area defined on outer side by red-brown scales ;
Pyralide: of the Subfamily Hydrocampinee. 373
a broad oblique silvery-white medial band from cell to inner margin,
defined on inner side by a curved red-brown line and on outer by a
diffused red-brown patch from submedian fold to uiner margin ; an
oblique silvery-white discoidal bar beyond the fovea; an oblique
conical silvery-white postmedial patch from costa to vein 3, defined
by red-brown and with some red-brown beyond it on costa; a
silvery-white subterminal band from costa to discal fold, defined by
red-brown extending to vein 2 where it ends in a point and with a
wedge-shaped brown mark below it above tornus, these brown
markings tinged with silvery below the band; a fine red-brown
terminal line; cilia silvery. Hind wing white; a _ black-brown
subbasal band from below costa to above outer margin; a yellow
band from middle of costa to tornus defined on each side by slightly
sinuous red-brown lines, the outer line with a brilliant silver line
beyond it; the area beyond the band irrorated with black scales ;
five black ocellate spots on a white band on termen between discal
and submedian folds with small brilliant silvery spots between
them; the termen yellow towards tornus; cilia silvery.
Hab. QurENstand, Townsville (Dodd), 1 3 type. Lup.
22 mm.
(5) Margarosticha argyrograpta, sp. n.
@. Head, thorax, and abdomen orange-yellow with a golden
gloss, the head, patagia, and base of abdomen with some white ;
palpi white with brown band at extremity of the 2nd joint and the
8rd ‘joint yellow; pectus, legs, and ventral surface of abdomen
white, the fore and mid tibize tinged with yellow and the former
black at extremity. Fore wing orange-yellow; a silvery-white
subbasal patch from middle of cell to inner margin where it extends
to the base, its outer edge angled outwards at median nervure ; an
oblique silvery-white medial band from discal fold to inner margin,
produced to streaks beyond lower angle of cell and below cell 2; an
oblique triangular silvery-white postmedial patch from costa to.
vein 4, its outer edge defined by brown ; a wedge-shaped silvery-
white subterminal band from costa to below vein 3, slightly defined
at sides by brown and its lower part metallic silvery; a metallic
silvery fascia above tornus; a terminal series of black points with
larger point at apex; cilia white tinged with yellow. Hind wing
orange-yellow, the base white ; a silvery-white medial band with
slightly waved edges, its inner edge defined by brown; a metallic
silvery postmedial lunule between veins 5 and 2 and a spot above
tornus ; three large black ocellate spots on termen between discal
and submedian folds with some silvery seales on their inner edge
and a line before them which is yellowish above and silvery white
below, extending to termen below the lowest spot, a small triangular
black spot above the uppermost spot on termen ; cilia silvery white
at base, white tinged with brown at tips.
Hab. Bismarck ARCHIPELAGO, Rook I. (Meek), 3 2 type,
Exp. 18-22 mm,
374 Sir G. F. Uampson on new
(1c) Cataclysta perirrorata, sp. n.
Head, thorax, and abdomen silvery white mixed with brown;
antennz white tinged with brown; palpi white tinged with yellow;
pectus, legs, and ventral surface of abdomen white tinged with
brown. Fore wing white, the medial area irrorated with large
black scales ; the base orange-yellow with some brown at costa ; an
inwardly oblique orange-yellow antemedial band slightly defined at
sides by brown; an oblique orange-yellow postmedial band from
below costa to termen at submedian fold where it is confluent with
a similar curved subterminal band, a silvery point below its extre-
mity with a yellow striga before it to tornus; an orange-yellow
terminal band from just below apex to vein 3 where it ends in a
point, defined on inner side by a brown line and with some silver
between it and the subterminal band except towards costa; cilia
white mixed with brown. Hind wing white, the medial area
irrorated with large black scales, narrowing to inner margin near
tornus; a faint oblique orange-yellow antemedial shade; two fine
very slightly waved black subterminal lines, slightly excurved
below costa and bent outwards to tornus; four ocellate black spots
on termen from below apex to submedian fold with small metallic
silver spots on the 1st and 4th and small silver spots between them
on termen.; cilia white with a brown line at base and some brown
at tips especially towards apex. .
Ab. 1. Abdomen orange-yellow mixed with brown; fore wing
with the costal area suffused with brown, the medial area more
thickly irrorated with black except towards costa, the medial band
on outer side and the subterminal band on inner side with some
silvery suffusion, the oblique postmedial band almost interrupted at
middle and with distinct bar from it to tornus; hind wing suffused
with brown except at base, the oblique orange-yellow antemedial
band distinct, defined by brown and bent inwards to inner margin,
the ocellate terminal black spots more confluent. Br. C. Africa,
Mashonaland.
Ab. 2. Fore wing with the bands and hind wing with the ante-
medial shade rufous. Cape Colony.
Hab. Goup Coast, Appan, 1 d, Bibianaha (Spurrell), 4 3,
29, Kumasi (Sanders), 1 ¢ type; 8S. Niger1a, Ebute Meta
(Boag),1 3,1 9; Br. C. Arrica, Zomba (Old), 1 9, Mt. Mlanje
(Neave), 4 5,2 9; Masnonatann (Dobbie), 2 3g; Care Cotony,
Annshaw (Miss F. Barrett), 1 ¢. Exp. 12-16 mm.
(le) Cataclysta nigristriata, sp. n.
Head, thorax, and abdomen silvery white suffused with red-
brown, the last more fulvous towards extremity; palpi brown
towards base, white towards extremity; pectus, legs, and ventral
surface of abdomen white tinged with brown. Fore wing white,
the medial area except towards costa irrorated with large black
scales and with fine black streaks beyond the cell and on vein 1
» ah bee
~ Pyralide of the Subfamily Hydrocampine. 375
and inner margin, the terminal area from costa beyond middle to
inner margin near tornus golden yellow; the base reddish brown
with inwardly oblique outer edge; a faint inwardly oblique
yellowish antemedial band, followed by a cupreous-brown medial
line, incurved below the cell; an oblique wedge-shaped white post-
medial patch from costa to discal fold, defined on inner side by
brown; a subterminal band from below costa to vein 4 where it
ends ina point, white above, silvery below, defined by black lines,
the line on inner side reaching the costa; an oblique silvery bar
from below vein 4 beyond the cell to just above tornus; cilia
white tinged with cupreous brown except at submedian fold and at
tips between veins 6 and 4. Hind wing white, the medial area
irrorated with large black scales and with slight streaks below costa
and triangular in shape, from costa to submedian fold; cupreous-
brown bars at and beyond end of cell; the inner area orange-
yellow from before middle to tornus with metallic streaks on it at
middle of inner margin and in terminal part of submedian fold; an
oblique black line from below apex to submedian fold beyond
middle; five ocellate black spots on termen from below apex to
above tornus with a white band before them and orange-yellow
between their upper parts and metallic silver spots on termen, the
first and third black spots smaller; cilia cupreous brown, white at
tips except at apex.
Hab. Durcu N. Gury, Ron I. (Doherty), 1 3,1 9, Kapaur
(Doherty), 1 3, Mimika (Woolaston), 1 3 type; Apmiraury Is.
(Meak), 1 $, 1 Q; LovrstapE Is., St. Aignan (Meek), 1 2.
Hap. 12 mm.
(1g) Cataclysta amboinalis, sp. n.
2. Head, thorax, and abdomen golden yellow mixed with white
and black; antennz brownish, yellow towards base; palpi yellow
tinged with brown ; pectus, legs, and ventral surface of abdomen
white tinged with brown. Fore wing white, the basal area and
costal area to end of cell tinged with cupreous brown, the medial
area irrorated with large black scales except towards costa, the
apical area from costa at end of cell to termen above tornus golden
yellow defined on inner side by brown ; antemedial line dark brown,
slightly curved ; an oblique wedge-shaped silvery-white postmedial
patch defined by brown from costa to vein 4; a curved subterminal
band from costa to vein 3, silvery white above, metallic silvery
below defined by black lines, the outer line not reaching the costa ;
an oblique metallic silvery spot defined by blackish above tornus ;
cilia cupreous brown at base, white tinged with brown at tips, at
submedian fold pure white to base. Hind wing white, the medial
area irrorated with large black scales except towards inner margin,
the terminal area cupreous brown; a slight black spot near base
below the cell; an indistinctly double oblique cupreous-brown
antemedial line; four partly confluent ocellate black spots with
metallic silver centres on termen from below apex to submedian
376 On new Pyralide of the Subfamily Hydrocampine.
fold, with some orange-yellow scales before the three lower spots
and metallic gold points between the spots on termen; cilia
cupreous brown at base, white tinged with brown at tips.
Hab. AmBotna (Doherty), 1 2 type. ap. 12 mm.
(1h) Cataclysta queenslandica, sp. n.
3. Head and thorax silvery white, the patagia yellowish at tips;
abdomen silvery white suffused with golden yellow; antenne
brownish with white points in front; palpi yellow tinged with
brown ; pectus, legs, and ventral surface of abdomen white tinged
with yellow, the fore femora above and tibia on inner side black.
Fore wing white, the basal area obliquely and the costal area to
end of cell cupreotis brown, the medial area irrorated with cupreous
brown except towards costa, the terminal area orange-yellow from
costa at end of cell to inner margin near tornus; a cupreous-brown
discoidal spot and two lines from lower angle of cell, diverging
towards inner margin ; an oblique wedge-shaped silvery-white post-
medial patch from costa to vein 4, some cupreous brown beyond it
on costa ; a subterminal band from below costa to vein 3, where it
ends in a point, white above and metallic silver below, the white
part defined at sides by cupreous brown on inner side extending
to the costa ; an oblique metallic silver spot above tornus ; a fine
dark terminal line; cilia cupreous brown, white at submedian fold.
Hind wing white, the postmedial area irrorated with large black
seales, triahgularly from costa where it extends to apex to sub-
median fold; the base tinged with yellow; a cupreous-brown ante-
medial bar in and below the cell; an oblique cupreous-brown
medial band from costa to submedian fold; the terminal half of
inner area orange-yellow with some metallic silver at middle
of inner margin; a fine sinuous black-brown subterminal line from
apex to above tornus ; five partly confluent ocellate black spots on
termen with metallic silver points between them and orange-yellow
points on termen ; cilia cupreous brown at base, white tinged with
brown at tips.
2. Head with dark bar behind antenne; thorax tinged with
ochreous brown ; abdomen yellower; fore wing with the base and
costal area redder brown, an oblique red-brown medial band, the
yellow terminal area defined on inner side by two red-brown lines,
angled inwards to lower angle of cell; hind wing with oblique
orange-yellow medial band defined at sides by cupreous-brown
lines, the postmedial area more strongly irrorated, the subterminal
line more distinct and defined on each side by yellowish white, the
terminal ocellate spots better defined and separate.
Hab. QuEENstand, Cooktown, Cedar Bay (leek), 1 3 type,
Kuranda (Dodd),1 9. Hxp., 5 14, 2 16 mm,
[To be continued, }
On the Genera Sardinella, Harengula, cc. 377
®
XXXIV.—A Revision of the Clipetd Fishes of the Genera
Sardinella, Harengula, de. By OC. Tare Recan, M.A.
(Published by permission of the Trustees of the British Museum.)
THE genus Clupea, as understood by Giinther, includes a
number of genera which may be arranged in two grotps:
one of these, characterized by a well-marked median notch in
the upper jaw, has been dealt with in a recent paper (supra,
p- 297) ; the other group includes genera without a distinct
notch in the upper jaw, namely, Clupea, Clupeonella, Sardina,
Sardinella, Opisthonema, Harengula, Lile, and Heringia.
Of these I have already published revisions of Clupea and
Sardina (‘ Annals,’ (8) xviii. 1916, p. 1, and xix. 1917, p. 226),
and I have nothing to add to Berg’s synopsis of Clupeonella
(Harengu/a, Berg, ‘ Annals,’ (8) xi. 1913, p. 480, and Poiss.
de l’eau douce de la Russie, p. 30, 1916). The other genera,
comprising species that are mostly tropical and_ strictly
marine, form the subject of the present paper.
SARDINELLA, Val. 1847,
Cuv. & Val. Hist. Nat. Poiss. xx. p. 2638,
Clupeonia, Cuy. & Val. ¢. c. p. 346,
This genus is closely related to Sardina, Antipa (Regan,
Ann. & Mag. Nat. Hist. (8) xviii. 1916, p. 11), from which
it differs especially in the absence of radiating grooves on the
operculum and in the structure of the posterior margin of the
_ branchial chamber, the vertical edge of the cleithrum being
covered by a dermal fold which bears two obtusely pointed
projections some distance apart, with a shallow concavity
between them. The vertebrae are fewer than in Sardina,
numbering 44 in §S. gibbosa, 45 in S. sindensis, and 46 in
S. longiceps and 8, maderensis.
The majority of the species are tropical, but those of the
Eastern Atlantic range into the Mediterranean; most of
them are of considerable economic value, both as food-fishes
and as a source of oil,
Synopsis of the Species.
I, Ventral scutes sharply keeled.
A. Pelvic fins 9-rayed ; a dark spot at edge of operculum.
Eye 33 to 43 in length of head, which is 33 to 4 in
length of fish; 110-160 gill-rakers on lower
part GEA@RtCRIOR BPCHy i. ii Soe ee ae 1. aurita,
Ann, & Mag. N. Hist. Ser. 8. Vol. xix. 25
378 Mr. C. T. Regan on the Clupeid Fishes
Eye 5 to G6 in length of head, which is 3 to 33 in
length of fish; 180-250 gill-rakers on lower part
Orenterior arch..i.; wfisstpeeees ui in 2. longiceps.
B. Pelvic fins 8-rayed ; a dark spot at base of anterior rays of dorsal
fin.
1. Mediterranean and West African species.
Depth 33 to 4 in length; 60 to 95 gill-rakers on
lower part of anterior arch (in specimens of 100
to. 300 (nimeyg, acces see ince ee weeeee o Maderensis.
Depth 33 to 33 in length; 110 to 130 gill-rakers on
lower part of anterior arch (in specimens of 100
to}200 ann) Fo 5Fk sh Ce.G Ph gde BGs beta ate 4. eba.
Depth 3 in length; 90 to 100 gill-rakers on lower
part of anterior arch (in specimens of 170 to
DOO RIE) Stes arate ge elec ieee OER Ere Bist obey 5. cameronensis.
2. Indo-Pacific species.
Depth 23 in length; 130 gill-rakers on lower part of
anterior arch...... Acre withe walenee eet. ayant eee te 6, day.
Depth 23 to 3; gill-rakers 55 to 65; diameter of eye
ato os if tenath ot held): § i.e. ae ible eet ee 7. brachysoma.
Depth 3; gill-rakers 48 to 55; diameter of eye 33 to
32.in lenoth of head... .§ 5.5 .—eeie Sees scot Oe perforata,
Depth Sto 34: pill-rakers 70 Oye Fos aes aes oui wos 9. fimbriata,
Depth 3} to 4; gill-rakers 58 to 62 ...........4.. 10. sindensis. -
Depth 33 to 4; gill-rakers 50 to 55 ......... wee. LL. gebbosa.
Depth 33 to 4; gill-rakers 88 to 44. .............. 12. melanura.
If. Ventral scutes feebly keeled.
\
Maxillary nearly or quite reaching to below eye; 36
to 40 gill-rakers on lower part of anterior arch ;
pelvics below anterior half of dorsal .......... 13. sim.
Maxillary not reaching eye; 27 to 31 gill-rakers on
Ss fal « - ? . : . .
lower part of anterior arch; pelvics below origin
Ol dorsdleecg : cava e oe eee area oy nee 14, clupeoides.
1. Sardinella aurita.'
2 Clupea ceruleo-vittata, Richards, Iehth. China, p. 305 (1846) *.
Sur finella aurita, Cuv. & Val. Hist. Nat. Poiss, xx. p, 263, pl. 594
(1847)
Surdinella anchovia, Cuv. & Val. t..c, p. 269.
Meletta mediterranea, Cuv. & Val, ¢. ¢. p. 369.
a Sardinella lemuru, Bleek. Nat. Tijdschr. Ned. Ind. v. 1853, p. 500.
? Sardinia pseudohispanica, Poey, Mem. Cuba, ii. p. 811 (1860).
Clupea aurita, Giinth. Cat. Fish. vil. p. 420 (1868).
Clupea anchovia, Giinth, t. ¢, p. 421.
Clupea melanosticia, Giinth. t. e. p. 430.
Clipea lemuru, Giinth. t. c. p. 430; Bleek. Atl. Ichth. vi. p. 108, Clup.
pl. ix. fig. 1 (1872). ;
? Clipea psendohispanica, Giinth. t. ¢. p. 442.
Clupea brasiliensis, Steind. Sitzungsh. Akad. Wien, lxxx. 1880, p. 182.
* Richardson's description is based on a coloured figure by Reeves,
probably, but not certainly, intended to represent this species.
iets
of the Genera Sardinella, Harengula, Le. 319
Clupanodon pseudohispanicus, Jord, & Everm. Bull. U.S. Nat. Mus.
xlvii. 1896, p, 423.
Sardinella anchovia, Jord. & Everm. t. c. p. 429.
Clupea immaculata, Kishinouye, Journ. Imp. Fisheries Bureau Tokyo,
xiv. 1907, p. 96, pl. xix. fig. 1.
AN sige ee Antipa, Denkschr. Akad. Wien, lxxviii. 1906, p. 46
+ 1. oo. LZ.
Charen longiceps, Weber & Beaufort, Fish. Indo-Austral. Archipelago,
ii. p. 82 (1918),
Depth of body 4 to 54 in the length, length of head 34
to4. Snout as jong as or longer than diameter of eye, which
is 34 to 44 in the length of head; maxillary extending to
below anterior 2 of eye; a patch of teeth on the tongue.
110 (young) to 160 gill-rakers on lower part of anterior
arch. About 48 scales in a longitudinal series, 12 to 14 ina
transverseseries; ventral scutessharply keeled, 18-20 + 13-15.
Dorsal 16-20. Anal15-19. Pelvics 9-rayed, inserted below
or behind middle of dorsal. A dark spot at edge of operculum.
Vertebrae 47 or 48. ;
Cape Cod to Rio Janeiro; Black Sea and Mediterranean ;
Indo-Australian Archipelago, China, and Southern Japan.
The description is based on eight specimens, 120 to 180 mm.
long, from Havana, Trinidad, and Rio de Janeiro (S. an-
chovia), several of 110 to 130 mm. from Algiers, the type of
S. lemuru from Java (145 mm. long) and two examples from
China, 180 mm. long (C. melanosticta, Giinth.). In a very
small fish (75 mm.) I count 80 gill-rakers on the lower part
of the anterior areh.
The discontinuous distribution of this species is remark-
able; in the Indian Ocean it is represented by the allied
S. longiceps.
2, Sardinella longiceps.
Sardinella longiceps, Cuv. & Val. Hist. Nat. Poiss, xx. p- 273 (1847).
Sardinella neohowii, Cuy. & Val. t. c. p. 274.
Alosa scombrina, Cuv. & Val. t.c. p. 442.
Cee Giinth. Cat. Fish. vii. p. 428 (1868); Day, Fish. India,
. Od.
Oluped scombrina, Giinth, ¢. ¢. p. 448.
Depth of body 4 to 42 in the length, length of head 3 to 33.
Snout longer than diameter of eye, which is 5 to 6 in the
length of head; maxillary extending to below anterior part
or nearly to middle of eye. 180 to 250 gill-rakers on lower
part of anterior arch. 46 to 48 scales in a longitudinal
series, 12 or 13 in a, transverse series ; ventral scutes sharply
keeled, 18-21 + 13-15... Dorsal 16-18. Anal14-16. Pelvies
9-rayed, below or beliind middle of dorsal. A dark spot at
edge of operculum. Vertebiz 47.
; 95%
880 Mr. C. T. Regan on the Clupeid Fishes
Tndian Ocean.
Several specimens, 120 to 180 mm. in total length, from
Mombasa, Muscat, and India.
3. Sardinella maderensis.
Clupea maderensis, Lowe, Trans. Zool. Soc. ii. 1839, p. 189. :
? Sardinella ygranigera, Cuv. & Valg Hist. Nat. Poiss. xx. p. 267 (1847).
Clupea maderensis (part.), Giinth. Cat. Fish. vii. p. 440 (1868).
Depth of body 32 to 4 in the length, length of head 33 to
41, Snout as long as or a little longer than diameter of eye,
which is 34 to 4 in length of head; maxillary extending to
below anterior + of eye. 60 to 95 gill-rakers on lower part
of anterior arch. 48 to 50 scales in a longitudinal, 12 or 13
in a transverse series; ventral scutes 19-20+14-16. Dorsal
18-19. Anal 18-19. Pelvies 8-rayed, a little in advance
of middle of dorsal. A dark spot at base of anterior dorsal
rays. Vertebre 48.
Eleven specimens, 110 to 300 mm. in total length, from
Madeira, Cape Verde Is., and Mogadore.
If S. granigera be this species, it occurs in the Medi-
terranean.
4. Sardinella eba.
Alosa eba, Cuv. & Val. Hist. Nat. Poiss. xx. p. 369 (1847).
Clupea maderensis (part.), Giinth. Cat. Fish. vii. p. 440 (1868).
Depth of body 34 to 32 in the length, length of head 33 to
4, Snout as long as diameter of eye, which is 33 to 4 in the
length of head ; maxillary extending to below anterior 4 of
eye or a little beyond. 110 to 130 gill-rakers on lower part
of anterior arch. 44 to 46 scales in a longitudinal, 11 to 13
in a transverse series; ventral scutes 18-19+4+14. Dorsal
18-20. Anal 17-22. Pelvics 8-rayed, below middle of
dorsal. A dark spot at base of anterior dorsal rays. Ver-
tebree 46.
Mediterranean ; West Africa.
Eight specimens, 110 to 200 mm. in total length, from
Keypt, Algiers, and Nigeria.
5. Sardinella cameronensis, sp. 0.
Clupea senegalensis (non Benn.), Giinth. Cat. Fish. vii. p. 441 (1868) *.
Depth of body 3 in the length, length of head 4. Snout
* Alosa senegalensis, Bennett (Proc. Zool, Soc. i. 1831, p. 147), is
probably a synonym of Sardina pilchardus.
of the Genera Sardinella, Harengula, Se. 381
as long as diameter of eye, which is 33 in length of head;
maxillary extending to below anterior 4 of eye. 90 to 100
gill-rakers on lower part of anterior arch. 44 scales in a
longitudinal, 13 in a transverse series ; ventral scutes 18-
19+14. Dorsal 18-19. Anal 20-21. Pelvics 8-rayed,
below middle of dorsal. Caudal lobes long, 3 length of fish.
A dark spot at base of anterior dorsal rays. Vertebre 46.
Two specimens, 170 and 200 mm. in total length, from
Camaroon.
6. Sardinella dayi, sp. n.
Depth of body 2? in the length, length of head 3¢.
Snout as long as diameter of eye, which is 32 in length of
head; maxillary extending to below anterior 4 of eye. 130
gill-rakers on lower part of anterior arch. 44 scales in a
longitudinal and 12 in a transverse series; ventral scutes
19+13. Dorsal 18, nearly equidistant from end of snout
and base of caudal. Anal19. Pelvics 8-rayed, below middle
of dorsal. Pectoral 2? length of head. Silvery; back
darker; a blackish spot at base of anterior dorsal rays ;
upper part of dorsal and edge of caudal dusky.
A single specimen, 125 mm. long, from Karwar, India,
presented by N. B. Kinnear, Esq. ‘The species is named in
memory of Dr. Francis Day.
7. Sardinella brachysoma.
? Kowala albella, Cuy. & Val. Hist. Nat. Poiss. xx. p. 362, pl. 602
(1847).
Sardinella brachysoma, Bleek. Verh. Bat. Gen. xxiv. 1852, Haringacht.
p. 19.
Hurengula hypselosoma, Bleek.. Nat. Tijdschr. Ned. Ind. viii. 1855,
427
Clupea brachysoma, Giinth, Cat. Fish. vii. p. 423 (1868) ; Bleek. Atl.
Ichth. vi. p. 104, Clup. pl. ix. fig. 4 (1872); Day, Fish. India, p. 635,
pl. clxii. fig. 3 (1878); Weber & Beaufort, Fish. Indo-Austral.
Archipel. ii. p. 70, fig. 25 (1913).
Clupea hypselosoma, Giinth. t. c. p. 481; Bleek. 7. ¢, pl. ix. fig. 2.
Depth of body 24 to 3 in the length, length of head 4 to
43. Snout shorter than diameter of eye, which is 3 to 33 in
Jength of head ; maxillary extending to below anterior } of
eye or a little beyond. 55 to 65 gill-rakers on lower part of
anterior arch. 40 to 44 scales in a longitudinal and 12 or 13
in » transverse series ; ventral scutes 17-20+12-13. Dorsal
14-20. Anal 18-22. Pelvics 8-rayed, below or in advance
of middle of dorsal. A dark spot at base of dorsal ; upper
pert of dorsal and ends of caudal lobes often dusky.
! India ; Malay Archipelago.
}
382 Mr. C. T. Regan on the Clupeid Fishes
Seven examples, 100 to 150 mm. in length, from Madras,
Java, and Amboina, including the types of the species and of
HI. hypselosoma.
8. Sardinella perforata.
Clupeonia perforata, Cantor, J. As. Soc, Bengal, xviii. 1850, p. 1276.
Clupalosa bulan, Bleek. Verh. Bat. Gen. xxii. 1849, Madura, p. 12.
Spratella kowala, Bleek. Nat. Tijdschr. Ned. Ind. ii. 1851, p. 492.
Clupea perforata, Giiuth. Cat. Fish. vii. p. 424 (1868) ; Bleek. Atl.
Ichth. vi. p. 110, Clup. pl. x. fig. 2 (1872); Weber & Beaufort,
Indo-Austral. Arch. Fish. ii. p. 74 (1918).
Clupea bulan, Bleek. Atl. Ichth. vi. p. 110, Clup. pl. viii. fig. 5 (1872).
Depth of body about 3 in the length, length of head 4
to 42. Snout as long as or a little shorter than diameter of
eye, piel ee to 32 in length of head; maxillary extending
to below anterior 4 of eye. 48 to 55 (58) gill-rakers on
lower part of anterior arch. About 44 scales ina longitudinal,
12 or 13 inatransverse series ; ventral scutes 17-20 + 10-13.
Dorsal 17-20. Anal 17-20. Pelvics 8-rayed, below or in
advance of middle of dorsal. A dark spot at base of anterior
dorsal rays.
Indian Ocean and Archipelago.
Several specimens, 90 to 130 mm. in total length, including
types of the species, of S. kowala, and C. bulan, from the
Persian Gulf and the Malay Archipelago.
9. Sardinella fimbriata.
Spratella fimbriata, Cuv. & Val. Hist. Nat. Poiss. xx. p. 359, pl. 600
(1847).
Korvala lauta, Cantor, J. As. Soc. Bengal, xviii. 1850, p. 1279.
Clupea fimbriata, Giinth. Cat. Fish. vii. p. 427 (1868).
Depth of body 3 to 34 in the length, length of head 4.
Snout as long as diameter of eye, which is 33 to 32 in length
of head; maxillary extending to below anterior } of eye or a
little beyond. 70 to 75 gill-rakers on lower part of anterior
arch, About 45 scales in a longitudinal and 12 in a trans-
verse series ; ventral scutes 18-19+12-13. Dorsal 18-19.
Anal 18-21. Pelvies 8-rayed, in advance of middle of
dorsal. A dark spot at base of anterior dorsal rays; upper
part of dorsal and posterior edge of caudal dusky.
Sea of Bengal.
Four specimens, 110 to 125 mm. long, from Akyab, Oris.a,
Malabar, and Madras (May), and the types of K. laua
(skins) from Pinang.
er
of the Genera Sardinella, Harengula, dc. 383
10. Sard nellu s ndensis.
Clupea sindensis, Day, Fish. India, p. 638, pl. elxifi, fig. 2 (1878).
Depth of body 34 to 4 in the length, length of head 32 to
4}. Snout as lone as or shorter than diameter of eye, which
is "31 to 33 in the. length of head ; maxillary extending to
below Saiexids sof eye. 58 to 66 etll-rakers on lower part
of anterior arch. 44 to 48 scales in a longitudinal, 11 to 13
ina transverse series 3 ventral scutes 17-19+ 12-15. Dorsal
17-19. Anal 18-21. Pelvics 8-rayed, below or in advance
of middle of dorsal. A dark spot at base of anterior dorsal
rays ; upper part of dorsal and ends of caudal lubes sometimes
blackish.
Indian Ocean and Archipelago.
Thirteen specimens, 95 to 130 mm. in total length, from
Sind, Bombay, Amboina, and Formosa.
11. Sardinella gibbosa.
? Clupanodon jussiewt, Lacep. Hist. Nat. Poiss. v. pp. 469, 474, pl. xi.
fig. 2 (1803).
? Clupeonia jussteut, Cuv. & Val. Hist. Nat. Poiss. p. 346 (1847);
Sauvage, Hist. Madagascar Poiss. p. 495,
? Clipeonia fasciata, Cuy. & Val. t.c. p. 349.
Clupea gibbosa, Bleek. Journ. Ind. Arch, iii. 1849, p. 72; and Atl.
Ichth. vi. p. 106, Clup. pl. viii. fig. 6 (1872).
Spratella tenbang, Bleek. Verh. Bat. Gen, xxiv. 1852, Haringacht.
28.
Chines tembang (part.), Giinth. Cat. Fish. vii. p. 426 (1868).
Clupea fimbriata (part.), Day, Fish. India, p. 637 Weber & Beaufert,
Fish. Indo-Austral. Arch. 1. p. 75, fig. 26 (1913).
Depth of body 35 to 4 in the length, length of head + to
4}. Snout as long as or longer than diameter. of eye, which
is 35 to a in length of head; maxillary extending to below
anterior + or 4 of eye.|. 90 to 55 gill-rakers on lower part of
antellor pie 44 to 48 scales in a longitudinal, 11 to 13 in
a transverse series; ventral scutes 18-20+-13-15. Dorsal
17-20. Anal17-19. Pelvics 8-rayed, somewhat in advance |
of middle of dorsal, A dark spot at base of anterior dorsal
lays ; upper part of dorsal and posterior edge of caudal often
usky.
Indian Ocean and Archipelago.
Kleven specimens, 100 to 160 mm. in total length, from
Durban, Mombasa, Ganjam, Madras, Siam, Celebes, Java,
and Amboyna, including the type of & tembang.
384 Mr. C. T. Regan on the Clupeid Fishes
12. Sardinella melanura.
Clupanodon sinensis, var., Lacep. Hist. Nat. Poiss. v. pl. xi. fig. 3
(1803). ee
Clupea melanura, Cuv. Régne Anim. ed. 2, ii. p. 318 (1829).
—= Clupeonia commersont, Cuy. & Val. Hist. Nat. bois, xx. p. 350 (1847) ;
Sauvage, Hist. Madagascar Poiss. p. 494 (1891).
Spr atella Jjimbriata, Bleek. Verh. Batay. Genootsch. xxiv. 1852,
Haringacht. p. 27.
Huarengula melanurus, Bleek. Nat. Tijdschr. Ned. Ind. v. 1853, p. 245.
Clupea atricauda, Giinth. Cat. Fish. vii. p. 426 (1868); Bleek. Atl.
Ichth. vi. p. 106, Clup. pl. x. fig. 5 (1878) ; Day, Fish. India, p. 636,
pl. clxiv. fig. 5 ue Weber & Beaufort, Fish. Indo-Austral.
Arch. ii. p. 80 (1918).
Clupea aul Tisch Bleek. Atl. Ichth. vi. p. 105, Clup. pl. xiii. fig. 5.
(1872).
? Harengula melanura, Sauvage, Hist. Madagascar Poiss. p. 492,
pl. xlviil. fig. 4.
Depth of body 3} to 4 in the length, length of head 4 to
Al, Snout as nh as orm little longer than diameter of eye,
winch’ is 34 to 4 in the length of head ; maxillary extending
to below anterior 4 of eye. 388 to 44 gill-rakers on lower
part of anterior arch. 44 to 46 scales in a longitudinal
series, 12 or 13 in a transverse series ; ventral scutes sharply
keeled, 19-20+138. Dorsal 18-19. Anal 16-18. Pelvics
8-rayed, below or a little in advance of middle of dorsal. A
dark spot at base of anterior dorsal rays; ends of caudal
lobes usually blackish.
Indian Ocean and Archipelago.
Four specimens, 115 to 160 mm. in total length, including
two received from Dr. Bleeker as ee melanurus and 8. fimbri-
ata, which appear to be the specimens figured as C. atricauda
and C. sundaica.
13. Sardinella strm.
Clupea sirm, Ripp. Neue Wirbelth. Fische, p. 77, pl. xxi. fig. 1
(1835-40); Ginth. Cat. Fish. vii. p. 425 (1868) ; Weber & Beautort,
Fish. Indo-Austral. Archipel. ii. p. 62 (1918).
Sardinella letogaster, Cuv. & Val. Hist, Nat. Poiss. xx. p. 270 (1847).
Sardinella leiogastroides, Bleek. Nat. Tijdschr. Ned, Ind. vu. 1864,
p. 250.
fluid. liogaster, Bleek. Atl. Ichth. vi. p. 102, Clup. pl. iv. fig. 6 (1872) ;
Klunzinger, Zool. Botan. Ges. Wien, xxi. 187], p. 598; Weber &
Beaufort, é. ¢. pabl
Clupea leiogastr otdes, Bleek. 7. c. Clup. pl. xiv. fig. 2.
Clupea pinguis, Ginth. Ann. & Mag. Nat. Hist, x. 1872 2, p. 425, and
Brenchley’s Cruise of the “Curacoa,’ p. 426 (1878); Weber &
Beaufort, ¢. c. p. 83.
Depth of body 43 to 5 in the length, length of head 4 i
41. Snout longer than diameter of eye, which is 3% to 4}
the length of “head ; maxillary nearly or quite reaulie
of the Genera Sardinella, Harengula, ée. 385
vertical from anterior margin of eye. 36 to 40 gill-rakers on
lower part of anterior arch. 42 to 45 scales in a longitudinal,
12 in a transverse series; ventral scutes 16-15+13-15.
Dorsal 17-19. Anal 17-20. Pelvics 8-rayed, in advance of
middle of dorsal.
Indian Ocean and Archipelago.
Six specimens, 105 to 185 mm. in total length, from
Zanzibar, Batavia, Celebes, and Misol, including types of
S. liogastroides and C. pinguis.
14. Sardinella clupeoides.
Amblygaster clupecides, Bleek. Journ. Ind. Arch, 1849, p. 73.
Clupea clupeoides, Giinth. Cat. Fish, vii. p. 425 (1868); Bleek. Atl.
Ichth. vi. p. 103, Clup. pl. xiv. fig. 1 (1872) ; Weber & Beaufort,
Fish. Indo-Austral. Arcbipel. il. p. 63 (1915).
Clupea okinawensis, Kishinouye, Journ. Imp. Fisheries Bureau, Tokyo,
xiv. 1907, p. 96, pl. xix. fig. 2.
Depth of body 32 to 44 in the length, length of head 4 to
4}, Snout as long as or a little longer than diameter of eye,
which is 35 to 4 in length of head ; maxillary not extending
to beloweye. 27 to 31 gill-rakers on lower part of anterior
arch. 42 to 44 scales in a longitudinal, 12 in a transverse
series; ventral scutes 16-17+12-14. Dorsal 17-19. Anal
16-18. Pelvics 8-1ayed, nearly below origin of dorsal.
Malay Archipelago to Riu-Kiu Islands,
‘wo specimens, 160 and 230 mm. long, the latter the type
of the species,
OPISTHONEMA, Gill, 1861.
Proc. Acad, Philadelphia, p. 37.
Differs from Sardinella in that the last ray of the dorsal
fin is prolonged into a filament.
Two species from Tropical America.
1. Opisthonema oglinum.
Clupea thrissa (non Linn.), Broussonet, Ichth, fase. 1 (1872); Giinth.
Cat. Fish, vii. p. 482 (1868).
Megalops oglina, Lesueur, J. Ac. Philad. i, 1817, p. 359.
Opisthonema oglinum, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii.
1896, p, 482,
Depth of body 24 to 382 in the length, length of head 4
to 42, Snout as long as or a little longer than diameter of
eye, which is 34 to 4 in length of head; maxillary extending
to below anterior 4 of eye. 65 (young) to 100 gill-rakers on
Jower part of anterior arch, 48 to 50 scales in a longitudinal,
386 Mr. C. T. Regan on the Clupeid Fishes
15 or 16 in a transverse series 3 ventral scutes 17-20 + 13-16.
Dorsal 18-20. Anal 22-25. - Pelvics 8-rayed, a little in
advance of middle of dorsal.
Carolina to Brazil.
Several examples, 85 to 275 mm. in total length.
2. Opisthonema libertatis.
Meletta libertatis, Giinth. Proc. Zool. Soc. 1866, p. 603.
Clupea libertatis, Giinth. Cat. Fish. vii. p. 483 (1868).
Opisthonema libertate, Jord. & Everm. Bull. U.S. Nat, Mus. xlvii.
1896, p. 433.
Clupea (Opisthonema) bulleri, Regan, Ann, & Mag. Nat. Hist. (7) xiii.
1904, p. 255.
Depth of body 3 to 34 in the length, length of head 33 to
41, Snout as long as or longer than diameter of eye, which
is 32 to 5 in the length of head; maxillary extending to
below anterior 4 or nearly to middle of eye. 85 (young) to
165 gill-rakers on lower part of anterior arch. 48 to 50
scales in a longitudinal, 14 to 16 in a transverse series ;
ventral scutes 17-19+14-16. Dorsal 17-19. Anal 19-21.
Pelvics 8-rayed, a little in advance of middle of dorsal.
Pacific coast of Mexico and Central America; Galapagos
Islands.
Several examples, 75 to 250 mm. in total length.
HARENGULA, Val. 1847.
Cuy. & Val. Hist. Nat. Poiss. xx. p. 277.
This genus has never been properly distinguished from
Sardinella, but they differ in some important characters.
Tn Sardinella, as in Sardina, the two last rays of the anal
fin are enlarged and the transverse grooves on the scales are
paired, their inner ends separated by an interspace. In
Harengula the posterior anal rays are equal and the trans-
verse grooves on the scales are continuous; moreover, the -
lower jaw is more prominent and the sheath at the base of
the dorsal fin is lower than in Sardinella. I count 39 ver-
tebrein H. dispilonotus, 40 in H. maculosa and H. castelnaut,
41 in A. pensacola, 42 in H. schrammi, 43 im L/, zunast, 43
or 44 in H. macrophthalma, and 44 in H. punctata.
Like Sardinella, this genus occurs in the tropical Indo-
Pacific; but whereas Sardinella has four West African and
Mediterranean and only one Antillean species, Harengula
has four species on the coasts of Tropical America, but is
absent from the eastern Atlantic,
of the Genera Sardinella, Harengula, &c. 387
The species are mostly smaller and of less economic value
than those of Sardinel/a, and some of them are reputed
poisonous [cf. Cuv. & Val. xx. p. 295 (H. humeralis=ma-
crophthalma) and p. 377 (Meletta venenosa= H. punctata)].
Synopsis of the Species.
I. American.
A. 27 to 33 gill-rakers on lower part of anterior arch.
I. Depth of operculum 2, diameter of eye 22 to 3 in length of
head; depth of body 3 to 33 in the length.
1. maculosa.
11. Depth of operculum more than 3} length of head, diameter of
eye 3 to 33,in length of head.
Anal of 16-19 rays. Depth 3-32 in the length,
Henle teeta Raa eet aso sips. cin step: ome si ape'e whee 2. macrophthalma.
Anal of 15 rays. Depth 2% (-33) in the length,
BAM wala MANTIS wrshalt Ad wth viccatde Gade 3. thrissina.
B. 33 to 36 gill-rakers on lower part of anterior arch; depth of
operculum 3 to 3 length of head; eye 22 to 3 in head; depth
of body 22 to 23 inthe length .....;.. 4, pensacole.
II, Indo-Pacific.
A. 35 gill-rakers on lower part of anterior arch ; 36 to 38 scales in
Bi Lomi HOTA Y SETIOS. ie. k ve age cee ye 5. dispilonotus.
B. 30 to 34 gill-rakers on lower part of anterior arch ; 40 to 45 scales
in a longitudinal series,
Depth of body 22 in the length ................ 6. koningsberger?.
Depth of body 83 to 4 in the length, length of head
33 to 4; eye 3 to 33 in length of head...... 7. punctata.
Depth of body 4 to 44 in the length, length of head
4 to 42; eye 33 to 82 in length of head .... 8, schrammi.
C. More than 40 gill-rakers on lower part of anterior arch,
1. Depth of body 23 to 3} in length; eye 3 in length of head;
45 gill-rakers on lower part of anterior arch.
9. castelnaut.
2. Depth of body 3 to 4 in length; eye 33 to 32 in length of head,
About 50 gill-rakers on lower part of anterior arch ;
40 to 42 scales in a longitudinal series; ends
of caudal lobesblackish . 1... . 6.4 cece wees 10. wittata,
“About 50 gill-rakers on lower part of anterior arch ;
44 to 46 scales in a longitudinal series;
caudal fin uniform +). oe a Apel 11. zunas?.
70 gill-rakers on lower part of anterior arch .... 12. nymphea.
1. Harengula maculosa.
Harengula maculosa, Cuy. & Val. Hist. Nat. Poiss, xx. 1847, p. 292,
Alosa apicalis, Mull, & Trosch. in Schomburgk, Hist. Barbadoes, p. 678
(1848).
Harengula jaguana, Poey, Rep. i. p. 190 (1866).
Clupea macrophthaima, Giinth, Cat. Fish. vii. p. 421,
388 Mr. C. T. Regan on the Clupeid Fishes
Sardinella sardina, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1896;
p. 480,
Sardinella macrophthalmus, Jord. & Everm, 1. ¢.
Depth of body 3 to 3} in the length, length of head 3} to
34. Snout shorter than or nearly as long as diameter’ of eye,
which is 22 to 3 in the length ot head and greater than its
distance from lower edge of preeoperculum; maxillary ex-
tending to below anterior part or middle of eye; depth of
operculum 3 length of head. 27 to 33 gill-rakers on lower
part of anterior arch. About 40 scales in a longitudinal and
11 in a transverse series ; ventral scutes 15-17+10-12.
Dorsal 17-19. Anal 17-19. Pelvics 8-rayed, below middle
of dorsal. Anterior part of dorsal blackish superiorly.
Vertebree 40.
Several examples, 90 to 200 mm. in total length, from
Florida, the Bermudas, and the West Indies.
2. Harengula macrophthalmus.
Clupea macrophthalma, Ranzani, Noy. Com. Ac. Se. Inst. Bonon. y.
1842, p. 820, pl. xxiii.
Harengula latulus, Cuv. & Val. Hist. Nat. Poiss. xx. p, 280, pl. 595
(1847).
Harengula clupeola, Cuv. & Val. t. ¢. p. 289.
Harengula humeralis, Cuv. & Val. t. ¢. p. 298.
Alosa striata, Cuv. & Val. t. ¢. p. 429. |
Alosa bishopt, Mill, & Trosch. in Schomburgk, Hist. Barbadoes, p. 675
(1848).
“= Harengula sardina, Poey, Mem. ii. p. 310 (1860).
868).
Clupea humeralis (part.), Giinth. Cat. Fish. vii. p. 422 (1
3. Nat. Mus, xlvii.
.4
Sardinella humeralis (part.), Jord. & Everm. Bull. U.}
1896, p. 431.
Depth of body 3 to 32 in the length, length of head 33 to
32. Snout shorter than diameter of eye, which is 3 to 33 in
the length of head and not greater than its distance from
lower edge of preeoperculum ; maxillary extending to below
anterior 4 of eye or beyond ; depth of operculum more than
4 length of head. 29 to 33 gill-rakers on lower part of
anterior arch. About 42 scales in a longitudinal and 11 ina
transverse series; ventral scutes 16-19+12-14. Dorsal
16-19. Anal 16-19. Pelvics 8-rayed, below middle of
dorsal. A dark humeral spot. Vertebra 43-44.
Atlantic coast of Tropical America®.
Numeious examples, 80 to 130 mm. in total length, from
Florida, the West Indies, Fernando Noronha, and Bahia.
* Valenciennes described H. latula as a European species; it is
probable that this was a mistake, although it is not impossible that the
species, may cross the Atlantic.
of the Genera Sardinella, Harengula, &e. 389
3. Harengula thrissina.
Clupea thrissina, Jord. & Gilb. Proc, U.S. Nat. Mus. 1882, p. 353.
Sardinella thrissina, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1896,
p- 480.
Depth of body 23 (-34) in the length, length of head 33.
Snout a little shorter than diameter of eye, W hich is 31 in the
Jength of head and rather less than its distance from lower
edge of preeoperculum ; maxillary extending a little beyond
anterior 4 of eye ; depth of operculum more than 2 length of
head. 30 gill-rakers on lower part of anterior arch. 40
scales in a fon gitudinal and 12 ina transverse series ; ventral
scutes 18+ 11. (16+13). Dorsal 17. Anal 15. Pelvies
8-rayed, below middle of dorsal. A dark humeral spot.
Pacific coast of Mexico.
A single specimen, 115 mm. in total length, from Jalisco.
Harengula pensacole.
Clupea humeralis (part.), Giinth. Cat. Fish. vii. p. 422 (1868).
Harengula pensacole, Goode & Bean, Proc, U.S. Nat. Mus. 1879, p. 152.
Sardinella humeralis (part.), Jord. & Everm. Bull. U.S, Nat. Mus. xlvii.
1896, p. 431.
Sardinella sardina, Jord, & Everm. ¢. ec. 1900, fig. 193.
Depth of body 23 to 22 in the length, length of head 33 to
34%. Snout shorter than diameter of. eye, which is 22 to Pin
length of head and equal to its distance from lower ‘edge of
preoperculum ; maxillary extending to below anterior 4 of
eye or beyond ; depth of operculum % to 2 length of head.
33 to 36 gill-rakers on lower part of ‘anterior arch. About
40 scales in a longitudinal and 11 in a transverse series ;
ventral scutes 16-17+13. Dorsal 16-18. Anal 16-18.
Pelvics 8-rayed, below middle of dorsal. Vertebra 41.
Four specimens, 80 to 140 1 mm. in total length, from
Florida and Trinidad.
5. Harengula dispilonotus.
Harengula dispilonotus, Bleek. Nat. Tijdschr. Ned. Ind. iii, 1852, p. 456,
Clupea dispilonotus, Giinth. Cat. Fish. vii. p. 429 (1868); Bleek. Atl,
Ichth. vi. p. 111, Clup. pl. i, fig. 3 (1872); Weber & Beaufort,
Fish. Indo-Austral. Arch. ii, p. 69 (1913).
Depth of body 3 to 34 in the length, length of head 32 to
4. Snout a little aioutce than diameter of eye, which is Ain
the length of head ; maxillary extending to below anterior }
of eye. 369 gill- rakers on lower part of “anterior arch. 36 to
38 scales in a longitudinal, 11 or 12 in a transverse series ;
390 Mr. C. T. Regan on the Clupeid Fishes
ventral scutes 14-16+11-13. Dorsal 18-20. Anal 16-18.
Pelvics below anterior 4 of dorsal. _ A dark spot or ocellus on
back at base of posterior dorsal rays, a second beliind it.
KE. Indian Archipelago.
Six specimens, 75 to 100 mm. in total length, including
the type of tie species.
6. Harengula koningsbergert.
Clupea koningsbergeri, Weber & Beaufort, Verh. Akad. Amsterdam,
xvii. no. 8, 1912, p.14; Fish. Indo-Austral. Arch. i. p. 72 (1918).
Depth of body 22 in the length, length of head 33.
Snout shorter than diameter of eye, which is 22 in length of
head ; maxillary extending to below anterior $ of eye ora
little beyond. 33 gill-rakers on lower part of anterior arch,
42 scales in a longitudinal and 12 in a transverse series ;
ventral scutes 17-18+11-12. Dorsal 18-19. Anal 20-21.
Pelvics below or a little in advance of middle of dorsal.
A specimen of 115 mm, from N.W. Australia, and one of
75 mm. from New Guinea; the species was described from
the Aru Is.
7. Harengula punctata.
Clupen punctata, Riippell, Neue Wirbelth. Fische, p. 78, pl. xxi. fig. 2
(1840).
Clupea quadrimaculata, Riippell, t. ¢. p. 78, pl. xxi. fig. 3.
Sardinella lineolata, Cuv. & Val. Hist. Nat. Poiss. xx. p. 272 (1847).
Harengula punctata, Cuv. & Val. t. c. p. 297; Sauvage, Hist. Mada-
gascar Poiss. p. 493 (1891).
Harengula bipunctata, Cuv. & Val. t. e. p. 298.
Melcetta obtusirostris, Cuv. & Val. t. ¢. p. 575.
Meletta venenosa, Cuv. & Val. t. c. p. 877.
Harengula moluccensis, Bleek. Nat. Tijdschr. Ned. Ind. iv, 1853, p. 609.
Harengula kunzet, Bleek. id. xii. 1857, p. 209.
Harengula spilurus, Guichenot, in Maillard, He de la Réunion Poiss.
p. 16 (1865) ; Sauvage, op. cit. p. 493, pl. xviii. fig. 3.
Clupea moluccensis, Giinth. Cat. Fish. vii. p. 427 (1868); Bleek, Atl.
Ichth. vi. p. 107, Clup. pl. v. fig. 2 (1872); Weber & Beaufort,
Fish. Indo-Austral. Arch. ii. p. 81 (1918).
Clupea venenosa, Giinth. t. c. p. 449; Weber & Beaufort, ¢. c. p. 77.
Clupea kunzei, Bleek. Atl. Ichth. vi. p. 107, Clup. pl. v. fig. 1 (1872) ;
Day, Fish India, p. 636.
Clupea dubia, Bleek. t. c. p. 108.
‘Harengia stereolepis, Ogilby, Proc. Linn. Soc. N.S. Wales, xxii. 1897,
. 769.
Chived mizun, Kishinouye, Journ. Imp. Fisheries Bureau, Tokyo, xiv.
1907, p. 98, pl. xx. fig. 3.
Depth of body 3} to 4 in the length, length of head 33 to 4.
Snout as long as or shorter than diamcter of eye, which
of the Genera Sardinella, Harengula, dec. 391
is 3 to 34 in the length of head, equal to or greater than its
distance from lower edge of preeoperculum ; maxillary ex-
tending to below anterior } of eye or a little beyond ; depth
of operculum about $ length of head. 30 to 34 gill-rakers on
lower part of anterior arch. 42 to 45 scales in a longitudinal,
11 or 12 ina transverse series; ventral scutes 16-20 + f1-14.
Dorsal 17-19. Anal 17-19. Pelvics below or a little in
advance of middle of dorsal. Anterior part of dorsal blackish
superiorly. Vertebra 44.
Tropical Indo-Pacific, from E. Africa to the Paumotu
Archipelago.
Numerous examples, 60 to 130 mm, in total length, in-
cluding the types of Z/. moluccensis and H. kunzet.
8. LHarengula schrammi.
Alosa schrammi, Bleek. Verh. Bat. Gen. xxii, 1849, Bali, p. 11.
Clupea schrammi, Bleek. Atl. Ichth. vi. p. 109, Clup. pl. xiv. fig. 3
(1872) ; Weber & Beaufort, Fish. Indo-Austral Arch. ii. p. 88 (1918).
Depth of body 4 to 44 in the length, length of head 4 to
45. Snouta little longer than diameter of eye, which is 34
to 33 in the length of head ; maxillary extending to below
anterior 3 of eye. 32 to 34 gill-rakers on lower part of ante-
‘rior arch. About 42 scales in a longitudinal and 12 in a
transverse series ; ventral sentes.17-18+12. Dorsal 18-19.
Anal 18-19. Pelvics 8-rayed, below or a little in advance of
middle of dorsal. Vertebree 42.
Malay Archipelago.
Four specimens, 100 to 115 mm. in total length, from
Misol and Goram ; with these I lave compared the type, a
small fish in bad condition.
9. Harengula castelnau’.
Kowala castenaui, Ogilby, Proe. Linn. Soc. N.S.Wales, xxii. 1897,
p. 66.
Depth of body 24 to 3} in the length, length of head about
4, Snout a little shorter than diameter of eye, which is 3 in
the length of head; maxillary extending to below anterior 4
or middle of eye. 45 gill-rakers on lower part of anterior
arch. 40 to 42 scales in a longitudinal, 12 in a transverse
series ; ventral scutes 16-19+10-13. Dorsal 17-19. Anak
18-21. Pelvics below middle or anterior 4 of dorsal. Tip.
of dorsal and ends of caudal lobes blackish. -
New South Wales.
Nine specimens, 130 to 180 mm. in total length.
392 Mr. C. T. Regan on the Clupeid Fishes
10. Harengula vittata.
Clupeonia vittata, Cuy. & Val. Hist. Nat. Poiss. xx. p. 352 (1847).
Alausa melanurus, Cuv. & Val. t. ¢. p. 441.
Clupeonia jussieui, Cuv. & Val. op. cit. pl. 599.
Clupea melanura, Giinth, Cat. Fish. vii. p. 449 (1868); Bleek. Atl,
Ichth. vi. p. 111, Clup. pl. xi. fig. 5 (1872); Weber & Beaufort,
Fish. Indo-Austral. Arch. ii. p. 72 (1913).
Harengula vanicoris, Jord. & Seale, Bull. U.S. Bureau Fisheries, xxv.
1906, p. 187.
Clupea rechingeri, Steind. Sitzungsb. Akad, Wien, cxv. 1906, p. 1424.
Depth of body 34 to 33 in the length, length of head 33 to
4, Snout as long as diameter of eye, which is 32 to 52 in
the length of head; maxillary extending to below anterior 4
of eye or beyond. 50 gill-rakers on lower part. of anterior
arch. 40 to 42 scales in a longitudinal, 12 in a transverse
series; ventral scutes 17+13. Dorsal 16. Anal 15-19.
Pelvics below anterior } of dorsal. Basal part of caudal
dusky ; ends of lobes blackish. 7
~ Indo-Pacific.
Two specimens, 80 and 90 mm. in total length, from
Celebes and Raiatea.
11. Harengula zunast.
Clupea kowal (non Riipp.), Schleg. Faun. Japon. Poiss. p. 255, pl. vil.
fig. 1 (1846).
Harengula zwnasi, Bleek. Verh. Bat. Gen. xxvi. 1854, p. 117.
Clupea kowal (part.), Giinth. Cat. Fish. vil. p. 450 (1868).
Clupea zunasi, Giinth. t. c. p. 451; Kishinouye, Journ. Imp. Fisheries
Bureau, Tokyo, xiv. 1907, p. 98, pl. xx. fig. 4.
Depth of body 31 to 34 in the length, length of head 4 to
41. Snout nearly as long as or shorter than diameter of eye,
which is 3} to 32 in the length of head; maxillary extending
to below anterior 3 of eye or beyond. About 50 gill-rakers
on lower part of anterior arch. 42 to 46 scales in a longi-
tudinal series, 11 or 12 in a transverse series; ventral scutes
17-20+11-15. Dorsal 17-19. Anal 17-20. — Pelvies .
below anterior 4 of dorsal.
China, Corea, and Southern Japan.
Several specimens, 90 to 140 mm. in total length, from
Amoy, China, and from Japan.
12. Harengula nymphea.
Clupea nymphea, Richards, Ichthyol. China, p. 804 (1848); Ginth.
Cat. Fish. vii. p. 428 (1868).
Depth of body 3} in the length, length of head 4, Snout
of the Genera Sardinella, Harengula, dc. 393
as long as diameter of eye, which is 34in fae length of head ;
maxillary extending to below anterior + of eye. 70 ell.
rakers on lower part of anterior arch. About 40 scales in a
Jongitudinal and 13 in a transverse series; ventral scutes
18+11. Dorsal 17. Anal20. Pelvies 8-rayed, a little in
advance of middle of dorsal.
China.
Here described from the type, 120 mm. in total length.
Lite, Jordan & Evermann, 1896.
Bull. U.S. Nat. Mus. xlvii. p. 429.
This genus is close to Harengula, but is distinguished by
the well-defined bluish-silvery lateral band and by ‘the absence
of scales on the lobes of the caudal fin. The dermal fold on
the edge of the cleithrum described in Sardinella, which is
developed to a greater or Jess extent in Harengula, is absent
‘in this genus.
Three species from America.
1. Lile stohifera.
Clupea stolifera, Jord. & Gilbert, Proc. U.S. Nat. Mus. 1881, p. 339.
Sardinella stolifera, Jord. & Everm. Bull. U.S. Nat. Mus. xlvii. 1896,
p- 431, and 1900, fig. 194.
Depth of body 31 to 4 in the length, length of head 4}
to 43. Diameter of eye 3 in length of head; maxillary
extending to below anterior + of eye. 36 gill-rakers on lower
part of anterior arch. About 40 scales in a longitudinal, 10
or 11 in a transverse series ; ventral scutes 16-18+10-12.
Dorsal 15-17 ; origin nearer to end of snout than to base of
caudal. Anal 16-19. Pelvics 8- rayed, below anterior rays
of dorsal. A silvery lateral band; ends of caudal lobes
blackish. Vertebree 40 (42).
Pacific coast of Mexico.
Hight specimens, 90 to 100 mm. long.
2. Lile piquitinga.
Sardinella piquitinga, Schreiner & Ribeiro, Arch. Mus. Rio Janeiro,
xii. 1903, p. 72.
Depth of body about 34 in the length, length of head 33
to 4. Diameter of eye 22 in length of head; maxillary
extending to below anterior + of eye. 33 gill-rakers on lower
part of anterior arch. About 40 scales in a longitudinal and
10 in a transverse series; ventral scutes }<17+ 10-11,
Ann. & Mag. N, Hist. Ser, 8. Vol, xix. 26
394 On the Genera Sardinella, Harengula, d&c.
Dorsal 15-18; origin nearer end of snout than base of caudal.
Anal 17-19. Pelvics 8-rayed, below anterior part of dorsal.
A silvery lateral band. . Vertebree 40. .
Coast of Brazil.
Three specimens, 80-90 mm. long, from Pernambuco.
3. Lile platana, sp. n.
Depth of body 3 in the length, length of head 4. Diameter
of eye 3 in length of head; maxillary extending to below
anterior + of eye; 23 gill-rakers on lower part of anterior
arch. About 40 scales in a longitudinal series; ventral
scutes 18+10.. Dorsal 14; origin equidistant from end of
snout and base of caudal. Anal 18. Pelvics below origin of
dorsal. A silvery lateral band.
La Plata.
A single specimen of 45 mm.
HERINGIA.
Heringia, Fowler, Proc. Acad. Philadelphia, 1xiii. 1911, p. 207.
Rhinosardinia, Kigenmann, Mem. Carnegie Mus. v. 1912, p. 445.
Form moderately elongate, strongly compressed ; abdomen
sharp-edged. Mouth small, toothless; lower jaw rather
prominent, upper without distinct notch; maxillary with a
retrorse spine near its proximal end; 2 supramaxillaries.
Operculum smooth ; suboperculum rounded. Dorsal median,
of 13 to 15 rays, without scaly sheath ; anal of 15 to 17 rays,
with a low scaly sheath; caudal forked, scaly at base, without
alar scales. Pelvics 8-rayed, a little in advance of dorsal.
38 to 40 scales in a longitudinal, 9 or 10 in a transverse
series ; each scale crossed by a transverse groove, from which
2 (fewer anteriorly, more posteriorly) run backwards to the
free edge, which is entire. Vertebree 39.
1. Heringia amazonica.
Clupea amazonica, Steind. Sitzungsb. Akad. Wien, lxxx. pt. 1, 1880,
p- 188.
Rhinosardinia serrata, Kigenm. Mem. Carnegie Mus. v. 1912, p. 445,
pl. Ixil. figs. 3, 4.
Depth 3 to 34 in length, length of head 44 to 5. Diameter .
of eye 3 to 3} in length of head; maxillary extending to
below anterior margin or anterior } of eye. 35 gill-rakers
on lower part of anterior arch. 38 to 40 scales in a longi-
tudinal, 9 or 10 in a transverse series ; ventral scutes keeled
oer
On new Weevils from India. 395
and pointed, 16-17+10-11. Dorsal 13-15 ; origin nearer
end of snout than base of caudal. Anal 15-17. Pelvics a
little in advance of dorsal. Silvery ; back bluish.
Amazon ; rivers of Guiana.
Four specimens, 60 mm. long, co-types of A. serrata,
from British Guiana.
2. Heringia bahiensis.
Pellonula bahiensis, Steind. Sitzungsb. Akad. Wien, lxxx. 1880, p. 181,
pl. iii. fig. 2.
Sardinella pernambucana, Schreiner & Ribeiro, Arch, Mus. Rio
Janeiro, xii. 1903, p. 72.
Maxillary extending to below anterior } of eye. 42 to 44
scales in a longitudinal series. Dorsal 16-17. Anal 16-18.
A more or less distinct bluish-silvery lateral band.
Joast of Brazil.
This species seems to be scarcely distinct from the preceding,
examples of which also show a faint lateral band in certain
lights.
XXXV.—On new Weevils of the Genus Mecysmoderes from
India. By Guy A. K. MarsHAtt, D.Sc.
Subfamily CevruorruyNnoHINa.
Mecysmoderes memecylonis, sp. n.*
g. Colour black ; the head with moderately dense seti-
form scales, those in the middle being brown and the others
white ; the pronotum sparsely set with inconspicuous dark
sete, and with a longitudinal patch of transverse white sete
on the anterior half on each side; the elytra with a longi-
tudinal stripe on the first interval bordering the thoracic
spine and composed of small, ovate, overlapping, yellowish-
white scales (the basal half of the stripe usually brownish),
being continued behind the spine as a double row of elongate
brown scales, partly replaced by white ones near the apex ;
the remaining intervals each bear a single row of similar
brown scales, with a few white ones here and there, except
along the immediate base, where all the scales are white; the
* The types of all the species described here are in the British
Museum. :
26*
396 Dr. G. A. K. Marshall on
lower surface fairly closely covered with separated white
scales throughout.
Head minutely shagreened, with an indistinct network of
raised lines and a strong longitudinal carina on the vertex ;
the forehead only slightly depressed below the level of the
eyes and with a faint central carina. Rostrum moderately
stout, very slightly widened from the middle to the apex; the
basal half shagreened, with a well-marked central carina and
another on each side of it, the apical half shining in the
middle and with several rows of very shallow punctures at
the sides. Antenne with the scape sharply pointed at the
apex; the funicle with joint 1 about as long as 2 and twice
as thick, 3 a trifle longer than 2, 4 shorter than 2, and 5 and
6 shorter still and bead-like, 5 as long as broad, 6 transverse.
Prothorax very strongly and regularly convex transversely,
the sides about equal to half the basal margin, almost straight
and with a moderate apical constriction, the front margin
produced into a sharp obtuse angle in the middle, the basal
spine very long (more than one-third the length of the elytra),
the basal margin forming on each side an angle of about 75°
with the main axis ; the upper surface finely shagreened and
dull, with a strong complete central carina and a large-meshed
reticulation of fine raised lines, without any conspicuous
depressions ; the dorsal outline moderately curved, sloping
gradually from the base to one-third, and thence much more
steeply. lytra about as long as their united width at the
shoulders and very broadly rounded behind, the humeral
slope forming an angle of about 45° with the suture; thie
strie broad, shining, and deeply punctate, the intervals.
rather narrower, carinate and minutely granulate; the dorsal
outline convex, deepest before the middle, sloping gradually
to the base and more steeply behind. Legs black, clothed
with separated setiform white scales, the tarsi piceous ; all
the femora with a sharp tooth, the dorsal apical fringe of the
hind tibia longer than the first tarsal joint, the claws unusually
small, Sternum: the space between the front coxee as broad
as the rostrum; the meso- and metasternum broadly exca-
vated, the end of the furrow sloping gradually up to the base
of the metasternum, and the mesosternum being almost
horizontal.
Length 13-14, breadth 1}-1} mm.
Mysore: Madhavgiri, on leaves of Memecylon umbellatum
(H. Mann, Pusa Coll.). Assam: Sylhet (type).
Allied to M. nigrorufa, Mots., but, apart from its red-
brown colouring, distinctly narrower form, and unarmed
femora, that species has the prothorax much more finely
new Weevils from India, 397
reticulate, its dorsal outline being almost flat, the front
margin rounded, and the central carina flattened; the elytra
bear a single row of white scales on the first interval behind
the thoracic spine, and the other intervals are set with
irregular minute dark sete; and the metasternum is only
shallowly excavated in front. ©The sternal furrow of MW. me-
mecylonis is very similar to that of MZ. carinata, Fst., but in
the latter the margins of the apex are overhanging.
Mecysmoderes verrucosa, sp. 0.
3. Black or brownish black, shining and sparsely set with
very short recumbent white sete ; the elytra with an elongate
patch of brown scales along each side of the thoracic spine ;
the tarsi testaceous.
Head unevenly reticulate, with a well-marked central
carina running from the vertex to the edge of the interocular
depression. ostrum stout, about as long as the front femur,
gradually widening from base to apex, somewhat compressed,
the greatest depth exceeding the width, the dorsal outline
strongly curved, but with a depression at the base and a deep
sinuation in the middle ; the upper surface rugosely punctate,
with a strong central carina (obsolete in the median depres-
sion and ceasing a little behind the insertion of the antenne)
and two finer irregular ones on each side. Antenne with the
apex of the scape not produced into a sharp point; the
funicle with the two basal joints equal in length, 3 rather
shorter, 4 again much shorter, 5 and 6 bead-like. Prothorax
with the sides about as long as half the basal margin, slightly
rounded in the posterior half, abruptly narrowed in front, the
apical constriction being broad and collar-like, the front
margin rounded dorsally and slightly produced in the middle ;
the whole of the central portion of the disk is raised into a
large boss-like prominence, the top of which bears an elevation
on each side, the space between them being longitudinally
flattened; the upper surface is coarsely reticulate, with-a
sharp central carina running from the apical edge to the tip
of the basal spine, and on each side of it on the apical collar
is another sharp carina—these being convergent behind in
the type and parallel in a second specimen. Lilyéra about as
long as their width at the shoulders, which are very promi-
nent and rounded, the humeral slope forming an angle of
about 70° with the main axis; the strie broad and shallow, —
rather sinuous, much more irregular than usual, and con-
taining very unequal punctures, the intervals very uneven,
bearing elongate tubercles and set with scattered minute
398 Dr. G. A. K. Marshall on
granules ; the more conspicuous tubercles are situated as
follows :—interval 2 with a small one at the top of the de-
clivity, on interval 3 a small one near the base and a very
large one near the middie, on interval 5 a large one near the
base and a small one close behind it, on interval 7 a medium
one near the shoulder and another about the middle, on
interval 9 a large one about the middle, and a large one just
behind the shoulder on interval 10; near the apex a trans-
verse row of tubercles on intervals 8-7, those on 5 and 6
being elongate and the others short ; behind these is a broad
transverse impression. Legs rather long, with scattered
recumbent short white sete ; the femora rugosely punctured,
armed with a stout tooth, and transversely impressed exter-
nally near the apex ; the tibiz rather slender, scarcely dilated
at the apex and there clothed externally with golden-brown
hairs, which form a dorsal fringe hardly as long as the first
tarsal joint. Sternum: the mesosternum sloping very steeply
and very shallowly impressed in the middle to receive the
apex of the rostrum, the metasternum not excavated, and the
space between the front coxze as wide as the rostrum.
Length 24-33, breadth 2-23 mm.
AssaM: Patkai Hills (W. Doherty).
A strikingly distinct species.
Mecysmoderes metasternalis, sp. n.
3. -Colour varying from black to red-brown, variegated
with brown, yellowish and white scales or sete; the head
with fairly dense narrow white scales; the prothorax with the
entire sides and lower surface bearing close whitish scales
and setee, a triangular patch of white sete on the apical third
of the median area (its base on the front margin), and some
pale setee on each side of the basal third of the central carina,
thus leaving a broad, roughly X-shaped, bare discal patch ;
the elytra with a very long raised stripe of scales on each
side of the thoracic spine extending for more than half
their length, the basal two-thirds being very dark brown, the
apical third whitish ; these scales are very closely packed and
obliquely raised or almost erect; a large lanceolate sub-
humeral patch of dense whitish scales (continuous with
similar scaling on the whole lower surface), the inner edge of
which reaches the sixth stria at the base and extends obliquely
backwards to nearly the middle of the lateral margin ; the
apical margin and the posterior half of the lateral margin
with an irregular edging of white scales, and a row of
yellowish scales on the basal fourth of the second interval ;
new Weevils from India. 399
the rest of the elytra clothed with dark brown scales varie-
gated with narrower light brown and white scales, and with
a conspicuous spot of broader white scales before the middle
on the eighth interval.
Head with the inner edge of the eyes strongly raised above
the general level; the forehead very narrow, not broader than
the antennal club, and broadly impressed behind ; the vertex
rugosely punctured, with a well-marked central carina and
densely clothed with elongate pale scales. Rostrum piceous
black, with the extreme apex reddish; long, slender, and
cylindrical, extending beyond the middle of the metasternum ;
rather dull and very finely aciculate, with two short indistinct
furrows on each side at the base, and no central carina; the
punctures indistinct at the base, shallow and widely separated
for most of the length, and deeper near the apex. Antenne
testaceous ; the scape without any apical process ; the three
basal joints of the funicle subequal in length, joints 4 and 5
a little longer than broad, 6almost globular. Prothorax with
the basal spine very long, about as long as the prothorax,
and extending along nearly two-thirds of the elytral suture ;
the sides (viewed vertically) almost straight and shorter than
half the basal margin ; the front margin somewhat produced
dorsally, the produced portion elevated and truncate in the
middle, so that when seen from in front it appears as a flat-
tened triangle ; the upper surface covered with large irregular
reticulate punctures, the median area raised and strongly
compressed in the anterior half, being bounded by a sharp
carina on each side to about the middle, the central carina
well-marked throughout ; the dorsal outline distinctly convex
and deepest about the middle ; the scales on the sides of the
prothorax are mostly setiform, but there is a patch of much
broader scales just below the outer carina and another patch
some distance below that. lytra about as long as broad ;
the striz broad and deep, with large punctures and without
rows of scales, except in the first stria; the intervals some-
what broader than the striz and irregularly granulate ; the
scales on the greater part of the surface are small, narrow,
parallel-sided, and truncate at the tip, but those in the raised
patch along the suture, in the basal stripe on interval 2, the
subhumeral patch, and the white border are broader and
elliptical in shape. Legs piceous, the tarsi paler; the femora
densely clothed with broad pale scales and all with a strong
tooth ; the dorsal apical fringe of the posterior tibie much
longer than the first tarsal joint. Sternum with the front
coxe widely separated; the mesosternum almost horizontal
and broadly and shallowly impressed; the metasternum with
400 Dr. G. A. K. Marshall on
a broad and deep furrow throughout, which is quite open at
the posterior margin.
Length 23, breadth 2mm.
Burma: Tavoy, Tenasserim (W. Doherty).
The most striking features of this species are the structure
of the front margin of the pronotum, the very long pro-
thoracic spine, the elevated sutural scales on the elytra, and
the open continuous furrow on the metasteraum.
Mecysmoderes subhumeralis, sp. n.
¢. Colour varying from piceous black to red-brown, the
upper surface sparsely clothed with yellowish hair-like
scales; the sutural patch of scales on the elytra extending
for about one-third of their length, the basal two-thirds of
the patch being black and the apical third whitish; on the
seventh interval of the elytra there is a small] spot of white
scales before the middle; the lower surface sparsely clothed
with pale scales, except for a dense patch on the upper half
of the mesosternal epimeron.
Head reticulate, the central carina continued well on to
the forehead, which is almost as broad as the base of the
rostrum and strongly impressed. Jtostrum short and very
stout, hardly longer than the front femur and only just
reaching the metasternum, parallel-sided in the basal two-
thirds, slightly wider at the apex, its dorsal outline strongly
convex, and its depth greater than its width in the middle;
the upper surface with indistinct confluent punctation, a
strong central carina for three-fourths its length, and a less
distinct lateral one. Antenne testaceous ; the scape without
an apical process ; the funicle with joints 1 and 2 subequal,
3 distinctly shorter, 4 longer than broad, 5 and 6 subglobular.
Prothorax with the sides equal to half the basal margin,
straight from the base to beyond the middle and then rather
abruptly constricted; the basal margin more nearly trans-
verse than in most species, the spine short, about one-fourth
the length of the suture; the front edge truncate dorsally and
narrowly marginate ; the whole discal area forming a rounded
elevation laterally compressed in the anterior half, irregularly
and unequally reticulate, with a strong and complete central
carina, on each side of it in the front half a short carina that
does not reach the margin, and beyond this another irregular
sinuate one that does reach the margin. Llytra about as
long as their greatest width, the striz broad and deep, with
separated punctures and each with a single row of distant
minute white sete ; the intervals costate and scarcely broader
new Weevils from India. 401
than the strize, each with a row of small granules bearing
short recumbent sete and some scattered yellowish hair-like
scales (especially in the basal half). Legs piceous, with the
tarsi paler ; the femora rugosely subgranulate, with sparse
whitish hair-like scales and each with a small tooth; the
dorsal apical fringe of the hind tibia not longer than the
first tarsal joint. Sternum with the front coxa widely
separated, but not enough to receive the whole rostrum ; the
mesosternum not impressed and sloping steeply ; the meta-
sternum simple. ;
Length 2?, breadth 12 mm.
ASSAM: Patkai Hills (W. Doherty, type) ; Sudiya
(Doherty).
Very similar superficially to 1M. memecylonis, sp.n., but
differing markedly in the structure of the sternum, non-
angulate front margin of the prothorax, short thoracic spine,
short third funicular joint, setigerous elytral strie, etc.
Mecysmoderes pusio, sp. n.
$ ¢. Colour piceous, with greyish-white scaling, the
rostrum and tarsi often paler; the scales on the head are
narrow and fairly dense ; on the pronotum they are more
hair-like and sparser on the disk, being shorter and denser at
the sides and margins and in the central furrow ; on the
elytra there is a short stripe of almost circular small scales on
the basal fourth of the first interval, the remaining intervals
each bearing a single regular row of very narrow and short
scales, except on intervals 9 and 10, at the bases of 3 and 5,
and on the humeral callus, where the scales are duplicated or
more numerous ; on the lower surface the scales are larger,
oval, and fairly close, but usually not quite contiguous.
Head shagreened and shallowly reticulate, with an in-
distinct central. carina on the vertex only, the forehead
shallowly impressed and broader than the club of the an-
tenna. Rostrum long, slender, and cylindrical, not quite
reaching the hind margin of the metasternum (d) or ex-
tending slightly beyond it (2); very faintly tricarinate in
the basal part, the more shining apical area with indistinct
shallow separated punctures. Antenne testaceous, the scape
with a short apical pointed process, the funicle with the two
basal joints subequal, the third slightly longer. Prothorax
with the sides about as long as half the basal margin, scarcely
rounded and very shallowly constricted at the apex; the
basal margin nearly transverse externally, the central spine
short, only about one-eighth the length of the suture, the
402 Dr. G. A. K. Marshall on
apical margin truncate dorsally ; the upper surface moderately
convex, with well-marked carinate reticulations, except on
the apical collar, a broad shallow central furrow from the
collar to beyond the middle, containing a deep rounded fovea
in front and changing near the base into a low carina that
runs on to the spine. Hilytra about as long as their greatest
breadth, the striz very broad, with deep separated punctures
and quite bare, the intervals subcarinate, each with a row of
spaced setigerous granules. Legs piceous, with the tarsi
paler, clothed with separated, long, narrow, pale scales ; the
femora with a minute tooth, which is situated much lower
down than usual, being about in the middle of the limb and
partly concealed by scales; the dorsal apical fringe of the
hind tibize hardly as long as the first tarsal joint. Sternum
with the front coxee widely separated for the reception of the
rostrum ; the mesosternum broadly and deeply excavated,
almost horizontal in the middle; the metasternum with a
broad central furrow throughout, which is not enclosed
behind.
Length 13, breadth 1,4, mm.
Burma: ‘Tavoy, Tenasserim (W. Doherty).
Very similar to M. carinatus, Fst., in which the thorax
is of quite the same type and the sculpturing of the elytra is
similar ; but that species differs markedly in its metasternum,
which has a deep overhanging excavation in its anterior part
only, the femora have no tooth, there is no carina on the
head, etc.
Mecysmoderes pectinipes, sp. n.
Colour black, variegated with grey and brown sete and
scales; the prothorax clothed above with recumbent sete
only, the brown ones being more numerous on the disk ; the
elytra with a sutural stripe of dense whitish scales extending
for more than one-third of the suture, the remaining intervals
bearing linear scales (not less than two, and more often three
deep), which are grey at the base and mostly dark brown
elsewhere, variegated here and there with grey scales; the
lower surface rather densely clothed with broad whitish
scales. Immature specimens are sometimes testaceous, with
the thoracic carina and spine black; and various intergrades
occur between this and the black mature form.
Head with coarse close confluent punctation, the central
carina indistinct or absent, the forehead not at all impressed.
Rostrum black, with the apex reddish; as long as the front
new Weevils from India. 403
femur (¢) or a little longer ( 2 ), cylindrical, slightly tapering
to the apex from the insertion of the antenne ; the upper
surface without any distinct carine, but closely and con-
fluently punctate from the base to the antennw, beyond which
the punctures are finer and isolated, especially in the ¢.
Antenne testaceous ; the scape without any apical process ;
the funicle with joints 1 and 2 subequal, 3 shorter, 4-6 bead-
like. Prothorax with the sides gently rounded and each
about as long as half the basal margin, the apical constriction
slight; the basal spine nearly one-third the length of the
suture, the apical dorsal margin feebly angulate in the
middle; the upper surface is regularly and gently convex,
without elevations or depressions, evenly set with longitu-
dinally confluent punctures, with a low, broad, complete
central carina and a fine, short, apical one at some distance on
each side of it ; the dorsal outline almost flat, deepest near
the base, and sloping forwards. Llytra as long as their
greatest width, the striz broad and deep, and containing
single rows of narrow scales; the intervals evidently broader
than the strie, flat, and closely and strongly punctate. Legs
black, with the tarsi red-brown, rather thinly clothed with
narrow pale scales ; the femora with a stout tooth; all the
tibize strongly dilated at the apex, which bears a double row
of stout testaceous spines instead of the usual bristles.
Sternum with the front coxze so narrowly separated that the
rostrum cannot lie between them; the mesosternum almost
vertical and not impressed; the metasternum simple.
Length 22-3}, breadth 13-2 mm.
Manpras: Nilgiri Hills (A. LZ. Andrewes).
A very distinct species on account of its dilated and pecti-
nate tibiz, closely approximated front coxz, and unimpressed
forehead. .
Mecysmoderes tenutrostris, sp. n.
@. Colour reddish brown above, with the head, the
thoracic carina and spine, and the basal margin of the elytra
blackish ; the head and pronotum with rather sparse yellowish
sete, the latter with a small patch of ovate whitish scales on
each side before the middle just below the outer carina; the
elytra with the sutural stripe of ovate scales extending over
nearly half the suture, the scales being blackish except for a
short distance at the apex, where they are whitish ; a few
ovate whitish scales at the base of interval 2 and near the
apex of the suture, the rest of the intervals being clothed
404 On new Weevils from India.
(usually about three deep) with intermingled yellowish and
blackish narrow setiform scales ; the lower surface blackish
(except the prosternum, which is reddish) and densely clothed
with broad ovate whitish scales except on the mesepimeron,
the lateral lobe of the mesosternum, and the front half of the
metepisternum, where the scales are markedly narrower and
have a brownish tinge.
Head coarsely and confluently punctate, with a carina on
the vertex; the forehead strongly depressed, very broad
behind, and narrowing in front to the width of the rostrum.
Rostrum extremely long and slender, cylindrical and mode-
rately curved, reaching to the hind margin of the first ventral
segment ; a smooth central line on the basal third and rows
of punctures on each side of it, the apical area smooth
and impunctate. -Antennw testaceous brown, inserted behind
the middle of the rostrum ; the scape without any apical
process ; the funicle with joints 1 and 3 subequal, 2 markedly
longer, 4 much longer than broad, 5 and 6 globular. Pro-
thorax with the sides as long as half the basal margin, gently
rounded and rather broadly constricted at the apex, the apical
margin being truncate (as seen from above), but raised in the
middle so as to form a vertical angle; the basal margin rather
less oblique on each side than usual, the spine very long and
slender, extending nearly for one-half the suture ; the upper
surface not very convex, coarsely and subreticulately punctate,
with a well-marked and complete central carina, and a promi-
nent carina on each side of it in the anterior half, below
which the sides are compressed. Jilytra as long as their
greatest width, the strice deep but not broad, each containing
a row of spaced yellowish scales; the intervals evidently
broader than the striz, flat, and coarsely punctate. Legs
rather long and slender, clothed with separated yellowish
setiform scales ; the femora with a stout tooth ; the dorsal
apical fringe of the hind tibie not longer than the first tarsal
joint. ,Sternum with the front coxee moderately widely sepa-
rated; the mesosternum quite flat and almost vertical ; the
metasternum simple.
Length 22-24, breadth 14-2 mm.
Mapras: Nilgiri Hills (1. L. Andrewes).
Differs from all the Indian species known to me by its
very long and slender rostrum ; the unusual length of the
second joint of the funicle is also noteworthy.
On a new Holothurian from Bermuda. 405
XXXVI.—Oecurrence of a Holothurian new to the Fauna of
Bermuda. By W. J. Crozier.
(Contributions from the Bermuda Biological Station for
Research.—No. 61.)
THe West-Indian affinities of the Bermudan fauna and flora
(Britton, 1912, p. 193) have been evident to every student
of these regions ; yet, as Pilsbry (1900, p. 494) remarks in
considering the Pulmonates, there is “ abundant evidence of
what we call chance, or the rigorous selective action of an
over-sea journey, in the Bermudian assemblage.” Con-
tinued collecting is disclosing further additions to the marine
population in the shape of species identical with well-known
West-Indian forms *. Inone such case, which is the subject
of this note, it seems to me that the addition may legiti-
mately be considered as of recent date.
The pedate Holothurians of Bermuda waters have been
collected ever since 1888 by Heilprin, Verrill, Clark (1901),
and others. Each of the five species previously reported
(Cucumaria punctata, Stichopus mobbii, Holothuria surinam-
ensis, H. captiva, and H. rathbuni) is well represented in
the West-Indian area. Certain conspicuous Antillean types
have, however, been lacking in the Bermudan collections ;
Actinopyga and several species of Holothuria are in this
category T.
I have had oceasion to examine with care several thousand
specimens of Stichopus mébii, Semp.—with which, on super-
ficial examination, Actinopyga might conceivably be con-
fused,—and have, indeed, given particular attention to Holo-
thurians collected at many points in Bermuda, No unusual
specimens were observed until July 3, 1916, when there was
secured from a depth of about 6 feet beneath low water, in
the channel entrance to Hungry Bay (on the exposed south
* Among the Enteropneusta, for example, of which an illustrated
account is in course of preparation, at least two of the four or more
species which I have found occur also in the Bahamas and at Jamaica,
as well (probably) as at other stations in the West Indies.
+ Of the five apodous species found here (Clark, 1907), only two.
(Chirodota rovifera and Synaptula hydriformis) are typically West-
Indian, while two others (Leptosynapta inherens and L. roseola) are
northern forms ; the remaining one (L. acanthia) appears to be peculiar
to the Bermudas.
406 On a new Holothurian from Bermuda.
shore), an individual which was at once seen to be peculiar.
The Holothurian was about 25 em. in length, and of a deep
yellowish-brown colour, unmarked by spots of any other
hue. Anal teeth were particularly prominent, and the pedi-
ciles and ventral surface were tinged with greenish pigment,
such as one is accustomed to see in Holothuria captiva and in
FH]. surinamensis (Crozier, 1915, p. 274). Stichopus has no
anal teeth, and totally lacks this green pigment. The speci-
men was unfortunately lost before detailed examination could
be given it. Attempts to discover an -additional example
have thus far been fruitless.
The single specimen observed has, however, considerable
z00-geographical interest. It seems probable, from the
external characters above mentioned, that it isan Actinopyga,
presumably A. agasstzi (Selenka) *, which is known from
the Bahamas, Florida, Tortugas, Barbadoes, Jamaica, and
Hayti (Sluiter, 1910). The inference seems fair that it
represents a recent arrivalin Bermudan waters, since previous
extensive and detailed collecting has failed to reveal its
presence, and since only a single specimen has been found.
Concerning the method whereby, on this assumption, it came
to Bermuda, one can only speculate. It may have been
transported either (as a larva) by ocean currents or (as an
adult) upon the bottom of a vessel. The latter seems the
less probable.
References.
Britton, N. L. 1912. “Botanical Exploration in Bermuda,” Journ.
N.Y. Bot. Gard. vol, xiii. pp. 189-194, 5 pl.
Crark, H. L. 1901. ‘ Bermudan Echinoderms.—A Report on Obser-
vations and Collections made in 1899,” Proc. Bost. Soc. Nat. Hist.
vol. xxix. pp. 339-545,
——. 1907. “The Apodons Holothurians: a Monograph of the
Synaptide and Molpadide,’ Smithsonian Contrib. Knowl.
vol. xxxv. 231 pp., 18 pl.
Crozigr, W. J. 1915, ‘“ The Sensory Reactions of Holothuria surimam-
ensis, Ludwig,” Zool. Jahrb., Abt. Physiol. Bd. xxxv. pp. 233-297.
Piuspry, H. A. 1900. “The Air-breathing Mollusks of the Ber-
mudas,” Trans. Conn, Acad. Arts & Sci. vol. x. pp. 491-509,
1. 62.
Suurrer, ©. P. 1910. “ Westindische Holothurien,” Zool. Jahrb.,
Suppl. 11, pp. 831-342.
* IT am indebted to Dr. H. L. Clark for a suggestion regarding the
probable identity of the specimen.
On a new Lizard and Two new Frogs. 407
XXXVII.— Descriptions of a new Lizard and Two new Frogs
discovered in West Africa by Dr. H. G. F. Spurrell. By
G. A. Boutencer, F.R.S.
(Published by permission of the Trustees of the British Museum.)
Lygosoma spurrelli.
Section Hmoa. Habit lacertiform ; the distance between
the end of the snout and the fore limb is contained once and
two-fifths in the distance between axilla and groin. Snout
short, obtuse. Lower eyelid with an undivided transparent
disk. Supranasals very small; frontonasal broader than
long, forming a very broad suture with the rostral and
narrowly in contact with the frontal, which is as long as the
frontoparietals; frontoparietals and interparietal distinct,
latter smaller; four supraoculars ; seven superciliaries ; sixth
upper labial largest and below the eye. LHar-opening mode-
‘ rately large, vertically oval, without lobules anteriorly. 30
smooth scales round the middle of the body, ventrals largest.
Marginal preanals not enlarged. The hind limb does not
reach the axilla. Digits rather short, feebly compressed ;
subdigital lamelle obtusely keeled, 13 under the fourth toe.
Tail once and a half the length of head and body, gradually
tapering from the base. Dark reddish brown above, nape,
back, and limbs with numerous small black and yellow spots ;
a blackish lateral streak from the nostril, through the eye, to
the base of the tail ; sides pale brown, without spots; lower
parts yellowish white.
mm,
HHO CHOUbL COVEN Ut) aerate arcaeushuve uedeyer 48
s * HORG UTD 62 onsen ol cai 17
IE CAGR AMS KBr t Pe Lyte ee sant ere 10
Widthotgheadue Wrais 8. dee eo 8
Hore ‘limb tec ts et h aldn aie on Poe tao 13
ELindblintily® toaae baat tai 18
OO betas o eer eens hetah heels taste eae 8
PRE a tits AN pr aes. iat aly dn Gel wait #et 74
A single specimen from Obuasi, 8. Ashantee.
Closely allied to LZ. breviceps, Peters*, but with fewer
scales round the body.
Rana leonensis.
Vomerine teeth in very short oblique series close to the
anterior corners of the choanze. Head much longer than
* Cf. Boulenger, Ann, Mus. Genova, (38) ii, 1906, p. 206.
408 On a new Lizard and Two new Frogs.
broad ; snout pointed, strongly projecting, once and a half
the length of the eye ; canthus rostralis distinct; loreal region
feebly oblique, feebly concave ; nostril nearer the end of the
snout than the eye; interorbital space a little broader than
the upper eyelid; tympanum very distinct, two-thirds the
diameter of the eye and close to it. Fingers obtuse, first,
second, and fourth equal ; subarticular tubercles moderately
large, moderately prominent. Hind hmb very long, the
tibio-tarsal articulation reaching far beyond the tip of the
snout ; tibia once and two-fifths in length from snout to
vent, longer than the fore limb or the foot. Toes obtuse,
broadly webbed, the web reaching the tips of all except the
fourth, of which the two distal phalanges are free; sub-
articular tubercles moderately large, moderately prominent ;
inner metatarsal tubercle elliptical, half the length of the
inner toe; no outer tubercle; no tarsal fold. Skin smooth
above, with six fine, feebly prominent, interrupted glandular
folds along the back; a stronger fold above the temporal
region ; lower parts smooth. Grey above; a whitish line
from the tip of the snout, along the canthus rostralis, to the
eye, continued on the temporal fold ; side of head, including
the tympanum, dark brown, black above, this dark band
continued on the side of the body; limbs with numerous
dark cross-bandss; hinder side of thighs black, variegated
with white ; throatand belly yellow; lower surface of thighs
and of inner side of legs bright pink; plantar surface dark
brown, the web variegated with white. Male with blackish
external vocal sacs, each protruding through a slit measuring
one-third the length of the mandibular ramus and terminating
close to the middle of the base of the arm.
From snout to vent 43 mm.
A single male specimen from Bibianaka, Sierra Leone.
Distinguished trom &. equiplicata, Werner, by the longer
snout, with less oblique and feebly concave loreal region, and
the longer fourth finger ; from R. longiceps, Peters, by the
web not extending to the tip of the fourth toe and the
shorter slit for the vocal sac ; from R. oayrhynchus, Sundev.,
by the well-marked canthus rostralis.
Rappia spurrelli.
Head a little longer than broad ; snout pointed, as long as
the orbit, projecting strongly beyond the mouth ; canthus
rostralis rounded ; loreal region nearly vertical and feebly
concave ; nostril a little nearer the tip of the snout than the
eye; interorbital region a little broader than the upper
eyelid; tympanum hidden. Fingers moderately elongate,
On new Species of Indo-Malayan Lepidoptera. 409
free ; disks rather large. Tibio-tarsal articulation reaching
the eye; tibia 21 times in length of head and body, con-
siderably longer than the foot ; toes two-thirds webbed, two
phalanges of fourth free; subarticular tubercles small and
feebly prominent. Skin smooth above, granular on the belly
and under the thighs. Reddish brown above, with four
blackish longitudinal streaks; a blackish streak from the
nostril to the eye and a dark brown temporal band ; lower
parts white. Male with a large external subgular vocal sac,
covered by a large round flat disk.
From snout to vent 28 mm.
A single specimen from Obuasi, 8. Ashantee.
Closely allied to R. oxyrhynchus, Blgr., from the Katanga,
but distinguished by the total absence of web between the
fingers and the presence of the gular disk, as well as by the
coloration.
XXXVITI.— New Species of Indo-Malayan Lepidoptera.
By Colonel C. Swinwos, M.A., F.L.S., &e.
Family Lycenide.
Arhopala dascia, nov.
?. Upperside much as in A. ganesa, Moore, but darker
in colour. Underside with the ground-colour greyish white,
markings chocolate-brown; fore wing with a rather broad
medial band from the costa to vein 2, the immediate base of
the wing chocolate-brown, this colour running up the costa
for a short distance, the space between the base and the
median band filled up with fine transverse bands close to
each other, a discal macular band of square spots which
become somewhat diffuse at the hinder angle, the third spot
from the costa placed outwards ; the marginal space dark ;
a double row of lunular marks near the margin: hind wing
with the whole space, with the exception of a large round
patch below the middle of the costa, covered with round spots
(white-edged) on a dark ground, very difficult to describe,
but exactly similar to Watson’s figure, plate A, fig. 6, 2,
Bo. N. H. Soe. x., of a form he likens to A. ganesa, remarking
that it deserves a distinctive name, but Watson’s figure,
like A. ganesa, has no tails, and dascia has tails as in
A. aberrans, Doherty. I cannot but think that Watson’s
figure represents a sp-cimen of dascva with the tails broken
off.
Expanse of wings 1,%, inch.
Hab. Toungoo, Burma.
Ann. & Mag. N. Hist. Ser, 8. Vol. xix. 27
410 Colonel C. Swinhoe on new
Family Hesperiide.
Hasora philetas.
Ismene philetas, Plitz, Stett. ent. Zeit. xly. p. 55 (1884), unpublished
plate no. 1159.
Hasora simplicissima, Swinhoe (part.), Trans. Ent. Soc. 1908, p. 34.
Hasora mixta, Fruhstorfer (part.), Iris, 1911, p. 68.
Flab. Philippines.
Fruhstorfer puts philetas as a synonym to mixta, Mab., but
mixta like simplicissima is a Parata, whereas philetas (which
Semper wrongly identified in Rhop. Phil. ),is a Hasora, without
the eae on the fore wing, though the markings are very
similar; I have philetas male from the Philippines which
exactly corresponds with Plotz’s figure.
Notocrypta tobrianda, nov.
&S 2. Somewhat similar to N. alwensis, Swinhoe, Ann. &
Mag. N. H. (7) xx. p. 434 (1907), from the Solomon Islands,
figured in Trans. Ent. Soc. 1908, pl. iii. fig. 11, and
N. wokana, Plétz (from Aru Isl.), Berl. ent. Zeit. xxix.
225 Gee a copy of Plétz’s figure given in the same
plate (fig. 9), but in neither sex are there any dots or any
other ae above or below, there being only the discal
white band of the fore wing which is very much narrower
than in either of the above-named species, is much more
curved, narrows hindwards in the male into a square spot, is
similar in the female, but is of the same width throughout.
Expanse of wings, ¢ 17%, 2 1,8) inch.
Hab. Kiriwini Isl., Tobriand group.
Jetion elia-eburus.
Hesperia eburus, Plotz, Berl. ent, Zeit. xxix. p. 226 (1885), un-=
published plate no. 1373.
Eetion elia-ayankara, Fruhstorfer, Iris, 1911, p. 19.
Hab. Malacca, Perak.
A somewhat vitiable form, though the type form elia, Hew.,
from Sumatra, is fairly constant.
Telicota bambuse kiriwinia, nov.
g. Resembles 7. pythias, Mabille, Pet. Nouve, ti. p. 374
(1878), from Java, Sumatra, and Nias; the markings are
very similar, but the colour is paler ; the fore wing is longer
and the hind wing produced hindwards.
9. Very different from the females of pythias. Upper-
side: fore wing nearly all black; an orange streak on the
basal half of the costa ; a small streak below its outer end ;
a streak on the median vein from near the base to the origin
Species of Indo-Malayan Lepidoptera. 411
of vein 2; a longer streak on the internal vein and a shorter
streak from near the base close along the hinder margin of
the wing; two orange spots in the’ disc, and two small spots
subapical: hind wing also nearly all black ; astreak of orange
hairs in the cell, one along the internal vein, and a discal
narrow band of four orange spots. Underside: both wings
black, marks orange: fore wing with a broad streak close to
the costa; from the base to beyond the cell-end, joined to a
spot at the upper end of the cell, three discal spots, the upper
one pushed outwards; a small dot again outwards, and three
subapical: hind wing with a spot at the end of the cell and
a discal band as on the upperside; antenne longer than.
usual in the group.
Expanse of wings, g 13, 2 1,4, inch.
Hab. Kiriwini Isl., Tobriand group.
Family Deilemeride.
Section I.
Veins 6 and 7 of hind wing not stalked, palpi short,
Deilemera paradelpha, nov.
3 %. Belongs to the pellex group: Section I. of my
“© Monograph of the Deilemeride”” *, Frons white ; head and
collar luteous ; thorax white; a black spot on the head, two
on the collar, and the thorax with three black spots down the
middle and three on each side; palpi luteous, last joint
black; abdomen dull greyish yellow, with pale brown seg-
mental bands: fore wing pale black, with a lilac tinge; spots
white, a small round spot in the middle of the cell in the
male, developed intoa short streak in the female ; a large
oval spot across the end of the cell, from vein 11 to vein 2;
three submarginal spots as in pellex ; a white streak on the
hinder margin from base to a little beyond the middle:
hind wing white; a pale black uniform marginal band,
running narrowly up the abdominal margin for a short
distance, and containing a white subapical spot and a medial
spot. Underside: wings as on the upperside ; legs ochreous ;
body white, without markings. ~
Expanse of wings, ¢ 2, 174 inch.
Hab. Fergussen Isl.
Deilemera bouruana, nov.
9. Allied to D. separata, Walker, xxxi. p. 204 (1864),
from Gilolo Island. Pectus white; palpi white, last joint
black; frons, head, and thorax white, collar tinged with
* Trans. Ent. Soc. 1908, pp. 53-84,
27%
412 Colonel ©. Swinhoe on new
ochreous ; frons with a black spot, a large one on the head,
yunning in between the antennze, two on each side of the
collar; a black streak down the middle of the thorax and one
on each side ; abdomen ochreous grey, with black macular
segmental bands, marked with white: fore wing black,
markings white; a thick streak in the cell, from the base to
its middle, another similar but thicker streak immediately
below it and extending a little beyond it; a streak on the
hinder margin, from the base to beyond the middle ; a large,
oval, discal, oblique spot from close to the costa, where it is
round and small to vein 2, nearer the hinder angle of the
wing than is usual in the pellex group, in one specimen it
runs below vein 2 and is attenuated ; three larger submarginal
spots. Underside: wings as above; body white with thin
segmental bands on the abdomen; legs white striped with
black.
Expanse of wings 1,6 inch.
Hab. Bouru Is]., South Moluccas.
Deilemera externa, nov.
3 3. Face white ; palpi black; frons, head, and thorax
ochreous ; a black spot on the frons, a large one on top of
head, covering nearly the whole space and running in between
the antenne, two on the collar, an angular spot behind them
followed by an oval black spot on each shoulder ; abdomen
ereyish ochreous, some white specks at the base and thin
darker segmental lines touched with white at the sides and a
row of black spots on each side: fore wing dark black,
markings white ; a disjointed thin streak from the base to
the middle of the cell; a small spot below it and a little
outside; an oval spot in the dise from vein 11 toa little
below vein 2, narrow in the type-specimen, broader in others ;
three submarginal spots, the two lower ones connected to-
gether in the type-specimen ; a white streak on the hinder
margin: hind wing with a rather broad and even marginal
black band running thinly up the abdominal margin to the
base, and narrowly along the costa, with a knob at its middle ;
two white spots in the band, and indications of a third spot
as in Aurivillius’s figure of pellec*, but not so distinct.
Underside: wings as above; legs and thorax ochreous;
abdomen white with black segmental bands.
Expanse of wings, ¢ 1,8, 2 -1y% inch.
Hab. Batjan Isl., Middle Moluccas.
* Rec, Crit. in Sy, Ak. Handl. xix. (5) pl. i. fig. 5 (1882).
Species of Indo-Malayan Lepidoptera. 413
Section I. B,
Antenne shortly pectinated.
Deilemera gonora, nov.
3. Nearest to D. oroya, Swinhoe, from Sula Besi. Palpi
white, last joint black; frons, head, and thorax white; frons
with a black spot, collar with two ; two square black spots
followed by around one on the thorax; abdomen white with
hardly any markings, tip ochreous ; wings white, markings
black: fore wing with the veins black ; a small patch on
the middle of the costa, another on the costa above the cell-
end and running into the bar at the end of the cell; a broad
black band on the outer margin containing three white sub-
apical spots close together, the middle one the largest, the
lower very small, followed by two spots against the black
patch above the cell-end, with two white linear marks below
them, and large white spot at the middle of the outer margin
with a white streak attached to its lower end: hind wing
with a fairly broad black outer marginal band, not continued
on the costa, its inner margin sinuous and two white spots in
it, apical and medial. Underside: wings as above; body
white without markings ; legs ochreous.
Expanse of wings 2 inches.
Hab. Stephensort, Dutch New Guinea.
Deilemera similis, nov.
3 2. Nearest to D. burica, Holland, from Bouru, of
which I have a fine series from Bouru; palpi white, last
joint black ; frons, head, and thorax pale ochreous, nearly
white ; frons with a black linear spot, another similar spot
on the head; thorax with two black spots on each side ;
abdomen pure white, with small black dorsal and lateral
spots: fore wing black, markings white; a broad spear-
shaped streak from the base, from the median vein to the
internal vein, containing a black dot atits base, two short sub-
costal streaks above it and two round spots outside it ; a broad
oblique band divided by the veins into five large spots, the
lowest round, the others elongate, the fourth from the costa
the largest, and extended both outwards and inwards with a
little streak hindwards from its interior end, the third wedged
in outwards between the second and the fourth, sometimes
with a very small spot attached to its upper side; a little
beyond the band is a subcostal spot with another below it;
a large double spot subapical and another near the middle of
the outer margin: hind wing with a rather broad black band
narrowing hindwards, and continued in almost a line on two-
thirds of the costa; a white spot on the costa below the
A414 Colonel C. Swinhoe on new
apex, a smaller spot just inside it, and a larger spot near the
margin below the middle. Underside: wings as above ;
body and legs white, tarsi black at the ends.
Expanse of wings, ¢ 2, 1;% to 2 inches.
Hab. Stephensort, Dutch New Guinea ; a fine series.
Section II. D.
Veins 6 and 7 of hind wing stalked, palpi longer ; male without
secondary sexual characters.
Deilemera cenis parva, nov.
9. Very similar to D. cenis, Cramer, from Sikhim, Silhet,
and Assam, but very much smaller, all the spots and bands
about half the size.
Expanse of wings 1,% inch.
Hab. Au-San, Central Formosa. |
‘T'wo females.
Deilemera poliodesma, nov. «
3 2. Head, collar, body, and palpi luteous, last joint
black ; a black spot on the frons, another on the top of the
head, two on the collar, a thick black stripe along the middle
of the thorax, and a narrower black stripe on each side;
abdomen with black segmental bands above, and black spots
on the segments beneath: fore wing black ; a broad discal
white band divided by the veins into six spots, much as in
D. baulus, Boisd., from Bouru, but narrower, the uppermost
spot small, the veins of the wing from the base to this
band dull yellowish: hind wing pure white, a broad marginal
black band, angled inwards on vein 2, narrowing on the
costa, and running upwards for ashort space on the abdominal
margin. Underside much as above, but paler, the inner
portion of the fore wing suffused with dull ochreous.
Expanse of wings, ¢ 2, 1; inch.
Hab. New Britain (New Mecklenburg).
Belongs to the daulus group. Ihave a great number of
examples of this group from the different islands, and,
though more or less resembling each other, each island
seems to contain a form of its own.
Deilemera homogona, nov.
g. Allied to D. lacticinia, Cramer ; the abdomen is with-
out the black segmental bands on the upperside, having
merely a row of black dots, decreasing in size hindwards;
the fore wing has the discal white band twice the width,
more compact, and on the upperside of the small outer white
spot (the third from the costa) there is another white spot
Species of Indo-Malayan Lepidoptera. ALS
‘attached ; the marginal black band of the hind wing is
much narrower and decreases in width hindwards.
Expanse of wings 15 inch.
Hab. Hué, Annam.
Deilemera perissa, nov.
3 2. Allied to D. luctwosa *, Vollenhoven, from Batjan
Island ; palpi, head, and body ochreous, last joint of the
palpi black; frons with a black spot in its centre, one on the
head, two on the collar ; three longitudinal stripes on the
throat, a spot at the base of the abdomen followed by seg-
mental black bands: fore wing black, a spear-shaped white
band from near the base, widening outwards to the base of
vein 2, where it is cut short; a short white streak from the
base along the costa, another on the median vein to the base
of vein 2, and a similar streak on the internal vein ; a fairly
broad discal white band of seven white spots joined together,
the fourth round and small, between the ends of the third
and fifth, the lowest still smaller and oval; in one example,
the band from the base is connected with the discal band by
a thin white streak and there is a white spot at the end of
the cell: hind wing white, with the usual black marginal
band, much as in ductuosa. Underside as on the upperside;
body and legs ochreous, the former spotted with black ; the
legs with black stripes.
Kixpanse of wings, gd ¢, 14 inch.
Hab. Obi Isl., Moluccas.
A fine series.
Deilemera delocyma, nov.
3S. The antenne long, the pectinations also somewhat
longer than is usual in the group ; pectus, frons, head, and
thorax dull orange-ochreous ; frons with a black spot, one on
the neck, two on the collar; thorax with black stripes on the
middle and on the sides ; abdomen dull orange-ochreous, with
black segmental bands, so broad as almost to obliterate the
ochreous colour: fore wing long; costa much arched before
‘the apex; colour black, a white stripe below the médian
vein from the base to the middle; a broad and somewhat
upright discal band divided by the veins into seven spots,
extending from near the costa to near the hinder margin, the
first three from the costa narrow, the fourth oval, wedged in
between the onter halves of the third and fifth, the sixth the
largest, the seventh small and round-edged: hind wing
white, a broad black marginal border, angled at vein 2,
and continued narrowly along the costa. Underside: body
* Tidj. von Dierk, i. p. 42 (1863),
416 On new Species of Indo-Malayan Lepidoptera.
orange-ochreous; abdomen with black segmental bands;
legs orange-cchreous striped with black.
Expanse of wings 2 inches.
Hab. Flores Isl.
Family Orthostixide.
Alex niasica, nov.
@. Frons and palpi black, the inner side of the first two
joints of the palpi white ; ground-colour of the head, body,
and wings greyish ochreous ; ; head and collar blackish
through thickly covered irrorations ; both wings uniformly
covered with short brown striations : fore wing with a pale
brown mark at the upper end of the cell, both wings with
indistinct traces of outwardly curved, pale brown, thin bands,
antemedial and discal; a thin, marginal, pale brown line and
ochreous cilia with grey tips. Underside as above, but the
striations and transverse lines thicker and more prominent.
Expanse of wings 2 inches.
Hab. Nias.
Quite distinctfromany other species of this genus known tome.
Family Geometridae.
Pingasa talagt, nov.
g. Frons, head, body, and wings greyish white, strongly
tinged with ochreous irtorations ; ; palpi with the third joint
black, elongate as in P. aravensis, Prout *; wings with the
markings pale grey: fore wing with a sinuous blackish
thick line closing the cell; a nearly straight short line
beyond it from the middle of interspace 3 to near the costa,
an indistinct outwardly curved subbasal line ; a more distinct
and thicker postdiscal line, more or less distinctly macular ;
a submarginal row of indistinct spots pricked with white on
their inner sides, and minute dots on the outer margin; hind
wing with a line closing the cell; the two outer marginal
bands diffuse and more or less conjoined together and with
the outer margin, the white inner marks on the very indistinct
submarginal ‘spots more prominent than they are on the
fore wing. Underside: wings paler and without irrorations ;
fore wing with a very large black spot at the end of the cell
and a large black upper discal patch attenuated hindwards ;
three white dots outside the upper part of the black patch ;
hind wing with a thick black lunular mark at the end of the
cell, and a large black upper discal patch as on the fore wing
but without the white dots.
Expanse of wings 1,‘ inch.
Hab, Tulagi, a small island off Ysabel Island in the
Solomen group Setar
* Noy. Zool. xxiii. p. 7 (1916).
On Cylindroiulus (Leucoiulus) nitidus (Verhoef’), 417
XXXIX.—Notes on Myriapoda.—V.* On Cylindroiulus
(Leucoiulus) nitidus (Verhoef). By Hitpa K. Braves,
M.Sc., L.R.C.P., M.R.C.S., and the Rev. S. GraHam
Birks, M.Sc.
Last year Dr. A. Randell Jackson recorded ¢ the finding by
one of ust at Winkhill, near Leek, Staffs, of Cylind?otulus
(Leucotulus) nitidus (Verhoeff). This seems to have been
the first record of the occurrence of this species in Britain,
and the following notes are based primarily upon our study
of material from the locality named in the light of the original
paper § in which Dr. C. W. Verhoeff describes Julus nitedus.
Of this description we have made the fullest use, and we here
express our indebtedness.
Our specimens were first examined by Dr. Jackson, and
then sent to Monsieur le Docteur Henry W. Brélemann of
Pau, who, with his usual kindness, dissected one of the males,
and identified it as the Julus nitidus described by Verhoeft.
1. OccURRENCE AND HABITAT.
Cylindroiulus nitidus was taken in September 1915 at
Winkhill, Staffs, by one of the writers (H. Kk. B.); three
males and two immature specimens were found together under
a stone on the grassy sides of the road leading from the
station to the village; no adult females were then taken.
In September 1916, when we had already made some study
of the limited material then at our disposal, we took many
specimens of both sexes in the same immediate neighbourhood.
In the field C. nitidus resembles C. stlvarum (Meinert) very
closely except in habitat. Verhoeff states (doc. cét.) that he
had noticed this animal only in woods or on heights. He
records it in the Rhine and Moselle districts: at Melbthal,
near Bonn, in a wood under leaves ; at Ippendorter Héhe,
near Bonn, at the edge of a wood under stones; at Cochem,
in woods and at Siebengebirge, Petersberg, in woods under
stones. ‘There are some trees in the neighbourhood of the
ground where the English specimens were collected at
Winkhill; but the very limited area of its occurrence is not
in close connection with woods, the district being hilly and
* The authors’ previous notes I.-IV. in this series appeared as follows :—
I., Lancs. & Ches. Nat., June 1916; IL, cid. July 1916; IIL, ‘ Irish
Naturalist,’ August 1916; IV., Lancs. & Ches. Nat., September 1916.
7 ‘On some Arthropods observed in 1915,” Lanes. & Ches. Nat., Feb.
1916, p. 391. .
{BK Be
§ “Kin Beitrag zur mitteleuropaischen Diplopoden-Fauna,” Berliner
Entomolog. Zeitschrift, xxxyi. 1, 18V1, pp. 116 et seg.
418 Miss H. K. Brade and the Rev. S. G. Birks on
rather bleak. Some little distance away, however, there are
several well-wooded areas.
The rocks of the neighbourhood are Carboniferous sand-
stones and shales. There is Carboniferous Limestone not
more than 3 miles distant in a southerly direction.
2 2. SYSTEMATIC POSITION.
This species takes its place in the family Julide, being
referred to the genus Cylindroiulus (Julus s, |.) and subgenus
Leucoiulus.
3. Some EXTERNAL CHARACTERS.
(a) Dimensions.
Dr. Verhoeff (loc. cit.) gives the following dimensions :—
Male: length 15-20 mm. ; breadth 1°0-1°3 mm.
Female: length 18-24 mm., less often up to 29 mm. (as
in the case of one example 2 mm. broad) ; breadth 1°3-2°0 mm.
‘The dimensions of three males taken at Winkhill in 1915
are :—
Length 14-20 mm.; breadth 1°0-1°3 mm.
And of three females taken at the same place in 1916 :—
Length 22-24 mm.; breadth 1°6-1'8 mm.
(b) Colour-markings ete.
The exoskeleton is brightly polished. Hach pleurotergite
of the trunk is composed, as usual, of a narrow anterior pro-
zonite (fig. 2, p) anda broader posterior metazonite (fig. 2, m).
In our specimens the general tone of colour is brownish,
but the shade varies very much in different specimens from
quite dark to fairly light; above the level of the foramina
the pleurotergites are relatively dark, and dorsally the seg-
ments are distinctly mottled by patches of a greyish tone ;
passing ventrally the somites get much lighter, and so the
mottled appearance is practically absorbed by the generai
tone of colour. At the level of the foramina a row of distinct
brown spots may be seen running along each side of the
body. These spots occur on each of the metazonites except
those of the first few and last few segments,
Frons and vertex without grooves.
The prozonites are smooth.
The metazonites are marked with longitudinal grooves set
widely apart, but these grooves are wanting on the dorsal
portions of the most anterior segments (fig. 1).
The ventral plates (sternites) are simple and insignificant.
Cylindroiulus (Leucoiulus) nitidus (Verhoef), 419
Fig. 1.
Fig. 1.—C. nitidus, anterior end of body of 3. Details of eye not shown.
On the collum segment the characteristic curved groove is
clearly seen. 3 is the third body segment. x 86. H.K. B.
del.
Fig. 2.—Posterior end of same. f, foramina repugnatoria ; p, prozonite ;
m, metazonite. x 36. H.K. B. del.
420 Miss H. K. Brade and the Rev. S. G. Birks on
Above the mouth four bristle-bearing dimples are present.
Crossing the head from side to side is a deeply pigmented
band on which the eyes are situated laterally (fig. 1).
The collum segment is large and a longitudinal furrow
runs along each of its lateral edges, which are slightly
curved. Above this longitudinal furrow and parallel to it
near the posterior border on each side two very much shorter
grooves sometimes occur. These shorter furrows appear to
correspond to the longitudinal grooves of other segments.
In Julus nitidus the foramina repugnatoria are small and
situated on the posterior border of the metazonite, and so lie
in close proximity to the sutures between the segments
(fig. 2, f).
The tergite of the anal segment is produced into a long
caudal horn which tapers gradually and is bluntly pointed at
the tip ; it is round in transverse section.
(c) Appendages.
We take.the three males collected in 1915 as examples :—
A ¢ of 20 mm. had 56 segments, 101 pairs of legs, and 2 legless terminal
seoments.
Aodgofl4 do. 46 do. ris do. 4 do.
Acgofl4 do. 48 do. 85 do. 2 do.
The number of segments and, therefore, the number of
pairs of legs vary considerably according to the maturity
of the specimens.
The appendages of the males collected in 1915 are arranged
as follows :—
Segments. Pairs of limbs.
Lok teen ° 1st One pair, modified, hook-shaped.
Avis a. Rain ntaled rae 2nd One pair, with pads on the 4th and
5th joints and a secretory organ
on the coxite.
Sirs w strides sia ek ie Genital apertures,
eaters thsi ae 3rd One pair.
eachemrere oe ) a Two pairs.
Gis 8 atis oe eee oe Two pairs.
7 8th Anterior gonopods.
Llano ah 9th Posterior gonopods,
10th, 11th,
8 and onward . aE
Last 2or4 .. Juegless.
Two pairs,
In the female the first and second pairs of legs are not
modified, and in place of the gouopods are two pairs of
walking-legs ; otherwise the arrangement is the same.-
Cylindroiulus (Leucoiulus) nitidus (Verhoef). 421
(d) Ocell.
Dr. Verhoeff dwells upon the great interest of the ocelli in
this species, and states that this character provides an inter-
mediate condition between Ummatoiulus and Aliaiulus. The
external surface of the field of the eye is not broken up by
the convexities of the individual ocelli, and with some illumi-
nations the microscope reveals the fact that the field is quite
smooth. With some illuminations it is impossible to see the
individual ocelli, with others they are quite distinctly visible.
Ocelli arrangement (1915 material) :—
Ina ¢ of 14 mm.: 4, 5, 6, 5, 3, 2 (25). In curved rows, somewhat
irregular.
Ina ¢ of 20 mm.: 3, 5, 6, 6, 4, 4, 2,1 (81). Irregular.
Ina ¢ of 14 mm.: 3, 4, 6, 5, 4, 1 (23). Irregular.
In an immature specimen: 5, 4, 3, 2,1 (15), Fairly regular.
do. do.» 4,5; 2, FCO). 0.
It will be seen that there is great variation in the number
of individual ocelli. This is also noticed in Dr. Verhoeft’s
description, where he records from 26-40 ocelli in different
specimens. As in the case of the walking-legs, the greater
number appear to occur in the more mature specimens,
Fig. 4.
Fig. 3.—First leg of male of C; nitidus. x 140. H. K. B. del.
Fig. 4.—Second leg of male, with segments numbered. s, secretory
organ (penis). x 140, H.K. B. del.
3. MopIFIep APPENDAGES OF THE MALE.
The first pair of legs are hook-shaped, forming an angle
at the outer bend as the tip turns inwards, and a small
prominence is present at this angle (fig. 3). The second pair
422 Miss H. K. Brade and the Rey. S.G. Birks on
of legs are also slightly modified, the joints being thicker
than those of the normal walking-legs, and pads are present
on the fourth and fifth joints—that is, the penultimate and
antepenultimate Joints (fig. 4). The claws are strong, and
the last joints are armed with strong bristle-like spines,
some of which overlap the claws. The coxite bears a
secretory organ (fig. 4, s). The copulatory feet or gonopods
are composed of the modified appendages (two pairs) of the
seventh body-segment. Their situation is denoted by a gap,
as these legs are retracted, in this group, within the somite to
Fig. 5.
Figs. 5-8.—Gonopods. A, anterior gonopod; B, posterior portion of
posterior gonopod; C, anterior branch of posterior gonopod ;
a, b,c, parts of CO; F, flagellum; d, hooked part of B; L-L,
point of measurement (see text).
Fig. 5.—Silhouette of anterior gonopods, anterior surface. x 140.
H. K. B. del.
Fig. 6.—Silhouette of posterior gonopods, posterior surface. xX 140.
H. K. B. del.
which they belong. They are complicated in character, and
the anterior and posterior gonopods of each side are firmly
attached to one another. The anterior pair of gonopods are *
seen from an anterior position as two lobe-like projections
rounded at the tip (fig. 5). From a posterior position the
posteriot pair appear as two pointed projections with the tips
turned inwards, broader at the base than at the tips (fig. 6).
Viewed from the side the external surfaces of the anterior and
posterior gonopods are seen (fig. 7). They are clearly sepa-
rated at the free ends and united at the bases. The anterior
gonopods are stronger and generally less transparent than the
posterior pair, which is delicate and very transparent, some
parts being irregular in outline.
Cylindroiulus (Leucoiulus) nitidus (Verhoef). 423
The anterior gonopods are leaf-like in form, and, being
convex posteriorly, their lateral edges wrap round and protect
the anterior branch of the posterior gonopods somewhat
(ig. *).
The posterior gonopods consist of a foliaceous anterior
branch and a subtrapezoidal posterior portion.
The anterior branch consists of three parts—a, b, and c—
which are very distinct and characteristic; @ is finger-shaped
and is the longest, standing out well beyond the others ; the
middle portion, 8, is the strongest, and is protected by very
Fig. 7.
~S
t
‘
‘
‘
‘
A
\
:
Anterior and posterior gonopods, external profile. x 140. H.K. B. del,
thick chitin; while ¢ is smaller and approaches in shape
more nearly to a.
The structure of the posterior gonopod is clearly seen in
fig. 8, which represents the inner view of the gonopods both
anterior and posterior of one side. In this case the appendages
have been flattened out somewhat to show the separate parts
of each. The most prominent part of the posterior portion of
the posterior gonopod is the hooked piece, d, which is sparsely
plumous at the tip, while a noticeable feature is the flagellum,
F, This, which is inserted upon the base of the anterior
424 On Cylindroiulus (Leucoiulus) nitidus (Verhoef’).
gonopod, curves round so that its tip appears in repose above
the delicate part of the posterior gonopod behind the hooked
portion d, ‘The actual breadth of the posterior gonopods at
L-L in the specimen figured is 0'210 mm.*, and that of the
whole organ (anterior and posterior gonopods of one side
together) is*4 mm,
Fig. 8.
q &
é F
He
fi at ree.
‘ Fares Le ae
‘ Pe ee | ae
cig
‘
<5)
octes
o 7.
Anterior and posterior gonopods, internal profile. x 140, H.K.B. del.
We are deeply indebted to Monsieur le Docteur Henry W..
Brélemann for his very careful examination and report upon
our material. Drawings of the gonopods which he furnished
have been invaluable in the preparation of the illustrations
which accompany this paper.
We must also express our best thanks to Miss Simpson, of
Darwen, for her assistance in translation.
The Victoria University of Manchester.
* Dr. Brélemann’s measurement,
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No. 114. JUNE 1917.
XL.—Notes on Collembola.—Part 4. The Classification of the
Collembola; with a List of Genera known to occur in the
British Isles. By Joun W. Suorsornam, N.D.A., Berk-
hampsted, Herts. ;
In my paper on “Some Irish Collembola” (1914), p. 59,
I remarked that the classification adopted was one which,
in the main, had been accepted by authors for the previous
seven years, and that Dr. Borner had recently proposed a
new system on which I should make some notes.
This paper was practically completed during the early
part of 1914, but my unexpected call to South Africa
prevented me publishing it, and I then intended waiting till
I returned to England. However, as I am about to publish
a preliminary account of the Collembola of Lancashire and
Cheshire, I feel it is just as well to list them according to
the new system, which is much to be preferred to any yet
proposed. I therefore give here aetranslation of part of
Borner’s paper (1913 4), pp. 318-322, which forms a key to
the new classification, and I append a list of the genera of
Collembola hitherto found in the British Isles.
My best thanks are due to Dr. A. D. Imms, of the
University of Manchester, for kindly seeing this paper
through the press and for criticisms and suggestions, and to
Mr. T. A. Coward, also of Manchester University, for much
kind assistance.
Ann, & Mag. N. Hist. Ser. 8. Vol, xix. 28
426 Mr. J. W. Shoebotham’s Notes on Collembola.
There have been many different arrangements of the
Collembola made during the last 75 years, and the number
of families recognized has varied from 3 to as many as 8.
Nicolet (1842) has the springtails divided into the
Smynthurelles,’ Podurelles, and Lipurelles, and Lubbock
(1862) called these Smynthuride, Poduride, and Lipuride,
while in his monograph (1873) he formed 6 families, viz.,
Smynthuride, Papiriide, Degeeriade, Poduride, Lipuride,
and Anouride. Various modifications were used by authors
till the end of the 19th century, when another family— |
Neelide—was made for the reception of the genus Neelus of
Folsom. Schiffer (1896), in his paper on “ The Collembola
from the Neighbourhood of Hamburg,” differentiated the
subfamilies Isotomine and Tomocerine, which now rank as
separate families.
During, the present century the work of Borner has done
much te advance our knowledge of the classification of the
order Collembola. In one of his earliest papers (1901 a)
he divided the Collembola into two suborders, the linear
kinds to be grouped under the name Arthropleona and the
globular forms he called Symphypleona. Keys were given
to the families and subfamilies of the Arthropleona, and
these, together with an account of the Symphypleona, were
given in more detail in his paper on “The Apterygotal
Fauna of Bremen” (1901 4). Then, in 1906, in his work
on “The Classification of the Collembola,’’ Borner discussed
the whole group and the relationship of the families, sub-
families, and tribes. He recognized the families Poduride,
Entomobryide, Neelide, and Sminthuride, and gave a synopsis
of the subfamilies and tribes. This system, with but little
variation, was used by authors for many years. Then, in
1913, when examining some species of Pseudosira and
Paronella from Java, Borner happened to find a peculiar
structure on the hinder trochanters, in the form of a number
of short, outstanding, pointed bristles, to which he gave the
name ‘‘ Trochanteral organ.” On looking through his collec-
tion of slides, he found that this structure was present in all
the true Entomobryine, but absent in the Tomocerine and
Isotomine. This discovery led Borner to propose a new
arrangement of families, which I give in this paper. He
firstly divided the Arthropleona into two natural sections
according to the structure of the prothorax (see below, in
Key to Families, etc.). The old family Poduride, which
corresponds to the new section Poduromorpha, was divided
into three, the subfamilies Hypogastrurine and Onychiurine
being raised to the family rank, and the name Poduride
~
Mr. J. W. Shoebotham’s Notes on Collembola. 427
restricted for the single genus Podura. The second section
Entomobryomorpha, which corresponds to the old Entomo-
bryide, was also split into three; the name Entomobryide
was retained for those species possessing the Trochanteral
organ, and the rest divided into two new families, Isotomide
and Tomoceride. The Sminthuride and Neelide remained
as before.
This classification gives us 8 families, and it may seem
a large number for so small an order, but there are many
districts in the world that have never been searched for
springtails, and others in only a haphazard manner, and,
doubtless, when the group has been more thoroughly
worked, there will be hundreds of new species discovered,
which will result in the formation of new genera, and probably
of the larger divisions also.
As an example of how a tribe has increased in size and
importance in recent years, take the Cyphoderini. This
tribe for many years contained only the one genus Cypho-
derus, Nicolet (1842), and that genus, as we know it to-day,
contained only two or three species up to the end of the 19th
century. Now, as a result of collections made in various
parts of the world, there are the additional genera, Cypho-
derodes of Silvestri (1911), Pseudocyphoderus, Imms (1912),
the peculiar genus Calobatella described by Borner (1913 a).
The genus Cyphoderus now contains a dozen or more species,
with the probability of the number being increased in the
near future.
Synopsis of Suborders, Sections, Families, Subfamilies, and
Tribes of the Order Collembola, taken from Bérner
(1913 4), pp. 318-322.
A. Body flattish-cylindrical, elongated, as a rule distinctly segmented,
with free thoracic and free abdominal segments; rarely the
abdominal segments 5 and 6 or 4-6 are fused together.
Suborder AntHROPLEONA, C. B., 1901.
I, Tergum of the prothorax similar to the terga of the other body-
segments, always, as in the case of these, possessing some hairs.
Furcula present or absent, in the first case lying under abdominal
tergum 4. Integument generally granular, mostly soft, seldom
with stouter chitinized sclerites. Ventral tube always short,
pocket-like, smooth-walled. Manubrium ventrally always without
hairs. Section PopuromorpPnA, C. B,, 1913.
a. Without pseudocelli. With or without eyes. Sensory organ otf
the third antennal segment only with sense-rods, without
sense-cones, without outer papille. Fourth antennal segment
without subapical sense-pit, always with retractile sense-knob.
28%
4.28
Mr. J. W. Shoebotham’s Notes on Collembola.
1. Head hypognathous. Eyes situated near the hind edge of the
heid. Dentes bowed in the horizontal plane, annulated
towards the end, over-reaching the ventral tube. Manubrium
in form recalling that of the Symphypleona, with a special
medial support-piece of the dentes.
1. Family Popurip# (C. B.), 1906.
2. Head obliquely prognathous. With or without eyes, these, if
ii.
present, situated in front of the middle of the head. Dentes
not annulated, fairly straight, seldom reaching past the
veutral tube, or the furcula more or less completely reduced.
When the fureula is present the manubrium is simple in
form, without the medial support-piece of the dentes.
2. Family HypoGasTRuRIDs& (C. B.).
. Mandibles with well-developed molar plate.
Subfamily HypocastruRin», C. B., 1906.
Mandibles without true molar plate or absent.
Subfamily AcnorocTina, C. B., 1906.
* Anal segment with undivided supra-anal valve. With or
without furcula.
Tribe PsEuDACHORUTINI, C. B., 1906.
*# Anal sezement with double-lobed broader supra-anal valve.
Without furcula.
Tribe AcHorutTInNI, C. B., 1906.
b. With pseudocelli. Without eyes. The sense-organ on antenna
III. provided with from 2-3 sense-cones, and often at the same
time with outer papille and protecting bristles, in addition to
the sense-rods. Post-antennal organ generally well-developed.
Antennal segment 4 generally with subapical sense-pit
(= Lipuride, Lubbock, 1869 1.)
3. Family Onycururip# (Lbk., C. B.).
Il. Tergum of the prothorax always membranous and without hairs.
Fureula generally present, and in the more recent forms of the
group pushed analwards, Integument generally smooth, mostly
with sclerites. Ventral tube short or elongated, sometimes with
a lateral blind sac. Manubrium ventrally, generally with hairs
or scales, seldom naked.
a.
Section ENromMopryoMorpHa, ©. 'B., 1913.
Trochanteral organ absent. Ventral edge of the claw simple,
without groove.
1. Third and fourth abdominal terga of nearly equal length or
the 4th longer, sometimes this (the 4th) fused with the 5th
and 6th (without scales*). Naked or ciliated sensory setz
(=bothriotriche) present or absent.
4, Family lsoromip& (Schffr., C. B., 1896-1903.)
1 This should be 1870. Lubbock’s “Notes on the Thysanura—
Part 4,” was published in 1870, not 1869.—J. W. 8.
* This is as given by Borner (19136); but, to include the genus
Oncopodura, Carl & Lebed., it should now read “Scales nearly always
absent, when present they are without longitudinal ribs.”—J. W. S.
1914,
,
Mr. J. W. Shoebotham’s Notes on Collembola. 4.29
i. Head obliquely prognathous, Antenne inserted in the
front half of the head. (Trachex absent.) Furcal seg-
ment without chitin-ridges, to which are articulated the
basal sclerites of the furcula if these (the chitin-ridges)
are present. J’urcula seldom absent, Post-antennal organ
generally present.
Subfamily Isorommunm, Schffr., 1896.
(Including the Isoromuriy, C. B., 1906.)
ii. Head hypognathous. Antenne inserted in the middle of
the head. (Head trachez present.) Furcal segment with
two stout chitin-ridges, which serve for the articulation
of the hasal sclerite of the furcula.
Subfamily AcraLeTin~, C. B., 1906.
2. Third abdominal tergum considerably longer than the fourth,
ull abdominal segments free. With longitudinally ribbed
scales, Post-antennal organ absent, Sensory sete present,
ciliated, Furcula always present.
5. Family Tomocrerip2 (Schfir., 1896),
i. Dentes annulated; mucro diminutive, without hairs, An-
tennal segments 3 and 4 about equally long.
Subfamily LEprvoPHORELLINz (C. B., 1906.)
ii, Dentes not annulated, 2-segmented; mucro longish, with
hairs. Third antennal segment strikingly elongated (this
annulated as well as the fourth),
Subfamily Tomocrerin& (C. B., 1906).
b. Trochanteral organ present (on the trochanters of the hind legs),
Ventral edge of the clawsas a rule with basal groove (“split”).
Hairs and scales (at least in part) ciliated. Fourth abdominal
tergum as a rule considerably longer than the third. Ciliated
sensory sete always present. Furcula always present.
6. Family Enromosryip» (C. B., 1906).
1. Dentes slender, annulated dorsally here and there. Mucro
small. With or without scales. With or without eyes.
Kmpodial appendage with 4 wing-corners.
Subfamily Enromopryin (C. B., 1906).
i, Antenna 4-segmented, with undivided basal segments.
Tribe EnromosryinI, C. B., 1906.
ii. Antenna 5- or 6-segmented, with the first or first and second
segments secondarily divided into two.
Tribe OHCHESELLINI, C. B., 1906,
[Here also belongs, presumably, the Heteromuricine of
Imms, 1912. The medial cercus described by Imms
should be nothing else than a peculiarly elongated supra-
anal valve. |
2. Dentes not annulated, not, or only a little, tapered towards
the end.
i. Dentes without dorsal ciliated scales or ciliated spines,
dorsally and dorso-laterally (often completely all round)
uniformly haired ; if dental spines are present, then some-
times they are modified into bristles at the ends of the
dentes. Empodial appendage with 4 strong wing-corners.
430
Mr. J. W. Shoebotham’s Notes on Collembola.
Mucrones plump. Coxe strikingly short. With or
without scales. The hitherto known forms with eyes and
free-living. Subfamily ParonELiinZ (C. B., 1906).
ii. Dentes with dorsal ciliated scales or ciliated spines, these at
the bases of the dentes pass over into ciliated bristles.
Coxe generally distinctly longer than the trochanters (also
in Troglopedetes ?), Hmpodial appendage with 3 wing-
corners, or more or less reduced. yeless and scaled.
Subfamily CypHopERIN# (C. B., 1906).
* With one entodorsal row of ciliated spines on the dentes.
Free-living in caves.
Tribe TROGLOPEDETINI, C.B., 19138.
** With a double row of ciliated scales on the dentes. Man-
dibles with or without molar plate. Head obliquely pro-
gnathous, or hypognathous. Claws normal or with a
bladder-like terminal portion. Mostly living in association
with ants or termites.
Tribe CypHopERrinI, C. B., 1918.
B. Body pear-shaped to almost globular, as a rule, with obsolete seg-
mentation of the thorax and the first 4 abdominal segments;
especially the abdominal terga and pleure not differentiated. The
ano-genital segments remain mostly indepéndent; but are occasionally
fused together. Manubrium with a medial support-piece of the
dentes, similar to the Poduride, ventrally always without hairs.
Pronotum (sometimes also the metanotum) without hairs.
Suborder SympHypizona, C. B., 1901.
a. Antenne inserted in or in front of the middle of the head, always
considerably shorter than the head-diagonal, 4-segmented ; head
without elevated vertex. Corpus tenaculi without bristles.
Cove elongated, also on the outer side distinctly longer than the
trochanteral segment, Ano-genital segment concealed under the
furcal segment (seen from above). Abdominal sensory sete
absent. 7. Family NrErLipz, Folsom, 1896.
b. Antenne inserted behind (over) the middle of the head, generally
considerably longer than the head-diagonal, not rarely with
subdivided segments. Head with distinctly elevated vertex over
the neck. Corpus tenaculi (generally) with bristles. Core not
elongated, on the outer side considerably shorter than the inner
side and than the trochanteral segment. Ano-genital segment
not concealed under the furcal segment. Abdominal sensory
setae present. 8. Family SminraHuripm®, Lubbock, 1862.
. Ventral sac (tube) even in full-grown animals smooth-walled
(seldom at the tips with small warts—after Linnaniemi).
Corpus tenaculi with lateral appendage (stylus?) at the base
of the rami. Integument granular. Trachee (always?)
present. Subfamily SminvHuriinwz, C. B., 1906
* Anal and genital segment fused, on each side with two (one
small) sensory sete.
Tribe SmuintHuRIDINI, C. B., 1913.
** Anal and genital segment separated, sometimes the latter
fused with the furcal segment. The genital segment with
only one sensory seta.
Tribe Katrannint, C. B., 1913.
Mr. J. W. Shoebotham’s Notes on Collembola. 431
2. Ventral sac (tube) in full-grown animals with warted walls.
Antenne always distinctly bent.
i. Antenne bent between the 3rd and 4th segments, 4th segment
longer than the 38rd, the latter always undivided. Furcal
segment without paired dorsal papille. Corpus tenaculi at
the base of the rami without lateral appendage. ‘Tracheze
present (always?). Genital and anal segment not fused.
Subfamily SminrHurin»®, C. B., 1906.
* Tibio-tarsus at the distal end on the hind side with 2-3
(seldom with a 4th on the front side) more or less closely
applied clubbed hairs, not, or only little, over-reaching the
claws. With or without empodial appendage. Mucronal
bristle absent. Tribe BouRLETIELLINI, C, B., 1913.
[Ilere also belongs the genus Corynephoria, Absolon
(1907), which is very nearly related to Bour/etiella, and
which only differs by the absence of the empodium and
through the dorsal clavate appendage. It is doubtful also
whether it possesses trachez ; abdominal sensory sete are,
however, present in normal numbers. |
*#* Tibio-tarsus without the described clubbed hairs, having
instead sometimes outstanding, finely knobbed, clavate
hairs, Mucro with or without bristle.
Tribe SminrHurtint, C. B., 1918.
ii. Antenne bent between the 2nd and 3rd seyments, 4th always
shorter than the 3rd. Furcal segment with one pair of
dorsal papillze. Corpus tenaculi as in 1 (i. e. Sminthuridine).
Trachez absent (always?). Genital and aval segment fused.
Subfamily Dicyrromina&, C. B., 1906.
The Genus Srra, Lubbock.
In 1870, in his “ Notes on the Thysanura—Part 4,”
Lubbock described several Collembola new to the English
Fauna, and one formed the type of a new genus which he
called Sira (Seira). In his monograph, three years later,
he described several species under Stra, including
S. domestica (Nic.).
S. nigromaculata, Lbk.
S. buskit Lbk.
In later years the genus was split up, and Schott proposed
the name Pseudosira for types like the S$. domestica (Nic.),
leaving the name Sira for those lke nigromaculata and buskit.
But this should not be so, for Lubbock expressly states that
domestica forms the type of his genus Sta. Therefore
Pseudosira must fall and Sura take its place, and for the
species hitherto included in Sira, I propose the new name
Willowsia *.
1 Named after my friend Mr, F, W. Willows, of Tsolo, South Africa.
432 Mr. J. W. Shoebotham’s Notes en Collembola.
Genus Srra, Lbk., mihi.
Seira, Lubbock (1870), p. 279.
= Pseudosira, Schott.
Dentes ventrally covered with scales.
Type. S. domestica (Nic.).
Genus WILLowslA, gen. nov.
Dentes ventrally only with ciliated hairs, without scales.
Type. W. (Seira) nigromaculata (Lbk.).
List of Genera of Collembola found in the British Isles.
Class INSECTA.
Subclass Apterygota, Oudns,
Order COLLEMBOLA, Lbk.
Suborder ARTHROPLEONA, C. B.
Section PODUROMORPHA, C. B.
I, Family. Poduride (C. B.).
1. Genus Popvra, Linn., The.
II. Family Hypogastruride (C. B.).
Subfamily HypPoGAsTRURIN 2, C. B.
2. Genus HypoeastrurRa, Bourl., C. B.
3. Genus XENYLLA, The.
4. Genus WituEemia, C. B.
5. Genus ——?
A genus comes here related to both Hypogastrura
and Xenylla. I have an English species in my
collection, which in many respects is intermediate
between these two genera, but so far 1 have not
described it. Possibly it is Beckerella, Linnaniemi.
Subfamily AcHorutin~, C. B.
Tribe PsEuDACHORUTINI, CO. B.
6. Genus PsEUDACHORUTES, The.
7. Genus Micranurta, C. B.
8. Genus Anvripas, Laboulb.
9. Genus Friesxa, D. T.
Mr. J. W. Shoebotham’s Notes on Collembola. 433
10. Genus CoonpracHorvres, Wahler.
[The genus Chondrachorutes of Wahlgren has
not been previously recorded from the British
Isles, but it nevertheless occurs there, for I took
specimens at Berkhamsted, Herts, in 1910, but
have not yet described them. ] ;
Tribe AcHoRUTINI, C.B,
11. Genus AcnorvTEs, Templ., C. B.
= Neanura, MacG.
III, Family Onychiuride (Lbk., C. B.).
12. Genus OnycuruRvS, Gerv., C. B.
13. Genus ProTarHorvura, Absln., C. B.
14. Genus TuLiperera, Lbk.
Section ENTOMOBRYOMORPHA, C. B.
IV. Family Isotomide (Schffr., C. B.).
Subfamily [sorominz, Schitr,
15. Genus Isoroma, Bourl., C. B.
16. Genus Acrentr, C. B.
17. Genus Protsoroma, O.B.
17 a, Genus ? ARcutsotoma, Linnaniemi.
{Linnaniemi has proposed the genus Archi-
sotoma for the reception of Proisotoma bessellsi
(=P. spitzbergenensis, Lbk.), but as I have neither
my specimens of dessed/si nor Linnaniemi’s descrip-
tion by me at the moment, I cannot say whether
it should be regarded as a separate genus dr asa
subgenus of Prowsotoma. |
18. Genus ANvROPHORUS, Nic.
19. Genus Foutsomra, Willem.
20. Genus TETRACANTHELLA, Schtt.
21. Genus Isoromonss, Axels-Linn.
22. Genus Isoromurvs, C. B.
28. Genus ONcopopuRA, Carl & Lebed.
Subfamily AcraLeTin_”, C. B.
[The subfamily <Actaletine is not represented in the
British Isles. ]
434 Mr. J. W. Shoebotham’s Notes on Collembola.
V. Family Tomoceridz (Schffr.).
Subfamily LePrpoPHORELLIN.2, C. B.
[The subfamily Lepidophorelline is not represented in
the British Isles. ]
Subfamily Zomocerin” (C. B.).
24. Genus Tomoczrvs, Nice.
24 a. Genus P Poconognatuvs, C. B.
[Borner has proposed the genus Pogonognathus
for the species T'omocerus (P.) longicornis Miill.). |
VI. Family Entomobryide (C. B.). |
Subfamily Hvromopryinx (C. B.).
Tribe ENTOMOBRYINYI, C.B.
25. Genus Enromosprya, Rond.
26. Genus Sinexua, Brook.
27. Genus Lepipocyrtvs, Bourl.
28. Genus Srra, Lbk., Sbhthm.
29. Genus WILLOWSIA, gen. nov.
Tribe ORCHESELLINYI, C. B.
30. Genus OrcHESELLA, Templ.
31. Genus HerEromurvs, Wankel.
Subfamily Paroneriin#, C. B.
[The subfamily Puronelline is not found in the British
Isles. } :
Subfamily OyevoperRn& (C. B.).
Tribe TROGLOPEDETINI, C. B.
[This tribe has not yet been found in the British
Isles. |
Tribe CypPHODERINI, C. B.
32. Genus CyPpHopERUS, Nic.
Suborder SY MPHYPLEONA, C. B.
VII. Family Neelidz, Fism.
33. Genus NEELUS, Flsm.
34. Genus MreaLtorHorax, Willem.
Mr. J. W. Shoebotham’s Notes on Collembola. 435
VIII. Family Sminthuridz, Lbk.
Subfamily SwrwravripinZ, C. B.
Tribe SMINTHURIDINI, C.B.
35. Genus SMINTHURIDES, C. B.
Tribe KATIANNINI, C.B.
36. Genus SMinrHuRINUS, C. B.
37. Genus ARRHOPALITES, C. B.
Subfamily Suzwravrin A, C.B.
Tribe BouURLETIELLINYI, C.B.
88. Genus BourLETIELuA, Banks, C. B.
Tribe SMinTHURINYI, C.B.
39. Genus SmintrHurRus, Latr., C. B.
40. Genus Attacma, C. B.
41. Genus SpHryorueca, C, B.
Subfamily DicyrtominAz, C. B,
42, Genus Dicyrroma, Bourl., C. B.
43. Genus Dicyrromrna, C. B.
44, Genus Prenorurix, C. B. ©
References.
Apssouton, K. (1907.) “Zwei neue Collembolen-Gattungen.” Wiener
Entomol. Zeitung, Jahrg. 1907, pp. 335-348, with 3 figs,
Borner, C. (1901 a.) “ Vorliufige Mittheilung iiber einige neue
Aphorurinen und zur Systematik der Collembola.” Zool, Anz,
vol. xxiv. no. 633, pp. 1-15.
(19016.) ‘Zur Kenntnis der Apterygoten-Fauna von Bremen
und der Nachbardistrikte. Beitrag zu einer Apterygoten Fauna
Mitteleuropas.” Abh. Nat. Ver. Bremen, vol. xvii, Heft 1,
pp- 1-140, pls. i., ii., text-figs. 1-63,
(1903.) “dNeue altweltliche Collembolen, nebst Bemerkungen
zur Systematik der Isotominen und Entomobryinen,” Sitzungs-
Ber. der Gesellsch. naturf, Freunde zu Berlin, 1908, no. 8, pp. 129-
182, 1 pl.
(1906.) “Das System der Collembolen, nebst Beschreibung
neuer Collembolen des Hamburger Naturhistorischen Museums. 5
Mitt. aus dem Naturhist. Museum, xxili—Jahrbuch der Hamburg.
Wissensch. Anst. xxiii. pp. 147- 188, with 4 text-figs.
(1913a.) “Neue Cyphoderinen.” Zool. Anz, rok xli. no. 6
p. 274-284, figs. 1-9: ;
(1913 0.) “Die Familien der Collembolen.” Zool. Anz, vol. xli.
no, 7, pp. 815-822.
436 Mr. R. E, Turner on Fossorial Hymenoptera.
Forsom, J. W. (1896.) “ Neelus murinus, representing a new Thysa-
nuran family.” Psyche, vol. vii. no. 242, pp. 391, 392, pl. B.
Imus, A. D. (1912.) “On some Collembola from India, Burma, and
Ceylon; with a Catalogue of the Oriental Species of the Order.”
Proc. Zool. Soc. London, 1912, pp. 80-125, pls. vi—xii.
Lussock, J. (1862.) “Notes on the Thysanura, Part 1. Smyn-
thuride.” Trans. Linn. Soc. vol. xxiii. pp. 429-448, pls. xly., xlvi.
—. (1870.) “Notes on the Thysanura, Part 4.” Trans. Linn. Soc.
vol. xxvil. pp. 277-297, pls. xlv., xlvi.
——. (1873.) Monograph of the Collembola and Thysanura. London,
Ray Society, 1873, pp. 1-276, pls. i.—lxxviii.
Nicotet, H. (1842.) “Recherches pour servir a l'histoire des Podu-
relles.” Mém. de la Soc. Helv. des Sc. Nat. vol. vi. pp. 1-88,
Is. i.—ix.
Sieeeaees C. (1896.) ‘ Die Collembola der Umgebung von Hamburg
und benachbanter Gebiete.” Mitth. aus dem Naturhist. Museum,
vol, xiii. pp. 149-216, pls. i.—iv.
SHoEBoTHAM, J. W. (1914.) “ Notes on Collembola.—Part. 2. Some
Trish Collembola, and Notes on the Genus Orchesella.” Ann. &
Mag. Nat. Hist. ser. 8, vol. xili., Jan. 1914, pp. 59-68, pl. iii.
Sitvestri, F. (1911.) “ Termitiofili raccolti dal Prof. K. Escherich a
Ceylon.” Zool. Jahrb, vol. xxx. Heft 4, pp. 401-418, pls. v.—xi.
XLI.—Notes on Fossorial Hymenoptera.— XXVIII. On new
Ethiopian Species of Bembex in the British Museum. By
Row.anp EK. Turner, F.Z.S., F.E.S.
Bembex obtusa, sp. n.
3. Niger; mandibulis, apice excepto, labro, clypeo, fronte macula
obliqua nigra utrinque, scapo, orbitis externis, pronoto margine
postico, callis humeralibus, prosterno, mesosterno antice, meso-
noto linea supra tegulas, scutello linea obliqua utrinque, post-
scutello margine postico, pedibusque flavis; femoribus anticis
supra nigro-lineatis, tarsis anticis infra nigro-maculatis; seg-
mento dorsali primo macula transversa utrinque fasciaque trans-
versa mediana angustissima, segmentis dorsalibus 2-6 fascia
transversa bisinuata, ventralibus 2-5 macula utrinque, segmento-
que ventrali primo fascia angusta apicali flavo-olivaceis; flagello
subtus ochraceo; segmento dorsali sexto apice, septimoque
dimidio basali ferrugineis; alis hyalinis, venis fuscis, thorace
duplo longioribus,
Long. 21 mm.
3. Clypeus very broadly triangularly flattened on the
apical half, labrum flattened at the base; a strong longitudinal
carina between the antenne; sixth, seventh, and eighth joints
of the flagellum each with two or three small spines beneath,
Mr. R. E. Turner on Fossorial Hymenoptera. 437
joints 9 and 10 subdenticulate beneath, penultimate joint
concave beneath, longer than the tenth, apical joint about
half as long again as the penultimate, concave beneath, blunt
and rather strongly curved at the apex. Fore tarsi rather
stout, with a strong tarsal comb, the apical joint flattened,
nearly as broad and Jess than half as long again as the
penultimate, with a small spine on the middle of the outer
margin ; anterior and intermediate femora not serrate ; inter-
mediate tibize produced into a distinct spine at the apex.
Seventh dorsal segment very broad, with parallel sides on
the basal portion to beyond the middle, the apex obtuse; the
surface of the segment finely punctured, with coarse punc-
tures intermixed near the apex. Second ventral segment
with a very large tubercle, which is broadly truncate at the
apex ; sixth ventral segment with a slightly raised, broadly
triangular area near the apex; seventh broad, with a longi-
tudinal carina on each side; eighth produced into a stout
blunt tooth. Median cell of the hind wing emitting only one
vein from the apex.
Hab. Nyasaland, Mlanje, 2300 ft. (S. A. Neave), October.
The apical dorsal segment is shaped somewhat as in the
variety of B. pugillatria figured by Handlirsch (Sitzungsb.
Akad. Wiss. Wien, cii. t. v. fig. 15), but the paraliel sides
are continued much nearer to the apex than in that figure.
In the antenne it approaches albofasciata, Sm. (=karschii,
Hand].), also in the short wings, but differs in the legs and
apical abdominal segments. The three intermediate joints of
the fore tarsi are moderately dilated, about as broad as long,
nearly as much dilated as in latctarsis, Hand).
Bembex johnstont, Turn.
Bembex johnstoni, Turn, Ann. & Mag. Nat. Hist. (8) x. p. 3872 (1912).
Gus
This belongs to the group of B. diversipennis, not of mébii,
to which I compared it in the description; it is, indeed,
probably only an extreme colour-variation of diversipennis.
In a considerable series of that species from H, Africa tlie
thorax is always without yellow markings in both sexes,
except in one female from Harar. The yellow markings on
the abdomen vary much, but do not form continuous fascize
as in most West-African specimens. The wings are hyaline
in the male, more or less strongly fuscous at the base in the
female in all specimens which I have seen.
438 Mr. R. E. Turner on Fossorial Hymenoptera.
Bembex albidula, sp. n.
g. Niger; abdomine ceruleo-tincto; mandibulis, apice excepto,
labro, clypeo macula magna basali nigra utrinque, scapo subtus,
orbitis, macula parva utrinque sub ocellis, prosterno, callis
humeralibus apice, segmento mediano angulis posticis, segmentis
dorsalibus 2-4 fascia late interrupta, primo macula magna
utrinque, quinto fascia anguste interrupta, sexto linea transversa
utrinque, segmentis yentralibus secundo tertioque macula parva
utrinque, femoribus anticis intermediisque subtus, tibiis supra
nigro-lineatis, tarsisque anticis intus albidis; alis hyalinis, venis
fuscis, antice latitudine thoracis duplo et dimidium longiores.
Long. 22 mm.
3. Eyes distinctly divergent towards the clypeus ; labrum
flattened, clypeus convex; carina between the antenne very
low. Antenne almost normal; eighth joint of the flagellum
with two very minute spines beneath ; ninth with a minute
spine near the base; apical joint slightly curved, rounded at
the apex, longer than the penultimate. Fore tarsi mode-
rately stout, the joints not dilated ; the basal joint with seven
spines, the basal spine small. Fore and intermediate femora
unarmed, intermediate tarsi simple, intermediate tibia not
produced at the apex and without a long apical spur.
Seventh dorsal segment subtriangular, narrowly rounded at
the apex, closely punctured at the base, sparsely at the apex.
Second ventral segment with a small tubercle near the apex ;
sixth produced into a stout blunt tooth at the apex; seventh
unarmed, without carine; eighth in the form of an acute
spine. Median cell of the hind wing emitting two veins from
the apex.
Hab. Sierra Leone, Mussaia (J. J. Simpson), April.
This fine species is somewhat intermediate between diversi-
pennis and monedula, but seems to belong to the group of the
latter, though distinguished from it by the spines on the
eighth and ninth joints of the flagellum, and by the much
narrower seventh dorsal segment. Handlirsch, in giving the
distinguishing characters of the group, says “ Mittelschenkel
geziilnt,” but in describing the species says “ femoribus inter-
mediis infra non dentatis.” The latter statement is correct,
B. ugandensis, Turn., is also near this species, but differs
in the structure of the antennee, the hollow grooves beneath
the three apical joints being well marked in ugandensis and
almost obsolete in the present species, the spines on the other
joints are also different. The colour-differences, though
great and apparently constant locally, cannot alone be relied
on in this group.
Mr. R. E. Turner on Fossorial Hymenoptera. 439
Bembex odontopyga, sp. n.
g. Niger; mandibulis, apice excepto, labro, clypeo macula basali
nigra utrinque, scapo subtus, orbitis, femoribus, tibiisque subtus,
tarsisque flavis; flagello subtus obscure brunneo; segmentis
dorsalibus 2-4 fascia angusta late interrupta, quinto sextoque
fascia: continua, septimo macula obliqua utrinque, segmentisque
ventralibus 2—5 fascia angusta continua apicali brunneo-flavis ;
segmentis dorsalibus obscure czruleo-micantibus ; segmento
septimo dorsali apice fusco-ferrugineo ; alis hyalinis, venis fuscis.
Long. 19 mm.
3d. Clypeus with a carina from the base to beyond the
middle, deflexed from the end of the carina to the apex, not
very strongly convex ; the carina between the -antenne very
distinct. Four apical antennal joints distinctly hollowed
beneath, stout, the apical joint blunt at the apex, a little
longer than the penultimate. Head, thorax, and median
segment clothed with long whitish pubescence, that on the
mesonotum shorter and brownish. Fore tarsi normal, the
basal joint with-seven spines ; fore and intermediate femora
not serrate. Median segment with a very feeble longitudinal
groove. Seventh dorsal segment with a short spine on each
side near the base, the apex rather broadly truncate, not
undulating at the sides. Ventral segments unarmed. Wings
about two and a half times as long as the breadth of the
thorax; median cell of the hind wing emitting two veins
from the apex. The eyes are distinctly divergent towards
the clypeus.
Hab. Nyasaland, Ngara (Dr. J. E. S. Old), October.
This belongs to the bidentata group, and apart from colour-
differences may be distinguished from mébii by the absence
of a strong tooth at the apex of the intermediate femora; from
bidentata by the more distinct carina of the clypeus, by the
colour of the pubescence, and by the broader apex and less
developed teeth of the seventh dorsal segment. B. scotti,
Turn., the only remaining African species of the group, may
be at once distinguished from this by the much finer and
sparser puncturation of the thorax and median segment. In
colour odontopyga resembles compedita, Turn., which is allied
to fuscipennis.
Bembex forcipata, Hand.
Bembex forcipata, Handl. Sitzungsh. Akad. Wiss. Wien, cii. p. 798
(1893). 3.
Bembex massaica, Cameron, Sjistedt, Kilimandjaro-Meru Exp, ii.
p. 290 (1910). ¢.
440 Mr. R. E. Turner on Fossorial Hymenoptera.
Bembewx liturata, sp. n.
3. Niger; labro albido, basi macula longitudinali brunnea; man-
dibulis, apice excepto, clypeo fascia transversa basali nigra, scapo
subtus, orbitis, pronoto margine postico, propleuris callis humer-
alibus, mesosterno antice, tegulis macula, linea supra tegulas,
scutello margine postico, pedibusque flavis; tibiis supra nigro-
lineatis; segmentis dorsalibus sex basalibus fascia flavo-olivacea ;
segmentis ventralibus primo apice, secundo fascia lata transversa,
3-6 macula utrinque flavis; alis hyalinis, venis fusco-ferrugineis.
9. Mari similis, clypeo dimidio basali nigro; segmento mediano
macula parva utrinque flavo; segmento ventrali secundo dimidio
apicali brunneo-ferrugineo, utrinque flavo-maculato.
Long., ¢ 17, 2 15 mm.
gd. Clypeus strongly convex; inner margin of the eyes
almost parallel ; no carina between the antenna; seventh
joint of the flagellum emarginate at the apex beneath, eighth
with a small spine beneath, ninth and tenth stout, penultimate
joint much broadened, longer than the subconical apical joint.
Tarsi slender, the spines of the comb of the anterior tarsi
slender ; intermediate femora very feebly serrate. Seventh
dorsal segment very broadly rounded or subtruncate at the
apex, with a marginal carina on each side near the base.
Second ventral segment with a strong tubercle, which is
rounded at the apex; sixth with a large raised semicircular
area. Wings rather short, only twice as long as the breadth
of the thorax ; median cell of the hind wing emitting only
one vein from the apex.
?. Basal joint of the fore tarsus with seven spines ; sixth
dorsal segment broadly triangular, very sparsely punctured in
the middle, more coarsely and closely on the sides ; second
ventral segment shining in the middle, with large sparse
punctures.
Hab. 8. Africa, Willowmore (Dr. H. Brauns), December.
The male genitalia are not strongly curved as in the me/a-
nopa group, the relationship being rather with the mediterranea
graup.
Bembex opima, sp. n.
3. Niger, clypeo, mandibulis, apice excepto, Jabroque albo-flavi-
dulis; flagello subtus brunneo-ferrugineo ; scapo, supra nigro
maculato, orbitis, pronoto postice, callis humeralibus, tegulis,
mesonoto linea supra tegulas, scutello margine postico, postscutello
margine postico, segmento mediano fascia angusta apicali, seg-
mento dorsali primo dimidio apicali, secundo antice nigro bi-
sinuato, 3-6 omnino, segmentis yentralibus 2-5 lateribus et
Fabricius’s Types of Odonata in the British Museum. 441
margine apicali, pedibusque flavis; alis hyalinis, venis fusco-
ferrugineis.
©. Mari similis ; macula flava sub ocellis, segmento dorsali secundo
flavo, macula parva nigra utrinque, 3-5 basi anguste nigris, sexto
nigro macula flava apicali utrinque; ale antice thoracis latitu-
dine duplo et dimidium longiores.
Long., ¢ 9,16 mm,
3. Clypeus strongly convex, labrum flattened, no carina
between the antenne ; seventh, eighth, and ninth joints of
the flagellum spined beneath, apical joint curved, truncate at
the apex, no longer than the penultimate. Anterior femora
unarmed, intermediate femora very obscurely serrate ; ante-
rior tarsi normal, the basal joint with six spines ; intermediate
tibie and tarsi normal. Second ventral segment with a
strong tubercle acute at the apex; sixth ventral segment
with a slightly raised triangular area; seventh dorsal
segment closely punctured, narrowly rounded at the apex,
the sides with short, stiff, black pubescence. Median cell
of the hind wing emitting two veins from the apex, the lower
vein ill-defined. Head and thorax thickly clothed with
grey pubescence.
?. Middle of the second ventral segment shining, with
deep sparse punctures ; sixth dorsal sezment subtriangular,
very narrowly rounded at the apex, closely punctured, with
stiff sete on the sides, subcarinate longitudinally in the
middle.
Hab. 8. Africa, Willowmore (Dr. H. Brauns), January
and February. ;
The seventh ventral segment of the male has a median
longitudinal carina.
‘This seems to be nearest to capicola, Handl., though
differing in the armature of the ventral segments of the male,
the shape and sculpture of the seventh dorsal segment, and
in other smaller details.
XLII.—On Fabrictus’s Types of Odonata in the British
Museum (Natural History). By HERBERT CAMPION.
As far as I am able to ascertain, the British Museum Collec-
tions include fourteen Dragonflies which have been described
or determined by Fabricius. All but two of them came in
Sir Joseph Banks’s Collection of insects, which was presented
by the Linnean Society in 1863. Although the welfare of
Ann. & Mag. N. Hist. Ser. 8. Vol. xix. 29
442 Mr. H. Campion on Fabricius’s Types of
the collection necessitated its removal from the large and
ornate cabinet in which it was then contained, it has fortu-
nately remained intact and separate from the main collections
of the Museurn.
With regard to the identification-labels relating to these
historical specimens, an attempt has been made to determine,
by the character of the handwriting itself, whether Fabricius
was the author of all or any of them. Through the good
offices of Mr. J. H. Durrant I have been able to examine the
caligraphy of two holograph letters written by Fabricius in
Paris in 1805. Although both: were produced in the same
year, there is a considerable difference between them in style
and even in the formation of letters—a fact which makes it
all the more difficult to distinguish Fabricius’s hand from
other writing of the same period. At the same time, these
Jabels are in two different styles of writing, at least, and the
two styles may even be detected upon the same label.
Nevertheless, I think it may be assumed that most, if not all,
of the drawer-labels in the Banks Collection are the work of
Fabricius himself. The case of the pin-labels in the General
Collection is much clearer, as they correspond exactly, both in
form and caligraphy, with the labels attached to certain
undoubted Fabrician types of Coleoptera to which Dr. C. d.
Gahan has kindly called my attention.
I. SPECIMENS IN THE BANKS COLLECTION.
The twelve Banksian dragonflies, which nominally repre-
sent as many species, are in a fair state of preservation, and,
with the exception of those labelled Libellula carolina, Aishna
grandis, Agrion virgo, and A, linearis, are the types of
species described by Fabricius at various dates. Five of
these are the types of nominal species merely, the valid
species represented by their types being Lebellula [ Neuro-
themis | stigmatizans, L. | Rhyothemis| notata, aud Agrion
[ Sapho | ciliata. Certain other types referred to by Fabricius
as being in the Banksian Collection are not now to be found
there, and I am unable to learn anything concerning their
fate. The missing insects are Libellula bifasciata (=? Li-
bellula lydia, Drury, ? ), L. variegata (=Lalpopleura lucia,
Drury, 2), and AHshna vuriegata. As regards the last-
named, the British Museum register of accessions expressly
mentions Ashna variegata, from Tierra del Fuego, as being
in the Banksian Collection at the time of its acquisition by
the Museum (1863). In 1887 McLachlan knew of its
disappearance, and wrote, ‘‘ In order to save disappointment,
Odonata in the British Museum. 443
I state that it no longer exists in Mus. Banks, and has
probably been long ago destroyed” (int. Mo. Mag. xxiv.
p-77). It is unlikely now that the obscurity which surrounds
the identity of this species, as well as the history of the type,
will ever be cleared up.
No fewer than five of the specimens under consideration
have obviously incorrect habitats assigned to them in Fa-
bricius’s writings. These are Libellula equestris (= Neuro-
themis tullia, Drury), L. ferruginata (= Crocothemis servilia,
Drury), Zshna grandis (= Aischna cyanea, Miill.), Agrion
ciliata (= Sapho ciliata, F.), and A. lineards (= Mecistogaster
linearis, F.). OE£ course, lapses of this description were not
at all infrequent at a time when little or no importance was
attached to the facts of geographical distribution.
Below each insect stands a large oblong drawer-label, with
a double black border, bearing the name of the genus and
species, as well as a reference to the published description.
The labels applying to Libellula stigmatizans, L. oculata, and
L. carolina have a portion of the reference printed in—z. e.,
“ Fab. Entom. p.” in the case of the two first-named, and
“Linn. S. N. p.” in the case of the last-named. The only
pin-labels of any description are fonr tickets marked with the
British Museum registration number, and five modern-looking
tickets bearing the name of the reputed country of origin.
Three manuscript genus-labels, dividing the collection into
the genera Libellula, A’shna, and Agrion, may also be of
post-Fabrician date.
In considering the specimens seriatim, Fabricius’s original
diagnosis of each of the eight types may be usefully quoted,
but for our present purposes his more detailed descriptions
need not be consulted, and will therefore be omitted.
(1) Libellula stigmatizans, F. ¢. Type.
(= Neurothemis stigmatizans, ’., 9 .)
Labels :— Libellula stigmatizans Fab. Entom. p. 421,
n. 5°’; square white ticket, ‘* Australia,” printed ; round
blue;ticket, ‘£2.
Diagnosis :—* L. flavescens, alis macula apiceque fuscis :
stigmate niveo. Habitat in nova Hollandia. Mus, Bank-
tanum.”’—F., Syst. Ent. p, 421, no. 5 (1775).
This specimen and the next were examined by De Selys,
and were identified by him as the ? and @ respectively of a
single species (Ann. Mus. Civ. Genova, xiv. pp. 292, 293;
1879). Although he adopted the name of the g (oculata)
as that of the species, the modern rule Rep Ddne page-
29
444 Mr. H. Campion on Fabricius’s Types of
precedence requires the application to the species of the first
name written by Fabricius (stigmatizans).
(2) Libellula oculata, F. ,jav. Type.
(= Neurothemis stigmatizans, F., 8.)
Labels :—*“ Libellula oculata Fab. Entom, p. 421, n. 6”;
square white ticket, ‘ Australia,” printed; round blue
ticket,.4: ee
Diagnosis :—‘* L. flavescens, alis anticis apice, posticis
margine aqueis: stigmate niveo. Habitat in nova Hollandia,
Mus. Bankianum.”—F., Syst. Ent. p. 421, no. 6 (1775).
Fabricius’s additional words, “ Preecedenti [ Z. stigmatizans |
nimis affinis, cujus forte mera varietas,” are worthy of note.
See also the remarks under (1) ZL. stigmatizans, above.
(3) Libellula indica, F. 9. Type.
(= Rhyothemis variegata, Linn.)
Label :—* Libellula indica Fabr. Sp. Ins. No. 8.”
Diagnosis :—‘‘ Li. alis flavo fuscoque variis apice albis,
posticis macula baseos cyanea. Libellula Arria. Drury Ins.
2. tab. 46. fig. 1. Habitat in Coromandel. Mus. Dom.
Banks.” —F., Spec. Ins. i. p. 521, no. 8 (1781).
The most noticeable difference between Lhyothemis varie-
gata variegata and Rh. variegata arria, the Indian and
Chinese forms of this variable species, is one of size, and in
this respect our type undoubtedly belongs to the smaller
Indian form, the abdomen measuring 20°5 mm. and the hind
wing 30°5 mm.
(4) Libellula notata, F. Type.
(= Rhyothemis notata, F.)
Labels :—“ Libellula notata Fabr. Mss. Ins. n. 19”;
oblong white ticket, ‘‘ Sierra Leone,” written.
Diagnosis :— L. alis planis nigris: maculis apiceque
albis. Habitat in Sierra Leon Africae Mus. Dom. Banks.’’—
F., Mant. Ins. 1. p. 337, no. 19 (1787).
This type is a normal specimen of the West-A frican species
figured and described by Ris as the true notata of Fabricius
(Coll. Selys, Libell., fasc. xv. p. 959, pl. vii.; 1913). The
abdomen is missing, and the sex has not been definitely
determined, although the wing-pattern rather indicates a
male. The hind wing measures 26 mm.
Odonata in the British Museum. 445
(5) Libellula equestris, F. 8. Type.
(= Neurothemis tullia, Drury.)
Labels :—‘ Libellula equestris Fabr. Sp. Ins. No. 20”;
square white ticket, ‘“ Africa,” written.
Diagnosis :— L. alis dimidiato nigris, fascia media nivea.
Habitat in Africa equinoctiali. Mus. Dom. Banks.”—F.,
Spec. Ins. i. p. 523, no. 20 (1781).
This insect agrees very well with Drury’s figure of his
Libellula tullia, adult ¢, from Bombay (Ill. Ex. Ent. ii
pl. xlvi. fig. 3; 1773). That figure, however, fails to show
the transverse white band in the wings, and is also a little
larger than Fabricius’s type, the hind wing measuring 22 mm.,
as compared with 21 mm. Im our type the last antenodal is
continuous in both fore wings. ‘The reference to Africa is, of
course, a mistake, as the genus Veurothemis is entirely
unknown from that continent.
(6) Libellula marginata, F. g. Type.
(=Palpopleura lucia, Drury.)
Labels :—* Libellula marginata Fabr. Sp. Ins. No. 24”;
square white ticket, ‘‘ Africa,” written; round blue ticket,
66 63 7
47°
Diagnosis :— L. alis nigris, anticis macula apicis, posticis
margine albis, Habitat in Africa equinoctiali. Mus. Dom.
Banks.”’—¥., Spec. Ins. i. p. 523, no. 24 (1781).
Comparing this specimen with the plate of coloured figures
of Palpopleura lucia in Ris’s monograph of the Libellulins
(Coll. Selys, Libell., fase. ix. pl. i.), we find that the pattern
of the fore wing is intermediate between the first and second
of those figures, while that of the hind wing is —e
between the second and third figures.
(7) Libellula ferruginata, F. g. Type.
(= Crocothemis servilia, Drury.)
Label :—* Libellula ferruginata Fabr. Sp. Tns Nor bh
The specific name was written originally ‘ ferruginea,” and
was altered subsequently to “ ferruginata.’
Diagnosis :— Li. alis basi eee corpore obscure
ferrugineo. Habitatad Cap. Bon. Spei. dus. Vom. Banks.”
—F., Spec. Ins. i. p. 521, no. 11 (1781).
ye “fer ruginata, F’., has ‘been erroneously identified with the
common African species Crocothemis erythrea, Brullé, because
the Cape of Good Hope was stated to be its habitat. ‘This
446 Mr. H. Campion on Fubricius’s Types of
locality, however, is manifestly a wrong one, as Fabricius’s
type clearly belongs to the Asiatic form Crocothemis servilia,
Drury, 1773. This is shown by the abdomen, which is
parallel-sided and devoid of mid-dorsal black spots, and by
the wings, which are narrow and rather smoky at the tips.
The abdomen measures 27°5 mm. and the hind wing 34 mm.
There are 11} antenodals in each fore wing. Drury’s figure
of Libellula servilia, from China (Ill. Ex. Ent. i. pl. xlvi.
tig. 6; 1770), agrees fairly well with the type of L. ferrugi-
nata, the main points of difference being that the abdomen is
too long and that the coloured area at the base of the fore
wings is too large. In 1793 (Ent. Syst. ii. p. 380) Fa-
bricius treated his L. ferruginata, as wellas L. servilia, Drury,
as synonyms of his L. ferruginea. In the original deserip-
tion of L. ferruginea, 1775, the habitat was given as
“ America,” but in 1793 the habitat was changed to “ China.”
(8) Libellula carolina, Linn. 6.
(=Tramea virginia, Ramb.)
Label :—“ Libellula carolina Linn, 8. N. p. 504. n. 17.”
‘The base of the abdomen is in poor condition, the contents
having apparently been eaten out by mites, and the hamules
have disappeared. Nevertheless, the character of the basal
spot in the hind wing is sufficient to show that the insect is
not the North-American Tramea carolina, Linn., but the
Chinese species now known as Yramea virginia, Ramb.
This Chinese species is usually referred to as Tramea chi-
nensis, De Geer, but, as a case of homonymy is involved,
Dr. Ris (Coll. Selys, Libell. fase. xvi. (1) p. 978; 1913) has
recently restored to it Rambur’s name of virginia. Re-
examination of Rambur’s type has revealed its Asiatic
origin, notwithstanding that author’s erroneous citation of its
habitat as “ Amérique septentrionale” (Ins. Névr. p. 34;
1842).
(9) shna grandis, Linn. 3s.
(= dischna cyanea, Mill.)
Label:—“ Aishna Grandis Fabr. Sp. Ins. No. 2.”
Lieference :—“ A, thorace lineis quatuor flavis, corpore
variegato, Habitat et in Insulis Sandwichii Mus. Dom.
Banks.”’—I*., Mant. Ins. i. p. 339, no. 2 (1787).
In this passage, which is quoted zn ewtenso, Fabricius
proposes to extend the distribution of 4. grandis, as given
in his earlier writings, so as to include the Sandwich Islands.
The locality is not repeated in Fabricius’s list of 1793, and
Odonata in the British Museum. 447
is, of course, entirely erroneous. The insect is, in fact, a g
of the common European species Aschna cyanea, Miill.,
which was not recognized as a distinct species until 1764.
It seems to have been confused by many of the older entomo-
logists with 42. juncea, Linn., or even, as in the present case,
with i. grandis, Linn.
(10) Agrion ciliata, F. 9. Type.
(=Sapho ciliata, ¥.)
Labels :—“ Agrion ciliata Fabr. Sp. Ins. No. 3”; round
blue ticket, “$7.”
Diagnosis :—‘ A. viridi enea, abdomine fusco, pedibus
ciliatis nigris. Habitat in Coromandel. dus. Dom. Banks.”
—F’., Spec. Ins. i. p. 528, no. 3 (1781).
Concerning this specimen De Selys wrote thus:—*“ J’ai
reconnu notre espéce dans l’exemplaire type de lAgrion
ciliatus male, de Fabricius, qui existe encore heureusement
dans la collection de Joseph Bancks, déposée a la Société
Linnéenne de Londres. Dans ses ouvrages, Fabricius in-
dique spar erreur Coromandel comme la patrie du cliatus,
mais j’ai examiné avec soin ’exemplaire type sous le rapport
de la réticulation, et je me suis assuré qu’elle est en tout con-
forme a celle des individus de Sierra Léone” (Monogr.
Calopt. p. 60, 1854). Fabricius’s type, however, is not a
male, as stated by De Selys, but a small example of the
female sex. The abdomen is 41 mm. in length and the hind
wing 37 mm.
(11) Agrion virgo, Linn. ¢.
(= Calopteryx splendens, Harr.)
Label :—“ Agrion Virgo Fabr. Sp. Ins. p. 526. n. 1.”
This is the common European species Calopteryx splendens,
Harr., which was regarded by Linnzeus as merely a form of
C. virgo. Fabricius apparently adopted the same view, and
never recognized Harris’s action in 1782 in separating the two
forms specifically.
(12) Agrion linearis, F. .
aoe (= Mecistogasier linearis, F.)
Label :— Agrion Linearis Fabr. Sp. Ins. No. 5.”
This specimen is not the type, the species having been
described in 1776 from material in the possession of Dr. Fother-
gill. The type cannot now be traced, and in its absence it is
impossible to say precisely what Fabricius’s species may be.
448 Mr. H. Campion on Fabrietus’s Types of
In 1781 he identified it with Mecistogaster lucretia, Drury,
and some colour is lent to this identification by the fact that
both species were described from Fothergill’s collection.
Drury stated that his lueretia came from the Cape of Good
Hope, while Fabricius gave India as the habitat of his
linearis; but, of course, Mecistogaster is exclusively a Neo-
tropical genus. Whatever may be the identity of the
Fabrician type, the species in the Banks Collection is quite
distinct from that figured by Drury. The specimen before
us was examined by De Selys, and referred by him to the
species which he described as J/. linearis, F. (Bull. Acad.
Belg. (2) x. p. 22, 1860). A note appended to that deserip-
tion may be usefully quoted here:—“ L’exemplaire de la
collection Banks & Londres, qui passe pour avoir été étiqueté
par Fabricius, est un male de cette espéce, 4 ptérostigma
brun (semi-adulte). Les figures de Drury et de Sulzer,
citées & ’appui dans |’ Hntomologia systemaiica, sont au con-
traire la lucretia. Quant a la description de Fabricius, elle
peut s’appliquer aux deux espéces. Si l’on devait prendre le
linearis de Fabricius pour synonyme de lucretéa (nom plus
ancien), il faudrait adopter pour notre espéce linearis le nom
de tullia, de Burmeister.”
II. SPECIMENS IN THE GENERAL COLLECTION OF THE
Britisu Museum.
In 1793 (Ent. Syst. ii.) Fabricius referred to three
dragonflies in the British Museum Collection. These were
Libellula trimaculata, De Geer (=L. lydia, Drury), p. 374,
no. 3; LL. sinuata (=Palpopleura lucia, Drury), p. 378,
no. 17; and L. vibrans, p. 380, no.30. The first is involved
in much obscurity, and the second has not been traced at all,
but LZ. vibrans has been identified with certainty. Unlike
the Banksian insects, the two Fabrician specimens now in
the General Collection carry a plain buff pin-label, with the
two upper corners cut off, and bearing the name of the
species in Fabricius’s handwriting.
(1) Libellula lata, F. 9¢.
(=L. lydia, Drury, ¢ .)
Label :—* Libellula lata Fab.”
Apparently this name was never published, and the only
reference to it which I can find is one contained in an inter-
leaved and annotated copy of Linneus’s ‘Systema Nature’
(ed. xii.). This book is preserved in the British Museum
Odonata in the British Museum. 449
(Natural History), but the authorship and date of the supple-
mentary descriptions with which it abounds are unknown
tome. Several manuscript additions to the genus Libellula
have been made, and among them is the following :—
‘lata. LIB. alis planis hyalinis fascia maculaque baseos
nigro- fusca, abdomine carinato glauco.
Mser.*
Hab. in Pensylvania, Marylandia, Carolina.”
These words, however, while agreeing sufficiently well with
the male of Z. lydia, do not apply to the Fabrician specimen,
which is a female of that species.
There seems to be some reason for believing that Libellula
bifasciata, F’., usually identified with LZ. pulchella, Drury,
may be, in reality, the 2 of LZ. lydia, Drury. The last-
named insect is very different in its wing-markings from its
corresponding @, which is undoubtedly the ZL. trimaculata of
De Geer and Fabricius, but is very like Z. pulchella, and
especially the ? of that species. It is true that in 1793
Fabricius identified his L. bifasetata, 1775, both with Drury’s
figure of L. pulchella and also with Petiver’s figure (Gazo-
phylacium, i. pl. xv. fig. 2) of what is clearly intended for
L. lydia, 2. ‘his is, however, evidently a case of confusion,
due to the similar appearance of the two insects, and the
difference in size was overlooked. But it isa very significant
fact that Fabricius compares both bzfaseiata and trimaculata,
in respect of build and size, with LZ. depressa, Linn., and it
may be more natural to look upon them as the ? and ¢,
respectively, of the same species, LZ. lydia, than to regard
bifasciata as L. pulchella, which is decidedly larger than
L. depressa, and trimaculata as L. lydia, 8, which is rather
smaller than L. depressa.
The fact that the specimen under consideration undoubtedly
belongs to ¢rtmaculata, and is the only one of the kind in the
British Museum which possesses any label in Fabricius’s
handwriting, would lead one to suppose that it is really the
type of his description of that species. But this view of the
matter seems to be negatived, both by the description itself,
and by the two figures (De Geer, Mém. Ins. iii. pl. xxvi.
fig. 2, and Petiver, Gazoph. i. pl. xv. fig. 1) cited in illustra-
tion of it, all of which apply to the male sex alone.
(2) Libellula vbrans, F. 9. Type.
Labels :—* Libellula vibrans Fab.” ; ‘“ Georgia.”
Diagnosis :—< L. alis planis albis: macula media atra
450 Mr. O. Thomas on
apicibusque ferrugineis. Habitat — — Mus. Britann.’—F.
Ent. Syst. ii. p. 380, no. 30 (1793).
Although Fabricius did not indicate any locality for this
type, it nevertheless carries a small round label inscribed
“Georgia.” The fact that the written surface of the label
had been placed in direct contact with the pectus of the
dragonfly no doubt led to its being overlooked. The specimen
is in an unusually good state of preservation, and the measure-
ments are as follows: —Abdomen 39°5 mm., hind wing 49 mm.,
pterostigma 7 mm. In the fore wings the triangles are
3-celled, the subtriangles are 6-celled, and the antenodals
number from 163 to 18. At the base of each wing a dark line
in the subcostal space reaches nearly to the third antenodal.
The nodal spot on all wings is small, and no markings of any
kind lie between that spot and the pterostigma. The brown
apical cloud on all wings is small, not reaching inwards much
beyond the distal end of the pterostigma.
y]
XLITI.—A new Vole from Palestine.
By OLDFIELD ‘THOMAS.
(Published by permission of the Trustees of the British Museum.)
In 1913 the British Museum received as a donation from
Mr. N. Charles Rothschild six voles from Ekron, south-east
of Jaffa, and these were provisionally put down as Microtus
syriacus, Brants.
Inquiry was, however, made of Prof. Matschie as to certain
details of the type of that species, and with the help of his
account I am now able to recognize that the Ekron vole is
distinct and should be described as new.
Microtus philistinus, sp. n.
Like M. lydius, Blackler, but bulle larger.
Size and general colour above quite as in MZ. lydius, the
back sandy brown, rather more buffy than Ridgway’s “ buffy
brown.” Sides more buffy, but not so strongly as in lydius.
Under surface washed with buffy, more so than in lydius, in
which the ends of the hairs are greyish white, less so than
in guentheri. Hands and feet buffy fawn. Tail as long as in
lydius, longer than in guentheri ; its upper surface terminally
aistinetly blackened, which is not the case in lydius ; its
lower surface pale buffy—white in lydius.
a new Vole from Palestine. A451
Skull and teeth like those of MZ. lydius, with the important
exception that the bulla, although unusually variable in size,
are conspicuously larger in most specimens and_ slightly
larger in all. Height from crown to molars markedly less
than in M. hartingi.
Dimensions of the type :—
Head and body 125 mm. ; tail 33; hind foot 20.
Skull: condylo-incisive length 29°7; zygomatic breadth
16°5 ; nasals 8°2 x 3:9; length of brain-case from postorbital
angle backwards 13:6; palatilar length 14°8; diastema 9 ;
palatal foramina 5°5; length of bulla from front of par-
occipital process in a straight line forwards 85; upper
molar series (crowns) 6°9.
Hab. Ekron, 8.E. of Jaffa, Palestine.
Type. Adult male. B.M. no. 14.1.16.1. Collected
1st December, 1913, by Tl’. Aharoni. Presented by the Hon.
N.C. Rothschild. Six specimens.
From A. guenthert, Danf. & Alst., this species is distin-
guished by its longer tail, and from JZ. lydius by the various
characters above enumerated, notably by its larger bulle.
With regard to the two voles from Palestine described
long ago by Brants & Wagner, “ Hypudeus syriacus” and
“* Hypudeus cinerascens,” the latter is soon disposed of, as it
is clearly a hamster (Cricetulus), and not a vole at all.
Mr. Aharoni has sent examples from Jaffa agreeing with the
description in every detail.
Microtus syriacus, from the Lebanon (fide Matschie), is
said to be a greyish, not a sandy-brown, species, and the
accounts of the type sent me by Prof. Matschie show it to
have been smaller than M. philistinus (upper tooth-row
5°7 mm., diastema 6°9), and to have been apparently of a
different group of voles. For he says of the teeth that m,
has only seven spaces, with four projecting angles on its outer
side, numbers never found in the present group, in which
nine spaces and five outer angles always occur.
_ Whether MM. syriacus may prove to be a young Chiono-
mys or some totally different form of vole, still remains to
be seen. Both Brants and Matschie have been struck by the
unusual length of its whiskers, the longest of which measures
36°55 mm. Those of J, philistinus are of quite moderate
length.
I have provisionally used a binomial for the Ekron vole,
but think it probable that it may grade into M. lydius and
guenthert, the latter the earliest described of the group.
A452 Mr. O. Thomas on the small Hamsters
XLIV.—On the small Hamsters that have been referred
to Cricetulus phzus and campbelli, By Otxprizip
Tomas.
(Published by permission of the Trustees of the British Museum.)
Tue small unstriped Hamsters with naked soles, which
occur over an area ranging from the Crimea, Asia Minor,
and Palestine on the west to the Altai, Kashehar, and
Ladak on the east, have all been eethier referred to one
species, called phaus, or occasionally split up into several on
size, a character that proves most illusory. Both their
nomenclature and characters still remain in much confusion.
Firstly, I regret to say that the name pheus does not
stand for any of them, as it is antedated by an earlier term.
In 1773* Pallas described, of this group, the species
migratorius (type-locality: R. Ural, 8.W. of Orenburg),
sungorus ¢ (RK. Irtish, Siberia), arenarius (Irtish), and
barabensis (Irtish) the first and fourth of which he chan ged
in the ‘Glires’ ¢ to accedula and furunculus, while he added to
them pheus (type-locality: Lower Volga, near Sarepta).
By a curious fatality, not unusual in nomenclature, this
name, the latest of all, got complete dominance over the
others, and has been universally used to the present time.
No doubt the wide utilization of Pallas’s well-known ‘ Glires’
was the cause of the mistakes involved.
Putting aside sungorus and barabensis (furunculus) as not
of the “pheus’’ group, we may first accept without hesi-
tation Dr. Satunin’s opinion § that accedula (i. e. migra-
torius) is the same as pheus, both being from the same
region of Southern Russia. This acceptance, however,
involves the use of the senior name, migratorius, and the
consequent disappearance of pheus. Dr. Satunin also
considers arenarius, from the Irtish, as the same species.
On laying out the whole Museum series assigned to
“ nhaus,” rather more than 80 in number, I find that while
the Central Asiatic forms are, as a general rule, larger, with
larger teeth and longer hair than those from the west, yet
that both skull and teeth vary in series of each form to such
an extent as tooverlap each other. I therefore provisionally
* Reise, ii. pp. 703-704 (1773).
+ Sungorus cannot be treated as a misprint for songarus, as has
commonly been done, for it not only occurs both in text and plate, but is
also used in the same form for another animal on p. 7380.
t-P. 8641779).
§ Mittheil. Kaukas. Mus. ii. p. 340 (1906).
rejyerred to Cricetulus pheeus and campbelli. 453
combine them all (except a set from Ladak) as one species,
which, as shown above, should bear the name migra-
torius.
The status of the Grecian form, Cricetulus atticus, and
that of Palestine, to which the name cinerascens* applies,
I do not propose at present to discuss, but I confess I doubt
very much if either of them deserves specific distinction
from C. migratorius.
Putting aside these, the subspecies of C. migratorius
which may be recognized appear to be as follows :—
C. migratorius migratorius, Pall.
Synn. accedula, Pall., pheus, Pall., eversmanni, Brandt.
Type-locality. R. Ural.
Range. 8. Russia, Caucasus region, Transcaspia, Persia,
and Asia Minor south of the coastal forest-region of the
north-east.
Size comparatively small; skull about 27 mm. in length,
its upper tooth-row 4 mm., the teeth themselves narrow
and light. Fur short, hairs of back about 9 mm. in length.
Colour above pale grey, generally with indistinct median
darker dorsal shading. Hairs of under surface with grey
bases, except that they are wholly white on an area on the
throat, which extends in a median point down to between
the fore legs.
C. migratorius vernula, subsp. n.
Type-locality. Khotz, near Trebizond.
Range no doubt all the forest coastal strip along the
south-east corner of the Black Sea.
Size averaging slightly larger than in true migratorius.
Fur of back 8-9 mm. in length. Colour darker through-
out, the upper surface near “ mouse-grey,” the median dark
dorsal area faintly evident. Hairs of under surface with a
shorter length of white above the slaty, and the white itself
not so pure, but faintly suffused with buffy ; white throat-area
less extended towards the chest, ending about halfway
towards the fore limbs. Ears with proectote markedly
blackish. Tail distinctly darker above, at least for its basal
half,
Dimensions of type :—
Head and body 115 mm.; tail 82; hind foot 17; ear 17°5.
Skull: greatest length 28:2 ; condylo-incisive length 26 ;
palatal foramina 5; upper molar series 42.
* Hypudeus cinerascens, Wagn. Wiegm. Arch, 1848, p. 184.
454 Mr. O. Thomas on the small Hamsters
Type. Adult female. B.M. no. 6.5.1.83. Original
number 2443. Collected 25th February, 1906, by A. Robert.
Presented by Oldfield Thomas. Seven skins and six speci-
mens in spirit. ;
A darker and more saturate race of C. migratorius, a
variation which occurs in other forms from the same district.
Its range eastwards iuto Trans-Caucasia remains to be
worked out, but a specimen in spirit from the Talysch Mts.
appears to be very similar to it. To the south, a specimen
from ‘60 miles north of Erzeroum” (Woosnam) would
also seem to be referable to the same species.
C. migratorius arenarius, Pall.
Type-locality. Baraba Steppes, Lower Irtish, Siberia.
Southwards to Lake Balkhash.
Under surface wholly white, the hairs white to their
bases.
A spirit-specimen from Mt. Bek Dauata, north of Lake
Balkhash (W. Bateson), has head and body 94 mm., tail 26,
hind foot 15. Its skull measures 28°5 mm. in total length,
its tooth-row 4°2 mm.
The Museum only contains the above-mentioned spirit-
specimen, whose agreement with Pallas’s description as to
the whiteness of the under surface leads me to assign it to
the true arenarius, and to consider the next form, of which
we have a large number of specimens, as distinct.
C. migratorius griseiventris, Sat.
Type-locality. R. Bis-shen-gol, Altain-nuru, Gobi Altai.
Range. Central Asia. Samarkand and eastwards along
the Thian Shan, filling up the area between the ranges of
arenarius and fulvus.
Size averaging larger and teeth heavier than in the
western forms. Fur longer—hairs of back 10 mm. or more.
General colour above pale grey, more or less strongly
suffused with buffy or fulvous. Under surface with the
hairs snowy white to the bases on throat and inguinal
region, but broadly slaty at base on the belly.
A male from Djarkent measures:—Head and _ body
120 mm.; tail 27; hind foot 16. Skull: greatest length
30; upper tooth-row 4:5.
Of this form the Museum possesses a fine series from
Djarkent, collected by W. Riickbeil, besides specimens
from Samarkand, Dzungaria, Hami and elsewhere in the
Carruthers collection. It differs from C. m. arenarius by
the presence of broad slaty bases on the belly hairs,
referred to Cricetulus pheus and campbelli. 455
C. migratorius fulvus, Blanf.
Type-locality. Kashgar.
Range between 35° and 41° N., 74° and 80° E., so far as
our specimens show, but probably extending considerably
further, especially towards the east.
Like C. m. griseiventris in all respects except that the
general colour is a little paler; the white of the under
surface passes higher up on the sides, and is less modified by
slaty bases, which are only present, and then very incon-
spicuously, on the centre of the belly.
Mr. Holmes-Tarn collected some specimens of this beauti-
ful little Hamster on the Karakash River, Chinese Turkestan,
and Mr. Carruthers an example on the north side of the
Karakoram Mts. The original specimens were obtained
during the Yarkand Mission, and the type is in Calcutta.
Subspecies fulvus and griseiventris are no doubt very
closely allied, but may apparently be distinguished by the
darker colour and greater amount of slaty on the belly of
the latter.
In the nearly or quite pure white belly fulvus agrees with
the typical arenarius of much further north, though the two
appear to be separated by the range of griseiventris. Speci-
mens from many further localities will be needed before the
exact relations of the three can be determined.
Although I provisionally accept Dr. Satunin’s view as to
the Central Asian Hamsters belonging to the same species
as C. migratorius, | think it not improbable that they really
ought to be divided into two species—a small-toothed one,
of which migratorius and vernula, and perhaps arenarius,
would be subspecies ; and a large-toothed Asiatic one, con-
sisting of the subspecies fulvus and griseiventris.
But the following animal must in any case be distin-
guished as a species :—
Cricetulus alticola, sp. n.
General characters and colour above about as in C. m.
fulcus, but the hairs of the under surface broadly slaty at
base, even on chest and throat, those of the chin being alone
white to their roots. Ears grey, not darkened on the pro-
ectote. Tail heavily haired, wholly white.
Skull distinguished from that of all forms of C. migra-
torius by its conspicuously smaller bulle, small both verti-
cally and horizontally. Palatal foramina comparatively
long, parallel-sided. ’
456 Mr. O. Thomas on small Hamsters.
Dimensions of type (measured in the flesh) :—
Head and body 98 mm. ; tail 381 ; hind foot 15:5; ear 15.
Skull: greatest length 27; condylo-incisive length 24;
palatilar length 11:3; palatal foramina 5°5; breadth of
bulla at right angles to its greatest diameter 3°3; upper
tooth-row 3:9.
Hab. Ladak. Type from Shushal, 13,500’ ; other speci-
mens from Durgu Vil and Khardong (Crump), and Teza,
Upper Sutlej Valley (Whitehead).
Type. Adult male. B.M. no. 6.10.3.18. Original
number 115. Collected 29th June, 1906, by C. M. Crump,
and presented to the National Museum by Col. Ward.
Four specimens in all.
This Hamster has a quite extraordinary resemblance to
the pale-coloured Voles of the genus Alticola (e. g.,
A. stracheyi, cricetulus, or phasma), with which it agrees in
proportions, length and colour of fur, and external appear-
ance generally ; it even has, unlike C. migratorius, their slaty-
_ based throat-hairs. In fact, the only points that show it is
not a Vole are the shorter and more “ pudsy ” feet and the
finely-haired untufted ears—neither of which affect the
general appearance.
C.m. fulvus is also very like Alticola phasma above, but
the resemblance is spoilt below by the nearly wholly white
under surface. :
This Ladak Hamster is probably most nearly allied to the
Tibetan C. lama, B.-Ham., but is considerably smaller and
has a shorter tail.
With regard to the systematic-arrangement of the small
Hamsters, I agree with Mr. Miller as to the advisability of
generically separating my Cricetulus bedfordie from the
others (genus Phodopus, Miller), and would, indeed, go
further by also distinguishing the species sungorus and
campbelli, which have an intermediate condition, both of
foot-structure and dentition. The new genus might be
called :—
CRICETISCUS, g. Nn.
Soles densely hairy, the three posterior of the six normal
pads completely suppressed, and the three distal ones very
small, hidden in the hair. Teeth neither so complicated as
those of Cricetulus, in whieh there is a broad notch, and
commonly a deep pit, between the outer and inner main.
cusps of each lamina, nor so simplified as those of Phodopus,
in which the notch is reduced and the pit is absent.
' Genotype: C. campbelli. (Cricetulus campbelli, Thos.)
On new Pyralide of the Subfamily Hydrocampina. 457
Other species: C. sungorus. (Mus sungorus, Pall.)
Although C.sungorus was known so much the earlier, it is
only represented in the material available to me by a single
dried skin, in which the characters are not very clearly dis-
cernible. With both skins, skulls, and a spirit-specimen of
C. campbelli for examination, I therefore think it advisable
to make that species the genotype.
The position of Satunin’s Cricetulus roborovskit is not
clear, as his expression “ Die Sohlen sind dicht mit weissem
Haar bekleidet”’ is applicable either to a Phodopus or a
Cricetiscus.
XLV.—Descriptions of New Pyralidie of the Subfamilies
Hydrocampine, Scopariane, &c. By Sir Georee F.
Hampson, Bart., F.Z.S., &c.
(Continued from p. 376. ]
(12) Cataclysta nyasalis, sp. n.
Head, thorax, and abdomen white mixed with dark brown witha
cupreous gloss; antenne brown ringed with white; palpi white,
black-brown above; fore and hind femora and tibie black-brown
above; claspers of male yellowish. Fore wing with the base white,
brown at costa; an inwardly oblique orange-yellow antemedial
band from cell to inner margin, defined by brown lines and with
white beyond the outer line; the medial area white very thickly
irrorated with dark cupreous brown; postmedial line white, slightly
incurved below vein 3 and ending at tornus, an orange-yellow band
beyond it from below costa to vein 8; the terminal area brown with
a white line from costa before apex to termen at vein 4, an orange-
yellow band beyond it on termen from apex to discal fold; cilia
dark brown with some white at tips. Hind wing white, very
thickly irrorated with dark cupreous brown from near base to the
subterminal line, which is black-brown defined on each side by
white, slightly incurved below vein 2; five partly conjoined ocellate
black spots on termen from below apex to above tornus, with
metallic silvery points between them, the anal spot larger with the
silvery point in its centre, some orange-yellow points beyond them
on termen; cilia cupreous brown at base, white slightly tinged
with brown at tips. }
Hab. Br. C. Arrica, Mt. Mlanje (Neave), 3 5, 1 Q type.
Exp. 10 mm.
*
Ann. & Mag. N. Hist, Ser. 8. Vol. xix. 30
458 Sir G. F. Hampson on new
(2a) Cataclysta atriterminalis, sp. n.
@. Head, thorax, and abdomen pale ochreous mixed with some
silvery white; palpi with the 8rd joint brownish ; pectus, legs, and
ventral surface of abdomen white. Fore wing white, the costal
area brownish ochreous ; an inwardly oblique orange-yellow ante-
medial band from discal fold to inner margin, defined by brownish
lines ; a short orange- ‘yellow fascia beyond the cell; brownish sub-
terminal spots at veins 6 and 2 and above tornus. Hind wing
white; an oblique orange-yellow medial band edged by brownish
lines, the yellow extending on inner margin to near tornus; the
terminal area broadly black with some whitish scales and some
orange scales on termen.
Hab. Cutepes, Saugir I. (Doherty), 1 2 type. Exp. 10 mm.
(47 a) Cataclysta obliquifascia, sp. n.
@. Head and thorax fulvous yellow mixed with grey-brown ;
abdomen brownish white, dorsally tinged with ochreous yellow
except towards base and with dark ‘brown segmental lines ; antenne
dark brown except at base; palpi yellowish, the 3rd joint dark
brown; pectus, legs, and ventral surface of abdomen white tinged
with brown, the fore femora towards extremities, the tibize and
tarsi dark brown, the tibiz banded with yellowish, the mid tibiz
with dark brown stripe above, the tarsi ringed with dark brown.
Fore wing fulvous yellow; small subbasal and antemedial brown
spots on costa and slight brown marks on inner margin before and
at middle; an obliquely curved reddish-brown fascia from costa
before apex to middle of discocellular, with a whitish fascia below
it; an oblique rather diffused reddish-brown line from before
termen at discal fold to inner margin before tornus, with some
whitish beyond it; a fulvous-yellow terminal band defined on inner
side by a reddish-brown line; cilia white at base, brown at tips.
Hind wing golden yellow, the costal area to below the cell and to
beyond middle white; an oblique dark brown medial band from
cell to inner margin, and a rather diffused oblique postmedial line
from vein 4 to inner margin ; an incurved subterminal brown line
from costa to discal fold; a series of small black-brown spots before
termen from below apex to vein 2, some metallic silver beyond
them on termen ; cilia yellow at base, with black line at middle,
the tips white tinged with brown.
Hab, Brazit, Pernambuco (Swale), 1 9 type. Hap. 16 mm.
(51) Cataclysta chionostola, sp. n.
@. Head and thorax white slightly tinged with rufous; abdo-
men white ; antenne brownish except towards base ; pectus, legs,
and ventral surface of abdomen white, the fore legs slightly tinged
with rufous and the tibiz with black-brown streak below. Fore
Pyralide of the Subfamily Hydrocampine. 459
wing silvery white; two cupreous-brown points at base; ante-
medial line cupreous brown, oblique to above vein 1, then inwardly
oblique, a patch of pale yellow before it below the cell; a very
oblique cupreous-brown line from middle of costa to below sub-
median fold well beyond middle, then recurved to inner margin, an
oblique striga beyond it in end of cell and another beyond the
cell, with a yellowish tinge between them ; a triangular postmedial
patch of the ground-colour from costa to below vein 4, defined at
sides by yellow-brown lines ; two cupreous-brown subterminal lines
with a yellowish tinge between them, from just below costa to just
above inner margin, incurved below vein 3; a cupreous-brown line
before termen. Hind wing silvery white; a faint yellow and
brown tinge in and below middle of cell; a straight oblique
cupreous-brown medial line ; postmedial line cupreous brown, in-
curved between veins 6 and 2; two cupreous-brown subterminal
lines with a yellowish tinge between them, the inner line slightly
incurved between veins 6 and 2, the outer angled outwards at
vein 6, then slightly waved and joining the inner line at submedian
fold ; a cupreous-brown subapical striga and a line before termen
from the angle of the outer subterminal line to vein 2.
Hab. Perv, Rio Pacaya, 1 2 type. Hxp. 18 mm.
(4a) Ambia chrysogramma, sp. n.
Head, thorax, and abdomen white, the last slightly tinged with
brown on dorsum; antennz with an ochreous tinge. Fore wing
silvery white, the costa tinged with orange-yellow towards base; a
small tuft of rufous scales below middle of costa with the orange-
yellow antemedial and medial lines arising below it and rather
diverging towards inner margin; a conical postmedial patch
defined by orange-yellow from costa to vein 5, with a brown point
on the line defining its outer side at costa; an orange-yellow sub-
terminal line, exeurved to submedian fold, where it is angled in-
wards, a fine yellow-brown line beyond it, incurved below vein 2,
the terminal area tinged with yellow. Hind wing silvery white; a
small orange-yellow discoidal spot; an orange-yellow postmedial
line, excurved to vein 4, then bent inwards to origin of vein 2 and
oblique to inner margin; an orange-yellow subterminal line, ex-
curved to vein 2, then incurved, a fine yellow-brown line beyond it;
the terminal area tinged with yellow.
Hab. Samoa Is., Pago (de la Garde), 1 5,1 Q type. Exp.
12 mm.
(9b) Ambia cymophoralis, sp. n.
3. Head white, the antenne tinged with ochreous, the palpi
yellow behind ; thorax and abdomen white, the tegule with sub-
dorsal orange-yellow stripes, the patagia orange-yellow above and
the dorsum of thorax orange-yellow except the metathorax, the
abdomen banded with orange-yellow; pectus, legs, and ventral
30*
460 Sir G. F. Hampson on new
surface of abdomen white, the fore femora above and tibiz on inner
side yellowish with some black at the femoro-tibial joint. Fore
wing orange-yellow with a fulvous tinge; some white at base in
and below the cell; an erect silvery-white subbasal band ; a silvery-
white band just before middle, defined on each side by dark brown
below the cell, excurved below the costa and above inner margin
and emitting a spur at discal fold to the white discoidal lunule
defined by black except above; the medial part of costa white; a
silvery-white wedge-shaped mark in discal fold before the post-
medial band, which is silvery white defined on each side by dark
brown, incurved below costa, then excurved to vein 3, below which
it is angled inwards, then erect with its outer edge excurved at
submedian fold; a silvery-white subterminal band from costa to
vein 1, defined on each side by dark brown, strongly on outer side,
its extremities at costa and above vein 1 dilated into spots, ex-
curved between those points; cilia white, chequered with brown at
apex and with orange-yellow at middle and tornus. Hind wing
orange-yellow with a slight fulvous tinge, the base white ; a silvery-
white antemedial band from cell to inner margin connected with a
silvery-white patch in end of cell with a black discoidal bar on its
outer edge ; a silvery-white postmedial band, excurved and defined
on outer side by brown to vein 2, then incurved; a silvery-white
subterminal band defined on each side by black from costa to vein 1,
its extremities on costa and above vein’1 expanding into spots,
excurved between those points; cilia white, chequered with red-
brown towards apex and at middle.
Hab. LovrsiavE Is., St. Aignan I. (Week), 3 3 type, Rossel I.
(Meek), 1 9. Hap. 16-18 mm.
(16a) Ambia rufitincta, sp. n.
@. Head, thorax, and abdomen white suffused with rufous, the
last slightly irrorated with dark brown towards base and strongly
with black towards extremity ; antenne white tinged with yellow ;
palpi white tinged with yellow and with some black above ; pectus,
legs, and ventral surface of abdomen yellowish white. Fore wing
white suffused with rufous and slightly irrorated with brown ; Pf
faint inwardly oblique rufous antemedial line; a curved black
medial line, rather diffused on inner side and incurved at median
nervure ; an oblique slightly curved blackish postmedial line from
costa to vein 2 above tornus, defined on outer side by white ; a fine
brown terminal line from costa to vein 2. Hind wing white
suffused with rufous and slightly irrorated with brown except on
inner area ; a patch of diffused black scales in and beyond end of
cell; postmedial line black, oblique and defined on outer side by
white to vein 4, then obsolete and retracted to beyond lower angle
of cell, then curved and rather diffused to inner margin; a fine
brown terminal line except toward tornus.
Hab. Perv, Cuzco Mts. (Garlepp), 1 Q type. Ep. 14 mm.
Pyral.de of the Subfamily Hydrocampine. 461
(17 a) Ambia hemigrammalis, sp. n.
®. Head, thorax, and abdomen white mixed with fulvous
yellow and irrorated with some black scales; palpi white banded
with black; pectus, legs, and ventral surface of abdomen white,
the fore tibize banded with black. Fore wing white; a subbasal
black bar from costa; an antemedial black line from costa to discal
fold and some scales at inner margin; a broad oblique orange-
yellow shade from below costa before the postmedial line to inner
margin before middle, irrorated with some black scales and with
black striz on it on each side of the discocellulars ; a strong oblique
black postmedial line from costa to discal fold, then an incurved
shade formed by blackish scales with the area beyond it orange-
yellow; an orange-yellow bar from costa before apex, thena double
eurved black line filled in with silvery white; the terminal area
orange-yellow, narrowing to a point below apex; cilia white, black
at base at apex and with a black patch between veins 4 and 2.
Hind wing white, the basal area irrorated with black and with a
black patch at end of cell; the terminal half suffused with orange-
yellow and slightly irrorated with brown; a dark brown shade
between veins 7 and 2 before the indistinct double dark postmedial
line filled in with white ; a narrow white band defined by slight
dark lines before the termen which is yellow, the band not ex-
tending to tornus; cilia white, dark brown at base from vein 4 to
tornus.
Hab. PEru, Yungas de la Paz (Seebold), 12 type. xp.
12 mm.
(17 6) Ambia sufetulodes, sp. n.
©. Head, thorax, and abdomen white mixed with dark brown,
the prothorax and patagia at middle with brown spots, the abdomen
suffused with brown towards extremity, leaving white segmental
lines ; palpi and maxillary palpi white banded with black; pectus,
legs, and ventral surface of abdomen white, the fore tibize, spurs,
and tarsi banded with black. Fore wing white, the basal and
terminal areas suffused with brown, the medial area with broad
brownish shade; a white subbasal band, defined on inner side by
black ; a white antemedial band, defined on each side by black and
incurved below submedian fold; two semicircular white marks
defined by black on medial part of costa, the first with black point
at costa; a small round black discoidal spot; a postmedial white
band defined on each side by black, excurved to vein 5, then in-
curved, expanding somewhat at costa; an oblique white streak
from apex and a blackish line before termen excurved at middle ;
a dark terminal line except at the excision at discal fold; cilia
white with some brown at apex and middle. Hind wing white, the
terminal area suffused with brown, broadly towards costa and
narrowing to tornus; some brown near base; a broad oblique
462 Sir G. F. Hampson on new
brown band from discal fold before the small blackish discoidal
spot to inner margin before the postmedial line, with a white bar
on it at inner margin; postmedial line black defined on outer side
by white, incurved below discal fold, then excurved; a black line
before termen ; a black terminal line from apex to vein 3 except at
the excision at discal fold, and black strie# at veins 2 and 1; cilia
white with some brown at apex and middle.
Hab. Peru, R. Pacaya, 1 9 type. Harp. 14 mm.
(226) Ambia fulvicolor, sp. n.
o. Head, thorax, and abdomen yellow suffused with fulvous,
the last with subdorsal white segmental bands ; frons and 8rd joint
of palpi white ; pectus, legs, and ventral surface of abdomen white
tinged with fulvous. Fore wing yellow suffused with fulvous; an
oblique subbasal silvery-white band from cell to inner margin, with
some red-brown before it; some dark brown on costa before the
antemedial silvery-white band, which is interrupted in the cell,
oblique towards costa and below the cell and defined by red-brown ;
the cell suffused with red-brown except towards base; a fulvous
lunule at end of cell defined by dark brown and with some white
beyond it; the fovea above end of cell white defined by dark brown
and with a silvery-white point above it on costa; some dark brown
on.costa before an oblique silvery-white postmedial band from costa
to vein 4 and a triangular mark from vein 2 to inner margin, both
defined on outer side by the dark brown postmedial line which is
angled inwards at vein 2, the costa beyond it dark brown; a
slightly sinuous dark brown subterminal line with a series of small
silvery-white spots before it from below costa to inner margin, the
hair on which is dark brown below it; cilia white mixed with some
yellow and chequered with dark brown below apex and at veins 4,
3, 2. Hind wing yellow suffused with fulvous along median ner-
vure and on terminal area, the base white; an oblique silvery-
white antemedial band defined by dark red-brown; a fulvous
discoidal spot defined by dark red-brown; postmedial line dark
brown defined on inner side by a silvery-white band, slightly in-
curved below vein 4; a sinuous dark brown subterminal line
defined on inner side by silvery-white spots, small to vein 5, then
interrupted to just above vein 8, larger and more diffused below
vein 38; cilia white mixed with some yellow, chequered with dark
brown at apex and between veins 5 and 2.
Hab. Br. N. Guinea, Kumusi R. (Meek), 1 3 type. Hap,
18 nm.
(23 6) Ambia albiflavalis, sp. n.
3. Head, thorax, and abdomen white tinged with yellowish;
antenne ochreous ; frons yellow; palpi yellow with the 3rd joint
white; pectus, legs, and yentral surface of abdomen white. Fore
Pyralide of the Subfamily Hydrocampine. 463
wing silvery white; the base orange-yellow with oblique outer
edge; an obliquely curved orange-yellow antemedial band; an
orange-yellow band from end of cell to inner margin, the end of
cell tinged with brown and the fovea above it with two brown
points on its upper edge, a yellow patch with a white spot on it
beyond it on costal area; an orange-yeilow subterminal band
detined at sides by brown, obliquely curved to vein 2, then bent
outwards to tornus, giving off on inner side between veins 4 and 2
a yellowish fascia tinged with brown to lower end of cell; a pale
brown terminal band. Hind wing silvery white ; an orange-yellow
antemedial band from cell to inner margin; a curved orange-yellow
postmedial band detined by red-brown from costa to vein 1, its
outer edge angled outwards at vein 4; a sinuous orange-yellow
subterminal band defined by red-brown and ending at tornus, its
outer edge excurved at discal fold to the narrow orange-yellow
terminal band defined on inner side by a red-brown line and ending
at submedian fold.
Hab. 8. Nigeria, Lagos (Dudgeon), 1 d type. Exp. 14 mm.
(23 f) Ambia niveiplagalis, sp. n.
do. Head, thorax, and abdomen white mixed with red-brown
-and dark brown, the head with dark line between antennz, the
tegule dorsally and patagia at middle with white patches, the
abdomen -pale towards extremity and with white bands; antennz
yellowish ringed with black ; palpi pale red-brown, the 8rd joint
white with red-brown band towards extremity; pectus, legs, and
ventral surface of abdomen white, the fore legs tinged with red-
brown, the abdomen with faint brownish bands. Fore wing
yellowish tinged with rufous and irrorated with dark brown; an
inwardly oblique blackish antemedial line, excurved above inner
margin and with a dark shade before it from subcostal nervure to
inner margin; an oblique slightly sinuous blackish medial line with
a white striga before it in and below the cell, where it is conjoined
to a silvery-white patch below the submedian fold extending to
inner margin and to the antemedial line ; a quadrate silvery-white
spot beyond the cell defined by blackish and a quadrate patch from
vein 2 to inner margin defined by blackish at sides, the fovea above
end of cell white ; an oblique elliptical silvery-white patch defined
by blackish from costa, to which it narrows, to vein 4; an apical
white spot, then a curved series of white spots defined by blackish,
minute to vein 3, the spots. below veins 3 and 2 larger; cilia red-
brown mixed with white. Hind wing yellowish tinged with rufous
. and irrorated with dark brown ; a broad subbasal silvery-white band
defined by dark brown and with red-brown spot on it at inner
margin; a rather lunulate silvery-white spot beyond the cell defined
by dark brown; a rounded postmedial silvery-white patch defined
by dark brown from costa to vein 4, a spot below vein 3, and a
eurved band from vein 2 to above tornus; a triangular silvery-
464 Sir G. F. Hampson on new
white apical spot, small conjoined subterminal spots above and
below vein 4and larger rather wedge-shaped spots below veins 3
and 2; cilia reddish brown mixed with some white.
Hab. Perv, Carabaya, Oconeque (Ockenden), 1 3 type. Hap.
20 mim.
(806) Ambia melanistis, sp. n.
2. Head, thorax, and abdomen black-brown mixed with some
white; pectus, legs, and ventral surface of abdomen white. Fore
wing very dark red-brown with a blackish tinge; an antemedial
white point on costa and medial bar from costa; a slightly ex-
curved punctiform white postmedial lme from costa to discal fold,
the line then almost obsolete and incurved below vein 4, with white
points above and below vein 1, a metallic silvery patch before it
between veins 3 and 1 and a small spot at middle of inner margin ;
a curved punctiform white subterminal line; cilia whitish. Hind
wing very dark red-brown with a blackish tinge ; the basal part of
costa white ; an antemedial white point on inner margin; a small
metallic silvery spot beyond the cell; an oblique postmedial white
band from costa to vein 4 and a minute spot above tornus; a
curved white subterminal line from costa to discal fold and a series
of stria between discal fold and vein 1; the termen more rufous;
cilia white.
Hab. Formosa, Kanshirei (Wileman),1 9 type. Hap.12 mm.
§(38 a) Ambia argentistriata, sp. n.
Fore wing of male on underside with large costal fold from
before middle to near apex; hind wing in both sexes on upperside
with tuft of long spatulate hairs below end of cell.
3. Head, thorax, and abdomen yellow, the tegule and patagia
with scarlet streaks, the abdomen dorsally suffused with rufous ;
palpi tinged with rufous; pectus, legs, and ventral surface of abdo-
men white tinged with rufous. Fore wing orange-yellow; the
basal area suffused with scarlet, irrorated with a few black scales
and with some silver scales below costa; a curved scarlet ante-
medial line; the medial area except towards costa and the post-
medial area below vein 4 with a silvery gloss finely striated with
dark brown; a scarlet line from upper angle of cell, oblique to
vein 8, then erect, with a yellow streak tinged with scarlet in sub-
median fold to the antemedial line and beyond it on the silvery
area, the area beyond the line, except the silvery area, yellow tinged
with scarlet with brilliant silver streaks defined by black scales in
the interspaces of the postmedial area from below costa to vein 4
and a streak of black scales below vein 4, its extremity connected
by a line formed by black scales with the inner margin and in-
curved at submedian fold; a curved brilliant silver line before
termen and a terminal series of black strie ; cilia with brown mixed
oe
Pyralide of the Subfamily Hydrocampine. 465
except at base. Hind wing white; a triangular area from origin
of vein 2 to termen between discal fold and vein 2 orange-yellow ;
the tuft of scales below end of cell black; a curved scarlet post-
medial line between veins 5 and 2; a brilliant silver line before
termen from costa to vein 5, then a series of small brilliant silver
spots with black points before and beyond them on each side of veins
4to 2; the termen narrowly orange-yellow to apex ; cilia white, their
bases orange-yellow with a brown line at base to vein 2.
Ab. 1. Head, thorax, abdomen, fore wing, and the triangular
patch on hind wing yellowish suffused with rufous, the scarlet
markings replaced by rufous, the fore wing with silvery marks
before the antemedial line in the interspaces.
Hab. W. Couomsta, Jiminez, 1 ¢ type, W.slopes,1 5. Eup.
20 mm.
(88 ¢) Ambia pheomeralis, sp. n.
@. Head, thorax, and abdomen grey-white mixed with brown ;
palpi, pectus, legs, and ventral surface of abdomen white tinged
with brown, the tarsi dark brown ringed with white. Fore wing
grey-white thickly irrorated with brown; an indistinct brown
medial line defined on inner side by whitish, excurved to median
nervure, then sinuous; a slight dark discoidal spot ; a large patch
of dark brown suffusion from costa to vein 2 before the postmedial
line which is white slightly defined on outer side by brown and
excurved from costa to vein 2 near termen, then almost obsolete
and retracted to below the angle of cell, then more distinct, defined
on inner side by brown and waved to inner margin; a series of dark
strize before termen froin below apex to above tornus. Hind wing
white thickly irrorated with dark brown, the base, cell, and costal
area to near apex white; the tuft of hairs below end of cell black-
brown ; subterminal line white, excurved to vein 2, then sinuous;
a series of ocellate black spots on termen from apex to submedian
fold, defined on inner side by white with a fine sinuous dark line
before it and with slight orange marks between them, the spots
from apex to discal fold minute, then larger and with a black line
beyond them on termen ; cilia white, metallic silver at base.
Hab. Boxtvia, Yungas de la Paz (Seebold), 1 2 type. Eup.
22 mm,
(4a) Oligostigma centrimacula, sp. n.
Antenne of male fringed with hair above at one-third from base ;
hind tibiz fringed with hair below towards base,
Head and thorax white, the head tinged with rufous behind, the
patagia with some black-brown on outer edge; abdomen white,
suffused with pale yellow except towards extremity; antennz
tinged with rufous; palpi with some black on extremity of 2nd
joint behind; pectus, legs, and ventral surface of abdomen white,
466 Sir G. F. Hampson on new
the fore femora above and tibie on inner side black, the tarsi ringed
with black. Fore wing orange-yellow, the costa red-brown to end
of cell; a white fascia in the cell from before middle extending to
well beyond the cell and with the dark cupreous red-brown dis-
coidal spot on it; a curved silvery-white subterminal band defined
on outer side by a black line from below costa to submedian fold in
which it is bent inwards as a streak; a fine brown terminal line;
cilia white. Hind wing with the basal half white with oblique
outer edge defined by a strong black-brown line between discal and
submedian folds; the terminal half orange-yellow with oblique
silvery-white subterminal bar from costa to vein 6 and subterminal
silvery-white band defined by dark brown lines between veins 4 and
1; four minute rather quadrate black spots on termen between
vein 5 and submedian fold; cilia white, metallic silvery at base
beyond the spots.
Hab. QUEENSLAND, Kuranda (Dodd), 1 3, 19 type. Hzp.
16 mm.
(Le) Oligostigma peruviensis, sp. n.
Head and thorax dark brown mixed with white, the patagia with
white streak on upper edge towards base, the metathorax edged
with white ; abdomen brown with white segmental bands; antennz
white ringed with brown; palpi brown with some white in front
and at tips; pectus, legs, and ventral surface of abdomen white,
the legs tinged with brown, the abdomen tinged with brown except
towards base and with white segmental lines. Fore wing cupreous
brown mixed with some ochreous; an obliquely curved somewhat
dentate blackish subbasal line defined on each side by slight white
marks except towards costa; antemedial line white defined on each
side by dark brown, oblique to median nervure, angled outwards at
median nervure and submedian fold, then very oblique, forked
white streaks beyond it at submedian fold; the medial area with
white spots on costa, in the cell, and at inner margin; a white
- medial line, oblique to submedian fold, then excurved; a rather
lunulate black discoidal spot defined by ochreous ; the postmedial
area with a loop formed by sinuous white lines from costa to
vein 2, enclosing a white band to vein 4 rather constricted below
costa ; a narrow white band from below costa to vein 1, its inner
edge irregular and incurved at discal and submedian folds before an
orange-yellow terminal band defined on inner side by a black line
and on outer by black striz on termen; cilia white with a brown
line near base and the tips with brown mixed. Hind wing cupreous
brown ; a sinuous white subbasal band; a white antemedial line;
a broad white medial band ; postmedial line white, angled inwards
below costa and outwards at vein 4, then bent inwards and sinuous
to inner margin; a white band with sinuous inner edge from discal
to submedian fold before the terminal orange-yellow band defined
on inner side by a black line; a series of five small ocellate black
Pyralids of the Subfamily Hydrocampine. 467
spots on termen from vein 7 to below 4, the spots below veins 7
and 5 double, each with a white point on them; cilia white mixed
with brown and with a brown line near base.
Ab. 1. Fore wing without the white in end of cell or in the
postmedial loop.
Hab. Peru, Carabaya, R. Huacamayo, La Union (Ockenden),
13,19, La Oroya (Ockenden), 23, 3 2 type. Exp., g 22-
26, 2 32-88 mm.
(1d) Oligostigma rufiterminalis, sp. n.
2. Head and thorax greyish suffused with red-brown ; abdomen
whitish tinged with red-brown and slightly irrorated with black ;
palpi white tinged with red-brown and irrorated with black ; pectus,
legs, and ventral surface of abdomen whitish, the legs tinged with
red-brown, the fore tibiz with white band at middle and black
band at extremity. Fore wing greyish suffused with red-brown ;
blackish points near base in and below the cell; a rather diffused
black antemedial line, excurved to below vein 1 and bent inwards
to inner margin ; the medial area with rufous streaks irrorated with
black in discal and submedian folds ; a somewhat inwardly oblique
black medial line, slightly excurved below costa; an elliptical
rufous discoidal spot defined by black; postmedial line black
slightly defined on outer side by whitish, very slightly waved
towards costa, then excurved to vein 2, where it is strongly angled
inwards, then oblique; a narrow silvery-white band from below
costa to vein 1, above which it forms a small spot with a deeper
red-brown shade before it, before the narrow orange-yellow terminal
band defined on inner side by a black line; cilia whitish tinged with
red-brown. Hind wing whitish tinged with red-brown, the area
beyond the postmedial line rufous ; a curved black subbasal line; a
round yellow discoidal spot defined by black; postmedial line
black, excurved to vein 3, then incurved; a narrow silvery-white
subterminal band defined on outer side by a black line with some
yellow beyond it; the termen narrowly whitish between veins 6
and 2, with four minute black spots on it; cilia whitish tinged
with red-brown.
Hab. MavacascaRr, 1 2 type. Hwvp. 24 mm.
(12a) Oligostigma piperitalis, sp. n.
2. Head and thorax grey tinged with rufous and irrorated with
dark brown; abdomen white with diffused rufous bands; antenne
dark brown, pale red-brown towards base ; palpi red-brown ; pectus,
legs, and ventral surface of abdomen white, the fore and mid legs
tinged with red-brown. Fore wing grey tinged with rufous and
thickly irrorated with dark brown; a short blackish streak in
middle of cell and small rather diffused discoidal spot; a narrow
white band from below costa to vein 1 before the fulvous-yellow
468 Sir G. F. Hampson on new
terminal band defined on inner side by a fine dark line; a fine dark
terminal line; cilia whitish tinged with rufous and with a fine dark
line near base. Hind wing pale rufous irrorated with dark brown ;
the base white; a narrow oblique white antemedial band, diffused
outwards at costa; a small blackish discoidal spot; a narrow white
postmedial band excurved to vein 4, then incurved ; a narrow white
band defined on inner side by diffused blackish and on outer by a
black line from below costa to inner margin before the fulvous-
yellow terminal band; a white terminal line with minute black
spots on termen below veins 7, 5, 4, 8, defined on inner side by a
black line which is slightly waved before the spots, the termen with
a fine black line at apex and below the spots; cilia white faintly
tinged with brown.
Hab. N. Nigeria, Zungeru (Macfie), 1 2 type. Exp. 18 mm.
(186) Oligostigma flavialbalis, sp. n.
2. Head, thorax, and abdomen white, the shoulders with some
rufous, the metathorax yellow behind; palpi and maxillary palpi
banded with rufous; fore tibize tinged with rufous with a white
spot at middle. Fore wing silvery white, the costa tinged with
rufous at base, then with yellow to beyond middle, the inner margin
with a pale yellow patch before middle; a small rufous-spot in
upper part of middle of cell; a yellow discoidal spot defined by
rufous scales; a silvery-white subterminal band defined by pale
brown lines with diffused pale yellow before it and the terminal
area beyond it pale yellow; cilia white tinged with rufous. Hind
wing white; the subbasal area pale yellow, diffused on inner side
and defined on outer bya pale brown line; a pale brown postmedial
line, excurved below costa, the area beyond it pale yellow with a
narrow silvery-white subterminal band on it; the termen with
minute black bars at veins 6 to 2; cilia white tinged with rufous.
Hab, Mapacascar, Betsileo (Cowan),1 2 type. Exp. 22mm.
_ (23.6) Oligostigma leucomma, sp. n.
3. Head and thorax fulvous yellow mixed with white ; abdomen
white suffused with fulvous yellow except at base; palpi white at
tips ; pectus, legs, and ventral surface of abdomen white suffused
with fulvous yellow. Fore wing white, the costal and terminal
areas suffused with fulvous yellow; black subbasal strize above and
below vein 1; an antemedial fulvous-yellow band, defined on outer
side by some dark scales below the cell; a slightly waved blackish
medial line somewhat excurved below the costa; the outer half of
medial area fulvous yellow, suffused wich blackish below the cell;
postmedial line blackish, defined on outer side by white to vein 4
and with a white spot before it beyond the cell, slightly incurved
below costa and excurved at middle, at vein 4 bent.inwards to lower
angle of cell, then waved to inner margin; a black subterminal line
Pyralidee of the Subfamily Hydrocampine. 469
defined on inner side by a narrow white band, incurved from costa
to discal fold and below vein 2, excurved at middle; a black terminal
line; cilia white, black at apex and middle. Hind wing white,
the terminal area suffused with fulvous yellow; a curved eid
antemedial line with some dark scales on it below the cell ;
blackish discoidal bar ; postmedial line fulvous yellow, excurved .
vein 4, then bent inwards to lower angle of cell, then with some
dark scales on it and excurved above inner margin; a slightly
sinuous black subterminal line with a narrow white band on its
inner side, excurved at middle ; cilia white with a blackish line near
base and some dark scales at tips to vein 2.
Q. Head, thorax, abdomen, and fore wing almost entirely
fulvous yellow, the last without the white spot before the post-
medial line and the white beyond the postmedial line and before
the subterminal line reduced.
Hab. QUEENSLAND, Stradbroke I. (Turner), 3 5, 1 Q type.
Exp. 16-18 mm.
© (230) Oligostigma fulvicolor, sp. n.
6. Head, thorax, and abdomen fulvous, the last with some
yellow mixed; frons white; tarsi white at base. Fore wing
fulvous mixed with some yellow ; a whitish antemedial spot on
costa with a slight dark streak Belaive it; an inwardly oblique
slightly sinuous silvery-white medial band defined on each side by
blackish except at costa and with a slight dark streak beyond it
below costa to an elongate white spot defined by dark scales above
end of cell; a white point defined by dark scales at upper angle of
cell and a slight oblique dark streak below lower angle with a
yellow mark above it extending to the postmedial narrow silvery-
white band defined on each side by black-brown except at costa,
obliquely curved to vein 2, then incurved; a subterminal series of
short dark streaks with some whitish on the streaks above and
below vein 2; cilia dark brown. Hind wing fulvous; a narrow
slightly sinuous silvery-white antemedial band defined on each side
by black-brown ; a similar postmedial band, obliquely curved to.
vein 1, then bent outwards to tornus ; a subterminal series of slight
rather wedge-shaped dark marks with some whitish in centres ;
cilia dark brown.
Hab. Peru, Carabaya, Oconeque (Ockenden), 3 3 type. Exp.
34-38 mm.
(286) Oligostigma flavipictalis, sp. n.
¢. Head, thorax, and abdomen white, the hind tarsi tinged
with yellow on inner side. Fore wing silvery white; a golden-
yellow patch at base of costa, slightly defined on outer side by
brown ; a curved golden-yellow antemedial band, slightly defined
on each side by brown towards costa, where it is somewhat dilated ;
470 Sir G. F. Hampson on new
a slight brown discoidal striga; an orange-yellow patch below end
of cell in submedian interspace, its inner edge connected by a bar
with inner margin and its outer edge defined by a slight brown
line continued to inner margin ; an oblique orange-yellow mark on
costa above end of cell defined on inner side by a brown striga ; an
orange-yellow postmedial patch below costa with which its inner
edge is connected by an oblique bar, defined at sides by slight
brown lines ; a curved orange-yellow subterminal band from costa
to below vein 4 where it is bent inwards, its inner edge defined by
a blackish line to discal fold and with a slight oblique brownish
line from the inner side of its recurved part to tornus; a narrow
orange-yellow terminal band defined on inner side by a fine black
line, curved inwards at tornus. Hind wing silvery white ; a curved
orange-yellow subbasal band defined on outer side by a slight brown
line; an orange-yellow medial band defined by slight brown lines,
its outer edge forming a hook at vein 4, then incurved; an orange-
yellow subterminal band defined by fine black lines, its inner edge
strongly incurved from below vein’ 4 to submedian fold; the
termen narrowly white with a fine terminal black line; cilia
chequered yellow and white with some blackish scales at tips.
Hab. Stnaarore (Meade-Waldo), 1-2 type. Exp. 14 mm.
(18a) Aulacodes hemimelena, sp. n.
2. Head and thorax black-brown ; abdomen yellowish tinged
with black-brown and with some white at base; pectus white;
legs and ventral surface of abdomen yellow, the fore legs tinged
with red-brown, the fore femora black-brown above. Fore wing
black-brown; a curved silvery-white band from below costa to
vein 1 before the golden-yellow terminal band defined on inner side
by black striz and on outer by a terminal series of black points
and striga at submedian interspace; cilia silvery white. Hind
wing white with a black-brown pateh at base, the terminal area
broadly golden yellow defined on inner side bya black line between
discal and submedian folds; a minute ocellate white spot defined
by black on termen at discal fold, then three minute black spots
with some red between them to vein 2; cilia silvery white, brown
at base heyond the spots.
Hab. Puitrpprnes, Manila (Ledyard), 1 2 type. Exp.
18 mm.
(23 b) Aulacodes quadriplagiata, sp. n.
3. Headand thorax fulvous yellow; abdomen white tinged with
yellow; palpi with some brown at side of 2nd joint; legs yellow,
the fore tibize with dark brown band at extremities, the tarsi ringed
with brown; pectus and ventral surface of abdomen white. Fore
wing with the basal half black-brown except the costal area which
is fulvous yellow at base, then white, and a conical white ante-
Pyralide: of the Subfamily Hydrocampine. 471
medial spot at inner margin; a medial white band leaving the
costal yellow; a semicircular deep chocolate-brown postmedial
patch from below costa to vein 3 defined on outer side and below by
a curved silvery-white band and shading to red-brown at costa
before the white band; a terminal yellow band defined on inner
side by a black line to submedian fold, the yellow band bent
inwards on inner area to near the basal black-brown area; a
terminal series of black points and small spot below apex ; cilia
silvery white, tinged with brown at apex. Hind wing silvery
white, the terminal area broadly bright yellow, extending on inner
area to near base ; the white area defined by a curved black line
between discal and submedian folds; subterminal black striz above
and below vein 2, then a curved silvery-white line to above tornus ;
a curved silvery-white line from costa before apex to termen at
discal fold; minute silvery-white ocellate spots defined by black
and with black points on their outer edge above and below vein 4
before termen ; a minute black spot below vein 3 and striga below
vein 2; cilia silvery white tinged with red-brown at base.
?. Fore wing with the basal half chocolate-brown, its upper
edge indented by an elongate white mark in the cell and with
white streak below it on inner margin, the whole costal area above
it yellow, the postmedial patch red-brown and extending to sub-
median fold.
Hab. D’Enrrecastreaux Is., Goodenough I. (Meek), 1 ¢,
2 9 type. Hxp. 20-26 mm.
(24a) Aulacodes costifascialis, sp. n.
Hind tibiz of male rather curved downwards and fringed with
hair throughout.
Head, thorax, and abdomen white suffused with yellow, the
abdomen yellower except at base; palpi red-brown towards tips ;
fore legs with the femora black-brown above, the tibize with black-
brown band at extremity. Fore wing golden yellow; a rufous
fascia on costa to end of cell, where it expands into a triangular
patch to lower angle of cell, a silvery-white fascia below it in and
just below the cell; a wedge-shaped silvery-white patch beyond the
cell from below costa to vein 2, defined by slight fuscous lines; a
curved silvery-white subterminal band from costa to above vein 1,
where it is somewhat bifid, defined by fine black lines except at
costa; a terminal series of black points and striga in submedian
interspace ; cilia silvery white. Hind wing silvery white, the inner
and terminal areas broadly golden yellow, the white area defined
by an oblique sinuous black postmedial line from vein 6 to sub-
median fold; an obliquely curved silvery-white line from costa
before apex to termen at discal fold; four minute ocellate silvery-
white spots before termen between vein 5 and submedian fold, the
two upper spots with black points on their outer edges, the two
lower with black points beyond them, some orange-red on termen
472 On new Pyralidee of the Subfamily Hydrocampine.
between the spots and two minute black points above them above
vein 5; cilia silvery white with some brown at base beyond the
spots.
Hab. D’Enrrecasteaux Is., Goodenough I. (Meek), 2 g,4 2
type; Brsmarck Arcu., Rook I. (dleek), 2 3,19. Eup. 20-
24 mm.
(246) Aulacodes nigriplagialis, sp. n.
Eristena trigonalis, ab. 1, Hmpsn, A. M. N. H. (7) xviii. p. 390 (1906).
Hind tibiz of male slightly fringed with hair above towards
extremity.
Head, thorax, and abdomen orange-yellow, the head and teguli
with some brown mixed ; palpi irrorated with dark brown ; pectus
and ventral surface of abdomen white suffused with orange-yellow,
legs orange-yellow, the fore tibiz with dark brown band at extre-
mity. Fore wing orange-yellow ; a very dark red-brown fascia on
costa to end of cell, the end of cell below it white; a large conical
very dark red-brown patch tinged with blackish from postmedial
part of costa to vein 2, defined on inner side by a curved dark line
met at vein 2 by another faint curved dark line traversing the
patch and with a slight greyish tinge between them, asilvery-white .
band defining the outer edge of the patch defined on outer side by a
fine curved black line; a terminal series of black points, forming a
minute spot.below apex and striga at submedian interspace ; cilia
silvery white. Hind wing orange-yellow, the costal area whitish
to beyond middle; an oblique black postmedial line between discal
and submedian folds; an oblique silvery-white line from costa
before apex to termen at discal fold ; four small black spots before
termen between discal and submedian folds, ‘the two upper spots
defined on inner side by silvery white, with a waved black line
before them diverging obliquely below the 2nd spot, some orange-
red on termen between the spots; cilia silvery white, brown at base
beyond the spots.
Hab. Duron N. Guinea, Fak-fak (Pratt), 1 ¢ type, Kapaur
(Doherty), 1 9. Exp. 18 mm.
(24 ¢) Aulacodes dolichoplagia, sp. n.
3g. Head white, the frons suffused with golden yellow, the back
of head with some brown; thorax silvery white, the shoulders, tips
of patagia, and metathorax dark brown; abdomen golden yellow,
white at base; antenne yellow; palpi red-brown, white at base ;
pectus, legs, and ventral surface of abdomen white, the legs tinged
with yellow, the fore femora above and tibiwe at extremities dark
brown. Fore wing dark brown; a silvery-white fascia below base
of cell conjoined to a patch in end of cell; a large oblique conical
silvery-white patch from postmedial part of costa to below vein 3
beyond the cell; a series of white striz before the narrow orange-
yellow terminal band defined on inner side by a black line; 4
Descriptions and Records of Bees. 473
terminal series of black points; cilia brown at base, silvery white
at tips. Hind wing silvery white; a dark brown patch at base ;
the terminal area golden yellow, expanding on inner area to middle,
defined on inner side by a series of dark points from below costa to
vein 2 and a striga at vein1; a series of silvery-white marks before
termen from below costa to submedian fold, defined by blackish, the
spot below costa round, the others elongate except the small spot
below vein 5; minute terminal black spots above and below vein 4
and slight strize towards apex and between veins 3 and 1; cilia
silvery white with some brown at apex and a brown line through
them from vein 5 to near tornus. .
Hab. Durcn N. Guinea, Fak-fak (Pratt), 1 g type. Eup.
24 mm.
[To be continued. |
XLVI.—Descriptions and Records of Bees —LXXV.
By T. D. A. Cocxrrett, University of Colorado,
Aylocopa draconis, sp. nu.
3 .—Length about 25 mm., anterior wings 18°5 mm.
Black, without any metallic tint; thorax thickly covered
(except bare space on disc) with reddish-ochreous velvety
hair; abdomen not banded. Eyes extremely large, con-
verging above; mandibles bidentate, with a yellow basal
patch ; tubercle of labrum small; clypeus ivory-colour,
more or less brownish, with a pair of black spots, the surface
of clypeus closely punctured, but an impunctate. median
ridge ; supraclypeal area almost pallid; ocelli large, far
down on front ; face and front with red-brown hair, darkest
around ocelli; top of head and cheeks with reddish-ochreous
hair; anterior femora swollen, without hair below; anterior
and middle tibiz with bright fulvous hair on outer side,
reddish on inner, and sooty behind ; anterior tarsi similarly
coloured, but from middle of basitarsus on there is creamy-
white hair on under side posteriorly, beneath the sooty, and
on apical part of basitarsus anteriorly and beneath the red
is very bright ; middle tarsi with reddish-black hair above
and behind, but red beneath ; hind femora broad, basally
keeled beneath, with a pustuliform swelling on the side of
the keel; hind tibize with a conspicuous apical lobe, much
broader than long, on imner side; hind basitarsi with
ochreous hair in front, black above and red behind, the
Ann. & Mag. N. Hist. Ser. 8, Vol. xix. on
474 Mr. T. D. A. Cockerell—Deseriptions and
under side presenting a large, bare, shining, elevated, wedge-
shaped surface; tegule black. Wings brown, subtrans-
lucent, apically suffused with rosy-purple ; venation very
different from that of X. sinensis, Smith, the third sub-
marginal cell not conspicuously broadened or bulging
apically, and the second much less elongated. Abdomen
with sooty hair on first segment, second with fulvous, sooty
only along apical margin, the rest with scanty hair except
at sides and apex; at sides it is sooty, except anteriorly on
segments 3 and 4, where is some fulvous ; at apex the hair
is long and reddish; beneath, the bind margins of the
segments are narrowly bright ferruginous and the hair is
fulvous.
Southern China (no other particulars known). In U.S.
National Museum.
Resembles X. appendiculata, Smith, but the hair is quite
differeutly coloured. There is also a general resemblance to,
the smaller X. collarvs.
Xylocopa punctifrons, sp. n.
$ .—Length about 21 mm., anterior wings 16 mm.
Clypeus (except lower margin, broadening laterally),
transverse supraclypeal mark and lateral face-marks (ending
very obtusely halfway up front) ivory-colour ; labrum black,
with a minute light point ; thorax without conspicuous light
hair, except at sides, where a broad band of greyish hair
extends from the tegule to the ventral surface. Abdomen
with a little pale hair on first segment. Wings dark reddish
fuliginous, with strong purple tints ; light hair on hind
tibiz, and middle and hind tarsi, as in X. ¢arsata, but it is
ochreous.
French Congo (Queensland Museum). Received by the
Museum from Le Moult of Paris.
Very close to X. tarsata, Smith, but considerably larger,
with the greater part of clypeus and sides of face densely
punctured. The colour of the hair on the legs agrees with
that of YX. tarsata, var. namutonensis, Strand, but that form
has the size of ¢arsata. It is possible that X. punctifrons
is the undescribed male of X. tuderculiceps, Ritsema, but the
legs have much more light hair than in the female of that
species, and the localities are far apart.
In the same lot came XY. carinata, Smith, Mesotrichia
preusta (Smith),and Crocisa excisa, Friese, from Dimbroko,
French W. Africa.
*
Records of Bees, 475
Centris maroniana, sp. 0.
d .—Length about 28 mm., anterior wings 22°5 mm.
Robust, black, including the legs, but anterior trochanters
and tibize suffused with chestnut«red. Eyes very large, con-
verging above; ocelli large, practically contiguous, and
lateral ones distant from eyes less than half diameter of one ;
sides of face, supraclypeal area and upper part of clypeus
ferruginous ; the rest of clypeus and the labrum chrome-
yellow ; scape short and stout, dark reddish, yellow in front;
mandibles elbowed near apex; hair of head clear ferru-
ginous ; disc of mesothorax and mammiform elevations of
scutellum shining; thorax densely covered with velvety
hair, black, with a faint rusty tint dorsally, except ante-
riorly, where it is rich deep red, the red gradually fading
into the black ; pleura with dark reddish hair ; anterior legs
with red hair, black on basitarsi; middle and hind legs with
long pure black hair; tegule ferruginous. Wings dark
fuliginous, brilliantly purple, with some shades of green.
Abdomen with short velvety hair, which is black except a
broad yellowish-white (olivaceous-tinted) band, occupying
second segment except base and third except extreme apex.
“ Guyane, Maroni ” (Queensland Museum; received from
Le Moult).
Related to C. smithiana, Friese (which J have from
F. Smith’s collection), but larger, with the hair of the
thorax dorsally black except in front. From the character
of the ocelli, it possibly flies in the evening or at night.
The bee-fauna of Maroni, as shown by the same collection,
includes the following :—Acanthopus splendidus, Vab., Aglaé
cerulea, Lep., Exerete frontalis, Guér., Oxea festiva, Sm.,
Xylocopa tarbata, Fab., Bombus incarum, Frankl., Centris
obsoleta, Lep., C. americana, Klug, Epicharis conica, Sm.,
E. schrottkyi, Friese, E. affinis, Sm., Ceratina leta, Spin.,
Lulema dimidiata, L., E. fasciata, Lep., H.mocsaryi, Friese,
E. smaragdina mexicana, Mocs., Euglossa brulle, Lep.,
Eufriesia pulchra, Sm., &c.
Pachymelus mediocincius, sp. 0.
d@.—Length 18 mm., length of anterior wing 14 mm,
Black, with tarsi dark red ; eyes large, slightly converging
above; clypeus prominent, but flattened on disc, yellow,
with upper and lateral margins and two conspicuous spots
black; labrum large, emarginate at apex, yellow, with
lateral and inferior margins narrowly black; muaudibles
31*
476 Mr. T. D. A. Cockerell—Deseriptions and
bidentate, the inner tooth short (style of P. hova), basal
part of mandibles mainly very pale yellowish ; scape with a
yellow stripe in front. Face, front, and vertex with long
black hair, but also white at sides of face and (appressed)
on each side of labrum ; occiput and cheeks with long white
hair ; mesothorax shining, but distinctly punctured ; scu-
tellum only feebly bigibbous; thorax with black hair,
becoming obscurely ochreous along anterior margin of
mesothorax, bright ochreous (forming a conspicuous wide
band) on metathorax, and pure white in middle of ventral
surface. Legs with mainly black hair; anterior tibiz with
a band of appressed golden pubescence in front; tegule
black. Wings dilute fuligimous; venation as in P. micre-
lephas, but second s.m. narrower. Abdomen closely punc-
tured, basal segment with black hair; apical margin of
second segment, and all of next three except base, covered
with appressed bright ferruginous pubescence; sixth seg-
ment with hair partly red and partly black; apical plate
emarginate.
Miarinarivo, Madagascar (Queensl. Museum ; from Le
Moult).
Similar to P. cambouei, Sauss., but that is a very much
larger species, with the scutellum different.
Hylesides concinnus (Fabricius).
Launceston, Tasmania, Feb. 15-16, 1916 (Ff. M. Littler).
Genus new to Tasmania. The female agrees with main-
land specimens ; but the male, compared with one from
Sydney, differs by the wholly black prothorax and the more
distinct punctures of second abdominal segment.
Megachile derelicta, Cockerell.
? St. Patrick’s R., Tasmania, 6.2.14 (Littler).
New to Tasmania.
-Mesotrichia bryorum (Fabricius).
Daru, Papua (Queensland Mus.).
The female has the wings. suffused with rosy-purplish,
whereas Australian examples usually have them much more
green,
Eulema amabilis, sp. n.
_fAt first sight exactly like 4. bruesi, Ckll., but
differing thus : green of mesothorax more brilliant, extend-
ing along the sides to the posterior end, where it is very
Records of Bees. ATT
bright and broadly margined mesad with blue ; scutellum
with lateral sulci strongly marked, the general surface of
scutellum dark purplish and shining, the lateral margins
thickened and shining steel-blue ; median smooth ridge of
labrum little broadened above ; apex of abdomen broadly
emarginate, formed as in E. manni, Ckll. From E. manni
it is at once known by the strong keel down middle of
elypeus, the dark black-haired first abdominal segment,
the blue margins of scutellum, &c. From E. smaragdina,
Perty, by the black hind tibiz, marked with green poste-
riorly (with a rather small but distinct tubercle above the
spurs), and tle entirely black hair of thorax. From &.
auripes, Gribodo, by the strong clypeal keel, the less promi-
nent lateral keels of labrum, and the hair of legs differently
coloured, the fringe on apical part of auterior tarsi ferru-
ginous, while the pubescence on outer face of middle basi-
tarsi is shining creamy-white. It is also distinct from £.
mexicana aud the various related forms described by Friese
and others. The mesopleura is dark blue.
Manaos, Brazil (Miss H. B. Merrill). U.S, Nat. Museum.
Mesonychium dugesi, sp. 0.
g .—Length about 15 mm.
Very robust, black, with the abdomen dark but. brillant
blue, the hind margins of the segments more or less green ;
dise of mesothorax dark purple-blue on each side of the
median sulcus ; pubescence at first sight appearing wholly
black, but it is mixed with ochreous on labrum and lower part
of clypeus, and there is a spot of thesame on each side of front;
on lower part of pleurais a little pale hair, and there is white
hair on anterior tibiz posteriorly ; second and third antennal
joints dark red in front; scutellum rather short, hairy, —
without conspicuous prominences; third s.m. strongly
narrowed above, but not nearly to a point ; spur of middle
tibia strongly bifid, one division with two or three spines ;
hind femora broad, with a very large tooth beneath near
base ; hind tibize with a polished red area at end ; venter of
abdomen with a large red area in subapical-region. The
wings are brownish subhyaline, with a dark apical cloud.
Guanajuato, Mexico (A. Dugés). U.S. National Museum.
This has the appearance of the species which Schrottky
places in his genus Cyphomelissa, but the third submarginal
cell is as in Melissa or Mesoplia. It will easily be known
‘from Mesonychium insigne (Melissa insignis, Sm.), from
Orizaba, by the absence of the bright yellow hair. The
A478 Mr. T. D. A. Cockerell—Deseriptions and
venation and middle spur separate it from M. ce@rulescens,
Lep.
Mesonychium decoratum (Smith).
Bocas del Toro, Panama, July 6, 1908 (W. Robinson).
This agrees with one from F. Smith’s collection, obtained
by Bates in 8. Paulo, Brazil. A form with broader pygidial
plate (?) comes trom Rio Mato, Venezuela, October
( Carriker).
Mesonychium azureum guatemalense (Cockerell).
Cacao, Trece Aguas, Guatemala, April 4 (Barber &
Schwarz).
This specimen shows that the type was partly denuded.
The disc of mesothorax and outer face of hind tibiz are
ornamented with green scale-hke hairs.
Mesonychium duckei (Friese).
Cabima, Panama, May 17, 1911 (Aug. Busck).
The third s.m. is very much broader below than in
M. decoratum, so that the venation approaches Cyphomelissa.
After reviewing the subject, I must agree ‘with Ducke that
Mesonychium is the proper name for this genus, including
Mesoplia and Melissa, and also Cyphomelissa as now inter-
preted by Schrottky.
Colletes cyaneseens, Haliday.
I have this from Santiago, Chile, and specimens marked
Southern Chile (M. J. Rivera) are in the U.S. National
Museum. C. atripilis, Vachal, is a synonym.
Triepeolus pruinosus, 8p. Ni.
2 (type).—Length about 9 mm.
Black, the thorax densely punctured and not shining ;
basal half of mandibles red ; labrum dark reddish ; clypeus
very minutely and densely punctured ; scape red at base
and more or less at apex ; second and third antennal joints
bright ferruginous ; a conspicuous patch of creamy-white
hair on each side of antennz ; mesothorax with a thin
pruinose pubescence, anterior margin with two nearly round
spots of yellow pubescence ; yellowish-white prothoracic
hair-band broadly interrupted in middle, ending laterally in
round spot on base of tubercles; tubercles reddish ; hind
margin of mesothorax with a creamy hair-band ; scutellum
a im
Records of Bees. 479
moderately bigibbous, axillz bluntly conical ; area of meta-
thorax dull and rough basally ; tegule bright ferruginous.
Wings strongly dusky at apex. Legs obscure ferrugmous,
spurs red. Abdomen with broad yellow bands on first two
segments, that on first anteriorly produced at sides into an
evanescent cloud, not a distinct tooth or band-like lobe ;
third and fourth segments with whitish bands, more or less
failing in middle; modified pygidial space subcircular, not
very large; last ventral segment produced beyond last
dorsal, the very broad end turned downward.
? .—Similar in appearance, but the legs are mainly black ;
the anterior tibiz, middle tibiz in front, hind tibiz at base,
and the tarsi (the hind ones not wholly) are red; face
densely covered with creamy-white hair; mandibles black
with a median red band; flagellum black except at base ;
yellow spots on anterior margin of mesothorax larger,
suffusedly elongate posteriorly ; abdomen with five yellow
bands (successively paler) and one white one, the first two
bands more or less suffused with brownish-orange ; ventral
fringes pale reddish at ends. The dark parts of abdomen
are hoary with a fine pale pubescence.
Carcarana, Argentina (L. Bruner, 15). U.S. Nat. Museum.
Resembles Epeolus burmeisteri, Friese, but considerably
larger, with darker legs, and the patch on anterior margin
of mesothorax divided into two spots. Hpeolus ( Doeringiella)
bizonatus (Holmbg.), from Bahia Blanca (Bruner), is super-
ficially very like 7. pruinosus, but larger, ad easily separated
by the extraordinary antenne.
Isepeolus vierecki, Jorgensen.
Bahia Blanca, Argentine (Bruner); San Juan, Argentine
(C. S. Reed).
It is permissible to correct the specific name, printed
“ yecki” in Jorgensen’s work.
Isepeolus bruneri, sp. n.
? —Length 10 mm.
Black, mandibles obscurely reddish in middle, but other-
wise tegument of head and thorax all black; thorax
variegated with white hair as in other species, with two
conspicuous black spots on scutellum, and others covering
axilla; disc of mesothorax shining, with well-separated
unctures; first two abdominal segments ornamented as in
I. cockerelli, Jorg., except that the inner processes on second
segment are longer; ‘third with a pair of large oblique
480 Descriptions and Records of Bees.
(quadrate) patches on hind margin, and each side with a
large complicated patch of white, presenting a deep sinus
anteriorly; fourth segment with a very large and thick
mark having two parts, like the letter H; fifth with two
large spots, not reaching apical margin ; sixth segment with
.a small outwardly directed basal spime on each side. _ Face
with white hair, partly black on lower part; front and
vertex with black hair; a band of white hair in front of
ocelli; scape red at base, middle covered with long white
hair, the broad apex intense black; flagellum red beneath ;
third antennal joint unusually short for the genus, not as
long as next three combined ; tegule red. Wings brownish .
on apical margin, stigma and nervures piceous; second s.m.
distinctly narrower above, receiving first r. n. before end.
Legs black with the usual white hair-marks, knees red ;
pleura with black hair.
Carcarana, Argentina (L. Bruner).
Allied to J. cockerelli, but known by the abdominal
pattern, venation, &c.
Lonchopria alopex, sp. un.
& .—Length about 14 mm.
Head, thorax, and legs black, with long and abundant
fox-red hair. Abdomen shining olive-green, with the same
red hair, except the last two segments, which are black, the
penultimate with black hair. Mandibles bidentate, reddish
apically ; malar space very short ; clypeus very smooth and
polished, the upper part with two rounded elevations, between
which is a depression bearing a tuft of very long red hair;
labrum bituberculate ; antenne very long, reaching to meta-
thorax, flagellum bright ferrugimous beneath except at base;
face very broad; mesothorax shining, with well-separated
punctures; area of metathorax smooth, with an obtuse
transverse ridge ; tegule black. Wings dusky, stigma dull
ferruginous; third s.m. very oblique ; apical segment of
abdomen keeled. Maxillary palpi with six subequal joints.
La Paz, Bolivia, Nov. 14, 1898. U.S. National Museum.
A remarkable species, superficially resembling L. thoracica
(Friese), but with much longer and more abundant hair on
abdomen, shorter stigma, quite different mandibles, &ce.
According to specimens received from Friese and Jensen- |
Haarup, it is L. armata, Fr., which is the male of L. chalybea,
Fr., not ZL. enea, Fr., as Friese first thought. L. marginata
(Spin.), described as a Colletes, the specific name preoccupied,
apparently becomes L. zonalis (Reed, 1892).
Mr. G. J. Arrow on the Khapra Beetle. 481
Svastra bombylans (Holmberg).
Bahia Blanca, Argentine (Bruner, 2).
Xenoglossa crawfordi, Cockerell.
Guanajuato, Mexico (A. Dugés).
‘olletes punctipennis, Cresson.
Brownsville, Texas, 1998 (Jones § Pratt).
New to the United States.
Pseudomelecta californica miranda (Fox).
Mexico (C. F. Baker collection, 2320).
Megachile anthracina, Suith.
Moulmein, I. Burma, Dec. 1910 (R. L. Woglum).
XLVIU.—The Khapra Beetle (Yrogoderma khapra, sp. n.),
an Indian Grain-pest. By GILBERT J. ARROW.
(Published by permission of the Trustees of the British Museum.)
Tuis very destructive wheat-pest has been studied in great
detail by Messrs. J. H. Barnes and A. J. Grove, who have
published figures and descriptions of it in all its stages in Mem.
Dept. Agric. India (Chemical Series), iv. 6, 1916, p. 172)
under the name Attagenus undulatus, Motsch. As already
stated in a footnote in the ‘ Review of Applied Entomology,’
~v. 1917, p. 126, the insect is really a species of Trogoderma
and appears to be without a specific name. Attagenus undu-
latus is quite a different insect, as I have established from
specimens in the British Museum received from Motschulsky
himself (see Ann. & Mag. Nat. Hist. (8) xv. 1915, p. 426).
Mr. Bainbrigge Fletcher has incorrectly reported me (Agric.
Research Inst. Pusa, Bull. 59, p. 14) as saying that the
insect common in stored wheat in Northern India should
be known by this name. On the contrary, the distribution
I recorded shows A. undulatus to belong to tropical and not
wheat-growing latitudes.
_ Specimens found in imported wheat have been received
at the British Museum during many years past, and I have
482 Mr. G. J. Arrow on the Khapra Beetle.
,
regarded them as probably identical with Trogoderma versi-
color, Creutz., but they have invariably been in such bad
condition that exact determination was impracticable. Under
the name of ‘ Kapra ” specimens were sent to the Museum by
the late E. T. Atkinson in 1888 and stated to be destructive
to wheat in godowns at Delhi. Recently I have been able to
examine perfect examples, bred in the greatest abundance from
samples of Karachi wheat collected by Mr. J. H. Durrant, and
the study of these has convinced me that the species is neither
T. versicolor, Creutz., nor 7. inclusum, Lec., the figures and
descriptions of which show them to be larger and darker
coloured, with different antenne, and possibly not distinct one
from the other. It is, therefore, necessary to give a new name
to this exceedingly serious pest, and I propose to adopt the
vernacular name by which, according to Messrs. Barnes and
Grove, it is known to Indian grain-dealers. It may be briefly
diagnosed as follows :—
Trogoderma khapra, sp. n.
Rufo-ferruginea, capite, pronoto corporeque subtus obscurioribus,
antennis pedibusque rufis, elytris vage fusco-bifasciatis ; ovalis,
nitida, corpore subtus equaliter, supra longius et magis irregu-
lariter griseo-pubescenti; antennis 11-articulatis, articulis 3-7
minutis, 8-11 sat magnis, clavam formantibus, foemine ovatam,
multo compactam, maris longiorem, apice producto et compresso.
Long. 1°75-3 mm.
Although I have seen an enormous number of specimens,
the largest scarcely exceeds 3 mm. in length, with the head
fully extended, and this is considerably less than the size
indicated for the European and N.-Ameriecan types of Creutzer
and Leconte. The elytra are of a rather light red-brown
shade, generally marked with two vaguely defined darker
transverse bands, and the head and pronotum are nearly
always distinctly darker than the elytra, but rarely black.
The surface is clothed with grey hairs, which are very easily
rubbed off, and the worn specimens found amongst the grain
are very smooth and shining. Upon the darker areas of the
elytra the hairs are finer and scantier. ‘The antennze and
legs are entirely light in colour.
The males are much smaller on the average than tlie
females and have rather longer antenne, the joints composing
the club, and especially the terminal one, being more
elongate.
This insect is found in enormous profusion in cargoes
of wheat from Karachi and Bombay ; but there is no evidence
that it is able to perpetuate itself in Hurope, nor has it been
found in grain imported from other regions than India.
Geological Society. 483
PROCEEDINGS OF LEARNED SOCIETIES.
GEOLOGICAL SOCIETY,
February 28th, 1917.—Dr. Alfred Harker, F.R.S., President,
in the Chair,
The following communication was read :—
‘Fourth Note on the Piltdown Gravel, with Evidence of a
Second Skull of Hoanthropus dawsoni. By Arthur Smith
Woodward, LL.D., F.R.S., V.P.G.S. With an Appendix on the
Form of the Frontal Pole of an Endocranial Cast of Hoanthropus
dawsont. By Prof. Grafton Elliot Smith, M.A., M.D., F.R.S.
Excavations last summer round the margin of the gravel-pit at
Piltdown (Sussex) supported the conclusion that the deposit is a
varied shingle-bank, and that the three layers containing Palxo-
lithic remains and derived Pliocene fossils are approximately of
the same age. Many elongated flints and pieces of Wealden
sandstone were observed in the bottom sandy clay with their long
axis more or less nearly vertical. No teeth or bones were found,
but one nodular flint obtained from the same layer as Hoanthropus,
seems to have been used by man as a hammer-stone. This is not
purposely shaped, but merely battered along faces that happened to
be useful when the stone was conveniently held in the hand.
Tn the winter of 1915 the late Mr. Charles Dawson discovered
in a plonghed field, about a mile distant from the original spot,
the inner supraorbital part of a frontal bone, the middle of an
occipital bone, and a left lower first molar tooth, all evidently
human. These are rolled fragments, and the first and third may
be referred with certainty to Hoanthropus dawsoni; but it is
doubtful whether they represent more than one individual. In
mineralized condition they agree with the remains of the type-
specimen. The piece of frontal bone exhibits the characteristic
texture and thickness, with only a very slight supraciliary ridge,
and a small development of air-sinuses. The occipital bone is
somewhat less thickened than that of the original specimen of
Eoanthropus, and bears the impression of a less unsymmetrical
brain. The external occipital protuberance is a little above the
upper limit of the cerebellum, as. in Neanderthal man; thus
differing from the condition both in KHoanthropus and in modern
“man. ‘The lower molar is exactly similar to the first lower molar
of Hoanthropus already described, but is more obliquely worn by
mastication. Detailed comparison shows that this tooth is human,
differing essentially from that of a chimpanzee in its more hypso-
dont crown, thicker enamel, and less prominence of the neck over
the root. The occurrence of the same type of frontal bone with
the same type of lower molar in two distinct localities, adds to the
probability of their belonging to one and the same species. With
these remains were found brown flints in great abundance, and
one rolled portion of a lower molar tooth of Rhinoceros in the
same highly-mineralized condition as the derived Pliocene teeth at
Piltdown.
In an Appendix, Prof. G. Elliot Smith expresses the opinion
that the endocranial cast of the fragment of frontal bone presents
features more primitive and more ape-like than those of any other
known member of the human family,
A484 Miscellaneous.
MISCELLANEOUS.
[ We have received from the Secretary to the International Commis-
_ sion on Zoological Nomenclature a circular letter giving 39 generic
names in Protozoa, Coelenterata, Trematoda, Cestoda, Cirripedia,
Tunicata, and Pisces, chiefly Linnzean, which have been proposed
for inclusion in the Official List of Zoological Names. Owing to its
length we are unable to publish the list in full, but a copy will be
sent to any person sufficiently interested on application to the Secre-
tary to the International Commission on Zoological Nomenclature,
Smithsonian Institution, United States National Museum, Wash-
ington, D.C., U.S.A. ]
Notice to the Zoological Profession of a Possible Suspension of the
International Rules of Zoological Nomenclature in the Cases of
Musca, Linneus, 1758, and Calliphora, Desvoidy, 1830.
In accordance with the Rules of the International Zoological
Congress, the attention of the zoological profession is invited to the
fact that Dr. L. O. Howard, W. Dwight Pierce, and twenty-one
other professional zoologists have requested the International
Commission on Zoological Nomenclature to exercise its plenary
power in the case of the Linnean genus Musca, 1758, and, under
suspension of the Rules, to declare J. domestica as type of this
genus, also, under suspension of the Rules, to validate reas
Desvoidy, 1830, with C. vomitoria as type.
The request is based on the grounds of practical utility, and an
almost unbroken history of cunsistent usage since 1758 in the case
of Musca, and since 1880 in the case of Calliphora, It is claimed
that a strict application of the Rules will produce greater confusion
than uniformity.
According to the premises at present before the Commission, if
the Rules are strictly applied, the generic name of Musca would
take either MW. cesar or M. vomitoria as type, and the species
M. domestica would be cited either in Conostoma, 1801 [?] (type
Ascaris conostoma=larva of JJ. domestica), or in Conosoma, 1802
(type Ascaris conosoma=larva of M. domestica), or in Promusca,
1915 (type M. domestica), thus resulting in a very regrettable”
change in the nomenclature of the species in qnestion as almost
universally used in entomological, zoological, medical, epidemio-
logical, and veterinary literature.
The Secretary of the Commission invites any person interested in
these cases of nomenclature to communicate his opinion on the
subject as soon as possible, and not later than. May 1, 1918, when
the subject will be submitted to the Commission for vote.
C. W. Srizes,
Secretary to Commission.
25th & EK. Streets, N.W.,
Washington, D.C.
INDEX ro VOL. XIX.
ACARINA, new, 136.
Aconemys, new species of, 281.
Actinopyga agassizii, on the occur-
rence of, in Bermuda, 406.
Alex, new species of, 416.
Ambia, new species of, 459.
Ananca, new species of, 170.
Andrena, new species of, 282.
Anobostra, characters of the new
genus, 97.
Anthophora, new species of, 287,
Aphytoceros, new species of, 342,
Argyractis, new species of, 363.
Arhopala, new species of, 409,
Arrow, G. J,, on melolonthine coleo-
ptera, 59; on the Khapra beetle,
481.
Arthrolips, new species of, 13.
Asthenopholis, key to the species of,
60,
Attatha, new species of, 337.
Aulacodes, new species of, 470.
Barnacles from the hull of the ‘ Terra
Nova,’ on, 229.
Bather, Dr. F. A., on triassic cri-
noids from New Zealand, 360.
Bathyparia, characters of the new
genus, 56
Batrachians, new, 407.
Bembex, new species of, 436.
Birks, Rey. S. G., on Cylindroiulus
nitidus, 417.
Books, new: —Boulenger’s Catalogue
of Freshwater Fishes of Africa,
160 ; Hampson’s Catalogue of the
Lepidoptera Phalzne,
ment, vol. i., 291.
Borradaile, L. A., on barnacles from
the hull of the ‘Terra Nova,’ 229,
Bostra, new species of, 90.
Boulenger, G. A., on new lizards of
the family Lacertidee, 277; on a
new lizard and two new frogs
from W. Africa, 407. '
Brade, Miss H. K., on Cylindroiylus
nitidus, 417.
Bradypus, new species of, 356.
Brevoortia, notes on species of, 301,
Cacoplesia, new species of, 163.
Calcagninus, new species of, 103.
Campion, H., on Fabricius’s types of
Odonata in the- British Museum,
441.
Cataclysta, new species of, 374, 457,
Cavia, new species of, 154.
Centris, new species of, 475.
Cerceris, new species of, 104.
Champion, G. C., on coleoptera from
the Seychelles and Aldabra, 161.
Chilton, Prof. C., on the New Zea-
land amphipod Hyale grenfelli,
273; on a new tuberculate ter-
restrial isopod from New Zealand,
327.
Chlorops, new species of, 51.
Chromatopterum, new species of, 49,
Chyliza, new species of, 270.
Cirphis, new species of, 336.
Cirripedes, notes on New Zealand,
229,
Supple-
486
Clupea, on the fishes of the genus,
226.
Cockerell, T. D. A., descriptions and
records of bees, 282, 473.
Coleoptera, new, 1, 59, 144, 161, 188,
395, 482. ;
Collembola, notes on, 425.
Corylophidz from the Seychelles
and Rangoon, on, 1.
Cricetiscus, characters of the new
genus, 456.
Cricetulus, new species of, 452.
Crick, G.C., on the belemnite animal,
291.
Crozier, W. J., on the occurrence of
a holothurian new to the fauna of
Bermuda, 405.
Crustacea, new, 229, 327, 405.
Cubaris, new species of, 327.
Curculionide, new Indian, 188.
Cylindroiulus nitidus, note on, 417,
Cynothrissa, characters of the new
genus, 203.
Dattinia, new species of, 96.
Daubania, characters of the new
oenus, 19.
Deilemera, new species of, 335, 411.
Diatomineura, new species of, 209.
Dicranocnemus, new species of, 62.
Dimorpha, new species of, 518.
Diptera, new, 33, 207, 225, 266.
Distant, W. L., on the homoptera
of Indo-China, 100.
Ectenopsis, new species of, 217.
Elalis, new species of, 76.
Elis, new species of, 517.
Encopognathus, new species of, 106.
Epiplema, new species of, 339,
Eremias, new species of, 279.
Erephopsis, new species of, 211.
Eristena, new species of, 368.
Ethmalosa, characters of the new
genus, 302.
Eulema, new species of, 476,
Eurina, new species of, 42.
Eurygenius, new species of, 174.
Fishes, new, 198, 297, 377.
Gargela, new species of, 362.
Geckobia, new species of, 138.
Geckobiella, characters of the new
genus, 138.
Geological Society, proceedings of
the, 230, 291, 357, 483.
Geropaschia, definition of the new
generic name, 361.
INDEX,
Gonialosa, characters of ‘the new
genus, 315,
Gouna, new species of, 61.
Groves, J., on Characee from the
Lower Headon beds, 231.
Hematopota, new species of, 225.
Hampson, Sir G. F., on new Pyra-
lide, 65, 361, 457.
Hapalemur group, on the lemurs of
the, 543 ; new species of, 348.
Haplegis, new species of, 47.
Harengula, synopsis of the species of,
386.
Hebomoia, new species of, 332,
Helomyza, new species of, 266.
Herculia, new species of, 81.
Heterochelus, new species of, 61.
Hilsa, characters of the new genus,
305.
Hirst, S., on some new mites, 136.
Holothurian, on the occurrence of a,
new to the fauna of Bermuda,
405.
Homonotus, new species of, 149,
Homoptera, new, 100.
Hoplionota, new species of, 144.
Huechys, new species of, 104,
Hyale grenfelli, note on; 273.
Hymenoptera, new, 104, 147, 282,
317, 456, 475.
Ichnotropis, new species of, 278.
Tsamia, new species of, 331.
Isepeolus, new species of, 479.
KXhapra beetle, note on the, 431.
Lacerta, new variety of, 277.
Lagaroceras, new species of, 44.
Lamb, C. G., on exotic Chloropidee,
33;-on exotic Helomyzidie, Scio-
myzidze, and Psilide, 266.
Lapworth, Dr. C., on graptolites
trom Peru, 295.
Lemurs of the Hapalemur group, on
the, 343.
Lepidoptera, new, 65, 331, 361, 409,
457.
Lewisium, new species of, 23.
Lile, new species of, 394.
Limnothrissa, characters of the new
genus, 207.
Lonchopria, new species of, 480.
Lorymodes, characters of the new
genus, 96.
Loxocera, notes on species of, 272.
Lygosoma, new species of, 407.
M‘Intosh, Prof., on the nervous
INDEX.
system and other points in the
structure of Owenia and Myrio-
chele, 233,
Mammals, new, 154, 280, 281, 348,
356, 450, 452.
Margaronia, new species of, 342.
Margarosticha, new species of, 372.
Marshall, Dr. G. A. K., on new
species of Indian Curculionidae,
188; on new weevils of the genus
Mecysmoderes, 395.
Maulik, 8., on Cassidine and Bru-
chide from the Seychelles and
Aldabra, 144.
Mecysmoderes,
395.
Meioderus, new species of, 16.
Meionops, characters of the new
genus, 191,
Mesonychium, new species of, 477.
Microthrissa, new species of, 202.
Microtus, new species of, 450.
Mogannia, new species of, 103.
Monomma, new species of, 162.
Mordella, new species of, 179.
Mordellistena, new species of, 183,
Murgisca, new species of, 562. ~
Mycteromimus, characters of the
new genus, 166.
Myriochele, notes on the nervous
system &e. of, 255.
Nematolosa, characters of the new
genus, 312.
Nematoserica, characters of the new
genus, 64,
Nomenclature, notice of a possible
suspension of the rules of, 484.
Notocrypta, new species of, 410,
Nymphula, new species of, 369.
Odaxothrissa, new species of, 205.
Odonata, on Fabricius’s types of, in
the British Museum, 441.
Oligostigma, new species of, 465.
Onychocnemis, characters of the new
genus, 193.
Ops, new species of, 50.
Orthoperus, new species of, 29.
Osericana, new varieties of, 341.
Owenia, notes on the nervous system
&e. of, 283.
Oxacis, notes on species of, 168.
Pachylophus, new species of, 37,
Pachymelus, new species of, 475,
Paractenia, new species of, 88.
Parectecephala, new species of, 52.
new species of,
487
Pellonula, new species of, 201.
Peltotrachelus, characters of the new
genus, 188.
Pemphigonotus, characters of the
new genus, 54.
Phzenomerus, new species of, 197.
Phryganomima, characters of the
new genus, 95.
Pimeliaphilus, new species of, 142.
Pingasa, new species of, 416,
Pison, new species of, 109.
Plutopaschia, characters of
new genus, 361.
Pocock, R. L., on the external cha-
racters of the Felidae, 1138; on
the lemurs of the Hapalemur
group, 345; Felide, on the ex-
ternal characters of the, 113.
Peecilothrissa, characters of the new
genus, 201.
the
_ Pomolobus, notes on species of, 299.
Potamothrissa, characters of the
new genus, 205,
Prosopigastra, new species of, 325,
Pterygosoma, new species of, 136,
Pyralide, new, 65, 361, 457,
Pyralis, new species of, 65,
Rana, new species of, 407.
Rappia, new species of, 408,
Regan, C. T., revision of the clupeid
fishes of the genus Pellonula and
of related genera in the rivers of
Africa, 198; on the fishes of the
genus Clupea, 226; on the clupeoid
fishes of the genera Promolobus,
Brevoortia, and Dorosoma, 297 ;
on the clupeid fishes of the genera
Sardinella, Harengula, &c., 377.
Reid, C., on Characeze from the
Lower Headon beds, 231.
Reptiles, new, 277, 407.
Rhopalum, new species of, 107.
Rhypobius, new species of, 27.
Ricardo, Miss G., on new species of
Tabanide from Australia and the
Fiji Islands, 207; on new Indian
species of Hematopota, 225.
Routledge, S., on the geology of
Easter Island, 357.
Sacada, new species of, 88.
Sacium, new species of, 7.
Salatura, new subspecies of, 381.
Sapyga, new species of, 318.
Sardinella, new species of, 380,
Scotcecus, new species of, 280,
488
Scott, H., on Corylophides from the
Seychelles and Rangoon, 1.
Sericoderus, new species of, 18.
Shoebotham, J. W., notes on Collem-
bola, 425.
Silvius, new species of, 212.
Sitina, new species of, 333.
Sloths, notes on three-toed, 352.
Smith, Dr. S., on a new coral genus,
230.
Sparrmannia, new species of, 59.
Stebbing, Rey, T. R. R., on 8.-Afri-
can Talitridz, 330.
Steleocerus, new species of, 39.
Stemmatophora, new species of, 76.
Stictodrya, new species of, 165.
Stictoptera, new species of, 337.
Stolothrissa, characters of the new
genus, 206,
Stopes, Dr. M. C., on mesozoic
cycads, 293.
Swinhoe, Col. C., on new Indo-
Malayan lepidoptera, 331, 409.
Sylepta, new species of, 342.
Tabanidz, on new species of, from
Australia and the Fiji Islands,
207.
Tabanus, new species of, 213.
Tachysphex, new species of, 320.
Talitride, notes on 8.-African, 330,
TN DEK
Tegulifera, new species of, 70.
Telicota, new variety of, 410.
Teluropus, characters of the new
genus, 195,
Terpnosia, new species of, 101.
Tetralonia, new species of, 287.
Thomas, O., on the species of Cavia,
152; on a new species of Scotce-
cus,’ 280; on a new species of
Aconemys from Southern Chili,
281; notes on three-toed sloths,
352; on a new vole from Pales-
tine, 450; on the small hamsters ~
that have been referred to Crice-
tulus pheeus and campbelli, 452,
Trechmann, C. T., on the trias of
New Zealand, 359.
Triepeolus, new species of, 478.
Trogoderma, new species of, 482.
Turner, R. E., notes on fossorial
hymenoptera, 104, 147, 317, 436.
Tyndis, new species of, 99.
Willowsia, definition of the new
generic name, 431.
Xylocopa, new species of, 475.
Xylophilus, new species of, 176.
Zitha, new species of, 95. f
Zoological nomenclature, notice of a
possible suspension of the rules of,
484,
END OF THE NINETEENTH VOLUME,
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SCOTT. Ann. & Mag. Nat. Hist. S. 5. Vol. XIX. Pisce
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SCOTT. Ann. & Mag. Nat. Hist. S. 8. Vol. XIX, Pl. HI.
H. Scott del. Camb. Univ. Press
CORYLOPHID BEETEES:
SCOTT. Ann. & Mag. Nat. Hist. 8. 8. Vol. XIX. Pl. IV.
H. Scott del. Camb. Univ. Press
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Notice to the Zoological Profession of a Possible Suspension of the
International Rules of Zoological Nomenclature in the Cases of
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