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FISCHER, P.-H., DuvAL, M. & Rarry, A. 1933. Etudes sur les échanges respiratoires des littorines. Archs
Zool. exp. gén. 74: 627-634.
Koun, A. J. 1960a. Ecological notes on Conus (Mollusca: Gastropoda) in the Trincomalee region of Ceylon.
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THIELE, J. 1910. Mollusca: B. Polyplacophora, Gastropoda marina, Bivalvia. In: SCHULTZE, L. Zoologische
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ANNALS OF THE SOUTH AFRICAN MUSEUM
ANNALE VAN DIE SUID-AFRIKAANSE MUSEUM
Volume 77 +#Band
December 1978 Desember
Part 5 Deel
UPPER CRETACEOUS SEDIMENTS FROM THE
IGODA RIVER MOUTH, EAST LONDON,
SOW TVA RIG”
By
HERBERT CHRISTIAN KLINGER
&
BRIAN E. LOCK
Cape Town Kaapstad
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UPPER CRETACEOUS SEDIMENTS FROM THE IGODA RIVER
MOUTH, EAST LONDON, SOUTH AFRICA
By
HERBERT CHRISTIAN KLINGER
South African Museum, Cape Town
&
BRIAN E. LOCK,
The University of Southwestern Louisiana, Lafayette
(With 7 figures)
LMS. accepted 20 September 1978]
ABSTRACT
The name Igoda Formation is proposed for a sedimentary sequence consisting mainly of
calcareous sandstones and arenaceous limestones exposed on the bluffs overlooking the
Igoda River Mouth near East London, South Africa. The invertebrate fauna indicates a Late
Cretaceous age, probably Late Campanian to Early Maastrichtian. Faunal paucity precludes
detailed comparison with other areas in southern Africa, but common faunal elements occur
at Lower Needs Camp (here regarded as a lateral facies equivalent of the Igoda Formation),
Pondoland, Zululand, Madagascar and Angola. Affinities with Madagascar are strongest.
CONTENTS
PAGE
INntroductiongwe-yar-e Mweta tee eee ial hn 71
The Igoda River Mouth section. . .... . 13
WnhesroadisectiOnmecs: isan eos ee) lee AS
Fauna and age of the Igoda Formation . .. . 75
Comparison with other areas in southern Africa .. 81
SUMMA TVs eee cee a erie Ain Vow tases Syl yhaeiy etabs 82
INCKNOWled SEMmentseame naan ee un yen Ney ieee unis ns 83
FE TErENCES he marry neg ee en er) cate Sheer 83
INTRODUCTION
Late Cretaceous fossiliferous limestones have long been known from the
eastern Cape from one of the small quarries on the farm Needs Camp (the
East or Lower Quarry) near East London (Lang 1908; Woods 1908; Chapman
1916). McGowran & Moore (1971) established a probable Upper Senonian
(Campanian to Maastrichtian) age for this deposit on the basis of the micro-
faunal content. Contrary to a report by King (1972), the second quarry at
Needs Camp (the West or Upper Quarry) is excavated in limestones of Tertiary
age (Lock 1973 and in preparation). Microfaunal investigations confirm this
Ann. S. Afr. Mus. 77 (5), 1978: 71-83, 7 figs.
71
YD ANNALS OF THE SOUTH AFRICAN MUSEUM
(W. G. Siesser pers. comm.), although reworked Cretaceous material occurs in
the limestones.
Until 1974 no other outcrops of Late Cretaceous marine sediments had
been reported from south of the Umzamba Estuary (Pondoland, Transkei).
Mountain (1974: 19) briefly described a new fossiliferous locality less than
15 km from Needs Camp as follows: The rock occurs in a ‘tiny exposure on
the East London—Kidd’s Beach road just west of the Goda River valley as the
road climbs up on to the coastal plain. It can be seen in the present road cutting
and consists of a friable limestone with abundant echinoid spines, Ostrea shells
and gastropod casts. It is possibly Cretaceous but no one has examined the
microfossils.’
Following landslides during the last few years, an extension of this occur-
rence became exposed in the bluffs overlooking the western shore of the lagoon
at Igoda Mouth, less than a kilometre from the roadside section mentioned by
Mountain (Fig. 1).
Tertiary and ?Pleistocene
calcareous sands
Igoda Formation
Beaufort sedimentary rocks
and Karroo dolerite
Fig. 1. Locality map, Igoda Formation. A. Roadside exposure reported by Mountain (1974);
note the new road under construction. B. Type section of the Igoda Formation. C. Reference
section. (See also Fig. 2.)
UPPER CRETACEOUS SEDIMENTS FROM THE IGODA RIVER MOUTH 13
THE IGODA MOUTH SECTION
The succession in the new exposures is summarized in Figure 2.
The Cretaceous sequence, here named the Igoda Formation, rests uncon-
formably on sandstones and mudstones of the Permian/Triassic Beaufort Group,
which have been intruded by dolerites of later Karoo age. The contact is one
of some relief. The type section (sections BI, B2) is a composite one (see Fig. |
for location and Fig. 2 for sections), and has at its base up to 50 cm of matrix-
supported small-pebble conglomerate, comprising well-rounded and _ well-
sorted brown-stained pebbles with a mean diameter of about 2 cm but with
individual pebbles up to about 5 cm long. Pebble lithologies consist of vein-
quartz and quartz arenites for the most part, although one pebble was a single,
well-rounded crystal of orthoclase 1,2 cm long.
The matrix is an arenaceous limestone with abundant glauconite. Some
shell fragments are present in this limestone matrix. The basal conglomerate
passes up into a sequence of arenaceous limestones and calcareous sandstones,
all with a high content of glauconite. As this section is followed up the slope,
exposures become very poor, and only a few of the more resistant calcareous
beds form ledges. At an altitude some 20m above the small-pebble conglomerate,
a second unconformity, at the base of the overlying Alexandria Formation (of
probable Tertiary age), is reached.
The middle portion of the Igoda Formation is better exposed some 30 m
to the south. In this second section a brown, small-pebble conglomerate,
indistinguishable from that just described, lies at the base of a similar sequence
of well-exposed glauconitic arenaceous limestones and calcareous sandstones.
As in the first section, these strata are fossiliferous, yielding a shelly fauna
dominated by ostreids which litter the surface. At this point, however, the
calcareous sequence has a much reduced thickness of just over 7 m, and the
small-pebble conglomerate overlies a 3-metre-thick sequence of white, poorly
consolidated, unfossiliferous sandstone and white, small-pebble conglomerates
which are in all other respects identical to the brown ones already described.
These three lithological associations are regarded as informal members,
and are known as the ‘white member’, ‘brown, small-pebble conglomerate
member’ and ‘calcareous member’ respectively.
The base of the Alexandria Formation is marked by another conglomerate
consisting of much larger pebbles (up to 30 cm in diameter) mostly of Beaufort
Group sandstone and siltstone. These pebbles are generally less resistant than
those found in the conglomerates of the Igoda Formation. The matrix of the
Alexandria Formation basal conglomerate is a coarse bioclastic limestone
containing fossil gastropods, especially Patella sp. and Conus sp., usually as
moulds. Glauconite is absent to rare. This unit is about 1,5 m thick and is
overlain by 10 m of cross-bedded, well-indurated, coarse bioclastic limestones.
Characteristically, the cross-bedding comprises a single tabular set, with
seaward dips of about 25°. Above this scarp-forming unit is a considerable
thickness (at least 30 m) of poorly consolidated calcareous aeolian sands.
ANNALS OF THE SOUTH AFRICAN MUSEUM
74
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UPPER CRETACEOUS SEDIMENTS FROM THE IGODA RIVER MOUTH Ud)
A second section was measured, 500 m closer to the sea (section C). Here
the white member is no longer present, and the calcareous member is even
further reduced in thickness (5,5 m). In other respects the sequence is similar
Geelkis. 2):
THE ROAD SECTION
The original outcrop described by Mountain (1974, see above) was identi-
fied, and the adjective ‘tiny’ found to be very appropriate. Only a few tens of
centimetres of very weathered glauconitic arenaceous limestone can be seen
overlying Beaufort Group sandstones and merging upwards into soil and surface
debris in the bank at the inland side of the road. The outcrop is very overgrown
and easily overlooked. Broken specimens of Rhynchostreon decussata and
Lopha spp., common forms in the main outcrop area, establish a correlation
with the lagoon-side exposures.
At the time of the most recent visit (May 1977), roadwork was in progress
for the straightening and improving of the main coastal road. Where this new
road reaches the approximate altitude of Mountain’s outcrop, it runs about
100 m inland of the latter, and cuts through a mass of dolerite. At the top of
the cutting at this point, a few rounded pebbles of Beaufort Group sandstone
in a calcareous cement were found. The size of these pebbles (20-30 cm) suggests
that they are remnants of the conglomerate from the base of the Alexandria
Formation. This would imply that the Igoda Formation pinches out at this
point (area A on map, Fig. 1).
FAUNA AND AGE OF THE IGODA FORMATION
In comparison with the Cretaceous sediments of Zululand and Transkei
(Kennedy & Klinger 1975 onwards), the invertebrate fauna of the [Igoda Forma-
tion is meagre and poorly preserved. Bivalves are the commonest group,
especially the ostreid forms, which have been preserved by virtue of their
unique shell mineralogy, followed by brachiopods, baculitid ammonites,
echinoids, and rare, normally coiled, ammonites, in that order of abundance.
At present, the material in the authors’ collections is too scant to merit
formal description and discussion, but a preliminary examination of the fauna
has revealed the presence of the following faunal elements:
FORAMINIFERA
Textularia sp.
COELENTERATA
‘Caryophyllia’ cf. arcotensis Forbes (Fig. 3)
ECHINODERMATA
Unidentifiable cidarid with uniserial pore pairs (Fig. 4B)
Crinoid stem ossicles of Pentacrinus type (Fig. 4A)
76 ANNALS OF THE SOUTH AFRICAN MUSEUM
levee, Shs ‘Caryophyllia’ arcotensis Forbes (Geology Department, Rhodes University.)
A-B. Lateral view. Scale bar 0,5 cm long. C. Dorsal view, stereopair. Scale bar 0,5 cm long.
UPPER CRETACEOUS SEDIMENTS FROM THE IGODA RIVER MOUTH HT
Fig. 4. A. Crinoid stem ossicle of the /socrinus type. B. Cidaris sp. indet.
Scale bars 0,5 cm long.
MOLLUSCA—CEPHALOPODA
Baculites subanceps Haughton (Fig. 5)
Eupachydiscus ? sp. (Fig. 6)
Pachydiscid sp. indet (compressed)
Saghalinites sp. cf. S. cala (Forbes) (Fig. 7A-B)
MOLLUSCA—GASTROPODA
Turritella (Zaria) cf. T. (Z.) besairiei Basse
MOLLUSCA-BIVALVIA
Rhynchostreon decussata (Goldfuss)
Lopha (Actinostreon) schnaebelei Basse (Fig. 7C—H)
‘Trigonia sp.
Spondylus douvillei Basse
Panopea cf. orientalis (Forbes)
‘Tnoceramus’ spp.
BRYOZOA
cf. Ceriopora micropora Goldfuss
‘“Membranipora cf. plebicola Brydone
BRACHIOPODA
cf. Terebratulina relicta Stoliczka
cf. Terebratula manuaensis Muir-Wood
cf. Rhynchonella natuans Stoliczka
Eolacazella affine (Bosquet)
78
ANNALS OF THE SOUTH AFRICAN MUSEUM
| J
Fig. 5. Baculites subanceps Haughton. A, C. SAM-—PCI5721. x 1,2.
B. SAM-PCI5723. x 1,0. D. SAM-PCI5728. x 1,0. E-H. SAM—PCI5906.
x 1,0. I-L. SAM-PCI5720. x 1,0.
UPPER CRETACEOUS SEDIMENTS FROM THE IGODA RIVER MOUTH 719
Fig. 6. Eupachydiscus ? sp. indet. SAM-—PCI5719. x 1,0.
Of the species identified, only a few can be used for accurate dating of the
Igoda Formation.
Baculites subanceps s.s. has been firmly dated as Late Campanian in
Angola (Howarth 1965), whilst the Pacific subspecies B. anceps pacificus occurs
in the Late Campanian of Japan and California. Saghalinites ranges from the
Santonian to Maastrichtian stages of the Late Cretaceous (Kennedy & Klinger
1977). S. cala, which the Igoda specimen resembles most, ranges from Cam-
panian IV to Maastrichtian II (sensu Kennedy & Klinger 1975) in Zululand but
is also known to occur in slightly older sediments in Pondoland (Transkei)
80 ANNALS OF THE SOUTH AFRICAN MUSEUM
Fig. 7. A-B. Saghalinites sp. cf. S. cala (Forbes). C-H. Lopha (Actinostreon) schnaebelei
(Basse). C. SAM-PCI5773. x 1,0. D. SAM-—PCI5749. x 1,0. E. SAM-—PCI5736. x 1,0.
F. SAM-PCI5750. x 1,0. G. SAM—PCI5752. x 1,0. H. SAM—PCI5744. x 1,0.
UPPER CRETACEOUS SEDIMENTS FROM THE IGODA RIVER MOUTH 81
(Middle Santonian—Lower Campanian) and southern India (Santonian or
Campanian). The ammonite genus Eupachydiscus is typically Late Cretaceous,
ranging from Coniacian to Maastrichtian, but the Igoda material is too poorly
preserved for specific identification.
Ceriopora micropora Goldfuss has been described from the type locality
of the Maastrichtian stage, but is also common in the Santonian and Campanian.
M. plebicola is from the Maastrichtian.
Eolacazella affine (Bosquet) occurs in the Maastrichtian of Western Europe,
and was also found at the Lower Needs Camp Quarry now dated as Upper
Campanian/Lower Maastrichtian by McGowran & Moore (1971).
These data all seem to suggest an age of Campanian/Maastrichtian for the
Igoda Formation with only a slight possibility of Santonian elements present.
COMPARISON WITH OTHER AREAS IN SOUTHERN AFRICA
The nearest onshore sediments of similar age occur at the Lower or Eastern
Quarry at Needs Camp. Eolacazella affine occurs at both the Igoda River Mouth
and at Lower Needs Camp, thus suggesting the two outcrops to be temporal
equivalents, though of different lithologies. In places, the outcrops at Lower
Needs Camp consist virtually of bryozoan limestone only. Exposures of similar
lithologies have since been found by one of the authors (H. C. K.) in roadside
excavations approximately half-way between the Upper and Lower Quarries,
illustrating that the Lower Needs Camp lithology has a much greater aerial
extent than previously suspected, and may yet prove to be a mapable unit to
confcrm with the requisites for formal recognition as a separate Formation by
the South African Committee for Stratigraphy.
The Lower Needs Camp sediments probably grade laterally into the slightly
deeper water facies of the Igoda Formation, but outcrops connecting the two
areas are, as yet, unknown.
Along the east coast of southern Africa, the closest Senonian sediments
occur at the Umzamba Estuary (Transkei), which have been firmly dated as
Mid-Santonian to Lower Campanian (Klinger & Kennedy 1977). Rare Saghali-
nites cala in the Umzamba Formation are comparable with Saghalinives sp. in
the Igoda Formation.
The closest, and thus far only known, onshore Upper Campanian/Lower
Maastrichtian sediments on the east coast of southern Africa occur in the
False Bay/St Lucia region of Zululand (Kennedy & Klinger 1975). Here Saghali-
nites cala occurs quite frequently with Nostoceras sp. and Eubaculites sp. in the
Upper Campanian and Lower Maastrichtian respectively. S. ca/la appears to
be the only identifiable faunal element in common between the two areas.
Lower Maastrichtian sediments are known from Cheringoma further north in
Mozambique, but to date only poorly-preserved Eubaculites species are known
(Crick 1923).
Much further north, the Campanian and Maastrichtian faunas of Mainti-
rano, Madagascar, as described by Basse (1931), strongly resemble those from
82 ANNALS OF THE SOUTH AFRICAN MUSEUM
Igoda especially as far as the thick-shelled ostreid forms are concerned, and to
a lesser extent with the Late Senonian fauna of the east coast Province of
Vatomandry described by Cottreau (1922), which is characterized by the paucity
of ammonites, similar to the Igoda Formation.
Towards the southern and western coast, the closest sediments of com-
parable age are found in the off-shore Alphard Group between Cape St Francis
and Cape Recife (Klinger et al. in prep.). Here, however, the dominant faunal
element is Eubaculites latecarinatus indicative of Early Maastrichtian age.
No macrofossils are common to both outcrops.
From here westwards, the first known Cretaceous on-shore sediments occur
near Bogenfels (South West Africa) (Klinger 1977), but these have been dated
definitely as Cenomanian, although some doubt still exists as to whether
Senonian strata are present on-shore or not.
The closest comparable deposits occur at Carimba in Angola, the type area
of Baculites subanceps, the latter being the only faunal element in common
between Igoda and Angola. The Cretaceous sediments at Carimba have been
firmly dated as Late Campanian by Howarth (1965).
SUMMARY
The name Igoda Formation is proposed for a sedimentary sequence
consisting mainly of calcareous sandstones and arenaceous limestones exposed
on the bluffs overlooking the Igoda River Mouth. The Formation is informally
divided into ‘white member’, ‘brown, small-pebble conglomerate member’ and
‘calcareous member’.
In comparison with the Cretaceous sediments of Natal and Transkei, the
Igoda fauna is meagre, consisting mainly of thick-shelled ostreid bivalves,
followed by brachiopods, baculitid ammonites, echinoids, and rare, normally
coiled ammonites, in that order of abundance.
These faunas point to an age of Late Campanian to Early Maastrichtian.
Due to the paucity of the faunas, comparisons with other temporally
equivalent areas are tenuous. The Lower Needs Camp Quarry deposits are of
similar age, and probably represent a shallower water and more restricted lateral
facies equivalent of the Igoda Formation. Saghalinites sp. cf. cala connects
the Igoda Formation to the Cretaceous deposits on the east coast of southern
Africa at the Umzamba Estuary and in the False Bay/Lake St Lucia region of
Zululand.
As far as the abundance of thick-shelled ostreids and relative scarcity of
ammonites is concerned, the Cretaceous deposits of Maintirano and Vato-
mandry in Madagascar show greatest affinity, but this should probably be
ascribed to the depositional environment rather than to more favourable
migration routes.
On the west coast of southern Africa, comparable deposits occur at Carimba
in Angola which provide a definite date for Baculites subanceps, i.e. Late
Campanian.
UPPER CRETACEOUS SEDIMENTS FROM THE IGODA RIVER MOUTH 83
ACKNOWLEDGEMENTS
We wish to express our gratitude towards the following persons for identi-
fying and commenting on some of the material: Mr I. McMillan (SOEKOR,
Johannesburg), Mr D. Salmon (Geological Survey, Cape Town) (Foraminifera),
Prof. Dr E. Voigt (Geologisch-Palaontologisches Institut, Hamburg—Bryozoa),
and Dr E. Owen (British Museum (Natural History), London—Brachiopoda).
Mr L. Matthews collected some of the specimens while a graduate student.
REFERENCES
Basse, E. 1931. Monographie Paléontologie De Crétacé de la Province de Maintirano, Mada-
gascar. Tananarive. Gouv. Gén. Madagascar et Dépend., Service des Mines.
CHAPMAN, F. 1916. Foraminifera and ostracoda from the Upper Cretaceous of Needs Camp,
Buffalo River, Cape Province. Ann. S. Afr. Mus. 12: 107-118.
CoTTREAU, J. 1922. Paléontologie de Madagascar. X. Fossiles crétacés de la cote orientale.
Annls Paléont. 11: 111-192.
Crick, G. C. 1923. Appendix A. On Upper Cretaceous Cephalopoda from Portuguese East
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S. Afr. 76: 1-5.
McGowran, B. & Moore, A. C. 1971. A reptilian tooth and Upper Cretaceous microfossils
from the Lower Quarry at Needs Camp. Trans. geol. Soc. S. Afr. 74: 103-105.
MOounrTAIN, E. 1974. The geology of the area around East London, Cape Province. An explanation
of sheet map 3227 D (East London), 3228 (Kei Mouth). Pretoria: Geological Survey.
Woops, H. 1908. Echinoidea, Brachiopoda and Lamellibranchiata from the Upper Cretaceous
limestone of Needs Camp, Buffalo River. Amn. S. Afr. Mus. 7: 13-19.
6. SYSTEMATIC papers must conform to the International code of zoological nomenclature
(particularly Articles 22 and 51). :
Names of new taxa, combinations, synonyms, etc., when used for the first time, must be
followed by the appropriate Latin (not English) abbreviation, e.g. gen. nov., sp. nov., comb.
nov., syn. noy., etc.
An author’s name when cited must follow the name of the taxon without intervening
punctuation and not be abbreviated; if the year is added, a comma must separate author’s
name and year. The author’s name (and date, if cited) must be placed-in parentheses if a
species or subspecies is transferred from its original genus. The name of a subsequent user of
a scientific name must be separated from the scientific name by a colon.
Synonymy arrangement should be according to chronology of names, i.e. all published
scientific names by which the species previously has been designated are listed in chronological
order, with all references to that name following in chronological order, e.g.:
Family Nuculanidae
Nuculana (Lembulus) bicuspidata (Gould, 1845)
Figs 14-15A
Nucula (Leda) bicuspidata Gould, 1845: 37.
Leda plicifera A. Adams, 1856: 50.
Laeda bicuspidata Hanley, 1859: 118, pl. 228 (fig. 73). Sowerby, 1871: pl. 2 (fig. 8a—b).
Nucula largillierti Philippi, 1861: 87.
Leda bicuspidata: Nicklés, 1950: 163, fig. 301; 1955: 110. Barnard, 1964: 234, figs 8-9.
Note punctuation in the above example:
comma separates author’s name and year
semicolon separates more than one reference by the same author
full stop separates references by different authors
figures of plates are enclosed in parentheses to distinguish them from text-figures
dash, not comma, separates consecutive numbers
Synonymy arrangement according to chronology of bibliographic references, whereby
the year is placed in front of each entry, and the synonym repeated in full for each entry, is
not acceptable.
In describing new species, one specimen must be designated as the holotype; other speci-
mens mentioned in the original description are to be designated paratypes; additional material
not regarded as paratypes should be listed separately. The complete data (registration number,
depository, description of specimen, locality, collector, date) of the holotype and paratypes
must be recorded, e.g.:
Holotype
SAM-—A13535 in the South African Museum, Cape Town. Adult female from mid-tide region, King’s Beach
Port Elizabeth (33°51’S 25°39’E), collected by A. Smith, 15 January 1973.
Note standard form of writing South African Museum registration numbers and date.
7. SPECIAL HOUSE RULES
Capital initial letters
(a) . The Figures, Maps and Tables of the paper when referred to in the text
e.g. *... the Figure depicting C. namacolus ...’; *. . . in C. namacolus (Fig. 10)...’
(b) The prefixes of prefixed surnames in all languages, when used in the text, if not preceded
by initials or full names
e.g. DuToit but A.L.du Toit; Von Huene but F. von Huene
(c) Scientific names, but not their vernacular derivatives
e.g. Therocephalia, but therocephalian
Punctuation should be loose, omitting all not strictly necessary
Reference to the author should be expressed in the third person
Roman numerals should be converted to arabic, except when forming part of the title of a
book or article, such as
“Revision of the Crustacea. Part VIII. The Amphipoda.’ ;
Specific name must not stand alone, but be preceded by the generic name or its abbreviation
to initial capital letter, provided the same generic name is used consecutively. }
Name of new genus or species is not to be included in the title: it should be included in the
abstract, counter to Recommendation 23 of the Code, to meet the requirements of
Biological Abstracts.
SMITHSONIAN INSTITUTION LIBRARIES
“HOUMA
3 9088 01206 6650
HERBERT CHRISTIAN KLINGER
&
~ BRIAN E. LOCK
UPPER CRETACEOUS SEDIMENTS FROM THE
IGODA RIVER MOUTH, EAST LONDON,
SOUTH AFRICA