BULLETIN
du MUSÉUM NATIONAL
d’HISTOIRE NATURELLE
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PUBLICATION BIMESTRIELLE
zoologie
82
JANVIER-FÉVRIER 1973
N° 108
BULLETIN
du
MUSÉUM NATIONAL D’HISTOIRE NATURELLE
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BULLETIN DU MUSÉUM NATIONAL D'HISTOIRE NATURELLE
3 e série, n° 108, janvier-février 1973, Zoologie 82
Tortricidae of the Andringitra Range,
Central Madagascar (Lepidoptera) 1 .
Part 1. Tortricinae
by Alexey Diakonoff *
Abstract. —• The présent paper contains records of 46 species, belonging to 13 généra, of
which one genus, 14 species and one subspecies are described as new. Sketches of head and wing
neuration of the new genus and figures of genitalia of new species are presented.
Résumé. — Dans la présente note 46 espèces de Tortricidae Tortricinae capturées dans le
massif de l’Andringitra (Madagascar Centre) sont étudiées. Elles appartiennent à treize genres,
dont un est nouveau. Quatorze espèces et une sous-espèce sont décrites comme nouvelles. Des
dessins de la tête et de la nervation des ailes du nouveau genre sont données, ainsi que les figures
des genitalia des nouvelles espèces.
Dr P. E. L. Viette kindly entrusted me with the studv of rich and excellently pre-
served material of the Muséum national d’Histoire naturelle of Paris, from the Andrin¬
gitra Range, Central Madagascar.
The greatest part of this material originates from two localities, viz. : (1) Andringi¬
tra méridional, chaîne de l’Andrinony, cirque de Manjarivolo, 1 640 m, 26.x-4.xi.1970
(P. Viette) and (2) Andringitra oriental, forêt d’Anjavidilava, 1 975, 1 990, 2 000 and
2 005 m, 19-25. xii and 1-15.1.1970-1971 (P. Griveaud). For the sake of brevity these
localities are indicated below simply as Manjarivolo and Anjavidilava, respectively, fol-
lowed by altitude and date, except under descriptions of new species. Ail other localities
are cited in full.
An interesting geomorphological, climatological and botanical survey of the Andrin¬
gitra Range of the hand of a group of scientists has appeared recently (Pauliax et al ., 1971).
Digitosa leptographa Diak.
Diakonoff, 1960 : 24, fig. 17-18, pl. 3, fig. 15, pl. 41, fig. 16-17. — 1961 : 152. — 1970 : 104.
Distribution. East Madagascar : Ambatondrazaka ; Andranomalaza ; Sandrangato
Forest. — North Madagascar : Tsaratanana Range.
1. Research project (Recherche Coopérative sur Programme il 0 225) of the Centre national de la Recher¬
che scientifique). See : Bull. Soc. Écologie, 1971, 2 (2-3) : 189-266.
* Department of Entomology, Rijksmuseum van Natuurlijke Historié, Raamsteeg, 2, Leiden, Nether-
lands.
108, 1
106
ALEXEY DIAKONOFF
Anjavidilava 1 975-2 005 m, 19-25.xn. 1970-1-15.1.1971 (P. Griveaud), 7 12 Ç,
genit. slide 8275.
Manjarivolo (P. Viette), 1
Digitosa metaxantha Diak.
Diakonoff, 1960 : 27, pl. 3, fig. 13-14 {$).
Distribution. East Madagascar : Andranomalaza.
Anjavidilava, 2 005 ni, 1-15.1.1971 (P. Griveaud), 1
Manjarivolo [P. Viette), 1 $, genit. slide 8532.
Parapandemis griveaudi Diak.
Diakonoff, 1960 : 45, fig. 42-43, pl. 6, fig. 33-33a, pl. 8, fig. 44-45, pl. 9, fig. 48. — 1961 : 152. —
1970 : 105.
Distribution. East Madagascar : Ambatondrazaka ; South Moramanga ; Itala-
viana ; Marojejy Range ; Sandrangato Forest. Central Madagascar : Betsileo country ;
Nature Reserve III. North Madagascar : Tsaratanana Range, 1 900-2 362 m.
Manjarivolo (P. Viette), 3
Parapandemis plutosema Diak.
Diakonoff, 1960 : 49, pl. 8, fig. 41-43. — 1961 : 152.
Distribution. Central Madagascar : Ankazobe district ; Ambatondrazaka district ;
Manjakandriana district ; Ambatolampy district ; Ambalavao district.
Anjavidilava, 1 975-2 005 m, 19.xn.1970-15.1.1971 (P. Griveaud), 12 <?, 3 Ç.
Manjarivolo (P. Viette), 1 <$.
Parapandemis retroflua Diak.
Diakonoff, 1960 : 53, fig. 48-49, pl. 10, fig. 55-56. — 1970 : 105.
Distribution. Central Madagascar : Ankaratra Range, Ambatolampy district.
East Madagascar : Marojejy Range ; Moramanga district. North Madagascar : Tsara¬
tanana Range, up to 2 000 m.
Anjavidilava, 1 990, 2 000 and 2 005 m, 19-25. xii and 1-15.1.1970-1971 ( P. Griveaud),
8 <J.
Manjarivolo (P. Viette), 3 <$, 1 $.
Parapandemis subovata Diak.
Diakonoff, 1970 : 109, fig. 6, pl. 7, fig. 4-5.
Distribution. North Madagascar : Tsaratanana Range, 1 900-2 050 m.
Manjarivolo (P. Viette), 2
TORTRICIDAE OF THE ANDRINGITRA RANGE
107
Parapandemis capnobathra (Meyr.)
Homona capnobathra Meyrick, 1930 : 605. — Viette, 1947 : 56. — Clarke, 1955 : 73. — 1958 :
148, pl. 74, fig. 2-2b.
Parapandemis capnobathra : Diakonoff, 1960 : 52, pl. 7, fig. 4, pl. 9, fig. 49-50 (rf). — 1970 : 106.
Parapandemis euphana Diakonoff, 1960 : 59, pl. 38, fig. 252-253 (Ç).
Distribution. East Madagascar : Moramanga, Ankasoka. North Madagascar :
Tsaratanana Range.
Manjarivolo (P. Viette), 2 Ç.
Parapandemis sclerophylla Diak.
Diakonoff, 1960 : 60, pl. 14, fig. 80-81. — 1970 : 111.
Distribution. Central Madagascar : Nature Reserve III, Ivelona Valley.
Anjavidilava, 1 990, 1 975 and 2 005 m, 19.xn.1970-15.1.1971 (P. Griveaud), 13
Parapandemis straminocula plectopa Diak.
Diakonoff, 1960 : 71, pl. 11, fig. 67-68, pl. 13, fig. 78. — 1970 : 112.
Distribution. Central Madagascar : Ankazobe, Ambohitantely ; Ambatolampy.
East Madagascar : Ankasoka. North Madagascar : Tsaratanana Range.
Anjavidilava, 1 990 m, 19-25 .xii.1970 (P. Griveaud), 1 <$.
Parapandemis euryloncha sp. nov.
(Fig. 1-2)
c? R> mm. Head whitish, vertex sufïused with pale ochreous. Palpus pale ochreous,
terminal segment rather long. Thorax pale grey. Abdomen whitish, touched with fuscous,
glossy.
Fore wing rather broad, subtruncate, costa arched anteriorly, almost straight, hardly
curved posteriorly, apex moderately pointed, termen sinuate above, vertical, moderately
rounded below. Ochreous-white with a golden gloss, markings light fuscous mixed with
ferruginous, strewn with a few black points. Rasai patch to about 1/4, edge angulate
in middle, moderately concave above and below this ; central fascia from well before middle,
oblique, gently dilated downwards but sufïused and obliterate below middle of cell, only
posterior half of fascia extending as far as end of cell, rather tinged fulvous, with three large
black dots in a horizontal row in middle of cell ; costal patch semioval, paler, grey-fuscous,
including two spots of ground colour on Costa, top touching upper angle of cell and with
that, the central fascia ; an oblique blotch from beyond middle of posterior edge of costal
patch to 2/3 of termen ; apex and upper half of termen sufïused with fuscous-ferruginous ;
entire wing finely marbled with darker fuscous, forming short strigulae, interconnecting
other markings. Cilia ochreous-fuscous (damaged).
108
ALEXEY DIAKONOFF
IIind wing pale ochreous with a golden gloss, apical third suffused with tawny and
indistinctly dotted with fuscous. Cilia pale ochreous, around apex and along upper half
of termen infuscated.
Male genitalia. Very near those of P. marginumbra Diak., but with a distinctly broa-
der, rhomboidal uncus and a slightly longer sacculus, so that lower angle (top) of valva
in more pointed.
Madagascar Centre, Andringitra oriental, forêt d’Anjavidilava, 2 005 m, 1-15.1.1971
(P. Griçeaud), 1 (J, holotype, genit. slide 8280.
A single, rather rubbed specimen, with the genitalia very close to those of marginum¬
bra. However, the broader uncus, together with the completely different colouring and
markings, compel me to separate this species.
Parapandemis xanthacra Diak.
Diakonoff, 1960 : 66, pl. 12, fîg. 69-70, pl. 13, fig. 77 (<?$). — 1970 : 111.
Distribution. East Madagascar : Moramanga. North Madagascar : Tsaratanana
Range.
Manjarivolo, 1 650 m, 26.x-4.xi.1970 (P. Viette & P. Griveaud), 1 2, genit. slide
8277.
Madagascar Nord, massif du Tsaratanana, piste de Mangindrano, au Maromokotra,
au sud du piton coté 2 831 m, 13-15.xi. 1966, 2 500 m, “ Mission du Tsaratanana xi.1966,
camp n°. 3 (P. Griveaud, P. Soga, P. Viette et D. Wintrebert) ”, 1 $, genit. slide 8276.
Note : This specimen lias been erroneously attributed by me to Niphothixa atava
Diak. (paratype : Diakonoff, 1970 : 114, line 10 from top).
Cuspidata (Cuspidata) ditoma peratra subsp. nov.
çj 16 mm. Head and thorax grey-purple. (Palpi missing.) Abdomen fuscous.
Fore wing truncate, broad, Costa strongly sinuate : obliquely arched anteriorly, roun-
ded-prominent at 2/5, concave and impressed before apex, apex pointed and produced,
termen sinuate, moderately oblique. Evenly deep blackish-purple, rather lustreless.
Central fascia faintly indicated by dull darker purple colour, from 3/5 of Costa to 3/4 of
dorsum, rather straight, narrower along upper third, moderately dilated in middle ; on
Costa this fascia followed by an oblique blotch of faintly glossy anthracite colour. a trifle
paler than ground colour and hardly visible in certain lights, reaching halfway across wing ;
other markings indefinite. Cilia concolorous.
Hind wing rather light fuscous-grey, darker fuscous-purplish towards apex and on
posterior half. Cilia pale fuscous, with a darker subbasa'l band (imperfect).
Male genitalia. Similar to those of the nominate form.
TORTRICJDAE OF THE ANDRINGITRA RANGE
109
Anjavidilava, 2 005 m, 1-15.1.1971 (P. Griveaud), 1 <$, holotype, genit. slide 8535.
The single specimen available is so distinct from the earlier described ditoma (a widely
distributed species of middle high altitudes, 800-950 m), with regard to the colouring, mar-
kings and the shape of the fore wing, that I do not venture to include it, but prefer to regard
the only male a melanistie subspecies of higher altitudes. The genitalia are in ail details
similar to those of ditoma. Future eollecting will provide additional proof, I liope.
Cuspidata (Cuspidata) castanea Diak.
(Fig- 3)
Diakonoff, 1960 : 187, pl. 15, fïg. 88-89 ($).
Distribution. Central Madagascar : Manjakandriana ; Ambatondrazaka.
Anjavidilava, 2 005 m, 1-15.1.1971 (P. Griveaud), 1 <§, gent. slide 8273.
Ç 20-22 mm, apallotype 1 . Head, palpus and thorax rather deep tawny-fulvous,
palpus actually dusted with dark grey (shorter and more triangular than in C. anthracitis).
Abdomen with venter dull light ochreous, sides glossy whitish, dorsum infuscated.
Fore wing oblong, rather dilated, Costa sinuate throughout (because of the prominent
costal cilia at apex), apex rectangular, termen slightly sinuate, vertical above, rounded
and slightly prominent below. Rather deep tawny-fulvous with vinaceous tinge, obscured
by grey-purplish darker sufïused marbling and strewn with minute transverse dark stri-
gulae ; a group of sparse scattered dark purple transverse strigulae of slightly elevated
scales over anterior half of dise above middle, not reaching Costa and an oblique and slightly
curved regular sériés of similar deep purple points from below 2/3 of Costa reaching to
middle of dise and apparently following anterior edge of (indefinite) costal patch ; a slightly
paler fulvous, curved band from 3/5 of costa, just preceding above mentioned sériés of dots ;
termen and tornus a trifle paler and faintly orangeish tinged. Cilia fulvous, lighter around
apex and along upper half of termen, basal third throughout deeper grey-purple.
Hind wing light yellow, brighter than in male, a trifle paler towards base and dorsum,
apex irregularly spotted with dark grey, spots quickly becoming sparse and small anteriorly
and along termen. Cilia pale yellow, along upper half of termen and around apex light
grey, with a blackish subbasal band.
Female genitalia : Characteristic by the throughout sclerotized sterigma, with lamella
postvaginalis at the sides high and broad, somewhat resembling that in C. ditoma Diak.
(but there sterigma subhyaline and lamella antevaginalis dilïerently shaped), and also
by the presence of a slender, serrulate dark cestum with about fîve coils (absent in most
other species) ; ductus busae also with five coils only.
Anjavidilava, 1 990 m, 19-25.xii.1970 (P. Griveaud), 1 Ç, apallotype, genit. slide
8531 ; the same, 1 Ç.
1. The new term apallotype (= apo-allotype) is proposed for the substitution of the debatable term
“ neallotype
108, 2
TORTRICIDAE OF THE ANDRINGITRA RANGE
111
Cuspidata (Cuspidata) micaria sp. nov.
(Fig. 4-6)
G 16-17 mm, $ 16-19 mm. Head and thorax deep fulvous. Palpus paler fulvous,
slightly sufïused with grey. Abdomen pale yellowish-ochreous.
Fore wing oblong-truncate, rather broad, dilated along basal third, Costa sinuate,
being arched along basal 2/3, apex gently prominent and pointed, termen sinuate, rounded
below, vertical. Ferruginous, indistinctly spotted with darker, roundish spots which
actually are formed by deep ferruginous scales with narrow whitish bases (giving striated
appearance to spots) ; a sufïused straight, oblique, pale fulvous fascia from Costa well
beyond middle, towards tornus, rather well-defined along upper half of wing, but edges
gently sufïused ; below this fascia disappears in pale fulvous posterior third or fourth of
wing which does not reach wing margin and is traversed by sufïused ferruginous streaks
along veins ; sometimes origin of fascia obscured by ground colour ; posterior fourth of
wing with numerous black crescentic strigulae tending to form oblique parallel sériés ;
anterior half of dorsum with variably extended, more or less contiguous spotting with bright
orange, strewn with reddish scales ; these spots round, arranged in more or less vertical
sériés, obliterate above fold, except a pair at 1/3 and a single spot at 2/3. Cilia deep ferru¬
ginous.
Male genitalia. Nearest to those of C. ditoma Diak., but difïering by the slightly more
gradually narrowed sacculus, with the concavity of its upper side more basal, and by the
less curved aedeagus.
Female genitalia difïering from those of ditoma by a much shorter and less coiled duc-
tus bursae.
Andringitra oriental, forêt d’Anjavidilava, 1 990 and 2 005 m, 19-25.xii.1970 and
1-15.1.1971, respectively (P. Griveaud ), 4 10 $. Closely allied with C. (C.) ditoma
Diak., but superficially quite distinct by much darker colouring, continuous fasciate dark
anterior edge of pale band (in ditoma reduced to a small costal triangle) and strong réduc¬
tion of posterior edge of that band (in ditoma usually a sériés of dark purple dots), and
presence of orangeish dotting on dorsum.
Madagascar Centre, Andringitra méridional, chaîne de l’Andrianony, cirque de Manja-
rivolo, 1 650 m, 26.x-4.xi.1970 (P. Viette), 1 <$, genit. slide 8332 ; the only specimen with
distinctly darker, fuscous-purplish hind wings, but otherwise similar to the holotype.
Cuspidata (Cuspidata) oligosperma Diak.
Diakonoff, 1960 : 83, fig. 54-55, 58-62, pl. 16, fig. 94. — 1970 : 114.
Distribution. East Madagascar : Analamazoatra ; Andranomalaza ; Sandrangato
Forest. North Madagascar : Montagne d’Ambre ; Tsaratanana Range. Central Mada¬
gascar : Ambatondrazaka district ; Ambatolampy district.
Anjavidilava, 2 005 m (P. Griveaud), 1 genit. slide 8272.
112
ALEXEY DIAKONOFF
Cuspidata (Cuspidata) anthracitis Diak.
Diakonoff, 1960 : 88, pl. 15, fig. 93. — 1970 : 144.
Distribution. East Madagascar : Ambatondrazaka. North Madagascar : Tsara-
tanana Range.
Anjavidilava, 1 975, 1 990, 2 005 m, 26-30. xii. 1970,19-25. xii. 1970, 1-15.1.1971 (P. Gri-
veaud), 6 <^, 4 $, genit. slide 7307 (J.
Cuspidata (Pilophorica) leptozona Diak.
Diakonoff, 1960 : 90, fig. 7-8, 67a, pl. 16, fig. 99-100. — 1963 : 134.
Distribution. East Madagascar : Analamazoatra, Sandrangato. Central Mada¬
gascar : Ankaratra Range.
Anjavidilava, 1 990 m, 19-25. xii. 1970 (P. Griveaud), 1 c?.
Metamesia synclysa sp. nov.
(Fig. 7-10)
cj 17-18 mm. Head whitish. Palpus light fuscous, inner side whitish. Thorax
pale grey, tegula infuscated. Abdomen pale fuscous-greyish, anal tuft whitish-ochreous
tinged.
Fore wing oblong-oval, Costa considerably and gradually curved from base to apex,
apex pointed and slightly prominent, termen strongly sinuate above, flatly rounded below,
oblique. Even, rather pale grey, Costa narrowly sufïused with dark grey. Markings
formed by grey-fuscous suffusion, but little contrasting. Basal patch to 1/4, hardly per¬
ceptible in holotype (indefînite in paratype), edge somewhat, irregular, outwards-oblique ;
central fascia from well before middle, to end of dorsum, moderately broad, slightly dilated,
anterior edge entire, sufïused, gently convex, posterior edge with a small tooth in middle,
gently concave and ill-defined above this, somewhat irregular below ; costal patch rather
small, oblong, obliquely triangular, top rounded ; this patch almost confluent with a large
triangular spot along termen, with truncate top (in paratype smaller, well-separate from
costal patch) ; these two markings rather sufïused. Cilia concolorous but paler and more
glossy.
Hind wing whitish, touched with creamy and very pale fuscous (in paratype with a
golden gloss). Cilia concolorous.
Male genitalia. Resembling those of M. nolens Diak. most and similarly rather strongly
sclerotized, but uncus longer, rounded, with a not dilated, but slightly notched top ; gna-
thos stronger, arms broader, along their bases and basal halves denticulate outwardly.
Labis oblong-oval, denticulate on top, denticles finer than in nolens. Valva similar. Aedea-
114
ALEXEY DIAKONOFF
gus long and straight, basal third slightly dilated, caecum rounded, little oblique ; a small
group of denticles on upper side just before top and bases of three large cornuti percep¬
tible.
Andringitra centre, cirque Boby, 2 500 m, 24-28.xi.1970 (P. Soga), 1 <$, holotype,
genit. slide 8495 ; 1 <$, paratype, genit. slide 8497.
Metamesia leptodelta sp. nov.
(Fig. 11-12)
Ç 16 mm. Ilead and palpus light brown. Thorax brown-fuscous. Abdomen grev.
Fore wing narrow, Costa curved along basal half, apex acutely pointed, termen con¬
cave, very oblique, lower third rounded-prominent. Pale ochreous, rather densely irro-
rated with light brown and light fuscous, markings rather dark brown. Basal patch to
1/4, darker ochreous-tawny, denser irrorated with fuscous, but still little contrasting ;
its edge with upper half little outwards-oblique and straight, lower half concave, almost
vertical, central fascia from beyond 2/5, moderate, hardly dilated and rather straight
to end of dorsum ; costal patch moderate and rather narrow, obliquely triangular, eonnec-
ted with fascia above its middle ; a slender outwards-concave preterminal fascia from
apex, gradually dilated downwards and merging in lower end of central fascia. Cilia
pale ochreous.
Ilind wing whitish-ochreous, vein 1 b and apex gently infuscated.
Female genitalia. Resembling those of M. peracuta Diak., but differing as foliows :
lobes of 9th sternite short, by far not meeting ventrallv (in peracuta touching each other
in middle). Lamella postvaginalis, a sclerotized bar, narrower in middle (in peracuta
membraneous) ; lamella antevaginalis rather similar, but at the sides upper edge distinctly
incised, incision curving downwards ; colliculum shorter. Signum shorter. Also resem¬
bling the genitalia of M. retrocitra Diak., but sterigma much more robust, colliculum wider
and especially : signum quite distinct, cone-shaped (in retrocitra blade-like).
Andringitra central, cirque Boby, 2 500 m, 24-28.x.1970 (P. Soga), 1 $, holotype,
genit. slide 8336. A slender, narrow-winged species, with long acute apex, superficially
surprisingly similar to Goniotorna (Tenuisaccula) mianta sp. n. male, but by doser inspec¬
tion revealing its true nature.
Anthophrys spectabilis Diak.
Diakonoff, 1960 : 122, fig. 143-154, pl. 22, fîg. 74. — 1970 : 116.
Distribution. North Madagascar : Tsaratanana Range. Central Madagascar :
Betsileo Country ; Andringitra Range, Anjavidilava. East Madagascar : Moramanga
district ; Ambatondrazaka district ; Marojejy Range.
TORTRICIDAE OF THE ANDRINGITRA RANGE
115
Anjavidilava, 1 975, 1 990, and 2 005 m (P. Griveaud). 32 <$.
Ouest du Vohidray (Andringitra), Ambalamarovandana, 1 530 m, 16-21.1.1971 (P. Gri¬
veaud), 1 <$.
Manjarivolo (P. Viette), 1
Goniotorna (Oestophyes) verticillata, group of species
This group of closely allied and rather similar species présents considérable difficul-
ties. Described after very few specimens, the species showed sulïicient génital différences
in the two sexes. Now that good sériés of each species are available, they prove that the
species indeed are distinct ; that they are little variable and that the génital characters
are constant. Still their discrimination is far from easy, because both the superficial
and the génital différences are fairly small. Therefore I am presenting elaborate drawings
of the genitalia.
In the males the most important parts are the shape of the transtilla with the labides,
the sacculus and the shape and armature of the aedeagus. In the females the shape of the
entire sterigma is characteristic.
The transtilla is mostly shaped as a slender transverse rod, with central part turning
down and acculade-shaped, while the labides, rising at each end of the transtilla, are dou¬
ble : a ventral arm, usually broader (in mount situated on top of the other arm) and a more
dorsal, often slender and dentate labis proper (in mount, underneath the ventral arm).
Each of the pairs has a spécifie shape.
Goniotorna (Oestophyes) praerupta Diak.
Diaiconoff, 1960 : 136, pl. 24, fig. 159, 159a.
Distribution. East Madagascar : Anosibe road, Sandrangato forest.
Anjavidilava, 1 990 m (P. Griveaud ), 1 (J, genit. slide 8337 ; 1 genit. slide 8534.
Goniotorna (Oestophyes) trigodes sp. nov.
(Fig. 43-46)
<$ 19 mm. Head pale tawny, faintly spotted darker. Palpus tawny, towards apex
sufîused with blackish. Thorax pale ochreous, anterior third suffused with fulvous-tawny.
Abdomen pale ochreous, rather suffused with greyish, anal tuft grey.
Fore wing truncate, rather broad, distinctly dilated, Costa arched almost throughout,
straight only before apex, apex subobtuse, a tri fie projecting, termen very gently sinuate
and vertical above, rounded beneath. Pale ochreous-tawny, markings much suffused
deeper fulvous-tawny, partly ill-defmed. Basal fourth of Costa with a suboval suffusion ;
central fascia from just before middle of Costa to 3/4 of dorsum, light tawny, densely but
116
ALEXEY DIAKONOFF
faintly marbled with deeper brownish-tawny ; narrow along upper fourth, dilated 4 X and
roundish in dise, moderately narrowed again on dorsum, anterior edge with a few irregular
slender projections below, posterior edge sufîused and somewhat irregularlv scalloped,
convex except above ; coarse and sparse transverse marbling on dorsum between base
and central fascia, partly connected with latter and with a more or less isolated spot on
upper edge of cell at 1/3 of wing ; costal patch ill-defined, represented by two tawny-ful-
vous costal spots, on ends of each veins 11 and 10 ; apex from 2/3 of Costa to tornus suffu-
sed with deeper tawny-fulvous and faintly marbled with still deeper fulvous, this darker
marbling tending to form two curved subapical séries of spots. Cilia light tawny-fulvous
(imperfect).
Ilind wing whitish-ochreous with a golden gloss, apex gently sufîused with deeper
ochreous-yellow. Cilia eoncolorous.
One paratype, genit. slide 8290 <$, 22 mm, evenly deeper vinaceous tinged, with mar¬
bling rather obscured and indefinite, but central fascia of similar shape, only darker and
well-defined ; posterior third of costa of similar cloudy deep vinaceous suffusion, irregular
vertical sériés of purple points before termen. Hind wing with somewhat deeper vina-
ceous-suffused apex. The genitalia similar to the type, except tip of lahis which is fînely
spinose (instead of single). Otherwise similar to the type, also the characteristic serrulate
edge of sacculus.
Male genitalia. Rather similar to those of G. (O.) irresoluta Diak., but with shorter
and more robust labis, strongly spined on top (in paratype with only a couple of spines),
and differing from ail other species of the subgenus by fmely serrate edge of sacculus. Pul-
vinus long-haired. Aedeagus with a dark and fiat apical bicuspidate process.
Madagascar Centre, Andringitra central, forêt d’Anjavidilava, 1 990 m, 19-25.xn. 1970
(P. Griveaud ), 1 <J, holotype, genit. slide 8525 ; 1 paratype, genit. slide 8519.
Madagascar Centre, Andringitra méridional, chaîne de l’Andrianony, cirque de Man-
jarivolo, 1 650 m, 26.x-4.xi.1970 (P. Viette ), 2 <§, paratypes, genit. slides 8290 and 8346.
A distinct species, recognizable by pale fulvous colouring and pale yellowish hind
wings.
Goniotorna (Oestophyes) irresoluta irresoluta Diak.
(Fig. 13-14, 18-19, 25-26)
Diakonoff, 1960 : 137, pl. 24, ûg. 158, 188a ((J).
Goniotorna ( Oestophyes ) mesostena Diakonoff, 1963 : 134, pl. I, fig. 3, pl. II, fig. 1-2 (Ç). Syn. nov.
Distribution. East Madagascar : Ampitameloka, 840 m ; Périnet ; Andranoma-
laza.
Anjavidilava, 1 975-1 990 m, 19-30.xn.1970 (P. Griveaud), 17 çj, 11 Ç, genit. slides,
£ 8340, $ 8293, 8352.
A somewhat variable species, distinct, however, by the well-defined edges of the three
usual markings : the basal patch, the central fascia and the costal patch. Especially
118
ALEXEY DIAKONOFF
Fig. 20-26. — Genitalia of Madagascan Tortricinae. 20, Goniotorna (G.) insatiata sp. nov., (J, holotype ;
21, the same, aedeagus ; 22, the same, Ç, allotype ; 23, the same, ductus bursae ; 24, the same, bursa ;
25, G. (Oestophyes) irresoluta Diak., <J, genit. slide 8340 ; 26, the same, aedeagus.
TORTRICIDAE OF THE ANDRINGITRA RANGE
119
the inner edge of the latter, being complété and well-defîned, forms a good characteristic
for the séparation of this species from rather similar G. (G.) mucida Diak., where ail the
markings are more or less suffused, most edges being either incomplète or entirely obli-
terate.
The species appears variable because of the edges of the usual markings being either
rather straight or more or less strongly undulate. So the edges of markings are tolerably
straight in the type specimen (<$) from 840 m, and also in the type ($) of G. (O.) mesostena
Diak., also from low altitude (Permet). Having been able to study a nice sériés of males
and females now, I am satisfied that mesostena is the female of the présent species and
the name, a junior synonym. Some males and one female of Anjavidilava hâve the edges
of markings ahnost as straight, but ail the remaining specimens hâve them undulate to
a variable extent. Ail specimens are rather darkly and distinctly marked, females darker
tlian males.
Goniotorna (Oestophyes) verticillata Diak.
(Fig. 47-48)
Diakonoff, 1960 : 138, pl. 24, fig. 157, 157a, 157b, pl. 25, fig. 161-162.
Distribution. Central Madagascar : Ankaratra Range ; Andringitra Range.
Anjavidilava, 1 975-2 005 m, 19 .xii.1970-15.1.1971 (P. Griveaud), 11 21 Ç.
Andringitra central, plateau d’Andohariana (Soaindrana), 2 030 m, 16-21.xi.1970
(P. Vielle), 2 cJ.
Goniotorna (Oestophyes) deinozona sp. nov.
(Fig. 27-32)
^ 19-20 mm. Head and palpus rather deep vinaceous-fulvous. Thorax deep vina-
ceous-brown, mixed with fuscous-grey, tip blackish. Abdomen tawny-vinaceous at base,
towards apex becoming densely suffused with anthracite-grey.
Fore wing oblong, moderately broad, Costa curved anteriorly, straight posteriorly,
subobtuse, termen gently sinuate and vertical above, rounded beneath, fulvous-vinaceous-
brown, moderately dusted and transversely strigulated with black, hardly so anteriorly,
markings formed by black dusting. Rasai patch over 1/4, hardly darker by very fine
blackish suffusion and délicate transverse strigulation, stronger on costal edge ; edge of
patch narrow, blackish, well-defined, acutely angulate above middle, rectangularly pro¬
minent on dorsum, emarginate just above angulation and prominence, respectively ; central
fascia from before middle of Costa, occupying about posterior third of dorsum, moderate
as far as edge of cell, thence twice as broad, anterior edge rather straight ; this fascia sligh-
tly obscured by black above, becoming gradually darker downwards, lower half almost
completely black ; anterior edge marked with a few single white scales ; costal patch repla-
ced by fine transverse strigulation along posterior 2/5 of costa over about upper half of
wing ; an oblique black suffused and spindle-shaped streak, resting on termen above tor-
TORTRICIDAE OF THE ANDRINGITRA RANGE
121
nus, including some three minute white scales (and being lower part of edge of costal patch).
Cilia concolorous, rather sufïused with blackish.
Hind wing rather dark purplish-grey, with little gloss, cilia grey, more glossy, a pale
basal and a grey subbasal lines.
Ç 21-22 mm. Deeper vinaceous-brown, stronger strigulated with blackish throughout,
basal patch without notches nor dorsal prominence ; central fascia somewhat more slender ;
posterior fascia replaced by a faint dark grey sufïused band from 3/5 of Costa to before
lower third of termen ; white scales very few, spread, dark-edged. Hind wing slightly
lighter and more glossy. Otherwise as male.
Male genitalia. Very similar to those of G. (O.) verticillata Diak., difïering chiefly
by the slender single-pointed and not dorsally dilated labis. Aedeagus less slender and
shorter than in that species, with a subapical sclerotized fold of the upper side, with a
finely serrate frontal edge.
Female genitalia. Allied to those in verticülata but quite distinct. Ostium broadly
funnel-shaped and tolerably symmetrical, moderately narrowed along lower third only
(in verticillata asymmetrical, strongly narrowed along lower half). Colliculum proper
short, hyaline body surrounding it limited, less circular.
Anjavidilava, 1 975-1 990 m, 19-26 .xii.1970 (P. Grweaud), 1 holotype, genit.
slide 8530 ; 1 Ç, allotype, genit. slide 8333 ; 4 Ç, paratypes.
Madagascar Centre, ouest du Vohidray, Ambalamarovandana, 1 530 m, 16-21.1.1971
(P. Grweaud ), 1 paratype, genit. slide 8534.
A deep vinaceous-brown species with well delined blackish central fascia. Judging
from the male genitalia, also close to G. (O.) benevolens Diak., from Périnet, but with diffe¬
rent labis and completely different markings.
Goniotorna (Oestophyes) vulpicolor Diak.
Diakonoff, 1960 : 144, pl. 26, fig. 172-173 ($).
Distribution. East Madagascar : Sambava district, Marojejy Range, 2 030 m.
Anjavidilava, 19-30.xii.1970, 1 975-1 990 m (P. Grweaud), 3 Ç, genit. slides 8500,
8501, 8502.
The female genitalia hâve a very characteristic long-erected ostium, removing the
species far from its congeners. Lamella postvaginalis in the présent sériés is less strongly
constricted at each side of the ostium than in the type, but otherwise the genitalia are
identical.
The species has a remarkable and puzzling similarity with the female of Metamesia
mianta sp. nov., perhaps owing to mimicry.
122
ALEXEY DIAKONOFF
Goniotorna (Goniotorna) mucida Diak.
(Fig. 15-17)
Goniotorna (Oestophyes) mucida diakonoff, 1960 : 142, pl. 27, fig. 178-179. — 1970 : 117.
Distribution. Central Madagascar : Ankaratra Range. North Madagascar : Tsa-
ratanana Range.
Anjavidilava, 1 975-2 005 m, 19.xn.1970-15.1.1971 (P. Griveaud), 1 apallotype,
genit. slide 8342 ; 14 2 $, genit. slides $ 8344, 8516, Ç 8343, 8526.
Having studied a long sériés of males of this species, I am satisfied that it belongs
to the subgenus Goniotorna Meyr. and not to Oestophyes Diak.
17-20 mm, apallotype. Head pale grey, palpus pale grey rather mixed with dark
grey, base of médian segment tawny. Thorax light purplish mixed with darker. Abdo¬
men pale ochreous, grey-whitish with a golden gloss, anal tuft somewhat deeper ochreous-
grey.
Fore wing ohlong-suhoval, moderately hroad, Costa arched thronghout appearmg
almost straight before apex, apex rectangular, slightly obtuse, termen vertical and gently
concave above, broadly rounded beneath. Pale tawny-fulvous, towards base suffused
with deeper fulvous, moderately and rather regularly strewn with slender black transverse
strigulae, oblique on anterior half of wing, on posterior becoming sparse sériés of black
points, parallel to posterior edge of central fascia and in some three broad curves below
apex and before termen. Basal patch indefinite, suffused with ferruginous-fulvous, little
darker ; central fascia well-defined, blackish and rather narrow along upper third, from
Costa before middle to 3/4 of dorsum, on lower 2/3 much dilated by a dark brown suffusion,
extending anteriorly along médian third of fascia, posteriorly broadly rounded and pro¬
minent along lower 2/3, but not reaching tornus ; costal patch indicated by some four
suffused black dots in a semioval, a fifth dot in middle and marginal ; this patch slightly
lîlled out with blackish. Cilia pale tawny.
Hind wing pale ochreous, except along Costa, suffused with pale fuscous-tawny, cilia
pale ochreous.
Male genitalia. Resembling those of G. (G.) chersopis Meyr. most, but with less roun¬
ded valva, more slender (althougli slightly variable) labis, and the aedeagus with a sub¬
apical spike at the left side of upper edge.
The species is considerably variable, paler or darker, with conspicuous or almost
entirely obliterate central fascia. However, the shape of fascia (when présent) and general
colouring remain constant. This, together with the short-spiked aedeagus, rather narrow
and simple sacculus (as depicted) and, of course, the normal male hind wing, allow of
an easy séparation from (apparently much more rare) G. (G.) chersopis.
TORTRICIDAE OF THE ANDRINGITRA RANGE
123
Goniotorna (Goniotorna) micrognatha Diak.
Diakonoff, 1960 : 147, pl. 27, fig. 176-177, pl. 28, fig. 186-187, pl. 30, ûg. 196. — 1963 : 136.
Distribution. East Madagascar : Périnet ; Ambatondrazaka. Central Madagascar :
Manjakandriana ; Betsileo Country, Ankazomivady. Southeast Madagascar : Andoha-
helo Range. North Madagascar : Montagne d’Ambre.
Anjavidilava, 1 990 m, 2 005 m (P. Griveaud ), 5 genit. slides 8345, 8349, 8350
and 8470.
Manjarivolo (P. Viette), 1 çj, genit. slide 8348.
Madagascar Centre, forêt Vakoana, Ambalamarovandana, 1 530 m, Andringitra-
Ambalavao, 21.1.1958 (P. Griveaud), 1 çj, genit. slide 8462.
Goniotorna (Goniotorna) megalogonia Diak.
Diakonoff, 1960 : 150, pl. 28, fig. 184-185, pl. 30, fig. 197-198.
Distribution. East Madagascar : Périnet ; Nosivola ; Ambatondrazaka ; 900-1 000 m ;
Marojejy Range, 2 030 m.
Anjavidilava, 1 975-2 005 m, 19 .xii.1970-15.1.1971 (P. Griveaud), 4
Goniotorna (Goniotorna) insatiata sp. nov.
(Fig. 20-24)
$ 17.5 mm. Head and face densely scaled, dark grey slightly mixed with paler pur-
plish. Palpus purple, fmely and closely dusted black. Thorax deep purple, apex fuscous-
whitish. Abdomen pale ochreous-fuscous, strewn with black. Posterior tibia pale grey,
densely scaled with long soft appressed scales, especially above.
Fore wing suboval, costa oblique but moderately arched to beyond middle and with
short, erected (and not rolled) costal edge (fold) from well beyond middle, straight before
apex, apex obtusely angular, termen short, straight and vertical above, rounded below.
Light ochreous, paler anteriorly but almost entirely obscured with transverse strigulation
and suffusion. Basal patch over 1/3, densely suffused with black, not reaching below vein
18, edge traceable, rather straight, with a triangular prominence in cell ; central fascia
dark grey, edges suffused black and tolerably parallel as far as cell ; in and below cell
fascia strongly projecting posteriorlv to before tornus, its edge almost horizontal above,
vertical and slightly concave posteriorly, below reaching tornus ; this fascia rather suffused
with ferruginous, veins faintly suffused with black, edges deep ferruginous ; anterior edge
of cell with a large angulation just above middle, moderately concave above and below
this ; space between basal patch and fascia on costal third suffused grey, below pale with
124
ALEXEY DIAKONOFF
three undulate black slightly inwards-oblique parting lines ; costal patch regularly semi-
oval, rather narrow, purplish, grey and ferruginous, with four black costal spots and ferru-
ginous marks along lower edge ; light ochreous ground colour so forming well defined cur-
ved streak from 2/3 of costa to apex and a broader vertical blotch along termen, the latter
including a submarginal dark purple patch of marbling and narrow lines along veins. Cilia
pale ochreous-grey, basal third light ochreous with grey bars.
Hind wing moderately broad, sub-semioval, termen oblique, little prominent in mid-
dle, dorsum with a triangular, not tightly rolled lobe, with posterior edge and inside pale
grey ; upper side of wing dull, rather deep fuscous-grey, strewn with small jet-black scales ;
a long pencil of fuscous-grey hairs from base not quite reaching tornus, paler at base. Cilia
glossy pale whitish-grey, with a pale basal and a slender grey subbasal line.
Male genitalia. Nearest to those of G. (G.) chersopis Meyr. (fig. 49-50) differingas follows.
Valva semioval, less spheroid ; sacculus sclerotized throughout, strongly angularly projecting
and apparently fiat, posterior edge of projection entirely straight (concave in ail other
species). Aedeagus with a large spike well before top of upper edge. Labis more slender.
Madagascar Centre, ouest du Vohidray, (Andringitra), Ambalamarovandana, 1 530 m,
16-21.xi.1971 (P. Griveàud), 1 çj, holotype, genit. slide 8529 ; 1 (J, paratype ; 1 $, allo¬
type, genit. slide 8536.
Ç 19 mm. Similar to the male, but with narrower fore wing. Basal patch with edge
indicated, angulate in middle, with a dark fuscous médian spot just above fold at 1/3 of
wing ; central fascia sufîused on lower half, posterior edge rather well defined, accentuated
by a very oblique and strongly convex continuons séries of dark points, a dark fuscous
spot on 3/4 of dorsum ; preterminal dark markings absent, except a conspicuous curved
sériés of dark dots from top of costal patch, paralled to edge of central fascia to tornus.
Hind wing bronze-fuscous, not black-dusted and therefore much lighter. Cilia pale
grey with a darker submedian band.
Female genitalia. Ostium bursae widely triangular, with sides gently convex, point
moderately rounded, conus of colliculum small, colliculum proper slender and tubular,
open frontally, gently dilated downwards, preceded by a rather wide hyaline dilatation
of ductus, colliculum with thick refracting walls. Corpus bursae rather small, signum
of normal shape.
A puzzling species, closely resembling G. (G.) megalogonia but smaller, and by doser
study differing also superficially by long-haired posterior tibiae (in megalogonia short-
haired, in chersopis normal), by black scales over hind wing and abdomen (absent in cherso¬
pis) and by different tornal lobe which is triangular and shorter, not oblong and projecting
posteriorly beyond tornus, as in megalogonia.
The subapical spike of the aedeagus is smaller in chersopis and actually is absent (or
not traceable) in the lectotype of that species (fig. 49-50), contrary to my original descrip¬
tion of the genitalia (1959) ; however, a small spike is présent in another syntype.
TORTRICIDAE OF THE ANDRINGITRA RANGE
125
Goniotorna (Goniotorna) erratica (Diak.)
Syndemis erratica Diakonofî, 1947 : 24 fig. 3-4. —- Viette, 1947 : 56.
Goniotorna (G.) erratica : Diakonoff, 1960 : 150, fig. 77, pi. 28, fig. 188-189. —- 1963 : 136. —
1970 : 117.
Distribution. Central Madagascar : Tsimbazaza park. East Madagascar : Mana-
kambahiny Est ; Marojejy Range. North Madagascar : Tsaratanana Range.
Anjavidilava, 1 975 m, 1 990 m (P. Grioeaud) , 4 <$.
Goniotorna (Tenuisaccula) rhodolemma Diak.
Diakonoff, 1960 : 154, pi. 29, fig. 192, pl. 30, fig. 199-200.
Distribution. Central Madagascar : Ankasoka ; Andringitra Range, Anjavidilava,
2 030 m.
Anjavidilava, 1 975, 1 990 and 2 005 m (P. Grioeaud ), 3 1 Ç.
Manjarivolo (P. Viette), 1 Ç.
Goniotorna (Tenuisaccula) polyops Diak.
Diakonoff, 1960 : 159, pl. 27, fig. 180-181, pl. 31, fig. 203.
Distribution. East Madagascar : Sandrangato forest. Central Madagascar : Andrin
gitra Range, Anjavidilava, 2 030 m, 19.1.1958 ; Ampitameloka, Moramanga ; Sambava
district, Marojejy Range.
Andringitra centre, plateau d’Andohariana (Soaindrana) 2 030 m, 10-21.xi. 1970
(P. Viette), 1 $, genit. slide 8291.
Goniotorna (Tenuisaccula) mianta sp. nov.
(Fig. 33-36)
16 mm, Ç 16.5-18.5 mm. Head light tawny, face whitish. Palpus light tawny.
Thorax light tawny, glossy. Abdomen pale fuscous.
Fore wing moderately broad, Costa curved throughout, apex pointed and slightly
produced, termen sinuate, moderately oblique. Glossy pale ochreous-tawny, markings
more or less suffused, brown to dark brown. Rasai patch to 1/4, only costal half as far
as fold developed, blackish-brown, edge well-defined, oblique and gently concave, lower
edge moderately suffused ; central fascia from well before middle, rather narrow, hardly
dilated, oblique and gently excurved (concave posteriorly), limited below by lower edge
33
Fig. 33-38. — Genitalia of Madagascan Tortricinae. 33, Goniotorna (Tenuisaccula) mianta sp. nov., <J,
holotype ; 34, the same, aedeagus ; 33, the same, $, allotype ; 36, the same, bursa ; 37, Acleris phane-
rocrypta sp. nov., holotype ; 38, the same, aedeagus, with juxta.
TORTRICIDAE OF THE ANDRINGITRA RANGE
127
of fold ; dark brown, edges faintly suffused with blackish-brown ; costal patch narrow and
oblong, gradually narrowed posteriorly, ends rounded : from before 2/3 to well before angle,
light brown ; a darker brown, well-defined preterminal fascia, from angle to dorsum before
tornus, gently extended along dorsum (thus indicating base of central fascia), posterior
cdge distinctly concave and fînely serrulate, anterior edge triangularly prominent above
middle, elsewhere somewhat irregular ; a dark brown dot on lower angle of cell and a paler
brown suffused line, parting space between central fascia and costal patch, strongly sinuate
and merging in suffusion around foot of preterminal fascia. Cilia whitish or pale tawny,
a brownish suffused bar apposite apex.
Hind wing pale fuscous-greyish, tinged tawny towards apex, fînely strewn witb sparse
black scales over basal third. Cilia glossy pale ochreous.
Male genitalia. Resembling those of G. (T.) leucophrys Diak., but very distinct.
Uncus more elongate, spoon-shaped ; transtilla witb labidcs very robust. Yalva semi-
circular, bigher than in other species, sacculus more rounded along lower edge, sclerotized
throughout, its upper edge forming a smooth sclerite, oblong-crescentic. Aedeagus scle¬
rotized, with a longitudinal split, subapical thorn small, straight, not upcurved.
Andringitra oriental, forêt d’Anjavidilava, 1 975 m, 26-30.xn. 1970 (P. Griveaud), 1
cJ, holotype, genit. slide 8467. Superficially resembling a Metamesia species.
Goniotorna (Tenuisaccula) chondrocentra sp. nov.
(Fig. 41-42)
18 mm. Head and palpus light tawny. Thorax whitish-grey, glossy, anterior
fourth light tawny. Abdomen glossy fuscous-grey.
Fore wing oblong-suboval, Costa curved throughout, apex tolerably pointed, termen
very gently sinuate, hardly oblique above, rounded below. Glossy rather light grey-
fuscous. Basal patch hardly to 1/4, slightly darker infuscated, edge marked with blackish
dots, with an acute prominence just above middle, otherwise almost straight and vertical,
preceded bv a submarginal black parallel and straight line ; central fascia indicated bv an
oblique dark grey band from before middle of Costa to upper edge of cell, edges slightly
marked with purplish small dots ; lower half of fascia hardly traceable, purplish ; three
purple black dots among middle of edge of basal patch, a subohlong larger spot beyond
middle of cell and a curved row of purple-black points from 3/5 of Costa to tornus ; another
sériés of smaller dots below apex and before termen ; three larger dots on central fascia
below Costa ; a row of suffused grey blotches along dorsum. Cilia light purple-fuscous
with a white basal line.
Hind wing glossy pale grey, apex slightly darker and with finest grey dotting. Cilia
concolorous.
Male genitalia. Resembling those of G. (T.) rhodoptila Diak., but with uncus elongate
and pointed, tegumen broader at base, gnathos very densely black-scaled (or spined).
128
ALEXEY DIAKONOFF
Madagascar Centre, Andringitra méridional, chaîne de l’Andrianony, cirque de Man-
jarivolo, 1 650 m, 26 x.-4.xi.1970 (P. Viette), 1 <§, holotype, genit. slide 8288; 1 çj, para-
type, genit. slide. 8331.
Distinct by greyish general appearance and dark discal dots ; the paratype is much
paler, but the dots are similar, only more contrasting.
Fig. 39-42. — Male genitalia of Madagascan Tortricinae. 39, Goniotorna (' Tenuisaccula) trignoma sp.
ilov., holotype ; 40, the saine, aedeagus ; 41, G. (T.) chondrocentra sp. nov., holotype ; 42, the same,
aedeagus.
Goniotorna (Tenuisaccula) trignoma sp. nov.
(Fig. 39-40)
$ 16 mm. Head white, scape of antenna ferruginous. Antenna ferruginous. Pal-
pus fulvous, at base becoming whitish. Thorax ferruginous mixed with whitish, tip white
(imperfect). Abdomen pale ochreous-fuscous, anal tuft fuscous, very large.
Fore wing oblong-truncate, costa arehed at base, straight posteriorly, apex subobtuse,
termen almost straight, hardly sinuate, little oblique. Very pale ochreous, with a strong
silky gloss, sparsely sprinkled with fulvous, markings brown. A dark brown oval spot along
basal 1/5 of costa, hardly reaching below base of fold ; central fascia formed by a rather
slender well-defîned oblique streak front middle of costa to lower edge of cell, light brown,
edges dotted and lower third suffused with dark brown ; anterior edge straight, posterior
moderately concave, top truncate by cell edge : lower half of fascia extremely faint, appa-
Fig. 43-46. — Male genitalia of Madagascan Tortricinae. 43, Goniotorna ( Oestophyes) trigodes sp. nov.
paratype, genit. slide 8290 ; 44, the same, aedeagus ; 45, the same, holotype ; 46, the same, aedea
gus.
Fig. 47-50. — Male genitalia of Madagascan Tortricinae. 47, Goniotorna ( Oestophyes) verticillata Diak.
genit. slide 5286 ; 48, the same, aedeagus ; 49, G. (G.) chersopis Meyr., lectotype ; 50, the same, aedea
130
ALEXEY DIAKONOFF
rently dilated and less oblique, costal patch oblong-semioval, rather narrow, edges and
two oblique transverse lines darker brown ; some irregular transverse blackish dotting
before termen, in right wing forming two parallel lines, below apex and above tornus, respecti-
vely ; terminal edge with minute blackish interneural marginal marks. Cilia concolorous.
Hind wing evenly pale ochreous-fuscous, less glossy. Cilia paler.
Male genitalia. Nearest to those of G. (T.) rhodolemma Diak., but very distinct by
an acute spike at the base of sacculus ; besides, by a more narrow, spindle-shaped uncus
and a longer, entirely spiny labis ; valva with regard to the shape, nearest to that in G.
(T.) rhodoptila Diak., but shorter, with a rounded bulge of the Costa, but differing at once
by the spike of the sacculus.
Females lacking.
Madagascar Centre, Andringitra oriental, forêt d’Anjavidilava, 1 975 m, 26-30.xn. 1970
(P. Griveaud), 1 <$, holotype, genit. slide 8523. A slender species with quite distinct
and characteristic markings.
Goniotorna (Serruligera) melanoconis Diak.
Diakonoff, 1960 : 164, pl. 33, fig. 214-216. — 1970 : 119.
Distribution. Central Madagascar : Betsileo Country, Ambatofitorahana forest.
East Madagascar : Analamazoatra forest ; Manakambahiny Est. North Madagascar :
Montagne d’Ambre.
Anjavidilava, 1 990 m, 2 005 m (P. Griveaud), 11
Manjarivolo (P. Viette), 1
Madagascar Centre, ouest du Vohidray (Andringitra), Ambalamarovandana, 1 530 m,
16-21 .i. 1971 (P. Griveaud), 1 $.
Epichoristodes (Epichoristodes) canonicum sp. nov.
(Fig. 51-52)
^ 21-24 mm. Head light ochreous-fulvous, face paler. Palpus light fulvous, pale
ochreous inside. Thorax light fulvous, pale ochreous on posterior half, tegulae suffused
with tawny. Abdomen pale ochreous.
Fore wing oblong-truncate, broad, little dilated, Costa strongly arched anteriorly,
straight posteriorly, apex moderately pointed, slightly prominent, termen sinuate above,
rounded-prominent below, vertical. Glossy pale ochreous, strewn with fine transverse
ferruginous strigulae, tending to form a réticulation before termen. Markings deep ferru-
ginous, with suffused dark brown edges, well-defined. Basal patch extending on Costa slightly
less than 1/5, on dorsum to before middle, light ochreous, with dark obliterate réticulation,
below fold deep ferruginous-brown with dark brown réticulation ; extreme base suffused
TORTRICIDAE OF THE ANDRINGITRA RANGE
131
dark brown, Costa with three triangular dots, posterior largest and marginal ; edge of patch
from this spot, concave and strongly oblique, with a subobtuse projection in middle, filled
ont with dark ferruginous-brown, edge below fold deeply emarginate ; central fascia rather
narrow, constricted above middle, from 1/3 of Costa to 2/3 of dorsum, upper half wedge-
shaped ; anterior edge of fascia well-defined, three t.imes scalloped, posterior edge along
lower 2/3 suffused and notched twice, extending along posterior 1/3 of dorsum to tornus
and becoming paler and greyish there ; costal patch at 3/5, moderate, subquadrate, gently
oblique, lower angles rounded ; followed by two marginal round dots, anterior with its
lower edge connected by a small bar with patch ; an orange small suffusion on dorsal lobe ;
a similar suffusion, paler and narrower, along posterior third of cell above its middle ; a
suffused ferruginous apical spot, extending over cilia. Cilia ferruginous in apex and along
lower 1/3 of termen, ochreous elsewhere, with a faint dark brown basal band, dissolved
into dark bars opposite ends of veins, towards and in tornus, bar on end of dorsum large
and distinct.
Hind wing pale golden-ochreous with a silky gloss, a trifle deeper ochreous posteriorly.
Cilia concolorous, around apex and along upper third of termen suffused vinaceous.
Male genitalia. Very similar to those of E. ( E .) leucocymba Diak., but with a narrower
valva, a less sclerotized sacculus which is not dilated in middle and a less prominent pul-
vinus.
Female unknown.
Andringitra oriental, forêt d’Anjavidilava, 1 990 m, 19-25.xii.1970 (P. Griveaud),
1 (J, holotype, genit. slide 8268 ; the same, 1 975 m, 26-30 .xii.1970 (P. Griveaud) ; the
saine, 2 000 m, 2-5.xn.1970 (P. Soga) and 1-15.1.1971 (P. Griveaud).
Andringitra méridional, chaîne de l’Andrianony, cirque de Manjarivolo, 1 650 m,
26.x-4.xi.1970 (P. Viette).
Holotype and 7 <$ paratypes altogether.
A very distinct species with the usual Archipine markings. The genitalia, however,
resemble those of the quite differently marked E. leucocymba closely.
Belongs to the group of E. ( E .) goniopa Diak. and macrosema Diak., difîering from
the former by the sacculus rounded and not prominent heyond base, and by the top of
valva more obtusely rounded and the aedeagus similar, but distinctly shorter ; and sepa-
rated from macrosema at once by the dilated aedeagus of the latter species.
Epichoristodes (Epichoristodes) atricaput sp. nov.
(Fig. 55-56)
Ç 17.5 mm. Head, palpus and collar deep ferruginous. Thorax light tawny-fulvous,
glossy, anterior edge, shoulder and basal half of tegula deep ferruginous. Abdomen pale
golden-ochreous, venter tawny-ochreous with a crimson iridescense.
TORTRICIDAE OF THE ANDRINGITRA RANGE
133
Fore wing oblong-truncate, Costa arched at base, straight posteriorly, moderately
but distinctly promineut at 3/4, apex obtuse-rectangular, termen gently concave, vertical
above, rounded beneath. Pale fulvous with a strong whitish gloss, densely strigulated
with transverse, broadly undulate, partly interrupted, orangeish-fulvous, very dull lines
of diverse width. Markings of the same colour, very dull. Anterior fourth of costal
edge pale orange, a small basal spot deep ferruginous ; central fascia indicated by a moderate
subtriangular patch, continued to dorsum well beyond middle by a single stria, gradually
concave and narrowly edged with glossy ground colour anteriorly, irregular posteriorly ;
costal patch large, from beyond middle of Costa to well before apex, semioval, below con-
nected with transverse striae and including a couple of small spots of glossy ground colour ;
Costa posteriorly orangeish-fulvous, dull ; transverse strigulation reduced along and below
basal fourth of Costa, less distinctly around costal patch and on terminal fourth of wing ;
the latter fourth with some three almost vertical thicker straight transverse striae, poste-
rior marginal, these striae irregular in right wing. Cilia dull orangeish-fulvous, with deep
ferruginous apical third.
Hind wing fulvous-orange, deeper coloured and tinged pinkish posteriorly, along Costa
pale ochreous. Cilia concolorous.
Female genitalia. Sterigma little sclerotized, transversely oblong, lamella postva-
ginalis in middle, narrow at the sides ; lamella antevaginalis little extended, irregularly
sub-linear. Colliculum membraneous, strongly wrinkled. Ductus bursae very long and
narrow, straight. Corpus bursae almost spherical. Signum rhomboidal, with serrulate
edge.
Madagascar Centre, Andringitra méridional, chaîne de l’Andrianony, cirque de Man-
jarivolo, 1 650 m, 26.x-4.xi.1970 (P. Viette), 1 Ç, holotype, genit. slide 8284.
A conspicuous insect with distinct but rather not instructive genitalia. The actual
position of the présent species is somewhat puzzling without the evidence of male génital
characters.
Epichoristodes (Epichoristodes) apileticum Diak.
Diakonoff, 1960 : 170, pl. 33, fig. 217-219, pl. 35, fïg. 227-228. — 1961 : 155. — 1970 : 120.
Distribution. Central Madagascar : Betsileo country ; Ankaratra Range. North
Madagascar : Tsaratanana Range. East Madagascar : Ampitameloka.
Manjarivolo (P. Viette), 1
Anjavidilava, 1 975 m, 1 990 m (P. Griveaud), 3 <$, 1 Ç.
Epichoristodes (Epichoristodes) ypsilon Diak.
Diakonoff, 1960 : 173, pl. 34, fig. 220-221.
Distribution. Manjakandriana.
Anjavidilava, 2 005 m, 1-15.1.1971 (P. Griveaud), 1 rf.
134
ALEXEY DIAKONOFF
Epichoristodes (Tubula) incertum Diak.
Diakonoff, 1960 : 171, pl. 34, fig. 224, 224a {incerta).
Epichoristodes ( Tubula) nervosum Diakonoff, 1970 : 120, pl. 10, fig. 1-2 (Ç). Syn. nov.
Distribution. Northeast Madagascar : Marojejy Range. Central Madagascar :
Ankaratra Range, 1 850-2 030 m ; Andringitra Range, Anjavidilava. North Madagascar :
Tsaratanana Range, 1 900-2 050 m.
Andringitra central, plateau d’Andohariana (Soaindrana), 2 030 m, 10-21.xi.1970
(P. Viette), 1 $.
Anjavidilava, 1 975-2 005 m, 19.xn.1970-15.1.1971 {P. Grü’eaud), 11 11 Ç, genit.
slides cJ 8330, $ 8269, 8285.
Manjarivolo {P. Viette), 3 5 Ç, genit. slide 8468
Having been able to study a nice sériés of males and females I am satisfied that “ ner¬
vosum ”, described after a single specimen, is but the female sex of the présent species,
which should be referrcd to subgenus Tubula , as is now proven by the female genitalia.
Gephyraspis insolita sp. nov.
(Fig. 53-54)
$ 21 mm. Ilead pale yellow. Palpus light tawny, base pale yellow, edge of this
colour oblique. Thorax pale yellow, apex fuscous-tawny. Abdomen and anal tuft white,
upper half sufîused with blackish.
Fore wing oval, Costa strongly curved anteriorly, and with a short upright. and open
fold obtusely rectangular, termen straight, vertical above, rounded beneath. Light tawny,
evenly and almost entirely infuscated ; very faint fuscous-grey darker shadowy streaks
obliquely across wing, from helow Costa at 1/4 and before middle, respectively (the second
streak apparently indicating central fascia) ; Costa with a rather well-defined pale yellow
streak from end of costal fold to apex, including 3-4 distant black points ; tornal area as
far as cell sufîused orange-tawnv and on posterior half darker infuscated. Cilia dark
fuscous.
Hind wing rather dark purple-fuscous with an indefinite reddish-vinaceous tinge,
edge clouded with dark fuscous ; dorsum strewn with dense dark fuscous scales. Cilia
fuscous-grey with a dark fuscous-grey subbasal shade (inperfect).
Male genitalia. Very similar to those of G. lutescens Diak. but difïering by the shape
of the médian process of the transtilla (broken at base, but présent in mount) ; with an
inverted-Y-shaped (instead of inverted-T-shaped) base and a fuscate top. Valva decidedly
broader, aedeagus obliquely hooked at top.
Females lacking.
Anjavidilava, 2 005 m, 1-15.1.1971 {P. Griveaud), 1 cJ, holotype, genit. slide 8281.
TORTRICIDAE OF THE ANDRINGITRA RANGE
135
Rather similar to the type-species, but with an elementary costal fold, generally paler
and with dorsum not darker suffused. Hind wing darker brownish. The transtilla and
the aedeagus are so different that there seems to be no doubt of the species being distinct.
Gephyraspis contranota sp. nov.
(Fig. 66-68)
Ç 21 mm. Head creamy-white. Palpus long, light ochreous-golden, tip of médian
and entire terminal segment brownish. Collar creamy-white. Thorax pale tawny. Abdo¬
men fuscous-ochreous.
Fore wing oblong, moderately broad, Costa curved anteriorly, sinuate posteriorly,
apex moderately pointed, slightly produced, termen sinuate, little oblique. Pale ochreous-
tawny, strewn ail over with short jet-black strigulae and dots. Basal patch to 1/6, suffu¬
sed light tawny-greyish, edge narrowly blackish, patch reaching halfway across wing ;
this patch followed on Costa by three narrow, interrupted lines ; central fascia broken in
two, costal part narrow, oblique, blackish, turning grey downwards, interrupted by cell,
from beyond 1/3 of costa ; lower half of fascia forming an obliquely oval large patch on end
of dorsum, chestnut-brown, mixed with black on lower half ; sotne light brown suffusion
extending along dorsum below fold, not reaching base ; costal patch elongate-triangular
rather narrowly depressed, équilatéral, brownish, closely strewn with black. Cilia (imper-
fect) brown.
Hind wing and cilia pale ochreous with a golden gloss, apex dotted with dark grey.
Female genitalia. Lobi anales finely granulate and shortly haired. Ninth stérilité
sclerotized and compressed laterally, dorso-ventral diameter being larges!. Lamella
postvaginalis, a brace-shaped, strong edge, in middle slightly rising, latéral angles long
and pending ; lamella antevaginalis, a simple, slender edge, angulate in middle. Colli-
culum wide, funnel-shaped, asymmetrical, constricted above end. Ductus bursae long
and slender. Corpus bursae small, spherical. Signum, a single granulate cusp.
Madagascar Centre, Andringitra oriental, forêt d’Anjavidilava, 2 005 m, 1-15.1.1971
( P. Griveaud), 1 Ç, holotype, genit. slide 8339 ; 1990 m, 19-25 .xii. 1970 (P. Griveaud), 1 <$,
paratype, slide 8581.
A distinctly marked species, but the position is uncertain without males. The para-
type is very pale, with ail markings présent but faded to a pale grey.
DORIDOSTOMA gen. nov.
Head with dense, closely appressed scales. Ocellus small, posterior. Proboscis
rather short. Antenna in male ciliated, cilia just over 1. Thorax without a crest. Legs
and abdomen normal.
Fore wing oblong-subtruncate, moderately broad. Vein 2 from middle of cell, 3
136
ALEXEY DIAKONOFF
from angle, 4 doser to 5, 7 separate to termen, 9 from 2/3 distance 10-8 ; 10 from 1/2 dis¬
tance 11-9, 11 from slightly beyond middle of cell.
Hind wing without a cubital pecten ; slightly over 1, broadly semioval, vein 2 from
3/5 or 2/3 of cell, 3 and 4 connate or almost connate, from angle, 5 closely approximated
at base, 6 and 7 shortly or moderately stalked.
Male genitalia. Tegumen broad and short. Uncus very broad, almost quadrate,
xruncate, angles rounded ; with two patches of rather thin quadrifid scales. Socius very
small, free. Gnathos moderate, inverted-trapezoidal, with acutely produced angles and
a slender hook. Transtilla dentate above, almost paired. Pulvinus absent. Valva small,
oblong-semioval ; sacculus indefinite. Aedeagus small, slender, moderately curved at
the extremities ; cornutus, one slender spine.
Type-species : D. denotata sp. nov.
Apparently nearest to Pandemis Hb., but with a complicated gnathos, thorny trans¬
tilla and small valva. Tliese characters and also indefinite sacculus separate this form
also from Parapandemis. The species has the appearance, rather, of an Epichoristodes,
but the genitalia are widely different.
There are no females available.
Doridostoma denotata sp. nov.
(Fig. 57-58, 61-63)
$ 18-19 mm. Head purplish-grey, on forehead suffused with fulvous. Palpus light
tawny-vinaceous, laterally with a darker médian suffused streak from base of médian seg¬
ment above to its apex below, terminal segment grey. Thorax vinaceous-brown, or pur-
plish. Abdomen fuscous, venter paler.
Fore wing oblong-suboval, Costa arched anteriorly, little curved posteriorly, apex
obtusely pointed, termen gently sinuate above, rounded and prominent below. Fulvous,
finely dusted purplish, posteriorly tending to form transverse strigulae. Markings deep
cofîee-brown, dull, well defined. Basal patch irregular, almost 1/3, strongly invaded by
ground colour, upper half indicated only by oblique and concave edge, lower half dark pur-
plish ; edge with tooth-like up-turned prominence at 1/3, thence inwards-oblique, to before
1/3 of dorsum ; base of Costa with a blackish streak ; central fascia from slightly beyond
middle, moderately broad, little dilated, to posterior 1/4 of dorsum ; its anterior edge
straight, extremities slightly extended anterad along wing edges ; posterior edge concave
along upper 1/3, strongly excavated above middle, excavation limited posteriorly by a
rising dark tooth, edge below this regularly convex and fascia moderately thickened, with
a black spot on dorsum ; costal patch large, along posterior third to apex, before apex inclu-
ding two small costal spots of ground colour, edged with black ; this patch with anterior
edge twice convex, appearing notched in middle, acute top of patch to middle of wing,
posterior edge less oblique, somewhat suffused, in left wing followed by a dark subterminal
TORTRICIDAE OF THE ANDRINGITRA RANGE
137
line, little outwards-oblique ; terminal part paler with darker, blackish dotting cilia (imper-
fect) fulvous, ferrnginous towards apex, with black marks towards dorsum.
Hind wing grey-fuscous with a golden gloss, lînely strewn with darker (tips of scales).
Cilia pale golden-fuscous with a blackish subbasal band.
Male genitalia. As described with the genus.
Female unknown.
Andringitra oriental, forêt d’Anjavidilava, 1990 m, 19-25 .xii.1970 (P. Griveaud ),
1 <$, holotype, genit. slide 8434 ; the same, 2 005 m, 1-15.1.1971 (P. Griveaud), 1 para-
type.
Fig. 57-60. — Sketches of heads and wing neuration of Doridostoma gen. nov., holotypes. 57, D. denotata
sp. nov., head ; 58, the same, wing neuration ; 59, D. stenomorpha sp. nov., head ; 60, the same, wing
neuration.
138
ALEXEY DIAKONOFF
Fig. 61-68. — Genitalia of Madagascar! Tortricinae. 61, Doridostoma denotata sp. nov., <J, holotype ;
62, the same, tip of a bristle of the uncus, stronger magnified ; 63, the same, aedeagus ; 64, D. steno m
morpha sp. nov., holotype ; 65, the same, aedeagus ; 66, Gephyraspis contranota sp. nov., $, holo¬
type ; 67, part of ductus bursae ; 68, bursa.
TORTRICIDAE OF THE ANDRINGITRA RANGE
139
Doridostoma stenomorpha sp. nov.
(Fig. 59-60, 64-65)
3 16 mm. Head, palpus and thorax whitish, finely dusted throughout with dark
fuscous, so as to appear pale grey. Abdomen whitish, dorsum infuscated, except the anal
tuft.
Fore wing oblong-suboval, rather narrow, Costa curved throughout, more curved at
extremities, apex pointed, termen sinuate, very oblique, long. Creamy-whitish, more
or less touched with fuscous, very finely but irregularly dusted with minute black points
and strewn with small groups of black hair-scales. Cilia long, concolorous, with a blackish
suffused subbasal band and a narrower and faint subapical line, except around tornus.
Hind wing glossy creamy-whitish, faintly tinged deeper creamy on apical half. Cilia
whitish, glossy.
Male genitalia. Tegumen robust, higher than broad. Uncus very broad, almost
quadrate, at the sides rounded-prominent, below with two brushes of bristles. Socius
very small, free. Gnathos hooked, rather strong, arms rather broad and curved, hook
oblong-oval. Yalva small, over 2 X as long as broad, suboval. Transtilla, a narrow
rod ; labis very large, semioval, rising, sparsely long-dentate. Aedeagus pistol-shaped,
upper part straight, with a short, narrowed caecum, top of aedeagus clavate.
Andringitra oriental, forêt d’Anjavidilava, 1 975 m, 26-30 .xii.1970 (P. Griveaud),
1 (J, holotype, genit. slide 8498. A puzzling species with the appearance of a Balioxena
iospila Meyr. (except for the palpi), with the male genitalia intermediate between the
preceding species and Peteliacma torrescens Meyr., but without anything of the peculia-
rities of the neuration of these two généra.
Balioxena iospila Meyr.
(Fig. 69)
Meyrick, 1912 : 13. — Viette, 1948 : 57. — Clarke, 1955 : 175. — 1958 : 59, pl. 29 fig. 1-ld. —
Diakonoff, 1960 : 179, fig. 80, pl. 36, fig. 236.
Distribution. Central Madagascar : Andringitra Range ; Antanarivo.
Manjarivolo (P. Viette), 2 <$.
Anjavidilava, 1 990 m, 19-25. xii. 1970 (P. Griveaud), 2
Andringitra central, plateau d’Andohariana (Soaindrana) 2 030 m, 10-21.xi.1970
(P. Viette), 2 $, genit. slide 8249 ; 1 $, apallotype, genit. slide 8251.
$ 22 mm, apallotype. Similar to male, but head pale ochreous, abdomen dark fuscous,
anal tuft slaty grey, glossy.
Fore wing more pointed. Pale ochreous, from below Costa to dorsum evenly suffused
with brighter ochreous, becoming brighter downwards, almost orange on dorsum ; cell
140
ALEXEY DIAKONOFF
throughout except on edges rather mixed with leaden-black scales, less numerous towards
base ; very sparse black scales strewn on both sides of fold, and a few from cell to apex.
Hind wing brighter pale ochreous, with stronger golden gloss.
Female genitalia. Sterigma little sclerotized, with oval latéral finely punctulate
tumescences. Ostium small, tubular and narrow, followed by a hyaline colliculum, below
dilated in a slightly sclerotized dilatation ; cestum, a straight, gradually dilated band.
Signum one, a simple hollow spike, with a small basal plate.
Fig. 69-70. — Female genitalia of Madagascan Tortricinae. 69, Balioxena iospila Meyr., $, apallotype ;
70, Acleris malagassana sp. nov., Ç, holotype.
Vialonga polyantha Diak.
Diakonoff, 1960 : 186, pl. 36, fig. 236-237.
Distribution. East Madagascar : Moramanga district.
TORTRICIDAE OF THE ANDRINGITRA RANGE
141
Anjavidilava, 1 990 m, 2 005 m (P. Griveaud), 11 Ç, gen. slide 8341.
Manjarivolo (P. Viette), 1 $.
The male is still unknown.
Bactrostoma cinis Diak.
Diakonoff, 1960 : 195, fîg. 86, pl. 37, fig. 241-242, pl. 38, fig. 249-250.
Distribution. Central Madagascar : Ankaratra Range.
Manjarivolo (P. Viette), 2 <$.
Furcinula punctulata Diak.
Diakonoff, 1960 : 199, text. fig. 88, pl. 38, fig. 254-255, pl. 39, fig. 256-258.
Distribution. East Madagascar : Marojejy Range, 1 600 m.
Anjavidilava, 1 975 m, 26-30 .xii. 1970 (P. Griveaud ), 1 <$, genit. slide 8522.
Acleris malagassana sp. nov.
(Fig. 70)
18 mm. Head, palpus rather light tawny-fuscous. Thorax tawny-fuscous. Abdo¬
men light golden-ochreous, venter tawny-ochreous with a crimson iridescence.
Fore wing elongate, moderately broad, Costa arched at base, gently curved posteriorly,
apex pointed, slightly prominent, termen sinuate, oblique. Deep tawny-fuscous with a
bronze gloss, gently sufïused throughout, strongly suffused over middle third with blackish,
a large, obliquely oval well-defined patch, occupying basal third of costa and basal seventh
of dorsum, edge over dise gradually convex, rather more rounded and prominent below
costa ; this patch pale ochieous, except along edge, rather densely sufïused with tawny
and strewn with slightly raised dark blackish-fuscous scales ; blackish suffusion posteriorly thin
and tending to be arranged along veins ; a minute blackish point halfway between apex
and cell, followed by more irregularly scattered minute dark points. Cilia pale ochreous
with blackish apical half (imperfect).
Hind wing pale ochreous, densely mottled throughout with pale grey, with a strong
silky gloss. Cilia sordid pale ochreous.
Female genitalia. Resembling those of the Palaearctic A. variegana Den. & Schiff.,
but quite distinct. Sterigma (evenly sclerotized) with latéral appendages vertical, slen-
der and gradually narrowed, ventral rim of ostium but slightly darker, colliculum almost
twice as long as appendages of sterigma, cylindrical, sclerotized, rather wide, gently curved
at lower end. Signum asteroid.
Madagascar Centre, Andringitra oriental, forêt d’Anjavidilava, 1 990 m, 19-25. xii. 1970
(P. Griveaud), 1 Ç, holotype, genit. slide 8284.
142
ALEXEY DIAKONOFF
A moderate rather narrow-winged species. The genitalia show a relation with the
A. thylacites Meyr. group of African species.
Acleris phanerocrypta sp. nov.
(Fig. 37-38)
about 16 mm. Head pale ochreous. Thorax pale ochreous, rather infuscated
(imperfect). Palpus rather long and slender, médian segment expanded and brushy towards
apex, terminal segment slender, exposed, moderate ; pale ochreous. Abdomen fuscous.
Fore wing moderately broad (tip of wings lacking), Costa moderately curved at base,
straight posteriorly. Pale ochreous, rather densely dusted, dotted and marked with dark
fuscous. Basal patch not reaching 1/4, regularly angulate in middle, edge straight above
middle, below strongly oblique, on dorsal edge vertical, narrow but distinct ; central fascia
indicated hy a moderate, outwards-concave streak from before middle of Costa, anterior
edge triangularly prominent ; costal patch ill-defined, indicated by some four costal stri-
gulae ail about 3/4, semioval ; an interrupted dark line from below costal patch, running
very oblique to halfway upper edge of cell and termen, thence little curved to termen
above tornus ; a similar dark line halfway between this and termen (damaged) ; a slender
marginal fuscous line along termen and in tornus ; Costa strigulated fuscous between mar-
kings ; a dark fuscous dot on 2/3 of fold. Cilia pale ochreous (imperfect).
Hind wing whitish-ochreous, rather dull, very faintly dotted fuscous (dotting on under-
side). Cilia concolorous.
Male genitalia. Tegumen rather slender and high. Gnathos darkly sclerotized,
subquadrate, the basis of the arms being connected by a dark transverse band-like subsca-
phium ; arms of gnathos strongly convex laterally, lower angles acutely produced, trans¬
verse part concave. Socius very long and narrow, with a lower, rigid part and a membra-
neous hairy part extended over it. Valva moderate, oblong-oval ; sacculus with a trian¬
gularly dilated base and a subapical long pending naked process ; cucullus hairy, rounded,
with a ventral triangular appendix (digitus). Juxta very broad, triangular. Aedeagus
slender, bent almost rectangularly, with two spikes (not deciduous cornuti) and a short
tooth below apex.
Madagascar Centre, Andringitra oriental, rivière est, Antsifotra, formation à Agau-
ria, 2 000 m, 2-5.xn.1970 (P. Soga), 1 <$, holotype, genit. slide 8533.
Unfortunately the single specimen is rather damaged, but the male genitalia are very
distinct. The species belongs to the A. thylacitis Meyr. group. Obviously it is not the sex
partner of A. tnalagassana sp. n.
Acknowledgements
The author is grateful for the opportunity to study this interesting material and for
the permission to retain certain duplicates for the collection of the Leiden Muséum.
TORTRICIDAE OF THE ANDRINGITRA RANGE
143
REFERENCES
Clarke, J. F. Gates, 1955. — Catalogue of the type spécimens of Microlepidoptera in the British
Muséum (Natural History) described by Edward Meyrick. 1 : i-vii, 1-332, 4 pl.
— 1958. — Loc. cit. 3 : 1-600, of which 1-298 plates.
Diakonoff, A., 1948. — Microlepidoptera from Madagascar. Mém. Inst, scient. Madagascar,
sér. A, 1 : 22-30, fîg. 1-8.
— 1959. — Mabille’s types of Malagassy Tortricidae. Rev. franç. Ent., 26 : 167-186.
— 1958. — Notes on Saalmüller’s types of Malagassy Microlepidoptera. Senckenberg. biol.,
39 : 89-90, pl. 4-5.
—- 1960. — Tortricidae from Madagascar. Part I. Verh. K. ned. Akad. Wet., Natuurk., ser. 2,
53 ( 2 ) : 1-209, 90 fig., 40 pl.
— 1961. -— Tortricidae from Madagascar in the Berlin Muséum. Deutsche ent. Ztschr., n. 5,
8 : 152-155.
— 1963. — Tortricidae (Lepid.) collected in Madagascar by Dr. Fred Keiser. Verh. Naturforsch.
Ges. Basel, 74 : 133-144, fig. 1-5, pl. l-III.
—- 1970. — Lepidoptera Tortricoidea from Tsaratanana Range (North Madagascar). Mém.
ORSTOM, 37 : 103-150, fig. 1-39.
Meyrick, E., 1912. — Exotic Microlepidoptera. Vol. 1 (1912-1916) : 1-640.
— 1930. — Exotic Microlepidoptera. Vol. 3 (1923-1930) : 1-640.
Paulian, R., J. M. Betsch, J. L. Guillaumet, Ch. Blanc, P. Griveaud, A. Descarpentries,
P. Viette, R. Albignae & M. Petit, 1971. — RCP 225. Étude des écosystèmes monta¬
gnards dans la région malgache. 1. — Le massif de l’Andringitra. 1970-1971. Géomorpho¬
logie, climatologie et groupements végétaux. Bull. Soc. Ecologie, 2 : 189-266, fig. 1-44.
Viette, P., 1948. — Catalogue des Microlépidoptères de Madagascar et des archipels environnants.
Mém. Inst, scient. Madagascar, sér. A, 1 : 31-75.
— 1972. — Contribution à l’étude des Satyrides du massif de l’Andringitra, Madagascar
Centre (Lep. Nymphalidae Satyrinae). Bull. Soc. ent. Fr., 76 1971 (1972) : 146-154, 6 fig.
Manuscrit déposé le 17 mars 1972.
Bull. Mus. Hist. nal., Paris, 3 e sér., n° 108, janv.-févr. 1973,
Zoologie 82 : 105-143.
Achevé d’imprimer le 31 octobre 1973.
IMPRIMERIE NATIONALE
3 564 001 5
Recommandations aux auteurs
Les articles à publier doivent être adressés directement au Secrétariat du Bulletin du
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gnés d’un résumé en une ou plusieurs langues. L’adresse du Laboratoire dans lequel le
travail a été effectué figurera sur la première page, en note infrapaginale.
Le texte doit être dactylographié à double interligne, avec une marge suffisante, recto
seulement. Pas de mots en majuscules, pas de soulignages (à l’exception des noms de genres
et d’espèces soulignés d’un trait).
Il convient de numéroter les tableaux et de leur donner un titre ; les tableaux
compliqués devront être préparés de façon à pouvoir être clichés comme une figure.
Les références bibliographiques apparaîtront selon les modèles suivants :
Bauchot, M.-L., J. Daget, J.-C. Hureau et Th. Monod, 1970. — Le problème des
« auteurs secondaires » en taxionomie. Bull. Mus. Iiist. nat., Paris, 2 e sér., 42 (2) : 301-304.
Tinbergen, N., 1952. — The study of instinct. Oxford, Clarendon Press, 228 p.
Les dessins et cartes doivent être faits sur bristol blanc ou calque, à l’encre de chine.
Envoyer les originaux. Les photographies seront le plus nettes possible, sur papier brillant,
et normalement contrastées. L’emplacement des figures sera indiqué dans la marge et les
légendes seront regroupées à la fin du texte, sur un feuillet séparé.
Un auteur ne pourra publier plus de 100 pages imprimées par an dans le Bulletin,
en une ou plusieurs fois.
Une seule épreuve sera envoyée à l’auteur qui devra la retourner dans les quatre jours
au Secrétariat, avec son manuscrit. Les « corrections d’auteurs » (modifications ou addi¬
tions de texte) trop nombreuses, et non justifiées par une information de dernière heure,
pourront être facturées aux auteurs.
Ceux-ci recevront gratuitement 50 exemplaires imprimés de leur travail. Ils pourront
obtenir à leur frais des fascicules supplémentaires en s’adressant à la Bibliothèque cen¬
trale du Muséum : 38, rue Geofïroy-Saint-IIilaire, 75005 Paris.