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AM THE
JOURNAL
OF THE
BOMBAY NATURAL HISTORY SOCIETY
INDEX AND TITLE PAGE
VOL. XXXII
PARTS 3 & 4
MADRAS
PRINTED AT THE DIOCESAN PRESS
1928
INSTRUCTIONS TO BINDER
The contents of these two parts should be arranged in the
following order when they are being bound :—
Title page... a es
Contents of Parts 3 and 4, Vol. XXXII
List of Contributors i, ... | To follow the frontis-
List of Plates... re ci piece in this order.
Index to Illustrations as
Errata se. oe P|
Index to species res ..- To goat the end of
two numbers.
THE
JOURNAL
OF THE
BOMBAY NATURAL HISTORY SOCIETY
EDITED BY
P. M. D. SANDERSON, F.zs., S. H. PRATER, c.M.z.s.
& SALIM A. ALI, m.B.0.0v.
VOL. XXXII (=
Parts 3 & 4
Containing 3 Coloured Plates, 65 Black and White Plates,
38 Text-figures and 3 Maps
Dates of Publication
Part III. (Pages 397 to 616) |... 15th January, 1928.
» IV. ( , 617 to 817)... 31st May, 1928.
LONDON AGENTS
DULAU & Co., Ltd., 32, Old Bond Street, London, W. I.
PRINTED AT THE DIOCESAN PRESS, MADRAS.
1928
CONDENTS OF VOLUME XXXII
No. 3
THE GAME BIRDS OF THE INDIAN EMPIRE. (W2th a coloured
plate.) OF THE EaSTeERN COMMON CRANE WMegalornis
grus lilfordt AND THE DEMOISELLE CRANE Anthropotdes
virgo. By E. C. Stuart Baker, P.Z.S., F.U.S., M.B.0O.U.,
H.F.A.O.U-; ee
REVISION OF THE aa Gi OF THE ROR TAS beg ESIDENCY. oe V.
By Wa biatver and, ©. McCain Jaz eeicecuseuwetacdctconece eke
PAGE
397
408
THE HOME OF THE EASTERN GORILLA. (Witha map and ten ~
plates.) By Marius Maxwell..
INDIAN DRAGONFLIES. Part XXIX, “(With ee blates.) By
Et.-Col. F, C. Fraser, 1.M,S.,-F.8.S:. ae
A SIND LAKE. (With six plates and yee Tae Lene ‘By
Salim A. All... :
THE LANGURS, OR par homens OF Sees ieee ‘(With
one coloured and one black and whtte plates and twenty-one
text-tieuyes.) By K. 1. Pocock, F.R-s..
A COLLECTING TRIP TO LapaKk. Part I. With a ioe ee,
iive plates.) “By V5. LVaPersonte... a
A List oF ORCHIDS, WITH SOME News Suneiaees FROM THE
HicH Wavy Mountains (Mapura District). (Wzth a
PGC) By OIE oS. ND, hale. Sy oh sometyade acy tea cari
AN ACCOUNT OF THE PEARL FISHERIES OF ‘TUTICORIN,
MarcCH AND APRIL, 1927. (With four plates.) By P. R.
Awati, I.E.S.. oa Se Re Ae ate Sati Mode ties cachaiae
MoDERN Was Me ODss pea I. (Wz2th stx plates and
ite textizgures.). By ©. H. Prater, C.M.Z.s.. ae
BomBAy NATURAL HISTORY SOCIETY’S Mian Sorat OF
INDIA, BURMA AND CEYLON. Report No. 46. By T. B.
A PRELIMINARY REPORT ON THE ANATOMY AND LIFE
HISTORY OF THE COMMON EDIBLE BACKWATER OYSTER,
Ostrea madrasensis. (With two plates). By S. T. Moses,
MBAy 2 FZ. 5 RoRsAcl.: .
ORNITHOLOGY IN THE ae By the ee ie [Col C. E. ane
DINERO Mauser 8 Sas eee ater eect ck fae< stidlem: ase dalsexs cate os adabeeed one She
@ BIW Vireo team erediw ooh tae iio owas aw dod eT EY cdva see ees ea kees
436
450
460
472
505
518
524
532
545
548
og
o/ 1
a5
589
CONTENTS OF VOLUME XXXII
iv
MISCELLANEOUS NOTES :—
I. Readers’ Views on ‘ Game Preservation in India. ’
By W. M. Logan Hoine.. ; ase
II. A Unique Lion BRO ter oi “(With a piney =
Salim A. Ali.. ve Vat a
III. White Tigers. (With a phen ape By FB. 5
Robinson.. : yak
IV. A Pussle eee aver aon Buffalo ‘Bait. ‘ay
Ss a\iro.C-Cliver: ;
V. Unusual Benaviour he Teeeen ithe Live Bait.
(With a plate.) By F. O. Cave...
Vi. ‘What is the Record Panther ?* by Ww. M. dean
Home.. : aSadleys SONS Tae eRe eee
Vito A Beane rer Flashi@ent: “(With a plate.) By
B. Avs. Brunskills ae :
VIII. Boldness of Banter ie W. M. ie ‘Homer
IX. Circumstantial Account of a Panther attached oy
Wild:Dogs: By Je McC Clive, Ce eee
X. Wild Dogs. By J. C. Faunthorpe.. M
XI. How Wild Dogs kill their Prey. Be A A. Daal
Brander... ie Sees eh eee
XII. The Record: Eoqcnoned tantslopee ( Tetraceros
guadricornts). (Witha photo.) By C McCann...
XIII. Abnormal Blackbuck and a Good Chinkara Head
from Bikanir. (Wz7th a plate.) By Surajmal
Singh... eae: jodie haNaiis eee
XIV. Clicking Meiee ae te Munn ey Charles
Theobald, F.zZ.S.. RPE ee et Ue ee
XV. The Bray of the Chital ee oe W. M. Logan
Home.. Ra ame rh ok
XVI. Musth in See: cae N. , Coen Rep eee hk eee a
XVII. - A Large Tusker Blephant. By A. W. John...
XVIII. A Good Nilgiri Tahr (Hemztragus hylocrius.) By
A. We JSonn.. Ee ete nts Ai te
OX Birth:<Ot Frente Calf at igo "(With a photo.)
By C.°E. -D: Mears... setae eae
XX. An‘ Example of an Assisted iDascase! es Hugh
Whistler, F.L.S., F.Z.S.. aoe Tae
XXI.- Extraordinary Pa enuciie saa hel Ruse. Cheniea
Scimitar Babbler Oo eee Me ce)
By Arthur H. Berriff... ivha as
XXII. Occurrence of the Wode ne in ane ‘Conteal
Provinces. By J. McC. Clive......
PAGE
592
Ss)e)
599
600
CONTENTS OF VOLUME XXXII
XXIII. Notes on Duck Shooting in the Roorkee District,
7 U.P. in the years 1903 to 1927. CE ee
By A. H. Cunningham... : SAG ge
-XXIV. Snipe and Woodcock in S. Hectic aS Ha, G.
- Phythian Adams.. Sere
XXV:. A Correction to Mr. ss 8 Osiastons ss NRE on
oe the Birds of Kashmir’. By Hugh Whistler......
Mex LE Aan Aibino Turtle: By B.A.~D’ Abreau, F:Z.S.....
XXVII. Notes on the Desert Monitor (Varanus griseus)
" and the Spiny Tailed Lizard ( Uvomastix
microlepis). By Norman L. Corkill... ce
XXVIII. Hamadryads in the Kumaon Terai. By, W. M.
Logan Home.. : eae Dee
XXIX. Mer. Slater’s Death a Saale! Bite. ins Charles
oe ‘+: : Theobald.. a
XXX. A Note on ne (ones sais onae. Tb yonis
- myctertzans) and Young. Cee photo.) By
C. McCann.. :
- XXXI. A Record She Nesneee ‘(With a Pein) ne
G. A. Hare.. :
XXX. “Aquatic anchor in hire eoliecion of the ye
| ‘cultural College, Coimbatore, S. India. By
CedticWDiOvCres win aliens eu eee ot been emu
EpR@ OI DENG Shae tele antec vc erst Somtinr pemsee teeeed anne ce oc oaesk es
No. 4.
THE our BIRDS OF THE INDIAN EMPIRE. (With a coloured
plate.) OF THE BLACK-NECKED CRANE Wegalornis nigrt-
colitis, “By B.C. Stuart Baker, ¥-Z.S.) 81S. M.B.O;U.,
BLIGH CA@r incre sinc te ieee errant, cota th anusgia wade eo ee alan Sais
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY, Part
Wik By re blauten S.J ePlicl Hal Gr 42 ds:4Mecale abe,
A COLLECTING Trip TO LApak. Part Il. (W2th two plates.)
Vg N aS: ECT SONG ek iio bata tel uel Wie seo cotenaa tyes
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA. Part
Il. (With two plates and three text- -figures.) By R. I.
HO COCR MREG: att secnen | eel ne soe een. Bumaeys! a, MEd
THE Bic BULL OF SEMBUTOLLAVU. (Witha plate.) ByC. P.
CLOGS Wii vase tens hot Canes Si eentekcss Sth catkins. uatuats:
INDIAN DRAGONFLIES. Part XXX. (With three plates.) By
ew oll. ©! Nraser, 1.M.S.5 FoEsS 20 occ de cen Lo. eakentencce
THE STUDY OF PLANT Lire. Partl. (With two De nid
Sieve lext-tioures.) By Charles’ McCann, ..-. ses. 0.09: ecs cc.
9
e
608
610
611
612
613
614
616
617
622
650
660
678
683
692
vi CONTENTS OF VOLUME XXXII
Notes AND DESCRIPTIONS OF FISHES FROM CEYLON. (With
two plates.) By Tieniy WV emer ae 9.) es ee eee
THE MAMMALS AND BIRDS OF KASHMIR AND THE ADJACENT
Hit. PROVINCES, “RartVil, By ColscA. Be ward &
FORAGING TERMITES. (Lutermes biformis, Wasm.) (With a
plate.) By R. W. G. Hingston, Major, I.M.s., ae
FURTHER NOTES ON BIRDS ABOUT SIMLA. By reese Whistler
LD Pits Gt 0) Ais Wa rere od te ney Runs chins Pry cone Ge HEVIA babi SA MeSt gy rl nact nie
SOME NEW SPECIES OF PLANTS FROM THE WESTERN GHATS.
(With a plaice.) By le Blatternand ©. McCann a.c.c- ace
BIRDS OF PREY AND THEIR Uses. (W/7th four plates.) By
Cy BH. Donald, 2rez:s< -M. BiQaUs 3 sa eee oe eee
A NOTE ON THE BIRDS OF THE PESHAWAR Districr. (Witha
map.) By Rev. F. S. Briggs and B. B. Osmaston,
M.B.O0:Ui; 2.BsS. ((RCtda) tea ena en me eee ee eens
MODERN Museum Mrtuops. Part Il. (Wzth two plates.)
By > Hie <Prater, G2 beZi6 eee ate ae an eet aan are ee
REVIEWS :—A eee oor OF THE BIRDS OF EASTERN CHINA.
Part LV.
avals: 6.0.0 e700 16,4 05a 8,0 bole; 'ete/e) 6 ake. 66.6 6 eiererepes @pelelere 98:9 aera
THE LITTORAL FAUNA OF KRUSADAL ISLAND 1 IN
THE GULF OF MANAAR WITH APPENDICES
ON THE VERTEBRATES AND PLANTS...........
With A CAMERA IN TIGHR: ANDi. .c.-ccu eee ee
DWELLERS IN THE JUNGLE 0 ae ee
EDITORIALS A ne ee cee J Pia Ade igs a ee
OBITUARY. 2S cee Pn nee ee
MISCELLANEOUS NOTES :—
I. ‘A Unique Lion Photograph.’ By F. W. Champion,
LFS and téeply by-SalimyAd Alt... sense oes
II. A Clouded Leopard (Felis nebulosa) attacking
Man. (With a plate.) By A. Browniow, Indian
Police sfc. 2 sas eae nt ele toa lee eee eer
III. Noosing Tigers. By H. A. Boas.. ae
lV. Reply’ to Mr: Phillips” Note onin: “371, ok
XXXII ve Colour of the Long-armed Shonthe
tailed Bat (Zaphozous longimanus.) By C.
MocCann, 33. gacscictt cages ee tees nena eee
V. Habits of the Porcupine. By Ralph C. Morris..
VI. An Elephant with extraordinary ‘ Glands’. ( With
a plate.) By A.S. Laurie... Ae
VII. Ons Natural Deaths’ in Wild aleohante ie
Ralph: ©.aMortis, «can ..2). ot cee ee eer oe
788
789
790
791
Tou
792
«194
VIII.
XII.
XIII.
XIN
XV.
XVI.
OVO
XVIII.
XIX.
XX:
OE
CONTENTS OF VOLUME XXXII
Clicking Noise made by Muntjac. By B. B.
Osmaston, Indian Forest Service (Retd.). .......
Grows and —<Koels; By E. <A. Andrews,
Entomologist, Indian Tea Association
The Rufous-breasted Blue Flycatcher ( Cyornts
hy perythra) at High Levels. By C. Boden Kloss
The Nidification of the Sind Hill Warbler (Suzya
crinitgera striatula). By H. W. Waite,
eereceec eee eee
F.Z.S y
BE BuON Win. Lil cia OLICE! nannies eer a seine cobs dette
On the Unusual Food-habits at Horibills, By
Satya “Churn. WawieMes., BL. Ph.D.,. F:Z.S.,
Wee By ORWeeth, area cats wanes eek he oka etary staat a8
The Black-capped Kingfisher (Halcyon pileata) i in
the Darbhanga District, Bihar. By Chas. M.
[ne@lis, F:Z.S:,. F-B.S., M.B.O.U.. aoe
A Quail’s Queer Retreat. Reseed: fon ihe
Sunday Times, Johannesburg, Transvaal,
INFO VERIDEE LS p20 ok NaN Mews ool ida eee. cel oes
Occurrence of the Sheldrake (7% ador na 2 tadorna) in
the Darbhanga District, Bihar. By Chas. M.
MMe liSy -Pe2(S4 Fae .Sa 5, Me BcOe Uni. foc tinctaek Se pew eatiee
Occurrence of the Sheldrake (Zadorna tadorna) in
the United Provinces. By Capt. BE. E. L.
Seats ots Cat) creas fee oes 20 oh els cbs ee te a,
Recovery from the Bite of a Phoorsa (&chis
By Lieut.-Col. A. A. Meaden,
The Mogul Emperors of India as Naturalists and
Sportsmen. By Sunder Lal Hora, D. Sc., F.t.s.,
F.z.S., Zoological Survey of India, Calcutta......
The Effects of Bee Venom. Reprinted from 7he
Indian Medical Gazette, Vol. LXII, No. 5, May
1927. By A. Bayley de Castro, Junior Medical
carinata).
eee eeeree © F&F eee F-8OF ses eoersr neers reres ZF © ee oF FF Hoe OOD
Plants and Insects. eye H. Jouguet... a
Weeds of the Indian Wayside. By H. idagaees
PROCEEDINGS AND STATEMENTS OF ACCOUNTS.
eoe eee eer ees O82 FT eFe
vii
PAGE
795
796
796
OSL
798
798
799
793
800
800
802
805
807
810
813
ALPHABETICAL LIST OF CONTRIBUTORS
VOLUME XXXII
Nos. 3 and 4
PAGE PAGE
ppAMS Major E. G. PHYTHIAN, BEATTER,- 7 Sede oe alae
I. A.; Snipe and Woodcock in F.L.S. and McCann, C. ; A list
South India : 606 of Orchids with some new
ALT @OALIM oN.4 VA Boing ale species from the High Wavy
(with six dates and three text- Mountain (Madura District)
figures ) a feet 7.20 2460 (wth a plate) : : 518
—_—— ; An Unique Lion ; Some new species
Photograph (wih a plate) 583 of Plants from the Western
— ; ‘A Unique Lion Ghats (wzth a plate) 733
Photograph ’ (a reply) 788. | Boas, H. Ag Noosing Tigers .:. “790
ANDREwS, E. A.; Crows and Bor., N. L.; Musth in Ele-
Koels : a .. 796 plant. yee: ; nee we <OUe
AWATI, PoORG tee 5oseean | BRANDER, A. A. Dunbar How
Account of the Pearl Fisheries Wild Dogs kill their Prey 591
of Tuticorin, March and April, Briccs, Rev. F. S. AnpD
1927 (with four plates) 524 OSMASTON, B.B.,M.B.O.U.,
BAKER, Bi Co STUART ers I.F.S. (Retd.) ; A Note on the
| DDR Cr a le Anes Wiles eOe Oia. Birds of the Peshawar District
H.F.A.O.U. ; The Game Birds (with a map) ae we «744
of the Indian Empire, Vol. BRowntow, A.; A clouded
V. The Waders and other Leopard (Felis nebulosa) attack-
Semi-sporting Birds. Part V. ing Man (wzth a plate) 789
(With a coloured plate). The BRUNSKILL, Major B.A.S.;
Demoiselle Crane, Eastern A Panther and Flashlight _
Common Crane ke om O97, (with a plate) . ios O08
-—___—- 7 Parte Vi aKa acne CASTRO, A. Bayley de.; The
a coloured plate). The Black- Effects of Bee Venom . Aire teil)
necked Crane, The Great CAVE, @ CAPD... uOce Unusual
White or Siberian Crane, The ; behaviour of a Tigress with
Hooded Crane ... 617 Live Bait... vee ese ROOT,
BERRIFF, ARTHUR H.; et. CHAMPION, F. W., I.F.S.; ‘A
oalinieray Pugnacity of the Unique Lion Photograph’ ... 788
Rusky-cheeked Scimitar Bab- CLIVE, Capt. J. Mc C., Mce>:
bler (Pomatorhinus erythro- A Tussle between Tiger Baa:
genys) ye Bete + gu OO Buttale Bait. so... eee 586
BRATTER seryn moe) ce Ph.D., : CircHeetneunl
Boe Sond se McCann, 2 Cx: Account of a Panther attacked
Revision of the Fauna of the by Wild Dogs 590
Bombay Presidency, Part V. a MO coutrenee? of tite
Gramineeé ; we 408 Wood Sine in the Central
——— ; Part VI. PLE, 622 Province ~., a 6
600
LIST OF CONTRIBUTORS
PAGE
CorKILL, NorMAN L.; Notes
on the Desert Monitor
(Varanus griseus) and the
Spiny-Tailed Lizard (Uromas-
tix microlepts) ... sae
CUNNINGHAM, Capt. A. H.;
Notes on Duck shooting in the
Roorkee District U.P., in the
years 1903 to 1927 (wzth a
graph) a ar
D’ABREU, E. a oF. Z. S. An
Albino Turtle ..- oe
DONALD, Celis: F.Z.S.,
NEB.O Us ; Birds of Prey
and their Uses (wth four
plates) ane See
DoveR-CEDRIC; Aquatic
Rhynchota in the Collection
of the Agricultural College,
Coimbatore, South India
EDITORIAL oe;
FAUNTHORPE, Coron a Ci
Wild Dogs
FowLeR, H. W.; Notes Pe
Descriptions of Fishes from
Ceylon (with two plates)
FRASER, Lt.-Cou. F.C., I1.M.5.,
F.E.S.; Indian Dragonflies
Part XXIX (with three plates)
oe Pt, KO (eh three
oe
RRy. Be; Cenper Nataral
History Society’s Mammal
Survey of India, Burma and
Ceylon, Report No. 46, on a
Collection from ‘Toungoo,
Burma ae ie
GOULDSBURY, C. 'P. The Big
Bull of Senmibnrollava (with a
tlate) Be aes
HARE, Lt.-CoL. G. A:;
Record Cubbany Masheer
Hincston, Mayor R. W. G.,
I.M.S.; Foraging Termites
(Eutermes biformes, Wasm.)
(with a plate) :
Homes, Mayor W. M. crate
The
Readers’ View on Game
Preservation in India é
; What is the Reco
Tiger
608
600
608
737
450
683
544
678
613
716
581
587
ix
PAGE
Home, Major ;
Panthers ...
Boldness of
; he Bray of the
Chitral Stag oe
; Hamadryads
the Kumaon Terai as
HorRA, SUNDER Lat, D. Sc.,
Peso HeZ.5.13> Che Mocul
Emperors of India as Natural-
ists and Sportsmen as
INGEIS, (C.mASG Ma) BAS.
F.E.S., M.B.O.U. ; The Black-
capped Kingfisher (Halcyon
pileata) in the Darbhanga Dis-
trict, Bihar
——ees
in
; Occurrence of the
Sheldrake (Zadorna tadorna)
in the Darbhanga District,
Rihar
John. AgwwW. s A Large: seer
Elephant ee wae see
——_ 4 Good ) Niloizi
Tahr (Hemitragus hylocrius)...
JouGuET, H.; Plants and
Insects ay aes ite
; Weeds of the
Indian Waveinee
KLoss, C, BODEN * The Rarouc:
breasted Blue mighnbone:
(Cyornis hyperythra) at high
levels fh ae ie
LAPERSONNE, V. S.:; A Col-
lecting Trip to Ladak, Part II
(w2th two plates)
LaurIg, A. S.; Elephant mie
Extraordinary ‘ Glands’
Law, Satva CHURN,
Beene Lee ge Me Ze See
M.B.O.U.; On the Unusual
Food-habits of Hornbills
Luarp, 1L1-Cor, (Cy, Eie(The
late) ; Ornithology in the Bible.
McCann, C. ; The Record Four-
horned Antelope (Vetraceros
guadricornts) (with a photo) ...
; A Note on the
Green Whip Snake (Dryophis
mycterizans) and Young Pe
; The Study of
Plant Life, Partl. (Wzth two
plates and five text-figures) ,..
a
M. A.,
588
594
610
802
798
799
596
597
807
810
796
650
792
798
593
592
612
6
‘CO
WO
x7 LIST OF CONTRIBUTROS
PAGE
McCanvn, C. ; Reply to Mr. Phil-
lips’ “Note sons ip. 3/71) Vol:
XXXII ve Colour of the Long-
armed Sheath-tailed Bat
(Zaphozous longimanus) ... 791
McCann; See Blatter, E.
MACKIT#,. n:-@o 1m. VE Ve;
R-AL.MéC. > = Indian “Snake
Venoms ... : Mes set iE
MAXWELL, erercee The Home
of the Eastern Gorilla (wth a
map and ten plates) Ex pe a'4 36
MesDEN, LT.-CoLt., R.A.M.C.;
Recovery from the Bite of a
Phoorsa (Echis carinata) ... 800
MEARS? Ci -E.- 1D. > Birth sof
Freak Calf at Indore (with a
photv) Ee oer ee oT
Morris, RALPH C, ; Habits of
the: Porcupite: 3: ste set OW
——— ; Elephant with
Extraordinary ‘ Glands’ saa 7/88)
“NiO? Nvat area
Deaths’ in Wild Elephants ... 794
MOSES. (5. 1s, NiAg,. SEZ. os:
Baa Ae eee. ~ Preliminary,
Report on the Anatomy and
Life-History of the Common
Edible Backwater Oyster
Ostrea madrasensis (with two
plates) —::: a 2 es Ot8
OBITUARIAL NOTICES 530, 786
OsmastTon; B. B:, -M.B:O:.U.,
Los. (Retds)s Clicking
Noise made by Muntjac wn OS
; wee Brices, Rey:
—————
| sesh
ROCOCK. OR. «Fy ESR aS che
Langurs, or Leaf Monkeys, of
British India, PartII (with
PAGE
two plates and three text-
figures) ... re a oe OO
PRATERS Oe. Hs Ce M.2 Z. eS.5
Modern Museum Methods,
Part I (with six plates and
five text-figures) Nese Oe
—— ————; Part II (with two
plates) ase es Oe
REVIEWS 57 laa 73
ROBINSON, F. B. . White Tigers. 584
SEARIGHT, CAPT. E. Bb Rie
Occurrence of the Silelarabe
(Zadorna tadcrna) in the
United Provinces ads 6 800
SINGH, SURAJMAL; Abnormal
Blackhuck and a Good Chin-
kara Head from Bikanir Sent O93
‘SUNDAY TIMES’, JOHANNES-
BURG; A =gQOuail’s'~ Oweex
Retreat ... Rae 99
THEOBALD, CHARLES, FB. Z:S.
Clicking Noise made oy
Muntjac ... ss 593
——.— —- ; Mr. Staters’ beet
by Snake bite es che soe) Od
“WAITER: HS W.) Fee ZeeSs
M:°B.O.U: ; fhe -Nidification
of the Sind Hill-Warbler
(Suya crinigera striatula) ... 797
WARD, Cot. A. E.; The Mam- ;
mals and Birds of Kashmir
and the Adjacent Hill Provin-
ces; Part Vil. =: 71
WHISTLER, HuGH ; An neamle
of an Assisted Passage ee DOS
; A Correction to
Mr. B. B. Osmaston’s Notes on
the Birds of Kashmir ... O07
; Further Notes on
Birds Snot Simla ee ix = 726
LISTE OF PLATES
VOLUME XXXxXIl
No. 3
PAGE
The Game Birds of the Indian Empire—
The Eastern Common Crane (Megalornis grus lilfordi). The Demoi-
selle Crane (Anthropoides virgo) ae ae seer OOF.
The Home of the Eastern Gorilla—
Plate I—No. 1. Mt. Mgahinga (11,400 feet) with Mt. Mukavura
(13,547 feet) in the background . .. 436
No. 2. Mr. James Hugh Barnes, the wien: com-
panion, with the Porters. The Precipitous
Peaks of Sabinio in the background 436
Plate II—No. 3. Panoramic View of Mt. Mikeno (14,385 fect)
and Mt. Karisimbi on the left (14,633 feet)... 438
No. 4. Panoramic View of Mt. Sabinio ... 438
Plate III—No. 5. A Typical Forest of Mt. Mikeno (alt. about
11,000 feet) showing Hagenia trees in tall
undergrowth ss 439
No. 6. Another Forest Scene on “Mt. Milcene at 11, 000
feet altitude, showing a portion frequented
by Gorillas.. 439
Plate IV—No. 7. A Forest of are Deepa on : Mt. Maeno
at an altitude of over 9,000 feet . .. 440
No. 8. Arboreal Heath (Arica arborea) and Senecio, a
Giant Groundsel (Senecio johnstonz) 440
Plate V—No. 9. The Summit of Mt. Mikeno (14,385 feet)
showing alpine vegetation on the slopes of
the peak ~ ... 44]
No. 10. A Startled Female Goniia tating off with
young on her back ... rote wes 441
Plate VI—No. 11. An adult Male Gorilla ... re .. 442
No.12. A pair of female Gorillas secured on
. Mt. Mikeno ae oon, 442
_ Plate’ VII—No. 13. Four captured young Gonles ees we 443
No. 14. One of the four young Gorillas, with its
guardian .., : 443
Plate VIII—No. 15. A Gorilla shelter in the roan me ofa iene ge
. tree Me 444
No. 16. A recently vacated hehe shelter of a family’ Gt
Gorillas showing the excrements of the
. Apes (X) in the right foreground .. 444
. Plate IX—No.17.' A Gorilla ‘Bed’ or ‘Nest’ formed in the
is crotch of a bifurcated tree on the western |
ae slopes of Mt. Karisimbi at an altitude’ of ©
i ee | some 11,000 feet os we, ww 445
xii LIST OF PLATES
Plate X—No. 18. Dead Leopard on the branch of a ‘large
Hlagenia tree, showing a vacated Gorilla
shelter below
Indian Dragonflies—
Plate I—1. Wings of Rhinocypha bisiguata, Selys, Male
2. 6 s btforata delimbata, Selys,
Male..
3. Wings of Rhinecy pha elenien Pers merche
Male ... ae
Plate II—1. Wings of Riper MEERA, herby: cite
2: ; Micromerus lineatus, Burn., Male
Sh ar Calocypha laidlawi (¥ras.) Sips
Plate JII—land2. Two Views of the Cauvery River near
Fraserpet, Coorg By abe
A Sind Lake—
Plate I— 1. Negotiating the Reedy Marginal Shallows
2. Nearing the Butts ies
Plate II— 3. The Fowlers ieaving the bane
4, The Fowlers about to enter a Flock
Plate III— 5. A successful Catch
6. The Gazis in action :
Plate IV— 7. Red-crested Pochards in a hues Net |
8. Removal of Captives from Dhubbi
Plate V— 9. An Egret Farm on the Manchar
10. Interior of the same
Plate VI—11. Fishing on the tienehae = aneine the Caen
12. Another Snap of the Landing Operations
The Langurs, or Leaf Monkeys, of British India—
Plate I— Newly-born Langurs 1. The auratus-Group; 2. The
aygula-Group ; 3. The Antellus-Group ae
Plate II—l. The Kangra Langur (P. entellus ajax) ...
2. The Nepal Langur (P. entellus achilles)
3. The Common Langur (P. entellus entellus)
Plate Iil1—Map of Kashmir and Ladak
A Collecting Trip to Ladak—
Plate I—1. House-boat arabella on Wular Lake
2. Sind Valley, one of Kashmir’s Picturesque Side
Valleys : ive see
Plate II—1. Baltal, 9,000 Feats at ne Head of the Sind
Valley
2. Kolahoi, 17,800 eae a Sonepicdous Pea in nthe Great
Himalayas, with its Glacier a LO;aat hie
Tree-line :
Plate III—1. A Morning’s Bag of Show: (oe: a Mauibecs
11,000 feet ‘ “ts
2. Market Place at Leh, and ee Gompat above the
Winter-Quarters of the Ecclesiastical Head of
Western Tibet
Plate IV—1. Looking through the Carerara of Leh to the Taeae
Range , are cs te
2. Leh in Spring ime abs ai aes
PAGE
446
450
450
450
451
451
451
456
462
462
463
463
464
464
465
465
466
466
467
467
474
478
478
478
502
506
- 506
507
507
510
510
511
511
Plate
LIST OF PLATES
V—1. Our Transport of Hybrid Yaks
2. Camp at Chang-Chen-Mo Valley, 14, 500 feet
Orchids from the High Wavy Mountain (Madura District)—
A. Odontochilus rotundtfolius
B. Diusperts neilgherrensis we hi ae Big
Pearl Fisheries of Tuticorin—
Plate
Plate
Plate
Plate
I—1. Pearl Bazaar Guarded by Police oe Lie
2. A corner of the General Bazaar, Pearl Fishery
Camp
II—3. Divers’ Encampment
4, Divers at Work
III—5. Division of spoils at Sea
6. Bags of Oysters
IV—7. The Return of the Divers
8. Washing the Pearl Oysters
Modern Museum Methods—
Plate
Plate
Plate
Plate
Plate
Plate
I—Plate of Domed Ceiling of Bird Gallery, American
Museum of Natural History, New York
II—Model of Exhibition Hall, designed for the New
Museum of Natural Sciences, Buffalo, New York
IlII—New Reptile Hall, American Museum of Natural
History, New York, showing ‘ Built-in’ Groups
IV—Model! of New Hall of African Mammals, American
Museum of Natural History, New York :
V—Curved Background in course of preparation for a
Group at the American Museum of Natural History,
New York
VI—A case of birds snowing the ole jens meted of
Exhibition
Life-History of the Common Edible Backwater Oye =
Plate 1—Anatomy of Ostrea madrasensts
2— re) oe) i)
Young Lion attacking a Zebra
A Tigress with its kill
A Panther and a Cow ‘< Kill’
Abnormal Blackbuck shot in Bikaner—
A Good Chinkara Head from Bikaner, Horns, 144” use
Record Cubbany Mahseer, weight 105 lbs.
No. 4
The Game Birds of the Indian Empire—
The Black-necked Crane (Megulornis nigricollis)
A Collecting Trip to Ladak—
(A) The Rocky Road to Yarkand
(B)
Our Camp at 15,500 feet in the Kareena
Stoliczka’s Tomb, Leh ... ="
The Langurs, or Leaf Monkeys, of Baten rade
Plate
late
Il1I—The Capped Langur and Young (P2thecus pileatus)
IV—The Dusky Langur (Pithecus obscurus)
The Big Bull of Sembutollavu
xiii
PAGE
514
514
921
221
526
926
527
227
528
528
— §29
529
536
So/
538
539
542
543
550
551
583
587
588
593
613
Ziv VAST OF PLATES
Indian Dragonflies—
Plate I—Micromerus greent, M. aurantiacus, M. andamanensts,
M. finalis, M. lineatus indica, M. lineatus lineatus,
Calocypha laidlawt, Libellago astatica, L. curta
Plate II—Two Views of the River Cauvery, Coorg-Mysore
Frontier en mee at ae
Plate Ill—-Libellagine Larve. Fig. 1. Rhinocypha ignipennis,
Fig. 2. Micromerus lineatus indica, Fig. 3. Rhino-
cy pha bisignata
The Study of Plant Life—
Different Types of Roots eet
(A) Aérial Adventitious Roots of fe Hanan awe Wen Lees bse
(B). Another type of Adventitious Root. The ‘Screw Pine’
(Pandanus sp.)
Fishes from Ceylon—
Plate (—Cypselurus hallbergi
Plate Il--Atherina duodecimaltis
Foraging Termites—
Eutermes btformis, Wasm. 6
New Species of Plants from the Western Ghats—
Barleria gibsonioides, Blatter, sp. nov.
Birds of Prey and their uses—
1. (A) Bonelli’s Eagle (Avzeretus fasciatus) ...
(B) Golden Eagle (Aguila chrysetus daphaneé)
2.(A) Eastern Goshawk (Astur gentilis schvedowt)
(B) Laggar Falcon (Falco jug ger) Ae
3. (A) Eastern Steppe-Eagle (Aguila nipalensis Abalone)
(B) Booted Eagle (Hieretus pennatus)
4.(A) Kestrel (Falco tinnunculus) A
(B) White-eyed Buzzard (Butastur teesa) ...
Modern Museum Methods—
1. A Lesson in Minerals—Children’s Museum, Brooklyn, New
York e's tee oe. we ac
2.(A) The Vivarium (a great centre of attraction to young
visitors )
(B) A corner of the walle used Lipa
Clouded Leopard (Felis nebulosa) ...
Elephant with extraordinary ‘ glands’
PAGE
686
688
690
700
701
701
706
707
718
733
738
738
739
739
740
740
741
741
768
769
769
789
792
TMA PO TELUSTRATIONS
VOLUME XXXIi
Nos. 3 and 4
PAGE PAGE
Anthropoides virgo, Pl. 397 5. A Typical Forest of Mt.
Aguila chrysetus daphane, Pl, Mikeno (11,000 ft.) .. 439
fig. 2 ; 738 6. Forest scene on Mt. Mikeno
——— nipalensis nipalensis, PI, (11000 ft:) PI. 439
fig. 1 ate 740 7. A forest of Hagenia aye.
Astur gentilts apeedowe. Pl. Gee sinica on Mt. Mikeno(9,000 ft.) 440
1 : oh es 739 8. Arboreal Heath (rica
Atherina duodecin:alis P\. 707 arborea) and Senecio, a giant
Barleria gibsonioides, Pl. 733 Groundsel (Senecio john-
Blackbuck (abnormal) shot in stont), Pl. : ee .. 440
Bikaner, Pl tig! 593 S. Summit of Mt. Mikeno
Bos gaurus, Pl. : 678 (14,385 ft.) 441
Butastur teesa, Pi. fig. B. a. 741 10. A Startled Gorilla, Panine
Calf born with bifurcated Week, off with her young on her
Photo on OO back,. Pl, 44]
Calocypha Mialeon “Pl, ae 3... “451 11. An adult maie Celie 442
Ae ae Pl. fig. 10... 686 12. A pair of female Gorillas
Chinkara. Head from Bikaner, secured on Mt. Mikeno, PI. 442
fig. 2 593 13. Four captured young
Cy pselurus fallbencs. Pl. .. 706 Gorillas eS ... 443
Disperis neilgherrensis, Pl. fig. B. 521 14. One of the four young
Dryophis mycterizans, and Gorillas with its guardian,
Young, Photo ; 612 Plies? qichin loud ge see ota
Elaphus indicus, Tiskerenomine 15. A Gorilla shelter in the
peculiar gland-like growths on hollow-base of alarge tree... 444
neck 8 792, 16, A recently vacated night
Eutermes Pies Pl. ee oe 718 shelter of a family of Gorillas,
Falco jugger, Pl. fig. 2 739 Ph Ge eae eg ae aed
—- tinnunculus, P\. fig. 1 741 17. A Gorilla ‘ Bed’ or ‘ Nest’
Felts nebulosa, Pl. seen i sh’) on the western slopes of Mt.
Gorilla, Home of the Eastern-- Karisimbi (11,000 ft.), Pl. ... 445
Map of Uganda ; 436 18. Dead Leopard on the
1. Mt. Mgahinga, (11,400 ft. ) branch of a large Hlagenta
with Mt. Muhavura (13,547 tree, showing a vacated
2 ft.) he ee; 436 Gorilla shelter below 446
oe MYrJ., EL, Barnes with the Hieretus fasciatus, Pl. fig. 1 738
ih Porters, Pl... of 436 — pennatius, Pl. fig>2- 740
-3. Mt. Mikeno (14,385 ft. ) and Indian Dragonflies—
Mt. Karisimbi (14,633 ft.) ... 438 Views of the Catvery Ree eee
4 Mt. Sabinio, Pl... 438 near Fraserpet, Coorg, Pl... 456
Xvi
Indian Dragonflies—
Two views of the River Cau-
very, Coorg-Mysore Frontier,
Favourite haunts of many
species of, Pl. wes
Kashmir and Ladak, map of.
Ladak, a collecting trip to—
House-Boat arabella on Wular
Lake, Pluie. 1
Sind Valley, Pl. fig. 2.
Baltal, 9,000 ft. at the Heae ot
the Sind Valley, Pl. fig. 1 ...
Kolakoi,17,800:ft., Plone. 1;
A morning’s Bag of Snow
Cock at Moulbeck, 11,000 ft.,
Pl. tis. rs
Market place at Leh ane Aine
Gompa, PI. fig. 2
Gateway of Leh, PI. fig. 1
Leh in Springtime, Pl. fig. 2.
‘Transport of Hybrid Yaks, PI.
pce OU es
Camp at Chang: bonene ‘Mo Val-
ley, 14,500 ft., Pl. fig. 2
Rocky Road to Varkaad Camp
at 15,500 ft. in the Kara-
KOfanm: ies on
Stoliczka’s Tomb
Langurs, newly born, Pl.
Libellago astatica, Pl. fig. 11
—__—_—_— curia, Pl. fig. 32
Lion, (young) attacking a Zebra
Macaca irus, Stomach of, fig. 1.
-- silenus, Hand of, fig.
— Foot - a
The Record Cunpaae:
Masheer,
Photo
Megalornis grus Die PL.
-—_—--——._nigricollis, Pl.
Micromerus andamanensis,
fig. 3 ‘
—_——--—— aurantiacus, Pl. fig. 2,
—__—_—_--—— finalts, Pl. fig. 4
CECT MP figs.)
PL
—_—-—— lineatus, P|. fig. 2...
—— —___ —__———_— indicus, Pl.
fig. 5. 3.
—_—_—_-—_—___——-——— Pl.
fic. 2. larva nc. ee
—_—_—__-- Bodies. Pl.
figs. 6-9
PAGE
688
502
506
506
507
507
510
510
511
oll
514
514
INDEX TO ILLUSTRATIONS
PAGE
Museum Methods, Modern—
Part of Domed Ceiling of Bird
Gallery, American Museum
of Natural History, New
VYoruc el: 536
Model of Exhibition Hall ne
signed for the New Museum
of Natural Sciences, Buffalo,
New York, PI. Pen OSG
New Reptile Hall, American
Museum of Natural History,
New York, showing ‘ Built-
in’. sroups, PI: 538
Model of New Hall of Atuican
Mammals, American Museum
of Natural History, New
York, Pl. 539
Window-seat for Victiore: co 539
Type of Desk Case in use in
the Geological Gallery,South
Kensington, fig. .. 540
Curved Background in course
of preparation for a group at
the American Museum of
Natural History, New York,
Pi; a eee nee soe, O42
A case of Birds showing the
shelfless method of Exhibi-
tion, Pl. 543
Side View of Group Cae. aoe 543
Plate I. Children’s Museum,
Brooklyn, New York Pl. 768
LI >(A); Viavarinms.< Pl: 769
(B) Library, Pl. eaata ge Act]
Odontochilus rotundifolius, Pl.
He paw hoy. ios RE see el
Ostrea madrasensis, Pls. 550, 551
Panther and a Cow ‘ Kill’ 588
Pearl Fisheries of Tuticorin :— $
Pearl Bazaar guarded a Police -
Plone 526
Corner of the Concea Dasaen,
Pearl Fishery Camp, PI. fig.
2 nee See so an E20
Divers Encampment, Pl. fig. 1. 527
at work, Pl. fig. 2 fee Ded
Division of spoils at sea, Pl.
sehen Wma = 528
Bags of Oyen Pl a 2 528
Returao of the Divers, Pl. fig.
i ave ct son) hese eo
INDEX TO ILLUSTRATIONS
PAGE
Washing the Pearl Oysters, Pl.
fig. 2 aes
Peshawar District, eifap of
Pithecus auratus, Pl. fig. 1
— entellus, Hand of, fig.
Foot of, fig. ..
—_—__—_——_-—_ P.. fig. 3 :
achates, skull,
fig.
———_—— — —_ achilles, Pl. ae
2 3
—_—_———— - ajax, Pl. fig. 1.
skull, fig.
——_—_—_—_——_——- entellus, PI. fig.
3 :
skulls, figs.
——_——__ — ———._ hector,
fig. :
—— —- —_—- inlus, skull, fig.
a —--——— -—— pallipes, skulls,
figs.
—— —_ —-._ priamellus, Frais,
figs.
———_———— schistaceus, skull op
ae aaee aa
———— femoralis, Pl. ae 2
Sa keatit, figs.
skulls, front and side views ...
——— obscurus, Pl,
pileatus,
es er
ee
skull,
adult and
young, Pl.
pileatus, figs.
—— ee
skull, front and side view
———— pyrrhus atrior, fig.
skull, front view :
—_— —— shanicus, fig.
skull, front view
———— sener nestor, skull, fig.
vetulans, stomach
of fig.
Plant Life, the Study of—
Dicotyledonous seed, section of
fig.
Menecotviedonone nee
tion of, fig.
Box of soil with seed, Ae
Box containing seedlings,
illustrating tendency to
sec-
485, 486,
529
744
474
473
474
474
489
478
478
482
478
487
483
491
497
495
478
474
676
668
662
662
671
671
499
472
694
694
696
XVil
PAGE
turn towards source of light,
fig’. 697
Seedling of a Bean Ponting
the parts of which it is com-
posed, fig. ; 698
Roots, Different pes: of, Pl,
figs. 1-10 700
Aerial adventitious Roots of the
Banyan, (Ficus bengalensts),
PY axe ae a LOL
Adventitious Root, another
type of, Pl. ao 701
Rhinocypha biforata delonbatn
Pipi 2 450
—__—_—_-——- bisignata ... 450
———_—___—__-—__-—-— Pl. fig. 3
Larva ... re aes ... 690
od tontpennis, Pl. fig. 1.
Larva ... os bis ... 690
—— perforata perforata,
Pl, ie. 3 — e tp 400
—— whiteheadi, Pl. fig. 1. 451
Sembutollavo, Big Bull, Pl. 678
Sind Lake, A
(a) Negotiating the Reedy
Marginal shallows 462
(6) Nearing the Butts, Pl. 462
(a) The Fowlers leaving the
Punt 463
(6) ————-—— about to enter
a. Flock, Pl. 463
(a) A Successful Catch 464
(6) The Gazis in action, Pl.... 464
(a) Red-crested Pochards in a
Dhubbi net 465
(6) Removal of ae om
Dhubbi, P .. 465
(a) An Egret ea on the
Manchar ».» 466
(6) Interior of the same, Pl.... 466
(a) Fishing on the Manchar—
Landing the catch 467
(6) Another snap of the Land-
ing Operations, Pl. .. 467
Tetraceros quadricornis, Photo
of Record Head... 592
Tiger (white variety) 584
Tigress and its kill 587
ERRATA
Vol. XXXII, Nos. 3 and 4
Page 514, line 1 for furruginea read ferruginea
506
566
566
656
656
659
749
740
749
+)
tbe
9)
> din
9
Me)
379:
99
39
38 for chenchrts read cenchris
20 for Cicona read Ciconia
30 for Cotyle read Cottle
10 for Pyrrospiza read Pyrrhospiza
9 from bottom for Allen Hume vead Alan Hume
2 for Octocris read Otocoris
1 for fulginosa read tuliginosa
38 for mustcapa read muscicapa
42 for musicapa read muscicapa.
INDEX OF SPECIES
Acanthis flavirostris baltistanicus
ladacencis
Accipiter nisus nisosimilis
-— virgatus affinis
Acridotheres ginginianus
tristis
———- tristis
Acrocephalus agricolus
—————— eo
PAGE
515,
2 CONCINeENS CONCINENS ...
Adansonia digitata
AXgithaliscus concinnus
Zfrides cylindricum
AEthopyga gouldize
—— siparaja
Agrion fenestrata
fulgipennis
lineata
perforatus
Agrostis ramosa
verticillata
Alauda arvensis dulcivox
- gulgula ... ;
Alcedo atthis as aie
——_——_-—— _pallasi
Alectoridia quartincaria
Alectoris graeca chukar
—---------—-——. pallescens
Alloteropsis
——--—— — cimicina
—--———-— similata
Alseonax latirostris
Alternanthera echinata
Amandava amandava ...
Ambassis gymnocephalus
——-—— safgha
-——- ——. uroteenia
Amia endeketzenia
haringtoni.
stentoreus brunnescens.
683,
Ammomanes deserti phoenicuroides.
Ammoperdix griseogularis
griseogularis ...
658
658
757
729
752
726
192
607
607
607
7S!
700
746
520
UCT
727
683
683
684
451
636
409
754
754
756
468
418
O12
572
635
635
635
749
811
752
708
708
708
708
755
758
Ammoperdix heyi
Amphilophis compressa
——_——-—— concanensis
—_——-—-——- ensiformis
-—- ——— glabra
—_——-——- k untzeana
odorata
————— _ —,—_- Pertusa
woodrowii
Anabas testudineus
Anezctochilus elatior
Anapheeis mesentina
Anas platyrhyncha
——- p. poecilorhyncha
Anatherum muricatum...
Andropogon acicularis
aciculatus
——— ---—— alectoridia
-annulatus
——-_-—_——_ aristulatus
armatus
arriani
- asper
—_—_— ———_ assimilis ...
—_——_——_ aucheri
—_—_ -—-— —*—-- genuinus
-— ——_——_ besukiensis
bicornis
breviaristatus
—_——-——-_ brevifolius
—_—— ——. calamus aromaticus
——_-_——_— caricosum
———_—-—— cerriferus .. ;
——_—— ciliolatus ceruleus
-——— increscens
—_——__——- circinnatus
——_———— citratus
—————_ citriodorum
—_——_———- compressus
—_-_-——_-——-. concanensis
—_—___—_———- contortus
ceokei
xx INDEX OF SPECIES
PAGE PAGE
Andropogon debilis.... oe ... 428 | Andropogon pachnodes oe beer ees
demissus ... ae ... 429 | ————-—— pachyarthrus ... we. 429
-— -———— echinatus es ... 417 | ———--——— parvifolius Ms w 428
——— echinulatus ee .. 411 | ——-—~—— pertusus ae ee 4)
——- ensiformis set 421, 422 | —_-___-____- —______ var. genuinus... 421
—— ———— festucoides ee .. 409 | ————~-——— ——-wightii.. 421
——_——— filiculmis ... aoe -. 420 | --———— polphyllus ak .. 416
——. Horidus 4:. ene we = 428: | -———-— —— polystachyos ..-622, 623, 624
ee foveolatus s .. 427 | ————— prionodes po we 417
——e glaber .. ae .. 422 | —————— pumilus ia ee 429
——_——-—— glabratus ... 3 . 411) — punctatus Sas sos 422
—__——_-——- gryllus.... i . 411)—— quadrivalva age .. 1630
———_-—— heteroclitus ae .. 631 | ———— ritchii one 622, 623
—— ———— hispidissimus ... we = 625 | —————— roxburghii i ons -, 432
—_—_——_--— hugelti. ... ve .. 421 | —————— royleanus Bs w= =6 4]
————-——— humilis ... see .. 623 | —————— schcenanthus 430, 432, 433
--—- ——_——-. intermedius ee w 422 | ——————- —__—----——- var. cesius 434
a subvar. gla- SS SS = - genuinus’- 433
ber .. 422 | —-— “= martini 433
———— —__ ——-—_ var. pune- SS ee ALS ay woe 417
tatus' .... 0 4225) =———-_- —_—_‘sprengvelit wee om ei: 81S)
——-——-—— ischynanthus... .. 632 | ———— -- squarrosus Af .. 409
—_—_——_-——_ jwarancusa nis . 431 | —-—_——_— — var. genuinus 409
or ae es PAnISCr hist AO eee strictus aA : 427
——__—__ ———_-—- proper Sot 432 ——_——_——- tenellus See Bae 428
a var. genuin- —_———_——- trinii fon le weep) 2413
us ... 432 | ——————triticeus ... Aa 622, 623
——_——_—_-—— —__--__ -——. —-tantum. 431 | ———- um bellatus ae wae 7 O31
—_———— kuntzeanus to we. 422 | ———--—— violaceus 23 win 418
—————_——— lancearius aa .. 412 | ———— wightanus a Same
————-——_ lanceolatus . ay. | WOOd Tonal ais wer AO)
Lt eee So -——~ subvar. ser- Androscepia tremula ei be) -O3:
rulatus .... 417 | Anisops sardea SAS is see 615
ee -——-var. genuinus 417 | ——-—— niveus oe He wi, OLS
—__——-——-laniger ... @ ... ... 430 | Anser anser ve sc oe. 260
eee LAWSON ee SLAs mae 409 ——-— indicus oe axe mal 470
———_—__— _ Jjananthus he .. 623 | Anthistiriaargentea ... i: re O28
i be martini ... Pee mee 433 —_———_— australis aaa ot see 629
SS —_ -— Mmonomeros de ets 631 —_—— ciliata oe wes 628, 630
————-—— monostachyus ... *%. 427 | ———-—— ceespitosa ne tte 51-45:
ee Ont anus are ... 420 | ————— cuspidata os «ss ~~ 628
= = jonticola S .. 413 | ————— cymbaria te i401 de OOU.
bE ee igor velutinids aloo depauperata 230 ee) ees
ee I TICaAtiS a nee. 409 oa desfontainei ate isle 628
— ——_——_ nardoides Ae we 433 | —————- forskahlii oes -- 628
———_———_—— nardus var. ceriferus ... 432 | ——-——-— glauca vee .. 628
as nutans... eo ... 630 | —~——— heteroclita oss cen, -- 63]
=. _-__ optusifoliue hee. Hse 1428 |e hispida bent |) users 028
——— odoratus ... = .. 424 | ————— imberbis oe -. 628
————-—— oliganthus i .. 623 | ———-——laxa tee -. 628
— Olivieri .y6 as = 0,2 430 |] —— paleacea rete = 628
—___ + schmidtii
INDEX OF SPECIES
PAGE
Anthistiria punctata 628
——————. scandens 630
———— semiberbis 630
—-——_-———. syriaca 628
——— —— tremula 631
——— — vulgaris a 628
Anthracoceros m. Ee labaticus TAs:
Anthropoides virgo 398, 469
Anthus campestris 754
richardi rufulus 754
spinoletta blakistoni 754
Apis melifica 805
Apluda gryilus 4}]
Apus melba ToL
Aquila chrysaétus dapharias 738
- clanga 20. 469
heliaca 469, 757
nepalensis 731
rapax vindhiana 757 |
Arboricola torqueola millardi Reed Loe
Ardea cinerea 563, 760
cinerea 471
grus 402
purpurea ... 563
virgo 398
Ardeola grayi 760 |
Argya caudata 746
_-—— earlii 746
pathraxon ciliaris ae 41
ae - quartinians 418
————- cloratus 418 |
———— cuspidatus 418 |
——_—— echinatus 417 |
-—__— —— inermis 416
-—_—__-—— jubatus 418
—_—_———- lanceolatus ... eae a7
_——— ——-— —- var. schinatis.. 417
——-— lancifolius 418
—-———— langsdorfii 418
-——_———- major 418 |
———_-——- meeboldii 418
——-———— microphyllus 418
—— ——- var. aneifoling 418
————_——- minor 418
-—__——— molle 418
——-——- multicaulis ...° 418
—_—— —— nudus 418
——— —— plumbeus 418
—_——. quartinianus 418
Ea SC uper 418
418
| Arthraxon serrulatus
submuticus ...
—_—_-——- violaceus
—— vriesii
Asio flammeus cee
Astur gentilis schaedowi
Athene glaux ie
Atherina duodecimalis ...
— forskalii
— lacunosa
Atta barbara
Axonopus
-— cimicinus
Barbus hexastichus
— tor
Barilius bola
Barleria gibsonioides
Batratherum echinatum
-———— ——-— lanceolatum
molle
schimperi ..
serrulatum
Belostoma indicum
Bos gaurus
Botaurus stellaris
Brachiaria
—$—— ———
distachya
——-——— distachyum ...
——_—_---— isachne
——_——. mutica
————— notochtona ...
——_—_——- prostrata
—-— ramosa
Bubo bubo bengalensis
Buteo ferox 5
Bulbophyllum milenemenee
Butastur teesa Pe
Buteo buteo rufiventer
Caccabis chukor ...
XX1
PAGE
TAQ;
.. 785, 802,
637,
638,
Te IO al AU;
Cacomantes merulinus passerinus
Calamina imberbis _
Calandrella rufescens persica...
Callacanthus burtoni
Callosiurus ferrugineus...
Calocypha laidlawi
Calopteryx lineata
Calotes versicolor
Camponotus compressus
Canis aureus
——- lupus
——- pallipes
683,
417
418
418
418
756
739
565
706
706
706
724
635
635
785
804
804
733
417
417
418
418
417
615
678
760
636
639
639
637
637
645
643
645
756
797
519
Tod,
471
559
fa7
628
755
727
546
457
684
809
724
713
712
713
Xxil
INDEX OF SPECIES
PaGE
Chrysopogon wightianus var. leucan-
thus
Chelidon urbica ...
Chrysoglossum ace
-—_—_——-—— hallbergii
Ciconia alba
— nigra
Cinnyris asiatica...
Circus zeruginosus
cyaneus
-——- macrourus
macrurus
Cirrhina mergata
— mrigala
Cirrhopetalum gamblei...
- —— thomsoni
Cisticola juncidis cursitans
Cleisostoma tenerum
-—— stocksii
Clione vastifica
Clupea ilisha
—-—
| Coelogyne corrugata
| ——--—— nervosa
Columba livia, subsp.
Conitermes similis
Coracias benghalensis penphalonee as
— garrula semenowi
Cordiochloa cimicina
—- fimbriata
Corvus corax aan
———- ——- tibetanus...
- cornix sharpii
— —+—- corone orientalis
| ———- splendens
- Cotile TIPEStiiSis:s
PAGE
Cannomys pater ... 547
en plumbescens 547 |
Capella gallinago gallinago 759
Capillipedium assimile ... 420 |
—_-——_-——_ filiculme 421 |
——-—— -—— hugelii 421 |
Capparis aphylla 808
—————. spinosa 808 |
---~ zeylanica 809
Caprimulgus sp... 756
Caranx ignobilis.. 707
- sexfasciatus 707 |
Carine brama - 737 t|
Carpodacus erythrinus .. 152 |
——- thurus blythi oe sO 4|
Casarca ferruginea 514, 760 |
Catla catla ba 802, 804 |
Centrophorum chinense : is 412
Cepalopyrus flammecips Tre oat 7208
Cephanodes hylas eee oleh
Cercopithecus cephalopterus ... = S01
—— kephalopterus ... isheeos4 98
—__——_—_ -—— senex oe 498, 500
——— vetulus .. 498, 501, 504
Certhia himalayana 508, 747
Ceryle rudis leucomelanura 468, 756
Cettia cetti cettioides fio
Chaimarrhornis leucocephala ... 748
Charadrius dubius 759
Chettusia leucura = 469
Chlamydotis undulata macqueeni 759
Chlidonias leucopareius 760
Chrysopogon aciculatus 412
asper 412
——_——— auccheri 416
—_—_——_-—— ciliolatus ... = 413
———_——_—_—_ ——-——_ auccheri eee AlS
—_——— ——_ ——-——-_ var. aucheri... 416 ©
creeds as 413
—————— esenbeckii 413
—__———- glabratus 411
—_——_——_ glaucopsis 421
——_—_—— gryllus - ... 411
————-—— increscens 413
—_—__———. lancearits 412
== ae On LantS, 413
= omnibusware 413
——__——— pictus © 420
ol payilucs 416
—_—_—_———. serrulatus 413
: 412
‘Wwightianus
coronoides intermedius
lawrencii .
ruficollis
--— frugilegus 360
———- monedula scemmeringii
——- + zugmayeri ...
Coturnix communis
———— coromandelica
Cot tmnix ae,
ee OEE
- Crinum eleonore
Cuculus canorus... .
Culicicapa ceylonensis ...
Cuncuma leucorypha
Cyanosylvia suecica pallidogularis
Cygnus bevicki Det gee pore =
413
2 366
519
519
566
566
755
469, 757
757
471
757
468
804
519
519
75]
521
408
vi 550
468, 803
—-§20
520
758
723
755
755
635
635
745
512
745
wa. 655
655, 745
rn 745
745
745
654, 745
ake w598
655, 745
2. 4566
802,
757
658, 749
7€0
—s.
INDEX OF SPECIES xxiii
PAGE PAGE
Cygnus cygnus se re entice 760) | Digitaria royleana oe oe “ae 633
~ olor ee. oe .. 760 | ————sanguinalis ... ws 632, 634
Cylindrostethus Reeders Mae ee OLE Ware = - var: Ciliaris - 633
Cymbopogon arabicus tas bees 4230 ————-~ tenuiflora ea oe w. 634
! arriani nae ue .. 430 —-——ternata he a wat Oe
SS —— CSIUS ae fae ve 434 ———— vaginata oak oe .. 641
—_——_——- circinnatus ee .. 430 Dipcadi saxorum sah ees ae. 730
—_—_———- citratus... wi en oe -- ursule ... se ee ind Vh35
eee jwarancusa ss ... 431 | Disperis neilgherrensis .. Re vu C2
— ei —— artini... ee we = 433 -- zeylanica © ae — ae gow
—_——-——_ martinianus a ... 433 | Dodoncea vicosa . oe oe ae WhO
schoenanthus ... ... 430 | Dryobates He pedy hier. sts ete Ae
Cynodon ternatum ae ... 632 | ——-—---scindianus ... i te bio)
Cynopterus brachyotis anealate ge Odd Dryophis mycterizans ... sy ell ie Ole
Cyon dukhunensis ys ee 714, 780 | Duranta plumieri sed es ee = CUS
Cyornis hyperythra Bue se .. 796 | Echinochloa ac xi wes see: 22040
Cypselurus hallbergi... oe aw) 705 -- - colona ‘ oa . 646
oxycephalus ae .. . 706 | —-—— ——-—— var. frumentacea. 647
—— sinius oh ses .. 706 | ——-—- commutata bs, ... 647
Dendrobium nanum ... a ee —_——-+——- cruciformis ie oe WOSt:
—_—_—__-—— nuttans.. .. 618 | —— - crus-galli ie ste. OEF
—_——___-—— -. ——- var. Fubrclapner 518 -_——--——-- frumentacea Ste .. 647
———_—— sp. age aa .. ol9 | ——-———- hispidula ... a vais, « O47
Dendrocygna javanica ... Bie ae 700 ——-——-scabra_... = .. 648
Derris robusta ... as a .. 496 -stagnina ... a .. 648
Diceeum ignipectum ... = eee ee Echis carinata... ios bs 800, 801
Dichanthium ae . ds ... 623 | Egretta alba bes sot ae 470, 760
anata ae 425, 426 | Eicharniacrassipies _... ace ... 810
—__——_——- armatum oe .. 425 | Hlaphiis indictis ... ae abs 792, 794
—_—= - caricosum ee .. 425 | Emberiza cia ee a a eed VATA
—_——_-——-- mccannii re = 425 | ~ ——- —— par es we as © Jol
—-- panchganiense ... .. 425 | -——-—— cioides vayans ay soar LAPS
— + - serrafalcoides ... ac. 7427 -——— ——-- fohkienensis se, AUS
Dichoceros bicornis bat 798 | ———-—- godlewski_... a ee, Me
Dicrurus leucocephalus innoicaadatns 750 | ——-—~—- icterica a 2 Se)
———-— macrocercus ... a? ... 450 | -—-——-- leucocephala oo fo 18
Digitaria affinis ah ae moa 2 O42 ——— —-- melanocephala or fon 18
—- appressa aie oe .. 642 | —-—— schcemilus pallidior... re:
—-— chrysoblephora s .. 633 | Emyda granosa intermedius ... J. | -608
———— ciliaris var. quadristachya... 633 | Engraulis commiersont... ee .. 104
--——-~+ commutata’ ... ee .. 633 | Enithares indica... ane a OS)
———— distachya ae se .. 639 | -—~——-—triangularis ... oes foe SOLS
—— — fimbriata ae its ... 633 | Eremophila alpestris deosai ... sss 7 G09
———— foliosa i Ve ea Ga — longirostris ... 512
—-— linearis oe sas .. 634 | Eremopogon foveolatus ee ae ed
———- longiflora see sea. .-Od4 paranipyeanunm ... 2H a7
—-— marginata var. Ginbtiata a. , Ooo. tia braccata ... a oe Some hsv ES)
———— paspaloides var. longipes ... 641 | —— nana... an a ne sa, 019
ao PCUICEllariS” ... ie .. 634 | —— pauciflora ... au obs sceay)
—— —— pennata aes A .. 634 | —— pseudoclavicaulis ... bee see ue OO
foo Pseudo-durva *... ae fen 034 reticosa A m st ae golG
XXKiV
PAGE
Eria arborea 439
Ericholoa .. 636
— annulata 636
——-—— hackelii 636
—— -— polystachya ... 636
-——- ramosa 636
Erolia minuta 759
Erythrina rhodGen (sae 651
—— rubicilla severtzovi ... oie HOO
——-—— rubicilloides lapersonnei ... 651
Eudynamis scolopaceus malayana 796
se -- scolopaceus. 756
Eunetta falcata ... 605
Eutermes biformis Gd
——-—— vastator 724
Exonantes ee 706
Falco chiquera enlatiers = a abe ele)
——~ jugger 739, 740, 758
——- peregrinator 725,
——- subbuteo Pre {0
- tinnunculus 741, 758
Felis chaus aa 18
nebulosa 789, 790
Ficus bengalensis 701
Franklinia buchanani (py!
Fringilla montifringilla ded
Fulica atra 759
atra 469
Galerida cristata cheadesla 755
-——--——- magna 755
Gallinago nemoricola 600
Gallinula chloropus ee wef hohe,
Gamphrena decumbens 810, 811
Gelochelidon nilotica 760
Gerris adelaidis 614
——-- anadyomene 614
—--- lucidus 709
—-- paludum ... 614
—-- spinole 614
Glaucidium brodiei 728
Glaucionetta clangula ... 605
Glossogobius biocellatus 710
Gomphrena globosa 811
Gorilla beringeri... “436-449
Grus communis ... 402, 563
——- leucogeranus 618
—- lilfordi 402
——- monacha 620
——- nigricollis ... 617
Gunomys varius ... 547
Gypeetus barbatus 558 757
INDEX OF SPECIES
PAGE
Gypetus barbatus hemachalanus 728
Gymnanthelia lanigera 431
Gymnema sylvestre 703
Gymnorhis xanthocollis 753
Gyps fulvus “ze 558
Habenaria peristyloides 522
aa stenostachya 522
Hagenia abyssinica 439
Halcyon pileata 798
-— smyrnensis fusca 799
——_—— smyrnensis 756
Haliastur indus ... 757
Helopas annulatus 636
~--- ——= leevis es 636
Hominieeiodee shaeinvinyien 704
————- --—— neglectus 705
———_—_- -——__ unifasciatus ... 705
Hemitragus hylocrius .. 597
Hetropogon 3 622
—_-——_--—-— contortus ... 623, 624
——_——_—_+—_ var. genuinus,. 624, 625
—_-—_ —___-—. glaber 624
———_—— genuinus sub-var.
hispidissimus 624, 625
a ee —— roxburghii ... 624
-——— -___—_ ——. ——— — secundus 624
—__—_—_-——-— typicus 624, 625
——_——— insignis 623
——-------—- oliganthus 622, 623
——_—— —— polystachyos 623
——-—_-——_ ritchiei 623
Hevea brasiliensis 703
Hiereetus fasciatus 728
pennatus 738
Hilsa ilisha wes.» SOS
Hirundo daurica 754, 773
———— nippalensis 773
—--— PUSticae ss : 566
— TUSEICA es 753
——--— smithi filifera 754
---—---.-— striolata 113
Holcus gryllus... 411
——-— pallidus 41]
~ pertusus 421
Horeites pallidus Pallas 726
Horornis pallidus 751
Hyena creata TVA
——-—- striata 711
Hydrometra greeni 614
——_-——--- — lineatus 614
aa vittata 614
INDEX OF SPECIES
PAGE
Hydroprogne caspia caspia 469
Hypacanthis spinoides 793
Hypogynium fovuolatum Soe LACT
Ictinzetus malayensis perniger 728, 738
Inocotis papillosus 470
Iphigenia stellata 734
Iseilema anthephoroides 626
———— laxum 627
—__—— wightii 627
Ixobrychus minutus 760
Tynx torquilla 5 eo
Josephia lanceolata 520
Labeo calbasu 804
——— dyocheilus i ... 804 |
——— rohita 458, 802, 804
Laccotrephes griseus a Ole
_ ruber 615
Lanius cristatus isabellinus 750
excubitor lahtora 750
schach erythronotus 750
vittatus 750
Lantana camara 812
Larus brunneicephala ... 516
Lawana conspirsa 806
Leiognathus splendens 707
Lethrinus reticulatus ae, 409
Libellago ; 683, 691
—- asiatica Seven 459
-———-- bisignata 455
——-—-- blandus 687
————-- lineata 684
——_——-- perforata 451
Limosa limosa limosa Po. opal)
Liopicus mahrattensis purocrier ates aoe, CTS)
Liparis viridiflora 518 |
- wightiana 518
- sp. 518
Lobivanellus indicus 759
Lophoceros birostris 798 |
Loranthus sp. 701
Luczea ciliata 418 |
Lindia maculata hess OU |
Liiscinia svecica ... mes 654
MED abot . —-654
————- ——- —- pallidogularis 654, 658
Lusciniota melanopogon mimica Zou
Lutjants kasmira 709
Macaca mulatta Des 545
——-—-silenus ... be 43
———-vius a 72
Macroniés seenghala BO
aes
KKV
PAGE
Malachra rotundifolia .:. 810, 811
Mareca penelope : 761
Margaritifera vulgaris ... ta, Dee
Marmaronetta ee $3 605, 761
Megalornis grus . -402
—_——- frond 402, 469
——- leucogeranus 618
—— — monachus 620
~ —— nigricollis 617
Merganser castor | 13
—— merganser aricntalts - 761
Merops or entalis... VSI)
persicus persicus 756
superciliosus javanicus... 795
Messor barbarus ‘ 724
Metaponia pusilla som 7% 7953
Metrocoris staii ; 614
Micromerus Andon iene 687
-——-- aurantiacus 690
——_—-——- finalis 688
—___-——-- greenei ss 069
——_—_—_———_-- lineatus 683, 686
—— ——- - blandus . 687
————____-—_—— indica yh xe)
—__—-- —— —-—- lineatus ... 684, 686
—- obscurus 684
- uxor 684
Micropus affinis ... 756
Milium cimicinum 635
— compressum 635
——— - crus-galli 647
— filiforme ... 634
— ramosum 636
— sanguinale 634
Milvus ater ay 558
migrans govinda ee)
regalis gree orto)
Molpastes heemorrhous irermenit: ea, 240
———--—— leucogenys humii 746, 747
——---—- ——-— ieucogenys 746, 747
——--——- leucotis _ 746, 747
Monocharia hasteefolia ... 810
Monodactylus argentens.. | 710
Monticola solitaria pandoo . 749
| Montifringilla brandti eT VOR ye 513, 650
Motacilla alba dukhunensis 468
——-——— —-— personata © - 154
ae cinerea caspica wy j -T-n 754
——-—- citreola _ eve 6 468
ee eT VINEE EES calcar ata . Peete =|
oa flava cre Uy 43 eo ABB
XXvi
PAGE
Motacilla ava thunbergi 754
Mugil longimanus 707
Murex tenui spina 550
Muscicapa hypoleuca : 749
—— striata neumanni ... 749
Myiophoneus temminckii 749
Nelumbium speciosum ... Bene
Neophron percnopterus 508; /0V
Nettion crecca crecca 761
Nucifraga hemispila 5 726
--—— multipunctata 507, 745
Nycticorax nycticorax ... 760
Nymphea lotus . 461
Nyroca fuligula ... 605
- marila marila 605
- rufa : 761
- —— rufa 470
Oberonia brunoniana 518
-—— lindleyana 518
——-—— iridifolia 518
——-—— verticillata 518
Odontochilus rotundifolius 2k
Odontotermes fex 719
CEcodoma sae ee
CEcophylla T23°RL29
(Enanthe capistrata 748
deserti 748
——-—— isabellina 748
——-——— opistholema ... see 148
picata ... coe 658, 748
——--———- xanthoprymna chrysopygia 748
(Enopopelia tranquebarica tranque-
barica 758
Onychotrechus oaaan 614
Oplismenus crus-gallz .. 547
-—— frumentacens 647
—_—_--—— limosus 647
———-+——-. scaber 648
-——- stagninus 648
Oreocincla dauma 749
-— mollissima simi dienes |
Oriolus oriolus kundoo ... 752
Orthopogon crus-gallt... 647
647
-—— stagninus ... 648
Ortygornis pondicerianus 758
Ostrea angulata .. 548
edulis TOON
cucullata ... .. 548
gryphoides var. cuttackensis . 548
madrasensis — 548
INDEX OF SPECIES
Ostrea virginiana
—-—— virginica. ...
Otis tarda. ane
Otocoris Sipcstic longirodeHoe
—— var. madrasensis ...
PacE
548
548
548
759
659
— (Eremophila) alpestiie déosai 659
Otus ascalaphus ... 565
- spilocephalus 728
Ovis ammon ee elt
Oxyura leucocephala 605, 761
Palzornis nipalensis eupatria 756
—-- torquatus : 756
Palumbus palumbus casiotis ... 758
Pandanous sp. 701
Pandion haliztus fog SY
Panicum affine 642, 644
— appressum 642
————. arabicum 646
——-—_ argyrotrichum 634
— —— arvense 638
———— awelianum 643
——— barbatum 643
——-— barbinode : 637
———— beckmanniz forme... 642
——~— brachylachnum 638
——-— brizeforme 642
— ——-— brizoides 642
——— burgi .. 648
—_— — cespitosum 643
—--—— calauzense 643
——— — carnostm 642
——-——. caucasicum 637
———- ciliare ... 633
——— cimicinum 635
cognatissimum 638
——— colonum 646
———— conjugatum 635
— — controversum 644
—— — crispum 643
—— cruciforme 31, SOS
——— crus-galli : 647, 648
——— var. colonus 646
—_—____ —_—_—__ -—__-—_. frumenta-
ceum 647
———_———___—___——_—— leiostachyum. 648
———_———__— maximum 648
wi —-———— stagninum ... 648
————o —— submuticum. 648
———- cuspidatum 646
—_-——— daltoni ..s 646
———-— distachyum 637, 639
ee ae
INDEX OF SPECIES
PAGE
Panicum equinum 637
—— —-— equitans ce 646
———— fimbriatum 633
——-— flavidum 641
——— floridum 642
fluitans.. ave O92
————. frumentacea ... 647
—_——- galli .. 648
———— geminatum 642, 644
———— glomeratum 642
-——-—— _ grossarium me. 1-043
—— —— helopus 2 vi 638, 644
—_——— —var.glabrescens ... 644
——— hirsutum 644
———— hispidulum 647
———— hispidum 647
—— — hostiti 647
———— insularum 647
————_ isachne... 633
————_ javanicum 645
kenegi 644
———— lielievrei 648
——_—— limosum 647
———— longiflorum 634
——— marathense 644
—-——— marginatum 643
-———— muticum 637
— —— notochtona ... Ree cea = O45
——————— NUMIdiIanum ... ... 637, 642, 646
oryzetorum 648
—_——- oryzinum 647
——— parvulum 634
——_—— paspaloides 642
—-—- patens ... 638
——-— pennatum ww. 634
———— petiveri sf i 638, 646
—— — nosteasintin var. GS 632
——-— pictum 648
—-—— procumbens 643
———— prostratum 643
——_—— pseudocolonum 646
pubinode 637
——— punctatum 642
————— purpurascens... 637
pyzmeum “2 203d
——_——— ramosum 637, 638
———— repens ... 643
———— reptans : ssaee “643
sanguinale var. alsnwsene
thum 633
ee ———— Ciliare 633
eee
XXVil
PAGE
Panicum sanguinale var. macrostach-
yum 633
————— sarmentosum ... 637
———— scabrum 648
———— setigerum j 644
——_——__——__-———- tomentosa 644
————- sieberi ... sae 043
———. stagninum ise 647, 648
—- —-—-—~ —— var. frumentacea 647
—--—— subquadriparum 639
——_—— ternatum 632
———— tetrastichon 646
————- trichopus 645
-—--—— truncatum 642
—-— turgidum 642
—-—— wumbrosum 638
————- viaticum 643
——— wightt... ioe ©1637,
— — zonale ... 646, 647
Papaver somniferum 703
Papilio agamemnon 811
—-——- aristolochize 811
- hector ane 812
Paradoxurus binneniods 546
niger ae .. 946
Parus major kashmiriensis 508, 512, 745
—-—-- melanolophus 508
——-- monticolus 745
——-- rufonuchalis ronuehalic 508
Paspalidium es 641
— flavidum ... 641
——— geminatum 642
punctatum 642
Paspalum ... ee 639
—-- annulatum 636
———--——. appressum 642
——--—— boryanum 641
-—-- brachiatum ... 641
——--—— brevifolium ... 634
——--——. canarze 640
———--——. compactunm ... 640
-———-- -- —— dilatatum .. 640
-—-----— distichum~ ... fe 640, 641
—_- — -- var. vaginatum. 640
——--—— filiculme 634
——-— -- foliosum 641
——-—— imperfectum ... 640
-—_——- -— inflatum 641
—_——-—— kleinianum 641
—_—— -— littorale ... 640
— longiflorum ... 634, 64]
INDEX OF SPECIES.
XXViii
PAGE
Paspalum miliaria 640
-— orbiculare §39
——--—— ovatum 640
——-— pedicellare 634
—— -— pedicellatum... 634
——--—— pennatum 634
——_-——_ pratense 640
——-— pseudo-durva 534
——-—— royleanum 634
———---- sanguinale 633
——___—____ -_-— - ——- var. ciliare 633
——-—— scrobiculatum : os 639
—-——_ —_ —_--—— var. com-
mersonii 639
+ —___ ——-——.- var. frumen-
taceum ... 639
—___—- —_— —--—_——- var. poly-
stachyum 639
—_——- — selloi... 640
——--— squamatum ... 641
——--—— ternaturm ay pe oe
——_--—— vaginatum as 640, 641
—-. — forma longipes. 640
Passer domesticus ees es),
——— hispaniolensis “753
Pastor roseus Zaz
Pelecanus crispus 562
——-- ---— onocrotalus ... 562
Perdix hodgsonie caraganaz... oly
Pericrocotus brevirostris brevirostris.. 750
Perristylus stenostachyus Be Ole
Petaurista cineraceus ae: 1 O46
—_—--—— philippensis... 546
Petrocossyphus cyanea... 566
Phalacrocorax carbo 562
-- pygmeeus 562
Phalaris zizanioides seen = 409
Pheidole ... he ae Ass 719, 724
Phoenicurus erythrogastra grandis 652
—-- erythronotus Li 748
—=-- ochrurus phcenicuroides
658, 748
Phylloscopus collybita tristis 751
—~--— subviridis Fo
Pica pica bactriana — oil
Picus squamatus 755
Pipatherum annulatum =... 1636
Pithecus auratus 660, 667
————_ aygula... 675
——— brahma 664
——'— cephalopterus 498
PAGE
Pithecus chrysogaster ... 2-2 666
———— crepusculus 667, 669
————- cristatus 667
———— durga ... a3 vee es, 1003
——— entellus 473, 476, 660
—-—- ——~—-- achates 476, 488, 489
—_—— ——-—--- achilles 476, 478
———_ —-—-- ajax ... 476, 480, 482
— -—-— -——-- dussumieri ... 477, 491
—_——- ——-—--- elissa 477, 493
—- -- entellus
476, 484, 485, 486, 487
-——-——- —__--- hector ... 476, 481, 483
———_— -- hypoleucos ... 477, 492
—_—— -- inlus ... 477, 490, 491
—_——— -—_—--- lanius ... 480
—_———_ -———--- ceneas 477, 492
—_—_—— -- pallipes 477, 497
——_——- -———--- priamellus 477, 494, 496
—— — -- schistaceus ... 476, 477
~—-——— femoralis [675
——_—— ——--— _keatii 676
——-——. germani 667
———— johnii ... S08,
—_—— margharita - | 667
——_—— melamerus 670
———— obscurus 667
———— phayrei 667
—-——— pileatus 660
————— -- brahma 664, 666
—_—— -- durga 663, 665
———— —-——--- pileatus __.... 661, 662, 665
—_———- ————--- saturatus 5, S003
—_—— ———--- shortridgei ... 663, 665
—__——— —_—---- tenebricus ... €64, 665
———— potenziani 666
-—— pruinosus 666
——_—— pyrrhus : 667
—_—— ———--- atrior 2;. 671,673, 734
Eatla di SePR -- barbei 768
as Le oe -- crepusculus . 672, 676
Se = AVICAUGA ee. 672, 674
—_—_—____——_ pha yyrei ... 669, 672, 674
ee Se San crortin 673
—shanicus _... 670, 671, 674
wos cehex 498, 660
a johnt : 503
eee 5 nestor .. ...499, 502, 504
—————— senex Pere es)!
—_— —————-_ vetulus ... 472, 501
—_——— shortridgei .. 665
ove \ onal
INDEX OF SPECIES
PAGE
Pithecus shortridgei belliger ... 663, 665
—vetulus 498, 499
= -— monticola 502
_ as nestor... 502
ee. -—__—_phillipsi 502
-—.-—— vetulus 501
Platalea leucorodia major 470
Plectranthus rugosus 808
——_—— - — ternifolius 808
Plegadis falcinellus 470
Pleuroplitis ciliata 418
——-——-— major 418
——-— -——— plumbea 418
——-——— schimperi 418
— ———-- vioJacea 418
Podiceps al bipennis 470
—-—— cristatus cristatus 652
—-—— ruficollis capensis 761
Pollinia brevifolia 428
———— fulva 413
——— gryllus 411
Pomatorhinus erythrogenys 599
Porphyrio poliocephalus poise phatn: 469
Presbytis albinus ... 900
——_——— barbei 668, 670
——_——. brahma ; : 664
———— cephalopterus var. monticnla 502
———— crepuscula 672
——__— —-— Sraieantinn 672
———— durga 663
entellus .. 484
SS — anchises 488, 493
——w—w— feimoralis 675
——_—_ johnii 503
———— lania 480
——-—— maurus see. 007
melamera 668, 670
— neglecta keatii 676
——_—— obscurus 669
— -—— carbo Heat = OWS
aia Phayrei 669, 672
——_—— pileatus 660, 663 |
priamus 495
— sanctorum 673
— schistaceus . 481
-———— shanicus 670, 671
— shortridgei 665
——— thersites 495
ursinus 502
Prinia gracilis lepida 751
inornata inornata 752 —
PAGE
Prunella atrogularis 749
———— rubeculoides 659
—_—— muraria 659
Pseudanthistiria heterodita zt 631
—. --—— hispida 631
—_——_--——-— umbellata wo, ~-631
Pseudogyps bengalensis 728, 757
Psilopogon schimperi 418
Pterocles corunatus atratus 758
— indicus : 758
Pteroclurus alchatus eamaseaene 758
- senegalensis 758
Ptilomera laticaudata 614
Ptyonoprogne rupestris 753
Pucrasia macrolopba biddulphi 507
Pygathrix cephalopterus 498
flavicauda 672
——_-——-— pileatus : 660
Pyrrhocorax graculus age tel olZ
-——-- pyrrhocorax himalayanus 512
Pyrrhospiza punicea humii 658
Pyrrhula aurantiaca 508
Rallus aquaticus korejewi 799
Ranatra elongata 615
Raphis ciliolata 413
coerulea 413
Rattus concolor 547
——w— rattus khyensis 547
rufescens 547
Ratufa gigantea 546
Rhaphis acicularis 412
—— -—~ echinulata 411
od gryllus 411
-——— — orientalis 412
——— trivalvis 412
— wightianus w. 412
Rhinocypha 683, 691
——- apicalis 451
——-——-- beesoni 453
——_——_——-- biforata abbreviata 454
——-—__——__——_—- beesoni 455
—__—_—__—___-—_-- biforata ... 453
—__—_—_-—_—_—_—- delimbata 453
——_———_- bisignata oe 455
-—_—— ——-- inas 451
—_—_——_-- laidlawi és 457
—_— ——- perforata Beatiien 452
ee — limbata... 451
—_—_ -—__—_—_ —, perforata 0° 452
——- whiteheadi 450, 451
Rhyacornis fulginosa 749
Rhynchops albicollis ever pk tOO
EXX INDEX OF SPECIES
PAGE
Riparia riparia indica ... one ces DS
Ritaarita---. ae ee oe 802, 893
Rostratula benghalensis benghalensis 759
Saccolabium filiforme ... Je A O20.
—-— gracile... ee eave
—_—_———— nilagiricum he ara 20)
—— pulchellum ae. ee O20
Sanguinaria vaginata ... oa ie Al
Satyrium nepalense Ase es EO
Saxicola caprata bicolor ae nanprean oF
———- torquata indica A ves) EAS
—_——_-——- leucura _... .. 448
Saxicoloides fulicata cambaiensis ... 749
Schizachyrium brevifolium ... we. 428
Schcenanthus amboinicum ... Aer iert: sey
Scolopasx rusticola ans oie ed EN)
Scotocerca inquieta striata... ‘ecto
Scotophilus castaneus ... pe. Boe ere): )
ecyllaiserrata: \\s.: bei aoe Pe sll)
Semnopithecus anchises Se we = 488
——__———--— argentatus hi 661, 663
——__——__———— barbei ... oe aan Ors
—— barbus ... at ‘ae 9068
—_—_—— dussumieri ... an eee
—_——_-——— entellus acs .. 484
—__——_ ——— femoralis i ZOO
— — holotephreus ... ne O69
—_—_—__ ——— hypoleucos ... x 492
——_———_-——— hypoleucus ... 221 488
——-——-_ — -—johnii ... ae Bee ar O03
—_——_——-— kelaarti ae fee, Soe
—_—— ———_ maurus a3 ace OOF
————- ——— neglectus “ee seta 07/9)
——_——_-_——_— nestor ... 0 Se Ue
—_—_——_——-_— nipalensis hey eee cw
-———.—___-——. obscurus ee 667, 672
——_——_-——_ pallipes a bee Ado
——_——_- ——— pileatus ae 660, 661
—_——_——_——__—— priam ... wey Seer 295
—__——_-——— _ pyrrhus re eh eOOT,
—_-_-— schistaceus.... 477, 478
——_——- senex ... ieepdet eta 00
Senecio johnstoni ee Poe seo 1439
Serinus pusillus Ss aap Boe en ed
Setaria hirsuta ... oe Ee an £045
Siganus javus.... such td sakes ada oval
——_——— —— Virgatus... as wr LO
Sillago sihama ais ae ee LOU.
Simia aygula on bes en 07544
——- entellus nee vee .. 484
Jona sac, oo a ee eS
Simia maura BAC ons S50 see
Siphia parva parva ai
Sitta europea kashmiriensis .., 5c
Sparus berda
Sphazerodema rusticum
Spilornis cheela
Spiranthes australis
Stachyridopsis pyrrhops
Sterna melanogaster
~——— seelia
Stipa arguens
——- paleacea BoD M3
Streptopelia decaocta decaocta
-——--~ senegalensis cam bayensis
Strix aluco biddulphi
— nivicola
-——- indranee indranee
Struthio camelus es ae aod
Sturnus vulgaris humii mas
~-——~-—-- nobilior ae
——_—-————-—- poltaratzskyi. . 726,
——-— - porphyronotus ;
Suya crinigera striatula : dos
Sylvia althea me ae eat 607,
— curruca affinis ... ant 607,
—— -- minula...
Syrrhaptes tibetanus
Tadorna cornuta Bae
——_—— tadorna uae ae 799,
Tamiops macclellandi...
Taphozous longimanus ... ae 546,
Tarsiger chryszeus whistleri
Tephrodornis pondicerianus
Terias hecabe 66
Terpsiphone paradisi ... 500 wes
Tetraceros quadricornis ay
Tetraogallushimalayanus ... 510,
——-- tibetanus es eas
——-——- tibetanus
Tetrarhynchus unionifactor
Themeda australis
= ciliata s
——-—— cymbaria .. soe 627,
—- —_—_—-— forskallii oe Mes 628,
—--— imberbis
—— polygamna
—— -— quadrivalvis ... aa 627,
-_—— —-- tremula re a Boo
- —-— triandra wn O27 SOZSs2 020m
Therapon jarbua a aes
‘Tichodroma muraria
Tinnunculus chenchris ...
Tomeutes phayrei phayrei
————- pygerythrus ...
Torgos calvus
Tringa glareola ...
- hypoleuca
-——--- ochropus
- totanus
Tristachya leucothrix
Tupaia belangeri belangeri
- —_—_——-—- Siccata
- ——-—-— tenaster
Turdoides somervillei
Turdus atrogularis
- viscivorus
Turtur communis
senegalensis
Upupa epops
- ——-- epops
Urginea polyantha
Urochloa ‘
- cimicina
————-- helopus
———-—-- marathensis
- panicoides
————- pubescens
INDEX OF SPECIES
PAGE
563,
558
546
547
728
SS
759
759
dey)
628
546
546
546
599
749
749
563
563
658
' 756
735
642
635
644
644
644
644
Urochloa reptans
—— —— setigera
-—— trichopus
Uroloncha malabarica ..
Uromastix microlepis ...
Vanda pulchella
Vanellus vanellus
Vanessa cardui
Varands griseus ... ‘
Vetiveria arundinacea ...
lawsoni
inuricata
——-— odorata
——--——- zizanioides
Viscum album
Viverra zibetha
- ———-- picta
—- —— —- prilinosa
Viverricula malaccensis
Vulpes alopex
- bengalensis
- ferrilatus ...
Zenarchopterus dispar ...
Zeuxine blatteri ... Ss
-- longilabris se =
Zosterops palpebrosa occidentalis
XXX
PAGE
643
644
645
752
608
520
759
811
608
409
409
409
409
409
701
546, 778
546
546
778
715, 716
715
716
705
§22
§22
759
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_ JOURNAL |
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PPA ee ah eens dower eaee oes aul ne ss owsicinioetuancleAuaienigasre nasi on siaswegners islstealtaisicrese
Tur LANGURS, OR LEAF MONKEYS, OF BrRiTIsH INDIA (With one
coloured and one black and white plates and twenty-one text-figures.)
By Re Te POCGCK, B.RiS: iti s.c tes csccstcereeeere ececsstraseds tstncnaces costes cei Sls
A CoLLECTING TRIP TO LADAK. PartI. (With a map and five plates.)
Vee Saale A ECLCOMME cas avretoes assem tater sbiancbtetn «Gn. ekiasewh emeee aa nacsestone tenses
A List OF ORCHIDS, WITH SOME NEW SPECIES FROM THE HIGH WAvyY
MounTAINS (MapurRA District). (With a plate.) By E. Blatter,
Seren PD AIG S 5 -obre petra «00d 0 Se'oGla'e cio nest wisiasslelohisinn's eSivis's vis oissets vas «now shiviaws "eos
AN ACCOUNT OF THE PEARL FISHERIES OF TUTICORIN, MARCH AND
Apri. 1927.2 (With four-plaes.) By Pi. KR. Awati, 1.2.8. ...:.08..
MODERN MusEuM METHODS. Part I. (With six plates and five text-
MDEULESE ant Oh abl eA PLALET 8 GC MeZiG.© a.couceniseshaesiesene ants PO oe ne
BoMBAY NATURAL HISTORY SOCIETY’S MAMMAL SURVEY OF INDIA,
BURMA AND CEYLON. Report No. 46. By T. B. Fry
Ceoercorersevreceses
A PRELIMINARY REPORT ON THE ANATOMY AND LIFE HISTORY OF
THE COMMON EDIBLE BACKWATER OYSTER, Ostrea madrasensis.
(Hith lwo plates.) -By S: 7. MOSES, M.A., F.Z.S., F.R.AsTuis. cesecses ss aes
ORNITHOLOGY IN THE BIBLE. By the Late Lt.-Col. C. E. Luard
eee eee ee eee eee ee ee ee ee ee ee ere er ey
FOC mM Peo tA OS OSOOHHT EH SESH TOE OO DETHS GOL SEH HEE Be DF EH OOHHAE LEH HHT OEE OOH EEO SEHBAC OL OOD
Ceo eee eee eeer FF OOH ors HEE FH FOr FEF Bree eFTs Boe SH EFF HHBH ooo TEH HOTEL EET OOTES EHO EE Os OOEEDE
MISCELLANEOUS NOTES :—
I.—Readers’ views on ‘Game Preservation in India.’ By W M.
Logan Home ...
rn oe ee ee oe ee ee ee ee 2 2
Il.—A Unique Lion Photograph. (Wz7th a plate.) By Salim
Aeeeut
COOCECOEET SOS THS HOHE ST HOOSHEHHOHHHHLG: SHLOHHESs CHL TZ FHEHHHTHHETMHTHHOHOH HO SB ES OED
II].—White Tigers. (Witha photograph.) By F.B. Robinson ......
IV.—A Tussle between Tiger and Buffalo Bait. By J. Mc C.
Clive
oe eeere 9 eee 246 ee veese eee cect eeeerseee Boe ee se - Cee r1 ee HT he ee BO BEES geeeer
450
460
586
il CONTIN TS “OF Vol. DXTT No.
PAGE
V.—Unusual Behaviour of a Tigress with Live Bait. (Wzth a
plater = “By FE AO CAvic ee seer eres ctniwn rs 20s celae see eseieo mares 587
VI.—What is the Record Panther ? By W. Me Logan Home ......... 587
VILA Panther and Flashiiont. = Cy77easplaie.) By, BY Also:
Bruriskalil i os Ue ke ccae vas eae eee eee eee me eeece tenia ae tems eee inareaine 588
VIII.—Boldness of Panthers, By W.M. Logan Home: (00.00... see coe en: 588
IX.—Circumstantial Account of a Panther attacked by Wild Dogs.
By JoweiC Cline. i. Gass. -eees Oe iia Ned Ren oe othe es ce ciple 590
Xe—Wild’ Dogs; . ByJ. Go launthonper sc ances tess brary se 590
XI.—How Wild Dogs kill their Prey. By A. A. Dunbar Brander ... 591
XII.—The Record Four-horned Antelope (7etraceros guadricornis).
(With a photo. \ = By Cy McCan te iawacncase een senes acne ees Se
XIJJ.—Abnormal Blackbuck and a Good Chinkara Head from Bikanir.
(Withia plate.) “By Surajimal sim elgaer ne een eae 593
XIV.—-Clicking Noise made by Muntjac.. By Charles Theobold,
BiZsSe' seed Sivaae abs sudeaeee Seat se a tieen satetias tee eeine te see ea eats 593
XV.— The Bray of the Chital Stag. By W. M. Lie Home Sentero E
XVi.—Musth in‘Hlephant By NUM BOts iia tates ee eee tence nes 594
XVIL.—A Large Tusker Elephant: By A. We Jon. 2.) 22 Gisescsceestns tee 596
XVIII.—A Good Nilgiri Tahr (Hemztragus hylocrius). Ry A. W. John.. 597
XIX.—Birth of Freak Calf at Indore. (Witha photo.) By C. E. D.
1) eth oy Ae AE ME ater Oy Rae Ronin ear iA RA caren tuna RI Re cha 597
XX. —An Example of an Assisted Passage. By Hugh Whistler,
BST 5-0 oZaScce eek oesey ems tone Cena a eee ee ee ee ee epee 598
XXI.—Extraordinary Pugnacity of the Rusty-Cheeked Scimitar
sabbler (Pomatorhinus erythrogenys). By Arthur H. Berriff 599
XXIL.—Oceurrence of the: Wood Snipe im, the Central Provinces:
By J Me GC Clive ee eet ka ole: eee re tee i ey Sa ele ae 600
XXIII.—Notes on Duck Shooting in the Roorkee District,-U. P., in the
years 1903 to 1927. (With a graph.) By A. H. Cun-
MNO HAM | 1. eee aedekeg foares oeque cede as ater oe eee ek sere ener aoe ee eee 600
XX1IV.—Snipe and Woodcock in S. India. By E.G. Phythian Adams.. 506
XXV.—A Correction to Mr. B. B. Osmaston’s ‘ Notes on the Birds of
Kashmir’) “By Huoh Winistler =a .s). cet eeeceee: eeentae eeeeece 607
XX ViE—An Albino Turtle. By he A lB’ A breat, IZ {Seen eee 608
XXVII.—Notes on the Desert Monitor (Varanus griseus) and the Spiny
Tailed Lizard (Uvromastix microlepis). By Norman
eee Corkille rtersc.. sce eeee saa @anls cag satu detoae Bede raouiamrenie s smsementaseet ess 608
XXVIII.—Hamadryads in the Kumaon Terai. By W.M. Logan Home. 610
XXIX.—Mr. Slater’s Death by Snake Bite. By Charles Theobold ......... 611
XXX.—A Note on the Green Whip Snake (Dryophis mycterizans) and
VYoune.. | (W227i a prolos) By. Ce Me Cangas ere, seta ate eee 612
XXXI.—A Record Cubbany Masheer (With a plate.) By G. he Hare. 3613
XXXII.—Aquatic Rhynchota in the Collection of the Agricultural
College,. Coimbatore, 8. India. .By Cedric Dover :)........5.5.. 614
PROCEEDINGS | sic g cncte cece ace tnouistcte ei tette ete Ace tell ae ete oe en ee eae pa ccceitaie’ cist ean ee 616
ee <r
te ‘azIS "YEN 6/ ‘azIs "FEN 6,
os4iA saprodouyquy Ipdojf1] Sn4u8 siusojesay|
ANVaD SlISssiOWSsd Sahl SANVYQ NOWWOD N&YSlsSvya AHL
“O00 “LSIP “LV AT AVENOGT ©
JOURNAL
OF THE
Bombay Natural History Society
JANUARY, 1928 VoL. XXXII No. 3
THE GAME BIRDS OF THE INDIAN EMPIRE
BY
BeGr SLUART BAKER, BZ.S:, F.L:S., M.B.O.U.,.H.F.A.O.U.
VoL. V
THE WADERS AND OTHER SEMI-SPORTING BIRDS
PART V
(With a coloured plate)
(Continued from page 245 of this Volume)
SUB-ORDER BALEARIC#
FAMILY BALEARICID
CRANES
The True Cranes are birds of large size with long necks and legs
but with a bill only equal to, or but little longer than, the head. The
nostril is enclosed by a membrane on the posterior side and it is
placed in the base of a groove which extends about half-way up the
upper mandible. There are 12 tail feathers and 11 primaries. The
tracheze pass into a hollow space between the long walls of the
sternal keel and are more or iess convoluted.
Our Indian Cranes were divided by Sharpe into the genera Grus,
Sarcogeranus, Antigone and Anthropoides.. ‘The name Grus is not
tenable and has to give way to Megalornis, whilst the differences
between this and Sarscogeranus seem hardly generic. On the other
hand Antigone, not recognized by Blandford, is now generally accept-
ed as sufficiently defined to form a good genus.
KEY To GENFRA
A. Crown of head bare; sides of head and
upper neck feathered By, ... Megalornis.
B. Crown of head bare, sides of head and
neck not feathered te . Antigone.
- C:; Crown of head feathered _... ... Anthropordes,
$98: JOURNAL, BOMBAY NATORAL HIST, SOCIETY, Vol secre
Genus—ANTHROPOIDES
Anthyropoides. —Vieill., Analyse, p. 59, 1816.
Type.—Ardea virgo Linn.
In this genus the head is feathered throughout and there are long
white plumes springing from behind the ear-coverts ; the feathers
of the lower neck are long and lanceolate, whilst the inner second-
aries are much lengthened but not disintegrated as in Grvus. The
bill and legs are shorter in proportion than they are in that
genus.
ANTHROPOIDES VIRGO
| The Demotselle Crane.
ortente. Restricted to India).
Anthropoides virgo. Blanf. and Oates, iv, p. 190.
Vernacular names.—Karkarra (Hind.); Ghants (Nepal); Karkuchi
(Mahr.) ; Kallam (Deccan); Garara (Uriya); Wada-Koraka (Tel.);
Kar-koncha (Can.) Ses Ont aie |
Description.—Forecrown tonape grey ; patch below the eye, a line
over the ear-coverts and long aigrette behind them pure white ;
remainder of head and neck black, the feathers of the lower fore-
neck very long and lanceolate falling over the breast; winglet,
greater coverts and wing quills blackish; ends of lengthened inner
secondary plumes black, grading into the pale French grey of the
rest of the plumage; the grey of the upper plumage extending on to
the base of the hind neck.
Colours of soft parts.—Iris red-brown (possibly young birds) to
crimson or red; bill sage-greenish with a red tip; legs and feet
black. ; fee ia
Measurements.—Wing 450 to 530 m.; tail 165 to 182 m.; tarsus
170 to 187m. ; culmen 63.10.70 a.
-- Young birds are like the adult, but have the head wholly grey ;
‘the black of the neck is browner behind, more grey in front, and
there are no lengthened plumes ; wings coloured as in the adult but
the inner secondaries but little lengthened.
Distribution.—Breeding in Southern Europe and the high plateaus
of Algeria; Central and Northern Asia as far East as Western
Mongolia. In Winter south to Northern Africa, Mesopotamia,
Palestine and India. Inthe last-mentioned country it has occurred
south as far as Kollegal in Coimbatore, but is rare south of the
Deccan. East it extends through Eastern Bengal and Assam to the
greater part of Burma, but it is not common east of Bihar.
' ~WVidttication.—No better description of the breeding of these
beautiful cranes exists than that given by Dr. A. S. Cullen in his
account of the Nesting of Birds in Bulgaria, though it is to be feared
that the birds no longer exist in such great numbers.
‘‘ This-bird arrives in the Dobrudsha in about the second week
of April.- It makes its appearance in flocks of from twenty to
thirty birds and upwards, and the form of these flocks very
much resembles: those of the Common Crane, and they almost
always fly very high, especially when passing wooded country;
THE GAME BIRDS OF THE INDIAN EMPIRE 399
but when crossing the plains they fly lower, and occasionally
come within gunshot. For the first week after their arrival
flocks of these birds mingle with those of the Common Crane,
and resort to any fields of newly-sown grain that they can
find, and to these they often do great damage. Regularly
once or twice during the day they repair to some lake or pond
ot fresh water to drink. Inthe evening the fields are desert-
ed, all the birds going to the nearest lakes or marshes, where,
I believe, they remain a]l night. About the second week after
their arrival the flocks break up into pairs, which disperse
themselves all-over the country. Very soon after the birds
have paired they begin to lay. Eggs may be found from
about the third week in April to very nearly the end of.May ;
but much depends onthe state of the weather. The nest of
the Demoiselle Crane is, without exception, made on the
..ground, usually amidst some kind of young grain, but. often
amongst grass on fallow land, and now and then, though more
rarely, amongst stubble. The nest—if indeed ‘such it_can be
- called—is made by the birds pulling up or treading down the
grain, grass, or stubble for the space of about two feet, and
scratching the shallowest possible hollow in the middle of-the
-bare patch thus formed. A few small straws and stones are
often found in and about this hollow, but whether they are
brought there by the birds, or found there by them, which _ Is
very probable, I cannot say with any certainty.’
‘“‘T believe myself, however, that they are placed there by the
birds. The eggs, which are never more nor less than. two. in
number, are a/ways placed side by side, in, the hollow already
described, with their small ends pointing in the same direction.
The male assists the female in hatching the eggs; indeed, I
-have every reason to think that he sits as much as the female.
To the uninitiated in the habits of this bird, its eggs are very
difficult to find. When an intruder approaches the spot where
the eggs lie, he at first sees nothing except perhaps a solitary
bird Standing on one leg as if sleeping. Should he not be a
very good observer, as he draws nearer he will probably only
notice a pair of birds walking rapidiy away, plucking the grass
as they go, apparently feeding eagerly, and he will most likely
account for the sudden appearance of the second bird by con-
cluding that it had been sitting on the ground resting, and will
not think anything about the matter, and pass on. Very few
persons indeed would at all suspect that the birds had a nest,
Often, nay, almost always, when the birds have placed some
distance between them and their nest, they will take flight, and
to all appearance desert the spot altogether, but no sooner is
the intruder’s back turned, than there they are again in a
wonderfully short space of time. One of the birds, I believe
the male, always keeps watch over the other, except in the
middle of the day in very hot weather, when the bird, which
acts as sentinel, deserts its post and goes to the nearest pond
_ or lake to drink. It does not, however, remain away very
long.’’
400 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
‘¢ Should the bird whose place it is to watch while the other sits
be absent, the sitting bird when disturbed is not quite so guarded
in its movements, and will allow a much nearer approach.
When the eggs are first laid the birds will leave them as soon as
an intruder comes in sight, but as the incubation advances they
become Jess shy, and will not leave their eggs so readily.
‘¢T cannot say whether the bird which watches stands close to
ora little distance fromthenest. I rather incline to the latter
opinion. Ifacrane be observed chasing other birds away from
any particular spot, it isa sure sign that it has a nest not far
off. This bird will give chase to eagles and great bustards
without the ieast fear, should they venture near its nest. For
all kinds of harriers and seagulls it has a great dislike. I
have been told by the Tartars that, should a dog by chance go
near the nest of this bird, both the birds will attack him, strik-
ing him with their beaks and wings, and making a great noise
all the time. I myself have never had an opportunity of
witnessing such an interesting encounter, but I am certain
that this bird has sufficient courage for one. The Demoiselle
crane is easily domesticated when young.”
Dybowski says that in Dauria ‘‘it nests onthe rocky banks of
rivers andrarely on bare mountains. The nest is made of small
stones fitting close to each other, the surface of the nest is flat or
deepening somewhat towards the centre; it chooses sometimes a
place which is a few inches higher than the surrounding ground,
and fills up all the crevices and openings with stones.”’
The breeding season lasts from the middle of May to the middle
of July, a month earlier in its most southern breeding haunts, even
later in the more northern. ‘The normal clutch of eggs is two only,
but three are occasionally laid. In appearance generally they are
small replicas of those of the Common Crane, but they are still
longer and more narrow in proportion to their size. The ground
colour varies from a pale olive yellow to a warm olive brown or
olive green whilst the markings consists of large primary blotches of
brown or purple brown and secondary blotches of neutral tint of pale
purplish-lavender. In most eggs these are fairly numerous over the
whole surface, but insome they are confined to the larger end where
they form an indefinite ring or cap. One hundred and twenty eggs
average 83°3 & 53°1 mm., maxima 91°4 & 55:1 and 84:2 & 56°6 mm. ;
minima 74:1 & 48°5 and 78:0 & 47:0 mm.
Habits.--The Demoiselle Crane arrives in India in October and
the first flocks which arrive seem to fly straight down to their
southern limits as it is said to arrive in the Deccan in the middle of
October, whilst in the north few birds seem to settle until the end
of that month, though they must pass over on their way south.
They return North in March and April, Whistler recording numerous
flocks from the Jhang District between March 8 and April 7. Both
coming and going they assemble in vast multitudes, and a most
interesting letter from Mr. A. A. Phillips, quoted by Whistler,
gives a graphic account of one of these assemblies :
‘On March 25, C. and I went out after them. They were a
sight worth seeing and hearing. There seemed ‘to be vast
THE GAME BIRDS OF THE INDIAN EMPIRE 401
divisions of them about, but we only shzkaved one division:
while they were feeding on the ground there was a broad band
of them for about 14 miles. When alarmed they bunched to-
gether and looked just like a white pebble beach about 100 by
500 yards in extent, and when they rose the noise was like the
roaring of the sea. We shot three which were all Demoiselles.”’
The dancing of this Crane is well known, Nordmann’s description
having been translated by Bree, as follows :—
‘« They arrive in the South of Russia about the beginning of
March, in flocks of between two and three hundred individuals.
Arrived at the end of their journey, the flock keeps together
for some time ; and even when they have dispersed in couples,
they reassemble every morning and evening, preferring in
calm weather to exercise themselves together, and amuse
themselves by dancing. For this purpose they choose a con-
venient place, generally the flatshore of a stream. There they
place themselves in a line, or in many rows, and begin their
games and extraordinary dances, which are not a little surpri-
sing to the spectator, and of which the account would be con-
sidered fabulous were it not attested by men worthy of belief.
They dance and jump around each other, bowing in a burlesque
manner, advancing their necks, raising the feathers of the
neck-tufts, and half unfolding the wings. In the meantime
another set are disputing ina race the prize for swiftness.
Arrived at the winning-post they turn back, and walk slowly,
and with gravity; all the rest of the company saluting them
with reiterated cries, inclinations of the head, and other
demonstrations, which are reciprocated. After having done
this for some time, they all rise in the air, where, slowly
sailing, they describe circles, like the swan and other cranes.
After some weeks these assemblies cease, and from that
time they are constantly seen walking in loving pairs
together.’’
In flight and voice these cranes are very similar to the Common
Crane ; they migrate in broad V shapes or in long waving lines, and
their call is a beautiful sonorous trumpet, somewhat less loud per-
haps than that of Wegalornis grus. Vidal says that they are very
slow rising off the ground, and considers the best way to get a shot
at a flock is to walk straight up to it without concealment, ‘‘ and
immediately the first bird flaps his wings preparatory to taking
flight, to run as fast as ever you can, straight at them. If you
have any breath left in you, after a short spurt of 60 or 70 yards you
are almost certain of a shot at the fag-end of the flock within killing
distance.”’
Most people, however, who shoot cranes do so from behind the
cover of a cart or of a led bullock, or if there be high crops about,
by stalking them under cover of these. They are extremely wary,
clever birds and, whether it is desired to shoot or trap them, they
are very difficult to bring to bag. They are, however, sometimes
caught in nets and, less often, in nooses, and when once caught
soon become affectionate pets, but they are very jealous of other
pets and jealous and irritable with small children.
402 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
As an article for the table the Demoiselle ranks very high, some
people considering it the best of all Indian Game-Birds. They feed
almost entirely on shoots, seeds and young crops, but also eat
insects, small reptiles, etc.
Genus.—MEGALORNIS
Megalornis.—Gray, List Gen. Birds, p. 85, 1841.
Type. :
In the genus Megalornis the crown of the head is bare but the
sides of the head and neck are feathered and not bare as in Antigone.
The wings are long with the inner secondaries lengthened and
considerably exceeding the primaries ; tail short ; the tibia naked on
the lower half ; toes short and strong with short, blunt claws. The
sexes are Aike in plumage.
Cranes are almost cosmopolitan in range, three genera and five
species being found in India.
Kky TO SPECIES
A. Plumage grey, tail grey.
(a) Throat grey ; white on neck confined to
a ime ccowi thieysides, 2: M. grus.
(6) Throat white ; greater part of sides and
backtot neck white Ue .. MM. monachus.
B. -Plumage grey, tail black ~..: i” M, nigvicolits.
C. Plumage white ... a ... MM. leucogeranus.
MEGALORNIS GRUS
Ardea grus Linn., Syst. Nat., 10th ed., 1, p..153.
Type locality.—Sweden.
The typical form differs from our Indian bird in its darker plum-
age more especially in the colour of the inner ornamental] seconda-
ries.
MEGALORNIS GRUS LILFORDI
The Eastern Common Crane
Grus lilfordi Sharpe, Cat. B. M., xxiii, p. 252, 1894 (BE. Siberia). .
Grus communis. Blant. and Oates, IVe 8p. eG.
Vernacular names.—Kurunch, Ki urch (Haind.) ; Awn7 Sind)\s 70 ase
(Deccan) ; Kelangi (Tel.).
Description. -—Lores and crown to nape bare black stan with scat-
tered black hairs; a broad band of bare skin brick-red to red,
across the nape, ae black hairs even more scarce than on the crown ;
a patch of grey feathers behind the bare red spaces; sides of face
grey ; ear-coverts, sides of head and neck white running back behind
the grey neck patch and down the hind neck; winglet, primaries,
the terminal portions of the primary coverts and the tips of the long
secondaries black; chin, throat and foreneck dark siaty brown;
remainder of plumage pale, almost silvery grey ; tail slightly darker
grey, blackish towards the tip. - .
Colours of soft pavis.—Iris orange-red to red-brown .or crimson ;
bill dull glaucous or dingy green, paler and more yellow towards
THE GAME BIRDS OF THE INDIAN EMPIRE 403
the tip; legs and feet black, the soles paler, more brown, greenish
- or even fleshy. —
~ Measurements—Wing 533 to 608 mm.; tail 192 to 205 mm. ; tarsus
225 to 258 mm. ; culmen 103 to 118 mm.
Young birds have the edges of the grey feathers isabelline or
rufescent isabelline ; the sides of the head and neck and hind neck
pale rusty rufous; feathers cover the red bare skin on the nape
whilst the crown also is more or less covered with the same; the
drooping inner secondaries are wanting.
Halt-grown birds have the edging to the feathers more rufous and
more conspicuous and the whole crown and neck clothed with rusty
coloured feathers.
Chicks in down are pale golden fulvous above: a broad line of deeper
tint runs down the centre of the back and two narrower, less defined
lines from the wings down each side of the body; centre of crown
and a mark above the eye dark golden rufous; below sandy buff,
albescent on the chin and foreneck and pale on the centre of the
abdomen. |
Distribution.—The Eastern Common Crane breeds in Eastern
Siberia from the Yenesi basin eastwards and in Turkestan. In winter
south to China and practically the whole of Northern India. It is
extremely common in North-west India and extends south through
the greater part of the Bombay Presidency and the Deccan. It has
occurred in Southern Orissa and I have seen it as far East as
Lakhimpur and Cachar in Assam on several occasions.
Nidtitication.—There is very little that I can find on record in
regard to the breeding of the Eastern form of the Common Crane
H. E. Dresser had eggs which he received from the Russian
collectors round about Tomsk which is considerably to the West of
the Yenesi. Of these he merely says that they differ in no way
from the European bird and that their breeding habits are similar.
I have also two pairs of eggs in my collection collected by
Smirnoff north of the town of Yenesi and on the east of that river.
A brief note which accompanies the eggs says: ‘‘ These birds breed
in small communities of half a dozen or so, the nests being placed
on the ground from 50 yards to half a mile apart, sometimes two
quite close together. The nest is a big untidy one made of all kinds
of things which the birds pick up from the ground such as rushes,
moss and lichen and otner scraps. It is sometimes quite high, as
much as a foot whilst across it sometimes measures over a metre.
The eggs always number 2.’’
The sort of country it selects as a breeding site is nearly always
such as contains swamps and lakes and it is said by Burturlin to
sometimes breed on the extreme edge of the lakes in among the
rushes and reeds, probably however, more often it breeds on
marshy ground some little distance from the lakes themselves.
_ The only eggs I have seen of this race have all been laid in May
or early June but they are said to breed sometimes as late as the
last week of the latter month. The eggs could not possibly be
distinguished from those of the Common European Crane. In
shape they are rather long drawn-out ovals, the smaller end blunted
although distinctly smaller than the other end. The texture is
404 JOURNAL, BOMBAY NATURAL ATST. SOCIETY, Vol. XXXII
rather coarse but the surface is quite smooth though with little, if
any, gloss and the shell is stout though not particularly so for such |
large eggs. The ground colour varies froma pale greenish-grey
or yellowish-grey to olive or yellow brown and the whole surface
is sparingly marked with reddish-brown streaks and blotches with
other secondary markings of paler tints of the same and dull
lavender grey. Occasionally an egg is found with rather darker
browner markings.
Six eggs average 93°38 K-59:6 m:,, maxima, 95-2 x 95:0am.-
minima 90:2 « 57:0 m,
Burturlin gives the breeding distribution of the eastern form of
the Common Crane as Altai, Turkistan, Dauria to Manchuria and
north from Siberia to the 66th degree of latitude.
Habits.—In India, the Common Crane is only known to us as a
winter migrant, arriving generally about the second or third week in
October; Hume records their arrival on one occasion as early as
October 3, whilst Doig says that in Sind, many birds arrive as
early as the end of August, though this must be very exceptional.
They begin to leave again by the middle of March, the bulk of the
birds moving about the end of that month anda few staying on
until the end of April. These late birds Hume thought were young
birds not anxious to breed and therefore not leaving as early as the
old ones. In the east of India the few Cranes that are to be seen
arrive in November and I do not think that I have ever seen them
after the second week in March. In North-west India they are
often found in very great numbers said sometimes to be in many
hundreds.
The Crane is undoubtedly one of our most wary game birds and
one of the most difficult to approach within gun shot.
Colonel Tickell refers to this bird and gives a most interesting
account of their habits which is quoted by Blyth in his monograph
on the Cranes. He says :— :
‘In serene weather in the cold season, when, though the skys
are cloudless blue, the mild atmosphere enables the sportsman
to roam aJi day long over miles of far-reaching plains, the
sonorous trumpet cail of the Crane is one of the most familiar
sounds that meet the ear. The cry resembles the syllable
‘kurrk’ a single loud brassy note, harsh and metallic, but
mellowed by the vast distance at which it is generally heard.
At such times the birds may be seen high in the air, advancing
in an angle or ina single diagonal line winnowing their way
across the pathless vault to distant regions, or soaring round
and round in irregular swarms, and independently, as if re-
connoitring the country below to select a fit place for their
nocturnal meal. When thus employed sometimes two or
more parties meet, and the clamour amongst them then
becomes extreme, like the mutual clacking and cackling of
assembled geese. Should the country, as beforesaid, be quite.
open, and no human being be observed moving on the plain,
they will sometimes descend to the ground in the day-time,
and on such occasions the downward sweep, from a height of
perhaps a mile in the air to the face of the earth, of two or
THE GAME BIRDS OF THE INDIAN EMPIRE 405
three hundreds of these large birds, is a majestic spectacle,
though of necessity only to be seen from afar.”’
‘© T once, indeed, met with a striking exception to this wary
habit of the Crane in keeping, during the day, out of gunshot
inthe air. In January, 1845, as I was one day riding along the
Trunk road (as the old Government road from Calcutta to
Delhi is called), in the district of Manbhoom, near the village
or Chowkee of Niamutpoor, I came in sight and within point
blank bail shot of a host of these birds collected in a rice
stubble field. There was a prodigious concourse of them
amounting probably to a thousand. They were making a
deafening clamour and seemed in great agitation, while every
now and then a party of them would take wing and fly off,
generally to the eastward, towards the Damoodur River, but
many went in other directions. The country in the vicinity
was open and wild, with bushes scattered about, and here and
there a cultivated patch; the hour early in the morning, by
sunrise they had all disappeared. I was told by the villagers
that as night fell the birds would return and re-assemble in
the same spot, and that they had been in the habit of doing so
for many years. I confess I was never more astonished at
anything within the circle of Ornithology than at this instance
of exception to the wild and wary habits of the Crane. They
could not have assembled in the field (which was not a large
one)to feed, asthe grain in the stubble would not have
afforded a night’s meal to a tenth part of the flock. The most
probable conjecture is that they fed elsewhere during the
night, as is the wont of the bird and repaired to that particu-
lar spot to rest till dawn, congregating in separate and
independent parties. But why they should have collected
night after night and have stopped so near a village, and
so liable to approach, is inconceivable. The people of the
vicinity are, it must be noted, no shikaries and the report of
a gun was probably a sound almost unknown in that part of
the district. The Crane, like most waterfowl, feeds at night,
more as amatter of precaution than by choice. No such
nocturnal habits are perceived in tame individuals, nor, is it
probable, in the wild steppes of Tibet or Siberia, where these
birds breed unmolested and roam about free from the sight
even of a human being.”’
‘¢ Tn India, after filling their crops with the gleanings from
the rice and vetch stubbles, the Cranes retire at or soon after
sunrise to sand flats or islets in large rivers, or to open
downs inland, where no coverts afford means of approach to
an enemy. And there they may be seen through a telescope,
preening themselves, lazily stretching forth their legs or wings,
or sitting dozing on their heels. One or two, however, keep
on the alert, their heads raised high; and these, to judge by
their size and flowing plumes, are old males.’’
As we have already seen the Crane is a magnificent flier, one
with great power and endurance and also capable of carrying out
both graceful and intricate evolutions at pleasure. It is also a fine
2
406 JOURNAL, BOMBAY. NATURAL ‘AIST. SOCLETY, Vol. XXXII
runner and a wounded bird will often give its would-be captors a
long chase before success crowns their efforts, indeed on marshy
ground a second shot is generally necessary to bring to bag a bird
that is merely winged. He is also a good swimmer but takes to
water of swimming depth only under pressure, though they are
very fond of wading about in shallow water when there is a supply
of food to tempt them. The Common Crane, like all cranes, when
only wounded, will put upa very fierce resistance to capture,
striking viciously with the bill but, as Hume points out, a very
slight tap with a stick on its long slender neck is quite sufficient
to place it hors de combat. Hume gives a remarkable example
of the force with which a Crane shot dead falls to the ground
He writes: ‘‘ As we walked back to the boat we became aware
that an enormous flock of Cranes that had been trumpeting all
the while, were practically over our heads. I wanted to unload
my big gtn and so fired at random straight up. I had hardly
recovered from the recoil when a peculiar rushing sound caught
my . ear, (luckily the sound ‘outsped-~ ‘the. bird),, I sjust < started
back to listen when with a ‘scrush’ a dead bird plopped on the
ground at my feet. The Crane had fallen back downwards, the
shoulders first striking the ground and was firmly embedded in the
clay. Wepulled it out and found a_ basin-like depression in the
ground, as faras I could judge inthe dark, fully 8 inches deep in
the deepest part.’’
Cranes feeding as they do so muchin cultivation, very soon get
used to the cultivators moving about in their small carts or
with their bullocks and frequently donot trouble to keep out of
gun-shot of these familiar objects. Accordingly, advantage is
taken of this both by Indian and European sportsmen who approach
under cover of bullock cart or bullock and thus get shots at birds
which it would be otherwise impossible to approach. At other
times birds may be driven towards the sportsmen bat to do this
with any success afew beaters only must be employed and
these must know their work well. For a line of beaters to
walk straight towards the birds is merely a signal for the latter to
rise ez masse and to take their departure high in the air and far out
of shot of the gunners. If the birds are to be persuaded to fly over
these latter within shot, the two or three beaters employed must
slowly and apparently casually work their way up towards the
birds so that these do not become really frightened, but gradually
move along inthe desired direction until finally they are within
about 100 yards of where tne sportsmen are hiding, when they may
be startled into flight. A flock, however, once thus moved and
shot at may be given up as hopeless for the rest of the day and if
more birds are wanted, the sportsmen must move on to fresh
ground.
Cranes are very omnivorous in their diet but probably on the
whole prefer a vegetarian meal toany other. Different shoots of
all kinds of crops, especially rice, are greedily eaten and in some
cases, when the flocks are numerous and large, they cause con-
siderable damage tocrops.. They are saidto be especially fond of
water melons and destroy even more of these than theyeat. In
THE GAME BIRDS OF THE INDIAN EMPIRE 407
addition to all kinds of aquatic plants, grain and shoots they will
eat any small reptile and all kinds of large insects and it has been
alleged that on occasions they even catch fish. They feed princi-
pally in the very early mornings and late evenings whilst, where
they are much disturbed, they feed a great deal during the night,
more especially when there is a moon. During the hot hours of
the day they resort to the banks of rivers and tanks where they
have a wide view all roundthem and there pass the time away
dozing and often standing on one leg in shallow water with their
head and neck tucked away into the feathers of the back. Directly
they flight towards their feeding grounds Cranes become very
noisy, constantly uttering aloud trumpet-like call but during the
day they seem to keep more or less silent.
Like all their family this bird indulges in dancing during the
breeding season and at odd moments even during the Winter
months, when the attitudes, bowings and scrapings indulged in
during these preformances are often very ludicrous.
A flight of these birds when moving for any distance is always
carried out inregular formation, either a long line or a wide‘ V’,
the leader at the apex apparently changing place every few minutes
with one of the other birds in the rear. When flying short dis-
tances from one feeding ground to another, this formation is not
always observed and the birds move in indiscriminate flock.
(Zo be continued)
REVISION OF
THE FLORA OF THE BOMBAY PRESIDENCY
BY
EK. BLATTER: §.J.40P0s, F.l.S.
PART V
GRAMINEAL
BY
E. BLATTER and C. McCann
(Continued from page 298 of this Volume.)
27. CLHISTACHNE, Benth. in Hook. Ic) Pl. xiv, 60,.¢.1379. Stapt ina.
Trop. Atr. ix. lod.
Tall, rather coarse grasses, annual according to Stapf, perennial according
to Hook. f. Leaves long, narrow, flat, with stout midribs. Panicles narrow,
more or less contracted, greyish or fulvously hairy. Spikelets solitary, all
alike, hermaphrodite, pedicelled on the tough rhachis of racemosely arranged
or panicled racemes, falling entire from the thickened tips of the pedicels.
Florets 2, lower reduced to an empty gliime, upper hermaphrodite. Involucral
glumes equal, very similar, with involute margins, more or less coriaceous,
delicately 7-9-nerved, muticous. Floral glumes hyaline, lower 2-nerved, upper
2-dentate or subentire, 3-nerved, with a twisted flexuous awn from the sinus or
tip ; pale very minute, ciliate. Lodicules 2, broad-cuneate, sparingly ciliate.
Stamens 3. Stigmas laterally exserted, plumose. Grain oblong to obovoid-
oblong’, very obtuse or truncate ; embryo half the length of the grain.
Species 3, one in West India and 2 in tropical Africa,
1. Cleistachne Stocksii, Hook. f. in F. B. I. vii, 163.
Descripiton : Stem tall, stout, simple. Leaves 3J-40 cm. by 12 mm., finely
acuminate, softly hairy, midrib stout, margins slightly thickened, ciliolate,
sheath terete, ligule oblong, coriaceous. Panicle 15-20 cm. long, long-
peduncled, sub-erect ; rhachis and branches sparsely ciliate, pedicles of spike-
lets strigose with bright yellow hairs. Spikelets 5mm. loug, crowded, dark
brown, callus short, bearded. Lower involucral glume dark brown, obscurely
many-uerved, hirsute, shining, upper like lower, but narrower nearly glabrous.
Lower floral glume 2-nerved, margins infolded, tip hispid, upper a twisted awn
16-25 mm. long, dilated at the base into a hyaline, entire, 3-nerved membrane,
embracing the minute, ovate, obtuse pale.
Locality : Tungar forest, Bassein (Bhide!).
Distribution : So far only found in Malabar on the Bababoodan Hills.
28. VETIVERIA, Thouars ex Virey in Journ. de Pharm. 1. ser., xiii, 499 ;
Stapfin Fl. Trop. Afr.ix, 156.
Coarse, perennial, glabrous grasses ; rhizomes stout ; culms stout, more or
less compressed below. Leaf-biades firm to hard, conduplicate in bud, then
flattening out, at least upwards, gradually passing into the sheath ; lower
sheaths much compressed, flabellate-imbricate. Panicles erect, long, of many-
rayed whorls of slender simple or rarely compound racemes, glabrous except
for the frequently bearded calli. Spikelets 2-nate, of each pair subsimilar,
differing in sex, one sessile, the other pedicelled, on the articulate fragile
rhachis of copiously whorled (rarely panicled) peduncled 3-to many-jointed
racemes, the sessile spikelets falling with the contiguous joint and the accom-
panying pedicelled spikelet or at least the accompanying pedicel; joints and
pedicels slender, stightly and gradually thickened upwards. Florets 2, lower
reduced to an empty glume, upper hermaphrodite in the sessile, male in the
pedicelled spikelets. Sessile spikelet laterally slightly compressed, awned or
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 409
awnless. Involucral glumes equal, lower more or less coriaceous or char-
taceous with a broad rounded back and subinflexed margins, usually muti-
cous, upper boat-shaped, keeled upwards, with broad hyaline ciliate margins,
muticous, mucronate or aristuiate. Floral glumes hyaline, of lower floret
2-nerved, of upper minutely 2-dentate, muticous or mucronulate or with a per-
fect or imperfect awn from the sinus. Pale minute, hyaline, nerveless. Lodi-
cules 2 glabrous. Stamens 3. Stigmas laterally exserted ; styles subterminal.
Grain oblong, slightly oblique at top. Pedicelled spikelet dorsally compressed ;
involucral glumes much thinner than in the sessile, like the floral glumes
usually awnless.
Species about 7 in the tropics of the Old World.
1. Leaves 5-13 cm. long. Panicle 15-18 cm. long ... 1. V. Lawsont.
2. Leaves 30-90 cm. long, Panicle up to over 30
cm. long aie on os: 2. VV. SISQnIOIdes.
1. Vetiveria Lawsoni, Blatter & McCann, mov. comb.—Andropogon Lawsont,
Hook; f. F.B.1. vis, 187.
Description: Rootstock stout, horizontal. Stem erect, simple, slender,
internodes very long. Leaves chiefly subradical, 5-13 cm. by 5mm., exactly
linear, rigid, curved, acute or obtuse, tips serrulate, base not contracted, mar-
gins ciliate, nerves 4-8, strong ; sheaths compressed, of lower very short, of
cauline very long, striate; ligule a ridge of hairs. Panicle 15-18cm. long,
narrow, elongate, branches or peduncles of spikes opposite and fascicled,
branchlets slender, puberulous with a white scurf. Spikes 6-12 mm. long, pale
reddish, erect ; joints 6-8, very obliquely truncate, tips obscurely ciliate, pedi-
cels nearly equalling the spikelet, slender, compressed. Sessile spikelets 4
mim. long, linear-lanceolate, callus bearded with silky hairs. Lower involucral
glume linear, rigid, coriaceous, tip obtuse, bristly, keels muricate, scaberulous
margins inflexed, upper involucral glume cymbiforin, tip 2-fid, awn longer
than the glume, base ciliate, keel pectinately ciliate above the middle. Lower
floral glume oblong, ciliate, nerveless, upper arched,‘ linear, obtusely 2-dentate,
awn very slender. Pale oblong, ciliate, nerveless. Anthers Jong. Pedicelled
spikelets male, longer and narrower than the sessile, callus naked ; lower in-
volucral glume 3-nerved, awned, keels pectinately ciliate, upper acuminate,
awned. Floral glumes oblong, obtuse, ciliate.
Locality: S. M. Country: Dharwar District, very common (Sedgwick
2170!) ; Dharwar (McCann A277 !).
2. Vetiveria zizanioides, Stapf in Kew Bull. (1906), 346-49, 362, in Fl. Trop.
Afr. ix, 157.--V. odorata, Virey in Journ. de Pharm. 1. ser. xiii, 499.—V. arundi-
nacea et muricata, Griseb., Fl. Brit., W. Ind. 559, 560.— Phalaris zizaniotdes,
Linn. Mant. Alt., 183.—Andropogon muricatus, Retz., Obs. iii, 43; Roxb. Fl.
ind. 1, 205° Gtah, Cat, Bonib, Pl. 238; Grif. Ic. Pl. As. t. 139) f.57, €..155,
f.1; Dalz. and Gibs. Bomb. Fl. 302; Duthie Grass. N. W. Ind. 90, Fodd.
Grass. N. Ind. 36, t. 24.—A. FHestucoides, J. S. Presl in C. B. Pres] Reliq.
Haenk. i, 340.—A. sguarrosus, Hack. (won Linn. f.) var. genuinus, Hack. in
Monogr. Androp. 542-44.—A. sguarrosus, Hook. f. (#om Linn. f.) in F. B. I,
vii, 186.—.4. sguarrosus, Cooke (mom Linn. f.) in Fl. Bomb. Pres. ii, 991.—
Agrostis verticillata, am. Ill. Gen. i, 162.—Anatherum muricatum, Beauv.
Agrost. Expl. Planch. 15.
J. D. Hooker and Cooke and many others have followed Hackel in calling
this plant Andropogon squarrosus, Linn, f. Stapf (in Kew Bull. 1906, 347) has
explained that this name applies to quite a different plant: ‘ No notice was
taken of Scheuchzer’s description or of Petiver’s and Du Bois’s specimens, and
when Linnzeus, about 1770,* received the grass from Koenig he described it
as something new under the name Phalarts zizantoides. Koenig, however, also
sent specimens of the grass to Retzius, who published it as dudropogon murt-
catus * in 1783. This name, which was suggested by Koenig himself, was
subsequently adopted by Roxburgh and most other botanists. More recently,®
however, it has been replaced by Andropogon sguarrosus, aname adopted by
the younger [Linnzeus * for a plant, also communicated by Koenig, who found it
1Linnzeus, Mant. Alt. (1771), 183.
Retz. Observ. iii (1783), 43.
3Hackel, Andropog. in DC. Monogr. Phaner. vi (1889), 542.
*Linn. f. Suppl. (1781), 433.
410 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
‘“cirea Zeylonam natans supra stagna profundiora,’’ and entirely distinct from
Andropogon muricatus. ‘The specimen is still in Linneus’ herbarium and was
correctly identified by R. Brown? with his Panicum abortivum, that is Chame
vaphis spinescens, a characteristic floating grass of the Indo-Malayan region.
Retzius? himself is responsible for the erroneous reduction of Andropogen
squarrosus to Andropogon muricatus, which recently has been revived, although
Roxburgh*® long ago drew attention to the confusion. ‘‘ Zzzanioides’’ being
the earliest specific epithet, it will have to be adopted for the ‘‘ Khas Khas,’’ so
that its name under Ve/zveria must be V. zizanioides.’
Vern. Names: Vala, Ushir, Valo, Bala, Khas Khas of the Anglo-Indians,
Description: Adensely tufted perennial grass. Rootstock branching with
spongy aromatic roots. Culms stout, up to over 1'8 m. high, usually sheathed
allalong. lLeaf-sheaths compressed, especially the lower which are sharply
keeled and fan-like, imbricate, very smooth, firm ; ligules reduced to ascarious
rim; blades narrowly linear, acute, 30-90 cm. long, 4:2-10°6 mm. wide, erect,
rigid, firm or somewhat spongy, usually glabrous, rarely more or less hairy
downwards on the face, pale green, midrib slender, lateral nerves close, 6 or
more on each side, rather stout, slightly prominent, margin spinously rough.
Panicle oblong, up to over 30 cm. long, usually contracted ; rhachis stout,
smooth ; whorls 6-10 with up to 20 rays; branches oblique to suberect, naked
for up to 5cm., filiform, slightly rough. Racemes up to 5 (rarely 7'5) cm. long,
very slender ; joints about as long as the sessile spikelets or sometimes distinctly
exceeding them, smooth or more or less rough, minutely and unequally ciliolate
at the slightly oblique tips; pedicels similar, but shorter. Sessile spikelet
linear-lanceolate to almost linear, acute or subacute, 4*2-4°8 mm. long, yellow-
ish, olive or violet-brown or purplish to almost black; callus obtuse, under
1mm. long, glabrous. Involucral glumes, acute, coriaceous, lower niuriculate
all over the back, 5-nerved, lateral nerves close, very fine; upper spinulously
muricate on the keel. Lower floral glume as long as the involucral glumes,
acute, reversedly ciliolate, upper up to 3°3 mm. long, narrow, oblong-lanceolate,
mucronulate, ciliate. Lodicules 2, quadrate and conspicuous, though sinall.
Styles and stigmas short. Stigmas purple. Anthers 2-3°3mm.long. Pedi-
celled spikelet sparingly aculeolate or almost smooth ; upper floral glume entire,
acute.
Locality: Gujarat: Road to Lasandra (Chibber!); Daman (Bhide)) ;
Ahmedabad, common in damp valleys (Sedgwick !).—Konkan : Ghatkoper,
Horse-shoe Valley (McCann 9957 !).—N. Kanara : Dandeli (Talbot 2209 !).
Cke. l.c. classes this species amongst non-indigenous plants. We are of
opinion that it is indigenous in most parts of the Presidency.
Distribution: Practically over the whole of India, and eastwards to Burma.
Occasionally cultivated. Lower Guinea in Tropical Africa. Throughout the
Malayan region only cultivated or as anescape. Introduced into the Mascare-
nes, the West Indies and Brazil.
Farly history and economic uses: See Stapf in Kew Bull., l.c.
29. ‘CHrySopoGon, Trin. Fund. Agrost. 187; Stapf in Fl. Trop. Afr.,
ix, S159;
Perennial (at least in the Old World). Leaf-blades narrow. Panicles
usually lax, of whorls of simple or basally divided filiform branches, rarely
the branches 2-nate or solitary. Spikelets in threes at the ends of the
branchlets of terminal panicles, one sessile, the other 2 pedicelled, the three
falling entire from the thickened, nearly always bearded, oblique tips of the
peduncles ; exceptionally 2-nate in 2-jointed racemes, one sessile, the other
pedicelied, each sessile spikelet falling with the contiguous joint and its
pedicelled companion, pedicels and joints, if present linear-filiform, never
longitudinally grooved or appendaged. Florets 2, lower reduced to an empty
glume, upper hermaphrodite in the sessile, male or neuter in the pedicelled
spikelet. Sessile spikelets usually laterally compressed, awned. Involucral
glumes subequal ; lower coriaceous or chartaceous, involute with a rounded’
*R. Brown Prodr. Fl. Nov. Holl. (1810), 193.
*Retz.,° l.c.,3v (1789), 21- .
°Roxburgh Fl. Ind. ed., Carey and Wall. I (1820), 270.
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 411
back or complicate and more or less keeled upwards, upper boat-shaped,
more or less keeled. Floral glumes hyaline, lower 2-nerved, upper linear,
entire or 2-dentate with a usually perfect awn from the sinus. Pale 0 or small,
hyaline, nerveless. Lodicules 2, small, glabrous. Stamens 3, Stigmas
exserted laterally low down. Grain linear, laterally compressed ; embryo half
the length of the grain ; scutellum linear-oblong. Pedicelled spikelet dorsally
compressed, awnless or aristulate.
Species about 18 in the hot parts of the Old World, only a few entering the
temperate zone. One in Florida and Cuba.
Cooke (ii, 984-986) describes 4 species of Andropogon belonging to the
section Chrysopogon : A. aciculatus, Retz., A. lancearius, Hook. f., 4. montt-
cola, Schult., and A. Auchert, Boiss.
To these we add 4 species not noted from the Presidency before : Chrysopogon
Wightianus, Ness, Ch. asper, Heyne, Chr. polyphyllus, Blatter and McCann, and
Ch. Gryllus, Trin.
A. Pedicels of the upper spikelets half as long as
the sessile spikelets or longer
I, Pedicels of upper spikelets glabrous or
nearly so
1. Stems erect. Leaves 15-45 cm. long ae le VER Gryllus:
2. Stems creeping below. Leaves 2-13 cm.
long ... a wis at On ACLCULOLIES.
il. Pedicels' of upper spikelets villous with
rusty rarely pale hairs
1. Lower involucral glume of pedicelled
spikelets long-awned, upper not or very
shortly awned
(a) Callus long villous all round 3, Gi asper.
(6) Callus giabrous in front ... 4, C. lanceartus.
2. Involucral glumes of pedicelled spikelets
both awned - 5. C. Wightianus.
B. Pedicel of upper spikelets not half as long
as the sessile spikelets
1. Lower sheaths compressed 6. C. montanus,
II. Lower sheaths terete
1. Leaves, peduncle and branches of panicle .
glabrous 7. C. polyphyllus.
2. Leaves, peduncle and branches of panicle
not glabrous oe ie an On Ce AUCHELT,
1. Chrysopogon Gryilus, Trin. Fund. Agrost. 188 ; Nees Gen. Fl. Germ. Mono-
cot. i, t. 93; Benth. Fl. Austral. vii, 537.—Andropogon Gryllus, Linn. Cent. Pl.
iso; black. Monogr. Androp. 550); Host. Gram. Austr: 11, 1, t21; Sibth, Fi,
Grec.. i. t. 67; Duthie Grass. N. W. Ind. 22, Fodd. Grass. N. Ind. 40; Hook.
f.in F. B. I. vii, 187; Collett Fl. Siml. 602, fig. 191.—A. echinulatus, glabra-
tus et Royleantus, Steud, Syn. Gram. 395, 397.--Chrysopogon glabratus, AMabor
in Mem. Acad. Petersb. ser. 6, ii (1833), 318. —khaphis Gryllus, Desv. Opusc.
69.—F. echinulata, Nees in Royle Ill. Bot. Himal. 417.—FPollinita Gryllus,
Spreng. Pugill. ii, 10; Reichb. Ic. Fl. Germ. t. 54.--4pluda Gryllus, Pres.
Cyp. & Gram. Sic. 55.—Holcus gryllus et pallidus, Br. Prodr. 199.
Description ; Stems simple, forming dense hard tufts, erect, 15cm. to 1°5
m. high; nodes smooth, Leaves 15-45 by 4-8 mm., linear, acute, glabrous or
hirsute, margins serrulate; sheath keeled above, glabrous or pubescent.
Panicle large, 12-20 cm., rhachis angular, scabrid, axils bearded, branches long,
5-10 cm., capillary, spreading, simple or branched, usually very many ina
whorl and bearing 2-4 spikes, tips obliquely truncate and densely bearded.
Sessile spikelets 5-8 mm., callus straight, acute. Lower involucral glume
coriaceous, 2-toothed, dorsally rounded with 2 muricate or mamillate keels or
channels, shining, smooth or scaberulous, margins broadly involute, upper
chartaceous, hyaline, lanceolate, mucronate or aristulate, awn equalling the
spikelet or shorter, keel and sides bristly above the middle. Lower flora] glume
linear-oblong, obtuse, nerveless, upper linear, minutely 2-toothed, awn minute
or 12-35mm. long. Pale small, oblong, glabrous. Pedicelled spikelets rather
longer than the sessile, terete, lanceolate, acuminate; pedicels glabrous or
ciliolate. Lower involucral gluine acuminate or aristulate, 5-9-nerved keels
412 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol XXXII
ciliate above, upper lanceolate, acuminate, ciliate. Floral glumes narrower,
ciliate, awn of upper half the size of the glume.
Locality: N. Kanara: WHalyal (Talbot 2088 !).
Distribution: Temperate Himalaya from Kashmir to Sikkim, 4,000-9,000
ft. Khasia Hills, 4,000-5,000 ft., westwards to N. Africa and S. Europe,
Australia.
2. Chrysopogon aciculatus, Trin. Fund. Agrost. 188; Duthie Grass. N. W.
Ind. 22 (acicularis), Fodd. Grass. N. Ind. 39; Benth. Fl. Hongk. 424, Fl.
Austral. vii, 538.—Andropogon aciculatus, Retz. Obs. v (1789), 22 ; Roxb. F1.
Ind..i, 262 ; Grah. 238; Hack) Monogr. Androp. 562); Hook.f. in F.3d .gvais
188; Cke. ii, 984.—A. acicularis, Willd. Sp. pl. iv, 906.—Ahaphis acicularis,
Desv. Opusc. 69.—A. trivalvis, Lour. Fl. Cochinch. 553 ; Trin. Sp. Gram. Ic.
t. 8, 9.-—Centrophorum chinense, Trin. Fund. Agrost. 106, t. 5.— Rheede Hort.
Mal. xii, t. 43.
Description: Cke.l.c.
Locality : Konkan: Alibag, sandy shore (Ezekiel!).—NV. Kanara; Karwar,
sea-shore (Sedgwick and Bell 5070!) ; Jog, hills (Hallberg and McCann A272 !).
Distribution: More or less throughout India, Ceylon, Tropical Asia,
Australia, Polynesia.
Uses: According to Haines the leaves which lie close to the ground escape
to a large extent the lips of cattle. The plant is a pest on account of the sharp
callus and small awns sticking to the clothes.
3. Chrysopogon asper, Heyne ex Wall. Cat. n. 8784.—Andropogon asper,
Heyne in Herb. Rottler ex Hook. f. in F.B.I. vii, 189.
Description : Stem 30-90 cm., leafy below, very slender above. Leaves
distichous, 30-45 cm. by 12-18 mm., broadly linear, acute cordate, coriaceous,
flat, smooth, li-nerved, midrib very slender, spinulose beneath, margins
spinulosely serrulate, and with a few long tubercle-based cilia towards the
broad semi-amplexicaul base ; sheaths broad, compressed, laxly hirsute, lower
ones 12 mm. broad, keeled, armed with scattered tubercle-based hairs.
Panicle 18 cm. long, narrow, of many whorls of short, unequal, simple, smooth,
erect branches bearing solitary rarely 2 erect spikes, tips very shortly bearded.
Sessile spikelets 6 mm. long, pale, coriaceous, callus up to 2 mm., long villous
all round. Lower involucral glume hispid beneath, the tip strongly compressed
above ; upper with the keel and sides more or less hispid above the middle, awn
as long as the glume or shorter. Lower floral glume shorter than the upper
involucral, narrow, obtuse, 2-nerved, ciliate, upper consisting of the linear,
hyaline, 3-nerved base of the awn, awn 35-50 mm. long. Pedicelled
spikelets 8 mm. long, narrowly lanceolate, 7-nerved, pale, shining, keels
ciliate ; pedicels very shortly rufous-villous on both margins, excised at
the tip in a semi-circle ; lower involucral glume thin, tip 2-dentate, nerves
strong, sub-equidistant, or the 3 lateral on each side submarginal, margins
narrowly incurved, keels ciliolate from base to tip, upper lanceolate, acuminate,
3-nerved, ciliolate. LI.ower floral glume linear-oblong, 2-nerved, ciliate, upper
narrowly lanceolate, 1-nerved.
Locality : N. Kanara: Tinai (Talbot 2564 !).
Distribution : Madras : Pulicat Hills.
4. Chrysopogon lancearius Haines in Haines Bot. Bihar and Orissa 1036.— Axndro-
pogon lanceartus, Hook. f.in F. B. I. vii, 190; Cke ii, 985.
Description . Cke. l.c.
Locality: Deccan; Panchgani (Blatter and Hallberg B 1230;).—S. AZ.
Country: Castle Rock (Woodrow). Seems to be very rare in the Presidency.
Distribution : Sikkim Himalaya, Behar, Chota Nagpur, Orissa, W. Penin-
sula.
Uses: A good fodder according to Haines.
5. Chrysopogon Wightianus, Nees ex Steud. Syn. Gram. 397.—Andropogon
Wightianus, Steud. Syn. Gram. 395; Hook. f. in F. B. [. vii, 191.—A. aristu-
latus, Hochst. ex Steud. Syn. Gram. 397; Hack. in Monogr. Androp. 556.—
A. breviaristatus, Steud. 1. c. 396.—Rkhaphis orientalis, Desv. Opusc. 69.—
Rk. Wightianus, Nees ex Steud. 1. c.
Description : Very variable in habit. Stems short or long, erect or ascending
from a short stout creeping stock. Leaves 7-25 cm. long, linear, acute, rigid,
from glabrous to pubescent on both surfaces and with sometimes tubercle-
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 413
based hairs, spinulosely serrulate ; sheaths glabrous, lower ones compressed ;
ligule very short, villous. Panicle 7-13 cm. long, contracted, lower branches
long, few in a whorl, rhachis and branches minutely hairy ; spikes solitary, green
or brownish. Sessile spikelets subcylindric, 4nim. long, callus long, densely
bearded with rusty hairs all round. Lower involucral glume laterally compres-
sed above, minutely truncate, glabrous below, hispid above, obscurely 4-nerved,
tip 2-dentate, upper chartaceous, hispid above onthe keel and sides, tip 2-
lobed, awn as long as the glume cr shorter. Lower floral glume linear-oblong,
2-nerved, ciliate, upper consisting of an awn with a narrowly dilated 2-lobed
base, awn 50-65 mm. long, column hispid. Pedicelled spikelets nearly 12 mm.
long, lanceolate, pubescent ; pedicel truncate, margins shortly villous. Lower
involucral giume glabrous or pubescent, 7-nerved, awn longer than the glume,
keels ciliate, upper lanceolate, 3-nerved, awn as lonz asthe glume or shorter,
Tower floral glume oblong, 2-nerved, ciliate, upper very narrow, ciliate,
nerveless.
Locality : S. M. Country ; Castle Rock (Bhide!). — NV. Kanara ; Jog to Sid-
derpur, open grass land (McCann A273!).
Distribution : Madras, Nilgiris, Burma, Assam. (Hackel mentions a speci-
men gathered in Ceylon but, according to Hooker f., it seems to be a starved
specimen of Chrysopogon zeylanicus, Thw.).
6. Chrysopogon montanus, Trin. in Spreng. Neue Entdeck, ii, 93; Haines Bot,
Bihar and Orissa 1037.—Andropogon monticola, Schult. Mant. (1824), 665 -
Kunth Enum. Pi. i, 506 ; Steud. Syn. Gram. 395; Hack. Monogr. Androp. 557
(excl. var. velutinizs); Hook f. in F. B. 1. vii, 192, cum omnibus var.; Cke. ii,
985 ; Holein Ind. Forest Mem. i (1911), 108.—A. Sprengeliz, Kunth Rev. Gram.
166.—Pollinia fulva, Spreng. Pugill. ii, 93—Andropogon Triniz, Steud. Syn,
Gram 395; Hack. Monogr. Androp. 558.-- A ciliolatus caruleus et
tucrescens, Steud. 1. c.— Chrysopogon ciliolatus, Boiss, Fl. Or. v. 458 (excl. var.
Aucheri, Boiss); Duthie Grass. N. W. Ind. 22.—C c@eruleus, Duthie 1. c. 23,
Fold. Grass. N. Ind. 39, t. 60.—C. increscens, Nees ex Steud. 1. c. 396.—C.
Wightianus var. leucanthus, Thw. Enum. Pl. Zeyl. 366.—C. serrulatus, Trin.
in Mem. Ac. Petersb. ser. 6, ii (1833), 318, Spec. Gram. t. 331.— C. Esenbeckit,
Arn. in Steud. Syn. 395.—Aaphis ciliolata et cerulea, Nees ex Steud. Syn.396, 395.
This synonymy requires an explanation. Chrysopogon montanus, Trin. as
understood in this place comprises Hackel’s two species Andropogon monticola
Schult. and A. Triniz, Steud., and is identical with Hook. f.’s. 4. monticula,
Schult. with all its varieties.
Hackel has two species and he distinguishes them by the following characters :
A. monticola : Upper involucral glume of sessile spikelet keeled, the keel
from the base up to 3-7 of its length densely pectinate ciliate with long, rigid,
rufous hairs, shortly white hispid in the upper 7.
A. Trinii: Upper involucral glume of sessile spikelet keeled below the apex
only, keel white-ciliate, the lower §-$ not keeled and glabrous.
Hook. f. in F. B. I. makes of these species two varieties : var monticola proper
and var. Triniz, and includes them under A. monticola, Schult., adding a third
variety : var. robustus.
At the same time Hooker confesses: ‘lam unable to classify the varieties of
this common and variable plant in accordance with geographical areas or
other considerations. This, if possible, must be effected by field-botanists in
India. There is every gradation from the coarsely hirsute keel of szontzcola,
to the perfectly smooth of some states of Zz7imzz; from the awnless to long
awned gl. I of the pedicelled spikelets, and from the glabrous to the pubescent
of the same organ ; the colour of which affords no character; nor does its
length, orthat of the cilia on its keels.’
Cooke (ii, 985) bas adopted the name A. monticola, Schult. with Hooker’s
description and evidently also the latter’s varieties. But his opinion does not
count in this case as he hasnot seen any specimens from the Presidency and
‘ was therefore ’’ as he says himself,‘ unable to fix definitely the variety to
which the Bombay species belong. They will probably belong to var. Trinzi
H.f.’ What induced Cooke to say that they probably belong to var. 7yzn7zz we
cannot understand, especially as Hooker came to the conclusion that he was
not able to classify the varieties according to geographical areas.
We have examined a great number of specimens from all parts of the Presi-
dency, except Sind, Cutch and Kathiawar and we have been able to separate
3
4144 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
many into the two varieties. They exhibit almost the same distribution and
often both are found in the same locality, with this exception that var. 7riniz
has not been observed in N. Kanara, Gujarat and the Konkan. But we must
also mention, and thisis the most important point, that we saw many specimens
all over the country which could net beclassed under either variety, and it
would require many new varieties if we wanted to give aname to all the
different variations. Andeven then they would be forms only and not
varieties.
Haines seems to have felt the same difficulty when he tried to classify the
montanus material of Bihar and Orissa. He distinguishes five forms. If we
wanted to follow the same method for our area, we doubt whether double the
number of forms would yield satisfactory results.
Stapf describes the specimens from Tropical Africa under the name of
C. montanus var. tremulus, Stapf. He calls it ‘ one of the several races which
constitute the rather polymorphic species C. montanus, Trin., whose area
includes Southern Africa, Madagascar and India. The var. tremulus ap-
proaches very closely the var. elatzor, Stapf, a large-spikeleted parallel to the
var. serrulatus, Stapf (Chrysopogon serrulatus, Trin.) and differs from it
apparently only in the almost quite smooth rhachis and branchlets (a few sharp-
pointed hairs may be found under a high power) and the pedicels, which are
glabrous almost up to the middle, and not ciliate from the base.’ It seems
to us (it may look like presumption on our part to criticize our veteran and
highly merited agrostologist) that it is somewhat risky to found new varieties
of a protean species on a few specimens only.
According to Hole all the 3 varieties mentioned by Hooker ‘appear to vary
greatly, as regards their habit and vigour of growth, in response to the moisture
conditions of the habitat and also according as whether, or not, the plants
are habitually grazed, cut for fodder, or periodically burnt. The colour of
the cilia of glume II of the sessile spikelet (pale or white in robustws and rufous
in monticola), accordingly, appears to be the chief difference in the habit, and
these forms appear to have different and fairly defined areas of distribution
(monticola occurring citiefly in Central and Southern India, while vobustus is
mainly found in N. India, in the outer N. W. Himalayas and Sub-Himalayan
tract).’ Hole who studied the varieties vobustus and Tyrinzi both from
herbarium specimens and in the field kas observed that the plant at Dehra Dun
graduaily and imperceptibly passes from the typical robustus to the typical
Trinit. We are justified in stating that a similar transition takes place between
monticola and Triniz in Western India. (Of Central and Southern India
we have no experience). We have therefore a gradual transition from
robusius to Triniz at Dehra Dun, and from TJyriniz to monticola in W. India
and consequently, we are not allowed to consider Hooker’s varieties as good
varieties.
Vern. Names: Sunthia Khad (Dohad), Agiva, Gogar, Ghora, Dand,
Pandhari Kusal (Poona), Kare Hullu (Bijapur).
Description: Avery variable perennial grass. Stems usually slender, erect
or geniculately ascending, glabrous, sometimes robust, simple or branched,
30 cm. to 1°2 m., but often attaining 2 m., slightly compressed, solid, develop-
ing usually axillary leafy and flowering branches from all the upper nodes except
the one next below thepanicle. (The branches growing within the sheaths
push the latter away from the stem which often results in a characteristic fan-
shaped appearance.) Blade of uppermost leaf of flowering stem usually
mucroniform, but attaining 8 cm., of lower leaves up to 43cm. long and 8 mm.
broad, linear acuminate, tapering from the base, scabrid on margins, sometimes
also scabrid dorsally on midrib, and scaberulous above, especially towards the
apex, often ciliate towards the base with tubercle-based hairs, at least when
young; sheath glabrous. compressed, keeled, especially of the lower leaves,
shorter or longer than the proper internode ; ligule a minute membranous rim.
Panicle 5-15 cm. long, ovate tosubcylindric, yellowish to purplish, of several whorls
of few or many capillary flexuous very unequal branches bearing solitary spikes,
branches of flowering panicle more or less horizontally spreading, of the fruiting
panicle erect and closely appressed to the rhachis, rhachis and branches smooth
or scaberulous. Spikelets in clusters of 3, a central sessile hermaphrodite one
with 2 lateral pedicelled male ones, the clusters being terminal and solitary on
the capillary branches of the panicle. Sessile spikelets laterally compressed,
4-7 mm, long, tip of peduncle brown-bearded, clavate, callus short, with ova]
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 415
scar and dense beard. Lower involucral glume laterally compressed, narrow-
oblong, embracing the margins of the upper, chartaceous 2-4—-nerved, hispidly
ciliate dorsally on keel towards the apex or almost glabrous, often scaberulous
dorsally on nerves and minutely pubescent with appressed hairs dorsally near
margin, apex subtruncate or 2-dentate. Upper involucral glume laterally com-
pressed, broader than lower, obtusely keeled, subcoriaceous, 3-nerved, margins
broad, hyaline, membranous, ciliate or not, very variable with regard to its
hairiness, sometimes almost glabrous, at other titnes hispidly ciliate dorsally on
keel with long white or rufous hairs more or less from base to apex, sometimes
also pubescent, or minutely villous dorsally on keel and lateral nerves, awned,
awn 2°5-6 mm. long, apex entire or 2-lobed. Lower floral glume 7 the length
of to subequal the upper involucral glume, linear, hyaline, ciliate, nerveless or
indistinctly 1-3- or more-nerved, apex obtuse. Upper floral glume consisting of
the narrow 3-nerved base of the awn, basal 3 or 3 hyaline, membranous,
upper portion chartaceous, awn geniculate, 10-18 mm. long, but also reaching
37 mm. (including the twisted column), margins ciliate or not, apex entire or
2-lobed. Pale sometimes present, very narrow, 1:°25mm. long: Lodicules 2,
cuneate, glabrous. Anthers3, upto3mm. long, yellow orpurple. Stigmas
2, laterally exserted at base of spikelet, yellow. Pedicelled spikelet dorsally
compressed, subequal the sessile spikelet ; pedicel less than half the sessile
spikelet, usually about 3 the spikelet, densely ciliate on both margins with stiff
rufous or white hairs, the upper of which are shorter than to subequal the
spikelet. Lower involucral glume lanceolate, membranous, 5-7-nerved,
minutely pubescent with appressed hairs dorsally, especially towards the apex,
or almost glabrous, sometimes ciliate dorsally on midrib and marginal nerves,
especially towards the apex acute or shortly awned. Upper involucral glume
subequal to the lower, 3-nerved, marginsincurved, long ciliate, apex acute or
mucronate, glabrous dorsally. Floral glumes linear, hyaline, ciliate, nerveless
or indistinctly nerved. Pale sometimes present, as in sessile spikelet, but
slightly longer, very narrow.
The flowers are much visited by smal! bees. sean
Locality : Gujarat: Mahal-Dangs, elevation 800 ft., rainfall 100” (Sedgwick
and Bell 5391 !) —AKhandesh: Tapti River (Blatter and Hallberg 5476!); Bhusawal
(McCann 5224 A!); Nandgaum, Bori River (Blatter and Hallberg 3827!) ;
Bori, Tapti Island (Blatter and Hallberg 5146!) ; Amalner, Bori River (Blatter
and Hallberg 4455!).--Konkan : Sion Creek (Sabnis A 231 !); Matheran
(D’Almeida 9958 !).—Deccan: Khandala to Karjat (Blatter and Hallberg
A. 232!) ; Kirkee (Talbot!) ; Pashan (Gammie!) ; Mangiri, 8 miles E. of Poona
(Gammie !); Katraj (Gammie !); Pasarni Ghat (Blatter and Hallberg
B 1209 !).—S. MW. Country > Dumbai (Talbot 2317!) ; Badami (Talbot 2926!) ;
N.W_ of Dharwar (Sedgwick 3141 !); Dharwar, dry pasture land, elevation
2 400 ft., rainfall 34” (Sedgwick 1817!) ; Konankeri, elevation 1,800 ft., rainfall
35” (Sedgwick and Bell 4439!); Haveri (Talbot 2189!).—Kanara; Jog to
Siddhapur, open grass land (Hallberg and McCann A274!). This species
forms patches of many individuals, or combines with other individuals to form
associations of a few species.
Distribution - Throughout India, especially in hilly tracts, from the N.W.
Himalaya southwards, ascending to 6,000 ft., extends to Ceylon, Burma,
Afghanistan, Tropical and S. Africa, Madgascar.
Uses : In Bihar and Orissa this grass is considered to bea valuable fodder,
and Hole, writing of the Siwalik Division, calls it one of the most valuable
fodder grasses. In Mount Abu, according to Lisboa, it is reckoned as a good
fodder grass and the grain is used as food by the natives, But the same writer,
under the name of Axdropogon serrulatus, Trin. (= Chrysopogon montanus
var. Frinii) remarks: ‘Said to be good fodder, used much in Poona, but
reports from other places unfavourable.’ (J. C. Lisboa, List of Bombay
Grasses, Bombay (1896), 81).
The last statement might find an explanation by a suggestive note made by
Hole (l.c. 111): ‘So faras the local (Dehra Dun) plant is concerned speci-
mens with the more hairy glume II tend to occur in localities where there is a
scarcity of available moisture, both on the dry ridges and slopes of the Siwalik
Hills and also (rarely) on water-logged soil, and the writer believes that the
characters which have been utilized to define these varieties vary in response to
the factors of the habitat and particularly in response to the available water
supply. Provided that the development of the plants has not been interfered
416 JOURNAL, BOMBAY NATURAL HIS17. SOCIETY, Vol. XXX1i
with by grazing, grass-cutting, or other agency, those plants with the more
hairy glume II are usually less robust and less coarse, or rank, than the others,
and they are therefore as arule most valued for fodder and are distinguished
locally by the vernacular name of dhaula, whereas the coarser plants with
smooth glume arecalled gurv/a. As this grass affords a valuable fodder and is
sometimes cultivated, in consequence, it is important to determine the extent to
which its characteristics are constant. If, as suggested above, they depend on
the available moisture, it is obvious that cultivation of this grass on good
agricultural land, with a large quantity of available moisture, would result in
producing an inferior class of rank, coarse fodder.’
7. Chrysopogon polyphylilus, Blatter & McCann, comb. nov.— Andropogon
polyphyllus, Hook. f. in F. B.I. vii, 194 (gee habet Hack. mss. in Herb.
Duthie).—A. Aucheri, var. polyphyllus, Hack.in Herb. Duthie.
Description : Stem 60-90 cm, high, as thick as a crow-quill or more, stiff,
simple or fastigiately branched, quite glabrous. Leaves crowded or not, 15-25
cm. by 2-4 mm., narrow, rigid, acuminate, flat, pale glaucous-green, glabrous
on both surfaces, coriaceous, midriband nerves very slender, margins minutely
scaberulous ; sheaths terete, appressed, hard. Panicle 10-13 cm. long, oblong,
subsecund, dense- flowered, very pale, branches 6-12 mm., very unequal, in many
closely approximate whorls, smooth, peduncle slender, quite glabrous.
Sessile spikelets 4mm. long drooping, white or pale purplish, callus long,
ieiooheels long, obtuse, bearded at the very base only with long fulvous hairs.
Glumes asin C. montanus. Lower involucral glume obtuse, glabrous, keel
ciliate towards the tip; upper not awned, keel glabrous or ciliate. Upper
floral glume with an awn 3-8 mm. long, nearly straight, pale. Pedicelled spike-
lets narrowly lanceolate, acuminate, glabrous, 7-nerved, eciliate, not awned,
rather longer than the sessile ; pedicels naked, villous at the tip only.
Can easily be distinguished from C. montanus by the stout naked callus
Which is bearded at the base only, and by the naked pedicels which are long-
villous only at the tip.
It differs from the next species, C. ducheri by its size, the long, glabrous,
glaucous leaves and the glabrous peduncle and branches of the panicle.
Locality : Gujarat > Porbandar (Bhide !) ; Dohad (Bhide!) ; Watrak River
on rocks (Sedgwick 1165!) ; Daman (Bhide).—Deccan - Dhond, river-bank
(Bhide !).
Distribution - Central Provinces. W. Peninsula.
8. Chrysopogon Aucheri, Stapf in Kew Bull. (1907), 211. —Andropogon Auchert,
Boiss. Diag. ser. 1, fasc. 5 (1844), 77; Hook. f. in F.B.I. vii, 155; Cke., ii, 986.-A.
Auchert, var. genuinus, Hack. Monogr. _Androp. 560. NG ‘hrysopogon ciliolatus,
var. Aucheri, Boiss. Fl. Or. v, 458.
Description : Cke.Akc,
Stapf thinks that C. Aucheri comprises several geographical] races and that
the one from which the species was first described extends from Arabia through
Southern Persia and Baluchistan to Sind. He characterizes it‘ by the lower
glume of the pedicelled spikelet being usually awnless or in any case much
more shortly awned thanthe upper, by the glume awns not being ciliate or
ciliate only at the base, and by the longer beards of the pedicels.’
Locality : Sind: Gizri (Sabnis B777!) ; Jemadar ka Landa near Karachi
(Stocks).
Distribution: Sind, Baluchistan, Afghanistan, S. Persia, Arabia (not
Africa).
30. ARTHRAXON Beauv. Agrost. 111; Cke. ii, 967.
Species about 20, in the tropical and subtropical regions of the Old World.
We retain the 6 species mentioned by Cke. lc. His Al. lanceolatus,
Hochst. will be slightly restricted under the name of 4. serrulatus, Hochst., the
name A. lancifolius, Hochst, will be substituted for A. microphillus, Hochst.,
and 4. guartinianus, Nash, will take the place of 4. czlzaris, Beauv.
Key as in Cke.
1. Arthraxon inermis, Hook. f. in F. B. I. vii, 145; Cke. ii, 968.
Description: Cke.l.c.
Locality : Konkan : Okda Forest, Thana District (Ryan 718!); Wada Range,
Thana District (Ryan 692!) ; Matheran (Woodrow !); Marmagoa (McCann !).
— Deccan : Mahableshwar (Sedgwick and Bell 4513!, Woodrow) ; Purandhar
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 417
(McCann 5592!); Khandala (McCann 9950 !, 9740!).—S. WZ. Country : Derikop
woods (Sedgwick 1845!) ; Castle Rock (Gammie 15678 !).
Distribution: W. Peninsula, apparently endemic.
2. Arthraxon serrulatus, Hochst. in Flora (1856), 188; Stapf in Fl. Trop. Afr.
ix, 163 —A. lanceolatus, Hack. in Monogr. Androp. 348 (excl. var. echinatus) ;
Duthie, Grass, N. W. Ind. 17; Hook.f. in F.B.I. vii, 143 (partim excl. Batra-
therum echinatum, Nees in Edinb. Phil. Journ. xviii, (1835) 181; Andropogon
echinatus, Heyne ex Steud. Nomencl. ed. 2, 91: Arthraxon echinatus, Hochst.
in Flora (1856), 188 ; Andropogon lanceolatus, Roxb. Fl. Ind. i, 257) ; Cke. ii,
968 (partim). A. lanceolatus var genuinus, subvar, serrulatus, Schweinf. in
Bull, Herb. Boiss., 2. ser. ii, 10.—Andropogon serrulatus, Link Hort. Berol. i,
241 (quoad specimen, descriptio partim erronea?); A. Rich. Tent. Fl. Abyss. ii,
458.—A prionodes, Steud. Syn. Pl. Glum. I, 383.—Satratherum lanceolatum,
Nees in Edinb. New Phil. Journ. xviii (1835), 181.-——. serrulatum, Hochst., ex
Steudlic.
This species includes all the material of the Nileland of Tropical Africa, of
tropical Arabia and the greater part of the Indian specimens which, up to now,
were ranged under Arthraxon lanceolatus, Hochst., as understood by most
authors.
It has been pointed out by Stapf that Arthraxon lanceolatus, Hochst., was
founded on Andropogon lanceolatus, Roxb., a Coromandel plant, which has
larger and wider long-awned spikelets with the lower involucral glume very
minutely imuricate towards the tips only. Neither of these names can,
consequently, be mentioned as synonyms of Arihraxon serrulatus.
Description.—A perennial grass; rhizome short, emitting fascicles of closely
set culms and innovation shoots, which are more or less thickened below and
covered with reddish silky cataphylls. Stems rather slender, up to 90 cm.
high, many-noded, usually finely pubescent, sometimes glabrous, with vegeta-
tive branches below and usually solitary flowering branches above, the latter
subfastigiate. Leaf-blades lanceolate to ovate-lanceolate, setaceously acumi-
nate, witha caudate amplexicaul base. glaucous or greenish, 2°5-5 cm. long, 12-
18 mm.wide, margins cartilaginous, ciliate with the cilia springing from tuber-
cles, smooth, very finely pubescent below or glabrous, primary lateral nerves
about 8-11 on each side, fine,slightly raised below; ligule 1-2 mm. long, rounded,
membranous; sheaths terete, tight, slightly shorter than the internodes or exceed-
ing them in the leafy shoots, more or less hairy with tubercle-based hairs and
often softly pubescent at the nodes, the uppermost frequently glabrous. Racemes
2-5 nate, slender, greenish or suffused with purple or violet, 3-over 5 cm. long on
a short common axis, the fascicles borne on a slender peduncle, shortly or far
exerted from the supporting bladeless or almost bladeless sheath ; rhachis fragile
shortly bearded at the nodes ; joints narrowly linear, 3°3-6°3 mm. long, hairy on
the back and sides, or the lowest almost glabrous, hairs increasing upwards to
more than 2 mm.; pedicels very similar, but much shorter and more slender.
Sessile spikelets narrowly lauceolate-linear to linear slightly tapering upwards
upto 74mm. long (not including the awn), glabrous; callus very short,
puberulous. Glumes 4. Lower involucral glume chartaceous, 6 mm. long,
lanceolate, with a minutely truncate hyaline tip, very convex on the back,which
is smooth or more often muricate along the 4 indistinct or faintly raised
inner, spinulously muricate along the outer nerves, margins hyaline, compara-
tively wide ; upper linear-lanceolate in profile, membranous, 3-nerved, glabrous
or nearly so. Lower floral glume linear, acute, up to 4 mm. long, hyaline,
nerveless or obscurely l-nerved at the base; upper membranous at the base,
hyaline upwards, narrowly lanceolate-linear in profile, 2-dentate, with the
teeth minute, sometimes produced into short capillary bristles, delicately
3-nerved, awned from near the base, awn 8-15 mm. long, very fine, kneed
about the middle, twisted below. Anthers up to3 mm. long. Grain bacilliform
about 4°2 mm. long. Pedicelled spikelet male, linear-lanceolate, acute, about
5 mm.long. Involucral glumes subequal, lower herbaceous-chartaceous, acute,
Scaberulous along the outer and slightly soon the 4-5 inner nerves or almost
smooth, upper slightly shorter, membranous, otherwise as in the sessile spikelet.
Lower floral glume as in the sessile spikelet, upper linear-ob!ong, obtuse,
nerveless, muticous.
Locality : Sind: (Stocks 642).—Gujarat- Morvi, Kathiawar (Woodrow).—
Deccan : Lonavla (Woodrow) ; Pashan, tank (Gammie!). Panchgani (Blatter
418 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Volt, XXXII
and Hallberg B1277!).—S. M. Country: S. W. of Dharwar (Sedgwick and
Bell 4436!) ; Derikop (Sedwick 2022!); Belgaum, Fort wall (Sedwick
3013 !).
Distribution ; More or less throughout India, Tropical Arabia and Tropical
Africa ( Nileland).
3. Arthraxon Meeboidii, Stapf in Cke. ii, 969, in Kew Bull. (1908), 449.
Locality: Western Ghats: Khandala, in open grassland on a hillside, 600 m.
(Meebold 9132) ; Khandala, common (McCann 9949!, 9948!); Lonavla,
eommon (Bhide!, McCann!); Panchgani (Blatter and Hallberg B1238!,
B1225!), Tableland (Blatter 3804 ;); Mahableshwar (Sedgwick and Bell 4523).
__N. Kanara: Tinai (Talbot 2569!).
Distribution - W. Peninsula, apparently endemic.
4. Arthraxen lancifolius, Hochst. in Flora (1856), 188; Stapf in FJ. Trop.
Afr. ix, 165.—-A. microphyllus. Hochst. in Flora (1856), 188; Hack. Monogr.
Androp. 351, imel. var. lanctfolius; Hook.f. in F.B.I. vii, 147; Cke. ii, 970 ;
Haines Bot. Bihar and Orissa 1026.—A. Schmidtii, Hochst. (1856), 189.—A.
minor, Hochst. l.c. 188, partim.—A. Schimperi, Hochst. l.c. partim.—A.
molle, Balt. f. Bot. Socotra 315.—A. ciliaris, Henriq. in Bolet. Soc.
Brot. xiii, 133, mom Beauv.—Andropogon lanctfolius, Trin. in Mem. Acad.
Petersb. 6 ser., ii (1833), 271.—A. molle, Duthie Grass. N.W. Ind. 17.—
A. multicaulis, Steud. Syn. Pl. Glum. i, 383.—Batratherum molle, Nees &
Arn. in Edinb. New Phil. Journ., xviii (1835), 181; Aitchis. Cat. Panj. Pl.
174.—B. Schimperi, Nees ex Hochst. l.c. 179 —Psilopogon Schimperi, Hochst.
ex A. Rich. Tent. Fl. Abyss. ii, 447.—APleuroplitis ciliata, J. Schmidt Beitr.
Fl. Cap. Verd. 152.—P. Schimpert, Regel in Bull. Acad. Petersb. x (1866),
369.—Lucea ciliata, Steud. l.c. 414.
Description : Cke. l.c.
Locality : Gujarat: Surat (Herb. Dehra Dun !, Cooke !).—Konkan : Tun-
gar hill, Thana District (Herb. Econ. Bot. Poona!); Parel (Herb. Econ, Bot,
Poona!); Kutla (Garade!); Salsette (Jacquemont 713).—Deccan : Trimbak,
Nasik District (Herb. S. X. C., Bombay!).—S. MW. Country: Belgaum
Fort walls (Sedgwick 3012!); W. of Dharwar, banks of road, in forest
(Sedgwick 1851!); Castle Rock (Bhide !).—Kanara: Karwar (Talbot 1308 !)
Yellapore (Talbot 2084 !).
Distribution : More or less throughout India, Ceylon to Tonkin and S. W.
China, tropical Africa (Upper Guinea, Nileland).
5. Arthraxon quartinianus, Nash in North Americ. Fl. xvii (1912), 99,
Merrill in Phillip, Journ. Sc. Bot. vii. (1912), 229; Stapf in Fl. Trop. Afr: ix
166.—A. major, Hochst. in Flora (1856), 188.—A. coloratus, Hochst. 1.c.—-
A. plumbeus, Hochst. l.c. 189.—.4. violaceus Hochst. l.c.—1. Schimperi,
Hochst. l.c. partim.—A. ciliaris, Rendle in Cat. Afr. Pl. Welw. ii, 138, xon
Beauv.—A. ciliaris, subsp. guartinianus, Hack. Monogr. Androp. (excl. vars.
Hookeri et glabrescens).— Alectoridia quartiniana, A. Rich. Tent. Fl. Abyss.
ii, 448, t. 99. -Lucea major, Steud. Syn. Pl. Glum. i, 414; Hochst. l.c. 179 —
L. plumbea et violacea, Steud. l.c.—L. Schimpert, Hochst. in Flora (1856), 180
(the genuine).— Pleuroplitis plumbea, Nees ex Steud. 1.c.—P. violacea, Nees
ex Steud. l.c.—P. major, Regelin Bull. Acad. Petersb. (1866), 369, partim,—
P. quartiniana, Regel !.c. 376,partim.— Andropogon violaceus, Heyne ex Steud.
Jd.e.—A. Alectoridia, Steud 1.c. 383.
We wish to substitute 4. guartinianus, Nash in place of J. ciliaris, Beauv.
This latter species which was described in 1812 has been emended repeatedly
since then, and that to such an extent that it is scarcely possible to recognize
the original plant. Hackel made five subspecies: Langsdorffit, submuticus,
nudus, Quartinianus, and Vriesii, and nine varieties. The subspecies were
arranged by him in two leading groups :
1. Joints of rhachis glabrous, or with a few scattered hairs towards the tip
(Langsdorfi1, submuticus, nudus).
2. Joints of rhachis, at least the upper, more or less ciliate (Quartinianus,
Vriesii).—To these two groups Hooker f. added athird one (F.B.I., vii, 146):
3. Spikes silkily villous, spikelets 5-7 mm. long, lower involucral glume
entire or minutely 2 toothed, awn 12 mm. or more. (This is Arthraxon
cus pidatus, Hochst., considered by Hackel as a distinct species.
Arthraxon quartinianus, Nash, and adopted by Stapf. c., covers part of the
subspecies Quartinianus in Hackel’s group 2, including var. Quartinianus
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 419
s. sty. and var. coloratus, but excluding var. Hfookeri and var. glabrescens.
Quartin’s specimen No. 14, collected in Abyssinia, and representing a form with
large spikelets, has, according to Stapf, to be considered as the basis of the
species, whilst Schimper’s 1532, the type of Arthraxon coloratus, stands for
the other extreme.
Arthroxan quartinianus, Nash, includes all the material of the Bombay Pre-
sidency that has come under our observation.
The rest of A. ciliaris, Beauv. et auctorum has to be studied separately, but
as the material lies outside the Presidency we leave it to other botanists to work
out their respective material.
Description: Annual. Stems very slender, ascending from a_ branched,
sometimes prostrate and rooting base, from 8-30 cm. high, smooth, glabrous or
finely puberulous below the inflorescence, with solitary or 2 to 3-nate branches
above, internodes mostly exserted. Leaf-blades ovate-lanceolate to oblong-
lanceolate from a cordate amplexicaul base, finely acuminate, 25 to over
50 mm. long by 6-12 mni. broad, flaccid, with scattered tubercle-based hairs on
both sides, ciliate towards the base, primary lateral nerves 3-4 on each side,
very fine ; ligules membranous, ciliolate, short, truncate ; sheaths lax, the upper
very often more or less inflated and bladeless, more or less hirsute in the upper
part with tubercle-based hairs and densely ciliate along the upper margins,
nodes shortly bearded. Racemes 1 to about Q, in fascicles, born on filiform,
ultimately long exserted peduncles, 2°5-15 cm. long, very slender, flexuous ;
rhachis fragile ; joints 3°3 mm. long, usually shortly ciliate, but the uppermost
cilia sometimes up to 16 mm. long, sometimes glabrous or nearly so ; pedicels
reduced to a minute point or subule. Spikelets solitary, sessile, oblong-lanceo-
late in profile, somewhat oblique, laterally compressed, including the very
minute glabrous or minutely puberulous callus, 3°3-4°2 mm.long. Involucral
glumes subequal, lower subchartaceous, acute, scaberulous along the very
slender 7-9 nerves or almost smooth towards the base, upper obliquely lanceoiate
to linear-lanceolate in profile, acute or minutely mucronate, 3- (rarely 5-)
nerved, eciliate. Lower floral glume oblong, obtuse, hyaline, obscurely
2-nerved, shorter by %than the involucral glumes, upper narrowly linear-
lanceolate in profile, 2-2°7 mm. long, awn from near the base, usually 6°3 mm.
long, more rarely down to 4°2 or up to 8'5 mm. long, very delicate, kneed and
twisted below the middle. Stamens 2.
Locality: Gujarat: Chamargaon (Woodrow).—Khandesh: common
(McCann !).—Konkan : very common (McCann !).—Deccan . Mahableshwar
(Sedgwick and Bell 4501!) ; Panchgani (Blatter and Hallberg B1244!, B1252!,
B1257!, B1266!, B1273!, B1290!) Khandala, very common (McCann!).—
S. MM. Country: S.W. of Dharwar (Sedgwick and Bell 4434!) ; Dharwar
(Sedgwick 3098!) ; Gadag (Talbot 2304!) ; Belgaum (Ritchie 796A).— Kanara:
Halyal (Talbot 2161!) ; Yellapore (Talbot 1057 !).
Distribution: From Bihar southwards to Ceylon, Tropical Africa (Nileland,
Upper and Lower Guinea, Mozambique District), introduced into Jamaica and
Guadeloupe.
6. Arthraxon jubatus, Hack. Monogr. Androp. (1889), 358 ; Hook. f. in F. B. I.
vii, 147; Cke. ii, 970.
Description : Cke. l.c.
Locality : Konkan: (Law, Stocks).—Deccan: Kori Fort, 12 miles south of
Lonavla (Woodrow) ; Khandala, damp rocks (Hallberg 9788!) ; Lohagad Fort,
top (McCann 9789 !).
Distribution : W. Peninsula, apparently endeinic.
31. CAPILLIPEDIUM, Stapf in Fl. Trop. Afr. ix, 169.
Annual or perennial grasses. Stems slender, simple or branched, sometimes
very copiously, often bearded at the nodes. Leaf-blade with a rather con-
spicuous white midrib. Panicles delicate, when much divided the branchlets at
length more or less divergent. Spikelets small, 2-nate, one sessile, the other
pedicelled, similar in shape but differing in sex, in 1-2- (rarely up to 8-) jointed
racemes at the ends of the capillary primary and secondary and often tertiary
or even quaternary branches of a loose panicle; joints and pedicels finely fili-
form, longitudinally grooved and hyaline in the groove, disarticulating
horizontally ; sessile and pedicelled spikelets deciduous, the former with the
adjacent joint and pedicel. Florets 2 in the sessile spikelet, lower reduced to
420 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Wolo!
an empty glume or quite suppressed in the pedicelled spikelets, upper her-
maphrodite ; one male or neuter, in the pedicelled spikelet. Sessile spikelet
dorsally compressed, awned, callus small, shortly bearded. Involucral
glumes equal, membranous to subherbaceous ; lower 2-keeled, with narrow
inflexed margins ; upper boat-shaped, 3-nerved, keeled, grooved on both sides
along the obtuse keel. Lower floral glume hyaline, nerveless, upper consisting
of a hyaline, linear stipe, firmer upwards, passing into a slender awn. Pale0O.
Lodicules 2, minute, glabrous. Stamens 3. Stigmas exserted laterally, longer
than the styles. Grain oblong-ellipsoid or oblong, dorsally slightly compressed;
embryo exceeding zof the grain. Pedicelled spikelet awnless, glume, if
present, hyaline, nerveless.
Species 6, in tropical and subtropical Asia, Polynesia and Australia, tropical
Africa.
Three species, described by Cooke (ii, 981, 982) under Andropogon assimilis,
Steud., 4. Hugeli, Hack., and A. fzliculmis, Hook. f. belong to this genus.
I. Stems more or less suffrutescent below, stiff,
erect
1. Nodes of stem glabrous ; callus shortly
bearded ay oe as ELEC. SASSOUIZ01S »
2. Nodes of stem bearded ; callus densely
villous Pr a seseQan iG. ah woeltr.
Il. Stems decumbent and interlaced, very
weak, filiform... ie Semon “GC. 1tleculines.
1. Capillipedium assimile, A. Camus, Graminées in H. Lecomte. FI.
Générale de l’Indo-Chine 7 (1922) 314; Haines Bot. Bihar and Orissa
(1924), 1028 (.4. Camus prior fecit combinationem novam).—Andropogon
assimilis, Steud. in Zoll. Syst. Verz. (1854), 58; Syn. Gram. (1855), 397;
Hook. f. in F. B. I. vii, 179; Cke. ii, 981.—4. montanus, Benth. Fl. Hongk.
423, non Roxb., excl. syn.; Hack. Monogr. Androp. 490, excl. syn.— Chrysopo-
gon pictus, Hance in Ann. Sc. Nat. ser. 5, v (1866), 252.—C. glaucopsis, Duthie
Grass. N. W. Ind. 22.—Raphis repens, Nees ex Steud. l.c.
Description : Cke. 1. c.
Locality : Khandesh: Toranmal, growing among Strobilanthus (McCann
9671!).—Konkan: Above Kenery Caves (McCann 9959!).—Deccan: in dry
forest, between Mahableshwar and Panchgani, at 4,000 ft. (Sedgwick and Bell
4738 !).—Kanara: Jugglepet, road side, common (Talbot 1386 !).
Distribution : Temperate Himalaya, Khasia, Behar, N. Bengal, Rajputana,
Central India, W. Peninsula, Java, China, Japan.
2. Capillipedium Hugelii, Blatter & McCann, nov. comb.—Andropogon
Hugeliit, Hack. Monogr. Androp. 492 ; Hook. f.in F. B. I. vii, 180 ; Cke. ii, 982.
Description: Cke. l.c.
Locality: Khandesh: 'Toranmal (McCann 9672!).—Konkan: Mulgaum
(McCann 3664!).—Deccan: Khandala, St. Xavier’s Villa (McCann 9423!) ;
dry forests between Mahableshwar and Panchgani (Sedgwick 4738!) ;
Panchgani (Blatter and Hallberg B1321!, McCann!).—S. MW. Country:
Deciduous forests west of Dharwar (Sedgwick 4498!).—Kanara : Halyal (Tal-
bot 2082!) ; Ecumbi to Mungod (Hallberg and McCann A288!).
Distribution : Rajputana, Central Provinces, Central India, W. Peninsula.
3. Capillipedium filiculme, Blatter & McCann, w#ov. comb.—Andropogon
filiculmis, Hook. f. in F. B. I. vii, 181; Cke ii, 982.
Description : Cke.1.c.
Locality: Konkan: Trombay (McCann A286!).—Deccan. Khandala to
Karjat (Blatter and Hallberg A287!) ; Igatpuri (Blatter and Hallberg 5117 !) ;
Poona, in rocky places (Jacquemont 310) ; Donshi, Mawal Districts (Woodrow
26); Purandhar (McCann 5570 !).—S. MW. Country: forests near Dharwar
(Sedgwick 1854!).
Distribution + W. Peninsula, apparently endemic.
32. AMPHILOPHIS, Nash in Britt. Man. Fl. N.U.S. ed. i, 71;
Stapf in Fl. Trop. Afr. ix, 171.
Perennial grasses. Stems slender, simple or branched, bearded or beardless
at the nodes. Panicles mostly subdigitate with a short primary axis, rarely the
racemes on branches of the second order; racemes always shortly peduncled,
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 421
Spikelets small, 2-nate, one sessile, the other pedicelled, similar in shape or
the pedicelled reduced ‘and smaller, the latter always different in sex except
sometimes the lowermost pair which may be homogamous (male or neuter), on
the fragile rhachis of many-jeinted shortly peduncled racemes ; joints and
pedicels filiform, longitudinally grooved and hyaline in the groove, disarticu-
lating horizontally ; sessile and pedicelled (always ?) spikelets deciduous;
the former with the adjacent joint and pedicel. Florets 2 in the sessile
spikelets, lower reduced to an empty glume, upper hermaphrodite, 2 or 1 in
the pedicelled spikelet, the lower male or neuter, the upper neuter or usually
quite suppressed. Sessile spikelet dorsally compressed, awned ; callus small,
shortly bearded. Involucral glumes equal, thinly chartaceous to membranous ;
lower 2-keeled, with narrow sharply inflexed margins ; upper boat-shaped,
3-nerved, acutely keeled. Lower floral glume hyaline, nerveless, upper a
hyaline linear stipe, firmer upwards, passing into aslender awn. Pales 0 or
very minute. Lodicules 2, minute, glabrous. Stamens 3. Stigmas exserted
laterally usually low down, longer than the styles. Grain oblong, obtuse,
dorsally slightly conipressed ; embryo about half the length of the grain.
Pedicelled spikelet awnless, glumes, if present, hyaline, nerveless.
Species probably over 25, mostly in Tropical Asia.
Cke. (ii, 977-981) mentions 8 species which belong to tlie section
Amphilophis of Andropogon : Andropogon compressus, Hook. f., A. Wooa-
rowit, Hook. f., A. pertusus, Willd., A. Kuntzeanus, Hack., A. ensiformis,
Hook. f., A. concanensis, Hook.f., A. tutermedius, Cooke (von R. Br.),
A. odoratus, Dna. Lisboa. Allthese are now being transferred to the new
genus Amphilophis, Nash.
Key to the species (after Cke.)
A. Racemes digitate or fasciculate, the lower longer than the rhachis of
the inflorescence
I. Lower involucral gluine of sessile spikelets
villous below the middle
1. Lower involucral glume not pitted
(2) Upper involucral glume mucronulate ... 1. <A. compressa.
(6) Upper involucral glume obtuse 2. A. Woodrowit.
2. Lower involucral glumes pitted ... 3. A. pertisa,
II. Lower involucral glume of sessile spikelets
glabrous below the middie (sparsely silky
in A. enstfornits)
l. Nodes of stem densely bearded ww. 4. A. Kuntzeana.
2. Nodes of stem glabrous
(a) Leaves reaching 12 mm. broad ww. O. A. ensiformis.
(6) Leaves reaching 3 mm. broad 6. A. concanensis.
B. Racemes panicled, the lower branches shorter
than the rhachis of the inflorescence
I. Non-aromatic ; sheaths terete ee .. ##. A. glabra.
ie sAromatic. sheaths compressed ite . 8 A. odorata.
1. Amphilophis compressa, Blatter & McCann, an nov. Andropogon
compressus, Hook. f. in F.B.I. vii (1896), 172; Cke. ii, 977.
Description ; Cke. 1.c.
Locality : Deccan: Khandala, plain behind the Saddle (Hallberg 9657 !,
Se Lonavla (McCann 9433!) : Mawal (Wocdrow!) ; Poona (Wood-
row).
Distribution : Western Peninsula, apparently endeinic.
2. Amphilophis Woodrowii, A. Camus. Andropogon Woodrowti, Hook. f. in
Bal, vit (1896), 173); Cke. ii, 978.
Description : Cke. 1. ¢.
Locality : Deccan: Khorbasa, Mawal Districts (Woodrow); Pand, 20
miles W. of Poona (Woodrow !).
Distribution ; Western Peninsula, apparently endemic.
3. Amphilophis pertusa, Stapf in Fl. Trop. Afr. ix, 175; Haines Bot.
Bihar and Orissa 1030.—Hfolcus pertusus, Linn. Mant. Alt. 301. — Andropogon
pertusus, Willd. Sp. Pl. iv, 922; Beauv. Agrost. 131, t. 23, fig. 2; Roxb. Fl.
Ind. i, 258; Hack. Monogr. Androp. 479 (vars. genuinus et Wightit) Duthie
4
422 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
Grass. N.W. Ind. 21 (excl. Syn.), Fodd. Grass. N. Ind. 38, t. 25; Boiss. Fl. Or.
vy. 464; Balf: £. Bot. Socotra’ 316; -Hook. £..:F..B.I,: vii, 173 ;7Cke.il;"-978:=—
Lepeocercis pertusa, Nees ex Steud. Syn. Pl. Glum. i, 364.
Vern. Names: Ghanga, marvel (Satara, Sholapur, Poona), payen, palva,
alvan.
Description: Cke. ii, 978. A very variable plant. We must confess we
find it impossible to follow various authors who have described a number of
varieties. It would be easy to increase their number on merely morphological
grounds, but the results would be highly unsatisfactory. It is only from
genetical tests that we can expect to get an insight into the natural variations
of this and other species of this genus.
Locality: Gujarat: Perim Island, Gulf of Cambay (Blatter 3814 !).—
Konkan: Daman (Bhide!); St. Xavier’s College compound, Bombay
(McCann 9630!, 9631!).—Deccan: Nasik (Bourke 11!); Rahuri (Nana
A278!) ; Khandala, Campoli (McCann 9961!) ; Pashan (Gammie!) ; Lonavla
(Woodrow) ; Agricultural College, Kirkee (Bhide!); Poona (Cooke) ;
Panchgani (Blatter and Hallberg B1236!), on edge of Tableland (Blatter
9962!), on roadside 4,000 ft. (Sedgwick and Bell 4699!) ; Satara (Lisboa) ;
Sholapur (Lisboa) ; Joonur (Talbot!).—S.47. Country: W. of Dharwar,
elevation 2,000 ft., rainfall 35” (Sedgwick and Bell 4494 !} ; Dharwar, elevation
2,400 ft., rainfall 34” (Sedgwick and Bell 4488!) ; Kunur, elevation 2,000 ft.,
rainfall 35” (Sedgwick and Bell 4953!) ; Badami (Talbot 2944!) ; Haveri
iain 2233!); Gokak (Sheodye!).—Kanara: Halyal (Talbot 2080!,
2106 !).
Distribution: More or less throughout India, chiefly in the drier parts,
Ceylon, Afghanistan, Arabia, tropical Africa (Upper Guinea, Nileland,
Mozambique District).
Uses: A good fodder grass. (See W. Burns, L. B. Kulkarni and S. R. God-
bole: A study of some Indian grasses and grasslands. Mem. Dept. Agr. in
India, xiv (1925), 47, 48.)
4. Amphilophis Kuntzeana, Haines Bot. Bihar and Orissa 1031.— Andropogon
Kuntzeanus, Hack. Monogr. Androp. 478; Hook. f. F.B.I. vii, 175 ; Cke. ii,
979;
Descripion «| Cke. lec.
Locality : Konkan: (Stocks).—Deccan » (Woodrow 153!) ; Mawal (Wood-
row).
Distribution : Bihar, Central Provinces, W. Peninsula.
5. Amphilophis ensiformis, Blatter & McCann, comb. nov.—Andropogon ensi-
formis, Hook. f. in F. B. I. vii, 175 ; Cke. ii, 979.
Description: Cke.1.c.
Locality : Deccan: Lonavla (Woodrow !).
Distribution: W. Peninsula, apparently endemic.
6. Amphilophis concanensis, Blatter & McCann, comb. nov.—Andropogon
concanensis, Hook. f. in F. B. I. vii, 174; Cke. ii, 980.
Description: Cke.l.c.
Locality : Konkan: Matheran (Woodrow !).—Deccan : Yenna River, Ling-
mala, Mahableshwar, 4,000 ft. elevation, 200” rainfall (Sedgwick and Bell
4652!) ; Tingmala, Mahableshwar (Blatter and Hallberg B1328 !) ; Khandala,
in water courses, common (McCann 9651!) ; Manmad, river-bed (Blatter
A283 !) ;—Kanara: Kalanadi River, Supa, on rocks, elevation 2,000 ft., rainfall
100 ” (Sedgwick and Bell 4857!) ; Halyal (Talbot 2221!) ; Goond (Talbot
2202!) ; Gersoppa Falls, on rocks in river bed (Hallberg and McCann A279 !).
Distribution : W. Peninsula, apparently endemic.
7. Amphilophis glabra, Stapf in Fl. Trop. Afr.ix, 172.—<Andropogon glabver,
Roxb. Fl. Ind. i, 271; Steud. Syn. Pl. Glum. i, 392.—A. punctatus, Trin.
Ic. t. 328, non Roxb.—A. intermedius var. punctatus, subvar. glaber, Hack.
Monogr. Androp. 487.—A. intermedius, K. Schum. in Engl. Pfl. Ost.-Afr.
C. 98; Hook. f in F. B.I. vii, 175; Cke. ii, 980.—A. imutermedius, var.
punctatus, Stapf in Dyer, Fl. Cap. vii, 345.—Amphilophis glabra. Haines
Bot. Bihar and: Orissa 1028, zon Stapf and partim.
It will be seen from the above synonymy that Stapf considers Andropogon
glaber, Roxb. as the type of the species. Of Andropogon intermedius, R.Br. as
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 423
conceived by Hackel he includes only the var. punctatus, subvar. glaber. Haines
in his Bot. Bihar and Orissa, 1028, adopts Stapf’s name Amphilophis glabra with
the following synonyms: Andropogon intermedius, R. Br. inc. A. glaber,
Roxb., A. punctatus, Roxb. and A. montanus, Roxb. ?
This is evidently not Stapf’s Amphilophis glabra. WHaines includes Andro-
pogon intermedius, R. Br., whilst Stapf confines his species to var. punctatus,
Haines has A. punctatus, Roxb. as asynonym, Stapf excludes it expressly.
Haines’ description is much wider than that given by Stapf and includes
Hackel’s vars. genuinus, Henket, punctatus and glaber, and one of his own
var. hirta. He says in a foot-note (p. 1029) : ‘ These varieties are after Hackel
and were described before I consulted Stapf’s Graminez in Fl. Trop. Africa,
where he reduces Hackel’s zztermedius to Roxburgh’s glaber. Thenames (e.g.
genuinus) in some cases become inapplicable if Roxburgh’s gladver is the type.
Var. Henketz is, | think, a distinct species and easily distinguished in the field.
It should be called Amphilophis Henke.’
Haines seems to overlook the fact that Stapf’s glabra is restricted to one of
Hackel’s varieties of Andropogon intermedius, viz., punctatus and that, conse-
quently not all of Hackel’s names can be applicable.
In our opinion Haines is not justified in calling his species Amphilophis
glabra, Stapf. As it includes practically the whole of Andropogon intermedius,
R. Br. taken in Hackel’s sense he might have called it Amphilophts intermedia,
Haines, if that name had not been preoccupied by Stapf (Fl. Trop. Afr. ix,
174).
We are not arguing the point whether Stapf or Haines is right in the
treatment of Andropogon intermedius, R. Br. et auctorum ; good reasons can
be adduced for both cases. All we wish tosay is that Haines’ Amphilophis
glabra is not A. glabra, Stapf.
Those who prefer to adopt Haines’ A. glabra and Hackel’s varieties might
consider a remark made by Hackel himself (l.c. 487): ‘ 4. fascicularts
Thwattes Enum. Pl. Zeyl. p. 437 non Roxb. complectitur varietates B
[Henket] et 8 [punctatus| formis inlermedits (spiculis in eadem panicula
mixtis) conjunctas. Etiam in Bengalta transitus et ipsa varietas B [Henket]
proventunt ; in Himalaya,e. gr. pr. Simlah forme inter § [punctatus] et
oc [genuinus| intermedie inveniuntur.’
We adopt Stapf’s conception of 4. glabra together with his description. Dr.
Stapf was kind enough to name some of our Bombay specimens,
Description: Perennial. Rhizome very short, hard, innovations extravagi-
nal, cataphylls ovate to lanceolate, acute, hard, smooth. Stems tufted, erect
or shortly ascending, to overlm. high, rather stout below, glabrous, 5-7-
noded, simple or very sparingly branched. Leaf-sheaths terete, glabrous,
smooth, the intermediate and upper mostly shorter than the internodes, nodes
glabrous or the upper bearded. Blades linear, long-tapering to a setaceous
point, hardly broader, not or slightiy contracted at the base, up to over 30 cm.
by 8°5 mm. (mostly narrower), pale green, often turning reddish or purplish,
glabrous, rarely with very fine, long, spreading, white hairs at and above the
ligule, smooth except at the margins, midrib rather stout to very stout down-
wards, lateral nerves 3-4 on each side ; ligules truncate, very short, scarious.
Panicle narrowly oblong, 10-23 cm. by 25-38 mm., erect, primary axis much
longer than the lowest racemes, somewhat stout and (at least when mature)
stiff, smooth, shortly bearded at the branch axils, otherwise glabrous and
smooth ; branches whorled, up to 6 in a whorl or semiverticillate, or 2-nate or
solitary from the weaker nodes, straight, 25-75 mm. long, the longer divided
from 12 mm. above the base, few to 7-noded, secondary branches simple.
Racemes 12-38 mm. long, straight or flexuous, usually purplish, inconspi-
cuously white-villous ; joints and pedicels 2-3°3 mm. long, ciliate, uppermost
cilia much longer than the rest, up tol mm. long. Sessile spikelet pale green
or purplish throughout, including the small minutely bearded callus 3°3-3°8
mm. long. Involucral glumes equal; lower truncate, slightly concave on the
back, chartaceous-membranous, hairy below the middle, more rarely glabrous,
keels rigidly ciliolate upwards, intracarinal nerves 4-5, very fine, evanescent
upwards with or without a pit above the middle: upper lanceolate, acute, 3-
nerved, keel scabrid upwards, margirs sparingly ciliate. Lower floral glume
oblong, 2°7 mm. long, hyaline, nerveless, ciliate, upper an awn 6:3-]2°7 mm.
long, brown below, pale above the bend. Pedicelled spikelet neuter, mostly
reduced to the glumes, narrowly linear-oblong to linear, 2°7-3'3 mm, long,
424 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
glabrous, of the same colour as the sessile. Lower involucral glume rigidly
and minutely ciliolate, 5-9-nerved, often rolled in, not pitted, upper hyaline,
nérveless, often minute.
Locality : Konkan: Penn (McCann A282 !).—Deccan : Pimpalgam, on the
brink of astream (Bhide!) ; Lonavla (Hallberg 9955!) ; Khandala, St. Mary’s
Ravine, on a water course (McCann 9435 !).
Distribution : More or less throughout india, Tropical Asia, N.E, Australia,
Tropical Africa (Upper and Lower Guinea, Mozambique District, Nileland)
Madagascar,
8. Amphilophis odorata, A. Camus. <Azdropogon odoratus, Dna. Lisboa in
Journ. Bomb. Nat. Hist. Soe. iv, (1889), 123, cumic. and vi (1891), 68,
203; Hook.f. in F.B,I. vii, 177; Cke. ii, 981.
Description : Cke. l.c.
Locality : Khandesh (Lisboa).—Konkan : Khardi (Mrs. Lisboa).—Deccan :
Igatpuri (McCann A281] !), Lonavla ; (Mrs. Lisboa, Bhide!) ; plain at foot of
Lohagad (McCann 9956!) ; Pand, 20 miles W. of Poona (Herb. Econ. Bot.
Poona!) ; Mawal, Poona District (Woodrow) .
Distribution : W. Peninsula, apparently endemic.
33. DICHANTHIUM, Willemet in Usteri Ann. xviii (1796), 11;
Stapf in Prain Fl. Trop. Afr. ix, 177.
Perennial or annual. Stems simple or branched, usually many-noded,
bearded or beardless at the nodes. Panicles usually subdigitate with a short
or very short primary axis, rarely the latter elongated ; racemes always shortly
peduncled. Spikelets small, 2-nate, one sessile, the other pedicelled, similar
in shape, different in sex, except the lowermost 1 or 2 pairs (sometimes 3 or 4)
of each raceme which are (with occasional exceptions in 0. annulatum)
homogamous (male or neuter), in many-jointed, shortly peduncled, subdigi-
tate, rarely subpanicled or racemosely arranged, racemes ; joints and pedicels
filiform, solid, disarticulating subhorizontally except the lowest barren pairs ;
fertile sessile and pedicelled spikelets deciduous, the former with the adjacent
joint and pedicel. Sessile spikelet dorsally compressed, awned except the
basal homogamous ones; callus small, shortly bearded. Involucral glumes
equal, thinly chartaceous, lower usually very obtuse, 2-keeled, with narrow
sharply inflexed margins, upper boat-shaped, 3-nerved, acutely keeled.
Lower floral glume hyaline, nerveless, upper reduced to the hyaline base of a
slender awn; pale minute or absent. Lodicules 2, minute, glabrous.
Stamens 3. Stigmas exserted laterally at or above the middle or near the tips.
Grain oblong, obtuse, dorsally compressed ; embryo rather more than half the
length of the grain. Pedicelled spikelet awnless. Lower involucral glume
oblong, many nerved, upper flat with sharply inflexed margins closing over the
hyaline lower floral glume if present and the stamens, upper floral glume
usually 0, never awned.
Species 10 in the tropical and warm-temperate regions of the Old World.
So far 4 species were known from the Presidency and described by Cooke
under Andropogon armatus, Hook. f., A. Cooket, Stapf, A. caricosus, Linn.
and 4. annulatus, Forsk. To these we have added 2 new species:
Dichanthium panchganiense, Blatter & McCann, and DD. McCannit,
Blatter.
Key to the species
A. Racemes digitate (sometimes solitary in
D. earicosus), 25 mm. long or longer
I, Lower involucral glumes of pedicelled
spikelets armed with marginal bul-
bous- based bristles
1. Lower involucral glume of pedicelled
spikelet always pitted 1. D. pancthganiense.
2. Lower involucral glume of pedicelled
spikelet not pitted
(a) Ligulé absent ..; a .. 2. 0. armatum.
(6) Ligule present ... ini ww 3. D. MeCannit.
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 425
Il Lower involucral glumes of pedicelled
spikelets not armed with marginal
bristles
1. Nodes of stem glabrous; Jigule a
shortly ciliate small membrane 3. 6 4. DD. Cavitésam:
2. Nodes of stem bearded ; ligule large,
membranous : » 8S. D. aennulatum.
B. Racemes solitary, less than 25 mm. long... 6. D. serratalccides.
1. Dichanthium panchganiense, Blatter & McCann in Journ. Bomb. Nat
Hist.so0c, xxxit (1927), 381.
Locality: Deccan: Panchgani (McCann!)
Distribution: W. Peninsula, apparently endemic.
2. Dichanthium armatum, Blatter & McCann, comb. nov.—Andropogon
armatws, Hook. f. in F. B: I. vii, 197: Cke. ii, 987.
Description: Cke. 1. c.
Locality : Konkan. Stocks (teste Hook. f.).—Deccan: Kalsabai Hills,
Nasik District (Patwardhan 1183 !); Gira Hill, Khandala (McCann 9430 ! 9431 !);
Lohagad (McCann 3871!); Panchgani (Blatter and Hallberg B1212!) ;
Pasarni Ghat (Blatter and Hallberg B1304!).
Distribution : W. Peninsula, apparently endemic.
3. Dichanthium McCannii, Blatter in Journ. Bomb. Nat. Hist. Soc. xxxii
(1927), 381.
Locality : Deccan: Panchgani (McCann ! ).
Distribution: W. Peninsula, apparently endemic.
4. Dichanthium caricosum, A. Camus: Note sur le genre Dichanthium
Willemet in Bull. Mus. Hist. Nat. (Paris) 27 (1921), 549.—Andropogon cart-
cosum, Linn. Sp. Pl. ed. 2 (1763), 1480; Hack. Monogr. Androp. 567 ; Hook.
f. in F. B. I. vii, 196; Cke. ii, 937; Haines Bot. Bihar and Orissa, 1039.—For
further synonyms see Hook. f.l.c.
Three authors have made the new combination Dichanthium caricusum :
(a) A. Camus. c. (6). Stapf in Ridley Fl, Malay Penins. v (1925), 210. (c)
Haines in Bot. Bihar and Orissa (1924), 1039. A. Camus has therefore to be
adopted as authority for D. caricosinm.
We are not trying to describe varieties or forms of this very variable species.
Vern. Names: Marvel (Mar.), Zinzvo (Guj.).
Description : Cke.l.c. :
This species can, according to Burns and others* be distinguished from
D. annulatum, Stapf by the following characters :—
Dichanthium cavicosum. Dichanthium anniulatum.
Habit : Big and tufted with ter- Medium size with termi-
minal and axillary in- nal inflorescences.
, florescences.
Nodal hairs on stem : Short. Long.
Colour of inflorescence: Light purplish-green. Purple.
Hairs of inflorescence: Short. Long.
Haines |. c. believes there is no single character by which DY. cari€osum can
be distinguished from 1). annulatum. ‘1 have,’ he says, ‘ tested all those given
in the F. B. I. and found them fail on specimens named by Sir J. D. Hooker
himself ; the key characters in Bengal plants are also unworkable as applied to
the same specimens, many of which have bearded nodes, and the character of
spiral or subdistichous spikelets is difficult to apply. Linnaeus described
Andropogon Caricosus .as with solitary spikes, and Willdenough (sic!) adds
‘leaves with sparse hairs and sheaths hirsute at the base’ (probably he refers
to the nodes).’
In his key-characters Haines says that in J. caricosum tne callus is glabrous.
This seems to be amistake. Hackel calls the callus Brevissime barbatus, Cooke
has ‘ bearded,’ Rangachari describes it as ‘ short and shortly hairy below.’ We
have always found it bearded.
Locality : Gujarat: Surat, road sides (Sedgwick 315!).—Khandesh ;
Toranmal (McCann 9669!) ; Tapti, N. E. of Bhusawal (Hallberg 5111 !) ;
* W. Burns, L. B. Kulkarniand S. R. Godbole : A Study of Some Indian
Grasses, in Mem. Dept Agr. Ind., xiv (1925), 46.
496 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
N. slope of Chanseli (McCann 9968! ).—Konkan.: Parsik, railway tract
(McCann 9633!).—Deccan: Junnar (Mamlatdar of Junnar!) ; Shevgaon
(Mamlatdar of Shevgaon!); Lohagad, half-way up (McCann 9627!) ;
Agricultural College Farm, Poona (Herb. Econ. Bot. Poona!) ; Khed
(Mainlatdar of Khed!) ; Purandhar (McCann 5570!) ; Bairawady, Purandhar
(McCann 5075! ) ; Panchgani (Blatter 3803! , Blatter and Hallberg Bi223! ) ;
between Mahableshwar and Panchgani, elevation 4,000 ft. (Sedgwick and
Bell 4743;).—S. J7. Country: Dharwar, elevation 2,400 ft., rainfall 34”
(Sedgwick 1828!).—Kanara: Halyal (Talbot 2427 !).
Distribution : India, Burma, Ceylon, Mauritius, China.
Uses: A good fodder grass.
5. Dichanthium annulatum, Stapf in Prain Fl. Trop. Afr. ix, 178; Haines Bot.
Bihar and Orissa 1039.—Andropogon annulatus, Forsk. Fl. Aegypt.-Arab,
(1775), 173 ; Duthie Fodd. Grass. N. Ind. 33, t. 20 ; Hack. Monogr. Androp. 570 ;
Hook. f. F.B.I. vii, 196; Cke. ii, 988.~ For further synonyms see Hook. f. 1. c.
Vern. Names: Zinjoo, handi daroya, daroya (Surat), dhrow (Broach),
zinzma (Charodi), jinjva (Panch-Mahals), marvel (Poona), sheda, sam-payen
palvan-hullu, marwalyan-hullu (Dharwar).
Description: Cke. 1. c.
Locality : Sind: Jamesabad (Sabnis B972!); Mirpurkhas (Sabnis B1028!,
B1185!) ; Hyderabad (Sabnis B51!); Pad-Idan (Sabnis B515!); Larkana
(Sabnis B462!) ; Nasarpur (Sabnis B1140!) ; Tatta, Kullan Kote Lake (Blatter
and McCann D667!) ; Tatta, tombs (Blatter and McCann D668 !) .—Gujarat :
Kabirwad (Chibber!) ; Nadiad Farm (Herb. Econ. Bot. Poona!) .—Khandesh :
Amboli, Bori River (Blatter and Hallberg 5148!) ; Dadgaum (McCann 9665!) ;
Toranmal (McCann 9670!) ; Bor, Bori River (Blatter and Hallberg 4428! );
Umalla, Tapti Bank (Blatter and Hallberg 5228! ).—Kunkan : Sion (McCann
3672 !) ; Sewri (McCann 3587!) ; Parsik, railway line (McCann 9654!) ; Cam-
poli (McCann 5356!).—Deccan : Kirkee to Poona, railway line (Garade 83!) ;
Cellege Garden, Poona (Garade!); Chattarshinji Hill, Poona (Ezekiel) ;
Trimbak (Chibber!) ; Khandala, very common (McCann 5297!); Manmad
Blatter 9970!) ; Purandhar, north foot (McCann 9421!).—S.M/. Country :
Devikop, elevation 1,800 ft., rainfall 40” (Sedgwick 3984 !).—Kanara - Halyal
(Talbot 2081 !) ; Kulgi (Talbot 2299!).
Distribution - Tropical Africa (Nileland, Mozambique District), from
Morocco through North Africa, the Orient and India to China and Australia,
Pacific Islands.
Uses ; Considered good fodder.
§. Dichanthium serrafalcoides, Blatter & McCann, xov. comb.—Andropogon
Cooket, Stapf ex Woodrow in Journ. Bomb. Nat. Hist. Soc. xiii, 438 (omen
tantum) ; Cke. ii, 986 (descriptio).—A.( Dichanthium ?) serrafalcoides, Cooke et
Stapf in Kew Bull. (1908), 450.
The systematic position of this species is somewhat doubtful. Cooke already
found that it does not agree with Hackel’s subgenus Dichanthium, but he adds:
‘It is the only subgenus into which it will fit at all.’ Cooke and Stapf, when
describing the same species under a different name, remark : ‘Ob spiculas secun-
darias intimas saepissime neutras caeterum eadem forma ac fertiles si vis ad
Dichanthiun referendus, sed nulli speciei arcte attinis spiculis maiuscults in
vacemos spiculis Serratalct haud dissimiles congestis insignis.’ Following
this suggestion we have put it under Dichanthium.
Description : Cke. ii, 9s6.
Lotality: Deccan : Lonavla (Bhide !); Khandala, Echo Point (McCann 9403!) ;
Kalanbai Hills (Patwardhan !); Sakar-Pathar near Lonavla (Woodrow !) ;
Mahableshwar (McCann!); Panchgani (McCann!, Blatter and Hallberg
B1250 !).
Distribution: W. Peninsula, apparently endemic.
34. EREMOPOGON, Stapf in Prain Fl. Trop. Afr. ix (1917), 182.
Perennial or annual grasses. Stems slender, simple below, more or less
branched above, the branches often gathered in fastigiate bundles, each sup-
ported by a bladeless sheath and terminated bya solitary raceme. Spikelets
small, 2-nate, one sessile, the other pedicelled, similar in shape, different in
sex, on the fragile rhachis of many-jointed solitary spatheate racemes which are
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 427
frequently gathered in fastigiate bundles, rarely the lowermost 1-3 pairs homo-
gamous ; joints and pedicels filiform, compressed, solid or slightly grooved,
disarticulating horizontally ; spikelets deciduous, the sessile with the adjacent
joint and pedicels. Sessile spikelet dorsally compressed, awned ; callus small,
shortly bearded. Involucral glumes equal, thinly chartaceous to membranous,
lower 2-keeled, with narrow inflexed margins, uppet boat-shaped, 3-nerved,
acutely keeled. Lower floral glume hyaline, nerveless, upper reduced to a hya-
line upwards firmer linear stipe passing into a slender awn. Stamens 3. Stig-
mas exserted laterally nearthe middle of the spikelet, longer than the styles.
Pedicelled spikelet awnless ; soniewhat similar to the sessile.
Species about 5, in the tropical and warm-temperate parts of the Old World.
Cooke knew one species from the Presidency which he described under the
name of Andropogon foveolatus, Del. Weadd Fremopogon Paranjpyeanum,
Blatter and McCann.
1. Lower involucral glume of sessile spikelet 4-nerv-
ed... a ane ee w. 1l. £. foveolatus.
2. Lower involucral glume of sessile spikelet faintly
5-7-nerved ... ticle iss w. 2. L. Paranjpyeanum
1. Eremopogon foveolatus, Stapf in Prain Fl. Trop. Afr. ix (1917), 183.—
Andropogon foveolatus, Del. Fl. Egypte 16, t. 8, fig. 2; Duthie Fodd. Grass.
N. Ind. 35, t. 22; Hack. Monogr. Androp. 402; Hook. f. F. B.1. vii, 168;
Cke. ii, 977.—A. strictus, Roxb. Fl. Ind. i, 260.—A. monostachyus, Spreng.
Pug. ii. 9.—Aypogynium foveolatum, Haines Bot. Bihar and Orissa (1924),
1041.
Description: Cke.l.-c.
Locality : Sind: Sehwan to Laki, foot of hills (Sabnis B 651!).—Gujarat -
Road to Lasandra (Chibber !) ; Bhuj Hill, Cutch (Blatter 3765 !).—Khandesh :-
Umalla, Tapti Bank (Blatter and Hallberg 5222! ); Bhusawal (McCann 4243 !);
Bor, Bori River (Blatter and Hallberg 4309!) ; Naradana (Blatter and Hallberg
5180 ;).—Konkan: Parel, poor specimen (McCann 5373 !).—Deccan.: Pan-
chgani, roadside, elevation 4,000 ft., rainfall 60” (Sedgwick and Bell
4735!) ; Yeola (Herb. Econ. Bot. Poona!) ; Arangaon, Ahmednagar (Ryan !) ;
Chattarshinji Hill (Ezekiel!) ; Deolali (Blatter 4550!) ; Manmad (Blatter
A261!) ; Panchgani (Blatter and Hallberg B1245!).—.S. 7. Country : Dharwar,
elevation 2,400 ft., rainfall 34 ” (Sedgwick 1825 !) ; Haveri (Talbot 2229 !).
Distribution: Tropical Africa (Upper Guinea, Nileland), Canaries, from
Egypt and Tropical Arabia to the drier parts of India.
2. Eremopogon Paranipyeanum, Blatter & McCann, comb. nov.—Andro-
pogon Paranjpyeanum, Bhide in Journ. and Proc. Asiat. Soc. Beng., New
series, vii (1911), 514.
Description: A delicate-looking grass. Stems slender, erect, 30-45 cm.
high ; upper nodes pubescent; leaves 2°5-7°5 cm. by 2mm., subcordate at
base, long-hairy on both sides, the margins thickened and minutely irregularly
repand and spinulosely serrulate; sheaths glabrous; ligule a short erose
membrane. Racemes solitary, 12-25 mm. long (without the awns), on a very
slender peduncle. Sessile spikelets 3 mm. long. Lower involucral glume
oblong, obtuse, faintly 5-7-nerved, glabrous, margins narrowly incurved, keels
shortly ciliate at the apex, upper just a little longer than the lower, 3-nerved,
oblong, apiculate. Lower floral glume shorter than the involucral glumes,
hyaline and with ciliate margins, epaleate, upper floral glume consisting of the
narrow base of the awn, just a little more than half as long as the lower in-
volucral glume, obscurely margined and 1-nerved with 2 obscure narrow lobes
at the apex, and an interposed slender, twisted, scabrid awn about 4 cm. long,
bearing a bisexual flower. Pedicelled spikelets about 4mm. long. Lower
involucral glume oblong, obtuse, 7-9-nerved, margins incurved and broadly
winged at the keels, wings shortly ciliate towards the apex, upper involucral
glume a little shorter than the lower, oblong, acute, 3-nerved, margins ciliate.
Lower floral glume shorter than the upper involucral, hyaline, ciliate, faintly
3-5-nerved, epaleate, male. Joints and pedicels compressed, obliquely truncate,
3-3 as long as thesessile spikelets and ciliate with short white hairs on both
sides,
Locality : S. M. Country : Castle Rock (Bhide !).
Distribution: So far endemic,
428 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXX
35. SCHIZACHYRIUM. Nees Agrost. Bras. 331. Stapf in Prain
Fl. Trop. Afr. ix, 1184.
Annual or perennial grasses, rarely suffrutescent, never very tall. Stenis
slender. Leaf-blades folded in bud, usually narrow. False panicles varying
from very loose and scanty to densely fascicled ; spathes mostly very narrow,
scarious, membranous or lower down herbaceous. Spikelets 2-nate, of each
pair differing in sex and mostly also more or less in shape and size, one sessile
the other pedicelled, on the articulate fragile rhachis cf many-jointed solitary
racemes terminating the culms and their branches, supported by spathes and
often collected into a false panicle, the sessile spikelets falling with the conti-
guous joint and the accompanying pedicelled spikelet ; joints and pedicels
thickened upwards, often rather stovt with ascarious cupuliform and more or
less dentate terminal appendage. Sessile spikelets dorsally compressed or some-
times in their lower half almost terete, awned ; callus short with a short beard
at the very base. Involucral glumes equal or subequal, lower chartaceous to
subcoriaceous, more or less convex or flat on the back with, at least from the
middle upwards, sharply inflexed and mostly narrow margins, 2-keeled, the
keels running out into teeth or mucros, upper thinner to membranous, nar-
rowly boat-shaped to dorsally flattened, keeled (at least upwards), 1-3-nerved,
the delicate margins ciliolate Floral glumes, ciliolate, hyaline, lower mem-
branous downwards and often purplish, 2-nerved, upper usually 2-fid or 2-den-
tate, rarely entire, awned, awn from the sinus or continuing the entire valve,
Pale 0 or a microscopic hyalinescale. Lodicules 2, minute, glabrous. Stamens
3, rarely 2. Stigmas laterally exserted low down; styles terminal. Grain
narrowly linear in outline or tapering upwards, subterete; embryo short.
Pedicelled spikelet similar to the sessile, but usually relatively broader and
flatter, or more or less reduced and then sometimes quite small. Involucral
glumes more or less membranous, the lower aristulateor muticous. Floral
glumes, if present hyaline, ciliate, muticous.
Species about 50, in the tropics of both hemispheres.
1. Schizachyrium brevifolium, Nees Agrost. Bras. 332; Miq. FI. Ind. Bat. iii,
495; Stapf in Prain Fl. Trop. Afr. ix, 187; Haines Bot. Bihar and Orissa 1042.
— Andropogon brevifolius, Sw. Prodr. Veg. Ind. Occ. 26, Fl. Ind. Occ. i, 209;
Hack. in. Monogr. Androp. vi, 363 (excl. var. fragilis) ; Duthie Grass. N. W.
Ind. 19, Fodd. Grass. N. Ind. 34; Hook.f F.B.I. vii, 165.—A. obtusifolius,
Poir. Encycl. Suppl. i, 583.—d. parvifolius, Roxb. Fl] Ind. i, 277.—A. tenellus
Pres]. Rel. Heenk. i, 335.-—-A. dedilis, Kunth Enum. Pl.i, 488.—A. floridus,
Trin.in Mem. Acad. Petersb. 6 ser. ii, 265.—FPollinia brevifolia, Spreng. Syst,
i, 288.
Description: Annual. Stems weak, up to60cm. long, usually ascending
from adecumbent base, rarely erect many-noded, branched from most of the
nodes: branches often 2-4-nate and dividing again, very slender to filiform,
geniculate, glabrous, the lower internodes usually much compressed. Leaf-
blades linear, constricted at the junction with the sheath, the lower and those of
the primary branches obtuse or sub-obtuse or suddenly narrowed to a sharp
point, 25-38 mm. long and 2-6'3mm. wide, the upper and those of the secon-
dary and tertiary branches much narrower and more acute, often glaucescent,
smooth except along the margins and the lower side of the midrib, nerves very
fine. Ligules membranous, very short, ciliolate. Sheaths compressed, the
lower keeled, somewhat lax, glabrous, smooth, the uppermost spathe-like ;
leaves like the spathe of the inflorescence ultimately turning reddish. Spathes
on subcapillary curved or flexuous branchlets, very narrow, acute, reaching to
the base of the racemes or slightly exceeding thei ; racemes borne on filiform
upwards clavate peduncles, slender from less than 12mm. to slightly over
25 mm. long, 5-11l-jointed ; joints 2-2°7 mm. long, widening upwards from a
slender base, tips 2-dentate glabrous, smooth ; pedicels as long as the joints,
very slender and only slightly thickened upwards. Sessile spikelets linear-
lanceolate, acuminate, more or less convex on the back, pale green, about
3°3 mim. long including the minute callus, which is more or less shortly bearded,
at least on the sides. Involucral glumes equal. lower thinly chartaceous, 2-
dentate, scaberulous along the keels, with very tine intracarinal nerves, smooth
or very minutely scaberulous on the back ; upper boat-shaped, acute, l-nerved,
ciliolate. Lower floral glume elliptic, obtuse, hyaline, nerveless, ciliolate,
upper 2-fid almost to the base, 1'6 mim, Jong, segments linear-oblong, sub-
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 429
obtuse, sparingly ciliolate, awn 8°5-12°7 mm. long. Anthers 0'5mm. long, deep
redor purple. Grain sublinear, tapering upwards. Pedicelled spikelet reduced
toa minute glume, often passing indistinctly into the pedicel and produced into
a bristle-like awn, about 4:2 mm. long.
Locality : S. M. Country : Badami (Talbot 2930 !).
Distribution : Widely distributed throughout the tiopics.
36. ANDROPOGON, Linn.; Stapf in Prain Fl. Trop. Afr. ix, 208.
After the restoration of Hackel’s subgenera to the rank of genera, the general
characters of Andropogon itself must be modified in many ways:
Mostly perennial grasses of various habit. Spikelets 2-nate, the sessile and
pedicellea differing trom one another in sex and more or less heteromorphous, all
pairs similar, or if the lowest sessile spikelet male or imyerfect then resemb-
ling the others. Spikes (spiciform racemes) many-jointed, fragile. paired (very
rarely solitary) or corymbose (digitate or subdigitate) on terminal pedurcles,
embraced below by a spathe-like leaf (spatheole), frequently 2 or more pairs
with their spathes subtended by a common spathe and soon to more compo-
site branching, the whole forming a false panicle; the sessile spikelets falling
with the contiguous joint and the accompanying pedicel; joints and pedicels
filiform or thickened upwards and then the tips frequently more or less cupular
or auricled Sessile spikelets dcrsally or iaterally compressed, rearly always
awned ; callusshort, mostly quite obtuse, shortly bearded. Involucral glumes
equal or subequal, subcoriaceous to membranous, lower flat or coucave or
channeled on the back. withatleast from the middle upwards sharply inflexed
margins, 2-keeled ; upper more or less boat-shaped, keeled upwards, 3-1-nerved,
sometimes aristulate. Flcral glumes ciliate or ciliolate, rarely glabrous, lower
hyaline, 2-nerved, upper 2-fid or 2-dentate, hyaline or firmer and some-
times substipitiform below the insertion of the awn. Pale a hyaline nerveless
scale or 0. Lodicules ?, minute, glabrous. Stamens 3. Stigmas lateially
exserted ; styles terminal. Grain narrowly lanceolate to oblong in outline,
subterete to planoconvex ; embryo about half the length of the grain. Pedicel-
led spikelets often very different from the sessile in shape and less soin size,
always miore or iess compressed dorsally, never concave or channelled on the
back, sometimes reduced and then often small or quite suppressed. Involucral
glumes herbaceous-chartaceous to niembranous, the lower muticous or very
rarely aristulate. Floral glumes, if present, hyaline. ciliate, muticous.
Species about 100, mostly in the tropics of both hemispheres.
1. Andropogon pumilus, Roxb. Fl. Ind. 1 (1832), 273; Steud, Syn. Gram. 388 ;
Hook f.in F.B.1. vii, 170; Cke. ii, 976; Haines Bot. Bihar and Orissa 1044.—
A. demzssus, Steud, l.c.—A. pachyarthrus, Hack. Morvogr. Androp. 449;
Duthie Grass. N. W. Ind. 21, Fodd. Grass. N. Ind. t 39.
Vern. Names: Zinzvo (Surat), berki, gondwal, la!lgavat, tambrut, gondal,
chiman chara, malakava.
Description » Cke. l.c.
Locality: Gujarat. Nadiad farm (Herb. Econ. Bot. Poona!) ; Surat
(Dalzell).—Ahandesh ; Bhusawal (McCann 5451!)! Bor, Bori River (Blatter
and Hallberg 5116!) ; to Naradana (Blatter and Hallberg 5207!).—Deccan -
Pashan (Gammie!) ; Bairawadi, Purardhar (McCann 5054!) ; Manmad (Blat-
ter A262!); Shinda (Sabnis A260!); Panchgani (Blatter and Hallbeig
B1268 !, B1274!, B1326!).—S. 7. Country: Haveri (Talbot 2230!) ; Dumbal
(Talbot 2318!) ; Harsol (Sedgwick 1083!) —Kanara : Karwar (Lisbea).
Distribution : Nepal, Bihar, Rajputana, Central P:ov., W. Peninsula.
37. CYMBOPOGON, Sprevg.; Stapf in Prain Fl. Trop. Afr. ix, 265.
Perennial, densely tufted and usually aromatic grasses. Leaves often very
coarse. Panicles frequently much compound and contracted, spatheate.
Spikelets 2-nate, those of each pair differing in sex and more or less in shape—
except those of lowest pair of the lower or of both racemes which are homoga-
mous (male or neuter)—one sessile, the other pedicelled on the articulate tragile
tha:his of many-jointed paired racemes, terminating the culms and their
bianches ; raceme-pairs supported by a spatheole, collected into often decom-
pound or supra-decompound spatheate panicle ; the fertile spikelets falling with
the contiguous joint and the accompanying pedicel ; joints and pedicels filiio1m
5)
430 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
or linear with frequently more or less cupular or auricled tips, those of the
lowest pair (raceme-base) often conspicuously swollen, oblong or barrel-
shaped and hard. Sessile spikelets (above the lowest) feraale or hermaphro-
dite, dorsally, rarely, laterally, compressed, awned (normally) ; callus very
short, obtuse, shortly bearded. Involucral glumes equal or subequal, more or
less chartaceous, lower almost flat or slightly depressed or narrowly grooved on
the back, with at least from the middle upwards sharply inflexed margins, 2-
keeled, upper more or less boat-shaped, keeled upwards, usually 1-nerved.
Floral glumes ciliate or ciliolate (sometimes cbscurely), lower entire, hyaline,
2-nerved, upper 2-fid or 2-lobed, hyaline, rarely firmer and almost stipe-like
below the insertion of the awn; column of awn, if any, smooth. Pale 0.
Lodicules 2, minute, glabrous. Stamens3. Stigmas laterally exserted ; styles
terminal. Grain oblong in outline, subterete to plano-convex in cross-section ;
embryo about half the length of the grain. Pedicelled spikelets usually slightly
different in shape and size from the sessile, but never depressed or grooved on
the back. Involucral glumes muticous, lower chartaceous to sub-chartaceous
upper thinner. Lower floral glume hyaline, 2-nerved, upper 0, but usually a
male flower present.
Species about 36, in the tropical, more rarely in the subtropical regions of the
Old World.
Cooke mentions one indigenous (Andropogon Jwarancusa, Jones) and one
cultivated species (Andropogon Schenanthus, Linn.) belonging to this genus.
We add two indigenous species and a cultivated one:
Key, mainly after Stapf -
A. Blades long, hard, rough-edged throughout, filiform to linear ;
lower involucral glume of sessile spikelet flat or concave between the keels
I. Raceme-joints villous all over, hairs long,
more or less concealing the sessile
spikelets ; awn usually a straight, very
short bristle
1. Basal leaf-sheaths in dense tufts, tightly
clasping, thickened below ; blades
more or less filiform and flexuous,
except when very short; raceme-
fascicles more or less simple we Ll. C. Schoenanthus.
2. Basal leaf-sheaths ultimately loosened
and curled; blades flat; raceme-
fascicles compound bs 2. 2. °C. Jwarancusa.
II. Raceme-joints bearded along the sides,
but hairs not concealing the sessile
spikelets; fertile spikelets awnless ... 3. C. cztratus.
B. Blades flat, 5-30 mm. wide, rounded to
subcordate and stem-clasping at the base,
of a soft texture, with smooth edges, at
least in the lower part ; lower involucral
glume of sessile spikelet with a narrow
groove from the middle downwards
corresponding to a keel inside
I. Blades 10-30 mm. wide (rarely under 10
mm.), somewhat fat, rich green, at
least above; panicles 10-30 cm. long,
turning reddish when mature oa ACM Martine.
II. Blades 2-6 mm. wide, thin, glaucous ;
panicles 10-20 cm. long, glaucous or
straw-colour when mature ... ptt se Oma StULS.
1. Cymbopogon Schenanthus, Spreng. Pug. ii (1815), 15, wow Schult.; Stapf
in Kew Bull. (1906), 303-313, 352-353, in Prain Fl. Trop. Afr. ix, 268.—C.
arabicus, Nees ex Steud. Syn. Pl. Glum. i (1855), 387.—C. Arriani, Aitch.
Cat. Punj. Pl. (1869), 174.—C. Curcinnatus, Hochst. ex Hack. in Monogr.
Androp. 599.—Andropogon Schenanthus, Linn. Sp. Pl. ed 1 (1753), 1046
(non Hackel et plurimorum auct.).—A. bicornis, Forsk. Fl. Aegypt.—Arab.
(1763), 173, non Linn.—A. laniger, Desf. Fl. Atl. ii (1800), 379; Boiss. FI.
Or. v, 465; Balfour Bot. Socotra, 316; Hack. in Monogr. Androp. 598;
Benth. in Hook. Ic, Pl. t. 1871.—A. Olévieri, Boiss. Diagn. Pl. Or. ser. +1,
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 431
fasc. v (1844), 76.—A. circinnatus. Hochst. e¢ Steud. Syn. Pl. Glum. i (1855),
387.—A. Arriani, Edgew. in Journ. Linn. Soc. vi (1862), 208.—A. Jwaran-
cusa, subsp. daniger, Hook.f. F.B.I. vii. 203; Cke. ii, 976 (var. tantum).—
Gymnanthelia lanigeru, Anderss. in Schweinf. Beitr. Fl. Aethiop. 306 (nomen
tantum).
For Mee a of the foundation of this species see Stapf in Kew Bull.
(1906), 303-305.
Description: Perennial, compactly cespitose, with numerous intravaginal
innovations, 15-45 cm. high. Culms erect, slender, few— to 4-noded and simple
below the inflorescence, terete, glabrous, very rarely with a few small hairs at
the nodes. Leaf-blades semiterete, filiform, wiry, flexuous, very firm and
often circinate upwards, rounded on the back, channelled on the face, or those
of the culms somewhat flatter and shorter, up to more than 23 cm. long, 1 mm.
in diameter, glabrous, finely scaberulous on the nerves below, though often
smooth to the touch, pale, glaucous, evenly 7-9-nerved, the midrib showing
only above as a broad, white band. Ligules membranous to scarious, oblong,
truncate, ciliate, up to 3°3 mm. long. Sheaths very firm, smooth, glabrous,
tight, those of the innovations and base of the culms widened at the base, very
tough and long-persistent, straw-coloured, up to 13 cm. long. Spatheate
panicle narrow, 8-30 cm. long, few—to 7-noded lower internodes 5-7°5 cm.
long, upper rapidly decreasing in length, slender, glabrous ; lowest primary
branch rarely undivided at the base, 3-2-noded and up to ]5cm. long, usually
forming up to 4-rayed tiers ; lowest subtending sheaths with foliaceous blades;
rays finely filiform, 2°5-3°7 cm. long, rarely to over 5cm., glabrous; spathes
narrowly lanceolate, subherbaceous, often tinged with pale purple, witha
short blade or the upper bladeless and produced into a setaceous point, 3°7-4°3
cm. long, glabrous. Spatheoles very narrow, acute or with a setaceous point,
12-25 mm. long, pale or straw-coloured ; peduncles finely filiform, widened up-
wards 3°3-4'2 mim. long, tips truncate. Racemes 2-nate, more or less divari-
cate, at length epinastically deflexed, 1-2 mm. long, white-villous, pale or
tinged with purple, one subsessile, the other with a bare base, 1-2 mm. long,
bases puberulous to pubescent in the fork, ciliate-bearded upwards, with
minutely cupular and denticulate tips, that of the subsessile raceme as well as
the adjacent pedicel stout, elliptic to elliptic-oblong in outline and convex on
the back, ultimately more oz less glabrescent and glossy ; fertile joints filiform,
slightly widened towards the oblique subcupular auricled tips, 2°7-3'3 mm.
long, densely hairy to villous from the back and the angles ; adjacent pedicels
similar to the joints but more slender. Womogamous pair of spikelets one at
the base of the sessile or of both racemes ; the sessile spikelet of the lowest but
one of the sessile raceme intermediate and imperfectly awned. Fertile spike-
lets linear-lanceolate, more or less acuminate, acute, including the callus
5'3-6°3 or even 7°4 mm. long, glabrous, pale green below, reddish upwards ;
callus short obtuse, shortly bearded. Involucral glumes equal, chartaceous,
lower nerveless and shallowly concave between the acute scaberulous keels,
minutely 2-denticulate, upper lanceolate-oblong in profile, acute, slightly
curved on the back, l-nerved, margins broadly hyaline upwards, ciliate.
Lower floral glume linear-oblong, nerveless, hyaline, ciliolate, slightly shorter
than the involucral glumes, upper very narrow, shortly 2-fid, cuneate-linear
and chartaceous below the insertion of the awn, less than 3°3 mm. long,
lobes broadly lanceolate, ciliate, awn up to 1 cm. long’, very fine, niore or less
keeled at and slightly twisted below the middle; column smooth. Anthers
2mm. long. Pedicelled spikelets male, linear-oblong, 4:2--6°3 inm, long,
glabrous, more reddish than the sessile ; involucral glumes subchartaceous,
with 5-9 evenly distant intracarinal nerves, the upper thinner, 3-nerved ; lower
floral glume linear-oblong, sub-2-nerved, ciliolate, 42 mm. long; upper
floret reduced to a male flower, or its glume present as a microscopic scale.
Locality: Sind: (Stocks 816, Woodrow); Jemadar ka Landa _ near
Karachi (Stocks).
Distrbution : From Morocco through N. Africa, Arabia, Persia, Afghani-
stan, Baluchistan, Punjab, Sind.
flistory and Uses: See Stapf in Kew Bull. (1906), 305-313, 353.
2. Cymbopogon Jwaranctisa, Schult. Mant. ii (1824), 458 ; Stapf in Kew Bull.
(1906), 354: Haines Bot. Bihar and Orissa 1045.— Andropogon Jwarancusa
Jones in Asiat. Research. iv (1795), 109; Cke. ii, 976.—A. Jwarancusa var.
432 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
genuinus, Hack. Monogr. Androp. 5)9.—A J. subsp. Jwarvancusa proper,
Hook. f. F. B. I. vii, 203.—A Jlaniger, Duthie Fodd. Grass. N. Ind. t. 23.
Vern. Names : Jwarankusa (i.e. fever-restrainer) (Sanskr.), khavi (Hindust.)
Description: Usually a tall grass, up to 18m. high, with very aromatic
roots, densely tufted, the stems from clusters of firm, persistent, finaliy loose
and open and tortuous leaf-sheaths, more or less widened below. Leaves flat,
up to 60cm. long and 5 mm. broad, narrowly linear, filiform above and ending
in along capillard tip, liguleQSmm. long, membranous. Panicles long,
narrow interruptey, with very compressed, short, fascicled branches bearing
spathes about 5 cm. long and spatheoles 6-18 mm. long. Racemes 1 4-18 cm.
long, often 5 joined, joints half as longas the uppermost villi. Spikelets 3-4
pairs, green, half hidden by the 5 mm. long villi, on the joints and pedicels.
Sessile spikelets 5 mm. long; lower involucral glume flat or concave
between the keels, which are neither winged nor margined (omitting
of course, the ordinary inflexed margins of the glume common to the genus)
or sometimes narrowly margined, scabrid or ciliolate, nerves 2-4 or 0 between
the keels. Joints of rhachis and pedicels subclavate, with toothed tips.
Pedicelled spikelets equal or rather longer than the sessile, narrowly lanceolate,
purplish ; lower involucral glume 7-9 nerved.
Note.—This species is nearly related to C. Schenanthus and the two, as
pointed out by Hackel (1. c. p. 600), are not always distinguishable with cert-
ainty. ‘At high altitudes, ’ says Stapf (1. c. 314), ‘ asin Kumaon and Spiti, or
inthe dryer parts of the Punjab, it (C. Jwarancusa) becomes dwarfed
and narrow-leafed and forms a ‘‘ transition state’’ to C. Schenanthus. The
latter is a characteristic desert plant, able to exist witha minimum supply of
water. On the other hand, C. Jwarancusa is dependent on an, at least
temporarily, abundant supply of water, and prefers the neighbourhood of
rivers, or actually grows in the beds of torrents. Itis not impossible, that the
distinguishing characters of C Jwarancusa as compared with C. Sche-
nanthus, that is the robust state, the long, flat and relatively broad leaves, and
the more composite panicles, are mainly due to eolaphic conditions.’
Locality : Stnd: Karachi (Dalzell and Gibson); Bholari (Bhide !); Shikarpur
(Bhide !); Umerkot, sandy plains (Sabnis B1082!); Gharo (Blatter and McCann
D669! D670!) ; Gholamalla (Blatter and McCann D671!).—Gujarat - Ahmeda-
bad (Dalzell and Gibson).
Distribution : Outer hillzone of the United Provinces, Kumaon, Garhwal
(up to 3,000 m. or over) and westwards as faras Peshawar, Jodhpur and Jais-
almer States, Sind, Bihar.
Uses : Stapf is of opinion that this grass is very probably used along with
C. Schaenanthus. See also Stapf (1. c. 313-314).
*3. Cymbopogon citratus, Stapf in Kew Bull. (1906), 357 in Prain Fl. Trop. Afr.
ix, 282.— Andropogon Schenanthus, Linn. Syst, ed. x (1759), 1304, not of Sp.
Pl.; Roxb. Fl. Ind. ed. Carey and Wall. i (1820), 278.— A. citratus DC. Cat.
Hort. Monsp. (1813), 78 ; Nees in Allgem. Gartenz. iii (1835), 266.—A. citriodo-
rum (sic!) Desf. in Tabl. Ecole Bot. ii (1815), 15.—A. Roxburghit, Nees in
Wight Cat. (1833), no. 1699 (nomen tantum) ; Steud Syn. Pl. Gluin. i,
(1855), 395---A. ceriferus Hack. in Mart. Fl. Bras ii, pt. iii (1883), 281.—A.
nardus, var. ceriferus, Hack. Monogr. Androp. (1889), 605. Sehwnanthus
amboinicum, Rumph. Herb. Amboin. v, 181, t. 72.
For the taxonomic position of this species see Stapf in Kew Bull. (1906),
330-333.
Vern. Names: Oleu cha, hirvacha (Mar.), lilicha (Guj.), vasane-hullu,
kavance hullu, majjige hullu (Canarese).
Description « A tall perennial, throwing up dense fascicles of leaves from a
short, oblique annulate, sparingly branched rhizome, usually barren, but
occasionally giving rise to a stout erect culm up to over 18 m. high, 7-8-noded
and simple below the panicle. Leaf-blades linear, long-attenuated towards the
base and tapering upwards to a long setaceous point, up to over 90 cm. long by
16-18 mm. wide, very firm, glaucous-green, glabrous, sinooth or more or less
rough upwards and along the margins ; midrib somewhat stout below,
whitish on the upper side; primary lateral nerves 4-6 on each side, raised
particularly above with 2-4 secondary nerves between them. Ligules very
short, scarious, rounded or truncate. Sheaths terete, those of the barren
shoots much widened at the base, and tightly clasping each other, narrow and
separating upwards, with rounded shoulders at the mouth, 10-30 cm. long,
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 433
subcoriaceous, quite glabrous and smooth, more or less cinnamon-coloured or
russet on the inside ; sheath of the culms tight, shorter than the internodes,
finely pubescent or velvety at the nodes . Spatheate panicle decompound to
subdecompound, loose, 30 to over 60cm. long, nodding; internodes 4 to over
6, the longest up to 20 or 22 cm. long, rapidly, decreasing in length upwards;
lowest primary branches undivided at the base, upto over 45 cm. long, and
up to 5- or 6-noded, the following forming mixed tiers of very unequal variously
compound and sim ple rays, ultimate tiers upto 4-rayed ; rays filiform and
glabrous; spathes narrow-lanceolate, acute or acuminate, 2° 5-5 cm. long with
narrow membranous margins. Spatheoles very narrow, linear-lanceolate to
aimost subulate when inrolled, 14-18 mm. long, acute or finely acuminate,
reddish to rich russet. Peduncles 6-10 mm. long, glabrous. Racemes 2-nate,
finally spreading at right angles or epinastically deflexed, moderately dense,
14-25 mm. long, pale, variously tinged with duli purple, loosely villous, one
subsessile, the other with a slender filiform bare base, almost 2 mm. long and
hairy, the pedicel of the homogamous pairalso slender, though short; fertile
joints filiform, slender, 2-3 mm. long, ciliate on both sides, tips obliquely
auriculate and cupular, adjacent pedicels very similar. Homogamous pair of
spikelets 1 at the base of the sessile raceme, its sessile member usually slightly
differing in shape from the fertile spikelets. Fertile spikelets linear to linear-
lanceolate, acutely acuminate, 5-6 mm. long, reddish, glabrous; callus short,
obtuse, minutely bearded. Involucral glumes subequal, lower subchartaceous,
slightly depressed towards the base, otherwise flaton the back, keels acute,
scaberulous above, intracarinal nerves 0 orl, short or indistinct, upper boat-
shaped, slightly curved on the back, acute, keeled upwards. Lower floral
glume hyaline, linear-oblong or almost linear, sub-2-nerved, ciliolate above,
slightly shorter than the involucral glumes, upper narrowly linear, acute, about
4mm. long, usually entire and awnless, rarely more or less 2-fid with a small
bristle from thesinus. Anthers 2mm. long. Pedicelled spikelets male or
neuter, linear to subulate-lanceolate, as long as the sessile, reddish, glabrous;
lower involucral glume 5-9-nerved, upper 3-nerved; lower floral glume
shorter to much shorter than the involucral glumes, hyaline, ciliolate, upper
very narrowly linear, nerveless if present at all.
Locality: Gardens in Bombay.
Distribution: This grass is only known in the cultivated state. Probably of
Indian origin, and now widely distributed over the tropics of both hemispheres.
See Stapf in Kew Bull. (1906), 334.
History and uses of the Lemon-grass : Stapf 1.c. 322-330. 334, 358.
4. Cymbopogon Martini, Stapf in Kew Bull. (1906), 359 ; Haines Bot. Biharand
Orissa 1046.—C. Martinianus, Schult. Mant. ii (1824), 459.—Andropogon
Martini, Roxb. Fl. Ind. i (1820), 280.--A. pachnodes, Trin. in Mem. Acad.
Petersb. ser. 6, ii (1833), 284. and Spec. Gram. Ic. (1836), tab. 327.—A.. Calamus
avomaticus, Royle, Essay Antiq. Hind. Med. (1837), 33 (momen tantum), Ilustr.
Bot. Himal. (1840), tab. 280.—A. nardoides, a Nees, Fl. Afr. Austr. (1841), 116.
—A. Schenanthus, Fluck. and Hanb. Pharmacogr. (1874), 660 (zow Linn.)—
A. Schenanthus, var. genuinus, Hack. Monogr. Androp. (1889), 609 (artim).
—A. Schenanthus, var. Martini, Hook. f. in F.B. I. vii (1897), 204 (exclus.
synon. referentibus ad plantas Africanas).—For the foundation of this species
and its synonymy see Stapf in Kew Bull. (1906), 335-337.
Vern. Names: Geranium grass, Rusa grass; rohisha, rosem (Sanskr.) ;
rusa, gandh-bel, mirchia gandh, tikhari (Hindust.): rohish, roshegavat
(Mar.) ; rhonse, rauns (Guj.) ; eunthi hullu, khasi hullu (Kanar.).
Description - A tall, perennial. sweet-scented grass, 1°5-2°4m. high. Stems
glabrous, straw-coloured, leafy. Leaves flat, usually broad, rounded or sub-
cordate at the base, more or less glaucous beneath, those below the inflore-
scence rarely under 23 cm. long by 1] cm. wide at the base, but often 2'5 cm,
wide below, tapering from a little above the base or from the middle to a firm
tip, glabrous except for the scabrid margins, margin sometimes smooth near
the base. Spikes 2-nate, 12-18 mm. long, oblique or divaricate or less often
deflexed. Peduncle about half the length of the 18-25 mm. long spatheole,
several spatheoles and their peduncles from a spathe of lower order, these
arranged in long usually narrowly oblong panicles not more than 3'5-5 cm.
wide, but sometimes panicle with many branches and broader. Joints and
pedicels slenderly clavate (excluding the much thickened lowest), about half as
434 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXI1
long os the sessile spikelets, tips with a lanceolate tooth or 3-toothed, margins
long-villous, 3°5-5 mm. long. Lower involucral glume (above the lowest spike-
let) with lanceolate centre becoming oblanceolate or oblong from the keels
being membranously winged above the middle, back with a vertical median
depression below the middle corresponding to a ridge inside ; upper cymbiform
with the dorsal keel winged above, minutely ciliate below.
Locality : Gujarat: Champanir (Chibber!) ; Ahmedabad, dry hills (Sedg-
wick 310!) : Junagad, Kathiawar (Blatter 3783!) ; Bhuj-Rhodir-Maha, Cutch
(Blatter 3649!) ; Anjar, Cutch (Blatter 3741 !).—Ahandesh : Road to Chinchpada
(Chibber!) ; Toranmal (McCann A235 !).--Konkan : Wada Range (Ryan
488!) ; Gokhirva, Bassein (Ryan 41!); Keltan (Ryan 392!) ; St. Xavier’s
College compound (McCann 446! !).-—-Deccan : Ganeshkhind Botanic Gardens
(Garade 435 !) ; Purandhar (Bhide !) ; Pashan (Gammie !) ; Modasa (Sedgwick
and Saxton!) ; Khandala, very common (McCann !) ; Igatpuri (Blatter and
Hallberg 4432!) ; Purandhar (McCann 5010 !) ; Kasara, Igatpuri Ghat (McCann
4343A !) ; Panchgani (Blatter and Hallberg B1248!, B1282 !, B1297 !, B1324 !).
—S. M.Country : Haveri (Talbot 2180!) ; Dharwar (Talbot 2616!) ; Badami
(Talbot 2928 !). [According to Malcolmson, ‘ the Rusa grass in the Deccan
affects particularly the trap, more or less avoiding the granite, so much so that
he was able to trace the green-stone dykes across the granite by the luxuriance
of the grass ’ (ex Stapf) ].
Distribution : From the Afghan frontier to the Rajmahal Hills in Bengal and
from the subtropical zone of the Himalaya to about 12° N., excluding the desert
region of the Punjab and the greater part of the northern Carnatic.
Stapf excludes also the outer slopes of the Western Ghats, but the localities
given above show that tne grass is well represented in that region. For the
history and uses of the Rusa grass oil Olewm Palmarose seu Geraniti Indici
(Palmarosa oil) see Stapf in Kew Bull. (1906), 338-341, 360.
5. Cymbopogon caesius, Stapf in Kew Bull. (1906), 360. ia Prain FI. Trop.
Afr. ix, 287.—Andropogon caseius, Nees in Wight. Cat. (1833) nos. 1700b (omen
tantum) and in Hook. and Arn. Bot. Beech. Voy. 244 cum descriptione
(partim).—A. Schenanthus, var. crvestus, Hack. in Monogr. Androp. 610;
Schweinf. in Bull. Herb. Boiss. ii, App. ii, 14; Hook. f. in F.B.I. 205, exlus.
fere omnibus synon.
For foundation of this species see Stapf in Kew Bull. (1906), 344.
Description : A perennial, tufted grass, up tol m. high, with intra— and ex-
travaginal innovation-shoots from a short rhizome. Culms erect or geniculate-
ascending, slender, more or less wiry, frequently branclied below, the branches
often in fascicles from the knees, often many-noded, terete, glabrous, smooth.
Leaf-blades linear from a scarcely narrcwed rarely slightly rounded base,
tapering to a long setaceous point, those of the culms up to over 15 (sometimes
almost 3C) cm. long, 2-6 (sometimes 10) mm. broad, of the innovations usually
much shorter, flat, bluish-glaucous, glabrous, smooth, midrib slender, primary
lateral nerves very fine, 3-4 on each side. Ligules very short, rounded,
scarious. Sheaths rather firm, tight, the lowest mostly short, those placed at
branching nodes at length thrown aside, inrolling or deciduous, glabrous,
smooth, usually much shorter than the internodes. Spathaceous panicle
narrow, mostly 7-15 cm. long, rarely much longer, sometimes reduced and
small, dense or interrupted ; internodes usually 4-6, the lowest rarely exceeding
athird of the panicle, the following gradually decreasing ; lowest primary
branch shortly exserted from its sheath, undivided at the base, or like the
following forming mixed or (upwards) simple-rayed tiers; rays of ultimate
tiers 5-3, finely filiform. 7-10 cm. long, glabrous ; lowest subtending sheaths
with foliaceous blades: spathes lanceolate, acuminate—2°5—4 cm. long, sub-
herbaceous, glaucous, sometimes turning reddish. Spatheoles narrowly lanceo-
late, acuminate, 14-16 mm. long, subherbaceous to scarious, turning dirty
straw-colour or slightly reddish ; peduncles filiform 5-6 mm. long, glabrous.
Racemes 2nate, obliqualy erect. 12-14 mm. long, greenish, more or less
white-villous, one subsessile, the other with a bare base, over 2 mm. long, finely
pubescent on the inner side, clliate and thickened upwards, base of the sub-
sessile raceme swollen, hard, fused with the equally swollen and hard adjacent
pedicel ; fertile joints filiform, about 2 mm. long, glabrous on the back,densely
ciliate on the sides, cilia snow-white, tips often cupular with a crenulate margin
or auricle ; adjacent pedicels very similar. Homogamous pair 1 at the base of
REVISTON OF THE FLORA OF THE BOMBAY PRESIDENCY 435
the sessile raceme. Fertile spikelets oblong, slightly wider above the middle,
subobtuse, 4 mm. long, greenish, glabrous ; callus very small, obtuse, minutely
bearded. Involucral glumes equal, subchartaceous., lower minutely truncate,
flat on the back with a fine median grove in the lower half, keels narrowly
winged from the middle upwards, intracarinal nerves 1 on each side towards
the keels, very fine ; upper narrow in profile, very acute, very narrowly winged
above the middle with ]1 delicate nerve on each side near the margin. Lower
floral glume delicate, oblong, minutely truncate, ciliolate, nerveless, upper
substipitiform, almost 3 mm _ long, 2-fid to the middle, segments subulate,
ciliolate, awn very fine, 10-14 mm. long, bent at and twisted below the middle.
Anthers almost 2mm. long. Pedicelled spikelets male, linear to lanceolate-
oblong, subobtuse, 4 mm. long, green, glabrous ; lower involucral glume
slightly convex on the back, subherbaceous, about 10-nerved, the inner 6 nerves
prominent, upper subhyaline, 3-nerved ; floral glume oblong, truncate, sub-2-
nerved, almost 4 mm. long.
Locality : Gujarat: On sandy and gravelly hills and banks, Ahmedabad and
Pranti}.
Distribution : Throughout the Carnatic, Gujarat, Arabia, Somaliland.
For history aud uses see Stapf Kew Bull. (1906), 342-345, 361.
(To be continued)
THE HOME OF THE EASTERN GORILLA
BY
Marius MAXWELL,
Author of ‘ Big Game Hunting with a Camera in Equatorial Africa’
(With a map and ten plates)
INTRODUCTION
Intending to visit a portion of the Eastern Congo and in parti-
cularthe Birunga (Virunga) mountains. I left Nairobi with my
companion, Mr. James Hugh Barnes, on February 11, 1925,
and travelled by motor car across Uganda. We joined our porters
at Kabale, a few days’ march from the border of the Belgian Congo,
and proceeded to Behungi (altitude 8,214 feet). From this spot,
situated on the eastern wall of the great Central Depression, we
enjoyed a splendid view over an extensive stretch of craterland,
partly on the border-line of Western Uganda and partly in the ad-
joining territory of the Congo. We also gained our first view of the
magnificent chain of mountains known as the Birunga volcanoes.
These mountains have long been known to form the main habitat of
the Birunga Gorilla! (Gorilla beringerzi), named after Oscar von
Beringe, who discovered the species in 1903. This is a very
different animal from the lowland gorilla found in other parts of the
Congo and West Africa, a young specimen of which could at one
time be seen in the Zoological Gardens in Regent’s Park. The
Kivu, Birunga, or Eastern Gorilla is a much finer creature, the adult
male being particularly impressive in appearance. The West
African Goriila differs from the Birunga Gorilla in many respects,
the hair, for instance, on the head and body being considerably
shorter in the Western form.
Our object was not only to acquaint ourselves with the home of
this comparatively little known ape, but aJso to study its habits
more closely. Incidentally, I wished to take a few snapshots of the
animal in its native surroundings. In accordance with these
intentions, over a month was spent on several of these volcanoes,
with an aneroid barometer and a hand-camera as our chief
equipment.
GEOGRAPHY OF THE BIRUNGA MOUNTAINS
It may perhaps be as well to preface this account of my experi-
ence with a brief description of the geographical position and the
physical aspect of these mountains. This introduction, illus-
trated with a few photographs, may help to establish a better under-
standing of the habitat of the Birunga Gorilla. (See Map.)
1 Also known as the Eastern or Mountain Gorilla,
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No. 1. Mt. Mgahinga (11,400 ft.) with Mt. Muhavura (13,547 ft.)
in the background.
(See Hxplanation of Plates.)
No, 2. Mr. James Hugh Barnes, the writer’s companion, with the Porters.
The precipitous peaks of Sabinio in the background.
(See Explanation of Plates.)
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THE HOME OF THE EASTERN GORILLA 437
The Bitunga Mountains proper, as the map indicates, are situated
to the north-east of Lake Kivu, and present, with the eight main
volcanoes, an imposing chain. This chain runs in a somewhat
south-westerly direction, almost at right angles across the main
depression or Central Rift, and roughly speaking separates the
basin of the Congo River from that of the Nile. Attheir base, and
extending over a large tract of country to the north as well as to the
south of these volcanoes, can be distinguished a multitude of small,
extinct craters overgrown with grass. These forma veritable
‘craterland’, and appear, when viewed from a great elevation
dotted like hillocks about the floor of this great depression. Inci-
dentally it may be observed that so much progress has been made
of late years inthe development of the Congo that the locality is
now easy of access.
The eight main volcanoes of the Birunga range to which reference
has just been made, are taking them in order from east to west :—
1. Muhavura, with an almost perfect regularity in shape, and
a summit rising to a height ! of 13,547 feet above sea-level.
2. Mgahinga, a truncated cone with slightly concave sides and
a height of 11,400 feet.
3. Sabinio, with its array of jagged and precipitous peaks, and
deeply furrowed slopes, attains a height of 11,960 feet.
4. Visoke, a truncated cone with almost straight sides, has a
height of 12,568 feet.
5. Karisimbi, with its unpretentious flat cone, is the highest
mountain in the whole range and reaches an altitude of 14,633 feet.
6. Mikeno, a massive mound with a rocky summit, as may be
seen from the illustration, is perhaps the most favoured home of the
Kivu or Mountain Gorilla, which is known by the local name of
Negéghi (pronounced Angéghz). Mikeno has a height of 14,385 feet.
Then follow the most westerly craters, Niragongo (11,250 feet) and
Namlagira, the lowest mountain of the whole range, with a height
of 9,766 feet. Niragongo and Namlagira stand side by side
together, but detached from the other six. The last two volcanoes
are at present the only ones that are still active. Niragongo appears
to be dying out.
The first six volcanoes can be divided into two distinct groups
connected by a long and comparatively low saddle, roughly eight
miles long, densely wooded, mainly with bamboo and tropical
undergrowth.
The eastern group consists of Muhavura, Mgahinga anc Sabinio,
which stand closely side by side in a line running approximately
from east to west. They form the present boundary between a
corner of Western Uganda and the mandated territory known as
Ruanda (Belgian occupation). Sabinio stands on the boundary of
Uganda and the Belgian Congo.
The second group, consisting of Visoke, Karisimbi and Mikeno,
is situated in Congo Territory, and the three volcanoes are situated
-1 The heights of the various volcanoes are taken from the Boundary Commis-
; South A. 35
sion Maps, 1923. (Provisional Issue) Sheet =a (ee
6
¢
438 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
roughly at the corners of an isosceles triangle. The saddle, men-
tioned just now, connects Sabinio of the first group with Visoke in
the second group.
These splendid mountains rise from the level of the highland
plains which are already from 5,090 to 6,000 feet above sea-level.
On aclear day they forma commanding array visible for many
miles around.
It is locally known that the Birunga Gorilla, or as the natives call
it, Megdghi, occurs in’ both the Sabinio and Mikeno groups of
volcanoes. The latter, however, is the more favoured habitat.
There can be no doubt that an occasional migration may take place
from the one group of mountains to the other, since the gorillas can
wander across the densely forested saddle between Visoke and
Sabinio without having to pass through inhabited valleys or piains.
There can therefore be no question of local distinctions between the
gorillas living in the two groups of mountains. Although it is not
improbable that these apes occasionally frequent the two most
westerly volcanoes Nirangongo and Namlagira, their presence there
is apparently, not so common that these volcanoes can be regarded
as permanent habitats.
The shooting of these great apes is now prohibited by the Congo
Government under penalty. Although the Birunga volcanoes have
recently, in 1924, been declared a closed sanctuary for gorillas, the
provincial Commissioner of the Kivu District, M. Ernest Ledocte,
very kindly allowed Mr. J. H. Barnes and myself to enter the
reserve, on condition that we would avoid killing any of the gorillas,
a condition we willingly accepted.
The absence of successful photographs showing the Birunga
Gorilla inits native forests is due mainly to the following causes:
Their normal habitat on these mountains lies at an altitude of
between 7,000 and 12,000 feet and they live among the densely
carpeted slopes, or in the thick bamboo forests, and the wooded
ravines and kloofs, where the light conditions are mostly too
unfavourable to allow for an instantaneous exposure. These
equatorial mountain regions have in general a humid and mist-laden
atmosphere, especially in the higher zones, above, say, 9,000
feet altitude. In these increased altitudes, the slopes and crags
are more frequently than not enveloped in mist, with the peaks
and the summits shrouded in dense clouds, or they are exposed to
rains, varying from a cold continuous drizzle, with heavy showers at
intervals, to a pelting tropical tornado and occasional hailstorms.
Secondly, the gorilla is averse to showing itself to human
beings. Its tendency is to shun observation and, if its privacy is
invaded, it retires into the densest parts of the forest undergrowth,
offering little opportunity for what can only be a hurried snapshot,
even with reasonable light conditions.
The vegetation on these Birunga volcanoes, especially on Mikeno’
and Karisimbi, can be roughly divided into three distinct zones.
The lower belt, say from approximately 7,000 to 9,000 feet,
consists of bamboo forests and large trees with dense under-
growth. Tangles of lianas and a variety of creeping plants hang
in curtains or festoons from the tree stems or from the lofty
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Journ. Bompay Nat. Hist. Soc.
No. 5. A Typical Forest of Mikeno (alt. about 11,000 ft.), showing
Hagenia trees in tall undergrowth.
(See Explanation of Plates.)
No, 6. Another Forest Scene on Mt. Mikeno at 11,000 ft. alt., showing
a portion frequented by Gorillas.
Oe
THE HOME OF THE EASTERN GORILLA 439
branches. In short, the scenery of this zone presents in many
places the primeval nature of a typical Congo Forest as it is so often
depicted in illustration.
The intermediate zone, say from 9,000 to about 12,000 feet
altitude, as typified by the vegetation of Karisimbi and Mikeno,
consists in the main of a vast undulating carpet of a chervil-like
plant and stinging nettles, interspersed with giant lobelias, docks,
sorrels, hemlocks and similar plants. Stately, broad-crowned
Hagenia trees (Hagenia abyssinica) rise from among this dense
undergrowth at such distances that in many places their crowns
touch one another. They form, as it were, a roof that covers the
undergrowth aia height of twenty-five to thirty feet from the ground,
while transmitting a certain amount of light, so that the mountainous
landscape below the trees remainsin moderate gloom. These Hagenia
trees, as may be noticed in the photographs, have mostly short,
thick stems with here and there very widely diverging branches of
great size. Some of these, especially the broken and decaying
branches, are heavily overhung with a variety of vines and creepers
and carry massive pads of rank moss and lichens. The base of the
stems of old trees frequently reach a circumference of fifteen feet ;
their roots are exposed and are often hollowed at the base. Cur-
tained as they are with trellises of luxuriant creepers, they form
niches serving as snug resting-places and night-shelters for the
gorillas of these regions. ‘The cautious stalker, if he is fortunate,
may occasionally finda gorilla peering out at him from among such
thickets. Owing to the height of the surrounding vegetation, the
animals are able to enter or leave these shelters without being
observed. The crevices and rents of these trees constitute the
favourite home of the mountain Hyrax, a small beast like a marmot
in appearance, which utters a peculiar cry.
The highest zone, commencing at an altitude of approximately
12,000 feet, shows an equatorial subalpine and alpine flora beginning
with patches of Giant Heath (£77ca arborea), the stems being ,fre-
quently covered with a ragged beard moss, which hangs in places
in long grey-green shreds. The height which these tree heaths
can reach is shown in the accompanying photograph. The foliage
-varies in colour from heather and dark brown to a light brown or
deepred. The giant heaths appear to grow best on shallow surface
soil resting upon the rugged volcanic rocks. ‘The rock is here and
there covered with great golden patches of moss or exposed in
scored and precipitous patches with glistening streaks of moisture
trickling from the sodden slopes and mountain ledges above.
Where the layer of soil is too thin to furnish sufficient hold to the
roots of these giant heaths, the heavy trees topple over. These
fallen trees, enveloped as they are in thick beard moss and lichens
present a curiously dismal conglomeration of dying and decaying
alpine vegetation. Higher up, the lobelias with their tall poker-
like tops and the giant grotundsels (Seveczo johnstonz) are the most
dominant features of the flora. In the highest regions the ground
- is covered with Alchemillas and thick drab moss.
After these remarks upon the local conditions, we will proceed
to discuss the habits of the gorilla itself.
440 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
GENERAL HABITS OF THE BIRUNGA GORILLA
In disposition, these apes are normally neither ferocious, nor
aggressive, nor do they even appear to be quarrelsome among
themselves. They naturally resent being suddenly and unexpected-
ly interfered with while dozing, or feeding in their secluded
retreat. Except under such provocation the beast is decidedly of a
retiring nature, and takes care to avoid human observation. On
the other hand it may well be imagined that the gorilla offers in its
appropriate mountain wilds an aspect quite different from that pre-
sented by an occasional immature specimen of the West African
lowland gorilla in captivity. In the former case, it is generally met
with in circumstances and surroundings that are apt to excite the
imagination of the pursuer, especially after he has read the various
stories of their ferocity and other alleged undesirable traits. On
close acquaintance with this inoffensive inhabitant of these altitudes,
it will be found that the gorilla differs in mentality very little from
the other apes. A more profound study. of the animals does,
however, show that certain primitive human instincts exist in these
anthropoids. On the other hand, it should be emphasized that there
is no suggestion of any skilled movements ; the idea of these apes
employing, in their wild state, sticks or stones as means of defence
or offence, would strike an observer, even after a most preliminary
acquaintance, as far from correct. Nature has provided these large
apes, especially the old males, with ample means for safeguarding
themselves against the attacks of carnivora. An occasional young
one may fall a victim to the leopard, traces of which animal are
fairly common in these regions, occurring even up to an altitude of
13,000 feet. The lion is not included in the fauna associated with
the gorilla of the Birunga volcanoes.
The first glimpse ofa gorilla in the more open parts of the
Hagenia and ‘ wild-chervil’ forests is as a rule the appearance of
a large, jet-black and shaggy head seton a pair of disproportionately
broad and square shoulders. Although the normal gait of the
gorilla is ostensibly not that of a biped, but that of a quadruped,
it frequently and unconsciously retains the erect posture. It may
on occasion be seen standing, or wading through the green vege-
tation, moving at intervals for afew steps in an erect attitude, and
reaching out for some attractive stalk. In its erect gait the gorilla
stoops forward perceptibly and shows in profile the back ofits
shaggy head in a straight line withits somewhat inclined back.
Gorillas squat about a good deal, and show a variety of comical
poses, especially when picking at the soft bark of the chervil-like
plants. They consume the pith of these plants, which, in certain
regions appears to constitute their main diet.
Curiosity will, as arule, prompt them to stand erect on their
short, bandy legs for quite along time and peer intently at the
intruder; in such cases, however, they rarely show more than their
head and shoulders, above the vegetation. The countenance of
this ape, when its curiosity only is aroused, bears a most peaceful
expression. ‘The neck being very short and massive, the head is
in consequence held somewhat rigidly. When scared or pursued
JouRN. Bompay Nar. Hist. Soc. Prats IV.
No. 7. <A Forest of Hagenia abyssinica on Mt. Mikeno at an altitude of
over 9,000 ft.
(See Haplanation of Plates).
No. 8. Arboreal Heath (Hrica arborea) and Senecio, a Giant Groundsel
(Senecio johnstoni). Note their respective heights by comparison
with the figure below the trees.
:
|
'
Journ. Bompay Nat. Hist. Soc. Puate V.
No. 9. The summit of Mt. Mikeno (14,385 ft.), showing alpine vegetation
on the slopes of the peak. -
(See Explanation of Plates.)
No. 10. A startled Female Gorilla (marked with cross in foreground)
making off with her young on her back.
(See Explanation of Plates).
THE HOME OF THE EASTERN GORILLA 44]
the gorilla does not take to the trees, but prefers to move silently
off on all fours through the dense under-vegetation, shunning
observation as much as the surroundings wil] permit. It is quickly
out of sight, anda fleeting glimpse ofits head and sloping back is
all that is presented to the observer. The arms being very long
in proportion to the bowed legs, it goes without saying that the
body stoops less during locomotion than it does with other apes.
The forepart of the body is supported by the knuckles of the hands
_ placed onthe ground. The adjoining photograph shows a startled
mother-gorilla in the act of escaping with her progeny seated like a
jockey and clinging to her back. I took this snapshot at a moment
when the animal mounted an obstruction across her path. She was
thus for an instant exposed to the camera, before disappearing
in the thick, yielding undergrowth. The attitude here depicted
is evidently the way they carry their infants when they are
prompted to move hurriedly through the dense ground vegetation.
At other times the young one may be seen encircling with its
arms the neck of the seated parent, sometimes with its little feet
clinging to the shaggy hair of its mother.
On the right of this picture, somewhat indistinctly, about half-
way up the mountain-slope, is a sub-adult gorilla in an agitated
condition and standing up among the dense forest undergrowth and
exposed to its waist. The circumstances were briefly these: I
happened to come suddenly upon a party of the feeding creatures.
Startled by my appearance, they uttered their discordant shrill and
reiterated cry of excitement previous to a hurried departure. The
_younger members are, on such occasions, apt to show their resent-
ment, or terror, by antics as well as by shrill cries, thus occasionally
giving the stalker time to take a hurried snapshot of his subject.
It will be noticed that the mist-laden atmosphere has somewhat
interfered with the clarity of the photograph, but in the circumstan-
ces it was all that the lens could be expected to give.
DIFFERENCES OF SEX AND AGE
The sexual differences in these apes are considerable. Whereas
the female gorilla, which is in build and height much inferior to
that of the male, invariably makes off with the stoop in the case of
a disturbance, the old male ‘of the family will, at close range,
behave quite differently. The bull gorilla of the party, usually a
very fine and powerful creature is on such occasions apt to give a
display of his violent resentment at the sudden intrusion. The
resentment is exhibited, first by a quick approach on all fours, to
the accompaniment of a sharp, abrupt and reiterated sound, some-
thing between a grunt and a bark, followed by an impressive ad-
vance in an erect posture, accompanied by a quick succession of
discordant screams not unlike the noises of a pig as it finds its way
to the slaughter-house. The suddenness, and the fact that these
demonstrations are generally carried out at uncomfortably close
range, in unexpected and awkward thickets, leaves one baffled for
the moment. Inthe cases experienced we found that once it has
expended the effect of its bluff and realized that its menacing
442 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXI1
approaches are of no availin driving the stranger away, it turns
promptly and joins the departing family group. The grotesque
appearance, at close quarters, of the stocky, bowlegged creature,
advancing at atoddle in an erect attitude is impressive ; with its
scalp drawn down and thus bristling its hairy crest it isa sight
enough to scare any wild creature of the forest. With the mouth
open and the retracted lips exposing the canines at each expiration,
the irritated animal then undoubtedly gives a pronounced ap-
pearance of ferocity. Heapproaches the intruder rapidly in his
stooping attitude, on all fours, and only rises on his feet at a few yards
from his adversary. The environment is usually such that it .
increases the effect of the sudden appearance from the undergrowth
of the great shaggy head and broad shoulders. I have not seen
gorillas beat their chest in such circumstances ; and I am not aware
that they ever do so when in movement, as their arms then seem
instinctively inclined to take advantage of any support. The noise
produced by thumping on the body is occasionally heard coming
from adult members of a party, and is inthe absence of intruders,
presumably purely an expression of exuberant spirits.
At a distance the old male may beat his chest in mild defiance
of an intruder, but the performance is, I have noticed, usually
carried out from some covert or in thickets where the creature
imagines himself concealed from observation. The performance
becomes sometimes ludicrous.
Whereas the tautiy paunched belly in the male is covered
with hair (as the adjoining photograph of gorilla clearly
shows), the chest is bare and _ wrinkled, attaining with
age a marked flabbiness; the armpits are likewise bare.
The sound of the beating on this part of its anatomy is in
the case of an old male nothing more formidable than that
which a portly person can produce by the loud and repeated thump-
ing of his naked chest with either the palm of his hand or with the
closed fist; the Kivu Gorilla appears to produce both these sounds.
The impressive forest-environment and the silence which reigns
over these man-forsaken regions has no doubt much to do with the
elaborate descriptions of these quite natural sounds.
The females are insignificant in appearance as compared to the
adult male, and are known to be from a foot to about a foot anda
half shorter than the males ofequal age. The female gorilla looks
in its wild state more podgy, or squatter, than the male.
The photograph of the dead male gorilla, as well as those that
follow of the captured young ones, were very kindly given to me
by Pére G. Provoost of the White Friars Mission at Lulenga, in the
Kivu District. The official seated beside the animal’s body is the
late Commandant Elias Arrhenius, a well-known Swedish gentleman
who was in the service of the Belgian Congo Government. Arrhe-
nius, who it will be remembered, accompanied Prince William of
Sweden to these parts, was among the first to send specimens to
Europe. This particular one, together with the pair of female
gorillas also shot by Arrhenius, appears to have reached the Swe-
dish Museum of Natural Science in the year 1916, although they
were secured before that date.
Journ. Bompay Nat, Hist. Soc, Prater VI.
No. 11. An adult male Gorilla.
(See Hxplanation of Plates.)
No. 12 A pair of female Gorillas secured on Mt. Mikeno,
(See Hxplanation of Plates.)
JOURN. BompBay Nat. Hist. Soc. Prate VII.
No. 13. Four captured young Gorillas.
(See Hxplanation of. Plates.)
No. 14. One of the four young Gorillas
with its guardian.
THE HOME OF THE EASTERN GORILLA 443
It will be seen from the photographs of the young gorillas, cap-
tured in the same region (Mount Mikeno), that they differ materially
from the adults. The very young ones have their whole body
covered with the coarse hair except the face and parts of the hands
and feet. The glabrous skin of the face is black and shiny. The
nostrils are large and prominent, and the deeply set eyes almost
hidden by the overhanging brows. The hair is jet-black and coarse
and resembles that of the common Himalayan Black Bear. In fact,
the first glimpse of a young female gorilla moving on all fours
- through the undergrowth reminds one much of the black bear in
India. One of the four young gorillas, a female, figuring in the
adjoining illustration, reachedthe Antwerp Zoological Gardens but
died unfortunately some time towards the end of 1924. One of the
warders of this Zoo, in charge of the Monkey House, informed me
that it had succumbed to an abscess of some kind in the throat, after
an unsuccessful operation. Its food in captivity consisted of bana-
nas, apples and vegetables. Celery, I should imagine, would be
the nearest approach to the gorillas’ natural foodin its mountain
home, which as already stated, is a kind of wild chervil with long
blanched and succulent stalks.
Another of the four specimens, if I am rightly informed, died
during the sea voyage, and the remaining two perished locally in
the Congo.
The capture of such young gorillas can easily be effected, in suit-
able localities, by creating a sudden commotion among a troop by
concerted shouting and the firing ofa rifle. Thus embarrassed, in
the confusion the young ones are separated from the others and
give an opportunity to the pursuers to close in and throw a sack, or
blanket over the terror-stricken creatures. Unfortunately such a
procedure is apt to involve the killing of one or two of the adults
among the party.
With adult individuals, especially the males of more advanced
age, the chest is bare to a well defined line about the waist, with
the skin wrinkled on the breast, forming in the old males distinct
creases. The back of the adult male exhibits a patch of grizzled
hair, which develops with advancing age into a broad and conspi-
cuous silver-grey band across the loins. ‘Thecrest of hair growing
on a fleshy swelling along the line of the sagittal suture, is very
prominent in the male, much, more so than in the female. This
sagittal crest is only erected to its full extent when the beast has
been molested and is thoroughly roused, either enraged or terrified.
The same habit is, of course, frequently exhibited by many other
mammals.
It will, however, be noticed that the gorilla does not, as monkeys
usually do, display their endeavour to terrify their antagonist by
approaching in abrupt starts. The adult male gorilla carries his
bluff further by a continuous advance in the upright attitude to
within very close proximity of the intruder. By such means he is
confident and no doubt accustomed to succeed in scaring the
occasional Batwa hunters that venture to seek their honey and other
edible forest resources in these regions. Ata distance he will not
embark upon this advance, but will usually content himself with
444 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
breast-beating and other signs of defiance and displeasure. In their
normal mood these great apes look phlegmatic and good-natured.
There isa varied individuality in their appearance and features.
Chest measurements of dead specimens at sixty inches in fully-
grown males have been recorded, and even a couple of inches more.
These measureinents would naturally be proportionately more for
the expanded chest of the live animal. Although their maximum
height is known to be in the neighbourhood of only six feet from
the crown to the heel, the weight of an adult male exceeds that of
two men of average weight. Some old males have been secured in
these regions which are stated to have weighed between four and
five hundred pounds. The short, bowed legs of the gorilla, sup-
porting the heavy, pot-bellied body, the excessively broad and
square shoulders and the long and powerful arms, reaching to the
knees when the creature is erect, give the Kivu Gorillaits singularly
stocky and massive appearance.
ARBOREAL HABITS
Although on the whole the Kivu Gorilla is not arboreal in its
habits, as can be seen at once from an examination of its hands and
feet,1 it should not be presumed that it is always a clumsy climber.
In the lower forest regions of Mount Mikeno, at an elevation of
approximately 8,000 feet, it is not unusual to see a young or a sub-
adult gorilla climbing atree some forty to fifty feet in height. In
such a case, the favoured observer will be astonished to perceive
the deliberate and, as it struck me at the time, methodical manner
in which these apes are able to ascend the tall trees, in much the
same way as amanclimbing up a pole, fitted with cross-spokes,
i.e. in an upright posture and using the human overhand move-
ments. They can remain comfortably perched among the top
branches, every now and then phlegmatically reaching out their long
hairy arms and plucking at some favourite morsel in the way of a
fruit or some young shoots. It is surprising to see how well they
blend with the foliage and how little of their body is exposed to the
observer, giving no chance of a successful snapshot. In the cases
witnessed, I have invariably seen them descend by the stem by
which they ascended, and in the same way. They seem to be far
from at ease when they are surprised in atree. In such a case they
will, after a moment devoted to satisfying their curiosity climb
down promptly and move off through the undergrowth.
They may at times remain hidden among the thick vegetation at
the base of some tree and utter their spasmodic bark of defiance
with the object of deterring the intruder trom further interference.
On one occasion I spent the greater part of an afternoon in the
1 Itis known that the hand of the gorilla is relatively shorter and broader
than in any of the other apes, the Orang-utan, or the Chimpanzee. It is the
same also with regard to the feet, which are almost flat and not arched as in
man, with the big toe widely separated from the other digits as in the other
apes. The photograph of the dead male shows that the fingers are short,
tapered, and thick at the base. In the foot, as the tracks of the gorilla on soft
ground will show, the heel is distinctly in evidence. The digits of the animal’s
foot are somewhat webbed at the base, making the small toes appear short
and stumpy. .
Journ. Bombay Nat. Hist. Soc. Piuate VIII.
No. 15. A Gorilla Shelter in the hollow base of a large tree.
(See Haplanation of Plates.)
No. 16. A recently vacated Night Shelter of a family of Gorillas showing
the excrements of the Apes (x) in the right foreground,
(See Haplanation of Plates.)
‘33 OOO'TT 9UIOS Jo opNgiy]e Ue 4B IqUTISIAey “4IT
JO Sedo[s usojsomM 04} UO 901} poyBoINziq V JO YOJOIN GYY UI pouUTIOJ ,, SANT ,, JO « P°Ds, BINOD Y “LT ‘ON
“XI Givig ‘009 ‘LSI “LVN AVANOG ‘Nunor¢
THE HOME OF THE EASTERN GORILLA 445
close vicinity of a troop of feeding gorillas, in the hope of being
able to secure a snapshot. The troop, which may have consisted
of a couple of dozen or more members, were busy feeding within
the confines of a very dense bamboo clump. Beyond hearing the
loud snapping or the splitting of stems at intervals, occasional
grunts, or a bark, we saw nothing but a stray glimpse of a peeping
head or a passing body. The flock, which we had disturbed earlier
in the day, was well aware of our presence but appeared now to pay
not the slightest heed to us. They pretended to be unaware of any
- scrutiny and wenton with their feeding, freely and without a care.
It appears quite evident that they are incapable of swinging
themselves from tree to tree as do the Orang-utans, for instance.
This is not difficult to understand, if we consider their external
anatomy. It is worth noting, however, that the younger gorillas
are capable of moving over the tops of dense bamboo thickets,
causing comparatively little damage to the trees, considering their
weight and bulk. In such case, they can spread themselves out
Sometimes with their arms apart, and crawl, so to speak, over the
bending clusters of bamboo tops, snapping an occasional weak
stem by their weight. The heavier males content themselves with
seeking their food at the base of the trees, causing at times aregular
din by the intermittent snapping of bamboo stems. The noise
created in a bamboo patch by a troop of a couple of dozen feeding
gorillas is not unlike that caused by a few elephants feeding in
similar forests. On the other hand, among open forest in the
higher altitudes, I found the gorillas singularly quiet and almost
silent, unless they happened to be suddenty startled at close range.
They remain quite silent throughout the night. A bamboo patch
evacuated by a large flock of about a couple of dozen individuals
bears ample testimony to the strength of which these animals are
possessed, especially the old males, not only in the shape of broken
bamboos some two and a half to three inches thick, but also of split
bamboo stems deliberately twisted.
Among the forests of the higher regions, the gorilla may show a
certain slowness, amounting almost to clumsiness, when they
proceed on all fours along the overhanging or slanting branches of
the Hagenia trees. This is readily understood if we consider the
animal’s configuration, and bear in mind the fact that the abdomen,
especially in large individuals of advanced age, protrudes considera-
bly, and gives a markedly pot-bellied appearance. The great weight
of these heavy apes must also be taken into account.
SLEEPING-PLACES, FOOD AND ENEMIES
Sleeping-places are commonly met with, and gorilla tracks are
plentiful in certain localities. These apes move about a good deal
and are constantly changing their feeding grounds. ‘Their un-
sheltered resting-places are made of twigs and leaves heaped up a
few inches from the ground, and generally bear the deep impression
of their posteriors, as they squat on these ‘ground-nests’. The
young ones, as the traces show, huddle at night round their parents.
It is not infrequent to see these nests fouled, and their vacated
sleeping-places are often littered with dung. Their proper night-
7
446 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
quarters are mostly sheltered, and the following photographs will
give an excellent idea of these retreats. The shelters are also
sometimes occupied in the day time at intervals during heavy
thunderstorms. The sleeping-places are, as a rule, not used on two
nights in succession. The gorillas of these regions appear to be
continually on the move from day to day, and seem to favour certain
surroundings. ‘This is acurious fact as there cannot be any question
of scarcity, nor any marked difference in the food on the slopes, or
ridges, in whichever direction they happen to turn. The shelters
have often the appearance of a natural wigwam formed at the
bifurcated or hollow base of some old Hagenia tree, with low and
massive overhanging branch; they are curtained with a profusion
of luxuriant vines and climbers. One of the photographs shows the
nature of this creeper-vegetation round a newly-vacated gorilla
night-shelter. In one corner may be noticed the excrement (marked
with a cross) of the departed occupants, reminiscent of the dung of a
horse. This was a snug and spacious retreat, which could have
offered a comfortable rain-shelter for some half-a-dozen men. In
some cases, the space is just enough to accommodate one large
male: here the old man gorilla squats on his ‘throne’ apart from
his family, surrounded by a drapery of creepers, which offers
effective protection against wind andrain. ‘The old animal is, more-
over, enabled to-peer from his eminence over the immediately
surrounding undergrowth, as the shelters are often situated on the
ridges or slopes.
Whereas the adult male gorilla seems to prefer the ground seats,
it not infrequently happens that the younger members of a flock
occupy small platforms, from four to six feet above ground level:
such little platforms, which resemble nests after they have been
occupied, are usually roughly built in the crotch of some large
Hagenia tree. |
The food of these animals consists mainly of tender bamboo
shoots, sorrels, hemlocks, etc., and especially the pith of the parsley-
like:plants already referred to as the wild chervil. Berries are also
eaten, and one finds, in addition, the remains of stalks of the
taller plants from which the bark has been neatly peeled off and
chewed.
As regards enemies of the gorilla, it may safely be accepted that
no herbivorous forest animal would feel inclined deliberately to
disturb a troop of gorillas, or even to remain for any length of time
in their immediate vicinity. On the other hand, I have reason to
believe, from the presence of leopards prowling on the skirts of a
party of gorillas that, as has been already mentioned, these wily
animals, with the patience and cunning characteristic of such
prowlers, prey upon anoccasional young gorilla. The gorillas of
these forests cannot be regarded as particularly wary. The adult
male gorillas are doubtless free from any kind of persecution by
beasts of prey, and therefore more unwary. The young gorillas
appear ignorant of their enemies; nevertheless they do not stray
far from their parents.
Apart from the strenuous exercise with which the stalking of
these great apes is invariably associated ina mountainous country
JOURN. BomBay Nat. Hist. Soc. PLATE X,
No. 18. Dead Leopard on the branch of a large Hagenia tree, showing a vacated Gorilla Shelter below.
THE HOME OF THE EASTERN GORILLA 447
covered with luxuriant vegetation, where, moreover, the atmosphere
is mist-laden and rarified, the shooting of these animals can present
no difficulty to the determined hunter. Few Europeans, however,
would feel inclined to take the life from these normally harmless
creatures, unless they were wanted fora definite purpose. The
gorilla is,in these regions, by no means as rare ashe has been
supposed to be. The difficulty of obtaining photographs does not
lie so much in actually surprising the animal in his mountain home as
in finding circumstances which will allow a snapshot to be taken
with areasonable hope of success. ‘The securing of a really effec-
tive picture remains at all times largely a matter of luck, although,
of course, the greater the patience and deter mination exercised the
greater the chance of success.
Nove ON THE GAIT OF THE GORILLA
‘As to the gait of the gorilla it is cléar that in the wild State but
few opportunities are obtainable for observation, owing to. the
difficult nature of the forest undergrowth.
Although rarely, they may yet at times expose themselves ange
completely when they happen to move across the trampled vegeta-
tion of their feeding tracks. Much can be learned from the animals’
spoor on the damp and soft humus soil.
In captivity the animal shows quite clearly the nature ef its
gait, as may be seen by watching the young West African specimen
which wasfora time at Regent’s Park. It was found to exhibit
three distinct gaits, namely: The walk on all fours, the upright
walk, and the run.
In the first instance there is nothing unusual about the motions
of their limbs as they are identical with those of a baboon moving
on all fours. The application of the extremities of its limbs to the
ground is, however, entirely different, since the gorilla, like the
Orang-utan and Chimpanzee, does not place the palms, but the
knuckles of the flexed hand on the ground. In the case of the feet,
it walks on the outer margin of the sole and heel with the toes
somewhat bent downwards.
Their walkin the upright attitude, in contradistinction to that on
al] fours, is in appearance not unlike the toddling gait of a pronoun-
cedly bandy-legged child.
Their accelerated mode of progression, or run on all fours resolves
itself into the following impacts, namely: right knuckles—left
knuckles—margin of right foot pad—margin of left food pad. In
fact, the impacts themselves succeed one another in much the same
way as in the canter of a four-footed animal, i.e., a transverse
motion. The arms of the gorilla being very long in proportion to
its bowed legs, the body naturally slopes at an angle towards its
posteriors, and consequently causes the head to remain more eleva-
ted than inthe baboon, for instance. The forward progression of
the body, in the run, is accompanied by a slight side swing, an
almost imperceptible side twist, of the animal’s hind quarters.
Although the great toe of the gorilla is widely separated from
the smaller digits, and forms with the latter a prehensile foot, it will
be noticed in the young specimen at the Zoo that in its erect, as
448 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
well as in its stooping gait the great toe is usually held in close to
the other digits. The large toe is held somewhat flexed, with the
ball turned inwards and slightly downward. The creature walks
normally on the outside margin of its heel and foot pad and holds
the sole slightly at a slant. Its tracks onthe soft mud usually show
adeep impression of the outer edge of the foot and heel, anda
faint impression of the small toes.
FULLER EXPLANATION OF PLATES
No. ]
Mount Mgahinga (11,400 ft.) with Mount Muhavura (132,547 ft.) in background.
Taken in the evening from the slopes of Mount Sabinio.
No. 2
Mr. James Hugh Barnes, Mr. Maxwell’s companion, with the porters. The
outline of the preciptous peaks of Sabinio, seen inthe background, bears resem-
blance to thatof a molartooth. This photograph was taken from a point on the
saddle between Sabinio and Mount Mgahinga, at an altitude of 8,600 feet. It
is remarkable with what scanty clothing these natives can face the raw cold on
these heights. The steep slopes of Sabinio are covered with bamboo to a
height of nearly 9,000 feet from which altitude commences the subalpine vege-
tation, first the arboreal heaths and finally the alpine groundsels with lobelias
among the Alchemilla rmeadows.
No. 3
Panoramic view of Mount Mikeno (14,385 ft.) and Mount Karisimbi on the left
(14,633 ft.), with the summit of the latter hidden inthe clouds. This photograph
was taken from a spot, to the east of Mikeno, with an altitude of 8,400 feet.
On these two mountains the bamboo forests cease at a height of some 9,000 ft.
and the vegetation from this height to about 12,000 ft. consists mainly of
Hagenia trees with widely spreading branches, amidst a meadow undergrowth
of chervil-like plants, nettles and other harbaceous growths, interspersed with
tall giant lobelias. Above 12,000 ft., the arboreal heaths, giant groundsels and
lobelias, among other alpine plants, make their appearance more common,
and take the place of the Hagenias; a soft carpet of Alchemillas and different
kinds of drab moss constitutes the ground vegetation. The three zones of flora
can roughly be distinguished on the photograph. ‘Traces of gorillas are met
with from an altitude of 7,000 ft. and are fairly common in places up to 12,000
feet. They become scarce above this height.
No. 4
Panoramic view of Mount Sabinio, taken from an altitude of some 8,000 ft.
above sea level, looking up into the north eastern gorge. Apart from numerous
glens and corries, the slopes of Sabinio are intersected by no less than three
immense gorges, one on the north-east, another on the south-east and a third
on the north-western slope.
The white patch of smoke, on the right-hand side of the throat of the gorge
shown about half-way up the mountain, indicates Mr. Maxwell’s night camp,
at an altitude over 9,000 ft.
No.5
Hagenia trees among the carpet of a soft undergrowth consisting of nettles,
docks, sorrels, hemlocks and mainly of a plant resembling a wild chervil, which
is a favourite food of the gorilla. This scene represents a typical forest of
Mikeno at an altitude of 11,000 ft. and thereabouts. The height of the under-
growth is indicated by the native standing on the right-hand side of the fore-
ground.
No. 6
Another forest scene on the same altitude and showing a portion frequented
by gorillas.
THE HOME OF THE EASTERN GORILLA 449
No. 7
A forest of Hagenta abyssinica with a view onthe western slope of Mikeno,
looking up towards itssummit. This view was taken at an altitude of a little
over 9,000 feet. The poker-like tops of a few giant lobelias are visible in the
foreground.
No. 8
Arboreal Heath (4rica arborea) and Senecio, or giant groundsels (Seecio
johnstont). Note their respective heights by comparison with the figure below
the trees.
No. 9.
The summit of Mt. Mikeno, height 14,385 ft. Taken from a point over
12,000 ft. in altitude to the south-east of this peak. The peak is occasionally
covered with snow at thesummit and frequently enveloped in clouds. In the
foreground can be seen the crowns of the last Hageniatrees. Note the alpine
vegetation on the slopes of the peak.
No. 10.
A startled female gorilla making off with her young on her back. Indistinct-
ly, on the right, about halfway up the slope may be seen a sub-adult Gorilla
showing the upper part of its body above the undergrowth. This photograph
was taken on the Western slope of Mt. Karisimbi, at an altitude of some
11,000 ft.
No; it.
An adult male gorilla, shot by the late Commandant Elias Arrhenius for the
Swedish Museum of Natural Science.
Norm .2;
A pair ot female gorillas, secured by the late Commandant Arrhenius and
destined for the Swedish Museum of Natural Science. The specimens were
secured on Mount Mikeno.
No. 13.
Four captured young gorillas, of which two were destined for the Antwerp
Zoological Gardens but only one of the two reached Antwerp. The remaining
two appear, from accounts given to me, to have died locally.
No. 14.
One of the four young gorillas with its guardian.
No. 15.
A gorilla shelter in the hollow base, and among the bifurcated roots of a
large tree. It forms a snug and spacious shelter for several people. Note the
vegetation round the roots and the luxuriant growth of the mosses.
Now-16:
A recently vacated night shelter of a family of gorillas, at the base of an old
Hagenia tree. Note the nature of the vegetation at this altitude (approxi-
mately 11,000 ft.) and the excrements of these apes on the foreground, to the
right of the trellis of climbers in the centre.
Noe dy,
A gorilla ‘Bed’ or ‘Nest’ on the western slopes of Karisimbi, formed in
the crotch of a bifurcated tree. ‘he ‘ Nest’ is about four feet from the ground.
As their traces point out the adults prefer their resting places on the ground,
The young ones occasionally occupy such elevated little platforms, a few feet
from the ground. This photograph was taken at an altitude of some 11,000
feet.
No. 18.
Dead leopard on the branch of a large Hagenia tree. Below the branch is a
vacated gorilla shelter.
INDIAN DRAGONFLIES
BY
F.C. FRASER, Lt.-COL., I.M.S., F.E.S.
Part XXIX
(With three plates.)
(Continued from page 319 of this Volume)
Genus—RHINOCYPHA Ramb. (cont.)
Group—ferforata.
Rhinocypha whiteheadi, Kirby, Ann. Mag. Nat. Hist, (7), ae v, p. 536, pl. xii,
fig. 4 (1894) ; Martin, Mission Pavie, Nevrop. (sep.) ieee Laid. Rec.
Ind. Mus. “vol. xili, pp. 38 and 39, (A917) me MEG pate (Ent.
Ser.), vol. Vii. No. 7, pl. viii, ig. 4 (1922)
Male. Abdomen 17 mm. Hindwing 21 mm.
Head: labium and labrum and rest of head velvety black marked with blue
as follows :—a reniform or oval spot on the outer side of each hinder ocellus,
asmaller postocular spot behind it and a medial linear occipital spot behind
tnem.
Prothorax black marked on each side with a rounded pale blue spot, anda
large blue spot on the dorsum of posterior lobe and hinder portion of middle
lobe, shaped like an arrow-head.
Thorax velvety black with a dark violet lustre, marked with azure blue and
chorome-yellow as follows :—the mesothoracic triangle pale blue, extending con-
siderably less than half-way up the dorsum of thorax, and very broad at base,
far more so than in others of the genus; an antehumeral stripe consisting of a
small isolated spot above and anarrow curved stripe on the lower half of
dorsum ; a fine posthumeral stripe closely apposed to the humeral suture, broken
above to leave a small isolated spot, behind which is another elongate larger
spot, all blue A broad interrupted stripe on lower part of sides, yellow, but
the portion on the metepimeron, blue anteriorly. Pruinosed white beneath.
Legs black, the two hinder pairs of tibia, and the femora pruinosed white.
Wings hyaline palely tinted with yellow at the bases, opaque blackish brown
apically, the hindwing marked on this part with three rows of vitreous spots.
Forewings with the apical portion opaque blackish brown as far inwards as,
rather more than half-way from apex to node, extending nearer the node along
costal border of wing, the inner border of the area serrate and running oblique-
ly backward and outwards; pterostigma black, the cells immediately beneath
it hyaline. Hindwing with opaque area darker and extending inwards for
about three-fourths the distance between apex and node, its inner border very.
irregular, indented and serrate. The vitreous spots a beautiful violet from what-
ever angle viewed. The apical spot single, 4 cell-rows deep by 6 cells wide,
irregularly shaped, situated 3 cell-rows behind pterostigma, which is black;
the middle row of spots 3 in number, all in alignment and of about equal
length, about 10 cells wide, the costal spot lying between /Azz and A7zzz, the
mid spot between //Azzz and Riv -+v, the hinder spot between Cuzz and JA ;
the proximal spot single, short, only 5 cells long, lying between /A7zz and
Riv + v, projecting slightly into the opaque area. All wings with 14 to
15 antenodal nervures; discoidal cell traversed 3 times in the forewing,
4 times in the hind ; wings petiolated to level of 2nd az; 21 to 26 postnodals.
Wings of approximately the sameshape and length, long and narrow, the hind
slightly broader than the fore.
Abdomen black marked with azure blue as follows :—segiments 1 to 4 with
lateral rounded spot at apical end of segments, growing successively smaller
Journ., Bombay Nat. Hist. Soc. PLATE |
ae
EU
as
mm 2
aS A TE Ee,
: ci
ae a
Ate
cid
Pr D
e
shu
ne
‘a8
1. Wincs or Lhinocypha bisignata, Selys, male.
2. Wrixcs or Rhinocypha biforata delimbata, Selys, male.
3. Wines oF Ahinocypha perforata perforata, Perch, male.
PLATE II.
Journ., Bombay Nat. Hist. Soc.
HH
3
ra
@
5H i LB
~ i fo
:
:
inocypha whiteheadi, Kirby, male.
WINGS OF &h
male.
male,
b
Wincs oF Micromerus lineatus, Burn.
Wines oF Calocypha laidlawt (Fras),
INDIAN DRAGONFLIES 451
from 1 to 4, segment 1 has also the apical border narrowly lined with blue, and
there is ashort ventro-lateral stripe on segments 2 and 3. Anal appendages
shaped as for genus; inferior appendages with a series of small robust spines
on upper surface.
Female. Unknown.
Distribution. Cachar, Assam. Very local but apparently not uncommon
where found, to judge by the numbers taken by Mr. Antrim. Martin reports it
from Tonkin, but Iwas unable to find any specimens in his collection in the
Paris Museum, and infer that he has mistaken arace of perfurata for this
insect. There is a single male from Sibsagar, Assam, in the Indian Museum
collection. Although belonging to group perforata, it is not very closely re-
lated to perforata itself, and cannot even be considered as a subspecies of that
insect as has been suggested. Its wings are not only broader than perforata
but are distinctly rounded at the apices. It ought not to be difficult to discover
the female of this beautiful insect.
Rhinocypha perforata perforata Percheron. (Agrion perforatus), Gen. Ins.
Neur. t, 2 (1835) ; Ramb. Ins. Nevrop., p. 235 (1842) ; Selys, Syn.
Cal., p. 63 (1853) ; Id. Mon. Cal., p. 219 (1854) ; Id. Bull. Acad. Belg.
(2), vol. xxxv, p. 487 (1873) ; Walk. (Lzbellago perforata), List. Neur.
Ins. B.M., vol. iv, p. 647, No. 7 (1853) ; Kirby, Cat. Odon., p. 114
(1890) ; Mart. Mission Pavie, Nevrop. (sep.) p. 17 (1912) ; Wiil. Proc.
U.S. Nat. Mus., vol. xxvii, p. 174 (1904) ; Krug. (A. apicalis) Stett.
Ent: Zeit. p. 79 (1898); Laid. (#. zzas) Proc. Zool, Soc., Lond,
pp. 88-90, pl. vi, fig. 6 (1902) ; Id. (#. apicalis) Fascic. Malayen-
sis (Zool), part 1. p.. 196; Id. Rec: Ind, Mus., vol. xii; pp. 38, 39,
(1917) ; Fras. Mem. Pusa (Ent. series), vol. vii, pl. viii, fig. 3 (1922).
Rhinocypha perforata perforata is not found within Indian limits, but is
represented by its subspecies or races /imbata and beatifica described below.
The type comes from Cochin China, and cotypes exist in the Maclachlan
collection from the Isle of Hainan, China, collected by Swinhce. It differs
from limbata by not having the border of the apex of hindwing vitrefied
hyaline and by the reduced opaque area in both fore and hindwings. In the
forewing this area occupies a little more than the apical fourth, but in the
hindwing extends to within four cells of the ncde. In /zmbata this area
extends nearly up to the node in the forewings, but is variable in the hind,
usually to about the same distance from the node, whereas in deafzifica it
extends right up to the node. Typein Paris Museum.
(In referring to my paperin the Memoirs of Pusa cited above, 1 note that
the figures have been wrongly numbered in the plate. Figure 4 is really
k. whiteheadi, and figure 3 should be F&. perforata limbata. ;
Rhinocypha perforata limbata, Selys, Bull. Acad. Belg. (2), vol. xlvii. p. 392,
(1879) ; Kirby, Cat. Odon. p. 114 (1890); Laid. Rec. Ind. Mus.,
VOL. Xi, pe oo (1917 ).
Male. Abdomen 16-18 mm. Hindwing 25 mm.
Head : labiuin dirty white, the apices of lobes black; labrum and rest of
head velvety black with a small rounded spot on the outer side of each
posterior ocellus, a larger rounded postocular spot and a linear medial spot on
the occiput, all bluish.
Prothorax black with the greater part of the posterior lobe rose pink or
lilaceous, and a large blue spot on either side of the middle lobe.
‘Thorax velvety black, the mesothoracic triangle extending rather more than
one-third up the dorsum, lilaceous in colour. On the outer side of this triangle,
and slightly longer than it, a large subtriangular azure blue spot with its
hinder border meeting the anterior part of the humeral suture. Above this
spot a small triangular upper humeral spot also blue, whilst laterally the greater
part of the sides are of the same colour, viz., nearly the whole of the mesopi-
meron, the upper two-thirds of the metepimeron and a narrow prolongation
along the hinder border of the humeral suture going as faras the root of
forewings. Beneath thorax 2-4 yellow spots which may be somewhat
obscured by pruinescence. In the Siamese and Annamese forms, there are
only two small rounded spots at the hinder part of thorax, whereas in speci-
mens from Assam, there are two additional larger oval spots in line with and
in front of these, .
452 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
Legs black, the flexor surfaces of the two hind pair of femora and tibiee
pruinosed white or creamy white.
Wings very long and very narrow, of equal breadth, hyaline in the basal
portions where they are palely tinted with yellow; opaque blackish brown in
the apical portions, marked with two series of vitreous spots in the hindwings.
Forewings with the outer third oniy opaque, this area stopping short of the
hinder border of wing, with a serrate border proximad and prolonged inwards
along the costal border between the costa and Az, to within 2-4 cells of the
node ; the hyaline border of wing posterior to the opaque area vitreous with
bluish, violaceous or green reflex which may extend as far as the extreme apex
of wing. Pterostigma black ; 15 antenodal nervures, 24 postnodals; discoidal
cell traversed twice only; petiolation ends at level of 2nd antenodal nervure.
Hindwing with opaque area extending inwards to within 1 to 4 cells of node,
the proximal border of this area ragged and indented deeply by a vitreous spot
8or9 cells long, between JAzzz and Riv-+v. The hinder border of wing
narrowly hyaline and vitreous, finely reticulated and extending round apices of
wing as far as pterostigma. ‘The apical series of vitreous spots very variable,
‘usually composed of 3 linear spots which decrease in length from costa to
hinder border of wing and which slightly overlap the level of inner end of
pterostigma. The costa] spot 11 to 16 cells long, lying, between /A7z and F222 ;
the hinder spot 3 cell-rows deep by 4 to 8 cells long, lying between //A7zzz and
Riv 4- v, whilst the middle spot, which is 2 to 3 cell-rows deep by 9to 10 cells
long, lies midway between the two others banded by short intercalated sectors ;
the inner series lying rather nearer the node than pterostigma, composed of 3
spots, a costal spot l cell-row wide by 10 cells long, lying between /Aiz and
Riiz, a medial spot 2 cell-rows wide by 10 to 15 cells long between /Azzz and
Riv +- v, and a hinder spot 1 cell row wide by 9 to 12 cells long between WA
and Cwiz. All these vitreous spots and areas glowing emerald green or
opalescent according to the angle from which viewed ; 2@ postnodal nervures,
14 to 15 antenodals ; discoidal cell traversed 3 to 4times ; petiolation the same
as for forewing ; pterostigma black.
Abdomen black marked laterally with azure blue as follows :—segment 1
with a large cuneiform spot on each side, 2 with a bilobed, longitudinal spot
extending from near the base to apical border, 3 to 4 with long wedge-shaped
spots with base of wedge at apical border of segments, segments 5 to9
with short triangular apical spots, that on 9 being very reduced.
Anal appendages black, superiors slender, cylindrical, semi-circular, a little
broadened at apex which is curved strongly in. Inferiors half the length of
superiors, moderately separated, bluntly pointed, a little denticulated at apex.
The superiors minutely spined on the outer sides near apex.
Female. Abdomen 18mm. Hindwing 26 mm.
Differs from male by having the wings entirely hyaline, tinted palely with
greenish yellow; pterostigma blackish brown, its outer half yellow at the
centre, especially in the hindwings ; 13 to 15 antenodal nervures in forewings,
26 to 30 postnodals.
Head similar to that of male with the addition of a large spot of yellow on
upper surface of epistome and two smaller ones behind it. The spots on vertex
and occiput also yellow, body-markings same as for male but yellow instead of
blue, the mesothoracic triangle finely outlined in yellow, its centre black.
Legs black, tibia and femora not pruinosed. Abdomen black with yellow
markings as for male, but with additional ventro-lateral stripes on segments
2to5. Vulvar scale as for genus.
Distribution. Limbata extends from Annam, through Siam to Burma and
Assam. The type in the Selysian collection, is from East Burma. The type
of perforata perforata is from Cochin China and is distinguishedfrom Jimbata
by the total absence of the hyaline and vitreous border of the hindwing, the
opaque area here coming right down to the border of wing, at least along the
extreme apex. The abdominal markings, said by Selys to be more restricted in
limbata, are not found to be so when aseries of specimens are examined. Inthe
forewing at least, the opaque areais greatly reduced as compared to /imbata or
beatifica.
Rhinocypha perforata beatifica, Fras. Rec. Ind. Mus., vol. xxix, pp. 86-87 (1927).
Similar in size and markings to /7mbatc, from which it differs by the greater
extent of the opaque area in all wings, extending in the fore and hind right up
INDIAN DRAGONFLIES 453
to the level of node. This is especially noticed in the forewings, where not
only does the costal streak extend to node, but also the body of the area, the
serrated border running from the node and extending as far asthe apex of
wing, leaving a rather broader area of hyaline border than is found in dimbata.
In the hindwing, the hyaline border broadens out at apex of wing, extending
right up to, or even overlapping, the outer end of pterostigma. This latter
has a narrow bluish centre in the hindwing along its outer half, and that of
forewing is pale for the same extent. In the outer series of vitreous spots, the
middle spot is 2 cell-rows deep and is fused completely with the costal spot, so
that in reality there are only two spots in this series.
Distribution. Naga Hills, Assam, from Aprilto June. ‘Type in the Fraser
collection. Seatifica differs from /zmbata to about the same extent as does the
latter from true perforata, so that it is quite possible, that given sufficient
material, it might be possible to construct a series showing agradual merging
of one into the others. For the present I consider it better to consider the
perforata series as one species with two subspecies.
Rhinocypha biforata biforata, Selys, Bull. Acad. Belg. (2) vol. vii, p. 446 (1859) ;
Id. ibid. (2), xlvii, p.392 (1879) (A. dzforata, race delimbata) ; Kirby,
Gat, Odon. “7p. li3;(1690); Waid. roc. Zool. Soc. Wound. (1), p. s&8.
(1902) ; Will. Proc. U. S. Nat. Mus., vol. xxviii, pp. 173 and 179, fig.
12, (1904) ; Laid. Rec. Ind. Mus., vol. xiii, pp. 37 and 38, (1917) ;
Fras. (2. deesoni) Mem. of Pusa. (Ent. Ser.), vol. vii, No. 7, pp. 61 to
63, (1922).
Male. Abdomen26mm. Hindwing 23 mm.
The species is not found within Indian limits but is represented by its sub-
species—ézforata delimbata Selys, biforata beesoni, Fras, and biforata abbreviata
subsp. nov. From all these 6zforata is easily distinguished by the greater extent
of the opaque area in the hindwings, where it extends right up tothe nodeora
little beyond it. The vitreous spots tend to be longer in ézforata and the reti-
culation somewhat closer. /&. deesoni falls closer to the parent species but
curiously enough, is separated from it geographically by the intervention of the
other two subspecies.
Rhinocypha biforata delimbata, Selys, Bull. Acad Belg. (2), vol. xlvii, p. 392
(1879) ; Kirby, Cat. Odon. p. 113 (1890) ; Will. Proc. U.S. Nat. Hist.
Mus., vol. xxviii, pp. 173 and 179, fig. 12 (1904) ; Laid. Rec. Ind. Mus.,
vol. xiii, pp. 37 and 38 (1917).
Male. Abdomen 19-20mm. Hindwing 23-26 mm.
Head velvety black marked with five bright ochreous spots, one-rounded or
oval on the outer sideof each posterior ocellus, around postocular spot on
each side anda medial oval occipital spot. Labium white, the lobes tipped
with black.
Prothorax black, the posterior lobe finely bordered with yellow and witha
large rosepink spot in its middle. A large bluish spot on each side near the
trochanters and a similar subdorsal spot on each side of middle lobe.
Thorax velvety black marked as follows :—the mesothoracic triangle rose-
pink or lilaceous, extending less than half-way up the dorsum of thorax but
continued as a fine line of the same colour along the middorsal carina as far as
the antealar sinus. On each side of the mesothoracic triangle, a large irregu-
larly ova! spot of the same colour, much larger than the triangle itself, a small
upper antehumeral spot, a broadish stripe behind the upper two-thirds of the
humeral suture, the greater part of the mesopimeron, and metepimeron, azure
blue. The hinder lateral suture, the lower part of the metepimeron, anda
quadrate area on the upper anterior part of mesopimeron all black, the latter
area bearing a small isolated blue spot. Beneath 6 large yellow spots separated
by black sutures and framed narrowly in black.
Legs black, the two hinder pairs of tibiz pulverulent white, the same two
pairs of femora pulverulent yellow on the flexor surfaces.
Wings hyaline, the greater part enfumed and tinted with yellow, which is
most intense near the bases. Forewings with apices tipped with blackish brown,
this marking gradually shading off into the hyaline area at outer end of ptero-
stigma (in ézforata biforata this area begins well proximal of the pterostigma
and covers about the outer fifth of the wing) ; discoidal cell traversed 3 times ;
14-16 antenodal nervures, 27-32 postnodals ; petiolation ends at the level of the
8
454 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
2nd antenodal nervure. Hindwings with about the apical fourth opaque black-
ish brown (the apical third or more in 6zforata biforata), this area marked with
a series of four linear vitreous spots, very irregular, each varying in length,
and the costal three often linked up by one or more connecting cells, the inner
level of the spots forming a concavity towards apex of wing, as dues also the
inner border of the opaque area, which extends slightly more basad along
hinder border of wing. Just proximal of the opaque area (projecting into it in
biforata biforata) a second row of vitreous spots.
The costal one of these spots lying slightly nearer the node than pterostigma,
composed of one row of cells, about 12 in number, lying between /Azz and R2iz,
the second spot shorter, lying more distad, 2 cell-rows deep by 8 or9 cells long.
lying between Aiv-+v and A722, the hinder spot at the same level as the costal,
9 cells long by 1 row wide, lying between WA and Cuzz. In the same space as
the medial spot of this series, but lying behind level of node, another vitreous
spot 1 row wide by 11-18 cells long. Pterostigma black; 15-16 antenodal
nervures, 26 postnodals ; discoidal cell traversed 3 times.
Abdomen black marked with azure blue as follows :—a large triangular spot
on each side of segment ], a midsubdorsal and an apicolateral spot on each
side of segment 2, as well as a ventro-lateral stripe below them ; similar spots
and stripes, but longer, on segments 3 to 5; segments 6 and 7 with the ventro-
lateral stripe only.
Anal appendages black, shaped as for genus.
Female. Abdomen 15-16 mm. Hindwing 23 mm.
Head black with the same markings as for male, but in addition, the bases
of mandibles, the basal segments of antennz, two large spots on frons and two
minute linear spots in front and to the outer side of the anterior ocellus all
ellow.
" Prothorax black with the following yellow markings :—the hinder border of
posterior lobe outwardly finely yellow, a narrow longitudinal medial line, often
broken into minute points on the dorsum of same lobe, a geminate point at the
anterior end of this on the medial lobe, a largish triangular spot on each side
of midlobe and a rounded spot on each side of the anterior end.
Thorax black, marked with blue and yellow as follows :—the mesothoracic
triangle finely mapped out in yellow, its centre black, a fine antehumeral line
incomplete above and strongly hooked outwards and backward in front, a fine
incomplete humeral line, all yellow. The sides similar to the male, blue.
Legs black, not pruinosed, the flexor surfaces of the hinder pairs of femora
yellow.
Abdomen black marked exactly as for male. Vulvar scale and anal ap-
pendages as for genus.
Distribution. Burmah and Assam, common in parts of the former but
apparently rare or local in the latter. I have specimens from Mergui, Lower
Burma collected in November, and from Maymyo, Upper Burma, collected in
July, the latter specimens by Col. F. Wall. It appears to be the commonest
Rhinocypa in the Maymyo district.
The measurements differ widely from those given by Selys for dzforata, but it
is obvious that these are incorrect, at least those for the abdomen. The Maymyo
specimens are very constant in their markings, especially with regard to the
extent of the opaque areas of wings, so that I think delimbata should at least
rank as a subspecies.
Rhinocypha biforata abbreviata subsp. nov.
In a male specimen from Assam, now in my own collection, and which I have
named adbdreviata, I note the following differences :—
The mesothoracic triangle is considerably smaller than in delimbata and the
carina above it is unmarked ; the antehumeral spot is absent; the forewings
have only the extreme tips opaque and the nodal index is considerably lower,
(12 antenodal nervures and 21 postnodals) ; the outer series of spots are discrete
and all are much shorter than for type (12 to 13 cells for costal spot in 6zforata,
only 6 in adérveviata, 18 cells in the second spot, only 14 in abbreviata, etc.).
The middle series of spots show the same difference, being only 4 to 5 cells in
length and falling well short of the opaque area, although in the hindwing,
this area is as extensive as in 6zforata btforata. Finally the proximal spot is
only 8 cells in length. The nodal index of hindwing lower, 11 antenodals and
23 postnodals,
INDIAN DRAGONFLIES 455
Abdomen with markings present on the first five segments only :—a large
triangular blue spot on the sides of segment 1, an apical rounded spot, a short
mid-latera] stripe and a ventro-lateral stripe on segment 2, segments 3 to 5 as
for 2 but the medial stripes absent and the other markings much reduced.
The sinall size of this insect, the abbreviated vitreous markings, and the
reduced abdominal markings are, I think, sufficient to form a well-defined
_subspecies, bearing the same relation to 6zforata as does trimaculata to
tonipennis.
Rhinocypha biforata beesoni, Fras, Mem. Dept. Agricult. Ind. (Ent. ser.) vol.
vii, No. 7, pp. 61 and 62 (1922).
Male. Abdomen 18mm. Hindwing 25 mm.
FR. beesoni shares the characters of 6zforata and delimbata, in that the apical
opaque area of the forewings is as extensive asin Ozforata, and the same area
in the hindwings is as reduced as in—de/imébata. In—the forewing the opaque
area extends 2 to 3 cells proximal of inner end of pterostigma, the inner margin
somewhat irregular, bevelled outwardly from costa posteriorwards and
prolonged slightly along the costa as far as half-way between pterostigma and
node between the costal and radial nervures. ‘There is also a slight prolonga-
tion along hinder margin of wing. Inthe hind wing the opaque area covers
nearly the outer third of wing and the middle series of spots extends right
up to it or even invades it for a short distance. The length of these spots is
slightly greater thanin adbreviata, but decidedly less thanin 62zforata or
delimbata. ‘The vitreous spots in all these subspecies have a metallic green or
coppery reflex in the opaque areas, violaceous in the hyaline area.
The discoidal cells are traversed 3 and4 times respectively, the forewing
has 14-15 antenodals, and 22-25 postnodals. The body markings are similar
to those of delimbata.
Distribution. Thetype in the Forest Research Institute, Dehra Dun, comes
from Lachiwala in that district, and was collected in the month of November.
I have also seen two pairs in the Pusa collection which were collected at Mergui,
Lower Burma, during June. ‘There areno records of its having been taken in
the vast stretch of country separating these two localities and it may be that
some error in tabulating has crept in, for Mr. C. F. Beeson collected in Lower
Burma as well as around Dehra Dun. It is easily distinguished trom delimbata
and aboveviata by the extent of the apical marking of forewing, and from
biforata by the much reduced opaque area in hindwing.
Rhinocypha bisignata, Selys. Syn. Calop. p.62. (1853); Id. Mon. Calop. p.
214, (1855) ; Walk. (Lzbellago bistgnata) List. Neur. B.M. vol. iv.
p- 648. No. 9, (1853) ; Kirby, Cat, Odon. p. 113, (1890); Laid. Ree. Ind.
Mus. vol. xiii, p: 38, (1917) ; Fras. Mem. Pusa. Dept. Agric. (Ent.
series) vol. vii, No. 8, pp. 80 and 81, pl. x (1922) ; Id. Rec. Ind. Mus.
vol. xxvi, p. 483 (1924).
Male. Abdomen 20mm. Hindwing 24-26 mm.
Head. Eyes brown, rest of head, including lips velvety black. A small spot
external to each posterior ocellus, a similar postocular spot on each side and
an occasional occipital spot all bright ochreous.
Prothorax black withalarge yellow spot oneach side of middle lobe, a
similar spot on the outer ends of posterior lobe and a large rose-pink spot
covering the greater part of dorstum of same lobe.
Thorax black marked as follows:—the mesothoracic triangle rose-pink,
extending for more than one-third, but for less than half-way up dorsum of
thorax ; a large oval spot of the same colour, pointed at both ends, bluatlyso
anteriorly, lying between the humeralsuture and mesothoracic triangle, but
well separated from latter, and of twice itslength; a small upper yellow
anterkumeral spot, a narrow yellow stripe bordering the upper half of humeral
suture behind; a broad very broken stripe on hinder half of mesopimeron and
the whole centre of metepimeron golden yellow. Beneath three pairs of yellow
spots, the hind the larger, the middle pair obscured. Legs black, the two
hinder pairs of tibicee and femora pruinosed white on flexor surfaces.
Wings long and narrow as in 6zforata, the hind slightly broader than the
fore, hyaline in the basal portions, where they are tinted with yellow, opaque
blackish brown in the apical areas, the hind bearing here a large vitreous
fuliginous spot. Forewings with the outer fourth or more opaque, this area
456 JOURNAL, BOMBAY NATURAL HIST.SOCIETY, Vol. XXXII
being briiliant coppery or with fiery reflex as in zguipennis, with its inner
border prolonged slightly basad between costa and Az and running obliquely
out towards the hinder border of wing. Hindwing with the inner third opaque
and marked with two series of vitreous spots which glow with a coppery reflex,
or violaceous, the inner spot always the latter colour. The outer spot produced
by a fusion of similar spots to those seen in 6zforata, but often completely or
partially divided into two or more spots, lying between (zz or WA or Riv+v,
and its outer border on a level with the middle of pterostigma. The inner
series composed of 3vitreous spots similar to those seen in dzforata but the
middle one usually prolongedinwards to fuse with the nodal spot seen in
biforata. Thefirst spot of this series one row deep by 8 cells long, lying
between /A72zz and Azv-+v, the middle spot 20-22 cells long when fused with
the nodal spot, otherwise only 10 cells long, lying between /Azzz and Riv+u,
the hinder spot 11-12 cells long, lying, between J7A and Cwzi. Pterostigma
black in all wings ; nodal index 12-14 antenodals and 28-31 postnodals in the
forewings, 12-14 antenodals and 22-28 postnodals inthe hind; discoidal cell
traversed 2-3 times in forewings, 3-6in the hind; petiolation begins slightly
proximad of the 2nd antenodal.
Abdomen black marked with yellow as follows :—segyment 1 with asub-
triangular lateral spot on each side; segment 2 with a midlateral stripe, an
apical spot and a ventro-lateral stripe ; 3 and 4 similar to 2, but the mid-lateral
spot obsolete, whilst 5 has the apico-lateral spot only. All other segments
unmarked.
Anal appendages as for genus, the inferiors minutely spined above.
Female. Abdomen 16mm. Hindwing 22 mm.
(Although so common an insect, and known forso long atime, the female
of this insect has hitherto escaped description).
Head. Labium bluish green tipped with black; labrum black with a large
oval or triangular yellow spot on each side ; cheeks broadly yellow, this colour
continued up as a narrow bordering to eyes as far back as the level of the
posterior ocellus, the basal segments of antenne, a small triangular spot low
down on sides of epistome, two large triangular spots on the upper surface of
frons, two transversely oval spots just behind these lying in front and to outer
side of anterior ocellus, all yellow; a small rounded spot on the outer side of
each posterior ocellus, a rounded postocular spot on each side and a mid-
occipital spot, triangular in shape all bright ochreous.
Prothorax biack with a large yellow spot on each side and the borders of the
posterior lobe finely yellow laterally.
Thorax as for male but the large antehumeral spot replaced by a fine ante-
humeral! line incomplete above, whilst the mesothoracic triangle is black, finely
outlined in yellow, as also the middorsal carina. Legs black, not pruinosed.
Wings entirely hyaline, tinted palely with yellow, the apices narrowly en-
fumed ; pterostigma black with pale creamy centre; discoidal cell traversed
twice, or more rarely, thrice in all wings ; 12-14 antenodal nervures, 21-27 post-
nodals in forewings, 19-24 in the hind.
Abdomen black marked with yellow as for male but rather more extensively,
thus segments 3 to Sare marked similarly to 2, and 6 and/7 similar, the
remainder unmarked.
Anal appendages and vulvar scale similar to genus.
Distribution. ‘This species is the sole representative of the genus in Southern
and Peninsular India. It is a widely distributed insect, thus I have records
of its capture in localities as wide apart as Cochin and the Central Provinces.
It is common throughout the Palnai, Nilgiri and Shevaroy Hills in the far south,
abundant in Coorg, less common at Khandala and Igatpuri on the ghats near
Bombay, !astly it is widespread throughout the Agency Tracts and Jeypore on
the East Coast. Its exact limits here are yet to be worked out but it probably
stops short of Bengal. It isa submontane insect, living and breeding between
2,000 to 5,500 feet altitude Its iarva was discovered inthe Nilgiris and—was one
of the first to be described of the genus. The females congregate on the
bare twigs of neighbouring trees in great numbers and are very rarely seen
pairing. Occasionally they may be seen ovipositing ona piece of dead twig
floating in mid-stream. ‘The males frequently disport in couples before the
females, circling and sparring with one another, their white pruinosed legs glis-
tening and thrust out like the arms of a wrestler seeking for an opening.
Meanwhile their wings flash like living jewels of fire.
Journ., Bombay Nat. Hist. Soc. Prats Ili,
EER EX
Wie. 2)
lanp 2. Two VIEWS OF THE CAUVERY RIVER NEAR FRASERPET, COORG,
WHERE Phinocypha bisignata AND Micromerus lineatus ABOUND. OTHER
DRAGONFLIES TAKEN IN THESE SPOTS WERE Chloroneura apicalis AND
guadrimaculata Zygonyx isis, Macromia indica, Macromidia donaldi, Onycho-
themis testacea ceylanica, ETC.
-
: :
-_ zi
aary 7
= 7 rs ;
rat ‘ A
ald . 4 :
1
. :
. - =
7 . an
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t
a
INDIAN DRAGONFIIES 457
Genus CALOCYPHA, gen. nov.
Moderately small but robust insects with characters similar to those ot
Rhinocypha, differing mainly in details of venation and the greater petiolation
of wings. Head, thorax, abdomen and legs similar to Rhinocypha. Wings
equal in length, similar in shape, apices markedly rounded, reticulation more
open than in Ahinocypha, hyaline with opaque apices; the arc very oblique,
lying almost in the same line as the costal border of disccidal cell, the latter
shorter than in most Ahinocypha and acutely pointed outwardly ; petiolation
beginning at or well distad of the inner end of discoidal cell; the nervure ac
situated more basad than in Aehinocypha ; anal appendages slenderer, the supe-
riors twice the length of segment 10, narrow, cylindrical, curving in at apices
which are bluntly pointed, furnished with a few hairs. Inferiors much shorter,
half the length of superiors, broad at base, directed directly back and parallel
to one another, furnished with a few minute spines on upper surface.
Female with apices of hindwings partially opaque, the venational details as
forthe male. Vulvar scaie and anal appendages as for genus Ahinocypha.
Calocypha laidlawi (Fras.) (AAznocypha laidlawt) Rec. Ind.Mus., vol. xxvi,
pp. 482, 483 (1924).
Male. Abdomen 17-18 mm. Hindwing 20 mm.
Head black marked with bright vermilion as follows :—a large oval spot on
dorsum of anterior part of epistome, a pair of large subtriangular spots behind
the former on frons, two large tongue-shaped spots on vertex nearly confluent
with former, closely apposed to the ocelli which they partially enclose. The
latter spots changing from vermilion to pale ochreous at their hinder ends; a
postocular spot on each side of occiput of the same colour. Eyes dark brown;
labium pale yellow ; labrum glossy black.
Prothorax black marked with creamy yellow as follows :—a narrow anterior
collar, an oval spot on each side of middle lobe, a spot on each side of posterior
lobe and a large spot on trochanters.
Thorax black, the mesothoracic triangle very narrow, very elongate, extend-
ing the whole length of dorsal carina, expanded in its lower third, with parallel
sides in its upper two-thirds, bright vermilion red. The central part of alar
sinus creamy yellow, laterally the lower parts of sides azure blue save for a
small triangular area posterior to the humeral suture and a small linear spot on
upper part of 2nd lateral suture. The dark area in front bearing a small upper
posthumeral blue spot. Beneath thorax six large rounded yellowish spots, the
anterior pair confluent.
Wings hyaline, the bases tinted with yellow, rather less than the apical thirds
of all wings opaque black with a dark violety metallic reflex, the margin of this
opaque area straight and sharply defined, not bearing any clear vitreous spots
or stripes. Pterostigma black, swollen outwardly, oblique at both ends.
Legs black, the flexor surfaces of the two hinder pairs of tibiz pruinosed
white, the same pairs of femora very slightly so.
Abdomen black marked with bright azure blue on segments 1to8. Seg-
ments 1 to 3 broadly blue on the sides from base to apical border, segments
4to 8 with a broad wedge-shaped spot tapering apicad but not reaching apical
border of segments, gradually diminishing in size from4to8. Segments
2 to § with narrow paired basal subdorsal blue lunules. Anal appendages black.
Female. Abdomenl17 mm. Hindwing 24 mm.
Differs from the male by the markings bright pale ochreous instead of ver-
milion and azure blue, and by its forewings entirely hyaline.
Head black marked with pale ochreous as follows :—two large oval spots on
labrum covering the greater part of its surface, the same series of spots on
upper surface of epistome and frons as seen in male, the bases of mandibles
and a narrow stripe alongside each eye, as faras level of ocelli, the basai seg-
ments ofantennz, asmallspot on each side of epistome, and lastly, the same
postocular spots as seen in the male,
Prothorax similar to male but the middorsum bearing a stripe which runs
from posterior lobe nearly to anterior end of prothorax.
Thorax black marked with greenish yellow as follows :—two minute spots on
the antealar sinus, the middorsa! carina finely, a fine antehumeral stripe, its
anterior end curling abruptly outward, a short posthumeral stripe, incomplete
below, astill shorter vestigial stripe just posterior to the upper part of first
458 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXI11
lateral suture, a long stripe traversing the whole length of sides, crossing both
sutures obliquely, its upper and lower borders very irregular and serrate.
Legs black, femora a dirty yellow on inner sides.
Wings hyaline, the hind with the apices broadly brown as far proximal to the
inner end of pterostigma asits own length, and of a much deeper tint in the
hinder two-thirds of wing. Anteriorly the cell-middles are hyaline, whilst at the
apex of wing, the brown area is replaced by a small opaque white area.
Pterostigma with inner half black, outer half whitish margined with black.
Antenodal nervures 10 to 11 in forewings, postnodals 18 to 19 (10 toll, and
18 to 24 in the male) ; discoidal cell of forewings traversed once only, rarely
twice, in both sexes, 2 to 3 times in the hindwings. Petiolation beginning at
inner end ot discoidal cell in the female.
Abdomen black marked with bright ochreous and greenish yellow :—segment
1 with a large lateral spot and a minute middorsal linear apical spot, all seg-
ments from 2 to 8 with the middorsal carina finely yellow and from 2to7 with
a long narrew stripe followed by a large apical spot on each side, whilst
beneath this marking, on all segments, is a superventral stripe ; remaining seg-
ments unmarked.
Anal appendages black, long, acuminate. Vulvar scale as for genus.
Distribution. Confined to South Kanara, to the net-work of rivers about
Sulia. I well remember the first discovery of this beautiful insect, when in
company with Mr. C. A. Souter, I was making my way up the bed of the river
in the Coorg—S. Kanara enclave. We had come to a fork in the stream and
agreed to part company, each to work his own branch of the fork. Shortly
afterwards I came on my first specimen which I mistook for a Pseudagrion
rubriceps. Its size and conspicuous colouring led me however to glance at it
a second time, when I at once recognized that it was something altogether new.
I took eight males in all and when I rejoined my fellow shikari, found that he
too had made the same discovery, having taken four more males (Nov. 9, 1923).
Mr. Souter discovered the female on this same riverin the following April, where
I again found it later. For a long time, the discovery of a second species of
Rhinocypha in South [India has been visualized by Dr. Laidlaw and myself, so
that it was very gratifying to come upon this insect. Dr. Laidlaw has pointed
out to me the marked differences in venation and has suggested the removal of
the insect from genus Ahinocypha, a suggestion here carried out. The type and
co-type in the British Museum. Paratypes in the Fraser, Ris, Laidlaw, Morton,
Pusa and Inglis’ collections. The insect is a shade lover, albeit its bright
colouring. It is found perched on half-submerged logs in midstream, or
resting on twigs overhanging the river. The reds and blues are very con-
spicuous when the insect is in flight, whilst the black tips form an a]most com-
plete circle round it as the wings whirl with cinema effect.
Genus, LIBELLAGO, Selys.
Selys, Syn. Cal., p. 57 (1853) ; Id. Mon. Cal. p. 226 (1854) ; Walk. List,
Neur. Ins. Br. Mus. vol iv, n. 643 (1853).
Moderately robust insects with characters very similar to genus Ahinocypha
but differing less from that genus than does Calocypha.
Head, thorax and abdomen similar to ARhinocypha, but mesothoracic
triangle undeveloped and uncoloured ; the abdomen, on the other hand, of the
males, usually brilliantly coloured. Wings entirely hyaline in both sexes,
narrow, the hind not noticeably broader than the fore, venation close, similar
to that of ARhinocypha, except J7A which is more or less zig-zagged for at least
the outer part of its course, so that the cells bordering it are pentagonal instead
of rectangular as in Rhinocypha ; petiolation ends well proximad of the inner
end of discoidal cell. Legs variable, similar to Ahinocypha in the single
Asiatic representative, similar to A&hinocypha, or widely dilated, in African
species and usually brilliantly coloured. Anal appendages and genitalia very
similar to Rhinocypha.
Distribution. Orly a single species has been recorded from Asia, ZL. aszatica
vittata, which extends through Burma and Tonkin, tothe Philippines where
it is represented by ZL. astati¢a astatica. The gerus although so closely allied
to Rhinocypha, is essentially an African one, and it is doubtful whether aszatzca
should be included in it, as the venational differences seperating it from
Rhinocypha are not nearly as marked as they are in the African species of the
INDIAN DRAGONFLIES 459
genus, nor is the abdomen marked as brilliantly asin the latter. The Asiatic
species may be regarded as a link bridging the gap between the two genera, and
raises a point of profound interest as to how, when and where the passage took
place. (Genotype—Libellago dispar Beauv.
Libellago asiatica vittata Selys, Ann. Mus. Civ. Genov. (2a), x, (Xxx),
pp. 58, 59 (1891); Will. Proc. U.S. Nat. Mus., vol. xxviii, p. 173
(1904) ; Mart. Mission Pavie, Neuropteres (sep.), p. 17 (1901) ; Laid.
Rec, Ind. Mus., vol. xiii, pp. 39, 40 (1917).
Male. Abdomen 23mm. Hindwing 33 mm.
Head black unmarked ; prothorax black unmarked.
Thorax black with a fine posthumeral yellow stripe and two oblique dark
ochreous bands on the sides. Legs black.
Wings hyaline suffused with brown at extreme bases, especially in the
subcostal and median spaces as far as the basa! antenodal nervure; 16
antenodal nervures, 19-22 postnodals in forewings, discoidal cells traversed
twice or thrice ; pterostigma black, about 2 mm. in length.
Abdomen black marked with reddish brown, segments 1 and 2 entirely black,
dorsum of segments 3 to 8 reddish brown, the sides yellow, as well as the
dorsal carina finely ; apices of segments narrowly, and the whole of segments
9 and 10 black. Anal appendages black, superiors twice as long as segment
10, slender, curving in towards their apices which area little spatulate ;
inferiors cylindrical, separated, curving in at apices, about half the length of
superiors.
Female. Abdomen 20 mm. Hindwing 30 mm.
Head. Labium black; labrum black witha large diffuse rounded central
spot of pale yellow ; bases of mandibles, a large spot on the sides of epistome,
another on the sides of frons, the basal segments of antennae, the cheeks
broadly, a small oblique spot on the outer sides of the posterior ocelli and a
rounded postocular spot on each side of occiput pale creamy yellow.
Prothorax black, the lateral ends of posterior lobe and a large spot on each
side of middle lobe pale yellow.
Thorax black marked with apale greenish yellow antehumeral stripe
incomplete above, where however, is an isolated spindle-shaped spot of
ochreous ; a bright ochreous elongate posthumeral upper spot, at the lower end
of which, and on the dorsal side of the humeral suture, is a somewhat smaller
triangular spot ; laterally the whole of the lower half of mesopimeron, and the
whole central portion of the metepimeron as well asa small curved stripe
behind the upper part of the first lateral suture bright ochreous. Beneath
thorax a broad transverse stripe behind the legs, followed successively by a
large arrow-head shaped spot and a broad terminal stripe all ochreous.
Legs black, coxe and trochanters yellow, proximal ends of all femora
yellowish brown.
Wings hyaline, the extremity of all apices clouded narrowly with blackish
brown ; 15 to 16 antenodal nervures in forewings, and 17 to 18 postnodals, 16
antenodals and postnodals in hindwings. Discoidal cells traversed thrice ; the
nervure ac well proximad of inner end of discoidal cell; petiolation begin-
ing at level of 2nd antenodal ; pterostigma blackish brown, longer than in
male.
Abdomen black marked with yellow as follows:—segments 3to 8 with a
narrow middorsal carinal stripe, segments 1 to 7 with the sides broadly yellow,
this colour expanding dorsalwards at apical ends of segments ; segment 8 with
a rounded apico-lateral spot.
Anal appendages black, long and narrow, pointed. Vulvar scale as for
Rhinocypha, black with a large medio-lateral bright vellow spot.
Distribution. Bbhamo, Burma, within Indian limits, June and July.
Reported also by Martin from Tonkin. Type inthe Selysian collection.
Appears to be a rare insect wherever taken.
(70 be continued.)
Note :—In part xxvii, Indian Dragonflies of this volume, by a printer’s error,
the figures have been transposed. Fig.1 should be Fig. 2as regards the
explanatory text, and vice versa.
A SIND LAKE
BY
SALIM A. ALI
Assistant Curator, Bombay Natural History Soctety
(With stx plates and three Text figures)
This is the narrative of a delightful New Year week spent on the
Manchar, perhaps the largest fresh water lake in India, and certainly
the most enthralling from the point of view of the sportsman and
ornithologist.
Situated in the Larkana District, the Manchar, like all the other
numerous dhunds (jheels) of Sind, owes its existence principally to
the annual inundations of the river Indus. A portion of the Indus
waters teaches the Mancnhar by a natural and very tortuous channel
known as the Western Nara, and again finds its way back to the
parent river through another—the Aral. This latter commences
at a point near the dusty little town of Sehwan whose chief pride at
the present day lies in the dilapidated ruins of a mud-and-brick fort
claiming an abstruse connection with the invasion of Alexander
the Great, and in the annual fair held at the dargah of a pzr
long-defunct but who continues to perform his miracles
notwithstanding.
The Manchar, in other words, may be regarded merely as a
local expansion of the Western Nara Canal; during inundations
when the level of the Indus rises much higher than that of the lake,
the Aral discharges water into the Manchar; when these floods
subside, the current is reversed and the water carried back to the
river. A vast expanse of land—20,000 acres or more—is exposed
by the post-monsoon drying or draining off of the lake which
is extremely fertile and valuable for the cultivation, of vadz crops
such as wheat aud gram. It is these cultivations, incidentally, that
afford attractive pasture to the innumerable hosts of geese and
cranes which constitute such a salient feature of the bird-life on
this wonderful lake.
Unfortunately for the bird-lover, by reason of this very fertility
of the soil left high and dry by the retiring floods, the days of the
Manchar as a wildfowl resort are numbered. A mammoth scheme
is afoot for draining off the lake more or less completely in the
cold weather so as to render the greater part of its bed available
for cultivation. The disappearance of these extensive feeding
grounds will doubtless produce a marked effect on the bird-life
not only of the district but of the province of Sind as a whole,
especially as the project is expected to come into operation
simultaneously with the mighty Sukkur Barrage and its intricate
network of canals, with the consequent opening up of enormous
tracts of hitherto unpopulated country.
As a whole, the waters of the Manchar are shallow, ranging froma
few inches in depth to eight or nine feet. The surface of the marginal]
a SIND LAKE 461
shallows is covered with tangles of the floating lotus plant (chiefly
Nymphea lotus and Nelumbium spectosum) which grow in unbounded
profusion in every dhund in Sind. These in the Manchar yield
a considerable revenue to Government. The roots of both, called
Lorh and Beeh are extensively used as food, as a substitute for
potatoes and in many other forms—whilst their seeds or nuts (Vago
and Pédbiro) are also much eaten and sold in the bazaars.
Apart from these littoral lotus tangles, the shallows abound in
reeds and sedges affording ideal shelter to countless myriads of
wildfowl of every description which, driven by the bitter cold and
scarcity of food from their northern homes, migrate southwards and
find an agreeable haven in their hospitable seclusion.
We were encamped in the District Local Board bungalow at Shah
Hassan, a tiny village on the western shore of the Manchar,
between its waters and the desolate looking Kirthar Range which
forms the natural boundary between Sind and Baluchistan. The
road to Shah Hassan from our iast camp ran for a good way by the
margin of the lake, and all along in the distance could be descried
millions of waterfowl, floating on the water lumped together, look-
ing like immense floating islands, while the terrific honking,
quacking and ‘spattering’ set up by the birds sounded like the
distant grind of some weird machinery. Every now and again there
passed over or in front of us within distinguishing range, flocks of
various species of duck, and skeins of Grey Lag Geese leisurely
winging their way lakewards from the direction of the Kirthar where
they retire nightly to feed on the beloved grit which the hills supply.
A chronicle of one of these memorable days spent on the lake
will suffice as sample.
We left the bungalow soon after daybreak in punts, each punt
manned by one gun, a shikari and three mohanas with poles. The
mohanas or ‘ mirbahrs’ (literally ‘ sea-Jords’’ or ‘Admirals’ ! ) are
Mohomedans of what one may call an amphibious species. They
spend their lives on or near the River Indus or one of the numerous
dhunds created by its monsoon inundations, and eke out a bare
livelihood by plying boats, gathering lotus seeds and roots, fishing
and capturing wildfowl in a number of ingenious ways, some of
which are hereafter described. Some of the more cnterprising ones
also go in for farming egrets, an industry which holds much promise
when the traffic in plumes is legalized. Fatigue is a thing unknown to
these ‘ sea-lords’ and it is remarkable how they will go on punting
almost incessantly all day without exhibiting any symptoms of it.
We glide placidly over the limpid waters in the direction of the
butts that have been prepared overnight in the neighbourhood of a
bank where Grey Lag Geese have been observed to congregate in
the mornings before repairing to their favourite feeding grounds
among the tender gram and wheat fields hard by the margin of the
lake. A piercing north-easterly wind is blowing which strikes our
faces like needles, and pierces through great-coat, tweed jacket, and
thick Jaeger vest to the very marrow of our bones. We sit on the
bow of our punt with our heads snugly buried io the ears in the
coljlar of the trusty great-coat and our hands thrust well down its
grateful pockets. In our course, we disperse immense flocks of
9
462 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXH
coots, with which are mingled odd parties of gadwall and other
species of duck. The Manchar is, and has long been, notorious for
the masses of coots it harbours, but their numbers are totally incon-
ceivable until you actually see them and can believe your eyes. Great
patches of the lake are literally blackened with their multitudes and
their ‘ spattering ’ on the water as they attempt to rise, is positively
bewildering to the new arrival. Inspite, however, of the wholesale
slaughter and destruction that proceeds apace from the time the
coots arrive till their return migration at the commencement of the
hot weather, their numbers appear to remain unaffected, and the
legions that visit the country again the following season show that
the coot is one of those fortunate species that is in some way ex-
ceptionally well-fitted in the struggle for existence, and stands in
no immediate danger of extinction.
But to continue with our narrative. Owing to exceptionally
heavy inundations of recent years, the greater part of the thicker
reed-beds on the Manchar which erstwhile provided such admirable
cover to the sportsman have disappeared, rendering satisfactory
sport none too simple a proposition. The butts or ‘hides’ in the
present instance consisted of a few twigs of tamarisk thrust into the
lake-bed to conceal the punts, there being no natural cover at hand.
These were arranged in a row at distances of 40 yards or more
from each other. We soon take up our positions, and at a given
signalthe drive commences. Some of the punts have gone ahead
to round off the birds, while others are stationed at a distance in the
rear to turn the birds back over the guns. The first to take alarm
are a party of Pintails—all males keeping together. They fly low
and unsuspectingly over the central ‘hide’ from where the right
rings out, followed by the left in quick succession. ‘These are
accommodatingly responded to by splashy thuds as two of the fat
birds pay the penalty of blissful ignorance. The flock breaks up
to right and left at lightning speed, and the birds lose no time in
rising well out of range.
The reports of the gun rouse the other habitués of the lake into
action and now follows a sight that can never be forgotten—a sight
that must leave an indelible impress on the mind of every sports-
man who shoots on the Manchar. The birds rise in their thousands
to the accompaniment ofa veritable babel of flapping, swishing,
spattering, honks, quacks and a variety of other noises. Nowcome
the pochards and the shoveilers, spotbills, mallard and teal ali in a
jumble, some swishing past in orderly flocks overhead, others flying
singly or in twos and threes. The clouds of mixed species that rise,
soon resolve themselves into little parties of their own. Unfortu-
nately the wind keeps up and spoils the fun. The birds get steadily
higher and higher and are soon quite out of range. Our gunners
have been feverishly active all this while, and the firing has been
fast and furious. The average of kills however, dwindles off appal-
lingly after the first few fusilades, and in the later stages of the shoot
the sportsman has to rely more upon his luck than on his skill.
In a very short time the birds have disappeared. The tiring now
becomes sporadic and far between as an occasional mallard or
white-eye returns blundering past to reconnoitre ground so lately
Journ., Bombay Nat. Hist. Soc. PLATE |
1. NEGOTIATING THE REEDY MARGINAL SHALLOWS.
Up sennpiin:
NS ANON OA XS.
2. NEARING THE BUTTS.
(Note masses of wildfowl in the background).
Journ., Bombay Nat. Hist. Soc.
4. THE FOWLERS ABOUT
TO ENTER A FLOCK.
Prate if
A SIND LAKE ; 463
evacuated. Very good sport may sometimes be had at this stage,
as the gunner need be in no hurry, and the tendency to ‘ brown’
flocks, which is perhaps the most important cause that contributes
towards unsatisfactory averages, is eliminated.
None of the Grey Lag Geese which formed such an imposing
array on the mud-banks in our front have come our way during the
drive except a pair, of which one fell and the other was disgracefully
missed. During operations, a great many skeins have been noticed
flying over, well out of range, in the direction of the young wheat
and gram on the farther margin of the lake. The masses that rose
at the first shot have also madea bee-line for these parts. A
hasty confab of the shikaries decides that our acquaintance with
the birds might be renewed with advantage, and we push off. Ali
the way in the punts, flock upon flock of the geese are observed
leisurely winging their way towards their rendezvous flying
studiedly out of range and frankly disdainful of our murderous
aspirations. Notwithstanding the obvious, the more enthusiastic
section of our party have been totally incapable of restraining the
trigger-finger. On the universally accepted and sound principle
that accidents wz// happen in the best regulated households, the
Mannlichers are brought into action, but magazine after magazine
has failed to disturb the complacence of the assaulted, who doubt-
less regard it all as a rather meaningless and totally unnecessary
demonstration on the part of the human biped.
True, one ‘accident’ did happen. A stray bullet caught the
leader of a wedge plumb in the centre of his fat body blowing his
back completely away, and the bird arrested dead in his flight as if
by a stroke of heart failure, hurtling down like a stone through 500
feet or more of space presented a remarkable spectacle. ‘This
event of course roused hopes which conduced to dissipate ammuni-
tion in the shortest possible time !
In an hour we are on the farther shore. Punts are discarded and
a rigorous tramp of about three miles over slushy ground brings us
within reach of our objective. By the time we arrive at our destination
it is 10.30. ‘lhe distant fields, as far as the eye can reach, are one
seething mass of jostling Grey Lags, but the total absence of
suitable cover presents a problem. ‘The shikaries however are up
to the game. They put their heads together and we soon find our-
selves hustled along to within a few hundred yards of the birds—
now crouching, now crawling on all fours, now halting motionless
then again running cautiously forward taking the fullest advantage
of every unevenness the ground offers. Here we are made to squat
behind diminutive tussocks of grass in ankle-deep water, all in aline
at intervals of a hundred yards or so. The beaters who have gone
ahead in the opposite direction, seeing us safely ensconsed, com-
mence walking towards the birds. The foremost to scent trouble
are the odd parties feeding on the outskirts who rise, as often as
not to settle again after circling the fields. When the men have
managed to edge themselves more or less into the centre of the
congregation, the shout is raised. The sight that now ensues com-
pletely baffles description. Momentarily your horizon is obscured
by dense masses of Grey Lag Geese and the din of their honking is
464 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
such as to render the various excited injunctions of your shikari
positively inaudible. The birds spread themseives over a vast
front. The guns have been placed directly in their line of retreat
to the lake, and you presently see a few thousands coming
your way. You crouch and prepare to send up your greetings,
cylinder and choke, but before you have been able to gauge the
range, the great multitude have swept over, and now come smailer
skeins which happily are flying lower. You hear shots from your
neighbours and occasional heavy splashes in response. The birds
swerve to right or left, but before they can rise much higher they
are directly over you. If you remain cool and collected and pick
out your birds before firing, a couple of rights and lefts will bring
you two brace without difficulty. If ambitiously inclined, however,
and aspiring to a record bag by ‘browning’ the flock, you wilias
likely as not be left cursing your stars with a clear blue sky above and
precious little to compensate for it in the nature of birds in hand !
This briefly is the typical programme of a day’s shooting on the
Manchar, which with sundry variations may be experienced by all
sportsmen who shoot on the lake.
A couple of days were spent in watching the wildfow] netting and
fishing operations of the mohanas, which proved both enjoyable and
instructive.
The flesh of the coot, though somewhat rank and fishy to ‘ sophis-
ticated’ palates is held at a premium by the Sindhi, especially of the
poorer classes. He prefers it toduck and other wildtowl, and the
coot accordingly is a coveted object of the mohana’s attentions and
devices.
There are numerous little villages and hamlets by the shores of
the Manchar and most of the larger dhunds in Sind, the inhabitants
of which subsist throughout the cold weather months either on the
flesh of this unfortunate bird or otherwise by the wherewithal pro-
vided by traffic therein. In the neighbourhood of every such
village may be seen masses of coots’ feathers which bespeak the
sad fate of the many gone before. Coots sell in the villages at
about two annas apiece, and owing to their ready demand, bring
brisk business to the fowler. A véry common method of capturing
the birds is for a man to wade into the shallows frequented by the
coots, wearing on his head in the form of a cap, the complete skin
of a duck with head and neck properly set up, and provided with
breathing and eye-holes for the fowler. Under cover of this decep-
tion he stalks gradualiy up right into the midst of an unsuspecting
flock, nothing visible except the harmless looking decoy on his
head. When well within the assembly, the fell work commences.
A bird is grabbed by the leg under water and in the twinkling of an
eye, before it has time to flutter, the victim disappears below the
surface. Its legs are bound up with water weeds and the captive
firmly secured to a belt round the fowler’s waist. The other mem-
bers of the flock have noticed nothing of this foul treachery, and are
as confiding as before. Presently a second bird disappears, a third
follows rapidly, and then a fourth. Shrewd suspicions now begin
to cross the minds of the silly coots and misgivings become appa-
rent as to things being as they should ; there is certainly something
‘NOILIV NI $1205 AH,
9
‘HOLVO InassHooaS YW °S
"20 "}SIP] yen) Aequiog ‘-uanor
Journ., Bombay Nat. Hist. Soc. PLATE IV
7. RED-CRESTED POCHARDS IN A Dhubbi NET.
8, REMOVAL OF CAPTIVES FROM Dhubét.
siaiade
A SIND LAKE: *.° °° hee 465
wrong somewhere! A number of the birds spatter away some
distance from this mysterious duck, but are too dense however to
detect that unlike other self-respecting ducks, this one of. their
suspicions that follows steadily, progresses backwards or sideways !
The eye- and breathe-holes for the fowler are placed near the vent
and undertail coverts of the skin, so that the decoy and the man
below are facing in opposite directions! When the fowler has
gathered half a dozen birds or so below the water, he fastens their
legs together and moving off some distance from the flock, he
allows his ring of captives to come to the surface for breath. It not
infrequently happens that when sport is good and the fowler
ambitious, some delay occurs in this very necessary operation with
the result that several of the hapless prisoners are drowned.
Leaving the knot of captured birds on the water, the wily mohana
again sidles into the flock and the process is repeated. Thus it is not
unasual for twenty or thirty birds to be taken by a single man in the
course of a morning’s work. Duck are also taken in this manner,
but the stupidity of the coot lends itself particularly to this
device, and they make by far the more ready victims. By way
of a variation, in place of the duck-skin cap a tame cormorant is
frequently perched on the man’s head, and the bird appears to parti-
cipate whole-heartedly and to enjoy the sport thoroughly. This
method of fowling is of great antiquity and has been pursued
through the ages practically unchanged in every detail. It was
greatly in vogue in the time of the Emperor Akbar.
A method of netting wildfow] on a somewhat larger scale is
known as the Diubdi. Here a net, about 30 feet 20 feet is stret-
ched on the shallow bed of the lake at a favourite feeding ground
of the birds. A quantity of grit from the neighbouring barren hills
is sprinkled over the net by way of bait. A drag-line, 200 yards or
more, is attached to one end of the net by the pulling of which the
net closes up in the form of a Gladstone bag.
Ly
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net closes A meels B and C neels D.
Fug. 1.
When
Drag Line
ST ae _ 7 =
466 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXX
The fowler withthe loose end of this line conceals himself and
his punt ameng the reeds at a distance and keeps a vigilant look-out.
While ‘tipping ’ about for food, the presence of the grit is dis-
covered by the birds and this marks a general scramble for the spot
within the sphere of influence of the hidden net. Now is the time
for the fowler to get busy: the line is pulled steadily and gradually
till one final and mighty heave leaves the birds entangled. One of
my photos shows thirty-five Red-crested Pochards taken ina net of
this size. The fowler now approaches, and removing one captive
after another, proceeds in the most matter of fact manner to inter-
lock their wings—twisting one wing round the other over the
back—and to dump the birds mercilessly into his punt prior to pack-
ing them up in crates for despatch to Karachi and other markets.
On a vaster scale, and for business on wholesale lines, the modus
operandi of the mohanas is as follows :—
A flight-net about 12 feet high and a quarter of a mile or more in
length is stretched on upright poles across a portion of the lake in
the proximity of some favourite feeding ground, with due regard to
direction of wind, etc. This net is turned up at the bottom, just
below the water-line, to form a continuous bag running all along the
the lower border. A quantity of the beloved grit is dropped along
the length of this net. Ducks, geese, coots and other wildfowl
gatherin their hundreds tofeed on this spot at night, and under
cover of the darkness a number of punts move up in line for-
mation from the opposite direction collecting and driving the un-
conscious birds towards the net. When the punts have advanced
to within thirty yards or so of this ‘barrier of twine’, a signal is
given and a terrific uproar set up. Lighted faggots and torches
are all of a sudden hurled at the birds who rise in distraction and
attempt to flee in the opposite direction. Before they are able to get
well under way and rise high enough to clear the cryptic net, they
dash into it headlong to be thrown down fluttering into the gaping
bag below, whence they are soonextracted by the mohanas who
hurry forward anon. In this manner athousand or more birds may
be taken in the course of a single drive.
A rather sporting method of procuring coots for the pot is
frequently indulged in by the mohanas. The gadgets employed in
this sport are a powerful bow made of bamboo and a number of
blunt, headless, unfeathered arrows, about three feet in length and
three-quarters of an inch thick. The hunters approach a flock of coots
in punts and divesting themselves of their clothing to very nearly the
irreducible minimum, launch into the waist-deep water where they
crouch submerged up to their chins in a row at intervals of fifteen
yards or thereabouts. The punts move off to round up the birds and
drive them towards the bowmen or Gazisas they are called. For
some time the coots continue to swim away from the advancing
punts and in the direction of the archers, but on being pursued
closer, they attempt to rise. They spatter along the surface before
taking off, and by the time they are over the line of bowmen they
are scarcely higher than fifteen or twenty feet. The men who
have hitherto been immersed to their chins with bow and arrow
in readiness under water, suddenly pop up and discharge their
Journ., Bombay Nat. Hist. Soc. PLATE V
10. INTERIOR OF THE SAME,
Journ., Bombay Nat. Hist. Soc.
11. FISHING ON THE MANCHAR—LANDING THE CATCH.
12. ANOTHER SNAP OF THE LANDING OPERATIONS.
467
He wiil spend the
A SIND LAKE
The arrows strike the birds crosswise, i.e., on the flat,
and often with such force as to take a wing clean off. The men are
~
four or five birds.
The mohana’s methods of fishing also afford interesting watching,
and the industry and hard labour entailed by some of the Jess whole-
sale systems leave one in strong doubt as to whether after all the
zeal displayed is not worthy of a better cause!
best part of a morning—his family in full strength participating —
such adepts at the game that misses are a rare occurrence, and a
volley from half a dozen bowmen seldom accounts for less than
rocking his boat from side to side and creating the most deafening
and unholy row with tom-toms and copper ‘dekchies’ in attempts
to drive the fish into the nets he has put up at a distance, and when
missiles.
Direction °f drive
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Fig. R
With
pull out a single fish or sometimes a couple—a catch not worth
everything has gone off well and ‘ according to programme ’, he will
more than four annas!
468 JOURNAL, BOMBAY NATURAL HIST, SOCIETY, Vol, XXXII
In one of the methods most commonly employed, the nets are
arranged as shown in Figure 2.
The fish are finally driven into the enclosed spaces marked A and
B whence the mohanas land them by diving, or ifthe water be
shallow, by means of a hand net like the one shown in the photo
(Noes)
The most important fish of Sind is the Pala (Clupca ilisha). Itis
a sea fish belonging to the same genus as the herring, and ascends
the Indus in the months of February and March for spawning in
enormous numbers. Fora short period at this time, this species
fairly swarms the Manchar. Next in importance come the Dambhro
(Labeo rohtta) which may grow to a length of three feet, and the Worz
(Cirrhina mirgala). Two specimens from the lake sent to the Indian
Museum, Calcutta, have been identified by Dr. S. L. Hora as these
species.
When the catch is not large enough to warrant its being carried to
market, the fish are kept over till the next day, the day after, or
even longer. The mohana, on withdrawing a fish from the landing
net, probes deep into the animal’s side just below the spinal chord
with a large skewer-like needle. Having thus paralysed his captive,
he proceeds in the most callous manner imaginable to drive this
same needle threaded with twine through one eye-socket and out
through the mouth. The fish along with other hapless ones is now
secured to the punt and the bunch is left floating listlessly on the
water in a comatose condition till gathered in and carried off to
market. It is thus that the mohana always manages to supply
‘fresh fish’!
Apart from duck-shooting, which is par excellence the sport of Sind
in general and of the Manchar in particular, by far the most en-
grossing charm of the Manchar lies in the unequalled variety of bird-
life it presents. I mention only a few of the species met with in
the course of a day’s casual observation when the main object was
shooting. This is only a very small fraction of the species I was
able to recognize, and in addition there were a host of others which
could not be identified for obvious reasons.
Flocks of Starlings, chiefly from five to twenty birds but some-
times hundreds, kept about the margins of the lake feeding on the
semi-marshy grassland whence the water had lately receded. A
specimen obtained proved to be Srurnus vulgaris nobilior.
Wagtails were plentiful, both along the margin of the lake and in
the shallows overgrown with the lotus plant, where they could settle
on and hop about from one floating leaf to another. The common-
est species—specimens of which were secured and have since been
identified by Dr. C. B. Ticehurst—were: (1) The Indian White
Wagtail (Aotacilla alba dukhunensts); (2) The Blue-headed Wagtail
(M. flava melanogriseus); and (3) The Yellow-headed Wagtail (JZ.
cttreola).
The Pied Kingfisher (Ceryle rudts leucomelanura) was perhaps the
commonest kingfisher. Seen mostly in parties of four or five.
Extremely pretty to watch as he hovers stationary over the water
and hurls himself head foremost at some unsuspecting fish from a
height of fifteen to twenty feet. The small Alcedo atthis pallasi was
aE Sa :
A SIND LAKE 469
also notuncommon. His tactics as a general rule differ fronr those
of his pied cousin. He sits on a branch rarely five or six feet above
the surface and drops perpendicularly on a passing fish with which
he flies off to a neighbouring perch.
The Indian Purple Moorhen (Porphyrio poliocephalus poliocephalus) :
This bird was an extremely common frequenter of the reed beds,
where it must breed. An immature male was caught by the
mohanas by surrounding a patch of reeds in which it had taken
refuge, being unable to fly well. It looked lean and lanky. The
iris were brown; bald patch on forehead and bill] black, slightly red-
dish on lower mandible. Legs and feet pale orange brown.
Coot (Fulica atra atra): As mentioned before, this is without
doubt the most abundant species onthelake. Besides the incessant
ravages of man, a great many fall victims to the numerous species
of accipitrine birds that ceaselessly beat over the waters of the
dhund. Iwas the fortunate witness of a raid ona flock of coot by
a large eagle, the species of which I was unable to ascertain owing
to the distance. It was I think either the Imperial Eagle (Aguzla
heliaca) or the Large-spotted Eagle (Aguzla clanga) both of which
were common about the locality. The plan of attack was interesting.
The eagle swooped down on the compact flock (about 200 birds)
and in this first bout broke it up. Several of the coots spattered
off, some dived below the surface while others swam away as fast
as possible from the scene of assault. The eagle renewed the attack
three or four times, till, having made confusion worse confounded, it
succeeded in isolating from the flock an individual which promptly
dived. This was the assailant’s opportunity. After a couple of
unsuccessful stoops, whenever the coot appeared on the surface for
breath, the eagle bore down on him like a hurricane and carried
him away in his talons.
In this particular case I was able to notice nothing of the singular
manner, often described, in which coots are said to defend them-
selves against eagles. It has been recorded that on the appearance
over them of one of these birds, the coots collect in a dense body
and when the eagle stoops at them, they throw up a sheet of water
with their feet and completely baffle the enemy.
No cranes, neither Megalornis grus liltordt nor Anthropotdes virgo,
were seen. I was informed that it was as yet somewhat early for
them and that they swarmed the localities, especially round our
latter camp, Janghar, in January and February when the crops were
fully grown.
White-tailed Lapwings (Chettusza leucura) were fairly common
along mud banks, feeding in companies of four or five birds.
The Black-tailed Godwit (Limosa limosa limosa): This bird
was extremely abundant on the shores of the lake near Shah Hassan.
Their numbers must run into tens of thousands, and large patches
near the margin of the water were literally obscured by their
multitudes.
The Caspian Tern (Hydroprogne caspia caspia): Numbers of
this beautiful tern were seen flying over the lake. When one of the
specimens I shot dropped into the water, it was immediately swoop-
ed upon by a Marsh Harrier (Czrcus eruginosus) which is perhaps the
10
470 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
most numerous species of the Accipitres about the lake. This bird
was also secured.
The Black Ibis (/uzocotis papillosus): Flocks of from twelve to
twenty birds, feeding on the drier portions of the lake margin.
The Glossy Ibis (Plegadis falcinellus): Parties of twelve to fifteen
birds appeared to be the usual thing. When flying away from or
coming towards one in their usual V formation, they are easily mis-
taken for geese at a distance.
The Large White Egret (Zgvetia alba): Only two pairs were
seen and one specimen secured. The birds were very shy and
wary. Measurements of this specimen were as follows: Bill from
forehead feathers, 127 mm., tarsus 214 mm., bill yellow; legs and
feet greenish-black.
The Bar-headed Goose (Amser indicus): It was significant that
during the three or four days spent at Shah Hassan not a single
bird of this species was seen. When camp was shifted to Jhangar,
about eight miles eastward, it was found that all the geese
frequenting this portion of the dhand were indicus and no Grey
Lags were observed. The two species appeared to have appor-
tioned off distinct zones. The numbers of these birds were consider-
ably less than those of the Grey Lag at Shah Hassan. Some large
flocks were seen feeding on the young wheat and gram by the dhand
in the early mornings and at sundown. ‘They feed throughout the
night and repair to the open water during the day where they ride
on the surface and rest.
Among the ducks, the commonest species by far appeared to be
the White-eyed Pochard (Wyvoca ruta rufa). This bird is an expert
diver, and unless squarely hit it dives and holds on to the sub-
merged water weeds andis then extremely difficult to recover.
The Little Grebe (Podiceps albtpennis): This little bird is also ex-
tremely abundant in the shallows. Flocks of fifty to sixty birds were
common, and the rapidity with which they dived was indeed remark-
able. When fired at, in many cases they had almost disappeared
below the surface before the charge reached them! My diary records
the following peculiarity as regards the colouration of the irides in
fresh specimens which in the Fauna (Azrds, Ist edn. vol. i, p. 475)
is stated to be ‘red-brown’.
tng bo fs fs
Some of the other birds seen on the lake and in its precincts
were :
The Spoonbill (Platalea leucorodia major); Large flocks on mud
banks,
A SIND LAKE 471
The Common Heron (Ardea cinerea cinerea): Quite common.
Seen singly.
The Desert Buzzard (Buteo buteo rutiventer): Seen usually in
pairs perched on mounds near the lake. Obtained a specimen.
The Pale Harrier (Circus macrourus): Extremely common.
Continuously beating over the shallows everywhere.
Before concluding, a few hints as to how a shoot on the Manchar
can be arranged may prove useful to intending sportsmen.
It is best to get into communication with the Deputy Collector
of Sehwan (Larkana District) and inform him well in advance of the
dates on which you propose to shoot. The stations to get down at
are either Sehwan or Bubak Road on the N.W. Rly. (Karachi-Lahore)
at both of which places tongas can be easily got. If you leave
Karachi at 11 p.m. you reach Sehwan at 9 a.m. the next day, and
Bubak Road is only 20 minutes further on. About Rs. 5 per day
for a punt and crew would be enough.
There are no bungalows on the Manchar. The P. W. D.
Bungalow at Bubak might be used. The Local Board Bungalow at
Shah Hassan where we stopped is a long way from the Railway.
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA
BY
R: 7. Pocock. F.R.Ss
(With 1 coloured and 1 black and white plate and 21 text figures)
THE DIFFERENCES BETWEEN THE LANGURS AND THE MACAQUES
Although the name Langur is, strictly speaking, the Hindi title for certain
Monkeys restricted to the northern parts of India, it has been extended during
recent years to include a great variety of related forms widely distributed in
the East. In this paper I have adopted it for all the eastern monkeys which
are not Macaques.
The Langurs are found. mostly in thickly forested districts, throughout
British India from Kashmir and Upper Burma in the north to Ceylon and
Tenasserim in the south. Outside that area they extend into southern China |
and Siam and through the Malay Peninsula to Borneo. There are several |
genera; but the only one which, as occurring within the British area, |
concerns us at the present time, is the genus now called Pzthecus, which was
formerly better known as Semnopithecus or Presbytts.
Except where regarded as sacred and encouraged to enter native villages,
Langurs avoid the presence of man and are essentially dwellers in the tree-tops,
feeding mainly upon leaves, shoots and buds and, in case of alarm, passing from
branch to branch with prodigious leaps or swinging from one branch to another
by means of their arms somewhat after the manner of Gibbons. ‘They are less
terrestrial and more arboreal than the Macaques and differ from them also in
diet. And the differences in these respects between these two groups of monkeys
are accompanied by important structural modifications connected with the
digestive organs and limbs. The Macaques have large cheek-pouches and
a comparatively small, simple stomach, whereas in the Langurs there are
no check-pouches and the stomach is capacious and sacculated. In the second
place the hands and feet of the Macaques are shorter and broader than in
the Langurs and have a larger thumb and great toe. (Text figs. 1A, iB, 2A, |
2B, 3A and 3B.) |
Fic. 1.—A. Simple stomach of the Common B. Sacculated stomach of Ceylonese Langur
Macaque (/JZacaca trus) from the Malay ( Pithecus senex vetulus).
Peninsula.
Half nat. size.-
Half nat. size
= — Wie al psy es
—— ~ : yyy
a
— ee \
K ek G (WY Qi, NESS
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a OS Ge ene ay WW
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B. Hand of Malabar Macaque (Macaca stlenus).
© Nas
si Gu Wr
Fic, 2.—A. Hand of Indian Langur (Pzthecus entellus).
Half nat. size.
Half nat. size,
Fic. 3.—A. Foot of Indian Langur {Pithecus entellus).
B. Foot of Malabar Macaque (Macaca silenus).
474. JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
These differences are expressed by referring the Macaques and Langurs to two
distinct families, the Macaques to the Cercopithecid@, which also includes
the Baboons, Mangabeys, and some other African Monkeys, and the Langurs to
the Colobid@, which also includes the Guerezas (Colobus) of Central Africa.
THE CLASSIFICATION OF THE LANGURS
For the original descriptions of the Langurs of British India we are indebted
to many authors, more particularly to Blyth; but the most important attempt
to deal with them comprehensively was made by Anderson, who not only had
in his hand Blyth’s specimens in the Calcutta Museum, but also made a point of
seeing, where possible, the speciinens described by French, Dutch and other
foreign zoologists and preserved in the Museums of various countries in Europe.
This work which is a monographic revision of all the species of Langurs, then
called Semnopithecus, known up to that time, was published in 1878 in the
Zoological Results of the two Expeditions to Western Yunnan. It was an
important treatise and formed the basis of Blanford’s account of the Langurs
contained in his classic volume of the Wammals of British India, published in
1891, and of Forbes’s account of these monkeys in vol. ii of his Handbook of
the Primates, 1894. But the arrangement and treatment of the species in the
three volumes cited does not attest any clear understanding of their mutual
affinities on the part of the authors; and this statement is still more true of
Elliot’s classification of the Langurs contained in vol. iii of his Review of the
Primates, published as recently as 1913.
On working through the collection in the British Museum, I find evidence
that the Langurs of the genus /Pzthecus may be divided primarily into three
groups sharply distinguishable by the colour of the newly-born young; and this
character is sometimes correlated with others in the skull, coloration or other
external features of the adult which make it possible to assign with some degree
of assurance to one or the other of the groups those species in which the
coloration of the young has not as yet been ascertained.
These groups may be defined as follows :—
Group 1.—Newly born young, a uniform dark blackish-brown all over.
(Plate I, fig. 3).
This group, which may be called the Auéellus-group, since the Entellus
Langur is the best known species contained in it, is restricted to India and
Ceylon and ranges in suitable localities throughout those countries from the
Himalayas southwards but does not, so far as is known, cross the Brahmaputra.
All the known Langurs of this group are discussed in this paper. I refer them
to two well-defined species, Pzthecus entellus and Pithecus senex, each of which
is represented by several local races or subspecies. The colour of the young
has not been recorded, so far as | am aware, in the typical form of either
of these; but it is known in the subspecies. dussumieri and achates, which are
admittedly closely related to typical entellus, and in the subspecies johnzz
which is admittedly closely related to typical senex.
Group II.—Newly born young, a uniform golden or orange red all over.
(Plate 1,sfic. 1):
This group, which may be called the Auratus-group after one of its earliest
known species, ranges from Assam, Upper Burma and Siam southwards through
the Malay States, Sumatra, Java and Borneo. So far as British India is
concerned it is represented by two distinct species, Pzthecus pileatus found in
Assam and other districts near at hand, and Pizthecus auratus which, as here
understood, extends from Upper Chindwin to Southern Tenasserim. Both these
species are represented within the British Indian area by several local races.
Group Il/.—Newly born young parti-coloured, white below, on the sides
of the body, the legs and the forepart of the head, but with a blackish or dark
brown spinal stripe of varying width extending from the back of the head
on to the tail and usually at all events spreading more or less on to the
outer side of the arms. (Plate I, fig. 2).
This group I call the Aygula-group, from the oldest known species belonging
to it. Containing several distinct species and subspecies in Borneo, Java,
Sumatra and the Malay Peninsula, this group is more southern in its distribution
than the 4uratus-group and only just enters our area in South Tenasserim
where it is represented by Pithecus femoralis, a species well known in the
Malay Peninsula.
Vi
|RN. BOMBAY NAT. HIST. SOC.
I. The Auratus-Group.
& SEABORNE, LTD.
LONDON,
NEWLY-BORN LANGURS
2. The Aygula-Group.
3. The Entellus-Group.
ad
=|
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 475
My conclusions regarding the numbers of different kinds of Langurs occurring
in British India and concerning the importance of the differences between
them vary considerably from those of my predecessors. It is needless here
to enter into the details of this subject since they are set forth in the following
pages ; but it must be explained that the divergence of opinion is due mainiy
to two things. In the first place, I use the word ‘species’ in a much
wider and more comprehensive sense than that adopted by those who
determined the langurs collected by the Mammal Survey, a great many of
the forms they considered as species being regarded by me as subspecies
or local races. In the second place, as compared with the pioneer workers,
Blyth, Anderson and Blanford, I have had the advantage of most of the
splendid material collected by the Mammal Survey, organized by the Bombay
Natural History Society. Where the authors mentioned had only one or
two, probably badly preserved skins and a few skulls to guide them, I
have had several, in some cases a dozen or more properly prepared, measured
and dated skins, from various localities, with skulls to match.
This material has revealed greater individual variation, especially perhaps in
the skulls, than was formerly supposed to exist; and it has established the
occurrence of a number of local races or subspecies linking forms to which,
in the absence, or rather ignorance, of interyrading types, full specific rank was
by common consent formerly accorded.
Although the material collected by the Survey has vastly extended our
knowledge of the Langurs of British India, it cannot be claimed that it has made
easier the task of naming them. It has, indeed, greatly increased the difficulty
of constructing analytical keys to facilitate their determination because it
has shown the impossibility of assigning definite and constant characters to
all the named representatives of this group. There is still much to be learnt
about them. Further collections in areas not yet investigated, like the eastern
parts of India from Orissa to south of Madras, will almost certainly bring
to light new local races and will perhaps show that some of the forms here
nominally distinguished are based upon characters not deserving that distinction.
For the most part I have had several specimens to guide me; but in others
I have been compelled to rely upon a single locally isolated example; and
in cases where I have given a name to one or two such examples, it has
been to draw especial attention to some particular character of interest,
although of doubtful systematic value.
SYSTEMATIC ACCOUNT OF THE SPECIES AND SUBSPECIES
The characters of the three groups above described into which the Langur
of British India fall may be expressed analytically as follows :—
(a2) Newly born young uniformly coloured above and
below ; never more than one frontal whorl in the
hair on the head ; nasal bones in the skull not
convexly bulging below the brow.
(6) Newly born young uniformly blackish brown.
(Himalayas to Ceylon) .. Lntellus-group.
(61) Newly born young uniformly golden yellow | or
golden red. (Assam to Tenasserim) .. Auratus-group
(a*) Newly born young particoloured, white with a
black or brown spinal stripe from the head to the
tail tip; a pair of frontal whorls; nasal bones
convexly bulging below the brow. (S. Tenasserim) d4ygula-group.
I, The ENTELLUS-group.
Characters as set forth above.
The Langurs of this group were referred by Blanford and other authors to six
or seven or even more species. But that method of treating them does not
express their true affinities. They are very obviously assignable to two categories
to which I give specific rank and their characters may be contrasted as follows:—
(2) Asingle whorl or parting on the forehead behind
the brow ; hair of the loins not shorter than else-
where on the back ; lower side of the body paler
than the upper; skull with prominent brows, the
plane of the orbit sloping downwards and back-
wards from its upper to its lower rim ... .. entellus,
476 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
(a*) No whorl on the forehead ; hair on the loins, or |
croup, shorter than on the back and typically
paler ; lower side of the body approximately the
same colour as the upper ; skull with less promi- |
nent brows, the plane of the orbit sloping down- |
wards and forwards from its upper to its lower |
TID. Her aoe ay aoe - SCHEX. |
1. Pithecus entellus, Dufresne.
Simia entellus, Dufresne Bull. Soc. Phil., Paris, i, p. 49, 1797. |
It is needless to repeat the distinguishing characters of this species as stated
above. Itis also needless to quote more than the original reference to the name,
because I use the specific title in amuch more comprehensive sense than my
predecessors to embrace the Langurs assigned by Blyth, Anderson, Blanford
and other authors to several distinct species, schzstaceus, entellus, hypoleucos,
anchises and priam, all of which, with the possible exception of azchises, I
regard merely as local races or subspecies of a single species for which estellus
appears to be the oldest name available. |
Distribution. North of the Ganges from Kashmir to Sikkim, possibly to |
Bhutan, and south of the Ganges from Cutch and Kathiawar to Bengal, thence
southwards into northern Ceylon.
KEY TO THE SUBSPECIES OF P. enfellus |
(a) Whiskers long and thick, mostly or completely hiding
the ears ; head and whiskers uniformly whitish or
pale buff and contrasted with tint of back. (N. of
Ganges).
(6) Coat long, shaggy and mane-like ; arms below
elbows black and strongly contrasted with
colour of body ont GIO.
(6+) Coat shorter, not shaggy and mane- -like ; arms
and hands not black and not sharply con-
trasted with colour of body.
(c) General colour dark earthy-brown ; coat
thick and woolly ... achilles.
(c1) General colour paler, slaty or greyish- -buff :
coat shorter and less woolly.
(2) Skull larger, with Pas nasal
region and larger teeth . hector.
(dq?) Skull smaller, with less. prominent
nasal region and smaller teeth we. SChistaceus.
(at) Whiskers shorter, not or only partly hiding ears ;
head and whiskers variable but never so white
and strongly contrasted as in section (a) (S. of
Ganges).
{e) Hands and feet black and generally sharply
contrasted with the arms and legs.
(f) Crown of head at most only a little paler
than nape and shoulders ; hands and feet
black or brown and typically sharply con-
trasted with arms and legs... entellus.
(f+) Crown of head clearly defined from the
shoulders and back by its paler tint.
(g) No tuft on crown; upper portion of
whiskers typically clouded with black.
(hk) Paler; crown of head uniformly
buffish in tint; tail and outside
and inside of legs below knees not
black. y
(z) Arms below elbows not black,
contrasted with the hands ... achates.
(71) Arms below the elbows black,
blending with tint of. hands.
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 477
(7) Paler ; cheeks pale like the
crown : dussumtiert.
(77) Darker ; upper part of
cheeks blackish .. 2ulus.
(A) Darker; crown of head suffused
with dark hairs ; tail and outside
and inside of leg below knee
black.
(k) Notso dark above or below ;
inner side of thigh clothed
with white hairs ; size smaller Ayvfoleucos.
(£*) Darker above and _ below ;
inner side of thigh mostly
clothed with black hairs;
size larger ies . C@neas.
(g*) -Agtiit On-the: crown ; upper portion
of whiskers pale like the crown ... elisa.
(e1) Hands and feet not black, practically uniform
in colour with the arms and legs. ;
(7) Crown of head pale, like the cheeks, and
contrasted with back ; notuft;sizesmaller priamellus.
(77) Crown of head typically darker than
cheeks, hardly contrasted with back,
and typically tufted ; size large .. pallipes:
In the construction of this key, my aim has been the juxtaposition of what
appear to be related forms, although it would perhaps have been easier to make
a frankly artificial analytical table by the rearrangement of the characters em-
ployed. But the actual or probable occurrence of intergrading specimens, it
must be remembered, makes the construction of such keys difficult and their use
somewhat unsatisfactory. Also it must be remembered that the distinctive
features of dussumierzt and hypoleucos, which I have not seen, have been inferred
from the descriptions of authors and that the form, also unknown to me, named
anchises by Blyth, which in the tint of its hands and feet is intermediate between
e and e?, has been omitted.
Subspecies schistaceus Hodgson.
Semnopithecus schistaceus, Hodgson, Journ. As. Soc., Bengal, ix, p. 1212,
Semnopithecus nipalensis, id. 161d.
Not Semnopithecus schistaceus of Blanford, Anderson and other authors, or at
least only in part.
Dark slaty above; below and entire head pale yellow; hands and feet
somewhat darkened ; concolorous. Hair of one sort, more or less wavy. Head
and body, 30 inches ; tail, 36 inches ; hind foot, 83 inches.
Distribution. Nepal; the Terai Forest.
The description of this race given aboveis taken from Hodgson’s original
diagnosis which, as indicated by the measurements, was based in all probability
upon an adult male. It is important to remember this because the description
cannot be applied to the adult males of any of the Himalayan Langurs in the
collection of the British Museum.
This series comprises several examples belonging to Hodgson’s collection
which are ticketed Nepal. No doubt Hodgson regarded them as specifically
identical with the form he described as schistaceus ; but by no stretch of the
imagination can they be described as ‘dark slaty above.’ They are most
emphatically brown, without any perceptible trace of dark, or even light, grey to
justify the epithet ‘ slaty.’ Moreover, the available evidence suggests that these
brown langurs, as stated below, live at high altitudes in Nepal and possibly
Sikkim. There is at all events no evidence that I can find of their occurrence at
the relatively low level of the Terai Forest ; and this was the opinion of Blanford,
who supposed the langurs of the Terai to be typical entellus and thus rejected
Hodgson’s original locality for schistaceus. :
The only specimen in the British Museum that was shot at a low level in Nepal-
is a female procured for the Mammal Survey by N. A. Baptista at Hazaria
Aya!
478 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
Pathergatti at an aititude of 300 feet. This has the clearly defined pale head,
the yellowish tinted under side and the uniform coloration of the upper side of
the body and outside of the limbs described by Hodgson for schistaceus and
Hinton and Fry? identified it as that form, probably quite correctly. It cannot,
it is true, be accurately described as ‘ dark slaty above,’ because the tint of the
upper side is pale and greyish-buff. The animal, however, isa female giving the
following approximate measurements in inches :--Head and Body, 23; Tail, 36;
Hind foot, 73; Ear, 13 ; and since the skull, although the teeth are fully up and
even worn, has the occipital suture open, the langur was probably not quite full-
sized and might wel! represent the female of schistaceus on the assumption that
Hodgson’s type of schistaceus was a male.
This skull is of particular interest because it is intermediate between the skulls
of typical extel/us of the low levels of Bengal and those of the large langurs
described below which inhabit the Himalayas at higher altitudes and have
always, but wrongly, been regarded as true schistaceus. Its measurements in
millimetres are as follows :—Total length, 115; lenyth of palate, 41; of upper
molars, 34; width across orbits, 64; across zygomata, 82. (Text fig. 4 B.)
It will thus be seen that it is noticeably smaller than the skulls of females of
the races from Chamba, Kumaon and Sikkim and has smaller teeth. On the
other hand, it hardly differs in these respects from the skulls of some examples of
entellus, although the face is more sloped and less vertical than in average
examples of that race owing to slightly greater prominence of the jaws, but the
suborbital area of the face does not recede more than it does in many examples
of entellus. The brow ridges are prominent and in the middle line rise vertical-
ly above the base of the nasals,
Pic. 4B.—Skull of Pithecus entellus schistaceus from
Hazaria Pathergatti in the Nepal Terai, 2 subadult.
Two-thirds nat. size.
Subspecies achilles, nov.
(Plated hos2.)
Semnopithecus schistaceus, Anderson, Blanford, Elliot and other authors, at
least in part.
Not S. schistaceus, Hodgson, 1841 (cf. swpra).
Distinguishable from schistacews by the nearly uniform chocolate or dusky
sepia brown colour of the upper side of the body, the tail and the outside of the
limbs. The upper surface, however, has a paler brown or buff sheen in
ee
* Jour., Bomb. Nat. Hist. Soc., vol, xxix, No. 2, p. 404, 1923,
PLATE 11.
JOURN. BomBay Nat. Hist. Soc.
THE Kanera Lanaur (P. entellus ajax)
THE NEPAL LANGUR
1;
lles)
entellus achi
P
(
2.
THE Common Laneur (P. entellus entellus)
3.
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THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 479
reflected light, the tip of the tail is sometimes pale and the hands and feet are
sometimes patched with black. Head and nape of neck and whiskers cream or
dirty white and contrasted with the brown hue of the shoulders. ‘The lower side
and the inner surfaces of the limbs whitish, sometimes tinged with buff.
Coat full, thick and long, but not markedly shagey, the hairs being tolerably
uniform in length and generally more or less wavy especially on the head.
Type. Adult male from the Satthar Hill (12,000 feet) in Gorkha, Nepal.
Distribution. Nepal at high altitudes ; and possibly Sikkim.
The only specimen certainly assignable to this race in the British Museura
which is labelled with full particulars is the type mentioned above, an adult
male shot for the Mammal Survey by N. A. Baptista at an altitude of 12,000 feet
on Satthar Hill, Gorkha, about 50 miles north-west of Khatmandu. Its
approximate measurements in inches are as follows :—
Head and Body © Tail Hind Foot Kar
30 364 | ge i
‘
These dimensions correspond tolerably closely with those recorded by
Hodgson for schistacews and with those of the males of the local race from
Chamba and Kangra described below.
The skull, too, although somewhat flatter along the top, is very like the skulls
of that subspecies, as the following dimensions in millimetres show :—
|
. f
Total length | Length of jLength of Upper
Palate Molars
; | Zygomatic
Orbital width | widen
In addition to the type, the British Museum possesses several skins of this
race belonging to Hodgson’s collection and ticketed Nepal. These, referred to
above under schistaceus, have always been regarded as schtstaceus. No doubt
they were considered to be identical with it by Hodgson himself and also by
Blanford who saw them. No doubt, too, they are the specimens, or some of the
Specimens, which induced Blanford to describe schtstaceus as ‘ earthy or
greyish-brown.’ Earthy-brown describes them well; but there is no grey in
the pelage of the upper side of the body. Blanford, however, like Anderson,
used the title schzstaceus in a comprehensive sense for all the big langurs living
at high altitudes in the Himalayas.
This race may also occur in Sikkim. There is at all events in the British
Museum a skin, alleged to have been collected in Sikkim by Mandelli, which is
in all important respects like the type and like Hodgson’s specimens. Moreover,
Elliot’s description of schistaceus which he said was ‘mars brown’ above, was
taken from a specimen in the Calcutta Museum ticketed Sikkim. And since
many of Hodgson’s specimens ticketed Nepal came, as Hinton has shuwn, from
Sikkim, Hodgson’s examples of achilles referred to above, may have come from
the latter country.
The available skull material of achilles is unfortunately scanty. Setting aside
the skull of the type, perhaps the most important is that of a young female
belonging to one of Hodgson’s specimens. A comparison between this skull
and that of the female described above as schzstaceus is interesting, It is much
younger as is shown not only by the distinctness of its sutures but more
particularly by the state of its dentition, the upper canine and last upper molar
only just appearing above the bone of the jaw, both being probably covered by
thegum when the monkey was alive. Nevertheless its measurements in
millimetres are as follows :—Total length, 112; length of palate, 40; of upper
molar series, 37; width across orbits, 69; across zygomata, 85. Thus it is
wider across the eyes and temples, has larger teeth andas longa palate, and
480 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
although a trifle shorter owing to the lesser development of the occiput, it would
certainly have exceeded the length of the skull of schistaceus with growth. The
profile too is very different and recalls that of the skull of the female of the race
from Kumaon, described below, in the prominence of the nose and the receding
brow.
Of Langurs from Kashinir there is very little material in the British Museum,
namely two skins only. These, however, appear to be identical with the Nepal
form achilles. This is perhaps surprising considering the difference between
achilles and the Langur of Chamba and Kanera described below.
I provisionally refer to achilles four skins from Sikkim which are a little
paler and greyer than skins from Nepal, but like them in the length and
texture of the winter coat. The dimensions of three out of these four female
specimens, collected in December and January for the Mammal Survey by
Mr. C. A. Crump, are as follows :—
| |
See Head & Hind | aS
Locality Body Tail foot | Kar | Weight
Chautang,5,350 ft, 2 fe. | 263°— | 33%—)| 87 —| 12” | ae
Lachen, 8,800 ft. 2 ea Solem 26h aye ee 35 Ibs.
Chautang, 5,356 ft.2 _... el BEE SO ees aS are ae
The only available adult skull is defective, one side of it having been shot
away. It isa little lower in its facial portion than the female skulls of hector
and ajax, but this may be an individual peculiarity because otherwise it is very
similar, except that the nasal region is less prominent that in hector and like
that of the Kangra examples of ajax. Its total length is 125 mm., and the
upper molar and premolar teeth measure 38 mm.
This is probably the Langur which has previously been recorded from
Sikkim as schistaceus. But, as stated above, there is in the British Museum a
skin labelled Sikkim which was collected by Mandelli and presented by
Blanford. But this skin is racially indistinguishable from Nepalese skins of
achilles ; and since, according to Blanford, Mandelli’s localities are not trust-
worthy, it is not improbable that the skin in question came from Nepal and
not from Sikkim. And the same may be said of the example of achilles in the
Caleutta Museum from Sikkim which Elliot described as schistaceus.
A distinguishable Himalayan race closely allied to P. entellus achilles has
been described and may here be mentioned, although it does not, so far as is
known at present, occur within the area of British India.
This is :—
Subspecies Zanzus, Elliot.
Presbytis lania, Elliot, Ann. Mag. Nat. Hist. (8), iv, p. 273, 1909; zd. Rev.
Primaiés, ii, p. 93, 1913 (under Pygathrix).
Colour very like that of typical achilles from Nepal but with aslightly greyer
sheen especially on the loins and hind legs, and with more white on the back
of the thighs. lie coat is a little longer, thicker, softer and more wavy.
Distribution. Chumbiin Tibet, 10,000 feet.
The dimensions in inches of the single fernale specimen collected by Major
F. M. Bailey are as follows :—
Head and body 25; hind foot 734; ear 1%.
The skull, which I have not seen, measures, according to Elliot, 125 mm.
in total length, and the zygomatic breadth is 102 mm.
Subspecies ajax, nov.
(Plate I], fig. 1.)
Pithecus entellus, Lindsay, Journ. Bomb. Nat. Hist. Soc., xxxi, p. 599, 1926
(not P. extellus, Dufresne.)
A big handsome langur as large as achilles but differing in the colour and
texture of its coat. The general colour of the upper side is dusky brownish-
grey, much greyer than in achilles, with the whitish cap on the crown typically
less distinctly defined owing to a greyish tinge on the nape blending more or
THE LANGURS, Ok LEAF MONKEYS, OF BRITISH INDIA 481
less with that of shoulders ; the arms below the elbows and the hands are black
or blackish brown in adults and the legs paler greyish-brown, darkened with
sooty grey on the thigh and knee in front, the feet at most a little darker than
the area below the knee, with the foot bordered externally with white and some
whitish hairs on the toes. The whiskers and throat, the under side of the body,
the inside of the liinbs and the backs of the thighs, and as much as four inches
of the tip of the tail are white.
The coat is loose and shaggy, forming a kind of mane over the shoulders and
upper part of the arms and on the flanks, and consisting largely of long greyish
white glistening hairs such as are seen in typical entellus.
Type. Adult male from Deolah, Chamba, 6,000 feet (H. W. Wells).
Distribution. Northern Punjab ; Chamba, from 6,000 to 7,500 feet ; Kangra,
from 2,450 to 9,500 feet ; Kulu, from 9,800 to 11,000 feet.
Young specimens differ considerably from adults in being much paler in
colour both on the back and onthe arms, and the coat consists of long thick
more woolly hair, the coarser glistening hairs of the coat of the adult being
practically absent. ‘The examples from Chamba were collected in December
and January by H. W. Wells. Those from Kangra, procured in March and
April, differ in having the pale cap on the head move clearly defined and in
having the arms browner. Three females from Kulu, collected in June, still
carry the winter coat but itis very shaggy and wavy. These specimens also
have paler arms. Such differences as those observed between the Chamba,
Kangra and Kulu individuals are probably seasonal.
The following are the approximate measurements in inches of some selected
individuals :—
| | |
Locality and Sex | “Head and Tail | aed Ear | Weight
‘ eRe oe 7
Chamba, Deolah, 6 000 ft., ¢ £3 |e BOs Bee ile LOS = 2 be
2 bara Tissa, 7; 500 fie Ce 30” B27 1h Se Zo 43 lbs,
we. ; ©halan Tissa; 6,700 it. cr 30” BB 0=—| FO 22 ia
Kangra Fort, 2 ASO it. ia 34a”) 847") 14” 46 lbs.
», » Chicaian, 9,000 ft., Q eo age Coa aon ca tan 28 lbs.
eee wamyala, Go00 ft, 2-201 23% SH esas. oi ielilas te i
Kangra, 2. 400 ft., ; a t Med 33” 73 14”
Kulu, Rahla 9,800 Et, One. ee nels 217 — 31 7 id”
ys ” ria me cae Ses ele 303” | 7 3
The skulls (Text fig. 5A.) do not differ in any important points from those of
achilles, as represented by the one from Gorkha and the fragments of those
collected by Hodgson referred to above. The following table shows their
dimensions in millimetres :—
— : |
Total | Length of | Uength of | Orbital | Zygomatic
length Palate | Upper Molars width width
ee ee an |
Locality and Sex
Chamba, ¢ 147 o/ 4] | 84 | ja0
ete = Ol 145 o4 40 | 82 111
pe es 141 34 40 84 109
Kangra, Jb 144 54 41 S4 109
3 2 124 46 40 72 94
rae g 120 43 40 13 94
ee ee
Subspecies A#ector, nov.
Presbytis schistaceus, Wroughton, Journ., Bom. Nat. Hist. Soe., xxiii,
p. 286, 1914 (not P. schistaceus, Hodgson).
Approximately the.same size as achilles and ajax but differing from both in
colour and in the shortness of the hairs of the winter coat. General
482 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
colour slaty-grey tinged with buff or pale brown ;_ the outside of the arms grey
below the elbow and a little darker than the back, but not nearly so dark as in
ajax; the outside of the legs paler than the back and with a sooty grey tint on
the thigh in front. The white cap on the head and nape sharply detined and
the under side either dirty white or more frequently suffused with buff or pale
red. The winter coat consisting of hairs of tolerably uniform length, shorter
than in achilles and not so thick and very much shorter than in ajax.
Type. Male (skin and skull) from Sitabani, Ramnagar, Kumaon, 2,000 feet.
B. M. No. 14.7.10.10.
Distribution. Kumaon up to 6,000 feet.
In general coloration the known examples of this race closely resemble
the specimen of schistaceus from Hazara Patherghatta described above. The
skulls of adult females of the two are, however, different in dimensions
and shape, those of hector being larger, with bigger teeth, less pronounced
brows and more prominent nose. But additional specimens from the Terai
Forest may prove hector to be identical with schistaceus, if the skulls are found
to intergrade.
The following table gives the approximate dimensions in inches of the skins
of this langur shot by C. A. Crump in November, January and February :—
Head and Tail | Hind
Locality and Sex Body
Kar Weight
!
Champawat, Almora, 6,000 ft., @ ... 274+ | 39 9+ | 2— git
Sitabani, Ramnagar 2,000 ft. ; a ee 26 3/ 84+), 2—) 38 lbs.
Dela, Ramnagar, 1,500 ft. ~ ae 24+ | 333—| 73+) 1$ | 39 1bs.
Ratighat, Naini Tal, 3,700 a 233-+ | 33 8 2—{ 31 lbs.
a TE ID PT LE SED
The two males, it will be seen, area little smaller than the adults from
Chamba ; but the skulls show them to be rather younger animals.
y ht My
UH VLA gy af
om
aA pa ns LY i
Hi me eT’ CJ
-- RE = Wy)
EZ2Z=8B eS 5
eae. 7
Fic. 5A.—Skull of Pithecus entellus ajax from Chamba, ¢ ad.
Two-thirds nat. size.
THE LANGURS, Ok LEAF MONKEYS, OF BRITISH INDIA 483
The skulls generally resemble those of achilles and ajax, but have the nose
more prominent. Thisis particularly noticeable in the skulls of two adult
females when compared with the skulls of adult females of ayax from Kangra.
It is also observable in two male skulls. These also are narrower than the
skulls of adult males of ajax ; but they are younger, the basioccipital suture
being still open. Their youth, no doubt, accounts for their narrowness; and
probably increase in the size of the brow ridges with advance of years would
lessen the appearance of prominence of the nose. (Text fig. 4A.)
A,
Fic. 4A.—Skull of Pithecus entellus hector from Naini Tal. 2 ad.
Two-thirds nat. size.
The dimensions in millimetres of four skulls are as follows :—
li
Total
Locality and Sex | : Length of Length of Orbital} Zygomatic
|
ength | Palate Upper Molars| width width
| e | | ff
Ramnagar ¢ || 135 | 50 3 ad 96
a Be ee ls 2130" 49 40 | 75 Y8
Naini Tal 9 a 123 46 40 70 94
Ramnagar @ | 119 46 41 | 70 90
Several skulls of adult males from Garhwal, presented to the British Museum
by B. B. Osmaston, may, for geographical reasons, be provisionally assigned to
this race; but unfortunately there are no skins from this district of Kumaon by
which the correctness of this inference can be tested. These skulls do not differ
appreciably from those of achilles and ajax.
The Langurs recorded by Blyth on Hutton’s authority as occurring in
Mussoorie and Simla up to 11,000 feet and described by Hutton as dark greyish
with pale hands and feet and white head, throat breast and tail-tip, certainly
differ from ajax from Chamba, but appear to agree very closely with this
Kumaon race, hector; and anumber of skulls of adult males presented by
Hutton to the British Museum, which may have been procured by him either at
Simla or Mussoorie, are not distinguishable from the skulls of adult males of
ajax and achilles. Nevertheless, as stated above, the skulls of adult females of
hector from Kumaon have, owing to the prominence of the nose, a different
appearance from adult female skulls of ajax,
484 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
Subspecies emtellus, Dufresne.
(Plate II, fig. 3.)
Simia entellus, Dufresne, Bull. Soc. Phil., Paris, 1, p. 49, 1797.
Semnopithecus or' Presbytis entellus of Blyth, Anderson, Blanford, etc.; and of
Dollman, Journ., Bom. Nat. Hist. Soc., xxii, No. 3, p. 438, 1913.
Smaller than the Himalayan races and distinguishable from them by the
uniform coloration of the head and shoulders and by the sharp contrast between
the black or brown and black hands and feet and the grey of the arms and legs
below the elbows and knee. General colour of upper side darker or lighter
shining sepia brown, becoming paler on the shoulders and head. Outer side of
arms to, or almost to, the wrists and of the legs down to, and often including,
part of the foot, much the same tint asthe back, as also is the greater part of
the tail ; but the leg below the knee is generally paler and greyer and the end
of the tail is usually nearly white. The whiskers, which do not conceal the ears,
the throat, the chest, the belly, the inside of the limbs and the back of the thighs
whitish or buffy- white.
Distribution. Northern parts of peninsular India south of the Ganges from
Bengal westwards to Kathiawar and Guzerat.
The Mamma! Survey secured a useful series of this langur from various
localities ranging from Bengal, whence the original example came, westwards
to Kathiawar and Guzerat. I have been unable definitely to establish the
existence of local races over the whole of this area which is probably tolerably
uniform in its physical features ; but the series of skins as a whole shows that
entellus is not nearly so sharply differentiated as is suggested by the writings
of Blanford and other authors. Blanford, for instance, following Blyth and
Anderson, lays great stress upon the uniform tint of the head and shoulders,
involving the absence of a distinct pale cap on the crown, and the sharp contrast
between the blackness of the hands and feet and the paler tint of the arms and
legs. Both these features are, however, variable. In aseries from Midnapore
(200 feet), for example, the head is paler than the shoulders, but the tints blend
on the nape; the hands and the feet are black in adult males but brown with
black fingers and toes in a young male and adult females. Similar variation
with age in the tints of the extremities is seen in other specimens. A female
from Junagadh in Kathiawar is darker above than the Midnapore examples but
has the feet grey with black toes and the hands and wrists black; and a female
from Deesa in Palanpur is paler than the Midnapore series but this langur was
shot in May when it was apparently still carrying its shagsy winter coat.
The following are the approximate dimensions in inches and_ the weights of
a few specimens picked haphazard :—
Locality ane Sex Rae Tail eee Ear | Weight
Hazaribagh, 1,000 ft., 4... ate 253 AP 8a e= 2
Midnapore, 200 ft., ¢ ae 25 44 S| 14-|. doebs.
Hoshangabad, 3,300 ft., ¢ es 25 42% | 8} | 2
Chikalda, Berar, 366 ft., ¢ ny 233 42 83
Junagadh, Kathiawar, 200 ft., ¢ ... 23 37 5, 73 13
Seonbadoh, Berar, 1,800 ft., 9 ey: Zo Lar Svene le ever 13
Midnapore, 200 ft., 9 ee ae Der 8 | 7 14 | 25 lbs.
Deesa, Palanpur, @ | | 7% 15
ae
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 485
These measurements bear out the statement made above regarding the
comparative dimensions of eztellus and the related langurs to the north of the
Ganges. In connection with Blanford’s statement that the ears of entellus are
larger than those of schistaceus, it must be remembered that he was unacquainted
with schistaceus in the restricted sense in which that term is used in this paper.
The ears on dried skins of ente/lus look larger mainly because they are not
concealed to the same extent by the whiskers. Nevertheless they are on the
average a little larger than the ears of the races from the high Himalayas.
The skulls vary greatly in shape. Blanford’s statement that the skull
differs from that of the mountain forms he took for schistaceus in having the
nose not projecting beyond a straight line tangential to the middle of the brows
and the alveolar border of the incisor teeth is not true. In two female skulls
from Hewra in Nimar and Seone, Malwain the Central Provinces, the nose
projects beyond that line. Similar variation in the shape of the cranial
portion may be seen by a comparison between the figure of an adult male
skull of ezte/lus published by Blanford where the upper surface is low and
flat, with a very long and projecting occipital region, and the skull of the
adult male from Hazaribagh which is more steeply sloped. Of particular which is
interest is the skull of an adult male from Deesa in Palanpur, which in its low
facial portion recalls the skull of the South Indian and Ceylonese race formerly
known as priam and described below as pallipes. Nevertheless, as Anderson
pointed out, the skulls of evtellus certainly differ on the whole from those of
the northern mountain races in having the line of the face more vertical, with
the nose and jaws less prominent. It also is alittle smaller and has smaller
teeth. But, as stated above, these differences will in all probability be found
to break down when more is known of true schistaceus from the Nepal Terai,
(hext fics. .5B;, 6A, 6B) 7A and 9A.)
SSS,
WSS
————
SS
=
SS
—
=
——!
—
FIG. 5B.—sxull of Pithecus entellus entellus from Hazaribagh, ¢ ad,
Two-thirds nat, size,
12
486 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. X.<XT1I
YZ
Fic. 6A,--Skull of ‘Pzthecus entellus entellus from Midnapore. 2 subadul¢
Two-thirds nat. size,
IND DZ.
sas
SMOSSEZ
Fic, 68.—Skull of Pzthecus entellus entellus from Palanpur. ¢ ad.
Two-thirds nat. size,
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 487
Fic, 7A.—Skull of Pzthecus entellus entellus from Hoshangabad, @ ad,
Two-thirds nat. size.
Fic 9A.—Skull of Pzthecus entellus entellus from Midnapore, ¢ subadult.
Two-thirds nat. size.
488 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
The following are the measurements in millimetres of some of the speci-
mens :—
ican 2 Total | Length of Length of | Orbital | Zygomatic
ocality sand ex length Palate Upper Molars | width width
Hazaribagh, @ ...| 131 51 | 34 82 106
a bare 128 47 36 78 102
Bengal, ¢ evel leo 46 SY 81 104
Deesa, 3 ae het a7 42 36 81 103
Midnapore, ¢...| +123 46 36 a 92
Chikalda, ¢ sncfoay 120 42 33 75 98
Junagadh,
Kathiawar, ¢ oy 0) 42 35 73 91
Midnapore, 2) o...| 2 130 36 34 67 89
Hoshangabad, 2...| 109 37 35 65 87
Junagadh, 2 rel LOG 37 33 72 88
With the exception of the second and third on the list, which were presented
to the British Museum by Mr. W. Theobald, these skulls were collected by the
Mammal Survey. The male examples from Midnapore and Junagadh are not
quite fully developed.
Biyth (Journ., As. Soc., Bengal, xiii, p. 470, 1844 and xvi, p. 733, 1847)
gave the name anchises to the skin of a Langur, from an tnknown locality in
the Deccan, which he described as like the darkest examples of entellus but
with the hands mixed white ard blackish, the feet whitish with dusky black
above the base of the toes and on the terminal phalanges and the leg from the
knee whitish. Coat generally much lcnger, the hair from 4 to 6 inches on the
sides of the body and not wavy as in entel/us ; the toes having long hairs like
those of aspaniel’s paw.
Out of the very large number of extellus and entellus-like langurs in the
British Museum procured by the Mammal Survey in India to the south of the
Ganges, there is not one that exactly fits the description of anchises. Nearest
to it comes a female, collected in November at Hewra (1,000 feet) in Nimar.
This resembles entel/us in the uniform tint of the head and shoulders; but
the coat, as in azchises, is long, there are longish hairs on the toes and the feet
are much paler, being grey, like theleg, with some black hairs on the knuckles.
The hands, however, are notso pale as in the type of anchises ; they are
distinctly brown, that tint blending with the colour of the arm, and have black
hair on the knuckles.
Very like this example from Nimar is one obtained by Mr. Prater at Lahi,
Seone Malwa, in the Central Provinces; but the hands and feet are a little
darker; and a second specimen from the same locality is still blacker on
its extremities and is hardly different from emfellus in that respect, but the
head is paler than the shoulders as in the more southern races below recorded.
These specimens, with the type of anchises, are peculiarly interesting be-
cause they show a complete gradation in the colours of the hands and feet
between entellus and the entellus-like form from Southern India and Ceylon
usually cited as przam or priamus which is described below under the name
pallipes. And the specimen with the darkest hands and feet and the paler
head similarly reveals a gradation between ezfellus proper and the race next
described.
The exact status of amchises must forthe present be left unsettled. It may
represent a distinct subspecies. At all events on the available evidence it can-
not be assigned to typical extellus; and equally certainly it cannot be
regarded as identical, as Blyth supposed, with the black-footed, black-handed
forms of southern India. Conceivably it may represent a variety or sport
of either.
Subspecies achates, nov.
Semnopithecus anchises var. Blyth, Journ. As. Soc., Bengal, xiii, p 471, 1844.
Presbytis entellus anchises, Dollman, Journ. Bom. Nat. Hist. Soc., xxii, No. 3,
p. 442, 1913. Not typical S. anchtises, (cf. supra.)
Semnopithecus hypoleucus, Blanford, at least in part. Not S. Aypoleucos, Blyth.
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 489
Closely related to typical enteldus and intergrading with it, but with the buff
or greyish-buff coloured crown of the head paler than the shoulders and differen-
tiated in tint, although not so sharply differentiated as in the races to the north
of the Ganges. Its liair also is more woolly. The general tint of the dorsal
surface is on the average soinewhat darker, the upper part of the cheek between
the eye and the ear is tinged with blackish-grey and the black of the hands and
feet is not so defined from the tint of the arms and legs owing to the somewhat
darker hue of the lower half of these limbs, especially of the legs below the
knees.
Distribution. Bellary, Dharwar and Kanara, from 1,500 tc 2,000 feet in
the Western Ghats; also Karwar, on the sea coast, to the west of the
Ghats.
Type. Adult {from Hausbavi, 2,000 feet, S.W. of Dharwar. B.M. No.
14.11.18.1.
The following are the measurements and weights of some of the specimens
collected by Mr. Shortridge for the Mammal Survey :—
: ind | |
Locality and Sex "Body | Tail | foot | Ear | Weight
Vijayanagar, Bellary, 1,600 ft., ¢ ...| 263”-—- | 40” ae 24% | 635. Tbs.
Fausbavi, o.. Dharwar, 2,000 ft, ¢... |; 27”%— 43”+" 8” | 2”— | 30 lbs.
Devikop, 26 miles 8. of Dharwar, |
2,000 10. t= Ses 2 2On OO ea a SOU
rate 26” | 38” 74” | Diem
Samasgi, Kanara boundary, | |
S. W. Dharwar, 2,000 ft., ¢ ...| 24” 34” | 7” . | ya” | 992 Ibs.
Biber raat ek 34”) 7” | 1g” | 193 ibs.
Vijayanagar, Bellary, 1,500ft., Q@ 1.) 26” 35” | 74"%—| 2” 27 lbs.
Alnavar, Dharwar, 2 oe cOK 345” | 73”°+| 2”7— om
Devikop, 26 miles S. of Dharwar |
ZOOO tte) a5 i eZ B7ne |) Fae) | -oH ae
Hausbavi, S. Dharwar, 2 pect 2agt |) Soe. || 7 127- |. 223 Ips,
Samasgi, Kanara boundary,
Ss. W. Dharwar; 2,000 ft., 2 2.) -228’—. | 314” | 64""- | 1k” | 17 Ibs;
POW
~
ec
Fic. 7B.—Skull of Pithecus entellus achates from Hausbavi. ¢ ad.
Two-thirds nat. size.
490 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Three examples collected at Potoli, N. Kanara (1,800 feet) are interesting.
Two of them resemble typical acha/es in colour ; but the third is darker, as dark
as the two examples from Jog described below, and is in this respect a transi-
tional specimen between the two races.
From Blyth’s original description of amchises it is clear that he knew by
hearsay from Eliot of the existence of exteliws-like langurs to the south of the
range of typical entel/us. Although he did not know the exact locality of his type
of anchises, he stated that that form represented entedlus in the Deccan and along
the foot of the Western Ghats. He also had a specimen from Coimbatore
which he regarded as a probable variety of ezchises from which it differed by
having black hands and feet as in entelius. ‘This langur no doubt belonged to
the race here described as achates. It was probably Blyth’s statement regarding
the distribution of auchises, and not his description of it, that induced Dollman
to identify as anchises the langurs here described asa new subspecies.
The skulls (Text fig. 7B) in form and also in individual variability are very
like those of typical entellus, but aresmaller, with a more vertical face, shorter
jaws and smaller teeth. The measurements in millimetres of a few of them
are as follows :—
SOTO OAD RR SO SN a TT ar
: Total Length of Length of | Orbital | Zygomatic
Be ay eax length Palate Upper Molars | widthi width
Hausbavi, ¢ oy ee 0 eal 41 32 74 93
Samasgi, ¢ seciee Ome 37 30 66 87
Potoli, N. Kanara, |
6) = AlZ | 40 34 70 oI
Devikop, 32 seat Vie 38 32 70 —
Vijayanagar, Q@.... 110 | 38 33 69 89
Alnavar, @ ae ALO} 4 39 SL 68 84
Samasgi, 2 gcc) LOZ) Yl 62 30 65 81
Karwar, & sae}, O00 34 31 61 82
These measurements show that whereas the skulls of the males are on the
average distinctly smaller than those of extellws, the skulls of the females are
quite as big. The sexual difference, therefore, in aczates is much less than in
entellus. Since moreover, the body measurements of the males attest approxi-
make equality in size between the males of achates and entellus, it is evident that
the males of achates have a relatively much smaller head.
Subspecies zuwlus, nov.
Resemnbling and intergrading with P. entellus achates, but smaller and with
closer, softer, less shaggy hair, and darker in tint, the back being deep brown,
against which the pale buff coloured head shows up conspicuously. Cheeks
with a considerable quantity of black hair between the eye and the ear, as in
achates. Arms in the male black from the elbow outside, a little paler in the
female. Legs deep greyish-brown outside. Tail blackish-brown with the termi-
nal third whitish. Lower surface and inside of arms and legs pale as in achaieés.
Distribution. Jog, Gersoppa Falls on the Kanara-Mysore Boundary, 1,300
feet. (G. C. Shortridge).
Type. 3h, BM. No. 14.11-18.7.
The two adult examples collected at this locality give the following approxti-
mate measurements in inches :—
Su oP Cae te Geis Ce oy cn hl geen | ane aon Panera
te Hand and) -, | Hind :
sex Body | Tail | Shae Kar | Weight
= | = ——
3 oP re se aZOE rim gt |Oe Hse sofa a 21 lbs.
Q ee ea see ee ete 50 4= O10 Disa 183 5
Except for the conspicuous paleness of the head, this langur much resembles
the race, Aypoleucos, in the darkness of the back, the black tint of the arms
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 491
and the proximal two-thirds of the tai] ; but in the paleness of the head, of the
tip of the tail, the grey hue of the outer side of the legs and the absence of
the blackness from the inner sides of the arms and legs it is like typical achates.
And the interest of the specimens lies in the link they form in the chain between
hypoleucos and entellus, (Text fig. 8A) hitherto regarded as well-defined species.
— Ges Y/);
WW Zr BaeBeg
= NZS
esa Y fos
o/s 1s
OAL
Fic. 8A.—Skull of Pithecus entellus iulus from Jog in Kanara. ¢ ad.
Two-thirds nat. size.
The skul!s measure. in millimetres, as follows :—
RE EEE 2 rare
{
ae Total | Length of Length of Orbital Zygomatic
length Palate Upper Molars | width | width
Me. eas is) 2 AO6 S15 30 65 84
ete. sae 103 36 3] 63 85
- Judging from the somewhat scanty material, there is scarcely any difference
in size between the male and female of this race.
Subspecies dussumtert, Geoftr.
Semnopithecus dussumiert, 1. Geoffroy, C. R. Acad. Sci., xv, p. 719, 1842,
and Arch, Mus., ii, p. 538, pl. xxx, 1843.
Back greyish-brown ; crown of head, cheeks, neck and flanks yellowish ;
outer surface of arm to the elbow a little darker than the back, from the elbow
to the finger-tips black ; outer surface of legs tothe ankle a little darker than
the back, feet black ; tail black turning to greyish: brown in the terminal third ;
under side and inside of limbs yellowish.
Distribution, Coast of Malabar.
Since the single known example of this langur was collected by a French-
man, it probably came from Mahé, the French Colony in Malabar.
No langurs quite resembling this race, as described and figured, were collected
by the Survey, those most like it being the specimens referred to achates and
tulus. The latter, indeed, so far as it is possible to judge, only differs from
dussumiert in being much darker on the back, which cannot be described as
greyish-brown, and in having the upper part of the cheek or whiskers between
the eye and ear washed with black pigment. Otherwise they appear to re-
semble one another in colour, particularly in the blackness of the arm below
the elbow. Close resemblance to dussumieri is also shown by achates, but in
492 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
the latter the ariui below the elbow is not black, being noticeably paler than the
hand, the upper part of the cheek is dark and the basa! two-thirds of the tail is
not black. It may be added that dussumieri differs from hypoleucos in being
much paler above and in having the area of the leg below the knee about the
same tint as the back and not black. Yet in spite of these obvious differences
between the description of dussumieri and the type of hypoleucos, which he had
in his hands and described, Anderson quoted dussumier? as an unquestionable
synonym of hypoleucos. Naturally his description of hkypoleucos does not fit
dussumiert. Blanford, on the other hand, gave a description of Aypoleucos
which fits dussumiert, but not the type of hypoleucos. Finally, Elliot’s state-
ment that the type of dussumuzert, which he saw in the Paris Museum, agrees
with his description of an apparently accurately determined specimen of
hy poleucos from Travancore must be untrue, if any reliance is to be placed on
Geoffroy’s description and figure of dussumieri.
Subspecies Aypoleucos, Blyth.
Semnopithecus hypoleucos, Blyth., Journ. As. Soc., Bengal, x, p. 839, 1841;
also xiii, p. 470, 1844; and xvi, p. 733, pl. xxvi, fig. 1, 1847 (Presbytis).
Anderson, Zool. Res. Yunnan, p. 20, 1878. Not S. hypoleucos, Blanford,
Mamm. Brit, India, p. 33, 1888, or at least only in part.
Back and shoulders deep dusky brown, head and whiskers brownish-white, a
little darker on crown ; outer surface of arms to elbows and of legs to knees the
same colour as the back but arms below the elbows and legs below the knees,
including the hands and feet, black; tail also black. Sides and lower surface
of body, inner side of arms to elbows and of legs to knees white; arms below
elbows and legs below knees mostly black with some whitish hairs on the arms
and on the front of the leg.
Distribution. Travancore.
No langurs exactly fitting this description, which is taken from Blyth’s
original diagnosis, were collected by the Mammal Survey. Blyth had one skin
of an old mate said to have come from Travancore and the dimensions he gave
of the head and body, 21 inches, and of the tail, 32 inches, are probably only
approximately correct. Nevertheless, Anderson, who extracted the skull from
Blyth’s type, stated that its total length is 106 mm., and since this type was an
old male, it is clear that Aypoleucos is a small race closely resembing in that
particular the race from Jog described as zwlus. It is also evidently very like
zulus in colour especially in the blackness of the outer side of the arm below the
elbow ; but it differs from zudws in having the legs below the knees black out-
side and inside, with apparently more black on the inside of the arm below the
elbow and a somewhat darker head.
Subspecies @neas, nov.
Evidently tolerably closely resembling hypoleucos but a larger and more
heavily pigmented form. Back dark brown ; the top of the head much darker
than in achates, iulus and priamellus and not nearly so sharply contrasted with
the shoulders but passing into reddish-buff in front. Arms very dark, deep
brown close up to the body but black in front on the shoulder and from above
the elbow tothe finger-tips on the inner as well as on the outer side. Outer
surface of legs like the arms, brown close to the hip but black in front and
from above the knee to the tips of the toes, black also on the inner side up to
the pubic region. Tail blackish throughout. Whiskers, throat, flanks and
lower surface buffy or reddish-white, this hue also extending along the inner
surface of the upper arm and obscurely down the middle of the pubic area
between the thighs,
Distribution. S. Coorg from Makut, below the Ghats, 250 feet, to Wottekolli,
2,000 feet (G. C. Shortridge).
Type. Adult male from Makut, B.M. No. 14.11.18.23.
The description given above was taken from a sub-adult male from Makut
and an adult female from Wottekolli. They resemble /ypoleucos in general
coloration but certainly differ from it in the blackness of the hairs on the inner
side of the thigh making a sharp contrast with the white of the belly. A young
male with the head and body 24 inches and the tail 32 inches is not so heavily
pigmented on the limbs and in this respect comes nearer “ypoleucos. ‘The head
of @neas also seems to be considerably darker than in hypoleucos ; and the only
ae
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 493
known example of the latter, au old male, is considerably smaller than the adult
male of @zeas as the following measurements show :—
Head and
. | Hind | ;
Locality and Sex Body | Tail | foot | Ear | Weight
|
tas vee e eo a es ha a ee S| ge PP ee ) |
Makut, 250 ft., ¢ ... a | 28” S27 he 13” | 253 1bs.
Wottekolli, 2,000 ft.,2 _... ia 250 Soe | 6” | qa” | 22 Se
The skulls of these two adult examples measure, in millimetres, as
follows :--
Bay Total Length of Length of | Orbital Zyyzomatic
f length Palate Upper Molars width | width
re 112 40 a2 68
2 106 38 RY? 66 86
It seems singular that two races of Pithecus entellus should occur in Coorg,
namely, e/zssa at Nagarhole in the south-eastern part of the province and
céneas at Makut and Wottekolli in the southern part, and both at nearly the
same elevation. Nevertheless the two are readily distinguishable by their
colour and it seems probable that neas inhabits a zone with much heavier
rainfall than edissa.
Subspecies e/zssa, nov.
Presbytis entellus anchises. Dollman, Journ. Bomb. Nat, Hist. Soc., xxii,
p. 494, 1913. Not auchises, Blyth.
Indistinguishable in general colour from some examples assigned to achates
and no two of the four specimens examined exactly alike. But differing
apparently from achates in having somewhat softer, silkier hair, a distinct tuft
or crest on the crown of the head aud in the uniformly white or whitish-tint of
the cheeks, the dusky hue between the eye and the ear, present in achates and
zulus, being absent.
Distribution. Nagarhole, S. E. Coorg (2,600 feet).
Type. Adult 2, B. M. No. 14,11.18.16.
The chief interest of this form lies in the presence of the crest or tuft on the
crown of the head, which was noticed by Mr. Shortridge, the collector, and is
apparent on all the skins. A similar crest has hitherto been regarded, so far as
the Indian and Ceylonese langurs are concerned, as distinctive of the race
described below as fallifes and generally known as priam or priamus. I do
not attach very much importance to this crest as distinguishing e/zssa from
achates ; but its development in e/7ssa unquestionably lessens the systematic
value that has been attached to the development of a similar crest in pallipes
and it is worthy of special attention on that account alone.
The measurements in inches and the weight of two adult females and of
an immature male are as follows :—
Sex | cece | Tail Hiud foot ! Ear Weight
raf DAY | 36” | 74” 13” 20 lbs.
Oe Dae ear |) BG Me 14” 23 Ibs.
ees" Be i give ||, gust 14” 184 lbs.
494 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
The following are the dimensions of the skulls of the same specimens :—
Sex Total jLength of} Length of Orbital Zygomatic
length Palate Upper Molars | width width
Pee Nees eee eS | ey | 63
QO. | - 103. 36am 33 | 67 79
Q So | 34 30 65 81
In the skull of the male the canine and the last molar, although through
the bone, are only half up; and since it may be assumed that the monkey
was not full sized, the conclusion that e/zssa is about equal to achates in size
is justifiable.
Subspecies priamellus, nov.
Colour of upper side sepia brown, similar to that of achates, but with the head
and cheeks dirty white, without any dark pigment on the cheeks between the
eye and ear, with the outer side of the arms and Jegs uniformly brown, about
the same tint as the back, this colour blending with that of the hands which are
not black and only slightly darker than the arm, while the feet are greyish-
brown and the same colour as the leg below the knee, the tail being the same
colour as the back with its terminal portion whitish. The back of the thighs,
the whole of the underside of the body and the inside of the limbs clothed with
white hairs.
Distribution. Shernelly in Cochin.
Type. Adult male, collected for the Survey by Mr. Riley O’Brien. B.M.
Now 21 21155:7;
One specimen only was obtained and no measurements were recorded.
The chief interest of this specimen lies in the resemblance it shows to achates
and pallipes. It approaches achates in the paleness of the head as compared
with the back but differs in the absence of blackness from the hands and feet,
and it resembles pal/ipes in the hands and feet being the same tint as the arm,
and legs but differs in the colouring of the head. ‘There is also no trace of a
crest on the head.
The skull has prominent brows and the interorbital nasal region is concave,
being vertical above and projecting below at an angle of about 135°; but the
most noticeable peculiarity about the skull is the shape of the nasal orifice which
instead of being long and narrow, as is typically the casein the other races of
entellus, is wide, the width considerably exceeding half the greatest height.
(Text figs. 8B and 9B.)
The dimensions of the skull are as follows :—
| |
|
Total Length of
Length of la Orbital Zyzomatic
length Palate | Upper Molars | width | width
7 | | |
105 | 38 Slee 69 52
Since the skull! attests the full maturity of the specimen, it is clear that this
race of langur is much smaller than pallipes, considerably smaller than achates
and as large only as zu/us ; but from the structure of the skull and the colour of
the hands and feet I am disposed to think this langur is more nearly related to
pallipes than to achates or tulus.
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 495
Fic. 8B.—Skull of Pzthecus entellus priametlus from Cochin. ¢ ad.
Two-thirds nat. size.
Ks
(j th aN Wh
Weis /
’
.
by
\
“shy
ugonne’l My} {}
Feat; i) 4
| ) 2 Nl y
|| +i, in 4 f
UTATTRDM |
| |
Fic. 9B.—Skull of Pithecus entellus briamellus from Cochin, -¢ ad.
Two-thirds nat. size. :
Subspecies pallipes, Blyth.
Semnopithecus pallipes, Blyth, dun. Mag. Nat. Aist., p. 312, 1844 (April).
Semnopithecus priam, Blyth, Journ. As. Soc., Bengal, xiii, pp. 470 and 476,
1844 (after October).
Presbytis priamus, Blyth, Journ. As. Soc., Bengal, xvi, p. 732, pl. liv,
fig. 1, 1847.
Semnopithecus priamus, Anderson, Blanford and most recent authors.
Presbytis thersites, Blyth, Journ. As. Soc., Bengal, xvi, p. 1271, pl. liv,
fig. 3, 1847.
Tolerably closely resembling typical ewte//us in size and general colour, the
back and outside of the limbs varying from uniform sepia brown to slaty-grey,
the crown of the head being brown and hardly contrasted with the tint of the
shoulders, although frequently contrasted with the whiter hue of the cheeks and
496 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
brow ; but the hands and the feet are practically the same colour as the arms
and the legs and not blackened as in emtellus. Also the hair of the crown is
typically raised into a compressed crest or tuft. Thetail is tke the back in
colour except its extreme tip which is grey or buffand the whole of the under
side, the inside of the limbs and the backs of the thighs are white or whitish.
Distribution. Southern India, the Coromandel Coast from Nellore south-
wards, to, according to Blanford, Trivandrum and up the Nilgiri Hills toa
height of 6.000 feet, as reported by Davison. Also Northern and Central Ceylon
as far south as Trincomali and the Kandyan Hills.
In regarding this langur as a distinct species from enfe//us, Blanford relied, so
far as external characters are concerned, upon the pale hands and feet and the
presence of the crest on the head. But the importance of these characters is
lessened by the presence of pale hands and feet in frzamellus and in the form
named azchises by Blyth, the presence ofa similar crest in e/zssa from $.E.
Coorg, and its absence in the Ceylonese examples of pallipes nanied thersiles by
Blyth.
The following table shows the approximate dimensions in inches and the
weights of some of the specimens obtained by the Mammal Survey :—
[Head and |
| |
Locality and Sex Body | Tail |Hind et Ear Weight
Cheddikulum, Ceylon, ¢} 25” 36” 74” 2” | 293 Ibs.
rs » d| - 232" 32” 75” 2”— | 252 Ibs.
Mahendragiri Range, S. | |
Travancore, ¢ 4 pais | 26% 74” Be | ae
Mankeni, Ceylon, 9 ... 20m | 3438” 67 14” 15> ibs:
. .. Oo a 2 30” 61” 1k” 13 Ibs.
The Ceylonese examples were collected by Major E. W. Mayor in the
northern and central provinces of the island, and the single example from
S. Travancore by Mr. R.S. Pillay at Avambo at the foot of the Mahendragiri
range.
These measurements show that the adult males are about the same size as
entellus and achaies, and suggest that the females are smaller; but the two
females measured are shown by their skulls to be scarcely full grown. Neverthe-
less, the skull of an adult female, recorded below, from the Nilgiri Hills, is
considerably smaller thau that of a male from the same locality, thus showing
that in pallipes the sexual! differences in size are as marked as in the skulls of
typical evtellus.
The measurements in millimetres of the available skulls are as follows :—
Loony and Sex| otal | Leng of | Lesa of] Oral Zygor
a ee eee ire ! :
Mahendragiri |
Range, ¢ Rit a 23 43 32 fee: 95
Nilgir? Hills, ¢ ...) 122 45 34 78 97
Cheddikulum, aed 117 42 31 74 90
Nilgiri Hills, 2 .../ 100 36 : 30 | 65 82
Mankeni,@ ...|_—-:100 32 30 64 75
oe. POP Sy OO Be 30 63 78
yn Sly 4 Waals ee 34 30 60 77
{
LR LT TE SE EE FT I STS TT I
THE LANGURS, OR LEAF MONKEYS, OF BRITISH [NDIA 497
- But the measurements convey no idea of the profound variation in the shape
of the skulls. The skull of the adult male, collected by Davison in the Nilgiri
Hills, which is certified by Thomas and Blanford to have been taken from the
head-skin of aspecimen of pallipfes (prianz), has the nose prominent, the back
of the cranium high, the plane of the occiput steeply sloped and the lower jaw
very massive, with the symphysis strongly inclined. But in the skull of the
male from the Mahendragiri Range the nose is almost vertical, the back of the
cranium is much less raised, the occipital! plane is less steeply sloped, the lower
jaw is less massive and its symphysis is more vertical. (Text figs. 10A
and 10b.)
Fic. 10A.—Pithecus entellus pallipes from the Nilgiri Hills. ¢ ad.
Two-thirds nat. size.
Fic. 10B.—Pithecus entellus pallipes from the Mahendragiri Range. ad.
Two-thirds nat. size.
498 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXI1
The female skull from the Nilgiri Hills (Davison) has the nose nearly vertical,
the cranial portion short high and rounded and the lower jaw massive, where-
as the female skulls from Mankeni in Ceylon have the facial portion less steep,
the cranial portion lower and flatter, especially in one of the specimens and
the lower jaw much less massive.
The skulls of adult males of pallzpes differ from those of typical etellws in
having the face lower and the upper jaw less massive; but they resemble in
that respect, as stated above, the skull of an adult male of entellus from Deesa
in Palanpur. This last-mentioned skull, indeed, is very like the skull of
pallipes from the Mahendragiri Range except for the prominence of the nose,
and in that respect it closely resembles the skull from the Nilgiri Hills.
The names pallipes and priam were given in the same year to this langur and
Anderson, Forbes and Elliot, assuming apparently that przam was the first to
be published, added padlipes to it asa synonym. But since pallipes was pub-
lished in April and przam not until after October, it is clear that pallipes is the
older name and by the law of priority must, if properly introduced, be adopted.
Blanford omitted to quote it, possibly regarding the name as invalid. But
Blyth applied ¢allifes to the ‘reputed’ Entellus monkey of Southern India
which is quite different from the true extellus. Technically perhaps it may be
claimed that this is not a description ; but since Blyth had a skin and knew it
came from Southern India, it is clear that by reputed he meant ‘ so-called ’
and not ‘ surmised’ ; and what he said about this langur was quite sufficient
to enable it to be identified with certainty. ‘This apparently was the opinion of
Anderson who quotes pallipfes in the synonymy of priamus without any query as
to the validity of the description or doubt as to the identity of the species. His
verdict, being that of the first reviser, I have adopted.
The only other name to be considered is ¢hersifes given by Blyth toa langur
from Nuera-Kalawa, west of Trincomalee in Ceylon, differing from fallipes
(przamus) in having no crest on the crown. Blanford interrogatively added
thersites to the synonymy ot the Ceylonese langur cephaloplerus, now known as
veltulus ; but Blyth was much too good a zoologist and knew the Indian langurs
too well to have overlooked the very obvious differences between the cepha-
lopterus and the entellus group of langurs; and it is quite clear from his figure
and description of ¢tersztes that it belongs, as he said, to the latter category.
It is most emphatically an exzellus-like, not a cephalopterus-like monkey, and I
have no doubt that it is identical with pallipes (priam), despite the absence of
the crest. Kelaart also obtained a specimen in Ceylon without a crest.
Two langurs alleged to be representatives of these so-called species are figured
side by side on Wolf’s plate of Ceylon Monkeys in Tennent’s Natural History
of Ceylon, 1861. Wolf was a first-rate animal artist and his illustrations on
this plate are so good that it may be inferred that they were drawn either from
living specimens or skins. Now itis interesting to note that the top left-hund
figure representing ¢hersites, alleged to have no crest, has a crest as large as
that in the Survey specimens, whereas the top right-hand figure, representing
priamus, has a high peaked crest larger than in any example of this langur I
have seen, although there is a good deal of individual variation in the size of
the crest. I think it probable that the crest is actually or potentially present
in all individuals of the race and that its occasional absence is due to
moulting.
Pithecus senex, Erxleben.
Cercopithecus senex, Erxleben, Syst. Regn. Anim., p. 24, 1777.
Cercopithecus vetulus, Erxleben, fom. cit. p. 25 (in part).
Cercopithecus kephalopterus, Zimmermann, 1780=—Presbytis, Semnopithecus.
Pygathrix or Pithecus cephalopterus, Anderson, Blanford, Elliot, Wroughton
and others (in part).
Pithecus vetulus, Hinton, Ann. Mag. Nat. Hist. (9), xi, p. 506, 1923
(in part).
Resembling, except in albino specimens, the Langurs of the Entellus-group
in the deep blackish-brown hue of the newly born young, but differing in the
absence of the frontal whorl-like parting on the forehead, the presence of a
patch of shorter paler hair upon the croup or loins, the black or brown tint of
the under side and, usually at all events, in the presence in the female of a
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 499
white or whitish patch of varying extent on the pubic region below the
callosities, and on the inner side of the thighs.
Distribution. Southern India and Ceylon.
The closeness of the affinity between the Langurs described below as
subspecies of Pithecus senex was well-known to Blyth, Anderson and other
authors. Hence the grouping of them under one specific name is not a novel
idea, although most modern authors have followed Blanford in admitting three
species.
Phe skulls, so far as my observations go, present a considerable amount of
individual and but little subspecific variation. The sketch of the side view of
the skull of P. senex nestor will give an idea of the general form. It will be
noticed that the brows are not nearly so pronounced as in the subspecies of
Pithecus entellus, so that the plane of the orbit is not vertical but inclined back-
wards. But the prominence of the brow and consequently the slope of the nose
and the inclination of the orbit vary a good dealindividually. (Text fig. 11A).
Fic. 11A.—Skull of Pithecus senex nestor from Panadura. ¢ ad.
Two-thirds nat. size.
The skulls may be here dealt with comprehensively. There will be no
occasion to do more than refer to them very briefly under the subspecific
headings.
The following are the measurements in millimetres of a few :—
Total
j |
: Q Length of Length of Orbital | Zygomatic
ecu and Sex length Palate Upper Molars; width | width
johnii, S$ (juv.), | |
Cochin im 108 32 30 65 | 80)
johnii, 2, Nilgiris. 108 36 on 64 83
johnii, 2, Cochin. 95 30 30 oe aN.
vetulus, &, Anasi- |
galla 1 103 34 26 61 74
nestor, 3. Pana- |
dura sea 98 33 26 60s 19
nestor, 2, Pana-
dura see 94° 32 26 607 4) 72
These measurements suggest that the South Indian subspecies, ju#miz, has a
considerably larger skull than the Ceylonese races vetudus and nestor ; but I
500 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
have seen a male skull belonging to a form from Ceylon clesely allied to vetulus
and nestor which gave the following measurements :—Total length 109, length
of palate 39, of upper molars 28, orbital width 65, zygomatic width 80.
KEY TO THE SUBSPECIES OF Pithecus senex.
a General colour white or whitish are gee AESCHEZG,
a* General colour brown or black.
6 Crown and whiskers tolerably uniformly coloured
buff or brown; the throat darker than the
crown ; general colour black; the tail also
mainly black . johnit.
6+ Crown and whiskers contrasted, crown brownish
or buff, whiskers white or whitish; throat
lighter than crown ; tail greyish to brownish,
not black.
¢ Coat thick and shaggy; head and croup not
sharply distinguished by colour from the
back ae monticola.
ce! Coat shorter and smoother : head and croup
more distinctly defined from colour of
back in adult.
dad General colour black, with head darker
but croup patch silvery grey vetulus.
d* General colour brown, with head paler
and croup patch grey to grey brown westor.
Subspecies sevex, Erxleben.
Cercopithecus senex, Erxleben, Syst. Regn. Anim. Mamm. p. 24, 1777.
Seninopithecus senex. Schlegel, Mus. Pays Bas, Simie, p. 53, 1876; Blanford,
Faun. Brit. Ind. Mamm., p. 35, 1888.
Presbyits albinus, Kelaart, Prodrom. Faun. Zeyl, p.7, 1852: 72d, Journ. As. Soc.,
Bengal, xx, p.A82,\852:;
Fur dense, wavy, of a nearly uniform white colour, with only a slight dash
of grey on the head; whiskers white, full and pointed laterally ; hands and
feet with long white hairs overhanging the tips of the finzers and toes. Face
and ears black, eyes brown ; palms and soles flesh-coloured.
Distribution. Ceylon, Matelle (Matale).
Erxleben gave the name sevex to a white Ceylonese monkey referred to by
Knox! as another sort of monkey, milk white both in body and face. This
no doubt was a pure albino possibly, perhaps probably, belonging to one of
the Ceylon races of Langurs recorded below; and since senex has page
priority over vefulus, the former name must, according to the rule, be
adopted, although it was unfortunately probably applied to a variety.
The available evidence does not at present enable us to say to which, if to
any, of the now well-known races sexex should be applied. In adopting
the name I have, therefore, followed Schlegel who seems to have been the
first reviser to affix senex toa definite type of Langur from Ceylon. ‘This
was represented by a specimen brought by Temminck from that island
and presented to the Leyden Museum. Schlegel described it as yellowish-
white, washed with brown on the back and with red on the upper
side of the head and nape. Blanford, who saw what was no doubt the same
specimen in Leyden, endorsed, with unimportant modifications, Schlegel’s
description of the colour, adding that the face and ears were black, the palms
and soles flesh-coloured, the fur dense and wavy, the whiskers full and the
toes well haired. He compared its dimensions to those of P. ursinus (=montt-
cola) and the thickness of the coat bears out the view that the two are similar.
Although the blackness of the face and ears and the wash of colour on the head
and back show that this specimen is not so complete an albino as the white
*Flistorical Relation of Ceylon, vol. i, Ch. VI, p. 25, 1681.
Mitt LANGORS, OR LEAP MONKEYS, OF BRITISH INDIA 501
langur recorded by Knox, Schlegel’s application of the name sevex need not
on that account be set aside.
But many years before Schlegel’s time, Kelaart gave the name albinus to a
whitish Langur received as a living animal from Matelle, and his description
applies so closely ina general way to Temminck’s specimen, although the
latter was paler, that there seems no reason to doubt that Temminck and
Kelaart had very similar varieties of the same Langur. I have, therefore, fol-
lowed Schlegel in adding aléznus to the synonymy of senex and have reproduced
Kelaart’s description and fixed upon Matelle as the typical locality for senex in
view of the possibility of the existence now, or in the part, of a very pale race
akin to the darker races, vetulus, nestor and monticola. This possibility must
be borne in mind because it is in a measure supported by an old record by
Spence Hardy, quoted by Sir J. Emerson Tennent, to the effect that a troop of
white monkeys was seen at Dambool on the rock upon which the great temple
stands and by Blanford’s statement that white monkeys might be seen occa-
sionally on the Ceylonese mountains in parties of three or four, always apart
from other monkeys.
On the other hand, Sir Emerson Tennent had a white langur, taken between
Ambepasse and Kornegalle, where they were said to be numerous; and this
specimen, according to him, had all the characters of cephulopterous, apart from
its colour, which was strikingly white with the face and eyes black. He was
told, moreover, that white monkeys had been seen near the Ridi-galle Wihara
in Seven Korles and also at Tangalle. This suggests that the lowland form,
probably vetu/us, is addicted to albinism.
Subspecies vetulus, Erxleben.
Cercopithecus vetulus, Erxleben, Syst. Regn. Anim. Mamm. 3, p. 25, 1777.
(= cephalopterus of later authors).
Pithecus vetulus vetulus, Hinton, Ann. Mag. Nat. Hist. (9), xi, p. 509.
1923.
Colour of back from nape to croup black finely speckled with grey; croup
grey or silvery, this tint spreading on to outside of thighs and root of tail; tail
greyish to greyish-brown ; arms and legs at least from the knees downwards
black like the back ; head brown above ; whiskers and throat mostly white
but whiskers brown at the ends and some brown on the middle of the throat ;
under side and inside of limbs black, but some white on the pubic region
beneath the callosities in the female.
Distribution. S. Ceylon from about Ranna through the Udugama Hills in
the Galla District and up the western coast to the Kala Ganga River ; probably
also in Ratnapura.,
This information regarding the range of this race was supplied by Mr.
W. W.A. Phillips to Mr. Hinton, who quoted it in his paper.
The following table gives the approximate dimensions in inches of some
examples of this race :—
|
Locality and Sex Bea Tail Hind foot, Weight
Kalutara, ¢ | 24 — 28” — 63” —
mnasigalla; cf .... ..| 2157+ 29” +- 64” — 123 lbs.
Matugama, ¢_... eh ee th BOR = 7” — |
Kalutara, © Bes es) fae | 2747+ 6c
Kalutara, Q ee molweiege | 280 6” — | lit tbs
Anasigalla, 9... stl ee | 30” | G7
502 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XAX2G7
Subspecies wzestor, Bennett.
Semnopithecus nestor, Bennett, Proc. Zool. Soc., p. 67, 1833.
Semnopithecus kelaarti, Schlegel, Monogr. des Singes, p. 52, 1876.
Pithecus vetulus nestor, Hinton, Ann. Mag. Nat. Aitst. (9), xi, p. 510, 1923.
Pithecus vetulus phillipst, Hinton, zdzd.
Differs from P. senex vetulus, in having the blackish hue of the upper and
under sides and limbs replaced by deeper or lighter sepia brown which, toning
more with the paler areas on the head and croup, lessens their conspicuousness.
The whiskers, however, are white and conspicuous, the head is, ochraceous or
buff, and the patch on the croup varies from silvery-grey to pale brown.
Distribution. Southern Ceylon ; the Panadura and Horana Districts, north of
the Kala Ganga River, and ranging up the west coast to a little way north of
Colombo and inland up the Kelani Valley, probably to the foot of the Adam’s
Peak Range.
This information regarding the range of this race was published by Mr. Hinton
from information received from Mr. W. W. A. Phillips ; and Mr. Hinton has
shown that it inhabits an area where the annual rainfall is less than in the area
inhabited by vetulus.
The type of zesfor, in the British Museum, is a half-grown langur, merely
ticketed Ceylon. I cannot, however, distinguish it by any reliable characters
from the series of skins, sent by Mr. Phillips, which were described by Mr.
Hinton as Pithecus vetulus phillipst.
The type of pxzllipsz, an adult male from Gonopola in the Panadura District,
has the back, the arms and the legs sepia brown, with the limbs darkening to-
wards the extremities and turning black on the hands and feet ; the croup patch
and tail are grey, the head buffy brown and paler than the back, the whiskers
and throat white and the lower side dark brown.
An adult female from Horana in the Panadura District is tolerably similar and
has a white patch below the callosities.
Anotiier adult female from Godigamiura in the Panadura District is darker
brown both above and below than the type and has the croup patch, not grey
and conspicuous, but pale brown as in the type of wesfor.
The approximate measurements in inches of three specimens from the Pana-
dura district are as follows :—
| Head and
, ; Hela Se
Locality and Sex | Body -| Tail Hind a Ear
Gonopola, 2 , in 20” 253” 64” 13”
Gonabendaluwa, Q ae 223” 31% 67 | 13”
Horanna, @ ae Po laetorg coo sis | ia)
ETI me — a (RT CRE LT ET
Subspecies monticola, Kelaart.
Presbytis cephalopterus var. monticola, Kelaart, Journ. Rov. As. Soe.
(Ceylon Branch), ii, No. 5, p. 321, 1850, zd. Prodr. Fauna Zeylan, p. 2, 1852.
Presbytis uwrsinus, Blyth, Journ. As. Soc., : Bengal, -xx, p. 155, 1851; and
of Anderson, Blantord, Elliot, Wroughton and other authors quoted under
Semnopithecus, Pygathrix or Pithecus.
A large mountain-form closely resembling P. vefulus nestor in colour but
with shaggier longer coat, and the pale patch on the croup usually at all
events less distinctly defined. Colour varying from deep blackish-brown to
sepia brown on the back and limbs. Summit of head and nape always
brown ; whiskers, throat ard chin grey or white and contrasted with the colour
of the head and chest. Face black, bordered with black hairs.and with a line
of black hairs between the eyes and expanding on the summit of the nose
between the nostrils.
Distribution. Mountains of Southern Ceylon.
Examples of this fine langur are rare in collections. ‘The British Museum
has four examples only. An adult male from Adam’s Peak (S. B. Bell) is
dark blackish-brown, blackening on the hands and feet. The croup patch
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA _ 503
is paler, sepia brown ; the whole of the crown of the head and the nape are
also paler sepia brown and, like the croup-patch, are sharply set off by the
darker hue of the back; the whiskers, throat and chin are grey ; the tail, which
in its basal half is like the croup patch, becomes greyer in its terminal portion.
Another adult male from Pattipola in the Central Provinces, where it was
shot at an altitude of 6,200 feet by Major E. W. Mayor, is considerably paler
brown than the one from Adam’s Peak. Hence the croup patch and the crown
of the head are much less conspicuous and the hands and feet are brown.
An adult female from Hakgalla Uva, 5,600 feet (A/ajur E&. W. Mayor) is
also paler than the male from Adam’s Peak but somewhat darker than the one
from Pattipola. The patch of white hair below the callosities is quite distinct.
The fourth example, ticketed merely Ceylon and received from Dr. Kelaart,
is too old and worn to call for comment.
Hitherto this langur has always been cited as wrstnus ; but monticola appears
to be the older of the two names. It was given by Kelaart to a langur from
the higher part of the Kandyan Province, Nuwara Eliya, which was described
as similar to typical cephalopterus (== vetulus) but without white on the inner
side of the thighs and on the rump. It was also said to be larger and darker
than its low country relative, with a rufous tinge on the neck and longer and
more wavy hairs.
The following approximate measurements in inches are available from two
examples collected by Major E. W. Mayor :—
Locality and Sex sae ee Tail nes | Weight
F 7 : 7
Pattipola, 6,210 ff 5, ch ... 23¢° 243” rg 20 lbs.
Hakgalla Uva, 2 ine 214”7+ 2050 63” 163 lbs.
The type of monticola, a female from Nuwara Eliya, was probably young
since, according to Kelaart, its head and body measured only 17in., the tail
being 26 in. and the hind foot 6in. But he sent, he tells us, a larger specimen
to Blyth, which was deseribed by Blyth as wrsinus, as Kelaart stated in 1852.
The largest specimen, so far as I am aware, of which the dimensions have
been recorded, was measured by Elliot. The head and body were 28 in., the
tail 31 in. and the hind foot 7 in. It also came from Nuwara Eliya; but the
measurements were taken from a dried and probably stretched skin.
Setting aside Elliot’s specimen, it will be observed that, according to Major
E. W. Mayor’s recorded measurements, monticola is no larger than P. s.
vetulus and P. s. nestor. It is true that made-up skins of smonticola look
larger, but this is no doubt largely due to the thickness of the coat. There
is nevertheless considerable cifference between P. s. monticola and P. s.
vetulus in weight, monticola: being much heavier. Moreover according to
the measurements recorded by Hinton in his paper quoted above, the skull
of monticola (=ursinus) is about 10 mm., nearly half an inch longer than
the skulls of vetwlus and nestor.
Subspecies johniz, Fischer.
Simia johnit, Fischer, Syn. JZamm., p. 25, 1829.
Semnopithecus, Presbytis or Pithecus qohnit or johni, Anderson, Blanford,
Elliot and most recent authors.
A mountain-race of large size, nearly as large as monticola but with the coat
shorter, closer, blacker and more glossy on the body and limbs, the general
colour being jet black, or nearly sv, above and below. It differs also from
Pithecus senex monticola in the uniform buff-brown tint of the long hairs
of the crown and of the whiskers and throat, there being no contrast in tint
between the whiskers and crown; the tail alsois nearly black and not very
noticeably paler than the middle of the back, as in the Ceylonese fornis.
Distribution. Mountains of Southern India, the Anamalai, Palni and
Nilgiri Hills and the Western Ghats from Coorg to Cape Comorin.
504 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
The following table gives the approximate dimensions in inches of examples
of this langur collected for the Mammal Survey by Mr. G. C. Shortridge and
Mr. Riley O’Brien :—
Locality and Sex | pee | Tail Hind foot | W
Brahmagiri Hills, 8.
Coorg, 2,780-5,138 ft.,¢ 26% 303” 7" — 214 Ibs.
Kumblacodie, Cochin, |
3°500-£t.,, ¢ zee 2a 344” 74”
Anamaad, S. Malabar,
3,200 ft., ¢ | 224” SWE | ES
Kukkal Shola, Palnis,
6,100 ft; 2 ae: 24” 32% | igs [25> bse
Made-up skins of this langur appear to belong to a considerably larger form
than the typical Ceylonese vefulus and nestor. But the dimensions show
that there is not very much difference. It is, however, a heavier animal as
indicated by the weight. I have not seen the skull of a fully adult male, the
one measured above 268 having the occipital suture open and the upper canines
hardly longer than in the female. But the skull of the fully adult would
certainly be larger than in the Ceylonese races vefulus and nestor and the
teeth are larger.
Although this langur has always hitherto been regarded as a species distinct
from the related Ceylonese langurs, it does not appear to me that its characteris-
tics should be given more than subspecific importance.
( Zo be continued)
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A; COLLECTING TRIP’ TO LADAK
BY
V. S. LAPERSONNE
Asst. Curator, Bombay Natural History Society
PART I
(With a map and tive plates)
It was a happy opportunity that allowed me to hitch my cart
behind that of Col. Meinertzhagen’s, when he organized a collecting
trip to Ladak and on to the borders of Chinese Tibet for the purpose
of acquiring a thorough knowledge of the resident bird life at the
higher altitudes, and also authentic proof of the migratory species
that cross the Great Himalayas and breed in the highlands of Ladak
and Little Tibet. The success of this collecting enterprise and the
subsequent recognition of new species added to the fauna of Ladak
was the work of a congenial employer and friend and a first-class
organizer. It is to him I owe whatever knowledge I have of
Systematic collecting and of preparing bird skins that will be a
‘thing of joy for ever.’
The path of a bird collector is not always strewn with roses. His
peregrinations may take him, at times, well beyond the pale, where
perhaps, through sheer malice, he may see a‘ snow man’ and come
back to us with photographic proofs of human foot-prints in the
snow.
On the other hand the field is infinitely varied. There are hunting
grounds to suit all tastes. The arid desert, the tropical forest with
its varied and multi-hued denizens, or the bleak glacier-hung moun-
tain top.
Col. Meinertzhagen and I left Pindi on February 21 and passing
up the picturesque Jhelum Valley arrived in Srinagar on February
27, having collected a good many migrants on the way.
We left Srinagar on March 23 and crossing over the Great
Himalayas at the depression known as the Zogi-la Pass, followed
the Indus Valley and reached Leh on the 6th May. Leaving Leh
on the 17th May we crossed over the Ladak range by the Changla
and Marsimik la passes and reached Changchen-Mo Valley on the
31st May.
Retracing our steps we made a wide detour, skirting the
Pangkong Lake to Shushal and thence crossing the Indus reached
Puga on June 20. Returning to Leh we struck north and arrived at
the foot of the Kara Koram Range on July 22.
We returned again to Leh and worked down the Indus to Skardu
in Baltistan on August 13. Thence we returned to Kashmir by way
of the Deosai Plateau and reached Srinagar on September 5.
During the period of collecting we covered over 2,000 miles and
in the course of our travel reached an altitude of 21,050 ft. in the
Kara Koram.
The Vale of Kashmir is described asa saucer-shaped valley between
the Great Himalayas and the Outer Himalayas or the Pir Panjal
506 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Mountains between the Great with a length of about 80 miles, a
breadth of 24 miles, and a mean altitude of 5,600 feet above sea-
level, corresponding in latitude to Palestine, the Mediterranean
and California. ‘Thus it has a blend of temperatures which should -
compare favourably with any other summer resort in the world.
For its genial beauty Kashmir should stand alone, for here you
have a combination of lakes, the quiet beauty of cultivated fields
and gardens, and a snow-capped mouutain range which completely
encircles the luxuriant valley.
It would be as great a mistake to generalize on the character
of the Kashmiri, as to accept the theory that all Kashmiri
women are beautiful. Yet if we must err, let it be on the side of
caution. Sir Francis Younghusband, for many years Resident of
Kashmir, has written a very interesting book on this province, and,
as a man who knows what he is talking about says, ‘A Kashmiri
soldier is almost a contradiction in terms. There is no such thing.
They will endure and suffer but they will not fight. And they are
very careful of the truth.’ Col. Meinertzhagen added a rider to
that by saying, ‘ They areso careful of the truth that they seldom
use it.’ However, it isto their splendid endurance there we owe the
success of our enterprise.
We had arrived in that ‘ Valley of Bliss’ just when the almond
blossoms were in bloom and Kashmir was warming up after a
rather severe winter. We had intended staying in an hotel, but this
became impossible. Being the earliest visitors we were preyed
upon by every shopkeeper who had anything to sell. The
climax of Col. Meinertzhagen’s patience seems to have been
reached when a none too clean fellow offered to give him a Turkish
bath !
- The ‘ Season’ had not begun when we arrived in Kashmir and
the houseboats were piled up along the banks. The Kashmiri boat-
man claims Noah as his ancestor and if, as Younghusband says,
‘they didn’t borrow the pattern of their boats from Noah, he must
certainly have borrowed it from them.’ They are known as
MANJIS and live entirely on their boats with their families and
cattle, and one can picture the squalor when all agree that the
Kashmiri could make Kashmir prefection if only he would wash.
We engaged a houseboat which gloried in the name of ‘ Arabella’
but nothing else happened; and slipping anchor from a regular
hulabaloo of ‘ Papier Mache Sir.’ ‘ Shampoo—furst Class 2?’ and
one or two pointed remarks as ‘ Hair cut and Shave Sir?’ we
worried our way down stream through the crowded waterways to
the accompaniment of frantic yells of our boatmen anda return
‘cuss fire from Kashmiri bargees’ wives, which if understood
suits the circumstances down to the ground.
We glided down the river passing the Maharaja’s Palace and just
beyond the precincts of the city a fair amount of dead horses and
cattle, which the Kashmiri delights in pushing into the river when
their days are done.
The most conspicuous object inthe immediate neighbourhood of
Srinagar is Takht-i-Suleiman, ‘The throne of Solomon’ a bare
rocky hill rising 1,000 feet from the floor of the valley, and sur-
Journ., Bombay Nat. Hist. Soc.
- Sinp VALLEY, ONE OF KASHMIR’S PICTURESQUE SIDE VALLEYS.
HLIM ‘SVAWIVWIF] LVEXD AHL NI MVAQ SNONDIASNOD V ‘LA 0O8‘ZT ‘IOHVIOY
— Ia xy re y ee See eZ ie, %
"205 “3S!IPT Wey) Aequiog “uinor
A. COLERGTING. TRIP TO LADAK 507
mounted with a Hindoo temple. The view from this height will
repay the climb. On one hand the climber looks down on the town
of Srinagar, the Venice of the East, with its numerous waterways
and gaily coloured boats, following the course of the Jhelum with
little hamlets and temples clustered on its banks, and then away-
over the valley the land rises to the pine-covered slopes and the
culminating touch to every Kashmir scene—the snows. Looking in
the opposite direction the view is not so extensive. In the fore-
ground the Dal Lake with stately poplars on its shores and on its
placid waters the famous Isles of Chenars, mirrored again in its
crystal clear depths. Fleecy clouds crept over the snow peaks that
strung out in a jagged line against an azure sky while a bluey haze
hung over the valley and the sparkling domes of the temples sent
back the rays of the setting sun.
We camped in the Jee of acraggy hill and had an uninterrupted
view across the still waters of the Wular Lake to the stupendous
snow-clad mountains, that stretched across the eastern sky. The
pass into Ladak and the land of the laughing Bhudda led over these
mountains, but was considered impassable at this time of the year.
We decided to wait a month, and fill in the time collecting birds in
the vicinity.
Few of the summer visitors, in the course of their migration from
the plains and foot hills had arrived, though in those quite historic
gardens of Shalimar where the ‘ playing fountains ’—never play, the
Paradise Flycatcher, or as he is known to the residents of Kashmir
the ‘Ribbon-bird’, with a flash of satin white, darted among the
dark green of the chenar trees.
At dawn the call of that game little fellow, the Chukar, who can
be relied upon to give a good morning’s sport, is echoed back and
forth among the hills. Higher up on the pine-covered tops are the
haunts of the Koklass (Pucrasta macrolopha biddulphz), and the Nut-
cracker (Nuctfraga multipunctata).
In the valley the House Crow is entirely replaced by the Jackdaw
during winter, and the sight of these birds leaving the valley in
thousands every morning for villages along the Jhelum Valley and
returning to roost among the chenar trees in the heart of the town,
can hardly be missed by any visitor.
With the rise in the temperature and the melting of the snows on
the heights, we decided to commence the trek that was to take us
into Ladak and allow us to dangle our feet into Chinese Tibet. On
the 25th March we bade farewell to the ‘ Arabella’ at the mouth
of the Sind River and moved up that beauteous valley, which at any
season surpasses in beauty any other side valley of Kashmir. A
thick forest of fir covered the northern slopes with glimpses of snow
showing through the dark green, and toy-like villages clinging to
the lower slopes.
Such natural beauty of scenery, the varied bird life and exhile-
rating climate enhanced our collecting spirit considerably. Even our
local shikari, who usuaily suffered from a stomach or a backache,
‘girded up his loins’ to bag the wary Monal Pheasant, though
these when brought in were too full of holes and lead to be of much
use for specimens or the pot.
508 JOURNAL, BOMBAY NATOGRAL FIST. SOCIETY, Vol. X2°2g7
Small parties of Tits, Tree-Creepers and Nuthatches, which
consisted of Parus major kaschmtriensis, Parus melanolophus, Parus
rufonuchalis rutonuchalis, Sitta europea kashmiriensts, and Certhia
himalayana(?) frequented the forests. The sub-specific rank of
this last birdis under discussion as being intermediate between the
Himalayan and Turkestan races.
The Orange Bullfinches (Pyrrhula auzvantiaca) were not un-
common in the fir forests. Col. Meinertzhagen writes of this bird in
the /ézs of July 1927, ‘The brilliant orange of their plumage
showed up like some precious stone on the drab dull green of the
ground on which they fed. Aftera moment they would fly up into
a tree, soon to return one by one and sit quite still munching small
seeds. Their dislike of movement when feeding and when in a tree
was due to their inherited consciousness that in such brilliant plums
age they would become visible and an easy prey to any feathered
enemy.’
As we moved higher up the vailey, forest bird life became
Scarce |
The few that braved the winter conditions at 8,000 feet were the
Brown Dipper, White-capped Redstart, Guldenstadt’s Afghan Red-
start, and higher still, the White-breasted Dipper.
On the 7th April we left Traghol, 9,500 feet. This was our fourth
day in this picturesque valley. Each day brought forth more
wonders in mountain scenery ; each turn in the road a surprise.
The thick belt of fir trees thinned out ; giving place to the slender
hardy birch. Snow lay thick everywhere, at times completely
bridging the river, and then one enters Sonamarg. Glaciers move
down the flanking mountains, and in spring, the snow having melt-
ed, the glacial formation reaching down to the tree line, is a mass
of dazzling ice and bare jagged rocks.
Kolahoi, 17,850 feet, its sharp peak conspicuous from any point
along the Sind Valley ridge is the highest point in the Great
Himalayas. The peak is six thousand feet above the glacial
formation, which stretches down to the tree line.
We arrived at Boltal 9,700 feet on the 10th April. It has a
delightful dak-bungalow at the head of the Sind Valley, and is
completely surrounded by the wildest and grandest scenery of
snow, mountain and mist.
This is the stage when the traveller realizes what he is up
against. The Zogi-la Pass which crosses the Great Himalayas
into Ladak and which is usually open to traffic after the 15th May,
was bunged high with snow; the only alternative was a snow-
bound, precipitous ravine that really was a frozen mountain torrent,
covered over with snow.
We remained three days here and every day were made
acquainted with the awe-inspiring sight of avalanches. With the
thawing of the snow, a few tons would rush down the slopes with
the roar of an express train.
The traveller can never be too cautious in his plans of an
assault on these mountain fastnesses during winter.
He must be up and away before dawn to avoid the avalanches,
his face and hands well greased to avoid frost-bite, his feet shod
AvCOLLEGTING TRIP TO LADAT- 509
with grass shoes fora firm foot-hold on the frozen surface of the
snow, and, as fancy dictates, a flask of rum or brandy, or if this is
‘haram,’ chocolate will be found an admirable substitute.
At an ungodly hour in the early dawn of Easter Sunday on
April 13 when the temperature was below zero—and our circulation
almost that—we set out for the assault on the snow-bound ravine.
Ten minutes brisk walk brought us to the foot of the ravine.
Here each man readjusted his load and snapped his belt into the
‘starvation hole.’
The coolies were dimly visible in the darkness which precedes
dawn as they grouped together for a final smoke. Their gnome-
like figures stood out against a sombre sky, and little points of
light from their cigarettes, stabbed into the darkness. Col.
Meinertzhagen gave the signal and we fell into single file and began
the assault.
That scramble up the ravine lasted a little over an hour and for
the greater part it was a ‘ fly-on-the-wall’ business. To add to the
discomfiture, a howling gale, accompanied by a blizzard, tore down
the closed-in ravine. ‘A gale,’ as our cook said, ‘ guaranteed to
disembowel a camel,’ or the equivalent to that in Hindustani.
Looking back on the events of that day, my diary helps with the
one elucidating word—‘ Hell,’ followed by notes of exclamations,
queries and asterisks, which apparently had its soothing effects, and
continues ‘ But still the wind blew, howling over the peaks, the
stinging snow finding its way down your neck, in your ears and
nostrils. The fine snow being blown from the heights, resembled,
as it caught the sun’s rays, a spreading white fire.’ We topped the
pass and paused for a ‘breather’ and watched the coolies ; these
gallant fellows who will not fight, but who can be thoroughly relied
upon in these mountains, were putting up a final spurt to gain the
top, and as they filed past with their 40 lbs. loads, we could not help
but cheer.
Physical disasters have made the Kashmiri selfish ; tyranny has
made him a liar, and superstition has made him timid. Yet, as he
swings along, to the lilt of a mountain song, indifferent to the cold
or steep ascents, we have nothing but admiration to offer.
We were now in that range of snows, the snowy range par ex-
céllence, which extends in a southern direction ina line of snowy
peaks, seen from Simla, to the famous peaks of Gangotri and Nanda
Devi in Garhwal. And we struggled along over a snow-bound
freezing cold valley, among the snow clad giants for three days,
seeing extremely little of bird life. Tie winter conditions prevail-
ing in the region, would hardly appeal to any birds, not even to
that curious bird which seems to spend its life either flying at
ridiculous altitudes or sitting on bare rocks—the Chough.
As we neared Ladak the temperature rose, though the altitude
(11,500) remained the same, and on the 16th April we were thank-
ful to see the last of the ‘snowy range par excellence,’ and which,
omitting those unrivalled mountains seen from Darjeeling, is second
to none.
We fully expected to see, after crossing the Himalayas, a broad
undulating plateau stretching away from our feet to the eastern
15
510 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
horizon ; instead our vision was plunged into a conglomeration of
barren hiils, parched valleys and gaunt Bhuddists’ monasteries,
which seem perched on every peak.
The track now followed the contour of the hills, with not atree to
break the monotony of barren hillside or a river cradled in the
valleys. At Molbek the atmosphere, people, and villages entirely
change. We were walking now on ‘the roof of the world,’ among
an ancient race, with still more ancient customs. Monuments, .
prayers and supplications carved and inscribed on stone, are piled
high along the caravan route which reaches to Leh, the capital of
Ladak,
We put in a few days at Molbek chiefly for the Himaiayan Snow
Cock (Tetraogallus himalayanus) which were quite common on the
hills, and secured quite a good series with the help of Tibetan
guides who take their pleasures in dead earnest, and appear to de-
light in seeing you, purple about the gills, climbing to 14,000 or
16,000 feet in pursuit of these birds.
We found that these descended tothe river at an early hour of the
morning for a drink, and, as the day warmed, moved higher up—at
times to the summit of the bare hills—where they roosted during the
warmer hours of the day, at elevations of 14,000 and 16,000 feet.
They rarely fly, and then only to cross the valley, if hard pressed by
asportsman. ‘They weigh about 5 lbs. and it takes a good knock to
bring them down. During the breeding season they utter a long
drawn-out whistle—a dismal sound well associated with the dreary
isolated haunts of this bird. The limits of the Himalayan species
appear to be at 16,000 feet, and roughly all that region south of the
main Ladak chain. Further north, the Tibetan Snow Cock replaces
the Himalayan.
To refer again to the Tibetan gun-bearer : he hasno equal in these
hills, possessing, as he does, an unflagging spirit, an unquenchable
zeal and a clean sense of humour—but keep to windward of him, for
if to his lee you can judge the full extent of his cleanliness! !
The road now follows the course of the Indus and we plunge again
into a labyrinth of confined valleys and gorges, shut in on either
hand by vertical walls of rock, and emerge with the Indus into a flat
bottomed broad valley that gently slopes to the foot of the Ladak
range, where the flat roofed houses of Leh crowd around the
monastery, aschickens do round a well-fed hen.
Leh, the great emporium of the trade which passes between India,
Turkestan and Chinese Tibet, has been the starting point of many
an adventurous journey into the last named country. The town does
not seem to have much of a historical past. The unwarlike Ladakis,
even with nature fighting on their side, were conquered by the
rulers of Kashmir. The Kashmiri troops left to garrison Leh were,
on one occasion, slaughtered by a strong force of Chinese from
Tibet, since when Ladak, though still a province of Kashmir, is
ecclesiastically subject to Lhassa, paying tribute, in the guise of
presents, to mollify the Chinese who are held in wholesome fear by
the Ladakhis to this day.
The Ladakhi, or ‘hybrid’ Tibetan is of a mild easy-going disposi-
tion, thoroughly honest, with the thinking capacity of a Yak,
Journ., Bombay Nat. Hist. Soc.
FEET.
MARKET PLACE AT LEH, AND THE ‘ GOMPA’ ABOVE, THE WINTER QUARTERS OF THE
ECCLESIASTICAL HEAD OF WESTERN TIBET.
HHI OL
Ha] AO
‘AONVY MVAVW'Y] a
AVMHLVS) HHL HOAOMHL ONIMOO’T
‘205 ‘slp ‘yey Aequiog “usanoe
A COLLEGTING TRIP 70 LADAK 511
The effects of high altitude may account for his wholesome horror
for any form of manual labour and he seems fortunate in possessing
a spouse who fills the roles of husbandman and handmaid. A
part of her dowry is a turquoise studded headgear—the remainder
of it—her slave-like willingness to work.
The price of a bride or brides, fluctuates with the supply, and on
our visit in 1925, the women constituted 8U per cent of the population
of Leh. What a happy hunting ground is Leh for our be-(k)-nighted
bachelors of Bombay !
Since time immemorial, man’s soul has yearned for a ball of some
kind. An ‘oblong’ one, a wooden one, or even a rag one. Buta ball
of some description it must be and until he gets it, he is never really
happy. The Ladakhi is no exception to the rule. Every fair sized
village has its polo ground and though the game, as played in the
plains of India, is atame affair to the one played here, there is a
general tendency to adhere to rules.
_ Throughout the hour’s play there is no change of ponies, the
game is fast and furious, and the ball never out of play.
Fouls are unknown; in a scrimmage, you may take a ‘ whack’
at your opponent or his horse. On occasions the game is played
down the broad main street, and norule exists preventing an attack
from a player who has lain in ambush in one of the shops flanking
the polo ground.
The word ‘ pulu’, which means ‘ ball’ in Tibetan has given rise to
the theory that polo originated among the Mongolians, but history
tells us that the game drifted from Persia through Turkestan into
China and Japan, thence from Tibet to Gilgit and so to the plains.
The earliest records of this ‘game of kings’ in Tibet seem to
tend more to feats of horsemanship and skill with stick and ball,
than to the scoring of goals.
The nature of the present game is a mixture of thetwo. The ball
is brought into play, in the event of a goal, ina most skilful manner.
A player gallops at full speed down the ‘ wing’ and, near the haif
way line, still at full gallop, tosses the ball into the air and smites it
square towards his opponent’s goal.
Being the centre of the trade of these parts, Leh boasts a reason-
able sized bazaar and market.
Goods are more often bartered for goods than sold forcash. The
Yarkandi from over the Kara Koram arrives in Spring with his
caravan of wooland dried fruits, exchanging these for silks and
wearing apparel from Kashmir. The Tibetan after a long and
arduous journey over wind-swept desolate plains, arrives with his
loads of Chinese brocade and quaint ornaments. The Kashmiri
floods the market with all conceivable goods, even to the extent of
Huropean provisions. The Ladakhicontributes an occasional Yak
tail.
Above the market and perched on an impassable rock are the
winter quarters of the ecclesiastic head of Western Tibet. Like
most monasteries it has an equally impassable ‘ stairway ’ leading up
to the entrance, but offers no other obstruction to sightseers.
Bhuddhism may not interest everyone of us, and few travellers
have the time to ‘worry’ about such matters. We accept the
§12 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
existence of those silent edifices as part of the abominable landscape
and pass on. Yetin those silent evenings, when the clouds have
crept up the valley, the monastery sticks out above like an isoiated
island in a billowy sea; when the still night air throbs with
the sonorous convocations from the deep-throated trumpets up in
the monastery, when the moon shines down on the unscrutable
face of Bhudda—it would almost seem as if Bhudda walks this earth
again.
We have not the desire to tackle Bhuddhism, nor the time to dip
into dimmest antiquity. With our object ever in view we begin
preparations to leave Leh for the borders of Tibet proper, the
region of the wild yak, the wild ass and the fabulous unicorn of
Tibet.
No jungle exists in the whole of Ladak; the few trees, mostly
willow and poplar, are planted in villages for building purposes
and fuel. The traveller will find no ‘ shady’ tree under which to
have his Junch between stages and, as he travels higher the gardens
around villages will thin out and disappear.
The bird life then, for the most part, is terrestrial in habits and
palaearctic in its zone of distribution. Thus the traveller meets,
when he crosses the great Himalayas, the Magpie (Pca pica
bactriana). Col. Meinertzhagen points out, in his paper on the Birds
of Ladak in the /ézs, that the Magpie is essentially a bird of the
wind-swept, open and almost treeless Ladak, in contradicton to
Baker’s habitat of this bird as being Kashmir, Kumaon and Gharwal.
Visitors to Kumaon and Gharwal will bear out the statement of
Col. Meinertzhagen that the Magpie is not a bird of ‘ well-wooded ’
parts. Observations in Mesopotamia, Persia and Baluchistan, all
countries that could do witha bit of vegetation, shew that the
Magpie frequents the upland steppes.
The Tibetan Raven (Corvus corax tibetanus) is met with all over
Ladak. His bell-like tones should be unmistakable to any visitor
to the highlands of Ladak.
The Red-billed Chough (Pyrrhocorax pyrrhocorax himalayanus)
extends over the greater part of Ladak, chiefly over 12,000 feet.
It occurs on either side of the Great Himalayas as low as 6,000 feet
in winter, up to 19,000 feet in summer.
The Yellow-billed Chough (Pyrrhocorax graculus forsytht) hardly
distinguishable, on the wing, from the above bird appears to fre-
quent the same localities—though not quite so common. Both these
birds were breeding at Leh at 11,600 feet. Horned Larks (Zvemo-
pbhila alpestris longtrostris) were common from 12,000 feet to the
snow line.
The Great Tit (Parus major kaschmiriensis), common on either
side of the Himalayas, extends for a considerable distance into
Ladak, wherever there were trees.
Snow-Cocks (Zetvaogallus himalayensis himalayensts) and the
Chukor (Alecloris graeca pallescens) as distinguishable from Alectorts
graecca chukar, a darker race, were common inthe alpine regions
of Ladak.
We left Leh on the 17th May for the valley of Chang-chen-Mo,
245 miles westward, which is the furthest point reached by any
————
A COLLECTING TRIP TO LADAK S13
traveller, for beyond lies a wilderness of the coldest, bleakest and
most desolate region conceivable,
We crossed over the Ladak range by the Chang-la pass, a regular
corker in the way of passes—and one which touched the scale at
18,000 feet.
We had camped just below the pass at 15,700 feet the evening
before the ascent and were fortunate in securing a fine series of
Brandt’s Mountain Finch (Montifringilla brandti hematopyga).
They appear to be birds of the extreme altitudes and are met with
usually on the fringe of the snows, upto 18,000 feet.
Before we crawl over the pass, I must dip my flag to Col.
Meinertzhagen, who, with the tenacity with which leaders are made,
got us over the pass when things looked their worst.
A relay of coolies were engaged to prevent any unnecessary
delay on the pass, for what with the height and the thawing of the
snow progress must be slow. There is an added danger after
sunset on such passes. A caravan caught after sundown at 18,000
feet, would find it difficult to get through without losing a large
part of its ‘cattle’ and men. Sucha catastrophe cannot be pre-
vented in view of the sudden drop in the temperature after sunset.
When tackling a high pass, you must, like Felix, ‘keep on
walking’. It is preferable, tobecoming a ruddy icicle, and helps
the circulation. Your progress may resemble the gait of a beetle
over the ridges and furrows of a ploughed field, but there is always
the consoling thought that all passes have a summit.
As much as I would like to forget this one instance of the effects of
high altitude, it may not come amiss to give first hand experience.
At altitudes over 16,000 feet, or at times 15,000 feet, it depends on
one’s physical condition, the traveller begins to feel the oppressive-
ness of the atmosphere. The mere act of turning over in bed isa
respiratory exertion; while ordinarily the respiration resembles
the effects of a second hand bicycle pump.
A generai listlessness is first observed, followed by a splitting
headache, as if a smack has been taken at the skull with an axe—
and the iron left in. Follows, if you have got it bad, a violent
desire to be ‘ sick.’
I doubt if there is any preventive for these effects, though
‘ Crystallized Ginger’ may be recommended.
There is also atendency to knock off ‘ food stuffs, ’—particularly
meat, though the craving for ‘ sweet stuffs ’, such as sugar and jam,
increases.
If on the other hand you are one of those who are blessed with
the innards of a Tibetan—the symptoms will be noticeable in your
camp. Your cook, if you haven’t left him to die on one of the
passes, usually finds he has a shocking grievance against the
Shikari or vice versa. This being appeased, he develops a com-
plication of disorders and a desire to peg out.
We crossed over the Changla pass and eleven miles beyond
camped in a willow garden at 14,500 feet. Just beyond the
pass we secured a Goosander (Merganser castor) which occurs
sparingly on large sized rivers above 10,000 feet. Later on (July)
we found the young in down on the Indus River below Leh.
514 JOURNAL, BOMBAY NATURAL HIST: SOCIETY, Vol. XX
The Brahminy Duck (Casarca furruginea) was first met at about
16,000 feet though they became commoner as we approached the
more isolated watered valleys.
We now entered the higher uplands or the ‘ Changlung ’—a name
given to all that region north of the Ladak Range, and the floors
of whose valleys rose from 14,500 feet to over 16,000 feet in the
Chang-chen-mo. ‘The mountainous chain that cuts across from
northwest to southeast is a continuation of the mighty Kara
Koram Range, which can be a real nuisance in its quiet
way.
The pass over this range, the Marsimik-la Pass, topped the scale
at 18,500 feet, and the ascent from 15,000 to the summit, in less
than two miles, is a steady pull.
We had left Phobrung, which is the most easterly limit of ‘ civiliza-
tion,’ in its broadest sense, on the 28th May and getting over the
pass in fine style camped at 17,700 feet on the site of a Tartar camp.
Our transport consisted entirely of yaks, or to be more correct
the hybrid known as ‘zhoe.’
Wild yaks occur on the Chinese-Tibetan border at extremely high
altitudes. Pocock mentions them as occurring in the Chang-chen-
mo valley at 20,000 feet in summer, descending toless snow covered
country at 15,000 in winter. According to local intelligence the
wild yak breeds between 20,000 and Say feet, and the young are
captured soon after birth.
The method of capture, and we give it for what it is worth, is for
a party to rush down on the parent yak, with tom-toms and cat-calls
thus driving it off. Itis hard to believe anybody ‘ rushing’ about
at these altitudes, burdened with tom-toms to say nothing of great
demands on the respiratory organs without cat-calls! The Tibetan
is a wonderful fellow, and if, as some affirm, his pig-tail is used as an
appendage by which Bhudda hauls up the ‘ failful’—to Nirvana, we
should think that even without it, his sheertoughness ought to get
him there.
At our camp at 17,700 feet it had snowed heavily all night and we
had become anxious for our yakmen. But any anxiety was unneces-
sary. These hardy mountaineers had bunched together and allowed
the snow to pile up over them and though not quite so comfortable
and hygienic were decidedly warmer than we in our flapping tents !
‘The ‘ Zhoe’ will be found the best mode of transport over heavy
snow and high passes. The novice will find he has not many
lengthy intervals of rest once the animals get amoveon. The
‘older beasts appear to be ever thinking of a point ahead which they
would rather have behind them.
On the 30th May we entered Chang-chen-mo valley. If there be
any colder bleaker spot in this world, let me not know it.
When I look at my diary written at these altitudes, the general
tone shares the pessimism of our cook. North, East or West was
an unbroken panorama of bleakness and desolation, devoid of vege-
tation or any sign of human habitation.
Then, as if not satisfied with her effects, Nature let loose a pierc-
ing wind down the narrow valley, which very nearly froze the
marrow in one’s bones.
Journ., Bombay Nat. Hist. Soc.
<
\
GU
ty,
y
CAMP AT CHANG-CHEN-MO VALLEY, 14.5JU FT
BREEDING GROUND OF BRAHMINY DUCK
A COLLECTING TRIP TO LADAK 515
However we seemed amply repaid for these discomfitures, from
a natural history point of view of course.
The Tibetan Snow-Cock ( 7etvaogallus tibetanus tibelanus,) replaced
the Himalayan species but were harder to obtain owing to the high
altitudes (17,000 to 20,000 feet) which this bird frequents. In
habits they do not differ from the other bird and appear not quite
so shy.
The semblance of the track we had followed now disappeared
entirely.
We observed a herd of Wild Asses that stood and took stock of
our party, while we stumbled over the boulders. It may not do
your temper much good to have a Wild Ass with his ears cocked,
looking down on you as you worry your way along, but you soon
get used to him and the ridiculous capers he cuts around the lag-
ging transport.
The sight of aman seems to excite his curiosity immensely, and
he will at times approach to within 25 yards for a thorough in-
spection of the biped.
I was able to observe a fine specimen at close quarters. After
semaphoring with his ears he decided to take a walk round and
then for no apparent reason, spun around and fled as if the furies
were behind him. One could understand this behaviour if he had
had a Tibetan to windward.
At the time I had put it down to my Kashmiri-cured posteen—but
sportsmen have commented on this behaviour and its disturbing
effects on other game—particularly Ovzs ammon and the Tibetan
Antelope.
The Tibetan Antelope, we find from ‘ Al Khanzir’s’ narrative in
one of Blackwood’s numbers to be the ‘ original’ unicorn of Tibet.
The horns of this animal are so placed that in profile they resemble
a single horn.
The Tibetans most naturally do not admit this and ‘ Al Khanzir ’
gives us the legend ‘that should a unicorn espy a virgin he will
instantly lay his head in her lap and look up into her eyes like a love
sick swain’.
Such an incident has never occurred and though maidens are not
plentiful in Tibet, they certainly do exist !
We shifted camp the next day, and ‘ discovered’ a delightful spot
on the lee ofa high cliff. A clear stream flowed past our camp,
while all around were bushes and a grassy lawn.
The Brahminy Duck had its young here, allin down. Theegegs
are laid in a hole on the hillside and the young as soon as hatched
are carried down to the water by the parent birds. A brood con-
sists of six to eight eggs.
A common bird. and one which turned out to be a new race,
was the Ladak Twite. Col. Meinertzhagen has named this Acanth?s
Hlavtrostris ladacensts and gives the following description :—
‘ Ladak birds are thick-billed as in rafosirigata, but differ from
that race in fresh plumage in having the upper plumage not so red.
In worn plumage Ladak birds are scarcely distinguishable from
rutostrigata. They differ from montanella in having a richer and
redder tone on the upper parts.
516 JOURNAL, BOMBAYINATOCRAL ‘HIST. SOGETY, Vol. X27
‘ A series of ten birds were obtained at Chimre (11,900 feet, May),
Tankse (12,900 feet, May), Chanchenmo (14,800 feet, June), Pang-
kong Lake (13,900 feet, June), and Moulbekh (10,350 feet, August),
Wings of five males 80-81 mm. and of five females 73-81 mm.
‘The Ladak Twite is common wherever met. It is apparently
confined to Eastern Ladak (my Moulbekh specimen being a soli-
tary bird andprobably a straggler from elsewhere). Observed at
Zingral, Tankse, Pamzal, Pangkong Lake, Shushal, Puga, Tso Kar,
Debring, and at Khardung. In summer they seldom come below
12,000 feet or over 15,000 feet.
‘Nests with eggs were found at Tankse on May 21, on the
Pangkong Lake on June 8, and on the Tso Kar on June 23,
Clutches numbered three eggs in three cases and four eggs in
three cases. Nests in low bushes (once 7 feet from the ground
in a willow-tree), with thick lining of yak-hair.’
We left Chang-chen-mo Valley on June 4 and retracing our
steps topped the Marsimik-la pass and descended to Phobrong ina
single day. This may not sound anything like a feat, but the
reader should bear in mind the rapid descent from 18,500 feet to
14,500 and the sudden change in the atmospheric conditions. If
there in anything worse than a never ending track that zigzags sky-
wards, it is arapid descent of more than five miles over boulders
and loose gravel.
We pitched our camp at Phobrong on a grassy lawn that sloped
down to the Pang-Kong Lake whence it rose again to the great
heights of the Ladak Range, now wreathed in fleecy clouds.
Evening finds contentment everywhere; the yaks quietly chew-
ing the cud, the men huddled together over their ‘ All steel’ pipes,
and floating on the evening air the soothing sounds of the hubble-
bubble from the kitchen tent. Forgotten are the grievances, a
bond of friendship such as never could have existed in the higher
altitudes, has brought together the cook, the ‘pot walluper’, the
shikari and one or two parasites, ‘‘‘ The Burra Saheb’ is quite
a descent stick, and not what we thought him at 17,000 feet; the
‘Chota Sahib’ uses some vile language but we know he doesn’t
mean it. Anyhow we are going back’’’.
The cook firmly makes up his mind to give us a real change from
the eternal ‘estew’. Alas! these resolutions vanish with the last
pull off the hookah.
Our road now skirted the west shore of the Pangkong Salt Lake
that lay between the outer ranges of the Ladak and Kara Koram
Mountains. The Lake is the largest in these regions being nearly
thirty miles in length and about five miles in breadth. Such is the
intensity of the cold during winter in these parts, that villagers are
able to drive herds of yaks and ponies across its frozen surface to
the hill ranges opposite.
The Brown-headed Gull (Lavus brunneicephala), a familiar bird in
the Bombay Harbour and along the west coast of India, crosses over
the Himalayas and breeds on such lakes. On our visit (June 8) the
Gulls had attained their breeding plumage and by the size of their
organs we concluded that the shingle banks were used as nesting
sites.
A COLLECTING TRIP TO LADAK 317
About midway down the shore of the lake, we pitched camp in
open bush-covered country and were fortunate in securing a series of
the Tibetan Partridge, which have proved a new species and named
by Col. Meinertzhagen as Perdix hodgsonie caraganae.
‘Ladak birds,’ he writes, ‘are generally paler above, especially
the chestnut nuchal collar. The dark brown markings on the upper
parts of P. 4. hodgsonie are replaced by chestnut markings in Ladak
birds, though this is not invariable. In size they do not differ from
P. h. hodgsonie.
Type in the Tring Museum: <, Shushal, 15,000 feet, Eastern
Ladak, June 13, 1925.
Distributig. Shores of Pangkong Lake to Rupshu and Hanle.
Is confined to the valleys where the Tibetan Gorse (Cargan)
grows.’
(Zo be continued)
16
A LIST OF ORCHIDS, WITH SOME NEW SPECIES FROM THE
HIGH WAVY MOUNTAIN (MADURA DISTRICT)
BY
BE. BLATTER, S.J., Ph.D., Fels.
(With a plate)
The following is a list of Orchids which were collected in the month of May
1917 by the late Prof. F. Hallberg and myself on the High Wavy Mountain at
an altitude of 4,500 to 5,500 ft. The area examined measures about 12 square
miles, covered with dense evergreen forest and some smaller patches of grass-
land, apparently forest-clearings.*
Before publishing this paper I sent my MS. and specimens to Mr. C. E. C.
Fischer of the Kew Herbarium who is just engaged in writing the continuation
of the unfinished work of the late Mr. Gamble’s Flora of the Presidency of
Madras. As the next issue of the Flora contains the Orchidacez and as Mr.
Fischer knows the Madras Flora by many years’ practical experience in that
Presidency, I could scarcely find anybody better qualified to criticize my list.
Though I cannot agree on every point with Mr. Fischer’s views, I am greatly
obliged to him for pointing out a number of mistakes I had made. He even
discovered a new species amongst my material whichI had failed to recognize
as such.
1. OBERONIA, Lindl.
1. Oberonia Brunoniana, Wight Ic. t. 1622.—Nos. 329, 330, ? 26551.
Distribution: W. Peninsula: Konkan, N. and S. Kanara, Nilgiris, Pulneys,
mountains of Malabar and Travancore.
2. Oberonia verticillata, Wight Ic. t. 1626.—Nos. 26549, 26554.
Distribution : WN. Kanara, Nilgiris, Pulneys, Travancore.
3. Oberonia iridifolia) Lindl. Gen. & Sp. Orchid. 15, var. denticulata,
Hook. f. in Hooker FI. Brit. Ind. v, 676.—O. denticulata, Wight Ic. t. 1625.—
Nos. 331, 26550.
Distribution: Of variety: Nilgiris, Pulneys.
4, OQberonia Lindleyana, Wight Ic. t. 1624.—No. 26552.
Distribution: Nilgiris, Travancore Mts.
2. LIPARIS, Rich.
1. Liparis viridiflora, Lindl. Gen. & Sp. Orchid. 31.—No. 26546.
Distribution: Sikkim Himalaya, Upper Assam, Khasia Hills, Nilgiris.
2. Liparis Wightiana, Thwaites Enum. 295.—No. 26547.
Distribution: Pulneys, Travancore, Ceylon.
3, Liparis sp.—No. 26545.
Only one specimen in fruit.
3. DENDROBIUM, Swartz.
1. Dendrobium nutans, Lindl. Gen. & Sp. Orchid. 90 (including D. Jerdoni-
anum, Wight Ic. t. 1644), var. rubrolabris, Blatter var. nov.—Sepala necnon
petala alba, labium rubrum maculis purpureis.
* See this Journal, vol. xxv (Sept. 1917), p. 290.
NEW SPECIES FROM HIGH WAVY MOUNTAINS 519
The lip has 3 very high, close, crisped lamellae. Margin of midlobe very
strongly crisped. Nos. 335, 25808.
Distribution of type: Coorg, Nilgiris, Ceylon.
2. Dendrobium nanum, Hook. f. Ic. Pl. t. 1583.—Nos. 26523, 25810.—Flowers
pale pink, except lip which has a purple spot on each side.
Distribution: Malabar.
3. Dendrobium sp. near nanum, Hook. f.— Too young for proper identification.
—No. 25811.
4, BULBOPHYLLUM, Thouars.
Bulbophyllum nilgherrense, Wight Ic. t. 1650.
The leaves are subacute, not retuse as shown in Wight’s figure.—No. 340.
Distribution : Nilgiris, Malabar.
5. CIRRHOPETALUM, Lindl.
Cirrhopetalum Gamblei, Hook.f.in Hook. ¢. Fl. Brit. Ind. v, 778.—C. Thomsont,
Hook.f. l.c.—No. 332.
I intended to transfer this species to Bulbophyllum, following Smith ; Bulbo-
phyllum Thou., sect. Cirrhopetalum, in Bull. Jard. Bot. Buitenz., ser. 2, viii
(Oct. 1912), 19-29. But Fischer tells me that Kranzlin,Schlechter and Rolfe have
accepted Cirrhopetalum as a good genus, and he thinks that the unequal sepals
make a good character to separate it from Bulbophyllum.
Distribution : Nilgris, Pulneys, Bababoodan Hills.
6. CHRYSOGLOSSUM, Blume.
1. Chrysogiossum Hallbergii, Blatter, sp. nov. (Orchidacew— HKpidendree—
Dendrobie@). Similis Chrysoglosso maculato, Hook.f. sed recedit ab eo habitu
epiphytico, mento saccato, disco labii 3 lamellis predito, bracteis multo
longioribus.
Epiphytic. Pseudobulb 1-leaved, leaf with 5 large and very numerous close
finer veins. Scape lateral, about 12-flowered, with 3 sheaths. Bracts longer
than pedicel and ovary. Flowers pedicellate. Perianth-segments all 9 mm.
long. Sepais and petals obovate-lanceolate to lanceolate-acute, the latter
broader, all 3-nerved. ‘The lateral sepals united to the upper half of the spur of
the lip, forming asmall saccate mentum, below which the proper spur may be
distinguished, which is subcylindrical, slightly incurved and not quite 1 mm.
long. Lip clawed, 3-lobed, midlobe much longer than the side-lobes, suborbi-
cular, obtuse, with a very broad claw, base of claw minutely auricled, side-lobes
subrectangular, obtuse. Disk with 3 very distinct lamellee. Column slender,
bent forward, winged in front, the wings becoming broader and thicker down-
wards, forming a long foot, each wing having a large auricle about half-way
down. Anthers small, terminal, opercular, with a fiontal cusp, 2-celled, cells
distinct, hemispherical. Pollinia 2, free, subglobuse, waxy.—No. 26488.
2. Chrysoglussum sp. Probably aform of C. maculatum WHook.f.—Epiphy-
tic, notin flower. Found together with the new species, but the finer nerves
are not so close, and the leaves are in tufts, not solitary on the pseudobulb.—
No. 26487.
3. ? Chrysoglossum sp.--No. 26487,
7. Erta, Lindl. .
1. Eria braccata, Lindl. Journ. Linn. Soc. iii, 46.—4. reticosa, Wight Ic. t.
1637.—No. 338.
Distribution : W. Ghats, Konkan, Nilgiris, Ceylon.
2. Eria pauciflera, Wight Ic.t. 1639.—No. 337.
Distribution » Nilgiris.
3. Eria nana, A. Rich. ?, but too incomplete for certainty.—No. 25809.
4. Eria pseudociavicaulis, Blatter, sp. nov. (Orchidace—Epidendree—Eriee,
Sect. Hymeneriz). Distinguitur ab Eria clavicauli, Wall. foliis longioribus
520 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
non acuminatis, internodiis longioribus, racemo unifloro, bracteis oblongis
acutis haud nervosis, sepalo dorsali elliptico-oblongo obtuso, petalis subobtusis.
Epiphytic. Stems tufted, clavate, terete, with numerous longitudinal
fissures above, 20 cm. long ; last internode 7 by 1'5 cm., swollen in the middle,
Slightly oblique; next internode 7 cm., thickest on top, the following 5 by 3
cm., cylindrical. Sheaths 0. Leaves 3, 16-17 by 4-4°5 cm., oblong-lanceolate,
acute, coriaceous ; nerves numerous, stronger ones regularly distributed, bet-
ween which there are a number of very fine close ones. Peduncle opposite the
middle leaf, 22 mm. long, terete, pedicel solitary, strongly ribbed upwards, both
thinly brown-pubescent ; at base of peduncle a small sheath ; at junction of
peduncle and pedicel two large bracts which are olive coloured, oblong, acute,
half-amplexicaul, glabrous, hardly nerved, one lower than the other ; opposite
the upper is asmall upcurved awn,possiblv a reduced second pedicel. Flowers
faintly fragrant, glabrous, mainly white, mentum tinged with pink. Dorsal
sepal 15 by 6 mm., elliptic-oblong, obtuse ; lateral sepais very broad at base, 15
mm. long, triangular, falcate, subacute, reflexed, mentum 8 mm. long. Petals
falcately oblong, 14 by 5 mm., subobtuse. Lip sessile at the base of the
column, incumbent, much shorter than the lateral sepals, broadly obovate,
subequally 3-lobed with the margins light-purplish fading inwards, all lobes
slighly crisped, lateral lobes bent inwards, midlobe minutely white-puberu-
lous. Disk with a white pubescent ridge, 2 mm. broad in the lower half, which
portion has in front 2 small lateral calli, the upper half much narrower,
yellowish, hardly reaching the subacute tip of the midlobe. Column white, bent
nearly at aright angle to the axis of the flower below the stigma and produced
into along foot forming the broad mentum together with the lateral sepals,
foot broadest at the bend, marked further out with a few faint buff spots.
Stigmatic cavity broad, below the horizontal rostellum. Anther opercular,
embraced by the flaps of the column, 2-celled, cells approximate at base behind.
Pollinia 3 in each cell, connected by their bases, 2 much larger than the third,
compressd, pyriform, pale cream-coloured.
Described from onespecimen which was brought to Bombay where it flowered
in the beginning of September 1917.—No. 554, kept in formalin at St. Xavier’s
College, Boinbay.
8. JOSEPHIA, Wight
Josephia lanceolata, Wight Ic. t. 1742.—No. 333.
Pistribution - Central Provinces, W. Ghats from the Konkan to Travancore,
Ceylon.
9, Ca@LOGYNE, Lindl.
Coelogyne nervosa, A. Rich. in Ann. Sc. Nat. ser. 2, xv, 16.—C. corrugata,
Wight.
Side lobes of lip much veined inside with orange-yellow. The 3 raised ridges
of the disk yellow in front ; rest of corolla white. Pollinia pale yellow. Wings
of column orange.
Our specimen flowered in Bombay in September 1917.—No. 339.
Distribution « Nilgiris, Pulneys, Travancore hills.
10. AFRIDES, Lour.
Aerides cylindricum, Lindl. in Wall. Cat. 7317.—Nos. 122, 25801, 25806, 25807.
Sepals and petals of No. 122 white, lip pale pink.—Not common.
Distribution - From the Coorg Hills to Travancore.
11. SACCOLABIUM, BI.
1. Saccolabium pulchellum, C.E.C. Fischer.—S. nz/agiricum, Hook. f. in
Hook. Fl. Brit. Ind. vi, 60.—Vanda pulchella, Wight Ic. t. 1671.—No. 342,
Distribution : Nilgiris.
2. Saccolabium filiforme, Lindl. in Journ. Jinn. Soc. iii, 36. The colour of the
flower seems to vary a good deal. In Wight’s plant they are rose-coloured, in
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NEW SPECIES FROM HIGH WAVY MOUNTAINS 521
Ceylon plants orange with red stripes. Our flowers are bright brick-red, lip
orange, midrib of petals very prominent, ending in a point short of margin,
lateral veins very obscure, close to midrib.
Hook.f. (Fl. Brit. Ind. vi, 54) mentions expressly that the sac of Saccolabium
is ‘ without a large scale or callus within under the column’ except in S.
longifolium. There is one, however, in our specimens as well as in Fyson’s t.
250 illustrating S. filiforme, Lindl. See also Wight Ic. 1685, text.
Fischer who examined some flowers of the same specimen could not find a
trace of acallus. I presume that itis sometimes present and sometimes not.
In any case Hooker’s general statement about the presence of the callus will
have to be slightly modified. No. 189.
3. Saccolabium gracile, Lind]. Gen. & Sp. Orchid. 225.—No. 26531.
Distribution ; Ceylon.
12. CLErsosToMA, BI.
Cleisostoma tenerum, Hook. f. in Hook. f. Fl. Brit. Ind., vi, 73.
Sepals and petals green with purplish longitudinal markings. Lip white, side-
lobes yellowish with purple markings. The whole flower, except the midlobe,
sometimes assuming an orange hue in the same plant. leaves /7-nerved.
Rostellum 2-lobed at base, each lobe with a horizontal linear appendage ; in the
space thus formed rests the gland.—Nos. 30, 25805, 25803, 25804.
Distribution : Nilgiris, Ceylon.
13. ANAECTOCHILUS, BI.
Aneectochilus elatior, Lind]. in Journ. Linn. Soc. i, 178..—No. 834.
Distribution ; Nilgiris.
14. ODONTOCHILUS, Bl.
Odontochilus rotundifolius, Blatter, sp. nov. (Orthidacee-—-Neottie—Spiran-
thee) Caulis terrestris nanus procumbens; folia pauca, parva, rotundo-
mucronata. Flores albi; sepalum dorsale necnon petala apice brunnea.
Sepala erecta, dorsale quidem 2 cm. attingens, 5-nervosum, lateralia autem
16mm.longa, oblonga, 5-nervosa. Petala 15 mm. longa oblongo-acuminata
subfaleata, 3-nervosa. Labium 14mm. iongum apice 2 lobis elongatis fere
horizontalibus ornatum. Columna brevis appendice frontali praedita. Rostelli
brachia longa, unoquoque brachio lobulo unico ventrali munito. Forma
foliorum necnon sepala 5-nervosa hanc speciem a ceteris speciebus affinibus
distinguunt.
Terrestrial. Stem nodose, procumbent, creeping below, bulbous-based-hairy,
15 cm. high. Leaves few, almost orbicular, mucronate, the largest seen 2 cm.
diam., glabrous, midrib distinct on lower surface, stalk about as long as blade,
bulbous-based-hairy, amplexicaul. Flower terminal, solitary, nodding ; bract
broadly ovate-acuminate, as long as the ovary. Sepals erect (not spreading),
green at base, rest whitish; dorsal sepal 19 mm. long, concave-hooded, 5-
nerved, tip reflexed, brown ; lateral sepals 16 mm. long, oblong, subobtuse,
5-nerved, connate at the base and concealing the minute spur of the lip. Petals
15 mm. long, oblong-acuminate, subfalcate, 3-nerved, white, tips brown. Lip
white, adnate to the base of the column, base with a minute subsaccate spur ;
claw broad, entire,4 mm. long, with a few long hairs, then the lip is bulging
out into 2 rounded lobes for a distance of 5mm. and again contracting, after
that narrow-linear for 5 mm. with 2 short elongate obtuse lobes spreading
almost horizontally at the apex. Column short, white, frontal appendage ot
column cup-shaped, lanceolate-acuminate when seen from the side, lying in
the folded claw of the lip; calli small. Rostellar arms long, slender, each
with a minute lobe on the ventral side. Anther lanceolate-acuminate, curved ;
pollinia broadly lanceolate in outline ; caudicles long, slender. Ovary 5 by
2mm., slighly broader upwards, densely pubescent.—Plate 1, A.
Described from one specimen which we found in damp evergreen forest.
Seems to be very rare —No. 25802.
522 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXX11
15. SPIRANTHES, Rich.
1. Spiranthes australis. Lindl. in Bot. Reg. t. 823.
Fischer thinks this must be asport of S. austvalis, Lindl. = S. sinensis, Ames,
Orchid. ii (1908), 53.
The flowers are polymorphous. The top flower is larger, sepals and petals
obtuse, sepals minutely serrulate. Rostellum mucronate at tip. Rostellum of
lower flowers smaller, minutely bifid at apex.—Flowers white, tinged with
pink.
Found near top of plateau.--No. 26528.
Distribution: Throughout India, Afghanistan, N. Asia, China, Java,
Australia, N. Zealand.
16. ZEUXINE, Lindl.
1. Zeuxine longilabris, Benth. in Gen. PI. iii, 600.
Differs from other known specimens in the colour of the flower: Sepals olive
with white margins. Lateral petals and lip white with olive base. Column
orange. Bractpinkish. Ovary olive with 3 lighter stripes. No. 26530.
2. Zeuxine Biatteri, C. E. C. Fischer in Kew Bull, 1927.
Sepals greenish with pale tips. Petals pale-pink. Sac orange. Limb
white.
Found in damp evergreen forest.—No. 343.
17. “PERISTYLUS; Bi:
1. Perristylus stenostachyus, Krzl.—Habenaria stenostachya, Benth. FI.
Hongk. 362.— Habenaria peristyloides, Wight Ic. t. 1702.—No. 26507.
Distribution: Sikkim Himalaya, Khasia Hills, Tenasserim;, Deccan
Peninsula, from the Konkan southward, China.
18. SATYvRIUM, Swartz.
1. Satyrium nepalense, Don Prodr. 26.
We found only one dry specimen in bare patches of the hill-top.—No. 344.
Distribution : Temperate Himalaya, from Kashmir to Sikkim, Khasia
Hills, Deccan Peninsuia, from the Konkan to Travancore, Ceylon.
19. DISPERIS, Swartz.
1. Disperis neilgherrensis, Wight Ic. t. 1719.—D. zeylanica, Trin. Cat. Pl.
Ceyl. 91.
ne many points in the morphology of this species are not, or only partially,
known, I give a full description of our specimen :
A small plant, 7-11 cm. high, all glabrous. Tuber entire,2 by lcm. Stem
rather stout, fleshy. Leaf solitary, very rarely 2, situated about 3 up thestem,
broadly ovate, acute, sessile, cordate, subentire, 5-7-nerved, amplexicaul.
Flowers few, subsessile, each with a leaf-like bract. Pedicel and ovary up to
15 mm. long. Dorsal sepal narowly lanceolate, 3-nerved, rather obtuse, forming
together with the petals a concave hood lcm. long and as broad, only glued
together, not united. Lateral sepals spreading, broadly falcately ovate, united
for about 3mm., 10 by5mm., subacute, with 5 nerves from the base 1, the
central one more prominent, each with asmallsaccate spur about 3mm. from
the base. Petals broadly semilunate, subobtuse, with several anastomosing
nerves and an oblique band marking the position of the dorsal sepal, margin
minutely papillose, slightly waved, Sepals and petals white at base, rest
greenish, veins purple. Lip : midlobe S-lobed. Claw erect, confluent with
column. Midlobe pendulous, resting on the claw, subacute, in the centre with
a short, laterally compressed, obtuse, oblong spur which is slightly tomentose.
The lip is folded along the side-lobes which, therefore, are channelled below,
they are strongly curved downwards and papillose and yellow-tomentose above,
united with the pendulous portion for some distance; the lower part of
midlobe (before meeting its side-lobes) lies along the large rostellum, but is
NEW SPECIES FROM HIGH WAVY MOUNTAINS O73
not united withit; the side-lobes surmount the column. Lateral lobes of lip
very short, spreading. Column resting on and united with claw of lip.
horizontal and having 2 papillose appendages at the base. Stigma consisting
of 2 small fleshy processes just above the place where the lateral lobes of the lip
are situated. Anther posticous, erect, attached below to the conniving
column and claw ; cells close together above, but distinct, with a short process
below, appear to be open trom the beginning. In front of anther and en-
closing its margins lies the large rostellum ; itis emarginate at the tip,
withering downwards with recurved margins and has at the base Z hyaline
flat appendages along which the caudicles of the pollinia rest and with a large
flat deltoid exposed gland at its apex. Caudicles not twisted, bright yellow ;
pollen-grains olive: green, 2-seriate, secund on the twisted axis, cuneiform, com-
pressed. Ovary up tol4mm. long, 6-ribbed, slightly twisted.—Plate 1. B.
Found among damp leaves in evergreen forest, not common. No. 345,
All the specimens mentioned in this paper are at Kew.
AN ACCOUNT OF THE PEARL FISHERIES OF TUTICORIN
MARCH AND APRIL 1927
BY
P.eRe AWATI, 1.4:S,
Professor of Zoology,
Royal Institute of Science, Bombay
(With four plates)
This pearl fishery was held, under the directorship of Dr. B. Sundara Raj,
M.A., Ph.D., Director of Fisheries to the Government of Madras, in the months
of March and April, 1927. The Government of Bombay had permitted me to go
to Tuticorin to observe and study the different processes involved in carrying out
a pearl fishery.
There have been many fisheries in previous years but never beiore was a fishery
held on such a grand scale as the present one as regards the extent and the
excellent organization of the camp and magnificent operatious at sea.
The story of the pearl fishery will be best understood if it is described under
three different aspects, the first of them dealing in outline with the life history
and the habits of the pear! oyster, the second with the formation of pearls and the
last describing in minute details the different operations of the fishery.
I. Pearl Oyster.—The celebrated pear! oyster of ‘Tuticorin, known as
Margaritifera vulgaris, belongs to the family of Aziculide having close
affinities with Scallops (/Pecten) and Mussels. It differs from Edibie oysters in
the possession of bundles of tough threads (byssus) by means of which it anchors
itself to the substratum.
Some of the general characters of the family areas follows :—
‘ The shell is usually inequivalve; the dorsal margin straight, often very
long and forming anterior and posterior wings or ears; the lateral teeth of the
hinge line are much prolonged and may be inconspicuous ; the mantle lobes are
not fused ; siphons are absent; the foot is moderately long, tongue-shaped with
a well developed byssus gland ; the posterior adductor muscle is very large and
nearly central, the anterior adductor muscle absent.’
The internal characters need not be mentioned here.
Life History and Habits.—The Pearl oyster is unisexual and remains so till
the end of. its life unlike what has been observed in the edible oyster which
changes its sex after each spawning period. It becomes sexually mature within
the first two years of its life and spawns twice a year in the beginning of the
South-west and North-east Monsoons. Its fecundity is enormous. Eggs are
extruded direct from the oviducts by means of the suprabranchial passage
to the sea and are fertilized in the open sea by spermatozoa emitted by ripe
males.
Eges undergoing development after fertilization become swimming embryos
within a very short time. The latter pass through different stages such as
trochosphere, veliger, etc., and are transformed within a week into spats, 1.e.,
very tiny oysters with syimnmetrical shells and delicate byssus threads. Before
the spat formation, the larvae orembryos are at the mercy of ocean drifts and
currents which carry them in all directions. As soon as the spat is
formed, it gets itself attached by means of the delicate byssus threads to any
floating or rooted algze. It seeks the shady under side of the branches and
sooner or later drops off and sinks to the bottom being dislodged by shaking or
by the disintegration of the floating weed. Once it is on the bottom it begins to
grow in size, all the time being anchored firmly to the substratum by means of
those byssus threads.
AN ACCOUNT OF THE PEARL FISHERIES OF TUTICORIN 525
Though the anchorage is firm, the Pearl oyster is by no means a sedentary and
helpless animal. It creeps slowly along thesurface by means of its long and
tongue-shaped foot when the byssus threads are damaged or uprooted. This
oyster has got the power .of forming over and over again byssus threads which
are secreted by a certain gland situated in the foot.
The shell of this oyster continues to grow in length till the third or the fourth
year of its life. After this period it thickens. greatly by the deposit of nacreous
material in its interior and especially in the neighbourhood of the hinge at the
dorsal edge. This is the time of rapid pearl formation. The oyster tissues are
then in the condition which lends to the secretion and deposition of limy material
either externally by addition to the thickness of the shell or internally as
successive coats deposited on any particle, such as a dead parasite, sand grain,
diatom, a fragment of nacre, ova or excreta, which has given rise to the necessary
stimulation. At about this time or a little later the oyster becomes fishable when
it measures about 4 by 3 inches. If it is not fished at the proper time (which is a
mysterious period) the oyster has got a tendency to disappear. Causes of its
disappearance are not yet discovered.
Longevity of the oyster is not definitely known but it appears that it does not
I've beyond six or seven years.
Banks or Paars.—The grounds on which these oysters are found aggregated
close together are known as Pearl oyster banks or paars (meaning rock or any
form of hard bottom in distinction to ‘manul’ which indicates loose or soft
sand). ‘‘ The Pearl bank plateau is bounded for the most part by 10 or 12
fathom line. Outside the paars, the ground shelves away rapidly to 20 to 30
fathoms which marks the top of the steep slope down to the mud-floored abyss
forming the central portion of the Gulf of Manaar. Along the line of this slope,
soundiness in some places jump in short distances to anything between 100 and
1,000 fathoms. Within 10 or 12 fathom line, onthe other hand, the ground is
for the most part very level.’’ At Tuticorin a distance of 10 to 12 miles is run
from shore before a depth of 10 fathoms is reached.
The surface of these plateaus is either sandy diversified here and there by
outcrops of so-called rocks generally in the form of flat or slightly inelined
ledges or low terraces, or entirely rocky with a mount of sand in crevices. The
rock in the latter case may sometime be covered by a thin layer of sand which
shifts from time to time with the currents.
The rock of the plateau inay be grit stone formed of yellow tinted quartz sand
cemented by carbonate of lime or it may be largely dead, worn and altered coral
or it may be a shelly mass, an agglomerate of dead valves of pearl oysters and
other molluscs.
‘A certain amount of sandy bottom interspersed with harder tracts and with
plenty of cultch, natural or artificiali.e., fragments of dead coral and of
calcretes, broken shells lying on the sand is probably the most favourable ground.
The cultch gives foothold for new broods of oysters and at the same time helps
to stiffen the shifting sand and prevents in some degree the wash out and
turnover of the surface which is caused by a heavy swell.’
These banks abound in rich flora and fauna besides the pearl oysters. The
microscopic spores and embryos of these plants and animals form the substantial
food of these oysters.
Enemies of the Pearl Oyster.—The Pearl oyster is a very delicate animal. Though
it is found in shallow waters up to 10 to 12 fathom line within as inany miles
from the shore, the sea water in which it is found must be clean i.e., free from
any sandy suspensions and also free from any dilutations of fresh water from the
land. These two factors alone will kill the oyster.
Besides the above there are other causes of death of these oysters ;
(1) Shifting of Sand.—This shifting which buries oysters and kills them is
caused by strong currents set up by the South-west monsoons and other storms.
(2) Predacious Fish.—Sharks, Rays and File fishes devour these oysters.
Though enemies, they form a necessary link in formation of pearls in oysters.
This connection will be explained later on.
(3) Boring Animals.—Boring sponges and boring molluscs, bore holes
into the shells. The former allow parasites and sand grains to enter through the
holes, into the soft body of the oyster and thus kill it indirectly, while the latter
suck the soft parts through these holes and swallow them.
(4) Miscellaneous causes such as overcrowding, over-fishing and epidemic
due to Protozoan or other parasites,
17
526 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Il. Pearls.—In India there is a popular belief that pearls are formed by
drops of rain falling at a particular period known as Swa/i. At this time it is
supposed that pearl oysters come up to the surface of the sea water, open their
shells and wait for the drops. As soon as these drops have fallen into them, the
shells are closed and the animals go to the bottom. ‘These drops become
afterwards pearls. Such and similar beliefs are still held by different communi-
ties all over the world.
Scientifically speaking, there are three important methods in the formation of
pearls ;
(1) Grains of sand, minute fragments of nacre, etc.
(2) Pathological secretions.
(3) Cestode embryos.
any one of the above entering into the body of the oyster causes irritation and
then stimulation to deposit pearly nacre around the source of irritation.
This nacre is deposited by an epithelial layer derived from the ectoderm (skin).
Almost all the fine pearls found free in the body are formed by deposition of
nacre around the cestode embryos which find entrance into the oyster. It will
be very interesting reading if a complete life cycle of this pearl-stimulating
cestode is described here.
‘The adult worm (7etrarhynchus untontfactor) from the body of a Shark or
Ray sets free its nunierous young embryos into the sea. Such free swimming
embryos gain entrance with food currents into the body of the oyster. They are
found in the mantle, in the gills, in the liver and also in the connective tissue of
the body wall in an encysted or encapsuled condition. If these infested oysters
are eaten by File fish (Balistes) Sharks or Rays, the cestode larve are transierred
to the body of the new host where they begin to grow. These cestodes are then
in arecognizable stage of Zetrarhynchus but they are not yet adult nor have they
reproductive organs. It is not yet definitely determined whether these immature
cestode forms reach their maturity in the same fish or in another new host, i.e.,
Shark or Ray which preys on the File fish.’
‘In those oysters which do not become the prey of the File fish, the cestode
larvae have no chance of completing their life cycle. They die and become
encapsuled in the layers of pearl nacre deposited by the living tissues of the
oyster upon the source of irritation caused by the cestode embryo which ulti-
mately becomes the centre of a brilliant and costly pearl.’
Thus it will be seen from the above that the predaceous fish form a neceSsary
link in formation of pearls. ‘ No rays, no pearls ; abundance of rays no oysters,
as the latter are destroyed by the former. But Nature has somehow restored
the balance between the oysters and the predaceous fish.’?
Different views as to pearl-formation can be best summed up in Prof.
Herdman’s words:—
(1) Majority of pearly excrescences on the interior of the shell are due to
the irritation caused by boring animals. In exceptional cases a free pearl may
be formed in this way. ‘There is a device by means of which these blisters or
excrescences are cut and filed into pearls of very low order. These pearls are
then known as Vottu or Blister pearl.
(2) Minute grains of sand or inorganic particles form the nucleus of pearls
under exceptional circumstances. When the shell is damaged, these sand
grains get into the interior and supply the irritation that gives rise to pearl
formation. ‘These are known as Ampullar pearls.
(3) Many pearls are found in the muscles close to surface. These are formed
around minute calcareous concretions, the Calcosperules, which are produced
in the tissues and form centres of irritation. These are known as Muscle pearls.
(4) Most of the fine pearls found free in the body contain the remains of
the cestode parasites (embryos). ‘They supply the requisite irritation for pearl
formation. ‘These are known as Cyst pearls.
Valuation of Pearls.—Valuation of pearls is a very complicated affair. The
principle underlying this valuation is to pass pearls through different sieves.
Each sieve is round about 6 to 8 inches in diameter and 2 to 3 inches high with
sized holes in the bottom. The size of the holes in each basket (sieve) is
different. The pearls that do not pass through the holes of any particular
sieve have a different value and different name. In this manner different pearls
Journ., Bombay Nat. Hist Soc.
ROO ame
POLICE
PEARL BAZAAR GUARDED BY
i
ARL FY
PE
)
E GENERAL BAZAAR
NER OF TH
A CoR
14s
Journ., Bombay
Nat. Hist. Soc.
5. DIVERS’ ENCAMPMENT.
4, DIVERS AT WORK.
AN ACCOUNT OF THE PEARL FISHERIES OF TUTICORIN Sei
Names of pearls depending on valuation, size, shape and lustre are different
in different parts of India. If mentioned here, it will cause more confusion and
will serve little or no purpose at all in this narration.*
III, Pear! Fishery.x—A pearl fishery involves many operations, some at sea
and others on land, but long before it actually begins, the banks or paars have
been surveyed many times in order to investigate spatfalls as well as numbers
of fishable oysters. If the survey results as to the quantity of fishable oysters,
and if the valuation reports of pearls contained therein are satisfactory, it is
then and then only decided to hold a fishery. In order to make it known to all
concerned, it is thoroughly advertised in Burma, Ceylon and other parts of India
and the world through various newspapers, so that rich pearl merchants will be
attracted to the place. In the meanwhile divers are being recruited, fishing
boats are being brought to the place and the camp is being constructed on the
spot where the fishery will be held.
Fishery Camp at Silavathurat and tts Admintstration.—The site of this
pearl fishery camp known as Silavathurai lies at a distance of nearly two and
half miles to the north of the town of Tuticorin. ‘It is the time-honoured spot
where fisheries from ancient Hindu and Portuguese days have been held.’?
The camp covers an area of more than 100 acres. The front i.e. the eastern
portion of the camp extends up to the open beach, while the rear (western)
portion lies amidst Palmyra trees. On the south side it is bounded by a serics
of small lagoons which separate the town from the camp, while tothe north lies
a piece of uncultivated land full of thorny bushes and cactus, a small back-
water channel and a salt factory, a mile orso away from the camp.
On the beach are built huts or quarters for officers, visitors and the staff. A
little towards tte south, but facing the beach, are constructed offices including
an auction hall in front of the old beacon. In the same line parallel with the
beach but far away towards the south is built a Kottu (store) for Government
shells, and still further away on the beach itself for nearly a mile are located
series of huts known as Thotties for rotting oysters.
. Just behind the offices is reserved an area for retail selling and cutting open
oysters by retail buyers. The area known as Sales Ground is fenced with wire
all round except at two opposite ends which are kept open for admission into
and exit from it.
The pear] bazaar extends on both sides of the road which runs east to west
from the Sales Ground. This bazaar consists of pear} shops opened by pearl
merchants coming here from different places but mostly from Bombay. These
shops are rented out to them and Government receive a good income out of it.
A road which runs at right angles to the west end of the pearl bazaar accom-
modates shops of green grocers and eating houses for various kinds of people
who frequent the camp. ‘There are also some cloth shops. These shops are
also rented out.
There are twoor three roads running east to west at right angles to the mid
portion ot the general bazaar on either side of which are built huts for divers,
different sects occupying different areas. These huts are given free to these divers.
Rehind all the above roads but along way off towards the west are sunk
e'ght tube wells which supply water to the whole camp. ‘They are worked
with a hand pump. Water is supplied in a bullock cart to the officers and the
staff, but other people have to take water from the pump which is worked by
Government coolies. A guard is placed at each tube well to see that no bathing,
washing or polluting the water in any way is done. The water arrangements are
in charge of a mechanic lent for the purpose by the Industrial Department of
Madras.
In the north-west corner, but away from the tube wells and also from the
camp, there are bathing places for divers and others. A contractor who is in
charge of these places collects a small fee for every bath. One can see a crowd
of bathers every evening when divers have returned ashore all tired and
wanting a cold bath to refresh themselves.
The whole camp is well guarded by constables who are in charge of one
Inspector of Police and his two assistants. Its health is also well looked after
1 The above account of the pearl oyster and the formation of pearls is
mainly based on Prof. Herdman’s Reports to the Government of Ceylon,
2 Report of the Pearl Fishery, Tuticorin, Madras, 1926-27,
528 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXI1
by Medical and Sanitary authorities. There is a small hospital with an out-
door dispensary attached to it. .
One Additional Superintendent who is recruited from the Revenue Depart-
ment looks after administration of the camp. He goes round the whole camp
twice a day morning and evening, supervising the general comfort.
The lighting is done by the department itself and it is in charge of the above
officer. The roads are well lit with Kitson lights.
Peaceful and harmonious working of the whole camp reflects a great capacity
for organization and tact on the part of the Director of Fisheries.
Operations up to Diving.—Nearly one thousand skin divers took part in
this fishery. There were Christian divers, mostly local i.e., from Tuticorin,
there were Mohamadan divers from a place called Kilakarai, now famous for
efficient divers, and there was also in addition to the above, a sprinkling of Arab
divers once famous for diving. These divers not possessing a boat of their
own, had to hire a fishing boat to take them out tothe pearl banks and bring
them back ashore after the day’s work. Every boat taking part in a fishery
has to obtain first a pass from the Customs Department which (pass) is handed
back to the same authorities after the fishery is over. These divers have also
to hire men for lowering them into the sea and haul them and their baskets up
during the period of diving. These men, known as mandacks, are really the
crew of the boats doing additional work and earning extra money.
Nearly seventy-eight boats were engaged in this fishery.
Before the actual start from the shore which was at about four o’clock in the
morning, the tindal of every fishing boat had to obtain a fresh permit every night
from the Beach Master as to the number of divers and mandacks his boat was
carrying to the place of fishing. The permits were made in different colours and
boats were accordingly divided into several groups, those of each group having a
permit of the same colour. ‘The purpose of this device will be clearly understood
when a certain operation i.e., the division of oysters at sea, is described
later on.
Every night in the camp a gun was fired at 11 p.mi., a second gun at 3a.m.
and the last at 4 a.m. when boats and divers were supposed to be ready for
departure from the shore. When everything was ready, the boats weighed anchor
and set sail for the place of fishing. If the winds were unfavourable, these boats
were generally towed either by the Government Trawler Lady Goschen or a
hired tug, for a distance of 10 or 12 miles to the places on the pearl banks already
marked out for fishing by the Superintendent in charge of the Pearl Banks. Once
the fishing boats arrived on the spot, they cast their anchors and the divers got
ready for diving. By this time there was sufficient light for diving purposes.
OPERATIONS FROM DIVING TO RETURNING OF DIVERS ASHORE
Complement of each boat.--In each beat there were two classes of men directly
concerned with diving, one of them being divers 10 to 12in number and the
other, as many mandacks. Besides,the above, there was the tindal of the boat
with six to twelve men as the crew according to the size of the boat. Sometimes
the mandacks themselves constitute in number the crew.
Methods of Diving.—There appeared to be no standardized method of diving.
‘The Arab divers had poles projecting over the sides of their boats by means of
which they hoisted themselves out of the water by an acrobatic movement. They
usually used horn clips on the nose and it was their custom to come to the
surface hanging on to the rope with the left hand and not letting it go in mid
water until they reached the surface.
‘ Most of the Tamil boats have temporary outriggers fitted to the sides of the
boats from which the divers are suspended. The Tamil divers do not wear
horn clips on their nose, but instead the nose is held between the thumb and
the fore finger during the rapid descent to the bottom. They let go the rope
in mid water after they have received an initial pull sufficient to bring them to
the surface. ‘The disadvantage of this method is that these divers reach the
surface some yards away from their boats and also spend more energy in
coming up.
‘ With rare exceptions both kinds of divers go down feet first with their feet
resting on the stone sinker. A few of them are seen to hold the sinker near
the chest and dive in head first. ’?
z Report of the Ci eylon “Pearl Fishery, 1 925-26.
Journ., Bombay Nat. Hist. Soc.
SPOILS AT SEA
DIVISION OF
a
BAGS OF Oy
(Each bag contains 500 oysters.)
6
Journ., Bombay Nat. Hist. Soc.
<n
7. THE RETURN OF THE DIVERS.
8. WASHING THE PEARL OYSTERS.
AN ACCOUNT OF THE PEARL FISHERIES OF TUTICORIN 329
Every diver carries two ropes with him to the bottom one of them attached to
the stone sinker and the other to the basket in which oysters picked from the
bottom are raised to the surface.
Both these ropes are hauled by the mandack. As soon as the diver reaches
the bottom, the stone sinker is let loose which is immediately hauled up
to the surface. In the meanwhile the diver is feverishly busy in picking up the
oysters or whatever comes to his hands and collecting them in the basket. He
pulls at the basket rope whenever he wishes to rise to the surface for fresh air.
Immediately the mandack begins to haul up the basket with the diver and
within a few seconds both of them are brought to the surtace.
After some rest the diver commences his work again.
Diving was done till a signal was given in the shape of firing of a gun ora
shrill siren from a trawler when diving from all the boats had to cease at once.
This took place at about 1 p.m.
In the meanwhile the Superintendent of the Pearl Banks was very busy in
supervising the fishing boats. If the latter had strayed out of their places, they
were towed back to their former or sometimes to new places. It is an advantage
that all boats should work together on a particular spot. He also saw to it that
diving ceased at once after the signal was given and that there was no illegal
cutting of oysters before the division was made.
Division of Oysters at Sea.—As soon as diving ceased, boats began to app-
roach, one by one, the Government schooners which were anchored on the fishing
ground. Each side of the deck of these schooners flew a flag of a particular
colour. Only those fishing boats approached that particular side which had
permits of the colour of the flag fying on thatside. This device saves a lot of
worry and labour and provides a check on the boats and the divers.
On each side of the deck were placed two officers one for making divisions of
the oysters brought by the divers and the other for checking and sealing the
divers’ shares in gunny bags. ‘There were nearly half a dozen menials to help
these officers.
As the boat came alongside the schooner, every diver, one after another,
handed his collection to the officer who madethree equal divisions, giving
one to the diver and appropriating the remaining twoon behalf of Govern-
ment. In the meanwhile the checker put the diver’s sharein a gunny bag,
sealed it and handed it over to the particular diver who went back to his boat.
When all the divers got back their shares and the tindal received his old permit
filled in with new details and countersigned by these officers, that boat was ready
to go back toshore. Other boats were dealt within a similar way. They were
all ready now to go back, ‘They reached the shore from two to three hours in
fair weather from the time they got back their permits.
Sometimes it was very difficult to make divisions at sea owing to the high
winds and choppyseas. Under these circumstances divisions were made on
land after the boats had gone back under escort and deposited their collections
in the Government Kottu or store.
The fishing boats arrived ashore in groups or one by one as the case may
be and there was usually a huge crowd on the beach to welcome them.
The staff of the Beach Master was very busy on the beach when the boats
arrived. ‘They were scrutinizing the sealed bags of divers, getting back the
permits already issued to them overnight and searching the boats for any mis-
appropriated oysters. When they were satisfied, they allowed the divers, the
mandacks, the crew and the tindal to go ashore.
Retail Selling and Retail Buying .—As already stated above, every diver has
to pay the tindal and the mandack, the former for the hire of the boat and the
latter for the services rendered at sea. He paid both the people in kind. He
made six equal divisions of his share, retaining four of them and giving the
remaining two to the tindal and the mandacks respectively.
These people now retired to the sales ground to sell their oysters. Retail
buyers could buy them from one oyster upwards. The average retail price
varied from twenty to forty oysters toa rupee. Onan average a diver made
about ten to twenty rupees a day.
A retail buyer cut open his oysters in another part of the same ground,
anxious to see what his oysters contained.
In the meanwhile the Government transport boats arrived with the
(sovernment share which was handed over tothe Kottu manager. These oysters
had been already counted and bagged, five hundred in each bag, at sea. They
530 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol: XXXII
remained inthe Kottu till they were delivered next day to those who had
bought them at the public auction.
Public Auction.—A public auction was held every night at 8 or 9 p.m. in the
auction hall. Half an hour or so before the auction began, a gun was fired
to apprise prospective bidders of the event.
The auction hall was built on the central portion of a raised platform. Inside
the hall were seated the Director and his staff, while in the open the public
waited for the auction to begin.
The Director sat by himself on a raised platform, while his staff consisting
of voucher makers, checkers and treasury clerks satin front of him facing the
public. There was a bamboo railing separating the public from the staff.
Atthe appointed hour the auction began, nobody was allowed to bid for
Jess than one thousand or more than twenty-five thousands oysters. One
of the peons standing in front of the staff, facing the public, announced the
numbers of the oysters Government had for sale that particular day. Onan
average it amounted to one Jakh and fifty thousand oysters daily.
Bidding per thousand went on till a certian price became stable for some
time when some bids were accepted at that price. The price usually ranged
between rupees thirty to fifty per thousand. Those whose bids were accepted
at that price were admitted one after another into the hall where every one of
them was required to get a voucher containing his name and the number of
oysters he had bought. He presented this voucher to a treasury clerk who wrote
the same details in his ledger and received a deposit, generally 1/10 of the whole
amount of the purchase and gave a receipt for the same.
Bidding went on in the manner described above, till all the oysters of that
day were sold out. :
Next day the buyers presented their deposit receipts to the cashier in the
treasury and got receipts for full payments which they presented to the Kottu
manager. The latter gave them delivery of their goods.
On an average Government got a net income of Rs. 5,000 to Rs. 6,000 per
day. But this income depended on various factors. Fair weather, calm sea,
favourable winds, no under currents, warm day so that divers found it comforta-
ble to dive, presence of efficient divers, place for fishing on the banks rightly
located, pearls properly valued, presence of bidders and pearl merchants, good
sanitation in the camp, absence of strikes, etc., all these conditions, indi-
vidually as well as collectively increased the income or decreased it if any of them
was adverse. Presence or absence ofthese conditions makes or mars pear] fisheries.
Rotting of Oysters in Thotties.—After the delivery of the goods, the buyers
took their loads of oysters to their respective Thotties. Use of these Thotties
was auctioned and Government got a good income out of it.
These hotties were located on the beach far away from the camp, since they
might become the source of an epidemic like Cholera as the oysters were being
rotted in them. ‘The four walls of each thotty were constructed of bamboo
thatties with asmall entrance but it wasopen tothe sky. Each hut or thotty
measured about 30 X 30 X 8 ft.
In these thotties then, the oysters were dumped in heaps for rotting purposes.
Various agencies worked together for macerating or rotting the soft parts of these
oysters. Heat engendered by the heap produces the optimum conditions under
which various kinds of flies, mostly Calliphora, Lucilia and Sarcophaga, lay
their eggs or larvee. The latter which are voracious feeders of offal complete,
with the help of bacteria, the maceration within four or five days. The soft
parts are now turned into homogeneously pulpy material which emits vicious
and repulsive odour.
After the fifth day these oysters or whatever remains of them were washed ina
small dug-out or a tin bath or in a still smaller vessel according to the number of
the oysters to be washed. After stirring or churning thecontents and throwing
away empty shells, the container was filled to the brim with sea water and kept
over night. The next morning the superboyant liquid was slowly decanted and
the contents again washed. This process was repeated so many times that
a fine powered stuff was left behind. It was then spread out on a black piece of
cloth and left there for drying. After it was thoroughly dried,.a vigorous
search for pearls was made on the spot or the whole stuff was taken home or to
a shop where thorough search was made at leisure.
Pearl Bazaar,—There were sixty to seventy pearl shops or huts located on
both sides of the main road.» ‘These shops were rented out to pearl merchants.
AN ACCOUNT OF THE PEARL FISHERIES OF TUTICORIN 531
In these shops pearls were sold or bought or merely exhibited for show. In
some of the shops fine holes were driiled into pearls which were then sewn into
threads and were exhibited for sale. There were also people in this bazaar who
bought blisters of the pearl shells and made cheap pearls out of them by
careful filing.
Lastly my cordial thanks are due to the Director of Fisheries and his whole
staff of the Pearl Fishery for the courtesy shown to me and for the trouble they
underwent in taking me over all the operations involved in this Fishery.
[ also thank the Government of Bombay which allowed me to study this
Fishery on the spot.
MODERN MUSEUM METHODS
BY
S. dle) PRATER, C.M:Z.S.
Curator, Bombay Natural History Society
Part |
(With six plates and five text figures)
Since one of the main objects for which a Museum is established
is the ‘ diffusion of instruction and rational entertainment among
the mass of the people’, it follows that, apart from the purely
scientific research and investigation which it carries out, the collec-
tions which are open to the public in its galleries, however rich,
important and complete they may be, are only of real value to the
extent to which they are made use of by the public and to the extent
to which the public is able to benefit by the instruction and enter-
tainment they are intended to afford.
Now the attitude of the public to the museum might often be ex-
pressed in the words of a certain New England farmer who, after
enumerating the many virtues of his deceased wife, concluded by
saying ‘somehow I never liked the woman’. The public realizes
more or less that the museum is an institution intended for its
benefit but somehow does not like it. May not the reason for this
attitude be explained by the fact that many museums are de-
humanized institutions, completely lacking in attractiveness because
of their unsympathetic treatment of the public and their unsym-
pathetic methods of presenting their treasures ?
Shortly after arriving in New York, I read an article in one of the
Sunday papers dealing mainly with the work the Museums were
doing for the people of the country. ‘ In America to-day,’ the writer
pointed out, ‘ the Museum fills the role of educator, entertainer and
nursery companion ; it has ceased to be a dusty warehouse of
antiquities and has become a bright and cheerful instruction centre
for the community.’ After a few months spent working in and
visiting many of the Museums in the country I realized more clearly
the truth of his contention. Like everything else in American life,
the Museum has developed into an institution with the motto
‘service to the public’. The national spirit is expressing itself in
these rapidly growing institutions—-efficiency, gaiety, and democracy
—these characteristics already differentiate the museums of America
from those of other lands. One finds efficiency in their careful study
of the comfort of the visitor while in the museum galleries and in
their almost perfect methods of charting, casing, and labelling. One
finds vividness and charm in their methods of exhibition, and
democracy in the wide appeal they make to the public at large. Art
and Science are learning to work together. The Science Museum
calls on the artist for ‘atmospheric effects of nature’ and the Art
Museum looks to the scientist to arrange its treasures in some in-
telligible progression. The result is the creation of museums which
are really attractive to the masses, of museums which look to and
receive public support. The impression one receives is that these
MODERN MUSEUM METHODS 533
museums are unique in their history, progress and purpose; they
cannot be compared with kindred institutions in the old world.
Their appeal is not so much to the advanced technician or craftsman
as to the man in the street. Except in the case where qualified
guides conduct parties of visitors over its galleries, the teaching of
the museum is silent teaching, and to be successful this teaching
demands aneffort on the part of the visitor. How far is this effort
forthcoming? Now one might say that 90 per cent. of the people who
visit a museum visit it mainly for pleasure and with the majority
the message, the museum has to offer, loses much of its
meaning unless it can provide something which arouses and holds
interest. To deluge people with facts and to offer too much to
their attention results in getting very little of it. The story of the
monkey who was unable to get his handful of nuts out ofa jar
until he had dropped some of them has its application here. The
neck of the jar is as large as the public interest, no larger—to grab
too much attention is to lose it. To get the public to look at and
absorb what you offer you must drop some of the nuts. One cannot
teach people by marshalling whole series of facts before them. It
is the principles that matter ; facts will soon be forgotten while the
principles remain. In offering the visitor principles which are new
and unknown to him it is necessary to commence with something
that is familiar and known to him. Our understanding and accept-
ance of a thing depends much on what we already have in our con-
sciousness. Introduce something which has no connection with our
past experience and we are not interested. To hold our interest
there must be a large ingredient of the familiar with the unfamiliar.
Edward Fisher in speaking of museums and their work said, ‘ The
great purpose of Museums is to stimulate the observant powers into
action. ‘The educational value of museums will be in exact pro-
portion to their powers of creating new thoughts in the mind’.
The object of the museums is therefore to lead the people to
better ideals and to offer them new view-points in a way which
they can understand and appreciate. The appeal is made more to
the senses than to the intellect but it is made in such a manner that
in addition to mere attractiveness it offers food for thought which
is the first step to a higher and more intellectual ideal.
There are many ways of telling the story of nature. Why not
choose the beautiful way ? The way which leads from an apprecia-
tion of its interest and its beauty to an understanding of its mean-
ing. It is the method which will make the widest appeal. A second
reason is the obtaining of public support. By giving the public
something which it can understand and appreciate and something
which it wants, the museums succeed in rousing its interest and
ultimately its support. And in this the American Museums have
been completely successful.
A study of the methods adopted by museums in the United States
in serving their public would perhaps repay the Museums of other
countries. The stress and strain of modern life, and the demand
that it makes on the time and attention of the average wage-earner,
does not leave much time for such side issues as Art and Natural His-
tory. It is a condition that is becoming increasingly prevalent not
18
934. JOURNAL, BOMBAY NATURAL’ HIST. SOCIETY, Vol. 2a,
only in America but also in the older countries. Dr. Van Straelen,
the Director of the Natural History Museum, Brussels, with whom
I discussed this aspect of the question believed that the democratic
spirit of modern times necessitated a change inthe attitude of
museums. The old museums were aristocratic, their show galleries
were merely in the nature of a concession, but modern conditions
necessitated a change in this policy. The museums were becoming
more and more dependent on public support, on money voted for
them by the people’s representatives, and unless museums did some-
thing for the people the money would not be forthcoming.
‘Applied science will find its votaries and its supporters ’ continued
the Doctor, ‘but for pure science we will have increasing difficulty
in finding money. We arrange our galleries on the plan of a text
book where specimens, models and explanatory labels trace in out-
line the various classes of the animal kingdom. This type of -
exhibition was necessary and useful for the more serious students, but
the great majority of our visitors are not serious students and for
them we want something else. There must be some ‘ light houses’
in a gallery to draw their attention and rouse their interest.’
At the Natural History Museum, Berlin, radical changes are being
made in the character and nature of the exhibits. There, too, the
museum is making a bid for popular interest and support by the
introduction of methods of exhibition which appeal to the lay visi-
tor. For if the institutions which stand for the cultural develop-
ment of the people are to succeed in their mission, it has become
necessary for them to alter their standards, to break down their
aloofness and restraint—that attitude of mind which looks on the
visiting public as something in the nature of a nuisance, and come
down to a more sympathetic consideration of the people’s wants.
Some may object. Why appeaito peoples’ wants? ‘Their wants
may be stupid, childish or even positively mischievous. What we
must appeal to is their reason. Professor Overstreet in his interest-
ing series of lectures on ‘ The Influencing of Human Behaviour’
gives us the answer ‘No appeal to reason can ever be successful
if it is not also an appeal to awant’. An understanding and
appreciation of the truth of this axiom will save much time in
argument. If one of the main motives of a Museum is education
not of the few Lut of the mass, and if this education is to be effective,
it requires the rousing of the popular interest in its activities as a
first step, and it is the duty of the Museum Director to make his
museum worth being interested in, to give his museum such vitality
and pertinence that people will want to be interested in it, will want
to visit it, and will realize the necessity of supporting it, and though
the people may want something quite inferior, his problem is to get
them to want something worthwhile, and to give them the opportu-
nity for these more worthwhile wants to multiply and grow.
I will now pass to a consideration of the methods adopted by
Museums in the United States and on the Continent to attract and
maintain public interest and support. The subject may be divided
under two headings :
1. The Museum Galleries as a whole.
2, The nature and methods of arrangement of the exhibits.
MODERN MUSEUM METHODS 535
1. GENERAL TREATMENT OF GALLERIES
The principal factors which react on the mind of the visitor on
entering a gallery are—
1. The architecture and colour of the walls and the show cases,
2. The show cases themselves and their general arrangement,
3. The exhibits.
The third factor being the most important, it necessarily follows
that the first and second factor should be subordinate to the most
effective presentation of the third.
We will leave the question of architecture aside for the present.
The museum curator is rarely ina position to decide the architec-
tural requirements of the building he is to use. The question
of the colour of its walls is a matter which might easily come
within his province to decide. It is not difficult to realize the
importance of colour asa means of attraction. Even as chiidren
we were attracted rather by colour than by form. We retain this
impulse throughout life, only we temper it by a new desire fora
pleasing arrangement or harmony of colours. Our objective should
therefore be a pleasing colour scheme, that harmonious blending of
show cases and walls which will be most effective in the presentation
of the exhibits shown in the gallery.
Colour Schemes.—AS a general rule quiet colours should be
utilized. The choice of colour will depend largely on the lighting of
the room and the objects to be displayed, but the rule in its selec-
tion should be the avoidance of an obtrusive scheme which will
draw to itself the attention of the visitor. At the Brooklyn Museum,
the show cases and walls in the hall of Invertebrates were painteda
bluish-grey ; the effect produced was restfui and pleasing. In the
Mammal Gallery at the Natural History Museum, Altona, Hamburg,
the colouring of the walls and show cases was very harmoniously
conceived. In the main Mammal Hall the walls had a 6’ 6” dado of
dusky slate blue relieved with a faint grey stripe, the upper portion
of the wall was carried out in Rejane green‘! and the contrasting
areas of colour were divided by narrow black lines, the pedestals
of the show cases corresponded in colour to the dado, their upper
{framework was in biack, while the backgrounds were grey stippled
with green. In the Bird Gallery in the same museum the dado
of the hall was in nickel green, the upper surface of the wall
cream buff with a white area where the wall joined the ceiling. The
pedestals of the show cases were treated in a similar tone of green
as the dado, while their frames were black.
The selection of a colour scheme should be governed by the type
of exhibit shown in the gallery. Certain exhibits require quiet
tones, others more brilliant ones to give them their full significance,
particularly where colour in the exhibits themselves is wanting.
A rather daring and original colour scheme is to be seen in the
Ethnological Gallery at the Brooklyn Museum. This hall is divided
by its architecture into three sections, the columns supporting the
ceiling forming the divisions. Each section is confined to illustrat-
ing the arts and crafts of a distinct race. In the first section the
* Ridgeway’s Colour Standards, pl. xxxtii,
536 JOURNAL, BOMBAY NATURAL HIST, SOCTETY, Vol. Os
show cases were green, in the second royal blue and in the third
rose pink. The plinths of the adjoining pillars were coloured to
correspond with the show cases. Thus in the same hall one had
three aisles of show cases paintedin different colours. After passing
through several galleries where drab or neutral tones predominated,
this origina: colour scheme had the effect of renewing and tre-arous-
ing one’s attention. The eye was greeted by something fresh and
colourful and the effect was not unpleasing. Rejane green makes
an excellent background for animals and birds. It is used in the
galleries of the Munich Museum and is perhaps the most successful
colour tone I have seen used in any Museum in connection with
these types of exhibits.
Atmosphere.—The colouring of the walls and the show cases to
produce a desired ‘ atmosphere’ is another point which deserves
consideration. Inthe Invertebrate hall at the Brooklyn Museum,
where the majority of the exhibits were illustrative of life in the
sea, the blue grey colouring of the walls gave a marine note to the
gallery which was further accentuated by mural paintings depicting
seascapes in various quarters of the globe. Inthe new Fish Hall
at the American Museum, the colour of the walls is grey witha
stippling of waved blue lines giving them a watery effect; the back-
grounds of the cases against which the fishes are shown, had the
same colour motif in a slightly darker tone, the whole object being
to give a ‘marine atmosphere’ to the gailery. Artificial light
through lanterns of greenish glass increased this effect. The whole
question of the colour in a museum gallery is a phase of museum
work which requires study, it is now in the experimental stage.
It marks an advance in museum methods. Its object is to please
and attract the visitor by creating an atmosphere which will produce
a subtle reaction on his mind and in any way prepare him to enjoy
and appreciate what he sees.
Mural Paintings.—A further development is the decoration of the
walls with appropriate mural paintings. Many museums are now
adopting this method of decorating their galleries. We find
galleries of Art, Archeology, Geology, Ethnology, Zoology and
Botany decorated in this manner. Apart from their purely decora-
tive effect, murals have a distinct educational value. ‘They are a
clear and simple means of transmitting ideas—they are tools for
communicating thoughts, stimulating interest and arousing feeling
and emotions. Where arange of exhibits is shown which requires
a mental effort on the part of the visitor to clothe them with the
interest they merit, these pictures will supply the required stimulus
—will supply vividness and drama to what might otherwise have
been a barrentheme. In the hall of the Age of Man in the American
Museum, where the exhibits illustrate successive stages in the history
of man, the dry bone atmosphere was to a large extent relieved by
the beautiful murals adorning the walls of the gallery. The flesh-
less skull of the Cro-magnon man springs into life in the mural
painting depicting the Cro-magnon artist at work painting his pro-
cession of mammoths in the cavern of Font-du Gaume in France.
At Altona, Hamburg, the walls of the Bird Gallery were ornament-
ed with paintings of individual birds or groups of birds ; such subjects
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MODERN MUSEUM METHODS 03/
as geese in flight ora soaring eagle gave a charming and appropriate
setting to the exhibits. The vaulted ceiling of the Bird Gallery at
the American Museum of Natural History is painted to represent
a sunset sky against which we see various birds in flight— actual
specimens are used and as these recede further into the distance
they are substituted with painted birds, the gradation in size and
the perspective is cleverly maintained and ihe effect altogether is
realistic and charming. (Vl7de Plate 1).
There are certain points to be remembered in decorating the wails
of a gallery with murals. Firstly, a mural must preserve the flatness
of the wall it decorates. It should not mar the architectural lines
or destroy the even level of the wall’s surface by giving it an effect
of contours. Mr. Charles Knight, one of the most successful animal
painters in America told me that he preserved the desired flatness
in his murals and yet gave them an effect of roundness by painting
them always with the light coming up from behind, so that the objects
were more or less silhouetted against the background and yet had
a feeling of depth.
Then again as regards colouy tones, these should always be ruied
by the type of exhitit in the gallery. Where one is exhibiting
skeletons or fossils and such like colourless material, a certain lati-
tude in colouring is permissible but in a gallerv where the exhibits
are themselves colourful, restraint is necessary in the colour of the
murals so that a harmonious effect might be obtained. In every
case the murals should be a subordinate feature and should not
become the centre of attraction. In the old African hall at the
American Museum the murals illustrating scenes of African life were
done completely in sepia. In the new Reptile Gallery which
contaiis many beautiful and vividly coloured groups, the murals are
being painted in quiet shades. Small colour sketches of these
murals were first done to scale, subsequently a full sized copy was
done on cheap muslin, this painting was fitted to the walls to see
whether it harmonized or clashed with the exhibits ; certain colours
were then reduced and other heightened till the desired effect was
achieved. I have written at some length on this subject because I
believe that our museum offers an excellent field to our school of
mural painters in Bombay, it will afford them the opportunity of
showing their skill to the people of their own city and of having
their work exhibited where it will be seen by thousands. Natural
History offers them an endless range of subjects while the Archzolo-
gical and Art sections could also be made a venue for their skill.
Arrangement of Show Cases.—The second factor which draws the
attention of the visitor on entering a gallery are the show cases and
their arrangement. A gallery crowded with tall show cases whose
massive wood work and external finish makes them unduly conspicu-
ous carries with it a decidedly forbidding aspect—it creates a feeling
of heaviness and artificiality which can be thoroughly depressing, and
its effect upon the visitor is to repel rather than attract. The over-
crowding of the gallery with show cases and the cases with exhibits
appals the visitor who quickly realizes that he will never be able to
absorb a tenth part of what is offered him.
The arrangement of the show cases and the type of show cases
5388 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
to be used in a particular gallery will depend largely on the
architectural features of the room, its lighting and the nature of the
exhibits to be displayed, but the general attractiveness of the room
and the effect of its arrangement upon the visitor is a point always
to be seriously considered. The accompanying photograph
(Plate II) illustrates a model exhibition hall designed for the new
Buffalo Museum. It will be seen that the cases are Jet into the walls
of the room—they are of the ‘ built in’ type, their breadth varies
according to the nature of the exhibits to be displayed. In the
present instance the cases along the sides are 18’’ deep, those at
the head of the gallery which are intended for groups are 6 feet
deep. This is one method of arrangement which is very successful
and very attractive. It gives an air of spaciousness even to a small
room and is adaptable to any type of exhibit.
A second method of arrangement is seen in the new Reptile Hail
in the American Museum. The floor of the gallery is occupied by a
series of small metal-framed box cases none of them exceeding 4
feetin height, the walls on one side have a series of ‘built in’ groups
(vide Plate II1) while the opposite wall has a series of vertical cases
arranged in the form of alcoves. The use of only low cases
for the floor gallery is an advantage in that, there is nothing
to obstruct the view of the visitor from the gallery as a whole and
that atmosphere of cumbersomeness and artificiality produced by
an assemblage of tall show cases is largely obviated. The use of
the centre of the gallery for some artistic and dominating exhibit is
an effective method of adding to itsattractiveness. This is seen to
good effect in the design for the new hall of African Mammals at
the American Museum of Natural History, New York, where the
central exhibit will take the form of a magnificent group of African
Elephants mounted as a memorial to the late President Roosevelt.
In the Mammal! Gallery at the Altona Museum, the centre of the floor
is occupied by the skeleton of a horse mounted ona high pedestal,
the walls are lined with ‘ built in’ cases, while the main floor space
is occupied by low box cases illustrating life histories of mammals.
It has been shown that by introducing the ‘built in’ type of show
case, the necessity for large cases on the floor of a gallery, is largely
obviated. High show cases on the floor of the gallery obtrude on
the visitor and obstruct the view, while the use of the floor space for
low cases only gives airiness and spaciousness to the hall and
facilitates guarding. The arrangement of cases as alcoves while it is
effective in focussing the attention of the visitor on one part of the
gallery at a time, has the disadvantage of making the policing of a
gallery difficult. The use of a prominent central exhibit actsas a
‘light house’ to attract the visitor.
Show Cases.—In addition to the problem of attractiveness the
Museum Curator has another problem—a very important problem
—and that is the prevention of fatigue. The visitor sees the
museum galleries not only with his eyes but also with his protest-
ing feet. Visiting a museum takes time and while the visitor takes
his time it will be more difficult for his brain to take in new im-
pressions, if the rest of his bodily organism is crying out for atten-
tion. Benches with restful backs are a great necessity in a museum.
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MODERN MUSEUM METHODS 539
The supply of seating is a necessity which is quite often made
conditional on available space, but if the comfort of the visitor is to
be seriously considered, then the provision of seating accommoda-
tion must be made an essential feature in the planning of a gallery.
At the Altona Museum, Hamburg, I saw a space beneath a
window between two show cases effectively used for a_ seat.
The second source of fatigue is eye strain, caused firstly because
the objects shown are placed either too high above or too low
below the Jevel of the eyes, secondly because the exhibits are
placed too far from the eyes and thirdly because objects are placed
in long monotonous rows.
The structure and design of museum show cases in which
museum exhibits are shown are an important factor in helping to
remove many of these disadvantages.
The older type of show cases were characterized by their heavy
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Fic. 1. WINDOW SEAT FOR VISITORS, ALTONA MUSEUM.
wooden frames, their ornate dust-catching design, abnormal length,
height and depth, low bases, horizontal sashes obstructing the
view and hinged or sliding doors. All of these features were a dis-
advantage from the visitors’ view point. The modern museum
case is distinguished by its narrow framework, reduced length,
depth and height, its relatively high base which does away with
the necessity for stooping, its large area of plate glass, no obstruct-
ing sashes and easily removable knock-down sides instead of
hinged or sliding doors.
All museum cases may be classed under two headings, i.e., cases
in which the exhibits are shown on a horizontal surface (horizontal
or desk or table cases) and cases in which the exhibits are shown
540 JOURNAL, BOMBAY NATORAL HIST. SOCGIZEDY, Vo Lee
on a vertical surface (vertical or wall cases) a comparison of the
advantages of the two types yields the following results :—
Vertical case. Horizontal case.
Maximum of exhibition space with Maximum floor space with
minimum or floor area. minimum exhibition area.
Clean glass (vertical glass is rarely Soiled glass (horizontal glass
touched by the visitor). is leaned upon and touched
by every visitor).
Comfort for the visitor. Discomfort, the body must be
bent at a fatiguing angle to
see the exhibits.
Well and evenly lighted objects, Visitor obstructs the light as
absence of eye-strain. he leans over the case.
Adaptable for any kind of object. Can only be used for small
objects.
iG, 2. TYPE OF DESK CASE IN USE IN THE GEOLOGICAL GALLERY,
SOUTH KENSINGTON,
MODERN MUSEUM METHODS 541
Owing to its many disadvantages the tendency to use the horizon-
tal or desk case is steadily decreasing, as exhibition cases of the
vertical type appear to be the most practical and economical for use
ina museum gallery. A very practical substitute for the ordinary
type of horizontal or desk case is the type of case illustrated in the
accompanying diagram; by increasing the height of the case and
subsequently the angle of the slope a larger exhibition area is made
available without the occupation of additional floor space. The
exhibits are thrown into better relief and the objects more evenly
lighted, the glass is not leaned upon and the necessity for stooping
is obviated. This type of case is in use in the Geological gallery at
South Kensington, London. The case has a solid base which
provides storage accommodation. The solid base has been much
in use in museum show cases as it provides storage accommodation
for exhibits not on display. Where a sufficiency of storage
accommodation is not available, this is imperative though not
always convenient, the disadvantage of the use of the high solid
base for horizontal cages is that it detracts from the appearance of the
case, is more expensive and makes policing of the gallery
more difficult. On horizontal cases of the desk and table type, where
the base is 32” from the ground, the use of a leg base is preferable.
In arriving at the most suitable height for the base of a
case the comfort of the visitor should always be taken into
consideration. The placing of small objects a few inches from the
floor level, as is often done in museums, is perfectly useless as the
visitor would either have to go onhands and knees or bend down
at anuncomfortable angle to see them, and this he is usually
unprepared to do with the result that the exhibits so placed are
rarely looked at. It would be better to sacrifice this space and
limit the area available for exhibition to a suitable height from the
ground wherethe exhibits would have a _ better chance of being
looked at. Theeyes of the average person are 60” from the
ground, the most comfortable visual distance averages 1 foot, the
body may be lowered 15° without stooping. To prevent eye
Strain on the part of the visitor andthe necessity for excessive
stooping the bases of aildesk and table cases should not be lower
than 32” from the ground and the objects within these cases should
not be further than 2-4” from the glass. In wall cases, the height
of the base should not be lower than 24” and the depth of these
cases should not be in ordinary circumstances exceed 18” while
the height at which objects are shown should not exceed 62”.
The modern museum group requires a special type of case.
The most effective case forthe purpose is the type which is
‘builtin’ and shows none or very little framework, the group
being viewed as through a window in the wall of the gallery.
This type of case is particularly suitable for large habitat or
zonal groups, but the possibility of its use will depend onthe
architecture of the hall, and in the construction of the modern
museum galleries special provision is made for the installation of
these built-in cases. The accompanying photo (Plate IV) shows
series of built-in groups in the design for the Hall of African
Mammals of the American Museum of Natural History, New York.
12
542 JOURNAL, BOMBAY NATURAL AIST. SCCIETY, Vol, oa
Lacking requisite architectural facilities, group cases may be instal-
led on the floor of a gallery and built in the alcoves formed between
large floor cases. One such group was recently installed in our
Museum. It is built between two large floor cases. Itis 14 ft. in
length, 7 ft. deep and 11 ft. high—the window measures 6 ft. & 4 ft.
RQ
Fic. 4. Ground Plan of Group Case
showing curve of Background.
Fic. 3. Front view of Group Case.
The panels on each side may be used
for photographs or transparencies.
As regards dimensions of these cases the most satisfactory ruie
is to have the length double the depth. As regards the background
its main portion is flat but where it approaches the sides of the case
it takes a gradual curve. ‘The sides of the case should not be too
long as it is difficult to avoid a ‘strained’ angle in the objects .
painted on it.
The ceiling or top of the case should be made as high as
possible so that on looking at the group the visitor does not see
the line where the painted background meets the top of the case.
Reflections from outside the case generally mar the effect of many
museum groups. To avoid reflections, the volume of light within the
case must be greater than the volume without, and in groups of this
type artificial illumination from within is nearly always necessary.
By its use any desired lighting effect may be achieved. The
lights are generally placed behind the upper portion of the front
panel of the case. To avoid shadows and to shed a diffused light
within the case, the reflectors of the lamps are directed towards the
top of the case from which the light is deflected on to the objects
below, or the light may be thrown on to a white painted board
attached on hinges and placed immediately in front of the lights.
By fixing this board at any required angle the volume of light
reflected into the case can be controlled.
Reflections may be further reduced by fixing the front glass of
the case at an angle, the degree of the slope being one inch to
every foot of height.
As regards the construction of the curved background this may
be done as follows. A form with the required curve is built by
running up 2” & 4” joists which are held together with cross
bracings (Plate V). This form is then covered over with pieces of
}
}
PLATE V
Amer. Mus, of Nat. Hist., New York.
ND IN COURSE OF PREPARATION FOR A GROUP AT THE AMERICAN MUSEUM
}
CURVED BACKGROU
By courtesy
Journ., Bombay Nat. Hist. Soc.
OF NATURAL HISTORY, NEw YORK.
“OSvIIYD ‘wnasnyy Pjaut
IA ALVIg
‘NOILISIHXY AO
GOHLAW SSHITIGHS AHL ONIMOHS saulg 4Ao
aASVO VW
"205 °3S1ff
Asaqanoo Ag
‘yeN) Aequiog “uaAnor
Painted Backgrourmda.
MODERN MUSEUM METHODS 543
plaster board which are screwed into it. The plaster board is made
by running plaster into flat or curved moulds as required and intro-
ducing layers of fibre and burlap into the plaster before it has set.
Fic. 5. Side view of Group Case.
When the whole form has been covered with plaster board the
joints between the boards are filled with plaster, the whole surface
is then sand papered and coated with shellac and when dry painted
with two to three coats of white zinc. An alternative method is to
cover the form with wire mesh, mache is then forced into the mesh
and spread evenly over it. It is then shellacked and allowed to
dry—when dry a second layer of wire mesh and mache’ is laid on
and then a third, each layer being shellacked and allowed to dry
before the next one is added. It is then white leaded and covered
with muslin as previously described. The photograph illustrates
a group case in construction at the American Museum of Natural
History, New York. A third method is to cover the curved form
with strips of 4” & 4” wood, then coat the whole surface with
shellac. When dry, mix varnish and white lead to a syrupy solution,
strain it through close-meshed wire netting and brush it over the
surface doing one section at a time, covering each section as it is
brushed with a layer of muslin which must be rolled on quite
smooth. When dry the whole surface is coated with white lead—-a
good quality of white lead must be used—the poorer qualities turn
yeliow after a time and have a bad effect on the painting.
Smaller groups are constructed on the same principle and will
often be found more practicable where conditions of space make
the installation of large groups impossible—a consecutive series of
sf. JOURNAL, BOMBAY WATURAL AIST. SOGKETY, Vol. SX0GTr
small groups illustrating the life histories of small mammals, birds,
reptiles, fishes or insects can be effectively arranged along the walls
of a gallery.
For the small museum one can do much more with small groups
and show much more than with large groups. ‘The background of
these groups, if curved, can be made of cork lino screwed to a curved
form, or of card board. At the Colorado Museum the backgrounds
of the cases consist of coloured photographic transparencies on
ground glass—the photograph illustrating the actual environment
of the species forming the subject of the group.
Shelves. —In many museums the use of shelves within the cases is
being done away with because they tend to produce a monotonous
lineal form of arrangement. ‘The shelf is being replaced by the
individual bracket which supports the specimen against the back-
grotind of the case. Most objects shown in museums lend theim-
selves to this style of exhibition. The accompanying photograph
(Plate VI) shows a case of birds in the Field Museum, Chicago,
where the customary shelf has been substituted by the bracket.
The Shrosbree bracket, a simple and efficient contrivance, which can
be accommodated to both small and large objects will be found most
useful for the purpose. It is supplied by the Public Museum and
Library, Milwaukee, Wisc. The use of the bracket offers greater
freedom of arrangement and better use of available space—the
objects are brought into sharp relief against the background and an
even distribution of lighting is assured. The labels are either fixed
to the pedestal supporting the specimen or are placed in metal
holders screwed into the back of the case.
We are adopting the shelfless method of arrangement in our Bird -
Gallery to very good effect.
For the background of our case we have used a screen made a 2
thicknesses of 3 ply wood fitted into a frame within the case so
that the screen with the exhibits on it is removable.
In arranging the birds ona screen it is first necessary to atte
how many birds are to be shown on the screen and the amount of
space each specimen will occupy—space being left if necessary for
subsequent additions—for this purpose rough sketches are made on
the screen indicating the position of the bird. In arranging the
» Specimens a lineal or monotonous arrangement should be avoided
and an attempt should be made to arrive at a free and harmonious
picture of the screen as a whole. When its genera design has been
completed the mounted birds are fixed to the screen so that changes
in arrangement if necessary may be made. When this is completed
the specimens are removed and the whole screen is wellsand paper-
ed and then painted with three coats of flat oil paint. The colour
is applied with an air brush—petrol is used as amedium. ‘This
ensures a fine even flat coat of colour. Divisions between families
are indicated by fine strips of beading. Variety may be given to
the exhibits by introducing individual painted backgrounds showing
the environment of a particular species and by the introduction of
nests, foliage, etc. The labels are held in metal holders screwed on
to the screen and coloured in a tone similar to the background.
(Lo be continued.)
BOMBAY NATURAL HISTORY SOCIETY’S MAMMAL SURVEY OF
INDIA, BURMA AND CEYLON
REPORT No. 46
ON A COLLECTION FROM TOUNGOO, BURMA
BY
To Ey PRY;
South Kensington Museum :
COLLECTION ess if VINO. 46.
LOCALITY srl .+ - Soungoo District.
DATE: ... sii ... November 1926-February 1927.
COLLECTED BY Se .. Mr. J.M.D. Mackenzie, 1.F.s.
The collection made by Mr. Mackenzie in the District of Toungoo is valuable
in that it supplements those made by him in Proine and Pegu, and serves to il-
lustrate fairly well the fauna of the country north of those districts and adjoining
the Shan States; in fact the work done by him in these three districts covers a
considerable area which was left untouched by the MamimalSurvey. Although
a good many animals known, or supposed to exist in Burma, as noted by
Mrs. Lindsay when reviewing the survey work, have so far eluded capture, yet
the general fauna of this country is fairly well known, and to this knowledge
Mr. Mackenzie has contributed in no simail measure.
The present collection represents 14 genera and 18 species none of which are
actually new, but there are many specimens which were much wanted, notably
Tomeuies ph.. phayret.
MACACA MULATTA, Zimm.
The Rhesus
: (Synonymy in No. 16.)
Toungoo, 2 1. |
Specimens of this inonkey were included in the collections made in Prome
and the Shan States respectively, by Messrs. Mackenzie and Shortridge, so that
its existence in the intervening district of ‘Toungoo was more than probable,
still it is satisfactory that this has been proved. Blanford stated that this species
was known throughout Upper Burma and as far south as Bombay in India proper.
On the one hand no specimens were found in Tenasserim, Burmah, nor on the
other in Dharwar, Kanara, etc., S. India, during the Mammal Survey operations
consequently it inay be inferred that this macaque has not wandered further
south than about 18° N. latitude, and the mits of its habitat have now been
fairly well established.
CYNOPTERUS BRACHYOTIS ANGULATUS, Mill.
The Malay Short-nosed Fruit Bat
(Synonymy in No. 17)
Toungoo, 9 1.
Corresponds in all respects with specimens from Tenasserim and the Mergut
Archipelago.
SCOTOPHILUS CASTANEUS, Horsf.
The Chesnut Scotophil
(Synonymy in No. 17)
Mounsoo, ¢ 2,9. 5:
Hitherto recorded only from the Malay Peninsula, Tenasserim and the Meregui
Archipelago ; it is therefore noteworthy that it occurs so much further north.
\
546 JOURNAL, BOMBAY NATURAL HIST, SOCIETY, Vol. XXXT/
‘TAPHOZOUS LONGIMANUS, Hardw.
The Long-armed Sheath-tailed Bat.
(Synonymy in No. 6)
Toungoo,g 1, 22.
The male specimen differs considerably from the two females in colour being
red instead of dark brown, but not in other respects. A similar variation has
been observed in specimens from the Central Provinces.
TUPAIA BELANGERI BELANGERI, Wagn.
The Burmese Tree Shrew
Toungoo, gl.
Specimens from Pegu and Prome were identified by Mr. Thomas as true
T. belangeri, and this one tallies with them in fur and size of skull. It would
appear that this species is confined to the above mentioned area since
Mr. Thomas has separated into sub-species the animals found north and south
of it, naming them respectively 7. del. siccata and T. bel. tenaster.
VIVERRA ZIBETHA, Lin.
The Large Indian Civet
ae specimen appears to be a true zz6etha. Mr. Wroughton gave the names
picta and V. 2. pruinosa to specimens from: Chindwin and the Shan
oe respectively, but they differ so slightly from zz6etha that it is doubtful if
they will ultimately be accepted as subspecies.
PARADOXURUS BIRMANICUS, Wr.
The Burmese Palm Civet
(Synonymy in No. 16)
Toungoo, fl.
A young specimen, but the black lines on the back by which it is distinguished
from P. niger, are clearly marked.
PETAURISTA CINERACEUS, Blyth.
The Tenasserim Brown Flying Squirrel
(Synonymy in No. 29)
Toungoo, Jl.
Blanford considered this Burmese Flying Squirrel to be identical with the
very similar, though rather larger, animal P. philippensis, but this name is now
held to apply to a southern Indian form. ‘There are considerable differences in
colour notably in the tails which are grey in cineraceus and black in philip-
pensis. ‘The Society is indebted to Mr. Mackenzie for three other specimens
from Pegu and Prome, no others having been obtained by the Survey.
RATUFA GIGANTEA, McCl.
The Assam Giant Squirrel
(Synonymy in No. 14)
20 miles N. of Toungoo, J 1, $ 2.
CALLOSCIURUS FERRUGINEUS, F. Cuv.
The Burmese Bay Squirrel
(Synonymy in No. 16)
25 miles N. of Toungoo, #1, 21.
TOMEUTES PHAYREI PHAYREI, Blyth.
Phayre’s Squirrel
(Synonymy in No. 14)
2 miles E. of Toungoo, ¢ 1, <2 2;
oe) 9) rel
40 99 97 3
30 re) om) 3
15 - Jo
11 3
99
>? 9?
MAMMAL SURVEY OF INDIA, BURMA AND CEYLON 547
No true specimens of 7. ph. phayret were obtained by the Survey ; those
recorded in Report No. 14, being subsequently recognized as 7. ph. bdlanfordt.
The former are more highly coloured above and below, while the dark stripes
on the under sides are absent or much less accentuated in the latter.
TOMEUTES PYGERYTHRUS, Geoff.
The Irrawady Squtrret
(Synonymy in No. 29)
30 miles west of Toungoo, ¢ 2, @ 2.
20 re re orl.
40 3 ae Cn Gee os
Variations in colour, from a bright olivaceous to a dull grey, are very notice-
able in these specimens ; this is due no doubt to slight local differences in en-
vironment (such as thick or open forest), as they were all taken within a com-
paratively few miles of each other.
TAMIOPS MACCLELLANDI, Blyth.
The Striped Burmese Squirrel
(Synonymy in No. 14)
5 and 7 miles E. of Toungoo—@ 1 and ¢ 3.
Tamiops m. barbei was obtained from the Shan States, Prome, Mergui
Archipelago and Tenasserim, thus indicating that Toungoo is probably about
the southern limit to which macclellandi proper extends.
GUNOMYS VARIUS, Thos.
The Malay Mole Rat
(Synonymy in No. 17)
Toungoo, #1,Q 1.
It may perhaps be worth while to point out that Toungoo is the most north-
erly district from which G. varius has been recorded ; but as this rodent may
easily have been carried northward by river-boats, etc., up-country correspon-
dents of the Society (if there are any) might be asked to keep a look out
for it.
RATTUS CONCOLOR, Blyth.
The Little Burmese Rat
(Synonymy in No. 16)
20 miles north-west of ‘Toungoo, ¢ ].
Pounsoo,) 4-7; 23.
RATTUS RATTUS KHYENSIS, Hinton.
The Shan Tree Bat
(Synonymy in No. 1)
20 miles north-west of Toungoo, 2 1.
In the earlier reports all the common house-rats received from Burmah were
recorded as &. r. rufescens variety white under parts.
RATTUS RATTUS RUFESCENS, Gray.
The Common Indian Rat
(Synonymy in No. })
Toungoo, @ 2.
CANNOMYS PATER, THOS.
The Bay Bamboo Rat
(Synonymy in No. 20)
25 miles N. of Toungoo, ¢ 2, @ 4.
These specimen are undoubtedly C. pater. Geographically they should belong
to one or the other of the subspecies C.c. plumbescens or C. pater, made by
Thomas, and the general colour clearly indicates that they should be classed
as the latter.
A PRELIMINARY REPORT ON THE ANATOMY AND
LIFE HISTORY OF THE COMMON EDIBLE BACKWATER
OYSTER, OSTREA\ MADKASENSTS
(4 paper read betore the Indian Science Congress, Lakore, January,
1927). |
BY
So. TD. MOSES, M.A.,.B.2:S«, FR: Asl
(With 2 plates)
This backwater oyster, common in Madras Presidency in all
estuaries and favoured spots along the coast, owes its widespread
presence to its being generally unaffected by a long range of varia-
tion in the salinity of the water it lives in; a sudden onset of fresh
water, as during floods, however kills thousands annually. Being
commoner than the fresh water mussel, it may with advantage be
used as a type for dissection in the class room.
The anatomy of the Tuticorin backwater oyster was one of the
earliest items of research I did for Mr. Hornell, and later I had
opportunities to examine oysters from Pulicat, Ennur, Cochin,
Cannanore and Calicut. Inthe plankton collections of these places
I found occasionally the trochophore and veliger stages of the oyster.
These notes are presented in this form not because they are
complete but that others with greater facilities may fill up the gaps.
in them, e.g., among others, trace in the laboratory the complete life-
history from the egg to the spat.
Mr. Hornell in 1909 believed this species to be identical with the
Ceylon oyster identified as Ostvea cucullata and said it was closely
akin to the American O. virginica and stated that the two seemed to
be geographical races of one species. O. cucullata is, however, the
rock oyster which is characterized by the deep ridges of the shell
ending in dentate edges which interlocked tightly when the valves
came together. In 1910, however, Mr. Hornell differentiated the
Sind species with a white muscular scar, from ours which has a
purplish-black scar, but said that both appeared to be related to the
Portugese mud oyster O. angulata.
The Chiika Lake oyster once named O. gryphoides var cuttackensts
is now called O. virginiana by Vredenburg who says all our back-
water oysters present no difference to the American species. Hor-
nell agreed with Vrendenburg in 1917. H.B. Preston, the expert
in charge of the publication of the volumes on Mollusca in the
‘Fauna of India’ series, declared in 1916 the extremely variable
backwater oyster of the East and West Coasts as O. madrasensis. The
close similarity of our species. with the American in its anatomy
and life history may justify its being named O. virginiana var
madrasensts.
External appearance.—Nothing can be seen externally except the
hard shell the sharp ends of which cut the feet of coolies and others.
THE COMMON EDIBLE BACKWATER OYSTER 549
wading on oyster beds. The shell, white and flinty in appearance,
is covered by a thin cuticle which when fresh is slimy to the touch
on account of its being encrusted with muddy algal felting. Though
slipper is the typical shape, variation is the rule, as oysters are of
irregular growth habit the shape depending on age, overcrowding
and other conditions of life.
The shell as in all Pseudo-lamellibranchs is inequivalve. The
valves are united anteriorly by a dorsal hinge ligament which being
elastic keeps the shell agape. The attached valve, the left, is
bigger and more convex and gives lodgment to the whole body
while the free right valve being flat functions as a lid.
The thickness of the shell also varies. Laminations and roughly
concentric lines of growth are seen externally. The latter start
from a swollen knob, the umbo, which as in other bivalves is the
early shell. It is recognizable by its dark color. The left valve
grows faster than the right.
The interior of the shell is like porcelain and white in color
except for the muscle scar and some patches along the margins
where it is purplish or purplish-black, with a touch of red.
When a piece of shell is broken off and the fracture examined
there are seen the three usual layers, a thin outer horny cuticle, a
thick—the thickness varies according to the varying number of strata
forming it at different places in the same shell—layer composed of
small columns of calcium carbonate set close together and a third
thin layer containing purplish and brown pigments in some parts.
It is the thick middle layer which is responsible for the reputation
of oyster shell grit to cure hens of laying soft eggs.
Soft parts —To examine the soft parts, the oyster should be
opened gently taking care not to damage the delicate organs. To
get specimens in a completely relaxed condition, they should be
killed overnight in a weak solution of Chloral hydrate. Soaking in
alcohol facilitates the dissection of nervous and other systems.
To open an oyster insert the point of a stout knife in the posterior
margin at a point about midway in the length. Then gradually
push the knife in helping this by slightly rocking the blade sideways.
Then when the knife is far enough pushed in cut through the muscle
and separate the free valve from the other.
The body thus exposed is seen to conform to the shape of the
valve. It is coated by a mantle which as in all bivalves is divided
into two flaps which enclose all the internal organs as if in a curtain.
It secretes the shell. Its free edges, purplish or dirty brown in
color, are fimbriated and bear two rows of very highly sensitive
short tentacles. Healthy oysters, when gaping, respond at once to
the slightest touch of the mantle edge with a needle by at once
tightening the valves.
Anterior to the muscle lies the visceral mass made up mainly
of the light colored reproductive organs and the dark digestive
glands. The visceral mass of spent oysters is dark colored, which
prejudices some, but wrongly, against using such oysters as
food.
Digestive system :—Anterior to the gills and between them and
the hinge are the two pairs of fleshy palps which enclose the oval
20
300 JOURNAL, BOMBAY NATURAL HIST. “SOGETY Wel. XOCaT
mouth. Food particles are brought in currents set by the action of
the ciliated gills and palps. The direction is easily seen by sprink-
ling powdered carmine in the gills of a freshly opened oyster. The
mouth leads through a short cesophagus into a stomach with thick
but translucent walls. The dark green liver around the stomach
sends many ducts to it. From the stomach leads the intestine which
bends on itself when at the end of the muscle on its ventral side and
coming back near its starting point describes a circle round the
stomach and goes out as the rectum to the dorsal side of the muscle,
to open out bya funnel-shaped anus. The rectum does not as in
the fresh water mussel, traverse the ventricle and so the way to the
heart is not through the stomach here. A crystalline style—absent
in spirit specimens—is seen in the first (pyloric) arm of the intes-
tines. The walls of the intestine and rectum are double indupli-
cated, as may also be seen from the imprimature on the feces.
The food of the oyster is mainly diatomic and is found in the
plankton of the water where it lives. About half the stomach
contents consists of unrecognizable vegetable debris but the follow-
ing were recognized :—Biddulphia, Rhizosolenia, Cheetoceras, Cos-
cinodiscus, Pleurosigma, Navicula, Ceratium, Foraminifera, Peridi--
nians, Dinophysis, sponge spicules, setee of Polychet larve and
Daphnids.
The enemies of the oyster are legion and they are active at all
stages of its life. Apart from some, e.g., other molluscs, barnacles
and aquatic flora, which crowd its beds to its disadvantage, there
are active enemies like crabs (Scylla serraia), starfishes (Luzdia
maculata and Astropectim), shellfish (AZurex tenuispina) and the boring:
sponge (Clione vastifica).
The oyster is edible, but Europeans who take it raw and in
its natural juice should see that the source is not a sewage
contaminated bed. The danger, though real, especiaily in a
country where washing after evacuation is done anywhere, is
exaggerated as oysters digest and kill injurious bacilli. Oysters
however muddy their original habitat, become clean if placed for
some days in a wholesome current and in factin the Madras
Aquarium clumps of live oysters when placed in tanks rendered
the water less turbid and more crystal-clear. Once a dozen oysters
from a suspected source as well as adozen clean ones were sent to
the King Institute, Guindy, for bacterial examination and both were
declared free.
Vascular system.—Anterior to the muscle and between it and the
visceral massis a transparent thin-walled sac, the pericardium,
containing the heart. There are two auricles and one ventricle.
The ventricle does not, as in Lamellidens, embrace the rectum. To
make out all the vessels, sections of hardened specimens are more
satisfactory than dissection or even injection. The aorta after
leaving the ventricle divides into two, one to supply the muscle
and the other to the visceral mass. The blood from these organs
go to the gills and after aeration flow into the auricles. The blood
is colorless.
The gills—Hanging down into the space between the two lobes
of the mantle are two pairs of gills. Their ventral edges are free,
PLATE I
ourn., Bombay Nat, Hist. Soe.
4
muscle
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ANATOMY OF Ostvea madrasensis
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ANATOMY OF Ostrea madrasensis
THE COMMON EDIBLE BACKWATER OYSTER 551
They are plaited and consist of a multitude of filaments closely cover-
ed with cilia. These cilia by their concerted action set the currents
going; the stream of water aerates the blood as well as supplies
food. The cilia also strain the food particles. The gills do not
serve as a broodpouch unlike in the freshwater mussel though it
does in another species O. edulis of Europe.
Muscular system.—The only muscle presentin the adult is the
posterior adductor, roughly rectangular with the dorsal side concave
and stretching transversely across the body from valve to valve.
The anterior of the two muscles present in the embryo atrophies
later. The mantle has avradiating bundle of muscular fibres which
withdraw the edge of the mantle from the margin of the shell anda
set of fibres distributed lengthwise along the edge, which by con-
traction make it crinkled. The muscle fibres as in all bivalves are
unstriped.
Excretory system.—The renal organs which are rudimentary,
consist of a pair of simple tubes lined by secretory cells. One open-
ing is into the pericardium while the other is into the mantle cavity,
as in other Pelecypods.
Nervous system.—lIt is very simple consisting of two pairs of gang-
lia (aggregations of nerve-cells). The anterior pair have two com-
misures one on each side uniting them to the posterior pair lying
below the muscle. ‘The anterior pair innervate the internal organs
while the mantle and muscles receive their nerve supply from the
posterior.
The two rows oi tentacles fringing the mantle are very sensitive
and function as tactile organs. Eyes are absent though their
function seems to be attended to by the marginal sense organs
which react quickly evento a shadow fiitting across the gape.
Neither otocysts noi osphradia are seen but oysters do hear and
detect changes in the salinity of the water. The veliger possesses
an otocyst.
Reproductive system.—The fatness of an oyster lies in the fulness
of the gonad which is found enveloping the visceral mass. The
sexes are separate. If the milky fluid from the gonad be examined
with a hand lens, that with particles present is the male’s and the
one without is the female’s. The gonads are large paired glands
and open out by ducts into the mantle cavity above the gills.
When well developed they are swollen and enwrap the dark diges-
tive glands—oysters are then very tasty to eat—the least pressure
making the cream colored fiuid exude. The ova and sperms pass
out into the water through the mantle cavity and fertilization is ex-
ternal and accidental. The oyster spawns all the year round though
two maximal seasons may be recognized, April to June and
September to November. Sexual activity is stimulated usually by
a reduction in the salinity and this depends on rainfall and sunshine.
The oyster passes through trochophore and veliger stages in its
life-history. The veliger possesses a foot which is functional till
the spat settles down to a fixed life. There is also an otocyst.
The growth after fixation is rapid, the oyster attaining a market-
able size in 1 years. The size attained within four months is
remarkable but the shell is thin. The thickness and weight of the
552 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
shell connote the age of the oyster better than its length and
breadth. The largest examined was from the Pulicat Lake and
the right valve measured 17” by 7”.
Among curious places where spat settled down, were the two
eyestalks of athus blinded crab Scylla serrata from Pulicat Lake.
The unfortunate crab had onits carapace many barnacies and on
its gills Lepads identified as Dichelaspts cor.
ORNITHOLOGY IN THE BIBLE
BY
an LATE ETr.eCou.€.E. Luarp
Ah, the land of the rustling of wings,
Which ts beyond the rivers of Ethiopia.
Before turning to the passages dealing with birds, a brief glance
at the geography of the region involved will be useful. ‘The area
with which we are concerned is roughly Palestine, that is Cannaan
and Syria and the neighbouring lands of Arabia, the Sinai Peninsula
and more distant Egypt.
Dean Stanley in his charming Szzaz and Palestine points out
how the Peninsula of Sinai is remarkable for possessing the three
grand features of earthly scenery, the sea, the desert and the moun-
tains. The great desert area is, however, cut into by three great
clefts, the Nile on the west, the Gulf of Suez in the centre and the
Gulf of Akaba on the east, which last merges into the deep valley of
the Arabah and yet deeper Jordan valley, further on. Palestine
forms a high level tract lying between the plains of Assyria and the
Mediterranean. Long before the war made ita reality, it was
familiar to all of us of the older generation from very early days,
as a land intimately connected with the absorption of tiresome
chronological and historical minutiz concerning an apparently very
overrated people. From north to south a great limestone ridge,
which splits into two parallel chains, traverses the whole country,
while four rivers the Orontes, Litani(or Leontes), the Barada (Abana
of the O.T.) and the Jordon flow through it. The last is the most
important and is the main artery of Palestine. On either side of it
run chains of mountains linking up with those of Sinai. Palestine
is thus curiously isolated. On the east is a desert, in the centre
the deep Jordon valley, a formidable trench, on the west the sea, on
the south desert again and on the north the Lebanon range. So
this small area, only some one hundred and forty miles long and
sixty wide, offers variations in climate, soil and vegetation which
make it an ideal place for animal life of all kinds. While the Jordon
valley falls to 2,300 feet below sea-level, the mountains of Lebanon
rise to over 2,000 feet above it, with snow-capped peaks. Thus
snow-covered hills, deep and fertile valleys, dry hot winds from the
deserts and cool sea breezes, offer unusual opportunities for bird
life, through the various plant and insect life the area produces.
Palestine lies in the Holarctic region of Newton or Palaearctic of
Sclater above the Ethiopian and Indian, and many bird forms
common in India are met with, such as bulbuls, the white-breasted
kingfisher, sun-birds, and many birds of prey.
The birds Moses encountered were probably much the same as
those met with in this region to-day.
554 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
The birds mentioned in the Bibie are not of course all those seen
by the writers but only those which entered most into their daily
lives either as beneficent or unkindly visitants, and those which
were striking and useful in metaphor.
To the Israelites lying out on the slopes of Mount Hermon or
watching their flocks and herds in the fertile plains, the birds of
prey were of peculiar interest, and but too familiar and dreaded
visitors. How often must the youthful David have watched
‘* the eagle that swoopeth on the prey.”’
We may, therefore, take the birds of prey together as a conspicuous
group. This is the more useful in that it 1s not always easy to say
if an eagle, kite or vulture is referred to. But is this surprising ?
Take any five of your friends out and see how many birds they can
distinguish even when not so closely connected as the birds of prey
or so similar in habits and appearance, that save in their size and
idiosyncracies of flight they offer no very distinctive signs.
I would here note that throughout this account the Revised
Version (R.V.) is referred to unless the Authorized Version (A.V.)
is especially noted.
The birds of prey mentioned are, egies the glede, falcons, gier-
eagle, the kite, hawk, osprey, ossifrage and vulture. The owls may
be dealt with separately,
The A.V. and the R.V. differ in the translation of the principal
passage which mentions these birds. I may add that throughout
this article I have quoted somewhat fully as many members of the
Society may be unfamiliar with our Bible, while my own experience
has proved how seldom my countrymen’s houses contain a copy of
the Revised Version.
The two passages lie in Leviticus and Deuteronomy and enume-
rate birds which were unclean and therefore not fit for food.
se. IV:
Lev. 11-13, 17:
And these ye shall have in abomination among fowls ; they shall not
be eaten, they are an abomination ; the eagle, and the gier-eagle, and
the osprey ; and the kite, and the faclon after its kind ; every raven after
its kind and the ostrich, and the night-hawk, and the sea-mew, and the
hawk after its kind ; and the little owl, and the cormorant, and the great
owl; and the horned owl, and the pelican, and the vulture ; and the
stork, the heron after its kind, and the hoopoe and the bat.
A.V. (as given)
Eagle, and the ossifrage, and the osprey and the vulture, and the kite
after his kind ; every raven after his kind ; and the owl, and the night-
hawk, and the cuckow, and the hawk after his kind, and the little owl,
and the cormorant, and the great owl, and the swan, and the pelican,
and the gier-eagle, and the stork, the heron after her kind, and the lap-
wing and the bat.
The changes are those of gier-eagle for ossifrage, kite for vulture,
falcon for kite, ostrich for owl, sea-mew for cuckow, horned owl for
swan, vulture for gier-eagle and hoopoe for lapwing. The bat seems
to have been looked upon as a bird.
ORNITHOLOGY IN THE BIBLE 555
In the corresponding passage in Deuteronomy (14-11) the lists
run thus:
Ra
Eagle, gier-eagle, osprey, glede, falcon, kite, raven, ostrich, night-
hawk, little owl, great owl, horned owl, pelican, vulture, cormorant,
stork, heron and hoopoe.
A.
Eagle, ossifrage, osprey, glede, kite, vulture, raven, owl, night-hawk,
cuckow, hawk, little owl, great ow!, swan, pelican, vier-eagle, cor-
morant, stork, heron and lapwing.
It should be borne in mind that the Hebrew word is of course the
same in the A.V. and R.V., only the interpretation is new.
BIRDS OF PREY
In his solitude and grief, and, be it added, the lecturing of his
prosy but well-meaning friends, Job seems to have turned instinc-
tively to nature.
Job 12-1:
No doubt ye are the people,
And wisdom shall die with you.
I have understanding as well as you
if But ask now the beasts, and they shall teach thee ;
And the fowls of the air, and they shall teach thee ;
Or speak to the earth, and it shall teach thee.
Some of the finest passages on birds are his. In chapters 36 and 37
Elihu pours out his long-winded discourse naively commencing with
Suffer me a little and I wili shew thee
For I have yet somewhat to say on God’s behalf.
Until God, losing patience, speaks from the storm which has
gathered as Elihu discoursed, and exciaims,
Who is he that darkeneth counsel by words without knowledge ?
and then asks Job who he is and of what account :
39-27. -
Doth the hawk soar by thy wisdom,
And stretch her wings toward the south ?
Doth the eagle mount up at thy command ?
And make her nest on high?
She dwelleth on the rock, and hath her lodging there,
Upon the crag of the rock and the stronghold.
From thence she spieth out the prey :
Her eyes behold it afar off.
Her young ones also suck up blood:
And where the slain is, there is she.
Possibly here some confusion has arisen with kites or vultures
but many eagles do eat carrion, at times. I have myself seen the
Indian tawny eagle (Aguzla vindhtana) sharing a dead cow with
kites, vultures and dogs.
556 JOURNAL, BOMBAY NAUTRAL AIST. SOCIETY, Vol. XXXII
The passage 1s an astonishing summary of eagle life. The
strength, swiftness and savage nature of these birds are constantly
used in metaphor:
Job 9-25 :
Now my days are swifter than a post :
They flee away, they see no good
They are passed away as the swift ships :
As the eagle that swoopeth on the prey.
The post is the ‘ harkara’ of India, each man doing his section
of ayjoumey =
Jev@ 51-31 :
One post shal] run to meet another.
Solomon used the eagle in metaphor:
Prov. 30-18 :
There be three things which are too wonderful for ime,
Yea, four which I know not:
The way of an eagle in the air,
The way of aserpent on a rock,
The way of a ship in the midst of the sea,
And the way of a man with a maid.
Then there is that fine passage in Jeremiah, in chapter 4, where
in verse 13 the swiftness of the eagle is referred to; it is dramatic
in form and for proper appreciation the speakers should be indicated:
Jer. 4-11:
10. Prophet.
Ah, Lord God! Surely thou hast greatly deceived this people and ©
Jerusalem, saying, ‘ Ye shall have peace’; whereas the sword reacheth
unto the soul. And at that time shall it be said to this people and to
Jerusalem.
A Cry +
A hot wind from the bare heights in the wilderness toward the daughter
of my people, not to fan, nor to cleanse.
The Lord :
A full wind from these shall come for me: now will I also utter judg-
ments against them. Behold, he shall come up as clouds and his chante
shall be as the whirlwind ; his horses swifter than eagles.
The People :
Woe unto us for we are spoiled :
Here whirlwind, horse and swooping eagle are brought together
as the swiftest things the writer knew.
In Habbakuk, which is really written throughout in dramatic form,
though this isnot brought out in our Bible, he refers to the approa-
ching Chaldean attack. The Prophet cries to the Lord in verses 1-4
and in verses 5-10; the Lord replies :
Vii10.3 7 HE. Ov 0-3
For, lo, I raise up the Chaldeans that bitter and hasty nation, which
march through the breadth of the earth to possess dwelling places which
are not theirs. They are terribleand dreadful: their judgment and their
dignity proceed from themselves. ‘Their horses also are swifter than leo-
pards, and are more fierce than evening wolves; and their horsemen
spread themselves: yea their horsemen come from far; they fly asan
eagle that hasteth to devour,
ORNITHOLOGY IN THE BIBLE 557
In some passages confusion between vultures and eagles can be
clearly detected. Thus:
Micah. 8-16:
Make thee bald, and poll thee for the children of thy delight,
Enlarge thy baldness as the eagle:
For they are gone into captivity from thee.
No eagles are bald, whereas vultures often are. Probably the
well-known passage in Matthew is another instance :
24-28 :
Wheresoever the carcase is there will the eagles be gathered together.
In both cases the marginal reading is ‘ vultures.’
In the wonderful song sung by Moses inthe 32nd chapter of
Deuteronomy the eagle is used in metaphor :
Went. 32-10, 11:
He found him in a desert land,
And in the waste howling wilderness ;
He compassed him about, he cared for him,
He kept him asthe apple of his eye:
As an eagle that stirreth up her nest,
That fluttereth over her young,
He spread abroad his wings, he took them,
He bare them on his pinions:
The A.V. has for the last line:
taketh them, beareth them on his wings
apparently with the idea that an eagle carried its young inthis way.
In that most wonderful of laments (quoted at the end of this article)
David compares Saul and Jonathan to eagles (II Sam. 8-17).
Like Moses, Isaiah uses eagles in referring to God’s care for
Israel: and the freeing of God’s servants :
Ts. 40-28 :
a % the everlasting God, the Lord, the Creator of the ends of
the earth, fainteth not neither is weary. ... Butthey that wait
upon the Lord shall renew their strength: they shall mount up with
wings as eagles : they shall run and not be weary, they shall walk and
not faint,
There are many other passages, which will be found inthe index
of references, but these are perhaps the most important.
Vultures are specifically referred toin certain passages, besides
the marginal alternative readings mentioned.
The vulture is one of the unclean birds, but the R.V. has gier-
eagle where the A.V. has vulture, and a marginal reading ‘ great
vulture’ against ‘eagle’. Inthe corresponding passage in Deute-
ronomy the R.V. has ‘ gier-eagle’ where the A.V. has ‘ ossifrage ’.
The vulture was of course as familiar an object to the Israelite as
it is to us in India.
The vulture occurs in no passages exceptin these lists, in the
R. V. as inthe two passages in the A. V. viz., Job 28-7 and Is.
34-15 we now have ‘bird of prey’ in the first and‘ kite’ inthe
second.
Without going into the question too closely it will be interésting
to consider the basis on which these translations are founded,
21
398 JOURNAL, BOMBAY NATURAL AIST. SCCVETY, Vol. 72ers
Generally the R.V. has accepted the premise that resemblance
between the modern Arabic name and the early Hebrew word is a
sure guide to identification of the bird.
The Hebrew word translated ‘eagle’ is ‘nesher’ from a root
meaning ‘to tear with the beak’. This corresponds with the
Arabic word ‘nasr’ (1) with an identical meaning which though
also used as a generic term, with qual:fying adjectives for eagles,
vultures, and ospreys, is usually appliedto the Gryphon vultures
(Gyps fulvus) and this is no doubt the bird meant. The passage in
Mich. 1-16 where the word‘ nesher’ is used alsoshews this while
the passages in Job 39-27, 30, Jer. 49-16 describe it well.
The Assyrian vulture god, Nisroch, morever, was the Gryphon
vulture, the marginal reading actually giving ‘ great vulture’. The
Hebrew word ‘ racham’ (Lev. 11-18; Deut. 14-17) is the same as
the Arabic ‘rakham’ and ‘rakhma ’(4*«+,) which the R.V. trans-
lates ‘ vulture” in place of the “gier-eagle’- of the AvV. ane
Arabic word designates the White Scavenger Vulture (Meophron
percnopterus) a near relative of the ‘ Pharaoh’s chicken’ of our
cantonments. The word ‘ vulture’ is a translation of three Hebrew
terms in the A. V., ‘da-ah’ (Lev. 11-14) and ‘ dayyah’ or ‘ dayyoth’
(Deut. 14-13; Is. 34-15). But in the R. V., it is rendered kite im
these places. The third term is ‘ ayyah’ (Lev. 11-14; Deut. 14-13;
Job 28-7) which the R. V. renders ‘falcon’. The latter certainly
means a small bird of prey and hence falcon is better ‘ hidayah’ in
Arabic is used of a kite. Possibly the ‘daah’ was the Red Kite
(Milvus regalts) and ‘ayyah’ the black (JZ. ater). |
The Gier-eagle or Ossifrage translates ‘peres (Lev. 11-13%
Deut. 14-12) or ‘the breaker’. The Lammageier or Bearded
Vulture (Gypetus barbatus) was said to throw tortoises, etc., down
from a height to break them, hence its name of ‘ bone-breaker ’,
The ‘ glede’ (Deut. 14-13) may be a kite or hawk. The word is
according to the Mew Oxford Dictionary used locally for buzzards,
the osprey, and the peregrine, but more properly for the kite
(47. regalis). The word is derived from a root meaning to ‘ glide’.
The Hebrew word so translated is ‘ raah’.
The word ‘hawk’ translates the Hebrew ‘nes’ (Gen. 40-10;
Lev. 11-16; Deut. 14-15; Job 39-26) and probably refers to the
Kestrils, of which 7. chenchrisis commonest: but it may refer to
the sparrow-hawk. But the termis generic for the smaller kinds
of bird of prey. Some eighteen species of hawk have been record-
ed from Palestine.
A few passages deal with these birds:
Job 39-26:
Doth the hawk soar by thy wisdom ?
And stretch her wings toward the south ?
Ts, 34-15:
There shall the kites be gathered, every one with her mate.
It is thus clear that it is no easy matter to distinguish the different
species from the words used.
ORNITHOLOGY IN THE BIBLE 559
In the New Testament in Matthew 24-28 and Luke 17-37 the word
translated eagle is ‘tos’ (aeros), though vultures are certainly
meant. In Revelations 4-7, 12-14 eagles may be intended.
Falcons besides being in the lists of the unclean are mentioned in
Job 28-7 where the A.V. has vulture. This last passage is very
obscure in the A. V., whereas the R.V. clearly shows that a mine is
being described, and well described :
Job 28, 1-11:
Surely there is a mine for silver,
And a place for gold which they refine.
Iron is taken out of the earth,
And brass is molten out of the stone.
Man setteth an end to darkness,
And searcheth out to the furthest bound
The stones of thick darkness and of the shadow of death.
He breaketh open a shaft away from where men sojourn ;
_ They are torgotten of the foot that passeth by ;
They hang afar from men, they swing to and fro.
As for the earth out of it cometh bread ;
And underneath it is turned up as it were by fire.
The stones thereof are the place of sapphires,
And it hath dust of gold.
That path no bird of prey knoweth,
Neither hath the falcon’s eye seen it :
The proud beasts have nof trodden it,
Nor hath the fierce lion passed thereby.
He putteth forth his hand upon the flinty rock ;
His eye seeth every precious thing.
He bindeth the streams that they trickle not ;
And the thing that is hid bringeth he forth to light.
One is apt to regret that the keen-sighted ‘vulture,’ as in the
A.V., was not retained where we now have falcon.
Game Birds.—The only game birds mentioned are the partridge
and the quail. It is curious that ducks are nowhere mentioned,
although they must have been common enough even if they did not
enter much into the life of the Jewish shepherd, and were perhaps
not eaten. It has been suggested that the ‘‘ fatted fowl,” of I Kings
4-23, may be a duck or a goose, as the latter were fattened by the
Egyptians.
Partridge is a translation of the Hebrew word ‘ qore,’ which
occurs twice only, in I Sam. 26-20, Jer. 17-11. The Hebrew name
iS OnOMmatopeic from the bird's call.
The ‘ partridge of the mountains’
(I Sam. 26-20) is probably Ammoperdix heyt which is very common
near the Dead Sea. It should, however, be noted that the Septua-
gint has, ‘is come out to seek my life’ where our Bible has ‘a
flea’. When hunted this bird conceals itself easily owing to its
protective colouring. The other partridge met with is the Red-
legged Chukor (Caccabis chukor) well known in India. Its cry 1s
heard everywhere. The Arabic name for this bird is ‘ karia’
(42,18). ae
The sandgrouse has been also suggested. In Eccus. 11-30 it is
mentioned as being used as a decoy ; or possibly for fighting :
Like as a partridge taken and kept in a cage so is the heart of the proud.
560 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Partridges are still used as decoys to catch Chukor.
The passage in Jeremiah has caused much comment: Jer. 17-11:
As the partridge that gathereth young which she hath not brought forth, so
is he that getteth riches, and not by right ; in the midst of his days they shall
leave him, and at his end he shall be a fool.
The words are spoken in a series of remarks not unlike proverbs.
The passage may be corrupt as there is nothing known of this bird
to lead one to suppose that partridge broods get mixed up.
Quail are first mentioned in Exodus 16-13:
And it came to pass at even that the quails came up, and covered the camp:
and in the morning the dew lay round about the camp.
Num. 13- 31:
And there went forth a wind from the Lord, and brought quails from the sea,
and let them fall by thecamp, about a day’s journey on this side, and a day’s
journey on the other side, round about the camp, and about two cubits above
the face of the earth. And the people rose up all that day, and all the night
and all the next day, and gathered the quails.
The same event is alluded to in Psalms 78-25 and 105-39:
The birds were migrating and driven by the high wind to rest
after crossing the sea. They come in March ordinarily in vast quan-
tities. The Hebrew for quail (Coturnix communis) is ‘ selav’ con-
nected with a root meaning ‘tobe plump’. The Arabic word is
similar, ‘salwa’ (Sglw)s Sandgrouse have been suggested here
also but so strong a flier would scarcely become exhausted in this
way. .
‘We may now pass on to birds of other classes. The words tran-
slated ‘ bird’ are in the Old Testament four in number (i) ‘ off,’
rendered by bird or fowl, the latter generally described as ‘ of
héaven’ or “of ‘the air’ (Gen. 1-21, 30;:Lev. 1-14; Jobia2-76
Ps.00-11etces)
(ii) ‘ayzt’, rendered ‘fowls’ or. ‘ birds.of prey’ in thetAi=V2
and R.V. respectively (Gen. 15-11, 13; Job. 28-7; Is. 46-11; Ez.
39-4; etc.) ;
(ill) * Zszppor’; > Atabie *-astun = (520!) is used of all small
twittering birds (Gen. 7-14; Lev. 14-4; Ps. 102-8; etc.);
(iv) ‘baal kanaph’ or ‘possessor of a wing’ (Pro. 1-17; Is.
10-14); -etc.):
In the New Testament two words are used ‘ ta petaina’ (ra
meterva) see Mt. 13-4; Lk. 13-19; and ornea (opvea) for ‘ birds of
prey’ (Rev. 18-2; 19-17, 21).
Birds were not unsympathetically noted as we find their salient
habits recorded. Migration is referred to, thus Isaiah evidently
takes this metaphor from large migrating flocks, when he says:
) eee oe
As birds flying so will the Lord of hosts protect Jerusalem.
Jer. 8-7 :
Yea, the storkin the heaven knoweth her appointed times, and the
turtle and the swallow and the crane observe the time of their coming ;
but my people know not the ordinance of the Lord (cf. Cant. 2-12.)
ORNITHOLOGY IN THE BIBLE 561
Care of their young (Deut. 32-11; Mic. 23-37; etc.) their helpless-
ness (Ps. 104-12). They were also tamed (James 3-7). Birds were
used as food when clean and not in the prohibited list, quail, par-
tridge, pigeons and doves are mentioned as eaten, hie the Tal-
mud gives a good deal of information on keeping birds for this pur-
pose. The very poor eat sparrows (Mt. 10-29), while the eggs of
clean birds were also used as food (Deut. 22-6; Is. 10-14; Lk. 11-12)
The verse in Job is doubtful :
Job 6-6:
Can that which hath no savour be eaten without salt ?
Or is there any taste in the white of an egg?
The marginal] reading ‘purslain’ for the white of an egg is ap-
parently correct and not asin the text.
Bittern.—-This is one of those cases, not many, in which the
Revisers seem to have gone astray. Where the A.V. has bittern
the R. V. has porcupine. As will be seen below it endows the
porcupine with vocal powers and an agility it certainly never
possessed.
Is. 14-23:
For Iwill also make it (Babylon) a possession for the bittern and
pools of water: and I will sweep it with the bosom of destruction.
A.V.
Zeph. 2-13:
And the flocks shall he down inthe midst of her (Nineveh), all the
beasts of the nations both the cormorant and the bittern shall eee in
the upper lintels of it ; their voice shall singin the windows.
ls, 34-11 :
But the cormorant and the bittern shall possess it; the owl and the
raven shal] dwell in it.
Whereas bitterns with their mysterious ‘booming’ call, cormo-
rants, ravens, and even the pelican, who takes the cormorant’s place
in the R.V., are suited to ruined places and pools of water and some
of them can also perch, no porcupine ever climbed on to a ‘ chapiter ’,
as the R.V. has it, or sang ina window! The Revisers took the
Hebrew word ‘ quippod’ as porcupine owing to its resemblance to
the Arabic ‘ kunfud’ (ai8).
Cocks and hens.—Domestic fowls came into use. it appears, about
4 ».c. since all earlier representations are of wild fowl, as the
drooping tail shows. The Old Testament nowhere mentions them
but we meet with them in the New Testament (Lk. 13-34). The
crowing of the cock is dramatically used in the well-known scene
with Peter (Mt. 14-30): this cock-crow was the third watch just
before dawn.
Cormorant and Pelican.—The word ‘qaath’ in Hebrew is tran-
slated cormorant in the A.V. but Pelican in the R.V. so that the
cormorant in the. R. V. is only mentioned in the lists of prohibited
birds. The root of the Hebrew word means ‘to vomit’ (Zeph.
2-t4;55 Is; 22-11;. 102-6). Actually the Pelician is not.‘ of the
wilderness’ in the sense of desert as it is a water haunting bird.
562 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Two species are common P. onocrotalus and P. crispus, the latter
having curled feathers on its head.
The word translated cormorant in the R.V. 1s ‘ shalakh’ that is
a ‘diver’ (Lev. 11-17; Deut. 14-17). Its harsh cry is well known:
There are two species Phalacrocorax carbo and P. pygmeus.
Crane and Swallow.—Translation has associated these very differ-
ent birds, and the swift. The word ‘ sus’ or ‘ sis (Is. 38-14; Jer. 8-6;
Zac. 10-3; Ps. 32-9))is inthe A: V. rendered by crane butinghe
R.V. by swallow. It should actually be swift. In the same pass-
ages the A.V. translates ‘ agur’ swallow and the R.V. crane.
ls. 38-14:
Like a swallow or a crane so did I chatter,
I did mourn as a dove.
Jer. 8-7.
and the swallow and the crane observe the time of their
coming.
In Psalm 84-3; 26-2: it is the word ‘deror’ that is transiated
rightly swallow : 84-3:
Yea, the sparrow hath found her an house,
And the swallow a nest for herself, where she may lay her young, -
Even thine altars, O Lord of hosts, my King and my God.
These references are generally appropriate except that the term
chatter is not very clear as though swallows might perhaps be said
to chatter, cranes utter a more sonorous and trumpet-like note.
Cranes (Grus communis) are met with in Winter in enormous .-
numbers. Dean Stanley recounts how near Wadi Haderah the sky
was literally darkened by a vast flight of red-legged cranes, with
black and white wings. In the case of swifts and swallows in India
one is familiar with their hordes building in temples and mosques.
Crow.—Not perhaps really distinguished from the raven. But it
is only mentioned in the Apocrypha, in Baruch 6-54.
Cuckow and Sea-mew.—The Cuckoo (Lev. 11-16: Deut. 14-15) has
vanished from the R.V. giving place to the Sea-mew. The Hebrew
word is ‘shahaph.’ It was a water bird and may have been a gull.
It is rightly placed in the list of the unclean which a cuckoo would
not be.
Dove and Turtle-dove.—Words used in the Hebrew are three, and
are not clearly distinguished as between doves and pigeons, which
is not surprising seeing that modern ornithology cannot successfully
differentiate between the two. The three words used in Hebrew for
these birds are :—(i) ‘ yonah’ (Lev. 5-7) which come from a root
meaning to mourn and refers more to pigeons; (ii) ‘Zor’ meaning
a turtle-dove (Lev. 12-6; Can. 2-12), an onomatopzic word repre-
senting its cry, and (iii) ‘ gozal’ really the young of any bird, a
‘ cheeper * (Deut. 22-11) but also used for young pigeon (Gen. 15-9).
The first mention of the dove is in the story of the Flood
(Gen. 8-8). Doves were evidently domesticated in early days.
They were used for sacrifice as were also young pigeons (Gen.15-9).
The brilliance of its colouring (Ps. 68-13), its harmlessness (Mt.
10-16), its helplessness (Ps. 74-19) are all referred to. It was used
ORNITHOLOGY IN THE BIBLE 563
as a common name and as a term of affection (Cant. 1-15; 4-1;
Se eli 6-9) ht ise alsou used as a type of. the Holy Spirit
(Mt. 3-16).
Dean Stanley describes how at Askalon the doves formerly sacred
to Venus still inhabited the haunts of their ancestors, though the
temple with its rites had long passed away. Ina siege the dung of
doves was eaten (I K 6-25).
Blue rock pigeons are met with in large flocks in the ‘ wadis’ or
gorges: read pigeons for doves below :
Ts. 60-8 :
Who are these that fly as a cloud,
And as the doves to their windows.
In Ps. 55-6 it should read (yonah) pigeons:
And I said, Oh had I wings like a dove
Then would I fly away and be at rest.
Lo, then would I wander far off,
I would lodge in the wilderness.
I would haste me to a shelter
From the storm, wind and tempest.
The arrival of these birds is referred to also (Jer. 8-7; Can. 2-11,
2);
ae of the turtle doves (7. communis) comes in April, another
(7. senegalensis) roams the streets of towns half tame.
Fowl.—The word ‘ayyit’ is translated ‘fowl’ in the A.V. in
certain places in which the R.V. has ‘ birds of prey’ (Gen. 15-11;
Job 28-7 ; Ez. 39-17-——birds) elsewhere fowl is retained.
Hoopoe and Lapwing.—The Lapwing of the A.V. has become
Ioopoe (Upwpa .epops) in the R.V. The Hebrew word. is
‘dukipath ’’ (Lev. 11-19; Deut. 14-18). It is only mentioned among
the unclean birds. The Hoopoe is a common spring visitor.
Heron.—This bird is only in the prohibited list (Lev. 9-19; Deut.
14-18). The R.V. has ‘ ibis’ as a marginal reading. Herons
are abundant (A. cinerea and A. purpurea). The ibis consorts with
herons.
Hunting.— It will not be out of place to make some reference to
hunting. Hunting for sport is a product of comparatively advanced
civilization. -Man could not afford to hunt for sport so long as his
food supply had to be obtained in the chase. Nimrod was not a
hunter for sport, but a mighty slayer of noxious wild animals.
The hunter is noted as uncouth (Gen. 10-9) but cunning. Birds
were hunted (Sam. 26-20) as the partridge, and according to the
law they had to be bled, similarly to the ‘ halal’ of the Musalman:
Lev, 17-13:
And whatsoever man there be of the house of Israel, or of the
strangers that sojourn among them, which taketh in hunting any beast
or fowl that may be eaten; he shall pour out the blood thereof, and
cover it with dust. (See Deut. 12-16).
Bows and arrows were used (Gen. 27-3; etc.) and these were at
times of bronze or bronze mounted (Ps. 18-34; Job 20-24) the arrows
were tipped with flints or iron. Slings were in common use, the
tribe of Benjamin excelling in their use (Jud. 20-16; 1 Chr. 12-2),
564 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
Nets (vesbeth-—Prov. 1-17; Hos. 5-1) and snares or nooses (moquesh—
Amos 3-5) are mentioned and clubs (Jer. 41-39).
Ants 3-5:
Can a bird fall in a snare upon the earth, where no gin is set for
him?
Ps. 69-20:
Let their table before them become a snare;
And when they are in peace, let it become a trap
Job 18-9 :
A gin shall take him by the heel,
And a snare shall lay hold on him.
The A.V. has ‘robber ’ for snare.
The noose was used as also traps (Job 18-10). The fowler used
such devices (Hos. 9-8). The word ‘cage’ in Jer. 5-27 should
be ‘trap’. Pits were used (Ez. 19-8). Decoys are mentioned
(Eccus. 11-30).
Nests.—Many reference are made to nests, and a warning is given
as to letting the parent birds go free (Deut. 22-6). Job refers to his
house as his nest (Job 29-18). The Hebrew words is ‘ gen’ (Gen.
6-14; Deut. 32-11; Nu. 24-21 ; Jer. 14-16 > elab. 2-9: Job 39-27).
Night-hawk.—The word rendered night-hawk is in Hebrew
‘techmas’ which merely means a bird of prey being connected
with a root‘to tear’. The night jar is in no sense a bird of prey
and identification is, therefore, uncertain. The Septuagint and
Vulgate translate it by owl (Lev. 11-16; Deut. 14-15).
Osprey.—The Osprey is only given in the prohibited lists in
Leviticus and Deuteronomy, and is a translation of the Hebrew
‘ozniyyah’. It is unlikely to have been the osprey as there were
no hunting grounds for it and probably it is another eagle ora
harrier. The word osprey (Fr. osfraze) is itself merely a corruption
of ossifraga, the name of the lammageier.
Ossifrage.—See birds of prey above.
Ostrich.—In the A.V. the Hebrew words ‘bath yaanah’ (Lev.
11-16 ; Deut. 14-15; Job 30-29; Is. 13-21 ; 34-13; 43-20 ; Jer. 50-39 ;
Mic. 1-8) are translated ‘owl’. But the word means ‘ daughter of
greed’. Two other words ‘yeenim’ (Lam. 4-3) and ‘reanim’
(Job 39-13) are translated ow] and peacock in the A.V., but should
be ostrich (Stvuthzo camelus) which is met with in Barbary, Syria,
Arabia and even Mesopotamia, while formerly it inhabited Egypt.
The ostrich has a very weird cty not, it is said, unlike a loue
groan.
Mic. 1-8:
For this I will wail and howl ;
I will go stripped and naked:
I will make a wailing like the jackals,
And a mourning like the ostriches,
Its preference for lonely spots is noted :
Job 30-29:
I am a brother to jackals
And a companion to ostriches
ORNITHOLOGY IN THE BIBLE 565
Then Isaiah in describing the destruction of Edom cries :
Ts. 34-13 :
And thorns shall come up in her places, nettles and thistles in the fort-
resses thereof; and it shall be an habitation of jackals, a court for
ostriches.
In that great lecture on natural history which comes in the 29th
Chapter of Job there is a dissertation on the ostrich :
Job 39-13:
The wing of the ostrich rejoiceth ;
But are her pinions and feather kindly ?
For she leaveth her eggs on the earth,
And warmeth them in the dust,
And forgetteth that the foot may crush them,
Or that the wild beast may trample them.
She is hardened against her young ones, as if they were not hers,
Though her labour be in vain, she is without fear ;
Because God hath deprived her of wisdom.
Neither hath he imparted to her understanding.
What time she lifteth up herself on high,
She scorneth the horse and his rider.
The ostrich is too curious an instance of bird life not to attract
attention, for a bird eight feet high which defends itself by kicking
like ahorse, has long eyelashes, only two toes and no wings to
speak of, except ornamental plumes is an anomaly. The ostrich is
polygamous, one cock having three or fourhens. ‘They lay ina
common nest, a shallow pit scraped by their feet in the sand, which
is heaped upon the sides asa _ protective wall. The cock begins
the brooding as soon as a dozen eggs orso are laid, especially at
night, by day their attendance is for watch and ward rather than
hatching which is done by the sun’srays. The recognition of
individual offspring by the mothers was thus not possible.
Owls.—The correction of owl to ostrich is given above. The
words rendered owl are ‘ yanstiph ’ (Lev. 11-17 ; Deut. 14-16 ; Is. 34-
11), the ‘great owl’ but the Septuagint reads ibis: ‘kos’ (ib. and
Ps. 102-6) little ‘owl’ and ow] ; ‘tinshemeth’ (Lev. 11-18 ; Deut. 14-
16; Is. 34-11) is translated ‘horned owl’ in the R.V. but swan in
the A.V. ‘qippoz’ (Is. 34-15) ‘great owl’ in A.V. and ‘arrow-
snake’ in R.V. but as obviously a bird is meantthe R.V. seems
inerror here; ‘lilith’ (Is. 34-14) in A.V. ‘ screech-owl’ becomes
‘nightmonster’ in R.V. asin margin of A.V. Lilith is a fabulous
monster, hairy and very fond of attacking infants, often appearing
in female shape. Owls are very common in Palestine. <Athene glaux,
the ‘bomeh’, is very tame and a great favourite, Otus ascalaphus,
étec.
Peacocks. —Mentioned in I. K. 10-22, 2 Chr. 9-21.
Once every year came the navy of Tarshish bringing gold and silver,
ivory and apes and peacocks.
The word is ‘ tukkiyyim ’ and as the bird was not indigenous but
imported the name is considered to be a corruption of the Tamil
word for this bird, ‘tokei’. The passage in Job 39-13 has ostrich
in R.V. for the peacock of A.V., the 'Hebrew word being
‘renanim ’.
Pelican.—See Cormorant.
tats
266 JOURNAL, BOMBAY WNATUCRAL HIST). SOCIETY. Vols2eodn
Porcupine.—See Bittern.
Raven.—It was an unclean bird (Lev. 11-15; Deut. 14-14) and no
doubt is not always distinguished from crows inthe Bible. They
are common in Palestine. The Hebrew word ‘oreb’ (Ar. Ghurab)
comes from a root meaning ‘tobe black’, and hence includes all
corvidae.
Sea-mew.—See Cuckow.
Speckled bird of prey.—This occurs in Jer. 12-9. The passage is:
Is mine heritage unto meas a speckled bird of prey?
Are the birds of prey against her round about ?
It is not very clear what is meant except that birds of prey are
inimical.
Stork.—The stork is among the unclean birds, but the crane is
not (Lev. 11-19; Deut. 14-18, Job 39-13; Ps. 104-17; Jer. 8-7; Zac.
5-9). The Hebrew word is‘ chasidah’ meaning ‘loving kindness.
It is a useful bird in clearing caterpillars and locusts off the field ’,
and is much prized in consequence, and nicknamed by the Arabs
‘Am Said’ or the ‘ Father of goodluck’. It was hence sacred and
is perhaps, therefore, mentioned to preserve it. The white stork
(Cicona alba) is the commonest though the black variety (C. x¢gra)
occurs.
Sparrow.—The word translated sparrow is‘tsippor’ (Ps. 84-3 ;
102-7—fow] in A. V.) which means only a small twittering bird (Ar.
asfar ( j)20!)). Small birds are not really distinguished and it is not
possible to say what bird is actually intended. The ‘sparrow on
the house top’ (Ps. 102-7) is possibly the Petrocossyphus cyanea,
which is very common, its monotonous note being heard everywhere. |
Swallow.—The Hebrew word ‘deror’ is rightly translated
swallow. See Crane. Many varieties are met with, A. rustica,
C. urbica, Cotyle rupestris, etc. Bee-eaters are possibly indicated by
the same word.
Swan.—This bird has vanished from the R.V. in Lev. 11-18 and
Deut. 14-16, and becomes ‘horned owl’ (‘tinshemeth’). But
probably the Ibis is meant or a coot ; the vulgate has ‘porphyrio’.
Before we leave the birds I wouid draw the attention of readers
to the great beauty of many of the passages in which the birds occur.
But it is essential, if their real beauty is to be seen, that they should
be read in the R. V., or better still in Professor Moulton’s ‘ Modern
Readers Bible’.
It is probable that no worse edited book ever issued from a press
than our Authorized Version, and itis a remarkable proof of our
unquestioning acceptance of the established, that for some three
hundred years we made no attempt to improve the form in which
the greatest treasure of the English language was placed before us.
As has been remarked, if Shakespeare, Bacon, Milton, Hallam, and
Tennyson were issued under one cover, no distinction being made
between prose and poetry, all tities being excised, the names of all
speakers omitted, speeches divided in the middle and sentences cut
up into a convenient length for a certain width of column, and the
chapters formed of an equal number of obtrusively numbered
verses, irrespective of sense or subject, such a work would scarcely
be an exaggeration of our Authorized Version, The Revised
ORNITHOLOGY IN THE BIBLE 567
Version, much as it leaves to be desired, is a great improvement ;
especially in the distinction made between prose and verse. But to
continue our subject.
These nomad open-air dwelling authors could scarcely have failed
to be influenced by the poetry of bird life, and so we find it.
In that wonderful song of Moses in the 32nd Chapter of Deutero-
nomy which begins :
Give ear, ye heavens, and I will speak ;
And let the earth hear the words of my mouth ;
My doctrine shall drop as the rain,
My speech shall distil as the dew,
As the small rain upon the tender grass,
And as the showers upon the herb.
and then he continues
es
For the Lord’s portion is his people ;
Jacob is the rock of his inheritance ;
He found him in a desert land,
And in the waste howling wilderness ;
He compassed him about, he cared for him,
He kept him as the apple of his eye ;
As the eagle that stirreth up her nest,
That fluttereth over her young,
He spread abroad his wings, he took them,
He bare them on his pinions ;
In the Song of Songs, that greatest of epithalamia, bird passages
are frequent; but to get the full sense of this dramatic poem the
speakers need to be indicated: the Bridegroom, the Bride, the King
and the Choruses. In the second phase, in which the Bride is recit-
ing the joys of courtship, we get a lovely spring song.
The Bride :
The voice of my beloved! Behold he cometh
Leaping upon the mountains
Skipping upon the hills.
My beloved is like a roe or a young hart
Behold he standeth behind our wall,
He looketh in at the windows
He sheweth himself through the lattice.
My beloved spake, and said unto me:
‘Rise up, my love, my fair one,
And come away.
For, lo, the winter is passed
The rain is over and gone
The flowers appear on the earth
The time of the singing birds is come,
And the voice of the turtle is heard in our land ;
The fig tree ripeneth her green figs,
And the vines are in blossom
They give forth their fragrance
Arise, my love, my fair one
And come away.
Oh my dove that art in the clefts of the rock,
In the covert of the steep place,
Let me see thy countenance,
Let me hear thy voice
For sweet is thy voice,
And thy countenance is comely.
One is tempted to quote for ever and it should all be read, but iead
intelligently.
568 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXL
In the Psalms, bird passages are many. We may take one instance ;
Ps. 91:
For he shall deliver thee from the snare of the fowler,
And from the noisome pestilence,
He shall cover thee with his pinions
And under his wings shalt thou take refuge :
and again in Isaiah, who often uses bird metaphors:
75. S*
Ah! theland of the rustling of wings
Which is beyond the rivers of Ethiopia ;
That sendeth ambassadors by the sea,
Even in vessels of papyrus upon the waters.
In Jeremiah. 4-23-26; in describing his vision of the destruction
of the earth, he cries :
I beheld the earth, and, lo, it was waste and void ; and the heavens and
they had no light. I beheld the mountains, and, lo, they trembled, and
all the hills moved to and fro. I beheld, and, lo, there was no man, and
all the birds of the heaven were fled. I beheld, and, lo, the fruitful field
was a wilderness, and all the cities thereof were broken down at the
presence of the Lord, and before his fierce anger.
One more passage and I must leave itto my readers to search for
themselves.
I will end with David’s lament for Saul and Jonathan:
1 Sam. 31-19:
Thy glory, O, Israel.
Is slain upon thy high eet 9
How are the mighty.
Fallen !
Tell it not in Gath,
Publish it not in the streets of Ashkelon ;
Lest the daughters of the Philistines rejoice,
Lest the daughters of the uncircumcized triumph.
Ye mountains of Gilboa, let there be no dew nor rain upon you,
Neither fields of offerings :
For the shield of the mighty was vilely cast away.
The shield of Saul, as of one not anointed with oils.
From the blood of the slain
From the feet of the mighty,
The bow of Jonathan turned not back,
And the sword of Saul returned not empty.
Saul and Jonathan were lovely and pleasant in their lives,
And in their death they were not divided
They were swifter than eagles
They were stronger than lions,
Ye daughters of Israel :
Weep over Saul,
Who clothed you in scarlet delicately
Who put ornaments of gold upon your apparel
How are the mighty.
Fallen in the midst of the battle
O, Jonathan,
Slain upon thy high places
Iam distressed for thee, my brother Jonathan
Very pleasant hast thou been unto me
Thy love tome was wonderful,
Passing the love of women.
How are the mighty.
Fallen !
And the weapons of war
Perished.
ORNITHOLOGY IN THE BIBLE 569
So whatever the technical imperfections of the translation we
should be thankful to the good fortune that gave us our Bible in the
age of Shakespeare and remember the words of Jesus, the son of
Sirach :—
‘Ye are entreated therefore to read with favour and attention, and
to pardon us if in any part of what wehave laboured to interpret we
may seem to have failed in some of the phrases. For things originally
spoken in the Hebrew have not the same force in them when they are
translated into another tongue.’
REFERENCES (REVISED VERSION)
GENERAL.—Watural History of the Bible. H. B. Tristram. Hebrew,
CoNCORDANCE—By G.M. Wigram. Dictzonary of the Bible.
BIRD, off : Gen. 1-21; 7-8, 14, etc.; Lev. 17-13; 20-25, etc.; Ps.
50-11, etc. (in general use).
ayit: Gen. 15-11; Is. 18-6; Job 28-7; Ez. 39-4.
UStppor 4 rene, ile y to=107; Wz, 17-23; Ps. 8-8; 11-1 (and
many other places).
baal kanaph: Gen. 1-21; 7-14; Ps. 78-27; 148-10; Job 39-13
(and many other places).
BITTERN (See: Porcupine): Is. 14-23; 34-11; Zep. 2-14. [so in
ALY. Only, |
CaGE: Jer. 5-27 (better ‘ trap ’).
Cocks (N. T. only): Mt. 26-34, 74,75; Mk. 13-35; 14-30, 68, 72;
Lk. 12-34, 60,61; Job 13-38, 18-27.
CORMORANT, shala@kh: Lev. 11-17; Deut. 14-17; Is. 34-11; Zep.
2-14 (see Pelican).
CRANE, agur: Is. 38-14; Jer. 8-7.
Crow (See Raven): Baruch 6-54.
Cuckow (See Sea-mew).
DovE AND TURTLE-DOVE, youdh: tor: gozal: Gen. 8-8, 9, 11;
Oa Slos9 eves I-i4 = 75-7, 11 < 12-6;, 8+ 14-22:, 30):
15-14, 29; 12-6; 14-22; Num. 6-10; Ps. 55-6; 68-13;
44-19'> Cant.” 1-15); 2-12) 14 ;-4-1 ; 5-2::12; 6-9: Ts.
38-14 ; 59-1] ;-60-8; Ez. 7-16; Nah. 2-7; Jer. 8-7;
48-28; Kos. 7-11; 11-11; Mat. 3-16; 10-16; 21-12;
Mack. 1-10) 2-24 + 3-22 In, Se 42-148 16;
EAGLE, nesher: Ex, 19-4; Lev. 11-13; Deut. 14-12; 28-49;
b2-4ks 2 Sam, 1-23 Job,)..9:26; 39-27 > Ps. .103-5:
Prov. 23-5; 30-17, 19; Is. 40-31; Jer. 4-13; 48-40 ;
49-16, 22; Lam. 4-19; Ez. 1-10; 10-14; 17-3, 7; Dan.
4-33; 7-4; Kos. 8-1; Obads 4; Mic. 1-16; Hab. 1-8;
Mat. 24-28; Lk. 17-37: Rev. 4-7; 12-14.
FALCON, ayyah (kite in A.V.): Lev. 11-14; Deut. 14-13; Job 28-17
(Vulture in A.V.)
Patrnp Fow. (Goose or Duck): 1. K. 22.
Fowc (See Birds).
HOWEER : Ps: 91-3; 124-7; . Prov. 6-5; Jer. 5-26; Hos. 9-8.
GIER-EAGLE, feres (Ossifrage in A. V.): Lev. 11-13; Deut. 14-12.
GLEDE, vahah: Deut. 14-13.
Hawk, Lev. 11-16; Deut. 14-15.
Hen: Mt. 23-37; Lk. 13-34.
570 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
HERON, anaphah: Lev. 11-19; Deut. 14-18.
Hoopok (Lapwing in A.V.): Lev. 11-19; Deut. 14-18.
HUNTING (See article): Lev. 11-19; Deut. 14-18: 1 Sam. 26-20,
Isis (See Owl).
KiTE, da-ah: Lev. 11-14; dayyah: Deut. 14-13.
Lapwinc: (A. V.) (See Hoopoe).
Nest: Gen. 6-143 Num. 24-21; Deut. 22:6 7732-19; Job i29-tee
32-27; Ps. 84-3; 104-175 Prov. 27-8. eleo) 101A
16-2; 34-15; Jer. 22-25; 42-28; 49-16; Ez. 31-6,
10-9.; (Ob.-4 : Hab. 2-9; Mat. 78220 -k 29°58:
Net (See Snares and Hunting).
NIGHT-HAWK, tachmas» Lev. 11-16; Deut. 14-15.
OSPREY, dzniyyah: Lev. 11-13; Deut. 14-12.
OSSIFRAGE (See Gier-eagle).
OstricH, bathya-anadh: Lev. 11-16; Deut. 14-15; Job 30-29; Is.
13-21; 34-13; 43-21; Jer. 50-39 ;° Micah 1-8; a-Aae
has owl.) ye-entm; La. 4-3; veanim: Job 39-13) A.
V. has peacock).
OwL, 4os: Lev. 11-17; Deut. 14-16; Ps. 102-6.
guippoz: Is. 34-15. [Great Owl in A.V.] Arrow-snake of
R.V. must be a bird.
tinshemeth : Lev. 11-18; Deut. 14-16. [Swanin A.V., horned
owl in R.V.]
liltih. is. 34-14 | Screech owl vot “AsV. |. an RV ice ndcann
Tonster.
gansuph: Lev. 11-17;. Deut. 14-16. In Is: 34-11 is) prez
bably the Ibis.
PEACOCK, 7Zkeiyy77- 1s Ke L022 47 AG. 29-2 15
PARTRIDGE, gove: Is. 26-20; Jer. 17-11; Sam. 26-20; Eccus.. 11-30;
PELICAN, gd-ath: Lev. 11-18; Deut. 14-17; Ps. 102-6; Is. 34-11]
Zep. 2-14.
PORCUPINE, gzppod (Bittern of A.V.): Is. 14-23; 24-11; Zep. 2-14.
QuallL, selar: Ex. 16-13; Num. 11-31; Ps. 106-409.
RAVEN, oveb: Gen. 8-7; Lev. 11-15; Deut. 14-14; I K. 17-4, 6;
Job 38-41; Ps. 147-9" Prov. 30-17 ;:°Cant. 5-10eaaise
34-11; Lk. 12-24.
SEA-MEW, shachaph: Lev. 11-18; Deut. 14-15.
SNARES (See Hunting): Ps. 91-3 ; 124-7; Prov. 7-23; Eccus. 9-12;
Hos. 9-8; Amos 3-5.
SPARROW, ¢észppor (any twittering bird): Lev. 14-4; Ps. 84-3;
102-7. > Mat? 1029:; Wks 12-6;
SPECKLED FOWL: Gen. 12-9.
STORK, chasidah: Lev. 11-19; Deut. 14-18; Job 39-13.
light of Ps, MOIA Ss wer. (38-7 | Zac. oes}
SWALLOW, devor: Ps. 84-3; Prov. 262: Is. 38-14; Jer. 8-7; "see
Crane).
SWAN, ¢znshemeth (See Ow] and Ibis): Lev. 11-18; Deut. 14-16.
TURTLEDOVE, tor: Gen. 15-9, Lev. 1-14; 5-7; 11; 12-6, 8; 14-22,
30;- 15-14, 29; Num. 6-10; Ps. 74-19: Cant; (2202s
Jer, 8-7; Lk. 2-24.
VULTURE, vacham: Lev. 11-18; Deut. 14-17.
REVIEWS
1. ‘STALKING BIG GAME WITH A CAMERA IN EQUATORIAL AFRICA
with a Monograph on the African Elephant.’ By Marius Maxwell. Royal
quarto, 200 pp., 113 plates—London: William Heinemann, Ltd. 1925., Price,
ozs. 62. net.
Mr. Maxwell has done yeoman service in the cause of the Natural History of
our larger game animals, and his book is virtually a plea for the protection
of wild life from ruthless and wanton destruction and from the complete exter-
mination with which a number of interesting forms are threatened in many
parts of the African continent.
The modern camera as a weapon for hunting game is fast gaining in popu-
larity with sportsmen all the world over, and this form of sport is one which
certainly deserves all possible encouragement. We hope the day is not far off
when it may supersede and completely replace the destructive rifle.
As the author points out, big game photography is an exhilerating sport and
provides just as much healthy recreation—and we may add, after a perusal of
his experiences, EXCITEMENT-—-somewhat more than the average sportsman
bargains for—as the hunting of game with a rifles Endless opportunities are at
the same time presented to the observer who wishes to study at first hand the
peculiarities of the larger creatures of the wilds commonly classed as dangerous,
Much useful knowledge may be gleaned by the naturalist regarding the ways
and habits of animals from photographs taken in their native surroundings and
under perfectly natural conditions, and to the modern artist, sculptor and
taxidermist such representations are invaluable for an accurate and truthful
portrayal of his subject.
The major part of the book deals in a facile and lucid manner with the
African elephant,—his ways, habits, gait, temperament and idiosyncracies.
Chapter V on the Senses of the African Elephant is particularly thorough and
illuminating and speaks much forthe care and attention bestowed by the
observer on the behaviour of his subject under varying conditions and circum-
stances, often extremely trying and tncomfortable. Supported as _ these
observations are by aseries of the finest photographs ever obtained of the
pachyderm, they supply the most valuable first-hand knowledge we possess
of these fast disappearing giants.
Excellent as have been the efforts of previous big game photographers, it
is significant that neither Carl Schillings nor Radclyffe Dugmore appear to
have been fortunate enough to succeed in photographing the African Elephant,
probably on account of the nature of the country the animal frequents. Schill-
ings with his telephoto outfit fully appreciated the difficulties involved, and
indeed it is only the modern photographic apparatus equipped with the
fastest of lenses that makes success possible in the gloomy ill-lit recesses of the
bush country which is their home in daytime. Dugmore, moreover, chiefly
employed fairly high power telephoto lenses which are wholly unsuited for
instantaneous work in indifferent light. It is interesting to note that Mr.
Maxwell advocates ordinary 3°5 to 4°5 (or faster) lenses with a focal length of
6 to 10 inches, such as he himself used, his plan being to stalk up carefully
as close to his sitters as possible. He believes that real success with elephants
lies only in the use of ordinary lenses at short range, as under the prevailing
conditions these give more detailed and faithtul results that can be obtained
from even telephoto lenses of low power. Maxwell’s views as regards a
suitable camera are apparently very different from Dugmore’s who fourteen years
ago (Camera Adventures in the African Wilds), considered that ‘ The ordinary
little hand camera with its short focus lens is practically useless. Only once
in a great while can one approach near enough to use it.’ According to
Dugmore the sort of camera necessary for the work is one of the long focus
reflex type ‘equipped with a convertible lens of high speed and a telephoto
lens of the greatest speed. The camera must be rigid enough to allow the
telephoto lens being used without danger of shaking.’
While acknowledging that both Schilling’s and Dugmore’s photographs are
unique in their own way, and the latter’s especially, as a whole, much
sharper and better as photographs, it must be remembered that Maxwell’s
object in making his pictures was of a more specific nature. He deplored the
$72 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol 2g
absence of photogiaphic illustrations in the epoch-making works of Selous and
Neumann, and set himself out to fillthe gap in the photographic records of the
larger game animals of Africa by securing a series of photographs illustrating
the life and habits of the African elephant in his native haunts, zot by using
such devices as camera-shelters, perches, flashlight or any other artificial means,
but by stalking the animals and portraying them exactly as they are seen by
the hunter of big game. The measure of his success can be readily gauged by
any one who glances over his wonderful array of photographs.
The photographer’s intrepidity leaves one agape. His experiences are
recorded in a simple matter-of-fact manner and without garnish or exaggeration
of the danger involved. In fact; the whole trend of Mr. Maxwell’s writings is
towards proving that the ferocity and truculence usually ascribed to even such
admittedly dangerous creatures as the African buffalo are often much overdone,
and that the natural tendency of wild animals is to leave man severely alone.
His opinion about the disposition of the African elephant is that as a general
rule, when not under severe provocation, heisa patient, retiring and even-
tempered beast and by no means as savage as heis usually depicted. He
admits however that ‘ Face to face meetings are at all times nasty incidents and
particularly speculative at close quarters.’ We like Mr. Maxwell’s prescription
of how to avoid the fury of a stampeding herd, and the advice is simple enough ;
it would appear that the elephant accommodatingly possesses an instinctive
aptitude for avoiding obstacles even in astampede. Well, the thing to do the
next time you find aherd bearing down on you isto stand behind the first
thorn tree of average size you come across, restrain your natura! inclination to
bolt, quietly watch the rampaging monsters brushing past with intermittent
deafening shrieks and shrill trumpetings, and safety is yours! Unfortunately,
however, ‘ There are of course exceptions to every rule’ !
One of the most remarkable pictures in the book, plate 5, Chapter III,.is of a
magnificent Masai bul! elephant taken at a distance of approximately eight yards
and under conditions that call forspecial mention. A face to face encounter
with the African elephant growing in size at each advancing stride must be one
of the finest and most impressive sights in nature, and, as the author says, ‘ To
watch for the first time the reflected image of an advancing elephant with ears
extended growing larger and larger on the focussing glass of a reflex camera is
perhaps the strangest of possible experiences, and one that is not easily forgot-
ten by the enthusiastic photographer.’ This particular bull, on detecting the
presence of the cameraman, deliberately moved forward towards him and
when some eight yards from the lens suddenly stopped. Life and death hung in
the balance. Suspense followed the click of the shutter. An eddy in the
wind-currents might have changed the course of events and sent the animal
either thundering down onthe intruder orcrashing through the bushin his
haste to get away !
The chapters on Giraffe, Rhinoceros and Hippopotamus are excellent, and
supported by a number of wonderful] photographs. [t is obvious that skilful
tactics are required to stalk long-sighted giraffes in the open bush country
which forms their usual habitat. The author has been able to secure a unique
series depicting the gait of the animal over whichthere has been some contro-
versy. His modus operandi wasto pursue the animals in a Ford Car and to
photograph them at close range with a short focus lens. Incidentally it is
interesting to learn that fora distance of a couple of miles or so when driven
to his utmost, 2 giraffe will keep up a speed of between 28 to 30 miles per hour
and that even at this speed they appear to glide along without effort.
The most remarkable achievement with rhino, in our opinion, was when the
author obtained four snapshcts in succession of a charging rhino in various
stages of the assault. The charge commenced at a distance of thirty yards,
when the first picture was taken. Three others were subsequently obtained as
the animal came on and was turned by a head shot from his companion’s rifle,
which covered the entire proceedings, and as the animal turned again and again
to repeat the attack on its aggressors. Any one who has used a reflex camera
will realize what a difficult proposition it is to change films, reset the shutter
and make exposures in the face of such an onslaught and in circumstances
which cannot but be exceedingly critical and trying.
The only grouse we have against Mr. Maxwell is that he omits to tell us what
make of film packs he was using. In view of the excellent results he has
obtained we think it was in fairness due to the makers of the film and to other
REVIEWS 573
enthusiastic photographers, whom the book may tempt to follow his lead, to
divulge this very important secret of his success.
We cannot too strongly recommend every sportsman and naturalist to
possess a copy of this book which will, we confidently believe, stand in a class
by itself for many a long day.
Sisters
2. POPULAR ZOOLOGY (in Marathi). By Vishnu Narayan Gokhale, B.ag.
B.Sc. (Lecturer, Agricultural College, Poona) and Vasude, Ganesh Desh-
pande, B.Ag. (Lecturer, Agricultural College, Poona).
Teachers of Nature Study in the Marathi-speaking areas of the Bombay
Presidency will welcome this volume. It deals thoroughly with animals of
various types in language easily understood by schoolchildren. The printing is
good. The drawings are numerous and excellent, and the size and price of the
book are moderate. The authors are two of the junior staff of the Poona
College of Agriculture and are to be congratulated on their very useful con-
tribution to vernacular scientific literature. We trust that the book will have
a large sale in the Marathi-speaking areas.
WB:
3. ‘A BIRD BOOK FOR THE POCKET.’ By Edmund Sandars. (Oxford
University Press. 7s. 6d. net.)
The little book that lies before us is of no importance from the point of view of
scientific ornithology. The author in his preface admits the fact and expressly
states that he does net claim to have added to existing knowledge. Hehas
merely tried to make a handy pocket book in which a resumé of the chief facts in
the life history of the ordinary British Birds is accompanied by ‘a just picture of
their beutiful forms and colouring’. We have therefore to judge the author's
book merely in the light of the success with which he has attained his own
objective. The illustrations are certainly no just picture of beauty of form or
colouring. With scarcely an exception theshape, attitude and drawing of the
birds is faulty while the colouring is crude and poorly reproduced.
The text is rather better. The short space of halfa page allotted to most
species is used to the best advantage with short crisp word pictures of the salient
features of the bird’s life, habits and characteristics. ‘hese are well expressed
and accurate, though they tend to become mechanical] and do not rise to special
cases. For instance neither illustration nor text would help a beginner to diffe:
rentiate between Willow-Wren and Chiffchaff except by their songs. Ten pages
are given to a general introduction which describes briefly the leading
characteristics of birds. This is helpful though it is dishgured by mistakes. For
instance under the head of moult, curriency is given to the mistaken belief in
change of colour without a moult by a change of pigment in feathers ; while the
statement that a stock-dove usually makes a hole in which to build its nest, and
that the ‘ the Bearded Tit and the Game Birds are scratchers of the grcund’ are
samples of inaccuracies that spoil the introduction.
Finally there isa general chapter ov eggs with indifferent coloured illustrations
of a few general types. The index omits the scientific names but gives English
and local names and a glossary of such words as seem to need explanation.
~The idea and design of the book are very useful, but the execution might well
have been improved upon.
H.W.
4. ‘TIGER AND OTHER GAME’ By Cot. A. E, STEWART. London:
Longmans, Green & Co. Ltd., 1927.
An excellent book. This and Dunbar Brander’s are the two best books I
have read. The wav in which the book has been written and the variety
of information and advice given by the author makes the work one of the
most useful a novice on shikar could have. Col. Stewart devotes Chapter
I to what he calls ‘ Preliminaries’ and in this very sound advice is given
on points of etiquette and politeness in correspondence with Government
Officials on the proposed shoot and in dealings with the local Forest Officers
and village Headmen, etc. The first portion of Chapter II also deals with
this, and excellent advice is given on the proper way of treating a jungle
23
574 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
native, and the author is quite correct in saying that it is here that
many soldier shikaries fail badly. The author has taken great pains
to explain the whole procedure and methods to be adopted in beating for tiger
and in sitting up for them. I notice that Col. Stewart takes an opposite view to
Geveral Wardrop with regard to following up wounded tiger. Col. Stewart
considers that atracker or two should go in front covered by the sportsman
with his rifle at the ready; General Wardrop holds that the sportsman should
be the one to goin front. I entirely side with Col. Stewart in this matter and
agree with his methods. In his chapter on sitting up for tiger the author writes
that a tiger when hit, if not knocked out, will usually charge in the direction he
is facing and that a sportsman must have a clear view and steady aim if firing
at a tiger facing him. As the book has been written, I take it, principally as
a guide to inexperienced shikaries I think perhaps that the author might have
said that on no account should a young shikari fire at a tiger facing him, but
that he should get his sight on to him and wait till he starts to move off before
firing. \tis hardly likely that a novice ona tiger shoot would be absolutely
steady in his aim, perhaps seeing his first tiger facing him, and a frontal shot in
this case might be disastrous. I am glad that Col. Stewart is not one of those
who absolutely ban the use of electric light when sitting up at night over a kill.
In many parts of India, especially in the South, and where beating is impracti-
cable, the use of artificial light is absolutely necessary.
Referring to the Sloth Bear the author remarks that it isa poor climber.
Though I have never heard of a man being dragged out of atree by a Sloth
Bear I should hardly describe it as a poorclimber. On the contrary I should
say that the Sloth Bear is an exceedingly good climber and the height they
will climb up a tree for figs and fruit is amazing.
Dealing with Bison the author writes that a tiger will think twice before he
will tackle a Bison. I should say Au/l Bison for | have known several cases of
cows being killed, and three cases of Solitary Bulls being killed by Tiger, but
I must admit that atiger that can tackle and kill a Solitary Bull must bea
large and powerful specimen.
I read the author’s paragraph on sitting up over a wallow for observation with
interest and pleasure, for this is what I enjoy doing exceedingly myself. If one
has a permanent machan erected on a tree over aswamp or wallow the game
get used to it and it attracts no attention whatever,
Writing on Grey Jungle Fowl the author remarks that he has never heard
them calling on the ground and that he does not think they do. He is however
quite mistaken here. Jungle Cock very often call on the ground during the
dayti:me. :
In his para on Taxidermists I think that the author’s advice would have
been better if he had said that heads and skins should be sent to the Bombay
Natural History Society, Bombay, who undertake taxidermy work for its
members, the standard of this work being equal to that produced anywhere,
or to Messrs. Van Ingen and Van Ingen, or Messrs. Theobald Bros., both of
Mysore. The Bombay Natural History Society employ the very latest methods
and pay great attention to details. Living in Sontkern India I personally deal
with Messrs. Van Ingen and Van Ingen and have always found them satisfactory,
they are a most up-to-date and skilled firm, and it can be truthfuily said that
the Bombay Natural History Society and this firm turn out work equal to, if
not better than taxidermists at home. This being so I see no reason for sports-
men to send their trophies home, especially as the author admits that duty has
to be paid on the trophies if they are brought out again.
There is one excellent preservative which I see the author does not mention
for preserving skins and that is Atlas ‘S’. Ithink there is nothing like it for
putting on skins if the animal is freshly skinned, to keep them in good condition
in hot jungles. This preservative is now used by the chief taxidermists and
tanneries in India.
The author’s advice on rifles and ammunition is very sound indeed, as alsois
his advice always to have a medicine chest in camp.
The Autobiography of Skaitan Bhalu and the illustrated interviews are very
good indeed and ferm a happily thought-out ending to a book which is full of
sound advice, useful information, and interesting anecdotes written in a plain
convincing style from start to fiaish. as
_.This book should be in the library of every sportsman. ;
Re Godh ont
EDITORIAL
EAieseh ye
In- the last issue of the Journal we published the proposed new constitution
of the Society and we invited criticism and in fact printed at the end of the
Bye-laws a slip which would enable members with very little trouble to themselves
to advise the Honorary Secretary what their opinion on the subject is.
Members will no doubt be anxious to know what the general opinion is—we
are sorry to say the general op:nion of members is “You can govern the Society
as you like, personally we do not care the value of the smallest Turkish co.n.’
Apathy—is it a tribute to the Committee? There has not been a s.ngle
objection. All who have sent in their forms have approved, but ‘all 1s
unfortunately not a word here which is an embracing one. It certainly does not
embrace ‘all’ the members of the Society but embraces a minute number. The
vast majority do not care.
If this attitude of the members towards the future constitution of the Society
be an expression of confidence in the Committee, the Committee accept it. but
hardly with gratitude. They would be more grateful if members took an interest
in what was going on; without interest being taken the Society is hardly lkely
to flourish.
Anyhow the response is sufficient to enable the Committee to go forward with
the scheme and we hope by the time the next Journal is issued, to be able to
announce that the Society is a registered Society and able to hold property
and Government Paper in its own name.
The new constitution necessitates a close investigation of our invested
securities—Do our investments equal the amount paid by living life members
or are they too small or are they in excess? The Treasurer's Statement of
Accounts should be in the next Journal, and will explain the actual financial
situation of the Society better than ever before. The monies invested in,
and derived from, the publications of the Society other than the Journal, and
the income and expenditure of the Taxidermy Department will be separated
from the regular income and expenditure of the Society under which head, as
regards expenditure, will be taken expenditure which refers to every member
of the Society—the issue of the Journal and the maintenance of the Society’s
Office staff, office and library. Without wishing to anticipate the Treasurer
we can state here that our investments more than equal the amount contri-
buted by living life members.
A question arises—if there be a surplus can that surplus be applied to any
object the furtherance of which lies within the constitution of the Society, and
if so, what would be the most pressing needs of the Society?
The most pressing need is a better illustrated Journal and perhaps the
encouragement by a modest honorarium of those who write about Natural
History subjects in a manner which appeals to the general member. The
Journal must be our main recruiter and our binding link, but we want something
more and we want a good museum—a museum for the student and research
worker as well as a museum for the general public.
The arrangement the Society entered into, now several years ago, with
the Bombay Government and the Trustees of the Prince of Wales’ Museum,
Bombay, has given members of the Society vastly improved conditions on
those of old—though it was the personal care bestowed on the old museum
that laid the foundaticns and enabled the superstructure to be erected. But
no one will pretend that the existing building is suitable for a Natural History
Museum nor that the accommodation provided is anything like sufficient for
present needs. Members will be glad to know therefore that there is a
possibility of the Trustees of the Prince of Wales’ Museum being able to
provide a new building for the needs of the Natural History Museum, and
that the building, while in keeping with the noble architectural design of the
late Mr. G. Whittett, will be one designed solely for the purpose for which
it is intended to be used, namely a museum for the instruction of the public
576 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXil
by the display of Natural History subjects in scenery characteristic of that in
which the fauna exhibited lives and has its being. For this we shall have
to thank not only the Trustees and the Architect, but the man who went to
America and Europe and learnt what a revolution in museum methods had
taken place, and having digested what he had learnt, proceeded, after the
manner of many good mothers, to give of the digested learning to the young
around him.
The Trustees of the Prince of Wales’ Museum will however experience
difficulty in finding the money for the actual building, and the whole scheme
may be held up unless some wealthy citizen of Bombay or resident in India
comes forward with a generous donation to the building fund as did the
first Sir Currimbhoy Ebrahim, Bart., for the existing museum. The fitting
up of the museum when built must, however, be the work of the Society and
here we may find the best use for surplus investments—always provided
of course that the membership is sufficiently large to enable us to carry
on the ordinary work of the Society without needing the interest derived
from surplus investments.
Whatever this surplus may be, it will be but a drop in the ocean compared
with what we need and we record therefore with great pleasure the appointment
of Sir David Ezra, Kt., as a Vice-Patron of our Society. Sir David is a citizen
of Calcutta who has done much for the Calcutta Zoo and for the Zoological
Gardens at Regents Park. It was Sir David who provided the funds for the
despatch to Bombay of the Elephant for the London Zoo which the British
India Steam Navigation Company so kindly took home free, and Sir Ezra’s
donation of Rs. 5,000 towards the furtherance of the Society’s work is not
only opportune and needed but of good omen.
We cannot say when this new building will be available but we are already
planning habitat groups—-groups which will enable the observer to realize
what the animal, bird, insect or. reptile is like in its natural surroundings,
and we are already in possession of promises of support from members who
are able to get us suitable trophies.
We welcome, and we aid, in the cause of Natural History those who come
to India and Burma on behalf of Museums situated in countries outside the
Empire; we realize that we shall benefit by studying their methods but why— ©
why—should the British Empire be left out of it? Are we to rest content
with the fact that if we want to see a good and representative exhibit of the
fauna of the Indian Empire we must go to America to see it? It rather looks
like it. The wild fauna of a country under modern conditions has no certainty
of continual existence. In what museums are there specimens of the almost
extinct Smaller One-horned Rhinoceros (Rhinoceros sondaicus) of the
Sundarbands, Tenasserim, and the Malay Peninsula? We know of none where
mounted specimens exist but there soon will be in America, thanks to the forth-
coming expedition of Col. Faunthorpe and Mr. Vernay. We hope that someone
connected with our Museum will be able to accompany these gentlemen or
their Artist Assistants, and if so, we look forward to the publication of an
interesting account of modern methods in the collecting field.
Our editorial in the last number of the Journal on Game Preservation nee
produced a certain amount of criticism—-not as much as we should have liked
in view of the great importance of the subject. Some of the criticism has
been adverse owing to the non-realization of the fact that in making out
a case for Game Preservation it was necessary to look at the question from
all sides, and an article such as we have written would have been incomplete
without some reference to the economic or, as we said, more ‘material motives’
for the protection of Game. The plea for protection of Big Game per se would
not necessarily carry the same weight with all classes of people.
The word ‘Game’ as considered in the framing of laws does not limit itself
to the class of animals usually sought after by the shikari but includes wild
life as a whole and would include the smaller carnivora which are hunted
and trapped for furs and which would equally require protection if they are
at all to be considered as an economic asset and they are seriously so considered
in other parts of the world but always in regions where they do not clash with
agricultural interests. We have stressed this point in our article where we
definitely state that the plea for protection cannot carry weight in urban lands
and cultivated areas. Our whole plea for protection is concentrated on waste
lands and in forest areas, where the interests of man are not brought into
See POOL LE DLO IAL. . 7 877
conflict with the wild species and there are sufficiently large areas in India
where these conditions prevail, which areas provide for the present the
strongest guarantee against the extermination of wild life. The article was
based mainly on correspondence received from members of the Society and
was hence restricted in the main to such provinces from which information was
available. We do not for a moment claim that it covers the question
completely or is free from defects, but its purpose is to rouse interest and
draw attention to a question which should be the concern of a Society such
as ours. .
We welcome all sane efforts to preserve the wild fauna of a country and in
our next issue we shall refer once more to the work of the Society for the
Preservation of the Fauna of the Empire with whom we intend affiliating.
In the present issue of the Journal we publish a review of Marius Maxwell’s
Book Stalking Big Game with a Camera in Equatorial Africa. A prospectus
of Mr. Maxwell’s book was forwarded to members with the October issue
of the Journal and in the present number we have the pleasure of giving
our readers a closer acquaintance with the author through the medium of
his very interesting paper on ‘The Home of the Eastern Gorilla’. Those who
have seen Mr. Maxwell!’s book or have the good fortune to possess a copy, will
realize that his magnificent series of portraits of African animal life require
no commendation from us. The astonishing success achieved by the author
led us to hope that someone would write the story of the Indian jungle as it
never has been written before, in pictures of equal beauty and merit. Mr. F.
W. Champion’s book With a Camera in Tiger-Land fulfils this hope.
Mr. Champion has done for India what Marius Maxwell, Dugmore, Schillin,ss
and others have done for Africa. He gives us intimate pictures of Indian
animals in a setting which carries with it the spirit of the Indian jungle more
effectively than pages of the printed word could have done. To those who love
the Indan jungles—be they those to whom its sights and sounds are now but
a happy memory or be they those who can still answer its call—we~ most
heartily commend Mr. Maxwell’s book and to the lucky latter we would-add
‘Go thou and do likewise’. The camera is increasingly becoming a
competitor to the rifle in the hunting of game. Hunting with a camera
has been rightly described as the expression of sport in its highest form.
It requires all the qualities of hardihood, of daring and resourcefulness, of
patient endurance and unflinching determination which are demanded from
the ordinary hunter. Five years of continuous effort, Mr. Champion admits,
have failed to produce for him a satisfactory picture of so common an animal
as the Indian Sambhar although he has stalked to within a few yards of these
magnificent creatures and could have shot a score of them in the same time.
Hunting with the camera does not have destruction as its aim; it offers instead
endless opportunities for the study, and enjoyable observation of wild life. It
is no part of the camera hunter’s business to frighten and disturb his quarry,
and the trophies he obtains are far more interesting than any that could have
fallen to his gun. The value and importance of his work in the faithful
portraiture of animal life in its natural setting must increase as the inexorable
march of civilization threatens, and will increasingly continue to threaten, wild
life even in its uttermost retreat.
It is but too readily apparent, even to one only casually interested in the
bird life of this country, how infinitismal is our present knowledge of
Bird migration in India and how negligible have been our endeavours so far
to further it, as compared with the efforts being made and the results achieved
in Europe and America. No systematic effort has so far been made towards
acquiring first-hand information regarding the movements of the hundreds of
species that visit us year after year, the influx commencing about September-
October and the exodus in March-April.
Individual observation is solely responsible for what little has been attained.
Such observation deserves every encouragement and, in spite of the difficulties
and shortcomings it presents, it is a mistake to underrate the value of such
information obtained by careful study. An accurate record of the appearance
and disappearance of a given species in any particular locality is manifestly
of great value especially if the records for that locality extended over a number
of seasons. By a systematic co-ordination of frequent and accurate notes of
this nature it is often possible to learn something of the movements of
578 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXX1]
migratory birds even though it is but very occasionally possible to see anything
of the migration in progress, and, unless it be on some isolated island on the
route taken by birds, preferably with a lighthouse, as for instance Heligoland
er Fair Isle (between the Orkney and Shetland group) it is extremely difficult
for individual observation to obtain direct evidence of migration; on the main
land, with its large and varied native population of birds, such a method can
at best yield very patchy results.
_ As distinct from the purely observational methods of study, which can clearly
be pursued only by a very few observers who are fortunate enough to be
stationed in favourable localities, there is the modern method of bird-marking.
The ‘ringing’ of birds consists of the fastening of a light aluminium ring of
appropriate size bearing a stamped number and address, on the instep region
or ‘shanks’ of trapped or netted birds. The birds are then released.
A small percentage of these ‘ringed’ birds are afterwards shot or recaptured
in distant parts of the globe and the rings are returned to the marking station
with data as to the exact locality and date of capture. When a _ large
number of such records have been collected and correlated, it is poss: ‘ble
gradually to build up knowledge of the routes followed in migration, and a
number of other important facts also which could not have been ascertained by
other means.
As an instance it may be cited that rings put on Storks in North Germany
have been returned from Damascus, Alexandria, the Blue Nine, the Kalahari
Desert, Basutoland and elsewhere, so that Dr. Thienemann, the Director of
the German Ornithological Society’ s* observation station at Rossitten on the
Baltic Coast (E. Prussia), was able to conclude definitely that there is an
autumnal migration of Storks from the shores of the Baltic to South Africa.
Some of the main problems whose solution we in India may in course of
time look for to this method of marking are :—
(1) To what extent do our migrants return to their birthplaces to breed
and under what circumstances are new areas colonized?
(2) Do birds take up definite quarters when visiting this country, and
if so, do they seek the same year after year?
(3) To what extent do the birds of a single species occupying the same
area during their winter visit, seek the same area for breeding in the northern
part of thelr range?
(4) Are definite routes followed by birds, and if so what are they, and
what is the nature of such routes (i.e. course of a river, etc.).
There are besides a great many other questions that need answering, and little
can be achieved unless there is a general realization of the fact that our knowledge
on any of the above heads is “practically nil, and unless a properly organized
and systematic effort is made on an extensive scale for the capture, marking
and release of our migrants throughout the country. It will be recalled that
possibly the only experiment of this kind in India was carried out early
in 1926 by the late Maharaja of Dhar, an ardent sportsman and naturalist.
The results of this experiment were published in the Journal (p. 1034 of
vol. xxxi) and a further recovery—the most interesting of the lot—was
recorded on p. 222 of vol. xxxii. The results must be regarded as highly
gratifying and indicate the scope and utility of the ringing method.
The Society proposes shortly (provided sufficient encouragement is forth-
coming) to launch on a campaign of bird marking, and we trust that all bird
lovers and members who have opportunities will come forward and co-operate
with us whole-heartedly. We propose to issue to members on request at a
nominal cost, aluminium stamped rings bearing a number and the Society’s
address, to be fastened round the legs of birds. Perhaps a beginning could
conveniently be made. with waterfowl.
Along with the rings, a schedule will be supplied to the member on which
data about all birds marked will have to be filled in by him such as (1) number
on ring, (2) species of bird, (3) date of marking and release, (4) locality of
marking and release and (5) sex and other remarks.
The completed schedules when returned to us will be posted up in a
special ledger. Any recoveries recorded will be entered up, and the marker
immediately informed.
We look to our sporting princes, District Officers and other members who
may be favourably stationed to co-operate in building up the much-needed
knowledge regarding the movements of our birds. All interested in the scheme
EDITORIAL 579
are cordially invited to communicate with the Honorary Secretary. Though
the time is short let us hope a few thousand birds will have been ringed before
the exodus commences in March and April.
We hope the present issue of the Journal will be appreciated but we have
misgivings as to the future. We have very little reserve matter beyond a
continuation of the interesting papers on Kashmir contributed by that veteran
naturalist Col. Ward. Members have never let us down in the past and we
appeal to them not to do so now.
We must congratulate our printers on the settlement of the industrial
troubles they have been going through, and we hope that the year 1928 will
witness the Journal issued with more regularity than it has been in 1927,
OBITUARY
CHARLES ECKFORD LUARD
With deep regret we record the death of Lt.-Col. C. E. Luard,
C.I.E., at his residence, ‘ Woodsend ’, Boars Hill, Oxford.
Lt.-Col. Luard, a son of the late Col. C. H. Luard, R.E., was educat-
ed at Marlborough and Christ Church, Oxford. Hecame out to India
in 1892 and after six years of service with the 8th Gurkhas was trans-
ferred to the Political Department as an assistant to the Agent to the
Governor-General in Central India. The major portion of his service
was spent in the Central Indian States and his long official connection
and close study of these States made him perhaps the foremost
authority of our times on this important group of Indian States.
Although more of a‘ Savant’ thana ‘ Political’ Col. Luard was
esteemed and respected by all those he came in contact with both in
his official and private capacity. He gained the warm regard of the
ruling princes and was quite as popular with his European colleagues.
Col. Luard wrote the Azstory of the States in Central India and
also compiled a valuable bibliography of that history. ‘The poriion
of the J/uzperial Gazetteer dealing with Central India was prepared
by him ; while his recognized intimacy with the history, people and
rulers of Central India led him to be deputed on three successive ~
occasions, viz., 1901, 1911 and 1921 to prepare the decennial census
reports. Col. Luard joined the Society in 1905 and was a frequent
contributor to the pages of its journal. In the present issue of the
Journal we publish the last of the contributions received from his
pen—‘ The Ornithology of the Bible’. Cther articles by him of a
Similar nature, ‘Deer Hunting in Shakespeare’s Day’ and
‘Shakespeare onthe Noble Art of Hawking ’ reveal his leaning
towards this rather fascinating line of research.
Col. Luard, who was made a C.1.E. in 1923, married in 1903
Alice Margaret, daughter of Mr. James Todd of Aberdeen and
leaves two sons and a daughter.
MISCELLANEOUS NOTES
I.—READERS’ VIEWS ON ‘GAME PRESERVATION
IN INDIA’
You have started a discussion in vol. xxxi of the Journal, Nos. 3
and 4, and have invited others to give their views on ‘ Game preser-
vation’. There can be no doubt that the preservation of Indian
Fauna is a question requiring immediate attention.
In your article in No. 3, you go straight to the root of the trouble
when you point out that it is the fact that game trophies, whether
hides or horns, are allowed to be sold by law, which is mainly
responsible for the heavy decrease of game in most districts.
There are two points I should like to touch on. You sum up as
follows :—
‘The impression gained from the letters read is that what is
principally needed is a law forbidding the sale of any part of a big
game animal.’ So farso good. I heartily agree with this ; dut¢ you
go on to say ‘ carnivora excepted ’.
Point No. 1. Why should Carnivora be excepted? I may say
without fear of much contradiction that carnivora, i.e., tiger, panther,
bear, afford just about ¢ke most exciting sport of any. Why then
should they be excepted from alaw which it is proposed to introduce
in order to preserve big game for sporting as well as for Naturai
History purposes ?
Another potent argument in favour of not excepting them from
sale is this: Big game sportsmen do not (at any rate the true sports-
men certainly would not) ever sell any of their trophies, least of all
tiger, panther or bear. It therefore follows that carnivora skins
soid must be those killed either by poachers or by people who are
not sportsmen at all but who merely shoot carnivora as matter of
business, when they have the luck to be placed in the middle of
the tiger and panther jungles, and are thereby selfishly depriving
real sportsmen, who have probably spent a lot of money in getting
to their blocks, of a chance of enjoying well-earned sport.
I therefore strongly urge that if any law is agitated for to be passed,
it should be made to include carnivora, or at any rate, tiger.
Point No. 2. You go onto say ‘Save by a Forest Officer in the
public interest.’ This requiries, I think, some elucidation. Per-
sonally, I would not leave any sort of loophole if the Jaw is to be
really effective. So many good laws have been rendered ineffective
by some such weak clause.
Let us examine this second exception. In what exact public interest
would it be to sell skins of any game animal, or horns? If it refers to
shed horns, I fear that it would be only too easy for forest guards
and coolies or othersto shoot animals for food and produce their
horns as picked up shed ones. I have not come across or heard of
any district where horns and hides lie about in such profusion that
it would be worth while collecting, advertising and selling them,
24
982° JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Volos
The amount realized I should say would not be commensurate
with the labour and time of the forest guards wasted in searching
the country for them.
I have recently been in one of the best districts in India for chital,
hog and barking deer. Though of course they had shed their horns
long ago, Chital were to be counted in herds of 200, swamp Deer I
saw in herds of 50 and 60, hogdeer in herds of 20(I counted 23 in one
herd, which must be very unusual) and Barking Deer 5 and 4 together.
Although I was moving all over the block for fifteen days all day
and every day, | picked up 1 chital head and 1 horn of a hog deer.
I should be very glad if you can:let me know what steps are being
taken to get Government to pass legislation on this important
subject.
Turning now to Mr. Brayne’s suggestion, in No. 4 of vol. xxxi,
page 1020. I certainly think something of the sort should be started.
The syndicate idea, Mr. Brayne says, has already been turned down
by the Forest Department, so I need not refer to it, except that
one should like to know their reasons ? As regards his second sugges-
tion, I think that a Game Preservation Society would be a first-rate
scheme ; a definite organization like this would be of immense value
in many ways.
For one thing, here you would have a big body of big game
sportsmen which would, by fear of public opinion, greatiy
curtail the present shooting of wretched immature sambur,
chital bucks, etc., etc. if it did not actually get legislation
passed on the subject. It has always been a matter of astonishment
to me that there are such excellent rules and penalties in the -
Kashmir Game Preservation Department and yet there are none in
British India. The most that forest shooting-block passes do is to
limit the number of heads one may shoot Nothing is said as to
size, an equally important point if not moreso. I have met young
sportsmen in blocks who had not the vaguest idea as to what size
a shootable sambur or chital was. The Kashmir Game Laws are
to my mind the model of what Game Laws should be.
The Game Preservation Society, again, would in time be able to
place the best material as watchers in the forests, with heavy
penalties (such as loss of pay and dismissal) in case of bribery by
poachers, etc.
Again, the Society would be able to assist its members in the
many ways which a sportsman requires and which are now denied
to him, unless he be a highly placed officiai such as the Governor of
a Province, or unless he be a Policeman, Commissioner or Forest
Officer. This particularly applies to the U.P., as only the above-
named people can obiain elephants or beaters and other assistance.
The ordinary sportsman (military or business man) is tremendously
handicapped if ke has to deal with a wounded tiger or panther. In
fact, it is 100 to 1 against his getting it. Recently J had to leave a
wounded panther (a very big one too) in my block, as I could not
secure the services of any men, elephants or buffaloes to follow it
up. I followed it up alone for hours, a most hazardous game, as
one had to force one’s way through thick long grass (over one’s
head) as quietly as possible, an very exhausting and dangerous
“VUdaZ V ONINOVLILVY NOIIT ONNOA
‘009 ‘LSIF] ‘LVN AVaWOg ‘Nanor
WISCHELANEOOS NOTES : 583
business. I had to give upin the end, leaving the panther as a potential
source of danger to any forest people or others in the neighbourhood.
Had I had an elephant, I could have followed the panther all day if
necessary and of course with vastly increased chances of seeing
it, looking down as one could have done into the grass. I may say
here that I tried my best to get even one elephant before entering
. my block, but neither the civil authorities, nor the Forest Department
could (or would) assist.
AJMER, W.M. LOGAN HOME,
July 11, 1927. Major,
4th Bombay Grenadiers.
[Although the extinction of the big carnivora would mean an
increase in those animals which, when in large numbers, area
curse to the agriculturist, we do not think the prohibition of tiger
shooting altogether by non-license holders or by any except sports-
men would be practical politics. It might be necessary—we can
imagine several instances when it would be in the public interest—
for the Forest Officer to shoot tigers and it would certainly be in
the public interest for him to seize tiger shot contrary to law.
Possessing the trophies, would it not be in the public interest for
him to sell them for the benefit of the public? Eps.]
Il.—A UNIQUE LION PHOTOGRAPH
(With a plate)
The accompanying plate of a young lion attacking a zebra is
perhaps the most unique photograph ever obtained of a lion in his
native environment. The photograph came into my possession
through the late Mr. C. A. Latif, a prominent citizen of Bombay,
who had resided in Zanzibar for a considerable number of years
towards the end of the last century.
From the particulars furnished by this gentleman it would appear
that the photo was taken by an engine driver on the then newiy
opened Uganda Railway, which as is well known, even now runs
through some of the finest game country in the world. The engine
driver got his camera ready to photograph a herd of zebras as they
stood a short distance from the line, gazing in wonderment at the
weird and unfamiliar object that had invaded their territory and rattl-
ed so glibly along the iron rails. The instant he pressed the shutter a
young lion sprang from the surrounding scrub on one of the zebras,
and this was the resultant exposure! If I have understood
correctly, Mr. Latif was a passenger on this very train, but even if
he were not, the driver was known to him personally and it was he
that presented Mr. Latif with a copy of this remarkable photograph.
The coincidence and the circumstances sound positively mira-
culous and I should hardly have credited the account had it come
from a less reliable source.
The photo was in Mr. Latif’s wonderful private collection of
curios collected all over the world and to my mind there appears no
reason to doubt its authenticity and genuineness
584 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXX11
Strangeiy enough | find the identical photograph considerabiy
touched up and reproduced in Hutchinson’s ‘ Animals of all Coun-
tries’ (part iv, p. 173). All that is there mentioned about it is ‘ By
permission of E. N. A.’ ;
It would be very interesting to learn who ‘E.N.A.’ is, how the
photograph came into his possession and if he has another version
as to how it was secured by the camera-man.
BoMBAY NATURAL History SOCIETY, SALIM A. ALI.
6 APOLLO STREFT,
September 5, 1927.
IIl.—WHITE TIGERS
(With a photograph)
I have recently had the luck to shoot a white tigress and would
like to know if any other members of the Society have ever done
so. I may mention that, not long after, a lady, Mrs. Murphy, wife of
Mr. P. W. Murphy of the I.C.S. of this Province, also had the good
fortune, when out with me to bag another one, also a tigress, and
very much the same colour asthe one I got. ‘The animal I shot
was pure white with black stripes on her body and russet brown
ones on the tail. Both were well grown beasts, thick set, and taping
respectively, 8’-54” and 8’-2” between uprights. They were shot in
the same jungles, but about twenty miles apart, in the district of
Bhagalpur.
I enclose a photo of the animal I shot for publication in your
journal if you iike. The taxidermists to whom the trophy was sent
report that during the year 1926 they received three white tiger
skins including mine (mine was shot on December 6, 1926), but
my skin is the only pure white one, the other two being cream
coloured. |
MISCELLANEOUS NOTES 585
The animal! was not an albino, as it did not have pink eyes, but the
ordinary coloured eyes of a tiger, and so did Mrs. Murphy’s. Is it
possible that there is a distinct breed of white tigers, or are they
only freaks? I have for years past been told by my shikaries that
there was a white tiger inthe jungles near here, quite twelve years
ago. Herders who saw the animal, described it as ek dum gatke
mitik satéd (white like acow). Inever believed them till I shot
the tigress which must be a descendant of the white one seen by
the herders years ago. Both the tigresses were young animals,
about three or four years old. I once did see a white animal pass a
little distance from a kill which I was sitting up over, and at the
time thought it must have been a hyzena. It was bad moonlight and
in thickish jungle, but I was rather suspicious at its making no noise
in the leaves, which a hyzena usually does with its clumsy tread. I
now think it must have been a white tiger. Both the above
tigresses were shot in a beat, and took a good deal of lead. I hit
my tigress from about twenty yards away through the right shoulder,
bowling her over; she however managed to scramble away, and I
missed with the second barrel. I followed her up soon after for
about 100 yards, but did not find much blood, and had to give up
for that day as it was getting late. The next day I found her
fully 1000 yards away from where I originally hit her and
polished her off with a shot through the neck. On skinning her I
found that my first shot had gone through the shoulder and the
bullet or rather parts of it were found under the skin on the opposite
side, her liver being in shreds. Surely it is exceptional for a
wounded tiger with such a fatal shot to go so far. The bullet used
was a Contractile from a smooth bore (12-bore).
I shall be much interested to hear what information you can give
me in regard to white tiger.
F. B. ROBINSON,
SAGRAMPUR,
BIHAR AND ORISSA,
January 28, 1927.
[Mr. Robinson’s tigress is yet another of the many examples of
complete or partial albinism recorded among tigers in recent years.
Partial albinism is defined as a condition in which certain areas of the
body remain marked by more or less well defined areas of colour
owing to the presence of more or less pigment. It is believed that
a pigmented animal owes its colour to the power that certain tissues
of its body possess to secrete tyrosinases (oxydizing ferments) and
chromogenic substances—the colouration being the result of the
action of the ferments upon one or more of the chromogens and the
theory is therefore put forward that an albino is an individual
whose skin lacks the power to secrete either the ferment or
the chromogens—it may hold one of these substances, never
both.
But the presence of both these colour determinants are not alone
sufficient to explain the development of colour which is arranged in
animals in a definite pattern, either self, spotted, striped, etc., and it
586 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
is believed that colour is correlated with some other factor which
determines the pattern—each speciai pattern having its own deter-
minant and it has been shown that, while an albino may not possess
the factors which determine colour, it may still possess factors which
determine pattern, both visibly in the body tissues or as a gametic
character ; when its presence will oniy be detected in subsequent
generations.
The frequent records of white tigers in recent years has been
remarkable. On looking through our Journals we find no tess
than seventeen distinct records of white tigers shot in India
during the last twenty years—a period during which certainly less
tigers were shot than formerly if we are to credit the records of the
old Sporting Magazines. The question arises as to whether these
frequent records can be looked upon as instances of variation tending
to the formation of a new variety orrace. If there is anything in the
theory of protective coiouration in reference to the tiger, then the
white colouring would be an obvious disadvantage militating
against the establishment of such a race, and yet these tigers appear
to thrive quite as well as the normal coloured animals from the fact
that their records are becoming more and more frequent.
Do white tigers breed white tigers is a question on which natura-
lists and sportsmen are, as in the case of black panthers, inclined to
differ. The white tigers of the Rewah State would appear to
belong to the same family and to have bred white for several
generations. Mr. Robinson assumes his tigress to be the offspri-
ing of white parents. The black panthers in the Kolhapur Zoo have
bred black panthers for several generations. EDs. ]|
IV.—A TUSSLE BETWEEN TIGER AND BUFFALO
BAIT
Whilst shooting in the Central Provinces in February this year,
a buffalo calf, tied up for a tiger bait, showed signs of a struggle
with a tiger when visited inthe morning. Unfortunately, having
injured my leg, I was unable to go and see that calf but several
independent witnesses and a reliable shikari were all questioned and
no discrepancies occurred in their answers.
The calf bore marks of claws and teeth all over its back and,
though it had subsequently to be destroyed, was still alive. Its
horns were covered with blood and tiger’s hair and it must have
given nearly as good as it took.
A tussle between a 24 year old tethered calf, and a tiger strikes
me as rare; the tiger may have been frightened off by passers by,
but even so the tussle lasted sufficiently long for the tiger to inflict
a large number of wounds and the incident is an example of the
stubborn courage of the domestic buffalo.
UNITED SERVICE CLUB,
Par MAtt, S.W., J. Wc. CCE IVE
June: 7.1927. Captain, M.C,
JOURN. BomBay Nat. Hist. Soc,
A TIGRESS AND ITs KILL.
MISCELLANEOUS NOTES 587
V.—UNUSUAL BEHAVIOUR OF A TIGRESS WITH LIVE
BAIT
(With a plate)
I have recently been shooting in the C. P. with Capt. Graham of
this Regiment. We found we had to deal with a tigress which
would not return to her kills, so we took it in turns to sit up over a
live bait. On the third night the tigress attacked the bait at about
12:30 a.m. Capt. Graham saw the tigress standing firmly with all
four feet on the ground and holding the side of the buffalo calf’s
throat in her jaws. In this position she pinned the animal’s head to
the ground.
It was in this position that Capt. Graham shot the tigress, which
fell dead on the spot.
In the morning I went out and photographed it, and I now enclose
the prints, which may possibly interest you or some of the readers
of your Journal.
THE RIFLE BRIGADE,
LANDIKOTAL, INDIA, : F. O. CAVE,
Sune Ss. 1927. Captain.
VI.—WHAT IS THE RECORD PANTHER?
I read a remarkable statement which I expect a good many other
members of the Society will also have read, in the issue of the
Sporting and Dramatic for July 6, 1927, as follows :— Rowland
Ward’s Records gives the length of a leopard killed in S. Nigeria
before skinning as 9 ft. 3 in. while the record Indian specimen
is one from Kashmir measuring 9 ft. lin. The average of a good
Indian panther, before being skinned ranges from 7 ft. 9 in. to
8 ft. 6 in.’ Now this statement is absolutely contradictory to
all previous records, observations and articles appearing in any of
the Society’s Journals and also in any standard work on Big Game
Shooting. I refer readers particularly to Col. A. E. Ward’s able
and authoritative articies in vol. xxix, No.1, page 25 and vol. xxxi,
Now) pace 3.
As I have quoted the Storting and Dramatic article, I propose
to do the same with Col. Ward’s observations.
‘There is no doubt that an animal 8 ft. length measured between
‘pegs, or even by taping from the nose between the ear and then
‘straight to the tip of tail, is very rare.’
Col. A. E. Ward then gives a table of record leopards the biggest
of which is 7 ft. 54 in.
Again ‘A full grown male leopard may be from 6 ft. 9 in. to
‘8 ft.; an eight-foot leopard measured between pegs is very un-
‘common. In Rowland Ward’s Records of Big Game a measure-
‘ment of 8 ft. 3 in. is given and 8 ft. is recorded of two specimens
‘at least; and then we have 7 ft. 6in., 7 ft. 54 inches.’
With all the above I most emphatically agree and all sportsmen to
whom I have spoken on the subject have the same opinion. It is
therefore most difficult to understand the statement made in the
Sporting and Dramatic and I suggest the Society might collect
588 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
further opinions on it and at the same time ask the Editor of the
Sporting and Dramatic to give further particulars, such as
(1) Who gave the information regarding these gigantic brutes
of leopards (bigger than any tigress ever known and
equalling most average tigers !)?
(2) Who actually measured them and how ?
(3) What other witnesses were present ?
(4) The date and place where shot or otherwise killed ?
5) Where are the skins now and what do they now measure?
(6) Will the owners very kindly allow two or three well-known
big game hunters to view the skins?
In this connection I might observe that the dressed skins would
normally measure, for the 9 ft. 3 in. one = 10 ft. 10 in. and the
Y ft. 1 in. = 10 ft. 8 inch. As regards the Kashmir one it is
difficult to understand how Col. Ward did not see or even hear of
it. Is itin the Kashmir Game Records ? This point could be verified
easily. If not, why was the owner so very retiring and modest
after shooting almost the world’s record panther ? !!!
As regards the statement that the ‘ average of a good Indian
panther before being skinned, ranges from 7 ft. 9 in. to 8 ft. 6 in.’,
this is frankly incredible and gives a totally erroneous idea which
I am sure the majority of big game sportsmen will not agree with.
AJMER, W. M. LOGAN HOME,
August 2, 1927. Major.
VII.—A PANTHER AND FLASHLIGHT
(With a plate)
This photograph of a panther was obtained in the foothills of
Garhwal. The panther was fired at from a machan when it was just
too dark to aim properly and within half an hour came back to its
‘kill’ in the open river-bed,!00 yards from the machan. It set off the
flashlight, which makes a noise like a grenade, and yet returned and
ate most of the ‘kill’ during the night.
Panthers in this part of Garhwal always return when they are fired
at and missed. If, however, on their first approach they see anyone
in a machan they will remain close by for an hour or so and then go
away and do not return even when the shzkarz has gone home.
BA. S.- BRUNSKiGE,
Major.
VIII.—BOLDNESS OF PANTHERS
I was out one morning alone after a Chital and was returning
at 7.30 a.m. along the edge of the jungle. To my right was
a vast expanse of open grassy plain known locally as a Chauri.
I saw some animal a long way off moving from left to right
out into the plain, when I got the glasses on to it, I saw to my
surprise it was a fine panther, its coat glistening in the bright sun.
I at once sat down and watched. It continued to walk along quite
unconcernedly getting further and further out into the open plain.
I guessed that it was making for a kill and this proved to be the
(‘stoqrenb-pulg s,aeyyueg oyy caoqe ysn{ savedde asou s,Mog oy)
;« TTIM,, MOO V GNVY AAHINVd V
‘009 ‘LSIFT ‘LVN AVaNWog ‘Nunor
MISCELLANEOUS NOTES 589
case. It disappeared into some rather higher grass and then
started returning with an animal in its jaws. I then commenced
running, crouching through the grass, which was only knee-deep and
managed to get within 300 yds. without the panther taking any notice
though I was in full view more or less the whole time. I then saw
that he would soon reach the edge of this jungle and so I took my
chance and fired, and missed (the bullet went high and behind him).
He dropped the kill and bounded off into the jungle. I got up and
walked about 50 yds. when I saw him coming stealing back. I at once
got into a sitting position again and fired when he had stopped to
pick up his kill; but again missed. This time I ran crouching as
hard as I could and made about 80 yds. and then sat down and
hardly got into position when he again came stalking tack out
into the open, picked up the kill and walked unconcernedly off.
I fired and this time hit him. He staggered off and when I reached
the spot, I found it was a full grown Kakur he had been carrying.
I went back to the bungalow and got three men and tracked the
animal. I think this is a remarkable case of boldness and fearless-
ness on the part of this panther.
On another occasion, a few days later, I sat over a fresh kill (a cow)
which had been dragged over 400 yards. I wasin position at 6 p.m.
and my manleft me at that time. At 6.10 a magnificent panther
arrived, quietly walked up to the kill, never looking to right or left,
and at once seized it in his powerful jaws and began dragging it off
backwards. I got him through the back of the neck and he died
then and there, biting his left fore arm in his jaws. My man had
only gone about 4 mile and in fact was still walking away when he
heard-the shot. The panther must have been sitting waiting within
earshot and close indeed.
While on the subject of panthers, I should like to protest
against a statement I read ina big game book recently that ‘ the
panther looks what he is, a perfect swine, while a tiger always looks
agentieman!’ I think that is a remarkable statement, and person-
ally I (and a great number of sportsmen agree with me) have
exactly the opposite opinion. In view of my recent experience
in the Terai, I consider a panther is a most sporting animal and
gives one a very good chance, whereas a tiger is a skulking
nocturnal beast, who has to be driven out to the guns ; he very rarely
ever comes to a kill in daylight, and curiously enough the book
I mentioned brings out this very point.
A tiger more often than not will not return to its kill even when
not tampered with, whereas the sporting panther comes back to his
kill even when one has dragged it 100 yards and often to only a
head and a few bones. Such is my experience and it is the
experience of a great many more experienced shikaris than myself.
Take again the case of man-eaters; far more tigers become man-
eaters than ever panthers do. I wonder if the author of that
statement considers man-eating tigers as gentlemen! !
AJMERE, W. M. LOGAN HOME,
July 8, 1927. Major,
4th Bombay Grenadiers,
ra
590 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
[X.—CIRCUMSTANTIAL ACCOUNT OF A PANTHER
ATTACKED BY WILD DOGS
The following may be of interest to members :— |
Whilst shooting in the Central Provinces a few years ago a friend
of mine found a panther in a dying condition. When dead, the
panther was brought into camp and examined and skinned. The
animal had been emasculated, the wound thus formed including the
anus, no other wounds being found.
On removing the skin it was found that practically the whole of
the entrails had been drawn out through the wound.
The only possible explanation was that this was the work of
Cuon; both my friend and his brother, who was present, are ex-
perienced shikaris and their veracity cannot be questioned.
The panther may, of course, have been in a feeble condition
when attacked but even so the occurrence is remarkable.
I cannot think of any other possible explanation. My informant
emphasized the fact that the panther was alive when found.
UNITED SERVICE CLUB, J: Me. C. CLIVE, mics
PArTAMALL, oO. Wels Captain.
June 16, 1927.
[In vol. v, p. 191 of the Journal, there is an account of a pair
of panthers being treed by wild dogs. The two panthers were in a
‘ Sallai’ tree, one above the other with a large pack of ten or twelve
couple of wild dogs moving about below. A similar instance is
recorded in vol. xviii, p. 194 of the Journal, in which a pack of wild
dogs was seen in pursuit of a female panther. R. C. Morris in a
note on wild dogs, vol. xxx, page 691, says he saw the remains of a
panther that had been killed by wild dogs. It seems well establish-
ed that panthers are occasionally attacked and killed by wild dogs—
bears have also been attacked—while the belief is current that even
tigers are not free from attack from these vermin—but the cases so
far recorded in reference to tigers are based mainly on hearsay
evidence. As regards the dogs having emasculated the panther,
see Note No. XI on the opposite page. Eps. |
X.—WILD DOGS
‘here has been some correspondence about wild dogs lately.
It has been alleged that there are two species (which I do not
believe) and mention has been made of the colour of the tip of the
tail. I have just received from the American Museum of Natural
History, New York, three specimens, about two-thirds grown which
were shot from the same small pack by Kunwar Dillipat Shah of
Khairigarh. One of these has a black tip to the tail, another has
a few white hairs, and the third has a distinct white tip to the tail
like that of the silver fox. These three dogs were presumably of
the same litter of puppies.
ORIENTAL CLUB, J.C. FAUNTHORPE,
LONDON, Colonel,
September 3, 1927.
MISCELLANEOUS NOTES 591
[Only two species of wild dog have hitherto been recognized :
Cuon dukhunensis, the wild dog of the whole of India and Cuox
rutilans, the wild dog of Burma andthe Malay countries. There is
however a belief prevalent in India from Kashmir to Cape Comorin
that there are more than two species and that in addition to the
‘Jangli Kuta’ of the plains there are two hill kinds, one larger and
the other smaller—the small breed is said to limit its range to the
low ravines and to restrict its attacks to sheep and goat while the
larger breed hunts in the higher mountains. It has been suggested
that the smaller breed is really a pack of females and cubs, the
mother training the young idea in the arts of hunting and naturally
blooding them on easily killed quarry, while the full-grown packs
roam the forest attacking what they may. Ebps.]
No. XI.--HOW WILD DOGS KILL THEIR PREY
A recent issue of the Journal, vol. xxxi, page 813, contained an
account of a sambhar stag in Burmah having been emasculated by
wild dogs and the affirmation of the local shikaries was that this was
their invariable practice. As the article implied a contradiction of
my remarks on the subject in ‘ Wild Animals in Central India’
which was referred to, perhaps you will be good enough to publish
what I believe to be the explanation of this widespread belief.
Anyone familiar with the East knows of the exaggerated impor-
tance attached to anything connected with the organs of
reproduction. The very idea of emasculation is horrific ! Occasional
and isolated instances of this having taken place are discussed with
bated breath, possibly dwelt upon with a lascivious satisfaction.
That emasculation ‘may’ take place immediately becomes that it
‘does’ take place. The mind of the jungli native construes
this as a dreadful and purposeful action on the part of the wild
dog.
Now for the fact that emasculation occasionally takes place and
the explanation thereof. When an animal is attacked at bay the
dogs launch themselves on the head and seize it clustered like ticks
on every conceivable place. If the pack happens to be a large one
there is no room on the head for those diffident in attack and so these
immediately attack the hind quarters. This is the normal procedure
generally of a pack of dogs. It constantly happens when a bobbery
pack attacks a jackal or a pie dog. The victim is seized at both
ends and stretched.
It would be surprising if under such circumstances some dog did
not find the scrotum and testicles of any stag a most convenient
mouthhold and that before the struggle ceased these did not
occasionaliy become detached. That emasculation is either a
common or a deliberate act I can deny from a wide personal
experience of the wild dog and its habits.
I have now explained how this operation is sometimes performed
fortuitously and I have suggested how an ‘occasional occurrence ’
has been converted into an ‘invariable practice’ in the natives’
mind.
592 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Anyone reading Wzld Animals in Central india will see that I
state as follows :—
‘The story to the effect that they (wild dogs) are in the habit of
emasculating the animals they attack can be discarded’’. This
sentence was purposely worded so as to make it clear that although
emasculation might sometimes take place it was not habitual. The
habit is denied, the act is not.
‘ Ivy Bank, ELGIn, A. A. DUNBAR BRANDER.
SCOTLAND,
August 14, 1927.
[The belief that the wild dog emasculates its quarry is very
prevalent in India, as also is the belief that they sprinkle bushes with
urine and drive their prey through them and then rush upon their
quarry which is blinded by the acid fluid! Emasculation does occur
but is, as pointed out by Mr. Dunbar Brander, the result of chance
and not of deliberate effort. Eps.]
No. XII.—THE RECORD FOUR-HORNED
ANTELOPE (7TETRACEROS QUADRICORNIS.)
(With a photo)
In February of this year we received avery fine head of this
animal for mounting, shot by Col. E. Conway-Gordon in the Central
Provinces. On examination the rear horns proved to be far above
the known record and the fore-horns also exceeded those of the best
head.
F,
BERD. E
Journ., Bombay Nat. Hist. Soc.
a
ABNORMAL BLACKBUCK SHOT IN BIKANER.
woe aS
oS OS a
A Goop CHINKARA HEAD FROM BIKANER. HORNS, 143”.
MISCELLANEOUS NOTES 593
The measurements were taken when the head arrived and then
again at the time of writing, when the head was completed and
thoroughly dry. It will be seen from the table given below that
there has been a considerable amount of shrinkage.
| Length of | Length of fore Circ terenge.
| rear horns. horns.
pean ane Lae
| R. | i | R. aes (o Rear. Fore.
|
® 34 1” 3m 4
On arrival tale fe gen Te 23 3 . a
On September 9, 1927 ee Oe aly OFS 1h 247 26 | 28" Je"
| |
The right rear horn is broken as will be seen from the accom.
panying photograph. The core of this horn travels right up its
length and measures 34 inches. The core of the left rear horn on
the other hand is very short being only 1# inches.
The rear horns spread outwards and owing to their being broken
the tip to tip measurements cannot be taken. The front horns
converge inwards and the tip to tip measurement between them
is one inch.
The best head in Rowland Ward’s ‘ Records of Big Game’ (1914)
measures: rear horn 44” : fore horn 23”,
BoMBAY, C. McCANN.
September 9, 1927.
XIII.—ABNORMAL BLACKBUCK AND A GOOD
CHINKARA HEAD FROM BIKANIR
(With a plate)
I am enclosing two photographs which I trust will be of interest
to you. The blackbuck carries an abnormal head, as is clearly seen
from the photograph, and was shot by His Highness the Maharajah
of Bikaner in the north of the Bikaner State at Hanumangarh. The
chinkara, which measured 144”, was shot in the north of the State
near Ganganagar by the Maharaj Kumar Sahib. It is practically a
record for Bikaner ; only one bigger head measuring 144” is known
to have been shot in the State so far.
BIKANIR, SURAJMAL SINGH,
RAJPUTANA, Secretary to
August 31, 1927. Svt Maharaj Kumar Sahib.
XIV.—CLICKING NOISE MADE BY MUNTJAC
I remember having read in some of your Journals different articles
attributing the curious clicking sound made by the Barking Deer to
various causes. While at the local Zoo sometime ago Iwas watch-
ing the Muntajc in the enclosure, when I suddenly heard the buck
making this peculiar noise. The little animal was on the other side
594 JOURNAL, BOMBAY NATCRAL AIST. SOCIETY, VolVXXxods
of the railing to me andnot more than about a yard away. It was
moving to and fro parallel with the railing and clicking away merrily.
I sat down and watched it most carefully for along timeand found
that the noise was distinctly produced by the animal’s mouth. The
clicking noise was made with the tongue and the lips at the same
time, much like aclucking hen. The mouth was not open but every
time the tongue clicked the ends of the lips were drawn back. I
asked the keeper, and he informed me that this often occurred and
was a sign of pleasure. Recently Mr. C. Raitt of Faith Estate,
Kutta P.O., S. Coorg, was with me at the Zoo and he also heard the
same noise and said it was made by the mouth.
I have seen it stated by some that the noise was made by moving
the tushes in their sockets. This is a physical impossibility, because
the tushes, although set loosely in their socket are yet imbedded ina
cushion of muscles and flesh; which excludes all possibility of any
clicking sound being produced, though the tushes could be moved
about to a certain extent by external means and not by its own
power. Whether the doe also makes this clicking sound I am
unable to state, but I do not see why she should not be able to.
I have also read that the clicking noise was made by the hoofs
but I do not think this is correct.
MyYSsoRE, CHARLES THEOBALD,
5S. INpDIA, | esas)
September 2, 1927.
XV.—THE BRAY OF THE CHITAL STAG
My attention has been drawn to Miscellaneous Note No. III, page
694 of vol. xxx, No. 3 (which I have somehow missed seeing
before). I refer to the note on the Chital.
Mr. Morris says, ‘I would not describe the call (szc) as in the
nature of a whistle at all.’ I presume he refers to the rutting season
‘roaring’ of the stags; personally, I quite agree with him. ‘The
chital stag’s voav in the rutting season is, at any rate, from the
Siwaliks to the Terai, very much the same as any other stag’s roar,
and certainly the Terai and Siwalik stags’ roar could not be called a
whistle, whereas their alarm note zs distinctly a kind of whistling
cry, which the does also utter. To my mind the chital stag’s roar
resembles the Kashmir Stag’s and Scottish Stag’s roar only it is
not so loud.
AJMER, W. M. LOGAN HOME,
August 25, 1927. Major,
¢th Bombay Grenadiers.
XVI.—MUSTH IN ELEPHANT
Col. G. H. Evans in his well-known book Elephants and their
Diseases opens his paragraph on musth with the words ‘ Male
elephants and very rarely females, on attaining maturity, are subject
to peculiar periodical paroxysms of excitement .. . to which the
name of musth is applied. .. .’
MISCELLANEOUS NOTES 595
In view of the above statement by such a well-known authority, it
may be of interest to your readers to know that during the Depart-
mental Kheddah Operations conducted by the Forest Department
in the Goalpara-Bhutan Mahal, Assam, 1925-26-27, two female
elephants exhibiting unmistakeable signs of musth have been caught.
The cases are as follows:
No. 1. A very large female elephant of the Kumeraband type
was caught in November 1925. This elephant had very large
tushes, and at first, from her size and build, might have been taken
foramakhana. Although full grown she had never had a calf, and
there is no doubt that she was barren. While in the stockade she
gave a good deal of trouble and attacked her fellows and the
koonkies, and eventually broke both of her tushes on a tree inside
the stockade.
This paroxysm of fury while in the stockade was succeeded at the
depot by an impassivity and ‘ don’t care’ attitude which was quite
as remarkable. The temporal glands were greatly swollen and
gave forth a profuse oily discharge. Her training proceeded with
remarkable rapidity and she was able to go out without koonkies ten
days after her capture. Shortly afterwards, however, symptoms of
blood poisoning were observed, due to the severe wounds the
broken-off ends of the tushes had caused in the mouth and the
animal was released in an effort to save her life.
No. 2. A very large female elephant of the Kumeraband type
caught in November 1926. This elephant was not of such fine’
build as the former and in addition had had a calf. Her temporal
glands were swollen and gave forth a copious discharge. She
seemed very drowsy while at the depot but her training proceeded
without incident and she left for her owner’s home about a month
after capture.
The question as to what causes the condition of masth in
elephants can be ascribed, is one that has not been Satisfactorily
answered. It is true that some bulls when in this condition court
the society of females, but it is by no means an invariable rule and
it cannot be dogmatically asserted that this condition is due to a
lack of sexual gratification.
As regards both of the cases cited above they did not show any
perceptible desire for male company at the time nor did tuskers or
maknas at the depot exhibit any interest in them.
One cannot lay down conclusions from two cases oniy, but it
seems clear that the symptoms of the condition in the male and
female is exactly the same. First the violent stage in which all
animals are attacked and pain does not seem to be felt. This is
succeeded by a drowsy stage in which the animal takes its fodder as
usual but seems oblivious of its surroundings, and the attack then
appears to gradually wear off. In both stages the temporal glands
are swollen and give out a characteristic discharge.
The first case apparently was captured in the earliest stages and
the second in the latter stages.
Both these animals were caught in the month of November
and in stockades which are a few miles off from one another.
All the other numbers of the respective herds were normal.
596 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Some years ago an Assamese gentleman, Rai Sahib Rajant
Kanta Chowdhury of Sorbhog owned a female elephant which was
locally known by a term which was the equivalent of ‘ female
tusker.’ This was a North Cachar elephant and owed its sobriquet
to the fact that it had longish tusks and went musth regularly each
year until its death recently.
Elephant attendants have told me that male elephants go musth
up to an advanced age, in fact none of them could recall an
elephant that had not gone musth even in its old age.
We shall not probably get an explanation of why female
elephants sometimes exhibit these secondary sexual characteristics
of the male until some anatomical light is shed upon the question.
The above lines are written in the hope that they may catch the
eye of those interested in the problem and that some information
leading to its elucidation may enlarge.
KOCHUGAON, INA eb OR:
ASSAM, Imperial Forest Service.
TARUATV 9-1 927;
XVII.—A LARGE TUSKER ELEPHANT
The following are the measurements of the tusker elephant I
wrote you about :—
Height at shoulder 10ft. 44”.
Length Girth Weight
Richtinsk..6 24 21% 85 lbs.
Left SER Go 6 ZEY 86 lbs.
Total pec WR elo:
The above were taken in the presence of witnesses and certified
measurements were sent to Rowland Ward. What struck me about
the tusks was their girth and symetry—a perfect pair! As regards
height, this is of course difficult to get accurately, but it was taken
between uprights, and the least I could makeit was 10’ 45”; taken
at the highest point of shoulder, I made it 10’ 9”, but did not
think this possible after all Sanderson says about their being no
such thing as a 10’ elephant. Hewas 65” round the forefoot.
I think there can be no doubt of his being a record for South India.
Apparently the Senior Ranee of Travancore thinks so too, for all
my offers to purchase the tusks have been refused !
CHITTAVURRAI, A. W. JOHN.
ELLAPATTI P.O., via PERIAKULAM,
SouTH [NpDIA,
July 27 1927.
(There is anote in our Journal, vol. xi, p. 335, by Lieut. 5.8. Flower
giving measurements of tusks of Indian Klephants in the posses-
sion of the King of Siam. The largest of these measured 9’ and
9’ 104” respectively ; they belonged to an elephant whichis said
to have been ninety years old when it died at Bangkok and are easily
the record pair for the Asiatic elephant. Eps. ]
MISCELLANEOUS NOTES 597
XVIII.—A GOOD NILGIRI TAHR (HAMITRAGUS
FINAEOC FEI OS)
The following measurements of a Nilgiri Tahr, or Ibex, may
interest you as they constitute a record for this district (High
Range, Travancore). He was shot in May 1899 by myself :—
Length 164” (163” when shot)
Girth at base 82”
CHITTAVURRAI, A. W. JOHN,
ELLAPATTI P.O., Via PERIAKULAM,
SoutTH INDIA,
July 26 1IZT.
[A fine head ofa Nilgiri Tahr was recently presented to the
Society by Capt. H. R. Irwin. It was shot in the Nelliampathy Hills,
South India, and measured 164”, girth 94” (vzde Journal, vol. xxxi,
page 520. Eps. |
XIX.—BIRTH OF FREAK CALF AT INDORE
(With a photo)
With reference to a paragraph which appeared in the Current
Topics column of the Zzmes of India last month, the following
30 HOURS AFTER BIRTH,
26
598 JOURNAL, BOMBAY NATURAL. AIST. SOCIETY, Vol. XXXII
notes and enclosed photographs may be of interest to readers of
‘Miscellaneous Notes’ in the Society’s Journal.
Early in the morning of August 19 last, at the house of
Baddrudin in the Palassia Mohalla, Indore, a calf was born with a
bifurcated head, having two separate muzzles each with mouth and
nostrils complete. The head had four eyes, two of which, one on
each side, appeared to be normal, while the other two, joined to-
gether in the middle of the forehead, were sightless and ‘ dead’
looking.
The calf had to be hand fed as its head was too heavy for it to
lift. It sucked with both mouths.
Its birth, on Krishna’s Day (/anma Ashtamz), caused great ex-
citement at Indore, and thousands of people from the city and the
surrounding country came to see it and make offerings—much to the
joy of the lucky owner.
Of the three photos enclosed, one was taken when the animal
was about thirty hours old, and the other two (one of which is a side
view in which the head appears to be quite normal), when it was
eight days old. The calf died on the twelfth day.
LAL BaGH, INDORE, C. E. D. MEARS.
September 19, 1927.
[We are unfortunately able to reproduce only one of the photo-
graphs. Eps. ]
XX.—AN EXAMPLE OF AN ASSISTED PASSAGE
On April 26, 1926, I sailed from Bombay in the S.S. City of
Exeter which left about midday. About 3.30 p.m. I discovered
that we had on board a singie House-Crow (Corvus splendens) which
had apparently failed to notice our departure in time. It looked
rather bewildered at finding itself at sea, and grew more and more
uneasy as dusk approached, launching out over the sea and then
regaining the ship with difficulty. It was a dark dusky looking bird
obviously of the typical race from Bombay, and not the pale
zugmayert which is the form at Karachi whence the Czty of Exeter
had come.
The crow stayed with the ship for the next few days roosting on
the rigging and receiving food and water by the good offices of the
crew. We did not putin at Aden. We passed Perim on May 2
about 7-8 a.m. and after that I watched the bird making several
attempts to fly towards the mainland which was visible. Its
courage always failed however and it returned to the ship. By
evening however it had disappeared and I was told that it
had apparently flown on to another ship. Unfortunately this
ship was steaming in the same direction as ourselves, or I might
have told a good story about corvine sagacity intaking a ship back
to India! As an observed example of how stragglers leave their
habitat, this seems worth recording.
BATTLE, SUSSEx,
August 8, 1927, HUGH WHISTLER,
VR Al. Sa0 ahez-S
MISCELLANEOUS NOTES ba Fiat wh 599
XXIL_EXTRAORDINARY PUGNACITY OF THE RUSTY-
CHEEKED SCIMITAR BABBLER (POMATORHINUS
ERYTHROGENYS)
Last Sunday I went with a couple of friends for a jaunt in the car
on the cart road in the direction of Tara-Devi. As it was a
beautiful evening in the monsoons, we pulled the car up at the
seventh milestone to admire the wonderful scenery, and above all
the wonderful cloud effects that are only to be seen at this particular
time of the year. .
The country in the immediate vicinity is scrub oak. I happened
to glance up the khud side, when I noticed two of the above-men-
tioned birds settle in a small oak tree about 90 feet above us:
Keeping my eyes fixed on them, | noticed they were engaged in
fighting. In afew seconds they both overblanced, and fell to the
ground, and owing to the gradient of the steep hillside, they com-
menced rolling down the khud in our direction. So busily engaged
were they in their fight, that they paid no attention to the car they
were fast approaching. My friend Mr. Neville Cotton realizing he
stood a chance of catching the birds, jumped out of the car, and
stood with his hands ready open. On they came until they reached
a stone wall about 10 feet high, which separates the road from the
hillside. As they arrived at this point, they actually fell into the
hands of my friend. On examining them I found that they still had
no intention of giving up their fight, for one bird held the other’s leg
between his beak, while the other had a firm grip of his opponent’s
throat. It was no easy matter to separate them, anyhow after a
little careful handling I managed to accomplish this, when both birds
gave vent to their feelings by jabbing at my fingers. After their
wrath appeared to have cooled down somewhat, I released them both,
only to find they still had no intention of making friends, for all the
trouble commenced afresh on a bush, about 100 feet down the
khudside. A heavy mist soon obscured them from my view. Both
birds were in deep moult. It would be interesting to know what
all the trouble was about, and as I write this I wonder if they
are still atit?
THE MALL, SIMLA, ARTHUR H. BERRIFF..- -
July 25, 2927.
[As arule, the best of good fellowship prevails among the babbler
fraternity, though its members are susceptible to occasional
paroxysms of violent rage when two or more members of a ‘ sister-
hood’ will go for one another with the utmost zestand fury. We
have witnessed a somewhat similar incident to the one described by
Mr. Berriff, the belligerents in this case being a couple of Bombay
Babblers (7usdoides somervillec). An individual was seen furiously
pursuing another on the ground over strewn foliage, stones and
other impediments. They subsequently hopped on to a branch about
6 feet high whence they dropped in a tangle on the ground below
lying on their backs and continuing to grapple fiercely with each
other using bill and claws. We were able to approach near enough
to cover the birds up with a sola topee without attracting notice.
600 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
They made good their escape, however, but no sooner were Ane
out than the chase recommenced in all earnest.
A third bird here joined the fray and after the original opponents
had come to grips and fallen to the ground a second time, the new-
comer dropped perpendicularly on them and also got knotted up!
EDs. |
XXII.—OCCURRENCE OF THE WOOD SNIPE IN THE
CENTRAL PROVINCES
In February last, while shooting in the Mandla District of the
Central Province;, about five miles trom Amarkantak, I saw a Wood
Snipe (Gallinago nemoricola). Forsyth mentions having seen one
once in the same district, otherwise I have never heard of the
occurrence of this kind in the Central Provinces.
I should be interested to hear whether any other members of the
Society have observed the species in the Provinces.
UNITED SERVICE CLUB, JeMCAC aC ink:
PALI MATEO. Wa, Captain, M.C.
June 7 J1e2%.
XXIII.—NOTES ON DUCK SHOOTING IN THE ROORKEE
DISTRICT, UseP., IN THE VBARSe1I03 Oe o27,
(With a graph)
1. The record extends over 21 seasons from 1903 to 1927, with
three gaps of single years, in 1909-10, 1915-16 and 1923-24. The
bags were ali made in the jheels formed by the Solani R., close to
its confluence with the Ganges R., about ]5 miles S.E. of Roorkee,
in the extreme N.W. of the plains of the U.P. (and some 35 miles
S.E. of Saharanpur). In the great floods of September 1924, both the
Solani R. and the Ganges R., were in exceptionally high flood, and
as a result almost all the jheels were completely blotted out by
being silted up by sand from the Solani R.; so that where there
used to be 6 ft. and 8 ft. of water in a jheel there is now dry land with
grass growing: this area wiil therefore never again be attractive to
duck, unless indeed another cataclysm occurs similar to 1924. The
duck shooting in this district is therefore now entirely ruined, and a
thing of the past, as proved by the figures of the years since 1924:
I have therefore thought that it may be valuable to record the figures
now before it is too late (especially as a record extending over 21
seasons in the same district is not frequently obtainable), as they
may be of value in the study of the distribution and migration of
the various species of duck in India. For the past three seasons 1924
to 1927, I estimate there have never been more than some 500
duck in all the area of the former jheels, where prior to 1924
there used to be from 6,000 to 10,000 at least all through a
season.
2. The figures given in my Table are taken from my game-
book, written up in ink at the time. I have always been in the
habit of vetting the bag carefully on return to camp each evening
MISCELLANEOUS NOTES 601
and morning, and recording the numbers of each species separately
in the game-book. Capt. E. EB. Searight, r.E.,has very kindly veri-
fied the figures in the Table with my original game-book.
The figures in the Table have been plotted as three curves on the
Chart shown, for easier analysis. The bags were all made by
‘flight shooting ’, i.e., on birds moving naturally to and from their
feeding grounds at evening and morning, and without any driving,
etc. ‘The duck shooting ‘season’ has been taken as from Sep-
tember 15 to April 15, but as arule very little good is done before
November 1 or after March 1 as the main body of the
migratory duck arrive in the Roorkee District between November
1 and 15 usually, and leave again by end of February. ‘The first
species to arrive in any numbers are the White-eyed Pochard,
Gadwall and the Common Teal, and the last are Wigeon and the
Red-Crested Pochard; the latter are often not ‘in’ till middle of
December, and they are also the first to leave. The last species to
leave are the Garganey Teal (who moreover do not arrive in this
district in numbers till February, presumably on their way back from
further south ; they do not seem to come to the Roorkee district on
their arrival in India): also the White-eye and Gadwall stay late,
some few even tiil April sometimes.
The ‘ evening flight’ for record purposes has been taken as from
3.30 p.m., till dark (about 6 p.m.); and the ‘ morning flight’ as
from dawn till 8.30 a.m. or 9a.m.; though on misty mornings the
birds will flight tili as late as 10 a.m. Shooting inthe middle of the
day merely spoils the evening flighting, and never leads to anything
ofa bag. There is usually a big afternoon flight which goes on
spasmodically between 3.30 p.m. and about 5p.m., apparently birds
stretching their wings after their mid-day siesta. And the real
‘evening flight’ lasts from about after 5 p.m., till well after dark,
these being duck going to feed.
3. The figures in the Table present several points of interest ;
the years 1906-7, 1907-8, 1908-9 were all exceptionally good
years ; and so were 1918-19 and 1919-20; also 1914-15 was good:
in these years unusual numbers of duck arriving in this district.
This is well! shown onthe chart: the red curve of ‘ Total Bag’
showing ‘peaks’ for these years, corroborated by the blue curve
showing the average bag per ‘flight’: from the green curve it
appears that big bags of over 50 per flight are more numerous in
the years when duck are more plentiful, as one might perhaps
expect. The years 1907, 1908 and again 1918 were exceptionally
dry years, when the monsoon practically failed in the Roorkee area;
so it seems as if in a dry year more duck come to India; though
another explanation may be that in a dry year the smaller outlying
jheels are dried up and so all the duck are concentrated in the larger
and deeper places, such as the Roorkee jheels used to be prior to
1924.
The 27th column of the Table, giving the average bag per
‘flight’ in a season, shows that one’s average expectation in the
Roorkee District was about 29°68, say 30 birds, and cutting out the
last three years, 1924-27, when the conditions had radically altered
for the worse, it would be about 40 per flight. And the last line of
602 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
the Table, showing the average numbers of each species, shows that
in any bag one might expect to get 37 per cent. Gadwall, 13 per
cent. Mallard, 11 per cent. White-eyes, 11 per cent. Common Teal
and 7 per cent. of Red-crested Pochard: though of course any
particular bag, owing to the nature of the place chosen as a stand,
might consist almost entirely of one species, e.g., all Mallard with a
few Common Teal if the place was in shallow water ; and all Red-
crested Pochards if the water were very deep, with perhaps a few
Gadwall.
4. Some points noted in my game-book that are not shown in
the Table, are of interest as regards the dates the duck arrive and
leave ; thus taking the different years :
1902-3: On October 18 (i.e., early in season), a morning bag
consisted of 12 White-eyes, 2 Common Teal, and 5
Cotton Teal.
1904-5: On October 22 there were no duck ‘in’ at all: on Nov-
ember 19 a bag of 43, showing the duck came in
between those dates: on March 5 (late in season)
a DAG OL. O1:
1906-7; Was an exceptionally good season: between April 8
and 11 (very late). 4Gadwall, 1 Red-crest Pochard,
2 White-eyes, and 13 Garganeys were got.
1907-8: On November 8 birds were few, only 20 bagged: on
January 27, 1908, all the Red-crests had left; (very
early). This was a very good season, anda very
dry year.
1908-9: On October 31 a bag of 24: while 69 were got on Nov-
ember 1, so the birds came in early this year: this
also was a very good season, and very dry.
1911-12: On March 9 a bag of 40: the duck were leaving, and
the weather very hot.
1912-13: A poor season: the Soilani R. altered its course owing
to big floods in 1912 monsoon.
1913-14: A good season: on November 8 abag of 19 only, birds
not yet ‘in’ properly ; on March 1 bag of 20, birds
leaving fast.
1914-15: On March 2 (full moon) three consecutive flights of 40,
41, 33: In monsoon of 1914 the Solani R. altered
its course radically and formed several new jheels:
a good season.
1917-18: On November 17 a bag of only 7: birds not yet
arrived; this was very late: on March 23 only
3, birds having left: a very poor season with few
duck. |
1918-19: A very good season with exceptional numbers of duck.
1918 was a very dry year with no monsoon: on
November Y a bag of 55, the birds being well -
fan. : a Z
MISCELLANEOUS NOTES 603
1919-20: On November 15 a bag of 101. A very dry year, with
very little water in jheels.
1920-21: On November 27 a bag of 91, this was also a very dry
year with no rain after August 20, 1920.
1922-23: -On March 9 a bag of 92, very late in season fora
big bag. Heavy monsoon in 1922, and heavy rain
in February 1923.
1924-25: A very bad year: In September 1924 exceptional floods
in Solani and Ganges R., nearly all the Solani jheels
were silted up solid, and no longer existed after
the floods.
1925-26 and 1926-27: Both very bad seasons: there were
hardly any birds throughout both seasons, owing
to there being no water area for them to settle
on.
5. Numbers of the ditterent spectes.—The last line of the Table (in
bold type), gives the percentage of each species shot, on the average
of the whole period of 21 seasons; it is interesting to compare
these figures given for the U.P., east (district not stated), in the
Table on page 59 of D. Dewar’s book The Ducks of India
(1925 edn.) ; his figures are the number of each species occurring per
1000 birds shot. It may be here noted that Roorkee is in the
extreme west of the U,P., and there seems to bea considerable
difference in the numbers of some of the species occurring in the
east and in the west of the U.P.: most notably in the case of the
Red-headed (or Common) Pochard, and the White-eyed Pochard
and Garganey Teal, and to a less extent with Gadwall, Pintail, Spot-
bill, and Common Teal.
|
U.P. West U.P. East REMARKS
Species | (Roorkee) (Dewar) Roorkee Nos. compared to
_ per 1000 per 1000 U.P. East Nos. are
Mallard aaa 130 110 Rather more.
Gadwall eeeu|? 3/3 150 23 times as many.
Red-crested Pochard ... 74 80 About the same.
Red-headed (Common) 56 ils) Nearly four times as many.
Pochard.
White-eyed Pochard ... 116 270 _Much less than half the num-
| bers.
Pintail he | 22 | 40 | Half as many.
Skoveller teal 37 | 30 _ About the same.
Spotbil! ete. 14 6 Twice as many.
Wigeon a 21 16 _About the same,
Common Teal ue | 119 200 , About half as many.
Garganey Teal ee | 8 | 50 Less than 1/6 the Nos.
In the Roorkee District, the Gadwall (37:3 per cent) are the most
numerous species, being three times as many as any other kind. Next
come Mallard (13 per cent), the White-eyed Pochard (11:6 per cent),
Common Teal (11:9 per cent), Red-crested Pochard (7:4 per cent),
Common Pochard (5°6 per cent)—then the Shoveller only 3°75 per
604 JOURNAL, BOMBAY NATURAL AIST: SOCIETY, Vol. XXXII
cent; while, Wigeon, Pintail, are only about 2 per cent each, and
Tufted Pochard and Spotbill only 1:45 per cent each.
In some years exceptional numbers of particular species seem to
have occurred in the Roorkee District e.g.
Mallard (normal 13 per cent). In 1906-07, 17 per cent; and in
19¢38-9,17 per cent were bagged.
Gadwall (normal 37:3 per cent). In 1906-7, 43:4 per cent; in
1919-20, 43 per cent: in 1920-21, 47 per cent; in 1921-22, 60 per
cent ; and in 1922-23, 47 per cent were bagged.
Pintati (normal 2:2 per cent). In 1907-8, 4:4 per cent; and
1918-19, 5 per cent.
Wigeon (normal 2:1 per cent) In 1911-12, 4:2 per cent; and
in 1918-19, 4°3 per cent.
Red-crested Pochard (normal 7°4 per cent). In 1920-21, 15-3 per
cent.
Red-headed (Common) Pochard (normal 5:6 per cent). In 1906-7,
8:5 per cent; in 1918-19, 9 per cent; and in 1920-21, 9:4 per cent.
White-eyed Pochard (normal 11:6 per cent). In 1908-9, 13:9 per
cent.
Tutted Pochard (normal 1:45 per cent). In 1913-14 an exceptional
increase of 10°4 per cent.
Common Teal (normal 11°88 per cent). In 1907-08, 20°3 per cent;
and in 1918-19, 16°6 per cent.
Apparently in the exceptionally good (and dry) years, such as
1906-7, 1907-8, 1908-9, 1918-19 the Maliard, Pintail, Wigeon,
Common and White-eyed Pochards, and the Common Teal, formed a
bigger percentage of the bags thanin morenormal vears: while the .
commonest duck, the Gadwall, formed an exceptionally large part
of the bags in only twoof the exceptionally good years (1406-7,
and 1919-20), and then again in three consecutive seasons (1920
to 1923) that were not remarkable for large numbers of duck.
6. Some further notes on the commoner species in the Roorkee
District.
Spotbill. There are always in every year a good many small
parties (two to six) about during the whole season, and they
probably remain all the hot weather. All I have shot were the
common form (Anas p. pecilorhynca).
Comb Duck. These are uncommon, but one saw one or two about
in most years, early in the season only ; though I shot a pair on
January Livd9i5.
Cotton Teal. These are not common, but early in November ; a
few are seen about in most years; these appear to leave the
district after November, presumably for the south.
Lesser Whistling Teal. Nearly every year Isaw a few flocks
about; which disappeared after end of November, presumably to
go east or south. I never happened to bag any in the Roorkee
District, but other guns bag a few most seasons, always early in
the season.
Pintail. ‘These are not numerous, but every year there are a few
flocks all through the season.
Wigeon. Also not numerous, but occur every season; they come
late (in December) and leave early (February).
| \ \ I | | a .
p } | I | | | | i | 1 I ae Ee L b Sal oan ee ec tee . ‘Suosvas
Ba = >
pe OSES" se tel 8 2 2 2 B@ PeeWee sss esa gs ae zy
eS ~ ~ S N nv Be ~~ 3S ) N 3 \ % ® Gq :
ys 8 rs * “ nd S 8 = =
S ! (4)
0
00L
oo
FAT; f
64 002
eze
~SF2 ~ ™ 192
ae = = SS yy 00F
a 8c \ g0e
re & LOE," eve we
Ny XN =
Ss s S 00v
ae “ue &. >
uv NY
8 “ = & F005
x ver > NS
y u » |
255" =
ols 5 = 009 0
oy 3
oO Ne 0597 >
see se Ly OOL
5 LS G
65. = 8 l
‘\ 5 “ $ Loog
Ne & & Sie 2 “els 3 00. SS
Pye ~ ~ 6Le €-6l_.. 4“ 8
= = N 261, omen w Red x Foz
“xe ra 6-81" BS j p= WY a a 006 SS
9-6 & mes \ 9:02 iS ; %
Ly ! 4 w / 9 o
8 x = ! a / = i 8
3 \ ei a s ! oor S Foe
SI \ =: %
| \ x
oy oO OS
8 \ S
' < bor
ich s
iy
ov]
1
Na Z
nx l ene aes!
! =
* | <e
“*eserecnccceerMee remo, Fe Sie) \ ! y 9 =
4 o is the \ yg =
yO, Sw : 2
By $19 : =
or t r
796, : °
4 .
“eae Bay, | 3
" Bees eat 5
9 o
Mate cesscscccesie ‘uosoas ous og 420 szybyf big Jo‘ou 70;01 F
ee = = = WS:095 19d ry bf our Hw on abosaay $
OS DOLAD, uu p2bbogq yonp 7979
burjooys yybyf 4g epou yonp Jo shvog Jo pioray
‘WoOYbUIUUuNnD HY JOD 49 Zz6s 0f C06L mos suosvas {Z Ut YN ‘aayd400y AoaU
: ISAO pue jo i ret
j Gosves Ul S}YysIp [eloy : : : : :
*IIAO
pue 0g jo s}ysIp : a? Sm c 00 ur
UOSvdS Url [BIOL ° .
"Yoel Uy
‘Alenuer uy : : 4 on et
|
|
; ae Uy oa a is
|
|
‘Iequls0aqy uy : : 5 ie N i) <
Big flights of 50 and over.
fan
—
—_
=
[=
=
22
T=)
‘ced
TABLE SHOWING DETAILS OF BAGS OF DUCK MADE BETWEEN 1903 & 1927 IN THE ROURKE DISTRICT, U.P.
SURFACE FEEDING Duck. | | | Ba a a Big flights of 50 and over. 8 REMARKS.
= —___—__—— wane —— = co < ——____ —_______-—_—_|] @ = =
= a
k shoot | 5 bo a OR a we lags a Ey av 2
De eH | & | g |sa] 3 Shee | | Bs fle le | a8 |. 8 Dates of
ken as i | x jez] « HOS og | es es ue fa |4). Hee | a bagging |
September g.- | a | ay laa ¢ | 9» 2] : Cis Ea st Gl|oen Ieee eee 25 uncommon ( m= Morning ‘ flight’, dawn (6 a.m.) till 9 a.m.
So April AR] Ss i |= | phy ia B® ) oa i sl Sh ect ail Qa i) Sole, F! ./ es mes >. || SBeeiss of i e= Evening ‘ flight ’, from 3.30 p.m. till dark
b = - : Hy : an D $ oe = . 9 S| 8 s/s |" = aK ik. 6 p.
ees) ils le lS) ee leg Sf yg Woes SB eh GES Cee) eral ae es | 2 Sea se oe Cy
a z Fingered et yy a iS 2 = = $ |} 33/25] ae] 25 |] 3 | ess EI ® |/as|/ 81s = | ana | oe S12 \a12\/3| ams) ee es
2 | 84 oI Ss 3 i = os bo ZO luvié =D 2 wists! | ig be BTS Ome lee: a as6/8s Z2l/Alwa |e [a | Shee | sa ou
season 2 = | oe = c} rel] tS a ae 6a }ag|o5] £8 5 |ot&| o CE lich 3 = a |6g4)]53 ale | es ce | eno] 25 Bo
Sas & 1016 eye |) eae I eS ES = 16 | 6 ols Sh ol) tepell ea. iene Mika NEL: cet Ale eee) eye fe ca
a | | ae ie | | | Bo N.B.— FIGURES OF THIS YEAR ‘91/95’ NOT
1894-1895 43 2 %.|| 69 90 61 | eel | 278 21 a os INCLUDED IN TOTALS BELOW.
| ! = —t— ———— ———
a = pe = = _—_—— i 7 sale ] | | ee |
ya13-1904 je 2) a 1] |) G35 8 37 1 3 2 12 | Sh oe Wc 17094 ed 3103337 se bane4 | Parse [Rca eset eons en 28 612-03 fae
| | |
icop-1905 61, 1 emo 51 6 128 Sle 18| 34 80 P|) ats 32 | Ipmergoga] martes e2d:IMiee. |x liecet petal sseilll az | 49 pI98-04{ tees
| | nA | 30
195-1906. 3 85 1 6 11 | 3 Oe) « cexrs 29 | 46 Bl | 64 | | | 261 12, | 21-7 1 1] 2 | GEARS rere ES
| | | | | | | 29-12-06
AI ~~ | i} e *12- 1-07 e}
1906-1907 Zk gl] 2) w2) a} 62) 6| 423) 6 | eevallie 72))) 4934 | 107K 9 | m2] 15 | 1) | 43] 1,095] 27] 405 2) sil cali 6 | 78 | ea A very good season.
| | |
| | | | | | | |
| | | | | | | \ | | | | f A very good season—a very dry year.
= 3 | | 4 4 | 33 | | 7 | | | 53:51] 5| 2 P| ee 8 1 §1562 | cecsee § Local record flight till 1919—made on evening of
1907-1908 7| 42 7\ 60 45 | 442 133 |tee 39 53 Gs SEU os 207 |... esate i v | 1,017 19 | 53°5 1 | 3 November 15, 1907.
| | | | | | | | | | |
| | | ap §| §$29-12-081| 65.00
g-1909 ... | 23| 2 {emer 79)| Sy |ec43i |e o5ylennesa7i|| Sileesailiey72)| os2)|iee nail rail) 116) (nea | ere yee tS LAH ie taiye 1,013)]| <273)| 37: SMe AN ceepl| sl <- Saale 704) ha 3-09 §\Spectes|not xecorded-Wes. very ig ooulyears
I } | | 3 | "
|
hart | | | Ft a 53 {| $20- 2-111) Only 18 days in December, J: d Feb
1910-1916 5| 22) | ee 1 | 132 TAN ess 65| 10 25ullyen 91 CO Sat rs Il eae lheccs ti] 347] 18] 19°3))...| 1 Pa UW ore 63 {| SaRsOt| nly 18 days in December, January and February.
| | | | | | H | F 7
1g]1-1912 | 7 56 | 2) 14 1 171 PSU! eco} EA) 20 63 i4 0 47 | Sh les oA a a 494 24] 20:6}... | 2 2 73 gk eos -= 2 | esse
| | |
| | | | | ( Heavy floods in February, 1913, and Solani R.—
n | | 7 aR 4 } MARI ~ 9: Pal hern as CO | eareo attend its course.
‘912-1913 ey | op |) He te 57 Ol om a 10S u os 14 | a). ae = sca | 308 BN) SEE ces || cp | er ae | Only } season after January 1, 1913.
| | | \
| } | |
9131914 4 | 3] 93} 9] 49] 26] 209] 19] | 51 79)\\) 37M\| ee 65| 3 esa | 22] 29-7]... | 3] | 3 85 : | ic:
| ee) | 1 ea | | ell ell | ag |g phl-1-1 5)
19-1915...) 1 ¢2| 8] 34 23| 28) 3| 284 22 67} 7) 113} 32 Z| TSS ili eee ace sci 779 23 | 33:9\| .-. | 1) 1) 3 | ” me || oe | a i 311-15
| | | | | | | | | | | | | | | | | | | |
| | | | | | |
| | | | | | | | | | | | § €28-1-17 1| onty } sez f
1916-1917 | 7 | 56} 1 | Gi er SIG ey es 625 eng Co} SU 3197] Samet | aes 19 | eter |e tl] 4a} 10] 4b) |] 3) | 3 | | 65 |) S071. (aly } season, after January 1, 1917.
| | | | | | |
97-1918. 2) 59) 2h as 1 90| 10] ... 2} 2 134 er24 re 15 | 2 | | vel econ lal 227/|| 12] 18:O\ peal |) U))-- |] 2. Wess BP eee
| | | | | | 135¢ | ( Very dry year, no monsoon, exceptionally good
"4 | @ | nae 9 5 let qecwet Wo | season.
1918-1919 1} 81 17)\|| 23) 28 202 235| (ers: 5 50 27 5 tee 923 ese ees oo =n eon 554 9| 61:5'| 4] 2)... | -- | ‘ { 100e | Only shot 4 season, up to December 7, 1918.
|
| | | mga ! Avenged Year aryyine monsoon hardly.
4 | 5 . p > se 7 5 -11- record up to 1927— di i f
1915-1900... 5) us 3] 18] 13 CN PSU) es 6| 26 74 ES re 53) eee ieee a0 sceyall ees 657) 12) 54:7] 4|..| 1]... ie Gi) <2 { joie; 1 16-11-1915 t st pe ekg ECO WES TEEC
(920-1921. ) 5) Gr Si) Bi) Ay Sea) FEI 33) | ese 25 | ea ea eel (tes es 570!| 16 |) s56f/) | | 3] 2)... | 5 | x ‘SUS | eee =-= |
| | | | | | | | | Ws
1921-1922 xO 5; 19 9 8 14 173 3 7 2 1 mm | 41 2 | 284 8 | 35°-5¢] \ 2 | 1 2 | Lee eros =
| | | | | | | | | | | ks
1922-1923... as | 4 60 10; 14 3 115 Li 3 | 2 | 10 | 4 | oy | 18 | 1| a Ih te Wee) seco 245 | 13 | 8B). | Lie. | | | 1 | + 85 sees Exceptional floods in Ganges and Solani R. in Sept-
| | | | | | | | | | | | | ember 1924 and most jheels silted up altogether.
| | | | | | | | | | | | | | | | IP) 4
| | | | | | | | | |
Wels... | | | 9) 17 | 18 | cco |) ea am Nees | | S| @ A) 2 [2 |e | | | | 275| 14] 196 | | 1} | a | ac2 ees A bad season with few birds.
1925-1926.) | | oy | 2 | 3 4 2| 3 47 All es 13 1 27 3 ono 2 WW coer |) eco od ch 112 8) 140 cL) ecsillieemaltireee 1 el 53. | seen Very bad season—only shot 4 season up to Decem-
| | H | | | | ber 28, 1925.
| | |
1926-1927 # | a Filla 1 2 6 2a ee 13 1 32 4 | 6 4 83| 14] 5-9 | | | 21 | tees | Exceptionally bad season—no birds at all.
a i ee eee] | Sj Ef = - — — — ! | =) | — — ESSE
T% | | I e 2 | =r _ , a'|| Ik ral ory | N.B.—Geese are not counted in Total bag of season
Totals | 56] 2)...] 4 | 83/}1,295| 144! 371] 222) 3.696) 211] ... | 738'| 555] 1,154] 140 1} 1,177; 80, 2| 6; 2] 6) 9,888 | 333 | 29-68] 15 | 19 | 16 Vea Oy eH oad Vek ej in Col. 25 opposite—not in the percentages.
| ae
ia } | — | —|-—— ae ak bea | =| |e 3 |) | = | —s —_ —= —<
Average al | | | eo | | | | | |
Precentage | ue | on | oe | 104) 84/1308 | 145 3°75 2:24 | 3734 213 TAB SON) 1165 | 44 Ol} 11°88 | +808 | -02 | -06 | -O1| 06} 10000] ... | .. | .- |---|. | | we | a ns ap || ax see
Species, | | } | | |
|
| | | | | | | | | ] i |
" ; d are not included in the totals at
Notes,—All the Spotbill shot were the common one (Anus pipoccilorhynca). Notes.—The figures for the year 1894-1895 are only given for comparison, and i b
All the White-eyed Pochard were the western form (Nyroca n. nyroca). ae bottoin of Le Ge in the average of species, as the year was distant from the series of twenty.
TNA Sa ee eee pie eae cere aa 2 no “er fedjby ithe: Bombay Mnsiboares are taken from my Game Book, kept in ink, and written up daily—I have always vetted
b fatlini raat i i i yeni’ d mornings.
Though no Lesser Whistling Teal were actually bagged, one or two flocks were generally seen each year the species bagged immediately on reaching camp, evenings an F if
in November (caving for the south at end of November)—and a few bagged each year by other All the duck were shot by ‘ flight shooting,’ as the birds flighted naturally to their feeding grounds
guns, morning and evening.
Specimens of following species were bagged near Roorkee between 1892 and 1905 by other guns to my
knowledge—Stiff-tailed Duck (O+yura leucocephala)—Marbled Duck (Marmaronetta angustirostris).
The scamp (Vyroca m. morila).
Compared with Col, Cunningham’s original Game Book,
E. E. G, SEARIGH'T,
Capl., R.E.,
Bee PEMeelaie ‘A. CUNNINGHAM,
Colonel,
April 20, 1927,
tt
MISCELLANEOUS NOTES 605
Shoveller. These are fairly nutmerous in the jheels every
season, occurring in scattered parties.
Red-crested Pochard. These occur every season, the big flocks
arrive late (in December) and leave early. Owing to their occurring
in big flocks, a big bag is often largely of this species.
Red-headed (Common) Pochard. Some come every year; but they
are variable in numbers from season to season, much more so than
the Red-crests.
White-eyed Pocharyd. Is common every year and ubiquitous
in the jheels: they come earliest of the migratory duck, and
leave late: all I have shot have been the western form (Vyroca
N.NYVOCA).
Tufted Pochard (Nyroca fuligula). These are never numerous ;
but a few small flocks come every season.
Brahminy. Quite a few of these are always about in the jheels,
though they are much more numerous in rivers such as the Ganges.
Common Teal. ‘These are ubiquitous and common every year ; this
year however, 1926-27, it was noticeable that there were hardly any.
Garganey Teal. Very few of these appear till February ; pre-
sumably on their return migration from the east; they are then
fairly common ; and stay last of the duck.
The Smew. A few scattered individuals occur most years.
The Goosander. 1 have never seen these in the jheels, but have
shot them in the Ganges R. near Hardwar, where they are fairly
common.
Geese. Both the Grey-lag, and the Bar-headed Geese occur in small
numbers in the jheels in most years: the former being far more
common than the latter.
7. The following duck are very rare in the Roorkee district :—
Golden Eye (Glaucionetta clangula). 1 shot one on February
20, 1911, and sent it to the Bombay Natural History Society for
verification.
Falcated Teal (Eunetta falcata). I shot one on January 31, 1915,
and one on January 28, 1917, both were sent to the Bombay
Natural History Society for verification.
Common Sheldrake (Tadorna cornuta). Isaw two of these birds
flying with and leading four Brahminies, unfortunately my shot
brought down the third bird in the line, a Brahminy !! so I was
unable the make certain: but I am certain they were Common
Sheldrake (of which I have shot several in Sindh), as the birds
were well in shot and the light was bright, about 8 a.m.
Marbled Teal (Marmaronetta angusttrostris).
Stitt-tatled Duck (Oxyura leucocephala).
The Scaup (Nyroca m. maria).
Specimens of these last three have to my knowledge been shot
in the Roorkee jheels by other guns between the years 1892 and
1905.
RoorKEE, U.P., A. H. CUNNINGHAM,
April 20, 1927. Colonel,
e7
608... JOCORNAL, BOMBAY NATORAY, WIST. SOCKET Vo Wool. XO]
XXIV.—SNIPE AND WOODCOCK IN S. INDIA.
The following extracts from my Game Registers of the last four
seasons, showing the different varieties of Snipe, and the number
of Woodcock shot, may be of interest :—
a Cooma! g 2. ;?
ls! 8 £\Elx/2
Locality Year | Month! z 2 he a ols Remarks.
o— = Co eal
eee llama
ae Se ae | || | is
| | |
Ootacamund ase 1923-4 aa | Ly 97s aia e eitst ShotOcotpens!o:
7 pe PG0GE S| «As SB eel elsleaet nor February 22.
2 VIO 26-6) Secs. | ae Slien, Deol eae First shot September 28.
| Last shot February 19.
| | Wood Snipe January 2.
Pf ... (1926-7)... |...| Q7|...| 2)...|...] First shot September, 28.
| | | | | | Last shot March 5.
| Wood Snipe January 29.
Cannanore seat 1925 Dec: 18 IN] Pees ba
P are 1926 |e) ops UO} ae 22 Fs ern IR
} 1926] Nov.|23| 130] 1\...! 21 2
. 1927, Jan.j38) 134)... 3|21
S. Mysore a 1926 Jan. 23 SO) het et
* i 1986) sce Ite] asl elenlel
. 111926-7) Dec.-|15] Bkastecs| dass
| | Feb. |
75 Dy
S S
Locality Year |3| Ist shot | Last shot = Remarks.
g Es
Ootacamund _... |1923-4.14) Nov. 5 { Jan. 26 | 3) Stopped shooting on
January 26.
a .. |1924-5/27; Dec. 12 | March9 | 5 se
+ .-» |1925-6/14, Dec. 16 | Feb. 19 | 7| A poor year.
of . |1926-7 24; Dec. 11 | March 2 a ae
NOTES
Ootacamund.—Fantails are distinctly rare on the Plateau ; in fact,
the one shot in 1923 is the only one I have heard of for some time.
Wood Snipe are commoner in the Wynaad than up here, where only
four or five are shot each season. I send for reference the wing of
one shot last January.
Cannanore.—Pintails largely predominate ; occasionally one of the
larger variety is noticed, e.g. six during 1927. I send for reference
the tail of the last Swinhoe shot.
S. Mysore.—The Fantails included several more or less typical
specimens of Radde’s snipe, but unfortunately no record of them
was made at the time.
MISCELLANEOUS NOTES 607
Woodcock.—These seem to come in later than formerly, possibly
visiting the Anamalais before arriving here. February is the best
month and nearly all are gone by March 15, or earlier if heavy
rain falls. I think 25 to 30 may be considered a good average
season’s bag, it being remembered that smalJl game shooting on
the Plateau is limited to two days a week. Three woodcock weighed
on January 5, i.e. well in the season, were 10, 11 and 11 oz.
respectively.
LOVEDALE, E.G. PHYTHIAN ADAMS,
NILGIRIS, Major, 1.A., (Retd.)
Fane 1, 1927,
XXV.—A CORRECTION TO Mr. B. B. OSMASTON’S
‘NOTES ON THE BIRDS OF KASHMIR’
In Vol. xxxi, part 4 of our Journal, Mr. B. B. Osmaston has com-
menced a series of very valuable and interesting notes on the birds
of Kashmir. He will perhaps forgive me for venturing to correct
two of his identifications, as mistakes quickly gain currency and are
endlessly repeated.
No. 75, Acrocephalus concinens concinens.—'The Chinese Paddy-field
Warbler.
A Paddy-field Warbler has been known to breed in Cashmere ever
since Brooks first found an empty nest at Shupyion on June 13, 1871,
and shot the male bird which is still in the British Museum. Yet
though we have for some years known it to be common and Rattray,
Wilson, Davidson, Bates and others have collected numbers of the
nests and eggs, we cannot get its name right. In the last edition
of the Fauna we find it shown under two species Acrocephalus agrt-
colus (which may nest in Turkestan or the plains of India but cer-
tainly does zof nest in Cashmere), and A. concinens concinens (which
in my opinion does not occur anywhere in Indian limits). It re-
mained however for Mr. Osmaston to collect a proper series of skins
though owing to his departure for Africa he was unable to examine
them critically and merely accepted the identification given in the
Fauna. I have seen his series and also various other birds collect-
ed in the Cashmere jheels in the breeding season, 17 inali. I
have no hesitation in saying that these birds agree with Harington
and Whitehead’s series inthe British Museum from the Kaghan
Valley, the type locality for Acrocephalus conctnens haringtonz, which
is therefore the correct name for the Paddy-field Warbler which
breeds so commonly in Cashmere.
No. 77. Sylvia curruca affinis. Indian Lesser White-throat.
Both Dr. C. B. Ticehurst and myself have endeavoured for some
years past to dissipate the legend that this species breeds within
Indian limits, but it is repeated again and again like the mistakes
over the Paddy-field Warbler. Mr. Osmaston kindly gave me two
White-throats coliected by him inthe Suru Valley to identify his
eggs. These are both specimens of Sy/vza althea, Hume’s White-
throat which is really the breeding Whitethroat of Cashmere. It
608 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
will be noticed in Colonel Meinertzhagen’s account of his trip to
Ladakh (/ézs., 1927, p. 421) that he found S. althwa to be the com-
mon breeding bird of the Indus, Shyok and Nubra Valleys. Mr.
Osmaston’s mistake was of course due to his hurried departure for
Africa before he could make a fresh examination of his skins, but I
can only end by urging that for the present no egg collector in India,
Cashmere or Baluchistan will name any Whitethroat’s eggs without
a skin from the nest to support his identification.
CALDBEC HOUSE, HUGH WHISTLER.
BATTLE,
August 30, 1927.
XXVI.—AN ALBINO TURTLE
On June 25, I secured a specimen oi the Common Mud Turtle of
these parts, Amyda granosa intermedius, Annandale, which is appar-
ently an albino. It was a half-grown specimen and was bright
yellow throughout with the exception of the limbs and neck which
were chrome yellow. The plastron had the usual colourless appear-
ance. It was taken in the Ambajheri Tank at Nagpur and is depo-
sited in the Nagpur Museum. (Reptile No. 274.)
NAGPUR, “C. P.,; BO AVD ABREU. Fz.s
August 17, 1927.
XXVII.—NOTES ON THE DESERT MONITOR
(VARANUS GAISEUS) AND THE SPINY TAILED
LIZARD (CROMASTIX MICGROLEPTS)
The following field notes may be of interest: they have been
compiled from observation and the experience of Bedouin hunters.
Most of the facts were obtained from Nahaita, an Araif tribesman,
who has spent most of his thirty years or so of life hunting the wild
animals of the Shamiyah Desert, and Radam, who has had a similar
career. For their truth I cannot vouch, but both men are now in
the Camel Police and are models of reliability when reporting, and
of observation and knowledge when anything relating to the local
stretches of Iraq and Najd is under discussion.
The Desert Monitor (V. griseus) locally known as the harwhal,
ranges in adults from twelve to twenty inches in length. It is so
called because it is never seen travelling at any other pace but that
of a fast wriggling run. I have myself, however, seen them travel-
ling at what could only be termed a slow stumbling crawl but this
was in about twelve square feet of confinement.
They are easily overtaken by a running man. When neared, they
stop, turning their heads with mouths wide open in the direction of
their pursuers. They then lash their tails furiously from side to
side and shoot their tongues in and out at the same time hissing
loudly. If foot or stick is approached more closely to them, they
will make sudden darts snapping fiercely at the offending object,
and. although the only result is bleeding and broken teeth, they
persist in doing the same thing repeatedly.
MISCELLANEOUS NOTES 609
I have found that the best way to catch them is by a sharp kick
under the belly which bowls them over, a foot placed quickly on the
head then enables one to pick them up by the tail when a noose may
be easily passed over the head and up to the waist. I have never
seen them biting the line that holds them captive.
These reptiles are found in the Shamiyah Desert ana aiso in the
Gizerah region between the Tigris and Euphrates. ‘They prefer
sandy places especially with mounds or undulations, and it is only
when these are present that they are found in Subakha, a salt mud
of treacherous consistency found in patches of varying size in the
local desert.
Though they sometimes dig their own burrows, they more usually
adopt the disused holes of other animals such as rats and jerboas.
When its own excavation, the shaft is driven in at an angle of about
forty-five degrees to a distance of roughly eighteen inches, the
sunken end being expanded into a chamber. The outlet remains
single.
They are said to feed on the surface only, never taking food below
ground. No vegetation is eaten and they dislike water. Their
food consists of crickets, small lizards and local desert snakes.
Of the pairing of the harwha/l nothing is known, they are always
seen single. ‘The female is similarly coloured to the male, that is
with straw coloured belly, and back of a greenish-grey ground with
darker mottling. The belly ofthe male [ have noticed has a pinkish
tinge in early summer. The female is slightly smaller.
It is said not to hibernate, and as a fact is seen much more fre-
quently in winter all day long than in summer when it is seen in
early morning and late evening only. It apparently objects to more
than moderate heat and thisis borne out by its behaviour in capti-
vity. At night and when not directly under the sun’s rays it lay
contentedly inthe centre of its cage, but when the sun shone directly
upon it, great unrest was evinced and it was not happy until a
crack or cranny had been found into which at least its head might
be pushed.
It is not venomous. Ihave been bitten and so has my spaniel.
There was a very small puncture, little blood and no subsequent re-
action of any sort. It is not eaten by the Arab.
The eggs are laid in the burrow chamber and number fifteen to
thirty. They are described as being of the size of a pigeon’s egg
and are white spotted with brown. ‘The young are similar in colour
and appearance to the adult.
Both belly and throat swell when the animal hisses, the maximum
of hiss and swelling occur together.
The only folklore obtainable was, that the tribes say that if the
teats of goats and ewes are found swollen, the recent pastures are
searched anda harwhal is invariably found in the vicinity. They
say that ithangs on to theteats and drinks the milk. ‘The tribes
admit that no teeth marks are ever found. In captivity they do not
touch milk.
The ‘thub’ or Spiny-tailed Lizard (UW. microlepis) has the head
of the ordinary lizard to the eye of the lay observer, but its body is
almost as broadas itis long. The tail is relatively short but
610 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
tremendously thick and carries fifteen to twenty horny spikes which
project backwards and slightly outwards from the main stem. I
cannot appreciate its utility; asa weapon itis absurd, and asa
boring instrument if it is true that the ‘ thub’ burrows head first, it
is at the wrong end of the machine.
My informants say that it gets its name from its never moving
from the one place. It is found all over the Shamiyah Desert but
only where stony cutcrops occur and is not found at all in the
Gizerah.
Its burrow descends vertically in a medium spiral to a depth of
about four feet, a horizontal off-shoot near the surface in the
female burrow containing the eggs. The burrows are occupied
singly and have one outlet.
Unlike the harwhal the ‘thub’ eats vegetation but of no
particular sort. It also eats crickets and it is generally averred that
the female thub watches her eggs hatch not from maternal
solicitude but in order to eat them.
They hibernate in December and January and are much slighter
in February.
They breed in April and May. The female is said at this season
to protrude her rear half from her burrow at the same time slowly
waving her tail. This attracts passing males who signify their
presence by hissing. Hunters often catch a female thub by
approaching gently while she is thus employed and imitating the
hiss of the male. The female leaves her hole and is then an easy
capture. Coitus is by ventral apposition; the pair do not associate
for a time longer than is necessary for the act. Most males are
darkly coloured similar to the harwhal but there are some much
lighter. All females are light.
The eggs which are white are thirty to forty in number and are
compared in size and shape to small dates. They are laid in the
female’s burrow and as stated previously are hatched for the
table. Whether she eats only a proportion or all that cannot
escape 1s unknown. Both reptiles and eggs are eaten by the
Arab.
They are not venomous, and ripe old age, longer than man, is
said to be their destiny if they escape the desert hunters. The
commonest method of catching them is to pour water down their
hoies when they promptly emerge.
Their throats and bellies distend when they hiss asin the har-
whals.
NASIRIYAH, NORMAN L. CORKILL.
IRAQ,
September 9, 1927.
XXVITIL.—HAMADRYADS IN THE KUMAON TERAI
I saw two hamadryads in my shooting block. Neither of them
seriously showed much fight. The first one made off through
jungle ; I was within 10 yards of it when first seen.
MISCELLANEOUS NOTES 611
The second was inmy back verandah. I heard a curious swishing
noise while having tea at 4.30 p.m. and on looking out of the
window saw a huge hamadryad slowly moving along the back
verandah. I called up the servants and we threatened it with
sticks and it made off rapidly. We threw big stones and one caught
it on the back, drawing blood. It reared up about 3 feet or more,
expanding its hood, but did not drive home its attack and finaliy
went up a bushy tree 15 feet off the ground. Meanwhile my man
brought a shot-gun and I shot it through the head. It measured
13 feet.
I may say that I saw two hamadryads last year in a block in the
Siwaliks and neither of them showed fight. One we killed, the
other was lying beside a stream when I came on it in the early
morning while out after achital. It swam the stream like greased
lightning and disappeared.
AJMER, W. M. LOGAN HOME,
Silty S 1927. Major,
4th Bombay Grenadiers.
[While the belief that the Hamadryad wiil attack human beings
with or without provocation is widely current, authentic records of
its doing so are extremely rare. In the majority of instances this
snake, like others of his kind, seeks safety in flight. Eps.]
XXIX.-MR. SLATER’S DEATH BY SNAKE BITE
Some months back you published in your Journal my letter
regarding the above, in which I stated that it was a well-known fact
in these parts that the snake which bit Mr. Slater of the Geological
Department was a King Cobra or Hamadryad. I gave the facts as
stated to me by the Range Forest Officer who was at that time in
charge of the Thirthahalli Range in the Shimoga District where
this incident took place. In alater issue of the Journal Mr. R. C.
Morris of Attikan Estate, Mysore District, contradicted my statement,
and said that it was not a King Cobra but a Russel’s Viper which
bit Mr. Slater and caused his death. Iam sorry I was not able to
reply to this contradiction earlier as I have only now been able to
get authentic confirmation of my previous statement.
Mysore, S. INDIA,
September 27, 1927. CHARLES THEOBALD.
[Mr. Theobald forwards us letters received by him from the
District Magistrate, Shimoga, the Director, Mysore Geological
Department, and the District Forest Officer, Shimoga. Though
none of these officials were present at the time of the occurrence
they conclude from the reports gathered at the time that the snake
which caused Mr. Slater’s death was a ‘ Kalinaga’ which according
to Mr. Theobald is the local name for Hamadryad. Enps.]
65128 JOURNAL, BOMBAY NATURAL AIST? SOGIE TY, Vol, 2Oea?
XXX.—A NOTE ON THE GREEN WHIP SNAKE
(DRYOPHIS MYCTERIZANS) AND YOUNG
(With a photo)
A short while ago we received in the Society’s office two
consignments of this beautiful and perhaps the most graceful of our
Indian snakes. Among them were a few gravid females. About
two weeks after their arrival one of them presented us with fourteen
young, twelve of which are alive and vigorous, one was born dead
while the last was expelled still enveloped in the fcvetal membrane.
They were born during the night of September 5 and were discovered
the next morning.
The mother taped 1262°5 mm. and the average length of the
young, taken approximately twelve hours after birth, was 345 mm.,
the longest was 368 mm. and the shortest was 339 mm.
In general colouring they are like the parent except that the
longitudinal line on either flank is white instead of yellow. There
is a distinct white line over the upperside of the orbit which travels
forward along the angle of the snout to the tip. This is apparently a
juvenile character as it is not present in the adult specimens. The
only yellow marking is a short line running backwards from the
base of the orbit to the point where the head is differentiated from
the body,
The first slough as is customary with snakes took place soon after
birth.
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Bed ntadim nome saw mee Pe Re Uhdet Meal isis Sasa pilar SEtiy
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Journ., Bombay Nat. Hist. Soc.
THE RECORD CUBBANY MAHSEER.
ht 105 lbs.
1g
We
MISCELLANEOUS NOTES 613
The specimens were measured four days after and though they had
had no food, showed an average length of 361 mm.—an average
increase of 16 mm. Thelongest measured 379 mm. and the shortest
342 mm.
On a former occasion a specimen which |! had in captivity gave
birth to twenty-three young, seventeen of which were alive and the
others born dead. On this occasion also they were born during the
night.
It is unfortunate that these snakes cannot be kept for any great
length of time in captivity as feeding them is a very difficult matter,
and though they endure a long time without food they soon begin
to look miserable and emaciated. They would perhaps live if they
could be kept in a large cage and given a supply of small birds and
mice from time to time.
The accompanying photograph shows a number of these creatures
in one of our cages.
BomMBAY NATURAL HISTORY SOCIETY,
6, APOLLO STREET, C. McCANN.
September 13, 1927.
{The mass of snakes shown in the photo were part of two
consignments of snakes sent from Hyderabad, (Deccan), to Delhi and
intercepted en voute by the postal authorities who, though they
tolerate most things, draw the line at live snakes. The sender
collected about 280 of these snakes for despatch through the post
from which we may conclude that the green Whip Snake is very
common in the neighbourhood of Hyderabad. Eps.]
XXXIL—THE RECORD CUBBANY MASHEER
(With a plate)
The mighty Masheer shown in the accompanying plate was taken
from the Cubbany River, South India (for which it is a record), on
November 6, 1926. It was landed in about 40 minutes using a Hardy
16 ft. split cane Hi-Regain rod, cutty-bunk line of 40 lbs. breaking
strain and Alcock’s special single hook.
Weight “ie ie ee, 105 Ibs.
Length ee: oe oa) 5 ft. 4 in.
Girth a a os oO ite Zi,
KIRKEE, G. A. HARE,
eawoust 1, 1927. Lt.-Col.
[Col. Hare has very kindly presented the skin of this magnificent
fish to the Museum of the Bombay Natural History Sociory. Eps.]
28
614. JOURNAL, BOMBAY NATURAL ‘HIS TASOCIE TV AVGl eos
XXXIT.—AQUATIC RHYNCHOTA IN THE COLLECTION
OF THE AGRICULTURAL COLLEGE, COIMBATORE.
5, INDTA
(Published wrth the permission ot the Director of Museums,
Sao aid LA..S.)
Rao Sahib Y. Ramachandra Rao, Acting Government Entomo-
logist, Coimbatore, has very kindly entrusted the collection of
aquatic Rhynchota in his care to me for identification. Though
there is nothing of outstanding interest among this material, the
foliowing list of the species contained in it seems to be worth
publishing, as it adds considerably to our knowledge of their geo-
graphical distribution. All these species, with the exception of the
first, are described by Distant in his volumes in the Fauna of
British India series.
Family HyDROMETRIDA
1. Hydrometra greeni, Kirk. Coimbatore, ‘at light’. (Coll. V.&.
December 11, 1914).
According to Bergroth (Zool. Med.; i, 2, p. 118, 1915). As oxeenz,
Kirk. is quite distinct from A. lzxeatus, Esch. (described by Distant
as AZ. vittala, Stal.), though Distant and others have regarded it as
a synonym, his opinion being based chiefly on the male genital
segments. A conspicuous character, however, is that the whitish
hemelytral vitta is broadly fuscously interrupted about four to six
times, while in A. /zneatus it is only narrowly interrupted behind the
middle by two transverse veinlets. As the examples before me are
females, I have based my identification mainly onthis character.
J. R. Ja Torre-Bueno (Sfolia Zeylanica, xiii, 2, p. 226, 1925), who
does not appear to have noticed Bergroth’s work, also regards
Hl. greent as a distinct species, though he does not give very
definite reasons for his opinion. WA. greenz is a common Ceylonese
insect.
2. Gerris anadyomene, Kirk. College Farm, Coimbatore. (Coll.
Sius./., October 11,1916).
This is another common Ceylonese species, which may be
expected to occur in §. India.
3. Gerris adelaidis, Dohrn. Pulney Hill, 5,000-6,000 feet. (Coll.
P. S. Nathan, May 10, 1917).
The specimen before me agrees exactly with the description
given by Distant, the semi lunate basal marking on the head being
very characteristic. This species is related to G. paludum and G.
spinole and would come in Distant’s generic section C. It seems to
be a little known form of which further material would be useful.
4. Onychotrechus vadda, Dist. Pulney Hill, 5,000-6,000 feet. Coll.
(9, 7Vathanz,.May.10. 1917)
5. Cylindrostethus productus, Spin. Cauvery Bed, ‘Trichinopoly.
(Coll. PS: Nathan, Mareh 20,4918).
6. Ptilomera laticaudata, Hardw. Coimbatore.
7. Metrocoris stali, Dohrn. Dodabetta, Nilgiri Hills, 8,000 feet.
MISCELLANEOUS NOTES 615
Family NEPID&
8. Laccotrephes ruber, Linn. Wynad, Malabar ; Ponanosat, Tanjore
District.
9. Laccotrephes griseus, Fabr. Guindy, Madras (Coll. Fletcher,
Aueuste 17, 1913) ; Coimbatore (Coll. fletcher, May 13, 1913 and
November 27, 1913°(Colk. C. A. S., March 27, 1915); Yemmiganur,
Bellary = Wistricts (Coll... C.°V. December 24,. 1912); Sirugappa,
Bellary District (Coll. D. 2., November 28, 1912).
10. Ranatra elongata, Fabr. Peermade, Travancore (Coll. R. S. V.
April2i-23; 1913); Combatore( (Coll. Y. #., May 18, 1913).
Family, BELOSTOMATIDA
11. Spherodema rusticum, Fabr. Mathimudongudem, Kistna Dis-
trict ; Coimbatore; Sewage Farm, Madras (Coll. flefcher, August
1913).
12. Belostoma indicum, Lep. and Serv. Kinavallore, S. Malabar
(Coll. P. S. Nathan, June 15, 1918); College Farm, Coimbatore.
Family NOTONECTIDA
13. Enithares indica, Fabr. Coimbatore.
14. Enithares triungularis, Guer. Peermade, Travancore (Coll. /saac,
April 21-28, 1918).
15. Anisops sardea, Herr. Schaff. Coimbatore, ‘ at light’.
16. Anisops niveus, Fabr. Peermade, Travancore (Coll. /saac,
April 21-28, 1918).
c/o F.M.S. MuSEuM, CEDRIC DOVER.
KUALA LUMPUR,
PROCEEDINGS
Proceedings of the Meeting held on September 27, 1927.
A meeting of Members of the Bombay Natural History Society and their
friends tock place on Tuesday, September 27, at 6.30 p.m., at the Prince of
Wales Museum. Mr. H. A, W. Brent presiding.
NEw MEMBERS
The following 51 members were elected since the last meeting :—Mr.
I. C. Petit, Bombay ; Mr. Ishwardas Lukhmidas, Bombay; Mr. Lakhmidas
G. G. Tejpal, Vile Parle, Bombay; Mr. A. P:Neatham, Bombay ; Mr:
Hooseinbhoy A. Lalljee, Bombay; Sir R. N. Mookerjee, K.c.1.E., K.C.V.O.,
Calcutta; Mr. A. W. John, Chittavarrai, S.I.; Mr. S. C. Petit, Bombay: Myr.
S.C. Barooah, Assam ; The Principal, Deccan College, Poona; Mr. Sultanbhoy
Bajeebhoy, Bombay ; Mr. A. K. Kothare, Bombay; Mr. H. B. Guzdar, Bombay;
Mr. C. H.. Phillips, Assam; Mr. E.R. Gfant, Washington, U.S.A. ; Mua:
P. M. Dalal, Bombay ; Mr. J. Burke, Bombay ; Mr. E. C. Benthall, Calcutta ;
Sir Onkar Mull Jatia, Kt., 0.8.z., Calcutta; The Principal, Maharfaja’s
College, Ernaculam, 8.1.; Mr. J. N. R. Mehta, Karachi; Mr. R.G. J. Crisp,
Itarsi, C.P.3 Mrs. A. J. Leech, Madras; Sir Cowasjee Jehangir (Jr.)) mt
K.C.S.1., C.1.E., Poona; Mr. B.S. N. Cooper, Bombay ; .- Phe. Ytvarail on
Limbdi, Limbdi, Kathinwar; Mr. Kaikhosru N. Chandabhoy, B A., F.A.A.,
Bombay; Major E.A. Breithaupt, Hyderabad, Déccani ; Dr. F: E. Davar,
L.D.s., R.C.S., L. M. &S.,J.P., Bombay ; Mr. Ebrahim Dawood Kazi, Bombay ;
Mr. C. E. Hewetson, I.F.Ss., Bombay; The Principal, Mrs. A. V.N. College,
Vizagapatam ; Mr. A.G.C. Elliott, Kishtwar, Jammu; The Librarian, Osmania
University College Library, Hyderabad, Deccan; Mr. D. N. Bahadurji,
Bombay; Mr. T. T. Kothavala, Dhulia; Mr. Hoshang N. E. Dinshaw,
Karachi; Mr. Fakirjee Cowasjee, Karachi; Mr. A, R. Hughes, Bombay ; The
Honorary Secretary, Ranikhet Club, Ranikhet, U.P.; Mr. Wm. Harper,
Bombay; Mr. Rustomjee D. Dalal, Bombay ; Khan Bahadur D. B. Cooper,
Satara; The Librarian, Lucknow Military Prison & Detention Barracks
Library, Lucknow, U.P.; Raja Sardar Singh Bahadur, Khetri, Rajputana; Mr.
Muhumud Faiyuz Khan, F.R.G.S., B.Z.S., Agra. U. P:; Mr. C. E.Atwoodr
Bombay; Mr. C. P. Gouldsbury, Mattupatti,. S.d.; Mr:C. P. Wallace
Bombay; Mr. B. P. Patel, Bombay; Mr. K. C. Dinshaw, J.p., Bombay.
The first case of a series to illustrate the classification of Indian Birds was
exhibited before the gathering.
Sir Reginald Spence read a paper on ‘Game Preservation in India’
compiled from an enormous amount of material received from members,
Forest Officers, Sportsmen and others competent to write on the subject from
all over the country.
The paper (since published in the Journal) dealt with the existing methods of
preserving game from destruction and suggested ways and means which might
be eraployed with advantage in future. Game animals should, on the whole, be
looked upon as the living assets of the country and therefore their develop-
ment, and preservation should be encouraged by every legitimate means. In
the paper attention was drawn to the various parts of the country which were
gradually but steadily being depleted of game and where more stringent
methods of control were called for.
This of course cannot be entirely left to the forest officials, generally
speaking are more concerned with increasing their forest produce revenue than
preserving game which is often injurious to the forest plantations.
After the paper was over, the Chairman introduced Mr. J. E. Barton, OF the
G.I. P. Railway, who very kindly prefaced the film ‘ Sport in India’ which
was subsequently shown, with his experiences with the cameramen and the
beasts depicted.
The film was kindly lent by the G. I. P. Railway Publicity Department and
was made at Betul, onthe G.I. P. line in the C. P. The object was te—show
the public of Bombay what possibilities there were in the way of Big Game
shooting within a twenty-four hours’ journey from the City.
The pictures were much appreciated and the Chairman moved a vote of thanks
both to the Publicity Department of the G. I. P. Railway and to Mr. Barton for
his prefatory remarks.
af
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Printed by George Kenneth at the Diocesan Press, Madras and published by
R. A. Spence for the Bombay Natural History Society, Bombay.
PPSOURNAL’ (7 s)
Geter foo ee
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EDITED BY a
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Bye © Stuart, baker, "E.Z.S., B.lsS.,:M,B,0.Ui., HoF.A.OLUs ilocsekee
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY, Part VI.
Eee ater Sd y bil: Deyuikis ly. Go he ccese. ves dbeien sehisescacd seas icievsccaesseeee?
A CoLLectTiInG Trip TO LApDAK. Part II. (With two plates.) By
Sete CESOIMI Careers ctse ch ccs tay vabuicaceiantess oie vies sVaslbigietecsoesichecshoaedtoeteck
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA. Part II.
(With two plates and three text-figures.) By R. 1. Pocock, F.R.S.
THE Bic BULL OF SEMBUTOLLAVU. (With a plate.) By C. P,
CE COMM SID Uy eterna acta esc veastsinds «aside Stelios ues ssiewess alleles pee veuwelceccess es
INDIAN DRAGONFLIES. Part XXX. (With three plates.) By Lt.-Col.
RCO ol HASSE ac PMG. 14s He PCSip sacesccet Secon veledos sc uats cues Vege vicoateceeasedns ced
THE STUDY OF PLANT LIFE. Partl. (With two plates and five text-
UOTE) ML ESV Ae MATES NLCC AMIN, Hoi ped ncedy eee dace cies osegdorcasiboseediseeseedss
NOTES AND DESCRIPTIONS OF FISHES FROM CEYLON. (With two
DUGLESe\ mY. ELCU TY. Wi, POWER” iistsosesceecetiaccn cstoatesdsccehovesee ree
THE MAMMALS AND BIRDS OF KASHMIR AND THE ADJACENT HILL
PROVINCES woceant, Ville sby Col. As He Warten oitescsctiissecccieceecec ase
FoRAGING TERMITES. (utermes biformis, Wasm.) (With a plate.)
yen Wis Gren EMIMOStOM Mal OF, I MeSc: «ges csclaetedeidh + oat Seta ves dbase edness
FURTHER NOTES ON BIRDS ABOUT SIMLA. By Hugh Whistler, F.L.s.,
ZS nee a eee ae se atinne seo ctnc eae erddncs talon siovabiaad eedcebourdemn cus sees sgesceastauee
SomME NEw SPECIES OF PLANTS FROM THE WESTERN GHATS. (With a
lale.). By Ey Blatter and, Cs McCantt =): Jics.csccssteasdesskeceecastessee ce os
BIRDS OF PREY AND THEIR USES. ( With four blates.) By C. H. Donald,
AES NTO: U Serer ies Serac at oibe tical ae lsil aca cle weeeceea sot e toivebblngelccedebcooesuwasasys
A NOTE ON THE BIRDS OF THE PESHAWAR District. (Witha map.)
By Rev. F. S. Briggs and B. B. Osmaston, M.B.O.U., I.F.S.,
WING) by aprdepeactdes sacle ts Rotiesierenwe ic easdhes Memes nee mau cenceags Nucvecrtaca zene
MoDERN MusEuM MetHops. Part Il. (With two plates.) By S. H.
PERUANO (Cp MisZ Soo ocsipccchcsesstiegse PAs cdiotinaacs cee mae ties tgece orm eseetes Ce Neeuee ss ary
REVIEWS—
A HANDBOOK OF THE BIRDS OF EASTERN CHINA. Part IV _........,
THE LITTORAL FAUNA OF KRUSADAI ISLAND IN THE GULF OF MANAAR
WITH APPENDICES ON THE VERTEBRATES AND PLANTS .,....ccccccceeees
RUE meAt G@ ANTE R Av EN “DIGHRe MAND) 9.32. 2ccdestsanccerdsecds ceeatrecccetcensasvossce
Hecate IES Me INR EEG g TWIN GIO EO © mucins tire as sui ci ge tess sale abecndcsesaecideletiesensaveevesnies
io BET ORIUAILA AiseenGCoeercReeeE rn ee eer eer Pe Oerere een acne ae ce aust ta Veet s ehcmainhe outs wes ues
SES IOILA SOIC VMMEN MR PIC ONE io sialic iuic> Fea sie oi Yale: ica dials bre Sais a Ceva:bsia sinsia yn cledisinte cy a'n'e ole ydeeledieesb'ee
MISCELLANEOUS NOTES :—
I.—‘ A Unique Lion Photograph.’ By F.W. Champion, I.F,S.
ANGEL PLYMON SII. As oANLT acdc: ceireal antels sects dns, ¥unesenet cecesmen
II.—A Clouded Leopard (Felzs nebulosa) eee Man. (Witha
pia By A. Brownlow; Jndian: iPONCEe |. f..0..0 di tecscase seas
PAGE
660
il CONTENTS OF Vol. XXXII, No. 4
Ill -—Noosine Tigers." By tet OAs) Scssncancteccceessesnes devahcastaer
IV.—Reply to Mr. Phillips’ Note on P. 371, Vol. XXXII ve Colour
of the Long-armed Sheath-tailed Bat (Zuphozous, longi-
Manus J .By C,-MeCanneaines. scc-.scecceceesees Secsaeseche <teceeinc cae ;
V.—Habits of the Porcupine. By Ralph C. Morris
VI.—An Elephant with Extraordinary ‘Glands’. (Witha plate.)
By MAC Sis AUT GS terse cc eeetenciesdoces sci dasieleaatestaeceeseseeteec nesses
VII.—On ‘ Natural Deaths’ in Wild Elephants. By Ralph C.
Morris
ee ee ee ee ee ee ee eee ee ee ee ee ee
VIII.—Clicking Noise made by Muntjac. By B. B. Osmaston,
Indian Forest Service (Retd.)
Cee ceo eoeseet¥secarce eer seseca'*® seerersees
IX.—Crows and Koels. By E. A. Andrews, Entomologist, Indian
Tea Association
HEC COSCO ESET F OOHOHEE LETHE EEE HESS ORE OH EE LOCH L EC EERECEOEE CEE
X.—The Rufous-Breasted Blue Flycatcher. (Cyornis hyperythra)
at High Levels. By C. Boden Kloss
COCO cee eseseteeoseorerccegsecese
XI.—The Nidification of the Sind Hill Warbler (Suya crinigera
styviatula). ‘By. El. W.) Waite, F.Z:S;, M.B.0,0.. “indian
Police
eeeerece SeaDoo ET ES SET ELT EHEC HET EOH LOOT ELSES ESET COCO ECC ECOL OE EEE
XI1.—On the Unusual Food-habits of Hornbills. By Satya Churn
Law, M.A., BoL., Ph: D.;f.Z.S.., M8 <O.U.
XIUI.—The Black-capped Kingfisher (Halcyon pileata) in the Dar-
bhanga District, Bihar. By Chas. M. Inglis, F.z.s., F.4.S.,
M.B.O.U.
CROKE COCO LERSERL EE EEEE BOT EEEEOES Bs BEOEES CEE TEe Leer eeEEEC TEE Seo eEOtE® eos
XIV.—A Quail’s Queer Retreat. Reprinted from the Sunday Times,
Johannesburg, Transvaal, November 13, 1927 .oc.c.cccccecee ce
XV.—-Occurrence of the Sheldrake ( Zadorna tadorna) in the Dar-
bhanga District, Bihar. By Chas. M. Inglis, F.z.s., F.E.S.,
M.B.O.U.
COO Fee eee Ber ee sect seeee Seterccerccceeseesos SSC Cee cereccccceeeecereD
XVI.—Occurrence of the Sheldrake (Zadorna tadorna) in the United
Provinces, “By f). Wo) Searight Capt as ee
XVII.—Recovery from the Bite of a Phoorsa (Echis carinata). By
A, A. Meaden, Lieut.-Col., R.A.M.c.
.
ee eS ee ee
XVIII.—The Mogul Emperors of India as Naturalists and Sportsmen.
By Sunder Lal Hora, D.Sc., F.L.S., F.z.S., Zoological Survey
of India, Calcutta
210 O 000 1010 BS, O.0 018 Bis 0 Bee 6: 0) 10/818 074 eee ,0)0)0010:0,4\0) 0.001010 vee plaice ecaisie in
XIX.—The Effects of Bee Venom. Reprinted from Zhe Indian
Medical Gazette, Vol. LXII, No. 5, May 1927, By A. Bayley
de Castro, Junior Medical Officer
XX.—Plants and Insects. By H. Jouguet
XXI.—Weeds of the Indian Wayside. By H. J ouguet
PROCEEDINGS AND STATEMENTS OF ACCOUNTS
fOr eee rea reese c sens tec ceecccres see
794
795
796
796
797
798
800
800
802
805
807
810
813
es ‘EN iy
Spoodlusiu siusojesayy
BSNVYUD GAWDSN-WMOVIG SAHL
“OOS “LSIP “IVND 4 VUNOT
qo?
JOURNAL
OF THE
Bombay Naturai History Society
May, 1928 VoL. XXXII No. 4
THE GAME BIRDS OF THE INDIAN EMPIRE
BY
a CaS TUART sbAKER, B.Z.S.,. EF. S., M:B-O:U., H.F.A.0.U.
VOIE., = W
THE WADERS AND OTHER SEMI-SPORTING BIRDS
EAR iy I
(With a coloured plate)
(Continued from page #07 of this Volume)
MEGALORNIS NIGRICOLLIS
The Black-necked Crane
Grus nigricollis Przewalski, Mongd. Tang., li, p. 135 (1876),
(Koko Nur.)
Vernacular names.—Tung-du, Trung-dung. (Tibet).
Description.—Lores and crown dull red bare skin, sparsely
covered with coarse black hairs: a patch of feathers below and
behind the eye white, sometimes extending in a narrow line over
the eye; rest of head and neck black ; wing-guills dull black ; the
innermost prolonged secondaries black; inner webs of outer
secondaries black or mottled with black and many of the greater
coverts over the secondaries more or less black on the inner webs ;
scapulars sometimes with a little black mottling on their terminal
quarters ; remainder of plumage pale ashy-grey, the shafts of the
feathers of the upper plumage showing as fine darker brown lines ;
tail dark blackish-grey, tipped and edged paler grey.
Colours of sott parts.x—‘‘ Iris yellow; bill horny-grey or horny-
green, more yellow towards the tip ; legs and feet black ’’ (Ludlow. )
Measurements.—Wing 625 to 668 mm.; tail 234 to 947 mm.; tarsus
223 to 252 mm.; culmen 124 to 137 mm.
Distribution.—Ladak, Tibet, Setchuan and Yunnan, where it was
procured by Forest,
618 JOURNAL, BOMBAY NATURAL, HIST. SOGIETY, Vol eon
Niditication.—The Black-necked Cranes’ nest and eggs were
first discovered by Capt. R. Steen in 1905-6 at the Hramtso Lake
in Tibet at an altitude of about 14,700 ft. Later Capt. J. Kennedy,
Major F. M. Bailey and others found it breeding in some numbers
about various Tibetan lakes and Osmaston also found it breeding
in the Ladak Lakes.
The nest varies considerably. Often it is a mere scratching in
the ground with a pretence of lining, just afew reeds and rushes
forming a bed for the eggs tolie on. At other times amore
pretentious nest is made and quite a pile of grass, rushes and other
weeds is accumulated having a good depression in the centre for the
eggs. The bird breeds on the same ground as the Bar-headed Goose
and sometimes its own nest is surrounded by the large down-
covered nests of these birds. The eggs are two in number and closely
resemble those of the Common Crane except in being much bigger.
Most of those I have seen arerather dingy olive-brown or olive-
green eggs scantily blotched with dull reddish-brown or _ purplish-
brown with secondary, or underlying, markings of purple grey or
reddish-grey. TTweive eggs average 101:2 & 64:1mm.; maxima 105-0
xX 63°4mm. and 103:2 & 69:1 mm.; minima 96:4 « 64:1 and 39-3
xX 59°6 mm.
The breeding season is from the end of May to the first week in
July, most eggs being laid between May 25 and June 15.
Habits.—Walton found this crane: very wild and difficult to
approach in the Gyantse plateau but much more tame round about
Lhasa, though it seems equally common in either place. During
the breeding season it is constantly found in company with Bar-
headed Geese and looks very curious as it stalks about, its black
head and neck towering above the very white geese. It keeps
entirely to the shores and islands ofthe lakes and to the wide open
plateaus, feeding both on vegetable shoots and on all kinds of
insects, smallreptiles, etc. Ludlow describes its call as very like —
that of the Common Craneand its flight is said to be powerful and
graceful.
MEGALORNIS LEUCOGERANUS
The Great White or Stberian Crane
Grus leucogeranus Pallas, Reise Reich. Rus., ii, p. 714 (1773);
(Irtin and Ob Rivers.) ; Blanf. and Oates, iti, p. 187.
Vernacular names.—Kare Khar. (N.W.P.), Tunki(Oudh), Chin
Kulang (Hansi, Hind.) ; Burmuch (Behar).
Description.—W inglet, primary coverts and primaries black ; fore-
head, fore-crown, face and sides of head bare except for a few white
bristles ; remaining plumage pure white.
Colours of soft parts.—Naked skin of head reddish; iris bright
pale yellow; bill umber brown; legs and feet pale reddish-pink.
(Hume).
Measurements.—Wing § 635 to 645 mm.; & 538 to 620 mm. ; tail
205 to 218 mm. ; tarsus 275 to 285 mm. ; culmen 185 to 197 mu
Young birds have the whole head clothed with white feathers and,
according to Hume, have the white plumage tinged with buff,
THE GAME BIRDS OF THE INDIAN EMPIRE 619
Distribution.—South-eastern Europe, Asia Minor, through Siberia
to Japan. In winter migrating to India and North China.
Niditication.—There is very little known of the breeding of this
fine crane. It is said by Godlewski to bethe most wary and shy
of all cranes, inhabiting the steppes near the large yivers, arriving
at its breeding haunts in April and leaving again in September.
Dresser gives the tollowing as known breeding places. Russia on
the lower Ural and in the Perm Government (both uncommon) ; in
the Tolol and Targai Governments, not uncommon; in the northern
parts of Turkestan ; Dauria on the Vilnui River ; in the delta of the
Kolzma; Amur, Ussurito N. E. Mongolia. As regards its nest
the only note I have is from Kuschel who in sending a pair of eggs,
writes, ‘‘ I have another pair, both taken by a Russian collector from
nests of rushes and reeds on the ground by a lake.’’ Thetwo eggs
Kent me measure 98:9 « 54-6. and 95:0 x 62:0 m. In colour
these have a rather bright reddish-olive ground, well blotched with
dark brown primary and purple-grey secondary markings. They
can be matched by many eggs of the Common Crane. An egg
received by Dresser from Bianchi was paler than most eggs of that
bird, but unusually heavily blotched, whilst another laid in Lord
Lilford’s grounds was a darker colour than most. These two eggs
measured 92°7 X 61:7 and 101°3 XK 63:2 mm. Other eggs laid in
the aviaries of Mr. St. Quentin seem to closely resemble these.
Finally, a pair of these eggs from Bianchi was also once exhibited
by Dresser ata meeting of the B.O.C.! but not described or
measured.
June seems to be the month when most eggs are laid, whilst two
seems to be the normal clutch. As, however, with many other
cranes, often only one young survives and Hume noted that the
old birds on arrival in India usually had only oze young one with
them.
Flabtts.—The Siberian Crane arrives in North-west India about
the middle of October, sometimes a little earlier, sometimes not
until the third week of that month. Further south they appear still
later, not reaching their extreme southern limits until the last week
in October or even early November. It regularly visits most parts
of North-west India and spreads in lessening numbers as far as
the Najatgurh Swamp near Delhi on the south to Jainagar in Be-
har on the east (where it was seen by Inglis) and to Kohee, twenty
miles south-east of Kamptee near Nagpur. Nowhere does it
arrive in vast multitudes like the Common and Demoiselle Cranes
but only in small flocks seldom numbering more than twenty
individuals and often consisting of one or two pairs and their young
ones.
This Crane is said to be the most elegant and graceful of all our
cranes. Hume says of it, ‘‘ It is the lily of birds and stand in what
position it may, the entire outline of its form presents the most
graceful and harmonious curves.’’
It is exceedingly wary and shy and the utmost patience and care
are required for a successful stalk. Moreover the birds keep
LUBULL: BO. Ce, VOL, =X1, pixod:
620 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
entirely, or almost so, to places where there are no facilities for
stalking. They stand or wade all day long in shallow swamps or
lakes well away from cover and if there are beds of tall reeds and
rushes round the edge of these same pieces of water, be sure the
birds will be nowhere near them. Hume, who tried all kinds of
ways to get within shot says there is but one way of doing so.
‘“ You obtain from one of the local fowlers a trained buffalo, and enter
the water a good quarter of a mile away from the bird, under cover
of this quadruped. It has, as usual, a string run through the nos-
trils and tied lightly together behind the horns. You hold this
string where it lies across the cheek with the left hand; your ex-
tended left arm is hidden behind the neck: your whole body is
bent, so that your head and shoulders are covered by the buffalo’s
shoulders, your body and the greater part of your legs, by its body.
Only your legs to a little above the knees show close to the hind
legs; and, as far as possible, you always keep the beast up to his
belly in water. Thus covered you slowly sidle up to the Cranes,
making the buffalo, now put hisnose up, head in air, now stop and
lower his head to the water, and generally dawdle and meander
about without apparently any fixed idea in his head, accordingly to
the natural manner and customs of a free and independant buffalo.
With a little practice it is easy thus to get within shot. You softly
let the cheek strings go and at once fire below the buffalo’s neck.
Before your gun is well off, your sporting companion, who has a
marked distrust of Europeans and white faces, and has been inces-
santly endeavouring to kick you throughout your whole promenade,
knocks you head over heels, and rushes off towards his dusky
owner, bellowing as if he, and not you, wete the injured party.”’
When not kiiled outright it defends itself stoutly with legs, wings
and bill,but chiefly with the latter, against either human or other foes.
They feed principally on aquatic plants and seldom resort to
cultivated dry plains, at all events during their stay in India, nor
have the remains of any animals or insects been found in their sto-
machs. Hume remarks on the remarkable number of small pebbles
often found in the stomachs of these birds. Out of an old male he
took enough to nearly fill a wine glass, the pebbies being mostly
quartz or green stone ranging in size from a pin’s head to an ordi-
nary pea.
Their flight is similar to that of the ordinary crane and though
they seldom soar, they do so occasionally with ease and grace.
Apparently they have no trumpet call like most cranes and their
cry has been likened to a repetition of the syllables kayekhur rapidly
repeated whilst their usual note when feeding and undisturbed is a
feeble chirrup.
MEGALORNIS MONACHUS
The Hooded Crane
Grus monacha Temm., Pl. Col., pl. 555 (1835) (Yesso and
Korea).
Vernacular nanes.—None recorded.
Description.—Lores, forehead and fore crown covered with black
bristles ; remainder of head and neck pure white; all the rest of the
THE GAME BIRDS OF THE INDIAN EMPIRE 621
plumage dark slaty-grey, more or less tinged with brown, especially
on the upper parts; edges of the feathers both above and below
fringed with grey, these almost disappearing in abraded plumage ;
quills, the decomposed ends of the drooping secondaries and the
tail blackish-brown.
Colours of soft parts.—‘* Iris yellow ; bill greenish tinged with red
towards the base ; feet dull reddish ’’ (Blyth).
Measurements.—Wing 510 to 545 mm. ; tail 155 to 180 mm. ; tarsus
208 to 220 mm.; culmen 92 to 110 mm.
Young birds appear to be a much paler grey; the head is well
covered with bristly feathers, grey with black shafts on the crown,
whiter on the forehead and black on the lores and in a patch on
either side of the forehead.
Distvibution.— Breeding in Japan and Eastern Siberia, migrating
south to China in winter. The only quite certain record of this
crane’s occurrence in India is that of a young bird shot by me in
North Cachar in December, 1899, but I saw several of these cranes
on two occasions in the lower reaches of the Subansiri in N.
Lakhimpur. Hume saw a small flock of cranes in Manipur which
must have been this species and other probable occurrences are
recorded by Anderson at Ponsee, west of Bhamo, and by Couchman
near Myothil in Upper Burma.
Nidttication.—I can find nothing on record beyond the statement
that it undoubtedly breeds in Dauria, Amur and in Eastern Siberia
and that it arrives in its breeding haunts in April and leaves again
in August.
Habits.—This crane is said to haunt open plains and marshes
but to be nowhere very numerous, whilst even when migrating it
collects in small parties only. According to David and Oustalet it
migrates in pairs or in flocks of half a dozen to a dozen. The flock
I saw in Cachar numbered seven, and those I saw on the Subansiri
were in two flocks of seven and eight respectively. In each of
these instances the birds were on the move and though I shot one
in Cachar I could get nowhere near those on the Subansiri, though
I spent a whole day trying to get at one flock which kept moving
down stream and settling every half mile or so. The other flock
rose when first disturbed and went right away at once, flying very
high in a line and loudly trumpeting at the start. The flock I saw
in Cachar seemed very tired as if they had flownfar. They pitched
very headlong on the banks of the stream I was boating down and
Stood still as if exhausted. Even after I had shot and secured one,
the remaining six kept pitching among the short reeds on the river
side, though they would not allow me to get within a hundred yards.
Their call seemed to me much like that of the Common Crane and
their flight also very similar. They rose with perfect ease from the
ground and at once rose to a considerable height.
David and Oustalet comment on the affection shown by a pair of
these birds, the one for the other, and narrate how the male of one
pair being shot, the female descended to the ground and attempted
to assist him to rise.
(To be continued.)
REVISION OF
THE FLORA OF THE BOMBAY PRESIDENCY
BY
Ey BLATTER, S:J.; PhSD., at--s.
PART VI
GRAMINEAt
BY
E. BLATTER and C. McCann
(Continued from p. 435 of this Volume)
38. HETEROPOGON, Pers. ; Stapf in Prain Fl. Trop. Afr. ix, 410.
Perennial or annual grasses, with simple or more often upwards branched
culms ; branches few to many, mostly flowering and gathered into a spatheate
panicle; racemes conspicuously dorsiventral, the bases of the male (or neuter)
spikelets subimbricate on the back of the raceme, their upper parts bending
forward around the sides, forming a hollow in which the fertile spikelets are
enclosed, with their awns exserted anticously and often intertwisted. Spikelets
2-nate, those of the lower (1 to many) pairs alike in sex and shape, male or
neuter, of the upper pairs differing in sex and shape, one of each pair sessile,
the other pedicelled on the many-jointed rhachis of solitary racemes, terminat-
ing the culms and their upper branches; rhachis, tough or upwards tardily
disarticulating and glabrous between the homogamous pairs, readily disarticulat-
ing above them; homogamous pairs long-persistent, the spikelets of the
heteroyamous pairs falling separately, the pedicelled with the pedicel, the
sessile with the adjacent joint and the adjacent pedicel or its base. Sessile
spikelets subcylindric, awned ; callus long, pungent, densely bearded upwards.
Involucral glumes equal, the lower coriaceous rarely chartaceous, more or less
tightly involute, quite keelless, nerves obscure, often connected by few trans-
verse nerves in the upper part ; upper with a deep longitudinal groove on each
side, coriaceous, rarely chartaceous, between them, thinner towards the margins,
membranous at the tips, 3-nerved. Lower floral glume hyaline, nerveless, upper
stipitiform from a hyaline very slender base, cartilaginous upwards and passing
into a usually stout geniculate awn. Pale small or absent. Lodicules large or more
or less reduced, to very minute. Stamens 3, often rudimentary or absent. Stigmas
exserted terminally or laterally. Grain more or less linear in outline, subterete,
slightly dorsally compressed ; embryo somewhat exceeding the middle of the
grain. Pedicelled spikelets male or neuter, dorsally flattened, usually slightly
asymmetric, and often somewhat twisted, muticous, imbricate. Lower involu-
cral glume herbaceous, many-nerved, winged upwards from one or both keels ;
upper membranous, lanceolate-oblong, acute, 3-nerved. Floral glumes
hyaline, 1-nerved, well developed or more or less reduced. Stamens 3 or 0.
Species about 6, in the tropical and subtropical regions of the whole world.
Cke. describes four species under Andropogon, viz. A. polystachyos, Roxb.,
A. triticeus, R. Br., A. Ritchiet, Hook. f., and A. contortus, Linn. We trans-
fer these to Heleropogon and add Heteropogon oliganthus, Blatter & McCann.
A. Not more than 30 cm. high
I. Upper involucral glume of pedicelled
spikelet 1-3-nerved es a. 1-37. f0licaniiins:
Il. Upper involucra! glume of pedicelled
spikelet 5-nerved ee .. 2 H. polystachyos.
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 623
B. More than 40 cm. high
I. Lower involucral glume dorsally deeply
grooved ‘ HT. insignis.
II. Lower involucral glume not dorsally
grooved
1. Ligule of several narrow membranous
segments sc 50 set 4s EL Rechter.
2 Juigule trineate; ciliolate .... ua Oa fle CONLOTIUS.
1. Heteropogon oliganthus, Blatter & McCann, nov. comb.— Andropogon
oliganthus, Hochst. ex Steud. Syn. Gram. 368; Hook. f. in F.B.I. vii, 201.
Description: A dwarf annual. Stems 7-15 cm. high, tufted, slender, some-
times creeping below, geniculate. Leaves 5-7°5 cm. long, quite flat, subensi-
form, acuminate, softly hairy, margins thickened, cartilaginous, sheaths
compressed ; ligule very short, membranous. Spikes usually very shortly
exserted from the long, narrow, glabrous spathes ; peduncle curved, puberulous.
Spikes 8-16 mm. long, with 3-6 pairs of glumes, lower spikelets neuter. Joints
and pedicels about half as long as the sessile spikelet, long-ciliate. Sessile
spikelet 5mm. long, oblong; callus short, obtuse. Lower involucral glume
obtuse, chartaceous, dorsally villous toward the base, not dorsally channelled,
shining, faintly nerved, with narrowly involute margins, ciliolate towards the
truncate tip, callus bearded, upper involucral glume obtuse, apiculate. Lower
floral glume very short, broadly oblong, ciliate, upper with a shining awn 3-4 cm.
long, column of awn glabrous. Pale very small. Anthers 1mm. long. Pedi-
celled spikelet, neuter, larger than the sessile, ovate-oblong. Lower involucral
glume herbaceous, obovate-oblong, truncate, distinctly 9-13-nerved, subemargi-
nate, keels scabrous, imargins ciliolate, hard!y incurved, upper shorter by },
membranous, oblong, obtuse, 1-3-nerved, ciliate. Pale of upper floral glume
very small.
Hackel, l.c., is of opinion that this species is intermediate between Dichan-
thium and Hetercpogon, but comes nearer the Hleleropogons, because the
sessile spikelets differ from the pedicelled ones in shape, nervation and the dorsal
furrow.
Locality: Deccan: Kalsubai (Patwardhan!); Mahableshwar, open edge
above the precipices looking from Dhobi’s Waterfall path to Elphinstone Point,
elevation 4,500 ft., rainfall 270 inches (Sedgwick & Bell 4608!) ; Panchgani
(Blatter & Hallberg B1221!, McCann!).
Distribution : Nilgiris.
2. Heteropogon polystachyos, Blatter & McCann. comb. nov.—Andropegon
polystachyos, Roxb. Fl. Ind. i (1832), 261; Steud. Syn. Gram. 367; Hook. f.
in F.B.I. vii, 989; Dalz. and Gibs. 301; Cke. ii, 989.—A. pumtlus, Roxb. Ic.
ined. t. 2021 (ex Hook. f.).
Description : Cke. 1.c.
Locality: Deccan: Khandala (Woodrow) ; Mahableshwar, western side of hill
(Dalzell & Gibson). We have not seen this species,
Distribution : Peninsular India.
3. Heteropogon insignis, Thw. Enuin. Pl. Zeyl. 437; Benth. Fl. Austral. vii.
517.—Andropogon triticeus, R. Br. Prodr. (1810), 201 ; Hack. Monogr. Androp,
588; Steud. Syn. Gram. 368; Hook. f. in F.B.I. vii, 200; Cke. ii, 989.—A.
ischynanthus et liananthus, Steud. 1|.c. 367.
Description : Cke. luc.
Locality : Konkan : Above Kenery Caves (McCann 9634 !).— Deccan : Around
Vital Hills, Poona (Bhide 782!) ; Chattarshinji Hill, Poona (Ezekiel!) ;
Khandala, very common on open hillside composed of rock fragments (McCann
9425!) ; Igatpuri (McCann 4338!) ; Mawal, Poona District (Woodrow).—S. J7/.
Country: Manoli (Talbot 3978 !).—Aanara: Anmod, bare hillsides, elevation
2,000 ft., rainfall 200 inches (Sedgwick 3324 !).
Distribution: India (Burma, C. Provinces, W. Peninsula), Ceylon, Malaya,
Australia.
4. Heteropogon Ritchiei, Blatter & McCann, comb. nov.-Andropogon Ritchiei,
meek. f. in f:B.1. vii, 201; Cke. ii, 990,
Description : Cke. 1. ¢,
624° JOURNAL, BOMBAY NATOCRAL AIST SOCEITY Vol ees
Locality : Deccan : Katraj Ghat, eleven miies S.E. of Poona (Gammie 1037 !) ;
hills near Poona (Woodrow).—S. M7. Country: Belgaum (Ritchie teste
Hook. f.).
Distribution : W. Peninsula, apparently endemic.
5. Heteropogon contortus, Roem. & Schult. Syst. Veg. ii, 836; Stapf in Prain ;
Fl. Trop. Afr. ix, 411.—Azndropogon contortus, Linn. Sp. Pl. (1753), 1045 ;
Hack. Monogr. Androp. 585 (excl. A. polystachyus, Roxb.) ; Hook. f. in F.B.I.
vii, 199 ; Cke. ii, 990. For other synonyms see Hook. f. and Stapf 11, cc.
Vern. Names :—Sunkhali, Nani Sunkhali (Dohad); Survalu (Charodi) ;
Kursali (Poona) ; Kusal, Sukhli kursali, Ganjali hullu (Belgaum) (ex Burns).
Known to Anglo-Indians as Spear grass.
Description : Cke.1.c.
Locality : Gujarat: Sevalia (Chibber!); road to Lasandra (Chibber !) ;
Sungiri (Gammie 15586!) ; Junagad Kathiawar (Blatter 3789!).— Khandesh :
Bhusawal (Gammie!) ; Toranmal (McCann 9640!).—Konkan: Dahe forests
(Ryan 717!) ; Osarvira forest, Mokhada range (Ryan 190!); Malabar hill
(McCann 3620!) ; Mulgaum (McCann 4245!) ; Parsik, railway line (McCann
9655!) ; above Kenery Caves (McCann 9662!).—JDeccan: Katraj Ghat,
11 miles S. E. of ‘Poona (Shevade!) ; Trimbak (Chibber!); Khandala, very
common all over the hills (McCann 9422!); Bairawadi, Purandhar (McCann
5059!) ; Rahuri (Nana A264!) ; Chattarshinji Hill, Poona (Ezekiel !); Manmad,
river bed (Blatter A269!) ; Igatpuri (McCann 4328!); Panchgani (Blatter &
Hallberg B1246!, B1296!, B1308!).—S.1/.Country: Dharwar, elevation
2,400 ft., rainfall 34 inches (Sedgwick 1820!) ; Castle Rock (Gammie 15686!) ;
Badami (Talbot 2925!).—Aanara: Yellapore (Talbot 734!).
Distribution : Mediterranean region and tropics and subtropics generally.
Uses: For an interesting account of this grass see W. Burns, L. B. Kulkarni
and S. R. Godbole: A study of some Indian Grasses and Grasslands, in Wem.
Dept. Agr. India xiv (1925), 28-44.
Varieties : Hackel (1.c.)distinguishes two varieties and,excluding polystachyus,
5 subvarieties. His first variety genwznus is characterized by the male spikelets
being more or less covered on the back or at least above or towards the margins
with white, patent tubercle-based bristles. This character together with the
degree of ramification of the culms forms the foundation of four subvarieties :
typicus, Roxburghii, hispidissimus and secundus.
The second variety glaber has the male spikelets glabrous. Both varieties
are represented in India, and all the subvarieties except secundus.
To Hooker f. (1. c.) these varieties and subvarieties appear ‘ to be too in-
constant for definition’, and, according to Stapf (l.c.) the inconstancy of those
characters ‘is so evident that it is not worth while to discriminate between the
forms corresponding to them.’ Haines, too, has abstained from distinguishing
varieties and forms.
Burns and others in the paper quoted above (p. 40) wrote in 1925: ‘In our
observations at Kalas and elsewhere we noticed variability within the species,
and early came to the conclusion that there must be definite varieties of Andro-
pogon contortus. We can say with confidence that there are at least two
varieties, differing markedly in size, habit, longevity, and in morphological
characters. Oneis small and annual, the other large and perennial.’ Since
then Patwardhan and Hedge have published a paper? in which they describe in
detail the morphology, anatomy, physiology and ecology of the two varieties.
As the authors themselves identify their perennial variety with Hackel’s
subvariety ¢ypzcws, and their annual variety with Hackel’s hispidissimus, we
shall refer to them in the following under Hackel’s names. The description is
taken entirely from the paper just mentioned.
Key to the varieties.
1. Lower involucral glume of pediceiled
spikelet sparsely hairy with long tuber-
cle-based deciduous hairs on the back,
in the upper part and towards the
margins ; lower part glabrous .. (a) var genuinus, subv.
ty picus.
1G. B. Patwardhan and G. R. Hedge, Two varieties of Andropogon contor-
des, Linn. In Journ, Ind, Bot. Soc, vi (1927)).213;
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 625
2. Lower involucral glume of pedicelled
spikelet densely hairy with tubercle-
based persistent hairs all over the back ;
hairs in lower part shorter than in
upper bis sae . (0) var. genuinus, subv.
his pidissinius.
(a) var. genuinus, suby. typicus, Patwardhan and Hedge 1.c.; Hack. in
Monogr. Androp. 586 (swb Androp.).
Description: Perennial. Stems densely tufted, 45cm.to 1:2 m., suberect
or decumbent, about 3 mm. thick at the base and much flattened, glabrous,
simple or subfastigiately branched from the nodes of the upper half; inter-
nodes not much longer than the leaf-sheaths. Leaves crowded at the base;
sheaths smooth, compressed, keeled, shortly auricled at the mouth; ligule
short, truncate, ciliolate; lamina 5-35 cm. long, acuminate or abruptly so, at
first folded inward, afterwards flat, rigid, suberect, ciliolate or sparingly hairy
on the upper surface, with tubercle-based hairs towards the base, rough to
the touch on both surfaces. In semi-dry or dry specimens the leaves may be
vinaceous to deep vinaceous in colour. Spike solitary, terminal, 2°5-7'5 cm.
long excluding the awn, with closely imbricating pairs of sessile and pedicelled .
spikelets. Spikelets subsecund, pedicelled ones all male or neuter ; of the ses-
sile spikelets the lower 2 to 8 male or neuter awnless, the upper awned, female ;
the lower awnless spikelets persistent, the upper awned ones. deciduous.
Sessile spikelet: Lower involucral glume narrow, linear-oblong, truncate
or rounded, brownish, many-nerved, hispid with short sparse hairs,
margin incurved, tip membranous; upper involucral glume linear, obtuse,
coriaceous, dark brown, hispidulous, 3-nervec, margins incurved. Lower
floral glume oblong, hyaline, thin, nerveless, short, truncate, paleate; upper
empty, reduced to an awn about 7‘5cm. long, column slightly hairy, callus
long, pointed, with atuft of reddish brown hairs. Ovary linear, stigmatic
branches from 6-8 mm. long, with thick-set hairs, carmine in colour, fading
towards the tips. Pedicelled spikelet: Lower involucra! glume lanceolate,
obliquely twisted, sparsely hairy with long, tubercle-based, deciduous hairs on
the back in the upper part and towards the margins, lower part glabrous, mar-
gins unequally winged ; upper involucral glume oblong-lanceolate, acuminate,
o-nerved, ciliate towards the tip, margins hyaline. Lower floral glume oblong-
hyaline, l-nerved, ciliate, epaleate, containing stamens ; upper empty, obovate,
lanceolate, hyaline, ciliate, nerveless. Stamens 3, filaments 3mm. long; anthers
oblong-lanceolate, cordate at base, pale yellow when young, light purplish
vinaceous; pollen grains round, not sculptured, dull grey. The sessile male
spikelets in the lower part are similar to the pedicelled spikelets above. Fruit
9-8 mm. long, thinly hispid, dark brown with 2 deep furrows on the ventral
side, armed with a pointed callus below and a long lash-like awn above.
Locality: ‘'Tegur (ex Burns and others).
Distribution: Tropics and subtropics generally, N. W. Himalaya, Afgha-
nistan (ex Hackel).
(6) var. genuinus, subv. hispidissimus. Patwardhan and Hedge l.c.; Hack.
in Monogr. Androp. 587 (swb Androp.).—Andropogon hispidissimus, Hochstett.
in Schimper Pl. Abyss. un. it. no. 1219.—A. desukiensis, Steud. in Zoll.
Syst. Verz. 59.
Description: Annual. Stems little tufted, 15-60 cm. high, erect or slightly
decumbent below, slender and slightly compressed near the base, glabrous,
simple or subfastigiately branched from the upper 2 or 3 nodes; internodes
much longer than the leaf-sheaths. Leaves little crowded atthe base ; sheaths
smooth, slightly keeled, shortly auricled at the mouth ; ligule short, truncate
and ciliolate ; lamina 2°5-20 cm. by 2°5-5 mm. wide, linear, acuminate or
abruptly so, ciliate or hairy onthe uprer surface up to nearly half its length
from the base with tubercle-based hairs, rough to the touch on both surfaces,
in semi-dry or dry specimens straw to very light purple in colour. Spike solitary,
terminal, 1-5 cm. long excluding the awns, with closely imbricating pairs of
sessile (lower) and pedicelled spikelets. Spikelets subsecund, pedicelled, all
male or neuter ; of sessile the lower 2-5 male or neuter, awnless, upper 4-12
female, awned. Those which are male or neuter persistent, the upper 4-12
pairs which include both males and females deciduous. Sessile spikelet :
Lower involucral glume narrow, linear-oblong, truncate or rounded, brownish,
2
626 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. aH
many-nerved, densely hairy with whitish long hairs, margins incurved, tips
membranous ; upper involucral glume linear, obtuse, coriaceous, dark brown,
hispiduious, 3-nerved with incurved margins. Lower floral glume oblong,
hyaline, thin, nerveless, short, truncate, epaleate, upper empty, reduced to an
awn, 3S'1 cm. long, columu densely hairy with whitish hairs, callus long,
pointed, bearded with light brown hairs. Ovary linear, stigmatic branches
4-5 mm. long with thinly set ox-blood red hairs. Pedicelled spikelet : Lower
involucral glume lanceolate, obliquely twisted, densely hairy with tubercle-
based, persistent hairs all over the back, hairs on lower part shorter than in
upper; upper involucral glume oblong-lanceolate, acuminate, 5-nerved,
ciliate, tip hairy, margins hyaline. Lower floral glume oblong, hyaline,
l-nerved, ciliate, upper oblong or obovate-lanceolate, hyaline, nerveless.
Stamens 3, filaments 4mm. long; anthers oblong, sagitate, pale yellow when
young, sulphur yellow when mature; pollen grains round, smooth, dull grey.
The sessile spikelets in the lower part are similar to the pedicelled spikelets
above. Fruit 5-8 mm. long, densely hairy with whitish soft hairs, light brown
with two deep furrows on the ventral side, armed with a pointed callus below
and a long, lash-like awn above.
Locality: ‘ Usually found on the very shallow parts of Deccan soils’ (ex
Burns and others, l.c.).
Distribution : Malabar, tropical Africa, Madagascar, Java, Phillippines.
39. IskiLEMA, Hack. ; Cke. ii, 995.
Cke. has described two species: /. Wightiz, Anders. and J. daxum, Hack.
We adda third one, /. anthephoroides, Hack.
The following key is after Hole? :—
A. Lower involucral glume of hermaphrodite
spikelet dorsally appressed, hairy at base and
ciliate on margins in basal } .. Ll. L. anthephoroides,
B. Lower involucral glume of hermaphrodite
spikelet glabrous dorsally at base and on
margins in basal 4.
I. Spathe and upper floral leaf not tubercled on
keel RE CI BOR AIL
II. Spathe and upper floral leaf tubercled on
keel ee AL AN
1. Iseilema anthephoroides, Hack. Monogr. Androp. 683; Hook. f. in. F.B.I.
vii, 219 ; Haines Bot. Bihar & Orissa, 1054.
Description: A much tufted annual grass, very leafy below, with many
stems 30-80 cm. high, Sometimes pink, nodes glabrous. Leaves mostly short,
the longer ones about 13 cm. by 5 mm., subobtuse, ciliate at base and tip of
sheaths, cilia with small tubercle-bases, blades with scabrid margins, nerves
usually fine and uniform; ligule of short fine hairs. Panicles long, rather
strict, but some of the spatheoles divergent, lower spathes foliaceous, upper
with shorter blades, base of blade and top of sheath with very long cilia ;
spathes often with many tubercles on the margins. Spatheoles cymbiform,
not acuminate, smooth and glabrous or minutely tubercled and scaberulous,
margins scarious, several spatheoles from each spathe or leaf-sheath. Peduncle
of cluster very short, 2°5-5 mm. long. Cluster scantily bearded at the base.
Involucral spikelets broadly oblong, 4 mm. long, rounded at tip, not or very
sparsely ciliate, their pediceis about 1mm. long and nearly as broad at top,
compressed, bearded. Glumes 2 only. Lower involucral glume with narrowly
inflexed margins, strongly 2-nerved on the back and almost sulcate on either
side of midrib, 2 other partial nerves between the strong ones, upper involucral
glume nearly as long, flat, oblong, obtuse, 1-nerved; floral glume absent.
Anthers yellow. Sessile spikelet 5-6 mm. long, suddeniy tapering part or
beak rather longer than the lower broader part. Lower involucral glume
2-cuspidate at tip, 4-nerved between keels, hispid-hairy on the back on the
wider portion, the beak scabrid or scabrellous ; upper involucral glume as long,
1R. S. Hole, The Indian Species of J/sezlema, Agric. Journ. of India,
Special Ind. Sc. Congress Number (1917), 125-131. See also A. Camus, Le
genre /sezlema, in Bull. Soc. Bot. France, series 4, xxiii (1923), 493,
i
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 627
narrowly lanceolate with prominent ciliate keel on lower third, scabrellous
above, margin inflexed. J.ower floral glume very narrow, 2-nerved, ciliate,
upper reduced to the membranous base of the awn, awn 12-i4 mm. long,
very slender, nearly smooth.
Locality: Khandesh: Near Naradana (Blatter & Hallberg 5206!) —
Deccan: Katraj Ghat (Gammie 929!) ; Deolali (Blatter & Hallberg A316!) ;
Chattarshinji Hill, Poona (Ezekiel!) ; Pashan (Gammie!).—S. WW. Country :
a fields EH. of Hubli (Sedgwick & Bell 5295!) ; VYelvigi (Sedgwick
2085 !).
Distribution: W. India and Deccan.
Uses: Asmaller yielder and an inferior fodder piant than /. laxum. See
W. Burns Bull. 78, Dept. Agric. Bombay, p. 11.
2. Iseilema laxum, Hack. Monogr. Androp. 682; Hook. f. F.B.I. vii, 218;
Cke. ii, 996. For synonyms see Hook. f.1.c
Description: Cke. 1. c.
Locality: Gujarat: Red upland near Talod (Sedgwick !).—handesh :
Bor, Bori River (Blatter & Hallberg 4429!) ; Toner, Tapti bank (Blatter &
Hallberg 5167!); Dadgaum (McCann A322!).--Konkan: Sion (McCann
3668 !); Thana (McCann 8728!); Parsik Hill (McCann A321!) ; Bhandup,
rice field (McCann A323!) ; Trombay {McCann A324 !).—Deccan : Grasslands
between Mahableshwar and Panchgani (Sedgwick & Bell 4742!) ; Nasik
(Bourke!) ; Lohagad, half way up (McCann A320!) ; Poona (Jacquemont
439).—S. M. Country: Dastikop (Sedgwick 2059!) ; Dharwar (Sedgwick
1826!) ; Belgaum (Ritchie 799).—Kanara: Halyal (Talbot 2087!) ; Sirsi
(Kulkarni!).
Distribution : Upper Gangetic Plain, Orissa, Deccan, W. Peninsula.
Uses : Considered to be the best fodder grass in Central and S. India.
3. Iseilema Wightii, Anders. in Nov. Act. Soc. Sc. Upsala ser. 3, 2 (1858), 251;
Iliack. Monogr. Androp. 679; Hook. f. F. B. I. vii, 218; Cke. ii, 996. For
synonyms see Hackel and Hook. f. 11. ce.
Vern, Names: Mabid (Dohad), Moshi (Surat), Gandhi (Charodi), Gandheli
(Panch Mahals), Sona, Tambrut, Tambit, Gondval, Gamsi, Mussau (ex Burns).
Description : Cke. 1.c.
Locality: Gujarat: Kharaghoda (Saxton 503C! ); in a very marshy valley
between Wastrapur and Thaltij (Sede wick 322!); Morvi, Kathiawar (Woodrow). —
Khandesh : Antroli, Bori River (Blatter & Hallberg 5150 !).—Deccan : Panch-
gani (Blatter & Hallberg B1294!); Poona (Woodrow).—S. MW. Country :
Kunnur (Sedgwick & Bell 4922!) ; Rhanibennur (Bhide!) ; Haveri (Talbot
2254!); Dharwar (Nana A325!); Belgaum (Wecodrow).—Kanara: Halyal
(Talbot 2143 !).
Distribution : Throughout India.
Uses: A fair fodder.
40. TiumMeEpbA, Forsk.; Cke. ii, 992.
Species about 16, in the tropical and subtropical regions of the Old World,
chiefly Indo-Malayan.
Cooke describes 4 species: 7. tmberbis, T. Cooke, 7. ciliata, Hack.,
T. cymbaria, Hack., and 7. tremula, Hack. Following the laws of priority we
have to substitute for the first two: 7. ¢rzandra, Forsk. and 7. guadrivalvis,
©; Kuntze.
I. Involucral spikelets truly verticillate
1. Involucral spikelets glabrous or more or
less irregularly beset with tubercle-based
hairs 1. 7. triandra.
2. Involucral spikelets. ‘always with a row of
stiff bristles along the flexures near the
tips, the bristles arising from coarse pale
tubercles 2. T. guadrivalvts,
Il. Involucral spikelets in closely. superposed
pairs
1. Inflorescence a decompound thyrsiform
panicle. Lower involucral glume of
bisexual! spikelets not channelled tah ae 3 Ls CYMOaTIG
628 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol2 XG
2. Inflorescence a racemiform prance:
Lower involucral glume of bisexual
spikelets deeply channelled ... ee the atvemulan
1. Themeda triandra, Forsk. Fl. Aegypt. Arab. cxxiii ef 178; Schweinf. in
Bull. Herb. Boiss. ii, Append. ii, 16,95; K. Schum. in Engl. Pfl. Ost.—Afr.
A. 51; Rendle in Cat. Afr. Pl. Welw. ii, 161; Pilg. in Engl. Pflanzenw. Afr. ii,
151, fig. 114; Pilg. in Mildbr. Wiss. Ergebn. Deutsch. Zentr.—Afr. Exped. ii, 45 ;
Eyles in Trans. Roy. Soc. S. Afr. v. 298; Stapf in Prain Fl. Trop. Afr. ix, 416,
partem tantum nostre speciei amplectens.-—-Themeda Forskalit, Hack. in Monogr.
Androp. 659, excl. syn. Anthistiria hispida, Thunb. gue est Tristachya
leucothrix, Trin. see. Stapf.; Duthie Fodd. Grass. N. Ind. 43; Durand &
S@hinz Consp. Fl. Afr. v. 731; K.Schum. 1.c. A. 23, 56, 79); Enel. Hlochee-
birgsfl. Trop. Afr. 115; Batt. & Trab. Fl. Anal) Algér. et Tunigis55. Stapn
in Kew Bull. (1907), 212; Pilg. in Engl. Pllanzenw. Afr. ii, 119.— Anthistiria
imberbis, Retz. Obs. iii, 11: Desf. in Journ. de Phys. x1, 293, t. 1; Thunb. FI.
Cap. i, 402, ed. Schult., 107 ; Kunth Enum. i, 481; Steud. Syn. Pl]. Glum. i, 401;
Stapf in Dyer Fl. Cap. vii, 366; Wood Natal Pl. ii, t. 133; Burtt Davy in
Transvaal Agric, Journ. 111, -287, t. 02%; Hook. 1. in EBT viiy (21 iri eee
Ceyl. v, 248; Prain Beng. Pl. 1207,.—A. glauca, Desf. Fl. Atlant. ii, 380, t. 254
excl. syn.— Stipa paleacea, Vahl; Coss, & Durieu Expl. Scient. Algér. ii, 52.—
A. Desfontainet, Kunth Rev. Gram. i, 161.—A. czliata, Retz. 1.c.(om Linn. f.);
Lamk. lll. t. 841, £. 2; Cav. Ic. 5, t. 459; Nees in Linnaea vii, 284 ezin Fl; Ate
Austr. 121; Anders. in Peters Reise Mossamb. Bot. 562; Oliv. in Trans. Linn.
Soc., xxix, 176% Roxb. Fl. Ind., 71,2475 Grab. Cat) BombiePl\ 23973 Dalz
Gibs. Bomb. Fl: 304; Thw. Enum. Pl. Zeyl’366; Trim. Cat. Ceyl, Pienose
Benth. FI. Austr. vii, 542.—4. Forskahliz, Kunth Rev. Gram. i, 162, Enum. Pl.
481.—A. vulgaris, Hack. in Engl. & Prantl. Nattrl. Pflanzenfam. ii, pars 2,
29.—A. punctata, Hochst. ex A. Rich. Tent. Fl. Abyss. ii, 448.—A. paleacea,
Ball. in Journ. Linn. Soc. Bot. xvi, 734.--A4. australis, R. Br. Prodr. 200.—
A. cuspidata, Anders.in Nov. Act. Upsal. 2 (1856), 229.—A. cespiteosa, Anders.
l.c. 241.--A. argentea, Nees Fl. Afr: Austr. 124.--Ad. depauperata, Anders, 1.c.
243.—A. syriaca, Boiss. Diagn. Pl. Or. ser. i, fasc. 13, 72.—Themeda polygama,
Gmel. Syst. 149.—Stzpa.arguens, Thunb. Prodr. 20 (zon Linn.).—Calamina
imberbis, Roem. & Schult. Syst. ii, 810.—Zhemeda imberbis, T. Cooke in
Cke. ii, 993.--Z. tmberbis, Haines in Bot. Bihar & Orissa, 1049.
Hooker f. who deals with thisspecies under Anthistiria, Linn., says ina
preliminary remark (F.B.I., vii, 211): ‘ Thespecies of this genus are most
difficult of discrimination, of which the best proof isthe irreconcilable conclu-
sions of two excellent botanists, both experts in the order of Graminec,
He then criticizes Hackel for having restored Forskahl’s name of 7Themeda
‘ because of its having four years of priority, and of Linnaeus’ description of
Anthistiria being very inaccurate.’ He admits the claim of priority, but as to
the other reason he rightly adds that if inaccurate description has to be consi-
dered, a host of the genera of old authors would haveto be invalidated. Hooker
finally decides in favour of Anthistiria because this genus ‘ had for upwards of
a century been adopted by all botanical writers.’ This reason, however, does
not hold good in view of the present rules of nomenclature. Hackel restored
Themeda in i889 and since then most systematists have followed his example.
It was easy to settle this point, but the real difficuity comes in when we have to
define and give aname toall the material gathered by Hackel (1. c. 659-664)
under his 7hemeda Forskalit, and by Hooker f. under Axthistiria tmberbis,
Retz. (F.B.1. vii, 211).
A glance at Hackel’s synonymy and localities shows that he has included all
the forms of this highly variable grass which are found in the tropical, sub-
tropical and sometimes in the temperate regions of the Old World. The same
applies to Hooker’s A. zmberbis, except that he separated Hackel’s var. dubia
/axa and restored it to its former specific rank of A. Jara, Anders. But this is
of minor importance in this connection. It does not change the fact that both
Hackel and Hooker describe the same material and of the same area though
under different names.
Hacke! justifies the adoption of the specific name 7hemeda Forskalii in this
way: ‘ Themeda triandra, Forsk. Fl. Aeg.-arab. p. 178, Anthist. Forskalii
Kunth Revis. Gram. 1, p. 162, generis typus, a Forskalio prope Bulgose in
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 629
Arabia felici lecta, probabiliter etiam varietas est Th. Forskalii meae; sed
descriptio rem dubiam relinquit, specimina authentica desiderantur. Nec
hucusque ullam hujus generis speciem in Arabia felici lectam vidi, etsi proba-
bile est, Th. Forskalii varietates in Syria Abyssiniaque crescentes etiam in Arabia
inveniri. Itaque nomen specificum ‘‘ triandra’’ tamquam dubium et rem indi-
cans qtiae in hoc et plerisque Graminearum generibus nullius est momenti,
seposui, ‘‘ Forskalii’’ a Kunthie datum non minus quidem dubium, sed generis
auctorem commemorans recepi.’
Hooker does notagree with Hackel, ‘ The earliest names of this plant are
Themeda triandra, Forsk., aud Aunthistiria imberbis, Retz. Hackel has abandon-
ed both, substituting first duthistiria vulgaris, and then Themeda Forskalit,
on the ground that A. zmberbis was perhaps not Forskahl’s 7. triandra (why
then call it Forskaliz?) of which no type specimen exists, and because ¢rzandra
indicates a character of noindividual value in grasses. In so doing he over-
looked Gmelin’s name of T. polygama (Syst. 149). Having regard to the wide
range of A. zmberbis, from Australiato Africa, its presence in Arabia might
well be anticipated ; and that itis anative ofthat country is now proved by
Schweinfurth’s finding Hackel’s var. glaucain that country. This makes the
var, (which is local, and not Indian) the type of the species, and if Forskahl’s
naime of 7henzeda is to be retained, necessitates a re-arrangement of the varieties.
To meit appears most expedient to retain Retz.’s name which applies to the
prevalent form over the area of distribution as the specific one. ’
Hooker’s reasons against Hackel’s view are certainly valid, but his own
Anthistiria imberbis does not restona firmer foundation. The fact that the
var. is local and not Indian should not prevent us from making it the type of
the species, and the other circumstance that a re-arrangement of the varieties
will become necessary if the name 7vemeda is retained, can only be a reason of
expediency.
So far we come to the conclusion that Hackel and Hooker deal with the same
material, but that neither name is satisfactory.
We come now to the latest publication affecting our question. Stapf l.c. has
adopted the name Zhemeda triandra, Forsk. (1775) instead of 7. Forskalii
(1889), ‘ as there is no doubt that the type of Forskal’s species, which apparently
has been lost, was one of the forms covered by the description given by Stapf’
(see Schweinfurth in Bull. Herb. Boiss. ii, App. IJ, 16).
But here Stapf creates a new difficulty. His description applies only to the
‘ African share of Hackel’s 7. Horskaliz’. In order not to be open to misstate-
ments we quote the whole passage in which Stapf explains his position: ‘ The
species, as defined here, is, however, taken in a sense somewhat narrower than
Hackel’s ; this restriction requires a short explanation. Hackel in his mono-
graph of the dAxdropogonee distinguishes eleven varieties and as many sub-
varieties or forms within his 7. Forskaliz, whilst other authors have at various
times described more than a dozen species, all of which come within the com-
pass. There can beno doubt as to the close affinity of these forms and the
question of their status is mainly one of expediency. A careful examination of
the large amount of materia! at Kew and the British Museum bas led to the
conclusion that for the present it will be most useful to detach, firstly, those
forms that are fairly uniform, and at the same time exclusive, over a large area ;
and secondly, those that, though of alimited range, stand out from the remain-
der by some character or characters. This leaves a residuum much less homo-
geneous than any of the segregates just referredto. It consists apparently of
more or less fixed races, mutants, hybrids and edaphic forms which from her-
berium material are the less separable because they are toa high degree inde-
pendent of geographical areas. At the same time, however, they aree all
African with an extension into Arabia, Syria andthe south-eastern corner of
Asia Minor, and, taken as a whole, represent practically the African share of
Hackel’s 7. Forskali. It isto this aggregate that the description and synonymy
given above apply.’
From the above it is evident that Stapf’s 7. ¢rizandra, Forsk. is not identical
with Hackel’s 7. Forskalii and Hooker’s A. imberbis, Retz., as it comprises
only the African element including ‘an extension into Arabia, Syria, and the
south-eastern corner of Asia Minor.’ Stapf’s synonymy !eads to the same con-
clusion, except for the inclusion of 7. zaberbis, T. Cooke (FI. Bombay ii, 993).
The Australian element Anthistiria australis, R. Br. has been separated by the
same author as Themeda australis, Stapf. It seems to us that Stapf’s treatment
630° JOURNAL, BOMBAY NATURAL AIST, SOCIETY, Vel Xoo
of 7. Triandra is somewhat arbitrary. We quite agree that the name should
remain and that itis the only correct name, but we cannot agree to its being
restricted to the African element only, and it is difficult to understand why the
Indian specimen should not go by thesame name. If we could distinguish
groups of varieties or forms that are confined to more or less definable geogra-
phical areas, it would be admissible to speak, e.g. of an African group and
call it 7. ¢riandra, because Arabia exhibits one of those African forms, or of an
Indian or Australian group, and name them accordingly. But experience
shows that with regard to the material under review there are.no such geogra-
phical areas which contain a group of varieties or forms that are peculiar to one
area exclusively. A glance at the localities given by Hackel under the different
varieties and sub-varieties will confirm our statement.
We are, therefore, of opinion that the name 7%emeda triandra, Forsk. should
embrace all the material that was described by Hackel under 7. Forskalii, by
Hooker under A. imzberbis, and by Cooke under 7. zmberbis.
Haines describes the material from Bihar and Orissa under the name of
T. imberbis, T. Cooke, and adds in brackets ‘partly’. His species, therefore,
is not Cooke’s 7. zmberbis, but must be given some other specific name as long
as Cooke’s namie stands. But if botanists adopt our 7. tviandra, Forsk. Haines’
name will be merged in it, and his material may be treated as a form or group
of forms under that species.
Description : See Hackel, Hook. f. and Cooke 11. cc.
Locality: Gujarat: Ahmedabad (Gammie 16391 !).—Khandesh: Toranmal
(McCann 9813 !).—Konkan : Mokhada range (Ryan 2626!) ; Mahaluxmi (Sabnis
A297!); Bhandup (McCann 9810!); Bassein (McCann 4475!).—Deccan :
Ganeshkhind Botanic Gardens (Garade!); Panchgani (Blatter 3806!. Blatter
& Hallberg B1326!); Khandala, common (McCann A291!); Bairawadi,
Purandhar (McCann 5069!); Igatpuri (McCann 4322!); Poondra (Talbot
4307 !).--S. M7. Country: Devaranji, elevation 1,800 ft., rainfall 90 inches,
(Sedgewick & Bell 4427!) ; Castle Rock (Gammie 15728!) ; Dudsagar Falls
(McCann A298!) ; Dharwar (Nana A289!).
Distribution : Africa, indo-Malaya, Australia.
2. Themeda quadrivalvis, O. Kuntze Rev. Gen. Pl. [I, 794; Stapf in Prain
Fl. Trop. Afr. ix, 420; Haines in Bot. Bihar & Orissa 1050.—Andropogon
guadrivalva, err. typog., Linn. Syst. ed. 13, 758.—Themeda ciliata, Hack. in
Monogr. Androp. 664; Cke. ii, 994.—Anthistiria ciliata, Linn. f. Suppl. 113;
Geertn. Fruct. ii, 465. t. 175 ; Lam. Til: t. 841, fie. 1; Beauv. Acrostst323/et. gee
Kunth Enum. i, 481; Steud. Syn. Pl. Glum.1, 401; Baker Fl. Maunit, 4437
Balf. f. Bot. Socotra 317. (partim) ; Duthie Grass. N. W. Ind. 42; Hook. f. in
F B.I.. vii; 213; Stapf in Dyer Fl. Cap. vii, 368:—4: scandens, Roxbe Pin Ing
i, 248; Duthie Fodd. Gr. N. W. Ind. t.61.—A. semiberbis, Nees. Fl. Afr.
Austr. 125.—Andropogon nutans, Linn. Mant. ii, 303.
Vern. Names: Bongrut (Sholapur); Bhathu (Surat); Bhati, Zini bathi,
Mothi bathi (Dohad) ; Bhatharu (Broach) (ex Burns).
Description : Cke. 1.c.
Locality : Gujarat: Ahmedabad (Gammie 16391!).—Kkandesh : Toranmal
(McCann 9817!).—Konkan: Parsik, railway line (McCann 9808!) ; Alibag,
water-works (Ezekiel!); S. Konkan (Stocks deste Cooke, Law); Salsette
(Jacquemont 717 feste Cooke).--Deccan: Pashan, near Poona (Gammie!) ;
road, Mahableshwar to Pratapgad (Bhide 1170!) ; Purandhar (McCann 5571!) ;
Bairwadi, Purandhar (McCann 507A!); Panchgani (Blatter & Hallberg
B1311!, B1325!).—S. W. Country: Konankeri, elevation 2,000 ft., rainfall 40
inches (Sedgwick & Bell 4943!) ; Dharwar. elevation 2,400 ft., rainfall 34
inches (Sedgwick & Bell 4486!); Dastikop elevation 2,500 ft., rainfall 35
inches (Sedgwick & Bell 2060!) ; Castle Rock (Gammie 15729!) ; Belgaum
(Ritchie 886 Zeste Cooke).—Kanara: Halyal (Talbot 2115!).
Distribution: N. W. India, Bengal, C. Provinces, W. Peninsula, Tenasserim.
Introduced in tropical and 8. Africa.
3. Themeda cymbaria, Hack. in Monogr. Androp. 668; Cke. ii, 994.—
Anthistiria cymbaria, Roxb. Hert. Beng. (1814), 6, e¢ Fl. Ind. i, 251 (excl.
syn.) ; Kunth Enum. Pl. i, 482 (excl. syn.) ; Grah. Cat. Bomb. Pl. 219; Daiz
& Gibs. Bomb, PF). 3043) Hook.f. inch Bos wit, 215% Trim. si Ceyliin, 225)
Description : Cke.1. c.
meyESION OF THE FLORA OF THE BOMBAY PRESIDENCY 631
Locality: Khandesh: Dadgaum (McCann 9815!); Toranmal (McCann
9818! ).—Konkan: common (Cooke); I am doubtful (McCann).—Deccan :
Lonavla (Garade!).—.S. 7. Country : Castle Rock (Gammie 15634 !).—Kunara :
Mirjan (Hallberg & McCann A295!).
Distribution : W. Peninsula, Ceylon.
4, Themeda tremula, Hack. Monogr. Androp. 667; Cke. ii, 995.-- Anthistiria
tremula, Nees ex Steud. Syn. Gram. 401 ; Hook. f.in F.B.I. vii, 214.—Andros-
cepia tremula, Anders. in Nov. Act. Upsal. ser. iii, ij (1856), 247.
Description : Cke. 1. c.
Locality: Konkan: Trombay (McCann A293 !); Ghatkoper, Horse-shoe
Valley (McCann A327!).—Deccan: Khandala (McCann 5359!) ; Igatpuri
(Blatter & Hallberg 5484!) ; Purandhar, northern foot (McCann 5041!) ;
Poona (Woodrow feste Cooke).—S. M7. Country : Devaranji, elevation 1,800 ft.,
rainfall 90 inches (Sedgwick & Bell 4428!) ; Castle Rock (Gammie 15687 !).—
Kanara: Devimani Ghat (Kulkarni!) ; Jugglepet (Talbot 1568!) ; Tinai Ghat
(Sedgwick & Bell 3196!) ; Sirsi to Siddhapur (Hallberg & McCann A294!).
Distribution: From the Central Provinces and the Konkan southwards,
Ceylon.
41, PSEUDANTHISTIRIA, Hook. f. ; Cke. ii, 992.
Species 4.—Indian.
Cooke has described one species: P. hispida, Hook. f. Weadd two others :
P. umbellata, Hook. f. and P. heteroclita, Hook. f.
I. Ligule exauriculate
1. Ligule atruncate glabrous membrane, much
divided to the base, the tips minutely ciliolate.
Sessile spikelets glabrous ees peed n Lod (LES DTCs
2. Ligule short, membranous. Sessile spikelets
not glabrous oe 2 is we. Ce We heterocizia.
II. Ligule auricled wae < aes .. 3. PF. umbellata.
J, Pseudanthistiria hispida, Hook. f. in F. B. I. vii, 219; Cke. ii, 992.
Description : Cke. 1.e.
Locality : Gujarat: Surat (Garade !).—Konkan : Dahe Forest (Ryan 705 !) ;
Matheran (D’ Almeida A329!) ; Ghatkoper, Horse-shoe Valley (McCann A326 !);
Sion (McCann 5247!) ; St. Xavier’s College compound, Bombay (McCann
4524!) ; Kalyan (Woodrow).—Deccan : Lonavla (Gammie!) ; Purandhar Fort
(Gammie 1010!); Igatpuri Ghats (McCann 4343!) ; Khandala, common,
railway line (McCann A331!) ; Panchgani (Woodrow) .—S. J7. Country : Castle
Rock (Gammie 15634, McCann A328!); Derikop (Sedgwick: 2061!) ; Londa
(Gammie 15863! , Woodrow).—Kanara: Birchy (Talbot 2096!) ; Yellapore
(Talbot 1522!) ; Devimani Ghat (Kulkarni!).
Distribution : C. Provinces, W. Peninsula.
2. Pseudanthistiria heteroclita, Hook. f. in F. B.I. vii, 219.—Andropogon
heteroclitus, Nees Fl. Afr. Austr. 115; Steud. Syn. Gram. 389; Hack. in
Monogr. Androp. 400.—A4. monomeros, Hochst. in Hvuhen. Pl. Ind. Or. no.
183.-—Anthistiria heteroclita, Roxb. FI. Ind. i. 249.
Description : Stems 30-80 cm. high, geniculate, slender, terete, smooth, sub-
simple or branched. Leaves linear, 15-30 cm. long, 3-5 mm. broad, glabrous
or more or less ciliate on both surfaces, nerves distinct, margins with long,
tubercle-based hairs or nearly glabrous; sheaths much shorter than the
internodes, quite glabrous; ligule short, membranous, exauriculate. Panicles
20-30 cm. long, leafy, compound, with many shortly peduncled fascicles,
fascicles of spikes about 12 mm. broad, proper Spathes 7-10 mm. long, hardly
longer than the spikes, towards the margin with long, tubercle-based bristles ;
spikes 6-8 mm. long. Sessile spikelets 3-4 mm. long, linear-oblong, hispidu-
lous all over. Lower involucral glume furrowed. Upper floral giume awned,
awn 18-24 min. long, thin. Pedicelled spikelets lanceolate, with a few long,
tubercle-based bristles, keel ciliate.
Locality : Konkan (Law ex Hook. f.).
Distribution : Bengal, Konkan, S. Kanara.
3. Pseudanthistiria umbellata, Hook. f. in F. B. I. vii, 220. — dndropogon
umbellatus, Hack. Monogr. Androp. 401.
632. JOURNAL; BOMBAY NATURAL AIST. SOCIETY, Vol. X2Cg
Description: A very slender, glabrous plant with filiform, prostrate, creeping
branched stems, rooting at the nodes, stems 30-60 cm. long, compressed.
Blade 2-5 cm. long, distant, linear-oblong, acute, rounded at base, sessile or
short-petioled, nerves distinct, very slender, with a few scattered cilia on both
surfaces, nearly smooth; sheaths shorter than the blade, often with tubercle-
based hairs above, rarely glabrous; ligule passing at the sides into 2 short,
herbaceous, fimbriate auricles of the sheath. Panicle leafy, very lax, interrupted,
12-20 cm. long ; fascicles of spikelets few, axillary, 6-12 mm. broad, glabrous
or with a few tubercle-based cilia on simple, rarely branched capillary pedun-
cles shorter than the leaves ; lower peduncles sometimes elongate, 2°5-7°5 cm.
long and bearing several fascicles ; outer spathes 8-25 mm. long; spikes 3-6 in
a fascicle, proper spathe 10-12 mm. long, rather longer than the spikes,
glabrous or towards the margin with a few bulbous-based hairs. Sessile
spikelets 3-4°5 mm. long, linear, scaberulous. Lower involucral glume
dorsally concave; awn of upper floral glume 12-14 mm. long, very thin.
Pedicelled spikelets linear-lanceolate, naked, awn 12-18 mm. long.
Locality: S. M. Country: Londa (Gammie 15869!).—Kanara: Birchy
(Talbot 2073 !).
Distribution : Deccan and W. Peninsula, Ceylon.
42. Di1GiTARtIA, Hall. Hist. Stirp. ii (1768), 244; Stapfin Prain Fl.
Wrop. Atrix; 422,
Species more than 100, in the warm parts of the whole world, but chiefly in
the Old World.
Cke. (ii, 940-942) describes 6 species. All are retained in this place, but the
name of Digitaria sanguinalis will be replaced by VD. marginata.
Key as in Cke.
1. Digitaria ternata, Stapf in Dyer FI. Cap. vii, 376, e¢ in Prain Fl. Trop. Afr,
ix, 452; Cke. ii, 940.—Paspalum ternatum, Hook. f.in F.B.I1. vii, 17.—
Panicum ternatum, Hochst. in Flora xxiv (1841), 1, Intell. 19; Hack. in OUest.
Bot. Zeitschr. (1901), 331.—P.. phe@nocarpum, var. gracile, Nees Fi. Afr. Austr.
23.—Cynodon ternatum, A. Rich. Tent. Fl. Abyss. ii, 405.
Description: Cke. 1. c. Occasionally long fine hairs are found on the
peduncles.
Locality: Deccan: Purandhar Fort (Bhide!).—S. M4. Country: Belgaum
(Herb. Bot. Gard. Cal.).
Distribution : India (Khasia Hills, Burma, W. Peninsula), Yunnan, tropical
and ‘S$. Africa.
2. Digitaria marginata, Link Hort. Berol.i, 229; Stapf in Prain Fl. Trop.
Afr. ix, 439.
Description: Annual, 30cm. tolm. high. Stems tufted, usually ascend-
ing from a geniculate or prostrate base, simple or branched from the lower
nodes, glabrous, few to many-noded. Leaves 5-15 cm. by 4-8 mm., linear or
linear-lanceolate from a slightly contracted and rounded base, acute, flat,
flaccid, glabrous or sparingly hairy particularly towards the mouth, margins
finely cartilaginous, rough and often crisp, midrib very slender, whitish ;
sheaths thin, subherbaceous, loose, glabrous, or more or less beset with
spreading tubercle-based hairs often forming a loose beard at the base ; ligules
truncate, membranous, up tooverl]mm.long. Spikes mostly 4-9, sessile,
subdigitate, solitary or 2-3—nate on a short, scaberulous common axis, erect or
spreading, rather slender, strict or slightly flexuous 5-15 cm long, often finely
pubescent at the base; rhachis almost straight, triquetrous, lateral angles
winged, herbaceous, scabrid, internodes up to more than 2mm. long. Pedicels
2-nate, onevery short, the other up to 15 mm. long, angular, scabrid.
Spikelets appressed, lanceolate, acutely acuminate, 2-4 mm. long, pale
greenish, rarely tinged with purple, variously hairy, rarely quite glabrous.
Lower involucral glume an ovate, obtuse to subacute membranous scale,
usually not over 0°3 mm. long, sometimes obsolete or quite suppressed ; upper
ovate-lanceolate, acute, equalling or considerably exceeding half of the upper
floral glume, rarely distinctly shorter, 3-nerved, with fine lines of hairs between
the nerves and along the margins, rarely quite glabrous. Lower floral glume
corresponding in outline and size to the spikelet, firmly membranous, 7-nerved,
rarely quite glabrous, usually with fine lines of hairs between the inner side-
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 633
nerves (of each half) and along the margins; upper floral glume oblong-
lanceolate, acutely acuminate, almost as long as the spikelet, thinly
chartaceous, pale or slightly purplish, brownish when ripe. Grain oblong,
plano-convex, whitish, scutellum Jess than half the length of the
rain. ‘
° This species is not identical with Paspalum sanguinale, Lamk. of the F.
B.I. or with Digitaria sanguinalts, Scop., in Cooke’s Flora or in Haines’
Bot. of bihar & Orissa. DD. sanguinalis, Scop. (sensu stricio) isa plant of
S. Europe and has not been found either in India or tropical Africa, as was
pointed out by Pilger (in Engl. Jahrb. xxx, 118) and Stapf. Most of the
numerous synonyms given by Hook. f. in the F. B. I. would have to be men-
tioned underthe different varieties. Here we have to deal only with one
variety which was described by Cooke as var. cz/7aris, Prain and which was
called var. fzmbérzata by Stapf.
var. fimbriata, Stapf, 1. c. 440.—Dzgitaria fimbriata, Link Hort. Berol. 1. c¢.
226.—D. commutata, Schult, Mant. ii, 262.—D. chrysoblephoru, Fig. & De
Not. in Mem. Acc. Tor. ser ii, xiv, 364.—D7gi/arta ciliaris, var. guadristachya,
Wight.--D. sanguinalts, var. ciliaris, Rendle in Cat. Afr. Pl. Welw. ii, 163,
and in Journ. Linn. Soc. Bot. xl, 228.—Digitoria sanguinalis, var. ciliaris,
Prain Beng. Pl. 1181; Cke. ii, 940—Panicum ciliare, Retz. Obs. iv, 16; Kunth
Faia. 1, 62; Roxb. Fl. Ind. i, 290 ; A. Rich. Tent. Fl. Abyss. ii, 360; Dalz. &
Gibs. Fl. Bomb. 290; Duthie Indig. Fodd. Grass. t.9.—P. fimbriatum, Pres}
Rel. Haenk. i, 298; Kunth Enum. i, 81.—FP. sanguinale var. ciliarve, Franch.
Contr. Fl. Congo Franc. 46.—P. sanguinale var. blepharanthum, Hack. in
Durand & Schintz Consp. Fl. Afr. v, 762.—P sarguinale, var. macrostachyum,
Hack. 1. c. 763.—FPaspalum sanguinale var, ciliare, Hook. f. in F.B. I.
wil, 15;
Description: Upper involucral glume usually much exceeding the middle
of the fertile floret and frequently equalling ¢ of its length. Indumentum of
spikelets uniform or more often more or less varied in the same inflorescence ;
hairs of the upper involucra] glume and lower floret partly in fine lines, all of
one kind, very fine, thin-walled, o-tuse-tipped, partly more thick-walled with
slightly clavate tips, and up tol mm. long, spreading out at maturity and
forming a rigid double fringe on each side of the spikelet, the inner fringe
often mixed witha varying number of tubercle-based acute yellow bristles
which ultimately also spread out at right angles.
It is well to remember what Stapf says ina note, ].c. 441: ‘ The peculiar
indumentum of the spikelets, which in the mature state leads to the formation
of spreading fringes, inay extend to all spikelets alike or it may be, at least in
its perfect development, confined to the long-pedicelled member of each pair
of spikelets or only to some of them, in which case the indumentum of the
fringeless spikelets approaches more or less that of var. Linkiz.’? Var. Linki2,
Stapfis Hooker f’s var. commutatum of Paspalum sanguinale in F.B. I.
vat. 15.
Vern. Names: Tara (Surat); Shikaol or Arotaro (Dohad); Chansarieu
(Broach) ; Taro, Modhan (Sind) ; Fakri, Fakria, Kurad, Suka, Revga, Dinohi,
Shikar koli, Kalam hullu (Bijapur) ; Shimpigyan hullu (Belgaum) (ex Burns).
Locality: Sind: Sanghar (Sabnis B903!) ; Mirpurkhas, in cultivated fields
(Sabnis B1209!) ; Bughar, Indus River (Blatter & McCann D686!) ; Ghula-
malla, garden (Blatter & McCann D687 !).—Guzarat: Cutch (Blatter 8742 !) ;
Ahinedabad (Herb. Econ. Bot. Poona!).—Khandesh: N. slope of Chanseli
(McCann 9535!) ; Bor, TaptilIsland, on sand and mud (Blatter & Hallberg
0463!) ; Muravat, Tapti Bank (Blatter & Hallberg 3839!) ; Umalla (Blatter
& Hallberg 5178!); Amalner (Blatter & Hallberg 4443!); Sumit (Blatter
& Hallberg 5158!) ; Bor, Bori River (Blatter & Hallberg 5213!) ; Dadgaum
(McCann 9531!)—Konkan: Victoria Gardens, Bombay (McCann 9831!) ;
Versova, Salsette (McCann 4308!); Malabar Hill (McCann 4300 !); very
common through the Islands of Bombay and Salsette (McCann !) ; Parsik,
railway line (McCann 9530!) ; Bassein (McCann 4485!) ; Alibag, sandy shore
(Ezekiel !).— Deccan: Khandala (McCann 3650!) ; Igatpuri (McCann $833!) ;
Purandhar (McCann 5606!) ; Chattarshinji (Ezekiel!) ; Pashan (Gammie!) ;
Deolali (Blatter & Hallberg 4556!) ; Gangapur (Blatter & Hallberg 4581!) ;
Panchgani, Maratha Well (Blatter & Hallberg B1233!).—S. 7. Country:
Haveri, dry ground, compound of P. W.D. (Talbot 2228!) ; Dharwar (Sedg-
3
634 JOURNAL, BROMBAY._NATURAL AIST. SOCIETY, Vol. XXX
wick!); Belgaum (Herb. Bot. Gard. Cal. !)—Xanara: Karwar (Talbot
1294!) ; Haiyal (Talbot 2153 !) ; Kulgi (Talbot 2279 !).
Distribution: Tropics of both hemispheres, rarely found beyond the tropics.
3. Digitaria pennata, Chiov. in Result. Scient. Miss. Stefanini—Paoli i, 183 ;
Cke. ii, 941, ervonee ascribens nomen Hookerof.; Stapf in Prain Fl. Trop.
Afr. ix, 472.—Panicum pennatum, Hochst. in Flora xxxviii (1855), 197;
Schweinf. in Bull. Herb. Boiss. ii, App. ii, 18.—FPaspalum pennatum, Hook. f.
in PeB.leviil, 1G.
Description: Cke.l.c.
Locality: Sind: Karachi (Stocks feste Hook. f.); Jemadar ka Landa,
near Karachi (Stocks); Tatta, Kullan Kote (Blatter & McCann D684 !) ;
Tatta (Blatter & McCann D685!).—Guwjarat: Porbandar (Bhide !) ; Morvi,
Kathiawar (Woodrow).
Distribution : Baluchistan, tropical Arabia and Africa.
4. Digitaria pedicellaris, Prain Beng. Pl. ,(1903), 1181; Cke. ii, 941; Haines
in Bot Bihar & Orissa 1009.—Paspalam pedicellare, Trin. ex Steud. Nom.
ed. 2 (1841), 272; Hook. f. in F.B.I., vii, 19.--P. pedzcellatum, Nees & Arn. in
Wight Cat. 2310; Duthie Grass. N. W. Ind. 1.—Milium sanguinale, Roxb. FI.
Indsl.. 30> (evel. Si 77, biti.)
Description: Cke.1.c.
Locality: Konkan: ~Thana (McCann 8725!) ; Sion, (Herb. SS. xX. (Calis
Mulyaum (McCann 3663 !).—Deccan : Chattarshinji Hill, Poona (Ezekiel !).—
S. M. Country: Hubli, elevation 2,000 ft.. rainfall 30 inches (Sedgwick &
Bell 4229 !).
Distribution: Throughout India.
5. Digitaria longiflora, Pers. Syn. i (1805), 85 (zon Trin.); Cke. ii, 941;
Stapf in Prain F). Trop. Afr. ix, 469; Haines in Bot. Bihar & Orissa 1008.—
Digitaria Pseudo-Durva, Schlechtend. in Linnea, xxvi, 458.—D. linearis,
Schult. f. Mant. ii, 264 (zom Roem. and Schult.).—D. tenuiflora, Stapfin Dyer
Fl. Cap. vii, 380 (zoz P. Beauv.).—Puaspalum longiflorum, Retz. Obs. iv, 15
(non Trin.) ; Baker Fl. Maurit. 431; Hook.f. in F.B.I. vii, 17 (partim).—P.
brevifolium, Fluegge Gram. Monogr. 150.—P. Pseudo-Durva, Nees Fl. Afr.
Austr. 21.—P. filiculme. Nees ex Thw. Enum. Pl. Zeyl. 358.—Panicum longi-
florum, Gmel. Syst. 158.—P. parvulum, Trin. Pan. Gen. 117.—P. argyrot-
richum, Durand & Schintz Consp. Fl. Afr. vy, 741 (mon Anders.).—-Milium
filiforme, Roxb, FI. Ind. i, 314.
Digitaria tenuiflora, P. Beauv. given as asynonym by Cke. seems to be a
different species. It is apparently a perennial of erect habit and with long
narrow leaves.
Description :. Cke. lic;
Locality: Deccan: Deolali (Blatter & Hallberg 9835 !).—S. 47. Country :
Dry uplands, Dharwar, elevation 2,400 ft., rainfall 34 inches (Sedgwick 2653 !) ;
Belgaum (Herb. Bot. Gard. Cal.).—Kanara: Halyal (Talbot 2310!) ; Londa
(Bhide!).
Distribution: Throughout India, Ceylon, tropical and S. Africa, Mada-
gascar, Mascarenes, Malaya.
6. Digitaria Royleana, Prain Beng. Pl. (1903), 1181; Cke. ii, 942; Haines in
Bot. Bihar & Orissa 1008.—Faspalum Royleanum, Nees ex Thw. Enum.
(1864), 358; Hook. fin BT. vai; 18% rin). Ceyl. v.25.
Description: Cke.1.c.
Locality: Konkan: St. Xavier’s: College compound (McCann 4533!) ;
Mulgaum, Salsette (McCann 9523 !).—Deccan: Khandala (McCann 3651!) ;
Lonavla (Herb. Econ. Bot.. Poona!) ; Purandhar Fort (Bhide!) ; Lina Hill,
Nasik District (Blatter & Hallberg 4542!); Lohagad, way up (McCann 9512 !);
Katraj Ghat, 11 miles S.E. of Poona (Bhide!); Panchgani, slopes below
Third Tableland (Blatter & Hallberg B1229!); Panchgani, Maratha Well
(Blatter & Hallberg B1224 !, B1281!) ; Mahableshwar, in a garden, elevation.
4,500 ft., rainfall 270 inches (Sedgwick & Bell 4584 !); Suvasni Ghat
(Woodrow).--S. JZ. Country : Dharwar, elevation 1,600 ft., rainfall 34 inches
(Sedgwick 2843!) ; Belgaum (Herb. Bot. Gard. Cal.!).—Kanara: (Talbot!)
Distribution: Hilly districts throughout India, Ceylon ; apparently not in
tropical Africa. . . ie
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 635
43. MALLOTEROPSIS, Presl, emend. Hitchc, in Contrib. U. S. Nat.
Herb. xii, 210; Pigpe in Prain Ef, Trop. Attsix, 462, ee eee
Chew il Jeo).
Stapf (1. c. 483) explains why he adopts Alloteropsis, Presl, as emended by
Hitchcock. ‘ As Hitchcock (Contrib. U.S. Nat. Herb. xii, 210) has pointed
out, Presl’s description and analyses of Alloteropsis are based on a composition
of a Panicoid and an Andropogonoid grass, whilst the original in Presl’s
herbarium is undoubtedly the plant described here as A. semzalata, and so is
also the habit figure (1) in Presl’s plate. The genus is therefore accepted here
with Hitchcock’s emendation.’
‘Another member of this genus, A. cimicina, was included by Palisot de
Beauvois (Agrost. 12) in his genus Avonopus under its earliest synonym
Milium cimicinum and as ‘‘ A Ccimicinus?’”’ on p. 154, and this led J. D.
Hooker (Fl. Brit. Ind. vii, 64) to use the name Axonopus in preference to
Alloteropsis, a view which was adopted by myself in F.. Cap. vii, 418. From
P. Beauvois’ diagnosis however, and from the fact that he quotes in the first
place Milium compressum as example for Axonopus, there can be no doubt
that he had primarily (/2lzum compressum in view when establishing his genus
Axonopus, and it is in that sense that the genus is understood in this work.
A. cimicina also forms the basis of another genus, Corzdochloa, Nees in Edinb.
News Phil Journ, xv, 381. A. Chase (in Proc. Biol. Soc. Washington, xxiv,
157) maintains this genus as distinct from <A//oteropsis, and I followed her
when drawing up the key of the genera of Tropical African grasses (p. 13) ;
but I have since come to the conclusion that the species referable to these
two group. are so similar in the peculiar structure of their spikelets that they
are better merged into one genus for which Al/oteropsis has priority over
Cordtochtoa.’
We have, therefore, to add the characteristics of the genus asgiven by
Stapf:
Perennial or annual. Jleaf-blades flat or more or less convolute; ligules
membranous, ciliate or ciliolate, short or reduced to amere rim. Racemes
sessile or pedune led, often more or less compound towards the base, digitate
or subdigitate on a more or less elongated common axis. Spikelets ovate or
elliptic to lanceolate-oblong ; acute or acuminate, mostly awned, slightly or
conspicuously compressed from the back, falling entire from the pedicels,
2-nate or fascicled, subsecund and abaxial on the triquetrous rhachis of more
or less spiciform racemes. Lower floret usually male, upper hermaphrodite.
Involucral glumes unequal, lower smaller, membranous to hyaline, 3-1-nerved,
very. acute, often inucronulate, upper equal or subequal to the spikelet,
membranous to chartaceous, 5-nerved, with the outer nerves submarginal,
densely ciliate along them. Lower floral glume resembling the upper involucral
glume, but ciliate, pale short, deeply 2-fid with conspicuously auricled flaps,
upper floral glume chartaceous, glabrous, delicately ciliolate upwards,
o-nerved, produced into a straight awn or mucronate, pale equal to the glume,
2-keeled, with broadly auricled flaps. Lodicules 2, broadly cuneate. Stamens
3. Styles distinct, stigmas laterally exserted. Grain enclosed by the glume
and pale, elliptic-oblong, dorsally much compressed ; scutellum about half the
length of the grain ; hilum basal, punctiform.
Species about 5.—In the tropics anc the warm temperate zone of the Old
World, 2 in India.
In the Bombay Presidency there is only one species :
1, Alloteropsis cimicina, Stapf in Prain Fl. Trop. Afr. ix, 487; Haines in Bot.
Bihar & Orissa 1009.—MWilium cimicinum, Linn. Mant. Alt. 184.— Panicum
cimicinum, Retz. Obs. ili, 9; Roxb. Fl. Ind. i, 295; Steud. Syn. PJ. Glum..i,
43.—P conjugatum, Dalz. & Gibs. Bomb. Fl. 291.—Axonopus cimicinus (?),
ibeanv. Acrost. 154; Hook. f. in F. B. 1. vii, 64; Hook. f.in Trim..Fl. Ceyl.
v, 166; Cke. ii, 925.—Uvochloa cimicina, Kunth Rev. Gram. i, 31, t. 103;
Dalz. & Gibs. 1. c 289.—Cordiochloa cimicina, Nees in Edinb. N. Phil.
Journ. xv (1833),.381; A. Chase in Proc. Biol. Soc. Wash. xxiv, .158.—C.
fimbriata, Nees ex Wight Cat. no. 16562 Aitehis. Cat. Panvab,, Pi, 158.
Description : Cke, lsc.
Locality : Konkan: Sewri (McCann 3586!) ; Mulgaum (McCann 3654 !).—
Deccan: Sinhagad forests (Rhide!); Lina Hill, Nasik District (Blatter &
636 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXX11
Hallberg 4583!) ; Panchgani (Hallberg !).—S. 7. Country: Dharwar (Sedg-
wick 2032!, Woodrow) ; Gokak (Shevade!); Badami (Woodrow).—-Kanara :
Halyal (Talbot 2294!) ; Kulgi (Talbot 2434 !).
Distribution : Tropical Africa, Madagascar, throughout India to Java.
44. ERIOCHLOA, H. B. & K.; Cke. ii, 944.
Species about 25.—In the warm parts of the whole world.
Cooke describes one species: &. polystachya, H. B. & K. which name has
to cede to &. ramosa, O. Kuntze.
1. Eriochloa ramosa, O. Kuntze Rev, Gen. Pl. ii, 775; Hack. in Bull. Acad.
Int. Bot. xvi; 19; Merrill in Philipp. Journ. Sc. i, Suppl: 348sestapiin Pra
Fl. Trop. Afr.ix, 498; Haines in Bot. Bihar & Orissa 1006.—Z. annulata,
Kunth Rev. Gram. i, 30, and Enum. i, 73; Duthie List Grass. N. W. Ind. 2;
Benth FI. Hongk. 409 ; Hack. in Engl. Jahrb. vi, 233.—£. polystachya, Duthie,
fll. Indig. Fodd. Grass. Ind. t. 41; Fodd. Grass. N. Ind. 2; Hook. f: in F-Ba:
vii, 20; Cke. ii, 944; ‘Rendle in Journ: Linn. Soc, Bot. xxxvi, 320 (vow H.R:
& K.!).—Milium ramosum, Retz. Obs. vi (1791), 22; Roxb. Fl. Ind. i, 317;
Griff. Notul. iii, 15; Ic. Pl. Asiat. t. 139, f. 60.—Agvostis ramosa, Poir. Encycl.
Suppl. i, 257.—Paspalum annulatum, Fluegge Monogr. Pasp. 133; Trin Sp.
Gram. Ic. t. 133.—Helopus levis, Trin. ex Spreng. Neue Entdeck. ii, 49, fig.
4.—H. annulatus, Steud. Syn. Glum. i, 99 (zou Nees) .-—~Pipatherum annulatum,
Presl, Rel. Haenk i, 221 (xom Raddi).
Note.—Masaja Honda in his Revisio Graminum Japonie (Bot. Mag. Tokyo
37 (1923), 113-124) is of opinion that Avzochloa ramosa, O. Kuntze, has to be ©
partly emended and changed to the new species Eriochloa Hackelit. Details
are wanting to form an opinion on this point.
Description : Cke. 1. c.
Localily: Sind: Umerkot, in a garden (Sabnis B718!) ; Jamesabad, on
banks of a watercourse (Sabnis B967!); Bughar, Indus River (Blatter &
McCann D691!); Tatta, Kullan Kote Lake (Blatter & McCann D692!) ; Tatta
(Blatter & McCann D693!).—Gujarat: Ahmedabad (Gammie 16408!).—
Konkan: Bassein (Bhide!) ; Antop Hill (McCann 3613!) ; Alibag, rice field
(Ezekiel, !) ; Bandra, on walls and in ditches (McCann !) ; Bombay (Hallberg
Al41!); Bombay, near Mahim (Woodrow, Lisboa); Bhandup, in an old
distillery compound, ina ditch (Hallberg A19!).—S. M/. Country: Shiggaon,
elevation 2,000 ft., rainfall 34 inches (Sedgwick 2356!) ; Kunnur, elevation
2,000 ft., rainfall 35 inches (Sedgwick & Rell 4937!) ; Ranibenur (Bhide!) ;
Dharwar (Sedgwick !).—Kanara: Halyal (Talbot !).
Distribution : Tropics of the Old World, introduced into Ascension Island,
St. Helena, Cuba.
45. BRACHIARIA Griseb. in Ledeb. Fl. Ross. iv, 469; Stapf in Prain FI.
Trop. Air. 1x,-000;
(Sect. Brachiaria and part of sect. Paspaloidee of Panicum of the F.B.1.).
Perennial or annual. Leaf-blades linear to lanceolate, usually flat ;
ligules reduced to a narrow ciliate or ciliolate rim. Racemes usually subsessile
and sclitary on a common axis, sometimes bare at the base owing to the
arrest of spikelets, rarely truly peduncled and panicled, simple or compound
near the base, rarely to or beyoud the middle; rhachis filiform, triquetrous
or more or less flattened and herbaceous with a wavy or zig-zag midrib,
which projects as a mostly acute keel on the face; pedicels solitary or in
pairs, alternately to the right and the left of the facial angle or the midrib, if
solitary all short or very short, if paired, the primary slightly to very much
longer ; spikelets closely appressed, always biseriate in the plane, but frequent-
ly becoming 1-seriate by the dovetailing of the alternate spikelets of the closely
approximate ranks, more or less contiguous with their sides or imbricate, form-
ing dense, spike-like racemes, or distant by almost their own length or more,
glabrous or hairy. Spikelets more or less elliptic or oblong, more or less flat-
tened or slightly depressed, convex on the base, falling entire from the pedicels,
1-2-, rarely more-nate, secund and adaxial (with lower involucral glume towards
the axis and the convex side of the upper floral glume away from the axis),
closely appressed to and 2-seriate on the triquetrous or flat rhachis of spiciform
racemes ; lower floret male or barren witha usually well-developed pale, very
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 637
rarely the latter suppressed. Involucral glumes dissiniilar and mostly very
unequal in length. Lower involucral glume shortest ; upper resembling and
more or less equalling the lower floral glume, 5-7- (rarely 9-) nerved. Lower
floral glume 5-, rarely 7-nerved, the lateral nerves placed towards the margins
and distant from the middle nerve; pale usually only slightly shorter than the
valve with well-developed inflexed flaps, or the latter vanishing above the middle;
upper floral glume oblong to elliptic in outline, emucronate, though sometimes
contracted into an apiculus, crustaceous or subcoriaceous with firm involute
margins, faintly 5-nerved ; pale almost as long as the glume, 2-keeled, its
sides tightly embraced by the valve. Lodicules 2, small, broadly cuneate.
Stamens 3. Styles distinct ; stigmas plumose, laterally exserted from the upper
part of the spikelet. Grain tightly enclosed by the glume and pale, more or
less flattened on both faces ; hilum subbasal, punctiform ; embryo half to over
? the length of the grain.
Species about 80. In the warm parts of the whole world, but chiefly in
Africa.
Cooke mentions 3 species which belong to this genus: Panicum TIsachne,
P. ramosum and P, muticum. 'To these we add Brachiaria distachya (Panicum
distachyum) .
A. Spikelets 0°5 mm. long orslightly more ... 1. &B. lsachne.
B. Spikelets 2°5-4 mm. long
E90 chito 1:8 pa hich ves aot Es ULI.
II. Less than 80 cm. high
1. Spikes S-many ... aise ee Os Oe POOSC,
2. Spikes 2-4 ste aes .. 4. B. distachya.
1. Brachiaria Isachne, Stapf in Prain Fl. Trop. Afr. ix, 552; Haines in Bot.
Bihar & Orissa 1004 (habet Roth per errorem).—B. cruciformis, Griseb. in
Ledeb. Fl. Ross. iv, 469.— Panicum /sachne, Roth ex Roem. & Schult. Syst.
ii, 458; Roth Nov. Pl. Sp. 54; Schult. Mant. ii, 252; Steud. Syn. Pl. G1. i, 57;
Hook. f, in F.B.I. vii, 28; Stapf in Dycr FI. Cap. vii, 390; Cke. ii, 931.—
P. cruciforme, Sibth. & Sm. Fl. Greca i, t. 59; Baker Fl. Maurit. 434;
Schweinf. in Bull. Herb. Boiss. ii, App. ii, 19; Duthie Grass. N.W. Ind. 3,
Indig. Fodd. Grass. t. 43, Fodd. Grass. N. Ind. 6; Boiss. Fl. Or. v, 437.—
P. caucasicum, Trin. Sp. Gram. Ic. t. 262.—P. Wightiz, Nees Fl. Afr. Austr. 29.—
P. pubinode, Hochst. ex. A. Rich. Tent. Fl. Abyss. ii, 363.—chinochloa cruci-
forms, Koch in Linnea, xxi, 437; Reichb. Ic. Fl. Germ. i, t. 29, fig. 1413.
Description : Cke. ii, 931.
Locality : Sind: (Herb. Econ. Bot. Poona!).—Gujarat: Surat (Chibber !,
Dalzell teste Cooke).—Khandesh: Sungiri (Gammie 16552!) ; Dhulia Farm
(Chibber !) ; Chanseli (McCann A92!) ; Nimb, Tapti Bank (Blatter & Hall-
berg 9571!) ; Dadgaum (McCann 9562!) ; Tapti, Bhusawal (Blatter & Hall-
berg 5156!) ; Umalla village (Blatter & Hallberg 5159!) ; Bor, Bori River
(Blatter & Hallberg 4424!).—Konkan: Clerk Rd., Bombay, along brackish
water (Sabnis 9565 !); very common in Bombay and Salsette Islands (McCann !).
—Deccan: Yeola (Herb. Econ. Bot. Poona!) ; Mangiri, 8 miles E. of Poona
(Gammie!) ; Sholapur (D’Almeida A91!); Deolali (Blatter 9570!, 9569!) ;
Khandala, common (McCann 9566!) ; Purandhar, N. foot (McCann 9568 !) ;
Panchgani, below Sidney Point (Blatter & Hallberg 1271!).—S. M. Country :
Nelogi, elevation 1,800 ft., rainfall 30 inches (Sedgwick 2134!) ; Haveri (Talbot
2150 !).—Kanara : Halyal (Talbot 2150 !).
Distribution : Throughout the plains of India in damp places, Ceylon, west-
wards to Italy and tropical and S. Africa.
*2. Brachiaria mutica, Stapf in Prain Fl. Trop. Afr. ix, 526.—Panicum
muticum, Forsk. Fl. Aegypt.—Arab. 20; Link Hort. Berol. I, 206 ; Hook. f. in
F, B. I. vii, 34, and in Trim: Fl. Ceyl. V, 140.— P. numidianum, Lam. I11.I,
172; Boiss. Fl. Or. v, 438.— P. purpurascens, Raddi ex Opiz in Flora (1822),
266.—P. barbinode, Trin. Sp. Gram. Ic. t. 318; Duthie List Grass. N. W. Ind.
2.— P. sarmentosum, Benth. in Hook. Niger Fl. 561 (70m Roxb.).— P. equinus,
Steud. Syn. Pl. Glum. i, 73.— P. molle, Griseb. Fl. Brit. West Ind. 547 (excl.
syn.); Baker Fl. Maurit. 436.
In adopting Forskal’s name ‘ muticuim’ for this species Stapf, according to
his own words, -has relied on Ascherson’s indentification (Asch. & Schweinf,
Ill, Fl. Egypt 160) of the type with the Algerian P. numidianum.
638 JOURNAL, BOMBAY NATURAL AIST. SOCIEIY, Vole axa
Popular Names : Water Grass, Mauritius Grass, Para Grass, Scotch Grass,
Buffalo Grass.
Descripiion ;: Perennial 1-2°5 m. high. Stems ascending from a sometimes
prostrate and copiously rooting base, stout, terete, usually many-noded and
sheathed high up, simple or sparingly branched, glabrous, often waxy, pruinose
below the nodes. Leaf-blades linear, up to 30cm. long, 6-10 mm. broad
glabrous or rarely more or less hirsute, margins scabrid. Panicle oblong to
ovate-oblong in outline subsecund or almost quaquaversal, 6-20 cm. long;
common rhachis terete to semiterete, more or less deeply channelled or triquet-
rous upwards, scabrid along the angles, glabrous. Racemes numerous,
solitary or irregularly approximate, sometimes paired or in false whorls, shortly
peduncied or subsessile, obliquely spreading, 7 (rarely 12) to 2°5 cim. long,
mostly compound, glabrous, greenish or tinged with purple ; rhachis flat, with a
slender, raised midribuptoimm. wide, villosulous atthe base, otherwise
vlabrous; secondary racemes usually very short, and 6-3-spiculate ; pedicels
solitary or paired, very short, or if paired then the longer up tol mm. long,
frequently with a few setules. Spikelets laterally contiguous or discontiguous,
those of the secondary racemes often imbricate, oblong or lanceolate-oblong,
acute, 3-3°5 mm. long, glabrous. Involucral glumes dissimilar, lower broad-
ovate, acute to subacute, from lessthan 3 to not quite$ the length of the
spikelet, faintly 3-5-nerved, often tinged with purple; upper corresponding in
outline and size to the spikelet, 5-7-nerved. Lower floral glume as long as the
upper involucral glume and similar to it; pale narrowly oblong, subacute,
almost as long as the glume, with narrow flaps; anthers 2mm. long; upper
floral glume slightly shorter than the spikelet, mostly 3mm. long, oblong,
subacute or minutely apiculate, pale yellowish, glume and pale crustaceous,
very finely transversely wrinkled or almost smooth. Stigmas blackish-purple,
very conspicuous.
Locality : Cultivated at Kirkee and Surat (Woodrow) and very likely in other
places.
Distribution: A native of S. America and W. Africa, but introduced else-
where (Stapf).
Uses: A fodder grass. See Kew Bull. (1894), 384.
3. Brachiaria ramosa, Stapf in Prain Fl. Trop. Afr. ix, 542; Haines Bot.
Bihar & Orissa 1005.—Panicum ramosum, Linn. Mant. (1767), 29; Steud.
syn. Pl. Glum.1, 97 ; Hook. tin. FB. yil,.36)( part) 3, Prim. el Ceyie aves
140; Prain Beng. Pl. 1175; Cke. ii, 932.—P. arvense,-Kunth Rev. Gram.
i, 391, t. 109.—/.. Petiveri, Dis. ii, 144. (parvtim); Baker Fl. Maurit. 434 ;
Aitchis. Cat. Panjab Pl. 160; Duthie Grass: N. W. Ind. 6;-Fodd. tGrase
N. Ind. 11; Boiss. Fl. Or. v, 439.—P. brachylachnum, Steud. lc. 62.—P.
cognatissimum, Steud. 1.c.69.—P. patens, Boj. Hort. Maurit. 365 (zom Linn.).—
P. pygmaéum, Boj. 1.c.—P. Helopus, Watt. Dict. Econ. Prod. vi, part 1, 10
(partim).—P. umbrosum, Retz. Obs. 4 (1786), 16; Roxb. FI. Ind. i, 297.
Description : Cke. ii, 932.—Stapf points out that this species occurs in a glab-
rous and apubescent state, and that the original specimen in Linneeus’ herbarium
represents the former. . ‘ The pubescence,’ he says, ‘ if present, extends gener-
ally to the culms, the leaves, the axes of the inflorescence and the spikelets, the
upper glume [upper involucral glume] and lower valve [lower floral glume].
On the blades it may be scanty and disappear with age. It does not seem to be
correlated with any other character, and the area of the glabrous and pubescent
states overlap completely, in fact both have been taken in the same collecting.’
He mentions another curious modification in which the lower floral glume is
more firmly membranous to crustaceous and faintly transversely rugose and
thus more or less resembles the upper floral glume (not the upper involucral
glume as is normally the case). It has been collected in India and W. Africa.
Locality : Sind: Chachtra (Mamlatdar of Chachra!) ; Shahabander (Karachi
P.O.C. of Shahabander!); Sangarh (Sabnis BY01!, B887!); Nasarpur,
clayey soil (Sabnis B1057!).—Gujarat: Ahmedabad (Herb. S.X.C. 2165!);
Mausari (Mamlatdar of Mausari!); Sumrasar, Cutch (Blatter 3756!).—
Khandesh:: Tamer, Tapti bank (Blatter & Hallberg 5172!) ; Antroli, Bori
River (Blatter & Hallberg 5149!); Toranmal (McCann A142!).—Konkan :
Malabar Hill (McCann !) ; Versova (McCann 9588!) ; Byculla (McCann 9586!) ;
Sion (McCann 8689!) ; Bandra Hill, in fallow fields (Vakil A115!).— Deccan :
Khandala (Sedgwick 2631!) ; Poona (Woodrow!): Lina Hill, Nasik District
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY _ 639
(Blatter & Hallberg A145 ').—S. AZ. Country: Dharwar, garden weed
(Sedgwick 2651!) ; Haveri (Talbot 2231!) ; Badami (Woodrow Zeste Cooke).
Distribution : Throughout India, Ceylon, Afghanistan, tropical Africa (Upper
Guinea, Cape Verd Islands).
4. Brachiaria distachya, Haines in Bot. Bihar & Orissa 1004 (per errorem
distachyum).—-Panicum distachyum, Linn. Mant. i, 138; Retz. Obs. iii, 17;
Damk. Ill. t. 43, f.2; Steud. Syn. Gram. 41; Aitchis, Cat. Panjab, Pl: 159;
Duthie Grass N. W. Ind. 3; Indig. Fodd. Grass. t. 42, Fodd. Grass. N. Ind. 6;
Benth. Fl. Austral. vii, 478; Hook. f. in F.B.I. vii, 37.— P. subguadriparum ,
Trin. Gram. Panic. 145, Sp. Gram. Ic. t. 186.—Dzigzitaria distachya, Pers. Syn.
1,:35.
Description: A slender, creeping grass, glabrous or panicle sparsely hairy.
Stems 30-60 cm. high. Leaves linear or lanceolate or linear-lanceolate, acu-
minate, 5-15 cm. by 3-6 mm., widest at the rounded or amplexicaul base, flat ;
sheaths ciliate or noton the margins, mouth hairy. Spikes 2-4, distant, 2 5-6°5
cm. long, rarely morethan 10 cm., erect, at last spreading ; rhachis slender,
glabrous. Spikelets variable in size, pale green, 3-4 mm. Jong, solitary, subses-
sile, spikately arranged in 2 (-1) series, ellipsoid, glabrous. Lower involucral
glume embracing the spikelet and margins overlapping below, § to nearly 4 the
spikelet, 5-7-nerved, obtuse or subacute ; upper involucral glume ovate, acute,
7-nerved, paleate or not, pale if present narrow, neuter. Lower floral glume
5-nerved, upper ellipsoid, obtuse or rounded, 2mm. long, brown and minutely
transversely lineolate or obscurely rugulose when ripe.
Locality : Gujarat : Ahmedabad, banksand margins of fields around Ahmeda-
bad (Sedgwick 239!).—S.M/. Country : Dharwar (Sedgwick 2840 !).
Distribution : India, Ceylon, China, Malaya, Australia.
46. PaspaLum, Linn. Syst. Nat. ed. x, 855; Cke. ii, 943.
Species over 200, chiefly in tropical America, only a few in the Old World.
Cooke describes 3 species: P. scrobiculatum, Linn., P. compactum, Roth.
and P. distichum, Linn., to which we add Paspalum dilatatum, Poir. P.
distichum has to be replaced by P. vaginatum, Sw., for reasons detailed below.
I. Leaves over 14 cm. long
1. Plant 60-90 cm. high oe .. Ll. P. serobiculatum.
2. Plant 1-1 San. high ane 2 2 L Atlatalium.
II. Leaves less than 11 cm. long
1. Leaves 2°5-7'5 cm long, 8-12 mm.
broad ee ad .. 3. FP. compactum.
2. Leaves 5-10 cm. long, 1:2-4 mm.
broad : ee 4. P. vaginatum.
1, Paspalum scrobiculatum, Linn. Mantiss. (1767), 29; Hook. f.in F. B. J. vii
10; Cke. ii, 943; Haines in Bot. Bihar & Orissa 1000.—P/. scrobiculatum,
Lion. var. Commersoniz, Stapf in Prain Fl. Trop. Afr. ix, 573.—P. scrobicu-
latum, Linn. var. frumentaceum, Stapfl.c. 575.—P. scrobiculatum, Linn. var.
polystachyum, Stapf 1. c. 576.—For further synonyms see Hook. f. and Stapf
ll. cc. Hook f. gives P. orbiculare, Forst. Prodr. 7 as a synonym, but Stapf
considers it as a distinct species of the Polynesian and Indo-Malayan regions
with an extension into New Zealand and Australia. Of Hook. f. syn. in the
F. B. I. we wish to enclose only those which apply to spontaneous forms of the
Old World.
As can be seen from tle above synonymy we are not following Stapf in
distinguishing several varieties or rather forms. He takes the cultivated forms
of India to be the original P. scrobiculatum of Linnueeus and calls it P. scrobicu-
latum var. frumeniaceum. All the spontaneous forms of P. scrobiculatum as
understood by most post-Linnzean authors are put by Stapf under P. scrobtcu-
latum, Linn. var. Commersonii, the type for this combination being P.
Commersonit, Lam. 111. i, 175, t. 43, fig. 1. The third form var. polvstachyum
does not seem to occur in India.
Description: Cke. 1. c.
Locality: Sind: Jamesabad, in bed of watercourse (Sabnis B979!) ; Bohara
(Blatter & McCann D689!).—Kowukan : Victoria Gardens, Bombay (McCann
4297!) ; Mulgaum, Salsette (McCann 3607 !) ; Parsik, between stones of railway
tract (McCann 9516!); Vihar Lake, Salsette (McCann 9517!).—Deccan :
(40 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXI1
Khandala, in watercourse, on sandy soil (McCann 9824!).—S. WZ. Country:
Konankeri, Dharwar District, elevation 1,800 ft., rainfall 40 inches (Sedgwick
& Bell 4966!) ; W. of Dharwar, elevation 2,000 ft. rainfall 40 inches (Sedg-
wick & Bell 4452!) ; Dastikop, elevation 2,500 ft., rainfall 35 inches (Sedgwick
2109!) ; Dharwar (Garade!) ; Belgaum (Herb. Bot. Gard. Cal.!).—Kanare :
Halyal (Talbot 2297!) ; Karwar (Herb. Econ. Bot. Poona 2297 !, 618!) ; Castle
Rock (Gammie!).
Distribution : Tropics of the Old World.
*2. Paspalum dilatatum,* Poir. Encycl. v, 35; New South Wales Agric:
Gaz. x (1899), 32, with plate; Trin. Diss. 11, 113, with plates; Agric. Grass.
U.S. 31-32; H. H. Mann in Dept. Agric. Bomb. Bull. 77, 68.—P. ovatum,
Nees ex ‘Trin. Gram. Panic. 113.-- P. pratense, Spreng Syst. i, 247.—P. Selloz,
Spreng. ex Nees Agrost. Bras. 43.
Popular Names: Golden Crown Grass, Hairy-flowered Paspalum.
Description: A tall, erect grass, about 1-1'5 m. high. Culm developing
from a thick rootstock with 3-5 leaves; leaf at base of culm often about 30 cm.
long, 8-12 mm. broad, smooth on both sides, rugose along the margins.
Raceme 12-3) cm. long, having 5-10 somewhat spreading spikes, which are
7 cm. or more in length, 2°5-5 cm. apart, upper ones gradually shorter.
Spikelets closely arranged in 4 rows, two on each side of the narrow and nearly
straight axis in alternate pairs, 15-2 mm. wide, and 3-4 mm. long, ovate,
acutely pointed, crowded and overlapping each other, compressed, margins
clothed in silky hairs. Involucral glumes ovate, acute, 5-nerved, nearly smooth
except the fringe of white hairs on the margin. Floral glumes thick, hard, and
firm, very minutely punctate. Pale fitting inside the flowering glume and
enclosing the stigmas and styles. Anthers linear. Styles 2; stigmas blackish-
purple, plumose.
Locality : Cultivated. See Mann, 1. c.
Distribution : Virginia, Mississippi, Louisiana, ‘lexas, S. America, especially
Brazil.
Uses: One of the best fodder grasses in N. America and is extensively
cultivated there. For a ful! account see Kew Bull. (1602), 1-4.
3. Paspalum compactum, Roth Nov. Pl. Sp. (1821), 36; Kunth Enum. PI. i, 61 ;
Steud. Syn. Gram. 31; Hook. f. in F. B. I. vii, 12; Cke. ii, $43.—P.. militaria,
C. Muell. in Bot. Zeit. xix (1861), 325.—P. Canare, Steud. 1. c. 58.—P. tmper-
fectum, Roxb. ex Kunth l.c.
Description : Cke. 1.c.
Locality : Konkan : Tiwari-Pada, Bassein (Herb. Econ. Bot. Poona 1€90!) ;
Matheran, to Louisa Point (D’Almeida A243!).—Deccan : Khandala, growing
in gravelly soil, very common (McCann 9823!); Igatpuri, very common
(McCann 4587 !); Panchgani, First Tableland (McCann B1300! B1265!), Second
Tableland (McCann B1242!, B1292!); Mahableshwar (Cooke, Woodrow).—
S. M. Country: Londa (Bhide!); Belgaum (Hole 15!).—Kanara: Yellapore
(Talbot 657!) ; Tinai (Talbot 2566!) ; Castle Rock, elevation 1,900 ft., rainfall
250 inches (Sedgwick 2752!) ; Karwar, Bingy Ghat (a very villious form,
McCann) (Talbot 1529!) ; Devimani, elevation 1,300 ft. (McCann A17!).
Distribution : W. Peninsula of India.
4. Paspalum vaginatum, Sw. Prodr. Veg. Ind. Occ. 21, Fl. Ind. Occ. i, 135;
Trin, Gram. Pan. 94; Panic. Gen. $3, Sp. Grams Ic. €/1120); Kunth enunier
52; Steud. Syn. Pl. Glum. i, 20.: Hitchcock & Chase in Contrib. U:.S. Nat
Herb. xviii, 307 (zon P. Beauv. FJ. Owar.) ; Stapf in Prain Fl. Trop. Afr. ix,
570.—P. vaginatum, forma longipes, Lange in Vidensk. Medd. Naturh. Foren.
Kjébenh. (1854), 42, Pug. Pl. Hisp. i, 28.—P. dzstichum, N. L. Burm, FI, Ind.
23; Gaertn. Fruct. 11,2, t. 80°; Kunth 1. c.j) Steud. lic. 29; Baker th Maurie
431; Benth. Fl. Austr. vii, 460; Hack. in Forschungsr. S. M.S. ‘ Gazelle’ iv,
6 ; Hook. f. in F. B. Lb vii, 12;" Rendle. in Journ. Linn. Soc:.Bot. xxxviny 319k
Cke. ii, 943 ; Ridley Mat. Fl. Mal. Pen. iii, 124 ; Stapf in Dyer F1. Cap. vii, 371;
Merrill in Philipp. Journ. Sc. i, Suppl. i, 346 (zou Linn.).—P. dztstichum, var.
vaginatum, Griseb. Fl. Brit- W. Ind. 541.—P. Jittorale, R. Br. Prodr. 188; Trin.
1 We have to thank Mr. K. Biswas, the Curator of the Herbarium of the
Sibpur Botanic Garden, for most of the information contained under this
species.
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 641
Gram. Pan. 95, Sp. Gram. Ic. t. 112.—P. longitlorum, P. Beauv. Fl. Owar. ii,
46, t. 85, fig. 2; Grah. Cat. Bomb. 234 (zon Retz.).—P. bvachiatum, Trin. ex
Nees Agrost. Bras. 62.—P. foliosum, Kunth Rev. Gram. i, 25.—P. sgvamatum,
Steud. 1. c.. 21-—FP. inflatum. A. Rich.in Ram. de Sagra Fl. Cub. iii, 298.—
P. kleintanum, Pres] Rel. Haenk. i, 209.—P. Boryanum, Presl i. c—Ligitaria
foltosa, Lag. Gen. ef Sp. Nov. 4.—Y. vaginata, Philippe FI]. d. Pyren. ii, 415.—
D. paspaloides var. longipes, Lange ex Willk & Lange Prodr. Fi. Hisp. i,
45.—Sanguinaria vaginata, Bub. Fl. Pyren. iv, 258.
To explain the change of name from P. distichum, Linn. to P. vaginaium,
Sw., and to elucidate certain points of the above synonyiny taken from Stapf, it
will suffice to quote the short explanation given in the Fl. Trop. Afr. p. 572:
‘Frequently confused with Paspalum distichum, Linn. (Herb. Linn. !) which
has broader and flatter leaves, mostly sessile or subsessile lateral spikes and
spikelets with firmer sides and a more convex upper glume, which is appressedly
and silky pubescent.’
Description: Cke. ii, 943.
Locality: Gujarat: Marshy edge of the Bokh, Prantij (Herb. Econ. Bot.
Poona!).—AKonkan: Marine Lines, Bombay (Hallberg 9514!) ; sea-shore,
Bombay (Woodrow) ; Malabar Hill, Bombay (McCann 3609!) ; St. Xavier’s
College compound (McCann 9830!) ; Alibag, sandy shore (Ezekiel!) ; Malwan
(Woodrow).
Distribution. ‘Tropics of the whole world, mostly on the sea-shore ; intro-
duced into Galicia (Spain) and the Western and Centra! Pyrenees.
47, PASPALIDIUM, Stapf in Prain Fl. Trop. Afr. ix (1917), 15 clavi et 1. c.
(1920), 582.
(Sect. Paspaloidee in F. B. 1. partim).
Perenniai, semiaquatic or terrestrial. Leaves linear, flat or involute. Ligule
a ciliolate rim. Racemes sessile or sub-Sessile and secund on the alternate
notches of a triquetrous common axis of a false compound spike, more or less
appressed to the more or less hollowed-out flanges of the latter. Rhachis ending
in a subulate point. Spikelets mostly conspicuously 2-seriate, nearly always
quite glabrous, ovate to ovate-oblong or ovate-lanceolate (when seen in front
view), awnless, falling entire from the pedicels, solitary, secund and abaxial on
the rhachis. Involucral glumes mostly dissimilar and very unequal in length ;
lower reduced toa small scale or upto, rarely over, half the length ofthe
spikelet, upper mostly almost equalling the spikelet, 5-7-nerved with the
nerves evenly distributed, rarely both glumes much reduced. Lower floral glume
similar to the upper involucral glume with the inner side-nerves more distant,
pale if present only slizitly shorter than its glume with well developed inflexed
flaps; upper floral glume oblong to elliptic in outline, acute to apiculate ;
emucronate, crustaceous with firm involute margins, faintly 5-nerved, pale
almost as long asits glume, 2-keeled, its sides tightly embraced by the glume
allalong. lLodicules 2, small, broadly cuneate. Stamens 3. Styles distinct ;
stigmas p!umose, laterally exserted from the upper part of the spikelet. Grain —
tightly enclosed by the more or less hardened glume and paie.
Species about 12.—In the warm countries of the whole world. Six are con-
fined to Australia and New Caledonia.
Ccoke (ii, 929) describes 3 species belonging to this genus: Panicum
flavidum, Retz., P. punctatum, Burm., and P. fluztans, Retz. We retain all, but
substitute the older name geminatum for fluitans.
I. Lower spikes shorter than the internodes. Upper
involucrai glume shorter than the upper floral
glume eas ey ans re 1. P. flavidum.,
II. Lower spikes as long as or longer than the internodes.
Upper involucral glume about 3 the upper floral
glume aes a sae Fi oe see
III. Lower spikes as long as or shorter than the internodes.
Upper involucral glume mostly as long as the
upper floral glume ibe st ‘at ae 3. P. geminatum.
1. Paspalidium flavidum—A. Camus in Lecomte Fl. l’Indo-Chine, vii 1, 419 ;
Haines in Bot. Bihar & Orissa 1001 (erronee attribuens combitnationem
Staptio). —Panicum flavidum, Retz. Obs. iv (1786), 15; Griff. Notul. iii, 33 ; Ic.
Pi As. t. 139; fis. 67; Duthie Grass, N. W. Ind. 3, Indig. Fodd.:' Grass; t: vi;
4
2. P. punctatum.,
642 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
Fodd. Grass. N. Ind. 7; Benth. Fl. Austr. vii, 474; Hook. f. in F.B-I. vii, 28 ;
Cke. ii, 929.—P.. brizoides, Jacq. Eclog. Gram. (1813), 2, t. 2; Roxb. Fl. Ind.
i, 293; Dalz. & Gibs. Bomb. Fl. 290; Duthie Grass. _N. W. Ind. 2; Aitchis.
Cat. Panjab Pl. 159; Baker Fl. Maurit 433.—P. floridum Royle Ill. Bot.
Himal. 420.
Description : Cke. ii, 929.
Locality : Sind : Sanghar (Sabnis B762!).—Gujarat: Near Surat (Dalzell &
Gibson teste Cooke); Morvi, Kathiawar (Woodrow feste Cooke) ; Porbandar,
Kathiawar (Woodrow teste Cooke).— Konkan: Mulgaum (McCann 9579!);
Thana (McCann 8726!); N. & S. Konkan (Iuaws_ teste Cooke).— Deccan :
Poona (Bhide!) ; Mr. Gammie’s compound, Kirkee (Bhide 894!) ; Khandala
(Woodrow teste Cooke!).— S. WM. Country: Konankeri, in a small tank,
elevation 1,800 ft., rainfall 40 inches (Sedgwick & Bell 4964!); Belgaum
(Herb. Econ. Bot. Poona!).—Kanara: Yellapore (Talbot!) ; Halyal (Talbot
2095 !).
Distribution : Plains of India, Ceylon, tropical Asia (not in tropical Africa as
reported by Hook. f. and Cooke).
2. Paspalidium punctatum, A. Camus in Lecomte Fl. ]’Indo-Chine, vii, 419:
Haines in Bot. Bihar & Orissa 1001 (erronee attribuens combinationem
Staptio).—Panicum punctatum, Burm. Fl. Ind. (1768), 26; Hook. f. in
BeBe) Wil, 20 ICke. 11020,
Description : Uke. 1.c. .
Locality : Deccan: Poona (Woodrow, Lisboa) ; Ahmednagar (Woodrow).
Distribution : Throughout India in marshes, Ceylon, Malaya (not in tropical
Africa).
3. Paspalidium geminatum, Stapf in Prain Fl. Trop. Afr. ix (1920), 583;
Haines in Bot. Bihar & Orissa 1002.— Panicum geminatum, Forsk. FI.
Aegypt.—Arab. 18; Schweinf. in Bull Herb. Boiss. ii, App. ii, 19; Hack. in
Bull Herb. Boiss. iv, App. iii, 14; Hitchcock & Chase in Contrib. U.S. Herb.
xv, 30.—P. fluitans, Retz. Obs. iti, 8e¢ v, 18; Willd. Sp. Pl. i, 338; Kunth Enum.
i, 78; Steud. Syn. Pl. Glum. i, 59; Miq. Fl. Ind: Bat: iii, 455; Baker Fl; Maumit:
433 ; Cke. ii, 929.—P. brizoides, Lam. IIl., 1, 170 (om Retz.).—P. paspaloides,
Pers. Syn. i, 81; Kunth Enum. i, 77; Steud. i.c. 60; Boiss. FI. Or. v, 436%
Balfour f. Fl. Socotra 310; Hook. f. in F.B.I. vii, 30, in Trim. Fl. Ceyl. v, 135.—
P. beckmannieforme, Mikan ex Trin. in Spreng. Neue Entdeck. ii, 83;
Spreng. Syst. i, 309.—P. truncatum, Trin. Diss. ti. 130, ef Sp. Gram. Ic. t. 168.
—P.affine, Nees Agrost. Bras. 113.—P. drizaeforme, Pres] Rel. Haenk. 302 ;
Steud. l.c. 60.—P. numtdianum, Sieb. ex Schult. Mant. ii, 267 (zom Lam.).
P. carnosum, Salzm. ex Steud. l.c.—P. appressum, Doell. in Mart. FI. Bras. ii,
ii, 184.—P. glomeraium, Buckl. Prel. Rep. Geol. Agr. Surv. Tex. app. 3 (zon
Moench.).—P. turgidum, Cheval Sudania 25.— Paspalum appressum, Lam. Ill.
i, 176.—Dzigitaria appressa. Pers. 1.c. 85.--D. affinis, Roem. & Schult. Syst.
ii, 470.
Description : Cke. ii, 929.
Locality: Sind: Munchar Lake (Stocks deste Cooke); Tatta (Blatter &
McCann D611!, D612!).—Gujarat : Porbandar (Bhide!) ; Lasundra (Chibber!);
Ahmedabad, canal baaks (Sedgwick !) ; Cutch, Anjar, tank (Blatter 3745 !).—
Khandesh: Dhulia (Chibber!) ; Borod, growing in water, partly submerged
(McCann A97 !).—Kunkan: Mahaluxmi (Sabnis 5449!) ; opposite Kantwadi,
sandy shore, Bandra ( Vakil A99!) ; Victoria Gardens (McCann A100!) ; common
in Bombay Isl. (McCann !).—Deccan: Bund Gardens, Poona (Garade 343!) ;
sides of stream Dangar Guy, Ahmednagar (T. Cooke 6!) ; Manmad, river bed
(Blatter A94!); Sholapur Tank, in water (D’Almeida A95!); Pashan
(Gammie!); Igatpuri (Blatter & Hallberg 5492!).—S. 47. Country : Dharwar
(Sedgwick 3692!) ; Shiggaon (Sedgwick 2081!).—axara: Halyal (Talbot
2149!).
Distribution : More or less throughout India, Ceylon, Afghanistan, Arabia,
tropical Africa and America.
48, UROCHLOA, Beauv. Agrost. 52, t. xi, fig.1; Stapf in Prain Fly Drep:
Afr. 1x,,°586.
Perennial or annual. Leaves linear to lanceolate, flat; ligules a ciliate rim.
Racemes sessile or subsessile on a common axis, simple or nearly so; rhachis
more or less triquetrous with a low almost straight or zigzag facial angle or keel,
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 643
rarely strap-shaped ; pedicels solitary or in pairs, alternately to the right and
the left of the facial angle, usually reduced to short disc-tipped stumps, or if
binate the primary slightly longer. Spikelets close, coutiguous or slightly dis-
contiguous, 2 or irregularly piuri-seriate, glabrous or hairy, broad-ovate to
elliptic or lanceolate-oblong, awaless, usually more or less flattened or slightly
depressed abaxially, convex on the back, falling entire from the pedicels, soli-
tary or binate or in fascicles of 3-4, secund and abaxial on the rhachis involucral
glumes similar and subequalor more often dissimilar and very unequal in length,
the ower being the shorter, upper resembling and more or less equalling the
lower floral glume, 5-11 (mostly 7-) nerved. Lower floral glume &-7-, rarely
more-nerved the inner lateral nerves somewhat distant from the midnerve, pale
subequal to the glume, with well-developed inflexed flaps and sharp sometimes
marginate keels ; upper floral glume elliptic to rotundate-elliptic in outline,
very obtuse, with usually scabrid or barbellate mucro, narrowly involute, 5-7-
nerved, pale almost as long as the glume, 2-keeled, the sides tightly embraced
by the valve all along. Lodicules 2, small, broadly cuneate. Starrens 3;
styles distinct ; stigma plumose, laterally exserted upwards. Grain tightly
enclosed by the glume and pale, broadly to rotundate-elliptic, dorsally com-
pressed.
‘* Very similar to Brechiaria, but with the orientation of the spikelets inverted
and a short fine mucro from the very obtuse apex of the fertile valve. Although
very similar in general appearance, none of the species of Urochloa can be said
to approach closely members of the genus Brachiaria. Their affinities are
clearly inter se, suggesting a distinct line of evolution. The occurrence of
parallel states, one with glabrous, the other with pubescent spikelets, but other-
wise indistinguishable, runs almost through the whole genus. To this may be
added the presence of a submarginal fringe in the lower floret, almost normal
in some and very rare in other species, and apparently in no case correlated
with other characters.’’ (Stapf l.c.).
Species about 18.—Hot parts of the Old World, one in America, but perhaps
introduced.
Cooke describes 3 species of Panicum which belong here: Panicum prostra-
tum, Lamk., P. settgerum, Retz., and P. Javanicum, Poir., to which we add
Urochloa marathensis, Henrard.
A. Spikelets up to 2 mm. long ne ae lO. veplans,
B. Spikelets 2°5-5 mm. long
I. Spikelets lanceolate, acuminate see ea "OSCE Sera.
II. Spikelets ovate to elliptic-oblong
1. Leaves about 3cm. long. Racemes 2 cm.
U. marathensts.
lone 2s wat . a an
2. Leaves 3°5-15 cm. long. Racemes 2°5-5 cm.
long 34. wee i .. 4 UO. Helopus.
1. Urochloareptans, Stapfin Prain FJ. Trop. Afr. ix, 601; Haines in Bot.
Bihar & Orissa 1003.—FPanicum reptans, Linn. Syst. Nat. ed. x, 870; Hitchc.
& Chase in Contrib. U.S. Nat. Herb. xv, 36, fig. 17, excl. P. grossarium.
—Fivepens, N. i, Burm. Fl. Ind. 26, t. 11, f. 1; Rottl. in Neue Schrift. iv, 182 ;
Roxb. Fl. Ind. i. 302; Boj. Hort. Maur. 364 (zow Linn.).—P. prostratum,
Lam. Ill. i, 171; Mig. Fl. Ind. Bat. iti, 446; Griseb. Fl. Brit. W. Ind. 546;
Schweinf. in Bull. Herb. Boiss. ii, App.ii, 20; Baker Fl. Maur. 435; Duthie
List: Grass. N. W. Ind. 6; Ill. Indig. Fodd. Grass. t. 45; Fodd. Grass. N. Ind.
Reem boss, El Or, v, 438; Dalz. & Gibs. 290; Hook.f. in F.B.1. vii, 33;
Cke. ii, 932; Merrill in Philipp. Journ. Se. i, 355.—P. darbatum, Lam. 1..c.
—P. cespitosum, Sw. Fl. Ind. Occ. i, 146. —P. Szederz, Link. Hort. Berc!. i, 207.
—P. procumbens, var. Nees Agrost. Bras.109.—P. crispum, Lianos, Fragm. 42.
~— P. insularum, Steud. Syr. Pl. Glum.i, 61.—P. calacczense, Steud.. l.c. 65.
—P. aurelianum, Halein Wood Classb. ed. iii, 787.—-P. viaticum, Salzm. ex Doell
in Mart. Fl. Bras. ii, ii, 155.—P. mareginatum, Vahlex. Hook. f. 1. c.—&4racht-
aria prostrata, Griseb. in Abh. Ges. Wiss. Goett. vii, 263.
Description: Cke. ii, 932, under Panicum prostratum.
Locality : Gujarat: Ahmedabad, famine grass plot, Redar (Herb. Gujarat
College!) ; Ghad, on black soil (Sedgwick 1124!).—Khandesh : Nandurbar,
(Mamlatdar of Nandurbar!) ; Umalla, Tapti bank (Blatter & Hallberg 5229 !);
Bor, Tapti (Blatter & Hallberg 4417!).—Konkan: Vasco da Gama (Bhide !):
Matunga, salt pans (Sabnis 9572!) ; Byculla (McCann 9577!).—Deccan : Kirkee
644 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXX11
(Bhide!); Poona (Herb. Econ. Bot. Poona!); Deolali (Blatter A107!) ;
Sholapur (D’Almeida Al08!, A109!).—S. M7. Country: S.W. of Dharwar,
elevation 1,800 ft., rainfall 90 inches (Sedgwick & Bell 4437!) ; Dharwar,
under trees on black soil, elevation 2,400 ft., rainfall 34 inches (Sedgwick 2882 !).
Distribution: Plains of India, Ceylon, Tropics generally, subtropical
America, also tropical Arabia and the Mascarenes.
2. Urochloa setigera, Stapf in Prain Fl. Trop. Afr. ix, 598; Haines in Bot.
Bihar & Orissa 1003.—FPanicum setigerum, Retz. Obs. iv, 15; Roxb. Fl. Ind.
i;.299 ; KunthsEnum.i, 90 @xcl. syn. nonnuliis) > Hookt: in 3. Br. il, of.
Trim: Fl. Ceyl..v, (41; Cke. 11, 933.--7. affine, ‘Poir. Eneycl: “Suppl. in, 278
(ex Kunth).
Description: Cke. ii, 933, under Panicum setigerum.
Locality : Sind: Mirpur Sakro (Blatter & McCaun D609!); Gharo (Blatter
& McCann D610!).-Gujarat: Rajkot, Kathiawar (Woodrow 45 deste Cuoke! ).—
Khandesh: Dhulia (Herb. Econ. Bot. Poona!).—Aonkan : Bassein (Paranjpe !).
—Deccazx : Ganeshkhind Bot. Gardens (Herb. Econ. Bot. Poona!) ; Mangiri
Farm, 11 miles S. E. of Poona (Herb. Econ. Bot. Poona !).
Distribution : India, Ceylon, Mauritius, tropica! and 8. Africa.
Note: We donotthink there is any good reason for retaining Hook. f.’s
var. tomentosa which was also mentioned by Cke. 1.c. We refer to the note
given above after the general characteristics of the genus and add what Stapf
says with regard to this species in particular: ‘The African specimens have
glabrous spikelets. In India, however, the pubescent form appears to be pre-
valent.’
3. Urochloa marathensis, Henr. in Mededeel. Rijks Herb. 43 (1922), 1-3, pl. 1.
— Panicum marathense, Henr. in Herb. Lugd. Bat.
Distribution: Annual, dwarf and robust, glaucous, branching from the
lower geniculations. Stems low, striate, quite glabrous, few-noded, nodes
pubescent. Leaf-sheaths terete or slightly compressed, striate, lower ones gap-
ing, upper ones tight, shorter than the internodes, hirsute with bulbous-based
hairs. Ligule very short, covered with long hairs. Blade cordate at the base,
linear-lanceolate, gradually acutate, more or less 3cm. long, 5-7 mm. broad,
flat, on both sides but especially on the upper sparingly covered with spreading
bulbous-based hairs, margin thickened, distinctly undulate, fimbriate with iong
tubercular hairs. Racemes 2-3, distant from each other more or less 1 cm.,
2 cm. long, stout, erect-patulous or finally reflexed ; rhachis subtrigcnous,
pilose at the base, more than twice as narrow as the spikelets slightly undulate,
the angles scaberulous, otherwise glabrous, giving off solitary pedicels. Spike-
lets biseriate, broadly elliptic, very acute, 4 mm. long, anteriorly almost flat,
posteriorly very convex, green, glabrous. Involucral glumes very unequal.
Lower almost 3 of the spikelet, subobtuse, 5-nerved, upper as long as the
spikelet, very acute, distinctly 7-9-nerved. Lower floral glume equal in
shape to the upper involucral glume, flat, 5-nerved, glabrous on the back,
tubercular-echinulate near the margin, except near the tip and base, paleate,
Upper floral glume shorter than the preceding glumes, elliptic with a
rounded tip, rigid, opaque, brown-straw-coloured, rugulose, long caudate, 5-
nerved, bullate below.
Locality: S.M. Country and N. Kanara (A. P. Young ex Henrard). We
have not seen the specimen.
Distribution : So far endemic.
Note: Henrard hasa variety from the same localities : var. Velutina which
differs from the type by the densely pubescent spikelets. What we saidina
note uuder the previous species can be applied here.
4. Urochloa Helopus, Stapfin Prain Fl. Trop. Afr. ix, 595; Haines in Bot.
Bihar & Orissa 1002.—U. pubescens, Kunth Rev. Gram.i, 31, Enum. Pl. i,
74.—U. panicoides, Schult. Mant. ii, 595 (zou Beauv.).—FPanicum Helopus,
Trin. in Sprene. Neue Hntdeck:*11, $4" Panic. (Gen. 150; -e7 “Gram le.wser
Descr. ii, t. 183; Nees Agrost. Bras. 117; Steud. Syn. Pl. Glum.i, 57; Duthie
List Grass. N. W. Ind. 4, Fodd. Grass. N. Ind. 8.—P. Helopus var. glabrescens,
K: Schum. in Engl. Pll. Ost.—Afr. C. 101; Stapf in Dyer Fl. Cap. vii, 392.—
P. hirsutum, Koen. ex Roxb. Fl. Ind, i, 300.—P. Kenigi, Spreng. Syst. i,
311.-—-P. hochstetterianum, A. Rich. Tent. Fl. Abyss. ii, 369.—P. geminatum,
Hochst. ex A. Rich. l.c. (zon Forsk.).—P. controversum, Steud. Syn. Pl.
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 645
Glum.i, 60; Schweinf. in Bull. Herb. Boiss. II, ii, 19.—Setaria? hirsuta,
Kunth Rev. Grain. i, 47, Enum. Pl. i, 157.—S. pzlifera, Spreng. Syst. iv, Cur.
Post. 33.—Panicum javanicum, Hook. f. in F.B.I. vii, 35 (wom Poir, partim) ;
Cke. ii, 933 (partz12).
This is the species which was described by Hook. f. and many others,
amongst them by Cooke, under the name of Panicum javanicum, Poir.
According to Stapf P. javanicum, frequently confused with U. Helopus, is U.
panicoides, Beauv., a distinct species. A numberof synonyms and references
have, according to the same authority, to be excluded from the F.B I. under
Panicum javanicum : Bentham’s Flora of Australia (vii, 476) mentions Panicum
Flelopus, but itis partly Brachiaria notochtona, Stapf (Panicum notochtonum,
Domin) and partly Arachiaria ramosa, Stapf. T. 7 in Duthie’s Fodd. Grass.
does not represent Panicum Helopus, but Brachiaria ramosa, Stapf. Urochloa
panicotdes, Beauv. isasynonym of Panicum javanicum, Poir. P. trichopus,
Hochst. is Uvochloa trichopus, Stapf.
As a number of foreign elements have crept into the usual descriptions of our
species we give Stapf’s description of P. Helopus.
Annual. Stems tufted, 30-60 cm. high, erect or geniculately ascending from
a short sometimes rooting base, frequently sparingly branched from the lower
nodes, 4-10-noded, intermediate internodes like the uppermost (peduncle) very
sparingly pubescent or almost glabrous. Leaf-blades lanceolate to linear-
lanceolate from a wider and semi-amplexicaul base, 3°5-15 cm. by 8-12 mm.
rarely up to 25cm. and then linear and narrowed towards the base, soft, flat,
pale or yellowish-green, loosely and often finely hirsute with tubercle-based
hairs, rarely almost glabrous, margins usually crisped or wavy and more or less
ciliate. Sheaths somewhat loose, pale, striate, densely ciliate upwards, more or
less shortly hirsute with the hairs tubercle-based, nodes pubescent to subvillous.
Ligules a densely ciliate rim. Inflorescence of mostly 4-7 erect or at length
more or less spreading stiff or slightly flexuous sessile or subsessile spiciform
secund racemes ; common axis 1:2-5 (rarely 75 cm.) long, subsemiterete below,
inuch flattened upwards, pubescent. Racemes moderately dense, 2-seriate,
simple, 25-5 cm. long, solitary or here and there approximate and then
unevenly distributed ; rhachis straight or slightly wavy, flat on the back, about
1mm. wide, villous atthe base, glabrous upwards, rarely sparingly hairy,
angles scabrid ; internodes up to 1 mm. long ; pedicels solitary, reduced to short
stout stumps with discoid tips, frequently bearing some long spreading hairs.
Spikelets laterally contiguous or subcontiguous ovate to elliptic-oblong, very
acute, 4-5 mm. long, greenish, glabrous or pubescent. Involucral glumes
dissimilar ; lower broad-ovate, subobtuse to acute, clasping at the base, 15-2
mm. long, glabrous or sparingly and minutely pubescent, 5-nerved upper one
corresponding in size and outline to the spikelet, prominently 7-11-(mostly 9-)
nerved, glabrous or pubescent. Lower floral glume very similar to the upper
involucral glume, but flat or slightly depressed, 5-7-nerved with the inner side-
nerves distant, glabrous or pubescent, pale oblong, acute, slightly shorter than
its glume. Anthers 2 mim. long. Upper floral glume rotundate-elliptic,
greenish to pale brown, 2°5-3 mm. long, pale finely transversely rugose or
granular, mucro up tol mm. long, sparingly barbellate. Grain rotundate-
elliptic on outline, much compressed, about 2 mm. long, yellowish or greenish.
Locality: Gujarat: Baroda (Cooke deste Cooke).—Deccan: Katraj Ghat,
11 miles S. E. of Poona (Shevade!); Poona (Woodrow!) ; Deolali (Blatter
A110!) ; Ganeshkhind Bot. Gardens (Patwardhan !) ; Chattarshinji (Bhide!) ;
Mangiri, near Poona (Gammie 15344!); Akola (Mamlatdar of Akola!) ;
Dapuri near Poona (Jacquemont 482!).—S. W/. Country: Dharwar, elevation
2,500 ft., rainfall 35inches (Sedgwick 2157!); Kilgerry (Talbot 2419 !) ;
Haveri (Talbot 2284!) ; Bijapur (Meebold 11201!) ; Badami (Woodrow feste
Cooke).— Kanara: Kulgi, elevation 2.000 ft. (Talbot 2283!) ; Nundgod (Mam-
ratdar of Nundgod).
Distribution: Plains of India, Ceylon, tropical and S. Africa, Mauritius.
49, Ecuinocuioa, Beauv. Agrost. 53, t. 11, fig. 2; Stapf in Prain
Bl Trop. Aire ix, 604;
Annualor perennial. Leaf-blades from a slightly constricted or equally wide
rarely much attenuated base. Ligules 0 or represented by a transverse fringe of
hairs. Panicles of crowded or loosely arranged secund speciform branches
646 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
mostly bearing spikelets from the base or near it. Spikelets ovate to elliptic- or
lanceolate-oblong, usually cuspidate or awned, very convex on the back, flat or
slightly depressed in front, falling entire from the pedicels, 2-nate or clustered,
secund and abaxial on the triquetrous rhachis of racemosely arranged false
spikes Involucral glumes unequal, membranous, the lower much shorter,
more or less ovate from a clasping base, 3-5-nerved, often mucronate, the upper
corresponding in length and outline to the spikelet (as seen from the back),
very concave, 5-7-nerved, acute, cuspidulate or cuspidate, rarely produced into
a short awn. Lower floret equalling the upper glume (excluding cusps or,
awns) ; lower floral glume very similar to the upper involucral glume, but flat
or depressed on the back and often with a more pronounced cusp or an awn ;
pale equal tothe body of the valve, or in barren florets more or less reduced,
hyaline, finely 2-keeled. Upper floral glume ovate to elliptic-oblong, apiculate
or obtuse, very convex on the back, subcoriaceous or crustaceous, polished,
faintly 5-nerved, margins firm, involute up to near the tip, then flat, not emb-
racing the tip of the pale, pale sub-equal to. the glume and similar in substance,
with rounded keels and flaps which thin out towards the flat slightly recurved
tips. Lodicules 2, cuneate, fleshy. Stamens 3. Styles distinct; stigmas
plumose, exserted from near the tips. Grain broad-elliptic dorsally flat,
ventrally convex ; hilum punctiform, subbasal.
Species about 20-25.—The warm regions of both hemispheres.
Note: Jt will be useful to remember what Stapf says regarding this genus
l.c.605: ‘ The segregation of the numerous fornis which make up the genus
Echinochloa and their reduction to more or less well definable species is still
unsatisfactory, mainly owing to their apparently endless variability and the
difficuity, if not impossibility, of discriminating between stable and unstable
modifications and the effects of hybridization. Here, as in other cases, observa-
tion in the field and experiment will have to decide.’
Cooke describes under Panicum 2 species which have to be referred to
Echinochloa: P.colonum and P. stagninum. We add &. Crus-Galli which
Cooke thought did not occur anywhere in the Bombay Presidency.
I. Lower involucral glume and upper floral glume
equally acute or cuspidate 1. &. Colona.
If. Lower involucral glume and upper floral glume
cuspidate or produced into an awn, the latter
more than the former.
1. Ligule 0 .... . 2: &. Crus-Gallt,
2. Ligule a fringe of stiff hairs or absent in the
uppermost leaves . a sac oe, ofl. Sta Gultga:
1. Echinochica colona, Link Hort. Berol. ii, 209; Parl. Pl. Nov. 40; Hitchc.
in Gray Man. Bot.ed..7,.138.,efin-Contrib..U; Si. Nats Herb: oxi 7135 evan
256, xvili, 345; Stapf in. Prain Fl. Drop. Airs 1x, 2607 ; Hainesminuebor
Bihar & Orissa 997.—Z. zonalis, Parl. Pl. Panorm. i, 119.— Panicum colonum,
Lion, Syst. Veg..ed. 10 (1759), 870, Sp. Pl. ed..u, 84; Jacq. Eclog,Gramiat:
32; Roxb. Fl. Ind, i, 299; Nees Agrost. Bras. 119; Steud Syn. Pl. Glum. ij,
46; Benth. Fl. Hongk. 411, et Fl. Austral. vii, 478; Griseb. Fl. Brit. W. Ind.
545 ; Baker Fl. Maurit, 438; Duthie List Grass, N. W. Ind. 3, Indig. Fodd,
Grass. t..4, Fodd. Grass. N. Ind. 4; Boiss. Fl. Or. v-435: Balfst. Botssocotra
310; Hack. in Bol. Soc. Brot. vi, 140 ; Schweinf. in Bull. Herb. Boiss. ii, App.
ii, 20,95; Hook. f.in F.B.I. vii, 32; Cke. ii, 931.—P. arabicum, Nees ex
Steud. Nomencl. ed. ii, 252, ef Syn, Pl. Glum. i, 63 (Jartim).—P. brizotdes,
Linn. Mant. ii, 184.—P. fetrastichon, Forsk. Fl. Aegypt.—Arab. 19.—P.
cuspidatum, Roxb. FI. Ind. i, 301; Duthie Grass. N. W. Ind. 3; Steud. 1. c.
47.—P. pseudocolonum, Roth. Nov. Sp. 47; Steud. l.c. 46.—P. zonale, Guss. Ind.
Sem. H.R. Bocc. 1825, e¢ Fl. Sic. Prodr. i, 82.—P. numidianum, Presl. Cyp. &
Gram. Sic. 19 (zom Lam.).—P. Daltoni, Parl. ex Webb in Hooker Niger FI.
185.— P. eguitans, Hochst. ex A. Rich. Tent. Fl. Abyss. ii, 365.—P. Crus-Galli
var. colonus, Coss. Glum. Expl. Alger. 28.—P. Peliveri, Kotsch. ex Griseb.
lsc. (707 rin...)
Description: Cke. ii, 931.
Locality : Sind: Karachi (Nankad !); Mirpurkhas (Bhide!, Sabnis B1176!) ;
Sind (Blatter!) ; Larkana, barren plains (Sabnis b93!, B95!, B457!) ; Sanghar
(Sabnis B899!, B894!) ; Nasarpur, clayey soil (Sabnis B1048!) ; Mirva Canal,
Khairpur Mirs (Sabnis B263!); Khairpur Mirs (Sabnis B337!) ; Sehwan to
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 647
Laki (Sabnis B63!) ; Hyderabad (Sabnis B49!) ; Pad-Idan (Sabnis B516 !} ;
Chuar Ch. (Blatter & McCann D616!, D621!) ; Baghar (Blatter & McCann
D617!); Mirpur Sakro (Blatter & McCann D618!, D620!); Ghulamalla
(Blatter & McCann D619); Shikarpur (Woodrow).—Guwjarat: Lasundra
(Chibber!); Anjor (Cutch), brackish water (Blatter 3743!) ; Ahmedabad
(Cowper!) ; Morvi, Kathiawar (Wcodrow).—Khandesh: Muravad, Tapti
bank (Blatter &- Hallberg 4435!); Bor, -Tapti Island (Blatter &
Hallberg 4439!); Dadgaum (McCann Al104!); N. slope of Chanseli
(McCann <Al105!).— Konkan: Bombay, Salt swamps (Woodrow J) ;
Vetora (Sabnis 33589!) ; Charni- Road, Bombay (Sabnis 4292!) ;
Bombay, very common (McCann!); Alibag, rice fields (Ezekiel!).— Deccan :
Purandhar Fort (Bhide!, McCann 5520!); Nira Canal, Poona District (Chib-
ber!); Khbandala, very common (McCann A101!); Sholapur (D’Almeida
Al02!); Igatpuri (Blatter & Hallberg 5487! ; McCann 4331!); Deolali (Blatter
A103!); Poona, canal (Ezekiel !).—S. 47. Country : Dharwar (Sedgwick 2655!) ;
Castle Rock (McCann Al106!); Londa (Woodrow).—Kanara: Dongi Nallah
(Talbot !); Karwar (Talbot 614!).
Distribution : Throughout the plains of India, Ceylon, all over the tropics and
the warm-temperate regions of the world. Probably of African and Indian
origin according to Stapf.
The sameauthor is of opinion that Panicum frumentaceum, Roxb. which
is grown in India asa grain crop, is evidently descended from “chinochloa
colona. Roxburgh’s species may, therefore, be treatea as a variety :
var. frumentacea, Blatter & McCann.—AFchinochloa frumentacea, Link. Hort.
Berol. i, 204; Aitchis. Cat. Panjab Pl. 161.— Panicum frumentaceum, Roxb.
iiads i; 304:;Schult. Mant. it, 230;.Trin. Sp.. Gram. Ic.,t. 164; Duthie
Grass. N. W. Ind. 4, Field & Gard. Crops 3. t. 24, Fodd. Grass. N. Ind. 8.—
P. Crus-Galli, var. frumentaceum, Trim. Cat. Ceyl. Pl. (1885), 104.—
Echinochloa Crus-Galli, var. frumentacea, Haines in Bot. Bihar & Orissa
998.—Panicum stagninum, Retz. var. frumentacea, Cooke in Cke. ii, 931.—
Oplismenus frumentaceus, Kunth Rev. Gram.i, 45, Enum. Pl. i, 146; Daiz.
& Gibs. Bomb. Fl. Suppl. 98.
Description : Tall, robust. Stems erect, from 60 to 120cm. high. Panicle
often nodding. Spikes secund, incurved, crowded. Spikelets mostly 3-nate,
unequally pedicelled, one at least sessile, varying from hispidulous to almost
glabrous, and from acute to cuspidulate or rarely distinctly cuspidate.
Locality : Cultivated in and near the Ghat districts.
2. Echinochloa Crus-Galli, P. Beauv. Agrost. 161; T. Nees Gen. FI. Gerin.
Monocot. i, t. 21; Reichenb:. Ic. Fl. Germ. i, t. 29, fig: 1411, 1412; Hitche. in
Contr. U.S. Nat. Herb. xii, 213 ; Stapf in Prain F]. Trop. Afr. ix, 610; Haines
in Bot. Bihar & Orissa, 998.—&. commututa, Schult. Mant. ii, 267.—
E.. hispidula, Nees in Royle il. Bot. Himal. 416; Dalz. & Gibs. Fl. Bomb.
Suppl. 98.—FPanicum Crus-Galli, Linn. Sp. Pl. ed.i, 56; Fl. Dan. t. 1564;
migsts Gram. Austr. ii, 15, t. 19; Knapp Gram... Brit. «i,. Trin.’ Sp. Gram:
faeet. 4161; 1625 Nees FI. “Afr. Austr. 58 3 Steud. Syn. Pl:: Glum,: 1; - 47:
Benth. Fl.. Austr. vii, 479 (partim); Duthie List. Grass. N.W. Ind. .3
Wwagecmy ts. A. fig: 1;) Boiss, .Pl- Ors v, 435; .Hook. f. in F:B.1. vit; 3
(partim) ; Stapf in Dyer Fl. Cap. vii, 397.—P. Aispidum, Forst. Prodr.
7; Nees Agrost. Bras. 257.—P. Aispidulum, Retz. Obs. v, 18; Lam. Ill. i, 171;
Roxb. Fl. Ind. i, 306; Nees Fl. Afr. Austr. 57; Royle J. c. 420; Stend. Syn.
Pl. Glum. i, 47.—-P. oryzinum, Gmel. Syst. i, 157.—P. stagninum. Host. Gram.
Austr. iii, t. 51 (zon Retz.).—P. Hostii, Marsch. Bieberst. Fl. Tauro-Cauc. iii,
5/7.—P. limosum, Presl ex Nees Agrost. Bras. 257.—Milium Crus-Galli, Moench
Meth. 202.—-Ofplismenus Crus-Galli, Dumort. Agrost. Belg. ; Kunth Rev.
Gram. i, 44, e¢ Enum. i, 143 (excl. syn. P. zonale).—O. limosus, Presl., Rel.
Haenk. i, 321: Kunth Enum. i, 144.—Orthopogon Crus-Galli, Spreng. Syst. i,
307.—O. Retzi1, Spreng. l.c.
Description: Annual, up to 1 m. high. Stems geniculately ascending,
branched below, compressed towards the base, glabrous and smooth, inter-
nodes enclosed or exserted. Leaf-blades linear, base scarcely narrowed,
narrowed to an acute point, 7-25 cm. by 6- over 12 mm., flat, subflaccid,
glabrous, more or less dull greyish-green, smooth or scaberuious below,
particularly towards the tip, margins finely cartilaginous, scabrid to almost
smooth, Sheaths somewhat loose, the lower often compressed, whitish and
648 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
thin, the upper subherbaceous, all smooth, glabrous and striate except the
basal which are pubescent above their insertion. Ligules 0, junction of blade
aud sheath glabrous inside marked by a brown zone. Panicles erect, strict or
flexuous, at length exserted, 7°5-20 cm. long; axis triquetrous, scabrid ;
branches few to about 15, solitary or 2-nate, suberect or spreading, distant
except the uppermost or all more or less approximate forming a ‘ lobed’
panicle, the lower 2°5-6°2 cm. long, forming rather stout dense mostly
many-ranked simple or subcomposite subsecund sessile false spikes ;
rhachis triquetrous, scabrid, coarsely bristly, particularly near the nodes;
pedicels fascicled or 2-nate, very short, up to 1 mm. long,.scabrid, bristly at
the base, tips subdiscoid. Spikelets crowded, ovate-eiliptic in outline, acute,
cuspidate or awned 2°5-3 mm. long, greenish or tinged with purple. Lower
involucral glume membranous, very broadly ovate, clasping at the base, obtuse
to subcuspidate, 1 mm. long, 5-nerved scaberulous; upper herbaceous-mem-
branous, very broadly ovate-oblong, concave, acute, cuspidate, as long as the
spikelet, 5- or (near the tip) 7-nerved, rigidly pubescent between the scabrid
and spinulose nerves. Lower floral glume similar to the upper involucral glume,
but flat or depressed on the back, cuspidate or produced into a scabrid often
long flexuous awn, 7-nerved (at least at the tip), pale elliptic, shorter by 4 than
its glume, keels scaberulous upwards; upper floret hermaphrodite, elliptic-
ovate in outline, cuspidate, over 2mm. !ong, whitish or yellowish, polished,
glume and pale subcoriaceous. Anthers oblong. Grain broad-elliptic in outline,
15mm. Jong:
Locality: Sind: Mirpurkhas (Bhide!); Ghulamalla (Blatter & McCann
D613!); Keti (Blatter & McCann D614!).—Gujarat: Stream near Prantij
(Sedgwick !) ; the Bokh, Prantij Taluka (Sedgwick 1144!).—Deccan: Poona
(Woodrow !).—S. M7. Country: Aluarar, elevation 2,000 ft., rainfall 35 inches
(Sed¢wick 3096 !) ; Bidi, elevation 2,500 ft., rainfall 50 inches (Sedgwick 3076 !).
— Kanara: Halyal (Talbot 2167 !).
Distribution : Common through the greater part of India and Malaya; as a
weed throughout the warm temperate countries of the northern hemisphere,
rather rare in the tropics of Africa and the New World and south of the Tropic of
Capricorn (Stapf).
Uses: See Duthie, Fodd. Grass. N. Ind. 6; Vasey Agric. Grass. Unit. States
ed. ii, 27; Maiden Man. Grass. N. 8. Wales, 38-41; Yearbook U. S. Dept.
Agric. (1902), 580-582.
3. Echinochloa stagnina, P. Beauv. Agrost. 161; Stapf in Prain Fl. Trop. Afr.
617 —E£. scabra, Roem. & Schult. Syst. ii, 479.—Panicum stagninum, Retz.
Obs.-v, 173 ‘Roxb. Fi. Ind:1;295:; (Grah:iCatal.c 2374Necs, “Actost. Brace: 26e
Trin. Pan. Gen. 128, ef in Mém. Acad. Pétersb. 6me sér. iii, 216; Steud. Syn.
Pl. Glum. i, 47; Stapf in Dyer Fl. Cap. vii, 394; Cke. ii, 930.—P. scabrum,
Lam. fll.i, 171,¢2 Bocyel.iv. 7445 Nees 7. ic/5 Steud, 1: ¢-—PGall7,- Thunb,
Prodr. 18, e¢ Fl. Cap. ed. i, 389, ed. Schult. 103.—P. Crus-Galli, Woodrow in
Journ. Bomb. Nat. Hist. Soc. xiit (1901), 433 (vom Linn.).—P Crus-Galli, var.
stagninum, Fenzl. in-Ind. Sem. Hort. Berol. 1850; Hook. f. in Trim. Fl.. Ceyl
v, 1385; Prain Beng. Pl. 1174.—P. Crus-Galli, vars maximum, submuticum et
leiostachyum, Franch. Contr. Fl. Congo Franc. in Bull. Soc. Hist. Nat. Autun.
viii, 347.—P. pictum, Nees Fl. Afr. Austr. 59 (mon Agrost Bras.).—P. Burgi,
A. Cheval. in Rev. Cult. Colon. vii, 513-520.—P. Lelievrez, A. Cheval. l.c.
516.—P. oryzetorum, A. Cheval. l.c.—Orthopogon stagninus, Spreng. Syst. i,
307.—Oplismenus stagninus, Kunth Rev. Gram. i, 44, e¢ Enum. i, 144 (partzm) ;
Dalz. & Gibs. Fl. Bomb. 292.—O. scaber, Kunth Rev. Gram l.c. 44, Enum.
lc. 145.
Description: Cke. ii, 930, under Panicum stagninum.
Locality: Sind: Ghulamalla (Blatter & McCann D615!).—Konkan : Virar,
on bank of a tank (McCann 9585!, 9584!).—Deccan: Igatpuri (Blatter &
Hallberg 5473!) ; Khandala, in water in the smaller village tank (McCann
27441!) ; Panchgani (Blatter & Hallberg B1241!).—S. WM. Country: Hulkop,
elevation 2,000 ft., rainfall 50 inches (Sedgwick 3175!) ; Chikkerur, Taluka
Kod, water hole by road (Sedgwick 1969!) ; Bomigatti tank (Sedgwick 3830!) ;
common in the Carnatic (Sedgwick).— Kanara : Pardhani (Talbot 3136!).
Distribution: More or less throughout India, Ceylon, tropica] and S. Africa.
Uses : Considering tropical African conditions Stapf observes (l.c 619):
‘ This grass deserves every attention on account of its locally abundant supply
REVISION OF THE FLORA OF THE BOMBAY PRESIDENCY 649
and high sugar-content. Chevalier states that itis the most useful of all the
plants growing in a wild state in the neighbourhcod of Timbuctu. Every part
of it is utilized. It yields excellent fudder, material for thatching and caulking,
is burned to produce a salt used inthe manufacture of soap and indigo, the
grains are eaten and the canes are gathered for extracting sugar or preparing
vingigre or a beverage resembling cider. Sir John Kirk also describes it as one
of the richest of fodder grasses. Although typically a perennial with long
rhizomes creeping in the mud of swam gs, lakes and rivers, it seems on tempor-
arily flooded land to flower frequently the first year and then to behave as an
annual.’
(Fo be continued).
A COLLECTING Rie TOL ADAK
BY
V. S. LAPERSONNE
Assistant Curator, Bombay Natural History Society
Parr II
(With two Plates)
(Continued trom page 517 ot this Volume)
Shushal (14,500 feet) our next halt, is a delightful spot. We had
camped in a willow garden, with the ever-present ‘Gompa’
(monastery) perched on a hillock. We were ‘ pleasantly’ surprised
to find next morning, that a heavy fall of snow had blotted out the
landscape, which was not usual for the time of the year at those
altitudes, unless something was radically wrong somewhere.
Half a mile behind our camp, the Bar-headed Geese (Anser indicus)
were breeding on a shallow ‘jheel’ and appeared to lay their eggs
on little grass-topped islands. On our arrival there were quantities
of egg-shells scattered about and only one young in down, which
was secured.
It is probable that the young are preyed upon by birds of prey.
The Tibetan, for a certainty will not molest these birds, believing
that with the hatching out of successful broods, the year’s harvest
is as good as assured. They even go sofar as to appoint a ‘ watch-
man’, who does not seem very enthusiastic over a job that keeps
him out all day, and will no doubt give any naturalist the necessary
help, for a consideration, and be glad to see the last of the ‘ darned
things’.
A suitable tip in the right quarter often got us much invaluable
information on the breeding sites of Owls, Partridges and Snow-
cocks, though some of the local sportsmen drew on their imagination
freely and appeared quite content merely with having strolled out
with a Sahib. Others again ssemed overjoyed with the prospect of
a morning with the guns and felt quite hurt if their lively chatter was
cut into by the interpreter, with the remark ‘ that they should keep
their reminiscences for their great-grandchildren on long winter
nights’.
The Large Eagle Owl breeds among bare rocky hills, usually on
the precipitous sides, in huge ‘rifts’. At Shushal we secured a
parent bird with two young in down. The Brahminy was aiso seen
leaving the hillside, and in among the reeds the Mallard had changed
the plumage by which we know him so well, for one quite insignifi-
cant.
Brandt’s Snow Finch (1/7. brandti hematopyga) appeared in large
flocks and as Col. Meinertzhagen observed, these were all females.
The males seemed to keep to the higher altitudes, and even as
late as June neither sex showed any signs of genitai development,
A COLLECTING TRIP TO. LADAK 651
Among the Rose Finches, Zrythrina rubicilla scvertzovi appeared
the commonest along the Pangkong Lake, frequenting the most
desolate regions.
Another species of Rose Finch, which roosted in hundreds on the
willow trees around our camp, has been identified as a new race.
Col. Meinertzhagen has named this A7ythrina rubicillotdes lapersonnet
and gives the following :—
‘Ladak specimens are generally paler above and below, the
upper parts not washed with such a deep crimson and the under
parts of a paler and more delicate rose-colour than occurs among
Kansu specimens. Females are also generally paler.
‘ A single female was obtained at Leh on May 7 and they occurred
in large numbers at Shushal in June, where many hundreds used to
roost in a willow grove in which I was camped. Birds seen at
Chumathang at 13,U50 feet above Leh on the Indus were also
believed to be of this species. ‘They are not soshyas &. 7”. severi-
zov7 and seem to prefer a Jess desolate type of country. They were
busy building at Shushal in June, many incomplete nests being
found in willow-trees and furze (Cavagana) from one to twelve feet
from the ground.
‘Birds recorded under Evythrina rhodochlamys by Hume and
Henderson (‘‘ Lahore to Yarkand,’’ p. 259) and by Sharpe (Second
Yarkand Mission, p. 42), are withcut doubt &. rubictlloides laper-
sonnet. The birds observed by Henderson at Tutyalak (13,000 feet)
and recorded as severtzovz were more likly to be Pyrrhospiza, which
is common there. It is not suitable country for Arythrina rubicilla
or rubicillordes.’
Leaving Shushal onthe 22nd June we cut across to the Indus over
a mountainous pile of rocks, which leaves one with the impression
that this particular part of the globe has been left unfinished by the
Great Architect and the material strewn about.
We follow the hot dantesque gorges of the Indus as they wind in
and out amidst the silent rocky mountains whose peaks cut a jagged
line against an azure sky. Flat-topped hamlets punctuate the mono-
tony of sun-baked rock and muddy water where the gorges open
out; but such openings are few. The general aspect is that of the
protracted neck of a bottle, so narrow in eels that it would not
be a feat to kick one’s hat across.
While on the theme, a brief aoe aA account of the
upper reaches of the Indus which of recent years, political move-
ments have made accessible, would not bore the reader more than
we have done already.
The remarkable phenomena of the upper reaches of the Indus
Valley are the clearness and transparency of the atmosphere and
the great difference between the sun’s temperature 72 vacuo and the
air temperature in the shade. This difference as recorded at Leh
occasionally exceeds 90°.
The sources of the Indus have been traced to the glaciers of the
Kailas group of peaks and except for the remarkable instance of
transverse drainage, pursues an even north-westerly course amidst
an array of mountain scenery which for its majesty of sheer altitude
is unmatched by any in the world.
652 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
At Leh the river has already pursued an even course of 300 miles
from its source and except for the divergence which occurs about
100 miles above Leh, continues on a comparatively gentle placid
course over its sandy bed, when it captures the Shyok River (a
tributary almost as large as itself) and fed by the monster glaciers
of the Kara-Koram Range. ‘The combined streams still hold on a
north-westerly trend for another 1C0 miles deep hidden under the
shadows of the vast array of snow-capped mountains of -Baltistan
until they arrive a few miles below Gilgit. Here the great change
of direction to the south-west occurs, which is thereafter maintained
till the Indus reaches the ocean.
The Indus of to-day isthe Indus of myth and legend but swept
clean of mystery and the hidden strongholds of the fanatics along
its banks—a’ hindrance from all time to the progress of this world.
Many important points are bridged by the suspension system,
though there still exists the ‘Cantilever’ and the rope-bridge which
sways uneasily over the roaring waters.
We dipped into Puga in Rupshu and continued up the sandy
vailey to the shores of the Polokonka Lake (14,000’), where we
found the Great Crested Grebe (Podiceps cristatus cristatus) breeding,
as well as a large colony of Brown-headed Gulls in an advanced
stage of genital development.
The eggs of the Crested Grebe are placed on a floating mass of
reeds and the eggs, 4 to 6, in number appear to take on the colour
of the decaying matter in the nest.
Several lakes in the east districts at 14,000 feet above sea level
have been of much greater extent and connected to the drainage
system ofthe country, but they are now mostly without outlet,
saline, and in process of desiccation.
The highest visible water mark of the Polokoncha Lake is quite
150 feet above its present level, and recent years seem to show a
much more rapid process towards drying.
We shifted camp to the northern end of the lake and pitched our
tents on the lawn-like banks of a mountain torrent, while all around
the Poppies and Primulas grew in wild profusion.
There was comparatively little of bird or animal life. Gulden-
stadt’s Redstart (Phenicurus erythrogastra grandis) seemed in sole
possession of the stream. The surrounding hills offered nothing
but bareness and desolation.
Small parties of Wild Asses, or Kiang'ss as they are more commonly
known, visited the shores of the lake in the early mornings and late
evenings, the dead stillness being occasionally broken by their
‘shrieking bray.’ There seems a good bit of discrepancy between
the accounts of the Kiang’s cry given by different sportsmen. We
could safely leave it as intermediate between the neigh of a horse
and the bray of an ass—-almost alike to the vocal efforts of a mule,
which at its very best sounds like nothing on earth.
Whether in the rarified atmosphere the Kiang 1s as fleet of foot
as the species from Sind, it is difficult to say. The former can
‘move’ on occasions taking 18,000 and 19,000 feet hills in grand
style.
Besides its dreariness, the shores of the Polokoncha hold for the
Journ., Bombay Nat. Hist. Soc.
Our Camp AT 15,500 FT. IN THE
KARAKORAM
me
bem ne
”
kore
A COLLECTING TRIP TO LADAK 653
traveller a certain sadness. It was here that the wife of a young
Englishman met a tragic end, by inadvertently lighting a fire in her
tent with the very obvious result, when the very first rule in camp-
ing is broken.
~ On the 28th June we left the Polokoncha Lake and crossing the
Taklung Pass (17,600 feet) which rises sheer from the shores of the
lake, descended rather abruptly into the Gya Gorge.
While on the Taklung Pass the writer brought off a record bag of
Snow-cock ( 7etraogallus tibetanus) obtaining six birds in a single shot!
The possibility of such a feat may be enhanced when it is known
that two adult birds had their heads together, while four young
squatted on the mother bird’s back.
Following the Gya Gorge, which with rare determination pierces
an arm of the I.adak range, we swing off to right angles and enter
once again the Indus Valley which still keeps its boulder-strewn
nature. You wonder if the track lost itself or from fright and loneli-
ness ran up a frightfully untidy glen to Hemis, or that Hemis
huddled itself among the bare rocky crags in trying to escape from
the encroachments of ‘ western civilization’, for even here the
missionary offers a very very old dog a new collar, expecting him
to do new tricks.
We wind up the frightfully untidy glen into the heart of the
mountains, where amid forbidding rocky hills, Hemis, the Mecca
of one of the noblest of religious faiths and simple philosophies
yet unfolded by man, stands gaunt and silent.
True to this land of contradictions and surprises, a delightful
garden of stately poplars and wild roses, whose delicate perfume
scented the balmy air, was cradled among the barren hills and set
like a first-water jewel in a cheaptrinket. A multi-coloured crowd
of Ladakis thronged the massive gates that led into the monastery ;
while taking his joys less seriously, that hardy son of desolate
wind-swept steppes, the Tibetan, could easily be picked out by his
easy swinging walk and his fine example of full-blooded manhocd.
From the babble of voices it seemed that all Ladak and his wife
had turned up to witness the annual Devil-dancing and Festival
held at Hemis. And so the sun set on this strange scene, casting
grotesque figures among the crags, until night shrouded the narrow
glen in a sombre mantle, the darkness punctuated with innumerable
camp fires.
It might have been a scene from the Arabian Nights, if such a
scene did not date back to 500 B.c., for Hemis is almost as old as
the hills, even older some say.
The religious displays or ‘ Devil-dances ’ enacted annually among
the Bhuddists are familiar sights anywhere between Ladak and
Bhutan.
Much has been written on the various conceptions of this mystic
religion. Mystic, perhaps from the theologist point of view but to
the weary traveller it appears a bewildering agglomeration of
superstitions pieced together into an amazing farce, in which the
‘ Sons of Belial have a glorious time’.
Still one is really impressed by the rhythmic dancing, the
gorgeous dresses and the grotesque masks worn by the dancers,
654 JOURNAL, BOMBAY NATURAL AIST, SOCIETY, Vol. XXX/1
which vie with each other in diabolic hideousness. Then, too, there
is tae perfect symphony produced by the treble and bass trumpets.
The latter are ten feet in length and emit a volume of sound not
unlike the amplified drone of a giant beetle in flight.
We left Hemis and arrived at Leh, by two easy stages on
July 5th. As yet the status of many of the migrants had not been
cleared up and the few days spent at Leh in preparations for our
journey to the Karakoram were helpful in clearing up one or two
problems. The Blue-throats (Luscinza svecica ssp.) which seem
fairly well distributed along the Indus and Shyok Rivers, wherever
suitable country is found, have for a considerable time confused
naturalists by their variable colourings on the throat.
Thanks to Col. Meinertzhagen’s systematic methods of collecting,
we obtained a remarkable fine series ot the Blue-throat which
breeds in Ladak. This has been identified as Luscinia svecica
abbottz, and is characterized by avariable colouring of red, some-
times reddish-white and white stars. Among our specimens of
Luscinta suecica abbotti there were five males and one female with
pure white stars, four males with pure red stars and four males and
two females a mixed red and white.
A red ‘starred’ male would have a white ‘ starred ’ female or
vice versa. ‘The type is a white-starred bird, and is known as the
‘ White-spotted Blue-throat.’ ;
Next we have Luscinia suecica pallidogularis (the Red-spotted
Blue-throat) which oftener than not is mistaken for the preceding
bird. Of pallidogularis Col. Meinertzhagen says, ‘ Whereas in
abbotti the “‘ star’’ is very variable, in this race it is always pure red,
sometimes a few white feathers are apparent. Underparts paler
than in addot¢ti and with seldom much black at the lower end of the
blue chest. Blue much paler. This is the only Blue-throat met
with in Kashmir in spring. First arrivals (all males) were noted on
March 23. This race does not breed (nor does any Blue-throat)
inthe Vale of Kashmir, though in Northern Kashmir, on the Gilgit
Road at 11,300 feet I found two pairs feeding young. The adults
are typical of this race.’
Thus we have Luscinia (Cyanosylvia) suecica abbotti breeding
throughout the Indus and Nubra Valleys—but not in Baltistan ; and
L.s. pallidogularis breeding in Baltistan (Western ‘Turkestan to
East Trans-caspia—Hartert).
These observations are clearenough and backed with specimens,
and should eliminate the difficulty in identifying the Blue-threats.
With the advent of spring the dreariness of winter seems never to
have existed. It is a country of sunshine and rivers and rich
fertile valleys ; and far away the great mountains hang inthe serene
sky, peaks, crests, and ridges of inaccessible and eternal snow.
The little hamlets in the sweep of the Indus Valley peep through
a smothering foliage of poplar and willow; while the cheerful
chatter of the House Sparrows adds liveliness to the drowsy
air.
It is surprising that the Jackdaw (Corvus monedula semmeringiz),
so abundant in the Vale of Kashmir, should not find its way up here
(Leh). Beyond a couple seen in Leh in May we never came across
A COLLECTING TRIP TO: LADAK 655
these birds anywhere in Ladak. Surely there is some mistake in
Baker’s statement ! that these birds breed in Ladak !
Nor does the Jungle Crow (Corvus coronotdes intermedius) extend
for any great distance into Ladak, though this bird has been recorded
as breeding in Ladak. What may have been mistaken for the
preceding bird is the Carrion Crow (Corvus corone orientalis), which
we found quite plentiful at Kargil in April and took several clutches
of eggs from nests in willow trees at Moulbeck (11,000 ft).
Elsewhere I have mentioned the House Crow (C. s. zugmeyerz) as
occurring in the Vale of Kashmir, and then only being isolated to
a few colonies, which Osmaston observes do not leave the valley
but brave the winter. Perhaps it may yet be discovered that the
Sind House Crow and the one in Kashmir may not belong to the
same race.
If there is a paradox among birds it is the scavenger of
Ladak—the Tibetan Raven (Corvus corax t7befanus). To see him
studiously peer into the bowels of a dead horse or walk gingerly in
the refuse of a village is to heartily dislike the fellow ; but to see
him at ‘ play’ is truly amazing. They wanton about in a thousand
different manners and postures, sometimes pursuing each other,
and making a mock fight; sometimes rising; sometimes falling
with closed wings; sometimes floating awhile in the air on their
backs; sometimes flying edgewise; sometimes whirling round
the monastery with vast rapidity; and all with an eagerness
and joyousness of motions and cries and screams that shcwed
the overflowings of happy sensations too plainly to be mistaken.
Besides these attractions, C.c. fébefanus has a‘most peculiar and
extraordinary metallic call wondrously like the tones of bells which
emitted from ahill-top has the resemblance to ‘‘ vesper belis’”’
heard at a distance.’
He also possesses a sense of humour. Observe his comical
imitations of an eagle missing its prey. He willsit calmly by and
‘ watch the fruitless efforts of an eagle or buzzard, and then with
twirls and turns and screams will show you how it shouldn’t be
done. But wedigress. Our preparations are almost complete for
the trip to the Karakoram. A slight change has occurred in our
dramatis persone. The shikari’s abhorrence for the cook and
his ancestors finds a head and he gives vent to his pent-up feelings.
Feelings which, he confidesin me, be has fostered ever since he
clapped eyes on the son of Sheitan. And, like all true Muslims
who at least a dozen times a day ‘seek refuge in Allah from
Sheitan the stoned’, he had no alternative but to resign.
Unfortunately this ‘ dafailleroyale’ took place on a plateau at
16,600 feet, where the only Tibetan Grouse (Syrrhaptes izbelanus) we
had seen, was secured. Ourskinning after the day’s march prevent-
ed either of us from a thorough search for more specimens. The
incident will prove the effects of high altitude on the equanimity of
the mind, of even those of little orno imagination.
The vacancy was soon filled up by a Yarkandi—a fellow who could
go over a pass without a sob and be game for some more! His is
2S, voli, pare 3G;
656 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
the type one meets along a line drawn from Yarkand, through
Afghanistan and Baluchistan, along the highlands of Persia, into
Kurdistan and soto Turkey; and like the soldier of that cuuntry,
he will fight or freeze, march and starve not only without a mutiny
but even without a sense of grievance against anybody. He is a
gentleman, and anyone acquainted with Muslim countries knows
that a Moslem gentieman is unsurpassed in any country. .
We left Leh on July 12, and camped that evening just below
the Kardong Pass at 15,000 feet. Next morning a diligent search
was made tor that beautiful Rose-finch (Pyrrospiza punicea humzz)
of which Col. Meinertzhagen secured one the evening we arrived.
I had marched up a side valley to the foot of a glacier and drawna
blank and then up a rugged hill to about 16,000 feet, where, pausing
from the effects of an oppressive atmosphere I looked back to where
Leh lay basking in the noonday sur, and then for no apparent reason
an unmitigated siab of dreariness came over me and I wished Leh,
Ladak, the snowy mountains and all naturalists in the nether
regions. High altitude again! If such feelings are allowed to run
riot anything may happen; but I had an employer who understood
better than I then thought and shall always feel grateful to him for
his tolerant attitude towards my misgivings. It is but human nature
that we consider our own experiences in travel and adventure as
unique (with special stress on the hardships), and completely ignore
the fact that others have been before us and others will follow.
If you have crossed the Great Himalayas in winter and wandered
through barren Ladak ; if you have crossed the Changla Pass at
18,500 feet and found another pass still higher, and your goalisa
desolate region which is called Chang-chen-mo ;—if these trifling
hardships damp your enthusiasm there is a quiet garden in Leh
among the tall silent poplars, with the wild roses rambling over
the swelling sod, that marks the trend of those pioneers who
have ‘ gone before ;’ it will make you ‘ gird up your loins’ for yet
another try.
Here among the lone graves, an obelisk israised by a thankful
Government as a small mark of its appreciation to the memory of
Ferdinand Stoliczka, that brilliant young scientist who gave of his
best in that bloodless field of Natural History.
‘ Ferdinand Stoliczka,’ so the inscriptionruns, ‘the eminent young
Geologist and Natural Historian accompanied the second Yar-
kand Mission in 1873 under Sir Douglas Forsyth. His health,
severely affected by his previous work, gave way under the strain
of strenuous work, he died at Shyok, in June 1874, ‘ while returning
loaded with the spoils and notes of nearly a year’s research in the
least known parts of Central Asia.’
‘The scientific results of this research,’ writes Allen Hume,
‘constitute an imperfect record of one single year’s work of our dear
friend and comrade Ferdinand Stoliczka. Imperfect,—because we
had only his collections anda very brief diary—a mere azde memoire
to work upon and so are able to present only the dry bones, as it
were, of the results of his labours ; which had he survived to deal
with them himself, would have risen up a living entity, vivified by
his genius. 7 i 3 oar
Imperfect,—but yet the best memorial that, under the circum-
BOMBAY NAT. HIST. SOc
JOURN.
Leh
zka’s ‘Tomb,
Cc
Stoli
A COLLECTING TRIP TO 'LADAK 657
stances, we survivors could raise to him, untimely hidden in that
lone Tibetan grave.
Imperfect,—but for all that, affording every here and there
glimpses of the perseverance and self-devotion which, informed
by wide scientific knowledge, and sanctified by a pure and
generous heart, constituted Stoliczka’s irresistible claims to respect
the admiration and the love of ali who ever came closely in contact
with hii.’
To those who are acquainted with the Palaearctic, that cruel relent-
less region, Stoliczka’s name shall go down as one of those heroes
whose bloodless blameless triumphs remain for ever the inheritance
and the blessing of mankind.
The road after crossing the Kardong Pass at 17,600 feet descends
abruptly into the sandy valley of the Nubra River. So abrupt was
the snow-covered descent that we expressed a wish we would never
have to assault this particular pass again. Fruitless wish!
The Nubra Valley, unlike the Indus, is fairly well cultivated—its
entire length being more orless planted with large gardens and the
gaps filled up with tamarisk bushes.
Panamik, 10,400 feet is the last village below the Karakoram.
From here the road turns into the Nubra Gorge and follows the roar-
ing river along its precipitous banks till an altitude of 13,000 feet is
reached and then takes one onto a meadow, surrounded by stupen-
dous mountains, with gigantic glaciers between the peaks.
The Karakoram Range forms the great water divide which
separates the Indus drainage from that of the Yarkand and other
Chinese-Tibetan rivers.
It is buttressed by K? (Godwin-Austen) 28,650 feet at its conver-
gence with the undulating elevated plateau of Yarkand. The
accompanying photograph gives some idea of the mountainous
regions and two enormous glaciers are clearly seen on either side
of the conical hill.
The pass over this range is crossed over a ten-mile glacier at
19,000 feet and lets one into Yarkand.
The snow had melted, only the very highest peaks retained their
winter mantle, and the glaciers sparkled in the bright sunshine, and
as the ice thaws huge boulders clatter down. ‘The pass itself winds
up the debris brought down by the melting of snow and ice, and, taken
all round is the most savage bit of primeval chaos in all the
mountain ranges we encountered.
Perhaps our protracted stay at high altitudes had weakened us
physically or perhaps this was our Pisgah, but after a‘ trial’ up the
slippery moraine, Col. Meinertzhagen decided to turn back and
we began the return journey as eagerly as horses heading for
Stables.
It was Col. Meinertzhagen’s intention to work the Shyok Valley
up to its junction with the Indus and crossing over thence to reach
Skardu. But this project had to be abandoned owing to the floods
in some of the rivers.
We wili not weary the reader with a description of the return
journey to Leh.
Let it suffice that as we confronted the steep ascent to the
6
658 JOURNAL, BOMBAY NATORAL, HIST. SOCIZATY, Volz XOOd7
Kardong Pass, nothing in the vocabulary could express my feelings.
It was the first pass wherein I lay hold of a Yak’s tail as a means
of locomotion and was hauled, much against my wil], at a pace
which caused alarming palpitations.
Skardu, which will be found upon the map, is the capital of
Baltistan (or Little Tibet as itis called), and rests amidst a fine array
of rugged mountains with the Indus sweeping past in a great horse-
shoe bend. We arrived here on August 13 after a hot trip
through the Indus Gorge, to find Skardu among one of the dirtiest
towns in the Himalayas. ‘The inhabitants, originally Bhuddists, are
now extremely bad imitations of Moslems, and like all proselytes are
heartily disliked by both sides. However, Baltistan amply repays
a visit. Its bird-life will settle many a knotty problem, being a
sort of half-way house between Kashmir and the desolate regions
that lie beyond Baltistan.
Most of the migrants were undergoing their autumn moult prior
to setting forth on their winter passage, during our visit.
Conspicuous among these was that familiar little fellow, the
Kashmir Red-start (Phenicurus ochrurus phenicurotdes), who seem-
ed to be fairly weii distributed throughout Ladak also. The
Eastern Red-spotted Bluethroat (Cyanosylvia suecica pallidogularis)
were busy feeding young, as were the Pied Chats (Cenanthe
picata).
The Hoopoes (Upupa epops) were in fresh plumage and ready to
take off. These are distributed throughout Ladak and are found
always near villages. The regularity of their arrival from the
plains to their breeding haunts in Ladak has fixed the date for the
commencement of spring ploughing in that country.
Cholera had been raging on Skardu on our arrival and we were
gladto geta moveon. On August 19 we left Skardu for the Deosai
Plateau by way of the Satpur Lake, which is cradled between two
rugged hills, quite 5,000 feet above Skardu, and worried our way up
to 11,500 feet when we sat down for tiffin.
Up to this stage Col. Meinertzhagen had shown no sign of the
catastrophe which befell him. We had even secured a new sub-
species of the Twite (Acanthis flavirostris baltistanicus), as distinct
from Acanthis tlavirostris ladacensts, which is also a new sub-species.
That same afternoon, while we camped at about 13,000 feet, Col.
Meinertzhagen had an attack of cholera and to add to this misfor-
tune a snow storm swept down the hill.
Next morning we crossed over at 14,000 feet and the Deosai
Plateau stretched away from our feet to the lilac hills, over an even
grassy meadow, strewn with flowers. It was a relief being able
to see such an expanse of open country after months of confined
valleys and dreadful gorges. Toseethe sun set without intervening
mountains or the golden orb of the moon rise over the rim of the
world.
Even though the cholera still gripped him, Col. Meinertzhagen
kept the same cheerfulness and keen interest he had always
shown.
The plateau for the most part was well stocked with higher alti-
tude birds. The Horned Larks were abundant and Col. Meinertzhagen
A COLLECTING TRIP TO LADAK 659
has separated these from Olocorzs alpestris longtrostris and named
them Octocris (Evemophila) alpestris deosat. In the /bzs (1927, p. 402)
he says: ‘A series of nine birds were obtained on the Deosai
Plateau at the end of August. They are mostly infull moult, only
three birds having completed their wing moult, (wings of three
127-131, culmens of six males 18-20 m.)
These birds run as large as /ongirostis but are both darker and
browner in fresh plumage. ‘They have the white gap between the
black of the throat and that of the cheeks.
Apparently confined to the Deosai plateau between 12,000 and
14,000 feet.’
The Robin Hedge-Sparrow (Prunella rubeculotdes) which was
commonly met with between 12,000 and 14,000 feet over the greater
part of Ladak was abundant on the plateau, and several nests were
seen with fresh eggs on July 8. Col. Meinertzhagen has sepa-
rated this form under P. ~. muraria. We crossed the plateau
in three inarching days; owing to Col. Meinertzhagen’s condition
matters had to be rushed a bit and on August 27 wedescended to
Burzil Chowki on the Gilgit Road. We put in four days here and
with Col. Meinertzhagen on his feet again, commenced the final
stages to the Vale of Kashmir.
It was a late autumn evening when we crossed the Razdhanangan
Pass and looked down on a sea of gardens and fields, and, sparkling
through the gathering mists, the gem-like lakes that make the Vale
of Kashmir ‘ a paradise on earth. ’
(Zhe end)
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA
BY
R; 1. POcocKF.-R: Ss;
Part. 3h
( With two plates and three text-figures.)
(Continued from page 504 of Vol. XXXT11)
II, The PyRRHUS-group.?
Distinguishable from the Entellus-group by tke uniform golden-red hue
which in the newly-born young pervades the whole of the body, limbs and tail
above and below. In cranial characters the species more resemble Pithecus
senex than Pithecus entellus.
Distribution. From Assam and Upper Burma southwards through Laos,
Siam and the Malay Peninsula to Sumatra, Java and Borneo.
The actual number of species is doubtful; but within the area of British
India I admit only two, which may be briefly contrasted as follows :—
(a) Alittle larger; hairon the crown directed backwards
from the brow and forming athick mat overhang-
ing the occiput and marked off from the short hair
clothing the cheeks; face entirely black Le
(at) Smaller; hair on crown variable but never forming
a mat as described above; eye-lids and usually the
lips flesh-tinted (in British Indian forms) .. pyrrhus,
pileatus.
Pithecus pileatus, Blyth.
Semnopithecus pileatus, Blyth., Journ. As. Soc. Bengal, xii, p. 174, 1843,
and xiii, p. 467, 1844.
Semnopithecus, Presbytis, Pygathkrix or Pithecus pileatus of Anderson,
Blanford, Forbes, Elliot and other authors.
Atolerably large langur characterized by the presence on the crown of the
head of a thick continuous mat of backwardly directed or nearly erect hairs
extending from the brow in front to the occiput behind and across the head
between the temples and ears. Laterally this matis defined by the shorter
hair clothing the temple behind the eye and by the more or less, often greatly,
elongated hairs of the cheek, and behind it overlaps the shorter hairs of the fore
part of the nape.
The colour is variable but the prevailing tint of the upper side is grey, some-
times suffused more or less with black, especially on the crown ; the lower side
is whitish, buff orsuffused with orange red, rarely grey like the upper side.
The face is entirely black as in the Entellus-group, neither the eyelids nor the
lips showing any trace of the flesh colour seen in all the British Indian repre-
sentatives of the following species, P. pyrrhis.
Distribution. Assam, north and south of the Brahmaputra and Upper
Chindwin to Tipperah, and Chittagong.
The evidence that this species belongs to the group with golden red newly
born young was suppiied by the birth of young of that colour from a pair kept
for many years in the Zoological Society’s Gardens, London.
Elliot classified P. pzleatus with P. entellus, stating the two to be allied.
‘They are certainly not closely allied. As living animals they havea very
different facial expression. P. evfellus has a narrow compressed nose, whereas
1 In the first part of this paper (Journ. Bomb. Nat. Hist. Soc. XXXII, p. 474),
this group was called the Auratus-group. But since the identity of Pzthecus
auratus in a matter of dispute, I think it advisableto change the name to the
Pyrrhus-group.
TEE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 661
in P. pileatus the nose is flat from side to side. The difference may be aptly
compared to the difference between a European and a Negro.
This is borne out by the shape of the facial portion of the skulls. In
P. pileatus the brow ridge is not prominent, the plane of the orbit is inclined
upwards and backwards and the nasal bones are wide. In those respects the
skull much more closely resembles the skull of P. sénex than the skull of
P. entellus.
In his paper, quoted below, on the langurs of this group, Hinton suggested
that the different forms, some of which were regarded as distinct species, would
very likely prove to be merely subspecies of a single species. Believing this
view to be quite correct, I have adopted it in this paper.
Key TO THE SUBSPECIES OF P. pileatus
(2) Whiskers shorter, grey, not noticeably lighter in tint
than the crown ... “se wes .. Shortridget,
(a1) Whiskers longer, whitish or reddish and rather
sharply contrasted with the darker tint of the
crown.
(6) Nosharp contrast in colour between the upper and
lower sides of the body and the outer and inner
surfaces of the limbs; throat and breast whiter
than the belly .. wee) «Ovahma.
(6+) Asharp contrast between the areas mentioned
above ; throat and breast not paler than the
belly. ve
(c) Lower side greyish white, tinged with buff
on whiskers, throat and breast . ptleatus.
(c1) At least the whiskers, throat and breast
bright rusty red.
(d) General colour paler above, head and
nape nearly uniform slaty grey, red
suffusing underside at least to the
groins .. durga
(d+) General colour darker above,
head blackish with grey patch
on nape; red of the under side
fading away on the abdomen. ... /enebricus.
Subspecies p2leatus, Blyth.
Semnopithecus pileatus, Blyth, Journ. As. Soc., Bengal, xii, p. 174, 1843.
Pithecus pileatus pileatus, Hinton, Journ. Bomb. Nat. Hist. Soe , xxix, No. 1,
pp. 79-80, 1923.
Semnopithecus argentatus Worsfield, Cat. Manm. East Ind. Co., p. 7, 1851.
Distinguished by the pale colour of the whiskers, throat, underside of the
body and inner side of the limbs which are sharply contrasted with the grey
hue of the upper parts but are white or whitish and at most faintly tinged with
buff or pale red.
Distribution. Assam from the Naga to the Garo Hills.
It is needless to repeat the history of this local race of Pzthecus pileatus which
was fully told by Hinton in 1923 when, on the evidence supplied by Blyth’s
original description based upon a langur of which the locality was unknown, he
restricted the subspecific title pzleatus to the Capped Langurs with pale under
parts.
The colour, however, varies in different individuals assigned to this race. For
example, a half- -grown female shot by H. W. Wells at Mokokchung in the
Naga Hills (5,000 feet) is pale slaty or smoky grey above and on tlhe outside of
the limbs and on the tail, with the head and hands a little darker. The
whiskers, the whole of the lower side and the inside of the limbs are white and
the white extends some distance up the flanks behind the arm-pit. On the hand
the white of the inner side passes above the thumb and the fingers are patched
with white. The feet are not darker above than the outside of the leg, and the
inner edge of the foot and the toes are white. This specimen appears to be
662 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
whiter below than Blyth’s type of pz/eatus, the under side of which he described
as ‘dull fulvous white’. This was also an immature female.
An adult male from Mokokchung differs from the female in having the areas
which in the female are white, tinged with butf or very pale red ; the hands and
feet are darker and there is no white on the fingers or toes, but the extension of
the white of the inner side of the leg along the margin of the foot above the
great toe is retained.
An adult male from Tura in the Garo Hills, 1,400 feet (H.W. Wells)
generally resembles the last-described but is paler on the legs and arms, the
outside of the legs below the knees and the top of the feet being much greyer.
—
ME See
—— Gg
a
py gS)
Fic. 1—A and B. Front and side views of skull of adult male Pithecus
pileatus pileatus from Tura in the Garo Hills.
An immature male from Lait Kynsao in the Kasi Hills, 2,000 feet (H. W.
Wells) is very like the adult male from Tura; and an immature male from
Konshong in the Jaintia Hills, 3,000 feet, has the outer side of the legs below the
JOURN. BomBay Nat. Hist. Soc. Puate III.
THE CAPPED LANGUR AND YOUNG
(Pithecus pileatus)
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA _ 663
knee greyer than in the older specimen from Lait Kynsao, and, as Hinton
pointed out, the terminal portion of the tail is buff instead of blackish.
The redder hue of the under side of these young males from the Khasi and
Jaintia Hills distinguishes them markedly from the immature but older female
from the Naga Hills. They serve toconnect the typical form of pzleatus with
the intensely red-bellied form recorded below as Pithecus pileatus durga. So
also in a measure does the type of argenfatus so faras can be judged froma
specimen in a very poor state of preservation. This langur, an immature
specimen, came from Sythet, and the hair that remains on its under side sug-
gests that, when freshly killed, the monkey was redder on the chest than are
the adult males from the Garo and Naga Hills described above. It is also
browner on the dorsal area. Since, nevertheless, the skin is not nearly so red
beneath as in the typical red-bellied Capped Langur described below, I follow
Wroughton in assigning argentatus to the synonymy of pzleatws proper, at all
events until fresh material of this langur from Sylhet shows that opinion to be
wrong, It may be proved that the type of argentatus is an immature specimen
of the red-bellied form which has lost its redness by fading.
The photograph of the Capped Langur Pithecus pileatus reproduced on PI.
III of this paper was taken from a specimen from Assam living in the Zoological
Gaardens, London.
‘Subspecies durga, Wroughton.
Presbytis pileatus, Blyth, Journ. As. Soc., Bengal, xvi, p. 735, pl. xxvi, fig. 3,
1847.
Presbytis durga, Wroughton, Journ. Bomb. Nat. Hist. Soc., xxiv, No. 4,
Da Gs55).1916.
Pithecus durga, Hinton, Journ. Bomb. Nat. Hist. Soc., xxix, No.1, p. 79,
1923.
Pithecus pileatus saturatus, Hinton, fom. cit., p. 81.
Distinguished from P. p. pileatus by the intensity of the redness of the areas
which in the typical form are at most faintly tinged with that colour.
Distribution. Assam, north of the Brahmaputra from Lakhimpur to the
Naga Hills and Cachar ; thence southwards into Tipperah and Chittagong.
From the evidence supplied by collectors it appears that this erythristic form
occurs at lower levels than the typical form with the paler under side. For
example, in the Naga Hills typical pzleatus was taken at Mokokchung at an
altitude of 5,000 feet and durga at Lakhuni at 2,000 feet.
In his tabular classification of the langurs of the pz/eatus group, Hinton,
following Wroughton, described the colour of the whiskers and under surface
of P. durgaas ‘ ochraceous’. ‘They are, on the contrary, markedly red, as red
to all intents and purposes as in the form henamed FP. pzleatus saturatus. The
general colour of the upper side of both forms he described as dusky brown.
As a matter of fact the type of durga is somewhat browner and dirtier looking
than the comparatively recently killed langurs referred to saturatus ; but ic
must be remembered it is a specimen that belonged to Dr. Reid and has been
many years in the British Museum and the difference may I think, be assigned
to that cause or to discoloration due to the original method of preservations,
such as that which gave rise to the difference between the skins of typical
Pithecus pileatus shortridget and those that Wroughton distinguished a
P. shortridgei belliger. Also, I think, that soiling, orthe make up, of the
skin of durga explains the apparent difference Hinton noticed between it and
saturatus in the width of the dusky brown dorsal area and the less sharp con-
trast when the skin is viewed from above, between it and the red hue of the
flanks. The teeth of the type of durga are, itis true, a little larger than in
saturatus ; but I feel tolerably sure that this will break down as a difference
when more specimens come to hand.
Now the label on the type of durga bears the locality Upper Assam, Cachar.
No doubt, however, as Wroughton decided, ‘Upper’ was a mistake for
‘Lower’. Taking, then the locality as probably Cachar, it is significant in
view of the claimed identity between durga and saturatus that examples
assigned to saturatus by Hinton were collected by the Mammal Survey at
Lanka and Lamsahkang in North Cachar. It does not seem probable that two
oo of P. pileatus, both intensely red below, occur in that district of
\ssam. :
664 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol, XXXII
Although I have not seen examples of this form from Tipperah and Chitta-
gong, I have added those countries to its distributional area because Blyth’s
description and coloured figure of specimens therein collected agree in all
essentials, as Hinton also thought, with those to which he gave the name
_—saturatus.
Blyth regarded these red langurs as identical with his typical pzleatus of
which he had previously seen a single specimen which was whitish below and
came from an unknown locality;and from the material in his hands he
inferred that the redness of the under side was characteristic of the male. In
this he was mistaken because in a pair collected by H. W. Wells at Golaghat
at 400 feet the male is not redder than the female.
This pair is also interesting in other ways. The male, although not redder
than the female, is darker grey above and onthe outside of the limbs and
blacker on the hands, feet and tail, the feet being quite black, whereasin the
female they are grey. Moreover, the female, apart from being brighter red,
especially upon the sides of the neck and upper part of the flanks, is very like
the adult male from Turain the Garo Hills, 1,400 feet, which has been re-
ferred above to the typical form pzleatus pileatus. Nevertheless, it is not so
deep ared as an adult male from Lanka, 400 feet, in North Cachar. It may
indeed, be described as intermediate between the male pz/eatus from Tura
and the male durga (saturatus) from Lanka.
Individual variation in redness is attested by other specimens. For instance
a female from Lamsahkang, 800 feet, in the Cachar Hills, has the red more
diffused than in the male from the same spot. It invades the outer side of the
leg replacing the normal grey of the limb which is retained in the male.
Subspecies fenebvicus, Hinton.
Pithecus ptleatus tenebricus, Winton, Journ. Bomb. Nat. Hist. Soc., xxix,
No, 1) peal, 1923.
Distinguishable from the two preceding local races by the darker tint of the
upper surface. The head, back, outer surface of the arm and hand, and the
base of the tail are ashy black; the leg is somewhat greyer and there isa _ half-
collar of grey tipped hairs defining the cap on the back of the head. The base
of the whiskers, the throat, chest and shoulders are almost as deep a red as in
P. pileatus durga, but the whiskers are especially grey as in P. pileatus brahma
and it is noticeable that the redness of the under side is much less pronounced
on the abdomen and inguinal region, which in the male are whitish and in the
female pale red.
A young specimen is darkisn grey above and on the outside of the /limbs, but
is whitish below with a yellow tinge on the cheeks and chest. It is darker
above than a young male of P. pzleatus pileatus of about the same age from
Konshnong in the Jaintia Hills.
Distribution. Assam, north of the Brahmaputra; North Kamrup on the
Matunga River, 1,200 feet and Bogra Nadi, 2,000-3,000 feet.
Subspecies 6vahma, Wroughton.
Presbytis brahma, Wroughton, Journ. Bomb. Nat. Hist. Soc., xxiv, No. 4,
p. 654, 1916 and xxv, No. 4, p. 559, 1918.
Pithecus brahma, Hinton, Journ. Bomb. Nat. Hist. Soc., xxix, No.1, p. 79, 1923.
Tolerably uniformly slate-grey on the summit of the head, the back and the
outside of the limbs and on the tail, but the limbs and tail darkening towards
their extremities. The inside of the limbs paler grey but not sharply distin-
guished from the tint of the outer surface, with no pale line, involving the
hallux on the inner side of the foot. Whiskers white and sharply contrasted
close tothe face as in the typical fcrm of the species, but ashy at the tips as in
tenebricus. The throat and chest whitish ; abdomen the same but tinged with
reddish buff especially on the flanks.
Distriéution. Seajuliin the Dafla Hills, Lakhimpur, Assam, north of the
Brahmaputra.
A single example only of this fine langur is known. Although regarded by
Wroughton and Hinton as representing a distinct species, its characters are, in
my opinion, of subspecific value ; and I do not doubt that intermediates between
it and the other races will be discovered.
‘THE LANGURS, OR LEAF MONKEYS, OF BRITISH [NDIA 665
The peculiarities in colour to be noted are the absence of sharp contrast in
tint between the outer and inner surfaces of the limbs such as is seen in pz/eatus,
durga and éenebricus, and the slight suffusion of the flanks and abdomen with
pale red, the throat, whiskers and chest being white. In the other races which
exhibit more or less pronounced erythrism, that tint is, at all events as a rule,
more pronounced upon the whiskers, throat and chest than on the abdomen,
- Subspecies shoriridget, Wroughton. ese
Presbytis shortridgei, Wroughton, Journ. Bomb. Nat. Hist, Soc., xxiv, No. 1,,
pz 06, 1915:
P: shortridgei belliger, id. loc. cit., p. 57.
Pithecus shortridgei, Hinton, Journ. Bomb. Nat. Hist. Soc., xxix, No.1, p. 79,
1923.
General colour very much as in bvahma, uniformly slate-grey all over the
dorsal surface and on the head, but the ventral surface grey and the whiskers
wholly grey and not appreciably differentiated in tint from thecrown. No trace
of erythrism anywhere in the pelage.
Skull as in the other races of P. pzleatus.
Distribution. Upper Chindwin: Homalin (400 feet), Minrin (450 feet) and
Hkamti (500 feet). .
This fine langur is at once distinguishable from the other known forms of the
species by the greyness of the whiskers which gives it a very different appearance.
Apart from that, however, and the absence of the rufescence of the ventral
surface, the colour is very like that of P. pzleatus brahma. But in the latter, as
in P. pileatus tenebricus, the tips of the whisker hairs are grey. Although no
intermediates between drahma and shortridgei are known, there is 10 doubt, in
my opinion, that they will be found, because the cclour-differences between
them are comparatively slight. Hence I regard shortridget merely as a local
race of P. pileatus.
The examples from Hkamti to which Wroughton gave the name éelliger are
browner in hue than the others; but this difference, as he afterwards learnt
from the collector, G. C. Shortridge, is due to discoloration from smoke in the
preparation of the skins.
Since the subspecies of Pithecus pileatus, above discussed, are to al] intents
and purposes alike in size, the approximate dimensions in inches of a few
representative examples may be conveniently set forth in one table :—
Locality and Sex oe Tail ae Kar | Weight
| |
pileatus— | |
Garo Hills, ¢ sol 28 41 73 1} ks
Jaintia Hills, 2 ve | 23 Bye ayes | 1} 213 Ibs.
durga—
Lakhimpur, ret Ee 27s 39 = 13 5
N. Cachar, g ae 23 41 a) 13 Z/ libs:
+] ’ coo | 20 eoe ae0 | ae0 -
tenebricus— |
N. Kamrup, ¢ 223 363 | 7 1z 193 Ibs.
3 ae 20 oO te sf 1} ‘a
ae 2 we 225 34 fi 13
5 2) se 18 | 283) 6% li
shortridgei— | |
Upper Chindwin, ¢ oe || 283 41 8—| 13 30 lbs
» ) 3 see 2h 40 8—| 1+. oot
’ om) e) ae 26 38 73 13 213 lbs
Sa ae
666 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
The dimensions of the skulls in milimetres are as follows :—
pileatus—
Naga Hills, ¢.., 118 35 29 75
Garo Hills, ¢... 115| 35] 28; 70] 89
Garo Hills, 2... | 102) 30; 28] 65). 75
|
durga— | |
N.Cachar, ¢ UCM ieee Neer ee faery
Cachar(*), gs] Z| 3 30 | 74} 86 | type of durga.
Lakhimpur, ale oi) 26 76 87 | type of saturatus.
Naga Hills, 9 111 | 35| , 30 | 6s | 79
tenebricus—
IN. Kamrup,; ¢ se) “21S }) 3532 28 66 SO" types. not. tull
| grown.
Peis fs 98 29 28 63 77
brahma— | | | |
Lakhimpur, ¢ . 110; 34 Ze 70
shortridgei — | |
Chindwin, -@ «: ie “OSH b SON SEZ9 91 | type of shortridg.
7 Gt AAS Ge Hale ueSl Fa 85 | .
55 fii) 711d fe SOM 29h NewS 92 | type of delliger.
- @ of 1047], : BBUl 28 Ia Blain agg |
The body-measurements taken from freshly killed animals show that the
males are larger than the females and that there isno great difference in size
between the adults of all the races. An apparent exception to this is supplied
by fenebricus in which an adult female is as large as the larger of the two males;
but examination of the skull of this male, the type, shows that the basioccipital
suture is still open, indicating that growth kad not ceased.
Of drahma unfortunately no measurements in the flesh were taken. The skull
which is that of an adult is decidedly small.
With regard tothe tooth-measurements it will be noted that there is con-
siderable difference between the types of durga and saturatus in that respect.
But the teeth in the type of satwratus are old and worn and have probably lost
a little in total lengtn. Another male belonging to Mr. Hinton’s series of
saturatus is intermediate between the two and a female of the same set from
Lakuniin the Naga Hills has the tooth-row as long as in the type of durga
although the teeth are a little narrower.
Note.—The langur now known as Pithecus potenziani, Bonaparte (= chry-
sogaster, Lichtenstein), which was quoted as a synonym of pzleatus by Ander-
son and was sceptically included by Blanford as a member of the fauna of
British India on the erroneous record ot the occurrence of the species in Tenas-
serim, is in reality an inhabitant of the Mentawei Islands off the west coast
of Sumatra.
It is a very interesting species related, in my opinion, to pzleatus. Its pre-
vailing colour is jet black above, with a considerable amount of rusty red hairs
on the ventral surface. The th: oat and cheeks are white, as in pz/eatus, and a
white band extends from the temples over the brow, as in immature examples
of P. pileatus, and, as in that species, the whiteness of the whiskers spreads
over the base of the ear. The newly-born young too is golden.
One would be almost inclined to regard this langur asa small, insular
melanistic race of pzleaius were it not that the whiskers are shorter and more
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 667
bushy and the crown of the head has no mat-like cap, but a compressed crest or
tuft as in some forms of Pithecus pyrrhus. The species serves in a measure to
link P. pileatus and P. pyrrhus.
Pithecus pyrrhus, Horst.
Semnopithecus pyrrhus, Horsfield, Zool. Res., Java, 1820, and of many subse-
quent authors, including Thomas and Wroughton, Proc. Zool. Soc.
London, 1909, p. 372. (Presbytis).
Semnopithecus or Presbytis maurus, E. Geoffroy, 1812, and of many subse-
queat autaors ; but not Sz#za maura of Schreber.
Resembling PP. pzleatus in the uniform golden or golden red colour of the
newly-born young but differing in being smaller and in the mode of growth of
the hair on the crown which never forms a mat or cap as described under that
species. Its arrangement is very variable. Occasionally there is a whorl on
the forehead and not infrequently there is an erect tuft or crest on the summit
of the head to which a good deal of importance has been attached, althoughin
certain cases there seems no doubt that its occurrence is individually variable.
Sometimes, again, the hair lies smoothly back from the brow to the nape of the
neck, at others it is upstanding and roughly arranged likeamane. The
whiskers are typically long and blend with the hairs of the crown. The colour
varies from subspecies to subspecies, but is ia some cases subject to individual
variation to a pronounced degree, but the inside of the thighs never shows the
whiteness so characteristic of the species of the following group, which Pyrrhus
overlaps in range.
Distribution. From Upper Burma southwards and eastwards into Annam,
Siam, the Malay Peninsula, the Sunda Islands and Borneo.
For the typical Javan form of this species Elliot (Rev. Primates, iii,
p. 75, 1913) adopted the name auratus Geoffrvy (1812), which has priority over
pyrrhus. But his statement that the type of auratus, which he saw in Paris,
is indistinguishable from examples of pyrrhus in the British Museum, is shown
by the description of awraius to be untrue. And since, in my opinion, all the
available evidence suggests that auratus belongs to the Aygula-group of
Langurs, I do not propose to discuss it further here,
In Java P. pyrrhus is commonly either red or jet black, being addicted
to erythrism or melanism. Buta third variety is blackish, grizzled to a varying
extent with buff or grey; and this variety passes into the Sumatran form
known as ¢ristatus or pruinosus which also occurs in the Malay Peninsula and
isin my opinion only subspecifically distinguishable froma form from Siam
and Cochiu China named by M. Edwardes germant but usually quoted as
germaint ; and througha form from Annam, named margharita by Elliot,
germani passes into the typically Siamese Langur assigned in this paper to
crepusculus which is closely akin to the Burmese phayrez. On the other hand,
in the Malay Peninsula cristaius passes into the Tenasserim form described
below as atrior which is very similar to another Burmese form, 6aréez, and to
the Malayan form odscurus and has been identified under both those
names.
This brief epitome of the evidence of intergradation between the Javan
Langur pyrrius and others occurring in more northern countries of tropical
Asia is the explanation of my adoption of the name fyrrhus as the specific
title for all those forms.
A few words inay be said about the Langur known as oéscurus, because
that name figures largely in the literature relating to the Langurs of British
Burma. As stated below, it wasapplied by Blyth, Thomas and Blantord to
Langurs as widely separated geoyraphically as Bhamo and Teaasserim and
this is eloqueat testimony of the closeness of the affinity between these
monkeys. Nowthenamne Semunopithccus obscurus was given by Reid in 1837
(Proc. Zool. Soc., p. 14) toa menagerie specimen of which the locality was
unknown. Fortunately Martin in 1838 saw the specimen aad identifiably
described it (Wag. Nat. Hist., ii, p. 440). Tue hair on the back of the head
formed a crest and was ashy intint, thus contrasting with the blackish-
brown or brownish-black hue of the rest of the head and body. The middle
line of the back was apparently browner, the outside of the thighs greyish
brown, the hands and teet black and the tail dark ashy grey.
668 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
This description applies tolerably closely to many examples in the British
Museum from various places in the Malay Peninsula, but the crest on the
head is not usually present. This race of Puithecus pyrrhus does not
apparently enter the British Indian area; but two closely related subspecies
occur in its southern portion.
A photograph of the Dusky Langur Pithecus pyrrhus obscurus that was
living in the ZOO Se Gardens, Lundon is reproduced on Pl. LV of this paper.
KkY TO THE BRITISH INDIAN RACES OF P. byrrhus
(2) A whorl or parting on the forehead behind the
brow . ; shanicus.
e(a*) No definite hair- whorl on forehead.
(6) No distinct cap of pale hair on the back of the
head and nape.
(c) General colour lighter or darker, but lustrous
grey, buff or brown in reflected light ; lips
flesh-coloured.
(dq) Grey, buff or greyish brown above, grey to
white below ; base of tail grey.
(e) White or nearly white below ; typically a
tuft on the head . phayret.
(e+) Grey below ; tuft on the head present or
absent crepusculus.
(d+) -Colour darker, brown above, deeper grey
below ; base of tail brown ; no tuft . . barbei,
(c1) “General colour deep brown to nearly black a
sometimes varied with grey Bakers hair
.. <.. less lustrous ; lips biack atrior.
-(6+): A sharply contrasted pale grey or butt cap on.
: the back of the head and nape.
(f). Body.and arms blacker, legs not sharply
nee. contrasted with loins fe sanctorum.
é _(f7).. Body and arms browner, legs grey and
sharply contrasted with loins ... flavicauda.
rae Subspecies darbei, Blyth. :
:- Pyesbytis barbei, Blyth, Journ. As. Soc., Bengal, xvi, p. 734, 18; id. Cat.
Mamm. Mus. As. Soc., p. 14, 1863; id. Mamm. Birds, Burma, p. 11, 1875...
- Semnopithecus barbet, Anders., Ann. Zool. Res., Yunnan, p. 12, 1878 -and
Cat. Mamm. Ind. Mus. Be 1, Be 48-49, 1881] (in pale Blanford, Mamm.- Brit.
India; p. 39, 1891.
-- Presbytis ‘melamera, Elliot, re Mag. Nat: jet (8 3), iv, p 267, maea and
Rev. Primates, p. 47, 1912.
With no whorl or parting inthe hairs of the forehead and no crest on the
erown of the head. Lips and eyelids flesh coloured and contrasting with the
blackish hue of the rest of the face. General colour dark brown above with a
pale buffsheen especially noticeable on the head and shoulders. The hind
limbs and tail only slightly paler than the back. ‘The underside dark grey.
Hands, feet and brow black.
Distribution. Interior of the Tipperah Hills and from the Kakhyen Hills as
far south as Bhamo.
- ‘There has been considerable doubt about the identity and geographical distri-
bution of thislangur. In his original description Blyth stated that the skins
came from the Ye Province of Tenasserim; but later, from information re-
ceived from the collector, he corrected this to the interior of the Tipperah
Hills. This alteration was accepted by Anderson and Blanford, although in
his description of the monkey Anderson cited Tenasserim as one of its areas.
He probably had in his hands specimens of the dark race from Northern
Tenasserim, described below, which is very like Jbardez. However that may
be, it appears to me that Blyth’s correction of the locality must be accepted,
especially as the Rev. J. Barbe, after whom this langur was named, is known
to have collected in Chittagong and the Tipperah Hills because he secured there
examples of Pithecus pileatus, a species which certainly is not found in Tenas-
JOURN. BomBay Nat, Hist. Soc. PuatTeE LV.
THE Dusky LANGUR
(Pithecus obscurus)
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 669
serim. I cannot, therefore, agree with Wroughton’s decision to take Tenasserim
as the locality of baréez.
The acceptance of the interior of the Tipperah Hills as the locality of the
type of durbez renders probable, on geographical grounds, the correctness of
Anderson’s identification as darbez of specimens which he collected in the
Kakhyen Hills and on the Irawaddy above Mandalay. Until evidence tothe
contrary is forthcoming, I shall assume he meant the state of Mandalay which
is just tothe northof Bhamo. Now ata place called Cadu Ciaung, Bhamo, L.
Fea long ago secured aspecies of langur, and one of his specimens, presented
to the British Museum aud identified by Thomas as véscurus, was subsequently
described by Elliot as representing a new species to which ke gave the name
melamerus; and this specimen agrees sufficiently closely with Anderson’s
description of the specimens he identified as dar6ez as to leave little Goubt that
it is the same kind of langur. Acting, therefore, on the assumption that
Anderson’s Kakhyen examples were correctly named édarbez, I have added
melamerus to the synonymy of davbez and have taken my diagnosis of barbei
from the type of melamerus, the only available specimen.
‘This specimen agrees with Blyth’s original, but brief, description of barber
in the absence of the crest, in having the face and the under side coloured as in
obscurus, inthe whiteness of the ‘interior base of the thigh’, and in the tail
being only slightly paler than the body ; the shoulders and outside of.the arms
cannot be described as silvered, but other subspecies of P. obscurus vary in the
grey or buff sheen of the upper side.
Anderson, whohad access tothe typical example of darbez in Calcutta an
presumably identified the langurs he procured in the Kakhyen Hills by compa-
rison with it, described the colouras blackish brown above with the sides and
back of the head, the shoulders, fore-limbs, the lower parts of the hind limbs
and the tailas greyish brown, the hands, feet, eyebrows and whiskers being
black and the under parts nearly thesame. But from his citation of Mount
Mooleyit in Tenasserim asa locality for darbez, it may be presumed that his
description applied also to examples from. that area- which, as stated below,
appear to me to represent a distinct local race. - ae ee
Blanford, to whom this langur was unknown, added to the. difficulty. of
identifying--it by placing it in his synopsisin atotally different section from
phayret. : . :
ne Subspecies piayrez, Blyth.
Presbytis obséurus, Biyth., Journ. As. Soc., Bengal, xiii, p. 466, 1844 (not
typical obscurius of Reid).
Presbytis phayret, Blyth., Journ. As. Soc., Bengal, xvi, p. 733, pl. xxxvil, fig.
3, 1847 ; also of Anderson and Blanford (in part).
Presbytis: phayrei, Wroughton, Journ. Bomb. Nat. Hist. Soc., xxiii, Na. F,
_p. 464, 1915 and of. cit., xxiv, p. 297, 1916. ae
Semnopithecus holotephreus, Anders., Zool. Kes., Yunnan, p. 27, 1877.
“Accrest or tuft on the crown of the head but no parting on the forehead ;
lips flesh coloured and eyes partly or wholly encircled by the same tint. The
general colour is variable, being lighter or darker brown above with a varying
extent of greyish sheen over the back and the outside of the limbs, the tail is
dark like the back but sometimes paler at the base or throughout. The under
side is distinctly white or whitish and contrasted strongly or moderately with
the colour of the flanks.
Although closely related to dur6e1, this race may be distinguished from it by
the whiteness of the under side and by the presence of the erect crest on the
crown of the head. oe
Distribution. Arakan; the Bassein district of Pegu ; Prome ; Mount Popa.
Under the name Presbytis, obscurus, Blyth gave a very good description of
this langur, having skins of adults and young collected in Arakan by Capt,
Phayre; and Blanford shot specimens in the Bassein district of Pegu west of
the Bassein river; but the specimens from Moulmein he referred to phayret
belong, I think, to another local race, crepusculus.
In the British Museum there is a half grown specimen from Arakan which
was received long ago fromthe Asiatic Society of Bengal and was for many
years exhibited in the public gallery. It may be regarded as topotypical,
since Blyth gave no definite spot in that district for his type specimen. It is
670 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
dull, lustreless and dark brown above and white below, the white contrasting
with the dark hue of the flanks; thus it agrees closely with Blyth’s coloured
figure of the type of phayreiz. An adult from the S. Zmayi Reserve, 500 feet
(J. M. D. Mackenzie) is also like Blytn’s type in those respects, beiag dark
brown above with the white of the ventral surface strongly pronounced and
sharply contrasted. It hasalso a considerable amount of white on the inner
side and back of the thighs. Also an immature individual from a spot 30
miles S.E. of Prome, 300 feet (J. M. D. Mackenzie) presents the same style
of coloration as the example from Arakan ; but an adult from the same
locality is not so white below and is greyer above. Individual variation
between specimens from the same place is aiso shown by two from Yin on
the Lower Chindwin collected by Mr. Macmillan. One is dark brown above
and on the outside of the thighs, but is not so white below as in the speci-
mens mentioned above; the other is greyer on the back and legs. This
specimen and the greyer of two examples taken 30 miles S.E. of Prome
link the typical darker forms with a good series collected by G.C. Short-
ridge at 4,961 and 4,981 feet on Mount Popa, and the one from Yin on the
Lower Chindwin (G. W. Dawson) mentioned above. These are brown with
a distinct grey sheen all over the upper side, the amount of the grey sheeu
varying individually ; the under side is white or whitish but not very strongly
contrasted with the flanks.
These Mount Popa langurs, as originally pointed out by Wroughton and
Shortridge, differ from the langurs from the Shan States to the north for which
I retain Wroughton’s name shanicus, in having no whorl of radiating hairs on
the forehead and in exhibiting the upstanding crest on the crown. They are
also somewhat paler above and whiter below. ‘They also differ from daréei, as
here understood, in being greyer above, whiter below, and in possessing the
crest. They come nearest to the form described below as Crepusculus, differing
from it appareatly only in having the under side whitish instead of ashy grey.
I have.added holotephreus to the synonymy of phayrez to eliminate it from the
list of so-called species of Pzthecus. The following considerations justify, in
my opinion, this course. (1) The description mentions no character by which
it can be definitely separated from phayret. (2) The locality of the specimen
was unknown. (3) According to Elliot the type was not found in the Calcu:ta
Museum where it was supposed to be preserved. Elliot suggested that it might
be a synonym of barézz ; but the ‘ pale yellowish grey’ tint of the under side
agrees better with phayrez. The type, moreover, hada crest on the crown of
the head which is present in pxayvez but was not recorded in the case of barbei.
Subspecies shanicus, Wroughton,
Presbytis barbei, Wroughton, Journ., Bomb. Nat. Hist. Soc., xxiii, p.
465, 1915.
Presbytis shanicus,;Wroughton, Journ., Bomb. Nat. Hist. Soc., xxv, p. 47, 1917.
Pithecus melamerus, Wroughton, Journ., Bomb. Nat. Hist. Soc., xxvii, p. 552,
1921. (Not melamerus, Elliot).
A dark brownish form closely resembling darbei, as represented by the type
of selamerus, in colour, but distinguished from it and from all the other British
Indian races of Pithecus pyrrhus by possessing on the forehead a naked spot
whence the hair radiates in al! directions as in the evzellus langurs.
Distribution. North Shan States in Upper Burma.
The example of this race which first came to hand and was identified as
barbet by Wroughton was collected by G. C. Shortridge at Ngapyinin opposite
Kyonk Myoung to the east of the lrawaddy. Shortridge noticed that it differ-
ed from langurs he collected on Mount Popa, which were identified as phayret,
in possessing the frontal whorl, in having no upstanding crest on the crown,
and less pallid skin round the eye, the area being reduced to half circles on the
inner side of those organs. Nevertheless, in his description of phkayrez, Blyth
stated very definitely that the eyes were surrounded on the inner side by flesh-
coloured skin.
When Shortridge subsequently secured similar specimens in the North Shan
States, at Séen, Gokteik (2,133 feet) and Pyaung yaung, Wroughton decided
to give a new name to the series because he ascertained from the late Dr.
Annandale that the type of darbez in the Calcutta Museum has no frontal
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 671
parting in the hairs. He therefore named them Presbytis shanicus, selecting
a Séen specimen as the type of the species,
aa,
if
U
ey
iit ANI
Wy,
Fic. 2.—Front view of skull of adult male Pithecus pyrrhus shanicus
from Gokteik in the North Shan States, two-thirds natural size.
B.—The same of Pithecus pyrrhus atrior from the Ye Forest, Tennasserim.
Later, however, he came to the conclusion that these North Shan langurs
were identical with the form from Cadu Ciaung, Bhamo, which Elliot had des-
cribed as Presbytis melamerus; and in Wroughton’s summary melamerus
appears in the table amongst the enfellus group which present the frontal
whorl. But asI have stated above, the type of smelamerus, which is in the
British Museum, has no parting on the forehead at all comparable to that
shown by adults and young from the North Shan States.
672 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
It must be admitted that this difference between langurs so alike in colour
and so close to one another in geographical distribution is very singular; but
on the available evidence, I do not think it would be justifiable to ignore the
character. I have, therefore, decided to admit shanicus as a subspecies of
Pithecus pyrrhus.
Subspecies crepusculus, Elliot.
Presbytis phayrei, Tickell, Journ., As. Soc., Bengal, xxviii, p. 428, 1859.
Presbvtis crepuscula and crepuscula wroughtoni, Elliot, Ann. Mag. Nat. Hist.,
(8), iv, pp. 271-272, 1909; and Rev. Primates, ii, pp. 84-85, 1912 (under
Pygathrix).
Intergrading with phayrez , but on the average paler and greyer above and
not so white below, the ventral surface being grey and sometimes hardly paler
than the dorsal.
Distribution. Northern Tenasserim and adjacent areas of Siam and Laos.
Two examples of this race preserved in the British Museum and collected by
Davison at an altitude of 5,000 feet on Mount Mooleyit, to the west of Moulmein,
were identified by Blanford as phayrez and reasonably considered by him to be
the same as the langur from near Moulmein referred to phayrez bv Tickell.
These specimens were named Presbytis crepuscula by Elliot. ‘The two differ
in tint being in different stages of pelage, as Elliot pointed out. One is toler-
ably uniformy brown all over the upper surface, and on the arms, hands and
feet some rusty brown hairs are retained, so that the extremities are not black
as is typically the case in P. pyrrhus. The other, in fresh pelage, is greyer,
with black hands and feet, and scarcely differs from Mount Popa examples of
pbhayrei except that the belly is greyer and the base of the tail the same tint as
the loins. There is also no upstanding crest on the head. Another example
collected by A. S. Vernay at an altitude of 1,000 feet at Lampha in Tenasserim
is almost identical with the greyer of the two Mount Mooleyit specimens.
This race also spreads into Siam. From this country the British Museum has
a large number of specimens, exhibiting considerable range in the tint of the
upper side from various localities. A brownish specimen from Pacheburi was
named crepuscula wroughtoni by Elliot, but J can find no evidence to justify
itsseparation from crepusculus. In aseries of three examples collected by
A.S. Vernay on the Mewong River 40 and 53 miles east of Um Pang at 800 and
1,000 feet, and one 28 miles sortheast of Um Pang at 1,750feet,no two are
quite alike, the tints of the back varving in greyness and brownness and sheen
and of the arms in darkness according apparently to the age of the pelage.
Another obtained by Mr. Vernay at Kempenpet on the Klong Klung River is
characterized by a marked buffy sheen over the neck and shoulders. Others
obtained by K. G. Gairdner in Seeswad at about 2,000 to 3,000 feet are dull,
dirty-looking skins, their tint being due probably either to the method of pre-
servation or to oldness of the pelage.
The only other specimen I need refer to is a beautiful skin wtth lone greyish
hair obtained by Messrs. J. Delacour and W. Lowe at Xien Quang Koa in
Laos. This has recently been identified by Thomas as Pithecus phayrei.
It may be added that the tuft on the head usually regarded as characteristic
of phayret mav be present or absent in these specimens Ic may be always
present and erect in the living animals, but it is not always preserved in dried
skins.
Subspecies flavicauda, Elliot.
Semnopithecus obscurus, Blanford, Mamm., Brit. India, p.4?, 1888 (in part).
Pygathrix tlavicauda, Elliot, Proc. U.S. Nat. Mus. xxxviii, p. 352, 1910,
and fev. Primates, ii, p. $0, 1912.
Closely related to typical obscurus and sanctorum, with a similar pale grey
or gieyish buff patch on the back of the head but differing in the very sharp
contrast between the grey or greyish buff tint of the tail and of the onter side
of the hind legsand the blackish loins. The sides of the back are blackish
brown, but the centre is browner and this tint, only somewhat paler, pervades
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA _ 673
the shoulders and arms, The head, apart from the back of the crown, is black,
as also are the hands and feet, and the under side is greyish brown.
Distribution. Southern Tenasserim, Mergui Archipelago and parts of
Peninsular Siam.
The British Museum possesses many examples of this race. One collected
by Davison at Bankasun (Bankachon) in S. Tenasserim was ascribed by
Blanford to obscurus. Others from the same locality were collected by G. C.
Shortridge who also got one in Tenasserim Town and another at Banlaw on
the Great Tenasserim River. These specimens exhibit variation in the dis-
tinctness of the pale areas, There is also an example from King Island,
Mergui, collected by Mr. C. Primrose; and Elliot records the race from Sir
William James Islands, Mergui. ‘The type of his f/avicauda came from Trong
in Lower Siam,
Subspecies sanctorum, Elliot.
Presbytis sanctorum, Elliot, Proc, U. S. Nat. Mus., xxxviii, p. 351, 1910 ;
REO, EP VINaies, Ds do, 1913.
Mostly blackish above with soine brown on the middle of the back behind the
shoulders and the outside of the limbs at most a little paler than the upper side
of the body, the hands and feet being, as usual, black. But the tail is pale
buffy grey and the patch on the head whitish. The lower side is uniformly
brownish grey.
Distribution. St. Matthew Island in the Mergui Archipelago.
I have not seen an example of this Langur, my description being adapted
from the remarks of Elliot who declared that in colour sanctorum is very like a
race inhabiting Terautau and Langkawi Island in the Straits of Malacca,
named by Thomas Presbytis obscurus carbo, except that the nuchal crest is
creamy white and the tail pale buffy grey. The specimens of caro in the British
Museum represent a dark insular race closely akin to the mainland race obscurus,
and, in view of Elliot’s statements, I have been compelled to judge of the
colour of sanctorum from them.
Subspecies atrior, nov.
A very dark form nearly uniformly coloured deep, dusky brown all over the
upper parts and the outside of the legs, with less sheen on the hairs; but the
outer side of the arms, especially about the elbows, is paler than the shoulders
and back. ‘The under side is dark, dusky greyish brown. ‘The tail is at most
only slightly paler than the body.
Distribution. Northern parts of Tenasserim and adjoining areas of Siam.
Type. Ye Forest, 500 feet, in the island of Moulmein in the Ataran Division
of Tenasserim. (G. F. R. B. Thurling. B. M.).
This langur may be the same form as the specimen, or specimens, from
Mount Mooleyit referred by Anderson to Semnopithecus barbei. In the dark-
ness of its tint it is very like the example from Bhamo described by Elliot as
melamerus, which I identify with darbez ; but it is somewhat darker above and
below, has little or no white on the back of the thigh, and the skin of the lips
and chin is black in all the examples seen and not flesh-coloured as in darbez,
phayret and other races described above.
In addition to the type above mentioned the British Museum has an example
collected by Davison at the foot of Mount Nwalabo, Tavoy, Tenasserim, which
is a little greyer on the tail and under side than in the type; alsoa specimen
from N. Lat. 14° 25’ and E. Long. 98° 45’, to the east of Tavoy, in Siam ; and
one from Maw to Maw Lai in Siam, both collected by K. G. Gairdner, and
another without locality obtained by the same collector. The example from
Nwalabo in Tavoy shows acertain amount of grizzling on the head, tail and
elsewhere and is of special interest as a connecting form between the type of
atriov and Malayan examples of cristatus.
The above described British Indian subspecies of P. pyrrhus are all
very similar in size. The dimensions in inches of a few of the largest
8
674 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
specimens measured in the flesh may beconveniently set forth in one
table 2
Locality and Sex “Body Tail | Hind foot| Ear | Weight
CURRIES PACU SN em,
shanicus | |
Gokteik, N. Shan States, ¢ 2a re oh 65 13 19 lbs
is - Q Zo Sle sal 63 iP ee Pic
phayret | |
Mt. Popa, Upper Burma, ¢ | 24 52 7 le eee
* 3 4 21s yr Sol 6 1 IS3e ts
crepusculus | | |
Lampha Tenasserim, 22 | aoe 6 pene Ge ull 5
tlavicauda | | | |
Bankachon, Tenasserim, ¢ 25 31 7 ie
i Ma oF 21) eecOe 63 Wee
The recorded measurements of other specimens of these subspecies agree so
closely with those quoted that it is needless to insert them in the table. Unfor-
tunately the only available specimen assigned to darbez was not measured in the
flesh, but the size and proportions of the skin are similar to those selected for
record.
The dimensions in millimeters of some of the skulls are as follows :—
ec TTD wi
Pee: | ze
| ety BSS 1 Se) ae SS ee ae |
St | tea | S8a)20 / s4 | Height and
Locality and Sex | S . a Ay bo" ae | a ‘Width of Orbit
es = | He | oPs | On | Br
yee ik Lew PRRs ae Be Mua b se
shanicus
Gokteik Oa, OL oo 14 65 79% CBE are aD?
- fe) 94 | 28 26 52 eal 24 <> 90
phayret |
S. Zamayi, ae AOS: Nn =35 29 62 81 7 eae aah /4) |
Chindwin, rs4 100 | 33 25 | 64 80 POV RDG Ya
flavicauda | | |
Tenasserim Town, ¢ | 105 33 27. 60 1S AN 26 area!
Bankachon, a 102 ope 26 |= 65 SO Mil: 2 See ee
" Si M0082 Dy Bair leaz5 OL CRO
iP 3 98 38 OS Hass} 74 21 ate oil
- Q 96 28 27 | ~«58 71 25 noe OT
- Q 92 | 30 269 Sore De eS
atyvior |
Ye Forest, Ge) 102 ee 28) ge5ra ars | 26 x 24
Nwalabo, a re ey Ame tia | 61 76 25. xoeZe,
These skull measurements attest general similarity in size between ali the
subspecies. An additional column showing the dimensions of the orbit has
been added because variation in the size of the orbits has been considered to be
a character ot systematic importance. The orbits, for example. are larger in
atrior than in shanicus or phayret. In the skull of atrzor from the Ye Forest,
Tenasserim, the orbit is 3mm. higher and 3mm. wider than in the skull of
phayret from the Southern Zamayi Reserve, although the general dimensions of
the skull of the latter exceed those of the former, except in the width acrossthe
orbits. And this difference in the size of the orbits gives avery different look
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 675
to the full face view of the skull, the skull with the large orbits being not only
wider across the orbits but narrower between them as well, and has more
prominent brow-ridges. These characters, however, like other cranial characters,
are individually variable, as is clearly shown by the records of the specimens of
flavicauda from Tenasserim. These skulls also indicate that other characters
which have been relied upon are unreliable. For example, the second on the
list, from Bankachon, has the palate wider and the tooth-rows more arched
than the first from Tenasserim Town, whereas the fourth on the list, a younger
animal, from Bankachon, reserables in these particulars the one from -
Tenasserim Town.
III. The AyGuLA-group
Differing from the Axfellws-group and the Pyrrhus-group in the colour of the
newly-born young which exhibits a definite pattern of white and black or brown,
extending as a median spinal stripe from the back of the head on to the upper
side of the tail and generally, perhaps always, passing on to the outer surface
of the arms, to a greater or less extent, forming a cross-shaped mark. In the
adults of most of the langurs of this group the white above and below is more
or less obliterated by the extension of the black or brown pigment, but the
inner side of the thigh is almost always white and sharply contrasted with the
tint of the outer side. Usually there is an erect tuft of hair on the back of the
head and a median whorl or a pair of whorls on the forehead. When there isa
pair of whorls the hair typically rises into a crest between them. The skull is
generally distinguisbable from that of the Axtellus- and Pyrrhus-groups by a
marked convex swelling on the base of the nasal bones.
Distribution. Indo-china and the Malay Peninsula through Sumatra, Java
and Borneo.
This group does not extend so far to the north asthe Pyrrhus-group but
overlaps its range in the countries above mentioned.
The name of the group is taken from a Javan species, Pithecus aygula,
described as Szmza aygula by Linnzeus in 1758.
The precise status of the many species and subspecies of this group has not
yet been ascertained. But since only one species the correct name of which
appears to be femoralis, occurs within the area of British India, the characters
of the other representatives of the group need not be considered here.
Pithecus femoralts (Horsfield), Martin.
Semnopithecus femoratis, Martin, in Charlesworth’s Mag. Nat. Hist., ii, p. 436,
1838 (first description of the langur named femovalis, but not described,
by Horsfield, Appendix, Life of Sir T. S. Raffles, p. 642, 1830).
Semnopithecus or Presbytis temoralis of Anderson, Blanford and other authors,
at least in part.
Semnoptithecus neglectus, Schlegei, Mus. Pays-Bas., Simi, p. 47, 1876.
The whole of the upper side is dark blackish brown becoming biack on the
extremities of the limbs and tail ; the face is black with the lips flesh-coloured
and clothed with white hair, and the eyelids also are to a certain extent pallid.
The under side is also mostly blackish brown, or deep brownish grey, frequent-
ly with a white stripe extending down the chest on to the belly which tends to
become lighter and may be quite white where it merges with the pubic region.
The inner side of the arms is frequently pale from the axilla to the elbow and
the inner side of the thighs is always clean white and this white may extend asa
narrow stripe down the inner side of the leg between the knee and the heel,
and on the thigh the white does not blend with the dark tint of the outer side
but is sharply marked off from it by a straight white line. The under side of
the tail is dark throughout like the upper.
The coat is shaggy. There is a thick erect tuft on the crown of the head and
a pair of whorls on the forehead, the hair rising into a median crest between
them.
Distribution. Southern Tenasserim, the Malay Peninsula, Peninsular Siam,
and possibly Sumatra.
Horsfield’s type of this species, an immature animal, isin the British Museum.
It is labelled Sumatra but very likely came from the Malay Peninsula. At all
events it very closely resembles examples from Perak, Selangore, Trang and
other places in Malaya.
G76 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXH
In addition to the typical form there are other subspecies in the Malay
Peninsula and closely related species og subspecies occur in Sumatra, Borneo
and the smaller adjoining islands.
Subspecies £eatiz, Rob. and Kloss.
Presbylis neglecta keatit, Robinson and Kloss, Journ., Fed. Malay States Mus.,
iv; pe 174, 1001,
Pithecus femoralis keatitt, Wroughton, Journ., Bomb. Nat. Hist. Soc., May
: TOUS: a7 02:
Very closely resembling the typical form, but with the lower side more pig-
mented, there being no white on the inner side of the arm between the axilla
and the elbow, no white line on the chest and hardly any white on the belly
which is pigmented backwards to the public region where its dark tint is
strongly contrasted with the white on the inner side of the thighs.
Distribution. 'Tenasserim : Malewoon and Bankachon ; also Kao Wang Hip
in Peninsular Siam.
In the British Museum there are examples of this subspecies, which is of
doubtful value, from the above-mentioned localities.
Ss
os
ESSE
~ es
‘ \
’
i
ve
eS 2s5e2=
Ka a
Mp
i ori
m1
ESS 19g
Fic. 3 A and B.—Front and side view of skull of adult inale Pithecus
femoralis keatit from Bankachon in Tenasserim.
THE LANGURS, OR LEAF MONKEYS, OF BRITISH INDIA 677
The approximate dimensions in inches of two examples are as follows :—
Locality and Sex. shell vite! Tail Hindfoot) Ear | Weight
Body |
—a _ —_ ‘y : — —
Malewoon, of 22+- Ve ol | 7 li 154 Ib
Bankachon, eo 222 33 | 7 14 143. 45
Locality and Sex
Bankachon,
Kao Wang Hip,
of
=
| Total
| length
)
ue
The measurements in millimetres of two skulls are :—
Length of} “Length of Orbital 'Zygomatic
Palate | Upper Molars| Width | Width
13 | 25 ees ee ae
2B 25 — 62 | 70
|
(The end)
THE BIG BULL OF SEMBUTOLLAVU
BY
C. P. GoOULDSBURY
: (With a plate)
Short green grass in the swamps, with elephant grass—over six
feet high as a rule—on the adjoining slopes, intersected here and
there by small patches of jungle, the latter taken into cardamom
cultivation by the neighbouring ryots—a tiny river running all along
the valley—-such was the home for many a year of the bull of this
story.
During the dry months ail the grass that would burn was set
alight, but the swamps always kept their cover of green and the
adjoining patches of cardamom jungle with the undergrowth alone
cleared—-afforded welcome shade for bison and incidentally elephant,
during the scorching midday heat.
The valley in which the bison roamed was roughly five miles in
length, and some two miles broad, broken up by numerous smali
orassland re-entrants.
From a ‘spotting’ point of view the country ijeft a lot to be
desired, as a herd of elephants might be, and very often was, in
close proximity, to the blissful ignorance of the stalker, who would
in all probability only discover it as the light was failing, and a
return to camp was contemplated. On several occasions a long
and tedious detour was therefore unavoidable, and with the short
Indian twilight closing down suddenly into darkness, with a herd
of cow elephants in the vicinity, with small calves at heel, or possibly
a solitary young tusker anxious to throw his weight about, the
return to camp was not always without incident.
Similarly it was inadvisable to leave camp in the mornings before
the light was sufficiently strong to allow one to see at least 100
yards all round. Failure to observe this rule, resulted ina friend
walking practically on to a round grey object which he took, in the
early morning light, to be a rock, and which suddenly turned itself
into a solitary bull elephant, and one too with an unsavoury reputa-
tion. M’s retirement from the scene was rapid, and not altogether
noiseless, but a nasty accident might very easily have occurred, had
he actually stumbled on the animal in the half light.
Bison, of sorts, were always in the vailey, but master bulls,.
although frequenting the entire length of it, were difficult to pick
up, unless they were with the cows during the rutting season—April
and May, and even during these months they would seldom lie up
during the hot hours in the midst of the herd, but usually cleared off
to an isolated patch of jungle soon after dawn, joining up with the
cows again about dusk. At other times of the year the master bull
was somewhere in the valley but, with the elephant grass, well over
Journ., Bombay Nat. Hist. Soc.
THE BiG BULL OF SEMBUTOLLAVU
Tip to tip across forehead we. AO
Widest spread (outside) weer AEC
Girth eae aae PYy) 20%.
See ese raeegmcar arias te
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HOD
oad
THE BIG BOLE OF SEMBUTOLLAVU 679
ones head it was more by luck than judgment that he was ever
spotted, and if he was, it was usually at dusk in the failing light,
and the presence of elephants necessitated the greatest care ina
stalk, even if the latter was at all possible before the light went, the
almost inevitable result being that the bull was left alone that night,
in the hope that he would be in the vicinity at early dawn next
morning—a hope which never materialized in the many attempts
I made during the hot weather months.
It was, to say the least of it, very tantalizing to get reports in
from my shikaris regarding the size of one old bull bison in particular
—solitary at the time—on three occasions when my men had been
down they had sat on a small rocky hill looking down on the old
bull who, as luck would have it, had come out to feed in a large
swamp, known as Sembutollavu, at five o’clock in the afternoon.
The following two week-ends, my only chance of getting away,
were spent fruitlessly in trying to get in touch with him. On one
occasion he was heard moving about in cardamom jungle near the
swamp, but did not come out before dark, and on the next, a
solitary tusker was near the ground usually frequented by the bull,
and it meant getting in between the elephant and the swamp, thereby
giving my wind to what we knew was an elephant with a decided
‘penchant ’ for chasing human beings, although at that time he
had not actually been proscribed by Government as a rouge.
The close season for big game in the district of which I write,
begins on May 3], and the bull being again reported near the Sembu-
tollavu swamp, in company with eight or nine cows, my shikaris
went down for four days to see if they could locate him, in which
case two were to keep in touch with him, the third returning to
me on the Friday. As luck would have it, the week-end fell on the
last two days of the month, which would enable me if necessary to
get in three stalks, Saturday afternoon, Sunday morning and
evening.
At Friday midday, along came the third shikari, and his expan-
sive grin was the forerunner to the welcome news that the bull had
been seen with his cows, on the Wednesday afternoon, in Sembu-
tollavu.
The grass on the ridge above the swamp was long, and the herd
had apparently been lying out for some hours, before it actually
came to the stream to drink. The men had again seen the bull on
Thursday evening but this time he had evidently left the cows
during the previous night, as he rejoined them from a cardamom
jungle some distance away, and some time after they themselves
had been out. On this particular evening the men had heard a
tiger, and sent word to me that with a tiger about, it was probable
that the big bull would now stay with the herd, and not leave the
cows by themselves. Chances therefore of at any rate seeing him
appeared good, and all arrangements were made for tents and kit
to be sent down on Saturday morning.
I started for camp early that afternoon, and reached it about
3.30 p.m. after a stiff climb up one ridge, and a steep descent down
another, the journey taking three hours, and the going as rough
as it could be. It wasa relief to eventually see the tents nestling
680 JOURNAL, BOMBAY NATURAL HIST. SOGIETY, Vol. XXX
under a small bluff, well out of elephant country, on the edge of a
stream, and a greater relief to meet Velandy, my head shikari,
with the news that he had again marked down the bull, with his
cows, in Sembutollavu, the previous evening.
He was very reticent regarding the size of the head, merely
remarking that through the telescope it looked a good one.
A hurried tea, the heavy rifle, a double barrelled -577 by Jeffery
pulled through, and its cartridges collected, the lantern tested, and
various odds and ends such as glasses, the telescope, and raincoat
distributed amongst the shikaris, a move was made to the swamp,
some two miles from camp.
Taking up my position with Velandy on a ridge commanding the
swamp, the second shikari was posted on a small hill to my rear,
to warn us in the event of elephants, which at the time were
numerous in the valley, feeding out of some oi the cardamom
jungles and thus cutting off our return to camp at dusk.
Hardly had Velandy and I settled down when the glasses revealed
seven or eight cow bison lying in high elephant grass, under some
stunted thorn trees. Fora long time it was difficult to pick out the
bull, but Velandy at last spotted him with the telescope, lying out
somewhat to our right. He was under a small tree, and beyond the
colour of his hide proving him a bull, it was impossible to get a
satisfactory view of his head. For over an hour we lay and watched
him. Occasionly a horn would show up above the high grass as he ©
swung his head at a tormenting fly, but before the glasses were on
him, the horn would sink back again.
At last the cows appeared to be inclined to feed. One by one
they got up, and moved slowly down the hill opposite us to the
swamp, through which ran asmall stream. Even now the high
grass gave me only occasional glimpses of them, as they moved
from clump to clump.
A glance at the bull, who was still lying down, and I turned
back to the cows, trying to make out exactly how many there were.
Whilst watching them through the glasses I felt Velandy’s hand
on my knee, and the telescope was pushed towards me.
‘Heis up, Dorai. Look at his head.’ Bringing the telescope on
to the bull, I gave a gasp, and whispered to Velandy ‘ He is well
‘over 40 inches. Why didn’t youtell me when I asked you in camp.’
‘IT knew he was, Dorai, but if I had said so, before you actually
‘gaw him yourself, you would have called mea liar,’ and Velandy
grinned broadly at me.
The bull stood for a moment rubbing his horns against the tree
under which he had been lying, and then sauntered down hill to-
wards his cows. Even though he towered above them, it was only
possible to see the ridge of his back over the elephant grass, but
as he was head on, I got a very fine view of his horns through the
telescope, and, in addition to their spread, the width between tips
struck me as being exceptional.
The cows, seemed in no hurry to come down to me, and as it was
much too far to risk a shot at the bull, covered as he was by the
elephant grass, there was nothing for it but to possess my soul in
patience.
THI BIG BULIVOR SEMBULOLELAVES Ss 0s. 681
As the bull closed up with the cows, the latter started grazing
quietly down towards me, evidently making for a small opening in
the grass at the edge of the stream, which the glasses showed me to
be a ford, from the numerous tracks, and mud on the blades of
grass. If they actualiy came down to the ford, they would be
within seventy yards of my position, but from their rate of pro-
gress, this would take a considerable time, and the wind was none
too steady. Above the ford and on the line they were taking was
a smail tree, near which the grass seemed shorter.
This I estimated to be about 120 yards from me, and after
searching it carefully through the telescope, I decided to take the
bull, if, and when, he got as far, and also if he turned broadside
tO. Mme.
The cows were zigzagging about grazing, ard I hoped, provided
the wind kept steady, that the bull would give me the chance I
wanted.
In the meantime I made myself as comfortable as possible, and
explaining the position to Velandy, told him to watch the result of
my shot when I did fire, and to be ready with spare cartridges.
After what seemed an interminable age the cows one by one,
crossed the shorter strip of grass, and the bull gradually approached
it. Onreaching the small tree, he lay downunder it. This was a
Shock, as I had ne idea how long he proposed remaining in this
position. It was quite impossible attempting to shoot as the grass
though much shorter here than where I had first seen him, was still
too long to enable me to get aclear sight on him, in addition to
which the target had now been considerably reduced in height.
Velandy suggested whistling to get him on to his feet, but in this
event I was certain he would face towards me—a difficult shot to
make certain of at the distance. As he lay be was broadside on,
and my only hope was to cover him, and wait. ‘The wind was
fairly steady, and I got Velandy to watch him with the telescope,
and tell me if he made any movement to getup. In the meantime
I got ready to cover him as soon as he showed any signs of moving.
The cows were by now gradually feeding away from him, which
evidently prevented him from settling down, and it was not long
before Velandy whispered that he was on his knees. As his back
heaved up I covered what I thought was his shoulder—-the grass
made things a bit difficult and holding my breath, let go at him. I
felt as I pulled the trigger that I had held a little low, and let him
have the second barrel as he swung round away from me, but this
was a palpable miss.
The buil went up the hill, keeping the small tree between us, and
Showing so little the effect of the first shot that Velandy declared I
had missed him. Two more barrels failed to hit him. and amidst
wails from Velandy that he was arecord bull, that I was wasting
ammunition on him, etc., etc., I put ina last despairing shot as he
was going uphill away from me, aiming between his horns, with
the 200 yards leaf up, and avery full sight. He bucked to this
shot, swayed, and then came rolling down the hill. There seemed
no life in him through the glasses, so letting the cows—which had
lumbered up the hill after the bull when I started shooting—clear
2
682 JOURNAL, BOMBAY NATURAL AIST, SOCIETY, Vol. XXXII
off, we went up to look athim. ‘The grass when we got into it, was
so long that we had to make a detour to the top of the ridge and then
come down on the tracks made by the buil when rolling. On reach-
ing him, a few sticks thrown at him elicited no response, but on
trying to get at his head, I found this tucked under him, and
impossible to move.
As the shikaris caste would no permit of their handling him, I
sent the second man, who had joined us when I started shooting, to
the nearest jungle for a length of green cane, which on arrival I
knotted round a hind leg, telling the mento heave at the other end.
This was almost too successful, as the slope of the ground very
nearly brought the bull ontop of us. However it had the desired
effect of releasing his head, and I gave another gasp when I put the
tape onit. The widest spread outside showed 431 inches. Girth 20
inches, tip to tip between horns 34inches, tip to tip of horns across
the forehead 75 inches, and length on outside curve of longer horn
302 inches.
My first shot was low down behind the shoulder, and my exceed-
ingly lucky last one had caught him near the spine, forward of the
rump, and as we discovered later, had travelled, and set up in the
lungs.
In addition I cut a spherical 12-bore bullet out from just under the
skin of the flank --a bullet evidently fired into him from a shot gun,
by a cardamom ryot, most of whom own guns, ostensibly to
protect themselves from wild elephants.
As it was then getting dark, and elephants were about, we left the
bull where he lay, to return and secure the head and mask early
next morning.
My return to camp was with mingled feelings. Joy at having
outwitted the old bull predominated, but at the same time there was
the inevitable feeling of sorrow that he would roam his territories
no more. Inthis particular case, not a vestige of his enormous body
was to be wasted, as a pencilled chit from camp to a friend on a
neighbouring estate, early the following morning, produced an
incredible number of pariah coolies, and the head and mask being
removed, I leit for my bungalow knowing that it would not be long
before every particle of meat, including the intestines, would be on
its way to gladden the hearts, and soothe the insides, of numerous
coolies, and provide them with at least one square meai.
The accompanying photograph gives a good idea of the bull’s
head and also of the flies that discovered him, though these proved
no deterrent to the swarms of coolies that later got down to work on
him.
A curious feature was the presence of raw red spots each the size
of half a crown on all four legs of the bull from ‘ stocking ’ to fet-
lock. These were denuded of hair and were raw though not
actually bleeding.
INDIAN DRAGONFLIES
BY
Lt.-CoL, F, C. FRASER, I.M.S., F.E.S.
Part XXX
(With three plates)
(Continued trom page 459 of this Volume.)
Genus—MICROMERUS, Ramb.
Micromerus, Ramb. Ins. Nevrop. p. 238. (1842) ;—Selys., Syn. Cal., p. 64
(1853) ;—Id. Mon. Cal. p. 233 (1858) ;—Walk. List. Neur. B. M. iv, p. 652
(1853) ;—Kirby, Cat. Odon., p. 115 (1880) ;—Will. Proc. U. S. Nat. Mus.
vol. xxviii, p. 172 (1904) ;—Ris. Ent. Mitteil. V. Nr. 9/12 (1916).
_ Characters as for the subfamily, the epistome however, much more developed
than in Rhinocypha, much more projected, so that a deep fissure intervenes
between the frons and nasus. Fore and hind-wings similarly shaped, very long
and narrow, reticulation moderately open ; fore-wings of male, in most species,
tipped with black, otherwise all wings of both sexes completely hyaline (except
in suellemani and bisignatus, both ex-Indian species, in which the wings are
marked with broad blackish brown fasciz as found in many species of Ahkinocy-
pha) ; antenodal nervures 4 to 7 in number; sectors of are arising from a
common point, where the arc is strongly angulated at alittle above its middle ;
ZA undulated from its origin ; petiolation ends proximad of the first antenodal
nervure ; intercalated nervures few in number and found only near apices of
wings ; A777 arising widely distad of node; pterostigma absent in forewings of
male except in those species where the apices are not tipped with black, present
and very elongate in the hind, and in all wings of the female.
Mesothoracic triangle absent or reduced, uncoloured ; legs long and slim,
usually all femora and tibiz pruinosed white on the flexor surfaces. Abdomen
short and stout, much shorter than wings, fusiform, depressed, narrowing very
gradually towards the 10th segment. Anal appendages very similar to those of
Rhinocypha, superiors twice the length of segment 10, pincer-like, cylindrical
and pojnted at apices ; inferiors obtuse, less than half the length of superiors.
Vulvar scale very robust, shaped as for Rhinocypha.
Larva very similar to that of Ahkinocypha (vide Plate III.)
Distribution. Southern Asia from Western India to Borneo and New Guinea.
Most Indian species appear to have origin from /ineatus, which is only found
typically from Lower Burma within Indian limits. Elsewhere it comes from
Java and Siam, several subspecies or races occurring throughout Burma,
Continental India and Ceylon. In the latter island, due to insulation, we find
asurprising development of the genus, the species found there not being
apparently closely related to lineutus.
Their habits do not differ in any respects from those of Rhinocypha, and they
are found in similar habitats.
Genotype—Microm-rus lineatus Burm.
_The present position of the nomenclature of MWicromerus and Litellago is
uncertain. The first mention of the latter genus is found in Selys, Mon. Lib.
Eur. p. 200 (1840), wherein he gives a short description of the genus and cites
Agrion lineata as the genotype, but also gives fenestrata and fulgipennis (now
classed as Rhinocypha. but at that time as Agrion.) as belonging to the same
genus. Lzneata had however been described by Burmeister as a Calopleryx
and Selys committed anerror in assigning it to Agrion. That he did so is
perfectly clear, from the fact that he refers to the Won. Lib. Eur., when des-
cribing genus Wicromerus in the Mon. Cal., p. 233. It seems perfectly clear
therefore that /ineafa as referred to inthe Mon. Lib. Hur. was Burmeister’s
Calopteryx lineata (Micromerus lineatus), and equally clear that it was given
684 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol XXXIi
as the genotype for Lzdellago, and this being the case, Wicromerus becomes a
mere synonym for Zzdellago, and the latter name, for what it now stands, is
therefore preoccupied, AsI have already committed myself, in the present
Monograph, to the present use of the names, a change of seats in midstream
might cause endless confusion and I have therefore decided to let things stand
as they are. In a future revision however, it will beeome necessary to rectify
these errors. (/icromerus becoming Libellago and a new name being chosen for
the latter, for which I would suggest Chlorocypha.
Key to Indian species of Wicromerus.
1, Abdomen marked with bright orange or brick-red... 2
Abdomen marked with bright citron yellow or
greenish i ey ae, bes 3
~
2. Greater part of abdomen bright orange; middorsal
carina of abdominal segments not marked with
black. (Burma.) Mik AIS lace? .. JW. aurantiacus.
Abdominal segments bearing large spots of brick-
red ; dorsal carina of all segments black. (Ceylon.) MM. greent.
3. Male with 6 antenodal nervures to all wings ae ee
Male with 5 antenodal nervures. (Burma, N.
India. ) ae Pee an M. lineatus lineatius.
4. Black tip to forewings of male 4.to $ mm. in length. o)
Black tip to forewings of male only 3 mm in length, ae ac
.5. Humeral stripe present at its middle ; posterior lobe
with a large dorsal spot; thoracic markings
orange. (Nicobars.) pad re a
Humeral stripe absent ; posterior lobe of prothorax
unmarked ; thoracic markings citron yellow or iat
green. (Andamans) — sis ... MW. andamanensis.
M, blandus.
6. Small insects, hindwing less than 18mm. in length ;
linear spot-on hinder border of occiput with a
inedian anterior prolongation ; abdominal mark- =a
ings citron yellow (Continental India and Ceylon.) WV. lineatus indica.
Larger insects with hindwing more than 20 mm. in ~ :
length ; linear spot on hinder border of- occiput
without median. prolongation ; abdominal -mark- er
-- - . ings bluish or bluish-green, -(Ceylon.) - 5S M. finalts.
... Micromerus lineatus lineatus, Burm. (Calopteryx lineata Burm.) Handb, Ent.
ii, p. 826 (1839),—Selys. (Agrion lineata) Mon. Lib. d?Europe, p..200
(1840), —Ramb: (MWicromerus lineatus) Ins. Nevrop., p. 238 (1842),—
' Selys. Syn. Cal., p. 65 (1853),—Id. Mon. Cal., p. 237 (1858) ,-—-Id. Bull.
Acad. Belg. (2), xxvii, p. 665 (1869).—Walk. (Lzbellago lineata) List.
Neur. Ins. B. M., p. 653 (1853), —Ramb. 1. c. (A/icromerus uxor) p. 239
(1842),—Kirby Cat. Odon. p. 115 (1890),--Selys. Ann. Mus, Civ.
Genov. (2) x (xxx), p. 62 (1891),—Laid. Fascic. Malay. (Zool), pt. 1,
p. 197 (1903), —Will. Proc. U.S; Nat. Mus., vol. xxviii, p. 171, figs. 5.and
6 (1904),—Laid. Rec. Ind. Mus., vol. xiii, p. 39 (1917),—F ras. ldzd., vol.
xxvi, p. 483 (1924).--Micromerus obscurus, Kirby Proc. Zool. Soc.
Lond., p. 328t. 33, f. 1(1886).—Id. Cat. Odon., p. 115 (1890).
Male. Abdomen 15mm. Hindwing 17 mm.
Head. Labium dull brownish yellow to blackish ; labrum blackish brown,
the anterior border finely yellow ; epistome black, the anterior facet blue black
metallic, above velvety black ; frons and vertex velvety black—with 2 transverse
oval citron yellow spots on former near middle line and a large oval spot on
outer side of each posterior ocellus ; occiput black with a large postocular <pot
subtriangular in shape, on each side separated from, more or less narrowly, a
median crown-shaped spot. The latter spot rather variable, in the type, trans-
versely elongate witha median spot in front of its middle, but in others, con-
fluent with this spot and with additional triangular points on either side.
Prothorax black marked with citron yellow as follows :—The border of the
posterior lobe very finely except below where itis expanded into a pyriform
INDIAN DRAGONFLIES . — 685
spot ; a large transversely oval spot on dorsum of posterior lobe, this part being
raised into an oval boss coated with fine hairs ; a narrow anterior collar, anda
large subtriangular spot on each side of middle lobe.
Thorax black marked with citron or greenish yellow as follows :—The whole
of antealar sinus, the middorsal carina finely, a narrow ragged antehumeral
stripe incomplete above, a small upper antehumeral spot, a norrow curved post-
humeral stripe incomplete below, Laterally nearly the whole cf the mesepi-
meron which shows a narrow wedge of black at its middle and a hook- -shaped
or triangular-shaped invasion of black above ; lastly the whole of metepimeron
except the anterior sutural line and the ventral border narrowly. Beneath
yellow.
Legs black, flexor surfaces of tibie and of the anterior pair of. femora prui-
nosed white.
Wings hyaline, the bases of all, nearly to level of node, palely tinted with
amber ; the apices of forewings (except i in tenerals where no marking is visible)
tipped with black as far proximad as inner end of pterostigma of hindwing but
falling short of hinder border of wing. Pterostigina absent in forewing, black
tumid and elongate in hindwing where it covers from 2 to 24 cells. Costa of
forewing gradually thickened towards the site where pterostigma would
ordinarily be; discoidal cell traversed once ; nodal index, 5 antenodal nervures,
8 to 10 postnodals.
Abdomen black and bright greenish or golden yellow, the latter colour pre-
vailing on the anterior segments, the former on the hinder. Segment 1 with
the apical border narrowly and the sides broadly yellow, segments 2 to 5 with
the latero-ventral borders and the intersegmental sutures narrowly black, the
carina also finely black from segment 2 to 4, orin tenerals from 2 to 6, but
arrested short of apical border where the black expands or is connected to two
wing-like projections, so that this together with the carinal stripe form inverted
capital ‘Ts’ (in some specimens, the armsof the‘ T”’ are slightly separated
from the stem, but this is rare). Segments 5 and 6 with two large dorsal vellow
spots tapering apicad, enclosed by an expansion and confluence of the carina!
T-spot with the apical ring and latero-ventra] black ; segments 7 and $ may
show smaller similar spots, but adults have these unmarked, as also 9 and 10.
Anal appendages black, superiors twice the length of segment 10, separated
widely at base, converging at apices which are turned slightly in, cylindrical
and very slightly tapered... Inferior appendages only half the length of supe-
riors, closely. apposed at bases and. again at.apices so as to enclose a minute
foramen, blunt, broad at base, narrowed thereafter.
Female. Abdomen 13to16 mm. Hindwing 18 to 22 mm.
A stouter, more robust insect than the male, marked very similarly but the
yellow more extensive.
Head with the same markings as for male but with the. following additional
spots :—The labrum and labium, the former with a small basal median-tongue
of black:; the anterior surface of epistome (which in the male is metallic blue) ;
the bases of mandibles, cheeks, basal joints of antennae, a large spot on either
side of epistome and two large diamond-shaped spots on front of frons, narrow-
ly separated from the smaller hinder pair.
Prothorax, thorax and legs as for male.
Abdomen black marked with yellow as follows :—The apical border and sides
of segment 1; the dorsal carina on segments 2to7; a longitudinal stripe on
dorsum of 8 and 9, whilst that of 10is unmarked. Laterally the seginents
yellow with a large apical spot notched infront on segments 2 to 7, bifid on
segment 8, and present only as asmall yellow spot on sides of 9 and 10.
(Briefly the abdomen may be said to be yellow with two longitudinal bands of
black and black articular rings at the intersegmental nodes.)
Anal appendages black, twice as longas segment 10, pointed, widely sepa-
rated throughout. Legs black, the flexor surfaces. of femora and tibiz yellow.
Wings entirely hyaline, the basal area, especially along the costa slightly
tinted with amber ; pterostigma, present in all wings, elongate, black or yellow
framed in black according to age, covering 2 or 3 cells ; 6 antenodal nervures,
9 to 10 postnodals.
Distribution. Throughout the Sondaic Archipelago and the adjacent main-
land, Java, Sumatra, Siam, Annam, Burma and North India. Kirby’s oéscurus,
described from a teneral specimen, from northern India, is identical to speci-
mens which I have received from Dehra Dun, Cachar and Upper Burma. It
686 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
differs only in the limitation of black markings due to tenerality and is
undoubtedly dzweatus. In Peninsular and Southern India we find a distinct
subspecies replacing typical Jzweatws, which I have named indica. ‘The two
Ceylon forms show sufficient differences as to deserve specific rank.
Micromerus lineatus indica, subsp. nov. (A/zcromerus lineatus) Fras. Rec. Ind.
Mus.. vol. xxvi, p. 483 and pl. xxiv, fig. 4 (1924),—Laid. Spolia zey-
lanica (M. lineatus, Burm.), vol. xii, part 47, pp. 354, 355 (1924).
Male. Abdomen 14to16mm. Hindwing 15 to 18 mm.
Head very similar to that of Zzmeatus, labium ochreous, labrum dark brown-
ish yellow, black at base; a more or less obscure spot on either side of
epistome, the anterior facet of which is metallic blue as in d¢meatus ; eyes brown
but with a bluish metallic lustre in certain lights; bases of mandibles and
cheeks narrowly yellow ; the oval spots on frons reniform in shape and often a
second pair of smaller rounded spots behind them ; crown-shaped mark on
occiput with an elongate median point, but no lateral points, or if present then
very short, usually connected to the postocular spots.
Prothorax, thorax and legs as for type /zmeatus, the femora pulverulent white
at basal halves only.
Wings as for /zneatus but the apical black spot of forewings distinctly longer
(absent in tenerals) ; nodal index differing slightly, in Kanara specimens 6
antenodals in the forewings, 7 in the hind, less commonly 5 to 6in the hind ;
in Deccani specimens, almost always 6 in ‘both wings and 10 to 14 postnodals.
The amber tinting at base usually very marked, upto or even beyond node ; dis-
coidal triangles traversed once ; pterostigma as in deneatus.
Abdomen. (As in Lidellago, so in Micromerus specialization seems to have
confined itself almost entirely to the colouring and markings of the abdomen.)
Segment 1 bright golden yellow with the base narrowly and asmall quadrate
black spot; segments 2 to6 bright citron yellow, the dorsum golden green;
often very green, 2 with a thick middorsal dumb-bell-shaped black marking
confluent with a narrow basal ring, the ends of which are prolonged along the
sides nearly to apical border, and separated very narrowly from the ventral
border, segrnents 3 to 5 with broad middorsal stripes confluent with narrow
basal rings and expanded somewhat apically, whilst laterally near the apical
border, asmall ventro-lateral spot of black is seen; segments 6and7 with
basal subdorsal wedge-shaped yellow spots enclosed by a fusion of the mid-
dorsal black with an elongated lateral spot, segments 8 to 10 black. Insome
specimens the middorsal marking of segment 2 is narrowly bisected with
yellow, and there is nearly always an indication of the beginning of such a line
apicad.
Anal appendages not differing from /zneatus.
Female. Abdomen13to17 mm. Hindwing 17 to 20 mm.
Head. Labium yellow ; labrum yellow with a small median basal point or
tongue of black; epistome entirely yellow in front, similarly coloured abcve
but here edged with black anteriorly and laterally, its dorsum clouded with
light brown andthe base narrowly dark brown witha tongue of the same
colour running obliquely forwards and outwards; frons with two large
quadrate citron yellow spots, separated narrowly by a black median stripe which
is confluent anteriorly and posteriorly with an equally narrow black stripe, the
whole thus forming a black anchor-like mark, accentuated by the outer corners
of the hinder portion angulated forwards like the flukes of an anchor ; markings
of rest of head similar to those of male but an additional pair of spots situated
between ocelli. Eyes brown,
Prothorax entirely similar to the male. Thorax similar but lateral markings
rather more extensive. Legs yellow, the femora lined externally with dark
brown.
Abdomen largely yellow with black markings as follows :—Segment 1 with a
large quadrate spot on dorsum, the base narrowly and the apical border
laterally; segment 2 to & with the dorsum very broadly but bisected by a
narrow middorsal carinal yellow line which runs as far as segment 9, but on the
last two segments is fusiform, tapering apically and basad ; laterally a stripe
more or less defined, broadening abruptly subapicad and not quite reaching
the base of segments. On segment 9 this line expands basad and becomes con-
fluent with the dorsal black to enclose a large rounded subdorsal yellow spot,
segment 10 black with two transversely elongate subdorsal yellow spots.
Journ., Bombay Nat. Hist. Soc, PLATE I,
7 8 oe eds il 12
» Micromerus greent, Laid. (Ceylon.)
Fic. 1
An Da aurantiacus, Selys. (Burma.)
3h andamanensis, Fras. (Andamans. )
4, finalis, Selys. (Ceylon.)
5. lineatus indica, subsp. nov. (Kanara.)
6. lineatus, Burm. (Dehra Dun.)
he (Cachar. )
8. (Java.)
9. ” ”” ” av’ (Siam.)
10. Calocypha laidlawi (Fras.) (Kanara.)
ll. Libellago astatica, Selys. (Burma.)
curta, Selys. (Uganda.)
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INDIAN DRAGONFLIES 687
Wings hyaline palely amber tinted ; pterostigma present in all wing, creamy
white in a black frame which expands somewhat proximad, over 2 cells; nodal
index,—6 to 7 antenodals, 9 to 10 postnodals ; discoidal cell traversed once in all
wings.
nee appendages blackish brown, pointed, tapered, widely divaricate, rather
longer than segment 10.
Vulvar scale robust, with a large spot of yellow laterally.
Distribution. Avery common insect throughout South India especially in the
Western Ghats and Deccan. It is common on the river banks at Poona and
Khandala, less common in Coorg, Kanara and Malabar.
Laidlaw reports it from Ceylon, Haragama, July, and remarks on its difference
from type Jineatus lineatus Burm., from Burma and Siam.
It appears to be double-brooded, as it may be found pretty well throughout
the year. The abdominal markings will easily distinguish it from typical
lineatus. Type and co-type in B. M. from Poona.
Micromerus lineatus blandus Selys, Syn. Cal., p. 64 (1853) ;—Id. Mon. Cal.,
p. 254 (1854) ; Walk. (Lzbellago blandus) List. Neur. Ins. B. M. iv,
p. 652 (1853) ;—Laid. Rec. Ind Mus., vol. xiii. p. 39 (1917).
Male. Abdomen 17-18 mm. Hindwing 20-21 mm.
Female. Abdomen 12-15mm. Hindwing 19-21 mm.
I have not seen this species, which in the Selysian description is said to
closely resemble /zmeatus, a fact which leads me to describe it as a subspecies
of the typical form from Java.
It is a more robust insect and differs from dzneatus as follows :—
The yellow stripe on the cheeks bordering the eyes absent ; prothorax black,
its middle lobe unmarked but the large dorsal tumid spot on the posterior lobe
extending forwards as a point; thoracic markings more orange than yellow ;
the humeral stripe incomplete above and below, present in its middle third only;
the anterior of the two broad lateral fasciz broken up into three large spots,
the upper very minute ; dorsal markings of abdomen broader; apical black
spot of forewings longer than broad, about 4 mm. in length; pterostigma
longer, covers about 5 cells in the hindwings; 6 antenodal nervures in all
wings.
Female very similar to that of /zmeatus. ‘The yellow markings more restricted
the humeral stripe absent, the anterior of the two lateral fascize divided up into
two spots ; the dorsal black of segments 3 to 6 narrower, nearly interrupted on
some, ground colour orange ; nodal index higher, 14 postnodal to hindwings,
12 to the fore ; pterostigma yellow, black, proximad, broader, over 3 to 5 cells.
Distribution. Confined to the Nicobars. Type in the Selvsian coll.
Micromerus andamanensis, Fras. Rec. Ind. Mus., vol. xxvi, p. 410 (1924),
Male. (Female unknown,) Abdomen16mm. Hindwing 20 mm.
Head. Eyes bluish grey; labium and labrum black ; rest of head jet black
with the following citron yellow markings ;—two narrow oval spots at base of
epistome above, a large ovalspot on the outer side of each posterior ccellus, a
similar postocular spot on each side, and a linear spot on the hinder border of
occiput.
ee black with a large yellow spot on each side.
Thorax black marked with golden greenish yellow as follows, two small spots
on the antealar sinus, a fine carinal dorsal line, two fine antehumeral stripes
incomplete above, an oblique linear spot on each side of the antealar sinus.
Laterally two irregular spots lying between the lateral sutures, the anterior
golden yellow, the posterior distinctly greenish, and a broad stripe occupying
the upper anterior three-fourths on the metepimeron.
Legs black, the hinder tibiz pulverulent white on the flexor surface. Wings
hyaline, faintly amber tinted at bases, forewings broadly tipped with black to 4
mm. in extent and metallic blue by reflected light ; hindwings enfumed at
apices ; pterostigma black, only present in hind wings; 5to 6 antenodal
nervures in all wings.
Abdomen golden yellow marked with black as follows :—Segment 1 narrowly
black at base, more broadly at sides; segment 2 with a broad dorsal stripe
expanded at basal and apical ends of segment, also a broad black stripe low
down on sides not extending as far as apical border of segment ; segments 3 to
6 with broad _middorsal stripes expanded only at apical border of segments
688 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
where it forms a narrow ring; segment 7 similar but on the sides, the apical
black extending forwards nearly as far as base of segment; segments 8 to 10
entirely black.
Anal appendages black, similar to those of /zweatus.
Distribution, Andamans, the type, a male, isin the Indian Museum, and
was taken by the late Dr. Annandale on the Western slopes of Mt. Harriet, No-
vember 30, 1923(fluttering over the surface of a pool below a waterfall). Altitude
500 ft. This very beautiful insect is easily distinguished by the broad metallic blue
tip to its wings, as wellas the extreme grade of melanism as compared to other
species of the genus, it is to be hoped that some collector on the spot will find
the female for us. Itshould not be difficult to secure a fine series as most spe-
cies are usually abundant where taken. The femaleis probably very similar
to dineatus and without a black tip to her wings.
Micromerus finalis Selys, Ball. Acad. Belg. (2) xxvii, p. 665 (1869) ,—Id.
lbid., XXxvi, p. 616 (1873),—Kirb,. Cat. Odon. 115 (1890),—Id. Journ.
Linn. Soc. (Zool) vol. xxiv, p. 556 (1894),—Laid., Spolia zeylanica
vol. xii, parl 47, pp. 351-352 (1924).
Male. Abdomen 16mm. Hindwing 21mm.
Head. Labium yellow, lobes tipped with black ; labrum glossy black, un-
marked ; epistome metallic blue or violety blue ; cheeks with a narrow yellow
stripe against the eyes ; frons black with a pair of large rectangular spots of
citron yellow ; vertex black with an oval citron yellow spot on the outer side of
each posterior ocellus ; occiput black with a rounded postocular spot on each
side and a linear spot along the hinder border, narrowly separated from the
former.
Prothorax black marked exactly asin dineatus, but the anterior of the two
lateral fascize with a large black spot below. Beneath black marked obscurely
with dark yellow areas.
Lees black, all tibia pulverulent white on flexor surfaces and all femora
pulverulent on same surfaces except for a short distance at the distal
extremities.
Wings hyaline palely amber tinted, especially the hinder ; forewings tipped
with black for about 3mm., pterostigma absent; hindwing with well-formed
elongate pterostigma, covering 3 cells ; 6 antenodal nervures in all wings, 10
postnodals in the hind ; discoidal cell traversed once in all wings.
Abdomen black marked with bright citron yellow laterally, changing to
green or bluish-green on dorsum. Segment 1 with the base narrowly and the
sides broadly black ; segment 2 with a large angulated subdorsal spot on either
side, the black forming a large T-shaped mark on dorsum, base, and sides
where the arms of the T turn back ; segments 3 to 4 very similar but the head
of the T-shaped mark very narrow, as also its lateral prolongations back, and
on the sides of the latter segment, the beginnings of a narrow lateral stripe
apicad ; segment 5 to 7 with subdorsal elongated spots divided, except on seg-
ment 5, by a narrow lateral black stripe; segments 8 to 10 unmarked.
Anal appendages black, similar in structure to those of /zneatus.
(The female has not hitherto been described.)
Female. Abdomen 16mm. Hindwing 25mm.
Head black marked with bright yellow ochre as follows —the labium except
for the tips of lobes ; the labrum except for the base which is narrowly black
and the border which is broadly black, and also a triangular prolongation from
the middle of the basal black which may be confluent with that of the fore-
border. (In very adult specimens, the yellow reduced to two minute points) ; a
pyriform spot on the dorsum of epistome, reduced in adult specimens to a tiny
linear mark. (The aiuterior facet of epistome metallic blue, but more dully so
than in male) ; the bases of mandibles broadly yellow, the basal segments of
antenne, the cheeks, two pairs of spots on frons in front of ocelli, an oval spot
on outer side of each hinder ocellus and similar spots on occiput as in male. (In
very adult specimens, these spots reduced to mere points, and the median
linear spot on hinder border of occiput entirely absent.)
Prothorax black with a linear or top-shaped spot on middorsum of posterior
lobe and a large spot on the sides of mid-lobe.
Thorax black marked very much as in the male but the markings more res-
tricted. ‘The antealar sinus with two small yellow spots, the middorsal carina
very finely, the humeral stripe either absent or represented by a chain of small
JOURN. BOMBAY Nat Hist. Soc. PieATe ul
Two views of the River Cauvery, Coorg-Mysore Frontier; favourite
haunts of Micromerus lineatus indica and Rhinocypha bisignata. Other
species taken here include Mesogomphus lineatus, Macromia cingulata
and indica, Macromidia donaldi, Onychothemis testacea, Zygonyz torrida
| isis. Chloroneura apicalis and quadrimaculata were found here also,
| the former lurking under the bigger trees, the latter perched on bare
rocks in mid-stream.
INDIAN DRAGONFLIES 689
spots, or a tiny linear upper spot ; the lateral fascize similar or the anterior one
bisected below and completely cut in two at its middle, the upper portion more
or less obsolete. Beneath brownish yellow with the sutures diffusely black.
Legs yellowish, black externally, the hinder pair of femora bright yellow
except at the distal ends.
Wings palely tinted and enfumed, the hinder pair occasionally tipped narrowly
and diffusely with blackish brown ; pterostigma creamy white framed in black
and clouded with black proximad ; discoidal cells traversed once or twice; 7
antenodal nervures to al] wings, 11 to 13 postnodals.
Abdomen black marked with bright citron yellow or greenish yellow as
follows,—the middorsal carina narrowly from segment 2 to 8; segment 1 as for
male, segments 2 to 8 with broad lateral stripes constricted subapicad, markedly
on segments 2 and 3, less so on 4 to 6, tapered apicad on segment 7 and_broad-
ening apicad on segment 8; alarge round spot replacing this stripe on seg-
ment 9, and a smaller spot on segment 10, absent in very adult specimens. A
chain of broad yellow spots on ventrum. Vulvar scale robust, shaped as for
lineatus.
Anal, appendages black, conical, pointed at apices, twice the length of
segment 10.
Distribution. Confined to the montane and submontane districts of Ceylon.
Col. F. Wall took a fine series at Banderawela and Nalande, the former 5,000 ft.,
during October, but it probably has a succession of broods throughout the
monsoons. I took this species in August at Dyatalava, and Col. Wall and Mr.
E. Green at Kandy, 2,000 ft. October-November.
Micromerus greeni, Laid. Spolza Zeylanica, vol. xii, part 47, pp. 352-354
(1924).
Male. Abdomen 14mm. Hindwing 17-19 mm.
Head black marked with bright ochre as follows :— Labium entirely black ;
labrum glossy black, unmarked ; epistome black, its anterior facet bluish-violet
metallic; frons with a single pair of spots, less commonly a second pair of very
minute points posterior to anterior pair; an oval spot on the outer side of each
hinder ocellus, a rounded postocular spot on each side of occiput and a linear
spot along its hinder border, which in some, has a tiny medial point anteriorly ;
the cheeks with a narrow yellow bordering against the eyes.
Prothorax black marked with citron yellow as follows :—a narrow anterior
collar followed Iaterally by asmall spot and then a large medio-lateral subtrian-
gular spot. A large spot on the dorsum of posterior lobe shaped like the ace-
of-clubs, and two smaller spots low down on its sides.
Thorax black marked as for /ineatus but the markings more reduced as
follows,—the humeral stripe present only as a tiny upper linear spot, the antealar
sinus with two small rounded spots, the upper part of the anterior lateral
fascia tapered away above and shaped like a sickle, the second or hinder fascia
framed heavily in black. Beneath black.
Legs black, all tibiae pulverulent white on flexor surfaces, all femora pulveru-
lent creamy yellow on distal halves of flexor surface.
Wings hyaline palely tinted with yellow, the hind palely enfumed, the apices
of forewings tipped with black for about 3 mm. and with a more or less marked
metallic lustre at this part ; the hinder with apices clouded with black at extreme
border. Pterostigma present only in hindwings, covers 3 to 33 cells, black.
Costa proximad to node, and the nervures behind it, bright blood-red, this
colour less marked in the hindwing and darkening proximad. Discoidal cells
with one traversing nervure ; 6 antenodal nervures in all wings, 10 to 12 post-
nodals in hindwings.
Abdomen black marked with bright citron yellow and blood-red as follows, —
segment 1 with an apical ring of citron yellow broadening laterally ; segment 2
with a large subdorsal spot not reaching the basal extremity and constricted
near its apical end, citron yellow ; segment 3 witha broader spot citron yellow
tinged with reddish, extending from base to apex of segment and abruptly
narrowed subapicad and, partially divided by a narrow longitudinal black
stripe running from its apical end ; segments 4 to 10 with the markings blood-
or brick-red, broad paired spots separated by the black dorsal carina, constricted
at the apical end and not quite reaching the apex of segments, thus forming
rather narrow inverted black capital T-like markings on each segment,
Segments 9 and 10 with only a small round apical spot on each side.
10
6909 JOURNAL, BOMBAY NATURAL FIST. SOCIETY, Vol. SOC,
Anal appendages black, rather slenderer than in other species, twice as long
as segment 10.
(The female has not hitherto been described.)
Female. Abdomen 12to13 mm. Hindwing 19 to 20 min.
Head marked very similarly to the male, differs as follows:—Labrum with a
pair of large triangular yellow spots ; epistome with a metallic colouring poorly
developed ; two small linear spots on dorsum of epistome, obsolete in very
adult specimens ; bordering medial spot on occiput absent.
Prothorax black with a narrow linear medial stripe on dorsum of posterior
lobe, a large spot low down on its sides, a large spot on each side of medial
lobe and, rarely, a fine anterior collar.
Thorax black marked exactly as for male, but in very adult specimens,
with the antero-lateral fascia cut in to two spots at its middle.
Legs black, some slight pruinescence on the flexor surface of femora, and
the same surface on anterior pair of tibiae pulverulent creamy white except at
distal ends.
Abdomen black marked with citron yellow, with a greenish tinge on the
anterior segments as follows:—Segment 1 with a lateral quadrate spot ;
segments 2 to 8 with the middorsal carina finely yellow and with lateral
stripes, which are constricted apicad on segments 2 to 5, tappered on 6 and/7,
reduced to a spot on the others or, in very adult speciinens absent on segments
9 and 19, less often on segment 8 and rarely present on segment 10.
Anal appendages and vulvar scale as for /zneatus.
Wings hyaline, not enfumed or tinted ; 5 to 6 antenodal nervures, 10 to 12
postncdals in all wings; discoidal cell traversed once; pterostigma white
framed in black and clouded at its proximal end.
Distribution. Col. F. Wall took a number of these insects at Nalande,
Ceylon, during September and October, whilst Mr. E. Green took specimens at
Haragama, Peradeniva and Kandy from June to December, so that it appears
to have a long season like most species of the genus. It closely resembles JZ.
aurantiacus, but the black markings are more extensive. Easily distinguished
from all other species by its red abdomen, strongly reminescent of the African
Libellagos.
Micromerus aurantiacus, Selys, Bull. Acad. Belg. (2), xvii, p. 448 (1859), —Id.
Ibid., (2), xlvii, p. 397 (1879),—Kirby, Cat. Odon., p. 115 (1890) ,—Laid.
Proc. Zool. Soc. Lond., p. 331 Sept. (1920),—Will. Proc. U. S. Nat.
Mus.; vol. xxvili; p.172,’fig. 7 (1904).
Male. Abdomen 12-15mm. Hindwing 15-16 mm.
Head black spotted with citron yellow as follows:—A lateral spot on each
side of epistome, which latter is metallic blue at its anterior extremity ; two
pairs of squarish spots on frons, a rounded spot on the outer side of each hinder
ocellus, a postocular rounded spot on each side of occiput anda linear spot on
its hinder border. In very adult specimens the latter spot and the hinder pair
of the frontal spot absent.
Prothorax black with a small geminate spot on the middorsum of posterior
lobe and a large spot on each side of mid-lobe, yellow.
Thorax black marked with citron yellow as follows,—the middorsal carina
finely, an antehumeral stripe broadening below, a fine humeral stripe incom-
plete below or almost obsolete, two small spots on antealar sinus, and two
broad fascize on the sides of thorax, the anterior of which is bifid below and
narrowed and sickle-shaped above.
Wings hyaline, tinted palely with amber at bases, the forewings tipped with
black for about 3mm. and without pterostigma. Costa reddish immediately
proximad of node ; hindwing with well-formed pterostigma, black, covers 2 to 3
cells ; discoidal cell traversed once in all wings; 5 antenedal nervures, 10
postnodal to. all wings. .
Abdomen bright orange, the articulations narrowly black, as also the ventra
border of segments. Segment 2 with an irregular black apical marking, not
meeting over dorsum but prolonged obliquely forwards on the sides to fuse with
black of ventral border. On segments 3 to 8a pair of minute black dorsal
subapical points; segment 9 with the latero-ventral border extending on to
dorsum especially apicad where it indents the orange ground colour ; segment
10 with its sides and apical border black,
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INDIAN DRAGONFLIES 691
Anal appendages black, shaped as for green1, twice the length of segmeuit 10.
Inferiors only half the length.
Female unknown.
Distribution. Only a single specimen of this insect has been taken within
India limits, this in Mergui, Lower Burma, a male by Mr. Elton Bott, now in
the Pusa Museum. It is not uncommon however in Malaysia and thence
throughout Southern Asia. Williamson reports it from Trong, Lower Siem,
Martin from Tonkin, Laidlaw from Borneo. The markings of the abdomen are
very reduced in the Burmese example. The Siamese example has the base and
sides of segment 1 black, as also a lateral black stripe on segments 3 to 8,
enclosing a basal spot of the ground colour on segments 3to 5 and incomplete
basad on segments 6to 8. There is also an interrupted black stripe on the sides
of segments 2 to 8 in the type from Singapore. Type in Selysian collection.
LIBELLAGINE. LARVAE.
Only three larve of the Lzbellagine, two Khinocypha and one of Microme-
rus, are known. All three agree in everything except minor details so that we
may say for certain that those of Lzdel/ago are not dissimilar.
Very sluggish in habits and cryptic in colouring, they are difficult creatures to
find. They cling to roots or broken-up debris in slow-running streams or at the
bottoms of pools in swifter running ones. 1 have found the easiest way to take
them is to dredge out a quantity of such debris and spread it on the foreshore of
the stream, exposed to the sun. So soon asit has drained and the creatures
have begun to feel the heat of their new environment, will they begin to stir from
their lethargy, and as a consequence, are at once easily detected.
They possess three spine-like caudal gills, which are undeveloped in the
earlier instars. All appear to have banded limbs and most, some cryptic brown
markings on the head, thorax and abdomen. ‘The mask is deeply fissured, the
edges of the fissure overlapping ; the lateral lobes are cheliform and furnished
with along movable hook. The whole mask is of remarkable length, extending
back, when in repose, to the hinder legs.
The gizzard has numerous folds, usually 16 in number, each bearing a row of
4to5teeth. The antenne long and 8-jointed, the basal one very short, the
penultimate of great length.
Autotomy of the caudal gills isaccomplished with the greatest ease, so that it
is difficult to obtain one with all intact; probably these are more for defence
than for respiratory use, and they certainly carry on quite weil without them.
For details of the respective larve, the reader is referred to Plate III.
(To be continued)
THE STUDY OF PLANT LIFE
BY
CHARLES McCann,
Asst. Curator, Bombay Natural History Society
Part fh.
(With lwo Plates and 5 text tigures)
The study of plant life, otherwise known as Botany, is very often
believed to be one of those ‘ dry-as-dust’ subjects which hold
no, or little, interest for the average man and one that is best left to
those cynics who find no pleasure in life; in tact, a botanist is very
often considered a ‘ dry-as-dust gentleman. ’
Just as we meet different types of people, so there are also differ-
ent types of botanists for, like other subjects, Botany is divided into
many different branches. First comes the Systematist who splits
the Vegetable Kingdom up into classes, orders and families and
ultimately into genera and species. To this type of Botanist we
owe the scientific names, and without him it would be almost impos-
sible to carry on, as classification and the fixing of the species must
necessarily form the basis for other phases of botanical work. ‘The
Systematist, while he is able to give us the names, may know
little of the functions and life of the plants he has examined ; this
is the department of the Physiologist who endeavours to explain
the functions of various plants and their organs in the light of
researches made by the Systematist. There are then the Specia-
lists who have an exhaustive knowledge of a single group or genus.
Their knowledge is very deep but often narrowed to the limits of
their special subjects. Lastly we may refer to the all-round botanist
who, although his knowledge may not go so deep as that of the
Specialists, in all likelihood gets more pleasure out of the subject
as a whole, and is better able to see the wonder and infinite variety
of the Vegetable Kingdom and to approach its true position in the
general scheme of lifeon our planet. He will have more ample
opportunity of studying plants in their natural environment, noticing
their habits, likes or disiikes and their special peculiarities which
are as infinitely varied as those manifested in animal life.
The Animal Kingdom, including man with all his achievements,
is almost entirely dependent on the Vegetable World for existence.
The existence of plants does not strike us so forcibly as does that
of animals, perhaps on account of the absence of locomotion
(though locomotion zs present in the lower microscopic forms of
plant life} and other activities similar to those of the Animal
World.
LAL STODY OR PLANS LIFE, 693
From the layman’s point of view, one of the drawbacks to the
study of botany in India is perhaps that the majority of books
available require a fore knowledge of the subject before they can be
used to any advantage. It is this drawback that is mainly respon-
sible for the fact that many in search of a hobby set aside botany
as a subject without vital interest.
This and the following articles are being written with a view to
awaken the interest of my readers to the ‘ little world’ within ours
to which we are so deeply indebted, and to lay before them as far
as possible, the Little Book of Nature from the reading of which so
much interest and pleasure may be derived.
THE CYCLE oF PLANT LIFE.
What is this cycle of plant life and how may we study it?
Some regard the Vegetable Kingdom as a dead domain from which
we learn but little ; but in reality it is a very living one from which
an infinite amount of knowledge may be gathered. Each little
plant has its own little story to tell of its place in nature. The
whole of the Animal Kingdom is dependent on plants either direct-
ly or indirectly for its food and to a great extent for the fresh air
which is so necessary for its existence. We could go on indefinitely
forging the links in that wonderful chain knownas the ‘ Economy of
Life’, but this would take us beyond the scope of the present article
and so only brief reference to these links wiil be made. In the course
of our story I will endeavour to cover the cycle of plant life from the
seed to the mature plant and back to the seed, and to reveal the
sequence of the life history.
To commence this cycle and to read the wordless story which
plants tell us every day of our lives, it would perhaps be best to
start from the seed and follow the development of the plant which
springs theretrom to the stage where the seed is returned by the
plant to earth again. The unanswerable question ‘Which came
first the egg or the hen’ ? or in our case, ‘ the seed or the tree ?’
might perhaps arise in the mind of my readers, but as this question
is just a matter of opinion and conjecture, I leave it to the reader to
decide which he would prefer to have first. For my purpose I
propose to start from the seed.
THE SEED.
What is a seed? To put it bluntly, a seed is that apparently
dead object which when given the requisite amount of moisture, deve-
lops and gradualiy begins to show signs of life—it grows! Where
does the life come from ? The so called seeds of plants contain
the germ (life) which after varied periods of rest (for this they
must have) and under favourable conditions spring into life as it
were, develop into new individuals, thus carrying on the growth of
the species—‘ pay the debt to their ancestors ’ as our Hindu friends
would say. The periods of rest vary, as I have said, sometimes
for many years. Barley and Oats (seeds) last over ten years. Seeds
containing oil lose their power of germination very soon because the
oil becomes rancid; in Willows, e.g., already after a few days.
Some seeds germinate in the fruit, i.e., in some Mangroves
694 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXI/
(Rhizophora). In the soil some seeds are preserved for one
hundred and fifty years. Heldreich has shown that seeds of Silene
(Campion) have retained their power of germination from 1500-
2000 years.
If we examine a seed carefully either with the unaided eye or
preferably with a hand lens, we shall find that it is composed of (1)
an outer skin, the testa or seed coat: (2) the seed leaf or leaves or
cotyledons, which compose the two halves of a seed (taking a pea
seed for example); (3) the germ or embryo which is carefuily
PLUMULE
/ RADICLE
MICROPYLE
SECTION OF DICOTYLEDONOUS SEED.
packed away and protected by the seed leaves. All plants are not
entirely dependent on this mode of reproduction, but it is the
general one among the flowering plants. It might be mentioned
here that the Vegetable Kingdom is divided into two large classes
(a) the Flowering Plants technically known as the Phanerogams ; (6)
the Non-flowering plants the Cryptogams. The last mentioned
TESTA
; an SS eee = - \
ae Sa SEED -~-LEAF
RADICLE '
4
PLUMULE
SECTION OF MONOCOTYLEDONOUS SEED.
contain the ferns, mosses, etc. The Flowering Plants are again
divided into two, the Dcotyledons, those possessing two seed leaves,
LILLE SLOLDY OFF PLAN ELE Ee 695
such as peas, beans and most of the large trees; this class possesses
perhaps the largest number of orders; and the Monocotyledons, those
which have only one seed leaf, such as the Orchids, Lilies, Grasses,
etc. Having seen the great divisions into which the plant world
is divided, which is not altogether out of place here, we shall return
to our story.
As I have said, the production of seed is the general rule but
there are some which propagate their species by a process known as
‘budding’. The Pine Apple, for instance, propagates its species by
producing top and lateral buds which themselves develop into new
plants. These buds are fostered by the parent plant till they drop
off and are then able to look after themseives. There are other
plants again which may be grown from parts of the leaves and
cuttings, such as Begonias. ‘The rose is a familiar example of plants
derived from cuttings, as the cultivated roses rarely seed. In some
plants as soon as apart of the stem or branch reaches the soil, it
develops adventitious roots (of which we shall see more when we
consider the roots of plants) and sooner or later severs all connection
with the parent plant to lead an independent life. The violet isa
good example of this.
Ferns have no visible ‘seeds’, but their place is taken by struc-
tures known as ‘spores’ which are developed in special organs
known as sforang7a on the under surface of the leaves or fronds or
have specially composed organs which bear them. The spores may
be easily seen if one examines the underside of the fronds, they form
little brown, black or yellowish masses distributed over the under-
side. In some cases the masses of sporangia are arranged in
definite patterns. Mosses develop little pin-head like organs, known
as capsules, which contain the spores. It will suffice to say that
most of the lower forms of plants bear spores and that there are
various types or spore-bearing structures to be met with amongst
them.
Having studied the different means of reproduction and propa-
gation to be met with in the Vegetable Kingdom, we shall now
consider the development or germination as it is called of a seed
and follow up its growth step by step until we reach it again at the
top of the tree. Let us suppose that our seed has reached a favour-
able locality and has received a sufficient supply of moisture. If we
watch it closely it will be noticed that it gradually begins to swell
up—the first signs of growth in cases of seeds with a thin seed coat
—till it bursts. The thin testa or seed coat which has up to now
enveloped the cotyledon (seed leaves) casket enclosing the embryo
can no longer contain the growing embryo, so it naturally splits at a
particular spot (of this spot we shall see more later on). The first
evidence of the plant which is emerging from the seed is a tiny white
conical structure—the first root, from which the elaborate root
system will be developed in the future. It immediately begins to
take a downward curve irrespective of the position of the seed before
or after germination. One may prove this by a simple experiment
with a few beans which have been kept in water overnight or until
they germinate. Should the reader however wish to pursue the
experiment further, all he need do is to get an empty cigar box
696 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
and remove one side and substitute a piece of glass sloping from
the top inwards (fig. 1). Fill the box with damp saw dust and place
a few of the beans in it near the top of the glass in different positions.
Through this glass screen the reader will be able to see the root
system as it develops from the germinating seeds, to observe what
are known as the root hairs (the functions of which I will explain
presently) and also how the roots invariably take a directly downward
direction irrespective of the position of their point of emergence
from the seed. The tendency of growing vertically downwards is
known as Geotropism, 1.€., growing towards the centre of the earth.
In many instances soon after the root has appeared and attached
itself to the substratum, the seed lifts itseif above the surface of
the soil, casts the testa and unfolds the two halves of which the seed
was composed, i.e., the seed leaves or cotyledons. Between them
will be seen a minute pair or more of other leaves, beautifully
folded, which will develop in course of time. In some cases the
seed-leaves remain thick and fleshy, while in others they expand and
take the form and colour of ordinary leaves, though generally very
different in shape to the leaves that follow. When they first appear
above the surface of the soilthey are white but as aresult of the action
of light they become green. The cause of this will be explained
when we take up the subject of the leaves. Seed-leaves are heavily
charged with reserve food for the young plant and act in the same
way as the yolk of an egg. After the leaves have appeared and
taken on the duties of supplying part of the necessary food, the seed-
leaves shrivel up and fade away, their duties being ended. The
vegetative part or shoot always grows upwards or towards the
source of light. This tendency which is just the opposite to that of
the root, for roots grow away from light, is known as Heliotropism,
i.e., growing towards the light or sun. A very simple experiment
will prove this :—Place a few developed seedlings in an absolutely
dark box with a small hole bored somewhere low down on one side
to let in light. After a day or two open the box and examine the
seedlings and it will be noticed that the plants have changed their
direction of growth from the upright towards the source of
ligt; ion .23
THE STUDY OF PLANT LIFE 697
As breathing is common to all living things when in an active
state (for breathing is almost at a standstill during hibernation),
it might be asked, do seeds |
breathe? The answer is not
diffcult—though in the case
of the seed it may not be
considered as breathing in
the strict sense of the word.
Breathing as is generally
understood is an interchange
of gases, usually through
organs specially adapted for
the purpose. Animals give
off carbon dioxide (Co,) and
inhale oxygen by day as well
as by night, plants on the
other hand, while they follow
the same process by night,
reverse it by day breathing
in carbon dioxide and exhal-
ing oxygen. Seeds give off Fic. 2.
Co, especially at the time of
germination. There isa tiny aperture at one end of the seed, known
as the micropyle through which this breathing, soto speak is carried
on. Itis here that the radicle or first root breaks through the testa.
The prompt extinguishing of a lighted taper inserted into a
bottle in which germinating seeds have been enclused, indicates
that the seeds have exhausted or breathed in the oxygen contained
in the bottle. Similarly the heat, the oppressiveness ard the
tendency to suffocation on entering a closed room in which grain is
stored is due to the breathing of the seed and the accumulation of
gases in the room.
The seeds of Monocotyledons, as has been said before, possess
only one seed-leaf and this does not as a rule appear above the sur-
face of the soil but nourishes the young plant from where it lies.
Now that we have seen the chief parts of whicha seed is compos-
ed, viz.: the testa, the seed-leaves and the germ or embryo, and
superficially studied its development, we shall proceed to follow up
its growth from the root and gradually climb up the tree till we
return to itand complete the cycle. The development of the lower
types of plants from the spores, we shall consider in a future
article.
Tam AINKoOon.
To go on with our story, we commence by carefully examining
the first root or radicle which, as I have already indicated, shows
itself first and appears as a white or creamy thread-like process
which descends into the earth. On appearance it is smooth but it
soon develops fine hair-like structures known as the voot-hairs. Root-
hairs do not grow all over the root, but are confined to the growing
end. As the root elongates the upper hairs are lost and are re-
placed by new ones near the growing tip. From the radicle
11
698 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
the great root systems of large trees are developed. As the root
branches in various directions, the growing extremities of each branch
are also clothed with root-hairs. The older parts of the roots become
darker in colour and
thicker in proportion,
assuming acorky cover-
ing to protect them. It
LEAVES may be asked, what are
; the root-hairs and what
useful purpose do they
serve ? Root-hairs are
unicellular prolongations
which are very essential
STEM to the life of every plant:
they are the ‘ procura-
tors,’ as it were, of a
great quantity of the
SEED-LEAF mineral food and
moisture thatis neces-
Sary to the plant. They
go into the microscopic
crevices between’ the
ROOT particles of soil and take
up the available moisture
and the particular salts
in solution in the soil
that are necessary to
RCOOT-HAIRS their growth. They do
not! sabsorb = Salleesthe
mineral salts which are
present inthe soil, but
A SEEDLING OF A BEAN SHOWING THE PARTS HENS ake Pe oO Sele:
Ge ears acc tion. An acid secretion
which exudes from the.
root-hairs is powerful enough to dissolve such salts as are not
soluble in water and to absorb them. At the same time this acid
secretion also helps the roots to push their way through hard soil
and even rock by disintegrating it, though: as arule the roots foliow
the line of least resistance.
The root tip which is the growing part is very delicate and tender
and is covered by a special structure the Roof-cap which protects the
growing end onits way through the soil. The root-cap consists of
layers of celis which are thickened to form this protecting cover.
As the outer layers of this root-cap are destroyed, new layers are
being formed on the inside. This root-cap is best displayed by the
Palms and especiaily in the case of the roots of the Screw-pine or
Pandanus which is fairly common along our shores, from the flower
of which the Keora scent is extracted. The part of the root which
is protected by the root-cap is devoid of root hairs. The region
which is covered by root-hairs is known as the ‘ root-hair region.’
As I have said before the root-hairs secrete an acid. Ifa piece of
polished marble was placed at the bottom of a box in which some
ye
THE STODY (OF PLANT: LIFE . 699
beans were allowed to germinate in a covering layer of saw dust, it
will be seen after a suitable lapse of time that the growing roots on
encountering the resistance of the marble in the downwaid course
have not only spread all over its surface but have left as a result of
the acids given out by them, a distinct impression of their outline
upon its polished surface.
Not the whole of the root is capable of elongation but the growing
part is restricted to about j;th of an inch from the tip. Theroot tip
is sensitive to various stimuli and will react accordingly. Water is
the chief attraction and roots may go many yards to obtain it. When
water is deep below the surface of the soil, roots will go down deep
but when it is near the surtace they spread out near the -surface.
The root systems in plants vary according to the amount of mois-
ture required and are very often governed by the position of the
leaves. In some plants the leaves are so arranged that when the
rain water or dew falls on them it is directed towards the stem ard
thence to the roots. In such cases the root system is restricted toa
very small area and very often consists of only a taproot or main
root. Good examples of these plants are Cannas, Lilies, etc., which
require a good supply of water at their particular seasons. In others
the branches are spread out and the moisture falls to the ground
from the tips of the leaves; the moisture is thus spread over a
greater area. Naturally the root system of these plants is in pro-
portion tothe crown. Most of the large trees are good examples of
this type of root system.
We have examined the root hairs and root cap; let us now have
a glance at the older portions of the root which were left behind by
the growing ends and have had time to enlarge. These parts have
long since lost their root hairs and thus no longer have the power
of absorption. Nevertheless they still continue to grow but not in
length. At first they become discoloured and put on an outer layer
of protective armour—cork—which is secreted by the outer cells and
which gradually bscomes thickerin order to protect the internal
‘channels’ which convey the sap* from the root hairs throughthe stem
to the leaves. They also increase in girth and become more and more
rigid to be able to support the tree. What purpose do roots serve?
Besides being the ducts through which the soluble food and moisture
is passed on to the stem and leaves they serve the purpose of ancho-
rage—of keeping the tree upright and preventing itfrom being blown
down by winds and storms to which it will be subject in the future.
These are the two main functions of the root, but they are not all.
They are also used as ‘ store-houses’ for surplus food and moisture
which is accumulated during a favourable season and enables the
plant to endure through the long periods of draught or to serve it
through its periods of rest—that is to say that they come up during
One season, flower, fruit and accumulate reserve material and then
die down again to the surface, of the ground only to reappear next
season. Accordingiy plants are classifed into different groups.
Plants which develop very rapidly during a favourable season from
seed and die down completely after producing seed are known as
1 The word ‘sap ’ is used here in its generally accepted meaning.
700 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXI1
Annuals—living but one season. Plants whose life is prolonged be-
yond a single year or over an indefinite number of years are called
Perennials. We know now that roots perform the chief functions, (1)
of procurators of food, (2) of anchorage, (3) of store-houses and
reservoirs. <A brief survey of different types of roots to be met
with and the varied shapes they assume according to the require-
ments of a particular species, will not be out of place here.
Among plants belonging to the Dicotyledons, the radicle or tap
root grows and gives off branches and these branch again in turn;
with the Monocotyledons, on the other hand, the radicle is short-
lived. Roots that appear from other parts of tne plant and are known
as adventitious roots. These are slender and thread like and are
spoken of as fzbrvous roots.
We find this type of root most common among all the Monocoty-
ledons, such as Grasses, Lilies, etc. Adventitious roots are also to
be met with among the Dicotyledons of which the Banyan (/icus
bengalensis) and others are good instances.
As we have seen, roots may act as a store-house and in conse-
quence assume enormous proportions and various shapes. ‘Thus we
have roots whose names are descriptive of their shape. The Carrot
is a good example of a conical root serving the purpose of storage,
the Beet assumes a spindle shape, while the NKRaddish is either
conical or globulur. Dahlias have swollen or tuberous roots. These
are a few examples among many which might be indicated here.
All these queer shapes originate from fibrous roots. Care must be
exercised however not to confound tuberous roots with a tuber, for
example the common potato, which is a fleshy underground stem
ana MOt a Tootat all.
An iodine test will show that such roots contain a high percentage
of starch, as iodine stains starch a bluish-purple. Beet contains
cane-sugar, while the roots of the Dahlia containinulin. The value
of these roots to the Animal Kingdom and man in particular is
readily apparent. .
As a protection against the depredations of field-mice, insect
larve and other underground animals, many food-storing roots
develop poisonous and nasty substances in their tissues, in the way
of noxious alkaloids, foetid gum, resins and other products well
known to druggists; it has been observed that such roots are very
seldom attacked.
Frequently plants produce more than one type of root, serving
different purposes, some being fibrous, absorbing roots, others
swollen and charged with reserve food. Some roots after greatly
elongating, contract and pull the stem lower down into the soil, such
as the Lilies and other familiar plants which have no apparent
stem but grow with their leaves in rosettes adpressed to the
ground.
Roots often go to avery great depth into the soil, much deeper
than people imagine, especially in hard and dry subsoil, in the quest
for water. Evenin rich garden soil the roots of wheat have been
traced to a perpendicular depth of five or six feet. This is however
trifling when compared with the depth to which the tap-root of a
Baobab-tree (Adansonia digitata) descends. Anderson in his account
Journ., Bombay Nat. Hist. Soc.
Ce 2
DIFFERENT TYPES OF ROOTS
Testicular root. 6.
Napiform root. he
Nodulose root. or
Moniliform root. oh
Globular root. 10.
Fibrous roots.
Tuberous facicular root.
Thickened Tap-root.
Annulated roots.
Fusiform root.
Journ., Bombay Nat. Hist. Soc.
ANOTHER TYPE OF ADVENTITIOUS Root.
THE
‘Screw PINE’ (Pandanus Sp.)
THE STUDY OF PLANT LIFE 701
of Senegal, records the root of one of these trees to have reached a
depth of one hundred and ten feet.
Then there are what are known as aerial roots. These are
exemplified by most of the epzphytic orchids, i.e., growing on trees
but not obtaining their nourishment from them. There are other
plants which have aerial roots such as the Banyan, already referred
to and also the Pandanus, but these answer mainly the purpose of
support.
The roots of many orchids are feed/ng aerial roots. They are
modified for absorption of nutriment from the surrounding atmos-
phere ; in some cases the orchids have no leaves, the roots obtaining
for the plant all that is requisite for its nourishment and support.
The roots hang downwards like white-threads or cords, the whiteness
being due to a papery membrane which covers the green chlorophyll-
containing cells of the true root. The envelope consists of
perforated cells and acts like a sponge. When they come into
contact with water they absorb it immediately until they fil out.
The inner living green cells are thus surrounded by a fluid envelope
and are able to obtain from it as much moisture as they need.
Besides, this sponge tissue has the power of condensing aqueous
vapours and other gases, so that the plant is entirely indepencent
of its host for nourishment. Trees growing in marshy ground also
send up aerial roots in order to obtain oxygen which is absent in the
water-logged soil. Familiar examples ot these roots are to be met
with in the different species of mangroves which are so common
along the marshy shores of the Islands of Bombay and Salsette.
Some large trees develop large root-expansions as additional]
supports and these are known as duttvess roots. These roots stand
out like great wings round the base of the tree and their develop-
ment is so enormous at times that a man standing between two of
these buttresses would be effectively screened.
The roots of some aquatic plants dangle in the water from which
they derive their nourishment, while others contain large air-spaces
which aid the plant to float.
Roots possess one almost constant character and that is, that they
produce members like themselves, i.e. root-branches, while the
stems produce dissimilar members—leaves. However, it frequently
occurs that roots giverise to leafy shoots—at times a boon to garden-
ers and at others acurse, especially in the case of weeds. It often
happens that the seemingly complete eradication of weeds in a
garden is followed after a few days by fresh outcrops and the area
that was apparently cleared is covered with weeds again much to
the disgust of the gardener. The reason for this is that part of the
roots were left behind in the soil and having the power to send up
shoots have covered the ground once more. ‘These shoots are
known as adventitious shoots or suckers.
There still remains one type of root, i.e., the roots of parasitic
planis. These plants are entirely dependent on their host for their
nourishment and send their roots into the food-canals of the branches
or roots of their host and suck them dry. A good example of this
is the Loranthus, a near relative of the Mistletoe (Viscum album )
which is commonly to be seen in great clusters on mango trees
702 JOURNAL, BOMBAY: NATORAL HIST, SOGIETV. Vali2eods
and others. This plant supplies us with an extremely interest-
ing example of the motility of young roots. The fruit contains
birdlime—-a peculiar viscous, tenacious and elastic substance—and
when the berry falls away it adheres to whatever it falls upon. The
seed is embedded in the viscid pulp and germinates commencing in
the following way. ‘ The radicle, ’ says Mr. N. 5. Brown, “at first
grows out, and when it has grown to about an inch in length, it
develops upon its extremity a flattened disc; the radicle then curves
about until the disc is applied to any object that is near at hand.
If the spot to which the disclias fastened is suitable, the germination
continues and no locomotion takes place ; but if the spot should not
be a favourable one, the germinating embryo has the power of
changing its position. This is accomplished by the adhesive
radicie raising the seed and advancing it to another spot, or to make
the purpose plainer, the disc at the end of the radicle adheres very
tightly to whatever it is applied to; the radicle itself straightens
and tears away the viscid berry from whatever it has adhered to and
raises it inthe air. ‘The radicle then again curves and the berry is
carried by it to another spot, where it adheres again. This disc
then releases itself, and by the curving about of the radicle is
advanced to another spot, where it again fixes itself. This,
Dr. Watt says, has been repeated several times, so that to a certain
extent the young embryo, still within the seed, moves about. It
seems to select certain places in preference to others, particularly
leaves. The berries on falling are almost certain to alight upon
leaves, and although many germinate there : they have been observed
to move from the leaves to the stem, and finally fasten there.
(Gardener’s Chronicle, 1881).
It might have been better perhaps if we had considered the sap
of plants at an earlier stage, but before we leave the roots we must
elaborate a little more on their functions as procurators of part of
the food of plants. Sap is the fluid which the roots absorb from the
soil or the leaves from the air and which contains in solution the true
nutritive principles. Water is the main substance of the sap. It has
been estimated that for every two hundred grains of water absorbed
and exhaled by a plant only one grain of inorganic matter has been
taken up and for every 2000 grains of water consumed one grain of
inorganic substance has been appropriated. Young and growing
parts of plants are well supplied with sap which fills up every avail-
able space in the plant-tissues which is not already occupied by
other substances. It is conveyed into the plant by the root hairs,
and does not undergo any marked change till it reaches the leaves.
Sap may be acid or sweet, clear or turbid, nutritious or otherwise so
that its value from an economic point of view is considerable. The
acid taste of most unripe fruit is due to the sap. Citric acid, the
most common form is found in the juice of lemons, oranges and
many of the commonest fruits, and it often exists together with other
acids, such as malic acid in the gooseberry, currant and several others.
The latter acid is found alone in apples and pears. It is due to the
action of these acids that knives are stained when used on various
fruit. Itis perhaps due to the presence of these acids that young
fruit are protected from the attacks of birds and animals which feed
LAE SLTODY OF LELAND LLRE 703
on the ripe fruit which is one of the factors in seed dispersal. Birds
and animals which eat the ripe fruit containing the fully developed
seeds often pass them out in their excreta sometimes miles from the
place where they were eaten. The seeds are usually undigested
and thus help nature in the dispersal of seed and the propagation of
the species in other localities.
In some cases the acid juices of plants destroy or deadenone’s sense
of taste if masticated, and this state may often last for hours accord-
ing to the quantity eaten. In some cases sugar tastes like sand in
the mouth after taking some of these plant acids by chewing the
leaves, flowers or seeds. A sweet orange under the same conditions
will taste sour ; the citric acid alone remains distinguishable ; sweet
and bitter tastes being destroyed. After a good dose of the leaf of
a tropical Asclepiad known as Gymanema sylvestre, quinine tastes like
chalk.
The sweetness of the sap of the Beet-root is due to the presence
of canose or cane-sugar dissolved init. It was first discovered by a
Prussian chemist named Margraff (about 1747), butit was not till
1809 when Napoleon forbade the importation of cane-sugar into
France from the West Indies, that the discovery was turned to prac-
tical account. An Imperial sugar factory was then established at
Rambouillet ; pupils were instructed in the process, premiums were
offered for the best samples of the sweetener, and in the course ofa
few years the manufacture of beet-sugar was prosperously set on
foot.
Cane-sugar is abundantly found in sugar-cane and the sugar-
maple. Itis also found in the nectaries or honey-secreting glands of
flowers where it furnishes the substance from which the bees gather
their honey.
It is secreted by the nectaries and the quantity is at its fullest
during the emission of the pollen but stops soon after fertilization of
the flower.
A careful distinction must be made between cane-sugar and
grape-sugar or glucose. These are differently produced by the
plant and when treated with the reagent known as Fehling’s solution
react differently—glucose gives a bulky yellow deposit which canose
does not.
Besides these juices there are the milky ones exuded by many
plants especially the milkworts (Euphorbias). These milky juices
are known as latex. Strange though it may seem, this milky juice
is clear while in the uninjured tissues but instantly becomes turbid
on exposure to the air. The colour of latex is generally white, but
yeilow, red and in rare. cases blue milk-saps are met with. The
microscope shows that it consists of a colourless fluid in which float
myriads of minute globules which give the sap its opaque appear-
ance. The common Euphorbia (often erroneously called Cactus)
frequently used as a hedging, is a good example of this type of
plant. India rubber which is obtained from a tropical American
tree (Hevea brasiliensis) and other trees, and opium from the opium
Poppy (Papaver somniterum) are dried juices of such plants.
(To be continued. )
NOTES AND DESCRIPTIONS OF FISHES FROM CEYLON
BY
HENRY W. FOWLER
OF THE ACADEMY OF NATURAL SCIENCES OF PHILADELPHIA
(With 2 Plates)
The collection reported below was purchased by the Academy of Natural
Sciences of Philadelphia in 1924 fron Dr. F. Hallberg, of Géteborg, Sweden.
All were obtained at Colombo. Several of the species are of special interest
and one appears to be new to science.
ENGRAULIDZ®
Engraulis commersontit (Lacépéde)
Depth 42 ; head 4, width 24. Snout 43 in head ; eye 3$, covered with adipose
membrane, greater than snout or interurbital ; maxillary reaches gill opening,
x in head; row of fine teeth on each maxillary and narrow bands on vomer
and palatines ; interorbital 4, slightly convex. Gill rakers 19+ 21, lanceolate,
greater than gill filaments or 1? in eye. Scales very caducous, mostly fallen,
33 (pockets) in lateral series from shoulder to caudal base, 9 transversely, 21?
predorsal : vertical strize 7 ; pectoral axillary scale 3 of fin, ventral scale § ; no
abdominal scutes ; venulose area at shoulder moderate. Dorsal 111, I, I insert-
ed little nearer caudal base than front eye edge, third simple ray 12 in head ;
Analtti, 17, I, inserted opposite last 2 in length of depressed dorsai, first
branched ray 1,%; caudal 32 1n combined head and body, forked; least depth
of caudal peduncle half its length or 2g in head; pectoral 14; ventral 2.
Largely pale brown. Faint silvery lateral band, less than eye in width, widest
on caudal peduncle. Iris pale slate. Dorsal and caudal gray, other fins
whitish.
One 110 tam
TACHYVSURIDZ
Hemipimelodus dussumtert (Valenciennes).
Depth 53; head4, width 1g. Snout 34 in head ; eye 4, 1} in snout, 1% in inter-
orbital ; mouth width 2,, in head ; band of villiform teeth in each jaw, palate
and tongue edentulous; maxillary barbel reaches # in pectoral spine, outer
mental 4 in pectoral spine, inner mental barbel not quite to pectoral spine ;
internasal space slightly less than interorbital, nostrils large, front one close to
edge of snout and posterior about midway in snout as seen in profile; inter-
orbital 22 in head, slightly convex ; occipital fontanel broad, reaches tu base of
occipital bony plate, which about over gill opening; occipital buckler with
median ridge, wide as long, long as snout. Gill opening with slight median
notch, free from isthmus. Dorsal I, 6, spine adipose like for greater portion
terminally, equals head ; adipose fin large, slightly over half of head length ;
Anal vy, II, first branched ray 12; caudal 1, forked; least depth of caudal
peduncle 2z in its iength or 3; in head; pectoral 1%, spine slightly over half
length of fin; ventral]. Back and head above pale gray brown, beiow
whitish. Snout, occipital fontanel and under surface of head whitish. Iris
leaden gray. Barbels dusky, pale basally. Fins all pale or whitish, upper
front edge of spinous dorsal or dorsal spine dusky. Caudal with gray terinin-
ally.
One 47mm. Evidently the young of this species, which changes considera-
bly in proportions with age.
NOTES AND DESCRIPTIONS OF FISHES FROM CEYLON 705
HEMIRAMPHIDZ
Hemiramphus neglectus Bleeker
Depth 83 from upper jaw tip, body subcylindrical ; head 42, width3. Snout
2 in rest of head, free portion little broader than long, width in snout to eye 14;
eye 3in head from snout tip, 1; in snout, slightly greater than interorbital ;
mouth cleft 2 to eye, lower jaw before snout tip 4 to caudal base ; maxillary # to
eye, vertical expansion slightly less than pupii, length 3? in head from upper
jaw tip ; interorbital 33, slightly depressed ; postorbital 2 of eye. Gill rakers
9-+ 19, lanceolate, equal gill filaments or 3 in eye. Scales 47 from above gill
opening to caudal base and 3? more on latter, predorsal 40 ?; all little adherent,
most having fallen ; basal radiating striz 3, circulimoderately small. Dorsalt,
13, first branched ray 24 in head from snout tip; Anal1, 13, first branched ray
3; caudal I, moderately forked or emarginate, lower lobe longer; least depth
of caudal peduncle 14 its length or 44 in head ; pectoral 14 ; ventral 2}, inserted
little nearer head than caudal base. Faded pale brownish, largely uniform.
Predorsal with 3 median close set parallel dusky lines. Narrow dusky band
from over gill opening to caudal base, nearly wide as pupil on tail aboveanal,
its lower edge with darker bordering line. Snout above and occiput with close
set dusky to blackish dots. Beak blackish-brown, continued back along each
side of mandible as blackish brown streak nearly to eye. Iris slaty. Fins all
pale. Dorsal and anal tinted with gray brown.
One 135mm. Also one for comparison from Padang, Sumatra, 136mm. It
differs in Depth 83; head 41; snout 23; eye 3, ; interorbital 4; gill rakers
6-+11; Dorsal 11, 13; Anal 1, 15.
Certainly very close to Hemiramphus unifasciatus (Ranzani) from the
Tropical Atlantic. Comparison with examples of this latter from Porto Rico
shows the American form with a shorter preorbital, which also a little wider
than long.
Zenarchopterus dispar (Valenciennes)
Depth 74 from upper jaw tip, body slightly compressed ; head 3g, width 2%.
Snout 1,% in rest of head, free upper portion broader than long, width in snout
to eye 12; eye 4 in head from snout tip, 13 in snout, 1§ in interorbital ; mouth
cleft 2 to eye, lower jaw before snout tip 23 to caudal base; maxillary largely
concealed, # to eye, nearly vertical, expansion ? of pupil, length 3 in head from
snout tip; interorbital 3, level. Gill rakers 6-+ 12, lanceolate, long as gill
filaments or 32 in eye. Scales 42 in lateral series from above gill opening to
caudal base and 3 more on latter, 4 above lateral line, 2 below, 38 predorsal ;
few basal scales at front of anal; 5 or 6 basal radiating strie marginally and
circuli fine. Dorsal 12, first ray 3} in head from upper jaw tip; anal I, 8,
first branched ray 2}; caudal 13, truncate; least depth of caudal peduncle 1§
in its length or 4 in head from snout tip; pectoral 12 ; ventral 33. Back brown,
with median dusky line on predorsal. Underlaid gray brown line from
shoulder to caudal, greatest width about 2 of pupil along side of tail. Beak
blackish. Snout dusky. Iris slate. Vertical fins grayish, more or less dusky
terminally. Paired fins pale.
One 150 mm.
Cypselurus hallbergi, new species
y. s
(Plate 1)
Depth 57; head 44, width 12. Snout 34 in head from snout tip; eye 33,
with free lids, 1} in interorbital ; maxillary concealed, # to eye, equals snout ;
teeth minute, sparse, form slight band in each jaw, none on palate or tongue ;
interorbital 2; in head, very slightly concave. Gill rakers 7 + 21, lanceolate,
equals gill filaments or 1$ in eye. Scales slightlv adherent, rather well
exposed ; 47 between gill opening above and caudal base and 3 more on
latter ; 8 above lateral line, 3 below to anal origin ; predorsal 30, extending
forward little before nostril; 5 or 6 basal radiating stria, edge lobate and
circuli finely concentric. Lateral line not reaching caudal or not extending
beyond tip of last anal ray ; tubes slender, simple. Dorsal1, II, 1, first ray
4 in total head length; Anal 1, 9, I, first branched ray 3%, fin inserted
opposite base of second branched dorsal ray ; caudal widely forked, upper
lobe 13 in that of lower which 34 in combined head and body ; least depth of
12
706 JOURNAL; BOMBAY NATORAL AIST. SOGIETY, Vol Xed7
caudal peduncle 12 or 3 in head ; pectoral not quite reaching caudal base, 13
in combined head and body, rays Iv, 13; ventral I, 5, spine nearly third of fin
which reaches back far as pectoral (right fin shorter), length 2¢ in combined
head and body, its insertion midway between hind eye edge and caudal base.
Back deep brown, lower sides and under surface whitish. Iris slate. Lower
jaw pale. Dorsal and caudal pale brown. Anal whitish. Pectoral neutral
brown medially, terminally dusky, pale basally and lower basal margin
whitish. Ventral neutral brown medially and terminally, with broad inner
whitish border, base and spine also pale.
One example, the type, 195 mm. (lower caudal tip damaged so estimated
at 207 mm.). Differs from all the species of the genus I have examined in the
increased rudimentary or simple rays in the pectoral and also in the increased
gill rakers. Apparently resembles Cypselurus simus (Valenciennes), but
unlike any I have seen in the above respects. It is also suggestive or
Cypselurus oxycephalus (Bleeker), but that species falls within the subgenus
Exonautes as the anal origin is opposite that of the dorsal.
I take pleasure in dedicating this species to Dr. F. Hallberg who collected
the type.
ANABANTIDZ
Anabas testudineus (Bloch)
One 70mm. Small dark spot on opercular flap, less than pupil. Caudal
base with small vertical brown bar.
ATHERINIDZ
Atherina duodecimalis, Valenciennes
(Plate I)
Depth 5%; head 3%, width 1$. Snout 3% in head from snout tip; eye 23,
greater than snout or interorbital ; maxillary reaches first fifth in eye, 23 in
head ; teeth small, conic, in bands in jaws, on vomer and palatines ; mandi-
bular rami moderately elevated inside mouth ; interorbital broadly concave,
2} in head. Gill rakers 5 + 21, lanceolate, slender, 23 in eye. Scales 38 from
above gill opening to caudal base and 4 more on latter, 7 transversely, 16
predorsal, 5 behind depressed spinous dorsal ; single row of scales on cheek ;
large scale below pectoral base nearly large as eye; scales with 3 basal
marginal points and 30 basal, vertical, parallel strie, apical region smooth
and with 18 small inconspicuous marginal lobes. Dorsal V—i, 10, 1, second
spine 2; in total head length, first branched ray 12 and spine of soft fin short
or 4 of first ray ; spinous dorsal origin midway between eye center and caudal
base; anal base about equals its distance from caudal; Anal I, 12, first
branched ray 12 and spine fits length; caudal equals head, well forked,
lower lobe little longer; least depth of caudal peduncle 33; pectoral 1¢;
ventral 1{. Uniformly pale brownish, with more or less whitish or silvery.
Top of head, end of mandible and snout dusted with dusky brown. Patch
of dusky dots on middle of opercle. Each scale of back above with submiargi-
nal are of dusky brown dots. Narrow band of dusky dots along scales at
base of anal. Gray band from pectoral axil to caudai base medially, little
less in width than pupil at greatest expansion which opposite soft dorsal
and anal. Fins all pale, with arc of dusky dots at pectoral base.
One 70 mm. Interest attaches to this specimen as the species was originally
described from Ceylon and that it does not appear to have been obtained there
since. ‘The type is very briefly described with Dorsal V—I, 9 and Anal I, 12, its
length 76 mm. Bleeker’s materials, as given by Weber and Beaufort are
apparently the same. It is nota Hepsetza, however, as catalogued by Jordan
and Hubbs. No previous writer has yet described it completely and as it has
not been figured the accompanying drawing is offered. The large infrapectoral
scale, the position of the vent at the first 2 of the ventral fins and black in color,
will serve to distinguish it from allied species as Atherina forskali, Riippell and
Atherina lacunosa, Schneider. Ogilby has indicated a large infrapectoral
scale for the last named, .
PLATE |
Bombay Nat. Hist. Soc.
Journ.,
Cypselurus hallbergi, New Species.
Henry W. Fowler del.
PLATE II.
Journ., Bombay Nat. Hist. Soc.
Valenciennes.
Atherina duodecimalits,
Henry W. Fowler del.
NOTES AND DESCRIPTIONS OF FISHES FROM CEYLON 107
MUGILIDA
Mugil longimanus, Gunther
Depth 32; head 33, width 13. Snout 4; eye 3%, greater than snout, 1¢ in
interorbital ; maxillary concealed, reaches eye, 33 in head; mandibular angle
forms broadly obtuse; lower preorbital edge finely serrate. Scales 31 from
above gill opening to caudal base, 12 transversely at soft dorsal, 25 predorsal, 5
behind depressed spinous dorsal; soft dorsal and anal covered with fine scales ;
axillary pectoral scale 3 of fin; scales with 6 or 7 basal radiating strie, no
apical points. Dorsa! VI—I, 8, I, first spine 1} in head, first branched ray lf ;
Anal Ill, 9, 1, first branched ray 12 with third spine 2 its length ; caudal lI,
deeply emarginate; caudal peduncle depth 23 ; pectoral 1;4,, reaches little beyond
dorsal origin ; ventral 13. Gray brown above, silvery white below. Dorsals,
caudal and pectorals with gray brown, last with small diffuse dusky spot at
origin. Hind caudal edge dusky. Lower fins whitish. Iris slaty.
One 91 mm.
CARANGIDA
Caranx tgnobilis (Forskal)
Depth 22; head 2,%, width 24. Snout 32 in head from snout tip; eye 374,
with narrow adipose lid behind, subequal with snout or interorbita] ; maxillary
reaches eye center, length 24 in head; interorbital 4, convex. Gill rakers
§ + 15, lanceolate. Scales 43 + 31, arch 12 in straight section of lateral line,
14 above, 22 below, scute depth 1¢ in eye; breast naked below to ventral,
except small patch of small scales posteromedially. Dorsal I, VIli, I, 19, 1,
third spine 22 in total head length, first branched ray 2, ; Anal fl—I, 16,
I, second spine 1} in eye, first branched ray 24, in head; caudal 13,
strongly forked; least depth of caudal peduncle 1#; pectoral 1,, reaches
fourth scute of lateral line; ventral 22. Back brownish, below silvery white.
Iris slate. Spinous dorsal dusted dusky. Soft dorsal and caudal with pale
brownish, other fins whitish.
One 94 mm.
Caranx sexfasciatus. (Quoy and Gaimard)
Depth 23 ; head 3, width 2g. Snout 33 in head from snout tip ; eye 4, adipose
lid covers last sixth of eye, subequal with snout or interorbital; maxillary
reaches opposite eye center, expansion l?in eye, length 2,4, in head ; row of
short conic teeth in each jaw and narrow inner band with same, also narrow
bands of very small teeth on vomer and palatines and tongue ; interorbital 4,
convexly elevated; preopercle flange with radiating and marginal venules.
Gill rakers 7+15, lanceolate. Scales 43-++-31, arch 12 in straight section of
lateral line ; scute depth 1§ in eye; breast densely scaled. Dorsal I, Vili, I, 20,
1, fourth spine 23 in total head length, first branched ray 1,9, ; Anal H—I, 16, 1,
second spine 5}, first branched ray 234, ; caudal 1}, strongly forked ; least depth
of caudal peduncle 6g; pectoral 1,4, reaches fourth scute of lateral line ;
ventral 2}. Back brown, below whitish. Back and sides with 6 or 7 broad
obscure darker vertical bands, broader than interspaces. Iris slate. End of
mandible brownish. Spinous dorsal neutral dusky. Soft dorsal and caudal
with pale dusky marginally, otherwise pale or whitish like other fins.
One 105 mm.
LEIOGNATHID
Letognathus splendens (Cuvier)
Three 41to 51mm. These agree with Day’s figure, though as they are
young the back is with Jarge diffuse dull brown blotches. Black blotch inclu-
ding 3 highest membranes of spinous dorsal terminally greatly contrasted. All
show a neutral dusky basal line along dorsals. Lower half of body sprinkled
with dusky or blackish dots. Iris slate.
708 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol, XXXII
CHEILODIPTERIDZ
Amia endeketenia (Bleeker)
Depth 3; head 3, width 25. Snout 4 in head from snout tip; eye 3}, greater
than snout or interorbital ; maxillary reaches opposite eye center, expansion
4 in eye, length 24 in head; bands of minute teeth in jaws, on vomer and
palatines ; interorbital 4¢ in head, nearly level; preopercle ridge entire, edge
finely serrate. Gill rakers 5 -+ 15, lanceolate, 24 in eye. Scales 21 in lateral
line to caudal base and 2 more on latter, 2 above, 6 below, 6 predorsal to
occiput, 2 rows on cheek ; 10 to 13 basal radiating striz, apical denticles 30 to
32, weak, small, in 3 transverse series and circuli moderate, Dorsal VI — I, 9,
second spine 1# in total head length, first ray 1§ with spine of soft dorsal 13
in first ray ; Anal II, 8, second spine 2%, first ray 12; caudal 14, forked; least
depth of caudal peduncle 23; pectoral 1%; ventral 1. Faded pale brown
generally. Traces of 4 dusky longitudinal narrow bands; second begins on
side of snout, crosses eye and postoctlar, where most distinct. Iris slate. Back
and head above with dull brown shades. Fins all more or less pale. Spinous
dorsal dusky terminally. Soft dorsal and anal each with diffuse dusky
subbasal longitudinal band. Round blackish spot little smaller than pupil at
caudal base medianly. Caudal margined above and below with pale dusky.
One 54mm. It differs a little from Day’s figure in that he shows the second
dorsal spine but half length of third, whereas in mine the second and third
spines are subequal.
AMBASSIDZ
Amobassis satgha (Forskal)
Depth 23; head 23, width 24. Snout 42 in head from snout tip; eye 3,
greater than snout or interorbital ; maxillary reaches opposite front pupil edge,
expansion 3 in eye, length 2} in head: interorbital 5, very slightly convex ;
preorbital edge entire, lower edge with 7 serre ; lower ridge and edge of
preopercle serrate. Gill rakers 9 + 17, lanceolate, slender, 1¢ in eye. Scales
30 in incomplete lateral line to caudal base and 3 more on latter, 4 above, 7
below, 22 predorsal, 3 rows on cheek ; 1 to 3 basal radiating striz, no apical
denticles and circuli moderate. Dorsal VII, 1,9, 1, second spine 13 in total
head length, first ray 24; Anal III, 9, 1, third spine 23, first ray 1, ; caudal
1,4,, deeply forked, lobes slender and pointed ; least depth of caudal peduncle
33; pectoral 14; ventral 123. Uniform pale brown, scales along edge of
predorsal and about back posteriorly above with deeper brown dustings.
Mandible and upper jaw brownish. Iris slate. Lateral gray line along tail
and caudal peduncle medially faint. End of second membrane of spinous
dorsal dusky, fins otherwise all pale.
One 88 mm.
Ambassis gymnocephalus (Lacépéde)
Depth 2#; head 23, width 22. Snout 4 in head from snout tip ; eye 33,
greater than snout or interorbital; maxillary reaches first fourth of eye,
expansion 32 in eye, length 2% in head ; interorbital 43, very slightly convex ;
preorbital ridge and edge, also lower ridge and edge of preopercle serrate.
Gill rakers 8 + 21, lanceolate, slender, 12 in eye. Scales 11 + 10 in interrupted
lateral line to caudal base and 2 more on latter, break below front of soft
dorsal, 3 scales above, 6 below, 17 predorsal, 2 rows on cheek; 5 basal
radiating strize, no apical denticles and circuli moderate. Dorsal Vil, I, 9, 1,
second spine 12 in total head length, second and third spines subequal, first
ray 2; Anal Ill, 9,1, third spine 14, first ray 1; caudal I, deeply forked, lobes
slender; least depth of caudal peduncle 23; pectoral 14; ventral 13.
Iniform pale brownish, scales along edge of back dusted with deeper brown,
also jaws dusted brown. Distinct dusky slate line along caudal peduncle and
tail medianly each side. Second dorsal membrane blackish. Fine pale. Iris
slate.
One 60 mm. Ambassis urotenia, described from Delagoa Bay by Fowler’ is
not Bleeker’s species, but a younger stage of the present, as it is described with
2 rows of scales on the cheek.
1 Proc. Acad Nat. Sci: Phita., 1925; p..220.
NOTES AND DESCRIPTIONS OF FISHES FROM CEYLON 709
LUTJANIDA
Lutjanus kasmira (Forskal)
Depth 23 ; head 22, width 2g. Snout 34; eye 34, 1,4, in snout, greater than
interorbital ; maxillary reaches 3 in eye, expansion 24 in eye, length 23 in
head ; interorbital 42, little convex ; preopercle edge not gashed behind, finely
denticulated and denticles larger around angle. Gill rakers 7+ 15, lanceolate.
Scales 43 in lateral line to caudal base and 7 more on latter, 7 above, 12 below,
16 predorsal, 5 rows on cheek ; 7 or 8 basal radiating striz, apical denticles 40
to 58, with 6 or 7 series of transverse basal elements and circuli fine. Dorsal
X, 13,1, fourth spine 22 in head, first ray 22; Anal Ill, 8,1, second spine 23,
second ray 23; caudal 14, slightly emarginate; least depth of caudal peduncle
374 ; pectoral 12; ventral 1%. Back brown, sides below and under surface paler
to whitish. Each row of scales with obscure pale median stripe. Diffuse dusky
stripe from snout tip through eye across postorbital. Diffuse blackish blotch
size of eye on lateral line below front of soft dorsal. Vertical fins pale brownish,
paired fins paler or whitish. Iris slate.
One 84 mm.
‘THERAPONIDZ
Therapon jarbua (Forskal)
One 100 mm.
-SPARIDZX
Sparus berda Forskal
One 80 mm. Molars in 3 rather irregular series in each jaw posteriorly.
Preorbital with its least width 12 in eye. Dark blotch at shoulder but little
distinct. Dorsal with more or less dusky on membranes terminally.
LETHRINIDZ
Lethrinus reticulatus, Valenciennes
Depth 22 to 2g; head 22 to 22, width 23 to 22. Snout 23; eye 3, 1f in snout,
greater than interorbital; maxillary not quite or to eye front, 3 in head ;
interorbital 4 to 4g, slightly convex. Gill rakers4-+ 7, short tubercles. Scales
45 in lateral! line to caudal base and 2 more on latter, 6 above, 13 below, 9
predorsal. Dorsal X,9,1, fourth spine 22 to 2¢ in head, first ray 22 to 23 ;
Anal Ill, 8, second spine 23 to 24, second ray 22; caudal 14 to 1g, emarginate ;
least depth of caudal peduncle 3 to 334,; pectoral 13 to 1g; ventral 14. Pale
olive gray, slightly paler on under surface. Sides and back with darker and
rather large reticulations, mostly with meshes deeper vertically. Iris slate.
Postocular more or less dark olive. Variably diffuse dark blotch above pectorai
base and another vertically at caudal base. Dorsals and caudal with grayish
tints, fins otherwise pale.
Two 68 to72mm. These appear to be the young of this species, originally
described from an example 102 mm. long obtained in New Guinea. Valencien-
nes says the head is reddish with 2 or 3 brown bands below the eye. Opercular
border yellow. Body paler than head, irregularly blotched with blackish.
Soft dorsal, anal and caudal finely streaked with dusky. Bleeker’s figure’
appears to be the same, though it is less variegated than my examples, or
without the dark suprapectoral blotch or the basal caudal bar. Lethrinus
meensit Bleeker also appears to be the adult stage. The species ranges from the
Seychelles to Oceania.
GERRIDZ
Gerres lucitdus Cuvier
One 70 mm. Snout short, 13 in large eye, which 23in head. Third dorsal
spine 13 in head, second spine but little shorter Though spinous dorsal dusky
terminally on 3 highest membranes the usual spots subbasal on membranes
above edge of basal scaly sheath are little evident.
SILLAGINIDZ
Sillago sthama (Forskal)
One 140 mm.
1 Atlas Ichth., 8, 1877-78, Pl. 320 (52), Fig. 1.
710 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
MONODACTYLIDZ
Monodactylus argenteus (Linneus)
One 64mm. Brown ocular band and narrower brown band from befo1e
spinous dorsal down on opercle.
SIGANIDZ
Siganus javus (Linnzus)
One 62 mm.
Siganus virgatus (Valenciennes)
One 85mm. This differs largely from Day’s figure in that the dark ocular
band from the spinous dorsal to the chin is very obsolete, though it has
traces of several dark interorbital lines. The head and body are rather coarsely
spotted, blotched or vermiculated with deeper brown. Spinous dorsal and
anal clouded with deeper brown, though on soft dorsal 3 rows of dusky spots
largely on each ray and most distinct basally on fin. On anal largely as 2 dark
basal bars. Caudal transversely and diffusely barred with dusky. Pectoral
pale. Ventral dusted with dusky.
GOBIIDZ
Glossogobius biocellatus (Valenciennes. )
Depth 52 ; head 23, width 2. Snout 33 in head from snout tip; eye 4g, 14 in
snout ; maxillary reaches nearly opposite hind eye edge, 2 in head; teeth
moderately large, well spaced, in bands in jaws, none on palate; tongue with
deep median notch in front ; interorbital narrow bony frenum, 24 in eye. Gill
rakers 2+ 7, short low rounded tubercles. Scales 30 from over gill opening
to caudal base and 3 more on latter; 8 scales transversely from soft dorsal
origin ; 14 predorsal ; 14 to 17 basal radiating striz, apical denticles 31+ 41 to
32+ 36 and circuli fine. Dorsal VI—I, 9, 1, fourth spine 23 in total head length,
eighth ray 13; Anall, 8, seventh ray 1;; caudal 1g, pointed; least depth of
caudal peduncle 3$; pectoral 13; ventral 13. Brown above, under surface
paler though more or less soiled ; back and sides mottled and marbled with
dusky. Iris slate. Spinous dorsal gray, clouded with dusky, leaving some
small white spots and rather large conspicuous black blotch on penultimate
membrane medially. Soft dorsal gray with dusky reticulations. Caudal pale
above lower portions gray with rather broad blackish transverse bars. Anal
blackish, with scattered small spot basally. Pectoral with scattered dusky
and blackish dots. Ventral neutral dusky termiually, outer borders pale or
gray.
One 93 nim.
THE MAMMALS AND BIRDS OF KASHMIR AND THE
ADJACENT HILL PROVINCES
BEING
NATURAL History NOTES
BY
Cot. A. E. WARD
PART VII
(Continued trom page 11 of Vol. XX X71)
Family—AHyenide
Genus—Hyana
This group is in India represented by
The Striped Hyzeena—AHyena striata
Vernacular names.—' Lakha-bagh, or bagha Lakva’—and most
commonly ‘ Chirak and Jhirak ’.
Colour.—Darkish grey, sometimes of a brownish hue, traversed
with tawny stripes, these are often of a blackish-brown colour, and
are on the body and limbs.
Dimensions. —Head and body about 34 feet, tail with hair under
14 feet. Skuli of a large male 8” basal length, zygomatic measure-
ments6:1%. Weight63lbs. This specimen was shot nearthe Jumna
bridge at Kalsi in the Western Dun.
It is not often that a hyzena can be weighed as the carcase is left
as a rule when shot.
Distribution.—Very rare in Kashmir but has been found on the
Murree road and in Hammel. It is fairly common in some of the
hill districts with which we are concerned, and practically throughout
India.
Hlabits.—-It is not uncommon to turn hyzenas out of short grass
whilst beating with elephants in the Terai; on more than one occasion
I have shot hyznas within a few feet of the elephant’s trunk,
killing dead with a charge of large shot. A big hyana was very
troublesome in a cattle-breeding farm. I campedin the farm and
spent two days looking for the animal and eventually got close to it.
It was eating the body of a calf within a few yards of the sheds, and
to my intense surprise I killed it dead with a charge of No. 5.
Whilst staying with a regiment in a Musketry Camp in the hills,
there was a good deal of trouble with the very disagreeable cries of
hyeenas at night. We used to turn out with guns, but although we
saw the animals we were unable to shoot owing to the tents being so
712 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol- Fea,
close together. They occupied any small and convenient patch of
ground nearby inspite of the fact that in camp were several dogs
which the men used to loose after barking deer or jackals.
We were sitting in the mess tent after our meal, when we heard
the men urging ontheirdogs. The sides of the tents were up and we
were iust turning outto see what was afoot, whenahyzena rushed
under the table and bolted out at the other side! Beyond a chair
being broken no harm was done. The hygna ran down a ravine
followed by the dogs, but apparently they did not care to face it.
- Dogs, unless very wary, are apt to be greatly mutilated by
hyzenas ; with their powerful jaws they can crush the leg of any
dog.
Hyzenas can be ridden down on a fast pony which is handy at
turning.
The fossils in the Siwalik Hills and elsewhere in the vicinity oie
to there having been other species of the Hyzenide in India, and it is
believed that H. crgata was amongst these. Besides these above,
fossils have been found in Madras. Incertain parts of the submontane .
districts, a class of hyenas exists which are small, and are far less
striped than those of the lower Punjab hills—these are only a local
variety of strzata, the only species now found in India.
OrRDER—C YNOJ/DEA.
Family—Canid@: Dogs, Wolves, Jackals, Foxes.
The differences in the footprint of a dog and a cat consist in the
two central toe pads in the former being in advance of the side pads,
making the footprint longer, and in the presence always of blunt
nail prints, owing to the claws being non-retractile. The dentition
consists amongst other teeth of four upper and iower premolars
respectively whilst the cat has three ee tale cus in the upper jaw and
two in the lower jaw.
The genera are known as Canzs, Cyon and Vulpes.
Cants includes the wolves, Cyon the wild dogs, Valdes the foxes.
The Wolf-—Canzs lupus
Vernacular names.—Ratnakin in Kashmir—Chanku in Tibetan—
Baglura in many jungles, where this name is given to both the
Panther and Wolf.
Colour._-Wolf skins are not as variable in colouration as is
generally supposed to be the case. Nearly all Tibetan wolves are
grey or greyish-white with slaty under fur; some show a brownish
tinge mixed with white hairs. The tailis tipped with black. A
few wolves are black due to melanism. Brownish-biack wolves
of the colour sometimes shown by a wild cat as far as I know are
rarely seen.
I have seen a black and agrey wolf with the same mother. Doubt-
less as was the case with the wolves which have been brought home
to England, black cubs have been born from a black parent.
Alsatian hounds have been bred with wolves ; the cubs are apt to
fight amongst themselves, but are quite tame and affectionate to
their owners. These cross-bred hounds are often difficult to rear,
THE MAMMALS AND BIRDS OF KASHMIR Bk)
they are great feeders, and are very delicate during the first year of
their lives in India.
Distribution.—The species C. laniger is from Tibet and Ladak ;
the pelts sold in Srinagar are those of this class.
Habtts.—_Wolves are by no means nocturnal although they rest
during the heat of the day. I have seen a she-wolf stand for hours
in the scorching sun of Ladak, with her cubs secreted in the hills
behind her, she was doing sentry-go between her offspring and the
intruders.
Wolves carry portions of their prey to their caves; I doubt the
wolf being much of a carrion feeder for the simple reason that they
hunt in small packs and eat all the flesh of the animals they have’
killed, and leave the bones to be picked by carrion-eating birds.
I cannot say much regarding the habits of Canzs lupus; they are
rarely seen except when travelling.
Dimensions of a very large Tibetan wolf: Head and Body 3’
10”, height 2’ 5”, weight 60 lbs. Blanford gives alarge skull as
8:7”, basal length and 5-5” in zygomatic breadth. I tried in vain
to shoot wolves in the hills between the Saseer Pass and the Shyok
but could never getnear them. Three wolves stood on a bare hill
and watched ; directly I moved they were off and when I followed
them over a low hill, they watched me from the slopes of the main
hill, and as soon as I showed myself, although I was lying prone on
the ground, they galloped away. ‘This occurred on the southern
slopes of the Karakoram where probably the wolves had rarely
seen a man.
The Indian Wolf.—Canzs pallipes
Vernacular names—PSheriya, Bagyar and Landga.
Probably the best known name is Bheriya whilst Langda is the
name in Gondwana.
Colour.—Grey with a very decided brownish hue; dark onthe back
—much browner than is C. lupus.
Dinvensions.— Much smaller than C. /ufus rarely exceeding 3 feet
apud Blanford skull has 6°85” basal length, and 4:4” broad—weight
from 40 to 45 lbs.
Distribution.—The plains of India, not ascending high into the
Himalayas; at one time common in Abbottabad and Hazara, rare
in the Punjab ranges, very rare in the Vale of Kashmir.
Flabits.—Gregarious to a limited extent, possibly they would be
more so, were they not continually being hunted down. Govern-
ment is always on the watch to destroy wolves as they are very
destructive and kill horses. cattle, sheep, goats or any live
domestic animals they come across. Antelopes are systematically
hunted by three or more wolves working in unison, whilst wolves
which take to killing women and children are much to be dreaded.
The largest wolves I have seen dwelt in the open ground in
front of the old Fort at Fatehgarh.
The Jackal.—Canzs aureus
In vernacular—Gzdar, Shial are probably the commonest names
—Shal in Kashmir—AHokol in Gondwana—J/higal according to
13
714, JOURNAL, BOMBAY NATURAL HIST. SOCIE IP. Vol. ood7
Blanford in Persia but this name is often used by Mahomedans.
There are many names besides these. In parts of the C.P. ‘ Chota-
Bagh’ is often the name.
Near Rewah territory I was taken on an elephant for miles, in
order to shoot a Chota-Bagh, which I naturally thought was a young
tiger or a Jeopard. On arriving at a small village I knew that I
had been sold; I was taken to a thorn bush and there was the
‘Jack’. This was in May! However the information was correct ;
the headman said the Bagh eat their ‘murghis’. I could not
help laughing, much to the disgust of the mahout.
Colour.—Pale rufuous—sometimes reddish. Albinos are very
rare. Blanford mentions ‘ coal black ’—often brown hairs on the
back tipped with black.
Dimensions.— Jackals are often very large in Kashmir, where they
are very destructive to sheep.
Head and body of a Jarge jackal 31” —weight up to 25 lbs. a fine
female weighed 19 ibs.—a male weighed 22 lbs.—both were killed
in the winter.
Distribution.—Throughout India, ascending the hills to about
5,000’ or more. A few days ago jackals were calling at 7,500’ and I
have known them to ascend to much higher elevations.
Jackals eat certain fruits notably Zzzyphus (ber) also Indian corn.
The vernacular name Pfzal is usually applied to the Jackal
when frightened. The name Phzaow is aiso used, it is a mistake
to think that these terms only apply toa‘ Jack ’ which is frightened
by atiger or leopard as they will Phzaow even when pigs or bears
or wild dogs are on the move.
Genus—Cyon
The Indian Wild Dog —Cyon dukhunensts
The only Vernacular names I know are Ram-Hun (Kashmiri)
and Jungli Kutta—Ban Kutta-—Bhansa. Bhansa in the Eastern
Himalayas.
Colour.—Red. I cannot call to mind ever seeing rufuous grey
or light brownish-grey (Blanford), but I have seen brownish-red
wild dogs; there are sometimes dark tips to the hairs on the back.
Lighter red below. The pups are sooty brown.
Dimensions. —Head and body 38”, tail with hair 14”—weight
28 lbs.—heaviest known weight 32 Ibs.
The habitat extends throughout India in forest land, ranging high
up in the Himalayas. In parts of the upper Indus Valley it is fairly
common, but is gradually being reduced in numbers. The wild dog
is very rare in the Vale of Kashmir, I have not seen one for many
years past, nor have I seen the skins in the shops except one or two
from Baltistan.
Habits.—Gregarious and predatory, living on any animal killed.
They hunt in packs and take Sambhar, Nilgai and all sorts of
deer, while cattle, sheep, goats and ponies fall, easy victims to
their rapacity. Even leopards are persecuted by wild dogs and
compelled to take to trees for safety. They are often heard whilst
THE MAMMALS AND BIRDS OF KASHMIR 715
hunting at night and over and over again have I listened to them
whilst lying awake in the jungles.
That wild dogs call when hunting is true as I have watched them
at afew yards whilst doing so. When running by scent they only
whimper, but when the prey is in sight and at hand, they often
break into an undescribable howl.
Cyon dukhunensts breed in colonies in burrows in soft ground, also
in caves or holes amongst rocks, hunting the surrounding jungles.
Having once found a colony, it is as well to set to work delibera-
tely and smoke them all out. ‘The young are probably smothered ;
those able to run about get into the surrounding grass and can be
beaten out by a line of elephants. They are hard to hit with a
bullet whilst dodging through the grass, and unless close, are still
more difficult to kill with shot.
A colony was breeding within about a mile of a jungle where
there were tigers. Strangely enough no foot prints of the dogs
were to be seen in the vicinity and it was evident that they gave the
big cats a wide berth and both refrained from interfering with each
other.
The breeding season is in the winter months as a rule, but 1 have
seen quite small pups in May.
The period of gestation is probably nine weeks.
The Indian Fox—Vulpes bengalensis
Vernacular names: Lomri—Lom in Hindustani, Lomrz and some-
times Lhal in Jammu, Kokrz and Khakvyi are names which are
sometimes heard in Northern India; I have heard the name Kokrz
in the Punjab.
Colour.—From light to reddish-grey. Jerdon remarks, ‘ The fur
just after it has assumed its winter coat Is very beautiful, a pure grey
in the body contrasting with the rufous limbs.’
I have often watched various foxes during the morning and even-
ing hours and invariably remarked that early in the morning V2/fes
bengalensts does not seem to be in the least hurry when moving off
to the burrows. It is seldom that it is to be seen carrying food,
homewards, but when so doing it proceeds very warily.
Dimensions.—V ary a good deal. Length, head and body 19 to 22”
generally about 22”; tail 12” weight of a full-grown specimen
6 to 7 lbs.
Distribution. —According to Jerdon ‘found throughout India’
rare in forest countries. Blanford and others put the limit at ‘ not
recorded west of Sind and the Punjab.’ Personally the extreme
west I have seen it is inthe Jammu Province, Sialkote and adjacent
localities.
The Common Fox—Vulfes alopex
Vernacular names: Lomrz. Laash is not the Kashmiri name for
the common fox, it is the name usually for any kind of cat. It is
quite impossible to give any standard of colour to the Himalayan
variety ; go to any furriers shop and you will find pelts of almost
any colouring. On the other hand V. flavescens is a superb fox of
almost a universal yellow, often witha tinge of golden-rufous. The
716 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
weight given by Jerdon as 14 ibs. is fairly correct, but 18 lbs. has
been recorded. The average weight of six maie foxes is 12 lbs.
The price of a pelt is about Rs. 25.
Turning to the Himalayan form. The colouring on the back
varies from red and yellow, to dark grey and very rarely to black.
The sides are pale. Ears are jet black outside. This portion of the
skin is stitched together and made into collarets, fetching of course
very high prices. The ears inside are generally buff. ‘The face
is rufuous, there is a large black spot in front of the eye and the
cheeks are white’ (Blanford). Tail nearly always grey; the tip of
the tail is white.
Hlabits:—These foxes dwell in burrows or amongst rocks at an
elevation of 5,000’ to about 8,000’. In the summer the cubs come out
inthe afternoon and play about on the swards of green turf; the
vixen generally rests half hidden by some bushes and at the
slightest sign of danger she utters a shrill bark repeating it at short
intervals, finally if any of the cubs are too keen at their play, the
mother races across the grass, the whoie family following and they
are soon lost to view. Probably none of them will be seen for
some days. I have never seen the mother and cubs dart into their
homes when frightened. Cubs wiil often play at the entrance ofa
cleft in rocks whilst the mother lies secreted under cover. Seldom
more than five are produced ata birth and by August they are
sufficiently well grown to be able to escape from most terriers.
The Small Tibetan Fox—Vulges ferrtlatus
Vernacular Names: J/gar (Tibetan) according to Blanford. I
have heard that name in Little Tibet. Lokrz is the common name.
Dimensions: Head and. body 197” to. 20” ;\ tall with*hait22:
Habitat.—Tibet, with a considerable distribution from the Northern
Sutlej to the East.
Colour.—-Reddish-yellow, tail plain grey, tip of the tail pure white,
breast and stomach pure white.
The pelts of this fox are not common in the furriers shops, but
from time to time a certain number can be selected which are so
much of one size, and approximately of the same colour, as to
justify the placing them as belonging to a race of foxes which come
from the same locality. The traders who bring down skins give but
little help. I think V. ferrz/atus comes from nearer to our borders
than do other varieties of V. alopex.
(Lo be continued.)
FORAGING TERMITES
(AUTERMES BIFORMTS, WASM)
BY
R. W. G. Hineston, MAJor, I.M.S.
(With a plate)
The place of investigation is Central India, the district of the
Bundelkhand gneiss. Here we stand on the primitive floor of the
Peninsula, those crystalline rocks on which have been deposited all
the later sedimentary beds. Itis atract of undulating uplands,
wide flat spaces, rocky ridges, crumbling granite hills. Vast areas
of it are bare, stony and neglected; other tracts are clcthed pro-
fusely in forest or thorny scrub.
Eutermes bitormis, a species of White Ant, is particularly partial
to this kind of soil. Inplaces I find it in great abundance, colonies
under almost every stone. Here we have a species of peculiar
interest. It raises no mounds, builds no elaborate nor conspicuous
city, possesses none of that architectural capacity so commonly met
with amongst the white ants. As an artificer it has notthe slightest
merit. Its interest lies in its absence of seclusion. The mound-
building termites hide beneath the soil. Whenthey move about
they always do so beneath tunnels. Rarely, if ever, do they come
into view. All their work takes placein the strictest seclusion,
which makes them particularly difficult to observe. We must disturb
them before seeing them at work. It is otherwise with FAutermes
btformis. This venturesome species has open-air habits, excavating,
foraging, guarding the termitarium in the full light of day. From
it we will learn some facts about termites which otherwise we would
never know.
As in all the species, we find workers and soldiers. The workers
are pale yellow, with shining heads that show not a trace of eyes.
The soldiers are of two distinct sizes, smallones that scarcely exceed
one-tenth of an inch, and large ones that are almost one-fifth of an
inch long. Inone point they differ remarkably from the workers.
Their mouth parts are very peculiar. They have no trace of jaws
or any type of biting organ; in their place is a conical pointed spear,
akind of tubular beak. Obviously their work is altogether different
from that which falls to the workerclass. In addition the community
has its males and females, very much larger than the workers, and
provided with eyes and wings.
THE NEST
Nothing special marks the Autermes nest, no mound, no heap, no
citadel of spires. All it doesis to make a superficial excavation
scarcely more than afew inches indepth. It seldom goes down
718. JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
beyond afoot. Ifthe insect has any preference in the matter, it
likes to establish itself underneath a stone. :
The nest consists ofa ramification of passages, a reticulated
system of galleries and chambers, their width being equal to about
that of a lead pencil. They appear to have no particular arrange-
ment, but just ramify and communicate in ail directions. The patch
of ground is literally honeycombed withthe network. When ex-
ploring it we seem to be cutting intoa sponge. Thewalls of the
passages are a little roughened and usually lined with a black
material. This pigment, at first sight, looks like some fungus; but
more probably it is the excreta of the termites which they smear
over the walls.
The galleries are always perfectly clean. What do we findin
them ? First of all, nurseries. Special chambers appear to be
reserved for this purpose. In them we find young of all ages and
sizes heaped together in a scrambling mass. Hundreds may be
kept in the largest of the nurseries. Others are met with scattered
through the galleries. Less often we come across sexual forms,
perhaps only afew inches beneath the soil. The only other stuff
I have recorded from these galleries are fragments of half-chewed
orass.
Il have said that these termites emerge into the open, so let us
try and catch them at nest-building work. How do they excavate
their network of galleries? It is not a sight that we can see at any
time. But, if fortunate, the following spectacle presents itself. A
stream of termites issues from the nest. All belong to the worker
class. Each carries a load of excavated material, a small damp
pellet of earth, whichit casts away on the rubbish heap. The
stream moves slowly, carefully, methodically. ‘Though all its
members are completely blind, yet they advance with the same
certainty as if they had perfect sight. It is obvious that their
antenne feel out the way. The load deposited, they return for
more. . In fact their manner of excavation is identical with that of
the ordinary ant.
But where are the soldiers in this important business? These
diggers and carriers are all workers. Not a single soldier takes
part in the toil. What is their function? Have they no part in the
making of the nest? They have a part, and a very remarkable one.
Look along the edge of the excavated earth, the material which the
workers are heaping into amound. There the soldiers have taken
their stand. They have formed a ring, a martial circle, a defensive
barrier around the heap. Within the circle is the stream of workers
carrying and casting down their loads. Both types of soldiers join
in this rampart, the large ones are twice the size of the small ones,
yet their duties appear to be the same.
Here is a social system most elaborately organized, the peaceful
toilers, the martial ring, the military machinery ready forwar. It
is worth looking at in closer detail. The martial ring isa circle of
sentinels. They are fairly evenly distributed: there may be fifty or
a hundred of them: they do not run about, but stand stationary in
the circle, each keeping to its allotted place. All face outward, their
dagger-shaped beaks away from the nest opening, their headsa
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FORAGING TERMITES 719
little elevated, antennze held erect, and trembling as though feeling
for danger inthe air. It may be only an insect spectacle, a vision
of little and unimportant things. But little things are big things
when viewed in proportion. And here we see termites with that
organized machinery which brutalizes civilized life.
Clearly this is a military system. Let us watch the barbarous
machinery at work. I disturb the excavators, the worthy toilers
that are carrying out the earth. Instantaneously a retreat is
sounded. Thelabourers make no attempt at resistance. Inhalfa
minute all are in the nest. Nota worker is visible; the martial
element holds the field. The soldiers, to aman, remain outside.
A mad excited frenzy seizes them. They rush about on the heap
of debris. Eager for battle, they seek out the enemy. The martial
ring disintegrates for fight.
This is the first lesson taught by Autermes. How highly developed
is the military machinery amongst creatures so low in the scale. It
reminds us of Hymenopteron society with their labourers, their
soldiers and their defensive methods. But the militarism of
Futermesis more impressive than that of ants. In Autermes the
castes, both in structure and strategy, are far more strikingly
distinct. Nevertheless ants have sometimes been observed making
the same kind of martial ring. But never, I feel sure, with the
perfection of Zutermes. Moggridge saw Atta barbara making rings
around its queens to protect them from lizards which would not
pass through the ring. Miss Cheesman in the Society Islands saw
something similar with Phezdole. She gave food to the ants. Their
soldiers made aring around it, kept back all other competitors while
the smaller workers took it piecemeal away.
In the higher forms of termites the military organization is well
displayed at the nuptial flight. Take Odoniotermes tee for example.
Workers first break openthe galleries. ‘hen the winged forms
squeeze their way out, hundreds emerging one after another and
ascending into the air. In an hour or more the eruption ceases.
Then the breaches have to be repaired. Workers come forward to
seal these openings. And now for the first time soldiers appear.
They donot come out completely from the galleries. ‘Their method
is to remain within the passage and from there line the breach. We
see their heads protruding above the margin, clearly keeping a
strict guard while the workers seal the opening with clay.
FORAGING EXPEDITIONS
We leave for a moment these military adventures and turn to
more productive manifestations of life. Hzlermes despatches
foraging columns, mainly for the purpose of collecting grass. The
expeditions take place in the monsoon evenings, an hour or two
before the sun sets. A sharp eye is needed to detect them; for the
column of pale flesh-coloured workers 1s almost invisible against the
soil. A course is made for the harvesting area, some suitable patch
of grass usually about three or four feet from the nest. A pathway
is formed on which the insects move. Their method is like that of
harvesting ants. Those facing outward are empty-handed; those
720 JOURNAL, BOMBAY NATCRAL- AIST. SOCIETY, Vol. XXXT7/
returning are Jaden with grass. The marchis orderly. In coming
and going they pass one another, and move without confusion along
the path. Hach individual with its leafy loadreminds us of the leaf-
cutting ants of America that advance beneath a Birnam wood. I
have seen many thousands on these foraging expeditions; the
whole nest seemed to have been abroad. On such occasions they
are extremely timid. ‘The slightest interference alarms the mass
which turns about and hastens indoors. In the harvesting area we
see quiet activity. Some individuals mount the blades, cut off the
tips of the vegetation and allow themselves to fall with the separat-
ed ends. Others find suitable fragments on the ground, either green
cuttings that have dropped from above, or pieces of dried stalks. In
the business of transport they are individualistic. Each sticks to
its own load. They cannot combine to shift alarge mass. Here
again they resemble the harvesters and certain of the insect-hunting
ants.
Now for the part taken by the soldiers. It is only the workers
that cut and carry grass. The soldiers are unfitted for transporting
operations, even if they had the instinct to work. Their mouth-parts,
as I have said, are shaped into a spear, an implement that could
never grip a load. But the soldiers are not absent from these
harvesting operations. As in excavating, so in foraging, the
warriors have important duties to fulfil. Soldiers mix themselves
in the harvesting column. It reminds one of social hymenoptera on
the march. Others stand along the foraging track, others scatter
around the collecting area, some ascend the blades of grass. Wher-
ever the toilers extend their enterprise, there the soldiers also appear.
Their business is to protect the foragers, in the same way as they
did the excavators, though the irregular nature of the ground does
not lend itself to the martial ring. Interfere with the steady stream
of industry. Immediately the soldiers are up in arms ready to
attack anything that comes. When alarmed, the foragers set about
retirement ; the stream of labourers pours back to the nest. But
the soldiers remain out until all have disappeared ; then the warriors
themselves withdraw, forming, as it were, a rearguard to the mass.
Another small experiment shows their military instinct. I kill a
worker while it is engaged in foraging. Another worker finds it,
drags it towards the termitarium. Soldiers assemble, close around
it; a martial ring, or is it a ring of mourners, is formed round the
moving corpse. The procession advances, afunereal spectacle,
the bier, the corpse, the military assemblage all go steadily into the
nest.
Why does Autermes require all this soldiering ? What is the
particular enemy they fear? I have little doubt that ants are the
danger. How often have we seen the ravager, Lobopelia, breaking
through the earthworks of a termites’ nest.
So dangerous to Autermes are insect-hunting ants that the species
often builds covered roadways along the surface of the soil. They
are nothing like the extensive ramifications of other termites, but
are just low tunnels fashioned out of earth. As arule they are short
and project from the nest opening ; seldom are they more than an
inch or two in length. Such a scanty surface architecture does little
FORAGING TERMITES 721
more than guard the entrance to the termitarium. It is all that
remains to the foraging species of other termites’ pyramids and
domes. ‘lhe building of these roadways is done in daylight ; in fact
building and provisioning may go on simultaneously. Division of
labour exists amongst termites with the same efficiency as in
ordinary ants. Wesee some workers carrying clay from the termi-
tarium, others moulding it to the roof of the passages, while others
at the same time collect grass. The foragers keep close to the end
of the tunnel, scarcely going farther than half an inch from it. They
get hold of the nearest blade of grass, gnaw it off, drag it into the
tube. The work is done with apparent timidity. The object of
the tunnel is to permit the workers to get without exposure to
the nearest patch of grass.
DUTIES OF SOLDIERS
This is clearly the most interesting point. The soldiers possess
two important functions, different from those of the ordinary worker.
(1) The function of their antenne. (2) The function of the pointed
beak.
First let us see the function of the antenne. J uproot a stone,
one beneath which a nest is established, and, in doing so, I break
through the reticnlated walls. Termites burst forth, a violent
eruption, which very quickly subsides. Then the work of repair
begins. The soldiers seek out the broken galleries; they gather
along the edge of the breaches. The ordinary workers for the
moment disappear, withdrawing themselves into the labyrinth of
tubes. Soon the workers reappear, bringing with them whatever
fragments of material they can find, pellets of earth, nodules of grit,
pieces of dry grass. A moistening with saliva makes these frag-
ments viscid. Then they are glued into the breach. ‘They are
not merely laid in position. The workers with an oscillating
movement of the head screw them forcibly into place.
Now let us watch the behaviour of the soldiers. In this work of
building they clearly have duties other than those of defence.
Collecting along the edge of the rupture, heads turned outward,
antennz expanded, they stand apart at fairly even distances so that
the antennz of each soldier just touches those of its companion on
either side. Jn this we have the strategical ring. But when we
watch the soldiers’ antennze we see that they do something more.
We notice that they are all the time swaying and touching ; they
give the impression of estimating distance and in some way measur-
ing the work. I little doubt that they are being used as calculating
organs, measuring out the extent of the damage and determining
the progress of repair. The soldiers in this way act as supervisors
while the workers apply the bricks. The breach now rapidly con-
tracts. Soon the antenne of the supervising soldiers can touch
from opposite sides of the rent. They now know that repair is
nearing completion. Hence they withdraw into the gallery; only
their heads appear in the breach. Their antennz still project,
gently swaying, and gauging how much remains to be done.
Brick after brick is brought by the workers ; in the end the soldiers
14
722 “JOURNAL, BOMBAY NATURAL FIST. SOGETY, Vol Zee
all but disappear. Only the tips of their antennz are visible when
the last few bricks are moulded to the wall.
Here we have a clear division of labour. Measuring and
calculating falls to the soldiers; labourers do the ordinary toil.
Now for the purpose of the pointed beak. Of course this is the
organ of defence. While the swarm is engaged at harvesting I
place an injured cricket near the nest. This to the termites is a
terrible invasion. All the workers rushfor the entrance. Soldiers
dash in and surround the invader. Some of them shoot jets of
poison from their beaks. All their military tactics are called forth.
When the workers withdraw the soldiers follow them. As soon as
all are safely housed the soldiers take up a position at the gate. A
martial group occupies the entrance, their bodies are kept in the
interior, their heads appear at the open door. In fact the gate is
lined with soldiers, and bristles with antennary threads. Now
comes the final act of defence. While the soldiers hold the entrance
the workers build a rampart across it. Loads of earth are carried
up until the door is firmly sealed. How human-like are these
military tactics. First an attack, then a rearguard action, then a
picket holding the last defences while a suitable rampart is thrown
up. Again we see how like they are to ordinary ants. J/essor
barbarus, for example, under similar circumstances defends the open
gate of its formicary by placing across it a barricade of stones.
I have said that, when attacking, they shoot out poison. This
brings me to the soldier’s spear. The weapon is not particularly
sharp, being really a cylindrical tube intended for ejecting spray.
When the soldier is annoyed it shoots out from the spear a jet of
clear sticky fluid, liquid on emergence, but hardening on exposure.
It oozes out like a rope of viscid treacle and then falls in glutinous
coils. It has no effect on litmus. Its smell is distinct. Sometimes it
comes out in a series of spurts as if driven with definite force.
Thus is the spear a defensive weapon, not, however, a penetrating
dagger; it is more in the nature of a poison syringe. It must be of
great value against natural enemies, of which the carnivorous ants
are chief. A rearguard of soldiers standing at a gateway or swarm-
ing on the breach of a broken gallery, with their poison spears all
directed outward, offer an impregnable front to attack. Woe betide
the enemy smeared with their discharge. Its joints, mouth-parts,
legs become clogged; gravel, leaves, everything sticks to it. If it
does not escape it gets glued to the ground.
SPECIAL SENSES
Touch.—The antennz, of course, are the tactile organs. Thrust
out in front, they touch every object and carefully explore the way.
For the soldiers they seem to serve aS measuring implements on
those occasions when the passages are under repair.
Smell.—Like ants, they possess a good sense of smell. In one
place the stream of foragers led over a smooth stone. I rubbed my
finger a number of times across their path. This disturbed them,
but only slightly. The line became broken where I had interfered.
New arrivals hesitated to cross. My rubbing had done something
FORAGING TERMITES 723
which checked their course. Soon, however, they regained
confidence. The stream continued as before. Ants, like JZessor,
would have been much more inconvenienced. Yet this incident
clearly shows that Autermes has some sense of smell.
When the foraging army was returning to the termitarium, I
placed a nodule of camphor in their path. They were instantly
aware of it, clearly disliked it. Before ever their antennze touched it
they halted and turned aside. Very soon the whole army was
deflected and had to reach the nest by a circuitous route.
Around the nest entrance I made aring of camphor. The army
was abroad. News of the interference quickly reached them.
Much alarmed, they called a retirement. But the camphor blocked
them. They would not approach it. Though madly determined to
reach the next aperture, yet they were utterly cut off. They were
as isolated from the termitarium as if it were surrounded by a
watery moat.
These simple experiments indicate clearly that termite activities
are mainly guided by their keen sense of smell.
Hearing .—Eutermes has a peculiar habit which suggests an auditory
sense. When alarmed, they erect themselves, shake their bodies,
rapidly and forcibly oscillate their antennz. Both soldiers and
workers adopt this manoeuvre. It is very evident when a nest is
disturbed, or when a soldier is irritated with a straw. The act isa
plan of communicating danger. When one termite erects itself, its
neighbours become alarined. Others at a distance catch the infec-
tion; if the army is abroad the news quickly spreads and in a few
moments all are racing for the nest. The communication is not
visual. Lutermesis blind. Can it have an auditory significance?
Certainly it is very like that of the Red Ant. Odcecophylla erects
itself, jerks its body, hammers the surface of a leaf with its abdomen.
A tapping noise is produced in this way, a trumpet that spreads
alarm. The vibrations of Autermes give no audible sound, at least
nothing that we can hear. Perhaps it is beyond the range of human
hearing. For we might anticipate auditory powers in insects
deprived of sight.
The suspicion of an auditory sense is strengthened by the fact
that Smeathman observed a marching Termite in Africa which
emitted a distinct noise. He saw columns of workers issuing from
ahole, marching under the direction of enormous soldiers which
climbed up the plants and made audible signals to the marching
column who responded with a hissing noise. Also Bristowe detected
two kinds of noises made by a Brazilian species of termite. He
refers to the soldiers of Conztermes similis. Some of them jerked
their bodies up and down and made a tapping noise by a movement
of the chin. Others by sharply crossing their mandibles produced a
noise like the snapping of thin glass. More than.likely Autermes
bzformits can emit some sound inaudible to us.
Sight.—A word disposes of this faculty. Both soldiers and
workers are totally blind.
Recognition of one another.—Ants, as is well known, are friendly
to their comrades, but hostile to members of another community
though belonging to the same species. Furthermore we have
724 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
reason to believe that each ant can distinguish every other ant
within the confines of its own nest. Is there anything of this kind
in the termite community ?
I take a worker from one nest and transfer it toa gang of the
same species employedat collecting grass. The gang refuses to
accept the worker. Indeed they show it every mark of hostility.
A soldier meets it. The alarm is sounded. Other soldiers arrive,
shoot their poison at it, and soon have it helpless onthe ground
smeared in viscid juice. Thus Aulermes like ants shows communal
independence. The members, though all in harmony with one
another, are hostile to the members of other nests.
COMPARISON WITH HYMENOPTERA
The chief point that strikes us in this record of behaviour is the
close resemblance in small details between Azutermes and the
Hymenoptera. A termite and an ant are as different in structure as
are a dragon-fly andabee. Yet see their detailed resemblance in
behaviour. I do not refer to their social organization, a resemblance
that has frequently been pointed out. JI am concerned only with
details of behaviour as exemplified in this daylight haunting species.
Their mode of excavation is exactly like that of ants, the gang of
porters carrying out the loads and pitching them down on the
rubbish heap. The protective ring of stalwart soldiers calls to
mind a nest of Camponotus compressus where idle monsters move
about the rubbish heap and smaller labourers eject the earth. We
have also seen that Atta and Pheidole do occasionally manufacture
aring. Disturb Autermes, andthe soldiers show fight; so do the
Camponotus monsters while the busy toilers hasten underground.’
How very like are their foraging expeditions to the seed-collecting
habits of WZessor barbarus. We have the common path, the harvest-
ing area, the coming and going with individual loads. Their cut-
ting of the vegetation is like the leaf-collecting Oecodoma where each
reaps its leafy fragment and bearsit away tothenest. Their march
is on the plan of the foraging Ecitons, the stream of workers, the
scattered soldiers, each with its duty to perform. The building of
arcades is done by &. vastator which spreads along the grounda
covered roadway underneath which the workers advance.
Ants are renowned for their division of labour. With Autermes we
have excavators, builders, provisioners, each toits owntask. Their
rampart of defence is the plan of MWessor barbarus which keeps out
the enemy witha barricade of stones. Ants are particular in dis-
posing of their dead. Weare even given an instance of a funeral
procession where a iong and organized stream of workers followed
the corpse to a special grave. Such a story might be thought to be
purely imaginary. Yet what have we seen in Hutermes biformts ?;
a funeral ring encircling the corpse and retiring with it into the nest.
Their sensory capacities are in many ways like those of ants. Their
sense of smell seems identical with that of Messor barbarus which
+ The habits of these Indian species of ants have been described in A
Naturalist in Himalaya and A Naturalist in Hindustan,
FORAGING TERMITES 725
camphor distracts in exactly the same way. They communicate
danger by shakings and jerkings, not at all unlike the plan that
Ocecophylla adopts. Lastly we see that recognition characteristic of
ants in which the members of a community are friendly to one
another but hostile to the members of other nests.
I conclude with a quotation from Professor Wheeler. He is
referring to termite communities far more highly organized than
mine. ‘When we reflect that ants and termites have been able,
through slow physiological and instinctive processes, independently
to evolve such strikingly analogous peculiarities as those I have
described, we can scarcely doubt that different human communities,
belonging to the same species and endowed with some intelligence,
may frequently have hit upon the same inventions.’
FURTHER NOTES ON BIRDS ABOUT SIMLA
BY
HuUGH WHISTLER, F.L.S., F.ZS.
In Vol. xxvi of the Journal, pages 770-775, I recorded a series of observatiots
made about Fagoo near Simlain 1918 and followed it upin Vol. xxvii, pages
94-111, with a further series of notes made in 1919 in Simla. As it was my good
fortune to be posted as Superintendent of Police, Simla, for the season of 1925
and as there is no liklihood that I shall ever visit the Simla Hilis again, I take this
opportunity of setting on record the more interesting of my notes made in that
year. I arrived at Simla on February 6, 1925 and left it again on October 23
of the same year. Before the season started I was able to make short tours to
Sabathu, Dagshai and Solon from February 27 to March 4 and to Narkunda,
Koteghar and Kotekhai in the second half of March. For the rest of the time I
was confined to Simla or localities within reach of a day’s expedition.
Nucifraga hemispila, Vigors. The Himalayan Nutcracker.
Common from about 6,000-9,000’ in the hills behind Simla as near as Masho-
bra and Mahasoo but not observed in Simla itself. I found it very common and
in pairs about Narkundah from March 13 to 16 and found a nest on March 16
built in the branches of a pine close to the trunk at no great height from the
ground. One bird was sitting in the nest and both made a great fuss when I
examined it, so I was surprised to find that it contained a single dead chick.
Both sexes apparently take part in incubation.
The flight is very similar to that of a Jay—very flapping with rounded wings
and the same rather hesitating manner combined with fair speed.
Stachyridopsis pyrrhops (Blyth). The Red-billed Babbler.
A few pairs were seen at Sabathu when I was there from February 23
to 27, One was seen at Kotekhai on March 20, a pair were seen at 7,000’
in the Chirot Nala, Mahasoo on June 14.
Cephalopyrus flammiceps flammiceps (Burton). The Fire Cap.
On March 21, two flocks were met with in some willows in the Giri Valley
above Parala; they were singing a low twittering song rather like a feeble
version of the song of the Goldfronted Serin (Serinus pusillus).
It was reported as first seen in Simla on March 27 and on April 1 a pair
were found building near Snowdon. ‘This nest contained four rather incubated
eggs on April 21.
On May 8 I found a pair feeding well-grown young in 7,500’ on Jakko. The
nest was in a hole in an Oaktree trunk about 20’ from the ground.
Horeites pallidus pallidus, Brooks. The Pale-bush Warbler.
Several were observed about Kotekhai in the Giri Valley on March 20-21
and the song was also heard there once.
This species breeds in fair numbers about 8,000’ at Mahasoo and Kufri and
from May to August the characteristic song may be freely heard in the thick
undergrowth that clothes the hillsides wherever the larger forest trees have been
cut. Itis probably double brooded as on August 9 I found a brood of newly
fledged young clicking and skulking in arough patch of cover in the cultivation
above the Kufri bazaar.
Sturnus vulgaris poltaratzskyi. Finsch. Finsch’s Starling.
A solitary female of this species was found about three miles below Kotekhai in
the Giri Valley on March 19, It was in poor plumage and was consorting with
Common Mynahs.
Acridotheres tristis (L.). ‘The Mynah.
Common at Dagshai 5,000’ and Sabathu in February, at 7,200’ at Koteghar
in March and at Kotekhai in March.
Tarsiger chryseus whistleri, Ticehurst. The Golden Bush-Robin.
A female was found at about 6,500’ in the Chirot Nala on March 29.
FURTHER NOTES ON BIRDS ABOUT SIMLA 727
Oreocincla mollissima simlaiensis, S-B. The Plain-backed Mountain-Thrush.
I shot a maleon Jakko at 8,000’ on February 14 ; another Mountain-Thrush
seen the same day and on February 11 also on Jakko was probably of the same
species.
Carpodacus thurus blythi (Piddulph). The White-browed Rose- Finch.
I shot an immature male (in the plumage resembling that of the female) at
8,000’ Kufri on February 22, 1925. It was a solitary bird and was feeding on
the ground in a light sallow thicket on the top of the Kufri ridge. This appears
to be the first record of this species in the Punjab Province.
Callacanthis burtoni (Gould). The Red-browed Finch.
Isaw a male on February 17 and two females on February 18, in both cases
feeding on the bank above the mall close toGorton Castle. A flock of about
thirty were seen by a friend on March 1, feeding by the side of the cartroad
below the Cecil Hotel. These birds are very tame and have the movements on
the ground of a heavy sparrow.
Serinus pusillus (Pallas). The Gold-fronted Serin.
On February 27, I found a flock on the cartroad about a mile from Sabathu.
From March 19 to 21,1 met withsome parties in the Giri Valley between
Parala and Kotkhai. A single bird was heard at 6,000’ on the southern face of
Tara Devi by the cartroad on April 10.
Fringilla montifringilla, L. The Brambling.
I have now to recant the former opinion expressed by me in the Journal
(Vol. xxx, p. 410) that there must be some mistake about Jerdon’s record that
the Brambling occurred at Simla.
On December 13, 1924, Mr. A. E. Jones obtained a female in his garden at
Marston Grange (7,000’).
Then on February 15 and 22,1 met personally with several in the culti-
vation at 8,000’ above Kufri bazaar. They were in company with the flocks
of Stoliczka’s Mountain-Finch that were common on the hillside amongst the
snow and owing to the restless behaviour of the latter species were hard to
obtain when with them. However, the Bramblings at times liked to rest quietly
in the leafy boughs of the Mountain Oaks and were then easy to approach.
When I visited the same place on March 10, I could only find a single female
in a huge flock of the Mountain Finches.
In all I obtained five specimens.
Athopyga gouldie (Vigors). Mrs. Gould’s Sunbird.
On April 5, I saw a brilliant adult male in the Chirot Nala at about
6,500’ ; it was hovering in the undergrowth apparently collecting spiders
threads for its nest and then dashed away intothe forest out of sight.
The call is a note likethe rapid opening and closing of a paur of
scissors asin the case of “. siparaja. Near the same spot on July 12, I
shot two young males. They were moving about some parasitic plants at the
top of a lofty oak tree, turning incessantly from side to side and uttering the
same scissor’s note.
Diceum ignipectum (Hodgson). The Fire-breasted Flowerpecker.
On February 24-25, 1 found several of these little birds about Sabathu
Cantonment.
Dryobates hyperythrus (Vigors). The Rufous-bellied Pied Woodpecker.
A male was seen in the pine woods of Narkundah at 9,000’ on March 16.
Apus melba (L). The Alpine Swift.
_ A few Alpine Swifts were noted in the vicinity of Simla on various dates from
April 11 to May 17.
On the autumn migration however they were observed in much larger numbers
than is usual about Simla, and for a longer period; the first birds were seen
on August 21 and the last on September 24; between those two dates I have
records of flocks seen on sixteen different days, on some days in very large num-
bers.
Cacomantis merulinus passerinus (Vahl.) The Indian Plaintive Cuckoo.
I heard this little Cuckoo on July 15 calling in a valley about 5,500’, below
Kasumpti.
728 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Strix aluco nivicola (Blyth). The Himalayan Wood-Owl.
On April 4 I found one of these delightful birds sitting in a holly oak bush
near the ground and it allowed me to look at it from a distance of about 4 feet.
It then flew off and settled near the ground in another tree. This was at 7,800’
on Jakko where the species may be heard calling at nights.
On July 12 I found two adults sitting in the undergrowth of wattle bushes at
7,300’ by the private road in the Chirot Nullah.
Strix indranee indranee (Sykes). The Brown Wood-Owl.
Only observed on one occasion (Apri! 16) in a nullah just above Annandale in
the late afternoon.
Otus spilocephaius (Blyth). The Spotted Himalayan Scops-Owl.
The characteristic call of this little owl was heard in the grounds of Vice-
regal Lodge on October 21.
Glaucidium brodiei (Burton). The Collared Pigmy Owlet.
This Pigmy Owlet must be quite common in and about Simla and the neigh-
bouring hills as | heard it calling on various dates from February 20 to May 17
after which the call was not heard. A female was shot at Dagshai 5,700’ on
March 2 and I observed it up to 9,000’ at Narkunda. It calls both by day and
night, and is active by daylight.
Torgos calvus (Scop). The King-Vulture.
Observed as follows :— February 27 Dharmpore, March 2 Dagshai, March 3
Solon, March 8 over Tara Devi and Simla W., March 10 beyond Sanjouli,
March 18, Theog, April 5 by the Tollbar. After that it was not seen.
Pseudogyps bengalensis (Gm.) The White-backed Vulture.
Observed as follows : -March 2 Dagshai, March 8 two or three on Tara Devi,
May 10 about Naldera.
Giypaetus barbatus hemachalanus, Hutton. The Lammergayer.
On September 24 an adult Lammergayer was brought in to Mr. A. E. Jones
by some hill men, who declared that they had caught it in some telegraph wires.
Although we could detect no sign of injury it was very quiet in demeanour
sitting passively on the floor, a little inclined to bite if molested but otherwise
still and silent ; it kept the head rather lowered and the mouth a little open.
If uninjured it may have felt conscious of the hopelessness of opening its huge
wings for flight in the restricted area of the room. Mr. Jones kindly allowed
me to present this bird to the Society and its portrait appears in our Journal
(Vol, xxxi, 198).
Hieraetus fasciatus (Vieill). Bonelli’s Eagle.
Bonelli’s Eagle was seen from the railway between Solon and Tara Devi
on March 2. Single birds, apparently of this species, were seen soaring over
the Hotel Cecil on March 9 and by Parala in the Giri Valley on March 21.
Spilornis cheela (Latham). The Crested Serpent Eagle.
Occasionally observed about the catchment area from April to July.
Ictinaetus maiayensis perniger (Hodgson). The Black Eagle.
As there is little on record regarding this fine Eagle in the Western Himalayas
it seems worth while recording my experience of it in detail.
On March 14and 16 I saw an Eagle, which was almost certainly of this_
species, soaring about Sidhpur 8,500’-9,000’ between Koteghar and Narkunda,
and on the latter date it also flew over Narkunda.
What was perhaps another was seen on May 10 near the Retreat, Mashobra.
On June 28 I had a splendid view of one hunting in the bottom of the Chirot
Nala (catchment area) at about 7,000’. It was very dark in appearance, save
for a light area in the primaries and the marked bars on the tail. The yellow
at the base of the beak was very noticeable. It was a wonderful flier covering
the sides of the nullah in circular beats, practically without flapping the wings.
The wings and tail looked very broad and rounded, the primaries spreading out
like the fingers of a hand.
On August 16 I secured a fine maleat Kufri 8,000’. This was in complete
moult with the organs very indistinct. In the crop was a naked nestling of
FURTHER NOTES ON BIRDS ABOUT SIMLA 729
some passerine bird, apparently Zrochalopieron lineatum and in the stomach
incubated eggs of a Laughing Thrusheither 7. vartegatum or T. erythrocepha-
lum. It seems fair therefore to assume that the marvellous powers of flight are
devoted in the summer months chiefly to bird nesting and that as the Eagle
quarters the hillsides it isscanning the bushes for nests. This is the only
specimen of the Black Eagle I have ever secured personally and the thrill that
the incident afforded will perhaps excuse the inclusion of the following extract
from my diary :—
‘* We emerged out of the forest on to a bare spur and looked down into the
head of the nullah which lay below us a thousand feet, breaking up into numerous
side ravines, partly clothed in forest,and partly bare or clothed with undergrowth,
in places almost precipitous. There, far below us was aBlack Eagle which
was hunting over the trees and undergrowth in regular beats, backwards and
forwards, sailing in and out of the nullahs and contours with the utmost ease
and grace. This he was doing in the head of the main nullah, and then he
started to beat up the side towards where we were resting, after one false
alarm in which he started towards our side and then returned to the main
nullah. I told my companions to remain absolutely still without moving hand or
foot as we were out inthe open as the Eagle worked along the sides of the
nullah, quartering the ground backwards and forwards but all the while
gradually advancing in our direction, I saw that his course was steadily
ascending and would probably bring him into the gap between some trees on my ~
left. Fora moment he went out of sight in a nullah soJ hastily shifted a yard
or two to cover that gap and waited with my 410 collecting gun (No. 8shot)
ready and cocked. In my new position I could no longer see the hillside and
watch the progress of the bird but a minute or two later it suddenly sailed into
the gap in front of me, and seeing me steered off to drift down hill. I fired and
hit the bird which staggered and again recovered, and turning made off in the
direction in which it had come; but it was obvious that it had been hard hit.
We watched it keenly as it went, and it disappeared into the deodars that
clothed a side nullah ; however it did not reappear on the far side of the nuilah,
nor could we see it elsewhere. Obviously it had stopped in the nullah, so
leaving Major C. to watch from our spur I made off down the hillside with my
orderly Tulsi Ram as fast as might be consistent with caution.
I entered the deodars very quietly with gun ready loaded and every nerve on
the gut vive to detect the wounded Eagle resting in a tree, or to hear the sound
of small birds mobbing it. The sunlight and the shadow were very peaceful
and quiet, and the only calls came from a large party of Grosbeaks in the trees
and some Variegated Laughing Thrushes—but these all were obviously concern-
ed only with their own affairs. There wasnosign of my quarry. I passed
through the nullah and emerged on to the ridge beyond and then signalled and
shouted to Major C. perched on the spur above, with his handkerchief fluttering
in the breeze to show his position. He replied that there was still no sign of the
bird. TulsiRam and I then worked back into the nullah, taking different
lines, and a welcome shout soon announced that Tulsi Ram had found the
Eagle lying dead in a tree.”’
Accipiter virgatus affinis, J. E. Gray. The Besra Sparrow-Hawk.
The late Mr. Basil-Edwards kindly allowed me to examine a beautiful adult
male Besra that he shot at Simla 6,000’, on June 28, 1925.
On June 18I1 founda nest of the Chestnut-bellied Rock-Thrush at 8,000
on a pineclad slope of the Chirot Nala. It contained two well-feathered chicks
which as I looked at them dashed out of the nest and down the hillside. At
once there was an agitated skirmish which resolved itself into the parent Rock-
Thrushes trying to save the chicks from a tiny hawk, which dashed at them in
a most determined manner regardless of my presence and successfully carried
one of them off. This Hawk could only have been the adult male of this spe-
cies, and I saw what was probably the same bird at the bottom ofthe nullah
on July 12.
Falco peregrinator, Sund. The Shahin Falcon.
In March I found a pair of Shahins frequenting the face of a mountain close
to Simla and on April 10 I discovered what was evidently the site of their
eyrie ip a precipice ; it was however quite inaccessible. The same birds or others
were seen over Simla occasionally in June and July.
15
730 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXTI
Falco subbuteo, L. The Hobby.
During the summer months the Hobby is distincly scarce in the neighbour-
hood of Simla and I did not hear of it until May 7, when one was reported as
seen flying over the Ridge. It does however breed inthe vicinity, as will be
seen from the following account taken from my diary :—
‘*On the evening of June 17, 1925, I went out to examine a certain ridge near
Simla, on which the [fobby Falcon has been known to breed for at least 10 or
12 years. The upper slopes of this ridge, looking towards the S. W. are
broken by small nullahs, and covered with a fairly thick growth of Deodar
forest, of no great age.
I started towards the top of the hill and commenced to work downwards in
hopes of finding the whereabouts of the Hobbies. I had not gone far before I
heard the screaming of a Falcon below me and was able to track it intoa
small nullah filled with Deodars. I accordingly took upa position on the
ridge at one side of the nullah and settled to watch. It was not long before I
made out a Hobby sitting in a Deodar tree where it was calling and after a short
time the other bird also appeared. Both Hobbies were very noisy screaming
at intervals the plaintive rising fee—tee—tee—tee which is common to most of
the Falcons, and varying it occasionally with a harsh single note pz¢t p7¢ or
chip chip chip uttered two or three times in succession. They kept on flying
. about in circles, sometimes over the trees, sometimes amongst them and often
low over the ground, settling here and there on different trees.
One of the Hobbies was carrying something in its talons—apparently a
small bird—which it eventually ate on a Deodar.
This behaviour went on for sometime and I then left the place without dis-
turbing the birds.
There was no doubt from their excitement that they were preparing to
breed, and I noticed a crow’s nest in very good condition which seemed a likely
nest for them to use, though they did not directly approach it while I was
there. It was about 40’ from the ground against the trunk of one of the
Deodars overhanging a mule road that wound through the nullah. This spot
is at an elevation of about 6,500’.
On June 24, 1925, I returned to the spot about 5p.m. walking along the mule
road and as I approached the site of the Crow’s nest the little male Hobby
was visible sitting on the topmost shoot of a tall Deodar about 20 yards away.
He paid no attention tome andthe female was not visible anywhere so I
climbed up on to the ridge, where [ had sat before, and made myself com-
fortable to watch.
The little Falcon was sitting there on his lofty perch in complete ease,
looking over the wide valley that stretched before him with portions of Simla
visible on the ridges across the other side. It was cool and pleasant with a
bright sun and though he was sitting quite very much at his ease he was all the
time on the alert, his head turning from side to side incessantly. After atime
he started to stretch himself and show systems of preparing to move, then he
flew off circled once in the air and disappeared over the ridge for his evening’s
hunting and I did not see him again.
I waited there for sometime longer but the female did not appear and
nothing happened. Then I climbed along the hillside to a spot whence I could
see onto the top of the crow’s nest noted at my previous visit, and there was
the female sitting in the nest, where she had obviously been all the time.
So without disturbing her I went away determined to allow ample time for the
eggs to be laid.
On June 27 I returned to take the eggs. As we arrived at the spot there
was complete silence andthe male could not be seen on any of the trees, nor
from below could we distinguish any bird in the nest. I hada sinking feeling
that something was wrong and that the prize was not to be mine : however I
told Tulsi Ram, my orderly, to go up the tree and he divested himself of his
haversack and coat and started : he was not more than a quarter of the way upto
the nest when the female Hobby flew out of the nest screaming once or twice
and settled on a bough of a Deodar some 30 yards away. There she sat more
or less unconcerned and took but little notice of the climber.
He reached the nest and reported three eggs and said that the nest was
quite small and that there was no food lying aboutit. I told him to take the eggs
and substitute for them a couple of hen’s eggs, slightly smeared with mud to
darken them, which we had brought forthe purpose. ‘This he did and des-
FURTHER NOTES ON BIRDS ABOUT SIMLA 431
cended again. Wethen stayed on the road below the nest examining and
packing the eggs but almost as soon as the climber was down the Hobby took
to wing and flying in a circle round the nest tree settled ina neighbouring
Deodar. This she repeated again, flying close to the hillside, and after one or
two more short flights she returned to the nest and settled herself down in it on
the hen’s eggs.
We then climbed up to the usual knoll and settled there to watch ; the female
Hobby was visible through the glasses sitting very deep in the nest cavity, her
back below the line of the sides of the nest; her head was a little higher but
hardly able to see over the side, her tail projecting rather upwards over one
side.
For a time nothing happened : then in the distance I heard the screaming
calland the male Hobby appeared from out over the valley flying rapidly
towards the site of the nest: as he approached it was evident that there was
food in his talons and he circled once past the nest calling, while the female
answered, and after that he settled on a neighbouring Deodar. Here he sat on
a horizontal bough and I could see he was carrying the partially plumed
body of a small bird. The female slipped off the nest and settled in another
neighbouring Deodar. The male kept on calling sometimes a sharp p7t p7t pit,
sometimes the familiar screaming call, while the female answered in the same
way and flew from perch to perch. Then the male took the bird in his beak
and the female flew on to the bough beside him, took it in her beak, transferred
it to her claws and flew away and settled in the Deodars further up the nullah
where we could not see her. The male relieved of his burden cleaned his beak
against the bough, preened his feathers, and then settled himself to sit patiently
on the bough.
In a comparatively short space of time the female re-appeared having eaten
the bird; she then took some exercise circling in the air and settling in the
various trees, calling a good deal in the process while the male sat quietly in
his place ; finally she flew down tothe nest and settled on the edge of it
where she stood looking about her; then she stepped down into the egg cavity
and stood there, and gradually settled down sometimes putting her head down
to look at or move the eggs ; and before she finally was satisfied she changed
her position in the nest by a half circle.
When she was definitely settled on the eggs, I went down and had the male
flushed and shot him, as the specimen was required. His crop was empty but
there were a few fragments of flesh and feather in the stomach probably eaten
while pluming the bird he had brought for the female. The report of the gun
brought the female off the nest calling but she settled at once in a neighbouring
tree and ultimately returned to the nest in which we left her sitting. Two days
later I sent the orderly to remove the hen’s eggs and he reported that the
female Hobby was not at the nest, so she probably had deserted it on finding
that the male had vanished.
In October there was a most marked migration of Hobbies in the Simla
Hills. On October 3 three were visible high over the Ridge at dusk. On
October 9 one was reported to meand I saw asecond bird that had been
shot.
On October 11 Hobbies were visible in the Chirot Nullah all day on the wing
most of the time, There were at least 5and probably more. I noticed that while
the bird was soaring the primaries slanted slightly downwards, almost below
the level of the back, while the bird moved in curves and circles climbing and
sinking at will in the air with scarcely an effort—an occasionally shifting of the
angles of the wings and two or three rapid strokes of the wings now and again
being all the effort involved. Thus the Hobby soared fora space, then there
would be a burst of activity and the bird flew straight with regular rapid wing
beats ; next followed a swift downward stoop of a hundred feet, and again the
bird was rising swiftly without effort. Thus were the Hobbies feeding or
playing ail day long, backwards and forwards, up and down, over the steep
bare hillsides and the deep cut ravines. I shot an adult and an immature bird ;
both were exceedingly fat and the former had its crop stuffed with remains of
insects, apparently cicadas.
After returning to Simla I saw two more Hobbies by the Grand Hotel in the
evening.
On October 13 I saw one— or perhaps more—at dusk soaring high over Gorton
Castle. On October 15 I saw two at dusk high over Snowdon. The same
732, JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
evening there were said to be about a dozen over Mashobra and one was shot
and sent to me to substantiate the record.
These birds must have moved on immediately as on October 181 saw none in
the course of along day overtowards Kufri, though the next evening at dusk
saw a single bird over the Ridge.
Arboricola torqueola millardi, Stuart Baker. The Hill-Partridge.
I attribute to this species a characteristic call of the Simla hills, a very gentle
melancholy poor or pheaw, wnich at closer quarters sounds more like guaaw ; it
is uttered singly at short intervals and the sound carries for a long distance, being
audible throughout a whole nullah. This call may be heard from 6,000 to 8,000’
in the Simla hills throughout May and June and the first half of July and occa-
sionally at other times. If my attribution is correct the Hill-Partridge must be
fairly common, though I only occasionally flushed a pair on forest-clad hill-
sides.
Journ., Bombay Nat. Hist. Soc.
Barleria Gibsontotdes, BLATTER, SP. NOV.
Del. Norah Coady.
SOME NEW SPECIES OF PLANTS FROM THE WESTERN GHATS
BY
EK. BLATTER AND C. McCann
(With a Plate)
Barleria Gibsonioides, Blatter, sf. zov. [Acanthacea sectionis Eubarlerie,
similis Barleriz Gibsoni, Dalz. a qua tamen recedit bracteis basi semper late
rotundis, sepalis interioribus glabris exceptis marginibus ciliolatis, corolla
minime glabra, seminibus discoideis, colore corollae].
A tufted undershrub, up tol m. high, very leafy. Stems quadrangular,
very obscurely winged, stiff, erect, light green ; internodes thickened above the
leaves. Leaves glabrous, except the margin which is finely spinous-hairy,
leathery or papery, somewhat rigid, lowest broadly ovate or almost rounded,
obtuse at the tip, lamina up to 14cm. by up to 10 cm., the rest ovate-oblong or
ob ong, acuminate, up to 20 cm. by 8 cm., very variable in size and shape,
shining-green above, glaucous beneath, base rounded or tapering, main
nerves up to 7 pairs, depressed and whitish above, prominent beneath ; stalks
of lowest leaves up to 4 cm. long, getting shorter upwards. Flowers in dense,
imbricate, terminal spikes, rarely solitary, axillary ; spikes more or less 10 cm.
long in flower. Bracts foliaceous, the lowest 1 or 2 pairs resembling the leaves
and up tol0 cm. long, the rest getting rapidly shorter upwards and shorter
than the calyx, broadly elliptic-lanceolate, acute, broadly rotund at the base,
pale-green. Bracteoles at base of stalkless flower, linear, boat-shaped, subacute,
glabrous on both surfaces, margin ciliolate, up to2 cm. by 2mm. Outer sepals
2, ellipsoid, rounded at base, up to 5cm. long and 2-23 cm. broad, often
exactly alike as to size and shape, but one generally slightly shorter and sub-
acute at the apex, the other acute, both distinctly nerved, quite glabrous, glau-
cous on the lower surface, pale green and shining on the upper; inner sepals
2, narrowly linear-lanceolate, acuminate, boat-shaped, up to 15 mm.’ long,
2-3 mm. broad, glabrous, with ciliolate margin. Corolla 6-10 cm. long, minute-
ly glandular-hairy on the margin and tube, sparingly so on the surface of the
lobes ; tube 4-5 cm. long, enlarged upwards, dark purple inside and outside,
lobes obovate or ellipsoid, rounded at the apex, delicately veined, mauve (light
and dark), the 2upper lobes each with a large elongated dark purple blotch
beginning at base of lobe and extending 15-20 mm., blotches sometimes faint
or absent. Stamens 2; filaments white in the lower part, mauve in the upper ;
anthers violet ; pollen globose, white. Staminodes 3, without anthers. Base of
stamens and staminodes minutely glandular-hairy. Style filamentous, white in
the lower part, mauve above. Stigina bifid. Unripe capsule oblong, tip conical,
shining, smooth. Seeds 4, disc-shaped, glabrous. [PLATE I.]
Very variable as to size and shape of leaves even on the same plant. In open
Situations the leaves are shorter, thicker, stiffer, and darker green on the
upper surface, whilst when growing amongst other vegetation, they are longer,
narrower, thinner and lighter green. In the latter surroundings the internodes
of the stem are considerably thinner and longer. ‘The flowers, too, vary a
good deal as to size and shape of the coroila-lobes. On drying the upper
side of the leaves turns black, the lower bluish-grey.
Locality : Below Godowli,3 miles from Panchgani, W. Ghats of Bombay
Presidency (Blatter Nos. 1, 2, 3, 5, 7 type, 4, 6, co-type). Wasin flower during
the first half of October 1927.
Crinum Eleonorae, Blatter & McCann, spf. nov. [Amaryllidacea Sectionis
Brachynematis, similis Crino brachyneniati, Herbert, a quo tamen differt collo
robusto longo, staminum filamentis longioribus, antheris duplo longioribus,
stylo quam filamentis multo longioribus].
Bulb ovoid or almost spherical, up to20 by 13cm., narrowed into a stout
, heck about 8 cm. long, clothed with old pale leaf-sheaths. Leaves appearing
734 JOURNAL, BOMBAY NATURAL HIST, SOGIETY, Vol eed?
after the flowers, ensiform, with obtuse apex up to 60cm. long and 5 cm. broad,
not very firm, fairly thin, dark green, glabrous, with the margins entire.
Scape lateral, up to 50 cm. long, reaching 2°5 cm. diam., cylindrical, but
slightly compressed, green, tinged with purple. Flowers white, strongly scented
upto 20 in an umbel, nodding. Bracts 2, broadly lanceolate, obtuse or acute,
with broadly inflexed and thickened margin, 9 by 3:3cm., green, tinged with
purple outside, pale inside. Bracteoles linear, up to9cm. lony, green, tinged
with purple. Pedicels up tolcm. long, stout. Perianth funnel-shaped, tube
green, tinged with purple, or greenish and becoming paler towards tip, up to
6°5 cm. long, cylindric at base, then becoming 4-sided with rounded angles ;
lobes up to 7'5 cm. long and 22cm. broad, lanceolate, the inner ones some-
times oblanceolate, cuspidate, reflexed and slightly concave at the tip, with a
longer cusp in bud, pure white, sometimes tinged with purple along the median
line on the back ; the buds are cream-coloured and the purple tinge is more
pronounced. Filaments on the throat of the perianth-tube, free, filiform,
ourple, 1 cm. long ; anthers in young buds yellow, then (whilst still in the bud)
zetting black and only yellow along the open slits, dorsifixed, up to 15 mm.
Jong, oblong, twisted when dry. Style filiform, included in the tube, up to
5 cm. long, white in the lower part, purple above. Fruit not seen.
The anthers open in the bud.
Locality : Lingmala near Mahableshwar, open hill-sides and on both sides of
the river above the Yenna Falls, also half-way between Panchgani and Maha-
bleshwar (McCann Nos. 7, 8, 9, 10, type).
Forma purpurea.—McCann found specimens in the same localities with the
perigone-segments oblanceolate, deeply tinged with purple on the dorsal side,
and the perigone-tube with the bracts uniformly brown-purple. (McCann No.
Ii; type.)
In flower: End of April and beginning of May.
iphigenia stellata, Blatter, sf. zov. [Liliacea, similis Iphigeniw, indice A. Gray,
differt tamen ab ea floribus duplo maijoribus, petalis minime unguiculatis, oblongis
vel oblongo-lanceolatis, vel oblongo-obovatis, non lineari-subulatis, mucronatis
vel acuminatis].
Corm subglobose, 10-18 min.in diam., narrowed into a short hypogezal neck,
tunicate with light-brown membranous sheaths. Stem up tol0 cm. high, rigid
to flexuous. Leaves alternate, 2-4, grass-like, sessile, the lower up to 12 by
1cm., all about the same size, linear or linear-lanceolate, tip acute-mucronate,
coriaceous, rigid, keeled, shining on both surfaces, base long-sheathing ; midrib
depressed on upper surface, nerves many, upto about 12, obscure above, very
prominent beneath. Flowers various hues from pale to dark mauve, cream-
coloured in bud, up to 7in aterminal short raceme. Bracts linear cr linear-
lanceolate, resembling theleaves, upto6 cm. long. Pedicels 12-20 mm. long,
stout, thickened below the flower, deeply 6-grooved, each ridge supporting a
petal. Perianth-segments up to 15 mm. long, up to 4 mm. broad in the middle,
oblong or oblong-lanceolate, or oblong-obovate, mucronate or acuminate, not
clawed, spreading. Stamens up to5 mm. long, purple; filaments rigid,
straight, slightly fattened on inner surface ; anthers 1-1°5 mm. long, versatile,
purple ; pollen yellow. Ovary green, slightly shorter than the stamens, oblong-
trigonous, 6-sulcate; styles 3, 2 mm. long, united atthe base for about % their
length, recurved, dark purple, stigmatic surface beginning with the curve of
the styles; ovules ovoid-oblong. Capsule almost globose, or obovate or oblong-
obovate, loculicidal, up to8 mm. long, deep grooves between the carpels,
shallow grooves on back of carpels, styles and stigmas persistent. Seeds subglo-
bose or ovoid, 2 mm. long, slightly irregular, flattened on one side, umbonate,
yellow testa very thin.
Like J. indica very variable and resembling it at first sight. Can always
be distinguished by the much larger flowers and by the shape of the clawless
petals. _
Locality : Tableland of Panchgani, W. Ghats of Bombay Presidency, fairly
common (Blatter Pl type), also found on Second and Third Tableland, on
Tableland near Mahomedan School.
Was found in flower about the middle of June, 1925. There had been some
heavy showers at the end of May and beginning of June.
Fruit noted about the middle of August.
NEW SPECIES OF PLANTS FROM THE WESTERN GHATS 735
Urginea polyantha, Blatter & McCann, sf. nov. [Liliacea similis Ureinece
indicee, Kunth, sed recedit ab ea pedicellis brevibus nunquam deflexis, racemo
denso multifloro, perianthio rotato, perianthii segmentis omnibus apice barbatis,
staminibus brevioribus, stylo cylindrico, capsula apice truncata].
Bulb globose-ovoid, tunicate, pale, up to 5 by 4°5 cm., neck 3°5 cm. long,
conical. Leaves appearing after the flowers, not observed. Scape up tc 60 cm.
high, up to 4mm. diam. at the base, slender, rigid, slightly bent at tip, wreenish
to “purple, dense-flowered. Flowers many, up to 50, about 1 cm. and less apart
on the raceme up to 25 cm. long. Pedicels filiform, up to 6 mm. long, spreading
in flower, ascending or appressed and almost double the length in fruit. Bracts
deltoid, about 1 mm. long, often still visible in fruit. Flower-buds brownish-
purple, obovate when very young, gradually becoming clavate. Perianth
rotate, pale yellowish-green inside, with a broad brownish band along the
centre outside ; segments oblong, 5-6mm. long, bearded at tip, outer ones
3 mm. broad, inner 2-2°5 mm. broad, obtuse. Stamens whitish, 2°5 mm. long ;
filainents flattened, getting very thin upwards; anthers versatile, cordate at
base. Ovary oblong, 6-grooved. Style short, stout, uniformly thick through-
out, 3-grooved. Stigma trigonous. Capsule up to 10 by 5 mm., broadly
ovoid-oblong or broadly ellipsoid, truncate at tip, coriaceous, triquetrous, each
carpel with an elevated groove in centre and with a slightly higher rib on
margin. Cells 2-10-seeded. Seeds broadly ellipsoid, sometimes irregularly
toothed at upper end, broadly winged, up to 6 by 5 mm., flat, shining black,
seed without wing pear-shaped, flat.
Tetramerous ovaries have been observed. The flowers open before dawn and
remain open till about noon. The perianth encloses the fruit till the latter
reaches half the final size and then falls away.
Can be distinguished from U. Wightiana, Hook. f. by the much denser
raceines, the shorter pedicels, the shorter and broader perianth-segments, the
flowers not drooping, the seeds not being orbicular, and by the deltoid bracts.
In U. coromandeliana, Hook. f. the flowers are drooping and have longer
pedicels, only the inner perianth-segments are bearded at the tip, and the style
is obconic.
Locality : Amongst short grass on Tableland of Panchgani, W. Ghats of
Bombay Presidency, 4,400 ft., very common in some patches, less common on
slope of Tableland (Blatter and McCann No. 101, type).
Flowered from the middle of March toend of April, 1926.
Dipcadi Ursulae, Blatter, sf. mov. [Pertinet ad familiam Liliacearum ;
accedit ad Dipcadi montanum, Baker, a quotamen recedit foliis latioribus,
scapo tereti, bracteis ovatis longe-acuminatis quain pedicelli multo longioribus,
pedicellis in vivo robustis minime filiformibus, perianthii lobis exterioribus
quam tubus multo longioribus. }.
Bulb globose, small, about 20 mm.in diam., tunicate, with many white
fibres from the base. Leaves 4-5, leathery, 15-30 cm. long, up to 7mm. broad,
linear, plicate, grass-green, sheathing at the underground base. Scape green,
terete, 15-30 cm. long, smooth, naked. Flowerswhen in bud in dense racemes
of 6-12 flowers, ultimately the raceme becoming lax and reaching up to 12 cm.
long. Bracts caducous, ovate, long-acuminate, 10-26 mm. long, reaching 2-3
mm. beyond the bud, centre green, margin in the lower part or the whole
scarious. Pedicels 6-9 mm. long, stout, not filiform. Flower-buds dark pink-
brown. Perianth 15 mm.long, 18mm. across, slightly fragrant, the 3 outer
lobes 9 by 3-4 mm., reaching 3-4 mm. beyond the tube, oblong, obtuse,
recurved from about the middle and bent downwards, with 5 close, parallel
nerves in the middle and with an indurated minutely hairy apex, lower surface
pink-brown or greenish along the centre with the margin whitish, upper surface
greenish or whitish or dirty-white, along the centre darker ; the 3 inner perianth-
lobes connate to about the middle, lower part oblong, 5mm. long, tip 3-4 mm.
long and as broad, recurved at right angle to tube, with an indurated apex
which bears a few very minute hairs, lower surface whitish, along midnerve
green or pink-brown, upper side white, or slightly tinged with green or pink-
brown. Stamens attached to the perianth tube; filaments in their free part
linear-lanceolate, flattened, 3 mm. long, white, bent twice ata right angle just
below the anther. Anthers 2 mm. long, oblong, flat, thin, green, reaching the
mouth of the tube, dehiscing longitudinally, introrsely. Ovary narrowly
obovoid, trigonous, 5 mm. long, 2-2°5 mm. broad in the upper part, green,
736 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol, XXXII
stalk 1mm.long. Style 5-6 mm. long, stout, slightly thickened at both ends,
papillose, especially in the upper half. Stigma indistinctly 3-lobed, 1:5 mm.
in diam., not thicker than the style. Young capsule much longer than broad.
Locality: In grass on Tableland of Panchgani, W. Ghats of Bombay
Presidency, 4,400 ft. (Blatter No. P74, type).
Flowered about the middle of August 1925.
Dipcadi saxorum, Blatter, sf. mwov. [Pertinens ad familiam Liliacearum
accedit ad Dipcadi concanense, Baker, differet tamen ab ea specie scapo necnon
racemo multo longioribus, racemo ca 20-floro, bracteis late ovatis et longe
acuminatis basi laciniolatis, perianthio parvo, segmentis ovato-obtusis,
staminum filamentis brevissimis, stylo multo breviore, seminibus paullisper
ellipticis vel fere orbicularibus].
Bulb tunicate. Scape about 50 cm. long, terete, smooth. Racemes up to 20
cm. long, rather lax, about 20-flowered. Bracts broadly ovate, long-acumi-
nate, scarious, slightly fringed near the base about 7 mm. long. Pedicels stout,
by zlonger than the bracts. Perianth 12-14 mm. long, tubular, slightly
trigonous, the 3 outer lobes connate for 3 their length, linear-oblong, obtuse,
recurved from about the middle with athick, glandular tip, the 3 inner
connate for almost § their length, 3mm. broad, ovate-obtuse, with a recurved
and slightly thickened tip, all lobes whitish with an olive-coloured central band.
Filaments about 1 mm. long, slightly flattened ; anthers 3 mm. long, versatile,
yellow. Pistill0 mm. long. Ovary5 mm. long, trigonous, clavate, shortly
but distinctly stipitate ; style very stout, 5 mm. long, minutely granular
upwards; stigma slightly thicker than style, 3-lobed. Capsule broader than
long, deeply 3-sulcate, loculicidally 3-valved, membranous, stipitate, stalk
stout, about 2°5 mm. long; cells 5-seeded, seeds slightly elliptical to nearly
orbicular in outline, reaching 5 mm. diam., compressed, with a raised margin,
the lowest and topmost in each cell plano-convex.
Locality : Rocky places above the Kanheri Caves in Saisette, about 1,000 ft.,
Bombay Presidency (Blatter and Hallberg No. 8S], type)
Flowered in August 1917.
BIRDS OF PREY AND THEIR USES
BY
C,H. DONALD), F.Z.S., MiB.0.U;
(With # plates)
From time immemorial the Raptores, as aclass, have been looked
upon as ‘ vermin’ to be destroyed at sight. The gamekeeper has
no use for them whatsoever, the poultry farmer (particularly in this
country), wages continual war on them, and even the Memsahib,
and her cook, chiefly the latter, have given of their best mutton-
chops or beef ‘i-stakes’ to satisfy the hunger of some marauding
kite.
In England, even the little kestrel is persecuted by the game-
keeper as a destroyer of pheasant chicks and partridges, and in this
country, asin parts of Ireland, ‘the HAWK’ is anathema to all
who have anything to do with the preservation of game.
Yet, there is another side to the story, andif we go into the
subject a little deeper than what appears onthe surface, and con-
sider the habits of the various species, one begins to wonder what
we should do without them ?
It, of course, must be thoroughly understood from the first that
no bird of prey will ever refuse a good opportunity of taking an
easy prey, such as a wounded or sick bird, or one in which the
flight is defective from an injury to the wing or otherwise. A
White-eyed Buzzard (Butastur teesa) who has spent his life catching
rats, frogs and insects, will immediately bestir himself should a
maimed partridge appear in the open within the range of his vision,
whereas normally, a whole covey might feed with impunity ina field
within fifty yards of his perch without his taking any notice of them
or the partridges of him.
Let us now try and see to what extent the birds of prey, asa
class, deserve this unenviable reputation.
From Mr. Stuart Baker’s Hand List of the Birds of India we
find there are 105 varieties of diurnal birds of prey. This
represents almost one-sixth of the world’s total number, so India
is well supplied.
Of this number many are winter visitors which arrive in the
autumn and disappear in the spring. Some are to be found in the
hills and not in the plains, and vice versa, whereas many which are
found in the north are wanting in the south, and southern birds are
very often unknown in the north.
From this it is pretty evident that no hard-and-fast rule can be
laid down for any particular locality, or even Province, and if we
are to discover to what extent the Raptores are injurious to our
16
738 JOURNAL, BOMBAY NATCERAL FIST. SOGETY, Vol. Cod?
game preserves, we must first find out what species are normally
found in that area.
To begin with let us make a list of the genera to be found in the
whole of India, from which we must eliminate those which are
harmless, and I have no doubt that many of our members will
probably be surprised at the result.
The Order Accipitres is divided into three families, viz.
Pandionide, Gypide and Falconide, the last being again
subdivided into two sub-families, Gypaétinze and Falconine. It is
unnecessary here to go into details regarding every species or
variety, but we can enumerate the genera and state how many
species each is comprised of, and these genera are :—
1. Pandion (Osprey), one; Vultures (4 gypius, Torgos, Gyps,
Pseudogyps, and Neophron) in all ten species: The Lammergeier
(Gypetus), one; True Eagles (Aguzla), six ; Hawk-eagles (Hzeraétus,
Lophetriorchis, Ictinaétus, Spizaétus), eleven; Serpent Eagles,
Buzzards, Fishing Eagles, Kites, Harriers, Honey-Buzzards, Bazas
and Falconets, in all forty-five species and varieties, beginning
with No. 1621 Czrcaétus gallicus to No. 1654, Buteo b. japonicus ;
No. 1670 to 1675 and again from 1692 to 1695.
This then leaves us with the hawks and falcons, i.e. As/ur,
Acctpiler and Falco, which between them account for thirty-one
species and subspecies.
Now out of this lot the following might safely be eliminated
without further consideration as not coming within the range of
game destroyers, viz. Osprey one, Vultures ten, the miscellaneous
assortment from Serpent Eagles to Faiconets forty-five, and the
Lammergeier one.
That is fifty-seven species, or over one half of the birds of prey
of India.
From this it must not be assumed that none of them ever catch a
game bird, as harriers catch quail or even a very occasional
partridge, and a fishing-eagle might often be seen flying off with
a duck (though probably never one that has not been in some way
maimed), but as a normal thing none of these birds are game
killers and the majority are absolutely incapable of catching any
game bird in full possession of its power of flight.
To this list we can add several others. The genus Aguila
contains six species of big powerful birds, but of these six only one,
the Golden Eagle (4. c. daphane) is a hunter, or capable of catching
any game bird. The remaining five might safely join the list of
the ‘harmless’. The hawk-eagles must all remain in the list of
suspects, and in fact are more or less, rather move than less all
distinctly injurious, with the possible exception of Wieraétus pennatus
and Jctinaélus m. perniger.
Among the hawks and falcons we have a number of keen
hunters, but how many would normally take a bird as big asa
partridge if smaller fry is available? Very few among the hawks,
at any rate. From the falcons we can ignore the merlins, hobbies
and kestrels as being too small to worry about, which leaves us
with six falcons which are all capable of doing considerable
damage,
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BIRDS OF PREY AND THEIR USES 739
This reduces the list to one eagle, nine hawk-eagles, six falcons
and perhaps as many hawks, say a little over a score of birds in all,
out of a total of 105.
Having arrived at the number that should be watched as provable
destroyers, the next question for the game preserver to consider is
how many of these ever visit, or are ever likely to visit his parti-
cular preserves. For this purpose we can divide them into two
classes, those that will be found in the plains, and those that live in
the higher ranges of the Himalayas. Of course, the list will vary
with each province, but if for the sake of argument we take the
Punjab, which I happen to know best, we should arrive at approxi-
mately the following conclusion. I say approximately because
stragglers and wanderers might turn up anywhere.
Variettes fiills Plains
Golden Eagle i| nil
Bonelli’s Eagle 1 1 (up to 8000’)
Changeable Hawk-Hagle (rare) nil (low hills) 1
Hodgson’s Hawk-Eagle if nil
Goshawk 1 nil
Shikras and Sparrow Hawks 5 h)
Falcons 4 6
13 G8)
The above then is a rough list of what might reasonably be
expected to appear in either the hills or the plains of any district in
the Punjab. It includes such birds as the Shanghar and Red-capped
falcons, which are rare at best, whereas many species are restricted
to particular tracts and are by no means universally met with
throughout the Province.
This means, that in a province like the Punjab we need only
bother about some thirteen species of the Raptores as being J/zkely
to destroy our game in any particular locality, and if each province
was taken separately, the result would be more or less similar.
Having arrived at the above conclusion for the province, let us
see how the thirteen birds effect our particular area, be it in the
plains or in the hills, so let us start with a game preserve in the
plains ; what do we find :—
(a) Bonelli’'s Eagle. They hunt in pairs and are undoubtedly
destructive.
(6) Booted Hagle. Rare at best and will seldom take a bird
as quick on the wing as a partridge.
(c) Changeable Hawk-Eagle. Only in the vicinity of low hills,
but very destructive where found.
(d) Shtkras and Sparrow Hawks. Not normally destructive
to birds the size and strength of a partridge, but will take young
chicks and cheepers.
(e) Falcons. Wf near water, the Siberian Peregrine will be a
constant visitor in the winter, but partridges are only in danger
when feeding far from cover.
(f) The Indian Peregrine Falcon. Not very common and birds
only in danger when away from cover.
740 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
(g) The Laggar falcon. Wunts in pairs and a pair may
become attached to a preserve and do considerable damage when
cheepers are about. ,
(h) The Saker Falcon. Is not likely to be found over scrub
jungle, and prefers open plains and sandy tracts. Can be ignored.
(2) Lhe Red-capped and Shanghar Falcons. Too rare to matter.
It will thus be seen that an intelligent interest in the doings of
the Raptores which frequent any area is all that is necessary to
find out the real culprits that are doing, or apt to do, damage but
to call the whole family ‘ vermin’ and try to exterminate -them is
the surest way of ruining any game preserve, as I shall endeavour
to show.
Real enemies of ground game.
The next question is, when does ground game need most pro-
tection? Obviously in the nesting season.
During the nesting season what is its worst enemy? Here we
have a number of things to contend with such as rats, mongooses,
the larger snakes, monitor lizards, and to a smaller extent foxes,
cats and perhaps jackals. Of these, again, which are the worst ?
Monitor lizards, rats (because of their numbers), snakes and mon-
gooses. The last, however, might be allowed considerable
latitude because of the good they do in keeping down snakes and
rats. Snakes also eat rats, but the actual number of rats killed
by snakes in the year must be, at best, comparatively small and
almost negligible.
Any of the above-mentioned will wipe out an entire clutch of
eggs in a few moments. It would take a hawk or a falcon, even
if it took to a game diet and ate nothing else, a week or ten days
to accomplish such a result.
On the other hand what keeps down rats and snakes, besides a
mongoose, cats and foxes? The birds of prey. There is nota
single bird of prey (except vultures), that will not catch and eat a
rat, given the chance; not even the noble Peregrine or the mighty
Goiden Eagle, will refuse the tit-bit, whereas the majority of them
practically live on them. Remove this host from any area and the
result will probably spell disaster.
I have recently been through the best partridge preserves in the
province, during the gun dog trials in Patiala and Jind, and the
fact was very noticeable that we came upon patch of jungle where
the birds were extremely plentiful, in fact. too plentiful for the
peace of mind of the working dogs, whereas we also came upon
patches where tor half an hour at a time, nota bird was flushed.
This fact led me to keep an eye on the nature of the ground and
I noticed that many of these barren beats were infested with rat
holes, and the mounds of the mole-rat were much in evidence.
Was this a coincidence, or does it bear out my theory that rats
are one of the biggest enemies that ground game has?
Throughout the six days, four in Jind and two in Patiala, that I
was there, I also kept a close eye on the birds of prey we en-
countered, and these are the species I saw :—Shikras, in plenty.
One chased a partridge that had been very slightly wounded for
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BIRDS OF PREY AND THEIR USES 741
quite a long way, but failed to catchit. I mnever saw another one
attempt to chase, though I twice saw partridges fly into the open
and pass a tree, into which a shikra had gone a few minutes before,
Without the latter taking any notice. One pair of Red-headed
Merlins, two pairs of Laggar Falcons, several White-eyed Buzzards,
a couple of Long-legged Buzzards, and a larger number of Tawny
and Steppe Eagles, and one Imperial Eagle. A Steppe Eagle carried
off a partridge that had been shot and was, in turn, pursued by no
less than four others and an Imperial, but which eventually got
the booty I did not discover. I noticed harriers on several occa-
sions, and a very fine adult Spotted Eagle took up a commanding
position over the water on which the water test took place, but
‘moved on’ as soon as the first shot was fired. I saw no sign of
any of the ‘nobler’ falcons and was somewhat surprised not to see
a single Bonelli’s Eagle.
The presence of eagles and buzzards is a fairly good indication
that rats are numerous and hunting (of rats) good.
In the hills, what is the worst enemy game birds have to contend
with in the breeding season? The Jungle Crow first and foremost,
and the Indian Marten. Then possibly foxes, cats and such like.
During the autumn and winter the Golden Eagle takes considerable
toll, and the Hodgson’s Hawk-Eagle and the Goshawk at all times,
but the Goshawk is by no means as common as it used to be, and
the Hodgson’s Hawk-Eagle is curiously given to certain localities
and completely wanting in others. It is very common, for instance,
in Bhadarwar and parts of Chamba and very rare in Kangra. Yet
itis very common again in parts of the Simla Hills such as the
Jubal and Taroach States, but rare on the Sutlej watershed. This
species is particularly destructive to pheasants of all kinds and his
method of hunting does not give them the smallest chance of
escape.
The Golden Eagle, on the other hand, does not appear to touch
game during the breeding season and every nest I have examined
(and I have taken four youngsters from their nests, as far apart as
one in Kashmir, two in Bushahr State (Simla Hills) and one in Tehri
Garhwal), has contained more remains of crows than anything else.
The reason for this is not hard to seek. He is a typical falcon in
his way of hunting, and prefers the open. His long wings are all
against his hunting among trees, and unless pheasants can be
flushed for him and rise above the trees, he is powerless. During
the summer months most pheasants keep to thick cover, except
Snow-cock and a few Monal which go out on to the Alpine pastures
to feed. At this period the Golden Eagle has his work cut out
providing food for a shrieking youngster in the nest, and it is not
quality he seeks but quantity, and there is no lack of crows.
Crows, with dozens of youngsters, go in large flocks to roost
among the oaks at high levels in the evening and come down in the
morning, and the eagle has no difficulty in getting as many as he
wants. In this respect he does a great deal of good, but there is
no denying the fact that any and every pheasant which has the
temerity to feed out in the open, in the autumn when the young
eagle accompanies its parents, is looking for trouble.
742 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
Bonelli’s Eagle, though unlike the other hawk-eagles in his
method of hunting, does considerable damage, particularly among
Chukor. They always hunt in pairs and are very persistent in their
attack. oS
Though we have already reduced the possibie destroyers of game to
thirteen, it must be remembered that this number is the maximum
of what might reasonably be expected to be seen over any particular
locality, and it can be very considerably reduced for really practical
purposes. For instance, in the plains, the possibility of five species
of shikras and sparrow-hawks appearing in any one locality is very
remote. Then again, during the most important period of game
preservation, viz. the breeding season, at most two species of
falcons will be found, the rest having left the country.
Thus the game preserver need only trouble himself with a very
small percentage of the total number, and then only if he finds that
some particular individuais have regularly attached themselves to
the vicinity and taken to a game diet. Rather than shoot the wrong
birds it would perhaps be better to shoot none at all.
After all if we look at it in another way we find that the
commonest hawk inthe plains of India, and the hills too for that
matter, is perhaps the Shikra. It takes considerable toll of the
smaller birds, and so do Red-headed Merlins, and yet do we find any
diminution in babblers, sparrows and our host of little garden birds ?
Most of us can probably name various places where game was
plentiful twenty or thirty years ago, inspite of the birds of prey,
which had been there with the game from time immemorial. Ican
even now find my way to places in the Himalayas where Monal and
all the other pheasants are extremely numerous, and appear to
thrive inspite of the presence of Golden Eagles, Goshawks and the
other birds of prey. But these are places where there is not a
village within twenty miles and the people in those villages have
not yet aspired to gun licenses.
It is man, with his guns, nets and nooses who does most of the
damage. Before he came on the scene as a destructive agent,
Nature preserved her balance. Now since man has upset the
balance it behoves him to remedy the evil that he has brought about,
if we would preserve our game, and to do that it is necessary to
consider the whole position intelligently, and discover the real
agents of destruction and wage war on them, and not seek to destroy
an entire family simply because one member has proved a game-
destroyer.
Is it not time that the Society took up the question of not only
game preservation but the preservation of our fauna generally ?
India is dependent on its agriculture, and the thousands of birds, of
which the zamindar knows nothing and cares less, are one of the
main factors which protect his crops. Out of over 2,000 odd species
to be found inthe Indian Empire,a good sixty per centum are
insectivorous and therefore the zamindar’s staunch allies. There
are only avery small percentage that are actively destructive.
Under the Arms Act we have numbers of would-be shikaries who
are out to shoot something for the sake of shooting, in season and
out of season.
BIRDS OF PREY AND THEIR USES 743
Under the Wild Birds and Animals Protection Act, Government
can stop the shooting of any particular species, but cannot declare
a general close season. Is this of any use? You cannot do
anything toa man unless you actually catch him in the act of
shooting a protected bird, and what isto prevent him shooting a
dozen protected birds when no one is near, and producing an
‘unprotected’ bird when questioned ?.
A general close season when the carrying of a shot gun is an
offence under the Act is essential, except in the vicinity of crops.
Crop protection licenses might be issued free of charge, but the
moment the man is found using his gun outside his crops the license
should be cancelled and the arm forfeited.
Something on these lines will have a very beneficial effect but the
present system spells extermination of game within the next two
decades or so.
The rules and regulations of the Forest Acts can only apply to
protected forests, but what about the rest of the country where
the ‘ would-be’ shikari, to say nothing of the many criminal tribes
with their nets and nooses, can destroy what they like with
impunity ?
A NOTE ON THE BIRDS OF THE PESHAWAR DISTRICT
BY
Rev. F.S. BRIGGS AND B. B. OSMASTON, M.B.O.U., 1.F.S. (RETD.)
(With a map.)
The following notes on the birds of the Peshawar District have been combined
from the observations of the Rev. F.S. Briggs and Mr. B. B. Osmaston by
Mr. Hugh Whistler. The former spent six years at Peshawar, from 1920 to 1926.
The latter was there merely for a single cold weather from October 27, 1925
to March 27, 1926, but the sliortness of his stay was compensated for by the care
with which he made a collection of skins to identify the cold weather avifauna.
These skins have been enumerated under their respective species, and the
identification of the various racial forms is largely due to them. It is, however,
realized that this list of Peshawar birds cannot be considered complete. It is
proffered both as a record of the notes made by these two observers, and as a
groundwork for a fuller investigation of the avifauna of what, from its position,
will undoubtedly prove to be one of the most interesting and prolific districts of
India to the students of the palaearctic area.
Peshawar is situated in the notth-west corner of British India, N. Lat. 34°,
E. Long. 713°, in a wide plain traversed by the Kabul River, and surrounded by
hills.
This valley or plain measures some forty-eight miles from east to west, and
about thirty from north to south. Itis roughly level, at an altitude of about
1,100 feet above the sea.
For six or seven miles in all directions from Peshawar the valley is very
fertile owing to a verv complete system of irrigation by canals from the Kabul
and Bara Rivers, and magnificent crops of wheat, barley, maize, rice and
sugar-cane are produced.
Outside the irrigated area the country is virtually a desert, owing to the short-
ness of the rainfall, Peshawar being outside the area served by the South-west
Monsoon.
These dry desert areas bear little or nothing in the way of trees, vegetation
being restricted to a sparse growth of thorny shrubs. Woods and tree-growth
are restricted to the irrigated tract, including Peshawar Cantonment, which is
well wooded with a variety of trees. Elsewhere in the irrigated tract we find
extensive orchards of pears, quinces, peaches, oranges and pomegranates, and
along streams willows and mulberries, with roadside avenues of tamarisk and
sheesham.
The climate is one of extremes. The winter months, from October to March,
are cold, with regular night frosts fron: December to February, but with plenty
of warin sunshine in the day. The summer heat is severe and prolonged, and
frequently the temperatures registered at Peshawar are as high as, or even higher
than those in any other part of India.
The country round Peshawar falls naturally into three classes :—
1. Theirrigated tract, including Peshawar itself and its immediate sur-
roundings, characterized by field crops and aconsiderable quantity of tree
growth.
2. Thedry, desert country, surrounding the first, and extending to the
hills.
3. The swamp areas, found scattered in (1), and also including very large
beds of bulrushes, reeds and tall grass in the neighbourhood of the Kabul
River and its branches and backwaters. Unfortunately, from the point of view
of the ornithologist, a very large proportion of the reeds are cut and dried for
fuel and other purposes just at the time when they would provide nesting-sites
for several species of birds.
The birds which affect these three types of country are mostly quite distinct.
Journ. Bombay Nat. Hist. Soc.
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-A NOTE ON THE BIRDS OF THE PESHAWAR DISTRICT 745
1. Corvus corax (L). The Raven,
Although rarely seenin Peshawar itself the Raven is found in winter throug-
out the Peshawar Valley, being common a few miles out, for instance in the
more open stretches round Nowshera and Risalpur, near Jamrud, and along
the Khaibar Pass and Kohat Roads. During the hot weather it is only to be
seen in the valley on occasional cool days following rain, but it is at this season
very numerous at Cherat, and a few may be seen in the Khaibar Pass. A pair
build regularly on the cliffs on the Marble Rocks, between Nowshera and
Risalpur. The nest contained four newly hatched young and two infertile eggs
on March 6,1923, and three fresh eggs on February 8,1925. The birds which
laid in this nest were certainly C. c. /aurencit, but some of the Ravens seen in
the valley and more especially in the Khaibar Pass are very rusty in appearance,
and it is possible that C. c. ruficollis either breeds in the Pass, or wanders into
the valley. No specimens were, however, procured.
2. Corvus coronoides intermedius (Adams). The Himalayan Jungle Crow.
3o January 2, 1926, ¢ February 7, 1926, Peshawar. A single bird was shot by
Osmaston at Peshawar on February 7, feeding on the carcase of a buffalo.
There is some doubt as to whether it is this species or the Carrion Crow (Corvus
corone) which Briggs noticed as a regular but never numerous winter visitor to
the Peshawar Valley. He records it as always commoner in March and April
than at any other time, and then apparently migrating in a westerly direction
as small parties may be seen making their way up the Khaibar Pass.
3. Cervus frugilegus (L). The Rook.
3o & February 5, 1926, Peshawar. Abundant winter visitor, arriving towards
the end of October (earliest date October 23), and gradually becoming more
numerous ; practically all are gone by the end of March ; latest date observed
April 5. During the daytime the Rooks feed in the irrigated fields, and are
then rarely seen in Cantonments or villages. They collect, however, in vast
flocks at night to roost in the avenue trees in the Cantonments of Peshawar and
Mardan, though none seem to sleep in Nowshera or Risalpur.
4. Corvus cornix sharpii. (Oates). The Eastern Hooded Crow.
& December 21, 1926, 2 January 21, 1926. Peshawar; ¢ December 29, 1925,
Pabbi. A regular winter visitor ; earliest date October 23, latest date March 23.
It is found in pairs, or rarely in small parties, often associated with Rooks,
and keeps almost entirely to the wilder, more open, and barren parts of the valley
specially in the vicinity of swamps or the river. It is the wildest and most
wary of all the local crows.
5. Corvus splendens zugmayeri (Laubm.) The Sind House-Crow.
& December 29, 1925, Peshawar. Exceedingly common and resident in and
around Peshawar. It roosts separately from the Rooks. Eggs are mostly laid
in the first week of July. In February, 1921 there appeared to be an epidemic
among the species, and considerable numbers fell dead from their sleeping-
perches.
6. Corvus monedula semmeringii (Fischer). The Eastern Jackdaw.
@ February 5, 1926, Peshawar. A winter visitor from November to early
March, varying in numbers greatly from year to year. In the winter of 1920-21
they were numerous, and very numerous again in the winter of 1925-26,
whereas in the winter of 1921-22 Briggs only saw them once, namely on
January 1. They feed in the fields with the Rooks, and also roost with them.
7. Nucifraga multipunctata (Gould). The Larger Spotted Nutcracker.
A straggler, probably of this species, which is the form found on the Sufed
Koh to the west and in Hazara to the north, was seen by Briggs in Peshawar
on April 23, 1920. The previous night had been very stormy.
8. Parus major kaschmiriensis (Hartert). "The Kashmir Grey Tit.
© October 31, 1925. Q@ November 6, 1925, Peshawar. A fairly common
winter visitor to the Peshawar Valley; earliest date October 21, latest date
March 30. It keeps chiefly to the wooded areas.
9. Parus monticolus (Vigors). ‘The Green-backed Tit.
A winter visitor, less numerous than the last species. Earliest date October
13, latest date March 14.
iy
745 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
10. A€githaliscus concinnus (Gould). The Red-headed Tit.
A party of about six were seen by Briggs at Cherat about 4,500 on Angust
22, 1920,
Il. Argya earlii (Blyth). The Striated Babbler,
Rather rare, but probably a resident. Anest with fresh eggs was taken by
Briggs in swampy ground at Rashkai near Mardan, on May 29, 1922.
12. Argya caudata. The Common Babbler.
& 2 November 16, 1925. ¢ Q@ November 18, 1925. @ December 14, 1925,
Peshawar.
A common resident, numerous in the tong grass and tamarisk near streams,
and, as elsewhere, found in loose parties. In winter it is more numerous in
gardens than in summer. While breeding it shows a preference for ever-
greens. Fresh eggs found between March 21 and August 19 but mainly in
April.
13. Molpastes hemorrhous intermedius(Jerdon). The Punjab Red-vented Bulbul.
3 2 November 16, 1925. Common in Peshawar and in the surrounding
woods. Doubtless resident, but Briggs notes :—‘ In the unusually dry and hot
summer of 1920, Red-vented Bulbuls became scarce about the middle of July
and remained so till the evening of September 29. On that evening, about
seven o’clock, a large flock of white termites appeared in Peshawar, with
numbers of Bulbuls in pursuit of them. For about twenty minutes not less
than thirty Bulbuls were busily employed in my garden feeding on the termites
which they caught in the air afterthe manner of Flycatchers. The next day
Bulbuls seemed to be about the same numbers as they had been the previous
winter, and I noticed no great fluctuation in numbers afterwards.’ Fresh eggs
found between March 13 and July 12.
14. Molpastes leucogenys leucogenys (Gray). The White-cheeked Bulbul.
& 6 November 13, 1925. 2 November 18, 1925. 4 December 7, 1925. ¢
December 13, 1925.
15. Molpastes leucogenys humii (Oates). Hume’s White-eared Bulbul.
3 2° January 11, 1926. Twenty miles south-east of Peshawar.
Osmaston collected a nice series of ten of these Bulbuls in the winter of
1925-26, and states that they are common, and found to some extent inthe dry
tracts, though mainly in the irrigated area. He suspects that they may be a
winter visitor. Briggs’ longer observations enable him to amplify this as
follows :—‘ Very numerous in winter, especially in the latter part. In the
summer this bird retires from the lower parts of the valley to the hills
surrounding it. I have found them breeding at Attock at about 1,000 feet,
below Cherat at about 2,000 feet and near Ali Musjid at about 2,500 feet. It is
surprising that the White-cheeked Bulbul of the Peshawar Valley should be
leucogenys and not Aum.’ He further adds that he saw two Bulbuls outside
Peshawar on the Jamrud road on January 18, 1920, which he identified as J/. Z.
loucotis.
An examination of Osmaston’s series shows that all the specimens from
Peshawar are clearly J/. /. leucogenys, and we are justified in describing that
form asa regular and common winter visitor to Peshawar. ‘Three birds, how-
ever, obtained in the dry foot-hills 20 miles to the south-east of Peshawar are
closer to 17. l. humit, and doubtless represent the resident form. We have been
compelled for the identification of these birds to examine the British Museum
series as well as thirty birds in Mr. Whistler’s collection, as the account of
these forms in the second edition of the Fauna is not very satisfactory, and
superficially they are difficult to understand owing to the amount of variation
in the individual. It may be as well, therefore, to explain the position.
The Yellow-vented Bulbuls of North-west India form an excellent example
of a species, divided into subspecies, and connected by intermediate forms.
Gray in 1830 described the hill bird from Darjeeling and in this species,
curiously enough, although the bird is so variable in the north-west generally,
birds from the Eastern Himalayas are precisely similar to birds from the
Western Himalayas as leucogenys, while Gould in 1836 described the plains
bird as deucotis; its type locality has since been restricted to Karachi,
Sind, .
A NOTE ON THE BIRDS OF THE PESHAWAR DISTRICT 747
If Sind and Himalayan birds alone are contrasted, we have two. very
distinct birds, which might well rank as full species, with no danger of
confusion between them.
Leucogenys is a large bird with a long, comparatively slender bill, compared
with the smaller /ewcotis and its short bill.
Leucogenys : Bill from skull ¢ 18-20 mm. ; 2 17°5-19'5.
Wing ¢ 88°5-98 ; 2 88-96°5
Leucotis. Bill from skull J 17 mm. ; Q 16°5-17:
Wing 87-88 ; 2 81°5-85
In leucogenys the crest is so long and so curved forward over the beak that
it resembles a fool’s cap in shape—a peculiarity most marked in life ; the colour
is hair brown, each feather being narrowly edged with greyish white. ‘here is
a white line above the lores. The upper plumage is earth-brown tinged with
olive-brown and even olive green. ‘he under tail coverts are bright sulphur
yellow.
In Jeucotis the crest is so short as to be almost non-existent ; it is entirely
black, and there is no white line above the lores. The white cheek-patch is
twice as large as in the other bird, and the black on the throat is more sharply
defined, contrasting with the whitish lower plumage. The upper plumage is
plain fawn brown. ‘The under tail coverts are rich saffron yellow.
But between these forms a regular gradation may be found in the triangle
of country which extends north-west of the Salt Kange to the Afghan ranges
and to the Himalayas. The elevation here runs between 1,000 and 3,000 feet
and the Bulbuls from this area approximate more closely to /ewcogenys than to
leucotis, in that all have the under tail coverts bright sulphur yellow, and in
their size. Thecrest and loral line are variable, as is also the colour of the
upper parts.
It so chances that in 1889 Oates picked out a single bird from this intermediate
area, from Jalalpur, Jhelum District, on the southern border of the triangle,
and made it the type of his Wolpastes humit, so this name is available for the
intermediate forms. It must be remembered, however, that this race 17. 1. humiz
being a true ‘intermediate is variable, approximating on either side to the two
races which it connects. This approximation follows fairly definite lines.
Birds from the Jhelum Salt Range agree fairly closely with Oates’ type, which
in its shorter black crest without pale edging and with practically no trace of
the white loral line, in the size of the white cheek-patch, and in the fawn colour-
ed upper plumage, is very close totrue /eucotis. Birds from Kohat and Bannu
are in turn closer to Aumiz as are Osmaston’s three birdsfrom the dry foothills
about Peshawar. At Attock, Campbellpur, and Rawalpindi, however, we
find, as might be expected, that the birds are much closer to the true
leucogenys, which also probably in winter descends from the Himalayas and
mixes with them. Indeed they areas close to dewcogenys as Oates’ type is to
leucotis, and a determined ‘splitter’ might name them so as to make two
groups of races merging into one another.
This explains the situation at Peshawar. The resident bird belongs to the
drier areas aud low ranges, and is of the intermediate race Aum7z ; itis, how-
ever, obscured in winter by a regular immigration to the fertile irrigated area
of the Himalayan race, leucogenys.
16. Certhia himalayana (L). The Tree-Creeper.
& November 20, 1925. 2 November 24, 1925, Peshawar. A fairly common
winter visitor, found in avenues, woods, and gardens, Earliest date seen
October 17, 1925, latest date March 14, 1925. Briggs found one caught in a
spider’s web and entirely unable to free itself,
17. Tichodroma muraria (L) The Wall-Creeper,
A numerous winter visitor to the sandy nullahs and rocky gorges that surround
the Peshawar Valley, such as the Khaibar at Ali Musjid, Attock and the
Kalapani River at Risalpur. Peshawar itself is not suitable to its requirements.
18. Saxicola caprata bicolor (Sykes). The Pied Busn-Chat.
& December 30, 1925,,Peshawar. Mainly a summer visitor, though a few birds
Winter in the valley. ‘The earliest and latest dates for the mass are February 28
and October 1, In the valley eggs may be found commonly between April 7
and May 17, but Briggs found two nests with eggs on the point of hatching
on July 11, 1922, The majority of the birds of this species appear, after breeding
748 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXX11-
in the valley, to take their young, as soon as they can fly, to the lower hills
round the Peshawar Valley.
19. Saxicola torquata indica. (Blyth). The Indian Stone-chat.
@ November 9, 1925, 2 November 20, 1925, g November 26, 1925, Peshawar.
Mainly a spring and passage migrant, noted by Briggs as passing in spring
between March 17 and May 24, and in autumn between September 24 and
October 10. A few however, remain in winter.
20. Saxicola torquata leucura (Blyth). The White-tailed Stone-chat.
3 December 12, 1925, Peshawar. Osmaston collected a pair at Peshawar on
December 9, 1925, and his statement that a Stone-chat is not uncommon through-
out the winter, especially in grassy areas near swamps probably refers partly to
this race.
21. (Enanthe picata (Blyth). The Pied Wheatear.
According to Briggs a regular winter visitor to the Valley, keeping mainly
to barren country : earliest and latest dates seen are September 8 and March 8.
It breeds in the Khaibar Pass, where Briggs on several occasions’ saw it carrying
nesting material, but never had the opportunity of examining a nest.
22. (Enanthe capistrata (Gould). The White-headed Wheatear.
& November 15, 1925, ¢ December 6, 1925. According to Briggs, breeds in all
the hills round the Peshawar Valley down to about 2,000 ft. A pair were watched
at work on a nest in the Manki Pass, a few miles S. of Nowshera, on March 21,
1923. Thenest was then almost complete. It is much commoner as a winter
visitor to the Peshawar Valley than the last species. Briggs notes that ‘ Indivi-
duals of this species seem to be much attached to limited localities in their
winter area. Forexample, onthe road between Nowshera and Risalpur I
could be certain of seeing four birds of this species any day in the cold weather.
Near a lime kiln there was always a pair of them, male and female, and within
fifty yards of a certain tree there was always asingle male. There was always
another single male in the compound of the house at Risalpur where I usually
stayed. I always looked for him on the telegraph wires outside the house on
my arrival, and nearly always saw him there.’ Briggs’ earliest date for the
species is September 18. He, like other observers in other areas, remarks on the
preponderance of males in winter both in @. pzcata and @. capistrata.
23. (Enanthe opistholeuca (Strickland). Strickland’s Wheatear.
& December 21, 1925, § December 11, 1925. Not recorded by Briggs, but said
by Osmaston to occur in the dry country in winter, though not so commonly
as the last species.
24. (Enanthe isabellina (Cretsz). The Isabelline Wheatear.
2 November 15,.1925, Peshawar. gJanuary 25, 1926, Risalpur. Fairly common
locally in winter, generally in the neighbourhood of cultivation, in the dry,
desert land at the foot of the hills.
25. (CEnanthe xanthoprymna chrysopygia (De Filippi). The Red-tailed Wheatear.
Q2 December 17, 1925, Bara Fort, five miles. south-west of Peshawar. Fairly
common in the Khaibar, and probably breeds there. In the winter it is not rare
in the dry, desert country below the foothills.
It is noteworthy that neither Osmaston nor Briggs record the desert Wheater,
CE. desertt, which is so common a winter visitor to the Punjab.
26. Phoenicurus erythronotus (Evesrm.) Eversman’s Redstart.
3d January 22, 1926, Dilazak, 9 December 7, 1925, 2 December 10, 1925, Pesh-
awar. Stated by Osmaston to be seen occasionally throughout the winter in the
irrigated area.
27. Pheenicurus ochrurus pheenicuroides (Moore). The Kashmir Redstart.
A common winter visitor, but for some reason less numerous than usual in
the winters of 1924-25 and 1925-26. Earliest date seen September 24, latest
date April 17. It occurs both in the wet and dry areas, but, according to
Briggs, shows a strong preference for the neighbourhood of Tamarisk trees.
28. Chaimarrhornis leucocephala (Vigors). The White-capped Redstart.
One was seen by Briggs in the Khaibar Pass on December 12, 1924.
A NOTE ON THE BIRDS OF THE PESHAWAR DISTRICT 749
29. Rhyacornis fulginosa (Vigors). The Plumbeous Redstart.
Recorded by Briggs as fairly common on the Khaibar Stream during tHe cold
weather.
he Cyanosylvia suecica pallidogularis (Sarudny). The Eastern Red-spotted Blue-
throat.
3 October 31, 1925, ¢ November 18, 1925, ¢¢ November 28, 1925, ¢ December
Oy 1925; Peshawar, 6 January 22, 1926, 2 December 29, 1925, Pabbi. Very
numerous on passage, passing’ through the valley between ‘April 10 and May 10,
in spring, and between September 14 and October 14in autumn. During the
spring migration they seem to pass through almost entirely in pairs, and may
frequently be seen courting. In autumn they are less numerous, and seem as a
rule to make a shorter stay. In the winter they are not uncommon in the
irrigated country in dense cover such as sugar-cane, and grass and bulrushes
especially near swamps.
31. Saxicoloides fulicata cambaiensis (Leth.). ‘Che indian Robin.
& November 15, 1925, Peshawar. A resident, but more numerous in summer
than in winter. In this area (asat Kohat), it avoids the neighbourhood of
houses, and prefers barren country, especially the stony Mahomedan
graveyards. Eggs found by Briggs between April 18 and May 24.
32. Turdus atrogularis (Temm.). The Black-throated Thrush.
& November 10, 1925, ¢ December 3, 1925, ¢ Q December 8, 1925, Peshawar.
A very common winter visitor, found chiefly in and around the station area and in
orchards. It is most numerous in February, and the latest date on which it is
recorded is March 28.
33. Turdus viscivorus (L). The Missel Thrush.
One was seen by Briggs at Peshawar on February 2, 1921.
34. Oreocincla dauma (Lath.). The Small-billed Mountain Thrush.
One was noted by Briggs on February 9, 1924.
35. Monticola solitaria pandoo (Sykes). The Blue Rock Thrush.
Not found in Peshawar according to Osmaston, but observed in the neigh-
bouring rocky hills, Khaibar, etc.
36. Myiophoneus temminckii (Vigors). The Himalayan Whistling Thrush.
Not found in Peshawar according to Osmaston, but seen on the streams in the
neighbouring foothills.
37. Prunella atrogularis (Brandt). The Black-throated Accentor.
¢ January 1, 1926, Peshawar. Not uncommon, according to Osmaston, in
January and February in Pirbala and other woods.
38. Musicapa striata neumanci (Poche). The Spotted Flycatcher.
Briggs records that he saw a flock of these Flycatchers (and was certain that
they were not Alseonax latirostris) feeding in his compound at Peshawar on
April 19, 1921, and again from October 6 to 16, 1921, and on April 4, 1922.
39. Musicapa hypoleuca (Pallas). The Pied Flycatcher.
Briggs writes :—‘ A few birds indistinguishable from the British species pass
through Peshawar almost every spring. Earliest and latest dates seen February
18 and March 30. The only year that I have seen the species in the autumn was
1925, when a pair fed in my compound almost every day from October 31, till I
left Peshawar on January 11.’ The species has not yet been included in the
Indian list, so future observers at Peshawar should make every effort to verify
this occurrence by obtaining specimens. The species does not appear to have
been recorded nearer than Tehran.*
40. Siphia parva parva (Bechst.). The Red-breasted Flycatcher.
3b October 29, 1925, {November 8, 1925, 2 December 1, 1925, Peshawar. Chiefly
a passage migrant, the majority passing through in spring between March 24.
and April 21. First noted on the autumn migration on October 21. After the
first rush their numbers gradually decrease until only a few remain for the
tS. BON AS., vol. xxvi1, ps 114,
750 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXI/
winter. They are found in the orchards and station area, and along all roads
bordered with Tamarisks. Red-breasted males are very rare amongst the winter
birds.
41. Culicicapa ceylonensis. The Grey-headed Flycatcher.
Q November 7, 1925, Peshawar. The above specimen was obtained by
Osmaston.
42. Terpsiphone paradisi (I.). The Paradise Flycatcher.
A few breed in the valley, as Briggs founda female feeding three young in
the trees overhanging the Grand Trunk Road, eight miles east of Peshawar on
June 20, 1923. it is, however, a passage migrant in spring and autumn.
Briggs states that they pass through in large numbers, but appear to stop only
one day. That day falls, as far as his observations go, between April 23 and
29, in thespring, and between September 19 and 29, in the autumn. The
race has not been identified.
43. Lanius excubitor lahtora (Sykes). The Indian Grey Shrike.
o ? January 13, 1926. Not common, but appears to be resident. It is confined
to barren stretches of country near the foot of the hills, showing a preference for
ground studded with stunted Ber trees.
44. Lanius vittatus (Valenc.). The Bay-backed Shrike.
Asummer visitor, very common in cultivated ground. Earliest date seen
March 23, latest date September 19. It has two broods, building a fresh nest
foreach. Eggs found between April 7 and July 7. Stragglers were seen by
Briggs near Taru Jabba on February 20, 1923 and January 25, 1924.
45. Lanius schach erythronotus (Vigors). The Rufous-backed Shrike.
& November 13, 1925, Peshawar. Resident, but much more common in
summer than in winter and a bird of the irrigated tract. A male of this species
was observed by Briggs to feed in his garden almost daily for the whole of the
six years he was in Peshawar. Forthe greater part of the year this bird was
alone, but, in the spring he was joined by a female, generally about the middle
of March. Each year they built either in his garden or the one next to it, and
usually succeeded in bringing offtwo broods of young. Once or twice they
used the same nest for both broods. The young of the first brood always dis-
appeared before the second brood had left the nest, and as soon as the second
brood was able to fly at all strongly. generally some time in August, they and the
hen disappeared. It is, of course, unsafe to generalize from a single case of
this sort, but it might be worth while to shoot a few Rufous-backed Shrikes in
winter with a view to discovering whether the decrease in their numbers each
winter is due to the migration of thefemales. It is to be noted that the one
specimen obtained by Osmaston, in November, was a male.
Osmaston remarks that the bird is a wonderful mimic ; he heard it giving in
quick succession an accurate imitation of the notes of the following :—House
Sparrow, Swallow, Indian Swift, Parrakeet, Mynah, Sandpiper, Bulbul and
Common Babbler. Eggs found between April 10 and July 1.
46. Lanius cristatus isabellinus (Hemp. and Ehr.). The Isabelline Shrike.
3 November 30, 1925, ¢ January 26, 1926, Peshawar. The above two
examples, procured by Osmaston, are the only records.
47. Tephrodornis pondiceriana (Gmel.). ‘The Common Woodshrike.
One was seen by Briggs on the outskirts of Peshawar on March 7, 1921.
48. Pericrocotus brevirostris brevirostris (Vigors). ‘The Short-billed Minivet.
© November 10, 1925, ¢ December 1, 1925, Peshawar. An erratic winter
visitor, appearing for aday or two at atimein small flocks, never more than
ten strong.
49. Dicrurus macrocercus ( Vieill.). ‘The King Crow.
A common summer visitor, laying in the first half of May. Earliest date
seen March 22, latest date October 1.
50. Dicrurus'leucocephalus longicaudatus (A. Hay). ‘The Ashy Drongo.
A pair was seen by Briggs in the Khaibar, May 28, 1925.
A NOTE ON THE BIRDS OF THE PESHAWAR DISTRICT 751
51. Acrocephaius stentoreus brunnescens (Jerdon). The Indian Great Reed
Warbler.
3d November 5, 1925. The above specimen shot by Osmaston was alone ina
swamp. Hedid not meet the species again till March 15, when he saw and
heard others, doubtless on migration.
52. Lusciniola melanopogon mimica (Madrasz). The Moustached Sedge
Warbler.
d November 5, 1925, 0? January 22,1926, Peshawar, 2 December 29, 1925,
Pabbi. Obtained by Osmaston, who found it common and noisy, though
difficult to observe in the swamps, especially in borrow-pits by the railway.
53. Cisticola juncidis cursitans (Frankl.). The Fantail Warbler.
3g November 12, 1925, Q December 4, 1925, Peshawar. Found by Osmaston
in some of the grassy swamps, but stated to be not very common.
54. Franklinia buchanani (Blyth). The Rufous-fronted Wren Warbler.
G November 15, 1925, Peshawar. The above specimen obtained by Osmaston
in the dry country supplies the only record.
55. Sylvia curruca affinis (Blyth). The Indian Lesser Whitethroat.
3 November 5, 1925, 2 November 6, 1925, ¢ December 11, 1925, Peshawar.
A fairly common winter visitor.
56. Sylvia curruca minula (Hume). The Small Whitethroat.
& December 11, 1925, Peshawar. The above specimen was obtained by
Osmaston.
Briggs reports that individuals of one or other of the above races of S. curruca
spend the summer in the Tamarisk scrub on the banks of the Kalapani River.
Presumably they are non-breeders.
57. Phylloscopus collybita tristis (Blyth). The Siberian Chiff-chaff.
& October 29, 1925, 2 November 3, 1925, 4 November 8, 1925, 9 December
11, 1925, Peshawar. A very common winter visitor, especially in the wet
country. It is to be found in every clump of trees, also in road-avenues of Shi-
sham and Tamarisk, and often feeding by streams. Thesong may be heard
in April before the birds leave for their summer quartters.
58. Phylloscopus subviridis (Brooks). Brooks’ Willow Warbler.
& November 11, 1925, Peshawar. A winter visitor, and probably not
uncommon.
59. Scotocerca inquieta striata (Brooks). The Streaked Scrub Warbler.
A resident and fairly numerous on the dry, rocky hill sides on the spur which
separates Kohat from Peshawar, also on dry ground in the vicinity of Risalpur.
Osmaston found a nest with six eggs in the former place on March 12, 1926, and
Briggs found a nest containing one young bird on the point of leaving the nest
near Risalpur on April 15, 1925.
60. Horornis pallidus (Brooks). ‘The Pale Bush Warbler.
Observed by Osmaston towards the end of February in the Pirbala woods, its
note being quite unmistakeable.
61. Cettia cetti cettioides (Hume). The Eastern Cetti’s Warbler.
do October 31, 1925, 4 November 23, 1925, ¢ January 9, 1926, ¢ January 12,
1926, ¢ January 25, 1926, ¢ January 26, 1926, Peshawar. Fairly common,
and widely distributed in every swampy patch, provided there is ample cover
of bulrushes, reeds, or long grass. They are rarely seen, being extremely shy
and careful not to expose themselves. Their presence is, however, revealed by a
rather loud ‘ click’ repeated at intervals from dense cover. The above series
was only obtained by the exercise of much patience and preseverance
62. Prinia gracilis lepida (Blyth). The Streaked Wren Warbler.
Q@ November 26, 1925, ¢Q November 30, 1925, Peshawar, ¢ January 28,
1926, Risalpur. Generally distributed in and around grassy swamps as well
asin cultivated fields.
752 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
63. Prinia inornata inornata (Sykes). The Indian Wren Warbler.
@ November 5, 1925, 9 November 10, 1925, 9 November 11, 1925, ¢ 2 Novem-
ber 16, 1925, § November 26, 1925, ¢ December 26, 1925. A common bird in
long grass in moist localities, occurring in the winter in small parties.
64. Oriolus oriolus kundoo (Sykes). The Indian Oriole.
Status uncertain. Briggs says :—‘ Common in April and May. A few pairs
may breed in the valley. One young bird seen on the outskirt of Peshawar,
August 31, 1921.’
65. Pastor roseus (L). The Rosy Starling.
& November 7, 1925, Peshawar. ‘ Passes through in large flocks in the
late summer and early autumn. The earliest flocks to arrive are composed
almost entirely of adult males. Inthe later flocks young birds and females
predominate. They appear tofeed mainly on the fruits of various species of
Ficus. Inspring they appear either to gototheir breeding area by another
route, or to pass through the valley without halting. I have only seen them
in the spring twice; on April 13, 1924, when a small flock left a group of
trees at dawn and flew steadily westward ; and in April 1925, when large flocks
passed through between the 21st and 25th of the month. Earliest date seen
on the southward migration July 14; latest date October 21.’—(Briggs). In
1925, however, Osmaston saw small flocks early in November.
66. Sturnus vulgaris porphyronotus (Sharpe). The Central Asian Starling.
3&3 November 18, 1925, { November 21, 1925, ¢ December 2, 1925. Peshawar.
67. Sturnus vulgaris poltaratzskyi (Finsch). Finsch’s Starling.
3 November 7, 1925, # November 21, 1925, 4 November 24, 1925, ¢ Decem-
ber 2, 1925, Peshawar.
68. Sturnus vulgaris humii (Brooks). Hume’s Starling.
Under the heading Sturnus humii the following appears in Vests and Eggs
of Indian Sirds, Hume and Oates, vol. i, p. 369 :—This Starling lays towards
the end of April at Peshawar, where I found it nesting in willow-trees in the
Cantonment compounds. (See also /6zs, 1870, p. 529.)
Both S. v. dorphyronotus and S. v. poltaratzskyi are numerous as winter visi-
tors, and they flock together. The earliest date on which Briggs saw Starlings
was October 12, and the latest date April 13, but most years all have left before
the end of March. Neither Osmaston nor Briggs identified S.v. humiz, and the
latter found no Starlings breeding in the valley, though after reading the above
quotation from Hume he looked for them for six successive seasons.
69. Acridotheres tristis tristis (L). The Common Mynah.
Exceedingly common and aresident. Eggs were found froin the middle of
April to the end of August, there being at least two broods. .
70. Acridotheres ginginianus (Latham). The Bank Mynah.
@ November 7, 1925, Peshawar. A summer visitor. Breeds in very large
colonies in the sand-cliffs near Risalpur. Fresh eggs found in great numbers
on May 9, 1922, and May 15, 1923. In the autumn mixes in small flocks with
A. tristis before departing. Earliest date seen March 22, latest date October
12 (Briggs). Osmaston, however, saw small parties about on his first arrival
in 1925, but not after the middle of November.
71. Uroloncha malabarica (L). The White-throated Munia.
@ November 6, 1925, Peshawar. A resident, but not very numerous.
Rarely seen near houses.
72. Amandava amandava(L). The Red Avadavat.
&% 2 November 14, 1925, Peshawar. ‘ Not common, but small parties seen
on grasslands on two or thee occasions.’ (Osmaston). Many are sold in the
bazaars at Peshawar, and Briggs remarks on the ease with which they escape
from their cages.
73. Caropdacus erythrinus (Pall). The Common Rosefinch.
‘Pass through Peshawar in flocks, staying for a few days in spring. I have
never seen them on their autumn journey. They seem to time their arrival at
Peshawar to coincide with the ripening of the mulberries, on which they feed.
Their numbers vary greatly from year to year. In 1924 the mulberry crop was
A NOTE ON THE BIRDS OF THE PESHAWAR DISTRICT 753
good, and the Rosefinches numerous: in 1925 the mulberry crop was poor,
and Rosefinches far less numerous. Earliest date seen April 18, latest date
April 30. (Briggs).
74. Metaponla pusilla (Pall.). The Gold-fronted Serin.
& November 25, 1925, Peshawar. The above young bird, secured by
Osmaston, was a solitary straggler.
75. Hypacanthis spinoides. (Vigors). ‘The Himalayan Greenfinch.
Q November 25, 1925. Osmaston secured the above specimen from a small
flock, the only lot seen.
76. Gymnorhis xanthocollis (Burton). The Yellow-throated Sparrow.
A common summer visitor, first observed on April 10. Numerous nests found
between May 16 and 23, all in holes in Tamarisk trees.
77. Passer domesticus (L). The House Sparrow.
@ December 6, 1925, ¢ @ December 13, 1925, fg ¢ 2Q January 4, 1926,
Peshawar. Avery numerous resident, breeding from the middle of March to
the end of september or beginning of October. Although commonly building
in houses, it frequently builds in trees in the Peshawar Valley, a trait shared by
the sparrows of the Quetta Valley, but not usual in the plains of India. Ber
trees are preferred. In winter large flocks are found far out froni the station in
the irrigated land.
78. Passer hispaniolensis (Tschusi). Tschusi’s Sparrow.
Recorded by Hume (/dzs., 1870) as being in breeding condition in the
Peshawar Valley in April. It was not observed by Osmaston or Briggs.
79. Emberiza leucocephala (Gmelin). The Pine Bunting.
& November 29, 1925, ¢ § December 30, 1925, ¢ ¢ January 4, 1925,
Peshawar. A fairly common winter visitor, in November December, and
January, occurring in rather large flocks both in swampy ground and _ in fields
in the cultivated tract.
80. Emberiza scheeniclus pallidior (Hartert). The Reed Bunting.
¢ January 9,1926, ¢ February 8, 1926, Peshawar. Found by Osmaston to
be locally common in winter occurring in small flocks in the wet country.
8!. Emberiza cia par (Hartert). The Meadow Bunting.
Q November 14, 1925, ¢ November 15, 1925. A fairly common winter visitor
both to the irrigated tract and tothe uncultivated country, especially up the
Khaibar and at Attock.
82. Emberiza melanocephala (Scop.) The Black-headed Bunting.
Briggs notes that a small flock fed daily in the barley fields outside Peshawar
from May 7 to May 20, 1920.
83. Emberizaicterica (Eversm). The Red-headed Bunting.
Passes through in large numbers in the spring, earliest and latest dates
April 23, and May 10. Not seen in the autumn, except fora single bird on
the outskirts of Peshawar Cantonment on August 13, 1923.
84. Riparia riparia indica (Ticehurst and Whistler). The Small Sand-Martin.
o? December 9, 1925, o? January 1, 1926, Peshawar. Omaston found this
form common along the Kabul River and its branches and canals in December,
when it was ‘singing’. Briggs says that itis a resident and numerous, but
subject to local movement within the Valley in relation to the supply of
insects. Eggs found February 14 to March4. They breed in suitable spots
in the banks of the Kabul River, and in large colonies in the same _ sand cliffs
near Risalpur which are later occupied by Bank Mynahs. He believes that
the Bank Mynhas enlarge and use the hole of the Sand Martins frequently.
85. Ptyonoprogne rupestris (Scop.). The Crag Martin.
A few were seen by Osmaston in November, 1925.
86. Hirundo rustica rustica (L). TheSwallow. .
3 b&b 2 December 6, 1925. Peshawar. Osmaston states that he found it
common tbroughout the winter months. The above five specimens were
secured at asingle shot as they sat ona telegraph wire. Briggs says :—Pass
through in large flocks in spring and autumn, the autumn flocks. frequently
18
754 JOURNAL, BOMBAY NATURAL AIST. SOCIAL] Vi Volyeexd 1
arriving after rain on the hills to the north west. Some are to be seen
throughout the year. Those which remain through the summer are probably
non-breeders, as I have never seen or heard of anest, though I once saw one
picking up mud from the side of a pool as though to build. It is also
possible, however, that some, if not all, of those seen in the summer breed in
the hilis, and come downto the valley during the day forfood. In summer
they always appear to be more numerous after rain.
87. Hirundo smithii filifera (Stephens). The Wire-tailed Swallow.
A fairly numerous summer visitor ; earliest date seen April 21. Eggs found
from May !, onward, and young seen in the nest as late as September 27. This
species sometimes uses the same nest for more than one season, if it is not
destroyed by floods. Briggs notes that every nest of this species that he
examined was in a position where it was liable to be flooded out, in low
culverts. In the summer of 1925, when an unusual number of heavy thunder-
storms visited the Peshawar Valley no nest escaped destruction before the
young were ready to fly with the exception of one found containing young on
September 27.
88. Hirundo daurica (L). The Striated Swallow.
A small flock of Striated Swallows (subsp. ?) was seen by Briggs hawking
for insects over the Mall at Nowshera on April 13, 1923.
89. Motacilla alba dukhunensis (Sykes). The Indian White Wagtail.
o? November 10, 1925, ¢ November 11, 1925, ¢ 2 December 15, 1925,
Peshawar. A very common winter visitor found in the irrigated area. Earliest
date seen September 10, latest date May 4.
90. Motacilla alba personata (Gould). The Masked Wagtail.
A fairly numerous winter visitor. Earliest date seen August 21, latest date
May. 11.
91. Motacilla cinerea caspica. (Gmelin). The Grey Wagtail,
A fairly numerous winter visitor. Earliest date seen September 22, latest
date May 9.
92. Motacilla flava thunbergi (Billberg). The Grey-headed Wagtail.
Large flocks pass through in the spring and autumn. and a few stay for the
winter. Earliest date seen September 6, latest date May13. In the autumn
all the flocks appear to be composed of both young and old.
93. Motacilla citreola calcarata. Hodgson’s Yellow-headed Wagtail.
& Deow2l,.1925:2 Dec. 21; A925:
Occasionally seen throughout the winter ; most numerous in March and
April. A few, probably non-breeders, stay all the year round by the Kalapani
River.
94. Authusrichardi rufulus (Vieill.). ‘The Indian Pipit.
o ? November 27, 1925, Peshawar. Not very common, but occasionally
seen in grasslands near cultivation. Briggs found a nest with two partially
incubated eggs on April 12, 1921.
95. Anthus campestris (L). The Tawny Pipit.
3d December 11, 1925, Peshawar. Osmaston says that it is not common, but
that he sawit occasionally near the meeting place of the wet and the dry
country.
96. Anthus spinoletta blakistoni (Swinhoe). The Water Pipit.
& November 2, 1925, ¢ November 11, 1925, c ? November 14, 1925, ¢ 2 De-
cember 15, 1925, ¢ December 31, 1925: Peshawar. An exceedingly common
winter visitor, found about swamps, and in moist irrigated fields.
97. Alauda gulgula (Frankl.). The Smaller Skylark.
Apparently breeds in the district as ou April 16. Briggs found a clutch
of five partly incubated eggs at Pajaggi which he attributes to this species.
98. Alauda arvensis dulcivox (Brooks). The Eastern Skylark.
. § December 12, 1925, Peshawar. A common winter visitor, found in flocks.
in the ffelds. -
ao
“at MOTE ON THE BIRDS OF THE PESHAWAR DISTRICT 755
99, Calandrella raytal adamsi (Hume). The Indus Sand Lark.
3d 2 December 21, 1925. . Pabbi ; February 25, 1926. Peshawar. Found by
Osmaston in flocks in December, feeding near swampy ground.
100. Calandreila rufescens persica (Sharpe). Sharpe’s Short-toed Lark.
. o-January 28, 1926, Risalpur. The above specimen was.obtained by
‘Osmastan from a small flock which was very wild.
101. Galerida cristata chendoola (Frankl.). Franklin’s Crested-Lark.
& November 3, 1925, ¢ 2 December 3, 1925, Peshawar. Osmaston says that
itis widely distributed though not common. Briggs says that it is resident and
numerous, but that a great number appear to move to the higher ground sur-
rounding the valley after bringing off their young. Eggs found throughout
April.
102. Galerida cristata magna (Hume). Hume’s Crested Lark.
Briggs says :—‘ A winter visitor, most numerous in spring and autumn when
on passage.’
103. Ammomanes deserti phoenicuroides (Blyth). ‘The Desert Finch-Lark.
-A resident, and fairly numerous on barren ground. Eggs found between
‘May 29 and July 4,
104. Zosterops palebrosa occidentaiis (Ticehurst). The White-eye.
‘--9 November 8, 1925. Fairly common in the wooded areas in winter. Briggs
remarks that it appears spasmodically in large flocks, especially after rain.
105. Cinnyris asiatica (Latham). The Purple Sunbird.
A common summer visitor. Earliest date seen March 17 (apart from a
straggler, a male in breeding plumage, seen by Briggs inthe Khaibar on
February 22, 1921). Latest date seen September 11. Briggs found fresh eggs
from April 29toJunel, and heard of earlier and later occurrences. He
notes that this species, if undisturbed, appears to use the same nest fora
second brood. ,
106. Picus squamatus (Vigors). The Scaly-bellied Green Woodpecker.
.Seen on two occasions by Osmaston in Pirbala wood in January 1926.
107. Dryobates scindianus (Horsf. and Moore). The Sind Pied Woodpecker.
re November 25, 1925, & & December 26, 1925, Peshawar. A common resi-
dent in all wooded areas in the irrigated tract. The nest-holes are commenced
in March.
108. Liopicus mahrattensis aurocristatus (Tickell), The Mahratta Woodpecker.
¢ January 1, 1926, Peshawar. A common resident, found in the same area as
the last species.
109. lynx torquilla (L). The Wryneck.
& November 13, 1925, 2 Novem be 20p dd2o, 2 January fs 1926, 2 January 22,
1926, Peshawar. A fairly common winter visitor, and rather more numerous on
passage according to Briggs.
- 110. Coracias benghaiensis benghaiensis (L). The Blue Jay.
A resident in small numbers ; its numbers are greatly augmented in summer
by an immigration about March. Eggs found between April 21 and May 25.
On May 15, 1923, Briggs found a Roller sitting on four of its own eggs and one
Bank Mynah’s egg, in a colony of the latter species.
W121. Coracias garrula semenowi (Loud. and Tsc). The Kashmir Roller.
Passes through in large numbers in spring and autumn. A few stay to breed
usually keeping well away from towns and villages. Eggs found May 10, 1923,
and May 15, 1923.
112. Merops orientalis (Latham). ‘The Bee-Eater.
A very numerous summer visitor, first observed by Osmaston on March 1¢.
Briggs gives his earliest date as Match 8, and latest date October ll. Eggs ate
. laid at the end of April or beginning of May.
113. Merops superciliosus javanicus (Horsf). ‘The Blue-tailed Bee-eater.
A summer visitor, less numerous than the last species. It avoids the neighk-
bourhood of houses. Earliest date seen April 2, latest date September 6.
756 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
114. Merops persicus persicus (Pall). The Blue-cheeked Bee-eater.
Briggs says :—‘ Only once seen, and then in large numbers, of which a
considerable proportion were young birds : September 6, 1925.’
115. Ceryle rudis leucomelanura (Reich). The Pied Kingfisher.
‘2 Decembe.y 17, 1925. A resident species, common on the Kabul River and
its branches. Breeding commences in March, and Briggs found eggs op
April 1, 1924. :
116. Alcedo atthis. The Common Kingfisher.
g November 20, 1925. Fairly common along streams and canals, and
apparently resident.
117. Halcyon smyrnensis smyrnensis (1.). The White-breasted Kingfisher.
3S November 8, 1925. A resident, and common throughout the irrigated
country and near swamps. Lays at the end of April and the beginning of
May.
118. Upupa epops epops. The European Hoopoe.
© December 28, 1925, Peshawar. Briggs states :—‘ Resident and common, .
but the individuals which spend the winter in and about Peshawar do not
seem to be the same as those which breed, asin the autumn and spring there
is a period of a few days in which in some years there appear to be no Hoopoes
at all, while in other years they appear to be far more numerous than at normal
times. The explanation of this would seem to be that the Hoopoes which
winter in Peshawar go further north or east to breed, and that their place is
taken by birds of the same subspecies from the south. Sometimes those from
the south arrive before, and sometimes after the departure of the others.
Eggs are laid in the second half of April. Osmaston remarks that Hoopoes
were not common throughout the winter, but that they became more num-
erous ia February and March, and that breeding commences in late March.
These notes both suggest the marked spring passage which one would expect
in the Peshawar Valley.
119. Micropus affinis (Gray). The Common Indian Swiit.
Numerous and resident. It breeds chiefly in April, May and September, but
occasionally in almost every month.
120. Caprimulgus sp. The Night jar.
Briggs notes that some species is not uncommon in the summer, and that he
picked up a fledgling in May 1923.
‘121. Cuculus canorus (L). The Cuckoo.
One was seen by Briggs on April 23, 1921.
122. Eudynamis scolopaceus scolopaceus. (L). The Koel.
A summer visitor less common inthe Peshawar District than further down
country. Earliest date heard May 12. Not heard after the end of August.
123. Palgornis nipalensis eupatria (Hodg.). The'Large Indian Parrakeet.
Common and resident. Osmaston obtained fresh eggs on March 2, from a
hole in a mosque in Peshawar City.
124. Palzornis torquatus (Bodd.). The Rose-ringed Parrakeet.
Resident and numerous.
125. Asio flammeus (Pont.). The Short-eared Owl.
November 17, 1925, Peshawar. Not uncommon throughout the winter in
grasslands.
126. Strix aluco biddulphi. The Wood-Owl.
A single bird was seen and shot by Osmaston on November 18, 1925.
127. Bubo bubo bengalensis (Frankl.). The Indian Great Horned Owl.
December 2, 1925, resident. Briggs found two young, not quite able to fly,
near the Kalapani River on May 17, 1922. At the same place on another
oceasion he watched an adult bird of the species being mobbed by two Kashmir
Rollers. The Owl was sitting in full sunlight at about 10.0 a.m. on a boulder
on top of the cliff above the river, and the two Rollers kept swooping at its
head. For about a quarter of an hour it took no notice of these demonstrations
_ A NOTE ON THE BIRDS OF THE PESHAWAR DISTRICT 757
beyond bowing its head slightly as the Rollers came near, after which it flew
away. Osmaston notes that it is fairly common wherever there are woods of
any extent.
128. Carine brama The Spotted Owlet.
January 6, 1926. The Peshawar Resident. Common around Peshawar
wherever there are trees. Eggs found April ] to 14.
129. Pandion haliaetus (L). The Osprey.
Seen on the Kabul River by Osmaston.
130. Gyps fulvus (Habl.) The Griffon Vulture.
Said by Osmaston to be common near Peshawar, but the exact form requires
verification.
131. Pseudogyps bengalensis (Gmel.). The White-backed Vuiture.
Appears spasmodically at all seasons though it is not known to breed in
the Valley.
132. Neophron percnopterus (L). The Neophron Vulture.
Fairly common and resident.
133. Gypaetus barbatus. The Lammergeyer.
Rarely seen near Peshawar. Common inthe neighbouring hills and in the
Khaibar Pass.
134. Aquila heliaca (Savigny). The Imperial Eagle.
One was seen by Osmaston and Briggs on December 21, 1925, near the Pabbi
Swamp.
135. Aquila nepalensis (Hodgs.) -The Steppe Eagle.
Not uncommon near Peshawar Cantonments in winter. (Osmaston).
136. Aquilarapax vindhiana (Frankl.) The Indian Tawny Eagle.
A resident and fairly common.
137. Butastur teesa (Frankl.). The White-eyed Buzzard.
One was seen by Osmaston in March 1926.
138. Cuncuma leucorypha (Pallas). Pallas’ Fishing Eagle.
One was noted by Osmaston in January 1926 feeding on a dead animal by
the Kabul River.
139. Haliastur indus (Bodd). The Brahminy Kite.
Occasionally seen, most frequently after rain towards the end of the summer.
(Briggs).
140. Milvus migrans govinda (Sykes). The Pariah Kite.
Very common and resident. Eggs nearly all laid in the first week of March,
This Species has a curious habit of picking up sticks as if to commence build-
ing operations about the middle of September, and continues to do this till the
weather turns colder in October, when it seems to do no more till nest-building
begins in earnest towards the end of January or beginning of February.
141. Circus eruginosus (L.). The Marsh Harrier.
Common near extensive swamps in winter.
142. Circus macrurus(S.G. Gmel.). The Pale Harrier.
Seen occasionally in winter. (Osmaston).
143. Circus cyaneus (L). The Hen Harrier.
Seen occasionally in winter. (Osmaston).
144. Buteo ferox (S. G. Gmel.). The Long-legged Buzzard.
d December 21, 1925, Pabbi. 2 December 30, 1925, Peshawar. A common
winter visitor to the irrigated country. Inthe dry foothills it may be resident
in small numbers, as Cock records having taken the eggs of this species about
three miles from Nowshera in 1872. (Nests and Eggs, vol. iii, p. 126).
145. Accipiter nisus nisosimilis. The Sparrow Hawk.
¢ November 23, 1925, Peshawar. A fairly common winter visitor,
758 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXI11
146. Faico jugger (Gray). The Lugger Falcon.
Said by Briggs to be resident, but apparently commoner in March and
April than at other times of the year.
147. Falco chiquera chiquera (Daudin). The Red-headed Merlin.
& December 22,1925, Peshawar. A fairly common resident, generally seen
in pairs. Eggs taken at Risalpur, March 2, 1925.
148. Falco tinnunculus (l.). The Kestrel.
A bird in juvenile plumage was shot by Osmaston on December 21, 1925,
near Pabbi. The species is fairly common in the valley in winter, keeping
mainly to desert and swampy country.
149. Columba livia subsp ? The Blue Rock Pigeon.
Fairly numerous at Attock, on the cliffs by the Kalapani River, and on rocks
near Warsak.
150. Paiumbus palumbus casiotis (Bonap.). The Eastern Wood Pigeon.
A few were seen by Osmaston in a large orchard on February 23, 1926.
151. Streptopelia senegalensis cambaiensis (Gmel.). The Little Brown Dove.
A resident, but more numerous in summer than in winter. Breeds throughout
the hot weather.
152. Streptopelia decaocto decaocto (Frival). The Indian Ring-Dove.
A very common resident in the irrigated tract. Breeds from the beginning of
April to the end of the hot weather.
153. (Enopopelia tranquebarica tranquebarica (Herm.). The Red Turtle Dove.
A regular but not very numerous summer visitor, Earliest date seen
March 14.
154. Pterocies indicus (Gmel.) The Painted Sandgrouse.
This species is apparently resident in small numbers at Rustom, some twenty
miles from Mardan, in the Buner foothills. As it has also been procured in the
Orakzai, about half way batween Bannu and Peshawar, and Mr. Donald pro-
cured it west of Shinwari, it is probably more generally distributed in the foot-
hills than is generally known.?
155. Pterocles coronatus atratus (Hartert). The Coronetted Sandgrouse.
This species is said to occur near Jamrud.°*
156. Pteroclurus alchatus caudacutus (S.G. Gmel.). The Large Pintail Sand-
grouse.
A few pairs are said to breed in the vicinity of Mardan in June.*
157. Pteroclurus senegalensis (Licht.). The Common Sandgrouse.
Capt. Cock’s notes on the breeding of the Common Sandgrouse will be found
in Hume’s ests and Eggs, second edition, vol iii, p. 362.
A few are shot between Nowshera and Attock every year.
158. Coturnix coturnix coturnix (L). The Common Quail.
Passes through in large numbers in April and September. A few stay on in
the autumn until well into December, and may, perhaps, if they escape being
netted, spend the whole winter in the valley.
159. Coturnix coromandelica (Gmel.) The Rain-Quail.
Said by Osmaston to be conimon in the fields in November.
160. Ammeoperdix griseogularis griseogularis (Brandt). ‘The See-see.
Resident in the low hills surrounding the valley. It may sometimes be found
on the outcrops of limestone which rise in the middle of the valley.
161. Ortygornis pondicerianus (Gmel.). The Grey Partridge.
Fairly numerous in the low hills surrounding the valley.
———-—_____-—_--— — ——— — a —-—— — — $$ $e
* Stuart Baker, Game Birds of India, vol. ii, p. 256. ? loc. cit., p. 268.
® loc. cit., p. 283,
A: NOTE ON THE BIRDS OF THE PESHAWAR DISTRICT. 759
162. Rallus aquaticus korejewi (Sarudny). The Turkestan Water-Rail.
@ November 24, 1925, ¢ January 26, 1926, Peshawar. Common in almost
every swamp throughout the winter months. Very shy, and flushed with great
difficulty. The call, heard at dawn and after sunset, is peculiar, somewhat
resembling the noise made by a small pig in trouble. (Osmaston).
163. Fulica atra(L). The Coot.
Briggs says that itis very numerous in all suitable localities at all seasons,
but repeated search failed to discover a nest. Osmaston, on the other hand,
considers it not common,
164. Gallinula chloropus (L). The Water-hen.
Fairly common in winter according to Osmaston.
105. Otis tarda (L). The Great Bustard.
Of the six examples of the Great Bustard which have been obtained in India
four can be claimed by the Peshawar District.
The first specimen, a female, was obtained on December 23, 1873, at
Hastnagur near Mardan and just north of the Kabul River; a party were
found in some fields of mustard and giant millet, and they were so shy that,
though they remained in the neighbourhood for some weeks, no other specimen
could be obtained. (Hume, /6zs, 1871, p. 404.)
Two other females were shot in the same locality on January 6, or 8, 1911.
There were about twenty-five of them, and they were observed from roughly
December 20, 1910, to the beginning of February. (Fzeld, February 11, 1911.)
Finally Sir George Roos-Keppel recorded, in a letter dated December 1, 1917,
to the B.N.H.S. (Journal, vol. xxv, p. 745) that one had just been brought in to
him from the neighbourhood of Peshawar.
166. Chlamydotis undulata macqueeni (Gray and Hardw). ‘The Houbara.
An account of Houbara hawking on the Jamrud Plain will be found in
Game Birds of India, vol. ii, p. 195. (Stuart-Baker.) They are commonly
hawked abeut Mardan by officers of the Guides.
167. Lobivanellus indicus (Bodd.). The Red-wattled Lapwing.
A common summer visitor, first observed by Osmaston on February 24, 1926,
Briggs observed it occasionally in the winter.
168. Vanellus vanellus (L). The Peewit.
& December 19, 1925, Peshawar. Common in flocks in winter on open and
wet land. Osmaston first observed on November 12, but found it most
abundant in December and January.
169. Charadrius dubius (Scop.). The Little Ring-Plover.
Common on sandbanks of the Kabul River and on the margins of swampy
pools in winter.
170. Tringa hypoleuca (L). The Common Sandpiper.
A fairly common winter visitor to the banks of sluggish streams or near pools
and marshy ground.
171. Tringa glareola (L). The Wood Sandpiper.
_A winter visitor, but not very common.
172. Tringa ochropus (L). The Green Sandpiper.
o? November 11, 1925, 2 November 25, 1925, Peshawar.
173. Tringa totanus (L). The Common Redshank.
Numerous in winter.
174. Erolia minuta (Leister). The Little Stint.
A winter visitor, never very numerous.
175. Scolopax rusticola (L). The Woodcock.
Not common, but occasionally seen in woods in Jantiary.
176. Capella gallinago gallinago (L). The Common Snipe.
Q December 4, 1925. A common winter visitor.
177. Rostratula benghalensis benghalensis (L). The Painted Snipe.
‘My collection contains two specimens shot at Peshawar, and sent to mein
the flesh under the impression that they were Woodcock.’ (Whistler).
760 JOURNAL, BOMBAY NATURAL HIST, SOCIETY, Vol. XXXII
178. Chlidonias leacopareius (Temm.). The Whiskered Tern.
Numerous in winter, and possibly a resident.
179. Gelochetidon nilotica. (Gmel.). The Gull-billed Tern.
One which had been captured by villagers just outside Peshawar was seen by
Briggs on September 1, 1921.
180. Sterna seena (Sykes). The Indian River Tern.
Resident on the Kabul River.
181. Sterna melanogaster (Temm.). The Black-bellied Tern.
Resident on the Kabul River.
182. Rhynchops aibicollis (Swains). The Scissor-bill.
Found by Briggs in large numbers ona sandbank in the Indus opposite
Attock on April 24, 1923. From the behaviour of the birdsit appeared that
they had eggs, but a prolonged search failed to discover any.
183. Ardea cinerea (L). The Common Heron.
Fairly numerous on the Kakul River at all seasons.
184. Egretta alba (L). The Large White Egret.
Occasionally seen on the Kabul River.
185. Ardeola grayi (Sykes). The Indian Pond Heron.
2 December 8, 1925. Common and apparently resident.
186. Nycticorax nycticorax (L). The Night Hercn.
Commoner in summer than in winter, but noted at all times of the year.
187. Ixobrychus minutus (L). The Little Bittern.
One was seen by Briggs on July 26, 1920.
188. Botaurus stellaris (L). The Bittern.
Occurs in most of the swamps in tall bulrushes.
189. Cygnus cygaus (L). The Whooper Swan.
Of the five specimens of this species which have been obtained in India two
were shot by Mr. M. Donlea out of a herd of seven, on December 12, 1910, near
Dera Momin, on the Kabul River. (Stuart-Baker, Game Birds of India, vol. i,
p.1G.)
199. Cygnus bewicki (Yarrell). The Bewick’s Swan.
Of the two specimens obtained in India one was shot by Major P. C. Elliott-
Lockhart near Mardan, on December 30, 1910 (loc. cit. p. 21.)
191. Cygnus olor (Gmel.). The Mute Swan.
One was shot about the end of March, 1910, by Capt. H O’Brien at Now-
shera. Itis of interest to note that all the above-mentioned specimens of Swans
obtained in the Peshawar District were obtained in the same year. (loc. cit.
p. 26).
192. Amser anser (L). The Grey Lag Goose.
Fly over in very large skeins in autumn, In spring they appear to collect on
the Kabul River previous to going north. Briggs notesthat the two occasions
on which Geese flying south seemed to be most plentiful were after rain. He
says:—‘On August 25, 1922, there was thunder and rainin the hills in the
evening. From 9p.m.to10p.m. Geese in large numbers were passing over
in a southerly direction almost continuously. The following year rain fell on
August 22 and 23, from 9pm.on the latter night tillI went to sleep large
numbers of Geese were passing overhead. Each of these entries represent the
first record of Geese for the year. In none ofthe other four years for which
I have records are Geese noted earlier than September 4.’
193. Dendrocygna javanica (Horsf.). The Lesser Whistling Teal.
Briggs records that he saw asmall flock on October 12, 1922.
194. Casarca ferruginea (Pall.). The Ruddy Sheldrake or Brahminy Duck.
Fairly numerous in winter.
195. Anas platyrhyncha (L). The Mallard.
Numerous in winter on the Kabul River.
A NOTE ON THE BIRDS OF THE PESHAWAR DISTRICT 761
196. Mareca penelope (L). The Wigeon.
Not numerous, but a few are shot every winter.
197. Nettion crecca crecca (L). The Teal.
Common in the winter on the larger jhils and streams.
198. Marmaronetta angustirosiris (Ménétries). The Marbled Duck.
This species was obtained by Logan-Home near Nowshera. (loc. cit. p. 243.)
199. Nyroca rufa (L). The White-eye.
A winter visitor to the Peshawar Valley, but never very numerous.
200. Oxyura leucocephaia (Hartert). The White-headed Duck.
Four specimens of this species have been obtained in the Peshawar Valley.
(loc.-cit. pp. 304; 305. )
201. Merganser merganser orientalis (Oates). Tho Eastern Goosander.
.. [have received them or known for certain of their having been
obtained from the Peshawar Valley.* (Hume, quoted in loc. cit. p. 319.)
262. Podiceps ruficollis capensis (Salvad.). The Little Grebe.
Fairly common in the Peshawar Valley.
There can be little doubt that other ducks besides those included in this last
occur in the Peshawar Valley, but, beside those observed by Osinaston and
Briggs, only such species as are specifically mentioned in Stuart Baker’s Game
Birds as having been obtained in the district have been included.
19
MODERN MUSEUM METHODS
BY
5. H,. PRATER, .C.M.Z.S:
Curator, Bombay Natural History Society.
Par? II
(With two plates.)
(Continued trom page 544 of this Volume.)
CHARACTER AND ARRANGEMENT OF EXHIBITS
The more generally accepted method of exhibiting Natural History
specimens is to arrange them in cases in accordance with their
respective orders, families and genera, classification being the basic
principle which governed the arrangement. ‘The system adopted is
an heirloom of the days when no distinct line of demarcation was
recognized between the reference or study collections of a Museum
and its show collections. The attainment of a clearer concept of the
aims and purpose of a Museum as an institution for research and
popular education necessitated a change in the treatment of its
collections. Such exhibits from the collection as were suitable for
purposes of display were installed in the Puolic Galleries while the
main portion of the collection was closely reserved in rooms
accessible only on application by students and research workers.
At the same time the exhibits shown in the public galleries conti-
nued to be arranged and.divided in much the same manner as those
stored in the study collection, i.e. they were shown for the most
part as individual exhibits arranged according to their réspective
orders, familiesand genera. This classified series was usually supple-
mented with additional material illustrating various phases such as
structure, modification, habits, etc., which formed an index or intro-
duction to the study of the main exhibit. To sumup, the specimens
with their labels formed an illustrated text book or guide to the
study of Natural History and revealed the extent to which science
had progressed in the study of these subjects, and the principles on
which this study was based.
The exhibits in the public galleries of a Museum arranged as
above indicated, are intended for the layman. Their arrangement
is based on principles which are the outcome of knowledge gained
by men who have made a special study of the subject and are
endeavouring to hand down to the public at large the knowledge so
gained.
The exhibits as planned have a powerful teaching value; their
effectiveness for the purpose, however, depends largely on how the
MODERN MUSEUM METHODS 763
visitor uses the material the Museum has to offer. The teaching
of the Museum is silent teaching—a teaching which demands an
effort on the part of the visitor for whom it is intended. How far
is this effort on the part of the visitor forthcoming and how far is
the influence of the Museum’s teaching carried to the general body
of visitors ? We might liken the visitor to a Museum to a traveller
in a strange country. The traveller who has previously acquainted
himself with the traditions, manners and customs of the country he
is visiting has his mind prepared to benefit by the impressions he
gains by his visit—-he might be likened to that class of visitor who,
by his knowledge of the principles and motives which underlie the
arrangement of the collections in the galleries, is able to understand
and appreciate them and to benefit by the lesson they convey. For
this quite small class of visitor a collection so arranged is sufficient
to provide what is required. A second class of traveller who
arrives in a country without any knowledge concerning it, but yet
makes an effort during his stay to acquaint himself with the fresh
view-points it offers, may be likened to that still smaller class of
visitor who, realizing there is something to be gained by a visit to
the Museum, makes an effort to absorb somewhat of its teaching: to
this class the Museum has still something to offer. Lastly there is
that large class of people who travel mainly for pleasure or amuse-
ment. They may be likened to ninety per cent. of the visitors to a
Museum who visit it mainly from the same motive, quite often
because there is nothing better to do, or to keep out of the wet, or for
some such reason: people who not only lack knowledge but also
the inclination to acquire it, and who form undoubtedly the largest
proportion of visitors to a Museum. For them the teaching the
Museum has to offer loses most of its force unless that teaching
provides something which can arouse and hold their interest and
lead them to absorb, unconsciously perhaps, somewhat of the
message it intends to convey. A Visitor of this type passing
through galleries of named and classified animals comes away at
the most with the impression that there are a great number of
different animals in the world. The special, laboriously prepared
cases illustrating particular phases of animal life, labelled in terms
he cannot possibly hope to understand, hardly bother him at all, as
conscious of his shortcomings, he probably does not even look at
them. It follows therefore that the teaching value of the exhibits
in such a Museum is lost on the majority of its visitors.
One of the main objects of a Museum being the diffusion of
knowledge, the measure of its success will depend largely on the
ability of the Museum to drive its message home to the largest
number. We are teachers not as aright but only when that which
we teach is effective in influencing those whom we teach. If we
are invariably confronted by indifference, boredom or even hostility,
then we have failed in our objective. If we are eager to capture
the interest and enthusiasm of people, to get them to think and
work along with us, and are able only to capture indifference, it
is obviously necessary for us to address ourselves correctly to the
problem before us, not by taiking vaguely about goals and ideals
but by finding out quite specifically which methods are to be
764. JOURNAL, BOMBAY NATURAL HIST: SOCIETY, Vol. XXXII
employed if we are to ‘get across’ to capture attention, to win
regard and induce people to think and act along with us.
The classes of visitors for whom.the Museum must provide are ;
(1); The-Research: Worker.
(2) Thecsstudent:
(3) Those who visit the Museum without any definite object.
The Research Worker.—The needs of the research worker do not
enter into the present discussion which concerns. itself exclusively
with that part of the Museum collections which are accessible to
the general public. The research or study collections of a Museum
which are intended for professional or advanced students are ins-
talled in rooms to which the general body of visitors are not
admitted, and their installation and preservation present problems
distinct from those which arise in the treatment of the exhibits in
the public galleries.
Students.—As regards the student visitors, one might divide them
roughly into three classes: (a) University Students, (6) Primary and
High School Students, (¢) Members of the general public who
are interested in Natural History.
University Students—The needs of the University student. are
particular and distinct from those of the general body of visitors.
He is entering upon an academic study of Zoology which cannot be
followed on the broad lines in which Natural History as a whole is
treated in the galleries of the Museum. -A_ practical. course of
Zoology as prescribed for University students implies the detailed
study of particular types of animals which are representative of a
given group—this study of a particular animal in the totality of
its organization enables the student to compare its parts and organs
with those of kindred species and helps him to arrive at an under-
standing of the distinctive characters of the group as a whole
and the characters special to the example itself. He thereby
obtains an idea of the connection between comparative anatomy.
and classification, the significance of family, of generic and specific
characters.
- The. students’ “needs could therefore be met by a specially
selected collection of representative types, the details of whose
internal and external anatomy are illustrated by actual specimens,
models, diagrams and then labels. These types could be arranged
in their correct relationship so as to furnish the student with
material necessary for the study of a particular group, and at the
same time afford him a synopsis of the animal or the vegetable
kingdom as a whole. As a supplement to the main exhibit he
would require a series of exhibits whose object would be to explain
and illustrate by actual specimens, diagrams, etc., the various
technical terms used in Zoological description. A collection of
this character while it is invaluable to the student entering upon the
serious study of a complex subject, goes considerably beyond the
needs of the general body of visitors, for the majority of whom it
would hold little interest. Such a collection would therefore best
serve its purpose by being kept in a room or rooms which are out-
side the main stream of circulation of the general body of visitors.
For the greater convenience of students and for obtaining the
MODERN MUSEUM METHODS 765
seclusion and quiet that is necessary, admission to these rooms
should preferably be by ticket. For greater facility, students should
be able to obtain admission cards from the teaching staff of their
colleges, while members of the visiting public, desirous of doing
so should te able to obtain them from the check rooms at the
entrance-to the Museum building.
- The students’ room at the Edinburgh Natural History Museum
might be taken as an example of how a collection of this nature
should be arranged. A series of vertical cases lining the walls of
the room contains carefully selected representative types, classi-
fied according to: correct sequence, affording a conspectus of the
various phyla of the animal kingdom from Protozoato Man. The
exhibits consist of actua] specimens, where necessary of enlarged
models and of diagrams and labels which are arranged to illustrate
the bearing of comparative anatomy on classification. A series of
supplementary cases with carefully labelled specimens and diagrams
serve as an index to the technical terms used in the description and
labels of the main exhibit. The very careful and attractive manner
in which the material in this room has been prepared and presented
is not the least part of its value as an educative exhibit. The
methods adopted in the treatment of a collection of this nature, the
design and fixtures of the show cases, the colour used for back-
grounds, mounts and labels, the care bestowed in the preparation
of the specimens and the methods of labelling, all have a powerful
influence for repulsion or attraction... The response obtained from
the visitor in examining the exhibits in a museum gallery is similar
to the effect produced on the on-looker at a shop window: he may
be repelled by a crude display or attracted by an arrangement
carefully conceived with the purpose to please. In fact, in the
presentation of their treasures, Museums can learn much from
the persuasive methods of the window display artist.
Primary and High School Students—To meet the needs of
primary and high school students, many Museums in America,
both of Natural History and Art, have adopted the idea of the
Children’s Museum—a series of rooms in the building, or perhaps
an annexe to the building itself, which is devoted primarily for the
use of the children. Miss Anna Billings Gallup, Curator-in-Chief
of the Brooklyn Children’s Museum, the pioneer and perhaps the
most successfully conducted institution of its kind, has defined for
me the policy and ideals of her Museum. She said that its purpose
was to give the children experience with real things, objects of
beauty and scientific interest. An atmosphere of friendly hospita-
lity and of understanding of a child’s point of view was the basis
of all its activities. Through its door children are invited to enter
freely and enjoy the story-telling exhibits prepared especially for
them. Objects are put into their hands to be handled, admired and
compared, arranged, discussed and rediscussed. Games played
with objects on display enter into every-day’s programme and help
to bring into action new ideas in Museum experience which are
helpful both to the children and to those who guide the Museum’s
activities. In providing for children’s needs there is wisdom in
learning from children what these needs actually are. A trained
766 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
guide presides over these activities, whether they be in relation to
a group or an individual, and endeavours to bring his instruction
into concord with the school life, the home life and other
recreational interests of the child. The child learns to look
upon the Museum as a pleasure resort and centre of informa-
tion. He is even provided with a library organized especially
for him.
Briefly the aim and purpose of the Children’s Museum is to afford
intellectual entertainment and instruction of a better type for
children. It aims at arousing their observant powers and offering
them new fields, interests, study, new hobbies and teaching them
in a way children like to be taught, arousing their interest and
putting them into the way of finding things out for themselves
with just that little help and encouragement that is needed. The
feeling about the institution is that it is regarded by its visitors as
something of their very own—-something with which grown-ups have
very little to do. They are welcomed within its walls and are en-
couraged to use and handle and study its collections, read in its
library such books as especially appeal to them, play its games and
enter into its out-door activities. ‘That feeling of awe and restraint
which inspires the child during a visit to a Museum is not en-
countered here.
The staff indirect control of this section of the Museum
is chosen with care, the selection being governed as much
by. the capacity of the candidate to ‘handle’ children as by
his or her own knowledge and ability. ‘We find out so much
about the child here,’ said Miss Gallup to me, ‘ we find out the
observant children—tbe child with a penchant for independent
enquiry and the child who is careless at observation. Such
a one, given a series of questions, will come back with only
half of them answered. He or she is encouraged to go back
and study the objects until the whole is reveaied. Sometimes
we find that a child has no colour sense at all—cannot distinguish
between reds and blues. The child’s needs are two-fold, mental
and physical. The Children’s Museum tries to provide for both
of them.
During the past year, in the Prince of Wales’ Museum we have
had for the first time the fortunate experience of handling some
6,000 children and have come to realize something of the interest
and enthusiasm which can be engendered in them by an effort
directed to meet their speciai needs.
To come to the more practical side of the question and to illus-
trate how a Museum may serve its younger student visitors, also
those among the general public whose interest is not merely super-
ficial, I do not think that I can do better than by taking the
Children’s Museum at Brooklyn as an example of how the work
may be carried out. The Children’s Museum at Brooklyn has
served and assisted in the founding of Children’s Museums in many
foreign countries or in adding them as real departments to already
existing institutions. It has been a source of inspiration in the
work we are attempting to do here in Bombay, and I gladly pay
this tribute to Miss Gallup, its genial Curator-in-Chief for the
MODERN MUSEUM METHODS 767
help and encouragement I received at her hands during my visit to
her institution.
The Children’s Museum at Brooklyn is situated in what was once
a residential building in a small public park. Its collections are
arranged to meet the following purpose :—
(A) To provide a general outline of the animal kingdom by a
especially arranged collection of types illustrative of various
classes of the animal kingdom, in order to supply a foundation for
the study of the subject by making the visitor familiar with the
appearance and characteristics of the principal animal classes.
(B) To encourage an interest in Natural History and provide
material for its study by the exhibition of a series of exhibits
representative of the local fauna and flora.
(C) To extend its teaching to the study of the earth’s crust by
providing exhibits illustrative of the various minerals of which it is
composed.
(D) To co-ordinate its teaching more helpfully with instruction
the child is given in the schools by the inclusion of exhibits illus-
trative of History and Geography.
To sum up briefly the collections are divided into a Zoological
and Botanical Section, a Mineral Section and Historical and
Geographical Sections.
THE TyPE CoLLECTION
Zoology is set forth in a type room of ample dimensions located
on the first floor. The various ciasses of the animal kingdom are
presented in specimens, drawings, pictures and models, assembled
in cases that line the wall and occupy the central floor space.
Beginning at one side, the sequence of development is carried
around the room and down and up the central cases so that the
skeieton of man, the highest unit, faces the opening study of the
amoeba. Oiue large label for each sub-phylum or sub-kingdom is
fastened onthe glass front of the case and similar labels of uniform,
smaller size are provided for each class. Strips of moulding carried
down the back wal] of the case and across its base separate the phyla;
black cords cutline the minor divisions thus, by size of label and
width of outline the observer will know at a glance which groups
are co-ordinate. Another distinguishing feature is the color of
the bases or pedestals of the exhibit for all the lower forms, for
creatures whose habitat is the water or the sandy shore, they are
grey—for the mammals they are black or:very dark. The labels and
their frames are similarly distinguished. In the whole arrange-
ment the display is restricted only to such specimens as will show
in clear outline the important successive stages in the develop-
ment of the subject. Excessive detail is avoided as this is confusing
to the youthful mind. Pictures are used to show tiny forms many
times magnified, and to explain a label further or to represent
some form absent from the sequence, some form which must be
listed because children are interested in it, such as the elephant
among the Ungulates, an order sufficiently represented in the
exkibit by two specimens,
768 JOURNAL, BOMBAY NATURAL. HIST. SOCIETY, Vol. XXXII
The labels are mostly couched in popular language but as is
generally unavoidable in labelling a type collection scientific nomen-
clature will sometimes predominate and the direct appeal of the
label to the child will consequently be lessened. ‘This is where the
work of a docent or guide is most welcome. While the Zoology
collection as a whole is for pupils of the higher grades, the teacher
of any lower grade may feel sure that her pupils will receive much
benefit from a visit to the museum. For very young children the
description specializes in well-known animals and calls attention to
colour, beauty of form, and other striking characters. The guide
thus finds something to interest and instruct the child what-
ever its age or grade may be till the story of the Ascent of
Man is almost unconsciously learned. The lecturer makes a point
of taiking about just such exhibits as the child himself has recog-
nized, taiking about these first and then going onto other familiar
animals, he is able to give the child a desire for more ‘ stories’.
The child asks questions and learns more about the exhibits, and
gradually acquires knowledge of many animal forms, knows some-
thing of classification, and story of the Evolution of Man. ‘The
important point in the arrangement of a systematic collection on
the above lines, is that the collection should be kept as small as
possible. The inclusion of an excessive number of exhibits only
tends to confuse the issue; it should be limited to the inclusion of
such well-marked forms as will suffice to indicate the limit of varia-
tion in a particular class of animals and illustrate the inter-relation-
ship of the animals included in that class. One should always
bear in mind that one is providing for the amateur and not for the
professional.
LOCAE- BIRD ‘LiFe
A second series of exhibits illustrates the bird life of the local
park. The birds are attractively mounted, shelves are not used as
they tend to produce a monotonous line in this form of exhibition.
The birds are fixed to the walls of the case on brackets and are
arranged in cases according to the season during which these birds
can be seen in the park. Thus you have cases of permanent
residents, of winter residents, occasional visitants, summer birds,
Spring and autumn migrants. The series gives a graphic idea of
local bird life as it illustrates species which can be actually seen in
the child’s neighbourhood. ‘The importance and the value of an
exhibit illustrative of the local fauna, is too obvious to be stressed.
An interest in nature and later in natural history commences or
arises from an interest in the animal or plant life of ones immediate
neighbourhood, and a collection of this description will not only
supply the stimulus to this interest but will also help in its develop-
ment. In addition, such a collection will meet the requirements
of those whose main object in visiting the museum is to identify
some animal or bird they have seen in their vicinity.
THE INSECT ROOM
This room contains an attractively mounted and displayed series
of insects in wall cases which illustrate various features of insect
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THE CHILDREN’S MUSEUM
THE VIVARIUM 1S A GREaT CENTRE OF ATTRACTION TO YOUNG VISITORS.
A CORNER OF THE WELL-USED LIBRARY.
By courtesy. Children’s Museum, Brooklyn.
MODERN MUSEUM METHODS ; 769
life. The collection being intended for children, color and beauty
of shape have influenced the selection of specimens for illustrating
the various studies, but at the same time the exhibits are arranged
on scientific lines so that their full educational value is maintained.
The child can here become familiar with local moths, butterflies and
beetles. A silk worm and silk exhibit, a hive of living bees and
many species of insects that do harm to local trees and crops are
included in the series.
THE MINERAL ROoM
The Mineral Room of the Museum is one of its most popular
features. It is divided into two sections—a series of students’
trays containing labelled examples of minerals which the child can
examine and handle in a way that children like to do, and a corres-
ponding series of finer specimens which are included in the main
exhibition. The object here is to give the child a good working
knowledge of minerals. Mineralogy would usually be considered
a dry as dust subject for the child, but when attractively presented
young people quickly become quite enthusiastic over the science
and few types of lone material are more 1n demand at the Children’s
Museum than the trays of rocks and crystals, while the mineral and
volcano room swarm with little folk. Mineral study seems to have
become a tradition with the Museum, ard it has developed a
department which numbers now among its alumnia mining engi-
neer, a mine operator in Mexico, and the late Curator of Mineralogy
at the Public Museum at Milwaukee. All these found their first
interest in minerals in the rooms of the Children’s Museum at
Brooklyn. Although the function of a Children’s Museum involves
cultural rather than professional training, it augurs well for the
subject concerned when such results are obtained.
Tur BOTANICAL SECTION
This section includes cases which form an introduction to the
study of Botany, the various parts of flowers and leaves, etc., are
explained in models and pictures.
THE HIsTORICAL SECTION
American History occupies five sections, European sovereigns
and navigators, American, Indians, settlement and colonial period,
the Wars, New York State and City, outlining significant events
in national development, all these are set forth in charmingly
realistic miniature groups or paintings.
THE GEOGRAPHY ROOM
Outlines primitive life in all parts of the world, polar scenes,
jungle and desert, illustrated in minature groups.
THE LIBRARY
The Library in the Children’s Museum contains 7,500 carefully
graded and selected books. It is open during the same hours as
the Museum, i.e. 9 a.m. to 5.30 p.m. weekdays, 2.30 to 5.30 p.m.
Sundays. It is a pioneer library inits own field and has a share in
20
770 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol: XXX//
all the activities of the Museum. It is one of the best and most
complete nature libraries in the country, and its books cover a wide
range of additionai topics, such as travel, history, biography,
mechanics and inventions which appeal to the interests of the
child. The number of readers exceed 60,000 each year, and quite
frequently there are from 400 to 600 children visiting the library in
aday. At such time, of course, the children come in relays. The
library occupies two rooms, a home-like cheery room for children,
and an adjoining reference room for teachers and older students.
As far as possible individual attention is given to the readers, first
assisting them in the selection of books and leading them to a love
of good reading, and second teaching them to get reliable inform-
ation from books. Information is offered about all the exhibits
found in the Museum, and pictures, collections of poems and plays
and other data are provided in connection with school studies. The
boys and girls who have been working industriously for a certificate
of credit from the Museum in animal study, history, geography, or
Some such subject, come to the library in search of informa-
tion on a variety of subjects such as protozoa and amphibia, animals
found in the South Sea Islands, trees and vines of Brazil, and the
Berbers of the Sahara Desert. Again the librarians get inquiries
from children about their own city. On such topics as the different
types of bridges in New York and their construction, and how
the subways and tunnels are built. To showhow the library is
being used by the cbildren I might say that a girl composed a little
play entitled: ‘Christmas in Mother Goose Land,’ which her
classmates supplemented with poems found in the library. Needless .
to say, the library offers considerable attraction to Boy Scouts and
Girl Guides. They come in to identify flowers, birds’ nests, mush-
rooms and shells, and to inquire about aquaria, snakes, and a
number of things which interest them.
Among the library’s activities is the holding of exhibitions of
paintings and pictures appropriate to time and season, thus the
birthdays of Washington and Lincoln were the occasion for the dis-
play of pictures relative to the life of these men, and attendance at
these exhibitions is always very large. In this way books and
periodicals were used by hundreds of boys and girls, quite a large
number of whom wanted books and stories about these national
heroes. These patriotic pictures attract the children and even the
adults, while reproductions in colour of paintings representing scenes
in the history of the country give them an idea of the story of their
land and what it means. In connection with these pictures the
library posts up lists of plays and pageants, flag drills, etc., appro-
priate to the celebration of America in the making. There is also
a list of stories of countries, comprising titles of books about
eighteen European countries which had some share in the growth of
America. I have written at length on the activities of this library
because it is not usual to find work of this nature in institutions such
as these. The policy adopted by the Children’s Museum Library
shows just how very active and helpful a library can be and to how
many different fields of interest and endeavour it can lead ;its
users,
MODERN MUSEUM METHODS ri
EDUCATIONAL WORK
Musuem Courses
Opportunity is given to the children to work at the Museum in
eight different subjects represented by its collections. A student
may begin to work at his convenicnce and spend as much time as he
pleases on any ccurse. The whole thing is left to the will of the
child and no blame whatsoever attaches to him if he stops off and
fails to complete a course, ‘ We leave them to develop their ideas
by themselves,’ said Miss Gallup. ‘ Sometimes they tire of a thing,
and leave off; but mostly they come back again.’ The completion of
each course entitles each child to a certificate of credit which counts
towards a medal. Abronze medal is awarded for fifty credits and a
silver medal tor a hundred credits. Courses may be chosen in any
order. The courses and amount of credits against them areas follows:
A child desiring to takea course in Minerals applies fora leaflet
giving the requirements for a certificate in mineral study. He then
asks for a ‘model notebook’ on mineral study which gives him
directions how his own notebook is to be kept. In answering the
questions he records his description in pencil in his own notebook.
He gets from the library a book on minerals and from the docent a
streak plate for testing minerals, a box of minerals giving the scale of
hardness of various minerals, and the first student’s tray of minerals
containing twenty named important minerals in compartments. Each
compartment bears aletter and a number which corresponds to those
on the specimen and in addition the name of the mineral and its chemi-
cal composition is attached to the compartment, i.e., the first tray is
_A, the first compartment would be Al and below it is a iabel marked
‘‘Aliron pyrites, Fe S82.’’ Withthis equipment the child commences
the study, his note book must be the record of his own observation.
He must not crib from a text-book and the value of the work to him
will depend upon his own independent action. The book is only to
be used as a reference—if he cannot find the answer to any question
at once about any particular specimen he is recommended to leave
a blank, for in some later tray he will probably find a specimen
which will give him the information he is after. It necessarily
follows that the first specimen and the first tray take quite a long
time to do, but once he gets into the way of working, things become
easier for him. After completing A! he takes up A? and so on,
then he goes on to the B series, where he will find some new
minerals and also some duplicates, which will add to the information
of those he examined first. After studying the first three trays it is
suggested to him that he should mix up the minerals of the next
tray and see how many he can identify at sight. I have taken
mineral study as an example, but it is obvious that the methods used
here can be applied to every other phase of life. The whole objective
of the courses isto help the child, firstly, by giving him an incentive
to work in the shape of a prize, secondly, permitting him to work inde-
pendently, to allow him to handle exhibits and find out things for
himself, with the least possible assistance. Naturally, these methods
give the child an interest in the work by putting him ‘ on his mettle,’
as it were.
772 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Museum Gantes
Among the features of the museum are the games it organizes
for its visitors, games which are both interesting and educational. A
child wishing to play the ‘ bird game’ is given pictures of twenty-five
birds exhibited in the gallery and he is asked to go out and pick these
out. The next stage is to give to him twenty-five names and tell him
to find these birds in the colJection; in this way he becomes familiar
with quite a number of birds and is able to tell them at sight. There
are various games of this type covering various phases of animal
life adapted for children of various grades. A child who can iden-
tify twenty-five birds is allowed to take a stuffed specimen in a
small glass case home for a few weeks. He has learned everything
about this specimen and tells his people about it, and takes it to
school and tel!s his classmates about it. They in their turn get
interested and want to have a bird of their own and so the interest
in the museum spreads.
Stereoscopic Views
Every afternoon during the week a crowd of from twenty to fifty
children congregate in the History Room where a dozen stereos-
copes and a store of views are brought out under direction of the
museum attendants and placed at their disposal. Geographical
views, pictures of wild birds and flowers, prove particularly popular,
and more than six hundred children enjoy the use of these stereos-
copes each month.
(To be continued)
REVIEWS
1. AHANDBOOK OF THE BIRDS OF EASTERN CHINA,
(Part 1V)., By, J. D. D..La Touche (Taylor and Francis, London ;
75. 0d. net.)
We have already had the pleasure of noticing the publication of
the first three parts of Mr. La Touche’s Handbook of the Birds of
Eastern China, and now have to welcome the appearance of Part IV,
dated August 1927. This deals with the families Ploceide, Fring il-
lidze, Bombycillidee and Hirundinidz which are well represented in
India; indeed many forms described by Mr. La Touche are identical
with our Indian birds, whilst others are closely allied to them.
Two new races are described in this part, Amdberiza ctotdes vagans
(type locality Amur Bay) and &. c. fohkienensis (type locality
Foochow). It is interesting to note also that the author accepts
Dr. Sushkin’s opinion that Amberiza godlewskiz should be treated
not asa race of Emberiza cia but as a separate species with races of
its Own.
A new feature in the present part is that under’various species
the author includes a brief resume of the races, including extra
limited forms: this is undoubtedly useful. Itisa pity however
that theplumage descriptions are somewhat irregular in method
of treatment. Some are very diffuse and tend to become. almost a
catalogue of specimens inthe author’s collection; others err by
incompleteness, mentioning only asingle sex. Allthe same we are
able to congratulate the author on the completion of this further
instalment of his book, which will beof the greatest possible value
to all who are interested in Chinese ornithology and especially to
those who look forward to the resumption of active fieldwork in
China when her present troubles subside.
Indian ornithologists should note that in treating of the Red-
rumped swallows (Hzrundo daurica) Mr. La Touche does not agree
with certain details of the distribution east of India of the subspecies
striolata and nipalenszs given in the new edition of the Fauna;
the study of these swallows in India is very perplexing and a
correct estimate of their distribution remains to be written.
Mr. La Touche’s remarks should therefore be consulted by students
of the group.
This part contains a map to be substituted for that already issued
in Part III in which an error has been discovered. It also contains
four halttone plates to illustrate types of country, and it is interest-
ing how similar much of the country appears to that of India.
EW
2. THE LITTORAL FAUNA OF KRUSADAI ISLAND IN
THE GULF OF MANAAR WITH APPENDICES ON THE
VERTEBRATES AND PLANTS. By Various Authors.
This interesting publication marks the revival of a serial publi-
cation—Bulletin of the Madras Government Museum—which, as is
stated in the editor’s foreword, has been in abeyance since 1907,
774. JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
The series will hereafter be issued in two sections, the Natural
History and General Sections, and the number under review forms
No. I of vol. iof the Natural History Section of the Mew Series of
this publication. As the general title of the work indicates it deals
with the Littoral Fauna of Krusadai Islandin the Gulf of Manaar,
and is a systematic account of practically all groups of Invertebrates
excluding Protozoa, Porifera, flatworms and some of the smaller
groups. From amongst the Chordates only the Urochordates have
been dealt with, while a list of the Vertebrates generally is included
in an appendix. An account of the Flora of the area is published as
a second appendix.
The volume has been prepared primarily for the use of the
students who will be working inthe Biological Station which is
proposed to be established on Krusadai Island and should serve as
a very valuable handbook. The visits of the students, as is pointed
out in the introduction can as a rule only be of short duration, and
if full value is to be obtained from them some ready means, by
which at least the cornmoner organisms that will be met with can
be identified, must be provided. In Europe this can to a great
extent be done with the aid of the many excellent monographs that
have been published on various groups. Butin India, few such
monographs have yet been written. To preparea complete and
self-sufficient monograph ofthe fauna and flora even of so small an
area as that under consideration, witha full account of the classifi-
cation of every group, would swell the work to aninordinate size
with matter which would be largely a repetition of the work of
others already published elsewhere. And, however convenient it
might be, it would be undesirable; in as much as it would help to
confirm the impression which students so often get that all know-
ledge is to be found between the covers of their text-books. On
the other hand, to write an account which could only be foilowed
by specialists or with the aid of books that cannot, for some time at
least, be available on the island, would be to defeat the object of
the work. We have therefore followed as far as possible the
classification set forth in some standard text-book such as the
Oxford and Cambridge Natural Histories. The particular book
followed has been mentioned under each group and should be con-
sulted for details. As, however, only the broad outlines of
classification are dealt with in such books, reference is given to
other works that have been found helpful in connexion with more
detailed classification, and it has usually been necessary to insert a
brief account of the distinctive characters of such of the smaller
subdivisions as are represented in the collections. These have
been kept as short and clear as possible, only the most useful
characters being included. But keys have as a rule been avoided
on account of the sense of completeness and finality which they are
apt, often falsely, to convey. Descriptions of species have also
been kept as brief as possible, especially in the case of those already
described in the publications of the Indian Museum, in Alcock’s
‘Materials for a Carcinological Fauna of India’ (1895-1900), in
1 Journ., Asiatic Soc., Bengal, 1895-1900.
REVIEWS 775
Stanley Gardiner’s ‘Fauna and Geography of the Maldive and
Laccadive Archipelagoes’, inthe reports of the Percy Sladen Trust
Expedition to the Indian Ocean! and in Herdman’s ‘Report
on the Pearl Oyster Fisheries of the Gulf of Manaar’, works
bearing in so definite a manner on the fauna under considera-
tion that they must obviously be available for reference along with
the standard text-books already mentioned in the library of the
Krusadai biological station.
The different chapters dealing with the various groups have been
written by different authors, but the greater part of the volume is
the work of Dr. F. H. Gravely, Superintendent of the Madras
Museum, who has also acted as the editor for the remaining contri-
butions, and Dr. B. Sundara Raj, Director of Fisheries, Madras.
In each case the commoner forms met with have been dealt with in
sufficient detail for identification and very instructive line and dot
drawings of the species have been published. Fairly extensive
literature references have been included at the end of each chapter
and these with the work under review should enable students to
identify most of the commoner forms, which are found in the region.
A general description of the island and other collecting grounds
in this region together with maps of the different parts is also
included.
The work is by no means the last word on the subject and as the
editor remarks wilj need revision after the various groups have been
revised by specialists.
The editors and the authors, however, deserve the thanks of
students and incidentally of others interested in Marine Biology on
producing a really useful handbook on the littoral fauna of the
isJand and the adjacent parts, and it is sincerely hoped that it will
stimulate more detailed research on Indian marine fauna of the
region.
The printing and general get up of the work is really excellent
and the price (Rs. 8) for a work of its size with 26 plates, is by no
means exorbitant.
Bulk.
3. WITH A CAMERA IN TIGERLAND. By F. W. Champion,
M.A. (Oxon) F.z.s.. Imperial Forest Service of India. [Chatto &
Windus, London, 1927. Price 30s. 228 pages, 73 illustrations. ]
Books dealing with sport and the natural history of the fauna of
India have hitherto almost without exception, been conceived and
written with the object of describing the methods and adventures
which lead to the destruction of animals.
In his book Wild Animals in Central India, reviewed in this
Journal (volume xxix, page,1032) Mr. Dunbar Branderrightly remarks
that when not bent on slaughter, one sees and observes far more of
the animals of the forest. Mr. Champion advances us much further
than that, and in the book before us, which he correctly claims to be
‘the first book ever published which is illustrated throughout with
+ Trans. Linn. Soc, London_1907 onwards.
7744 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
photographs of wild animals just as they live their every-day lives in
the great Indian jungles’, does a real service to the wiid animals he
loves so well, for we venture to predict that an increasing number
of sportsmen will endeavour to live up to his teaching and example.
‘Tt is better’ we have recently read somewhere ‘to hold the
hand of Nature and to look into her eyes than to alienate and
slaughter her.’ That is the spirit in which this book has been
conceived and written.
The first three chapters give us charmingly written pictures of
‘A Day in the Jungle’, ‘ A Night inthe Jungle’, and an account of
what is truly described as ‘ An Extraordinary Experience’. Those of
his readers wno have had the fortune to wanderon elephant back
through the submontane forests of the Himalayas, must vividly
recail to mind days similar in a general way to that so faithfuily
presented to us. The coming upon the tiger and his kill has
happened to some of us, but to few of us has been vouchsafed the
sight of the wild elephant: and it is quite certain that no one of us
has photographed him in his natural surroundings as has our
persevering author.
The extraordinary experience related in Chapter iii must be quite
unique, and such photographs as plates xv, xvi, xvii, are unlikely to
be again obtained.
We take the volume in the order in which it is arranged, but keep
the elephant to the last. With the chapter on monkeys there are
seven excellent photographs depicting the common Brown or Ben-
gal Monkey and the Langoor, these being the two species of pri-
mates found in the forests of the United Provinces. With the
remarks of the author as to the ‘cruel iron bars’ of the cage of
capitivity most of us must be wholly in accord, and we hope that
the time will come when such things will be condemned by public
Opinion.
Mr. Champion expresses himself as absolutely convinced that a
langoor, if he calls four or five times, does so because he can actually
see a tiger or a leopard. Onthis point we would remark that we
have seen langoors give the alarm call at sight of a yellow dog-
and this not once only but on several occasions—while the smaller
monkey never took any notice of it. Otherwise also, in our experi-
ence, the langoor is less reliable than his smaller relative, who never
deceives the sportsman. ‘The aboriginal tribes of India recognize
that the langoor is not always to ke relied upon and wiil give the
alarm when rhesus would not. Both species provide excellent training
for the would-be photographer of animal life in the forests.
Of tiger in general the author writes with the knowledge born of
careful observance, and has much that is interesting to relate. He
rightly concludes that the sense of smell possessed by tigers is weak :
‘practically non-existent’ in his opinion; but we would not put it
so low as that, and would say that itis not more highly developed
than that of man.
As to the ‘ twelve-foot tiger’ the ‘ probability ’ that many of these
lengths were taken from skins after they had been removed from the
carcasses was amusingly confirmed to us only three years ago. A
sportsman who shot three tigers in one beat reported the fact to the
REVIEWS 777
civil officer of the District, stating in all seriousness in his letter
that the animals he had shot measured 13 ft. 6in., 12 ft. 9in., and
11 ft. 7 in., respectively. It was ascertained that these lengths
were obtained by taping the fresh skins, pulled out serpent-wise to
their fullest extent! That method of measurement is no doubt the
orgin of the ‘ twelve-foot’ tiger.
The sport of photographing tiger is fully described. The author,
having fixed for himseif the high standard of obtaining photographs
of animals under strictly natural conditions, and ‘not where he is
the pursuer or pursued’, rules out the employment of beating for
the production of photographs of tigers ; and explaining that he has
never yet had a tiger come near a machan in daylight, in a light
good enough for a picture to be produced, he proceeds to describe
two other methods.
One is ‘ ghooming ’ (not ‘ ghumming ’) an anglicized colloquialism
of a Hindustani word, or wandering about in hopes of a‘ subject’; and |
the other the stalking of a tiger on his kill. A well-trained elephant,
with an equally well-trained mahawat (we like that spelling of the
word) is essential to either method. But the experience of the
author, after years of continuous effort, is that but few pictures are
obtained in proportion to the amount of energy expended; and
he comes to the conclusion, which is borne out by the vivid
‘action’ of his flashlight photographs, that this is the best method
of all.
Success, however, the author remarks, is only to be obtained by
dint of perseverance and determination to succeed in spite of repeat-
ed failures andthe risk of contracting severe attacks of malarial
fever. In some other paris of India the fever risks would not be so
great. Justly, however, can the author claim that automatic flash-
light photography is ‘ not too easy to be sporting ’.
In the enlarged copy of the head of the tiger of plate xiv, xxi
which forms the frontispiece of the book, we see the size to which
the pupil expands in the dark and can realize how it is that, however
fine the trip-wire may be, it will most likely be detected, as is shown
at plate xxx. Also we realize the delight of the author when he
developed this plate !
We are given an interesting chapter on tigers and their prey, the
author coming to sound and right conclusions on this as on other
matters he discusses. Tigers do not always seize their prey in the
same manner. ‘Circumstances alter cases’ and, as indicated by a
photograph on p. 587 of Vol. xxxii of this Journal and witnessed by
the present writer under electric light, a tiger, when the prey is easily
within his power, does not necessarily break the animal’s neck, but
seizes it in a vice-like grip until life is extinct.
In these days of modern photography the aim of all enthusiasts
is to obtain a photograph of tiger in the act of killing his prey. Mr.
Champion has not yet succeeded in obtaining such a picture and it
can be fairly safely asserted that only by cinema photography, car-
ried out by daylight under previously prepared conditions, will
such a photograph be obtained. Nevertheless we hope that our
persevering author’s efforts in this direction will be crowned with
suCCeSS.
ey
778 JOURNAL, BOMBAY NATURAL AIST, SOCIETY, Vol. XXXII
Of leopards and the smaller cats we are given some excellent
pictures and interesting reading. It is remarked that in the detec-
tion of the trip-wire, however fine it may be, the leopard is even
superior to the tiger. That is an instance of knowledge coming tous
through the photographing of animals in their natural surroundings.
The author might, with accuracy, have asserted more positively
that the leopard cat is less common than the jungle cat (Felis chaus).
The photograph at plate xxxiv is that of the Small Indian Civet
(Viverricula malaccensts) and not of the Large Indian Civet (Viverra
zibetha) as it is without the distinctive Vzverva crest along the
spine.
Under the heading ‘ Hyzenas and Others’ we see pictures of
hyzenas and jackals only. The absence of a photograph of the
animal popularly known as the wild dog is to be regretted ; but, as
the author says, it would take many, many years successfully to por-
tray photographically all the interesting creatures of the jungles.
Mr. Champion is correct in stating that there are large numbers
of hyzenas in the forests. They are very common animals. We
are at one with the author in doubting that a leopard will abandon
his kill to a hyzena, also we agree that the hyzena of the forests is a
particularly silent animal. The killing of tethered goats by
hyzenas is within our experience, also we have personal knowledge
of a hyzena having killed and eaten a woman and a boy and attacked
another woman.
Of the rodents, photographs of the Indian Palm Squirrel and of
the porcupine are given. In view of the certain knowledge we
have that leopards frequently kill porcupines for food it is not
probable, as the author remarks, that they often inhabit the same
dwelling.
Of the deer tribe we have good photographs of the stag and hind
chital, but none of the noble sambur stag, or of the ubiquitous
barking deer. ‘Theabsence of pictures of these animals, so common
in the forests known to the author, emphasizes the difficulties in the
way of successful animal photography under natural conditions.
Of-‘ Antelopes and Others’ there are pictures of blue bull, and a
silhouette of a distant gural; also a photograph of two wild boars
in a characteristic attitude. Mr. Champion is correct in stating that
the serow is to be found at elevations considerably below 6,UCO feet.
The wild elephant has, perforce, to be photographed by day, and
in doing this Mr. Champion has been eminently successful. He
gives us much interesting information as to the manners and habits
of both tame and wildelephants. Both the chapters are exceedingly
interesting, and the photographs among the most striking in the
book.
In justice to the author we must touch but lightly on all he tells
us and leave the reader to his certain enjoyment of it. He will
learn much of what the author terms the fascinating pursuit of
photographing wild elephants, will read of some highly exciting
incidents, and generally learn much that he did not know before.
The final chapter of the book is on photography. Willing and
helpful-expert advice on cameras, lenses, plates, etc., is given; also
especially valuable practical instruction as to the flashlight
REVIEWS 779
photography by means of which the author has been able to secure,
and produce for our instruction and enjoyment, so many excellent
pictures.
However, he makes one statement which calls for comment.
his. issn regard to Marius Maxwell’s ‘ Big Game Hunting with a
Camera in Equatorial Africa’. According to Mr. Champion ‘ nearly
all his (Maxwell’s) pictures were produced with an ordinary 8 in.
lens, and strange to say, without a reflex camera’’. We cannot under-
stand how Mr. Champion can have derived the imipression that
Maxwell did not employ a reflex camera in face of what that author
Says at the very commencement of the book, in his lucid and
instructive chapter on ‘ A Camera Huntsman’s Equipment’ (page 3,
para 3;. Plate I of this chapter, moreover, depicts the camera
chiefly used by Maxwell, which is manifestly a short focus reflex.
We do not know what Mr. Champion’s sources of information
may be for stating that Maxwell’s method of photographing big
game was ‘to stalk up to his subject and then to expose piates at
close range, often by holding the camera over his head and centering
the animal by means of a direct vision view finder’’. From a perusal
of Maxwell’s book one certainly does not gain the impression that
this method was often adopted. It may perhaps have been only in
a very exceptional instance that it had to be resorted to.
Or the game animals of India the author has photographed seven
in their natural surroundings and at close range. In the mountains,
plains, and forests of India and Burma there are more than thirty-
seven! So there is plenty of room for those sportsmen who will
abandon the rifle in favour of the camera and preserve pictures
instead of horns, and hides, and hoofs, of the game animals of this
country. Through animal photography such as isput before us by
Mr. Champion, the sportsman will obtain records more permanent
than perishable trophies ; the naturalist, the sculptor, and the painter
will all have pleasure in and profit by his labours.
The duty of a reviewer is to criticize, withholding nothing
adverse if such be found; but we close this book with no note of
dissent, and feel sure that a wide circle of readers will enjoy it
with very deep interest. More especially will it be appreciated by
those who themselves know and love the vast jungles of India and
have at heart the preservation of their wild inhabitants from useless
destruction.
RB
4. DWELLERS IN THE JUNGLE. By Lieut.-Col. Gordon
Casserly. [Ward Lock & Co., Ltd., London and Melbourne, 1925.
255 pages, 16 full-page illustrations by Warwick Reynolds. 5s. net.]
5. IN THE GREEN JUNGLE. By Lieut.-Col. Gordon
Casserly. [Ward Lock & Co., Ltd., London and Melbourne. 255
pages, 16 full-page illustrations by Warwick Reynolds. 5s. net.]-
The stories of animal and bird life in ‘ Dwellers in the Jungle’,
given to us by Lieut.-Col. Gordon Casserly, are exceedingly
well told and hold the attention of the reader from cover to cover,
The word painting of the scenery of the great forests of India is
780 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
good, the style easy and attractive, and the natural history in the
main quite accurate.
We are told of the baby wild elephant and how his mother saved
him from the murderous onslaught ofatiger : andof a tame elephant
(this being a story built upon an actual occurrence) roaming wild for
the best part of a year and returning to her bondage. The use of
the word ‘ galloping ’ to express the utmost pace of an elephant is
misleading to those readers who have not seen this animal moving
out of a walk. The two stories of wild and tame langoors are
natural to life and realistically told, as also that of the jungle cock
and of the weaver bird, and in ‘ The Warden of the Ford’ the way
of the man-eating crocodile is faithfully presented to the reader.
In ‘The Hated Horde’ the story of the wild dog of the Indian
jungles is well depicted. The author describes these animals as
hunting in full cry, and mostly, though not always, at night. We
believe it is a fact that Cxon dukhunensis hunts entirely by day, and on
the several occasions on which we have actually seen them in pursuit
of their quarry they have been running quite mute. No doubt there
would be some eager whimpering when the climax of the hunt is
reached.
Of the many other natural history incidents, and the like, related
to us, we doubt the king cobra attaining a length of 18 feet; and
have not before known that the python, or any other snake, first
secures its prey by flinging a coil after the manner of a lasso. The
hyzena does not wear a perpetual snarl, but, like all other animals,
bares his teeth only when he has reason to do so. Theterm ‘ buck’
we would reserve for the male of the goat species and think it a
misnomer for the deer tribe.
With these few criticisms in mind we commend the book whole-
heartedly to all lovers of nature study, and the glamour of the great
forests of India and their denizens will be with them as they read.
‘In the Green Jungle’ is not a wholly accurate title as the open-
ing story is that of a camel of the Great Sahara Desert, and a very
good one it is. The life of the ubiquitous jackal is excellently
well related. The ‘ Law of the Jungle’ is the story of a man-eating
tiger. His fitting end is doubltess based upon an actual incident of
the kind. Throughout the forests of India are many ancient ruins
and what the wild life may be amid some of these is vividly
described.
Of the gallant wild boar, the ruthless python, the swift hunting-
leopard, we are given absorbing tales. The story of the white ant
is good, and in ‘ The Mockers’ the langoor again provides interést-
ing material.
There is but little room for criticism. We would not call the
‘ Did-he-do-it ?’ which is a lapwing, a long-legged fowl.
The book is of merit equal to that of the former volume, and we
close it with similar commendation to the readers old and young
alike.
R.B.
EDITORIAL
We have recently welcomed—and very warmly—another expedition sent out
to India to obtain further material for the Natural History Museum in New
York, and by careful o bservation by trained artists in the actual haunts of our
big game to learn how best to exhibit the denizens of our jungles to the public
of America.
The expedition was organised some years ago by Mr. Arthur S. Vernay, a
business man of London and New York and incidentally an enthusiastic big
game hunter and naturalist, in collaboration with Col. J. C. Faunthorpe, late
Commissioner of the Lucknow Division, United Provinces. Its object was to
secure a representative collection of the game animals of the Indian Empire
for the American Museum of Natural History, New York. Through the
active support of the Government of India the expedition received exceptional
facilities for achieving its purpose, with the result that a series of very beautiful
groups illustrating the game animals of this country are now being prepared
for the Hall of Asiatic Maminals at the New York Museum. To complete the
work and in order that the groups of animals exhibited might be shown in their
natural setting, Mr. Vernay arranged in December last year for two experts
from the staff of the American Museum, Mr. Albert Butler and Mr. C. C. Rosenk-
ranz, anoted landscape painter, to visit India with a view to studying the actual
areas where the animals were obtained, collecting forms of plant life typical
of these areas and painting pictures and taking sketches of the country in which
the animals lived. Mr. Vernay considered that the Museums in India should
also benefit by this work and he accordingly very generously invited Mr. S. H.
Prater, the Society’s Curator, to accompany this expedition and also provided
the funds for Mr. K. B. Savardekar our Museum artist, who has painted some
excellent backgrounds for groups in the Museum at Bombay, te be a member of
the party. Mr. Vernay’s generosity has not only enabled the staff to acquire
first hand knowledge of up-to-date methods in the field but has also enabled
our Museum to secure a very beautiful series of paintings and photographs
taken in the forests of India and Burma which will serve as studies for realistic
settings for groups which the Curator is now preparing.
In addition to the above Mr. Vernay has presented the Society with a copy of
a remarkable film recording the work the Vernay-Faunthorpe expedition in
India. Taken at a cost of nearly £6,000, the film records scenes of animal life
taken from Nature in the forests of Nepal, Burma, Central and Southern India.
Rhinos, Wild Elephants, Tigers, various forms of Deer and even some of the
smaller and less known animals of the Indian Jungle play the leading role in
this film which is the finest record of aninial life ever made in this country.
It is remarkable that a man who has had but a remote connection with this
city has seen fit to do so much for one of its public institutions. Museums have
been ever dependent for their origin, their growth and development, on private
enterprise and on the support of citizens, who take a pride in their city and its
institutions. Ifa stranger can do so much, surely there must be many in this
city who would cone forward in the same way to help their own Museum if
they realised how rapidly it was taking its place as one of the finest institutions
of its kind in the East, and how increasingly it was fulfilling its mission as a
centre of culture, of popular education and entertainment,
Mr. Prater was away about two months and was accompanied by our artist
whom, thanks also to the generosity of Mr. F. V. Evans—one of our Vice-
Patrons—we have been ableto send on the journey. We say thanks to Mr.
Evans as he is providing the salary for a second artist to prepare models of the
fish of our coasts and rivers and at the moment this permitted of our sending
Mr. Savardekar with Mr. Prater, When in Mysore the party was looked
after by one of our members— Mr. R. C. Morris of Mysore to whoin our thanks
are due.
Weare very glad that we were able at the present time to give Mr. Prater and
our artist this opportunity, as our hopes of being able shortly to erect a modern
782 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
Natural History Museum are steadily increasing. In our last editorial we had
the pleasure of thanking Sir David Ezra for his donation of Rs, 5,000. Now
we have asimilar pleasure in recording a donation of £500 from Mr. Vernay
towards the cost of equipping the New Museum. Mr. Vernay thus not only
gives our Curator the opportunity of learning how to do it, but provides us with
the means to enable the Curator to follow the teaching of Mr. Squeers and
having learnt the meaning to carry out the job.
Mr. F. V. Evans who gave us the beautiful model of a fly, enlarged twenty-
eight times has followed up his gift by presenting us with similar models of the
Anopheles and Stegomia mosquitoes, both the work, as with the fly, of the two
sisters the Misses Edwardes. As in the former case, so now, we have to thank
the P. & O. Steam Navigation Company for bringing these models out free of
freight charges. Possibly because the P. & O. did this in the case of the fly,
Messrs. Mackinnon Mackenzie & Co., not to be outdone, took home for ws for
the London Zoo a live elephant free of freight, only charging for the carriage
of Homo sapiens who went in attendance. We wonder if any one will present
the Zoo with a pair of Rhinos so that we can say to them ‘ You brought out
two mosquitoes free, surely it is not too much to ask you should take back free
also two Rhinoceroses !’
Our NEw CONSTITUTION
A Special General Meeting of Members of the Society was held on January
31, 1928, when the following resolutions were passed :—
1. That it is expedient that the Society be registered as a Society under
Act XXI of 1860.
2. That the draft Memorandum of Association and Rules now laid before
the Meeting and which were published in the Society’s Journal of October
20, 1927, be hereby approved as the Memorandum of Association and Rules
of the Society.
3. That the Members of the Executive Committee and Advisory one
mittee therein named be approved.
4..~That Sir Reginald ‘Spence, .Mr.H2 A. OW. Brent and Mire) 375
Greaves or any two of them, be hereby empowered to take all necessary steps
to effect the registration of the Society and to transfer to it all the property
and assets including securities and cash belonging to the unregistered Society
and for that purpose to execute on behalf of all the members of the unregistered
Society all deeds, transfers and other documents which may be SSSA or
expedient.
The Rules and Regulations as published on pages 385-393 of vol. xxxii were
passed with the following slight amendments. Rule 7 was altered to read :
. ‘The capital obtained from life-membership contributions and compounded
‘subscriptions shall not be used as revenue Eut shall be invested in Govern-
“ment securities and the interest thereon only utilized as revenue. PROVIDED
‘ALwAys. that if and whenever the market value of the investments shall
‘equal or exceed the amount of the contributions and compounded subscrip-
‘tions paid by the existing life-members and the members who have coin-
‘pounded their subscriptions then and in such case it shall not be cbligatory
‘to invest any further sums received by way of contributions from life-members
‘ or compounded subscriptions but the same may be used as revenue.’
At the present time our investments in Government paper exceed the amount
paid by life-members now on the roll and we wish to be at liberty to use any
surplus as members may, through the Committee, consider to be in the best
interests of the Society. For instance it might be in the best interests of the
Society to spend some of the surplus life-membership contributions on the
equipment of the new Museum.
The other alteration was to Rule 54 and was to the effect that
‘No payment shall be made by the Honorary Treasurer without the
‘ authorization of the Honorary Secretary, or in his absence, a mein ber of the
‘ Committee.’
THe Socrety’s ASSETS
The registration of the Society necessitated the statement of the property
handed over by the old unregistered Society to the new and registered
EDITORIAL 783
Society and members may be interested in the valuations which were given,
namely :—
Securities as on December 31, 1927 (Face value) Rs. 90,000.
Fixed Deposits, Bank Balances and Cash toa ay 19 5156-12-85;
Research Collections ... eee oat pe 09,0007)
Library Books sae “es me », 20,000 |
Publications ses oe ae 5, 17,500 }}Nominal value
Journals of the Society nie me ,, 15,000 | for Insurance.
Furniture awe 3 ado: |
THE SOCIETY’S PROPOSED ‘ BOOK OF BIRDS’
In 1926 we started work on the painting of the plates for the Society’s popular
illustrated book on the Common Birds of the Plains of India. We are glad
to record that our artist has now finished the illustrations in colour of 200
species which completes our programme. Capt. Pates has provided us with
several very interesting photographs of Bird Life and a chapter on Bird Photo-
graphy. The work is therefore nearly ready for the Press and would be so now
but that our staff, having so little to do!, have only completed the letterpress
for notes and descriptions of about forty species. However, we hope to be able
to make a definite announcement as regards price in our next issue,
THE GAME BOOKS
Writing of price reminds us that the reduction in the price to members of
vols. i and ii of the Game Girds has caused a revival in demand and has evidently
impressed on several the advantages of membership, but we must ask
members, who are keen on the publication of another volume, to continue
their efforts to induce other members, who have not done so, to buy and those
who are not members to become so in order that they should be able to buy
cheaply also.
GAME PRESERVATION
In No. 2 of this volume we gave our views on Game Preservation in India
and in the editorial which appeared in No. 3 we referred toacertain amount
of criticism which we had received. We give below in full acriticism by
Lt.-Col. C. H. Stockley, a Member of our Advisory Committee who would
like to see the sale of game or of the skins or heads of gaine prohibited. Col.
Stockley writes under date January 19, 1928 :—
‘My idea on Game Preservation is that concentration on the sale question
is essential, as other questions involve dispensal of force and are open to
attack: the sale question is unassailable on the crop destruction plea, as the
cultivator still retains the right to kill marauding beasts. It is so-called ‘‘waste’’
lands which attract the poacher. It is there he makes his biggest profits and
actually unites in small bands witha regular supply system. Witness the
results in the Northern Circars and any accessible ‘‘ waste’’ lands. Nowhere
in India does the so-called ‘‘ Zemindari ’’ jungle hold any game but a few scat-
tered remnants. I shot all over the country near Nowgong twenty-five years
ago and got grand sport I have just been out for ten days and the country is
almost bare of mammalian life. I confess to being almost fanatical on the
‘‘sale’’ question, as I have lately seen some of the results and heard more.
The elephant question is one that ruins much of our efforts tosave game. I had
the figures of the Kheddah sales and outgoings, but have lost them. The
damage done to crops alone is immense compared with the small results of the
sales, particularly in S. India; while the Forest Department in Kanara in 1922
temporarily offered a reward for the destruction of elephants, but had to dis-
continue it for want of funds. The Canara elephants were constantly being
reinforced by protected elephants from Mysore and possibly even further south
I have seen a herd of over eighty elephants within a couple of miles of a
railway station in the Coimbatore District, while the rice fields of a Burmese
village after the visit of quite a small herd in the rains, have been a really dis
tressing sight, considering how poor the jungle folk usually are. ‘The Kashmir
game is doing well, except Markhor. These are rapidly decreasing everywhere
they exist, and are vanishing from the Pir Panjal and Kaj-i-Nag. Astor is bad,
Chitral worse, and the N. W. Frontier Province will soon lose the last of the
straight-horned race.’
784 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXX
GAME PRESERVATION AND OTHER KINDRED SOCIETIES
We referred to the ‘Society for the Preservation of the Fauna of the
Empire’ in the last editorial and stated we intended affiliating our Society
to the one at home. We gave a resumé of the objects of the Society in
No 1, volo xxxil:
We now have pleasure in referring to the work done by the Nilgiri Game
Association during the year ending June 30, 1927. We notice that as from
October 1, 1926, Government transferred the management of the plateau fishery
to the Association subject to the following conditions :—
(1) The Association shall regulate fishing in the plateau waters in accor-
dance with the rules and regulations which the Government may prescribe from
time to time.
(2) The Association will be at liberty to appropriate all receipts from
licenses which they may issue for fishing in these waters, but shall be responsi-
ble for the efficient management of the fisheries and the working of these fish-
erties shall not be a charge on the provincial revenues except to the extent
indicated in condition (4) below.
(3) In the event of mismanagement or of infringement of any of the condi-
tions mentioned above, the Governinent shall be at liberty to resume the control]
of the fisheries without payment of any compensation ; the decision of Govern-
ment as to whether there has been any mismanagement or infringement shall be
finai.
(4) The Government will provide for the supervision of fisheries as
well as the game protection work of the Association free of charge thereby
also relieving the association of the payment of Rs. 50 per mensem which
it has been making to the Inspector of Fisheries the Nilgiris for the latter
work.
The question of continuing the payment of rewards for the destruction
of wild dogs was discussed and it was decided that the payment of the
rewards should continue, although in July 1926 the Government declined
to grant to the Association fines levied under Rule 6 of the Nilgiri Game
Rules :=—
The following were the principal alterations made in the shooting rules :—
(1) Small game shooting shall open with pigeon shooting on September 1,
for the sake of uniformity.
(2) That the killing of all monkeys be prohibited.
(3) That the shooting of all Ibex except bucks possessing a well-defined
saddle mark be prohibited.
(4) That only one sambhur stag be allowed on season license. On the
monthly license one shall still be allowed subject to the condition that it shall
not be shot on the plateau.
One of the principal changes in the fishing rules was :—
Permitting the killing of trout of 8 inches or less, such trout not to count
against the limit allowed under the license.
We quote the following notes by members of the Association. Major
E. G. Phythian Adams (a member of our Society) wrote :—
‘ The present condition of the big game in the area continues to be satisfac-
tory, but more effective steps require to be taken to stop poaching which is on the
increase. The fact that the Nilgiri Game Association has set aside a sum for
reporting cases of poaching should be made known to all watchers and Forest
Guards and also to the Police whose assistance is needed in checking the sale of
game during the close season.
Wild dogs are still too numerous to allow the reward to be discontinued.
There was a large pack at Mudumalai in May, while others were working Masi-
nigudy and Anaikatty as usual. A small pack was reported several times on
the plateau towards Mukerty.
The small Game season appears to have been an average one.
The Peermund and Bangitappal streams need restocking ; there appear to be
no fish at all in the latter. There are several other good streams in the Kundahs
which might be stocked with advantage against the time when the Pykara
Hydro-Electric scheme will ruin the trout fishing in the Mukerty River. I do
not think that any appreciable improvement in the fishing can be expected till
steps are taken to reduce the present Jarge number of otters which infest all
the streams. Their food certainly consists largely of crabs, but the latter
BIRDS OF KASHMIR—-AN APPEAL
See
For some years past I have been collecting records and notes
with a view to writing a detailed avifauna of Kashmir. This work
I now hope to take in hand at the end of the summer on the con-
clusion of a final visit to Kashmir. May I ask any of our members
who have any notes and observations on Kashmir birds if they would
be kind enough to contribute towards the completeness of such a
book, either by sending such to the Journal for publication these
under their own name in the very immediate future or by sending
them to me? I should of course be careful to acknowledge my
indebtedness to any one willing to help in this manner.
By Kashmir I mean any of the territories under His Highness
the Maharajah of Kashmir and Jammu following the ordinary
political boundary and including therefore not only Kashmir proper
and Jammu, but also Ladakh, Baltistan, Astor and Gilgit. Notes
referring to any area will be most acceptable, as will be readily
understood when [ explain that some of the lowest and ornithologi-
cally least interesting areas are less represented in my notes than
the valleys most difficult of access. For instance, there is less on
record about the plains of Jammu and the Jhelum valley cart road
than about Gilgit or the Nubra Valley.
Detailed lists and observations or isolated records of a single
species will be equally acceptable, and in particular 1 should be glad
of notes on the various game birds, especially those of interest to
the sportsman as well as to the ornithologist.
If any one can help me with information I shall be very grateful.
CALDBEC HOUSE,
BATTLE,
SUSSEX, HUGH WHISTLER.
April 1, 1928.
EDITORIAL 785
form part of the fishes’ food supply and at the same time keep the number of
fish down by destroying ova also those fish which the otters do take are the
largest.’
Mr. J.B. Vernede, wrote :—-
‘All game is decreasing in the Kotagiri District and on the North,
east slopes. Sambhur have decreased very markedly and as far as 1 am
aware not a single shootable stag was seen during the season. Small
game has also decreased largely, jungle fowl and hares in particular being
very scarce.
I cannot ascribe any particular reasons for this decrease though poaching
which is rife, may be a contributory cause, also some increase in vermin.
Writing of the year under review in general the Committee record—
(a) Shooting—
The year has been on the wholean average one and there is little of in-
terest to record. In accordance with a resolution of the Committee at its
meeting held on December 13, 1926 the services of allGame watchers were
dispensed with from January 1927 as it was considered that they were of little
if any use in checking poaching or for any other purpose.
(b) Fishing —
There appears to be a distinct falling off in the quality of the fishing both
as regards numbers and size of fish though perhaps not in the average size of
the fish in the streams.’
The following amounts were spent in destruction of Vermin—,
RS:
Wild Dogs aes ... 360
21 Otters eee a tO
Other Vermin ... w.. 7/4
CENTRAL PROVINCES ANGLING ASSOCIATION
We welcome the publication of the Ist Annual Report of the Central Provin-
ces Angling Association from which we take the following extracts as regards
the birth and activities of the Association.—
Early in 1927 a small number of enthusiastic anglers met togetherin Jubbul-
pore and discussed the advantages of forrning an Association. It was decided
to call a meeting of all those interested in angling, and, if the necessary
support was forthcoming, to formally consider the question of forming an
angling association.
This meeting was held at the Nurbudda Club, Jubbuipore, on March 1927,
at which 13 gentlemen were present.
It was decided that the main objects of the Association should be to collect
and maintain all records relating to Fishes and Fishing and generally to con-
sider the adoption of any course or methods which might further the advance-
ment of or improve the sport of Angling in the Central Provinces.
As regards progress the report states—
Rivers.— Owing to the fact that the Association only commenced to function
practically at the close of the fishing season, it was not found possible to
arrange any Association ‘ baiting’. This will, however, start as soon as it is
practicable after the monsoon waters have sufficiently subsided. The number
of ‘runs’ to be baited will, of course, depend on the number of ‘ baiting’
subscribers.
Tanks.—Iit is to be regretted that, up to the present, it has not been possible
to make any satisfactory arrangement with the irrigation authorities for the
preferential recognition of the Association in respect of the tanks under their
charge.
With regard to identification. It is interesting to note that allthe larger carp
caught in the water-works area during the period for which this water was
open to the Association have been proved to be Barbus hexastichus (Day’s
Fishes of India, p. 565, plate cxxxvi, fig. 4), and not ‘ Barbus tor’ (the
ordinary Mahseer). Day records these fish as growing up to 3 feet in length.
The only other fish that have so far been formally submitted for identifica-
tion proved to be two Macrones seenghala (one ot the Tengara or cat-fish
group), which were caught with asmall spoon ina pool inthe R. Gaur, up-
stream of the village of Kosamghat lying about a mile north of Jubbulpore-
Mandla road between the seventh and eighth mile-stones. The two specimens
recorded measured 144 and 133 inches respectively.
ee
OBITUARY
JOHN WILLIAM YERBURY
We regret to record the death on November 10 last as the
result of being knocked down by a motor car, of Lieut.-Col.
J. W. Yerbury, late R.A. Col. Yerbury who was aged eighty at the
time of his death, was born on March 30, 1847, at Serampore in
Bengal, and was the son of Lieut.-Col. John William Yerbury, of
the 3rd King’s Own Light Dragoons and afterwards of Belcombe,
Bradford-on-Avon. He obtained his commission in the Royal
Artillery in 1868 and up to his retirement in 1892, served in various
parts of the world. He was for many years stationed at Aden, and
from his papers on the Birds of this settlement (which appeared
in the /dzs for 1886 and the Society’s Journal for 1897) he was well-
known to ornithologists.
Col. Yerbury was one of the earliest members of the Bombay
Natural History Society, being on the rolls in 1886 when the
Society issued its first Journal. Besides ornithology, he was keenly
interested in Mammals, Reptiles, Lepidoptera and Diptera, of all of
which he made considerable collections, especially of the last named
which he eventually presented to the British Museum. He also
collected shells at Aden, several of which were described by the late
E. A. Smith (Proc. Zool. Soc. London, June 16, 1891) as new species
and named after him. After his retirement his chief interest appears
to have been confined to Diptera on which group of insects he
wrote several valuable papers.
His most important contributions to the Journal of this Society
are as follows :—
‘The Butterflies of Aden and Neighbourhood, with Some
Notes on Their Habits, Food Plants, etc.,’ vol. vii, p. 207.
‘ A List-of the Birds of Aden,’ vol. x1, 1p. 7/5.
His appeal for assistance in collecting Gad Flies, Bot Flies and
Warble Flies (vol. xiii, p. 683) contains some very useful instruc-
tions for collecting these insects.
Officers quartered at, and residents residing in Aden have cause
to be indebted to the late Col. Yerbury for leaving behind for their
use withthe Political Resident at Aden, a set of valuable pamphlets
on the Flora and Fauna of the settlement ‘as complete as I have
been able to make it.’
EUGEN M. VAN INGEN
The death occurred at Mysore on February 27 of Eugen M.
Van Ingen the well known sportsman and taxidermist, after a
severe operation. Mr. Van Ingen joined the Bombay Natural
History Society in 1913 and was known among a very wide circle of
OBITOCARY 787
friends, as the very best of good fellows. Mr. Van Ingen’s fame
as an artist in taxidermy was only equalled by his keenness and
proficiency as a shikari and fisherman. Some of the biggest known
records in jungle and stream were annexed by him and his advice
and assistance were freely sought in big game expeditions
organized in the State on the occasion of the visit of distinguished
guests, notably the Prince of Wales, the Earl of Reading and H. E.
Lord Irwin. His death, which creates a great gap in the personnel
of Indian shikar, will be deeply deplored by hosts of sportsmen
throughout the country.
MISCELLANEOUS NOTES
I.—THE TRUTH ABOUT ‘A UNIQUE LION PHOTOGRAPH’
We are glad to notice that the photograph of a young Lion
attacking a Zebra, published on page 583 of this Volume has
aroused such considerable interest amongst our readers.
Writing on the 20th of March, Mr. F. W. Champion, the well-known
wild animal photographer and author of W7zth a Camera in Tiger-
land, puts forward a number of well-considered and cogent
reasons for his belief that the photograph is one of those ingenious
fakes which are unfortunately much too common with Natural
History subjects. Quite naturally, however, he could not be alto-
gether certain that it was a ‘ triumph of the faker’s art’, and admit-
ted that if genuine, it must in his opinion be ‘by far the finest
natural history photograph ever taken by anyone or likely to be
taken.’
Sometime back we had written to Messrs. Hutchinson & Co.,
the publishers of Azzmals of all Countries, for information regarding
the same photograph reproduced in their book, and referred to in
the note which accompanied our plate. To this we recently heard
from Mr. Wm. Martin-Hurst, Managing Editor of the Exclusive
News Agency, London, (‘ E.N.A.’) who kindly wrote to say that he
could give us no particulars beyond the fact that their original photo-
graph was a very old copy, having been in their possession for at
least 25 years; that as far as he knew it came _ originally
from an American source and was said to have been taken by a
well-known American Big Game Hunter; and that various people
more or less qualified to speak had cast considerable doubt on the
authenticity of the picture.
Still more interesting are the cuttings from Zhe Camera for April
and June 1927 received by the last mail through the kindness of one
of our members, Mr. A. J. Page, which help conclusively to run the
photograph to earth. The April (1927) number of Zhe Camera
in noticing a monthly Magazine of photography issued by the
Soviet from Moscow entitled ‘ Sovietskoe Foto’, reproduced from
its pages the same Lion-Zebra photo with the following graphic and
soul-stirring account: ‘One of the pictures which we endeavour to
reproduce is without exception the most dramatic and tragic nature
picture that we have yet seen. It is said to have been taken by a
hunter on the plains (or steppes) of Africa. It shows the triumphal
ride of a lioness on the back of a Zebra, terror depicted on the
unfortunate animal’s features as in vain he flies to escape from the
fate that awaits him. It reminds onein its wild way of the triumphal
march of an emperor through the streets of Rome celebrating his
victory. Nothing was spared to display the barbaric glory of the
victor, even to the march of his conquered victim, barefooted and in
chains. Here is the Queen of the jungle, emblem of ferocity and
cruelty, and it requires no imagination to realize that in an instant
(II ON 230N snosuelpasstf, 99g) “(vsojnqau sya) predoay papno[y eq,
‘DOS 'LSIH “LYN AVENOG ‘Naenor
MISCELLANEOUS NOTES 789
the Zebra must collapse and that the Lioness’s fangs will be in its
throat.’
The publication of the Soviet photograph in 7he Camera promptly
elicited the following letter to the editor from a good lady in Bake-
well, Derbyshire, whose name is not given as the gentleman she
refers to (whom we by no means envy!) has quite wisely preferred
to remain silent in spite of being addressed on the subject-
‘Dear Sir—It may be of interest to you to know that the actual
photographer of the incident you reproduce of the Lioness and
Zebra, resides here in Bakewell. Having had in my possession a
copy of this, I was more than astonished to find it on the front of
your magazine when in a Sheffield photo shop last month. I was
in conversation with the gentleman only yesterday and he assures
me that the incident did really happen as depicted, and that he was
a member of the hunting party. It certainly did not happen recent-
ly, as he has left South Africa since 1916. How it has got to Soviet
Russia has yet to be explained.’
While these amusing and amazing claims and counterclaims were
in progress, the matter seems to have been brought to the notice of
the ‘iconoclastic’ Dr. G. Arnold, the Curator and Zoologist of the
Natural History Museum, Bulawayo, whose conclusive letter on the
subject expresses our sentiments admirably. Dr. Arnold writes
under date 29th April 1927:
‘Dear Sir—It is quite correct that the photograph on the cover
of Zhe Camera is taken from part of a group in this Museum.
Some years ago a photo was taken of this group, and, with a
faked background added, was published in the United States with
the information that it was a snapshot taken by an intrepid hunter.
‘Liar’ would perhaps have been a more correct description of the
intrepid gentleman. The photo on the cover is not new to me,
as it was published elsewhere some years ago, and broughr to my
notice.’
Our thanks are due to those members who have communicated to
us their views on the photograph, and helped towards the final
elucidation of the mystery.
EDITORS.
Il.—A CLOUDED LEOPARD (#EL/S NEBULOSA)
ATTACKING MAN
(With a_ plate)
I enclose a photograph of an animal which was brought in by a
Karen boy who stated that while cutting wood near his village he
saw the animal approaching him and when it saw him it crouched
and approached him cautiously ‘pushing bundles of branches and
leaves in front ofit.’ It sprang at the boy who split its skull with a
dah (knife). I find on inquiry that this is generally believed to be the
usual method this animal adopts when approaching its prey whether
as a means of concealment or to puzzle its prey I amunable to say.
Perhaps you could enlighten me on this point.
790 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
The animal is a species I have never seen or heard of before ; it
is known locally as ‘ In Cha.’ .
This particular individual is known to have taken several head of
cattle in the vicinity.
Tavoy, BuRMA, A. BROWNLOW,
December 10, 1927. L[ndian Police.
|The identity of the animalin Mr. Brownlow’s note and photo-
graph as the Clouded Lecpard (felis nebulosa) is confirmed by the
skin sent us for examination. This beautiful and somewhat rare
cat inhabits the hilly portions of Burma, Siam, Malay Peninsula,
Sumatra, Java and Borneo and also occurs in Assam, Sikkim,
Bhutan and the Nepal Himalayas. It attains a length of from 6 to
63 feet of which 23 to 3 feet is taken up by thelong and thickly
furred tail.
Our knowledge of the habits of this animal is very scanty and
Mr. Browniow’s note forms an interesting addition. Itis believed
principally to be an arboreal species feeding on small mammals
and birds which it captures among the branches. Eps. ]
IlI.—NOOSING TIGERS
Among the coolies on these hills there has been a custom
(which I have strenuously tried to stop) of setting traps and wire
nooses for the catching of Sambur, Barking Deer and Pig.
I was informed one day last month that a tiger was heard
‘Making plenty big noise’ at aplace about four miles from my bunga-
low. On arriving at the scene [found him to be about 98 per cent
dead and proceeded to put him out of his misery.
On examination I found that the tiger had been caught around
the neck by a wire noose, presumably placed for the unwary
sambur. The wire had been placed acrossa well-trodden sambur
track.
The wire in question had a total length of 10 ft. 7 in. was 5 ply of
approximately 20 gauge. The far end was affixed to small sapling,
at a height of about 3 ft. from the ground.
The beast was a full grown male, in healthy condition, measuring
9 ft. 8 in.— ‘ fair do’s’ --between pegs.
Last year a similar case occurred on a neighbouring estate, the
wire in this instance was ordinary telegraph wire. The beast, I
am told, measured, 9 ft. 7 in.
Yet another case occurred last year in which a tigress was
noosed, of the particulars of which I am afraid I am ignorant.
How is it that a noose, presumably placed for snaring sambur,
will catch a tiger which usually walks, and even stalks with his
head down?
PALAGAPANDY,
NELLIAMPATHIES, H. A. BOAS.
November 22, 1927.
MISCELLANEOUS NOTES 791
IV.—REPLY TO MR. PHILLIPS’ NOTE ON P. 371,
VoL. XXXIT RA-COLOUR OF THE LONG-ARMED SHEATH-
TAILED BAT (TAarAOCZOUS LONGIMANTGS)
Iam quite in agreement with Mr. Phillips that the young of
T. longimanus are coloured like the females (and vo/ like the males)
as is the general rule with all juvenile animals.
It is very apparent that he has confused the statement of Blanford
with mine, as will be clearly seen from the paras which I reproduce
below :—
Mr. Phillips writes, ‘Ido not, however, agree with the sugges-
tion that the young of this bat are pale fulvescent and become
gradually black with age. I am convinced from my experience
(soam I.,C. McC.) with species in Ceylon, that the opposite is in
reality the truth, that is to say, that the young are darker in the first
coat and become gradually lighter and more brightly coloured with
age, the adult male becoming light reddish (almost cinnamon) brown
eventually.’
The following is an extract from my note in the Journal vol. xxxi
p. 1030 :—
‘All the three males were much lighter in colour than the
females, being a yellowish-brown: the females on the other hand
were dark blackish-brown, almost black. Blanford, inthe Fauna of
British India, mentions that Blyth observed the young individuals
to be pale fulvescent and that they become gradually darker with
age. Whether these paler ones were males or females he does not
say, for they might have all been males. One male taken by me the
day before was even paler than those secured in the Museum, being
a very pale fulvescent tint.
On looking throughthe materialin the Society’s collection I found
this difference to be the general rule, only a few being like the
females which are probably young specimens. One at least is very
young and coloured like the female. In almost all cases I could
pick out the males from the females by the difference of colour
alone.’
With regard to the question, ‘Whether these paler ones
for they might have all been males?’ I doubted Blyth’s statement
regarding the age of the bats. I did not mean to suggest for a
moment that the young were coloured like the males! It is quite
clear from my statements in the second para‘... only a few
being like the females which are probably young specimens,’
and ‘one atleast is very young and is coloured like the female,’
that I was of Mr. Phillips’ opinion that the young were coloured
like the females and not like the males.
BoMBAY, C. MCCANN.
January 10, 1928.
V.—HABITS OF THE PORCUPINE
Mr. C. McCann’s note on the Habits of the Porcupine in
vol. xxxil, No. 1, has somewhat surprised me. Porcupine exist
in large numbers on these hills (the Billigirirangans) and it is
>
792, JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
very noticeable that not only their little paths, on the hill-sides, but
also paths formed by deer and other game are strewn with their
droppings but I have never noticed cone-shaped accumulations of
their excreta. The little Four-horned Antelope, which has only two
horns in these parts, and is known as the Pokey, is addicted to this
habit and the above is a good description of the little mounds of
excreta formed. by them: but similar formations in the case of
porcupines are not seen here.
HONNAMETTI ESTATE, RALPH C. MORRIS.
ATTIKAN-P2@:
via Mysore,
December 11, 1927.
VIL—AN ELEPHANT WITH EXTRAORDINARY ‘GLANDS’
(With a plate.)
In answer to your letter of December 29, regarding the
Bellagalla Rogue Eiephant which I shot and sent you the snap of,
which had two extraordinary glands on the neck, which I thought
might be of interest to you, I will briefly give you some particulars
which may be of further interest.
This elephant used to frequent the forests between Coimbatore
District and Mysore State, adjoining the Billigirirangan Hills. I
shot it in the valley about half-way between Attikan and Kollegal.
During the past two years or more, it had become a great menace
to the people living in the wilds round about and it took to destroy-
ing their crops and no one dared to go anywhere near to drive it
away without being chased by it. It made short work of a few
unfortunates whom it got hold of and from all accounts it simply
trampled them out flat. It was proscribed a rogue about two years
back and though many set out after it, they did not have the
fortune to meet it. Mr. Ralph Morris of Attikan who I believe is
well known to you, promised to send me a wire if any reliable
khubber reached him about rogue elephants, of which there were
many, making their appearance off and on round about Attikan, and
as luck would have it he wired to me on October 24, and I
managed to get to him on October 26. We got into touch
with the rogue on 29th and after much tracking we came upon
him in an opening in the thick jungle, quite suddenly. I feel sure
he had his eye on us. He stood quite still about 40 yards off
giving usa broadside shot. Morris let me have the shot and I got
in a well placed shot between the eye and ear which stunned him
and Morris followed up with another shot immediately after and he
swung round falling on his side on the slope of the hill as the
picture shows and I placed another good shot at the base of the
skull which ended his career. It was all over in a minute or so.
On going up to him I noticed the two glands (I call them glands
but may be I am wrong).
The one you see on the upper side (right) was about 3 to 4inches
long, whereas the one on the underside was quite, 9 inches long.
They were both very slimy in appearance and to feel. They were
‘yisue] ul sayout 6 ynoqe JaMoyT pue sayour + saddy
‘(X) yoou uo syIMoIs dyT][-puR[s selpNded oy SuIMOYsS JOYsNy, s,atneyT “So YW ‘ATW
‘90S ‘“LSIH ‘LYN AVEWog ‘Nunor
MISCELLANEOUS NOTES 793
coated with a whitish fluid (slime) and seemed to be merely two
bags or envelopes, which you could distend, being very elastic.
They were black in colour. The lower one looks white due to the
sun shining down on it and probably the reflection from the slime
causes it to look white in the photo.
The light was very difficult being about 8.45 a.m. and the sun
was just peeping over the hiil through the thick growth. The
photo I send (reproduced herewith) was the best cf three snapshots.
By the way I know Mr. Ralph Morris cut off the lower gland with
the idea of sending it to you. Ido not knowif he actually did so
for I had to leave hurriedly.
The people of the forests who accompanied us told us that they
had never seen any elephani with this extraordinary formation
under the neck and I have made enquiries of others who have had a
lot to do with elephants and they say the same. It will be interest-
ing to learn if possible what function they could possibly perform.
COIMBATORE, A. 5. LAURIE.
S. INDIA,
January 4, 1928.
[Regarding, the above tusker and the extraordinary ‘ glands’
Mr. Ralph C. Morris writes us as follows :—
‘Iwas with Mr. Laurie at the time he shot the elephant and
we were both amazed to see, when we went up to the fallen
tusker, thatit carried a most extraordinary pair of appendages
on its neck, one on either side behind the lower jaw-bone as can
be clearly seenfrom the photograph. The appendages were inthe
shape of sausages, the upper one being about 5 inches long, and
the lower a good 9 or 10 inches. ‘To the touch they were tender
and appeared to be covered with a skin of unusual thinness, the
skin being of a raw appearance, mottled grey-pink in colour and
moist. Apart from the fact that the elephant was clearly in ‘‘ Musth’’,
there being a flow from the ‘“‘ Musth”’ gland orifices, I cannot
think what was the cause of these extraordinary growths, nor can
I assign any reason for them. I referred the question to Mr.
Saunders, Principal of the Veterinary College in Madras, but
Mr. Saunders couid advance no theory on the matter. He had seen
nothing of the kind before, and although I have been in at the death
of eleven or twelve proscribed elephants I have never seen a similar
case before, nor have I seen growths of this kind mentioned in any
work on elephants. I had the larger of the two appendages cut off
and gave instructions for it to be pickled preparatory to sending
it to the Madras Veterinary College for examination. Unfortunately
my instructions were misunderstood by my skinner who, instead of
preserving the whole appendage, skinned it, threw away tlie inside,
and pickled the skin! Can you ascribe any possible reason for these
growths, and have you heard of a similar case before ?’
Mr. A. J. W. Milroy, 1.F.S., of Assam, who has had considerable
experience of elephants, both tame and wild, writes to say that he
has never come across any glands of this sort.
23
7941 JOURNAL, BOMBAY NATURAL HIST, SOCIETY, Vol. XXXI1
Mr. N.L. Bor, 1.F.s., of Goalpara District, Assam, another authority
on elephants writes in this connection as follows :—
‘As far as my experience goes I have never seen growths of the type
mentioned by your correspondent. Tumours of a fatty or serous
nature are not uncommon and | have seen them both in tame and
wild elephants. These tumours which are soft and elastic are not
infrequently found on the folds of the skin on the neck. Those
which are serous in type contain a fluid which is rather slimy and of
a barley-water colour.
It seems to me that the growths in question, I do not think they
can be glands, are of a similar nature which may have possibly
become chronic or malignant.’
It is extremely regrettable that owing to a misunderstanding the
contents of the ‘ glands’ should have been lost. It would be interes-
ting to hear what Zoologists have to say about these growths or
appendages. Eps. ]
VIL—ON ‘NATURAL DEATHS’ IN WILD ELEPHANTS
With regard to Mr. F. W. Champion’s interesting article on
‘Wild Elephants in the United Provinces’ in vol. xxxii, No. 1, in
which he quotes instances of elephants having been found dead,
the case of the elephant found dead inthe Lansdowne Division
certainly seems to be one of death from old age. Is there not just
a possibility however that it had been poisoned by villagers ?
Mr. Champion mentions that the graziers had known the elephant
for a year, and that it was too old and weak to run away with the
other wild elephants when driven from the crops of the netghtouring
villagers. This may be so: but I have known of elephants
frequently refusing to be driven off from villagers’ crops, remain-
ing in cover close to the fields when the rest of the herd had retreated
back into the jungles. I once came upon the decomposed carcases
of three cow elephants at the foot of these hills. The iocal forest
subordinate informed me that he thought the elephants had proba-
bly died of anthrax—though he could advance no evidence in support
of his theory: no other elephants had been reported sick or died:
nor did I hear of any other deaths. The elephants were lying dead
within a few yards of each other. Ido not think that the anthrax
theory was possible, nor do I consider it at all likely that the
elephants had died of natural causes. I was assured later by my
trackers, that the ryots in the fields these eiephants had been in the
habit of visiting and causing considerable damage to, kad put out
a number of poisoned pumpkins, etc., which these elephants had
eaten. Here there was the possible cause of their death. Mr.
Champion speaks of the elephant having been moved from its first
position. This is quite possible. I have seen a wild elephant
trying tomove a tusker (a proscribed rogue) I had shot: and in
the case of a herd tusker shot by the late Mr. R. H. Morris the carcass
was actually shifted a few yards by other elephants of the herd
which returned to the spot Jater. Atuskershot by Mr. A. L. Laurie
of Coimbatore in November 1927, when Iwas with him, ona slope
above a stream was moved a few yards down to the edge of the
MISCELLANEOUS NOTES 795
stream by elephants in a herd which visited the place a day or two
later. This elephant was clearly in ‘ Musth’, there being a discharge
from the ‘ Musth’ gland orifice between the eye and the ear on
either side of the head. Itis remarkable how quickly the skeleton
of an elephant will disappear, and Mr. Champion in no way
exaggerates when he says that ‘a single season may easily remove
the entire body and much of the skeleton of an animal even as large
as an elephant.’ A couple of years is sufficient to remove all
traces of a dead elephant—except perhaps the skuil which may
be found hidden away in long grass and dense undergrowth for a
year ortwo longer. Protection has undoubtedly contributed to the
increase of elephants in South India. Probably the Kheddas held
by the Mysore Government do more than anything else to keep
their numbers down in these parts, but even then I consider that
elephants are rapidly increasing. When I was in camp last March
there were at one time four separate and distinct herds of elephants
near our camp and three or four solitaries besides.
I have just received Mr. Champion’s book Wzuth a Camera in
Tiger Land. Vhe photos shown of Tiger, Panther, Biephant,
Deer, etc., in their natural state are wonderful and the reading
enthralling. Mr. Champion writes as only a keen naturalist and
lover of jungle and its denizens could write. The photographs are
superb, and one can imagine the trouble and painstaking care
involved in the achieving of such thrilling results.:
HONNAMETTI ESTATE, RALPH C.-MORRIS,
ATTIKAN (P;O.,
Via Mysore,
December 11, 1927.
VIII.—CLICKING NOISE MADE BY MUNTJAC
In your issue of August 1, 1927 ‘ Miscellaneous Notes ’ I have read
a note with the above title by Mr. Randolph C. Morris.
I am surprised to learn that Col. Burton has never heard the
Muntjac (the Khakar or Barking Deer of Northern India) emit this
clicking noise. I spent many years inthe forests of the United
Provinces, between 1889 and 1918, and very frequently heard the
noise described, especially when the Khakar was disturbed at cluse
quarters.
The noise is, as stated by Mr. Morris, only produced as the
animal bounds off. I believe it to be restricted to the male only
but I cannot be absolutely certain of this.
The noise is a rapidly repeated cluck—cluck—cluck. ... Generally
only six to eight repetitions, and I have always believed to be pro-
duced in some way by impact of the animals’ teeth. It is far too
rapid (about three to the second) to be produced by the feet andas a
matter of fact a very similar sound may be produced by striking
two pieces of bone together.
Ba Bb, OSMASTON;
indian forest Service (Retd.)
796 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
IxX.—CROWS AND KOELS
M. Jouguet’s letter in the number of the Journal published on
October 20 last reminds me of an occurrence which came to
my notice some three months ago, which may be worthy of
note.
Crows having become an insufferable nuisance round the bun-
galow, | had taken to shooting them, in the forlorn hope of driving
them away. Asis the way with crows, they had become sufficiently
knowing to keep just out of range, and we had reached the stage
when, by careful stalking, I occasionally got one which was either
too daring or too slow.
One evening, shortly before sundown, this game was in progress,
all the birds being between me and the sun, when suddenly there
was a loud ‘caa’ from the far side of a Derrts robusta tree near by.
Looking up, I could just make out a black body through the
feathery foliage, and shot it.
On picking up the bird, I was astonished to find that it was a
female koel (Audynamzs scolopaceus malayana?). It had appeared
to be black from being against the sun, and its ‘caa’ was sucha
perfect imitation of the real thing that, although I could not
make out the shape of the bird, I had had no doubt that it was a
crow.
Has this occurrence been recorded before ? If so, is the object
of itknown? Had it been a different time of year one might have
supposed that the koel was endeavouring to mix with the crowd to
accompany the crows to the roosting place. There was no sign of
a male koel, though I watched until the birds began to clear away.
Moreover, it is the only time in six years of residence in the
bungalow that I have known of a koel anywhere near the
compound.
TocKLAI EXPERIMENTAL STATION, E. A. ANDREWS,
CINNAMARA P. O. & T.O. Entomologist,
ASSAM, Indian Tea Association.
December 19, 1927.
X.—THE RUFOUS-BREASTED BLUE FLYCATCHER
(CYORNIS AY PERYTARA) AY HIGH LEVELS
While examining recently the series of Cyornis hyperythra (Blyth)
belonging to the Bombay Natural History Society’s Museum
I found two specimens differing markedly from the rest which were
in agreement with authors’ descriptions: Oates, Baker, etc.
The two skins are generally paler greyer-blue above; much less
rufous below and perhaps have more white on the tail than in
examples representing the typical race which has Darjiling for type
locality. They are also paler and greyer than the south-eastern
forms.
MISCELLANEOUS NOTES 797
One of the birds was taken at 6,000 ft. on Mt. Victoria, Burma, and
the other at Dakuri, 10,200 ft., on the Pindar R., north of Almora in
Kumaon.
These localities are practically at the extremities of the range of
C.h. hyperythra so it seems out of the question to regard them as
representatives of a distinct race unless that should be a high-level
form.
This note is written to suggest that those possessing other high-
level specimens should compare them with more _ topotypical
examples.
SINGAPORE, C. BODEN KLOSS.
December 20, 1927.
XI.—THE NIDIFICATION OF THE SIND HILL WARBLER
(SUYA CRINIGERA STRIATULA).
In the Fauna of British India, Birds, Revised Edition, vol. ii,
p. 520, the nidification of the Sind Hill Warbler (Suya crinigera
striatula) is shown as not recorded, and the following note may
therefore be of interest.
During the last week of July of this year I found the bird com-
mon on the hill of Sakesar, some thirty miles east of the Indus and
at the extreme west of the Punjab Salt Range, of which it forms
the highest point (4,992 feet).
Its favourite haunts appeared to be steep slopes covered with
long grass and well supplied with bushes of various kinds. Nest-
ing operations were in progress and several nests were found,
mostly in course of construction.
In one nest a bird was sitting on two eggs on July 24, and
again on 28th, when the eggs were taken and the male parent
secured. The eggs, which were slightly incubated, have a ground
colour of very pale pink and are marked with pale reddish blotches
scattered over the whole surface and most numerous at the broader
end, where they form an ill-defined ring. The nest was situated,
barely twelve inches from the ground, ina‘ Sanatha’ (Dodonea
viscosa) bush, through which some tufts of grass were growing. It
was an oval-domed structure, made of strips of grass, and the lower
portion was matted with seed-down. The inner lining was of fine
grass, and the entrance faced north.
Another nest, similarly built and situated, contained three half-
fledged young on July 28.
A small series of the birds was collected and sent to Mr. H.
Whistler who kindly identified them.
I have not met with this Warbler in any other part of the Salt
Range.
SARGODA, H. W. WAITE, F.z.s., M.B.0.U.,
PUNJAB, Indian Police.
October 15, 1927.
798 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
XII.—ON THE UNUSUAL FOOD-HABITS OF HORNBILLS
Apropos the subject of the unusual food-habit of Dzchoceros
bicornis, Linn., as recorded by Mr. W. S. Wood in J.B.N.4.S. (vol.
XXXVii, page 374) it is worthwhile to cite an instance of the
piscivorous habit of Lophoceros birostris (Scop.) which came under
my personal observation. It was in 1926 when at Hazaribagh I
bagged alive a specimen, very slightly injured by my gun-shots.
Anxious as | was to keep it in captivity, 1 was confronted with a
vexed problem as to itsdiet. Knowing it to bea frugivorous bird I
tried all sorts of fruits and berries, which I found it, to my amaze-
ment, to avoid studiously. It was literally starving itself for a
couple of days and would touch nothing except water. When I tried
to force a piece of ripe banana down its throat it would simply dis-
gorge the whole thing out. At last when some small fish were
presented before it, it readily paid attention to the fry, and after
throwing these up one by one in the air, it would catch them in its
bill and gulp down. The fact of this bird having any greed for fish
has never been recorded anywhere. In this case aiso the question
arises whether it is a practice commonly indulged in by these birds
in their wild state or merely a case of individual idiosyncrasy
noticeable in captivity. The fish-eating habit has only been
observed in the case of Azthracoceros m. malabaricus (Gmelin). The
other members of the Bucerolide appear to be largely frugivorous,
although a varied menu consisting of smaller reptiles and rodents,
snails and various insects is also more or less induiged in.
R.B. Sanyal, a former Superintendent of the Calcutta Zoo, prescribed
for the captive Hornbills a carnivorous dietary, within which small
birds were included (vzde his Handbook of the management of
animals in captivity in Lower Bengal). It is, therefore, easy to
realize that with a hungry Hornbill nothing goes amiss provided its
size falls within manageable limits. I have even seen Lophoceros
btrostris (Scop.) devouring the petals of Lauhinia flowers, which
appear to form a major part of the bird’s diet in places where there
is a comparative dearth of fruit trees and berries. The omnivorous
habits of the Hornbills are no doubt responsible for their occasional
outbursts of criminality (which finds a full play in captivity when
weaker companions are housed with them) but how far the predatory
instinct is let loose so as to play its part in Nature’s economy is a
point which cannot be answered until all the data are available as to
the food habits of each species.
CALCUTTA. SATYA CHURN LAW,
M.A., B-L.; PH.D., F/Z.S., M.D.0.U,
XII].—THE BLACK-CAPPED KINGFISHER (HALCYON
PILEATA) IN THE DARBHANGA
DISTRICT Bin AR
I have on several occasions recorded the occurrence of this bird in
Bihar in these pages, most of them being got in February. It is
rare but not asrare as | at first took it tobe. I now record another
which I shot in September.
MISCELLANEOUS NOTES 799
On the afternoon of September 17, 1922, while out in boat
fishing with my brother on the Bhagmati River near Bunhar factory
we got within five yards of one of these birds. The river is an
Open one running through cultivation and the Kingfisher was ona
bush overhanging the water. It took absolutely no notice of the
many small fish which came to the surface nor of us in the boat till
we got within that distance of it; even then it only flew across to
the other side of the river. We disturbed it on severai occasions
and what I noticed most was that it flew off silently on each occa-
sion quite unlike an Indian White-breasied Kingfisher (Alalcyon
smyrnensts fusca) which went off uttering its loud, harshcry. We
came across it again on the 19th, this time seated on the
bank and finally I shot it onthe 20th. It was silent on all three days
so [have never heard its cry. Itis easily recognized on the wing by
the buffy lower plumage, the upper plumage looks black except
when the sun shines on it, then it glistens. The contents of the
stomach consisted of a small crab; other specimens examined had
fed on fish and beetles. The local Aftrshzkars consider it only an
albino of the white breasted species and call it Ad/ak Tankz.
DARIJILING,
September 28, 1927. CHAS. M. INGLIS, F.z.s., F.E.S., M.B.O.U.
XIV.—A QUAIL’S QUEER RETREAT
(Reprinted trom the Sunday Times, Johannesburg, Transvaal,
November 13, 1927.)
A singuiar case is reported from Somerset East, C. P. A shearer
working at Hope Vale, a farm in the district, was busy on a heavy
woolled ram of Messrs. W. G. Jordaan and Sons, when he felt a
hard lump in the wool on the belly.
On opening up the fleece he found a live, fully-grown quail !
It is supposed that the bird had crept for shelter into the long
wool while the ram was lying down, and had remained there when
it was driven to the shearing shed. It is stated that there were
several witnesses of the find.
Has anything like this been found elsewhere ?
XV.—OCCURRENCE OF THE SHELDRAKE (TADORNA
LADORNA) IN THE DARBHANGA
Dist RICl BIAAK
There have been notes in late Journals of the occurrence of this
duck in various parts of Bihar, so it may perhaps be interesting to
gather together data of those got by me; some of these have
already been noted by me in this Journal, my first record being in
Vole exXV.. Ds ooo Of 1905, During that year I got one from
Mr. Edgell and another was snared not far from Baghownie. In
1905 I got four specimens, two drakes from the Benoa chaur on
May 1, and June 5, and two ducks, one on November 6, near
Baghownie and the other on the 30th of that month from the Maiser
chaur. {don’t appear to have got any more til 1915 when one was
800 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXII
brought in alive by a Wirshtkar on July 28. It was in good condi-
tion and didn’t appear to have been wounded at any time. During
the following year I got a fine drake near Baghownie on May
12, with a well developed knob on its bill. It was on a small] patch
of mud and water, more mud than water. ‘The last record I
have is of a duck got at the Benoa chaury on June 24, 1923 another
late date for it to be found. After that I left the district but have
no doubt that had I remained there I would have been able to add
to these records. Besides these nine specimens I have also heard
of one or two others being shot.
DARJILING,
September 28, 1927. CHAS. M. INGLIS, F.z.s., F.z.s., M.B.O.U.
XVI.—OCCURRENCE OF THE SHELDRAKE (7TADORNA
TADORNA) IN THE UNITED PROVINCES.
On December 25, 1927 at Saman Jal, Mainpuri District, United
Provinces, there were shot two specimens of the Sheldrake
(Tadorna tadorna).
ROORKEE,
January 4, 1928. BE. E. L. SEARIGHT, CaZi.,
R.E.
XVIL—RECOVERY FROM THE BITE OF A PHOORSA
(ECHIS CARINATA.)
No. 4179383. Fslr. Brannar, lst Royal Welch Fusiliers, had been
employed on fatigue, with other men, in knocking down a wall.
The wall had been knocked down and the men were carrying away
the stones. Brannar lifted up a large stone and was bitten by the
snake, which was underneath. This was at 5.10 p.m. on October 4,
19-7. He reached hospital about 5.30. There were two well-
defined punctures on the dorsum of the terminal phalanx of the
right middle finger, which were slightly oozing. The orderly assis-
tant surgeon incised them and rubbed in permanganate crystals.
He ‘vas seen by the orderly officer about twenty minutes later and
was given 40 c. c. of antivenin serum, subcutaneously, in the flank.
He did not show the faintest trace of constitutional disturbance.
He showed us a small gland in his right axilla, and said that he
felt a slight tingling running up from the finger to the gland, and
added that the tingling was not painful. He was detained for 36
hours, and then discharged to ‘ attend’. I saw him, this morning.
He assured me that he had not felt the slightest ill-effects, at any
time, and he is an intelligent man. There was slight thickening
over the incisions, and one was oozing a trace of serum. The
finger was not painful,
MISCELLANEOUS NOTES 801
The stone had already been disturbed with a crow-bar, before
Brannar picked it up. It seems to me that the absence’ of all signs
of poisoning may be due tothe snake having struck at the crow-bar
and emptied its poison sacs, before Brannar lifted the stone.
BRITISH MILITARY HOSPITAL,
NASIRABAD, A. A. MEADEN,
October 9, 1927. Lt.-Col., R.A.M.C.
[The snake that bit Fslr. Brannar has since been sent to us and
identified as the Phoorsa (Echts carinata). We append a note
on the subject of Indian snake venoms by Lt.-Col. F. P. Mackie,
LM.S., Director of the Haffkine Institute, Bombay, which is of great
interest. He writes :—
‘The mortality from snake bite in India is due in the large
majority of cases to one of four species. The cobraand the krait
which are known as colubrine snakes, and the Russell’s viper and the
Echis (or phoorsa) which are viperine.
The poison of colubrine snakes affects chiefly the nervous system
and produces paralysis of various muscles, staggering gait or incoor-
dination of speech followed by respiratory paralysis, convulsions
and death by respiratory failure. The symptoms of viperine poison
are severe and persistent pain at the position of the bite and oozing
of blood from the punctures. Death may take place from paralysis
of the heart or from clotting of blood in the vessels if the dose is
large or is injected into a vein but if life is prolonged hemorrhages -
take place from the mouth, nose or intestines or in various part of
the body. Mortification and gangrene occur at the site of the fang
marks and the finger or hand may be lost in this way even after the
general symptoms have passed off.
The minimum fatal dose'of cobra venom is about 15 maul tani
(about 1/5th grain) and the snake gives from 5-10 lethal doses at
one bite. The krait can deliver five times the lethal dose and the
Russell’s viper and Echis about twice the lethal dose at one bite.
The Echis however rarely gives a sufficient dose to kill an adult
man.
Death from a cobra bite has occurred in as short a time as 20
minutes, but may be delayed as long as 30 hours.
The antivenine produced at Kasauliis made from the venoms of
cobras and Russell’s vipers which are collected at the Haffkine
Institute and as these poisons are absolutely specific in their action
the antivenine is quite useless against the bites of kraits and
phoorsas.
Antivenine should always be given into the veins and not under
the skin and if given in the following doses will be instrumental in
saving many lives especially if given within a short time of the
bite :—
Russell’s viper bites 80-160 cc. of antivenine
Cobra bites 100-400 cc. ,,
(Acton & Knowles)
Permanganate of potash has little or no effect if rubbed into the
bite but generally increases the amount of necrosis and tissue
destruction.’ Eps. ]
24
802 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXI1
XVIII.—THE MOGUL EMPERORS OF INDIA AS
NATURALISTS AND SPORTSMEN.
(Published with the permission of the Director,
Zoological Survey ot India).
Mr. Salim A. Ali has done a great service to science in bringing
together all references to animals contained in the works of the
Mogul Emperors of India and in arranging them in a systematic way.
No. 2 of volume xxxii of the Journal of the Bombay Natural History
Society recently published, contains an account of fishes (pp. 268, 269)
as described by these emperors. The descriptions are in almost all
cases sufficient to enable the animals to be identified fairly closely.
Mr. Ali has, however, not been able to do full justice to these des-
criptions in his interpretations and I therefore propose to add a few
comments to the various passages quoted by Mr. Ali.
Babur refers to the fish of India as follows: ‘ Their flesh is
delicate and they have few small bones. They are surprisingly
active. On one occasicn a net was laid in the river from side to side.
Each side of the net was then raised a gaz (23 feet) above the water,
yet many of the fish leaped one after the other a full gaz over the
net and escaped. There are besides in many rivers of Hindustan
small fishes which if they hear a harsh sound or the treading of a
foot instantly leap a gaz or a gaz and a half out of the water.’
There is no doubt that in the description of the flesh a reference
is made to our larger marketable fish such as Ladbeo rohita, Catla
catla, Cirrhina mrigala, Barbus tor, etc. The active species referred
to as leaping is probably Catla catla (Ham. Buch.). Buchanan
Hamilton writes about this fish as follows: ‘It is a very strong
active animal, and often leaps over the seine of the fishermen, on
which account, when fishing for the catla, they usually follow the net
in canoes, and make a noise by shouting and splashing with their
paddles.’! I watched big specimens of cat/a leaping over nets at
Niazbeg near Lahore in 1919. It is a common sight in Bengal
(probably in other parts of India also) to see small fish leaping about
on the surface of the water during a heavy rainfall, probably they
react to the noise or disturbance produced by the falling rain. These
small fish are usually the young of the species enumerated above.
Another reference in Babur’s account runs as follows: ‘ One fish
is the kakeh. On a line with its two ears issue two bones, three
fingers-breadth in length. When caught it shakes these two bones
which return a singular sound, whence they have given this fish its
name of kakeh.’ About the identity of this fish Mr. Ali remarks,
‘The identity of a fish described by Babur in the following terms
isa mystery. Sir Lucas King has made no comment on it, and it is
surprising that such a singular animal, if it exists, should not be
generally known. I can find no reference to anything like it in
Annandale’s paper either.’ I have, however, no hesitation in stat-
ing that this mysterious creature is A7zta rzta (Ham. Buch.), which
1 Day, Fishes of India, p. 553 (1878).
— v
MISCELLANEOUS NOTES ~~ 803
is a common fish at Lahore in winter and is called Ahagga. One
often hears hawkers in the streets selling Navi dé Khagge (Khagga
fish of the Ravi River). Ina collection of drawings by Sir Alexan-
der Burnes made during his expedition to Cabul in 1837-38 and
recently described by me in the Journal of the Asiatic Society of Bengal
I‘ find that this fish is called Aagdé at Hyderabad and there seems
to be no doubt that Kakeh is the same as Kagd. Moreover, the
structure and the habit described by Babur exactly tally with those
of Rita rita. In this species the pectorai fins are provided with very
strong spines which are denticulated at both edges. They corres-
pond to the two bones described by Babur and their length would
depend on the size of the fish examined. These pectoral spines
along with the powerful dorsal spine are used as organs of offence
and defence and it is well known that the jagged spines of some of
our Siluroids (cat-fishes) occasion intense inflammation and some-
times inflict dangerous wounds. During the winter season the
individuals of this species hide themselves among pebbles and rocks
in crevices and lie in such situations more or less in a state of tor-
por. At this season they are usually caught in great numbers by
hand by diving to their places of retreat and holding them, when
caught, in such a position that they are unable to move their spines.
This is accomplished by holding the fish between the fingers in such
a way that all the three spines stand erect. The spines of the cat-
fishes are very much feared and they are broken off as soon as the
fish are caught. ,
At Lahore Aita rita is sometimes called 7y7zkanda, in which
reference is made to the three spines of the fish and in the vernacular
name Khagga | believe reference is made to the jagged condition
of the spine, because the thorny budding leaf of the date-palm tree
is also called Ahagga, at ieast in the Gujranwaia District of the
Punjab. The dialect changes from district to district in the Punjab
and I know that in this part of India the fishermen are often unable
to explain the meanings of the vernacular names of fish.
_ The pectoral spines of Azta rita are provided with a set of very
strong muscles and their proximal ends are specially modified to form
a movable joint with the pectoral girdle.» When the fish vigorously
shakes its pectoral spines, on being annoyed, a peculiar noise is
produced by the movements of the condyles of the spines in the
articular grooves. ‘These facts are closely in accord with Babur’s
description of Kakeh.
The Pulwah of Sind is now called Ailsa tlisha and not Clupea
tlisha. Mr. Ali observes that this species ‘ regularly ascends the
Indus in February and March in enormous swarms for the purpose
of spawning.’ While travelling in the Punjab in 1926 I was informed
by a reliable authority that these fish do not ascend in the Indus to
the distance and in the quantities which are referred to in earlier
records, and he actually feared that this highly-prized fish of Sind
was in the process of abandoning the Indus altogether. Exact
information on this point will be of immense economic importance.
- 1 Hora, Journ. As. Soc. Bengal (n.s.) voi. xxii, p. 123 (1927). .
® Hora, Rec. Ind. Mus., vol. xxii, p. 30 (1921). : a
304 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXX11
Jehangir’s reference to the blind fishes of Andha Nag in Kashmir
is very interesting. I donot knowof blind fish from the freshwaters
of India and it is to be hoped that any one who finds them in Kash-
mir or elsewhere in India will send them to the Indian Museum,
Calcutta, for determination.
Jehangir mentions that, ‘ The best fish in Hindustan is the Rohu
and after that the Barin. Both have scales and in appearance and
shape are like each other. Everyone cannot at once distinguish
between them. The difference in their flesh also is very small but
the connoiseur discovers that the flesh of Rohu is rather more agree-
able of the two.’ Mr. Ali identifies Rohu as Labeo calbasu and
suggests that the other may be the Indian Trout (Barilius bola).
-Rohu is a name which is sometimes indiscriminately applied to the
larger species of carp inhabiting our rivers, these may be Ladeo or
Barbus. Ifind (Hora, 1927, p. 124) that at Peshawar Rohoo is applied
to a species of Ladeo probably dyocheilus (McClell.) and at Hasan
Abdal Rahu is used for a species of Barbus, probably tor. Else-
where in northern India and in the provinces of Bengal, Assam and
Bihar and Orissa Rohz is used for Labeo vohita and to the best of my
knowledge is never used for Labeo calbasu, which on account of its
characteristic black colour is known as AKalabasu in Bengal. In the
Punjab and Sind it is called Dz and Di-hee respectively. Roh (Labeo
rohita) is considered to be a great delicacy and the Emperor Jehangir
in all probability refers to this fish.
The vernacular name arin is new to me andit is only by a process
of elimination that I can identify it as Civrhina mrigala. There are
four scaly fishes in the Indian rivers that are highly prized for the
flavour of their flesh and which grow to a fairly large size e.g.,
Labeo rohita, Catla catla, Cirrhina mrigala and Barbus tor. Caitla
can be readily distinguished by its characteristic upturned mouth.
Connoiseurs always find that the flesh of Barbus tor is much better
than of any other fresh water fish of India and moreover the form
of Barbus tor is not as deep as that of Ladeo rohita and the two can
also be distinguished by the form of their heads. Cirrhina mrigala
is very much like a Ladeo and one is likely to be confused by the
two forms; so it is probable that Barzz refers to Cirrhina mrigala.
It could have no reference to the Indian Trout (Barz/ius bola) which
in its characteristic shape, markings and the position of the fins, etc.,
is quite distinct from any species of Ladeo or the other carp named
above.
There is no doubt that the Mogul Emperors of India (Babur and
Jehangir) were careful students of nature and the descriptions of the
animals that they have left behind them, though meagre, fully testify
to this. I hope students interested in other groups of animals will
further elucidate the facts so ably brought to light by Mr. Salim
A. Ali.
ZOOLOGICAL DEPARTMENT, SUNDER LAL HORA,
UNIVERSITY OF EDINBURGH, D:Sc., F-L:S.,. B.Z.Ss
December 24, 1927. Zoological Survey of India, Calcutta.
MISCELLANEOUS NOTES 805
XIX.—THE EFFECTS OF BEE VENOM
Reprinted trom ‘ The Indian Medical Gazette,’ Vol. LXIT
No. 5, May 1927
Under the abovetitle Dr. A. R. D’Abreau has an article in the
Indian Medical Gazette for November 1926, and as a few very strik-
ing cases have come under my personal observation, I am publish-
ing this note.
The species generally credited with evil effects is Apis mellifica.
L. Brant and Ratzeburg were the first to study bee venom in
1833. Paul Bertin 1865 and Carlet in 1884 also studied it, but it
was not till 1897 that the chemical nature of the poison was carefully
studied by Langer.
This poison when freshly extracted is a transparent acid fluid
with a bitter taste and an aromatic smell. The acid reaction is
believed to be due to formic acid.
Langer’s experiments show ihat the active principle is not an
albuminous but an organic base.
The poison is destroyed, or its activity lessened by the action of
oxidizing agents.
Castellani and Chalmers (2nd edition) write ‘Intravenous injec-
tions into dogs produce convulsions, trismus, nystagmus, emprost-
hotonus, death from respiratory failure. This action is believed by
some authors to be due to a neuro toxin, but the blood after death
is very fluid, and the red corpuscles are destroyed, indicating a
marked hemolysis, while all the organs except the spleen show
heemorrhage and hyperemia, so that the effect may not be due so
much to the action on the nervous system, as to that on the blood.
With regard to the hemolytic action Morgenroth and Capri have
Shown that there is a similarity between this poison and cobra
venom.’
The constituents of this poison are :—
1. An inflammatory poison, said to be derived from the acid
gland.
2. Neurotoxins ; (a2) convulsive, (said to be derived from the
alkaline gland); (6) narcotic (secreted by the acid gland).
3. Heemolysins.
Let us now for a moment study the poison apparatus of a bee.
From the posterior endof the abdominal segment projects a
chitinous sheath which contains two barbed darts which form the
sting. Into this sheath and connected with the darts two (sometimes
three) ducts from glands open. The principal duct belongs to the
acid gland and opens anteriorly into a sac which is a reservoir for
the poison and which Jeads to a slender coiled tubular gland which
may be bifid or even divided into three. Thisis a very long gland
ramifying amongst the contents of the abdomen. The second open-
ing belongsto a small irregular tubular gland called the alkaline
gland, or gland of Dufour.
Now bearing in mind that different trains of symptoms are caused
as the result of stings, I think it is safe to infer that the two darts
can be used independently of one another. When on the other
hand a bee is very angry and attacks with force, both darts are used
806 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXX11
and the full amount of poison possible for ejection is injected. It
should also be remembered (at least this is my idea), that ail the
bees which apparently attack one do not sting, but the majority
generally settle for a second or so and then fly off.
The following cases will I hope illustrate my points :—
Case 1.—In September last an elderly woman in the female ward was
stung by a bee at the external canthus of the left eye one afternoon. The Sub-
Assistant Surgeon who happened to be in the ward at the time reported that the
bee deliberately attacked this patient and would not be driven away until it had
stung her. In about an hour’s time there was a considerable amount of swel-
ling and ecchymosis over the forehead, left temple, left side of the face, and
both eyelids were puffy. At first sight the patient presented the appearance of
erysipelas. There was a great deal of pain, a rise of temperature to 101°F. and
pulse of 112 per minute. By next morning all the physical signs had greatly
diminished and by evening there was no need to consider this part of the
patient’s illness.
Case 2.—A few days after this occurrence, my wife, son and myself were
sitting in the garden, when my son was stung on the back of the neck; but all
that happened in his case was the formation ofa little lump (nodule) anda
slight red blush about 2 inches in circumference, and this all disappeared in a
couple of hours.
This bee did not make a determined attack.
Case 3.—In December 1917 when we were encamped on the Rouvouma
River in Portuguese East Africa, aswarm of bees one morning attacked the
camp, and ‘ M’, a young chemist who had only recently joined us as a dispenser
from Scotland, was most furiously set upon. Some hundreds of bees at a time
settled on his head, face, neck, andhands. He became frantic in his endea-
vours to drive them off and in fact, fright and pain were soon followed by ex-
haustion, and later on by a good deal of collapse.
~ In about 3 or 4 minutes the bees had all flown away, and then we extracted
about 40 or 50 stings from ‘M’ mostly from the back of the neck. The face,
neck and scalp soon became very red and swollen, and later on a slight cedema
was noticed.
This stinging had taken place at 9a.m.and by 2 p.m. ‘M’ had tempera-
ture of 102°F. and complained of a constricted feeling in the head. During the
day he had one or two loose stools, which I attributed to fright as he was a very
nervous man. Next morning he felt a ‘ bitcheap’, the neck still showed a
slight swelling, but by evening this had gone and he felt quite fit.
Judging from the number cf bees that settled on this man, the number of
broken stings we found, his method of knocking them off and, finally, the symp-
toms of poisoning he exhibited, Ido not believe that less than one-eighth or
more of the bees that settled on him, stung him.
I had often been told that if a swarm of bees are around and you happen to
be their line of fight just stand perfectly still and quiet and they will-not attack,
while the moment you start driving them off by swinging your arms about,-the
stinging begins, and this was amply proved in this instance. I was not more
than four yards away from ‘ M ’ when the bees were flying past and I just shout-
ed out to him to keep still and then remained quiet and motionless. Innumer-
able bees came around me and some actually settled on meand though the
temptation to drive them off was great, I did not do so but waited for them to
go off on their own, while ‘M ’ on the other hand completely lost his head, with
the result that the bees soon concentrated all their energies on him.
Case 4.— Another instance occurred one afternoon under the Kosi Bridge, a
few miles out of Kursala in the Bhagalpore District in January 1919. A
party of us were in butts scattered along a wide strip of sand awaiting the re-
turn of demoiselle cranes from the fields. No one had informed us that there
was a huge beehive under the bridge, and‘ K’ of the Police had his butt not
more than 30 yards away from this. As the cranes came sailing past a regular-
feu dé joe started but very soon gave place to most awful yells from‘K’. He
was soon racing across the strip of sand to the river with a thick swarm of bees
about, around, and behind him. Seeing his plight, one of the party to divert
the attention of the bees fired a charge of small shot into them and at the same
time got his gun boy to throw as much sand as possible into the air about 10 or
MISCELLANEOUS NOTES - 807.
15 yards away from him. Ido not know the virtue of this procedure but it
certainly had the desired effect, for almost immediately the bees left him and
flew back to the hive.
Now comes the interesting point. ‘K’ had one sting on the forehead but
from the back of the neck to over the occipital region he must have had fifty or
more (I am writing from memory).
The back of the neck is for some reason a specially selected site for stinging.
It may be that when attacked one ‘ ducks’ one’s head to protect the face and
eyes, but this by no means affords a full explanation.
‘The tsetse fly also selects the back of the neck and the popliteal space.
‘K’ complained of a burning sensation and a certain degree of vertigo. In
a short time the affected area was swollen, red and hot, but ‘ K’ was as hard as
nails and no further symptoms developed.
Port BLairr, A. BAYLEY bE CASTRO,
May 5, 1927. Juntor Medical Officer.
XX.—PLANTS AND INSECTS
Jane Austen in one of her novels released a simple but effective
thought by making one of the characters exclaim: ‘And now you
love a hyacinth, so much the better, you have gained a new
source of enjoyment, and it is as well to have as many holds
upon happiness as possible.’ In other words the observation of
the symbiosis or close relationship between insects and plants and
the place of a plant in its natural classification may sometimes be at
once inferred by the insects who areits guests. Thus a beautiful
golden green shield bug (Chrysocoris) in Southern India is fre-
quently seen on the leaves of Jatropha curchas and in Calcutta a
brilliantly coloured bug of the same genus is found onthe leaves of
Acalypha indica also a plant of the Euphorbiaciz. Sometimes an
interesting phenemenon of insect life is witnessed and related by a
lucky observer in vivid, impressionistic prose leaving his readers
in the dark as to the true identity of the objects described unless
gifted with the powers of observation of a Sherlock Holmes.
Extracts in illustration will be quoted in full from their excep-
tional interest for the guidance of others in the future who have the
opportunity of observing similar phenomena in Indian insect life in
the localities referred to by the writers quoted. Mark Thornhill in
Haunts and Hobbies of an Indian Offictal remarks, ‘ There are in
India many varieties of insects that when they place themselves in
certain positions on plants or shrubs appear to be part of the vege-
tation. Many of them have been described but to realize their
wonder they must be seen—one moment a twig, a leaf, or a flower
the next moment a moth ora grasshopper. The most marvellous of
these transformations that I ever beheld was at the commencement
of the ascent from the valley of the Dunto Mussoorie. The lowest
part of the road is exceedingly bare, hot and dusty. Along its outer
edge there grew a row of extremely sma! ‘shrubs hardly much more
than half a yard in height. The shrubs were always covered with
a profusion of white flowers much resembling ‘immortelles’. In
appearance they seemed as hard and dry as the road itself. I had
often noticed these flowers and one day I enquired of my servant
what was the name of these flowers and where else they grew for I
had never seen any exactly like them before. The man looked sur-
prised at my question and enquired what flowers I referred to, for
808 JOURNAL, BOMBAY NATURAL AIST. SOCIETY, Vol. XXXII
he added there were none there. I pointed to the shrubs. He
smiled, walked to one of them and touched it with his stick.
Instantly the flowers vanished. I saw a tuft of bare twigs, and the
air was filled with a perfect cloud of small white moths. They
flitted about for atime and gradually alighted and the twigs were
covered with white flowers as before.’ No doubt Thornhill when
he wrote these lines fancied what he saw to be a case of protective
insect mimicry but it is a characteristic of some members ofthe
order Rhynchota (which includes the cicadas, lantern-flies, the quaint
little-horned monsters known as Membracids, and other small
moth-like forms) to collect in gregarious patches on the surface of
the epidermis of the plant selected as a host and whose life juice -
they are tapping and the insects seen by Thornhill were not moths
but Lawana conspursa. 'The specific scientific name a ‘ scattering
together’ is very apt from the simultaneous fight of the insects
when suddenly disturbed. The familiar brown mango-hopper has
a similar habit of sudden dispersion when rudely disturbed on
mango leaves. The host plant was probably Plectranthus ternt-
folius or rugosus, a small shrub of the order Labiata common on
base hill sides. The ieafless condition of the twigs indicated the
sapping of the plants by their insect guests. Tragicindeed was the
drama of insect life on a species of wild caper (either Capparis
aphylla or spinosa) witnessed by the eyes of Phil Robinson in the
vicinity of Delhi and graphically described by him in the following
words: ‘Everyone has read of the great flights of butterflies that
have astonished travellers. Great flights that cloud the sun as they
pass overhead, that drive their way even against the wind and do
not hesitate at the ocean itself. Right out in the ocean these
flights of suicides are sometimes met with, but whence and whither
they come and go men of science cannot tell. Now at the foot of
the gate-way near the Kutab-Minar, elevenmiles from Delhi grows
in profusion a thorny shrub, the same that amuses the camels and
goats among the ruins of the Futtehpur Sikri Hill, and on the parti-
cular morning we were there the whole thicket was alive with white
butterflies. I went down to examine it, and such a sight I never
saw before. Each bush was alive with the insect in every stage,
the chrysalides were so thick in places that they touched each other,
and while I watched they hatched by scores and hundreds. I picked
a spray of them, and in half an hour the wings were strong and the
butterflies were own. Inmy hands there remained a twig witha
dozen empty shells. This multiplication of life was simply awful.
Every other leaf had a caterpillar feeding on it, and among the cater-
pillars the mothers of the next generation were busy laying their
eggs. The bushes were literally alive. And nota butterfly left the
place of its birth. As soon as its wings were strong enough to
carry it, it hovered about the shell it had just left, and in a minute
or two found its mate and thus all on one inch of a twig the life was
lived from egg to caterpillar, from caterpillar to chrysalid, and from
chrysalid to butterfly. Death was all prepared for them by the side
of their cradles. Noticing that in very many cases the butterflies
did not move when I came near I picked up one with my hand from
the spot on whichit seemed to be resting, and then I discovered
MISCELLANEOUS NOTES 809
the horrible secret of its apathy. A spider was holding it to the
branch—a beautiful pale green spider—and my eyes opened. I soon
became aware of the multitudes of the same kind that swarmed
among the butterflies. _I waved my hand over a bush. A hundred
butterflies might rise from a branch but a hundred would remain
motionless. A spider had hold of each. To give you an idea of
the enormous creation and destruction that was going on in this
patch of scrub that morning I saw what I mistook for a birds’ nest,
a globular mass of web pendant from atwig. On taking it off I
found it was only the web of acommon spider and that it weighed
as nearly as possible half a pound. It was all dust and spider’s web
and dead butterflies. And you may calculate if you like how many
dead and sucked-up butterflies go to make up half a pound. Again
in a corner just as you have seen dried leaves drifted together by
the wind, I saw and sifted with my hands a thick drift of butter-
flies’ wings. ‘There were the remains of tens of thousands.’ The
butterfly observed by Robinson was Anapheis mesentina or a closely
allied pierid. In Western India it is partial to fluttering among the
showy white and canary yellow flowers of Capparts zeylanica, a
habit of which the tree lizard, popularly known as a blood-sucker, is
aware. This lizard Calotes versicolor lies in wait on the leafy twigs
of the plant on the look-out for unwary insects. The lower surface
of the hind wings of Anapheis are sometimes of a yellow tinge.
It is perhaps noteworthy that the two lower petals of Cappfaris
zeylanica flowers are also yellow and this blending of colours in
flower and butterfly must be of a protective nature to an insect
poised with closed wings onthe corolla of a flower. The green
crab-spiders of the genus Thamisus are perhaps far more deadly
enemies to lepidopterine life than lizards or birds. From their
cryptic colouring they are difficult to discern either on or in the
axils of leaves and sometimes mixed up with the bracts or involucre
surrounding a cluster of flowers. On one occasion a beautiful day-
flying humming bird moth (Cephanodes hylas) was observed hanging
by its long haustellum (proboscis) from the leaf bracts surrrounding
the flower head of a common garden Zinnea. Closer examination
revealed the anterior end of the moth’s proboscis in the clutches of
a green crab-spider snugly ensconced within the flowery receptacle.
Humming bird moths are fond of hovering over the blue flowers of
Duranta plumieri and it was decidedly unusual for one to have
fallen a victim at the shrine of a Zinnea. There is also a pale sickly
white form of the crab-spider just referred to whose get-up must
assimilate with the leaves of the white floretted Zinnea and thus
render it just as deadly as the green form just referred to. These
Spiders are in fact good instances of what is known as ‘ aggressive
protective mimicry.’ Their colouring likewise protects them from
the attacks of their deadly enemies the graceful restless wasps of
the family Pompilide who are perpetually on the look-out for
spiders and penetrate the inmost recesses of plants in eager quest
for their juicy fat-bodied victims.
Y,M.C.A., H. JOUGUET.
WODEHOUSE ROAp.
25
810 JOURNAL, BOMBAY NATURAL HIS. SOCIETY, Vol. XXXII
XXI.—WEEDS OF THE INDIAN WAYSIDE
To most minds the vegetable organism known as a weed is like
the scientific definition of dirt—‘ matter in the wrong place’ when it
obtains a footing in some garden and then over-runs the surround-
ing country, which it may or may not beautify according to a human
standard of esthetics. Butitis in the nature of a weed to grow
where it is not wanted. Each of the three great sea-port towns of
India has imported its own particular weed. In the swampy
country of Calcutta, Assam, and Eastern Bengal it is the lovely
flowered water-hyacinth (Zicharnia crasstpies), that has obtained a
firm footing, in Madras Gamphrena decumbens is dominant, and in
Bombay a species of mallow (Malachra rotundifolia). The first
named, the water hyacinth, has been met with in Madras growing in
Sparse patches in pools of stagnant water in the Spur Tank ard at Per-
ambur near the large local workshops of that town. On one occasion
it choked up the waters of a small tank in the suburb of Royapettah.
The water hyacinth grows profusely along the banks of the beauti-
ful Ulsoor Lake in Bangalore where however it is being eradicated,
and as regards Bombay it has been seen flourishing in the waters of
an old tank on the right hand side of the road from Mora to Uran
also in another tank behind the southern slope of the lesser
Karanja Hill across the harbour of Bombay. On the whole it does
best in Bengal although itis one of those weeds which would spread
anywhere where there is a sufficiency of shallow water to enable it
to thrive and propagate. So far as is known the only insects that
visit the thick clustering flowers are flies'and small bees. Butterflies
however sometimes visit the blue florets of Monocharia hastefolia
but none have so far been seen settling on the far more conspicuous
flowers of the water hyacinth. Florets or minute flowers are more
to the taste of butterflies, and it must be assumed in the case of the
water hyacinth that fertilization is effected through the agency of
flies and small tees. Aithough so frequently referred to in the
papers as the pest known as the lilac devil and in measures proposed
for its eradication on the waters of districts where it has become a
rampant weed, there is strange to say no descriptive account of the
plant in any of the older hand-books on Indian botany, a sure indica-
tion of its recent introduction into India by some enthusiastic horti-
culturist who imported the plant on account of the beauty of its
flowers (and in Calcutta the water hyacinth is sometimes sold as an
orchid by wily Indian plant vendors to those who are not acquainted
with its history !) Once seen it is an easy plant to identify. Great
rosettes of leaves, growing thickly in shallow water shaped like the
heads of rampant green cobras from which is thrown out a flower-
ing stem of about a foot in height enveloped in light lilac blue
flowers each flower bearing on the inner side of one of the petals a
pollen-yellow hieroglyphic in shape not unlike the head of a spear,
can be none other but the famous water hyacinth. The buoyant
freely floating bladder-like central root-stalk of the plant fixes itself
to the muddy bottom of a tank by numerous hair-like roots which
when dried look not unlike dense and clotted masses of course human
hair intermixed with the remains of this weed when uprooted and
MISCELLANEOUS NOTES 81i
carelessly thrown on the bank of a tank whose surface is being
cleared of the pest. It does not appear so far to have been recorded
from Northern India.
On alighting at the railway Central Station at Madras and taking
a walk in the grounds of the adjoining Peonles’ Park, it is at
once noticed that large spaces of waste ground are overrun by an
insignificant weed of about from 6 inches to a foot in height. The
leaves of the piant are of a narrow elliptical contour and of a downy
or cottony whiteness on the lower surface, the inflorescence being
a terminal white cone-shaped tuft of minute florets. This is
Gomphrena decumbens, a member of the same genus as the showy
purplish-red flowering G. globosa, the globe amaranth of gardens.
It is pretty evident that the former must have been accidentally
introduced into India from its native land Mexico. It is said that
the little weed came to India in a round about way from Australia
through being included in the fodder grasses loaded on boats con-
veying horses from some Australian port to India and that it has
only established itself at Madras within the last forty years. Nobody
seems to know much about it and it is undoubtedly of recent intro-
duction as it has not yet found its way into the pages of Indian
botanical handbooks. It occurs but sparingly in Bangalore, where
on the other hand another South American weed may be noticed
named Alternanthera echinata. Creeping along some of the road-
sides with leaves of a rounded oval shape and throwing out a spiny
inflorescence between the axils which gives it its specific hedge-hog
name. De Candolle pithily includes it in his Prodromus under a
list of species ‘non satisnotez’, but one would like to know some-
thing about a plant whose native home is the Argentine Republic
that is found straggling about roadsides in Bangalore and Madras.
It is of course undoubtedly of accidental introduction having been
shipped along with more showy consignments of foreign plants.
Malachra rotundifolia is presumably the large rough-leaved
mallow bearing conspicuous yellow flowers growing in dense gre-
- garious patches on low waste Jands in and around Bombay. Some-
times white flowers are seen amongst those of normal yellow but
the latter it may be added are invariably white inside the base of the
corolla of the flower which probably in an unsatisfactory way
accounts for the tendency towards albinism in some of the flowers
of the plant; Nairne in his flowering plants of Western India
gives the impression that the flowers grow in dense heads. It is
true that the involucre of three reniform leaves surround a capi-
tulum of five or six sessile flower heads, but of these only one or
at most two burst into bloom within the same involucre. In favour-
able situations this plant if not too crowded will attain a height
of four feet, but if growing closely together in dense patches
it seldom exceeds a stature of more than two feet. The flowers are
visited by butterflies and blue carpenter bees, of the former the
species noticed were the Suiphur Yellow, Zerzas hecabe, small blues
(family Lyczenida), the cosmopolitan Painted Lady (Vanessa
cardut), the green Swallow Tail (Papilio agamemnon), and examples
of Papilio aristolochie were observed flying over waste lands where
Malachra rotundifolia was growing profusely.
812 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXI1
The weedy shrubs known as Lanitanas (family Verbenacee), now
growing wild in most parts of India are also of South American
origin. They are easily recognizable from their small white, pink,
red and orange coloured florets which grow in dense clusters and
which are so attractive to butterflies. There was a spot on the
Cooum River at Madras where its banks were over-run with a white
and pink flowering Lantana camara and on fine mornings these
shrubs were visited by scores.of fine Trojan butterflies (Papzlio
hector). It was noticed that these insects were flying in a due
north to south direction during the months of September and
October and some assisted by a favourable following wind must
have travelled a good way following the contour of the east coast.
The larvee of this butterfly have been procured at Sivachellum, one
of the high peaks overlooking Waltair about 400 miles to the north
of Madras. When crossing over from Ceylon to India, this butterfly
was extremely common in the adjacent country beyond Dhunoshkodi
and Paumbaum. It is also a common Ceylon butterfly. In
Bombay the common Lantana is the orange and yellow flowering
variety which does not attract as many butterflies as the white and
pink flowering form at Madras. Have then butterflies a colour
preference sense? It is hard to say. Fritz Muller noticed in South
America a Lantana with florets of changeable colour which was
attractive to butterflies. He observed that each kind of butterfly
was very partial to a particular colour shade in the florets which
changed from yellow to orange and from orange to purple in all
three stages of colouring. Ofcourse it must be borne in mind that
he observed insects and plants intheir original naturalhome. In
India the Lantana has been introduced and it can only be asserted
with doubtful certainty that butterflies prefer the white and pink
flowering form.
j H. JOUGUET.
VMC.As
WODEHOUSE ROAD.
PROCEEDINGS
ANNUAL GENERAL MEETING
The Annual Generai Meeting of the members of the Bombay Natural His-
tory Society was held on Wednesday, February 29, at6p.m., in the Prince
of Wales Museum with Mr. H. A. W. Brent in the chair.
The Honorary Secretary announced thirty-one new members since the last
meeting.
~The appointment of officers for the ensuing year was as follows :—
President: H.E. The Right Hon’ble Lt.-Col. Sir Leslie Wilson, P.C.,
Geers. M.G. 1955.0:
Vice-Presidents:; H.H. The Maharao of Cutch, G.C.S.I., G.C.I.E. ;
Revd: ©. Blatter, S.J., Ph. D., F.L.S:, and the Hon’ble Mr. J. E: B. Hotson,
Coel-O.B. ,,.:Cis:
Executive Committee—Bombay: Mr.R. D. Bell, C.1.E., 1.C.S.; Mr. A.M.
Clarke; Sir Currimbhoy Ebrahim, Bart.; Mr. Alwyn Ezra, - F.R.G.S.,
PRE Gls hoes, Lot. V.N. plate, Bsc. > Lt.-Colo Far. Mackie, 1.M.S. ;
Mr. P. M.D. Sanderson, F.Z.S.; Mr. A. Todd; Mr. J. B. Greaves (Honorary
Treasurer) and Sir Reginald A. Spence, Kt. (Hfonorary Secretary).
Advisory Committee—Moffusil.—Mr. T. Bainbrigge Fletcher, F.E.S.
(Pusa); Mr. TY. R. Bell, C.1.E,, 1-F.S. (Retd.) (Sarwar); Col, W- H. Evans,
C.r. Bi, kh. (Peshawar); Lt.-Col: F.C, Fraser, M.S. (Waltair) ; Dr F. H.
Gravely, D. Sc. (Madras); Mr. C. M. Inglis, M.B.O.U., C.M.Z.S. (Darjee-
lino) Lt.-Col. C,, MH. Stockley,.D.S.0., M.C., O:B.E. (Jhansi) ; Lt.-Col. RvB.
Seymour-Sewell, I.M.S. (Calcutta).
THE HONORARY TREASURER’S REPORT
- The Honorary Treasurer, Mr. J. B. Greaves, presented the audited accounts
for the past year for the approval of members andin a brief explanation
stated :—
‘You will notice that this year in addition to the usual statement of receipts
and payments, the auditors have prepared a Balance Sheet showing the liabi-
lities and assets of the Society as on December 31, 1927.
With your permission I should like to take the Balance Sheet first.
peas first of all with the assets side of the Balance Sheet, Furniture stands
at Rs. 2,500, a conservative figure when one takes into account that this figure
includes the show cases containing the reference collection and the book cases
containing the library.
Game Books Account.—The Game Books have been separated from the
ordinary publications of the Society, because the Society in this case is an equal
partner with the author with regard to any profit which may be made on the
sale of the Game Books,
During the year under review Rs. 2,000 worth of Game Books have been sold
and this year we hope to do better. We have, as you will observe, created a
reserve equal to the balance of the total stock. There is however still a sum
of Rs. 5,310 due on account of Game Books. Notwithstanding this if the sales
continue at their present rate it is to be hoped that a large portion of this
reserve will ultimately be unnecessary.
Other publications.—The figure of Rs. 400 represents only those publications
which have been in stock for less than two years. The remaining publications
have all been written off although small sums of money continue to come in
from these sources.
Expenses on preparation of Bird Charts.—As you are aware, the Society
proposes during the current year to publish a series of Bird Charts and Books
and the figure of Rs. 697-13-0 represents preliminary expenses in connection
with the preparation of these publications.
San ces to staff and on account of Members.—Rs, 78-7-3, A nominal
amount,
814 JOURNAL, BOMBAY NATURAL HIST. SOCIETY, Vol. XXXIi
Investments. —Stand at the very satisfactory figure of Rs. 78,362-8-0 and this
figure is considerably below today’s market value.
Cash Balanéces.—A satisfactory figure of Rs. 19,156-12-8.
This concludes the asset side of our Balance Sheet making our total assets
RS.) 01495-8-117
You will notice that we have not taken into account our very complete
Research collection, or our stock of old Journals, 01 our very valuable library.
Liabilities.—Consist of Printing charges. Ks 6,000 is due on account of the
current year’s Journal and will be paid early in 1928. At the beginning of 1927
we paid Rs. 2907/11 on account of Journals publisned ‘n 1926. ‘The remaining
liabilities are all on account of publications.
Donations for Specific Objects Unexpended.—This is a sum which the
Society is holding as a trustee.
Life (embership Fees.—Rs. 42,600 which we have not spent and which
represents the whole of the capital sum paid by life members who are still alive,
and alsothe.... . of institutions, clubs, etc., compounded subscriptions.
We are left with surplus assets of Ks. 38,116-14-4.
I think you will agree that the Balance Sheet shows that the Society is in a
very sound position.
Dealing now with the receipts and expenditure account. This year we have
split this up under separate headings.
General Account.—In this has been included all the main activities of the
Society in connection with the Journal, etc. As you will notice we have finished
up the year with a statisfactory surplus on this head of Rs. 1,663-7-3.
Publication Account.— Here we have a smail deficit of Rs. sue 8-10.
Special Expenditure in connection with Prince of Wales Museum and Nature
Study.— The whole of the expenditure under tnis head has been met by special
donations including during the year, very handsome donations of Rs. 5,000 each
from H. H. The Maharaja of Jodhpur and Sir David Ezra, Kt., towards show
cases, models, etc., and we finish up the year with a satisfactory surplus of
Rs. 7,968-10-7, the sum shown in the balance sheet as being held in trust by
the Society.
Cash Account.—This account is merely a summary of the other account with
the addition of Entrance fees and Life Membership Fees received during the
year with approximately the same amount invested on the other side, and of
course the summary of our cash position.
In corclusion 1 would mention that the Society now consists of 188 Life
Members, 1,123 Ordinary Members who have paid their subscriptions and 62
Ordinary Members who have not paid their subscriptions, making a total of
1,373 members in all. Wuring the year we enrolled 12 Life Members, | old
member rejoined, and 146 new Ordinary Members were elected. There was a
net increase in membership of 35 during the year as I regret to state we lost 76
members through resignation.’
It was proposed by Chairman and seconded by Honorary Secretary that the
audited accounts of the Society for the past year be approved—Carried
unanimously.
On the motion of Mr. H. A. W. Brent a hearty vote of thanks for his services
was passed to Mr. J. B. Greaves.
=
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BOOKS FOR SALE
©) Journals Bombay Natural History Society ‘erie 3
complete set of 31‘ Volumes; also long runs,
odd Vols. & parts; (2) Hume & Marshall's
Game Birds of India 3 Vols. perfect. Rs.200.
Apply to—Mr. BILLIMORIA,
Marston Road, KARACHI.
FOR SALE
The plowing Volumes of the Society’s Journal, the property of a member
leaving India, are for sale :—
“Wor <1. Nos. 1, 2 & 3.
ONO XE A ANOS, Te 3, 4 & 5,
Mol Vili: No, 3. | | Mol. XXUM. Nos2d, 4,85,
‘Vol. XV. Nos. 3,4 & 5. | Wel, XIV Nog 15 22354 & 5,
Vol OX VE > Nose?) 3:5. 1 Vol. ~XXW- SNos“4;8) 3, 4-85,
Vole XVII> Nos. 2, 3, 4 @ 5. Vol.c XXVIL- No. 5:
Vol. XVII. Nos. 2,3 & 4. | Vol.: XXVIL. Nos..1, 3,4 &5.
Mor RIX Nes 12,3 6557) Vol oRRVIM. Nos, ty 2'6e42
Wole COE! Nos. 1, 2):3,4 & 5) | Voly XIX. “Nos. 12,3 & 4.
Wool: RX SONOS) 13a & 5. Vole KS Nos Ae
C Price Rs. 200 for the lot.
Apply to—THE ENGLISH HEAD CLERK, ;
Collector’s Office, Madura, South India.
AMMUNITION 52848 ">
“ASK FOR CATALOGUE
M. SALEH
Se 337, ABDUL REHMAN St.,
& CO., ~~ Bomaay, 3
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