THE
NAUTILUS
A MONTHLY JOURNAL
DEVOTED TO THE INTERESTS OF
CONCHOLOGISTS
VOL. XX
MAY, 1906, to APRIL, 1OO7.
EDITORS AND PUBLISHERS :
H. A. PILSBRY, Curator of the Department of Mollusca, Academy of Natural Sciences,
PHILADELPHIA.
C. W. JOHNSON, Curator of the Boston Society of Natural History,
BOSTON.
INDEX
TO
THE NAUTILUS, VOL XX.
INDEX TO SUBJECTS, GENERA AND SPECIES.
Abalones and the earthquake 135
Abal'Ones and the penal code of California . . .85
Alabama, some shells of Mississippi and . . 34, 40, 52
Amphigyra Pilsbry, n. gen. ...... 49
Amphigyra alabamensis Pilsbry, n. sp. (PL III, figs. 1, 2). 50
Anodonta imbeeillis Say 69
Anostoma, on the generic name 11
Ashmunella rhyssa Ckll. (PL VIII, figs. 3-5). . . 134
Ashmunella rhyssa edentata 135
Ashmunella rhyssa hyporhyssa. ..... 135
Ashmunella rhyssa townsendi Bartsch (PL VIII, figs.
1, 2) 133
Ashmunella, notes on some New Mexican. . . . 133
Athleta rarispina 131, 142
Basommatophora, two new American genera of . .49
Bifidaria agna Pilsbry & Vanatta (figs. 1, 2, 3) . . 141
Boreocheton 64
Brachypodella, a synonym of 47
British Columbia shells 95
Brooklyn Conchological Club 89
(iii)
IV THE NAUTILUS.
Bulimulus dormani . . 6
Busycon montforti Aldrich, n. sp. (PI. VI) . . . 121
Calliostoma carnicolor Preston, n. sp. (PI. VIII, figs. 5, 6) . 140
Cardium from Puget Sound, a new. .... Ill
Cardium (Cerastoderma) fucanum Ball, n. sp. . . 112
Cayuga Lake, land shells from the east shore of . . 100
Cazenovia, N. Y., mollusks of 97
Chiton marmoreus var. caeruleus Winkley . . .66
Chitons, remarks on certain New England . . .62
Chloritis edwardsi ........ 47
Cleopatra multiliarata Ancey, n. sp. . ; . . 45
Cleopatra smithi Ancey, n. sp 45
Cochliopa rowelli in California 10
Correspondence ........ 142
Cypraea bernardinae Preston, n. sp. (PL VIII, fig. 8) . 139
Cyrena (Polymesoda) panamensis 3
Don ax cayennensis ........
Donax denticulatus ........ 2
Drymaaus dominicus 7, 24
Drymaeus dormani . . . . . . . 7, 24
Drymaeus multilineatus 24
Dyakia, note on 47
Epiphragmophora (Micrarionta) hutsoni Clapp, n. sp.
(PI. IX) . .... 136
Epiphragmophora (Micrarionta) hutsoni, on the soft
anatomy of ........ 138
Epiphragmophora kelletti and E. stearnsiana . . .13
Epiphragmophora stearnsiana Gabb .... 13, 61
Epiphragmophora tudiculata, a healthy colony of (PL
IV) ... ... . 61
Everglades, collecting in the ...... 9
Eurycaelon crassa Hald. ....... 42
Fulgur, from Florida, a new fossil (PL VI) . . . 121
Glabaris Gray, or Patularia Swainson, note on the genus. 39
Glyptostoma newberryanum ... .61
Haliotis californiensis 86
Helix arbustorum in Iceland . . .94
Helix hortensis in Newfoundland . . 94
THE NAUTILUS. V
Helix hortensis on Magdalen Island. . . . . 105
Helix hortensis, on the distribution in North America
of . I 73, 95
Helix jejuna Say, transferred to Praticolella . . .32
Helix subglobosa 73, 77
Insufficiency of an operculum as a basis of classification
in the round-mouth shells . . . .91, 102, 113
Lepyrium, note on . . . . ... .51
Lepyrium showalteri 51
Lymnasa dalli Baker, n. sp 125
Lynmeea danielsi Baker, n. sp. 55
Lynmsea leai Baker, n. sp. . . 126
Lymnaga megasoma Say 36, 83
Manatee snail, Bulimulus dormani . . . .6
Microstoina hanleyanum Jous. . ... . . .47
Mississippi, some shells of Alabama and . . 34, 40, 52
Mitra picta Reeve, note on 57
Mumford collection of shells 119
Nebraska, a list of shells from 81
Neoplanorbis Pilsbry, n. gen 50
Neoplanorbis tantillus Pilsbry, n. sp. (PI. Ill, figs. 3-5). 51
Neritina showalteri 51
Nicaragua, shell collecting on the coast of ... 1
Notes. . . . 9, 24, 36, 46, 71, 83, 94, 105, 118, 144
Oklahoma, mollusks of 16
Omphalotropis macromphala Bavay = 0. quadrasi V.
Moll. . . ..... 47
Oysters carried by seaweed ...... 94
Paravitrea aldriehiana, n. sp. (PI. V, figs. 8-11) . 109, 111
Paravitrea from Alabama, new species of ... 109
Patularia swainson, notes on the genus Glabaris Gray or. 39
Pearls, output of American 9
Pisidium fragillimum Sterki, n. sp. .... 18
Pisidium friersoni Sterki, n. sp. . . . . .20
Pisidium levissimum Sterki, n. sp. . . . . 18
Pisidium minusculum Sterki, n. sp. . . 17
Pisidium neglectum Sterki, n. sp 87
Pisidium neglectum var. ( ?) corpulentum Sterki, n. var. 88
VI THE NAUTILUS.
Pisidium planatum Ancey 46
Pisidium proximum Sterki, n. sp. . 5
Pisidium subrotundum Sterki, n. sp. . 19
Pisidium succineum Sterki, n. sp 99
Pisidium superior Sterki, n. sp 98
Pi tar (Hysteroeoncha) dione 2
Plejona, Bolten 130, 143
Pleurotoma (Genota) carp enteri ana Gabb (PI. II, figs.
1-3) . . . . . .37
Pleurotoma (Genota) cooperi Arnold (PI. II, fig. 10). . 38
Pleurotoma (Genota) riversiana Raymond (PI. II, fig. 9). 39
Pleurotoma (Genota) stearnsiana Raymond (PI. II, figs.
4-6) . ' . 38
Pleurotoma (Genota) tryoniana Gabb (PI. II, figs. 7, 8). 38
Pleurotoma, the West American species of ... 37
Polygyra (Stenotrema) brevipila Clapp, n. sp. (PI. V,
fig. 1-4) ... .110
Polygyra^ ( Stenotrema) altispira P&lsbry (PH. V, figs. 5-7) . Ill
Pomatiopsis Winkleyi Pilsbry 43
Popular conchology 71
Praticolella jejuna Say 33, 46
Pseudopartula, note on . . . , . .47
Psilocochlis Dall, note on the genus 128
Psilocochlis mccalliei Dall 128
Publications received . 11, 36, 47, 71, 84, 96, 106, 119, 143
Quadrula trapezoides Lea. 68
Ravenel 'collection 36
Richfield Springs, New York, shells of . . . .89
Scala berryi Dall, n. sp 127
Scala (Cerostrema) montereyensis Dall, n. sp. . . 128
Scala from California, three new species of . . . 127
Scala lowei Dall, n. sp. 44
Scala rectilaminata Dall, n. sp. 127
Scallops, migrations of 83
Segmentina armigera Say (PI. VII, figs. 1-3). . . 123
Segmentina crassilabris Walker, n. sp. (PI. VII, figs. 4-6). 122
Segmentina wheatleyi Lea (PI. VII, figs. 7-9). . . 123
Sitala, a new Chinese 5
THE NAUTILUS. vii
Sitala hangchowensis Pils., n. sp 5
Sphaerium from Illinois, a new . . . . . 21
Sphaerium hendersoni Sterki, n. sp 69
Sphaerium, on the pathology of 11
Sphaerium solidulum 11
Sphaerium stamineum forbesi Baker, n. var. . . .21
Stenotrema altispira Pilsbry (PI. V, figs. 5-7). . . Ill
Stenotrema brevipila Clapp, n. sp. (PL V, figs. 1-4). . 110
Stenotrema from Alabama, new species of ... 109
Strigilla carnaria 1
Subspecies, mutations and forms . . . 58
Tellina (Eurytellina) angulosa. ... 1
Tivela mactroides 3
Tomogeres Montf 11
Tonicella blaneyi Dall 62, 65
Tonicella ruber var. index Balch, n. var. . . 66
Traehydermon ruber. ... ... 62
Unionidae, some observations on the ova of . 68
Unionidae of Kansas. . . . 144
Valvata bicarinata Lea (PI. I, fig.14) . .29
Valvata bicarinata connectans Walker, n. var. . 30
Valvata bicarinata perdepressa Walker, n. var. (PI. I,
figs. 15-16). . 30
Valvata, notes on ... 25
Valvata lewisii Currier (PI. I, figs. 12-13 . 29
Valvata sincera danielsi Walker, n. var. (PI. I, figs. 10,
11) . ... .28
Valvata sincera Say (PI. I, figs. 1-6) . . . .26
Valvata sincera nylanderi Dall (PI. I, figs. 7-9) . 28
Vitrea approxima and V. vanattai, note on . . 57
Vitrea (Paravitrea) aldrichiana Clapp, n. sp. (PI. V, figs.
8-11) . ... .109
Volutidae, a review of the American . . . 129, 142
Ward, Henry A. (obituary) ... .48
Winkley collection of shells 118
Wisconsin shells, list of ....
Zonitoides alliaria in Colorado. . . 144
THE NAUTILUS.
INDEX TO AUTHORS.
Aldrich, T. H. . . . .121
Ancey, C. F 45, 47
Baker, Frank C 21, 55, 125
Balch, F. N 62
Chadwick, G. H 22
Clapp, Geo. H 13, 105, 109, 136
Cockerell, T. D. A. . 58, 94
Dall, Win. H 39, 44, 111, 127, 128, 142
Ferriss, Jas. H. ..... .16, 133
Fluck, W. H 1
Frierson, L. S. 68
Grataeap, L. P 91, 102, 112
Henderson, Jr., John B .97
Henderson, Junius 144
Hinkley, A. A 34, 40, 52
Johnson, Charles W 46, 73, 95
Kelsey, F. W 61
Latchford, F. R 83
Nason, Win. A. 141
Pilsbry, Henry A. 5, 11, 32, 47, 49, 51, 57, 72, 119, 133, 138, 140
Preston, H. B 139
Raymond, W. J 37
Reynolds, A. G. 9
Rons, Sloman 57
Rowell, Rev. J 10
Simpson, Charles T 24
Smith, Burnett ... 129
Smith, Maxwell 89
Sterki, V 5, 11, 17, 69, 87, 98
Vanatta, E. G 95, 140
Walker, Bryant .... 25, 57, 81, 106, 122
Wheat, Silas C 89, 100
Williamson, Mrs. M. Burton 71, 85
Stearns, R. E. C. . 135
THE NAUTILUS.
VOL. XX. MAY, 1906. No. 1.
SHELL-COLLECTING ON THE MOSQUITO COAST OF NICARAGUA— VI.
BY W. H. FLUCK.
Tellina (Eurytellina] angulosa Gmel. Abundant on the beach all
along the coast from Bluefields to Cape Gracias a, Dios. The dead
shells found were almost always in perfect condition, not even the
hinge ligament injured. Live shells in the shallow water on quiet
days. This is the most highly colored bivalve on the cost.
Tellina ( CycloteUina) fausta Donovan. Plentiful in the shallow
water about the Man of War Keys.
Sanguinolaria (Psammotella) operculata Gmel. A common shell
on the sandy beaches of the Mosquito Coast. Walpa Siksa is a good
locality for it. This very beautiful shell is not as brilliant as T.
angulosa, but is larger.
Sanguinolaria rosea Lam. This beautiful pink shell is by no means
numerous along the east coast of Nicaragua. I searched the beach
and shallow water almost daily for four years, but succeeded in find-
ing four specimens only. It inhabits deep water, possibly. It is
altogether too fragile to stand the pounding of the surf.
Asaphis deflorata Linn. Abundant in the shallow sea and pools
at King's Keys and the other islands along the coast. It seems to be
variable in color, being pink, cream, blue, and white.
StrigiUa carnaria L. Very abundant. I found two sizes ; large,
full-grown forms fairly plentiful, and young, brighter colored half-
sized specimens, in large quantities. They occurred thus so regu-
larly that I was persuaded they were two distinct species. I felt the
more certain of this since in the larger forms the ligament was in
2 THE NAUTILUS.
every case more deeply seated. However, both Mr. C. W. Johnson
and Dr. W. H. Dall assured me that the smaller form was young
and the larger " typical."
Iphigenia braziliensis Lam. My specimens, with the exception
of a few dead shells, were found at or near the mouth of the Wavva
river, where it is plentiful. It is a dweller of estuaries.
Donax denticulatus L. This is by far the most abundant bivalve
on the Caribbean coast of Nicaragua. All the native children,
Indian, Creole, as well as the Spanish speaking ones gather it in
large basketfuls for the family larder. At low tide these beautiful
" wedges " are to be seen all over the beach in favorable places, sit-
ting up in the sand, thin end (posterior) down, with about one-third
of the broad anterior exposed, the shell a little open and the siphons
at work with every rush of the water as the waves come in and roll
out. Donax is either stupid or without fear, for it seldom trys to
escape by digging, on being approached, as do the Solens. The shell
is very variable in color and radiations. I have a series of about 25,
all different. Wounta Haulover.
Donax cayennensis Lam. Inhabits the same localities and the
same sandy stations between tides as the former, but it is not any-
thing like as numerous. It is a larger shell than denticulatus, with a
more angular anterior ridge, and is more uniform in color, not vary-
ing much from pure white, although some specimens tend to bluish,
yellow, or purple. Prinzapolka was the first locality at which I
found this shell, but one always find them wherever D. denticulatus
is to be had. A very good authority wrote me some time ago : " In
going over the Donacidce in the museum, I find no characters to
separate Lamark's Donax cayennensis from the Donax striata L."
I have never seen the latter form. Can these names be synonyms ?
Both D. cayennensis and D. denticulatus are called alii by the native
Indian population. The former they regard as the male and the
latter as the female. Aid means dimple. Is not that more beauti-
ful than wedge f
Pitar (Hysleroconclia) dione L. Very abundant. I shall never
forget finding my first specimen of this beautiful, sulcate, spinose
shell. The Indian boy who was with me further increased the
pleasure of the find by telling me that they were manas, without
number. I soon found that he was not far from the truth. Wounta
Haulover seems an ideal place for this species, the shore and bottom
THE NAUTILUS. 3
being sandy, the sea open with no bars, and the only rocks too far
out to be of any injury to their fragile spines. Wounta Haulover.
Chione cancellata. A variety. Much worn odd valves only.
Wounta Haulover, on the beach.
Cyrena (Polymesoda) panamensis Prime. This is the common
bivalve of all the lagoons along the Caribbean coast of Nicaragua.
It is not always found in the mud, although in the Bluefields lagoon
it thrives in that kind of a station. At Wounta Haulover it lives in
the lagoon back from fche sea, on a sandy bottom, while at Karata it
is to be found on the gravel. I have eaten very good chowder made
of klito, as the Indians call it. The umbones are always, so far as I
have observed, corroded in adult specimens.
Tivela mactroides Born. This pretty little clam is quite common
on the shore and in the shallow water along the open sea. The shell
is solid and looks as though it might stand rougher water than it
usually gets on the Mosquito Coast. Dr. Dall, in his " Synopsis of
the Family Veneridae " is quite right in saying: " This species may
be white, or chestnut-brown or with brown rays on a lighter ground."
I have specimens that alternate with white and brown concentrically.
Echinochama arcinella L. A few dead shells, with one exception
all left valves.
Chama congreyata Conr. Dead shells everywhere on the beach,
especially left valves attached to old Pecten, Area or Chione shells.
Codakia orbicularis L. Plentiful about Man of War Keys ; I
also have a few from Pearl Key.
Area umbonata Lam. Man of War Keys. I found but one speci-
men, although the Indian turtle-hunters, who frequent the Keys, told
me that it was a very common shell at times.
Scapharca ( Cunearca) incongrua, var. braziliana Lam. Wounta
Haulover. Plentiful at all times on the beach, generally with hinge
ligament perfect. Also found alive in the shallow water on quiet
days.
Area (Argina) campechensis Dillwyn. Walpa Siksa, on the sea-
beach, both north and south of the river mouth.
Area occidentalis Phil. Bilwi, just north of the Wawa river
mouth, on the open beach.
Melina ephippium L. Man of War Keys. Abundant.
Pinna muricata L. I never found a living shell, but good speci-
mens of dead ones were to be found at almost any time on the beach,
4 THE NAUTILUS.
especially about the sand-bars and in the coves at and near the mouth
of rivers, as at Wawa, Wounta, and Wai pa Siksa.
Pecten dislocatus Say. Wounta Haulover. A few single valves.
Plicatula ramosa Lam. Wounta Haulover. Found attached to
soft corals, sponges, and like marine, growth. Numerous.
Ostrea virginica Gmel. A very abundant shell in all the lagoons
and estuaries, not far from the sea. It seems to multiply very fast,
filling up harbors and channels. It does not get as large as our
northern oysters, possibly on account of the large quantities of fresh
water that cover them for protracted periods during the rainy season.
Discinisca antitlarum D'Orb. Wounta Haulover. I found six or
eight on an old Pinna. These little brachiopods remind one of
miniature Anomia simplex.
Helicina amoena Pfr. In native clearings in the forest along the
Wounta (or Kukallaya) river, northwest of the village of Kukallaya.
Helicina dalli Bartsch. Near Wani, Nicaragua.
Nephronaias flucki Bartsch. From the Wounta river, a day's
paddling by canoe to the northwest of the village of Kukallaya.
Streptostyla ftucki Bartsch. Near Wani.
Neocyclotus ^Neocyclotus) chrysacme Bartsch. Near Wani.
Chrondropoma callipeplum Bartsch. Near Wani.
The region about Wani is quite different from the low, sandy,
coastal-plain. It takes many days' canoe paddling, according to the
volume of water in the river, to reach it. It is in what the Indians
call the " hill country."
The five last-named shells are new. Dr. Paul Bartsch, who has
seen my entire collection, is at this writing publishing descriptions of
them.
I have a few more Mosquito Coast shells which I might add to
this list, but will not do so now. At some future time I shall ask for
space for another article.
I am convinced that the shells I collected in the lowlands of east-
ern Nicaragua do not by any means exhaust the molluscan fauna of
that region. The highlands of the interior, the mangrove swamps,
rivers, lagoons, sea beaches, and the keys, all have treasures for the
man who has time to search for them. As for myself, I picked up
only such shells as I stumbled over in the strenuous pursuit of other
duties.
THE NAUTILUS.
A NEW CHINESE SITALA.
BY H. A. PILSBKT.
Sitala hangchoivensis n. sp. Shell subperforate, trochiform with
rounded periphery, pale brown, surface dull, smoothish, under the
compound microscope showing dense, vertical striation, most distinct
on the penultimate whorl, and very fine, close spiral impressed lines,
the base distinctly striate spirally, with finer spirals in the intervals
between coarser lines. Spire conic with nearly straight outlines.
"Whorls 6, decidedly convex, the last rounded peripherally and convex
beneath. Aperture somewhat oblique, lunate, the margins thin and
simple ; columella concave below, vertical with reflexed edge above.
Alt. 3.7, diam. 4.1 mm.
Hangchow, China. Types no. 91131 A. N. S. P., from no. 1074
of Mr. Hirase's collection.
It is not easy to decide whether this species should be considered
a Sitala or a Kaliella, but on account of the fine spiral striation it
probably accords best with the first. It is larger than Kaliella de-
pressa and K. monticola Mlldff. with fewer whorls, 6 instead of 6^,
and it has a dull, not glossy surface.
NEW SPECIES OF PISIDIUM.
BY V. STERKI.
PISIDIUM PROXIMUM, n. sp. Mussel of medium size, moderately
to rather well inflated, outlines oval without any projecting angles,
posterior margin slightly subtruncate in some specimens, beaks
somewhat posterior, rather broad to narrow, rounded, or their cen-
ters more or less flattened, with a ridge around the flattened area in
some specimens ; surface with irregular, fine to rather coarse striae
and several lines of growth, shining, color light yellowish-horn to
brownish ; shell opaque to subtransluceut, thin ; hinge slight, cardinal
teeth of various shapes ; ligament rather long.
Long. 5.6 alt. 4.8, diam. 3.4 m. (Wash.)
Long. 6 alt. 5 diam. 3.8 m. (B. C.)
Long. 4.1 alt. 3.5, diam. 2.5 m. (B. C.)
Habitat : Washington, British Columbia : Walla Walla sent by
B. Walker, a good number of specimens deep horn to brownish,
THE NAUTILUS.
beaks rather broad and moderately elevated ; " a small pond, Second
Summit, B. C., at alt., 6000 ft." (collected by O. W. Spreadborough,
1905, sent by J. F. Whiteaves), about eighty specimens; somewhat
larger than the Walla Walla form, of light color becoming brownish
over the beaks in older specimens, then with irregular, alternating
zones of lighter and darker ; the beaks are more elevated and in
many specimens narrower. Although of rather different appearance,
the two Pisidia seem to be of the same species. '' Pond at head of
North Fork of Caribou Creek, East of Burton City, West Kootenay,
B. C." (by same); considerably smaller, but seem identical ; the
beaks are full and prominent, the disks rather flat towards the in-
ferior margin.
This Pisidium is under doubt as to its affinities. While apparently
somewhat like some forms of dbditum Hald., it has also resemblance
with a South American Pisidium which I take for boliviense Sturany.
More materials from other places may help answering this question.
Our Pacific Slope Pisidia, and more so those of Central and South
America are still very insufficiently known.
THE MANATEE SNAIL BULIMULUS DORMANT.1
The sooty mould of the orange, Meliola, is one of the most serious
results of white fly infestation of citrus groves, and is an element in
the injury to various plants from aphids and from some of the scale
insects, especially the Lecaniums, mealy-bugs, wax scales, and cot-
tony cushion scale. The fungus is not itself a parasite on the plant,
but a saprophyte, deriving its sustenance from the sweet honey dew
secreted by these insects. The injury to the plant results from the
smothering action of the fungus, the heavy coating of fungal threads
interfering with the healthful action of sunlight on the leaves. The
appearance of the various ornamental and hedge plants is also dis-
figured by the dark fungus. The sooty mould is especially bad fol-
lowing the white fly attack, owing to the large amount of honey dew
secreted by these insects. As the white fly larvae attach themselves
to the under side of the leaves, the honey dew exuded by them falls
to the top side of the leaves beneath, thus affording favorable oppor-
tunity for the growth of the fungus. So constant is the association
1 Press Bulletin No. 59, Florida Agricultural Experiment Station. Depart-
ment of Entomology.
THE NAUTILUS. 7
of the fungus and the white fly that badly infested groves and hedges
may be recognized at some distance by the heavily coated dark foli-
age. The fungus develops on the fruit as well as on the leaves and
stem, and washing becomes necessary, resulting not only in an added
expense, but also in increased danger of decay in shipping. In this
connection the habits of the tree snail, Bulimulus dormant, J are of
the greatest interest. This snail has been found in the orange
groves in Manatee county feeding upon the sooty mould. Just how
long the snail has been present on orange trees in this county it is
impossible to say. It was observed as long as two years ago by Mr.
F. D. Waite, at Palmetto. It seems to have been present in small
numbers in other groves at this time, but attracted no further atten-
tion until the present summer. The snail is now widely distributed
in Manatee county, occuring in many groves on both sides of the
Manatee river. The work of the snails is very characteristic.
When well started they occur in great numbers on the tree, spread-
ing over it from base to top. Its favorite food seems to be the sooty
mould. The fungus is cleaned from the leaves, stems, and fruit.
The leaves thus cleaned have a glossy, shiny appearance as though
free from white fly. The fruit thus cleaned has a better color and
probably ripens earlier. In addition to the fungus, the snail takes
algte and some lichens from the stem and trunk, giving the trunk a
much cleaner and fresher look. The trees that are cleaned stand
out conspicuously from the surrounding trees by their bright foliage
and clean trunks. The snails increase rapidly under favorable con-
ditions. The eggs are probably deposited in protected places about
the trunks of the trees, possibly also about the base of the tree under
leaves and other rubbish. That the snails are capable of doing ef-
fective work, when present in sufficient numbers, has been shown in
numerous groves in Manatee county during the present summer.
Such trees in these groves as are well stocked with the snails have
been thoroughly cleaned, the fruit not requiring washing.
The snails are of medium size, measuring when full grown, three-
fourths to one inch in length. The shells are smooth, white, or
corneous-white, and with about four bands of brown spots. Old
1 This species is now placed in the genus Drymseus. A closely allied species,
D. dominicus Rve. (B. marielimis Poey), is also common throughout southern
Florida, and frequently confounded with dormani. It probably has similar
feeding habits. — EDITORS.
THE NAUTILUS.
shells have often a somewhat corroded surface, the bands becoming
indistinct or absent. When the conditions are unfavorable, either cold
or dry, the snails take refuge in the hollows of the trees or under
leaves accumulated in the forks, or elsewhere, or under sacks at the
base of the trees when these are provided. It thus becomes an easy
matter to transfer them from tree to tree. A few snails placed by
Mr. Wade Harrison in one of his trees in March, increased in such
numbers as to free the tree of sooty mould by mid-summer of the same
year. The snails are known to range with some variation from the
mouth of the St. John's river on the north, to the Caloosahatchie river
on the south. The species is probably native to Florida, as speci-
mens in small numbers were observed about the hummocks and else-
where as long as fifty years ago. Its habit of feeding on the sooty
mould of the orange, however, was not known until within the past two
years.
In view of the fondness of the snails for the injurious sooty mould
fungus, it becomes of first importance to observe their treatment of
the beneficial fungi. Among the fungi parasitic on orange insects
there are four species of inestimable value to orange growers. These
are the brown fungus well known as an effective parasite on the
white fly larvte ; the red-pink fungus also parasitic on the white fly ;
the red fungus, and the gray-headed fungus, both parasitic on the
common scales. The brown fungus so effective in control of the white
fly, as is well-known, throws out spreading hyphas for some distance
around the body of the dead leaves. It seems that the snails occasion,
ally feed to a limited extent on these spreading hyphse, but evidently
not enough to interfere with the spread of the fungus, since this fungus
is doing particularly effective work in the groves in the Manatee region
where the snails occur ; the red-pink fungus is also abundant in the
groves where the snails occur and is untouched by them. The snails
have not been observed to feed on either the red or the gray fungi
parasitic on the common orange scales, and it is probable that they
have no taste for these parasitic fungi. Colonies of the snails are
being started in parts of the State where the white fly injury is severe
and where conditions seem favorable for the growth of the snail. It
is desirable that close attention be given to the habits of the snail, as
well as to means of protecting colonies against unfavorable conditions.
A few sacks thrown around the tree seems to afford a needed protec-
tion against unusual cold. It is probable that sprays can not be used
THE NAUTILUS.
on trees stocked with the snails without injury to the colony, for,
although protected by the shell from the immediate effect, sufficient
spray probably clings to the sooty mould on which they feed to de-
stroy them. The beneficial parasitic fungi and the snails may be
allowed to work together on unsprayed trees. The snail is here
spoken of as the Manatee snail since while recorded as occurring in
other parts of the State it was found working on the orange groves
first in the Manatee region.
NOTES.
Collecting in the Everglades. I have recently returned from a col-
lecting trip in the Everglades. Most of the collecting was done at a
point where the Big Cypress Swamp and the Everglades meet, about
80 miles from Myers and 35 miles from Jmmokalee P. O. Drymaeus
dominicus Reeve was found in the air-plants. Dead shells of Glandina
truncata var. minor, were plentiful, but live ones scarce. A few
Zonitoides arboreus were also found in air-plants. Polyyyra cereolus
var. carpenteriana and uvulifera I did not collect. Beautiful speci-
mens of the glossy, dark, amber-colored Physa cubensis Pfr., were
found on the under side of the leaves of water plants. Planorbis
intercalaris Pils., and P. tumidus were very plentiful, the latter in
the more shallow water. Ancylus peninsulae Pils. & Johns., was
found on decayed wood in the Cypress Swamp. Atnpullarin depressa
is abundant on the edge of the Everglades. The shells are heaped
around isolated cypress trees, where they are dropped by the
hawks which feed upon them ; a bright yellow variety without bands
is occasionally found. A small Utiio was common in a stream near
Immokalee. A. G. REYNOLDS.
OUTPUT OF AMERICAN PEARLS — Dealers in American pearls
state that the past season showed a diminished output of fine gems,
but a gain in the traffic in baroques or pearl formations of irregular
shapes.
The most valuable pearl discovered last season was taken from the
"Wabash river, near Mount Carmel, 111. This was a pink-white gem
of eighty grains, and is valued by its owner at $8000.
A pearl valued at $5000 was found in a mussel shell taken from
the Skillet Fork river, Carmi, 111. Other pearls, worth from $500
to $3000, were found last year ; but such fortunate discoveries were
considerably fewer in number than in former years.
10 THE NAUTILUS.
Nearly every year witnesses a shifting of the pearl-hunting indus-
try. The Kankakee river, Indiana, was the newest field last sea-
son, and some valuable gems were found there.
Of late, French buyers have been actively represented in the
American market. The activity of these foreign buyers, together
with the diminished supply, was largely responsible for the fact that
prices at the close of the season ranged from 25 to 35 per cent, higher
than they did a year before.
Despite the fact that American pearls now command a higher
price than ever, the volume of business has shrunk because of the
destruction of so many of the mussel beds.
In one year the output of the rivers of Arkansas was valued at
$2,000,000. Last year, some experienced dealers assert, the value
of the whole American product, exclusive of baroques, was not worth
more than $500,000.
Dealers charge that many fine pearls are rendered valueless by
the practice of button houses, and those who fish for them, of throwing
mussels into hot-water vats, in order to open great quantities of shells
rapidly. These button houses seek the mother-of-pearl in the shells.
Baroques, or pearls of irregular shapes, have attained a prominent
place in commerce. Some of them have a sheen or opalescence
equal to the quality of the finest gems.
Until recently, pearl hunters sought only symmetrical shapes, and
the baroques were left as playthings for children along the streams,
or were cast aside.
The influence of the " new art " originated a demand for these
angular, irregular and eccentric formations.
Jewelers now use the baroques in making scarf pins, chrysanthe-
mums, figures and ornaments.
The long and narrow shapes, especially, lend themselves with
readiness to the jeweler's art. As many of these odd formations
were found to have attractive color, lustre or iridescence, the public
soon found a liking for them. — North American.
COCHLIOPA ROAVELLI IN CALIFORNIA. — In response to an inquiry
in the December NAUTILUS concerning the habitat of this species
a single note has been received, referring to the original finding of the
shell. It has apparently not been found since. The note follows :
" Cochliopa Rowelli, was named from shells collected by me, near
Baulinas Bay (not Clear Lake) Marin Co., California." — J. Rowell.
THE NAUTILUS. 11
ON THE GENERIC NAME ANOSTOMA — At the time I prepared a
monograph of this genus (Man. of Conch., XIV, p. 109, 1901) I did
not notice that two modifications of the same name were already in
use : Anostomus Klein in Walbaum, Pet. Arted., Gen. Pise., Ill, p.
659, 1792, and Anastomus Bonnaterre, Encycl. Meth., Ornith., p.
xciii, 1790. Whether these conflict with the molluscan Anostoma
depends upon whether the different spelling be held to constitute a
different name — a question still in dispute. Tomogeres Montf. is the
next name applied to the molluscan Anostoma. — H. A. P.
ON THE PATHOLOGY OF SPH^ERIUM. — Among thousands of
Sphcerium solidulum Pr., in company with several other species, from
Des Moines, Iowa, collected and sent for examination by Mr. T. van
Hyning, there were several dozen specimens of special interest, ab-
normal and evidently of a pathological nature. They were consid-
erably smaller than the average, almost globular, the beaks being
nearly as broad as in the normal form — apparently representing a
distinct species. At the anterior margin of the shell there was a
more or less marked scar, contiguous to the anterior adductor muscle
and the lateral teeth. This was evidently caused in all specimens
by one and the same agency, probably a parasite affecting the mantle
edges in some way and thus causing the deformity of the shell. And
the effect was not only local, as is evident by the fact that a coarse
line of growth, from that place, all around both valves is noticeable
in nearly all specimens thus affected, and the normal growth of the
whole mussel was checked. Some larger specimens, of more or less
irregular growth, show the same scar at the same place, in some in-
stances only slightly marked.
Similar mussels have been noticed before, from other places, but
never in such numbers. Also some Pisidia were seen similarly af-
fected. It is hoped that fresh, living specimens will be obtained, in
order to ascertain the cause of the deformity V. STERKI.
PUBLICATIONS RECEIVED.
CATALOGO DE LOS MOLUSCOS TESTACEOS DE LAS ISLAS FILIPI-
NAS, JOLO Y MARIANAS. 1, MOLUSCOS MAKINGS, por J. G. Hidalgo,
1905, xvi + 408 pp. Since 1840, when the description of Cuming's
rich collection was begun, the Philippines have been noted for the
great number and beauty of their marine shells. A very large num-
12 THE NAUTILUS.
ber of species was first described from there, and in critical researches
on other Indo-Pacific faunas, constant reference to and comparison
with Philippine topotypes is necessary. Several years ago Elera
published a catalogue of the fauna, compiled from previous mono-
graphs, etc., but without new material or critical knowledge of the
mollusks themselves. Dr. Hidalgo has given many years to the
study of the Philippine fauna and has condensed his results in the
catalogue before us. 3121 species are admitted, and 704 others,
cited from the Philippines on insufficient grounds, are enumer-
ated. Those species actually seen by the author are indicated by a
*, and it will be noticed that a large majority are so marked. This
gives the records an authenticity far beyond that of a mere compila-
tion, and is a feature of special value since many of the former
Philippine citations rested upon Cumingian records only. A figure
of each species is cited. This is not only a convenience, but of great
value as showing the exact form intended, a significant point in these
days of nomenclature changes. In his views of specific limits, Hi-
dalgo takes the middle course, conservative as Crosse and von
Martens may be said to have been, but not reactionary as Tryon was
in the earlier volumes of his Manual. The use of genera is open to
some criticism, from the retention of names such as Triton, Vertagus,
etc., now generally held to be untenable. The localities of each
form are fully recorded, most of these records being new.
The typography of the volume is excellent. A portrait of the
author is given as frontispiece.
Dr. Hidalgo proposes to give in the second part of the Catalogo,
an enumeration of the land and fresh water mollusks. These cata-
logues do not replace the more extensive monograghic account of the
fauna which is appearing at intervals in the handsomely illustrated
Obras Malacologicas of the same author. The elaborate design of
the latter renders its publication a work of many years. Meantime
those interested in Indo-Pacific faunas will find the Catalogo one of
the most useful of the many scholarly works we owe to the disting-
uished Professor of Malacology of the Madrid Museum. H. A. P.
CATALOGUE OF THE LAND- AND FRESH-WATER MOLLUSCA OF
TAIWAN (FORMOSA). — By Henry A. Pilsbry and Y. Hirase (Proc.
A. N. S., Phila., 1905). In the island of Formosa 128 species of
land shells are now known, 28 of them described in this paper.
There are 26 species of fresh water mollusks. The affinities of the
mollusks are chiefly with those of China, though there is some affinity
to the Ryukyuan fauna.
THE NAUTILUS.
VOL. XX. JUNE, 1906. No. 2.
EPIPHRAGMOPHORA KELLETTII (FBS.) AND E. STEARNSIANA (GABB).
BY GEO. H. CLAPP.
Recent writers on American conchology have united these two
species, the latter as a variety of the former, but a careful study of
large series of both shows that Gabb was correct when he identified
and named stearnsiana as a distinct species.
Forbes' original description of Helix kellettii was published in P.
Z. 8., 1850, p. 55, and the type is figured on PI. IX, 2a, b. He
was, however, badly mixed on his localities as he says : " Of the
genus Helix there are nine species. Of these H. toivnsendiana,
nuttaHiana and colnmbiana are certainly from the neighborhood of
the Columbia river. Helix kettettii and pandorse, botli new, are prob-
ably from the same country, though the box in which they were con-
tained was marked ' Santa Barbara.' Helix areolata bears no indi-
cation of its locality."
Through the kindly offices of John Ponsonby, Esq., of London,
England, I was enabled to enlist the services of Edgar A. Smith,
Curator of Conchology, British Museum, and he sent me a colored
drawing of the type, which agrees very closely with Forbes' figure,
and at the same time he checked up Forbes' measurements and
found them correct, but was unable to count 6 whorls, unless any-
thing over 5 is called 6.
"With this as a starting point and a large series of shells from
Santa Catalina Is., and the mainland from San Diego to Kosario
and Cedros (Cerros) Is., Lower California. I have tried to sepa-
rate the two species with the following result:
14 THE NAUTILUS.
In looking up the literature, I found that there was considerable
diversity of opinion and that authors were not always careful to
follow the original description. Binney in Terr. Moll. IV, 17,
copied Forbes' description and remarks verbatim and copied Forbes'
fig. 2b on his PI. LXXVI, fig. 12.
Pfeiflfer in Mon. Hel. Viv. Ill, 182 changed Forbes' description
which gave the size as 22 x 19 x 19 m.m., whorls 6, to size 24 x 20 x
16 m.m., whorls 5.
Tryon in Mon. Terr. Moll. U. S., p. GG (Am. Jour. Conch. II, 317,
PI. VI, fig. 1.) translates Forbes' description, but his fig. 1, PI. VI
looks as if it might have been drawn from a specimen of E. arrosa
marinensis.
Binney in Terr. Moll. V, 361 translates Forbes' description rer-
batim, but figures a large depressed shell from Santa Catalina Is., fig.
242, and says : — " I am positive that it is correctly referred to kel-
letti." In his remarks he refers to the peculiar sculpture where he
gays: — " There are traces on different parts of each shell of three
different kinds of sculpturing; the wrinkles of growth, revolving
impressed lines, and a series of minute granulations running obli-
quely, sometimes almost perpendicularly, to the incremental wrin-
kles." This is an important observation.
This description and remarks are copied in Binney's Manual, pp.
149_150. There is a typographical error on p. 150 where he says
that Forbes' original figure is copied in Terr. Moll. V; the reference
should be to Vol. IV; PI. LXXVI, fig. 12.
Without attempting to give a full bibliography it is only necessary
to say that Cooper, Gabb, Newcomb, Carpenter, Stearns and Hemp-
hill have all written on the subject in various publications.
Gabb published the description of Helix sfearnswna in Am. Jour.
Conch. Ill, 235, PI. 1G, fig. 1. He gave the number of whorls as
5^, but did not state the size. The type is in the collection of the
Acad. Nat. Sciences of Philadelphia.
Binney and Bland in L. & F.-W. Sh., I, 177, changed Gabbs'
description somewhat, giving the number of whorls as 5 and adding
the size 22x17 x 12 mm. This attitude must be the length of the
axis, as the flattest shell that I have seen is 15^ mm. high, a single
specimen, while the others run from 16 to 21-| mm.
In Terr. Moll. V, 3G2, Binney repeated the description of L. &
F-W. Sh. adding notes on the anatomy and comparing with that of
THE NAUTILUS. 15
kellettii. This is copied verbatim in Manual Amer. Land Shells, p.
151.
In examining a large series of kelleltii from Santa Catalina I found
one shell which agrees exactly with Forbes' original description and
figure, and the later figures sent me by Mr. Smith, in size and color,
but it has 5 whorls instead of G. All of this lot, about 90 shells,
have the peculiar sculpture noted by Binney while none of the
stearnsiana from the mainland of L. Cal. have it, and as Mr. Ponsonby
informs me that this sculpture appears on Forbes' type, I think we
can safely limit kellettii to Santa Catalina Is.
1 2 3
Fig. 1. Embryonic whorls of E. kellettii Fbs. "The isthmus," Santa Catalina I.;
shell 26 mm. diam.
Pipe. 2. E. stearnsiana Gabb, San Diego ; shell 26 mm. diam.
Fig. 3 shows Fig. 1 superposed upon Fig. 2, the latter in stippled line. All
figures X 10.
The sculpture of the embryonic whorls of the two species also in-
dicates that they are distinct, that of kellettii consisting of very weak
radial lines, not waved, so that the whorls appear almost smooth,
while in stearnsiana it is a very closely set series of wavy lines, gen-
erally bifurcate at the suture, giving the surface a granulated appear-
ance under low magnification. The accompanying figures show the
difference in size of the embryonic shells of the two species, figs. 1, 2,
and in fig. 3 the drawings are superimposed to bring out the difference
more distinctly.
A series of 58 kelleltii from Santa Catalina shows a variation in
greater diameter of from 21^ to 27 mm., whorls fVom 5 to 5-|. 26
stearnsiana, mostly from San Diego, vary from 20 to 25^ mm.,
greater diameter, whorls 5 to 6.
Figuring both series into percentages: —
Of 58 kellettii 27 or 4G.5 per cent, have gr. diam. 25 to 26 mm.
16 THE NAUTILUS.
Of 58 " 43 or 74.1 per cent, have gr. diam. 24 mm., and over.
Of 26 stearnsiana 12 or 46.1 per cent, have gr. diam. 23 to 24 mm.
Of 26 " 21 or 80.7 per cent, have gr. diam. 24 mm. and
under.
Of 58 kellettii 56 or 96.5 per cent, have whorls 5 to 5 +.
Of 26 stearnsiana 19 or 73.1 per cent, whorls 5^ to 5|.
A series of 31 kellettii, selected by Hemphill to show variation in
size, shape and color, but not included in above lot, are from 20^ to
31 mm. gr. diam., whorls 4^ to 5^; 23 or 74.2 per cent.; one 24 mm.
and over and 18 or 58.6 per cent, have 5 to 5 + whorls.
Summing up : E. kellettii is the species found on Santa Catalina
Island, having a large embryonic shell with smooth whorls, surface of
all the later whorls faintly granulated ; generally over 24 mm.,
greatest diam.; whorls 5. (On the mainland at Ft. Vincent, Los
Angeles Co., Hemphill found dead shells which agree exactly with
those from Santa Catalina, but could find none alive). E. stearnsiana
is confined to the mainland and the islands of Lower California.
Embryonic shell smaller, sculptured with wavy lines giving it a
granulated appearance, remaining whorls smooth except for lines of
growth and sometimes faint revolving lines ; greater diam. generally
under 24 mm., whorls 5^ to 5|.
There is a wide variation in size, color and shape in both species
and a number of the forms of kellettii have been named by Hemp-
hill. Measurements below show largest, smallest, most elevated and
most depressed of 89 kellettii and 26 stearnsiana in my collection.
E. kellettii, largest, 31 x 26 x 23 mm. whorls 5£.
" smallest, 20^ x 17^ x 16^ mm. whorls — 5.
" elevated, 26 x 23^ x 25 mm. whorls 5^.
" depressed, 26 x 21 x 18 mm. whorls 5.
E. stearnsiana, largest, 25^x 22 x 21^ mm. whorls 6.
" smallest, 20 x 17 x 15£ mm. whorls 5 +.
" elevated, 21 x 20^ x 2<H mm. whorls 5£.
" depressed, 23 x 19^ x 16 mm. whorls 5^.
MOLLUSKS OF OKLAHOMA.
BY JAS. H. FERK1SS.
Few if any shells have been recorded from Oklahoma territory.
No state or territory in the Union has figured so little in concholo-
THE NAUTILUS. 17
gical literature. In 1897 I collected a few hours in Oklahoma City,
obtaining the following species :
Polygyra texasiana (Moric). Planorbis bicarinatus (Say).
Polygyra monodon (Rack.). Planorbis parvus (Say).
Helicodiscus lineatus (Say). Physa sp.
Zonitoides minuscula (Binn.). Lampsilis anodontoides (Lea).
Zonitoides arborea (Say). Lampsilis purpuratus (Lam.).
Zonitoides nitida (Miill.). Lampsilis gracilis (Bar.).
Euconulus fulvus (Miill.). Lampsilis parvus (Lea).
Strobilops amnis (Pils.). Quadrula lachrymosa (Lea).
Pupoides marginatus (Say). Quadrula pustulosus (Lea).
Bifidaria armigera (Say). Tritogonia tuberculata (Bar.).
Bifidaria contracts (Say). Symphynota complanata (Bar.).
Lymnfea probably techella(Hald.). Sphagnum sp.
Planorbis Irivolvis (Say). Pisidium sp.
NEW SPECIES OF PISIDIUM.
BY V. STERKI.
Pis. minusculum, n. sp. Mussel minute, slightly oblique, medium
inflated; superior margin short, moderately curved, bounded by
slightly projecting, rounded angles; supero-anterior and posterior
slopes little curved or straight, posterior end rounded, anterior a
rounded angle situated much below the median longitudinal line,
inferior margin rather well curved; beakes slightly posterior, rather
large, rounded or somewhat flattened, moderately projecting over
the hinge margin; surface with very fine, slight striae, somewhat
shining, color pale horn, shell translucent; hinge rather stout, car-
dinal teeth: the right strongly curved, its posterior part much
thicker and grooved, left anterior rather short, well curved, the
posterior longer, oblique, curved; "laterals" strong, the outer ones
in the right valve well formed; ligament, short, thick.
Long. 2.2, alt. 1.8, diam. 1.5 mill.
Habitat: Fox river, Wisconsin, collected by the late Geo. H.
Marston; types in the collection of the University of Wisconsin and
that of the Carnegie Museum.
A number of specimens of this little Pisidium was received in
1895 and then recognized as a new species, and again in 1904, when
18 THE NAUTILUS.
the Marston collection was revised. In spite of its small size, it
ranges under the group Rivulina, and has resemblance, in shape,
with forms of P. compressum Pr. and with handwerkii St.; from the
latter it is easily distinguished: it is smaller, more oblique, its beaks
are comparatively broader, more posterior, and less elevated, its stria
much finer and slighter.
Pis. frctffilltmum, n. sp. Mussel rather small, inequipartite, ob-
lique, well inflated ; superior margin short, nearly straight, bounded
by angles from which the supero-anterior and posterior slopes fall
down nearly straight at first, then passing over the rounded anterior
and posterior ends into the moderately curved inferior margin ; an-
terior part longer than posterior and anterior end narrower ; beaks
somewhat broad and full, well projecting over the superior margin,
flattened on top : surface with very fine striae and a few slightly
marked lines of growth, dullish, pale horn-colored ; shell subtranslu-
cent, very thin and fragile ; hinge very slight, short, but well formed,
plate narrow ; cardinal teeth thin, lamellar, nearly straight and longi-
tudinal, the right long, its posterior end not or slightly thicker,
the left ones rather long, nearly parallel ; " laterals " slight, thin,
the anterior short, their cusps close to the cardinals, especially so the
left which is high, sharp pointed, abrupt, the right comparatively
large ; the outer ones of the right valve small but distinct and rather
long ; ligament slight.
Long. 3, alt. 2.6, diam. 1.8 mill.
Habitat: Lake Geneva, Wisconsin, sent for examination by Mr.
F. C. Baker. Types are in the collection of the Chicago Acad. Sc.,
and that of the Carnegie Museum.
Although only ten specimens were seen, this Pisidium is quite
distinct by its shape and the formation of the hinge ; the shell is the
thinnest and most fragile yet seen in Pisidia. The mussel has some
resemblance in shape with P. walkeri St., but is much smaller,
shorter, especially its anterior part, the surface sculpture is finer
and slighter, and the hinge is quite different. Its closer affinities
are yet unknown.
Pis. levissimum. n. sp. Mussel small, somewhat elongate, slightly
oblique, moderately inflated; superior margin moderately curved,
posterior rounded, sometimes with a slightly projecting, rounded
angle at the scutum; inferior slightly curved, supero-anterior slope,
at some distance from the beaks, in some specimens straight or
THE NAUTILUS. 19
nearly so, in others obsolete; beaks rather broad, rounded or slightly
flattened, little prominent over the upper margin; surface with very
fine, slight, irregular strise, shining; color pale horn to light amber,
shell translucent to transparent; hinge very slight, long, curved,
plate narrow; cardinal teeth: the right rather long, straight or
slightly curved, longitudinal, its posterior end more elevated, slightly
thicker and projecting downward over the hinge plate, the left ones
rather long, nearly longitudinal and parallel, lamellar, the posterior
(upper) slightly curved; "lateral teeth": the posterior much longer
than the anterior, slight, curved, the right not raised in a cusp, the
left with a slight cusp, the right anterior with a rounded, the left with
a sharp and abrupt cusp, outer ones in the right valve wanting, or a
mere trace of the anterior; ligament slight.
Long. 2. 5, alt. 2.0, diam. 1.4 m., the largest; average: 2.3 m. long.
Habitat: Elk Rapids, Michigan. About 300 specimens were seen,
at all stages of growth, most of them dead and bleached shells, sent
for examination by Mr. F. C. Baker. Types are in the collection
of the Chicago Acad. Sc. and that of the Carnegie Museum.
In shape and appearance, this Pisidium resembles a form of tenw's-
simum St., but is much smaller and slighter, and the formation of
the hinge is radically different, as ascertained on quite a number of
specimens of both species. In fact, the " lateral teeth " are quite
different from those of all other Pisidia seen.
PISIDIUM SUBROTUNDUM, n. sp. Mussel small, moderately and
regularly to rather well inflated ; outlines oval to short oblong to
nearly circular; superior margin straight or nearly so, as especially
noticeable in younger specimens ; beaks slightly posterior, rather
broad (in most forms), moderately projecting over the hinge margin,
thus completing the round outlines, except for the slightly project-
ing angles at the scutum and scutellum ; surface with very fine, sub-
regular to irregular striae and usually a few well marked lines of
growth, dullish to somewhat shining; color horn or light amber to
straw to whitish ; shell translucent (in the horn colored) to opaque
(in the light colored specimens), thin ; hinge slight and teeth small ;
ligament rather short and slight.
Long. 3.2, alt. 2.7, diam. 1.8 (100: 84.4 : ,56.3) average.
Long. 3.6, alt. 3, diam. 2.3 (100 : 83.3 : 63.9) Albany riv.
Long. 2.8, alt. 2.5, diam. 1.8 (100 : 89.3 : 64.3) Isle Royale.
Hab.: New England, Anticosti Is., to Michigan. Jupiter river,
20 THE NAUTILUS.
Anticosti Is., Albany river, Ontario (W. Mclnnes ed., both sent by
J. F. Whiteaves); "Vineyard," Pawtuxet, R. I., (J. F. Perry);
several places in Aroostook co., Me., (O. P. Nylander); Pine river,
Marquette co., and Oswald Lake, Mich. (B. Walker); Isle Royale,
Mich. (University of Michigan ; some other specimens from waters
on the same island were under doubt). Specimens from Union
springs, Ontonagon Co., Mich. (B. Walker), are of somewhat differ-
ent shape: the beaks are more posterior and narrower, and the an-
terior end is less broadly rounded. One good specimen was among
fossils from Arnold's Lake, Washtenaw Co., Mich. (B. Walker.)
Examples of this Pisidium had been in hand for years, un-
named, or a part doubtfully referred to other species. Since last
year, several good lots from widely distant places came in and proved
the species to be valid.
PISIDIUM FRIERSONI n. sp. Mussel rather small, moderately in-
flated, outlines somewhat angular ; superior margin slightly curved
or nearly straight, with projecting, rounded angles where passing into
the supero-anterior and the posterior, which are well marked, and
nearly straight, inferior moderately curved ; beaks scarcely posterior,
projecting over the hinge margin, rather narrow, with a somewhat
flattened central area ; surface with very fine, subregular strias, dull-
ish to somewhat shining, color pale horn to straw or whitish, shell
translucent to opaque, thin, hinge comparatively rather stout ; right
cardinal tooth curved, its posterior end thicker and grooved, left
anterior quite short, somewhat V-shaped, posterior rather short, rather
steep oblique ; outer laterals of the right valve distinct ; ligament short.
Long. 4, alt. 3.5, diam. 2.5 mill.
Hab. : Louisiana, Texas, New Mexico, Colorado. — In 1899, Mr.
L. G. Frierson sent me quite a number of specimens at all stages of
growth, collected at Frierson, La. ; they were regarded as repre-
senting a distinct species, but shelved. Since then I have seen the
same from Dallas (Cincinnati Museum) and Comal Co., Tex., the
Gallinas river, Las Vegas, N. Mex., and a few from Denver, Colo.,
(T. D. A. Cockerell).
The species seem to range under the group of P. noveboracense
Pr., and so far as known, shows little variation. It has been named
in honor of Mr. L. G. Frierson.
ERRATUM. — NAUTILUS, February issue, p. 119, line 2 from top:
after even, there should be a comma.
New Philadelphia, Ohio.
THE NAUTILUS. 21
A NEW SPHAEBIUM FROM ILLINOIS.
FRANK COLLINS BAKER.
Spheerium stamineum forbesi nov. var.
Shell of good size, inflated, solid, subequilateral, trigonal ; um-
bones much elevated, rounded, somewhat inflated, placed a little an-
terior of the center of the shell, marked by very fine, concentric
lines of growth (sometimes coarser), the beaks very closely approxi-
mating ; dorsal and ventral margins well rounded; anterior end flatly
rounded, posterior end plough-shaped ; both ends have a somewhat
truncated appearance ; umbonal slopes convexly rounded ; surface
inclined to be shining, lines of growth rather crowded, tine in typical
specimens, coarser in others ; color light greenish or yellowish-horn,
lighter on the umbones, indistinctly rayed in some specimens ; liga-
ment weak, short, brownish in color ; cardinal teeth similar in form
and position to those of stamineum, the hinge-line not quite so thick
as in stamineum ; lateral teeth not quite so solid as in stamineum, the
posterior laterals also being shorter, not reaching so high up into the
arch of the hinge-plait, the comparative distance between the an-
terior and posterior laterals being greater in forbesi than in stamineum ;
muscle scars and pallial line rather distinct; nacre faint bluish-white,
with occasional darker zones.
Length 14.50; height 11.50; breadth 8.00 mill, types.
" 12.00 " 10.00 " 7.00 u "
" 12.00 " 9.50 " 6.75 " Havana.
" 11.00 " 8.50 " 6.50 "
" 11.50 " 8.00 " 6.50 " "
Thompson's Lake, Fulton Co. (types) ; Matanzas Bay, Havana,
Mason Co.; Little Fox River, White Co. Types: — Illinois State
Laboratory of Natural History ; topotypes, Chicago Academy of
Sciences, Academy of Natural Sciences of Philadelphia.
This apparently distinct variety of stamineum may be known by
its peculiar trigonal shape, plough-shaped posterior end and elevated,
inflated umbones. The ventral and dorsal margins are much -more
rounded than in stamineum and the lateral teeth are farther apart.
The umbonal sculpture is typically very fine, but is also as coarse as
typical stamineum in some specimens.
The variety will probably prove to be a common form in many
localities and will be easily recognized and separated from typical
22 THE NAUTILUS.
stamineum. Twenty-two specimens from three localities show very
little variation.
I take great pleasure in naming this variety in honor of Prof. S.
A. Forbes, Director of the Illinois State Laboratory of Natural
History.
LIST OF WISCONSIN SHELLS.
C. H. CHADWICK.
(Continued)
C. FRESH-WATER UNIVALVES.
Carychium exiguum Say. Milwaukee.
" " exile H. C. Lea. Milwaukee.
Limnsea stagnalis appressa Say. Menomonee River ; Oak Creek
near South Milwaukee ; Okauchee Lake and Dela-
field, Waukesha Co., Two Rivers, Manitowoc Co. ;
Lake Winnebago near High Cliff.
" columella Say. Mill-pond at Delafield, Waukesha Co.
'" megasoma Say. Molas Creek, Manitowoc Co.
" reflexa Say. Milwaukee and vicinity (abundant); Oak
Creek, South Milwaukee ; Sand Ridge Creek, Kenosha
Co.; Delafield, Waukesha Co.
" palustris Miiller. Vicinity of Milwaukee ; North shore
of Lake Winnebago. (" Var. micltiganensis " Walker
is included).
catascopium Say. Lake Michigan at Milwaukee.
" " var. approaching L. emarginata Say. Lake
Micl).
'* caperata Say. Vicinity of Milwaukee; Lake Winnebago.
" " umbilicata Adams. Milwaukee and southwest-
ward ; Sand Ridge Creek, Kenosha.
" humilis Say. Milwaukee (scarce).
" desidiosa Say. Milwaukee and vicinity (abundant); Two
Rivers, Manitowoc Co.; North shore of Lake Winne-
bago.
Planorbis trii'olvis Sny. Milwaukee (common); Delafield and
Okauchee, Waukesha Co.; Two Rivers, Manitowoc
Co.
" frivol vis (large form). Molas Creek, Manitowoc Co.
THE NAUTILUS. 23
Planorbis bicarinatus Say. Milwaukee and vie.; Golden Lake,
Waukesha Co.; Little Cedar Lake, Washington Co.,
Lake Winnebago.
'< bicarinatus striatus Baker. Milwaukee (fossil).
" campanulatus Say. Milwaukee aud vicinity (common);
Delafield, Okauchee and Golden Lake, Waukesha Co.;
Little Cedar Lake, Washington Co.; Two Rivers,
Manitowoc Co.; Crystal Lake near Elkhart Lake.
Planorbis hirsutus Gould. Two rivers, Manitowoc Co.
" deflectus Say. Vicinity of Milwaukee; Cedar Lake,
Washington Co.; Two Rivers, Manitowoc Co.
" parvus Say. Milwaukee and southwestward.
Seymentina armigera Say. Milwaukee and southwestward; Two
Rivers, Manitowoc Co.
Ancylus rivularis Say. Root River, Milwaukee Co.
" paralellus Raid., var. St. Martin's, Milwaukee Co.
" shimeki Pils. Near Wauwatosa, (one shell).
Physa sayii Tappan. Milwaukee; Golden Lake, Waukesha Co.;
Lake Winnebago near High Cliff.
" gyrl'nt* Say. Vicinity of Milwaukee and southwestward
(abundant;) Sand Ridge Creek near Kenosha;
Two Rivers, Manitowoc Co.; Lake Winnebago.
" "• elliptica Lea. Vicinity of Milwaukee.
" intcgra. Haldeman. Milwaukee and vicinity (common;)
Golden Lake, Waukeslia Co.
ApJexa hypnorum Linne. Milwaukee.
Pleurocera subulare pallidum Lea (?) Lake Winnebago near
High Cliff.
" elevatum Say. Milwaukee River at Lindwurm.
Goniobasis livescens Menke. Milwaukee River and Lake Michigan.
Amnicola limosa Say. Milwaukee River; Lake Michigan; Lake
Winnebago near High Cliff.
<k '' parva Lea. Near Milwaukee.
" " porafa Say. Lake Michigan at Milwaukee.
" lustrica Pils. Honey Creek near Layton Park, Mil-
waukee.
" cincinnatiensisLiQa. Milwaukee and Menomonee rivers;
Lake Winnebago near High Cliff.
" euirtrginata Kti -ter. Lake Winnebago near High Cliff.
24 THE NAUTILUS.
Somatogyrus subglobosus Say. Root River and Kinnikinnick
River, Milwaukee Co.; Kenoslia; Lake Winne-
bago.
Pomatiopsis lapidaria Say. Near Wauwatosa, Milwaukee Co.
Valvata tricarinata Say. Milwaukee and vicinity (abundant);
Lake Winnebago.
" " simplex Gould. Lake Michigan at Mil-
waukee
" bicarinata normal is Walker? North Milwaukee.
" sincera Say. Near Milwaukee.
Vivipara contectoides W. G. B. Kinnikinnick River (one shell.)
Gampeloma rufum Haldeman. Okauchee Lake, Waukesha Co.;
Little Cedar Lake, Washington Co.; Lake Winne-
bago.
" decisum Say. Milwaukee and vicinity; Root River,
Milwaukee Co.; Two Rivers, Manitovvoc Co.; Lake
Winnebago.
" subsolldurn Anth. Milwaukee and vicinity; Oak
Creek, South Milwaukee; Molas Creek, Manitowoc
Co.
Lioplax subcarinata Say. Lenosha; Lake Winnebago.
NOTES.
In the May number of the NAUTILUS in an extract from Bulletin
49, Florida Agricultural Experiment station, the writer stated that
he did not know how long the ^Manatee snail (DrymoEus dormum")
had been living on orange trees in Manatee County.
I lived in that county near Bradentown from 1882 until 1886,
and during all my residence there diligently collected the land,
fresh-water and marine mollusks of that region. During my resi-
dence there I collected Drym&us dormani in moderate numbers liv-
ing on the orange trees of a grove in hammock land west of Braden-
town, and rarely on trees in a grove north of Manatee river. I also
found a few specimens in the Foster Orange Grove south of Mana-
tee. I found several hundred dead shells in the heavy hammock
among live oaks north of the Manatee river, and rarely in the air
pines on live oaks elsewhere, but I never considered it abundant any-
where.
It may be of interest to state that Mr. E. J. Brown, a neighbor of
mine, has found a few specimens of the nearly allied species (D.
dominions) living on his grapefruit and orange trees, and two or three
specimens of D. multilineatus on his trees. Drymczvs domlnicus is
rather a scarce species in this region. CIIAS. T. SIMPSON.
Little River, Florida, May 21, 1906.
THE NAUTILUS, XX
PLATE I.
9
8
WALKER, ON VALVATA.
THE NAUTILUS.
VOL. XX. JULY, 19O6. No. 3.
NOTES ON VALVATA.
BY BRYANT WALKER.
In 1824 Say described an ecarinate form of Valvata from the
"Northwest Territory "as V. sincera. In 1856 Dr. James Lewis
described a somewhat similar shell from the Little Lakes, N. Y., as
V. striata. W. G. Binney (L. & F. W. Shells III, 13.) in 1865,
had "no doubt of its identity with V. sincera." In this decision
Dr. Lewis seems to have acquiesced as it is not mentioned at all in
his " L. & F. W. Shells of New York " (1874), nor was it listed in
his collection.
In 1868, however, A. 0. Currier (Shell-bearing Moll, of Mich. p.
9.) declared Lewis' species to be a valid one and striata being pre-
occupied proposed for it the name of lewisii.
Binney's opinion seems to have generally prevailed among Amer-
ican conchologists until within the last year, when Dr. W. H.
Dall (Alaska XIII, pp. 22 & 23) declared them to be specifically
distinct.
From a critical study of several hundred specimens from more
than fifty different localities, I have come to a similar conclusion,
viz : that there are two distinct species which have hitherto passed
as Say's sincera. The one, smaller, more elevated, with a smaller
umbilicus, the other, larger, more depressed and widely umbilicated.
Both vary in sculpture from being nearly smooth, with subobsolete
strias to very strongly sculptured forms in which the striae are ele-
vated in " thin, sharp lamellae. "
Owing to the prevalent confusion of the two forms, the existing
26 THE NAUTILUS.
records afford but little information as to whether their distribution
is the same or not. Taking them together, sincera is apparently a
northern form ranging through Canada and the northern United
States, from the Atlantic to the Pacific. Its range to the south
seems comparatively restricted, especially in the pastern states,
northern New York, Michigan, northern Illinois and Indiana, and
Nebraska being apparently the southern limit, as it is not listed from
Philadelphia (Shick) nor Allegheny Co., Pa. (Stupakoff), Cincin-
nati (Harper & Wetherby), nor Tuscarawas Co., Ohio (Sterki), nor
in any of the Iowa,, Missouri, Arkansas or Kansas lists. I am in-
formed by Mr. F. C. Baker that the southern range (south to Geor-
gia and Louisiana) attributed to this species in his " Mollusca of the
Chicago Area," was a typographical error and should have been
placed under V. tricarinata. In the extreme west, it would seem
to range further south, as Dall quotes the broadly umbilicated form
from the San Bernardino Mountains, Cal., and Utah. Ingersoll
also (Rep. U. S. G. & G. Survey, 1874, p. 390) quotes sincera from
Colorado and Utah. The only Utah specimens I have seen, are
rather referable to V. humerolis Say. and it is possible that all these
extreme southwestern citations should be revised.
VALVATA SINCERA Say, PI. I, figs. 1-6.
As Say's type of V. sincera has not been preserved, in determin-
ing which of the two forms, which have hitherto passed under
that name, is to be regarded as sincera we are necessarily re-
stricted to his original description and figure (a very poor one)
and such legitimate deductions as can be drawn from the circum-
stances under which his description was prepared, and his other
writings. When Say described his new species in 1824, the only
American species known to him was the typical V. tricarinata
and his purpose was to differentiate these two forms. Owing, no
doubt, to the striking and characteristic sculpture of Y. tricarinata,
Say omitted any description of general shape. He states that V-
sincera is " subglobose-conic " and " very similar " to tricarinata,
but with a " rather larger " umbilicus. And later in describing his
V. humeralis, he states that it is " subglobose " and " more depressed "
than sincera. It hardly seems possible that Say with his remarkable
acumen and critical appreciation of minute differences, would have
failed to call attention to the depressed form and very wide umbili-
THE NAUTILUS. 27
cus in differentiating his new species from tricarinata, had he had
before him a specimen of lewisii. Then too, his figure, whatever
else may be said of it, was certainly never intended to represent a
shell of that form. Say further describes his species as having the
whorls " finely and regularly wrinkled across." An examination of
Say's descriptions of other species, especially his earlier ones, will
show that he commonly used the terms " wrinkles " and " wrinkled "
in the same sense that today we would use stria? and striate. Thus
Polygyra albolabris, thyroides, profunda and sayii, Pyramidula soli-
taria, Gircinaria concava and Bulimulus dealbatus are characterized
as " wrinkled." Bulimulus multilatus is " striated with regular ele-
vated lines or wrinkles." And Valvata humeralis is stated to be
" wrinkled across or rather with slightly raised lines."
Taking all these things into consideration, we should say that
Say's sincera was a subglobose-conic shell, of about the same size and
shape of V- tricarinata, but with a rather larger umbilicus and with
accurately rounded, finely and regularly striated whorls. Shells
answering very exactly to these requirements were collected by the
Michigan Fish Commission in 1894, at High Island Harbor, Beaver
Islands, Lake Michigan, and were listed as that species, though with
no appreciation of its distinction from lewisii (Bull. Mich. Fish
Com. No. 6, 99 ; Naut. IX, 5). And there would seem to be no
reasonable objection to these being considered as typical examples of
Say's species.
As shown by the figures (figs. 1-3) the general shape of the shell,
with its " accurately rounded " whorls and round, deep umbilicus is
in exact accord with the original diagnosis. The surface is sculptured
with fine, close, regular, thread-like stria?, with a marked tendency
in many examples to rise in thin, sharp ribs approximating to those
of nylanderi. But in no instance are these more elevated ribs as
strongly developed as in that form, nor are they so widely spaced.
The specimen figured has 3| whorls and measures : Alt. 4, diam.
5 mm.
Many of the specimens have the apical whorls more or less deeply
tinged with dull red, the rest of the shell is of a pale greenish, with
the apical whorls nearly white in the unicolored examples.
There is some slight variation in height and correlated with that,
in the width of the umbilicus, but not sufficient to connect them
specifically with the widely umbilicated leioisii.
28 THE NAUTILUS.
Through the kindness of Dr. Whiteaves I have been enabled to
examine some of the Anticosti shells referred to by Dr. Dall. Com-
pared with the High Island specimens, these (figs. 4-6) are thinner*
less heavily striated, with a rather wider umbilicus, and the body
whorl is less inflated, but they evidently belong to the same species.
As above defined, V. sincera appears to be a less abundant species
than lewisii, but of considerable range, though apparently rather local
in its habitats. Specimens have been examined from the following
localities, in addition to those above mentioned : River Rouge, Wayne
county ; Mill Lake, Lake county and Lake Michigan, off New York
Point, Charlevoix county, Michigan ; Pigeon Lake, Ontario and
Milwaukee, Wis. Some immature specimens from Lake George,
N. Y., should also probably be included. It has also been found in
the marl deposits at Cement City, Jackson county; Ann Arbor;
Monitor, Bay county, and Pine Lake, Charlevoix county, Michigan,
and Caribou, Me. As all the marl specimen s are decorticated, it is
impossible to say whether they were originally typical sincera or var.
nylanderi.
V. SINCERA NTLANDERI Dall. PI. I, figS. 7-9.
Judging from the suite examined, nylanderi (figs. 7 and 8) in its
original locality is a well marked constant form, and standing alone
might well be accorded specific rank. But as stated above, the
specimens from High Island Harbor, show decided tendency to ac-
centuate the stria?, and in a considerable number from Isle Royale,
Lake Superior (collection of 1905), where typical sincera is not
represented, all the gradations from close, elevated ribs to the wide
spaced lamellae of typical nylanderi are represented. So that there
would seem to be no question, but that it is to be considered as a
varietal form of Say's species. This form in its various aspects has
also been found in Cliff river and Pine Lake, Marquette county,
Michigan ; Little Lakes, N. Y., and Lake Geneva, Wis. An un-
usually elevated example from Isle Royale, is illustrated by fig. 9.
V. SINCERA DANIELSI, n. v. PI. I, figs. 10 and 11.
Shell larger and proportionately higher than the typical form ;
whorls 4, finely and regularly striate, very convex with a deeply im-
pressed suture. Aperture entire, scarcely appressed to the body
whorl.
Alt. 6, diam. 5.75 mm.
THE NAUTILUS. 29
Cannon Lake, Rice county, Minn., Types No. 22475, coll.
Walker. Cotypes in the collections of L. E. Daniels and the Phila-
delphia Academy. Sixteen specimens of this fine large form were
collected by Mr. Daniels. They differ uniformly from the High
Island typical form in the particulars mentioned.
VALVATA LEWISII Currier. PI. I, figs. 12 and 13.
Dr. Lewis distinguished his V- striata by reason of its having the
" epidermis brown and very regularly striate." In the same paper
(Proc. P. A. N. S., 1856, 260) he describes " V. sincera" from the
same locality as having the " shell smooth and polished, white and
translucent." While unfortunately he does not seem to keep any
examples of his typical striata in his own collection, there is a set
from the Little Lakes labelled " sincera " which agrees with his de-
scription and which are the same as the form herein described as F
bicarinata perdepressa. This leaves practically no doubt as to his
striata being the ordinary striate form.
F lewisii in its aggregate form is exceedingly variable in regard
to sculpture, ranging from the smooth or obsoletely striate var. heli-
coidea to a heavily ribbed form in which the ribs become low, closely
spaced lamella? very similar to the Isle Royale form of sincera
nylanderi. In well developed examples of this form, the ribs give a
brownish appearance to the shell, especially when not well cleaned,
and it is possible that such specimens were the types of Lewis' species,
and if varietal distinction were to be made between this and the more
common form in which the sculpture is " fine and close like the wind-
ing of thread on a spool," it should be considered as the typical
lewisii.1 From the material before me, however, I fail to find suffi-
cient evidence of any racial distinction in this particular, (as there is
apparently in the case of V. sincera nylanderi) to justify such action.
The two forms apparently live together and grade indefinitely into
each other.
V. BICARINATA Lea. PI. I, fig. 14.
When I prepared my former paper on Valvata (NAUT., XV, p. 123)
I had not seen Prof. B. Shimek's " Mollusca of Eastern Iowa,"
(Bui. Lab. Nat. Hist. State Univ., la., I, p. 56, 1888) in which the
specific validity of this species is upheld.
1 This will have to be determined from examination of authentic author's
specimens which I have not been able to obtain.
30 THE NAUTILUS.
For comparison with the other forms herein discussed, a typical
example is figured.
V' bicarinata has recently been collected by Messrs. H. H. Smith
and A. A. Hinkley at several localities in Alabama, greatly extend-
ing its range towards the south.
V. BICARINATA CONNECTANS n. V.
Unicarinate, only the superior carina present, peripheral and basal
carinae obsolete, periphery rounded, base rounded or subangulated.
Type No. 24142, coll. Walker. Lake Michigan, New Buffalo, Mich.,
also Lake Ontario, N. Y. Cotype in collection of L. E. Daniels,
LaPorte, Ind.
Apparently of rather rare occurrence as only two specimens, wera
collected by Mr. Daniels in a considerable amount of material from
several localities along the south shore of Lake Michigan. A single
specimen also was found in a small set of V. bicarinata perdepressa
from Lake Ontario, N. Y., received from Mr. F. C. Baker. This
specimen is of special interest as illustrating the relation of this form
and the succeeding one. Beginning as a typical connectans, the
carina disappears about the beginning of the last whorl which is that
of a typical perdepressa.
V. BICARINATA PERDEPRESSA n. V. PI. I, figS. 15 and 16.
Shell broadly umbilicated, very much depressed, often planorboid.
Ecarinate, whorls regularly rounded, occasionally subangulated
around the umbilicus or at the periphery, smooth or obsoletely striate,
frequently obscurely malleated, especially on the lower half of the
whorl, very pale horn-colored, unicolored or with the apical whorls
more or less tinged with dull purple or red. Types (No. 24123,
coll. Walker) from Lake Michigan at Michigan City, Ind. Also
from Lake Michigan at Chicago, Ills., Millers, Ind., New Buffalo,
Mich., Lake Erie at Sandusky, Ohio, Lake Ontario at Jefferson
county and ?, N. Y., and Little Lakes, N. Y. Cotypes in
the collection of L. E. Daniels, LaPorte, Ind., and the Philadelphia
Academy.
This form is very abundant along the south shore of Lake Michi-
gan. It would be taken for a smooth form of V, lewisii, were it not
for the connecting links with V. bicarinata afforded by the occasional
subangulated individuals and bicarinata connectans. It is very vari-
THE NAUTILUS. 31
able in shape, ranging from planorboid with the apex depressed
below the level of the body whorl to specimens with the apex quite
as much elevated as in the ordinary specimens of lewisii. In sculp-
ture it varies from almost perfectly smooth to a striation nearly as
strong as that of typical lewisii. Indeed the more elevated and
strongly striated examples standing alone would unhesitatingly be re-
ferred to that species.
Dr. Dall, who kindly compared specimens with his V~ lewisii heli-
coidea writes : " The figured specimen (helicoidea} should have (to
be adult) about one-half more whorl than it has in the figure. Some
of yours, if the same view be taken, are almost exactly like the speci-
men figured. But, on comparison of adults, it is evident that the
lewisii helicoidea has a smaller umbilicus ; and, taken in profile, is less
flattened. Your specimens, too, are on the whole smoother and
more polished. The value of these differences remains to be deter-
mined."
The wider umbilicus in the Lake Michigan shells is undoubtedly
correlated with the more depressed shape. The specimens collected
by Lewis in the Little Lakes, N. Y., are indistinguishable from those
from Lake Michigan. The Lake Erie and Lake Ontario specimens
are uniformly smaller, but otherwise substantially the same.
The difficulty in finding any tangible distinction between the more
elevated, substriated individuals of this form and the obsoletely
striated examples of V- lewisii is similar to that experienced in at-
tempting to separate V- tricarinata simplex from the smooth or obso-
letely striated forms of V. sincera. And this naturally raises a query
as to whether there are really more than two polymorphic species
differing constantly in shape, but both varying in sculpture, all the
way from tricarinate to smooth, and from smooth to ribbed-striate.
Both series are complete and parallelism is absolutely exact.
Group of Group of
V. tricarinata. V. bicarinata.
Tricarinate, V- tricarinata. V- bicarinata nor-
malis.
Bicarinate, Y. tricarinata confusa. V- bicarinata.
Unicarinate, V. tricarinata unicarinata. V- bicarinata con-
nectans.
Smooth, Y> tricarinata simplex. V. bicarinata per-
depressa.
32 THE NAUTILUS.
Obsoletely striate, V. sincera (Anticosti). V. lewisiihelicoidea.
Thread-striate, V. sincera. V. lewisii (var. ?)
Ribbed-striate, V. sincera nylanderi. V. leivisii (typical ?)
In view of the recognized variability of all these forms, the in-
ference to be drawn would seem inevitable.
EXPLANATION OF PLATE I.
All the figures are drawn on the same scale (X 5) except 4, 5 and
6 which were drawn by Miss Helen Winchester, and these are sub-
stantially the same.
Fig. 1-3. V. sincera, High Island Harbor, Lake Michigan.
" 4—6. V. sincera, Anticosti.
" 7—9. V. sincera nylanderi, Portage Lake, Aroostook Co., Me.
" 9. V. sincera nylanderi, Isle Royale, Mich.
" 10-11. V. sincera danielsi, Cannon Lake, Rice Co., Minn.
" 12-13. V. lewisii, Traverse City, Mich.
" 14. V. bicarinata, Philadelphia, Pa.
" 15-16. V. bicarinata perdepressa, Lake Mich., Michigan
City, Ind.
HELIX JEJUNA SAY TRANSFERRED TO PRATICOLELLA.
BY HENRY A. PILSBRY.
Recently I received from Mr. George H. Clapp some living ex-
amples of this small snail, with a request that they be examined
anatomically to determine the generic position of the form, which
has of recent years been grouped with the small Polygyras of the
" Mesodon " type.
The snails were collected at Lemon City, Dade Co., Florida, by
Mr. Chas. T. Simpson. Externally the soft parts do not differ from
Polygyra and many other genera of Helices.
The terminal ducts of the genitalia are shown in Fig. 1. The
penis bears the retractor and vas deferens at its apex. Near the
base a very large gland or " appendix " is inserted (a in the figure).
This seems to be a nearly solid but soft and readily broken gland.
The vagina is more than half as long as the penis. The spermatheca
THE NAUTILUS.
33
is large and oblong, upon an extremely short duct. Other organs as
usual in Helices.
No Polygyra or " Mesodon " lias a gland or appendix upon the
penis, but in the Mexican genus Praticolella, represented in Texas
by P. griseola and P. berlandieriana, a similar structure is found.
I do not hesitate to refer jejuna to the genus Praticolella on the evi-
dence of this peculiar organ, which has been noticed in no other
genus of Helices within our limits. The Floridian snail differs from
the Texan and Mexican forms by having an apparently simple penis
retractor muscle, while in the other species it is split and has a triple
Fig. 1. P, penis; p, r, its retractor; a, penial gland ; vd, vas deferens ; ut,
uterus ; sp, spernatheca ; vag, vagina.
insertion on penis and vas deferens. This simple condition was
probably the ancestral structure of the genus, since it is that of all
other related Helices.
The shell of this species, which we will now call Praticolella je-
juna (Say), agrees somewhat better with that of Praticolella than
with any other of our genera of Helices. Not much is known of the
habits of the snail. It is quite a local form, which I never happened
upon at Miami or elsewhere in some half-dozen visits to Florida.
Mr. S. N. Rhoads found it on the " under side of young cabbage
palms" at Miami. Further information upon the situations it affects
will be of interest In Texas, Praticolella berlandieriana lives on
bushes with Bulimulus dealbatus mooreanus, but during prolonged
dry weather it burrows in the ground.
34 THE NAUTILUS.
SOME SHELLS OF MISSISSIPPI AND ALABAMA.
BY A. A. HINKLEY.
In the fall of 1904, with the aid of some of my conchological
friends, for the purpose of collecting in that region, a trip was made
to Abbeville, Grenada, Durant and Columbus, Mississippi, and
Tuscaloosa, Boligee, Tuscumbia and Florence, Alabama.
The streams along the Illinois Central railroad where stops were
made, have sandy beds and not favorable for molluscan life, with the
exception of two specimens of Pleurocera elevatum, no representatives
of the StrepomatidcB were found in these streams.
In determination of material collected thanks are due Messrs.
Bryant Walker, Wm. A. Marsh, L. S. Frierson, Dr. V. Sterki, F.
C. Baker and H. A. Pilsbry.
Family HEI.ICINID^E.
Helicina orbiculata Say. Boligee, Ala., dead specimens only.
Family HELICID.E.
Polygyra leporina Gould. Abbeville and Columbus, Miss., Boligee,
Ala.
Polygyra troostiana Lea. Florence, Ala.
Polygyra fraudulenta Pilsbry. Florence, Ala.
Polygyra inflecta Say. A common species found at all localities.
Polygyra exoleta Binney. Florence, Ala.
Polygyra palliata Say. Abbeville, Miss., Boligee and Tuscaloosa,
Ala.
Polygyra obstricta Say. Columbus, Miss., Florence, Ala.
Polygyra appressa Say. Florence, Ala.
Polygyra appressa perigrapta Pilsbry. Grenada and Columbus,
Miss.
Polygyra thyroides Say. Abbeville, Grenada and Columbus,
Miss., Tuscaloosa, Boligee and Florence, Ala. At Boligee they
were numerous along the railroad right of way. Ic the same situa-
tion though not as numerous were P. clausa and B. dealbatus.
Polygyra clausa Say. Boligee, Ala., varies a little in size and
thickness of the shell.
Polygyra spinosa Lea. Florence, Ala. Common on both sides
of the river.
THE NAUTILUS. 35
Polygyra barbigera Redfield. Tuscaloosa, Ala., a rare species.
Polygyra subglobosa Pilsbry. Columbus, Miss., Tuscaloosa and
Florence, Ala.
Polygyra monodon Rack. Abbeville and Columbus, Miss.,
Florence, Ala.
Family BULIMULID^E.
Bulimulus dealbatus Say. Boligee, Ala. Dead specimens were
very numerous in some of the fields.
Family PUPILLID^E.
Strobilopsis labyrinthica Say. Boligee, Ala., Abbeville, Durant
and Columbus, Miss.
Pupoides fallax Say. Boligee and Florence, Ala.
Bifidaria armifera Say. Boligee, Ala.
Bifidaria contracta Say. Boligee, Ala.
Bifidaria procera Gould. Boligee and Florence, Ala. At the
former place it was found in tufts of grass in an open field, at the
latter under stones in an open close pastured place along the river.
Family GLANDINID^E.
Glandina truncata Brug. Boligee, Ala. The remains of nests of
eggs were often found about old logs and young shells were quite
common. A mature horn-colored specimen was an interesting varia-
tion.
Family CIRCINARIID^E.
Circinaria concava Say. Tuscaloosa, Boligee and Florence, Ala.
Family
Omphalina fuliginosa Griff. Florence, Ala.
Omphalina laevigata Pfr. Abbeville, Grenada and Columbus,
Miss., Tuscaloosa, Boligee and Florence, Ala.
Vitrea carolinensis Ckll. Boligee and Florence, Ala.
Euconulus fulvus Mull. Abbeville, Miss., Boligee, Ala.
Zonitoides arborea Say. Abbeville, Grenada Durant and Colum-
bus, Miss., Boligee, Ala.
Zonitoides minuscula Binney. Boligee, Ala.
Gastrodonta demissa Binney. Columbus, Miss., Tuscaloosa, Bo-
ligee and Florence, Ala.
Gastrodonta interna Say. Tuscaloosa and Florence, Ala.
36 THE NAUTILUS.
Family ENDODONTID^E.
Pyramidula alternata, Say. Abbeville, Grenada and Columbus,
Miss., Tuscaloosa and Boligee, Ala.
(To be continued}.
NOTES.
THE RAVENEL COLLECTION — The museum of the College of
Charleston, S. C., has acquired the valuable conchological collection
of the late Dr. Edmund Raven el, of Charleston, which contains some
3,500 species of land, fresh-water and marine shells.
LYMN^EA MEGASOMA SAY. — Fine specimens of this rare species
have recently been taken at Lake Minnetonka, Minn., by Mrs. A.
W. Abbott of Minneapolis.
PUBLICATIONS RECEIVED.
NOTES ON JAPANESE, INDO-PACIFIC, AND AMERICAN PYRAMID-
ELLIDJE. By Wm. H. Dall and Paul Bartsch, (Proc. U. S. Nat.
Mus., XXX, pages 321-369, plates XVII-XXVI, 1906). This
valuable work on the PyramideUidae is based in part on the collec-
tion of the museum of Berlin loaned to the U. S. National Museum
for study. Some 22 new species are described and figured, and a
number of new names are proposed to replace those pre-occupied.
Three new subgenera Nisiturris, Bobella and Egilina are also defined.
DESCRIPTION OF Two NEW NAIADS. By Paul Bartsch, (Proc.
U. S. Nat. Mus., XXX, pages 393-396, Plates 27-29, 1906). The
species are Nephronaias flucki from Nicaragua and Diplodon cmapen-
sis from Argentina.
RECENT MOLLUSCA : A CATALOGUE OP OPERCULATE LAND
SHELLS. By G. B. Sowerby and H. Fulton. London, 1906. A
useful list of genera and species arranged " with slight modifications "
according to Kobelt and MollendorfF's catalogue.
ANATOMY OF ACM.EA TESTUDINALIS MULLER, Part I. Intro-
ductory material. External anatomy. By M. A. Willcox. Reprint
the American Naturalist, Vol. XL, pp. 171-187, 1906.
THE NAUTILUS, XX.
PLATE II
RAYMOND: WEST AMERICAN SPECIES OF GENOTA
THE NAUTILUS.
VOL. XX. AUGUST, 19O6. No. 4.
THE WEST AMERICAN SPECIES OF PLEUROTOMA, SUBGENUS GENOTA.
BY WILLIAM JAMES RAYMOND.
Five species of the subgenus Genota, section Dolichotoma, have
been described from the Pacific coast of North America. Three of
the species have been figured ; two have not hitherto. Specimens of
all five, including the types of four, being temporarily in possession
of the writer, it seemed desirable to bring them together in a single
photographic plate. Apparently derived from a common stock, and
arising in late tertiary time, the nominal species are closely allied.
Yet there is convenience in retaining the present specific designa-
tions of forms which are readily distinguishable. Two of the species
have not thus far been found living. As the deeper water along the
coast is explored, they may possibly be discovered in the dredge,
together with other forms not now known.
Pleurotoma ( Genota) carpenleriana Gabb.
Pleurotoma (Surcula) carpenteriana Gabb. Proc. Cal. Acad. Nat.
Sci., 1865, p. 183 ; Pal. Cal., vol. 2, p. 5, pi. 1, fig. 8.
Pliocene — San Fernando and Santa Rosa (Gabb) ; San Diego
(Ball).
Pleistocene — Santa Barbara and San Pedro (Gabb); San Pedro,
San Diego and Ventura (Arnold) ; Santa Monica (Rivers).
Living — Drake's Bay, 30 fathoms (Arnheim); Monterey to San
Diego (Cooper); San Pedro, Santa Catalina Island and San Diego,
10 to 100 fathoms (Raymond); Cerros Island (Dall).
This is the largest and most plentiful species of the group. It is
38 THE NAUTILUS.
characterized by a regularly fusiform outline, eight whorls, fine spiral
sculpture, slight concavity near the suture, slight convexity of the
anterior part of the upper whorls, the obtuse shoulder thus formed
being below the middle of each whorl and sometimes ornamented
with inconspicuous nodes. Plate II, fig. 1, illustrates Gabb's type,
found in the pleistocene of Santa Barbara. It is the property of the
University of California. Fig. 3 shows an example close to type.
It was dredged in about 100 fathoms, off Santa Catalina Island.
Length 80 mm. In fig. 2 the nodes are distinguishable, which when
better developed are a diagnostic feature of the next species.
Length 54 mm.
Pleurotoma (Genota} tryoniana Gabb.
Pleurotoma (Surcula) tryoniana Gabb, Pal. Cal., vol. 2, p. 6, pi. 1,
fig. 9.
Pleistocene — San Pedro (Gabb); Santa Barbara and San Pedro
(Cooper); Santa Monica (Rivers).
Living — San Diego (Cooper); San Pedro (Oldroyd).
Typically the angular, nodose whorls, eight in number, separate
this species from Genota carpenteriana, but intermediate forms like the
example shown in fig. 2 are found rarely, and indicate the close alli-
ance of the two species. Plate II, fig 7, illustrates Gabb's type,
found in the pleistocene of San Pedro. Fig. 8 represents a specimen
found at Santa Monica. Its length is 62 mm.
Pleurotoma (Genota) cooperi Arnold. Mem. Cal. Acad. Sci., Vol.
Ill, p. 203, pi. vii, fig. 3.
Pleistocene. — San Pedro (Arnold); Santa Monica (Rivers).
Characterized by the angular whorls, seven or eight in number,
decidedly concave above the nodose angle, with relatively sharp
sculpture. The nodes are more numerous than in the last species
and become less prominent toward the aperture. Aperture less than
waif the length of the shell. Plate II, fig. 10, illustrates the single
specimen found in the pleistocene of Santa Monica.
Pleurotoma (Genota} stearnsiana Raymond. NAUTILUS, Vol.
XVIII, p. 1.
Living. — San Diego and Santa Catalina Island, 25 to 40 fathoms
(Raymond).
THE NAUTILUS. 39
The species is characterized by its small size, eight or more
whorls, relatively wide form and broad color bands. Plate II, fig. 6,
represents the type. Length 30.5 mm. An old specimen, length
41.5 mm., is shown in fig. 5, and a younger example in fig. 4. All
are from San Diego.
Pleurotoma ( Genota~) riversiana Raymond. NAUTILUS, Vol. XVIII,
p. 14.
Pliocene. — Santa Monica (Rivers).
Characterized by narrow form, sharply expressed sculpture and
obtuse angle above the middle of the whorls. Plate II, fig. 9,
illustrates the type which thus far is the only specimen found.
Length 59 mm.
University of California, July, 1906.
NOTE ON THE GENUS GLABARIS GRAY OR PATULARIA SWAINSON,
BY WILLIAM HEALY DALL.
The genus Patularia Swainson, appears in his Malacology (1840)
pp. 287 and 381. There are two species of which the first is Anodon
ovatus Swainson, Exotic Conchology pi. xxxvi, 1823, (2nd ed. by
Hanley, p. 30, 1841) not Iridina ovata Swainson, Phil. Mag., 1823.
The second species A, rotundatus Swainson, is doubtfully referred to
Anodonta by Simpson in his Synopsis, p. 638. If it be as he sup-
poses synonymous with A. woodiana Lea, Swainson's name dating
from 1823, will of course take precedence. In 1841 Swainson cites
under his Patularia ovata Anodonta trapesialis Lamarck, and Han-
ley points out that the latter specific name, being four years older,
must take precedence of ovatus Swainson. Anodonta trapesialis,
however, in modern classification, is a typical Glabaris of Gray.
But Glabaris Gray, dates only from 1847, when the name was
applied to A. exotica Lamarck, by Gray, in his list of generic syn-
onyms, Proc. Zool. Soc. Lond., 1847, p. 197.
It would seem, therefore, that, since the first species and type of
Patularia is a Glabaris, that the former name should be substituted
for the latter in our systems. An examination of the nomenclators
fails to show any earlier use of the generic name Patularia in zoology
and there seems to be no reason which would militate against its
adoption.
40 THE NAUTILUS.
A careful examination of the text of the second edition of the
Exotic Conehology shows that this is the only name included in it
which is likely to affect the existing nomenclature.
SOME SHELLS OF MISSISSIPPI AND ALABAMA.
BY A. A. II1NKLEY.
(Concluded from July Number , p. 36).
Family SUCCINEID^E.
Succinea concordialis Gould. Durant, Miss., one dead shell was
all found. In the summer of 1904 this species was plentiful at the
R. R. bridge across Little Muddy near Du Bois, 111.
Succinea aurea Lea. Florence, Ala.
Family LIMNAEID^E.
Lymncea desidiosa Say. Tuscaloosa and Florence, Ala.
Lymner.a caper ata Say. Boligee, Ala.
Lymncea cohimella Say. Florence, Ala.
Physa gyrina Say. Florence, Ala.
Physa heterostropha Say. Tuscumbia, Ala.
Physa pomilia Conrad. A rill below Columbus, Miss.
Physa troostiana Lea. Spring above Florence, Ala.
Planorbis trivolvis var. glabratus Say. Shoal creek, Florence,
Ala.
Planorbis trivolvis Say. Durant, Miss. One specimen.
Planorbis dilatatus Gould. Grenada, Miss.
Segmentina wheatleyi Lea. Boligee, Ala.
Ancylns elatior Anthony. Tennessee river, Florence, Ala.
Ancylus kirklandi Walker. Tombigbee and Yalabusha rivers.
Ancylus rhodaceus Walker. Tennessee river, Florence, Ala.
Family AURICULID^E.
Carychium exiguum Say. Abbeville, Miss. Only found two
specimens.
Family STREPOMATIDJE.
Io spinosa Lea. Tennessee river, Florence, Ala. One specimen
only of this fine species was found.
THE NAUTILUS. 41
Angitrema salebrosa Conrad. Tennessee river, Florence, common.
Angitrema subglobosa Lea. Tennessee river, Florence, Ala.
Angitrema tuomeyi Lea. Tennessee river, Florence, Ala. A few
were found in 1894 but none this time.
Angitrema duttoniana Lea ?. Tennessee river, Florence, Ala.
Angitrema lima Conrad. Cypress Creek, Florence, Ala. A few
faintly banded.
Angitrema verrucosa Raf. Tennessee river, Florence, Ala.
Lithasia Jlorentina Lea. Cypress and Shoal creeks, Florence,
Ala. Found only a few.
Lithasia nuceola Anthony. Tennessee river, Florence, Ala.
Doubtful determination.
Lithasia curta Lea. Tennessee river and Shoal creek, Florence,
Ala.
Strephobasis plena Anthony. Tennessee river, Florence, Ala.
Strephobasis curta Hald. Tennessee river and Shoal creek, Flo-
rence, Ala.
Pleurocera alveare. Tennessee river, Cypress and Shoal creeks,
Florence, Ala.
Pleurocera undulatum Say. Tennessee river, Florence, Ala.
Pleurocera excuralum Conrad. Tennessee river, Florence, Ala.
Pleurocera moniliferum Lea. Tennessee river, Florence, Ala.
Pleurocera ponderosum Anthony. Tennessee river, Florence, Ala.
Pleurocera moriforme Lea. Tennesee river, Florence, Ala.
Pleurocera thorntonii Lea. Tennessee river, Florence, Ala.
Pleurocera annuliferum Conrad. Black Warrior, Tuscaloosa, Ala.
Pleurocera elevatum Lea. Yallabusha river, Granada, Miss.
Only two specimens found, they were covered with a thick deposit
which being removed revealed a polished, olive colored epidermus
and two strong bands.
Pleurocera trivittatum Lea. Tombigbee river, Columbus, Miss.
Pleurocera incurvum Lea. Cypress creek, Florence, Ala.
Pleurocera alabamensis Lea. Cypress creek, Florence, Ala.
Pleurocera olivaceum Lea. Tombigbee river, Columbus, Miss.
Pleurocera lativittatum Lea. Shoal creek, Florence, Ala.
Goniobasis acuta Lea. Florence, Ala.
Goniobasis formosa Conrad. Florence, Ala.
Goniobasis hydeii Conrad. Black Warrior, Tuscaloosa, Ala.
Goniobasis interveniens Lea. Florence, Ala.
42 THE NAUTILUS.
Goniobasis nassula Conrad. Spring in Tuscumbia, Ala.
Goniobasis thorntonii Lea. Spring in Tuscumbia, Ala.
Goniobasis aterina Lea. Florence, Ala. Found with Pomatiopsis
Mnkleyi.
Goniobasis nitens Lea. Florence, Ala. This determination is
doubtful.
Goniobasis adusta Anthony. Spring near Cypress creek, Flo-
rence, Ala.
Goniobasis paupercula Lea. Small stream above Florence, Ala.
Euryccelon crassa Hald. Tennessee river, Florence, Ala. Only
one very young shell found. When at this place in 1894 this form
was found of all sizes with three wide bands and the carina of the
young shells remarkably developed, Pilsbry says this is Eu. anthonyi.
Anculosa carinata Brug. Tennessee river, Florence, Ala. A
few specimens can be referred to no other species and are put under
this name with some doubt.
Anculosa subglobosa Say. Tennessee river and Shoal creek, Flo-
rence, Ala.
Anculosa praerosa, Say. Tennessee river and Cypress creek,
Florence, Ala.
Anculosa tintinnabulum Tennessee river and Shoal creek, Florence,
Ala.
Family VIVIPARIDVE.
Vioipara intertexta Say. Rill near Durant, Miss., and a rill near
Boligee, Ala., both places in heavy timber.
Campeloma ponderosum Say. Tennessee river, Florence and Tom-
bigbee river west of Boligee, Ala.
Campeloma decisurn Say. Tennessee river, Florence, Ala.
Campeloma decisum var. heterostropha. One specimen. Tennessee
river.
Campeloma coarctatum Lea. Tallahatchie. Yalabusha, Big Black
and Tombigbee rivers.
Campeloma decampi Currier. Tennessee river, Florence, Ala.
Family AMNICOLID^E.
Amnicola cincinnatiensis Anth. Big Black and Tombigbee rivers.
Atnnicola limosa Say. Tallahatchie, Yalabusha, Big Black and
Tombigbee rivers.
Amnicola auyustina Pilsbry. Tuscumbia, Ala. Found in the
THE NAUTILUS. 43
stream running from the spring, they were in algse and on the muddy
bottom.
Somatogyrus subglobosa Say. Tallahatchie and Yalabusha rivers.
Somatogyrus georgianus Walker. Tennessee river, Florence, Ala.
Somatogyrus substriatus Walker. Tombigbee and Tennessee
rivers. The most plentiful species of the genus in these two streams.
Somatogyrus humerosus Walker. Tennessee river, Florence, Ala.
Somatogyrus quadratus Walker. Tennessee river and Shoal crock.
Somatogyrus strengi Pilsbry and Walker. Tennessee river and
Shoal creek.
Somatogyrus biangutatus Walker. Tennessee river, Florence,
Ala.
Somatogyrus excavatus Walker. Shoal creek, Florence, Ala.
Somatogyrus tennesseensis Walker. Shoal creek.
Somatogyrus purnilus Conrad. Tennessee river, Florence, Ala.
Pyrgulopsis mississippiensis Pilsbry. Shoal creek. This little
species was numerous a short distance above the canal aqueduct,
where sediment had collected on the rock bottom, a few were found
on algae. They were only found in mid-stream.
Pomatiopsis lapidaria Say. Found on leaves after a shower, in a
moist situation, always shaded by timber and scrub palmetto. Boligee,
Ala.
Pomatiopsis hinkleyi Pilsbry. This species was very numerous in
the same places where the types were found in 1894. It was also
found across the river high up on the river bluff, on and in damp
moss but not in a wet situation like the others. See vol. X, p. 37
of the NAUTILUS.
Family CORBICULID^E.
Calyculina transversa Say. Yalabusha, Black and Tombigbee
rivers.
Calyculina occidentalis Prime. Creek, Boligee, Ala.
Sphaerium striatinum Lam. Big Black river, Durant, Miss.,
Boligee, Alabama, one specimen at last place.
Sphaerium stamineum Conrad. Young specimens, Grenada, and
one mature at Durant, Miss.
Pisidium virginicum Gmel. Durant, Miss.
Pisidium compression Prime. Tuscumbia spring; Black War-
rior, Tuscaloosa, and Shoal creek, Florence, Ala.
44 THE NAUTILUS.
Pisidium variabilis Prime. Big Black river, Durnnt, Miss.
Pisidium noveboracense Prime. Variety, spring, Tuscumbia, Ala.
Pisidium peraltum Sterki. Yalabusha river, Grenada, Miss.
Pisidium neglectum Sterki. Ms, name ; Black Warrior, Tusca
loosa, Ala.
Pisidium singleyi Sterki. Black Warrior, Tuscaloosa, Ala.
Pisidium atlanticum Sterki. Floating creek, Columbus, Miss.
(To be continued.}
A NEW SCALA FROM CALIFORNIA.
BY WILLIAM HEALEY DALL.
Scala lowei n. sp.
Shell small, conic, with five or more rapidly increasing whorls
after the (lost) nucleus ; color white, whorls very convex with deep
sutures and a small, spiral umbilicus ; there is no basal disk or cord ;
sculpture of about twenty-seven rather thick, strongly reflected,
smooth, close-set varices, and very close, fine, spiral threads, covering
the whole whorl between the varices, and separated by about equal
sulci ; aperture sub-circular, slightly higher than wide, the reflected
margin wide at the outer lip, patulous at the inner base, narrow
between the shoulder and the preceding whorl, and at the shoulder
produced into a short, rather stout spine which, repeated on successive
varices, coronates the whorls. Length of shell (without nucleus),
7.0; max. diameter, 4.0 ; max. diameter of aperture, 2.5 mm.
Habitat: dredged off Avalon, Catalina Island, Gala., in 40-60
fathoms, by H. N. Lowe, in 1903.
This species recalls S. bellastriata Carpenter, in general character,
but is smaller ; has more numerous, close-set and more strongly re-
flected varices ; and has the spiral striation and threading much
finer than in bellastriata of equal size.
The first shell belonging to Mr. Lowe was returned to him after
a figure had been drawn from the specimen. A second specimen was
sent by him, from the same vicinity, in 1906, which is retained in
the National Museum, No. 191548. This also has lost its nucleus.
The six remaining whorls measure 7.0 mm. in length.
THE NAUTILUS. 45
DESCRIPTIONS OF TWO NEW CLEOPATRA AND A PISIDITJM.
BY C. F. ANCEY.
1. Cleopatra multilirata, Anc.
Testa fusco-olivacea, indumento ferrugineo tecta, modice sed pro
genere aperte perforata, solidula, concolor vel circa umbilicum brun-
neo tincta,ovato-turbinata, liris acutis permultis (circa 8-9 in penul-
timo, 24-25 in ultimo anfractu) angulata. Spira conoidea, erosa.
Anfractus superst. 3^ rotundati, sutura profunda, ultimus latus,
ovatus. Apertura ovata, fere recta, superne subangulata, margini-
bus callo appresso nitido junctis, columellari vix patulo. Opercu-
lum fuscum, extus medio valde concavum, nucleo paucispirato, sub-
centrali, attamen propius marginem columellarem sito.
Long. 9, diam. 7, alt. apert. 5 mill.
Hab. Vinaninony, Madagascar (teste Ph. Dautzenberg).
A very distict species quite different from other forms recorded from
Madagascar, such as C. amoena, Morelet, C. moniliata, Morelet, C.
grandidieri, Crosse et Fischer and C. Trabonjyensis, Smith. All of
these but the latter were not correctly located in their proper genus,
as the first was referred to Melania, the second to Paludina and the
third to Paludomus by the describers. The present species is some-
what related to Cl. Trabonjyensis^ but the lirae are much more nu-
merous.
2. Cleopatra Smithi, Anc.
Testa ovato-acuminata, anguste perforata, subtenuis, flavescens vel
flavo-virens, epidermide tenui induta, liris fuscis infra peripheriam
ultimi anfractus evanescentibus regulariter exarata. Spira conoidea,
truncata. Anfractus circiter 7 convexi, sed haud tumido-ventricosi,
superiores, liris 4 cincti, concolores (lires haud coloratis), ultimus
ovatus, liris 5-6 aequidistantibus supra insertionem aperturae et ssepe
una infra medium cinctus et 2-3 parvulis circa perforationem sculp-
tus, inter liras striis increment! conspicuis eximie peraratus, zona
peripherica castanea, alteraque circa basin ornatus. Apertura ovalis,
superne et infra paulo acuminata, totius testae dimidio multo minor.
Peristoma tenue, marginibus callo tenui junctis, columellari simplici,
non expanso.
Long, (testse erosae, 4 anfr.) 11 diam. 7, alt. apert. 5, lat. ejusd.
31 mill.
Hab. River Chozi, which flows into the Chambe'zi, region of lake
Bangweolo, British Central Africa (Rev. Fath. M. Guilleme", 1905).
46 THE NAUTILUS.
Strikingly like Cl. Mweruensis, Smith (Proc. Zool. Soc. London,
1893, pi. lix, fig. 10), but smaller, differently colored and more regu-
larly lirate. The locality is very distant from lake Mweru, although
belonging to the same geographical region.
For this and numerous other interesting shells collected during his
journeys in Central Africa I am indebted to the Fath. M. Guilleme.
The material collected by him will be reported upon in the near
future. As a species of Cleopatra was already named after him, I
am respectfully dedicating this new species to E. A. Smith, Esq., of
the British Museum.
3. Pisidium planatum, Anc.
Concha minuta, lenticularis, parum inflata, compressa, nitidula,
tenuis, pallide cinerea, striis concentricis exilibus crebre sub lente
sculpta, umbonibus laevibus, depressis, inaequilatera. Pars antica
rotundata, postica fere duplo major, regulariter arcuata. Umbones
late depressi, obtusissimi, inermes, Icevigati. Dens cardinalis parvus,
laterales debiles. Ligamentum parvum, lineare.
Diam. 3£, alt. 2^, crass. 1^ mill.
Hab. Andriba, Central Madagascar (teste Dautzenberg).
A more depressed form than P. Madagascariense Smith, the
only other species of the genus described from the island.
NOTES.
ON THE HABITS OF PRATICOLELLA JEJUNA : — In the vicinity of
St. Augustine, Florida, I found this species locally only west of the
city in the white sandy, scrub oak section, near the pines, under boards
and leaves, in fact this was the only shell found there. They were
undersize, and very thin, probably due to the unfavorable condition
for molluscan life. Mr. A. G. Reynolds writes me from Disston
City, Hillsboro county, Fla. : " It is found under dead leaves and
trash, occasionally under boards, and more rarely on pine stumps."
Mr. Charles T. Simpson in his valuable paper, " Contributions to the
Mollusca of Florida," (Proc. Davenport Acad. Nat. Sci., v, p. 66,
1886), says : " The only mollusk ever found in high, dry pine woods
in Florida. I have seen it crawling on barren dry sand near
Braidentown, and around my dwelling, and along the road in
Fogartville." — C. W. JOHNSON.
THE NAUTILUS. 47
CHLORITIS EDWARDI, Gude, ascribed with some little doubt to
the Moluccas, is found in the Island of Ceram, where it was found by
the Rev. Father Le Coq d'Armanville, who lost his life in New
Guinea. My specimens were received from the Abb£ Mege,
Omphalutropis macromphala, Bavay (Journ. de Conch., 1905, vol.
liii, p. 250, figs. 9, 10, 11), is the same as 0. Qitndrasi, v. Moll.,
and therefore becomes a synonym of the latter. The shell is from
the Island of Guajan (or Guam), Ladrone Islands. Von Mollen-
dorff s paper appears to have escaped the notice of Mr. Bavay. — C.
F. ANCEY.
NOTE ON DTAKIA AND PSEUDOPARTULA — A recent article by
Lieut. -Col. H. H. Godwin Austen (Proc. Malac. Soc. Lond., vii, p.
93) has called my attention to the anatomical description of Dyakia
nasuta (Helix nasuta Metcalfe) by the same author (P. Z. S., 1891),
which I had OTerlooked when treating of this species in Manual of
Conchology, xiv, p. 12. It appears that the snail belongs to the
Zonitidce. There cannot be much doubt that the Sumatran and
Javan species forming the group Pseudopartula, and which I asso-
ciated with nasuta, will also be transferred to Zonitidce, since their
resemblance to nasuta seems too strongly marked to be adventitious.
— H. A. PlLSBRY.
A SYNONYM OF BRACHYPODELLA. — In my monograph of Uro-
coptidce I overlooked the name Microstoma hanleyannm Jousseaume.
Mem. Societe Zoologique de France for 1889, p. 247. This is
CyHndreUa hanleyana of Pfeiffer, a species of Brachypodella. The
name Microstoma will become a synonym of Brachypodetta. It was
probably not intended as a new generic name, being very likely an
error for Mychostorna — H. A. PILSBRY.
PUBLICATIONS RECEIVED.
HELICINA OCCULTA Say (Proc. Davenport Acad. of Sciences, ix,
pp. 173-180). ADDITIONAL NOTE ON HELICINA OCCULTA (Jour-
nal of Geology, xiii, 1905, pp. 232-237). By B. Shimek. The
rather extensive records of this species in both corichological and
geological literature are reviewed in these papers, with copious notes
and comparisons of recent and fossil shells. The alleged depauper-
48 THE NAUTILUS.
ation of loess as compared with modern shells is disproved; "while
there are variations in size in both recent and fossil shells, there is
no marked depauperation of the fossils." H. occulta is uniformly
found living on well-wooded grounds, leading Prof. Shimek to con-
alude that the forest areas in the loess region were formerly of much
greater extent.
ON THE ANATOMY OF CERTAIN SPECIES OF SOLENID^E. ON
THE ANATOMY OF ENSIS MAGNUS SCHUM. By H. H. Bloomer.
Journ. of Malacol., xii, pt. 4. Several oriental and tropical Ameri-
can species of Solenida are described anatomically in these papers.
Mr. Bloomer has now devoted ten papers to the morphology of the
Solenidtz, to the great advantage of all interested in Pelecypod
anatomy.
HENEY A. WARD.
We deeply regret to record the death of Professor Henry A. Ward,
who was killed by an automobile at Buffalo, N. Y., on July 4. He
was born at Rochester, N. Y., in 1834. Studied at Williams Col-
lege and Rochester University, and became an assisiant of Louis
Agassiz in 1854. He was Professor of Natural History at the
Rochester University from 1860 to 1865. From 1870 to 1900 Prof.
Ward traveled in all parts of the world, making large and valuable
collections which were shipped to his Natural History Establishment
at Rochester and from there distributed among the museums, uni-
versities, colleges and private collections throughout the country.
Prof. Ward has perhaps done as much to popularize and encourage
the study of natural history than any one man. Mr. Wm. T. Horna-
day paid him a glowing tribute, under the well earned title,
" The King of Museum-builders." It is probably safe to say that
through his agency more shells have been brought into this country
than by any other means. His " Catalogue of Specimens of Mol-
lusca" forms a useful hand-book for beginners. At various time his
Establishment has employed men who have subsequently become
prominent in various branches of the natural sciences.
For some years past Prof. Ward has devoted his energies chiefly
to collecting meteorites, bringing together the largest collection in
the world. An account of it may be found in the July number of
the Popular Science Monthly.
THE NAUTILUS, XX.
PLATE III.
AMPHIGYRA AND N EOPLANORBIS.
THE NAUTILUS.
Vor,. XX. SEPTEMBER, 1OO6. No. 5.
TWO NEW AMEBICAN GENERA OF BA80MMATOPHORA.
BY HENRY A. PILSBRY.
In the course of a collecting expedition to Alabama in October
and November, 1903, Mr. A. A. Hinkley found a number of mol-
Insks of unusual interest and importance.1 Among them were two
species of spiral-shelled gastropods quite unlike any fresh-water
snails hitherto known in this country, having affinities with the
Ancytidce.
Like Ancylus, the new forms are essentially sinistral in their soft
parts, but the shells are dextral. They are very small snails, and
were found on or usually under rocks in swift water, and evidently
do not depend upon free air for respiration, having a false gill in the
pallial chamber.
Genus AMPHIGYRA nov.
The shell is minute, Neritoid or Crepidula-like, with a small
depressed lateral spire, apparently dextral, composed of about l£
very rapidly enlarging whorls, the last very convex dorsally, im-
perforate ; the apex is smooth, and the last whorl is spirally striate.
The aperture is very large, oblique, transversely oval, the peristome
continuous and free, thin. Carity of the spire very small, a thin,
broad, concave columellar plate projecting across the end next the
spire, as in Orepidula or Latia.
The soft parts are sinistral, externally Limnseoid, with large
black eyes near the inner bases of the short blunt cylindric tentacles.
lSeo lists of the species collected, in NAUTILUS for August and September,
1904.
50 THE NAUTILUS.
Two adductor muscles, kidney-shaped in section, one on each side,
replace the usual columellar muscle.
The radula has 18, 1, 18 teeth, arranged about as in Lymncea.
Centrals with a single cusp, the laterals bicuspid, the outer cusp
smaller. The transition teeth have four or five cusps. The mar-
ginal teeth are low, wide and separated, with four or five cusps.
PI. Ill, fig. 6, teeth of A. alabamensis.
There is a short false gill hanging in the pallia! cavity.
The shell has some resemblance to Crepidula and Latia. Lepy-
rium and Pompholyx are also slightly similar; but the resemblance
is no doubt superficial. Pompholyx, like Amphigyra, is sinistral
with an ultrasinistral shell. It has no plate or lamina across the
visceral cavity. The soft anatomy of Amphigyra, so far as worked
out, seems to show most affinity with the Ancylidx.
The deck of Amphigyra is present at all stages of growth ob-
served, in young as well as mature shells. In Gundlachia no septum
is developed until a period of hybernation or aestivation is reached.
The shell is more solid than that of Ancylus or Gundlachia.
Amphigyra alabamensis n. sp. PI. Ill, figs. 1, 2.
The shell is shaped like a convex Crepidula, closely, finely and
sharply striate spirally, and of a pale yellowish-corneous tint. The
last whorl flares in a raised ledge at the baso-columellar region, the
back being very convex. The spire is slightly sunken, depressed.
The raised parietal margin of the lip is abruptly kinked where it
passes across the preceding whorl. The columellar plate or deck
extends over nearly one-third the total transverse length of the
aperture. Alt. 1.1, diam. 2 mm.
Wetumpka, Alabama, on the under surfaces of rocks in swift
water.
Genus NEOPLANORBIS n. gen.
The shell is minute, subdiscoidal, nearly flat above, convex below,
perforate, carinate at the periphery, composed of about two rapidly
enlarging whorls, the apex impressed and turned in. The aperture
is very oblique, wider than high, a little dilated at the base. Peris-
tome thin, not continuous, the columellar margin straight and
broadly dilated, somewhat thickened within.
The dentition and so far as known, the soft anatomy, is similar to
Amphigyra.
Type N. tantillus.
THE NAUTILUS. 51
The type of this group was at first taken for a Planorbis of the
P. dilatatus group,1 but when the dentition was examined it became
obvious at once that the snail could not belong to the Planorbinse.
The radula is not materially unlike that of Amphigyra, but the con-
chological features of the two groups are quite diverse, Amphigyra
being Crepiduloid with a distinct " deck " or columellar plate, while
Neoplanorbis is Planorbis-shaped, perforate, with a carinate periphery
and no " deck."
Neoplanorlis tantillus n. sp. PI. Ill, figs. 3, 4, 5.
Shell very narrowly perforate, slightly convex above, very con-
vex below, with a strongly projecting rounded keel at the periphery ;
light brown; surface slightly shining, sculptured with very obliquely
radial growth-lines and raised spiral stride, rather coarse for a shell
of this size. Whorls 2, rapidly enlarging, the apex somewhat
sunken; first whorl very convex, the second much less so, slowly
descending in front. Aperture very oblique, shaped like a gothic-
arched door, the upper and lower margins arcuate, the outer margin
angular, the columellar margin dilated, straight and vertical, with a
rather wide whitish callous within. Alt. .8, diam. 1.7 mm.
The specimens occurred at Wetumpka, Alabama with the preced-
ing species. This is one of the smallest fresh-water mollusks yet
found in America.
NOTE ON LEPYEIUM.
BY H. A. PILSBRY.
In connection with the genus Amphigyra I have had occasion to
examine the specimens of Lepyrium in the collection of the Academy.
It appears that the Lepyrium from the Cahawba river differs from
L. showalteri (Neritina showalteri Lea) of the Coosa, in being much
smaller, alt. 2.7, diam. 3 mm., with a straighter columellar edge and
without a raised outer margin of the columellar area. It will prob-
ably prove specifically distinct, but as I have been able to compare
only one of the original lot of L. showalteri, it may be best to rank
the Cahawba form ai a subspecies under the name Lepyrium showal-
teri cahawbensis.
1 It was reported as Planorbis tantillus in Mr. Hinkley'3 list, Nautilus XVIII,
p. 54.
52 THE NAUTILUS.
SOME SHELLS OF MISSISSIPPI AND ALABAMA.
BY A. A. HINKLEY.
(Concluded from August Number, p. 44-)
Family UNIONIDJJ.
Truncitta triquetra Raf. Tennessee river, Florence, Ala.
Truncilla brevidens Lea. Tennessee river, Florence, Ala.
Truncilla sulcata Lea. Tennessee river, Florence, Ala.
Truncilla haysiana Lea. Tennessee river, Florence, Ala.
Truncilla propinqua Lea. Tennessee river, Florence, Ala.
Truncilla personata Say. Tennessee river, Florence, Ala.
Truncilla biemar ginata Lea. Tennessee river, Florence, Ala.
Truncilla jlorentina Lea. Tennessee river, Florence, Ala.
Truncilla perplexa Lea. Tennessee river, Florence, Ala.
Truncilla turgidula Lea. Shoal creek, Florence, Ala.
Micromya cselata Conrad. Tennessee river, Florence, Ala.
Lampsilis excavatus Lea. Tallahatchie, Big Black and Tombigbee
rivers and Floating creek, Columbus, Miss.
Lampsilis ovatus Say. Tennessee river, Florence, Ala.
Lampsilis multiradiatus Lea. Tennessee river, Florence, Ala.
Lampsilis biangulatus Lea. Tennessee river, Florence, Ala.
Lampsilis hydianus Lea. Tallahatchie, Yalabusha, Big Black
and Tombigbee rivers.
Lampsilis affinis Lea. Tombigbee river and Floating creek,
Columbus, Miss.
Lampsilis orbiculatus Hildreth. Tennessee river, Florence, Ala.
Lampsilis anodontoides Lea. Tombigbee river, Columbus, Miss.,
and Yalabusha and Tennessee rivers.
Lampsilis fallaciosus Simpson. Tallahatchie, Yalabusha and Big
Black rivers in Mississippi.
Lampsilis rectus Lamarck. Tombigbee and Tennessee rivers.
Lampsilis lineosus Conrad. Tallahatchie, Big Black and Tom-
bigbee rivers.
Lampsilis concestator Lea. Yalabusha river, Grenada, Miss.
Lampsilis ogeecheensis Conrad. Tombigbee river, Columbus,
Miss.
THE NAUTILUS. 53
Lampsilis apicinus Lea. Tombigbee river, Columbus, Miss.
Lampsilis trabalis Conrad. Tennessee river. One specimen.
Lampsilis parvus Barnes. Tombigbee river, Columbus, Miaa.
Lampsilis cromwellii Lea. Yalabusha river, Grenada, Miss.
Lampsilis alatus Say. Tombigbee and Tennessee rivers.
Lampsilis purpuratus Lamarck. Tallahatchie, Yallabusha and
Big Black rivers.
Lampsilis gracilis Barnes. All the rivers visited.
Lampsilis leptodon Rafinesque. Tennessee river. Two poor
specimens.
Lampsilis claibornensis Lea. Floating creek, Columbus, Miss.
Medionidus acutissimus Lea. Tombigbee river and Floating
creek, Columbus, Miss.
Medionidus subtentus Say. Tennessee river, Florence, Ala.
Obovaria retusa Lamarck. Tennessee river, Florence, Ala.
Obovaria tinkeri Wright. Yalabusha river, Grenada, Miss.
Obovaria unicolor Lea. Tallahatchie, Yallabusha, Big Black and
Tombigbee rivers, very plenty in the last stream at Columbus, Miss.
Obovaria ellipsis Lea. Tennessee river, Florence, Ala.
Obovaria castanea Lea. Tallahatchie, Yalabusha, Big Black and
Tombigbee rivers.
Plagiola securis Lea. Tombigbee and Tennessee rivers.
Plagiola clegans Lea. Tennessee river, Florence, Ala.
Plagiola donaciformis Lea. Tallahatchie, Yalabusha, Big Black,
Tombigbee and Tennessee rivers.
Tritogonia tuberculata Barnes. Tallahatchie, Yalabusha, Big
Black and Tombigbee rivers. In the Black a few specimens were
found with purple nacre.
Cyprogena irrorata Lea. Tennessee river, Florence, Ala.
Obliquaria reflexa Rafinesque. Tombigbee and Tennessee rivers.
Ptchobranchus phaseolus Hildreth. Tennessee river, Florence, Ala.
Dromus dromus Lea. Tennessee river, Florence, Ala.
Strophitus spillmanii Lea. Yalabusha river, Grenada, Miss.
Strophitus tombigbeensis Lea. Tombigbee river, Columbus, Miss.
Strophitus elliotii Lea. Tallahatchie river, Abbeville, Miss.
Strophitus schafferianus Lea. Tennessee river, Florence, Ala.
Anodonta grandis Say. Tallahatchie and Yalabusha rivers.
Arcidens confragosus Say. Yalabusha, Big Black and Tombigbee
rivers.
54 THE NAUTILUS.
Symphynota costata Rafinesque. Tennessee river. Only dead
and poor ones found.
Symphynota complanata Barnes. Tombigbee river, Columbus,
Miss.
Margaritana monodonta Say. Tennessee river. Large numbers
were piled up by rocks, logs and in places on the river bank, where
the musk rats had opened them. The living ones were plen-
tiful under rocks in the shallow water, as many as two hundred
being found under one slab.
Unio gibbapus Barnes. Tombigbee and Tennessee rivers.
Unio crassidens Lam. Tombigbee and Tennessee rivers.
Unio arctatus Conrad. Tombigbee river, Columbus, Miss.
Unio tetralasmus geometricus Lea. Yalabusha river, Grenada, and
a rill near Durant, Miss.
Pleurobema hohtonensis Lea. Tennessee river, Florence, Ala.
Pleurobema edgariana Lea. Tennessee river, Florence, Ala.
Pleurobema tumescens Lea. Tennessee river, Florence, Ala.
Pleurobema dolabelloides Lea. Tennessee river, Florence, Ala.
Pleurobema decisa Lea. Tombigbee river, Columbus, Miss.
Pleurobema curta Lea. Big Black and Tombigbee rivers.
Pleurobema taitiana Lea. Tombigbee river near Boligee, Ala.
Pleurobema mix Lea. Tombigbee river, Columbus, Miss.
Pleurobema bulbosa Lea. Tombigbee river, Columbus, Miss.
Pleurobema nuculopses Conrad. Tombigbee river, Columbus, Miss.
Pleurobema aesopus Green. Tennessee river, Florence, Ala.
Quadrula asper Lea. Tallahatchie, Yalabusha, Big Black and
Tombigbee rivers.
Quadrula plicata Say. Tennessee river, Florence, Ala.
Quadrula perplicata Conrad. Yalabusha and Black rivers.
Quadrula latecostata Lea. Tombigbee river, Columbus, Miss.
Quadrula heros Say. Tallahatchie, Yalabusha and Tennessee
rivers.
Quadrula trapezoides Lea. Yalabusha, Big Black and Tombigbee
rivers.
Quadrula cylindrica Say. Tennessee river, Florence, Ala.
Quadrula tuetanevra Raf. Tombigbee and Tennessee rivers.
Common in botli streams.
Quadrula intermedia Conrad. Tennessee river.
Quadrula slopes Lea. Tombigbee river, Columbus, Miss.
THE NAUTILUS. 55
Quadrula rumphiana Lea. Tombigbee river near Boligee, Ala.
Quadrula pustulosa Lea. Tallahatchie, Yalabusha, Big BlacTi,
Tombiebee and Tennessee rivers. Those from tbe Yalabusha, Tal-
D
lahatchie and Big Black have few pustules, and occasionally one is
smooth.
Quadrula rubiginosa Len. Tallahatchie river, Abbeville, Miss.
Quadrula cermet Conrad. Yalabusha and Tombigbee rivers.
Quadrula hebetata Conrad. Big Black river, Durant, Miss.
Quadrula rubida Lea. Big Black riter, Durant, Miss.
Quadrula obliqua Lamarck. Tennessee river, Florence, Ala.
Quadrula solida Lea. Tennessee river, Florence, Ala.
Quadrula plena Lea. Tombigbee and Tennessee rivers.
Quadrula pyramidata Lea. Tennessee river, Florence, Ala.
Quadrula subrotundus Lea. Tennessee river, Florence, Ala.
Quadrula ebenus Lea. Tombigbee and Tennessee rivers. Common.
Quadrula granifera Lea. Tennessee river, Florence, Ala.
LYMNJEA DANIELSI SP. NOV.
BY FRANK COLLINS BAKER.
Lymnsea daniehi sp. nov.
Shell elongated, attenuated, of medium thickness ; color ranging
from very light-yellowish horn to dark chestnut or purple ; surface
shining, growth lines generally fine and close-set, but occasionally,
as near the aperture, coarse and often raised into ridges ; last whorl
often malleated ; spiral sculpture of fine impressed lines ; apex
smooth, spermaceti-white in color; whorls 7, flat sided or slightly
convex, slowly increasing in diameter ; spire long and sharply
attenuated, longer than the aperture ; sutures impressed ; aperture
ovate, often a little triangular, somewhat flaring ; peristome sharp,
simple, bordered by a dark purple or chocolate band in many speci-
mens ; parietal wall with a thin callus, generally closely appressed
to the shell and almost closing the umbilical region, leaving a small
chink; axis gyrate as in stagnalis, forming a heavy, ascending plait;
interior of aperture varying from white to dark purple.
Length 28.5, breadth 12, aperture length 14, breadth 6.5 mm.
56 THE NAUTILUS.
Length 27.5, breadth 11.5, aperture length 12.5, breadth 6.0 mm.
Length 29.01, breadth 11.25, aperture length 13, breadth 6.0 mm.
Length 25.5, breadth 10, aperture length 11.5, breadth 6.0 mm.
Length 23, breadth 9, aperture length 9.5, breadth 4.75 mm.
Length 20, breadth 10, aperture length 10.5, breadth 5.5 mm.
Types : Chicago Academy of Sciences, L. E. Daniels; cotypes,
Academy of Natural Sciences, Philadelphia; State Museum, Indian-
apolis, Ind.
Range : Lake Maxinkuk«e, Indiana, collected by Mr. L. E.
Daniels.
This distinct species may be known by its acutely pyramidal spire
and small, flaring aperture. It somewhat resembles elongate forms
of stagnalis. Some time ago Mr. Daniels sent six specimens of this
Lymncea for identification. It remained unnamed until a recent
visit to Washington, when a large collection of the same form was
seen which had been secured by Dr. Paul Bartsch, of the Smithson-
ian Institution, and its novelty was thereby established. Later,
Mr. Daniels sent several hundred specimens for examination. Lake
Maxinkukee also contains Lymneea palustris, desidiosa and parva.
The juvenile palustris are liable to be mixed with danielsi if care is
not taken to observe the form of the shell particularly the form of
the axis.
The material collected by Dr. Bartsch contains one monstrosity in
which the aperture is entire and separated from the body whorl.
The lot recently sent by Mr. Daniels is wonderfully uniform, show-
ing that the species is quite stable. A few curious variations occur
in which the spire is shortened and the aperture is enlarged and ex-
panded. In some specimens the columella plait is exaggerated to
such an extent that it forms a pseudo-tooth. In other specimens the
lower part of the aperture is somewhat effuse. The variation in
color is remarkable, and it would be of interest to know the cause of
the rich chestnut and purple color. The normal form is horn-col-
ored like stagnalis.
The species is named in honor of Mr. L. E. Daniels, of La Porte,
Indiana, whose assiduous collecting has added many new and inter-
esting shells to the fauna of the middle west.
THE NAUTILUS. 57
NOTE ON MITRA PICTA REEVE.
BY SLOMAN ROUS.
Mitra picta Reeve (Conch. Icon., p. 123, 1844) has been placed
by Tryon in the synonymy of M. barbadensis. This, I think, is an
error. A very cursory examination of the two shells will at once
show them to be distinct. The sculpture of barbadensis consists of
spiral raised lines, with fine microscopic, spiral and longitudinal
lines decussating the spaces between them. In picta the spiral lines
are impressed, closely, deeply foveolated, microscopically striated
longitudinally and pitted where they cross the revolving lines. The
fine spiral lines of barbadensis are absent in picta.
In color M. picta is a much darker brown, with the white patches
much more conspicuous, being larger and more numerous often in
wide longitudinal lines from end to end of the shell, but broken at
the suture, or often massed on the suture so that the white predom-
inates. I think usually one-fourth of the surface is white.
The folds on the columella of M. barbadensis are five in number,
sometimes a sixth being just perceptible, in picta there are only four,
the fourth being very small. I do not recollect ever to have noticed
a fifth fold, but if it occurs it will be as rare as six in barbadensis.
Tryon also says " Sowerby erroneously refers the latter (picta) to
the Cape of Good Hope." This is not an error of Mr. Sowerby.
I was resident in the Cape over forty years, during which I had the
pleasure of supplying him with many examples, and I have many
still in my possession, most, or I think all, of them collected in
Algoa Bay.
There can be no doubt that M. picta Reeve, is a valid species, and
that its habitat is the Cape of Good Hope. That other writers have
followed Tryon in this synonymy is sufficient reason for bringing this
error to the notice of conchologists.
NOTE ON VITREA APPROXIMA AND V VANATTAI.
BY H. A. PIL3BRY AND BYRANT WALKER.
Owing to the fact that the proof of the article in which these
species were described (Proc. A. N. S. P. 1902, pp. 431-3) was not
read by both us, some errors unintentionally crept in, which were
58 THE NAUTILUS.
not discovered until too late to be corrected in the original publi-
cation.
The localities of the two species were reversed and the proper
credit was not given for the discovery of V- approxima. The types
of that species were not collected by Ferriss and Walker and did
not come from Wilson's Cove. They were collected by Henry
Hemphill many years before and were distributed by him as " V.
wheatleyi Bid." As Hemphill's stay on Mt. Mitchell was very
limited (See Proc. P. A. N. S. 1900, page 149) they were no doubt
collected either along the trail to the summit or on Mt. Mitchell
itself. Collectors who received specimens from Hemphill as V.
wheatleyi would do well to reexamine them carefully.
V. vanattai was collected by Walker in Wilson's Cove.
SUBSPECIES, MUTATIONS AND FORMS.
BY T. D. A. COCKERELL.
Messrs. Pilsbry and Ferriss in discussing the variations of Bul-
imnlus alternates (Proc. Acad. Nat. Sci. Phila., 1906, p. 140)
say: " we can find no ground for dividing them [the different forms
of subsp. marice~\ into several races, as Prof. Cockerell proposes."
It seems opportune to protest against this interpretation of my
paper of 1891; the more so, since Prof. Pilsbry has before tried to
insist upon the erroneous nature of my proposed classification.
There is not, and never has been, the least divergence of opinion as
to the actual facts. In 1891, of course, we did not know as much
about the. subject as we do to-day, thanks to the excellent re-
searches of Messrs. Pilsbry and Ferriss; but now, as before, the
objections to my nomenclature have no valid basis, except in a mere
difference of opinion as to what should receive a name.
I grew up in England, accustomed to the methods of the Jeffrey-
sian school, which paid little attention to slightly differentiated races,
but bestowed names on all sporadic or individual variations which
were striking enough to be easily recognized. Professor Pilsbry
pays little attention to sporadic variations, but is very much in-
terested in geographical races or subspecies. It was undoubtedly a
THE NAUTILUS. 59
serious fault in the English work, that we were so little alert for
racial characters, and we were also to blame for confusing under the
name " variety " several different phenomena. I have watched the
development of the American researches with admiration, and cer-
tainly have no fault to find with the methods adopted; but at the
same time I believe we make a great mistake if we assume that
sporadic varieties are not worth noting and recording. Professor
Pilsbry will probably concede all this, in principle, but will object
to giving them names. It is unavoidable, I suppose, that there
should be differences of opinion about this; but it is certainly true
that only when they are named and find a place in the manuals do
they get properly noticed and recorded.
The best example of the English method which has so far appeared
is found in Taylor's " Monograph of the Land and Freshwater Mol-
lusca of the British Isles," now in course of publication. I can
hardly believe that any naturalist can study this work without ad-
miring its exhaustive treatment of variation, and perceiving the
value, from the standpoint of evolution, of the orderly presentation
of so great a mass of information. At the same time, it is not with-
out its faults, one of which is the confusion together of different races
and mutations, treating them all as " varieties," without discrimina-
tion. Thus under Testacella scutulum, the geographical form major,
from Algeria, appears as a variety; while its mutation albino, is
made to include all pallid forms, whether from Algeria or elsewhere.
It is evident that albina should be treated under major as a muta-
tion ; and the name pallida, proposed for the pale mutation of the
typical scutulum, should be so applied. Following the Jeffreysian
method, the term " monstrosity " is applied in an illogical manner to
sinistral mutations and various distortions of the shell, which have
nothing particular in common.
Deviations from the normal may be conveniently included under
three heads :
1. Subspecies, as generally understood in this country ; forms oc-
cupying a distinct territory (or it may be ecological position) of their
own, but intergrading with the species.
2. Mutations, the so-called spontaneous variations, arising sporad-
ically from some modification of the germ-plasm, and often persisting
in limited strains. These include, among other things, sinistral and
albino variations ; and it is desirable that analogous variations of
60 THE NAUTILUS.
different species should receive the same mutational name : e. g.,
simstrorsum for all sinistral mutations.
The interest attaching to these variations is very great at the
present time, when so much is being said about the theories of de
Vries and the Mendelian phenomena. We have, moreover, pretty
good evidence that such " sports " have given rise to genuine specific
characters, certain shells (e. g., Vitrea crystalline,} being permanently
" albino," while whole genera have become sinistral. The sinistral
mutations are perhaps of especial significance, inasmuch as in the
nature of the case they must arise " all at once."
3. Forms, resulting from the immediate effect of the environment
upon the individual, and presumably not inheritable in the proper
sense of the word. I notice that Messrs. Pilsbry and Ferriss, in the
work already cited (p. 169) name a " form " of Goniobasis.1 These
variations might be held to have the least significance of the three,
but they are often striking, and are usually of interest from a
physiological if not from an evolutional standpoint. It must be
remembered, also, that while they are said to be immediately due to
the environment, this is only true in a partial sense ; there must
already exist the inheritable power of responding in this particular
way to the conditions found. This power of response, as exhibited
under diverse environments, is itself a thing of great interest to
every biologist, and no one can doubt its significance in relation to
the persistence and spread of species.
The term " variety " may well be retained for use in cases which
cannot clearly be referred to one of the above categories. The
simple trinomial may be used for the subspecies ; in the case of the
others, " var.," " mut." or " f." should be placed before the last
name.
It should be added that not unfrequently " mutations " belong to
particular races, which are distinguishable only from their power of
producing them. Thus, I cannot see that the Cochlicopa lubrica of
the Rocky Mountains and of England are in any wise different
shells ; but in England certain mutations are frequently produced,
which are never seen in the Rocky Mountains.
1 Omitting, however, to name the very distinct mutations, figs. 27 and 28,
p. 167. According to the method I advocate, fig. 28 would be mut. spiralis,
and the same name would be applied to similar mutations of other species,
except when a different one had been earlier provided.
THE NAUTILUS, XX.
PLATE IV.
KELSEY: COLONY OF EPIPHRAGMOPHORA TUDICULATA.
THE NAUTILUS.
Voi,. XX. OCTOBER, 19O6. No. 6.
A HEALTHY COLONY OF EPIPHRAGMOPHORA TTJDICULATA.
BY F. W. KELSEY.
Wliile enjoying a short vacation, during the month of July of this
year, I sojourned with an old-time friend, Mr. I. J. Frazee, in
Pamoosa canon, about fifteen miles north of Escondido, in San
Diego County, and whenever I went prowling around among the hills
with my camera I always kept my weather eye open for anything
that carried a shell on its back.
My efforts were rewarded by a goodly find of Glyptostoma new-
berryanum Binney, and Epiphragmophora tudiculata Binney. The
latter, in one instance, were so plentiful as to deserve more than
passing notice, and I went to a good deal of trouble to enable me to
get a photograph to present to the readers of the NAUTILUS.
By means of bars, 1 moved a rock weighing over a ton, back from
the face of the cliff, so as to allow the light to enter and to give an
unobstructed view of the colony of helices hibernating in the lower
part of the cleft between the two masses of rock, and occupying a
space of not more than two square feet.
Of course, only a portion of the group shows in the picture, but
after making the exposure, I collected fifty-eight adult shells and left
more than half that number of immature ones for seed.
With one exception, I have never before seen snails in their wild
state so plentiful. This was in the summer of 1902 when I collected
over one thousand live Epiphragmophora stearnsiana Gabb, in a
space not over fifty yards square at Pacific Beach, near San Diego.
62 THE NAUTILUS.
EEMAEZS ON CEETAIN NEW ENGLAND CHITONS WITH DESCEIPTION
OF A NEW VAEIETY.
BY FRANCIS N. BALCH.
The recent announcement by Ball ['05 a. and b.] of a new Chiton
from New England — the first local addition to the group in many
years — makes this a favorable moment to introduce an allied novelty
with a few remarks.
Tonicetta Blaneyi Dall, was founded on a single specimen dredged
by Dwight Blaney, Esq., of Boston, off Ironbound Island, French-
man's Bay, Maine, in twenty fathoms.
Its diagnostic marks consist substantially in conspicuously radially
ribbed lateral areas and terminal valves.
Dall says it seems: " somewhat intermediate between Tonicella
and Trachydermon, with a leaning toward the former, while it
seems to be most nearly related specifically to Tonicella marmorea"
an opinion in which I concur.
But this " intermediate " character raises the whole question of the
true relation of these forms, which I now propose to discuss.
Trachydermon ruber and Tonicella marmorea are both common
north of Cape Cod in from five to one hundred fathoms; the former,
in my experience, outnumbering the latter (which favors the deeper
waters,) at least ten to one.
They much resemble one another and in fact Gould ['70] (who
retained them both in the Linnean genus Chiton) says of ruber: " It
is not difficult to distinguish at sight well marked individuals of this
species from those of (7. fulminatus " (i. e., Tonicella marmorea.)
" But there are intermediate specimens which it is not easy to pro-
nounce upon. In general, this species is smaller, more solid, more
convex, the valves more beaked, lines of growth more deep, the zig-
zag lines never appearing, though the posterior margin of the valve
is sometimes dotted with white and red. The impunctured or un-
granulated surface, however, is the best, as it is a constant, charac-
teristic."
But Gould's diagnostic points will not stand. The color-patterns
do duplicate, and the surface of ruber is punctate, as Dall has him-
self pointed out ['79]. I may add that in the coarseness of the punc-
tation the two overlap.
The fact is, as Gould indicates, that while the extremes of the two
THE NAUTILUS. 63
forms are well separated the series yet approach very near if they do
not overlap.
In spite of this the two are to-day almost universally placed by
American and British malacologists in two distinct genera — genera
which, in Pilsbry's monograph [Pilsbry '92] are rather widely sepa-
rated.
I propose very briefly to review their recent generic history.
Both were included in Gray's genus Ischnochiton, from which Car-
penter ['63] split off Trachydermon as a sub-genus, naming no type
though his own retiporosus was the first of the species referred to it.
Neither ruber nor marmoreus was mentioned. He later declared
it corresponded to the "second section" of Gray's Ischnochiton
« * * * scales of mantle minute, granule-like," and raised it to the
rank of a genus.
In 1873 [Carpenter '73] he published brief and informal, but im-
portant, notes of his observations on the Chitons of our New Eng-
land Coast made during work with our Fish Commission expeditions.
Of C. ruber he said : " It belongs to Gray's genus Ischnochiton,
* * * ' section f, mantle scales minute, granular ; ' but as the gill-
rows are short * * * it is necessary to establish a fresh genus,
Trachydermon * * *."
Of G. albus — now by far the commonest species all along our shore,
outnumbering ruber and marmorea combined a great many times
over and scarcely to be missed by any dredger — he said : " I twice
captured a live specimen ; but each time it eluded the aftersearch.
I do not doubt that this is also a Trachydermon * * *." While not
germane to the present inquiry this is interestingly suggestive of
changes in the CJiiton census.
Of G. marmoreus, he said : [it] " is Tonicia of H. Adams and
Gray, simply because the girdle is smooth. The true southern
Tonicise, however, have pectinated insertion-plates and ambient gills,
like the typical Chitons ; while the northern species so-called have
sharp plates and short gills. They differ, in fact, from Trachy-
dermon simply in the girdle being destitute of the minute scales. I
distinguish the group as ToniceUa."
This is the origin of the latter genus, which we now know is far
removed from true Tonicia.
Tonicella, then, was originally nothing but a scaleless-girdled
Trachydermon, with marmorea as its type-species. I have looked in
64 THE NAUTILUS.
vain through Dall ['79,] Fischer ['87,] Pilsbry ['92] and other
leading modern authorities for some other tangible distinction between
the two genera, but all the cited authors retain the two nearly in the
exact Carpenterian sense above given, though the single diagnostic dif-
ference results in their rather wide separation under Pilsbry's scheme.
The trouble with this single diagnostic difference is that according
to my observations it does not exist in the type-species, marmorea.
According to my observations neither ruber rior marmoreus has
scales, strictly speaking. In both (in ruber always, in marmorea
usually) the girdle is set with minute, stumpy spines. In ruber these
have some such shape as a thimble or short finger-cot and are thick-
set, giving a shagreen-like surface, while in marmorea they are pro-
portionately longer and thinner, commonly absolutely much smaller,
usually (if not always) much less thickly set, often very sparse
indeed and apparently sometines lacking entirely.
In other words my observations bear out the beautiful accuracy of
Sars [ '78,] Tab. 8, figs. 3 a — 1. and 1 a — 1., on the basis of which
he erected the genus Boreochiton expressly to contain our two forms,
in ignorance of Carpenter's prior Tracliydermon.
If Sars' figures and my observations are correct there is no more
ground for generically separating ruber and marmorea on the basis
of girdle-character [and no other has been suggested] than there
would be for putting No. 3 and No. 1 sandpapers in different genera.
If indeed the two species do not actually intergrade in this character,
at least the difference between the coarsest and finest (or naked)
marmorea is far greater than the difference between the coarsest
marmorea and the finest ruber.
The upshot is that I feel confident marmorea must be removed to
Trachydermon and stand close beside T. ruber. What is to become
of Tonicella after the removal of its type-species to a prior genus I
leave to others.
Admitting that ruber and marmorea stand close beside one another
in the same genus, how are they specifically distinguished ? I think
the answer must be — only by what Dall calls " the sum of the char-
acters." Gould's supposed diagnostic differences have been dealt
with already. Great as is the apparent difference between the rough
red-and-white striped narrow girdle of an extreme ruber and the
smooth, leathery, green, wide girdle of an extreme marmorea they
almost if not entirely overlap on this point also.
THE NAUTILUS. 65
The most reliable features known to me are the arrangement of
the notches and radial canals of the anterior valves (which in mar-
rnorea are confined to a central segment of say 100°, while in ruber
they spread through nearly 180°, the limiting pair being obscure) ;
and the form of the tegumentum of the mid-valves (which in mar-
morea has a rather sharply convex anterior edge, and in ruber con-
cave or sinuated). These characters are ascertained in so few speci-
mens that I am not sure how constant they are.
I have no doubt, however, that ruber marmorea, taking the "sum
of the characters," are quite valid species as such things go.
With these conclusions behind us, let us take up the new form* to
be discussed.
The affinities of T. Blaneyi may be analyzed as follows1: girdle-
covering, like marrnorea; girdle-color, like marmorea; punctation,
like ruber; notches and canals (anterior valve,) like marmorea; form
of tegumentum (mid-valves,) like ruber.
I may add that, examining large series for ribbed specimens, I
find distinct though slight " ribbing " of the anterior valve in several
specimens of ruber but none in marmoreal
What, now, is T. Blaneyi ? It may be pathological, but nothing
suggests it. It may be a unique specimen of an established distinct
species, but if so, since said to be conspicuous and occurring in much-
worked waters, it must be highly local or excessively rare. It may
be a " connected variant " of marmorea, but the intermediate stages
of " ribbing " are missing, occuring rather in ruber; it may be a
" connected variant" of ruber, but seems closer to marmorea; it may
be a "disconnected variant," "sport" or " mutant," which will
either become extinct or found a species and which merely happens
to come from the ruber end of the marmorea series. Finally, it may
be a hybrid, with the ruber tendency to wrinkling (possibly acces-
tral ?) intensified by the cross as is not uncommon in such cases.
My own provisional judgment would be for the latter.
While Mr. Blaney was working in Frenchman's Bay I was at
work (in the summer of 1899 and 1901) in the next bay west — Blue
Hill Bay. My richest ground was a small, stony area in about 12
fathoms off Harriman's Point where T. ruber was abundant while
1 Based entirely on Dr. Dall's descriptions and figures. The unique specimen
is in the U. S. Nat. Mus.
"Series much smaller.
66 THE NAUTILUS.
T. marmorea was rather uncommon — as indeed I found it through-
out. On this spot 1 dredged the form described below — one adult
and one half grown in 1899, and a young specimen in 1901.
Tonicella ruler (Lowe) var. index, nov.
In every respect typical except in color,1 which is in life an ivory-
white slightly tinged with green (taking on in alcohol or formol the
exact appearance of old ivory tinged faintly with yellow,) highly
polished and absolutely unmarked except for a few extremely faint
and minute light pink dots on the posterior edges of the mid-valves
and a very conspicuous dark purple elongated triangle pointing
posteriorly on the keel of the fifth valve. Girdle very pale, other-
wise typical.
A color variety connected with the typical form by intermediate
stages is perhaps not worth describing, but this form, while clearly
close to typical ruber, appears to be quite discontinuous. Dr. Dall
pronounces it new to him, which is practically to say new to science.2
In spite of the fact that the mark on the fifth valve is often recog-
nizable in typical specimens (and is probably present in all, though
disguised by the strong color-patterns) and in spite of the fact that
very pale, almost white, valves frequently occur and sometimes (as
Dall has noted) to the number of four or five in a single specimen,
yet the present variety can be distinguished from any other specimen
of ruber I have ever seen, across an ordinary room. It is not a mere
case of faintness of the rosy marking!?. The markings are wiped
out and a totally new color scheme (viz. green) introduced. My
three specimens, dredged two years apart, are absolutely indis-
tinguishable except for size. The first specimen dredged was adult
(probably hatched the year previous) the last very young (probably
breeding the next year.) It is, therefore, not connected with age
and has persisted on the same spot probably for at least four years;
but is not a true geographical race (in the sense, for instance in
which ornithologists use the term) since it is a cohabitant with the
typical form.
What, then, is T. ruber var. index? It may be pathological, or a
" physiological variety " — an albino in short — but it's local persist-
1 Radula not examined.
2 In this connection Chiton marmoreus var. cceruleus Winkley should be con-
sidered. See NAUTILUS VIII, p. 78, 1894.— ED.
THE NAUTILUS. 67
ence for at least four years looks otherwise. It is scarcely a
" variety " in the ordinary sense, for the intermediate terms are com-
pletely lacking. It is a " sport " or " mutant " in the sense of a dis-
continuous variant, breeding true and founding a new centre of varia-
tion ? To my mind the evidence, while wholly inconclusive,
suggests that possibility.
T. Blaneyi and T. ruber var. index, add two extremely interesting
problems to New England malacology-problems which are all the
more interesting because further work by Mr. Blaney or some other
fortunately situated student, should go far to yield a solution in the
course of a few years.
I may add that the feature from which var. index takes its name —
the striking dark backward-pointing triangle on the keel of the fifth
valve — suggests an interesting inquiry of a different nature. It ap-
pears to be a very wide spread and definite feature, appearing in
various species in various groups, now more, now less disguised by
conflicting color-patterns. It is by no means confimxl to the fifth
valve though commonest there. I find no notice of this odd Chiton
character in such examination of the literature as I have been able
to make. The fifth valve appears to have no specially significant
topographical relation with the internal anatomy, nor, indeed, is the
hollow of the keel in any of the valves occupied by any special organ
so far as I know. Has this mark, then, relation to some specializa-
tion of the complicated tegumentary system of organs?1
Jamaica Plain, Mass., September, 1906.
PAPERS CITED.
CARPENTER, PHILIP P.
[763] Supplementary report on the present state of our knowledge with
regard to the Mollusca of the west coast of North America. Rep. Brit.
Assoc. Adv. Sci., 1863, pp. 517-686.
['73] On the generic affinities of New England Chitons. Bull. Essex Inst.,
V. No. 9, pp. 152-154, Sep. 1873, (Reprinted in Ann. & Mag. Nat. Hist.,
4th Ser., v. XIII, pp. 119-123, 1874.)
1 Postscript. Since the above was written I have noted that in the ''List of
British Marine Mollusca" published in 1902 by a committee of the Concho-
logical Society of Great Britain and Ireland — apparently a very careful revision
in the course of which generic questions were well considered — T. ruber and
T. marmorea are, indeed, united in one genus, but that genus is Tonicella.
How this result was reached I do not know.
68 THE NAUTILUS.
DALL, WM. H.
[' 79] Report on the Limpets and Chitons of the Alaskan and Arctic Regions,
with descriptions of Genera and Species believed to be new. Proc. U. S.
N. M., pp. 281-344, pi. i-v., 1879. (Republished as Art. IV in " Sci.
Results of Exploration of Alaska.")
['81] On the genera of Chitons. Proc. U. S. N. M., pp. 279-291, 1881.
['05 a] A new Chiton from the New England Coast. Proc. Biol. Soc.
Wash, xviii, pp. 203-204, Sept. 2, 1905.
[ '05 b] A new Chiton from the New England Coast. NAUTILUS, xix. No.
8, pp. 88-90, pi. IV, Dec. 1905.
FISCHER, PAUL.
['87] Manuel de Conchyliologie, etc., Paris, 1887.
GOULD, AUGUSTUS A.
[ '70] Report on the Invertebrata of Massachusetts. 2d. ed. (Binney's)
Boston, 1870.
PILSBRY, HENRY A.
['92] Manual of Conchology, structural and systematic. Vol. XIV. Poly-
placophora. Philadelphia, 1892.
POSSELT, HENRY J.
['98] GronlandsBrachiopoder ogBloddyr. Conspectus Faun. Groenlandic.,
i, pp. i-xix, 1-298, pi. 1-ii and map. Copenhagen (?) 1898.
SARS, G. 0.
[ '78] Mollusca Regionis Arcticae Norvegias Bidrag til kundskab. om Norg.
Ark. Faun., I, pp. I-XVI, 1-466, pi. 1-34 and I-XVIII and Map. Chris-
tiana, 1878.
SOME OBSERVATIONS ON THE OVA OF UNIONIDJE.
BY L. S. FRIERSON.
When the ova of those species of Unionidse, with which I am
acquainted, are first noted in the marsupia, they are spherical in
shape, and consist of a single mass of yolk, surrounded with a clear
fluid, probably albumen, all confined in a spherical egg-shell mem-
brane. Soon after fertilization takes place, the yolk undergoes
segmentation, and goes through a " mulberry " stage, and finishes
with the familiar glochidium. Several interesting facts have been
noted, which may prove of interest. Some young ova of a Quadrula
trapezoides Lea were under observation, and being called oft' a short
time, on my return I noticed that some of the ova had changed their
THE NAUTILUS. 69
shapes in the interim. Some of these ova resembled dumb bells in
shape (except that the two balls were in close juxtaposition).
Close observation showed that on the side of the spherical vitellus
would appear a slight bump or projection. This would slowly grow
until it had become as large as the remainder of the yolk, thus pro-
ducing the dumb bell, apparently the fluid contents of the yolk would
pour itself from one ball into another, and a motion similar to that
of the well known Amoeba would result.
The other case was that of the ova of Anodonta imbecilUs Say,
which were in the mulberry stage. These were revolving about an
axis, making in some instances six complete turns per minute, while
the axis was fixed for each individual, the direction of this axis bore
no relation to anything that I could see, but were in every direction.
Some were horizontal, others perpendicular and many oblique.
This revolution was confined to the inside of the shell. The egg
as a whole remaining unmoved, neither the cause or object of this
rotation could be noted or conjectured by me. Possibly when packed
together in the marsupium, the motion would influence the shell, and
so in a measure prevent congestion in the narrow gill passages.
A solution of salt would in a minute or two cause a visible con-
traction of the vitellus, and the stoppage of the motion. Being
curious to know if alcohol would stimulate their motions I placed a
drop on the slide. In an instant an appearance resembling a violent
effervescence took place, and at its close every egg-shell had burst
open\ Apparently the endosmosis had so far exceeded the exosmosis
as to burst the membranes.
Frierson, La., August, 1906.
SPHJEKIUM HENDERSONI N. SP.
BY V. STERKT.
Mussel large, almost equipartite, well inflated ; superior margin
curved, with slightly marked rounded, or no angles where passing
into the supero-anterior and posterior slopes which are slightly
marked; scutum and especially scutellum distinct; anterior and
posterior ends rounded, the latter scarcely drawn downward ; beaks
near or in the middle in half-grown and adult specimens, markedly
70 THE NAUTILUS,.
anterior in the young, well prominent over the hinge margin, slightly
inclined towards the anterior, moderately large, rounded in the adult,
but with a small, more or less flattened area near the center, in some
specimens bounded by a slight ridge, most noticeable in the young ;
surface with moderately coarse, subregular sulcation, rather sharp to
shallow, rugulose, slightly shining, with one or sometimes two well-
marked, dark lines of growth ; color generally grayish in the adult,
lighter over the beaks and with light, yellowish zones along the mar-
gins, often with dark mottlings ; in some specimens there are irreg-
ular zones of bluish ; shell moderately thick, substance white, muscle
insertions distinct, those of the protractor pedis separated from the
anterior adductors ; hinge moderately stout, curved, formed rather
as in Sph. solidulum Pr., plate narrow, cardinal teeth short and
slight ; ligament rather long, covered, or a narrow median line un-
covered in old specimens.
Soft parts, in alcohol, pale, the syphons with a yellowish tinge ;
palpi large ; outer branchiae comparatively small ; metapodium dis-
tinct, but rather small, with the retractor pedis ; branchial cavities
with numbers of young at different stages, the largest 5 mill, long,
seems to be free in the cavity (not in a marsupium).
Size: long. 17.5, alt. 14, diam. 9.5 mill. (= 100:80:54.3); one
specimen 18 mill. long.
Habitat : Water holes, Crow Creek, 25 miles N. E. of Greeley,
Colorado, collected by Mr. Junius Henderson, curator of the museum
of the University of Colorado. There were over 120 specimens in
the lot, of all stages of growth, few of them full-grown ; some also
were in alcohol.
So far as can be judged, the present Spharium is distinct from all
described species, and except for the largest forms of S. simile Say,
is the largest. The mussel is higher than in simile, the beaks nar-
rower and more prominent, the shell and hinge are stouter, the sul-
cation coarser, and the color and surface appearance quite different.
There is a Sphcerium from several parts of the Mississippi valley,
somewhat smaller and more inflated, and partly of different outlines,
regarded as distinct since 1896, but shelved. It may be a form of
the present species, which was named in honor of its discoverer.
A few specimens were affected in the same way as those of S. sol-
idulum Pr. from Iowa, recently described in the NAUTILUS.
THE NAUTILUS. 71
NOTES.
POPULAR CONCHOLOGY — Karl Soffel, of Paris, the well-known
naturalist, has discovered that snails snore. He was experimenting
with several specimens, which he had placed in a glass jar in his
library, and one evening while writing he noticed a peculiar noise
issuing therefrom. It sounded like a person snoring in the next
room. M. Soffel approached and found that the snails were sleeping
soundly and snoring peacefully, the loudest snorer of them all being
the one that had lived among grape vines. — NEAT YORK AMERICAN.
PUBLICATION RECEIVED.
THE TERTIARY AND QUARTERNARY PECTENS OF CALIFORNIA.
By Ralph Arnold. (U. S. Geol. Survey, Professional paper No.
47, Series C., Syst. Geol. and Paleont., 76 — Pp. 264, plates LIU ;
plate I, a sketch map of California). Students of West American
Mollusks are greatly indebted to Dr. Ralph Arnold for his mono-
graph on this subject. In the introduction he says, of the first part,
it " Is a brief outline of the different Tertiary and Pleistocene for-
mations of California, giving the type localities, where, when and
by whom first described, their salient characters, where they and
their supposed equivalents are known to occur, the species of Pecten
found in them and their typical fauna as far as known." These
faunal lists will be very useful to students. The second part of the
work describes and illustrates the Tertiary, Pleistocene, and recent
Pectens, from Alaska to the Gulf of California, including the latter.
The classification follows Dr. Dall's " Tertiary Fauna of Florida."
The number of Pectens listed for California is 93, of these fifty
species and varieties are described as new.
The well-known form of Pecten ezquisulcatus Cpr. is figured as a
variety of Pecten circularis Sowerby, instead of P. ventricosus Sby.
Pecten hindsii Cpr. is listed as a var. of Pecten hastatus Cpr. rather
than hericeus, as lately listed, the latter now ranking as a variety
also of hastatus. Dr. Arnold writes the name as hericius Gould
rather than the better known form " hericeus." Dr. Dall's Ms.
name of Pecten (C.) hericius var. albidus and P. (Pseudamusium^)
randolphi Dall, var. tillamookensis, are two new varietal names of
Pectens known only among recent forms.
72 THE NAUTILUS.
The bibliography ranges from 1829 to 1905.
A list of bulletins issued by the U. S. Geol. Sur., under Series
C., Systematic Geology and Paleontology, is appended to this val-
uable monograph. — MRS. M. BURTON WILLIAMSON.
THE UROCOPTID MOLLUSKS FROM THE MAINLAND OF AMERICA
IN THE COLLECTION OF THE U. S. NATIONAL MUSEUM. By Paul
Bartsch. (Proc. U. S. Nat. Mus. 1906, pp. 109-160, 3 plates). This
paper deals chiefly with Mexican species, describing a number of new
species and subspecies, and several new subgenera of great interest.
The United States forms represented in the National Museum are
also reviewed, and two described as new: Holospira goldfussi
anacachensis, from the Anacacha Mts.,near Cline and Spofford, and
from near Eagle Pass, Texas. It differs from yoldfussi by the fewer
and stronger ribs. Holospira (Baplocion) tantalus, from ''some-
where in Arizona or New Mexico." No comparisons are given, but
it is said to be distinct from H. pilsbryi. It is what has been reported
as pilsbryi from the above territories. H. pilsbryi is to be removed
from the U. S. list.
The following errors or emendations require notice because they
enter into nomenclature:
Microceramus texasianus (p. 158) is used for M. texanus Pils.
Microceramus mexicauus (p. 159) should be credited to von
Martens, not Pfeiffer.
Ceelocentrum eiseni (p. 119) is used in place of C. eisenianum Pils.
If these changes are intended as corrections or emendations, that
fact should, we think, have been stated; but the introduction of new
forms or spellings of specific names on any grounds is to be deprecated.
-H. A. P.
NOTES ON WISCONSIN MOLLUSCA. By George H. Chadwick
(Bull. Wis. Nat. Hist. Soc., IV, pp. 67-99, 1906.) A valuable
annotated list containing 115 species, giving their distribution
throughout the State, bibliography, etc.
NOTES ON SOME LAND AND FRESH WATER SHELLS FROM
BRITISH COLUMBIA. By J. F. Whiteaves (The Ottawa Naturalist,
XX, pp. 115-119, 1906). An interesting faunal list containing
upwards of thirty species.
THE NAUTILUS.
VOL. XX. NOVEMBER, 1906. No. 7.
ON THE DISTRIBUTION OF HELIX HORTEN8IS MUELLER, IN NORTH
AMERICA.
BY CHARLES W. JOHNSON.
The object of this paper is to record in full our present knowledge
of tlie distribution of this species in North America, and to bring to-
gether the scattered literature bearing on the subject.
This species which presents such an interesting problem in geo-
graphical distribution was first recorded from North America in
1829 by a Mrs. Sheppard, of Woodfield, in an article " On the recent
shells which characterize Quebec and its environs." In this she
states that Helix hortensis is " found on the bank near the plain of
Abraham, common in the spring." In 1837 2 Amos Binney de-
scribed the "olivaceous yellow," non-banded variety as Helix sub-
globosa, stating that it " is common on the lower parts of Cape Cod
and Cape Ann, and is very abundant on Salt Island, a rocky unin-
habited island near Gloucester." This seems to be the first New
England record, for it is not mentioned by either John M. Earle or
Col. Jos. G. Totten in their lists of Massachusetts and New England
shells in 1833.3
Dr. A. A. Gould in 1841,* under Helix hortensis, adds to the
1 Transactions of the Literary and Historical Society of Quebec, I, p. 193'
1829.
J Boston Journal of Natural History, I, p. 488, pi. 17, 1837.
1 Hitchcock's Kept, on the Geology, etc., Mass., pp. 557, 559, 1833.
*Report on the Invertebrata of Massachusetts, p. 172, 1841.
74 THE NAUTILUS
localities above cited, "the region of Portland, Maine," and also
gives the St. Lawrence. In 1843 Dr. J. W. Mighels, in his cata-
logue of the shells of Maine,1 refers to H. hortensis as follows :
" Captain Walden has recently detected this species on a little island
in Casco Bay. It must be very plentiful as he obtained more than
five hundred specimens in a few hours. * * : Most of them have
five dark brown bands on a yellow ground ; some four, some two, and
a few none. Some have light fawn-colored bands on a white ground ;
in others, in place of colored bands the yellow ground is interrupted
with nearly transparent zones, one beautiful specimen is hetero-
strophed." The latter probably represents the mutation sinistrosum
of British authors, and seems to be the only American record. In
the same year James E. DeKay a under H. sulglolosa says: " I am
indebted to Col. Totten of the United States engineers for my ac-
quaintance with this species, which he found near the shore of the
St. Lawrence, two hundred miles below Quebec."
In 1851 s Amos Binney adds to his previous remarks on its geo-
graphical distribution, as follows : " It is also said to occur in the
northern part of Vermont, in Maine, Canada on the St. Lawrence,
Nova Scotia, and the islands of St. Pierre and Miquelon." In re-
ferring to changes which have taken place he says: "The prevalent
characters of this, and probably of other species in a given locality,
seem to undergo a considerable change from time to time. When I
first visited Salt Island, where the species abounds, ten years ago,
it was impossible to find a single specimen with either lines or bands.
One uniform color prevailed throughout. At the present time the
banded varieties are said to be not uncommon." To this Dr. Gould
appends the following note : " They have recently been discovered
by Dr. Samuel Cabot, in great numbers, on House Island, another
of the little islets in the vicinity of Cape Ann, where all of them are
of the banded variety. On the Outer Gooseberry, another neighbor-
ing islet, he found still another variety.
Mr. Samuel Tufts, Jr., in " A List of Shells Collected at Swamp-
scot, Lynn and Vicinity" (Proc. Essex Inst., I, p. 32, 1853), says :
1 Boston Journal of Natural History, IV, p. 332, 1843.
'Natural History of New York, Mollusca, Pt. V, p. 32, 1843.
1 The Terrestrial Air-breathing Mollusks of the United States (edited by A.
A. Gould), II, p. 112, 1851.
THE NAUTILUS. 75
"Helix hortensis may be added to this list on the authority of Wm. A.
Phillips, Esq."
Prof. Edward S. Morse in 1864, in his valuable paper, " Observa.
tions on the Terrestrial Pulmoniferaof Maine, etc.,1 places the species
under the genus Tachea with the following note : " This species has
been found in abundance on several islands from Casco Bay to Grand
Manan; Mr. Fuller found them on one of the extreme outer islands
of Casco Bay in great profusion."
In 1868 O. A. L. Mb'rch, under Helicogena hortensis in his paper
" On the Land and Fresh-water Mollusca of Greenland," ' says :
" Dr. Beck writes in his manuscript above this species : ' Wormskiold
has told me that he has found on the leaves of the small shrubs of
Salix lanatus in the vicinity of the interior of the Gulf of Tgaliko a
banded snail not unlike our garden snail. Two years ago (1844?)
I obtained a dead specimen from Greenland, probably introduced.'
It is certainly found in Iceland." See, also, Morch, "On the Land
and Fresh-water Mollusca of Iceland," page 42, of the same journal.
In 1869 W. G. Binney and T. Bland in " Land and Fresh-water
Shells of North America," 3 refer to the distribution of this species
as follows : " An European species introduced by commerce (?) to
the northeastern portion of North America. It is found on the
islands along the coast from Newfoundland to Cape Cod, and on the
mainland plentifully at Gaspe, C. E., also along the St. Lawrence,
Vermont (?), Connecticut (?), etc."
The above records give all we know of what may be called the
early history of this species in America, and clearly show that at
that time its distribution was practically as it is at present. The
following records are, therefore, probably only the results of more
careiul collecting over the same area. In taking up in detail the
distribution of H, hortensis, 1 will commence at the most southern
locality and go northward.
The Connecticut record is very doubtful, being based entirely on
J. H. Linsley's Catalogue of the Shells of Connecticut, 1845.*
r. Portland Society of Natural History, I, p. 10, 1864; also American
Naturalist, I, p. 186, fig. 16, 1867.
3 American Journal of Conchology, IV, p. 38, 1868.
3 Smithsonian Miscellaneous Collection, VIII, p. 181, 1869.
4 American Journal Science, XLVIII, p. 280, 1845.
76 THE NAUTILUS.
" Helix subylobosa (?) Binney, Weston, Gould, 172." The late
John H. Thomson, in his paper on •' The Land Mollusca of Bristol
County, Massachusetts," * says : " I have found large numbers of
the bright lemon-yellow variety of this shell on Martha's Vineyard
near Gay Head, never on the mainland. Of course introduced from
Europe. I tried some in my garden [New Bedford], but the slugs
Limax, Arion, etc., devoured the young, and I could never succeed
in getting matured specimens." Rev. H. W. Winkley informs me
that Rev. Geo. D. Reid has found it in the shell heaps on Martha's
Vineyard.
In 1870 Dr. H. A. Pilsbry records this species from Nantucket.1
They were taken by the late Dr. Harrison Allen at Sciasconset, the
four specimens showing the following variations : uniform yellow
(var. lutea), four and five bands, and one with five translucent almost
colorless bands. During the same year Dr. Benj. Sharp also found
it in the town of Nantucket, and later, on the adjacent island of
Tuckernuck, all of the uniform lemon-yellow variety.
In 1904 the writer reported the finding of this species at Chatham,
Mass.* Although I am aware that it was collected there long
before, it apparently has not been recorded. They were all very
pale yellow and bandless, the var. subalbida Locard. Some of the
older specimens had entirely lost their epidermis, and were chalky-
white, but still as active as their offspring. They were abundant on
the steep bluff in front of the light-houses, although I did not dis-
cover their presence until a rainy day brought them from their hid-
ing place ; not even a dead shell indicated their existence. Mr.
L. R. Reynolds informs me that he has collected H. hortensis on
Morris Island, a wooded island just south of Chatham.
There is a tray of specimens in the museum of the Boston Society
of Natural History from Provincetown, Mass., collected by Mr. L.
L. Thaxter. All are the bright yellow, bandless variety (lutea).
A few uniform light yellow specimens were found by Mr. A P.
Morse among some cedars near " Old Harbor," Cohasset, Mass.
The next localities to consider are those of the Cape Ann region.
W. G. Binney in his Manual of American Land Shells,* page 496,
1 Journal of Conchology (British), IV, p. 373, 1885.
»The NAUTILUS, IV, pp. 24 and 48, 1890.
1 The NAUTILUS, XVIII, p. 45, 1904.
* Bulletin, U. S. National Museum, No. 28, pp. 467 and 496, 1885.
THE NAUTILUS. 77
gives "Eagle Island, Marhlehead ; House Island, Manchester;
Kettle Island, Cape Ann." It is also to be found on the mainland
at Manchester, Magnolia, Gloucester and Rockport.
Prof. T. D. A. Cockerell, in a paper on " Helix hortensis in
America," * describes the following varieties from Magnolia : pallida
Ckll., " pale purplish or purplish-brown, without bands ;" quinque-
vittata Moq., " yellow with five bands ;" rufozonata Ckll., straw-
colored with red-brown bands, five in number ; also the varieties
subalbida, lutea and subglobosa. Later Prof. Cockerell, on the
" Variations of Helix hortensis at Rockport, Mass.," ' describes the
following varieties : sub glob o sa ; arenicola, "bands colorless, translu-
cent, 12345 ; " subalbida and lutea, the latter showing some fifteen
different band formulae including 00000, 12345, 123(45), (123)(45),
10345, 12045. The latter specimens were collected by Mr. G. H.
Clapp, who is quoted as follows : " The bandless or very faintly
banded forms were by far the most common. Typical shells were
scarce, even less plentiful than the transparent banded variety. On
rainy days the shells were out by the hundreds."
The first locality north of Cape Ann from which this species is re-
corded is some of the outer islands of Casco Bay. Mrs. A. S.
Packard has kindly donated to the Boston Society of Natural His-
tory a large series of this species from the collection of the late Prof.
Packard. These were collected on Brown Cow Island, Casco Bay.
The olive-green variety (subgloboscf) predominates, grading insensibly
icto the yellow (lutea)', the banded forms represent about ten per
cent, while a corresponding number could be referred to the variety
arenicola. Rev. H. W. Winkley has found it on Inner Green Island,
and Mr. W. H. Weeks, Jr., on Cliff or Crotch Island. In a letter
from Mr. H. K. Morrell, he writes : " I was at Seguin two days this
week and the keeper, Capt. H. L. Spinney, happened to say that
Helix hortensis was found on Pumpkin Knob, on the east side of
Sheepscot Bay, and was abundant on Matinicus." Mr. Owen Bryant
found the five-banded form on Seal Rock, one of the Matinicus
Group. Mr. Arthur H. Norton also reported it from Seal Rock, and
has found it on Mosquito or Little Egg Rock, Muscongus Bay. I
have a specimen in my collection labeled " Spruce Head, Me." Mr.
1 The NAUTILUS, III, p. 139, 1890.
a The NAUTILUS, XIII, p. 32, 1899.
78 THE NAUTILUS.
Morris Schick collected it at Bar Harbor. Mr. H. S. Colton and later
Mr. Dwight Blaney record it as " common" on Little Duck Island,
near Frenchman's Bay.1
It is also reported from Grand Manan, Halifax and Cape Breton,
while from the Gaspe region it has been collected by a number of
persons. Dr. John M. Clarke found it very common on the lime-
stone area at Perce. Mr. A. W. Hanham, in an article on the
" Mollusca of the Gaspe region,"2 says : "•Helix hortensis was very
common on the hillside, generally buried in the sand, several varie-
ties were taken, the plain form seemed to be the most abundant."
They were collected at Barachois, and in a beautiful series which he
sent to me at the time, I find several specimens in which the bands
are all united (12345). It has also been found at Gaspe Basin by
Mr. F. R. Latchford. Abbe Provancher lists it from the Province
of Quebec as " H. nemoralis." 3
Specimens from Wreck Harbor, East Cape, Anticosta. collected
by members of the " Arethusa" expedition in 1880, are in the col-
lection of the Boston Society of Natural History. In Newfoundland
the species seerns to extend along the greater portion of the west
coast. In a recent letter from Mr. L. P. Gratacap he says: "I
found H. hortensis in considerable numbers in Newfoundland, in
damp, woody ravines, in the two Codroy valleys, principally in the
Little Codroy. They were deep in color, with and without bands.
I did not see it on the east coast." Mr. Owen Bryant also failed to
find it on the east coast. Mr. G. H. Clapp*says: "A friend has
just brought me a living example of //. hortensis (12345) from the
headwaters of Robinson's River, west coast of Newfoundland."
This locality is not far above the Great Codroy. Mr. John Bryant,
Jr., found it at the Serpentine River, June 20, 1905, and Dr. John
Bryant at the mouth of East River, Hawkes Bay. During the past
summer (July 8, 1906) Drs. C. W. Townsend and G. M. Allen
also collected a beautiful series from the same locality on Hawkes
Bay. They found them only at the one place, feeding OH the " cow
parsnip." All were the bright yellow variety (lutea) with (12345)
and without bands.
1 The NAUTILUS, XVII, p. 99, and XVIII, p. 46, 1904.
J The NAUTILUS, VII, p. 65, 1893.
8 Le Naturaliste Canadien, XIX, p. 186, 1890.
*The NAUTILUS, XIV, p. 72, 1900.
THE NAUTILUS. 79
Dr. Wro. H. Dall in his work on Land and Fresh-water Mollusca,1
gives Labrador, but the exact locality is not recorded. Only the
single specimen referred to by Morch has been found in Greenland.
Dr. Dall thinks that it is doubtless an accidental importation. In
Iceland the species still exists, as will be seen from the following
letter from Mr. Gratacap : " I found hortensis this last summer in
Iceland, where it assumes a very dark tint, with the longitudinal
yellow threads strongly marked and the revolving dark bands re-
duced to one. They were fragile and very scarce. I have only three
from Seydisfiord, on the east coast."
The origin of this species in America has been the subject of con-
siderable discussion. Amos Binney, in describing subglobosa, and
comparing it with the European hortensis, and the possibility of its
being introduced by commerce, says : " It would be difficult, how-
ever, to account for their inhabiting the barren and retired situations
at the extremity of Cape Cod, and the rocky islands in the neighbor-
hood of Cape Ann, while in the intermediate country they are not
found." Later (1851) he believed its origin in this country to be
due to commercial intercourse with Europe. Prof. Edw. S. Morse
in 1864 (loc. cit.) asks this question : "If this species is really iden-
tical with the Tdchea hortensis of Europe, it seems a little singular
that here it should only be found on islands frequently barren and
far out from the land, * * while in the old country they become a
nuisance in gardens." This is only another way of saying that if this
species was introduced by commerce, why do we not find it in the
gardens of Portland, Salem and Boston, instead of on the barren
islands and exposed headlands ?
W. G. Binney has always questioned its introduction by commerce,
and later (1890) informs Prof. Cockerell (NAUTILUS, III, 139)
that he regards the species as naturally present in America, an
opinion in which Prof. Cockerell also concurs and refutes the theory
" that the hardy Norsemen of old may have carried the snails about
for food, and so imported it where they went." It seems too bad to
shatter this fabulous, though pertinent story, before the poet has had
a chance to versify. Dr. Pilsbry did not, however, coincide with
Prof. Cockerell's idea that it is a native American (see NAUTILUS,
IV, 24, 1890), and later in the Manual of Conchology (IX, p. 321,
1894), says : " H. hortensis inhabits many of the islands off the New
1 Harriman Alaska Expedition, XIII, p. 20, 1905.
80 THE NAUTILUS.
England coast, and being found in pre-Columbian kitchen-midden
deposits, cannot be regarded as a recent immigrant. Possibly it may
be the sole survivor of that Viking incursion in the eleventh cen-
tury." Rev. Henry "W. Winkley, in an interesting article, " Helix
hortensis in New England" (NAUTILUS, XVII, p. 121, 1904), sug-
gests that it is a survivor of the pre-glacial period. "When we con-
sider its present distribution and power to withstand even the
climate of Iceland, this seems to be the only solution of the problem,
which is further strengthened by its discovery in the Pleistocene by
Dr. Dall (loc. cit.), who says : " The wide distribution of the species,
often on uninhabitable islets off a coast little frequented, and its
presence, which I have verified, in the glacial Pleistocene of Maine,
tends to confirm the view that it is a prehistoric immigrant if an
immigrant at all."
The question which now arises is this : Why does Helix hortensis
continue to occupy the outer islands and headlands and not spread
further inland ? I think this can be quite readily explained, as far
as the New England coast is concerned, by taking into consideration
the geological character of the coast and the conditions most favor-
able to the life of land mollueks. The New England coast is com-
posed almost exclusively of granitic rocks, or, on Cape Cod, of sand
and gravel, both very unfavorable geological conditions for mollusca,
owing to the absence of lime ; even the more common species of the
eastern United States are comparatively few on the coastal area.
Again, the atmospheric conditions even slightly removed from the
outer shore line are very different from the islands and headlands,
continually bathed by the ocean's spray. These barren locations
make up in lime and moisture what they lack in vegetation; the
islets are also probably much more free from mice, rats and forest
fires, which on the mainland destroy large numbers of snails. Poly-
gyra albolabris and Pyramidula alternata are much more abundant
on the islands than on the adjacent mainland, nor do these remarks
apply only to the New England coast. The sandy pine and scrub
oak barrens of southern New Jersey are void of land shells, while on
the islands separated from the mainland by a wide expanse of salt
marsh, and numerous creeks and bays, Polygyra albolabris var.
maritima is quite abundant. The distribution of hortensis along the
broad estuary of the St. Lawrence is undoubtedly due to favorable
physical conditions.
THE NAUTILUS. 81
A LIST OF SHELLS FKOM NEBRASKA.
BY BRYANT WALKER.
The shells enumerated in the following list were all collected by
Dr. R. H. Wolcott, of the University of Nebraska, Lincoln, Neb.
Many of the localities represented are in counties not covered by
Aughey's List (Bull. U. S. Geol. Survey, 18, p. 697), so that while
not lar^e, the collection makes a considerable addition to our knowl-
D *
edge of the distribution of the different species in the State.
Dr. Pilsbry has kindly identified the Vertigo and Bifidaria.
Polygyra profunda Say. Omaha.
Polygyra multilineata Say.
Polygyra multilineata alba Witter.
Polygyra multilineata rubra Witter.
Omaha. A rather small, solid and elevated form. The four
specimens measure 2Qi x 15£, 20§ x 15, 20 x 15£, 20£ x 14 mm.
Bellevue. A single, small, globose, greenish-brown, unicolored
specimen measuring 16§ x 12^ mm.
Polygyra fraterna Say. Omaha. A small elevated, perforate
form varying from 12f-13^ x 4f-5^ mm. Salt Creek, Roca. Simi-
lar in shape, but rather larger, 13-14 x 5-5-^ mm.
Bifidaria contracta Say. Roca.
Bifidaria pentodon tappaniana Ad. Monroe Canyon, Sioux Co.
Vertigo ovata Say. Monroe Canyon.
Vitrea hammonis Strom. Roca.
Euconulus chersinus polygyratus Pils. Omaha, Roca and Monroe
Canyon.
Zonitoides arborea Say. Lincoln, Roca and Monroe Canyon.
Pyramidula cronkhitei anthonyi Pils. Monroe Canyon.
Vallonia parvula Sterki. Roca.
Succinea grosvenori Lea. Salt Basin, Lincoln. A small, rather
thick-shelled form, largest specimen 10x8 mm.
Hackberry Lake, Cherry Co. A large, thin-shelled form, quite
variable in contour, the two largest specimens measuring 14| x 10^,
aperture 7^x11 and 15^x10, aperture 7x10 mm. Immature
shells are very similar to those from Lincoln, but more inflated.
Succinea retusa Lea. Hackberry Lake and Marsh Lake, Cherry
County.
Succinea avara Say. Roca and Monroe Canyon.
82 THE NAUTILUS.
Lymnsea reflexa Say. St. Michaels, Buffalo Co., and Bellevue.
Lymnsea caperata Say. West Point, Cuming Co., and Dewey's
Lake, Cherry Co.
Lymnsea desidiosa Say. Monroe Canyon.
Lymnsea palustrts Mull. Dewey's Lake and Marsh Lake, Cherry
County.
Physa anatina Lea. Lincoln ; Washington, Cedar Co. ; Roca and
Platte River.
Physa gyrina Say. West Point, St. Michaels and South Bend.
Physa gyrina hildrethiana Lea. Lincoln and Hat Creek Valley,
opposite Monroe Canyon.
Physa gyrina oleacea Tryon. Bellevue.
Physa heterostropha Say ? Omaha. A few very young specimens
seem referable to this species.
Physa ancillaria Say. Hackberry Lake, Dewey's Lake and
" Sink " near Dewey's Lake, Cherry Co. Fine, large specimens.
Those from the u Sink " are elegantly striped like Lymn<za reflexa
zebra, a rare occurrence in this genus.
St. Michael. A more typical example.
Physa sayi warreniana Lea. Lincoln.
Physa Sp. ? Dewey's Lake, Cherry Co. A single specimen re-
sembling P. aplectoides Sterki, but with a short, obtuse spire and
well-impressed suture.
Aplexa hypnorum L. Dewey's Lake and Marsh Lake, Cherry Co.
Planorbis trivolvis Say. Bellevue, St. Michael, Lincoln, Hack-
berry Lake and Dewey's Lake, Cherry Co., Omaha and South Bend.
Planorbis bicarinatus Say. Sidney, Cheyenne Co. One example
is slightly transversely striate, approaching var. striatus.
Planorbis parims Say. Omaha and Lincoln.
Segmentina armigera Say. Omaha.
Ancylus shimekii Pils. Galloway.
Ancylus rivularis Say. Deadman's Run, Lincoln.
Ancylus tardus Say. Lincoln.
Ancylus Sp. ? Lincoln.
Amnicola limosa Say. Lincoln.
Sphterium simile Say. Lincoln.
Sphcerium stamineum Con. Salt Creek, Lincoln.
Sphcerium striatinum Lam. Lincoln.
Musculium jayanum Prime. Lincoln.
THE NAUTILUS.
Mmculium trantversum Prime. South Bend, Cass Co., and "Wood-
lawn Creek, Lincoln.
Musculium secure Prime. Lincoln.
Pisidium Sp. ? Omaha.
Lampsilis luteolus Lam. Blue River, Crete.
Lampsilis purvus Bar. Lincoln.
Lampsilis sitbrostratus Say. Lincoln.
Unio tetralasmus Say. Lincoln.
Anodonta grandis Say. Lincoln.
NOTES.
LYMNAEA MEGASOMA. This species is abundant in Cobalt Lake,
the centre of the wonderful silver-cobalt mines that now hold the
attention of the world. No shell is found in an environment of such
enormous qualities of precious mineral. Untold millions in value of
silver ores, running in many cases over $5,000 to the ton, surround
the placid lake, the waters of which are so strongly impregnated with
arsenic as to cause serious intestinal trouble to those who use it.
L. megasoma however lives and thrives, attaining large size and pre-
serving to a ripe old age the beautiful brown epidermis cha.'acteristic
of young shells. I have also found L. megasoma in Lake Temagami,
the terrestrial paradise of the Algonquins — the North West Arm of
Lake Nipissing, and at the mouth of French River. It doubtless
occurs in suitable waters across New Ontario from the Albany River
to Lake Abittibi. — F. R. LATCHFORD, Ottawa, Ont.
MIGRATION OF SCALLOPS. — The fact that beds of scallops in New
Bedford and Fairhaven waters and along the Cape shore are being
rapidly thinned out, and in some places completely exhausted, while
new beds are being constantly located about Nantucket, is alleged to
be causing the Massachusetts Fish and Game Commission to perfect
a system of procuring more definite data regarding the movements
of this toothsome shellfish.
Although the Nantucket fishermen will not commence dredging
before November 1st, they are keeping up a continual watch for new
scallop beds, and almost daily one of the boatmen will report large
beds of the shellfish on flats and shoals which only a week or so be-
fore were barren. And the old beds which give promise of such a
84 THE NAUTILUS.
rich harvest again this year are being watched with interest. The
scallops at Nantucket are in prime condition this year, and the
" eyes " are much larger than usual.
The Fish Commission experts are now desirous of obtaining defi-
nite data concerning the movements of scallops, and have liberated
in Nantucket harbor a number of the shellfish to which copper tags
have been attached. For the success of this experiment the com-
missioners look to the fishermen themselves, who are requested,
whenever a tagged scallop is found, to record the same to the experts
on their next visit to the island. It is known that the scallop is cap-
able of moving through the water quite rapidly by simply opening
and closing its shell, but how far it can go in a certain length of
time is what the Fish Commission wants to find out. Perhaps it
may develop through this experiment that scallops migrate from the
Cape shore across the sound to Nantucket. — (Boston Evening
Transcript.)
THE SENIOR EDITOR OF THE NAUTILUS, in company with Mr.
Jas. H. Ferriss, is on a collecting trip in Arizona. He expects to be
absent about six weeks.
PUBLICATIONS RECEIVED.
THE BEHAVIOR OF THE POND SNAIL. Lymnseus elodes Say. By
Herbert E. Walter (Cold Spring Harbor, Monographs, vi). An
interesting series of experiments and observations.
NEW MOLLUSCA FROM THE PERSIAN GULF, ETC. By J. Cosmo
Melvill (Proc. Mai. Soc., London, vii, p. 69). Thirty-one new spe-
cies of Gastropoda and one new Scaphopod are described and figured.
CAPULUS LISSUS SMITH, AS TYPE OF A PROPOSED NEW SUB-
GENL-S (MALLUVIUM) OF AMALTHEA. By J. Cosmo Melvill (Proc.
Mai. Soc., London, vii, p. 81). Under this subgenus is also placed
the Arnalthea benthophila Dall.
MOLLUSCA FROM THREE HUNDRED FATHOMS OFF SYDNEY. By
C. Hedley and "W. F. Petterd (Records of the Australian Mus., vi,
p. 212). About 100 species are recorded, of which 11 are new.
THE NAUTILUS.
Vor,. XX. DECEMBER, 1906. No. 8.
ABALONES AND THE PENAL CODE OF CALIFORNIA.
BY MRS. M. BURTON WILLIAMSON.
A rule, or tape-measure, is not considered necessary when collect-
ing shells along the beach, but if one would not pay too dear for his
shells on the Pacific Coast of North America it were well to take
some cognizance of the size of the Halioti detached from the rocky
coast, especially during certain months of the year.
Some years ago the writer presented to the American Association
of Conchologists a series of young Haliotis cracherodii Leech ; but
one would not dare to undertake it now that the law was passed pro-
hibiting the collecting of such shells ; for it is the young or small
specimens that the statute protects.
In the Penal Code of California, section 628, the law prohibiting
the collecting of abalones1 was incorporated and approved February
12, 1903, as an amendment to the section for the protection and
preservation of lobsters, crabs, etc. The law as then amended in
regard to abalones reads, "or any abalone shells or abalones the size
of which shall measure less than fifteen inches around the outer edge
of the shell." In 1905 this statute was amended, the size of the
shell allowed by law being three inches less in circumference for the
black abalone. It now reads as follows :
" Every person who between the 1st day of April and the 15th day
1 Monterey county appears to have taken the initiative in regulating aba-
lone-fisliing by passing an ordinance. See Dr. R. E. C. Stearns' article in
THE NAUTILUS for Nov., 1899, Vol. xiii, p. 81.
86 THK NAUTILUS.
of September of each year, buys, sells, takes, catches, kills or has in
his possession * * * any abalones or abalone shells of the kind
known to commerce as the black abalones (Haliotis Cu/ifornica),
the shell of which shall measure less than twelve inches around the
outer edge of the shell, or any other abalone shells, or abalones, the
shell of which shall measure less than fifteen inches around the outer
edge of the shell, is guilty of a misdemeanor."
Is this law enforced?
II. I. Piitchard, Deputy State Fish Commissioner, says : "Sec-
tion G28 has been enforced the same as any other section of the fish
and game laws."
In answer to my inquiry for THE NAUTILUS, he writes: " I can
only give you the names and dates of my personal cases : August 13,
1904, at Anacapa Island Bay, Webster of Venture, small shells,
fined $20.00, first offense. Same clay and place, Henry Ireland, of
Los Angeles, same fine. September 13, 1906, James Greenbeck, at
Redondo, small shells, fined $20.00. There has been a gang of
Japanese arrested near San Clemente Island, and again near Santa
Cruz Island and heavily find both times, but cannot give names and
dates; they used a diving suit and took 45,000 shells in 60 days."
Instead of mentioning " Haliotis californica" as the name of the
black abalone, our lawmakers would have clone better had they left
the shell with the familiar name of commerce instead of giving the
scientific one. It is to be presumed the name "Haliotis californica"
is intended for Haliotis californiensis? but this latter name belongs
to a rare variety of the black abalone (Hatiolis crac/tcrodii), not, to
my knowledge, collected north of the Mexican line. No concholo-
gist would testily that the common young black abalones were //.
californica or H. califormensis, but there is a saving clause in the
statute that would prohibit a culprit from escape because of the
technicality of a wrong name. It is this: " or any other abalone
shells, or absilones," evidently intended for the larger red and green
species (Haliotis nifescens, Haliotis fitlyens, etc.).
Abalone fisheries are confined to California. One is located a
few miles from Monterey, another at Whites' Point in Los Angeles
county. Dried ^abalones are prepared and shipped to China from
1 Dr. H. A. Pilsbry on Haliotis cracherodii var. Calif or nien sis Swains. THE
NAUTILUS, Vol. xii, no. 7. p. 79.
THE NAUTILUS. 87
San Diego. The Japanese are the men employed as divers. In
1900 the diving armor with air pumps came into use. Before that
the product from the Los Angeles county abalone fishery was in one
year, 1899, " GO, 000 pounds of dried abalones and 30,000 pounds of
shells of $7,800 value."1
It is said that the Chinese and Japanese are the consumers of this
white, boneless mollusk (the foot of the animal is the part used, and
this is white, no matter what the color of the shell may be); but I
have known of persons eating this shell-fish — not previously dried
as prepared for shipping — perfectly unconscious that the name given
to the soup, or fritters, masked the real one. Cans of daintily pre-
pared abalones, with another name, are among grocery supplies and
the contents of a can with butter, milk, salt and pepper, all heated
in a chafing-dish, is odorous and tender enough to please the palate
of any fish consumer.
Hollywood, Los Angeles, Cat., November 9, 1906.
NEW PISIDIA.
BY V. STERKI.
Pis. neglectum, n. sp. Mussel of small size, slightly to scarcely
oblique, rather rounded in outlines, moderately to rather well in-
flated ; superior margin slightly curved or nearly straight, with more
or less marked angles where passing into the anterior and posterior,
supero-anterior slope generally marked, slightly curved, anterior end
rounded angular, posterior part subtruncate, inferior margin moder-
ately curved ; beaks slightly posterior, rather narrow, rounded or
somewhat flattened on top, moderately projecting over the superior
margin ; surface with fine, subregular to irregular, crowded, con-
centric striae, microscopically rugulose, dullish or somewhat shining;
over the beaks there are much finer concentric strife to the center,
and the surface is not rngulose, more shining; color light to yellow-
ish horn to whitish, shell translucent to rather opaque, rather thin;
hinge rather slight, well formed, plate comparatively broad ; right
cardinal tooth rather short, strongly curved, not reaching the lower
. Kept. Fish Commission, 1901.
88 THE NAUTILUS.
edge of the plate, its posterior end much thicker, deeply grooved or
bifid ; there is an impressed groove between it and the edge of the
plate; left anterior strongly curved to sharply angular and bent up-
ward, the posterior oblique, curved ; '* lateral teeth " moderately
stout, the outer ones of the right valve distinct, the anterior left
cusp pointed, abrupt, the surfaces of all cusps rugulose ; ligament
rather strong.
Long. 3, alt. 2.6, diam. 1.8 in., types, and average.
Soft parts not examined.
Habitat : North America ; widely distributed and common in
Ohio, Michigan and Illinois, and probably over the Nearctic prov-
ince, seen also from New York, Canada and Alabama; living in all
kinds of waters. The specimens regarded as types are from a brook
at Krumroy, Summit Co., Ohio, 1000 (No. 3265, collection of
Cycladidoe).
This Pisidium was regarded as probably distinct years ago ; since
then the material has been looked over and compared dozens of
times. It seems to be the central form of a group which is one of
the most difficult to deal with of all our Pisidta. There are a host
of forms ranging under it or near it, really or apparently, and if only
a few specimens from various places were on hand, they might well
be regarded as distinct. But among the thousands seen, there are
more or less plainly intermediate forms. Some are still under doubt
and scrutiny, and waiting for additional material in order to ascer-
tain their true positions. Some forms have a slight resemblance
to some small forms of the polymorphous " abdtlum Hald.," others
to small forms of noveboracense Pr. It may also be said that the
more typical specimens resemble Pis. pusillum Gmel. more than any
other of our Pisidia does.
Var. (?) corpulentum n. Larger, more inflated, somewhat more
elongate and oblique, beaks generally somewhat more posterior,
large, prominent ; concentric striae coarser, surface more dull, color
whitish to yellowish, to grayish, and even bluish with lighter zones
along the margins.
Hab.: Grand River, Grand Rapids, Mich., collected repeatedly
by Dr. R. J. Kirkland in good numbers ; a similar Pisidium is from
Sillycash creek, near Joliet, 111., collected by Messrs. J. H. Ferriss
and J. H. Handwerk. It seems that these forms are connected with
Pis. neglectum, although the Grand River, Mich, form, is considerably
different and appears to be distinct.
THE NAUTILUS. 89
Pi's, trapezoideum St. (described in the NAUTILUS, IX, p. 124,
1896), from the middle Atlantic States, and probably the same from
Grand Rapids, Michigan, is very different from neglectum, yet there
may be intermediate forms connecting the two. Should that prove
true, trapezoideum would probably better be regarded as a specially
differentiated form, a variety of neglectum, and not vice versa.
Priority cannot be considered if there are better reasons for a differ-
ent point of view.
THE BROOKLYN CONCHOLOGICAL CLUB.
To the Nautilus :
The Brooklyn Conchological Club was formed over two years ago
by a number of collectors and admirers of shells. Several of the
members have extensive and valuable collections, the product of
many years of careful selection and world-wide correspondence with
collectors. THE NAUTILUS is a welcome visitor at the club. Two
of our members' names are found on its advertising pages, and
occasionally an article appears from the pen of our esteemed friend
and conchologist, Mr. Sloman Rons. We hope to hear of similar
clubs in other cities, and from individual collectors.
At the October meeting of the Brooklyn Conchological Club, Mr.
Maxwell Smith exhibited specimens of the shells collected by him
last summer in Otsego County, N. Y., also a map of the stations
visited. Mr. Smith gave an interesting account of his experience in
collecting. A list of the shells exhibited includes 54 species.
SILAS C. WHEAT.
SHELLS OF RICHFIELD SPRINGS, NEW YORK AND VICINITY.
BY MAXWELL SMITH.
The following species were all found in Otsego Co., with the ex-
ception of those from Weaver and Young's Lakes (Little Lakes),
Herkimer Co. The collection was made in July and August, 1906.
It is interesting to note that Lymnsea yracilts Jay was reported from
Canadarago Lake previous to 1870 by a number of authors. I
hunted diligently for this species but did not find it. From Allan
90 THE NAUTILUS.
Lake the water flows into Otsego Lake and a number of species
occur which are not found in Canarlarago. From the latter the
streams empty into the Susquehanna River and species finally inter-
mingle.
Amnicola limosa Say. In every lake and a large number of
streams.
Pomatiopsis lapidaria Say. Canadarago Lake.
Valvata tricarinata Say. Weaver, Canadarago and Allan Lakes.
Somatogyrus integer Sa}'. Sunset Creek.
Campeloma decisum Say. Canadarago Lake.
Omphalina fuliginosa Griflf. Sunset Hill, near Canadarago Lake.
Omphalina inornata Say. Sunset Hill.
Zonitoides nitidus Miiller. Woods near Richfield Spa.
Zonitoides arboreus Say. Sunset Hill.
Gfstrodonta ligera Say. Sunset Hill.
Pyramidula allernata Say. Everywhere with other species (one
albino farm).
Pyramidula striatella Anthony. East of Sunset Hill.
Polygyra tridentata Say. Sunset Hill, west of Canadarago Lake.
Polygyra Sayi Binney. Sunset Hill, east of Canadarago Lake,
near Oak Creek.
Polyyyra albolabris Say. Cooperstown, near Canadarago Lake,
Sunset Hill.
Polygyra palUata Say. Washed down Town Creek.
Polygyra tlyroides Say. East of Canadarago Lake.
Polygyra monodon Rack., \ar.fraterna Say. Woods near Rich-
field Spa.
Vallonia pulcheJla Miiller. In fields, everywhere.
Circinaria concara Say. Sun?et Hill, near Allan Lake.
Vertigo ovata Say. Under bark, near Richfield Spa.
Vertigo milium Gould. Near Sunset Creek.
Cocldlcopa lubrica Miiller. Cooperstown (Council Rock), Rich-
field Spa. (town), near Young's Lake.
Carycliium exigmim Say. Near Sunset Creek.
Succinea ovalis Gld. Near Canadarago Lake, near Oak Creek,
Cooperstown.
Succinea obliqua Say. Sunset Hill, in fields.
Succinea totteniana Lea. Near Allan Lake.
Succinea retusa Lea, var. peariensis Walker. Near Oak Creek.
THE NAUTILUS. 91
Lynwsea, stngnalis Linn. Canadarago Lake.
Lymiifp.a cohundlo. Say. Allan, Weaver Lakes.
Lynmcea palustris Miiller var. Canadarago Lake, Otsego Lake.
Fliysa ancillaria Say. Allan Lake.
Pliysa Iteteroslroplia Say. Canadarago and Otsego Lakes and
many si reams.
Planorbis fn'roh-i's Say. Allan Lake.
rianorbis dilatoius Say. Canadaraga Lake.
Planorbis rampanulatus Say. Canadarago Lake.
rinnorbis bicarinatus Say. Sunset Creek, Canadarago Lake.
Planorbis exacutus Say. Near Weaver Lake.
Plnnorbis deflectus .Say. Young's Lake.
Planorbis ulbns Miiller. Allan Lake, Otsego Lake.
Ancylus rioihiris Say. Oak Creek.
Anzylus tnrdus Say. Weaver Lake.
AUCI//US fuscus Adams. Allan Lake.
Ancy fits paralle.lus Hald. Town Creek.
Sphcerium simile Say. Sunset Creek, Town Creek, Allan Lake,
Weaver Lake.
Sphcerium rhomboideum Say. Allan Lake.
Spluerium striatinum Lam. Sunset Creek.
Plsidiutn ventricosum Prime. Canadarago Lake.
Pisidiitm novcboracensc. Prime. Canadarago Lake.
Lampsilis radiatus Gmelin. Canadarago Lake.
Slrophitus edentulus Say. Sunset Creek.
Ano'Ionta marghmta Say. Weaver Lake.
Alasmidonta undulata Say. Sunset Creek.
Unio complmtitus Solander. Oak Creek.
A NOTE UPON THE INSUFFICIENCY OF THE OFEKCULUM AS A BASIS
OF CLASSIFICATION IN BOUND-MOUTHED SHELLS.
I5Y L. P. GRATACAP.
In 1801 Lamarck established the genus Cyclostoma which he
made for the reception of a very miscellaneous and unclassified group
of shells, characterized however by certain common features as
the entire round aperture, continuous peristome, and operculum.
92 THE NAUTILUS.
Among this unassorted assemblage were placed marine, fluviatile
and terrestrial shells. Later Lamarck withdrew the marine and
fluviatile shells, and in 1819 he limited the genus Cyclostoma to
terrestrial species.
In 1829 the Rev. M. G. Berkeley described the anatomical struct-
ure of Cyclostoma elegans (Zoological Journal, vol. iv, p. 278), and
alluded to the operculum as " ovate spiral, calcareous." This de-
scription was, up to that time, the most extended and accurate that
had been published, of the soft parts of the animal of this group of
mollusca.
In the second edition of the Histoire NatureUe des Animaux sans
Vertebres, edited by Deshayes and Milne-Edwards (1838), the
diagnosis is Testa varia ; anfraclibus cylindraceis. Apertura circin-
ata, regularis : marginibus orbiculatim connexis, aetute pat&iti-reflexis.
Operculum.
In his observations Lamarck alludes to the similarly circular and
entire margins of the aperture of Pulndina, but remarks that in adult
cyclostomas the edge of the peristome is reflected, whereas in
Palud'ma and generally in fluviatile shells these edges are sharp and
plain.
The comments of the editors exhibit the diversity of views then
held by naturalists as to the affinities of these interesting shells;
some gave full weight to the fact of their aerial respiration and
grouped them in a special order, in which their manner of respira-
tion, their terrestrial habit, and the possession of an operculum
formed distinguishing or separative features; while others consider-
ing their respiration unimportant, dwelt upon morphological re-
semblances to Turbo, Ti'oc/tus and Scalaria. These resemblances
were the two tentacles, the absence of eye-stalks, the eyes placed at
the base of the tentacles, and the respiration anteriorly open. This
latter view was advocated by Cuvier. It was further emphasized by
their unisexual nature, and in the armature of their lingual ribbon,
which however rather coincides with that of pectinibranchiate gas-
teropods. Milne-Edwards and Deshayes regarded the cyclostomas
as terrestrial Turbos breathing air.
Amongst the forty-five species enumerated by Lamarck in 1838,
are representatives of the genera Pteroryclaa, Cyclophorus, Clioano-
poma,) Ttidora, Chondropoma, Helicina, Cyclvtns, Otopoma, Lepto-
poma, Truncatella, Megalomastoma, and Realia.
THE NAUTILUS.
Draparnaud had first separated the marine round-mouthed shells
from their supposed terrestrial congeners, which led Lamarck to
erect his genus Delphinula for the reception of some of the marine
forms, and later Puhidina for others, which genus was substituted for
his own Vivipara. Hdicina although proposed as a genus by Lam-
arck was not grouped by him near the cyclostomous genera. Its
operculiferous character was known, but in spite of this fact ita
position was assigned in the family of the Colimaces (Pulmonifera),
amongst the helices, bulimi and pupas. Ferussac had first recog-
nized that the genera Helicina and Cyclosloma were closely related,
and had, in deference to their similar breathing organization, placed
them at the end of the air-breathing gasteropods.
Reeve united Pnpina, Tntncatella, Cyclostoma, and Hdicina in the
single family Cyclostomacea. Menke as early as 1828 appears to
have separated the operculate shells into two families, typified by
Helicina and Cyclostoma. Dr. Gray (1842) first pointed out the
significant morphological distinction between Cyclostoma and He-
lict'nrt, and assigned to the family Helicinidae the three genera
Hdicina, Luci'dclla, and Alcadia, while Swainson (1840) had
grouped together Helicina Lam., Paclnjloma Swains., Oligyra Say,
Trochatella Swains., and LucideHa Swains. In the monograph
(1846) in Raster's Conchylien Cabinet upon "Die gedeckelten
Lungenschnecken," by L. Pfeiffer, the family Helicinacea was re-
garded as composed of the genera Trochatella Swains., Lucidella
Swains., Helicina Lam., and the family Cyclostomacea of Cyclostoma
Lam., C/toannpoma PIV., Cyclophorus Montf., Leploma Pfr., Megalo-
mastonia Guilding, Pupina Vignard, Cal/ia Gray, Pomalias Studer,
Aulopoma Trosch., Craspedopoma Pfr., Myxosloma Trosch., Ptero-
cyclas Bens., Acicula Hartmann, Geomelania Pfr., Hydrocena
Parreyss.
The generic divisions thus slowly evolved had been largely based
upon the characters of the opercula, and it seems that the credit of
emphasizing this feature was due to J. E. Gray, who in 1825 pub-
lished in the Zoological Journal and Philosophical Transactions the
results of his observations on their structure, formation and growth,
and insisted on their affording " characters for the division of families
and genera as the shell of the gasteropods themselves, and that to
neglect them in the description of the genus or species is quite as
rational as to describe only the single valve of a bivalve shell." The
94 THE NAUTILUS.
closing words of tliis quotation allude to Gray's opinion that the
operculum of the gasteropods was homologous or identical with the
second valve of a lamellibranch.
( To be continued.}
HELIX HORTENSIS IN NEWFOUNDLAND.
BY T. D. A. COCKERELL.
Mr. L. P. Gratacap has very kindly permitted me to examine a
dozen specimens of Helix hortemis which he collected at Little Codroy
river, Newfoundland, as reported in NAUTILUS, November, p. 78.
They are thin, and the dark bands when present are dull reddish-
brown, not black. The forms represented are :
(1) Clear yellow, handless = lutea Moquin. Two.
(2) Greenish-yellow, Landless = subylobosa Binney. One. This
seems to have been stained owing to the decay of the animal, and
may originally have been more nearly a pure yellow.
(3) Yellow, five-banded = quinquevittata Moquin. Five.
(4) Yellow, formula ( 123)45 = pauluccia Locard. One.
(5) Yellow, all the bands united = bouchardia Moquin. Three.
In the British museum there is an example of the variation
vallotia Moquin, from Labrador. It is yellow, with formula, O345.
Mr. Gratacap has also permitted me to see the shells collected at
Seydisfiord, Iceland, as reported in NAUTILUS, p. 79. They are
Helix arbustorum, rather thin, but otherwise typical.
NOTES.
OYSTERS CARRIED BY SEAWEED — Some time ago an oyster-
breeder in Morbilian, France, named Marline, called the attention of
the French Academic des Sciences to the appearance of unknown algae
that threatened to ruin the oyster-beds established at the mouth of
the river Va lines. These algae (which the breeders called ballons —
balloons) assume the form of little brownish-green leather bottles or
wineskins, which stick to the oysters, and which, microscopic at the
start, very soon reach the size of a large hen's-egg. Formed of a
very thin, elastic and rather frail coat, these bottles, usually full of
THE NAUTILUS. 95
water, fall in upon themselves at the moment of low tide. They be-
come empty then by the rents in their exterior; but, in virtue of
their elasticity, they fill up again with air. At the return of the
tide, they thus form a float more than sufficient to raise up the oyster
that serves them as support. Therefore at each great tide, when the
beds are wholly uncovered, the oysters are seen to disappear in the
olfing upon this automobile algae.
According to M. Bornet, we have the Colpomenia sinuosa, abound-
ing notably in the Mediterranean in the tracts adjacent to the At-
lantic. It was pointed out for the first time at Cadiz at the begin-
ning of the last century, and has never been seen farther north. It
no doubt came upon the hull of a vessel, and, having found in the
gulf of Morbihan suitable water, it multiplied there.
Hitherto no other eflf'eciive means has been found of combating
this a'ga than to sweep the beds with prickly fagots. It is to be
hoped that a rigorous winter will be sufficient to cause it to dis-
appear.— Scientific A mericun.
BRITISH COLUMBIA SHELLS. — Mr. Stewardson Brown, of the
Academy of Natural Sciences of Philadelphia, collected the follow-
ing species of land shells while on a botanical expedition in the
Canadian Rocky Mountains. At Field, B. C.: Tlnjsanophora in-
gersolli Bid., Vertigo goiddii Binn, Vertigo v. clulior Sierki, Vertigo
modestn Say, with the form parietalis Anc., Vitrea /tammonis Stiom,
Vitrea binneyana Mse., Eucomd'is J'ldms Drap., Zonitoidcs ar'jorea
Say, Pyramidula c. anthonyi Pi Is., Splnjradium edentidum Drap.,
Snccinea avara Say, and at Banff', Alberta : Vertigo modesta Say,
Vitrea biimeynna Morse, JEuconulns fidcus Drap., Pyramidula c.
anthonyi Pils., Splnjradium edentalum Drap — E. G. VANATTA.
ADDITIONAL LOCALITIES FOR HELIX IIORTENSIS — Since the
publication of my article on the distribution of Helix lioitensis in the
November NAUTILUS, I have received the following additional
records. From Curtain Island, Richmond Bay, Prince Edward
Island, a specimen of the plain yellow variety was received by Rev.
Henry AV. Winkley from a Mr. Ives. In 1902 Mr. Geo. H. Clapp
collected the banded variety (1234;")) at Cape Porpoise, about three
miles from Kennebunkport, Maine. Mr. Francis N. Balch has found
it at Orleans, Massachusetts, and also at Cohasset.
96 THE NAUTILUS.
I overlooked Mr. L. P. Gratacap's record of " Hare Island," St.
Lawrence River, given in his catalogue of the Bimiey and Bland
collection of the terrestrial air-breathing mollusks, etc. (Bull. Amer.
Mus. Nat. Hist., XIV, p. 39G, 1901). This species seems to be
more widely distributed than has been generally supposed. It is to
be hoped that these articles will lead to a still further knowledge of
its geographical distribution. — C. W. JOHNSON.
PUBLICATIONS RECEIVED.
STUDIES ON AUSTRALIAN MOLLUSCA, PT. IX. By C. Hedley
(Proc. Linn. Soc., N. S. Wales, pt. iv, 1905). Twelve new species
are described and figured, with critical notes on other species.
SOUTH AUSTRALIAN NUDIBRANCHS, AND AN ENUMERATION OP
THE KNOWN AUSTRALIAN SPECIES. By Herbert Basedow and
Chas. Hedley (Trans. Royal Soc., S. Australia, xxix, p. 134).
About 80 species are recorded, six new species described, and a
number of others redescribed and illustrated by 12 plates of beauti-
fully-colored figures.
REPORT ON MOLLUSCA COLLECTED BY MR. HERBERT BASEDOW
ON THE SOUTH AUSTRALIAN GOVERNMENT EXPEDITION 1903.
By Charles Hedley (Trans. Roy. Soc., S. Australia, 19«'5). A
valuable contribution to knowledge of the snails of the " Eremian
Region" of Central Australia. "Though quite unrelated to the
forms that people the arid regions of Asia, Africa or America, these
Australian shells ivpeat in their chalky texture and rough sculpture
the features of foreign species subjected to similar environment."
Several new species of Thersites and Xanthomelon are described, and
a beautifully-drawn plate illustrates the principal forms.
A CATALOGUE OF THE MOLLUSCA OF ILLINOIS. By Frank
Collins Baker (Bull. Illinois State Lab. Nat. Hist., vii, pp. 53-136,
Sept., 1906). An excellent catalogue, giving in full the distribu-
tion of each species throughout the State. The number recorded
is 241 species of fluviatile and 91 species of terrestrial mollusks. A
very exhaustive bibliography is also given.
THE NAUTILUS.
VOL. XX. JANUARY, 19O7. No. 9.
MOLLUSKS OF CAZENOVIA, N. Y.
BY JOHN B. HENDERSON, JK.
The village of Cazenovia, Madison Co., N. Y., about 20 miles
southeast of Syracuse, is upon the northern edge of the elevated
plateau of central New York. It is surrounded by hills more or less
wooded which in conformation form series of north and south valleys
of considerable depth, in one of which lies Cazenovia lake, with the
village at its southern end. At the altitude of Cazenova (1100 ft.)
the sedimentary rocks are shaly, but about two hundred feet below a
white limestone appears. The hills are grooved by deep ravines,
cutting many feet down into the limestone. These •' Gulfs " as they
are locally called, are heavily wooded, always damp, and offer excel-
lent collecting-grounds. Within a radius of a mile about the village
many types of station can be found with favorable soil, forest, stone,
and moisture conditions.
Messrs. Pilsbry, Walker, Clapp and the writer enjoyed a three
days' collecting trip through the Cazenovia valley, and the following
catch, remarkable in the number and quality of specimens, was made.
Comparatively little time was given to the lake.
Vallonia pulchella Mull. Polygyra thyroides Say.
" excentrica Sterki. " fraterna Say.
Polygyra tridentata Say (small). Strobilops labyrinthica Say.
" sayi Binn. Bifidaria contracta Say.
" albolabris Say. Vertigo pygmsea Drap.
" " dentutx (Walker). " gonldi Binn.
" dentifera Binn. " ventricosa elatior Sterki.
" palliata Say. Gochlicopa lubrica Miill.
98
1IIE NAUTILUS.
Circinaria concava Say.
Omphalina fuliginosa Griff.
" inornata Say.
Vitrina limpida Gld.
Vitrea cellaria Miill.
" hammonis Strom.
" indentata Say.
•' rhoadsi Pils.
Euconulus chersinus polygyratus
Pils.
Zonitoides nitida Miill.
". arbor ea Say.
" minusculd Binn.
Gastrodonta intertexta Binn.
" lie/era Say.
Ayriolimax campestris Binn.
" agrestis L.
Philomycus carolinensis Bosc.
Ariou fasciatns Nils, (very
abundant)
Pyramidula alternata Say.
perspectiva Say.
" striatetta Anth.
Helicodiscus lineatus Say.
Succinea retusa Lea.
" " pcoriensis Wolf.
" ovalis Say.
<k " var. (a striking
form, apparently
new).
" avara Say.
Lt'mnea reflex a Say.
Lhnnea catascopium Say.
" umbrosa Say.
'* emarginata Say.
" desidiosa Say.
" palustris L.
" humilis Say.
'' cohimella Say.
" owascoensis Baker.
Planorbis trivolvis Say.
" binneyiTryon.
" campanulatus Say.
" bicarinatus Say.
" deflectus Say.
'' pnrvus Say.
" exacutus Say.
Vcilvata (ricarinata Say.
Physa gyrina Say.
" srr^n Tapp.
" Integra Hald.
" heterostrjpha Say.
Aplexa hypnirum L.
Ancylus rivularis Say.
Pisidium variabile Prime.
Sphserium striatinum Lam.
" uccideiitale Prime.
" simile Say.
" fobale Prime.
" stamineum Con.
Calyculina securis Prime.
" transversa Say.
" partumeia Say.
NEW PISIDIA.
BY V. STERKI.
/Vs. superius n. sp. Mussel rather small, well inflated, ovoid in
outlines, rounded or subangular ; beaks rather posterior, broad,
rounded or somewhat flattened on top, projecting over the superior
THE NAUTILUS. 99
margin which is slightly curved, with slightly projecting, rounded
angles at the scutum and scutellum, the latter often covered in lateral
aspect, in full-grown mussels ; supero-anterior slope well marked,
slightly curved to nearly straight, posterior part subtruncate to
rounded ; young specimens are generally more angular in outlines
than adults ; surface with very fine, crowded, concentric stria? and
generally a few well marked lines of growth, somewhat shining ;
color light to deeper horn, generally with lighter zones along the
margins ; shell rather thin, more or less translucent ; hinge slight,
well formed, plate narrow, right cardinal tooth moderately to strongly
curved, its posterior part thick and grooved ; a deep, impressed
groove in the plate between its inferior edge and the cardinal tooth ;
left anterior strongly curved, posterior oblique, slightly curved to
nearly straight ; " lateral teeth " with short, slight, thin cusps, the
left ones and the right anterior pointed, the outer anterior of the
right valve distinct, the posterior quite small ; ligament rather slight.
Long. 3.2, alt. 2.8, diam. 2.2 m. (100 : 37.5 : 69) average.
Long. 3.5, alt. 3, diam. 2.4 m.
Soft parts not examined.
Habitat : Mountain Lake and Pine River, Marquette county,
Michigan, on the south shore of Lake Superior, Mono Lake, Musk-
egon county, Michigan, collected by Mr. Bryant Walker. A
Pisidium from the Isle Royale, in Lake Superior, and some speci-
mens from other places in Michigan, seem to range under the same.
Several hundred specimens at all stages of growth, collected in
Mountain Lake from 1895 to 1905, are remarkably uniform and
characteristic in appearance, although showing slight differences in
outline. There is none of the described species under which the
present Pisidium might be ranged, and it is similar only to tra/ie-
zoideum, being of about the same size and approximately the same
shape. But the latter mussel is much more angular in outline, its
supero-anterior slope is steeper, the anterior end is more angular
and more below the longitudinal median line, the beaks are narrower,
the surface more dull, and the hinge much stouter. Although
known for over ten years, this Pisidhmt has not been published
before, as I thought it might be connected with some other species.
Pis. succineum n. sp. Mussel of moderate size, moderately in-
flated, somewhat elongate, little oblique ; beaks slightly posterior,
rounded or slightly flattened, somewhat projecting over the upper
100 THE NAUTILUS.
margin, which is slightly curved, supero-anterior slope slightly
marked, curved to nearly straight, anterior end rounded angular,
posterior part subtruncate, inferior margin moderately curved ; sur-
face with fine to very fine irregular strife, shining, shell thin, trans-
parent to translucent, of wine to deep amher color ; hinge of the
same formation as that of P. noveboracense, but less curved and
slighter; ligament rather short and strong.
Long. 3.6, alt. 3, diam. 2 m. (100 : 83 : 5.5.5) average.
Long. 4, alt. 3.3, cliam. 2.4 in.
Soft parts not examined.
Habitat : North America, probably of wide distribution ; seen from
the Atlantic States from Maine to Virginia ; New York (near Mo-
hawk), Ohio, Michigan, Indiana ; common e. g. in the vicinity of
Washington, D. C., and Alexandria, Va., where it was collected in
1896 by the writer, and in the vicinity of New Philadelphia, O.
In shape it has a resemblance to P. noveboracense Pr., with which
it seems to range under the same group, but is smaller, the beaks are
less prominent, and it is at once recognized by its transparent shell,
its color and glossy surface. In the latter respect, it is also
similar to P. splendidulum St., but averages larger and its shape
is different. Like a number of other Pisidia, this has been a stumb-
ling-block for over ten years; but it seems constantly distinct, and
is of wide distribution.
Erratum. On p. 88 of the December number line 4 from bottom,
correct Silly cash to Lily cask creek.
LAND SHELLS FROM EAST SHORE OF CAYUGA LAKE.
BY SILAS C. WHEAT.
The following list of land shells were found by the writer in July
and August, 1905, on the east shore of Cayuga Lake, between Ilhaca
and Portland Point, N. Y., and between the lake and the top of the
cliffs :
Omphalina fuliginosa Griffith. Abundant, woods.
Omphtdina inornata Say. Common, woods.
Vitrea ferrea Morse. Rare, one found.
THE NAUTILUS. 101
Zouito/des nitida Miilier. Aliundant in low, wet fields.
Zonitoides arborea Say. Common.
Euconulus fulvns Miilier. Rare, found four.
Gastrodonta ligera Say. Abundant, woods.
Gastrodonta multidentata Binney. Rare.
Pyramidula alternatu Say. Abundant everywhere. One speci-
men is turreted, having a sharp angle at the periphery, and the
suture placed far below the angle.
Pyramidula striatella Anthony. Rare.
Pyramidula perspectiva Say. Rare, west side of lake.
Helicodiscus lineatus Say. Rare.
Polygyra tridentata Say. Common.
Polygyra sayi Binney. Rare, 8 on 1 sq. yd. No others found.
Polygyra albolabris Say. Abundant.
Polygyra albolabris dentota Walker. Rare.
Polygyra exoleta Binney. Rare.
Polygyra palUala Say. Common.
Polygyra thyroides Say. Common.
Polygyra Mrsuta Say. Rare.
Polygyra monodon Racket. Rare.
Polyyyra monodon fraterna Say. Common.
Vallonia pulchetta Miilier. Common.
Circinaria concava Say. Rare.
Piipoides marginata Say. Rare.
Vertigo ovata Say. Rare.
Vertigo ventricosa Say. Rare.
Cochlicopa lubrica Miilier. Abundant.
Bifidaria corticaria Say. Rare.
Bifidaria contracta Say. Common.
Bifidaria armifera Say. Rare.
Bifidaria pentodon Say. Rare.
Strobilops labyrinthica Say. Rare.
Succinea obliqua Say. Abundant.
Succinea obliqua totteniana Lea. Rare.
Succinea retusa Lea. Common.
Succinea avara Say. Common.
102 THE NAUTILUS.
A NOTE UPON THE INSUFFICIENCY OF THE OPERCULUM AS A BASIS
OF CLASSIFICATION IN ROUND-MOUTHED SHELLS.
(Continued).
BY L. P. GRATACAP.
The naturalists who subsequently worked upon the classification
of the mollusca were all sensibly influenced by these observations,
and the operculum became a diagnostic note in the separation of
genera. Troschel (1847) and Pfeiffer (1852), thoroughly applied
this method of discrimination, and augmented the number of genera.
M. Petit de la Saussaye (1850) questioned the significance of the
operculum as a real generic feature, and especially so far as the
nature of the substance of the operculum had been used by Pfeiffer
to distinguish genera. ' He says (Journal de Conchyliologie, Vol. I.,
1850) "the nature more or less calcareous or corneous of the oper-
cula arises rather from accidental circumstances, such as the habita-
tion of these animals, their nourishment, the character of the ground
upon which they live, the force of the sun, etc. As to the variable
form of the volutions of the spire, observed in these accessory parts,
it doubtless arises from the modifications, that the tissue secreting
them presents ; modifications which do not seem adequate for the
establishment of generic groups."
Benson reiterated the importance of the operculate features of the
round-mouthed shells, and the system of classification incorporated
in H. and A. Adams, " Genera of Recent Mollusca" (1858) still
further imbedded in the science the critical relations of the form
and substance of the operculum to the natural limitations of the
genera.
In the latest important systematic treatise, Tryon's Structural and
Systematic Conchology, the operculum is perhaps less exhaustively
used for descriptive purposes, but the author remarks that the oper-
culum " presents many beautiful modifications of structure, character-
istic of the smaller groups, which are often peculiar to limited
regions as in the Helicidae."
In an examination of this group of shells in the general collection
of the American Museum of Natural History and representing the
Jay, Haines, and Constable cabinets, the somewhat exaggerated im-
portance of the separative features of their opercula seemed apparent.
THK NAUTILUS. 103
It is quite evident that the opercula vary in substance and form ;
that these variations are related to groups and genera ; that in some
instances (Opisthoporus, Stoastoma) they present in one family a
unique and generally well-maintained uniformity of structure. But
their determinative character has been overestimated. The promi-
nent characters of the opercula are repeated in diverse sections of
the entire group, and the elements of their composition are more
nearly significant of habitat than they are of systematic differences
in the shells themselves.
It is noteworthy in this connection to recall the demonstration
made by Blanford (Annals & Mag. Nat. Hist., 1869), that the oper-
cula of Georissa (Blanford's genus) and Hydrocena Parreyss are
almost identical, whereas the former shell is entirely terrestrial, and
the latter marine.
(A note of importunity to traveling or exploring naturalists may
be appropriately inserted here. The character of the environment
of species or genera of land shells is generally not closely enough
observed. The moisture, temperature, mineral nature of the ground,
seasonal features, and frequency and amount of sunlight, the vege-
tation, are all. influential factors in the physiological life of shells.
They do, in land shells especially, have a clearly marked relation to
the coloring, thickness and operculum of the shells, and they are not
as particularly dwelt upon by collectors as they might be, both for
the purpose of elucidating the effect of environment and the some-
times misleading rules of systematists.)
Poey has called attention (Memorias sobre la Historic, Natural de
la hla de Cuba] to a striking illustration of local influence in the
sierras of the Isle of Pines. The Sierras de Casas and de Caballos
are respectively a short distance to the west and east of Nueva-
Gerona ; in the latter region the shells display regular spires, and
the crustaceans are destitute of spinose surface, as Trochatella stellata,
Pineria beathiana, Cyclostoma pupoides, among the shells ; Oniscus
sp.? among the crustaceans. In the former area the Trochatella is
represented by a denticulate, roughened, and loosely coiled shell
(constellata); the Pineria resembles a screw (terebra); the Cyclostoma
separates its last volution (moreletianum), and the Oniscus presents
spiny asperities, preserving otherwise a very similar appearance to
the de Casas species.
The geological formations are identical, and Poey has suggested
104 THE NAUTILUS.
the differing intensity of the sun's rays as a partial explanation of
these surprising differences.
This same author has protested against any sweeping conclusions
as to the importance in the operculate shells of the operculum, in es-
tablishing genera, and agrees with Petit de la Saussaye in consider-
ing this feature of quite subordinate value, remarking, " porque varia
el operculo de los Cyclostomas en los grupos mas aproximados por
el caracal y las costumbres del animal. No suce de asi en los tur-
bineos, entre los cuales se nota que los que viven apegados a los
arrecifes tienen el operculo corneo, y los que se apartan a mayor
profundidad lo tienen calcareo : de aqui un excelente caracter para
dif'erenciar las Litorinas de los verdaderos turbos y trocas."
Poey has also pointed out the confusion that results from the
adoption by different authors of varying or diverse morphological
characters for the classification of this group.
A Cyclostoma becomes a Chondropoma because of a corneous
operculum, or the same shell a Megalostoma by its thickened peris-
tome, a character participated in by the Cyclop/torus of Montfort,
and the Tropidophora of Troschel.
It is perhaps interesting to recall that Poey in a protest against
classification based upon the shelly parts of these mollusca, has
called attention to the method of progression of the Cyclostomas,
their divided pedal muscle allowing the movement forward of one-
half of the foot alternately with the other. It is not at all likely
that this feature would ever be useful in separating the round-
mouthed shells, but Poey has made of this observation an opportunity
to emphasize the preponderant significance of the physiology of mol-
luscs over the form, etc., of their shells for purposes of classification.
Cyclophorus has not a divided foot. Pfieffer, in his Monographia
Pneumonopomorum Viventium, analyzes the Cyclostomacea as com'
posed of three series, in each of which the form of the operculum is
regarded as the separative character. The entire systematic table
is as follows, so far as the opercula are here discussed.
FIRST SERIES.
Operculum circular, frequently concave exteriorly, formed of many slowly-
increasing whorls. Nucleus central.
I. Operculum shelly (testaceous).
Genera, Gyclotus, Alycseus, Diplommatina.
Oyclotus. Operculum orbicular, shelly, arctispiral, externally rather con-
THE NAUTILUS. 105
cave, with the margin of the whorl thickened, or raised in an incurved lamina,
nucleus subcentral.
Alycxus. Operculum circular, subtestaceous, obsoletely multispiral.
Diplommatina. Operculum subtestaceous, thin, with few whorls, outer edge
with thin lamella.
II. Operculum cartilaginous?
Genus, Pterocyclos. Operculum subcartilaginous, multispiral, interiorly
concave.
III. Operculum corneous.
Genera, Craspedopoma^ Aulopoma, Cyclophorus, Leptopoma, Meffalomastoma,
Calaulus, Pupinella, Pupina, Registoma, CalUa.
Craspedopoma. Operculum horny, solid, closely whorled. Nucleus central,
external lamina plane, internal furnished in its last whorl with a circular
prominence, centre profoundly concave.
Aulopoma. Operculum horny, arctispiral, planorbal composed of two laminae,
with an intermediate channel, with the last whorl furnished interiorly with a
circular groove. (The outer edge larger than the aperture and reflexed over
the peristome of the shell. — Adams.)
Cyclophorus. Operculum horny, thin, arctispiral, externally more or less
concave.
Leptopoma. Operculum membranous, arct'spiral, flat.
Megalomastoma. Operculum subcircular, thin, horny, arctispiral, planular.
Calaulus. Operculum orbicular, plane, corneous, arctispiral.
Pupinella. Operculum horny, arctispiral.
Pupina. Operculum thin, membranous, arctispiral, subplanum.
Rcgistoma. Operculum circular, thin, horny, arctispiral.
Callia. Operculum thin, membranous, arctispiral.
{To be continued.)
NOTES.
HELIX HORTENSIS ON MAGDALEN ISLAND — In 1901 the Car-
negie Museum of Pittsburg sent an expedition to the Magdalen
Islands and Labrador to collect birds and eggs, and on June 23d,
while at Grand Entry, Magdalen Island, one of the party collected
a lot of Helix hortensis Miill., which show a remarkable tendency
towards a single type of banding. There are about 330 shells in the
lot, counting young and old, and all are the form with yellow ground.
The following table, based on matured shells only, gives the number
of each form :
186—12345.
48— (12)345.
106 THE NAUTILUS.
1 — 1-2045.
22 — 00300. No. 3 very faint, translucent.
8—00000.
In a large proportion of the 186, 12345, bands 1 and 2 are very
close together, but separated by a distinct but very narrow light line.
In many of the (12)345 the union of 1 and 2 is indicated by a
lighter space in the center of the band. It will be interesting to get
further collections from this locality and see if the proportion of
(12)345 shells increases, as this lot seems to indicate.
The shells are rather thin and very uniform in size, the average
being about 18X15X14 mm. The largest shell measures
21 X 18 X 16^ mm., and in this shell bands 1 2 and 3 are unusually
wide. The smallest shell measures 16| X 14 X 12 mm.
Since my note in THE NAUTILUS, xiv, p. 72, I have received 8
additional H. hortensis from near the headwaters of Robinson's River,
Newfoundland, as follows : Four (12345), one 1(234)5, three 00000.
— GEO. H. CLAPP.
PUBLICATIONS RECEIVED.
MOLLTSCA OF THE SOUTHWESTERN STATES. II. By H. A.
Pilsbry and J. H. Ferriss. Proc. Acad. Nat. Science, Philadelphia,
1906, pp. 123-175.
This important paper completes the review of the southwestern
mollusca collected by the authors in Texas, New Mexico and Ari-
zona, with the exception of Omphalina and the Unionidse. Although
any attempt to a complete faunal monograph is expressly disclaimed,
this paper and its predecessor (P. A. N. S. P., 1905, p. 211) will be
found to contain practically all the reliable data extant upon the
fauna of southeastern Arizona and western Texas. The molluscan
fauna of this region, like the reptilian, shows that the transconti-
nental zones of distribution as established by Merriam cannot be
maintained in the Mediocolumbian region for these lower forms of
life, but that " these zones are secondary divisions of vertical life
areas of which the molluscan faunas were evolved in large part
independently."
While the range of practically all the Austroriparian species is
limited on the wpst by the rise between the 96th and 98th meridians
of longitude, marking the approximate limit of the Cretacious forma-
THE NAUTILUS. 107
tion and the Sonoran area in Texas, there is along its border an ap-
parent, rather than a real, intermingling of the fauna, owing to the
fact that the eastern species follow up along the river valleys, while
at the same time " the fauna on the bluffs or away from the streams
is frankly Sonoran." The scope of the present paper is, in the
main, restricted to the species of the Sonoran region, only such of
the Austroriparian species are mentioned as were actually collected
by the authors.
The list of the forms considered characteristic of the Sonoran
region in Texas includes twenty-one genera and thirty species. Of
the former, nine are not known to occur in the Austroriparian or
humid region of Texas. And of the latter, nine species and varieties
are described as new. The larger and more important part of the
paper is taken up with a monographic revision of the Texan Buli-
muli, which have so long been a source of exasperation and despair
to the American collector. It appears that the typical forms of B.
olternatus and schiedeanus do not, so far as known, occur in the
United States. The Texan series is restricted to B. dealbatus and
B. alternatus marise. The specific distinction of dealbatus and
alternates is clearly established by the details of the genitalia. The
typical form of dealbatus ranges from Alabama and Kentucky west to
Kansas. West of the Mississippi from southwestern Missouri
through Arkansas and the Indian Territory to Texas a number of
well characterized local races have been differentiated, of which six
are recognized and fully described and figured. B. alternatus marise
ranges through a wide territory along the Rio Grande from the Gulf
as far west as the Pecos river. Its exact western range has not been
determined. While there are some general differences between the
eastern (typical) and western forms, the intergradation is so com-
plete that the authors do not deem it advisable to attempt to separate
them into the distinct races.
Another important part of the paper is a revision of the genus
Helicodiscus, of which four species and one variety are recognized.
It is something of a shock, like that of losing an old friend, to learn
that lineatus of Say must be given up, but fortunately it is replaced
by another name, parallelus, of the same author.
Other corrections in nomenclature to be noted are, that Helix sayi
of Binney was preoccupied by Wood for an uncertain species of
Polyqyra, but probably auriformis ; and our well-known eastern
108 THE NAUTILUS.
species will be known as P. sayana Pils., hereafter, and that Pyra-
midula striatella Anthony for like reason must give way to P. cronk-
hitei anthonyi Pils.
Among the large number of new forms of land species described
the most remarkable is Bifidaria tuba, the type of a new submenus
Chaenaxis, characterized by its "large hollow axis, open below and
about one-third the total diameter of the shell," found in the drift of
the San Pedro river, Cochise county, Arizona.
Radiodiscus millecostatus from the Huachuca Mountains, Arizona
and Mexico, a minute Endodontid, is likewise both a new species
and the type of a new genus.
The fresh-water forms of the region are also fully discussed, and
afford several novelties in Planorbis, Vcdvata, Amnicola and Palude-
strina, all minute, and some of them the smallest species yet dis-
covered. The several varieties of Lymncea bulimoutes, — sonomuensis
Hemp., techella Hald., and cockerelli (new) — are fully differentiated
and figured, but unfortunately the typical form is neither figured nor
discussed comparatively.
In Segmentt'na, attention is called to the difference in the character
of the apertural lamellae in the North American forms included in
s. g. Planorbula Hald., and in the Antillian and Mexican group rep-
resented in our fauna by S. obstructa (Morel.). In S. armigera and
wheatleyi the lamellae, though differing in development, are " funda-
mentally identical." The Section Haldemanina recently established
by Dall (Alaska, xiii, 97, 1905) for the latter species would therefore
seem to be a synonym of Planorbula Hald.
The most important item in this portion of the paper is the dis-
covery of a species of Cochliopa ((?. riograndensis) in the drift of the
Rio San Felipe, Val Verde County, Texas. The occurrence of C.
rowelli Tryon, a Central American species, in California has always
been doubted. The present discovery confirms the genus as a mem-
ber of our fauna.
The authors intimate that their discussion of the southwestern
mollusks will be concluded by a third paper. Its appearance will be
eagerly looked for by all students of our North American fauna.
But, unless their well-known skill in the field has deserted them, it
is confidently expected that the expedition of 1906 to the Grand
Canyon will yield results quite as important as did their former ones,
and that the fauna of the southwest will in the near future be still
further illuminated by their labor?. BRYANT WALKER.
THE NAUTILUS XX.
PLATE V.
STENOTREMA AND PARAV1TREA.
THE NAUTILUS.
Vou XX. FEBRUARY, 19O7. No. 10.
NEW SPECIES OF STENOTREMA AND PAEAVITBEA FROM ALABAMA.
BY GEORGE H. CLAPP.
VITREA (PARAVITREA) ALDRICHIANA n. sp. PI. V, figs. 8, 9, 10, 11.
Shell small, widely, perspective^ umbilicate, flattened, slightly
convex above and below, the periphery well rounded ; greenish-
white almost transparent, highly polished with very faint growth
lines and on the body -whorl a few impressed lines spaced irregularly.
Whorls five, those of the spire rounded at the well-impressed suture;
umbilicus contained about three times in the diameter of the shell
and showing all of the volutions ; peristome forming two-thirds of a
circle ; lip simple.
Slightly below the periphery, and in the last half of the body
whorl are about four teeth which are longer, vertically, than wide,
and, apparently, project at both the upper and lower ends; i.e.,
they are double pointed.
Greater diam. 2, lesser 1.9, alt. 1 mm.
Type from the slope of the Cumberland Plateau in Jackson
County, Ala., close to the state line, and about 2 miles S. E. of An-
derson, Tenn. Collected by Herbert H. Smith, who found but four
specimens of this excessively rare species, the others being from
" Buck Creek Cove," Franklin County, Tenn., " Cove in Valley of
Little Crow Creek," and " Bennett's Cove, near State Line," both
Jackson County, Ala.
These four localities are near together on the Cumberland Plateau,
and along the Tenn. -Ala. boundary.
It is interesting to note that Mr. Smith collected both V. mufti-
110 THE NAUTILUS.
f/entata and lamellidens in the same general locality, although lie did
not find the latter in Alabama.
The species is so distinct from all others of the genus that it can
not be compared with any of them. The fact of finding four speci-
mens at different places and all of practically the same size, shows
that it is not likely to be the young of a larger species.
I take great pleasure in naming it after Hon. T. H. Aldrich so
well-known by his work on the fossil mollusca of the South.
Type in my collection ; the other specimens in the collections of
Bryant Walker, John B. Henderson, Jr., and T. H. Aldrich.
POLYGYRA (STENOTREMA) BREVIPILA n. sp. PI. V, figs. 1, 2, 3, 4.
Shell imperforate, globose, thin, light reddish-horn color ; densely
hirsute with fine, short hairs. AVhorls five, those of the spire convex
with a well-impressed suture ; the body-whorl very convex, equally
rounded above and below, deeply impressed in the umbilical region,
abruptly deflected at the aperture and contracted behind the lip.
Aperture transverse, narrow, widening anteriorly; parietal tooth
large, strong but narrow, erect, with the sides nearly at right angles
to the whorl and projecting beyond the lip, with which it is not
parallel, but diverges for three-fourths of its length, when it is abruptly
bent inward and downward, terminating opposite the second notch
in the lip, its distal or outer extremity connected with the end of the
peristome by a ridge of callus, the axial end sweeping around and
"pocketing " the basal end of the lip ; outer lip reflected back against
tha body-whorl, but with its sharp edge free from the whorl its entire
length; very much thickened along its inner edge which forms a
raised margin around the wide notch, and a well-developed tooth or
fold beyond it, after which the margin is incurved around the outer
extremity of the parietal tooth. Fulcrum long.
An average shell measures, diam. 8^, alt. 6 mm.
The largest seen measures 9x6, and the smallest 7| x 5| mm.
Collected by Herbert H. Smith on Horseblock Mountain, Talla-
dega Co., Ala., at an altitude of about 2,000 feet. On some of the
U. S. Geological Survey sheets this mountain is called " Talladega,"
but " Horseblock " is the local name.
" The mountain sides, near the top, are littered, or rather piled
with talus, big and little rocks ; the shells are found almost invari-
ably on the lower sides of these rocks, and generally they chose
THE NAUTILUS. Ill
the biggest and heaviest ; you turn over perhaps fifty to find one
shell. They are obtained by sheer hard work." H. H. S.
This is one of the most striking species of the Stenotrema hirsuta
group, and differs from all others in having a continuous free lip ; the
lip-notch is also of an entirely different type, as it will be noticed by
a reference to the figures that the notch is not an indentation in the
center of the lip, but the lip forms a curve from the columellar end
to the highest part of the notch when it is suddenly deflected and
bent forward. The parietal tooth differs in being more perpendicular
to the body-whorl, not bent toward the Up, as in Stenotrema, where
the tooth is very convex on its outer side and concave on the side
next the lip, which it frequently overhangs. On looking into the
aperture, through the notch, the whole of the long fulcrum can be
plainly seen.
The only species with which this is likely to be confused is P.
altispira Pils., but the resemblance, due to the wide notch in each
case, is only superficial ; for, as pointed out above, the notch is of a
different type. The hairs are also much finer, closer and shorter,
and the shape of the shell is entirely different.
Type in my collection ; cotypes in the collections of the Academy
of Natural Sciences, Phila., Bryant Walker, T, H. Aldrich and
John B. Henderson, Jr.
Reference to Figures.
Figs. 1, 2, 3, 4. Polygyra (Stenotrema) brevifnla Clapp.
Figs. 5, 6, 7. Polygyra (Stenotrema'} altispira Pils. Roan Mtn.,
N. C., specimen 9x6 mm., size of type.
Figs. 8, 9, 10. Vilrea (Paravitrea) aldrichiana Clapp. Type.
Fig. 11. Apparent shape of teeth of V. aldrichiana.
A NEW CAEDIUM FROM PTJGET SOUND.
BY WILLIAM HEALEY DALL.
Among the specimens of Cardium received from the Pacific coast
during a series of years was occasionally a specimen with more num-
erous ribs, smoother surface and small, compact shell, which was
doubtfully referred to the young of C. californien.se Deshayes, better
known under Gould's name of blandum. Recently in an endeavor
112 THK NAUTILUS.
to identify one of these shells, sent in from Monterey by Mr. Berry,
all the available specimens were brought together and carefully com-
pared, leading to the conclusion that the shell is a distinct species.
Cardium ( Cerastoderma ) fucanum n. sp.
Shell small, plump, compact, solid for its size, covered with a
smooth brown or gray periostracum usually more or less eroded ; form
varying from nearly circular to slightly ovate, with rather prominent
slightly prosoccelous beaks, situated a little anterior to the middle of
the shell ; valves moderately convex, equal and nearly equilateral,
sculptured with numerous (40 to 58) similar, small, low, flatfish radial
ribs separated by much narrower interspaces ; these ribs are not
nodulous nor keeled, they are sometimes slightly rippled by the in-
cremental lines, and, as usual, are smaller, fainter and more crowded
distally ; the interspaces are rounded ; near the dorsal margins on
each side of the beaks there is an ill-defined narrow space which is
devoid of ribs ; the average number of ribs is about 50 ; the inner
margins of the valves are crenulated ; the hinge delicate and normal ;
the inner surface of the valves white, polished ; the outer surface
usually shows three or four concentric sulci due to resting-stages.
Length 38, height 33, diameter 18 mm., of a fully adult specimen ;
a shorter variety measures 25.5, 24.5 and 16 mm. for the same di-
mensions.
From Bering Sea (in 70 fathoms, temperature 39.0°) south to
Monterey Bay, California (in 20 fathoms); the depths varying from
20 to 70 fathoms and the temperatures from 39.0° to 52.3°. Col-
lected by Kennerley, Swan, the U. S. F. Com. Str. Albatross, and
Mr. S. S. Berry, of Stanford University.
This species somewhat resembles C. hayesii Stimpson from the
Arctic Sea but has more numerous and more uniform ribbing. It
wants the angulation (in fresh specimens fringed with periostracum)
which is characteristic of G. ciliatum Fabr. in the young stages, and
has more, and more crowded, ribs, and a more solid shell. It is
most nearly allied to C. californiense but has a thicker and more
ventricose shell, is less attenuated posteriorly and less oval in form,
the ribs are more closely adjacent and more uniform in size than in
californiense ; though the number is about the same the diminutive
size of fucanum makes them appear much more numerous. Most of
the 14 specimens examined came from the Straits of Fuca.
THE NAUTILUS. 113
A NOTE UPON THE INSUFFICIENCY OF THE OPERCULUM AS A BASIS
OF CLASSIFICATION IN ROUND-MOUTHED SHELLS.
( Concluded) .
BY L. P. GRATACAP.
SECOND SERIES.
Operculum subcircular, few whorls, rather quickly widening, nucleus sub-
central.
I. Operculum horny.
Genera, Jamaicia, Licina, Choanopoma.
Jamaicia. Operculum horny, exteriorly convex, with few rugose whorls,
obliquely striated, sublammellose.
Licina. Operculum not described.
Choanopoma. Operculum horny, subcircular, whorls rather quickly widen-
ing, with free external margin, acute, frequently raised into lofty lamella,
nucleus subeccentric.
II. Operculum cartilaginous.
Genus, Adamsiella.
Adamsiella. Operculum circular, thin, subcartilaginous, with few whorls,
gradually increasing, with somewhat free edges, nucleus subcentral.
• THIRD SERIES.
Operculum oval or angulated, few whorls, more or less quickly increasing,
nucleus eccentric.
I. Operculum shelly.
Genera, Lithidion, Otopoma, Cyclostomus, Tudora, Leonia.
Lithidion. Operculum subcircular, shelly, whorls rather rapidly increasing,
with a subcentral keel, strong, convex, furnished with simple margins.
Otopoma. Operculum horny, solid, few whorls, convex center, simple
margin.
Cyctoslomus. Operculum suboval, horny, plane, four to five whorls, gradu-
ally increasing, simple margin, nucleus subeccentric.
Tudora. Operculum oval, horny, plane, two or three whorls rapidly increas-
ing, obliquely arcuate, striate, or sulcate, nucleus quite eccentric, with the left
and lower margin closely fitting to the peristomes.
Leonia. Operculum oval, shelly, externally perconvex, unispiral, nucleus
placed near the columellar margin.
II. Operculum cartilaginous, covered externally with a thin shelly layer.
Cistula. Operculum oval, thin, cartilaginous, externally furnished with a
thin horny layer, with a few whorls gradually increasing, margin generally
free, nucleus eccentric.
114 THE NAUTILUS.
III. Operculum entirely cartilaginous.
Chondropoma. Operculum oval, subcartilaginous, plane, few whorls, rapidly
increasing, nucleus generally quite eccentric.
IV. Operculum double interiorly concamerated.
Pomatias. Operculum cartilaginous, few-whorled, made up of two laminae,
interiorly concamerated.
V. Operculum corneous.
Genera, Realia, Omphalotropis, Bourciera.
Realia. Operculum thin, horny, few-whorled.
Omphalotropis. Operculum thin, horny, few-whorled.
Bourciera. Operculum oval, somewhat solid, horny, with few rapidly in-
creasing whorls.
Later authors have extended this list of genera, but Pfeiffer's
synopsis practically embraces the important and distinguishing gen-
era. The force assigned to the Operculum as separative of the genera
naturally appears exaggerated, when their characters are thus iso-
lated, but in view of the purpose of this paper to emphasize their
secondary, or in cases, entirely negligible weight, this isolation serves
the more explicit object of fixing attention solely upon the operculate
features.
To begin with, in the genus Cyclotus, there exist differences in
the opercula of many species almost as great as that between the
opercula of recognized genera. In the species C. corrugatum, Swb.
the whorls of the laminae in the operculum are margined by erect
incurved slightly striate free fillets, the whole operculum presenting
the appearance of a watch-spring; whereas in typical examples of the
operculum of Cyclotus, the laminae are flat with edges strictly in
contact or slightly overlapping, the latter feature becoming extreme
in seminudum. There is here no essentially different principle of
construction involved but the formal contrast in appearance might as
safely be invoked to make another genus in the case of C. corru-
gatum (as it has been) so far as opercula offer signs of generic dis-
tinction. Troschel and H. and A. Adams have indeed placed this
shell, along with asperulus, cingulatus, crassus, etc., in all twenty-
five species, in the subgenus Aperostoma.
The shelly substance of the operculum of Cyclotus is however a
quite constant feature. The shelly opercula are found in such small
shells as fodiens, hunanus, parvulus, minimus, where it might be ex-
pected that the calcareous secretions would be less complete.
THE NAUTILUS. 115
The opercula of the small shells Alycaeus and Dtplotumatinaare not
often found in shells in collections. Where I could examine them
under a one-inch objective they certainly offer no determinative dif-
ferences from the opercula of Cyclophorus, and like the latter are
horny (corneous). Compare for instance the opercula of Alycaeus
rathou/siana, Hende, Gyclophorus parapsis, Benson.
The genus Pterocyclos is certainly closely allied to genera Opis-
thoporus, Spiraculum, Shiostoma, but the opercula of these four
genera exhibit two contrasted forms. Taking Pterocyclos angulif-
erus Soul, as an example of one form, the operculum forms a calcare-
ous button, concave upon the outer side, interiorly filmed by a horny
scale covering its entire width, grooved at the sides, and showing on
its exposed surface closely wound narrow whorls, throughout ob-
liquely striate. This configuration and structure is quite as naturally
and truly referable to the opercula of some species of Cyclotus as C.
auriculata, Kob., etc.
This form of operculum is found in some species of Opisthoporus
(vide biciliatus, Mouss., birostra, Pfr.) and the distinction made by
H. and A. Adams between the opercula of Opisthoporus and Pterocy-
clas is certainly misleading if universally applied. Another more
common type of operculum, seen in Spiraculum, Rhiostoma, some
species of Opisthoporus and Pterocyclos is a spiral lamina, concave
interiorly, more or less deep with the free edges of the laminae erect
or explanate and horizontal, arising screw-wise steeply around a
solid nucleus (Rhiostoma} or more spreading with corneous intercal-
ation (Spiraculum}. The morphology of the shells themselves sep-
arates these genera, though it is probable the distinctions should form
subgenera, but the opercula present two forms, which while partially
restricted are surely not enough so to offer any basis for strict
classification.
But the confusion, so far as opercula are considered, does not end
here. The opercula of the second type of Pterocyclos, Opisthoporus,
Spiraculum, Rhiostoma, in substance and structure, is practically re-
peated in those of Choanopoma. Both groups show the exsert, erect,
or spreading laminae, the spiral curvature, the union of calcareous
and corneous texture, though the inner surface of the operculura in
Choanopoma is usually flat, and in some instances as C. pulchrum,
Gray, the coarse expansive character of the whorls contrasts with
the analogous feature in Pterocyclas, etc.
116 THE NAUTILUS.
The operculum of Leptopoma is essentially that of many species of
Oyclophorus (compare Leptopoma sericatum, Pfr. with C.fulguratus,
Pfr. and C. borneense, Mtf.). It is membranous, corneous, arcti-
spiral suppressedly striate, on the laminae, concave, with a polished
inner surface. The fimbriated shaggy exfoliated surfaces of many
opercula of the larger Oyclophorus (see G. arthritis, G. nilagirianus,
G. oculus-capri, C. tuba, C. validus, etc.) are age characters, and
seem also to have some relation to the moist habitats of the animals.
This type of operculum is repeated in Megalomastoma, though the
shell in the latter case is pronouncedly and divergently different (see
M. ventricosum, M. verruculosum.}
Gataulus is in its operculate character identical with the fore-
going. Although from the size of the apertures the opercula are
small, they are in construction, substance, expression, and micro-
scopic features, inseparable from the opercula of Cyclophorus, Mega-
lomastoma, and Leptopoma. The flatness usually observable in the
opercula of the last two genera is shown in Gyclophorus in small
shells, the concavity of the opercula of the latter genus being the
result of the natural curvature produced by the protruding fibres of
the larger foot in larger species.
The operculum of Pupina is unmistakably indicated in structure,
nature, and physical appearance with all the foregoing, its tenuity alone
arid flatness offering only the most evanescent and unreal discrimina-
tion from the other genera. Pupinella, Registoma, Gallia, are quite
inseparable in their operculate character. Aulopoma is u shell very
near in external characters to Pterocyclos, Opisthoporus, Spiraculum,
Rhiostoma, omitting the lip expansion and spiracle, but the oper~
culum is sharply contrasted. The operculum in Aulopoma is corneous,
consisting of a flat spiral made up of tubular whorls coarsely striate
and embracing, cap-like, the margins of the shell aperture. If the
distinction of this genus is based upon the operculum it rather forcibly
separates a shell from its natural generic position.
In structure, at least, the sub-orbicular operculum of Otopoma
naticoides Rec., with its cycloidal development with the tangential
strise leaving the whorls and extending outward, but forming flexu-
ous lines on the limits of the last whorl, is repeated in the authentic
opercula of Cyclophorus metabletus Crosse & Fischer.
The instances of exact repetition in the opercula of Cyclotus and
Gyclophorus ase numerous. The operculum of Gyclotus translucidus
THE NAUTILUS.
117
Sowb. is certainly reproduced in those of Cyclostorna xanthocheilus,
C. barclayanus, G. campanulatus. The Hat thin spiral, almost equal
with the slightly overlapping ascent outward to the last whorl, striate
on its exterior edges, is closely repeated in each.
In Adamsiella gray ana Pfr., A. variabile Ad., there is a spiral oper-
culum with exsert laminae, somewhat multifolded or lamellose on
edges, but structurally distinctly like Choanopoma and Opisthoporus.
The operculum of Tudora and Cistula with their rapidly increasing
whoris, the last expansive and filling one-third of the aperture, nuc-
leus subcentral, are practically identical. The thin substance of the
operculum of C/iondropoma gives it some essential differentiation, but
in the theory of its form it is exactly like Tudora and Cistula. The
outer surface of the operculum of Tudora is frequently ( T. shepardi-
anus Ad., T. pupoides Morelet) fibrous-rayed, whereas in megacheilus
its surface is smooth, or obsoletely striate which form is closely imi-
tated by the operculum of Ckondropoma.
The following table exhibits the substance of the operculum in the
genera of round-mouthed shells, as observed in the collection of the
American Museum of Natural History, and as given by Pfeiffer.
The two columns afford slight differences, but such differences might
readily be referred to individual variations in the shells examined by
two observers, using different groups of specimens.
SUBSTANCE OF THE OPERCDLUM.
Observed in Collection.
Testaceous to subtestaceous. Sub-
testaceous in C. crassus, Ad. C.
triliratum Pfr., C. rugatus Guppy,
C. suturale Swb.
Corneous.
Corneous.
Testaceous to Sub-testaceous in P.
rupestris Bens.
Not seen.
Corneous.
Corneous.
Corneous. Membranous.
Corneous.
Corneous.
Corneous.
Corneous.
Cyclotits.
Alycieus.
Diplommatina.
Pterocyclos.
Craspedopoma.
Aulopoma.
Cyclophorus.
Leptopoma.
Megalomastoma.
Cataulus.
Pupinella.
Pupina.
Pfeiffer.
Testaceous.
Sub-testaceous.
Sub-testaceous.
Sub-cartilaginous.
Corneous.
Corneous.
Corneous.
Membranous.
Corneous.
Corneous.
Corneous.
Membranous.
118
THE NAUTILUS.
Corneous.
Corneo- testaceous.
Not seen.
Testaceous to Corneo-testaceous.
In C. majusculum Morelet the
corneous layer is quite intimately
and equally developed with the
testaceous.
Corneous.
Testaceous.
Testaceous.
Corneous layer quite thick in in-
stances as C. bicarinatum Swb.,
C. filosum Swb., C. madagascar-
iense Gray, C. rangelinum, C.
xanthocheilus.
Testaceous to Testaceo-corneous.
Testaceous.
Testaceous.
Corneo-cartilaginous.
Not seen.
Not seen.
Not seen.
Not seen.
Testaceous to Sub testaceous.
Testaceous.
Testaceous to Sub-testaceous.
Testaceous.
Registoma.
Callia,
Jamaicia.
Licina.
Choanopoma.
Adamsiella.
LiUddion.
Otopoma.
Cyclostomus.
Tudora.
Lconia.
Cistula.
Chondropoma.
Pomalias.
Realia.
Oinphalotropis.
Bourciera.
Ilybocystis .
Cyclotopsis.
Ctenopoma.
Diplopoma.
Corneous.
Membranous.
Corneous.
?
Corneous.
Sub-cartilaginoua.
Corneous.
Corneous.
Corneous.
Corneous.
Testaceous.
Cartilaginous.
Sub-cartilaginous.
Cartilaginous.
Corneous.
Corneous.
Corneous.
Collections of the Cyclostomacea are often sadly deficient in oper-
cula, and this very interesting feature is as frequently absent as
present. It seems most probable that opercula have no exact generic
value, and should not be regarded with the extreme importance as-
signed to them by Pfeiffer, H. & A. Adams, and other authors.
It is hoped that a more detailed and illustrated study of them will
be made in the future.
NOTES.
THE WINKLEY COLLECTION OF SHELLS — "A most gratifying
gift was received last June from the Rev. Henry W. Winkley, of
Branford, Conn.; it consists of a large series, some 1600 species and
THE NAUTILUS. 119
varieties of land shells from all parts of the world, except New Eng-
land. His New England collection Mr. Winkley retains for study,
but he writes that he thinks in time it too ' will find its way to the
Museum.' The shells received are in excellent condition. Mr.
Winkley, as a member of the class of 1881, recalls with pleasure his
undergraduate work in the Museum, and gives the collection as his
contribution toward the twenty-fifth anniversary celebration of the
class. It is hoped that Mr. Winkley's gift will prove an incentive
to other graduates to associate their college work with the interests
of science and the Museum."-— (From the Annual Report of the Cura-
tor, Museum of Comparative Zoology, 1905-1906.)
THE MUMFORD COLLECTION or SEIELLS — Through a bequest of
the late Mr. Henry Mumford, the Museum of the Brooklyn Institute
of Arts and Sciences has secured an unusually fine collection of
shells, comprising about 15,000 specimens, representing 2,400 species;
this will be known as the Phebe L. Mumford Collection. It was
originally brought together by Mr. Isaiah Greegor, for many years
a resident of Jacksonville, Florida, who made a special effort to se-
cure the best possible specimens. This makes the collection par-
ticularly valuable for a museum, since the shells are in their natural
condition, retaining the outer layer or •' epidermis," which is usually
removed to show the brilliant coloring beneath, a proceeding that
makes the shell " look pretty," but utterly destroys its scientific
value ; there are, however, series of shells cleaned and polished so
that the student and the lover of the beautiful may alike find what
they wish.
Two years ago the Museum received a valuable gift of shells from
Mr. Julius Brunn, so that the division of mollusks is now very well
represented. — ( The Museum News.}
PUBLICATIONS RECEIVED.
AN ILLUSTRATED CATALOGUE OF THE MOLLUSC A OF MICHI-
GAN : Part I, Terrestrial Pulmonata (Land Snails). By Bryant
Walker (From Report of the Michigan State Board of Geological
Survey, 1906). "This catalogue has been prepared to supply the
need for a convenient manual of Michigan mollusks for the use of
teachers and students of biology throughout the state." It contains
120 THE NAUTILUS.
chapters on the history of Michigan conchology, the relations of the
Michigan fauna, on collecting and preserving mollusks, on classi-
fication, measurements and descriptive terms. While these subjects
are concisely treated, yet Mr. Walker has condensed into his pages
the practical results reached by the modern group of American
workers in both field and laboratory, so far as applicable to the
Michigan fauna. There are few collectors who will not profit by
reading the " Hints on collecting mollusca," and "Arrangement of
the collection." The classification and nomenclature of the cata-
logue are up to date.
The list comprises 81 species, all described and illustrated by cuts
in the text, most of them borrowed from Binney's standard series of
figures, though with a considerable number of original drawings.
Full references are given to previous Michigan literature, as well as
to the original descriptions, and to Mr. Binney's Manual. Excellent
keys to the species are given.
It is no disparagement of other lists to say that the Michigan
catalogue as here presented is the most thoroughly worked-up state
catalogue we have had. Every species has been the subject of
special investigation, as to its characters and identity, and its distri-
bution in Michigan. The time and labor involved in working up a
state fauna in this thorough manner will be appreciated only by those
who have undertaken the rigid verification of every record in the
fauna of an equal area.
A summary of the work on distribution done up to this time is
given in the tabular census, giving the distribution of all the species
by counties. Records have been received from all but 16 of the 84
counties in the state. It will be noted that the comparatively re-
mote and inaccessible Upper Peninsula is well represented in the
Census, and the Boreal character of its fauna is well shown by the
absence of nearly all of the Polygyras, larger Zonitidx, etc.
Species interesting because of their rarity or on account of notable
extensions of their known range, are VHrect wheatleyi and rhoadsi,
Pallifera hemphilli, Succinea retusa peoriensis (here for the first time
figured), Vertigo morsei, Cochlicopa lubrica morseana, etc. Excel-
lent and much-needed original figures of Strobilops labyrinthica,
S. virgo and S. affinis are given.
Mr. Walker is at work upon the fresh-water mollusks, to form the
second part of the catalogue. We will all look with interest for his
treatment of the aquatic pulmonates. — H. A. P.
THE NAUTILUS.
VOL. XX. MARCH, 19O7. No. 11.
A NEW FOSSIL BUSYCON (FULGUK) FEOM FLOEIDA.
BY T. H. ALDRICH.
BUSYCON MONTFORTI n. sp. Plate VI.
Shell large, solid. Whorls about seven ; those of the spire with
slight nodules, which on the body whorl develop progressively into
small, medium, and then large spines, ten in number on the specimen
figured ; these spines are flattened above, and partly so below, and
differ from those of Busycon eliceans Montf., by being filled by the
successive growth additions. Surface covered with raised lines, ex-
cept on the middle of the body whorl. Aperture within still yellow-
ish-brown. Canal contracted and twisted somewhat to the left.
Inner lip with a strong callus carrying a fold at the angle with the
canal ; outer lip corrugated within. The specimen has the spire
slightly broken, but it is probably pointed like Busycon eliceans. The
canal is considerably shortened by breakage.
Locality : Shoal River, Walton County, West Florida.
Remarks : The geological horizon is in the Upper Oligocene of
Dall. This species is especially interesting because it shows that
Busycon eliceans Montf. is a distinct form with an Oligocene (?) ances-
tor. There are a number of minor differences between the two
species, but when placed side by side they show without question
their intimate relations. The descent of the group will need to be
cousiderably modified to conform to the new discovery.
122 THE NAUTILUS.
A NEW SPECIES OF SEGMENTINA.
BY BKYANT WALKER.
SEGMENTINA CRASSILABRIS n. sp., PI. VII, figs. 4-6.
Segmentina wheatleyi, Witter, J. of Conch., I, 388 (1878).
«' ? Walker, Naut. VI, 137 (1893).
Walker, Rev. Mich. Moll., 18 (1895).
Shell dextral, broadly umbilicate, planorboid, smooth, slightly
wrinkled by subobsolete growth lines and microscopic transverse
lines and undulations ; light horn colored ; whorls 4^, regularly in-
creasing, those of the apex flattened and concavely depressed, ante-
penult and body whorl rounded above, the latter deflected and slightly
expanded towards the aperture, and sharply contracted at the lip,
forming an obtuse crest just behind the edge of the lip, obtusely
angled below, around the deep, subfunicular umbilicus ; aperture
oblique, subtriangular, greatly contracted by a heavy callous
ridge just within the lip, the face of which is concave, the edge of
the lip being sharp and black ; the extremities of the labial ridge are
connected by a slight callus across the parietal wall ; apertural
lamellae six ; two parietal and four (palatal) on the outer wall.
The principal parietal lamella is a thin, broad, sigmoid blade rising
from the lower third of the parietal wall, and extending upward and
backward nearly to the suture, it is widest in the center, the ante-
rior extremity is subtruncate and rather abruptly bent outwards,
posteriorly it slopes gradually to its termination ; below this and
just behind the curve in the anterior portion and about halfway be-
tween it and the base is a short, sharp denticle projecting obliquely
forward and downward ; the lower palatal fold extends transversely
nearly across the entire base of the whorl and is a low, strong fold,
the upper surface of which is parallel with the base, the inner ex-
tremity is opposite the lower parietal lamella and rises abruptly at
right angles to the base, the outer extremity is rounded and bent for-
wards, sloping down to its termination, viewed externally the base of
this fold roughly resembles a figure-7 ; immediately above this,
about halfway between it and the lower extremity of the third pala-
tal fold, and about at the periphery of the whorl is a short, trans-
THE NAUTILUS. 123
versely diagonal lamella, the anterior end of which is lower than the
posterior, which is slightly bent upwards ; above the anterior extrem-
ity of this fold is the third palatal fold, a short, slightly oblique
lamella, the posterior end of which is slightly curved backwards and
slopes more gradually to its termination than the anterior extremity;
above this and at its posterior extremity is a short, transverse denti-
cle, which lies immediately below the superior arch of the whorl.
Alt. 3, greater diam. 7^, lesser diam. 6 mm. Types (No. 2998
coll. Walker) from Hamtramck, Wayne Co., Mich. Also from
Kent and Monroe counties, Mich.; Muscatine, la.; Knox county,
Vincennesand Brownston, Ind.
This very distinct species was first seen in 1878, when specimens
from Muscatine, la., were received from Prof. F. M. Witter, labelled
S. wheatleyi Lea, on the authority of the late Dr. Jas. Lewis. Sub-
sequently, when the species were detected in Michigan, in reliance
on this identification it was so listed in the two papers cited above.
The recent receipt of the true S. wheatleyi from Princeton and Boli-
gee, Alabama, collected by H. H. Smith and A. A. Hinkley, has
shown that the reference of this form to that species was erroneous.
Although occupying an intermediate position between the well-
known S. armigera and S. wheatleyi, crassilabris, is quite distinct from
either. In external appearance, compared with armigera (Fig. 1-3),
it is smaller and more compactly coiled and higher in proportion to its
width, the umbilicus is smaller and deeper, and the angulation of
base of the whorl around the umbilicus more pronounced. In armi-
gera, the whorls are more regularly rounded and the aperture lacks
the crest behind the lip, and is not at all contracted, the thickening
within the lip is very slight and does not sensibly diminish its calibre.
In wheatleyi (Fig. 7-9) the distinctive features of crassilabris are
all intensified. The apical whorls are more depressed, the body-
whorl is obtusely angulated above and shai'ply carinated around the
umbilicus, which is funicular, the inner whorls scarcely, if at all, ob-
truding beyond the curve of the body-whorl ; the aperture is more
oblique and more contracted in front of the crest, which is more
prominent. The inner ridge of callus is less developed and lacks
the regular concave slope to the edge of lip characteristic of crassi-
labris.
As recently stated by Pilsbry and Ferriss (Proc. A. N. S. P.,
1906, p. 166) in regard to S. armigera and wheatleyi, the arrange-
124 THE NAUTILUS.
ment of the apertural lamelire in these three species is fundamentally
the same. Their number, position and character are the same in
all. The differences, which are, however, specifically characteristic,
are only in the degree of development.
In armigera, the principal parietal lamella is much shorter, less
oblique and its anterior end is less curved and truncated than that of
crassilabris, the palatal folds are all less developed, and the spaces
between them consequently greater ; the upper extremity of the basal
fold is less deflected and scarcely noti cable externally. In both
these species, there is a marked tendency to develop an additional
fold in connection with the central palatal giving it a V-shape, which
is quite noticeable from the exterior.
In wheatleyi, the position of the principal parietal lamella is more
like that of armigera than of crassilabris, being, taken as a whole,
less oblique and the anterior end is less bent forward than in the
latter species, but the posterior extremity is more prolonged than in
armigera, and is continued for a short distance parallel with the
suture ; the basal palatal is much wider than in either of the other
species and is abruptly contracted at its base, while the upper ex-
tremity is deflected as in crassilabris ; the second palatal fold is re-
markably developed, rising abruptly near the base of the whorl ; it
runs back for a short distance parallel with the base, and curves up-
ward, terminating just below the superior carina, having much the
shape of a boomerang. The third palatal is only slightly oblique,
the lower end is bent backwards and close to the upper extremity is
a small knob, not sufficiently differentiated from it to be called a dis-
tinct denticle, but evidently similar to the accessory fold noticed in
the other species ; the upper extremities of this fold and of the basal
palatal are about on the same level. The fourth palatal is situated
directly under the superior carina and is a short, straight, transverse,
club-shaped fold, the posterior extremity being the thickest.
EXPLANATION OF PLATE VII.
Figs. 1-3. S. armigera Say, Mohawk, N. Y.
Figs. 4-6. £. crassilabris (Type), Hamtramck, Mich.
Figs. 7-9. S. wheatleyi Lea, Princeton, Ala.
THE NAUTILUS. 125
DESCRIPTIONS OF NEW SPECIES OF LYMNJEA.
BY FRANK C. BAKER.
LYMN^EA DALLI n. sp.
Lymnsea parva Baker, NAUTILUS, XIX, p. 52, 1905. (Not of
Lea.)
Lymnsea dalli Baker, Bull. Ills. State Lab. N. H., VII, p. 104,
1906.
Shell very small, thin, ovate-conic, turreted ; color greenish or
whitish-horn ; surface dull to shining, marked by heavy, crowded
growth-lines, which are elevated into distinct ribs in some specimens ;
protoconch very small, flatly rounded, light-horn colored ; whorls 4^
to 5, rounded and distinctly shouldered ; spire generally obtusely
conic, turreted, a trifle longer than the aperture ; sutures very deeply
impressed ; aperture elongate, ovate or elliptical, the peristome con-
tinuous in many specimens ; outer lip acute ; inner lip forming a
rather flat extension over the umbilical region, leaving a rather pro-
nounced chink ; the lower part of the aperture is somewhat effuse ; the
columellar extension of the inner lip is appressed so as to form a
pseudo-plait ; the inner edge of the outer lip frequently forms a rib-
like ridge in very old specimens.
Length 3.25, breadth 2, aperture length 1.5, breadth 1 mm. Lake
James, Ind.
Length 4, breadth 2, aperture length 2, breadth 1 mm. Lake
James, Ind.
Length 3, breadth 2, aperture length 1.25, breadth, .9 mm. Lake
James, Ind.
Length 4.25, breadth 2, aperture length 1.75, breadth 1 mm.
Lake James, Ind.
Length 4.5, breadth 2.5, aperture length 2, breadth 1.1 mm.
(scalariform).
Length 3.75, breadth 2.75, aperture length 2, breadth 1.25 mm.
Rockford.
Length 4, breadth 2.1, aperture length 2, breadth 1.1 mm. Rock-
ford.
Types : Chicago Academy of Sciences ; cotypes, Academy of Nat-
ural Sciences, Philadelphia ; Smithsonian Institution, Washington.
Type Locality: Lake James, Steuben Co., Indiana.
126 THE NAUTILUS.
Range: Indiana to Montana, south to New Mexico and Texas.
Station : Same as humilis, curta, and the other small Lymnaeas.
Remarks : This little species, the smallest of our American Lym-
naeas, is related to parva, appearing at first sight to be a small exam-
ple of that species. It differs from parva in its smaller size, rounder
and more turreted whorls, more slender outline, longer and narrower
aperture and smaller and less conspicuous umbilicus. The whorls
are inclined to be shouldered also, a feature not emphasized in parva.
There is some variation in the length of the spire, many specimens
having a rather long spire, while in others the spire is much
depressed, and the whorls are humped, this last being in slightly
abnormal forms.
In THE NAUTILUS (xx, p. 52), this species was described as the
parva of Lea, but a careful study of Lea's types in the Smithsonian
Institution makes it evident that it is not that species, but a new one,
hitherto unnoticed. Young specimens of parva are similar to dalli,
but may be distinguished by the lesser number of whorls in shells of
the same size and in the different shape of the whorls. The two
small forms, parva and dalli, are related, but are easily separated
when attention is given to the details of form and size.
This peculiar little species is named in honor of Dr. "William H.
Dall, Curator of the Division of Mollusks of the Smithsonian Insti-
tution.
LYMN.EA LEAI n. sp.
Shell large, ovate, thin ; color, yellowish-horn ; surface with fine
lines of growth crossed by equally fine spiral lines ; whorls 6, flatly
rounded, very rapidly increasing in diameter, the last whorl very
large ; spire short, broadly, ovately conical ; sutures well impressed ;
aperture elliptical or elongate-ovate, narrowed above, longer than the
spire ; outer lip thin, only slightly thickened by a longitudinal rib ;
inner lip thin, narrow, appressed to the umbilical region so as to
leave a well-marked chink ; callus on the parietal wall thin, wide,
well marked ; columella twisted, with a strongly developed ascend-
ing plait.
Length 30, width 13.5, aperture length 18, width 8.5 mm. Type.
Length 28, width 13.5, aperture length 17, width 8 mm. Type.
Length 31, width 14.5, aperture length 18.5, width 9.5 mm. Type.
Types: The Chicago Academy of Sciences; 1 specimen, cotypes,
collection W. A. Nason, 2 specimens.
'!HE NAUTILUS XX.
PLATE VI.
ALDRICH : BUSYCON MONTFORTI.
THE NAUTILUS, XX.
PLATE VII.
8
3
6
9
1-3. -SEGMENTINA ARMIGERA. 4-6. S. CRASSILA t- RIS.
7-9. S. WHEATLEYI.
THE NAUTILUS. 127
Type locality : Near San Francisco, California.
Range: California, near San Francisco (W. A. Nason); San Joa-
quin River (Mrs. E. E. Bush, Phil. Acad.); Merced Lake (Stearns,
Dull, Smithsonian Institution); between Animas and La Plata,
Colorado (Ingersoll).
Remarks : L. leaii seems to be a very distinct species, easily recog-
nized by its short spire, very large aperture and heavy columellar
plait. It is more nearly related to L. proximo, rowellii Tryon than
to any other species, but seems to be distinct. It has probably been
heretofore identified as a form of the protean species palustris, but it
is unquestionably distinct from any form of that species. It also re-
sembles L. reflexa hemphilliana Baker, but differs in being very much
broader and has a differently shaped and larger aperture. It is
named in honor of one of America's best known conchologists, Dr.
Isaac Lea.
THEEE NEW SPECIES OF SCALA FKOM CALIFORNIA.
BY WILLIAM HEALEY DALL.
During the last year or two, among the minute shells, collected by
various correspondents on the Californian coast, there have been
several undescribed forms of Scala, descriptions of which follow.
Scala berryi n. sp.
Shell small, white, six-whorled : nuclear whorls polished, smooth,
without varices ; later whorlo with rather strong, reflected, faintly
axially striated varices, on the last whorl about 20 or 21 in number ;
these varices are quite close-set, not angulated in front of the suture
and markedly reflected, with the interspaces smooth ; base with no
basal disk or cord, the coil imperforate, the peristome rather callous
on the pillar side. Long. 3.5., max. diam. 1.75 mm.
Dredged in 200 fathoms, San Pedro Bay, Cala., Mrs. Oldroyd ;
and in 12 fathoms off Del Monte, Monterey Bay by S. S. Berry.
U. S. Nat. Mus. 107,724.
This species is, perhaps, nearest to S. clathratula A. Adams, and
to the next species.
Scala rectilaminata n. sp.
Shell very similar to S. berryi, with the same number of whorls
and varices, but having the nuclear shell smaller, the test less heavy,
128 THE NAUTILUS.
the varices nearly vertical to the surface of the whorl, instead of
reflected, which makes them appear sparser, though really the same
number occur on the whorl ; the varices are narrower and the peris-
tome less heavy and wide. Long. 3.25, max. diam. 1.6 mm.
Dredged in twelve fathoms mud, Monterey Bay, S. S. Berry.
U. S. Nat. Mus. 110,430.
The specimens seen are of a more yellowish-creamy-white than
S. berryi which has a bluish subtranslucent appearance.
Scala (Cirostrema) montereyensis n. sp.
Shell small (probably not full grown), the nucleus lost but with
five subsequent rapidly increasing whorls ; shell substance in two
layers, the inner translucent white, solid, the outer opaque white,
frothy, porous, with numerous puncticulations arranged in harmony
with the incremental lines, except on the basal disk where they form
spiral lines ; varices low, solid, with a spongy surface, nine in num-
ber ; basal disk conspicuous, slightly concave ; bordered by a con-
spicuous cord ; aperture gibbous, patulous near the imperforate axis.
Long. 2.5, diam. 1.5 mm.
Dredged in 25 fathoms mud, off Del Monte, in Monterey Bay,
Cala., by S. S. Berry. U. S. N. Mus. 110431.
This shell represented by two specimens, of which one is in Mr.
Berry's collection, is doubtless immature, but there is no other species
known north of Cape St. Lucas belonging to this special group, and,
though the characters require rather high magnification to see them
clearly, it cannot be confounded with any other Californian species.
NOTE OK THE GENUS PSILOCOCHLIS LALL.
BY WILLIAM H. DALL.
This curious Turbinella, collected by Mr. McCallie in the Eocene
of Georgia, was described in THE NAUTILUS for May, 1904, p. 9.
Additional material shows that the mature shell is covered with a
coating of enamel which extends to the very apex of the spire, ob-
scuring the sutures, a feature not hitherto reported in connection
with the genus Turbinella or any of its close allies ; so that Psilo-
cochlis described as a subgenus, seems fully entitled to generic rank.
Only the typical species, P. mccalliei Dall, is at present known.
THE NAUTILUS. 129
PUBLICATIONS RECEIVED.
A REVIEW OF THE AMERICAN VOLUTID^E. By "Wm. H. Ball.
(From the Smithsonian Miscellaneous Collections, vol. 48, 1907.)
This paper is the revision promised by the author in his " Notes on
some names in the Volutidai," in THE NAUTILUS for April, 1906.
Some years ago Dr. Dall outlined two important groups of the Vol-
utidse as follows :
1. Subfamily Scaphellinse, with a membranous protoconch which
is lost early in the intracapsular development and is replaced by a
shelly envelope, the secondary nature of which is evident in well pre-
served specimens.
2. Subfamily Vohitinse, a shelly protoconch. In these forms there
is no membranous stage, the protoconch being shelly throughout its
history. This shelly apex is never naturally lost.
His recent researches have shown that the type of the genus Sca-
phella belongs to the Volutina, and therefore the subfamily name
ScaphellincK has been given up and Caricellinse substituted for it. In
his review Dr. Dall employs not only the apical shell characters
which he regarded as important in his earlier papers, but also many
features of the soft anatomy. Stress is laid on " the presence or ab-
sence of a caecum to the ossophagus ; the characters of the radula; of
the verge, or external male organ ; and the presence or absence of an
operculum." A new three-fold division of the Vohitidee, here out-
lined, is regarded as provisional. It may be briefly summed up as
follows :
1. Subfamily Votutince. A shelly, persistent protoconch, sessile
eyes, operculum usually absent and the radula teeth usually in one
tricuspid series.
2. Subfamily Caricellinse. A membranous protoconch, operculum
absent, radula variable or absent. Otherwise as in Volutince.
3. Subfamily Volutomitrinse. Protoconch supposedly shelly ; adult
unicolored, with a conspicuous periostracum ; small stalked eyes,
" radula of a single long series, the separate teeth unicuspidate, with
deeply arcuate bases." No operculum.
The genera and species are distributed among the subfamilies as
follows :
Volutince : Valuta (Linne) Lamarck, with the following species :
musica L., virescens, Sol., ebrosa L.
Lyria Gray with the species beauii Fischer and Bernard!.
130 THE NAUTILUS.
Enseta Adams with the species barnesii Gray, cumingi Brod.,
pedersenii Verrill, cylleniformis Sowb., archeri Angas, reevei Dall,
guildingii Sowb.
Plejona Bolten containing species " conchologically related to the
F. spinosa Lamarck."
Caricellinae : Adelomelon Dall, with the species ancilla Sol., sub-
nodosa Leach, benthalis Dall, martensi Strebel, magellanica Lamarck,
ornata Lahille, beckii Brod., tuberculata Swainson, brasiliana Sol.,
ferussacii Donovan, paradoxa Lahille, stearnsii Dall, philippiana
Dall.
Zidona H. and A. Adams with the species angulata Swainson.
Tractolira Dall with the species sparta Dall.
Aurinia H. and A. Adams with the species dubia Brod., robusta
Dall, gouldiana Dall.
Maculopeplum Dall with the species junonia Hwass, dohrni Sowb.
Volutomitrina, : Volutomitra Gray with the species gronlandica
Beck and alaskana Dall.
Owing to the scarcity of specimens, we do not realize that on the
coasts of North and South America there are 33 recorded species.
There are considered to be two centers of distribution for the Amer-
ican species, the southern end of South America is taken as one
center and the Antilles as the other. The paucity of species along
the Pacific coast north of south latitude 40° is remarked upon.
Under the genus Aurinia, typified by A. dubia, Dr. Dall says :
" This genus is the degenerate descendant in one line, as Maculopep-
lum is a normal descendant in another, from the Eocene Caricella.
Its most prominent feature is the enfeebled plaits of the pillar,
usually of diminished number as well as size, its thin shell and prom-
inent Caricella-nucleus. The absence of the radula it shares with
Maculopeplum, though Holla, evidently a close relative, and even
more degenerate as regards the shell, has retained the radula." The
type of the genus Maculopeplum is the well-known Valuta junonia
Hwass.
In his note on the Volutida which appeared in THE NAUTILUS
for April, 1906, Dr. Dall calls attention to the fact that the type of
Swainson's genus Volutillthes is not Voluta spinosa but Voluta mur-
icina Lam. This point was ably confirmed by Mr. R. Bullen New-
ton1 in June, 1906.
1<(Note on Swainson's Genus Volutilithes," Proc. Malac. Soc., Loud., June,
1906, p. 100.
THE NAUTILUS. 131
Volutilithes muricina is evidently well removed from the forms
usually described as Volutilithes splnosus and Volutilithes petrosus.
The use of the generic name Plejona to designate such forms as the
the latter, seems, however, to be without warrant. As Mr. Newton
has pointed out, Bolten refers his Plejona fossilis to four figures in
d'Argenville. These four figures represent not only four different
species, but four different genera!
The very existence of the name Plejona rests, therefore, on a poor
basis, and it is hard to see how its type P. fossilis, can be restricted
to V. spinosa.
But to return to the application of Plejona to American Eocene
forms, the study1 of very large and carefully collected series from our
Gulf States has convinced the reviewer that the forms described as
Athleta tuomeyi by Conrad are nothing but races of his Voluta petrosa
which have been subjected to an unfavorable environment. They
are of one genetic stock, and the differences can in no sense be regarded
as generic; in fact they are not even specific.
M. Cossmann * in reviewing this study pointed out that if its re-
sults are to be accepted, Athleta must be substituted for Volutilithes
Swainson (1840 non 1831).
Athleta rarispina is the type of Athleta and is probably related to
Lamarck's Voluta spinosa in about the same way that A. tuomeyi is
related to Conrad's Voluta petrosa. With our present knowledge it
seems better to adopt Athleta for Conrad's Voluta petrosa, its varia-
tions, mutations and allies, than to revive Plejona.
In conclusion it is noted with regret that Dr. Dall has not pro-
vided this interesting and important paper with figures, which would
have been of the greatest assistance to workers having only small
collections or small libraries at their command. — BURNETT SMITH.
FOSSIL MOLLUSCA FROM THE JOHN DAT AND MASCALL BEDS
OF OREGON (Univ. of California Publications, Bull. Dept. of Geol-
ogy, V, no. 3, pp. 67-70). By Robert E. C. Stearns. The new
species are Epiphragmophora dubiosa, Pyramidula lecontei and Lym-
ncea maxima. The first is apparently related to fidelis or mormorum ;
'Smith, Burnett: " Phytogeny of the Races of Volutilithes petrosus." Proc.
Acad. Nat. Sci., Phila., May, 1906, p. 52.
* Revue Critique de Pateozoologie, p. 222, October, 1906.
132 THE NAUTILUS.
the second to Oreohelix strigosa. Together with Dr. Stearns' previous
report on John Day land shells (Science, 1902, 153), this paper is of
great interest, showing the antiquity of the present generic groups of
land shells, which were manifested in forms not unlike those now ex-
isting in the same region, as far back as the miocene — H. A. P.
CATALOGUE OF THE MOLLUSCA OF ILLINOIS. By Frank Collins
Baker (Bull, of the 111. State Laboratory of Natural History, VII,
article 6, 1906). This catalogue " is intended to record every refer-
ence to the mollusca occurring in the state, which may be found in
the published lists, and also all scattered references wherever pub-
lished." Under each species the author quotes every locality from
which specimens have been recorded, giving the authority for each
in parenthesis. 241 aquatic and 91 terrestrial species are recorded.
No less than 28 species and varieties have been based upon Illinois
specimens, seven of them being now considered synonymous. From
only ten counties, says Mr. Baker, have careful lists been made. In
this catalogue records have been obtained from 69 counties, of from 1
to 145 species. What has been done, and the great amount of work
still to be done, is graphically shown in the map. The richest portions
of the State, along the Wabash and Ohio rivers, are still very little
known; but in the upper counties several energetic naturalists have
thoroughly worked up the fauna, so that this first attempt at a State
catalogue is very creditable to the author, as well as the other natur-
alists, Messrs. Ferriss, Hinkley, Handwerk, Marsh, Nason, Strode,
Hart and others, who have contributed records.
It is inevitable in a first attempt to cover so large an area, that
some doubtful records, demanding careful verification, must be in-
cluded, and this catalogue is naturally no exception. Baker and
his co-workers will doubtless now set about the task of investigating
these questionable records. A paper by Prof. Shimek on his inves-
tigation of Pyrguiopsis in Rock River seems to have been overlooked.
Together with Baker's " Mollusks of the Chicago Area," this cata-
logue forms an excellent basis for further work in Illinois.
THE FOSSIL MOLLUSCA OF FLORISSANT, COLORADO. — By T. D.
A. Cockerell (Bull. Amer. Mus. Nat. Hist,, xxii, 1906). The fol-
lowing species are described as new. Omphalina (?) laminarum,
Planorbis florissantensis, Lymn&a sieverti, scudderi, Sphcerium jlor-
issantense. The new name Columna haydeniana is proposed for
Bulimus teres M. & H., not Olivier.
THE NAUTILUS.
VOL. XX. APRIL, 19O7. No. 12.
NOTES ON SOME NEW MEXICAN ASHMUNELLAS.
BT H. A. PILSBRT AND JAS. H. FERRISS.
Since the publication of the results of our work on Ashmunella in
1905,1 various correspondents have contributed specimens which
throw light upon a number of dubious points relative to the group of
species inhabiting the Sierra Blanca and Sacramento ranges, in New
Mexico, east of the Rio Grande.
There can be no doubt that a systematic campaign in this region
by collectors who are not afraid of hard work, and who get live
shells, would be well rewarded by a most interesting catch. It is
absolutely essential, however, to get to the mountain-tops, and to dig
deep in the rock-heaps. You will need a Ferriss hoe and stout
gloves, and you must forget fatigue.
ASHMUNELLA RHTSSA TOWNSENPI (Bartsch). Plate VIII, figs. 1, 2.
Ashmunella townsendi Bartsch. Smiths. Misc. Coll., xlvii, p. 13
(Aug. 6, 1904).
Ashmunella rhyssa townsendi Bartsch, Pilsbry. Proc. A. N. S.,
Phila., 1905, p. 231.
This race differs from A. r. miorhyssa chiefly by its strongly-
ribbed surface, like A. altissinm, which, however, is far smaller,
more depressed and more openly umbilicate. Three of the original
lot of townsendi, taken by Prof. Townsend in the Sierra Blanca
'Mollusca of the Southwestern States, Proc. A. N. S., Phila., 1905, pp.
223-253.
134 THE NAUTILUS.
above Ruidoso, N. M., sent by Prof. Cockerell, one with the spire
broken, measure :
Alt. 8.8, — 8 mm.
Diam. 14.1, 14.8, 13 mm.
Whorls 5£, — 5£.
Ribs on the last whorl 60, 48, 46.
The number of ribs on the last whorl is only approximately stated,
since on account of irregularities and minor wrinkles or ripples it is
impossible to make a quite definite count.
The weak trace of a parietal tooth is visible in one of the specimens.
The two specimens examined by Mr. Bartsch measure 8.2 x 15 mm.
The number of ribs is not stated.
These specimens are intermediate in character between the smaller
and more strongly-wrinkled examples of A. rhyssa, collected by Mr.
Ashmun, and the following form.
Thirteen examples, all dead and bleached, were taken by Mr. F. J.
Phillips on the north slope of a small brook near Nogal Creek, at an
elevation of 7500 ft., Lincoln Co., N. M., and recently sent to one
of us, one being drawn in figs. 1, 2 of plate VIII. In general char-
acter the lot is quite uniform. There are 30 to 35 ribs on the last
whorl ; in a few cases as many as 40, by splitting or breaking up of a
number of ribs. Ten of them, unbroken, measure as follows :
Alt. 8.1, 8, 7.9, 7.5, 8, 7.8, 7.8, 7.5, 7.1, 6.6 mm.
Diam. 13, 12.9, 12.8, 12.25, 12.2, 12, 12, 12, 12, 11.5 mm.
In 8 out of 10 fully adult shells a parietal tooth is developed, varying
from a mere trace in some to a well-developed though small tooth in
three or four. In two fully adult shells there is no trace of a tooth.
A low basal tooth is present in all, but in two or three it is barely
perceptible. The largest shell has 5^ whorls, the smallest barely 5.
This colony has the character of townsendi — small size and heavily-
ribbed surface — more emphatically expressed than in the original
townsendi, and compared directly with rhyssa it appears to be quite
distinct; yet it is connected with rhyssa through the more finely-
ribbed forms from above Ruidoso, so that the use of a trinomial
seems to express its relations better than an arbitrary specific
separation.
ASHMUNELLA RHTSSA (Ckll.) Plate VIII, figs. 3, 4, 5.
Professor Cockerell having found the unique type specimen of
THE NAUTILUS. 135
A. r. hyporhyssa kindly sent it for study. It is wholly bleached,
has lost the cuticle, and measures, alt. 9, diam. 15 mm., umbilicus
3.1 mm., and has 5^ whorls. There is a very slight basal lip-tooth,
and a small oblique parietal tooth, about 1.8 mm. long. The
sculpture is decidedly finer than in A* rhyssa, agreeing closely with
specimens of A. r. miorhyssa collected by Ashmun, which, however,
differ by having a slighly smaller umbilicus, 2.3 to 2.5 mm. wide in
a shell of nearly 16 mm. diam., and the aperture in miorhyssa is
slightly larger. I can find no other differences between miorhyssa
and hyporhyssa.
A. r. hyporhyssa was taken by Prof. C. H. T. Townsend " on the
lower slopes of White Mt., above head of Ruidoso, Aspen belt, at
about 9500 ft."
A. rhyssa and A. r. miorhyssa are from the Sierra Blanca.
Ashmunella rhyssa edentata Ckll.
A. rhyssa hyporhyssa Pils. Proc. A. N. S., Phila., 1905, pp.
228-231, pi. 12, f. 7-13, exclusive of synonym "Polygyra r. hypo-
rhyssa " and quoted description of same.
Similar to A. r. miorhyssa, but frequently more depressed, with
wider umbilicus, and generally without a parietal tooth, which is
present in less than 5 per cent, of the specimens examined.
Sacramento Mountains, at various elevations, as given in the paper
cited above. While but weakly differentiated from the Sierra Blanca
forms, it is well to have a name for the Sacramento race, common in
James Canyon near Cloudcropt.
I formerly united this race and A. r. hyporhyssa, in the absence of
the type of the latter.
ABALONES AND THE EARTHQUAKE.
BY ROBERT E. C. STEARNS.
The Japanese abalone fishermen who have made their head-
quarters at Carmel Bay, a few miles south of Monterey, had planned
to extend their operations to the more southerly part of the coast, on
the shore of San Luis Obispo County. This was in August. Upon
examination later on of the rocky region near Morro, where they
expected to make a great haul, it was found that the sea-bed was
coated with a greasy slime of a supposed bituminous character, which
136 THE NAUTILUS.
had killed the abalones. Abalones were found in abundance for
miles along the shore, so it was reported, but all dead. As the
" meats," as the dried soft parts are called, constitute the chief profit
of the fishery, no doubt the Japanese were disappointed.
The earthquake of April 18th, which was felt very slightly at
Morro, was credited with the killing, though it may have been caused
by some subsequent seismic disturbance. The abalones that occur
along this part of the shore belong principally to the species Haliotis
rufescens and H. cracherodii.
The Japanese parties engaged in the fishery at various places, in
some instances, use a diving-suit for working in or below the lamina-
cian zone, and are stripping some localities quite thoroughly. The
Chinese are said to confine their collecting generally to shallow
water. The greater part of the dried meats, if not all, are exported
to China.
There is one cannery in or near San Pedro where the abalones are
packed, the can-labels bearing the trade name of " Eno-labo," which
it will at once be seen is merely Obalone spelled backwards. Here
is an instance on a small scale where commercialism overreaches
itself. The dear public don't know what " Eno-labo " means, and,
as my grocer tells me, don't buy the goods. Fresh-packed "Eno-
labo," however, is quite palatable and wholesome food, good enough
for anybody.
Los Angeles, Cal., February 1, 1907.
EPIPHRAGMOPHOBA (MICRAEIONTA) HUTSONI N, 8P.
BY GEO. H. CLAPP.
Shell openly umbilicate ; rather thin but strong, smooth and shin-
ing when fresh ; reddish -horn color on the upper surface, much paler
to creamy-white below, with a rather narrow brown band, about
1 mm. wide, above the periphery, visible only on the last 1^ whorls,
the band margined above and below by wider white bands, the lower
shading off into the light-colored base. Whorls about 4^, the inner
ones convex, the last somewhat flattened above, well rounded below
and at the periphery. The embryonic shell consists of If whorls,
THE NAUTILUS. 137
terminating with an inconspicuous whitish line ; the first ^ whorl is
depressed and highly polished ; the remaining embryonic whorls are
evenly and densely covered with elongate-oval papillae arranged in
spiral lines. In the following 1^-2 whorls, beginning the neanic
stage, the epiconch bears exceedingly delicate, very short hairs ar-
ranged in obliquely descending series, but less regular than the
papillae of the embryonic whorls. These are entirely lost in mature
shells, although the hair-scars are generally visible on fresh speci-
mens. The body-whorl shows the usual slight growth-lines, but is
otherwise smooth except for varical impressions showing resting
periods, of which there are usually 2 on the body-whorl and others
clear up to the nuclear whorls. The body-whorl enlarges rapidly,
and is suddenly deflected and expanded at the aperture, which is
large, almost round, oblique, with a slightly reflected and thickened
margin ; lips widely expanded at the columella and partly reflected
over the umbilicus. Ends of lip converging, forming about four-
fifths of a circle. Gr. diam. 14^, lesser 11^, alt. 8 mm. Aperture
7 mm. high, 7 mm. wide. Umbilicus 2^ mm.
The figured specimen measures, alt. 7.5, diam. 14 mm. The
largest and smallest shells that I have seen measure 15 and 12^
mm. diam. respectively; both are dead shells.
Collected by Geo. S. Hutson, after whom the species is named,
about 8 miles from Quartzsite, Yuma Co., Ariz., in the foothills, at
an alt. of about 1600 ft.
Types, No. 5659 of my collection ; co-types, coll. A. N. S. P.
The sculpture of the embryonic whorls of this beautiful little shell
puts it in the group of Sonorella wolcottiana Bartsch (Smiths. Misc.
Coll., xlvii, p. 187, pi. xxviii). A large series from young shells of
2^ whorls to adults show the sculpture and epiconch in all stages,
the latter being so thin and delicate, however, that it apparently
entirely disappears by the time the shell has completed the third
whorl. The shells are remarkably uniform in size and color, and
most fresh specimens show occasional translucent spots scattered
over the body-whorl. The resting periods are shown by well-
marked varical impressions at all stages of growth, as is common in
most species from a semi-arid region. The body of the animal is
intensely black, with a gray mantle.
138 THE NAUTILUS.
ON THE SOFT ANATOMY OF E. (MICKAEIONTA) HUTSONI.
BY H. A. PILSBRY.
The animal is purplish -black, with gray mantle-edge. The sur-
face is finely and evenly granulose. There is a weak impressed line
on the ridge of the tail, and a pair of contiguous dorsal grooves,
scarcely noticeable except between the tentacles. Foot-margin very
narrow, hardly differentiated. The sole is conspicuously tripartite
in color, the sides being purplish or slaty-black, while the middle
field is of a much paler gray tint. The pale area is somewhat wider
than the dark sides.
The genitalia, fig. 6, show a very short penis, swollen basally, an
epiphallus of about the same length or somewhat larger, and a
flagellum longer than either. The long retractor muscle is inserted
on the epiphallus, and distally on the lung floor. At the base of the
vagina a well-developed dart sack (d. s.) is inserted ; on each side of
its base enter the ducts of the mucous glands, which ascend as high
as the dart sack and then descend, their flattened, glandular portions
being bound to the vagina and atrium. The general relations of the
dart sack and mucous glands are shown diagramatically in fig. 8.
As usual, the two glands are unequal. The spermatheca is ovate,
on a long, slender duct. Length of penis 2 mm. ; of epiphallus 2.8 ;
flagellum 4 mm. ; length of spermatheca and duct 16.5 mm.
The right ocular retractor passes between the cf and 9 branches
of the genitalia.
The general features of the alimentary tract are as usual in the
Helices. The crop is long, the salivary glands equally lengthened,
lying on the upper side of the crop, and concrescent throughout their
length.
The jaw is of the usual arcuate shape, with only three ribs, grouped
in the median third (fig. 7).
The radula has 17,10,1,10,17 teeth (pi IX, fig. 5). The cen-
tral and lateral teeth have simply mesocones without side cusps.
The marginal teeth have the ectocones developed, and in the outer
ones the larger cusp is bifid. I noticed no teeth with the ectocone
bifid. The transition from lateral to marginal teeth is quite gradual.
The venation of the lung is faintly outlined with gray pigment.
The kidney is about half as long as the lung, and nearly twice the
length of the pericardium.
THE NAUTILUS, XX.
PLATE VIII.
8
PILSBRY: ASHMUNELLA FROM NEW MEXICO.
PRESTON: CALLIOSTOMA AND CYPR^EA.
THE NAUTILUS XX.
PLATE IX.
mm i
c /
v
8
CLAPP: EPIPHRAGMOPHORA HUTSONI.
THE NAUTILUS. 139
While the shell of this species has all the appearance of a Sonor-
ella, the soft anatomy shows that it belongs to the subgenus Micra-
rionta, of Epiphragmophora, represented by E. gabbi, facia, steamsi-
ana, etc., the genitalia being typical for that group. In Sonorella I
have figured the anatomy of eight species and subspecies. All agree
in having a well-developed penis, and the flagellum is excessively
short or wanting, usually about half a millimeter long, in one case
one mm. long, but its length is always a very small fraction of that
of the epiphallus. In no Sonorella is there any trace of dart sack or
mucous glands. This is very unlike these organs in E. hutsoni.
The unexpected internal anatomy of this species emphasizes the
uncertainty of dealing with Sonorella-libe Helices without examina-
tion of the soft anatomy. How many of the supposed Sonorellas of
southeastern California may really prove to belong to Micrarionta is
problematic, but perhaps all those with the embryonic sculpture like
E. hutsoni will eventually be removed from Sonoralla.
EXPLANATION OF PLATE IX.
Figs. 1-3. Epiphragmophora hutsoni, shell.
4. Apex, enlarged.
5. Teeth.
6. Genitalia.
7. Jaw.
8. Diagram of dart sack and mucous glands, d. s., dart sack ;
epi., epiphallus ; ft., flagellum ; m. gl., mucous glands.
DESCRIPTIONS OF CYPRAEA BERNARDINO AND CALLIOSTOMA
CARNICOLOR N. SP.
BY H. B. PRESTON.
CYPRAEA BERNARDINO n. sp. PI. VIII, fig. 8.
Shell oval, with a well-defined dorsal line dividing into a small
fork near the anterior extremity; dorsal surface brownish-gray
flecked with white spots and streaks ; sides white, sparsely spotted
with chestnut, a number of dashes of the same color appearing at the
anterior extremity and slightly crenulated at the margins of the dor-
sal surface ; extremities obtuse ; base white ; columella somewhat
straight, bearing seventeen rather fine, white teeth ; peristome very
slightly curved, having fifteen teeth ; aperture narrow.
140 THE NAUTILUS.
Length 28.50 mm., greatest breadth 20 mm.
Hab.: Celebes (?).
A striking shell, whose nearest ally is perhaps Cypraea turdus Lk.
From this, however, it may be easily distinguished by the remark-
able white flecking on the dorsal surface, by the finer and more
numerous teeth on the columella and the narrower and straighter
aperture ; moreover, it is much flatter than is the case with Cypraea
turdus.
CALLIOSTOMA CARNICOLOR n. sp. PI. VIII, figs. 6, 7.
Shell conical, keeled, imperforate, somewhat glossy, flesh-colored ;
spire concave ; whorls 8-9, the last three rapidly increasing and
much flattened, sculptured with nodulous spiral ridges, almost every
alternate nodule being of a rich chestnut-brown ; sutures not well
defined above, and only slightly impressed between the last whorls ;
base very inflated and sculptured with closely-set spiral ridges inter-
cepted by lines of growth, thus presenting a coarsely, granular
appearance ; columella arched, reflexed outwards, forming a thick
callosity over the umbilical area, a slight callosity extending upwards
to the lips above ; peristome simple ; aperture subquadrate ; interior
of shell nacreous and irridescent.
Alt. 41 mm., diam. maj. 53 mm.
Aperture, alt. 25 mm., diam. maj. 25 mm.
Hab. : Celebes (?).
The above shell at first sight much resembles Calliostoma cun-
ninghami Gray, from New Zealand ; the spire is, however, more
concave, the last whorls are much more flattened and expanded out-
wards, and the nodulous spiral ridges are far less numerous than is
the case with that species ; moreover, the inflation of the base easily
separates it from C. cunninghami.
DESCBIFTION OF A NEW BIFIDAEIA (B. AGNA).
BT H. A. PILSBRY AND E. O. VANATTA.
When working on Bifidaria pentodon some time ago we found one
specimen of a species evidently distinct though related to that, from
" Silver Lake, Kansas," collected by Mr. J. B. Quintard. Recently
a few more examples of the same form were found in river d£bri»
THE NAUTILUS.
141
taken last October at Trinidad, in southeastern Colorado, by Mr.
J, H. Ferriss and the senior author.
BlFIDARIA AGNA n. Sp.
The shell is rimate, cylindric, the last three whorls of about equal
diameter, the summit very obtuse ; surface nearly smooth; of a sperm-
aceti-whitish color. There are 4| convex whorls, the last with a
strong rounded crest behind the outer lip, rather close to it above,
but being more oblique, it is further behind the lip at the base.
Behind this crest the whorl is a little flattened laterally, and shows
the lower palatal plica, though as a white spot. The aperture is
brought forward nearly in line with the ventral convexity of the
whorls. The peristome is thin, slightly expanded, strengthened a
short distance within with a low callous rib. The parietal lamella
appears distorted and angular in front view ; seen from the base it is
seen to consist of an angular and a parietal lamella, completely
united inwardly. The columellar lamella is sinuated or bent and
strongly developed. The basal fold is strong, arising on the callus
but extending inward beyond it. The lower palatal fold is large and
pliciform, more or less immersed, standing chiefly inward from the
palatal callus. The upper palatal fold is much smaller, and stands
rather near the lower, upon the palatal callus. There is a small or
minute suprapalatal fold, also on the callus.
Length 1.75, diam. 1 mm.
Trinidad, Colorado, type no. 93052 A. N. S. P. ; also Silver Lake,
Kansas.
This species resembles some forms of B. pentodon (Say), especially
those from northern Alabama, in which the lower palatal fold is long
142 THE NAUTILUS.
and enters somewhat beyond the palatal callus ; but it differs by the
distinctly bifid parietal tooth. The very strong crest behind the lip
is characteristic of B. agna. Like the related forms, this species has
the somewhat translucent whiteness of spermaceti or paraffin.
In the specimen from Kansas the teeth are somewhat smaller than
in the type. It may be less mature.
COKBESPONDENCE.
Editor Nautilus : I have to thank Mr. Burnett Smith for his ex-
tended review of my paper on American Volutidse in the March
NAUTILUS. There are, however, a few points on which further
light is desirable, and I wish to note them briefly.
1. So far from this being the general revision of the family upon
which I have been for some time at work the recent publication is
only a small part of it, which I thought I had made clear in my
remarks on page 341. My regret at not being able to illustrate the
paper is quite as great as that of my reviewer, but only those cog-
nizant of the facts know, since we lost Dr. McConnell, how many
fruitless, or almost fruitless, efforts have been made to obtain a com-
petent draughtsman. An important paper has been for more than a
year at a complete standstill, owing to the absence of an artist who
could do the drawings.
2. I pointed out in 1890 the identity of the so-called Athleta
tuomeyi with Volutilithes petrosa Conrad, and ascribed the deformity
to some special conditions of the environment. We have some two
thousand specimens in the National Museum, a part of which are
normal. But while the most conspicuously deformed specimens are
from Wood's Bluff and the lower Eocene generally, we have also
distorted forms from Jackson, Miss., and several other Jacksonian
localities, and one specimen from the Claibornes ands. These I shall
be happy to show Mr. Smith if he can pay us a visit.
That the true Athleta rarispina bears any such relation to Valuta
spinosa Lam. as V. tuomeyi does to petrosa is impossible, for Mr.
Smith's suggestion is incompatible with the fact that A. rarispina is
confined to the upper Oligocene, Miocene and Pliocene of Europe,
while V- spinosa is restricted to middle Eocene (Calcaire Grossiere).
There is no species contemporaneous with the Athleta which could
take the place of V- spinosa in such a relation, and, further, the two
species of Athleta are normal and not abnormal shells.
THE NAUTILUS. 148
3. The remarks about Plejona which were made by both Mr.
R. B. Newton and Mr. Smith indicate unfamiliarity with nomencla-
torial questions, which indeed are sometimes sufficiently puzzling.
But there is no ambiguity or difficulty in the case of Plejona, if the
rules are complied with. Bolten's genus (for the period remarkably
homogeneous) contains 24 species, all of which are identifiable, in
terms of the older authors, and only one of which is a fossil. The
first (fossil) species is based upon four very fair figures of 4< spiny
murices," which Argenville referred to one species, but which repre-
sent three or four, as species are now reckoned. If we prefer to take
our type from the first species in Bolten's list, it reduces itself to a
simple process of elimination which leaves us with V- spinosa (one
of those figured) as the type. If we regard the solution as vague,
there still remains my absolute right to revive Bolten's name for any
species contained in his list, and for which there is no prior valid
name. This I did several months before Mr. Newton proposed
Volulospina for the same shell.
If we reject Bolten's P. fossilis as vague, we are left to take
Voluta ebraea, his second species, as type, which involves the loss of
the generic name Voluta (Lam.), as now universally used, which it
seems to me would be foolish, since the preceding method enables us
to retain Voluta for V- musica and its allies without upsetting any
accepted name.
I regard Athleta (rarispina) Conrad as a good genus, perfectly
distinct from Plejona (spinosa) Bolten, as well as from the F. petrosa
deformities. The latter were erroneously referred to the genus
Athleta, which they really only superficially resemble, and conse-
quently any attempt to use Athleta for the spinosa group is fallacious.
Athleta does not occur in America, but there are probably three
European species. WM. H. DALL.
Smithsonian Institution, March 6, 1907.
PUBLICATIONS RECEIVED.
WEST AMERICAN MITRID^E By Mrs. M. Burton Williamson
(Proc. Biol. Soc., Washington, xix, Dec., 1906). Mitra idae, fultoni,
lowi, and the Peruvian M. orientalis are discussed at length and
with the exception of M. lowi, figured. A useful work, since these
black Mitres have been involved in some confusion.
144 THE NAUTILUS.
THE UNIONID^E OF KANSAS, part I, by Richard E. Scammon,
(Kansas University Science Bulletin, iii, no. 9). " In this catalogue
are listed 61 species and 5 subspecies. There is no doubt that future
collecting will add many more species to the list. The author has
tried to make certain of the identification and localities of every
species listed, and with three exceptions has examined native speci-
mens of each." Nearly all of the species are illustrated by line
drawings occupying 23 plates, and all are described. Since only
about 40 species were previously known, it will be seen that one-third
of those now recorded are new to the state. Mr. Scammon proposes
two new descriptive terms : " interdentum" for that part of the
hinge-plate lying between the pseudocardinal and lateral teeth, and
" branchial outline" to designate the slight groove in the cavity of
the more solid Unios, caused by the edge of the branchiae. It is
proposed in future papers to discuss the geographic distribution and
soft anatomy of the species.
NOTES.
In a recent letter Prof. Josiah Keep says : " The principal part of
my edition of ' West American Shells ' was destroyed in the great
San Francisco fire, so there are no more copies in the book stores.
Fortunately the cuts were stored in my home, and may be used again
if there should be a considerable demand for a revised edition. Of
this matter, however, I do not feel certain, though I have received
some very kind letters upon the subject." We are sorry to learn of
Prof. Keep's great loss, and hope that he will receive sufficient
encouragement to undertake a new edition of his work.
ZONITOIDES ALLIAR1A IN COLORADO A COUple of weeks ago
Guy H. Mason found in the Knudson greenhouse, Boulder, Colo., a
specimen of Zonitoides alliaria Drap., a species well known to Euro-
pean conchologists, but of which American records seem to be rare,
and confined, so far as they have come to my attention, to green-
houses. It occurs to me that the rarity of records may have resulted
from American conchologists unfamiliar with the species placing it
in their collections under some other name, particularly Z. cellaria.
We are indebted to Dr. Wm. H. Dall for confirmation of the identi-
fication, he having compared it with British specimens as named by
Jeffreys. It was at once recognized by Prof. Cockerell as of that
species, but his long absence from Europe made him feel somewhat
uncertain about it without opportunity to make direct comparisons.
The specimen is now in the University of Colorado Museum —
JUNIUS HENDERSON.
MBL WHOI LIBRARY
WH 17UL 0